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Full text of "An illustrated flora of the northern United States, Canada and the British possessions : from Newfoundland to the parallel of the southern boundary of Virginia and from the Atlantic Ocean westward to the 102nd meridian"

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ILLUSTRATED FLORA 


OF THE 


NORTHERN UNITED STATES, CANADA 
AND) THE BRPMSH POSSESSIONS 
ae NEWFOUNDLAND TO THE PARALLEL OF THE SOUTHERN BOUNDARY OF 


VIRGINIA, AND FROM THE ATLANTIC OCEAN WESTWARD 
TO THE 102p MERIDIAN 


BY 


ee tHaNI PLY LORD BRITTON, Pu.D.,.Sc:D; LES: 


DIRECTOR-IN-CHIEF OF THE NEW YORK BOTANICAL GARDEN; PROFESSOR IN COLUMBIA UNIVERSITY 


AND 


HON. ADDISON BROWN, A.5.,, LL. 


PRESIDENT OF THE NEW YORK BOTANICAL GARDEN 


THE DESCRIPTIVE TEXT 


CHIEFLY PREPARED BY PROFESSOR BRITTON, WITH THE ASSISTANCE OF SPECIALISTS IN 
SEVERAL GROUPS; THE FIGURES ALSO DRAWN UNDER HIS SUPERVISION 


SECOND EDITION—REVISED AND ENLARGED 


IN THREE VOLUMES 7 bd tee 
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Vom EA, a 4 ad 
GENTIANACEAE TO COMPOSITAE 
GENTIAN TO THISTLE 


NEW YORK 
CHARLES SCRIBNER’S SONS 
1913 


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CopyRIGHT, 1913 


By Natuanrt L. Britton anp HELEN C. Brown, a 


Executrix of the Estate of 


Appison Brown, deceased. 


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15. GENTIANACEAE 


CONTENTS OF VOLUME ITI. 


16. MENYANTHACEAE 17 
17. APOCYNACEAE 19 
18. ASCLEPIADACEAE 23 
19. DicHONDRACEAE 39 
20. CONVOLVULACEAE 40 
21. CUSCUTACEAE 48 
22. PoLEMONIACEAE 52 


23. HyDROPHYLLACEAE 65 


24. BORAGINACEAE 
25. VERBENACEAE 


Petals wholly or partly united, rarely separate or wanting (continued) 


26. 
27. 


Gamopetalae (continued) 


. LABIATAE 

. SOLANACEAE 

. SCROPHULARIACEAE 
. LENTIBULARIACEAE 
. OROBANCHACEAE 

. BIGNONIACEAE 

. MARTYNIACEAE 

. ACANTHACEAE 

. PHRYMACEAE 

. PLANTAGINACEAE 
. RUBIACEAE 


1-560 


99 
154 
174 
224 
233 
236 
239 


239° 


241 
245 
250 


ENGLISH FAMILY NAMES 


15. GENTIAN FAMILY I 
16. BUCKBEAN FAMILY 17 
17. DocGBANE FAMILY 19 


18. MILKWEED FAM 


iLY . 23 


19. DicHOoNDRA FAMILY 39 


20. MorNING-GLORY 


FAMILY 40 


21. DopDER FAMILY 
22. PHLox FAMILY 


23. WATER-LEAF FAMILY 65 


24. BoraAGE FAMILY 


25. VERVAIN FAMILY 05 


SUMMARY 


560 


ENGLISH INDEX AND PoPpuLAR PLAntT NAMES 


28. 
20. 
30. 
Bie 


22) 


GENERAL INDEX OF LATIN GENERA AND SPECIES 


Mint FAMILY 
Potato FAMILY 
Ficwort FAMILY 


99 
154 
174 


BLADDERWORT FAMILY 224 
BrooM-RApPE FAMILY 233 


TRUMPET-CREEPER 
FAMILY 

Unicorn PLANT 
FAMiLY 


. ACANTHUS FAMILY 
4. LopsEED FAMILY 
. PLANTAIN FAMILY 


° is used after figures to indicate feet. 

“is used after figures to indicate inches. 
“is used after figures to indicate lines, or twelfths of an inch. 
” over syllables indicates the accent, and the short English sound of the vowel. 
“ over syllables indicates the accent, and the long, broad, open or close English sound of 


the vowel. 


In THE METRIC SYSTEM. ; 
The metre = 39.37 inches, or 3 feet 3.37 inches. 


The decimetre = 3.94 inches. 
The centimetre = 3 of an inch, or 4i lines, 
The millimetre = z, of an inch, or # a line. 


24 millimeters = 1 line. 


236 


239 
239 
241 
245 


SYMBOLS USED 


iL nearly 


. CAPRIFOLIACEAE 
. ADOXACEAE 

. VALERIANACEAE 
. DIPSACEAE 

. CUCURBITACEAE 

. CAMPANULACEAE 
. LOBELIACEAE 

. CICHORIACEAE 

. AMBROSIACEAE 

. COMPOSITAE 


I-560 


. Mapper FAMILY 
. HONEYSUCKLE 


FAMILY 


. MoscHATEL FAMILY 
. VALERIAN FAMILY 

. TEASEL FAMILY 

. GourpD FAMILY 

. BELL-FLOWER FAMILY 
. LoBeLt1a FAMILY 

. CHIcorY FAMILY 
. RAGWEED FAMILY 
. THISTLE FAMILY 


561 


601 


256 


267 
283 
284 
288 
290 
203 
299 
304 
338 
347 


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Family 15. GENTIANACEAE Dumort. Anal. Fam. 20. 1829. 
GENTIAN FAMILY 

Bitter mostly quite glabrous herbs, with opposite (rarely verticillate) exstipu- 
late entire leaves, reduced to scales in Bartonia, and regular perfect flowers in 
terminal or axillary clusters, or solitary at the ends of the stem or branches. 
Calyx inferior, persistent, 4—-12-lobed, -toothed or -divided (of 2 sepals in Obo- 
laria), the lobes imbricated or not meeting in the bud. Corolla gamopetalous, 
funnelform, campanulate, club-shaped or rotate, often marcescent, 4—12-lobed or 
-parted, the lobes convolute or imbricated in the bud. Stamens as many as the 
lobes of the corolla, alternate with them, inserted on the tube or throat; anthers 
2-celled, longitudinally dehiscent ; filaments filiform, or dilated at the base. Disk 
none, or inconspicuous. Ovary superior in our genera, 1-celled or partly 2-celled ; 
ovules numerous, anatropous or amphitropous ; style simple, or none; stigma entire, 
or 2-lobed, or 2-cleft. Capsule mostly dehiscent by 2 valves. Seeds globose, 
angular or compressed ; endosperm fleshy, copious; embryo small, terete or conic. 


About 70 genera and 700 species, widely distributed, most abundant in temperate regions. 
Leaves normal; corolla-lobes convolute in the bud. _ 
Style filiform ; anthers usually twisting or recurving when old. 


Corolla salverform. 1. Centaurium, 
Corolla rotate. 2. Sabbatia. 
Corolla'campanulate-funnelform. 3. Eustoma, 


Style short, stout or none; anthers remaining straight. 
Corolla without nectariferous pits, glands or scales. 
Corolla funnelform, campanulate or clavate. 
Corolla without plaits in the sinuses; calyx without an interior membrane, 
4. Gentiana. 
Corolla with plaits in the sinuses; calyx with an interior membrane. 5. Dasystephana 


Corolla rotate. 6. Pleurogyna, 
1-2 nectariferous pits, glands or scales at the base of each corolla-lobe. 
Corolla rotate, a fringed gland at each lobe. 7. Frasera. 
Corolla campanulate, spurred at the base. 8. Halenia. 
Leaves, at least those of the stem, reduced to scales; corolla-lobes imbricated in the bud. 
Calyx of 2 foliaceous spatulate sepals ; upper leaves normal, 9. Obolaria, 
Calyx of 4 lanceolate sepals; leaves all reduced to scales. 10. Bartonia, 


1. CENTAURIUM Hill. Brit. Herb. 62. 1756. 


[ERYTHRAEA Neck. Elem. 2: I0. 1790.] 

Herbs, mostly annual or biennial, with sessile or amplexicaul leaves, and small or middle 
sized, commonly numerous, pink, white or yellow flowers in cymes or spikes. Calyx tubular, 
5-4-lobed or -divided, the lobes or segments narrow, keeled. Corolla salverform, 5-4-lobed, 
the tube long or short, the lobes spreading, contorted, convolute in the bud. Stamens 5 or 4, 
inserted on the corolla-tube; filaments short-filiform; anthers linear or oblong, becoming 
spirally twisted. Ovary 1-celled, the placentae sometimes intruded; style filiform; stigma 
2-lobed. Capsule oblong-ovoid or fusiform, 2-valved. Seed-coat reticulated. [Latin, 100 
gold pieces, with reference to its supposed medicinal value. ] 


About 25 species, natives of the Old World, western North and South America, and in the West 
Indies. Besides the following, about 8 others occur in the western and southwestern parts of the 
United States, Type species: Gentiana Centaurium L, 


Flowers spicate-racemose. 1. C. spicatum, 
Flowers cymose or cymose-paniculate. 
Basal leaves tufted. 2. C. Centaurium. 


No tuft of basal leaves. 
Corolla-lobes 11%4”-2%” long. 


2 GENTIANACEAE. Vot. III. 


Flowers short-pedicelled ; naturalized species. 3. C. pulchellum. 
Flowers slender-pedicelled ; native western species. 
Leaves oblong to linear-oblong. 4. C. exaltatum. 
Upper leaves mere subulate bracts. | 5. C. texense. 
Corolla-lobes 3%”-5” long. 6. C. calycosum, 


1. Centaurium spicatum (L.) Fernald. Spiked 
Centaury. Fig. 3330: 
Gentiana spicata L, Sp. Pl. 230. 1753. 
Erythraea spicata Pers. Syn. 1: 283. 1805. 


Centaurium spicatum Fernald, Rhodora 10: 54. 1908. 


Annual, glabrous, erect, strict, usually branched, 6-18’ 
high, Leaves oblong or lanceolate-oblong, sessile, obtusish 
at the apex, clasping at the base, 3’-13’ long, 2’—7” wide; 
flowers pink, sessile, distant and spicate-racemose on the 
mostly simple and leafless branches, about 8” long; tube of 
the corolla somewhat longer than the subulate calyx-seg- 
ments, 2-3 times as long as the linear-oblong lobes; capsule 
4’-5” long. 


Coast of Nantucket, and at Portsmouth, Va. Naturalized 
from Europe. May-Sept. 


2. Centaurium Centatrium (L.) W. F. Wight. 
Lesser or European Centaury. Bitter-herb. 
Bloodwort. Fig. 3331. 


Gentiana Centaurium L. Sp. Pl. 229. 1753. 
Erythraea Centaurium Pers. Syn. 1: 283. 1805. 
C. Centaurium W. F. Wight, Contr. Nat. Herb. 11: 449. 1906. 
Annual, glabrous, erect, usually branched, 6-15’ high. 
Leaves oblong, apex obtuse, the base narrowed; the lower 
forming a basal tuft, 1’-22’ long, 3’-6’ wide; stem leaves 
smaller, distant, rounded at the sessile or slightly clasping 
base: flowers numerous, 6-8” long, nearly sessile, in com- 
pound terminal mostly dense bracteolate cymes; corolla- 
lobes obtuse, 23”-3” long, about as long as the calyx-seg- 
ments and one-third to one-half as long as the corolla- 
tube; stigmas oval. 


In waste places, Nova Scotia and Quebec to Massachusetts, ; DP 
Illinois and Michigan. Naturalized from Europe. Earth-gall. Sanctuary. June—Sept. 


3. Centaurium pulchéllum (Sw-.) Druce. Branching Centaury. Fig. 3332. 


Gentiana pulchella Sw. Act. Holm. 1783: 84. f. 8, 9. 1783. 
Gentiana ramosissima Vill. Hist. Pl. Dauph. 2: 530. 1787. 
Erythraea ramosissima Pers. Syn, 1: 283. 1805. 

Erythraea pulchella Fries, Novit. 74. 1828. 

C. pulchellum Druce, Fl. Oxf. 342. 1897. 


Annual, glabrous, much branched, 3-8’ high. Leaves 
oval, ovate or lanceolate, the lower mostly obtuse, 3-8” 
long, the upper usually acutish or acute and smaller; 
no basal tuft of leaves; flowers pink, cymose-paniculate, 
all or nearly all of them short-pedicelled, 5”-6” long; 
tube of the corolla 13-2 times longer than the calyx- 
segments, its lobes oblong, obtuse, 14’-2” long; stigma 
oval; anthers oblong. 

In fields and waste places, southern New York to Penn- 


sylvania, Illinois and Maryland; also in the West Indies, 
Naturalized from Europe. June—Sept. 


GENUS I. GENTIAN FAMILY. 3 


4. Centaurium exaltatum (Griseb.) W. F. Wight. 
Tall or Western Centaury. Fig. 3333. 


Cicendia exaltata Griseb. in Hook. Fl. Bor. Am, 2: 69, pl, 157.' 
1834. 
Erythraea Douglasii A. Gray, Bot. Cal. 1: 480. 1876. 


Erythraea exaltata Coville, Contr. Nat. Herb. 4: 150. 18093. 
C. exaltatum W. F. Wight, Contr. Nat. Herb. 11: 449. 1906. 


Annual, erect, glabrous, branched, 6-18’ high, the 
branches few, erect, slender. Leaves oblong or linear- 
oblong, sessile, mostly acute at both ends, 5-10” long, 
distant, the basal not tufted; flowers few, terminal and 
axillary, all slender-pedicelled, 6’-8” long; tube of the 
corolla about one-third longer than the calyx-segments, 
the lobes oblong, obtuse, 14’-2” long. 

In sandy soil, western Nebraska to Wyoming, Washington, yy, —— 
Arizona and California. May-—Sept. ; 
- 5. Centaurium texénse (Griseb.) Fernald. 
Y Texan Centaury. Fig. 3334. 


i i Erythraea texensis Griseb; Hook. Fl. Bor, Am. 2: 58. 
4 vy) 1838. 
f < VY C. texense Fernald, Rhodora 10: 54. 1908. 
¢ \ <= Annual, corymbosely branched above, slender, 2’-&’ 
} high. Stem-leaves linear or linear-lanceolate, 8” 
INV long or less, acute, sessile, the upper ones reduced 
ff to subulate bracts; pedicels slender, as long as the 
| Vi calyx or longer; calyx 4-5” long, its lobes subulate; 
y} corolla light rose color, its tube longer than the 
calyx, its acute oblong to oblong-lanceolate lobes 
about half as long as the tube; capsule longer than 
the calyx. 


In rocky soil, Missouri to Texas. May-—Sept. 


) 


6. Centaurium calycosum (Buckley) Fernald. Buck- 
ley’s Centaury. Fig. 3335. 


Erythraea calycosa Buckley, Proc. Acad. Phil. 1862: 7. 1863. 
Centaurium calycosum Fernald, Rhodora 10: 54. 1908. 


Annual, corymbosely branched or sometimes simple, 2° high 
or less, the branches ascending or spreading. Leaves oblong 
to spatulate or linear, 4’-13’ long, acute, sessile; pedicels as 
long as the calyx or longer; calyx 4’-5” long, its lobes nar- 
rowly linear; corolla pink, its tube a little longer than the . 
calyx, its oval or oblong obtuse lobes nearly as long as the 


tube. 


In wet or moist soil, Missouri to Texas, Mexico and New Mexico. 
April-June. 


2. SABBATIA Adans. Fam. Pl. 2: 503. 1763. 


Annual or biennial erect usually branched glabrous herbs, with opposite or sometimes 
verticillate sessile or rarely petioled or clasping leaves, and rather large terminal and solitary 
or cymose pink rose or white flowers. Calyx 4-12-parted or -divided, the tube campanulate, 
sometimes very short, the lobes or segments usually narrow. Corolla rotate, deeply 4-12- 
parted. Stamens 4-12, inserted on the short tube of the corolla; filaments filiform, short; 


4 GENTIANACEAE. Vot. III. 


anthers linear or oblong, curved, revolute or coiled in anthesis. Ovary 1-celled, the placentae 
intruded; style 2-cleft or 2-parted, its lobes filiform, stigmatic along their inner sides. Cap- 
sule ovoid or globose, 2-valved, many-seeded. Seeds small, reticulated. [In honor of L. 
and C. Sabbati, Italian botanists, according to Salisbury, Parad. Lond. /. 32, therefore 
Sabbatia, though Adanson’s spelling was Sabatia.] 

About 18 species, natives of eastern North America, the West Indies and Mexico. Besides the 
following, some 6 others occur in the southern United States. Type species: Chironia dodecandra L, 


Flowers normally 4—5-parted, sometimes 6—7-parted, 
Branches opposite. 
Style 2-parted to below the middle or nearly to the base; flowers white. 


Leaves lanceolate or ovate, acute; flowers 8”—12” broad. . S. lanceolata, 


Lan] 


Leaves linear-oblong or lanceolate, obtuse; flowers 6’—9” broad. 2. S. paniculata, 
Style 2-cleft to about the middle; flowers normally pink. 

Leaves linear-lanceolate, sessile ; stem slightly 4-angled. 3. S. brachiata. 

Leaves ovate, cordate-clasping ; stem strongly 4-angled. 4. S. angularis. 


Branches alternate, the lower sometimes opposite in nos. 5 and 6, 


Calyx-segments foliaceous, longer than the corolla. 5. 5S. calycina. 
Calyx-segments linear or lanceolate, not longer than the corolla. } 
Calyx-tube 5-ribbed; flowers 1-2’ broad; leaves ovate to oblong. 6. S. campestris, 


Calyx-tube scarcely ribbed ; flowers 17-14’ broad; leaves linear to lanceolate. 
Calyx shorter than the corolla; style 2-parted. 
Flower pink with a yellow eye (rarely white) ; corolla-segments obovate. 
7. S. stellaris. 
Flowers white, very numerous; corolla-segments spatulate or oblanceolate. 
8. S. Elliotiti. 


Calyx-segments filiform, as long as the corolla; style 2-cleft. 9. S. campanulata. 
Flowers normally 8—12-parted, 114’-2¥4’ broad. to. S. dodecandra, 


1. Sabbatia lanceolata (Walt.) T. & G. 
Lance-leaved Sabbatia. Fig. 3336. 


Chironia lanceolata Walt. Fl. Car. 95. 1788. 
S. lanceolata T, & G.; A. Gray, Man. 356. 1848. 


Stem branched above, or simple, slender, some- 
what 4-angled, or terete below, 1°-3° high, the 
branches all opposite. Leaves lanceolate to ovate, 
acute, or the lower sometimes obtuse, 3-5-nerved, 
1’-2’ long, or the lowest shorter, the uppermost 
reduced to narrow bracts; flowers white, fading 
yellowish, 8”’-12” broad, usually numerous in 
bracteolate corymbed cymes; pedicels slender, 
2-7" long; calyx-lobes filiform-linear, much 
shorter than the corolla; corolla-segments oblong 
or slightly obovate; anthers recurved; style 
2-parted; capsule ovoid, about 3” high. 


In pine-barren swamps, New Jersey to Florida. 
May-Sept. 


5 
2. Sabbatia paniculata (Michx.) Pursh. 
Branching Sabbatia. Fig. 3337. 


C. paniculata Michx. Fl. Bor. Am, 1: 146. 1803. 
S. paniculata Pursh, Fl, Am, Sept. 138. 1814. 


Stem usually freely branching, 4-angled, 
1°-23° high, the branches all opposite. Leaves 
linear, linear-oblong, or lanceolate, obtuse, 
+’-13’ long, the lower commonly shorter and 
broader, the uppermost small and bract-like; 
flowers white, 6-9” broad, usually very nu- 
merous in corymbed cymes; pedicels mostly 
short, the central flowers of the cymes often 
nearly sessile; calyx-lobes linear, not more 
than one-half the length of the corolla; co- 
rolla-segments spatulate-oblong; anthers re- 
curved or coiled; style 2-parted; capsule ob- 
long, about 3” high. 


In dry or moist soil, Virginia to Florida. May-— 
Sept. 


GENUS 2. GENTIAN FAMILY. 5 


3. Sabbatia brachiata Ell. Narrow- 
leaved Sabbatia. Fig. 3338. 


Chironia angularis var, angustifolia Michx. FI. 
Bor. Am. 1: 146. 1803. 
S. brachiata Ell. Bot. S. C. & Ga. 1: 284. 1817. 
S. angustifolia Britton, Mem, Torr. Club 5: 259. IK 


1894. 


Stem slender, branched above, slightly 4- 
angled, 1°-2° high, the branches all opposite. 
Leaves linear, linear-oblong, or linear-lanceo- 
late, obtuse, or the upper acute, sessile, 1-2’ 
long, the uppermost small and_ bract-like; 
flowers pink with a yellowish or greenish eye, 
few in the racemed or short-corymbed cymes, = 
or solitary at the ends of the branches, about 
1’ broad; calyx-lobes linear, usually more than 
one-half the length of the corolla; corolla- 
segments obovate-oblong; style 2-cleft to about 
the middle; capsule oblong, 3-4” high. 


In dry or moist soil, Indiana to Louisiana, east ) 
to North Carolina and Florida. May-—Sept. A 


4. Sabbatia angularis (L.) Pursh. Bitter-bloom. Rose-Pink. Square-stemmed 
Sabbatia. Fig: 3330. 


Chironia angularis L. Sp. Pl. 190. 1753. 
S. angularis Pursh, Fl. Am. Sept. 137. 1814. 


Stem usually rather stout and much branched, 
sharply 4-angled, 2°-3° high, the branches all 
opposite or the lowest rarely alternate. Leaves 
ovate, acute at the apex, cordate-clasping at the 
base, 9-18” long, or the lower oblong and ob- 
tuse, those of the branches smaller; flowers rose- 
pink, with a central greenish star, occasionally 
white, 1’—13’ broad, usually solitary at the ends 
of the branches; calyx-lobes linear, one-half the 
length of the corolla, or less; corolla-segments 
obovate; style 2-cleft; capsule oblong, about 3” 
high. 

In rich soil, often in thickets, New York and 
Pennsylvania to western Ontario, Michigan, Flor- 
ida, Arkansas, Oklahoma and Louisiana. Bitter 
clover. Pink-bloom. American centaury. July—Aug. 
Flowers fragrant. 


g 


5. Sabbatia calycina (Lam.) Heller. 
Coast Sabbatia. Fig. 3340. 

Gentiana calycina Lam, Encycl. 2: 638. 1786. 
C. calycosa Michx. Fl. Bor. Am. 1: 146. 1803. 
Sabbatia calycosa Pursh, Fl. Am. Sept. 138. 1814. 
S. calycina Heller, Bull. Torr. Club 21: 24. 1804. 

Stem somewhat 4-angled, freely branched, 
6-12’ high, the branches alternate, or the 
lowest sometimes opposite. Leaves oblong or 
some of them slightly obovate, obtuse or acute, 
3-nerved, 1’-2’ long, narrowed to the sessile 
base or the lower into petioles; flowers usu- 
ally few, solitary at the ends of the branches 
or peduncles, 1’-1%’ broad; calyx-lobes linear 
or spatulate, leaf-like, longer than the spatu- 
late segments of the pink rose-purple or whit- 
ish corolla; style 2-parted, capsule ovoid- 
oblong, 3-4” high. 

In moist soil, Virginia to Florida, near the 
coast. Cuba; Santo Domingo. June—Aug. 


6 GENTIANACEAE. 


high. 


7. Sabbatia stellaris Pursh. Sea or Marsh 
Pink? Figy 3342; 


Sabbatia stellaris Pursh, Fl. Am. Sept. 1: 137. 1814. 
Chironia stellata Muhl. Cat. Ed. 2, 23. 1818. 


Stem nearly terete, or slightly 4-angled, usually 
much branched, 6’—2° high, the branches alternate. 
Leaves lanceolate-oblong to linear, sessile, 2’—2’ 
long, 2’—5” wide, or the lowest smaller, some- 
what obovate and obtuse, the uppermost narrowly 
linear and bract-like; flowers 9’—-18” broad, usu- 
ally numerous, solitary at the ends of the branches 
or slender peduncles; calyx not ribbed, its lobes 
narrowly linear, shorter than or nearly equalling 
the oblong or obovate corolla-segments; corolla 
pink to white with a yellowish starry eye bor- 
dered with red; style 2-cleft to below the middle; 
capsule about 22” high. 

In salt-meadows, coast of Massachusetts to Flor- 


ida. Recorded from Maine. Called also Rose-of- 
Plymouth, July—Sept. 


6. Sabbatia campéstris Nutt. 


opposite. 


Swertia difformis L. Sp. Pl. 226. 


1817. 


S. Elliottii Steud. Nomencl. Ed. 2, 2: 489. 


ridged, 


Florida. 


Vor. lie 


Prairie 


Sabbatia campestris Nutt. Trans, Amer. Phil, Soc, 
(II.) 5: 197, 


Stem 4-angled, branched, 6’-15’ high, the 
branches alternate or dichotomous, or the lowest 
Leaves ovate, oblong or lanceolate, 
mostly obtuse at the apex, sessile or slightly 
clasping and subcordate at the base, 3-1’ long; 
flowers solitary at the ends of the branches and 
peduncles, 1’-2’ broad; peduncles, 1-2’ long; 
calyx 5-ribbed or almost 5-winged, its lobes lan- 
ceolate, acute, 6’—-12” long, about as long as the 
obovate lilac corolla-segments, becoming rather 
rigidin fruit; style 2-cleft; capsule oblong, 3-4” 


On prairies, Missouri and Kansas to Texas, 


8. Sabbatia Ellidttii Steud. Elliott's 


Sabbatia paniculata Ell. Bot. S. C. & Ga. 1: 282. 
Not Pursh, 1814. 


1841. 


Stem paniculately branched, terete or slightly 
the branches alternate. 
Lower leaves obovate or lanceolate-oblong, 
obtuse, 6’-9” long, the upper and those of the 
branches narrowly linear or subulate; flowers 
very numerous, white, solitary at the ends of 
the branches and short peduncles, 1’ broad or 
less; calyx not ribbed, its lobes subulate-linear, 
one-half as long as the spatulate or oblanceo- 
late segments of the corolla, or less; style 
2-parted; capsule about 22” high. 


In pine-barrens, Virginia and North Carolina to 
Quinine-flower. 


GENUS 2. GENTIAN FAMILY. 7 


Chironia campanulata L. Sp. Pl. 190. 1753. 
Chironia gracilis Michx. Fl. Bor. Am. 1: 146, 1803. 
Sabbatia gracilis Salisb. Parad. Lond. pl. 32. 1806. 
Sabbatia campanulata Torr. Fl, U. S. 1: 217. 1824, 
Similar to the preceding species. Stem usually 
very slender and much branched, 1°-2° high, the 
branches alternate. Leaves linear, or linear-lan- 
ceolate, 1-12’ long, sessile, acute, or the lowest 
much shorter, obtuse, oblong or oblanceolate, 
sometimes narrowed into short petioles, the up- 
permost almost filiform; flowers pink with a 
yellow eye, about 1’ broad, solitary at the ends 
of the branches and peduncles, mostly 5-parted; 
calyx-lobes filiform-linear, equalling the oblong- 
obovate corolla-segments, or somewhat shorter; 
style 2-cleft to about the middle; capsule obovoid, 


1 


about 22” high. 
In salt marshes and along brackish rivers, rarely 
in fresh-water swamps, Nantucket to Florida and 


Louisiana. Also on the summits of the southern 
Alleghanies. Bahamas; Cuba. May—Aug. 


10. Sabbatia dodecandra (L.) B.S.P. Large Marsh Pink. Fig. 3345. 


Chironia dodecandra L. Sp. Pl. 190. 1753. 

Chironia chloroides Michx. Fl. Bor. Am. 1: 147. 1803. 
Sabbatia chloroides Pursh, Fl. Am. Sept. 138. 1814. 
Sabbatia dodecandra B.S,P. Prel. Cat. N. Y. 36. 1888. 


Stem 1°-2° high, little branched or simple, terete 
or nearly so, the branches alternate. Basal leaves 
spatulate, obtuse, 13’-3’ long; stem leaves lanceolate 
or oblong-lanceolate, acute, the uppermost usually 
narrowly linear; flowers few, pink, sometimes white, 
solitary at the ends of the branches or peduncles, 
13’-23’ broad; calyx-lobes narrowly linear, about 
one-half as long as the 8-12 spatulate-obovate co- 
rolla-segments; anthers coiled; style deeply 2-cleft, 
its divisions clavate; capsule globose-oval, 3” high. 

In sandy borders of ponds and along salt marshes, 
Massachusetts to North Carolina, near the coast. July— 


Sept. Plants of the Gulf States, previously referred to 
this species, prove to be distinct. 


3. EUSTOMA Salisb. Parad. Lond. pl. 34. 1806. 


Erect usually branched glaucous annual herbs, with opposite sessile or clasping entire 
leaves. Flowers large, blue, purple or white, long-peduncled, axillary and terminal, solitary 
or paniculate, Calyx deeply 5-6-cleft, the lobes lanceolate, acuminate, keeled. Corolla broadly 
campanulate, deeply 5-6-lobed, the lobes oblong or obovate, usually erose-denticulate, convo- 
lute in the bud. Stamens 5-6, inserted on the throat of the corolla; filaments filiform; 
anthers oblong, versatile, at length recurved, or remaining nearly straight. Ovary 1-celled; 
style filiform; stigma 2-lamellate. Capsule oblong or ovoid, 2-valved. Seeds small, numer- 
ous, foveolate. [Greek, open-mouth, referring to the corolla.] 


Four species, natives of the southern United States, New Mexico and the West Indies. Type 
species: Eustoma silenifolium Salisb. 


8 GENTIANACEAE. Vor. III. 


1. Eustoma Russellianum (Hook.) Griseb. 
Russell’s Eustoma. Fig. 3346. 


Lisianthus Russellianus Hook. Bot. Mag. pl. 3626. 18309. 


Lisianthus glaucifolius Nutt. Trans. Am. Phil. Soc. (II.) 
5: 197. 1833-37. Not. Jacq. 1786. 


E. Russellianum Griseb. in DC. Prodr. 9: 51. 1845. 


Stem terete, rather stout, 1°-23° high. Leaves 
oblong or ovate-oblong, 3—-5-nerved, the upper usually 
acute at the apex, cordate-clasping at the base, 13-3’ 
long, the lower usually obtuse at the apex and nar- 
rowed to a sessile or slightly auricled base; pedun- 
cles 1-4’ long, stout, bracted at the base, the bracts 
lanceolate-subulate, Small; flowers 2’-3’ broad; 
calyx-lobes long-acuminate, shorter than the purple 
corolla; corolla-lobes obovate, about 4 times as long 
as the tube; style slender; anthers remaining nearly 
straight; capsule oblong, pointed. 


On prairies, Nebraska to Louisiana, Colorado, Texas, 
New Mexico and Mexico. Canada pest. May—Aug. 


4. GENTIANA [Tourn.] L. Sp. Pl. 227. 1753. 


Erect mostly glabrous herbs, with opposite or rarely verticillate, entire sessile or short- 
petioled leaves. Flowers blue, purple, yellow or white, solitary or clustered, terminal or 
axillary. Calyx tubular, 4-7- (usually 5-) cleft. Corolla tubular, clavate, campanulate, sal- 
verform or funnelform (rotate in some exotic species), 4-7-lobed, often gland-bearing within, 
the lobes entire or fimbriate. Stamens as many as the lobes of the corolla and inserted in its 
tube, included; anthers connate into a tube, or separate, not recurved or coiled. Ovary 
1-celled; ovules very numerous; style short or none; stigma cleft into 2 lamellae. Capsule 
sessile or stipitate, 2-valved. Seeds numerous, sometimes covering the whole inner wall of 
the capsule, wingless or winged. [Named for King Gentius of Illyria.] ; 

About 150 species, mostly natives of the north temperate and arctic zones and the Andes of 
South America. Besides the following, some 20 others occur in the western parts of North America, 
Type species: Gentiana lutea L 


Corolla-lobes fringed or serrate ; flowers 17-3’ long. 


Corolla enclosed in the swollen wing-angled calyx. 1. G. ventricosa. 
Corolla conspicuously longer than the wingless calyx. a 
Corolla-lobes fringed all around their summits; leaves lanceolate. 2, G.crinita. 
Corolla-lobes fringed mainly on the margins ; leaves linear. 3. G. procera. 
Corolla-lobes with entire or rarely denticulate margins ; flowers 6”—12” long. : 
Corolla-lobes fimbriate-crested at the base, acute. 4. G. acuta, 


Corolla-lobes naked, subulate-acuminate. ‘ 
Calyx-lobes unequal ; flowers mostly 4-parted. 5. G. propingua. | 
Calyx-lobes equal ; flowers 5-parted. 6. G. quinquefolia, 


1. Gentiana ventricédsa Griseb. Swollen Gen- 
tians * Fig.2347: 


Gentiana ventricosa Griseb. in Hook. Fl. Bor. Am. 2: 65. 
pl. 152. 1834. 


Annual; stem strict, terete, branched above, about 1° 
high. Basal leaves obovate, small, those of the stem 
ovate-oblong, obtuse or acute at the apex, rounded or , 
subcordate at the base, 1’ long, 4”-6” wide; flowers few, 
solitary at the ends of stout peduncles, 4-parted; calyx 
inflated, ovoid, about 9” high, 4”-5” thick, wing-angled, 
enclosing the corolla; corolla-lobes oblong, obtuse, lac- 
erate-serrate; ovary elliptic-oblong. 


~ 


Grand Rapids of Saskatchewan, between Cumberland 
ee and Hudson Bay (Drummond). Not recently col- 
ected, 


GENUS 4. GENTIAN FAMILY. 9 


2. Gentiana crinita Froel. Fringed Gen- 
tian. Fig. 3348. 


Gentiana crinita Froel. Gen. 112. 1796, 


: Annual or biennial; stem leafy, usually branch- 
ed, 1°-34° high, the branches erect, somewhat 
4-angled. Basal and lower leaves obovate, ob- 


; tuse, the upper lanceolate or oblong-lanceolate, 
} 1-2’ long, acute or acuminate at the apex, sessile 
: by a rounded or subcordate base; flowers mostly 
- 4-parted, several or numerous, about 2’ high, 


solitary at the ends of the usually elongated 
’ peduncles; calyx-lobes lanceolate, acuminate, un- 
equal, their midribs decurrent on the angles of 
; the tube; corolla bright blue, rarely white, nar- 
rowly campanulate, its lobes obovate, rounded, 
conspicuously fringed all around their summits, 
scarcely fringed on the sides, spreading when 
mature; capsule spindle-shaped, stipitate; seeds 
scaly-hispid. 


In moist woods and meadows, Quebec to Ontario, 
Minnesota, south to Georgia and Iowa. Sept.—Oct. 


\g 
3. Gentiana procera Holm. Smaller 
Fringed Gentian. Fig. 3349. 
Gentiana procera Holm, Ottawa Nat. 15: 11. tIgor. 


Annual; similar to the preceding species but 
smaller; stem simple, or little branched, 3’—18' 
high. Basal and lower leaves spatulate, obtuse, 
the upper linear or linear-lanceolate, 1’-23’ long, 
2”-4” wide; flowers 1-6, solitary at the ends of 
elongated erect peduncles, mostly 4-parted, about 
1%’ high; calyx-lobes lanceolate, acuminate, their 
midribs decurrent on the tube; corolla narrowly 
campanulate, bright blue, its lobes spatulate- 
oblong, strongly fringed on both sides, entire or 
somewhat fimbriate or toothed around the apex; 
capsule short-stipitate; seeds scaly-hispid. 

In wet places, New York and Ontario to Minne- 
sota, South Dakota and Manitoba. July—Sept. Pre- 
viously confused with G. detonsa Rottb. and with G. 
serrata Gunner, Old World species. Northwestern 
and Rocky Mountain plants formerly referred to 
this species prove to be distinct from it. 

Gentiana nesdphila Holm, of Anticosti, differs in 
merely denticulate corolla-lobes, 


3 
5 


4. Gentiana acuta Michx. Northern Gentian. 
Fig. 3350. 


Gentiana acuta Michx. Fl. Bor, Am. 1: 177. 1803. 
Gentiana Amarella var. acuta Herder, Act. Hort. Petrop. 
I: 428. 1872. 


Annual; stem leafy, slightly wing-angled, simple or 
branched, 6-20’ high. Basal and lower leaves spatu- 
late or obovate, obtuse, the upper lanceolate, acuminate 
or acute at the apex, rounded or subcordate at the 
base, sessile, or somewhat clasping, 4’—2’ long; flowers 
numerous, racemose-spicate, 5-8’ high, the pedicels 
2-6” long, leafy-bracted at the base; calyx deeply 
5-parted (rarely 4-parted), its lobes lanceolate; corolla 
tubular-campanulate, 5-lobed (rarely 4-lobed), blue, its 
lobes lanceolate, acute, each with a fimbriate crown at 
the base; capsule sessile. 


In moist or wet places, Labrador to Alaska, Maine, 
Minnesota, south in the Rocky Mountains to Arizona and 
Mexico. Also in Europe and Asia. Closely#resembles the 
Old World G. Amarella, Felwort. Bastard-gentian, Bald- 
money. Summer. 


10 GENTIANACEAE. Vo. III. 


5. Gentiana propinqua Richards. Four-parted 
Gentian. Fig. 3351.- 
Gentiana propinqua Richards. Frank. Journ. 734. 1823. 


Stems slender, usually branched from the base and 
sometimes also above, slightly wing-angled, 2-7’ high. 
Basal leaves spatulate, obtuse, the upper distant, oblong 
or lanceolate, 3-12’ long, acute or obtusish at the apex, 
rounded at the slightly clasping base, obscurely 3-nerved ; 
flowers few or several, solitary on slender peduncles, 
mostly 4-parted, 8-10” high; calyx 4-cleft, 2 of the 
lobes oblong, the others linear-lanceolate; corolla blue 
or bluish, glandular within at the base, its lobes ovate- 
lanceolate, entire or denticulate; capsule linear, at length 
a little longer than the corolla. 


Labrador to Alaska and British Columbia. Summer. 


2 
3 


6. Gentiana quinquefolia L. Stiff Gentian. 
Ague-weed. Fig. 3352. 


Gentiana quinquefolia L. Sp. Pl. 230. 1753. 
Gentiana quinqueflora Lam. Encycl. 2: 643. 1786. 


Annual; stem ridged, usually branched, 2’-23° 
high, quadrangular. Basal leaves spatulate, obtuse, 
those of the stem ovate, acute at the apex, clasping 
and subcordate or rounded at the base, 3-7-nerved, 
3-2’ long; flowers in clusters of 1-7 at the ends 
of the branches, or also axillary; pedicels slender, 
2’-7” long; calyx one-fourth to one-third the length 
of the corolla, its lobes narrow, or sometimes folia- 
ceous, acute; corolla tubular-funnelform, 5/10” 
long, blue or yellowish, glandular within at the base, 
its lobes triangular, very acute, entire; capsule stipi- 
tate; seeds globose, wingless. 


In dry or moist scil, Maine and Ontario to Michigan, 
south to Florida and Missouri. Ascends to 6300 ft. in North Carolina. Consists of several or 


many races, differing in size of plant, size of corolla; the calyx-lobes narrow or broad. Five- 
flowered gentian. Gall-of-the-Earth. Gall-weed. Aug.—Oct. 


5. DASYSTEPHANA [Reneal.] Adans. Fam. Pl. 2: 502. 1763. 


Mostly perennial herbs with opposite entire leaves, and large sessile or short-stalked 
flowers in terminal and axillary clusters, or sometimes solitary, usually 2-bracted under the 
calyx. Calyx mostly 5-lobed, with an interior membrane. Corolla mostly 5-lobed, clavate 
to funnelform, with thin lobed or toothed plaits in the sinuses, but without glands. Stamens 
as many as the corolla-lobes, the anthers cohering in a ring or separate. Ovary 1-celled; 
ovules very numerous; style short; capsule stipitate. [Greek, rough garland.] 

About 75 species, natives of the north temperate zone. Besides the following, several others 
occur in western North America. Type species: Gentiana asclepiadea L. 

Margins of leaves and calyx-lobes scabrous or ciliate; seeds winged. 
Anthers separate, or merely connivent. 
Stems usually clustered ; calyx-lobes unequal; corolla narrowly funnelform. 1. D. affinis. 
Stems mostly solitary ; calyx-lobes equal; corolla campanulate-funnelform. 2. D. puberula, 
Anthers cohering in a ring or short tube. 
Corolla-lobes distinct, longer than or equalling the plaits. 
Flowers 1-4; corolla campanulate-funnelform, its lobes 2-3 times as long as the plaits. 
3. D. parvifolia. 
Flowers several or numerous; corolla club-shaped, its lobes not much longer than the 
plaits. 4. D. Saponaria. 
Corolla-lobes none or minute, the plaits very broad. 5. D. Andrewsit. 
Margins of leaves and calyx-lobes smooth or nearly so. 
Flowers clustered, sessile, 2-bracteolate under the calyx. 
Corolla-lobes ovate, twice as long as the plaits; leaves broad, acuminate; seeds winged. 
6. D. flavida, 
Corolla-lobes rounded, little longer than the plaits; leaves narrow; seeds winged. 
7. D. linearis. 
Corolla-lobes ovate, acute, much longer than the broad plaits; leaves broad ; seeds vied 
. Grayi. 
Corolla-lobes triangular-lanceolate ; leaves obovate ; seeds wingless. 9. D. villosa. | 
Flowers solitary, peduncled, not bracteolate ; leaves linear. 10. D. Porphyrio. 


GENUS 5. GENTIAN FAMILY. irae 


1. Dasystephana affinis (Griseb.) Rydb. Ob- 
long-leaved Gentian. Fig. 3353. 


Gentiana affinis Griseb. in Hook. Fl. Bor. Am. 2: 56, 1834. 
D. affinis Rydb, Bull, Torr. Club 33: 149. 1906, 


Perennial; stems clustered from deep roots, minutely 
puberulent, simple, 618’ high. Leaves linear-oblong 
to lanceolate-oblong, obtuse or acutish, rounded or 
narrowed at the base, firm, roughish-margined, indis- 
tinctly nerved, 3-12’ long, the floral smaller; flowers 
few, numerous, or rarely solitary, 5-parted, sessile and 
solitary or clustered in the axils of the upper leaves, 
about 1’ high, not bracted under the calyx; calyx-lobes 
linear or subulate, unequal, the longer about equalling 
the tube, the smaller sometimes minute; corolla nar- 
rowly funnelform, blue, its lobes ovate, acute or mu- 
cronate, entire, spreading, with laciniate appendages in 
the sinuses; anthers separate; seeds broadly winged. 


In moist soil, Minnesota to British Columbia, south in 
the Rocky Mountains to New Mexico. Aug.—Oct. 


2. Dasystephana pubérula (Michx.) Small. 
Downy Gentian. Fig. 3354. 


Gentiana tuberula Michx. Fl. Bor. Am, 1: 176. 1803. 
D. puberula Small, Fl, SE. U. S. 930. 1903. 

Perennial; stems usually solitary, leafy, 8-18’ high, 
minutely puberulent or glabrous, nearly terete. Leaves 
firm, lanceolate, or the lower oblong, roughish- 
margined, indistinctly nerved, pale beneath, nar- 
rowed or rounded at the base, 1-3’ long; flowers 
sessile or nearly so in the upper axils, rarely soli- 
tary and terminal, 2-bracteolate under the calyx, 
13’-2’ high; calyx-lobes linear-lanceolate, equal, about 
as long as the tube, rough; corolla campanulate- 
funnelform, 2-3 times as long as the calyx, its lobes 
ovate, entire, spreading, much longer than the cleft 
or laciniate appendages; anthers at first connivent, 
soon separate; seeds oblong, broadly winged. 

On prairies, Maryland to Ohio, Minnesota, South 
Dakota, Georgia and Kansas, Aug.—Oct. 


3. Dasystephana parvifolia (Chapm.) 
Small. Elliott’s Gentian. Fig. 3355. 

?Gentiana rigida Raf. Med. Fl. 1: 212. 1832. 
Gentiana Elliottii Chapm, Fl. S. States, 356. 1860. 

Not Raf, 
?Gentiana scaberrima Kusnezow, Act. Hort. Petrop. 

13: 59. 1893. 
D. parvifolia Small, Fl. SE. U. S. 930. 1903. 
D. latifolia Small, Fl. SE. U. S. 930. 1903. 

Perennial; roots acluster of thick fibres; stems 
slender, leafy, terete, minutely rough-puberulent, 
simple, 8’-2° high. Leaves ovate or lanceolate, 
acute or acutish at the apex, rounded or narrowed 
at the base, thin, roughish-margined, 1-2’ long, 
3-nerved, or the lower much smaller and obtuse; 
flowers 1-4, terminal, or rarely I or 2 in the 
upper axils, about 2’ long, sessile, 2-bracteolate 
under the glabrous calyx; calyx-lobes oblong or 
lanceolate, foliaceous, longer than the tube; co- 
rolla campanulate-funnelform, blue, its lobes 
ovate, obtuse, sometimes mucronate, entire, 3-4” 
long, about twice as long as the fimbriate or 
toothed appendages; anthers cohering in a tube; 
seeds oblong, broadly winged. 

In moist soil, Virginia to Florida. Sampson snake-root. Sept.—Oct. 

Gentiana decora Pollard, of the same range, differs in having the calyx-tube pubescent. 


12 GENTIANACEAE. Vou. III. 


4. Dasystephana Saponaria (L.) Small, 
Soapwort or Blue-Gentian. Fig. 3356. 
Gentiana Saponaria L. Sp. Pl. 228 1753. 
Gentiana Catesbaei Walt. Fl. Car. 109. 1788. 
D. Saponaria Small, Fl. SE. U. S. 930. 1903. 
Perennial; stem erect or ascending, terete, 
slender, simple or with short erect axillary 
branches, glabrous, or minutely rough-puber- 
ulent above, 1°-23° high, leafy. Leaves lan- 
ceolate, ovate-lanceolate or oblong, usually 
acute at both ends, 3-5-nerved, roughish-mar- 
gined, 2’-4’ long, the lowest obovate and 
smaller; flowers 1-2’ high, in sessile terminal 
and usually also axillary clusters of I-5, 
2-bracteolate under the calyx; calyx-lobes 
oblong or spatulate, ciliolate; corolla blue, 
club-shaped, its lobes erect, obtuse, equalling 


or longer than the cleft or lacerate append- 
yh Zé ages; anthers cohering in a tube; capsule stip- 

2 itate; seeds broadly winged. 
— In wet soil, Ontario to Minnesota, Connecticut, 


Florida and Louisiana. Calathian violet. Harvest- 
bells. Rough or marsh-gentian. Sampson snake- 
Z root. Aug,—Oct. 


5. Dasystephana Andréwsii (Griseb.) Small. 
Closed Blue or Blind Gentian. Fig. 3357. 

?Gentiana alba Muhl. Cat, Ed. 2, 29. 1818. 

?Gentiana clausa Raf. Med. Fl. 1: 210. 1832. 

G. Andrewsii Griseb. in Hook. Fl. Bor. Am. 2:55. 1834. 

D. Andrewsii Small, Fl, SE. U. S. 930. i903. 


° ° 


Perennial; stout, glabrous, 1°-2° high, simple, 
leafy. Leaves ovate to lanceolate, 3-7-nerved, acu- 
minate at the apex, narrowed or sometimes rounded 
at the base, 2-4’ long, rough-margined, the lowest 
oblong or obovate, smaller; flowers 1-13’ high 
in a terminal sessile cluster and commonly I or 2 
in the upper axils, 2-bracteolate under the calyx; 
calyx-lobes lanceolate or ovate, ciliolate, usually 
spreading; corolla oblong, club-shaped, blue, or 
occasionally white, nearly or quite closed, its lobes 
obsolete, the intervening appendages very broad, 
light colored, opposite the stamens; anthers coher- | 
ing in a tube; capsule stipitate; seeds oblong, 
winged. 

In moist soil, Quebec to Manitoba, Georgia and Ne- 
braska. Cloistered-heart. Bottle- or barrel-gentian. 
Aug.—Oct. 


6. Dasystephana flavida (A. Gray) Brit- 
ton. Yellowish Gentian. Fig. 3358. 


G. flavida A. Gray, Am, Journ. Sci. (II.) 1:80. 1846. 
G. alba A, Gray, Man. 360. 1848. Not Muhl, 1818. 


Perennial; stem stout, terete, glabrous, sim- 
ple, erect or ascending, 1°-3° high. Leaves ovate- 
lanceolate or oblong-lanceolate, smooth-mar- 
gined, acuminate at the apex, subcordate, clasp- 
ing or rounded at the base, 2-5’ long, 1-2’ wide; 
flowers several or numerous in a sessile terminal 
cluster and sometimes 1 or 2 in the upper axils, 
13’-2’ high, 2-bracteolate under the calyx; 
calyx-lobes ovate, acute, smooth or minutely 
rough-margined, shorter than the tube, at length 
spreading; corolla campanulate-funnelform, 
open, greenish or yellowish white, its lobes 
ovate, acute or obtuse, twice as long as the 
erose-denticulate appendages; anthers cohering 
in a tube; capsule stipitate; seeds winged. 


Tn moist soil, Ontario to Minnesota, Missouri, 
Virginia and Kentucky. Aug.—Oct. 


. 
| 
| 


ee ee eee 


ee Ss ON a. Oe Se 


GENUS 5. GENTIAN FAMILY. 13 


7. Dasystephana linearis (Froel.) Britton. 
Narrow-leaved Gentian. Fig. 3359. 


Gentiana linearis Froel, Gent. 37. 1796. 

Gentiana Saponaria var. linearis Griseb. in Hook, FI. 
Bor, Am. 2: 55. 1834. 

Gentiana rubricaulis Schwein. in Keating’s Narr. 
Long’s Exp. 2: 384. 1824. 

Gentiana linearis var, lanceolata A. Gray, Syn. Fl. 2: 
Inantet 23, Lo7o. 


Perennial; glabrous throughout; stem slender, 
terete, simple, 62° high, leafy. Leaves linear 
or linear-lanceolate, smooth-margined, acute at 
both ends, 3-nerved, 13-3’ long, 2-5’ wide; 
flowers 1-1’ high in a terminal cluster of I-5 
and sometimes also in the upper axils; calyx- 
lobes linear or lanceolate, shorter than the tube; 
corolla erect, blue, oblong-funnelform, open, its 
lobes erect, rounded, 1’-2” long, slightly longer 
than the entire or I-2-toothed appendages; an- 
thers coherent in a tube, or at length distinct; 
capsule stipitate; seeds winged. 

In bogs and on mountains, New Brunswick and 
Ontario to Maryland and Minnesota. Ascends to 
5000 ft. in the Adirondacks. Aug.—Sept. 


8. Dasystephana Grayi (Kusnezow) Brit- 
ton. Gray’s Gentian. Fig. 3360. 


Gentiana linearis var. latifolia A, Gray, Proc. Am. 
Acad. 22: 309. 1887. 

Gentiana Grayi Kusnezow, Act. Hort. Petrop. 13: 59. 
1893. 


Perennial; glabrous; stem terete, 1°-2° high. 
Leaves rather distant, lanceolate or ovate-lanceo- 
late, acuminate, rounded at the base, smooth- 
margined, 2-3’ long, the lower narrower and 
obtuse; flowers about 12’ high in a sessile term- 
inal cluster, 2-bracteolate under the calyx; calyx- 
lobes unequal, the larger about as long as the 
tube; corolla greenish blue or bright blue, club- 
shaped, its lobes ovate, acute or acutish, much 
longer than the broad 1I-2-toothed appendages; 
anthers coherent, or at length distinct; seeds 
oblong, winged. 


In wet soil, New Brunswick to western Ontario, 
Michigan and Minnesota. Referred in our first edi- 
tion to Gentiana rubricaulis Schwein., which proves 
to be a synonym of the preceding species, Recorded 
from central New York. Aug.—Sept. 


g. Dasystephana vill6sa (L.) Small. Striped 
Gentian. Fig. 3361. 


Gentiana villosa L. sp. Pl. 228. 1753. 
Gentiana ochroleuca Froel. Gent. 35. 1796. 
D. villosa Small, Fl, SE. U. S. 931. 1903. 


Perennial, glabrous or nearly so; stem simple, slen- 
der, terete, 6-18’ high. Leaves obovate, obtuse or the 
upper acute, narrowed at the base, faintly 5-nerved, 
1-3’ long, the lower much smaller; flowers several in 
a terminal sessile cluster and sometimes also in the 
upper axils, nearly 2’ long, 2-bracteolate under the 
calyx; calyx-lobes unequal, linear, longer than the tube; 
corolla greenish white, striped within, oblong-funnel- 
form, open, its lobes triangular-ovate or ovate-lanceo- 
late, erect, much longer than the oblique entire or 
I-2-toothed appendages; seeds oval, wingless. 

In shaded places, southern New Jersey and Pennsylvania 


to Florida and Louisiana. Marsh- or straw-colored gentian. 
Sampson snake-root) Sept.—Nov. 


14 GENTIANACEAE. Vot. III. 


10. Dasystephana Porphyrio (J. F.Gmel.) Small. 
One-flowered Gentian. Fig. 3362. 


Gentiana purpurea Walt. Fl. Car. 109. 1788. Not. L. 1753. 
Gentiana Porphyrio J. F. Gmel. Syst. 2: 462. 1791. 
Gentiana angustifolia Michx. Fl. Bor, Am, 1: 177. 1803. 
D. Porphyrio Small, Fl. SE. U. S. 931. 1903. 


Perennial, glabrous; stem erect. or ascending, simple or 
branched, 6-18’ high. Leaves linear, firm, acute or blunt 
at the apex, 1-2’ long, 1”-2” wide, or the upper and 
lower shorter; flowers solitary at the ends of the stem 
or branches, short-peduncled, about 2’ high, not bracteo- 
late under the calyx; calyx-lobes linear, longer than the 
tube; corolla funnelform, bright blue, sometimes brown- 
dotted within, its lobes ovate, acutish, spreading, 5-7” 
long, three times as long as the conspicuously laciniate 
appendages, or more; seeds oblong, wingless. 


In moist pine barrens, New Jersey to Florida. Aug.—Oct. 


6. PLEUROGYNA Eschol. Linnaea 1: 187. 1826. 


Slender usually branched annual glabrous herbs, with opposite leaves, and rather large 
blue or white flowers in terminal narrow racemes or panicles, or solitary at the ends of the 
slender peduncles. Calyx deeply 4-5-parted; segments narrow, often unequal. Corolla rotate, 
4-5-parted; lobes vate or lanceolate, convolute, acute, with a pair of narrow appendages at 
the base. Stamens 4 or 5, inserted on the corolla-tube near its base; filaments slender or 
filiform; anthers ovate, sagittate, straight. Ovary 1-celled; ovules numerous; style none; 
stigma decurrent along the sutures of the ovary. Capsule 2-valved. Seeds small and numer- 
ous. [Greek, referring to the lateral stigmatic surfaces.] 

About 7 species, of the colder parts of the northern hemisphere, only the following typical one 
in North America, ‘ 


1. Pleurogyna rotata (L.) Griseb. Marsh Felwort. 
Fig. 3363. 


wertia rotata L. Sp. Pl. 226. 1753. 
Pleurogyne rotata Griseb, Gent. 309. 1839. 


Stem erect, usually 6-15’ high, sometimes lower, simple, 
or with nearly erect branches. Leaves linear to lanceolate, 
3-2’ long, 1-2” wide, or the basal spatulate or oblong, 
shorter and sometimes broader; sepals narrowly linear to 
lanceolate, about the length of the lanceolate to oblong- 
lanceolate corolla-segments which are 4”-7” long; capsule 
narrowly oblong. 


Quebec, Greenland and Labrador to Alaska, south in the 
Rocky Mountains to Colorado. Recorded from the White 
Mountains of New Hampshire. Also in Europe and Asia. 
Summer. 


7. FRASERA Walt. Fl. Car. 87. 1788. 


Perennial or biennial erect glabrous mostly tall herbs, with opposite or verticillate leaves, 
and rather large white yellowish or bluish flowers, in terminal cymose panicles or thyrses. 
Calyx 4-parted, the segments narrow. Corolla rotate, 4-parted, the lobes convolute in the 
bud, each bearing 1 or 2 fimbriate or fringed glands within. Stamens 4, inserted on the base 
of the short corolla-tube; filaments subulate or filiform, sometimes united at the base; anthers 
oblong, versatile. Ovary ovoid, I-celled; style slender or short, but distinct; stigma 2-lobed 
or nearly entire. Capsule ovoid, coriaceous, somewhat compressed, 2-valved, few-seeded. 
Seeds flattened, smooth, margined or narrowly winged. [In honor of John Fraser, a botan- 
ical collector. ] 

About 15 species, natives of North America, all but the following typical one far western. 


GENUS 7. GENTIAN FAMILY. 15 


1. Frasera carolinénsis Walt. American 
Columbo. Fig. 3364. 


Frasera carolinensis Walt. Fl. Car. 87. 1788. 


Stem 3°-7° high, stout. Leaves mostly verti- 
cillate in 4’s, those of the stem lanceolate or 
oblong, acute or acutish, 3-6’ long, the basal ones 
spatulate or oblanceolate, usually much larger, 
the uppermost small and bract-like; inflorescence 
thyrsoid-paniculate, large, often 2° long; flowers 
slender-pedicelled, about 1’ broad; corolla yel- 
lowish white with brown-purple dots, its lobes 
broadly oblong, bearing a large circular long- 
fringed gland at or below the middle; style 2”-3” 
long; stigma 2-lobed; capsule much compressed, 
6’-8” high, longer than the calyx. 


In dry soil, western New York and Ontario to 
Wisconsin, south to Georgia and Tennessee. Yellow 
gentian. Pyramid-flower or -plant. June—Aug. 


8. HALENIA Borck. in Roem. Archiv 1: 25. 1796. 


[TETRAGONANTHUS S. G. Gmel. Fl. Sib. 4: 114. Hyponym. 1769.] 

Annual or perennial usually tufted glabrous herbs, with opposite leaves, and middle-sized 
white yellowish purple or blue flowers in terminal and axillary often panicled cymes. Calyx 
deeply 4-cleft or 4-parted, the segments lanceolate or oblong. Corolla campanulate, 4-5-cleft, 
the lobes convolute in the bud, each with a hollow spur or projection below, which is glandu- 
lar, at the bottom within, or sometimes spurless. Stamens 4 or 5, inserted near the base of 
the corolla; filaments filiform or subulate; anthers oblong, versatile. Ovary 1-celled, the 
placentae more or less intruded; ovules numerous; style very short, sometimes none; stigma 
2-lobed. Capsule ovoid or oblong, 2-vaived. Seeds globose-ovoid to oblong, compressed, 
smooth. [In honor of Jonas Halen, 1727-1810, a pupil of Linnaeus.] 


_About 30 species, natives of mountainous regions of North America, South America and Asia. 
Besides the following, another occurs in the southwestern United States. Type species: Halenia 
sibirica Borck. 


1. Halenia defléxa (J. E. Smith) Griseb. Spurred Gentian. Fig. 3365. 
Swertia deflera J. E. Smith in Rees’ Cyclop. no. 8. 1816, 
Halenia deflexa Griseb. in Hook. Fl. Bor. Am, 2: 67. pi. 
I55. 1834. 
H. Brentoniana Griseb. in Hook. Fl. Bor. Am. 2: 68, 1834. 
H, heterantha Griseb. loc. cit. 1834. 
Tetragonanthus deflexus Kuntze, Rev. Gen. Pl. 431. 1891, 
Annual or biennial; stem simple or branched, slen- 
der, erect, 6-20’ high, usually with long internodes. 
Basal leaves obovate or spatulate, obtuse, narrowed 
into petioles; stem-leaves ovate or lanceolate, acute, 
sessile, 3-5-nerved, 1’-2’ long, the uppermost much 
smaller; calyx-segments lanceolate or spatulate, acute 
or acuminate; corolla purplish or white, about 4” 
high, its lobes ovate, acute, the spurs deflexed or 
descending, one-fourth to one-half the length of the 
corolla or none; capsule narrowly oblong, 6’-7” 
long, about twice as long as the calyx. 


In moist woods and thickets, Newfoundland and Lab- 
rador to Massachusetts, New York, Saskatchewan, Mon- 
tana, Michigan and South Dakota. Races differ in size 
of the plant and of the flowers and in the development 
of the corolla-spurs. Recorded from the “ Indian Ter- 
ritory ” (Oklahoma), apparently erroneously. July—Aug. 


9. OBOLARIA L. Sp. Pl. 632. 1753. 


A low glabrous perennial herb, the stem simple or branched, the lower leaves reduced 
to opposite scales, the upper foliaceous, subtending the racemose-spicate or thyrsoid white 
or purplish flowers. Calyx of 2 spatulate sepals. Corolla oblong-campanulate, 4-cleft, the 
lobes imbricated, at least in the bud. Stamens 4, inserted in the sinuses of the corolla; fila- 


16 GENTIANACEAE. Vo. III. 


ments slightly longer than the ovate sagittate anthers. Ovary 1-celled, with 4 internal 
placental projections; ovules numerous; style distinct; stigma 2-lamellate. Capsule ovoid 
2-valved or irregularly bursting. Seeds minute, covering the whole interior of the capsule. 
[Greek, obolus, a coin, alluding to the thick round leaves.] 


A monotypic genus of eastern North America. 


1. Obolaria virginica L. Pennywort. Fig. 3366. 
Obolaria virginica L. Sp. Pl. 632. 1753. 


Stems 3’-6’ high from a perennial base with thick fibrous 
roots, bearing 2-6 pairs of thick small obtuse scales in 
place of leaves. Floral leaves broadly obovate-cuneate, 
obtuse, purplish, 4-6” long; flowers sessile or nearly so 
in the axils, in clusters of 1-4 (usually 3), and terminal; 
corolla about 5” long, cleft to about the middle, the lobes 
oblong, obtuse, entire, or denticulate; stamens included; 
capsule 22” long. 


In rich woods and thickets, New Jersey to Georgia, west to 
Illinois and Texas. Ascends to 2600 ft.in Virginia, April-May. 


10. BARTONIA Muhl.; Willd. Neue Schrift. Ges. Nat. Fr. Berlin 3: 444. 1801. 


Slender or filiform erect glabrous annual or biennial herbs, the leaves reduced to minute 
opposite subulate scales, or some of them alternate, and white purplish or yellowish racemose 
or paniculate flowers, or these rarely solitary and terminal. Calyx deeply 4-parted, the seg- 
ments lanceolate, acuminate, keeled. Corolla campanulate, deeply 4-cleft, the lobes imbricated 
in the bud. Stamens 4, inserted at the sinuses of corolla; filaments slender, longer than the 
ovate sagittate anthers. Ovary I-celled, the placentae intruded; ovules numerous; style very 
short or none; stigma 2-lobed. Capsule ovoid-oblong, compressed, acute, 2-valved. Seeds 
minute, covering the whole inner surface of the capsule. [In honor of Professor Benjamin 
Smith Barton, 1766-1815, of Philadelphia.] 


Four species, natives of eastern North America. Type species: Bartonia tenella Willd. 


Corolla yellowish, 11%4”—2” long. 


Corolla-lobes oblong, abruptly tipped, erose. 1, B. virginica. 
Corolla-lobes lanceolate, acute or acuminate, entire. 2. B. paniculata. 
Corolla white, 3’—4” long, its lobes spatulate. 3. B. verna. 


1. Bartonia virginica (L.) B.S.P. Yellow Bartonia. 
Fig. 3367. 

Sagina virginica L. Sp. Pl. 128. 1753. 

B. tenella Willd. Neue Schrift. Ges. Nat. Fr. Berlin 3: 445. 1801. 

Bartonia virginica B.S.P. Prel. Cat. N. Y. 36. 1888. 

Stem rather stiff, almost filiform, 4’-15’ high, simple, or 
with few erect branches above, 5-angled, yellowish green, 
sometimes twisted. Subulate scales 1-2” long, appressed, 
mostly opposite, the basal pairs close together, the upper dis- 
tant; flowers mostly opposite; pedicels ascending or erect, 
2”-6” long; corolla greenish yellow or whitish, 13”-2” long, 
its lobes oblong, obtuse, denticulate or erose, somewhat ex- 
ceeding the calyx; stamens included; ovary 4-sided; stigma 


1" 1" 


about 2” long; capsule about 12” long. 


In moist soil, Nova Scotia to Florida, Michigan, Minnesota and 
Louisiana. Screw-stem. July—Sept. 


eae 


GENUS IO. GENTIAN FAMILY. 17 


2. Bartonia paniculata (Michx.) Robin- 
son. Branched Bartonia. Fig. 3368. 
Centaurella paniculata Michx. Fl. Bor, Am. 1: 98. 


1803. 

EG. Moser Steud. & Hochst. ; Griseb. Gent. 308. 1839 
B. paniculata Robinson, Rhodora 10: 35. 1908 
B, lanceolata Small, Fl. SE. U. S. 932. 1903. 

Slender, 8-16’ high, mostly paniculately branch- 
ed above, the branches and scales mainly alter- 
nate, the slender pedicels spreading or ascending. 
Corolla yellowish-white or greenish, about twice 
as long as the calyx, its lanceolate acute or acumi- 
nate lobes 1’—12” long; anthers yellow. 

In wet soil, Massachusetts to Florida, Arkansas and 
Louisiana. Aug.—Oct. 

Bartonia iodandra Robinson, of bogs in Newfound- 
land and Nova Scotia, has larger purplish flowers 
with broader ovate-lanceolate corolla-lobes and pur- 
ple-brown anthers. 


3. Bartonia vérna (Michx.) Muhl. White Bartonia. 
Fig. 3360. 


Centaurella verna Michx. Fl. Bor. Am, 1: 97. pl. 12. f. 1. 1803. 
Bartonia verna Muhl. Cat. 16. 1813. 


Stem thicker and less rigid than that of the two preceding 
species, usually simple, 2-15’ high, racemosely or corymbosely 
I-several-flowered. Bracts distant, appressed, some of them 
commonly alternate; flowers solitary at the ends of the elon- 
gated erect peduncles, 3’-4” long; peduncles sometimes 2’ 
long; corolla white, its lobes spatulate or obovate, obtuse, 
denticulate or entire, about 3 os ice length of the calyx; 
ovary compressed ; capsule about 23” high. 


In moist sand, southern Virginia to Florida and Lowisiana. 
March-—May. 


Family 16. MENYANTHACEAE G. Don, Gard. Dict. 4: 167. 1837. 


BucKBEAN FAMILY. 


Perennial aquatic or marsh herbs, with basal or alternate entire crenate or 
3-foliolate leaves, and clustered regular perfect flowers. Calyx inferior, deeply 
5-parted, persistent. Corolla funnelform to rotate, 5-lobed or 5-cleft, the lobes 
induplicate-valvate, at least in the bud. Stamens 5, borne on the corolla, and 
alternate with its lobes; filaments mostly short; anther-sacs longitudinally dehis- 
cent; pollen-grains 3-angled. Ovary 1-celled, the 2 placentae sometimes intruded ; 
style long, short or none. Fruit a capsule, or indehiscent. 


Five genera and about 35 species, widely distributed. 


Leaves 3-foliolate; swamp plant. _ 1. Menyanthes. 
Leaves simple, entire, cordate, floating. 2. Nymphoides. 


1. MENYANTHES [Tourn.] L. Sp. Pl. 145. 1753. 


A perennial glabrous marsh herb, with creeping rootstocks, alternate long-petioled 
3-foliolate basal leaves, and white or purplish flowers, racemose or panicled on long lateral 
scapes or peduncles. Calyx 5-parted, the segments oblong or lanceolate. Corolla short- 
funnelform, 5-cleft, its lobes induplicate-valvate, fimbriate or bearded within, spreading. 
Stamens 5, inserted on the tube of the corolla; filaments filiform; anthers sagittate. Disk 
of 5 hypogynous glands. Ovary 1-celled; style subulate, persistent; stigma 2-lamellate. Cap- 
sule oval, indehiscent or finally rupturing. Seeds compressed-globose, shining. [Greek, per- 
haps month-flower, name used by Theophrastus. ] 

A monotypic genus of the cooler parts of the northern hemisphere. 

2 


18 ~MENYANTHACEAE. Vor. IIT. 


1. Menyanthes trifoliata L. Buckbean. 
Marsh or Bean Trefoil. Bog-bean 
or -myrtle. Fig. 3370. 

Menyanthes trifoliata L. Sp. Pl. 145. 1753. 


Rootstock thick, scaly, sometimes 1° long, mark- 
ed by the scars of bases of former petioles. Leaves 
3-foliolate; petioles sheathing at the base, 2-10’ 
long; leaflets oblong or obovate, entire, obtuse at 
the apex, narrowed to the sessile base, pinnately 
veined, 13’-3’ long; raceme borne on a long 
scape-like naked peduncle, arising from the root- 
stock, 10-20-flowered; pedicels stout, 3’—12” long, 
bracteolate at the base; flowers 5-6” long; calyx 
shorter than the white or purplish corolla, which 
is bearded with white hairs within; stamens 
shorter than the corolla and style exserted, or 
longer and style nearly included; capsule ovoid, 
obtuse, about 4” long. 

In bogs, Greenland to Alaska, south to Long Island, 
Pennsylvania, Minnesota, Nebraska and California. 
Also in Europe and Asia. Water- or bitter trefoil. 


Water-shamrock. Moon-flower. Marsh-clover. Bitter 
worm. Bog-nut. Brook-bean. May-July. 


2. NYMPHOIDES Hill, Brit. Herb. 77. 1756. 
[LIMNANTHEMUM S. G. Gmelin, Nov. Act. Acad. Petrop. 14: 527. pl. 17. f. 2. 1769.] 


Aquatic perennial herbs, with slender rootstocks. Leaves floating, petioled, ovate or 
orbicular, deeply cordate, entire or repand, or the primary ones different; flowers yellow or 
white, polygamous, umbellate at the summit of filiform stems at the bases of the petioles, or 
axillary, often accompanied by a cluster of thick elongated root-like tubers. Calyx 5-parted. 
Corolla nearly rotate, deeply 5-cleft, the lobes induplicate-valvate in the bud, sometimes 
fimbriate on the margins, glandular at the base. Stamens 5, inserted on the base of the 
corolla; filaments short; anthers sagittate, versatile. Ovary 1-celled; style short or none; 
stigma 2-lamellate. Capsule ovoid or oblong, indehiscent or irregularly bursting. Seeds 
numerous or few, smooth or rough. [Greek, like Nymphaea.] 


About 20 species, widely distributed in temperate and tropical regions. The following are the 
only ones known to occur in North America. Type species: Nymphoides flava Hill. 


Flowers white, accompanied by tufts of root-like tubers; native species. 


Floating leaves 1’—2’ long; flowers 3”—6” broad; seeds smooth, 1. N.lacunosum, 
Floating leaves 2’-6’ long; flowers 6”—10” broad; seeds rough. 2. N. aquaticum, 
Flowers not accompanied by tufts of tubers; corolla bright yellow, 1’ broad or more; introduced 

species, 3. N.nymphaeoides, 


1. Nymphoides lacundsum (Vent.) Kuntze. 
Floating Heart. Fig. 3371. 

Villarsia lacunosa Vent, Choix des Plantes, 9. 1803. 
Limnanthemum lacunosum Griseb. Gent. 347. 1839. : 
Nymphoides lacunosum Kuntze, Rev. Gen. Pl. 429. 1891. 

Rootstock buried in the mud, the roots long and 
fibrous. Primary leaves membranous, submerged, short- 
petioled; stems (stolons) filiform, greatly elongated, 
sometimes 10° long, bearing a short-petioled floating 
leaf, an umbel of flowers and a cluster of narrow tubers 
at its summit, or some of the floating leaves on petioles 
as long as the stems; floating leaves ovate-orbicular, 
purplish beneath, 1-2’ long, the basal sinus narrow or 
broad; pedicels slender; tubers linear-conic, 4/1 long; 
flowers nearly white, 3-6” broad; style none; capsule 
ovoid, covered by the connivent calyx-segments; seeds 
numerous, smooth. 


Z In ponds, Nova Scotia to Florida, west to Ontario, Min- 
(ip nesota and Louisiana. July—Aug. 


GENUS 2, 


2. Nymphoides aquaticum (Walt.) 
Kuntze. Larger Floating Heart. 


BUCKBEAN FAMILY. 


0 On 
8.2 


vos" 
eo oO, 


_Villarsia aquatica Gmel. Syst. 1: 447. 1791. 


_ Leaves alternate ; erect herbs. 


Fig. 3372. 


Anonymos aquatica Walt. Fl. Car. 109. 1788, 


vn 
on 
eo .0°R 


On 
rn 
3.9 
C 
‘0 
So. 
cc 
So! 
c 


Menyanthes trachysperma Michx, Fl. Bor. Am. 1: 

126. 1803. 

Limnanthemum trachyspermum A. Gray, Man. Ed. 

5, 390. 1867. 

Limnanthemum aquaticum Britton, Trans. N, Y. 

Acad. Sci. g: 12. 1889. 

Nymphoides aquaticum Kuntze, Rev. Gen. Pl. 

429. 1891. 

Similar to the preceding species but stouter 
and larger. Floating leaves cordate-orbicular, 
thick, entire or repand, 2’-6’ long, spongy, and 
with the petioles and stolons densely covered 
with minute pits; primary leaves spatulate; 
pedicels slender, 1-3’ long; tubers linear- 
oblong, thicker; corolla white, 6’—10” broad; 
style none; seeds rough; capsule longer than 
the calyx. 

In ponds, southern New Jersey and Delaware 
to Florida and Texas. May—Aug. 


O; 
want) 


3. Nymphoides nymphaeoides (L.) 
Britton. Water Lily. Floating 
Fleart.. ic. 3373: 


Menyanthes nymphaeoides L. Sp. Pl. 145. 1753. 


Limnanthemum nymphaeoides Hoffm. & Link, 
"OR Port, Deng44.0 C00: 


Stems long, stout, creeping or buried in 
the mud, ascending to the surface of the 
water, branched, the upper nodes bearing 
a petioled leaf and a cluster of flowers, or 
the upper internodes very short and their 
leaves apparently opposite. Petioles stout, 
mostly longer than the ovate-orbicular firm 
blades, which are 27-4’ broad; pedicels 
stout, becoming 3’-6’ long, not accompanied 
by tufts of tubers; flowers bright yellow, 
1’ in diameter, or more; corolla segments 
short-fringed; seeds with fimbriate margins. 


Naturalized in ponds, District of Columbia. 
Native of Europe and Asia. May-July. 


Family 17, APOCYNACEAE Lindl. Nat. Syst. Ed. 2, 299. 1836. 
DocBANE FAmIty. 

Perennial herbs, shrubs, vines, or some tropical genera trees, mostly with an 
acrid milky juice, with simple opposite alternate or verticillate exstipulate leaves, 
and perfect regular 5-parted cymose solitary or paniculate flowers. Calyx infe- 
rior, persistent, the lobes imbricated in the bud. Corolla gamopetalous, its lobes 
convolute in the bud and often twisted. Stamens as many as the lobes of the 
corolla, alternate with them, inserted on the tube or throat; anthers. linear-oblong, 
or sagittate, 2-celled; pollen-grains simple, often glutinous. Ovary superior, or 


‘its base adherent to the calyx, of 2 distinct carpels, or 1-celled, with 2 parietal 


placentae, or 2-celled; ovules few or numerous, anatropous or amphitropous; style 

simple, or 2-divided ; stigma simple. Fruit usually of 2 follicles or drupes. Seeds 

often appendaged by a coma; endosperm fleshy, not copious; embryo straight; 

cotyledons flat or concave; radicle terete, usually shorter than the cotyledons. 
About 130 genera and 1100 species, very widely distributed, mostly in tropical regions. 


1. Amsonia, 
Leaves opposite; vines or herbs. 
Flowers large, axillary, solitary. 2. Vinca, 


20 APOCYNACEAE. Vot. III. 


Flowers small, cymose. 
Erect or diffuse herbs; corolla campanulate. 3. Apocynum. 
High-climbing vines; corolla funnelform. 4. Trachelospermum. 


1. AMSONIA Walt. Fl. Car. 08. 1788. 


Perennial herbs, with alternate membranous leaves, and rather large blue or bluish flowers, 
in terminal thyrsoid or corymbose cymes. Calyx 5-parted, the segments narrow, acuminate. 
Corolla mostly salverform, the tube cylindric, but somewhat dilated at the summit, villous 
within. Stamens inserted on the throat of the corolla, included; anthers ovate or oblong. 
Disk none. Ovary of 2 carpels, connected at the top by the filiform style; ovules in 2 rows 
in each cavity, numerous; stigma appendaged by a reflexed membrane. Fruit of 2 cylindric 
several-seeded follicles. Seeds cylindric or oblong, obliquely truncate at each end, not 
appendaged. [Named for Charles Amson of South Carolina.] 


About 8 species, natives of North America and eastern Asia. Besides the following, 5 others 
occur in the southern and southwestern United States. Type species: Amsonia Tabernaemontana 
Walt. 


1. Amsonia Amsonia (L.) Britton. Am- 


sonia. Fig. 3374. 

Tabernaemontana Amsonia L. Sp. Pl. Ed, 2, 308. 

1762. 
Amsonia Tabernaemontana Walt. Fl. Car. 98. 1788. 
A. salicifolia Pursh, Fl. Am. Sept. 184. 1814. 
A, Amsonia Britton, Mem, Torr. Club 5: 262. 1894. 

Glabrous or nearly so, simple, or branched 
above, 2°-4° high. Leaves ovate, ovate-lanceolate 
or lanceolate, entire, acuminate at the apex, nar- 
rowed at the base, sometimes pubescent beneath, 
2’-4’ long, 4’-2’ wide; petioles 2’-4” long; flowers 
numerous; pedicels bracteolate at the base; calyx 
about 1” long, its segments subulate; corolla 
—9” long, beaked by the convolute limb in the 
bud, its lobes linear and about as long as the 
tube; follicles 2’-4’ long, about 2” thick, attenuate 
at the apex, glabrous, divergent or ascending; 
seeds papillose. 

In moist soil, New Jersey to Illinois, Kentucky, 
Missouri, Florida and Texas. Consists of several 
races, differing in leaf-form. April—July. 


2.. VINCAGE Sp. Pl. 2095" 17a 


Erect or trailing herbs, some species slightly woody, with opposite leaves, and large soli- 
tary blue pink or white axillary flowers. Calyx 5-parted, the segments narrow, acuminate. 
Corolla salverform, the tube cylindric, or expanded above, pubescent within, the lobes convo- 
lute, at least in the bud, oblique. Stamens included. Disk of 2 glands, alternate with the 
2 carpels. Ovules several in each carpel; style filiform; stigma annular, its apex penicillate. 
Follicles 2, erect or spreading, cylindric, several-seeded. Seeds oblong-cylindric, truncate at 
each end, not appendaged. [The Latin name.] 


About 12 species, natives of the Old World. Type 
species: Vinca major L. 


1. Vinca minor L. Periwinkle. Myrtle. 
Fig. 3375. 
Vinca minor L, Sp. Pl. 209. 1753. 


Perennial, trailing, glabrous; stems 62° long. 
Leaves oblong to ovate, entire, firm, green both 
sides, obtuse or acutish at the apex, narrowed at 
the base, short-petioled, 1-23’ long, 3’-1’ wide; 
flowers not numerous, solitary in some of the 
axils, blue, 9-15” broad; peduncles slender, 4’—13’ 
long; calyx very deeply parted, the segments subu- 
late-lanceolate, glabrous, about 14’ long; corolla- 
tube expanded above, as long as or slightly longer 
than the obovate, nearly truncate lobes; anther- 
sacs witha broad connective; follicles few-seeded. 


Escaped from gardens to roadsides and woods, On- 
tario to Connecticut, southern New York and Geor- 
gia. Native of Europe. Leaves shining. Also called 
running myrtle or small periwinkle. Feb.—May. 


GENUS 3. DOGBANE FAMILY. 21 


3. APOCYNUM L. Sp. Pl. 213. 1753. 


Perennial branching herbs, with opposite entire leaves, and small white or pink flowers 
in terminal and sometimes axillary corymbed cymes. Calyx 5-parted, the segments acute. 
Corolla urceolate to campanulate, the tube bearing within 5 small appendages alternate with 
the stamens, the limb 5-lobed. Stamens inserted on the base of the corolla; anthers sagit- 
tate, connivent around the stigma and slightly adherent to it. Disk 5-lobed. Ovary of 2 
carpels; ovules numerous in each carpel; stigma ovoid, obtuse, obscurely 2-lobed. Follicles 
slender, elongated, terete. Seeds numerous, small, tipped with a long coma. [Greek, dogbane.] 

About 11 species, of the north temperate zone, most abundant in North America. Type species: 
Apocynum androsaemifolium L. 

Corolla: campanulate, not angled, 4”-4'%4” long, pink, its lobes widely spreading or recurved. 
) 1. A. androsaemifolium. 
Corolla urceolate to short-tubular, or narrowly campanulate, angled, 144”-3%” long, greenish, 
white, or faintly pink, its lobes a little spreading, or erect. 
Corolla narrowly campanulate, 2'%4”—3'%4” long, lobesnearly equalthetube. 2, A. medium. 
Corolla urceolate to short-tubular, 1%”—3” long, lobes much shorter than the tube. 
Calyx-lobes ovate, much shorter than the corolla tube. 3. A. Milleri. 
Calyx-lobes lanceolate, about as long as the corolla-tube. 
Leaves and cymes glabrous, or sparingly pubescent. 
Leaves all narrowed at the base and distinctly petioled ; flowers greenish; plant rather 


bright green. 4. A. cannabinum, 
At least the lower leaves sessile, and mostly rounded or subcordate at the base; plant 
pale glaucous green; flowers white. 5. A. sibiricum. 
Léaves, cymes, and often the whole plant densely pubescent. 6. A. pubescens. 


1, Apocynum androsaemifolium L. 
Spreading Dogbane. Fig. 3376. 
Apocynum androsaemifolium L. Sp. Pl. 213. 1753. 

Syst. Ed. 10, 946. 1759. . 
A, divergens Greene, Leaflets 1: 56. 1904. 

Rootstock horizontal ; stem 1°-4° high; branches 
broadly spreading, mostly glabrous. Leaves ovate 
or oval, acute or obtuse and mucronate at the 
apex, rounded or narrowed at the base, glabrous 
above, pale and usually more or less pubescent 
beneath, 2’-4’ long, 1’-22’ wide; petioles 2’—4” 
long; cymes loose; pedicels 2’-3” long, subulate- 
bracted at the base; flowers about 4” broad; calyx- 
segments shorter than the tube of the pinkish 
corolla; corolla-lobes revolute; follicles about 4’ 
long, narrowed at the apex. 


In fields and thickets, Anticosti to British Colum- 
bia, south to Georgia, Missouri, Nebraska and Ari- 
zona. Ascends to 3500 ft.in Virginia. Honey-bloom. 
Bitter-root or -dogbane. Rheumatism-wood. Wild 
ipecac. Wandering milk-weed. Western wall-flower. 
Fly-trap. June-July. Linnaeus inadvertently failed 
to affix a binominal specific name in the first edition 
of his “Species Plantarum,” but corrected this six 
years later. 


2. Apocynum médium Greene. Intermediate Dogbane. 
Fig. 3377. 
Apocynum medium Greene, Pittonia 3: 29. 1897. 


Apocynum speciosum G. S. Miller, Proc. Biol. Soc. Wash. 13: 83. 
1899. 


Rootstock horizontal; stem stout, 4° high or less, the branches 
ascending, glabrous. Leaves oblong to oval or elliptic, mucronulate, 
ascending, 24-4’ long, somewhat pubescent beneath; petioles 
2”-4” long; cymes terminal, compact; pedicels 14-24” long; 
flowers erect; calyx-segments ovate, about half as long as the 
5-angled corolla-tube; corolla white or pinkish, 23’—34” long, 
urceolate-campanulate, its lobes acutish, somewhat spreading, 
nearly as long as the tube; follicles 3-4’ long. 


Fields and hillsides, Quebec to the District of Columbia, west to 
Iowa, June-Aug. 


22 APOCYNACEAE. Vor. 111. 


3. Apocynum Milleri Britton. Miller’s 
Dogbane. Fig. 3378. 


Apocynum Milleri Britton, Manual 739. Igot. 


Stem slender, 3° high or less, the branches 
spreading. Leaves oblong to ovate-lanceolate, 
23’-32' long, pubescent beneath, the pubescent 
petioles 13’—3” long; cymes small, terminal or 
also in the upper axils, the pedicels 1-13” 
long; flowers nearly erect; corolla pinkish, 
23-3” long, its rounded segments spreading, 
much shorter than the tube, which is longer 
than the ovate calyx-segments; follicles about 
4’ long. 


Dry soil, New York to Maryland and the Dis- 
trict of Columbia. June-July. 


4. Apocynum cannadbinum L. Indian Hemp. 
Amy-root. Fig. 3370. 


Apocynum cannabinum L, Sp. Pl. 213. 1753- 
A. cannabinum glaberrimum DC. Prodr. 8: 439. 1844. 
A. nemorale G. S. Miller, Proc. Biol. Soc. Wash. 13: 87. 


1899. 
Apocynum urceolifer G. S. Miller, loc. cit. 


Root deep, vertical, soon branching. Stem exten- 
sively branched, the branches erect or ascending, 
glabrous or nearly so, more or less glaucous. Leaves 
oblong, lanceolate-oblong or ovate-oblong, actite or 
obtuse and mucronate at the apex, narrowed or 
rounded at the base, glabrous above, sometimes pu- 
bescent beneath, 2’-6’ long, 3’-3’ wide; petioles 1-6” 
long, or sometimes none; cymes dense; pedicels 
short, bracteolate at the base; calyx-segments about 
as long as the tube of the greenish-white corolla; 
corolla-lobes nearly erect; follicles similar to those 
of the preceding species. 


In fields and thickets, Connecticut to Wisconsin, Ala- 
bama, Tennessee, Missouri and Kansas, perhaps extend- 
ing farther north. Rheumatism-root. Wild cotton. 


5. Apocynum sibiricum Jacq. Clasping- 
leaved Dogbane. Fig. 3380. 


A. sibiricum Jacq. Hort. Vind. 3: 37. pl. 66. 1776. 

A. hypericifolium Ait. Hort. Kew. 1: 304. 1789. © 

A. cannabinum var. hybericifolium A, Gray, Man, 
365. 1848. 

Apocynum album Greene, Pittonia 3: 230. 1897. 


Glabrous, pale green, often glaucous; stem 
1°-2° high, the branches ascending. Leaves ob- 
long, oblong-lanceolate to oval, 1’-3’ long, 3-12’ 
wide, obtuse or acutish at the apex, cordate- 
clasping, rounded, truncate, or most of the upper 
narrowed at the base, short-petioled, or sessile, 
the primary venation forming broad angles with 
the midvein; cymes many-flowered, dense to 
loose; pedicels mostly not longer than the flow- 
ers, bracteolate; calyx-segments about as long as 
the corolla-tube, lanceolate, acute; corolla-lobes 
nearly erect; follicles 2-34’ long. 


Mostly along streams, Quebec to British Coiumbia, 
Long Island, Ohio, Kansas and New Mexico. St. 
John’s-dogbane. June—-Aug,. 


GENUS 3. DOGBANE FAMILY. 23 


6. Apocynum pubéscens R. Br. Velvet Dog- 
bane. Fig.- 3381. 


A, pubescens R. Br. Mem, Wern. Soc. 1: 68. 1811. 
Apocynum cannabinum var. pubescens A. DC, Prodr. 8: 
440. 1844. 


Whole plant, including the pedicels and calyx, 
densely velvety-pubescent, or the stem sometimes 
glabrate. Branches ascending; leaves oval to elliptic, 
obtuse or acute at the apex, strongly mucronate, 
obtuse or obtusish at the base, the veins impressed 
in the pubescence of the lower surface; petioles 1’”—2” 
long; cymes dense; calyx-segments about as long as 
the tube of the corolla, lanceolate, acute; corolla- 
lobes erect; follicles about 4’ long. 


In dry sandy soil, Ontario to Rhode Island, Maryland, 


Alabama, Iowa and Kansas. April-Aug. Perhaps a pu- 
bescent race of A. cannabinum L, 


4. TRACHELOSPERMUM Lemaire, 
Wiaedeetieur 1: pl Or. 1851. 


Twining woody vines (some exotic species nearly erect shrubs), with opposite entire 
deciduous leaves, and small yellow greenish or white flowers in terminal and axillary com- 
pound cymes. Calyx small, deeply 5-parted, glandular within, the segments narrow. Corolla 
funnelform or salverform, the tube nearly cylindric, expanded above, the lobes convolute, 
more or less twisted. Stamens included, or short exserted; anthers sagittate, acuminate, 
connivent around the stigma and slightly adherent to it. Disk of 5 glandular lobes. Ovary 
of 2 carpels; ovules numerous in each carpel; style slender, its apex thickened below the 
narrow ring of the ovoid stigma. Follicles much elongated, slender. Seeds linear or oblong, 
not beaked, long-comose at the apex. [Greek, neck-seed, but the seed is not beaked.] 


About 6 species, natives of eastern Asia and North 
America. Type species: Trachelospermum jasminoides 
Lemaire, of eastern Asia. The following is the only 
known North American species, 


1. Trachelospermum différme (Walt.) A. 
Gray. Trachelospermum. Fig. 3382. 


Echites difformis Walt. Fl. Car. 98. 1788. 
Forsteronia difformis A. DC. Prodr. 8: 437. 1844. 
T. difforme A, Gray, Syn. Fl. 2: Part 1, 85. 1878. 

A high-climbing vine, the stems 3’ in diameter or 
more, the twigs pubescent or glabrous. Leaves thin, 
ovate, oval or lanceolate, acuminate or acute at the 
apex, narrowed or rarely rounded at the base, 14’-3’ 


long, 4’-2’ wide; petioles 2’-4” long; peduncles slen- 
der, shorter than the leaves; pedicels and branches 
of the cyme bracteolate at the base; flowers yellow 
or cream-color, 4’—-5” long; lobes of the corolla 
ovate, spreading, shorter than the tube; follicles very 
slender, 5-9’ long, scarcely 2” thick; many-seeded. 


In moist woods and along streams, Delaware to Flor- 
ida, Texas and Mexico, mostly near the coast, north to 
Arkansas and Missouri. June-Aug. 


Family 18. ASCLEPIADACEAE Lindl. Nat. Syst. Ed. 2, 302. 1836.* 


MILKWEED FAMILY. 

Perennial herbs, vines or shrubs, mostly with milky juice, with opposite alter- 
nate or verticillate exstipulate leaves, and mostly umbellate perfect regular flowers. 
Calyx inferior, its tube very short, or none, its segments imbricated or separate 
in the bud. Corolla campanulate,.urceolate, rotate or funnelform, 5-lobed or 
5-cleft, the segments commonly reflexed, mostly valvate in the bud. A 5-lobed 
or 5-parted crown (corona) between the corolla and the stamens and adnate to 
one or the other. Stamens 5, inserted on the corolla, usually near its base; fila- 
ments short, stout, mostly monadelphous, or distinct; anthers attached by their 


* Text revised for our first edition by Miss ANNA Murray Var, here somewhat modified. 


24 ASCLEPIADACEAE. Vou; Ty 


bases to the filaments, introrsely 2-celled, connivent around the stigma, or more 
or less united with each other ; anther-sacs tipped with an inflexed or erect scarious 
membrane, or unappendaged at the top, sometimes appendaged at the base; pollen 
coherent into waxy or granular masses, one or rarely two such masses in each sac, 
connected with the stigma in pairs or fours, by 5 glandular corpuscles alternate with 
the anthers. Disk none. Ovary of 2 carpels; styles 2, short, connected at the 
summit by the peltate discoid stigma; ovules numerous in each carpel, mostly 
anatropous, pendulous. Fruit of 2 several-many-seeded follicles. Seeds com- 
pressed, usually appendaged by a long coma; endosperm cartilaginous, mostly 
thin; embryo nearly as long as the seed; cotyledons flat. 


About 220 genera and 2000 species, of very wide geographic distribution, most abundant in 
tropical or warm-temperate regions. 


Erect or decumbent herbs. 


Corona-hoods each with an incurved horn within; eaves mostly opposite. 1. Asclepias. 
Corona-hoods prominently crested within; leaves alternate. 2. Asclepiodora. 
Corona-hoods unappendaged or with a thickened crest-like keel. 3. Acerates. 
Twining vines, 
Corolla-lobes erect ; corona-lobes 1—2-awned., 4. Gonolobus. 
Corolla rotate. 
Anthers tipped with a scarious membrane; pollen-masses pendulous, 5. Cynanchum. 
Anthers merely tipped ; pollen-masses horizontal. 6. Vincetoxicum. 


a 


1. ASCLEPIAS L. Sp. Pl. 214. 1753. 


Perennial erect or decumbent herbs, with opposite verticillate or rarely alternate entire 
leaves, and middle-sized or small flowers in terminal or axillary umbels. Calyx 5-parted or 
5-divided, usually small, the segments or sepals acute, often glandular within. Corolla deeply 
5-parted, the segments mostly valvate, reflexed in anthesis. Corona-column generally present. 
Corona of 5 concave erect or spreading hoods, each bearing within a slender or subulate 
incurved horn, either included or exserted. Filaments connate into a tube; anthers tipped 
with an inflexed membrane, winged, the wings broadened below the middle; pollen-masses 
solitary in each sac, pendulous on their caudicles. Stigma nearly flat, 5-angled or 5-lobed. 
Follicles acuminate. Seeeds comose in all but one species. [Dedicated to Aesculapius. ] 

About 95 species, mostly natives of the New World; besides the following, some 25 others occur 
in southern and western North America. Known as Milkweed, Silkweed, or Swallow-wort. Type 
species: Asclepias syriaca L. 

* Corolla and corona orange; leaves alternate or opposite. 
Stem erect or ascending ; leaves nearly all alternate. 1. A. tuberosa. 
Stems reclining; leaves, at least the upper, opposite, oblong or oval. 2. A. decumbens. 
** Corolla bright red or purple (sometimes greenish in A. rubra) leaves opposite. 
Flowers 4”—-6” broad; corona-hoods 2”—3” high. 
Leaves lanceolate or linear; hoods oblong, obtuse. 
Leaves ovate or ovate-lanceolate ; hoods lanceolate. ~ 
Leaves oblong, ovate or ovate-oblong; hoods oblong, acutish. 
Flowers 2”—3” broad; corona-hoods 1”-1'4” high, 
Plant nearly or quite glabrous ; leaves lanceolate or oblong-lanceolate. . A. incarnata. 
Plant pubescent ; leaves oblong. . A. pulchra. 
*** Corolla greenish, purplish, yellowish or white; leaves opposite or verticillate. 
Leaves ovate, oblong, ovate-lanceolate, obovate or orbicular. 
Plants glabrous throughout, or minutely pubescent above. 
Leaves sessile, clasping or very short-petioled. 
Peduncles of the solitary or several umbels short. 
Leaves ovate-oblong; hoods 2-auriculate at the base. 8. A. Sullivantii. 
Leaves nearly orbicular ; hoods truncate. g. A. latifolia, 
Peduncle of the usually solitary umbel elongated. 
Leaves wavy-margined. 


. A. lanceolata, 
. A. rubra. 
. A. purpurascens. 


NO UA 


Leaves cordate-clasping. 10. A. amplexicaulis. 
Leaves rounded at the base, short-petioled. 11. A. intermedia, 
Leaves sessile, flat; horn not exceeding the hood. 12, A. Meadi. 
Leaves manifestly petioled. 
Corolla greenish; umbels loose, the pedicels drooping. 13. A. exaltata. 
Corolla white; umbels dense. 14. A. variegata. 


Corolla pink ; some of the leaves verticillate in 4’s. 15. A. quadrifolia, 
Plants. at least the lower surfaces of the leaves, canescent or tomentose. 
Follicles tomentose, covered with soft spinose processes. ; 
Corona-hoods obtuse, short. 16. A. syriaca. 
Corona-hoods elongated, lanceolate. 17. A. speciosa. 
Follicles with no spinose processes, glabrous or pubescent. 
Leaves wavy-margined ; corolla-segments 4”—5” long. 
Leaves flat; corolla-segments 2”—3” long. 
Leaves lanceolate, oblong-lanceolate or linear. 
Leaves opposite, lanceolate or oblong-lanceolate. 
Leaves thick, short-petioled'; inflorescence woolly. 20, A. brachystephana. 
Leaves thin, slender-petioled ; inflorescence downy. 21. A. perennis. 


18. A. arenaria. 
19. A. ovalifolia. 


eee: MILKWEED FAMILY. . 25 


Leaves mostly verticillate in 3’s—6’s, narrowly linear. 
Hoods entire. 
Hoods dorsally hastate-sagittate. 

Leaves scattered, densely crowded, filiform-linear. 


1. Asclepias tuberosa L. Butterfly-weed 
or -flower. Pleurisy-root. Fig. 3383. 


Asclepias tuberosa L. Sp. Pl. 217. 1753. 


Hirsute-pubescent; stems stout, simple, or 
branched near the summit, ascending or erect, 
very leafy, 1°-2° high, the milky sap scanty. 
Leaves alternate, lanceolate or oblong, acute or 
sometimes obtuse at the apex, narrowed, rounded 
or cordate at the base, sessile or short-petioled, 
2’-6’ long, 2’—12” wide; umbels cymose, terminal, 
many-flowered; peduncles shorter than the leaves ; 
pedicels 2’-1’ long; corolla-segments about 3” 
long, greenish orange; corona-column about 4” 
long; hoods erect, oblong, bright orange, or yel- 
low, 2-3 times as long as the stamens, longer than 
the filiform horns; fruiting pedicels decurved; 
follicles nearly erect, finely pubescent, 4’-5’ long. 

In dry fields, Maine and Ontario to Minnesota, 
Florida, Texas, Chihuahua and Arizona. Consists of 
numerous races, differing in shape and size of the 
leaves and color of the flowers. June-Sept. Wind- 
or orange-root. Canada-, flux-, tuber- or white-root. 
Orange swallow-wort. Yellow milkweed. Indian-posy. 


3. Asclepias lanceolata Walt. Few-flowered 
Milkweed. Fig. 3385. 

Asclepias lanceolata Walt. Fl. Car. 105. 1788. 
A. paupercula Michx. Fl. Bor. Am, 1: 118. 1803. 

Glabrous; stem slender, usually simple, naked 
above, 2°-4° high. Leaves opposite, distant, linear 
or narrowly lanceolate, elongated, acuminate, nar- 
rowed at the base, short-petioled, 4’-10’ long, 20a" 
wide, roughish on the margins, the primary nerves 
widely spreading; umbels few-flowered, solitary or 
2-4 at the summit; peduncles about equalling the 
slender puberulent pedicels; corolla-segments oblong, 
4”-5” long, deep red; column thick, about 1” high; 
hoods obovate or oblong, obtuse, orange, 2-toothed 
near the base, nearly twice the length of the anthers 
and longer than the subulate incurved horn; fruit- 
ing pedicels decurved; follicles erect, minutely 
puberulent, fusiform, about 4’ long. 

In swamps, southern New Jersey to Florida and 
Texas, mostly near the coast. June-Aug. 


flowered, 


22, A. verticillata. 
23. A. galioides. 
24. A. pumila, 


) 


(\ Nf Ye / Pp 
WM ie Ppy 
\ We 


" 


wiZe 
WE 


2. Asclepias decimbens L. Decumbent 
Butterfly-weed. Fig. 3384. 
Asclepias decumbens L. Sp. Pl. 216. 1753. 


Hirsute-pubescent; stems decumbent, 2°-3° 
long, the ends ascending or erect. Leaves sessile 
or short-petioled, oblong or elliptic, obtuse at the 
apex, narrowed and often inequilateral at the 
base, 1’-3’ long, 4’-23’ wide, the upper opposite, 
the lower commonly alternate, the uppermost 
very small; umbels several or numerous, many- 
racemose along the branches, one 
usually in each of the upper axils; peduncles 
stout, short; pedicels slender, pubescent, about 
¥’ long; corolla-segments oblong, acutish, dark 
orange, about 3” long; column about 4” high, 
the hoods erect, oblong, orange, longer than the 
subulate horn; follicles slender. 

In dry fields, Illinois and Ohio to North Carolina 
and Florida. 


Creeping milkweed. June—Aug. 


ASCLEPIADACEAE. Vor, IEE 


4. Asclepias rubra L. Red Milkweed. 
Fig. 3386. 


Asclepias rubra L. Sp. Pl. 217. 1753. 


Nearly glabrous throughout; stem usually sim- 
ple, 1°-4° high. Leaves opposite, rather distant, 
short-petioled, ovate, lanceolate or the lower some- 
times oblong, rounded or subcordate at the base, 
gradually acuminate, rather firm, 3-8’ long, 1-23’ 
wide, the primary nerves wide-spreading; umbels 
1-4, many-flowered; peduncles shorter than or 
equalling the upper leaves; pedicels slender, downy, 
3’-1’ long; corolla-segments and hoods lanceolate- 
oblong, purplish red, or the hoods orange-red, 
3-4" long, or flowers sometimes greenish; horns 
of the hoods very slender, nearly straight; fruiting 
pedicels deflexed, the follicles erect, spindle-shaped, 
glabrous, about 4’ long. 


In moist soil, New Jersey and Pennsylvania to Flor- 
jy ida, Missouri, Louisiana and Texas. June-July. 
5. Asclepias purpurascens L. Purple Milkweed. 


Asclepias purpurascens L. Sp. Pl. 214. 1753. 
?Asclepias amoena L. Sp. Pl. 214. 1753. 


Stem stout, puberulent or glabrous, usually sim- 
ple, 2°-4° high, leafy to the top. Leaves ovate, 
elliptic or oblong, short-petioled, acute or obtuse 
and mucronulate at the apex, narrowed or round- 
ed at the base, nearly glabrous above, finely 
tomentose: beneath, 3-8’ long, 12-3’ wide, the 
primary nerves very wide-spreading; umbels 
many-flowered, borne in several of the upper 
axils, or sometimes solitary; peduncles stout; 
pedicels slender, puberulent, 1-14’ long; corolla 
deep purple, its segments oblong to oblong-lan- 
ceolate, about 3” long; column very short and 
thick; hoods oblong or ovate, nearly twice as 
long as the anthers, pale red or purple, the horns 
broad at the insertion, short-subulate and incurved 
at the apex; fruiting pedicels deflexed, the 
downy follicles nearly erect, 4-5’ long. 


In dry fields and thickets, New Hampshire to 
North Carolina, west to southern Ontario, Minne- 
sota and Arkansas. Ascends to 2000 ft. in the Cats- 
kills. June—-Aug. 


sWittinn, 6. Asclepias incarnata L. Swamp Milk- 
SKM OL ; 
SRY (PASA weed. Fig. 3388. 
4 —C™WwWH EES 
eK Asclepias incarnata L. Sp. Pl. 215. 1753. 


Stem slender, glabrous, or puberulent in 2 lines 
above, branched or rarely simple, 2°-4° high, 
leafy to the top. Leaves lanceolate or oblong- 
lanceolate, acuminate at the apex, narrowed, ob- 
tuse or sometimes subcordate at the base, 3-6’ 
long, 3’-13’ wide, the primary nerves not wide- 
spreading; petioles 3-6” long; umbels usually 
numerous, corymbed, many-flowered; pedicels 
pubescent, 5-10” long; corolla red or rose- 
purple, rarely white, its lobes oblong, about 2” 
long; column more than one-half as long as the 
obtuse pink or purplish hoods; horns slender, in- 
curved, longer than the hoods; anther-wings en- 
tire, or obscurely notched at the base; fruiting 
pedicels erect or incurved; follicles erect, 2’-33’ 
long, sparingly puberulent. 


In swamps, New Brunswick to Ontario, Saskatch- 
ewan, Tennessee, Louisiana and Colorado. Ascends 
to 3000 ft. in West Virginia. Rose- or swamp-silk- 
weed. Waternerve-root. WhiteIndian-hemp. July—Sept. 


GENUS I. MILKWEED FAMILY. | 


7- Asclepias pilchra Ehrh. Hairy Milk- 
weed. Fig. 3380. 


Asclepias pulchra Ehrh.; Willd. Sp. Pl, 1: 1267. 1798. 
A, incarnata var. pulchra Pers. Syn, 1: 276. 1805. 


Similar to the preceding species and perhaps hy- 
bridizing with it where the two grow together; 
stem stout, tomentose-pubescent, usually branched, 
2°-33° high, leafy to the top. Leaves broadly lan- 
ceolate, acute, acuminate or some of them obtusish 
at the apex, subcordate, rounded, or the upper nar- 
rowed at the base, puberulent or glabrous above, 
pubescent, at least on the veins beneath, 3-5’ long, 
3/-2’ wide; petioles usually stout and short; flowers 
similar to those of A. incarnata, but the corolla 
commonly lighter red or pink, rarely white; pedun- 
cles and pedicels tomentose; fruiting pedicels erect 
of incurved; follicles erect, densely pubescent, 2’—3’ 
ong. 


In moist fields and swamps, Nova Scotia to Minne- 
sota, south to Georgia. WhiteIndian-hemp. July—Sept. 


8. Asclepias Sullivantii Engelm. Sulli- 
vant’s Milkweed. Fig. 3390. 


A, Sullivantii Engelm.; A. Gray, Man, 366. 1848. 


Glabrous throughout; stem stout, simple or 
sometimes branched above, 2°-4° high, leafy to 
the top. Leaves thick, sessile, or on petioles less 
than 1” long, oblong or ovate-oblong, usually 
obtuse and mucronulate at the apex, subcordate, 
rounded or slightly clasping at the base, 4-6’ 
long, 13-3’ wide, the primary nerves very wide- 
spreading; umbels terminal and sometimes also 
in the upper axils, many-flowered; peduncles 
shorter than the leaves; corolla-segments oval- 
oblong, 5-6” long, purplish; column very short 
and thick; hoods oval, obtuse or truncate, gib- 
bous at each side near the base, longer than the 
anthers and the subulate incurved horn; follicles 
erect, glabrous, 3-4’ long, usually with blunt 
processes near the apex. 


In moist soil, southern Ontario to Ohio, Minne- 
sota, Nebraska, Missouri and Kansas. July—Sept. 


g. Asclepias latifolia (Torr.) Raf. Broad-leaved Milkweed. Fig. 3301. 


Asclepias obtusifolia var. latifolia Torr. Ann. Lyc. N. Y. 
2: 117. 1826. 


Asclepias latifolia Raf, Atl. Journ, 146. 1832-33. 
A, Jamesii Torr. Bot. Mex, Bound, Surv. 162. 1859. 


Minutely puberulent wnen young, glabrous when 
old; stem stout, usually simple, 1°-22° high, very 
leafy. - Leaves very thick, oval to orbicular, sessile 
or nearly so, commonly broadly emarginate and 
mucronulate at the apex and cordate or subcordate 
at the base, 4-6’ long and nearly as wide, primary 
nerves very wide-spreading; umbels 2-4, many- 
flowered, short-peduncled in the upper axils or rarely 
terminal; pedicels slender, canescent, nearly 1’ long; 
corolla-segments ovate, acute, 4’-6” long, greenish; 
column short and thick; hoods truncate, about equal- 
ling the anthers, the horn projecting from a short 
crest over the edge of the stigma; follicles erect on 
eae pedicels, ovoid, acutish, 2-3’ long, about 1’ 
thick. 


On dry plains, Nebraska to Colorado, Texas and Ari- 
zona. July—Sept. 


28 ASCLEPIADACEAE. Vor. 111. 


10. Asclepias amplexicaulis J. E. Smith, Blunt-leaved Milkweed. Fig. 3392. 


A, amplexicaulis J. E. Smith, Georgia Insects 1: 35 
DL Fea 1707: 


A. obtusifolia Michx. Fl. Bor. Am. 1: 115. 1803. 


Nearly glabrous, pale green, somewhat glau- 
cous; stem stout, erect or ascending, 2°-3° high. 
Leaves sessile, oblong or ovate-oblong, obtuse 
and mucronulate at the apex, cordate-clasping at 
the base, 3’-5’ long, 1-13’ wide, margins wavy- 
crisped; umbels many-flowered, usually solitary 
on the long terminal peduncle, rarely with a 
second shorter-peduncled one at its base; pedi- 
cels slender, downy, about 1’ long; corolla-seg- 
ments oblong, greenish-purple, about 4” long; 
column thick; hoods pink, nearly truncate and 
toothed at the summit, shorter than the subulate 
incurved horn, longer than the anthers; follicles 
erect on the stout decurved fruiting pedicels, 
downy, 4-6’ long. 


In dry fields, mostly in sandy soil, New Hamp- 
shire to Florida, northern New York, Minnesota, 
Nebraska and Texas. Ascends to 3000 ft. in Vir- 
ginia. May-—Aug. 

A. Bicknéllii Vail, Bull. Torr. Club 31: 458, is ap- 
parently a hybrid between A. amplexicaulis J. E. 
Smith and A. eraltata (L.) Muhl. 


11. Asclepias intermedia Vail. Intermediate 
Milkweed. Fig. 3393. 


A, intermedia Vail, Bull, Torr. Club 31: 459. 1904. 


Stem erect, glabrous, purplish, not glaucous, about 
1° high. Leaves oblong-elliptic, glabrous above, mi- 
nutely pubescent beneath, 6’ long or less, obtuse at 
both ends or the upper subcordate at the base, the 
petioles very short; umbels 2 or more, terminal, pe- 
duncled, the peduncles and slender pedicels pubes- 
cent; corolla green-purple, its oblong-lanceolate seg- 
ments 3-4” long, the erect hoods ovate-lanceolate, 
obtuse, about 3” Jong, pink-purple, with a darker 
stripe on the back, the margins with an erect tooth 
above the middle, the horn slender. 

Lawrence, Long Island. Possibly a hybrid between 
A, syriaca and A, amplexicaulis, 

A 


12. Asclepias Méadii Torr. Mead’s Milkweed. 
Fig. 3394. 
A, Meadii Torr.; A. Gray, Man. Ed. 2, Add. 704. 1856. 


Nearly glabrous throughout, pale green or glaucous; 
stem simple, or rarely branched above, 1°-2° high. 
Leaves opposite, sessile, flat, mostly distant, ovate, 
ovate-lanceolate, lanceolate or the lower oblong, acute 
or sometimes obtuse at the apex, the margins scabrous ; 
umbel solitary, terminal, several-flowered, borne on a 
peduncle 3-6’ long; corolla-segments greenish, ovate, 
acute, 3”’-4” long; column very short, thicker than 
high; hoods ovate, purplish, nearly twice as long as 
the anthers, rounded and truncate at the summit, 
longer than the subulate inflexed horn, with a small 
tooth at each side on the inner infolded margin; folli- 
cles erect on decurved pedicels, minutely puberulent, 
narrow, 4’-5’ long. 


In dry soil, Illinois to Iowa and Wisconsin. June—Aug. 


GENUS I. MILKWEED FAMILY. 29 


13. Asclepias exaltata (L.) Muhl. Poke or 
Tall Milkweed. Fig. 3395. 


A, Syriaca var. exaltata L, Sp. Pl. Ed. 2, 313. 1762. 
Asclepias exaltata Muhl. Cat. 28. 1813. 
A, phytolaccoides Pursh, Fl, Am. Sept. 180. 1814, 


Nearly glabrous throughout, with two opposite 
lines of pubescence on the usually simple stem, 3°-6° 
high. Leaves opposite, thin or membranous, oval, 
ovate or oblong, acuminate at both ends, 4-0’ long, 
13-4’ wide, the lower sometimes obovate, obtuse, 
shorter; petioles #/-1’ long; peduncles 1-3’ long; 
umbels usually several; pedicels slender, drooping 
or spreading, 1’-2’ long, puberulent; corolla green- 
purple, the segments ovate or oblong, obtusish, 
3-4” long; column short; hoods white or pink, 
slightly shorter than the anthers, much shorter than 
the subulate horn, at the summit truncate and entire 
or erose, with I or 2 slender teeth on each of the 
inner margins; follicles erect on the deflexed pedi- 
cels, downy, long-acuminate, 4’-6’ long. 

In thickets and woods, Maine to Minnesota, Georgia, 
Missouri and Arkansas. <Ascends to 5500 ft, in North 
Carolina. June-Aug. 


14. Asclepias variegata L. White Milk-weed. Fig. 3396. 
Nea) MOY ¢ Asclepias variegata L. Sp. Pl. 217. 1753. 


Stem glabrous below, pubescent above when 
SY/CE \"\AEZs young, simple, 1°-3° high. Leaves opposite, thick, 
a: SS a oval, ovate, oblong or the lower somewhat obo- 
x vate, obtuse and cuspidate or acutish at the apex, 
narrowed or rounded at the base, dark green 
above, pale beneath, 3-6’ long, 17-3’ wide, the 
middle ones sometimes verticillate in 4’s; petioles 
3-12” long; umbels 1-4, terminal, or rarely I or 
2 in the upper axils, densely many-flowered; pe- 
duncles 1-2’ long; pedicels 4’-13’ long, erect or 
ascending, usually densely puberulent; corolla- 
segments ovate or oval, about 3” long, white, or 
purple near the base; column very short and 
thick, purplish; hoods globose-obovoid, obtuse, 
spreading, longer than the anthers, about equal- 
ling the semi-lunate horizontally pointed horn; 
follicles downy, erect on the deflexed fruiting 
pedicels, 4-5’ long. 
In dry woods or thickets, Connecticut (?), south- 
ern New York to Illinois, Arkansas, Florida and 
Texas, June-July. 


15. Asclepias quadrifolia Jacq. Four- 
leaved Milkweed. Fig. 3397. 
A, quadrifolia Jacq. Obs. Part 2. 8. pl. 33. 1767. 


Stem slender, simple, 1°-2° high, usually leaf-} 
less below. Leaves thin, sparingly pubescent on 
the veins beneath, ovate to lanceolate, 2’-6’ long,} 
‘2 wide, acute or acuminate, narrowed or 
rounded at the base, or the lowest pair much 
smaller, obovate and obtuse, the upper and lower 
opposite, the middle ones usually verticillate in 
4’s; umbels 1-4, terminal, or rarely in the upper 
axils; peduncles slender, 3’-23’ long; pedicels 
about 1’ long; corolla pink or nearly white, its 
lobes lanceolate-oblong, 2”-3’’ long; column short; 
hoods white, obtuse at the apex, broadly 2-toothed 
above the base, twice as long as the anthers and 
the short incurved horn; follicles erect on the 
erect fruiting pedicels, 3’-5’ long, glabrous. 


Woods and thickets, Maine and Ontario to Minne- 
sota, Alabama and Arkansas. May-July. 


3° ASCLEPIADACEAE. Vor. IIT. 


16. Asclepias syriaca L. Common Milk- 
weed. Silkweed. Fig. 3398. 


Asclepias syriaca L. Sp. Pl. 214. 1753. 
Asclepias Cornuti Dec. in DC. Prodr. 8: 564. 1844. 

Stem stout, usually simple, 3°-5° high, finely 
pubescent at least above. Leaves oblong, oval or 
ovate, finely and densely pubescent beneath, soon 
glabrous above, acute or obtuse and cuspidate at 
the apex, obtuse, narrowed or subcordate at 
the base, 4’-9’ long, 2’-43’ wide, the primary nerves 
wide-spreading; petioles stout, 3-8” long; umbels 
several or numerous; peduncles pubescent or tomen- 
tose, 13-33’ long; pedicels 1-2’ long; corolla green- 
purple or greenish-white, its segments oblong-lan- 
ceolate, 3-4” long; column short and thick, the 
hoods ovate-lanceolate with a tooth on each side, 
longer than the anthers and the incurved horn; fol- 
licles 3-5’ long, erect on recurved pedicels, tomen- . 
tose and covered with short soft processes. 

In fields and waste’ places, New Brunswick to Sas- 
katchewan, North Carolina and Kansas. Leaves rarely 
lanceolate. Silky swallow-wort. Virginia silk, Wild 
cotton. June—Aug. 

Asclepias kansana Vail, of Kansas, differs by erect-spreading hoods of the corolla and more 
densely tomentose follicles. 


17. Asclepias specidsa Torr. Showy Milk- 
weed. Fig. 3399. 


Asclepias speciosa Torr. Ann, Lyc. N. Y. 2: 218. 1826. 
A, Douglasit Hook. Fl. Bor. Am. 2: 53. pl. 152. 1834. 

White-tomentose or canescent all over, or gla- 
brate below, pale; stem simple, stout, 1°-23° high. 
Leaves thick, broadly ovate or oval, obtuse and cus- 
pidate or acute at the apex, subcordate, rounded or 
narrowed at the base, petioled, 3’-8’ long, 2’-4’ wide; 
peduncles 1~—3’ long; umbels several or rarely soli- 
tary, many-flowered; pedicels stout, 9-18” long; 
corolla purple-green, its segments oblong or ovate- 
oblong, 4”-6” long, tomentose on the outer face; 
column very short or none; hoods lanceolate, 5-7” 
long, obtusish, expanded and with 2 blunt teeth be- 
low, the apex ligulate, 5-7 times as long as the 
anthers; horn short, inflexed; follicles erect or 
spreading on the recurved fruiting pedicels, 3-4’ 
long, densely woolly and covered with soft spinose 
processes. 

In moist soil, Minnesota to British Columbia, south 
to Kansas, Utah and California. May-July. 


Ys 

18. Asclepias arenaria Torr. Sand Milk- 
weed. Fig. 3400. 

A, arenaria Torr. Bot. Mex. Bound. Surv. 162. 18509. 


Densely tomentose-canescent all over, stems 
simple, ascending or erect, stout, 1°-2° high. 
Leaves obovate or oval, wavy-margined, thick, 
obtuse or retuse and cuspidate at the apex, trun- 
cate, subcordate, obtuse, or rarely some of them 
narrowed at the base, 2-4’ long, 13-3’ wide, the 
angle of the primary nervation broad; umbels 
densely many-flowered, short-peduncled or ses- 
sile; corolla greenish-white, its segments oval- 
oblong, 4”-5” long; column 1’-2” high; hoods 
oblong, truncate at each end, oblique at the apex, 
longer than the anthers, with a broad tooth on 
each side within; horn semi-lunate with an ab- 
ruptly incurved subulate apex; follicles puberu- 
lent, 4-5’ long, erect on the decurved fruiting 
pedicels. ; 

On sand-bars and hills along rivers, Nebraska and 
Colorado to Mexico and New Mexico. June—Sept. 


GENUS I. MILKWEED FAMILY. 31 


19. Asclepias ovalifolia Dec. Oval-leaved 
or Dwarf Milkweed. Fig. 3401. 


_Asclepias ovalifolia Dec. in DC, Prodr. 8: 567. 1844. 


Finely tomentose all over; stem simple, usually 
slender, erect, 10—2° high. Leaves oval, ovate, 
oblong or ovate-lanceolate, acute or obtuse and 
mucronulate at the apex, rounded or narrowed at 
the. base, 2-3’ long, 3’-13’ wide, short-petioled, 
the upper surfaces becoming glabrate at maturity; 
umbels solitary or few, several-many-flowered; 
peduncles short; corolla greenish-white or pur- 
plish, its segments ovate-oblong, obtuse, 2’~—3” 
long; column very short; hoods oval-oblong, yel- 
lowish, nearly twice as long as the anthers, bear- 
ing a large acute tooth on each of the inner mar- 
gins; horn subulate, incurved over the stigma; 
follicles ascending on the reflexed fruiting pedi- 
cels, pubescent. 


In woods and on prairies, Illinois to North Dakota, 
Wisconsin, Manitoba and Athabasca. June-July. 


20. Asclepias brachystéphana |*ngelm. 
Short-crowned Milkweed. Fig. 3402. 


Asclepias brachystephana Engelm.; Torr. Bot. Mex. 
Bound. Surv. 163. 1859. 


Puberulent when young, soon glabrate; stems 
clustered, often branched, spreading or ascend- 
ing, 6-12’ long. Leaves mostly opposi'e, lanceo- 
late or linear-lanceolate, thick, long-acuminate at 
the apex, rounded, subcordate or narrowed at the 
base, 2’-5’ long, 2-6” wide, or the lowest shorter; 
petioles 1’-4” long; umbels several, terminal and 
axillary, few-flowered; peduncles short; pedicels 
densely woolly, equalling or longer than the pe 
duncles; corolla greenish-purple, its segments 
about 2” long; column very short or none; hoods 
ovate, obtuse, shorter than the anthers, the short 
erect-incurved horn slightly exserted; follicles 
erect on the spreading or decurved fruiting pedi- 
cels, downy or hoary, acuminate, 2-33’ long 


i 


WS 
Xd 


21. Asclepias perénnis Walt. Thin-leaved 
Milkweed. Fig. 3403. 


NY 
NN 


In dry soil, Kansas (according to B. B. Smyth) 
Wyoming to Texas, Arizona and Mexico. |une-—Aug, 


Asclepias perennis Walt. Fl. Car. 107. 1788. 


Puberulent above, glabrous below; stem slen- 
der, simple or branched, erect, 1°-3° high. Leaves 
thin, opposite, lanceolate, oblong or ovate-lanceo- 
late, slender-petioled, acuminate or acute at both 
ends, 2’-6’ long, #’-1’ wide, nearly glabrous; um- 
bels solitary or several and corymbose; peduncles 
1-2’ long; pedicels slender, 3’-1’ long; flowers 
small, white; corolla-segments oblong, 1°-2' 
long; column about 4” high; hoods oval, erect, 
entire, about as long as the anthers, shorter than 
the subulate-filiform incurved horn; follicles gla- 
brous, erect on the nearly erect fruiting pedicels; 
seeds 5-6” long, 33’-42” wide, thin, usually with- 
out coma. 


On river-shores and in wet places, North Carolina 
to Indiana, Illinois and Missouri, south to Florida 
and Texas. May-Aug. 


ASCLEPIADACEAE. Vo. III. 


22. Asclepias verticillata L. Whorled Milk- 
weed. Fig. 3404. 
Asclepias verticillata L. Sp. Pl. 217. 1753. 


Roots slender, fascicled; stem slender, simple or 
branched, pubescent in lines above, leafy, 1°-23° 
high. Leaves narrowly linear, sessile, verticillate in 

3’s-7’s or some of them alternate, glabrous or very 
} nearly so, their margins narrowly revolute; umbels 
numerous, many-flowered; peduncles slender, 2-13’ 
long; pedicels almost filiform, shorter than the pe- 
duncles; corolla greenish white, its segments oblong, 
13-2” long; column about 3” high; hoods white, 
oval, entire, about equalling the anthers, much 
shorter than the subulate incurved horn; follicles 
erect on the erect fruiting pedicels, narrowly spindle- 
shaped, glabrous, 2’-3’ long. ; 

In dry fields and on hills, Maine and southern On- 


tario to Saskatchewan, south to Florida, Mexico and 
New Mexico. July—Sept. 


S 


23. Asclepias galioides H.B.K. Bedstraw Milk- i "i 
Gyy2 


weed. Fig. 3405. 
Asclepias galioides H.B.K. Nov. Gen. 3: 188, 1818. 


Glabrous, except the minutely pubescent stems and pedi- 
cels. Stems erect, 1° high or more, from a horizontal root- 
stock; leaves erect or spreading, in whorls of 2-6, narrowly 
linear, 2’-3’ long, the margins revolute; peduncles longer 
than the pedicels and shorter than the leaves; umbels 
9-13” in diameter; flowers greenish-white; corolla-seg- 
ments 2” long; hoods as high as the anthers, broadly 
rounded at the summit, dorsally hastate-sagittate, the ven- 
tral margins slightly involute, entire; horn arising from 
the base of the hood, long-exserted over the anthers; 
anther-wings minutely notched at the base; follicles erect 
on erect fruiting pedicels, attenuated, 2’-2?’ long, glabrous 
or minutely puberulent. 


Kansas to Colorado, Arizona and Mexico. May-July. 


24. Asclepias pumila (A. Gray) Vail. 
Low Milkweed. Fig. 3406. 


Asclepias pumila Vail; Britton & Brown, Ill, Fl. 3: 
12, 1898. 


Asclepias verticillata var. pumila A, Gray, Proc. 
Am, Acad. 12: 71. 1876. 


Stems 4’-10’ high, tufted from a woody root. 
Leaves very numerous, crowded, sometimes ob- 
scurely whorled, filiform-linear, 17-2’ long, 
smooth or minutely roughened, the margins rev- 
olute; umbels 2-several, short-peduncled, few- 
flowered; pedicels filiform, puberulent, 3’-4” 
long; corolla greenish white, its segments ob- 
long, 13’-2” long; column short; hoods white, 
erect, oblong, entire, equalling the anthers, 
shorter than the slender incurved horn; follicles 
erect on erect fruiting pedicels, narrowly spindle- 
shaped, 13’-3’ long, finely puberulent. 


Dry plains, Iowa to South Dakota, Arkansas, 
Texas, Wyoming, Colorado and New Mexico. 


GENUS 2. MILKWEED FAMILY. 33 


‘2, ASCLEPIODORA A. Gray, Proc. Am. Acad. 12: 66. 1876. 


Erect or decumbent perennial herbs, similar to Asclepias, with alternate or opposite 
entire leaves, and rather large flowers in terminal solitary or corymbed umbels. Sepals lan- 
ceolate. Corolla rotate, its segments spreading. Hoods oblong, inserted over the whole of 
the very short corona-column, curved upward, obtuse, crested within, at least in the upper 
part, slightly longer than the anther; at the sinuses between the hoods a small lobe or 
appendage, alternate with the anther-wings, simulating an inner crown. Anthers tipped with 
a scarious membrane, their wings horny, narrowed below, sometimes angled above the middle. 
Pollen-masses pendulous, pyriform, longer than their caudicles. Follicles ovoid or oblong, 
acuminate, with or without soft spinose processes, erect or ascending on the decurved or 
twice bent fruiting pedicels. Seeds comose. [Greek, gift of Aesculapius. | 


Five or six species, natives of the southern United States and Mexico, 
dora viridis (Walt.) A. Gray. 


Glabrous or nearly so; leaves oblong to ovate-lanceolate ; umbels corymbose. 
Stem rough-puberulent ; leaves lanceolate or linear, acuminate ; umbels solitary. 


Type species: Asclepio- 


1. A. viridis. 
2. A. decumbens. 


1. Asclepiodora viridis ( Walt.) A. Gray. 
Oblong-leaved Milkweed. Fig. 3407. 


Asclepias viridis Walt. Fl. Car. 107. 1788. 


Asclepiodora viridis A. Gray, Proc. Am. Acad. 12: 
66. 1876. 


Stem erect, puberulent above, simple, 1°-2° 
high. Leaves oblong to ovate-lanceolate, rather 
thin, obtuse and mucronulate or acute at the 
apex, narrowed or rounded at the base, short- 
petioled, 23-5’ long, 4-12’ wide; umbels 2-4, 
or sometimes solitary; peduncles 14’-2’ long; 
pedicels slender, about 2’ long; corolla green- 
ish, its segments, when expanded, oblong, ob- 
tuse or acute, 4-6” long, 2-3 times as long as 
the purplish or violet entire-margined hoods; 
anther-wings narrow, scarcely angled above; 
fruiting pedicels twice bent; follicles ascend- 
ing, puberulent, 2’-3’ long, sometimes with soft 
spinose projections. 


In dry soil, Illinois to Kansas, Texas, South 


Carolina and Florida. May-July. 
ry 2. Asclepiodora decimbens ( Nutt.) 
SG Berg Em A. Gray. Decumbent Milkweed. 
r\s 


"4 
oi 
Ee 


Fig. 3408. 


Ananthrix decumbens Nutt. Trans, Am. Phil. 
Soc. (IL) 57202... 2835-97. 

Asclepiodora decumbens A. Gray, Proc. Am. 
Acad. 12: 66. 1876. 


Stems decumbent or ascending, rough- 
puberulent, 10’-2° long. Leaves firm, linear 
to lanceolate, glabrous above, puberulent on 
the veins beneath, acuminate at the apex, 
narrowed at the base, 3-7’ long, 2°-8” 
wide; umbel solitary, many-flowered; pe- 
duncle 17-5’ long; pedicels stout, 4-1’ long; 
corolla depressed-globose in the bud, green- 
ish, its segments, when expanded, ovate or 
broadly oval, longer than the hoods; hoods 
purple, obtusely 3-lobed on the ventral 
margins, about 3” long, their tips incurved; 
anther-wings broad, angled above; follicles 
nearly erect on the recurved fruited pedi- 
cels, 3-4’ long, puberulent, at least when 
young, with or without soft projections. 


In dry soil, Kansas to Texas and Mexico, 
west to Utah and Arizona. April—June. 


34 ASCLEPIADACEAE. VoL. III. 


3. ACERATES Ell. Bot. S. C. & Ga. 1: 316. 1817. 


Perennial herbs, similar to Asclepias, with alternate or opposite thick leaves, and green 
or purplish flowers in terminal or axillary and short-peduncled or sessile umbels. Calyx 
5-parted or 5-divided, the segments acute, glandular within. Corolla deeply 5-cleft, the seg- 
ments valvate, reflexed in anthesis. Corona-column very short. Corona of 5 involute-concave 
or somewhat pitcher-shaped hoods, neither horned nor crested within or in one species having 
a small interior crest and usually a few small processes at the base of the anther-wings, 
forming an obscure inner crown. Pollen-masses solitary in each sac, oblong, pendulous. 
Stigma 5-lobed. [Greek, without horn, referring to the crown.] ’ 


About 7 species, natives of North America. Type species: Acerates longifolia (Michx.) Ell. 


Umbels sessile, or very nearly so, mostly axillary. 
Leaves oval to linear; hoods entire at the apex, 1. A. viridiflora. 
Leaves narrowly linear ; hoods 3-toothed, 2. A. angustifolia, 
Umbels, at least the lower, distinctly peduncled. 
- Plants glabrous, or nearly so; umbels usually several; leaves narrow. 


Hoods obtuse, entire; column 12” long; stem roughish puberulent. 3. A. floridana, 
Hoods emarginate ; column very short; stem glabrous. 4. A. auriculata. 
Plant hirsute; umbel solitary, terminal; leaves ovate to oblong. 5. A. lanuginosa. 


1. Acerates viridiflora (Raf.) Eaton. Green Milkweed. Fig. 3409. 


Asclepias viridiflora Raf. Med. Rep. (II.) 5: 360. 1808, 
Acerates viridiflora Eaton, Man. Ed. 5, 90. 1829. 


Puberulent or tomentulose, at least when young; 
stems simple, reclined or ascending, rather stiff, 
1°-3° high. Leaves slightly rough, alternate or op- 
posite, thick, oval, oblong or ovate to lanceolate or 
linear, 1-5’ long, 4#/—2’ wide, short-petioled, the mar- 
gins usually undulate; umbels several, or rarely 
solitary, axillary, densely many-flowered, sessile or 
very nearly so; pedicels very slender, tomentose, 
4-8” long; flowers green; corolla-segments nar- 
rowly oblong, 2’-3” long; column very short or 
none; hoods lanceolate-oblong, obtuse, minutely 
2-auricled at the base; mass of anthers longer than 
thick; anther-wings tapering below, semi-rhomboid 
above; follicles puberulent, 2’—4’ long. 


In dry, sandy or rocky soil, Massachusetts to south- 
ern Ontario, Saskatchewan, Florida and Texas. Con- 
sists of several races, differing mainly in leaf-form, 
June—-Sept. 


2. Acerates angustifolia ( Nutt.) Dec. 
Narrow-leaved Milkweed. Fig. 3410. 


Polyotus angustifolius Nutt. Trans, Am. Phil. Soc. 
CII.) 5: 201. 1833-37. 


A. angustifolia Dec. in DC. Prodr. 8: 522. 1844. 


Asclepias stenophylla A. Gray, Proc. Am, Acad, 12: 
72. 1876, ; 


Stems mostly several together, erect, straight, 
1°-2° high, puberulent above, glabrate below. 
Leaves opposite, or some of the lower alternate, 
sessile, narrowly linear, 2-5’ long, glabrous, the 
revolute margins and the thick midvein rough 
beneath; umbels 10-15-flowered, short-peduncled 
or subsessile, axillary, usually numerous; pedicels 
puberulent; corolla-segments oblong, greenish; 
hoods white, not exceeding the anthers, 3-toothed 
at the apex, the acute middle tooth merely a 
prolongation of the thickened crest-like midvein, 
shorter than the obtuse lateral ones; anther- 
wings notched at about the middle; follicles 
slender, erect, about 3’ long or more. 


On dry plains, Missouri, Nebraska and Colorado to 
Texas, 


GENUS 3. MILKWEED FAMILY. 35 


3. Acerates floridana (Lam.) A.S. Hitche. 
Florida Milkweed. Fig. 3411. 


FOYLE Sion Sone cee 


tory, 
Pal 


Cs) 


Asclepias floridana Lam. Encycl. 1: 284. 1783. 

Acerates longifolia Ell. Bot. S.C. & Ga.1: 317. 1817. 

Acerates floridana. A. S. Hitche, Trans. St. Louis 
Acad. 5: 508 1891. 


rts 
yen 


iz 
gi; 
abet 
Be | 
= 


Rough-puberulent; stems slender, simple or 
little branched, erect or ascending, 1°-3° high. 
Leaves mostly alternate, linear or rarely linear- 
lanceolate, acute or acuminate, short-petioled, 
2’-8’ long, 13”-6” wide, commonly rough-cilio- 
late on the margins and midrib; umbels several 
or solitary, peduncled, usually many-flowered; 
peduncles 3-15” long; pedicels slender, hirsute, 
3-1’ long; corolla greenish white, its segments 
narrowly oblong, about 2” long; column short 
but distinct; hoods oblong, obtuse, entire, shorter 
than the anthers; anther-wings narrowed to the 
base; follicles densely puberulent, 4’-5’ long. 


Moist soil, Ohio to southern Ontario and Minne- 
sota, North Carolina, Florida and Texas. June-Sept. 


4. Acerates auriculata Engelm. <Auri- 
cled Milkweed. Fig. 3412. 


Aceraies auricuiata Engelm. Bot. Mex. Bound. 
Surv. 160. 1859. 

Asclepias auriculata Holzinger, Bot. Gaz. 17: 
125. 1892. ‘ 


Stem glabrous, often glaucous, sinuous 
above, rarely branched below, usually stout, 
1°-3° high. Leaves mostly alternate, narrowly 
linear, glabrous, short-petioled, 3-8 long, 
1” pada : : 

3-23" wide, becoming leathery, the: rough 
margins not revolute; umbels commonly sey- 
eral, densely many-flowered, pedincled; pe- 

: duncles 2’—1’ long, pubescent; pedicels slender, 

: pubescent; flowers greenish white tinged with 
— \ dull purple; corolla-segments oblong, 2-23” 
long; column short, but distinct; hoods yellow, 
often with a purplish keel, entire, or emargi- 
nately truncate at the apex, not exceeding the 
anthers, the involute margins spreading at the 
base into auricles; follicles 2-3’ long, curved. 


CORP 
TS 
<A 


st 
oe 


vy 


. 


f& 
* 
fd) 
a a 

Wi 


In dry soil, Nebraska and Colorado to Texas 
and New Mexico. June-Sept. 


5. Acerates lanuginosa (Nutt.) Dec. 
. Woolly Milkweed. Fig. 3413. 


Asclepias lanuginosa Nutt. Gen, 1: 168. 1818. 
Acerates lanuginosa Dec. in DC. Prodr.8: 523. 1844. 


Hirsute all over; stems erect, slender, simple, 
6-18 high. Leaves oblong, ovate-lanceolate or | 
lanceolate, obtuse at the apex, narrowed or 
rounded at the base, short-petioled, 1-4’ long, 
4’-15” wide; umbel solitary, terminal, densely 
many-flowered, peduncled; peduncle stout, densely 
hirsute, 3’-13’ long; pedicels slender, puberulent 
or hirsute; corolla greenish, its segments oblong, 
about 23” long; column none; hoods purplish, 
oblong, obtuse, entire, with a flat fold or auricle 
on the infolded lower ventral margins, shorter 
than the anthers; anther-wings broadest below 
the middle. 


On prairies, northern Illinois to Minnesota, Ne- 
braska and Wyoming. June—Aug. 


36 ASCLEPIADACEAE. Vor, III. 


Va 
4. GONOLOBUS Michx. Fl. Bor. Am. 1: 119. 1803. 
[Ampetanus Raf.; Britton, Bull. Torr. Club 21: 314. 1804.] 
[EnsLenra Nutt. Gen. 1: 164. 1818. Not Raf. 1817.] 

Perennial twining herbaceous vines, with petioled opposite cordate thin leaves, and small 
whitish flowers in axillary peduncled clusters. Calyx 5-parted, minutely glandular within, 
the segments lanceolate. Corolla campanulate, deeply 5-cleft, the lobes slightly contorted, 
nearly erect. Crown nearly sessile, of 5 membranous truncate lobes, each appendaged by a 
simple or 2-cleft awn. Stamens inserted at the base of the corolla, the filaments connate 
into a short tube; anthers terminated by an inflexed membrane; pollen-masses solitary in 
each sac, ellipsoid, pendulous. Stigma conic, slightly 2-lobed. Follicles thick, acuminate. 
Seeds comose. [Greek, referring.to the somewhat angled pod.] 


Three species, natives of America. Type species: 
Gonolobus laevis Michx. 


a" 


1. Gonolobus laévis Michx. Sand Vine. 
Enslen’s-vine. Fig. 3414. 
Gonolobus laevis Michx. Fl. Bor, Am, 1: 119. 1803. 


Enslenia albida Nutt. Gen. 1: 164. 1818. 
A. albidus Britton, Bull. Torr. Club 21: 314. 1894, 


Stem sparingly puberulent, at least above, high- 
climbing, slender. Leaves slender-petioled, ovate, 
gradually acuminate, deeply cordate, palmately 
veined, glabrous or very nearly so, entire, 3-7’ long, 
13-5’ wide; petioles 1-4’ long; cymes numerous, 
densely flowered; peduncles stout, 3’—2’ long; flow- 
ers 2’-3” long; corolla-segments lanceolate, acute, 
twice as long as those of the calyx and exceeding 
the 2-cleft awns of the corona-lobes; follicles on 
ascending fruiting pedicels, 4-6’ long, somewhat 
angled, glabrous when mature. 


Along river-banks and in thickets, Pennsylvania to 
Illinois, Kansas, Florida and Texas. June—Aug. 


ZB 5. CYNANCHUM L. Sp. Pl. 212, “eee 
[VinceTox1icum Moench, Meth. 717. 1794.] 

Perennial twining herbaceous or slightly woody vines (some species erect herbs), with 
opposite or rarely verticillate or alternate leaves, and small yellowish green or purplish flow- 
ers in axillary cymes. Calyx 5-parted, minutely glandular within. Corolla rotate, deeply 
5-cleft, the segments spreading, somewhat twisted. Crown flat or cup-like, entire, 5-lobed or 
5-parted, the lobes not appendaged. Stamens attached to the base of the corolla, their fila- 
ments connate into a tube; anthers appendaged by an inflexed membrane. Pollen-masses 
solitary in each sac, pendulous. Stigma flat or conic. Follicles acuminate, glabrous. Seeds 
comose. [Greek, dog-strangling.] 

About 100 species, natives of both the Old World 
and the New. Besides the following, 2 native species 


occur in the southeastern United States. Type spe- 
cies: Cynanchum acutum L. 


1. Cynanchum nigrum (L.) Pers. Black 
Swallow-wort. Fig. 3415. 

Asclepias nigra L. Sp. Pl. 216. 1753. 

Vincetoxicum nigrum Moench, Meth, 317. 1794. 

Cynanchum nigrum Pers. Syn. 1: 274. 1805. 

Twining, or at first erect, puberulent, slender, 
2°-5° high. Leaves ovate or ovate-lanceolate, en- 
tire, thin, acuminate at the apex, rounded at the 
base, pinnately veined, petioled, 2’-5’ long, 4’-23’ 
wide; petioles 2”-12” long; pedicels 14”-3” long; 
flowers dark purple, about 23” broad; corolla- 
segments pubescent within; crown fleshy, 5-lobed ; 
follicles on nearly straight fruiting pedicels, about 
2’ long, glabrous. 

In waste places, escaped from gardens, Massa- 
chusetts to Pennsylvania and Ohio and in British 
Columbia. Introduced from Europe. June—Sept. 

Cynanchum VincetOxicum (L.) Pers., with green- 
ish-white glabrous corollas, another Old World 
species, is recorded as escaped from cultivation in 
southern Ontario. 


GENUus 6. MILKWEED FAMILY. 37 


6. VINCETOXICUM Walt. Fl. Car. 104. 1788, 


Twining or trailing perennial vines, with opposite usually cordate leaves, and rather large 
purple, brown, white or greenish flowers in axillary cyme-like umbels or fascicles. Calyx 
5-parted or deeply 5-cleft, mostly 5-glandular within. Corolla rotate, very deeply 5-parted, 
the tube very short, the segments convolute in the bud. Corona (crown) annular or cup- 
shaped, entire, lobed or divided, adnate to the corolla. Stamens inserted on the base of the 
corolla, the filaments connate into a tube; anthers not appendaged, merely tipped, borne along 
or just under the margin of the flat-topped stigma, the sacs more or less transversely dehis- 
cent. Pollen-masses solitary in each sac, horizontal or nearly so. Follicles thick, acuminate, 
smooth, angled or tuberculate. Seeds comose. [Greek, subduing poison.] 

About 75 species, natives of America. Besides the following, some 10 others occur in the 
southern and southwestern United States. Type species: Vincetoxicum gonocarpos Walt. 


Crown annular, 1o-crenate ; follicles angled, not warty. 
Corolla about twice as long as the calyx. 1. V. suberosum. 
Corolla 3-4 times as long as the calyx. 2. V.gonocarpos, 
Crown cup-shaped, about as high as the anthers; follicles warty. 
Flowers purple to dull yellow. 


Corolla-segments oblong, 3”—4” long ; crown crenate. 3. V. hirsutum. 
Corolla-segments linear or linear-oblong, 5”—7” long. 
Crown merely crenate. 4. V. obliquum. 


Crown toothed or lobed. 
Crown 5-lobed, with a subulate 2-cleft tooth in each sinus, 
Crown to-toothed, the alternate teeth thinner and longer. 
Flowers white ; crown deeply cleft. 


V. carolinense. 
V. Shortii. 
V. Baldwinianum. 


oy 


1. Vincetoxicum suberosum (L.) Britton. Coast Vincetoxicum. Fig. 3416. 


Cynanchum suberosum L. Sp. Pl, 212. 1753. 
G. suberosus R, Br. in Ait. Hort. Kew. Ed. 2: 82, 1811. 
V, suberosum Britton, Mem, Torr, Club 5: 266, 1894. 


Stem pubescent or glabrous, slender, twining. 
Leaves thin, 2-5’ long, 1-3’ wide, ovate or ovate- 
oval, acute or abruptly acuminate at the apex, cor- 
date at the base; petioles 4’-2’ long; umbels com- 
monly few-flowered; peduncles 4-1’ long; pedicels 
_43’-1’ long, fleshy, nearly glabrous; corolla brown- 

purple, broadly conic in the twisted bud, its segments 
lanceolate or ovate-lanceolate, acute, pubescent or 
granulose within, 3-4” long, about twice as long as 
the calyx; crown an annular fleshy undulately 
to-crenate disk; follicles glabrous, 3-5-angled, when 
young fleshy, when mature dry and spongy, 4-6’ 
long, 1’ in diameter or more. 


In thickets, Virginia to Florida, mainly near the coast. 
May-July. 


leaved Angle-pod. Fig. 3417. 

Vincetoxicum gonocarpos Walt. Fl. Car. 104, 1788. 
G. macrophyllus Michx. Fl. Bor, Am, 1: 119. 1803. 
Gonolobus laevis var, macrophyllus A, Gray, Syn, Fl, 2: 

Part 2, Tos. IS7te 

Glabrous or pubescent, stems slender, climbing 
high. Leaves broadly ovate, thin, 3-8’ long, 2’-6’ 
wide, acuminate at the apex, deeply cordate at the 
base, the sinus narrow or the rounded auricles over- 
lapping; petioles 1-4’ long; umbels few-flowered; 
peduncles 17-3’ long; pedicels rather stout, glabrous 
or nearly so; corolla conic in the bud, not twisted, its 
segments lanceolate, glabrous, 4-5” long, 3-4 times 
as long as the calyx; crown a low obtusely undulate 
disk; follicles glabrous, similar to those of the pre- 
ceding species but usually shorter. 


Along rivers and in moist thickets, Virginia to South 
Carolina, Georgia, Indiana Missouri and Texas. 


38 ASCLEPIADACEAE. Vor. III. 


3. Vincetoxicum hirsitum (Michx.) Britton. 
Hairy Vincetoxicum. Fig. 3418. 

Gonolobus hirsutus Michx. Fl. Bor. Am. 1: 119. 1803. 
V. hirsutum Britton, Mem, Torr. Club 5: 266. 1894. 

Stem downy, slender. Leaves ovate, acuminate at 
the apex, deeply cordate at the base, downy, 2-4’ 
long, 1’-2%’ wide, the sinus narrow, or the lobes 
sometimes overlapping; petioles slender, pubescent, 
3’-2’ long; peduncles usually about equalling the 
petioles, sometimes longer; umbels few-several- 
flowered; corolla brown-purple to greenish yellow, 
ovoid in the bud, its segments oblong, very obtuse, 
3-4” long, minutely puberulent without, about 4 
times as long as the densely pubescent calyx; crown 
cup-shaped, fleshy, about as high as the anthers, the 
margin 10-crenate; follicles lanceolate, 3’-5’ long, 
muricate, puberulent; seeds entire. 


In thickets, Maryland to Florida, west to Tennessee. 
Running milkweed. Negro-vine. July—Aug. 


4. Vincetoxicum obliquum (Jacq.) Britton. Large-flowered Vincetoxicum. 
Fig. 3419. 

Cynanchum hirtum L. Sp. Pl. 212. 1753? 
Cynanchum obliquum Jacq. Coll. 1: 148. 1786. 
G. obliquus R. Br.; R. & S. Syst. 6: 64: 1820. 
V. obliquum Britton, Mem, Torr. Club 5: 266. 1894. 

Stem puberulent or hirsute, slender. Leaves 
pubescent, broadly ovate, short-acuminate at the 
apex, deeply cordate at the base with an open or 
closed sinus, 2’-8 long, 13’-6’ wide; petioles 
rather stout, pubescent, 1-4’ long; umbels slen- 
der-peduncled, few-several-flowered; pedicels very 
slender, 1-2’ iong; corolla narrowly conic in the 
bud, red-purple within, greenish and minutely 
pubescent without, its segments elongated-linear, 
obtuse, 6-7 times as long as the hirsute calyx; 
crown cup-shaped, as high as the anthers, fleshy, 
its margin Io-crenulate, the intermediate crenu- 
lations sometimes 2-dentate; follicles ovoid-lan- 
ceolate, 2’-3’ long, muricate. 


In thickets, Pennsylvania to Ohio, Missouri, Vir- 
ginia and Kentucky. July—Aug. 


ee) 
5. Vincetoxicum carolinénse (Jacq.) Britton. 
Carolina Vincetoxicum. Fig. 3420. 
Cynanchum carolinense Jacq. Coll. 2: 228. 1788. 


G. carolinensis R. Br.; R. & S. Syst. 6: 62. 1820. 
V. carolinense Britton, Mem. Torr. Club 5: 265. 1894. 


Stem hirsute. Leaves broadly ovate, acute or 


base with a narrow or closed sinus, 3-7’ long, 2-53” 
wide, pubescent, at least beneath; petioles hirsute, 
14’-4’ long; peduncles 2’-4’ long; pedicels very slen- 
der, 1’ long or more; corolla brown-purple, oblong- 
conic in the bud, puberulent without, its segments 
linear-oblong or linear-lanceolate, obtusish, 5’-6” 
long, 5-6 times longer than the hirsute calyx; crown 
cup-shaped, scarcely fleshy, 5-lobed, with a subulate 

_longer 2-cleft erect tooth in each sinus; follicles 
muricate. 


In thickets, Virginia to Missouri, south to South Caro- 
lina and Louisiana. May-July. 


short-acuminate at the apex, deeply cordate at the © 


GENUS 6. MILKWEED FAMILY. 39 


6. Vinceoxicum Snortii (A. Gray) Britton. Short’s Vincetoxicum. Fig. 3421. 


Gonolobus obliquus var. Shertii A. Gray, Syn, Fl, 2: 
part i, 104, 1878. = 

G. Shortii A. Gray, loc, cit. Ed. 2, 404. 1886. 

V. Short Britton, Mem. Torr. Club 5: 266. 1894. 


Stem pubescent, or hirsute with spreading 
hairs. Leaves downy, broadly ovate, acute or 
short-acuminate at the apex, deeply cordate at 
the base and when old with a narrow or closed 
-sinus, 4-7’ long, 14-52’ wide; petioles stout, 
pubescent, 14’-3’ long; peduncles usually longer 
than the petioles; umbels several-flowered; pedi- 
cels 1’ long or more; corolla oblong-conic in 
the bud, dark crimson-purple, its lobes linear, 
5-7" lorig, 5-7 times as long as the hirsute 
calyx; crown cup-shaped, fleshy, as high as the 
anthers, its margin about 10-toothed, the alter- 
nate teeth thinner and longer, emarginate or 
2-parted, the others broader, thicker, with an 
obscure internal crest or ridge below the sum- 
mit; follicles warty. 

In thickets, Pennsylvania to eastern Kentucky 


and Georgia. Flowers with the odor of the straw- 
berry-shrub, June—Aug. 


7. Vincetoxicum Baldwinianum (Sweet) Britton. Baldwin’s Vincetoxicum. 


Fig. 3422. 
Gonolobus Baldwinianus Sweet; A. Gray, Syn. Fl. 2: 

Part 1, 104. 1876. 

Vincetoxicum Baldwinianum Britton, Mem. Torr. Club 

5: 265. 18094. 

Stem pubescent and hirsute. Leaves downy, broadly 
ovate, acute, or short-acuminate at the apex, deeply 
cordate at the base, 3-6’ long, or more; petioles hir- 
sute, 1-2’ long; peduncles 6-12” long, usually longer 
than the pedicels; umbels several-many-flowered; 
corolla white or cream-color, the lobes thin, oblong, 
or becoming spatulate, 4’—5” long; crown thin, the 5 
broader lobes quadrate, emarginate, or obscurely 
toothed; in their sinuses a pair of very slender 
linear-subulate teeth of more than double their 
length, much surpassing the stigma. 


Missouri and Arkansas to Georgia. May—June, 


Periploca graéca L., silk-vine, a handsome woody 
climber, with glabrous ovate-oblong leaves and brown- 
ish flowers in umbels, the obtuse corolla-segments vil- 
lous on the inner side has been collected as an escape 
from cultivation, 


Family 19. DICHONDRACEAE Dumort. Anal. Fam. 20, 25. 1829. 


DIcCHONDRA FAMILY. 


Consists only of the following genus: 


1. DICHONDRA Forst. Char. Gen. Pl. 39. pl. jo. 1776. 


Prostrate or creeping slender annual (sometimes perennial?) silky-pubescent or glabrous 
herbs, with nearly orbicular cordate or reniform petioled entire leaves, and very small soli- 
tary axillary peduncled flowers. Sepals nearly equal, oblong or spatulate. Corolla open- 
campanulate, deeply 5-parted, the lobes induplicate in the bud. Stamens shorter than the 
corolla; filaments filiform. Ovary villous, deeply 2-parted, each lobe 2-celled; styles 2, simple, 
arising from the bases of the ovary-lobes; stigmas capitate. Fruit of 2 pubescent 2-valved 
or indehiscent 1-2-seeded capsules. [Greek, two-grained, referring to the capsules.] 

About § species, natives of warm and tropical regions. Besides the following, another occurs 
in the southwest. Type species: Dichondra repens Forst. 


40 DICHONDRACEAE, Vot. III. 


1. Dichondra carolinénsis Michx. Dichondra. 
Fig. 2423. 


Dichondra carolinensis Michx. Fl, Bor. Am. 1: 136. 1803. 


Somewhat pubescent, or glabrous; stems almost fili- 
form, creeping, rooting at the nodes, 6’-2° long. Leaves 
orbicular to reniform, deeply cordate, #12’ in diameter, 
palmately veined; petiole often much longer than the 
blade; flowers 1’-2” broad; peduncles filiform; sepals 
obtuse, spatulate or obovate; corolla yellow to white, 
shorter than the sepals, its lobes ovate to oblong; cap- 
sule 1” high or less. 


In moist or wet places, Virginia to Texas and Mexico, 
near the coast. Widely distributed in Central and South 
America. Has been regarded as referable to the Old 
World D. repens Forst.; the specific name evolvulacea was 
used for it in our first edition, in error. 


Family 20. CONVOLVULACEAE Vent. Tabl. 2: 394. 1799. 


MorNING-GLORY FAMILY. 


Herbs, some tropical species shrubs or trees, the stems twining, ascending, 
trailing or erect, with alternate exstipulate entire dentate lobed or dissected leaves, 
and regular perfect axillary cymose or solitary flowers. Calyx inferior, 5-parted 
or 5-divided, usually persistent, the segments or sepals imbricated. Coralla gamo- 
petalous, funnelform, salverform, campanulate, tubular or rarely subrotate, the 
limb 5-angled, 5-lobed or entire. Stamens 5, inserted low down on the tube of 
the corolla and alternate with its lobes, all anther-bearing, the filaments filiform, 
or dilated at the base, equal or unequal; anthers 2-celled, the sacs longitudinally 
dehiscent. Disk annular or none. Ovary superior, sessile, 2—3-celled, with 2 
ovules in each cavity, or falsely 4-6-celled with a single ovule in each cavity, 
entire or 2—4-divided; styles 1-3, terminal, or arising from between the ovary- 
divisions; ovules anatropous. Fruit a 2-4-valved capsule or of 2-4 distinct car- 
pels, in our species. Seeds erect, the testa villous, pubescent or glabrous; embryo 
plaited or crumpled; cotyledons foliaceous; endosperm fleshy or cartilaginous, 
usually scanty. 

About 45 genera and probably 1000 species, of wide geographic distribution, most abundant 
in the tropics. 


Style 2-cleft or 2-divided. 
Style 2-cleft or 2-parted. 1. Stylisma. 
Style 2-divided to the ovary, each division 2-cleft. 2. Evolvulus. 
Style entire up to the stigma. 
Stigma or stigmas capitate or globose. 


Corolla salverform; stamens and style exserted. 3. Quamoclit. 
Corolla funnelform or campanulate ; stamens and style included. 4. Ipomoea. 
Stigmas 2, filiform to oblong. 5. Convolvulus. 


1. STYLISMA Raf. Neog. 2. 1825. 


Herbs, mostly perennial and procumbent, with entire short-petioled or sessile leaves, and 
1-5-flowered axillary peduncles; flowers white, purple, pink, or yellow. Sepals acute or 
obtuse. Corolla campanulate or funnelform-campanulate; limb plaited, 5-angled or slightly 
5-lobed. Stamens included; filaments filiform, or dilated at the base. Ovary 2-celled; style 
2-cleft or 2-parted; stigmas capitate. Capsule globose to ovoid, 2-celled, 2-4-valved. Style 
rarely 3-parted and ovary 3-celled. Seeds 1-4, glabrous or pubescent. [Greek, referring to 
the 2-parted style.] 


Seven known species, of the southeastern United States and Mexico. In our first edition this 
genus was referred to the Australian Breweria R. Br. Type species: Convolvulus aquaticus Walt. 


Sepals acute or acuminate; leaves oblong, elliptic or linear. 


Corolla white; filaments pubescent; plant pubescent or puberulent. Z-.Ss humistrata, 
Corolla purple; filaments glabrous; plants silky-tomentose. 2. S. aquatica, 
Sepals obtuse ; leaves narrowly linear. 3. S. Pickeringu. 


= 


GENUS I. MORNING-GLORY FAMILY. 41 


ay (FG a 1. Stylisma humistrata (Walt.) Chapm. 
> aR: : - 
vi Southern Breweria. Fig. 3424. 
VV ; Ai Convolvulus humistratus Walt. Fl. Car. 94. 1788. 
af Stylisma humistrata Chapm, Fl. S. States, 346. 1860. 
= Bonamia humistrata A, Gray, Man. Ed. 5, 376. 1867. 
Breweria humistrata A. Gray, Syn. Fl. 2: Part 1, 217. 1878. 
Pubescent or puberulent; stems slender, 1°-2° long, 


simple, or with a few long branches. Leaves elliptic, 
oblong-elliptic, or ovate-oblong, obtuse and mucronulate 
or some of them emarginate at the apex, subcordate, 
rounded or narrowed at the base, 4-1’ wide, 1-2’ long; 
petioles 1-3” long; peduncles slender, longer than the 
leaves, 1-7-flowered, minutely bracted at the summit; 
sepals glabrous or puberulent, oblong, acuminate, 2’—3” 
\ long; corolla white, 6-8” long; filaments pubescent; 
2 style 2-cleft; capsule ovoid, acute, glabrous, about as 
3 long as the calyx. 
In dry pine barrens, Virginia to Florida and Louisiana. 
ee May—Aug. 


‘D> 


2. Stylisma aquatica (Walt.) Chapm. Water 
Breweria. Fig. 3425. 


Convolvulus aquaticus Walt. Fl. Car. 94. 1788. 
Stylisma aquatica Chapm, FI. S, States, 346. 1860. 
Bonamia aquatica A. Gray, Man. Ed. 5, 376. 1867. 
Breweria aquatica A. Gray, Syn. Fl. 2: Part 1, 217. 1878. 


Finely and densely silky-tomentose, branched, the branches 
long and slender. Leaves oblong, elliptic or oblong-lanceo- 
late, obtuse at both ends, mucronate or emarginate at the 
apex, sometimes subcordate at the base, 4’-13’ long, 2”-8” 
wide; peduncles 1-3-flowered, longer than the leaves, mi- 
nutely bracted at the summit; sepals densely silky-tomen- 
tose, oblong, acute or acuminate, about 2” long; corolla 
purple or pink, 5’-7” long; filaments glabrous; style 
2-parted nearly to the base. 


In wet soil, especially in pine barrens, Missouri to Texas, 
east to North Carolina and Florida. May—Aug. 


3. Stylisma Pickeringii (M. A. Curtis) A. Gray. Pickering’s Breweria. 
Fig. 3426, 


Convolvulus Pickeringii M. A. Curtis, Bost. Journ. Nat. 
Hist. 1) 1205, Los. 


Stylisma Pickeringiit A, Gray, Man. Ed. 2, 335. 1856. 
Bonamia Pickeringii A. Gray, Man. Ed. 5, 376. 1867. 
Breweria Pickeringii A, Gray, Syn, Fl.2: Part 1,217. 1878. 


Etem pubescent or puberulent, very slender, simple 
or branched, 1°-2° long. Leaves puberulent or glabrous, 
narrowly linear, obtuse or acutish at the apex, narrowed 
at the base, 1-23’ long, 4”—-2” wide, the lowest sometimes 
narrowly spatulate; petioles very short; peduncles slen- 
der, about as long as the leaves, with 1 or 2 linear bracts 
at the summit which are usually longer than the pedicels 
and calyx; sepals pubescent or hirsute, ovate to oval, 
obtuse, about 2” long; corolla white, about 1’ long; fila- 
ments nearly glabrous; style 2-cleft, above, exserted; 
capsule ovoid, acute, pubescent, longer than the calyx. 


In dry pine barrens, New Jersey to North Carolina; 
Illinois to Iowa, Louisiana and Texas. June—Aug. 


2, EVOLVULUS L. Sp. Pl. Ed. 2, 39%. 1762. 


Erect or diffuse branching, mostly silky-pubescent or pilose, annual or perennial herbs, 
with small usually entire leaves, and axillary solitary, racemose or paniculate, small blue 
pink or white flowers. Sepals nearly equal, acute or obtuse. Corolla funnelform, campanu- 
late or rotate, the limb plaited, 5-angled or 5-lobed. Stamens included or exserted; filaments 


42 CONVOLVULACEAE. Vor. III. 


filiform; anthers ovate or oblong. Ovary entire, 2-celled; style 2-divided to the base, or 
near it, each division deeply 2-cleft; stigmas linear-filiform. Capsule 2-celled, globose to 
ovoid, 2-4-valved, 1-4-seeded. Seeds glabrous. [Latin, unrolling.]. 


About 85 species, natives of warm and tropical regions. Besides the following, some 7 others 
occur in the southern United States. Type species: Evolvulus nummularius L, 


1. Evolvulus pilosus Nutt. Evolvulus. 
Fig: 24277 


Evolvulus argenteus Pursh, Fl. Am. Sept. 187. 1814. 
Not R. Br. 1810. 


Evolvulus pilosus Nutt. Gen. 1: 174. 1818. 


Perennial, densely silky-pubescent or villous; 
stems ascending or erect, 3-0’ high, very leafy. 
Leaves sessile, oblong, lanceolate or spatulate, 3-9” 
long, 1-3” wide, acute or obtuse at the apex, nar- 
rowed at the base; flowers solitary and nearly ses- 
sile in the axils; peduncles 2-bracted at the base, 
recurved in fruit, 1-2” long; sepals lanceolate, 
acute or acuminate; corolla funnelform-campanu- 
late, purple or blue, 3-6” broad; capsule 13”-2” in 
diameter, about as long as the sepals. 


On dry plains, North Dakota to Missouri, Nebraska, 
Mexico and Arizona. May-July. 


3. QUAMOCLIT [Tourn.] Moench, Meth. 453. 1704. 


Twining herbaceous vines, with petioled entire lobed or pinnately parted leaves, and 
cymose racemose or solitary peduncled axillary flowers. Sepals 5, herbaceous, equal, acumi- 
nate, mucronate or appendaged. Corolla salverform (usually scarlet in the following species), 
the tube narrow, somewhat dilated above, mostly longer than the spreading 5-lobed limb. 
Stamens and simple style more or less exserted; stigma capitate; ovary 2-celled or falsely 
4-celled, 4-ovuled. Fruit usually 4-celled and 4-seeded. [Greek, dwarf kidney-bean.] 


About 10 species, of warm and tropical regions, only the following in North America. Type 
species: Ipomoea coccinea L, 


Leaves pinnately parted into very narrow segments. tO: Quamoclit. 
Leaves cordate, acuminate, entire or angulate-lobed. 2. Q. coccinea, 


1. Quamoclit Quamoclit (L.) Britton. Cypress Vine. Indian Pink. Fig. 3428. 


Ipomoea Quamoclit L, Sp. Pl. 159. 1753: 
QO. vulgaris Choisy in DC. Prodr. 9: 336. 1845. 
Q. Quamoclit Britton, in Britt. & Brown, Ill. Fl. 3: 

22. 1808. 

Annual, glabrous; stem slender, twining to a 
height of 10°-20°. Leaves ovate in outline, pe- 
tioled or nearly sessile, 27’ long, pinnately parted 
nearly to the midvein into narrowly linear entire 
segments less than 1” wide; peduncles slender, 
commonly much longer than the leaves, 1-6-flow- 
ered; pedicels 1’ long or more, thickening in 
fruit; sepals oblong, obtuse, usually mucronulate, 
“3” long; corclla scarlet, rarely white, salver- 
form, 1-12’ long, the tube expanded above, the 
limb nearly flat, the lobes ovate, acutish; stamens 
and style exserted; ovary 4-celled; ovule I in each 
cell; capsule ovoid, 4-valved, about 5” high, twice 
as long as the sepals. 


2 


In waste and cultivated ground, Virginia to Florida, 
Kansas and Texas. Sparingly escaped from gardens 
farther north. Naturalized from tropical America. 
July-Oct. American red bell-flower. Sweet-william- 
of-the-Barbadoes. Cupid’s-flower. Red jasmine. 


OO ed 


ae = 


cha 


GENUS 3. MORNING-GLORY FAMILY. 43 


2. Quamoclit coccinea (L.) Moench. Small Red Morning-glory. Fig. 3429. 


Ipomoea coccinea L. Sp. Pl. 160. 1753. 
I. hederaefolia L. Syst. Ed. 10, 925. 1759. 


Quamoclit coccinea Moench, Meth, 453. 1794. 


Annual, glabrous or puberulent, stem twining 
to a height of several feet or trailing. Leaves 
ovate to orbicular, deeply cordate, long-acumi- 
nate, 2’-6’ long, entire or angulate-lobed, slender- 
petioled; peduncles few-several-flowered, usu- 
ally not longer than the leaves; sepals oblong, 
obtuse, about 2” long, subulate-appendaged ; co- 
-rolla scarlet, salverform, 10’—20” long, the limb 
obscurely 5-lobed; stamens and style slightly 
exserted; ovary 4-celled with 1 ovule in each 
cell; capsule globose, 4-valved, 3-4” in diameter. 


Along river-banks and in waste places, Rhode 
Island to Pennsylvania, Florida, Ohio, Missouri, 
Texas and Arizona, Naturalized from tropical 
‘America, or native in the Southwest. A hybrid of 
this species with the preceding is sometimes culti- 
vated. American jasmine. July—Oct. 


4. IPOMOEA L. Sp. Pl. 159. 1753. 


Twining trailing ascending or rarely erect herbs, annual or perennial, with large showy 
axillary solitary or cymose flowers. Sepals equal or unequal. Corolla funnelform or cam- 
panulate, the limb entire, 5-angled or 5-lobed, the tube more or less plaited. Stamens equal 
or unequal, included; filaments filiform, or dilated at the base; anthers ovate, oblong, or 
linear. Ovary entire, globose or ovoid, 2-4-celled, 4-6-ovuled; style filiform, included; stigmas 
I or 2, capitate or globose. Capsule globose or ovoid, usually septifragally 2-4-valved, 2-4- 
seeded. [Greek, worm-like. | 

About 400 species, of wide geographic distribution. Besides the following, some 30 others 
occur in southern and western North America. Known as Morning-Glory or False Bindweed. | 
Type species: Jpomoea pes-tigrinis L. 

Ovary 2-celled (rarely 4-celled) ; stigma entire or 2-lobed. 

Leaves cordate; stems trailing or twining. 

Perennial from an enormous rcot; corolla 2’—3’ long. 
Annual; roots fibrous; corolla 4”—6” long, white. 
Annual; corolla 1’-1 4’ long, pink or purple. 

Leaves linear; stems ascending or erect. 

Ovary 3-celled ; stigmas 3; leaves cordate. (Genus PHARBITIS,) 
Leaves entire; corolla 2’-2™%’ long. 
Leaves deeply 3-lobed, corolla 1’-1%’ long. 


. I. pandurata. 
. I. lacunosa. 

. I. trichocarpa. 
. I. leptophylia. 


. I. purpurea. 
. I hederacea. 


Aun BwWHH 


1. Ipomoea pandurata (L.) Meyer. Wild Potato Vine. Fig. 3430. 


Convolvulus panduratus L, Sp. Pl. 153. 1753. 
I, pandurata Meyer, Prim, Fl. Esseq. 100. 1818. 


Perennial from an enormous fleshy root, glabrous 
or puberulent; stems trailing or feebly climbing, 
2°-12° long. Leaves broadly ovate, cordate, acumi- 
nate at the apex, 2-6’ long, slender-petioled, entire, 
sometimes contracted in the middle, or some of the 
later ones rarely angulate-dentate or 3-lobed; pe- 
duncles 1-5-flowered, much elongated in fruit; 
sepals oblong, obtuse or acutish, 6-8” long, gla- 
brous; corolla funnelform, white, or with pinkish 
purple stripes in the throat, 2-3’ long, the limb 
5-lobed; ovary 2-celled; capsule ovoid, 2-valved, 
2-4-seeded, the seeds densely woolly on the margins 
and pubescent on the sides. 


In dry soil, in fields or on hills, Ontario to Con- 
necticut, Florida, Michigan, Kansas and Texas. Oc- 
curs rarely with double flowers. Man-of-the-Earth. 
Mecha-meck (Indian). Wild sweet potato. Man-root. 
Wild jalap. Scammony. May-Sept. 


44 CONVOLVULACEAE. Vow. III. 


2. Ipomoea lacundsa L. Small-flowered 
White Morning-glory. Fig. 3431. 


Ipomoea lacunosa L, Sp. Pl. 161, 1753. 


Annual, pubescent or hirsute, rarely glabrous; 
stem twining, 2°-10° long. Leaves slender-petioled, 
broadly ovate, cordate, acute or acuminate at the 
apex, entire, angled or 3-lobed, 2-4’ long, the 
lobes acute; peduncles 1-3-flowered, shorter than 
the leaves; pedicels slender; sepals oblong or lan- 
ceolate, acute or acuminate, pubescent or ciliate, 
about 5” long; corolla funnelform, 6’—10” long, 
white, or the limb purple; ovary 2-celled; stigma 
capitate; capsule globose, 2-valved, shorter than 
or about equalling the sepals. 


> In moist soil, Pennsylvania to South Carolina, Illi- 
nois, Missouri, Kansas and Texas. In ballast at 
Atiantic seaports. Whitestar. Morning-glory. July— 
Sept. 


3. Ipomoea trichocarpa Ell. Small-flowered 
Pink Morning-glory. Fig. 3432. 


Convolvulus carolinus L. Sp. Pl. 154. 1753. 
Ipomoea trichocarpa Ell. Bot. S. C, & Ga. 1: 258. 1817. 
Ipomoea commutata R, & S. Syst. 4: 228. 1819. 


Ipomoea carolina Pursh, Fl, Am. Sept.145. 1814. NotL. 

1753. : 

Similar in habit to the preceding species, but the 
leaves usually more lobed; peduncles often longer 
than the leaves, 1-3-flowered; sepals lanceolate or 
oblong-lanceolate, acuminate, pubescent or ciliate; 
corolla 1’-13’ long, pink or purple; capsule glabrous 
or pubescent. 


Kansas to Texas, east to South Carolina and Florida. 


4. Ipomoea leptophylla Torr. Bush Morning- 
glory. Fig. 3433. 
Ipomoea leptophylla Torr. in Frem, Rep. 95. 1845. 


Perennial from an enormous root, which some- 
times weighs 25 Ibs., glabrous throughout; stems 
erect, ascending or reclining, rather stout, 2°-4° 
long, much branched. Leaves narrowly linear, en- 
tire, acute, 2-5’ long, 1’-3” wide; petioles very 
short; peduncles stout, nearly erect, usually shorter 
than the leaves, 1-4-flowered; pedicels shorter than 
the peduncles; sepals broadly ovate, obtuse, 3-4” 
long, or the outer shorter; corolla funnelform, pur- 
ple or pink, about 3’ long, the limb scarcely lobed; 
capsule ovoid, acute, 8-12” long, 2-celled, much 
longer than the sepals; seeds pubescent. 


In dry soil, South Dakota to Nebraska, Wyoming, 
Texas and New Mexico. Man-root. May-July. 


GENUS 4. MORNING-GLORY FAMILY. 45 


5. Ipomoea purpurea (L.) Lam. Morning- 
glory. Fig. 3434. 


Convolvulus purpureus L. Sp. Pl. Ed. 2, 219. 1762. 

Ipomoea purpurea Lam. Tabl. Encycl. 1: 466. 1791. 

pce bits purpurea Voigt. Hort. Sub. Calcutta 354. 
1845. 


Annual, pubescent; stem retrorsely hairy, twin- 
ing or trailing, 4°-10° long. Leaves broadly ovate, 
deeply cordate, acute or acuminate, 2’-4’ wide, 
slender-petioled; peduncles slender, 1-5-flowered, 
often longer than the petioles; sepals lanceolate 
or oblong, acute, pubescent or hirsute near the 
base, 6-8” long; corolla funnelform, blue, pur- 
ple, pink, variegated or white, 2-22’ long; ovary 
3-celled (rarely 2-celled) ; stigmas 3 (rarely 2); 
capsule depressed-globose, about 5” in diameter, 
shorter than the sepals. 


In waste places, commonly escaped from gardens, 
Nova Scotia to Florida, west to Ontario, Nebraska 
and Texas. There is a double-flowered form in cul- 
tivation. Adventive or naturalized from. tropical 
America. Ropewind. July—Oct. 


6. Ipomoea hederacea Jacq. Ivy-leaved 
Morning-glory. Fig. 3435. 


Ipomoea hederacea Jacq. Icon. Rar. pl. 36. 1781. 
Peres hederacea Choisy, Mem. Soc. Gen. 6: 440. 

1833. 

Annual, pubescent; stem twining or climbing to 
a height of 2°-5°, slender, retrorsely hairy. Leaves 
ovate-orbicular in outline, long-petioled, deeply 
3-lobed, cordate at the base, 2’-5’ long, the lobes 
ovate, acuminate, entire, or the lateral ones some- 
times repand or dentate; peduncles 1-3-flowered, 
much shorter than the petioles; flowers opening in 
early morning, soon closing; sepals lanceolate with 
long linear often recurved tips, densely hirsute be- 
low, sparingly so above, 8’—12” long; corolla funnel- 
form, the tube usually nearly white, the limb light 
blue or purple, 1-13" long; ovary 3-celled; stigmas 
3; capsule depressed-globose, 3-valved, about as 
long as the lanceolate portion of the sepals. 

In fields and waste places, Maine to Florida, Penn- 
sylvania, Nebraska and Mexico. Naturalized or adven- 
tive from tropical America. July—Oct. 


6. CONVOLVULUS L. Sp. Pl. 153. 1753: 


Herbs (the following species perennials with slender roots or rootstocks) with trailing, 
twining or erect stems. Leaves entire dentate or lobed, mostly cordate or sagittate and 
petioled. Flowers axillary, solitary or clustered, large, pink, purple or white. Sepals nearly 
equal or the outer larger, the calyx bractless or with a pair of bracts at its base. Corolla 
funnelform or campanulate, the limb plaited, 5-angled, 5-lobed, or entire. Stamens inserted 
on the tube of the corolla, included; filaments filiform, or dilated at the base. Ovary 1-2- 
celled, 4-ovuled; style filiform; stigmas 2, filiform, oblong, or ovoid. Capsule globose or 
nearly so, 1-4-celled, 2-4-valved. Seeds glabrous. [Latin, to roll together, or entwine.] 


About 200 species, of wide distribution in tropical and temperate regions, Besides the follow- 
ing, some 30 others occur in the southern and wesiern United States. Type species: Convolvulus 
sepium L, 


Calyx with two large bracts at the base, which enclose it. 
Stems trailing or climbing. ; 
Peduncles long, much longer than the petioles. 
Stems 3°-10° long; leaves hastate, the auricles often dentate. 
Stems 1°-3° long; leaves sagittate, the auricles rounded, entire, 
Peduncles short, mostly not longer than the petioles. 
Stem erect or ascending; flowers white; bracts not cordate. 
Calyx not bracted ; peduncle bracted at the summit. 
Glabrous or nearly so; leaves entire, auriculate. 
Canescent ; leaves with 2-4 basal lobes, 


C. sepium. 

C. repens, 

C. fraterniflorus, 
. C. spithamaeus. 


PwdhH 


. C. arvensis, 
. Cy incanus. 


aur 


46 CONVOLVULACEAE. Vor. LII. 


1. Convolvulus sépium L. Hedge or 

Great Bindweed. Lily-bind. Fig. 3436. 
Convolvulus sepium L. Sp. Pl. 153. 1753. 
Convolvulus sepium var. americanus Sims, Bot. 

Mag. pl. 732. 1804. 

Calystegia sepium R. Br. Prodr, Fl. Nov. Holl. 1: 

483. 1810. 

Glabrous or sparingly pubescent; stems ex- 
tensively trailing or high-twining, 3°-10° long. 
Leaves slender-petioled, triangular in outline, 
hastate, 2’-5’ long, acute or acuminate at the 
apex, the basal lobes divergent, usually acute, 
angulate-dentate or entire; petioles 3-2’ long; 
peduncles 1-flowered, longer than the petioles, 
often 2-3 times as long; flowers pink with 
white stripes or white throughout, about 2’ 
long; bracts at the base of the corolla, large, 
ovate, acute or obtuse, cordate; stigmas oblong. 

In fields and thickets, usually in moist soil, 
Newfoundland to North Carolina, British Colum- 
bia, Illinois, Nebraska and New Mexico. Also in 
Europe and Asia. June-Aug. Bell-bind. Wood- 
bind. Pear- or Devil’s-vine. Lady’s-nightcap. 
Hedge- or harvest-lily. Rutland beauty. Wood- 
bine. German scammony. Creepers. Bracted- 
bindweed. 

Convolvulus japonicus Thunb. FI. Jap. 85. 1784, a species with narrow hastate leaves and 
smaller pink flowers, cultivated in a double-flowered form, has in this form escaped from cultiva- 
tion from southeastern New York to the District of Columbia and Missouri. 


2. Convolvulus répens L. Trailing or 
Hedge Bindweed. Fig. 3437. 
Convoivulus repens L. Sp. Pl. 153. 1753. 
Convolvulus sepium var. repens A, Gray, Syn. FI. 2: 

Part 1, 2050 evovios 

More or less pubescent or tomentose; stem 
trailing or twining, 1°-3° long, simple, or spar- 
ingly branched. Leaves ovate or oblong, petioled, 
1-2’ long, obtuse, acute or abruptly acuminate at 
the apex, sagittate or cordate at the base, entire, 
the basal lobes rounded, scarcely or not at all 
divergent; petioles 2-1’ long; peduncles 1-flow- 
ered, equalling or longer than the leaves; flowers 
white (sometimes pink?) about 2’ long; calyx en- 
closed by 2 ovate acute or obtusish slightly cor- 
date bracts; stigmas oblong. 

In moist and dry soil, Quebec to Florida and 
Louisiana. Recorded from the Great Lake region. 
May-Aug. 

C. intérior House, of the western plains, with 
broader leaves and smaller corollas, is found in Kan- © 
sas and Nebraska. 


3. Convolvulus fraterniflorus MacKenzie & Bush. 
Short-stalked Bindweed. Fig. 3438. 


C. Sepium fraterniflorus Mack. & Bush, Fl. Jackson Co. 153. 
1902. 


C. fraterniflorus Mack. & Bush, Rep. Mo. Bot. Gard. 16: 104. 
1905. 


Sparingly pubescent; stems trailing or twining, much 
branched, 3°-6° long. Leaves hastate or hastate-sagittate, 
short-pubescent on both sides, 4’ long or less, acute at the 
apex, the basal lobes entire or dentate, spreading; pedun- 
cles often 2 in each axil, wing-angled, mostly not longer 
than the petioles; bracts large, cordate, pubescent, con- 
cealing the sepals at flowering time; sepals glabrous, ob- 
tusish, 5”-8” long; corolla white, about 2’ long. 


Dry banks and prairies, western Missouri; recorded eastward 
to the District of Columbia. July—Sept. 


GENUs 6. MORNING-GLORY FAMILY. . 47 


4. Convolvulus spithamaéus L. Upright 
or Low Bindweed. Fig. 3439. 


Convolvulus spithamaeus L. Sp. Pl. 158. 1753. 
Calystegia spithamaea Pursh, Fl. Am. Sept. 143. 


Lol 4. 

Volvulus spithamaeus Kuntze, Rev. Gen. Pl. 447. 
18g1. 

Convolvulus camporum Greene, Pittonia 3: 328. 
1898. 


Pubescent, or glabrate; stem erect or ascend- 
ing, straight, or the summit sometimes feebly 
twining, 6-12’ high. Leaves oval, short-petioled 
or the uppermost sessile, usually obtuse at both 
ends, sometimes acutish at the apex, and subcor- 
date at the base, 1—2’ long, 3’-12’ wide; pedun- 
cles 1-flowered, longer than the leaves: flowers 
white, nearly 2’ long; calyx enclosed by 2 large 
oval acutish bracts which are narrowed at both 
ends and not cordate at the base; stigmas ob- 
long, thick. 


In dry sandy or rocky fields or on banks, Nova 
Scotia to Ontario, Manitoba, Florida and Ken- 
tucky. Dwarf morning-glory. Low or _ bracted- 
bindweed. May-—Aug. 


5- Convolvulus arvénsis L. Small Bindweed. 
Fig. 3440. 
Convolvulus arvensis L. Sp. Pl. 153. 1753. 


Glabrous, or nearly so; stems trailing or decum- 
bent, very slender, 1°-23° long, simple or branched. 
Leaves ‘slender-petioled, ovate or oblong, entire, ob- 
tusish and mucronulate or acutish at the apex, sagit- 
tate or somewhat hastate at the base, 1-2’ long, the 
basal lobes spreading, acute or obtuse; peduncles 
1-4-flowered (commonly 2-flowered), shorter than 
the leaves, 1-3-bracted at the summit, usually with 
another bracct on one of the pedicels; sepals oblong, 
obtuse, 12” long; corolla pink or nearly white, 8-12” 
broad; calyx not bracted at the base; stigmas linear. 

In fields and waste places, Nova Scotia to Ontario, 
Montana, New Jersey, Pennsylvania, Kansas, New 
Mexico and California. Naturalized from Europe. Na- 
tive also of Asia. May-Sept. Hedge-bells. Bearbind. 
Corn-lily. Withwind. Bellbine. Corn-bind. Lap-love, 
Sheep-bine. 


6. Convolvulus incanus Vahl. Hoary Bind- 
weed. Fig. 3441. 
Convolvulus inconus Vahl, Symb. Bot. 3: 23. 1794. 


Finely and densely canescent, pale, or some- 
times greener; stems procumbent or trailing, 
usually branched, 1°-3° long. Leaves rather short- 
petioled, lanceolate, ovate to linear in outline, 
usually with 2-4 divergent lobes at the base, or 
the lower pair of lobes reflexed, otherwise entire 
or irregularly dentate, obtuse and mucronulate at 
the apex, 1-2’ long; peduncles 1-2-flowered, as 
long as or longer than the leaves, minutely 
bracted at the summit; pedicels 3’-6” long; 
sepals oblong, obtuse or mucronulate, about 3 
long; corolla white to rose-color; stigmas nar- 
rowly linear; capsule globose, about as long as 
the sepals. 


In waste places, near Lincoln, Neb. (according to 
Webber). In dry soil, Kansas and Arkansas to 
Texas, Arizona and Mexico. Also in southern South 
America. April—Aug. 


48 CUSCUTACEAE. Vor. III. 


Family 21. CUSCUTACEAE Dumort, Anal. Fam. 20. 1829. 
DoppeR FAMILY. 

White or yellow slender parasites, dextrorsely twining, the leaves reduced to 
minute alternate scales, the small white, yellowish or pinkish flowers cymosely 
clustered. Calyx inferior, 5-lobed or 5-parted (rarely 4-lobed or 4-parted), or 
of 5 distinct sepals. Corolla campanulate, ovoid, urceolate or cylindric, 5-lobed 
(rarely 4-lobed), the lobes imbricated in the bud, the tube bearing as many 
fimbriate or crenulate scales as there are lobes and alternate with them, or these 
sometimes obsolete. Stamens as many as the corolla-lobes and alternate with 
them, inserted in the throat or sinuses above the scales, short-exserted or included ; 
filaments short or slender; anthers short, ovate or oval, obtuse, 2-celled, the sacs 
longitudinally dehiscent. Ovary globose to oblong, 2-celled; ovules 2 in each 
cavity ; styles 2, terminal, separate, or rarely united below ; stigmas linear or capi- 
tate. Capsule globose or ovoid, circumscissile, irregularly bursting or indehiscent 
1-4-seeded. Seeds glabrous, globose or angular; embryo linear, terete, curved 
or spiral, its apex bearing 1-4 minute scales, endosperm fleshy ; cotyledons none. 


1, CUSCUTA [Tourn.] L. Sp. Pl. 124. 1753. 


Characters of the family. The filiform twining stems are parasitic on herbs and shrubs 
by numerous minute suckers. The seeds germinate in the soil and the plantlet attaches itself 
to its host, its root and lower portion soon perishing. The subsequent nutrition of the 
parasite is apparently wholly through its suckers. [Name from the Arabic.] 

About 100 species, of wide geographic distribution. Besides the following, some 15 ‘others 
occur in the southern and western parts of North America. Known as Dodder, or Strangle-weed. 
Type species: Cuscuta europaea L. 

* Corolla-scales crenulate; stigmas slender; capsule circumscissile; introduced species. 
Scales crenulate above, not incurved. 1. C. Epilinum. 
Scales crenulate all around, strongly incurved. 2. C. Epithymum. 

** Corolla-scales fringed; stigmas capitate; capsule indehiscent; native species. 
Sepals united below into a gamosepalous calyx. 
Flowers very nearly sessile; corolla persistent at the base of the capsule. 
Corolla-scales ovate, fringed all around; calyx-lobes obtuse. 3. C. arvensis, 
Corolla-scales abortive, or of a few processes ; calyx-lobes acutish. 4. C. Polygonorum. 
Flowers distinctly pedicelled ; corolla enclosing or capping the capsule, or at length deciduous. 
Tips of the corolla-lobes incurved or reflexed. 
Scales ovate, fringed all around ; capsule enclosed by the corolla. 5. C.indecora. 
Scales abortive, or of a few slender processes ; corolla capping the capsule. 6. C. Coryli. 
Corolla-lobes spreading or recurved. 


Scales small, irregularly fringed; capsule depressed-globose. 7. C. Cephalanthi. 
Scales long, fringed mainly above ; capsule pointed. 
Corolla 114” long; capsule globose, short-pointed. 8. C. Gronovii, 
Flowers 2”—3” long; capsule oval, long-pointed. g. C. rostrata. 
Sepals separate, subtended by similar bracts. 
Flowers cymose, pedicelled ; scales short; bracts entire. to. C. cuspidata. 
Flowers closely sessile in dense clusters ; bracts serrulate. 
Bracts few, broad, appressed; styles as long as the ovary. 11. C. compacta. 


Bracts numerous, narrow, their tips recurved; styles longer than the ovary. 12. C. paradoxa, 


1. Cuscuta Epilinum Weihe. Flax Dodder. 
Fig. 3442. 


Cuscuta Epilinum Weihe, Archiv. Apoth. 8: 54. 1824, 
Cuscuta densiflora Soyer-Willem, Act. Soc, Linn. Paris 
4: 281. 1826. 


Stems very slender, yellow or red; flowers sessile 
in dense clusters, yellowish white, about 1%” long. 
Calyx hemispheric, 5-lobed, the lobes acute, nearly 
as long as the corolla-tube; corolla yellowish-white, 
short, cylindric, becoming urceolate, 5-lobed, the 
lobes ovate, acutish, spreading, its scales short, erect, 
less than one-half the length of the tube, 2-cleft or 
emarginate, crenulate above, the crenulations not ex- 
tending to the base; stigmas linear-filiform ; capsule 
circumscissile, the withering-persistent corolla borne 
on its summit. 


On flax, Nova Scotia to New Jersey and Pennsylvania. 
Introduced from Europe. Native also of Asia. July- 
Aug. 


GENUS I. DODDER FAMILY. 49 


2. Cuscuta Epithymum Murr. Thyme Dod- 
der. Lesser Lucerne or Clover Dodder. 
Fig. 3443. 

Cuscuta Epithymum Murr. in L. Syst. Ed. 13, 140. 1774. 

Cuscuta Trifolii Bab. Phytol. 1: 467. 1843. 


Stems filiform, red; flowers sessile in small dense 
clusters, pinkish, about 1” long. Calyx variable, 
4-5-lobed, more than one-half the length of the cylin- 
dric corolla-tube, the lobes acute; corolla 4~5-lobed, 
the lobes erect, about one-half as long as the tube, 
acute, its scales strongly incurved, crenulate nearly 
or quite to the base; stigmas filiform; capsule cir- 
cumscissile, capped by the withering corolla. 

Usually on clover, Maine and Ontario to South Da- 
kota and Pennsylvania. Introduced from Europe, where 


it occurs on thyme, clover and other low plants. Hail- 
weed. Hairweed. July—Sept. 


Cuscuta europaéa L., another Old World species, 
with obtuse calyx-lobes and shorter styles, is recorded 
from Maine. 


\ 4 
3. Cuscuta arvénsis Beyrich. Field 


oy, Dodder. Love-vine. Fig. 3444. 
We Cuscuta arvensis Beyrich; Hook. Fl. Bor. Am, 2: 


77. As synonym, 1834. 


Plant pale yellow; stems filiform, the flowers 
nearly sessile in small clusters. Calyx broad, 
5-lobed, the lobes broad, obtuse; corolla nearly 
campanulate, 5-lobed, the lobes acute or acumi- 
nate, as long as the tube, their tips reflexed, 
its scales large, ovate, as long as or longer 
than the tube, densely fringed all around with 
short irregular processes; stamens not ex- 
serted; style shorter than the ovary; stigmas 
capitate; capsule depressed-globose, indehis- 
cent, the withering corolla and usually the 
stamens persistent at its base. 

On various herbs and low shrubs, Massachusetts 
to Manitoba, Florida, Texas, Mexico and Cali- 


fornia. Also in the West Indies and South Amer- 
ica. July—Aug. 


4. Cuscuta Polygonorum Engelm. Smart- | 
weed Dodder. Fig. 3445. 


Cuscuta Polygonorum Engelm. Am. Journ, Sci. 43: 

342. pl. 6. f. 26-29. 1842. 
C. chlorocarpa Engelm.; A. Gray. Man. 350. 1848. 

Plant orange-yellow; stems slender but rather 
coarse; flowers sessile or nearly so in dense clus- 
ters. Calyx short, 4-5-lobed, the lobes ovate- 
oblong, acute or acutish; lobes of the corolla 4 
or 5, triangular-ovate, acute, mostly as long as 
the tube, the scales usually obsolete, wanting, or 
consisting of only 2 or 3 slender processes on 
each side of the attached lower portion of the 
filament; filaments mostly slender; styles shorter 
than the ovary; stigmas capitate; capsule globose, 
the withering corolla persistent at its base. 

On Polygonum and other herbs, Pennsylvania and 
Delaware to Minnesota, Wisconsin and Arkansas, 


July-Sept. Has been referred to the South Amer- 
ican C. obtusiflora H.B.K. 


4 


5° CUSCUTACEAE. Vor. III. 


5. Cuscuta indecéra Choisy. Pretty 
Dodder. Fig. 3446. 


Cuscuta indecora Choisy, Mem. Soc. Gen. g: 278. 
pl. 3. f. 5. 184i. 

C. pulcherrima Scheele, Linnaea 21: 750. 1848. 

Cuscuta decora Choisy; Engelm. Trans, St. Louis 
Acad. 1: 501. 1859. 


Stems rather stout; flowers 13” long, pedi- 
celled in loose cymes, more or less papillose. 
Calyx 5-lobed, the lobes ovate to lanceolate, 
acute, mostly shorter than the corolla-tube; 
corolla campanulate, 5-lobed, the lobes trian- 
gular, minutely crenulate, spreading, nearly as 
long as the tube, their tips inflexed; scales 
ovate, erect, irregularly fringed with short 
processes all around; stamens slightly exserted 
or included; stigmas capitate; capsule oblong, 
acute, enveloped by the withering corolla. 


On various herbs and low shrubs, Illinois to 
Nebraska, south to Florida, Texas and Mexico, 
in several races. Also in the West Indies and 
South America. Corolla white; stigmas often 
yellow or purple. June—Aug. 


6. Cuscuta Céryli Engelm. Hazel Dod- 


der. Fig. 3447. 


Cuscuta Coryli Engelm. Am. Journ. Sci. 43: 337. 
fe 7-II. 1842. 

Cuscuta inflexa Engelm, Trans, St. Louis Acad. 1: 
502. O50. 


Stems coarse; flowers about 1” long, pedi- 
celled in loose or rather dense cymes. Calyx 
4-5-lobed, the lobes triangular or triangular- 
lanceolate, acutish, about as long as the corolla- 
tube; corolla campanulate, 4-5-lobed, the lobes 
minutely crenulate, nearly erect, triangular, 
acute, about as long as the tube, their tips 
inflexed; scales small, oval, obtuse, often with 
only a few processes on each side; stamens 
scarcely exserted; styles shorter than the 
ovary; stigmas capitate; capsule oblong, point- 
ed, enveloped or at length capped by the with- 
ering corolla. 


On the hazels and other shrubs or tall herbs, 
Connecticut to Virginia, South Dakota and Arkan- 
sas. July—Aug. | 


in 


ep 
J 


Mare 
et Hee 


ie 


| 


7. Cuscuta Cephalanthi Engelm. Button- 
bush Dodder. Fig. 3448. 


Cuscuta Cephalanthi Engelm. Am. Journ. Sci. 43: 336. 
pl. 6. f. 1-6. 1842. 


Cuscuta tenuiflora Engelm.; A. Gray, Man. 350. 1848. 


Plant yellow, stems rather coarse; flowers about 
1” long, short-pedicelled, clustered; calyx 5-lobed, 
the lobes ovate, obtuse, shorter than the corolla-tube ; 
corolla cylindric-campanulate, its lobes ovate, obtuse 
and rounded, spreading, one-half the length of the 
tube or less; scales about as long as the lobes, 
fringed mainly toward the apex with irregular pro- 
cesses; stamens included; styles slender, about as 
long as the ovary, shorter than the ripe capsule; 
stigmas capitate; capsule depressed-globose, 13” in 
diameter, surrounded or capped by the withering, at 
length deciduous corolla. 


On shrubs and tall herbs, Pennsylvania to Minnesota, 
Texas and Arizona. July—Aug. 


GENUS I. DODDER FAMILY. 51 


8. Cuscuta Grondvii Willd. Gronovius’ 
Dodder. Love-vine. Fig. 3449. 


C. Gronovii Willd.; R. & S, Syst. 6: 205. 1820. 
Cuscuta vulgivaga Engelm, Am. Journ, Sci. 43: 
338. pl. 6. f. 12-16. 1842. 


Stems yellow to orange, slender, high-climb- 
ing; flowers usually short-pedicelled, numerous 
in dense cymes. Calyx not bracted, its lobes 
ovate, obtuse, shorter than the corolla-tube; 
corolla campanulate, about 13” long, the lobes 
ovate, obtuse, rounded, spreading, nearly as 
long as the tube, the scales narrow, equalling 
or longer than the tube, thickly fringed about 
the summit and sparingly along the sides with 
long slender processes; styles slender, not as 
long as the ovary; stigmas capitate; capsule 
globose, short-pointed or pointless, 13” in diam- 
eter, enveloped or capped by the withering 
deciduous corolla. 


On herbs and low shrubs, Nova Scotia to Mani- 
toba, Montana, Florida and Texas. Scald-weed, 
Devil’s-gut July—Aug. 


g. Cuscuta rostrata Shuttlw. Beaked 
Dodder. Fig. 3450. 


Cuscuta rostrata Shuttlw.; Engelm. Bost. Journ. 
Nat. Hist. 5: 225. 1845. 


Stems coarse, yellowish white; flowers larger 
than in any of our other species, loosely cy- 
mose, pedicelled. Calyx gamosepalous, 5-lobed, 
the lobes ovate-triangular, shorter than the 
corolla-tube; corolla campanulate, 2’—3” long, 
white, its lobes broadly ovate, obtuse, about as 
long as the calyx-lobes; scales narrow, some- 
times spatulate, shorter than the tube, heavily 
fringed at the summit and sparingly along the 
sides with long slender processes; stamens in- 
cluded; styles slender, about as long as the 
flask-shaped ovary; stigmas capitate; capsule 
oval, long-beaked. 


On herbs and shrubs, Maryland to South Caro- 
lina and Georgia, July—Sept. 


to. Cuscuta cuspidata Engelm. Cuspidate 
Dodder. Fig. 3451. 


Cuscuta cuspidata Engelm. Bost. Journ. Nat. Hist. 5: 
224.) 1845. 


Plant yellowish; stems slender; flowers about 
12” long in loose panicled cymes. Calyx of 5 
distinct entire sepals, shorter than the corolla- 
tube, with 2-4 similar bracts at its base and often 
others on the pedicels; sepals orbicular to lanceo- 
late, cuspidate, mucronate or acuminate; corolla 
nearly salverform, its lobes triangular-lanceolate 
ot oblong, acute or cuspidate, spreading, about 
one-half the length of the tube; scales narrow, 
usually less than one-half as long as the tube, 
fringed all around with short irregular processes ; 
stamens not exserted; styles very selnder, longer 
- than the ovary; stigmas capitate; capsule bearing 
the withered corolla on its summit. 


- On coarse herbs, Nebraska to Missouri and Texas. 
July—Sept. 


»CUSCUTACEAE. Vot. III. 


11. Cuscuta compacta Juss. Compact 
Dodder. Love-vine. Fig. 3452. 
Cuscuta compacta Juss.; Choisy, Mem. Soc. Gen. g: 

285. betes 2s LSAT. 

Plant yellowish white, stems rather stout; flow- 
ers about 2” long, closely sessile in dense clusters. 
Calyx of 5 (rarely 4) distinct oval crenulate ob- 
tuse sepals, subtended by 3-5 similar rhombic- 
orbicular appressed serrulate bracts; corolla sal- 
verform, persistent, the tube cylindric, its 5 
(rarely 4) lobes oblong or ovate, obtuse, spread- 
ing, much shorter than the tube, the scales nar- 
row, one-half the length of the tube, fringed with 
numerous long processes; stamens included; 
styles slender; capsule oblong, enveloped or 
capped by the withering corolla. 

On shrubs, Ontario to Massachusetts, New York 
and Alabama, west to Kansas and Texas. July—Sept. 


12. Cuscuta paradoxa Raf. Glomerate 
or American Dodder. Fig. 3453. 


Cuscuta paradoxa Raf. Ann. Nat. 13. 1820. 
Cuscuta glomerata Choisy, Mem. Soc, Gen. 9: 184. 
De As fete) LSAT. 


Plant yellowish white, stems slender; flowers 
sessile, 13” long, exceedingly numerous in dense 
confluent clusters covering portions of the stem 
of the host-plant. Calyx of 5 distinct concave 
oblong obtuse serrulate sepals, subtended by 
8-15 narrower serrulate much imbricated bracts 
with recurved tips; corolla tube oblong-cylin- 
dric, its lobes oblong-lanceolate or triangular- 
lanceolate, obtuse, spreading or recurved, per- 
sistent; scales copiously fringed at the summit 
and sparingly along the sides with numerous 
long processes; styles 2-4 times as long as the 
ovary; capsule capped by the withering corolla. 


On tall herbs, mainly Compositae, Ohio to South 
Dakota, Nebraska and Texas. July—Sept. 


Family 22. POLEMONIACEAE DC. FI. Franc. 3: 645. 1805. 
: PHLox FamIty. 

Herbs, some species slightly woody, with alternate or opposite entire lobed or 
dissected leaves. Flowers perfect, corymbose-capitate, cymose or paniculate, reg- 
ular, or nearly regular. Calyx inferior, persistent, tubular or campanulate, 5-cleft, 
the lobes or teeth slightly imbricated. Corolla gamopetalous, funnelform, saucer- 
shaped, campanulate or rotate, the limb 5-parted, convolute in the bud. Stamens 
5, inserted on the tube of the corolla and alternate with its lobes; filaments slender 
or filiform; anthers ovate, oblong or linear, versatile, 2-celled, the sacs longitudi- 
nally dehiscent. Ovary superior, mostly 3-celled; ovules 2- in each cavity, 
amphitropous; style simple filiform; stigmas 3, linear. Capsule mostly loculi- 
cidally 3-valved. Seeds various, sometimes winged, sometimes enveloped in muci- 
lage and emitting spiral tubes when wetted ; endosperm abundant; embryo straight ; 
cotyledons flat ; radicle inferior. 

About 20 genera and over 200 species, most abundant in western America. 


Calyx distended and at length ruptured by the ripening capsule. 
Calyx scarious between the lobes. 
Corolla salverform; leaves opposite, entire. 
Seeds not mucilaginous when wetted; mostly perennials with large flowers; leaves 
opposite. 1. Phlox. 
Seeds mucilaginous when wetted; annuals; floral leaves alternate; flowers small. 
2. Microsteris. 
Corolla funnelform, tubular, salverform or campanulate; leaves alternate or ones 
3. Gilia, 
Calyx not scarious between the lobes; leaves alternate, deeply cleft. 4. Leptodactylon. 


GENUS I. PHLOX FAMILY. 53 


Calyx not distended nor ruptured by the capsule; leaves alternate. 
Calyx-teeth herbaceous, not spinulose-tipped. 


Stamens declined ; leaves pinnate. 5. Polemonium, 
Stamens straight and leaves entire in our species, 6. Collomia. 
Calyx-teeth spinulose-tipped ; leaves pinnatifid. 7. Naverretia. 


1. PHLOX L. Sp. Pl. 1st. 1753. 


Perennial or rarely annual, erect or diffuse herbs; with opposite entire leaves, or some of 
the upper ones alternate, and large blue purple red or white flowers, in terminal cymes or 
cymose panicles. Calyx tubular or tubular-campanulate, 5-ribbed, 5-cleft, the lobes acute or 
acuminate, mostly scarious-margined and the sinuses commonly scarious. Corolla salver- 
form, the tube narrow, the limb 5-lobed; lobes obovate, orbicular or obcordate, spreading. 
Stamens straight, short, unequally inserted on the corolla-tube, included. Ovary oblong or 
ovoid, 3-celled; style usually slender; ovules 1-4 in each cavity. Capsule ovoid, 3-valved, at 
length distending and rupturing the calyx-tube. Seeds usually only 1 in each cavity of the 
capsule, ovoid, wingless or narrowly winged, not emitting spiral threads when wetted. 
[Greek, flame. ] 

About 40 species, natives of North America and Russian Asia, many of them widely cultivated. 
Besides the following, some 24 others occur in the southern and western parts of North Anierica. 
Type species: Phlox glaberrima L, 

Leaves flat, ovate, oblong, lanceolate or linear. 
Cymes panicled ; flowers short-pedicelled or sessile. 
Calyx-teeth subulate. 


Stem glabrous or puberulent ; leaves lanceolate to oblong, 1. P. paniculata. 
Stem villous, glandular above; leaves ovate to ovate-lanceolate. 2. P. amplifolia. 
Calyx-teeth lanceolate, acute ; leaves lanceolate or ovate, acuminate, 3. P. maculata, 


Cymes corymbose, simple, or flowers scattered. 
Flowering stems erect or ascending, simple. 
Plants glabrous or nearly so. 
Leaves ovate or oblong; calyx-teeth acute. 
Leaves lanceolate or linear; calyx-teeth subulate-lanceolate. 
Plants pubescent, hirsute or villous. 
Stems erect or ascending; no prostrate sterile shoots. 
Leaves linear or lanceolate, acuminate, spreading. 6 
Leaves linear-oblong, acute or obtuse, nearly erect. 7 
Stems ascending or reclining ; sterile shoots prostrate. 
Lower leaves and those of the sterile shoots oblong or ovate. 8. P. divaricata, 
Lower leaves and those of the sterile shoots obovate. g. P. stolonifera, 
Stems diffusely branched, usually creeping ; leaves narrow. 


. P. ovata. 
. P. glaberrima. 


nf 


. P. pilosa. 
. P. amoena, 


Corolla-lobes cleft to or about the middle. 10. P. bifida. 
Corolla-lobes cleft only at the apex. 11. P. Stellaria, 
Corolla-lobes rounded ; western. 12. P. Kelseyi. 


Leaves subulate, fascicled or crowded; plants low. 
Stems creeping or ascending; flowers cymose; eastern. 


Corolla-lobes shallowly emarginate ; plant not glandular. 13. P. subulata. 
Corolla-lobes deeply emarginate ; upper part of plant glandular, 14. P. Brittonii. 
Densely tufted ; flowers mostly solitary ; western. 
Leaves densely white-woolly, 1” long; plant moss-like, 15. P. bryoides, 
Leaves less woolly or merely ciliate, 2”—6” long. 
Corolla-tube shorter than or equalling the calyx. 16. P. Hoodii, 
Corolla-tube longer than the calyx. 17. P. Douglasii. 


1. Phlox paniculata L. Garden Phlox. 
Fig. 3454. 


Phlox paniculata L. Sp. Pl. 151. 1753. 


Stem erect, stout or slender, simple or branched 
above, glabrous or puberulent, 2°-6° high. Leaves 
thin, sessile or short-petioled, oblong to oblong-lanceo- 
late, acute or acuminate at the apex, narrowed at the 
base, or the uppermost subcordate, 2-6’ long, 4-13’ 
wide; flowers short-pedicelled in compact paniculate 
cymules, the inflorescence often 12’ long; calyx-teeth 
subulate, glabrous, puberulent or glandular, more than 
one-half as long as the tube; corolla pink, purple or 
white, its lobes broadly obovate, rounded, entire, shorter 
than its tube; capsule oval, obtuse, slightly longer than 
the ruptured calyx-tube. 


In woods and thickets, Pennsylvania to Florida, Illinois, 
Kansas and Louisiana. Freely escaped from gardens in 
the north and east. Consists of many races, differing in 
leaf-form, size and color of flowers, and in pubescence. 
July—Sept. 


54 POLEMONIACEAE. Vot. IIT. 


2. Phlox amplifolia Britton. Large-leaved 
Phlox. Fig. 3455. 


Phlox amplifolia Britton, Man. 757. t1g901. 


Stem villous or glandular-villous, at least above, 
2°-33° high. Leaves large and broad, 23-6’ long, 
13-23’ wide, roughish above, the upper sessile, the 
lower ones, or some of them, narrowed, usually 
abruptly, into winged petioles which are sometimes 
one-third as long as the blade; flowers similar to 
those of P. paniculata, the inflorescence often 1° 
long; calyx glandular-villous; corolla-tube glabrous, 
ae lobes obovate, rounded or retuse; capsules 4’-5” 
ong. 


Woods and thickets, Indiana to Missouri, Kentucky 
and Tennessee, June—Aug. 


3. Phlox maculata L. Wild Sweet- 
William. Fig. 3456. 


Phlox maculata L, Sp. Pl. 152. 1753. 
Phlox suaveolens Ait. Hort. Kew. 1: 206, 1789. 


Stem slender, erect, simple or branched above, 
glabrous or puberulent, usually flecked with pur- 
ple, 13°-3° high. Leaves lanceolate or the upper 
ovate-lanceolate, glabrous, rather firm, long- 
acuminate, sessile, rounded or subcordate at the 
base, 2’-5’ long, widest just above the base, the 
lowest sometimes linear-lanceolate; flowers 
short-pedicelled; the compact cymules forming 
an elongated narrow thyrsoid panicle; calyx- 
teeth triangular-lanceolate, acute, or acuminate, 
about one-fourth the length of the tube; corolla 
pink or purple, rarely white, its lobes rounded, 
shorter than the tube; capsule similar to that of 
the two preceding species. 


In moist woods and along streams, Connecticut 
to Florida, Ohio, Minnesota and Mississippi. Oc- 
casionally escaped from gardens further north. 
P. maculata var. candida Michx. (P. suaveolens 
Ait.) is a race with white flowers and unspotted 
stem, occurring with the type. June—Aug. 


4. Phlox ovata L. Mountain Phlox. 
Fig. 3457. 
Phlox ovata L, Sp. Pl. 152. 1753. 
Phlox carolina L. Sp. Pl. Ed. 2, 216. 1762. 


Glabrous or nearly so throughout; stems sim- 
ple, slender, ascending from a decumbent base, 
1°-2° high. Leaves rather firm, the upper ovate 
or ovate-lanceolate, sessile by a rounded or sub- 
cordate base, acute at the apex, 1’-2’ long, the 
lower and basal ones longer, oblong or ovate- 
oblong, acute at both ends, narrowed into slender 
often margined petioles; flowers short-pedicelled 
in corymbed or sometimes simple cymes; calyx- 
teeth lanceolate or triangular-lanceolate, acute, or 
acuminate, one-third to one-half the length of 
the tube; corolla pink or red, its lobes obovate, 
rounded, entire. 


In woods, Pennsylvania to North Carolina, Geor- 
gia and Alabama, mostly in the mountains. May—Aug. 


_ 


GENUS I... PHLOX FAMILY. 


5. Phlox glabérrima L. Smooth Phlox. 
Fig. 3458. 


Phlox glaberrima L. Sp. Pl. 152. 1753. 


Glabrous or nearly so throughout; stem simple, 
slender, erect or ascending, 1°-3° high. Leaves lan- 
ceolate or linear, rather firm, mostly 1I-nerved, 
acuminate at the apex, narrowed at the base, 13’-4’ 
long, 2’-6” wide, sessile, or the lowest linear or ob- 
long, obtusish, shorter, and short-petioled; flowers 
short-pedicelled, the cymules corymbed; calyx-teeth 
subulate-lanceolate, one-third to one-half the length 
of the tube; corolla commonly pink, its lobes obo- 
vate, rounded or obcordate, longer than the tube. 


In open woods and on prairies, Virginia to Illinois, 
Wisconsin, Florida, Mississippi, Missouri and Arkansas, 
Ascends to 2200 ft. in Virginia. May-July. 


— 


6. Phlox pilésa L. Downy or Prairie 
Phlox. Fig. 3459. 
Phlox pilosa L. Sp. Pl. 152. 1753- 


Soft downy or hairy, often glandular; stem 
erect or ascending, simple or branched, slen- 
der, 1°-2° high. Leaves linear or lanceolate, 
spreading or divaricate, long-acuminate, 1-4’ 


1/7" 


long, 14”-4” wide, sessile, the base narrowed 
or rounded; cymules corymbed; flowers short 
pedicelled; calyx glandular, viscid,. its teeth 
setaceous-subulate, longer than the tube; co- 
rolla pink, purple or white, its lobes obovate, 
entire, the tube usually pubescent; capsule 
shorter than the calyx. 

In dry soil, Ontario to Manitoba, Connecticut, 
New Jersey, Florida, Arkansas and Texas. Hairy 
phlox. Sweet-william, April-June. 


Phlox argillacea Clute & Ferriss is a recently 
described relative or race of this species, grow- 
ing on prairies in Indiana and Illinois. 


7. Phlox amoéna Sims. Hairy Phlox. 
Fig. 3460. 


Phlox amoena Sims, Bot. Mag. pi. 1308. 1810. 


Usually quite hairy; stems simple, slender, ascend- 
ing, 6-18’ high. Leaves linear-oblong to ovate- 
oblong, sessile, acute or obtuse at the apex, mostly 
narrowed at the base, nearly erect, 3-2’ long, 13’’-24” 
wide, the lowest much shorter; flowers very nearly 
sessile in a dénse terminal simple or somewhat com- 
pound cyme, which is subtended by the uppermost 
pair of leaves; calyx hirsute, its teeth subulate, as 
long as or shorter than the tube; corolla pink or 
white, its lobes obovate, entire or rarely emarginate, 
shorter than the glabrous tube. 


In dry soil, Virginia to Kentucky, Tennessee, Florida 
and Alabama. April-June. 


g. Phlox stolonifera Sims. Crawling 


Phlox. Fig. 3462. 


Phlox stolonifera Sims, Bot. Mag. pl. 563. 
Phlox reptans Michx. Fl. Bor, Am, 1: 145. 


POLEMONIACEAE. Vo. III. 


8. Phlox divaricata L. Wild Blue 
Phlox. Fig. 3461. 


Phlox divaricata L. Sp. Pl. 152. 1753. 


Finely viscid-pubescent; stems ascending or 
diffuse, slender, producing creeping or ascend- 
ing leafy shoots from the base. Leaves of the 
sterile shoots oblong or ovate, obtuse, 1-2’ 
long, those of the flowering stems lanceolate, 
ovate-lanceolate, or oblong, mostly acute or 
acutish; flowers pedicelled in open corymbed 
cymules, faintly fragrant; calyx-teeth subu- 
late, longer than the tube; corolla bluish, its 
lobes obcordate, emarginate or entire, not 
much longer than the tube, sometimes shorter ; 
capsule oblong-globose, about 2” high. 


In moist woods, Quebec to Ontario, Minnesota, 
Pennsylvania, Florida, Louisiana and Arkansas. 
Ascends to 3700 ft. in Virginia. Sometimes called 
wild sweet william. April-June. 


1802. 
1803. 


Hirsute or pubescent; stems slender, diffuse, 
producing sterile creeping leafy shoots from the 
base. Leaves of the sterile shoots obovate, obtuse 
at the apex, 1-3’ long, narrowed at the base into 
petioles; flowering stems 4-10’ high, their leaves 
oblong or lanceolate, acute or obtuse, smaller; 
flowers in a simple or barely compound cyme, 
slender-pedicelled; calyx-teeth linear-subulate, as 
long as the tube or longer; corolla pink, purple 
or violet, its lobes rounded, mostly entire, about 
one-half the length of the tube; capsule subglo- 
bose, 12” high. - 


In woods, Pennsylvania to Georgia and Kentucky, 


mainly in the mountains. Ascends to 4500 ft. in 
Virginia. April-June. 


10. Phlox bifida Beck. Cleft Phlox. 
Fig. 3463. 
P. bifida Beck, Am. Journ. Sci. 11: 170. 1826. 


Puberulent or pubescent; stems diffuse, 
somewhat woody, much branched, slender, 
often 1° long, the branches erect or ascend- 
ing, 4-8 high. Leaves of sterile shoots 
linear, sessile, 1-2’ long, 1-2” wide, acute, 
those of flowering branches linear-oblong or 
lanceolate, much shorter; flowers in simple 
cymes or solitary in the axils, slender-pedi- 
celled; pedicels 3-12” long; calyx-teeth lan- 
ceolate-subulate, somewhat longer than the 
tube; corolla pale purple, its lobes as long as 
the tube or somewhat shorter, cuneate, cleft 
to about the middle into linear or oblong 
obtuse diverging segments; capsule oblong- 
globose, 1”-12” high. 


In dry places, Indiana to Tennessee, Michigan 
and Missouri, April-June. 


GENUS I. PHLOX FAMILY. 


11. Phlox Stellaria A. Gray. Chick- 
weed Phlox. Fig. 3464. 


Phlox Stellaria A. Gray, Proc. Am. Acad, 8: 252. 
1870. 


Glabrous or puberulent; stems diffuse, some- 
what woody, much branched, the branches 
nearly erect, 3-8 high. Leaves all linear, or 
linear-lanceolate, sessile, acute, 17-2’ long, 
1-14” wide; flowers in simple cymes or soli- 
tary in the axils, slender-pedicelled; calyx- 
teeth subulate-lanceolate, shorter than the 
tube; corolla pale blue or nearly white, its 
lobes cuneate, 2-lobed at the apex, nearly as 
long as the tube. 


On cliffs, southern Illinois and Kentucky and 
in Tennessee. April-May. 


Q 


i 
12. Phlox Kélseyi Britton. Kelsey’s 
Phlox. Fig. 3465. 

P, Kelseyi Britton, Bull. Torr. Club 19: 225. 1892. 

Many-stemmed from a woody root, the 
stems spreading, creeping, or ascending, some- 
times 8 long, glabrous, or slightly pubescent 
above, very leafy. Leaves oblong, or linear- 
oblong, sessile, glabrous, or nearly so 3-12” 
long, 1-2” wide, or the upper longer and 
narrower, thick, rigid, the apex spinose- 
mucronate, the revolute margins ciliate; flow- 
ers sessile or short-peduncled; peduncles and 
calyx somewhat glandular-pubescent or gla- 
brous; calyx-teeth subulate, as long as the 
tube or longer; corolla-tube somewhat exceed- 
ing the calyx, the limb about 8” broad, blue 
or lilac, the obovate-cuneate lobes rounded or 
truncate. 

North Dakota to Nebraska, Montana and Wyo- 
ming. May-June. 


OC 
ISDS: 


Ss 


2.W Si 


y 
ve 


13. Phlox subulata L. Ground or Moss 
Pink. Fig. 3466. 


Phlox subulata L. Sp. Pl. 152. 1753+ 


Pubescent or becoming glabrate. Stems tufted, 
forming mats, diffuse, much branched, the 
branches 2’-6’ long. Leaves persistent, subulate- 
linear, linear-lanceolate or linear-oblong, acute or 
acuminate, 4-10” long, 3’-1” wide, spreading, 
ciliate, rigid, commonly fascicled at the nodes; 
flowers in simple cymes,  slender-pedicelled ; 
calyx-teeth subulate from a broader base, about 
as long as the tube; corolla pink, purple or white, 
with a darker eye, its lobes emarginate or entire, 
shorter than the tude; capsule oblong, nearly 2” 
high. 


In dry sandy or rocky soil, New York to Florida, 
west to Michigan and Kentucky. Ascends to 3500 
ft. in West Virginia. Wild or mountain-pink. Flower- ¢ 
ing moss, April-June. 


58 POLEMONIACEAE. Vor. III. 


14. Phlox Brittonii Small. Britton’s Phlox. 
Fig. 3467. 


Phlox Brittonii Small, Bull. Torr. Club 27: 279. 1900. 


Glandular-pilose, deep green; stems copiously 
branched, forming wide mats. Leaves numerous, 
with small ones often clustered in the axils of the 
larger, subulate or narrowly linear-subulate, 23’—5” 
long, ciliate, especially near the base; calyx glandu- 
lar-pubescent like the branches, the lobes subulate, 
about as long as the tube; corolla mainly white, the 
limb 12-13 mm. broad, its lobes cuneate, much shorter 
than the tube, with 2 pale magenta spots at the base, 
cleft by a V-shaped sinus, a minute tooth in each 
sinus; capsule oblong, 2” high. 


On dry mountain slopes, Virginia, West Virginia and 
C North Carolina, May. 


15. Phlox bryoides Nutt. Moss Phlox. 
Fig. 3468. 


Phlox bryoides Nutt. Journ. Acad. Phila, (II.) 1: 153. 
1848, . 


Depressed, shrubby, moss-like, densely branched 
from a deep woody root, forming compact tufts 2-3” 
high. Leaves minute (about 1” long), closely im- 
bricated in 4 ranks, copiously white-woolly, triangu- 
lar-lanceolate, pale, acute, the margins infolded; 
flowers solitary and sessile at the ends of the 
branches, about 23” long; tube of the corolla longer 
than the calyx, its lobes broadly cuneate, entire. 


On dry hills, western Nebraska, Colorado and Wyo- 
ming. May-July. 


16. Phlox Hoddii Richards. Hood’s Phlox. 
Fig. 3460. 


Phlox Hoodii Richards. App. Frank. Journ. 733. pl. 28. 
1823. 


Densely tufted and branched from a woody root, 
2’-4’ high. Leaves imbricated, erect, rigid, subulate, 
mucronate, somewhat woolly or ciliate, becoming 
glabrate, 2”-6” long; flowers solitary and sessile at 
the ends of the branches, about 5” long; calyx-teeth 
lanceolate, acuminate, rigid, longer than the tube; 
tube of the corolla shorter than or equalling the 
Y. VA calyx, its lobes obovate, entire. 


2 
» rey In dry sandy or rocky soil, North Dakota to Mani- 
al, Alberta, western Nebraska and Wyoming. May- 
july. 


GENUS I. PHLOX FAMILY. 


17. Phlox Douglasii Hook. Douglas’ 
Phlox. Fig. 3470. 


P, Douglasti Hook. Fl. Bor, Am. 2: 73. pl, 158. 1834. 

Phlox Douglasii andicola Britton, Mem. Torr. Club 
5: 269. 1894. 

Phlox Douglasui longifolia A. Gray, Proc. Am. Acad. 
8: 254. 1870. Not P. longifolia Nutt. 


Similar to the preceding species; leaves pubes- 
cent or glabrous, less imbricated, sometimes 
spreading, rigid, usually fascicled at the nodes, 
4-12” long. Flowers solitary and sessile or 
short-pedicelled at the ends of the branches, 
5-8” long; calyx-teeth narrowly lanceolate, 
acuminate, about equalling the tube; tube of the 
purple or white corolla longer than the calyx, its 
lobes obovate, entire. 


Dry soil, Nebraska and Montana to Utah, Cali- 
fornia and British Columbia, May-July. 


2. MICROSTERIS Greene, Pittonia 3: 300. 1808. 


Much-branched annual herbs, with entire leaves, all but the floral ones opposite, the small 
flowers solitary or in pairs in the upper axils. Calyx mostly campanulate, 5-cleft, scarious 
between the lobes. Corolla salverform, with a slender tube and a 5-lobed limb. Stamens 
short. Ovary 3-celled. Capsule at length distending and rupturing the calyx-tube. Seeds 
few- and large, mucilaginous when wetted, but not emitting spiral tubes. [Greek, small Steris.] 

About 6 species, of western North America. Type species: Microsteris grécilis (Dougl.) Greene. 


1. Microsteris micrantha (Kellogg) Greene. 
Small-flowered Microsteris. Fig. 3471. 


Collomia micrantha Kellogg, Proc. Cal. Acad. 3: 18. 


1863. 
M, micrantha Greene, Pittonia 3: 303. 1898. 


Annual, pubescent, at length corymbosely much 
branched, 2-6’ high, the branches ascending. Lower 
and basal leaves oblong to spatulate, obtuse, com- 
monly opposite and nearly sessile, the upper linear 
or lanceolate, sessile, 3-1’ long, 1-2” wide, opposite 
or alternate, entire; cymes I-5-flowered; calyx-lobes 
linear-subulate, usually longer than the tube; corolla 
about 4” long, the tube yellowish, narrow, equalling 
or slightly longer than the calyx, the lobes purple or 
violet, short; ovules 2 or 3 in each cavity; capsule 
oblong, obtuse, as long as or longer than the calyx- 
tube. 


WW In dry or moist soil, western Nebraska to Wyoming, 

Colorado, New Mexico, Arizona and California. In our 

first edition referred to the northwestern M, gracilis 
ss (Dougl.) Greene. April—Aug. 


3. GILIA R. & P. Prodr. Fl. Per. 47. 1798. 


Herbs with opposite or alternate, entire pinnatifid palmatifid or dissected leaves. Flowers 
small or large, solitary, cymose, capitate, thyrsoid, or paniculate. Calyx campanulate or 
tubular, 5-toothed or 5-cleft, the sinuses scarious. Corolla funnelform, tubular, campanulate, 
rotate or salverform, 5-lobed, the lobes ovate, oblong, or obovate. Stamens equally or 
unequally inserted on the corolla, included or exserted. Ovary oblong or ovoid, 3-celled; 
ovules solitary or several in each cavity. Capsule ovoid or oblong, 3-celled, at length dis- 
tending and rupturing the calyx. Seed-coat commonly mucilaginous when wetted, in some 
species emitting thread-like tubes. [Named for Philip Gil, a Spanish botanist. ] 

About 100 species, natives of America. Besides the following, many others occur in the south- 
ern and western parts of North America. Type species: Gilia /aciniata R. & P. 

Corolla funnel-form to salverform; leaves pinnatifid or pinnately divided, the segments linear, not 
rigid, ; 

Flowers thyrsoid-paniculate or corymbose-paniculate. 

Corolla 1-2’ long; plants 1°-4° tall. 
Flowers paniculate, white. 
Flowers narrowly thyrsoid, red. 

Corolla 3’—5” long, violet or blue. 

Flowers narrowly thyrsoid-spicate. 


. G. longiflora, 
. G. aggregata. 
. G. pinnatifida, 
. G. spicata, 


WRN He 


60 POLEMONIACEAE. Vor. IIL. 


Flowers in dense or capitate cymes, or heads; flower-clusters leafy-bracted. 
Perennial; corolla-tube not longer than the calyx. 5. G. iberidifolia. 
Annual; corolla-tube 2~3 times as long as the calyx. 6. G. pumila, 


Corolla rotate; leaf-segments acicular. 


7. G.acerosa, 


1. Gilia longiflora (Torr.) Don. White-flowered Gilia. Fig. 3472. 


2. Gilia aggregata (Pursh) Spreng. Scar- 
let Gilia. Fig. 3473. 
Cantua aggregata Pursh, Fl. Am. Sept. 147. 


Gilia aggregata Spreng, Syst. 1: 626. 


Biennial, pubescent or puberulent; stem simple 


Cantua longifiora Torr. Ann. Lyc, 2: 221. 1827. 
Gilia longiflora Don, Gard. Dict. 4: 245. 1838. 
Collomia longifl. A. Gray, Proc. Am. Acad. 8: 261. 1870. 

Annual, glabrous, paniculately branched, 1°-2° 
high. Leaves all alternate, sessile, 1-23’ long, pin- 
nately divided into linear-filiform segments or the 
uppermost entire; flowers numerous, white, panicu- 
late, about 2’ long; calyx-teeth triangular-lanceolate, 
acuminate, shorter than the tube; corolla salverform, 
its tube narrow, 4 or 5 times as long as the orbicular 
or ovate, rounded or pointed, spreading lobes; sta- 
mens unequally inserted; ovules 8-12 in each cell; 
capsule narrowly oblong, exceeding or equalling the 
calyx; seed-coat mucilaginous and emitting spiral 
threads when wetted. 

In dry soil, Nebraska and Colorado to Texas and Ari- 
zona. May-—Sept. 

Gilia rubra (L.) Heller (Gilia coronopifélia Pers.), a 
related species with less spreading corolla-lobes, is com- 
monly cultivated, and rarely escapes from gardens to 
roadsides and waste grounds, 


1814. 


or sparingly branched, 2°-4° high, leafy at least 
below. Leaves alternate, the basal often tufted, 
mostly petioled, 1-3’ long, pinnately parted into 
narrowly linear segments; inflorescence narrowly 
thyrsoid-paniculate, often 12’ long; flowers ses- 
sile or very nearly so in small peduncled clusters, 
scarlet or red; corolla tubular-funnelform, the 
tube 1-13’ long, slightly thicker upward, the limb 
cleft into ovate or lanceolate acute or acuminate 
spreading or recurved lobes; stamens unequally 
or about equally inserted in the throat; ovules 
numerous; seeds mucilaginous and emitting spiral 


threads when wetted. 


In dry soil, western Nebraska (according to 
Coulter) to Texas and Mexico, west to British 


Columbia and California. June—Aug. 


3. Gilia pinnatifida Nutt. Small-flowered 


Gilia. Fig. 3474. 
Gilia pinnatifida Nutt.; A. Gray, Proc. Am, Acad, 8: 

276. 1870. 

Biennial or perennial from a deep root, much 
branched, viscid-glandular, 62° high. Leaves 
thick, pinnatifid, the basal tufted, 1-3’ long, the 
segments linear-oblong, sometimes toothed, ob- 
tuse or acutish, 2-6” long, those of the stem 
alternate, smaller, the uppermost minute and en- 
tire; flowers very numerous, small, paniculate, 
some sessile, some petioled; calyx 5-lobed, the 
lobes lanceolate to ovate; corolla salverform, 
3-5” long, the limb violet or blue, its white tube 
longer than the calyx and its obovate lobes; sta- 
mens exserted; seeds not mucilaginous nor emit- 
ting spiral threads when wetted. 

In sandy soil, western Nebraska and Wyoming to 
New Mexico. Has been mistaken for G. inconspicua, 
June-Aug. 


GENUS 3. PHLOX FAMILY. 


4. Gilia spicata Nutt. Spicate Gilia. Fig. 3475. 


Gilia spicata Nutt. Journ, Acad. Nat. Sci. Phila. (II.) 1: 
156. 1848. 


Perennial, woolly-tomentose; stems erect, rather stout, 
simple, solitary, or 2-4 from the woody root, 6-18’ high. 
Leaves alternate, narrowly linear, pinnately parted into 
3-5 linear segments, or some or many of them entire, 
1-2’ long; flowers in an elongated narrow spicate 
thyrsus, sessile in small clusters, purplish, 4’-6” long; 
tube of the corolla somewhat exceeding the calyx, con- 
siderably longer than the ovate-oblong lobes; calyx- 
lobes acuminate; anthers equally inserted in the throat. 


In dry soil, western Nebraska to Wyoming and Utah. 
May-—Aug. 


5. Gilia iberidif6lia Benth. Round-headed 
Gilia. Fig. 3476. 


Gilia iberidifolia Benth. in Hook. Kew. Journ. Bot. 
BN S2005 Os Ws 


Perennial by a deep root, woolly-tomentose, at 
least when young, branched from the base or also 
above, 3-18 high. Leaves mostly petioled, 4’-2’ 
long, pinnately divided into 3-9 narrowly linear 
sharp-pointed segments, or the uppermost entire; 
flowers white, densely capitate-clustered, 2”—-3” 
long, the clusters bracted by the upper leaves, 
3’-1" broad, sometimes corymbed; calyx-lobes 
awn-like; corolla-tube about the length of the 
calyx, slightly longer than the oval lobes; fila- 
ments equally inserted in or below the sinuses of 
the corolla; ovules I-4 in each cavity. 

In dry soil, South Dakota to Nebraska, Colorado, 


California, Montana and Idaho. In our first edition 
included in the similar G. congesta Hook. May-—Aug. 


6. Gilia pumila Nutt. Low Gilia. Fig. 3477. 


Gilia pumila Nutt. Journ. Acad. Nat. Sci, Phila. (1I.) 
T0566, 1648. 


Annual, branched from the base and sometimes 
also above, woolly at least when young, 3’-8’ high. 
Leaves alternate, thick, 3’-1’ long, pinnately di- 
vided into linear mucronulate sometimes lobed 
segments, or the uppermost entire; flowers in 
dense or at length looser simple or compound 
cymes, sessile; corolla 3’—4” long, its tube about 
3 times the length of the lobes and twice as long 
as the calyx; calyx-lobes awn-like; stamens in- 
serted in or below the sinuses of the corolla, 
somewhat exserted; ovules 5 or 6 in each cavity. 


In dry soil, western Nebraska to Texas, Idaho, 
Nevada and New Mexico. April-June. 


Gilia tricolor Benth., of California, admitted into 
our first edition as recorded escaped frem gardens to 
roadsides at Lincoln, Nebraska, is not known to have 
become established within our area, 


62 POLEMONIACEAE. Vor. III. 


7. Gilia acerdsa (A. Gray) Britton. Needle-leaved 
Gilia. Fig. 3478. 


G, rigidula var. acerosa A. Gray, Proc. Am. Acad. 8: 280. 1870, 


Gilia acerosa Britton, Man, 761. 1901. 


Perennial, from a woody base, 4-7’ high, glandular-pubes- 
cent, bushy-branched, the slender branches erect-ascending. 
Leaves 1’ long or less, pinnately parted into 3-7 acicular en- 
tire segments; flowers scattered, on slender pedicels 1’ long 
or less; corolla rotate, about 10” wide, its lobes rounded at 
the apex, abruptly contracted at the base; filaments filiform; 
anthers oblong; capsule about as long as the calyx-tube. 


Rocky and gravelly soil, Kansas to Texas, Mexico and New 
Mexico. April-Aug. 


4. LEPTODACTYLON H. & A. Bot. Beech. Voy. 369. 1841. 


Low perennial herbs, somewhat woody, with alternate deeply pinnatifid or palmatifid 
leaves, their segments subulate and spinescent, the rather large flowers terminal, clustered or 
solitary. Calyx 4-5-lobed, the lobes spinescent, the sinuses not scarious but membranous. 
Corolla funnelform, 4-5-lobed, the lobes somewhat spreading. Stamens short. Capsule at 
length distending the calyx-tube, many-seeded. [Greek, referring to the deeply divided 
leaves. | 


Three or four species, natives of western North America. Type species: Leptodactylon cali- 
fornicum H. & A. 


1. Leptodactylon caespitésum Nutt. Tufted Sharp-leaved Gilia. Fig. 3479. 


L. caespitosum Nutt. Journ, Acad. Phila, I].1:157. 1847. 


Gilia pungens caespitosa A, Gray, Proc. Am. Acad. 8: 
268. 1870. 


Gilia caespitosa A. Nelson, Bull. Torr. Club 25: 546. 
1898. Not A. Gray. 


Much branched and tufted from a thick buried 
woody base, 3-5’ high. Leaves densely fascicledS 
and imbricated, 3-5-parted, 4” long or less, the seg- 
ments subulate, spinulose-tipped, glabrous, or the 
margins ciliate; bases of the old leaves persistent; 
calyx about half as long as the corolla-tube, 4-lobed, 
the lobes subulate; corolla white or yellowish, sal- 
verform, the tube about 6” long, the limb 4-lobed; 
stamens 4. 


2 Dry bluffs, western Nebraska and Wyoming. June-— 
ept. 


Leptodactylon pingens Nutt. [Gilia piingens (Torr.) Benth.], of western North America, illus- 
trated in our first edition, where the above species was regarded as a variety of it, is not known 
to occur within our area. 


GENUS 5. PHLOX FAMILY. 63 


5. POLEMONIUM [Tourn.] L. Sp. Pl. 162. 1753. 


Perennial or rarely annual herbs, with alternate pinnate membranous leaves, and mostly 
large cymose-paniculate or thyrsoid flowers. Calyx herbaceous, not angled nor ribbed, cam- 
panulate, 5-cleft to about the middle, accrescent in fruit, the segments lanceolate or ovate, 
entire, erect or connivent over the capsule. Corolla tubular-campanulate or funnelform, 
rarely rotate, blue, white or yellow, the limb 5-lobed. Stamens about equally inserted near 
the base of the corolla, declined, the filaments slender, often pilose at the base. Ovary ovoid; 
ovules few or several in each cavity. Capsule ovoid, obtuse, 3-valved. Seeds wingless, or 
narrowly winged, mucilaginous and emitting spiral threads when wetted. [Name not 
explained.] 

About 15 species, natives of the cooler parts of the north temperate zone. Besides the follow- 


ing, some 10 others occur in the western parts of North America. Type species: Polemonium 
coeruleum L. 


Anthers exserted ; flowers 8’-10” broad; stem erect. 1. P. Van Bruntiae. 
Anthers included ; flowers 5”—6” broad ; stem reclining. 2. P.reptans, 


1. Polemonium Van Brintiae Britton. WZ 
American Jacob’s Ladder. Fig. 3480. 7 \NV 
Polemonium coeruleum A, Gray, Man. Ed. 4, App. 1863. 

NotI. 1753. 

Polemonium Van Bruntiae Britton, Bull. Torr. Club 19: 

2240 pl. 3t. 1892, 

Rootstock stout, horizontal, clothed with fibrous 
roots. Stems erect, glabrous below, somewhat 
glandular-pubescent above, 13°-23° high, leafy to 
the top; leaflets of the lower leaves short-stalked 
or sessile, ovate or lanceolate, acute, 4-134’ long, 
those of the upper fewer, the uppermost leaves 
3-5-foliolate or simple; cymose clusters panicled or 
solitary, rather loosely 3-5-flowered; pedicels 2’-4” 
long; flowers bluish-purple, 8-10” broad; corolla- 
lobes rounded; calyx 5-lobed to about the middle, 
much enlarged in fruit, the lobes acute; stamens 
exserted; ovules 3 or 4 in each cavity; capsule sev- 
eral-seeded. 


In swamps and along streams, Vermont and northern 
New York to Maryland. Differs from the Old World 
P, coeruleum L. in its stout rootstocks, more leafy stem, 
exserted stamens, and rounded corolla-lobes. May-— 


July. 


2. Polemonium réptans L. Greek Valerian. 
Blue-bell. Fig. 3481. 
Polemonium reptans L. Syst. Ed. 10, no. 1. 1759. 


Glabrous or very nearly so throughout, usually 
not more than 1° high; stems weak, slender, at 
length reclining or diffuse, the rootstock short. 
Leaflets oblong, ovate-oblong, or lanceolate-oblong, 
}’-1’ long, the uppermost leaves 3-5-foliolate or 
simple; flowers blue, 5-8” broad; calyx 5-lobed, its 
lobes obtuse or acute; stamens not exserted; ovules 
3 or 4 in each cavity; seeds about 3 in each capsule. 

In woods, New York to Minnesota, south to Georgia 
and Kansas. Ascends to 2200 ft. in Virginia. April- 
May. American abscess-root. Sweat-root. American- 
or creeping great-valerian. 


6. COLLOMIA Nutt. Gen. 1: 126. 1818. 


Annual or rarely perennial herbs, with alternate mostly entire leaves, and purple white 
or reddish capitate or cymose flowers. Calyx obpyramidal or cup-shaped, 5-cleft, scarious in 
the sinuses, accrescent in fruit, not distended by nor ruptured by the ripening capsule, its 


64 POLEMONIACEAE. Vo. III. 


lobes lanceolate or triangular, entire, erect, the sinuses often at length enlarged into a revo- 
lute lobe. Corolla tubular-funnelform or salverform, the limb 5-lobed, spreading, the lobes 
obtuse. Stamens unequally inserted on the tube of the corolla, mostly straight, the filaments 
unequal. Ovules 1 or few in each cavity. Capsule oval to obovoid. Seeds of most species 
mucilaginous and emitting spiral threads when wetted. [Greek, gluten, referring to the 
glutinous seeds when wetted. ] 


About 15 species, natives of western America. Besides the following typical one, some 10 
others occur in the western United States and British Columbia. 


1. Collomia linearis Nutt. Narrow-leaved 
Collomia. Fig. 3482. 


Collomia linearis Nutt. Gen, 1: 126. 1818. 
Gilia linearis A, Gray, Proc. Am. Acad, 17: 223. 1882. 

Annual, viscid-puberulent; stem erect, leafy, sim- 
ple or branched, slender, 3-18’ high. Leaves linear- 
oblong, lanceolate or linear-lanceolate, entire, acumi- 
nate at the apex, narrowed at the base, sessile, or 
the lower short-petioled, 1’-23’ long, 14”-6” wide; 
flowers 5’-7” long, numerous in terminal capitate 
leafy-bracted clusters; calyx-lobes triangular-lanceo- 
late, acute; corolla light purple or nearly white, the 
tube very slender, longer than the calyx, the lobes 
1-2” long; capsule at maturity about as long as 
the calyx. 

In dry soil, Manitoba to Minnesota and Nebraska, 


west to British Columbia, Arizona and California, Also 
in Quebec and New Brunswick. May—Aug. 


7, NAVARRETIA R. & P. Fl. Per. 2: 8. 1790. 


Annual glabrous or viscid-pubescent herbs, with alternate spinose-pinnatifid leaves, or 
the lowest entire, and numerous small flowers in dense terminal bracted clusters. Calyx 
prismatic or obpyramidal, the tube 5-angled, 5-cleft, not accrescent in fruit, not distended by 
nor ruptured by the ripening capsule, the sinuses scarious, the lobes mostly unequal, erect or 
spreading, spiny-tipped, entire, or often toothed. Corolla tubular-funnelform or salverform, 
5-lobed, the lobes oval or oblong. Stamens straight or declined, equally inserted in or below 
the throat of the corolla. Ovary 2-3-celled; ovules solitary, few or several in each cavity. 
Capsule 1-3-celled, dehiscent or indehiscent. Seeds mostly mucilaginous and emitting spiral 
threads when wetted. [In honor of Navarrete, a Spanish physician.] 


About 24 species, natives of western America. Besides the following, some 20 others occur in 
the western United States. Type species: Navarretia involucrata R. & P 


1. Navarretia minima Nutt. Small Navar- 
retia. Fig. 3483. 


Navarretia minima Nutt. Journ. Acad. Nat. Sci. Phila. 
(II.) 1: 160, 1848. 


MISS 
AS 


a 


Gilia minima A, Gray, Proc. Am, Acad. 8: 269. 1870. 


Depressed, tufted, somewhat pubescent; stem usu- 
ally branched, 13’ high. Leaves sessile, 4’-1’ long, 
I-2-pinnatifid into almost filiform rigid acicular seg- 
ments; flowers about 2” long, white, densely capi- 
tate; calyx-lobes awl-shaped, mostly toothed, about 
as long as the tube and equalling the corolla, the 
sinuses more or less white-pubescent; calyx-tube 
about equalling the indehiscent 1-6-seeded capsule. 


In dry soil, Nebraska and South Dakota to Washing- 
ton and Arizona. Summer. 


GENUS I. WATER-LEAF FAMILY. 65 
Family 23. HYDROPHYLLACEAE Lindl. Nat. Syst. Ed. 2, 271. 1836. 


WATER-LEAF FAMILY. 

Herbs, mostly hirsute, pubescent or scabrous, with alternate or basal, rarely 
opposite leaves, and perfect regular 5-parted flowers, in scorpioid cymes, spikes 
or racemes, or rarely solitary. Calyx inferior, deeply cleft or divided, the sinuses 
sometimes appendaged. Corolla gamopetalous, funnelform, salverform, campan- 
ulate, or rotate. Stamens 5, inserted on the tube or base of the corolla, and 
alternate with its lobes; filaments filiform; anthers ovate, oblong, or linear, mostly 
versatile, 2-celled, the sacs longitudinally dehiscent. Diskannularornone. Ovary 
superior, 2-celled, or 1-celled with 2 placentae; styles 2, separate, or partly united; 
stigmas small, terminal; ovules few or numerous, anatropous or amphitropous. 
Capsule 1—2-celled, mostly loculicidally 2-valved, rarely septicidally or irregularly 
dehiscent. Seeds oblong, globose, or angular, usually pitted, rugose or reticu- 
lated; endosperm fleshy or cartilaginous; embryo small; cotyledons half-terete 
or plano-convex. 

About 17 genera and 175 species, mostly natives of western North America, 

Styles united below; ovary 1-celled; leaves mostly lobed or dentate. 

Corolla-lobes convolute (rarely imbricated) in the bud; placentae dilated. 


Stamens exserted. 1. Hydrophyllum. 
Stamens not exserted. 
Sinuses of the calyx appendaged. 2. Nemophila, 
Calyx much enlarged in fruit, its sinuses not appendaged. 3. Nyctelea. 
Corolla-lobes imbricated in the bud; placentae narrow. 4. Phacelia, 
Styles distinct to the base; ovary 2-celled; leaves entire. 5. Nama, 


1. HYDROPHYLLUM [Tourn.] L. Sp. Pl. 146. 1753. 


Perennial or biennial herbs, with large lobed pinnatifid or p:nnately divided leaves, and 
rather large, white blue or purple flowers, in terminal or lateral peduncled more or less 
scorpioid cymes. Calyx deeply 5-parted, the segments lanceolate or subulate, the sinuses 
naked or appendaged. Corolla tubular-campanulate or campanulate, 5-lobed, the lobes con- 
volute in the bud, each with a linear appendage within, which extends to the base of the 
corolla and is incurved into a groove. Stamens 5, exserted; filaments pilose below or at the 
base; anthers linear or oblong, versatile. Ovary 1-celled, hispid-pubescent; placentae fleshy, 
dilated so as to nearly fill the cavity, free from the ovary-wall except at the top and bottom, 
each enclosing 2 ovules; styles united nearly to the summit. Capsule 2-valved. Seeds 1-4, 
globose-obovoid. [Greek, water-leaf, referring to the supposed cavity for water in each leaf.] 


About 12 species, natives of North America. Type species: Hydrophyllum virginianum L. 
Leaves, at least the lower, pinnatifid or pinnately divided. 


Calyx not appendaged in the sinuses or scarcely so, 


Plant sparingly pubescent ; leaf-segments acute, 1. H. virginianum. 
Plant villous-hirsute ; leaf-segments blunt. 2. H. macrophyllum. 
Calyx with a reflexed appendage in each sinus. 3. H. appendiculatum. 


Leaves palmately 5—9-lobed. ; 4. H. canadense. 


1. Hydrophyllum virginianum L. Vir- 
ginia Water-leaf. Fig. 3484. 
H, virginianum L. Sp. Pl. 146. 17536 


Perennial by scaly rootstocks; stems slender, 
glabrous or nearly so, simple or sparingly 
branched, ascending or erect, rather weak, 
1°-3° long. Lower and basal leaves long- 
petioled, 6-10’ long, pinnately divided into 5-7 
oblong ovate or ovate-lanceolate, acute or 
acutish, sharply toothed or incised segments 
1-2’ long, glabrous or with few scattered hairs ; 
upper leaves similar, short-petioled, smaller, 
with fewer segments; cymes slender-pedun- 
cled, simple or forked, dense or at length 
open; flowers white or violet, purple, darkest 
at high altitudes, short-pedicelled; pedicels 
strigose-pubescent; calyx-segments narrowly 
linear, hispid, spreading, the sinuses not ap- 
pendaged; corolla about 4” long, its segments 
erect; capsule globose, nearly 2” in diameter. 

In woods, Quebec to Ontario, South Dakota, 
South Carolina and Kansas. Ascends to 5000 ft. 
in North Carolina. Brook-flower. May-—Aug. 


5 


66 HY DROPHYLLACEAE. Vor. III. 


H. patens Britton, of Minnesota, differs in having the calyx-segments appressed to the corolla, 
the corolla-segments with spreading tips. 
F ate te ie “7 pe 2. Hydrophyllum macrophyllum Nutt. 
Pe My Large-leaved Water-leaf. Fig. 3485. 
H. macrophyllum Nutt. Journ. Phila. Acad. 7: 111. 

1834. 

Perennial by scaly rootstocks, villous-hirsute 
all over; stem rather stout, 2°-3° high. Lower 
leaves long-petioled, 8-12’ long, deeply pinnatifid 
or pinnately divided into 7-13 oval or ovate obtuse 
coarsely dentate segments 1-3’ long; upper leaves 
similar, smaller, shorter-petioled and with fewer 
segments; cymes long-peduncled, simple or fork- 
ed, very dense; peduncles stout; pedicels short; 
flowers nearly white; calyx cleft to below the 
middle, its lobes lanceolate, erect, white-hispid, 
the sinuses not appendaged; corolla 5-6” long; 
anthers oblong; capsule globose, densely white- 
hispid, 13” in diameter, enclosed by the calyx. 


In rich woods, Virginia to Ohio, Illinois, Alabama 
and Tennessee, Ascends to 4ooo ft. in Virginia. 


ieee paar: 
sre 


Panna TERY 


Paat 
\\? SV 7 
3. Hydrephyllum appendiculatum Michx. Nv S < 
Appendaged Water-leaf. Fig. 3486. a NG 
H. appendiculatum Michx. Fl. Bor. Am, 1: 134. 1803. : 


Biennial, rough-hairy all over; stem slender, simple 
or usually branched, weak, 1°-2° long, somewhat 
viscid above. Lower and basal leaves long-petioled, 
pinnatifid or pinnately divided into 5-7 ovate or oval, 
acute or obtusish, irregularly dentate or incised, mem- 
branous segments; upper leaves smaller and shorter- 
petioled, ovate to orbicular, variously lobed, the lobes 
acute and dentate; cymes loose, branched, the slender 
pedicels 4-10” long; calyx parted to near the base, 
enlarging in fruit, the segments triangular-lanceolate, 
acuminate, spreading, with a short reflexed appendage 
in each sinus; corolla violet or purple, 6-7” long; 
stamens little exserted; capsule about 14” in diameter. 

In woods, Ontario to New York, North Carolina, west to 
Minnesota and Kansas. May—June. 


4. Hydrophyllum canadénse L. Broad- 
leaved Water-leaf. Fig. 3487. 
H. canadense L. Syst. Ed. 10, 919. 1759. 

Perennial by scaly rootstocks; stems rather 
slender, commonly simple, glabrous or nearly so, 
1°-23° high. Leaves nearly orbicular, cordate, 
sparingly pubescent, at least above, palmately 
s-o-lobed, the lower long-petioled, often 12’ 
broad, occasionally with 1 or 2 pairs of small 
segments on the petiole; upper leaves smaller, 
but usually 4’-7’ broad; lobes ovate, acuminate, 
dentate; peduncles shorter than the leaves; cymes 
dense or becoming loose, simple or forked; pedi- 
cels short, nearly glabrous; calyx cleft nearly to 
the base, its segments linear-lanceolate, acute, 
nearly or quite glabrous, sometimes with a minute 
tooth in each sinus; corolla campanulate-rotate, 
white to purplish anthers linear-oblong; capsule 
2” in diameter. 

In woods, Vermont to New York, North Carolina, 
Ontario, Illinois and Kentucky. Ascends to 4000 ft. in Virginia. June—Aug. : 


GENUS 2. WATER-LEAF FAMILY. 67 


2. NEMOPHILA Nutt. Jeurn.»Phil, Acad3+/779, (1822. 


Annual diffuse pubescent slender and fragile herbs, with alternate or opposite mostly 
pinnatifid or lobed leaves. Flowers white, blue or variegated, solitary, peduncled, lateral or 
terminal. Calyx deeply 5-cleft or 5-parted, with a reflexed or spreading appendage in each 
sinus. Corolla campanulate or rotate-campanulate, mostly longer than the calyx, usually 
with 1o small appendages within at the base, the lobes convolute inthe bud. Stamens included; 
anthers ovate or oblong. Ovary tI-celled with placentae similar to those of Hydrophyllum; 
styles partly united; ovules 2-12 on each placenta. Capsule 2-valved. Seeds 1-4. [Greek, 
grove-loving. ] 

About 10 species, natives of North America, mostly Californian. Type species: Nemophila 
phacelioides Nutt. 


1. Nemophila micrécalyx (Nutt.) F. & M. 
Small-flowered Nemophila. Fig. 3488. 
Ellisia microcalyx Nutt. Trans. Am. Phil. Soc, (II.) 5: 

19I. 1833-37. 
Nemophila microcalyx F.& M. Sert. Petrop. pi. 6. 1846. 

Stems very slender, diffuse, branched, 2’-15’ long. 
Leaves membranous, petioled, 1-23’ long, pinnatifid 
or pinnately divided into 3-5 obovate cuneate or 
oblique, obtuse 2-3-dentate or -lobed, approximate or 
confluent segments, the upper all alternate, the lowest 
opposite; peduncles slender, 4’—12” long, opposite 
the leaves, shorter than or equalling the petioles; 
flowers white or blue, 13-2” long; appendages in 
the sinuses of the calyx minute; calyx scarcely en- 
larged in fruit; ovules 2 on each placenta; corolla- 
appendages obsolete or none; anthers oval; capsule 


about 13” in diameter, much longer than the calyx, conf} Y 
1-2-seeded. Be ; 
In woods, Virginia to Florida, west to Arkansas and 
Texas. April-June. : : 
~ 4 YH 
3. NYCTELEA Scop. Introd. 183. 1777. 3 Pa = 


[MacrocaLyx Trew, Nov. Act. Nat. Cur. 2: 330-332. pl. 7. f. 7. Hyponym. 1761.] 
[Errisia L. Sp. Pl. Ed. 2, 1662. 1763. Not Syst. Ed. 10, 1121. 1750.] 

Annual hirsute or pubescent branching slender herbs, with opposite or alternate, pin- 
nately divided or 1-3-pinnatifid leaves, and solitary or racemose small white or bluish flowers. 
Calyx 5-lobed or 5-parted, spreading, much enlarged in fruit, destitute of appendages in the 
sinuses. Corolla campanulate or nearly cylindric, shorter than or slightly exceeding the calyx, 
usually with 5 minute appendages on the tube within, its lobes convolute or imbricated in’the 
bud. Stamens included; anthers oval or oblong. Ovary 1-celled; styles united below; ovules 
2-4 on each of the placentae, which are similar to those of the two preceding genera. [Name 
unexplained. ] 


About 3 species, natives of North America, the fol- 
lowing typical. 


1. Nyctelea Nyctélea (L.) Britton. 
Nyctelea. Fig. 34809. 


Ipomoea Nyctelea L. Sp. Pl. 160. 1753. 

Polemonium (?) Nyctelea L. Sp. Pl. Ed. 2,231. 1762. 

Ellisia Nyctelea L, Sp. Pl. Ed. 2, 1662. 1763. 

Macrocalyx Nyctelea Kuntze, Rev. Gen. Pl. 434. 
1891. 


Sparingly hirsute-pubescent; stem several times 
forked, 4-12’ high. Leaves pinnately divided, 
petioled, 2’-4’ long, ovate-oblong in outline, the 
upper alternate, the lower opposite, the segments 
oblong or lanceolate, dentate, entire or- lobed; 
peduncles slender, 1-flowered, opposite the leaves; 
calyx in flower about 2” long, about equalling 
the corolla, enlarging, widely spreading and be- 
coming 8-14” broad in fruit, its lobes lanceolate 
or ovate-lanceolate, acuminate; fruit at length 
pendulous; capsule globose, 2”-3” in diameter. 


In moist soil, New Jersey to Minnesota, Saskatche- 
wan, Virginia, Nebraska and Kansas. April—July. 


68 HYDROPHYLLACEAE. Vou.1i: 


4. PHACELIA Juss.; J. F. Gmel. Syst. 330. 1791. 


Annual, biennial or perennial, mostly hirsute hispid or scabrous herbs, with alternate entire 
dentate lobed pinnatifid or dissected leaves, the lowest rarely opposite, and blue purple violet 
or white flowers in terminal scorpioid cymes or racemes. Calyx 5-lobed, somewhat enlarging 
in fruit; sinuses not appendaged. Corolla campanulate, nearly rotate, tubular or funnelform 
the tube sometimes appendaged within, opposite the lobes, the lobes imbricated in the buds 
Stamens exserted or included, attached near the base of the corolla; anthers mostly ovate. 
Ovary t-celled, the 2 placentae narrow, affixed to the walls; styles united below; ovules 2 or 
several on each placenta. Capsule 1-celled, or falsely nearly 2-celled by the intrusion of the 
placentae, 2-valved. Seeds usually reticulated. [Greek, a cluster, referring to the clustered 
flowers of some species.] 

About 90 species, natives of the New World. Besides the following, some 60 others occur in 
the western parts of North America. Type species: Phacelia secunda J. F. Gmel. 


Corolla-lobes entire. 
Corolla manifestly appendaged within, between the stamens. 


Leaves entire. 1. P.leucophyilla. 
Leaves crenate-dentate. : 2. P.integrifolia. 
Leaves pinnately divided, or pinnatifid, the segments incised. 
Racemes loose ; pedicels slender; ovules 2 on each placenta, 3. P. bipinnatifida. 
Racemes dense; pedicels short ; ovules numerous. 4. P. Franklinii, 


Appendages of the corolla inconspicuous or none, 
Filaments pubescent ; calyx-lobes oblong. 


Puberulent ; flowers 4”-5” broad. 5. P. dubia. 
Hirsute ; flowers 6”—7” broad. 6. P. hirsuta. 
Filaments glabrous ; calyx-lobes linear. 7. P. Covillei. 
Corolla nearly rotate, its lobes fimbriate. 
Lobes of the leaves and calyx acute. 8. P. Purshii. 
Lobes of the leaves and calyx obtuse. g. P. fimbriata. 


1. Phacelia leucophylla Torr. Silky Phacelia. Fig. 3490. 


Phacelia leucophylla Torr. Frem, Rep. 93- 1845. 


Perennial by a stout rootstock, pale, densely silky- 
pubescent, the hairs appressed or ascending. Stem 
simple or branched, 1°-13° high; leaves lanceolate to 
oblong, entire, pinnately veined, 2’-4' long, 47-12” 
wide, the lower long-petioled, the upper sessile or 
nearly so; spike-like branches of the scorpioid cymes 
very dense, nearly straight and 1-3’ long when ex- 
panded; flowers sessile, very numerous, about 4” high; 
calyx-lobes hispid, oblong-lanceolate or linear, some- 
what shorter than the white or bluish, 5-lobed corolla ; 
corolla-appendages conspicuous, in pairs between the 
filaments; filaments exserted, glabrous; ovules 2 on 
each placenta; capsule ovoid. 

In dry soil, South Dakota to Idaho, British Columbia, 
Nebraska and Colorado. May-Aug. The species has 
been taken for P. heterophylla Pursh, of the far west, 
which has spreading brown hairs, some of the leaves usu- 
ally pinnatifid, and pilose filaments. 


s VY = N— 
2. Phacelia integrifolia Torr. Crenate- 
leaved Phacelia. Fig. 3491. 
P. integrifolia Torr. Ann, Lyc. N. Y. 2: 222. pl. 3. 1827. 


Annual or biennial; stem erect or ascending, rather 
stout, very leafy, commonly branched above, viscid- 
hirsute, 62° high. Leaves finely strigose-pubescent, 
ovate-oblong or oblong-lanceolate, irregularly cre- 
nate-dentate, obtuse at the apex, rounded or cordate 
at the base, 1’-2}’ long, petioled or the uppermost 
sessile; spike-like branches of the scorpioid cymes 
dense, 2’-4’ long when expanded; flowers sessile, 
about 4” long; calyx-segments oblong, acute; corolla 
fubular-campanulate, white or blue, its tube longer 
than the calyx; filaments glabrous, exserted; ovules 
2 on each placenta; capsule ovoid, obtuse. 


In saline soil, western Kansas (according to BB: 
Smyth) ; Oklahoma to Colorado, Mexico, Utah and Ari- 
zona. April—Sept. 


GENUS 4. WATER-LEAF FAMILY. 69 


3. Phacelia bipinnatifida Michx. Loose- 


flowered Phacelia. Fig. 3492. | WO y QA 


Phacelia bipinnatifida Michx. Fl. Bor. Am. 1: 134. pl. 16. 
1803. 


ee 


Biennial, hirsute-pubescent; stem erect, usually 
much branched, glandular-viscid above, 1°-2° high. 
Leaves slender-petioled, 2’-5’ long, pinnately divided 
or deeply pinnatifid into 3-7 ovate or oblong acute 
or acutish, dentate or incised segments, or these 
again pinnatifid; flowers blue or violet, 6’-8” broad, 
numerous, slender-pedicelled in loose racemes, the 
inflorescence only slightly scorpioid; pedicels 4’—10” 
long, recurved in fruit; calyx-segments linear; ap- 
pendages of the rotate-campanulate corolla in pairs 
between the stamens, conspicuous, villous on the 
margins, corolla-lobes entire; filaments pilose, ex- 
serted; ovules 2 on each placenta; capsule globose. 

In moist thickets and along streams, Ohio to Illinois, 


Missouri, Georgia, Alabama and Tennessee. Ascends 
to 4000 ft. in North Carolina. April-June. 


4. Phacelia Franklinii (R. Br.) A. Gray. 
Franklin’s Phacelia. Fig. 3493. 
Eutoca Franklinii R. Br. App. Frank. Journ. 51. pl. 27. 


1823. 
Phac tia Franklinii A. Gray, Man. Ed. 2, 329. 1856. 

Annual, villous-pubescent; stem erect, 6-18 
high, simple, or corymbosely branched at the sum- 
mit. Leaves 147-3’ long, pinnately parted into 
7-15 linear or linear-oblong acute entire dentate 
or incised segments; flowers blue or nearly white, 
short-pedicelled in dense scorpioid racemes; calyx- 
segments linear-lanceolate, acute; longer than the 
tube of tne rotate-campanulate corolla; append- 
ages of the corolla free at the apex: anthers 
scarcely exserted; filaments glabrous, or nearly 
so; styles united nearly to the summit; ovules 
numerous on each placenta; capsule ovoid, acute. 


Western Ontaric, Michigan and Minnesota to Brit- 
ish Columbia, Wyoming and Idaho. Summer, 


5. Phacelia dubia (L.) Small. Small-flowered Phacelia. Fig. 34 


Polemonium dubium L. Sp. Pl. 163. 1753. 
Phacelia parviflora Pursh, Fl. Am. Sept. 140. 1814. a AN) 
Phacelia dubia Small, Bull. Torr. Club 21: 303. 1804. Oe 


O4. 


Annual, puberulent or glabrate, branched from the 
base, the branches very slender, erect or ascending, 
5-12’ high. Lower and basal leaves petioled, 1-2” 
long, pinnatifid or pinnately divided into 3-5 oblong 
obtuse entire or dentate segments, or rarely merely 
’ dentate, or even entire; upper leaves much smaller, 
sessile, less divided; flowers light blue or white, 
racemose, 4”=5” broad; racemes 5=15-flowered, elon- 
gated in fruit; pedicels 3”’-7” long; calyx-lobes 
oblong or oblong-lanceolate; corolla rotate-campanu- 
late, the appendages obsolete; filaments pubescent ; 
anthers slightly exserted; ovules 4-8 on each pla- 
centa; capsule globose, 13” in diameter, 6-12-seeded ; 
fruiting pedicels ascending. 

In moist soil, New York and Pennsylvania to Georgia, 
Missouri, Kansas and Texas. Ascends to 2000 ft, in 
Virginia. April-June. 


7o HYDROPHYLLACEAE. Vor. III. 


an 6. Phacelia hirsuta Nutt. Hairy Phacelia. 
Os AN le: : 

YS 2D Fig. 3495. 

MFRS Phacelia. hirsuta Nutt. Trans, Am. Phil, Soe, (1) 

5: 191. 1834-37. 

Phacelia parviflora var. hirsuta A, Gray, Proc. Am, 
ead. 1O-s2TeeeTe75. 

Similar to the preceding species but usually 
stouter and larger, hirsute-pubescent. Leaves 
petioled, pinnatifid or deeply pinnately divided 
into 5-9 oblong obtuse usually entire segments, 
or the upper 3-5-lobed or entire, sessile; racemes 
rather dense, scorpioid when unfolding; flowers 
blue, 6-7” broad; calyx-segments oblong-lanceo- 
late, hirsute; corolla rotate-campanulate, its ap- 
pendages short; ovules 3-4 on each placenta; 
stamens scarcely exserted; capsule globose, 4-8- 
seeded. 


In dry soil, Virginia to Georgia, west to Missouri, 
Kansas and Texas. April-June. 


7. Phacelia Covillei S. Wats. Coville’s 
Phacelia. Fig. 3496. 
Phacelia Covillei S, Wats. in A. Gray, Man, Ed. 

6, 360. 1890. 

Annual, similar to the two preceding species, 
branched from the base, pubescent; branches 
very slender, weak, 6-12’ long. Leaves deeply 
pinnatifid or pinnately divided into 3-7 oblong 
or obovate, obtuse segments; racemes only 
1-5-flowered; pedicels filiform, 6-8” long; 
calyx-segments linear, elongating in fruit; 
corolla tubular-campanulate, about 3” long and 
broad when expanded; filaments glabrous; 


anthers not exserted; appendages of the co- \ 
M, 


rolla obsolete; capsule globose, 13”-2” in diam- 
eter; fruiting pedicels recurved. 


Along the Potomac River above Washington, vA 
D. C., and in Illinois. April-May. 


8. Phacelia Parshii Buckl. Pursh’s Phacelia. Vig. 3497. 


Phacelia Purshii Buckl. Am. Journ. Sci. 45: 171. 
1843. 


Annual, pubescent; stem erect, usually much 
branched, 6-18’ high. Lower and basal leaves 
petioled, 13-3’ long, pinnately parted or pin- 
natifid into 9-15 oblong or lanceolate, acute or 
obtusish, entire or sometimes incised segments ; 
upper leaves sessile, with fewer segments; ra- 
cemes strongly 1-sided, 10~-20-flowered, much 
elongated in fruit; pedicels 3’-12” long; calyx- 
segments lanceolate or linear-lanceolate; corolla 
5-6” broad, blue or white, nearly rotate, not 
appendaged within, its lobes fimbriate; filaments 
slightly exceeding the corolla; ovules 2 on each 
placenta; capsule globose-ovoid. 


In moist woods or thickets, Pennsylvania to Min- 
nesota, south to North Carolina, Alabama and Mis- 
souri. Ascends to 2000 ft. in Virginia. April-June. 


GENUS 4. WATER-LEAF FAMILY. 73 


-g. Phacelia fimbriata Michx. Fringed or 
. Mountain Phacelia. Fig. 3498. 


Phacelia fimbriata Michx. Fl. Bor. Am. 1: 134. 1803. 


Annual, sparingly pubescent; stems simple or 
branched, ascending or diffuse, 6’-20’ long. Basal 
and lower leaves slender-petioled, 2’-4’ long, pin- 
nately divided into 5-9 orbicular obovate or oblong 
obtuse dentate or entire segments; upper leaves 
sessile, their segments oblong or lanceolate, acute; 
clusters loose, 3-12-flowered; pedicels 4”-15” long; 
flowers 4’—-5” broad, white; calyx-segments linear to 
spatulate, obtuse; corolla rotate-campanulate, not 
appendaged within, its lobes strongly fimbriate; fila- 
ments pilose, about equalling the corolla; ovules 2 
- on each placenta; capsule depressed-globose, 2” in 
diameter. 


In woods, mountains of Virginia to Alabama. May-— 
June. 


XN 
5. NAMA-E. Sp: PI 226." “17353: 
[Hyprotea L. Sp. Pl. Ed. 2, 328. 1762.] 

Perennial herbs, some tropical species shrubby, with alternate entire leaves, sometimes 
with spines in their axils, and blue cymose-clustered or racemose flowers. Calyx-segments 
distinct to the base, ovate or lanceolate. Corolla rotate-campanulate, not appendaged within, 
5-cleft, the lobes imbricated in the bud. Stamens inserted on the base of the corolla; fila- 
ments filiform, dilated below; anthers sagittate. Ovary 2-celled (rarely 3-celled) ; ovules 
numerous in each cavity, on fleshy adherent placentae; styles 2, rarely 3, slender or filiform, 
distinct to the base; stigma capitellate. Capsule globose or ovoid, septicidally or irregularly 
dehiscent. 


About 15 species, natives of warm and tropical regoins of both the Old World and the New. 
Besides the following, another occurs in the southern United States. Type species: Nama 
zeylanica L. 


Leaves lanceolate; flowers mostly in axillary clusters. 
Glabrous, or very nearly so, throughout ; sepals ovate to ovate-lanceolate. 1. N. affinis. 
Villous, at least above, and on the calyx; sepals lanceolate. 2, N. quadrivalvis. 
Leaves ovate; flowers mostly in terminal clusters ; sepals villous. 3. N. ovata. 


1. Nama affinis (A. Gray) Kuntze. 
Smooth Nama. Fig. 3499. 


Hydrolea affinis A, Gray, Man. Ed. 5, 370. 1867. 
Nama affinis Kuntze, Rev. Gen, Pl. 435. 1891. 


Glabrous or very nearly so throughout, 
with or without slender spines in the axils; 
stems ascending, 1°-24° high. Leaves ob- 
long-lanceolate, petioled, acute or acumi- 
nate at both ends, 2-5’ long, 4’-8” wide; 
flowers 6-7” broad, short-pedicelled, in 
rather dense peduncled leafy-bracted axil- 
lary clusters; sepals ovate or ovate-lanceo- 
late, acute or acuminate, about equalling the 
corolla; capsule 2” in diameter when ma- 
ture, somewhat longer than the styles, 
shorter than the sepals. 


In wet places, southern Illinois to Missouri, 
Louisiana and Texas. June—Aug. 


72 HYDROPHYLLACEAE. Vor. III. 


2. Nama quadrivalvis (Walt.) Kuntze. 
Hairy Nama. Fig. 3500. 


Hydrolea quadrivalvis Walt. Fl. Car. t10. 1788. 
H, caroliniana Michx. Fl. Bor. Am.1: 177. 1803. 
N. quadrivalvis Kuntze, Rev. Gen. Pl. 435. 18091. 


Similar to the preceding species, but pubes- 
cent, at least above, and on the calyx, with 
spreading hairs, usually bearing slender spines 
in the axils; stem ascending, 1°-2° high. Leaves 
lanceolate, acute or acuminate, glabrous, or 
sparingly pubescent, petioled, 2’-5’ long, 3’-8” 
wide; lower petioles 4’ long, or more; flowers 
in axillary clusters; sepals lanceolate or linear- 
lanceolate, acuminate, about as long as the 
corolla; capsule 2”-3” in diameter, longer than 
the styles, about the length of the sepals. 


In_wet soil, southeastern Virginia to Florida 
and Louisiana. June—Aug. 


3. Nama ovata (Nutt.) Britton. Ovate- 
leaved Nama. Fig. 3501. 


Hydrolea ovata Nutt. Trans. Am. Phil. Soc, (II.) 
5: 196, 1833-37. 
N, ovata Britton, Mem. Torr, Club 5: 272. 1894. 
Stem erect or nearly so, 1°-3° high, usually 
branched near the summit, puberulent, or 
somewhat hirsute, at least above, usually spine- 
bearing in most of the axils. Leaves ovate, 
rarely ovate-lanceolate, puberulent or glabrcus, 
short-petioled, or the upper almost sessile, 
1’-23’ long, 2’-12’ wide, acute at the apex, nar- 
rowed or rounded at the base; flowers often 
1’ broad or more, in terminal clusters; 
sepals lanceolate, acuminate, very villous, 
shorter than the corolla, longer than the cap- 
sule; styles longer than the sepals. 


In wet soil, Georgia to Missouri, Louisiana and 
Texas. May-—Sept. 


Family 24. BORAGINACEAE Lindl. Nat. Syst. Ed. 2, 274. 1836. 


BorRAGE FAMILY. 


Annual, biennial or perennial herbs, shrubs, or some tropical species trees. 
Leaves alternate, very rarely opposite or verticillate, exstipulate, mostly entire 
and hispid, pubescent, scabrous or setose. Flowers perfect, usually regular, mostly 
blue, in one-sided scorpioid spikes, racemes, cymes, or sometimes scattered. Calyx 
inferior, mostly 5-lobed, 5-cleft, or 5-parted, usually persistent, its lobes valvate. 
Corolla gamopetalous, mostly regular and 5-lobed, sometimes crested or append- 
aged in the throat, rarely irregular, its lobes imbricated, convolute, plicate or 
induplicate in the bud. Stamens as many as the corolla-lobes and alternate with 
them, inserted on the tube or throat; filaments slender or short; anthers 2-celled, 
the sacs longitudinally dehiscent. Disk annular, entire, or 5-lobed, or none, com- 
monly inconspicuous. Ovary superior, of 2 2-ovuled carpels, entire, or the carpels 
commonly deeply 2-lobed, making it appear as of 4 I-ovuled carpels; style simple, 
entire or 2-cleft in our genera; ovules anatropous or amphitropous. Fruit mostly 
of 4 1-seeded nutlets, or of 2 2-seeded carpels. Endosperm none; embryo straight 
or curved; cotyledons mostly flat or plano-convex; radicle short. 


About 85 genera and 1500 species, of wide geographic distribution. 


GENUS I. BORAGE FAMILY. | 73 


* Ovary entire or 2-4-grooved; style terminal. 1. Heliotropium. 
** Ovary 4-divided or deeply 4-lobed, the style arising from the center. 
Flowers regular. 
Nutlets armed with barbed prickles. 
Nutlets spreading or divergent, covered by the prickles. 2. Cynoglossum. 
Nutlets erect or incurved, the prickles on their backs or margins. 3. Lappula. 
Nutlets unarmed. 
Nutlets attached laterally to the receptacle, sometimes just above their bases. 
* Fruiting calyx not greatly enlarged nor membranous. 
Corolla small, usually white; receptacle conic or elongated. 
Annuals; calyx nearly closed in fruit; inflorescence naked or bracteolate. 


Lowest leaves mostly opposite; calyx persistent. 4. Allocarya. 
Leaves all alternate; calyx at length deciduous. 5. Cryptantha. 
Perennials or biennials; calyx-segments more or less spreading in fruit; inflo- 

rescence leafy. 6. Oreocarya. 


Corolla tubular-funnelform ; receptacle flat or convex. 
Corolla blue, rarely white; nutlets attached just above their bases. 


Maritime; nutlets fleshy, smooth and shining. 7. Pneumaria. 
Not maritime; nutlets wrinkled when mature and dry. 8. Mertensia. 
Corolla yellow; nutlets laterally attached. g. Amsinckia. 
Fruiting calyx much enlarged, membranous, veiny. 10. Asperugo. 
Nutlets attached to the receptacle by their very bases. 
Scar of attachment small, flat. 
Corolla salverform or funnelform, its lobes rounded, spreading. 
Racemes not bracted; corolla-tube short. 11. Myosotis. 
Racemes bracted ; corolla-tube cylindric, usually slender. 12. Lithospermum. 
Corolla tubular, its lobes erect, acute. 13. Onosmodium. 
Scar of attachment large, concave. 
Corolla tubular, 5-toothed. 14. Symphytum. 
Corolla rotate; anthers erect in a cone. 15. Borago. 
Flowers irregular. 
Stamens included ; throat of the corolla closed by scales. 16. Lycopsis. 
Stamens exserted; throat of the corolla dilated, open. 17. Echium. 


1. HELIOTROPIUM [Tourn.] L. Sp. Pl. 130. 1753. 


Herbs or shrubs, with alternate mostly entire and petioled leaves, and small blue or white 
flowers, in scorpioid spikes, or scattered. Calyx-lobes or -segments lanceolate or linear. 
Corolla salverform or funnelform, naked in the throat, its tube cylindric, its lobes imbricated, 
plicate or induplicate in the bud, spreading in flower. Stamens included; filaments short, or 
none. Style terminal, short or slender; stigma conic or annular. Fruit 2-4-lobed, separating 
into 4 I-seeded nutlets, or into 2 2-seeded carpels. Ovary entire, or 2-4-grooved. [Greek, 
sun-turning, 7. e., turning to or with the sun.] 


About 125 species, widely distributed in warm-temperate and tropicai regions. Besides the fol- 
lowing, some ro others occur in the southern and western parts of North America. The species are 
called Turnsole. Type species: Heliotropium europaeum L. 


Fruit 4-lobed, each lobe becoming a 1-seeded nutlet. 
Flowers in scorpioid spikes. 
Plant rough-puberulent ; leaves oval. 1. H. europaeum. 
Glabrous, fleshy ; leaves linear to spatulate. k 
Leaves linear to oblanceolate; corolla 2” broad. 2. H. curassavicum, 
Leaves broadly spatulate; corolla 3”—4” broad. 3. H. spathulatum. 
Flowers solitary, terminating short branches. 4. H. tenellum, 
5- 
6. 


Cryptantha. 
H. indicum. 


1. Heliotropium europaéum L. European 
Heliotrope. Fig. 3502. 
Heliotropium europaeum L. Sp. Pl. 130. 1753. 


Annual, much branched, rough-puberulent, 6-18’ high. 
Leaves oval, 1-2’ long, obtuse at the apex, narrowed at 
the base, slender-petioled, pinnately veined: flowers 
white, 1-2” broad, in dense I-sided scorpioid, bractless 
spikes; terminal spikes in pairs, the lateral ones com- 
monly solitary, becoming 1-3’ long in fruit; calyx- 
segments lanceolate to linear-lanceolate, shorter than 
the corolla-tube; anthers distinct, obtuse; stigma-tip 
long-conic; fruit depressed-globose, pubescent, 4-lobed, 
at length separating into 4 nutlets. 

In waste places, Massachusetts to New York, Pennsyl- 


vania and Florida. Adventive or naturalized from Europe. 
June-Oct. 


BORAGINACEAE. Vor, IIT. 


2. Heliotropium curassavicum L. Sea- 
side Heliotrope. Fig. 3503. 
Heliotropium curassavicum L. Sp. Pl. 130. 1753. 


Annual, fleshy, glabrous throughout, more or 
less glaucous, branched, diffuse, the branches 
6’-18’ long. Leaves linear, or linear-oblong, 
entire, very inconspicuously veined, 17-2’ long, 
13-3” wide, obtuse at the apex, narrowed into 
petioles, or the upper sessile, sometimes with 
smaller ones fascicled in the axils; scorpioid 
spikes densely flowered, bractless, mostly in pairs; 
flowers about 2” broad; calyx-segments lanceo- 
late, acute; corolla white with a yellow eye or 
changing to blue; stigma umbrella-shaped; anthers 
acuminate; fruit globose, at length separating 
into 4 nutlets. 

On sandy seashores, Delaware to Texas and Mex- 
ico. Widely distributed in saline and maritime soil 
in the West Indies, South America and the Old 


World. In ballast about the northern seaports. 
May-Sept. 


3. Heliotropium spathulatum Rydb. Spatulate- 
leaved Heliotrope. Fig. 3504. 


H. spathulatum Rydb. Bull. Torr. Club 30: 262. 1903. 


Perennial, glabrous, fleshy, glaucous, branched, 1°-13° 
high, the branches ascending. Leaves spatulate, indis- 
tinctly veined, 2’ long or less, obtuse at the apex; 
scorpioid spikes 2-5; flowers 3’-4” broad, white, or 
bluish; calyx-segments ovate-lanceolate, acute; fruit 
rather larger than that of H. curassavicum. 


Prairies, plains and meadows, Iowa to North Dakota, 


Assiniboia, Chihuahua and California. 


June-Sept. 


4. Heliotropium tenéllum (Nutt.) Torr. 
Slender Heliotrope. Fig. 3505. 


Lithospermum tenellum Nutt. Trans. Am. Phil. Soc. (II.) 
5: 188.. 1833-37. 
H. tenellum Torr. in Marcy’s Rep. 304. pl. 14. 1853. 
Annual, strigose-canescent; stem erect, slender, 
paniculately branched, commonly leafless below, 
6-18’ high. Leaves linear, entire, 3’-14’ long, 1-2” 
wide, narrowed at both ends, sessile, or the lower 
petioled; flowers white, about 24” long, sessile at the 
ends of short lateral branches, bracted by I or 2 
leaves; calyx-segments unequal, the 2 or 3 larger 
ones about as long as the corolla; corolla-tube canes- 
cent, slightly longer than the limb, its lobes entire; 
anthers obtuse; stigma subulate-tipped; fruit de- 
pressed, 4-lobed, strigose-pubescent, separating into 
4 I-seeded nutlets. 


In dry soil, Kentucky to Kansas, south to Alabama, 
Texas and New Mexico. April—Aug. 


GENUS I. BORAGE FAMILY. i is 


5. Heliotropium convolvulaceum (Nutt.) A. Gray. Bindweed Heliotrope. 
Fig. 3506. 


Euploca convolvulacea Nutt. Trans. Am. Phil. Soc. (II.) 

5: 189. 1833-27. 

Et, Pegivmraceuns A. Gray, Mem. Am. Acad. 6: 403. 

1857. 

Annual, strigose-canescent, usually much branched, 
6-15’ high, the branches ascending. Leaves oblong, 
ovate, or lanceolate, entire, short-petioled, obtuse or 
acute at the apex, narrowed at the base, 3’-13’ long; 
flowers numerous, fragrant, very short-peduncled, 
terminal and lateral, mostly solitary and opposite the 
leaves; calyx-segments lanceolate, acuminate, equal; 
corolla white, strigose, 8’—10” long, about 6” broad, 
its tube narrowed at the throat, longer than the calyx 
' and the angulate-lobed limb; anthers inserted on the 
tube of the corolla, slightly cohering by their tips; 
style filiform; stigma with a tuft of bristly hairs; 
fruit 2-lobed, pubescent, each lobe splitting into 2 
1-seeded nutlets. 


In dry sandy soil, Nebraska to Texas, Utah and 
Mexico. July—Sept. 


6. Heliotropium indicum L. Indian Helio- 
trope. Fig: 3507: 
Heliotropium indicum L. Sp. Pl. 134. 1753. 


Alen 


D x 
CO 


ing 


Annual, more or less hirsute or hispid; stem com- 
monly branched, 1°-3° high. Leaves ovate or oval, 
obtuse or acute at the apex, obtuse rounded or sub- 
cordate at the base, 2’-6’ long, 1-33’ wide, repand or 
undulate, borne on margined petioles 3’-23’ long; 
flowers blue, 2”-3” broad, sessile in terminal dense 
bractless usually solitary scorpioid spikes which be- 
come 3-6’ long in fruit; calyx-segments lanceolate, 
acute, shorter than the strigose corolla-tube; style 
very short, deciduous; fruit deeply 2-lobed, glabrous, 
the lobes divergent, each finally splitting into 2 
nutlets, each of which is ribbed on the back. 


In waste places, North Carolina to Kentucky, Illinois, 
Missouri, Florida and Texas. Naturalized from India. 
Also in ballast about the northern seaports. Widely 
distributed in warm regions as a weed. Turnsole. 
May-Nov. 


2. CYNOGLOSSUM [Tourn.] L. Sp. Pl. 134. 1753. 


Hirsute or hispid (rarely glabrous) mostly tall herbs, with alternate entire leaves, the 
basal long-petioled, and purple blue or white flowers in panicled, more or less scorpioid 
racemes. Calyx 5-cleft or 5-parted, enlarged and spreading or reflexed in fruit. Corolla 
funnelform or salverform, the tube short, the throat closed by 5 scales opposite the imbri- 
cated rounded lobes. Stamens included; filaments short; anthers ovate or oblong. Ovary 
deeply 4-lobed, separating into 4 diverging nutlets in fruit; style mostly slender. Nutlets 
oblique, flat or convex above, attached laterally to the convex or conic receptacle, covered 
with short barbed prickles. [Greek, dog’s tongue. ] 

About 75 species of wide geographic distribution. Besides the following, some 3 others occur 
in western North America. Type species: Cynoglossum officinale L. 

Stem leafy to the top; flowers reddish, purple or white; nutlets flat. 1. C. officinale. 
Stem leafless above; flowers blue; nutlets convex. Wid 

Flowers about 5” broad; nutlets about 4” long. 2. C. virginianum. 

Flowers about 34” broad; nutlets about 212” long. 3. C. boreale. 


76 BORAGINACEAE. Vou. II. 


1. Cynoglossum officinale L. Hound’s-tongue. Gipsy Flower. Fig. 3508. 


Cynoglossum officinale L. Sp. Pl. 134. 1753. 


Biennial, pubescent; stem erect, leafy to 
the top, stout, usually branched, 13°-3° 
a high. Basal and lower leaves oblong or 
i <i pal: oblong-lanceolate, slender-petioled, some- 

Als; times obtuse, 6-12’ long, 1-3’ wide; upper 
leaves lanceolate, acute or acuminate, ses- 
sile, or the uppermost clasping; racemes 
several or numerous, bractless or sparingly 
bracted, simple or branched, much elon- 
gated in fruit; pedicels 3-6” long; calyx- 
segments ovate-lanceolate, acute; corolla 
reddish-purple or rarely white, about 4” 
broad; fruit pyramidal, about 5” broad, 
each of the 4 nutlets forming a side of the 
pyramid, flat on their upper faces, mar- 
gined, splitting away at maturity, but hang- 
ing attached to portions of the subulate 
style. 

In fields and waste places, Quebec and On- 
tario to Manitoba, South Carolina, Alabama, 
Kansas and Montana. Often a troublesome 
weed. Naturalized from Europe. Native 


also of Asia. Called also dog’s-tongue, rose 
noble. Canadian or dog-bur. Sheep-lice. Tory-weed. Wood-mat. May-—Sept. 


2. Cynoglossum virginianum L. Wild Comfrey. Fig. 3509. 


Cynoglossum virginianum L. Sp. Pl. 134. 

1753. 

Perennial, hirsute; stem usually sim- 
ple, leafless above, stout, 13°-23° high. 
Basal and lower leaves oval or oblong, 
4’-12’ long, obtuse at the apex, nar- 
rowed into petioles; mnpper leaves ob- 
long, or ovate-lanceolate, sessile and 
clasping by a cordate base, acute, nearly 
as large, or the one or two uppermost 
quite small; racemes 2-6, corymbose, 
bractless, long-peduncled; flowers blue, 
about 5”.broad; corolla-lobes obtuse; 
calyx-segments oblong-lanceolate, ob- 
tuse, about 2” long at flowering time; 
fruit depressed, 4” broad, the nutlets 
convex on the upper face, not margined, 
separating and falling away at matur- 
ity, about 4” long. 

In woods, New Jersey to Kentucky, Mis- 3 
souri, Florida, Louisiana and Kansas. 5 


Ascends to 2500 ft. in Virginia. Dog-bur. 
April-May. 


GENUS 2. BORAGE FAMILY. a 


3. Cynoglossum boreale Fernald. North- 
ern Wild Comfrey. Fig. 3510. 


Cynoglossum boreale Fernald, Rhodora 7: 250. 1906. 


Similar to the preceding species, the stem 
more slender, villous below, appressed-pubescent 
above. Upper leaves clasping the stem; lower and 
basal leaves oblong, acute or acutish, long-peti- 
oled; racemes few, the flowers distant; calyx- 
segments only about 1” long at flowering time; 
corolla 3-4” broad; nutlets 2’-23” long. 


Woods and banks, Quebec to Ontario,-Connecticut, 
New York and Minnesota. In our first edition in- 
cluded in the preceding species, of which it may be 
a northern race. May—June. 


. eZ 
3. LAPPULA [Rivin.] Moench, Meth. 416. 1794. 
[EcHINOSPERMUM Sw.; Lehm. Asperif. 113. 1818.] 


Annual or perennial rough-pubescent or canescent erect branching herbs, with alternate 
narrow entire leaves, and small or minute blue or white flowers, in terminal bracted or 
bractless racemes. Calyx deeply 5-cleft or 5-parted, the segments narrow. Corolla salver- 
form or funnelform, the tube very short, the throat closed by 5 scales, the lobes obtuse, 
spreading, imbricated in the bud. Stamens included; filaments very short. Ovary 4-lobed; 
style short. Nutlets 4, erect or incurved, laterally attached to the receptacle, at length 
separating, the margins or backs armed with stout often flattened barbed prickles, the sides 
usually papillose or tuberculate. [Diminutive of the Latin /appa, a bur.] 


About 40 species, mostly natives of the north temperate zone. Besides the following, several 
others occur in western North America. Type species: Lappula Myosotis Moench. 


Racemes bracted ; fruiting pedicels not deflexed. 


Prickles in 2 rows on the margins of the nutlets, distinct. 1. L. Lappula. 

Prickles in 1 row on the margins, more or less confluent. 2. L. texana. 
Racemes bracted only at the base; fruiting pedicels deflexed. 

Stem-leaves ovate-oblong, the basal cordate; fruit globose. 3. L. virginiana. 


Leaves oblong, oblong-lanceolate or linear; fruit pyramidal. 
Flowers 3”—5” broad; fruit about 3” broad. 


: is . L. floribunda. 
Flowers 1”—2” broad; fruit about 2” broad. 


. L. deflexa. 


ap 


1. Lappula Lappula (L.) Karst. European Stickseed. Burseed. Fig. 3511. 


Myosotis Lappula L. Sp. Pl. 131. 1753. 
Lappula echinata Gilib. Fl. Lithuan. 1: 25. 1781. 
Lappula Myosotis Moench, Meth. 417. 1794. 
Echinospermum Lappula Lehm. Asperif. 121. 1818. 
Lappula Lappula Karst. Deutsch. Fl. 979. 1880-83. 

Annual, pale, leafy, hispid or appressed-pubes- 
cent, branched, 1°-2° high, the branches erect. 
Leaves linear, linear-oblong or the lowest spatu- 
late, sessile or the lower narrowed into petioles, 
ascending or erect, obtuse or obtusish at the apex, 
4’-14’ long; racemes leafy-bracted, more or less 
I-sided; pedicels very short, stout, not deflexed 
in fruit; calyx-segments lanceolate, becoming 
unequal and spreading; corolla blue, about 1” 
broad; fruit globose-oval, 14” in diameter; the 
nutlets papillose or also prickly on the back, the 
margins armed with 2 rows of slender distinct 
prickles. 

In waste places, Nova Scotia to British Columbia, 
south to New Jersey and Kansas. Naturalized from 


Europe. Native also of Asia. Stick-tight. Small 
sheep-bur. May-Sept. 


78 BORAGINACEAE. Vow. IIL: 


2. Lappula texana (Scheele) Britton. Hairy 
Stickseed. Fig. 3512. 
Cynoglossum pilosum Nutt. Gen. 1: 114. 1818. Not 

R. & P. 1794. 

Echinospermum texanum Scheele, Linnaea 25: 260. 1852. 
Echinospermum Redowskii var. cupulatum A. Gray in 

Brewer & Wats. Bot. Cal. 1: 530. 1876. 

Lappula texana Britton, Mem. Torr. Club 5: 273. 1894. 
L. Redowskii occidentalis Rydb. Contr. Nat. Herb. 3: 

170. 1895. : 

Annual, similar to the preceding species, 6—2° 
high, paniculately branched, the branches ascending 
or erect. Leaves linear or linear-oblong, mostly 
obtuse, the lower narrowed into petioles; racemes 
leafy-bracted; pedicels short, not deflexed in fruit; 
flowers about 1” broad; nutlets papillose-tuberculate 
on the back, the margins armed with a single row of 
flat, usually:more or less confluent bristles, or these 
united into a cup. 

In dry soil, Ontario to Manitoba, British Columbia, 
south to Nebraska, Texas and Arizona. Found also in 
ballast at Atlantic seaports. Nutlets with nearly dis- 
tinct bristles and others with bristles united into a cup 
sometimes occur on the same fruit. April—Aug. 


3. Lappula virginiana (L.) Greene. Virginia Stickseed. Fig. 3513. 
Myosotis virginiana L. Sp. Pl. 131. 1753. a® 

Cynoglossum Morisoni DC. Prodr. 10: 155. 1846. ai WE - 

E. virginicum Lehm. Asperif. 120. 1818. BX 
Lappula virginiana Greene, Pittonia 2: 182. 1891. 

Biennial, pubescent; stem paniculately branched, 
2°-4° high, the branches slender, spreading. Basal 
leaves (seldom present at flowering time) ovate or 
nearly orbicular, cordate, long-petioled, mostly 
obtuse; stem leaves ovate-oblong or oval, acute or 
acuminate at the apex, narrowed to the base, peti- 
oled, 3-8’ long, 1-4’ wide, the uppermost smaller, 
sessile; racemes very slender, divergent, bracted at 
the base, the bracts similar to the upper leaves; 
pedicels slender, short, recurved in fruit; corolla 
nearly white, about 1” broad; fruit globose, nearly 
2” in diameter; nutlets covered on the margins and 
usually also on the back by the slender distinct flat- 
tened barbed prickles, the backs commonly also 
more or less papillose. 

In dry woods and thickets, New Brunswick to On- 
tario, Minnesota, Georgia, Louisiana, Nebraska and 
Kansas. Called beggar’s-ticks or -lice. Virginia mouse- 
ear. Soldiers. Stick-tight. Dysentery-root or -weed. 
Tane—Sept. 


4. Lappula floribinda (Lehm.) Greene. 
Large-flowered Stickseed. Fig. 3514. 


Echinospermum floribundum WLehm. in Hook. FI. 
Bor. Am. 2: 84. pl.:164. 1834. 
Lappula floribunda Greene, Pittonia 2: 182. 1891. 


Biennial or perennial, rough-pubescent; stem 
stout, paniculately branched, 2°-5° high, the 
branches nearly erect. Leaves oblong, oblong- 
lanceolate, or linear-lanceolate, 2-4’ long, 2’—10” 
wide, sessile, acute or obtuse at the apex, or the 
lower narrowed into petioles; racemes numerous, 
erect or nearly so, very densely flowered, bracted 
at the base, many of them in pairs; pedicels 2’—4” 
long, reflexed in fruit; flowers blue or white, 
3-5” broad; fruit pyramidal, about 3” broad; 
nutlets keeled, papillose-tuberculate on the back, 
the margins armed with a single row of flat 
prickles, which are sometimes confluent at the 
base. 


Western Ontario and Minnesota to Saskatchewan, 
British Columbia, south to New Mexico and Califor- 
nia. June—Aug. 


GENUS 3. BORAGE FAMILY. 


5. Lappula defléxa (Wahl.) Garcke. Nod- 
ding Stickseed. Fig. 3515. 
Echinospermum deflexum var. americanum A, Gray, 

Proc. Am. Acad. 17: 224. 1882. 

Lappula deflexa Garcke, Fl. Deutsch. Ed. 6, 275. 1863. 
= Pa americana Rydberg, Bull. Torr. Club 24: 294. 

1897. 

Annual, rough-puberulent; stem slender, erect, 
paniculately branched, 1°-3° high, the branches 
spreading or ascending. Leaves oblong or oblong- 
lanceolate, mostly narrowed at both ends, 2’-4’ long, 
23-6” wide, the lower petioled, the upper sessile; 
racemes slender, many-flowered; pedicels slender, 
2”-4” long, deflexed in fruit; corolla white or 
bluish, about 1” broad; fruit pyramidal, about 2” 
broad; nutlets keeled, papillose-tuberculate on the 
back, rarely with a few prickles on the keel, the 
margins armed with a single row of flat prickles, 
these sometimes few. 

In thickets, Quebec to Manitoba, British Columbia, 
Towa, Nebraska and Wyoming. Also in Europe and 
Asia. May-—Aug. 


4. ALLOCARYA Greene, Pittonia, 1: 12) 4887. 


Mostly annual low herbs, with linear entire leaves, the lowest often opposite, and small 
flowers in terminal spikes or racemes. Pedicels thickened at the summit, persistent. Calyx 
5-divided, persistent, the segments narrow. Corolla salverform, white, yellow in the throat. 
Stamens included. Ovary 4-divided; style short. Nutlets crustaceous, smooth, or rough, 
attached at their base or below the middle to the receptacle, the scar of attachment concave 
or raised. [Greek, different nuts.] 

“ee About 25 species, natives of western North Amer- 


ica. Type species: Allocarya lithocarya (A. Gray) 
Greene. 


1. Allocarya scopul6rum Greene. Moun- 
tain Allocarya. Fig. 3516. 


Eritrichium californicum var. subglochidiatum A, 
Gray, Bot. Cal. 1: 526. In part. 1876. 


Allocarya scopulorum Greene, Pittonia 1: 16. 1887. 


Somewhat succulent, pubescent with scattered 
stiff appressed hairs, branched, the slender 
spreading branches 17-8’ long. Leaves 6-18” 
long, 1-13” wide, sessile or very short-petioled; 
flowers about 1” broad, distant, borne in most of 
the axils, very short-pedicelled; floral bracts 
similar to the leaves, but shorter; calyx segments 
linear-lanceolate; nutlets reticulate on the back, 
lightly grooved on the ventral side. 


Western Nebraska to Montana, Wyoming and 
Colorado. June—Sept. 


4 ~ 
5. CRYPTANTHA Lehm. Sem. Hort. Hamburg. 1832. F. & M. Ind. Sem. 
Hort. Petrop. 2: 35. 1836. 
[Krynitzk1a F. & M. Ind. Sem. Hort. Petrop. 7: 52. 1841.] 

Low annual setose or hispid branched herbs, with narrow alternate entire leaves, and 
small mostly white flowers, in scorpioid bractless or bracteolate spikes. Calyx 5-parted or 
5-cleft, at length deciduous from the spike, the lobes or segments erect, mostly connivent in 
fruit. Corolla small, funnelform, usually with 5 scales closing the throat, the lobes imbri- 
‘cated in the bud. Stamens included; filaments short. Ovary 4-divided; style short; stigma 
capitellate. Nutlets erect, rounded on the back, not keeled, the margins obtuse, acute or 
wing-margined, attached laterally to the conic or elongated receptacle, the scar of attach- 
ment mostly longer than broad. [Greek, hidden-flowered. ] 

About 50 species, natives of North and South America, mostly of the western United States. 
Type species: Cryptantha glomerata Lehm, 


Nutlets, at least some of them, with short processes. 1. C, crassisepala, 
All four nutlets smooth and shining. 2. C. Fendleri. 


80 BORAGINACEAE. Vou. III. 


1. Cryptantha crassisépala (T. & G.) Greene. Thick-sepaled Cryptanthe. 
Pigs 2507. 


Eritrichium crassisepalum T. & G. Pac. R. R. 
Rep: 22171. 1654. 

Krynitzkia crassisepala A. Gray, Proc. Am. 
Acad. 20: 268. 1885. 

Cryptanthe crassisepala Greene, Pittonia 1: 


Du2. Lesze 

Densely hispid, at length much branched, 
3-6 high. Leaves linear, or linear- 
spatulate, 4’-12’ long, sessile, or the lower 
narrowed into petioles; spikes very densely 
flowered; flowers about 2” broad, sessile, 
bracteolate, the bractlets slightly longer 
than the calyx; fruiting calyx 3’ long, 
closing over the fruit, its segments linear, 
obtusish, their midribs much thickened; 
fruit of 3 finely muricate nutlets, and I 
larger smooth and shining nutlet about 1” 
long, attached to the receptacle from the 
base to near the middle. 


In dry soil, Saskatchewan to Nebraska, 
Kansas, Texas and New Mexico. Rosita. 
June—Aug. 


2. Cryptantha Féndleri (A. Gray) 
Greene. Fendler’s Cryptanthe. 
Fig. 3518. 


Krynitzkia Fendleri A. Gray, Proc. Am. Acad. 
20: 268. 1885. 

Cryptanthe Fendleri Greene, Pittonia I: 120. 
1887. 


Erect, hispid; stem slender, paniculately 
branched, 6-15’ high. Leaves linear, or the 
lowest linear-spatulate, 1’-23’ long; spikes 
slender, bracteolate only at the base; flowers 
sessile, 1-2” broad; fruiting calyx nearly 
closed, its segments linear, leaf-like, herba- 
ceous, about 2” long; nutlets 4, all alike, 
brown, smooth and shining, less than 1” long, 
attached to the receptacle from the base to 
about the middle. 


In dry soil, Saskatchewan to Washington, south 
to Nebraska and Arizona. June—Aug. 


6. OREOCARYA Greene, Pittonia, 1: 57. 1887. 


Perennial or biennial hispid or strigose-pubescent herbs, mostly with thick woody roots, 
alternate or basal narrow leaves, and small white racemose-paniculate or densely thyrsoid 
flowers. Calyx very deeply 5-parted or 5-divided, the segments lanceolate, more or less 
spreading or recurved in fruit. Corolla funnelform or salverform, mostly crested in the 
throat, s-lobed. Stamens included. Ovary 4-divided; style mostly short. Nutlets 4, later- 
ally attached to the receptacle, not keeled, their margins acute or winged. [Greek, moun- 
tain nut.] Called White Forget-me-not. 

About 9 species, natives of western North America and Mexico. Type species: Oreocarya 
suffruticosa (Torr.) Greene. 


Inflorescence racemose-paniculate ; nutlets smooth. 1. O. suffruticosa. 
Inflorescence thyrsoid or thyrsoid-glomerate ; nutlets rough. 
Corolla-tube not longer than the calyx, little longer than the lobes. 
Densely rough-hairy, 6’—18’ high. 2. O. glomerata. 
Silvery appressed-pubescent, 3’—6’ high. 3. O. sericea. 
Corolla-tube longer than the calyx, 2 or 3 times as long as the lobes. 4. O. fulvocanescens. 


GENUS 6. BORAGE FAMILY. 81 


1. Oreocarya suffruticdsa (Torr.) Greene. 
Shrubby Oreocarya. Fig. 3519. 
peo ecots suffruticosa Torr. Ann. Lye. N. Y. 2: 225. 
1627. 
Epirichinm Jamesii Torr. in Marcy’s Rep. 294. 1853. 
seontieria Jamesii A. Gray, Proc. Am. Acad. 20: 278. 
I . 
OR ca suffruticosa Greene, Pittonia 1: 57. 1887. 
Perennial, rather stout, branched from the 
base and sometimes also above, strigose-pubescent 
or somewhat hirsute, 5’-12’ high. Upper leaves 
linear, 2-1’ long, the lower oblanceolate, some- 
what longer, obtuse or acute; racemes slender, 
panicled; pedicels about 1” long; calyx canescent 
and somewhat hispid, the segments slightly 
spreading, or erect in fruit; bractlets longer than 
the fruiting calyx; corolla 14’-3” broad; its tube 
about equalling the calyx; nutlets smooth, nearly 
1” long, shining, closely fitting together, trian- 
gular, acute-margined, nearly as wide as high. 
In dry soil, western Nebraska and Wyoming to 
Texas and Arizona. May—Aug. 


2. Oreocarya glomerata (Pursh) Greene. 
Clustered Oreocarya. Fig. 3520. 
Cuncclosswm glomeratum Pursh, Fl. Am. Sept. 729. 
aes yi glomeratum DC. Prodr. 10: 131. 1846. 
Krynitzkia glomerata A. Gray, Proc. Am. Acad. 20: 

279. 1885. 
O. glomerata Greene, Pittonia 1: 58. 1887. 

Perennial or biennial, densely hispid; stem 
erect, stout, simple or branched, 6-18’ high. 
Leaves spatulate or the upper linear, obtuse, 
1-23’ long, the basal commonly tufted; inflores- 
cence of thyrsoid clusters, the short dense lateral 
spike-like clusters mostly longer than the sub- 
tending bracts; calyx densely bristly; corolla 
3-5” broad; fruit pyramidal, the nutlets trian- 
gular-ovate, acute, acutely margined, papillose 
on the back. 

In dry soil, Manitoba to Nebraska, New Mexico 
and Utah. May-—Sept. 

Oreocarya thyrsifldra Greene, a related species 
of the Rocky Mountain region, enters our limits in 
western Nebraska. 


3. Oreocarya sericea (A. Gray) Greene. 
Low Oreocarya. Fig. 3521. 


Eritrichium glomeratum var. humile A. Gray, Proc. 
Am. Acad. 10: 61. 1874. Not &. humile DC. 
Krymitzkia sericea A. Gray, Proc. Am. Acad. 20: 279. 

188 
Oreocarya sericea Greene, Pittonia 1: 58. 1887. 


Perennial, low, tufted from the woody root; 
stems usually simple, 3-6’ high, silvery appressed- 
pubescent, or hirsute above. Leaves _linear- 
spatulate, %—-1’ long, 1’-13” wide, obtuse or 
acutish, imbricated on the short sterile shoots 
and at the bases of the flowering stems; inflores- 
cence thyrsoid or glomerate, usually short; calyx 
densely hispid; corolla 2’-3” broad, its tube not 
longer than the calyx; style short; nutlets acutely 
margined, acute, papillose on the back. 


In dry soil, Northwest Territory to Nebraska and 
Utah. May-Sept. 


6 


BORAGINACEAE. Vor, ike 


4. Oreocarya fulvocanéscens (A. Gray) 
Greene. Tawny Oreocarya. Fig. 3522. 


Eritrichium fulvocanescens A. Gray, Proc. Am. Acad. 
LOO, HES7A: 

Eritrichium glomeratum var. (?) fulvocanescens S. 
Wats. Bot. King’s Exp. 243. pl. 23. f. 7. 1871. 

Oreocarya fulvocanescens Greene, Pittonia 1: 58. 1887. 


Perennial, tufted, similar to the preceding species 
but densely strigose or hirsute. Leaves spatulate, 
or oblanceolate, obtuse, the lower and basal ones 
1-13’ long; inflorescence of thyrsoid clusters; calyx 
densely setose with yellowish hairs; corolla about 
2” broad, its tube longer than the calyx, 2 or 3 
times the length of the lobes; style filiform; nutlets 
acutely margined, tuberculate on the back. 


In dry soil, western Nebraska (according to Web- 
ber), Wyoming to Nevada and New Mexico. May—Aug. 


7. PNEUMARIA Hill, Veg. Syst. 7: 40. pl. 37. 1764. 

A perennial fleshy glabrous glaucous diffusely branched herb, with alternate entire 
leaves, and small blue pinkish or white flowers in loose terminal leafy-bracted racemes. 
Calyx-lobes triangular-ovate or lanceolate, somewhat enlarging in fruit. Corolla tubular- 
campanulate, crested in the throat, 5-lobed, the lobes imbricated in the bud, slightly spread- 
ing. Filaments slender, scarcely exserted. Ovary 4-divided; style slender. Nutlets erect, 
fleshy, attached just above their bases to the somewhat elevated receptacle, smooth, shining, 
acutish-margined, becoming utricle-like when mature. 


A monotypic genus of sea-beaches of the north 
temperate zone. 


1. Pneumaria maritima (L.) Hill. Sea 
Lungwort. Sea Bugloss. Oyster Plant. 


Big, 3522: 
Pulmonaria maritima L. Sp. Pl. 136. 1753. 
P. maritima Hill, Veg. Syst. 40. pl. 37. f. 3. 1764. 
Mertensia maritima S. F. Gray, Nat. Arr. Brit. Pl. 

354. 1821. 

Pale green, the branches spreading or ascend- 
ing, 3-15’ long. Leaves thick, ovate, obovate, or 
oblong, 1’-4’ long, acute or obtuse at the apex, 
narrowed at the base, the lower and basal ones 
contracted into margined petioles, the uppermost 
smaller; flowers blue or nearly white, about 3” 
long, all pedicelled; pedicels very slender, 5-18” 
long; calyx shorter than the corolla-tube; corolla 
with a crest in the throat opposite each lobe; 
nutlets about as long as the calyx-lobes when 
mature. 

On sea-beaches, Long Island (?), Massachusetts to 
Newfoundland and Greenland, Oregon to Alaska. 
Also on the coasts of Europe and Asia. May-—Sept. 


8. MERTENSIA Roth, Catal. Bot. 1: 34. 1797. 


Perennial glabrous or pubescent herbs, with alternate sometimes punctate leaves, and 
rather large blue purple or white flowers, in panicles, cymes, or racemes. Calyx-lobes lan- 
ceolate or linear, little enlarged in fruit. Corolla tubular-funnelform or trumpet-shaped, 
crested or unappendaged in the throat, its lobes obtuse, imbricated, little spreading. Stamens 
inserted on the tube of the corolla, included, or scarcely exserted; filaments flattened, or 
filiform; anthers oblong or linear, obtuse. Ovary 4-divided; style filiform. Nutlets erect, 
coriaceous, wrinkled when mature, attached above their bases to the convex or nearly flat 
receptacle. [In honor of Prof. C. F. Mertens, a German botanist. ] 


About 40 species, natives of the northern hemisphere. Besides the following, many occur 
in the western part of North America. The species are called Smooth Lungwort. Type species: 
Mertensia pulmonarioides Roth. 


Corolla trumpet-shaped, not crested in the throat, the limb barely 5-lobed. 1. M. virginica. 
Corolla funnelform-campanulate, crested in the throat, the limb manifestly 5-lobed. 
Stem-leaves ovate or ovate-lanceolate, acuminate. 2. M. paniculata. 


Stem-leaves oblong or lanceolate, obtuse or acute. 3. M. lanceolata. 


Genus 8. BORAGE FAMILY. . 83 


1. Mertensia virginica (L.) DC. Virginia 
Cowslip. Tree Lungwort. Roanoke- 
bells. Fig. 3524. 


Pulmonaria virginica 1.. Sp. Pl. 135. 1753. 
Mertensia virginica DC. Prodr. 10: 88. 1846. 


Glabrous; stem erect, or ascending, simple or 
sometimes branched, 1°-2° high, rather stout. 
Leaves oblong, oval, or obovate, pinnately 
veined, obtuse at the apex, 2’-5’ long, the upper- 
most sessile, the lower narrowed into margined 
petioles; racemes short, corymb-like; pedicels 
2”-6” long; flowers blue-purple, very showy, 
about I’ long; calyx-lobes oblong-lanceolate, 
obtusish, 1” long or less; corolla trumpet-shaped 
or nearly salverform, its tube cylindric, a little 
expanded above, longer than the 5-lobed plaited 
limb, pubescent at the base within, not crested in 
the throat; disk with two opposite linear lobes; 
filaments filiform, much longer than the anthers; 
nutlets not shining, rounded. 


In low meadows and along streams, southern On- 
tario to New Jersey and South Carolina, Minnesota, 
Nebraska and Kansas. Blue bells. March—May. 


re 2. Mertensia paniculata (Ait.) G. Don. 
Tall Lungwort. Fig. 3525. 


P. paniculata Ait. Hort. Kew. 1: 181. 1789. 
M. paniculata G. Don, Gen. Syst. 4: 318. 1838. 

Roughish-pubescent, dark green; stem erect, 
branched above, 12°-3° high, the branches slen- 
der. Leaves thin, pinnately veined, those of the 
stem ovate or ovate-lanceolate, acuminate at the 
apex, narrowed at the base, 2-5’ long, the lower 
narrowed into slender petioles; basal leaves ovate, 
rounded or cordate at the base; racemes several- 
flowered, panicled; pedicels filiform, 4”—10” long; 
flowers purple-blue, 6’-7” long; calyx-lobes lan- 
ceolate,acute; corolla tubular-campanulate, crested 
in the throat, the tube about twice as long as the 
calyx and exceeding the 5-lobed limb; filaments 
flattened, slightly longer than the anthers; style 
filiform, usually somewhat exserted; nutlets 
rounded. 

In woods or thickets, Hudson Bay to Alaska, south 
to Michigan, Nebraska, Idaho and Washington. 
July-Aug. 


Y 


3. Mertensia lanceolata (Pursh) DC. 
Lance-leaved Lungwort. Fig. 3520. 


P. lanceolata Pursh, Fl. Am. Sept. 729. 1814. 
Mertensia lanceolata DC. Prodr. 10: 88. 1846. 
M. linearis Greene, Pittonia 3: 197. 1897. 


Glabrous or somewhat hirsute; stem simple or 
branched, slender, 6-18’ high. Leaves papillose, 
indistinctly veined, light green, the upper lanceo- 
late, acute, sessile or slightly clasping at the base, 
the lower oblong or oblanceolate, obtuse, 3-4 
long, narrowed into margined petioles; racemes 
few-flowered, usually panicled; flowers blue, 5 - 
6” long; pedicels 3-7” long; calyx-lobes lanceo- 
late, obtuse, or acutish; corolla tubular-campanu- 
late, the tube longer than the calyx and longer 
than the 5-lobed limb, hairy at the base within, 
the throat crested; filaments a little longer than 
the anthers; style filiform, scarcely exserted. 


In thickets, western Nebraska . to Manitoba, 
Wyoming, Idaho and New Mexico. Races differ 
in amount or absence of pubescence and in width 
of leaves. June—Aug. 


84 BORAGINACEAE. Vor. III. 


Q- AMSINCKIA Lehm. Del. Sem. Hort. Hamb. 71 153i 


Annual hispid or setose herbs, with narrow leaves, the yellow flowers in scorpioid spikes. 
Calyx 5-parted, the segments linear. Corolla salverform or funnelform, the tube slender, 
naked or minutely crested in the throat, the 5 lobes spreading. Stamens 5, borne on the 
corolla-tube, included. Ovary deeply 4-lobed. Nutlets ovoid, rough, laterally attached to 
the receptacle below the middie. [In honor of William Amsinck, a burgomaster of Hamburg 
and friend of the Hamburg botanical garden.] 


About 15 species, natives of western North America and Chile, the following typical. 


1. Amsinckia lycopsioides Lehm. Amsinckia. 
Fig. 3527. 


Lithospermum lycopsioides Lehm. Pug.2: 28. 1830. 
Amsinckia lycopsioides Lehm.; DC. Prodr.10: 117. 1846. 


Diffusely branched, loosely hispid with long, bristly 
hairs, the branches often 1° long, decumbent or ascend- 
ing. Leaves lanceolate, oblong-lanceolate or ovate- 
lanceolate, repand-dentate or entire, 3° long or less, 
sessile; scorpioid spikes short in flower, elongating in 
fruit, the lower flowers bracteolate, the upper ones com- 
monly bractless; corolla about 4” long, its tube some- 
what longer than the calyx; nutlets rugose-reticulate. 

Waste grounds, Massachusetts and Connecticut. Ad- 
ventive from California. May-July. 


Amsinckia intermédia F. & M., an erect Californian 
species, with orange-yellow flowers and linear leaves, has 
been found in eastern Long Island and Nantucket. 


10. ASPERUGO [Tourn.] L. Sp. Pl. 138. 1753. 


An annual rough-hispid procumbent herb, with alternate entire leaves, or the uppermost 
sometimes opposite, and small blue or nearly white flowers, short-pedicelled and 1-3 together 
in the upper axils. Calyx campanulate, unequally 5-cleft, much enlarged and folded together 
in fruit, the lobes incised-dentate. Corolla tubular-campanulate, 5-lobed, the lobes imbricated. 
Stamens 5, included, inserted on the corolla-tube; filaments very short. Ovary 4-divided; 
style short; stigma capitate. Nutlets 4, ovoid, erect, granular-tuberculate, keeled, laterally 
attached above the middle to the elongated-conic receptacle. [Latin, rough, referring to 
the leaves. ] 


A monotypic genus of Europe and Asia. 
1. Asperugo procimbens L. German Mad- 49 99 
wort. Catchweed. Fig. 3528. 


Asperugo procumbens L. Sp. Pl. 138. 1753. NED 

~ FEZ 

Stems slender, branched, diffusely procumbent, ‘ Zo 
6’-18’ long, very rough with stiff bristly hairs. Leaves f 


Zz 


. 
ae 


oblong, lanceolate, or the lower spatulate, obtuse or | RY PA Bi, 
acutish at the apex, 4’-12’ long, the lower narrowed YN bien 
into margined petioles; flowers very short-pedicelled, y; 7 Wy 
about 1” broad, blue, the pedicels recurved in fruit; Mls ey 


fruiting calyx dry and membranous, strongly veined, 
4-6” broad; nutlets obliquely ovoid. Z 


In waste places and ballast, Massachusetts to south- LZ LY 

ern New York, New Jersey, District of Columbia and : Li 2 LY 
Minnesota. Adventive from Europe. Called also small Se BE 

wild bugloss and great goose-grass. May—Aug. : ; 


GENUS II. BORAGE FAMILY. 85 


11. MYOSOTIS [Dill.] L. Sp. Pl. 131. 1753. 


Low annual biennial or perennial, more or less pubescent, branching, diffuse or erect 
herbs, with alternate entire leaves, and small blue pink or white flowers in many-flowered 
elongated bractless more or less I-sided racemes, or these sometimes leafy at the base. 
Calyx 5-cleft, the lobes narrow, spreading or erect in fruit. Corolla salverform, the limb 
5-lobed, the lobes convolute in the bud, rounded, the throat crested. Stamens 5, included, 
inserted on the corolla-tube; filaments filiform; anthers obtuse. Ovary 4-divided, style fili- 
form. Nutlets erect, glabrous or pilose, attached by their bases to the receptacle, the scar of 
attachment small, flat. [Greek, mouse-ear. | 

About 35 species of wide geographic distribution. Besides the following, 1 or 2 others occur im 


the southern and western parts of North America. Called forget-me-not and scorpion-grass. 
Type species: Myosotis scorpioides L. 


Hairs of the calyx all straight ; perennial swamp or brook plants. 


Calyx-lobes shorter than the tube; corolla 3”’—4” broad. 1. M. scorpioides. 
Calyx-lobes as long as the tube; corolla 2”—3”' broad. 2. M.laxa. 

Hairs of the calyx, or some of them, with hooked tips; annuals or biennials. 
Fruiting pedicels longer than the calyx. 3. M. arvensis. 


Fruiting pedicels not longer than the calyx. 
Calyx-lobes equal; corolla blue or yellowish, changing to violet and blue. 
Corolla yellowish, changing to violet and blue; style longer than the nutlets. 
. 4. M. versicolor. 
Corolla blue; style not longer than the nutlets. 5. M. micrantha. 
Calyx-lobes unequal; corolla white. 6. M. virginica. 


1. Myosotis scorpioides L. Forget-me-not. 
Mouse-ear Scorpion-grass. Fig. 3529. 
Myosotis scorpioides var..palustris L. Sp. Pl. 131. 1753. 

Myosotis palustris Lam. Fl. Fr. 2: 283. 1778. 

Appressed-pubescent, perennial, with slender root- 
stocks or stolons; stems slender, decumbent or as- 
cending, rooting at the lower nodes, 6-18 long. 
Leaves oblong, oblanceolate or oblong-lanceolate, ob- 
tuse, narrowed at the base, 1-3’ long, 2”-6” wide, 
those of the stem sessile or very nearly so, or the 
lower petioled; racemes loosely many-flowered; pedi- 
cels longer than the calyx; calyx with straight ap- 
pressed hairs, its lobes equal, triangular-ovate, acute, 
shorter than the tube, spreading in fruit; corolla 
blue with a yellow eye, the limb flat, 3-4” broad; 
nutlets angled and keeled on the inner side. 


In brooks and marshes, Newfoundland to New York, 
Pennsylvania and Tennessee. Escaped from cultiva- 
tion. Native of Europe and Asia. Called also marsh 
scorpion-grass, snake-grass and love-me. May-July. 


2. Myosotis laxa Lehm. Smaller Forget- 
me-not. Fig. 3530. 


Myosotis lara Lehm. Asperif. 83. 1818. 


Myosotis palustris var. lava A. Gray, Man. Ed. 5, 
365. 1867. 


Perennial, appressed-pubescent, similar to the 
preceding species; stems decumbent, spreading, 
rooting at the nodes, 6-20’ long. Leaves oblong, 
oblong-lanceolate or spatulate, obtuse; racemes 
very loosely many-fiowered; pedicels spreading, 
much longer than the fruiting calyx; hairs of the 
calyx straight, appressed, its lobes equal, ovate- 
lanceolate, acutish, spreading in fruit, quite as 
long as the tube; corolla blue with a yellow eye, 
its limb concave, about 2” broad; nutlets convex 
on both the inner and outer sides. 


In wet muddy places, Newfoundland to Ontario, 
south to Virginia and Tennessee. Also in Europe. 
Ascends to 3500 ft. in Virginia. May-July. 


BORAGINACEAE, Vor, IIT. 


3. Myosotis arvénsis (L.) Hill. Field Scor- 
pion-grass or Mouse-ear. Fig. 3531. 


Myosotis scorpioides var. arvensis L. Sp. Pl. 131. 1753. 
Myosotis arvensis Hill, Veg. Syst. 7: 55. 1764. 


Annual or biennial, hirsute-pubescent; stem erect, 
branched, 6-18 high. Basal and lower leaves ob- 
lanceolate, obtuse, petioled or sessile; stem leaves 
mostly oblong or oblong-lanceolate, obtuse or acutish 
at the apex, narrowed to the sessile base, 412’ long, 
2-4” wide; racemes loosely flowered; fruiting pedi- 
cels longer than the calyx; hairs of the calyx, or 
some of them, with minutely hooked tips, the lobes 
equal, erect, or connivent in fruit, triangular-lanceo- 
late, acute, about as long as the tube; corolla blue 
or white, the limb concave, 1’-13” broad; nutlets 
convex on the outer side, somewhat keeled on the 
inner. 


In fields, Newfoundland to western Ontario and 
Minnesota, south to West Virginia. Perhaps not 
indigenous. Also in Europe. June—Aug. 


4. Myosotis versicolor (Pers.) J. E. Smith. Yellow and Blue Scorpion-grass. 
BIg. 2532: 
M. sormenisis var. (?) versicolor Pers. Syn. 1: 156. 
1805. 


Myosotis versicolor J. E. Smith, Engl. Bot. pl. 480. 
1813. 


Annual, hirsute-pubescent, with mostly straight 
hairs, often much branched above; stems slender, 
erect or ascending, 4-12’ high. Leaves oblong, 
obtuse or obtusish, sessile, or nearly so, or the 
lower spatulate and narrowed into margined peti- 
oles; racemes slender, mostly naked below; pedi- 
cels shorter than the fruiting calyx, appressed- 
pubescent, erect; calyx equally 5-cleft, the lobes 
iinear-lanceolate, erect or connivent in fruit, 
longer than or equalling the tube, the hairs, or 
some of them, with minutely hooked tips; corolla 
pale yellow changing to violet and blue, its limb 
about 1” broad; nutlets convex on the outer, 
slightly keeled on the inner side, shorter than 
the style. 


In fields and along roadsides, southern New York 
and Delaware. Naturalized from Europe. May- 


5. Myosotis micrantha Pall. Blue Scorpion- 
grass. Bigg5 33 


Myosotis micrantha Pall.; Lehm. Neue Schr. Naturf. Ges. 
Halle 37: 24. 1817. 


Annual or biennial, 8’ high or less, branched from 
near the base, the pubescence of both straight and 
hooked hairs. Leaves oblong to lanceolate, mostly ses- 
sile, obtuse or obtusish; racemes slender, distantly flow- 
ered to the base; pedicels much shorter than the calyx, 
pubescent, and usually with some hooked hairs; calyx 
equally 5-cleft, about as long as the corolla-tube; style 
not longer than the nutlets. 


Fields and roadsides, Ontario to Massachusetts and Ohio. 
May-July. Naturalized from Europe. Has been mistaken in 
America for M. collina Hoffm. 


_GENUS II, BORAGE FAMILY. 


6. Myosotis virginica (L.) B.S.P. Spring 
or Early Scorpion-grass. Fig. 3534. 

Lycopsis virginica L. Sp. Pl. 139. 1753. 
Myosotis verna Nutt. Gen. 2: Add. 1818. 
Myosotis virginica B.S.P. Prel. Cat. N. Y. 37. 1888. 

Annual or biennial, hirsute-pubescent or hispid, 
erect, branched, 3-15’ high, the branches erect. Leaves 
oblong or linear-oblong, sessile, 3’—-12” long, obtuse, 
or the lower spatulate and narrowed into short peti- 
oles; racemes usually bracted at the base, strict; 
pedicels ascending or erect, or slightly spreading at 
the apex; shorter than the fruiting calyx; calyx some- 
what 2-lipped, unequally 5-cleft, the lobes lanceolate, 
acute, longer than the tube, connivent in fruit, very 
hispid, the hafrs, or most of them, with minutely 
hooked tips; corolla white, the limb 14” broad or 
less; nutlets convex on the back, slightly keeled and 
margined on the inner side. 

On dry hills and banks, Maine and Ontario to Minne- 
sota, Florida and Texas. Forget-me-not. April-June. 

Myosotis macrospérma Engelm., of the Southern 
States, with larger flowers and fruit, the ripe calyx 
nodding or spreading, ranging north to Virginia and 
Kentucky, appears to be a race of this species. 


iz, LITHOSPERMUM [Tourn.] L. Sp. Pl. 132. 1753. 


Annual or perennial, erect branching or rarely simple, pubescent hirsute or hispid herbs, 
with alternate entire leaves, and small or large, white yellow or blue flowers in leafy-bracted 
spikes or racemes. Calyx 5-parted or 5-cleft, the segments or lobes narrow. Corolla funnel- 
form or salverform, 5-lobed, naked, pubescent or crested in the throat, the lobes entire or 
erose-denticulate, the tube sometimes pubescent at the base within. Stamens 5, included, 
inserted on the throat of the corolla; filaments short. Ovary 4-divided; style slender, or 
filiform; stigma capitate, or 2-lobed. Nutlets 4, or fewer, erect, white, smooth and shining, 
or brown and wrinkled, attached by their bases to the nearly flat receptacle, the scar of 
attachment not concave. [Greek, stone-seed, from the hard nutlets.] 


About 40 species, natives of the northern hemisphere, a few in South America and Africa. 
Besides the following, some 7 others occur in the southern and southwestern parts of the United 
States. Type species: Lithospermum officinale L. 

Corolla white or yellowish, its tube shorter than or equalling the calyx; flowers distant. 


Nutlets brown, wrinkled and pitted; annual or biennial. 1. L. arvense. 
Nutlets white, smooth and shining; perennials. 
Leaves ianceolate, acute; nutlets ovoid. 2. L. officinale. 
Leaves ovate, acuminate; nutlets globose-ovoid. 3. L. latifolium. 


Corolla dull yellow, its tube longer than the calyx; leaves lanceolate; flowers dense. 4. L. pilosum. 
Corolla bright yellow, its tube much longer than the calyx; flowers dense; red-rooted perennials. 
Corolla-lobes entire; flowers all complete. 


Hispid-pubescent ; corolla-tube bearded at the base within. 5. L. carolinense. 
Hirsute, somewhat canescent; corolla-tube not bearded at the base. 6. L. canescens. 
Corolla-lobes erose-denticulate; later flowers cleistogamous. 7. L. linearifolium. 


1. Lithospermum arvénse L. Bastard 
Alkanet. Corn Gromwell. Fig. 3535. 
Lithospermum arvense L. Sp. Pl. 132. 1753. 


Annual or biennial, appressed-pubescent; stem 
erect, usually branched, 6—20' high. Leaves bright 
green, lanceolate, linear or linear-oblong, sessile 
or the lowest short-petioled, mostly appressed, 
obtuse or acutish at the apex, narrowed at the 
base, indistinctly veined, 4-14’ long, 4-3” wide, 
the uppermost smaller; flowers sessile or very 
nearly so in the spikes, becoming distant, white, 
about 3” long; calyx-segments linear-lanceolate, 
longer than or equalling the corolla-tube; corolla 
funnelform, puberulent in the throat but not 
crested; nutlets brown, wrinkled and pitted, gla- 
brous, about 1” high, convex on the back, keeled 
on the inner side, one-third to one-half the length 
of the calyx-segments. 

In waste places and fields, Quebec to Ontario and 
Michigan, south to Georgia and Kansas. Natural- 
ized from Europe. Native also of Asia. Pearl-plant. 
Salfern-stoneseed. May-—Aug. 


88 BORAGINACEAE. Vor. IIT, 


2. Lithospermum officinale L. Gromwell. 
Fig. 3536. 
Lithospermum officinale L. Sp. Pl. 132. 1753. 


Perennial, finely puberulent; stem usually much 
branched, 2°-4° high, leafy. Leaves lanceolate or 
oblong-lanceolate, acute at the apex, narrowed at 
the base, few-veined, sessile, 13-4’ long, 3-12” 
wide, the upper surface rough, the lower pubes- 
cent; flowers yellowish-white, about 2” long, ses- 
sile; calyx-segments linear-lanceolate, about equal- 
ling the corolla-tube; corolla funnelform, crested 
in the throat; style about as long as the stamens; 
nutlets, when mature, white, smooth, shining, 
about 12” high, ovoid, obtuse, more than one-half 
as long as the calyx-segments, seldom all ripening. 

In fields and waste places, Quebec to southern New 
York, New Jersey and Minnesota. Plant grayish. 


Naturalized from Europe. Native also of Asia. 
Graymile. Littlewale. Pearl-plant. May—Aug. 


3. Lithospermum latifolium Michx. 
American Gromwell. Fig. 3537. 


Lithospermium latifolium Michx. Fl. Bor. Am. 1: 131. 

1803. 

Perennial, rough-puberulent; stem branched, 
2°-3° high, the branches long and slender. Leaves 
ovate or ovate-lanceolate, acuminate at the apex, 
pinnately veined, 2’-5’ long, 1-2’ wide, or the 
uppermost smaller, the lowest obtuse; flowers 
yellowish white or pale yellow, 2’-3” long, few, 
solitary, distan*; calyx-segments linear-lanceo- 
late, about as long as the corolla or a little 
longer; corolla funnelform, crested in the throat; 
style shorter than the stamens; nutlets white, 
shining, globose-ovoid, about 2” long, more than 
one-half as long as the calyx-segments. 

In dry thickets and fields, Quebec to New York, 


Minnesota, Kansas, Virginia, Tennessee and Arkan- 
sas. May. 


4. Lithospermum pilosum Nutt. Woolly 
Gromwell. Fig. 3538. 


Lithospermum pilosum Nutt. Journ. Phil. Acad. 7: 43. 
1834. 


Lithospermum Torreyi Nutt. loc. cit. 44. 1834. 


Perennial from thick roots, hirsute, -rather pale 
green; stems usually stout and clustered, very leafy, 
8’-18’ high. Leaves lanceolate or linear-lanceolate, 
2'-4’ long, 2”-5” wide, gradually acuminate to the 
apex, narrowed at the base, sessile, indistinctly 
veined; flowers dull yellow, very numerous and 
crowded in a terminal leafy thyrsus; calyx-seg- 
ments densely hirsute, shorter than the cylindric 
corolla-tube; corolla salverform, the throat puberu- 
lent below each lobe; style longer than the filaments ; 
nutlets ovoid, acute, white, shining, about 2” long. 

Western Nebraska (according to Williams) ; Wyo- 


ming to Montana, Alberta, British Columbia and Cali- 
fornia. May-July. 


Gaus 12) BORAGE FAMILY. 89 


5. Lithospermum carolinénse ( Walt.) MacM. 
Hairy or Gmelin’s Puccoon. Fig. 3539. 


Anonymos carolinensis Walt. Fl. Car. 91. 1788. 
Batschia carolinensis Gmel. Syst. 2: Part 1, 315. 1791. 
Lithospermum carolinianum Lam. Tabl. Encycl. 1: 397. 

1791. 
Lithospermum hirtum Lehm. Asperif. 305. 1818. 
Lithospermum carolinense MacM. Met. Minn. 438.. 1892. 

Perennial, hispid-pubescent, or scabrous; stems 
usually clustered, rather stout, simple, or branched 
above, 1°-23° high, very leafy. Leaves narrowly 
lanceolate, sessile, obtuse or acute at the apex, nar- 
rowed at the base, 2-3’ long, the lowest commonly 
reduced to appressed scales, the uppermost oblong; 
flowers 6-8” long, in dense short terminal leafy 
racemes, dimorphous; pedicels 1-3” long; calyx- 
segments linear-lanceolate, shorter than the tube of 
the orange-yellow salverform corolla; corolla-lobes 
entire, rounded, the throat crested, the tube bearded 
at the base within by Io hirsute teeth; nutlets white, 
shining, about 2” high, ovoid, very much shorter 
than the calyx-segments. 

In dry woods, western New York to Florida, Minne- 
sota, Montana and New Mexico. April—June. 


6. Lithospermum canéscens ( Michx.) 
Lehm. Hoary Puccoon. Fig. 3540. 


Batschia canescens Michx. Fl. Bor. Am. 1: 130. Pl. 
T4. 1803. 
Lithospermum canescens Lehm. Asperif. 305. 1818. 


Perennial, hirsute, somewhat canescent, at least 
when young; stems solitary or clustered, simple 
or often branched, 6-18 high. Leaves oblong, 
linear-oblong, or linear, obtuse or acutish at the 
apex, sessile by a narrowed base, 2-13’ long, 
2’-5’’ wide, the lowest often reduced to appressed 
scales; flowers about 6” long, sesssle, numerous 
in dense short leafy racemes, dimorphous; calyx- 
segments linear-lanceolate, shorter than the tube 
of the orange-yellow salverform corolla; corolla 
crested in the throat, its lobes rounded, entire, its 
tube glandular but not bearded at the base within; 
nutlets white, smooth, shining, acutish, shorter 
than the calyx-segments. 

In dry soil, Ontario to western New Jersey and 
Alabama, Saskatchewan, North Dakota and Texas. 
April—June. 

7. Lithospermum linearifolium Goldie. 

Narrow-leaved Puccoon. Fig. 3541. 
L. angustifolium Michx. Fl. Bor. Am. 1: 130. 1803. 

Not Forsk. 


L. linearifolium Goldie, Edinb. Phil. Journ. 1822: 
Roo 


Perennial by a deep root, strigose-pubescent and 
scabrous; stem branched, 62° high, the branches 
erect or ascending. Leaves linear, sessile, acute 
or acutish, 3’-2’ long, 14’-2}” wide; flowers of 
two kinds, in terminal leafy racemes; corolla of 
the earlier ones salverform, about 1’ long, bright 
yellow, the tube 3-5 times as long as the linear- 
lanceolate calyx-segments, the lobes erose-dentic- 
ulate, the throat crested, the base of the tube not 
bearded within; later flowers (sometimes all of 
them) much smaller, pale yellow, cleistogamous, 
abundantly fertile, their pedicels recurved in fruit; 
nutlets white, smooth, shining, ovoid, 14’-2” high, 
more or less pitted, keeled on the inner side. 

In dry soil, especially on prairies, Ontario and 


Indiana to Illinois, Kansas and Texas, west to Brit- k 
ish Columbia, Utah and Arizona. Yellow puccoon. April-July. 


9° BORAGINACEAE. VoL. III. 


T° ONOSMODIUM Michx. Bl) Bor Am) 24,122), 108: 


Perennial stout hispid or hirsute branching herbs, with alternate entire strongly veined 
leaves, and rather small yellowish or greenish white proterogynous flowers, in terminal leafy- 
bracted scorpioid spikes or racemes. ‘Calyx deeply 5-parted, the segments narrow. Corolla 
tubular or tubular-funnelform, 5-lobed, the lobes erect, the throat not appendaged, the sinuses 
slightly inflexed, the tube with a glandular 10-lobed band within at the base. Stamens 5, 
inserted on the tube or throat of the corolla, included; filaments short. Ovary 4-parted; 
style filiform, exserted. Nutlets 4, or commonly only 1 or 2 perfecting, ovoid, sometimes 
sparingly pitted, shining, smooth, white, attached by the base to the nearly flat receptacle, the 
scar of attachment small, flat. [Greek, like onosma, or ass-smell.] 


: About 10 species, natives of North America and Mexico. Besides the following, 3 others occur 
in the southern and southwestern United States. Type species: Onosmodium hispidum Michx. 


Corolla-lobes 2-3 times as long as wide. 1. O. virginianum. 
Corolla-lobes scarcely longer than wide. 
Stem glabrous below. 2. O. subsetosum. 
Stem hirsute or pubescent to the base. 
Pubescence silky ; nutlets distinctly pitted. 3. O. molle. 
Pubescence hirsute to strigose; nutlets indistinctly pitted. 
Nutlets not constricted. 4. O. occidentale. 
Nutlets distinctly constricted just above the base. 5. O. hispidissimum. 


xr. Onosmodium virginianum (L.) DC. Vir- 
ginia False Gromwell. Fig. 3542. 

Lithospermum virginianum L. Sp. Pl. 132. 1753. 

Onosmodium virginianum DC. Prodr. 10: 70. 1846. 


hairs; stem rather slender, usually branched above, 
1°-23° high. Leaves oblong, oval, or oblong-lanceo- 
late, obtuse or acutish, sessile, 1-33’ long, or the 
lower oblanceolate and narrowed into petioles; calyx- 
segments linear-lanceolate, acuminate; corolla cylin- 
dric or nearly so, yellowish-white, about 4” long, 
the lobes narrowly lanceolate, acuminate, 2 or 3 times 
as long as wide, nearly as long as the tube, strigose 
without; nutlets ovoid, obtuse or obtusish, pitted, 
1-13” long. 

In dry thickets or on hillsides, Massachusetts to Penn- 


sylvania, Florida and Louisiana. Ascends to 3000 ft. in 
Virginia. Wild job’s-tears. May-July. 


2. Onosmodium subsetosum Mack. & Bush. 
Ozark False Gromwell. Fig. 3543. 


O. subsetosum Mack. & Bush; Small, Fl. SE. U. S. troor. 
1903. 


Stem erect, glabrous, or with a few scattered ap- 
pressed hairs above, somewhat branched, 3° high o1 
less, the branches appressed-pubescent. Leaves lanceo- 
late, acute, papillose and appressed-hispid above, whitish 
appressed-pubescent beneath, the larger about 33’ long; 
bracts 4-1’ long; calyx-lobes oblong, obtuse, 3” long; 
corolla about 5’ long, canescent, its lobes triangular, 
acute, about 1” long; fruiting pedicels 2’-3” long; nut- 
lets whitish, ovoid, 13” long, obtuse or acutish, not con- 
stricted, sparingly pitted. 


Barrens, Ozark Mountains, Missouri and Arkansas. 
June—Aug. 


Densely appressed-hispid or strigose, with stiff 


GENUS 13. BORAGE FAMILY. 


3. Onosmodium modlle Michx. Soft-hairy False 
Gromwell. Fig. 3544. 


O. molle Michx. Fl. Bor. Am. 1: 133. 1803. 


Stem erect, branched above, about 2° high, hispid-pubes- 
cent or strigose, the branches soft-pubescent. Leaves 
lanceolate to ovate-lanceolate, the larger about 2’ long, 
densely soft-pubescent on both sides; bracts similar to the 
leaves, 1’ long or less; calyx-lobes linear-oblong, 3” long, 
obtusish; corolla 4-6” long, its lobes triangular, acute, 
1-13” long, pubescent outside; nutlets about 1” long, 
usually distinctly pitted. 


Barrens, Kentucky, southern Illinois and Tennessee. May- 
July. 


4. Onosmodium occidentale Mackenzie. 
Western False Gromwell. Fig. 3545. 


O. occidentale Mackenzie, Bull. Torr. Club 32: 


502. 1905. 
O. occidentale sylvestre Mackenzie, loc. cit. 504. 
1905. 


Stem 1°-33° high, branched above or also 
from the base, strigose or hirsute-pubescent. 
Leaves lanceolate to ovate-lanceolate, acute, 
appressed-pubescent on both sides, or the hairs 
somewhat spreading, 2-3’ long, strongly vein- 
ed; bracts similar to the leaves but much 
smaller; calyx-lobes lanceolate, acute to ob- 
tuse, 3-6” long; corolla 6-10” long, canes- 
cent all over outside, its lobes 13’-2” long, 
broadly triangular and acute; nutlets ovoid, 
acutish, about 2” long, dull, scarcely if at all 
pitted, not constricted at the base. 


On prairies and plains, Illinois to North Da- 
kota, Manitoba, Alberta, Kansas, Texas and New 
Mexico. Included in O. molle Michx., in our first 
edition, and by previous authors. May-July. 


5. Onosmodium hispidissimum Mackenzie. 
Shaggy False Gromwell. Fig. 3546. 
Onosmodium hispidissimum Mackenzie, Bull. Torr. Club 

32: 500. 1905. 

Spreading-hirsute with rough bristly hairs; stem 
stout, usually much branched, 1°-4° high. Leaves lan- 
ceolate, ovate-lanceolate or oblong, acute or acuminate 
at the apex, narrowed to the sessile base, 5-9-ribbed, 
2’-43’ long, 4’-1}’ wide; flowers very numerous and 
crowded; pedicels 1’-2”’ long in fruit; calyx-segments 
linear, somewhat shorter than the corolla-tube; corolla 
yellowish-white, pubescent outside, 5-9” long, its lobes 
triangular-lanceolate, acute, one-third to one-half as 
long as the tube; nutlets obtuse, about 13” long, dis- 
tinctly constricted at the base, little if at all pitted. 


In dry fields or thickets, or on banks, Ontario and west- 
ern New York to Minnesota, Missouri, Georgia and Texas. 
Ascends to 2200 ft. in Virginia. Previously referred to 
Onosmodium carolinianum (Lam.) DC. May-July. 


g2 BORAGINACEAE. Vor. III. 


14. SYMPHYTUM [Tourn.] L. Sp. Pl. 136. 1753. 


Erect coarse hairy perennial branching herbs, with thick mucilaginous roots, alternate 
entire leaves, those of the stem mostly clasping, the uppermost tending to be opposite, the 
lower long-petioled. Flowers yellow, blue, or purple, in terminal simple or forked scorpioid 
racemes. Calyx deeply 5-cleft. Corolla tubular, slightly dilated above, 5-toothed or 5-lobed, 
the lobes short, the throat with 5 crests below the lobes. Stamens 5, included, inserted on 
the corolla-tube; filaments slender. Ovary 4-divided; style filiform. Nutlets 4, obliquely 
ovoid, slightly incurved, wrinkled, inserted by their bases on the flat receptacle, the scar of 
the attachment broad, concave, dentate. [Greek, grow-together, from its supposed healing 
virtues. | 

About 15 species, natives of the Old World. Type species: Symphytum officinale L. 


Leaf-bases decurrent. ; 1. S. officinale. — 
Leaf-bases not decurrent, or but slightly decurrent. 2. S. asperrimum, 


1. Symphytum officinale L. Comfrey. Healing-herb. Fig. 3547. 
Symphytum officinale L. Sp. Pl. 136. 1753. 


Roots thick, deep; stem erect, branched, 2°-3° 
high. Leaves lanceolate, ovate-lanceolate, or the 
lower ovate, pinnately veined, 3-10’ long, acute 
or acuminate at the apex, narrowed into mar- 
gined petioles, or the uppermost smaller and 
sessile, decurrent on the stem; petioles of the 
basal leaves sometimes 12’ long; flowers numer- 
ous, in dense racemes or clusters; pedicels 2’—4” 
long; calyx-segments ovate or ovate-lanceolate, 
acute or acuminate, much shorter than the co- 
rolla; corolla yellowish or purplish, 6’—10” long; 
nutlets brown, shining, slightly wrinkled, 2” 
high. 

In waste places, Newfoundland to Minnesota, 
south to Virginia and North Carolina. Naturalized 
or adventive from Europe. Native also of Asia. 
June-Aug. Back- or black-wort. Bruisewort. Knit- 
back. Boneset. Consound. Gum-plant. 

Symphytum tuberésum L., with thickened tuber- 
ous roots, the nutlets granular-tuberculate, not shin- 
ing, has been found in sandy meadows in Con- 
necticut. 


2. Symphytum aspérimum Donn. Rough 
Comfrey. Fig. 3548. 
S. asperrimum Donn; Sims, Bot. Mag. 24: pl. 929. 

1806. 

Similar to S. officinale, but the pubescence 
rougher, the hairs stiff and reflexed. Leaves ovate- 
lanceolate to oblong-lanceolate, long-acuminate at 
the apex, narrowed at the base, all but the upper- 
most petioled, slightly or not at all decurrent, the 
lower often 8’ long; flower-clusters rather loose; 
calyx about half as long as the corolla-tube, its 
segments hispid; corolla bluish-purple. 


Waste grounds, Massachusetts to Maryland. Ad- 
ventive or naturalized from Europe. June—-Aug. 


15. BORAGO [Tonrn: | Sesek ors7-. 17538: 


Hirsute or hispid annual or biennial branching herbs, with alternate entire leaves, and 
showy blue flowers, in terminal loose leafy racemes. Calyx deeply 5-cleft or 5-parted. 
Corolla rotate, the tube very short, the throat closed by scales, the limb 5-lobed, the lobes 
imbricated, acute. Stamens 5, inserted on the corolla-tube; filaments dilated below, narrowed 
above into a slender appendage; anthers linear, erect, and connivent into a cone. Ovary 
4-divided; style filiform. Nutlets 4, ovoid, erect, attached by their bases to the flat receptacle, 
the scar of attachment large, concave. [Middle Latin, burra, rough hair, alluding to the 
foliage. ] 3 

Three species, natives of the Mediterranean region, the following typical. 


GENUS I5. BORAGE FAMILY. 93 


1. Borago officinalis L. Borage. 
Fig. 3549. 
Borago officinalis L. Sp. Pl. 137. 1753. 


Stem erect, branched, 1°-23° high, the 
branches spreading or ascending. Leaves ob- 
long to obovate, acute or obtuse at the apex, 
2’-5’ long, narrowed into margined petioles, 
or the upper smaller, ovate-lanceolate, sessile 
or partly clasping; flowers 8’-10” broad, 
pedicels rather stout, 14-2’ long, spreading or 
recurving; calyx-segments lanceolate, nearly 
erect in fruit; corolla bright blue, the lobes 
ovate-lanceolate; the cone of anthers darker, 
about 3” long; nutlets 2” high. 


In waste places, escaped from gardens, Nova 
Scotia to Ontario and Pennsylvania, but prob- 
ably not persistent within our range. Native of 
southern Europe. June—Sept. 


16. LYCOPSIS L. Sp -Pl. 138. 1753. 


Annual bristly-hispid branched erect or diffuse herbs, with alternate leaves, and small 
blue or bluish flowers, in dense leafy-bracted terminal spike-like scorpioid racemes. Calyx 
5-parted. Corolla slightly irregular, salverform, the tube curved, the limb somewhat unequally 
5-lobed, the lobes obtuse, imbricated, the throat closed by hispid scales. Stamens 5, included, 
inserted on the tube of the corolla; filaments short; anthers obtuse at each end. Ovary 
4-divided; style filiform. Nutlets 4, wrinkled, erect, attached by their bases to the flat recep- 
tacle, the scar of attachment concave. [Greek, wolf-face.] 

About 4 species, natives of the Old World, the following typical. 


pal Y 


1. Lycopsis arvénsis L. Small Bugloss. 
Fig. 3550. 
Lycopsis arvensis L. Sp. Pl. 139. 1753. 


Stem erect or ascending, at length divergently 
or diffusely branched, 1°-2° high, the branches 
becoming procumbent. Leaves lanceolate, nar- 
rowly oblong or the lower oblanceolate, obtuse, 
1-2’ long, undulate or dentate, sessile, or the 
lower narrowed into petioles, the upper much 
smaller and acute or acutish; flowers numerous, 
crowded, 2-3” broad, very  short-pedicelled; 
calyx-segments lanceolate, acute, nearly as long 
as the curved corolla-tube; nutlets shorter than 
the calyx. 

In fields and waste places, Nova Scotia to Ontario, 
Minnesota, Pennsylvania and Virginia. Naturalized 


or adventive from Europe. Native also of Asia. 
June-Sept. 


Dye ECHIUM | Pours. | “Le Sp Pl 136.” 1752: 


Biennial or perennial mostly bristly-hirsute branching herbs, with alternate leaves, and 
rather large blue violet or rarely white flowers, in leafy-bracted scorpioid spikes. Calyx 
5-parted, the segments narrow. Corolla tubular-funnelform, irregular, the limb unequally 
5-lobed, the lobes rounded, spreading, the throat not appendaged. Stamens 5, inserted low 
down on the tube of the corolla, unequal, at least the longer ones exserted; filaments slender, 
dilated at the base; anthers ovate or oblong. Ovary 4-divided; style filiform, 2-cleft at the 
summit. Nutlets 4, erect, ovoid, rugose, attached by their bases to the flat receptacle, the 
scar of attachment not concave. [Greek, a viper.] 


About 30 species, natives of the Old World. Type species: Echium itélicum L. 


94 BORAGINACEAE. Vor Lie 


1. Echium vulgare L. Viper’s Bugloss. Blue- 
weed. Fig. 3551. 


Echium vulgare L. Sp. Pl. 140. 1753. 


Bristly-hairy, biennial; stem erect, at length much 
branched, 1°-23° high. Leaves oblong, linear-oblong, 
or linear-lanceolate, obtuse or acute, entire, 2-6’ long, 
sessile, or the lower and basal ones narrowed into 
petioles; flower-buds pink; flowers bright blue, vary- 
ing to violet purple, 8’-12” long, numerous in short 
I-sided spikes, forming a narrow thyrsus; calyx-seg- 
ments much shorter than the corolla; limb of the 
corolla oblique, the lobes very unequal. 

In fields and waste places, Nova Scotia to North Caro- 
lina, Ontario and Nebraska. A troublesome weed in some 
sections of the North. Naturalized from Europe. Native 
also in Asia. June-July. Viper’s-herb. Viper’s- -grass. 
Snake-flower. Blue thistle. Blue stem or cat’s-tails. Blue 
devils. Adder’s-wort. 


Family 25. VERBENACEAE J. St. Hil. Expos. Fam. 1: 245. 1805. 
VERVAIN FAMILY. 

Herbs, shrubs or some tropical genera trees, with opposite verticillate or rarely 
alternate leaves, and perfect, more or less irregular, or sometimes regular flowers, 
in terminal or axillary spikes, racemes, cymes or panicles. Calyx inferior, mostly 
persistent, usually 4—-5-lobed or 4-—5-cleft. Corolla gamopetalous, regular, or 
2-lipped, the tube usually cylindric and the limb 4-5-cleft. Stamens 4, didy- 
namous, rarely only 2, or as many as the corolla-lobes, inserted on the corolla and 
alternate with its lobes; anthers 2-celled, the sacs longitudinally dehiscent. Ovary 
superior, 2—4-celled (rarely 8—-10-celled), composed of 2 carpels, each carpel with 
2 anatropous or amphitropous ovules, thus in 4-celled ovaries 1 ovule in each 
cavity ; style terminal, simple; stigmas I or 2. Fruit dry, separating at maturity 
into 2-4 nutlets, or a drupe containing the 2-4 nutlets. Endosperm little or none, 
or rarely fleshy ; embryo straight. 

_About 75 genera and 1300 species, of wide geographic distribution in temperate and warm 
regions. 
pipcrs in heads or spikes; ovary 2-4-celled; fruit of 2 or 4 erect nutlets; ours herbs. 

Corolla-limb 5-lobed, regular or nearly so; nutlets 4. 1. Verbena. 


Corolla-limb 4-lobed, 2-lipped; nutlets 2. 2. Lippia. 
Flowers in axillary cymes; shrubs; fruit drupaceous. 3. Callicarpa. 


t. ' VERBENA | Tourn.| L. Sp,-Pl 182 1752 

Herbs (some exotic species shrubby), mostly with opposite leaves, and variously colored 
bracted flowers, in terminal solitary corymbed or panicled spikes. Calyx usually tubular, 
5-angled, more or less unequally 5-toothed. Corolla salverform or funnelform, the tube 
straight or somewhat curved, the limb spreading, 5-lobed, slightly 2-lipped or regular. 
Stamens 4, didynamous, or very rarely only 2, included; connective of the anthers unap- 
pendaged, or sometimes provided with a gland. Ovary 4-celled; ovule I in each cavity; style 
usually short, 2-lobed at the summit, one of the lobes stigmatic. Fruit dry, mostly enclosed 
by the calyx, at length separating into 4, I-seeded linear or linear-oblong crustaceous smooth 
papillose or rugose nutlets. [Latin name of a sacred herb.] 

About 100 species, natives of America, ora single one indigenous in the Mediterranean region. 
Besides the following, some 15 others occur in the southern and western parts of North America. 
Type species: Verbena officinalis L. 

Flowers 2”-s” long, in narrow spikes; anthers unappendaged. 
Spikes filiform or slender; bracts shorter than the flowers. 
Spikes filiform ; fruit scattered ; corolla usually white. 
Leaves incised or pinnatifid; diffuse annual; fruit short. 1. V. officinalis. 
Leaves serrate (rarely incised) ; ; erect perennial ; fruit oblong. 2. V.urticifolia. 
Spikes slender; fruit densely imbricated ; corolla blue. 
Plants glabrous or sparingly rough-pubescent ; corolla 2”—3” long. 


Leaves lanceolate, acuminate, petioled. 
Leaves linear or spatulate-lanceolate, mostly obtuse and sessil 


. V. hastata. 
V. angustifolia. 


Nui w 


Plants densely soft-pubescent; corolla 4’”—5” long. . V. stricta: 
Spikes thick, dense ; bracts longer than the flowers. V.. bracteosa. 
Flowers 7”—12” long,in short dense elongating spikes ; connective of the longer stamens appendaged. 
Corolla-limb 6’—12” broad; bracts mostly shorter than the calyx. 7. V. canadensis. 


Corolla-limb 4”—7” broad ; ‘bracts equalling or exceeding the calyx. 8. V. bipinnatifida. 


GENUS I. VERVAIN FAMILY. 


1. Verbena officinalis L. European Ver- 
vain. Herb-of-the-Cross. Berbine. 


. Fig. 3552. 
Verbena officinalis L. Sp. Pl. 20. 1753. 

Annual ; stem 4-sided, slender, glabrous or nearly 
so, ascending or spreading, diffusely branched, 
1°-3° high. Leaves minutely pubescent, the lower 
deeply incised or I-2 pinnatifid, ovate, oblong, or 
obovate in outline, 1-3’ long, narrowed into mar- 
gined petioles, the teeth acute; upper leaves 
linear or lanceolate, acute, entire, sessile; spikes 
several or numerous, filiform, at length 4’-5’ long; 
fruits less than 1” high, scattered along the spikes, 
not at all imbricated; bracts ovate, acuminate, 
shorter than the 5-toothed calyx; corolla pur- 
plish or white, the limb 1’—2” broad. 

In waste and cultivated ground, Maine to Florida, 
Tennessee and Texas. Also on the Pacific Coast 
and in the West Indies. Naturalized from the Old 
World. Sometimes a troublesome weed. Herb-grace. 
Holy-herb. Enchanter’s-plant. Juno’s-tears. Pigeon’s- 
grass. Simpler’s-joy. June—Sept. 


2. Verbena urticifolia L. White or Nettle-leaved Vervain. 


Verbena urticifolia L. Sp. Pl. 20. 1753. 
V. urticifolia riparia Britton, Mem. Torr. Club 5: 276. 


1894. 
V. riparia Raf.; Small & Heller, Mem. Torr. Club 3: 12. 
1892. 


Perennial, usually pubescent; stem slender, strict, 
erect, 4-sided, paniculately branched above, 3°-5° 
high, the branches upright. Leaves ovate, oblong, 
or oblong-lanceolate, all petioled, or the uppermost 
sessile, serrate-dentate all around, or incised, some- 
times 3-cleft near the base, thin, acute or acuminate, 
mostly rounded at the base, 13-5’ long; spikes nu- 
merous, filiform, erect, or spreading, at length 4-6’ 
long; fruits oblong, scattered, not at all imbricated, 
about 1” high; bracts ovate, acuminate, shorter than 
the calyx; corolla white, blue or pale purple, its limb 
about 1” broad. 


In fields and waste places, New Brunswick to South 
Dakota, Kansas, Florida and Texas. MHybridizes with 
V. bracteosa, V. hastata and V. stricta. June—Sept. 


Verbena carolinénsis (Walt.) Gmel., with sessile, 
spatulate to oblong leaves and larger bluish flowers, 
native of the Southeastern States, is recorded as occur- 
ring north to Virginia. 


3. Verbena hastata L. Blue or False Ver- 
vain. Wild Hyssop. Fig. 3554. 

Verbena hastata L. Sp. Pl. 20. 1753. 

Verbena pinnatifida Lam. Tabl. Encycl. 1: 57. 1791. 

Verbena paniculata Lam. Encycl. 8: 548. 1808. 

Verbena hastata pinnatifida Britton, Mem. Torr. Club 

5: 276. 1894. 

Perennial, roughish-puberulent ; stem erect, strict, 
4-sided, usually branched above, 3°-7° high. Leaves 
oblong-lanceolate or lanceolate, petioled, acute or 
acuminate at the apex, narrowed at the base, ser- 
rate or incised-dentate with acute teeth, sometimes 
pinnatifid, 3-6’ long, the lower sometimes hastately 
3-lobed at the base; spikes numerous, panicled, slen- 
der, usually peduncled, 2’-6’ long; fruits densely 
imbricated on the spikes, 1-14” high; bracts ovate, 
acuminate, shorter than the calyx; corolla blue, 
white, or sometimes pink, its limb about 14” broad. 

In moist fields, meadows and in waste places, Nova 
Scotia to British Columbia, Florida, Nebraska and Ari- 
zona. Hybridizes with V. stricta and V. bracteosa. 
American vervain. Purvain. Iron-weed. June-Sept. 


4. 


V~. 


iE 


th 


sp 


V. 


Mullen-leaved Vervain. Fig. 3556. 
V. stricta Vent. Descr. Pl. Jard. Cels. pl. 53. 1800. 
Verbena rigens Michx. Fl. Bor. Am. 2: 14. 1803. 


Perennial, densely soft-pubescent all over 
with whitish hairs; stem stout, obtusely 4-an- 
gled, simple, or branched above, strict, very 
leafy, 1°-23° high. Leaves ovate, oval, or 
oblong, very short-petioled, acute or obtuse 
at the apex, narrowed at the base, promi- 
nently veined, incised-serrate or laciniate, 1’—4’ 
long; spikes solitary, or several, mostly sessile, 
dense, stout, becoming 6-12’ long in fruit; 
fruits much imbricated, 2”-23” high; bracts 
lanceolate-subulate, nearly as long as the ca- 
lyx; corolla purplish blue, 4”-5” long, its limb 
nearly as broad. 


In dry soil, Ontario and Ohio to Minnesota, 
South Dakota and Wyoming, south to Tennessee, 
Texas and New Mexico. Naturalized as a weed 
euunes east. Hybridizes with VY. bracteosa. June- 

ept. : 


5 cae 


5 
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ae 


<a 
YESS 


eo, ce 


G 


ot 


BAS 
S 

NS 

\ aS 
VN" 


fg 7 
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WUALA Bg 
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“WA SY 
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\ 


VERBENACEAE. 


Minnesota, Kansas and Arkansas. 


Vor. III. 


Verbena angustifolia Michx. Narrow- 
leaved Vervain. Fig. 3555. 
angustifolia Michx. Fl. Bor. Am. 2: 14. 1803. 


Perennial, roughish-puberulent or pubescent} 


stem slender, simple or branched, 4-sided above, 


-2° high. Leaves linear, spatulate or lanceo- 


late, obtuse or subacute at the apex, cuneate at 


e base and tapering into short petioles, serrate 


or serrulate, veiny, 12-3’ long, 2’-5” wide; 


ikes mostly solitary at the ends of the branches, 


usually peduncled, slender, dense, 2-5’ long; 
fruits overlapping or the lower somewhat dis- 
tant, 
equalling or shorter than the calyx; corolla pur- 
ple or blue, about 3” long, its limb about as 
broad, the lobes obovate or oblong. 


1%” high; bracts lanceolate, acuminate, 


In dry fields, Massachusetts to Florida, west to 
Hybridizes with 
June-Aug. 


stricta and V. bracteosa. 


6. Verbena bractedsa Michx. Large- 
bracted Vervain. Fig. 3557. 


V. bracteosa Michx. Fl. Bor. Am. 2: 13. 1803. 


Perennial, hirsute-pubescent;. stem 4-sided, 
much branched from the base, the branches 
decumbent or ascending, slender, 6-15’ long. 
Leaves ovate, oval, or obovate in outline, pin- 
nately incised or pinnatifid, 1-3’ long, more 
or less cuneate at the base and narrowed into 
short petioles, the lobes mostly dentate; spikes 
sessile, stout, dense, becoming 4-6’ long in 
fruit; bracts conspicuous, linear-lanceolate, 
rather rigid, longer than the flowers and fruits, 
the lower ones often incised; corolla purplish 
blue, about 2” long. 


On prairies and in waste places, Minnesota and 
Illinois to Virginia, Alabama and Florida, west to 
British Columbia, Arizona and California. In 
ballast grounds at Atlantic seaports. Hybridizes 
with V. canadensis. Prostrate vervain. May—Aug. 


GENUS I. VERVAIN FAMILY. 97 


7. Verbena canadénsis (L.) Britton. 

Large-flowered Verbena. Fig. 3558. 
Buchnera canadensis L. Mant. 88. 1767. 

V. Aubletia Jacq. Hort. V. 2: 82. pl. 176. 1772. 
powarte carolinensis J. G. Gmel. Syst. 2: 920. 

I ° 
Berbera canadensis Britton, Mem. Torr. Club 

5: 276. 1804. 

Perennial, pubescent or glabrate; stem slen- 
der, usually branched, 8-20’ high, the branches 
ascending. Leaves membranous, ovate in out- 
line, petioled, 1-3’ long, truncate or broadly 
cuneate at the base, irregularly toothed, or 
pinnately incised, often 3-cleft, the lobes den- 
tate; spikes peduncled, solitary at the ends 
of the branches, dense, short and capitate 
when in early flower, becoming 2-4’ long in 
fruit; bracts linear-subulate, mostly shorter 
than the calyx; calyx-teeth filiform-subulate ; 
corolla 10-12” long, blue, purple, white or 
in cultivation variegated, its limb 7-12” 
broad, the lobes oblong or obovate, emargi- 
nate or obcordate; fruit 23’-3” high. 

In dry soil, Illinois to Tennessee, Virginia and 
Florida, west to Kansas and Texas. This and 
the next the source of many garden and other 
hybrids.. Cut-leaved races have been referred 
to V. Drummondii (Lindl.) Baxter. May—Aug. 


——— 


eer ae eee 


SqSSSSSSSSEES 


8. Verbena bipinnatifida Nutt. Small- 
flowered Verbena. Fig. 3559. 


Verbena bipinnatifida Nutt. Journ. Acad. Phila. 
27 tes toot. 

Glandularia bipinnatifida Nutt. Trans. Am. Phil. 
Soc, (iI) 5% 28465 1933—-37- 

Verbena ambrosiaefolia Rydb.; Small, Fl. SE. 
Ue Se rom. "19035 


Perennial, producing suckers, hirsute or 
hispid; stems rather stout, mostly branched, 
erect, 6-18’ high. Leaves firm, petioled or 
the uppermost sessile, broadly ovate in out- 
line, deeply 1-2-pinnatifid into linear or linear- 
NA y oblong, obtuse or subacute lobes and_seg- 
MWR VG, 2 \ iy ‘ ments; spikes peduncled or sessile, solitary 
XX YL E / at the ends of the branches, thick, dense, at 

N first short and capitate, becoming 2-4’ long 

} in fruit; bracts linear-subulate, about as long 
as or somewhat exceeding the calyx; calyx- 
teeth filiform-subulate; corolla 6-9” long, 
purple or lilac, the limb 4’-7” broad, the lobes 
emarginate or obcordate; fruit 14’—2” long. 

On dry plains and prairies, South Dakota to 
Missouri, Texas and Chihuahua, west to Colo- 
rado and Arizona. May-—Sept. 


2. LIPPIA Houst.; L. Sp. Pl. 633. 1753. 


Perennial herbs, or shrubs, with opposite, sometimes verticillate, or rarely alternate leaves, 
and small bracted flowers, in axillary or terminal, mostly peduncled spikes or heads. Calyx 
small, membranous, ovoid, campanulate or compressed and 2-winged, 2-4-toothed or 2-4-cleft. 
Corolla-tube straight or incurved, cylindric, the limb oblique, spreading, somewhat 2-lipped, 
4-cleft, the lobes broad, often retuse or eroded. Stamens 4, didynamous, included or exserted ; 
anthers ovate, not appendaged, the sacs nearly parallel. Ovary 2-celled; ovules I in each 
cavity; style short; stigma oblique or recurved. Fruit dry, with a membranous exocarp, at 
length separating into 4 nutlets. [In honor of Auguste Lippi, 1678-1703, French naturalist. ] 

About 110 species, most abundant in tropical and subtropical America, a few African. Besides 
the following, which by some authors are separated as a distinct genus (PHyLa Lour.), about 6 
others occur in the southern United States. Type species: Lippia americana L. 


Leaves linear-cuneate to spatulate, 2-8-toothed ; peduncles little exceeding leaves. 1. L. cuneifolia. 
Leaves sharply serrate ; peduncles much longer than leaves. 
Leaves oblong or-lanceolate, mostly acute. 2. L. lanceolata. 
Leaves spatulate or obovate, mostly obtuse. 3. L. nodiflora. 


VERBENACEAE. VoL. IIT. 


1. Lippia cuneifolia (Torr.) Steud. Wedge- 
leaved Fog-fruit. Fig. 3560. 


\ pale cuneifolia Torr. Ann, Lyc. N. Y. 2: 234. 

1827. 

Lippia cuneifolia Steud.; Torr. in Marcy’s Rep. 293. 
Pls i722 1853. 


. Pale, minutely puberulent with forked hairs or 
glabrous, diffusely branched from the woody base; 
branches terete, slender, rigid, procumbent, some- 
what zigzag, with short erect branchlets at the 
nodes. Leaves linear-cuneate, sessile, obscurely 
veined, rigid, 1’-12’ long, 2-3” wide, with 2-8 
sharp teeth above the middle or rarely entire, 
acutish at the apex; peduncles shorter than or 
somewhat exceeding the leaves; head at first 
globose, becoming cylindric and 6-8” long; 
bracts cuneate, abruptly acuminate from the trun- 
cate or retuse summit; calyx flattened, 2-cleft, the 
lobes 2-toothed or emarginate; corolla-tube longer 
than the calyx; fruit oblong. 


On plains, South Dakota, Nebraska and Colorado 
to Texas, Mexico and Arizona. May—Aug. 


2. Lippia lanceolata Michx. Fog-fruit. 
Fig. 3561. 
L. lanceolata Michx. Fl. Bor. Am. 2: 15. 1803. 


Green, glabrous, or very sparingly pubes- 
cent with forked hairs; stems slender, weak, 
procumbent or ascending, sometimes rooting 
at the nodes, simple, or little branched, 1°-2° 
long. Leaves thin, oblong, ovate, or oblong- 
lanceolate, pinnately veined, short-petioled, 
acute or subacute at the apex, sharply serrate 
to below the middle, narrowed to the some- 
what cuneate base, 1-3’ long, 3-15” wide; 
peduncles slender, some or all of them longer 
than the leaves; heads at first globose, becam- 
ing cylindric and about 2’ long in fruit; bracts 
acute; calyx flattened, 2-cleft; corolla pale 
blue, scarcely longer than the calyx; fruit 
globose. 


In moist soil, Ontario to Minnesota, New Jer- 
sey, Illinois, Kansas, Florida, Texas and north- 
ern Mexico. Also in California. Frog-fruit. 
June-Aug. 


3. Lippia nodiflora (L.) Michx. Spatu- 
late-leaved Fog-fruit. Fig. 3562. 

Verbena nodiflora L. Sp. Pl. 20. 1753. 

Lippia nodiflora Michx FI. Bor. Am. 2: 15. 1803. 


Minutely and rather densely puberulent with 
short appressed hairs, creeping, or some of the 
branches ascending, 1°-3° long. Leaves thickish, 
spatulate, cblanceolate, or obovate, 6’—22’ long, 
3-12” wide, mostly obtuse at the apex, narrowed 
into a long or short cuneate entire base, sharply 
serrate above the middle; peduncles slender, 1’-6’ 
long, much longer than the leaves; heads at length 
cylindric and 5-12” long, 3-4” thick; calyx flat- 
tened, 2-cleft; corolla purple to white. 


In wet or moist soil, South Carolina to southern 
Missouri, Florida and Texas. Also in California, 
Central America, the West Indies, and apparently 
the same species in the warmer regions of the Old 
World. May-—Sept. 


GENUS 3. VERVAIN FAMILY. oe) 


3. CALLICARPA L. Sp. Pl. 111. 1753. 


Shrubs or trees, with opposite leaves, and small blue purple or white flowers in axillary 
cymes. Calyx short, campanulate, 4-toothed (rarely 5-toothed), or truncate. Corolla-tube 
short, expanded above, the limb spreading, 4-cleft (rarely 5-cleft), the lobes equal, imbri- 
cated in the bud. Stamens 4, equal, exserted; anthers ovate or oval, their sacs parallel. 
Ovary incompletely 2-celled; ovules 2 in each cavity, laterally attached, amphitropous; style 
slender; stigma capitate, or 2-lobed. Fruit a berry-like drupe, much longer than the calyx, 
containing I-4 nutlets. [Greek, handsome fruit.] 


About 45 species, the following typical one of southeastern North America, the others Asiatic, 
African and tropical American. 


1. Callicarpa americana L. French or 
Bermuda Mulberry. Fig. 3563. 


Callicarpa americana L. Sp. Pl. 111. 1753. 


A shrub, 2°-5° high, the twigs, petioles and 
young leaves stellate-scurfy, the mature leaves 
becoming nearly glabrous and glandular-dotted. 
Twigs terete; leaves thin, ovate, pinnately veined, 
slender-petioled, acute or acuminate at the apex, 
crenate-dentate nearly to the entire base, 3-6 
long, 12-3’ wide; cymes many-flowered, short- 
peduneled; pedicels very short; calyx-teeth much 
shorter than the tube; corolla pale blue, about 
14” long; fruit violet-blue, globose, 14” in diam- 
eter, very conspicuous in autumn. 


In moist thickets, Virginia to Florida, Missouri, 
Arkansas and Texas. Bermuda. June-July. Sour- 
bush. 


Callicarpa purpurea Juss., an Asiatic shrub often 
planted for ornament, with pink flowers and long 
leaves, has been observed in a swamp at Wilmington, 
Delaware. 


Family 26. LABIATAE B. Juss. Hort. Trian. 1750. 
Mint FAmMILy. 


Aromatic punctate herbs, or shrubs (a few tropical species trees), mostly with 
4-sided stems and simple opposite leaves; stipulesnone. Flowers irregular, perfect, 
variously clustered, the inflorescence typically cymose, usually bracteolate. Calyx 
inferior, persistent, regular or 2-lipped, 5-toothed or 5-lobed (rarely 4-toothed), 
mostly nerved. Corolla with a short or long tube, the limb 4-5-lobed, mostly 
2-lipped, regular in a few genera; upper lip 2-lobed, or sometimes entire; lower 
lip mostly 3-lobed. Stamens borne on the corolla-tube, typically 4 and didy- 
namous, sometimes 2 with or without staminodia, rarely equal ; filaments separate, 
mostly slender, alternate with the corolla-lobes; anthers 2-celled, introrse, or con- 
fluently 1-celled, or sometimes of a single sac. Disk usually present, fleshy. 
Ovary 4-lobed or 4-parted, superior, each lobe or division with 1 mostly anatropous 
ovule; style arising from the center of the lobed or parted ovary, 2-lobed at the 
summit. Fruit of 4 1-seeded nutlets. Seed erect (transverse in Scutellaria) ; 
endosperm scanty, or usually none; embryo mostly straight ; radicle short, inferior. 


About 160 genera and 3200 species, of wide distribution in temperate and tropical regions. The 
family is also known as LAMIACEAE. The foliage abounds in volatile oils. 


A. Ovary 4-lobed, the style not basal; nutlets laterally attached. (AyucEar.) 
Corolla-limb very irregular, apparently 1-lipped, or the other lip very short; stamens exserted, 


Upper lip of corolla short, truncate. 1. Ajuga. 

Upper lip of corolla 2-lobed, or all the lobes united into the lower lip. 2. Teucrium. 
Corolla-limb nearly equally 5-lobed. 

Corolla-lobes spreading; stamens short-exserted. 3. Isanthus. 

Corolla-lobes declined ; stamens long-exserted. 4. Trichostema. 


B. Ovary 4-parted, the style basal; nutlets basally attached. 
* Calyx with a protuberance on the upper side. (ScuTELLARIEAE.) 5. Scutellaria. 
** Calyx not gibbous on the upper side. (STACHYEAE.) 
Stamens and style very short, included in the corolla-tube. 
Anther-sacs parellel. 6. Marrubium. 
Anther-sacs, at least of the upper stamens, divergent. 7. Sideritis. 
Stamens longer, not included in the corolla-tube. 


100 LABIATAE. 


{ Corolla strongly 2-lipped ; lips unlike, the upper concave. 


(a.) Anther-bearing stamens 4. 


Posterior (upper) pair of stamens longer than the anterior. : 
Anther-sacs parallel or nearly so. 
Tall erect herbs; posterior stamens declined, anterior ascending. 
Trailing herb; stamens all ascending under upper lip of corolla. 
Anther-sacs divergent. 
Calyx tubular, nearly equally 5-toothed, not 2-lipped; plant erect. 
Calyx distinctly 2-lipped, or unequally 5-toothed. 
Trailing herb; calyx unequally 5-toothed. 
Erect herbs; calyx 2-lipped. 
Posterior pair of stamens shorter than the anterior. 
Calyx distinctly 2-lipped, closed in fruit. 
Calyx 3-10-toothed, not distinctly 2-lipped, open in fruit. 
Calyx membranous, inflated in fruit, faintly nerved. 
Calyx nearly equally 5-toothed, or 5-lobed. 
Calyx 4-lobed. 


Calyx not membranous, not inflated in fruit, distinctly 5—1o-nerved. 


Style-branches very unequal. 
Style-branches equal, or nearly so. 
Anther-sacs transversely 2-valved. 
Anther-sacs not transversely 2-valved, parallel or divergent. 
Nutlets 3-sided, truncate. 
Calyx-teeth not spiny-tipped. 
Calyx-teeth spiny-tipped. 
Nutlets ovoid, rounded above. 
Calyx with a spreading 5-toothed limb. 
Calyx-limb not spreading. 


Corolla-tube not longer than calyx; anther-sacs divergent. 


21. 


Voi. ITI. 


Agastache. 
Meehania. 


. Nepeta. 


. Glecoma. 
. Moldavica. ” 


. Prunella, 

. Dracocephalum. 
. Synandra. 

. Phlomis. 

. Galeopsis. 

. Lamium. 

. Leonurus. 


. Ballota. 


Stachys. 


Corolla-tube exserted ; lower petioles very long; anther-sacs parallel. 


(b.) Anther-bearing stamens 2. 


22. 


Betonica. 


Connective of the anther very long, bearing a perfect sac at one end, and a rudimentary one, or 


none, at the other. 
Connective very short, the anther-sacs confluent. 
Calyx tubular, 15-nerved, equally 5-toothed. 
Calyx ovoid-tubular, 13-nerved, 2-lipped. 


Be 


24. 
25. Blephilia. 


Salvia. 


Monarda. 


+7 Corolla 2-lipped, or regular; upper lip, when present, flat, or only slightly concave. 


(a.) Flowers in axillary whorls or clusters, or these forming terminal spikes. 


Corolla 2-lipped. 


Stamens curved, often converging, or ascending under the upper lip of the corolla. 


Anther-bearing stamens 2. 
Anther-bearing stamens 4. 
Corolla-tube upwardly curved, exserted. 
Corolla-tube straight. 
Calyx 1o-nerved, campanulate, about equally 5-toothed. 
Calyx mostly 13-nerved, tubular, 2-lipped. 
Stamens straight, often diverging. 
Calyx 15-nerved. 
Calyx 1o-13-nerved. 
Anther-bearing stamens 4. 
Anther-sacs divergent. 
Calyx equally 5-toothed; erect herbs. 
Calyx 2-lipped ; creeping herbs. 
Anther-sacs parallel. 
Anther-bearing stamens 2. 
Corolla regular, 4—-5-lobed. 
Anther-bearing stamens 2; plants not aromatic. 
Anther-bearing stamens 4; aromatic fragrant herbs. 


26. 


yf 


28. 
29. 


30. 


Hedeoma. 
Melissa. 


Satureia. 
Clinopodium. 


Hyssopus. 


. Origanum. 
. Thymus. 
. Koellia. 


Cunila. 


. Lycopus. 
. Mentha. 


(b.) Flowers in terminal panicled racemes or spikes; corolla 2-lipped. 


Anther-bearing stamens 2; lower lip of corolla long, fimbriate ; native. 
Anther-bearing stamens 4; lower lip of corolla not fimbriate; introduced. 
Flowers racemose. 
Flowers densely spiked. 


1. AJUGA L. Sp. Pl. 561. 1753. 


37- 


38. 
39. Elsholtzia. 


Collinsonia. 


Perilla. 


Annual or perennial, often stoloniferous herbs, mostly with dentate leaves, and rather 
large verticillate-clustered flowers in terminal spikes, or in the upper axils. 
campanulate, 1o-many-nerved, 5-toothed or 5-lobed, the teeth or lobes nearly equal. Corolla- 
limb 2-lipped, the upper lip short, truncate or emarginate, the lower spreading, with 2 small 


lateral lobes and a much larger emarginate or 2-cleft middle one. 


Calyx ovoid or 


Stamens 4, didynamous, 


somewhat exserted beyond the upper lip of the corolla, the anterior pair the longer; anther- 


GENUS I. MINT FAMILY. IOI 


sacs divergent, only slightly confluent at the base. Ovary not deeply 4-lobed. Nutlets obovoid, 
rugose-reticulate. [Greek, without a yoke; from the seeming absence of the upper lip of 
the corolla. | 

About 40 species, natives of the Old World. Type species: Ajuga reptans L. 


Sparingly pubescent, or glabrous, stoloniferous. 1. A. reptans. 
Pubescent with long hairs, not stoloniferous. 2. A. genevensis. 


1. Ajuga réptans L. Bugle. 


Ajuga reptans L. Sp. Pl. 561. 1753. 


Perennial, sparingly pubescent or glabrous, 
producing slender creeping stolons sometimes 
1° long; stem erect, rather stout, 6—15’ tall. 
Basal leaves tufted, obovate, rounded at the 
apex, crenate or undulate, 1-3’ long, tapering 
into margined petioles; leaves of the stem ob- 
long or oblanceolate, much smaller, sessile or 
. nearly so, those of the stolons mostly petioled; 
upper flower-clusters often forming a short 
spike, the lower commonly distant and axil- 
lary; corolla blue or nearly white, about 2’ 
long. 

In fields, Quebec and Maine to southern New 
York, locally naturalized from Europe. Brown 


bugle. Middle comfrey. Carpenter’s-herb. Sickle- 
wort. May-June. 


2. Ajuga genevénsis L. Erect Bugle. 
Fig. 3565. 
Ajuga genevensis L. Sp. Pl. 561. 1753. 


Perennial, not stoloniferous; stems usually sev- 
eral from the root, ascending, rather stout, long- 
pubescent. Leaves long-pubescent, the basal ones 
long-petioled, spatulate or oblanceolate, obtuse, 
sparingly toothed or entire, 2-4’ long, the upper 
sessile, obovate to ovate, usually coarsely den- 
tate, much smaller; bracts pubescent, the upper 
shorter than the flowers; inflorescence mostly 
dense; corolla 4’—3’ long. 

In fields and waste grounds, Maine to New York 
and Pennsylvania. Adventive from Europe. May- 


2. TEUCRIUM [Tourn.] L. Sp. Pl. 562. 1753. 


Herbs or shrubs, with dentate entire or laciniate leaves, and rather small pink, white or 
purplish flowers, in terminal bracted spikes or heads, or verticillate in the upper axils. Calyx 
tubular-campanulate, 10-nerved, equally or unequally 5-toothed. Corolla-tube short, the limb 
irregularly 5-lobed, the 2 short upper lobes oblong, declined or erect, the lateral lobe declined, 
more or less united with the upper ones, the lower lobe broader, also declined. Stamens 4, 
didynamous, exserted between the 2 upper lobes of the corolla, the anterior pair the longer; 
anther-sacs divergent, confluent at the base. Ovary 4-lobed; style 2-cleft at the summit. 
Nutlets obovoid, rugose-reticulated. [Named from the Trojan king, Teucer.] 

Over 100 species, of wide distribution in temperate and tropical regions. Besides the follow- 


ing, 2 or 3 others occur in the southern and southwestern United States. Type species: Teucrium 
friticans L. 


102 LABIATAE. 


VoL. tie 


*PERENNIAL SPECIES. 


1. Leaves toothed. 


7 Flowers in terminal dense spike-like panicles. 
Calyx and bracts canescent, without long hairs. 


Leaves, at least the lower, obtuse or ro 


is 
Leaves narrowed at the base, rugose-veined, mostly narrowly lanceolate. 


Calyx and bracts villous and often glandular-pubescent. 


77 Flowers in secund terminal spikes. 
Leaves laciniate ; flowers axillary. 


iS) 


** ANNUAL SPECIES, with pinnatifid leaves and axillary flowers. 


PO Ag 
iv ALN fff, 
~ Ks 
ISS 


Dy, 


ESV h. 
ence 


2. Teucrium littorale Bicknell. Narrow-leaved Germander. 


Teucrium littorale Bicknell, Bull. Torr. Club 28: 
169. Igor. 


T. canadense var. littorale Fernald, Rhodora 10: 
84. 1908. 


Pale and canescent, 2° high or less, erect or 
assurgent, often with ascending branches. 
Leaves thickish and rugose-veiny, narrowly ob- 
long or sometimes broader, narrowed into the 
petiole, closely fine-serrate or becoming un- 
equally dentate-serrate, 23-4’ long, 2’-13’ wide; 
petioles 23-5” long; spikes narrow, often in- 
terrupted; bracts about the length of the ca- 
lyx; calyx small, 2’—23” high, becoming some- 
what gibbous-urceolate, the teeth short, the 
upper ones obtuse; corolla pale pink, about 
8” long, loosely pilose without. 


On or near the coast, Maine to Florida and 
Texas, north to Arkansas and Oklahoma. In- 
cluded in our first edition in T. canadense L., and 
there figured for that species. July—Aug. 


unded at the base, not rugose-veined. 
T. canadense. 


2. T. littoraie. 

3. T. occidentale. 
4. T. Scorodonia. 
5. I. laciniatum. 
6 


. I. Botrys. 


1. Teucrium canadénse L. American 
Germander or Wood Sage. Fig. 3566. 


Teucrium canadense L. Sp. Pl. 564. 1753. 


Teucrium virginicum L. Sp. Pl. 564. 1753. 


Perennial, appressed-pubescent or canescent; 
stem erect, simple or somewhat branched, 
rather slender, 1°-2° tall. Leaves lanceolate, 
oblong-lanceolate or ovate-lanceolate, acumi- 
nate at the apex, irregularly dentate, mostly 
rounded at the base, short-petioled, 13-5’ 
long, 3’-2’ wide, glabrous or sparingly pubes- 
cent above, densely canescent beneath; spike 
usually dense, becoming 6-12’ long in fruit, 
bracts canescent, the lower sometimes folia- 
ceous, the upper commonly not longer than the 
canescent calyx; flowers 6’-10” long, very 
short-pedicelled; calyx about 3” long in fruit, 
its three upper teeth obtuse or subacute. 


In moist thickets or along marshes, New Eng- 
land to Ontario, Minnesota, Florida, Kansas and 
Texas. Ascends to 2600 ft. in Virginia. Ground- 
pine. June—Sept. 


Fig. 3567. 


GENUS 2. MINT FAMILY. 103 


3. Teucrium occidentale A. Gray. Hairy W 
Germander. Fig. 3568. i 
y 


Teucrium occidentale A. Gray, Syn. Fl. 2: 349. 1878. 


\ 
\ 
- T. boreale Bicknell, Bull. Torr. Club 28: 171. 1901. Si 
AY 


Perennial, villous or pubescent; stem erect, rather HN 
ASN 
Was 


stout, usually much branched, 1°-3° high, the branches 
ascending. Leaves lanceolate or ovate-lanceolate, 
thin, acute or acuminate at the apex, sharply den- 
tate, mostly rounded at the base, usually slender- 
petioled, 1’—32’ long, 2’-14’ wide; spikes dense, be- 
coming 3-8’ long in fruit; bracts lanceolate-subu- 
late or the lower sometimes larger, villous and often 
glandular; calyx and axis of the spike villous- 
pubescent and often glandular, the 3 upper calyx- 
teeth acute or acutish; corolla 4-6” long. 


In moist soil, Maine and Ontario to eastern Pennsyl- 
vania, British Columbia, Ohio, Nebraska, New Mexico 
and California. July—Sept. 


4. Teucrium Scorodénia L. Wood Germander. 
Fig. 35609. 
Teucrium Scorodonia L. Sp. Pl. 564. 1753. 


Perennial, villous-pubescent, 2° high or less, the branches erect- 
ascending. Leaves mostly ovate, 1-23’ long, obtuse or acute at 
the apex, cordate or subtruncate at the base, crenate, the petioles 
23-7” long; racemes narrow, rather loosely flowered, often 5’ 
long, the flowers secund, mostly in pairs, the pedicels shorter 
than the calyx, equalling or shorter than the ovate, acute or 
acuminate bracts; calyx veiny, the upper tooth broad; corolla 
light yellow, 3-4” long. 


A weed in cultivated fields, Ontario, and reported from Ohio. Ad- 
yentive from Europe. June-Sept. 


5. Teucrium laciniatum Torr. Cut-leaved 
Germander. Fig. 3570. 


Teucrium laciniatum Torr. Ann. Lyc. N. Y. 2: 231. 
1828. 


Melosmon laciniatum Small, Fl. SE. U. S. 10109. 

1903. 

Diffusely branched from a woody perennial 
root, 1° high or less, densely leafy, glabrous, or 
nearly so. Leaves 13’ long or less, pinnately 
parted into 3-7 stiff, linear, entire, toothed or 
lobed segments; flowers solitary in the upper 
axils, as long as the subtending leaves or shorter, 
short-peduncled; calyx deeply 5-parted, the lobes 
narrowly lanceolate, nearly equal; corolla pale 
blue or lilac, about 9” long, its lower lobes much 
longer than the calyx. 


Plains, Kansas and Colorado to Texas and Ari- 
zona. May-Aug. 


104 LABIATAE. Vor. ITT. 


sa y 6. Teucrium botrys L. Cut-leaved Annual Ger- 
Ae mander. Fig. 3571. 
Zi \p Teucrium botrys L. Sp. Pl. 562. 1753. 
aa Annual, villous-pubescent, branched, 1° high or less. 


Leaves slender-petioled, deeply pinnatifid into oblong, 
entire or toothed lobes, the basal ones with petioles 
longer than the blades; flowers whorled in the upper 
axils; pedicels shorter than the calyx; calyx campanu- 
late, gibbous, about 8” long in fruit, veiny, its teeth 
triangular-ovate, acute, nearly equal; corolla purplish, 
about 6” long. 


Waste grounds, Massachusetts and Ohio. Naturalized 
from Europe. July—Sept. 


3. ISANTHUS Michx. Fl. Bor. Am. 2: 3. 
PLO. WESOR: 


An annual erect finely viscid-pubescent much-branched 
herb, with narrow entire or few-toothed leaves, and 
small blue flowers in loose axillary cymes. Calyx 
broadly campanulate, 10-nerved, nearly equally 5-toothed, 
the teeth lanceolate. Corolla-tube not longer than the 
calyx, enlarged into the throat, the limb nearly equally 
5-cleft into obovate somewhat spreading lobes. Stamens 4, didynamous, incurved-ascending, 
not longer than the corolla, the anterior pair slightly the longer; anther-sacs divergent at 
maturity. Ovary deeply 4-lobed; style minutely 2-cleft at the summit. Nutlets rugose- 
reticulated. [Greek, equal-flower, the corolla-lobes being nearly equal.] 

A monotypic genus of eastern North America. 


1. Isanthus brachiatus (L.) B.S.P. 
False Pennyroyal. Fig. 3572. 


Trichostema brachiatum L. Sp. Pl. 598. 1753. 

Tsanthus coeruleus Michx. Fl. Bor. Am, 2: 4. 
pl. 30. 1803. 

I. brachiatus B.S.P. Prel. Cat. N. Y. 44. 1888. 


Stem slender, much branched, 620’ tall, 
the branches spreading. Leaves oblong or 
elliptic-lanceolate, acute at each end, entire, 
or with a few sharp teeth, 3-nerved, short- 
petioled, 1-2’ long, 2”-6” wide; axillary 
cymes I—3-flowered; pedicels very slender, 
some of themas long as the fruiting calyx; 
calyx-lobes acute or acuminate, longer than 
or equalling the tube; corolla 2’—3” long, 
the fruiting calyx 3” long. 


In sandy soil, especially along streams, 
Quebec and Ontario to Minnesota, Vermont, 
Georgia, Kansas and Texas. Flux-weed. Blue 
gentian. July—Sept. 


4. TRICHOSTEMA [Gronov.] L. 
Sp. Plisgs. 4753: : 


Annual or perennial erect branching herbs, some western species shrubby, with lanceolate 
oblong or linear entire or slightly repand leaves. Flowers small, or middle-sized, pink, blue, 
purple, or white, paniculate, or in axillary loose or dense cymes. Calyx campanulate, very 
unequally 5-lobed in our species, the lobes ovate or lanceolate, the 3 upper much longer than 
the 2 lower. Corolla-tube slender, exserted or included, the limb somewhat oblique and 
deeply 5-cleft into oblong more or less declined segments. Stamens 4, didynamous, ascend- 
ing, curved, the anterior pair the longer, the filaments filiform, spirally coiled in the bud, 
long-exserted; anther-sacs divaricate, more or less confluent at the base. Ovary deeply 
4-lobed; style 2-cleft at the summit. Nutlets obovoid, reticulated. [Greek, hair-stamen, 
referring to the slender filaments. ] . 

About 10 species, natives of North America. Type species: Trichostema dichotomum L. 


Leaves oblong or lanceolate; plant minutely viscid-pubescent. 1. T. dichotomum. 
Leaves linear; plant puberulent or glabrous. 2. T. lineare. 


GENUS 4. MINT FAMILY. 105 


1. Trichostema dich6tomum L. Blue Curls. Bastard Pennyroyal. Fig. 3573. 
Trichostema dichotomum L. Sp. Pl. 598. 1753. am 

Annual, minutely yiscid-pubescent; stem slen- ” 
der, rather stiff, much branched, 6’-2° high, the v4 \\ 
branches spreading or ascending. Leaves oblong 
or oblong-lanceolate, membranous, obtuse or sub- 
acute at the apex, narrowed at the base into short 
petioles, 1-3’ long, 3-10” wide, the upper grad- 
ually smaller; flowers paniculate, 6’-9” long, 
borne I-3 together on 2-bracteolate peduncles; 
calyx oblique, very unequally 5-lobed, the 3 upper 
lobes much longer and more united than the 2 
lower ones; corolla blue, pink or rarely nearly 
white, the limb longer than the tube; stamens 
blue or violet. 

In dry fields, Maine to Florida, Vermont, Pennsyl- 
vania, Missouri and Texas. The lateral flowers be- 
come inverted by torsion of the pedicels. July—Oct. 

ah & 


2. Trichostema lineare Nutt. Narrow-leaved 
Blue Curls. Fig. 3574. 


T. brachiatum Lam. Encycl. 8: 84. 1808. Not L. 1753. 
Trichostema lineare Nutt. Gen. 2: 39. 1818. 


Puberulent or glabrous, not viscid or scarcely so; 
stem very slender, at length widely branched, 6-18 
high, the branches ascending. Leaves linear, obtuse 
or subacute, sessile or very short-petioled, 4’—2’ long, 
1-2” wide, sometimes with smaller ones or short 
leafy branches in their axils; flowers very similar to 


those of the preceding species, sometimes larger. 


In sandy fields and dry pine barrens, Connecticut to 
Georgia and Louisiana, mostly near the coast. July—Aug. 


5, SCUTELLARIA [Rivin.] L. Sp. Pl. 598. 1753. 


Annual or perennial bitter herbs, some species shrubby. Flowers blue to violet, in 
terminal or axillary bracted mostly secund spike-like racemes, or solitary or 2-3 together in 
the axils. Calyx campanulate, gibbous, 2-lipped, the lips entire, the upper one with a crest or 
protuberance upon its back and often deciduous in fruit, the lower one persistent. Corolla 
much exserted, recurved-ascending, dilated above into the throat, glabrous within, the limb 
2-lipped; upper lip arched, entire or emarginate; lower lip spreading or deflexed, its lateral 
lobes small and somewhat connected with the upper, its middle lobe broad, sometimes emar- 
ginate, the margins mostly recurved. Stamens 4, didynamous, all anther-bearing, ascending 
under the upper lip, the upper pair somewhat the shorter, their anthers 2-celled, ciliate; 
anthers of the lower pair of stamens I-celled, also ciliate. Style unequally 2-cleft at the 
apex; ovary deeply 4-parted. Nutlets subglobose or depressed, papillose or tuberculate, borne 
on a short or elongated gynobase. [Latin,a dish, from the appendage to the fruiting calyx.] 


About 100 species of wide geographic distribution. Besides the following, some 15 others 
occur in the southern and western parts of North America, all known as Skulleap, or Helmet- 
flower. Typespecies: Scutellaria peregrina L. 

* Nutlets wingless, very slightly elevated on the short gynobase. 
Flowers 3”-5” long, in axillary and sometimes terminal secund racemes. 1. S. lateriflora. 
Flowers 6”—15” long, in terminal often panicled racemes. 
Plant glabrous or very nearly so; leaves broad. 
Plants pubescent, puberulent or pilose. 
Leaves all except the floral crenate or dentate, broad. 
Canescent, not glandular; corolla canescent. 3. S. incana. 
Densely glandular-pubescent ; corolla puberulent. 4. S. cordifolia. 
Pubescent below, glandular above; corolla nearly glabrous. 5. S. pilosa. 
Leaves all except the lowest entire, narrow. 6. S. integrifolia. 
Flowers solitary in the axils or sometimes also in terminal bracted racemes. 


ty 
nH 


. Serrata. 


Perennial from a thick woody root. >. S. resinosa. 
Fibrous-rooted; perennial by rootstocks or stolons. 
Flowers 2”—4” long. 8. S. parvula. 


Flowers 8”-13” long. 


106 LABIATAE. Vor. III. 


Minutely and densely glandular-pubescent, resiniferous. 9g. S. Brittonii. 
Densely cinerous-pubescent, pale. 10. S. Bushii. 
Glabrous or merely slightly puberulent. 
Leaves ovate, slender-petioled, cordate, obtuse. 11. S. saxatilis, 
Leaves lanceolate or ovate-lanceolate, nearly sessile, acute. 12. S. galericulata. 


** Nutlets membranous-winged, elevated on the slender gynobase; flowers axillary. 
13. 9. Nervosa. 


1. Scutellaria lateriflora L. Mad-dogor Blue 
Skullcap. Fig. 3575. 
Scutellaria lateriflora L. Sp. Pl. 598. 1753. 


Perennial by slender stolons, glabrous throughout 
or puberulent above; stem slender, leafy, erect or 
ascending, commonly branched, 4’-23° high. Leaves 
ovate, ovate-oblong or ovate-lanceolate, thin, slender- 
petioled, acute or acuminate at the apex, coarsely 
dentate-serrate, obtuse, rounded or subcordate at the 
base, 1-3’ long, the upper gradually smaller, the 
uppermost sometimes entire; racemes narrow, se- 
cund, axillary or often also terminal and leafy- 
bracted, several-many-flowered; flowers 3-5” long; 
calyx short; corolla blue, varying to nearly white, 
its lips about equal, one-fifth as long as the tube; 
nutlets borne on a very short gynobase. 

In wet places, Newfoundland to Ontario and British 
Columbia, Florida, Mississippi, New Mexico and Ore- 
gon. July-Sept. Blue pimpernel. Side-flowering scull- 
cap. Madweed. . Hoodwort. 


2. Scutellaria serrata Andr. Showy Skull- 


cap. Fig. 3576. 
Scutellaria serrata Andr. Bot. Rep. pl. 494. 1809. 
S. laevigata Aiken; Eaton, Man. Ed. 6, 333. 1833. 


Perennial, glabrous, or puberulent above; stem 
slender, erect, simple or branched, 1°-2° high. Leaves 
ovate or elliptic, slender-petioled, acute at the apex, 
narrowed, or the lowest rounded or subcordate at 
the base, crenate or dentate, 2-4’ long, the upper- 
most reduced to small floral bracts; racemes almost 
always simple and terminal, loosely flowered, the 
flowers opposite; fruiting calyx about 3” long; co- 
rolla 1’ long, blue, minutely puberulent, its tube nar- 


row, gradually expanded above into the throat, its | 
rather narrow upper lip a little shorter than the 
lower; nutlets borne on a short gynobase. 

In woods, southern New York and Pennsylvania to 
South Carolina, Illinois, Kentucky and Tennessee. One 
of the handsomest of the American species. Ascends to 
3000 ft. in Virginia. May—June. 


3. Scutellaria incana Muhl. Downy Skull- 
cap. Hig 2577. 


Scutellaria incana Muhl. Cat. 56. 1813. 


\ Scutellaria canescens Nutt. Gen. 2: 38. 1818. 
BESS. serrata Spreng. Syst. 2: 703. 182s. Not Andr. 1809. 


Lie, adh Perennial, finely and densely whitish downy, or 

Z the upper surfaces of the leaves glabrous; stem 
rather strict, erect, usually much branched above, 
2°-4° high. Leaves ovate, oval, or oblong, rather 
firm, slender-petioled, acute at the apex, crenate- 
dentate, narrowed, rounded or the lower subcordate 
at the base, 3-42’ long; racemes terminal, usually 
numerous and_ panicled, several-many-flowered ; 
fruiting calyx 13’-2’ long; corolla 9’—10” long, ca- 
nescent; upper lip of the corolla slightly longer 
than the lower; gynobase very short. 


In moist woods and thickets, Ontario to Michigan, 
Kansas, Georgia, Florida and Alabama. June—Aug. 


GENUS 5. MINT FAMILY. 107 


4. Scutellaria cordifolia Muhl. Heart-leaved 
Skullcap. Fig. 3578. 


Scutellaria cordifolia Muhl. Cat. 56. 1813. 
Scutellaria versicolor Nutt. Gen. 2: 38. 1818. 


Perennial, densely glandular-pubescent ; stem erect, 
usually stout, 1°-3° high, often simple. Leaves 
prominently veined, slender-petioled, broadly ovate, 
crenate-dentate all around, 2-4’ long, all but the 
uppermost cordate at the base; racemes terminal, 
narrow, solitary or panicled; bracts ovate, mostly 
entire, commonly longer than the pedicels; fruiting 
calyx nearly 3” long; corolla puberulent, 10’-12” 
long, blue with the lower side lighter or white, its 
tube narrow, its throat moderately dilated, its lat- 
eral lobes about as long as the upper lip; gynobase 
short. 


In woods and _ thickets, especially along streams, 
Pennsylvania to Florida, west to Minnesota, Kansas, 
oo and Texas. Includes several races. June— 

ug. 


5. Scutellaria pilosa Michx. Hairy Skullcap. Fig. 3579. 


Scutellaria pilosa Michx. Fl. Bor. Am, 2: 11. 1803. 
Scuteilaria ovalifolia Pers. Syn. 2: 136. 1807. 

S. hirsuta Short, Transyl. Journ. Med. 8: 582. 1836. 
Scutellaria pilosa hirsuta A. Gray, Syn. Fl. 2: Part 

T, 3705 Loos 

Perennial, stem slender, simple or branched, 
hairy or downy below, sometimes hirsute, glan- 
dular-pubescent above, 1°-3° high. Leaves ovate, 
oval, or oblong, petioled, obtuse, or the upper 
subacute ‘at the apex, crenate, 1-3’ long, nar- 
rowed or rounded at the base or the lower sub- 
cordate; racemes terminal, solitary or panicled, 
sometimes also in the upper axils; bracts oblong 
or spatulate, entire, longer than the pedicels; 
fruiting calyx about 3” long; corolla blue, 6’-8” 
long, minutely puberulent or glabrous, its lower 
lip and lateral lobes somewhat shorter than the 
arched upper one; gynobase short. 

In dry sandy woods and thickets, southern New 
York and .Pennsylvania to Michigan, Missouri, 
Florida and Texas. Races differ in size and in pubes- 
cence. Ascends to 4000 ft. in North Carolina. May- 
July. 

6. Scutellaria integrifolia L. Larger or 

Hyssop Skullcap. Fig. 3580. 
Scutellaria integrifolia L. Sp. Pl. 599. 1753. 
Scutellaria hyssopifolia L. Sp. Pl. 599. 1753. 

Perennial, hoary with a minute down; stem 
slender, erect, rather strict, simple or branched, 
6’-23° high. Leaves thin, linear to oblong, peti- 
oled, or the upper sessile, obtuse at the apex, 
entire, 1-2’ long, 2’-6” wide, or the lower ovate, 
lanceolate or nearly orbicular, obtuse and some- 
times subcordate at the base, often crenate-den- 
tate or incised; racemes solitary or several, term- 
inal; bracts linear-oblong, subacute, longer than 
the pedicels; fruiting calyx 2-3” long; corolla 
blue, or whitish underneath, 107-15” long, its 
large lips nearly equal; gynobase short. 


In fields, woods and thickets, Massachusetts to 
West Virginia, Tennessee, Arkansas, Florida, Louisi- 
ana and Texas. Consists of several races. May- 
Aug. Large-flowered scullcap. 


108 LABIATAE. VoL. III. 


Scutellaria Drumm6ndii Benth., a low annual villous species with flowers solitary in the axils, 
admitted into our first edition as recorded from Kansas, is not definitely known north of Texas. 


7. Scutellaria resinodsa Torr. Resinous 
Skullcap. Fig. 3581. 


S. restnosa Dorr, Ann, Myc. iN. Ya 2) 2325teege 
S. Wrightit A. Gray, Proc. Am. Acad. 8: 370. 1872. 


Perennial from a thick woody root, minutely 
canescent or puberulent and usually resiniferous ; 
stems rather slender, rigid, tufted, leafy, ascend- 
ing, 6-10’ high. Leaves ovate, oval, or oblong, 
sessile, or the lower short-petioled, entire, obtuse 
at the apex, mostly narrowed at the base, 3-6” 
long; flowers solitary in the axils; fruiting calyx 
nearly 3” long; corolla violet or nearly white, 
very pubescent, 6-8” long, its tube narrow and 
lips nearly equal; gynobase short. 


On dry plains, Nebraska and Kansas to Texas and 
Arizona. May-—Aug. 


8. Scutellaria parvula Michx. Small Skull- 
cap. Fig. 3582. 

S. parvula Michx. Fl. Bor. Am. 2: 11. 1803. 

Scutellaria ambigua Nutt. Gen. 2: 37. 1818. 

Si parweis var. mollis A. Gray, Syn. Fl. 2': 380. 

15706. 

SY campestris Britton, Mem. Torr. Club 5: 283. 

1894. 

Glabrous, or quite densely pubescent, sometimes 
slightly glandular, perennial by slender tuberous- 
thickened rootstocks; stems erect or ascending, 
very slender, usually branched, 3-12’ tall. Leaves 
ovate, oval or lanceolate, or the lower nearly 
orbicular, entire and sessile, or the lower dentate 
and petioled, 3’—-12” long; flowers solitary in 
the axils; fruiting calyx about 2” long; corolla 
2-4” long, violet, pubescent; gynobase short. 

In sandy soil, Quebec to Ontario, South Dakota, 
Florida,. Nebraska and Texas. Races differ in pu- 
bescence and in leaf-form. April—July. Little scull- 
cap. 


g. Scutellaria Brittonii Porter. Britton’s 
Skullcap. Fig. 3583. 


Scutellaria resinosa A. Gray, Syn. Fl. 2: Part 1, 381. 
1878. Not Torr. 1827. 


Scutellaria Brittonii Porter, Bull. Torr. Club 21: 177. 
1894. 


Perennial by tuberous-thickened rootstocks, vis- 
cidly glandular, pubescent or puberulent, branched 
from the base; stems erect, 4-8’ high, leafy. 
Leaves oblong or oval, sessile and entire or the 
lowest short-petioled and slightly crenulate, ob- 
tuse at the apex, rather prominently veined on 
the lower surface, 6-12” long, the upper scarcely 
smaller; flowers solitary in the axils; pedicels 
mostly shorter than the calyx; corolla pubescent, 
blue, 10-15” long, the tube narrow below, en- 
larged above into the throat; gynobase short. 


Nebraska (according to Coulter); Colorado and 
Wyoming. June-July. 


GENUS 5. MINT FAMILY. 


to. Scutellaria Bushii Britton. Bush’s Skullcap. 
Fig. 3584. 


fcutellaria Bushii Britton, Manual, 785. 1901. 


Roots thick-fibrous; stems several or numerous, 
tufted, erect or nearly so, finely cinereous-pubescent, 
‘12’ high. Leaves oblanceolate to oblong-oblanceo- 
late, obtuse, entire, sessile, #/-13’ long, 2-4” wide, the 
veins rather prominent; flowers solitary in the axils, 
short-peduncled, about 1’ long; corolla short-pubescent. 


109 


May-June. 


Barrens, southern Missouri. 


—-> 


11. Scutellaria saxatilis Riddell. Rock 
Skullcap. Fig. 3585. 
S. saxatilis Riddell, Suppl. Cat. Pl. Ohio, 14. 1836. 


Perennial by filiform runners or stolons, 
glabrate or sparingly puberulent; stem slen- 
der, weak, ascending or reclining, simple or 
branched, 6-20’ long. Leaves ovate, slender- 
petioled, thin, coarsely crenate, obtuse at the 
apex, cordate at the base, 1’-2’ long, or the 
lower nearly orbicular, and the upper lanceo- 
late, subacute and entire; flowers solitary in 
the upper axils, or clustered in a terminal 
leafy-bracted loose raceme; bracts longer than 
the pedicels; fruiting calyx about 2” long; co- 
rolla light blue, very nearly glabrous, 8-10” 
long; gynobase short. 

On moist banks and in thickets, Delaware to 
South Carolina, Ohio and Tennessee. Ascends 
to 3000 ft. in Virginia. May~July. 


12. Scutellaria galericulata L. Hooded 
Willow-herb. Marsh or European 
Skullcap. Fig. 3586. 


Scutellaria galericulata L. Sp. Pl. 599. 1753. 


Perennial by filiform stolons, not tuber-bear- 
ing, puberulent or pubescent; stem erect, usually 
branched, 1°-3° high. Leaves oblong-lanceolate 
to ovate-oblong, thin, short-petioled, or the upper 
sessile, acute at the apex, dentate with low teeth 
or the upper entire, subcordate or rounded at 
the base, 1-23’ long, the uppermost usually much 
smaller and bract-like; flowers solitary in the 
axils; peduncles shorter than the calyx; corolla 
blue, puberulent, nearly or quite 1’ long, with a 
slender tube and slightly enlarged throat; gyno- 
base short. 


In swamps and along streams, Newfoundland to 
Mackenzie, Alaska, New Jersey, the mountains of 
North Carolina, Ohio, Nebraska, Arizona and Wash- 
ington, Also in Europe and Asia. June—Sept. 


Scutellaria Churchilliana Fernald, of Maine and 

ew Brunswick, has smaller flowers, solitary in the 
axils, its leaves much like those of S. /aterifiora, 
and is, perhaps, a hybrid. 


1io LABIATAE. Vor. Til. 


13. Scutellaria nervosa Pursh. Veined 
Skullcap. Fig. 3587. 


S. nervosa Pursh, Fl. Am. Sept. 412. 1814. 
Scutellaria teucriifolia J. E. Smith in Rees’ Cycl. 
32: no. 15. 1816. 


Perennial by filiform stolons; stem glabrous 
or sparingly pubescent, erect, slender, simple 
or sometimes branched, 62° high. Leaves 
thin, glabrous, or sometimes decidedly pubes- 
cent, the lower slender-petioled, nearly orbicu-- - 
lar, crenate, often subcordate at the base, the 
middle ones larger, ovate, 12’ long, sessile or 
nearly so, obtuse or acute, coarsely dentate or 
crenate, the upper lanceolate or ovate-lanceo- 
late, acute, commonly entire; flowers solitary 
in the axils; fruiting calyx about 2” long; 
corolla blue, 4’—-5”" long, puberulent, the lower 
lip longer than the concave upper one; nutlets 
membranous-winged, borne on a slender gyno- 
base. 

In moist woods and thickets, southern Ontario, 
New York and New Jersey to Illinois and Mis- 
souri, North Carolina and Tennessee. Ascends 
to 3000 ft. in Virginia. May—Aug. 


6. MARRUBIUM [Tourn.] L. Sp. Pl. 582. © 1753. 


Perennial branching mostly woolly herbs, with petioled dentate rugose leaves, and small 
white or purplish flowers in dense axillary clusters, the juice bitter. Calyx tubular, 5-10- 
nerved, regularly 5-10-toothed, the teeth nearly equal, or the alternate ones shorter, acute or 
aristate, spreading or recurved in fruit. Corolla-limb 2-lipped, the upper lip erect, entire or 
emarginate, the lower spreading, 3-cleft, its broader middle lobe commonly emarginate. 
Stamens 4, didynamous, included, the posterior pair the shorter; anthers 2-celled. Style 
2-cleft at the summit, the lobes short. Ovary deeply 4-lobed. Nutlets ovoid, smooth. [Name 
Middle Latin, perhaps from the Hebrew, referring to its bitter qualities. ] 


About 40 species, natives of the Old World, the 
following typical. 


1. Marrubium vulgare L. White or 
Common Hoarhound. Fig. 3588. 


Marrubium vulgare L. Sp. Pl. 583. 1753. 


Stem erect, stout, woolly, especially below, 
1°-3° high, the branches ascending. Leaves 
oval, broadly ovate or nearly orbicular, rugose- 
veined, obtuse at the apex, crenate-dentate, 
rounded, narrowed or subcordate at the base, 
1-2’ long, rough, whitish above, woolly be- 
neath; petioles 2-1’ long, usually exceeding 
the flowers; clusters all axillary, densely many- 
flowered; flowers whitish; calyx-teeth usually 
10, subulate, more or less recurved, glabrous 
above, woolly below. 


In waste places, Maine and Ontario to Minne- 
sota and British Columbia, North Carolina, Ala- 
bama, Texas, Mexico and California. Also in South 
America. Naturalized from Europe. Native also of 
Asia. Old names, houndbene, marrube, marvel. 


7. SIDERITIS [Wourn As. Sp.Pl. 574... aaye 


Annual or perennial, usually pubescent herbs or shrubs, with opposite, entire or toothed 
leaves, and small white, pink, or yellowish flowers in axillary clusters which are separated 
or contiguous. Calyx tubular, 5-10-nerved, slightly 2-lipped, the teeth spine-tipped, the 
upper middle one broader than the others. Corolla-limb exceeding the calyx, 2-lipped; upper 
lip entire or lobed; lower lip longer than the upper, with a broad middle lobe. Stamens 4, 


GENUS 7. ' MINT FAMILY. III 


didynamous, included, the anterior pair the longer; anthers 2-celled, or those of the anterior 
pair more or less imperfect, the sacs divaricate. Style 2-cleit at the summit, lobes obtuse. 
Ovary 4-lobed. Nutlets obovoid, smooth. [Greek, iron, referring to :ts nedicinal use.] 


About 45 species, native of the Old World, most numerous 
in the Orient. Type species: Sideritis hirsuta L. 


1. Sideritis romana L. Simple-beaked Iron-wort. 
Fig. 3589. 
Sideritis romana L. Sp. Pl. 575. 1753. 


Annual, softly pubescent with spreading hairs, the stem 
erect, 4’-1° tall, usually branched. Leaves obovate to 
oblong, 4-13’ long, crenate, sessile cr nearly so, ultimately 
somewhat veiny; clusters few-flowered; flowers white or 
pinkish; calyx-teeth shorter than the tube, the middle one 
of the upper lip ovate, all subulate-tipped; corolla slightly 
exceeding the calyx, the middle lobe of the lower lip 
reniform. 


In fields, southeastern Pennsylvania. Naturalized from the 
Mediterranean region. Also in Bermuda. June—Aug. 


8. AGASTACHE Clayt.; Gron. Fl. Virg. 88. 
[Vieckia Raf. Med. Rep. (II.) 5: 308. 1808.] 
[LopHANTHUS Benth. Bot. Reg. 15: under fl. 1282. 1829. Not Adans. 1763.] 


Tall erect perennial herbs, with serrate, mainly ovate, petioled leaves, and yellowish 
purplish or blue flowers, verticillate-clustered in thick dense or interrupted bracted terminal 
spikes. Calyx narrowly campanulate, somewhat oblique, slightly 2-lipped, 5-toothed, the 
teeth of the upper lip somewhat larger than those of the lower, or all about equal. Corolla 
strongly 2-lipped, the tube as long as the calyx;-upper lip erect, 2-lobed; lower lip spreading, 
3-lobed, its middle lobe broader than the lateral ones and crenulate. Stamens 4, all anther- 
bearing, didynamous, the upper pair the longer; anthers 2-celled, their sacs nearly parallel. 
Ovary deeply 4-parted ; style 2-cleft at the summit. Nutlets ovoid, smooth. [Greek, many spikes.] 


‘About 8 species, natives of North America. Besides the following, 2 or 3 others occur in the 
western United States. Type species: Hyssopus nepetoides L. 


Glabrous or very nearly so, stout; corolla greenish-yellow. 1. A. nepetoides. 
Pubescent, stout; corolla purplish; leaves green both sides. 2. A. scrophulariaefolia. 


Glabrous or slightly pubescent, slender ; corolla blue; leaves pale beneath. 3. A. anethiodora. 


1. Agastache nepetoides (L.) Kuntze. 
Catnep Giant-Hyssop. Fig. 3590. 


Hyssopus nepetoides L. Sp. Pl. 569. 1753. 

Lophanthus nepetoides Benth. Bot. Reg. under 
pl. 1282. 1820. 

Vileckia nepetoides Raf. Fl. Tell. 3: 89. 1836. 

‘A, nepetoides Kuntze, Rev. Gen. Pl. 511. 1801. 


Glabrous, or slightly puberulent; stem 
stout, 2°-5° high, branched, at least above, 
sharply 4-angled. Leaves ovate or ovate- 
oblong, acuminate or acute at the apex, 
rounded, cordate or the upper narrowed at 
the base, mostly thin, coarsely dentate, 2’-6’ 
long; spikes 3-18’ long, usually very dense; 
petioles of the lowest leaves often 2’ long; 
bracts ovate, acute or acuminate; calyx- 
teeth oblong or ovate-oblong, obtuse or sub- 
acute, sometimes purplish; corolla greenish 
yellow, about 4” long, scarcely exceeding 
the calyx. 

In woods and thickets, Quebec and Vermont 


to South Dakota, south to Georgia, Tennessee, 
Kansas and Arkansas. July—Sept. 


112 LABIATAE. Vor. III. 


Vg 

. np WF 2. Agastache scrophulariaefolia 
\ nl (ZZ (Willd.) Kuntze. Figwort 
Giant-Hyssop. Fig. 3591. 


Hyssopus scrophulariaefolius Willd. Sp. Pl. 3: 
4 r8or. 

Lophanthus scrophulariaefolius Benth. Bot. 
Reg. under pl. 1282. 1829. 

Vleckia scrophulariaefolia Raf. Fl. Tell. 3: 89. 
1836. 

Agastache scrophulariaefolia Kuntze, Rev. 
Gen. Pls rs. 2 1ser- 


Similar to the preceding species, but com- 
monly taller, strong-scented, the obtusely 
4-angled stem, the petioles and lower sur- 
faces of the leaves more or less pubescent, 
sometimes villous. Leaves nearly identical 
with those of V. nepetoides in size and 
outline; spike sometimes interrupted, 3’-18’ 
long; bracts broadly ovate, abruptly acumi- 
nate; calyx-teeth lanceolate or ovate-lan- 
ceolate, very acute or sometimes acuminate, 
whitish or purplish; corolla purplish, 5-6” 
long, considerably exceeding the calyx. 

In woods and thickets, New Hampshire to 


Ontario, Wisconsin, North Carolina, Ken- 
tucky and Missouri. July—Oct. 


3. Agastache anethiodora (Nutt.) 


Britton. Fragrant Giant-Hyssop. 
Fig. 3592. 
Hyssopus anethiodorus Nutt. Fras. Cat. 1813. 
Hyssopus anisatus Nutt. Gen. 2: 27. 1818. 
Lopanithus anisaius Benth. Bot. Reg. under /l. 
1282. 1829. 
Vieckia anisata Raf. Fl. Tell. 3: 89. 1836. 
Vleckia anethiodora Greene, Mem. Torr. Club 
5: 282. 1894. 
A. anethiodora Britton in Britt. & Brown, Ill. 
Fi. 3: 85. 18098. 


Glabrous, or minutely puberulent; stem 
rather slender, leafy, usually branched, 2°- 
4° high. Leaves ovate or triangular-ovate, 
firm, mostly short-petioled, acute or acumi- 
nate at the apex, truncate, obtuse or some- 
times subcordate at the base, sharply ser- 
rate, green above, pale and minutely canes- 
cent beneath, 27-3’ long, anise-scented; 
spikes dense or interrupt: 1 seldom 6’ long; 
bracts broadly ovate, abru, tly acuminate; 
calyx-teeth ovate to lanceolate, acute, pur- 
ple; corolla blue, 4”-5” long, somewhat 
exceeding the calyx. 


On prairies and plains, Minnesota to Mani- 
toba, Alberta, Illinois, Nebraska and Colorado. 2 
July-Sept. Anise-hyssop. ie 


9. MEEHANIA Britton, Bull. Torr. Club 21: 32. 1894. 


A low pubescent spreading or decumbent herb, with long-petioled cordate leaves, trailing 
leafy stolons, and large blue flowers in terminal secund bracted spikes. Calyx campanulate, 
15-nerved, slightly 2-lipped, its teeth all lanceolate, acute, the 3 upper longer than the 2 lower. 
Corolla much exserted, puberulent without, pubescent within, the tube narrow at the base, 
gradually widely ampliate into the throat, the limb 2-lipped; upper lip 2-lobed, arched, the 
lobes ovate, obtuse; lower lip about equalling the upper, spreading, 3-lobed, the middle lobe 
emarginate, broader than the lateral ones. Stamens 4, didynamous, all anther-bearing, 
included, ascending under the upper lip, the upper pair longer than the lower; anthers 
2-celled, the sacs nearly parallel. Ovary deeply 4-lobed; style equally 2-cleft at the summit. 
Nutlets oblong, smooth. [Named for Thomas Meehan, of Philadelphia, 1826-1901, botanist 
and horticulturist.] 


A monotypic genus of eastern North America. 


a 


GENUS 9. MINT FAMILY. 


1. Meehania cordata (Nutt.) Britton. 
Meehania. Fig. 3593. 


Dracocephalum cordatum Nutt. Gen. 2: 35. 1818. 

Cedronella cordata Benth. Lab. 502. 1834. 

Meehania cordata Britton, Bull. Torr. Club 21: 33. 
pl. 173. 1894. 


Flowering stems ascending, 3-8’ high; stolons 
very slender, leafy throughout, sometimes 2° 
long. Leaves all broadly ovate or ovate-orbicu- 
lar, thin, obtuse or subacute at the apex, crenate 
all around, cordate at the base, sparingly pubes- 
cent with scattered hairs on both surfaces, or 
nearly glabrous beneath, green on both sides, 
1-2’ long, the basal sinus broad; spikes 17-4’ 
long; bracts ovate or oblong, acute, membranous, 
the lower sometimes crenulate and surpassing 
the calyx; bractlets small, lanceolate; calyx about 
5” long, puberulent, its longer teeth about one- 
half the length of the tube; corolla 1-13’ long, 
showy. 

In rich moist woods and thickets, southwestern 
Pennsylvania to Illinois, Tennessee and North 
Carolina. May-July. 


Px 
10. NEPETA [Rivin.] L. Sp. Pl. 570. 1753. 


Herbs, with dentate or incised leaves, and mostly white or blue rather small flowers in 
verticillate clusters, usually crowded in terminal spikes, or axillary and cymose. Calyx tubu- 
lar, somewhat oblique at the mouth, 15-nerved, usually incurved, 5-toothed, scarcely 2-lipped, 
but the upper teeth usually loriger than the lower. Corolla-tube enlarged above, the limb 
strongly 2-lipped; upper lip erect, emarginate or 2-lobed; lower lip spreading, 3-lobed, the 
middle lobe larger than the lateral ones. Stamens 4, all anther bearing, didynamous, ascend- 
ing under the upper lip, the lower pair the shorter; anthers 2-celled, the sacs divaricate. 
Ovary deeply 4-parted; style 2-cleft at the summit. Nutlets ovoid, compressed, smooth. 
[Ancient Latin name of catnep. | 


‘About 150 species, natives of Europe and Asia, the following typical. 


1. Nepeta Cataria L. Catmint. Catnep. 
Nep. Fig. 3594. 
Nepeta Cataria L. Sp. Pl. 570. 1753. 


Perennial, densely canescent, pale green; stem 
rather stout, erect, branched, 2°-3° high, the 
branches straight, ascending. Leaves ovate to 
oblong, petioled, acute at the apex, coarsely cre- 
nate-dentate, mostly cordate at the base, 17-3’ 
long, greener above tt 1 beneath; flower-clusters 
spiked at the ends c. the stem and branches, the 
spikes 1-5’ long; bracts small, foliaceous; bract- 
lets subulate; calyx puberulent, its teeth subu- 
late, the upper about one-half the length of the 
tube; corolla nearly white, or pale purple, dark- 
dotted, puberulent without, 5’-6” long, its lobe a 
little longer than the calyx, the broad middle lobe 
of its lower lip crenulate. 


In waste places, New Brunswick and Quebec to 
South Dakota, Oregon, South Carolina, Kansas and 
Utah; also in Cuba. Naturalized from Europe. 
Native also of Asia. July—Nov. 


11. GLECOMA L. Sp. Pl. 578. 1753. 


Low diffuse creeping herbs, with long-petioled nearly orbicular or reniform crenate 
leaves, and rather large blue or violet flowers in small axillary verticillate clusters. Calyx 
oblong-tubular, 15-nerved, oblique at the throat, not 2-lipped, unequally 5-toothed. Corolla- 
tube exserted, enlarged above, the limb 2-lipped; upper lip erect, 2-lobed or emarginate;.the 
lower lip spreading, 3-lobed, the middle lobe broad, emarginate, the side lobes small. Stamens 
4, didynamous, all anther-bearing, ascending under the upper lip of the corolla, not exserted, 
the upper pair the longer; anther-sacs divergent. Ovary deeply 4-parted. Nutlets ovoid, 
smooth. [Greek name for thyme or pennyroyal.] 

About 6 species of Europe and Asia, the following typical. 

8 


114 LABIATAE. Vor. His 


1. Glecoma hederacea L. Ground 
Ivy. Gill-over-the-Ground. 
Field Balm. Fig. 3595. 


Glecoma hederacea L. Sp. Pl. 578. 1753. 
N. Glechoma Benth. Lab. Gen. & Sp. 485. — 


1834. 
N. hetieracea BSP. Prel: ‘Cat. Newyerage 
I A 

Perennial, pubescent, the creeping stems 
leafy, sometimes 18’ long, the branches 
ascending. Lower petioles commonly 
longer than the leaves; leaves green 
both sides, 4-12’ in diameter; clusters 
few-flowered, the flowers 7’—10” long, 
short-pedicelled; _bractlets _— subulate, 
shorter than the calyx; calyx puberulent, 
its teeth acute or lanceolate-acuminate, 
about one-third as long as the tube; 
corolla-tube 2-3 times as long as the 
calyx; upper pair of stamens much 
longer than the lower. ’ 

In waste places, woods and thickets, New- 
foundland to Ontario, Minnesota, Oregon, 
Georgia, Tennessee, Kansas and Colorado. 
Old names, alehoof, cat’s-foot, gill, gill-ale, 
gill-go-by-the-ground, hayhofe, haymaids, 
hove, tunhoof, creeping charlie, robin-run- 
away, gill-run-over, crow-vituals, wild 
snake-root, hedge-maids. March—May. 


12. MOLDAVICA [Tourn.] Adans. Fam: Pl. 2: ;190" 27ae 


Herbs, with dentate entire or incised leaves, and blue or purple flowers in axillary and 
terminal bracted clusters, the bracts pectinate in our species. Calyx tubular, 15-nerved, 
straight or incurved, 5-toothed, the upper tooth much larger than the others, or 2-lipped with 
the 3 upper teeth more or less united. Corolla expanded above, its limb 2-lipped; upper lip 
erect, emarginate; lower lip spreading, 3-lobed, the middle lobe larger than the lateral ones, 
sometimes 2-cleft. Stamens 4, didynamous, ascending under the upper lip, the upper pair 
longer than the lower; anthers 2-celled, the sacs divaricate; style 2-cleft at the summit; ovary 
deeply 4-parted. Nutlets ovoid, smooth. [From Moldavia.] 


About 35 species, natives of the northern hemisphere. Only the following are known in North 
America. Type species: Dracocephalum Moldavica L. 
Corolla 2-3 times as long as the calyx; clusters mostly axillary. 1. M. parviflora. 
Corolla scarcely exceeding the calyx; clusters mostly terminal, dense. _ 2. M. Moldavica. 


1. Moldavica parviflora ( Nutt.) Brit- 


Ely, 
ton. American Dragon-head. ii Wines 
. a | US 
Fig. 3506. Nees 
Dracocephalum parviflorum Nutt. Gen. 2: 35. ee ah 
1818. CA): hyp , 
Annual or biennial, somewhat pubescent, WIZ 
or glabrous; stem rather stout, usually ZZ 
branched, 624° high. Leaves lanceolate, 
ovate, or oblong, slender-petioled, serrate, N 
or the lower incised, acute or obtuse at the 


apex, rounded or narrowed at the base, 
thin, 1-3’ long; clusters dense, many-flow- l\ 
ered, crowded in dense terminal spikes, 
and sometimes also in the upper axils; 


bracts ovate to oblong, pectinate with awn- a 

pointed teeth, shorter than or equalling the 

calyx; pedicels 1’—2” long; upper tooth of \ é 

the calyx ovate-oblong, longer than the aN 
narrower lower and lateral ones, all acumi- 


nate; corolla light blue, scarcely longer 
than the calyx. 


In dry gravelly or rocky soil, Quebec and 2 
Ontario to Alaska, New York, Iowa, Missouri ay 
and Arizona. May—Aug. 


GENUS I2. MINT FAMILY. 


2. Moldavica Moldavica (L.) 
Britton. Moldavian Dragon- 
head or Balm. Fig. 3597. 


Dracocephalum Moldavica L. Sp. Pl. 595. 
1753. 


EEE 
WS WS 


SS 


Annual, puberulent; stem erect, usu- 
ally widely branched, 1°-23° high. Leaves 
oblong or linear-oblong, dentate or some- 
what incised, obtuse at the apex, usually 
narrowed at the base, 1’-2’ long, 2-6” 
wide; clusters loose, few-flowered, com- 
monly numerous, mostly axillary; bracts 
narrowly oblong, usually shorter than 
the calyx, deeply pectinate with aristate 
teeth; pedicels 2’-4” long; calyx slightly 
curved, the 2 lower teeth somewhat 
shorter than the 3 broader equal upper 
ones; corolla 2-3 times as long as the 
calyx. 


In a cafion near Spring View, Nebr. 
Also in northern Mexico. Introduced 
from central Europe. June—Aug. 


13. PRUNELLA L. Sp. Pl. 600. 1753. 


Perennial simple or sometimes branched herbs, with petioled leaves, and rather small 
clustered purple or white flowers, in terminal and sometimes also axillary, dense bracted 
spikes or heads. Calyx oblong, reticulate-veined, about 10-nerved, deeply 2-lipped, closed in 
fruit; upper lip nearly truncate, or with 3 short teeth; lower lip 2-cleft, its teeth lanceolate. 
Corolla-tube inflated, slightly narrowed at the mouth, its limb strongly 2-lipped; upper lip 
entire, arched; lower lip spreading, 3-lobed. Stamens 4, didynamous, ascending under the 
upper lip of the corolla, the lower pair the longer; filaments of the longer stamens 2-toothed 
at the summit, one of the teeth bearing the anther, the other sterile; anthers 2-celled, the 
sacs divergent or divaricate. Ovary deeply 4-parted. Nutlets ovoid, smooth. [Origin of 
name doubtful; often spelled Brunella, the pre-Linnaean form.] 


About 5 species, of wide geographic distribution. Only the following typical one occurs in 
North America. i 


1. Prunella vulgaris L. Self-heal. Heal-all. Dragon-head. Fig. 3508. 
Sy) Prunella vulgaris L. Sp. Pl. 600. 1753. 


Pubescent or nearly glabrous; stem slen- 
der, procumbent or ascending or erect, 
usually simple, but sometimes considerably 
branched, 2’-2° high. Leaves ovate, oblong 
or oblong-lanceolate, obtuse or subacute at 
the apex, usually narrowed at the base, 
entire, crenate, dentate, or incised, 17-4’ 
long, the lowest commonly shorter and 
sometimes subcordate; spikes terminal, ses- 
sile or short-peduncled, very dense, 4-1’ 
long in flower, becoming 2’-4’ long in 
fruit; bracts broadly ovate-orbicular, cus- 
pidate, more or less ciliate; corolla violet, 
purple, or sometimes white, 4-6” long, 
about twice as long asthe purplish or green 
calyx; calyx-teeth often ciliate. 


In fields, woods and waste places throughout 
nearly the whole of North America. Native 
and naturalized from Europe. Native also of 
Asia. Thimble-flower. All-heal. Brown-wort. 
Carpenter’s-herb or -weed. Hook-heal or 
-weed. Heart-of-the-earth. Sicklewort. Blue- 
curls. May-—Oct. 


Prunella laciniata L., differing by pin- 
natifid or deeply incised leaves, found near 
Washington, D. C., and illustrated in our first 
edition, is a race of this species. 


116 LABIATAE. Vor. III. 


14. DRACOCEPHALUM [Tourn.] L. Sp. Pl. 504. 1753. 
[Puysostecia Benth. Lab. Gen. & Sp. 504. 1834.] 


Erect perennial glabrous or puberulent herbs, with serrate, dentate or entire leaves, and 
large or middle-sized, bracted purple violet, pink or white flowers in terminal spikes, or 
spike-like racemes. Calyx campanulate or oblong, membranous, swollen and remaining open 
in fruit, faintly reticulate-veined and 10-nerved, equally 5-toothed. Corolla much longer than 
the calyx, its tube gradually much enlarged upward, its limb strongly 2-lipped; upper lip 
concave, rounded, nearly or quite entire; lower lip spreading, 3-lobed, the middle lobe com- 
monly emarginate. Stamens 4, didynamous, ascending under the upper lip of the corolla, the 
lower pair the longer; filaments pubescent; anthers all alike, 2-celled, the sacs nearly paraliel, 
the margins of their valves commonly spinulose or denticulate. Ovary 4-parted. Nutlets 
ovoid-triquetrous, smooth. [Greek, dragon-head.] 


About 7 species, natives of North America, known as False Dragon-head or Lion’s-heart. Type 
species: Dracocephalum virginianum L. : 


Flowers 9” long, or more; leaves firm. 
Spike dense, many-flowered. 
Spike loose, few-flowered ; leaf-serrations mostly blunt. 
Flowers 5”—7” long; leaves thin. ; 
Spike loose; 4-8’ long; leaves few and distant. 3. D. intermedium. 
Spike dense, :’-4’ long; stem leafy. 4. D. Nuttallii. 


D. virginianum. 
. D. denticulatum. 


bi 


1. Dracocephalum virginianum L. Dragon-head. Obedient Plant. Lion’s 
Heart. Fig. 3500: 


Dracocephalum virginianum L. Sp. Pl. 594. 1753. 


Dracocephalum speciosum Sweet, Brit. Fl. Gard. 
pl. 93. 1825. 


Physostegia virginiana Benth. Lab. Gen. & Sp. 

504. 1834. 

Stem erect or ascending, rather stout, sim- 
ple or branched above, 1°-4° tall. Leaves 
firm, lanceolate, oblong-lanceolate, or linear- 
lanceolate, acuminate at the apex, sharply 
serrate or serrulate, narrowed at the base, 
the upper all sessile, 2’-5’ long, 2’-7” wide, 
the lowest petioled; bracts lanceolate, shorter 
than the calyx; spikes dense, becoming 4-8 
long in fruit, many-flowered; flowering calyx 
campanulate or somewhat turbinate, its teeth 
ovate, acute, about one-half as long as the 
tube; fruiting calyx oblong, 4-5” long, the 
teeth much shorter than the tube; corolla 
pale purple or rose, about 1’ long, often var- 
iegated with white, temporarily remaining in 
whatever position it is placed. 


In moist soil, Quebec to Ontario, Minnesota, Arkansas, Florida, Louisiana and Texas; escaped 
from gardens eastward. Races differ in width and serration of leaves and in size of flowers. 
July—Sept. 


Physostegia Digitalis Small, with broadly oblong to elliptic repand or undulate leaves, of the 
Southern States, perhaps extends northward into Missouri. 


GENUS I4. MINT FAMILY, 


2. Dracocephalum denticulatum Ait. Few- 
flowered Lion’s Heart. Fig. 3600. 


Prasium purpureum Walt. Fl. Car. 166. 1788? 
Drac. denticulatum Ait. Hort. Kew. 2: 317. 17809. 
P. virginiana var. denticulata A. Gray, Syn. Fl. 2': 383. 


1878. 
P. denticulata Britton, Mem. Torr. Club 5: 284. 1894. 


Stem slender, ascending or erect, simple, or little 
branched, 1°-2° high. Leaves firm or rather thin, 
oblong, linear-oblong, or oblanceolate, obtuse or 
acute at the apex, narrowed at the base, crenulate, 
obtusely dentate, or entire, 1-3’ long, 2’-6” wide, 
the upper sessile, the lower slender-petioled; spike 
loosely few-several-flowered; bracts lanceolate, little 
longer than the fruiting pedicels; flowering calyx 
oval-campanulate, its teeth acute, about one-third 
as long as the tube; fruiting calyx oblong, 3-4” 
long; corolla rose-pink, nearly or quite 1’ long. 


In moist soil, Pennsylvania to Illinois, Florida and 
Texas. June—Aug. 


3. Dracocephalum intermédium Nutt. 
Slender Lion’s Heart. Fig. 3601. 


Dracocephalum intermedium Nutt. Trans. Am. Phil. 
Soc, CL) si: 187. o1833—37- 

Physostegia intermedia A. Gray, Proc. Am. Acad. 8: 
B7ilee Logo 


Stem very slender, usually quite simple, 1°-3° 
high. Leaves usually few pairs, remote, thin, 
mostly shorter than the internodes, narrowly 
lanceolate or linear, acute or acuminate at the 
apex, repand-denticulate, little narrowed at the 
base, all sessile, or the lowest petioled, 23’ long, 
2’’-4” wide; spikes very slender, remotely many- 
flowered, 4-8’ long in fruit; lower bracts often 
nearly as long as the campanulate calyx; calyx- 
teeth acute, shorter than the tube; fruiting calyx 
broadly oval, 2-24” long; corolla much dilated 
above, 5’—7” long. 


On prairies, western Kentucky to Missouri, Louis- 
iana, Arkansas and Texas. May-July. 


4. Dracocephalum Nuttallii Britton. Pur- 
ple or Western Lion’s Heart. 
Fig. 3602. 
Physostegia parviflora Nutt.; Benth. in DC. Prodr. 
I2: 434. Assynonym. 1848. A. Gray, Proc. Am. 


Acad. 8: 371. Not Dracocephalum parviflorum 
Nutt. 


Stem rather stout, usually simple, 1°-3° high. 
Leaves lanceolate, oblong-lanceolate, or ovate- 
lanceolate, acute, acuminate, or the lower obtuse 
at the apex, sharply serrate or dentate, somewhat 
narrowed at the base, all sessile or the lowest 
petioled, thin, 3-4’ long, 3-10” wide; spikes 
densely several-many-flowered, 17-4’ long; bracts 
ovate or ovate-lanceolate, acute, shorter than the 
calyx; flowering calyx campanulate, its teeth 
ovate, obtuse or subacute, about one-third as long 
as the tube; fruiting calyx globose-oblong, 2’’-3” 
long; corolla purple, 5’—7” long. 

-In moist soil, Wisconsin and Minnesota to Ne- 
braska, North Dakota, Saskatchewan, British Colum- 
bia and Oregon. June--Aug. 


118 LABIATAE. VoL, III. 


1s. SYNANDRA Nutt. Gen. 2: 29. 1818. 


An annual or. biennial, somewhat hirsute, simple or little branched herb, with long- 
petioled ovate cordate crenate leaves, and large whitish flowers in a terminal leafy-bracted 
spike. Calyx campanulate-oblong, membranous, deeply 4-cleft, inflated in fruit, faintly and 
irregularly veined, the lobes narrowly lanceolate, the two upper shorter than the lower. 
Corolla much longer than the calyx, its tube narrow below, much expanded above, 2-lipped; 
upper lip concave, entire; lower lip spreading, 3-lobed. Stamens 4, didynamous, ascending 
under the upper lip of the corolla; filaments villous; anthers glabrous, 2-celled, the sacs 
divaricate, the contiguous ones of the upper pair of stamens sterile and connate. Ovary 
deeply 4-lobed; style unequally 2-cleft at the summit. Nutlets ovoid, smooth, sharply angled. 
[ Greek, stamens-together. | 


A monotypic genus of southeastern North America. 


1. Synandra hispidula (Michx.) Britton. 
Synandra. Fig. 3603. 


Lamium hispidulum Michx. Fl. Bor. Am. 2: 4. 1803.’ 
Synandra grandiflora Nutt. Gen. 2: 29. 1818. 
=» Torreya grandiflora Raf. Am. Month. Mag. 3: 356. 1818. 
' S. hispidula Britton, Mem. Torr. Club 5: 285. 1804. 
“ Stem rather slender, erect or ascending, weak, 
1°-23° long, striate. Leaves thin, the lower and 
basal ones broadly ovate, or nearly orbicular, pal- 
mately veined, acute or obtuse at the apex, deeply 
cordate at the base, the blade 2’-4’ long, and com- 
monly shorter than the petiole; floral leaves sessile, 
ovate or ovate-lanceolate, acute or acuminate, the 
flowers solitary in their axils, the uppermost leaves 
very small; calyx hirsute, its lobes about as long as 
the tube; corolla 1-13’ long, showy, the lower lip 
with purple lines. . 


Along streams and in wet woods, Ohio to Illinois, 
Virginia and Tennessee. Ascends to. 3,500 ft. in Vir- 
ginia. May—June. 


16. PHLOMIS [Tourn.] L. Sp. Pl. 584. 1753. 


Tall perennial herbs, or shrubs. Calyx tubular or tubular-campanulate, 5-10-nerved, the 
limb mostly equally 5-toothed. Corolla-tube usually with a woolly ring within, shorter than 
or exceeding the calyx, the limb strongly 2-lipped; upper lip erect, concave, arched or some- 
times keeled, entire or emarginate; lower lip spreading, 3-cleft. Stamens 4, didynamous, 
ascending under the upper lip of the corolla, the anterior pair the longer and their filaments 
with hooked appendages at the base; anther-sacs divergent. Ovary deeply 4-lobed; style 
subulate, 2-cleft at the summit, one of the lobes smaller than the other. Nutlets ovoid, 
glabrous, or pubescent above. [Greek, mullen, in allusion to the thick woolly leaves of some 


De 


species. | 
About 50 species, natives of the Old World. Type 
species: Phlomis fruticosa L. \I: | 
7 th Kol 

. x Wig 

1. Phlomis tuberdsa L. Jerusalem Sage. <= 
Sage-leaf Mullen. Fig. 3604. 
wy i 


y 
Phlomis tuberosa L. Sp. Pl. 586. 1753. 


Herbaceous from a thickened root; stem stout, 
purplish, glabrous or loosely pubescent above, 
usually much branched, 3°-6° tall, the branches 
nearly erect. Lower leaves triangular-ovate, long- 
petioled, acuminate or acute at the apex, coarsely 
dentate or incised-dentate, rather thick, deeply 
cordate at the base, strongly veined, 5-10’ long, 
3-6’ wide; upper leaves lanceolate, short-petioled 
or sessile, truncate or sometimes narrowed at the 
base, the uppermost (floral) very small; clusters 
densely many-flowered; bractiets subulate, ciliate- 
hirsute or nearly glabrous; calyx 5’-6” long, its 
teeth setaceous with a broader base, spreading; 
corolla 10-12” long, pale purple or white, twice 
as long as the calyx, densely pubescent, and the 
margins of its upper lip fringed with long hairs. 

In waste places, south shore of Lake Ontario. Naturalized from southern Europe. June-Sept. 


GENUS 17. MINT FAMILY. = 


57. GALEOPSIS L. op.) Ply 579% 1753: 


Erect annual branching herbs, with broad or narrow leaves, and rather small yellow red 
purple or mottled verticillate-clustered flowers in the upper axils, or forming terminal dense 
or interrupted spikes. Calyx campanulate or tubular-campanulate, 5-I0-nerved, 5-toothed, 
the teeth nearly equal, spinulose. Corolla-tube narrow, the throat expanded, the limb strongly 
2-lipped; upper lip erect, concave, entire; lower lip spreading, 3-cleft, the middle lobe obcor- 
date or emarginate. Stamens 4, didynamous, ascending under the upper lip of the corolla, 
the anterior pair the longer; anthers 2-celled, the sacs transversely 2-valved, the inner valve 
ciliate, the outer smooth, larger. Ovary deeply 4-parted; style 2-cleft at the summit. Nut- 
lets ovoid, slightly flattened, smooth. [Greek, weasel-like. | 

About 6 species, natives of Europe and Asia. Type species: Galeopsis Tetrahit L. 


Plant puberulent ; leaves linear to ovate-lanceolate. 1. G. Ladanum., 
Plant hispid; leaves ovate. 2. G. Tetrahit. 


1. Galeopsis Ladanum L. Red Hemp- 
Nettle. Ironwort. Fig. 3605. 
Galeopsis Ladanum L. Sp. Pl. 579. 1753. 


Puberulent, sometimes glandular above; stem 
slender, erect, much branched, 6-18’ high, the 
branches ascending. Leaves linear, ovate-ob- 
long, or ovate-lanceolate, mostly short-peti- 
oled, acute at both ends, dentate, or nearly 
entire, 1-2” long, 2’-8”” wide; flower-clusters 
mainly axillary, distant; calyx-teeth subulate- 
lanceolate, shorter than or equalling the tube; 
corolla 6-8” long, red or purple to white and 
variegated, twice the length of the calyx. 

In waste places and on ballast, New Brunswick 
to Michigan, New Jersey and Indiana. Adventive 
or naturalized from Europe. Native also of Asia. 
Dog-nettle. July—Oct. 


\|F H 
2. Galeopsis Tetrahit L. Hemp-Nettle. Hemp 
Dead Nettle. Ironwort. Fig. 3600. 
Galeopsis Tetrahit L. Sp. Pl. 579. 1753. 

A coarse and rough-hairy herb, the stem rather 
stout, branched, 1°-3° high, swollen under the joints. 
Leaves ovate, membranous, slender-petioled, acumi- 
nate at the apex, rounded or narrowed at the base, 
coarsely dentate, 2’-5’ long, 4’-24’ wide; flower-clus- 
ters axillary, dense, or in a short leafy-bracted spike ; 
calyx-teeth needle-pointed, bristly, as long as or 
longer than the tube; corolla 8’—12” long, pink or 
pale purple variegated with white, about twice the 
length of the calyx. 

In waste places, Newfoundland to British Columbia 
and Alaska, south to North Carolina, West Virginia and 
Michigan. Naturalized from Europe. Native also of 
Asia. Bee-, dog- or blind-nettle. Stinging or flowering 
nettle. Nettle-, wild- or bastard-hemp. Simon’s-weed. 
June—Sept. 


18. LEONURUS L. Sp. Pl. 584. 1753. 


Tall erect herbs, with palmately cleft, parted or dentate leaves, and small white or pink 
flowers verticillate in dense axillary clusters. Calyx tubular-campanulate, 5-nerved, nearly 
wegular and equally 5-toothed, the teeth rigid, subulate or aristate. Tube of the corolla 
included or slightly exserted, its limb 2-lipped; upper lip erect, concave or nearly flat, entire; 
lower lip spreading or deflexed, 3-lobed, the middle lobe broad, obcordate or emarginate. 
Stamens 4, didynamous, the anterior pair the longer, ascending under the upper lip of the 
corolla; anthers 2-celled, the sacs mostly parallel. Ovary deeply 4-parted; style 2-cleft at 
the summit. Nutlets 3-sided, smooth. [Greek, lion’s-tail.] 

About 10 species, natives of Europe and ‘Asia. Type species: Leonurus Cardiaca L. 


Lower leaves palmately 2-5-cleft, the upper 3-cleft._ 1. L. Cardiaca. 
Leaves deeply 3-parted, the segments cleft and incised. 2. L. sibiricus. 
Leaves coarsely dentate or incised-dentate. 3. L. Marrubiastrum. 


120 LABIATAE. Vo. III. 


1. Leonurus Cardiaca L. Motherwort. 
Fig. 3607. 
Leonurus Cardiaca L. Sp. Pl. 584. 1753. 


Perennial, puberulent ; stem rather stout, strict, 
commonly branched, 2°-5° tall, the branches 
straight and ascending. Leaves membranous, 
slender-petioled, the lower nearly orbicular, pal- 
mately 3-5-cleft, 2-4’ broad, the lobes acumi- 
nate, incised or dentate; upper (floral) leaves 
narrower, oblong-lanceolate or rhombic, 3-cleit, 
or the uppermost merely 3-toothed; flower-clus- 
ters numerous, exceeded by the petioles; calyx- 
teeth lanceolate, subulate, somewhat spreading, 
nearly as long as the tube; corolla pink, purple 
or white, 3’-5” long, its tube with an oblique 
ring of hairs within, iis upper lip slightly con- 
cave, densely white-woolly without, the lower lip 
mottled; anther-sacs parallel. 


In waste places, Nova Scotia to North Carolina, 
South Dakota, Montana, Kansas and Utah. Na- 
turalized from Europe. Native also of Asia. Also 
called cowthwort. Lion’s-ear. June—Sept. 


2. Leonurus sibiricus L. Siberian Mother- 
wort or Lion’s-tail. Fig. 3608. 
Leonurus, sibiricus L. Sp. Pl. 584. 1753. 


Biennial, puberulent or glabrate; stem stout, 
branched, 2°-6° high, the branches slender. Leaves 
long-petioled, deeply 3-parted into ovate or lan- Ny 
ceolate, more or less cuneate, acute or acuminate HS 
deeply cleft and incised segments, the lobes lan- SS wha Z, 
ceolate or linear, acute; lower leaves sometimes MH NY 
6’ wide, the uppermost linear or lanceolate, slightly Fm 
toothed or entire; clusters numerous, dense, usu- 
ally all axillary; calyx campanulate, 3” long, 
glabrous or minutely puberulent, its bristle-shaped 
teeth slightly spreading, shorter than the tube; 
corolla purple or red, densely puberuient without, 
4-6” long, its tube naked within, the upper lip 
arched; anther-sacs divergent. 


In waste and cultivated soil, southern Pennsylva- 
nia and Delaware. Bermuda. WNaturalized from 
eastern Asia. Widely distributed in tropical Amer- 
ica as a weed. May-—Sept. 


3. Leonurus Marrubiastrum L. Hoar- 
hound Motherwort or Lion’s-tail. 
Fig. 3609. 

Leonurus Marrubiastrum L. Sp. Pl. 584. 1753. 


Biennial, puberulent or pubescent; stem stout, 
branched, 2°-5° high. Leaves petioled, ovate 
or ovate-oblong, acute or obtuse at the apex, 
narrowed at the base, coarsely crenate or in- 
cised-dentate, 1’-3’ long, 3-12’ wide, the upper 
narrower; flower-clusters dense, numerous, 
axillary; calyx finely puberulent or glabrate, 
its bristle-shaped somewhat spreading teeth 
mostly shorter than the tube; corolla nearly 
white, glabrate, about 4” long,its tube scarcely 
exceeding the calyx, naked within, its lower 
lip ascending. 


In waste places, southern Pennsylvania and 
Delaware. Recorded from New Jersey. Natural- 
ized from Europe. Native also of Asia. June- 
Sept. 


GENUS IQ. MINT FAMILY. i2I 


19. LAMIUM [Tourn.] L. Sp. Pl. 579. 1753. 


Annual or perennial mostly diffuse herbs, with crenate dentate or incised, usually cordate 
leaves, and rather small flowers, verticillate in axillary and terminal clusters. Calyx tubular- 
campanulate, about 5-nerved, 5-toothed, the teeth sharp, equal or the upper ones longer. 
Tube of the corolla mostly longer than the calyx, dilated above, its limb 2-lipped; upper lip 
concave, erect, usually entire, narrowed at the base; lower lip spreading, 3-cleft, the middle 
lobe emarginate, contracted at the base, the lateral ones sometimes each with a tooth-like 
appendage. Stamens 4, didynamous, ascending under the upper lip of the corolla, the anterior 
pair the longer; anthers 2-celled, the sacs divaricate, often hirsute on the back. Ovary deeply 
4-parted; style 2-cleft at the summit. Nutlets smooth or tuberculate. [Greek, throat, from 
the ringent corolla.] 

About 40 species, natives of the Old World, known as Dead-Nettle or Hedge Dead-Nettle. 
Type species: Lamium purpureum L. 


Upper leaves sessile or clasping. 1. L. amplexicaule. 
Leaves all petioled. 
Flowers red or purple. 


Corolla 6”—9” long; leaves not blotched. 2. L. purpureum. 
Corolla 10”—-12” long; leaves commonly blotched. 3. L. maculatum. 
Flowers white. 4, L. album. 


1. Lamium amplexicatle L. MHenbit. 
Greater Henbit. Henbit Dead 
Nettle. Fig. 3610. 


Lanuum amplexicaule L. Sp. Pl. 579. 1753. 


Biennial or annual, sparingly pubescent; 
stems branched from the base or also from 
the lower axils, slender, ascending or decum- 
bent, 6-18’ long. Leaves orbicular or nearly 
so, coarsely crenate, 4’-13’ wide, rounded at 
the apex, the lower slender-petioled, mostly 
cordate, the upper sessile and more or less 
clasping; flowers rather few in axillary and 
terminal clusters; calyx pubescent, its teeth 
erect, nearly as long as the tube; corolla pur- 
plish or red, 6-8” long, its tube very slender, 
the lateral lobes of its lower lip very small, 
the middle one spotted; upper lip somewhat 
_ pubescent; flowers sometimes cleistogamous. 2 


In waste and cultivated ground, New Bruns- ay 
wick to Ontario, Minnesota, British Columbia, 
Florida, Arkansas and California and in Ber- W 
muda and Jamaica, Naturalized from Europe. 
Native also of Asia. Feb—Oct. 


2. Lamium purpureum L. Red 
Dead Nettle. Fig. 3611. 


Lamium purpureum L. Sp. Pl. 579. 1753. 


Annual, slightly pubescent, branched from 
the base and sometimes also above; stems 
stout or slender, decumbent, 6-18’ long. Leaves 
crenate or crenulate, the lower orbicular or 
broadly ovate, slender-petioled, rounded at the 
apex, cordate at the base, the upper ovate, 
short-petioled, sometimes acute at the apex, 
3/-1}’ long; flowers in axillary and terminal 
clusters; calyx teeth narrowly lanceolate, acu- 
minate, slightly longer than the tube, spread- 
ing, at least in fruit; corolla purple-red, rarely 
exceeding 34’ long, its tube rather stout, the 
lateral lobes of its lower lip reduced to I or 2 
short teeth, its middle lobe spotted; upper lip 
very pubescent. 

In waste and cultivated soil, Newfoundland to 
Pennsylvania, North Carolina and Missouri. Also 
in ballast about the northern seaports. Natural- 
ized or adventive from Europe. Native also of 
Asia. Old names, red or sweet archangel; day-, 
dog-, french- or deaf-nettle. Rabbit-meat. April- 

ct. 


122 LABIATAE. Vor. ITI. 


Lamium hybridum Vill., occasionally found in waste and cultivated grounds, introduced from 
Europe, differs by its more deeply and incisely toothed leaves. 


3. Lamium maculatum L. Spotted Dead Nettle. Variegated Dead Nettle. 
Fig. 3612. 


L. maculatum L. Sp. Pl. Ed. 2, 809. 1763. 


Perennial, somewhat pubescent; stems 
mostly slender, commonly branched, de- 
cumbent or ascending, 8’-13° long. Leaves 
crenate or incised-crenate, all petioled, usu- 
ally longitudinally blotched along the mid- 
rib, broadly ovate or  triangular-ovate, 
acute or obtuse, truncate or cordate at the 
base, 1-2’ long, or some of the lower ones 
much smaller and nearly orbicular; clusters 
few-flowered, mainly axillary; calyx-teeth 
lanceolate-subulate, as long as or longer 
than the tube, spreading; corolla 10-12” 
long, purple-red, its tube short, contracted 
near the base, with a transverse ring of 
hairs within, the lateral lobes of its lower 
lip very small. 


Along roadsides, escaped from gardens, 
Maine and Vermont to Virginia. Native of 
Europe and Asia. May-—Oct. 


4. Lamium album L. White Dead 
Nettle. Fig. 3613. 
Lamium album L. Sp. Pl. 579. 1753. 


Perennial, pubescent; stems decumbent or 
ascending, rather stout, simple or branched, 
1°-12° long. Leaves ovate, crenate, dentate 
or incised, all petioled, acute or acuminate at 
the apex, cordate or truncate at the base, 
1-3’ long, or the lower shorter and obtuse; 
clusters mostly axillary; calyx-teeth very 
slender, subulate, spreading, usually longer 
than the tube; corolla white, about 1’ long, 
its tube short, stout, contracted near the base, 
with an oblique ring of hairs within, the lat- 
eral lobes of its lower lip each with a slender 
tooth. 

In waste places, Ontario to Massachusetts and 
Virginia. Also in ballast about the northern sea- 
ports. Naturalized or adventive from Europe. 
Old names, white archangel; day-, blind-, dumb- 


or bee-nettle; snake-flower, suck-bottle. April— 
Oct. 


20. BALLOTA L. Sp. Pl. 582. 1753. 


Perennial pubescent or tomentose herbs, some species shrubby, with dentate or crenate 
leaves, and small bracted flowers in axillary clusters. Calyx tubular-funnelform, 10-nerved, 
5-10-toothed, the teeth dilated at the base, or sometimes connate into a spreading limb. 
Corolla-tube about as long as the calyx, provided with a ring of hairs within, the limb 
strongly 2-lipped; upper lip erect, concave, emarginate, lower lip spreading, 3-lobed, the 
middle lobe emarginate or obcordate. Stamens 4, didynamous, ascending under the upper 
lip of the corolla, the anterior pair the longer; anther-sacs divergent at maturity. Ovary 
deeply 4-lobed; style 2-cleft at the summit. Nutlets ovoid, smooth. [The Greek name.] 


About 30 species, natives of the Old World, most numerous in the Mediterranean region, the 
following typical. 


GENUS 20. MINT FAMILY. 123 


1. Ballota nigra L. Black or Fetid Hoarhound. 
Fig. 3614. 
Ballota nigra L. Sp. Pl. 582. 1753. 


Herbaceous, puberulent or pubescent, ill-scented; 
stem usually branched, erect, 12°-3° high, its hairs 
mostly reflexed. Leaves slender-petioled, ovate, or the 
lower nearly orbicular, acute or obtuse at the apex, 
coarsely dentate, thin, narrowed, truncate or subcor- 
date at the base, 1-2’ long; clusters numerous, several- 
flowered, dense; bractlets subulate, somewhat shorter 
than the calyx; calyx about 4” long, its teeth lanceo- 
late, sharp, bristle-pointed, spreading in fruit; corolla 
6-9” long, reddish-purple to whitish, its upper lip 
pubescent on both sides; nutlets shining. 

In waste places, eastern Massachusetts to Pennsylva- 


nia. Naturalized from Europe. June-Sept. Black arch- 
angel. Hairhound. Henbit. Bastard-hoarhound. 


21. STACHYS [Tourn | L2 Sp: Pl 580, tazse: 


Annual or perennial glabrous pubescent or hirsute herbs, with small or rather large 
purple yellow red or white flowers, loosely verticillate-clustered in terminal dense or inter- 
rupted spikes, or also in the upper axils. Calyx mostly campanulate, 5-10-nerved, 5-toothed, 
the teeth nearly equal in our species. Corolla purple in our species, its tube narrow, not 
exceeding the calyx, the limb strongly 2-lipped; upper lip erect, concave, entire or emarginate; 
lower lip spreading, 3-cleft, the middle lobe broader than the lateral ones, sometimes 2-lobed. , 
Stamens 4, didynamous, ascending under the upper lip of the corolla, the anterior pair the 
longer, sometimes deflexed or twisted after anthesis; anthers contiguous in pairs, 2-celled, 
the sacs mostly divergent. Ovary deeply 4-lobed; style 2-cleft at the summit into subulate 
lobes. Nutlets ovoid or oblong. [Greek, a spike, from the spicate inflorescence. ] 


About 160 species, of wide geographic distribution in the north temperate zone, a few in South 


America and South -Africa. Besides the following, some 20 others occur in the southern and 
southwestern United States. Type species: Stachys germanica L. 


‘A. Perennials. 


1. Leaves, or some of them narrowed or acute at the base, linear to oblong or lanceolate. 
Stem glabrous or nearly so, sometimes sparingly hirsute on the angles; leaves nearly or quite 


entire. Bae 
Leaves 1”—21%4” long, linear. 1. S. hyssopifolia. 
Leaves 2”—5” wide, oblong to linear-oblong. 2. S. atlantica. 


Stem retrorsely hirsute; leaves serrate. ; 
Stem hirsute only on the angles; leaves slightly pubescent. 3. S. ambigua. 
Stem densely hirsute all over; leaves densely pubescent. 4. S. arenicola. 


2. Leaves rounded, cordate or truncate at the base, oblong, ovate or lanceolate. 
Leaves all subsessile or short-petioled. jf 
Glabrous or very nearly so, the stem-angles sparsely bristly. 
Stem retrorsely hirsute ; leaves pubescent. 
Leaves lanceolate to oblong-lanceolate. 
Leaves ovate to oblong-ovate. 
Leaves, at least the lower, slender-petioled. 
Stem glabrous, or nearly so. 
Stem retrorsely hirsute or villous. 
Petioles less than one-fourth as long as the blades. 


. S. latidens. 


. S. palustris. 
. S. Nuttallii. 


anTrn un 


. S. tenuifolia. 


Pubescence hirsute. 9. S. aspera. 
Pubescence short, dense. 10. S. salvioides. 
Petioles, at least those of lower leaves, about one-half as long as the deeply cordate 

blades. 11. S. cordata. 


B. Annuals. 


Plant low, hirsute. | 12. S. arvensis. 
Plant tall, densely villous. 13. S. germanica. 


LABIATAE. Vor. ITT. 


1. Stachys hyssopifélia Michx. Hyssop 
Hedge Nettle. Fig. 3615. 


et S. palustris Walt. Fl. Car. 162. 1788. Not L. 1753. 
iy S. hyssopifolia Michx. Fl. Bor. Am. 2: 4. 1803. 
Perennial, glabrous or very nearly so through- 
out, sometimes slightly hirsute at the nodes; 
stem slender, usually branched, erect or nearly 
so, 1°-14° long. Leaves thin, linear, acute at 
both ends, or the uppermost rounded at the 
base, short-petioled or sessile, entire, or spar- 
ingly denticulate with low teeth, 17-2’ long, 
1-23” wide, the uppermost reduced to short 
floral bracts; clusters few-—several-flowered, 
forming an interrupted spike; calyx glabrous 
or slightly hirsute, 2-3” long, its teeth lan- 
ceolate-subulate, nearly as long as the tube; 
corolla about 7” long, light purple, glabrous. 


In fields and thickets, Massachusetts to Florida, 
Indiana, Michigan and Virginia. July—Sept. 


2. Stachys atlantica Britton. Coast Hedge Nettle. 
Fig. 3616. 


Stachys atlantica Britton, Man. 792. 1901. 


Perennial, glabrous or with a few hairs at the nodes of 
the stem. weak, diffuse, 8-16’ long. Leaves thin, oblong 
or linear-oblong, obtuse or obtusish at the apex, narrowed, 
or the upper sometimes rounded at the base, remotely den- 
ticulate or entire, spreading, 2’ long or less, 2”—5’’ wide; 
fruiting calyx glabrous, broadly campanulate, about 23” 
long, its teeth triangular-ovate, acuminate, more than half 
as long as the tube; corolla purplish. 


In wet meadows and marshes, Long Island to eastern Penn- 
sylvania. Perhaps a wet-ground race of the preceding species. 
Aug.-Sept. 


3. Stachys ambigua (A. Gray) Britton. 
Dense-flowered Hedge Nettle. Fig. 3617. 
Stachys hyssopifolia var. ambigua A. Gray, Syn. FI. 
Pre IEEE in esky  ansiyst 
S. ambigua Britton, Mem. Torr. Club 5: 285. 1894. 
Perennial; stem slender, erect, retrorsely his- 
pid, at least below, simple or sparingly branched, 
1°-2° high. Leaves oblong, oblong-lanceolate, 
or linear, pubescent or glabrate, acuminate or 
acute at the apex, narrowed at the base, 2’—10” 
wide, 1-3’ long, serrulate; clusters in a terminal 
rather dense spike, and usually also in the upper 
axils; calyx more or less hirsute, 23” long, its 
lanceolate-subulate teeth more than one-half as 
long as the tube; corolla nearly as in S. hyssopi- 
folia. 


In moist soil, Massachusetts to Pennsylvania, 
Georgia, Wisconsin and Kentucky. July—Aug. 


GENUS 21. MINT FAMILY. 125 


4. Stachys arenicola Britton. Sand Hedge 
Nettle. Fig. 3618. 


Stachys arenicola Britton, Man. 792. 1901. 


Perennial, very densely pubescent, 13°-3° high. 
Leaves lanceolate, narrowed at the base, sessile, 
finely crenate-dentate, acuminate, 23’-4’ long, the 
upper much smaller; upper bracts little longer than 
the flowers; calyx-teeth subulate-acicular, about as 
long as the tube; corolle about 5” long. 


In sandy soil, southern New York to Illinois and 
Michigan. July—Sept. 


5. Stachys latidens Small. Broad-toothed 
Hedge Nettle. Fig. 3619. 


Stachys latidens Small; Britton, Man. 793. 1901. 


Perennial; stem glabrous almost to the inflorescence, 
erect, 1°-23° tall, mostly simple, slightly rough on the 
angles. Leaves thin, various, the lower oval or ovate 
to oblong, 14-4’ long, the upper longer, narrower, 
oblong-elliptic to lanceolate, acute or acuminate, all 
crenate-serrate with rather small teeth, rather abruptly 
narrowed or subcordate at the base, short-petioled; in- 
florescence closely pubescent, the clusters few; calyx 
minutely pubescent, often sessile, 23’”—3” long, the tube 
campanulate, the teeth triangular, very short; corolla 
about 5” long, purplish. 


On mountain slopes and summits, Virginia to North 
Carolina and Tennessee. June—Aug. 


6. Stachys paltstris L. Hedge Nettle. 
Marsh or Clown’s Woundwort. Fig. 3620. 


Stachys palustris L. Sp. Pl. 580. 1753. 


Perennial, hirsute or pubescent all over; stem 
erect, strict, simple or somewhat branched, com- 
monly slender, and retrorse-hispid on the angles, 
1°-4° high. Leaves firm, lanceolate, oblong, or 
oblong-lanceolate, sessile, or very short-petioled, 
acuminate or acute at the apex, truncate, cordate 
or subcordate at the base, 2’-5’ long, 3’-1’ wide, 
crenulate or dentate; flower-clusters forming an 
elongated interrupted spike, sometimes also in 
the upper axils; flowers 6-10 in a whorl; calyx 
pubescent, its subulate teeth more than one-half 
as long as the tube; corolla purplish to pale red, 
purple spotted, 6’-8” long, its upper lip pubescent. 

In moist soil, Newfoundland to Oregon, south to 
southern New York, Illinois, Michigan, and in the 
Rocky Mountains to New Mexico. Also in Europe 
and Asia. June—Sept. Old names, clown’s-heal or 


all-heal. Cock-head. Dead nettle. Rough weed. 
June-Sept. 


126 LABIATAE. Vot. III. 


7. Stachys Nuttallii Shuttlw. Nuttall’s Hedge 
Nettle. Fig. 3621. 


Stachys Nuttallii Shuttlw.; DC. Prodr. 12: 469. 1848. 


Perennial, conspicuously hirsute, bright green; stem 
stiff, erect, 14°-33° tall, simple. Leaves thinnish, ob- 
long, oblong-lanceolate or ovate-lanceolate, acuminate, 
serrate-dentate, rounded or truncate at the base, short- 
petioled, 24’ long; spike interrupted; bracts surpass- 
ing the calyx, the upper ones with 3 tooth-like lobes; 
calyx-teeth triangular-lanceolate, acuminate, about +4 
as long as the tube; corolla purple, about 5” long, pu- 
bescent; nutlets about 1” long. 


In woods and on mountain slopes, Maryland and Vir- 
ginia to Tennessee. June—Aug. 


YY 
8. Stachys tenuifolia Willd. Smooth Wie 


YA 


Hedge Nettle. Fig. 3622. 


Stachys tenuifolia Willd. Sp. Pl.3: 100. 18o0t. 
S. glabra Ridd. Suppl. Cat. Ohio Pl. 16. 1836. 
S. cincinnatensis Kuntze, Rev. Gen. Pl. 531. 

1891. : 


Perennial; stem quite smooth, or slightly 
scabrous on the angles, slender, erect or 
ascending, usually branched, 1°-23° high. 
Leaves lanceolate, oblong, or ovate-lanceo- 
late, slender-petioled, thin, acuminate at the 
apex, obtuse or subcordate at the base, 
sharply dentate or denticulate, dark green, 
2’-5’ long, 3-2’ wide; clusters several or 
numerous in terminal spikes, or also in the 
upper axils; calyx glabrous, or sparingly 
hirsute, 2” long, its teeth lanceolate, acute, 
one-half as long as the tube or more; co- 
rolla about 6’-8” ! 


long, pale red and purple. 


In moist fields and thickets, New York to Illinois, Kansas, North Carolina and Louisiana. 
Ascends to 4000 ft. in North Carolina. June—Aug. 


g. Stachys A4spera Michx. Rough Hedge 
Nettle or Woundwort. Fig. 3623. 


Stachys aspera Michx. Fl. Bor. Am. 2: 5. 1803. 
Stachys hispida Pursh, Fl. Am. Sept. 407. 1814. 
S. palustris var. aspera A. Gray, Man. Ed. 2, 317. 1856. 


Perennial, rough hairy; stem erect or ascending, 
simple or branched, 2°-4° high, commonly retrorse- 
hispid on the angles. Leaves firm, oblong, oblong- 
lanceolate, or ovate-oblong, crenate-dentate, acute or 
acuminate at the apex, truncate, rounded or cordate 
at the base, 2’-6’ long, 4’-22’ wide, the lower slender- 
petioled, the upper short-petioled; spike terminal, 
mostly interrupted; clusters sometimes also in the 
upper axils; calyx about 3” long, hirsute or glabrate, 
its teeth triangular-lanceolate, acuminate, about one- 
half as long as the tube; corolla red-purple, about 3’ 
long, its upper lip pubescent. 


In moist soil, Ontario to Massachusetts, Florida, Min- 
nesota and Louisiana. Ascends to 5300 ft. in Virginia. 
June-Sept. Base hoarhound. 


GENUS 21. MINT FAMILY. 127 


10. Stachys salvioides Small. Sage-like 
Hedge Nettle. Fig. 3624. 


Stachys salvioides Small; Fl. SE. U.S. 1032. 1903. 


Perennial, rather finely and often closely puberu- 
lent or puberulent-pubescent; stem erect, 1°-3° tall, 
commonly branched, rough along the angles. Leaves 
firm, oblong to oblong-ovate or rarely ovate, 23’—5’ 
long, or shorter on the lower part of the stem, 
pubescent on both sides, acute or acutish, crenate, 
truncate or cordate at the base, slender-petioled; 
clusters several; calyx sessile or nearly so, becom- 
ing 22” long, finely pubescent, the tube campanulate- 
turbinate, the teeth triangular at mutarity; corolla 
nearly 5” long, pale purple. 


In stony soil, Virginia and West Virginia to Tennes- 
see. June-Aug. : 


11. Stachys cordata Riddell. Light-green 
Hedge Nettle. Fig. 3625. 


Stachys cordata Riddell, Suppl. Cat. Ohio Pl. 15. 1836. 
pee palustris var. cordata A. Gray, Man. Ed. 2, 317. 

1856. 

Perennial, hirsute, pale green; stem slender, weak,” 
mostly simple, ascending or reclining, 2°-3° long. 
Leaves membranous, flaccid, ovate, oblong or ovate- 
lanceolate, acuminate, acute or the lowest obtuse 
at the apex, nearly all of them cordate at the base, 
dentate or crenate all around, long-petioled, 3’-6’ 
long, 1-3’ wide, the lowest petioles nearly as long 
as the blades; spike interrupted; calyx-teeth subu- 
late-lanceolate, about half the length of the tube; 
corolla purplish, pubescent or puberulent, about 5” 
long. 

In woods and thickets, Ohio, Indiana and Illinois to 
North Carolina and Tennessee. Ascends to 2100 ft. in 
Virginia. July—Aug. 


12. Stachys arvénsis L. Corn or Field Woundwort. Fig. 3626. 
Stachys arvensis L. Sp. Pl. Ed. 2, 814. 1763. 


Annual, hirsute ; stem very slender, diffusely branch- 
ed, decumbent or ascending, 3’-2° long. Leaves 
ovate or ovate-oblong, thin, long-petioled, obtuse at 
the apex, crenate all around, cordate or the upper 
rounded at the base, about 1’ long; lower petioles 
commonly as long as the blades; clusters 4-6-flow- 
ered, borne in the upper axils and in short terminal 
spikes; calyx about 3” long, its teeth lanceolate, 
acuminate, nearly as long as the tube; corolla pur- 
plish, 3’-5” long. 

In waste places, Maine and Massachusetts to Ne- 
braska, in ballast about the eastern seaports, and in the 


West Indies, Mexico and South America. Naturalized 
from Europe. July—Oct. 


Stachys Aannua L., another European annual species, 
the stems erect, glabrous or nearly so, the yellow corolla 
much- longer than the calyx, has been found in ballast 
and waste grounds about the seaports. 


128 LABIATAE. Vor. ik 


13. Stachys germanica L. Downy Wound- 
wort. Mouse-ear. Fig. 3627. 


Stachys germanica L. Sp. Pl. 581. 1753. 


Annual; stem erect, simple, or somewhat branched, 
1°-3° high, densely villous. Leaves oval, ovate or 
lanceolate, crenate-dentate, villous, the lower round- 
ed or subcordate at the base, long-petioled, mostly 
obtuse at the apex, the upper short-petioled or ses- 
sile, narrowed at both ends; clusters of flowers. 
A dense, borne in most of the axils; bracts lanceolate, 
Ses) half as long as the calyx; calyx-teeth ovate, acumi- 
"] nate, awned; corolla purple, its tube about as long 
! as the calyx. 


Roadsides near Guelph, Ontario. Adventive from 
Europe. July—Sept. 


22, BETONICA [Tourn.] L. Sp. Pl. 573. 
a J 1753- 


Annual or perennial herbs, similar to Stachys. Lower leaves very long-petioled. Ver- 
ticils many-flowered, in terminal spikes. Calyx nearly equally 5-toothed, 5-10-nerved. Corolla 
purple, the tube exceeding the calyx, the limb strongly 2-lipped; upper lip concave; lower 
3-cleft, spreading. Stamens and pistil as in Stachys, but the anther-sacs parallel in some 


species. Nutlets ovoid, rounded above. [The classical 1) 
Latin name of wood betony.] N17 
Ten species, or more, natives of Europe and Asia, the NES 
“ following typical. AN Fi 
. aMccz's - a 
1. Betonica officinalis L. Betony. Wood Betony. ~# Ue i 
Bishop’s-wort. Fig. 3628. Fp Bees 
Betonica officinalis L. Sp. Pl. 573. 1753. NGS: 
Stachys Betonica Benth. Lab. Gen. & Sp. 532. 1834. AWE 
Perennial, pilose or glabrate, deep green; stem slen- &% SM AN 
der, erect, usually simple, 1°-3° tall. Leaves oblong or p ay 
ovate, obtuse at the apex, crenate all around, firm, cor- oF Ae 
date or truncate at the base, the basal and lower ones  N 


very long-petioled, the blades 3-6’ long, the upper dis- 
tant, short-petioled or nearly sessile; lower petioles 
14-4 times as long as the blades; spike short, dense; 
bracts ovate, mucronate, about as long as the calyx; 
calyx-teeth acicular, half as long as the tube or more; 
corolla-tube exserted. 


In a thicket at Newton, Mass. Fugitive from Europe. 
Herb christopher. Wildhop. July—Sept. 


23. SALVIA [Tourn.] L. Sp. Pl. 23. 1753. 


Herbs, or some species shrubs, with clustered usually showy flowers, the clusters mostly 
spiked, racemed, or panicled. Calyx ovoid, tubular or campanulate, mostly naked in the 
throat, 2-lipped; upper lip entire or 3-toothed; lower lip 2-cleft or 2-toothed. Corolla 
strongly 2-lipped; upper lip usually concave, sometimes arched, entire, emarginate or 2-lobed; 
lower lip spreading or drooping, 3-cleft or 3-lobed. Anther-bearing stamens 2 (the posterior 
pair wanting or rudimentary) ; filaments usually short; connective of the anthers transverse, 
linear or filiform, bearing a perfect anther-sac on its upper end, its lower end dilated, capi- 
tate or sometimes bearing a small or rudimentary one. Ovary deeply 4-parted; style 2-cleft 
at the summit. Nutlets smooth, usually developing mucilage and spiral tubes when wetted. 
[Latin, salvus, safe from its healing virtues.] 

About 500 species, of wide distribution in temperate and tropical regions. Besides the follow- 
ing, some 25 others occur in southern and western North'America. Type species: Salvia officinalis L. 


Leaves mostly basal, only 1-3 pairs on the stem. 
Leaves lyrate-pinnatifid or repand ; upper corolla-lip short. 1. S. lyrata. 
Leaves crenulate; upper lip arched, longer than the lower. 2. S. pratensis. 
Stem leafy, bearing several pairs of leaves. 
Leaves narrowly oblong, or lanceolate. 
Corolla 10’—15” long, its tube exserted. 
Corolla 4”—6” long, its tube not exserted. 
Leaves ovate, or broadly oval. 
Upper corolla-lip short, not exceeding the lower. 


. S. Pitcheri. 
. S. lanceifolia. 


fw 


GENUS 23. MINT FAMILY. 


Leaves merely crenate or crenulate; fruiting calyx spreading. 
Leaves pinnatifid, sinuate or incised; fruiting calyx deflexed. 
Upper lip of corolla arched, longer than the lower. 


1. Salvia lyrata L. Lyre-leaved Sage. 
Wild Sage. Cancer-weed. Fig. 3629. 
Salvia lyrata L. Sp. Pl. 23. 1753. 


Perennial or biennial, hirsute or pubescent; 
stem slender, simple, or sparingly branched, erect, 
1°-3° high, bearing I or 2 distant pairs of small 
leaves (rarely leafless), and several rather distant 
whorls of large violet flowers. Basal leaves tufted, 
long-petioled, obovate or broadly oblong, lyrate- 
pinnatifid or repand-dentate, thin, 3-8’ long; stem- 
leaves similar, or narrower and entire, sessile, or 
short-petioled; clusters distant, about 6-flowered ; 
calyx campanulate, the teeth of its upper lip subu- 
late, those of the lower longer, aristulate; corolla 
about I’ long, the tube very narrow below, the 
upper lip much smaller than the lower; fila- 
ments slender; anther-sacs borne on both the 
upper and lower ends of the connective, the lower 
one often smaller. 

In dry, mostly sandy woods and thickets, Con- 


necticut to Florida, west to Illinois, Arkansas and 
Texas. Corolla rarely undeveloped. May-July. 


I 

2. Salvia praténsis L. Meadow Sage. 
Fig. 3630. 

Salvia pratensis L. Sp. Pl. 25. 1753. 

Perennial, pubescent or puberulent; stem erect, 
rather stout, simple or little branched, sparingly 
leafy. Basal leaves long-petioled, ovate, oblong or 
ovate-lanceolate, irregularly crenulate, obtuse at 
the apex, rounded or cordate at the base, thick, 
rugose, 2-7’ long; stem-leaves much smaller, nar- 
rower, commonly acute, sessile or nearly so; 
clusters spicate, the spike elongated, interrupted ; 
calyx campanulate, glandular-pubescent, the teeth 
of the upper lip minute, those of the lower long, 
subulate; corolla purple, minutely glandular, its 
upper lip strongly arched, mostly longer than the 
lower; lower end of the connective with a small 
or imperfect anther-sac. 

Atlantic Co., N. J. Fugitive or adventive from 
Europe. May-July. 


3. Salvia Pitcheri Torr. Pitcher’s Sage. 
Mall Sage.) Fig: 2631. 

Salvia Pitcheri Torr.; Benth. Lab. 251. 1833. 

Salvia azurea var. grandiflora Benth. in DC. 

Prodr. 12: 302. 1848. 

Perennial, downy; stem stout, branched or 
simple, erect, 2°-5° high; branches nearly erect. 
Leaves linear or linear-oblong, dentate to en- 
tire, sessile, or narrowed at the base into short 
petioles, firm, 2’-5’ long, 2’-8” wide, the up- 
permost reduced to small bracts; clusters in 
long dense terminal spikes, or the lower ones 
distant; calyx oblong-campanulate, densely and 
finely woolly, about 3” long, its upper lip en- 
tire, obtuse, the lower with 2 ovate acute teeth; 
corolla blue or white, finely pubescent without, 
1’ long, its lower lip broad, sinuately 3-lobed, 
longer than the concave upper one; lower ends 
of the connective dilated, often adherent to 
each other, destitute of anther-sacs. 

On dry plains, Illinois and Minnesota to Mis- 
souri, Kansas, Colorado and Texas. July—Sept. 


9 


130 


5. Salvia urticifolia L. Nettle-leaved or 
Wild Sage. 

Salvia urticifolia L. Sp. Pl. 24. 
Perennial, pubescent, or nearly glabrous; stem 


rather slender, ascending or 


glandular above, adit 
erect, 1°-2° high. Leaves thin, ovate, 2-4’ long, 


acute at the apex, abruptly contracted below into 


LABIATAE. Vo. ITT, 


4. Salvia lanceifolia Poir. Lance-leaved Sage. Fig. 3632. 


S. lanceolata Brouss. App. Elench. Pl. Hort. Monsp. 15. 
1805. Not Lam. 
2S. reflexa Hornem. Enum. Pl. Hort. Hafn. 34. 1807. 
S. lanceifolia Poir. in Lam. Encycl. Suppl. 5: 49. 1817. 
Annual, puberulent or glabrous; stem leafy, usually 
much branched, erect or diffuse, 6-18 high. - Leaves 
oblong, linear-oblong or oblong-lanceolate, petioled, 
mostly obtuse at the apex and narrowed at the base, 
crenulate-dentate or entire, 1-2’ long, 2-5” wide, the 
upper reduced to lanceolate-subulate, rather persistent 
bracts; flowers mostly opposite, but sometimes 3-4 
together in the terminal spike-like racemes; pedicels 
shorter than the campanulate calyx; calyx 2’—3” long, 
its upper lip ovate, entire, the lower 2-cleft, the teeth 
ovate, mucronate; corolla blue, about 4” long, its lower 
lip narrow, twice as long as the upper; lower ends of 
the connectives dilated; style nearly or quite glabrous. 
On plains, Indiana to Nebraska, Montana, Colorado, 
Texas, Arizona and Mexico. Recorded as introduced in 
Ohio. May—Sept. 


irregularly dentate or crenate-dentate, usually al 


margined petioles; clusters several-flowered, in 
terminal interrupted spikes; bracts early decidu- 
ous; pedicels about as long as the calyx; calyx 
oblong-campanulate, about 3” long, the upper lip 
minutely 3-toothed, the lower 2-cleft, its teeth 
triangular-lanceolate, 


fruit; corolla puberulent without, 6-8” long, a 
blue and white, the lower lip broad, 3-lobed, AY) 
twice as long as the upper; lower ends of the Si 
connectives dilated; style bearded. 

In woods and thickets, Pennsylvania to Kentucky, 
April-June. 


south to Georgia and Louisiana. 


\ 


spreading in 


a 4 
We 


YP 


6. Salvia Verbenaca L. Wild Sage. 
Wild Clary. Fig. 3634. 
Salvia Verbenaca L. Sp. Pl. 25. 1753. 


Perennial; stem glandular-pubescent, erect, 
simple or sparingly branched, 1°-2° high. Leaves 
ovate, ovate-lanceolate or oblong, coarsely and 
irregularly incised-dentate or pinnatifid, peti- 
oled, or the uppermost sessile, the lower 3’-8' 
long, obtuse at the apex, cordate at the base, 
nearly glabrous, the upper acute, much smaller; 
floral bracts broadly ovate, short; clusters sev- 
eral-flowered in elongated terminal interrupted 
spikes; pedicels shorter than the calyx; calyx 
deflexed in fruit, 3-4” long, its upper lip re- 
curved-spreading, with 3 minute connivent 
teeth, the lower one with 2 lanceolate acumi- 
nate mucronate teeth; corolla blue, about 4” 
long, its upper lip nearly straight, scarcely 
longer than the lower; lower ends of the con- 
nectives dilated and adnate to each other. 

In waste places, Ohio to South Carolina and 
Georgia. Naturalized from Europe. Native also 
of Asia. Eye-seed. Vervain. June—Aug. 

Salvia verticillata L., a European species, 
with rough lyrate leaves, has been found wild in 
Pennsylvania. 


GENUS 23. MINT FAMILY. 131 


7. Salvia Sclarea L. Clary. Clear-eye. 
See-bright. Fig. 3635. 
Salvia Sclarea L. Sp. Pl. 27. 1753. 


Biennial or annual, glandular-pubescent; stem 
stout, erect, 2°-34° high. Leaves broadly ovate, 
rugose, acute or obtuse at the apex, cordate at the 
base, irregularly denticulate, the lower long-peti- 
oled, 6-8’ long, the upper small, short-petioled 
or sessile; clusters several-flowered, numerous in 
terminal spikes; bracts broad, ovate, acuminate, ». 
commonly longer than the calyx, pink or white; AG 
- calyx campanulate, deflexed-spreading in fruit, 
4-5” long, its lips about equal, the teeth all subu- 
late-acicular; corolla blue and white, about 1’ 
long; upper lip arched, laterally compressed, 
longer than the lower one. 

In fields, Pennsylvania to South Carolina, escaped 
from gardens. Naturalized from Europe. The 
mucilage of the seeds used to clear specks from the 
eye. June—-Aug. 


SA 
AS. oy, ZS 


24. MONARDA L. Sp. Pl. 22. 1753. 


Perennial or annual erect aromatic herbs, with dentate or serrate leaves, and rather 
large white red purple yellowish or mottled flowers, in dense capitate clusters, mostly bracteate 
and bracteolate, terminal and sometimes also axillary, the bracts sometimes brightly colored. 
Calyx tubular, narrow, 15-nerved, nearly or quite equally 5-toothed, mostly villous in the 
throat. Corolla glabrous within, usually puberulent or glandular without, the tube slightly 
dilated above, the limb 2-lipped; upper lip erect or arched, emarginate or entire; lower lip 
spreading, 3-lobed, the middle lobe larger or longer than the others. Anther-bearing stamens 
2, ascending, usually exserted, the posterior pair (staminodia) rudimentary or. wanting; 
anthers linear, versatile, 2-celled, the sacs divaricate, more or less confluent at the base. 
Ovary deeply 4-parted; style 2-cleft at the apex; nutlets ovoid, smooth. [In honor of Nicolas 
Monardes, a Spanish physician and botanist of the sixteenth century. ] 


About 12 species, natives of North America and Mexico. Type species: Monarda fistulosa L. 
Flower-clusters solitary, terminal (rarely also in the uppermost axils) ; stamens exserted. 
Leaves manifestly petioled, the petioles commonly slender. 


Corolla scarlet, 1%4’—2’ long; bracts red. 1. M. didyma. 
Corolla white, pink, or purple, 1’-1%’ long. 
Leaves membranous; corolla slightly pubescent, 10’—12” long. 2. M.clinopodia. 


Leaves thin or firm; corolla pubescent, 1’/-1%4’ long. 
Pubescence spreading; leaves thin. 
Corolla cream-color, pink, or purplish. 
Corolla or bracts deep purple or purple-red. 
Pubescence short, canescent; leaves firm, pale. 
Leaves sessile, or very short-petioled. 
Flower-clusters both axillary and terminal; stamens not exserted. 
Calyx-teeth triangular; corolla yellowish, mottled. 
Calyx-teeth subulate-aristate ; corolla white or purple, not mottled. 
Bracts lanceolate, gradually acuminate. 
Bracts oblong to oval, not acuminate. 


. M. fistulosa. 

. M. media. 

. M. mollis. 

M. Bradburiana. 


nui tw 


. M. punctata. 


ao wn 


. M. pectinata. 
M. dispersa. 


2 


1. Monarda didyma L. Oswego Tea. 
American Bee Balm. Fig. 3636. 
Monarda didyma L. Sp. Pl. 22. 1753. 


Perennial; stem stout, villous-pubescent, or 
glabrate, 2°-3° high. Leaves thin, ovate or ovate- 
lanceolate, petioled, dark green, usually pubescent 
beneath, hairy above, acuminate at apex, rounded 
or narrowed at the base, sharply serrate, 3'-6 
long, 17-3’ wide; lower petioles often 1’ long; 
flower-clusters almost always solitary, terminal; 
bracts commonly red; calyx glabrous without, 
glabrous or Slightly hirsute in the throat, its teeth 
subulate, nearly erect, about as long as the diam- 
eter of the tube; corolla scarlet, 14’-2’ long, 
puberulent; stamens exserted. 

In moist soil, especially along streams, Quebec to 
Ontario and Michigan, south to Georgia and Ten- 
nessee. Ascends to 5200 ft. in North Carolina. Red, 
low or fragrant balm. MHorse- or mountain-mint. In- 
dian’s-plume. July—Sept. 


132 


LABIATAE. Vor. III. 


2. Monarda clinopddia L. Basal Balm. 


Fig. 3637. 
Monarda clinopodia L. Sp. Pl. 22. 1753. 
Pycnanthemum Monardella Michx. Fl. Bor. Am. 2: 8. 
pl. 34. 1803. 


Perennial; stem slender, simple, or with few long 
ascending branches, glabrous or somewhat villous, 
1°-3° high. Leaves lanceolate, ovate or ovate-lan- 
ceolate, membranous, bright green, mostly slender- 
petioled, more or less villous on the veins beneath 
and puberulent above, acuminate at the apex, round- 
ed, narrowed or rarely subcordate at the base,sharply 
serrate, 2-4’ long, +’-2’ wide; clusters solitary, ter- 
minal; bracts pale or white; calyx curved, nearly 
glabrous without, somewhat hirsute in the throat, 
its teeth subulate, slightly spreading, about as long 
as the diameter of the tube; corolla whitish or yel- 
lowish-pink, slightly pubescent, 10’—-12” long; stamens 
exserted. 

In woods and thickets, New York to Illinois, Georgia 
and Kentucky. Recorded from Ontario. Ascends to 
5000 ft. in North Carolina. June—Aug. 

ee 


3. Monarda fistulosa L. Wild Bergamot. 
Fig. 3638. 
Monarda fistulosa L. Sp. Pl. 22. 1753. 


Perennial, villous-pubescent or glabrate; stem 
slender, usually branched, 2°-3° high. Leaves 
thin but not membranous, green, usually slender- 
petioled, lanceolate, ovate or ovate-lanceolate, 
acuminate at the apex, serrate, rounded, narrowed 
or sometimes cordate at the base, 12-4’ long, 
3’-23’ wide; clusters solitary and terminal, or 
rarely also in the uppermost axils; bracts whitish 
or purplish; calyx puberulent or glabrous, densely 
villous in the throat, its subulate teeth rarely 
longer -than the diameter of the tube; corolla 
pubescent, especially on the upper lip, yellowish- 
pink, lilac or purplish, 1-13’ long; stamens ex- 
serted. 


On dry hills and in thickets, Maine and Ontario 
to Minnesota, Florida, Louisiana and Kansas. As- 
cends to 2500 ft. in Virginia. Oswego-tea. June- 
Sept. 


4. Monarda média Willd. Purple Ber- 
gamot. Fig. 3639. 


Monarda media Willd. Enum. 32. 1800. 

Monarda fistulosa var. rubra A. Gray, Syn. FI. 2: 
Part 1, 374s oe. 

M. fistulosa var. media A. Gray, loc. cit. 1878. 


Perennial, sparingly hairy or glabrate; stem 
stout, commonly branched, 2°-33° high. Leaves 
thin, but not membranous, dark green, ovate 
or ovate-lanceolate, or the uppermost lanceo- 
late, slender-petioled, acuminate at the apex, 
sharply toothed, usually rounded at the base, 
3-5’ long, 1’-3’ wide; flower-clusters terminal, 
solitary, large; bracts deep purple, very con- 
spicuous; calyx curved, glabrous; or very 
nearly so, slightly villous in the throat, teeth 
slightly spreading; corolla purple or purple- 
red, about 13’ long, its upper lip pubescent; 
stamens exserted. 


In moist thickets, Maine and Ontario to Penn- 
sylvania and Virginia. June—Aug. 


GEnus 24. MINT FAMILY. 133 


5. Monarda mollis L. Pale Wild Berga- 
mot. Fig. 3640. 


Monarda mollis L. Amoen. ‘Acad. 3: 399. 1787. 
Monarda scabra Beck, Am. Journ. Sci. 10: 260. 1826. 


Perennial; stem slender, puberulent at least 
above, usually branched, 1°-32° high. Leaves 
thick or firm, pale, usually short-petioled, acumi- 
nate or acute at the apex, rounded, narrowed or 
cordate at the base, sharply or sparingly serrate, 
canescent or puberulent, rarely nearly glabrous, 
sometimes with a few scattered spreading hairs 
on the veins or petiole, 1-33’ long, 3’-1’ wide; 
flower-clusters terminal, solitary; bracts green or 
slightly pink, calyx puberulent, often hairy at the 
summit, densely villous in the throat, its short 
pointed teeth nearly erect; corolla yellowish, 
lilac or pink, about 12’ long, pubescent, some- 
times glandular; stamens exserted. 


. On prairies and plains, mostly in dry soil, Maine to 
Ontario, British Columbia, Alabama, Nebraska, Mis- 
souri, Texas and Colorado. June—-Aug. 


6. Monarda Bradburiana Beck. Brad- 
bury Monarda. Fig. 3641. 


Monarda Bradburiana Beck, Am. Journ. Sci. 10: 
260. 1826. 


Perennial, sparingly villous or glabrate; 
stem slender, often simple, 1°-2° high. Leaves 
rather thin, bright green, ovate or ovate-lan- 
ceolate, sessile, or very nearly so, or partly 
clasping by the cordate or subcordate base, 
acuminate at the apex, serrate, 2-33’ long; 
flower-clusters solitary and terminal; bracts 
green or purplish; calyx glabrous outside, hir- 
sute within and narrowed at the throat, its 
teeth long, bristle-pointed, divergent, longer 
than the diameter of the tube; corolla pink or 
nearly white, about 1’ long, its upper lip pu- 
bescent or puberulent, the lower commonly 
purple-spotted; stamens exserted. 


On dry hills or in thickets, Indiana to Alabama, 
west to Kansas and Arkansas. May-July. 


7. Monarda punctata L. Horse-mint. 
Fig. 3642. 
Monarda punctata L. Sp. Pl. 22. 1753. 


Perennial, usually rather densely pubescent or 
downy; stem usually much branched, 2°-3° high. S 
Leaves lanceolate, linear-lanceolate or narrowly _ 
oblong, serrate with low teeth, or nearly entire, ee 
usually acute at both ends, green, manifestly peti- 
oled, 1-3’ long, 2”-7’ wide, often with smaller 
ones fascicled in their axils; flower-clusters axil- 
lary and terminal,. numerous; bracts white or 
purplish, conspicuous, acute; calyx puberulent, 
villous in the throat, its teeth short, triangular- 
lanceolate, acute, not longer than the diameter 
of the tube; corolla yellowish, purple-spotted, 
about 1’ long, the stamens equalling or slightly 
surpassing its pubescent upper lip. 


In dry fields, southern New York to Florida, west 
to Minnesota, Kansas and Texas. Rignum. July—Oct. 


134 LABIATAE. Vor. tie 


8. Monarda pectinata Nutt. Plains Lemon Monarda. Fig. 3643. 


M. pectinata Nutt. Proc. Acad. Phila. (II.) 1: 182. 
1848. 


Annual, puberulent ; stem stout, simple or branch- 
ed, 1°-2° high. Leaves lanceolate or oblong-lan- 
ceolate, mostly sharply serrate or serrulate, acute 
at the apex, narrowed at the base, 17-3’ long, 
2-6’ wide; flower-clusters axillary and terminal, 
several or numerous; bracts pale, usually grayish 
or brownish, gradually awned at the tip; calyx- 
tube nearly glabrous, the throat densely villous, 
the teeth bristle-pointed, barbed, lax, nearly half 
as long as the tube; corolla pink or nearly white, 
not spotted, nearly or quite glabrous, 8-10” 
long; stamens not exserted. 


On dry plains, Nebraska and Colorado to Texas 
and Arizona. Prairie-bergamot. June—Sept. 


Monarda citrioddra Cerv., of Mexico, to which 
this was referred in our first edition, differs by long 
narrow reflexed bracts and shorter calyx-teeth. 


Monarda tenuiaristata (A. Gray) Small [M. aris- 
tata Nutt., not Hook.] of the south-central States, 
with narrower bracts and longer plumose calyx-teeth, 
ranges northward into Kansas. 


g. Monarda dispérsa Small. Purple 
Lemon Monarda. Fig. 3644. 


M. dispersa Small, Fl. SE. U. S. 1038. 1903. 


Annual, puberulent; stem stout, usually 
branched, 1°-23° high. Leaves oblong-spatu- 
late, oblanceolate or oblong, or narrowly ob- 
long to linear on the upper part of the stem, 
1’-4}’ long, shallowly serrate; flower clusters 
axillary and terminal, often numerous and 
usually conspicuous by the broad abruptly 
bristle-tipped purple bracts; calyx-tube longer 
than in M. pectinata, the teeth usually longer \ 


EE 


Ay 


and more slender, fully half as long as the 
tube; corolla pale, usually pink, puberulent, 
11-13” long; stamens mostly not exceeding 
the upper lip. 


Wh 
On plains and prairies and in cultivated grounds, y 
Missouri and Kansas to Georgia, Florida, Texas, 3 
New Mexico and adjacent Mexico. May—Aug. se 


25. BLEPHILIA Raf. Journ. Phys. 89:98. 18109. 


Perennial hirsute or pubescent erect herbs, with axillary and terminal dense glomerules 
of rather small purplish or bluish flowers, or the glomerules in terminal more or less inter- 
rupted spikes. Calyx tubular, 13-nerved, not villous in the throat, 2-lipped, the upper lip 
3-toothed, the lower 2-toothed, the teeth all aristate or those of the lower lip subulate. 
Corolla glabrous within, the tube expanded above, the limb 2-lipped; upper lip erect, entire; 
lower lip 3-lobed, the notched middle lobe narrower than the lateral ones. Anther-bearing 
(anterior) stamens 2, ascending, exserted or included; posterior stamens reduced to filiform 
staminodia, or none; anthers 2-celled, the sacs divaricate, somewhat confluent at the base. 
Ovary deeply 4-parted; style 2-cleft at the apex. Nutlets ovoid, smooth. [Greek, eyelash, 
from the fringed calyx-teeth.] 3 


Two species, natives of eastern North America. Type species: Blephilia ciliata (L.) Raf. 


Upper leaves lanceolate or oblong, sessile or short-petioled, slightly serrate. 1. B. ciliata. 
Leaves ovate or ovate-lanceolate, long-petioled, sharply serrate. 2. B. hirsuta. 


GENUS 25. MINT FAMILY. 135 


1. Blephilia ciliata (L.) Raf. Downy Ble- 
philia. Fig. 3645. 

Monarda ciliata L. Spe belies: 753. 

Blephilia ciliata Raf. Journ. Phys. 89: 98. 1819. 

Stem puberulent, or short-villous above, commonly 
simple, 1°-2° high. Lower leaves and those of sterile 
shoots ovate or oval, slender-petioled, crenate-den- 
ticulate, 1-2’ long, the upper lanceolate or oblong- 
lanceolate, short-petioled or sessile, mostly acute, 
longer and narrower, nearly entire; flower-clusters 
in a terminal spike and in the uppermost axils; 
outer bracts ovate to lanceolate, acuminate, usually 
purplish, pinnately veined, ciliate; calyx hirsute, the 
teeth of the upper lip about one-third longer than 
those of the lower; corolla purple, villous-pubescent, 
5-6” long; stamens exserted or included. 


In dry woods and thickets, Vermont to Minnesota, 
south to Georgia, Alabama and Missouri. June—Aug. 
Ohio horse-mint. 


hilia hirsuta (Pursh) Torr. Hairy Blephilia. Fig. 3646. 
Monarda hirsuta Pursh, Fl. Am. Sept. 19. 1814. 
Blephilia nepetoides Raf. Journ. Phys. 89: 98. 1819. 
Blephilia hirsuta Torr. Fl. U. S. 27. 1824. 


Stem villous-pubescent, or glabrous, usually branch- 
ed, 13°-3° high. Leaves membranous, ovate or ovate- 
lanceolate, acuminate at the apex, rounded, cordate 
or narrowed at the base, sharply serrate, slender- 
petioled, 2-4’ long, or the lower shorter and broader; 
flower-clusters axillary, or in a short terminal spike; 
outer bracts lanceolate or linear-lanceolate, long- 
acuminate, hirsute; calyx-tube nearly glabrous, its 
teeth very villous, those of the upper lip much ex- 
ceeding the lower; corolla pubescent, pale purple, 
rather conspicuously darker-spotted, 4’-5” long. 

In woods and thickets, Quebec and Vermont to Min- 


nesota, Kansas, Georgia and Texas. Ascends to 4000 ft. 
in North Carolina. June—Sept. 


26. HEDEOMA Pers. Syn. 23r 9 5607, 


Annual or perennial, strongly aromatic and pungent herbs, with small entire or crenulate 
leaves, and small blue or purple flowers in axillary clusters, these crowded into terminal, 
leafy-bracted spikes or racemes. Calyx tubular, 13-nerved, villous in the throat, the mouth 
mostly contracted in fruit, gibbous on the lower side at the base, or nearly terete, 2-lipped, 
or nearly equally 5-toothed, the upper lip 3-toothed, the lower 2-cleft. Corolla-limb 2-lipped, 
the upper lip erect, entire, emarginate or 2-lobed, the lower spreading, 3-cleft. Perfect stamens 
2, ascending under the upper lip, their anthers 2-celled, the sacs divergent or divaricate. 
Sterile stamens (staminodia) 2, minute, or none, very rarely anther-bearing. Ovary deeply 
4-parted; style 2-cleft at the summit, glabrous, Nutlets ovoid, smooth. [Greek, sweet smell.] 

About 15 species, natives of America. Besides the following, some 8 others occur in the south- 
ern and southwestern States. Sometimes called Mock Pennyroyal. Type species here taken as 
Hedeoma pulegioides (L.) Pers. 


Teeth of the upper lip of the calyx triangular; leaves serrate; annual. 1. H. pulegioides. 
Teeth of both lips of the calyx subulate; leaves entire. aki 
Calyx-teeth all nearly equal; annual. 2. H. hispida. 


Teeth of the lower lip nearly twice as long as the upper; perennial. 3. H.longifiora. 


136 LABIATAE, | Vor. III. 


1. Hedeoma pulegioides (L.) Pers. American Pennyroyal. Fig. 3647. 


Melissa pulegioides L. Sp. Pl. 593. 1753. 
Cunila pulegioides L. Sp. Pl. Ed. 2, 30. 1762. 
Hedeoma pulegioides Pers. Syn. 2: 131. 1807. 


Annual; stem very slender, erect, much branched, 
finely soft-pubescent, 6-18’ high, the branches ascend- 
ing. Leaves ovate to obovate-oblong, petioled, sparingly 
serrate, mostly obtuse at the apex and narrowed at the 
base, glabrous, or sparingly pubescent, thin, 3-12’ long, 
2-8" wide, the upper smaller; clusters few-flowered, 
axillary, rather loose; pedicels pubescent, shorter than 
or equalling the calyx; calyx pubescent, gibbous, oblong- 
ovoid in fruit, its 3 upper teeth triangular, acute, not 
exceeding the 2 subulate hispid lower ones; corolla 
bluish-purple, about 3” long; rudimentary stamens 
manifest, capitate at the summit, or rarely anther- 
bearing. 

In dry fields, Cape Breton Island to Ontario and Minne- 
sota, Florida, Alabama, Arkansas and Nebraska. Also 
called tick-weed and squaw-mint. July—Sept. 


2. Hedeoma hispida Pursh. Rough Pennyroyal. Fig. 3648. 


Hedeoma hispida Pursh, Fl. Am. Sept. 414. 1814. 
Hedeoma hirta Nutt. Gen. 1: 16. 1818. 


Annual; stem erect, branched, slender, 3-8’ high, 
pubescent, the branches erect-ascending, very leafy 
and copiously flowered. Leaves linear, entire, firm, 
sessile, or the lower short-petioled, blunt or subacute 
at the apex, narrowed at the base, more or less 
hispid-ciliate but otherwise mostly glabrous, 3’—1’ 
long, about 1” wide, the lower much shorter and 
smaller; clusters axillary, numerous, crowded, sev- 
eral-flowered; pedicels pubescent, shorter than the 
calyx; bracts subulate, very hispid, about equalling 
the calyx; calyx oblong, gibbous, hispid, its teeth 
all subulate, nearly equal in length, upwardly curved 
in fruit, about one-half as long as the tube, the 2 
lower ones somewhat narrower and more hispid 
than the upper; corolla about 3” long, bluish-purple; 
sterile stamens rudimentary or none. 


On dry plains, Ontario and New York to Saskatche- 
wan, Illinois, Louisiana, Arkansas and Colorado. May-'Aug. 


3. Hedeoma longiflora Rydb. Long-flowered 
Pennyroyal. Fig. 3649. 
Hedeoma longiflora Rydb. Bull. Torr. Club 36: 685. 1909. 


Perennial from a woody base, with an ashy down 
nearly all over; stems much branched, slender, erect, 
6-18’ high, the branches ascending. Leaves oblong or 
linear-oblong, entire, short-petioled or sessile, obtuse at 
the apex, narrowed at the base, spreading, 5-10” long, 
1’-23” wide, the lowest shorter; clusters axillary, 
loosely few-flowered; pedicels puberulent, about one- 
half as long as the calyx and equalling or longer than 
the subulate bracts; calyx oblong, hirsute, slightly 
gibbous, its teeth all subulate, upwardly curved and 
connivent in fruit, the 2 lower nearly twice as long 
as the 3 upper; corolla purple, 4-6” long; sterile 
stamens rudimentary or none. 

i th Dakota to Nebraska, Kansas and 
pen ee act i our first ‘edition, in H. Drummondit 
Benth. of the Southwest. April-Aug. 


GENUS 27. MINT FAMILY. 137 


27. MELISSA [Tourn.] L. Sp. Pl. 592. 1753. 


Leafy branching herbs, with broad dentate leaves, and rather small white or yellowish 
axillary clustered somewhat secund flowers. Calyx oblong-campanulate, deflexed in fruit, 
13-nerved, nearly naked in the throat, 2-lipped; upper lip flat, 3-toothed, the lower 2-parted. 
Corolla exserted, its tube curved-ascending, enlarged above, naked within, the limb 2-lipped; 
upper lip erect, emarginate; lower lip 3-cleft, spreading. Stamens 4, didynamous, connivent 
and ascending under the upper lip of the corolla; anthers 2-celled, their sacs divaricate. 
Ovary deeply 4-parted; style 2-cleft at the summit, the lobes subulate. Nutlets ovoid, smooth. 
[ Greek, bee.] 


About 4 species, natives of Europe and western 
Asia, the following typical. 


1. Melissa officinalis L. Garden or Lemon 
Balm. Bee-balm. Fig. 3650. 


Melissa officinalis L. Sp. Pl. 592. 1753. 


Perennial, pubescent; stem rather stout, erect 
or ascending, 1°-23° high. Leaves ovate, petioled, 
mostly obtuse at both ends, sometimes cordate, 
pinnately veined, coarsely dentate or crenate- 
dentate, 1-23’ long; flowers several in the axil- 
lary clusters; pedicels shorter than the calyx; 
calyx about 3” long, the teeth of its lower lip 
slightly exceeding those of the upper; corolla 
white, 5-7” long. 

In waste places, thickets and woods, Maine to 
Georgia, West Virginia, Missouri and Arkansas, 
also in Oregon and California. Naturalized from 
Europe. Plant lemon-scented. Balm-leaf or -mint. 


Honey-plant. Pimentary. Goose-tongue. Dropsy- 
plant. Lemon-lobelia. Sweet-mary. June—Aug. 


28. SATUREIA [Tourn.] L. Sp. Pl. 567. 1753. 


Herbs or shrubs, with small entire leaves, sometimes with smaller ones fascicled in their 
axils, and bracted purple flowers in dense terminal or axillary clusters. Calyx campanulate, 
mostly 1o-nerved, 5-toothed, naked or rarely villous in the throat. Corolla-limb 2-lipped, the 
upper lip erect, flat, entire or emarginate, the lower spreading, 3-cleft. Stamens 4, connivent 
under the upper lip of the corolla; anthers 2-celled, the sacs parallel or divaricate. Ovary 
deeply 4-parted; style 2-cleft at the summit. Nutlets oblong or oval. [The classical Latin 
name of the plant.] 


Ny 


About 18 species, the following typical one intro- 
duced as a garden herb from Europe, one of doubt- 
ful affinity in Florida, the others of the Mediterra- 


\\ Ni nean region. 

si 

aN \ a ee, 

SN 1. Satureia horténsis L. Savory. Sum- 


mer Savory. - Fig. 3651. 
Satureia hortensis L. Sp. Pl. 568. 1753. 


Annual, puberulent; stems erect, slender, much 
branched, 67-18’ high. Leaves linear or linear- 
oblong, short-petioled, entire, acute at both ends, 
3’-13’ long, 1-2” wide; clusters 3-5” in diam- 
eter, terminal and in many of the upper axils; 
bracts linear, small, minute, or wanting; calyx 
about equalling the corolla-tube, somewhat pu- 
bescent, its teeth subulate, about as long as the 
tube, ciliate; corolla little longer than the calyx; 
stamens scarcely exserted. 


7 


_In waste places, New Brunswick and Ontario to 
Kentucky, west to Nevada. Naturalized or ad- 
ventive from Europe. July—Sept. 


138 LABIATAE. Vor. III. 


29. CLINOPODIUM L. Sp. Pl. 587. 1753. 
[CALAMINTHA Moench, Meth. 408. 1794.] 

Herbs, or low shrubs, with entire or sparingly dentate leaves, and rather large flowers 
variously clustered. Calyx tubular or oblong, mostly gibbous at the base, about 13-nerved, 
2-lipped, naked or villous in the throat, the upper lip 3-toothed, the lower 2-cleft. Corolla 
usually expanded at the throat, the tube straight, mostly longer than the calyx, the limb 
2-lipped; upper lip erect, entire or emarginate; lower lip spreading, 3-cleft. Stamens 4, all 
anther-bearing, didynamous, ascending under the upper lip of the corolla, somewhat con- 
nivent in pairs, the longer mostly exserted; anthers 2-celled, the sacs divergent or divaricate. 
Ovary deeply 4-parted; style glabrous, 2-cleft at the summit. Nutlets ovoid, smooth. [Greek, 
bed-foot, the flowers likened to a bed-castor. | 

About 60 species, natives of the north temperate zone. Besides the following, 4 others occur 
in the southeastern United States and in California. The genus has been included in Satureia by 
authors. Type species: Clinopodium vulgare L. 

* Flower-clusters dense, axillary and terminal, setaceous-bracted. 1. C. vulgare. 
** Flower-clusters loose, axillary, or forming terminal thyrses; bracts small. 
Plants pubescent ; introduced species. 


Clusters peduncled ; calyx not gibbous; upper leaves very small; perennial. 2. C. Nepeta. 

Clusters sessile ; calyx very gibbous; plant leafy, annual. 3. C. Acinos. 
Plants glabrous; native species. 

Leaves linear or the lower spatulate, entire; corolla 4” long. 4. C. glabrum. 

Leaves oblong or oblong-lanceolate, serrate; corolla 6’—7” long. 5. C. glabellum. 


1. Clinopodium vulgare L. Field or Wild Basil. Basil-weed. Fig. 3652. 


Clinopodium vulgare L. Sp. Pl. 587. 1753. 
Melissa Clinopodium Benth. Lab. Gen. & Sp. 393. 1834. 
Calamintha Clinop. Benth. in DC. Prodr. 12: 233. 1848. 


Perennial by short creeping stolons, hirsute; stem 
slender, erect from an ascending base, usually branch- 
ed, sometimes simple, 1°-2° high. Leaves ovate or 
ovate-lanceolate, petioled, obtuse or acutish, entire, 
undulate or crenate-dentate, rounded, truncate or 
sometimes narrowed at the base, thin, 1—23’:long; 
flowers in dense axillary and terminal capitate- clus- 
ters about I’ in diameter; bracts setaceous, hirsute- 
ciliate, usually as long as the calyx-tube; calyx pu- 
bescent, somewhat gibbous, the setaceous teeth of 
its lower lip rather longer than the broader ones of 
the upper; corolla purple, pink, or white, little ex- 
ceeding the calyx-teeth. 

In woods and thickets, Newfoundland to North Caro- 
lina, Tennessee, Minnesota and Manitoba, in the Rocky 
Mountains to New Mexico and Arizona. Ascends to 
4ooo ft. in Virginia. Also in Europe and Asia. Stone- 
basil. Bed’s-foot. Field- or horse-thyme. Dog-mint. 
June—Oct. - 


Ko SS 
NAN AW 5 7 
Sty 7 NV 

Z, 


2. Clinopodium Népeta (L.) Kuntze. Field Balm. Field or Lesser Calamint. 
Basil-thyme. Fig. 3653. 


Melissa Nepeta L. Sp. Pl. 593. 1753. 

Cal. Nepeta Link & Hoffmansg. FI. Port. 1: 14. 1809. 
Clinopodium Nepeta Kuntze, Rev. Gen. Pl. 515. 1891. 
Satureia Nepeta Scheele, Flora 26: 577. 1843. 


Perennial by a woody root and short rootstocks, 
villous or pubescent; stem rather stout, at length 
much branched, the branches nearly straight, ascend- 
ing. Leaves broadly ovate, petioled, obtuse or acute, 
crenulate with few low teeth, rounded or narrowed 
at the base, the lower 3’-1’ long, the upper much 
smaller and bract-like; flowers few in the numerous 
loose peduncled axillary cymes, forming an almost 
naked elongated thyrsus; bracts very small, linear; 
calyx not gibbous, villous in the throat, about 13” 
long, the teeth of its lower lip twice as long as those 
of the upper; corolla light purple or almost white, 
about 4” long. 


In fields and waste places, Maryland to South Caro- 
lina, Alabama, Indiana, Kentucky and Arkansas. Ber- 
muda. Naturalized from Europe. Native also of Asia. 
June—Sept. 


GENUS 20. . MINT FAMILY. 139 


Clinopodium Calamintha (L.) Kuntze, the cala- /} 
mint of the Old World, with larger leaves and flow- YY} 
ers, admitted into our first edition, is not known in VN, 
the wild state within our area. WZ, 

Be 


a Ss 
3. Clinopodium Acinos (L.) Kuntze. yp 
Basil-thyme. Basil Balm. Fig. 3654.  S==" —— 


Thymus Acinos L. Sp. Pl. 591. 1753. 

Melissa Acinos Benth. Lab. Gen. & Sp. 389. 1834. 
Cal. Acinos Benth. in DC. Prodr. 12: 230. 1848. 
Clin. Acinos Kuntze, Rev. Gen. Pl. 513. 1891. 


Y 

Annual, pubescent; stems branched from the ‘ ava 
base, very slender, 6-8’ high. Leaves oblong ahh yee 
or ovate-oblong, petioled, acutish at both ends SX WED 
or the lower obtuse, crenulate or entire, 4-8” \ | or 
long; flowers about 6 in the axils, the clusters 
sessile; bracts shorter than the pedicels; calyx 
gibbous on the lower side, rough-hairy, longer \ 
than its pedicel, contracted at the throat, its subu- SV VW 
late teeth somewhat unequal in length; corolla GEL 
purplish, 13-2 times as long as the calyx. 

In waste places, Ontario to Massachusetts and 


New Jersey. Adventive or naturalized from Europe. oa 
Mother-of-thyme. Polly mountain. May-—Aug. 


VEG 


4. Clinopodium glabrum (Nutt.) Kuntze. 
Low Calamint or Bed’s-foot. Fig. 3655. 


Hedeoma glabra Nutt. Gen. 1: 16. 1818. 
Cal. Nuttallii Benth. in DC. Prodr. 12: 230. 1848. 
Calamintha glabella var. Nuttallii A. Gray, Man. 

Bidh27 3070 nose : 
Clin. glabrum Kuntze, Rev. Gen. Pl. 515. 1891. 

Perennial, glabrous, stoloniferous; stem very 
slender, at length much branched, erect or as- 
cending, 4-12’ high. Leaves of the flowering 
branches linear, entire, very short-petioled or 
sessile, mostly obtuse at the apex, obscurely 
veined, 4’-9” long, 3-1’ wide, the margins 
slightly revolute; lower leaves and those of the 
stolons shorter and broader, distinctly petioled ; 
flowers 1-4 in the axils; bracts minute; pedicels 
filiform, mostly longer than the calyx; calyx not 
gibbous, its throat pubescent in a ring within, 
its lower teeth somewhat longer than the upper; 
corolla purple, about 4” long. 

On rocks and banks, Ontario to western New 
York, Illinois, Minnesota, Missouri, Arkansas and 
Texas. May-Aug. 


5. Clinopodium glabéllum (Michx.) ae 
~ Kuntze. Slender Calamint or Bed’s- Gy Y, 
foot. Fig. 3656. e1 


Cunila glabella Michx. Fl. Bor. Am. 1: 13. 1803. fg” 
is ha glabella Benth. in DC, Prodr. 12: 230. _——A 


1848. 
Clin. glabellum Kuntze, Rev. Gen. Pl. 515. 1891. 


Perennial, stoloniferous, glabrous; stems 
weak, spreading or decumbent, at length freely 
branched, elongated, slender, 82° long. Leaves 
membranous, oblong, short-petioled, obtuse or 
the uppermost subacute at the apex, narrowed 
to a cuneate base, distinctly serrate with low 
teeth, 1’-2’ long, 2”-8” wide, the lowest and 
those of the stolons sometimes proportionately 
broader and shorter; axils 2-5-flowered; pedi- 
cels filiform, commenly twice as long as the 
calyx; bracts minute; calyx not gibbous, its 
throat pubescent in a ring within, its teeth 
nearly equal; corolla purplish, 6’—7” long. 

On river banks, Indiana to Kentucky, Tennes- 
see and Arkansas. May-July. 


140 LABIATAE. Vor. IIT. 


30. HYSSOPUS [Tourn.] L. Sp. Pl. 569. 1753. 


A perennial erect herb, the stem somewhat woody at the base, with narrow, entire leaves, 
and small bracted purple or blue flowers, in dense clusters in the upper axils, and forming 
elongated terminal more or less interrupted spikes. Calyx tubular, 15-nerved, about equally 
5-toothed, not hairy in the throat. Corolla-limb 2-lipped, the upper lip erect, emarginate, 
the lower spreading, 3-cleft, the middle lobe 2-lobed. Stamens 4, didynamous, the 2 longer 
ones exserted, divergent; anthers 2-celled, the sacs divaricate. Ovary deeply 4-parted; style 
2-cleft at the summit. Nutlets ovoid, somewhat 3-sided, nearly smooth. [Greek, an aromatic 
herb. | 


A monotypic genus of Europe and Asia. 


1. Hyssopus officinalis L. Hyssop. 
Fig. 3657. 
Hyssopus officinalis L. Sp. Pl. 569. 1753. 


Stems usually several together from the woody 
base, slender, strict, puberulent, simple or branch- 
ed, 1°-3° high, the branches upright or ascend- 
ing. Leaves linear to oblong, sessile or very 
nearly so, firm, acute at both ends or the lower 
obtuse at the apex, puberulent or glabrate, faintly 
veined, 13’-2’ long, 1”—3’’ wide, sometimes with 
smaller ones or short leafy branches in their 
axils; spike sometimes I-sided, dense, 3-1’ broad; 
pedicels short, puberulent; outer bracts as long 
as the calyx; calyx-teeth lanceolate, acute, one- 
fourth to one-third as long as the tube; corolla 
4’-5”’ long, its tube exceeding the calyx. 

Along roadsides and in waste places, Ontario and 


Maine to North Carolina, and on the Pacific Coast. 
Naturalized from Europe. June—Sept. 


31. ORIGANUM [Tourn.] L. Sp. Pl. 588. 1753. 


Perennial branching herbs, some species shrubby, with rather small crenate-dentate or 
entire leaves, and small bracted pink or purple flowers, in dense terminal glomerules. Calyx 
ovoid or campanulate, villous in the throat, about 13-nerved, 5-toothed or more or less 
2-lipped. Corolla-limb 2-lipped, the upper lip erect, emarginate or 2-lobed, the lower longer, 
spreading, 3-cleft. Stamens 4, didynamous, ascending; anthers 2-celled, the sacs divergent. 
Style 2-cleft at the summit; ovary deeply 4-parted. Nutlets ovoid or oblong, smooth. 
[Greek, mountain-joy.] 


About 30 species, natives of the Old World, the following typical. 


1. Origanum vulgare L. Wild Marjoram. 
Winter Sweet. Organy. Fig. 3658. 
Origanum vulgare L. Sp. Pl. 590. 1753. 


Perennial from nearly horizontal rootstocks, villous 
or hirsute; stem erect, slender, 1°-23° high. Leaves 
ovate, petioled, obtuse or subacute at the apex, rounded 
or subcordate at the base, crenate or entire, 1-13’ long, 
often with smaller ones, or short leafy branches, in 
their axils; flower-clusters often 2’ broad; bracts pur- 
plish, ovate or oval, about equalling the nearly regularly 
5-toothed calyx; corolla pink, purple or nearly white, 
longer than the calyx, the upper lobe broad; all four 
stamens, or the two longer, exserted. 

In fields and waste places, Ontario to New Jersey and 


Pennsylvania. Naturalized from Europe. Native also of 
Asia. Called also organs, pot-marjoram. July—Sept. 


GENUS 32. MINT FAMILY. 141 


32. THYMUS [Tourn.] L. Sp. Pl. 500. 1753. 


Perennial herbs, or low shrubby creeping plants, with small mostly entire leaves, and 
small purple flowers clustered in terminal glomerules, or in the axils of the leaves. Calyx 
ovoid, villous in the throat, 10-13-nerved, 2-lipped, the upper lip erect-spreading, 3-toothed, 
the lower 2-toothed, its teeth long and slender. Corolla-limb 2-lipped, the upper lip erect, 
emarginate, the lower spreading, 3-cleft. Stamens 4, more or less didynamous, mostly ex- 
serted; anthers 2-celled, the sacs parallel or divergent. Ovary deeply 4-parted; style 2-cleft 
at the summit. Nutlets ovoid or oblong, smooth. [Greek, incense. ] 


About 50 species, natives of the Old World, mostly European. Type species: Thymus vulgaris L. 


1. Thymus Serpyllum L. Wild or Creeping 
Thyme. Mother of Thyme. Fig. 3659. 
Thymus Serpyllum L. Sp. Pl. 590. 1753. 


" 

Stems more or less pubescent in lines, very slender, Nas 
procumbent, tough, much branched, 4-12’ long, com- is 
monly forming dense mats. Leaves oblong or ovate- } 
oblong, petioled, obtuse at the apex, usually narrowed 
at the base, entire, glabrous, or sometimes ciliate, 2-5” 
long; bracts similar to the leaves, but smaller; flowers 
numerous in verticillate clusters crowded in dense short 
terminal spikes, or also in the upper axils; calyx dis- 
tinctly 2-lipped, the tube usually pubescent and the teeth 
ciliate; corolla longer than the calyx. 


In thickets, woods, and along roadsides, Nova Scotia to 
southern New York and North Carolina. Naturalized from 
Europe. Native also of Asia. June-Sept. Old English 
names, brotherwort, hillwort, penny-mountain, shepherd’s- 
thyme. 


Blah KOELLIA Moench, Meth. 407. 1794. 
[BRACHYSTEMON and PycNANTHEMUM Michx. Fl. Bor. Am. 2: 5, 7. 1803.] 


Perennial erect, mostly branched, glabrous cinereous canescent or pubescent herbs, with 
small white or purple-dotted flowers, in terminal or sometimes also axillary capitate glom- 
erules or cymose clusters. Calyx ovoid, oblong or tubular, 1o-13-nerved, equally or more 
or less unequally 5-toothed, not villous in the throat, the 2 upper teeth sometimes united 
below. Corolla 2-lipped, the upper lip emarginate or entire, the lower 3-cleft, its lobes 
obtuse. Stamens 4, didynamous, nearly equal, or the lower pair a little longer; anther-sacs 
parallel. Ovary deeply 4-parted; style slender. Nutlets smooth, pubescent, or roughened. 
[Named for. J. L. C. Koelle, a German botanist of the eighteenth century. ] 

About 17 species, natives of North'America. Besides the following, 1 occurs in California and 


several in the Southern States. Type species: Koellia capitata Moench. Mostly very fragrant. 
Sometimes called Basil, the proper name of Calamint or Ocymum. 


* Leaves prevailingly linear, linear-lanceolate or oblong-lanceolate. 


Calyx-teeth ovate-triangular, acute, one-fourth as long as the tube. 1. K. virginiana. 
Calyx-teeth subulate, lance-subulate or bristle-tipped. 
Leaves linear or lanceolate; calyx-teeth subulate or lanceolate. 
Leaves linear or narrowly linear-lanceolate, entire. 2. K. flexuosa. 
Leaves lanceolate, entire, or usually serrate. 
Bracts appressed, erect; clusters dense. 
Hirsute or pilose; leaves mainly entire. K. pilosa. 
Puberulent, short-pubescent, or glabrate; leaves mostly denisoilate: 
= K. verticillata. 
Bracts spreading; clusters loose. K. clinopodioides. 
Leaves oblong to linear- oblong, obtuse or subacute; calyx-teeth awn-like. 6 K. hyssopifolia. 


** Leaves prevailingly ovate, ovate-oblong or ovate-lanceolate. 
Calyx-teeth bristle-tipped or subulate. 


Bracts appressed; clusters dense; calyx-teeth bristle-pointed. 7. K. aristata. 
Bracts spreading; clusters loose; calyx-teeth subulate. 
Calyx-teeth about one-half as long as the tube. 8. K. incana, 
Calyx-teeth as long as the tube. 9. K. pycnanthemoides. 
Calyx-teeth triangular, triangular-lanceolate or lanceolate. 
Bracts spreading; clusters loose; calyx-teeth short. 10. K. albescens. 
Bracts appressed; clusters dense. : i 
Bracts canescent ; leaves firm, acute. 11. K. mutica. 
Bracts ciliate or villous ; leaves membranous, acuminate. 12. K. montana. 


142 LABIATAE. Voi. III. 


1. Koellia virginiana (L.) MacM. Virginia Mountain-Mint. Fig. 3660. 


Satureja virginiana L. Sp. Pl. 567. 1753. 
P. lanceolatum Pursh, Fl. Am. Sept. 409. 1814. 
Koellia virginiana MacM. Met. Minn. 452. 1892. 


Stem strict, rather stout, glabrous or pubescent, 
1°-3° high. Leaves lanceolate or linear-lanceo- 
late, fragrant, very short-petioled or sessile, firm, 
entire, acuminate at the apex, glabrous or some- 
what puberulent beneath, or the uppermost densely 
canescent, 1-2’ long, 2-5” wide, often with short 
leafy branches in their axils; glomerules dense, 
4-6” in diameter, terminal, corymbed, canescent; 
bracts appressed, rigid, acute, acuminate or subu- 
late-tipped, not exceeding the clusters; calyx 
cylindraceous, or expanded above, canescent, its 
teeth triangular-ovate, equal or nearly so, acute, 
little longer than wide, about one-fourth as long 
as the tube; corolla pubescent without, purple- 
spotted, its tube longer than the calyx. 


In dry fields and thickets, Quebec and Ontario to 
Minnesota, south to Georgia, Alabama and Kansas. 
Virginia or mountain thyme. Prairie-hyssop. Penny- 
royal. Basil. July—Sept. 


Satureja Thymus virginicus L. Mant. 2: 409: 1771. Not 
S. virginica L. 1753. 

Origanum flexuosum Walt. Fl. Car. 165. 1788. 

Koellia capitata Moench, Meth. 408. 1794. 

Pycnanthemum linifolium Pursh, Fl. Am. Sept. 409. 1814. 

P. flexuosum B.S.P. Prel. Cat. U. S. 42. 1888. 

Koellia flexuosa MacM. Met. Minn: 452. 1892. 


Stem slender, stiff, nearly glabrous throughout, 13°-23° 
high. Leaves linear or linear-lanceolate, entire, glabrous 
or the uppermost puberulent, sessile, or the lower very 
short-petioled, 1-2’ long, 2’—13” wide, rather firm, often 
with short leafy branches in their axils; glomerules dense, 
terminal, corymbed, 3-5” broad, canescent; bracts ap- 
pressed, acuminate, or subulate-tipped, not longer than 
the clusters; calyx cylindraceous, canescent, its teeth subu- 
late and rigid, equal or nearly so, 3-4 times longer than 
broad, about one-third as long as the tube; corolla-tube 
longer than the calyx. 

In fields and thickets, Maine to Florida, New York, Minne- 
sota, Kansas and Texas. Little fragrant. July—Sept. 


3. Koellia pilésa (Nutt.) Britton. Hairy 


RUSS : Lt Se, Mountain-Mint. Fig. 3662. 
Vi VAT a QZ tag © Pycnanthemum pilosum Nutt. Gen. 2: 33. 1818. 
Pine : = QV, Pycnanthemum muticum var. pilosum A. Gray, Syn. FI. 
YESAS 2: Part 1, 355. 1878. 
71 Koellia pilosa Britton, Mem. Torr. Club 5: 279. 1894. 
> 


Pubescent, at least above, 1°-23° high. Leaves 
lanceolate, very short-petioled or sessile, entire or 
very sparingly denticulate, 1-2’ long, 3’-6” wide, 
firm, acuminate at the apex, mostly narrowed at the 
base, commonly with smaller ones, or short leafy 
shoots in their axils; glomerules dense, numerous, 
términal, villous or hirsute-canescent, about 4” in 
diameter; bracts lanceolate, acuminate, equalling or 
exceeding the clusters; calyx cylindraceous, narrow, 
canescent, its teeth lanceolate-subulate, equal, often 
ciliate, about 3 times as long as wide and one-fourth 
the length of the tube; corolla pubescent, its tube 
little longer than the calyx; stamens exserted. 


On prairies and in dry woods, Ontario to Pennsyl- 
vania, Georgia, Iowa, Kansas and Arkansas. July—Sept. 


3 


Koellia léptodon (A. Gray) Small, of the North Carolina mountains, with bristly-ciliate calyx- 
teeth, is: ecorded as extending to Ohio and Missouri. 


GENUS 33. MINT FAMILY. 143 


4. Koellia verticillata (Michx.) Kuntze. Torrey’s Mountain-Mint. Fig. 3663. 


Brachystemon verticillatum Michx. Fl. Bor. Am. 2: 
Oupl. si. 1803: 


P. Torreyi Benth. Lab. Gen. & Sp. 329. 1834. EVAR 99, CONN TTY 
Koellia verticillata Kuntze, Rev. Gen. Pl. 520. 1891. < eS aap @ 
ri] ji ——- a P 
Puberulent, glabrate or pubescent; stem slender, | Z 


1°-22° high. Leaves lanceolate, oblong-lanceolate 
or linear-lanceolate- (rarely ovate-lanceolate), 
short-petioled or sessile, serrulate or entire, acute 
or acuminate at the apex, rounded or narrowed 
at the base, 1-3’ long, 3-10’ wide, the upper- 
most sometimes canescent; flower-clusters dense, 
canescent, 5’-6” broad, terminal, corymbose and 
commonly also in some of the upper axils; bracts 
appressed, lanceolate, acuminate, ciliate, equalling 
or longer than the clusters; calyx canescent, its 
teeth subulate or lance-subulate, ciliate, 2-3 times 
as long as wide, one-fourth to one-third as long 
as the tube; corolla pubescent, its tube rather ex- 
ceeding the calyx. 


In dry fields and thickets, Vermont to Virginia, 
west to Missouri. July—Sept. 


5. Koellia clinopodioides (T. & G.) Kuntze. 
Basil Mountain-Mint. Fig. 3664. 


Pycnanthemum clinopodioides T. & G.; A. Gray, Am. 
Journ. Sci. 42: 45. 1842. 
Koellia clinopodioides Kuntze, Rev. Gen. Pl. 520. 1891. 
Pubescent or puberulent; stem slender, 1°-23° high. 
Leaves lanceolate or oblong-lanceolate, rather thin, 
short-petioled, sharply serrate, or the upper entire, 
12’-3' long, 5-12” wide, none of them canescent; 
flower-clusters loose, terminal and axillary, about 1’ 
broad; bracts linear-acuminate or subulate-tipped, 
not exceeding the clusters, some or all of them 
spreading; calyx finely canescent or glabrate, its 
teeth subulate, sometimes with a few long hairs, 
slightly unequal, about one-third the length of the 
tube; corolla-tube longer than the calyx. 


In dry soil, Connecticut to Pennsylvania, Virginia and 
Tennessee. Ascends to 5000 ft. in Virginia. Aug.—Sept. 


(Benth.) Britton. Hyssop Mountain-Mint. Fig. 366s. 


P. hyssopifolium Benth. Lab. Gen. & Sp. 329. 1834. 
Pycnanthemum aristatum var. hyssopifolium A. Gray, 


Syilol, 2) Part t, 354. 1878. 
K. hyssopifolia Britton, Mem. ‘lorr. Club 5: 279. 1894. 


Puberulent or glabrate; stem slender, stiff, 1°-3° 
high. Leaves oblong, linear-oblong, or lanceolate- 
oblong, short-petioled, or the upper sessile, obtuse 
or subacute at the apex, narrowed at the base, 
entire or denticulate, 3’-14’ long, 2”-6” wide, 
glabrous or minutely canescent; flower-clusters 
dense, minutely canescent, not at all villous, ter- 
minal, and usually also in the upper axils, often 
1’ broad; bracts linear-oblong, narrowed at each 
end, terminated by an awn almost as long as the 
body; calyx cylindraceous, glabrous or very nearly 
so, prominently nerved, its teeth bristle-pointed, 
slightly widened below, nearly as long as the 2 
tube; corolla-tube not longer than calyx. 1 


In dry soil, Virginia to Florida. June—Aug. 


144 LABIATAE. Vor. IIL. 


7. Koellia aristata (Michx.) Kuntze. Awned 
Mountain-Mint. Fig. 3666. 


Pyc. aristatum Michx. Fl. Bor. Am. 2: 8. pl. 33. 1803. 
Koellia aristata Kuntze, Rev. Gen. Pl. 520. 1891. 

Similar to the preceding species; stem slender, 
stiff, minutely canescent, 13°-23° high. Leaves ovate, 
or some of them ovate-lanceolate, short-petioled, 
sharply serrate, serrulate, or the upper entire, acute 
at the apex, rounded at the base, 1’-2’ long, 4’-12” 
wide, the uppermost usually minutely canescent; 
inflorescence as in the preceding species; bracts long- 
awned, appressed, the awn about one-third the length 
of the body; calyx canescent, its teeth equal, bristle- 
pointed, widened below, one-third to one-half as 
long as the tube; corolla-tube about equalling the 
calyx. 

In dry pine barrens, New Jersey to Florida and Louisi- 
ana, mostly near the coast. Wild basil. July—Sept. 


8. Koellia incana (L.) Kuntze. Hoary Mountain-Mint. Fig. 3667. 


Clinopodium incanum L. Sp. Pl. 588. 1753. 7 ; 
Pycnanthemum incanum Michx. Fl. Bor. Am.2: 7. 1803. WY Ne Maso RPO 
Koellia incana Kuntze, Rev. Gen. Pl. 520. 1891. . i 


Stem pubescent, or glabrous below, stout, 13°-3° 
high. Leaves thin, ovate to ovate-lanceolate, peti- 
oled, acute at the apex, sharply serrate or serrulate, 
white-canescent beneath, puberulent or glabrous 
above, 14-3’ long, #13’ wide, or the uppermost 
smaller and sometimes canescent on both sides; 
clusters loose, terminal and in the upper axils, 1’-13’ 
broad, canescent, the flowers sometimes secund on 
their branches; bracts linear, or the outer broader, 
canescent or slightly villous, spreading, mostly 
shorter than the clusters; calyx canescent, slightly 
2-lipped, its teeth subulate, somewhat unequal, the 
longer one-fourth to one-half as long as the tube, 
rarely villous; corolla-tube equalling or longer than 
the calyx. 


Dry thickets and hillsides, Maine to Ontario, Florida, 
Alabama and Missouri. Calamint. Wild basil. Aug.—Oct. 


9. Koellia pycnanthemoides (Leavenw.) Kuntze. Southern Mountain-Mint. 
Fig. 3668. 


Tullia pycnanthemoides Leavenw. Am. Journ. Sci. 20: 
343. pl. 5. 1830. 

P. Tullia Benth. Lab. Gen. & Sp. 328. 1834. 

K. pycnanthemoides Kuntze, Rev. Gen. Pl. 520. 1891. 

P. pycnanthemoides Fernald, Rhodora 10: 86. 1908. 


Stem rather stout, pubescent nearly to the base, 
2°-3° high. Leaves membranous, petioled, mostly 
ovate-lanceolate, acuminate or acute at the apex, 
narrowed at the base, sharply serrate, pubescent 
beneath, puberulent or glabrate and dark green 
above, 12’—33’ long, 2’-14’ wide, the lower green, the 
upper smaller and white-canescent on both sides; 
clusters loose, villous and canescent, terminal and 
axillary, 1-2’ broad, the flowers often secund; 
bracts linear-oblong, acuminate or subulate-tipped, 
villous, spreading; calyx-teeth very unequal, subu- 
late, villous, the longer about equalling the tube; 
corolla-tube not exceeding the calyx. 


In dry woods and on hills, Virginia to Georgia, Ken- 
tucky and Tennessee. Calamint. July—Sept. 


GENUS 33. MINT FAMILY. 145 


10. Koellia albéscens (T. & G.) Kuntze. 
White-leaved Mountain-Mint. Fig. 3669. 


P, albescens T. & G.; A. Gray, Am. Journ. Sci. 42: 


PAS tS42. 
Koellia albescens Kuntze, Rev. Gen. Pl. 520. 1891. 


Stem slender, soft-pubescent nearly to the base. 
1°-2° high. Leaves ovate to ovate-lanceolate, peti- 
oled, acute or subacuminate at the apex, narrowed 
or sometimes rounded at the base, sharply serrate 
or nearly entire, 1-23’ long, 4’-13’ wide, white- 
canescent beneath, green above, or the upper canes- 
cent on both sides; clusters loose, terminal and 
axillary, densely canescent, not at all villous, at 
length about 1’ broad; bracts linear, or the outer 
broader, spreading, sometimes exceeding the clus- 
ters; calyx densely canescent, its teeth triangular, 
obtuse or acute, slightly unequal, one-fifth to one- 
fourth as long as the tube; corolla-tube longer 
than the calyx. 

In dry woods and thickets, southern Virginia to 


Kentucky, Missouri, Arkansas, Florida and Texas. 
July—Sept. 


11. Koellia mutica (Michx.) Britton. Short- 
toothed Mountain-Mint. Fig. 3670. 


Brachystemon muticum Michx. Fl. Bor. ‘Am. 2: 6. pl. 32. 


1803. 
Koellia mutica Britton, Mem. Torr. Club 4: 145. 1804. 


Stem stiff, puberulent, or glabrous below, 1°-23° 
high. Leaves short-petioled or sessile, strongly veined, 
ovate or ovate-lanceolate, firm, acute at the apex, 
rounded or sometimes subcordate at the base, sharply 
serrate or serrulate, 12-3’ long, 3/-13’ wide, the lower 
mostly glabrous, the uppermost white-canescent on both 
sides, much smaller; flower-clusters capitate, dense, ter- 
minal, corymbose, often also in the upper axils, pubes- 
cent or canescent, 4-6” broad; bracts appressed, lan- 
ceolate-subulate or the outer broader; calyx pubescent, 
its teeth nearly equal, triangular-ovate or triangular- 
lanceolate, not much longer than wide, about one-fifth 
as long as the tube; corolla-tube exceeding the calyx. 

In sandy soil, Maine to Virginia and Florida, west to 
Pennsylvania and Missouri. Calamint. July—Sept. 


12. Koellia montana (Michx.) Kuntze. Thin- 
leaved Mountain-Mint. Fig. 3671. 
Pycnanthemum montanum Michx. Fl. Bor. Am. 2: 8. 1803. 


Monardella montana Benth. Lab. Gen. & Sp. 331. 1834. 
Koellia montana Kuntze, Rev. Gen. Pl. 520. 1891. 


sy, 


ie 


Stem slender, glabrous or nearly so throughout, 2°-3° 
high. Leaves distinctly petioled, membranous, glabrous, | 
ovate-lanceolate or the upper lanceolate, long-acuminate 
at the apex, usually narrowed at the base, sharply serrate, 
2’-5’ long, 3’-2’ wide, none of them canescent; flower- 
clusters rather dense, terminal and in the upper axils, 4’-1’ 
broad; bracts appressed, lanceolate or linear-lanceolate, 
acuminate, not exceeding the fully developed clusters, 
bearded more or less with long hairs; calyx tubular, 
glabrous or puberulent, or villous above, its teeth triangular- 
subulate, equal, one-fifth to one-fourth as long as the 
tube; corolla-tube longer than the calyx. 


In woods, mountains of southern Virginia to Georgia, Ten- 
nessee and Alabama. July—Sept. 


34. CUNILA L. Syst. Ed. 10, 1359. 1759. 


Perennial branching herbs, or low shrubs, with dentate or entire leaves, and rather 
small, clustered, purple or white flowers. Calyx tubular-ovoid, 10-13-nerved, villous in the 
throat, equally 5-toothed. Coro]la 2-lipped, longer than the calyx, the upper lip erect, emar- 
ginate, the lower spreading, 3-cleft. Anther-bearing stamens 2, long-exserted, straight, the 


Io 


146 LABIATAE. Vor. III. 


posterior pair rudimentary, or wanting; anther-sacs parallel. Ovary deeply 4-parted; style 
slender, 2-cleft at the summit. Nutlets smooth; scar of attachment basal and small. [Latin 
name of some plant.] t 

About 15 species, natives of America. The following typical speciés is the only one known 
ia the United States. 

1. Cunila origanoides (L.) Britton. Stone Mint. 
Sweet Horse-Mint. American Dittany. 
Wild Basil. Fig. 3672. 

Satureia origanoides L. Sp. Pl. 568. 1753. 
Cunila Mariana L. Syst. Ed. 10, 1359. 1759. 


Hedyosmos origanoides Kuntze, Rev. Gen. Pl. 520. 1891. 
Cunila origanoides Britton, Mem. Torr. Club 5: 278. 1894. 


Plant very aromatic; stem slender, stiff, branched, 
glabrous, or pubescent at the nodes, erect, 8-20’ high, 
the branches ascending. Leaves ovate, sessile or very 
short-petioled, acute at the apex, sharply serrate, round- 
ed, truncate or subcordate at the base, 4’-13’ long, 
densely punctate; flowers nearly 3’ long, numerous in 
terminal loose cymose clusters; corolla purple-pink, one- 
half as long as the stamens; posterior pair of stamens 
usually rudimentary. 


In dry woods and thickets, southern New York to Florida, 
west to Ohio, Missouri, Arkansas and Texas. Aug.—Sept. 


35. LYCOPUS [Tourn.] L. Sp. Pl. 21. 1753. 


Herbs, perennial by slender stolons or suckers, with erect or diffuse stems, petioled or 
sessile leaves, and small white or purple flowers, bracted and verticillate in dense axillary 
clusters. Calyx campanulate, regular or nearly so, 4-5-toothed, not bearded in the throat, 
the teeth obtuse or acute. Corolla funnelform-campanulate to cylindric, equalling or longer 
than the calyx, the limb nearly equally 4-cleft, or one of the lobes broader and emarginate. 
Perfect stamens 2, anterior, the posterior pair rudimentary, or altogether wanting; anther- 
sacs parallel. Ovary deeply 4-parted; style slender, 2-cleft at the summit. Nutlets truncate 
at the summit, narrowed below, trigonous, smooth, their margins thickened. [Greek, wolf-foot.] 

About 15 species of the north temperate zone. Besides the following, two or three others occur 
in western North America. Type species: Lycopus europaeus L. 

Calyx-teeth 4 or 5, ovate, shorter than the nutlets. a 
Base of the stem not tuberous; leaves ovate or ovate-lanceolate. 1. L. virginicus. 
Base of the stem tuberous-thickened ; leaves oblong to lanceolate. 2. L. uniflorus. 

Calyx-teeth mostly 5, lanceolate or subulate, longer than the nutlets. 
Bracts minute; corolla twice as long as the calyx. 

Leaves sessile. 
Leaves narrowed into a manifest petiole. 

Bracts lanceolate or:subulate; corolla not twice as long as the calyx. 
Leaves pinnatifid or deeply incised. bint 
Leaves merely coarsely dentate or serrate (lower rarely incised). 

Leaves oblong or oblong-lanceolate, serrate. 
Leaves ovate, coarsely dentate. 


. L. sessilifolius. 
. L. rubellus. 


. L. americanys. 


. L. asper. 
. L. europaeus. 


3 
4 
5 
6 
7 


1. Lycopus virginicus L. Bugle-weed. Bugle- 
wort. Fig. 3673. 
Lycopus virginicus L. Sp. Pl. 21. 1753. 


Perennial by long filiform leafy stolons, glabrous or 
puberulent; stem slender, erect or ascending, simple or 
branched, 6’-2° high. Leaves ovate or ovate-lanceolate, 
acuminate at the apex, sharply dentate, narrowed or 
cuneate at the base, petioled, or the upper sessile, dark 
green or purple, 13’-3’ long, 3’-13’ wide; bracts short, 
oblong; calyx-teeth 4, or sometimes 5, ovate or ovate- 
lanceolate, obtuse or subacute; corolla about 1” broad, 
narrow, nearly twice as long as the calyx, or longer; 
rudimentary posterior stamens minute; nutlets longer 
than or about equalling the calyx. 

In wet soil, New Hampshire to Florida, Alabama, Mis- 


souri and Nebraska. Northern Asia. Sometimes called 
wood betony. July—Sept. 


GENUS 35. MINT FAMILY. 147 


me Lycopus uniflorus Michx. Northern 
Bugle-weed. Fig. 3674. 


Lycopus uniflorus Michx. Fl. Bor. Am. 1: 14. 1803. 
Lycopus communis Bicknell, Britton Man. 803. 1901. 


Green or sometimes purplish, mostly less pu- 
berulent than L. virginicus; stems rather slender, 
simple or sparingly branched, 4’—23° tall, rather 
acutely angled, tuberous at the base, the stolons 
rarely tuber-bearing. Leaves oblong or oblong- 
lanceolate, acute or acuminate at the apex, ser- 
rate, #/-32’ long, sessile or nearly so; calyx-teeth 
4 or 5, triangular, ovate or ovate-oblong, rather 
obtuse; corolla about 13” long, less than twice as 
long as the calyx; rudimentary posterior stamens We i 


VO 


obsolete or minute; nutlets about as long as the \ 
calyx, oblique at the apex. 


= 


In low grounds, Newfoundland to British Colum- 
bia, North Carolina, Nebraska and Oregon. Sum- 


mer and fall. 

Lycopus membranaceus Bicknell, with thinner, (~S 
often coarsely-toothed, longer-petioled and larger 
leaves, appears to be a race of this species. 


3- Lycopus sessilifolius A. Gray. Sessile- 
leaved Water Hoarhound. Fig. 3675. 


Lycopus europaeus var. sessilifolius A. Gray, Man. Ed. 


5, 345. 1867. 
Lycopus sessilifolius A. Gray, Proc. Am. Acad. 8: 285. 
1870. 


Glabrous, or puberulent above, tuberiferous, per- 
ennial by stolons and suckers; stem simple, or at 
length branched, erect, 1°-2° high. Leaves ovate- 
lanceolate or oblong-lanceolate, closely sessile, acute 
or acuminate at the apex, sharply serrate, some- 
what narrowed at the base, 1-2’ long; bracts very 
‘small, acute; calyx-teeth usually 5, subulate, rigid, 
nearly as long as the tube; corolla twice as long as 
the calyx; rudimentary posteriorstamens oval; nut- 
lets shorter than the calyx. 


In wet soil, eastern Massachusetts to Florida and 
Mississippi, near the coast. Aug.—Oct. 


4. Lycopus rubéllus Moench. Stalked \ 
Water Hoarhound. Fig. 3676. 


Lycopus rubellus Moench, Meth. Suppl. 146. 1802. 
Lycopus europaeus var. integrifolius A. Gray, Man, 
Ed. 5, 346. 1867. 


Glabrous or minutely puberulent, perennial by 
leafy stolons; stem erect or ascending, simple or 
at length freely branched, 1°-3° high. Leaves 
ovate to oblong-lanceolate, or narrower, acumi- 
nate at the apex, usually dentate, narrowed or 
cuneate at the base, 2-5’ long, 4’-14’ wide, usually 
tapering into a conspicuous petiole; bracts mi- 
nute, acute or acuminate; calyx-teeth triangular- 
subulate, herbaceous, one-half as long as the 
tube or more; corolla longer than the calyx; 
rudimentary posterior stamens oval or oblong; 
nutlets much shorter than the calyx. 


In wet soil, southern Vermont to Florida, Minne- 
sota, Arkansas and Louisiana. Gipsywort. July- 
Oct. 


148 LABIATAE. Vor. III. 


5. Lycopus americanus Muhl. Cut-leaved 
Water Hoarhound. Fig. 3677. 


L. americanus Muhl.; Bart. Fl. Phil. Prodr.15. 1815. 
Lycopus sinuatus Ell. Bot. S. C. & Ga. 1: 26. 1817. 


Lycopaeus europaeus var. sinuatus A. Gray, Man. 
Ed. 5, 346. 1867. 


Puberulent or glabrous, perennial by suckers; 
stem stiff, erect, simple or branched, 1°-2° high. 
Leaves lanceolate or ovate-lanceolate in outline, 
petioled, acuminate at the apex, incised, pinna- 
tifid or the uppermost merely serrate, 2-4’ long; 
bracts subulate, the outer ones sometimes exceed- 
ing the calyx; calyx-teeth triangular-subulate, 
cuspidate, rigid; corolla little exceeding the ca- 
lyx; rudimentary posterior stamens thickened at 
their tips; nutlets much shorter than the calyx. 

In wet soil, Newfoundland to British Columbia, 


south to Florida, Texas, Utah and California. Gipsy- 
wort. Bitter bugle. Paul’s-betony. June—Oct. 


6. Lycopus asper Greene. Western Water Hoarhound. Fig. 3678. 
Lycopus asper Greene, Pittonia 3: 339. 18098. 
Lycopus lucidus var. americanus A. Gray, Proc. Am. 

Acad. 8: 286. 1870. 

Pubescent or glabrate, perennial by stolons; stem 
usually stout, erect, strict, leafy, simple, or some- 
times branched, 1°-3° high. Leaves oblong-lanceo- 
late, acute at the apex, narrowed or rounded at the 
base, sessile, or very short-petioled, 2’-6’ long, 2’-12 
wide, sharply serrate with acute ascending teeth; 
bracts ovate or lanceolate, acuminate-subulate, the 
outer ones often as long as the flowers; calyx-teeth 
3, subulate-lanceolate, nearly as long as the tube; 
corolla little longer than the calyx; rudimentary 
stamens slender, thickened at the tips; nutlets much 
shorter than the calyx. 

In wet soil, Michigan to Kansas, west to Manitoba, 
British Columbia, California and Arizona. Regarded in 
the first edition of this work as the same as L. lucidus 
Turcz. of NW. America and NE. ‘Asia. July—Sept. 


3 


7. Lycopus europaéus L. Water or Marsh 
Hoarhound. Gipsy-wort. Gipsy-herb. 
Fig. 3679. 

Lycopus europaeus L. Sp. Pl. 21. 1753. 


Puberulent or pubescent, perennial by suckers; 
stems stout, at length widely branched, 1°-23° 
high. Leaves ovate, ovate-oblong, or oblong- 
lanceolate, short-petioled, or the upper sometimes 
sessile, coarsely dentate, or the lower incised at 
the base, 1-3’ long, 2’-1’ wide; bracts subulate- 
lanceolate, the outer shorter than or equalling 
the flowers; calyx-teeth subulate-spinulose; co- 
rolla scarcely longer than the calyx; rudimentary 
posterior stamens obsolete; nutlets shorter than 
the calyx. 

In waste places, Massachusetts to Virginia. Natu- 
ralized from Europe. Greenarchangel. Bitter bugle- 
weed. July—Sept. 


36. MENTHA [Tourn.] L. Sp. Pl. 576. 1753. 


Erect or diffuse odorous herbs, with simple sessile or petioled mostly punctate leaves, 
and small whorled purple pink or white flowers, the whorls axillary or in terminal dense or 
interrupted spikes. Calyx campanulate to tubular, 1o-nerved, regular, or slightly 2-lipped, 


GENUS 36. MINT FAMILY. 149 


5-toothed. Corolla-tube shorter than the calyx, the limb 4-cleft, somewhat irregular, the 
posterior lobe usually somewhat broader than the others, entire or emarginate. Stamens 4, 
equal, erect, included or exserted, sometimes imperfect; filaments glabrous; anthers 2-celled, 
the sacs parallel. Ovary 4-parted; style 2-cleft at the summit. Nutlets ovoid, smooth. 
[Name used by Theophrastus; from the nymph Minthe.] 

About 30 species, natives of the north temperate zone. The more or less characteristic odors 
of the species change during the progress of the life of the plant. Type species: Mentha spicata L. 

* Whorls of flowers in terminal spikes, or some in the upper axils. 

Plants glabrous or very nearly so. 


Spikes slim, narrow, mostly interrupted; leaves sessile, or nearly so. 1. M. spicata. 
Spikes thick, mostly dense, at first short ; leaves petioled. 
Leaves lanceolate or oblong, acute. 2. M. piperita. 
Leaves ovate, obtuse, or the upper acute, subcordate. 3. M. citrata. 


Plants villous, hirsute or canescent, at least at the nodes. 
Spikes slim or narrow,.often interrupted. 
Leaves lanceolate or ovate-lanceolate, acute. 4. M. longifolia. 
Leaves elliptic or ovate-oblong, obtuse, reticulated beneath. 5. M. rotundifolia. 
Spikes thick (6”), dense, elongated or short. 
Leaves sessile; spikes 1’—3’ long; plant canescent. 
Leaves distinctly petioled, or the uppermost sessile; spikes short. 
Leaves simply serrate. 7. M. aquatica. 
Leaves mostly incised, the margins crisped and wavy. . M. cripsa. 


** Whorls of flowers all axillary. 


Upper leaves much smaller than the lower. 9. M. Cardiaca. 
Upper leaves not conspicuously reduced. 
Stem pubescent. 


n 


. M. alopecuroides. 


ice) 


Leaves rounded or obtuse at the base. 10. M. arvensis. | 
Leaves narrowed, mostly cuneate at the base. 11. M. canadensis. 
Stem glabrous or nearly so. 12. M. gentilis. 


1. Mentha spicata L. Spearmint. Lamb or 
Common Mint. OurLady’s Mint. Fig. 3680. 


Mentha spicata L. Sp. Pl. 576. 1753. 
Mentha spicata var. viridis L. loc. cit. 1753. 
Mentha viridis L. Sp. Pl. Ed. 2, 804. 1763. 


Glabrous, perennial by leafy stolons; stem erect? 
branched, 1°-13° high. Leaves lanceolate, sessile or 
short-petioled, sharply serrate, acute or acuminate 
at the apex, narrowed at the base, the largest about 
23’ long; whorls of flowers in terminal narrow acute 
usually interrupted spikes, which become 2’-4’ long 
in fruit, the one terminating the stem surpassing the 
lateral ones; bracts subulate-lanceolate, ciliate, some 
of them usually longer than the flowers; calyx cam- 
panulate, its teeth hirsute or glabrate, subulate, nearly 
as long as the tube; corolla glabrous. 


In moist fields or waste places, Nova Scotia to On- 
tario, Minnesota, Washington, Florida, Texas and Cali- 
fornia. Naturalized from Europe. Also in Bermuda. 
Native also of Asia. Garden-, brown- or mackerel-mint. 
Sage-of-bethlehem. July—Sept. 


Hs asi 2. Mentha piperita L. Peppermint. Fig. 3681. 
WS (7 Wie 
wit MG WG Mentha piperita L. Sp. Pl. 576. 1753. 
adi O47) UNE? ; 
EQ) Ah We WF. Perennial by subterranean suckers; stems glabrous, 
SON Nip Mie mostly erect, branched, 1°-3° high. Leaves lanceo- 
EW a Ve? late, petioled, dark green, acute at the apex, rounded 
Y Z\ Str a a FAX or narrowed at the base, rather firm, sharply serrate, 
WR \\\ YN glabrous on both sides, or pubescent on the veins 
; beneath, the larger 14-3’ long, 1-14’ wide; whorls 


, of flowers in terminal dense or interrupted spikes, 
WEE which are thick and obtuse, and become 17-3’ long 


eZ 
= 
TESS 


ty ip . in fruit, the middle one at length overtopped by the 
gle lateral ones; bracts lanceolate, acuminate, not longer 


than the flowers, or the lower occasionally folia- 
ceous; calyx tubular-campanulate, glabrous below, 
its teeth subulate, ciliate, one-half as long as the 
tube or more; corolla glabrous; style occasionally 
3-cleft. 


KES 
Ss YF 
Y) 


S$ SQA In wet soil, Nova Scotia to Ontario and Minnesota, 
3 south to Florida, Tennessee and ‘Arkansas. Also in 
5 Califormia, Bermuda and Jamaica. Naturalized from 


Europe. Lamb-orbrandy-mint. July—Sept. 


150 


LABIATAE. Vot. III. 


3. Mentha citrata Ehrh. Bergamot Mint. 
Fig.- 3682. 


Mentha citrata Ehrh. Beitr. 7: 150. 1792. 


Perennial by leafy stolons, glabrous throughout; 
stem weak, branched, decumbent or ascending, 1°—2° 
long. Leaves petioled, thin, ovate or ovate-orbicular, 
obtuse or the upper acute at the apex, rounded or 
subcordate at the base, sharply serrate with low 
teeth, the larger about 2’ long, 1-12’ wide; whorls 
of flowers in terminal dense thick obtuse spikes, and 
commonly also in the uppermost axils; spikes scarcely 
more than 1’ long in fruit; calyx glabrous, its teeth 
subulate, one-half as long as the tube, or longer; 
corolla glabrous. 


In wet soil, Connecticut to New York, Ohio, Michigan 
and Missouri. Naturalized from Europe. Also in Ber- 
muda and Porto Rico. July—Sept. 


4. Mentha longifolia (L.) Huds. Horse 
Mint. Fig. 3683. 


Mentha spicata var. longifolia L. Sp. Pl. 576. 1753. 
Mentha longifolia Huds. Fl. Angl. 221. 1762. 
Mentha sylvestris L. Sp. Pl. Ed. 2, 804. 1763. 


Perennial by suckers, canescent or puberulent 
nearly all over; stems mostly erect, branched, or 
simple, 1°-23° high. Leaves lanceolate, ovate- 
lanceolate, or oblong-lanceolate, sessile, or very 
short-petioled, acute at the apex, usually rounded 
at the base, sharply serrate, 1-3’ long, 3-13’ wide, 
sometimes glabrous above; whorls of flowers in 
terminal narrow dense or interrupted acute spikes, 
which become 2-5’ long in fruit; bracts lanceo- 
late-subulate, the lower equalling or longer than 
the flowers; calyx tomentose or canescent, its 
teeth subulate, one-half as long as the campanu- 
late tube; corolla puberulent. 


In waste places, Connecticut to Delaware, New Jer- 
sey and Ohio. Naturalized from Europe. Brook- or 
fish-mint. Water or European horse-mint. July—Oct. 


CORON LP 
RATES A 
CR 


WS 


5. Mentha rotundifolia (L.) Huds. 
Round-leaved Mint. Fig. 3684. 


Mentha spicata var. rotundifolia L. Sp. Pl. 576. 
1753. 


Mentha rotundifolia Huds. Fl. Angl. 221. 1762. 


tomentose-puberulent, somewhat viscid; stems 
ascending or erect, simple or branched, usu- 
ally slender, 13°-23° high. Leaves elliptic, or 
ovate-oblong, short-petioled, or sessile and 
somewhat clasping by the subcordate or 
rounded base, obtuse at the apex, crenate- 
serrate with low teeth, 1/2’ long, 9’-15” 
wide, more or less rugose-reticulated be- 
neath; whorls of flowers in terminal dense 
or interrupted spikes which elongate to 2’—4’ 
in fruit; bracts lanceolate, acuminate, com- 
monly shorter than the flowers; calyx-teeth 
setaceous, usually about one-half as long as 
the tube; corolla puberulent. 


In waste places, Maine to Florida, Ohio, Ar- 
kansas, Texas and Mexico. Bermuda. Pata- 
gonia- or apple-mint. Horse-mint. Wild mint. 
Naturalized from Europe. July—Sept. 


Perennial by leafy stolons, canescent or * 


GENUS 36. — MINT FAMILY. 


6. Mentha alopecuroides Hull Woolly Mint. 
Fig. 3685. 


Mentha alopecuroides Hull. Brit. Fl. 221. 1799. 


Perennial by suckers, white-woolly; stem stout, 
leafy, erect or ascending, simple or branched, 14°-3° 
high. Leaves broadly oval, sessile, or partly clasping 
by a subcordate or rarely rounded base, obtuse at the 
apex, sharply and rather coarsely serrate, pinnately- 
veined, the lower 2’-3’ long, 12’"-2’ wide; spikes rather 
thick, dense, stout, obtuse, 2-3’ long in fruit; bracts 
lanceolate, shorter than the flowers; calyx-teeth seta- 
ceous, one-half as long as the campanulate tube, or 
more; corolla pubescent. 


Along roadsides, Connecticut to New Jersey, Pennsyl- 
vania, Wisconsin and Missouri. Naturalized from 
Europe. July—Oct. 


7- Mentha aquatica L. Water Mint. Fish 
Mint. Fig. 3686. 
Mentha aquatica L. Sp. Pl. 576. 1753. 


Perennial by suckers, hirsute or pubescent, rarely 
glabrate; stem stout, erect, leafy, usually branched, 
13°-23° high, its hairs reflexed. Leaves broadly 
ovate, petioled, acute, subacute or the lower obtuse 
at the apex, rounded, subcordate or rarely narrowed 
at the base, sharply serrate, the larger 13’-3’ long 
and nearly as wide; whorls of flowers in terminal 
dense short thick rounded spikes, and usually also 
in the upper axils; spikes seldom more than 1’ long 
in fruit; bracts lanceolate, shorter than the flowers; 
calyx hirsute, its teeth lanceolate-subulate or tri- 
angular-lanceolate, one-third to one-half as long as 
the nearly cylindric tube; corolla sparingly pubescent. 


In wet places, Nova Scotia to Pennsylvania and 
Georgia. Naturalized from Europe. Aug.—Oct. 


8. Mentha crispa L. Crisped-leaved, 
Curled or Cross Mint. Fig. 3687. 


Mentha crispa L. Sp. Pl. 576. 1753. 
Mentha aquatica var. crispa Benth. Lab. Gen. & 

Sp. 177. “TO33: 

Sparingly pilose-pubescent at least at the 
nodes, petioles and veins of the lower surfaces 
of the leaves; stem rather weak, usually much 
branched, 13°-3° long. Leaves distinctly peti- 
oled, or the uppermost sessile, ovate in out- 
line, mostly acute at the apex, rounded, trun- 
cate or subcordate at the base, their margins 
crisped, wavy and incised, or the uppermost 
merely sharply serrate; whorls of flowers in 
dense thick rounded terminal spikes, which 
become 17-13’ long in fruit; calyx sparingly 
pubescent or glabrous, its teeth subulate, more 
than one-half as long as the campanulate tube; 
corolla glabrous. 

In swamps and roadside ditches, Connecticut 


to New Jersey and Pennsylvania. Balm-mint. 
Aug.—Oct. 


152 LABIATAE. Vor. IIT. 


g. Mentha Cardiaca Gerarde. Small-leaved 
Mint. Fig. 3688. 


Mentha Cardiaca Gerarde; Baker, Journ. Bot.2: 245. 1865. 


Pubescent or glabrate, often much branched, 13°-23° 
high, the upper branches ascending. Leaves lanceolate 
to oblong-lanceolate or ovate-lanceolate, acuminate or 
acute at the apex, the lower 2’-3’ long, the upper much 
smaller, all sharply serrate; flowers whorled in the 
upper axils; calyx-teeth subulate, about one-half as long 
as the tube. 


Wet grounds, mostly along rivers and streams, Nova 
Scotia to New Jersey, Pennsylvania and District of Colum- 
bia. July—Sept. 


10. Mentha arvénsis L. Corn Mint. Field 
Mint. Fig. 3689. 
Mentha arvensis L. Sp. Pl. 577. 1753. 


Perennial by suckers, pubescent or glabrate; stems 
erect or ascending, simple or branched, 62° high, slen- 
der. Leaves oblong-lanceolate, oval or ovate, petioled, 
acute at the apex or the lower obtuse, crenate-serrate 
with bluntish teeth, rounded at the base, 1’-23’ long, 
3/-y’ wide, the upper not much smaller than the lower; 
whorls of flowers all axillary, usually about equalling 
the petioles; calyx pubescent, campanulate, its teeth 
triangular, about as long as the width of their base, 
acute or sometimes obtuse, one-third as long as the tube. 

In dry waste places, New Brunswick to northern New 
York, New Jersey, Pennsylvania and Florida. Also in 


California and Mexico. Naturalized from Europe. Lamb’s- 
tongue. Wild pennyroyal. July—Sept. 


11. Mentha canadénsis L. American Wild Mint. Fig. 3690. 


Mentha canadensis L. Sp. Pl. 577. 1753. 

Mentha borealis Michx. Fl. Bor. Am. 2: 2. 1803. 

Mentha canadensis var. glabrata Benth. in DC. 
Prodi) 22) 17a5e 1edce 


Perennial by suckers; stem more or less pubes- 
cent with spreading or scarcely reflexed hairs, or 
glabrate, erect, simple, or branched, usually slen- 
der, 6’-23° high. Leaves oblong or ovate-oblong, 
or oblong-lanceolate, slender-petioled, acute at the 
apex, or the lower obtuse, sharply serrate, nar- 
rowed to a somewhat cuneate acute or obtuse 
base, glabrous or very sparingly pubescent, the 
larger 2’-3’ long, 3”-1’ wide; whorls of flowers 
all axillary, often shortér than the petioles; calyx 
oblong-campanulate, densely or sparingly pubes- 
cent all over, its teeth one-fourth to one-third-as 
long as the tube. 


In moist soil, New Brunswick to Manitoba, Brit- 
ish Columbia, Virginia, Nebraska, New Mexico and 
Nevada. Consists of several races, differing in 
pubescence, leaf-form and size. Odor like Penny- 
royal. July—Oct. 


GENUS 36. MINT FAMILY. 


12. Mentha gentilis L. Creeping or Downy 
Whorled Mint. Spearmint. Fig. 3691. 


Mentha gentilis L. Sp. Pl. 577. 1753. 
Mentha sativa L. Sp. Pl. Ed. 2, 805. 1763. 


Perennial by suckers; stem rather stout, ascend- 
ing or erect, branched, glabrous or puberulent 
with short reflexed hairs, 1°-2° high. Leaves 
ovate or oval, short-petioled, sparingly pubescent 
with scattered hairs on both surfaces, often blotch- 
ed, pinnately veined, acute at both ends, sharply 
serrate, the larger 13’-2’ long, the upper some- 
times much smaller than the lower; whorls of 
flowers all axillary; pedicels glabrous; calyx cam- 
panulate, glabrous below, its teeth subulate, ciliate, 
one-half as long as the tube; corolla glabrous. 


In waste places and along streams, Nova Scotia to 
northern New York, Iowa, North Carolina and Ten- 
nessee. Naturalized from Europe. Aug.—Oct. 


37. COLLINSONIA L. Sp. Fl. 28. 
1753. 


Tall perennial aromatic herbs, with large membranous petioled leaves, and terminal 
loosely panicled racemes of small, yellowish, mostly opposite flowers. Calyx campanulate, 
short, 1o-nerved, 2-lipped, declined in fruit, usually pubescent in the throat; upper lip 3-toothed; 
lower 2-cleft. Corolla much longer than the calyx, obliquely campanulate, 5-lobed, 4 of the 
lobes nearly equal, the 5th pendent or declined, fimbriate or lacerate, much larger, appearing 
like a lower lip. Anther-bearing stamens 2, not declined, much exserted, coiled before 
antithesis; bases of the filaments connected by a woolly ring; anthers 2-celled, or the sacs 
at length partially confluent. Ovary deeply 4-parted. Nutlets smooth, globose. [Named for 
Peter Collinson, 1693-1768, an English botanist, and correspondent of Linnaeus. | 

Three species, natives of eastern North America, the following typical. 


1. Collinsonia canadénsis L. MHorse- or Ox-balm. Citronella. Rich-weed. 
Fig. 3692. 


N . Collinsonia canadensis L. Sp. Pl. 28. 1753. 


Stem stout, erect or ascending, branch- 
ed, 2°-5° high, glabrous, or glandular- 
pubescent above. Leaves ovate or ovate- 
oblong, acuminate at the apex, narrowed, 
obtuse or sometimes cordate at the base, 
the lower slender-petioled, 6-10’ long, the 
ypper nearly sessile, much smaller, all 
‘oarsely dentate; racemes numerous, in 
terminal panicles sometimes 1° long; pedi- 
cels ascending, 3-6” long in fruit, subu- 
late-bracteolate at the base; flowers lemon- 
scented; calyx-teeth subulate, those of the 
lower lip much longer than the upper; 
corolla light yellow, 5-7” long; anther- 
bearing stamens 2, the upper pair rudi- 
mentary; fruiting calyx ribbed, 3’—4” long. 


In moist woods, Quebec and Ontario to 
Wisconsin, south -to Florida, Alabama and 
Arkansas. Root large, thick, woody. Knob- 
or horse-weed. Knob-grass or -root. Collin- 
son’s-flower. Richleaf. Stone-root. July—Oct. 


38. PERILLA Ard.; L. Gen. Pl. Ed. 6, Add. 578. 1764. 


Annual herbs, with petioled purple or discolored leaves, and small flowers in loose 
bracted racemes. Calyx campanulate, 1o-nerved, 5-cleft, nearly regular in flower, enlarging, 
declined and becoming 2-lipped in fruit, the upper lip 3-toothed, the lower 2-cleft, the throat 
not bearded. Corolla-tube not longer than the calyx, the throat obliquely campanulate, the 
limb 5-cleft, the lower lobe slightly the larger. Stamens 4, nearly equal, or the posterior 


154 ; LABIATAE. Vor. IIT. 


pair shorter, erect, divergent; anthers 2-celled. Style deeply 2-cleft; ovary 4-parted. Nutlets 
globose, reticulated. [The native name in India.] 


One or 2 species, natives of Asia, the following 
typical. 


1. Perilla frutéscens (L.) Britton. 
Perilla. Beef-steak Plant. Fig. 3693. 


Ocimum frutescens L. Sp. Pl. 597. 1753. - 
Perilla ocimoides L. Gen. Ed. 6, Add. 578. 1764. 
P.frutescens Britton, Mem. Torr. Cl. 5: 277. 1894. 


Purple or purple-green, sparingly pubes- 
cent; stem stout, erect, much branched, 1°-3° 
high, leafy. Leaves long-petioled, broadly 
ovate, acuminate at the apex, narrowed at 
the base, coarsely dentate or incised, 3-6’ 
long and nearly as wide; racemes terminal 
and axillary, many-flowered, 3-6’ long; pedi- 
cels spreading, 13’-3” long in fruit; calyx 
minute in flower, much enlarged, gibbous at 
the base and densely pilose-pubescent in 
fruit; corolla purple or white, 12”’ long, with 
a woolly ring within. 

In waste places, escaped from gardens, Con- 
necticut to Florida, Illinois, Missouri and Texas. 
Native of India. July—Oct. 


39. ELSHOLTZIA Willd. in Roem. & Ust. Mag. Bot. 11: 3. 1790. 

Herbs, with thin mostly petioled leaves, and small or minute clustered flowers, in ter- 
minal bracted spikes. Calyx campanulate or ovoid, 10-nerved, scarcely oblique, enlarging 
in fruit, not bearded in the throat, 5-toothed, the teeth nearly equal. Corolla-tube little 
longer than the calyx, straight, or a little curved, the limb oblique, or slightly 2-lipped, 
4-lobed; upper lobe erect, concave, emarginate, the 3 others spreading. Stamens 4, divergent, 
didynamous, ascending, exserted, the upper pair shorter; anthers 2-celled, or the sacs more 
or less confluent. Style 2-cleft at the summit. Ovary 4-parted. Nutlets ovoid or 
tuberculate, or nearly smooth. [Named in honor of , 
J. S. Elsholtz, a Prussian botanist. ] 


About 20 species, natives of Asia. Type species: 
Elsholtzia cristata Willd. 


1. Elsholtzia Patrinii (Lepech.) Garcke. 
Elsholtzia. Fig. 3694. 


Mentha Patrinii Lepech. Nov. Act. Petrop. 13: 336. 1802. 
E. cristata Willd. in Roem. & Ust. Mag. Bot. 11: 3. 1790. 
Elsholtzia Patrinii Garcke, Garcke, Fl. Deutsch. Ed. 4, 

257. 1858. 

Annual, glabrous or nearly so; stems weak, erect 
or ascending, at length widely branched, 1°-2° high. 
Leaves long-petioled, ovate or oblong, acute or 
acuminate at the apex, narrowed at the base, crenate- 
dentate, 1-3’ long; spikes terminal, very dense, 1/3’ 
high, about 3’ thick; flowers several in the axils of 
each of the broadly ovate membranous green reticu- 
lated mucronate bracts; calyx hirsute, shorter than 
the bract; corolla 1” long, pale purple. 


Notre Dame du Lac, Temiscouata Co., Quebec. Natu- 
ralized from Asia. July—Aug. 


Family 27, SOLANACEAE Pers. Syn. 1: 214. 1805.- 
Potato FAMILY. 

Herbs, shrubs, vines, or some tropical species trees, with alternate or rarely 
opposite, exstipulate entire dentate lobed or dissected leaves, and perfect regular 
or nearly regular cymose flowers. Calyx inferior, gamosepalous, mostly 5-lobed. 
Corolla gamopetalous, rotate, campanulate, funnelform, salverform or tubular, 
mostly 5-lobed, the lobes induplicate-valvate or plicate in the bud. Stamens as 
many as the lobes of the corolla and alternate with them, inserted on the tube, 
all equal and perfect in the following genera, except in Petunia, where 5 are didy- 
namous and the fifth smaller or obsolete ; anthers 2-celled, apically or longitudinally 


GENUS I. POTATO FAMILY. 155 


dehiscent. Disk present, or none. Ovary entire, superior, 2-celled (rarely 3-5- 
celled) ; ovules numerous on the axile placentae, anatropous or amphitropous; 
style slender, simple; stigma terminal; fruit a berry or capsule. Seeds numerous, 
the testa sometimes roughened ; embryo terete, spiral, curved, or nearly straight; 
endosperm fleshy ; cotyledons semiterete. 
About 75 genera and 1750 species, widely distributed, most abundant in tropical regions. 
* Fruit a pulpy berry; corolla plicate, its lobes generally induplicate. 
Anthers unconnected, destitute of terminal pores, dehiscent. 
Fruiting calyx bladdery-inflated. 
Fruiting calyx 5-angled and deeply 5-parted; ovary 3-5-celled. 1. Physalodes. 
Fruiting calyx 5-lobed, not parted, 10-ribbed, often 5—1o-angled, reticulated, wholly enclosing 


the berry ; ovary 2-celled. 
Corolla open-campanulate, yellowish or whitish, often with a dark center: seeds with a 


thin margin, finely pitted. 2. Physalis. 
_ Corolla flat-rotate, violet or purple; seeds thick, rugose-tuberculate. 3. Quincula. 
etd Saeeonc what enlarged, but closely fitted to the fruit, thin, obscurely veiny, open at 
the mouth. 


Corolla rotate, whitish ; lobes of fruiting calyx much exceeding the berry. 4. Leucophysalis. 
Corolla rotate, whitish, sometimes tinged with purple; fruiting calyx not exceeding the 
berry. 5. Chamaesaracha. 
Anthers connivent or slightly connate; fruiting calyx not enlarged. 
Anthers short or oblong, opening by a terminal pore or short slit in our species. 6. Solanum. 


Anthers long, tapering from base to summit, longitudinally dehiscent. 7. Lycopersicon. 
** Fruit a nearly dry berry; corolla campanulate, little or not at all plicate, its lobes imbricated. 
: 8. Lycium. 


*** Fruit a capsule; corol lform. 
Capsule circumscissile toward the top, whic arates asalid;corollairregular. 9. Hyoscyamus. 


Capsule opening by valves. 


Capsule generally prickly. 10. Datura. 
Capsule not prickly. 
Flowers paniculate or racemose; stamens nearly uniform in length. 11. Nicotiana. 
Flowers solitary ; stamens very unequal. 12. Petunia. 


1. PHYSALODES Boehm. in Ludwig, Def. 41. 1760, 
[Nicanpra Adans. Fam. Pl. 2: 219. 1763.] 

An annual erect branching glabrous herb, with alternate petioled thin sinuate-dentate or 
lobed leaves, and large light blue peduncled nodding flowers, solitary in the axils. Calyx 
5-parted, 5-angled, much inflated in fruit, its segments ovate, connivent, cordate or sagittate 
at the base, strongly reticulated. Corolla broadly campanulate, plicate in the bud, slightly 
5-lobed. Stamens 5, included, inserted on the corolla near its base; filaments filiform, dilated 
and pilose below; anthers oblong, the sacs longitudinally dehiscent. Ovary 3-5-celled; style 
slender; stigma 3-5-lobed. Berry globose, nearly dry, enclosed in the calyx. [Greek, Physalis- 
like.] 


A monotypic Peruvian genus. 


1. Physalodes physalodes (L.) Britton. 
Apple-of-Peru. Fig. 3695. 
Atropa physalodes L. Sp. Pl. 181. 1753. 
Physalodes peruvianum Kuntze, Rev. Gen. Pl. 452. 

1891. 

P. physalodes Britton, Mem. Torr. Club 5: 287. 18094. 

Stem angled, 2°-5° high. Leaves ovate or 
oblong, acuminate but blunt-pointed, narrowed at 
the base, 3-8’ long, 1-4’ wide; petioles longer 
than the peduncles; flowers 17-14’ long and 
broad; corolla-limb almost entire; fruiting calyx 
1’-14’ long and thick, its segments acute at the 
apex, their basal auricles acute or cuspidate; berry 
about 4’ in diameter, loosely surrounded by the 
calyx. 

In waste places, escaped from gardens, Nova Sco- 
tia to Ontario, Florida, Tennessee and Missouri. 
Adventive from Peru. Plant with the aspect of a 
large Physalis. Leaves similar to those of Stramo- 
nium. July—Sept. 


2. PHYSALIS L. Sp. Pl. 182. 1753.* 


Annual or perennial herbs, sometimes a little woody below, with entire or sinuately 
toothed leaves. Peduncles slender, in ours solitary from the axils of the leaves. Calyx 
campanulate, 5-toothed, in fruit enlarged and bladdery-inflated, membranous, 5-angled, or 
prominently 10-ribbed and reticulate, wholly inclosing the pulpy berry, its teeth mostly 


* Text contributed to our first edition by Dr. P. A. Rypperc, here somewhat revised. 


- 


156 SOLANACEAE. Vor. III. 


connivent. Corolla yellowish or whitish, often with a darker brownish or purplish center, 
open-campanulate, or rarely campanulate-rotate, plicate. Stamens inserted near the base of 
the corolla; anthers oblong, opening by a longitudinal slit. Style slender, somewhat bent; 
stigma minutely 2-cleft. Seeds numerous, kidney-shaped, flattened, with a thin edge, finely 
pitted. [Greek, bladder, referring to the inflated calyx.] 

Probably 75 species, or more. Two are of European origin, and about half a dozen are natives 
of India and Australia, the rest are American; about 30 occur in the United States. Type 
species: Physalis Alkekéngi L. 

* Annuals with branched fibrous roots. 
+ Plants more or less pubescent (except in races of P. barbadensis.) 


Fruiting calyx sharply 5-angled, more or less acuminate at the summit and sunken at the base; 
calyx-lobes (at flowering time) lanceolate or acuminate, as long as the tube or longer. , 
Leaves ovate, oblique, acute or acuminate, subentire at the base; upper part repand or suben- 
tire ; fruiting calyx small and short ; stem slender, diffuse, sharply angled. 1. P. pubescens. 
Leaves cordate, oblique, strongly sinuate to the base; stem stout, obtusely angled; fruiting 


calyx rounded. 2. P. pruinosa. 
Leaves cordate, scarcely oblique, more or less abruptly acuminate, acutely repand-dentate; stem 
tall, acutely angled; fruiting calyx larger, long-acuminate. 3. P. barbadensis. 
Fruiting calyx obtusely or indistinctly 5-10-angled; calyx-lobes (at flowering time) triangular, 
generally shorter than the tube. 4. P. missouriensis. 


77 Plants glabrous, or the upper part sparingly beset with short hairs, or a little puberulent 
when young; fruiting calyx obtusely 5—10-angled, not sunken at the base. 
Corolla yellow, sometimes with the center a little darker but never brown or purple. 
Peduncles generally much longer than the fruiting calyx; leaves sinuately toothed or subentire. 


d ie 5. P. pendula. 
Peduncles scarcely exceeding the fruiting calyx; leaves sharply dentate. 6. P. angulata. 
Corolla yellow, with a brown or purple center. 7. P.ixocarpa. 


** Perennial by thick roots and rootstocks. 
7 Pubescence not stellate (although in P. pumila of branched hairs). 


Pubescence on the leaves none, on the upper part of the stem and the calyx sparse and short, if any. 
Fruiting calyx ovoid, nearly filled by the berry, scarcely sunken at the base. 


Leaves ovate-lanceolate to broadly ovate, usually thin. 8. P. subglabrata. 
Leaves lanceolate, oblanceolate, or linear. 9. P. longifolia. 
Fruiting calyx pyramidal, very much inflated and deeply sunken at the base; leaves broadly 
ovatc, usually coarsely dentate. 10. P. macrophysa. 


Pubescence sparse, consisting of flat, sometimes jointed, and in P. pumila branched hairs; in 
P. virginiana sometimes a little viscid. 
Fruiting calyx ovoid, scarcely angled and scarcely sunken at the base; leaves thick, obovate or 
spatulate to rhomboid, subentire. 
Leaves obovate or spatulate; hairs all simple. 11. P. lanceolata. 
Leaves broader, often rhomboid; hairs on the lower surface branched. 12. P. pumila. 
Fruiting calyx pyramidal, more or less 5-angled and deeply sunken at the base; leaves ovate to 


lanceolate, generally more or less dentate. 13. P. virginiana. 
Pubescence dense, short, more or less viscid or glandular, often mixed with long flat jointed hairs, 
Leaves large; blade generally over 2’ long and more or less cordate. 14. P. heterophylla. 
Leaves less than 2’ long, rounded ovate or rhombic, scarcely at all cordate at the base; calyx, 
peduncles and younger branches with long white flat and jointed hairs. 15. P. comata. 
Leaves small, 1’-124’ in diameter, nearly orbicular, sometimes a little cordate at the base, not 
coarsely toothed; stem diffuse or prostrate. 16. P. rotundata. 


++ Pubescence dense, cinereous, beautifully stellate. 17. P. viscosa. 


1. Physalis pubéscens L. Low Hairy 
Ground-Cherry. Fig. 3696. 


Physalis pubescens L. Sp. Pl. 183. 1753. 


Annual; stem generally diffuse or spreading, 
much branched, angled, often a little swollen at 
the nodes, villous-pubescent or sometimes nearly 
glabrous; leaves thin, 1-22’ long, ovate, acute 
or acuminate, at the base oblique, slightly cor- 
date and generally entire, upward repand-den- 
ticulate or entire, pubescent, sometimes becoming 
nearly glabrous except along the veins; peduncles 
short, 1-2” long, or in fruit about 5”; calyx- 
lobes narrow but not with a subulate tip; corolla 
3-5’ in diameter, yellow with dark centre; an- 
thers usually purplish; fruiting calyx membra- 
nous, 10’-15” long, pyramidal, ovoid-acuminate, 
more or less retuse at the base. 


In sandy soil, Pennsylvania to Florida, Arkansas 
and California; also in Mexico, the West Indies, 
Central and South America and India. Called also 
dwarf cape-gooseberry and strawberry-tomato. July— 
Sept. 


GENUS 2. POTATO FAMILY, 15 / 


2. Physalis pruinosa L. Tall Hairy 
Ground-Cherry. Fig. 3697. 


Physalis pruinosa L. Sp. Pl. 184. 1753. 
P. pubescens Dunal, in DC. Prodr. 13: part 1, 

446. 1852. 

Annual; stem stout, generally erect, and 
more hairy than the preceding and the two 
following species; stem obtusely angled, finely 
villous or somewhat viscid; leaves firm, 14’-4’ 
long, finely pubescent, ovate, cordate, generally 
very oblique at the base, and deeply sinuately 
toothed with broad and-often obtuse teeth; 
peduncles 1-2” long, in fruit about 5”; calyx 
villous or viscid; lobes as long as the tube, 
narrow but not subulate-tipped; corolla 2’—4” 
in diameter; anthers yellow, or tinged with 
purple; fruiting calyx a little firmer and more 
pubescent than in the preceding, reticulate, 
10’—-15” long, ovoid, sunken at the base; berry 
yellow or green. 


In cultivated soil, Massachusetts to Ontario, 
Florida, Iowa, Missouri and Colorado. July—Sept. 


3. Physalis barbadénsis Jacq. Barbadoes 
Ground-Cherry. Fig. 3608. 


Physalis barbadensis Jacq. Misc. 2: 359. 1781. 
Physalis obscura Michx. Fl. Bor. Am. 1: 149. 1803. 
Physalis barbadensis obscura Rydberg, Mem. Torr. Club 

4')-327.. 1806. 

Annual; stem tall and erect or widely spreading, 
acutely 3-4-angled, pubescent, viscid, or nearly gla- 
brous; leaves 13-22’ long, ovate or heart-shaped, acute, 
or abruptly acuminate, sharply repand-dentate, pubes- 
cent with short hairs; peduncles short, 13’—-2” long, but 
in fruit sometimes 10”; calyx generally densely viscid- 
hirsute, lobes lanceolate, acuminate, but not subulate- 
tipped; corolla 23’—5’’ in diameter; anthers generally 
purplish; fruiting calyx longer than in the two pre- 
ceding species, 1-14’ long, acuminate and reticulate, 
retuse at the base. 

Sandy soil, Pennsylvania to Illinois, Missouri, Florida, 
Mexico, the West Indies and South America. July—Sept. 


\S 


4. Physalis missouriénsis Mack. & Bush. 
Missouri Ground-Cherry. Fig. 3699. 


Physalis missouriensis Mack. & Bush, Fl. Jackson 
Gos 1672 “1902: 


Annual; stem spreading, often zigzag, branched, 
striate, or slightly angled, villous with short hairs. 
Leaves 3’-33’ long, ovate, oblique and cuneate, 
obtuse, or cordate at the base, acute but not acu- 
minate, repand or sinuately dentate, hairy, at least 
on the veins; peduncles 4’—23” long, erect, in fruit 
23-5", reflexed, shorter than the fruiting calyx; 
calyx villous, lobes shorter than the tude, triangu- 
lar; corolla 14’-4” in diameter, yellow; fruiting 
calyx 7’-10” long, round-ovoid, nearly filled by 
the berry, scarcely sunken or commonly rounded 
at the base. 

Missouri and Kansas to Arkansas and Oklahoma. 


July—Sept. Referred in our first edition to the 
tropical P. Lagdscae R. & S. 


158 SOLANACEAE. Vor. III. 


5. Physalis péndula Rydberg. Lance-leaved 
Ground-Cherry. Fig. 3700. 


Physalis pendula Rydberg ; Small, Fl. SE. U. S. 983. 1903. 


Annual, stem erect, generally 13° high, branched, 
angled, glabrous; leaves lanceolate or ovate-lanceo- 
late, thin, usually coarsely toothed; calyx cylindric- 
campanulate, its lobes broadly triangular, shorter 
than the tube; peduncles filiform, about 1’ long, erect 
with nodding flower, in fruit 12-2’ long and re- 
flexed; corolla 3-4” in diameter, campanulate, yel- 
low, without a dark spot; anthers yellow, more or 
less tinged with purple, fruiting calyx about 10” long, 
rounded ovoid, indistinctly to-angled and purple 
veined, nearly filled by the berry. 


Illinois to Kansas and Texas. July—Sept. Referred in 
the first edition to P. lanceifolia Nees. 


Physalis angulata L. Sp. Pl. 183. 1753. 


Annual, erect, 13°-3° high, glabrous; stem 
angular, usually much branched; leaves ovate, with 
more or less cuneate base, somewhat sinuately 
toothed with long-acuminate teeth; blades 2’—23’ 
Jong, on slender petioles 1-2’ long, thin, the veins 
not prominent; peduncles slender, 10’—15” long, 
erect, in fruit often reflexed but seldom exceed- 
ing the fruiting calyx in length; calyx smooth, 
lobes triangular to lanceolate, generally shorter 
than the tube; corolla 23’—5” in diameter; anthers 
more or less purplish tinged; fruiting calyx about 
17 long, ovoid, 5-10-angled, sometimes purple- 
veined, nearly filled by the yellow berry. 


In rich soil, Pennsylvania to Illinois, Minnesota, 
Missouri, Texas, Central America, Brazil and the 
West Indies. Also in India. July—Sept. 


7. Physalis ixocarpa Brot. Tomatillo. 
Mexican Ground-Cherry. Strawberry 
Tomato. Fig. 3702. 


Physalis ixocarpa Brot.; Horneman, Hort. Hafn. 

Suppl. 26. 1819. 
P. aequata Jacq. f.; Nees, Linnaea 6: 470: 1831. 

Annual, stem at first erect, later widely spread- 
ing, much branched, angled, glabrous, or the 
younger parts sparingly hairy; leaves from cor- 
date to ovate, with a cuneate base which is some- 
what oblique, sinuately dentate or entire, 1-23’ 
long; peduncles short, 1-23” long; calyx sparingly 
hairy, its lobes short, broadly triangular, shorter 
than the tube; corolla bright yellow with purple 
throat, 5’-7” (sometimes nearly 10”) in diameter; 
fruiting calyx round-ovoid, obscurely 10-angled, 
often purple-veined, filled by the purple berry, 
which sometimes bursts it. 

Native of Mexico. It is often cultivated for its 


fruit and frequently escapes from cultivation, New 
York to Texas and California. 


GENUS 2. POTATO FAMILY. 159 


8. Physalis subglabrata Mackenzie and 
Bush. Smooth Ground-Cherry. 
Fig. 3703. 


?Physalis philadelphica Lam. Encycl. 2: ror. 1786. 
P. subglabrata Mackenzie & Bush, Trans. Acad. St. 
Louis 12: 86. 1902. 


Perennial from a deep rootstock, tall, erect, 23°-5° 
high; stem angled, dichotomously branched, gla- 
brous, or sometimes slightly pubescent with sparse 
and short hairs on the upper parts; blades ovate to 
ovate-lanceolate, often very oblique at the base and 
more or less acuminate, entire or repand-denticulate, 
23-4’ long, on petioles 13’-23’ long, often in pairs; 
peduncles slender, 5’—10” long, generally longer than 
the flower; calyx glabrous, or minutely ciliolate, 
lobes ovate-lanceolate or trangular, sometmes broadly 
ovate and unequal, generally equalling the tube; 
corolla yellow or greenish yellow with purplish 
throat, #’-1’ in diameter; anthers tinged with pur- 
ple; fruiting calyx at first somewhat 10-angled and 
sunken at the base, at last often filled with or burst 
by the large red or purple berry. 


Ontario and Rhode Island to Georgia, Minnesota, Kentucky and Colorado. July—Sept. 


g. Physalis longifolia Nutt. Long-leaved Ground-Cherry. Fig. 3704. 
| FY 


Physalis longifolia Nutt. Trans. Am. Phil. Soc. (II.) 5: 
193. 1833-37. 


Physalis lanceolata var. laevigata A. Gray, Proc. Am. 
Acad. 10: 68. 1874. 


Perennial from a thick rootstock; stem in the com- 
mon form stout and tall, 13°-3° high, slightly angled, 
branched above, the branches . strict, glabrous. 
Leaves lanceolate, oblanceolate, or linear, tapering 
into a short stout petiole 5’—10” long, subentire or 
repand; peduncles 5-10” long, in fruit often re- 
curved; calyx generally glabrous, its lobes triangular- 
lanceolate, about the length of the tube; corolla 6”— 
10” in diameter, yellow with a dark, commonly 
brownish center; anthers yellow, tinged with purple; 
fruiting calyx ovoid, about 17’ long, not sunken at 
the base; berry yellow, the lower portion and the 
stipe glutinous. 


In rich soil, Iowa to South Dakota, Montana, Arkan- 
sas, Utah and Mexico. July—Sept. 


10. Physalis macréphysa Rydb. Large- 
bladder Ground-Cherry. Fig. 3705. 


P. macrophysa Rydberg, Bull. Torr. Club 22: 308. 1895. 


Perennial; rootstock rather thick and fleshy; stem 
erect, 12°-3° high, comparatively slender, angled, 
perfectly glabrous, or the upper parts sparingly pu- 
bescent with very short hairs. Leaves large, thin, 
14’-33’ long, 1’-2’ wide, the lower obtuse, the upper 
acute or acuminate; petioles slender, 10’—20” long; 
peduncles 5’-8” long, erect, in fruit reflexed; calyx 
smooth, its lobes ovate-triangular or broadly lanceo- 
late, generally a little shorter than the tube; corolla 
yellow with a dark center, about 10” in diameter; 
anthers generally yellow, sometimes tinged with pur- 
ple; fruiting calyx large, 1}’-13’ long, 1-11’ in diam- 
eter, pyramidal to ovoid-conic, indistinctly 1o-angled, 
deeply sunken at the base; berry small, in the center 
of the calyx. 


In rich soil, Kansas and Arkansas to Texas. May- 
July. Rare. 


160 SOLANACEAE. Vo. IIT. 


11. Physalis lanceolata Michx. Prairie 
Ground-Cherry. Fig. 3706. 
Physalis lanceolata Michx. Fl. Bor. Am. 1: 149. 1803. 


Physalis pennsylvanica var. lanceolata A. Gray, Man. 
Ed. 5, 382. 1867. 


Perennial; rootstock apparently slender and creep- 
ing; stem about 13° high, first erect, later spreading 
or diffuse, only slightly angled, sparingly hirsute with 
flat hairs. Leaves lanceolate, oblanceolate or spatu- 
late, tapering into the petiole, acute or obtuse, nearly 
always entire, rarely wavy, but never sinuately tooth- 
ed, thickish, sparingly hairy with short hairs; pe- 
duncles 5-10” long, in fruit reflexed; calyx strigose 
or villous, rarely glabrous, its lobes triangular-lan- 
ceolate; corolla dullish yellow with a brownish cen- 
ter, about 8” in diameter; fruiting calyx round- 
ovoid, not sunken at the base, indistinctly to-angled; 
berry yellow or greenish yellow. 


On dry prairies, South Carolina to Illinois, South Da- 
kota, Wyoming, Kansas and New Mexico. July—Sept. 


12. Physalis pumila Nutt. Low Ground-Cherry. Fig. 3707. 


Physalis pumila Nutt. Trans. Am. Phil. Soc. (II.) 5: 
193. 1834. ; 

Physalis lanceolata var. hirta A. Gray, Proc. Am. Acad. 
10: 68. 1874. 


Perennial from a slender rootstock, 13°-3° high; 
stem hirsute, obscurely angled. Leaves thick, broadly 
ovate to oblong, acute at both ends and somewhat 
rhomboid, the lower often obtuse and obovate, gen- 
erally much larger than in the preceding; blades 
2’-4’ long, entire or seldom sinuate, on petioles 
10-15” long, strigose with many-branched hairs 
especially on the lower surface; peduncles 5’—10” 
long, in fruit reflexed and 14’-2’ long; calyx 
densely hirsute, not stellate-pubescent, its lobes tri- 
angular, generally a little shorter than the tube; 
corolla yellow with brown center, 8-10” in diam- 
eter; fruiting calyx usually more elongated than in 
the preceding, 13-2’ long, oblong-ovoid, a little 
sunken at the base, indistinctly 1o-angled. 


Plains and prairies, Illinois to Colorado and Texas. 
July—Sept. 


13. Physalis virginiana Mill. Virginia Ground-Cherry. Fig. 3708. 


A. Physalis virginiana Mill. Gard. Dict. Ed. 8, no. 4. 1768. 

E a, “4 Physalis pennsylvanica A. Gray, Man. Ed. 5, 382. 1867. 

LNEé4 Tied yA Not Liwr753- 

\. ‘\ foe Physalis virginiana intermedia Rydberg, Mem. Torr. Club 
oy a fe g 4: 345. 18096. 


Perennial; rootstock thick and somewhat fleshy; 
stem 13°-3° high, erect, dichotomously branched, 
somewhat angular, more or less strigose-hairy with 
flat hairs, sometimes a little glandular, or sometimes 
nearly glabrous. Leaves ovate-lanceolate, tapering to 
both ends, 13’-23’ long, generally more or less sin- 
uately dentate, often yellowish green; peduncles 5” 


=1097 
long, generally erect, in fruit curved but scarcely re- 
flexed; calyx strigose, hirsute, or at least puberulent, 
its lobes triangular or broadly lanceolate, nearly equal- 
ling the tube; corolla sulphur-yellow with purplish 
spots, 3-1’ in diameter; anthers yellow; fruiting calyx 
pyramidal-ovoid, 5-angled, sunken at the base; berry 
reddish. 


Rich soil, especially in open places, Ontario to Manitoba, 
; w 4 Connecticut, Florida, Louisiana and Texas. Consists of 
numerous races, differing in pubescence. July-—Sept. Wild cherry. 


GENUS 2. POTATO FAMILY. 161 


14. Physalis heterophylla Nees. Clammy Ground-Cherry. Fig. 3709. 
Physalis viscosa Pursh, Fl. Am. Sept. 157. 1814. Not 


- 1753- 
Physalis heterophylla Nees, Linnaea 6: 463. 1831. 
Physalis virginiana A. Gray, Syn. Fl. 2: Part 1, 235. 
1878. Not Mill. 1768. 


Perennial from a slender creeping rootstock, 14°-3° 
tall, at first erect, later generally decumbent and 
spreading, viscid and glandular, and villous with 
long spreading jointed flat hairs; leaves large, blade 
generally over 2’ long, usually broadly cordate, often 
acute and very rarely with an elongated tip, thick, 
more or less sinuately toothed, or sometimes suben- 
tire; calyx long-villous, lobes triangular, generally 
shorter than the tube; corolla 8’—-10” in diameter, 
greenish yellow with a brownish or purplish center; 
anthers mostly yellow; berry yellow. 

In rich soil, especially where the surface has been 
disturbed, New Brunswick to Saskatchewan, Florida, 


Colorado and Texas. The most common of our species, 
and includes several races. 


Physalis peruviana L., a native of South America, is cultivated for its fruit and often escapes. 
It resembles P. heterophylla, but differs in the leaves, which have a long tip, and in the pubes- 
cence, which is shorter, denser, and not at all viscid. Cape-gooseberry. Strawberry-tomato. 
Peruvian ground-cherry. Husk-tomato. 


15. Pnysalis comata Rydberg. Hillside 
Ground-Cherry. Fig. 3710. 
P. comata Rydberg, Bull. Torr. Club 22: 306. 1895. 


Perennial, erect, about 12° high; pubescence fine 
and short, that on the calyx, peduncles and upper 
branches mixed with long white flat jointed hairs. 
Like P. heterophylla Nees (P. virginiana Gray, not 
Mill.), but leaves smaller, blade not over 2’ long, 
round-ovate, scarcely at all cordate at the base, about 
2’ long, thin, somewhat repand-dentate, or nearly 
entire; petioles as long as the leaves; peduncles as 
long as the fruiting calyx, or longer; corolla green- 
ish yellow, with brown center, 6”-10” in diameter; 
fruiting calyx of thin texture, round-ovoid, some- 
what 10-angled, scarcely sunken at the base. 


Hillsides of Nebraska, Kansas, Colorado and Texas. 
Rare. 


16. Physalis rotundata Rydberg. Round- 
leaved Ground-Cherry. Fig. 3711. 


Physalis hederaefolia Holzinger, Cont. U.S. Nat. Herb. 
I: 212. 1892. Not Gray. 
P. rotundata Rydberg, Mem. Torr. Club 4: 352. 1896. 
Diffuse and spreading, zigzag, generally dichoto- 
mously much branched, from a perennial rootstock, 
densely and finely viscid-pubescent, usually more 
glandular than the preceding. Leaves nearly orbicu- 
lar with more or less cordate base, 1-13’ in diam- 
eter, with small teeth; petioles short, more or less 
winged; peduncles short, in fruit scarcely more 
than half the length of the calyx; corolla 8” in 
diameter, greenish yellow with a brownish center; 
fruiting calyx ovoid, slightly angled, scarcely sunken 
at the base. 


Dry plains, South Dakota to Texas and New Mexico. 
July—Sept. 


It 


162 SOLANACEAE. Vor. III. 


17. Physalis viscdsa L. Stellate Ground- 
Cherry. Yellow-henbane. Fig. 3712. 


Physalis viscosa L. Sp. Pl. 183. 1753. 
Physalis pennsylvanica L. Sp. Pl. Ed. 2, 1670. 1763. 


Perennial from a slender creeping rootstock; stems 
slender, creeping, with a dense ashy stellate pubes- 
cence, or in age rarely glabrate. Leaves elliptic, 
oval or ovate, obtuse, thinish, entire or undulate, in 
the typical South American race often cordate at 
the base, but rarely so in our plant; peduncles 3-1’ 
long; calyx stellate-pubescent, its lobes triangular, 
generally shorter than the tube; corolla greenish 
yellow with a darker center, 8’-10” in diameter; 
fruiting calyx 10-15” long, round-ovoid, scarcely 
sunken at the base; berry orange or yellow. 


On sea beaches, or in sand near the coast, Virginia 
and North Carolinato Florida. Eastern South America. 


Physalis Alkekéngi L., Strawberry tomato or Winter cherry, is a native of Europe and Asia, 
often cultivated for its fruit and sometimes escapes from cultivation. The flowers are whitish, 
the limb distinctly 5-lobed; leaves broadly deltoid, acute at both ends, repand or angulately 


toothed. 


3. QUINCULA Raf. Atl. Journ. 145. 1832. 


A low and diffuse somewhat scurfy herb, with a stout perennial root. Leaves from sinuate 
to pinnatifid, somewhat fleshy. Peduncles most commonly in pairs from the axils of the 
leaves, sometimes solitary, or in fascicles of 3-5. Calyx campanulate, 5-toothed, in fruit 
inflated, sharply 5-angular and reticulate, enclosing the fruit, the lobes connivent. Corolla 
flat-rotate, pentagonal in outline, veiny, violet or purplish. Anthers opening by a longi- 
tudinal slit. Seeds comparatively few, kidney-shaped, somewhat flattened, with thick margins, 
rugose-tuberculate. [Name unexplained. ] 


A monotypic genus of central North America. 


1. Quincula lobata (Torr.) Raf. Purple- 
flowered Ground-Cherry. Fig. 3713. 


Physalis lobata Torr. Ann. Lyc. N. Y. 2: 226. 1827. 
Quincula lobata Raf. Atl. Journ. 145. 1832. 


Perennial, low, spreading or prostrate, more or 
less scurfy-puberulent; stem obtusely angled and 
striate, much branched. Leaves oblanceolate or 
spatulate to oblong, sinuately toothed, or pinnatifid 
with rounded lobes, or rarely subentire, cuneate at 
the base, thickish and veiny, tapering into margined 
petioles; peduncles 1-2’ long,in fruit reflexed; calyx- 
lobes triangular, acute, shorter than the tube; corolla 
purplish, 10’-15” in diameter; anthers yellow, tinged 
with purple; fruiting calyx about as wide as long, 
sharply 5-angled, sunken at the base. 


On high plains, Kansas to California, Texas, New 
Mexico and Mexico. May-—Sept. 


4. LEUCOPHYSALIS Rydberg, Mem. Torr. Club 4: 365. 1806. 


A tall erect viscid and villous annual, with entire leaves, the blade decurrent on the 
petiole. Peduncles generally in fascicles of 2-4 in the axils. Calyx campanulate, 5-lobed, at 
first a little inflated, but soon filled by and closely fitted to the berry, thin, neither angled 
nor ribbed, faintly veiny, open at the mouth, the lobes exceeding the fruit. Corolla rotate, 
white, sometimes tinged with purple and generally creamy or yellow in the center, the limb 
plicate. Stamens inserted near the base of the corolla; filaments long and slender; anthers 
oblong, opening by a longitudinal slit. Style and stigmas as in Physalis. Seeds kidney- 
shaped, flattened, punctate. [Greek, white Physalis.] 


A monotypic genus of northern North America. 


—— 


GENUS 4. POTATO FAMILY. 163 


1. Leucophysalis grandiflora ( Hook.) 
Rydberg. Large White Ground- 
Cherry. Fig. 3714. 

P. grandiflora Hook. Fl. Bor..Am. 2: 90. 1834. 
Leucophysalis grandiflora Rydberg, Mem. Torr. 
Club 4: 366. 1896. ; 
Erect, tall, 13°-3° -high; stem somewhat 
angled, striate, more or less villous. Leaves 
large, 4’-8’ long, ovate to lanceolate-ovate, 
generally acute and entire, somewhat decur- 
rent on the petiole, more or less villous and 
viscid, especially on the veins of the lower 
surface; peduncles several from each axil, 
‘4’ long, villous; calyx villous, its lobes 
lanceolate, equalling the tube; corolla large, 
14-13’ in diameter, rotate, white with a more 
or less yellowish center; filaments slender; 
anthers short, yellow, often tinged with pur- 
ple; fruiting calyx ovoid, early filled by the 

berry. 


Sandy soil, Quebec to Saskatchewan, Mich- 
igan and Minnesota. May-July. 


5. CHAMAESARACHA A. Gray, Bot. Cal. 1: 540. 1876. 


Perennials, with entire to pinnatifid leaves, the blades decurrent on the petioles. Peduncles 
solitary, or in fascicles of 2-4 in the axils. Calyx campanulate, 5-lobed, in fruit somewhat 
enlarged, but not bladdery-inflated, close-fitting to the berry, thin, not angled nor ribbed, and 
faintly if at all veiny, open at the mouth, not exceeding the berry. Corolla rotate, white or 
cream-colored, often tinged with purple, the limb plicate. Stamens inserted near the base 
of the corolla; filaments long and slender; anthers oblong, opening by longitudinal slits ; 
style and stigma as in Physalis. Seeds kidney-shaped, flattened, rugose-favose or punctate. 
[Ground-Saracha, the latter a genus named in honor of Isidore Saracha, a Spanish Bene- 
dictine botanist. | 

An American genus, consisting of half a dozen species, natives of Mexico and the south- 
western United States. Type species: Chamaesaracha Coronopus (Dunal) A. Gray. 


Pubescence dense, puberulent and hirsute. 1. C. conioides. 
Pubescence sparse, puberulent or stellate, hirsute (if at all) only on the calyx. 2. C. Coronopus. 


1. Chamaesaracha conioides (Moricand) Brit- 
ton. Hairy Chamaesaracha. Fig. 3715. 


Solanum conioides Moric.; Dunal in DC. Prodr. 13: Part 
Withaia (i) sordiile Dunal, Joc. cit., 456. 1852. 
Chamaesaracha sordida A. Gray, Bot. Cal. 1: 540. 1876. 
Chamaesaracha conioides Britton, Mem. Torr. Club 5: 

287. 1895. 

Much branched from a perennial base, at first up- 
right, at length spreading, cinereous-puberulent with 
short branched somewhat glutinous or viscid hairs, 
generally also viscidly hirsute or villous with long and 
branched hairs, especially on the calyx; leaves oblan- 
ceolate to obovate-rhombic, usually acutish and taper- 
ing into a short petiole, generally deeply lobed, but 
varying from subentire to pinnatifid; calyx-lobes tri- 
angular, generally acutish; corolla about }#’ in diam- 
eter, white, cream-colored or sometimes violet-pur- 
plish; berry 24’-4” in diameter. 


In dry clayey soil, southern Kansas to California and 
Mexico. May-Sept. 


LF 2. Chamaesaracha Cordnopus (Dunal) 


A. Gray. Smoothish Chamaesaracha. 


\ of 7 Fig. 3716. 
VZ4 me Bilal Et vi Dunal in DC. Prodr. 13: Part 1, 
’ E 4. 1852. 
y C. Coronopus A. Gray, Bot. Cal. 1: 540. 1876. 
Branched and diffuse from a perennial base; 
ee stem obtusely angled; pubescence on the stem 
and leaves more or less roughish pruinose or 
stellate, often scarcely any; on the calyx stellate 
or sometimes hirsute. Leaves linear or lanceo- 
late, tapering at the base, more or less sinuately 
lobed, occasionally subentire, sometimes pinna- 
tifid; calyx-lobes triangular, acute; corolla white 
or ochroleucous, the appendages of the throat 
often protuberant; berry 22’-4” in diameter, 
nearly white. 


In. clayey soil, Kansas to Utah, California and 
Mexico. May—Sept. 


6. SOLANUM [Tourn.] L. Sp. Pl. 184. 
1753. 


Herbs or shrubs, often stellate-pubescent, sometimes climbing. Flowers cymose, umbel- 
liform, paniculate, or racemose, white, blue, purple, or yellow. Calyx campanulate or rotate, 
mostly 5-toothed or 5-cleft. Corolla rotate, the limb plaited, 5-angled or 5-lobed, the tube 
very short. Stamens inserted on the throat of the corolla; filaments short; anthers linear or 
oblong, acute or acuminate, connate or connivent into a cone, the cells dehiscent by a terminal 
pore, or sometimes by a short introrse terminal slit, or sometimes also longitudinally. Ovary 
usually 2-celled; stigma small. Berry mostly globose, the calyx either persistent at its base 
or enclosing it. [Name, according to Wettstein, from solamen, quieting. | 


About 1000 species, of wide geographic distribution, most abundant in tropical America. 
Besides the following, some 20 others occur in the southern and western United States. Type 
species: Solanum nigrum L. 

* Glabrous or pubescent herbs, not prickly. 
Plants green; pubescence simple, or some of it stellate; flowers white. 


Leaves repand or entire; ripe berries black. 1. S. nigrum. 
Leaves deeply pinnatifid ; ripe berries green. 2. S. triflorum. 
Plant silvery stellate-canescent ; flowers violet. 4. S. elaeagnifolium. 
** Stellate-pubescent and prickly herbs. 
Berry not enclosed by the calyx; perennials. 
Hirsute; leaves ovate or oblong, sinuate or pinnatifid. 3. S. carolinense. 
- Densely silvery-canescent ; leaves linear or oblong, repand or entire. 4. S. elaeagnifolium. 
Pubescent ; leaves ovate, 5—7-lobed. 5. S. Torrey. 
Berry partly or wholly invested by the spiny calyx; annuals. 
Lowest anther larger than the other four. 
Plant densely stellate-pubescent ; corolla yellow. 6. S. rostratum. 
Plant glandular-pubescent, with few stellate hairs ; corolla violet. 7. S. citrullifolium. 
Anthers all equal. 8. S. sisymbrifolium. 


*** Climbing vine, not prickly; leaves hastate or 3-lobed. 9. S. Dulcamara. 


v 


1. Solanum nigrum L. Black, Deadly or 
Garden Nightshade. Morel. Fig. 3717. 
Solanum nigrum L. Sp. PI. 186. 1753. 


Annual, glabrous, or somewhat pubescent with 
simple hairs, green; stem erect, branched, 1°-22° 
high. Leaves ovate, petioled, more or less inequi- 
lateral, 1’-3’ long, entire, undulate, or dentate, thin, 
acute, acuminate or acutish at the apex, narrowed 
or rounded at the base; peduncles lateral, umbel- 
lately 3-10-flowered, 3’-13’ long; pedicels 3-7” long; 
flowers white, 4”-5” broad; calyx-lobes oblong, ob- 
tuse, spreading, much shorter than the corolla, per- 
sistent at the base of the berry; filaments somewhat 
pubescent; anthers obtuse; berries black when ripe, 
smooth and glabrous, globose, 4”—-5” in diameter, on 
nodding peduncles. 

In waste places, commonly in cultivated soil, Nova 
Scotia to the Northwest Territory, south to Florida and 
Texas. Widely distributed in nearly all countries as a 
weed, and includes numerous races, differing principally 
in leaf-form and pubescence. Petty-morel. Duscle. 
Hound’s-berry. July—Oct. 


SOLANACEAE. VoL. III. 


GENUS 6. POTATO FAMILY. 165 


Solanum villdsum (L.) Mill., with coarsely den- 
tate leaves, the pubescence villous and somewhat 
viscid, has been found in ballast about the seaports. 


2. Solanum triflorum Nutt. Cut-leaved 
Nightshade. Fig. 3718. 


Solanum triflorum Nutt. Gen. 1: 128. 1818. 


Annual, sparingly pubescent with simple hairs, 
or glabrous; stem branched, 1°-3° high. Leaves 
pinnatifid or some of them pinnately lobed, acute 
at the apex, petioled, 2-4’ long, the lobes trian- 
gular-lanceolate, acute or obtuse, entire or dentate, 
the sinuses rounded; peduncles lateral, 1-3-flow- 
ered, 2-1’ long; pedicels 3’—7” long, reflexed in 
fruit; calyx-segments lanceolate, shorter than the 
corolla, persistent at the base of the berry; co- 
rolla white, 4-5” broad; anthers obtuse; berries 
green when mature, globose, smooth and glabrous, 
about 5” in diameter. 

On prairies and in waste places, Ontario to Mani- 
toba, Kansas, New Mexico and Arizona. Introduced 
in Missouri. May-—Oct. 


3. Solanum carolinénse L. Horse-Nettle. Sand-Brier. Fig. 3719. 
Solanum carolinensis L. Sp. Pl. 184. 1753. 


Perennial, green, finely stellate-pubescent 
with 4-8-rayed hairs; stem erect, branched, 
1°-4° high, the branches, petioles, midveins 
and sometimes the lateral veins of the leaves 
armed with straight subulate yellow prickles. 
Leaves oblong or ovate, repand, lobed, or pin- 
natifid, 2’-6’ long, the lobes obtuse or acutish; 
petioles 3’—10” long; flowers cymose-racemose, 
appearing terminal, but really lateral, as is 
manifest in fruit; pedicels 3-7” long, re- 
curved in fruit; calyx-lobes lanceolate, acumi- 
nate, about one-half the length of the corolla, 
persistent at the base of the berry; corolla- 
lobes ovate-lanceolate, acute; anthers elon- 
gated; berries orange-yellow, smooth and 
glabrous, 8’—10” in diameter. 


In dry fields and in waste places, southern On- 
*, tario to Vermont, Massachusetts and Florida, west 
: AY to Illinois, Nebraska and Texas. Adventive in its 
} northeastern range. Apple-of-sodom. Radical- 
weed. Bull-nettle. Tread-softly. May—Sept. 


4. Solanum eiaeagnifolium Cay. Silver- 
leaved Nightshade. Fig. 3720. 


S. elaeagnifolium Cav. Icon. 3: 22. pl. 243. 1794. 


Perennial, densely and finely stellate-pubescent, 
silvery-canescent all over; stem branched, 1°-3° 
high, armed with very slender sharp prickles, or 
these wanting. Leaves lanceolate, oblong, or 
linear, petioled, 1-4’ long, 3’-12” wide, mostly 
obtuse at the apex, narrowed or rounded at the 
base, repand-dentate or entire; flowers cymose, 
8-12” broad, violet or blue; peduncles short and 
stout, appearing terminal, but soon evidently lat- 
eral; calyx-lobes lanceolate or linear-Inceolate, 
acute; anthers linear; ovary white-tomentose; 
berries globose, yellow or darker, smooth and 
glabrous, 4’-6” in diameter. 


On dry plains and prairies, Missouri and Kansas 
to Texas and Arizona. Trompillos. May—Sept. 


166 SOLANACEAE. VoL. III. 


5. Solanum Torreyi A. Gray. Torrey’s 
Nightshade. Fig. 3721. 


S. Torreyi A. Gray, Proc. Am. 'Acad. 6: 44. 1862. 


Perennial, hoary with a stellate pubescence of 
8-12-rayed hairs, more or less armed with small 
subulate prickles. Leaves ovate in outline, 3’-6’ 
long, sinuately 5-7-lobed, the lobes entire or undu- 
late, obtuse; cymes appearing terminal, soon evi- 
dently lateral, branched, loosely several-flowered ; 
flowers showy, large; calyx-lobes ovate, abruptly 
long-acuminate, persistent at the base of the 
berry; corolla violet, 1-13’ broad, its lobes ovate, 
acute; berry globose, smooth and glabrous, I’ or 
more in diameter, yellow when ripe. 


On dry plains and prairies, Kansas to Texas. Re- 
ported from Missouri. 


6. Solanum rostratum Dunal. Sand 
Bur. Beaked or Prickly Nightshade. 
Buffalo-bur. Fig. 3722. 


Solanum rostratum Dunal, Sol. 234. pl. 24. 1813. 
S. heterandrum Pursh, Fl. Am. Sept. 156. pi. 7. 
1814. 


Annual, densely stellate-pubescent with 5-8- 
trayed hairs, usually copiously armed with 
yellow subulate prickles; stem erect, branched, 
1°-23° high. Leaves ovate or oval in outline, 
irregularly pinnately 5-7-lobed or 1-2-pinna- 
tifid, 2’-5’ long, petioled, the lobes mostly 
oblong, obtuse; flowers racemose, yellow, about 
1’ broad; racemes lateral; pedicels stout, 3’-6” 
long, erect both in flower and fruit; calyx 
densely prickly, surrounding and wholly en- 
closing the berry, the prickles becoming as 
long as the fruit, or longer; calyx-lobes lan- 
ceolate, acuminate; corolla about 1’ broad, 
slightly irregular, its lobes ovate, acute; sta- 
mens and style declined, the lowest stamen 
longer with an incurved beak; fruit, includ- 
ing its prickles, 1’ in diameter or more. 


+ 4 


On prairies, South Dakota to Texas and Mexico. Occasional in waste places, Ontario to New 


Hampshire, Tennessee and Florida, adventive from the west. Texas-nettle. Prickly potato. May-— 
Sept. The original food of the Colorado beetle. 


7. Solanum citrullifolium Braun. Melon- 
leaved Nightshade. Fig. 3723. 


Solanum citrullifolium Braun, Ind. Sem. Frib. 1849. 


Annual, glandular-pubescent, or a few 4-5-rayed 
hairs on the leaves, copiously armed with slender 
yellow subulate prickles, diffusely branched, 1°-3° 
high. Leaves irregularly bipinnatifid, resembling 
in outline those of the watermelon, 2-6’ long; ra- 
cemes lateral, several-flowered; flowers 1-13’ broad, 
violet; stamens and style declined; lowest anther 
violet, larger than the four other yellow ones; co- 
rolla somewhat irregular, its lobes ovate, acuminate ; 
fruit similar to that of the preceding species. 


In dry soil, Iowa and Kansas to Texas, Mexico and 
New Mexico. Referred. in our first edition, to S. 
heterodoxum Dunal. July—Sept. 


GENUS 6. POTATO FAMILY. 167 


8. Solanum sisymbriifolium Lam. Viscid 
Nightshade. Fig. 3724. 


Solanum sisymbriifolium Lam. Ill. 2: 25. 1793. 


Annual, branched, 2°-4° high, villous-pubescent eI oa 
with long viscid hairs and armed all over with bright % 
yellow prickles. Leaves thin, deeply pinnatifid into 
oblong toothed or sinuate lobes; flowers 14-13’ 
broad, light blue or white; stamens and style nearly 
erect; anthers all equal, yellow; corolla slightly 
irregular, its lobes deltoid or ovate-deltoid, acute or 
obtusish; fruit included ‘in the accrescent calyx 


which has a prickly tube and thinnish lobes. 


In waste places and on ballast, especially about sea- 
ports, from Massachusetts to the Gulf States. Intro- 
duced from tropical America. June—Sept. 


- yS ante 


g- Solanum Dulcamara L. Climbing or Bitter Nightshade. Blue Bindweed. 
Fellenwort. Bittersweet. Dogwood. Fig. 3725. 


Solanum Dulcamara L. Sp. Pl. 185. 1753. 


Perennial, pubescent with simple hairs or gla- 
brate, stem climbing or straggling, somewhat 
woody below, branched, 2°-8° long. Leaves peti- 
oled, ovate or hastate in outline, 2’—4’ long, 1-23’ 
wide, acute or acuminate at the apex, usually 
slightly cordate at the base, some of them entire, 
some with a lobe on one side near the base, some 
deeply 3-lobed or 3-divided, with the terminal 
segment much the largest; cymes compound, lat- 
eral; pedicels slender, articulated at the base, 
spreading or drooping; flowers blue, purple or 
white, 5’-7” broad; calyx-lobes short, oblong, 
obtuse, persistent at the base of the berry; corolla 
deeply 5-cleft, its lobes triangular-lanceolate, acu- 
minate; berry oval or globose, red. 

In waste places or in moist thickets, sometimes 
appearing as if indigenous, Nova Scotia to Minne- 
sota, Washington, New Jersey, Pennsylvania and 
Kansas. May—Sept. Woody nightshade. Poison- 


flower. Poison- or snake-berry. Scarlet berry. Natu- 
ralized from Europe. Native also of Asia. 


Solanum triquetrum Cav., a Texan and Mexican nearly glabrous herb, scarcely climbing, with 
somewhat ridged stems, 3-lobed deltoid-cordate or hastate leaves, lateral few-flowered cymes and 
globose red berries, is reported from Kansas. 

Solanum Melongéna L., the egg-plant, with blue or purplish flowers, and an ovoid or obovoid 
berry up to 6’ long, is occasionally found in waste grounds. It is native of Asia. 
species: Lycium afrum L. 


7, LYCOPERSICON Mill. Gard. Dict. Abr. Ed. 4. 1754. 


Annual, or rarely perennial, coarse branching or feebly climbing herbs, with 1-2-pinnately 
divided leaves, and lateral irregular raceme-like cymes of small yellowish flowers opposite 
the leaves. Calyx 5-parted, or rarely 6-parted, the segments linear or lanceolate. Corolla 
rotate, the tube very short, the limb 5-cleft or rarely 6-cleft, plicate. Stamens 5 (rarely 6), 
inserted on the throat of the corolla; filaments very short; anthers elongated, connate or 
connivent, introrsely longitudinally dehiscent. Ovary 2-3-celled; style simple; stigma small, 
capitate. Berry in the wild plants globose or pyriform, much modified in cultivation, the 
calyx persistent at its base. [Greek, wolf-peach.] 


About 4 species, natives of North America, the following typical. 


168 SOLANACEAE. VoL. IIT. 


1. Lycopersicon Lycopérsicon (L.) 
Karst. Tomato. Love Apple. 
Cherry Tomato. Fig. 3726. 

Solanum Lycopersicum L. Sp. Pl. 185. 1753. 

L. esculentum Mill. Gard. Dict. Ed. 8. 1768. 

Lycopersicum Lycopersicum Karst. Deutsch. Fl. 
966. 1880-83. 


Viscid-pubescent, much branched, 1°-3° 


high, the branches spreading. Leaves peti- 
oled, pinnately divided, 6-18’ long, the seg- 
ments stalked, the larger 7-9, ovate or ovate- 
lanceolate, mostly acute, dentate, lobed or 
again divided, 2’-4’ long, with several or 
numerous smaller, sometimes very small ones 
interspersed; clusters several-flowered; pe- 
duncles 1-3’ long; flowers 5”-8” broad; calyx- 
segments about equalling the corolla; berry 
the well-known tomato or love-apple. 


Escaped from cultivation and _ occasionally 
spontaneous from New York and Pennsylvania 
southward. Jews’ ear. June—Sept. 


8. LYCIUM L. Sp. Pl. ror. 1753. 


Shrubs, or woody vines, often spiny, with small alternate entire leaves, commonly with 
smaller ones fascicled in their axils, and white greenish or purple, axillary or terminal, soli- 
tary or clustered flowers. Calyx campanulate, 3-5-lobed or -toothed, not “enlarged in fruit, 
persistent at the base of the berry. Corolla funnelform, salverform, or campanulate, the tube 
short or slender, the limb 5-lobed (rarely 4-lobed), the lobes obtuse. Stamens 5 (rarely 4), 
exserted, or included; filaments filiform, sometimes dilated at the base; anther-sacs longi- 
tudinally dehiscent. Ovary 2-celled; style filiform; stigma capitate or 2-lobed. Berry glo- 
bose, ovoid, or oblong. [Named from the country Lycia.] 


About 75 species, widely distributed in temperate and warm regions. Besides the following, 
introduced from Europe, some 17 native species occur in the western parts of North America. Type 
species: Lycium afreum L. 


1. Lycium halimifélium Mill. Matrimony Vine. Box-thorn. Fig. 3727. 
Lycium halimifolium Mill. Gard. Dict. Ed. 8, no. 6. 1768. IN 1, 
( 


Lycium Barbarum var. vulgare Ait. f. Hort. Kew. Ed. 2, 
PAO Ge Site Oi ts 
Lycium vulgare Dunal in DC. Prodr.13: Part 1,509. 1852. 


Glabrous, spiny or unarmed; stems slender, climb- 
ing or trailing, branched, 6°-25° long, the branches 
somewhat angled, the spines, when present, slender, 
about 2’ long. Leaves lanceolate, oblong, or spatu- 
late, acute or obtuse at the apex, narrowed into short 
petioles, firm, 2-13’ long, 2’-4” wide; flowers 2-5 
together in the axils, or solitary; peduncles filiform, 
spreading, 6-12” long; calyx-lobes ovate, acute, or 
obtuse, 13” long; corolla funnelform, purplish chang- 
ing to greenish, 4’-6” broad, its lobes ovate-oblong; 
stamens slightly exserted; berry oval, orange-red. 

In thickets and waste places, escaped from gardens, 
Ontario to Virginia, Minnesota and Kansas. Introduced 


from Europe. Bastard jessamine. Jasmine. Jackson-vine. 
May-—Aug. 


9. HYOSCYAMUS [Tourn.] L. Sp. Pl. 179. 1753. 


Erect coarse viscid-pubescent narcotic annual biennial or perennial herbs, with alternate 
mostly lobed or pinnatifid leaves, and large nearly regular flowers, the lower solitary in the 
axils, the upper in a more or less I-sided spike or raceme. Calyx urn-shaped or narrowly 
campanulate, 5-cleft, striate, enlarged and enclosing the capsule in fruit. Corolla funnel- 
form, the limb somewhat oblique, 5-cleft, the lobes more or less unequal, spreading. Stamens 
declined, mostly exserted; filaments filiform; anthers oblong or ovate, their sacs longitudi- 


GENUS 9. POTATO FAMILY. 169 


nally dehiscent. Ovary 2-celled; style slender; stigma capitate. Capsule 2-celled, circum- 
scissile above the middle. [Greek, hog-bean. ] 


About 15 species, natives of the Mediterranean region, the following typical. 


1. Hyoscyamus niger L. Black Hen- 
bane. Hog’s-bean. Fig. 3728. 
Hyoscyamus niger L. Sp. Pl. 179. 1753. — 

Annual or biennial, villous and viscid, of 
an ill odor; stem stout, 1°-23° high. Leaves 
ovate, lanceolate, or oblong in outline, 3’—7’ 
long, acute or acuminate at the apex, sessile, 
or the upper clasping the stem, irregularly 
lobed, cleft or pinnatifid; flowers very short- 
pedicelled, 1-2’ broad; _ calyx-lobes triangular- 
ovate, acute; corolla greenish-yellow, strongly 
reticulated with purple veins, its lobes ovate, 
obtusish; capsule globose-oblong, about 5” 
high. 

In waste places, Nova Scotia to Ontario, New 
York and Michigan. Naturalized from Europe. 
Fetid nightshade. Insane-root. Belene. Chenile. 
Poison-tobacco. June—Sept. 


10. DATURA L. Sp. Pl. 179. 1753. 


Annual or perennial erect tall branching 
narcotic herbs, some tropical species shrubs 
or trees, with alternate petioled entire sinuate- 
dentate or lobed leaves, and large solitary erect 
short-peduncled white purple or violet flow- 
ers. Calyx elongated-tubular or prismatic, its 
apex 5-cleft or spathe-like, in the following species circumscissile near the base which is per- 
sistent and subtends the globose ovoid prickly capsule. Corolla funnelform, the limb plaited, 
5-lobed, the lobes broad, acuminate. Stamens included or little exserted; filaments filiform, 
very long, inserted at or below the middle of the corolla-tube. Ovary 2-celled, or falsely 
4-celled; style filiform; stigma slightly 2-lobed. Capsule 4-valved from the top, or bursting 
irregularly. [The Hindoo name, dhatura.] 

About 12 species, of wide geographic distribution. Type species: Datura Stramonium L. 
The following are introduced weeds. 


Glabrous or very sparingly pubescent; leaves lobed, calyx prismatic. 1. D. Stramonium. 
Finely glandular-pubescent; leaves entire or undulate; calyx tubular. 2. D. Metel. 


1. Datura Stramonium L. Stramonium. 
Jamestown or Jimson-weed. Thorn- 
Apple. Fig. 3729. 


Datura Stramonium L. Sp. Pl. 179. 1753. 
Datura Tatula L. Sp. Pl. Ed. 2, 256. 1762. 


Annual, glabrous or the young parts spar- 
ingly pubescent; stem green to purple, stout, 
1°-5° high. Leaves thin, ovate in outline, acute 
or acuminate at the apex, mostly narrowed at 
the base, 3-8’ long, irregularly sinuate-lobed, 
the lobes acute; petioles 1-4’ long; flowers 
white or violet, about 4’ high, the limb 14’-2’ 
broad; calyx prismatic, less than one-half the 
length of the corolla; capsule ovoid, densely 
prickly,about 2’ high, the lower prickles shorter 
than the upper or all about equal. 


In fields and waste places, Nova Scotia to Flor- 
ida, west to Minnesota and Texas. Naturalized 
from tropical regions. June-Sept. Peru-, mad- 
or devil’s-apple. Devil’s-trumpet. Jamestown-lily. 
Fire-weed. Dewtry. Races differ in color of 
flowers and in length of the prickles on the pods. 


SOLANACEAE. Vor. III. 


2. Datura Métel L. Entire-leaved 
Thorn-Apple. Fig. 3730. 


Datura Metel L. Sp. Pl. 179. 1753. 


Annual, densely and finely glandular-pubes- 
cent; stem stout, much branched, 4°-8° high. 
Leaves broadly ovate, acute at the apex, ine- 
quilateral, rounded or subcordate at the base, 
4-10’ long, entire or merely undulate; petioles 
1-3’ long; flowers white, 6-7’ high, the limb 
3-4 broad; calyx tubular, about one-half as 
long as the corolla; capsule globose or ovoid- 
globose, obtuse, prickly and pubescent, 1-13’ 
in diameter. 


In waste places, escaped from gardens, Rhode 
Island to Florida. Native of tropical America. 
July—Sept. 


11. NICOTIANA [Tourn.] L. Sp. Pl. 180. 1753. 


Annual or perennial viscid-pubescent acrid narcotic herbs or shrubs, with large alternate 
entire or slightly undulate leaves, and rather large white yellow greenish or purplish flowers, 
in terminal, often bracted, racemes or panicles. Calyx tubular-campanulate or ovoid, 5-cleft. 
Corolla funnelform, salverform, or nearly tubular, the tube usually longer than the limb, 
5-lobed, the lobes spreading. Stamens 5, inserted on the tube of the corolla; filaments 
filiform; anthers ovate or oblong, their sacs longitudinally dehiscent. Ovary 2-celled (rarely 
4-celled); style slender; stigma capitate. Capsule 2-valved, or sometimes 4-valved at the 
summit. Seeds very numerous, small. [Named for John Nicot, French ambassodar to 
Portugal, who sent some species to Catherine de Medici, about 1560.] 

About 50 species, mostly natives of America. Besides the following, some ‘to others occur in 
the southern and western United States. Type species: Nicotiana Tabacum L. 


Corolla 1’ long, the tube cylindric; calyx-lobes triangular. ; 1. N. rustica. 
Corolla 4’-6’ long. the tube very slender; calyx-lobes linear or narrowly ; 
lanceolate. 2. N.longiflora. 


1. Nicotiana rustica L. Wild To- 
bacco: - Pigs: 3731. 


Nicotiana rustica L. Sp. Pl. 180. 1753. 


Annual; stem rather slender, 2°-4° high. 
Leaves broadly ovate, thin, entire, slender- 
petioled, 2’-8’ long, 17-6’ wide; petioles 
3’-53’ long; flowers greenish-yellow, about 
1’ long, panicled; pedicels 3-6” long, rather 
stout; calyx-lobes broadly triangular, acute, 
shorter than the tube; corolla-tube cylin- 
dric, somewhat enlarged above, the lobes 
short, obtuse, slightly spreading; capsule 
globose, glabrous, about 5” in diameter, 
2-valved, longer than the calyx. 


In fields and waste places, escaped from 
gardens, Ontario to Minnesota, south to south- 
ern New York and Florida. Cultivated by the 
Indians, its origin unknown. Leaves greenish 
when dry. Indian, Syrian or real-tobacco. 
June-Sept. 


GENUS II. POTATO FAMILY. 171 


2. Nicotiana longiflora Cav. Long-flowered 
Tobacco. Fig. 3732. 
Nicotiana longifiora Cav. Descr. Pl. 106. 1802. 

Annual, minutely rough-puberulent and viscid; stem 
erect, slender, branched, 13°-3° high. Basal leaves 
ovate-lanceolate or broadly oblanceolate, obtuse, 610’ 
long, 1-3’ wide, tapering into slender winged petioles; 
stem-leaves linear or lanceolate, sessile, 2’-4’ long; 
flowers in terminal racemes, short-pedicelled, 4-6’ long; 
calyx oblong, pubescent, its narrow lobes nearly as long 
as the tube; corolla white or purplish, viscid, the tube 
slender, 1’-12” in diameter, expanding above, the lobes 
ovate-lanceolate, acute; capsule oblong. 


Near Harrisburg and Easton, Pa. Escaped from gar- 
dens. Native of South America. Aug.—Oct. 


12, PETUNIA Juss. Ann. Mus. Paris 2: 215. 
pl. 47. - 1803; 


Viscid-pubescent annual or perennial branching herbs, 
with entire leaves, and axillary or terminal solitary white 
violet or purple flowers (in cultivation sometimes variegated). Calyx deeply 5-cleft or 
5-parted, the segments narrow. Corolla funnelform or salverform, the limb plicate, spread- 
ing, slightly irregular. Stamens 5, inserted on the throat of the corolla, 4 of them didy- 
namous, perfect, the fifth smaller or obsolete; filaments slender; anthers ovoid, 2-lobed. 
Disk fleshy. Ovary 2-celled; ovules numerous in each cavity; style filiform; stigma 2-lamel- 
late. Capsule 2-celled, 2-valved, the valves entire. Seeds small, the testa rugose. [Petun, 
an Indian name of tobacco. | 

About 12 species, natives of South America. Type species: Petunia parviflora Juss. 


Corolla white, its tube cylindric. 1. P. axillaris. 
Corolla violet-purple, its tube campanulate. 2. P.violacea. 


1. Petunia axillaris (Lam.) B.S.P. White 
Petunia. Fig. 3733. 
Nicotiana axillaris Lam. Encycl. 4: 480. 1797. 
Petunia nyctaginiflora Juss. Ann. Mus. Paris, 2: 215. 
DL 47 fee LBOgs 
Petunia axillaris B.S.P. Prel. Cat. N. Y. 38. 1888. 
Very viscid; stem stout, about 1° high. Leaves 
ovate to obovate, obtuse or blunt-pointed at the 
apex, sessile, or the lower narrowed into short mar- 
gined petioles, 3-4’ long, 1-2’ wide; peduncles 
slender, 2’-4’ long, often longer than the leaves; 
calyx-segments linear-oblong, obtuse; corolla white, 
its tube cylindric, slightly enlarged above, 1’-1} 
long, 3-4 times as long as the calyx, its limb about 
2’ broad, the lobes rounded. 
In waste places, escaped from gardens, southern New 
York and Pennsylvania. Native of Brazil. July—Sept. 


2. Petunia violacea Lindl. Violet Petunia 


Fig. 3734. 
Petunia violacea Lindl. Bot. Reg. pl. 1626. 1833. 

Similar to the preceding species, but usually 
rather lower, and the stem slender. Leaves ovate 
or obovate, all but the uppermost petioled, mostly 
obtuse, 1-23’ long; peduncles slender, 1-2’ long; 
calyx-segments linear, subacute, or obtuse; co- 
rolla violet-purple, its tube campanulate, 9’—15” 
long, the limb less abruptly spreading, 17-13’ 
broad, the lobes subacute. 

In waste places, escaped from gardens, southern 
New York and Pennsylvania. Native of southern 
Brazil and Paraguay. June-—Sept. 

Petunia parviflora Juss., a prostrate pubescent 
annual, with small linear to spatulate leaves, and a 
funnelform corolla 4”—5” long, has been found on 
ballast about the eastern seaports, from South 
America. 


if SCROPHULARIACEAE. Vou. Wie 


Family 28. SCROPHULARIACEAE Lindl. Nat. Syst. Ed. 2, 288. 1826. 
Ficwort FAMILY. 

Herbs, shrubs or trees, with opposite or alternate exstipulate leaves, and 
perfect mostly complete and irregular flowers (corolla wanting in one species of 
Synthyris). Calyx inferior, persistent, 4—5-toothed, -cleft, or -divided, or some- 
times split on the lower side, or on both sides, the lobes or segments valvate, 
imbricate or distinct in the bud. Corolla gamopetalous, the limb 2-lipped, or 
nearly regular. Stamens 2, 4 or 5, didynamous, or nearly equal, inserted on the 
corolla and alternate with its lobes; anthers 2-celled, the sacs equal, or unequal, 
or sometimes confluent into one. Disk present or obsolete. VPistil 1, entire or 
2-lobed ; ovary superior, 2-celled, or rarely 1-celled; ovules mostly numerous, rarely 
few, anatropous or amphitropous, borne on axile placentae; style slender, simple; 
stigma entire, 2-lobed or 2-lamellate. [Fruit mostly capsular and septicidally or 
loculicidally dehiscent. Seeds mostly numerous, the testa reticulated, pitted, 
striate, ribbed, or nearly smooth; endosperm fleshy; embryo small, straight or 
slightly curved; cotyledons little broader than the radicle. 

About 165 genera and 2700 species, widely distributed, most abundant in temperate regions. 
1. Anther-bearing stamens 5; corolla rotate; leaves alternate. (VeERBASCEAE.) 1. Verbascum. 

2. Anther-bearing stamens 2 or 4; leaves opposite, verticillate or alternate. 


* Corolla spurred, saccate or gibbous on the lower side at the base. (ANTIRRHINEAE.) 


Corolla spurred at the base. 
Leaves palmately 3-5-veined. 2. Cymbalaria. 
Leaves pinnately veined. 
Flowers solitary in the axils. 


Throat of the corolla closed by the palate; leaves broad. 3. Kickxia. 
Throat of the corolla not closed by the palate; leaves narrow. 4. Chaenorrhinum. 
Flowers in terminal racemes. 5. Linaria. 
Corolla saccate or gibous at the base. 6. Antirrhinum, 


** Corolla neither spurred, saccate nor gibbous on the lower side. 
+ Anther-bearing stamens 4, the fifth sterile or rudimentary. (CHELONEAE.) 


Sterile stamen a scale adnate to the upper side of the corolla. 7. Scrophularia. 
Sterile stamen elongated, longer or shorter than the others. 
Corolla tubular, 2-lipped, the lobes of the lower lip flat. 


Sterile stamens shorter than the others; seeds winged. 8. Chelone. 
Sterile stamen about equalling the others; seeds wingless. 9. Pentstemon. 
Corolla 2-cleft, declined ; middle lobe of the lower lip conduplicate. 10. Collinsia. 


++ Stamens 4, all anther-bearing, or 2 sterile, or 2 only. 
a. Stamens 4, all anther-bearing; large Asiatic tree. (PAULOWNIEAE.) 
. 11. Paulownia. 
b. Stamens 4, all anther-bearing; herbs; corolla 2-lipped; stamens not enclosed 
in upper lip of corolla. (MIMULEAE.) 


Calyx prismatic, 5-angled, 5-toothed. 12. Mimulus. 
Calyx 5-parted, not prismatic. 
Calyx-segments equal; leaves pinnatifid in our species. 13. Conobea. 
Calyx-segments unequal, the upper one the largest. 
Corolla nearly regular, about equally 5-lobed. 14. Bramia. 
Corolla manifestly 2-lipped. 
Leaves palmately nerved; flower with an hypogynous disk. 15. Hydrotrida. 
Leaves pinnately veined ; no hypogynous disk. 16. Mecardonia. 


c. Stamens 4, 2 anther-bearing and 2 sterile, or 2 only; corolla obviously 2-lipped. (GRATIOLEAE.) 


Calyx 5-parted ; upper lip of corolla present. 
Sterile filaments short or none. 


Anther-sacs transverse, separated. 17. Gratiola. 
Anther-sacs parallel, contiguous. 18. Sophrononthe. 
Sterile filaments slender, 2-lobed. 19. Ilysanthes. 
Calyx 4-toothed; upper lip of corolla obsolete ; low mud plant. 20. Micranthemum. 


d. Stamens 4, all anther-bearing; corolla nearly regular; flowers on scapes. (LIMOSELLEAE.) 
21. Limosella. 
e. Stamens 2 only (rarely 4 in Synthyris) ; corolla rotate, salverform, tubular, or 
none. (VERONICEAE.) 
Leaves alternate; flowers spicate; corolla 2—3-lobed or none. 22. Synthyris. 
Leaves, at least the lower, opposite or verticillate; corolla 4-lobed. 
Corolla rotate or salverform; capsule obcordate or emarginate, compressed. 23. Veronica. 


Corolla tubular-funnelform ; capsple ovoid, not compressed. 24. Leptandra. 
f. Stamens 4, all anther-bearing; corolla campanulate, salverform or funnelform, scarcely 2-lipped. 
Leaves alternate; flowers in 1-sided racemes. (DIGITALEAE.) 25. Digitalis. 
Leaves, at least the lower, opposite. (BUCHNEREAE.) 

Corolla salverform; flowers in a long spike. 26. Buchnera. 


Corolla campanulate or funnelform. ‘ 
Stamens nearly equal; calyx-lobes as long as the tube. 27. Afzelia. 


Key To GENERA. FIGWORT FAMILY. 173 


Stamens strongly didynamous, unequal ; calyx-teeth shorter than the tube. 


Anthers awned at the base; corolla yellow. 28. Dasystoma. 
Anthers awnless; corolla purple, pink or rarely white. 
Anthers all alike ; lowers pedicelled ; leaves not auricled. 29. Agalinis. 


Anthers of the shorter stamens smaller; leaves auricled at base. 30. Otophylia. 
g. Stamens 4, all anther-bearing, ascending under the upper lip of the corolla. (EUPHRASIEAE.) 


Ovules several or numerous; capsule many-seeded. 
Anther-sacs dissimilar, the inner one pendulous by its apex; leaves mostly alternate. 


Upper lip of the corolla much longer than the lower. 31. Castilleja. 

Upper lip of the corolla scarcely longer than the lower. 32. Orthocarpus. 
Anther-sacs similar and parallel; leaves mostly opposite. 

Calyx 2-bracteolate at the base, oblique, 5-toothed. 33. Schwalbea. 


Calyx not bracteolate, 4—5-toothed, or cleft or split. 
Upper lip of the corolla 2-lobed, its margins recurved; calyx 4-cleft. 34. Euphrasia. 
Margins of the upper lip of the corolla not recurved. 
Calyx scarcely or not at all inflated in fruit; galea entire. 
Calyx 4-toothed or 4-cleft; capsule straight. 


Seeds spreading, numerous. 35. Bartsia. 
Seeds pendulous, few. 36. Odontites. 
Calyx split on the lower side or on both sides; capsule oblique. 
Galea short-beaked or beakless. 37. Pedicularis. 
Galea filiform-beaked. 38. Elephantella. 
Calyx ovoid, much inflated and veiny in fruit. 39. Rhinanthus. 


Ovules only 1 or 2 in each cell of the ovary ; capsule 1-4-seeded ; leaves opposite. 40. Melampyrum. 


1. VERBASCUM [Tourn.] L. Sp. Pl. 177. 1753. 


Biennial or rarely perennial, mostly tall and erect herbs, with alternate dentate pinnatifid 
or entire leaves, and rather large yellow purple red or white flowers, in terminal spikes, 
racemes or panicles. Calyx deeply 5-cleft or 5-parted. Corolla flat-rotate or slightly con- 
cave, 5-lobed, the lobes a little unequal, the upper exterior, at least in the bud. Stamens 5, 
inserted on the base of the corolla, unequal, all anther-bearing; filaments of the 3 upper 
stamens, or of all 5, pilose; anther-sacs confluent into one. Ovules numerous; style dilated 
and flattened at the summit. Capsule globose to oblong, septicidally 2-valved, many-seeded, 
the valves usually 2-cleft at the apex. Seeds rugose, not winged. [The Latin name of the 
great mullen; used by Pliny.] 


About 125 species, natives of the Old World. Besides the following, another is naturalized in 
the western United States. Type species: Verbascum Thapsus L. 


Plants densely woolly; flowers in dense terminal spikes, or spike-like racemes. 
Leaves strongly decurrent on the stem. 
Leaves not decurrent, or but slightly so. 

Leaves white-tomentose beneath; flowers in large terminal panicles. 

Plant glabrous or sparingly glandular; flowers racemose. 


. V. Thapsus. 
V. phlomoides. 
V. Lychnitis. 

. V. Blattaria. 


BwWhH 


1. Verbascum Thapsus L. Great Mullen. Velvet or Mullen Dock. Fig. 3735. 
Verbascum Thapsus L. Sp. Pl. 177. 1753. 


Erect, stout, simple or with some erect 
branches, densely woolly all over with 
branched hairs; stem 2°-7° high, wing- 
angled by the bases of the decurrent leaves. 
Leaves oblong, thick, acute, narrowed at 
the base, dentate or denticulate, 4-12’ long, 
the basal ones borne on margined petioles; 
flowers yellow, 8-12” broad, sessile, nu- 
merous in dense elongated spikes rarely 
branched above; stamens unequal, the three 
upper shorter with white hairy filaments 
and short anthers, the two lower glabrous 
or nearly so with larger anthers; capsule 
about 3” high, slightly longer than the 
woolly calyx. 


In fields and waste places, Nova Scotia to 
South Dakota, California, Florida and Kansas. 
Often a troublesome weed. Naturalized from 
Europe. Native also of Asia. Among some 40 
English names are hedge-, hig- or high-taper. 
Candlewick. Cow’s or bullock’s-lungwort. 
Aaron’s- or Adam’s-rod or -flannel. Feltwort. 
Hare’s-beard. Jacob’s-, Jupiter’s- or Peter’s- 
staff. Ice-leaf. Torches. Flannel-leaf. Old 
man’s-flannel or blanket-leaf. Woolen, 7. e., 
mullen. Shepherd’s-club. Velvet-plant. June— 
Sept. 


174 SCROPHULARIACEAE. VoL. ITI. 


2. Verbascum phlomoides L. Clasp- 
ing-leaved Mullen. Fig. 3736. 


Verbascum phlomoides L.’ Sp. Pl. 1194. 1753. 


Stem rather stout, usually simple, 1°-4° 
high. Leaves oblong to ovate-lanceolate, 
crenate, crenulate, or entire, woolly-tomen- 
tose on both sides, sessile or somewhat 
clasping, or slightly decurrent on the stem, 
or the lower often petioled with truncate 
or subcordate bases; flowers yellow, or 
cream-color, 1’ broad or more, usually in a 
solitary elongated tomentose spike-like ra- 
ceme; pedicels clustered, shorter than the 
calyx; stamens as in V. Thapsus; capsule 
4-5” long, exceeding the tomentose calyx. 


Eastern Massachusetts to Kentucky. Ad- 
ventive or fugitive from Europe or eastern 
Asia. June—Aug. 


{ } 
3. Verbascum Lychnitis L. White Mullen. 


Fig. 3737. 
Verbascum Lychnitis L. Sp. Pl. 177. 1753. 

Stem angled, rather stout, paniculately branched 
above, 2°-42° high, densely covered, as well as 
the lower surfaces of the leaves, with a white 
canescent nearly stellate pubescence. Leaves ob- 
long, ovate or oblong-lanceolate, crenate-dentate, 
2’-7’ long, the upper acute, sessile, but not decur- 
rent on the stem, the lower obtuse or acute at the 
apex and narrowed into margined petioles; flow- 
ers in a large terminal panicle, racemose on its 
branches, white or cream-color, 5-6” broad, 
nearly sessile; pilose hairs of the 3 shorter fila- 
ments white; capsule about 2” high, equalling or 
exceeding the calyx. 

In fields and waste places, Ontario to New Jersey 
and Pennsylvania. Reported from Kansas. Natural- 
ized from Europe. Native also of Asia. June—Sept. 
Its down once used for lighting, hence Lichnitis, 
lamp. 


4. Verbascum Blattaria L. Moth Mullen. Fig. 3738. 
Verbascum Blatteria L. Sp. Pl. 178. 1753. 


Stem erect, strict, slender, terete, glabrous or 
sparingly glandular-pubescent, usually quite simple, 
2°-6° high. Leaves oblong, ovate or lanceolate, 
dentate, laciniate, or pinnatifid, acute or acuminate, 
the upper 3’-23’ long, truncate or cordate-clasping 
at the base, the lower and basal ones sessile or 
somewhat petioled, sometimes 1° long, seldom pres- 

‘ ent at flowering time; raceme 1°-2° long, loose; 
pedicels spreading, 2’-1’ long, bracted at the base; 
corolla yellow or white, about 1’ broad, with brown 
marks on the back; filaments all pilose with violet 
hairs; capsule depressed-globose, 3” in diameter, 
longer than the calyx. 

In fields and waste places, Quebec to Florida, west to 
Oregon and California. Naturalized from Europe. 
Native also of Asia. June-Nov. Said to repel the 
cockroach (Blatta), whence the name Blattaria; fre- 
quented by moths, hence moth-mullen. 

Verbascum virgatum Stokes, a similar Old World 
species, found on Cape Breton Island and naturalized 
in the Pacific States, has fruiting pedicels shorter than 
the capsules. 


GENUS 2. FIGWORT FAMILY. 175 


2. CYMBALARIA Medic. Phil. Bot. 2: 70. 1791. 


Perennial creeping or spreading herbs, with long-petioled, mostly lobed, palmately veined 
leaves, and solitary axillary white to violet flowers. Calyx 5-parted. Corolla irregular, 
2-lipped, short-spurred ; upper lip 2-lobed, lower lip 3-lobed; throat nearly or quite closed by 
the palate. Stamens 4, didynamous, ascending, included, the filaments filiform. Style very 
slender. Capsule dehiscent by 2 terminal 3-toothed pores. Seeds numerous, small. [From 
the Greek for cymbal.] 


About 9 species, natives of the Old World, the following typical. 


1. Cymbalaria Cymbalaria (L.) Wettst. Kenil- 
worth or Coliseum Ivy. Fig. 3739. 


Antirrhinum Cymbalaria L. Sp. Pl. 612. 1753. 

Linaria Cymbalaria Mill. Gard. Dict. Ed. 8, no. 17. 1768. 

Cymbalaria Cymbalaria Wettst. in Engl. & Prantl, Nat. Pf. 
Fam. 4: Abt. 3b, 58. 1891. 


Perennial, glabrous; stem trailing, branched, often 
rooting at the nodes, 3’-12’ long. Leaves slender-petioled, 
reniform-orbicular, palmately 3-5-veined, 3-5-lobed, }’-1’ 
in diameter, the lobes broad and obtuse; petioles usually 
as long as the blade; flowers axillary, solitary, blue or 
lilac, 4-5” long; peduncles slender, recurved, shorter 
than the petioles; calyx-segments lanceolate, acute; 
palate yellowish; capsule globose, several-seeded; seeds 
rugose, wingless. 

Waste places and roadsides, adventive from Europe, On- 
tario to New Jersey and Pennsylvania, and in seaport 
ballast. Other English names are ivy-leaved toadflax, ivy- 
weed. Climbing or roving sailor. Aaron’s-beard. Wander- 
ing jew. Mother-of-thousands. Oxford-weed. Pennywort. 
June—Aug. 


3. KICKXIA Dumort. FI. Belg. 35. | 1827. 
[ELatino1pDEs Wettst. in Engl. & Prantl, Nat. Pfl. Fam. 4: Abt. 3b, 58. 1801. ] 


Mostly annual spreading or creeping herbs, with pinnately veined, short-petioled entire 
toothed or lobed leaves, and solitary axillary white yellow or variegated flowers. Calyx 
5-parted. Corolla irregular, spurred, 2-lipped, the throat closed by the palate. Stamens 4, 
didynamous, included; filaments filiform. Capsule subglobose, or ovoid, opening by I or 2 
terminal slits, pores or valves. Seeds numerous, ovoid, mostly rough or tubercled. [In 
honor of Jean Kickx, 1775-1831, professor in Brussels.] 


About 25 species, natives of the Old World. Type species: Antirrhinum Elatine L. 


Leaves ovate-orbicular, cordate or rounded at the base. 1. K. spuria. 
Leaves hastate. 2. K. Elatine, 


1. Kickxia sptria (L.) Dumort. Round-leaved Toad-Flax. Fig. 3740. 


Antirrhinum spurium L. Sp. Pl. 613. 1753. 

Linaria spuria Mill. Gard. Dict. Ed. 8, no. 15. 1768. 

Kickxia spuria Dumont. FI. Belg. 35. 1827. 

Elatinoides spuria Wettst. in Engl. & Prantl, Nat. Pf. 
Fam. 4: Abt. 3b, 58. 18091. 


Annual, pubescent all over; stems prostrate, branched 
or simple, 3’-2° long. Leaves short-petioled, ovate- 
orbicular, entire, or sometimes dentate, mucronulate at 
the apex, cordate or rounded at the base, }’-1’ in diam- 
eter; petioles 1’-2” long; flowers solitary in the axils, 
small; peduncles filiform, very pubescent, often much 
longer than the leaves; calyx-segments ovate, acute at 
the apex, cordate or rounded at the base, one-half as 
long as the corolla; corolla yellowish with a purple 
upper lip, the spur curved, about as long as the tube; 
capsule subglobose, shorter than the calyx; seeds ru- 
gose, not winged. 


_In waste places and ballast, New York to North Caro- 

lina and Missouri. Adventive from Europe. This and the 

pent called also cancerwort and female-fluellin. June— 
ept. 


176 SCROPHULARIACEAE., Vor. ITT. 


2. Kickxia Elatine (L.) Dumort. Sharp-pointed 
Fluellin or Toad-Flax. Fig. 3741. 

Antirrhinum Elatine L. Sp. Pl. 612. 1753. 
Linaria Elatina Mill. Gard. Dict. Ed. 8, no. 16. 1768. 
Kickxia Elatine Dumont. Fl. Belg. 35. 1827. 
Elatinoides Elatine Wettst. in Engl. & Prantl, Nat, Ea: 

Ram 4 Abt. 3b; 56. s1sone 

Annual, pubescent; stems prostrate, usually branched, 
slender, 6’-2° long. Leaves short-petioled, ovate, 2’—1" 
long, acute or acutish at the apex, triangular, hastate, 
truncate, or subcordate at the base, the basal auricles 
divergent, acute; petioles 1’—3’’ long; flowers solitary 
in the axils, about 3” long; peduncles filiform, gla- 
brous, or somewhat hairy, usually longer than the 
leaves; calyx-segments narrowly lanceolate, acute; co- 
rolla yellowish, purplish beneath, its spur slender, 
straight, declined; capsule subglobose, shorter than the 
calyx; seeds wingless. 

In sandy waste places, Canada (?); Massachusetts to 
Georgia and Missouri. Naturalized from Europe. WNa- 
tive also of Asia. Called also canker-root. June—Sept. 


4. CHAENORRHINUM DC] Lange; Willk. & Lange, Prodr. Fl. Hisp. 2: 
577: ES7O: 


Herbs with alternate, usually entire leaves, and violet, blue or white axillary flowers. 
Calyx 5-parted, the segments narrow. Corolla similar to that of Linaria, but with an open 
throat and a straight upper lip. Stamens 4, didynamous; filaments slender. Style filiform. 
Capsule inequilateral, one carpel larger than the other. Seeds ovoid or cuneate, ribbed. 
[Greek, open nose, referring to the open corolla-throat. ] 


About 20 species, chiefly in the Mediterranean region and Asia. Type species: Antirrhinwmn 
minus L. 


1. Chaenorrhinum minus (L.) Lange. Small Snap-dragon. Fig. 3742. 


Antirrhinum minus L. Sp. Pl. Ed. 2, ee 1763. 

Linaria minor Desf. FI. Atlant. 2: 46. 1800 

Chaenorrhinum minus Lange; Willk. & Lange, Prodr. Fl. Hisp. 
2570 seh o7O: 


Annual, glandular-pubescent all over; stem 5-13’ tall, ae (¢, 
often branched. Leaves linear-spatulate to linear, mostly — v4 
obtuse, 5-15” long, narrowed at the base; flowers shorter a \\ 
than the pedicels, blue or bluish, 23-4” long; calyx-seg- Xg/ Wy ‘ 
ments linear to linear-spatulate, somewhat shorter than the WS \ 
corolla; spur short and stout, much shorter than the body NN AW 
of the corolla; capsule globose-ovoid. ¢ 


Waste grounds and ballast, New Brunswick to New York, 
Pennsylvania and Michigan. Adventive from Europe. Se y 
5. LINARIA [Tourn.] Mill. Gard. Dict. Abr. SA 


Ed. 4. 1754. Sy 


Herbs, some exotic species shrubby, with alternate entire 
dentate or lobed leaves, or the lower and those of sterile 
shoots opposite or verticillate, and yellow white blue purple 
or variegated flowers, in terminal bracted racemes or spikes. 
Calyx 5-parted, the segments imbricated. Corolla irregu- 
lar, spurred at the base, or the spur rarely obsolete, 2-lipped, 
the upper lip erect, 2-lobed, covering the lower in the bud, 
the lower spreading, 3-lobed, its base produced into a palate 
often nearly closing the throat. Stamens 4, didynamous, 
ascending, included; filaments and style filiform. Capsule ovoid or globose, opening by 1 
or more mostly 3-toothed pores or slits below the summit. Seeds numerous, wingless or 
winged, angled or rugose. [Latin, linuwm, flax, which some species resemble. ] 

About 150 species, of wide geographic distribution, most abundant in the Old World. Besides 
the following, another species occurs in Florida. The corolla, especially the terminal one of the 
raceme, occasionally has 5 spurs and is regularly 5-lobed, and is then said to be in the Felon state. 
Type species: Antirrhinum Linaria L 
Flowers yellow, 8’—-15” long; leaves linear; flowers 12”-15” long. 

Leaves alternate. 1. L. Linaria. 

Lower leaves whorled. 2. L. supina. 
Flowers blue to white, 3”—6” long. 

Spur of corolla filiform, curved; native species. 

Spur of the corolla short, conic; European adventive species. 


L. canadensis. 
L. repens. 


Be 


GENUS 5. FIGWORT FAMILY. 177 


1. Linaria Linaria (L.) Karst. Ranstead. Butter-and-Eggs. Fig. 3743. 


Antirrhinum Linaria L. Sp. Pl. 616. 1753. 
Linaria vulgaris Hill, Brit. Herb. 108. 1756. 
Linaria Linaria Karst. Deutsch. Fl. 947. 1880-83. 


Perennial by short rootstocks, pale green and slightly 
glaucous; stems slender, erect, very leafy, glabrous, or 
sparingly glandular-pubescent above, simple or with 
few erect branches, 1°-3° high. Leaves linear, sessile, 
entire, acute at both ends, mostly alternate, 3’-12’ long, 
1-13” wide; flowers densely racemose, light yellow, 
1-1?’ long, the spur of the erect corolla somewhat 
darker, the palate orange-colored; pedicels 2’-4” long, 
nearly erect; calyx-segments oblong, acutish, about 13” 
long; spur subulate, nearly as long as the body of the 
corolla; middle lobe of the lower lip shorter than the 
other two; capsule ovoid, the seeds rugose, winged. 

In fields and waste places, Newfoundland to Oregon, 
Virginia and New Mexico. Naturalized from Europe. 
Native also of Asia. June-Oct. Brideweed. Flaxweed. 
Eggsand bacon. Yellow toad-flax. Impudent lawyer. Jacob’s-ladder. 
Rancid. Wild flax or tobacco. Devil’s flax. Snap-dragon. Devil’s- 
flower. Dead men’s bones. Bread and butter. Continental weed. 


ee Rabbit-flower. Widely distributed in temperate regions as 
a weed. 


Linaria genistaefolia (L.) Mill., found many years ago at the 
northern part of New York Island, and admitted into the first edi- 
tion of this work, has not recently been collected in America. 


2. Linaria supina Desf. Supine Linaria. Fig. 3744. 
Linaria supina Desf. Fl. Atlant. 2: 44. 1800. 


Perennial by short rootstocks, bright green; stems few or 
numerous, decumbent, 4-0’ long, glabrous or slightly glandular- 
pubescent. Leaves linear-spatulate to narrowly linear, the lower 
ones whorled, the upper alternate, mostly 5-15” long; flowers 
few together in short racemes, nearly similar to those of L. 
Linaria, but smaller; capsule globose or ovoid-globose. 


Waste places and ballast, northern Atlantic seaboard. Naturalized 
from Europe. 


3. Linaria canadénsis (L.) Dumort. Blue or Wild Toad-Flax. Fig. 3745. 
Antirrhinum canadense L. Sp. Pl. 618. 1753. 


\ 
Linaria canadensis Dumont, Bot. Cult. 2: 96. 1802. 


. 
Biennial or annual, glabrous, green, sometimes CaN 


fleshy; flowering stems erect or ascending, very i 
slender, simple, or branched, 4’—23° high, the sterile Ps 
shoots spreading or procumbent, very leafy. Leaves = yy 


linear or linear-oblong, 4-15” long, 3-1” wide, 
entire, sessile, those of the sterile shoots, or some 
of them, usually opposite; flowers 3-4” long, in 
slender long racemes; pedicels 2’—3” long, erect and 
appressed in fruit, minutely bracted at the base; 
calyx-segments lanceolate, acute or acuminate, about 
as long as the capsule; spur of the corolla filiform, 
curved, as long as the tube or longer; palate a white 
convex 2-ridged projection; capsule opening by 2 
apical valves, each valve becoming 3-toothed; seeds 
angled, wingless. 

In dry soil, Nova Scotia to Florida, west to Minne- 


sota, Oregon, Texas and California. Also in Central 
and South America. A dwarf form with no corolla is frequent. May-Sept. 


12 


178 SCROPHULARIACEAE. Vor. III. 


4. Linaria répens (L.) Mill. Pale- 
blue Toad-Flax. Fig. 3746. 
Antirrhinum repens L. Sp. Pl. 614. 1753. 


Linaria repens Mill. Gard. Dict. Ed. 8, no. 6. 
1768. 


L. striata DC. Fl. France, 3: 586. 1805. 


Glabrous, perennial by a horizontal or 
creeping rootstock; stem erect, or the base 
decumbent, 8-30’ high, usually branched, 
the branches slender. Leaves linear, en- 
tire, short-petioled or sessile, 2’-2’ long, 
1-2” wide, narrowed to both ends, the 
lower crowded, sometimes whorled, the 
upper more scattered; flowers in slender 
terminal elongating racemes; pedicels 2’— 
5” long; bracts narrowly linear, acute; 
corolla nearly white, but striped with blue 
or purple, about 6” long; spur short, conic; 
capsule subglobose; seeds wrinkled, wing- 
less. 

Newfoundland, and in ballast about the 


Atlantic seaports. Adventive from Europe. 
Summer. 


6. ANTIRRHINUM [Tourn.] L. Sp. Pl. 612. 1753. 


Annual or perennial herbs, with alternate leaves, or the lower and those of sterile shoots 
opposite, and mostly large red purple yellow or white flowers, in terminal racemes, or soli- 
tary in the upper axils. Calyx 5-parted, the segments imbricated. Corolla irregular, gibbous, 
or saccate, but not spurred at the base, 2-lipped, the upper lip erect, 2-lobed, the lower spread- 
ing, 3-lobed, its base produced into a palate nearly or quite closing the throat. Stamens 4, 
didynamous, included; filaments filiform, or dilated at the summit. Style filiform. Capsule 
ovoid or globose, opening by chinks or pores below. the summit. Seeds numerous, oblong, 
truncate, rugose or smooth, not winged. [Greek, nose-like.] 


About 40 species, natives of Europe, Asia and western North America. Besides the following 
introduced species, some 18 others inhabit the western United States. Type species: Antirrhinum 
majus L, 


Flowers 1’-1¥4’ long; calyx-segments ovate, much shorter than the corolla. 1. A. majus. 
Flowers 5”—7” long; calyx-segments linear, as long as the corolla. 2. A. Orontium. 


1. Antirrhinum majus L. Great Snap- 
dragon. Lion’s-mouth. Fig. 3747. 


Antirrhinum majus L. Sp. Pl. 617. 1753. 


Perennial, glabrous below, usually. more or 
less glandular-pubescent above; stem branched 
or simple, 1°-3° high. Leaves lanceolate, linear 
or oblong-lanceolate, entire, short-petioled, acute 
at both ends, rather firm, glabrous, 17-3’ long, 
1-5” wide; flowers racemose, purplish-red (of 
a variety of colors in cultivated forms), 1-13’ 
long; pedicels rather stout, 3’-6” long, erect in 
fruit; calyx-segments oval to ovate, obtuse, 2”— 
3” long; capsule obliquely ovoid, 4-5” high, 
opening by 2 pores just below the summit or 
at length apically 2-valved, much longer than 
the calyx. 

In waste places, sparingly escaped from gardens 
in the Atlantic States. Adventive from Europe. 
Other English names are rabbit’s mouth, bonny rab- 
bits, calf-snout, dragon’s-, tiger’s-, dog’s- or toad’s- 
mouth. Bulldogs. Lion’s-snap. June—Sept. 


GENUs 6. FIGWORT FAMILY. 179 


2. Antirrhinum Ordéntium L. Lesser 
Snapdragon. Fig. 3748. 
Antirrhinum Orontium L. Sp. Pl. 617. 1753. 


Annual, glabrous or pubescent; stem erect, 
simple, or branched, slender, about 1° high. 
Leaves narrowly linear, or the lower linear- 
spatulate, almost sessile, narrowed at both 
ends, 1-2’ long, 1”-2” wide; flowers solitary 
in the upper axils, purple, mostly distant, 
5-7" long; peduncles shorter than the flow- 
ers; calyx-segments linear, somewhat unequal, 
as long as the corolla, elongated in fruit so 
as much to exceed the-pubescent capsule. 

Fields and waste places, Ontario, New Eng- 
land, New York, Vancouver Island and Jamaica. 
Adventive from Europe. Native also of Asia. 
Corn-snapdragon. June—Aug. 


7, SCROPHULARIA [Tourn.] L. 
Sees el Orit 53: 

Perennial strong-smelling herbs, some ex- ete 
otic species shrubby, with mostly opposite 
large leaves, and small purple greenish or yel- $ 
low proterogynous flowers, in terminal pani- 
cled cymes or thyrses. Calyx 5-parted or 5-cleft, the segments or lobes mostly obtuse. 
Corolla irregular, the tube globose to oblong, not gibbous nor spurred at the base, the limb 
5-lobed, the 2 upper lobes longer, erect, the lateral ones ascending, the lower spreading or 
reflexed. Stamens 5, 4 of them anther-bearing and didynamous, declined, mostly included, 
their anther-sacs confluent into one, the fifth sterile, reduced to a scale on the roof of the 
corolla tube. Style filiform; stigma capitate or truncate. Capsule ovoid, septicidally dehis- 
cent. Seeds rugose, not winged. [Named for its repute as a remedy for scrofula.] 


About 120 species, natives of the northern hemisphere, most abundant in southern Europe. 
Besides the following, 2 or 3 others occur in the western United States. Type species: Scrophularia 
nodosa L. 


Corolla dull outside; sterile stamen deep purple. 1. S. marylandica. 
Corolla shining outside; sterile stamen greenish yellow. 
Upper lip of the corolla as long as the tube; panicle-branches sparingly glandular; leaf-blades 


not hastate-incised at the base. 2. S. leporella. 
Upper lip of the corolla much shorter than the tube; panicle-branches densely glandular; leaf- 
blades, especially the lower ones, incised-hastate at the base. 3. S. occidentalis. 


1. Scrophularia marylandica L. Mary- 
land Figwort, Heal-all or Pilewort. 
Fig. 3749. 

Scrophularia marylandica L. Sp. Pl. 619. 1753. 


Scrophularia nodosa var. marylandica A. Gray, 
Syn. Fl. 2: Part 1, 258. 1878. 


Glabrous below, somewhat glandular-pubes- 
cent above; stem slender, 4-angled with grooved 
sides, usually widely branched, erect, 3°-10° 
high. Leaves membranous, slender-petioled, 
usually puberulent beneath, ovate or ovate- 
lanceolate, acuminate at the apex, sharply ser- 
rate, narrowed, truncate or subcordate at the 
base, 3-12’ long; flowers greenish-purple, 
3-4” long, very numerous in the nearly leaf- 
less thyrses; bractlets mostly opposite, pedi- 
cels slender, ascending, 4’-12 long; calyx- 
lobes broadly ovate, obtuse, about the length 
of the tube; corolla green, dull without, brown- 
ish purple and shining within, little contracted 
at the throat, the two lateral lobes slightly 
spreading, the upper lip erect, its lobes short, 
rounded; capsule subglobose, with a slender 
tip; sterile stamen deep purple. 


In woods and thickets, Maine to South Dakota, North Carolina, Georgia and Tennessee. 
Scrofula-plant. Carpenter’s-square. Ascends to 4ooo ft. in North Carolina. July—Sept. 


180 SCROPHULARIACEAE. Vor. IIT. 


Scrophularia neglécta Rydb. differs from S. marylandica in the pubescent leaves, the larger 
corollas and larger capsules; it occurs in the western part of our range. 


2. Scrophularia leporélia Bicknell. Hare 


Figwort. Fig. 3750. 
Scrophularia leporella Bicknell, Bull. Torr. Club 23: 

B17 ee logo: 

Stem puberulent below, viscid-glandular above, 
sharply 4-angled with flat sides, 3°-8° tall, simple, 
or somewhat branched. Leaves short-petioled, 
ovate to lanceolate, acuminate at the apex, mostly 
narrowed at the base, but sometimes subcordate, 
glabrous on both sides when mature, usually in- 
cised-dentate, 2-10’ long; flowers 4”-5” long, in 
elongated narrow thyrses; bractlets mostly alter- 
nate; calyx-lobes ovate, obtuse, or acute; corolla 
contracted at the throat, green to purple and 
shining without, dull within, the two lateral lobes 
erect; lobes of the upper lip often narrowly 
oblong; sterile stamen greenish yellow; capsule 
ovoid-conic. 

- In woods and along roadsides, Vermont to Minne- 


sota, Virginia and Kansas. Ascends to 3500 ft. in 
Virginia. May-July. 


3. Scrophularia occidentalis (Rydb.) Bicknell. Western Figwort. Fig. 3751. 


Scrophularia nodosa occidentalis Rydb. Contr. Nat. Herb. 


22) Iso ital le 
Scrophularia occidentalis Bicknell, Bull. Torr. Club 23: 


315. 1896. 

Similar to S. leporella in habit, but with densely 
glandular panicle-branches. Stem more or less glandu- 
lar; leaf-blades ovate to lanceolate or oblong-lanceolate, 
incised or incised-serrate, at least those of the lower 
leaves hastately incised at the base; corolla 4”-5” long, 
the upper lip much shorter than the tube; capsules 
ovoid, 33’—-4” long. 


In low grounds and thickets, South Dakota to Oklahoma, 
Washington and California. June—Aug. 


8. CHELONE [Tourn] 1 Sp. Pl-Grrs 1753; 


Perennial, mostly glabrous branched or simple herbs, 
with opposite serrate petioled leaves, and large white 
red or purple flowers, in terminal and axillary dense 
spikes. Calyx 5-parted, bracted at the base, the seg- 
ments ovate or lanceolate. Corolla irregular, the tube 
elongated, enlarged above, the limb 2-lipped; upper lip 
concave, emarginate or entire, exterior in the bud; 
lower lip spreading, woolly within, 3-lobed, its lateral lobes sometimes longer than the middle 
one. Stamens 5, included, 4 of them antheriferous, didynamous, the fifth sterile, smaller; 
filaments slender, woolly; anthers woolly, cordate. Style filiform; stigma small, capitate. 
Capsule ovoid, septicidally dehiscent. Seeds numerous, compressed, winged. [Greek, tor- 
toise, the head of which the corolla resembles. ] 


Three species, natives of eastern North America. Type species: Chelone glabra L. 


Corolla white to purplish; bracts not ciliolate. 1. C. glabra. 
Corolla red or rose-purple; bracts ciliolate. 
Leaves oblong or lanceolate. 2. C. obliqua. 


Leaves ovate, acuminate; mountain plant. 3. C.Lyoni. 


GENUs 8. FIGWORT FAMILY. 181 


3 


1. Chelone glabra L. Snake-head. 
Turtle-head. Fig. 3752. es aa, 


Chelone glabra L. Sp. Pl. 611. 1753. \ 


Stem slender, erect, obtusely 4-sided, 
simple or sometimes branched, strict, 1°-3° \ 
high, the branches erect. Leaves linear- 2 


lanceolate to ovate-lanceolate, sharply ser- 
rate with low appressed teeth, acuminate 
at the apex, narrowed at the base, short- 
petioled, 3-6’ long, 3’-12’ wide, the prin- 
cipal veins about Io on each side of the 
midvein; flowers white or faintly pink, 
about 1’ long; bracts glabrous, not ciliolate ; 
calyx-segments ovate-oblong, obtuse; cap- 
sule ovoid, obtuse, about 2’ high, twice as 
long as the calyx. 


Les 


In swamps and along streams, Newfound- 
land to Florida, Manitoba, Alabama and Kan- 
sas. Ascends to 3000 ft. in the Adirondacks. 
Shell-flower. Cod-head. Bitter-herb. Balmony. 
Salt-rheum weed. Turtle-bloom. Fish-mouth. 
povice leaves sometimes broadly oval. July— 

ept. 


2. Chelone obliqua L. Red Turtle-head. 
Fig. 3753. 
Chelone obliqua L. Syst. Ed. 11, no. 4. 1767. 


Stem slender, ascending, 1°—2° high, usually branch- 
ed, the branches spreading or ascending. Leaves ob- 
long, or broadly lanceolate, acuminate at the apex, 
mostly narrowed at the base, petioled, sharply ser- 
rate with somewhat spreading teeth, or laciniate, 2’-6’ 
long, 3-23’ wide; petioles 2-6” long; principal veins 
about 10 on each side; flowers red or rose-purple, 
about 1’ long; bracts and calyx-segments ciliolate 
and usually puberulent; capsule similar to that of 
the preceding species. 


In wet thickets and along streams, Virginia to Illinois, 
south to Florida. July—Sept. 


\ 


3. Chelone Lyoni Pursh. Lyon’s Turtle-head. NY GY '' /; 
Fig. 3754. Ni \\ 


Wo aie : 


Chelone Lyoni Pursh, Fl. Am. Sept. 737. 1814. 


Stem slender, erect or nearly so, simple or branched, 
1°-3° high.’ Leaves ovate, acuminate at the apex, 
rounded, truncate or subcordate at the base; 3’-7 
long, 1-4’ wide, usually slender-petioled, sharply ser- 
rate with divergent teeth, the principal veins 8-10 on 
each side; flowers red or rose-purple, about 1’ long; 


bracts and calyx-segments ciliolate and puberulent. 


In swamps and wet thickets, mountains of Virginia, 
- North Carolina and Tennessee to Georgia. July—Sept. 


182 SCROPHULARIACEAE. Vor. IIT. 


9. PENTSTEMON Mitchell; Soland. in Ait. Hort. Kew. 3: 511. 1789. 


Perennial herbs, mostly branched from the base only, with opposite or rarely verticillate 
leaves, or the upper occasionally alternate, and large, usually showy, blue purple red or white 
flowers, in terminal thyrses, panicles, or racemes. Calyx 5-parted, the segments imbricated 
Corolla irregular, the tube elongated, more or less enlarged above, the limb 2-lipped; upper 
lip 2-lobed; lower lop 3-lobed. Stamens 5, included, 4 of them antheriferous fa didy- 
namous, the fifth sterile, as long as or shorter than the others; anther-sacs divergent or 
connivent. Style filiform; stigma capitate. Capsule ovoid, oblong, or globose, septicidally 
dehiscent. Seeds numerous, angled or even, wingless. [Greek, five stamens. | 


About 100 species, natives of North America and Mexico. Type species: Pentstemon pubes- 
cens Soland. 


* More or less pubescent or glandular, at least the calyx and pedicels. 


Corolla %’-114’ long; leaves entire, serrate, or denticulate. 
Stem pubescent or puberulent nearly or quite to the base. 
Thyrsus open, panicle-like. 
Corolla large, over 10” long; stem hirsute or canescent, often glandular. 
Corolla densely bearded in the throat; outer calyx-segments ovate or ovate-lanceolate. 
: 1. P. hirsutus. 
Corolla scarcely beatded in the throat;. outer calyx-segments lanceolate or linear- 


lanceolate. 2. P. canescens. 
Corolla small, less than 10” long; stem puberulent. 3. P. pallidus. 
Thyrsus narrow, raceme-like or spike-like. 
Corolla-tube abruptly enlarged; sterile filament densely woolly. 4. P. erianthera. 
Corolla-tube gradually enlarged; sterile filament slightly bearded. 5. P. albidus. 


Only the inflorescence, or pedicels, or calyx pubescent. 
Thyrsus open, panicle-like. 
Stem leaves oblong, ovate, or lanceolate. 
Calyx-lobes lanceolate to ovate, much less than 5” long. 
Corolla purplish; tube not gibbous above the point of enlargement. 
6. P. Pentstemon. 
Corolla white or pinkish; tube gibbous above the point of enlargement. 
7. P. Digitalis. 
Calyx-lobes linear-subulate from a narrowly lanceolate base, becoming fully 5” long. 
P. calycosus. 


Stem leaves linear-lanceolate. 9. P. gracilis. 
Thyrsus narrow, interrupted; calyx viscid. 10. P. tubiflorus. 
Corolla 2’ long, the tube much enlarged above; leaves dentate. 11. P. Cobaea. 


** Completely glabrous throughout, mostly glaucous. 


Leaves lanceolate, oblong, ovate, obovate, or orbicular. 


Stem leaves rounded, clasping; flowers 2’ long. 12. P. grandiflorus. 
Stem leaves acute or acuminate; flowers 9”—15” long. 
Corolla 9’—10” long; stem leaves lanceolate. 13. P. acwminatus. 
Corolla 17—-1%’ long; stem leaves mostly oblong. 14. P. glaber. 
Leaves linear or linear-lanceolate; flowers densely thyrsoid. 
Bracts lanceolate, small; flowers 6”—8” long. 15. P. angustifolius. 
Bracts ovate, acuminate, large; flowers 1’ long or more. 16. P. Haydent. 


1. Pentstemon hirsttus (L.) Willd. 
Hairy Beard-tongue. Fig. 3755. 


Chelone hirsuta L. Sp. Pl. 611. 1753. 
P. pubescens Soland. in Ait. Hort. Kew. 3: 360. 17809. 
Pentstemon hirsutus Willd. Sp. Pl. 3: 227. 1801. 


Stem slender, erect, downy nearly or quite to 
the base, 1°-3° high. Leaves puberulent or gla- 
brous, denticulate or the uppermost entire, the 
basal oblong or ovate, obtusish at the apex, 2-43’ 
long, 2’-2’ wide, narrowed into petioles, the up- 
per sessile, lanceolate, mostly acuminate, sessile 
or slightly clasping; inflorescence thyrsoid, rather 
loose, glandular-pubescent; pedicels mostly short ; 
corolla purplish or violet, the tube gradually 
dilated above, 2-grooved on the lower side, about 
1’ long, the throat nearly closed by the villous 
palate at the base of the lower lip; sterile fila- 
ment densely -bearded for about one-half its 
length. 

In dry woods and thickets, Maine to Ontario, 


Florida, Minnesota, Alabama and Missouri. Er- 
roneously recorded from Texas. May-July. 


GENUS 9. . FIGWORT FAMILY. 183 


2. Pentstemon canéscens Britton. Gray 
Beard-tongue. Fig. 3756. 


Pentstemon laevigatus var. canescens Britton, Mem. 
Torr. Club 2: 30. 1890. 
P. canescens Britton, Mem. Torr. Club 5: 291. 1804. 


Densely and finely canescent or puberulent, or 
the leaves sometimes nearly glabrous; stem rather 
stout, 1°-3° high. Leaves denticulate, the lower 
and basal ones oval, obtuse, narrowed into long 
margined petioles, the next I or 2 pairs con- 
tracted below the middle and somewhat fiddle- 
shaped, 3-6’ long, the upper ovate or ovate-lan- 
ceolate, acuminate, clasping; thyrsus elongated, 
open, glandular-pubescent, leafy-bracted below; 
pedicels very short; calyx-segments lanceolate, 
acuminate; corolla purple or nearly white, about 
1’ long, slightly or not at all bearded in the 
throat; sterile filament slightly bearded for about 
one-third its length; capsule ovoid, glabrous, 
.longer than the calyx. 


In dry woods, Virginia, West Virginia and North 
Carolina to western Kentucky. Recorded from Mis- 
souri. May-June. 


3. Pentstemon pallidus Small. Pale Beard- 
tongue. Fig. 3757. 


Pentstemon pallidus Small, Fl. SE. U. S. 1060. 1903. 


Puberulent, or sometimes canescent-puberulent; stem 
3°-9° tall. Basal and lower stem-leaves with oblong 
elliptic or nearly spatulate blades; upper stem-leaves 
few, the blades oblong, lanceolate or linear-lanceolate, 
rather remotely toothed, partly clasping; panicles nar- 
row; calyx-segments ovate, becoming triangular, acute, 
closely puberulent or pubescent during anthesis; corolla 
white or purplish, 9’—10” long, the tube rather grad- 
ually dilated, the throat sparingly bearded; sterile fila- 
ment bearded with very short hairs. 


In sandy soil or swamps, Connecticut and New York to 
Missouri, Florida and Oklahoma. June-July. 


4. Pentstemon erianthera Pursh. Crested 
Beard-tongue. Fig. 3758. 


Pentstemonerianthera Pursh, Fl. Am. Sept.737. 1814. 
Pentstemon cristatus Nutt. Gen. 2: 52. 1818. 


Puberulent below, glandular-villous and_ viscid 
above; stem rather stout, leafy, 6-18’ high. Leaves 
firm, entire or repand, the lower and basal ones 
oblong or spatulate, obtuse or acutish, 24’ long, 
narrowed into margined petioles, the upper sessile 
or somewhat clasping, acute or acuminate; thyrsus 
dense, narrow, leafy-bracted; flowers almost ses- 
sile; calyx-segments ,linear-lanceolate, acuminate, 
villous when young; corolla about 1’ long, rather 
abruptly dilated above, red or purple, its lower 
lip villous within; sterile filament densely long- 
bearded. 


On plains and bluffs, South Dakota to Manitoba, 
Montana, Nebraska and Nevada. May-July. 


; 
[> 


leaves 


as lon 
lated 


short 
On 


on Pentstémon (L.) Brit- 
Smooth Beard-tongue. 
Fig. 3760. 

Chelone Pentstemon L. Sp. Pl. Ed. 2, 850. 1763. 
Pentstemon laevigatus Soland. in Ait. Hort. Kew. 

2: 300. 1789. 
P.Pentstemon Britt. Mem. Torr. Club 5: 291. 1894. 

Glabrous, except the somewhat glandular- 
pubescent inflorescence; stem slender, 2°-3° 
high. Basal and lower leaves oblong or oval, 
obtuse, 3-6’ long, narrowed into margined 
petioles, denticulate; upper leaves sessile or 
slightly clasping, acute, oblong, or lanceolate, 
denticulate; thyrsus open, usually many-flow- 
ered; flowers nearly sessile; calyx-segments 
lanceolate, acute, short; corolla purple or pur- 
plish, 8’—10” long, the tube gradually enlarged 
above, the throat wide open, scarcely or not at 
all bearded, the lobes spreading; sterile fila- 
ment thinly bearded for about one-half its 
length, or more densely bearded above. 


Woods and thickets, Pennsylvania to Florida, 
from cultivation further north and east. May-July. 


6. Pentstem 
ton. 


I 


i/ 


SCROPHULARIACEAE. 


5. Pentstemon albidus Nutt. 


Pentstemon albidus Nutt. Gen. 2: 53. 


boia, Colorado, Nebraska and Texas. 


Vor. IIT. 


White-flowered 


Fig. 3759- 
1818. 


Beard-tongue. 


Stems puberulent below, densely glandular-pubes- 
cent above, rather stout, 6-10’ high. Basal and lower 


spatulate or oblong, obtusish, mostly entire, 


the upper lanceolate or oblong, sessile, denticulate, 
acute or acuminate, 13’-23’ long, 3” 
narrow, raceme-like, leafy-bracted, interrupted; ca- 
lyx-segments lanceolate, acuminate, viscid, one-half 


a” 


wide; thyrsus 


g as the corolla-tube; corolla white or nearly 


so, 8-10” long, funnelform, the tube gradually di- 


upward, the limb nearly equally 5-lobed, the 


lobes spreading; sterile filament slightly bearded with 


hairs. 


plains, Minnesota and South Dakota to Assini- 
June—-Aug. 


= 


Kentucky and Louisiana. 


Si, 


Occasionally escaped 


7. Pentstemon Digitalis (Sweet) Nutt. 
Foxglove Beard-tongue. Fig. 3761. 


Chelone Digitalis Sweet, Brit. Fl. Gard. 2: pl. 120. 


1825-27. 
Pentstemon Digitalis Nutt. Trans. Am. Phil. Soc. 
(IL) §: 18x. 1833-37. ad 


Pentstemon laevigatus var. Digitalis A. Gray, Syn. 
Fl. 2: Part 1, 268, 1878. : 


Glabrous, except the glandular-pubescent in- 
florescence; stem rather stout, <2°-5° high. 
Lower and basal leaves, oblong-or oval, obtuse 
or acutish, entire or’ repand, 2-7’ long, nar- 
rowed into margined petioles; upper leaves 
ovate, lanceolate or ovate-lanceolate, sessile 
and more or less cordate-clasping at the base, 
acuminate, sharply denticulate; thyrsus open, 
many-flowered; pedicels 17-3” long; calyx- 
segments lanceolate, corolla white, 1-17’ long, 
the tube abruptly dilated, the limb moderately 
2-lipped, the throat open; sterile filament 
bearded above. 

In fields and thickets, Maine to New York, IIli- 
nois, Kansas, Virginia and Arkansas. Doubtless 
escaped from cultivation in its northeastern range. 
May-July. 


fe 
= 


\ie 
% 
¥ 


GENUS 9. FIGWORT FAMILY. 185 


8. Pentstemon calycosus Small. Long-sepaled 
Beard-tongue. Fig. 3762. 


Pentstemon calycosus Small, Bull. Torr. Club 25: 470. 
1898. 


Glabrous except a few scattered hairs in the in- 
florescence and lines of puberulence on the inter- 
nodes, deep green. Leaves various, the basal spatulate, 
2’-6’ long, entire or undulate, with winged petioles; 
lower stem-leaves similar to the basal, the upper lan- 
ceolate or ovate-lanceolate, acute or acuminate, finely 
repand or distinctly serrate, rounded at the base and 
clasping; calyx-segments with a lanceolate base and 
a slender tip, often becoming 5” long; corolla purple, 
12’-15”" long; sterile filament sparingly bearded. 


On bluffs, Kentucky to Missouri, Alabama and Arkan- 
sas. May—June. 


g. Pentstemon gracilis Nutt. Slender 
Beard-tongue. Fig. 3763. 
Pentstemon gracilis Nutt. Gen. 2: 52. 1818. 


Glabrous or very nearly so up to the glandular- 
pubescent inflorescence; stem slender, strict, 6-18’ 
high. Basal and lower leaves linear-oblong or 
spatulate, mostly obtuse, denticulate, or entire, 
1-3’ long, narrowed into margined petioles; upper 
leaves sessile, linear-lanceolate or the uppermost 
lanceolate, acuminate, denticulate; thyrsus open, 
several-many-flowered; pedicels 2”-4” long; ca- 
lyx-segments lanceolate, acute or acuminate; co- 
rolla purple, 9-12” long, its tube gradually en- 
larged above, its throat wide open; sterile filament 
bearded for about one-half its length; capsule one- 
third longer than the calyx. 


On moist prairies, Manitoba to Minnesota, Mis- 
souri, Athabasca, Oklahoma and Colorado. May-July. 


10. Pentstemon tubiflorus Nutt. Funnel- 
form Beard-tongue. Fig. 3764. 


Pentstemon tubiflorus Nutt. Trans. Am. Phil. Soc. 

(Il.) 5: 181. 1833-37. 

Glabrous, except the viscid-pubescent calyx and 
pedicels; stem slender, strict, 2°-33° high, leafless 
above. Leaves oblong, ovate, or lanceolate, 1-4’ 
long, obtttsish or acute, the basal narrowed into 
broad marginéd petioles, the upper sessile or clasp- 
ing, entire or merely undulate; thyrsus narrow, 
interrupted; the clusters several-flowered; pedicels 
1-3” long, calyx-segments ovate, acuminate, short, 
striate-nerved; corolla nearly funnelform, white 
or purplish, less than 1’ long, nearly as broad 
when expanded, its tube gradually enlarged, the 
limb nearly equally five-lobed, the lobes spread- 
ing; sterile filament short bearded above; capsule 
ovoid, acute, about twice as long as the calyx. 


S= 


LX 


In moist soil, Missouri, Kansas and Arkansas. 
May-July. 


186 SCROPHULARIACEAE. Vor. III. 


11. Pentstemon Cobaéa Nutt. Cobaea 
Beard-tongue. Fig. 3765. 


Pentstemon Cobaea Nutt. Trans. Am. Phil. Soe. (1I.) 
5: 182. 1833-37. 


Stem stout, densely and finely pubescent below, 
glandular-pubescent above, 1°-2° high. Leaves 
oblong to ovate, firm, 3-5’ long, dentate, the lower 
\) mostly glabrous and narrowed into margined peti- 
Y oles, the upper sessile or cordate-clasping, usually 
pubescent; thyrsus short, several-many-flowered, 
open; flowers about 2’ long; calyx-segments lan- 
ceolate, acuminate, 5’-7” long; corolla purple, 
ptberulent without, glabrous within, its tube nar- 
row up to the top of the calyx, then abruptly 
dilated and campanulate, the limb scarcely 2-lipped, 
the lobes short, rounded, spreading; sterile fila- 
ment sparingly bearded; capsule ovoid, acute, pu- 
bescent, reticulate-veined, as long as the calyx. 


On dry prairies, Missouri and Kansas to Texas. 
Recorded from Ohio. May-July. 


12. Pentstemon grandiflorus Nutt. 
Large-flowered Beard-tongue. 
Fig. 3766. 
P. grandiflorus Nutt. in Fras. Cat. 1813. 


Glabrous and somewhat glaucous ; stem stout, 
2°-4° high. Leaves all entire and obtuse, the 
basal ones obovate, narrowed into broad peti- 
oles, those of the lower part of the stem ses- 
sile, oblong or oval, 1’-23’ long, the upper 
nearly orbicular, cordate-clasping, shorter; 
thyrsus open, leafy-bracted, the bracts orbicu- 
lar, cordate; pedicels 2-6” long; flowers 
nearly 2’ long; calyx-segments lanceolate, 
acute, 3-4” long; corolla lavender-blue, the 
tube rather abruptly dilated above the calyx, 
the limb somewhat 2-lipped; sterile filament 
incurved, villous and capitate at the summit; 
capsule acute, 8’-10” high, three times as long 
as the calyx. 

On prairies, Illinois to Minnesota, North Da- 
kota, Kansas and Colorado. Locally introduced 
eastward. June—Aug. 


13. Pentstemon acuminatus Dougl. Sharp- 
leaved Beard-tongue. Fig. 3767. 


Pentstemon acuminatus Dougl.; Lindl. Bot. Reg. pl. 
7285. “1829. 


Glabrous and glaucous; stem rather stout, strict, 
6’-2° high, leafy. Leaves firm, entire, the lower 
and basal ones oblong or spatulate, obtuse or 
acute, narrowed into petioles, the upper sessile or 
clasping, lanceolate or ovate-lanceolate, 2a 
long; thyrsus narrow, sometimes I-sided, usually 
leafy-bracted below; pedicels becoming 47-8” 
long in fruit; calyx-segments lanceolate, acute, 
2-3” long; corolla blue, 9-10” ‘long, its tube 
rather gradually dilated, the limb 2-lipped; sterile 
filament bearded along the dilated summit; cap- 
sule acute, twice as long as the calyx. 


In dry soil, Minnesota to Nebraska, Texas, Mani- 
toba, Alberta, Oregon and New Mexico. St. Joseph’s- 
wand. May—Aug. 


GENUS 9. FIGWORT FAMILY. 187 


14. Pentstemon glaber Pursh. Large 
Smooth Beard-tongue. Fig. 3768. 


Pentstemon glaber Pursh, Fl. Am. Sept. 738. 1814. 


Glabrous, somewhat glaucous; stem ascending 
or erect, rather stout, leafy, 1°-2° high. Leaves 
entire, firm, the basal and lower ones narrowed 
into petioles, the middle ones oblong or oblong- 
lanceolate, acuminate, sessile, the upper lanceo- 
late, acuminate, sessile, scarcely clasping; thyrsus 
narrow, elongated, densely many-flowered; pedi- 
cels 3-7” lotig in fruit; calyx-segments ovate- 
lanceolate, scarious-margined, abruptly acumi- 
nate, 3-4” long,their margins commonly eroded; 
corolla blue or purple, 1-13’ long, rather abruptly 
expanded above the calyx, the limb somewhat 
2-lipped, the lobes rounded; sterile filament 
bearded at the slightly enlarged summit; capsule 
narrowly ovoid, acute, about twice as long as 
the calyx. 


Tn moist soil, South Dakota to Nebraska, Wyo- 
ming and Arizona. Far western plants previously 
referred to this species prove to be distinct. May—Aug. 

KTP 
15. Pentstemon angustifdlius Pursh. 


Pale-blue Beard-tongue. Fig. 37609. 


P. angustifolius Pursh, Fl. Am. Sept. 738. 1814. 
Pentstemon coeruleus Nutt. Gen. 2: 52. 1818. 


Glabrous and glaucous; stem slender, erect, 
leafy, 6-15’ high. Leaves all linear or linear- 
lanceolate, entire, the lower narrowed into 
petioles, obtusish at the apex, the upper ses- 
sile, acute, 13-23’ long, 14”-2” wide; thyrsus 
narrow, spike-like, mostly dense; bracts lan- 
ceolate, acuminate; pedicels very short; ca- 
lyx-segments  linear-lanceolate, acuminate, 


| 2-3” long; corolla blue, or nearly white, 
( 8’’-r0” long, the tube gradually enlarged, the 
limb somewhat 2-lipped; sterile filament 
bearded at the summit. 
In dry soil, western Nebraska to North Da- 
kota and Montana. May-July. 


16. Pentstemon Haydeni S. Wats. Hay- 
den’s Beard-tongue. Fig. 3770. 


P. Haydeni S. Wats. Bot. Gaz. 16: 311. 1891. 


Glabrous, not at all glaucous or slightly so; 
stem decumbent, simple or branched, leafy, 1°-2° 
high. Leaves linear or elongated-lanceolate, en- 
tire, sessile and slightly clasping, acute, acumi- 
nate, or the lowest obtusish at the apex, 2’—5’ 
long, 1-5” wide; thyrsus narrow, dense; bracts 
ovate or ovate-lanceolate, large, cordate-clasp- 
ing, acute, or acuminate; fruiting pedicels 2”—3” 
long; calyx-segments lanceolate, striate-nerved, 
acuminate, 37-5” long; corolla blue, 1’ long or 
more, the tube broadly dilated above the calyx, 
the limb nearly equally 5-lobed; capsule acute, 
twice as long as the calyx. 


In moist soil, Nebraska, Kansas and Wyoming. 


188 SCROPHULARIACEAE. Vou. TH 


10. COLLINSIA Nutt. Journ. Acad. Phil. 1: 190, pl. 0: © 1817. 


Winter-annual or biennial herbs, with opposite or verticillate leaves, and blue pink white 
or variegated flowers, verticillate, or solitary inthe axils. Calyx campanulate, 5-cleft. Corolla 
irregular, the tube short, the limb 2-lipped; upper lip 2-cleft, the lobes erect or recurved; 
lower lip larger, 3-lobed, the lateral lobes spreading or drooping, flat, the middle one condu- 
plicate, keel-like, enclosing the 4 declined stamens and the filiform style. Stamens didy- 
namous. Corolla with a gland on the upper side of the tube near the base. Filaments fili- 
form; anther-sacs confluent at the apex. Stigma small, capitate or 2-lobed. Capsule ovoid 
or globose, septicidally 2-valved, the valves 2-cleft. Seeds few, large, peltate, concave on 
the inner side. [Named for Zaccheus Collins, botanist, of Philadelphia, 1764-1831. ] 


About 20 species, natives of North America. Type species: Collinsia verna Nutt. 


Corolla 5”—8” long, the throat shorter than the limb. 
Leaves, at least the lower, ovate or oblong; corolla-lobes notched. 1. C. verna. 
Leaves lanceolate; corolla-lobes obcordate. 2. C. violacea. 
Corolla 2”—3” long, the throat longer than the limb. 3. C. tenella. 


1. Collinsia vérna Nutt. Blue-eyed Mary. Innocence. Broad-leaved Collinsia. 
Fig. 3771. 
C. verna Nutt. Journ. Acad. Phil. 1: 190. pl. 9. 1817. 


Glabrous or puberulent; stem slender, weak, 
6’-2° long, simple or branched. Leaves thin, oppo- 
site, the lower broadly ovate or orbicular, obtuse 
at the apex, rounded, narrowed or subcordate at 
the base, crenate or entire, slender-petioled; mid- 
dle leaves sessile or cordate-clasping, ovate or 
oblong, obtuse, dentate, 1-2’ long, floral leaves 
ovate to spatulate, mostly acute, dentate or entire; 
upper whorls 4-6-flowered; peduncles 3’-1’ long; 
corolla 6-8” long, its throat equalling or shorter 
than the calyx, its lower lip blue, the upper purple 
or nearly white, the lobes emarginate or truncate; 
capsule globose, 23”—3” in diameter, shorter than 
the linear calyx-lobes. 

In moist woods and thickets, Ontario and western 


New York to Wisconsin, south to Pennsylvania, 
Kentucky and Kansas. April-June. 


The California Collinsia bicolor Benth., which 
differs from this by short-peduncled flowers, is re- 
corded as found introduced in Illinois. 


2. Collinsia violacea Nutt. Violet or Nar- 
row-leaved Collinsia. Fig. 3772. 


Collinsia violacea Nutt. Trans. Am. Phil. Soc. (II.) 
5: 179. 1833-37. 


Similar to the preceding species, stem slender, 
erect, usually branched, 6-15’ high. Leaves lan- 
ceolate or oblong-lanceolate, rather thick, entire or 
denticulate, obtuse or obtusish, the lower opposite, 
petioled, the middle similar, sessile, 1-2’ long, 3-5” 
wide, the floral linear or linear-lanceolate, opposite 
or verticillate; upper whorls 2-5-flowered: corolla 
5-6” long, violet, its lobes obcordate or emargi- 
nate; capsule globose, about 2” in diameter, shorter 
than the lanceolate acute calyx-lobes. 


Rich soil, Missouri, Kansas and Arkansas to Texas. 
April—May. 


GENUS I0. FIGWORT FAMILY. 


3. Collinsia tenélla (Pursh) Piper. 
Small-flowered Collinsia. Fig. 3773. 


Antirrhinum tenellum Pursh, Fl. Am. Sept. 

421. 1814. 

Collinsia parviflora Dougl.; Lindl. Bot. Reg. 

pl. 1082. 1827. j 
C. tenella Piper, Contr. Nat. Herb. 11: 496. 

1906. 

Puberulent, at length diffusely branched; 
stems very slender, 3-15’ long. Leaves 
oblong or lanceolate mostly obtuse at the 
apex and narrowed at the base, 3’-1’ long, 
entire, or sparingly toothed, the lower op- 
posite, sometimes broader, petioled, the 
floral sessile, opposite or verticillate; upper 
whorls 2-6-flowered; pedicels commonly 
longer than the flowers; corolla 3-4” long, 
about twice as long as the calyx, blue or 
whitish, the throat longer than the limb; 
capsule globose, 1-13” in diameter, little 
shorter than the lanceolate calyx-lobes. 

In moist places, Ontario to British Colum- 
bia, Michigan, Colorado, Arizona and Utah. 
April-June. : 


11. PAULOWNIA Sieb. & Zuce. Fl. Jap. 1: 25. pl. 10. 1835. 


A large tree, with the aspect of Catalpa, with broad opposite entire or 3-lobed, petioled 
pubescent leaves, and large violet flowers in terminal panicles. Calyx deeply 5-cleft, the 
lobes short. Corolla irregular, the tube elongated, enlarged above, the 5 lobes spreading, 
somewhat unequal. Stamens 4, didynamous, included; anther-sacs divaricate. Style slender, 
slightly thickened toward the summit, stigmatic on the inner side. Capsule coriaceous, ovoid, 
acute, loculicidally dehiscent. Seeds numerous, striate, winged. Flowers expanding before 

A the leaves appear. [Named for Anna Paulowna, daughter 
of the Czar Paul I.] 


A monotypic Japanese genus. 


1. Paulownia tomentosa (Thunb.) Baill. Pau- 
lownia. Fig. 3774. 

Bignonia tomentosa Thunb. Fl. Jap. 252. 1784. 

Paulownia imperialis Sieb. & Zucc. Fl. Jap. 1: 27. 1835. 

Paulownia tomentosa Baill. Hist. Pl. 9: 434. 1888. 

A tree with thin flaky bark, reaching a maximum 
height of about 70° and a trunk diameter of 4°, the 
branches stout, spreading. Leaves broadly ovate, 6-15’ 
long, 4-8’ wide, long-petioled, canescent on both sides 
when young, glabrate above when old, the petioles 
terete; flowers about 23’ long, numerous in large erect 
terminal panicles; pedicels stout, densely tomentose; 
calyx 5-lobed, the lobes thick, tomentose; corolla slightly 
irregular, puberulent without; capsule 2’ high, 1’ in 
diameter. 


Escaped from cultivation, southern New York and New 
Jersey to Georgia. May-July. 


12, MIMULUS L. Sp. Pl. 634. 1753. 


Erect or decumbent herbs, with opposite mostly dentate leaves. Flowers axillary, soli- 
tary, peduncled, pink, violet, or yellow, usually showy. Calyx prismatic, 5-angled, 5-toothed, 
the upper tooth usually the largest. Corolla irregular, its tube cylindric with a pair of ridges 
on the lower side within, its limb 2-lipped; upper lip erect or reflexed, 2-lobed; lower lip 
spreading, 3-lobed, the lobes rounded. Stamens 4, didynamous, inserted on the corolla-tube; 
anther-sacs divergent, or sometimes confluent at the summit. Style filiform; stigma 2-lamel- 
late. Capsule oblong or linear, loculicidally dehiscent, many-seeded, enclosed by the calyx. 
[Diminutive of mimus, a mimic actor. ] 


About 40 species, natives of America. Besides the following, some 30 others occur in the 
western United States and British Columbia. Type species: Mimulus ringens L 
Corolla violet, or rarely white; eastern species. 
Leaves sessile, clasping ; peduncles longer than the calyx. 1. M. ringens. 
Leaves petioled ; peduncles shorter than the calyx. 2. M. alatus. 
Corolla yellow ; western; two species adventive in the East. 
Plants glabrous or glabrate. 


190 SCROPHULARIACEAE. Vor. IIT. 


Erect; branches spreading ; leaves ovate; flowers 1’ long. 3. M. Langsdor ffi. 
Diffuse ; leaves nearly orbicular; flowers about 6” long. 4. M. Geyeri. 
Plant villous and viscid, diffuse, musk-scented. 5. M. moschatus. 


1. Mimulus ringens L. Square-stemmed 
Monkey-flower. Fig. 3775. 
Mimulus ringens L. Sp. Pl. 634. 1753. 


Glabrous, perennial by rootstocks ; stem erect, 4-sided 
or somewhat 4-winged, usually much branched, 1°-3° 
high. Leaves oblong, lanceolate, or oblong-lanceolate, 
pinnately veined, acuminate or acute at apex, serrate, 
auriculate-clasping at the base, or the lower merely 
sessile, 2’-4’ long, 4’-1’ wide; peduncles slender, 1’—2’ 
long in fruit, 2-4 times as long as the calyx; calyx- 
teeth lanceolate-subulate; corolla violet, rarely white, 
about I’ long, the throat narrow, exceeding the calyx; 
base of lower lip puberulent within; fruiting calyx 
oblong, 6’-8” long; seeds oblong, minute, reticulated. 

In swamps and along streams, Nova Scotia to Vir- 


ginia, Tennessee, Manitoba, Nebraska and Texas. As- 
cends to 3000 ft. in Virginia. June—Sept. 


i 


2. Mimulus alatus Soland. Sharp-winged 
Monkey-flower. Fig. 3776. 


Mimulus alatus Soland. in Ait. Hort. Kew. 2: 361. 

17809. 

Similar to the preceding species, glabrous; stem 
sharply 4-angled, the angles more or less winged. 
Leaves ovate, ovate-lanceolate, or oblong, acute 
or acuminate at the apex, dentate-serrate, nar- 
rowed at the base, petioled, 2’-5’ long, 9”-18” 
wide; petioles 3-1’ long, narrowly margined; 
peduncles stout, shorter than the calyx; corolla 
violet, 1’ long; calyx-teeth short, broad, abruptly 
mucronulate; seeds smooth. 


In swamps, Ontario to Connecticut, Illinois, Kan- 
sas, Georgia and Texas. June—Sept. 


3. Mimulus Langsdorfii Donn. 
Langsdorff’s Yellow Monkey- 
flower. Fig. 3777. 

Mimulus Langsdorffii Donn; Sims. Bot. Mag. 

bl. T50r. TeT2: 


Mimulus guttatus DC. Cat. Hort. Monsp. 127. 
1818. 


Perennial by stolons, glabrous or puberu- 
lent; stem rather stout, simple or branched; 
branches spreading. Leaves ovate to obo- 
vate, dentate or denticulate, obtuse at the 
apex, rounded, narrowed or cordate at the 
base, 17-3’ long, the lower short-petioled, 
the upper smaller, sessile or clasping; pe- 
duncles shorter than or equalling the flow- 
ers; calyx oblique; corolla yellow, 1-2’ 
long, the lower lip bearded at the base, 
often blotched with red or purple; fruiting 
calyx about 2’ long; seeds longitudinally 
striate. 


In wet meadows, Norfolk, Conn., and south- 
ern New York. Introduced from California. 
Summer. 


GENUS I0, FIGWORT FAMILY. 


4. Mimulus Géyeri Torr. Geyer’s Yellow 
Monkey-flower. Fig. 3778. 
M. Geyeri Torr. in Nicollet, Rep. Up. Miss.157. 1843. 
Mimulus Jamesit T. & G.; Benth. in DC. Prodr. 1o: 
371. 1846. 
M. glabratus var. Jamesii A. Gray, Syn. Fl. N. A. Ed. 
22> A477. 2 1O80 
Perennial by stolons, glabrous or nearly so; 
stems slender, creeping, diffusely branched, root- 
ing at the nodes, 6-18 long. Leaves broadly 
ovate, orbicular or broader, very obtuse, denticu- 
fate or entire, membranous, truncate, subcordate, 
or rarely narrowed at the base, palmately veined, 
short-petioled or the upper sessile, 4-1’ in diam- 
eter; peduncles slender, longer than the oblique 
calyx in fruit, about as long as the leaves; calyx 
scarcely toothed, 3-4” long at maturity; corolla 
yellow, 4”-6” long, the lower lip bearded at the 
base within, the throat broad; seeds nearly smooth. 
In brooks and swamps, Ontario to Illinois, Ne- 
braska, Wyoming and Colorado. June—Sept. 


5. Mimulus moschatus Dougl. Musk-flower. 
ahd Musk-plant. Fig. 3779. 
ELLY M. moschatus Dougl.; Lindl. Bot. Reg. pl. rr78. 1827. 
Perennial, villous-pubescent, viscid, musk-scented; 
stems creeping and ascending, branched, slender, 6-12’ 
long. Leaves ovate or oblong, short-petioled, acute or 
obtuse at the apex, denticulate, rounded or subcordate 
at the base, 1-2’ long, 5-12” wide; peduncles slender, 
longer than the calyx; flowers 1’ long or less; calyx- 
teeth lanceolate, acuminate, nearly equal; corolla yel- 
low, 2-3 times as long as the calyx. 
In wet places, Newfoundland to Pennsylvania, Ontario 


and Michigan; apparently introduced from western North 
America, where it is widely distributed. June-Sept. 
Mazus japonicus (Thunb.) Kuntze, a low herb with 
mostly basal, slender-petioled obovate repand leaves and 
racemose violet flowers, the campanulate calyx not angled, 
has been found at Washington, D. C., and abundantly 
about New Orleans, La. It is native of eastern Asia. 


rm CONOBEA. Aubl, Pl. Guian. 2: 639. pl. 2502, 1775. 

Herbs, with opposite pinnately parted pinnatifid incised or serrate leaves, and small blue 
or white peduncled flowers, solitary or two together in the axils. Calyx 5-parted, the seg- 
ments narrow, equal. Corolla irregular, the tube cylindric, the limb 2-lipped; upper lip emargi- 
nate or 2-lobed; lower lip 3-lobed. Stamens 4, didynamous, ascending, included; filaments 
filiform; anther-sacs parallel, not confluent. Style incurved at the summit; stigma 2-lamel- 
late. Capsule globose, oblong or linear, septicidally dehiscent, the valves entire or 2-cleft. 
Seeds numerous, oblong, striate. [Guiana name.] 


About 8 species, natives of America. Besides the fol- 
lowing, another occurs in the southwestern United 
States. Type species: Conobea aquatica Aubl. 


1. Conobea multifida (Michx.) Benth. 
Conobea. Fig. 3780. 
Capraria multifida Michx. Fl. Bor. Am. 2: 22. pl. 35. 
8 


1803. 
Conobea multifida Benth. in DC. Prodr. 10: 391. 1846. 


Annual, finely viscid-pubescent; stem at length 
diffusely branched, 4’-8’ high, very leafy. Leaves 
petioled, 4’-1’ long, pinnately parted into 3-7 linear 
or linear-oblong obtuse entire or incised segments; 
flowers greenish-white, 2-23” long, mostly solitary 
in the opposite axils, about as long as their filiform 
peduncles; calyx-segments linear-subulate, slightly 
shorter than the corolla; capsule narrowly ovoid, 
glabrous, about equalling the calyx. 

Along streams and rivers, Ontario to Ohio, Iowa, 
Kansas, Kentucky, Alabama and Texas. Introduced 
along the Delaware below Philadelphia. June—Sept. 


192 SCROPHULARIACEAE. Vor. III. 


14. BRAMIA Lam. Encycl. 1: 450. 1783. 


[Monniera P. Br. Civ. & Nat. Hist. Jam. 260. pl. 28. f. 3. Hyponym. 1755. Not L. 1759.] 
[Herpestis Gaertn. Fruct. & Sem. 3: 186. pl. 214. 1805.] 


Diffuse or prostrate herbs, with opposite mostly entire obtuse leaves, and small peduncled 
flowers, mostly solitary in the axils. Calyx subtended by 2 bracts, 5-parted, the upper seg- 
ment the broadest. Corolla blue or white, nearly regular, the tube cylindric, the limb nearly 
equally 5-lobed. Stamens 4, didynamous, included. Style slender; stigma capitate, or 2- 
lobed. Capsule globose or ovoid, septicidally dehiscent. Seeds numerous. [From Brami, 
a Malabar name.] . 


About 20 species, natives of warm and tropical regions. Type species: Bramia indica Lam. 


Leaves spatulate or cuneate ; capsules acuminate at the apex. 1. B. Monniera. 
Leaves obovate or orbicular-obovate; capsules blunt at the apex. 2. B. rotundifolia. 


1. Bramia Monniéra (L.) Drake. Monnier’s Hedge-Hyssop. Fig. 3781. 


Gratiola Monniera L. Cent. Pl. 2. 1756. 
Herpestis Monniera H.B.K. Nov. Gen. 2: 366. 1817. 
M. Monniera Britton, Mem. Torr. Club 5: 292. 1894. 
Bacopa Monniera Wettst. in Engler & Prantl, Nat. 

Baa) T8or- 
Bramia Monniera Drake, FI. Polyn. France. 142. 1892. 

Perennial, glabrous, fleshy; stem creeping, root- 
ing at the nodes, branched, 6-18’ long. Leaves 
spatulate or cuneate-obcordate, sessile, rounded 
at the apex, entire, or sparingly denticulate, 3’— 
10” long, 1-24” wide; peduncles mainly in alter- 
nate axils, slender, 2-bracteolate at the summit, in 
fruit longer than the leaves; flowers pale blue, 5” 
long; upper calyx-segment ovate, acute; corolla 
obscurely 2-lipped; stamens nearly equal; stigma 
slightly 2-lobed; capsule ovoid, acute, shorter 
than the calyx. 

On shores, near the coast, Maryland to Florida, 
Texas and Mexico. Widely distributed in tropical 


regions of the Old World and the New. June—Oct. 
Herb-grace. Water-hyssop. 


2. Bramia rotundifolia (Michx.) Britton. Round-leaved Hedge-Hyssop. 
Fig. 3782. 

M. rotundifolia Michx. Fl. Bor. Am. 2: 22. 1803. 

i ie rotundifolia Pursh, Fl. Am. Sept. 418. 

Biot rotundifolia Wettst. in Engler & Prantl, 

Nut. Pfl. 42°: 76. 1891. 

Perennial by stolons, succulent; stems creep- 
ing and spreading, branched or simple, villous- 
pubescent, 1°-2° long. Leaves obovate or or- 
bicular, palmately veined, entire, or slightly 
undulate, narrowed to a sessile or clasping 
base, 2-1’ broad, glabrous, not punctate; pe- 
duncles stout, shorter than the leaves, solitary 
or 2 together in the axils, longer than the 
flowers; flowers blue, 3-4” long; upper calyx- 
segment oval; corolla 2-lipped, longer than 


the calyx; stamens approximate in pairs; Z 
stigma 2-lobed; disk obsolete; capsule oblong, j \ 
obtuse, 14” high, at length 4-valved, shorter \ 
than the calyx. 

On muddy shores, Illinois to South Dakota, cy 


Tennessee and Texas. June—Sept. 


GENUS I5. FIGWORT FAMILY. 193 


15. HYDROTRIDA Willd. 


Perennial aromatic succulent creeping herbs, usually pubescent. Stems terete. Leaves 
opposite; blades broadest below the middle, palmately nerved, entire or shallowly toothed, 
punctate, partly clasping. Flowers solitary on short axillary peduncles. Calyx subtended 
by 2 small bractlets. Sepals nearly distinct, the outer ones cordate, the upper one broadest. 
Corolla blue or white, manifestly 2-lipped, the upper lip merely notched. Hypogynous disk 
present. Stamens 4, included; filaments adnate to near the throat of the corolla-tube. Cap- 
sule ovoid to conic, septicidally dehiscent, the valves cleft. Seeds numerous. [Name 
unexplained. ] 


About 8 species, of the southeastern United States, the West Indies and South America. Type 
species: Herpestis obovata Poepp. (See Linnaea 5: 107.) 


1. Hydrotrida caroliniana (Walt.) Small. 
Blue Hedge-Hyssop. Fig. 3783. 
Obolaria caroliniana Walt. Fl. Car. 166. 1788. 
Monniera amplexicaulis Michx. Fl. Bor. Am.2: 22. 1803. 
Herpestis amplexiculis Pursh, Fl. Am. Sept. 418. 1814. 


M. caroliniana Kuntze, Rev. Gen. Pl. 463. 1891. 
Septilia caroliniana Small, Fl. SE. U. S. 1064. 1903. 


Perennial by stolons, more or less pubescent, fleshy ; 
stems creeping and ascending, rooting at the lower 
nodes, 6—2° long, simple, or sparingly branched, 
leafy. Leaves ovate to oval, sessile and clasping by 
a subcordate base, obtuse at the apex, parallel-veined, 
punctate, entire, the margins ciliolate or naked; pe- 
duncles shorter than the leaves; usually shorter 
than the calyx; upper calyx-segment ovate, cordate; 
flowers blue, 4’-5” long, ephemeral; disk 10-12- 
toothed; stamens approximate in pairs. 


In wet pine barrens, New Jersey to Florida and 
Louisiana. June—Oct. SH 


; Ui 
16. MECARDONIA R. & P. Syst. 164. 17098. 


Perennial relatively rigid herbs, with 4-angled erect or diffuse, but rarely creeping stems. 
Leaves opposite; blades toothed, narrowed at the base, pinnately nerved. Flowers solitary 
on slender axillary peduncles subtended by two small bractlets. Calyx not subtended by bract- 
lets; sepals 5, unequal, the upper one broadest; corolla white, purple or yellow, 2-lipped, the 
upper lip notched Stamens adnate to near the corolla-throat. Hypogynous disk wanting. 
Stigma 2-lobed. Capsule septicidal, the valves cleft. Seeds numerous. [Named in honor of 
Anton Meca y Cardona, a founder of the botanical garden at Barcelona. ] 


; About 10 species, natives of warm and tropical America. Besides the following, some 4 others 
inhabit the southeastern United States. Type species: Mecardonia ovata R. & P. 


1. Mecardonia acuminata (Walt.) Small. Purple Hedge-Hyssop. Fig. 3784. 


Gratiola acuminata Walt. Fl. Car. 61. 1788. 

Matourea nigrescens Benth. Comp. Bot. Mag. 1: 173. 
1835. 

Herpestis nigrescens Benth. Comp. Bot. Mag. 2: 56. 
1836. 


Monniera acuminata Kuntze, Rev. Gen. Pl. 463. 1801. 
Mecardonia acuminata Small, Fl. SE. U. S. 106s. 
1903. 


Perennial, glabrous; stem erect, branched above, 
1°-2° high, very leafy. Leaves oblong or oblong- 
lanceolate, serrate, at least above the middle, 
short-petioled or sessile, obtuse or acute at the 
apex, narrowed or somewhat cuneate at the base, 
1-2’ long, faintly veined; peduncles in alternate 
and opposite axils, ascending, in fruit longer than 
the leaves, not bracteolate at the summit; flowers 
5-6” long, purple or purplish; upper calyx-seg- 
ment lanceolate, acute; corolla 2-lipped, the lower 
lip longer than the upper; stamens approximate 
in pairs; capsule oblong, 4-valved, 3-4” high, 
about equalling the calyx. 


In wet soil, Maryland to Florida, Missouri and 
Texas, mostly near the coast. Plant blackening in 
drying. June—Sept. 


13 


194 SCROPHULARIACEAE. Vouz ite 


17. GRATIOLA L. Sp. Pl. 17. 1753. 


Erect or diffuse glabrous or glandular-pubescent herbs, with opposite sessile, entire or 
dentate leaves, and yellow or whitish peduncled flowers solitary in the axils. Peduncles 
2-bracteolate &4t the summit in the following species. Calyx 5-parted, the segments narrow, 
slightly unequal. Corolla irregular, its tube cylindric, its limb more or less 2-lipped; upper 
lip entire, emarginate, or 2-cleft; lower lip 3-lobed. Perfect stamens 2, the anterior pair 
wanting, or represented by rudiments; filaments filiform; anther-sacs distinct, transverse and 
separated by a broad connective. Style filiform; stigma dilated, slightly 2-lobed. Capsule 
loculicidally and septicidally dehiscent, ovoid or globose, 4-valved. Seeds numerous, longi- 
tudinally and transversely striate. [Latin, grace or favor, from its reputed healing properties. ] 

About 30 species, of wide geographic distribution in temperate and warm regions. Besides the 
on ae 9 others occur in southern and western North America. Type species: Gratiola 
Oficinalr1s 5 


Sterile filaments minute or none. 


Glandular-puberulent ; flowers 4”—5” long; capsule ovoid. 1. G. virginiana. 

Glabrous; flowers 7” long; capsule globose. : 2. G. sphaerocarpa. 
Sterile filaments 2, slender, capitate at the summit. 

Leaves lanceolate, entire or remotely denticulate. 3. G. aurea. 

Leaves ovate or oblong, sharply serrate. 4. G. viscosa. 


1. Gratiola virginiana L. Clammy Hedge- 
Hyssop. Fig. 3785. 
Gratiola virginiana L. Sp. Pl. 17. 1753. 


Annual; stem erect, at length widely branched, 
glandular-puberulent, at least above, 3-12’ high. 
Leaves oblong or oblong-lanceolate, sessile, nar- 
rowed to both ends, denticulate, 17-2’ long, 2-5” 
wide, glabrous or nearly so; peduncles slender, 
glandular, shorter than or equalling the leaves; 
flowers 4-5” long; bractlets as long as the calyx, 
or longer; calyx about one-half as long as the yel- 
lowish corolla-tube; limb of the corolla short, white; 
sterile filaments minute or none; anther-sacs trans- 
verse, separated by a broad connective; capsule 
broadly ovoid, 2” high, as long as the calyx. 

In wet places, Quebec to British Columbia, south to 


Florida, Texas and California. Ascends to 3000 ftant 
Virginia. Water jessamine. May—Oct. 


2. Gratiola sphaerocarpa Ell. Round- 
fruited Hedge-Hyssop. Fig. 3786. 


Gratiola sphaerocarpa Ell. Bot. S. C. & Ga. I: 14. 

1816. 

Annual (or perennial?), glabrous; stem as- 
cending or erect, rather stout, simple or branched, 
6-12’ high. Leaves oblong or obovate-oblong, 
sessile, dentate or denticulate, 3-5-nerved, acute 
or obtusish at the apex, narrowed at the base, 
1-2’ long, 3’-8” wide; peduncles stout, little cr 
not at all longer than the calyx; bractlets about 
equalling the calyx; calyx-lobes linear; calyx 
nearly one-half as long as the corolla; flowers 
about 7” long; corolla-tube yellow, the limb 
paler; anther-sacs broad, transverse; sterile fila- 
ments wanting; capsule globose, 3’’°in diameter. 


N 
UN 3 


In wet places, southern New Jersey to Florida, 
Illinois, Missouri, Texas and Mexico. June—Sept. 


GENUS 17. FIGWORT FAMILY. 195 


3. Gratiola aurea Muhl. Goldenpert. 
Golden Hedge-Hyssop. Fig. 3787. 


Gratiola aurea Muhl. Cat. 2. 1813. 


Perennial, glandular-puberulent above, or gla- 
brate; stems decumbent, creeping or ascending, 
simple or branched, 4-12’ long, somewhat 4-sided. 
Leaves lanceolate to linear-oblong, 3’-1’ long, 
1-3” wide, sparingly denticulate, scarcely nar- 
rowed to the sessile and somewhat clasping 
base; peduncles filiform, in fruit equalling or 
longer than the calyx; corolla bright yellow, 
6-7” long, 3 times as long as the calyx; sterile 
filaments 2, capitate at the summit; anther-sacs 
of the fertile stamens broad, transverse; capsule 
globose-ovoid, shorter than or equalling the calyx. 


In sandy wet places, Quebec and Ontario to New 
Jersey and Virginia. Recorded from Florida. June— 
Sept. 


4. Gratiola viscésa Schwein. Viscid 
Hedge-Hyssop. Fig. 3788. 


Gratiola viscosa Schwein.; LeConte, Ann. Lyc. N. 
Ye) r0G. reac: 


Annual; stem weak, finely viscid-pubescent, 
slender, commonly simple, 6-18’ long. Leaves 
ovate, ovate-oblong or ovate-lanceolate, sharply 
serrate, acute at the apex, sessile, cordate-clasp- 
ing at the base, 3’-1’ long; peduncles slender, 
shorter than or exceeding the leaves; bractlets 
and calyx-segments foliaceous, entire or dentate, 
one-third to one-half as long as the yellow or 
purplish corolla; flowers 5”-6” iong; sterile fila- 
ments 2, capitate at the summit; anther-sacs of 
the fertile stamens transverse, separated by the 
broad connective; capsule subglobose, shorter 
than the calyx. 


In brooks and swamps, Delaware to Kentucky, 
Georgia and Tennessee. May-Sept. 


18. SOPHRONANTHE Benth.; Lindl. Introd. Nat. Syst. Ed. 2, 445 1836. 


Annual or perennial rigid caulescent herbs, with rough-pubescent foliage. Leaves oppo- 
site; blades entire or toothed, sessile. Flowers solitary in the axils, short-peduncled or nearly 
sessile. Calyx sessile in 2 bractlets; sepals 5, nearly distinct. Corolla white or purplish, 
2-lipped. Stamens 2, included. Staminodia filiform, capitate at the apex. Anther-sacs par- 
allel, contiguous. Capsule somewhat elongated, acuminate. [Greek, referring to the included 
anthers. | 


Two known species of eastern North America. Type species: Sophronanthe hispida Benth. 


SCROPHULARIACEAE. VoL. IIT. 


1. Sophronanthe pilosa (Michx.) Small. 
Hairy Hedge-Hyssop. Fig. 3789. 


Gratiola pilosa Michx. Fl. Bor. Am. 1: 7. 1803. 
Sophronanthe pilosa Small, Fl. SE. U. S. 1067. 1903. 


Perennial (?), hirsute; stems slender, erect, 
strict, simple, or branched, 6—2° high. Leaves 
ovate or ovate-lanceolate, dentate or denticulate, 
sessile, acute or obtusish at the apex, rounded, 
truncate or subcordate at the base, 4-1’ long; 
flowers nearly or quite sessile, commonly numer- 
ous, about 4” long; corolla purplish or white, 
slightly longer than the calyx and _ bractlets; 
sterile filaments 2, capitate at the summit; anther- 
sacs of the fertile stamens parallel, contiguous; 
capsule oblong-conic, acuminate, about the length 
of the calyx, or shorter. 


In dry soil, southern New Jersey to Florida, Arkan- 
sas and Texas. May—Aug. 


19. ILYSANTHES Raf. Ann. Nat. 13. 1820. 


Annual or biennial glabrous slender branching herbs, with opposite, mostly dentate and 
sessile leaves, and small purplish peduncled flowers solitary in the axils. Peduncles not 
bracteolate. Calyx 5-parted, the segments linear. Corolla irregular, the tube somewhat 
expanded above, the limb 2-lipped; upper lip 2-cleft, erect; lower lip larger, 3-lobed, spread- 
ing. Fertile stamens 2, included, their anther-sacs divergent; sterile stamens 2, 2-lobed, one 
of the lobes capitate, glandular, the other glabrous, shorter. Style slender; stigma slightly 
2-lobed. Capsule oblong or ovoid, septicidally dehiscent. Seeds numerous, wrinkled. [Greek, 
mud-flower. | 


About 10 species, of wide geographic distribution. Besides the following, 2 or 3 others occur 
in the southeastern United States. Type species: [/ysanthes riparia Raf. 


Peduncles longer than the leaves; calyx-segments shorter than the capsule. 1. I. dubia, 
Peduncles shorter than the leaves; calyx-segments mostly as long as the capsule, or longer. 
2. IJ. attenuata, 


1. Ilysanthes ditbia (L.) Barnhart. Long-stalked False Pimpernel. Fig. 3790. 


Q ’ 
Gratiola dubia L. Sp. Pl. 17. 1753. Pas, \ 


Capraria gratioloides L. Sp. Pl. Ed. 2, 876. — 1763. 
Ilysanthes riparia Raf. Ann. Nat. 13. 1820. 


Ilysanthes gratioloides Benth. in DC. Prodr. 10: ( 
8 y) 


419. 46. 
Ilysanthes dubia Barnhart, Bull. Torr. Club 26: 

376. 1899. 

Stem slender, mostly erect, at length dif- Sy 
fusely branched, 3-8’ long. Leaves ovate, 
ovate-oblong, or the lower obovate, sessile or \ 
very nearly so, or slightly clasping at the base, 


remotely denticulate or entire, obtuse or acutish 
at the apex, thickish, 3-7-nerved, 3’-1’ long, 
the upper ones commonly much smaller; pe- 
duncles slender, considerably exceeding the 
leaves; flowers 37-5” long; calyx-segments 
linear, about one-half the length of the corolla, 
shorter than the capsule; capsule narrowly 
ovoid-oblong, bluntish, 2-3” high; seeds 1” 
long, reddish, the ends usually truncate. 


In wet places, Massachusetts to Florida, west 
to Ontario, Minnesota and Texas. Also on the 
Pacific Coast, and in the West Indies and South A ( 
America. July—Sept. D) 7. 


GENUS I9, FIGWORT FAMILY. 


2. Ilysanthes attenuata (Muhl.) Small. 
Short-stalked False Pimpernel. 
Fig. 3701. 
Lindernia attenuata Muhl. Cat. 59. 1813 ( 


Ilysanthes gratioloides curtipedicellata Bush, Bull. cA 
Torr. Club 21: 494. 189 


I. attenuata Small, Bull. Tor. Club 23: 297. 1896. Ee, 
Stem erect or ascending, 3-16’ long, the branches 
spreading. Leaves oblong to ovate, or sometimes , 
obovate, 2-12’ long, thinnish, obtuse, serrate with {f 
a few low teeth, 3-5-nerved, narrowed into short \ 
petioles, or sessile; peduncles shorter than the | | 
leaves; calyx-segments linear-subulate, as long as P| ¢ 
the capsule, or longer; corolla 2-6” long; cap- 
sule narrowly ovoid, about 2” long, pointed; seeds 
slightly curved, 14’—2” long, yellowish brown, the 
ends usually rounded. 

In wet places, New Brunswick and Ontario to Wis- 
consin, south to Florida and Arkansas. Ascends to 4 


2000 ft. in Virginia. Perhaps not specifically dis- 
tinct from the preceding. May—Oct. 


20. MICRANTHEMUM Michx. Fl. Bor. Am. r: to. pl. 12. 1803. 


Creeping or ascending, branched small leafy annual glabrous herbs, with opposite obovate 
oval or orbicular sessile entire leaves, and minute white or purplish short-peduncled flowers, 
solitary in some of the axils. Calyx 4-5-lobed or 4-5-parted. Corolla very irregular, the 
tube short, the upper lip shorter than the lower, or wanting, the lower 3-lobed, spreading or 
ascending, the middle lobe the largest. Stamens 2, anterior; filaments short, somewhat 
dilated or appendaged at the base; anthers small, their sacs distinct, parallel, or slightly 
divergent. Style short; stigma 2-lobed. Capsules globose, 2-celled by a membranous parti- 
tion or becoming I-celled. Seeds several or numerous, minute. [Greek, small flower.] 


About 16 species, natives of America. Besides the following, another occurs in the southern 
United States. Type species: Micranthemum orbiculatum Michx. 


1. Micranthemum micranthemoides (Nutt.) Wettst. Nuttall’s Micranthemum. 
Fig. 3702. 


Hemianthus micranthemoides Nutt. Journ. Acad. Phil. 
Ts ©l9o. pls 6. 1817. 

Micranthemum Nuttallii A. Gray, Man. Ed. 5, 331. 1867. 

Micranthemum micranthemoides Wettst. in Engl. & 
Prantl, Nat. Pfl. Fam. 4: Abt. 3b. 77. 1891. 


Somewhat fleshy; stem filiform, creeping, the 
branches ascending, 4-23’ high. Leaves obovate to 
oval, obtuse, 1-24” long; flowers about 3” long, 
borne on peduncles of about the same length; calyx 
campanulate in flower, obovoid in fruit, 4-lobed, 
usually split along one side; peduncles recurved in 
fruit; upper lip of the corolla nearly obsolete; mid- 
dle lobe of the lower lip longer than the lateral ones; 
appendages at the bases of the stamens nearly as 
long as the filaments; capsule obovoid-globose, 4” in 
diameter, as long as the calyx. 


IN) 


In tidal mud, New Jersey to Florida. Aug —Oct. 
This species is the type of the genus Hemianthus Nutt., 
which has been regarded as distinct from Micranthe- 
mum by other authors, a view which may be maintained. 


21. LIMOSELLA L. Sp. Pl. 631. 1753. 


Low glabrous succulent floating or creeping, tufted annual herbs (or perennial by stolons?), 
with filiform stems rooting at their nodes, basal slender-petioled entire leaves, and filiform 
1-flowered scape-like peduncles, the flowers small, white, pink, or purple. Calyx campanu- 
late, 5-lobed. Corolla nearly regular, open-campanulate, the tube short, the limb 5-cleft. 
Stamens 4, inserted on the corolla-tube, scarcely exserted; filaments short: anther-sacs con- 
fluent. Style short; stigma capitate. Ovary 2-celled at the base, I-celled above. Capsule 
globose or oblong, becoming I-celled, many-seeded. [Greek, mud seated.] 

About 6 species, of wide geographic distribution. Type species: Limosella aquatica L. 


198 SCROPHULARIACEAE. Vor. III. 


1. Limosella aquatica L. Mudweed. Mudwort. Fig. 3793. 


Limosella aquatica L. Sp. Pl. 631. 1753. 
TEN tenuifolia Hoftm. Deutsch. Fl. 29. 
180 


4. 
Limosella australis R. Br. Prodr. Fl. Nov. Holl. 
I: 443. 1810. 


Leaves 17-5’ long, the blade oblong, linear- 
oblong, narrowly linear or spatulate, ob- 
tuse, one-fourth or one-third as long as 
the filiférm petiole. Peduncles shorter than 
the leaves, arising with the petioles from 
the base of the plant or from nodes of the 
creeping or floating stem; corolla pink or 
white, about 1” broad, scarcely longer than 
the calyx; calyx-lobes ovate, acute or acut- 
ish, about the length of the tube; stamens 
inserted high up on the corolla-tube; fila- 
ments somewhat longer than the anthers; 
capsule globose or oblong-globose, obtuse, 
12” high, longer than the calyx. 

On muddy shores and in brooks, Labrador 
and Hudson Bay to the Northwest Territory, 
south to New Jersey, and in the Rocky Moun- 
tains to Colorado, and in the Sierra Nevada to 


California. Also in Europe, Australia and 
South America. June—Aug. 


22. SYNTHYRIS Benth. in DC. Prodr. 10: 454. 1846. 


Perennial herbs, with thick rootstocks, simple erect stems, large petioled basal leaves, 
those of the stem much smaller, alternate, sessile, or clasping, bract-like. Flowers small, pink 
or purple, in terminal dense elongated spikes or racemes. Calyx 4-5-parted, the segments 
oblong or linear. Corolla oblong or campanulate, 2-4-lobed, or parted, or wanting, the lateral 
lobes, when present, exterior in the bud. Stamens 2 (occasionally 4), posterior, inserted on 
the corolla, or on the outer side of the hypogynous disk, exserted; filaments slender; anther- 
sacs parallel or divergent, not confluent. Ovary 2-celled or rarely 3-celled; style filiform; 
stigma capitate. Capsule compressed, obtuse, or emarginate, many-seeded, loculicidally dehis- 
cent. Seeds flat, oval, or orbicular. [Greek, closed doors, referring to the capsule-valves. ] 
Our species were referred, in the first edition of this work, to the Old World genus Wulfenia. 


About 10 species, natives of North America and Europe. Type species: Synthyris reni- 
formis Benth. 


Corolla present, usually 2-lobed. r. S. Bullit. 
Corolla none. 2. S. rubra. 


1. Synthyris Bullii (Eaton) Heller. Bull’s Synthyris. Fig. 3794. 
Gymnandra Bullii Eaton; Eaton & Wright, 259. 1840. PoP 
Synthyris Houghtoniana Benth. in DC. Prodr. 10: 454. 

1846. 
Wulfenia Houghtoniana Greene, Erythea 2: 83. 1894. 
Gymnandra Bullii Barnhart, Bull. Torr. Club 26: 378. 1899. 
Synthyris Bullii Heller, Muhlenbergia 1: 4. 1900. 


Pubescent; stem stout, 1°-23° high. Basal leaves 
ovate or orbicular, rounded at the apex, truncate, cor- 
date or reniform at the base, crenulate all around, 2-5’ 
long, 5-7-nerved, petioled, the petiole usually shorter 
than the blade; stem-leaves small, 4’-1’ long, sessile 
or slightly clasping, crenulate, obtuse, or acute, passing 
gradually into the bracts of the dense spike; flowers 
greenish yellow, 2’-3” long; corolla present, variously 
2-4-lobed (commonly 2-lobed), little, if any, longer than 
the calyx, its lobes obtuse, the stamens inserted on its 
base; spike much elongated in fruit; capsule emargi- 
nate, slightly exceeding the calyx. 


On dry prairies, Ohio to Minnesota, Michigan and Iowa. 
May-July. 


GENUS 22. FIGWORT FAMILY. 199 


2. Synthyris rubra (Hook.) Benth. Western Synthyris. 
Gymnandra rubra Hook. Fl. Bor. Amer. 2: 103. pl. 172. Oe 
1838. S 2 


Fig. 3795. 


Synthyris rubra Benth. in DC. Prodr. 10: 455. 1846. 


Wulfenia rubra Greene, Erythea 2: 83. 1894. 
Besseya rubra Rydberg, Bull. Torr. Club 30: 280. © 1903. 


Similar to the preceding species but lower, pubescent 
or tomentose, seldom over 1° high. Basal leaves ovate 
or oblong, obtuse or acute at the apex, narrowed, trun- 
cate or cordate at the base, 14’-3’ long, crenulate, peti- 
oled, indistinctly nerved; stem-leaves ovate or lanceo- 
late, acute, sessile, crenulate, or entire, 7’-1’ long; spike 
very dense, 1-2’ long in flower, 2-5’ long in fruit, its 
bracts purplish; corolla none; stamens inserted on the 
outer side of the hypogynous disk; capsule little com- 


pressed, emarginate, slightly longer than the calyx. 


In dry soil, South Dakota to Nebraska, British Columbia and Utah. May-June. 


23. VERONICA [Tourn.] L. Sp. Pl. 9. 1753. 


Annual or perennial herbs (some exotic species shrubs or trees), with opposite and alter- 
nate, rarely verticillate leaves, and mostly small blue purple pink or white flowers, terminal 
or axillary, racemose, spicate, or solitary. Calyx mostly 4-parted, sometimes 5-parted, the seg- 
ments oblong or ovate. Corolla rotate, its tube very short, deeply and more or less unequally 
4-lobed (rarely 5-lobed), the lower lobe commonly the narrowest. Stamens 2, divergent, 
inserted on either side and at the base of the upper corolla-lobe; anthers obtuse, their sacs 
confluent at the summit; filaments slender. Ovary 2-celled; style slender; stigma capitate; 
ovules few or numerous in each cavity. Capsule more or less compressed, sometimes very 
flat, emarginate, obcordate, or 2-lobed, loculicidally dehiscent. Seeds smooth or rough, flat, 
plano-convex, or excavated on the inner side. [Named for St. Veronica. ] 


About 200 species, of wide geographic distribution. Besides the following, 3 others occur in 
northwest America. Type species: Veronica officinalis L. 


* Flowers racemose in the axils of the leaves, bracteolate. 


Glabrous, or minutely glandular above (No. 3 rarely hairy) ; brook or swamp plants. 
Leaves ovate, oval, oblong, or oblong-lanceolate ; capsule compressed. 
Stem leaves sessile, partly clasping, serrulate or entire. 1. V. Anagallis-aquatica. 
All the leaves petioled, serrate. 2. V. americana. 
Leaves linear or linear-lanceolate ; capsule very flat. 3. V. scutellata. 
Pubescent, dry soil plants; leaves crenate or dentate. 
Leaves oval or obovate, petioled ; pedicels shorter than the calyx. 


t i V. officinalis. 
Leaves ovate, nearly or quite sessile ; pedicels longer than the calyx. 


V. Chamaedrys. 


ns 


** Flowers in terminal spikes or racemes, or solitary in the axils. 


Flowers in terminal spikes or racemes. 


Leaves all sessile ; capsule elliptic, emarginate. 6. V. Wormskioldii. 

Lower leaves petioled; capsule orbicular, obcordate. 7. V. serpyllifolia. 
Flowers solitary in most of the axils; peduncles shorter than the leaves. 

Erect; glabrous or glandular; capsule emarginate. 8. V. peregrina. 

Diffuse; pubescent; capsule obcordate. 9. V. arvensis. 


Flowers solitary in the axils; peduncles as long as the leaves, or longer. 
Leaves ovate or oblong, crenate or dentate. 
Corolla not longer than the calyx; capsule narrowly emarginate. 10. V’. agrestis. 
Corolla longer than the calyx; capsule broadly emarginate. 11. V. Tournefortii. 
Leaves orbicular, or broader, 3-s5-lobed or -crenate. 12. V.hederaefolia. 


200 SCROPHULARIACEAE. Vor iit 


1. Veronica Anagallis-aquatica L. Water 
Speedwell or Pimpernel. Fig. 3796. 
Veronica Anagallis-aquatica L. Sp. Pl. 12. 1753. 


Perennial by stolons or leafy shoots developed in 
autumn; stem rather stout, glabrous, or glandular- 
puberulent above, erect or decumbent, often rooting 
at the lower nodes, usually branched, 1°-3° high. 
Leaves of sterile autumn shoots orbicular to obovate, 
obtuse, serrulate, narrowed into margined petioles, 
those of the flowering stems ovate, oblong, or lan- 
ceolate, sessile and more or less clasping or the 
lowest short-petioled, serrulate or entire, 13-4’ long, 
4’-2’ wide; racemes peduncled, borne in ‘most of the 
axils, 2-5’ long; bractlets shorter than or exceeding 
the pedicels; flowers blue, or purplish striped, 2” 
broad; capsule compressed, not very flat, nearly or- 
bicular, 2-lobed, emarginate, 12” high; seeds flat. 


In brooks and swamps, Nova Scotia to British Co- 
lumbia, south to North Carolina, Nebraska and New 
Mexico. Also in Europe and Asia. The plant of the 
Atlantic Coast appears as if introduced. Ascends to 4000 ft. in Virginia. May—Sept. 


2. Veronica americana Schwein. American Brooklime. Fig. 3797. 


V. americana Schwein. ; Benth. in DC. Prodr. 10: 468. 1846. 


Similar to the preceding species, perennial by stolons 
or leafy shoots, glabrous throughout; stem decumbent, 
usually branched, rooting at the lower nodes, 6’—3° long. 
Leaves oblong, ovate or oblong-lanceolate, all distinctly 
petioled, sharply serrate, truncate, rounded, or subcor- 
date at the base, obtuse or acutish at the apex, 1-3’ 
long, #’/-1’ wide; racemes peduncled, borne in most of 
the axils, loose, elongated, sometimes 6’ long; bractlets 
shorter than the pedicels; flower blue or nearly white, 
usually striped with purple, 2” broad; capsule nearly 
orbicular, compressed, but not very flat, emarginate, 13” 
high; seeds flat. 


In brooks and swamps, Anticosti to Alaska, south to 
Pennsylvania, Nebraska, New Mexico and California. 
Ascends to 2600 ft. in the Catskills. Wallink. Blue-bells. 
April—Sept. 


Veronica Beccabinga L., a European brooklime, similar 
to V. americana in habit, but with crenate or low-serrate 
broad tipped leaves, is naturalized about Quebec and has been found on ballast about seaports in 
New York and New Jersey. 


3. Veronica scutellata L. Marsh or Skullcap 
fs Speedwell. Fig. 3798. 


Veronica scutellata L. Sp. Pl. 12. 1753. 


Glabrous, or very sparingly pubescent, rarely quite 
hairy, perennial by leafy shoots or stolons; stems slen- 
der, decumbent or ascending, leafy, simple or branched, 
commonly rooting at the lower nodes, 62° high. Leaves 
linear or linear-lanceolate, sessile and slightly clasping, 
remotely denticulate, acute, 1-3’ long, 1’-3” wide; ra- 
cemes borne in nearly all the axils, or only in the alter- 
nate ones, equalling or longer than the leaves; bractlets 
much shorter than the filiform spreading pedicels; flow- 
ers blue, 2-3” broad; capsule broader than high, very 
flat, deeply emarginate at the summit, slightly so at the 
base, 2-23” broad; seeds flat. ; 


In swamps, Labrador to British Columbia, south to south- 
ern New York, Minnesota and California. Also in Europe 
and Asia. May-Sept. 


GENUS 23. FIGWORT FAMILY. 201 


4. Veronica officinalis L. Common Speed- 
well. Fluellin. Gipsy-weed. Fig. 3799. 


Veronica officinalis L. Sp. Pl. 11. 1753. 


Perennial by stolons, pubescent all over; stem as- 
cending, 3-10’ high. Leaves oblong, oval, or obo- 
vate, petioled, 4’—2’ long, obtuse at the apex, serrate, 
narrowed into the petioles; racemes spike-like, nar- 
row, dense, elongated, often borne only in alternate 
axils, much longer than the leaves; subulate bract- 
lets and the calyx longer than the pedicels; flowers 
pale blue, 2’-3” broad; capsule obovate-cuneate, 
compressed, broadly emarginate, 2” high, 12” broad; 
seeds numerous, flat. 


In dry fields and woods, Nova Scotia to Ontario, 
South Dakota, North Carolina and Tennessee. As- 
cends to 5600 ft.in Virginia. Also in Europe and Asia. 
Appears, in most places, as if introduced. Paul’s-betony. 
Ground-hele. Upland speedwell. May—Aug. 


Veronica longifdlia L., an erect European species 
with lanceolate petioled acuminate sharply serrate 
leaves, has been found in waste grounds and fields 
from Nova Scotia to New York. 


\)) WE 
SY Ae 
=n SY 


5. Veronica Chamaédrys L. Germander 
Speedwell. Fig. 3800. 


Veronica Chamaedrys L. Sp. Pl. 13. 1753. 


Perennial; stem ascending, simple or branched, 
slender, pubescent in two lines, 4-12’ high. Leaves 
ovate, sessile, or very nearly so, pubescent, trun- 
cate, rounded or cordate at the base, incised- 
dentate, obtuse at the apex, +”-17’ long; racemes 
borne in opposite or alternate axils, peduncied, 
more or less pubescent, loose, 2-6’ long; pedicels 
filiform, longer than the calyx and usually longer 
than the bractlets; flowers light blue, 3’-4” 
broad; capsule obcordate, narrowed at the base; 


seeds numerous, flattish. 


In fields and waste places, Nova Scotia and 
Quebec to southern New York and Pennsylvania. 
Naturalized from Europe. Other English names 
are blue eye, eyebright, angel’s-eye, god’s-eye, bird’s- 
eye, cat’s-eye, base vervain. Forget-me-not. May-— 
July. 

Veronica Tetcrium L., also European, similar to 
V. Chamaedrys, but with oblong to lanceolate 
crenate leaves, has been found in New England and 
Ohio. 


6. Veronica Wormskioldii R. & S. Worms- 
kiold’s Speedwell. Fig. 3801. 


V. Wormskioldii R. & S. Syst. 1: 101. 1817. 


Perennial, pubescent or nearly glabrous; stems 
ascending or erect, slender, usually simple, 2’—12’ 
high. Leaves oblong, ovate, or elliptic, sessile, mostly 
rounded at both ends, crenulate or entire, 4-1’ long; 
flowers in a short narrow raceme at the end of the 
stem, light blue, 2-3” broad; pedicels shorter than 
the calyx in flower, much shorter than the bractlets, 
2’-3” long in fruit; capsule ellipsoid or slightly obo- 
void, moderately compressed, emarginate, 2”—3” high; 
seeds numerous, flattish. 


Labrador; mountains of Quebec and New England 
to Alaska, south in the Rocky Mountains to Colorado 
and in the Sierra Nevada to Nevada. Summer. Re- 
ferred in our first edition to V. alpina L 


202 SCROPHULARIACEAE. Vor. III. 


7. Veronica serpyllifolia L. Thyme-leaved 
Speedwell. Fig. 3802. 


Veronica serpyllifolia L. Sp. Pl. 12. 1753. 


Perennial, puberulent or glabrous; stems slender, 
decumbent, branched, the branches ascending or 
erect, 2’-10’ high. Leaves all opposite and petioled, 
or the uppermost sessile, oblong, oval, or ovate, #’—3’ 
long, crenulate or entire; flowers in short narrow 
racemes at the end of the stem and branches; pedi- 
cels equalling or longer than the calyx, usually shorter 
than the bractlets; corolla blue with darker stripes, 
sometimes white, 2-4” broad; capsule broader than 
high, broadly obcordate or emarginate at the summit, 
about 1” long, about equalling the calyx; seeds flat, 
numerous. 

In fields and thickets, Labrador to Alaska, south to 
Georgia, New Mexico and California. Also in Europe, 
Asia and South America. Ascends to 2600 ft. in the 
Catskills. Paul’s-betony. April—Aug. 


Veronica humifiisa Dickson, differing in larger flow- 
ers and more pubescent inflorescence, is apparently a 
northern race of this species, also occurring in Europe. 


8. Veronica peregrina L. Purslane Speedwell. 
Neckweed. Fig. 3803. 


Veronica peregrina L. Sp. Pl. 14. 1753. 


Vs 
WZ 
Annual, glabrous, or glandular-puberulent; stem N Uf 
erect or ascending, simple or branched, 3-12’ high. Y| 
Leaves cblong, oval, linear or slightly spatulate, 3-10” NY 
long, obtuse or acutish, the lowest opposite, short- ‘ 


WY 
a 


petioled, or sessile, broader than the upper and usually Vy p 
denticulate, the upper alternate, sessile, mostly entire, \\) Wy P 
each with a short-pedicelled flower in its axil; flowers WV Y Y 


nearly white, about 1” broad; pedicels much shorter i) 
than the calyx; capsule nearly orbicular, obcordate, a 
usually a little shorter than the calyx, 1’-13” high, > 

many-seeded, the seeds flat. Uf IN 


In moist places, and common as a weed in cultivated . N 
soil, Nova Scotia to British Columbia, south to Florida, 


Mexico and California. Also in Central and South Amer- S 
‘ea, distributed as/a(weed in the Old) Werld.) May-Oct: T— 


4 74) Welw 


9. Veronica arvénsis L. Corn or Wall 
Speedwell. Fig. 3804. 
Veronica arvensis L. Sp. Pl. 13. 1753. 


Annual, pubescent; stem slender, at first sim- 
ple and erect, at length much branched and dif- 
fuse, 3-10’ long. Lower leaves ovate or oval, 
opposite, obtuse at both ends, crenate or crenu- 
late, 2-6” long, the lowest petioled; upper leaves 
sessile, alternate, ovate or lanceolate, acute or 
acutish, commonly entire, each with a short pedi- 
celled minute flower in its axil; pedicels shorter 
than the calyx; corolla blue, or nearly white, 1” 
broad or less; capsule broadly obovate, obcordate, 
1” high. 

In fields, woods and waste places and in culti- 
vated soil, Nova Scotia to Ontario and Minnesota, 
south to Florida, Kansas and Texas. Also in Ber- 


muda. Naturalized from Europe. Native also of 
Asia. March-—Sept. 


GENUS 23. FIGWORT FAMILY. 203 


10. Veronica agréstis L. Procumbent, Field 
or Garden Speedwell. Fig. 3805. 
Veronica agrestis L. Sp. Pl. 13. 1753. 


Annual, pubescent; stems creeping or procumbent, 
very slender, branched, 3’-8’ long, the branches as- 
cending or spreading. Leaves broadly ovate or oval, 
obtuse at the apex, rounded, truncate or subcordate 
at the base, crenate, all short-petioled, or the upper- 
most sessile, the lowest opposite, the upper alternate 
and each with a slender-peduncled small blue flower 
in its axil; peduncles equalling or longer than the 
leaves; corolla not exceeding the calyx; capsule 
broader than high, compressed, but not very flat, nar- 
rowly emarginate at the summit, 1” high, 2” broad; 
seeds few, hollowed out on the inner side. 


In fields and waste places, Nova Scotia to New Jersey 
and Louisiana. Naturalized from Europe. Native also 
of Asia. Other English names are germander-chickweed 
and winter-weed. May-Sept. e 


11. Veronica Tournefortii Gmelin. Tournefort’s Speedwell. Byzantine Speed- 
well. Fig. 3806. 


Veronica Tournefortii Gmelin, Fl. Bad. 1: 39. 1805. 

Veronica agrestis var. byzantina Sibth. & Smith, 
Fl. Graec. 1: pl. 8. 1806. 

V. Buxbaumii Tenore, Fl. Nap. 1: 7. pl. r. 1811. 

V. byzantina B.S.P. Prel. Cat. N. Y. 40. 1888. 


Annual, pubescent; stems diffusely branched, 
spreading or ascending, 6-15’ long. Leaves 
ovate or oval, short-petioled, obtuse or acutish, 
sometimes narrowed at the base, crenate-dentate 
or somewhat incised, 4-12” long, the lowest 
opposite, the upper all alternate and each with 
a slender-peduncled rather large blue flower in 
its axil; peduncles filiform, as long as the leaves 
or longer; corolla exceeding the calyx, 3-4” 
broad; capsule twice as broad as high, 3” broad, 
with a wide and shallow emargination at the 
summit; seeds few or several, hollowed out on 
the inner side. 


In waste places, Nova Scotia to southern New 
York and Ohio; Colorado to California. Adventive 
or naturalized from Europe. Native also of Asia. 
Bird’s-eye. Cat’s-eye. May—Sept. 


12. Veronica hederaefolia L. Ivy-leaved 
i ‘ 
Speedwell. Fig. 3807. 
Veronica hederaefolia L. Sp. Pl. 13. 1753. 


Annual, pubescent; stems slender, diffusely 
branched, 3-18’ long. Leaves orbicular or 
broader, truncate or subcordate at the base, 
3-5-lobed or 3-5-crenate, petioled, #’/-1’ in diam- 
eter, the lower opposite, the upper all alternate 
and with slender peduncled small blue flowers 
in their axils; peduncles filiform, often longer 
than the leaves; corolla 2” broad, scarcely 
longer than the calyx; capsule little compressed, 
2-lobed, broader than high, shorter than the 
densely ciliate sepals, 2-4-seeded; seeds exca- 
vated on the inner side. 


In thickets, fields and waste places, southern 
New York, Pennsylvania and New Jersey to South 
Carolina. Naturalized from Europe. Native also 
of Asia. April-Oct. Ivy-chickweed. Mother-of- 
wheat. Small henbit. Winter-weed. Morgeline. 


204 SCROPHULARIACEAE. Vor. III. 


24. LEPTANDRA Nutt. Gen. 1: 7. 1818. 


Tall stout erect perennial herbs, with verticillate or opposite leaves, and small minutely 
bracted white or blue flowers, in dense peduncled spike-like racemes, terminal, or in the’ 
upper axils. Calyx 4-parted, short. Corolla tubular, or salverform, nearly regular, 4-lobed, 
the tube cylindric, longer than the lobes. Stamens 2, exserted, inserted low down on the 
corolla-tube; filaments filiform; anthers obtuse, short. Style about as long as the stamens, 
stigma minute. Capsule narrowly ovoid, scarcely compressed, not emarginate nor obcordate, 
4-valved at the apex. Seeds numerous, oval, minutely reticulated. [Greek, slender stamens, 
referring to the filaments. | 


Two species, the following typical one native of eastern North America, the other of north- 
eastern Asia. 


1. Leptandra virginica (L.) Nutt. Beaumont’s-, 
Bowman’s- or Culver’s-root. Fig. 3808. 


Veronica virginica L. Sp. Pl. 9. 1753. 
Leptandra virginica Nutt. Gen. 1: 7. 1818. 


Stem glabrous, or very nearly so, simple, strict, 2°-7° 
high. Leaves verticillate in 3’s—o’s or some of the upper- 
most opposite, lanceolate or oblong-lanceolate, short- 
petioled, long-acuminate at the apex, sharply serrulate, 
narrowed at the base, pinnately veined, glabrous both 
sides, or pubescent beneath, 3-6’ long, 3-1’ wide; spike- 
like racemes several or rarely solitary, 3’-0’ long, very 
dense, the terminal one first developing; pedicels and 
bractlets about as long as the calyx; calyx-segments 
ovate-lanceolate, acute; corolla tubular, white or bluish, 
2” long; capsule ovoid-oblong, 1-13” long, 2-3 times 
as long as the calyx. 


I 


Qa 


S > \ 
SS , 
SW 
S77 
QO 
. 45 
D 


Wi 


In meadows, moist woods and thickets, Ontario to Mani- 
toba, Massachusetts, Alabama and Texas. Recorded from 
Nova Scotia. Ascends to 2700 ft. in Virginia. Black-root. 
Culver’s-physic. Brinton’s-root. Oxadaddy. Quitch. Tall 
speedwell. June—Sept. 


25. DIGITALIS (Mourn L.Sp. Rivc2naeatnise: 


Tall biennial or perennial herbs, the stems simple, or branched at the base, with alternate 
dentate or entire leaves, and large showy purple yellowish or white flowers, in long terminal 
commonly 1-sided racemes. Calyx 5-parted, the segments imbricated. Corolla declined, 
somewhat irregular, the tube contracted above the ovary, then rather abruptly expanded, 
longer than the 4-5-lobed slightly 2-lipped limb; upper lip broadly emarginate or 2-cleft; 
lower lip 3-lobed, the middle lobe largest, the lateral ones exterior in the bud. Stamens 4, 
didynamous, ascending, mostly included; anthers approximate in pairs. Style slender; stigma 
2-lobed. Capsule ovoid, septicidally dehiscent. Seeds numerous, rugose. 
the finger of a glove, which the flowers resemble. | 


About 20 species, native of Europe and Asia, the 
following typical. 


1. Digitalis purpurea L. Purple Fox- 
glove. Thimbles. Fairy Cap. Fig. 3809. 


Digitalis purpurea L. Sp. Pl. 621. 1753. 


Usually biennial, pubescent; stem stout, erect, 
2°-5° high. Basal and lower leaves ovate or 
ovate-lanceolate, 6-10’ long, slender-petioled, 
acute at the apex, narrowed at the base, dentate; 
upper leaves similar, smaller, sessile; racemes I° 
long or more, dense, I-sided; flowers purple to 
white, 12’-2’ long, drooping; upper calyx-segment 
narrower than the four other foliaceous ones; 
corolla spotted within. 


Cape Breton Island and New York, naturalized 
from Europe, sparingly escaped’ from cultivation; 
also from Washington to California. June—Aug. 
Among some 60 English names are folk’s-glove [by 
corruption fox-glove], 7. e., fairy’s-glove, fairy- 
thimbles, -fingers, -weed, fairy-bells, pop-dock, or 
-glove. Rabbit’s-flower. Cottagers. Lion’s-mouth. \ ; 
Scotch mercury. Throatwort. Lady-fingers, -glove, -thimble. Dog’s-finger. Witches’-thimbles. 


GENUS 26. FIGWORT FAMILY. 205 


26. BUCHNERA L. Sp. Pl. 630. 1753. 


Erect, perennial or biennial, simple or branched, strict hispid or scabrous herbs, black- 
ening in drying, the lower leaves opposite, the upper sometimes alternate. Flowers rather 
large, white, blue, or purple, in dense terminal bracted spikes, the lower commonly distant. 
Calyx tubular, or oblong, 5-10-nerved, 5-toothed. Corolla salverform, its tube cylindric, 
somewhat curved, its limb deeply and nearly equally 5-cleft, spreading, the lateral lobes 
exterior in the bud. Stamens 4, didynamous, included; anther-sacs confluent into 1. Style 
slender, thickened or club-shaped above; stigma small, entire or emarginate. Capsule oblong 
or ovoid, loculicidally dehiscent. Seeds numerous, reticulated. [Named for J. G. Buchner.] 


About 30 species, natives of warm and temperate regions. Besides the following, another 
occurs in the southern United States. Type species: Buchnera americana L. 


1. Buchnera americana L. Blue-hearts. 
Fig. 3810. 
Buchnera americana L. Sp. Pl. 630. 1753. 

Hispid and rough; stem slender, stiff, 1°-23° high. 
Leaves usually all opposite, prominently veined, the 
lowest obovate or oblong, obtuse, narrowed into very 
short petioles, the middle ones oblong or oblong-lan- 
ceolate, dentate, obtuse, or acute at the apex, narrowed 
at the base, sessile, the upper lanceolate or linear-lan- 
ceolate, entire or nearly so; spike peduncled, 6-10’ long 
in fruit, the flowers mostly opposite, nearly 1’ long; 
bractlets shorter than the calyx; calyx strigose; corolla 
purple, its lobes obovate, obtuse, 3-4” broad; capsule 
slightly oblique, 4” high, a little longer than the calyx. 

In sandy or gravelly soil, New Jersey to western New 
York, southern Ontario and Minnesota, south to Virginia, 
Louisiana, Kansas and Arkansas. June—Sept. 


: Als 
27. AFZELIA J. G. Gmel. Syst. Naf. 2: 927. 1796. 2 
[SeymeRIA Pursh, Fl. Am. Sept. 736. 1814.] 

Erect stout branched annual or perennial herbs, mostly with opposite leaves, at least the 
lower I-2-pinnately parted or dissected, and yellow flowers solitary in the axils, or in ter- 
minal bracted spikes or racemes. Calyx campanulate, 5-cleft or 5-parted. Corolla slightly 
irregular, campanulate or rotate, the tube short, broad, the limb 5-lobed, the lower lobe 
exterior in the bud. Stamens 4, slightly unequal, scarcely or not at all exserted; filaments 
short, villous, at least near the base; anthers 2-celled, the sacs parallel, distinct. Style short 
or slender. Capsule globose or ovoid, acute and more or less compressed at the summit. 
Seeds numerous, reticulated. [Named for Adam Afzelius, 1750-1812, professor at Upsala.] 


About 10 species, natives of North America, Mexico and Madagascar. Besides the following, 
4 others inhabit the southern United States. Type species: Anonymos cassioides Walt. 


IZ 1. Afzelia macrophylla (Nutt.) Kuntze. 


Mullen Foxglove. Fig. 3811. 


Seymeria macrophylla Nutt. Gen. 2: 49. 1818. 
Gerardia macrophylla Benth. Comp. Bot. Mag. 1: 205. 


1835. 
A, anrophella Kuntze, Rev. Gen. Pl. 457. 1801. 

Annual (?), puberulent or glabrate; stem spar- 
ingly branched, or simple, 4°-6° high. Lower 
leaves long-petioled, pinnately parted, 615’ long, 
their segments lanceolate, coarsely dentate, irregu- 
larly incised, or pinnatifid; upper leaves short- 
petioled or sessile, oblong or lanceolate, 1’—3’ long, 
entire, obtuse or acutish at the apex, narrowed at 
the base, each with a sessile flower in its axil; 
flowers 5’-7” long; calyx-lobes lanceolate or 
ovate, acute, about as long as the tube; corolla 
2-3 times as long as the calyx, woolly in the 
throat; style short, club-shaped; capsule globose- 
ovoid, 3-4” high, twice as long as the calyx. 

In moist thickets and along streams, Ohio to 
Towa, Nebraska, Kentucky and Texas. Aug.—Oct. 


28. DASYSTOMA Raf. Journ. Phys. 89: 99. 18109. 
Large erect simple or branched, glandular-puberulent, pubescent or glabrous, annual or 
perennial herbs, partly parasitic on the roots of other plants, with opposite whorled or some 
alternate leaves, and large showy yellow flowers, in terminal mostly leafy-bracted racemes 


206 SCROPHULARIACEAE. Vor. III. 


or panicles. Calyx campanulate or turbinate, 5-lobed, the lobes longer than or equalling 
the tube, sometimes foliaceous. Corolla slightly irregular, funnelform, or campanulate- 
funnelform, the tube villous or pubescent within, the limb spreading, 5-lobed. Stamens 4, 
didynamous, included, villous or pubescent; filaments slender; anthers all alike, their sacs 
distinct, parallel, awned at the base. Style filiform. Capsule oblong, acute, loculicidally 
dehiscent, longer than the calyx. [Greek, thick or hairy mouth, referring to the corolla.] 


Six species, natives of eastern North America. Type species: Dasistoma aurea Raf. 


Plant glandular-pubescent ; corolla pubescent without. 1. D pedicularia. 
Puberulent, cinereous or glabrous; corolla glabrous without. 
Cinereous-puberulent. 
Leaves entire, dentate, or some of the lower pinnatifid, firm. 
Leaves entire, undulate or the lower pinnatifid, with entire lobes. 
Upper leaves mostly serrate, lower pinnatifid with toothed lobes. 
Leaves, at least all but the uppermost, pinnatifid. 
Glabrous or very nearly so throughtout. 
Leaves all pinnatifid, thin. 
Leaves entire, or the lowest dentate or incised. 


D. flava. 
. D. serrata. 
. D. grandiflora. 


wh 


. D. laevigata. 
. D. virginica. 


Hu 


1. Dasystoma pedicularia (L.) Benth. 
Fern-leaved or Lousewort False Fox- 
glove. Fever-weed. Fig. 3812. 

Gerardia pedicularia L. Sp. Pl. 611. 1753. 
Dasystoma pedicu'aria Benth. in DC. Prodr. 

IO: 521. 1846. 

Annual or biennial, more or less glandu- 
lar-pubescent, viscid, and with some longer 
hairs; stem rather slender, much branched, 
leafy, 1°-4° high. Leaves sessile, or the 
lower petioled, 1-2-pinnatifid, ovate or 
* ovate-lanceolate in outline, usually broadest 
at the base, 1-3’ long, the segments incised 
or crenate-dentate; pedicels slender, ascend- 
ing, mostly longer than the calyx, 1-2’ long 
in fruit; calyx-lobes oblong, foliaceous, 
usually incised or pinnatifid, 3’-4” long, 
corolla 1-12’ long, pubescent without, limb 
about 1’ broad; capsule pubescent, 5-6” 
long, beak flat. 

In dry woods and thickets, Maine and On- 
tario to Minnesota, Florida and Missouri. 
Races differ in pubescence and in leaf-divi- 
sion. Lousewort. Bushy gerardia. Aug— 
Sept. 


Dasystoma pectinata (Nutt.) Benth. ofthe Southern States, ranging north to Kentucky 
and Missouri, appears to be a very glandular race. 


2. Dasystoma flava (L.) Wood. Downy 
False Foxglove. Fig. 3813. 


Gerardia flava L. Sp. Pl. 610. 1753. 

D. aurea Raf. Journ. Phys. 89: 99. 1819. 

D. pubescens Benth. in DC. Prodr. 10: 520. 1846. 
Dasystoma flava Wood, Bot. & Flor. 230. 1873. 


Perennial, downy, grayish; stem strict, erect, 
simple, or with a few nearly erect branches, 2°—4° 
high, Leaves oblong, lanceolate or ovate-lanceo- 
late, usually opposite, rarely whorled in 3’s, firm, 
entire, or the lower sinuate-dentate or sometimes 
pinnatifid, 3-6’ long, short-petioled, the lobes ob- 
tuse; the upper much smaller and sessile, passing 
into the bracts of the raceme; pedicels stout, usu- 
ally shorter than the calyx even in fruit; calyx- 
lobes lonceolate, entire, about as long as the tube; 
corolla 13’-2’ long, ‘glabrous outside, its tube 
much expanded above; capsule 8’—10” long, pu- 
bescent, twice as long as the calyx. 

In dry woods and thickets, Maine to Ontario and 


Wisconsin, south to southern New York, Georgia 
and Mississippi. Yellow foxglove. July—Aug. 


GENuS 28. FIGWORT FAMILY. 207 


3. Dasystoma serrata (Benth.) Small. 
Serrate False Foxglove. Fig. 3814. 


D. Drummondii serrata Benth. in DC. Prodr. 10: 
Bat, Lodo. 


G. grandiflora integriuscula A. Gray, Syn. FI. N. 
IN, Ge Bos 1A 


Gerardia grandiflora serrata Robinson, in A. Gray, 
Man. Ed. 7, 730. 1908. 


Dasystoma serrata Small, Bull. Torr. Club 28: 451. 
1901. 


Perennial, grayish puberulent or finely pubes- 
cent; stems 1°-32° tall, often widely branched. 
Leaf-blades of the lower part of the stem pin- 
natifid and their segments entire, merely serrate 
and much smaller above; calyx-tube 23-3” 
broad during anthesis; calyx-lobes linear-lanceo- 
late to lanceolate, entire; corolla 1-12’ long; 
capsules ovoid or globose-ovoid, 5’’-73” long, 
short-beaked. 


In dry soil, Missouri to Louisiana and Texas. 
July—Sept. 


4. Dasystoma grandiflora (Benth.) Wood. 
Western False Foxglove. Fig. 3815. 


Gerardia grandiflora Benth. Comp. Bot. Mag. 1: 206. 
1835. 

D. Drummondii Benth. in DC. Prodr. 10: 520. 1846. 

D. grandiflora Wood, Bot. & Flor. 231. 1873. 


Perennial, cinereous-puberulent and roughish; 
stem much branched, very leafy to the top, 2°-3° 
high, the branches ascending or spreading. Leaves 
short-petioled, thin, ovate or ovate-lanceolate: in 
outline, all pinnatifid or deeply incised, 2-4’ long, 
the lobes acute or obtuse, serrate, or nearly entire; 
upper leaves smaller, sessile; pedicels, even in fruit, 
shorter than the calyx; calyx-lobes oblong or ovate, 
dentate or entire, about as long as the tube; corolla 
14-2’ long, glabrous without, its tube much ex- 
panded above. 


In dry woods and thickets, Minnesota and Wisconsin 
to Tennessee, Kansas and Texas. July—Aug. 


5. Dasystoma laevigata Raf. Entire- 
leaved False Foxglove. Fig. 3816. 


Gerardia laevigata Raf. Ann. Nat. 13. 1820. 

Dasystoma quercifolia var. integrifolia Benth. in 
We sProdr. 10:/520). redo. 

Dasystoma laevigata Raf.; Chapm. Fl. S. States, 
Ed. 2, 636. 1883. 


Perennial, glabrous or very nearly so, not 
glaucous; stem strict, simple, or sparingly 
branched, 1°-3° high, the branches ascending. 
Leaves usually all petioled, lanceolate or ovate- 
lanceolate, 13’-4’ long, entire, or the lowest 
dentate or incised; pedicels shorter than the 
calyx, or 1n fruit longer; calyx-lobes ovate- 
lanceolate, equalling or shorter than the tube; 
corolla glabrous without, 1-14’ long, the limb 
fully as broad, the tube much expanded above; 
capsule glabrous, twice as long as the calyx. 


In dry thickets, Pennsylvania to Michigan, 
Georgia and Missouri. July—Aug. 


208 SCROPHULARIACEAE. Vo. IIT 


6. Dasystoma virginica (L.) Britton. Smooth False Foxglove. Fig. 3817. 


Rhinanthus virginicus L. Sp. Pl. 603. 1753. 

Gerardia quercifolia Pursh, Fl. Am. Sept. 423. pil. I9. 
1814. 

D. quercifolia Benth. in DC. Prodr. 10: 520. 1846. 

D. virginica Britton, Mem. Torr. Club 5: 295. 18094 


Perennial, glabrous and glaucous; stem strict, 
rather stout, usually branched, 3°-6° high, the 
branches ascending. Leaves usually all petioled, 
ovate or ovate-lanceolate in outline, the lower 
I-2-pinnatifid, 4’-6’ long, the upper pinnatifid or 
deeply incised, the lobes lanceolate or oblong, 
acute, entire, or dentate; fruiting pedicels longer 
than the calyx; calyx-lobes ovate or ovate-lanceo- 
late, acute, entire, about equalling the tube; co- 
rolla 14-2’ long, glabrous outside, its tube not 
widely expanded above; capsule glabrous, twice 
as long as the calyx. 


In dry or moist woods, Maine to Minnesota, south 
to Florida and Illinois. Golden-oak. July—Sept. 


Dasystoma calycoésa Mackenzie & Bush has been 
| distinguished from D. virginica by its more finely 

LS es divided leaves, its elongate spreading branches -and 
its long and narrow calyx-lobes; it occurs in Missouri and Arkansas. 


29. AGALINIS Raf. N. Fl. 2: 61. 1836. 


Erect branching annual or perennial herbs, some South American species shrubby, mainly 
with opposite and sessile leaves. Flowers showy, usually large, purple, violet, yellow, red, or 
rarely white, racemose, or paniculate, or solitary and axillary. Calyx campanulate, 5-toothed, 
or 5-lobed. Corolla somewhat irregular, campanulate, or funnelform, the tube broad, short, 
or elongated, the limb 5-lobed, slightly 2-lipped, the lower lobes exterior in the bud. Stamens 
4, didynamous, included; filaments more or less pubescent; anthers 2-celled, their sacs obtuse 
or mucronate at the base, style filiform. Capsule globose or ovoid, loculicidally dehiscent, 
many-seeded. Seeds numerous, mostly angled. [Greek, remarkable flax.] 


About 45 species, natives of America. Besides the following, some 10 others occur in the 
southern United States. Most of the species blacken in drying. Type species: Agalinis palustris 
Raf. (Gerardia purpurea L.) The generic name Gerardia (Plumier) L., used for these plants 
in our first edition, is typified by Gerardia tuberosa L. (Stenandrim tuberosum (L.) Britton, of 
the Acanthaceae). 


Pedicels in flower shorter than the calyx, or but 1-2 times as long. 
Corolla 10”—-13” long. 


Calyx-teeth minute; root perennial. 1. A. linifolia. 
Calyx-teeth triangular, lanceolate or oblong, acute; annuals. 
Leaves very scabrous, filiform; capsule oblong. 2. A. aspera. 
Leaves slightly scabrous, linear; capsule globose. 
Calyx-teeth as long as the tube or longer. 3. A. heterophylla. 


Calyx-teeth shorter than the tube. 
Stem smooth or nearly so; branches spreading; ‘eaves rarely clustered in axils. 


4. A. purpurea. 
Stem scabrous; branches virgate; leaves much clustered in axils. 
5. A. fasciculata. 
Corolla 5”-8” long. 
Calyx-teeth triangular-subulate, acute. 6. A. paupercula. 
Calyx-teeth broad, short, obtuse. 7. A. maritima. 
Pedicels in flower 2—6 times as long as the calyx. 
Leaves flat, linear, spreading or ascending; capsule globose. 
Leaves %4”—-1” wide, %4’-11%4’ long; pedicels spreading. 8. A. tenuifolia. 
Leaves 1”-2” wide, 11%4’—3’ long; pedicels ascending. g. A. Besseyana. 
Leaves subulate, or filiform, the margins often revolute. 
Corolla-lobes obcordate or deeply emarginate. 10. A. parvifolia. 
Corolla-lobes rounded or slightly emarginate. 
Corolla 5”—7” long; pedicels erect-ascending. 11. A. Skinneriana. 
Corolla 8’—10” long; pedicels spreading-ascending. 
Calyx-teeth broadly triangular. 12, A.setacea. — 
Calyx-teeth linear-subulate. 13. A. Gattingeri. 


GENUS 29 FIGWORT FAMILY. 


1. Agalinis linifolia (Nutt.) Britton Flax-leaved 
Agalinis. Fig. 3818. 


Gerardia linifolia Nutt. Gen. 2: 47. 1818 


Perennial, glabrous and smooth; stem branched, 2°-3° 
high. Leaves narrowly linear, rather thick, 1-2’ long, 
1-14” wide, erect the upper much smaller and subulate; 
pedicels erect, in flower equalling or a little longer than 
the calyx, longer in fruit; calyx campanulate, truncate, its 
teeth minute; corolla purple, about 1’ long, narrower than 
that of the two following species, villous within, the lobes 
ciliate; filaments and anthers densely villous; anther-sacs 
mucronate at the base; capsule globose, 2’-3” in diameter, 
but little longer than the calyx. 


In moist pine barrens, Delaware to Florida. Aug.—Sept. 


2. Agalinis aspera (Dougl.) Britton. Rough 
Purple Agalinis. Fig 3819. 
G aspera Dougl.; Benth. in DC. Prodr. 10: 517. 1846. 


Annual, 1°-2° high, hispidulous-scabrous with rough 
stiff short whitish hairs, branched, the branches nearly 
erect. Leaves narrowly linear, 1-13’ long, less than 1” 
wide, erect or ascending; pedicels equalling or becom- 
ing a little longer than the turbinate calyx; calyx-teeth 
triangular-ovate or triangular-lanceolate, acute, one- 
fourth to one-third as long as the tube; corolla deep 
, purple, about 1’ long, nearly or quite glabrous within, 
# very pubescent without, the lobes ciliate; filaments vil- 
7, lous; anthers all alike, obtuse at the base; capsule ob- 
f long, 3-4” high, considerably longer than the calyx. 


On dry plains and prairies, Indiana to North Dakota, 
Missouri, Kansas and Arkansas. Recorded from Colorado. 


Aug.—Oct. 
QF 
WI 


3. Agalinis heterophylla (Nutt.) Small. 
Prairie Agalinis. Fig. 3820. 


Gerardia heterophylla Nutt. Trans. Am. Phil. Soc. 5: 
180. 1837. 


Gerardig crustata Greene, Leaflets 2: 108. 1910. 


Annual, more or less scabrous; stems 62° tall, 
branching above, the branches ascending. Leaves 
linear, or narrowly linear-lanceolate below, 4’-2’ long, 
rarely 2” broad, acute, erect or erect-ascending, de- 
cidedly scabrous along the margins; pedicels shorter 
than the calyx, enlarged upward; calyx campanulate, 
3-4” long, its teeth lanceolate or subulate-lanceolate, 
about as long as the tube; corolla rose-purple, about 
¥ long; capsules globose-oblong or globose-ovoid, 
23’’-3”” long. 


On. prairies and in wet woods, Missouri and Arkansas 
to Texas. Aug.—Oct. 


14 


210 SCROPHULARIACEAE. Vor. ELT, 


4. Agalinis purpurea (L.) Britton. Large 
Purple Agalinis. Fig. 3821. 

Gerardia purpurea L. Sp. Pl. 610. 1753. 
G. racemulosa Pennel, Torreya 11: 15. 1911. 

Annual, glabrous, smooth, or roughish; stem 
slender, branched, 1°-23° high, the branches 
spreading. Leaves narrowly linear, usually widely 
spreading, 1-13’ long, about 1” wide, rarely with 
smaller ones fascicled in their axils; flowers race- 
mose on the branches, purple (rarely white), 
about 1’ long and broad; pedicels shorter than 
or but little longer than the campanulate calyx, 
even in fruit; calyx-teeth triangular-lanceolate or 
ovate-oblong, acute, one-third to one-half the 
length of the tube; corolla much expanded above, 
villous or nearly glabrous within, pubescent with- 
out, the lobes ciliolate; anthers all alike, the sacs 
mucronulate at the base; filaments villous; cap- 
sule globose, 2”-3” in diameter, longer than the 
calyx. ( 

In moist fields and meadows, Maine to Florida, 
Indiana, Wisconsin, Missouri and Texas. Aug.—Oct. 


5. Agalinis fasciculata (Ell.) Raf. Fascicled 
Agalinis. Fig. 3822. 


Gerardia fasciculata Ell. Bot. S. C. & Ga. 2: 115. 
Gerardia fasciculata Ell. Bot. S. C. & Ga. 2: 115. 1824. 


Annual, finely pubescent and scabrous; stems 12°-4° tall, 
often puberulent or sparingly hispidulous, fastigiately branch- 
ed. Leaves numerous, the larger ones with conspicuous clus- 
ters of smaller ones in their axils, linear, mostly less than 1” 
wide, acute, very scabrous; flowers racemose, often numer- 
ous, purple, about 1’ long; pedicel shorter than the calyx; 
calyx-tube campanulate, 14”-1}” high, the teeth usually mi- 
nute, often less than 2?” long; corolla abruptly expanded 
above the short tube, minutely pubescent without, copiously 
pubescent within, the lobes ciliolate; capsule 2’-23” in diam- 
eter, becoming longer than the calyx. 


In marshes or sometimes in dry soil, Virginia to Florida and 
Texas. Aug.—Oct. 


6. Agalinis paupércula (A Gray) Britton. 
Small-flowered Agalinis. Fig. 3823. - 


Gerardia purpurea var. paupercula A. Gray, Syn. 2: 
Part 1, 293. 1878, 

Gerardia intermedia Porter; A. Gray, loc..cit. As 
synonym. 1878. 

G. paupercula Britton, Mem. Torr. Club 5: 295. 1894. 


Annual, glabrous and smooth or very nearly 
so; stems strict, branched above, 6-18’ high, the 
branches nearly erect. Leaves narrowly linear, 
3-1’ long, 3”-1” wide, spreading or ascending; 
pedicels equalling the calyx, or longer in fruit; 
calyx campanulate, its teeth about one-half the 
length of the tube, triangular-lanceolate, acute, or 
acuminate; corolla 6’—10” long, puberulent, rose 
purple, its limb about as broad, somewhat villous 
in the throat within, the lobes ciliate; stamens 
very villous; anther-sacs mucronulate at the base; 
capsule globose-oblong, 3” high, longer than the 
calyx. 

In bogs and low meadows, Nova Scotia to New 


Jersey, Tennessee, Manitoba and Wisconsin; eppar- 
ently also in Georgia and South Carolina. July—Sept. 


GENUS 20. FIGWORT FAMILY. 2G 


7. Agalinis maritima Raf. Sea-side or Salt- 
marsh Agalinis. Fig. 3824. 


Gerardia maritima Raf. Med. Rep. (I1.) 5: 361. 1808. 
A. maritima Raf. New FI. N. A. 2: 62. 1836. 


Annual, smooth and glabrous, fleshy; stem erect, 
usually branched, 4-16’ high, the branches ascend- 
ing. Leaves linear, thick, 3’-1’ long, 1” wide or less, 
obtuse, spreading, the upermost very small and subu- 
late, scarcely longer than the pedicels; pedicels in 
flower shorter than or equalling the calyx, about 
twice as long in fruit; calyx-teeth broad, short, ob- 
tuse; corolla rose purple, 5”—-9” long, glabrous, the 
limb about as broad; anther-sacs mucronulate at the 
base; capsule globose-oblong, 2-3” high. 

In salt marshes, Maine to Florida. July-Aug. The 


southern plant has larger flowers than the northern. 
Flowers before the upland species. ct 


8. Agalinis tenuifolia (Vahl.) Raf. Slender Agalinis. Fig. 3825. 
Gerardia tenuifolia Vahl, Symb. Bot. 3: 79. 1794. 


A. tenuifolia Raf. New Fl. N. A. 2: 64. 1836. 


Annual, glabrous; stem very slender, panicu- 
lately branched, 6-24’ high, the branches spread- 
ing or ascending. Leaves very narrowly linear, 
flat, acute, 4’-17’ long, 2’-1” wide, spreading; 
pedicels mostly equalling or longer than the flow- 
ers; calyx campanulate, its teeth very short, 
pointed; corolla light purple, spotted, rarely 
white, 6-9” long, vertically compressed when 
fully expanded, minutely puberulent, or glabrous; 
anther-sacs mucronate at the base; capsule glo- 
bose or slightly obovoid, 2”-23” in diameter, 
longer than the calyx. 


In dry woods and thickets, Quebec to Georgia, 
west to western Ontario, Kansas and Texas. Aug.— 


Oct. 


g. Agalinis Besseyana Britton. Bessey’s 
Agalinis. Fig. 3826. 


Gerardia tenuifolia var. macrophylla Benth. Comp. 
Bot. Mag. 1: 209. 1835. Not G. macrophylla Benth. 


Gerardia Besseyana Britton, Mem. Torr. Club 5: 295. 
1894. 


Annual; stem glabrous, rather stout, strict, 
branched, 1°-2° high, the branches ascending or 
nearly erect. Leaves linear, 1’-2’ long, 1-2” 
wide, scabrous, acute, ascending; pedicels ascerd- 
ing, longer than the flowers; calyx campanulate, 
its teeth triangular-subulate, one-third to one- 
half the length of the tube; corolla purple, 5-6” 
long; capsule globose, 2’-3” in diameter, exceed- 
ing the calyx. 

On dry hills and prairies, Ontario to South Da- 
kota, Colorado, Louisiana and Kansas. Recorded 
east to Connecticut. July—Sept. 


212 SCROPHULARIACEAE. Vo. III. 


to. Agalinis parvifolia (Chapm.) Small. 
Ten-lobed Agalinis. Fig. 3827. 


Gerardia parvifolia Chapm. Fl. S. U. S. 300. 1860. 
Gerardia decemloba Greene, Pittonia 4: 51. 1899. 


Annual, pale green, usually roughish; stems 823° 
tall, simple or branched above, striate-angled, the 
J branches mostly ascending. Leaves few, remote, 

a linear-spatulate, to narrowly linear, more or less 

4 Cl revolute, 22’-10” long or rarely slightly longer, 

scabrous-pubescent above; pedicel exceeding the ca- 

lyx ; calyx campanulate or turbinate-campanulate, 1’— 

134” high, the lobes minute, triangular; corolla light 

rose-colored, about 3’ long, or less, the lobes obcor- 

date or deeply emarginate; capsules globose-ovoid, 
or oval, 13’-2” long, apiculate. 


In dry sandy soil or on banks, Massachusetts to Flor- 
ida and Louisiana. Aug.—Oct. 


11. Agalinis Skinneriana (Wood) Britton. Skinner’s Agalinis. Fig. 3828. 
Gerardia Skinneriana Wood, Classbook 408. 1847. a 
?G. tenuifolia asperula A. Gray, Bot. Gaz. 4: 153. 1879. . 8) 
Annual, roughish; stem strict, striate, branched, (#) oe \ CF IS 
or sometimes simple, 6-18’ high, very slender, the x \y \\ 
branches erect or ascending. Leaves setaceous, as- Vi, | < 
cending or commonly erect and appressed, 2’—-1’ long, \ 
3” wide or less, the uppermost minute; pedicels WV 
longer than the calyx, scarcely longer than the flow- ZB 
ers, 2-4 times the length of the capsule; calyx-teeth 
minute; corolla light purple, 5’-6” long and about 


as broad, glabrous without, its lobes ciliolate; capsule | 
oblong, 2”-3” high, considerably longer than the 
calyx. 


In dry sandy woods and thickets, Indiana to Minne- 
sota and Kansas. Recorded from Ontario. Aug.—Oct. 

Gerardia viridis Small, differing by more widely 
spreading pedicels and narrow calyx-teeth often one- 
half as long as the tube, inhabits the western Gulf 
States and is recorded as extending northward into Mis- 
souri. 


\ 


12. Agalinis setacea (Walt.) Raf. Thread- 
leaved Agalinis. Fig. 3820. 


Gerardia setacea Walt. Fl. Car. 170. 1788. 
A. setacea Raf. New FI. N. A. 2: 64. 1836. 
G. Holmiana Greene, Pittonia 4: 52. 1899. 


Annual, smooth or slightly scabrous; stems 1°-2° 
tall, loosely branching, the branches mostly ascend- 
ing. Leaves rather numerous, mostly opposite, linear- 
filiform or setaceous-filiform, +12’ long, scabrous 
or nearly smooth; pedicels very slender, #’-12’ long; 
calyx campanulate, 13’-2” long, the lobes broadly 
triangular, about 4 as long as the tube; corolla rose 
purple, 5’-8” long or rarely larger; capsules subglo- 


bose or globose-oval, 2-23” in diameter. 


In dry soil or pine barrens, New Jersey to Florida and 
Texas. Aug.—Oct. 


GENUS 29. FIGWORT FAMILY. 213 
Q 

13. Agalinis Gattingeri Small. Gattinger’s f 

Agalinis. Fig. 3830. 


Gerardia Gattingeri Small, Fl. SE. U. S. 1078. 1903. 


Annual, smooth or slightly scabrous; stem 82° 
tall, wiry, with smooth very slender long branches. 
Leaves numerous, linear-filiform or almost fili- 
form, 4’-1?’ long, acute, smooth or slightly rough- 
ened; pedicels spreading or ascending, filiform, 
3’-1’ long, less than twice the length of the leaves; 
calyx campanulate, its teeth subulate or linear- 
subulate, much shorter than the tube; corolla rose 
purple, 8’-10” long; capsules subglobose, 12’—2” in 
diameter. 


In dry soil or woods, Wisconsin and Iowa to Ten- 
nessee and Texas. Aug.—Oct. 


go. OTOPHYLLA Benth. in DC. Prodr. 10: 512. 1846. 


Annual caulescent herbs, with hirsute-pubescent foliage. Leaves opposite; blades entire 
or pinnately divided, all or some of them auricled at the base, sessile. Flowers in terminal 
spikes. Calyx of 5 partially united sepals. Corolla purple or rarely white; tube broadly 
dilated at the throat; lobes spreading. Stamens 4, didynamous, included; filaments glabrous 
or nearly so; anthers awnless, those of the shorter stamens much smaller than those of the 
longer. Style slender. Stigma entire. Capsule oval or globose-oval, often minutely pointed. 
Seeds angled. [Greek, meaning lobed leaf.] 


Two species in eastern North America. Type species: Gerardia auriculata Michx. 


Leaf-blades, or some of them, with 2 auricles at the base; corolla 8’—10” long; capsules over 


ong. : [ 1. O. auriculata. 
Leaf-blades parted into 3-7 linear segments; corolla 1/114’ long; capsules about %4’ long. 


2. O. densiflora. 


1. Otophylla auriculata (Michx.) Small. Auricled Gerardia. Fig. 3831. 


G. auriculata Michx. Fl. Bor. Am. 2: 20. 1803. 
Otophylla auriculata Small, Fl. SE. U. S. 1075. 1903. 


Annual, scabrous; stem slender, simple, or 
branched above, hirsute, 1°-2° high. Leaves lan- 
ceolate or ovate-lanceolate, acuminate at the apex, 
sessile, mostly rounded and 2-lobed at the base, 
or quite entire, 1-2’ long, the basal lobes oblong 
or lanceolate, obtusish, short; flowers solitary in 
the upper axils, sessile, purple, 8’-10” long; ca- 
lyx 5-cleft, its lobes lanceolate, acute, slightly 
unequal, as long as or longer than the tube; co- 
rolla densely puberulent outside, glabrous within; 
filaments glabrous; anther-sacs obtuse at the 
base; capsule oval-oblong, about }# high, a little 
shorter than the calyx. 


In moist open soil, Pennsylvania to Illinois, Min- 
nesota, North Carolina and Kansas. Adventive at 
Woodbridge, N. J. July—Sept. 


214 SCROPHULARIACEAE. Vo. III. 


2. Otophylla densiflora (Benth.) Small. 
Cut-leaved Gerardia. Fig. 3832. 


G. densiflora Benth. Comp. Bot. Mag. 1: 206. 1835. 
Otophylla densiflora Small, Fl. SE. U. S. 1075. 1903. 


Annual, scabrous and short-hispid; stems stiff, 
erect, branched, or simple, 1°-23° high, very leafy. 
Leaves sessile, ovate in outline, ascending, about 
1’ long, pinnately parted nearly to the midvein 
into 3-7 narrowly linear acute rigid segments less 
than 1” wide; flowers 17-11’ long, rose purple, 
sessile in the upper axils; calyx 5-cleft, its lobes 
linear, acuminate, ciliate, about as long as the 
tube; corolla glabrous both outside and within; 
filaments glabrous or villous; capsule about ? 
high, shorter than the calyx. 


On dry prairies, Kansas to Texas. Aug.—Oct. 


31. CASTILLEJA Mutis; L. f. Suppl. 
A7:. D780. 


Herbs, parasitic on the roots of other plants, with alternate leaves, and red yellow purple 
or white flowers, in dense leafy-bracted spikes, the bracts often brightly colored and larger 
than the flowers. Calyx tubular, laterally compressed, cleft at the summit on the upper side, 
or also on the lower, the lobes entire or 2-toothed. Corolla very irregular, its tube not longer 
than the calyx, its limb 2-lipped; upper lip (galea) arched, elongated, concave or keeled. 
laterally compressed, entire, enclosing the 4 didynamous stamens; lower lip short, 3-lobed. 
Anther-sacs oblong or linear, unequal, the outer one attached to the filament by its middle, 
the inner one pendulous from its apex. Style filiform; stigma entire or 2-lobed. Capsule 
ovoid or oblong, loculicidally dehiscent, many-seeded. Seeds reticulated. [Named for Cas- 
tillejo, a Spanish botanist. ] 


About 50 species, mostly natives of the New World. In addition to the following, about 30 
others occur in the western parts of North America. Type species: Castilleja fissifolia L. f. 


* Plants villous-pubescent. 
Bracts broad, dilated, lobed, or entire. 


Stem leaves deeply and irregularly cleft into narrow segments. 1. C. coccinea. 
Leaves linear-lanceolate, entire, or rarely with a few lobes. 2. C. indivisa. 
Bracts linear or linear-lanceolate, entire. 3. C. minor. 
** Plants glabrous, woolly at the summit, or cinereous-puberulent. 
Glabrous, or tomentose at the summit; leaves lanceolate, mostly entire. 4. C. acuminata. 
Cinereous-puberulent, pale; stem leaves cleft. 5. C. sessilifiora. 


1. Castilleja coccinea (L.) Spreng. Scarlet 
Painted-cup. Indian Paint-brush or 
Pink. Prairie-fire. Fig. 3833. 


Bartsia coccinea L. Sp. Pl. 602. 1753. 
Castilleja coccinea Spreng. Syst. 2: 775. 1825. 


Annual or biennial, villous-pubescent; stem 
rather slender, simple, or with few erect branches, 
1°-2° high. Leaves sessile, parallel-veined, the 
basal oblong, obovate, or linear, tufted, mostly 
entire, 1’-3’ long, those of the stem deeply 3-5- 
cleft into linear obtusish segments, the bracts 
broader and shorter, 3-5-lobed or cleft, bright 
red or scarlet, conspicuous; flowers sessile, 10’— 
12” long, usually not exceeding the bracts; calyx 
cleft both above and below into 2 dilated entire 
or retuse oblong and obtuse lobes, sometimes 
scarlet; corolla greenish-yellow, its tube shorter 
than the calyx, its upper lip much longer than 
the lower; capsule oblong, acute, 5-6” long. 


In meadows and moist thickets, Maine and Ontario 
to Manitoba, North Carolina, Tennessee, Kansas and 
Texas. Ascends to 4000 ft. in Virginia. Bracts and 
calyx rarely yellow. Red indians. Election-posies. 
Wickawee. Bloody-warrior. Nose-bleed. May-July. 


GENUS 31. FIGWORT FAMILY. 


2. Castilleja indivisa Engelm. Entire-leaved 
Painted-cup. Fig. 3834. 


Castilleja indivisa Engelm.; Engelm. & Gray, Bost. 
Jqurn. Nat. Hist. 5: 255. 1845. 


Winter-annual; stem villous-pubescent, usually 
simple, 8-18’ high. Leaves sessile, parallel-veined, 
linear to linear-lanceolate, 1’—4’ long, 13’-3” wide, 
entire, or rarely with 2-4 lateral lobes; no tuft of 
basal leaves; bracts dilated, obovate to spatulate, 
bright red; flowers sessile, about 1’ long or less, 
not longer than the bracts; calyx cleft as in the 
preceding species, and corolla similar. 


In sandy soil, Kansas to Texas. Spring. 


3. Castilleja minor A. Gray. Small-flow- 
ered Painted-cup. Fig. 3835. 


Castilleja affinis var. minor A. Gray, Bot. Mex. 

Bound. Surv. 119. 1859. 

Castilieja minor A. Gray, in Brew. & Wats. Bot. Cal. 

L573. 1076. 

Annual, villous-pubescent; stem slender, strict, 
simple, or with I or 2 erect branches, 1°-23° 
high. Leaves all linear-lanceolate and entire, 
parallel-veined, sessile, acuminate, 2’-3’ long, the 
bracts similar, smaller, red or red-tipped, very 
narrow, equalling or longer than the short-pedi- 
celled flowers; calyx green, cleft on both sides to 
about the middle, the lobes lanceolate, acute, en- 
tire, or 2-toothed; corolla yellow, 6’—-10” long, its 
upper lip much longer than the small lower one; 
capsule oblong, acute, 6-8” long. 


In moist soil, Nebraska to New Mexico, west to 
Nevada and Arizona. Indian-pink. May-July. 


4. Castilleja acuminata (Pursh) Spreng. 
Lance-leaved Painted-cup. Fig. 3836. 


Bartsia acuminata Pursh, Fl. Am. Sept. 429. 1814. 

Castilleja acuminata Spreng. Syst. 2: 775. 1825. 

C. septentrionalis Lindl. Bot. Reg. pl. 925. 1825. 

Castilleja pallida var. septentrionalis A. Gray, in 
Brew. & Wats. Bot. Cal. 1: 575. 1876. 


Perennial, glabrous or loosely tomentose above ; 
stems slender, commonly clustered, 62° high, 
usually simple. Leaves sessile, 3-5-nerved, mostly 
quite entire, the lower linear, the upper lanceo- 
late, acuminate or acute at the apex, somewhat 
narrowed at the base, 2-4’ long; bracts oblong, 
oval, or obovate, obtuse. dentate, or entire, yel- 
lowish, greenish-white or purple, as long as the 
sessile flowers; calyx cleft on both sides to about 
the middle, the lobes lanceolate, usually again 
2-cleft; corolla 6’-8” long, its upper lip 2-4 times 
as long as the lower; capsule oblong, 6’-8” high. 


In moist soil, Newfoundland and Labrador to 
Hudson Bay, the mountains of New England, On- 
tario and Minnesota. Plants previously referred to 
this species from farther west are now regarded as 
distinct from it. June-Aug. Pale painted-cup. 


SCROPHULARIACEAE. Vor. ITI. 


5. Castilleja sessiliflora Pursh. Downy 
Painted-cup. Fig. 3837. 


Castilleja sessiliflora Pursh, Am. Sept. 738. 1814. 


Perennial, cinereous-puberulent all over; 
stems stout, simple, or branched from near 
the base, 6-15’ high, densely leafy. Leaves 
sessile, 1-2’ long, the lowest commonly linear, 
obtuse and entire, the others laciniate into 
narrow, entire or cleft segments; bracts green, 
similar to the upper leaves, shorter than the 
sessile flowers; calyx deeper cleft on the lower 
side than on the upper, its lobes linear-lanceo- 
late, acute; corolla yellowish, 14’ long, the 
upper lip about twice as long as the lower, the 
lobes of the latter linear; capsule oblong- 
lanceolate, acute, 6-8” long. 


On dry prairies, Illinois to Manitoba, Sas- 
katchewan, Nebraska, Wyoming and Texas. May-— 
July. 


32. ORTHOCARPUS Nutt. Gen. 2:56. 1818. 


Annual or rarely perennial herbs, mostly with alternate leaves, and yellow white or pur- 
plish flowers, in bracted usually dense spikes, the bracts sometimes brightly colored. Calyx 
tubular or tubular-campanulate, 4-cleft, or sometimes split down both sides. Corolla very 
irregular, the tube slender, the limb 2-lipped; upper lip little if any longer than the 3-lobed 
I-3-saccate lower one. Stamens 4, didynamous, ascending under the upper lip; anther-sacs 
dissimilar, the outer one affixed by its middle, the inner pendulous from its upper end, com- 
monly smaller. Style filiform; stigma entire. Capsule oblong, loculicidally dehiscent, many- 
seeded. Seeds reticulated. [Greek, erect-fruit.] 


About 30 species, natives of America, mostly of the western United States, 1 or 2 Andean. 
Type species: Orthocarpus luteus Nutt. “e 


1. Orthocarpus luteus Nutt. Yellow Ortho- 
carpus. Fig. 3838. 
Orthocarpus luteus Nutt. Gen. 2: 57. 1818. 


Annual, rough-pubescent or puberulent; stem strict, 
erect, branched above, or simple, 6-18 high, densely 
leafy. Leaves erect or ascending, linear or lanceolate, 
entire, or sometimes 3-cleft, 1-12’ long, 1’-2” wide, 
sessile, long-acuminate; bracts of the dense spike lan- 
ceolate, broader and shorter than the leaves, entire or 
3-cleft, acute, green, mostly longer than the flowers; 
flowers bright yellow, 4-5” long; calyx-teeth acute, 
shorter than the tube; corolla about twice as long as 
the calyx, puberulent without, its upper lip ovate, ob- 
tuse, about as long as the saccate 3-toothed lower one; 
capsule about as long as the calyx-tube. 


On dry plains and prairies, Manitoba to Minnesota and 
Brace: west to British Columbia and California. July— 
ept. 


33. SCHWALBEA [Gronov.] L. Sp. Pl. 606. 1753. 


A perennial erect finely pubescent and minutely glandular, simple or sparingly branched, 
leafy herb, with sessile entire 3-nerved leaves, and rather large yellowish-purple flowers in a 
terminal bracted spike. Calyx tubular, somewhat oblique, 10-12-ribbed, 2-bracteolate at the 
base, 5-toothed, the upper tooth much the smallest, the 2 lower ones partly connate; corolla 
very irregular, the tube cylindric, the limb 2-lipped; upper lip arched, concave, entire; lower 
lip somewhat shorter, 3-lobed, 2-plaited. Stamens 4, didynamous, ascending within the upper 
lip of the corolla; anther-sacs equal. Style filiform. Capsule oblong, many-seeded. Seeds 
linear, with a loose reticulated testa. [Named for C. G. Schwalbe, of Holland, who wrote 
(1719) on Farther India.] 


A monotypic genus of eastern North America. 


GENUS 33. FIGWORT FAMILY. 217 


1. Schwalbea americana L. Chaff-seed. 
Fig. 3839. 


Schwalbea americana L. Sp. Pl. 606. 1753. 


Stem slender, strict, 1°-2° high. Leaves ob- 
long or ovate-oblong, entire, acute at both ends, 
1-13’ long, 2”-8” wide, the upper gradually 
smaller and passing into the bracts of the rather 
loose spike; flowers very nearly sessile, 1-12’ 
long, longer than the bracts; bractlets at the base 
of the calyx linear, shorter than its tube; corolla- 
tube slightly exceeding the lower lobes. of the 
calyx, these connate to near their apices; capsule 
enclosed by the calyx. 


In wet sandy soil, eastern Massachusetts to Flor- 
ida and Louisiana, near the coast. May-July. 


VAAN 
34. EUPHRASIA [Tourn.] L. Sp. Pl. 604. 1753. 


Annual or perennial low mostly branched herbs, parasitic on other plants, with opposite 
dentate or incised leaves, and small blue purplish yellow or white often variegated flowers in 
terminal leafy-bracted spikes. Calyx not bracteolate at the base, campanulate or tubular, 
4-cleft (rarely 5-cleft with one of the lobes much smaller than the others). Corolla very 
irregular, 2-lipped, the upper lip erect, scarcely concave, 2-lobed, its margins recurved; lower 
lip larger, 3-lobed, spreading, its lobes either emarginate or obtuse. Stamens 4, didynamous, 
ascending under the upper lip of the corolla; anther-sacs equal and parallel, mucronate at the 
base. Capsule oblong, loculicidally dehiscent, many-seeded. Seeds oblong, longitudinally 
ribbed. [Greek, delight. ] 

About 110 species, natives of temperate and cold regions of both the northern and southern 
hemispheres. Besides the following, another occurs in northwestern North America. Type species: 
Euphrasia officinalis L. 


Flowers 2%4”-4” long. 


Leaves markedly pubescent on both sides. 1. E. arctica. 

Leaves glabrate or sparingly pubescent. 2. E. americana. 
Flowers 11%4’—2” long. 

Stem 3’-20’ long; flowers racemose-spicate. 3. E. Randit. 

Stem 1/2’ high; flowers subcapitate. 4. E. Oakesii. 


1. Euphrasia arctica Lange. Glandular Eye- 
bright. Fig. 3840. 


Euphrasia arctica Lange, Bot. Tidskr. 4: 47. 1870. 


Euphrasia latifolia Pursh, Fl. Am. Sept. 430. 1814. 
Not Willd. 


Annual; stem erect, simple, or with a few erect 
branches from near the base, pubescent with crisped 
hairs, 2’-7’ high. Leaves ovate to obovate, obtuse, 
pubescent on both sides, 2-5-toothed on each margin, 
the teeth sharp or blunt; spike 1-4’ long, the bracts 
imbricated, at least above, broadly oval to orbicular, 
cuneate at the base, sharply toothed, glandular- 
pubescent beneath or also on the margins; calyx- 
teeth acute; corolla 3’-4” long, lilac, or variegated, 
the lobes of its lower lip nearly parallel; capsule 
oblong-elliptic, ciliate on the margins, and more or 
less pubescent or pilose, about as long as the calyx. 


Bluffs and slopes, Greenland to Hudson Bay, New 
Brunswick, Maine and Minnesota. Has been erro- 
neously referred to E. hirtella Jordan. Summer. 


SCROPHULARIACEAE. Vo. III. 


2. Euphrasia americana Wettst. Hairy Eye- 
bright. Eyebright. Fig. 3841. 

Euphrasia americana Wettst. Mon. Euph. 127. 18096. 
E. canadensis Townsend, Journ. Bot. 36: 1. 1808. 

Annual, stem pubescent with crisped hairs, often 
at length much branched, 4-10’ high. Leaves gla- 
brous, or sparingly pubescent, ovate to oval, obtuse or 
acutish at the apex, 3-5-toothed on each side, 3-6” 
long, 13’—42” wide, the teeth acute; bracts cuneate 
or obtuse at the base, dentate, pubescent, not glandu- 
lar, slightly shorter than the flowers; calyx-teeth 
lanceolate, acuminate; corolla 3-4” long, purplish 
or nearly white, the somewhat spreading lobes of 
its lower lip emarginate; capsule narrow, cuneate, 
2”-3” high, about equalling the calyx. 


In fields and on hills, Maine and New Hampshire to 
New Rrunswick and Newfoundland. Summer. 


Euphrasia officinalis L. is not known from North 
America. 


3. Euphrasia Randii Robinson. Rand’s Eye- 
bright. Fig. 3842. 


Euphrasia Randii Robinson, Rhodora 3: 273. 1901. 
E. Randii var. (?) Farlowii Robinson, loc. cit. 274. 1901. 


Annual; stem simple or branched, erect or decumbent, 
3-20’ long, purplish, covered with whitish hairs. Leaves 
ovate to suborbicular or fan-shaped, finely pubescent on 
both sides, 4-6-toothed on each margin, the teeth obtuse 
or acutish; flowers spicate-racemose; calyx pubescent, 
its teeth triangular-lanceolate; corolla about 2” long, 
violet to nearly white, with a yellow eye; capsule oblong- 
elliptic, ciliate, notched. 


Thickets and grassy places, Newfoundland to Quebec and 
Maine. July—Aug. 


4. Euphrasia Oakesii Wettst. Oakes’ Eyebright. Fig. 3843. 
Euphrasia Oakesu Wettst. Mon. Euph. 142. 1896. 


Stem erect or ascending, very slender or filiform, simple, 
1-2’ high, somewhat pubescent. Stem leaves 2 or 3 pairs. 
Bracts and leaves orbicular or broadly oval, narrowed or 
nearly truncate at the base. 2-3” long, obtuse, pubescent, 
not glandular, with 2-5 blunt teeth on each side; spike 
short, capitate, only 3-5” long, the bracts densely imbri- 
cated; calyx-teeth triangular-lanceolate, short; corolla 
14”-2” long, purplish-white with a yellow eye; capsule 
oblong-elliptic, ciliate, longer than the calyx. 


White Mountains of New Hampshire and on Mt. Katahdin, 
Maine. July—Aug. 
Euphrasia Williamsii Robinson, found on Mt. Washington, 
e N. H., differs in being nearly glabrous, with a brown-purple 
corolla. 


35. BARTSIA L. Sp. Pl. 602. 1753. 


Annual or perennial herbs, partly parasitic on the roots of other plants, with opposite 
leaves, and purple pink red or yellow flowers. in terminal leafy-bracted spikes. Calyx cam- 
panulate or tubular, 4-toothed or 4-cleft. Corolla very irregular, the tube straight or recurved, 
the limb 2-lipped; upper lip erect, concave, entire, the margins not recurved; lower lip spread- 


GENUS 35. ‘  FIGWORT FAMILY. 219 


ing, 3-lobed. Stamens 4, didynamous, ascending under the upper lip of the corolla; anther- 
sacs similar, parallel. Capsule globose, oblong, or ovoid, loculicidally dehiscent, several— 
many-seeded. Seeds horizontal, striate, or ribbed. [Named for John Bartsch, a Prussian 
botanist, died 1738. ] 


About 6 species of the northern hemisphere. Only the following is known to occur in North 
America. Type species: Bartsia viscosa L. 


1. Bartsia alpina L. Alpine Bartsia. 
Fig. 3844. 
Bartsia alpina L. Sp. Pl. 602. 1753. 


Perennial by short rootstocks, pubescent; stem erect, 
leafy, simple, or rarely with 1 or 2 short branches, 4’—10’ 
high. Leaves sessile, ovate, or ovate-oblong, crenate- 
dentate, obtuse or acutish, rounded and sometimes 
slightly clasping at the base, 2’-1’ long; bracts similar, 
smaller, mostly shorter than the flowers; spike 17-2’ 
long; flowers 8-10” long, sometimes borne also in the 
upper axils; calyx 4-cleft nearly to the middle; corolla 
purple, its tube much longer than the calyx; anthers 
pubescent, at least on the back; capsule ovoid-oblong, 
equalling or longer than the calyx. 


Labrador to Greenland and the Arctic Sea. Also in 
‘Europe. Summer. 


36. ODONTITES Gmel. Fl. Sib. 3: 213. 1768. 


Annual erect herbs, half parasitic on the roots of other plants, with small opposite leaves, 
and yellow or red flowers in terminal bracted spikes or racemes. Calyx 4-toothed. Corolla 
with a narrow tube and a strongly 2-lipped limb, the upper lip concave, entire, or 2-lobed, the 
lower 3-lobed, spreading. Stamens 4, didynamous, ascending; anther-sacs similar. Capsule 
mostly subglobose, loculicidally dehiscent. Seeds few, pendulous. [Greek, referring to its 
supposed value as a cure for toothache. ] 


About 20 species, mostly natives of the Mediterranean region, the following typical. 


1. Odontites Odontites (L.) Wettst. Red 
Bartsia. Red Eyebright. Fig. 3845. 
Euphrasia Odontites L. Sp. Pl. 604. 1753. 
Bartsia Odontites Huds. Fl. Angl. Ed. 2268. 1778. 
Odontites Odontites Wettst. in Engl. & Prantl, Nat. PA. 

Fam. 4: Abt. 3b, 102. 1891. 

Annual, appressed-pubescent, roughish; stem slen- 
der, at length much branched, 6-15’ high, the branches 
erect or ascending. Leaves sessile, lanceolate, or 
oblong-lanceolate, acute or acuminate at the apex, 
serrate with low distant teeth, slightly narrowed at 
the base, 4-13’ long, 1-3’ wide; spikes slender, 
becoming 27-5’ long in fruit, somewhat 1-sided; 
bracts similar to the leaves, but smaller; flowers nu- 
merous, 4-5” long; calyx 4-cleft; corolla red or 
pink, its tube somewhat longer than the calyx; an- 
thers slightly pubescent; capsule oblong, shorter 
than the calyx. 


In fields and waste places, coast of Maine to Nova 
Scotia. Naturalized from Europe. Native also of Asia. 
June-Sept. 


37. PEDICULARIS [Tourn.] L. Sp. Pl. 607. 1753. 


Herbs, with alternate opposite or rarely verticillate, pinnately lobed cleft or pinnatifid 
leaves, and yellow red purple or white flowers, in terminal spikes or spike-like racemes. 
Calyx tubular, cleft on the lower side or sometimes also on the upper, or 2-5-toothed. Corolla 
strongly 2-lipped, the tube cylindric, the upper lip (galea) laterally compressed, concave or 


220 SCROPHULARIACEAE. Vor. III. 


conduplicate, sometimes short-beaked ; lower lip erect or ascending, 3-lobed, the lobes spread- 
ng or reflexed, the middle one the smallest. Stamens 4, didynamous, ascending within the 
upper lip of the corolla; anthers approximate in pairs, their sacs transverse, equal, parallel, 
obtuse or rarely mucronate at the base. Capsule compressed, oblique or curved, beaked, 
many-seeded, loculicidally dehiscent. Seeds reticulate, pitted, striate or ribbed. [Latin, per- 
taining to lice, long supposed to breed lice in sheep that feed on these plants. | 


About 125 species, mostly natives of the northern hemisphere. In addition to the following, 
some 30 others inhabit the western parts of North America. Mostly known as Lousewort; a few 
red-flowered species are called Red Rattle. Type species: Pedicularis sylvatica L. 


Beak of the galea conic, decurved, 1” long. 1. P. lapponica. 
Beak of the galea very short, or none. 
Annuals or biennials; stems leafy, freely branching; northern. 
Puberulent ; upper leaves crenulate, lower pinnatifid. 2. P. euphrasioides. 
Glabrous or very nearly so; leaves all pinnatifid. 3. P. palustris. 
Perennials; stems leafy, simple (rarely branched in No. 6). 
Corolla yellow, or the galea red; plants 6’-3° high; eastern species. 
Leaves pinnately lobed; capsule ovate, scarcely longer than the calyx. 
4. P. lanceolata. ° 
Leaves pinnately parted; capsule lanceolate, 3 times as long as the calyx. 
5. P. canadensis. 


Lower leaves pinnately divided ; capsule ovate. 6. P. Furbishiae. 
Galea crimson or purple; plant 1’—4’ high; arctic. 7. P. flammea. 
Perennial; stem scapose, or 1-leaved; flowers capitate; arctic. 8. P. capitata. 


1. Pedicularis lappénica L. Lapland Pedicularis. Fig. 3846. 
Pedicularis lapponica L. Sp. Pl. 609. » 1753. 


Perennial, puberulent; stems simple, or sparingly 
branched, leafy, 4’-8’ high. Leaves sessile, or very 
short-petioled, alternate or the lowest opposite, lanceo- 
late or linear-lanceolate, obtuse or acutish at the apex, 
3/-13’ long, pinnately incised into numerous approxi- 
mate oblong serrulate lobes; spike short, the flowers 
almost capitate, light yellow, 6’-7” long; calyx cleft on 
the lower side, 2-toothed on the upper; galea erect, 
arched, tipped by an abruptly spreading or recurved 
conic beak about 1” long. 


In open places, Labrador and Greenland to the Arctic 
Sea. Also in Arctic Europe and Asia. Summer. 


Pedicularis pedicellata Bunge, an Alaskan species is re- 
corded by Bunge from Labrador. It is distinguished from 
the above by its scapose stem, deeply pinnatifid leaves and 
pedicellate lower flowers. We have not seen specimens 
from the eastern side of the continent. 


P. euphrasoides Steph.; Willd. Sp. Pl. 3: 204. 1801. 


Biennial or annual, puberulent; stem branched, 
6-15’ high, the branches ascending. Lower leaves 
petioled, lanceolate or oblong-lanceolate in outline, 
2’-4’ long, 4’-8” wide, pinnatifid into oblong, obtuse, 
crenate-dentate segments; upper leaves sessile, linear 
or linear-oblong, smaller, merely crenulate; flowers 
in a short terminal spike and solitary in the upper 
axils; calyx cleft on the lower side, 2-3-toothed on 
the upper, shorter than the corolla-tube; corolla yel- 
low, or the galea purplish, about 6” long; galea as 
long as the tube, tipped with a very short truncate 
beak, minutely 2-toothed on the lower side at the 
apex; capsule apparently shorter than the calyx. 


Labrador to Greenland, the Arctic Sea, Alaska and 
British Columbia. Also in northern Asia. Summer. 


GENUS 37. FIGWORT FAMILY. 221 


3. Pedicularis palustris L. Purple Pedicularis. Marsh Lousewort. Fig. 3848. 


Pedicularis palustris L. 607. 1753. 
?P. parviflora J. E. Smith in Rees’ Cyclop. 26: No. 4. 


1814. 
ane Wlassoviana Stev. Mem. Soc. Nat. Mosc. 

620: P/O, 1. 1823. 
Pedicularis palustris var. Wlassoviana Bunge; 

Ledeb. FI. Ross. 3: 283. 1847-49. 
_ Biennial or annual, glabrous throughout, or the 
petiole-bases ciliate; stem erect, much branched, 
1°-2° high. Leaves 1’-2’ long, alternate, or some 
of them opposite, oblong-lanceolate in outline, 
all pinnatifid into oblong crenate or incised seg- 
ments; flowers in terminal rather loose spikes 
and solitary in the upper axils, 7-9” long; calyx 
2-cleft, the lobes with an incised crested border; 
corolla purple, sometimes white, its tube twice as 
long as the calyx, the lip much shorter, the galea 
arched at the top, blunt or apiculate, not beaked, 
bearing a pair of minute teeth below its summit; 
capsule obliquely ovate, twice as long as the calyx 
when mature. 

In wet situations, Labrador to Alaska, south to 
Quebec, the Northwest Territory and Oregon. Eu- - 
rope. Summer. Red rattle, Cow’s-wort. 


4. Pedicularis lanceolata Michx. Swamp 
Lousewort. Fig. 3849. 


Pedicularis lanceolata Michx. Fl. Bor. Am. 2: 18. 1803. 
Pedicularis auriculata J. E. Smith in Rees’ Cyclop. 26: 
No. 4. 1814. 


Perennial, glabrous or very nearly so throughout; 
stem stout, simple, or branched above, 1°-3° high, the 
branches erect. Leaves alternate and opposite, lanceo- 
late, or linear-lanceolate, 2-5’ long, pinnately lobed, 
the lower petioled, the upper sessile, the lobes oblong, 
obtuse, short, crenate-dentate, the margins cartilagi- 
nous; spikes short; calyx 2-lobed, the lobes with folia- 
ceous margins; corolla yellow, 8’—10” long, the galea 
arched, terminated by a very short truncate beak, the 
lower lip erect-ascending; capsule ovate, little exceed- 
ing the calyx, about 5” high. 

in swamps, Ontario to Connecticut, North Carolina, 
Manitoba, Ohio, Michigan, South Dakota and Nebraska. 
Aug.—Oct. 


Ld 
5. Pedicularis canadénsis L. Wood or Head Betony. Lousewort. Fig. 3850. 


Pedicularis canadensis L. Mant. 86. 1767. 


Perennial, hirsute, pubescent, or glabrate be- 
low; stems commonly tufted, ascending or 
erect, 6-18’ high, simple. Leaves alternate or 
some of them opposite, oblong-lanceolate, 3’—-5’ 
long, all but the uppermost slender-petioled, 
pinnately parted into oblong, obtuse, incised or 
dentate lobes; flowers spicate, the spike short 
in flower, 5’-8’ long in fruit, the lower bracts 
usually foliaceous, the others small; calyx cleft 
on the lower side; 2-3-crenate on the upper, 
oblique; corolla yellow, or reddish (rarely 
white), 7-10” long, the tube much longer than 
the calyx, the galea arched, incurved, not beaked, 
minutely 2-toothed below the apex; capsule 
lanceolate, oblique, 7-8” long, 2” wide, about 
3 times as long as the calyx. 


In dry woods and thickets, Nova Scotia to Mani- 
toba, Florida, Mississippi, Kansas, Colorado and 
North Mexico. Ascends to 3000 ft. in Virginia. 
High heal-all. Beefsteak-plant. Lousewort-fox- 
glove. Snaffles. April—June. 


222 SCROPHULARIACEAE. Vor: be 


6. Pedicularis Furbishiae S. Wats. Miss 
Furbish’s Pedicularis. . Fig. 3851. 


P. Furbishiae S. Wats. Proc. Am. Acad. 17: 375. 1882 


Perennial, pubescent, at least above; stem strict, 
simple, 2°-3° high. Leaves lanceolate, alternate, or 
some of them opposite, the lower long-petioled, 4’-6” 
long, pinnately divided into ovate or oblong, pinna- 
tifid or incised segments, the upper sessile, pinnately 
parted or lobed; calyx 5-lobed, the lobes entire or 
dentate; narrow, somewhat unequal; corolla yellow, 
7-9” long, the galea arched, truncate, not beaked, 
2-cuspidate at the apex; capsule ovate. 


In swamps and along streams, Maine and New Bruns- 
wick. July—Sept. 


7. Pedicularis lammea L. Red-tipped 
Pedicularis. Fig. 3852. 
Pedicularis flammea L. Sp. Pl. 609. 1753. 


Perennial, glabrous, or somewhat woolly; 
stem simple, 2’-4’ high, with several linear- 
oblong, -pinnately parted leaves. Basal and 
lower leaves slender-petioled, 17-13’ long, the 
uppermost sessile, the lobes ovate or oblong, 
incised-serrate; flowers about 6” long, pedi- 
celled in a short spike-like raceme, longer than 
the narrow bracts; calyx 5-toothed, the teeth 
lanceolate, acute, unequal; corolla-tube and the 
lower lip greenish yellow, the galea slightly 
arched, very blunt, much longer than the lower 
lip, its summit crimson of purple; capsule lan- 
ceolate, 6-8” long, 2-3 times as long as the 
calyx; fruiting pedicels 3-5” long. 

Labrador and Greenland, west to Alaska. Also 
in arctic and alpine Europe. Summer. 


8. Pedicularis capitata Adams. Capitate Pedicularis. 
Fig. 3853. 
Pedicularis capitata Adams, Mem. Soc. Nat. Mose. 5: 100. 1817. 


Perennial, pubescent or glabrous; stem scapose, leafless, or 
1-leaved, 1’-5’ high. Leaves slender-petioled, often shorter 
than the scape, pinnately divided, the segments ovate or ob- 
long, incised; flowers several in a capitate cluster at the end 
of the scape, 1-13’ long; calyx 5-cleft, the lobes foliaceous, 
incised or crenate;.corolla described as white; galea scarcely 
broadened above, slightly curved, very obtuse, twice as long 
as the lower lip; capsule oblong, a little longer than the calyx, 
beaked on the outer side near the summit. 


Arctic America; Hudson Bay to Alaska. Summer. 


38. ELEPHANTELLA Rydb. Mem. N. Y. Bot. Gard. 1: 362. 1900. 


Herbs similar to Pedicularis in habit. Leaves pinnately parted or pinnately divided. 
Inflorescence erect. Calyx 2-lipped, often campanulate. Corolla very strongly 2-lipped, the tube 
short, the upper lip (galea) produced into an elongated slender beak which is soon turned 
upward, the lower lip very broad. [Greek, little elephant, referring to the resemblance of 
the galea of the corolla to an elephant’s head.] 

Two or three species of northern regions. Type species: Elephantella groenlandica (Retz.) Rydb. 


GENUS 38. FIGWORT FAMILY. 223 


1. Elephantella groenlandica (Retz.) Rydb. Long-beaked Pedicularis. 
Fig. 3854. 
P. groenlandica Retz. Fl. Scand. Ed. 2, 145. 1795. 
Elephantella groenlandica Rydb. Mem. N. Y. Bot. Gard. 1: 

363. 1900. 

Perennial, glabrous; stem simple, erect, 1°-13° high. 
Leaves alternate, lanceolate in outline, acute or acumi- 
nate, pinnately parted or the lower pinnately divided 
into lanceolate, acute, crenulate or incised segments, 
the upper sessile, the lower slender-petioled, 2’-6’ long; 
spike 1-6’ long, very dense; calyx 5-toothed, nearly as 
long as the corolla-tube, the teeth short, acutish; co- 
rolla red or purple, the galea produced into a filiform 
beak 6-8” long, which is decurved against the lower 
lip and upwardly recurved beyond it; body of the 
corolla 23’-3” long; capsule obliquely ovate, about 
3” long. 

In wet soil, Labrador, Greenland and Hudson Bay to 
Athabasca, British Columbia, south in the Rocky Moun- 


tains to New Mexico, and in the Sierra Nevada to Cali- 
fornia. Summer. 


39. RHINANTHUS L. Sp. Pl. 603. 1753. 


Annual erect mostly branched herbs, with opposite leaves, and yellow blue violet or 
variegated flowers, in terminal I-sided leafy-bracted spikes, or solitary in the upper axils. 
Calyx compressed, 4-toothed, much inflated, membranous and conspicuously veiny in fruit. 
Corolla very irregular, 2-lipped, the upper lip (galea) compressed, arched, minutely 2-toothed 
below the entire apex, the lower lip 3-lobed, shorter, the lobes spreading. Stamens 4, didy- 
namous, ascending under the galea; anthers pilose, the sacs obtuse at the base, transverse, 
distinct. Capsule orbicular, flat, loculicidally dehiscent, several-seeded. Seeds nearly orbicu- 
lar, winged. [Greek, nose-flower, from the beaked corolla.] 


About 3 species, natives of the northern hemisphere. Type species: Rhinanthus Crista-galli L. 


1. Rhinanthus Crista-galli L. Rattle. Rattle-box. Yellow or Penny Rattle. 
Fig. 3855: 

Rhinanthus Crista-galli L. Sp. Pl. 603. 1753. 
Rhinanthus minor Ehrh. Beitr. 6: 44. 17091. 

Glabrous, or pubescent above; stem slender, usu- 
ally branched, 6-18’ high, the branches erect or 
ascending. Leaves lanceolate or oblong-lanceolate, 
sessile, coarsely serrate-dentate, acute or obtuse, 1’—2’ 
long, 2-4” wide; bracts broader, ovate, or ovate- 
lanceolate, incised-dentate, the teeth acuminate or 
subulate-tipped ; flowers yellow, 6-8” long; corolla- 
tube longer than the calyx, commonly with a purple 
spot on one or both lips, the teeth of the upper lip 
broad; fruiting calyx ovate-orbicular, 4’-6” in diam- 
eter; capsule orbicular, or broader, nearly as broad 
as the calyx, very flat, not oblique. 

Newfoundland and Labrador to Alaska and Oregon, 
iM south to Quebec, the White Mountains of New Hamp- 
shire, New York and in the Rocky Mountains to New 
Mexico; on the Atlantic Coast from Connecticut to New 
Brunswick. Common in northern Europe and Asia. 
Consists of several races, sometimes regarded as species, 


differing in habit and in color of the corolla-lips. Called 
yellow cockscomb, also rattle-bags. Penny-grass. Money-grass. June—Aug. 


; Rhinthus major Ehrh., found many years ago in fields at Plymouth, Massachusetts, differs 
in having the teeth of the upper corolla-lip elongated, its flowers somewhat larger. It is a native 
of Europe. 


224 SCROPHULARIACEAE. Vou. Th 


40. MELAMPYRUM [Tourn.] L. Sp. Pl. 605. 1753. 


Annual branching herbs, with opposite leaves, and small white yellow violet or variegated 
flowers, solitary in the upper axils, or in terminal bracted spikes. Calyx 4-toothed, the 2 
upper teeth somewhat the longer. Corolla irregular, 2-lipped, the tube narrow, gradually 
enlarged above, the upper lip compressed, obtuse or emarginate with a groove behind the 
margins, or these recurved or with a tooth on each side; lower lip spreading or ascending, 
3-toothed, 2-grooved beneath. Stamens 4, didynamous, ascending under the upper lip; anther- 
sacs distinct, parallel obtuse or mucronulate at the base. Capsule flat, oblique, loculicidally 
dehiscent, 2-4-seeded. Seeds smooth, strophiolate. [Greek, black wheat.] 


About 10 species, all of the northern hemisphere. Only the following are known in North 
America. Type species: Melampyrum arvense L. 
Leaves lanceolate or linear-lanceolate, the floral 2-4-toothed at the base; capsule much longer than 
wide, long-beaked. 1. M. lineare. 
Leaves ovate, all entire; capsule slightly longer than wide, short-beaked. 2. M. latifolium. 


1. Melampyrum lineare Lam. Narrow-leaved Cow-Wheat. Fig. 3856. 


Melampyrum lineare Lam. Encycl. 4: 22. 1797. 
M. americanum Michx. Fl. Bor. Am. 2: 16. 1803. 

Puberulent; stem slender, obscurely 4-sided 
above, at length widely branched, 6—13° high. 
Leaves lanceolate or linear-lanceolate to ovate, 
short-petioled, acuminate or acute at the apex, 
narrowed, obtuse, or the upper truncate at the 
base, 1’-23’ long, 13”-6” wide, the lower entire, 
the upper floral ones ovate or lanceolate, with 
2-6 bristle-pointed teeth near the base or entire; 
flowers short-peduncled, 4’-6” long; calyx about 
one-third the length of the corolla, its subulate 
teeth longer than its tube; corolla white or whitish, 
puberulent, the lower lip yellow; capsule 47-5” 
long, about 2” wide, twice as long as the calyx, 
long-beaked. 

In dry woods and thickets, Nova Scotia to British 
Columbia, south to Georgia, Tennessee, lowa, Mon- 


tana and Idaho. Consists of several races, differing 
in leaf-form. Ascends 3500 ft. in Virginia. May-— Aug. 


2. Melampyrum lIatifolium Muhl. Broad- 
leaved Cow-Wheat. Fig. 3857. 


Melampyrum latifolium Muhl. Cat. 57. 1813. 


Similar to the preceding, widely branched, 1°-12° 
high; but the leaves all entire, short-petioled, the lowest 
small, spatulate, obtuse, the middle ones lanceolate or 
ovate, acuminate, narrowed at the base, 2’-3’ long, the 
floral ovate or ovate-lanceolate, shorter, acute, mostly 
rounded at the base; corolla purple, veiny, larger; cap- 
sule little longer than wide, short-beaked. 


In dry woods, Delaware (according to Muhlenberg) ; 
mountains of Virginia to Georgia and Tennessee. June— 
Aug. 


GENUS I. BLADDERWORT FAMILY. 225 


Family 29. LENTIBULARIACEAE Lindl. Veg. Kingd. 686. 1847.* 
BLADDERWORT FAMILY. 


Aquatic plants, or if terrestrial usually on moist ground, the leaves in a basal 
rosette, or borne along floating stems, or more or less root-like, or wanting. In- 
florescence scapose. Flowers solitary or racemose, perfect, irregular. Calyx 
inferior, 2—5-lobed or parted, persistent. Corolla ‘hypogynous, gamopetalous, 
2- lipped, the upper lip entire or 2-lobed, the lower lip entire or 3-lobed, usually 
with a nectariferous spur. Stamens 2, borne upon the base of the corolla; anther- 
sacs confluent into I. Ovary superior, usually subglobose, 1-celled ; placenta sub- 
globose, central, free; ovules 2 or more, usually very numerous; style short or 
obsolete; stigma 2-lipped, the anterior lip larger, lamelliform. Fruit a capsule, 
bursting irregularly, or dehiscent by valves. Seeds variously appendaged or sculp- 
tured; embryo in the axis, often imperfectly developed; endosperm none. 


About 16 genera and 300 species, of world-wide distribution. 


Scapes without bracts or scales, 1-flowered ; leaves in a basal rosette. Pinguicula. 
Scapes with one or more bracts, with or without scales; leaves usually alternate, offen dissected 
or root-like and bladder-bearing. 
Bracts at the base of the pedicels without bractlets; calyx not enclosing the fruit. 
Branches verticillate, and verticillately or oppositely decompound; lateral lobes of lower lip 


of corolla saccate. 2. Vesiculina. 
Branches alternate or none; lateral lobes of lower lip of corolla not saccate. 
Bracts, and scales if present, flat, basally attached ; aquatic. 3. Utricularia. 
Bract solitary, tubular, surrounding the scape; scales none. 4. Lecticula. 
Bracts and scales peltate ; terrestrial. 5. Setiscapella. 
Bracts at the base of the pedicels accompanied by a pair of bractlets; calyx enclosing the fruit; 
terrestrial. 6. Stomoisia. 


1. PINGUICULA [Tourn.] L. Sp. Pl. 17. 1753. 


Acaulescent herbs, with fibrous roots, naked I-flowered circinate scapes, and leaves in 
a basal rosette, the upper surface commonly glandular and covered with a viscid secretion. 
Calyx 5-lobed, more or less 2-lipped, the upper lip 3-lobed, the lower 2-lobed. Corolla 5-lobed, 
more or less 2-lipped, the upper lip 2-lobed, the lower 3-lobed; base of the corolla saccate 
and contracted into a nectariferous spur. Capsule 2-valved. Seeds oblong, reticulate. [Latin, 
pinguis, fat, from the apparent greasiness of the leaves of several species. ] 

About 35 species, of wide distribution in the northern hemisphere, and southward along the 


Andes to Patagonia. Besides the following, 4 others are found in the southeastern United States. 
Type species: Pinguicula vulgaris L. 


Scapes villous, 1’—2’ high; corolla pale violet, less than 6” long. TP villosa. 
Scapes glabrous or nearly so, 2-6’ high; corolla violet- purple, more than 7” long. 2. P. vulgaris. 


1. Pinguicula villosa L. Hairy Butterwort. 
Fig. 3858. 


Pinguicula villosa L. Sp. Pl. 17. 1753. 
Pinguicula acutifolia Michx. Fl. Bor. Am. 1: 11. 1803. 


Scapes slender, densely villous, 1-2’ high, becoming 14’-5’ 
high in fruit. Leaves 3-6, the blades oval, obtuse or emargi- 
nate, 3-6” long, the margins usually inrolled; calyx minute, 
4”_2” long, the lobes acute, the 2 lower ones. smaller and 
partly united; corolla pale violet with a yellowish striped 
throat, 4”-6” long including the slender obtuse spur, 2-lipped, 
the upper lip erect, 2-lobed, the lower spreading, 3-lobed; cap- 
sule subglobose, about 1” in diameter. 


In bogs, circumpolar; southward in America to Labrador, Hud- 
son Bay, Yukon, and Alaska. June-July. 


* Text contributed by Dr. JooN HENDLEY BARNHART. 
15 


226 LENTIBULARIACEAE. Vor. ITI. 


2. Pinguicula vulgaris L. Common Butter- 
wort. Bog Violet. Fig. 3859. 
Pinguicula vulgaris L. Sp. Pl. 17. 1753. 


Scapes glabrous or nearly so, 1-6’ high, little if 
at all elongating in fruit. Leaves 3-7, the blades 
ovate to elliptic, obtuse, 2’-12’ long, the margins 
usually inrolled; calyx 13-23” long, the lobes ob- 
tuse, the 2 lower ones more or less united; corolla 
violet-purple, 7-10” long including the subulate 
acute spur, 2-lipped, the lips equally spreading, the 
upper 2-lobed, the lower 3-lobed; capsule ovoid, 
3-4” long, 23’-3”" in diameter. 

On wet rocks or gravelly places, circumpolar; south- 
ward in America to Newfoundland, New Brunswick, 
Vermont, northern New York, Michigan, Minnesota, 
Montana, and British Columbia. June-July. Bean- 
weed. Yorkshire sanicle. Sheep-root or -rot. Rot-grass. 
Sheepweed. Steep or earning-grass, from its use in 
curdling milk. : 

Pinguicula alpina L., reported from Labrador on 
account of a single specimen said to have been col- 
lected there many decades ago, probably does not occur 
in North America. It has flowers about the size of 
those of P. vulgaris, but nearly white and with a very 
much shorter obtuse spur. 


2. VESICULINA Raf. Fl. Tellur. 4: 109. 1838. 


Aquatic herbs, with horizontal submerged stems, the branches verticillate, and verticil- 
lately or oppositely decompound. Leaves, at least in the adult plant, none. Bladders terminal 
on the ultimate branches, the mouth naked or with a single median hairy proboscis. In- 
florescence racemose, I-4-flowered; scales on the lower portion of the scape none; pedicels 
from the axils of bracts, without bractlets, erect in fruit. Calyx 2-lobed, the lobes herba- 
ceous, concave. Corolla strongly 2-lipped, the upper lip not lobed, the lower lip 3-lobed, the 
lateral lobes saccate and together constituting a prominent 2-lobed palate, the middle lobe 
flat, comparatively inconspicuous. Anthers not lobed. Capsule many-seeded. Seeds tuber- 
culate. [Latin, vesicula, a little bladder.] 

About 6 species, confined to the New World. Type species: Utricularia saccata LeConte. 


1. Vesiculina purptrea (Walt.) Raf. 
Purple Bladderwort. Fig. 3860. 
Utricularia purpurea Walt. Fl. Car. 64. 1788. 
U. saccata LeConte; Ell. Bot. S.C. & Ga. 1: 21. 1816. 
Vesiculina saccata Raf. Fl. Tellur. 4: 109. 1838. 
Vesiculina purpurea Raf. loc. cit. 1838. 

Stems 1°-3° long, the internodes 1’-2’ long, the 
primary branches in whorls of 5-7, verticillately 
decompound. Leaves none; bladders 1-13” long, 
without bristles, the exterior surface of the valve 
with a central tuft of glandular hairs; scape 2’-6’ 


high, 2-4-flowered; bracts membranaceous, basally NY) La at WA 
attached, or more commonly with a free portion ay P, ty ff pez 
below the line of insertion; pedicels 3’—4” long, q YW SS Ki WY yb 
i it; c= Ms AN OR | YAR KVL 9 A 
half longer in fruit; calyx-lobes subequal, 14 SY At Wy SV ee 


long; corolla red-purple, the upper lip subrhom- ( 
boid, about 4” long and 6” broad, concave, the 
lower lip 4-6” long and broad, with a yellow 
spot at the base; spur conic, appressed to and 
shorter than the lower lip; capsule 14” in diam- 
eter; seeds numerous, minute, tuberculate-spiny, 
especially toward each end. 

In ponds, Maine to Florida and Louisiana, near 
the coast, also Michigan and Indiana to Minnesota. 
Hooded or horned milfoil. July—Sept. 


3. UTRICULARIA L. Sp. Pl. 18. 1753. 


Aquatic herbs, with horizontal submerged leafy stems. Leaves alternate, dissected, some- 
times root-like, 2-8-parted from the very base, and thus often appearing opposite or verti- 
cillate, the segments dichotomously or pinnately dissected, some or all of them bladder-bearing. 
Bladders with a pair of bristles and more or less other armature about the mouth. Inflores- 
cence racemose, the raceme sometimes reduced to a single flower; scales on the lower portion 
of the scape, when present, basally attached, sometimes’ replaced by a whorl of inflated 


GENUS 3. BLADDERWORT FAMILY. 227 


floats; pedicels from the axils of basally inserted but sometimes auriculate bracts, without 
bractlets. Calyx 2-lobed, the lobes concave, herbaceous, usually spreading under the mature 
capsule. Corolla strongly 2-lipped, the palate at the base of the lower lip, prominent, usually 
2-lobed. Anthers not lobed. Capsule few-many-seeded. Seeds more or less peltate, flat- 
topped, the margin variously winged or wingless. [Latin, utriculus, a little bag.] 


About 75 species, of world-wide distribution. Besides the following, 2 or 3 others occur in 
the southeastern United States. Type species: Utricularia vulgaris L. 


Scape without floats. 
Stems creeping on the bottom in shallow water; some or all of the leaves root-like. 
Spur and palate conspicuous; pedicels ascending in fruit. 
Segments of leaves all capillary ; lips of corolla nearly equal in Seca 
Spur stout, conic, shorter than the lower lip. . U. gibba. 
Spur slender, equalling or exceeding the lower lip. 
Spur tapering from base to apex; leaves all alike, bladder-bearing; scapes 2’-5’ 


high. 2. U. pumila. 

Spur conic at base, linear above; leaves not all bladder-bearing; scapes 4’—16’ 
high. a brosa. 
Segments e some leaves linear, flat, bristly-serrulate; upper lip of corolla Shake half 
length of lower lip. 4. U. intermedia. 


Spur a mere sac; palate obsolete; pedicels recurved in fruit. 5. U. minor. 
Stems free-floating, ‘except fora single point of attachment. 
Scapes 2-5-flowered, without scales; cleistogamous flowers also present. 6. U. geminiscapa. 
Scapes 6—20- flowered, with 1-5 scales ; cleistogamous flowers none. 7, U. macrorhiza. 
Scape with a whorl of more or less united, conspicuous, inflated floats. 
Scape from floats to lowest pedicel 2’ long or less; corolla 6”—9” long. 8 
Scape from floats to lowest pedicel 3’ long or more; corolla forest long. 9 


. U. radiata. 
. U. inflata. 


1. Utricularia gibba L. Humped or Swollen- 
spurred Bladderwort. Fig. 3861. 


Utricularia gibba L. Sp. Pl. 18. 1753. 
U. fornicata LeConte, Ann. Lyc. N. Y. 1: 76. 1824. 


Stems creeping on the bottom in shallow water, 
radiating from the base of the scape; leaves alter- 
nate, mostly once or twice dichotomous, the seg- 
ments few, capillary, bladder-bearing. Scapes soli- 
tary, ?’-4’ high, 1-3-flowered, with 1 scale or none; 
pedicels 1-5” long, erect-ascending; calyx-lobes 
1-13” long; corolla yellow, the upper lip subtrian- 
gular, 2’-3” long, 3’-4” broad, the lower about 3” 
long and broad, the palate prominent, 2-lobed; spur 
conic, very obtuse, shorter than the lower lip; cap- 
sule globose, 13-2” in diameter. 


In shallow water, Maine to Michigan, south to Flor- 
ida and Texas. June—Sept. 


2. Utricularia pumila Walt. Two-flowered 
Bladderwort. Fig. 3862. 
Utricularia pumila Walt. FI. Ge 64. 1788. 


Y 5 Utricularia biflora Lam. Ill. 1: 50. 179 
U. longirostris LeConte ; Fil. Bot. S. G '& Gare 2nr. 
1816. 
e U. macrorhyncha Barnh. Bull. Torr. Club 25: 515. 
1898. 


Stems creeping on the bottom in shallow water, 
radiating from the base of the scape; leaves 
alternate, dichotomously divided, the segments 
capillary, bladder-bearing. Scapes solitary or two 
together, 2-5’ high, 1-4-flowered, with I-3 scales 
or none; pedicels 23”-8” long, erect-ascending ; 
corolla yellow, the upper lip more or less 3-lobed, 
4-5” long, 6-7” broad, the lower 4’-5” long 

and broad, the palate prominent, 2-lobed; spur 

~ slenderly conic or subulate, obtuse or emarginate, 
equalling or usually exceeding the lower lip; 
capsule globose, 14” in diameter. 

In shallow water, Massachusetts to Florida and 
Louisiana, near the coast. July—Aug. 


LENTIBULARIACEAE. Vor. ITI. 


3. Utricularia fibrosa Walt. Fibrous Blad- 
derwort. Fig. 3863. 


Utricularia fibrosa Walt. Fl. Car. 64. 1788. 
U. siriata LeConte; Torr. Cat. Pl. N. Y. 89. x8x0; 


Stems creeping on the bottom in shallow water, 
radiating from the base of the scape. Leaves 
alternate, trichotomous at base, the rays di- or 
trichotomously divided, the segments capillary, 
fewer and more or less root-like when bladder- 
bearing. Scapes solitary or 2 together, 4’-16' 
high, 2-6-flowered, with I or 2 scales or none; 
pedicels 23”-6” long, one-half longer in fruit, 
ascending; calyx-lobes 1-13” long, a little longer 
in fruit; corolla yellow, the upper lip broadly 
triangular, faintly 3-lobed, 33’-5” long, 67-7” 
broad, radiately striate, the lower slightly 3-lobed, 
33-5” long and broad, the palate prominent, 
2-lobed; spur conic at base, slender above, obtuse 
or emarginate, appressed to and equalling or. ex- 


ceeding the lower lip; capsules 2’—22” in diameter. 


In shallow water, Long Island to Florida and Mis- 
sissippi, near the coast. June—Aug. 


4. Utricularia intermédia Hayne. Flat- 
leaved Bladderwort. Fig. 3864. 
U. intermedia Hayne, in Schrad. Journ. Bot. 1800': 

18. 1801. 

Stems creeping on the bottom in shallow water, 
radiating from the base of the scape. Leaves 
alternate, those on some branches or portions of 
branches 23’-7” long, trichotomous at base, the 
rays di- or trichotomously divided, with linear, 
flat, bristly-serrulate segments, and without blad- 
ders, those on other branches or portions of 
branches shorter and root-like, with fewer capil- 
lary segments and a few large bladders up to 
23” long; scapes solitary, 2’-8’ high, 1-4-flowered, 
with I or several auriculate scales; pedicels 4-10” 
long, erect-ascending; calyx-lobes 1’-13” long, a 
little larger in fruit; corolla yellow, the upper 
lip broadly triangular, 23’”-3” long, 33’—4” broad, 
the lower slightly 3-lobed, 5’-6” long and broad, 
the palate prominent; spur conic at base, cylindric 
above, acute, about # as long as the lower lip; 
capsules 14” in diameter. 

In shallow water, Newfoundland to British Co- 
lumbia, south to New Jersey, Indiana and California. 
Also in Europe. May—Aug. Commonly propagated 
by the velvety winter-buds. : 


5. Utricularia minor L. Lesser Bladder- 
wort. Fig. 3865. 
Utricularia minor L..Sp. Pl. 18. 1753. 


Stems creeping on the bottom in shallow water, 
4-12’ long, sparingly branched. Leaves alter- 
nate, with few divisions, usually only 47-12” 
long, and bearing 1-5 bladders each; larger blad- 
derless leaves, 1-3” long, with flat segments, 
sometimes occurring on special branches or seg- 
ments of the main stem; scapes solitary, filiform, 
2’-6' high, 3-6-flowered, with 2-5 minute auricu- 
late scales; pedicels capillary, 1-4” long, re- 
curved in fruit; calyx-lobes about 2” long; co- 
rolla pale yellow, the upper lip: minute, 17-2” 
long and half as wide, the lower 2-4” long, the 
palate nearly obsolete; spur very short and sac- 
cate; capsules about 2” in diameter. 

In shallow water, circumpolar, southward in 
America to Connecticut, New York, Pennsylvania, 
Ohio, Indiana, Colorado and California. June— 
July. Often propagated by winter-buds. 


GENUS 3. BLADDERWORT FAMILY. 229 


6. Utricularia geminiscapa Benj. Hid- 
den-fruited Bladderwort. Fig. 3866. 


Utricularia geminiscapa Benj. Linnaea 20: 305. 


1847. 
U. Pi idestina Nutt.; A. Gray, Man. 287. 1848. 
Stems about 8—10’ long, floating horizontally 
beneath the surface of the water, sparingly 
branched. Leaves alternate, about 10” long, 
4-7-dichotomous, and bladderless, or more or 
less reduced and bladder-bearing; scape slen- 


der, 2’-5’ high, 2-5-flowered, without scales; WAS 
cleistogamous flowers solitary on short pedun- We 
cles, 1 at the base of the scape and others scat- New 
tered along the stems, often in pairs; pedicels R Gita I 
i Beal, . ioe ROCESS x b 
of conspicuous flowers 2-3” long; corolla We BES hs 
yellow, 3” long or more, the lower lip longer 4 t m= al) MI 
and broader than the upper, 3-lobed, with a OE PA) 
prominent palate; spur a little shorter than Vg ' 
the lower lip, obtuse. ti Iwi 
In shallow water, New Brunswick to Virginia. CGN } M4 
Hooded or horned milfoil. July—Aug. ANN 8 'Sa8"? ln Se Gg a PUVA 
AyY TNR RA SASS a, = Sa 


7. Utricularia macrorhiza LeConte. 
Greater Bladderwort. Hooded 
Water Milfoil. Pop-weed. 


Fig. 3867. 
Utricularia macrorhiza LeConte, Ann. Lyc. N. Y. 
L735 Loe. 
U. vulgaris var. americana A. Gray, Man. Ed. 5, 
318. 1867. 


Stems 1°-3° long, floating horizontally be- 
neath the surface of the water, sparingly 
branched. Leaves alternate, #’—2’ long, dichot- 
omous at the base, each ray pseudo-pinnately 
divided, bladder-bearing, the bladders some- 
times 2” long; scape stout, 3-24’ high, 6-20- 
flowered, with 1-5 auriculate scales; pedicels 
3-8” long, becoming 5’—-10” long and recurved 
at maturity of fruit; corolla yellow, 7’-10” 
long, the lower lip a little longer and much 
broader than the upper, with a spreading, un- 
dulate, slightly 3-lobed border, and a promi- 
nent palate; spur shorter than the lower lip, 
SO subulate, upwardly curved, acute or obtuse. 
@wZ In stagnant water or sluggish streams, New- 

ANS foundland to Yukon, south to Maryland, Missouri, 
Oklahoma, Arizona, and Lower California. May-Aug. Often propagated by winter-buds. Very 
variable, but appears to differ constantly from the related and equally variable European species, 
Utricularia vulgaris L., by the longer stems, the 
shape and direction of the spur, and the minute- 
ness of the appendages (rudimentary stolons) at 
the base of the scape. 


8. Utricularia radiata Small. Small 
Swollen Bladderwort. Fig. 3868. 
Utricularia inflata var. minor Chapm. FI. S. U. S. 

282. 1860. Not U. minor L. 1753. 
Utricularia radiata Small, Fl. SE. U.S. 1090. 1903. 

Stems long, floating horizontally beneath 
the surface of the water. Leaves alternate, 
6-10-dichotomous, usually less than 14’ long, 
bladder-bearing; scape 1-4-flowered, with a 
whorl of 4-7 inflated floats, above the floats 
13-4’ high; floats up to 14’ long, lobed and 
finely dissected toward the apex; pedicels 4’— 
10” long; corolla 6-9” long, the upper lip 
suborbicular, undulate, the lower lip about the 
same length but broader, 3-lobed, with a 
prominent 2-lobed palate; spur conic, obtuse, 
appressed to and shorter than the lower lip. 

In ponds, Maine to Florida and Texas, near the 
coast. March—Sept. Ww 


230 LENTIBULARIACEAE. Vor. ITI. 


g. Utricularia inflata Walt. Large Swollen 
Bladderwort. Fig. 3869. 


Uiricularia inflata Walt. Fl. Car. 64. 1788. 
U. ceratophylla Michx. Fl. Bor. Am. 1: 12. 1803. 


Stems long, floating horizontally beneath the sur- 
face of the water. Leaves alternate, 10-12-dichoto- 
mous, usually more than 2’ long, bladder-bearing; 
scape 4-12-flowered, with a whorl of 4-9 inflated 
floats, above the floats 5’-12’ high; floats 2-32’ long, 
lobed and dissected from near the middle; pedicels 
8-16” long; corolla 10’-12” long, the upper lip 
subtriangular, undulate, often emarginate, the lower 
lip a little longer and much broader, undulate, faintly 
3-lobed, with a prominent 2-lobed palate; spur conic, 
toothed, appressed to and about half the length of 
the lower lip. 


In ponds, Delaware to Florida, near the coast. May. 


4. LECTICULA Barnhart. 


Herbs, with horizontal usually submerged leafy stems. Leaves alternate, 3-parted from 
the very base, the middle lobe erect and linear, the lateral {obes capillary and root-like, bladder- 
bearing; bladders slightly beaked, but without bristles. Inflorescence strictly 1-flowered, the 
pedicel continuous with the scape, its point of origin marked by the solitary bract, appearing 
like a scale above the middle of the scape; true scales none; bract basally inserted, amplexi- 
caul and tubular, the free margin truncate, more or less deeply 2-notched, without bractlets. 
Calyx 2-parted, the lobes concave, herbaceous, appressed to the mature capsule. Corolla 
very strongly 2-lipped, the palate a mere convexity at the base of the lower lip. Anthers not 
lobed. Capsule many-seeded. [Latin, a couch, from the transverse position of the corolla.] 


Two species, the following, and another, in tropical South America. Type species: Utricu- 
laria resupinata B. D. Greene. 


1. Lecticula resupinata (B. D. Greene) 
Barnhart. Reclined Bladderwort. 
Fig. 3870: 


Utricularia resupinata B. D. Greene; Bigel. FI. 

Bost. Ed. 3, 10. 1840. 

Utricularia Greenei Oakes, Hovey’s Mag. Hort. 7: 

180. 1841. 

Scape and pedicel slender, 1’-4’ high, becoming 
much elongated, 4-6’ high in fruit, the bract 
2-1” long. Flower half-reversed so as to rest 
transversely upon the summit of the pedicel; 
calyx-lobes subequal, about 1” long; corolla 
purple, 4”-6” long, the upper lip narrowly oblong- 
spatulate, the lower spreading, entire; spur conic- 
cylindric, obtuse, the tip distant from the lower 
lip and bent upward; capsule globose, 12’—2” in 
diameter. 

Margins of ponds and lakes, New Brunswick to 
western Ontario and Pennsylvania, and South Caro- 
lina to Florida. Rare and local. July—Aug. 


5. SETISCAPELLA Barnhart. 


Terrestrial herbs, with short root-like branches from the base of the scape. Leaves deli- 
cate, some basal, erect, with linear blades, usually evanescent and rarely seen, others root-like, 
borne on the root-like branches, and bladder-bearing; bladders minute, 2-horned at the apex. 
Inflorescence racemose, the raceme sometimes reduced to a single flower, when several- 
flowered usually becoming zig-zag above at maturity; scales on the lower portion of the 
scape several, scarious, peltate; pedicels from the axils of scarious peltate bracts, without 
bractlets. Calyx 2-parted, the lobes scarious, strongly longitudinally ribbed, spreading under 
or clasping the base of the mature capsule. Corolla strongly 2-lipped, the lower lip commonly 


GENUS 5. BLADDERWORT FAMILY. 231 


strongly and divergently 3-lobed; palate usually very prominent and 2-lobed. Anthers not 
lobed. Capsule many-seeded. Seeds prismatic, reticulate. [Latin, seta, a bristle, and scapus, 
scape, from the stiff, bristle-like scapes. ] 

About 12 species, chiefly American; 1 species in tropical Africa and 2 in Asia. Type species: 
Utricularia subulata L. 


Corolla yellow, the lower lip conspicuous, 2”—-4” long; spur conic. 1. S. subulata. 
Corolla white or purplish, both lips minute, less than 44” long; spur saccate. 2. S.cleistogama. 


1. Setiscapella subulata (L.) Barnhart. Zig- 
- zag or Tiny Bladderwort. Fig. 3871. 


Utricularia subulata L. Sp. Pl. 18. 1753. 
Utricularia setacea Michx. Fl. Bor. Am. 1: 12. 1803. 

Scape filiform, stiff, 14’-9’ high, 1-12-flowered, the 
pedicels capillary, 13-3” long. Calyx-lobes minute, 
about 3” long, becoming 1” long in fruit; corolla pale 
yellow, 3-6” long, the upper lip ovate, obtuse, the 
lower 2”-4” long, strongly and divergently 3-lobed, 
with a prominent faintly 2-lobed palate; spur flat- 
tened-conic, obtuse, ‘but acute in outline when viewed 
from the side, appressed to and about equalling the 
lower lip; capsule globose, 1” in diameter; seeds 
irregularly ovoid. 

In wet sandy soil, Nantucket to Florida, west to Ar- 
kansas and Texas. Also in the West Indies. July—Aug. 


2. Setiscapella cleistogama (A. Gray) Barn- 


hart. Pin or Closed Bladderwort. Fig. 3872. 
U. subulata var. cleistogama A. Gray, Syn. Fl. 2': 317. 1878. 
U. cleistogama Britton, Trans. N. Y. Acad. Sci.g: 12. 1880. 

Scape filiform, stiff, 4-23’. high, 1-3-flowered, the 
pedicels 1-3” long. Calyx-lobes minute, the upper 
faintly 7-nerved, the lower strongly 5-nerved; corolla 
dirty-white or purplish, +” in diameter or less, cleistoga- 
mous (?), consisting chiefly of the large saccate spur, 
the lips minute, obtuse, the lower faintly 3-lobed; cap- 
sule globose, 2” in diameter; seeds very minute and 
numerous, irregularly ovoid, ro” long. 

In wet soil, eastern Massachusetts, Long Island and New 
Jersey. Rare and local. July—Aug. 


6. STOMOISIA Raf. Fl. Tellur. 4: 108. 1838. 


Terrestrial scapose herbs, with tufts of root-like bladder-bearing branches arising from 
near the base of the scape, and few extremely delicate leafy branches radiating horizontally 
from the same region, so inconspicuous and so readily detached from the scape that they are 
rarely seen. Leaves delicate, some linear, erect, and grass-like, others colorless and root-like, 
bladder-bearing. Bladders minute and rudimentary, beaked but without bristles. Inflores- 
cence racemose or subspicate, sometimes reduced to a single flower; scales on the lower 
portion of the scape several, basally inserted; pedicels from the axils of basally inserted 
bracts, with an inner pair of opposite lateral bractlets. Calyx 2-parted, the lobes thin, veiny, 
appressed to and exceeding the mature capsule. Corolla 2-lipped, the lips nearly distinct, the 
upper narrow, erect, with a distinct claw, the lower with a prominent galeate palate, the edge 
of the aperture of the spur usually ciliate. Anthers vertically 2-lobed. Capsule many-seeded. 
Seeds very minute, subglobose or prismatic, reticulate or areolate. [Greek, hairy mouth, 
from the ciliate aperture of the spur in the type species. ] : 

About 50 species, of wide geographic distribution. Type species: Utricularia cornuta Michx. 
Corolla much exceeding the calyx. 


Lower lip of corolla 6”—-8” long; spur 3%4”—6” long. 1, S. cornuta. 
Lower lip of corolla 4”—-5” long; spur 2%’—4” long. 2. S. juncea. 
Corolla shorter than or about equalling the calyx. 3. S. virgatula. 


232 LENTIBULARIACEAE. Vor. III. 


TEX L y 1. Stomoisia corntta (Michx.) Raf. Horned 
{| Bladderwort. Fig. 3873. 


4 
ry Utricularia cornuta Michx. Fl. Bor. Am. 1: 12. 1803. 


G KS Stomoisia cornuta Raf. Fl. Tellur. 4: 108. 1838. 


Scape strict, brownish, 13’-12’ high, 1-5-flowered, 
the flowers usually approximate near the summit, the 
scales several or numerous, rarely exceeding 1” in 
length, acute. Bracts 4’—1” long, acute; bractlets the 
same length, but narrower; pedicels rarely exceeding 
the bracts; calyx yellowish, the upper lobe acumi- 
nate, 2’-3” long, the lower acute, often much shorter; 
corolla yellow, the lower lip 6’-8” long, with a 
spreading margin around the galeate palate; spur 
subulate, pendent, 32”-6” long; capsule subglobose, 
3’"-2”" in diameter, closely invested by the calyx- 
lobes and slightly beaked by the upper one. 


On wet borders of ponds, or in bogs, Newfoundland 
to Minnesota, south to Florida and Texas. Also in the 
Bahamas and Cuba. July—Aug. 


2. Stomoisia jancea (Vahl.) Barnhart. 
Rush Bladderwort. Fig. 3874. 
Utricularia juncea Vahl, Enum. 1: 202. 1804. 
Utricularia personata LeConte; Ell. Bot. S. C. & Ga. 

149 a) rites). 

Scape strict, brownish, 4-16’ high, 1-12-flow- 
ered, the flowers usually scattered along the 
upper part, the scales several or numerous, 3’-3” 
long, acute. Bracts #’-1” long, acute; bractlets 
the same length, but narrower; pedicels rarely 
exceeding the bracts; calyx yellowish, the upper 
lobe acuminate, 2-23” long, the lower acute, 
much shorter; corolla yellow, the lower lip 4-5” 
long, with little or no spreading margin; spur 
subulate, pendent, 23’-4” long; capsule subglo- 
bose, 1-13” in diameter, closely invested by the 
calyx-lobes and slightly beaked by the upper one. 

On wet borders of ponds, and in bogs, New York 
to Florida to Mississippi, chiefly near the coast. 
ise in the West Indies and South America. July— 

ug. 


2. Stomoisia virgatula Barnhart. Fairy- 


7 wand Bladderwort. Fig. 3875. 
yy) 
‘Ss Utricularia simplex C. Wright, in Sauvalle, Anal. 
re) Acad. Habana 6: 315. 1870. Not R. Br. 1812. 
i U. virgatula Barnh. Bull. Torr. Club 34: 580. 
1908. 


Scape wiry, strict, brownish, 1-8’ high, 1-6- 
flowered, the scales several, minute, acute. 
| Bracts less than 4” long, acute; bracts the 

same length, but narrower; pedicels shorter 

than the bracts; calyx purplish, the upper lobe 

acuminate, 2” long, the lower acute, much 

shorter; corolla yellow, the upper lip barely 

if at all exceeding the upper calyx-lobe, the 

lower of about the same length, with a minute 

‘= tuft of hairs in the throat; spur conic, pen- 
dent, acute, 1-13” long; capsule globose, 1” 


SF in diameter or less, closely invested by the 
L\ SAS UF CS calyx-lobes and beaked by the upper one. 
{ 7) 
\ 


On wet borders of ponds, New York, New Jer- 
WENB sey, Florida and Mississippi. Also in Cuba. 
ww Rare and local. Sept. 


GENUS I. ~ BROOM-RAPE FAMILY. 223 


Family 30. OROBANCHACEAE Lindl. Nat. Syst. Ed. 2, 287. 1830. 
BROOM-RAPE FAMILY. 


Erect, simple or branched, brown yellowish purplish or nearly white root- 
parasites, the leaves reduced to alternate appressed scales, the flowers perfect, 
irregular (rarely cleistogamous ), sessile in terminal bracted spikes, or solitary and 
peduncled in the axils of the scales. Calyx inferior, gamosepalous, 4—5-toothed, 
4-5-cleft, or split nearly or quite to the base on one or both sides. Corolla gamo- 
petalous, more or less oblique, the tube cylindric, or expanded above, the limb 
2-lipped, 5-lobed. Stamens 4, didynamous, inserted on the tube of the corolla and 
alternate with its lobes, a fifth rudimentary one occasionally present; filaments 
slender; anthers 2-celled, the sacs parallel, equal. Ovary superior, r-celled, the 
four placentae parietal; ovules numerous, antropous; style slender ; stigma discoid, 
2-lobed, or sometimes 4-lobed. Capsule 1-celled, 2-valved. Seeds numerous, 
reticulated, wrinkled or striate; embryo minute; cotyledons scarcely differentiated. 


About 11 genera and over 200 species, of wide geographic distribution, mostly in the northern 
hemisphere. 
Flowers all complete and perfect. 

Calyx 2-5-lobed or toothed. 


Calyx about equally 5-cleft; no bractlets on peduncles nor calyx. 1. Thalesia. 
Calyx unequally toothed, or split on both sides; flowers bracteolate. 2. Orobanche. 
Calyx spathe-like, split on the lower side, 3—4-toothed on the upper. 3. Conopholis. 
Lower flowers cleistogamous, fertile; upper complete, mostly sterile. 4. Leptamnium. 


1. THALESIA Raf. Am. Month. Mag. 2: 267. 1818. 
[ANOPLANTHUS Endl. Icon. Gen. Pl. 12. pl. 72. 1838.] 
: [ApHYLLon A. Gray, Man. 290. 1848.] 

Glandular or viscid-pubescent simple-stemmed herbs, parasitic on the roots of various 
plants, with scattered scales, and long-peduncled yellowish white or violet, complete and per- 
fect flowers without bractlets. Calyx campanulate or hemispheric, nearly equally 5-cleft, the 
lobes acute or acuminate. Corolla oblique, the tube elongated, curved, the limb slightly 
2-lipped, the upper lip erect-spreading, 2-lobed, the lower spreading, 3-lobed, the lobes all 
nearly equal. Stamens included; anther-sacs mucronate at the base. Ovary ovoid; placentae 
equidistant, or contiguous in pairs; style slender, deciduous; stigma peltate, or transversely 
2-lamellate. [Dedicated to Thales.] 

About 5 species, natives of North America. Type species: Orobanche uniflora L. 


Stem very short; peduncles 1-4, erect; calyx-lobes lanceolate, acuminate. 1. T. uniflora. 
Stem erect, 2’-5’ high; peduncles several ; calyx-lobes broad, acute. 2. T. fasciculata, 


1. Thalesia unifléra (L.) Britton. Pale or Naked Broom-rape. One-flowered 
Broom-rape. Squaw-drops. Cancer-root. Fig. 3876. 


Orobanche uniflora L. Sp. Pl. 633. 1753. 
Anoplanthus uniflorus Endl. Icon. Gen. Pl. 12. pi. 
He inelelere 
Aphyllon uniflorum T. & G.; A. Gray, Man. 290. 
1848. 
T. uniflora Britton, Mem. Torr. Club 5: 298. 1894. 
Stem usually less than 1’ long, nearly subter- 
ranean, bearing several ovate-oblong scales and 
1-4 slender erect scape-like glandular-puberulent 
naked 1-flowered peduncles 3’-8 high. Calyx 
campanulate, pubescent, glandular, 4’’-5” high, 
less than one-half the length of the corolla, its 
lobes as long as the tube or longer, lanceolate, 
acuminate; corolla white or violet, puberulent 
without, 8-12” long, the curved tube about 3 
times as long as the limb, the short lobes oval 
or obovate, obtuse; placentae nearly equidistant ; 
capsule ovoid, longer than the calyx. 
In woods and thickets, parasitic on the roots of 
various herbs, Newfoundland to Ontario, South 
Carolina and Texas. Far western plants, formerly 


referred to this species, prove to be distinct. 
Pipes. April-June. 


234 OROBANCHACEAE. Von Hi 


2. Thalesia fasciculata (Nutt.) Britton. — 


Clustered or Yellow Cancer-root. 
Fig. 3877. 

Orobanche fasciculata Nutt. Gen. 2: 59. 1818. 

Anoplanthus fasciculatus Walp. Rep. 3: 480. 
1844-45. 

Aphylion fasciculatum A. Gray, Syn. Fl. 2: Part 
iy Ie Ttsiykeys 

Thalesia fasciculata Britton, Mem. Torr. Club 5: 
298. 1894. 


Stem erect, 2-4’ high, densely glandular- 
pubescent, bearing several scales and 3-15 
naked 1-flowered peduncles 17-4’ long. Calyx 
glandular, broadly campanulate, 3-5” high, 
about one-third the length of the corolla, its 
lobes triangular-lanceolate or triangular-ovate, 
acute, equalling or shorter than the tube; co- 
rolla nearly 1’ long, purplish to yellow, puberu- 
lent without, the curved tube 3 times as long 
‘as the limb, the lobes oblong, obtuse, the limb 
more manifestly 2-lipped than in the preced- 
ing species; capsule ovoid to globose. 


In sandy soil, parasitic on the roots of various 
plants, mostly composites, northern Indiana _ to 
Minnesota, Yukon and British Columbia, Ne- 
braska, Arizona and California. April—Aug. 

Thalesia lutea (Parry) Rydb. [Thalesia fasciculata lutea (Parry) Britton] is a race with light 
yellow flowers, growing on grasses in western Nebraska and Wyoming. 


2. OROBANCHE [Tourn.] L. Sp. Pl. 632. 1753. 


Glandular-pubescent, erect simple or branched, reddish yellowish violet or nearly white 
herbs, parasitic on the roots of various plants, with scattered scales, and spicate or racemose, 
complete and perfect, bracted and sometimes bracteolate flowers. Calyx split both above 
and below, nearly or quite to the base, the divisions 2-cleft or rarely entire, or more or less 
unequally 2-5-toothed. Corolla oblique, strongly 2-lipped; upper lip erect, emarginate or 
2-lobed; lower lip spreading, 3-lobed. Stamens included; anther-sacs mostly mucronate at 
the base. Placentae equidistant, or approximate in pairs. Style slender, commonly persistent 
until after the dehiscence of the capsule; stigma peltate to funnelform, entire, or laterally 
2-lamellate. [Greek, Choke-vetch.] 


About 90 species, natives of the Old World and western America. Besides the following some 
6 others occur in the western parts of North America. Type species: Orobanche major i 


Calyx 4-toothed ; stems mostly branched. 1. O. ramosa. 
Calyx split on both sides ; stem simple. 2. O. minor, — 
Calyx 5-cleft ; stem simple. 3. O. ludoviciana. 


1. Orobanche ramésa L. Hemp or Branched 
Broom-rape. Fig. 3878. 


Orobanche ramosa L. Sp. Pl. 633. 1753: 


Plant yellowish; stem rather slender, branched, or 
rarely simple, 3’-15’ high, the scales few and distant, 
2”-5” long. Spike loosely many-flowered, denser 
above than below, the lowest flowers short-pedi- 
celled; bracts usually 3, the longest about equalling 
the calyx; calyx 4-toothed, the teeth triangular- 
ovate, acute, or acuminate, about as long as the tube; 
corolla 5”-9” long, the tube yellow, slightly con- 
stricted above the ovary, the limb bluish. 


Pasasitic on the roots of tomato, hemp and tobacco, 
New Jersey, Illinois, Kentucky. Adventive or natural- 
ized from Europe. Summer. Strangle-tare. 


Orobanche purpirea Jacq., another European species, 
with violet flowers and a 5-toothed calyx, is recorded as 
found on Achillea in lawns a: Wingham, Ontario. 


GENUS 2. BROOM-RAPE FAMILY. 235 


2. Orobanche minor J. E. Smith. Lesser 
or Clover Broom-rape. Herb-bane. 
Fig. 3870. 

O. minor J. E. Smith, Engl. Bot. p/..422. 1797. 


Plant yellowish-brown; stem rather stout, sim- 
ple, 4-20’ high; lower scales numerous, ovate- 
oblong, the upper lanceolate, acute, scattered, 
3-10” long. Spike dense, or the lower flowers 
separated, 3-8 long; bracts I or 2, lanceolate, 
qualling or longer than the flowers; flowers 
5’-9” long; calyx split both above and below, 
each of the lateral segments 2-cleft, the teeth lan- 
ceolate-subulate; corolla-tube yellowish, scarcely 
constricted above the ovary, the limb bluish. 

Parasitic on the roots of clover, New Jersey to 


Virginia. Naturalized from Europe. Called also 
devil’s-root and hell-root. Strangle-tare. May-July. 


3. Orobanche ludoviciana Nutt. Louisiana 
Broom-rape. Fig. 3880. 

Orobanche ludoviciana Nutt. Gen. 2: 58. 1818. 

Aphyllon ludovicianum A. Gray, Bot. Cal. 1: 585. 1876. 

Myzorrhiza ludoviciana Rydb.; Small, Fl. SE. U. S. 

1093. 1903. 

Stems stout, simple, solitary or clustered, viscid- 
puberulent, 4’-12’ high, scaly. Flowers 6-8” long, 
very numerous in dense terminal spikes, I-2-bracteo- 
late under the calyx; calyx 5-cleft, the lobes some- 
what unequal, linear-lanceolate, acuminate, longer 
than the corolla-tube, or shorter; corolla 2-lipped, 
purplish, its tube narrow, about twice as long as the 
limb, the teeth of its lips acute; anthers woolly; 
capsule ovoid-oblong, shorter than the calyx. 


; In sandy soil, Illinois to South Dakota, Saskatche- 
wan, Nebraska, Texas, Arizona and California. Strangle tare. June—Aug. 


3. CONOPHOLIS Wallr. Orobanch. 73s 11825: 


An erect stout simple glabrous, densely scaly, light brown herb, parasitic on the roots of 
trees, with yellowish flowers 2-bracteolate under the calyx, in a thick dense bracted spike, 
the bracts similar to the scales of the stem. Calyx oblique, deeply split on the lower side, 
3-4-toothed on the upper. Corolla strongly 2-lipped, the tube slightly curved, the upper lip 
concave, nearly erect, emarginate, the lower spreading, 3-lobed. Stamens exserted; anther- 
sacs bristly pubescent. Placentae about equidistant; stigma capitate, obscurely 2-lamellate. 
Capsule ovoid-globose. [Greek, signifying a 
scaly cone. ] 


Three known species, the following typical one 
of eastern North America, the other southwestern 
and Mexican. 


1. Conopholis americana (L. f.) Wallr. 
Squaw-root. Fig. 3881. 


Orobanche americana L. f. Suppl. 88. 1767. 
Conopholis americana Wallr. Orobanch. 78. 1825. 


Plants 3-10’ high from a thickened base, light 
brown, usually clustered, covered all over with 
stiff imbricated scales. Upper scales lanceolate 
or ovate, acute, 6’-10” long, the lowest much 
shorter; flowers about 4’ long, exceedingly nu- 
merous in the dense spike which is 6’—10” thick; 
corolla pale yellow, somewhat exceeding the ca- 
lyx; anthers sagittate; capsule ovoid-globose, 
Aen” high. 

In rich woods at bases of trees, Maine to On- 
tario, Michigan, Florida, Alabama and Tennessee. 
Cancer-root. Earth-club. Clap-wort. April—Aug. 


236 OROBANCHACEAE. Vor. III. 


4. LEPTAMNIUM Raf. Am. Month. Mag. 2: 267. Feb. 1818. 
[Epiracus Nutt. Gen. 2: 60. 1818.] 

An erect slender glabrous, purplish or yellowish, rather stiff branching herb, parasitic on 
the roots of the beech, with few small scattered scales, and sessile dimorphous flowers, dis- 
tantly spicate on the branches, the lower cleistogamous, abundantly fertile, the upper com- 
plete but mostly sterile. Calyx short, nearly equally 5-toothed. Corolla of the upper flowers 
cylindric, slightly flattened laterally, the tube much longer than the 4-lobed limb, the upper 
lobe concave, larger than the 3 lower ones; stamens not exserted; anther-sacs mucronulate 
at the base; style filiform, 2-lobed; ovary with an adnate gland on the upper side near the 
base. Corolla of the lower flowers minute, not unfolding, borne like a hood on the summit 
of the ovoid ovary; style very short; placentae contiguous in pairs; capsule at length 2-valved 
at the summit. [Greek, referring to the small calyx.] 


A monotypic genus of eastern North America. 


1. Leptamnium virginianum (L.) Raf. 
Beech-drops. Fig. 3882. 


Orobanche virginiana L. Sp. Pl. 633. 1753. 
Epifagus americana Nutt. Gen. 2: 60. 1818. 
Brae virginiana Bart. Comp. Fl. Phil. 2: 50. 

ISIS. 

Leptamnium virginianum Raf.; A. Gray, Syn. FI. 

2: Part 1, 314. 1878. As synonym. 

Plant 6’-2° high from a thick scaly base, the 
roots brittle, fibrous. Stem paniculately branch- 
ed, the branches straight, ascending; scales 
very few and small; corolla of the upper 
flowers 4”’-5” long and 1” thick, striped with 
purple and white, the limb not at all spread- 
ing; stamens about as long as the corolla; 
style slightly exserted; lower flowers 14” long, 
the corolla resembling the calyptra of a moss; 
capsule 3” high, somewhat oblique and com- 
pressed, many-seeded. 

In beech woods, Nova Scotia and New Bruns- 
wick to Florida, west to Ontario, Michigan, Mis- 
souri and Louisiana. Cancer-root or -drops. 
Virginia brown-rape. Aug.—Oct. 


Family 31. BIGNONIACEAE Pers. Syn. 2: 168. 1807. 
TRUMPET-CREEPER FAMILY. 

Trees, shrubs or woody vines, a few exotic species herbs, with opposite (rarely 
alternate) compound or simple leaves, and mostly large and showy, clustered 
terminal or axillary, more or less irregular flowers. Calyx inferior, gamosepalous. 
Corolla gamopetalous, funnelform, campanulate, or tubular, 5-lobed, somewhat 
2-lipped, at least in the bud. Anther-bearing stamens 2 or 4, inserted on the tube 
of the corolla and alternate with its lobes; anthers 2-celled, the sacs longitudinally 
dehiscent. Disk annular or cup-like. Ovary sessile or stalked, mostly 2-celled; 
placentae parietal, or on the partition-wall of the ovary; ovules very numerous, 
horizontal, anatropous; style slender; stigma terminal, 2-lobed. Capsule 2-valved, 
loculicidally; septifragally or septicidally dehiscent. Seeds flat, transverse, winged 
in our genera; endosperm none; cotyledons broad and flat, emarginate or 2-lobed; 
radicle short, straight. 


About 60 genera and over 500 species of wide geographic distribution in tropical regions, a few 
in the north and south temperate zones. 
Leaves compound; anther-bearing stamens 4; our species vines. 
Calyx-limb undulate; capsule flattened parallel with its partition-wall. 1. Anisostichus. 
Calyx s5-toothed ; capsule compressed at right angles to its partition-wall. 2. Bignonia. 
Leaves simple ; anther-bearing stamens mostly 2; trees. 3. Catalpa. 


1. ANISOSTICHUS Bureau, Mon. Bignon. 43. 1864. 


Woody vines, with opposite 2-foliolate leaves, the terminal leaflet reduced to a tendril, 
and large flowers in axillary cymes. Calyx campanulate, the limb merely undulate, truncate 
or slightly 5-toothed. Tube of the corolla much expanded above the calyx, the limb some- 
what 2-lipped, 5-lobed, the lobes rounded. Anther-bearing stamens 4, didynamous, included, 


GENUS I. TRUMPET-CREEPER FAMILY. 237 


inserted near the base of the corolla; anther-sacs glabrous, divergent. Capsule linear, flat- 
tened parallel with the thin partition, septifragally dehiscent, the margins of the valves more 
or less thickened. Seeds in 2 unequal rows on both margins of the partition, winged, much 
broader than high, the wing entire, or erose at the end. [Greek, unequal-ranked.] 

An apparently monotypic genus. Type species: Bignonia capreolata L. 


1. Anisostichus capreolata (L.) Bureau. 
Tendrilled Trumpet-flower. Cross- 
vine. Fig. 3883. 

? Bignonia crucigera L. Sp. Pl. 624. 1753. 
Bignonia capreolata L. loc. cit. 1753. 
Doxantha capreolata Miers. Proc. Roy. Hort. Soc. 3: 

190. 1863. 

Anisostichus capreolata Bureau, Mon. Bigon. Atlas 8, 

pt. 6. 1864. 

A glabrous woody vine, often climbing to the 
height of 40°-60°, the stems’sometimes 4’ in diameter, 
exhibiting a conspicuous cross in the transverse sec- 
tion. Leaves petioled, commonly with small, simple, 
stipule-like ones in their axils, 2-foliolate, terminated 
by a branched tendril; leaflets stalked, oblong or 
ovate, entire, acute or acuminate at the apex, cordate 
at the base, pinnately veined, 3-7’ long; cymes nu- 
merous, short-peduncled, 2-5-flowered; pedicels 1’—2’ 
long; calyx membranous; corolla 2’ long, orange and 
puberulent without, yellow within; capsule 5’—7’ long, 
8-10” broad, very flat, each valve longitudinally 
I-nerved; seeds broadly winged laterally, narrowly 
winged above’and below, 12’ broad. 

In moist woods, Virginia to Florida, Louisiana, Ohio 
and southern Illinois. Quarter vine. April—June. 


2. BIGNONIA L. Sp. Pl. 633. 1753. 


Climbing woody vines with aerial rootlets, with opposite pinnately compound leaves, and 
large showy red or orange flowers, in terminal corymbs. Calyx tubular-campanulate, some- 
what unequally 5-toothed. Corolla-tube elongated, enlarged above the calyx, narrowly cam- 
panulate, the limb slightly 2-lipped, 5-lobed, the lobes spreading. Anther-bearing stamens 4, 
didynamous, ascending under the upper lip; anther-sacs divergent, glabrous or slightly pubes- 
cent. Capsule elongated, slightly compressed at right angles to the partition, loculicidally and 
septicidally dehiscent. Seeds in several rows on each side of the margins of the partition, 
flat, winged, the wing translucent. [Named after the Abbe Bignon, 1662-1743, librarian to 
to Louis XV.] 


Two known species, the following typical one, the other Japanese. The name Tecoma, used 
for this vine in our first edition, belongs properly to a genus of pinnate-leaved yellow-flowered 
shrubs of tropical America, typified by Tecoma stans (L.) H.B.K. 


1. Bignonia radicans L. Trumpet-flower. 

Trumpet-creeper. Foxglove. Fig. 3884. 
Bignonia radicans L. Sp. Pl. 624. 1753. 

Tecoma radicans DC. Prodr. 9: 223. 1845. 
Campsis radicans Seem. Journ. Bot. 5: 362. 1867. 

A woody vine, climbing to the height of 20°-40° 
or prostrate if meeting no support. Leaves petioled, 
odd-pinnate, not tendril-bearing, 8-15’ long; leaflets 
7-11, ovate to lanceolate, short-stalked, sharply ser- 
rate, reticulate-veined, glabrous, or pubescent on the 
veins beneath, acute or acuminate at the apex, nar- 
rowed at the base, 14’-3’ long; flowers corymbose, 
short-pedicelled ; corymbs 2-0-flowered; calyx coria- 
ceous, about 1’ long; corolla scarlet, 23’ long. the 
tube veined within, 3 times as long as the limb, an- 
ther-sacs glabrous; stigma spatulate; capsule stalked, 
4’-6’ long, 10” in diameter, narrowed at both ends, 
little flattened, ridged above and below by the mar- 
gins of the valves; seeds in several rows on each 
surface of the broad partition, broadly winged lat- 
erally, the wing eroded. 

L In moist woods and thickets, southern New Jersey and 
S== Pennsylvania to Florida and Texas, north to [Illinois 

— and Iowa. Escaped from cultivation further north. 

Trumpet-vine. Trumpet-ash. Cow-itch. Cross-vine. Aug.—Sept. 


238 BIGNONIACEAE. Vor. IIT. 


2 CATALPA Scop. Introd: 170) 917711. 


Trees, or some exotic species shrubs, with opposite or rarely verticillate simple petioled 
leaves, and large showy white or mottled flowers in terminal panicles or corymbs. Calyx . 
closed in the bud, splitting irregularly or into 2 lips in opening. Corolla-tube campanulate 
or obconic, oblique, expanded above, 2-lipped, 5-lobed, the lobes all spreading, their margins 
crisped. Anther-bearing stamens 2, ascending under the upper lip of the corolla; anther-sacs 
glabrous, linear or oblong, divergent; sterile stamens (staminodia) 3, short (or occasionally 
4 perfect didynamous stamens and I staminodium). Disk obsolete. Ovary sessile, 2-celled ; 
ovules in 2-several rows on the sides of the partition. Capsule elongated-linear, terete, 
loculicidally dehiscent. Seeds flat, the large lateral wings dissected into capillary processes. 
[The American Indian name of the first species below.] 


About 5 species, the following in eastern North America, 2 in eastern Asia. Type species: 
Bignonia Catalpa L. West Indian trees referred to this genus prove to be distinct. 


Corolla thickly spotted within, 1’-114’ long, the lobes crimped. 1. C. Catalpa. 
Corolla little spotted, but purple-lined, 2’ long, the lobes nearly flat. 2. C. speciosa. 
tion northward as far as Pennsylvania and southern 


New York. Wood brown, soft, weak, durable in contact 


} . . ° 
with the soil. Weight per cubic foot 28 Ibs. June-July. Cigar-tree. Indian cigar-tree. Catawba. 


1. Catalpa Catalpa (L.) Karst. Catalpa. 
Indian or Smoking Bean. Candle-tree. 
Bean-tree. Fig. 3885. 

Bignonia Catalpa L. Sp. Pl. 622. 1753. 
Catalpa bignomioides Walt. Fl. Car. 64. 1788. 
Catalpa Catalpa Karst. Deutsch. Fl 927. 1880-83. 

A tree, with thin flaky bark, reaching a maximum 
height of about 60° and a trunk diameter of 4°, the 
branches spreading. Leaves strong-scented, broadly 
ovate, entire, or 3-lobed, acute or acuminate at the 
apex, densely pubescent beneath, becoming glabrous 
above, obtuse at the base, 6-12’ long, the lobes, when 
present, acuminate; petioles stout, nearly as long as 
the blade; flowers white, numerous, mottled with 
yellow and purple within, 1-12’ long, in large ter- 
minal erect panicles; capsules 6’-18’ long, 4”-7” 
thick, thin-walled, drooping, the partition narrow. 


In woods in the Gulf States. Escaped from cultiva- 


2. Catalpa specidsa Warder. Catawba 
Tree. Larger Indian Bean. Western 
Catalpa. Fig. 38860. 


Catalpa cordifolia Duham. Nouveau 2: fl. 5. 
1802. Not Moench, 1794. 


Catalpa speciosa Warder; Engelm. Coult. Bot. 
Gaza5e) toss 


A tree, with thick rough bark, reaching a 
maximum height of 120° and a diameter trunk 
of 44°, similar to the preceding species. Leaves 
not unpleasantly scented, broadly ovate, com- 
monly entire, long-acuminate at the apex; 
panicles few-flowered; corolla faintly mottled 
within; capsule thick-walled, 8-20’ long, nearly 
10” in diameter. 


In woods, southern Indiana to Tennessee, west 
to Missouri and Arkansas. Wood brown, soft, 
weak, durable. Weight per cubic foot 26 Ibs. 
Mey June. Cigar-tree. Hardy catalpa. Shawnee- 
wood. 


GENUS I. UNICORN-PLANT FAMILY. 239 


Family 32. MARTYNIACEAE Link, Handb. 1: 504. 1820. 
UNICORN-PLANT FAMILY. 


Herbs, with opposite leaves, or the upper sometimes alternate, and perfect 
irregular flowers, racemose in our species. Calyx inferior, 4-5-cleft or 4-5-parted 
or sometimes split to the base on the lower side. Corolla gamopetalous, irregular, 
the tube oblique, often decurved, the limb slightly 2-lipped, 5-lobed, the lobes 
nearly equal, the 2 upper ones exterior in the bud. Anther-bearing stamens 4, 
didynamous, or the posterior pair sterile; anthers 2-celled, the sacs longitudinally 
dehiscent. Ovary 1-celled, with 2 parietal placentae expanded into broad surfaces, 
or 2—4-celled by the intrusion of the placentae or by false partitions ; ovules numer- 
ous or few in each cavity of the ovary, anatropous; style slender; stigma 2-lobed 
or 2-lamellate. Fruit various in the different genera. Seeds oblong or orbicular, 
compressed, wingless or narrowly winged; endosperm none; embryo large; cotyle- 
dons fleshy, flat ; radicle short and straight. 

Three genera and about 12 species, mainly tropical. 


1. MARTYNIA L. Sp. Pl. 618. 1753. 


Coarse diffusely branched glandular-pubescent and viscid strong-scented herbs, with oppo- 
site or alternate long-petioled leaves, and large violet purple whitish or mottled flowers in 
short terminal racemes. Calyx 1-2-bracteolate at the base, campanulate, inflated, unequally 
5-cleft, deciduous. Corolla funnelform-campanulate, oblique, decurved, the 5 lobes nearly 
equal, spreading. Stamens 4 in our species; anthers gland-tipped, their sacs divergent. Ovary 
1-celled, the 2 parietal placentae intruded and expanded in the center of the cavity into broad 
surfaces bearing the ovules in I or 2 rows. Fruit an incurved beaked loculicidally 2-valved 
capsule, the exocarp somewhat fleshy, the endocarp fibrous, woody, crested below or also 
above, 4-celled by the extension of the placentae. Seeds numerous, tuberculate. [Named for 
John Martyn, 1693-1768, professor of botany at Cambridge, England. ] 


About 8 species, natives of America. Besides the following, 2 others occur in the south- 
western United States. Type species: Martynia annua L. 


1. Martynia louisiana Mill. Unicorn-Plant. 
Elephant’s Trunk. Martinoe. Double- 
claw. Fig. 3887. 


M. louisiana Mill. Gard. Dict. Ed. 8, no. 3. 1768. 
Martynia proboscidea Glox. Obs. 14. 1785. 


ol, AY 
Gey : H 
Annual, densely glandular-pubescent all over; MY 
stem stout, much branched, the branches pros- hf, L ¥ 
trate or ascending, 1°-3° long. Leaves broadly Gf SZ Ai 
ovate to orbicular, rounded at the apex, cordate vt 
at the base, repand, undulate or entire, 3/-12’ in 
diameter, the petiole stout, mostly longer than 
the blade; bractlets at the base of the calyx ob- 
long or linear, deciduous; calyx somewhat cleft 
on the lower side; racemes several-flowered ; 
pedicels slender; corolla whitish or yellowish, 
mottled with purple or yellow within, 14’-2’ long, 
the limb nearly as broad, the lobes obtuse; sta- 
mens all anther-bearing; fruit strongly curved, 
4-6’ long when mature, the beak longer than the 
body, splitting into 2 elastically diverging seg- 
ments, the endocarp crested on the under side 
only. 
In waste places, escaped from gardens, Maine to 
western New York, New Jersey and Georgia. Native 
from Indiana to Iowa, Utah, Texas and New Mexico. July—Sept. 


Family 33. ACANTHACEAE J. St. Hil. Expos. Fam. 1: 236. 180s. 


ACANTHUS FAMILY. 

Herbs, or some tropical genera shrubs or small trees, with opposite simple 
exstipulate leaves, and irregular or nearly regular perfect flowers. Calyx inferior, 
persistent, 4—5-parted or 4—5-cleft, the sepals or segments imbricated, equal or 
unequal. Corolla gamopetalous, nearly regularly 5-lobed with the lobes convolute 
in the bud, or conspicuously 2-lipped. Anther-bearing stamens 4, didynamous, 


240 ACANTHACEAE. Vo. IIT, 


or 2 only; anthers mostly 2-celled, the sacs longitudinally dehiscent. Disk annular 
or cup-like. Ovary 2-celled; ovules 2-10 in each cavity, anatropous or amphitro- 
pous; style filiform, simple; stigmas I or 2. Capsule dry, 2-celled, loculicidally 
elastically 2-valved. Seeds globose or orbicular, not winged, borne on curved pro- 
jections (retinacula) from the placentae, the testa close, mostly roughened, often 
developing spiral threads and mucilage when wetted. I¢ndosperm in the following 
genera none; cotyledons flat, commonly cordate. 


About 175 genera and 2000 species, natives of temperate and tropical regions of the Old World 
and the New. 


Corolla convolute in the bud, nearly regular; stamens 4. 


Ovules 2 in each cavity ; capsule 2—4-seeded. 1. Dyschoriste. 

Ovules 3-10 in each cavity ; capsule 6—20-seeded. 2. Ruellia. 
Corolla imbricated in the bud, strongly 2-lipped; stamens 2. 

Lower lip of the corolla 3-cleft; flowers bracted, not involucrate. 3. Dianthera. 

Lower lip of the corolla entire or 3-toothed; flowers involucrate. 4. Diapedium. 


1. DYSCHORISTE Nees, in Wall. Pl. As. Rat. 3:/81- aeg2 
[CaLOpHANES Don; Sweet, Brit. Fl. Gard. (11). pl. 787. 1833.] 

Erect or procumbent perennial herbs or shrubs, with entire leaves (smaller ones some- 
tibes fascicled in their axils), and blue or purple, rather large, bracted flowers, axillary, 
solitary or clustered. Calyx deeply 5-cleft, the lobes setaceous. Corolla funnelform, the 
tube slightly curved, enlarged above, or cylindraceous, the limb spreading, 5-lobed, somewhat 
2-lipped, the lobes rounded, sinistrorsely convolute in the bud. Stamens 4, didynamous, 
included, all anther-bearing in our species; anther-sacs mucronate at the base. Ovules 2 in 
each cell of the ovary; summit of the style recurved; stigma simple, or of 2 unequal lobes. 
Capsule oblong, linear, narrowed at the base, 2-4-seeded, in some species coherent. Seeds flat, 
orbicular, attached by their edges to the retinacula. [Greek, referring to the coherent capsule- 
valves. ] 


About 30 species, of wide distribution in warm and tropical regions. Besides the following, 
4 others occur in the southern and southwestern United States. Type species: Dyschoriste 
depressa Nees. 


1. Dyschoriste oblongifolia (Michx.) 
Kuntze. Dyschoriste. Fig. 3888. 
Ruellia biflora L. Sp. Pl. 633. 1753? 
R. oblongifolia Michx. Fl. Bor. Am. 2: 23. 1803. 
Calophanes oblongifolia Don; Sweet, Brit. Fl. 
Gard. (II.) pl. r8r. 1833. 
Ea biflorus Nees, Linnaea 16: 294. 
I 2 
D. EU onetelee Kuntze, Rev. Gen. Pl. 486. 1891. 
Rootstocks horizontal, slender; stems slen- 
der, erect, simple, or branched below, pubes- 
cent or puberulent, obtusely 4-angled, 6’-15’ 
high, rather stiff. Leaves ascending or erect, 
oblong or oval, rounded at the apex, somewhat 
narrowed at the base, very short-petioled, or 
sessile, pubescent or glabrate, 8-15” long; 
flowers commonly solitary in the axils; calyx- 
segments filiform, hirsute, exceeding the ob- 
long, obtuse bractlets; corolla blue, or mottled 
with purple, 8’—-12” long, slightly 2-lipped, its 
tube enlarged above; capsule oblong, about 
one-half the length of the calyx; anther-sacs 
mucronate-aristate at the base. 
In sandy pine barrens, Virginia to Florida, 
mainly near the coast. June—Sept. 


2. RUELLIA [Plumier] L. Sp. Pl. 634. 1753. 


Perennial herbs, or shrubs, mostly pubescent, with entire or rarely dentate leaves, and 
large violet blue white or yellow flowers, solitary or clustered in the axils, or cymose in 
terminal panicles, or sometimes cleistogamous and inconspicuous. Calyx 5-cleft, or 5-parted, 
the segments narrow. Corolla funnelform or salverform, the tube usually narrow, slightly 
enlarged above, the limb spreading, 5-lobed, the lobes obtuse, mostly nearly equal, sinistrorsely 
convolute in the bud. Stamens 4, included or exserted; anther-sacs not mucronate at the 
base. Ovules 3-10 in each cavity of the ovary; apex of the style recurved; stigma simple, or 
of 2 unequal lobes. Capsule oblong or club-shaped, compressed or terete, 6-20-seeded. Seeds 
compressed, ovate or orbicular, attached by their edges to the retinacula. [Named for I. 
Ruel or de la Ruelle, 1474-1537, an early French herbalist. ] 


GENUS 2. ACANTHUS FAMILY. 241 


About 200 species, mainly of tropical America, a few in Africa, Asia and Australia. Besides 
the following, some 6 others occur in the southern and southwestern United States. Type species: 
Ruellia tuberosa L. 


Flowers sessile or nearly so. ; 
Calyx-segments linear-lanceolate, scarcely exceeding the capsule. 1. R. strepens. 
Calyx-segments filiform-linear, exceeding the capsule. 

Leaves sessile or very short-petioled. 
Leaves slender-petioled. 
Flowers peduncled; peduncle with two large bracts at the summit. 


. R. ciliosa, 
. R. parviflora. 
. R. pedunculata. 


BwWh 


1. Ruellia strépens L. Smooth or Short-tube Ruellia. Fig. 3889. 
Ruellia strepens L. Sp. Pl. 634. 1753. 


Dipteracanthus micranthus Engelm. & Gray, Bost. 
Journ. Nat. Hist. 5: 257. 1845. 


Ruellia strepens var. cleistantha A. Gray, Syn. FI. 
eeeeatt m3275 tae: 


Glabrate or somewhat pubescent; stem erect, 
slender, simple, or branched, 4-sided, »1°-4° 
high, the branches ascending. Leaves oblong, 
oval, or ovate, petioled, acute or subacute at 
the apex, narrowed at the base, 3-6’ long; peti- 
oles 2’-10” long; flowers solitary or several 
together in the axils,some often cleistogamous ; 
calyx-segments linear-lanceolate, shorter than 
the corolla-tube, slightly pubescent or ciliate, 
8-12” long, #”-1” wide; corolla blue, 13’-2’ 
long, the limb nearly as broad, the tube about 
as long as the throat and limb; capsule club- 
shaped, longer than or equalling the calyx. 


In dry woods, Pennsylvania to Wisconsin, Kan- 
sas, Florida and Texas. May-July. 


2. Ruellia cilidsa Pursh. Hairy or Long- 
tube Ruellia. Fig. 3890. 


Ruellia ciliosa Pursh, Fl. Am. Sept. 420. 1814. 


Hirsute or pubescent, at least above; stem 
ascending, rather stout, 1°-23° high. Leaves 
hairy, ciliate, oblong, oval, or ovate, sessile or 
nearly so, obtuse or subacute at the apex, nar- 
rowed at the base, 14-3’ long; flowers clus- 
tered or solitary in the axils, sometimes cleis- 
togamous; calyx-segments filiform, hirsute, 
10-12” long, about #” wide; corolla blue, 
14’-2’ long, the tube equalling or longer than 
the obconic throat and nearly regular limb 
and about twice as long as the calyx; capsule 
shorter than the calyx. 


In dry soil, southern New Jersey and Pennsyl- 
vania to Florida, west to Michigan, Kansas and 
Louisiana. June—Sept. 


16 


ACANTHACEAE. Vo. III. 


3. Ruellia parviflora (Nees) Britton. 
Slender Hairy Ruellia. Fig. 3891. 


Dipteracanthus ciliosus parviflorus Nees, Linnaea 
16: 294. 1842. 

R. cilosa ambigua A. Gray, Syn. FI. 2': 326. 1878. 

Ruellia ciliosa parviflora Britton, in Britton & 
Brown, Ill. Fl. Ed. 1, 3: 203. 1898. 

R. parvifiora Britton, Man. 854. 1901. 


Sparingly finely pubescent, but green; stem 
erect, 6-18’ tall, simple or sometimes spar- 
ingly branched. Leaves minutely pubescent, 
ciliolate, oblong to oblong-lanceolate, or some- 
times oval, 14-4’ long, obtuse or .acutish, un- 
dulate, rather acuminately narrowed at the 
base, manifestly petioled; flowers solitary or 
clustered in the axils, the pedicels very short; 
calyx-segments linear-filiform, bristly, 8”’-12” 
long; corolla blue, 13-2’ long, the tube some- 
what longer than the throat and somewhat ex- 
ceeding the calyx, the limb #’-12’ broad; cap- 
sule oblong above the stipe-like base, barely 
8” long. 

In sandy soil, Maryland to Indiana, Florida and 
Texas. May—Aug. 


4. Ruellia pedunculata Torr. 
Ruellia. Fig. 3892. 


Ruellia pedunculata Torr.; A. Gray, Syn. FI. 
Dea hey S20: ema Gs 


Finely pubescent; stem erect, 1°—23° tall, 
the branches spreading. Leaves ovate to 
oblong-lanceolate, acute or acuminate at 
the apex, narrowed at the base, short- 
petioled, the larger 2’-3’ long; peduncles 
slender, spreading, 1-3’ long, with 2 leaf- 
like bracts at the summit which subtend a 
solitary flower, or 2 or 3 slender-pedicelled 
ones with pedicels similarly bracted; calyx- 
segments awn-like, equalling the narrow 
corolla-tube, or shorter; corolla funnel- 
form, 14’-2’ long; capsule about 10” long, 
puberulent, longer than the calyx. 


In dry soil, Illinois and Missouri to Arkan- 
sas and Louisiana. June—Sept. 


3. DIANTHERA L. Sp. Pl. 27. 
1753. 


Herbs mostly perennial, with entire or rarely dentate leaves, and small or large very 
irregular flowers, variously clustered or solitary in the axils. Calyx deeply 4~-5-parted, the 
segments narrow. Corolla-tube slender, short or elongated, curved or nearly straight, the 
limb conspicuously 2-lipped; upper lip interior in the bud, erect or ascending, concave, entire, 
or 2-dentate; lower lip spreading, 3-cleft. Stamens 2, inserted on the throat of the corolla, 
not exceeding the upper lip; anther-sacs ovate or oblong, slightly divergent, not mucronate, 
separated by a rathed broad connective. Ovules 2 in each cavity of the ovary; style slender; 
stigma entire, or 2-lobed. Capsule contracted at the base into a long stipe, about 4-seeded. 
Seeds flat, orbicular or ovate, the placentae not separating from the walls of the capsule. 
[Greek, double anthers. ] 


About 100 species, natives of tropical America, a few in tropical Asia and Africa. Besides the 
following, chee ae occur in the southern and southwestern United States. Type species: Dianthera 
americana L. 


Flowers capitate, the heads dense, at length oblong. 1. D. americana. 
Flowers in loose spikes. 
Flowers in pairs in the spikes. 2. D. ovata. 


Flowers scattered singly along the spikes. 3. D. lanceolata. 


GENUS 3. ACANTHUS FAMILY. 


1. Dianthera americana L. Dense- 
flowered Water Willow. Fig. 3893. 


Dianthera americana L. Sp. Pl. 27. 1753. \ 


Perennial, glabrous; stem erect, grooved and 
angled, slender, usually simple, 1°-3° high, or #4 
sometimes 6° long when growing in water. @ 
Leaves lanceolate or linear-lanceolate, grad- 
ually acuminate, 3-6’ long, 3-8” wide, entire, 
narrowed at the base into short petioles, or 
sessile; flowers violet, or nearly white, eapi- 
tate-spicate at the ends of slender axillary 
peduncles which are shorter than or equal to 
the leaves; bractlets linear-subulate, shorter 
than the flowers; corolla 5’—6” long, its tube 
shorter than the lips, the base of the lower 
lip rough and palate-like; capsule 6” long, 
exceeding the calyx, its stipe about the length 
of the slightly compressed body. 

In water and wet places, Quebec to Ontario, 
Michigan, Georgia and Texas. May—Aug. 


FL 


PP 
ly 
WHTNN 


D 


S 


2. Dianthera ovata Walt. Loose-flow- 
ered Water Willow. Fig. 3894. 
Dianthera ovata Walt. Fl. Car. 63. 1788. 
Perennial, glabrous; stem ascending or erect 
from a horizontal base, slender, 6-20’ high, 
simple, or sparingly branched. Leaves short- 
petioled, or sessile, ovate, oblong or oval, 1’—3’ 
long, 8-18” wide; flowers opposite in loose 
slender-peduncled axillary spikes, which be- 
come I-3 long; peduncles shorter than or 
but little exceeding the leaves; calyx-segments 
narrowly linear, much longer than the bracts 
and bractlets; corolla pale purple, 4-5” long; 


Narrow-leaved Water Willow. Fig. 3895. 


D. ovata lanceolata Chapm. Fl. S. States 304. 1860. 
D. lanceolata Small; Britton, Man. 855. tgo1. 

Perennial, puberulent; stem erect or spreading, 
more or less branched, 4-12’ long. Leaves linear 
to linear-elliptic or elliptic-lanceolate, 14’-4’ long, 
more or less acuminate, undulate, sessile or nearly 
so; flowers in interrupted slender spikes 14-4’ 
long; calyx-segments narrowly linear, 23’—33” 
long; corolla whitish or pale-purple, about 5” 
long, the lips nearly as long as the tube, which 
is saccate near the base, the upper lip truncate 
or retuse, the lower one 3-lobed, the middle lobe 
truncate or retuse, the lateral ones obtuse; cap- 
sule 7-8” long, the body as long as the stipe-like 
base or shorter. 


capsule about 6” long. 

In wet soil, especially along streams, southern 
Virginia to Florida. Records from Arkansas and 
Texas apply to the following species. June—Aug. 

“A 


In swamps and low grounds, Missouri to Tennessee, Florida and Texas. June—Sept. 


4. DIAPEDIUM Konig; Konig & Sims, Ann. Bot. 2: 189. 1806. 


{DrctreTerRA Juss. Ann. Mus. Paris, 9: 267. 1807.] 
Erect or diffuse branched pubescent or glabrous herbs, with entire petioled leaves, and 
blue red or violet flowers, subtended by involucres of 2-4 distinct or connate bracts, the 


244 ACANTHACEAE. Vo. ITI. 


inflorescence mostly cymose or spicate, the involucres subtending 1 flower or several. Calyx 
4-5-cleft, the lobes linear or subulate. Corolla-tube slender, slightly enlarged above, the 
limb conspicuously 2-lipped ; upper lip erect, concave, interior in the bud, entire or 2-3-toothed ; 
lower lip spreading, entire or 3-toothed. Stamens 2; anther-sacs parallel, sometimes unequal, 
separated by a narrow connective. Style filiform; ovules 2 in each cavity of the ovary. 
Capsule flattened, ovate or suborbicular, sessile or stipitate, 2-4-seeded. Placentae separating 
elastically from the walls of the capsule. Seeds compressed, nearly orbicular. 


About 60 species, natives of warm and tropical regions. Besides the following, 4 others occur 
in the southern and southwestern United States. Type species: Justicia chinensis L. 


1. Diapedium brachiatum (Pursh) 
Kuntze. Diapedium. Fig. 3896. 
Justicia brachiata Pursh, Fl. Am. Sept. 13. 1814. 
Dicliptera brachiata Spreng. Syst. 1: 86. 1825. 
D. brachiatum Kuntze, Rev. Gen. Pl. 485. 1891. 

Annual (?), glabrate, or pubescent; stem 
slender, 6-grooved, erect, much branched, 1°-2° 
high. Leaves ovate, membranous, long-petioled, 
acuminate or acute at the apex, narrowed or 
rounded at the base, 3-6’ long, 1-3’ wide; 
inflorescence paniculate,° the involucres I-4- 
flowered, each of 2 oblong or obovate, obtuse 
or mucronate, opposite bracts; corolla 8”’—12” 
long, pink or purple, the lips about as long as 
the slender tube; upper lip 2-3-toothed, the 
lower entire; capsule oblong, 2’-3” high, a 
little longer than the involucre, the valves 
slightly divergent in dehiscence, the placentae 
remaining attached to their summits. 


In moist thickets, North Carolina to Florida, 
Missouri, Kansas and Texas. July—Oct. 


Family 34. PHRYMACEAE Schauer 
in DC. Prodr. rr: 520, “weag 


LopsEED FAMILY. 


An erect perennial herb with divaricate branches, opposite membranous simple 
leaves, and small irregular purplish flowers, distant in slender elongated spikes. 
Calyx cylindric, 2-lipped; upper lip 2-cleft, the teeth setaceous; lower lip much 
shorter, 3-toothed, the teeth subulate. Corolla-tube cylindric, the limb 2-lipped; 
upper lip erect, concave, emarginate; lower lip larger, spreading, convex, 3-lobed, 
the lobes obtuse. Stamens 4, didynamous, included. Ovary oblique, 1-celled; 
ovule 1, orthotropous, ascending; style slender; stigma 2-lobed. Calyx reflexed 
in fruit, enclosing the dry achene, becoming prominently ribbed, closed and its 
teeth hooked at the ends. Cotyledons convolute; radicle superior. 

Consists of the following monotypic genus of eastern 


North America, eastern and central Asia. [Name un- 
explained. ] 


1. PHRYMA L. Sp. Pl. 601. 1753. 


1. Phryma Leptostachya L. Lopseed. 
Fig. 3897. 
Phyrma Leptostachya L. Sp. Pl. 601. 1753. 


Puberulent; stem 13°-3° high, somewhat 4-sided, 
sometimes constricted above the nodes, branched 
above, the branches slender, elongated, divergent. 
Leaves ovate, acute or acuminate at the apex, obtuse 
of narrowed at the base, very thin, coarsely dentate, 
2’-6’ long, the lower petioled, the upper often nearly 
sessile; spikes very narrow, 3-6’ long; flowers about 
3” long, mostly opposite, distant, borne on very short 
minutely 2-bracteolate pedicels, at first erect, soon 
spreading, the calyx, after flowering, abruptly re- 
flexed against the axis of the spike. 

In woods and thickets, New Brunswick to Manitoba, 
Florida and Kansas. Bermuda. Eastern Asia. June—Aug. 


GENUS 1. PLANTAIN FAMILY. 245 


Family 35. PLANTAGINACEAE Lindl. Nat. Syst. Ed. 2, 267. 1836. 
PLANTAIN FAMILY. 

Annual or perennial, mostly acaulescent or short-stemmed, rarely stoloniferous 
herbs, with basal, or, in the caulescent species, opposite or alternate leaves, and 
small perfect polygamous or monoecious flowers, bracteolate in dense terminal 
long-scaped spikes or heads, or rarely solitary. Calyx 4-parted, inferior, per- 
sistent, the segments imbricated. Corolla hypogynous, scarious or membranous, 
mostly marcescent, 4-lobed. Stamens 4 or 2 (only I in an Andean genus), inserted 
on the tube or throat of the corolla; filaments filiform, exserted or included; 
anthers versatile, 2-celled, the sacs longitudinally dehiscent. Ovary sessile, supe- 
rior, 1—2-celled, or falsely 3—-4-celled. Style filiform, simple, mostly longitudinally 
stigmatic. Ovules 1-several in each cavity of the ovary, peltate, amphitropous. 
Fruit a pyxis, circumscissile at or below the middle, or an indehiscent nutlet. 
Seeds 1-several in each cavity of the fruit; endosperm fleshy ; cotyledons narrow. 

Three genera and over 225 species, of wide geographic distribution. 


Flowers in terminal spikes or heads; fruit a pyxis. 1. Plantago. 
Flowers monoecious, the staminate solitary, peduncled, the pistillate sessile among the linear leaves; 
fruit indehiscent. 2. Littorella. 


1. PLANTAGO [Tourn.] L. Sp. Pl. 112. 1753. 


Acaulescent or leafy-stemmed herbs, the scapes arising from the axils of the basal or 
alternate leaves, bearing ‘terminal spikes or heads of small greenish or purplish flowers 
(flowers solitary in a few exotic species). Calyx-segments equal, or two of them larger. 
Corolla salverform, the tube cylindric, or constricted at the throat, the limb spreading in 
anthesis. Fruit a pyxis, mostly 2-celled. Seeds various. [The Latin name.] ; 

Over 200 species, of wide distribution. Besides the following, some 6 others occur in western 
North America. Known as Plantain, Ribwort or Roadweed. Type species: Plantago major L. 

* Plants acaulescent; flowers spicate or capitate at the ends of scapes. 
Corolla-lobes spreading or reflexed in fruit, not closed over the top of the pyxis. 
Leaves ovate, lanceolate or oblong. 
Seeds several or many in each pyxis. 
Pyxis ovoid, circumscissile at about the middle. 1. P. major. 
Pyxis oblong, circumscissile much below the middle. 2. P. Rugelii. 
Seeds 2-4 in each pyxis. 
Leaves all narrowed at the base, parallel-ribbed. 


Seeds excavated on the inner side. 3. P. lanceolata. 
Seeds flat or but slightly concave on the inner side. 
Spike very dense; leaves pubescent. 4. P. media. 
Lower flowers scattered ; leaves glabrous or very nearly so. 5. P. eriopoda. 
Leaves, or some of them, cordate; veins starting from the midrib. 6. P. cordata. 
Leaves linear or filiform. 
Leaves fleshy ; plant maritime. 7. P. maritima, 
Leaves not fleshy ; plants not maritime. 
Spike densely tomentose; bracts usually not longer than flowers. 8. P. Purshii. 
Plant green and glabrate; bracts much longer than the flowers. g. P: aristata. 
Corolla-lobes erect and closed over the top of the pyxis. 
Leaves spatulate to obovate; stamens 4. 10. P. virginica. 
Leaves linear-filiform ; stamens 2. 
Capsule about 4-seeded, slightly exceeding the calyx. 11. P. pusilla. 
Capsule 7—30-seeded, twice as long as the calyx. 12. P. heterophylla. 
** Stem erect, leafy; flowers capitate at ends of axillary peduncles. 13. P. arenaria. 


1. Plantago major L. Common or Greater 
Plantain. Dooryard Plantain. Fig. 3808. 


Plantago major L. Sp. Pl. 112. 1753. 

Perennial, glabrous or somewhat pubescent; rootstock 
short, thick, erect. Leaves long-petioled, rather firm in 
texture, mostly ovate, obtuse or acutish, entire, or coarsely 
dentate, 1-10’ long, 3-I1-ribbed; scapes 2’-3° high; spike 
linear-cylindric, usually very dense, commonly blunt, 2’—10' 
long, 3’’-4” thick; flowers perfect, proterogynous; sepals 
broadly ovate to obovate, scarious on the margins, one- 
half to two-thirds as long as the ovoid obtuse or subacute, 
5-16-seeded pyxis, which is circumscissile at about the 
middle; stamens 4. 

In waste places, nearly throughout North America. In part 
naturalized from Europe, but indigenous in the North and on 
salt meadows. Also in the West Indies. Small leaves are occa- 
sionally borne near the bases of the spikes; spikes rarely 
branched. May-—Sept. Way-side or round-leaf plantain. Broad- 
leaf. Hen-plant. Lamb’s-foot. Way-bread. Healing-blade. 


246 PLANTAGINACEAE. Vot. III. 


Plantago halophila Bicknell, of saline situations along the Atlantic coast, is densely pubescent, 
but otherwise like this species. A similar race occurs about lakes in northern New York. 


Plantago asiatica L. is a boreal race with thinner 
nearly erect Jeaves. 


2. Plantago Rugeélii Dcne. Rugel’s or Pale 
Plantain. Fig. 3899. 
Plantago Rugelii in DC. Prodr. 13: Part 1, 700. 1852. 

Similar to the preceding species, the spikes ‘tess 
dense, at least toward the base, usually long-atten- 
uate at the summit, sometimes 5” thick. Sepals ob- 
long, prominently keeled on the back, the margins 
green or scarious; pyxis oblong-cylindric, 2’-3” 
long, twice as long as the sepals, circumscissile much 
below the middle and entirely within the calyx, 
4-10-seeded; stamens 4. 

In fields, woods and waste places, New Brunswick to 
Ontario, South Dakota, Florida, Kansas and Texas. 
Petioles commonly purple at the base. Usually brighter 
green and with thinner leaves than P. major. This 
species, or the preceding one, was known to the Indians 
as “ White-man’s-foot.” Silk-plant. June—Sept. 

3. Plantago lanceolata L. Ribwort. Rib- 

grass. English, Buck, or Buckthorn 


Snake, Lance-leaved, or Ripple Plantain. Fig. 3900. 
Plantago lanceolata L. Sp. Pi aa32) 2753. i 


Perennial or biennial, more or less pubescent; 
rootstock mostly short, with tufts of brown hairs 
at the bases of the leaves. Leaves narrowly ob- 
long-lanceolate, mostly erect, shorter than the 
scapes, entire, acute or acuminate at the apex, 
gradually narrowed into petioles, 3-5-ribbed, 2 
12’ long, 3-12” wide; scapes slender, channelled, 
sometimes 23° tall; spikes very dense, at first 
short and ovoid, becoming cylindric, blunt and 
3’-4’ long in fruit, 4-6” thick; flowers perfect, 
proterogynous; sepals ovate, with a narrow green 
midrib and broad scarious margins, the two lower 
ones commonly united; corolla glabrous, its tube 
very short; filaments white; pyxis oblong, very 
obtuse, 2-seeded, slightly longer than the calyx; 
seeds deeply excavated on the face. 


In fields and waste places, New Brunswick to the 
Northwest Territory, British Columbia, Florida and 
Kansas. Bermuda. Naturalized from Europe; native 
also of Asia. Sepals rarely metamorphosed into 
small leaves. April—Nov. Black-jacks. Jack-straws. 
Dog’s-ribs. Cocks. Kemps. Leechwort. Ram’s-tongue. Rattail. Windles. Long-plantain. Ripple- 
grass. Kempseed. Headsman. Hen-plant. Clock. Chimney-sweeps. Cats’-cradles. 


4. Plantago média L. Hoary Plantain. 
Lamb’s-tongue. Healing herb. Fig. 3901. 
Plantago media L. Sp. Pl. 113. 1753. 


Perennial, intermediate in aspect between P. major 
and P. lanceolata, the short rootstock clothed with 
brown hairs among the bases of the leaves. Leaves 
spreading, ovate, broadly oblong or elliptic, obtuse or 
acute at the apex, entire, or repand-dentate, densely 
and finely canescent, 5-7-ribbed, narrowed at the base 
into margined, usually short petioles; scapes slender, 
much longer than the leaves, 1°-2° tall; spikes very 
dense, cylindric and 1-3’ long in fruit, about 3” 
thick; flowers perfect, white; sepals all distinct, ob- 
long, with a narrow green midrib and broad scarious 
margins; corolla glabrous; stamens pink or purple; 
pyxis oblong, obtuse, about as long as the calyx, 
2-4-seeded, the seeds merely concave, stamens 4. 

In waste places, Maine, Rhode Island, Ontario and 
New York. Adveéntive from Europe. Native also o 
Asia. May-Sept. Fire-leaves. Fire-weed. Lamb’s-lettuce. 


GENUS I. PLANTAIN FAMILY. 


5. Plantago eridpoda Torr. Saline Plantain. 
Fig. 3902. 

Plantago glabra Nutt. Gen. 1: 100. 1818? 

Plantago eriopoda Torr. Ann. Lyc. N. Y.2: 237. 1827. 

Perennial, succulent ; rootstock long, usually densely 
covered with long brown hairs among the bases of 
the leaves. Leaves oblong, oblong-lanceolate, or ob- 
lanceolate, entire, or repand-dentate, acute at the 
apex, narrowed into petioles, 5-9-ribbed, 3-12’ long, 
3’-1}’ wide, glabrous or very nearly so; scapes stout, 
more or less pubescent, longer than the leaves, 6’-18’ 
high; spikes 1’-5’ long, dense above, the lower flowers 
scattered; flowers perfect; sepals distinct, oblong- 
obovate with a narrow green midrib and broad scari- 
ous margins; corolla glabrous, its lobes spreading or 
reflexed; pyxis ovoid-oblong, very obtuse, one-third 
longer than the calyx, 2-4-seeded, circumscissile be- 
low the middle, seeds nearly flat. 

In maritime or saline soil, Nova Scotia and Quebec 
to Minnesota, Athabasca, Cclorado and California. 
June-Sept. 

Plantago sparsiflora Michx., of the Southeastern 
States, admitted into our first edition as found in southern 
Illinois, is not known to range north of North Carolina. 


very stout. 


July. 


6. Plantago cordata Lam. 


margined above; 


Heart-leaved 
Plantain. Water Plantain. Fig. 3903. 


Plantago cordata Lam. Tabl. Encycl. 1: 338. 1791. 


Perennial, glabrous, purple-green ; rootstock short, 
Leaves broadly ovate or nearly orbicu- 
lar, pinnately veined, obtuse or acute at the apex, 
entire or dentate, rounded, abruptly narrowed, or 
cordate at the base, often 10’ long; petioles stout, 
stout, longer than the 
leaves; spikes loosely flowered, sometimes I° long, 
with interrupted clusters; flowers perfect; sepals 
ovate to obovate, obtuse, green; corolla-lobes spread- 
ing; pyxis ovoid-globose, obtuse, circumscissile at 
or slightly below the middle, 1-4-seeded; seeds not 
excavated on the face. 


Tn swamps and along streams, Ontario to New York, 
Alabama, Minnesota, Missouri and Louisiana. March— 


7. Plantago maritima L. Sea or Seaside Plantain. Fig. 3904. 


Plantago maritima L. Sp. Pl. 114. 1753. 
Plantago decipiens Barneoud, Mon. Plantag. 16. 1845. 
P. borealis Lange, Fl. Dan. pl. 2707. 


Annual, biennial, or perennial, fleshy; rootstock 
stout or slender, sometimes with tufts of whitish 
hairs among the bases of the leaves. Leaves linear, 
glabrous, very obscurely nerved, sessile, or narrowed 
into short margined petioles, 2-10’ long, entire, or 
with a few small teeth, 1-23” wide; scapes slender, 
more or less pubescent, longer than or equalling the 
leaves; spikes dense, linear-cylindric, blunt, 1-5’ 
long; flowers perfect; sepals ovate-lanceolate to 
nearly orbicular, green, somewhat keeled; corolla 
pubescent without, its lobes spreading; pyxis ovoid- 
oblong, obtuse, 2-4-seeded, circumscissile at about 
the middle, nearly twice as long as the calyx; seeds 
nearly flat on the face. 


In salt marshes and on sea-shores, Greenland and 
Labrador to New Jersey, and on the Pacific Coast from 
Alaska to California. Also on the coasts of Europe and 
Asia. Consists of several slightly differing races. 
Buckshorn. Gibbals. Sea-kemps. June—Sept. 


248 PLANTAGINACEAE. Vor. III, 


8. Plantago Purshii R.& S. Pursh’s Plantain. Fig. 3905. 

se Plantago Purshii R. & S. Syst. 3: 120. 1818. 

E Plantago gnaphalioides Nutt. Gen. 1: 100. 1818. 
Plantago patagonica var. gnaphalioides A. Gray, Man. 

Ed. 2, 269. 1856. 

Annual, woolly or silky all over, pale green; 
scapes slender, 2-15’ tall, longer than the leaves. 
Leaves ascending, linear, acute or acuminate at 
the apex, narrowed into margined petioles, I-3- _ 
nerved, 13-4” wide, entire, or very rarely with 
a few small teeth; spikes very dense, cylindric, 
obtuse, 1-5’ long, about 3” in diameter, exceed- 
ingly woolly; bracts rigid, equalling or slightly 
exceeding the flowers; flowers perfect but heter- 
ogonous, many of them cleistogamous; sepals 
oblong, obtuse, scarious-margined; corolla-lobes 
broadly ovate, spreading; stamens 4; pyxis ob- 
long, obtuse, 13” long, little exceeding the calyx, 
2-seeded, circumscissile at about the middle; seeds 
convex on the back, deeply concave on the face. 

On dry plains and prairies, Indiana to western On- 
tario, British Columbia, Texas and northern Mexico. 
Locally adventive eastward. May-Aug. Salt-and- 
pepper-plant. 


g. Plantago aristata Michx. Large-bracted 
Plantain. Fig. 3906. 

Plantago aristata Michx. Fl. Bor. Am. 1: 95. 1803. 

Plantago patagonica var. aristata A. Gray, Man. Ed. 2, 

269. 1856. 

Annual, dark green, villous, or glabrate; scapes 
stout, erect, 6-18’ tall, exceeding the leaves. Leaves 
linear, acuminate at the apex, entire, narrowed intc 
slender petioles, sometimes prominently 3-ribbed 
13-4” wide; spikes very dense, cylindric, 1-6’ long. 
pubescent but not woolly; bracts puberulent, linear, 
elongated, ascending, the lower often Io times as 
long as the flowers; flowers very similar to those 
of the preceding species; pyxis 2-seeded; the seeds 
concave on the face. 

On dry plains and prairies, Illinois to South Dakota, 
Louisiana and Texas, west to British Columbia and New 
Mexico. Also widely adventive as a weed in the East- 
ern States from Maine to Georgia, its eastern natural 
limits now difficult to determine. May—Oct. 

Plantago spinuldsa Dene., differing in its pale green 
color and mostly shorter involucral bracts, enters the 
western part of our area in Nebraska. 


10, Plantago virginica L. Dwarf or White Dwarf Plantain. Fig. 3907. 
Plantago virginica L. Sp. Pl. 113. 1753. 

Annual or biennial, pubescent or villous; scapes 
erect, slender, 17-18’ high, much longer than the 
leaves. Leaves spatulate or obovate, obtuse or 
acutish, thin, entire, or repand-denticulate, nar- 
rowed into margined petioles, or almost sessile, 
varying greatly in size, 3-5-nerved, ascending or 
spreading; spikes very dense, or the lower flowers 
scattered, linear-cylindric, obtuse, 3’—4” thick, usu- 
ally 1-4’ long, but in dwarf forms reduced to 2-6 
flowers; flowers imperfectly dioecious; corolla- 
lobes of the fertile plants erect and connivent on 
the top of the pyxis, those of the sterile widely 
spreading; stamens 4; pyxis oblong, about as long 
as the calyx, appearing beaked by the connivent 
corolla-lobes, 2—4-seeded, the seeds yellow to brown. 

In dry soil, Rhode Island to Florida, Illinois, Mich- 
igan, Missouri, Arizona and northern Mexico. Ber- 
muda. March—July. 

Plantago rhodospérma Dcne., of the Southwest, 
with larger flowers and larger reddish seeds, is re- 
corded as adventive in Missouri. 


GENUS I. PLANTAIN FAMILY. 


11. Plantago pusilla Nutt. Slender Plantain. 
Fig. 3908. 
Plantago pusilla Nutt. Gen. 1: 100, 1818, 


Annual, puberulent; scapes filiform, 2-7’ high, 
longer than the linear-filiform, mostly entire, blunt- 
pointed obscurely t-nerved leaves. Leaves about 2” 
wide; spikes slender, linear, rather loosely flowered, 
2-3’ long, 13-2” thick; flowers imperfectly dioecious 
or polygamous; sepals oblong, obtuse, about as long 
as the bract, scarious-margined; corolla-lobes of the 
more fertile plants becoming erect over the pyxis; 
stamens 2; pyxis ovoid-oblong, obtuse, one-fourth to 
one-third longer than the calyx, about 4-seeded, cir- 
cumscissile at about the middle; seeds nearly flat on 
both sides. 


ty 
is 
fi 
te 
rest 
y 


In dry sandy soil, Massachusetts to Georgia, Illinois, 
Kansas and Texas. April—Aug. 


Plantago elongata Pursh, to which this was referred in our first edition, differs by larger 
seeds and saccate bracts, and enters our western limits in Nebraska. 


12. Plantago heterophylla Nutt. Many- 
seeded Plantain. Fig. 3909. 


Plantago heterophylla Nutt. Trans. Am. Phil. Soe. (1I1.) 

5: 177. 1833-37. 

Annual, similar to the preceding species, but gla- 
brous or slightly puberulent; scapes ascending or 
spreading, equalling or exceeding the leaves, 2’—-10’ 
long. Leaves narrowly linear or filiform, the larger 
about 2” wide, entire or often with several distant 
small teeth or linear lobes; spikes loose, linear, 3’—5’ 
long; sepals oblong, obtuse, scarious-margined, mostly 
shorter than the bract; corolla-lobes in the more fer- 
tile plants becoming erect over the pyxis; stamens 2; 
pyxis oblong, subacute, about twice as long as the 
calyx, 7-30-seeded, circumscissile rather below the 
middle; seeds somewhat angled, scarcely concave on 
the face. 


In moist soil, New Jersey to Florida, Illinois, Arkan- 
sas, Texas and apparently introduced in California. 
April—July. 


13. Plantago arenaria W. & K Sand Plantain. 
Fig. 3910. 


Plantago arenaria W. & K. PI. Rar. Hung. 1: 51. pl. 51. 
1802. 


Annual, pubescent, somewhat viscid; stem simple, or 
commonly becoming much branched, leafy, 3-15’ high. 
Leaves opposite, or whorled, narrowly linear, entire, 
sessile, 1-3’ long, about 1” wide; peduncles axillary, 
often umbellate at the ends of the stem and branches, 
slender, as long as the leaves or longer; heads of flowers 
conic, oval, or subglobose, 5’—-10” long, about 5” thick; 
lower bracts acute or acuminate; calyx-lobes unequal; 
corolla-lobes ovate to lanceolate, acute; capsule 2-seeded. 


Fields, Dayton, Ohio, and Lancaster County, Pennsylva- 
nia. Adventive from central Europe. Summer. 


250 RUBIACEAE. Voi. IIT. 


2. LITTORELLA L. Mant. 2: 295. 1771. 


A low perennial succulent herb, with linear entire basal leaves and monoecious flowers, 
the staminate solitary or two together at the summits of slender scapes, the pistillate sessile 
among the leaves. Sepals 4 Corolla of the staminate flowers with a somewhat urceolate 
tube, and a spreading 4-lobed limb. Corolla of the pistillate flowers urn-shaped, 3-4-toothed. 
Staminate flowers with 4 long-exserted stamens, their filaments filiform, the anthers ovate. 
Pistillate flowers with a single ovary and a long-exserted filiform style. Fruit an indehiscent 
1-seeded nutlet. [Latin, shore. ] 

Two known species, the following typical, the other in southern South America. 


1. Littorella unifldra (L.) Ascherson. Plan- 
tain Shore-weed. Shore-grass. Fig. 3911. 
Plantago uniflora L. Sp. Pl. 115. 1753. 
Littorella lacustris L. Mant. 2: 295. 1771. 
Littorella uniflora Ascherson, Fl. Brand. 544. 1864. 
Tufted, usually growing in mats; leaves bright 
green, 1-3’ long, ?’-1”” wide, spreading or ascending, 
mostly longer than the scapes of the staminate flow- 
ers, which bear a small bract at about the middle; 
sepals lanceolate, mostly obtuse, with a dark green 
midrib and lighter margins, sometimes only 3 in the 
fertile flowers; stamens conspicuous, 4’-6” long; 
corolla-lobes ovate, subacute; pistillate flowers very 
small; nutlet about 1” long. 
Borders of lakes and ponds, Maine and Vermont to 
Minnesota, Newfoundland, Nova Scotia and Ontario. 
July—Aug. 


Family 36. RUBIACEAE B. Juss. Hort. 
Trian. 1759. 
MADDER FAMILY. 

Herbs, shrubs, or trees, with simple, opposite or sometimes verticillate, mostly 
stipulate leaves, and perfect, often dimorphous or trimorphous, regular and nearly 
symmetrical flowers. Calyx-tube adnate to the ovary, its limb various Corolla 
gamopetalous, funnelform, club-shaped, campanulate, or rotate, 4-5-lobed, often 
pubescent within. Stamens as many as the lobes of the corolla and alternate with 
them, inserted on its tube or throat; anthers mostly linear-oblong. Ovary I-I0- 
celled ; style short or elongated, simple or lobed; ovules I-~ in each cavity. Fruit 
a capsule, berry, or drupe. Seeds various; seed-coat membranous or crustaceous ; 
endosperm fleshy or horny (wanting in some exotic genera) ; cotyledons ovate, 
cordate, or foliaceous. 


About 340 genera and about 6000 species, of very wide geographic distribution, most abundant 
in tropical regions. Known as Madderworts. 
* Leaves opposite, stipulate (sometimes verticillate in No. 3). 
Ovules numerous in each cavity of the ovary ; herbs. 
Top of the capsule free from the calyx; seeds few, peltate. 1. Houstonia. 
Capsule wholly adnate to the calyx ; seeds minute, angular. 2. Oldenlandia. 
Ovules 1 in each cavity of the ovary. 
Shrubs or small trees; flowers in dense globular heads. 
Low evergreen herbs; flowers 2 together, their ovaries united. 
Herbs; flowers axillary, nearly sessile, distinct. 
Capsule separating into 2 carpels, one dehiscent, the other indehiscent. . Spermacoce. ~ 
Capsule separating into 2 (or 3) indehiscent carpels. Diodia. 
** Leaves appearing verticillate; herbs (some of the leaves rarely opposite in No. 7); 
ovule 1 in each cavity of the ovary. 


. Cephalanthus. 
. Mitchella. 


nur Ww 


Corolla rotate; calyx teeth minute or none. 7. Galium. 
Corolla funnelform. 
Flowers in involucrate heads. 8. Sherardia. 
Flowers in panicles. a Asberula. 


1. HOUSTONIA L. Sp. Pl. 105. 1753. 


Erect or diffuse, usually tufted herbs, with opposite entire often ciliate leaves, and small 
blue purple or white, mostly dimorphous flowers. Calyx-tube globose or ovoid, 4-lobed, the 
lobes distant. Corolla funnelform or salverform, 4-lobed, the lobes valvate, the throat glabrous 
or pubescent. Stamens 4, inserted on the tube or throat of the corolla; anthers linear or 
oblong. Ovary 2-celled. Style slender; ovules numerous in each cavity; stigmas 2, linear. 
Capsule partly inferior, globose-didymous, or emarginate at the apex, loculicidally dehiscent 
above, its summit free from the calyx. Seeds few or several in each cavity, peltate, more 


GENUS I. MADDER FAMILY. 251 


or less concave, not angled. Seed-coat reticulate or roughened; endosperm horny; embryo 
club-shaped. [Named in honor of Dr. William Houston, botanist and collector in South 
America, died 1733.] 

About 25 species, natives of North America and Mexico. Type species: Houstonia coerulea L. 


* Plants 1-7’ high; peduncles 1-flowered. 
7 Peduncles filiform, 12%’ long. 


Erect ; leaves obovate or spatulate, narrowed into petioles. 1. H. coerulea. 
Diffuse or spreading; leaves nearly orbicular. . A. serpyllifolia. 
77 Peduncles 3”-18” long, stouter. 

Calyx-lobes narrow, about equalling the capsule. 3. H. patens. 
Calyx-lobes broad, much exceeding the capsule. 4. H. minima. 
** Plants 4’-18’ high; flowers cymose. 
Calyx-lobes lanceolate-subulate, 2 to 3 times as long as the capsule. 5. H. lanceolata. 
Calyx-lobes linear-subulate, scarcely longer than the capsule. 
Leaves broad, ovate, or ovate-lanceolate. 6. H. purpurea. 
Leaves oblong or spatulate, ciliate. 7. H. ciliolata. 
Leaves linear-lanceolate or oblanceolate, not ciliate. 8. H. longifolia. 
Leaves filiform or narrowly linear. 
Flowers loosely cymose on filiform pedicels; leaves not fascicled. 9. H. tenutfolia. 
Flowers densely cymose on very short pedicels ; leaves usually fascicled. 10. H. angustifolia. 


1. Houstonia coertlea L. Bluets. 
Innocence. Eyebright. Fig. 3912. 


Houstonia coerulea L. Sp. Pl. 105. 1753. 
Hedyotis coerulea Hook. Fl. Bor. Am. 1: 286. 


1833. 

@izentindia coerulea A. Gray, Man. Ed. 2, 

Tine LOS 0: : 

Erect, 3-7’ high, glabrous, or nearly so, 
perennial by slender rootstocks and form- 
ing dense tufts. Lower and basal leaves 
spatulate or oblanceolate, about 6” long, 
sometimes hirsute or ciliate, narrowed into 
a petiole, the upper oblong, sessile; flowers 
solitary on filiform terminal and axillary 
peduncles; corolla salverform, violet, blue, 
or white with a yellow center, 4’-6” broad, 
its tube slender and about the length of the 
lobes or longer; capsule didymous, com- 
pressed, about 2” broad and broader than 
long, the upper half free from the calyx 
and shorter than its lobes. 

In open grassy places, or on wet rocks, 
Miquelon and Nova Scotia to Quebec, New 
York, Michigan, Georgia, Alabama, Tennessee 
and Missouri. April—July, or producing a few 
flowers through the summer. Includes several 
races. Called also quaker-ladies, quaker- 


x 


y 


y 7 
\ | 


EA bonnets, Venus’-pride. Bright eyes. Angel- 
: V7 i= eyes. Blue-eyed-grass or -babies. Wild forget- 
ISVIA — me-not. Nuns. Star-of-Bethlehem. Little 

j y, washerwoman. 


2. Houstonia serpyllifolia Michx. 
Thyme-leaved Bluets. 


Fig. 3913. 
Houstonia serpyllifolia Michx. Fl. Bor. Am. 
E/$s. ) 1502) 
Hedyotis serpyllifolia T. & G. Fl. N. A. 2: 39. 
r841. 


Perennial; stems prostrate or diffuse, 
slender, glabrous, 4-10’ long. Leaves or- 
bicular or broadly oval, abruptly petioled, 
3-4” long, sometimes hispidulous; or those 
of the flowering stems narrower, distant; 
flowers on terminal and axillary filiform 
peduncles; corolla usually deep blue, 4-6” 
broad, its tube rather shorter than the 
lobes; capsule similar to that of the pre- 
ceding species but usually slightly larger, 
nearly as long as the calyx. 

Somerset County, Pennsylvania, to the high 
mountains of Virginia and West Virginia, 
South Carolina, Georgia and east Tennessee. 
May. 


252 RUBIACEAE. ~ Vor. III. 


3. Houstonia patens Ell. Small Bluets. Fig. 3914. 
Houstonia Linnaei var. minor Michx. FI. Bor. 

AT Isao S035 
H. patens Ell. Bot. S. C. & Ga. 1: 191. 1821. 
Houstonia minor Britton, Mem. Torr. Club 5: 

302. 1894. 

Annual, glabrous or nearly so, branched 
from the base, 1-6’ high. Lower and basal 
leaves oval or ovate, 5’-6” long, narrowed 
into petioles often of their own length, the 
upper narrower and sessile; peduncles ax- 
illary and terminal, erect-divergent, 3-18” 
long, 1-flowered; corolla violet-blue or pur- 
ple, 3-4” broad, its tube about equalling 
the lobes or somewhat longer; capsule com- 

- pressed, didymous, 23’-3” broad, its upper 
part free from the calyx and about equal- 
ling or exceeding the subulate lobes. 

In dry soil, Virginia to Florida, Illinois, 


Arkansas and Texas. March-April. Star- 
violet (Texas). 


4. Houstonia minima Beck. Least Bluets. 
Fig. 3915. 
Houstonia minima Beck, Am. Journ. Sci. 10: 262. 1826., 
Hedyotis minima T. & G. Fl. N. A. 2: 38. 1841. 
Oldenlandia minima A. Gray, Man. Ed. 2, 173. 1856. 

Annual, spreading or diffuse, 1-22’ high, roughish. Lower 
and basal leaves oval or ovate, with petioles shorter than or 
equalling the blade, the upper oblong, sessile; peduncles axil- 
lary and terminal, rather stout, 3-12” long; flowers 4’-5” 
broad; corolla violet or purple, the tube about as long as the 
lobes; capsule didymous, compressed, about 3” broad, its upper 
part free from the calyx and considerably exceeded by the 
lanceolate foliaceous lobes. 


In dry soil, Illinois to Kansas, Arkansas, Tennessee and Texas. 
March-April. 


5. Houstonia lanceolata (Poir.) Britton. Calycose 
Houstonia. Fig. 3916. 


Hedyotis lanceolata Poir. in Lam. Encycl. Suppl. 3: 14. 1813. 
Houstonia purpurea calycosa A. Gray, Syn. Fl. 17: 26. 1878. 
Houstonia lanceolata Britton, Man. 861. tgot. 


Houstonia calycosa Mohr, Contr. Nat. Herb. 6: 739. 1901. 


Perennial, glabrous or pubescent, rather stout, 6-16’ high. 
Leaves lanceolate to linear-lanceolate, firm in texture, not cil- 
iate, sessile, or the lowest spatulate and narrowed into petioles; 
corolla-tube little exceeding the calyx; calyx-lobes lanceolate 
or lanceolate-subulate, 5” long or less, much exceeding the 
capsule; capsule ovoid-globose, about 2” thick. 


In dry soil, Illinois to North Carolina, Alabama, Tennessee and 
Oklahoma. Recorded from Maine. May-—June. 


GENUS I. MADDER FAMILY. 253 


6. Houstonia purpurea L. Large Houstonia. 
Venus’-pride. Fig. 3917. 


Houstonia purpurea L. Sp. Pl. 105. 1753. 

Hedyotis purpurea T. & G. Fl. N. A. 2: 40. 1841. 
Oldenlandia purpurea A. Gray, Man. Ed. 2, 173. 1856. 
Houstonia purpurea pubescens Britton, Mem. Torr. Club 4: 


125. 1894. 

Perennial, stout, erect, tufted, branched or simple, 
glabrous or pubescent, 4-18 high. Leaves ovate or 
ovate-lanceolate, sessile, or the lower ones. short- 
petioled, 3-5-nerved and pinnately veined, obtuse or 
acute, 2’-2’ long, 3’—15” wide, the margins often ciliate; 
flowers in terminal cymose elusters; pedicels 1-4” 
long; corolla purple or lilac, funnelform, 3’—4” long, 
the tube at least twice as long as the lobes; capsule 
compressed-globose, 13” broad, somewhat didymous, its 
upper half free and considerably shorter than the subu- 
late-linear calyx-lobes. 


In open places, Maryland to Iowa, Kentucky, Missouri, 
Georgia, Alabama and Mississippi, especially in the moun- 
tains. May-—Sept. 


oY 


7- Houstonia ciliolata Torr. Fringed Hous- 
tonia. Fig. 3918. 


Houstonia ciliolata Torr. Fl. N. U.S. 1: 173. 1824. 
Houstonia purpurea var. ciliolata A. Gray, Man. Ed. 5, 

212. 18672 

Perennial, tufted, erect or ascending, 4’-7’ high. 
Lower and basal leaves petioled, thick, 1-nerved, obo- 
vate or oblanceolate, obtuse, 6’—10” long, their margins 
conspicuously ciliate; stem leaves oblong or oblanceo- 
late, sessile or nearly so; flowers in corymbed cymes; 
pedicels filiform, 1-4” long; corolla funnelform, lilac 
or pale purple, about 3” long, the lobes about one- 
third the length of the tube; capsule little compressed, 
obscurely didymous, 13” wide, overtopped by the linear- 
lanceolate calyx-lobes. 

On rocks and shores, Ontario to Minnesota, south to 
Pennsylvania, West Virginia, Kentucky and Arkansas. 
May-Aug. 


Y 
sy 


V, 


W223 


8. Houstonia longifolia Gaertn. Long- 

leaved Houstonia. Fig. 3910. 

Houstonia longifolia Gaertn. Fruct. 1: 226. pl. 49. f. 8. : \ Vy SS, 
88 


1788. 
Houstonia purpurea var. longifolia A. Gray, Man. Ed. NS Ww 
Sere. Loos, = ay 
WY : 
Perennial, usually tufted, erect, glabrous or pu- + 
berulent, 5-10’ high. Basal leaves spatulate or ob- 
lanceolate, obtuse, not ciliate, very short-petioled; \ 


stem leaves linear or linear-oblong, acute or obtuse, 
I-nerved, 6-12” long, 1-23” wide; flowers in \ 
corymbed cymes; calyx-lobes subulate; corolla pale 
purple or nearly white, 23”—3” long, its lobes about NN 7 
one-third the length of the tube; capsule little com- 
pressed, globose, ovoid, about 1” in diameter, its upper 
half free and much exceeded by the subulate calyx- 
lobes. 

In dry open places, Maine and Ontario to Manitoba, 


eres Georgia, Mississippi and Missouri. May-— d 
ept. ; HN 


254 RUBIACEAE. Vou. III. 


g. Houstonia tenuifolia Nutt. Slender-leaved 
Houstonia. Fig. 3920. 


Houstonia tenuifolia Nutt. Gen. 1: 95. 1818. 
Houstonia purpurea var. tenuifolia A. Gray, Syn. Flor. 1: 

Parte 2o. 18755 

Perennial, somewhat tufted, very slender and widely 
branching, erect, glabrous, 6’-1° high, sometimes finely 
pubescent below. Basal and lowest stem leaves ovate 
or oval, obtuse, petioled, 4’-6” long; upper leaves nar- 
rowly linear or filiform, blunt-pointed, 6-15” long, 
2’’-12”" wide; flowers in loose corymbose cymes; pedi- 
cels filiform, 2’-6” long; corolla purple, narrow, 2”-3” 
long, its lobes short; capsule compressed-globose, didy- 
mous, about 1” in diameter, its upper half free and only 
slightly exceeded by the subulate calyx-lobes. 

In dry soil, Pennsylvania to Ohio, North Carolina and 
Tennessee. May-—July. 


to. Houstonia angustifolia Michx. Narrow- aN 


leaved Houstonia. Fig. 3921. 4A \ 


Houstonia angustifolia Michx. Fl. Bor. Am. 1: 85. 1803. 
Oldenlandia angustifolia A. Gray, Pl. Wright. 2: 68. 1853. 


Perennial by a deep root, erect, stiff, glabrous, usually 
branched, 1°-2° high. Leaves linear, 6’—18” long, 1-3” 
wide, or the lowest narrowly spatulate, usually with 
numerous smaller ones fascicled in the axils, or on 
short axillary branches; flowers in terminal dense cy- 
mose clusters; pedicels short; corolla white or pur- 
plish, between funnelform and salverform, about 2” 
long, its lobes shorter than the tube; capsule com- 
pressed-obovoid, 13” wide, its summit free and scarcely 
exceeded by the calyx-lobes. 

In dry open places, Illinois to Kansas, Texas, Tennessee 
and Florida. May-July. Star-violet. Venus’-pride. 


2. ‘OLDENLANDIA: [Plumier] Ie. Sp; Fl. 110, ase 
Erect or diffuse slender herbs, with opposite leaves, and small axillary or terminal soli- 
tary or clustered, white or pink flowers. Calyx-tube obovoid or subglobose, the limb 4-toothed. 
Corolla rotate or salverform, 4-lobed. Stamens 4, inserted on the throat of the corolla; 
anthers oblong. Ovary 2-celled; ovules numerous in each cavity; style slender, 2-lobed. 
Capsule small, ovoid, top-shaped, or hemispheric, wholly adnate to the calyx-tube, loculicidally 
dehiscent at the summit, several or many-seeded. Seeds angular, not peltate; endosperm 

fleshy; embryo club-shaped. [Named for H. B. Oldenland, a Danish botanist. ] 


About 175 species, mostly of tropical distribution, most abundant in Asia. Besides the follow- 
ing, 2 others occur in the Southern States and 1 in New Mexico. Type species: Oldenlandia 


corymbosa L. 


1. Oldenlandia uniflora L. Clustered Bluets. 
Fig. 3922. 


Oldenlandia uniflora L. Sp. Pl. 119. 1753. 
Oldenlandia glomerata Michx. Fl. Bor. Am. 1: 83. 1803. 


Annual, weak, usually tufted, more or less hirsute- 
pubescent, diffuse or ascending; stems 1-15’ long. Leaves 
short-petioled or sessile, mostly thin, entire, 3-5-nerved, 
ovate, oblong, or oval, acute at the apex, narrowed at the 
base, 3-1’ long; flowers sessile or nearly so, white, about 
1” broad, terminal and axillary, clustered or solitary; 
calyx hirsute, hemispheric in fruit, the ovate or oval lobes 
erect and nearly equalling the tube. 


In low grounds, southern New York to Florida and Texas, 
north to Arkansas and reported from Missouri. Also in Cuba 
and Jamaica. June—Sept. 


GENUS 3. MADDER FAMILY. 255 


3. CEPHALANTHUS L. Sp. Pl. 95. 1753. 


Shrubs, or small trees, with opposite or verticillate short-petioled entire leaves, and 
terminal or axillary, densely capitate, bracteolate small white or yellow flowers. Calyx-tube 
obpyramidal, its limb with 4 obtuse lobes. Corolla tubular-funnelform, with 4 short erect 
or spreading lobes. Stamens 4, inserted on the throat of the corolla; filaments very short; 
anthers oblong, 2-cuspidate at the base. Ovary 2-celled; ovules solitary in each cavity, pen- 
dulous; style filiform, exserted; stigma capitate. Fruit dry, obpyramidal, 1-2-seeded. Endo- 
sperm cartilaginous; cotyledons linear-oblong. [Greek, head-flower. ] 


About 6 species, natives of America and Asia. The following, here taken as typical, is the only 
one known to occur in North America, unless the southwestern and Mexican plant proves to be 
distinct. 


1. Cephalanthus occidentalis L. Button- 
bush. Button-tree. Honey-balls. 
Globe-flower. Fig. 3923. 

Cephalanthus occidentalis L. Sp. Pl. 95. 1753. 

A shrub 3°-12° high, or sometimes a tree, 
up to 20° high, with opposite or verticillate 
leaves and branches, glabrous, or somewhat 
pubescent. Leaves petioled, ovate or oval, en- 
tire, acuminate or acute at the apex, rounded 
or narrowed at the base, 3’-6’ long, 1’—23’ wide; 
peduncles 1’~3’ long; heads globose, about 1’ 
in diameter, the receptacle pubescent; flowers 
sessile, white, 4”-6” long; style very slender, 
about twice the length of the corolla; calyx- 
tube prolonged beyond the ovary. 


In swamps, and low grounds, New Brunswick 
to western Ontario and Wisconsin, Florida, Texas, 
Arizona and California. Pond dogwood. Button- 
wood shrub. Box. Pin-ball. Little snowball. 
Button- or crane willow. Swamp-wood. River- 
or crouper-bush. June--Sept. 


4. MITCHELLA L. Sp. Pl. rit. 1753. 


Creeping herbs, with opposite petioled, entire or undulate, evergreen leaves, and white 
axillary or terminal peduncled geminate dimorphous flowers, their ovaries united. Calyx- 
tube ovoid, the limb 3-6-lobed (usually 4-lobed). Corolla funnelform, usually 4-lobed, the 
lobes recurved, bearded on the inner side. Stamens as many as the lobes of the corolla and 
inserted on its throat; filaments short and style exserted, or filaments exserted and style short. 
Ovary 4-celled; stigmas 4, short, filiform; ovules I in each cavity, erect, anatropous. Fruit 
composed of 2 united drupes usually containing 8 roundish nutlets. Seed erect; cotyledons 
short, obtuse; embryo minute. [Named after Dr. John Mitchell, botanist and correspondent 
of Linnaeus in Virginia. ] 

Two species, the following typical one North American, the other Japanese. 


1. Mitchella répens L. Partridge-berry. Twin-berry. Fig. 3924. 
Mitchella repens L. Sp. Pl. 111. 1753. 


Stems slender, trailing, rooting at the 
nodes, 6-12’ long, branching, glabrous, or 
very slightly pubescent. Leaves ovate-or- 
bicular, petioled, obtuse at the apex, round- 
ed or somewhat cordate at the base, 3-10” 
long, pinnately veined, dark green, shining; 
peduncles shorter than the leaves, bearing 
2 sessile white flowers at the summit; co- 
rolla 5’-6” long; drupes red (rarely white), 
broader than high, 2’—4” in diameter, per- 
sistent through the winter, edible. 


In woods, Nova Scotia to Florida, west to 
western Ontario, Minnesota, Arkansas and 
Texas. April-June, sometimes flowering a 
second time in the autumn. Hive- or squaw- 
vine. Checker-berry. Deer-berry. Fox- or box- 
berry. Partridge-vine. Winter-clover. Chicken-, 
cow-, pigeon-, snake- or tea-berry. Two-eyed- 
or one-berry. Squaw-plum. Leaves often whit- 
ish-veined ; flower-buds pink. Ascends to 5000 
ft. in Virginia. 


256 RUBIACEAE. Vor. III, 


5. SPERMACOCE L. Sp. Pl. 102. 1753. 


Herbs, with 4-sided stems, opposite pinnately veined stipulate leaves, and small white 
flowers, in dense axillary and terminal clusters. Calyx-tube obovoid or obconic, its limb 
4-toothed. Corolla funnelform, 4-lobed. Stamens 4, inserted on the tube of the corolla; 
anthers oblong or linear. Ovary 2-celled; ovules 1 in each cavity; style slender; stigma 
capitate, or slightly 2-lobed. Capsule coriaceous, didymous, of 2 carpels, one dehiscent, the 
other indehiscent. Seeds oblong, convex on the back; endosperm horny; embryo central; 
cotyledons foliaceous. [Greek, seed-point, from the sharp calyx-teeth surmounting the carpels.] 


Two or three species, natives of America. Type species: 
Spermacoce tenuior L. 


1. Spermacoce glabra Michx. Smooth Button- 
weed. Fig. 3925. 
Spermacoce glabra Michx. Fl. Bor. Am. 1: 82. 1803. 


Glabrous, decumbent or ascending, rather stout; stems 
10-20’ long. Leaves lanceolate or elliptic-lanceolate, peti- 
oled, or the uppermost sessile, 1-3’ long, 4-12” wide, acute 
at each end, the margins rough; corolla pubescent in the 
throat, about 12” long, scarcely exceeding the ovate-lanceo- 
late acute calyx teeth; stamens and style included; capsule 
obovoid, about 2” long, glabrous; seeds black, punctate. 


On _river-banks and in wet soil, southern Ohio and Ken- 
tucky to Florida, Kansas and Texas. June—Sept. 


ously stipulate leaves, and small axillary white lilac or purple flowers. Calyx-tube obconic 
or obovoid, the limb 2-4-lobed (sometimes 1-6-lobed), often with minute teeth between the 
lobes. Corolla funnelform or salverform, mostly 4-lobed. Stamens usually 4, inserted on 
the throat of the corolla; filaments slender; anthers versatile, oblong-linear, exserted. Ovary 
2-celled (rarely 3-4-celled) ; ovules 1 in each cavity; style filiform, simple or 2-cleft; stigmas 
2. Fruit crustaceous or somewhat fleshy, oblong, obovoid, or subglobose, 2-celled, finally 
separating into 2 indehiscent carpels. Seeds oblong, convex on the back; endosperm horny ; 
cotylendons foliaceous; embryo straight. [Greek, thoroughfare, where the species are fre- 
quently found. ] 


About 35 species, mostly American. Besides the following, another occurs in the Southern 
States. Type species: Diodia virginiana L 


Leaves linear-lanceolate ; style entire; stigmas capitate. * 1. D. teres. | 
Leaves lanceolate or oval; style 2-cleft; stigmas filiform. 2. D. virginiana. 
: . DS T \ y \ / 
1. Diodia téres Walt. Rough Button-weed. WZ \\ WA 


Fig. 3926. \ WA 
ee 


Diodia teres Walt. Fl. Car. 87. 1788. 
Spermacoce diodina Michx. Fl. Bor. Am. 1: 82. 1803. 


Rigid, usually rough, much branched from near the 
base, the branches prostrate or ascending, 4-sided above, 
4’-30’ long. Leaves linear or linear-lanceolate, very 
rough, 4’-12’ long, 14’"-3” wide, acute, the margins revo- 
lute when dry; flowers lilac or purple, 2”-3” long, usually 
solitary in the axils; style entire; stigmas capitate; fruit 
obovoid or top-shaped, hispid, about 2” high, the usually 
4 persistent calyx-lobes ovate to lanceolate. 


In dry or sandy soil, Connecticut to Florida, west to 
Illinois, Kansas, Texas, New Mexico and Sonora. July— 
Sept. Poverty-, or poor-land-weed. Poor Joe. 


GENUS 6. MADDER FAMILY. 257 


2. Diodia virginiana L. Larger Button- 
weed. Fig. 3927. 


Diodia virginiana L. Sp. Pl. 104. 1753. . 


Hispid-pubescent or glabrate, much branched 
from near the base, the branches procumbent 
or ascending, 1°-2° long. Leaves lanceolate to 
narrowly oval, narrowed at the base, acute, or 
the lowest obtuse at the apex, 17-3’ long; flow- 
ers I or 2 in each axil, about 6” long, the 
corolla-tube very slender; fruit somewhat 
fleshy, but becoming dry, hirsute or glabrous, 
oval, 3’-4” high, furrowed, crowned with the 
2 or 3 persistent lanceolate calyx-lobes. 


In moist soil, southern New Jersey to Florida, 
west to Arkansas and Texas. June—Aug. 


as GALIUM L. Spores BOSs. +1753: 


Annual or perennial herbs, with 4-angled slender stems and branches, apparently verticil- 
late leaves, and small white, green, yellow or purple flowers, mostly in axillary or terminal 
cymes or panicles, the pedicels usually jointed with the calyx. Flowers perfect, or in some 
species dioecious. Calyx-tube ovoid or globose, the limb minutely toothed, or none. Corolla 
rotate, 4-lobed (rarely 3-lobed). Stamens 4, rarely 3; filaments short; anthers exserted. 
Ovary 2-celled; ovules I in each cavity. Styles 2, short; stigmas capitate. Fruit didymous, 
dry or fleshy, smooth, tuberculate, or hispid, separating into 2 indehiscent carpels, or some- 
times only 1 of the carpels maturing. Seed convex on the back, concave on the face, or 
spherical and hollow; endosperm horny; embryo curved; cotyledons foliaceous. [Greek, 
milk, from the use of G. verum for curdling.] 


About 250 species, of wide geographic distribution. Besides the following, some 35 others 
occur in the southern and western parts of North America. The leaves are really opposite, the 
intervening members of the ventricles being stipules. Type species: Galium Molliigo L. 


I. Fruit dry. 
A. Flowers yellow. 1. G. verum. 
B. Flowers white, green or purple. 
1. Annuals. 


Flowers in axillary cymules, or panicled. 
Fruit granular or tubercled, not bristly. 
Fruit slightly granular, or smooth, 1 mm. broad; pedicels not recurved; stem very slender. 
2. G. parisiense. 
Fruit granular-tubercled, 3 mm. broad; fruiting pedicels recurved; stem stout. 
3. G. tricorne. 
Fruit densely bristly-hispid. 
Cymes few-flowered ; leaves 2-8 cm. long; fruit fully 4 mm. broad. 4. G. Aparine. 
Cymes mostly several-flowered ; leaves 1-2.5 cm. long; fruit smaller. 5. G. Vaillantit. 
Flowers solitary in the axils, subtended by 2 foliaceous bracts; fruit bristly. 6. G. virgatum. 


2. Perennials. 
* Fruit bristly hispid. 
a. Leaves in 4’s, 1-nerved. 7. G. pilosum. 
b. Leaves in 4’s, 3-nerved. 


Leaves lanceolate, oval, or ovate; flowers in open cymes. 
Upper leaves lanceolate to ovate-lanceolate, acuminate. 8. G. lanceolatum. 
Upper leaves ovate, oblong, oval, ovate-lanceolate or obovate, obtuse. 
Corolla usually hirsute; plant mostly pubescent; leaves oblong to ovate-lanceolate. 
9. G. circaezans. 


Corolla glabrous; plant little pubescent ; some leaves obovate. 10. G. kamtschaticum. 
Leaves linear to lanceolate; flowers in terminal panicles. 11. G. boreale. 
c. Leaves in 6’s. 12. G. triflorum. 


** Fruit smooth or warty. 
+ Flowers brown-purple. 


Leaves lanceolate, 3-nerved; fruit smooth. 13. G. latifolium. 
Leaves narrowly lanceolate, 1-nerved; fruit warty. 14. G. arkansanum 


17 


258 


Leaves lanceolate, acuminate. 


RUBIACEAE. Vor. III, 


77 Flowers white or greenish. 
Stems smooth; introduced species. 
Leaves linear to oblanceolate, cuspidate. 


15. G. Mollugo. 
16. G. sylvaticum. 


Stems mostly more or less retrorsely scabrous; native species. 


Leaves obtuse; stems slightly scabrous. 
Plants of wet soil, not shining. 


Flowers solitary, or few in small simple cymes. 


Corolla-lobes mostly 4, acute. 


Fruit 14%” in diameter; leaves ascending or spreading. 17. G. tinctorium. 
Fruit 4%” in diameter; leaves mostly reflexed. 18. G. labradoricum. 
Corolla-lobes mostly 3, obtuse. : 
Pedicels rough, curved; flowers mostly solitary. 19. G. trifidum. 
Pedicels smooth, straight; flowers 2 or 3 together. 20. G. Claytoni. 
Flowers numerous in forked cymes. 21. G. palustre. 
Shining plant of dry woodlands. 22. G. concinnum, 
Leaves cuspidate-acute ; stems retrorsely hispid. 23. G. asprellum. 
II. Fruit fleshy. 24. G. bermudense. 


1. Galium vérum L. Yellow Bedstraw. Lady’s Bedstraw. Fig. 3928. 


Galium verum L. Sp. Pl. 107. 1753. 


Perennial from a somewhat woody base, erect or 
ascending, simple or branched, 6’-23° high. Stems 
smooth or minutely roughened; leaves in 6’s or 8's, 
narrowly linear, 4’—-12” long, about 2” wide, rough 
on the margins, at length deflexed; flowers yellow, 
the cymes in dense narrow panicles; lower branches 
of the panicles longer than the internodes at anthe- 
sis; fruit usually glabrous, less than 1” broad. 


In waste places and fields, Maine and Ontario to Mas- 
sachusetts, southern New York, New Jersey and Penn- 
sylvania. Adventive ornaturalized from Europe. Native 
also of Asia. May—Sept. Cheese-rennet. Curdwort. Bed- 
flower. Fleawort. Maids’-hair. Yellow cleavers. Our 
Lady’s-bedstraw. 


Galium Wirtgeni F. Schultz, differs in having the 
lower branches of the panicle very short at anthesis, and 
is recorded as established in a meadow at Norfolk, 
Connecticut. 


Wy 


2. Galium parisiénse L. Wall Bedstraw. 


Fig. 3929. 


Galium parisiense L. Sp. Pl. 108. 


Galium anglicum Huds. Fl. Angl. Ed. 2, 69. 


Annual, erect or ascending, very slender, much branched ; 
stem rough on the angles, 6-12’ high. Leaves in verticils 
of about 6 (4-7), linear or linear-lanceolate, cuspidate, 
minutely scabrous on the margins and midrib, 2’-5” long; 
cymes several-flowered, axillary and terminal on filiform 
peduncles; flowers minute, greenish-white; fruit glabrous, 
finely granular, less than 3” wide. 


Along roadsides, Virginia and Tennessee. Adventive or 
naturalized from Europe. June—Aug. 


BNY Be 
\ aa 
WA WV 2s 
W YY 
Bo V W oA 


YS 


1778. 


GENUS 7. MADDER FAMILY. 259 


3. Galium tricérne Stokes. Rough-fruited Corn 
Bedstraw. Fig. 3930. 


Galium tricorne Stokes; With. Bot. Arr. Brit. Pl. Ed. 2, 1: 
mess) 1787. 


Rather stout, decumbent or ascending, 6-12’ high, 
simple, or little branched. Stem rough with reflexed 
prickles; leaves in 6’s or 8’s, linear or narrowly oblan- 
ceolate, 1’ long or less, 14’-2” wide, mucronate, rough 
on the margins and midrib; peduncles axillary, shorter 
than the leaves; pedicels thickened and curved down- 
ward in fruit; cymes axillary, usually 3- (1-3-) flow- 
ered; fruit tuberculate or granular, not hispid, 4’’-5” 
broad. 


In waste places, Ontario, and in ballast about the eastern 
seaports. May—Aug. 


4. Galium Aparine L. Cleavers. Goose-grass. 
Cleaver-wort. Fig. 3931. 


Galium Aparine L. Sp. Pl. 108. 1753. 


Annual, weak, scrambling over bushes, 2°-5° long, 
the stems retrorsely hispid on the angles. Leaves in 

. 6's or 8s, oblanceolate to linear, cuspidate at the apex, 
yx :1’-3’ long, 2’-5” wide, the margins and midrib very 
rough; flowers in 1-3-flowered cymes in the upper axils; 
peduncles 57-12” long; fruiting pedicels straight; 
fruit 2”-3” broad, densely covered with short hooked 
bristles. 


In various situations, New Brunswick to Ontario, South 
Dakota, Florida and Texas. Bermuda. Apparently in part 
naturalized from Europe. Widely distributed in temperate 
regions as a weed. May—Sept. Among some 7o other Eng- 
lish names are catchweed, beggar-lice, burhead, claver-grass, 
cling-rascal, scratch-grass, wild hedge-burs, hairif or airif, 
stick-a-back, or stickle-back, gosling-grass, gosling-weed, 
turkey-grass, pigtail, grip or grip-grass, loveman, sweet- 
hearts, scratch-weed, poor robin. 


5. Galium Vaillantii DC. Vaillant’s Goose-grass 
or Cleavers. Fig. 3932. 


Galium Vaillantii*DC. Fl. France 4: 263. 1805. 
Galium Aparine var. Vaillantii Koch, Fl. Germ. 330. 1837. 


Similar to the preceding species but smaller, the stem 
equally rough-angled. Leaves smaller, 1’ in length or 
less, linear-oblong or slightly oblanceolate, cuspidate- 
pointed, rough on the margins and midrib; cymes 2-0- 
flowered; fruit 1-14” broad, usually less hispid. 


In low grounds, Ontario to British Columbia, Missouri, 
Arizona and to California. Europe. The European G. 
spurium L., to which this plant was referred in the first 
edition, appears to have uniformly smooth fruit. 


RUBIACEAE. Vorstik 


6. Galium virgatum Nutt. Southwestern 
Bedstraw. Fig. 3933. 


Galium virgatum Nutt.; T.& G. Fl. N. A.2: 20. 1841. 


Annual, 4-12’ high, usually hispid, sometimes 
nearly glabrous; stem very slender, 4-angled, 
branched from the base, or simple. Leaves in 4’s, 
oblong or linear-oblong, 23’—5” long, 1” wide, or 
less, obtuse or acutish; peduncles axillary, 1-flow- 
ered, less than 1” long, recurved in fruit; flower 
white, subtended by 2 large oblong to lanceolate 
bracts which closely resemble the leaves; fruit 
about 1” in diameter, covered with slender barbed 
bristles. ‘ 


Dry prairies, barrens and glades, Tennessee and 
Missouri to Arkansas, Louisiana and Texas. April— 


June. 
7. Galium pilosum Ait. Hairy Bed- oS Fg 
VY Ay 
straw. Fig. 3934. r WY x n 

Galium pilosum Ait. Hort. Kew. 1: 145. 17809. at VES Si) 
Galium puncticulosum Michx. Fl. Bor. Am. 1: 80. IN p 5, 7 ayy 

1303, : Rye Shep. 
Galium pilosum puncticulosum T. & G. Fl. N. A. O WA ISR? 


PAY Me iteys ite 

Perennial, more or less hirsute-pubescent or 
glabrate; stems ascending, branched, 1°-23° 
long. Leaves in 4’s, oval or oval-ovate, punc- 
tate, I-nerved, obtuse, or obscurely 3-nerved 
at the base, mucronulate, 4-12” long, 2’-5” 
wide, the lower usually smaller; peduncles ax- 
illary and terminal; cymes numerous but few- 
flowered; pedicels 1-6” long, flowers yellow- 
ish-purple; fruit dry, densely hispid, nearly 2” 
in diameter. 


In dry or sandy soil, New Hampshire to On- 
tario, Michigan, Kansas, Florida and Texas. June— 
Aug. 


8. Galium lanceolatum Torr. Torrey’s 
Wild Liquorice. Fig. 3935. 
Galium circaezans var. lanceolatum Torr. Cat. Pl. N. 
J, 2}, | Tiss 
Galium lanceolatum Torr. Fl. U. S. 168. 1824. 
Galium Torreyi Bigel. Fl. Bost. Ed. 2, 56. 1824. 


Perennial, glabrous or nearly so, the stems 
minutely roughened, simple or often branched. 
1°-2° high. Leaves in -4’s, lanceolate or ovate- 
lanceolate, acutish or acuminate, 3-nerved, more 
or less ciliate on the margins and nerves, 1’-23’ 
long, 5-11” wide, the lower smaller and obtuse 
or obtusish; cymes rather few-flowered, loose, 
widely branched; flowers sessile or very nearly 
so; corolla glabrous, yellowish green to purple, 
its lobes acuminate; fruit dry, hispid with long 
hairs, 2”-22” broad. 

In dry woods, Quebec and Ontario to Minnesota, 


south to North Carolina and Kentucky. Ascends to 
P 4ooo ft. in Virginia. June—Aug. 


GENUS 7. MADDER FAMILY. 261 


g. Galium circaézans Michx. Wild 
Liquorice. Cross-Cleavers. 
Fig. 3936. 


G. circaezans Michx. Fl. Bor. Am. 1: 80. 1803. 


Galium circaezans glabellum Britton, Mem. Torr. 
Club 5: 303. 1894. 


Perennial, more or less pubescent, or gla- 
brate, branched, 4°-2° high. Leaves in 4’s, 
oval, oval-lanceolate or ovate, obtuse or ob- 
tusish at the apex, 3-nerved, 6-18” long, 
4-8” wide, usually somewhat, pubescent on 
both surfaces, ciliolate, the lower smaller; 
cymes divaricately branched; flowers sessile 
or nearly so, greenish; corolla hirsute with- 
out, or glabrous, its lobes acute; fruit hispid, 
similar to that of the preceding species, at 
length deflexed. 


In dry woods, Quebec and Ontario to Minne- 
sota, Florida, Kansas and Texas. May-July. 


10. Galium kamtschaticum Steller. North- 
ern Wild Liquorice. Fig. 3937. 
Galium kamtschaticum Steller; R. & S. Mant. 3: 186. 


1627. 
Cilan Littellii Oakes, Hovey’s Mag. 7: 179. 1841. 
Galium circaezans var. montanum T. & G. Fl. N. A. 

PAS Birly  aheylses 

Similar to the preceding species, but weak, 
smaller, stems 4-15’ long. Leaves in 4’s, broadly 
oval, orbicular, or obovate, thin, 3-nerved, obtuse, 
mucronulate, 6’-18” long, 4-12” wide, glabrate, 
or, pubescent with short scattered hairs on the 
upper surface and on the nerves beneath, some- 
times ciliate; flowers few, all on pedicels 2’-6” 
long; corolla glabrous, yellowish-green, its lobes 
acutish; fruit hispid, 2” broad. 

In mountainous regions, Cape Breton Island, Que- 
bec, northern New England and northern New York. 
Also in northeastern Asia. Summer. 


Z Mn XZ oSasrand 
11. Galium boreale L. Northern Bed- A sh “yon 
straw. Fig. 3938. 5S) | a= 
Galium boreale L. Sp. Pl. 108. 1753. V2 “po 20S INIZ N aoe 


Galium septentrionale R.& S. Syst. 3: 253. 1818. 


_ Erect, perennial, smooth and glabrous, strict, 

simple, or branched, leafy, 1°-23° high. Leaves 
in 4’s, lanceolate or linear, 3-nerved, obtuse 
or acute, I-23’ long, 1’—3” wide, the margins 
sometimes ciliate; panicles terminal, dense, 
many-flowered, the flowers white, panicled, in 
small compact cymes; fruit hispid, at least 
when young, sometimes becoming glabrate 
when mature, about 1” broad. 


In rocky soil or along streams, Quebec to 
Alaska, south to New Jersey, Pennsylvania, 
Michigan, Missouri, New Mexico and California. 
Also in Europe and northern Asia. May—Aug. 


262 RUBIACEAE. Vo. III. 


12. Galium trifldrum Michx. Sweet- 
scented or Fragrant Bedstraw. 


Fig. 3939. 
Galium triflorum Michx. Fl. Bor. Am. 1: 80. 1803. 


Perennial, diffuse, procumbent, or ascending, 
glabrous or nearly so, shining, fragrant in drying, 
the stems and margins of the leaves sometimes 
roughened. Leaves in 6’s, narrowly oval or 
slightly oblanceolate, I-nerved, cuspidate at the 
apex, narrowed at the base, 17-33’ long, 2-6” 
wide; peduncles slender, terminal and axillary, 
often exceeding the leaves, 3-flowered or branched 
into 3 pedicels which are 1-3-flowered; flowers 
greenish; fruit 12’-2” broad, hispid with hooked 
hairs; seed almost spherical, the groove obsolete. 

In woods, Greenland to Alaska, south to Florida, 
Louisiana, Colorado and California. Also in north- 


ern Europe, Japan and the Himalayas. June—Aug. 
Three-flowered bedstraw. 


13. Galium latifolium Michx. Purple 
Bedstraw. Fig. 3940. 


Galium latifolium Michx. Fl. Bor. Am. 1: 79. 

1803. : 

Perennial, erect, smooth and glabrous (rarely 
hispid), branched, 1°-2° high. Leaves in 4’s, 
lanceolate, 3-nerved, acute or acuminate at 
the apex, narrowed or rounded at the base, 
1’-2’ long, 4-8” wide, the midrib sometimes 
pubescent beneath, the margins minutely 
roughened, the lower smaller and sometimes 
opposite; peduncles axillary and terminal, 
slender, but usually shorter than the leaves; 
flowers purple; cymes loosely many-flowered, 
the pedicels slender, 2”-6” long, smooth or 
very nearly so; fruit smooth, slightly fleshy, 
2” broad, usually only one of the carpels de- 
veloping. 

In dry woods in mountainous regions, Penn- 
sylvania to Tennessee and Georgia. May—Aug. 


Wa 14. Galium arkansanum A. Gray. Ar- 
mt kansas Bedstraw. Fig. 3941. 


Galium arkansanum A. Gray, Proc. Am. Acad. 
19: 80. 1883. 


Similar to the preceding species but usually 
lower, the leaves linear-lanceolate, 6’—12” long, 
1-3” wide, the lateral nerves obscure or none, 
the midrib sometimes pubescent beneath, and 
the margins ciliate; fruiting pedicels roughish, 
3-12” long; flowers brown-purple, the numer- 
ous cymes loosely several-many-flowered; fruit 
glabrous, warty, somewhat fleshy, each carpel 
1-13” in diameter. 


Southern Missouri, Arkansas and Oklahoma. 
June-July. 


GENUS 7. MADDER FAMILY. 263, 


15. Galium Mollugo L. Wild Madder. White 
or Great Hedge Bedstraw. Fig. 3942. 
Galium Mollugo L. Sp. Pl. 107. 1753. 


Perennial, glabrous or nearly so throughout. Stems 
smooth, erect, or diffusely branched, 1°-3° long; leaves 
in 6’s or 8’s, oblanceolate or linear, cuspidate at the apex, 
6-15" long, 1-2” wide, sometimes roughish on the 
margins; flowers small, white, very numerous in termi- 
nal panicled cymes; pedicels filiform, divaricate; fruit 
smooth and glabrous, nearly 1” broad. 

In fields and waste places, Newfoundland to Vermont, 
Ohio, Pennsylvania, New Jersey and Virginia. Naturalized 


from Europe. Called also whip-tongue. Infants’- or babies’- 
breath. May-—Sept. 


Galium eréctum Huds., also European, differs slightly 
by having somewhat larger flowers on ascending pedicels, 
and is adventive in fields from Quebec to Connecticut and 
New York. 


16. Galium sylvaticum L. Wood Bedstraw. 
Fig. 3943. 


G. sylvaticum L. Sp. Pl. Ed. 2, 155. 1762. 


Perennial, erect, 2°-3° tall; stems several or many, 
shining, obtusely 4-angled, glabrous, or slightly pu- 
bescent, not scabrous. Leaves lanceolate or oblong- 
lanceolate, pale beneath, whorled in 8’s or 6’s, or 
those of the branches in 4’s, or opposite, the larger 
sometimes 2’ long; panicles large; pedicels filiform, 
erect-spreading in fruit; flowers white; corolla-lobes 
apiculate; fruit smooth. 


Fields and thickets,.Maine and Vermont, escaped from 
cultivation. Native of Europe. June—July. 


17. Galium tinctorium L. Stiff Marsh Bed-straw. Wild Madder. Fig. 3944. 


Galium tinctorium L. Sp. Pl. 106. 1753. ‘ é ~\ 
Galium trifidum var. latifolium Torr. Fl. N. & Mid. 4 
States, 78. 1826. Z 


Galium tinctorium filifolium Wiegand, Bull. Torr. Club la a oD 
24: 397. 1897. Y 
Perennial; stem erect, 6-15’ high, rather stiff, y 

branched almost to the base, the branches com- / 

monly solitary, strict (not irregularly diffuse), sev- ~| iy 


eral times. forked; stem 4-angled, nearly glabrous; 
leaves commonly in 4’s, linear to lanceolate, 4’—1' 
long, mostly broadest below the middle, obtuse, 
cuneate at the base, dark green and dull, not papil- 
lose, I-nerved, the margins and midrib roughish; 
flowers terminal in clusters of 2 or 3; pedicels slen- 
der, not much divaricate in fruit; corolla white, 
large, 1-1?” broad, 4-parted, its lobes oblong, acute; 
disk large; fruit smooth; seed spherical, hollow. 
annular in cross-section. 


Damp shady places, wet meadows and swamps, Quebec to North Carolina, Florida, Tennessee, 
Michigan, Nebraska and Arizona. May-July. 


RUBIACEAE. 


Vou. IIT. 


Sy ° Ag = 
aes 18. Galium labradéricum Wiegand. Labrador 
“\ 9 Marsh Bedstraw. Fig. 3945. 
< Galium tinctorium labradoricum Wiegand, Bull. Torr. Club 
24: 398. 1897. 
fl) 
ix Se Galium labradoricum Wiegand, Rhodora 6: 21. 1904. 
es Ga 
SK ARS oS Perennial, with very slender rootstocks; stems weak, 
BS as smooth, slender, more or less branched, 2-12’ high. 
| — y, Y pay - 
Ne ANS \p y, Leaves #—%’ long, linear-oblanceolate, narrowed at the 
HN | “OSS base, becoming reflexed, scabrous on the margins and 
——/ (y oS midvein beneath; flowers solitary, about 1” broad, or in 
O NV Y/, simple cymes; corolla white, mostly 4-parted; fruit 
OU 4 = smooth, seed annular in cross-section. 
DO 5 Swat 
yy &- 
3. Oo OO In mossy bogs, Newfoundland to Wisconsin, Connecticut, 
4 os IN western Massachusetts and New York. June—Aug. 
1g. Galium trifidum L. Small Bedstraw. Small Cleavers. Fig. 3946. 


Galium trifidum L. Sp. Pl. 105. 1753. 


G. trifidum var. pusillum A. Gray, Man. Ed. 5, 209. 1867. 


Perennial by slender rootstocks, very slender and weak; 
stem ascending, 16’ long or less, much branched and inter- 
tangled; stem sharply 4-angled, rough; branches com- 
monly in 2’s; leaves in 4’s, linear-spatulate, 23-7” long, 
obtuse, cuneate at the base, I-nerved, dark green and dull 
on both surfaces, scarcely papillose, the margins and mid- 
rib retrorse-scabrous; flowers small, on lateral or termi- 
nal pedicels which are capillary and much longer than the 
leaves, commonly two at each node or three terminal; 
corolla very small, white, ?’” long, trifid, its lobes broadly 
oval, very obtuse; fruit glabrous; seed spherical and hol- 
low, annular in cross-section 


7, 


\\\ : PYF 


aa “SOF 
| VY i 
< 
G9), iN \ 
y, 
va 


Sphagnous bogs and cold swamps, Newfoundland to New 
York, British Columbia, Ohio, Nebraska and Colorado. Europe 
and Asia. Summer. 


20. Galium Claytoni Michx. 


Perennial ; 


Clayton’s Bedstraw. Fig. 3947. 
Galium Claytoni Michx. Fl. Bor. Am. 1: 78. 
Galium tinctorium Bigelow, Fl. Bost. Ed. 2, 54. 


1803. 
1824. 


stem erect or ascending, more diffuse 


2 


€ 


when old, 62° high; stem slender or sometimes quite 
stout, sharply 4-angled, more or less rough, the diffuse 
branches in 2’s; leaves of medium size, 4’-8” long, 
commonly in 5’s or 6’s, linear-spatulate or spatulate- 
oblong, obtuse, cuneately narrowed into a short petiole, 
rather firm in texture, scabrous on the margin and 
midrib, dark green and dull above, not papillose, dis- 
colored in drying; flowers in clusters of 2’s or 3’s, term- 
inal, provided with I or 2 minute bracts; pedicels 
straight, in fruit strongly divaricate, glabrous and rather 
stout; corolla minute, white, 3-parted, the lobes broadly 
oval, obtuse; fruit glabrous; seed spherical and hollow, 
annular in cross-section. 


Swamps, Quebec to New York, North Carolina, Michigan, Missouri, Nebraska and Texas. 


May-July. 


GENUS 7. MADDER FAMILY. 265 


21. Galium palistre L. Marsh Bedstraw. Fig. 3948. 


Galium palustre L. Sp. Pl. 105. 1753. ey, & 3 
Galium trifidum var. bifolium Macoun, Cat. Can. x ie ( re 
Plants 202. 1884? ; WZ JE 


Perennial, stem erect and rather slender, about 
16’ high; internodes very long (middle one 23’-3’ 
long) ; short branches mostly in 2’s. Stem sharply 
4-angled, glabrous or a little rough; leaves in 
typical specimens rather small, in 2’s to 6’s, linear- 
elliptic to spatulate, cuneate at the base, obtuse, 
3-8” long, 1-2” wide, the rather firm margins 
and the midrib slightly scabrous, not papillose; 
flowers numerous in terminal and lateral cymes; 
bracteoles in the inflorescence minute; pedicels in 
flower ascending, 12’-23” long, in fruit strongly 
divaricate; corolla large, white, 1-1?” broad, 
4-parted, the lobes oblong, acute; disk almost 
obsolete; fruit glabrous; endosperm of the seed 
grooved on the inner face, in cross-section lunate. 


In damp shady or open places along roadsides and 
ditches, or in the margins of swamps. Newfound- 
land, Prince Edward Island and Quebec, to Con- 
necticut and New York. Also in Europe. Summer. 


+ (08 


22. Galium concinnum Torr. & Gray. 
Shining Bedstraw. Fig. 3949. 


Galium parviflorum Raf. Med. Rep. (II.) 5: 360. 
1808? 


Galium concinnum T. & G. FI. N. A. 2: 23. 1841. 


Perennial, glabrous, shining, usually much 
branched, the angles of the stem and edges of 
the leaves minutely scabrous. Leaves usually 
all in 6’s, linear or sometimes broader above 
the middle, narrowed at the base, blunt-pointed 
or minutely cuspidate, 4’—6” long, 1-13” wide, 
green in drying; peduncles filiform; pedicels 
short; flowers minute, white, numerous in open 
cymes ; fruit small, glabrous; endosperm deeply 
grooved. 

In dry woodlands, western New Jersey to Vir- 
ginia, west to Minnesota, Kansas and Arkansas. 
June—Aug. 


23. Galium aspréllum Michx. Rough 
Bedstraw. Fig. 3950. 

Galium asprellum Michx. Fl. Bor. Am. 1: 78. 1803. 

Perennial, weak, much branched and reclining 
on bushes, or sometimes erect; stem retrorsely 
hispid, 2°-6° long. Leaves in 6’s or 5’s, or those 4 
of the branches rarely in 4’s, narrowly oval or 
slightly oblanceolate, cuspidate at the apex, nar- 
rowed at the base, sometimes so much so as to 
appear petioled, 4’-8” long, 17-2” wide, their 
margins and midribs rough; cymes terminal and 
axillary, several-many-flowered; flowers white; 
fruit smooth and glabrous, about 1” broad; endo- 
sperm with a shallow groove. 


In moist soil, Newfoundland to western Ontario, 
south to North Carolina, Illinois, Wisconsin and 
Nebraska. Called also pointed cleavers. Ascends 
to 3500 ft. in the Adirondacks. June—Aug. 


266 RUBIACEAE. Vo. IIT, 


24. Galium bermudénse L. Ccast Bed- 
straw. Fig. 3951. 


Galium bermudense L. Sp. Pl. 105. 1753. 
alt hispidulum Michx. Fl. Bor. Am. 1: 79. 

18503. 

Relbuniuen bermudense Britten, Journ. Bot. 47: 

42. 1909. 

Perennial, much branched, hirsute, hispid 
or nearly glabrous, 1°-2° high. Leaves in 4’s, 
I-nerved, oval, mucronate, rather thick, 3’—10” 
long, 13’—4” wide, the margins more or less 
revolute in drying; flowers few, terminating 
the branchlets, white; pedicels 3’-4’ long, 
rather stout, becoming deflexed in fruit, some- 
times I-bracteolate; fruit fleshy, purplish, mi- 
nutely pubescent, becoming glabrate, about 2” 
broad. 


In dry or sandy soil, southern New Jersey to 
pegs and Georgia. Bermuda; Bahamas. May- 
ug. 


8. SHERARDIA [Dill.] L. Sp. Pl. 102. 1753. 


Slender annual procumbent or diffuse herbs, with verticillate spiny-pointed leaves, and 
small nearly sessile pink or blue flowers, in terminal and axillary involucrate heads. Calyx- 
tube obovoid, its limb 4-6-lobed, the lobes lanceolate, persistent. Corolla funnelform, 4-5-lobed, 
the tube as long as the lobes or longer. Stamens 4 or 5, inserted on the tube of the corolla; 
filaments slender; anthers small, oblong, exserted. Ovary 2-celled; style 2-cleft at the sum- 
mit; ovules I in each cavity. Fruit didymous, the carpels indehiscent. Seed erect. [Named 
for Dr. William Sherard, 1659-1728, patron of Dillenius.] 

A monotypic genus of the Old World. 


1. Sherardia arvénsis L. Blue Field. 
Madder. Herb Sherard. Spurwort. 
Fig. 3952. 

Sherardia arvensis L. Sp. Pl. 102. 1753. 


Tufted, roughish, stems numerous, prostrate, 
ascending, or decumbent, 310’ long. Leaves in 
4’s, 5’s or 6’s, the upper linear or lanceolate, acute 
and sharp-pointed, rough-ciliate on the margins, 
3-8” long, 1’’-2” wide, the lower often obovate, 
mucronate; flowers in slender-peduncled involu- 
crate heads, the involucre deeply 6-8-lobed, the 
lobes lanceolate, sharp-pointed; corolla-lobes 
spreading; fruit crowned with the 4-6 lanceolate 
calyx-teeth. 

In waste places, Ontario to eastern Massachusetts 


and New Jersey. Also in Bermuda. Adventive from 
Europe. June-July. 


9. ASPERULA‘TL) Sp; Piz103. 4753; 

Erect or ascending branching perennial herbs, with 4-angled stems, verticillate leaves, and 
small white pink or blue flowers in terminal or axillary, mostly cymose clusters. Calyx-tube 
somewhat didymous, the limb obsolete. Corolla funnelform, 4-lobed. Stamens 4, inserted 
on the tube or throat of the corolla; anthers linear or oblong. Ovary 2-celled; ovules I in 
each cavity; style 2-cleft. Fruit globose-didymous, the carpels indehiscent. Seed adherent 
to the pericarp; endosperm fleshy; embryo curved. [Latin diminutive of asper, rough, refer- 
ring to the leaves. ] 

About 80 species, natives of the Old World. Type species: Asperula odorata L. 


Leaves oblong-lanceolate to obovate; fruit hispid. 1. A. odorata. 
Leaves linear, 1” wide or less; fruit smooth. 2. A. galioides. 


to 
Oo’ 
“TI 


GENUS 9. MADDER FAMILY. 


1. Asperula odorata L. Sweet Woodruff. 


Fig. 3953. 
Asperula odorata L. Sp. Pl. 103. 1753. 

Stems erect, slender, smooth. Leaves usually 
in 8’s (6’s-o’s), thin, oblong-lanceolate, acute or 
obtuse, mucronate, I-nerved, roughish on the mar- 
gins, 6-18” long, the lower smaller, often obo- 
vate or oblanceolate; peduncles terminal and ax- 
illary, slender; cymes several-flowered; flowers 
white or pinkish, 12” long; pedicels 1-2” long; 
fruit very hispid, about 1” broad. 

In waste places, New Brunswick, N. J. Fugitive 
from Europe. Other English names are hay-plant, 
mugwet or mugget, rockweed, sweet hairhoof, wood- 
rip, woodrowel, star-grass, and sweet-grass. May-July. 

Asperula arvénsis L., another European species, 
with terminal capitate flowers, and linear obtuse 


leaves, has been found in waste places on Staten 
Island. 


2. Asperula galioides Bieb, Bedstraw Asperula. 
Fig. 3954. 


Asperula galioides Bieb. Fl. Taur. Cauc. 1: 191. 1808. 


Glaucous, glabrous, stems erect or ascending, 23° high or 
less. Leaves linear, rigid, involute-margined, 3-12’ long, about 
a” . : oy) ’ ; ’ - 
1” wide, whorled in 5’s—to’s (often in &’s), subulate-tipped or 
mucronate; cymes panicled; flowers white; fruit smooth. 


In fields, Connecticut to Michigan. Adventive from Europe. 
May-July. 


w 


Family 37. CAPRIFOLIACEAE Vent. Tabl. 2: 593. 1799. 
HONEYSUCKLE FAMILY 

Shrubs, trees, vines, or perennial herbs, with opposite simple or pinnate leaves, 
and perfect, regular or irregular, mostly cymose flowers. Stipules none, or some- 
times present. Calyx-tube adnate to the ovary, its limb 3—5-toothed or 3~5-lobed. 
Corolla gamopetalous, rotate, campanulate, funnelform, urn-shaped, or tubular, the 
tube often gibbous at the base, the limb 5-lobed, sometimes 2-lipped. Stamens 5 
(very rarely 4), inserted on the tube of the corolla and alternate with its lobes; 
anthers oblong or linear, versatile. Ovary inferior, 1—6-celled; style slender; 
stigma capitate, or 2—-5-lobed, the lobes stigmatic at the summit; ovules anatropous, 
I or several in each cavity. Fruit a 1-6-celled berry, drupe, or capsule. Seeds 
oblong, globose, or angular ; seed-coat membranous or crustaceous, smooth or can- 
cellate; embryo usually small, placed near the hilum; radicle terete; cotyledons 
ovate. 


About 10 genera and 300 species, mostly natives of the northern hemisphere, a few in South 
America and Australia. 


Corolla rotate or urn-shaped; flowers in compound cymes; styles deeply 2—5-lobed; shrubs or trees. 


Leaves pinnate; drupe 3—5-seeded. 1. Sambucus. 

Leaves simple; drupe 1-seeded. 2. Viburnum. 
Corolla tubular or campanulate, often 2-lipped; style slender. 

Erect perennial herbs; leaves connate. 3. Triosteum. 


Creeping, somewhat woody herb; flowers long-peduncled, geminate. 4. Linnaea. 


268 CAPRIFOLIACEAE, Vor. II. 


Shrubs or vines. 
Fruit a few-seeded berry. 


Corolla short, campanulate, regular, or nearly so. 5. Symphoricarpos. 
Corolla more or less irregular, tubular or campanulate. 6. Lonicera. 
Fruit a 2-celled capsule; corolla funnelform. 7. Diervilla. 


1. SAMBUCUS [Tourn.] L. Sp. Pl. 269. 1753. 


Shrubs or trees (or some exotic species perennial herbs), with opposite pinnate leaves, 
serrate or laciniate leaflets, and small white or pinkish flowers in compound depressed or 
thyrsoid cymes. Calyx-tube ovoid or turbinate, 3-5-toothed or 3-5-lobed. Corolla rotate or 
slightly campanulate, regular, 3-5-lobed. Stamens 5, inserted at the base of the corolla; 
filaments slender; anthers oblong. Ovary 3-5-celled; style short, 3-parted; ovules I in each 
cavity, pendulous. Drupe berry-like, containing 3-5, I-seeded nutlets. Endosperm fleshy; 
embryo nearly as long as the seed. [Latin name of the elder. ] 

About 25 species, of wide geographic distribution. In addition to the following, about 10 others 
occur in western North America. Type species: Sambucus nigra L. 

Cyme convex; fruit purplish black. 1. S. canadensis. 
Cyme thyrsoid-paniculate, longer than broad; fruit red.. 2. S. racemosa. 


1. Sambucus canadénsis L. Amer- 
ican Elder. Sweet or Common 
Elder. Fig. 3955. 

Sambucus canadensis L. Sp. Pl. 269. 1753. 


A shrub, 4°-10° high, glabrous or very 
nearly so, the stems but little woody, the 
younger ones with large white pith. Leaflets 
5-I1, usually 7, ovate or oval, acuminate or 
acute at the apex, short-stalked, glabrous 
above, sometimes pubescent beneath, 2-5’ 
long, sharply serrate, sometimes stipellate; 
cymes convex, broader than high; flowers 
white, about 14” broad; drupe deep purple 
or black, nearly 3” in diameter; nutlets 
roughened. 

In moist soil, Nova Scotia to Florida, west 
to Manitoba, Kansas and Texas. Also in the 
West Indies. Ascends to 4000 ft. in North 
Carolina. Called also elder-blow, elder-berry. 
The flowers and fruit have strong medicinal 
properties. Leaves heavy-scented when crushed, 
nae of young shoots often stipulate. June— 
uly. 


2. Sambucus racemosa L. Red- 
berried Elder. Fig. 3956. 


S. racemosa L. Sp. Pl. 270. 1753. 
S. pubens Michx. Fl. Bor. Am. 1: 181. 1803. 


A shrub, 2°-12° high, the twigs and leaves 
commonly pubescent; stems woody, the 
younger with reddish-brown pith. Leaflets 
5-7, ovate-lanceolate or oval, acuminate at 
the apex, often narrowed and usually in- 
equilateral at the base, 2-5’ long, not stipel- 
late, sharply serrate; cymes thyrsoid, longer 
than broad; flowers whitish, turning brown 
in drying; drupe scarlet or red, 2’-3” in 
diameter; nutlets very minutely roughened. 


In rocky places, Newfoundland to Alaska, 
British Columbia, Georgia, Michigan, Colo- 
rado and California. Fruit rarely white. April— 
May. Called also mountain elder. Poison- 
elder (Me.).  Boor- or bore-tree. Boutry. 
Ascends to 5000 ft. in Virginia. Sambucus 
pibens dissécta Britton, Mem. Torr. Club 5: 
304. 1894, is a race with leaflets laciniate. 
Lake Superior and Pennsylvania. 


Sambucus laciniata Mill., a cut-leaved race 
of the related European S. nigra L., has been 
found at Cape May, N. J., perhaps escaped 
from cultivation. | 


GENUS 2. HONEYSUCKLE FAMILY. 269 


2. VIBURNUM [Tourn.] L. Sp. Pl. 267. 1753. 


Shrubs or trees, with entire dentate or lobed, sometimes stipulate leaves, and white or 
rarely pink flowers in compound cymes, the outer flowers sometimes radiant and neutral. 
Calyx-tube ovoid or turbinate, its limb short, 5-toothed. Corolla rotate or short-campanulate 
in our species, regular, 5-lobed. Stamens 5, inserted on the tube of the corolla; anthers 
oblong, exserted. Ovary 1-3-celled; style short, 3-lobed or 3-parted; ovules solitary in each 
cavity, pendulous. Drupe ovoid or globose, sometimes flattened, I-seeded. Seed compressed; 
endosperm fleshy; embryo minute. [The ancient Latin name.] 

About 100 species, of wide geographic distribution. Besides the following, about 5 others 
occur in the southern and western parts of North America. Type species: Viburnum Tinus L. 

* Outer flowers of the cyme large, radiant; drupe red. 
Leaves doubly serrate, pinnately veined. 1. V. alnifolium. 
Leaves 3-lobed, palmately veined. 2. V. Opulus. 
** None of the flowers radiant; drupe blue or black (red in no. 3). 
I. Leaves palmately veined, or 3-ribbed. 

Cymes %’-1’ broad, the rays short; drupe red. 3. V. pauciflorum. 
Cymes 1%4’-2%’ broad, the rays slender ; drupe nearly black. 4. V.acerifolium. 
2. Leaves pinnately veined. 

a. Leaves coarsely dentate, the veins mostly prominent beneath. 


Leaves very short-petioled, pubescent. 5. V. pubescens. 
Petioles 3”—20” long. 
Leaves glabrous, or with tufts of hairs in the axils beneath. 6. V. dentatum. 


Leaves pubescent beneath, the pubescence more or less stellate. 
Drupe globose-ovoid; eastern and southern. 
Veins of the leaves not very prominent. 7. V.scabrellum. 
Veins very prominent on the under sides of the leaves. 8. V. venosum. 
Drupe oblong, twice as long as thick; western. g. V.molle. 


b. Leaves entire, crenulate, or serrulate, the veins not prominent. 


Native shrubs; drupes blue or black. 
Cymes manifestly peduncled. 


Peduncles shorter than the cyme; leaves crenulate. 10. V. cassinoides. 
Peduncle equalling or longer than the cyme; leaves mostly entire. 11. V. nudum. 
Cymes sessile, or nearly so. 
Leaves prominently acuminate. 12. V. Leuwtago. 
Leaves obtuse, or merely acute. 
Leaves and scarcely winged petioles glabrous, or nearly so. 13. V. prunifolium. 
Veins of lower leaf-surfaces and winged petioles tomentose. 14. V.rufidulum. 
European shrub, or small tree, escaped from cultivation; drupes red. 15. V. Lantana. 


1. Viburnum alnifolium Marsh. Hobble-bush. American Wayfaring Tree. 
Moose-bush or -berry. Fig. 3957. 


V.alnifolium Marsh. Arb. Am.102. 1785. 
Viburnum lantanoides Michx. FI, Bor. 

‘Agee 79, 1803: 

A shrub, with smooth purplish bark, 
sometimes reaching a height of 10°, 
widely and irregularly branching, the 
branches often procumbent and root- 
ing, the youngest twigs scurfy. Leaves 
orbicular, or very broadly ovate, 
strongly innately veined, short- 
acuminate or acute at the apex, 
usually cordate at the base, finely 
stellate-pubescent, or at length gla- 
brous above, scurfy with stellate pu- 
bescence on the veins beneath, finely 
serrate all around, 3-8 broad; peti- 
oles 4’-14’ long; cymes sessile, 3’-5’ 
broad, the exterior flowers usually 
radiant and neutral, about 1’ broad; 
drupes red, becoming purple, ovoid- 
oblong, 5-6” long; stone 3-grooved 
on one side, I-grooved on the other. 

In low woods, New Brunswick to 
North Carolina, Ontaric, Tennessee 
and Michigan. Leaves of shoots from 
cut stumps thin, ovate, corsely toothed. 
May-June. Tangle-legs or -foot. Dog- 
wood. Dogberry. Trip-toe. Witch-hopple 
or-hobble. Winter-buds naked. 


270 CAPRIFOLIACEAE. Vor. IIL. 


2. Viburnum Opulus L. Cranberry- 
tree. Wild Guelder-rose.. High 
Bush-cranberry. Fig. 3958. 

Viburnum Opulus L. Sp. Pl. 268. 1753. 
V. trilobum Marsh. Arb. Am. 162. 1785. 
V. americanum Mill. Gard. Dict. Ed. 8, no. 

8. 1768. 

A shrub sometimes 12° high, with nearly 
erect smooth branches. Leaves broadly 
ovate, sometimes broader than long, gla- 
brous, or with scattered hairs above, 
more or less pubescent on the veins be- 
neath, rather deeply 3-lobed, rounded or 
truncate and 3-ribbed at the base, the 
lobes divergent, acuminate, coarsely den- 
tate; petioles 3-1’ long, glandular above; 
cymes peduncled, 3-4’ in diameter, the 
exterior flowers radiant, neutral, 4-1’ 
broad; drupes globose, or oval, 4”—5” in 
diameter, red, very acid, translucent; 
stone orbicular, flat, not grooved. 

In low grounds, Newfoundland to Brit- 
ish Columbia, New Jersey, Michigan, Iowa, 
South Dakotaand Oregon. Also in Europe 
and Asia. Among many English names are 
marsh-, rose- or water-elder, white dog- 

EZ wood, whitten-tree, dog rowan-tree, gaiter- 
tree or gatten, cherry-wood, May-rose,, squaw-bush, cramp-bark. In cultivation, the snowball. 
Gadrise. Red elder. Love-roses. Witch-hopple 
or -hobble. Pincushion-tree. June-July. 

3. Viburnum pauciflorum Pylaie. 


Few-flowered Cranberry-tree. Fig.3959. 
V.pauciflorum Pylaie;T.&G.FI.N.A.2:17. 1841. 
Viburnum Opulus var. eradiatum Oakes, Hovey’s 

Mag. 7: 183. 1841. 

A straggling shrub, 2°-6° high, with twigs 
and petioles glabrous or nearly so. Leaves 
broadly oval, obovate, or broader than long, 
5-ribbed, truncate or somewhat cordate at 
the base, mostly with 3 rather shallow lobes 
above the middle, coarsely and unequally 
dentate, glabrous above, more or less pu- 
bescent on the veins beneath, 13-3’ broad; 
cymes peduncled, short-rayed, 3-1’ broad; 
flowers all perfect and small; drupes glo- 
bose to ovoid, light red, acid, 4’-5” long; 
stone flat, orbicular, scarcely grooved. 

In cold mountain woods, Newfoundland to 
Alaska, south to Maine, New Hampshire, Ver- 
mont, Pennsylvania, in the Rocky Mountains to 
Colorado, and to Washington. June-July. 


a lees || go 
4. Viburnum acerifolium L. Maple- 
leaved Arrow-wood. Fig. 3960. 

V. acerifolium L. Sp. Pl. 268. 1753. 

A shrub 3°-6° high, with smooth gray 
slender branches, and somewhat pubescent 
twigs and petioles. - Leaves ovate, orbicu- 
lar, or broader than long, cordate or trun- - 
cate at the base, pubescent on both sides, 
or becoming glabrate, 2’-5’ broad, mostly 
rather deeply 3-lobed, coarsely dentate, the 
lobes acute or acuminate; petioles 2’-1’ 
long; cymes long-peduncled, 12-3’ broad; 
flowers all perfect, 2-3’ broad; drupe 
nearly black, 3-4” long, the stone lenticu- 
lar, faintly 2-ridged on one side and 2- 
grooved on the other. 

In dry or rocky woods, New Brunswick to 
Georgia, Alabama, Ontario, Michigan and 
Minnesota. Upper leaves sometimes merely 
toothed, not lobed. May-June. Squash-berry. 
Maple-leaf guelder-rose. Dockmakie. 


GENUS 2. HONEYSUCKLE FAMILY. 271 


5. Viburnum pubéscens (Ait.) Pursh. 
Downy-leaved Arrow-wood 
Fig. 3961. 
Viburnum dentatum var. pubescens Ait. Hort. Kew. 

UE Sy ieRe Bi Astley 
V. pubescens Pursh, Fl. Am. Sept. 202. 1814. 

A shrub, 2°-5° high, with numerous straight 
and slender gray branches. Leaves sessile, or 
on petioles less than 3” long, ovate or oval, 
rounded or slightly cordate at the base, acute or 
acuminate at the apex, coarsely dentate, 13’-3' 
long, densely velvety-pubescent beneath, glabrous, 
or with scattered hairs above, or rarely glabrate 
on both surfaces; cymes peduncled, 12’-23’ broad, 
the flowers all perfect; drupes oval, nearly black, 
about 4” long; stone slightly 2-grooved on both 
faces. 

Rocky woods and banks, Quebec and Ontario to 
Manitoba, south, especially along the Alleghanies to 
Georgia and to Illinois, lowa, Michigan and Wyoming. 
The leaves of shoots are sometimes entire or nearly 
so. June-July. 


6. Viburnum dentatum L. Arrow- 
wood. Fig. 3962. 
Viburnum dentatum L. Sp. Pl. 268. 1753. 

A shrub with slender glabrous gray 
branches, sometimes reaching a height of 
15°. Twigs and petioles glabrous; petioles 
3-12” long; leaves ovate, broadly oval or 
orbicular, rounded or slightly cordate at 
the base, acute or short-acuminate at the 
apex, prominently pinnately veined, coarsely 
dentate all around, 133° broad, glabrous 
on both sides, or sometimes pubescent with 
simple hairs in the axils of the veins be- 
neath; cymes long-peduncled, 2-3’ broad; 
flowers all perfect; drupe globose-ovoid, 
about 3” in diameter, blue, becoming nearly 
black; stone rather deeply grooved on one 
side, rounded on the other. 

In moist soil, New Brunswick to Ontario, 
south along the mountains to Georgia and to 
: western New York, Michigan and Minnesota. 
¥ Called also mealy-tree. Withe-rod or -wood. 
May-June. 


7. Viburnum scabréllum (T. & G.) 
Chapm. Roughish Arrow-wood. 


Fig. 3963. 


V. dentatum semitomentosum Michx. FI. Bor. 
Ami 3i79; 1803. f 
V. dentatum var. (?) scabrellum T. & G. FI. 
~IN S Ne” Mieye 
V. scabrellum Chapm. FI. S. States 172. 1860. 
V. semitomentosum Rehder, Rhodora 6: 59. 
1904. 


Similar to the preceding species but the 
twigs, petioles, rays of the cyme and lower 
surfaces of the leaves more or less densely 
stellate-pubescent; petioles short and stout- 
er; leaves usually larger, crenate or dentate, 
- commonly somewhat pubescent above; drupe 
globose-ovoid, blue, 4” in diameter, its stone 
similar to that of V. dentatum. 


Woodlands and river banks, southern Penn- 
sylvania to Kentucky, Florida and Texas. Re- 
ferred, in our first edition, following previous 
authors, to V. molle Michx., a species long 
misunderstood. 


272 CAPRIFOLIACEAE. Vor. III. 


8. Viburnum venosum Britton. Veiny 
Arrow-wood. Fig. 3964. 
Viburnum venosum Britton, Man. 871. 1901. 
V. venosum Canbyi Rehder, Rhodora 6: 60. 

1904. 

A shrub, 6° high or less, the bark grayish- 
brown, the young twigs stellate-pubescent 
or stellate-tomentose. Leaves ovate to 
orbicular, 2’-5’ long, firm, coarsely and 
sharply dentate, glabrous or with sparse 
pubescence on the upper side when young, 
stellate-pubescent, at least on the veins, 
beneath, the petioles 1’—?’ long; cymes long- 
stalked, often 3-4’ broad, stellate-pubescent 
or glabrate; drupe globose or short-oval. 
3-4” in diameter, nearly black. 

Thickets, eastern Massachusetts to Ne 
Jersey, Pennsylvania and Virginia. Shoots 


sometimes bear reniform, very thin subcordate 
leaves with minute distant teeth. June-July. 


g. Viburnum mdlle Michx.  Soft- 
leaved Arrow-wood. Fig. 3965. 
V. moile Michx. Fl. Bor. Am. 1: 180. 1803. 
Viburnum Demetrionis Deane & Robinson, 

Bot. Gaz. 22: 167. pl. 8. 1806. 

A shrub about 12° high, the older twigs 
at length grayish black, the bark exfoliating. 
Bud-scales acutish, ciliolate; leaves broadly 
ovate or nearly orbicular, short-acuminate 
at the apex, cordate or truncate at the base, 
3-5’ long, coarsely dentate, glabrous and 
bright green above, soft-pubescent and paler 
beneath, some of the pubescence stellate; 
petioles 8’-20” long; stipules linear-filiform, 
2”-5” long; cymes terminal, peduncled, 
4-7-rayed, glandular-puberulent ; calyx-teeth 
ciliate; drupe oblong, obtuse at both ends, 
about 5” long and 23” broad, much flattened, 
with 2 grooves when dry. 

Bluffs and rocky woods, Kentucky, Missouri 


and Iowa. June. 
fy) 


to. Viburnum cassinoides L. 
Withe-rod. Appalachian Tea. 
Fig. 3960. 

Viburnum cassinoides L, Sp. Pl. Ed. 2, 

384. 1762. 

Viburnum nudum var. cassinoides T. & 

G. BION. A: 2:14. 2184ae 

A shrub, 2°-12° high, with ascend- 
ing gray branches, the twigs some- 
what scurfy, or glabrous. Leaves 
ovate or oval, thick, pinnately veined, 
narrowed or sometimes rounded at 
the base, acute or blunt-acuminate at 
the apex, 1-3’ long, usually crenu- 
late, rarely entire, glabrous or very 
nearly so on both sides; peduncle 
shorter than or equalling the cyme; 
drupe pink, becoming dark blue, 
globose to ovoid, 3’-5” long; stone 
round or oval, flattened. 

In swamps and wet soil, Newfound- 
land to Manitoba, Minnesota, New 
Jersey, Georgia and Alabama. June— 
July. False Paraguay-tea. 


GENUS 2. 


11. Viburnum nudum L. Larger or 
Naked Withe-rod. Fig. 3967. 


Viburnum nudum L. Sp. Pl. 268. 1753. 
Viburnum nudum var. Claytoni T. & G. Fl. N. A. 
Set. OAT. 


Similar to the preceding species, but usually 
a larger shrub, sometimes 15° high. Leaves 
oval, oval-lanceolate, or obovate, entire or 
obscurely crenulate, mostly larger (sometimes 
9’ long), narrowed at the base, acute or obtuse 
at the apex, more prominently veined, some- 
times scurfy on the upper surface; peduncle 
equalling or exceeding the cyme. 


Connecticut to Florida, west to 
Kentucky and Louisiana. Blooms a little later 
than the preceding species. Bilberry. 
berry. Possum- or Shawnee-haw. 


In swamps, 


Nanny- 


, Wk ryaays 
AHR ESA 3-4) CBOes 
Ae 4, <o 
AINA see, Sainieesions 
SE i ROO We: 
wy LSI54) Mee b 


CAS 
GQ 


@ Hy Rel 40) 
Wn VISA 
CNS §25° 


HONEYSUCKLE FAMILY. 


WZ, Oty. 


sa i 


12. Viburnum Lentago L. Nanny- 
berry. Sheep- or Sweet-berry. 
Sweet Viburnum. Fig. 3968. 

Viburnum Lentago L. Sp. Pl. 268. 1753. 


A shrub, or often a small tree, some- 
times 30° high and with a trunk diameter 
of 10°. Winter buds acuminate, gla- 
brous; leaves slender-petioled, ovate, 
mostly rounded at the base, acuminate 
at the apex, 2’-4’ long, glabrous on both 
sides, or rarely a little pubescent beneath, 
sharply serrulate; petioles often broad- 
ened and wavy-margined, 9’—12” long; 
cyme sessile, several-rayed, 2’-5’ broad; 


\ PIS NG ARE, drupes oval to subglobose, bluish-black 

DEON Sores with a bloom, sweet and edible, 5’-6” 
ean ae ais [AO een ove long; stone very flat, circular or oval. 

ASS SA BIp ERE In rich soil, Quebec to Hudson Bay, 

AY; <e= Manitoba, New Jersey, along the Allegha- 


13. Viburnum prunifolium L. Black 
Haw. Stag-bush. Sloe. Fig. 3969. 


Viburnum prunifolium L. Sp. Pl. 268. 1753. 


A shrub or small tree somewhat similar to 
the preceding species; but the winter buds 
smaller, less acute, often reddish-pubescent. 
Leaves shorter-petioled, ovate or broadly oval, 
obtuse or acutish but not acuminate at the 
apex, narrowed or rounded at the base, 1-3’ 
long, finely serrulate, glabrous or nearly so; 
petioles rarely margined; cyme sessile, several- 
rayed, 2’-4’ broad; flowers expanding with the 
leaves or a little before them; drupe oval, 
bluish-black and glaucous, 4-5” long; stone 
very flat on one side, slightly convex on the 
other, oval. : 


In dry soil, Connecticut to Georgia, west to 
Michigan, Kansas and Texas. Wood hard, red- 
dish-brown; weight per cubic foot 52 lbs. April— 
June. Sheep- or nanny-berry. Fruit ripe in Sep- 
tember, sweet and edible. A race, Viburnum 
prunifolium globdsum Nash, has the drupe glo- 
bose, about 3” in diameter, smaller; New Jersey 
and eastern Pennsylvania. 


18 


nies to Georgia, and to Indiana, Kansas and 
Colorado. | Wood orange-brown, hard; 
weight 45 lbs. to the cubic foot. May-June. 
Fruit ripe in October. Nanny-bush. Black 
thorn or haw. Nanny-plum. Tea-plant 
(Wis.). Wild raisin. 


274 CAPRIFOLIACEAE. Vor. III. 


14. Viburnum rufidulum Raf. Southern 
Black Haw. Fig. 3970. 


Viburnum rufidulum Raf. Alsog. Am. 56. 1838. 
Viburnum prunifolium var. ferrugineum T. & G. FI. 

N. A. 2:15. 1841. Not V. ferrugineum Raf, 1838. 
Viburnum rufotomentosum Small, Bull. Torr. Club 

23: 410. 1896. 

A small tree, becoming 20° high. Leaves 
elliptic to obovate, mostly obtuse at the apex, 
finely and sharply serrate or serrulate, narrowed 
or obtuse at the base, the veins brown-tomentose 
beneath; petioles 3’-8” long, winged, brown- 
tomentose; cymes large, sessile, or very short- 
peduncled, the principal rays 3-5, mostly 4; flow- 
ers 3-33” broad; drupe oval, 5-7” long, blue 
with a bloom; seed nearly orbicular. 
prenz6, In woods and thickets, New Jersey to Missouri, 

QP VV 


Kansas, Florida and Texas. Ascends to 3500 ft. in 
Virginia. April-May. Fruit ripe Aug.—Sept. 


cD. dj ab 
sevianeeutasce 
a} 
J 


Viburnum obovatum Walt., admitted into our 
first edition as recorded from Virginia, is not defi- 
nitely known to range north of South Carolina. 


15. Viburnum LantanaL. Way- 
faring Tree. Fig. 3971. 


Viburnum Lantana L. Sp. Pl. 268. 1753. 


aS 


A shrub, or small tree, sometimes 
12 ft. high, widely branched, the 
winter-buds naked. Young twigs, 
buds and petioles densely stellate- 
tomentose. Leaves ovate to ovate- 
elliptic, serrulate, dark green and 
loosely stellate-pubescent or glabrous 
above, paler and more or less stel- 
late-tomentose beneath, 2’-4’ long, 
rounded or acutish at the apex, sub- 
cordate at the base, the petioles stout 
and short; cymes _ short-stalked, 
stellate-tomentose, densely many- 
flowered, the flowers all alike, 3-4” 
broad; drupe red, oval, 4-5” long, 
its stone grooved. 


U 


LAVE iM 


// 


» 


) 
Wi 


\ 
p Ny. i 


7h 
r 


1 


i> 


LASS 


—fX 


Ej 
a 
7p 


ro4 


Roadsides, escaped from cultivation L : : 
in New England. Native of Europe and Asia. The naked winter-buds ally this species to V. 
alnifolium Marsh. May-July. 


3. TRIOSTEUM L. Sp. Pl. 176. 1753. 


Perennial herbs, with simple terete stems and opposite connate-perfoliate or sessile leaves 
narrowed below the middle. Flowers axillary, perfect, solitary or clustered, sessile, yellowish, 
green, or purplish, 2-bracted. Calyx-tube ovoid, its limb 5-lobed, the lobes elongated, per- 
sistent and sometimes foliaceous in our species. Corolla-tube narrow or campanulate, gibbous 
at the base, the limb oblique. Stamens 5, inserted on the corolla-tube; filaments very short; 
anthers linear, included. Ovary 3-5-celled; ovules 1 in each cavity; style filiform; stigma 
3-5-lobed. Drupe coriaceous, orange or red, enclosing 2-3 (rarely 4-5) 1-seeded nutlets. 
Endosperm fleshy; embryo minute. [Greek, three-bone, from the 3 bony nutlets. ] 


Six known species, the following of eastern North America, two Japanese, one Himalayan. 
Type species: Triosteum perfoliatum L. 


Leaves ovate or oval; flowers purplish or dull red. 
Leaves, or some of them, connate-perfoliate; fruit orange-yellow. 1. T. perfoliatum. 
Leaves narrowed to a sessile base; fruit orange-red. 2. T. aurantiacum. 
Leaves lanceolate or oval-lanceolate; flowers yellowish. 3. T. angustifolium. 


GENUS 3. HONEYSUCKLE FAMILY. 275, 


1. Triosteum perfoliatum L. Fever- | 
wort. Horse-Gentian. Fig. 3972. 


Triosteum perfoliatum L. Sp. Pl. 176. 1753. 


Stem erect, stout, finely glandular-pubescent, 
2°-4° high. Leaves ovate to broadly oval, 4’-0’ 
long, 2-4’ wide, acute or acuminate at the 
apex, abruptly narrowed at the base, connate- 
perfoliate, soft-pubescent beneath, somewhat 
hairy above, the margins entire or sinuate; 
bracts linear; corolla dull purplish-brown, 
greenish below, 6-10” long, viscid-pubescent, 
about the length of the calyx-lobes, the limb 
nearly regular; filaments bearded; drupe 4’’-6” 
long, obovoid-globose, orange-yellow, densely 
and finely pubescent; nutlets usually 3. 


In rich soil, Massachusetts to Alabama, Ken- 
tucky and Kansas. Fever-root. Wild or wood 
ipecac. Tinker’s-weed. Wild coffee. Horse-gin- 
seng. White gentian. Genson (N.C.). Ascends F e a 
3000 ft. in Virginia. May-July. 2. Triosteum aurantiacum Bicknell. 


Scarlet-fruited Horse-Gentian. 
Fig. 3973. 


T. aurantiacum Bicknell, Torreya I: 26. 1901. 

Stems stout, erect, 2°-4° tall, glandular- 
puberulent to hirsute. Leaves thin, ovate, 
ovate-oblong or oblong-lanceolate, 5’-10’ long, 
acuminate at the apex, more or less dilated at 
the base, but not connate-perfoliate, minutely 
soft-pubescent beneath, thinly appressed- 
pubescent above or nearly glabrous; corolla 
dull-red, 7-10” long, often shorter than the 
calyx-lobes; filaments bearded throughout or 
nearly so; drupe oblong-ovoid or obovoid, 
6-7” long, orange-red, densely short-pubes- 
cent; nutlets usually 3. 

In rich woods and thickets, New Brunswick to 
Quebec, Minnesota, Missouri and North Carolina. 
May-June. 


Triosteum angustifolium L. Yellow 
or Narrow-leaved Horse-Gentian. 


Fig. 3974. 
Triosteum angustifolium L. Sp. Pl. 176. 1753. 


Stem slender and hirsute-pubescent, 1°-3° 
high. Leaves lanceolate or oval-lanceolate, 
acute or acuminate at the apex, 3’-5’ long, 
2-13’ wide, rough-pubescent, tapering to the 
sessile base, or the lower smaller, obtuse and 
spatulate; corolla yellowish, 6’—-7” long; flow- 
ers commonly solitary in the axils. 


In rich soil, Connecticut and Long Island to 
New Jersey, Pennsylvania, Virginia, Alabama, 
Illinois and Louisiana. May—Aug. 


4. LINNAEA [Gronov.] L. Sp. Pl. 631. 1753. 


Creeping, somewhat woody herbs, with opposite evergreen petioled obovate or orbicular 
leaves, and perfect pink or purplish flowers borne in pairs at the summit of elongated ter- 
minal peduncles. Calyx-tube ovoid, the limb 5-lobed. Corolla campanulate or funnelform, 
5-lobed, the lobes imbricate. Stamens 4, inserted near the base of the corolla-tube, didyn- 
amous, included. Ovary 3-celled, 2 of the cavities with several abortive ovules, the other 
with 1 perfect pendulous ovule. Fruit nearly globose, 3-celled, 2 of the cells empty, the 


276 CAPRIFOLIACEAE. Vor. III. 


other with a single oblong seed. Endosperm fleshy; embryo cylindric. [Named by Grono- 
vius for Linnaeus, with whom the plant was a favorite.] 


Three or four species, of the north temperate zone. 
Type species: Linnaea boredlis L., of Europe. 


1. Linnaea americana Forbes. Twin-flower. 
Ground-vine. Deer-vine. Fig. 3975. 
Linnaea americana Forbes, Hort. Woburn. 135. 1825. 
L. borealis var. americana Rehder, Rhodora 6: 56. 1904. 
Branches slender, slightly pubescent, trailing, 62° 
long. Petioles 1’-2” long; leaves obscurely crenate, 
thick, 3-8” wide, sometimes wider than long; 
peduncles slender, erect, 2-bracted at the summit, 
2-flowered (or rarely proliferously 4-flowered) ; 
pedicels filiform, 3’—10” long, 2-bracteolate at the 
summit; flowers nodding, 4”-6” long, fragrant; 
corolla funnelform; calyx-segments about 1” long; 
ovary subtended by a pair of ovate glandular scales 
which are connivent over the fruit or adnate to it. 
In cold woods, mountains of Maryland, New Jersey, 
Long Island, north to Newfoundland, west through Brit- 
ish America to Alaska and Vancouver, south to Mich- 
igan, in the Rocky Mountains to Colorado and in the 
Sierra Nevada to California. Has been considered 
identical with the similar L. borealis L. of Europe and 
Asia, which has a nearly bell-shaped corolla and longer 

calyx. Twin sisters. Two-eyed berries. June-Aug. 


5. SYMPHORICARPOS [Dill.] Ludwig, Def. 35. 1760. 


Shrubs, with opposite deciduous short-petioled simple leaves, and small white or pink, 
perfect flowers, in axillary or terminal clusters. Calyx-tube nearly globular, the limb 
4-5-toothed. Corolla campanulate or salverform, regular, or sometimes gibbous at the base, 
4-5-lobed, glabrous or pilose in the throat; stamens 4 or 5, inserted on the corolla. Ovary 
4-celled, 2 of the cavities containing several abortive ovules, the other two each with a single 
suspended ovule; style filiform; stigma capitate, or 2-lobed. Fruit an ovoid or. globose 
4-celled 2-seeded berry. Seeds oblong; endosperm fleshy; embryo minute. [Greek, fruit 
borne together, from the clustered berries. ] 


About 10 species, natives of North America and the mountains of Mexico. Known as SE 
Peter’s-wort. Type species: Lonicera Symphoricarpos L. 
Fruit white; style glabrous. 
Stamens and style included; clusters usually few-flowered. 1. S. racemosus. 
Stamens and style somewhat exserted ; clusters many-flowered. 2. S. occidentalis. 
Fruit red; style bearded. , 3. S. Symphoricarpos. 


1. Symphoricarpos racemésus Michx. Snowberry. Wax-berry. Fig. 3976. 
Symphoricarpos racemosus Michx: FI. Bor. 

ATH ets OF eT OOs 
S. racemosus var. pauciflorus Robbins; A. 

Gray, Man. Ed. 5, 203. 1867. 

S. paucifiorus Britton, Mem. Torr. Club 5: 

305. 1894. 

An erect or diffuse shrub, 1°-4° high, 
glabrous, or usually so, the branches 
slender. Petioles about 2” long; leaves 
oval, obtuse at each end, sometimes 
pubescent or whitened beneath, 3’—2’ 
long, entire, undulate, or those of young 
shoots sometimes dentate; axillary 
clusters few-flowered, the terminal one 
mostly interruptedly spicate; corolla 
campanulate, about 3” long, slightly gib- 
bous at the base, bearded within; style 
glabrous; stamens and style included; 
berry snow-white, globose, loosely cellu- 
lar, 23’-5”’ in diameter. 

In rocky places and on river shores, 
Nova Scotia and Quebec to British Colum- Yyzes 
bia, south to Pennsylvania, Kentucky, Min- {/”/”~= 
nesota, South Dakota, Montana and inf, 
California. Commonly planted and some-\\" 
times escaped from cultivation. Races differ 
in size, habit and pubescence. Snowdrop- 
berry. Egg-plant. June—Sept. 


in 


GENUS 5. HONEYSUCKLE FAMILY. 


3. Symphoricarpos occidentalis 
Hook. Wolfberry. Fig. 3977. 


Symphoricarpos occidentalis Hook. Fl. Bor. 
Aine 285. 91833. 


Similar to S. racemosus but stouter, with 
larger leaves, 1-3’ long, more or less pubes- 
cent beneath, entire, or often undulate- 
crenate; petioles 2’-3” long; axillary 
clusters spicate, many-flowered, 67-12” 
long; corolla funnelform-campanulate, 3” 
long, lobed to beyond the middle; stamens 
and glabrous style somewhat exserted; 
berry nearly globular, white, 47-5” in 
diameter. 


Rocky situations, Illinois, Michigan and 
Minnesota to British Columbia, Kansas and 
Colorado. Buck-bush. June—July. 


4. Symphoricarpos Symphoricarpos 
(L.) MacM. Coral-berry. Indian 
Currant. Fig. 3978. 

Lonicera Symphoricarpos L. Sp. Pl. 175. 1753. 

S. orbiculatus Moench, Meth. 503. 1794. 
Symphoricarpos vulgaris Michx. Fl. Bor. Am. 1: 

106. 1803. 

Symphoricarpos Symphoricarpos MacM. Bull. 

Torr. Club 19: 15. 1892. 

A shrub, 2°-5° high, the branches erect or 
ascending, purplish, usually pubescent. Petioles 
1-2” long; leaves oval or ovate, entire or 
undulate, mostly obtuse at each end, glabrous 
or nearly so above, usually soft-pubescent 
beneath, 1-14’ long; clusters dense, many- 
3. flowered, at length spicate, shorter than the 
4. leaves; corolla campanulate, sparingly pubes- 

cent within, pinkish, about 2” long; style 

bearded; stamens included; berry purplish 
red, ovoid-globose, 12-2” long. 

Along rivers and in rocky places, banks of the Delaware in New Jersey and Pennsylvania, west 
to western New York and South Dakota, south to Georgia, Kansas and Texas. Aliso sparingly 
escaped from cultivation farther east. Fruit persistent after the leaves have fallen. Buck-bush. 
Turkey- or snap-berry. July. 


6. LONICERA L. Sp. Pl. 173. 1753. 


Erect or climbing shrubs, with opposite mostly entire leaves; flowers spicate, capitate 
or geminate, usually somewhat irregular. Calyx-tube ovoid or nearly globular, the limb 
slightly 5-toothed. Corolla tubular, funnelform, or campanulate, often gibbous at the base, 
the limb 5-lobed, more or less oblique, or 2-lipped. Stamens 5, inserted on the tube of the 
corolla; anthers linear or oblong. Ovary 2-3-celled; ovules numerous in each cavity, pen- 
dulous; style slender; stigma capitate. Berry fleshy, 2-3-celled or rarely 1-celled, few-seeded. 
Seeds ovoid or oblong with fleshy endosperm and a terete embryo. [Named for Adam 
Lonitzer, 1528-1586, a German botanist. | 


About 160 species, natives of the north temperate zone, a few in tropical regions. Besides the 
following, some 10 others occur in the western parts of North America. Type species: Lonicera 
Caprifolium L. 

* Climbing or trailing vines; flowers in heads or interrupted spikes; 
upper leaves connate-perfoliate. 
Corolla 2-lipped, the upper lip 4-lobed, the lower entire. 
Corolla glabrous within. 1. L. Caprifolium. 
Corolla pubescent within. 
Leaves pubescent, at least beneath; corolla yellow. 
Leaves pubescent on both sides, at least when young, ciliate; corolla slightly gibbous 
at base. 2. L. hirsuta. 
Leaves glabrous above, pubescent beneath; corolla-tube strongly gibbous at the base. 
3. L. glaucescens. 


Leaves glabrous on both sides, very glaucous beneath. 
Corolla greenish-yellow, the tube somewhat gibbous. 
Corolla-tube 3”-5” long; filaments hirsute at the base. 4. L. dioica. 


278 CAPRIFOLIACEAE, Vor. IIT. 


Corolla-tube 5”—7” long; filaments nearly glabrous. 5. L. Sullivantii. 
Corolla bright yellow or orange, its slender tube not gibbous. 6. L. flava. 


Corolla tubular, the short limb nearly equally 5-lobed. 


7. L. sempervirens. 


** Climbing vine; flowers in pairs on short axillary peduncles. 


8. L. japonica. 


*** Shrubs; flowers in pairs on axillary bracted pedurcles. 


Bracts of the peduncle subulate, linear, minute, or none. 


Leaves rarely cordate, more or less pubescent, or ciliate. 
Leaves pale, or glaucous, thick, strongly reticulate-veined. 


Peduncles shorter than the flowers; fruit blue; leaves ciliate. g. L. coerulea. 
Peduncles equalling the flowers; fruit red; leaves not ciliate. 10. L. oblongifolia. 
Leaves bright green, thin, ciliate, not strongly reticulate; fruit red. 11. L. canadensis. 
Leaves pale, densely pubescent beneath, even when old. 12. L. Xylosteum. 
Leaves cordate, glabrous. 13. L. tatarica. 
Bracts of the peduncle broad, foliaceous. 14. L. involucrata. 


2. Lonicera hirstta Eaton. Hairy 
Honeysuckle. Fig. 3980. 


L. hirsuta Eaton, Man. Ed. 2, 307. 1818. 


Twining, the branches hirsute and glandu- 
lar-pubescent. Upper one or two pairs of 
leaves connate-perfoliate, the others oval or 
ovate, short-petioled or sessile, softly pu- 
bescent beneath, dark green and appressed- 
pubescent above, ciliate, obtuse or obtusish 
at the apex, rounded or narrowed at the 
base, 2’-33’ long; flowers verticillate in 
short terminal interrupted spikes; corolla 
pubescent within, 1-14’ long, viscid-pubes- 
cent without, orange-yellow, turning red- 
dish, the tube slender, somewhat gibbous at 
the base, the limb strongly 2-lipped, about 
as long as the tube or shorter; filaments 
hirsute below. 


In woodlands, Vermont and Ontario to 
Manitoba, Pennsylvania, Ohio and Michigan. 
Rough woodbine. June-July. 


1. Lonicera Caprifolium L. Italian 
or Perfoliate Honeysuckle. 


Fig. 3979. 


L. Caprifolium L. Sp. Pl. 173. 1753. 

L. grata Ait. Hort. Kew. 1: 231. 1789. 

Caprifolium gratum Pursh, Fl. Am. Sept. 
161. re814. 


Climbing high, glabrous and somewhat 
glaucous. Upper one to three pairs of 
leaves connate-perfoliate, glaucous be- 
neath, the others sessile or short-peti- 
oled, oval or obovate, all rounded at 
the base, entire; flowers in terminal 
capitate sessile clusters; corolla gla- 
brous within, 1’-13’ long, purple with- 
out, the limb white within, strongly 
2-lipped; upper lip 4-lobed, the lower 
one narrow, reflexed; tube slightly 
curved, not gibbous; stamens and style 
much exserted; berries red. 


Thickets, New York, New Jersey and 
Pennsylvania to Michigan, Missouri, and 
in the Southern States. Escaped from 
cultivation and naturalized. Native of 
Europe. Called also american or fragrant 
woodbine. May-June. 


GENUS 6. HONEYSUCKLE FAMILY. | 279 
Vy 


WT. 


3. Lonicera glaucéscens Rydb. Douglas’ 
Honeysuckle. Fig. 3981. 


Lonicera Douglasii Hook. Fl. Bor. Am. 1: 282. 
1833. Not Caprifolium Douglasii Lind. 1830. 
Lonicera glaucescens Rydb. Bull. Torr. Club 24: 


go. 1897. 


Similar to the preceding species, the branches 
glabrous. Leaves glabrous above, pubescent, at 
least on the veins, beneath, 13-2’ long, chartace- 
ous-margined, not ciliate, usually only the upper 
pair connate-perfoliate; flowers verticillate in 
a short terminal interrupted spike; corolla yel- 
low, changing to reddish, pubescent or puberu- 
lent without, pubescent within, 1’ long, or less, 
the tube rather strongly gibbous at the base, 
the 2-lipped limb shorter than the tube; sta- 
mens nearly glabrous, or somewhat pubescent; 
style hirsute; both exserted; ovary sometimes 
hirsute. 


Ontario to Alberta, Pennsylvania, North Caro 
lina, Ohio and Nebraska. May—June. 


4. Lonicera didica L. Smoothed- 
leaved or Glaucous Honeysuckle. 
Fig. 3982. 

Lonicera dioica L. Syst. Ed. 12, 165. 1767. 


L. glauca Hill, Hort. Kew. 446. pl. 18. 1769. 
L. parviflora Lam. Encycl. 1: 728. 1783. 


Glabrous throughout, twining or shrubby, 
3°-10° long. Leaves very glaucous beneath, 
14’-3’ long, the upper connate-perfoliate, 
oval, obtuse, the lower sessile or short- 
petioled, narrower; flowers several in a 
terminal cluster, yellowish green and tinged 

with purple, glabrous without, pubescent 
within, the tube 3-4” long, gibbous at the 
base, scarcely longer than the 2-lipped limb; 
stamens hirsute below, exserted with the 
style; berries red, 3’—4” in diameter. 


In rocky ana usually dry situations, Quebec 
to Manitoba, south, especially along the moun- 
tains to North Carolina, and to Ohio and Mis- 
souri. Ascends to 3500 ft. in North Carolina. 
All the leaves of young shoots are sometimes 
connate-perfoliate. Small yellow or crimson 
honeysuckle. Small woodbine. May-June. 


5. Lonicera Sullivantii A. Gray. Sul- 
livant’s Honeysuckle. Fig. 3983. 


Lonicera Sullivantii A. Gray, Proc. Amer. Acad. 
19: 76. 1883. 


Similar to the preceding species, very glau- 
cous. Leaves oval or obovate, glaucous and 
commonly puberulent beneath, obtuse; flowers 
larger than those of the preceding species, the 
tube 5’-7” long, slightly exceeding the limb, 
pale yellow; stamens usually nearly glabrous; 
fruit yellow, 3” in diameter. 


In woodlands, Tennessee, Ohio and western 
Ontario to Iowa, Wisconsin and Minnesota. 
Recorded from Manitoba. May-June. 


CAPRIFOLIACEAE, Vor. III. 


6. Lonicera flava Sims. Yellow Honeysuckle. 
Fig. 3984. 


Lonicera flava Sims, Bot. Mag. pl. 1318 1810. 


Twining to a height of several feet, or trailing, 
glabrous. Leaves broadly oval, or elliptic, entire, 
obtuse at the apex, narrowed or rounded at the base, 
short-petioled, or the upper sessile, green above, 
glaucous beneath, the pairs subtending flowers 
connate-perfoliate; flowers bright orange-yellow, 
fragrant, in a terminal interrupted spike; corolla 
1-13’ long, the slender tube pubescent above within, 
not gibbous at the base, the limb strongly 2-lipped, 
about half as long as the tube; filaments and style 
glabrous, exserted; fruit about 3” in diameter. 


North Carolina to Kentucky, Missouri, Georgia and 


Alabama. 


7. Lonicera sempérvirens L. Trumpet or 
Coral Honeysuckle. Fig. 3985. 


Lonicera sempervirens L. Sp. Pl. 173. 1753. 
Lonicera sempervirens hirsutula Rehder, Rep. Mo. 
Bot Gard. 14: 169 1903. 


Glabrous or somewhat pubescent, high climbing, 
evergreen in the South. Leaves oval, obtuse, 2’—3’ 
long, or the lower ones smaller, narrower and 
acutish, the upper pairs connate-perfoliate, all 
conspicuously glaucous and sometimes slightly 
pubescent beneath, dark green above; flowers 
verticillate in terminal interrupted spikes; corolla 
scarlet or yellow, 1’-13’ long, glabrous or some- 
what pubescent, the tube narrow, slightly expanded 
above, the limb short and nearly regular; stamens 
and style scarcely exserted; berries scarlet, about 


47 


3” in diameter. 

In low grounds, or on hillsides, Maine to Florida, 
New Hampshire, New York, Nebraska and Texas. 
Woodbine (N. C.). Scarlet trumpet-honeysuckle. 
April-S ept. 


8. Lonicera japonica Thunb. Japanese 
or Chinese Honeysuckle. Fig. 3986. 


Lonicera japonica Thunb. FI. Jap. 89. 1784 


Pubescent, climbing high or trailing. Leaves 
all short-petioled, ovate, entire, 1-3’ long, 
acute at the apex, rounded at the base, dark 
green and glabrous above, pale and usually 
sparingly pubescent beneath; flowers in pairs 
from the upper axils, peduncled, leafy-bracted 
at the base, white or pink, fading to yellow, 
pubescent without, the tube nearly 1’ long, 
longer than the strongly 2-lipped limb; sta- 
mens and style exserted; berries black, 3-4” 
in diameter. 


Freely escaped from cultivation, Connecticut, 
New York and Pennsylvania to North Carolina, 
Florida and West Virginia. Naturalized from 
eastern Asia. June—Aug. 


ee 


es |. 


5 
% 


GENUs 6. 


g. Lonicera coertlea L. Blue or Moun- 
tain Fly-Honeysuckle. Fig. 3987. 


Lonicera coerulea L Sp. Pl. 174. 1753. 
Lonicera villosa Muhl. Cat. 23. 1813. 


Erect, shrubby, 1°-3° high, the twigs some- 
times slightly pubescent. Leaves oval or 
obovate, 1-12’ long, very obtuse at the apex, 
rounded or narrowed at the base, thick, con- 
spicuously reticulate-veined, pale and more or 
less pubescent beneath, glabrous above, at least 
when mature, ciliate on the margins; flowers 
in pairs in the axils, short-peduncled, subulate- 
bracted, yellow, 6-8” long; corolla pubescent, 
or glabrate, the tube gibbous at the base, the 
limb nearly regular, its lobes oblong, equalling 
or slightly exceeding the tube; ovaries of the 
two flowers becoming united and forming an 
oblong or nearly globose, bluish-black 2-eyed 
berry, about 22” in diameter. 

In low grounds, Newfoundland to Alaska, south 
to Rhode Island, Pennsylvania, Wisconsin, Wyo- 
ming and California. Europe and Asia. June. 


Lonicera Morrowi A. Gray, a Japanese species, 
with red berries and nearly white flowers, has 
escaped from cultivation in Massachusetts. 

fi 


Ir. Lonicera canadénsis Marsh. Amer- 
ican Fly-Honeysuckle. Fig. 3989. 
Vaccinium album L. Sp. Pl. 350. 1753. Not L. 

alba L. 

Lonicera canadensis Marsh. Arb. 81. 1785. 
Lonicera ciliata Muhl. Cat. 23. 1813. 

Shrubby, 3°-5° high, the twigs glabrous. 
Petioles 2’-3” long, very slender; leaves thin, 
1-4’ long, bright green on both sides, ovate or 
sometimes oval, acute or acutish at the apex, 
rounded or cordate at the base, villous- 
pubescent beneath when young, glabrous or 
nearly so when mature, but the margins 
strongly ciliate; flowers in pairs from the 
axils, greenish-yellow, about 8’ long; pedun- 
cles long-filiform; bracts very small, subulate; 
corolla-limb nearly regular, its lobes short; 
berries separate, ovoid, light red, about 3” thick. 
In moist woods, Nova Scotia to Saskatchewan, 


Connecticut, Pennsylvania, Indiana, Michigan and 
Minnesota. Medaddy-bush. May. 


HONEYSUCKLE FAMILY. 281 


to. Lonicera oblongifolia (Goldie) 
Hook. Swamp Fly-Honeysuckle. 
Fig. 3988. 
Xylosteum oblongifolium Goldie, Edinb. 

Phil. Journ. 6: 323. 1822. 

Lonicera oblongifolia Hook. Fl. Bor. Am. 

I: 284. pl. Too. 1833. 

L. altissima Jennings, Ann. Carn. Mus. 4: 

74. pl. 20. 1906. 

Leaves 1’-2’ long, reticulate-veined, 
glabrous or nearly so on both sides when 
mature, downy-pubescent when young, 
not ciliate; flowers in pairs on long 
slender peduncles; corolla yellow, or 
purplish within, 8’—-9” long, gibbous at 
the base, deeply 2-lipped; bracts minute 
or none; ovaries remaining distinct, or 
becoming united, the berries red or 
crimson. 

In swamps, New Brunswick and Quebec 
to Manitoba, New York, Pennsylvania, 
Michigan and Minnesota. May—June. 


282 CAPRIFOLIACEAE. 


pal 


gla 


as 
ish 


Ma 


13. Lonicera tatarica L. Tartarian 
Bush-Honeysuckle. Fig. 3991. 


Lonicera tatarica L. Sp. Pl. 173. 1753. 


A glabrous shrub, 5°-10° high: Leaves 
ovate, rather thin, not conspicuously reticulate- 
veined, 1-3’ long, acute or obtusish at the apex, 
cordate at the base, not ciliate; flowers in 
pairs on slender axillary peduncles; corolla 
pink to white, 7”-8” long, the tube gibbous at 
the base, the limb irregularly and deeply 
5-lobed, somewhat 2-lipped; peduncles 1’ long; 
bracts linear, sometimes as long as the corolla- 
tube; stamens and style scarcely exserted; 
berries separate, red. 

Escaped from cultivation, Ontario, Maine and 
Vermont to southern New York, New Jersey and 


Kentucky. May. Native of Asia. Garden fly- 
honeysuckle. 


Vo. III. 


12. Lonicera Xylésteum L. Fly-Honey- 


suckle. 


Fig. 3990. 


Lonicera Xylosteum L. Sp. Pl. 174. 1753. 

A shrub, 3°-7° high, the foliage densely ap- 
pressed-pubescent when young. 
oval, or obovate, entire, short-petioled, rather 


Leaves ovate, 


e green, obtuse, or the upper acute at the 


apex, obtuse, subcordate or narrowed at the base, 


brous above when mature, persistently pubes- 


cent beneath, 17-3’ long; petioles 2”-4” long; 
peduncles axillary, 2-flowered, 4’-8” long, about 


long as the. flowers, or longer; flowers yellow- 
white; bracts linear-subulate; berries scarlet. 


Escaped from cultivation, Rhode Island, New 
York and New Jersey. Native of Europe and Asia. 


y—June. 


14. Lonicera involucrata (Richards. ) 
Banks. Involucred Fly-Honeysuckle. 
, Fig. 3992. 


Xylosieum involucratum Richards. App. Frank. 
Journ Edi 2165 s1s23. 

Lonicera involucrata Banks; Richards. loc. cit. 
1823. 


Distegia involucrata Rydb. Bull. Torr. Club 33: 


152. 1906. 


A glabrate or pubescent shrub, 3°-10° high. 
Leaves short-petioled, ovate, oval, or obovate, 
2’-6’ long, acute or acuminate at the apex, 
narrowed or rounded at the base, more or less 
pubescent, at least when young; peduncles axil- 
lary, 1-2’ long, 2-3-flowered; bracts foliaceous, 
ovate or oval, often cordate; bractlets also 
large, at length surrounding the fruit; flowers 
yellow; corolla pubescent, funnelform, the 
limb nearly equally 5-lobed; lobes short, little 
spreading ; stamens and style slightly exserted ; 
berries separate, globose, or oval, nearly black, 
about 4” in diameter. 


In woodlands, New Brunswick and Quebec to western Ontario and Michigan, west to British 


Columbia and Alaska, south to Arizona, Utah and 


California. June-July. 


GENUS 7. ' HONEYSUCKLE FAMILY. 283 


7. DIERVILLA [Tourn.] Mill. Gard. Dict. Abr. Ed. 8. 1754. 


Shrubs, with opposite leaves, and yellow axillary and terminal cymose or solitary flowers. 
Calyx-tube slender, elongated, narrowed below, the limb with 5 linear persistent lobes. Corolla 
narrowly funnelform, the tube slightly gibbous at the base, the limb nearly regular, 5-lobed. 
Stamens 5, inserted on the corolla; anthers linear. Ovary 2-celled; ovules numerous in each 
cavity; style filiform; stigma capitate. Fruit a linear-oblong capsule, narrowed or beaked at 
the summit, septicidally 2-valved, many-seeded. Seed coat minutely reticulated; endosperm 
fleshy; embryo minute. [Named for Dr. Dierville, who brought the plant to Tournefort.] 


Three species, the following typical one, the others in the mountains of the Southern States. 
’ The Japanese and Chinese Weigelas, often referred to this genus, are here regarded as distinct. 


1. Diervilla Diervilla (L.) MacM. Bush- 
Honeysuckle. Fig. 3993. 
Lonicera Diervilla L. Sp. Pl. 175. 1753. 
Diervilla Lonicera Mill. Gard. Dict. Ed. 8. 1768. 
Diervilla trifida Moench, Meth. 492. 1794. 
D. Diervilla MacM. Bull. Torr. Club 19: 15. 1892. 
A shrub, 2°-4° high, glabrous or nearly so 
throughout, with terete branches. Leaves ‘short- 
petioled, ovate or oval, acuminate at the apex, 
usually rounded at the base, 2-5’ long, irregularly 
crenulate and often slightly ciliate on the margins; 
peduncles terminal, or in the upper axils, slender, 
1-5-flowered; flowers about 9” long; corolla more 
or less pubescent both without and within, regular 
or slightly irregular, 3 of its lobes somewhat 
united; capsule glabrous, linear-oblong, slender, 
beaked, crowned with the persistent calyx-lobes. 
In dry or rocky woodlands, Newfoundland to Mani- 


toba, North Carolina, Michigan and Wisconsin. 
Gravel-weed. Life-of-man. May-June. 


Family 38. ADOXACEAE Fritsch; Engl. & Prantl, Nat. Pfl. Fam. 4*: 170. 1891. 
MoscHATEL FAMILY. 

A glabrous perennial slender herb, with scaly or tuberiferous rootstocks, basal and 
opposite ternately compound leaves, and small green flowers in terminal capitate 
clusters. Calyx-tube hemispheric, adnate to the ovary, its limb 2-3-toothed. Corolla 
rotate, regular, 4-6-lobed. Stamens twice as. many as the lobes of the corolla, 
inserted in pairs on its tube; filaments short; anthers peltate, 1-celled. Ovary 
3-5-celled; style 3-5-parted; ovules I in each cavity, pendulous. Fruit a small 
drupe with 3-5-nutlets. Endosperm carti- 
laginous. 


The family contains only the following monotypic 
genus of the north temperate zone. 


1. ADOXA L. Sp. Pl. 367. 1753. 


_ Characters of the family. [Greek, without glory, 
1. €., insignificant. | 


t. Adoxa Moschatéllina L. Musk-root. 
Hollow-root. Moschatel. Fig. 3994. 
Adoxa Moschatellina L. Sp. Pl. 367. 1753. 


Stems simple, weak, erect, 3-6’ high, bearing 
a pair of opposite ternate leaves usually above the 
middle. Basal leaves 1-4, long-petioled, ternately 
compound, the segments broadly ovate or orbicu- 
lar, obtuse, thin, 3-cleft or 3-parted, the lobes 
obtuse and mucronulate; head 3-4” in diameter, 
composed of 3-6-flowers; corolla of the terminal 
flower 4-5-lobed, those of the others usually 
5-6-lobed; drupe green, bearing the persistent 

calyx-lobes above the middle. 

‘In shaded rocky places, Arctic America, south to 
Iowa, Wisconsin, South Dakota and Colorado. Also 
in northern Europe and Asia. Other English names © 
are bulbous fumitory, glory-less, musk-crowfoot or 
wood-crowfoot. Odor musky. May. 


254 VALERIANACEAE. Vot. III. 


Family 39. VALERIANACEAE Batsch, Tabl. Aff. 227. 1802. 
VALERIAN FAMILY. 


Herbs with opposite leaves, no stipules, and usually small perfect or polygamo- 
dioecious flowers, in corymbed panicled or capitate cymes. Calyx-tube adnate to 
the ovary, its limb inconspicuous or none in flower, often becoming prominent in 
fruit. Corolla gamopetalous, epigynous, somewhat irregular, its tube narrowed, 
and sometimes gibbous or spurred at the base, its limb spreading, mostly 5-lobed. 
Stamens I—4, inserted on the corolla and alternate with its lobes, usually exserted. 
Ovary inferior, I-3-celled, one of the cavities containing a single anatropous ovule, 
the others empty. Fruit indehiscent, dry, containing a single suspended seed. 
Endosperm little or none; embryo straight; cotyledons oblong. 


About 9 genera and 300 species, of wide distribution, most abundant in the northern hemisphere. 


Fruit 1-celled; persistent calyx-lobes becoming awn-like; tall herbs. 
é F 1. Valeriana. 
Fruit 3-celled; calyx-lobes minute or none; low herbs. 
2. Valerianella. 


1. VALERIANA [Tourn.] L. Sp. Pl. 31. 1753. 


Perennial, strong-smelling, mostly tall herbs, the leaves mainly basal and the cymose 
flowers paniculate in our species. Calyx-limb of 5-15 bristle-like plumose teeth, short and 
inrolled in flower, but elongated, rolled outward and conspicuous in fruit. Corolla funnel- 
form or tubular, usually more or less gibbous at the base, the limb nearly equally 5-lobed. 
Stamens commonly 3. Style entire, or minutely 2-3-lobed at the summit. Fruit compressed, 
t-celled, 1-nerved on the back, 3-nerved on the front. [Name Middle Latin, from valere, 
to be strong. ] 

About 175 species, mostly in the temperate and colder parts of the north temperate zone and 


the Andes of South America. Besides the following, 5 others occur in southern and western North 
America. Type species: Valeriana pyrendaica L. 


‘ 


Corolla-tube very slender, 6”’—10” long; basal leaves cordate. 1. V. pauciflora. 
Corolla-tube 1”—3” long; basal leaves not cordate. 
Leaves thick, parallel-veined, entire, or the segments not dentate; roots fusiform. 
2. V. edulis. 
Leaves thin, reticulate-veined, the segments dentate; roots fibrous. 
Lower leaves spatulate, often entire; plants glabrous. 
Segments of middle stem-leaves 9-13, usually sinuate-dentate; corolla 1.7”—2.5” wide. 
: 3. V.uliginosa, 
Segments of middle stem-leaves 5-7, entire; corolla 1.5” wide orless. 4. V.septentrionalis. 
All the leaves pinnately divided ; plants pubescent, especially at the nodes. 
s. V. officinalis. 


1. Valeriana pauciflora Michx. Large-flowered Valerian. Fig. 3995. 
V. pauciflora Michx. Fl. Bor. Am. 1: 18. 1803. 


Rootstocks slender, usually horizontal. 
Stem glabrous, erect or ascending, 1°-3° 
high, often sending out runners from the 
base; leaves thin, the basal ones slender- 
petioled, simple, or sometimes with a pair 
of small leaflets on the petiole, broadly 
ovate, cordate, acute at the apex, the mar- 
gins crenate or dentate; stem leaves pin- 
nately 3-7-divided, the terminal segment 
larger than the others; cymes terminal, 
clustered; flowers few or numerous; co- 
rolla pink, its tube very slender, 6-10” 
long; bracts linear; fruit oblong or oblong- 
lanceolate, about 3” long, glabrous or 
puberulent; bristles of the calyx at length 
elongated and plumose. 

In moist soil, Pennsylvania to West Vir- 


ginia, Illinois, Kentucky, Tennessee and Mis- 
souri. American wild valerian. May—June. 


GENUS I. VALERIAN FAMILY. 285 


2. Valeriana édulis Nutt. Edible Valer- 
ian. Tobacco-root. Fig. 3996. 
V. edulis Nutt. in T. & G. Fl. N. A. 2: 48. 1841. 

Erect, 1°-4° high, from a deep fusiform 
carrot-shaped root. Stem glabrous, or nearly 
so, the young leaves commonly more or less 
pubescent and the older ones finely ciliate, 
sometimes glabrous; basal leaves spatulate or 
oblanceolate, thick, 312’ long, 2’—10” wide, 
obtuse at the apex, narrowed into a margined 
petiole, parallel-veined, entire or with a few 
obtuse entire lobes; stem leaves few, sessile, 
pinnately-parted into linear or lanceolate seg- 
ments; flowers yellowish-white, small (2’), 
polygamo-dioecious, paniculate, the inflores- 
cence at length widely branching; bracts 

-lanceolate, short; fruit narrowly ovate, gla- 
brous or nearly so, 2” long, at length exceeded 
by the plumose calyx-teeth. 

In wet open places, Ontario to British Colum- 
bia, south to Ohio, Iowa, Wisconsin, and in the 
Rocky Mountains to Arizona and New Mexico. 
Called also oregon tobacco; the root cooked for 
food. May—Aug. 3. Valeriana uligindsa (T. & G.) Rydb. 

Marsh or Swamp Valerian. Fig 3997. 
Valeriana dioica Pursh, Fl. Am. Sept. 727. 1814. 

Iie IE. GAY 
V. sylvatica uliginosa T.& G. Fl. N. A.2: 47. 1841. 
V. uliginosa Rydb.; Britton, Man. 878. too. 

Erect, glabrous or very nearly so through- 
out, 8~-23° high. Rootstocks creeping or 
ascending; basal leaves thin, petioled, oblong 
or spatulate, obtuse, entire, or with a few 
obtuse lobes, reticulate-veined, 2’-10’ long, 
3-18 wide; stem leaves 2-4 pairs, petioled, 
pinnately parted into 3-15 ovate to lanceolate. 
dentate or entire, acute or obtuse segments; in- 
florescence cymose-paniculate, at length loosely 
branched; flowers pink or nearly white, 3-4” 
long, about 2” wide; bracts linear-lanceolate; 
fruit ovate, glabrous, 12” long. 

In wet soil, Quebec to New York, Ontario and 
Michigan. American wild valerian. Referred, in 
our first edition, to the following northern and 
western species. May—Aug. 


4. Valeriana septentrionalis Rydb. North- 
ern Valerian. Fig. 3908. 


Valeriana sylvatica Banks; Richards. App. Frank. 
Journ. Ed. 2, 2. 1823. Not F. W. Schmidt. 


Valeriana septentrionalis Rydb. Mem. N. Y. Bot. 
Gard. 1: 376. 1900. 


Erect, 8-16’ high, glabrous, or the inflores- 
cence minutely pubescent. Basal leaves spatu- 
late or oval, 4’ long or less, entire; stem leaves 
usually 3 pairs, the segments 5-7, oval to linear- 
lanceolate, entire, or wundulate-margined; in- 
florescence cymose-paniculate, dense; flowers 
white, about 14” wide; fruit 14-2” long, gla- 
brous. 


In wet soil, Newfoundland to British Columbia, 
south in the Rocky Mountains to New Mexico. 
Summer. 


VALERIANACEAE. Vor. IIT. 


nine 
rere 5. Valeriana officinalis L. Common, 
pene Garden or Great Wild Valerian. 
t Vandal-root. Fig. 3999. 


Valeriana officinalis L. Sp. Pl. 31. 1753. 


Erect, 2°-5° high, more or less pubes- 
cent, especially at the nodes. Leaves all 
pinnately parted into 7-25 thin reticulate- 
veined lanceolate acute or acuminate seg- 
ments, sharply dentate, or those of the 
upper leaves entire, usually with some 
scattered hairs beneath; flowers pink or 
nearly white, about 2” long; inflorescence 
of several rather compact corymbed cymes; 
bracts linear-lanceolate, rather large; fruit 
glabrous, ovate, about 12” long. 


Escaped from gardens to roadsides in New 
York, Ohio and New Jersey. Native of Eu- 
rope and Asia. Old names,’ cats’-valerian, 
setwell, cut-heal, all-heal. Garden-, summer- 
ce hardy-heliotrope. St. George’s-herb. June— 

ug. 


2. VALERIANELLA [Tourn.] Mill. Gard. Dict. Abr. Ed. 4. 1754. 


Annual dichotomously branched herbs, the basal leaves tufted, entire, those of the stem 
sessile, often dentate, the flowers in terminal, compact or capitate, in our species corymbed 
or panicled cymes. Corolla small, white, blue, or pink, nearlys regular. Calyx-limb short 
or obsolete in flower, in fruit various, not divided into filiform plumose segments, often none. 
Corolla-tube narrowed at the base, the limb spreading, 5-lobed. Stamens 3; style minutely 
3-lobed at the summit. Fruit 3-celled, 2 of the cells empty, and in our species about as large 
as the fertile one. [Name a diminutive of Valerian.] 

About 50 species, natives of the northern hemisphere, most abundant in the Mediterranean 
region. Besides the following, 8 others occur in the western parts of North America. Type species: 
Valeriana Locusta L. 

Corolla funnelform, the short tube not longer than the limb or about equalling it. 


Fruit flattened, twice as broad as thick; corolla blue. 1. V. Locusta. 
Fruit triangular-pyramidal; corolla white. 2. V.chenopodifolia. 
Fruit oblong-tetragonal or ovoid-tetragonal, grooved. 
Groove of the fruit broad and shallow. . V. radiata. 
V. stenocarpa. 


. V.Woodsiana. 


Fruit globose or saucer-shaped. 
. V.longiflora. 


3 
Groove of the fruit narrow. 4. 
d 5 
Corolla salverform, purplish, the slender tube much longer than the limb. 6 


1. Valerianella Locista (L.) Bettke. European Corn Salad. Fig. 4000. 


Valeriana Locusta and yar. olitoria L. Sp. Pl. 


se yV) YMA) 0) 

33- 1753- ot), Yon er 

Valerianella olitoria Poll. Hist. Pl. Palat. 1: Ky RY ery 
30. - 1776. on 


Valerianella Locusta Bettke. Anim. Val. tro. 
1826. 


Glabrous, or pubescent at the nodes, 6 
12’ high, usually branched from the base 
and repeatedly forked. Basal leaves spatu- 
late or oblanceolate, rounded and obtuse 
at the apex, 1}’-2’ long, 3-5” wide, entire; 
upper stem leaves oblong-lanceolate, usu- 
ally dentate; peduncles short; cymes 3-6” 
broad, almost capitate; bracts linear or 
linear-oblong; corolla blue, about 1” long; 
fruit flattened, rounded on the edges, 1” 
long, glabrous, twice as broad as thick, 
depressed-orbicular in outline, the two 
empty cavities smaller than the fertile one, 
which has a corky mass at its back. 

In fields and waste places, Maine to On- 
tario, Idaho, Arkansas, New Jersey, Pennsyl- 


vania, Virginia and Louisiana. Naturalized ie 
from Europe. The plant is cultivated and the leaves used for salad under the name of fétticus. 


White pot-herb, lamb’s-lettuce, milk-grass. April-July. 


i ae ee 


GENUS 2. VALERIAN FAMILY. 237 


2. Valerianella chenopodif6lia (Pursh) DC. Goose-foot Corn Salad. Fig. 4oot. 


Fedia chenopodifolia Pursh, Fl. Am. Sept. 727. EN) \e@ 
1814. A i) \ = 


V. chenopodifolia DC. Prodr. 4: 629. 1830. 
Fedia Fagopyrum T. & G. Fl. N. A. 2: 51. 1841. 


Glabrous, 1°-2° high. Leaves entire, or the 
basal and lower ones repand, spatulate, ob- 
tuse; upper stem leaves oblong or lanceolate, 
1-3’ long; cymes dense, 6’-8” broad, at length | 
slender-peduncled; bracts lanceolate or oblong- | 
lanceolate; corolla white, about 1” long; fruit SF AA 
triangular-pyramidal, 2” long, 1” thick, gla- | 
brous or minutely pubescent, the two empty 


cavities narrower than the fertile one but about > 3 
as deep. : es yy 5 

In moist soil, western New York to Virginia, Ab ao SQ 

yj nf ~: 
Me oe ER oes ie yj g Al 
ae Bi Za RY YW WES : ; 
3. Valerianella radiata (L.) Dufr. Beaked 
Corn Salad. Fig. 4002. 

Valeriana Locusta var. radiata L. Sp. Pl. 34. 1753. 
Fedia radiata Michx. Fl. Bor. Am. 1: 118. 1803. 
Valerianella radiata Dufr. Hist. Val. 57. 1811. 

Glabrous, or minutely pubescent below, 6-18’ 
high. Basal and lower leaves spatulate, obtuse, 
entire, the upper lanceolate, usually dentate; 
cymes 4-6” broad, dense; bracts small, lanceo- 
late or oblong-lanceolate ; corolla white, 1” long; 
fruit narrowly ovate-tetragonal, finely pubescent 
or sometimes glabrous, 1” long, 4” thick, the 
empty cavities as thick as or thicker than the 
beaked fertile one and separated from each other 
by a broad shallow groove. 

In moist soil, Massachusetts to Florida, west to 


Minnesota, Michigan, Missouri and Texas. Called 
also lamb’s-lettuce. May-July. 


Siam, aioe 
Phy, Np ee a oe 
: N Wes ee =, 7 
4. Valerianella stenocarpa ( Engelm. ) BY) \ 4. 
Krok. Narrow-celled Corn Salad. SY 


Fig. 4003. 
bee 
Fedia stenocarpa Engelm. Bost. Journ. Nat. W y 


Hist. 6: 216. 1857. IW 
Valerianella stenocarpa Krok, Kongl. Svensk. 
Akad. Handl. 5: 64. 1866. 
Similar to the preceding species and 
perhaps better regarded as a race of it. AY 
Wak 


Fruit oblong-tetragonal, slightly smaller, iors 
glabrous or sometimes pubescent; sterile S 
cavities not as thick as the oblong seed- 

bearing one, and separated from each aN 


other by a narrow groove. 


Prairies and woodlands, Kansas and Mis- 
souri to Texas. March—June. 


288 VALERIANACEAE. Vou. ILL. 


5. Valerianella Woodsiana (T.&G.) Walp. 
Woods’ Corn Salad. Fig. 4004. 

Fedia Woodsiana T.& G. Fl. N. A. 2: 52. 1841. 
Valerianella Woodsiana Walp. Rep. 2: 527. 1843. . 
F. umbilicata Sulliv. Am. Journ. Sci. 42: 50. 1842. 
Fedia patallaria Sulliv.; A. Gray, Man. 183. 1848. 

Usually larger than any of the preceding spe- 
cies, sometimes 3° high, glabrous or very nearly 
so throughout. Basal and lower leaves spatulate, 
obtuse, entire; upper leaves lanceolate or linear- 
oblong, usually dentate; cymes 3-6” broad, few- 
flowered; bracts comparatively large, lancéolate; 
corolla white, about 1” long; fruit glabrous, 
nearly globular to saucer-shaped, about 1” in 
diameter, the empty cavities inflated, introrse with 
a depression or concavity between them, as broad 
as the fertile one. 


In moist soil, New York, Pennsylvania and Ohio 
to Tennessee and Texas. May-July. 


6. Valerianella longifldra (T. & G.) 
Walp. Long-flowered Corn Salad. 
Fig. 4005. 

Fedia longifiora T. & G. FI. N. A. 2:51. 1841. 
Valerianella longiflora Walp. Rep. 2: 527. 1843. 
Glabrous; stem usually several times forked, 
6’-12’ high. Leaves very obtuse, the basal ones 
spatulate, 1-23’ long, 4”-8” wide, those of the 
stem oblong or spatulate-oblong, smaller, some- 
what clasping; cymes dense, corymbed, com- 
monly numerous, several-many-flowered; co- 
rolla salverform, pink or purplish, about 6” 
long, the almost filiform tube 3-4 times as long 
as the somewhat irregular 5-parted limb, the 
lobes linear-oblong; bracts with small gland- 
tipped teeth; fruit broadly ovate or nearly 
orbicular in outline, the empty cavities diver- 
gent, larger than the oblong seed-bearing one. 


In moist rocky situations, Missouri and Arkan- 
sas. April-May. 


Family 40. DIPSACACEAE Lindl. Veg. Kingd. 699. 1847. 
TEASEL FAMILY. 

Perennial biennial or annual herbs, with opposite or rarely verticillate leaves, 
and perfect gamopetalous flowers in dense involucrate heads. Stipules none. 
Flowers borne on an elongated or globose receptacle, bracted and involucellate. 
Calyx-tube adnate to the ovary, its limb cup-shaped, disk-shaped, or divided into 
spreading bristles. Corolla epigynous, the tube usually enlarged at the throat, the 
limb 2—-5-lobed. Stamens 2-4, inserted on the tube of the corolla and alternate 
with its lobes; filaments distinct ; anthers versatile, longitudinally dehiscent. Ovary 
inferior, 1-celled, style filiform; stigma undivided, terminal, or oblique and lateral ; 
ovule 1, anatropous. Fruit an achene, its apex crowned with the persistent calyx- 
lobes. Seed-coat membranous; endosperm fleshy; embryo straight. 

About 7 genera and 140 species, natives of the Old World. 


Scales of the elongated receptacle prickly pointed. 1. Dipsacus. 
Scales of the receptacle not prickly, herbaceous, capillary, or none. 
No receptacular scales. 2. Scabiosa. 
Scales of the receptacle about as long as the flowers. 3. Succisa. 


1. DIPSACUS [Tourn.] L. Sp. Pl.97. 1753. 


Rough-hairy or prickly tall erect biennial or perennial herbs, with opposite dentate entire 
or pinnatifid, usually large leaves, and blue or lilac flowers in dense terminal peduncled oblong 
heads in our species. Bracts of the involucre and scales of the receptacle rigid or spiny 
pointed. Involucels 4-8-ribbed with a somewhat spreading border. Limb of the calyx cup- 


GENUS I. TEASEL FAMILY. 289 


shaped, 4-toothed or 4-lobed. Corolla oblique or 2-lipped, 4-lobed. Stamens 4. Stigma 
oblique or lateral. Achene free from or adnate to the involucel. [Greek, to thirst, the 
leaves of some species holding water. ] 

About 15 species, natives of the Old World. Type species: Dipsacus fullonum L. 


Scales of the receptacle straight-pointed. 1. D. sylvestris. 
Scales of the receptacle hooked at the apex. , 2. D. fullonum. 


Dipsacus fullonum L. Sp. Pl. 97. In part. 1753. 
Dipsasus sylvestris Huds. Fl. Angl. 49. 1762. 
Biennial, stout, with numerous short prickles on 
the stem, branches, peduncles, midribs: of the leaves 
and involucre, otherwise glabrous or nearly so, 
3°-6° high. Leaves sessile, or the upper ones con- 
nate-perfoliate, lanceolate or oblong, the upper 
acuminate and generally entire, the lower obtuse 
or obtusish, crenate or sometimes pinnatifid at 
the base, often 1° long; heads at first ovoid, be- 
coming cylindric, at length 3-4’ long; flowers lilac, 
4-6” long; leaves of the involucre linear, curved 
upward, as long as the head or longer; scales of 
the receptacle ovate, tipped with a long straight 
subulate barbed awn, usually exceeding the flowers. 
In waste places, Maine and Ontario to North Caro- 
lina, west to Michigan. Naturalized from Europe and 
native also of Asia. July-Sept. Other English names 
are Venus-bath or -cup; wood- or church-brooms; 
shepherds’-staff ; card- or water-thistle; gipsy-combs; 


hutton-weed. Indian’s-thistle. Prickly-back. Adam’s- 
flannel. 


2. Dipsacus fullonum L. Fuller’s or 
Draper's Weasel. Fuller's, Vhistle. 


Fig. 4007. 


Dipsacus fullonum L. Sp. Pl. 97. 1753. 
Dipsacus fullonum var. sativus L. Sp. Pl. Ed. 2, 1677. 

1763. 

Similar to the preceding species. Leaves of the 
involucre, or some of them, shorter than the heads, 
spreading or at length reflexed; scales of the 
receptacle with hooked tips, about equalling the 
flowers, which are usually paler than in D. syl- 
vestris. 

About wool mills, Eastern and Middle States, rare. 
Fugitive from Europe, and perhaps nowhere per- 
manently established within our range. Other Eng- 
lish names are clothiers’ brush, Venus’-bath or -cup. 
Generally regarded as probably a cultivated variety 


of the preceding species, as it is not found wild, ex- 
cept as an evident escape. 


Dipsacus laciniatus L., with pinnatifid or bipinnatifid ciliate leaves, those of the involucre 
spreading, has been found at Albany, New York. Fugitive or adventive from Europe. 


2. SCABIOSA [Tourn.] L. Sp. Pl. 98 1753. 


Herbs, with opposite leaves, no prickles, and blue pink or white flowers in peduncled 
involucrate heads. Bracts of the involucre herbaceous, separate, or slightly united at the 
base. Scales of the pubescent receptacle none. Involucels compressed, the margins often 
minutely 4-toothed. Calyx-limb 5-10-awned. Limb of the corolla 4-5-cleft, oblique or 2-lipped. 
Stamens 4 (rarely 2). Stigma oblique or lateral. Achene more or less adnate to the invo- 
lucel, crowned with the persistent calyx. [Latin, scale, from its repute as a remedy for scaly 
eruptions | 

About 75 species, natives of the Old World. Type species: Scabiosa arvensis L. 


19 


tw 


go DIPSACEAE, Vor. ITI. 


I. Scabiosa arvénsis L. Field Scabious. 
Fig. 4008. 


Scabiosa arvensis L. Sp. Pl. 99. 1753. 
Knautia arvensis T. Coult. Dips. 29. 1823. 


Perennial, pubescent, simple or little branched, 
1°-3° high. Basal and lower leaves petioled, lan- 
seolate, acute or acuminate, entire, lobed, or pin- 
natifid, 3-8’ long; upper leaves sessile, often 
deeply pinnatifid; heads long-peduncled, depressed- 
globose, 1’-12’ broad; flowers lilac purple, about 
6” long; receptacle depressed-hemispheric, not 
scaly, covered with hairs between the flowers; 
achene angled, crowned with the 8 or Io linear- 
subulate calyx-teeth. 

In cultivated fields and waste places, Quebec to 
Massachusetts, Vermont, New York and Pennsyl- 
vania. Adventive from Europe. Other English 


names are blue buttons, blue caps, gypsy- or egyptian- 
rose, pincushion. June—Sept. 


3. SUCCISA (Vaill.) Moench. Meth. 488. 1794. 


Herbs, similar to Scabiosa, with opposite leaves, the flowers in long-peduncled heads, 
subtended by a several-many-leaved involucre. Scales of the receptacle herbaceous or capil- 
lary. Involucels grooved, the margins 4-lobed or 4-toothed. Calyx-limb 5-toothed or 5-awned. 
Corolla oblique, 4-5-lobed. Stamens 4. Achene crowned with the persistent calyx. [From 
the Latin, to bite off, the rootstock in some species being short and blunt. ] 


About 4 species, mostly natives of southern Europe. Type species Scabiosa Succisa L. 


1. Succisa australis (Wulf.) Reichenb. Southern Scabious. Fig. 4009. 


Scabiosa australis Wulf. in Roem. Arch. 3: Part 3, 

316. 1803. 

Succisa australis Reichenb. Fl. Germ. Excurs. 196. 

1830. 

Perennial, puberulent or pubescent, at least 
above ; stem slender, branched, 12°-3° high. Basal 
leaves oblanceolate to oblong, mostly obtuse, 4-12’ 
long, the petiole often as long as the blade or 
longer; stem leaves distant, lanceolate or oblong- 
lanceolate, entire, or toothed, short-petioled, or 
the upper sessile, acute or acuminate; heads of 
pale blue-purple flowers long-peduncled, rather 
less than 1’ in diameter, oblong-ovoid in fruit; 
receptacle scaly, the scales about as long as the 
glabrous, 8-ribbed involucels or longer; calyx 5- 
toothed; achene crowned with 5 calyx-teeth. 

Fields and meadows. Naturalized from Europe in 


Pennsylvania, central New York and Massachusetts. 
Pincushion-flower. Summer. 


Succisa Succisa (L.) Britton (Succisa pratensis 
Moench; Scabiosa Succisa L.) with villous 4-angled 
involucels, the calyx-limb 5-awned, has been found in fields at Louisburg, Cape Breton Island. 


Family 41. CUCURBITACEAE B. Juss. Hort. Triany a7 5e: 
GourD FaMIty. 


Climbing or trailing, herbaceous vines, usually with tendrils. Leaves alternate, 
petioled, generally palmately lobed or dissected. Flowers solitary or racemose, 
monoecious or dioecious. Calyx-tube adnate to the ovary, its limb campanulate 
or tubular, usually 5-lobed, the lobes imbricated. Petals usually 5, inserted on 
the limb of the calyx, separate, or united into a gamopetalous corolla. Stamens 
mostly 3 (sometimes 1), 2 of them with 2-celled anthers, the other with a 1-celled 
anther ; filaments short, often somewhat monadelphous. Ovary 1-3-celled; style 


GENUS I. GOURD FAMILY. 291 


terminal, simple or lobed; ovules few or numerous, anatropous. Fruit a pepo, 
indehiscent, or rarely dehiscent at the summit, or bursting irregularly; or some- 
times dry and membranous Seeds usually flat; endosperm none. 


About 90 genera and 700 species, mainly natives of tropical regions, a few in the temperate zones. 


Flowers large, yellow ; prostrate vine. : 1. Pepo. 
Flowers small, white or greenish ; climbing vines. 
Fruit glabrous; seeds numerous, horizontal. 2. Melothria. 


Fruit prickly ; seeds 1 or few, erect or pendulous. 
Fruit dehiscent at the apex or bursting irregularly ; several-seeded. 
Leaves 3-7-lobed ; anthers 3. 3. Micrampelis. 
Leaves digitately compound ; anther 1. 4. Cyclanthera. 
Fruit indehiscent, 1-seeded. 5. OICyOs. 


1. PEPO [Tourn.] Mill. Gard. Dict. Abr. Ed. 4. 1754. 


Rough prostrate vines, rooting at the nodes, with branched tendrils, usually lobed leaves 
mostly cordate at the base, and large yellow axillary monoecious flowers. Calyx-tube cam- 
panulate, usually 5-lobed. Corolla campanulate, 5-lobed to about the middle, the lobes recurv- 
ing. Staminate flowers with three stamens, the anthers linear, more or less united and no 
pistil. Pistillate flowers with 1 pistil; ovary oblong with 3-5 many-ovuled placentae; style 
short, thick; stigmas 3-5, each 2-lobed, papillose; staminodia 3. Fruit large, fleshy, with a 
thick rind, many-seeded, indehiscent. [From the Greek name of some large fruit.] 


About 10 species, natives of America, Asiaand Africa. Besides the following, some 6 others 
occur in the southern and southwestern United States. Type species: Cucurbita Pepo L. 


1. Pepo foetidissima (H.B.K.) Britton. Missouri Gourd. Calabazilla. Wild 
Pumpkin. Fig. 4010. 


Cucurbita foetidissima H.B.K. Nov. Gen. 2: 123. 
eee perennis James in Long’s Exp. 2: 20. 1823. 
Cucurbita perennis A. Gray, Bost. Journ. Nat. Hist. 

625793. “1850: 

Stem stout, rough, hirsute, trailing to a length 
of 15°-25°. Root large, carrot-shaped. Petioles 
stout, 3-8’ long, very rough; leaves ovate-trian- 
gular, thick and somewhat fleshy, cordate or trun- 
cate at the base, acute at the apex, 4-12’ long, 
usually slightly 3-5-lobed, denticulate, rough above, 
canescent beneath; peduncles 1-2’ long; flowers 
mostly solitary; corolla 23’-4’ long; pepo globose 
or globose-ovoid, 2’-3’ in diameter, smooth, its 
pulp fibrous and bitter. 


Dry soil, Missouri and Nebraska to Texas and 
Mexico, west to southern California. May-—Sept. 


Pumpkins, Squashes, Cucumbers and Melons, cul- 
tivated in. many races belong to this genus. Gourds 
belong to Cucurbita Lagenaria L., the type of the 
genus Cucurbita. 


Citrullus Citrillus (L.) Karst., the Watermelon, is 
found escaped from cultivation along river-shores in Virginia and West Virginia, and southward. 


2. MELOTHRIA L. Sp. Pl. 35. 1753. 


Slender, mostly climbing vines, with simple or rarely bifid tendrils, lobed or entire thin 
leaves, and small white or yellow monoecious flowers, the staminate clustered, the pistillate 
often solitary. Calyx campanulate, 5-toothed. Corolla campanulate, deeply 5-parted. Stamens 
3 in the staminate flowers, the anthers distinct or slightly united, the pistil wanting or rudi- 
mentary. Fertile flowers with 1 pistil; ovary ovoid, constricted below the corolla; placentae 
3; ovules numerous; style short; stigmas 3, linear. Fruit small, berry-like, pulpy, many- 
seeded. [From the Greek for some vine, probably Bryonia cretica.] 


About 70 species, natives of warm and tropical regions, most abundant in the Old World. 
Besides the following typical one, 3 or 4 others occur in the southern United States. 


CUCURBITACEAE. Vor. III. 


1. Melothria péndula L. Creeping Cucumber. 
Fig. 4011. 


Melothria pendula L. Sp. Pl. 35. 1753. 


Root perennial. Stem slender, climbing to a height of 
3°-5°, branched, glabrous, grooved; petioles 3’-23’ long; 
leaves nearly orbicular in outline, finely pubescent or sca- 
brous on both sides, cordate at the base, 5-lobed or 5-angled, 
denticulate or dentate; tendrils puberulent; staminate flow- 
ers 4-7, racemose, borne on a peduncle 34’-1’ long; fertile 
flowers solitary, slender-peduncled; corolla greenish white, 


about 2” broad; fruit smooth, ovoid, green, 4”-6” long. 


_In thickets, Pennsylvania (Schweinitz, according to Cog- 
niaux) ; Virginia to Florida, west to Indiana, Kentucky, Mis- 
souri and northern Mexico. June—Sept. 


3. MICRAMPELIS Raf. Med. Rep. (II.) 5: 350. 1808. 
[Ecuinocystis T: & G. Fl. N. A. 1: 542. 1840.] 

Mostly annual climbing herbs, with branched tendrils, lobed or angled leaves, and small 
white monoecious flowers. Calyx-tube campanulate, 5-6-lobed. Corolla very deeply 5-6-parted. 
Stamens 3 in the staminate flowers, the anthers more or less coherent. Pistillate flowers 
with a 2-celled ovary; oyules 2 in each cavity; style very short; stigma hemispheric or lobed. 
Fruit fleshy, or dry at maturity, densely spiny, 1-2-celled, usually with 2 seeds in each cavity, 
dehiscent at the summit. Testa of the seed roughened. [Greek, small-vine.] 


About 25 species, natives of America. Besides the following typical one, about ro others occur 
in the western United States. ; 


1. Micrampelis lobata (Michx.) Greene. Wild Balsam Apple. Mock Apple. 
Mock Orange. Fig. 4012. 

Momordica echinata Muhl. Trans. Am. Phil. Soc. 3: 

180. Name only. 1793. 
Sicyos lobata Michx. Fl. Bor. Am. 2: 217. 1803. 
Echinocystis lobata T. & G. Fl. N. A. 1: 542. 1840. 
Micrampelis ijobata Greene, Pittonia 2: 128. 1890. 

Stem nearly glabrous, angular and grooved, 
branching, climbing to a height of 15°-25°, 
sometimes villous-pubescent at the nodes. Peti- 
oles 1-3’ long; leaves thin, roughish on both 
sides, deeply cordate at the base, 3-7-lobed to 
about the middle, the lobes triangular-lanceolate, 
acute or acuminate, the margins remotely serru- 
late; staminate flowers very numerous in nar- 
row compound racemes; pistillate flowers soli- 
tary, or rarely 2 together; fruit ovoid, green, 
about 2’ long, armed with slender spines; seeds 
flat. ‘ 

Along rivers, and in waste places, New Brunswick 
to Ontario, Manitoba, Montana, Virginia, Pennsyl- 
vania, Kentucky, Kansas and Texas. Eastward, 


mostly occurring as an introduced plant. Wild 
cucumber. Creeper. Creeping Jenny. July—Sept. 


4. CYCLANTHERA Schrad. Ind. Sem. Hort. Goett. 1831. 


Climbing, annual or perennial, mostly glabrous vines, with forked or simple tendrils, 
usually digitately compound leaves and small white or greenish monoecious flowers. Calyx 
cup-shaped, 5-toothed. Corolla rotate, deeply 5-parted. Staminate flowers racemose or pani- 
cled, the stamens united into a central column; anther 1, annular in our species. Pistillate 
flowers solitary; ovary obliquely ovoid, beaked, 1-3-celled, with 2 ovules in each cavity; style 
short; stigma large, hemispheric. Fruit spiny, obliquely ovoid, beaked, at length irregularly 
dchiscent, few-seeded. [Greek, circle-anther.] 

About 40 species, natives of America. Type species: Cyclanthera peddta Schrad. 


GENUS 4. GOURD FAMILY. 293 


1. Cyclanthera dissécta (T.& G.) Arn. Cut- 
leaved Cyclanthera. Fig. 4013. 


Discanthera dissecta T. & G. Fl. N. A. 1: 697. 1840. 
Cyclanthera dissecta Arn. in Hook. Journ. Bot. 3: 280. 


1841. 


Annual; stem grooved and angular, glabrous, 
branching, climbing to a height of 3°-4°, or strag- 
gling. Petioles 1’-2’ long; leaves digitately 3-7- 
foliolate, the leaflets oval or oblong, usually acute at 
each end, 4-2’ long, rough on both sides, dentate, 
or somewhat lobed; staminate flowers racemose, 
borne on a peduncle 32’ long; pistillate flowers 
solitary, very short-peduncled; fruit narrowed at 
the base, slightly oblique, about 1’ long, armed 
with slender spines. 


Thickets, Kansas to Texas, Louisiana and northern 
Mexico. July—Sept. : 


5. SICYOS L. Sp. Pl. 1013. 1753. 


Annual climbing vines, with branched tendrils, angled or lobed leaves, and small white 
or green monoecious flowers. Calyx-tube campanulate or cup-shaped, 5-toothed. Corolla 
campanulate or rotate, 5-parted nearly to the base. Staminate flowers corymbose or race- 
mose, with 3 stamens, the filaments united into a short column, the anthers coherent; pistil 
wanting. Pistillate flowers several together in capitate long-peduncled clusters, with no 
stamens ; ovary oblong or fusiform, I-celled; ovule 1, pendulous; style short, slender; stigmas 
usually 3. Fruit spiny, indehiscent, 1-seeded. [Greek, a cucumber or gourd.] 

About 35 species, natives of America and Australasia. Besides the following typical species, 
2 others occur in the southwestern states. 

1. Sicyos angulatus L. One-seeded Bur- 
Cucumber. Star Cucumber. Fig. 4ora. 


Sicyos angulatus L. Sp. Pl. 1013. 1753. 


Stem angled, more or less viscid-pubescent, climb- 
ing to a height. of 15°-25°, or trailing. Petioles 
stout, 1-4’ long, pubescent; leaves nearly orbicular, 
rough on both sides, rather thin, deeply cordate at 
the base, 5-angled or 5-lobed, the lobes acute or 
acuminate, the margins denticulate; staminate flow- 
ers loosely corymbose or racemose, borne on elon- 
gated peduncles; fertile flowers capitate, their pe- 
duncles shorter; fruits sessile, 3-10 together, yellow- 
ish, about 3’ long, pubescent, armed with slender 
rough spines. 


— 


Along river banks and in moist places, Quebec and 
Ontario to Florida, west to South Dakota, Kansas and 
Texas. Naturalized in eastern Europe. Called also 
nimble kate, wild cucumber. Leaves sometimes 10’ across. 
Tune—Sept. 


Family 42. CAMPANULACEAE Juss. Gen. 163. 17809. 
BELLFLOWER FAMILY. 


Herbs (some tropical species shrubs or even trees), with alternate exstipulate 
leaves, usually milky juice, and racemose spicate paniculate or solitary perfect 
flowers. Calyx-tube adnate to the ovary, its limb mostly 5-lobed or 5-parted, the 
lobes equal or slightly unequal, valvate or imbricate in the bud, commonly per- 
sistent. Corolla gamopetalous, regular, inserted at the line where the calyx be- 
comes free from the ovary, its tube entire, or deeply cleft on one side, its limb 
5-lobed. Stamens 5, alternate with the corolla-lobes, inserted with the corolla; 
filaments separate; anthers 2-celled, introrse, separate or connate. Ovary 2-5- 
celled (rarely 6—10-celled, with the placentae projecting from the axis, or 1-celled 


294 CUCURBITACEAE. Vor. III. 


with two parietal placentae; style simple; stigma mostly 2—-5-lobed ; ovules anatro- 
pous. Fruit a capsule. Seeds very numerous and small; embryo minute, straight ; 
endosperm fleshy. 


About 40 genera and over 1000 species, of wide geographic distribution. 


Capsule opening by lateral pores or valves. 


Corolla campanulate, rarely rotate ; flowers all complete. 1. Campanula. 
Corolla rotate; earlier flowers cleistogamous. 2. Specularia. 
Capsule opening by terminal pores or valves. 3. Jasione. 


1. CAMPANULA [Tourn.] L. Sp. Pl. 163. 1753. 


Perennial or annual herbs, with alternate or basal leaves. Flowers large or small, soli- 
tary, racemose, paniculate, or glomerate, regular, complete, blue, violet, or white. Calyx-tube 
hemispheric, turbinate, obovoid, or prismatic, adnate to the ovary, the limb deeply 5-lobed or 
5-parted (rarely 3-4-parted). Corolla campanulate or rotate, 5-lobed or 5-parted. Stamens 
5, free from the corolla; filaments usually dilated at the base; anthers separate. Ovary 
inferior, 3-5-celled; stigma 3-5-lobed. Capsule wholly or partly inferior, crowned by the 
persistent calyx-lobes, opening on the sides, either near the top, middle or bottom by 3-5 
small valves or perforations, or tending to be indehiscent in some species. [Diminutive of 
the Latin campana, a bell.] 


About 250 species, natives of the northern hemisphere. Besides the following, some 8 others 
occur in the southern and western parts of North America; all known as Bell-flower. Type species: 
Campanula latifolia L. 


*Corolla campanulate; flowers solitary, racemose, glomerate, or panicled. 


Flower solitary at the end of the stem; arctic and alpine plants. 
Corolla 4”-6” long; capsule-openings near the summit. 
Corolla 6’-12” long; capsule-openings near the base. 

Flowers racemose, glomerate, or paniculate. 

Corolia 7”-15” long. 
Stem leaves linear, the basal orbicular, mostly cordate: 2. C. rotundifolia. 
Leaves all ovate to lanceolate; plants pubescent or scabrous. 
Flowers pedicelled, or clustered. 


4 


. C. uniflora. 
C. rotundifolia. 


iS) 


Calyx and corolla glabrous, or calyx finely pubescent. 3. C. rapunculoides. 
Calyx and corolla bristly-hairy. 4. C. Trachelium. 
Flowers sessile in terminal and axillary clusters. 5. C. glomerata. 


Corolla 2”’—5” long. 
Plants rough; style not exserted. 
Corolla white, or tinged with blue, 21%4”-4” long; leaves mostly linear-lanceolate, 
crenulate. 6. C. aparinoides. 
Corolla blue, 5”-6” long; leaves linear, denticulate with minute callous teeth. 
7. C. uliginosa. 
Plants smooth, glabrous, slightly viscid; style long-exserted. 8. C. divaricata. 


** Corolla rotate; flowers spicate. 9. C. americana. 


1. Campanula uniflora L. Arctic Harebell or Bell- 
flower. Fig. 4015. 


Campanula uniflora L. Sp. Pl. 163. 1753. 


Perennial, glabrous or nearly so; stem simple, 1-flow- 
ered, 1’-6’ high. Leaves linear or linear-oblong, acute, 
sessile, thickish, entire or sparingly dentate, 9-18” long, 
or the lower and basal ones spatulate, obtuse and narrowed 
into petioles; flower erect; calyx-tube turbinate, glabrous 
or pubescent, shorter than or equalling the lobes; corolla 
campanulate, 4”-6” long, blue; capsule cylindric or club- 
shaped, about 6” long, erect, opening by valves near the 
summit. 


Labrador and Arctic America to Alaska, south in the Rocky 
Mountains to Colorado. Also in northern Europe and Asia. 
Summer. 


GENUS I. BELLFLOWER FAMILY. 295 


2. Campanula rotundifolia L. Harebell. 
Blue Bells of Scotland. Fig. 4016. 

Campanula rotundifolia L. Sp. Pl. 163. 1753. 

Campanula rotundifolia velutina DC. Fl. France 
6: 432. 1815. 

C. linifolia var. Langsdorfiana A. DC. Prodr. 7: 


AZT.) 1530. fey / 
Campanula rotundifolia Langsdorfiana Britton, 


Mem. Torr. Club 5: 309. 1894. 
Perennial by slender rootstocks, glabrous or 
nearly so or sometimes pubescent or canes- 
cent; stems erect or diffuse, often several 
from the same root, simple or branched, 6—3° 
high. Basal leaves nearly orbicular or broadly 
ovate, usually cordate, slender-petioled, +’—1’ 
wide, dentate or entire, often wanting at flow- 
ering time; stem leaves linear or linear-oblong, 
acute, mostly entire, sessile, or the lower nar- 
rowed into short petioles and somewhat spatu- 
late; flowers several or numerous, racemose or 
sometimes solitary, drooping or spreading, 
slender-pedicelled; calyx-lobes subulate to fili- 
form, spreading, longer than the short-turbi- 
nate tube; corolla blue, campanulate, 7-12” 
long; capsule obconic or ovoid, pendulous, 
ribbed, opening by short clefts near the base. 
On moist rocks and in meadows, Labrador to Alaska, south to New Jersey, Pennsylvania, Illi- 
nois, Nebraska, in the Rocky Mountains to Arizona and in the Sierra Nevada to California. Also 
in Europe and Asia. Consists of many races, differing in pubescence, number and size of flowers: 
arctic and alpine plants are usually 1-few-flowered. Other English names are thimbles, lady’s- 
thimble, heath- or witches’-bells, round-leaved bellflower. June—Sept. 
Campanula patula L., which is retrorse-scabrous on the stems and leaf-margins and nerves, 
re basal leaves obovate to spatulate, has been found in fields in Connecticut, introduced from 
urope. 


3. Campanula rapunculoides L. Creep- 
ing or European Bellflower. Fig. 4017. 
Campanula rapunculoides L. Sp. Pl. 165. 1753. 

Perennial by slender rootstocks; stem gla- 
brous or pubescent, simple or rarely branched, 
leafy, erect, rather stout, 1°-3° high. Leaves 
pubescent or puberulent, crenate-denticulate, 
ovate or ovate-lanceolate, acute or acuminate, 
the lower and basal ones mostly cordate, 3’-6’ 
long, 1-2’ wide, slender-petioled, the upper 
short-petioled or sessile, smaller; flowers short- 
pedicelled, drooping, 17-13’ long in an elon- 
gated bracted 1-sided raceme; corolla campan- 
ulate, blue to violet, rather deeply 5-lobed, 
much longer than the linear spreading calyx- 
lobes; capsule globose, nodding, about 4” in 
diameter, opening by pores near the base. 

In fields and along roadsides, New Brunswick 
to Ontario, southern New York, Pennsylvania and 
Ohio. Naturalized from Europe. July—Sept. 


4. Campanula Trachélium L. Nettle-leaved Bell- 
flower. Fig. 4018. 
C. Trachelium L. Sp. Pl. 166. 1753. 

Perennial; stem rather stout, little branched, usually 
bearing scattered hairs, 3° high or less; basal leaves 
sparingly bristly-pubescent, ovate to reniform, cordate, 
slender-petioled; stem leaves ovate-lanceolate to lanceo- 
late, coarsely irregularly serrate, 23’-5’ long, acute or 
acuminate at the apex, narrowed at the base, short- 
petioled or the upper sessile; flowers nodding in termi- 
nal leafy-bracted racemes; calyx bristly-hairy or gla- 
brate; corolla campanulate, 1-13’ long; capsule opening 
by basal pores. 

Roadsides and thickets, Quebec to southern New York and 
Ohio. Naturalized from Europe. Canterbury bells. July—Sept. 


CAMPANULACEAE. Vo. IIT. 


5. Campanula glomerata L. Clus- 
tered Bellflower. Dane’s Blood. 


Fig. 4019. 
Campanula glomerata L. Sp. Pl. 166. 1753. 


Perennial by short rootstocks; stem stout, 
simple, erect, pubescent, leafy, 1°-2° high. 
Leaves pubescent on both sides, crenulate, 
the lower and basal ones oblong or ovate, 
mostly obtuse, sometimes cordate, slender- 
petioled, 2-4’ long, the upper lanceolate or 
ovate-lanceolate, acute, sessile or clasping, 
smaller; flowers about 1’ long, sessile, erect 
and spreading in terminal and axillary glom- 
erules; corolla campanulate, blue, rather 
deeply 5-lobed; calyx-lobes lanceolate, acu- 
minate; capsule ovoid or oblong, erect, about 
3” high, opening near the base. 


In fields and along roadsides, eastern Massa- 
chusetts and Quebec. Naturalized from Eu- 
trope. Sometimes called canterbury bells, a 
name more properly belonging to C. medium 
and C. Trachelium. June—Aug. 


6. Campanula aparinoides Pursh. Marsh 
or Bedstraw Bellflower. Fig. 4020. 


Campanula aparinoides Pursh, Fl. Am. Sept.159. 1814. 


Perennial; stems very slender or filiform, weak, 
reclining or diffuse, rough with short retrorse bris- 
tles, leafy, paniculately branched, 62° long. 
Leaves lanceolate, or linear-lanceolate, sessile, 
sparingly crenulate with low teeth, or entire, 
rough on the margins and midrib, acute at both 
ends, 2-13’ long, 1”-3” wide; flowers leafy- 
paniculate, 22-4” long; pedicels filiform, diver- 
gent; buds drooping; corolla open-campanulate, 
deeply 5-cleft, white or bluish-tinged, 23”—-4” long, 
its tube equalling or longer than the triangular- 
lanceolate acute calyx-lobes; style included; cap- 
sule subglobose, opening near the base. 


In grassy swamps, Maine to Georgia, Kentucky and 
Colorado. Called also slender bellflower. June—Aug. 


7. Campanula uligindsa Rydb. Blue Marsh 
Bellflower. Fig. 4021. 


Campanula uliginosa Rydb.; Britton, Man. 885. 1901. 


Perennial; similar to the preceding but branches less 
spreading; stem 1°-2° long, retrorsely hispidulous on 
the angles. Leaves linear, 1’-23’ long, 3-22” wide, 
retrorsely hispidulous on the margins and midribs, 
acute, minutely denticulate with callous teeth or entire; 
corolla blue with darker veins, cleft to below the mid- 
dle into lanceolate lobes; capsule subglobose, about 22 


long and nearly as broad, opening near the base. 


In wet meadows, New Brunswick to Saskatchewan, New 
York, Indiana and Nebraska. June-Aug. 


GENUS I. BELLFLOWER FAMILY. 297 


8. Campanula divaricata Michx. Pani- 
cled Bellflower. Fig. 4022. 


Campanula divaricata Michx. Fl. Bor. Am. 1: 109. 
1803. 
Campanula flexuosa Michx. loc. cit. 1803? 


Perennial, glabrous but sometimes viscid; 
stem erect, paniculately branched, slender, 
1°-3° high. Leaves lanceolate, ovate or oblong- 
lanceolate, the uppermost sometimes linear, 
sharply serrate, acuminate at the apex, nar- 
rowed to the base, the upper sessile, the lower 
petioled, 2’-3’ long, 3-12” wide, or the lowest 
commonly shorter and broader; flowers very 
numerous in compound panicles, drooping, 
slender-pedicelled; corolla light blue, cam- 
panulate, about 3” long; calyx-lobes lanceo- 
late, acute, scarcely spreading, often dentate; 
style long-exserted; capsule turbinate, about 
24” long, opening near the middle. 


On rocky banks, mountains of Virginia and 
West Virginia to Kentucky, Georgia and Tennes- 
see. Ascends to 2500 ft.in North Carolina. June— 
Sept. 


g. Campanula americana L. Tall 
Bellflower. Fig. 4023. 


Campanula americana L. Sp. Pl. 164. 1753. 


Annual or biennial, more or less pubes- 
cent; stem erect or nearly so, rather slen- 
der, simple or rarely with a few long 
branches, 2°-6° high. Leaves thin, ovate, 
oblong, or lanceolate, serrate, acuminate 
at the apex, narrowed at the base, petioled. 
or the upper sessile, 3-6’ long, the lowest 
sometimes cordate; flowers in a loose or 
dense terminal sometimes leafy spike, which 
is often 1°-2° long; lower bracts foliaceous, 
the upper subulate; corolla rotate, blue, or 
nearly white, about 1’ broad, deeply 5-cleft; 
calyx-lobes linear-subulate, spreading, style 
declined and curved upward, long-exserted ; 
capsule narrowly turbinate, ribbed, erect, 
4’-5”" long, opening near the summit. 


In moist thickets and woods, New Bruns- 
wick to Ontario and South Dakota, south to 
Florida, Kentucky, Kansas and Arkansas. 
Rare near the coast in the Middle States and 
New England. Ascends to 3000 ft. in West 
Virginia. July—Sept. 


2. SPECULARIA Heist.; Fabr. Enum. Pl. Hort. Helmst. 225. 1763. 
[Lecouzta Durand, Fl. Bourg. 2: 26. 1782.] 


Annual herbs, with alternate toothed or entire leaves, the stem and branches long, slen- 
der. Flowers axillary, sessile or nearly so, 2-bracted, or the upper panicled in some exotic 
species, the earlier (lower) ones small, cleistogamous, the later with a blue or purple nearly 
rotate corolla. Calyx-tube narrow, the lobes in the earlier flowers 3 or 4, in the later 4 or 5. 
Corolla 5-lobed or 5-parted, the lobes imbricated in the bud. Filaments flat; anthers sepa- 
rate, linear. Ovary 3-celled (rarely 2- or 4-celled) ; ovules numerous; stigma usually 3-lobed. 
Capsule prismatic, cylindric, or narrowly obconic, opening by lateral valves. Seeds ovoid, 
oblong, or lenticular. [From Speculum Veneris, the Latin name of the type species. | 

About 10 species, natives of the northern hemisphere, one extending into South America. 
Type species: Campanula Spéculum L.; S. Spéculum (L.) DC., of Europe, which is adventive in 
Lancaster County, Pennsylvania. 


Capsule narrowly oblong. 
Leaves sessile ; capsule-valves near the top. 1. S. biflora. 
Leaves cordate-clasping ; capsule-valves at about the middle. 2. S. perfoliata. 
Capsule linear-cylindric ; leaves sessile ; western. 3. S. leptocarpa. 


CAMPANULACEAE. Vo. III. 


1. Specularia biflora (R.& P.) F.& M. Small 
Venus’ Looking-glass. Fig. 4024. 


—o ee 


Campanula biflora R. & P. Fl. Per. 2: 55. pl. 200. J. 6. 
1799. 


S. biflora F. & M. Ind. Sem. Hort. Petrop. 1: 17. 1835. 
1 
Legouzia biflora Britton, Mem. Torr. Club 5: 309. 1894. 


Glabrous, or nearly so; stem simple or branched, 
very slender, roughish on the angles, 62° high. 
Leaves ovate, oblong, or the upper lanceolate, ses- 
sile, acute or obtuse at the apex, crenate with a few 
teeth, or entire, 4-10” long, or the upper smaller; 
earlier flowers with 3 or 4 ovate to lanceolate calyx- 
lobes, those of the later flowers 4 or 5, lanceolate- 
subulate, longer; capsule oblong-cylindric, 3’—5” 
long, opening by valves close under the calyx-teeth. 


In dry soil, Virginia to Kentucky, Missouri, Kansas, ~ 
Florida and Texas. Also in Oregon, California and 
South America. April—July. 


2. Specularia perfoliata (L.) A. DC. 
Venus’ Looking-glass. Fig. 4025. 

Campanula perfoliata L. Sp. Pl. 169. 1753. 
S. perfoliata A. DC. Mon. Campan. 351. 1830. 
L. perfoliata Britton, Mem. Torr. Club 5: 309. 1894. 

More or less pubescent; stem densely leafy, 
simple or branched from near the base, slender. 
rather weak, sometimes prostrate, retrorse-hispid 
on the angles, or nearly smooth, 6-24’ long. Leaves 
orbicular or broadly ovate, strongly cordate-clasp- 
ing or the lower merely sessile, crenate-dentate 
or sometimes entire, #/-1’ wide; flowers solitary 
or 2-3 together in the axils, sessile, the later 
(upper) ones with 5 (rarely 4) triangular-lanceo- 
late acuminate rigid calyx-lobes, and a rotate blue 
or violet corolla 5-10” broad, the earlier ones 
with 3-4 shorter calyx-lobes longer than the rudi- 
mentary corolla; capsule oblong, or narrowly tur- 
binate, 2”-3” long, finally opening at about the 
middle; seeds lenticular. 

In dry woods, Maine and Ontario to British Co- 
lumbia, south to Florida, Louisiana, Mexico, Arizona 
and Oregon. Called also clasping bellflower. May-— 
Sept. Also in the mountains of Jamaica and Santo 


. 

. 

Domingo. | 
3. Specularia leptocarpa (Nutt.) A. Gray. 


Western Venus’ Looking-glass. Fig. 4020. 


Campylocera leptocarpa Nutt. Trans. Am. Phil. Soc. 
(iis) 3s 2575 ero4s* 


S. leptocarpa A. Gray, Proc. Am. Acad. 11: 82. 1876. 
L. leptocarpa Britton, Mem. Torr. Club 5: 309. 1804. 


Hirsute, or nearly glabrous; stem slender, simple, 
or branched from the base, 6-15’ high. Leaves 
linear-lanceolate to oblong, sessile, not clasping, acute 
at both ends, or the lowest obtuse at the apex, entire 
or sparingly denticulate, #’-1’ long, 132” wide; 
flowers sessile and usually solitary in the axils, the 
later ones with 4-5 subulate calyx-lobes and a rotate 
corolla 5-9” broad, the earlier ones with 3 shorter 
calyx-lobes and rudimentary corolla; capsule linear- 
cylindric, 4”-8” long, less than 1” thick; the upper at. 
length opening near the summit; seeds oblong. 


In dry soil, western Missouri and Kansas to Montana, 
Colorado and Texas. May—Aug. 


GENUS 3. BELLFLOWER FAMILY. 299 


3. JASIONE L. Sp. Pl. 928. 1753. 


Herbs, the flowers in terminal heads subtended by a many-leaved involucre. Calyx-tube 
campanulate, adnate to the ovary, the limb 5-divided. Corolla at first tubular, splitting later 
to the base into 5 linear or linear-oblanceolate segments. Filaments subulate. Anthers united 
at the base, free above. Ovary inferior, 2-celled; stigma club-shaped. Capsule 2-celled, 
dehiscent by two terminal pores. [Greek name of some 
medicinal plant. ] 


Five species of central Europe and the Mediterranean Re- 
gion, the following typical. 


I. Jasione montana L. Sheep’s-bit. Fig. 4027. 
Jasione montana L. Sp. Pl. 928. 1753. 


Annual or biennial, with several stems from a simple 
root; stems 6-12’ high, branched above, leafy below, de- 
cumbent, erect or ascending, the branches spreading. 
Leaves linear, rough-hairy, sessile; flowers in long-pedun- 
ae hemispherical heads; corolla blue, seldom white or 
pink. 


In waste places, Massachusetts to southern New York. Ad- 
ventive from Europe. June—Sept. 


Family 43. LOBELIACEAE Dumort. Comm. 
Bot. 57.. 1822: 


Lospetta FAMILY. 


Herbs, or in tropical regions rarely shrubs or trees, often with milky sap which 
contains a narcotic-acid poison, with alternate, exstipulate, simple, entire, toothed 
or pinnately parted leaves and solitary, spicate, racemose or paniculate flowers. 
Calyx-tube adnate to the ovary, its limb 5-lobed or 5-parted, the lobes equal or 
unequal. Corolla gamopetalous, irregular, often bilabiate, its tube open on one 
side nearly or quite to the base, its limb 5-lobed; stamens 5, inserted with the corolla ; 
filaments sometimes cohering into a tube; anthers united. Ovary 2—5-celled; style 
single; stigma fringed; ovules numerous, sessile, horizontal, anatropous. Fruit 
a 1—5-celled capsule, or a berry. Seeds numerous, with a smooth or furrowed 
testa. Endosperm fleshy. 


About 20 genera and 600 species, of wide geographical distribution. 


1. LOBELIA L. Sp. Pl. 929. 1753. 


Herbs (some tropical species shrubs), with alternate or basal leaves and racemose spicate 
or paniculate, often leafy bracted, red, yellow, blue or white flowers. Calyx-tube turbinate, 
hemispheric or ovoid, adnate to the ovary. Corolla-tube straight, oblique or incurved, divided 
to the base on one side, 2-lipped in our species, the lobe on each side of the cleft erect or 
recurved, turned away from the other three which are somewhat united, the sinuses inclining 
to extend to the base of the corolla at maturity so as to divide it into 5 petals. Stamens free 
from the corolla-tube, monadelphous, at least above, two or all the 5 anthers with a tuft of 
hairs at the tips, three of them usually larger than the other two, all united into a tube or 
ring around the style. Ovary 2-celled, the 2 placentae many-ovuled; stigma 2-lobed or 2-cleft. 
Capsule loculicidally 2-valved. [Named after Matthias de L’Obel, 1538-1616, a Flemish 
botanist. ] 

About 250 species, of wide geographic distribution. Besides the following, some 16 others 
occur in the southern and western United States. Type species: Lobelia Dortmanna L. 

* Aquatic; stem simple, nearly naked; flowers light blue. 
Leaves terete, hollow, obtuse, tufted at the base. 1. L. Dortmanna. 
Leaves flat, linear-oblong or spatulate, entire or glandular-denticulate. 2. L. paludosa. 
** Terrestrial plants of wet or dry soil; stems leafy. 


1. Corolla-tube 5”—12” long. 
Flowers bright scarlet (rarely white) ; corolla-tube 10”—12” long. 3. L. cardinalis. 
Flowers blue, white, or blue and white; corolla-tube 5”—7” long. 
Leaves ovate, lanceolate, or the lower ones obovate. 
Leaves glabrous or sparingly pubescent. 


Calyx-lobes hirsute; sinuses with large deflexed auricles. 4. L. syphilitica. 
Calyx-lobes glabrous or glandular, usually without auricles. 
Leaves ovate to ovate-lanceolate, spreading. 5. L. amoena. 
Leaves linear to narrowly lanceolate, erect. 6. L. elongata. 
Leaves densely puberulent ; calyx hirsute; auricles small. 7. L. puberula. 
Leaves elongated-linear, strongly glandular-denticulate. 8. L. glanduosa, 


300 LOBELIACEAE. Vor. III. 


2. Corolla-tube only 2”-4” long. 
Stems mostly simple; flowers in terminal spike-like racemes. 


Sinuses of the calyx without auricles. g. L. spicata. 
Sinuses of the calyx with reflexed subulate auricles. 10. L. leptostachys. 
Stems mostly paniculately branched; flowers in loose racemes. 
Stem stout, pubescent; leaves ovate or oblong, dentate. 11. L. inflata. 
Stems slender, glabrous; stem-leaves narrow, the basal wider. 
Pedicels mostly longer than flowers, 2-bracteolate near the middle. 12. L. Kalmii. 
Pedicels not longer than flowers, not bracteolate, or only so at the base. 
Corolla 2%”-3%” long; calyx-tube hemispheric in fruit. 13. L. Nuttallii. 
Corolla 44%4”-5%4” long; calyx-tube turbinate. 14. L. Canbyt. 


1. Lobelia Dortmanna L. Water Lobelia. Water 
Gladiole. Fig. 4028. 


Lobelia Dortmanna L. Sp. Pl. 929. 1753. 


Perennial, aquatic, glabrous throughout, somewhat 
fleshy; roots numerous, white, fibrous; stem slender, 
simple, erect, hollow, minutely scaly, 6-18’ high. Leaves 
all submersed and tufted at the base of the stem, terete, 
hollow, obtuse, longitudinally divided by a partition, 
1-2’ long, about 2” thick; flowers in a loose terminal 
raceme, blue, 6’—-8” long; pedicels filiform, shorter than 
or equalling the flowers; calyx-lobes subulate or lanceo- 
late, shorter than the tube, the sinuses usually not at 
‘all appendaged; corolla-tube 3-4” long, its lower lip 
glabrous or nearly so. 


Borders of ponds, usually in sandy soil, sometimes wholly 
emersed when the water is low, New Jersey and Pennsyl- 
vania to Newfoundland, Wisconsin, Washington and British 
Columbia. Also in Europe. July—Sept. 


2. Lobelia paludésa Nutt. Swamp Lobelia. 
Fig. 4029. 
Lobelia paludosa Nutt. Gen. 2: 75. 1818. 


Perennial, aquatic, glabrous throughout; roots few and 
thick; stem nearly naked, slender, simple, or branched 
above, 1°-4° high. Leaves mostly tufted at the base, flat, 
narrowly oblong or spatulate, emersed, obtuse or acutish, 
entire or repand-denticulate and glandular, those of the 
stem few, small and sessile, the basal ones 2-9’ long, 2’-4” 
wide, narrowed into petioles; flowers pale blue, racemose, 
5-6” long; calyx-lobes narrowly lanceolate, about as long 
as the tube, the sinuses commonly not at all appendaged; 
corolla-tube 3’-4” long, its lower lip pubescent at the base. 


In swamps and ponds, Delaware to Florida and Louisiana 
mostly near the coast. May-July. 


3. Lobelia cardinalis L. Cardinal-flower. 
Red Lobelia. Red Betty. Fig. 4030. 


Lobelia cardinalis L. Sp. Pl. 930. 1753. 


Perennial by offsets; stem slightly pubescent, or 
glabrous, leafy, simple or rarely branched, 2°—43° 
high. Leaves oblong, oval, ovate-lanceolate, or 
lanceolate, thin, glabrous or sparingly pubescent, 
2’-6’ long, #/-12’ wide, acuminate or acute at both 
ends, crenulate or denticulate, the upper sessile, 
the lower petioled; flowers racemose, commonly 
numerous, bright scarlet or red (rarely white), 
1-13’ long; bracts usually glandular; calyx gla- 
brous or pubescent, its lobes linear, elongated, 
acute; corolla-tube nearly or quite 1’ long; larger 
anthers glabrous. 


In moist soil, New Brunswick to Florida, Ontario, 
Kansas, Colorado and Texas. Slink-weed. Hog’s- 
physic. July—Sept. 


_—_—_ 


GENUS I. LOBELIA FAMILY. 


4. Lobelia syphilitica L. Great Lobelia. 
Blue Cardinal-flower. Fig. 4031. 


Lobelia syphilitica L. Sp. Pl. 931. 1753. 
Lobelia syphilitica ludoviciana A. DC. Prodr. 7: 377. 
1839. 


Perennial by short offsets; stem sparingly pubes- 
cent, rather stout, very leafy, usually simple, 1°-3° | 
high. Leaves glabrous or sparingly puberulent, 2’-6’ 
long, 3’-2’ wide, oval, oblong, or lanceolate, acute 
or acuminate at the apex, narrowed at the base, re- 
pand-denticulate, irregularly crenate-dentate or near- 
ly entire, sessile, or the lower obovate, obtuse and 
narrowed into petioles; flowers bright blue, or occa- 
sionally white, 10’—12” long, densely racemose, leafy- 
bracted; calyx-lobes lanceolate, acuminate, pubescent 
or ciliate, the sinuses appendaged by large deflexed 
auricles; corolla-tube 5”—6” long, about 2” thick, the 
lobes of its larger lip oblong-oval, obtuse or acutish, 
glabrous; larger anthers glabrous. 


In moist soil, Maine and Ontario to South Dakota, 
Colorado, Georgia, Louisiana and Kansas. MHybridizes 
with the preceding species. High-belia. July—Oct. 


5. Lobelia amoéna Michx. Southern Lobelia. 
Fig. 4032. 


Lobelia amoena Michx. Fl. Bor. Am. 2: 152. 1803. 
Lobelia amoena glandulifera A. Gray, Syn. Fl. 2: 4. 1878. 


Nearly glabrous throughout, perennial; stem simple, 
slender, leafy, 1°-4° high. Leaves thin, ovate to oblong- 
lanceolate, narrowly oblong, or oval, repand-dentate or 
denticulate, the lower petioled and mostly obtuse, 2’-6’ 
long, the upper sessile, acute or acutish, smaller; flowers 
blue, racemose, nearly 1’ long; bracts narrow and small, 
or the lower foliaceous, glandular; pedicels 1-23” long; 
calyx-lobes linear-subulate, acuminate, glabrous, glandu- 
lar, elongated, the sinuses usually not auricled; corolla- 
tube 5-7” long, 13”-2” thick; larger anthers glabrous. or 
puberulent at the tip; lobes of the larger lip of the co- 


rolla broadly ovate to oval, obtuse, glabrous. 


In swamps, Delaware to Florida.and Alabama. July—Sept. 


6. Lobelia elongata Small. Long-leaved 
Lobelia. Fig. 4033. 


L. elongata Small, Fl. SE. U. S. 1144. 1903. 


Perennial, at least by offsets, glabrous or nearly 
so; stem simple, 1°-3° tall, commonly wand-like. 
Leaves rather few, erect or ascending, linear or 
nearly so, #—4’ long, mostly acute, serrate or 
dentate-serrate with gland-tipped teeth, sessile or 
narrowed into petiole-like bases; flowers deep- 
blue in rather closely flowered, but not densely 
flowered, one-sided racemes 4-12’ long; bracts 
linear to lanceolate, serrate with gland-tipped 
teeth, the lower ones sometimes surpassing the 
corollas; calyx-lobes elongate, linear-subulate or 
linear-setaceous, entire, as long as the corolla or 
shorter, without auricles at the sinuses; corolla- 
tube 5-8” long, lobes of the lower lip oval or 
ovate, glabrous. 


In low grounds or swamps, Virginia to Florida and 
Louisiana. Aug.—Oct. 


LOBELIACEAE. Vor. III. 


7. Lobelia pubérula Michx. Downy Lobelia. 
Fig. 4034. 
Lobelia puberula Michx. Fl. Bor. Am. 2: 152. 1803. 


Perennial, densely and finely puberulent all over, 
slightly viscid; stem simple, or rarely with a few 
branches, stout or slender, leafy, 1°-3° high. Leaves 
oval, oblong, ovate, or obovate, 1-2’ long, rather thick, 
the lower petioled, all obtuse or the uppermost acute, 
denticulate or crenate-dentate, the teeth often glandular; 
flowers blue, 8’—10” long, in long spike-like racemes; 
lower bracts, or sometimes all of them, foliaceous, 
glandular; pedicels very short; calyx hirsute or pubes- 
cent, its lobes narrowly lanceolate, elongated, usually 
with small short rounded auricles at the sinuses; cor- 
olla-tube about 5” long, 1-13” thick, the lobes of its 
larger lip broadly ovate, glabrous; larger anthers mi- 
nutely bearded. 


In moist sandy soil, southern New Jersey to Florida, 
Illinois, lowa, Kansas and Texas. Ascends to 3500 ft. in 
North Carolina. Aug.—Oct. 


8. Lobelia glandulosa Walt. Glandular Lobelia. 
Fig. 4035. 
Lobelia glandulosa Walt. Fl. Car. 218. 1788. 


Perennial; stem slender, leafy below, nearly naked 
above, simple, glabrous, or sparingly pubescent, 1°-4° 
high. Leaves elongated-linear to narrowly lanceolate, 
thick, glabrous, strongly glandular-dentate, the lower 
petioled, obtuse, often 7’ long and 4” wide, the upper 
sessile, acutish, shorter; flowers racemose-spicate, secund, 
often few and distant, blue, nearly 1’ long; bracts narrow, 
glandular; peduncles very short, sometimes with a pair of 
glands near the base; calyx-tube often densely hirsute, its 
lobes subulate, the sinuses not appendaged; corolla-tube 
5-6” long, 1”-14” thick, about twice as long as the calyx- 
lobes; larger lip of the corolla pubescent within at the 
base, its lobes ovate, acutish; anthers all bearded at the tip. 


ee re 


In swamps near the coast, southern Virginia to Florida. 
July-Sept. 


g. Lobelia spicata Lam. Pale Spiked Lobelia. Fig. 4036. 


Lobelia spicata Lam. Encycl. 3: 587. 1789. 
L. spicata hirtella A. Gray, Syn. Fl. 2: 6. 1878. 
L. spicata parviflora A. Gray, Syn. Fl. 2: 6. 1878. 


Perennial or biennial, puberulent, smooth or roughish; 
stem strict, simple, leafy, 1°-4° high. Leaves thickish, 
pale green, repand-dentate, crenulate, or entire, the 
basal ones commonly tufted, broadly oblong, oval, or 
obovate, very obtuse, narrowed into short petioles, 
1-33’ long; 1-2’ wide; stem leaves sessile, oblong, 
lanceolate, or spatulate, obtuse, the uppermost gradually 
smaller and acutish; flowers pale blue, 3’-5” long, 
densely or distantly racemose-spicate, the inflorescence 
sometimes 2° long; bracts linear, entire; pedicels very 
short, ascending; calyx-tube turbinate, usually glabrous, 
shorter than its subulate spreading, sometimes hirsute 
or ciliate lobes, the sinuses usually not at all append- 
aged; corolla-tube about 23” long and 1” thick. 


In dry, mostly sandy soil, or in meadows, Prince Edward 
Island to Saskatchewan, North Carolina, Alabama, Louisi- 
ana and Arkansas. Races differ in pubescence and in size 
of flowers. Ascends to 2500 ft. in Virginia. June—Aug. 


GENUS I. LOBELIA FAMILY. 


10 Lobelia leptostachys A. DC. Spiked 
Lobelia. Fig. 4037. 
Lobelia leptostachys A. DC. Prodr. 7: 376. 18309. 


Similar to the preceding species; stem usually by) 
stouter, puberulent or glabrous, 2°-4° high. Basal any 
leaves oval or obovate, obtuse; stem leaves spatu- ONG; 
late, oblong, or lanceolate, obtuse, sometimes uy 
slightly scabrous, denticulate or entire, or the 
uppermost narrower and acute; spike-like raceme 
elongated, usually dense; bracts linear, glabrous; 
pedicels very short; calyx-lobes linear-subulate, 
nearly as long as the corolla-tube, the sinuses 
with subulate deflexed auricles; flowers blue, 
"5! long. 


In dry soil, Virginia to Ohio and Illinois, Georgia 
and Kansas. June—Aug. 


tr. Lobelia inflata L. Indian or Wild 
Tobacco. Eyebright. Fig. 4038. 
Lobelia inflata L. Sp. Pl. 931. 1753. 

Annual, pubescent or hirsute, very acrid; 
stem leafy, commonly paniculately branched, 
1°-3° high. Leaves thin, repand-dentate or 
denticulate, the lower oval or obovate, obtuse, 
1’-23’ long, narrowed into short petioles, the 
upper sessile, oval, oblong, ovate, or ovate- 
lanceolate, obtuse, or the uppermost acute; 
flowers light blue, 2-3” long, usually distant 
in somewhat spike-like racemes; lower bracts 
foliaceous, the upper subulate; pedicels 2-5” 
long in fruit; calyx glabrous or nearly so, 
its subulate lobes about as long as the corolla; 
capsule inflated, 3-4” long, finely transversely 
veined between the ribs. 

In fields and thickets, usually in dry soil, Labra- 
dor to Saskatchewan, Georgia, Kansas and Arkan- 
sas. Gag-root. Puke-weed. Asthma-weed. Low- 
belia. Emetic-weed. Bladder-pod lobelia. July-— 
Nov. 


12. Lobelia Kalmii L. Brook or Kalm’s 
Lobelia. Fig. 40309. 
Lobelia Kalmii L. Sp. Pl. 930. 1753. 


Perennial by short offsets, glabrous through- 
out, or sparingly pubescent below; stem leafy, 
erect, paniculately branched, rarely simple, 
slender, 6-20’ high... Lower and basal leaves 
spatulate, obtuse, narrowed into short petioles, 
sparingly repand-denticuiate, or entire, 6’—12” 
long, 14’-24’ wide; upper leaves sessile, usually 
longer and narrower, linear, linear-oblong, or 
narrowly spatulate, the uppermost acute; flow- 
ers light blue, 4’-5” long, in loose racemes; 
lower bracts linear-lanceolate, the upper subu- 
late; pedicels nearly filiform, 4’—12” long, usu- 
ally 2-glandular or 2-bracteolate near the middle; 
calyx-lobes lanceolate-subulate, longer than the 
turbinate tube, the sinuses not appendaged; cap- 
sule wholly inferior, not inflated, campanulate or 
subglobose, about 2” long. 

On wet banks, and in wet meadows, Nova Scotia 


to New Jersey, west to Ontario, Manitoba, Ohio, 
Michigan and Iowa. July—Sept. : 


304 LOBELIACEAE. Vor. III. 


13. Lobelia Nuttallii R. & S. Nuttall’s Lo- 
belia. Fig. 4040. 


Lobelia gracilis Nutt. Gen. 2: 77. 1818. Not Andr. 
Lobelia Nuttallii R. & S. Syst. 5: 39. 1819. 


Annual, or perhaps biennial, glabrous throughout, 
or puberulent below; stem weak, usually reclining, 
very slender, loosely branched, at least when old, 
1°-3° long. Basal leaves spatulate to oval, obtuse, 
mostly petioled, 6’—-12” long, slightly repand, or en- 
tire; stem leaves distant, linear, linear-oblong, or 
slightly spatulate, longer and narrower, entire or 
sparingly glandular-denticulate; flowers 23’—4” long, 
pale blue, loosely racemose; bracts linear or the 
upper subulate; pedicels filiform, 2-4” long, naked, 
or minutely 2-bracteolate near the base; calyx-lobes 
subulate, longer than the depressed-hemispheric 
strongly ribbed tube, the sinuses unappendaged; cap- 
sule depressed-globose, half-inferior, about 1” long. 


In sandy soil, Long Island to Pennsylvania, Florida 
and Georgia. June—Sept. 


14. Lobelia Canbyi A. Gray. Canby’s Lo- 
belia. Fig. 4041. 
Lobelia Canbyi A. Gray, Man. Ed. 5, 284. 1867. 


Annual, slightly puberulent, usually glabrous; stem 
erect, slender, paniculately branched, or simple, 2°-3° 
high. Stem leaves linear or linear-oblong, 2’—13’ 
long, 3-13” wide, the lower obtuse, sometimes 
slightly repand-denticulate, the upper acute, nar- 
rower, entire; flowers racemose, blue, 4’-53” long; 
lower bracts linear, the upper subulate; pedicels 
erect or ascending, naked, filtform, 1-3” long; 
calyx-lobes_ linear-subulate, glandular-denticulate, 
equalling or somewhat longer than the narrowly 
turbinate tube, mostly shorter than the tube of the 
corolla; capsule oblong-turbinate, 2” long, shorter 
than the calyx-tube. 


Swamps, New Jersey to South Carolina. July—Sept. 


Family 44. CICHORIACEAE Reichenb. Fl. Excurs. 248. 1831. 
CuHicory FAMILy. 


Herbs (two Pacific Island genera trees), almost always with milky, acrid or 
bitter juice, alternate or basal leaves, and yellow, rarely pink, blue purple or white 
flowers in involucrate heads (anthodia). Bracts of the involucre in I to several 
series. Receptacle of the head flat or flattish, naked, scaly (paleaceous ), smooth, 
pitted, or honeycombed. Flowers all alike (heads homogamous), perfect. Calyx- 
tube completely adnate to the ovary, its limb (pappus) of scales, or simple or 
plumose bristles, or both, or wanting. Corolla gamopetalous, with a short or long 
tube, and a strap-shaped (ligulate) usually 5-toothed limb (ray). Anthers con- 
nate into a tube around the style, the sacs sagittate or auricled at the base, not 
tailed, usually appendaged at the summit, the sitnple pollen-grains usually 12-sided. 
Ovary 1-celled; ovule 1, anatropous; style very slender, 2-cleft, or 2-lobed, the 
lobes minutely papillose. Fruit an achene. Seed erect; endosperm none; radicle 
narrower than the cotyledons. 


About 70 genera and 1500 species, of wide geographic distribution. The family is also known 
as LIGULIFLORAE, and is often.regarded as a tribe of the COMPOSITAE. 


* Pappus of scales, or of scales and bristles, or none. 


Flowers blue or white; pappus of blunt scales. 1. Cichorium. 
Flowers yellow. 
Bracts of the involucre membranous, or herbaceous. 
Pappus none; achenes 20-30-nerved. 2. Lapsana. 
Pappus none; achenes 8-10-ribbed. 3. Serinea. 


GENUS |. CHICORY FAMILY. 305 


Pappus of rounded scales, with or without an inner series of bristles. 
Bracts of the involucre 9-18, reflexed in fruit, not keeled; pappus-bristles usually more 
numerous than the scales. 


Annuals; pappus-scales 5, obovate or rounded. 4. Krigia. 
Perennials ; pappus-scales 10-15, linear or oblong, minute. 5. Cynthia. 
Bracts of the involucre 5-8, erect in fruit, keeled; pappus-bristles and scales 5. 
6. Cymbia. 
Bracts of the involucre thickened and keeled after flowering ; pappus none. 7. Arnoseris. 
** Pappus, at least some of it, of plumose bristles. 
Receptacle chaffy. 8. Hypochaeris. 


Receptacle naked. 
Plume-branches of the pappus not interwebbed. 


Flowers yellow; plants scapose, the leaves basal. 9. Apargia. 
Flowers yellow ; plants leafy-stemmed. 10. Picris. 
Flowers pink. ‘ 11. Ptiloria: 
Plume-branches of the pappus interwebbed. 12. Tragopogon. 
*** Pappus of simple bristles or of some soft white scales. 
{ Receptacle with a few deciduous bristles; flowers yellow 13. Malacothri-x. 


tt Receptacle naked. 


1. Achenes spinulose, or with short processes near the summit. 
Heads few- (6-15-) flowered, yellow; stem branching. 14. Chondrilla. 
Heads many-flowered, yellow, solitary on scapes. 15. Leontodon, 
2. Achenes smooth, or papillose, not spinulose toward the summit. 
(a) Achenes flattened. 
Achenes truncate, not beaked; flowers yellow. 16. Sonchus. 
Achenes narrowed at the summit, or beaked; flowers blue or yellow. 17. Lactuca. 
(b) Achenes cylindric, or prismatic. 


Achenes terete, not narrowed either at the base or summit; flowers pink or purple. 18. Lygodesmia. 
Achenes narrowed at the base, narrowed or beaked at the summit; flowers yellow. 
Achenes slender-beaked. 


Pappus-bristles not surrounded by a villous ring at base. 19. Agoseris. 
Pappus-bristles surrounded by a villous white ring at base. 20. Sitilias. 
Achenes merely narrowed above, not beaked. 21. Crepis. 
Achenes narrowed at the base, otherwise columnar, truncate (except in 2 species of Hieracium). 
Flowers yellow, orange, or red. 22. Hieracium. 
Flowers white, cream-color, or purple. 23. Nabalus. 


1. CICHORIUM [Tourn.] L. Sp. Pl. 813. 1753. 


Erect branching herbs, with alternate and basal leaves, those of the stem and branches 
usually very small and bract-like, and large heads of blue purple pink or white flowers, 
peduncled, or in sessile clusters along the branches. Involucre of 2 series of herbaceous 
bracts, the outer somewhat spreading, the inner erect and subtending, or partly enclosing, 
the outer achenes. Receptacle flat, naked, or slightly fimbrillate. Rays truncate and 
5-toothed at the apex. Anthers sagittate at the base. Style-branches slender, obtusish., 
Achenes 5-angled or 5-ribbed, truncate, not beaked. Pappus of 2 or 3 series of short blunt 
scales. [From the Arabic name. | : 


About 8 species, natives of the Old World, the following typical. 


1. Cichorium Intybus L. Chicory. Wild Succory. 
Blue Sailors. Bunk. Fig. 4042. 


Vichorium Intybus L. Sp. Pl. 813. 1753. 
Cichorium Intybus divaricatum DC, Prodr. 7: 84. 1838. 
Perennial from a long deep tap-root; stems slightly 
hispid, stiff, much branched, 1°-3° high. Basal leaves 
spreading, runcinate-pinnatifid, dentate or lobed, spatulate, 
3-6’ long, narrowed into long petioles; upper leaves much 
smaller, lanceolate or oblong, lobed or entire, clasping and 
auricled at the base; heads numerous, 1-12’ broad, 1-4 \ 
together in sessile clusters on the nearly naked or bracted 
branches, or sometimes peduncled; inner bracts of the 
involucre about 8; flowers blue, or sometimes white. 
Roadsides, fields and waste places, Nova Scotia to Minne- 
sota, Washington, North Carolina, Kansas, Colorado and Cali- 
fornia. Bermuda. The ground-up root is used as a substitute 
or adulterant for coffee. Heads usually closed by noon. The 
flowers sometimes bear supplementary rays. Blue daisy or 
dandelion. Coffee-weed. Bachelor’s-buttons. Consists of sev- 
eral races. July—Oct. 


20 


306 CICHORIACEAE. Vor. III. 


2. LAPSANA L. Sp. Pl. 811. 1753. 


Annual erect branching herbs, with alternate dentate or pinnatifid leaves, and small 
panicled slender-peduncled heads of yellow flowers. Involucre nearly cylindric, its principal 
bracts in I series, nearly equal, with a few exterior small ones at the base. Receptacle flat, 
naked. Rays truncate and 5-toothed at the apex. Anthers sagittate at the base. Style- 
branches slender. Achenes obovate-oblong, 20-30-nerved, somewhat flattened, narrowed 
below, rounded at the summit. Pappus none. [Greek, Jampsana, the name of a crucifer.] 

About 9 species, natives of the Old World, the following typical. 


1. Lapsana communis L. Nipplewort. Succory 
Dock-cress. Fig. 4043. 


Lapsana communis L. Sp. Pl. 811: 1753. 


Stem paniculately branched, glabrous above, more or 
less hispid-pubescent below, 1°-33° high. Lower leaves 
ovate, repand-dentate, obtuse, thin, pubescent, or glabrate, 
petioled, 2-4’ long, often with 2-6 lobes on the petiole, 
the uppermost oblong or lanceolate, sessile, acute, much 
smaller, mostly entire; heads very numerous, 3-6” broad; 
involucre oblong-cylindric, 2’-3” high, and of about 8 
linear glaucous principal bracts and several very small 
outer ones. 


Along roadsides and in waste places, Quebec and Ontario 
to New Jersey, Pennsylvania and Michigan. Also on the 
Pacific Coast and in Jamaica. Naturalized from Europe. 
Called also bolgan-leaves, ballogan. June—Sept. 


3. SERINIA Raf. Fl. Ludov. 149. 1817. 


[Apocon Ell. Bot. S. C. & Ga. 2: 267. 1824.] 

Low glaucescent branching annual herbs, with alternate clasping entire or lobed leaves, 
or those of the stem sometimes appearing as if opposite, and few small long-peduncled heads 
of yellow flowers. Involucre broadly campanulate, its bracts about 8, equal, membranous, 
becoming concave after flowering. Receptacle flat, naked. Rays truncate and 5-toothed at 
the apex. Anthers sagittate at the base. Style-branches slender. Achenes obovoid, 
8-10-ribbed, contracted at the base, rounded at the summit. Pappus none, or a mere vestige. 
[Greek, small chicory.] 


Three known species, natives of the southern United States, 
the following typical. 


1. Serinia oppositifolia (Raf.) Kuntze. Serinia. 


Fig. 4044. 
Krigia oppositifolia Raf. Fl. Ludov. 57. 1817. 
Apogon humilis Ell. Bot. S. C. & Ga. 2: 267. 1824. 
Serinia oppositifolia Kuntze, Rev. Gen. Pl. 364. 1891. 

Glabrous throughout, or slightly glandular-pubescent along 
the ends of the peduncles, branched from the base, 4-10’ 
high. Basal and lower leaves petioled, oblong-lanceolate or 
spatulate in outline, acute or obtuse, entire, lobed or pin- 
natifid, 3’-5’ long, 2”-6” wide; upper leaves mainly sessile, 
clasping, alternate, or appearing as if opposite, usually entire, 
smaller; peduncles very slender, sometimes 4’ long; heads 
13-2” broad; bracts of the involucre acute or acuminate, 
about the length of the rays. 

Kansas to Texas, east to North Carolina and Florida. Recorded 
from Missouri. March—May. 


4. KRIGIA Schreb. Gen. Pl. 532. 1791. 


An annual herb, with scapose stems, basal, entire, sinuate-dentate or pinnatifid leaves, 
and a small or middle-sized head of yellow flowers, solitary at the end of the scape. Invo- 
lucre campanulate, its herbaceous bracts 9-18, reflexed in fruit, in 2 series, with no exterior 
shorter ones. Receptacle flat, naked. Rays truncate and 5-toothed at the apex. Anthers 
sagittate at the base. Style-branches slender, obtusish. Achenes turbinate, 15-20-ribbed, 
truncate. Pappus in 2 series, the outer of 5 thin broad rounded scales, the inner of 10 or 
more slender naked bristles. [In honor of David Krig, who collected plants in Maryland 
early in the eighteenth century. ] 


A monotypic genus of North America. This and the two following genera were included in 
Adopogon Neck., in our first edition, but that genus is not typified, and the name probably belongs 
to an Old World plant. 


GENUS 4. CHICORY FAMILY. 307 


1. Krigia virginica (L.) Willd. Carolinia Dwarf Dandelion. Krigia. 
Fig. 4045. 
Hyoseris virginica L. Sp. Pl. 809. 1753. 
Hyoseris caroliniana Walt. Fl. Car. 194. 1788. 


Krigia virginica Willd. Sp. Pl. 3: 1618, 1804. Le 
Krigia caroliniana Nutt. Gen. 2: 126. 1818. AN 


Adopogon carolinianum Britton, Mei. Torr. Club 5: 
346. 1894. 


i 


Annual, acaulescent; scapes usually several 
from the same root, very slender, glabrous or 
hispidulous, monocephalous, 1’-15’ high, simple, or 
sometimes branched at or near the base. Leaves 
commonly all basal, tufted, spatulate, lanceolate 
or linear, pinnatifid, sinuate, lobed, dentate or 
rarely entire, 1-6’ long, narrowed at the base into 
usually margined petioles; head 3’-7” broad; 
involucre of 9-18 linear-lanceolate bracts, reflexed 
after the fall of the narrowly turbinate somewhat 
5-angled achenes; pappus of 5 round short scales 
and 10 or more long capillary bristles. 


In dry, sandy soil, Maine to Ontario and Minne- 
sota, Florida and Texas; also in Washington. April— 
Aug. 


5. CYNTHIA D. Don, Edinb. Phil. Journ. 12: 305. 1829. 


Perennial herbs, with tufted basal leaves, the large many-flowered heads at the ends of 
simple or branched scapes. Flowers orange or yellow. Bracts of the involucre 9-18, reflexed 
in fruit, lanceolate to linear-lanceolate, not keeled. Pappus of 10-15 small scales and as 
many long bristles or more. [Mythological name. ] 

Four species, natives of North America. Type species: Cynthia virginica (L.) D. Don. 


Caulescent, branched above. 1. C. virginica. 
Acaulescent, monocephalous. : 2. C. Dandelion. 


1. Cynthia virginica (L.) D. Don. Cynthia. Virginia Goatsbeard. 
Fig. 4046. 


Tragopogon virginicum L. Sp. Pl. 789. 1753. 

Krigia amplexicaulis Nutt. Gen. 2: 127. 1818. 

Cynthia virginica D. Don, Edinb. Phil. Journ. 12: 309. 1829. 
Adopogon virginicum Kuntze, Rev. Gen. Pl. 304. 1891. 
Cynthia falcata Standley, Contr. Nat. Herb. 13: 356. t1g91It- 


Perennial, glabrous and glaucous; stem 1°-24° high, 
1-leaved and branched above, bearing 1-6 long-peduncled 
heads at its summit and usually 1 oblong, entire or 
toothed clasping leaf below the middle. Basal leaves 
tufted, runcinate, sinuate, denticulate, or entire, 2’—7’ 
long, narrowed into margined petioles, heads about 13’ 
broad; involucre of 9-15 lanceolate nerveless bracts, 
3-4” long, reflexed in fruit; achenes nearly oblong; 
pappus of 10-15 small oblong scales and an equal or 
greater number of capillary bristles; flowers orange to 
reddish orange. 


In moist woods and meadows, Massachusetts to southern 
Ontario and Manitoba, Georgia, Kentucky, Missouri and 
Colorado. Ascends to 4000 ft. in Virginia. False dandelion. May—Oct. 


308 CICHORIACEAE. Vot. III. 


2. Cynthia Dandélion (L.) DC. Dwarf 
Dandelion or Goatsbeard. Fig. 4047. 
Tragopogon Dandelion L. Sp. Pl. Ed. 2, 1111. 1763. 
Krigia Dandelion Nutt. Gen. 2: 127. 1818. 
Cynthia Dandelion DC. Prodr. 7: 89. 1838. 


seer eee: Dandelion Kuntze, Rev. Gen. Pl. 304. 
18g1. 

Perennial, acaulescent, glabrous and some- 
what glaucous; scape 6-18’ high, slender leaf- 
less, with a single head. Stolons filiform, bear- 
ing globose tubers; leaves all basal, tufted, 
linear-lanceolate to spatulate, entire, denticulate, 
sinuate, or pinnatifid, narrowed at the base, 
3-6’ long, 2-5” wide; head about 1’ broad; 
involucre nearly 3’ high; pappus similar to that 
of the preceding species. 


In moist soil, Maryland to Florida, Illinois, Mis- 
souri, Kansas and Texas. April—June. 


a 
DR . 6. CYMBIA (T. & G.) Standley, Contr. 
MA\ ; Nat. Herb. 13: 354. agin: 

An annual acaulescent herb, the leaves forming rosettes, the scapes monocephalous. 
Involucral bracts 5-8, ovate to ovate-lanceolate, remaining erect in fruit. Receptacle naked. 
Rays yellow, 5-toothed. Achenes turbinate, ribbed. Pappus an outer series of 5 obovate 
scales, and an inner series of 5 bristles, much longer than the scales. [Greek, referring to 
the cup-shaped fruiting involucre. ] 

A monotypic genus of the south-central United States. 


we 
1. Cymbia occidentalis ( Nutt.) Standley. A) 
Western Dwarf Dandelion. Fig. 4048. NX 


Krigia occidentalis Nutt. Journ. Acad. Phila. 7: 104. 
1834. 


Adopogon occidentale Kuntze, Rev. Gen. Pl. 304. 
1891. 


C. occidentalis Standley, Contr. Nat. Herb. 13: 354. 
IQIlI. 


Scapes tufted, 2’-8’ high, usually glandular, at | 
least above, sometimes glabrous, bearing a single 
head 5’-10” broad. Leaves basal, lanceolate to 
obovate, entire to pinnatifid, mostly shorter than 
the scapes; involucre 2-3” high, firm and keeled 
at maturity, remaining erect; achenes transversely 
wrinkled; pappus of 5 obovate scales and 5 alter- 
nating bristles, or these wanting. 


Prairies, southern Missouri and Kansas to Texas. 
April-May. 


7. ARNOSERIS Gaertn. Fr. & Sem. 2: 355. pl. 157. 1701. 


A low annual scapose herb, glabrous, or nearly so, with tufted basal dentate or nearly 
entire leaves. Scapes several or numerous, simple or branched, upwardly thickened below 
the solitary heads of yellow flowers. Involucre campanulate, its bracts in I series, narrow, 
equal, thickened and keeled on the back after flowering, rarely with a few outer minute 
ones. Receptacle flat, pitted, not chaffy. Anthers sagittate. Style-branches obtuse. Achenes 
oval, 8-10-ribbed, narrowed below, truncate, or with a denticulate margin. Pappus none. 
(Greek, lamb-succory. | 

A monotypic genus of western Europe. 


GENUS 7. CHICORY FAMILY. 


1. Arnoseris minima (L.) Dumort. Lamb 
Succory. Fig. 4040. V7, 
Hyoseris minima L. Sp. Pl. 879. 1753. Mi 


Arnoseris pusilla Gaertn. Fr. & Sem. 2: 355. 1791. 
Arnoseris minima Dumort. FI. Belg. 63. 1827. 


Scapes slender, 3’-12’ high, leafless, simple, i] Nii 
or with 1-4 branches mostly above the middle, ANG 
gradually thickened and hollow upward for a space | i 
of an inch or more below the heads. Leaves 
oblanceolate, obovate, or oblong, 1-3’ long, 3’—10” 
wide, usually coarsely and sharply toothed, nar- 
rowed into margined petioles; heads 8” broad, or 
less; bracts of the involucre linear-lanceolate, 
acuminate, strongly keeled after flowering, 2’-4 
long, curving over the achenes. 

Fields and waste grounds, Maine to Ontario and 


Michigan. Adventive from Europe. Also called dwarf 
swine’s- or hog’s-succory ; dwarf nipplewort. Summer. 


8. HYPOCHAERIS [Vaill.] L. Sp. 
Bi Sie; .1753- ' 


Mostly perennial herbs, with scapose, often 
branched stems and mostly basal tufted leaves, 
pinnatifid to entire, those of the scapes few, scale-like, very small, or none. Heads mostly 
large, long-peduncled. Flowers yellow. Involucre oblong-cylindric to campanulate, its bracts 
herbaceous, imbricated in several series, appressed, the outer smaller. Receptacle flat, chaffy. 
Anthers sagittate. Style-branches slender, obtusish. Achenes oblong to linear, 10-ribbed, 
somewhat narrowed below, contracted above into a long or short beak, or the outer truncate. 
Pappus of I row of plumose bristles, sometimes with some shorter simple ones. [Greek, for 
pigs, which are fond of its roots.] 

5 pout 50 species, natives of Europe, Asia and South America. Type species: Hypochaeris 
glabra L. 


1. Hypochaeris radicata L. Long- 
rooted Cat’s-ear. Gosmore. 
Fig. 4050. 
Hypochaeris radicata L. Sp. Pl. 811. 1753. 

Perennial; stems several together, glabrous, 
slender, 1°-2° high, bracted, or rarely simple, 
bearing a few scales. Leaves spreading on the 
ground, oblanceolate to obovate in outline, 
pinnatifid-lobed to dentate, 2’-6’ long, hirsute 
on both sides; involucre oblong-cylindric, 
about 1’ high, its bracts glabrous, or sparingly 
pubescent; heads 1’ broad or more; chaff of 
the receptacle of narrow soft pointed scales; 
achenes rough, all with very slender beaks 
longer than the body; flowers longer than the 
involucre. 

In waste places, Ontario to New York, southern 
New Jersey and Pennsylvania. Also in Califor- 
nia, Oregon, Washington and Colorado. Jamaica. 
Adventive or naturalized from Europe. Native 
also of Asia. May—Oct. 

Hypochaeris glabra L., the smooth cat’s-ear, a 
smaller species, with nearly or quite glabrous 
leaves, flowers scarcely longer than the involucre, 

é and the outer achenes truncate, has been found 
in Maine and Ontario, and in Georgia, and is naturalized on the Pacific Coast. 


Q. - APARGIA Scopamt Cam: Ed) 2) as-rig. ‘1772. 


Perennial scapose herbs, with tufted basal, mostly pinnatifid leaves, branched and scaly, 
or simple and naked scapes, and large heads of yellow flowers, solitary at the end of the 
scape or of its branches. Involucre ovoid or oblong, its principal bracts in I or 2 series, 
nearly equal, with several series of short outer ones. Receptacle flat, fimbrillate, villous, or 
somewhat honeycombed. Rays truncate and 5-toothed at the apex. Anthers sagittate at the 
base. Style-branches slender. Achenes oblong or linear, finely striate, contracted or beaked 
at the summit. Pappus of 1 or 2 series of plumose persistent brownish bristles, somewhat 
broadened at the base, or the outer scale-like and simple. [Greek, from the growth of these 
plants on unused land.] 


310 CICHORIACEAE. Vou. Lit 


About 45 species, natives of the Old World. Type species: Apargia incana Scop. 


Scape scaly-bracted, mostly branched; pappus of plumose bristles only. 1. A. autumnalis. 
Scape bractless, monocephalous; pappus of two kinds. 
Pappus of marginal and inner flowers dissimilar. 2. A. nudicaulis. 
Pappus of all flowers alike. 3. A. hispida. 


1. Apargia autumnale (L.) Hoffm. Fall 
Dandelion. Autumnal Hawkbit. Lion’s- 
tooth. Fig. 4051. 


Leontodon autumnale L. Sp. Pl. 798. 1753. 


Apargia autumnalis Hoffm. Deutsch. Fl. Ed. 2, 2: 
113. 1800. 


Plant glabrous or nearly so, or the involucres 
and ends of peduncles black-pubescent; scape 
slender, usually branched and scaly, 62° high. 
Leaves narrowly oblong to linear-lanceolate, pin- 
natifid into narrow lobes, or some of them coarsely 
dentate, 3-8’ long, 3-12” wide, acuminate at the 
apex, narrowed into rather short petioles; heads 
several, rarely solitary, about 12’-15” broad; in- 
volucre oblong; achenes tapering into a short 
beak, or the outer ones nearly terete; pappus- 
bristles all plumose. 


In fields and along roadsides, Newfoundland to 
Ontario, New Jersey, Pennsylvania and Michigan. 
Naturalized from Europe. Native also of Asia. 
June-Nov. Dog-dandelion. Arnica-bud. 


2. Apargia nudicaillis (L.) Britton. Rough , 
or Hairy Hawkbit. Fig. 4052. 


Crepis nudicaulis L. Sp. Pl. 805. 1753. 
Leontodon hirtum L. Sp. Pl. Ed. 2, 1123. 1763. 
Leontodon nudicaule Banks; Lowe, Trans. Camb. 

Phil Soca: 28: 183i. 

Plant more or less hirsute; scape simple, slender, 
4-12’ high, minutely scaly, or naked. Leaves 
linear-oblong to narrowly spatulate, acute or 
obtuse, not acuminate, nearly entire, coarsely 
sinuate-dentate or sometimes pinnatifid, 2-5’ long, 
3-8” wide, narrowed into petioles; head solitary 
at the end of the scape, 5’—10” broad; involucre 
canescent or pubescent; outer achenes with a 
pappus of simple narrow scales, the inner ones 
with a pappus of plumose bristles. 

In ballast and waste places, Connecticut to New 
Jersey and Pennsylvania, and on Vancouver Island. 
Adventive from Europe. June-Oct. 


3. Apargia hispida (L.) Willd. Common 
Hawkbit. Fig. 4053. 
Leontodon hispidus L. Sp. Pl. 799. 1753. 
Leontodon hastile L. Sp. Pl. Ed. 2, 1123. 1763. 
A. hispida Willd. Sp. Pl. 3: 1552. 1804. 

Glabrous, or bristly-hispid. Scape stout or slender, 
4’-25' high, bractless, monocephalous; head nodding 
before flowering; leaves 2’-0’ long, oblong-lanceolate 
in outline, coarsely and sharply dentate or subpin- 
natifid; head 13’ broad or-less; involucre 6”-7” high; 


pappus an outer series of short and an inner series 
of long plumose bristles, alike in all the flowers. 


Fields and waste grounds, Rhode Island to Pennsyl- 
vania, Ontario and Ohio. July—Sept. 


GENUS Io. CHICORY FAMILY. 3I1 


10. PICRIS L. Sp. Pl. 792. 1753. 


Erect hispid, mostly branching, leafy herbs, with alternate leaves (in our species), and 
rather large, usually corymbose or paniculate heads of yellow flowers. Involucre campanulate 
or cup-shaped, its principal bracts in I series, nearly equal, with 2-3 series of small or large 
exterior spreading ones. Receptacle flat, short-fimbrillate. Rays truncate and 5-toothed at 
the apex. Anthers sagittate at the base. Style-branches slender. Achenes linear or oblong, 
somewhat incurved, terete or angled, 5-10-ribbed and transversely wrinkled, narrowed at the 
base and summit, or beaked in some species. Pappus of I or 2 series of slender plumose 
bristles. [Greek, bitter. ] 


About 35 species, natives of the Old World, one perhaps indigenous in Alaska. Type species: 
Picris aspienioides L. 


Outer involucral bracts linear; achenes not beaked. 1. P. hieracioides. 


Outer involucral bracts ovate, foliaceous ; achenes short-beaked. 2. P. echioides. 
Wrayrf 
Ragu 


Mx 
1. Picris hieracioides L. Hawkweed 7h 


ie 
Picris. Fig. 4054. } 


ip 
i 


Picris hieracioides L. Sp. Pl. 792. 1753. 


Biennial, more or less hispid, much 
branched, 1°-3° high. Leaves, lanceolate 
or oblong-lanceolate, dentate, or nearly en- 
tire, the basal narrowed into petioles, acute, 
often 6’ long, those of the stem mostly 
sessile and smaller; heads numerous, 3’-1’ 
broad; involucre 4’-6” high, its outer 
bracts linear, subulate, spreading, the inner 
linear-lanceolate, acuminate ; pappus copious, 
nearly white. 


In waste places, Illinois, Pennsylvania, New 
Jersey, and in ballast about the seaports. 
Adventive from Europe. Native also of Asia. 
nor bitter. June-Sept. Old name lang-de- 
beef. 


3 
5 


2. Picris echioides L. Bristly Ox- 
tongue. Fig. 4055. 


Picris echinoides L. Sp. Pl. 792. 1753. 


Helmintha echinoides Gaertn. Fruct. & Sem. 2: 
368. 1802. 


Annual or biennial, branched, hispid; stem 
about 22° high. Basal and lower leaves spatu- 
late or oblong, obtuse, repand-dentate, 2’-6’ 
long, narrowed into petioles; upper leaves 
sessile and clasping, oblong or lanceolate, 
smaller, the uppermost mainly acute and en- 
tire; heads numerous, rather crowded, short- 
peduncled, about 4’ broad; outer bracts of the 
involucre 4 or 5, foliaceous, ovate, acute, 
hispid-ciliate, the inner ones lanceolate, mem- 
branous; achenes beaked. 


In waste places, Nova Scotia and Ontario to 
Pennsylvania, and in ballast about the seaports. 
Also in California. Adventive from Europe. July- 
Sept. Called also bugloss and bugloss-picris. 


312 CICHORTACEAE: Vo. III. 


11. PTILORIA Raf. Atl. Journ. 145. 1832. 
[STEPHANOMERIA Nutt. Trans. Amer. Phil. Soc. (II.) 7: 427. 1841.] 

Annual or perennial, mostly glabrous, often glaucous herbs, with erect, simple or branched, 
usually rigid stems, alternate or basal, entire dentate or runcinate-pinnatifid leaves, those of 
the stem and branches often reduced to subulate scales, and small erect heads of pink flowers, 
paniculate, or solitary at the ends of the branches, opening in the morning. Involucre cylin- 
dric or oblong, its principal bracts few, equal, scarious-margined, slightly united at the base, 
with numerous short exterior ones and sometimes a few of intermediate length. Receptacle 
flat, naked. Anthers sagittate at the base. Style-branches slender. Achenes oblong or 
linear, terete or columnar, 5-ribbed, truncate or beaked at the summit, the ribs smooth or 
rugose. Pappus of 1 series of rather rigid plumose bristles. [Greek, referring to the 
feathery pappus. ] 


About. 20 species, natives of western and central North America. Type species: Ptiloria 
pauciflora (Torr.) Raf. 


Involucre about 5” high; pappus brownish, plumose to below the middle. 1. P. pauciflora. 
Involucre about 4” high; pappus white, plumose almost to the base. 2. P. ramosa. 
‘| ao) aN 2, SN a oP yee 
“I Y ) UN \\ ‘ A\ \\ veh 1. Ptiloria pauciflora (Torr.) Raf. 
NY) p AY } rom VY VY Brown-plumed Ptiloria. Fig. 4056. 

\ \ i 7 4 yf ) ey i q I . \/ 

¥ \ He VY YG if y Y , Prenanthes? pauciflora Torr. Ann. Lyc. N. Y. 

‘ \ if Ji 2): 210. 1027. 


Ptiloria pauciflora Raf. Atl. Journ. 145. 1832. 


Stephanomeria runcinata Nutt. Trans. Am. 

Phil. Soc. (II.) 7: 427. 1841. 

Perennial; stem rather stout, striate, 
rigid, divergently branched, 1°-2° high. 
Basal and lower leaves runcinate-pinnatifid, 
1-23’ long, 3-6” wide, the upper all short 
and narrowly linear or reduced to scales; 
heads somewhat racemose-paniculate along 
the branches, usually about 5-flowered; 
involucre 4-5” high; rays 1-2” long; 
pappus brownish, plumose to below the 
middle. 


Plains, Nebraska, Kansas to Wyoming, 
Texas and California. Summer. 


eal? 


2. Ptiloria ramdésa Rydb. White- 
plumed Ptiloria. Fig. 4057. 


Ptiloria ramosa Rydb. Mem. N. Y. Bot. Gard. 
I: 453. 1900. 


Similar to the preceding species, but com- 
monly lower, bushy-branched, the branches 
ascending. Basal leaves runcinate-pinnati- 
fid, those of the stem linear or filiform, 
entire, or sometimes runcinate-dentate, the 
uppermost reduced to small scales; heads 
numerous, usually solitary at the ends of 
the branchlets; involucre about 4” high; 
pappus bright white, very plumose to near 
the base. 


Plains and dry, rocky soil, western Ne- 
braska to Wyoming, Montana and Colorado. 
In first edition of this work not distinguished 
from the western Ptiloria tenuifolia (Torr.) 
Raf. May—Aug. 


GENUS 12. CHICORY FAMILY. 313 


12, TRAGOPOGON [Tourn.] L. Sp. Pl. 789. 1753. 


Biennial or perennial, erect usually branched, somewhat succulent herbs, with slender 
fleshy tap-roots, alternate entire linear-lanceolate long-acuminate leaves, clasping at the base, 
and long-peduncled large heads of yellow or purple flowers, opening in the early morning, 
usually closed by noon. Involucre cylindric or narrowly campanulate, its bracts in I series, 
nearly equal, acuminate, united at the base. Rays truncate and 5-toothed at the apex. Anthers 
sagittate at the base. Style-branches slender. Achenes linear, terete, or 5-angled, 5-10-ribbed, 
terminated by slender beaks, or the outermost beakless. Pappus-bristles in 1 series, plumose, 
connate at the base, the plume-branches interwebbed. [Greek, goats-beard.] 


About 35 species, natives of the Old World. Type species: Tragopogon pratensis L. 


Flowers yellow; involucral bracts equalling or shorter than the rays. 1. T. pratensis. 
Flowers purple; involucral bracts much longer than the rays. 2. T. porrifolius. 


\ 


\ 


| 


1. Tragopogon praténsis L. Yellow 
Goat’s-beard. Meadow Salsify. . 
Fig. 4058. 


Tragopogon pratensis L. Sp. Pl. 789. 1753: 


Stem branched, 14°-3° high. Leaves keeled, 
tapering from the broad, more or less clasping 
base to a very long acuminate tip, the lower 
sometimes 10’ long and 1’ wide; peduncles 
thickened at the top; heads 1-23’ broad; bracts 
of the involucre about 8, lanceolate, acuminate, 
shorter than or equalling (rarely exceeding) 
the yellow rays; marginal achenes striate, 
smooth or roughened. 


In fields and waste places, Nova Scotia to New 
Jersey, Ontario, Ohio, Manitoba and Colorado. 
Naturalized from Europe. June—Oct. Called also 
buck’s-beard, noon-flower, star-of-jerusalem, noon- 
tide, joseph’s flower, go-to-bed-at-noon. 


2. Tragopogon porrifolius L. Oyster 
Plant. Salsify. Purple Goat’s-beard. 
Fig. 4059. 
Tragopogon porrifolius L. Sp. Pl. 789. 1753. 


Taller, sometimes 43° high. Peduncles very 
much thickened and hollow for I to 3 inches 
below the heads; heads 2-4’ broad, very showy; 
bracts of the involucre linear-lanceolate, acumi- 
nate, usually much longer than the purple rays; 
achenes sometimes 2’ long, the outer ones covered 
with scale-like tubercles, especially on the ribs 
below. 


In fields and waste places, Ontario to New Jersey, 
Virginia, Minnesota, British Columbia, Nebraska and 
California, mostly escaped from gardens, where it is 
common. Native of Europe. Called also vegetable 
oyster, jerusalem-star, nap-at-noon, oyster-root. Nat- 
uralized as a weed on the Pacific Coast. The 
root is the familiar vegetable known as oyster-plant. 
An apparent hybrid between this and the preceding 
species has been noticed at New Brunswick, N. J. 
June—Oct. 


314 CICHORIACEAE. : Voi. III. 


13. MALACOTHRIX DC. Prodr. 7: 192. 1838. 


Annual or perennial, branching or scapose herbs, with alternate or basal, mostly pinnatifid 
leaves, and long-peduncled panicled or solitary heads of yellow or rarely white flowers. 
Involucre campanulate, its principal bracts in I or 2 series, equal or nearly so, with several 
series of shorter exterior ones. Receptacle flat, naked or bristly. Rays truncate and 5-toothed 
at the apex. Anthers sagittate at the base. Style-branches slender. Achenes oblong or 
linear, glabrous, 10-15-ribbed, 4 or 5 of the ribs usually more prominent than the others, 
truncate, or margined and 4-5-toothed at the summit. Pappus-bristles in 2 series, the inner 
naked or minutely serrulate, slender, coherent at the base and deciduous in a ring, the outer 
few (1-8), more persistent, or all deciduous in our species. [Greek, soft-hair, in allusion to 
the soft pappus.] 


About 15 species, natives of the western and southwestern United States and lower California. 
Type species: Malacothrix californica DC. 


1. Malacothrix sonchoides (Nutt.) T. & G. 
Malacothrix. Fig. 4060. 


Leptoseris sonchoides Nutt. Trans. Am. Phil. Soe. (II.) 7: 
439. 1841. 


Malacothrix sonchoides T. & G. Fl. N. A. 2: 486. 1843. 


Annual, glabrous throughout, or slightly glandular; 
stem branched, 6-12’ high. Leaves somewhat fleshy, 
oblong or linear-oblong in outline, pinnatifid and the 
lobes dentate with mucronate-pointed teeth, the basal 
ones 13-3 long, narrowed into short broad petioles, 
those of the stem smaller, sessile; heads several or 
numerous, 8-13” broad; principal bracts of the invo- 
lucre linear, acute, scarious-margined, the outer short, 
oblong, obtuse, or acutish; achenes_ linear-oblong, 
margined at the summit by a 15-denticulate white 
border; pappus-bristles all deciduous. 


On dry plains, western Nebraska and Kansas to Cali- 
fornia and Arizona. May—Aug. 


14. CHONDRILLA [ Tourn: | L.Sp. PL 7o6. 1753: 


Perennial herbs, with stiff divaricately branched stems, the basal leaves large and mostly 
pinnatifid, those of the stem small, narrow, alternate, and few middle sized heads of yellow 
flowers mostly solitary at the ends of the branches. Involucre cylindric, several-flowered, 
its inner bracts in I or 2 series, nearly equal, with several series of small or minute outer 
ones. Receptacle flat, naked. Rays truncate and 5-toothed at the summit. Anthers sagittate 
at the base. Style-branches slender. Achenes oblong or linear, 4-5-angled, many-ribbed, 
more or less spiny near the summit, abruptly contracted into a beak. Pappus of copious 
soft white simple bristles. [Greek, lump, from the gummy matter borne on the stems of 


some species. | \\ Wy 


About 18 species, natives of the Old World, the fol- \\ 1) 
lowing typical. NY 


1. Chondrilla jancea L. Gum Succory. 
Fig. 4061. 


Chondrilla juncea L. Sp. Pl. 796. 1753. 


Stem rush-like, hirsute at the base, glabrous above, 
much branched, 1°-3° high. Basal leaves runcinate- 
pinnatifid, those of the stem linear or linear-lanceolate, 
acute, dentate or entire, sessile, 4-13’ long, 3’-13” 
wide; heads terminal and lateral on the branches, 
short-peduncled or sessile, 4-6” broad; involucre 
glabrous or nearly so, about 4” high, its inner bracts 
narrowly linear; achenes muricate and spiny near 
the summit, slightly shorter than the filiform beak. 


In dry fields and waste places, Delaware to Maryland 
and Virginia. Naturalized from Europe. July—Aug. 
Naked-weed. Skeleton-weed. Devil’s-grass. Hog-bite. 


GENUS I5. CHICORY FAMILY. 315 


15. LEONTODON L. Sp. Pl. 798. 1753. 
[TarAxacum (Hall.) Ludwig, Def. 175. 1760.] 


Perennial acaulescent herbs, with basal tufted pinnatifid or sinuate-dentate leaves, and 
large heads of yellow flowers, solitary, or very rarely 2 or 3 together at the ends of naked 
hollow scapes. Involucre oblong or campanulate, its inner bracts in I series, nearly equal, 
slightly united at the base, the outer of several series of shorter somewhat spreading ones, 
often reflexed at maturity. Receptacle flat, naked. Rays truncate and 5-toothed at the summit. 
Anthers sagittate at the base. Style-branches slender, obtusish. Achenes oblong or linear- 
fusiform, 4-5-angled, 5-10-nerved, roughened or spinulose, at least above, tapering into a 
very slender beak. Pappus of numerous filiform unequal simple persistent bristles. [Greek, 
lion’s-tooth. ] 

About 20 species, natives of the northern hemisphere and southern South America. Type 
species: Leontodon Taraxacum L. 


Outer involucral bracts reflexed ; achenes greenish brown, the beak 2-3 times their length. 
1. L. Taraxacum. 
Outer involucral bracts spreading or ascending. 
Achenes greenish, the beak 2-3 times their length. 2. L. latilobum. 
Achenes red, the beak not more than twice their length. 3. L. erythrospermum. 


1. Leontodon Taraxacum L. Dandelion. Blowball. Fig. 4062. 


Leontodon Taraxacum L. Sp. Pl. 798. 1753. 

-T. officinale Weber, Prim. Pl. Holst. 56. 1780. 

T. Dens-leonis Desf. Fl. Atlant. 2: 228. 1800. 

T. Taraxacum Karst. Deutsch. Fl. 1138. 1880-83. 


Root thick, deep, often 10’ long, bitter. Leaves Y 
oblong to spatulate in outline, usually pubes- 
cent, at least when young, acute or obtuse, 
pinnatifid, sinuate-dentate or rarely nearly 
entire, rather succulent, 3-10’ long, 4’-23’ wide, 
narrowed into petioles; scape erect, 2’-18’ high; 
head 17-2’ broad; containing very numerous 
golden-yellow flowers (150-200), inner bracts 
of the involucre linear or linear-lanceolate, the 
outer similar, shorter, not glaucous, reflexed, 
all acute; achenes greenish-brown, fusiform, 
spinulose above, narrowed into a filiform beak 
2-3 times their length, which support the 
copious white pappus, the fruiting mass of 
which becomes globose when ripe. 


NG 


} 


Yi) 


In fields and waste places, naturalized as a 
weed from Europe. Also in Asia and distributed 
as a weed in all civilized parts of the world. Jan.— 
Dec. Called also lion’s-tooth, cankerwort, milk- 
witch- or yellow-gowan, Irish daisy, monk’s-head, 
priest’s-crown, puff-ball. Arnica. 


2. Leontodon latilobum (DC.) Britton. 
Mountain Dandelion. Fig. 4063. 


Taraxacum latilobum DC. Prodr. 7: 146. 1838. 


T. Taraxacum alpinum Porter, Mem. Torr. Club 
5: 349. 18094. 


Similar to the preceding but scape lower, 
2’-7’ high. Leaves sinuately lobed with broadly 
triangular lobes, but less deeply so than in 
L. Taraxacum and rarely as far as half way 
to the midrib, or often merely dentate or sub- 
entire; heads smaller, or about 1’ wide; bracts 
fewer, the outer broadly ovate, appressed or 
merely spreading. 


In moist places, Newfoundland and Labrador 
to Quebec. Europe. Summer. Rocky Mountain 
and northwestern plants formerly regarded as 
included in this species are now considered dis- 
tinct. 


316 CICHORIACEAE. Vor. III. 


3. Leontodon erythrospérmum (Andrz.) Britton. Red-seeded Dandelion. 
Fig. 4064. 
ROOM, 


x Si (NAR? 
eS 


Taraxacum erythrospermum Andrz. in Bess. Enum. 
Pi Valhe7 see tects 


Similar to the two preceding species, the leaves 
glabrous, very deeply runcinate-pinnatifid or pin- 
nately divided into narrower triangular-lanceolate 
usually long-pointed segments; scapes glabrous, 
or pubescent above; heads rarely more than 1’ 
broad, 70-90-flowered; bracts of the involucre 
glaucous, the outer lanceolate, spreading or as- 
cending, the inner linear, longer, each usually 
with an appendage just below the tip; flowers 
sulphur-yellow, the outer rays purplish without; 
achenes narrower, bright red, or red-brown, spin- 
ulose above, the filiform beak not more than twice 
their length; pappus dirty white. 


In fields and woods, Maine and Vermont to New 
York, Pennsylvania, Ontario, Alberta, Illinois and 
Wyoming. Naturalized from Europe. April-June. 
April-June. 


16. SONCHUS [Tourn.] L. Sp. Pl. 793. 1753. 


Annual or perennial succulent herbs, with alternate, mostly auriculate-clasping, entire 
dentate lobed or pinnatifid, prickly-margined leaves, and large or middle-sized, peduncled 
corymbose or paniculate heads of yellow flowers. Involucre ovoid or campanulate, usually 
becoming thickened and more or less conic at the base when old, its bracts herbaceous or 
membranous, imbricated in several series, the outer successively smaller. Receptacle flat, 
naked. Rays truncate and 5-toothed at the apex. Anthers sagittate at the base. Style- 
branches slender. Achenes oval, oblong, or linear, more or less flattened, 10-20-ribbed, 
somewhat narrowed at the base, truncate. Pappus of very copious soft white simple capillary 
bristles, usually falling away connected, sometimes with I or 2 stouter ones which fall sepa- 
rately. [The Greek name of the Sow-thistle.] 


About 45 species, natives of the Old World. Besides the following, another occurs on the 
Pacific Coast. Type species: Sonchus oleraceus L. 


Involucre glandular-pubescent ; heads nearly 1’ high. 1. S. arvensis. 
Involucre glabrous; heads 6”’—8” high. 5 
Auricles of the leaves acute; achenes striate and transversely wrinkled. 2. S. oleraceus. 
Auricles rounded; achenes ribbed, not transversely wrinkled. 3. S. asper. 


1. Sonchus arvénsis L. Corn Sow-Thistle. 
Milk Thistle. Fig. 4065. 


Sonchus arvensis L. Sp. Pl. 793. 1753. 


Perennial by deep roots and creeping rootstocks, 
glabrous; stem leafy below, paniculately branched 
and nearly naked above, 2°-4° high. Lower and 
basal leaves runcinate-pinnatifid, often 12’ long, 
spinulose-dentate, narrowed into short petioles, the 
upper pinnatifid or entire, lanceolate, clasping; heads 
several or numerous, corymbose-paniculate, 1’—2’ 
broad, bright yellow, very showy; involucre nearly 
1’ high, its bracts as also the peduncles glandular- 
bristly; achenes oblong, compressed, with about 10 
rugose longitudinal ribs. 


In low grounds, fields and waste places, New Jersey 
to Quebec, Newfoundland, Minnesota, Colorado and 
British Columbia. Naturalized from Europe. Native 
also of Asia. July—Oct. Dindle. Gutweed. Swine-thistle. 
Tree sow-thistle. 


GENUS I6. CHICORY FAMILY. 317 


2. Sonchus oleraceus L. Annual Sow- 
Thistle. Hare’s Lettuce. Fig. 4066. 


Sonchus oleraceus L. Sp. Pl. 794. 1753. 


3 
t (f ) 


jks Sy fi 


U 
Annual, with fibrous roots; stem leafy be- \ 
low, nearly simple, 1°-10° high. Basal and 
lower leaves petioled, lyrate-pinnatifid, 4’—10’ 
long, the terminal segment commonly large 
and triangular, the margins denticulate with 
mucronate or scarcely spiny teeth; upper leaves 
pinnatifid, clasping by an auricled or sagittate 
base, the auricles acute or acuminate; upper- 
most leaves often lanceolate and entire; heads 
several or numerous, pale yellow, 9-15” 
broad; involucre glabrous, 6-8” high; achenes 
flat, longitudinally ribbed and _ transversely 
rugose. 


\ 


is ae “ } ! 


In fields and waste places, a common weed in 
most cultivated parts of the globe except the ex- 
treme north. Also in Central and South America. 
Naturalized from Europe. MHare’s-colewort or 
-thistle. Milk-weed. Miulk-thistle. Milky tassel. 
Swinies. The leaves used as a salad and as a pot herb. May-—Noy. 


3. Sonchus asper (L.) Hill. Spiny or Sharp- 
fringed Sow-Thistle. Fig. 4067. 


Sonchus oleraceus var. asper L. Sp. Pl. 794. 1753. 
Sonchus asper Hill, Herb. Brit. 47. 1769. 


Annual, similar to the preceding species; leaves 
undivided, lobed or sometimes pinnatifid, spinulose- 
dentate to spinulose-denticulate, the lower and basal 
ones obovate or spatulate, petioled, the upper oblong 
or lanceolate, clasping by an auricled base, the 
auricles rounded; heads several or numerous, 1’ 
broad or less; flowers pale yellow; involucre gla- 
brous, about 6” high; achenes flat, longitudinally 
ribbed. 


In waste places throughout most of our area and in 
tropical and South America. Widely distributed as a 
weed in nearly all cultivated parts of the earth. Nat- 
uralized from Europe. May—Novy. 


17. LACTUCA [Rominieee Sp. Pl. vos. 1753. 


Tall leafy herbs, with small panicled heads of yellow, white, pink, or blue flowers, and 
alternate leaves. Involucre cylindric, its bracts imbricated in several series, the outer shorter. 
or of I or 2 series of principal nearly equal inner bracts, and several rows of short outer ones. 
Receptacle flat, naked. Rays truncate and 5-toothed at the summit. Anthers sagittate at 
the base. Style-branches mostly slender. Achenes oval, oblong or linear, flat, 3-5-ribbed 
on each face, narrowed above or tipped by a filiform beak, which is somewhat expanded 
at the summit into a small disk bearing the copious soft capillary white or brown pappus- 
bristles. [The Ancient Latin name, from Jac, milk, referring to the milky juice.] 

About 100 species, natives of the northern hemisphere. Type species: Lactuca sativa L. 

A. Achenes filiform-beaked; rays mainly yellow. 
Introduced European species; heads few-flowered. 


Panicle widely branching; achene about as long as its beak. 1. L. virosa. 

Panicle-branches nearly erect ; achene shorter than its beak. 2. L. saligna. 
Native species; heads several- to many-flowered. 

Leaves, or their lobes, spinulose-denticulate ; stem leafy below. 3. L. ludoviciana. 


Leaves entire to pinnatifid, the teeth not spinulose; stem leafy to the inflorescence. 
Leaves hirsute or bristly on the veins beneath. 
Plant hirsute below; leaves pinnatifid or the upper entire. 4. L. hirsuta. 
Plant hirsute up to the inflorescence; leaves merely dentate or denticulate. 
5. L. Steelei. 


318 CICHORIACEAE Vor. III. 


Leaves glabrous. 
Leaves entire to pinnatifid, not sagittate at base. 6. L. canadensis. 
Leaves entire or denticulate, sagittate at base. 7. L. sagittifolia. 


B. Achenes truncate, or narrowed into stout beaks; rays blue to white. 


Perennial ; achenes flattened. ’ 8. L. pulchella, 
Annuals or biennials ; achenes swollen. 
Pappus bright white. 


Achenes beakless; leaves dentate, acuminate. g. L. villosa. 
Achenes beaked ; leaves pinnatifid. 10. L. floridana. 
Pappus brown; achenes short-beaked. 11. L. spicata. 
r yee ty 1. Lactuca virdsa L. Prickly or Wild 
wats gus 
ie mM ff Lettuce. Fig. 4068. 
oA \ ao 
Z, NY i} tt, Lactuca virosa L. Sp. Pl. 795. 1753. 


Lactuca Serriola L. Cent. Pl. 2: 29. 1756. 
Lactuca Scariola L. Sp. Pl. Ed. 2, 1119. 1763. 


Biennial, green and glaucous; stem stiff, leafy, 
usually paniculately branched, glabrous through- 


is 
<HN mae out, or bristly-hirsute at the base, 2°-7° high. 
4g Leaves oblong or oblong-lanceolate, spinulose- 
c margined, denticulate or pinnatifid, sessile or 
Z auriculate-clasping, the lowest sometimes 10° 
SC BAY long and 3’ wide, the upper much smaller; 
af ME midrib spinulose or hispid; heads 2’—4” broad, 
\ 


6-12-flowered ; very numerous in an open pan- 
icle; involucre cylindric, 1-14” thick, its outer 
bracts about one-third the nee of the inner; 
rays yellow; achenes obovate-oblong, about as 


long as the filiform beak; pappus white. 


In fields and waste places, Vermont to Georgia, 
west to the Pacific Coast. A troublesome weed. 
Naturalized from Europe. Consists of several 
races, differing in the form, lobing and position of the leaves. Aug.—Sept. Milk- or horse-thistle. 


lw 


2. Lactuca saligna L. Willow Let- 
tuce. Fig. 4069. 


Lactuca saligna L. Sp. Pl. 796. 1753. 


Biennial, light green ; stem slender, smooth 
or nearly so, 2° high or more, the very 
slender branches erect-ascending. Leaves 
glabrous, linear to oblong, 6’ long or less, 
runcinate-pinnatifid with pointed divergent 
lobes, or entire, sagittate at the base, the 
midvein sometimes spinulose; panicles nar- 
row; heads about 3” broad, few-flowered ; 
involucre subcylindric; outer bracts much 
shorter than the inner; achenes oblong, 
about one-half as long as the filiform beak; 
pappus white. 


Waste and cultivated grounds, Ohio. Nat- 
uralized from Europe. July—Aug. 


GENUS I7. CHICORY FAMILY. 


3. Lactuca ludoviciana ( Nutt.) DC. West- - 
ern Lettuce. Fig. 4070. 


Sonchus ludovicianus Nutt. Gen. 2: 125. 1818. ye \ 


Lactuca ludoviciana DC. Prodr. 7: 141. 838. 


Biennial, glabrous throughout, leafy uptoinflor- 4) 
escence, paniculately branched, 2°+5° high. Leaves Hi) 
oblong to ovate-oblong, acute or acutish, 2-4’ long, 4 Hy 
auriculate-clasping, spinulose-denticulate, sinuate- 
lobed, or pinnatifid with spinulose segments; heads ; \ 
3’-5”’ broad, numerous in an open panicle, their B: 4 


wa 
peduncles bracteolate; involucre cylindric or ovoid- y sae 
cylindric, glabrous, 8’—9” high, its bracts succes- 2 Z 
sively shorter and broader, the lower ones ovate; ‘ — so F 
rays yellow; achenes oval to obovate, flat, about 


» 
the length of their filiform beak; pappus white. 7 : 
| A bse ea\ 
Plains and banks, Iowa, Minnesota and South Da- 3 
kota to Kansas and.Texas. July—Sept. boy 


4. Lactuca hirsuta Muhl. Hairy or Red 
Wood-Lettuce. Fig. 4071. 


Lactuca hirsuta Muhl. Cat. 69. 1813. 

Lactuca sanguinea Bigel. Fl. Bost. Ed. 2, 287. 1824. 

Lactuca elongata var. sanguinea T. & G. Fl. N. A. 2: 
496. 1843. 


Stem 1°-6° high, naked and paniculately branch- 
ed above, usually hirsute, at least below. Leaves, 
or most of them, sinuate-pinnatifid, pubescent on 
both sides, or on the midrib beneath, those of the 
stem mostly sessile or auriculate-clasping, 3’-7’ 
long, the uppermost sometimes lanceolate and 
entire, the basal petioled; heads numerous, 2’—3” 
broad; involucre glabrous, 5’—9” high; outer bracts 
shorter than the inner; rays reddish-yellow or 
paler; achenes oblong-oval, flat, about the length 
of the beak; pappus white. 


In dry soil, Quebec to Ontario, Minnesota, Ala- 
bama and Texas. Stem, peduncles and involucre 
often red or purple. July—Sept. 


5. Lactuca Steélei Britton. Steele’s Wild 
ettuce. Fig. 4072: 


L. Steelei Britton, Man. 899. 1901. 


Stem and branches hirsute. Leaves obovate to oval, 
the larger about 8’ long and 4’ wide, sessile, irregu- 
larly dentate, not lobed, hispid on the veins beneath, 
short-acuminate; involucre about 6” high; rays not 
seen; achenes oval, 3-ribbed, about 14” long, the fili- 
form beak slightly shorter; pappus bright white. 


Near Washington, D. C., and, apparently, in Delaware. 
July—Aug. 


320 CICHORIACEAE. Vo. III. 


6. Lactuca canadénsis L. Wild or Tall Lettuce. Wild Opium. Fig. 4073. 
Lactuca canadensis L. Sp. Pl. 796. 1753. 
= \\ | ) Lactuca elongata Muhl.; Willd. Sp. Pl. 1525. 1804. 
Lactuca canadensis montana Britton, in Britton and 
Brown, Ill. Fl. 3: 274. 1808. 


- 
x 
XN 


WY W Pa 


yy, 


H NY; GSE 
I & 
Biennial or annual, glabrous throughout, some- 


what glaucous; stem leafy up to the inflorescence, 
2 we 3°-10° high, branching above into a narrow pan- 
Wa V\ icle. Leaves mostly sinuate-pinnatifid, those of 
q \ the stem sessile or auriculate-clasping, 2’-8’ long, 


the uppermost smaller, often lanceolate, acumi- 
nate and entire, sometimes all lanceolate and en- 
tire, the basal often 12’ long, narrowed into peti- 
oles; heads numerous, 2’—-3” broad; involucre 
cylindric, 4’-6” high, its outer bracts shorter than 
the inner; rays yellow; achenes oval, flat, about 
as long as the filiform beak; pappus white. 

In moist, open places, Nova Scotia to British Co- 
lumbia, Georgia, Alabama, Louisiana, Arkansas and 
Colorado. Santo Domingo. Trumpetweed, trumpet- 
milkweed, trumpets, and fire-weed. Butter- or horse- 
weed. Devil’s-weed. Devil’s-iron-weed. June—Novy. 

Lactuca Morssii Robinson, with blue rays, and 
achenes 3 or 4 times longer than their filiform beaks, 
may be a hybrid with L. spicata. 


7. Lactuca sagittifolia Eli. Arrow-leaved 
Lettuce. Fig. 4074. 

L. sagittifolia Ell. Bot. S. C. & Ga. 2: 253. 1821-24. 
Lactuca integrifolia Bigel. Fl. Bost. Ed. 2, 287. 1824. 

Not Nutt. 1818. 
L. elongata var. integ. T.& G. Fl. N. A. 2: 496. 1843. 

Biennial; stem glabrous throughout, or hirsute 
below, leafy nearly up to the usually paniculate 
inflorescence, 2°-6° high. Leaves oblong, oblong- 
lanceolate or lanceolate, acuminate or acute, en- 
tire or denticulate, the lower rarely pinnatifid, 
sometimes spinulose on the margins, those of the 
stem sessile or sagittate-clasping, 3-6’ long, 2’—12’ 
wide, the basal and lower ones petioled; heads 
commonly very numerous, 2’-4” broad; involucre 
cylindric, 5’-7” high, the outer bracts shorter than 
the inner; rays yellow or reddish; achenes oval, 
flat, rather longer than their filiform beaks; pap- 
pus white. 

In dry, open soil, New Brunswick and Ontario to 
Georgia, Idaho and Kansas. Called also devil’s-iron- 
weed. July—Sept. 


\\ 7 = 
\ \ Wi 7 l{] 8. Lactuca pulchélla (Pursh) DC. Large- 


\\a NZ 
\ yp WA fi} flowered Blue Lettuce. Fig. 4075. 
Wig Wy Sonchus pulchellus Pursh, Fl. Am. Sept. 502. 1814. 
Wf ZT Lactuca integrifolia Nutt. Gen. 2: 124. 1818. 
> J, Za Lactuca pulchella DC. Prodr. 7: 134. 1838. 
Pe | Perennial, glabrous throughout, somewhat glau- 
YIN ! ; — i cous; stem rather slender, leafy up to the corym- 
| SS \ bose-paniculate inflorescence, 1°-3° high. Leaves 
\ linear-lanceolate, lanceolate or oblong, acute, en- 
tire, dentate, lobed or pinnatifid, those of the stem 
sessile or partly clasping, 2’-8’ long; 2’-18” wide, 
the lowest and basal ones sometimes petioled; 
heads mostly numerous, 6-10” broad; branches 
! and peduncles scaly; involucre well imbricated, 
8-10” high, its outer bracts successively shorter, 
ovate-lanceolate; rays bright blue or violet; 
achenes oblong-lanceolate, flat, twice as long as 
their tapering beaks; pappus white. 
In moist soil, western Ontario to British Columbia, 
Michigan, Iowa, Kansas, New Mexico and California. 
June—Sept. 
Lactuca campéstris Greene, of the prairie region, 
is described as differing from this by yellow rays. 


WH] 
iI 


GENUs 17. CHICORY FAMILY. 321 


g. Lactuca villosa Jacq. MHairy-veined 
Blue Lettuce. Fig. 4076. 

L. villosa Jacq. Hort. Schoen. 3: 62. pl. 367. 1708. 
Sonchus acuminatus Willd. Sp. Pl. 3: 1521. 1804. 
Mulgedium acuminatum DC. Prodr. 7: 249. 1838. 
L. acuminata A. Gray, Proc. Am. Acad. 19: 73. 1883. 

Annual or biennial; stem glabrous, leafy up to 
the paniculate inflorescence, 2°-6° high. Leaves 
oblong, ovate or. lanceolate, acuminate, acutely den- 
tate or the teeth mucronate-tipped, glabrous above, 
pubescent with short stiff hairs on the veins be- 
neath, sessile and slightly clasping at the base, or A 
petioled, 4’-6’ long, 1-23’ wide, the lowest some- SS 
times lobed at the base; heads numerous, 3’—5” 
broad; peduncles usually minutely scaly; rays 
blue; involucre about 5” high, its outer bracts 
much shorter than the inner, some or all of them 
obtuse; achenes thick, oblong, little flattened, nar- 
rowed above; pappus white. 

In thickets, New York to Illinois, Nebraska, south 
iS Florida, Georgia and Kentucky. July-Sept. False 
ettuce. 


10. Lactuca floridana (L.) Gaertn. False or 
Florida Lettuce. Fig. 4077. 

Sonchus floridanus L. Sp. Pl. 794. 1753. 
Lactuca floridana Gaertn. Fruct. & Sem. 2: 362. 17091. 
Mulgedium floridanum DC. Prodr. 7: 349. 1791. 

Annual or biennial; stem glabrous, rather stout, 
leafy up to the large, paniculate inflorescence, 3°-7° 
high. Leaves deeply lyrate-pinnatifid, or sometimes 
cordate-ovate, sessile or petioled, 4-12’ long, glabrous 
above, pubescent on the veins beneath, the terminal 
segment usually broad, triangular, acute or acumi- 
nate, the lateral ones lanceolate to oval, acute, all 
usually dentate, or the leaves irregularly lobed; 
heads numerous, 3-5” broad; peduncles commonly 
scaly; rays blue; involucre about 6” high, its outer 
bracts much smaller than the inner; achenes thick, 
somewhat compressed, narrowed above into a short 
beak; pappus white. 

In moist, open places, southern New York and Penn- 
sylvania to Illinois, Kansas, Florida and Texas. Porto 
Rico. July—Sept. 


11. Lactuca spicata (Lam.) Hitchce. WP 
Tall Blue Lettuce. Fig. 4078. iy 


Sonchus spicatus Lam. Encycl. 3: 401. 1789. 
Mulgedium leucophaeum DC. Prodr.7: 250. 1838. 
Lactuca leucophaea var. integrifolia A. Gray, 

Sys Blox: Part 2, 444. 1884. 

Lactuca spicata integrifolia Britton, Mem. Torr. 

Club 5: 350. 1894. 

IES See Hitche. ; Britt. & Brown, Il. Fl. 3: 276. 

1898. 

Annual or biennial; stem usually stout, gla- 
brous, 3°-12° high, leafy up to the large, 
rather dense panicle. Leaves deeply pinnatifid 
or lobed to entire, sharply dentate with mu- 
cronate-pointed teeth, sessile, or the lower 
narrowed into margined petioles, glabrous on 
both sides, or pubescent on the veins beneath, 
5-12’ long, 2’-6’ wide; heads very numerous, 
about 2” broad; peduncles minutely scaly; 
tays blue to white; achenes oblong, com- 
pressed, narrowed above into a short neck; 
pappus brown. 

In moist soil, Newfoundland to Manitoba, 
North Carolina, Tennessee, Iowa, South Dakota 
and Colorado. Ascends to 2000 ft. in North 
Carolina. Races differ in leaf-form and in color 
of the flowers. Milk-weed. July—Oct. 


21 ‘XX 


322 CICHORIACEAE. Vor. III. 


18. LYGODESMIA D. Don, Edinb. Phil. Journ. 6: 311. 1829. 


Perennial or annual glabrous rigid branching herbs, with linear leaves, or the basal and 
lower ones sometimes broader and pinnatifid, those of the stem very narrow and entire or 
reduced to linear scales, and middle-sized 3-12-flowered heads of pink or purple flowers, 
solitary and erect at the ends of the stem and branches, or sometimes racemose. Involucre 
cylindric, its principal bracts 5-8, linear, scarious-margined, equal, slightly united at the base, 
with several very short outer ones. Receptacle flat, naked. Rays truncate and 5-toothed at 
the apex. Anthers sagittate at the base. Style-branches slender. Achenes linear, smooth 
or striate. Pappus of copious somewhat unequal simple bristles. [Greek, twig-bundle, from 
the numerous branches. ] ‘ 

About 6 species, natives of western and southern North America. Type species: Prenanthes 
juncea Pursh. 


Heads solitary at the ends of the branches; leaves linear or subulate. 1. L. juncea. 
Heads racemose along the branches; leaves elongated-linear. 2. L. rostrata. 


1. Lygodesmia jancea (Pursh) D. 
Don. Rush-like Lygodesmia. 


Fig. 4079. 


Prenanthes juncea Pursh, Fl. Am. Sept. 498. 
1814. 


Lygodesmia juncea D. Don; Hook. Fl. Bor. 
AMESTeS205., 1S23- ‘ 


Perennial by a thick woody root; stems 
stiff, striate, much branched, 8-18’ high, 
the branches erect. Lower leaves linear- 
lanceolate, rigid, entire, acute or acuminate, 
4’-2’ long, 3’”-1%” wide, the upper similar 
but smaller, or reduced to subulate scales; 
heads 6-8” broad, mostly 5-flowered, soli- 
tary at the ends of the branches; involucre 
6-8” high; achenes narrowly columnar or 
slightly tapering, truncate at the summit, 
about 8-nerved or ribbed, 23-33” long; 
pappus light brown. 


Plains, Minnesota to Saskatchewan, Mon- 
tana, Wisconsin, Missouri, Nebraska, Kansas 
and Arizona. Often infested by a globose gall 
2”-5” in diameter. June—Aug. 


2. Lygodesmia rostrata A. Gray. 
Beaked Lygodesmia. Fig. 4080. 


L. juncea var. rostrata A. Gray, Proc. Phil. 
Acad. 1863: 69. 1863. 


Lygodesmia rostrata A. Gray, Proc. Am. Acad. 
Oi 2172 1874: 


Annual, less rigid; stem striate, leafy, 
paniculately branched, 1°-3° high. Leaves 
elongated-linear, acuminate, entire, 3-nerved, 
the lower 3-7’ long, 1-14” wide, the upper- 
most very small and subulate; heads numer- 
ous, 7-10-flowered, about 3’ broad, racemose 
along the branches on scaly short erect pe- 
duncles; involucre 5-7” high; achenes nar- 
rowly fusiform, narrowed or somewhat 
beaked at the summit, 5-8-ribbed or -striate, 


4,7 


4’”-5” long, longer than the whitish pappus. 


Plains and canyons, South Dakota to Sas- 
katchewan, Nebraska, Kansas, Colorado and 
Wyoming. Aug.—Sept. 


GENUS IO. CHICORY FAMILY. (323 


19. AGOSERIS Raf.; D. Dietr. Syn. Pl. 4: 1332. 1847. 
[Troxrmon Nutt. Fras. Cat. 1813. Not Gaertn. 1791.] 


Perennial or annual herbs, mostly acaulescent, with tufted usually sessile basal leaves, 
and solitary heads of yellow or rarely purple flowers at the end of a naked or bracted scape. 
Involucre campanulate or oblong, its bracts imbricated in several series, appressed, or with 
spreading tips, membranous or herbaceous, not thickened after flowering, the outer ones 
gradually shorter and broader. Receptacle flat, naked or foveolate. Rays truncate and 
5-toothed at the apex. Anthers sagittate at the base. Style-branches slender. Achenes 
oblong, obovate, or linear, 10-ribbed, not flattened, beaked or beakless. Pappus of copious 
slender simple white bristles or soft unequal narrow scales. [Greek, head- or chief-succory.} 


About 30 species, natives of westefn and southern North America and southern South America. 
Besides the following, some 20 others occur in the western parts of the United States. Type 
species: Agoseris cuspidata (Pursh) D. Dietr. 


Achenes beaked. 


Head 1’—2’ broad, achenes 5”—6” long. 1. A. glauca. 
Head 17-2’ broad; achenes 5”—6” long. 2. A. parviflora. 
Achenes beakless. 3. A. cuspidata. 


1. Agoseris glauca (Pursh) D. Dietr. 
Large-flowered Agoseris. Fig. 4081. 


T. glaucum Pursh, Fl. Am. Sept. 505. 1814. 
Agoseris glauca D. Dietr. Syn. Pl. 4: 1332. 1847. 


Perennial, pale or glaucous, glabrous through- 
out or a little woolly below. Leaves linear, 
lanceolate, or oblong, entire, dentate or pinna- 
tifid, 2’-10’ long, 2’—10” wide, acuminate at 
the apex, narrowed at the base, sometimes into 
margined petioles; scapes stout, glabrous or 
slightly pubescent, longer than the leaves, often 
13° high; head 12’ broad; involucre oblong- 
campanulate, or broader in fruit, commonly 
quite glabrous, its bracts lanceolate, acuminate, 
often hyaline-margined ; achenes conspicuously 
beaked, 5-6” long, when mature longer than 
the copious pappus of rather rigid scabrous or 
denticulate bristles. 


Minnesota to South Dakota, Saskatchewan, 
south to Kansas (according to Smyth), Colorado 
and Utah. May-July. 


2. Agoseris parviflora (Nutt.) D. Dietr. 
Small-flowered Agoseris. Fig. 4082. 


Troximon parviflorum Nutt. Trans. Am. Phil. Soc. 7: 
434. 1841. 

Troximon glaucum var. parviflorum A. Gray, Syn. FI. 
PePatt 2; 437, 1884. 

Agoseris parviflora D. Dietr. Syn. Pl. 4: 1332. 1847. 


Perennial, glabrous throughout; scape slender, 
much longer than the leaves, 5’-15’ high. Leaves 
narrowly linear, acuminate, entire, 3-8’ long, 
1-23” wide; head 1’ broad or less; involucre ob- 
long-ovoid, becoming nearly hemispheric in fruit, 
6-8” high, glabrous, its bracts lanceolate, acumi- 
nate; achenes conspicuously beaked, about 4” long; 
pappus of numerous unequal very slender bristles. 


Plains, North Dakota to Nebraska, Manitoba, AI- 
berta, Idaho and New Mexico. Called also false dande- 
lion. May-July. 


324 CICHORIACEAE. . Vor. IIT. 


3. Agoseris cuspidata (Pursh) D. Dietr. 
Prairie False Dandelion. Fig. 4083. 


Troximon cuspidatum Pursh, Fl. Am.Sept.742. 1814. 
Troximon marginatum Nutt. Gen. 2: 128. 1818. 

Agoseris cuspidata D. Dietr. Syn. Pl. 4: 1332. 1847. 
Nothocalais cuspidata Greene, Bull. Cal. Acad. (I1.) 


2:55. 1886 

Leaves linear, long-acuminate, thick, pubescent 
or glabrate, 4’-8’ long, 2’-5”” wide, somewhat con- 
duplicate, their margins conspicuously white-to- 
mentose and crisped, or entire. Scape stout, to- 
mentose, at least above, shorter than or equalling 
the leaves; head 172’ broad; involucre usually 
quite glabrous, nearly 1’ high; achenes slightly 
contracted at the summit; about 3” long, beakless; 
pappus of 40-50 unequal bristles. 

In dry soil, on prairies and rocky hills, Illinois to 


North Dakota, Nebraska, Kansas and Colorado. 
Called also false dandelion. April-June. 


20. SITILIAS Raf. New Fl. N. A. 4:85. 1836. 
[PyrrHopappus DC. Prodr. 7: 144. , 1838.] : 


Annual or perennial herbs, with alternate or basal leaves, and mostly large, solitary or few 
heads of yellow flowers, borne on long, usually bracted peduncles. Involucre oblong or cam- 
panulate, its principal bracts in I series, nearly equal, slightly united at the base, with several 
series of smaller outer ones. Rays truncate and 5-toothed at the summit. Anthers sagittate 
at the base. Style-branches short, obtusish. Achenes oblong or fusiform, mostly 5-ribbed, 
roughened or hirsute, abruptly narrowed into a long filiform beak. Pappus of numerous 
soft simple brownish somewhat unequal bristles, surrounded at the base by a villous white 
ring. [Name unexplained.] 

Six known species, natives of North America and Mexico. Besides the following, 3 others 
occur in the southwestern United States. Type species: Sitilias caroliniana (Walt.) Raf. : 


Stem leafy, usually branched ; plant glabrous, or nearly so. 1. S. caroliniana. 
Scape naked, monocephalous ; plant hirsute, or pubescent. 2. S. grandiflora. 


1. Sitilias caroliniana (Walt.) Raf. Leafy-stemmed False Dandelion. Fig. 4084. 
Leontodon carolinianum Walt. Fl. Car. 192. 1788. 
S. caroliniana Raf. New FI. N. A. Part 4, 85. 1836. 
Pyrrhopappus carolinianus DC. Prodr. 7: 144. 1838. 


Annual or biennial; glabrous or nearly so; stem 
leafy, usually branched, 2°-5° high. Basal leaves 
oblong or oblong-lanceolate, pinnatifid, lobed, 
coarsely dentate or some of them entire, acute, 
acuminate, or obtusish, 3’-8’ long, 4-13’ wide, 
narrowed into margined petioles; stem leaves ses- 
sile or partly clasping, the upper usually lanceo- 
late, entire and acuminate; peduncles usually pu- 
berulent; heads 1 or several, 1-13’ broad; invo- 
lucre commonly puberulent or pubescent, about 1’ 
high, its outer bracts setaceous or subulate, spread- 
ing, the inner corniculate at the apex; achenes 
2-3” long, tipped with a filiform beak of about 
three times their length. 


In dry fields, Delaware to Florida, Kentucky, Mis- 
souri, Louisiana and Texas. April—July. 


GENUS 20. CHICORY FAMILY. 325 


2. Sitilias grandiflora ( Nutt.) Greene. 
Rough False Dandelion. Fig. 4085. 


Barkhausia grandiflora Nutt. Journ. Phila. Acad. 7: 
69. 1834. 


Pyrrhopappus scaposus DC. Prodr. 7: 144. 1838. 
Sitilias grandiflora Greene, Pittonia 2: 180. 1891. 


Hirsute or pubescent; root tuberous-thickened. 
Leaves all basal, oblong or spatulate in outline, 
deeply pinnatifid, 3-7’ long, 1-14’ wide, narrowed 
into margined petioles; scape naked or sometimes 
with a small leaf near its base; head solitary, 
1-2’ broad; outer bracts of the involucre small, 
short, subulate, the inner ones obscurely cornicu- 
late at the tip. 


On prairies, Kansas to Texas. April-June. 


at. CREPIS L. Sp. Pl. 805. 1753. 


Perennial, biennial or annual herbs, with alternate or basal, mostly toothed or pinnatifid 
leaves, and small or middle-sized heads, usually paniculate or corymbose, of yellow or orange 
flowers. Involucre cylindric, campanulate, or swollen at the base, its principal bracts in I 
series, equal, with a number of exterior smaller ones. Receptacle mostly flat, naked or short- 
fimbrillate. Rays truncate and 5-toothed at the apex. Anthers sagittate at the base. Style- 
branches slender. Achenes linear-oblong, 10-20-ribbed or nerved, not transversely rugose, 
narrowed at the base and apex, beakless in our species. Pappus copious, of very slender 
white bristles. [Greek, sandal; application not explained. ] 

About 200 species, natives of the northern hemisphere. Besides the following, about 20 others 
occur in western North America. Type species: Crepis tectorum L. 

Involucre glabrous. 
Involucre cylindric; plant pubescent below; introduced. 1. C. pulchra. 
Involucre campanulate; plant glaucous; native, western. 2. C. glauca. 
Involucre pubescent, glandular, or canescent. 
Foliage not canescent nor scurfy, sometimes hirsute. 
Stems naked, or with 1 or 2 small leaves; western, native. 
Stems leafy ; introduced annuals or biennials. 
Stem leaves narrow, revolute-margined, sessile. 
Stem leaves lanceolate, clasping, not revolute-margined. 


. C. runcinata. 


. C. tectorum. 


_ WwW 


Involucre 3’—4” high; achenes 1o-striate. 5. C. capillaris. 
Involucre 4”—6” high; achenes 13-striate. 6. C. biennis. 
Foliage cinereous, canescent, or scurfy, sometimes also hirsute. 
Inner bracts of the involucre 5-8; flowers 5-8. 7. C. intermedia. 
Inner bracts of the involucre 9-24; flowers 10-30. 8. C. occidentalis. 


1. Crepis pilchra L. Small-flowered Hawks- 
beard. Fig. 4086. 


Crepis pulchra L. Sp. Pl. 806. 1753. 


Annual; stem 2°-43° high, leafy and pubescent 
below, mostly glabrous, naked, and _ paniculately 
branched above. Stem leaves oblong or oblong- 
lanceolate, dentate, clasping by an auricled base or 
truncate, pubescent, 3-4’ long, acute or obtuse, or 
the uppermost lanceolate, acuminate and entire; 
basal leaves runcinate, narrowed into petioles; heads 
very numerous, about: 3” broad, in a large naked 
panicle; involucre narrow, cylindric, glabrous, about 
5” high, its principal bracts 12-15, linear, acuminate, 
the outer ones very short, ovate, appressed; achenes 
linear, faintly 1o-nerved, slightly narrowed above. 


Along railroad near Culpepper, Virginia, very abun- 
dant in 1890. Naturalized or adventive from Conti- 
nental Europe. May-July. 


326 CICHORIACEAE. Vor. III. 


: WWZ> 
4 er WZ YY hs 
VN w\ B 
\\ ITN ZB 


aN 


3. Crepis runcinata (James) T. & G. 
Naked Stemmed Hawksbeard. 


Fig. 4088. 


Hieracium runcinatum James in Long’s Exp. 1: 453. 


1823. 
Crepis runcinata T. & G. Fl. N. A. 2: 487. 


Crepis perplerans Rydb. Bull. Torr. Club 32: 134. 


1906. 


Perennial, similar to the preceding species, but 


2. Crepis glauca (Nutt.) T.&G. Glaucous 
Hawksbeard. Fig. 4087. 


Crepidium glaucum Nutt. Trans. Am. Phil. Soc. (II.) 
7: 436. 1841. 


Crepis glauca T.& G. Fl. N. A. 2: 488. 1843. 


Perennial; scapose, or rarely with I or 2 leaves 
on the stem, 1°-23° high, glabrous and glaucous 
throughout. Basal leaves spatulate, oblanceolate, 
or obovate, acute or obtuse at the apex, gradually 
narrowed into margined petioles, entire, dentate, 
or pinnatifid, 2’-6’ long, 4-1’ wide; heads not 
numerous, long-peduncled, 6-12” broad; pedun- 
cles glabrous; involucre campanulate, its principal 
bracts lanceolate, acute, the outer ones very short, 
ovate, appressed; achenes oblong-cylindric, slightly 
narrowed above, strongly 10-ribbed. 


In moist and saline soil, Nebraska to Saskatche- 
wan, Utah and Nevada. July—Aug. 


1843. 


not glaucous or scarcely so, often pubescent below ; 
stem leafless or with I or 2 small leaves, 1°-3° 
high. Basal leaves spatulate, obovate, or oblong, 
obtuse or acute, entire, repand, dentate, or rarely 
runcinate-pinnatifid, 2’-6’ long, 2’-12’ wide; heads 
several, long-peduncled, nearly 1’ broad; pedun- 
cles glabrous or glandular-pubescent; involucre 
campanulate, pubescent or glandular, its principal 
bracts linear-lanceolate, acute, outer ones short, 
lanceolate, appressed; achenes linear-oblong, some- 


what narrowed above, 10-ribbed. 


In moist soil, Iowa to North Dakota, 


Alberta, Utah and Montana. June—July. 


Crepis riparia A. Nelson, with broadly obovate 
basal leaves and larger flowers, is recorded from Ne- 


braska. 


Manitoba, 


4. Crepis tectorum L. Narrow-leaved Hawks- 
beard. Fig. 4089. 


Crepis tectorum L. Sp. Pl. 807. 1753. 


Annual: stem slender, puberulent or pubescent, 
leafy, branched, 1°-2° high. Basal leaves lanceolate, 
dentate, or runcinate-pinnatifid, 4-6’ long; stem 
leaves sessile, sometimes slightly sagittate at the 
base, linear, entire, dentate, or lobed, their margins 
revolute; heads numerous, corymbose, 6’—10” broad ; 
involucre narrowly campanulate, canescent or pubes- 
cent, 3’-5” high, its principal bracts lanceolate, acu- 
minate, downy within, the exterior ones linear, 
spreading; peduncles usually canescent; achenes I0- 
ribbed, narrowed above into a short beak, the ribs 
minutely scabrous. 


In waste places and on ballast, New York and New 
Jersey to Connecticut, Ontario, Michigan and Nebraska. 
Naturalized from Europe. June-July. 


GENUS 21. CHICORY FAMILY. 327 


5. Crepis capillaris (L.) Wallr. Smooth Hawksbeard. Fig. 4090. 


Lapsana capillaris L. Sp. Pl. 812. 1753. , 
Crepis virens 1. Sp. Pl. Ed. 2, 1134. 1763. iV 
Crepis polymorpha Wallr. Sched. Crit. 426. 1822. 
Crepis capillaris Wallr. Fl. Hereyn. 287. 1840. 
Annual; stem stout or slender, leafy, corym- 
bosely branched above, glabrous or somewhat 
hirsute below, 1°-23° high. Basal leaves spatu- 
late, pinnatifid, or dentate, sometimes 8’ long and 
2’ wide, narrowed into petioles; stem leaves lan- 
ceolate or oblong, clasping by a sagittate base, 
flat, the upper mostly very small and usually en- 
tire; heads numerous, 5’—-8” broad, slender-pedun- 
cled; peduncles glabrous or glandular; involucre 
oblong, more or less pubescent or glandular, 3-4” 
high, its principal bracts lanceolate, glabrous 
within, the outer mostly appressed; achenes I0- 
ribbed, smooth, slightly narrowed at both ends. 


In fields and waste places, Connecticut, New York, 
New Jersey and Pennsylvania, and in ballast about 
the seaports. Also on the Pacific Coast. Adventive 
from Europe. July—Sept. 


6. Crepis biénnis L. Rough Hawksbeard. 


Fig. 4091. 
Crepis biennis L. Sp. Pl. 807. 1753. 


Biennial, or sometimes annual; stem pubescent 
or hirsute, leafy, at least below, branched above, 
2°-3° high. Leaves runcinate-pinnatifid, usually 
hirsute, 2’-6’ long, oblong or spatulate, the lower 
and basal ories narrowed into petioles and some- 
times merely dentate, the uppermost lanceolate, 
clasping, their margins not revolute; heads sev- 
eral, subcorymbose, 1-12’ broad; involucre canes- 
cent or pubescent, 4”-6” high, its principal bracts 
linear-lanceolate, downy within, the outer ones 
linear-oblong or lanceolate, spreading; achenes 
oblong, slightly narrowed above, 13-striate, gla- 
brous. 


In waste places, Vermont to Pennsylvania, Mich- 
igan, and in ballast about the seaports. Naturalized 
or adventive from Europe. June—Aug. 


7. Crepis intermédia A. Gray. Small-flowered 
Gray Hawksbeard. Fig. 4002. 
Crepis intermedia A. Gray, Syn. Fl. 1: Part 2, 432. 1884. 


Perennial, cinerous-puberulent or scurfy; stem rather 
slender, 1-3-leaved, 1°-2° high, naked and branched 
above. Basal leaves lanceolate or oblong-lanceolate, 
laciniate-pinnatifid, long-acuminate at the apex, nar- 
rowed at the base, 4-6’ long; stem leaves lanceolate, 
sessile, less divided; heads several, 6-9” broad, 5-8- 
flowered; involucre oblong-campanulate, its principal 
bracts 5-8, lanceolate or linear-oblong, canescent, acut- 
ish, somewhat keeled by the thickened midrib when old, 
the outer ones few and short; achenes oblong, narrowed 
above, not beaked, strongly 10-ribbed. 

In dry soil, western Nebraska (according to Williams) ; 


Colorado to California, Montana and British Columbia. 
May-Sept. 


328 CICHORIACEAE., Vou. ik 


8. Crepis occidentalis Nutt. Large-flowered 
Gray Hawksbeard. Fig. 4093. 


Crepis occidentalis Nutt. Journ. Acad. Phila. 7: 29. 1834. 


Perennial, scurfy and canescent, sometimes also hir- 
sute; stem rather stout, leafy, branched, 6-18’ high. 
Leaves oblong-lanceolate or oblong in outline, laciniate- 
pinnatifid, acute or acuminate, the lower and basal ones 
6-10’ long, narrowed into petioles, the upper sessile and 
slightly clasping; heads few or several, corymbose, 
stout-peduncled, about i’ broad, 10-30-flowered; invo- 
lucre oblong-campanulate, canescent, its principal bracts 
9-24, linear, acute; achenes oblong, sharply 1o-ribbed, 
glabrous. 


Plains, western Nebraska (according to Gray) ; Colorado 
to California, north to Montana and British Columbia. 
May-July. 


22. HIERACIUM [Tourn.] L. Sp. Pl. 799. 1753. 


Perennial hispid scabrous glandular or glabrous herbs, with alternate or basal leaves, 
and small middle-sized or large, solitary corymbose or paniculate heads of yellow orange or 
red flowers. Involucre cylindric, campanulate, or nearly hemispheric, its principal bracts in 
1-3 series, the outer either regularly and gradually smaller or abruptly much smaller, none 
of them much thickened at the base after flowering. Receptacle flat, naked or short-fimbril- 
late. Rays truncate and 5-toothed at the apex. Anthers sagittate at the base. Style-branches 
slender. Achenes oblong, columnar, or fusiform, not beaked, 10-15-ribbed, terete or 4-5- 
angled. Pappus copious, or 1-2 rows of simple rather stiff persistent brown or brownish 
bristles. [Greek, hawk.] 

Not fewer than 300 species, natives of the north temperate zone and the Andes of South Amer- 


ica. Besides the following, some 20 others occur in western North America. Known as Hawkweed, 
Hawk-bit, or Speerhawk. Type species: Hieracium murorum L. 


A. Rootstock short, erect; plants without stolons. 
a. Heads 1'—2’ broad. 


Stem with 1-5 leaves only; basal leaves tufted ; introduced species. 


Leaves cordate or subcordate; scape naked or 1-leaved. 1. H. murorum. 

Leaves narrowed at the base; stem several-leaved. 2. H.vulgatum. 
Stem very leafy up to the inflorescence ; native species. 

Leaves short, ovate to ovate-lanceolate, rounded or clasping at the base. 3. H. canadense. 

Leaves elongated, linear-lanceolate, narrowed at the base. 4. H. scabriusculum. 


b. Heads less than 1’ broad. 


* Stem teat Deans nearly or quite up to the inflorescence, the upper leaves sometimes very small 
and distant. 
Pubescence of abundant brownish or whitish hairs %4’—1’ long; inflorescence elongated. 
5. H. longipilum. 
Pubescence of short hairs, or nearly or quite wanting. 
Leaves lanceolate to oblong-lanceolate, acute or acuminate, glabrous. 6. H. paniculatum. 
Leaves elliptic to obovate, mostly obtuse. 
Stem hispid-pubescent, densely glandular-hispid above; the peduncles stout, spreading. 
7. H, scabrum. 
Stem loosely pubescent; pedicels slender. 
Inflorescence elongated; achenes spindle-shaped. 8. H. Gronovit. 
Inflorescence corymbiform; achenes columnar, truncate. g. H. marianum. 
** Leaves all basal or 1 or 2 borne on the stem above. 
Basal leaves elliptic to obovate or oblong-spatulate ; native species. 
Pedicels and involucres glabrous or nearly so; leaves glabrous or loosely pubescent. 
10. H. venosum. 
Pedicels and involucres glandular-pubescent ; leaves villous. 11. H. Greenii. 
Basal leaves narrowly oblanceolate, introduced. 12. H. florentinum. 


B. Rootstock elongated, slender; plants mostly stoloniferous, scapose. 


Scape bearing a single head, rarely 2 to 4. 13. H, Pilosella. 
Heads several or many, corymbose. 
Flowers yellow. 
Plant glaucous; leaves glabrous or nearly so above. 14. H. floribundum. 
Plant not glaucous; leaves hirsute on both sides. 15. H. pratense. 
Flowers orange. 16. H. aurantiacum. 


GENUS 22. CHICORY FAMILY. 329 


1. Hieracium murorum L. Wall Hawk- ww Sy 
\ 


weed. Fig. 4094. TNS Hi, 


Hieracium murorum L. Sp. Pl. 802. 1753. 


Stem pubescent or glabrate, simple, or with I or 
2 branches, 1°-22° high. Basal leaves thin, ovate or 
oblong, obtuse or acute, cordate or truncate at the 
base, or abruptly narrowed into petioles, coarsely 
dentate or laciniate, at least near the base, 2-4’ long, 
1’-2’ wide, the petioles villous; stem leaves I or 2, 
short-petioled or sessile, sometimes none; heads 
2-several, corymbose, about I’ broad; peduncles as- 
cending, usually glandular; involucre 4’—5” high, its 
bracts linear-lanceolate, acute, glandular-pubescent, 
‘imbricated in 2 or 3 series; achenes columnar, trun- 
cate; pappus of slender nearly white bristles. 

Woodlands near Brooklyn, N. Y., Northampton, 


Mass., and about Quebec. Adventive or fugitive from 
Europe. French or golden lungwort. June—Aug. 


2. Hieracium vulgatum Fries. Hawkweed. 
Fig. 4095. 
H. molle Pursh, Fl. Am. Sept. 503. 1814. Not Jacq. 1774. 
Hi. vulgatum Fries, Fl. Hall. 128. 1817-18. 

Similar to the preceding species, sometimes taller 
and slightly glaucous; stem 2-5-leaved, pubescent or 
glabrate. Basal leaves oblong or lanceolate, acute at both 
ends, or some of them obtuse at the apex, coarsely den- 
tate or denticulate, petioled, 2’-5’ long, 4’-12’ wide, often 
mottled; stem leaves similar, short-petioled or sessile; 
petioles more or less pubescent; heads several, corym- 
bose, smaller than those of H. murorum or as large; 
peduncles mostly glandular, straight; bracts of the in- 
volucre imbricated in 2 or 3 series, linear, acuminate, 
mostly glandular; achenes columnar, truncate; pappus 
copious. 

Labrador and Newfoundland to Quebec, and in southern 


¢ ; New York and New Jersey. Naturalized from Europe. 
Also in Greenland, northern Europe and Asia. July—Sept. 


3. Hieracium canadénse Michx. Canada 
Hawkweed. Fig. 4096. 


H. canadense Michx. Fl. Bor. Am. 2: 86. 1803. 


Stem erect, firm, glabrate or pubescent, leafy, 
1°-5° high. Leaves numerous, ovate, oblong- 
lanceolate, ovate-oblong, or lanceolate, acute or 
acuminate at the apex, rounded, sessile, and, at 
least the upper ones, clasping at the base, 1’—3’ 
long, 3-12” wide, serrate or incised, the mar- 
gins sometimes ciliolate, glabrous or pubescent 
beneath, the lowest somewhat spatulate and 
petioled; no tuft of basal leaves at flowering 
time; heads usually numerous, corymbose- 
paniculate, about 1’ broad; involucre about 6” 
high, pubescent or puberulent, its bracts imbri- 
cated in 2-3 series, the outer spreading; flowers 
yellow; achenes columnar, truncate; pappus 
copious, brown. 

In dry woods and thickets, Newfoundland and 
Nova Scotia to Ontario, British Columbia, New 


Jersey, Indiana, Michigan, South Dakota and 
Oregon. July-Sept. High dandelion. 


Doe CICHORIACEAE. Vor. III. 


4. Hieracium scabritsculum Schwein. 
Narrow-leaved Hawkweed. Fig. 4097. 


H. scabriusculum Schwein. in Long’s Exp. 2: 394. 


y 9 


Yi) 


KX Vz 4 1824. 
INST Stem rather slender, glabrous or puberulent, 


sometimes hispid below, usually very leafy nearly 
or quite up to the inflorescence, usually simple, 
1°-23° high. Leaves lanceolate or linear-lanceo- 
late or the lowest spatulate, entire, denticulate or 
sometimes laciniate-dentate, acute or acuminate, 
narrowed to a sessile base, 1-3’ long, 2”-7” wide, 
glabrous above, mostly somewhat pubescent be- 
neath, the margins commonly ciliolate; no tuft 
of basal leaves at flowering time; heads few or 
several, corymbose, about I’ broad; peduncles 
rather stout, canescent; involucres 5-7” high, 
glabrous or somewhat pubescent, its bracts. im-+ 
bricated in 2-3 series, the outer spreading; flow- 
ers bright yellow; achenes columnar, truncate; 
pappus copious, brownish. 

Minnesota to Nebraska, Saskatchewan, British 
Columbia and Oregon. Included in our first edition 
in the similar Old World H. uwmbellatum L. Appar- 
ently erroneously recorded from the St. Lawrence 
River. June—Aug. 


i 


5. Hieracium longipilum Torr. Long-bearded 
Hawkweed. Fig. 4008. 


H. longipilum Torr.; Hook. Fl. Bor. Am. 1: 298. 1833. 
Hieracium barbatum Nutt. Journ. Phil. Acad. 7: 70. 
1834. Not Tausch. 1828. 


Stem, at least its lower portion, and leaves densely 
covered with long brown rather rigid bristly hairs 
+’-1’ long, arising from papillae; stem, very leafy 
below, stiff, simple, 2°-33° high. Basal and lower 
leaves spatulate or oblong, obtuse, mostly entire, 
4-8’ long, narrowed into margined petioles, the 
upper lanceolate or spatulate, mostly sessile, the S 
uppermost small and bract-like; heads not very 
numerous, racemose or racemose-paniculate, 8’—10” 
broad; peduncles short, stout, tomentose and glandu- 
lar; involucre 4”-5” high, its principal bracts in I 
series, linear-lanceolate, acuminate, with several 
short subulate outer ones; flowers yellow; achenes 
fusiform with a slightly tapering summit; pappus 
brown. 

Prairies and dry woods, Ontario to Minnesota, Illi- 
nois, Kansas and Texas. July—Sept. 


6. Hieracium paniculatum L. Panicled 
Hawkweed. Fig. 4099. 
Hieracium paniculatum L. Sp. Pl. 802. 1753. 


Glabrous throughout, or somewhat pilose-pu- 
bescent below, stem paniculately branched above, 
leafy, slender, 1°-3° high. Leaves thin, lanceo- 
late or oblong-lanceolate, acute or acuminate at 
the apex, narrowed to a sessile base, or the lowest 
into petioles, denticulate or dentate, 2-6’ long, 
3-12” wide; no tuft of basal leaves at flowering 
time; heads 5-7” broad, commonly numerous, 
corymbose-paniculate, 12-20-flowered; peduncles 
slender, often drooping, quite glabrous or some- 
times glandular; involucre about 3” high, glabrous 
or nearly so, its principal bracts in 1 series, linear, 
acute with a few very small outer ones at the 
base; flowers yellow; achenes columnar, truncate ; 
pappus brown, not very copious. 


In dry woods, Nova Scotia to Ontario, Michigan, 
Georgia, Alabama and Tennessee. Ascends to 4600 
ft. in Virginia. July—Sept. 


GENUS 22. CHICORY FAMILY, _ 331 


7. Hieracium scabrum Michx. Rough 
Hawkweed. Fig. 4100. 
H. scabrum Michx. Fl. Bor. Am. 2: 86. 1803. 


Stem stout, leafy, mostly hirsute or hispid be- 
low and glandular-pubescent above, strict, 1°-4° 
high. “Leaves hirsute, obovate, oblong, or broadly 
spatulate, 2’-4’ long, 1’-2’ wide, obtuse at the apex, 
narrowed to the sessile base or the lowest into 
margined petioles, denticulate; no tuft of basal 
leaves at flowering time; heads usually numerous, 
6-8” broad, corymbose- or racemose-paniculate ; 
peduncles stout, densely glandular; involucre 4”-5” 
high, glandular, its principal bracts in 1 series, 
linear, acute with a few very small outer ones; 
flowers yellow; achenes columnar, truncate; pap- 
pus brown. 

In dry woods and clearings, Nova Scotia to Minne- 
sota, Georgia, lowa, and recorded from Nebraska and 
Kansas. July—Sept. 


8. Hieracium Gronovii L. Gronovius’ or 
Hairy Hawkweed. Fig. 4101. 
Hieracium Gronovii L. Sp. Pl. 802. 1753. 


Stem stiff, mostly slender, leafy and villous or 
hirsute, at least below, sometimes nearly leafless, 
1°-3° high. Leaves villous or hirsute, the basal 
and lower ones obovate or spatulate, denticulate 
or entire, obtuse, 2-6’ long, usually narrowed into 
petioles; stem leaves mostly sessile, oblong or oval, 
obtuse or acute, narrowed or broad at the base, 
the upper gradually smaller; heads numerous, 
racemose-paniculate, 5-8” broad; peduncles glan- 
dular and canescent, slender; involucre about 4” 
high, somewhat canescent, its principal bracts in I 
series, linear-lanceolate, acute, with several very 
small outer ones; flowers yellow; achenes spindle- 
shaped, with a tapering summit; pappus brown. 

In dry soil, Massachusetts to Ontario, Illinois, 
Kansas, Florida, Louisiana and Texas. Santo Do- 
mingo. July-Sept. Cat’s-ear. 


g. Hieracium marianum Willd. Mary- 
land Hawkweed. Fig. 4102. 
H. marianum Willd. Sp. Pl. 3: 1572. 1804. 


Stems usually solitary, slender, pilose-pubescent, 
at least below, paniculately branched above, 2°-3° 
high, bearing 2-7 leaves. Basal leaves obovate, 
oblanceolate or oblong, ascending or erect, obtuse, ' 
narrowed at the base, sessile or petioled, hirsute, 
at least on the veins beneath, entire or glandular- 
denticulate, 2’-8’ long, 1-2’ wide, not purple-veined, 
mostly glabrous above, those of the stem similar, 
mostly smaller; heads commonly numerous, co- 
rymbose-paniculate, 6’—10” broad, 15-40-flowered, 
slender-peduncled; peduncles more or less glan- 
dular and sometimes canescent; involucre about 
4” high, its principal bracts in 1 series, linear- 
lanceolate, acute, or acuminate, glabrous or nearly 
so, with a few short outer ones; achenes col- 
umnar, truncate; pappus brown, not copious. 

In dry woods and thickets, New Hampshire to 
southern New: York, Pennsylvania, Ohio, Kentucky, 
Alabama and Florida. May-July. 

Hieracium alleghaniénse Britton, of the mountains of West Virginia, has the stem. more 
abundantly leafy and no tuft of basal leaves at flowering time. 


332 CICHORIACEAE. Vot. III. 


10. Hieracium venosum L. Rattlesnake-weed. Poor Robin’s Plantain. 


Fig. 4103. 
Hieracium venosum L. Sp. Pl. 800. 1753. 


Stems solitary or several from the same root, 
slender, glabrous, or with a few hispid hairs near 
the base, or also above, leafless or with 1-3 leaves, 
paniculately branched above, 1°-3° high. Basal 
leaves tufted, spreading on the ground, obovate, oval 
or oblong-spatulate, mostly obtuse, narrowed at the 
base, sessile or petioled, 1-4’ long, 2’-13’ wide, usu- 
ally purple-veined, glabrous or more or less hirsute, 
pale beneath, some or all of them glandular-der*icu- 
late; heads commonly numerous, 5’-8” broad, .5- 
40-flowered, slender-peduncled; peduncles glabrous, 
or slightly glandular; involucre about 3” high, its 
principal bracts in 1 series, glabrous or nearly so, 
with a few short outer ones; achenes columnar, trun- 
cate; pappus brown, not copious. 

Dry woods and thickets, Maine to Ontario and Mani- 
toba, south to Georgia, Kentucky and Nebraska. Ascends 
to 4200 ft. in North Carolina. Early or vein-leaf hawk- 
weed. Striped bloodwort. Snake-plantain. Hawkbit. 
Adder’s-tongue. May—Oct. 


11. Hieracium Greénii Porter and Britton. 
Green’s Hawkweed. Fig. 4104. 
Pilosella spathulata Sch. Bip. Flora 45: 439. 1862. Not 

Hieracium spathulatum Scheele, 1863. 

Hieracium marianum var. spathulatum A. Gray, Syn. FI. 

1: Part 2, 446. 1886. 
dale Green Porter and Britton, Bull. Torr. Club 20: 120. 

1893. 

Stem entirely glabrous up to the branches, rather 
slender, leafless or rarely with 1 or 2 leaves. 13°-23° 
high. Basal leaves tufted, ascending, spatulate, ob- 
long, or obovate, obtuse, narrowed at the base, mostly 
petioled, glandular-denticulate or entire, villous- 
pubescent or somewhat hispid, 4-7’ long, 3’-2’ wide; 
heads corymbose-paniculate, several or numerous, 
30-40-flowered, slender-peduncled, 8-10” broad; 
peduncles and branches canescent-tomentose and 
glandular; involucre 5” high, its principal bracts in 
I series, linear, acute, densely pilose-glandular; 
flowers bright yellow; achenes columnar, truncate; 
pappus brownish, not copious. 

In dry soil, mountains of Pennsylvania to Ohio, Vir- 
ginia and West Virginia. May—June. 


12. Hieracium florentinum All. King- 
devil. Fig. 4105. 
H, florentinum All. Fl. Ped. 1: 213. 1785. 

Stolons wanting; stem glabrous, or somewhat 
hispid, glaucous, slender, 12°-3° high, bearing 1-3 
leaves near the base. Basal leaves tufted, nar- 
rowly oblong, oblanceolate, or spatulate, entire, 
obtuse or acute at the apex, narrowed below into 
margined petioles, more or less hirsute with stiff 
hairs, or glabrous, 2’-4’ long, 3’-7” wide; heads 
several or numerous, corymbose, 4-6” broad; 
peduncles mostly short, pilose and glandular; 
involucre about 3” high, its bracts linear, acute 
or acuminate, pilose and somewhat glandular, 
imbricated in about 2 series; flowers yellow; 
achenes oblong, truncate; pappus a row of slen- 
der brownish bristles. 

In fields, meadows and along roadsides, New York 
and Ontario to Quebec and Maine; a troublesome 
weed. Naturalized from Europe. Referred in our | 
first edition to H. praealtum, also native of Europe, ° 
which differs in having long leafy branches from the 
base, and is recorded as established in a meadow at 
Andover, Massachusetts. June—Sept. 


GENUS 22. CHICORY FAMILY. 333 


13. Hieracium Pilosélla L. Mouse-ear Hawk- ane 
: NZ 
weed. Fig. 4106. KY ZZ 
Hieracium Pilosella L. Sp. Pl. 800. 1753. ait 
Hieracium Pilosella peleterianum Mer. Nouv. FI. Paris, Ed. Yj N) 


2, 230. 1821. 

Stoloniferous, pilose-pubescent throughout; stolons 
leafy, rooting, slender, 3-12’ long. Scape slender, 
erect, 4-15’ high, leafless, with a single head, or some- 
times 2-4; leaves oblong or spatulate, entire, obtuse or 
acutish at the apex, narrowed into petioles, often white- 
tomentose beneath, 14-3’ long, 4’-8’ wide; head 1’ 
broad or more; flowers yellow; principal bracts of the 
involucre in I or 2 series, linear, acuminate, pubescent, 
usually with I or 2 exterior ones; achenes oblong, trun- 
cate; pappus a single row of slender bristles. 

Dooryards and fields, Prince Edward Island to Ontario, ‘ 
New York, Pennsylvania and Michigan. Adventive from 


Europe. Ling-gowans. Felon-herb. Mouse-bloodwort. May-— 
Sept. 


14. Hieracium floribindum Wimm. & 
Grab. Smoothish Hawkweed. 
Fig. 4107. 
Hieracum floribundum Wimm. & Grab. FI. Siles. 27: 

204. 1829. 

Stoloniferous; plant glaucous-green; scape 
loosely hirsute and more or less glandular- 
pubescent, slender, 1°-23° high, the stolons some- 
times 8 long. Basal leaves tufted, narrowly 
oblanceolate, acutish or obtuse, 2’-6’ long, gla- 
brous or very nearly so above, the margins and 
midvein beneath more or less hirsute; stem- 
leaves none, or rarely I or 2 near the base; 
flowers several, 1’ broad or less, corymbose; 
peduncles glandular; rays bright yellow; bracts 
of the involucre in about 2 series, hirsute. 


In fields, New Brunswick and Maine to New York. 
Naturalized from Europe. June—Aug. 


15. Hieracium praténse Tausch. Fieid 
Hawkweed. Fig. 4108. 


H. pratense Tausch, Flora 11: Part 1, Erg. 56. 1828. 


Stoloniferous, hirsute or pilose with long hairs, 
those of the stem blackish. Stem scapose, simple, 
1°-2° high, often bearing 1 or 2 leaves below the 
middle; basal leaves numerous, tufted, light green, 
oblanceolate to oblong, obtuse, 2-5’ long, 5’’-10” 
wide, narrowed into margined petioles, or to a 
sessile base, entire, or with few distant minute 
glandular teeth, hirsute on both sides; heads ? VAG 
several or numerous, corymbose-paniculate, 10° > ‘Y Sy, fe 
wide, or less; flowers yellow; peduncles glandular £y ae: 
and often tomentose; bracts of the involucre 
linear-lanceolate, acuminate, glandular and pilose; 
achenes columnar, truncate. 


[ry 
— 


tS 4 
eae 


nj— 
Bt 
tos 
“%, 


\ JB atte 


Fields and roadsides, Quebec to southern New 
York and Pennsylvania. Naturalized or adventive 
from Europe. 


334 CICHORIACEAE. Vot. III. 


16. Hieracium aurantiacum L. Orange or 
Tawny Hawkweed. Golden Mouse-Ear 
Hawkweed. Fig. 4109. 


Hieracium aurantiacum L. Sp. Pl. 801. 1753. 


Stoloniferous; stem leafless or rarely with I or 
2 small sessile leaves, hirsute, slender, 6-20’ high. 
Basal leaves hirsute, tufted, spatulate or oblong, 
_obtuse, narrowed at the base, entire, or sometimes 
slightly denticulate, 2’-5’ long, 3-1’ wide; heads 
several, short-peduncled, corymbose, 7-12” broad; 
peduncles glandular-pubescent; involucre 4-5” 
/ high, its bracts linear-lanceolate, acuminate, im- 
+ bricated in 2 or 3 series, hirsute and sometimes 
\' glandular; flowers orange or red; achenes oblong, 
truncate; pappus a single row of slender brownish 
bristles. 


In fields, woods and along roadsides, New Brans- 
wick and Ontario to New York, New Jersey and 
Pennsylvania. Naturalized from Europe. Grim-the- 
collier. Devil’s- or Flora’s-paint-brush. Red daisy. 

_ Missionary-weed. June—Sept. 


23. NABALUS Cass. Dict. Sci. Nat. 34:94. 1825. 


Perennial, herbs with alternate, mostly petioled, dentate lobed or pinnatifid leaves, or the 
upper auriculate and clasping, and numerous small heads of ligulate white yellowish or 
purplish flowers in open or spike-like terminal panicles, or also in axillary clusters, usually 
drooping. Involucre cylindric, usually narrow, its principal bracts in I or 2 series, nearly 
equal, with a few smaller exterior ones at the base. Receptacle flat, naked. Rays truncate 
and 5-toothed at the summit. Style-branches slender. Achenes oblong or narrowly columnar, 
truncate, terete or 4-5-angled, mostly 10-ribbed. Pappus of copious rather rigid simple white 
to reddish-brown bristles. [Modern Latin, from an Indian name for Rattlesnake-root.] 


About 25 species, natives of America and Asia. Besides the following, two others occur in 
northwest America and one on the high mountains of North Carolina. Known by the general 
name of Rattlesnake-root or Drop-flower. Type species: Nabalus trifoliatus Cass. The European 
and African genus Prenanthes L. is here regarded as distinct from this. 


* Bracts of the involucre glabrous, or with a few scattered hairs. 


Heads s-7-flowered ; involucre very narrow, light green, 1” thick ; pappus light straw-color or brown. 
1. N. altissimus. 
Heads 8-16-flowered; involucre broader, green, purple or glaucous, 1%4”-3” thick. 
Leaves, or some of them, lobed, divided, or pinnatifid (sometimes entire in No. 3); involucre 
subcylindric, about 1%” thick. 
Pappus deep cinnamon-brown. 2. N. albus. 
Pappus straw-color or light brown. 
Inflorescence paniculate. 
Involucral bracts with some stiff hairs, obviously shorter than the pappus; panicle- 


branches divergent. 3. N.serpentarius. 
Involucral bracts glabrous, equalling the pappus; panicle-branches ascending, or 
upcurved. 4. N. trifoliolatus. 
Inflorescence thyrsoid or glomerate. 
Leaves palmately lobed or divided ; northern. 5. N. nants. 
Leaves pinnately lobed or pinnatifid ; southern. 6. N. virgatus. 
Leaves entire or denticulate; plant low, alpine; involucre narrowly campanulate, 21%4”-3” thick. 
7. N. Boottii. 


** Bracts of the involucre hirsute-pubescent. 


Inflorescence narrowly thyrsoid ; heads 8—16-flowered. 
Leaves and stem rough-puberulent or scabrous. 8. N. asper. 
Leaves and stem glabrous, glaucous. 9g. N.racemosus. 
Inflorescence corymbose-paniculate ; heads 20—25-flowered. 10. N.crepidineus. 


“ 4s a 


GENUS 23. CHICORY FAMILY. 335 


1. Nabalus altissimus (L.) Hook. Tall 
White Lettuce. Fig. 4110. 


Prenanthes altissima L. Sp. Pl. 797. 1753. 
N. altissimus Hook, Fl. Bor. Am. 1: 294. 1833. 


Glabrous, or sometimes hispidulous, not 
glaucous; stem slender, 3°-7° high, green, or 
sometimes purplish. Leaves thin, hastate, 
cordate, ovate, or the uppermost lanceolate, 
entire, denticulate, dentate or palmately lobed 
or divided, most of them long-petioled, the 
larger sometimes 6’ long; heads very numer- 
ous, in a narrow panicle, and often in axillary 
clusters, 5-7-flowered, pendulous, about 2” 
broad; inflorescence often narrow; involucre 
narrowly cylindric, 5-6” long, about 1” thick, 
green, glabrous, its principal bracts about 5; 
flowers greenish or yellowish white; pappus 
light straw-color, or cinnamon-brown. 


In woods and thickets, Newfoundland to Mani- 
toba, Missouri, Georgia and Louisiana. Lion’s- 
foot. Rattlesnake-root. Ascends to 2500 ft. in 
the Catskills. Wild lettuce. Joy-leaf. Milk-weed. 
Bird-bell. Races differ in leaf-form, pubescence 
and in color of the pappus. July—Oct. 


(WV y — (Om fa 2. Nabalus albus(L.) Hook. Rattlesnake- 
Gi (\ Vf Ay \ Lav root. White Lettuce. Fig. 4111. 


Prenanthes alba L. Sp. Pl. 798. 1753. 
Gir Nabalus albus Hook. Fl. Bor. Am. 1: 294. 1833. 


Glabrous and glaucous; stem commonly purple, 
2°-5° high. Leaves hastate, ovate, cordate, den- 
ticulate, dentate, lobed, or palmately divided, or 
the upper lanceolate, entire, thicker than those of 
the preceding species, the larger sometimes 8’ 
long; heads numerous, pendulous, 8-15-flowered, 
about 3” broad, paniculate, or thyrsoid, and often 
in axillary clusters; involucre glabrous, or with a 
few scattered hairs, glaucous, 5-7” high, about 
12” thick, its principal bracts about 8, purplish, 
with minute outer ones; flowers greenish or yel- 
lowish white, fragrant; pappus cinnamon-brown. 


7 


In woods, Maine and Ontario to Manitoba, Sas- 
katchewan, Georgia, Kentucky, Wisconsin and North 
Dakota. Lion’s-foot. White cankerweed. Wild lettuce. 
Milk-weed. Joy-leaf. Cancer-weed. Aug.—Sept. 


3. Nabalus serpentarius (Pursh) Hook. 
Lion’s-foot. Gall-of-the-Earth. Fig. 4112. 


Prenanthes serpentaria Pursh, Fl. Am. Sept. 499. pl. 
PAL TOA: 

Nabalus integrifolius Cass. Dict. Sci. Nat.34: 95. 1825. 

_ Nabalus serpentarius Hook. Fl. Bor. Am.1: 294. 1833. 


Glabrous or sparingly pubescent, green; stem 
stout or slender, not glaucous, 1°-4° high. Leaves 
rather firm, similar to those of the preceding 
species, equally variable in outline, pinnatifid or 
pinnately lobed, palmately divided, or merely 
dentate, or entire; inflorescence paniculate, the 
branches divaricate, upcurved; heads numerous, 
about 3” broad, 8-12-flowered, pendulous, panicu- 
late, and commonly also in axillary clusters; in- 
volucre more or less bristly-hispid, rarely gla- 
brous, green or purplish, about 13” thick, 5’—7” 
long, its principal bracts about 8, shorter than the 
pappus, with several minute lanceolate outer ones; 
flowers whitish or cream-color, rarely yellow; 
achenes about 3” long; pappus light brown or straw-color. 


336 CICHORIACEAE. Vor. IIL. 


In fields and thickets, Massachusetts to southern New York, Florida, Alabama, Kentucky and 
Mississippi. Canker-weed. Rattlesnake-root. White lettuce. Snake-gentian. July—Oct. 

Nabalus cylindricus Small, of the southern mountains, ranging north to Kentucky, differs by 
an elongated narrow panicle. 


4. Nabalus trifoliolatus Cass. Tall Rattle- 
snake-root. Fig. 4113. 


Nabalus trifolilatus Cass. Dict. Sci. Nat. 34: 95. 1825. 
Prenanthes trifoliata Fernald; Brainerd, Jones & Eggles- 
ton, Fl. Verm. 89. 1900. 


Glabrous throughout; stem usually stout, 3°-9° 
high. Leaves thinnish, the lower long-petioled, usu- 
ally 3-divided with the divisions stalked or sessile, 
the upper short-petioled or sessile, all commonly 
lobed or dentate, but the upper sometimes lanceo- 
late, acuminate and entire; inflorescence paniculate, 
the panicle-branches ascending, or nearly erect; 
heads few in the clusters, drooping, 7-12-flowered ; 
involucre pale green or purplish, glabrous, about 13” 
thick and 6” long, its principal bracts 6-8, equalling 
the pappus, the short outer ones ovate to lanceolate; © 
flowers whitish or pale yellow; achenes 2’-3” long; 
pappus light brown. 

In woods and thickets, Newfoundland to Pennsylva- 
nia, Indiana, Delaware and Tennessee. Aug.—Oct. 


5. Nabalus nanus (Bigel.) DC. Low Rattlesnake-root, or Lion’s-foot. Fig. 4114. 
Hreponines alba var. nana Bigel. Fl. Bost. Ed. 2, 286. 

1824. 
Nabalus nanus DC. Prodr. 7: 241. 1838. 
Prenanthes nana Torr.; Robinson & Fernald in A. 

Gray, Man. Ed. 7, 871. 1908. 
Prenanthes serpentaria var. nana A, Gray, Syn. FI. 

Tew eanoee els A roode 

Glabrous throughout; stem simple, erect, 4’-16’ 
high. Basal and lower leaves slender-petioled, 
3-divided, or sometimes broadly hastate, the divi- 
sions variously lobed, toothed, or entire, usually 4 
sessile, occasionally stalked; upper leaves much 
smaller, entire, toothed, or lobed, sessile, or short- 
petioled; inflorescence thyrsoid, glomerate-spicate 
or racemose, rarely with I or 2 short ascending 
branches; involucre dark purple-brown or nearly 
black, glabrous, 4-6” long, its inner bracts 6-8, 
slightly ciliate at the apex, about as long as the 
usually bright brownish pappus; outer bracts lan- 
ceolate to ovate-lanceolate. 

Alpine summits of the Adirondacks and the moun- 
tains of New England; Nova Scotia to Labrador and 
Newfoundland. Aug.—Sept. ; 


6. Nabalus virgatus (Michx.) DC. Slender 
Rattlesnake-root. Fig. 4115. 


Prenanthes virgata Michx. Fl. Bor. Am. 2: 84. 1803. 
Nabalus virgatus DC. Prodr. 7: 242. 1838. 


Glabrous, somewhat glaucous; stem strict, mostly 
simple, 2°-4° high. Leaves lanceolate or oblong- 
lanceolate, the basal and lower ones sinuate-pin- 
natifid or pinnately parted, petioled, often 10’ long, 
the lobes entire or dentate, distant; upper leaves all 
sessile, pinnately lobed, or the uppermost entire, 
very small and bract-like; heads very numerous, 
pendulous, about 2” broad, in a narrow, simple or 
branched, terminal thyrsus, often unilateral; invo- 
lucre purplish, about 13” thick and 5” long, its 
principal bracts about 8, with several minute outer 
ones; flowers white or pinkish; pappus straw-color. 


In moist sandy soil, New Jersey to Florida, near the 
coast. Called also De Witt’s-snakeroot. Sept.—Oct. 


GENUS 23. CHICORY FAMILY. 337 


7. Nabalus Boottii DC. Boott’s Rattle- 
snake-root. Fig. 41106. 


Nabalus Boottii DC. Prodr. 7: 241. 1838. 
Prenanthes Boottii A. Gray, Syn. Fl. 1: Part 2, 435. 
1884. 


Glabrous below, commonly pubescent above; 
stem simple, 4’-12’ high. Leaves thickish, the 
basal and lower ones ovate, hastate, or deltoid, 
petioled, mostly obtuse, entire, or denticulate, 1’-2’ 
long, the upper ovate or oblong, usually entire, 
short-petioled or sessile, much smaller; heads 
several or numerous, 10-18-flowered, erect, spread- 
ing, or some of them pendulous, racemose or 
somewhat thyrsoid, 4-5” broad; involucre cam- 
panulate-oblong, 23”-3” thick, 4”-7” long, dark 
purplish-green, its principal bracts 8-10, obtuse or 
obtusish, with several shorter outer ones; flowers 
whitish, odorous; pappus brownish. 


Alpine summits of the mountains of northern New 
England and New York. July—Aug. 


8. Nabalus asper (Michx.) T. & G. Rough 
White-lettuce. Fig. 4117. 


Prenanthes aspera Michx. Fl. Bor. Am. 2: 83. 1803. 
Nabalus asper T. & G. Fl. N. A. 2: 483. 1843. 


Scabrous or rough-puberulent; stem virgate, simple, 
2°-4° high. Leaves firm, oval, oblong, or oblanceolate, 
those of the stem all closely sessile, acute or acutish, 
dentate, denticulate, or the uppermost entire, the lower 
sometimes clasping, 2’-3’ long, 4-1’ wide, the basal 
tapering into winged petioles, commonly obtuse; heads 
very numerous, erect, spreading, or slightly drooping, 
3-4” broad, 12-16-flowered, in a long narrow thyrsus; 
involucre oblong, 2-22” thick, 5’-7” high, very hirsute, 
its principal bracts 8-10, with several short outer ones; 
flowers light yellow; pappus straw-color. 


On dry prairies, Ohio to South Dakota, Nebraska, Ken- 
tucky, Louisiana and Kansas. Rattlesnake-root. Aug.—Sept. 


g. Nabalus racemosus (Michx.) DC. Glaucous White-lettuce. Fig. 4118. 


Prenanthes racemosa Michx. Fl. Bor. Am. 2: 83. 1803. 


Nabalus racemosus DC. Prodr. 7: 242. 1838. AY 
Nabalus racemosus pinnatifidus Britton; Britt. & Brown, W) ) 
MU BIE 3": 2or. ~ 1898. ESSN 
Prenanthes racemosus var. pinnatifida A. Gray, Syn. FI. a KiNG 
bart 2) 433. 1884. \(l 4 
Stem virgate, rather stout, glabrous and somewhat eA\ 
glaucous; stem striate, 2°-6° high. Leaves thickish, SS 
glabrous and glaucous, the lower and basal ones oval, Zan 
oblong, oblanceolate, or obovate, dentate, denticulate, SAY 


pinnatifid or pinnately lobed, 4’-8’ long, mostly ob- 
tuse, tapering into long margined petioles; upper 
leaves sessile, smaller and partly clasping, lanceo- 
late to ovate-lanceolate, denticulate, entire, or pin- 
natifid, mostly acute; heads very numerous, erect, 
spreading, or slightly drooping, 12-16-flowered, 2”’-3” 
broad, in a long narrow thyrsus; involucre oblong- 
cylindric, hirsute, 5-6” long, 14”-23” thick, longer 
than the hirsute peduncle, its principal bracts 8-10, 
with several small outer ones; flowers purplish; 
pappus straw-color. 

In moist open places, New Brunswick and Quebec to 
Maine, Manitoba, Alberta, southern New York, New 
Jersey, Iowa, Missouri and Colorado. Aug.—Sept. 5 

Prenanthes mainénsis A. Gray, from northern Maine and New Brunswick, is probably a hybrid 
between N. racemosus and N. trifoliolatus. 
22 


338 CICHORIACEAE. Vor. III. 


10. Nabalus crepidineus (Michx.) DC. Corymbed Rattlesnake-root. Fig. 4119. 


Prenanthes crepidinea Michx. Fl. Bor. Am. 2: 84. 1803. 
Nabalus crepidineus DC. Prodr. 7: 242. 1838. 

Stem glabrous or very nearly so below, corym- 
bosely branched and sometimes puberulent above, 
stout, 5°-9° high. Leaves thin, the basal and lower 
ones hastate, ovate, oblong, or deltoid, sharply den- 
tate, lobed, or incised, sometimes Io’ long, usually 
with broadly winged petioles; the upper much 
smaller, sessile or short-petioled, narrowed at the 
base, not clasping, ovate, deltoid, or lanceolate, 
acute; heads numerous, pendulous, short-peduncled, 
corymbose-paniculate, 4-6” broad, 20-35-flowered; 
involucre oblong or oblong-campanulate, hirsute, 
5-8” long, about 3” thick, dark green or purplish, 
its principal bracts 12-15, with several very short 
outer ones; flowers cream color; pappus brown. 

In fields and thickets, western Pennsylvania and New 


York to Kentucky, west to Minnesota and Kansas. 
Aug.—Oct. 


Family 45. AMBROSIACEAE Reichenb. Consp. 112. 1828. 


RAGWEED FAMILY. 


Annual or perennial herbs, monoecious, or sometimes dioecious, many of them 
weeds, some shrubby, with alternate leaves, or the lower opposite, and small heads 
of greenish or white flowers subtended by an involucre of few, separate or united 
bracts, the pistillate heads sometimes larger and nut-like or bur-like. Staminate 
and pistillate flowers in the same heads, or in separate heads. Receptacle chaffy. 
Pistillate flowers with no corolla, or this reduced to a short tube or ring; calyx 
adnate to the 1-celled ovary, its limb none, or a mere border; style 2-cleft. Stami- 
nate flowers with a funnelform tubular or obconic 4—5-lobed corolla; stamens 
mostly 5, separate, or their anthers merely connivent, not truly syngenesious, with 
short inflexed appendages; ovary rudimentary; summit of the style often hairy 
or penicillate. 


Eight genera and about 60 species, mostly natives of America, a few only of the Old World. 


Staminate and pistillate flowers in the same heads; involucre of a few rounded bracts. 1. Iva. 
Staminate and pistillate flowers in separate heads, the staminate mostly uppermost; involucre of 
the pistillate heads bur-like or nut-like. 
Involucral bracts of the staminate heads united. 
Involucre of the pistillate heads with several tubercles or prickles in a single series. 
2. Ambrosia. 
Involucre of the pistillate heads with numerous prickles in several series. 3. Gaertneria. 
Involucral bracts of the staminate heads separate; involucre of pistillate heads an oblong bur. 
4. Xanthium. 


TS VA. L Sp. Pl..988.. 1753: 


Puberulent or scabrous herbs, with thick opposite leaves, or the upper alternate, and 
small nodding, axillary and solitary, spicate racemose or paniculate heads of greenish flowers. 
Involucre hemispheric or cup-shaped, its bracts few, rounded. Receptacle chaffy, the linear 
or spatulate chaff enveloping the flowers. Marginal flowers 1-6, pistillate, fertile, their corollas 
short, tubular or none. Disk-flowers perfect, sterile, their corollas funnelform, 5-lobed, their 
styles undivided, dilated at the apex. Anthers entire at the base, yellow, scarcely coherent 
with each other, tipped with mucronate appendages. Achenes compressed, obovoid, glabrous. 
Pappus none. [Named after Ajuga Iva, from its similar smell.] 

About 15 species, natives of America. Besides the following, 7 others occur in the southern 
and western United States. Type species: Iva annua L. 


Heads spicate or racemose, each subtended by a linear or oblong leaf. 
Heads solitary, pedicelled. 
Bracts of the involucre 4-5 ; heads 114”—2” high. 

Leaves serrate, oval or oblong; eastern. 1. I. frutescens. 

Leaves entire or nearly so, obovate or oblong; western. nS fe axillaris. 
Bracts of the involucre 6-9; heads 3”—4” high; southeastern. 3. I. imbricata. 
Heads spicate-paniculate ; leaves dentate. 4. I. ciliata. 
Heads spicate-paniculate, not subtended by leaves. 5. I. xanthiifolia. 


GENUs I. RAGWEED FAMILY. 339 


1. Iva frutéscens L. Marsh Elder. High- 
water Shrub. Fig. 4120. 


Iva frutescens L. Sp. Pl. 989. 1753. 
Iva oraria Bartlett, Rhodora 8: 26. 1906. 


Perennial, shrubby or herbaceous, somewhat 
fleshy; stem’ paniculately branched above, mi- 
nutely pubescent, or sometimes glabrous below, 
3°-12° high. Leaves oval, oblong, or oblong- 


wl} 
We 


lanceolate, all the lower ones opposite, short- ‘ <S W 

petioled, 3-nerved, acute or obtusish, serrate, nar- \\ We 

rowed at the base, the lower 4’-6’ long, 1’-2’ wide, Sp - 
the upper smaller and narrower, passing gradu- Sy" 

ally into those of the racemose inflorescence = 

which are much longer than the short-pedicelled \ 
heads; involucre depressed-hemispheric, its bracts 


about 5, orbicular-obovate, separate; fertile flow- 
ers about 5, their corollas tubular. 


Along salt marshes and on muddy sea-shores, 
Massachusetts to Florida and Texas, the northern 
plant (J. oraria) mainly broader-leaved and less 
shrubby than the southern. Jesuits’- or false Jesuits’- 
bark, July—Sept. 


2. Iva axillaris Pursh. Small-flowered Marsh 
Elder. Fig. 4121. 


Iva axillaris Pursh, Fl. Am. Sept. 743. 1814. 


» Perennial by woody roots; stems herbaceous, ascend- 
7 ing, glabrous or sparingly pubescent, simple or branched, 
1°-2° high. Leaves sessile, entire or very nearly so, 
obtuse, faintly 3-nerved, obovate, oblong, or linear- 
oblong, 4-13’ long, thick, somewhat fleshy, glabrous 
or pubescent, the lower opposite, the upper alternate 
and smaller, passing gradually into those of the in- 
florescence; heads mostly solitary in the axils of the 
leaves, 2-3” broad, short-peduncled; involucre hemi- 
spheric, about 12” high; its bracts about 5, connate at 
the base, or united nearly to the summit; pistillate 
flowers 4 or 5, their corollas tubular. 

In saline or alkaline soil, Manitoba and North Dakota to 


western Nebraska, New Mexico, British Columbia and Cali- 
fornia. May—Sept. 


Elder. Fig. 4122. 
Iva imbricata Walt. Fl. Car. 232. 1788. 


Perennial by woody roots, glabrous or nearly so 
throughout, fleshy; stem 1°-2° high, simple, or 
sparingly branched. Leaves all but the lowest 
alternate, sessile, oblong-spatulate, or lanceolate, ob- 
tusish, mucronulate, entire, or rarely serrate, obscurely 
3-nerved, the larger 1-2’ long, 3-5’ wide; heads 
about 4” broad, short-peduncled or nearly sessile, 
the upper often longer than their subtending leaves; 
involucre broadly campanulate, its bracts 6-9, not 
united, somewhat imbricated in 2 series; fertile 
flowers 2-4, their corollas tubular, the staminate 
ones much more numerous; chaff of the receptacle 
spatulate. 


On sandy sea-shores, southeastern Virginia to Florida 
and Louisiana. Bahamas; Cuba. July—Oct. 


340 _ AMBROSIACEAE. Vor. III. 


4. Iva ciliata Willd. Rough Marsh Elder. 
Fig. 4123. 

Iva annua Michx. Fl. Bor. Am. 2: 184. 1803. Not L. 

Iva ciliata Willd. Sp. Pl. 3: 2386. 1804. 


Annual, hispid-pubescent; stem simple, or 
branched, 2°-6° high. Leaves nearly all oppo- 
site, ovate, petioled, scarcely fleshy, puberulent 
beneath, acuminate at the apex, abruptly or 
gradually narrowed at the base, coarsely and ir- 
regularly dentate, 3-nerved, the lower 45’ long; 
heads spicate-paniculate, about 1” broad; spikes 
dense or interrupted, erect, 2’-8’ long; upper 
leaves linear-lanceolate, hispid, squarrose, much 
longer than the heads; bracts of the involucre 
3-5, distinct or united below, hispid; fertile flow- 
ers 3-5, their corollas slender; staminate flowers 
10-15. 


In moist soil, Illinois to Nebraska, south to Louisi- 
ana and New Mexico. Plant with the aspect of an 
Ambrosia. Aug.—Oct. 


ERA Gah? 
ASO Fas 
D (Ff y 


va xanthiifolia Nutt. Gen. 2: 185. 1818. 


Cyclachaena xanthiifolia Fresen. Ind. Sem. Hort. 
Franc. 4. 1836. 


Annual; stem much branched, pubescent or 
puberulent above, glabrous below, 3°-6° high. 
Leaves nearly all opposite, broadly ovate, long- 
petioled, acuminate at the apex, abruptly or 
gradually contracted at the base, coarsely and 
irregularly dentate, pale and canescent or pu- 
berulent beneath, roughish above, 3-ribbed, the 
lower often 6’ long and wide; inflorescence 
spicate-paniculate, terminal and axillary, ample, 
naked; heads sessile or minutely peduncled, 
1” broad or less; bracts of the involucre in 
2 series of 5, the outer ovate, the inner obovate 
or truncate, concave, subtending the usually 5 
achenes; corollas of the fertile flowers rudi- 
mentary or none; staminate flowers 10-15, their 
corollas obconic. 


In moist soil, or sometimes in waste places, 
Ontario to Assiniboia, south to Wisconsin, Ne- 3 
braska, New Mexico and Utah. In waste grounds a 
farther east. Plant with the aspect of a Cheno- | 
podium. July—Sept. 


2. AMBROSIA [Tourn.] L. Sp. Pl. 987. 1753. 


“Monoecious (rarely dioecious) branching herbs, with alternate or opposite, mostly lobed 
or divided leaves, and small heads of green flowers, the staminate spicate or racemose, the 
pistillate solitary or clustered in the upper axils. Involucre of the pistillate heads globose- 
ovoid or top-shaped, closed, 1-flowered, usually armed with 4-8 tubercles or spines; corolla 
none; stamens none; style-branches filiform; achenes ovoid or obovoid; pappus none. Invo- 
lucre of the staminate heads mostly hemispheric or saucer-shaped, 5-12-lobed, open, many- 
flowered; receptacle nearly flat, naked, or with filiform chaff; corolla funnelform, 5-toothed ; 
anthers scarcely coherent, mucronate-tipped; style undivided, penicillate at the summit. 
[The ancient classical name.] 


About 15 species, mostly natives of America. Besides the following, some 5 others occur in 
the southern and western United States. Type species: Ambrosia maritima L 


Sterile heads sessile; a lanceolate hispid lobe on inner border of involucre. 1. A. bidentata. 
Sterile heads short-pedicelled, involucre depressed-hemispheric. ; 
Leaves opposite, palmately 3—-5-lobed, or undivided ; receptacle naked. 2. A. trifida. 
Leaves opposite and alternate, 1-2-pinnatifid ; receptacle chaffy. i 
Annual; leaves thin; fruiting involucre spiny. 3. A. elatior. 


Perennial; leaves thick; fruiting involucre naked or tubercled. 4. A. psilostachya. 


GENUS 2. RAGWEED FAMILY. 


1. Ambrosia bidentata Michx. 


leaved Ragweed. Fig. 4125. 
Ambrosia bidentata Michx. Fl. Bor. Am. 2: 182. 1803. 


Annual, hirsute, usually much branched, very 
leafy, 1°-3° high. Leaves lanceolate, mainly 
alternate, sessile and somewhat cordate-clasping 
at the base, acuminate at the apex, I-nerved, 1’-3’ 
long, 2’-4” wide, usually with 1 or 2 sharp lobes 
at the base and a few minute sharp teeth above, 
or the upper ones quite entire, rough and hirsute 
or ciliate; spikes of staminate heads dense, 3-7’ 
long, their involucres turbinate, bearing a long 
lanceolate hispid reflexed lobe appearing like a 
bract on the inner border, their receptacles chaffy ; 
fertile heads » solitary, or clustered, oblong, 
long, bearing 4 sharp spines. 


4-angled, 3-4 


Prairies, Illinois to Missouri, Kansas, 


and Texas. July—Sept. 


Ambrosia trifida integrifolia T. & G. Fl. N. A. 2 


Lance- 


Louisiana 


2. Ambrosia trifida L. WHorse-cane. Bitter- 
weed. Great Ragweed. Fig. 4126. 


Ambrosia trifida L. Sp. Pl. 987. 1753. 
A, integrifolia Muhl.; Willd. Sp. Pl. 4: 375. 180s. 
: 290. 
1841. ; 
~ Annual, scabrous or hispid, or nearly glabrous, 
branched, 3°-17° high. Leaves all opposite, petioled, 
3-nerved, deeply 3-5-lobed, or undivided, the lobes 
lanceolate or ovate, serrate, acute or acuminate; 
lower leaves often 1° wide; racemes of sterile heads 
3-10’ long, their involucres saucer-shaped, 3-ribbed 
on the outer side, crenate-margined or truncate, their 
receptacles naked; fertile heads usually clustered in 
the axils of the upper bract-like leaves, turbinate to 
obovoid, 5-7-ribbed, conic-beaked, 3-4” long, each 
rib bearing a tubercle near the summit. 


In moist soil, Quebec to Florida, west to Manitoba, 
Nebraska, Colorado and New Mexico. Tall ambrosia. 


Richweed. Wild hemp. Horse-weed. Buffalo-weed. Hay-fever weed. July—Oct. 


3. Ambrosia elatior L. Ragweed. Roman 
Wormwood. Hogweed. Wild Tansy. 


Fig. 4127. 


Ambrosia elatior L. Sp. Pl. 987. 1753. 
Ambrosia artemisiaefolia L. Sp. Pl. 988. 


bescent. 


In dry soil, often a pernicious weed in cultivated 
fields, Nova Scotia to Florida, west to British Colum- 
bia and Mexico. Bermuda. Introduced into Europe asa 
weed. Consists of several slightly differing races. Also 
called bitterweed, stickweed, stammerwort, carrot-weed, 
black, or tassel-weed, hay-fever weed. July—Oct. 


1753- 


Annual, pubescent, puberulent or hirsute, panicu- 
lately branched, 1°-6° high. Leaves thin, 1-2-pin- 
natifid, petioled, 2-4’ long, the upper alternate, the 
lower mostly opposite, pale or canescent beneath, 
the lobes oblong or lanceolate, obtuse or acute; 
uppermost leaves of the branches sometimes linear- 
lanceolate and entire; racemes of sterile heads very 
numerous, 1-6’ long, the involucres hemispheric, 
crenate, the receptacle chaffy; fertile heads obovoid 
or subglobose, mostly clustered, 14’—2” long, short- 
beaked, 4-6-spined near the summit, sparingly pu- 


Wy 


342 . AMBROSIACEAE. Vor. III. 


4. Ambrosia psilostachya DC. Western Rag- 
weed. Fig. 4128. 


Ambrosia psilostachya DC. Prodr. 5: 526. 1836. 


Similar to the preceding species, but perennial by 
long rootstocks, the leaves thick, the pubescence stri- 
gose or hispid. Stems usually much branched, 2°-6° 
high, rather stout; leaves I-2-pinnatifid, 2’-5’ long, 
the lobes acutish; racemes of sterile heads several or 
numerous, 2-6’ long, the involucres campanulate, the 
receptacles chaffy ; fertile heads mostly solitary, ovoid 
or obovoid, reticulated, short-pointed, unarmed, or 


1" 


with about 4 short tubercles, pubescent, 12’-2” long. 


In moist open soil, Illinois to Saskatchewan, Texas, 
Mexico and California. July—Oct. 


3. GAERTNERIA Med. Act. Pal. 3: 244. 
1785. Phil. Bot. 45. 176¢ 
[FRANSERIA Cav. Icon. 2: 78. pl. 200. 1793.] 


Hispid or tomentose branching herbs, with the aspect of Ambrosias, sometimes woody at 
the base, with mostly alternate lobed or divided leaves, and small monoecious greenish heads 
of discoidal flowers, the staminate in terminal spikes or racemes, the pistillate solitary or 
clustered in the upper axils. Involucre of the pistillate heads ovoid or globose, closed, 
1-4-celled, 1-4-beaked, armed with several rows of spines and forming a bur in fruit; corolla 
none or rudimentary; style deeply bifid, its branches exserted; stamens none; achenes obo- 
void, thick, solitary in the cells; pappus none. Staminate heads sessile, or short-peduncled, 
their involucres broadly hemispheric, open, 5-12-lobed; receptacle chaffy; corolla regular, the 
tube ‘short, the limb 3-lobed; style undivided; anthers scarcely coherent; mucronate-tipped. 
[In honor of Joseph Gaertner, 1732-1791, German botanist. ] 


About 25 species, natives of America. In addition to the following, some 12 others occur in the 
western and southwestern United States. Type species: Xanthium fruticosum L.-f. 


Plant hirsute; annual; spines of the fruiting involucre long, flat. ; 1. G. acanthicarpa. 
Leaves densely white-tomentose beneath; spines short, conic; perennials. 
Leaves bipinnatifid. . 2. G. discolor. 
Leaves pinnately divided, the terminal segment large. 3. G. tomentosa. 


1. Gaertneria acanthicarpa ( Hook.) 
Britton. Hooker’s Gaertneria. 


Fig. 4129. 


Ambrosia acanthicarpa Hook. Fl. Bor. Am. 1: 309. 
1833. 

Franseria Hookeriana Nutt. Trans. Am. Phil. Soc. 
CLUinezeis45y Tsai. 

Gaertneria acanthicarpa Britton, Mem. Torr. Club 


5 332. 18945 
Franseria acanthicarpa Coville, Contr. Nat. Herb. 

4-120. 1895; 

Annual, erect or diffuse, paniculately branched, 
1°-2° high; stem hirsute or hispid. Lower and 
basal leaves slender-petioled, bipinnatifid, 2’—4’ 
long, the upper short-petioled or sessile, once- 
pinnatifid, or merely lobed; racemes of sterile 
heads usually numerous, 17-3’ long; fruiting 
involucres clustered in the axils, 3-4” long, 
commonly 1-flowered, armed with numerous 
long flat straight spines. 


In moist soil, Saskatchewan to western Ne- 
braska and Texas, west to British Columbia and 
California. Sand-bur. Recorded from Minnesota. July—Sept. : 


Gaertneria tenuifolia (A. Gray) Kuntze, a southwestern perennial species with pinnately 
dissected leaves, the fruiting involucre only about 1” long, is recorded as extending eastward into 
Kansas. : 


GENUS 3. RAGWEED FAMILY. 


2. Gaertneria discolor (Nutt.) Kuntze. 
White-leaved Gaertneria. Fig. 4130. 


Ambrosia tomentosa Nutt. Gen. 2: 186.- 1818. Not 
Gaertneria tomentosa (A. Gray) Kuntze. 


Franseria discolor Nutt. Trans. Am. Phil. Soc. (II.) 
7: 345. 1841. \ 


Gaerineria discolor Kuntze, Rev. Gen. Pl. 339. 1891. 


Erect or ascending from perennial rootstocks, 
branched, about 1° high. Leaves-nearly all bipin- 
natifid, petioled, densely white-tomentose beneath, 
green and pubescent or glabrate above, 2’-5’ long; 
sterile racemes narrow, commonly solitary, 1’-2’ 
long; fruiting involucres clustered in the axils, 7YV 
finely canescent, about 3” long, mostly 2-flowered, 
armed with short sharp conic spines. 


In dry soil, South Dakota, Nebraska, Wyoming, 
Kansas, Colorado and New Mexico. Aug.— —Sept. 


3. Gaertneria tomentosa (A. Gray) 
Kuntze. Woolly Gaertneria. 
Fig. 4131. 
ran eG tomentosa A. Gray, Mem. Am. Acad. 4: 8o. 
1849. 
G. tomentosa Kuntze, Rev. Gen. Pl. 339. 1891. 
Gaertneria Grayi A. Nelson, Bot. Gaz. 34: 35. 1902. 


Erect from a deep perénnial root, usually 
branched at the base, 1°-3° high. Leaves pin- 
nately lobed or divided, finely and densely to- 
mentose on both sides, or ashy above, the terminal 
segment lanceolate or oblong-lanceolate, acumi- 
nate, serrulate or entire, very much larger than 
the 2-6 rather distinct narrow lateral ones; sterile 
racemes solitary, 2’-4’ long; fruiting involucres 
selitary, or 2-3 together in the upper axils, ovoid, 
finely canescent or glabrate, 2-flowered, about 3” 
long, armed with subulate-conic, very acute, 
sometimes curved or hooked spines. 


On rich prairies and along rivers, western Nebraska, Kansas and Colorado. Aug.—Sept. 


4. XANTHIUM [Tourn.] L. Sp. Pl. 987. 1783. 


Monoecious annual branching coarse rough or spiny herbs, with alternate lobed or dentate 
leaves, and rather small heads of greenish discoid flowers, the staminate ones capitate- 
clustered at the ends of the branches, the pistillate axillary. Staminate heads with a short 
involucre of 1 to 3 series of distinct bracts; receptacle cylindric, chaffy; corollas tubular, 
5-toothed; anthers not coherent, mucronate at the apex; filaments monadelphous; style slen- 
der, undivided. Pistillate heads of an ovoid or oblong, closed involucre, covered with hooked 
spines, !-2-beaked, 2-celled, each cavity containing one obovoid or oblong achene; corolla 
none; pappus none; style 2-cleft, its branches exserted.« [Greek, yellow, from its yielding 
a yellow hair-dye.] 

About 15 species, of wide geographic distribution. Type species: Xanthium strumarium L. 
Leaves lanceolate, acute at both ends; axils bearing 3-divided yellow spines. 

. X. spinosum. 
Leav. es orbicular or broadly ovate, cordate to truncate at base; no axillary omits 
Bur, or its prickles, or both, more or less hispid-pubescent ; beaks incurved. 


Body of the bur ovoid to oval, twice as long as thick or shorter. 
Bur 1’ long or more, the prickles 4”-5” long. 2. X. speciosum. 


344 AMBROSIACEAE. Vor. III. 


“wy 
i 


Bur 10” iong or less, the prickles 2’—3” long. 
Bur densely prickly, its pubescence brown. 
Bur loosely prickly, its pubescence yellowish. 
Body of the bur oblong, more than twice as long as thick. 
Prickles longer than the diameter of the body of the bur. 
Prickles shorter than the diameter of the body of the bur. 


3. X. echinatum. 
4 
5 
Bur narrowly oblong. 6. X. pennsylvanicum. 
7 
8 


. X. glanduliferum, 


. X. inflexrum. 


Bur broadly oblong. . X. commune. 
Bur and its prickles glabrous, or merely puberulent ; beaks nearly straight. . X. americanum. 


1. Xanthium spindsum L. Spiny or 
Thorny Clotbur, Clotweed or Bur- 
weed. Pig i432: 

Xanthium spinosum L. Sp. Pl. 987. 1753. 


Stem pubescent or puberulent, much branched, 
ascending or erect, 1°-3° high. Leaves lanceo- 
late or ovate-lanceolate, acute or acuminate, 
lobed, or the upper entire, narrowed at the 
base, short-petioled, white-canescent beneath 
and on the whitish veins of the upper surface, 
2’-5’ long; axils each with a short-stalked 
3-pronged yellow spine nearly 1’ long; ripe 
fertile involucre (bur) oblong-cylindric, 4”-6” 
long, about 2” in diameter, pubescent, armed 
with short subulate rather inconspicuous beaks, 
and numerous glabrous spines about 1” long. 

In waste grounds, Maine to Ontario, Florida, 
Illinois, West Virginia, Missouri, Texas, New 
Mexico and California. Widely distributed as a 
weed in tropical America. Naturalized from 


Europe or Asia. Cocklebur. Dagger-cocklebur. 
Aug.—Nov. 


2. Xanthium speciosum Kearney. Great 
Clotbur. Fig. 4133. 


Xanthium speciosum Kearney, Bull. Torr. Club 
24: 574. 1897. 


Very stout, 3°-43° high. Stem sharply 
angled above; lower petioles 4’-6’ long; leaf- 
blades broadly triangular-ovate, the larger 6-8’ 
wide, 3-5-lobed, dentate, scabrous on both sur- 
faces; burs commonly clustered, oblong to 
ovoid-oblong, 1’ long or more, the stout beaks 
5-6” long, somewhat incurved, strongly hooked 
at the apex, equalling or a little longer than the 
dense subulate uncinate prickles, which are 
hispid to above the middle, and 4’-5” long. 


Moist and waste grounds, North Dakota to 
Wisconsin, Tennessee, Montana, Nebraska and 
Texas, and locally in waste places eastward. 
Aug.—Sept. 


GENUS 4. RAGWEED FAMILY. 


3. Xanthium echinatum Murr. Beach Clotbur. 
Fig. 4134. 

X. echinatum Murr. Comm. Goett. 6: 32, pl. 4. 1783. 

X. macuiatum Raf. Am. Month. Mag. 344. 1818. 

X. oviforme Wallr. Beitr. Bot. 1: 240. 1842. 


Stem rough, purplish or purple-blotched, 1°-2° high. 
Leaves firm, scabrous, with scattered short papillose 
hairs, obtusely toothed and lobed, somewhat resinous- 
glandular beneath; burs commonly clustered in the axils, 
ovoid to oval, 7-11” long, 4”-6” thick, glandular; prickles 
very dense, densely hispid from the base to the middle 
or beyond, subulate, hooked, the longer about 22” long, 
and about equalling the stout hispid beaks. 

Sea, lake and river beaches, occasionally in waste grounds, 


North Carolina to Nova Scotia, New York, Minnesota and 
North Dakota. - Recorded west to Saskatchewan. Aug.—Sept. 


345 


4. Xanthium glanduliferum Greene. 
Glandular Clotbur. Fig. 4135. 
Xanthium glanduliferum Greene, Pittonia 4: 61. 

i899. 

Similar to X. echinatum. Leaves very thick 
and scabrous with short stout papillae; burs 
oval, 5-8” long, 3’-43” thick, yellow, the 
prickles scattered, bristly-hispid nearly to the 
hooked apex, scarcely as long as the conic- 
subulate short-bristly beaks. 


In dry soil, North Dakota to Assiniboia, Brit- 
ish Columbia and Nebraska. Adventive in Mis- 
souri. June—Sept. 


Xanthium Macoiunii Britton, known only 
from Lake Winnipeg, Manitoba, differs by a 
longer bur, 10” long and 4” thick. 


5. Xanthium infléxum Mackenzie & Bush. 
Missouri Clotbur. Fig. 41306. 


Xanthium inflexum Mackenzie & Bush, Rep. Mo. Bot. 
Gard. 16: 106. 1905. 


Glabrate, or papillose-roughened above, 3°-43° 
high. Leaves long-petioled, broadly ovate, more 
or less cordate, mostly 3-lobed, crenate-dentate ; 
burs 1’ long or less, the body oblong, more than 
twice as long as thick, 3-33” in diameter, glan- 
dular-pubescent; prickles hooked, stiff, longer 
than the diameter of the bur, glandular-pubescent 
below, glabrous above; beaks stout, about 5” long, 
bent at the middle, strongly inflexed, hooked. 


Sandy river-bottoms, Courtney, Missouri. Aug.— 
Sept. 


AMBROSIACEAE. Vor. III. 


6. Xanthium pennsylvanicum Wallr. 
Pennsylvania Clotbur. Fig. 4137. 


Xanthium pennsylvanicum Wallr. Beitr. Bot. 1: 

236. 1842. 

Stem comparatively slender, smooth below, 
roughish above, 1°-3° high. Leaves thin, long- 
petioled, sharply toothed and some of them 
3-5-lobed, smoothish, or the upper surface 
scabrate, glandular; burs clustered in the axils, 
7-9” long, about one-third as thick, puberu- 
lent and resinous-glandular, and commonly 
with a few longer hairs; prickles numerous, 
subulate, hooked, more or less hispid or gla- 
brate, the longer ones nearly as. long as the 
diameter of the bur; beaks slender, but stouter 
than the prickles, incurved and hooked. 

Moist gorunds, Quebec to Minnesota, New 
York, Missouri and Colorado. Aug.—Oct. Re- 
ferred in our first edition to Xanthiwm canadense 
Mill., a name which has been variously applied to 
different plants by authors. 


7. Xanthium commune Britton. Cockle- 
bur or Clotbur. Fig. 4138. 


Xanthium commune Britton, Manual 912. 1901. 


Stem rather slender, 1°-2°, roughish. Leaves 
broadly ovate, more or less lobed, scabrous, espe- 
cially above; burs commonly solitary in the axils, 
oblong, 7”-12” long, half as thick, or less, the 
subulate-conic beaks slightly incurved, hooked at 
the apex, about as long as the subulate uncinate 
prickles, which are hispid to about the middle with 
brown hairs and shorter than the diameter of 
the bur. 


Moist grounds, Quebec to Alberta, Maryland, Mis- 
souri, Utah and Arizona. Aug.—Oct. 


8. Xanthium americanum Walt. Amer- 
ican Cocklebur or Burweed. 
Fig. 4139. 
X. americanum Walt. Fl. Car. 231. 1788. 
X. macrocarpum giabratum DC. Prodr. 5: 523. 


1836. 
X. glabratum Britton, Manual 912. 1got. 


Rough, 1°-63° high. Leaves slender-petioled, 
broadly ovate to orbicular, 3-ribbed and cor- 
date or cordate-reniform at the base, the lower 
often 10’ wide, the margins dentate, or more 
or less 3-5-lobed, both surfaces roughish and 
green; bur oblong, glabrous or merely puberu- 
lent, 6-9” long, about+3” in diameter, its 2 
sharp conical-subulate 2-toothed beaks straight 
or nearly so, equalling or slightly longer than 
the glabrous spines. 


Moist grounds, Ontario to Florida, Michigan, 
Tennessee and Kansas. Referred, in our first edi- 
tion, to the Old World X. strumarium L., from 
which it proves to be distinct. Sheep- or clot-bur. 
Button-bur. Small or lesser burdock. Aug.—Oct. 


Xanthium orientale L. (X. canadense Mill.), an 
Old World tropical species, is naturalized in the 
West Indies. 


Kry To GENERA. THISTLE FAMILY. 347 


Family 46. COMPOSITAE Adans. Fam. Pl. 2: 103. 1763. 
THISTLE FAMILY, 


Herbs, rarely shrubs (some tropical forms trees), with watery or resinous 
(rarely milky) sap, and opposite alternate or basal exstipulate leaves. Flowers 
perfect, pistillate, or neutral, or sometimes monoecious or dioecious, borne cn a 
common receptacle, forming heads, subtended by an involucre of few to many bracts 
arranged in one or more series. Receptacle naked, or with chaffy scales subtend- 
ing the flowers, smooth, or variously pitted or honeycombed. Calyx-tube com- 
pletely adnate to the ovary, the limb (pappus) of bristles, awns, teeth, scales, or 
crown-like, or cup-like, or wanting. Corolla tubular, usually 5-lobed or 5-cleft, the 
lobes valvate, or that of the marginal flowers of the head expanded into a ligule 
(ray ) ; when the ray-flowers are absent the head is said to_be discoid; when present, 
radiate ; the tubular flowers form the disk. Stamens usually 5, borne on the corolla 
and alternate with its lobes, their anthers united into a tube (syngenesious ), often 
appendaged at the apex, sometimes sagittate or tailed at the base; pollen-grains 
globose, often rough or prickly. Ovary 1-celled; ovule 1, anatropous; style of 
fertile flowers 2-cleft, its branches variously appendaged, or unappendaged ; stigmas 
marginal; style of sterile flowers commonly undivided. Fruit an achene. Seed 
erect ; endosperm none; embryo straight ; hypocotyl inferior. 


; About 800 genera and not less than 10,000 species, of wide geographic distribution. The family 
is also known as CARDUACEAE, AGGREGATAE, and by the English name of Asterworts. In Kuhnia, the 
anthers are distinct, or nearly so. 


* Anthers not tailed at the base; stigmatic lines of the style-branches only at the base, or not extending 
beyond the middle; flowers all tubular and perfect, never yellow; rays none. 


Style-branches filiform or subulate, hispidulous ; receptacle naked. Tribe 1. VERNONIEAE. 

Style-branches thickened upward, obtuse, papillose. Tribe 2. EUPATORIEAE. 

** Anthers tailed at the base, unappendaged at the tip; heads small; rays none (except in Inula where they 
are large, yellow). Tribe 4. INULEAE. 


*** Anthers not tailed at the base; stigmatic lines of the style-branches in the perfect flowers extending to 
the summit; flowers tubular only, or tubular and radiate, often yellow. 


1. Receptacle naked (see also Nos. 85-88 of Tribe 7). 
a. Bracts of the involucre usually well imbricated. 


Style-branches of the perfect flowers flattened, with terminal appendages. Tribe 3. ASTEREAE. 
Style-branches truncate, or with hairy tips. Tribe 6. HELENIEAE. 
b. Bracts little imbricated, or not at all; pappus of soft bristles. 
Tribe 8. SENECIONEAE. 
2. Receptacle chaffy (except in Nos. 85-88). 
Bracts of the involucre herbaceous or foliaceous ; not scarious. Tribe 5. HELIANTHEAE. 
Bracts of the involucre dry, scarious. Tribe 7. ANTHEMIDEAE. 
**** Anthers long-tailed at the base, with elongated appendages at the tip; heads large; rays none (in 
Centaurea often with enlarged marginal flowers); bracts imbricated. 
Tribe 9. CYNAREAE. 


Tribe 1. VERNONIEAE. 


Pappus double, the inner of rough capillary bristles, the outer of scales, or short bristles; heads 
not glomerate. 1. Vernonia. 
Pappus a single series of bristles ; heads glomerate, subtended by sessile bracts. 2. Elephantopus. 


Tribe 2. EUPATORIEAE. 
* Achenes 3-5-angled, not ribbed. 


Pappus of 5 broad obtuse scales ; aquatic herb with linear whorled leaves. 3. Sclerolepis. 
Pappus of numerous capillary bristles. 
Involucre of more.than 4 bracts; erect herbs. 4. Eupatorium. 
Involucre of 4 bracts; our species twining herbaceous vines. 5. Mikania. 


** Achenes 8-10-ribbed, or 8-10-striate. 
i. bracts . the involucre in several series, the outer successively shorter. 
Bracts of the involucre strongly striate-nerved ; heads panicled or corymbed in our species. 


Pappus-bristles rough or serrate; involucral bracts numerous. 6. Coleosanthus. 
Pappus-bristles plumose ; involucral bracts few. 7. Kuhma. 
Bracts of the involucre faintly striate, if at all; heads spiked or racemed. 8. Lacinaria. _ 
2. Bratts of the involucre in only 2 or 3 series; all nearly equal. 
i 9. Triltsa, 


Tribe 3.. ASTEREAE. 
A. Ray-flowers yellow (white in one species of Solidago), or wanting; plants not dioecious. 
* Pappus of scales, or awns, or wanting, never of numerous capillary bristles. 


Heads small, not over 2” high; leaves narrowly linear, entire. 
Perennial herbs, or shrubs; all the flowers fertile. 10. Gutierrezia. 


348 COMPOSITAE. Vo. III. 


Annual herbs; disk-flowers sterile. L 11. Amphiachyris. 
Heads large, showy; leaves oblong to lanceolate, spinulose-dentate. 12. Grindelia. 
** Pappus of either the radiate or tubular flowers, or both, of numerous capillary bristles, with or’ without 
an outer series of shorter ones, or of scales. 


+ Pappus, at least of the disk-flowers, double, an inner series of capillary bristles, and an outer one 

of scales or short bristles; heads large. 
Achenes of the ray-flowers thick, the pappus obsolete, or of a few caducous bristles; achenes of the 

disk-flowers flat. : 13. Heterotheca. 

Achenes of both ray-flowers and disk-flowers flattened. 14. Chrysopsis. 

tt Pappus wholly of capillary bristles. 

1. Heads wholly of disk-flowers (in our species) ; rays none; leaves narrowly linear. 
a. Perennial herbs; style-tips not exserted; eastern. 15. Chondrophora. 
b. Shrubs; style-tips mostly exserted ; western. 
Involucral bracts gradually narrowed to the tip, keeled, chartaceous. 16. Chrysothamnus. 
Involucral bracts cuspidate, flat, herbaceous. 17. Oonopsis. 
2. Heads with both ray-flowers and disk-flowers (in our species). 
a. None of the leaves cordate; pappus mostly as long as the achene, or longer. 

Leaves bristly-serrate or pinnatifid. 


Achenes glabrous; pappus-bristles deciduous. 18. Prionopsis. 
Achenes white-tomentose ; pappus-bristles persistent. 19. Sideranthus. 
Leaves entire, or toothed, not bristly-serrate. 
Leaves narrow, coriaceous, evergreen, entire; low western undershrubs. 20. Stenotus. 
Annual or perennial herbs; leaves not evergreen. 
Pappus-bristles unequal; heads loosely panicled; annual. 21. Isopappus. 


Pappus-bristles equal; heads variously clustered ; perennial herbs. 
Receptacle alveolate ; rays mostly not more numerous than the disk-flowers. 
; ; 22. Solidago. 
Receptacle fimbrillate; rays more numerous than the disk-flowers; heads corymbose- 
paniculate. 23. Euthamia. 
b. Lower and basal leaves cordate; pappus-bristles shorter than the achene. 
24. Brachychaeta. 
B. Ray-flowers present, not yellow in any of our species. 
* Pappus a mere crown, or of a few awns or bristles, or wanting, never of numerous capillary bristles. 
1. Receptacle conic. 
Outer bracts of the involucre shorter than the inner; achenes prismatic. 25. Aphanostephus. 
Bracts of the involucre all about equal; achenes flattened. 26. Bellis. 
2. Receptacle flat, or somewhat convex. 


Achenes of the disk-flowers compressed, of the ray-flowers 3-angled; perennial herbs with large 


heads. 27. Townsendia. 
Achenes fusiform, terete; annual herbs with small heads. 28. Chaetopappa. 
Achenes obovate, flattened, with thickened or winged margins; perennials. 29. Boltonia. 


** Pappus of numerous capillary bristles. 
1. Pappus a single series of capillary bristles; sometimes with an outer series of shorter ones. 
a. Bracts of the involucre in 2 to many series. 
Bracts mostly in 2-5 series; teeth or lobes of the leaves not bristle-tipped. 


Involucre narrow, its bracts firm ; rays few, white. 30. Sericocarpus. 
Involucre turbinate to hemispheric, bracts mostly thin; rays usually numerous, white to purple. 
Disk-flowers turning red, purple, or brown. 31. Aster. 


Disk-flowers permanently white; leaves subulate to spatulate, crowded, rigid. 
32. Leucelene. 

Involucre turbinate ; rays not exceeding the mature pappus, or none. 33. Brachyactis. 
Bracts in many series; teeth or lobes of the leaves bristle-tipped in our species. 34. Machaeranthera. 

b. Bracts of the involucre mostly in 1 or 2 series, very narrow; heads mostly long-peduncled. 
Rays longer than the diameter of the disk. 35. Erigeron. 
Rays not longer than the diameter of the disk; heads very small, panicled. 36. Leptilon. 

2. Pappus distinctly double, the inner bristles long, the outer shorter. : 

Leaves lanceolate, ovate, or obovate; rays white. 37. Doellingeria. 
Leaves narrowly linear; rays violet. 38. Ionactis. 


C. Ray-flowers none; dioecious shrubs; pappus capillary. : 
39. Baccharis. 


Tribe 4. INULEAE. 
* Heads small, rays none; flowers white, or whitish. 


1. Receptacle chaffy. ; 
Receptacle convex ; pappus none. } ; 40. Filago. 
Receptacle subulate ; pappus of the inner flowers of rough capillary bristles. 41. Gifola. 
2. Receptacle naked. 
: a. Pappus, at least that of pistillate flowers, of capillary bristles. 
Bracts of the involucre not scarious ; plants pubescent or glabrous, not woolly. 42. Pluchea. 
Bracts of the involucre scarious, mostly white or pink; plants woolly. 
Plants dioecious, or polygamo-dioecious. 


Pappus-bristles of staminate flowers thickened above. 43. Antennaria. 
Pappus-bristles not thickened ; stem leafy. 44. Anaphalis. 
Plants not dioecious; flowers all fertile. 45. Gnaphalium. 


b. Pappus none; leaves broad, alternate, woolly beneath. 
46. Adenocaulon. 
** Heads large; ray-flowers yellow. 47. Inula. 


Key To GENERA. THISTLE FAMILY. 


Tribe 5. HELIANTHEAE. 


A. Disk-flowers perfect, but sterile. 


Achenes thick, short, not flattened ; pappus none. 
Achenes merely subtended by the inner involucral bracts. 
Achenes embraced and enclosed by the inner involucral bracts. 
Involucral bracts unarmed. 
Involucral bracts armed with hooked prickles. 
Achenes flattened. 
Ray-flowers in 2 or 3 series; achenes falling away free. 


48. Polymnia. 


349 


49. Melampodium. 
50. Acanthospermum. 


51. Silphium. 


Ray-flowers in 1 series; achenes adnate to 2 or 3 scales of the receptacle, falling away with them. 


Rays large, yellow. 
Leaves opposite and basal. os 
Leaves alternate. 


Pappus none, or of 2 caducous awns. gee 
Pappus a persistent irregularly cleft crown. 54. 
Rays small, the head appearing discoid. ; 55: 


B. Disk-flowers fertile. 


* Ray-flowers persistent upon the achenes. 


Achenes compressed, or 3-angled; leaves entire. 56. 

Ackenes short, thick; leaves toothed. Sie 
** Ray-flowers deciduous, or none. 

{ Pappus a cup, or crown, or of a few teeth, awns, or bristles. 


1. Achenes, at least those of disk-flowers, not compressed (except in Ratibida 
scales of the receptacle mostly concave, or clasping. 


Scales of the receptacle small, awn-like or bristle-like ; rays white, short. 58. 


Scales of the receptacle broad, larger. 
Involucre of 4 large somewhat united bracts. 59. 
Involucre of several or numerous separate bracts. 
Receptacle conic, or columnar. 
Ray-flowers fertile, or wanting; leaves opposite. 60. 
Ray-flowers sterile, or neutral; leaves mostly alternate. 
Rays yellow. 
Achenes 4-angled or terete. 


Achenes 4-angled. 61. 
Achenes terete; leaves cordate-clasping. 62. 
Achenes compressed, winged. 63. 
Rays rose-purple or yellow. 64. 


Receptacle flat, or convex (low-conic in species of Nos. 66 and 68). 


Low fleshy sea-coast shrubs. 65. 


Tall herbs, not fleshy. 


Achenes not much flattened, not winged, nor margined. 66. 
Achenes of disk-flowers flattened and margined, or winged. 
Involucre of a few deflexed bracts. 67. 
Involucre of 2 series or more of appressed or spreading bracts. 
Perennials ; bracts erect or appressed. 68. 
Annuals; bracts spreading. 69. 


Chrysogonum. 


Berlandiera. 
Engelmannia. 
Parthenium. 


Crassina. 
Heliopsis. 


and Phaethusa) ; 


Verbesina. 


Tetragonotheca. 


Spilanthes. 


Rudbeckia. 
Dracopis. 
Ratibida. 
Echinacea. 
Borrichia. 
Helianthus. 
Ridan. 


Phaethusa. 
Ximenesia. 


2. Achenes very flat; scales of the receptacle flat, or but slightly concave. 


a.. Bracts of the involucre all separate. 


Pappus of 2 short teeth or awns, or a mere border, or none. 70. 
Pappus of 2-6 awns or teeth, upwardly or downwardly barbed or hispid. 
Achenes flat, or angled. WX. 
Achenes terete ; aquatic, the submerged leaves filiform-dissected. vie. 


b. Inner bracts of the involucre united to about the middle. 73. 
tt Pappus of numerous scales. 
Leaves opposite, toothed; ray-flowers fertile; rays small. 74. 
Leaves alternate, entire. 


Rays large, neutral; receptacle deeply honey-combed. 7p 
Rays none; scales of the receptacle narrow, rigid. 76. 


Tribe 6. HELENIEAE. 


Coreopsis. 
Bidens. 
Megalodonta. 
Thelesperma. 
Galinsoga. 


Endorima. 
Marshallia. 


A. Ray-flowers persistent on the achenes, falling away with them, papery. 


: i 77: 
B. Ray-flowers deciduous, or wanting. 
* Plants not dotted with oil-glands. 


Psilostrophe. 


a. Pappus none. 


78. Flaveria. 


b. Pappus present (in all our species), of separate scales or bristles. 
1. Bracts of the involucre petal-like, colored, their margins and apices scarious. 


Leaves, at least the lower, pinnately parted, or pinnatifid; rays none; corolla-lobes of disk-flowers 
79. Hymenopappus. 


ovate. 


Leaves entire; rays present, or none; corolla-lobes of disk-flowers linear. 


80. Othake. 


2. Bracts of the involucre herbaceous, not scarious-tipped, nor petal-like, appressed, or spreading. 


Receptacle naked. 
Bracts of the involucre appressed. 
Achenes 4-angled, linear or oblong. 
Achenes 5—10-ribbed or 5—10-angled, top-shaped. 
Involucral bracts separate to the base. 


81. Picradeniopsis. 


82. Tetraneurts. 


350 COMPOSITAE. Vot. IIT. 
Outer involucral bracts connate. 83. Hymenoxys. 

Bracts of the involucre spreading, or reflexed at maturity. 84. Helenium. 

Receptacle with bristle-like chaff. 85. Galliardia. 
** Plants dotted with oil-glands, especially the leaves and involucre. 

Involucral bracts more or less united; style-branches of the disk-flowers elongated. 

Involucral bracts united at the base. 86. Boebera. 

Involucral bracts united high up into a cup. 87. Thymophylla, 
Involucral bracts separate ; style-branches of the disk- flowers very short. 88. Pectis. 

Tribe 7. ANTHEMIDEAE. 
* Receptacle chaffy. 
Achenes flattened ; involucre obovoid to campanulate; heads small. 89. Achillea. 
Achenes terete; involucre hemispheric; heads large. 90. Anthemis. 
** Receptacle not chaffy, naked, or sometimes hairy. 
1. Ray-flowers usually present, sometimes wanting. 
Receptacle flat to hemispheric ; bracts of the involucre in several series. ot. Chrysanthemum. 
Receptacle conic to ovoid; bracts in few series. 92. Matricaria. 
2. Ray-flowers none; heads small. ; 
Heads corymbed; pappus a short crown; flowers yellow. 93. Tanacetum. 
Heads racemose, spicate or panicled; pappus none. 94. Artemisia. 
Tribe 8. SENECIONEAE. 

Leaves all basal; heads on scapes. 

Heads solitary ; flowers yellow. 95. Tussilago. 

Heads corymbed; flowers white or purple. 96. Petasites. 
Leaves opposite; rays yellow. 

Involucre of several thin herbaceous bracts. 97. Arnica. . 

Involucre of 4 or 5 broad fleshy bracts. 98. Haploesthes. 
Leaves alternate. 

Flowers white, whitish or pinkish; rays none. : 
Marginal flowers pistillate; disk-flowers perfect. 99. Erechtites. 
Flowers all perfect. ‘ 

Involucre of about 5 bracts; sap milky. 100. Mesadenia. 
Involucre of about 12 bracts and several smaller outer ones. 101. Synosma. 
Flowers yellow; ray-flowers mostly present. 102. Senecio. 
Tribe 9. CYNAREAE. 
* Achenes inserted on the receptacle by their bases, not oblique. 
Receptacle densely bristly. 

Filaments separate. : j 
Involucral bracts hooked at the tip; leaves not bristly. 103. Arctium. 
Involucral bracts not hooked; leaves bristly. Vea 

Pappus-bristles plumose. 104. Cirsium. 

Pappus-bristles not plumose. 105. Carduus. 

Filaments united below. 106. Mariana. 
Receptacle fleshy, not bristly. 107. Onopordon. 
** Achenes obliquely inserted on the receptacle. 
Heads not subtended by bristly leaves; involucral bracts often bristly. 108. Centaurea, 
Heads sessile, subtended by bristly leaves. 109. Cnicus. 
Xx 
1. VERNONIA Schreb. Gen. Pl. 2: 541. 1791. 


Erect branching perennial herbs, or. some tropical species shrubby, with alternate (very 
rarely oposite), in our species sessile leaves, and discoid cymose-paniculate heads of purple 
pink or white tubular flowers. Involucre hemispheric, campanulate or oblong-cylindric, its 
bracts imbricated in several or many series. Receptacle flat, naked. Corolla regular, 5-cleft. 
Anthers sagittate at the base, not caudate. Style-branches subulate, hispidulous their whole 
length. Achenes 8-10-ribbed, truncate. Pappus of our species in 2 series, the inner of 
numerous roughened capillary bristles, the outer of much shorter small scales or stout 
bristles. [Named after William Vernon, English botanist. ] 


More than 500 species, of wide distribution in warm-temperate regions, most abundant in South 
America. Besides the following, several others occur in the southern and southwestern United 
States. Type species: Serratula noveboracensis L. / 

Heads large, nearly 1’ broad; involucral bracts with long filiform tips. 
Heads smaller, 6” broad or less. 
Involucral bracts with filiform tips. 
Leaves lanceolate, relatively narrow; pappus purple, rarely green. 2: 
Leaves oval to lanceolate, relatively broad ; pappus yellowish. Bs 
Involucral bracts acute or obtuse, not filiform- ‘tipped. 
Leaves linear, 1-nerved. 4. 
Leaves lanceolate to oblong-lanceolate. 
Leaves glabrous or merely puberulent beneath. 
Heads loosely cymose. 5. V. altissima. 
Heads densely cymose. 6. V. fasciculata. 
Leaves tomentose beneath. : 
Involucral bracts squarrose, acuminate. V. Baldwinii. 
Involucral bracts obtuse or acute, appressed. . V. missurica. 


1. V.crinita. 


V.noveboracensis. 
V. glauca. 


V. marginata. 


om 


GENUS I. THISTLE FAMILY. 351 


op NASA 
eaeaes _ a We 
1. Vernonia crinita Raf. Great Iron WMi2 4 A 


weed. Fig. 4140. 


\! WIA 
SG 


BNW 
WH 


V. crinata Raf. New Flora N. A. 4:77. 1836. 


Vernonia arkansana DC. Prodr. 7: 264. 1838. 
NE MAUNA i 
i LONE fe 
St) t Wate fa 


x Ii 
= i] 
WZ 


Cacalia arkansana Kuntze, Rev. Gen. Pl. 969. 1891. 


Stout, glabrate or finely rough-pubescent, 8°- 
12° high, simple or little branched. Leaves nar- 
rowly lanceolate, finely denticulate, acuminate, 
3-12’ long, 3-12” wide; heads stout-peduncled, 
the peduncles thickened above; involucre hemi- 
spheric, 9”-12” broad, 50-80-flowered; bracts 
green, or the upper reddish, very squarrose, all 
filiform-subulate from a broader base and equal- 
ling the head, the inner ones somewhat wider 
below; achenes glabrous or hispidulous on the 
ribs; pappus purplish. 


On prairies and along streams, Missouri to Kan- 
sas and Texas. Aug.—Oct. 


\y (N y hi fi y d 
X UN i NY f 
(7 NG Ye 
i 


2. Vernonia noveboracénsis (L.) Willd. New York Iron-weed. Flat Top. 


Fig. 4141. 
Serratula noveboracensis L. Sp. Pl. 818. 1753. 
V. noveboracensis Willd. Sp. Pl. 3: 1632. 1804. 
C. noveboracensis Kuntze, Rev. Gen. Pl. 323. 1891. 
Vernonia noveboracensis tomentosa Britton, Mem. 
Torr. Club 5: 311. 1894. 


Roughish-pubescent or glabrate, 3°-9° high. 
smn Leaves lanceolate or narrowly oblong, serrulate, 
| Vs 3-10 long, 5-12” wide, acuminate or acute; 
LZ heads peduncled; involucre hemispheric, 20-40- 
Vion flowered, 4-5” in diameter; bracts brownish- 
purple or greenish, ovate or ovate-lanceolate, with 
subulate spreading tips usually twice or three 
times their own length, or some of the lower 
linear-subulate, the upper sometimes merely acute; 
flowers deep purple, rarely white; achenes his- 
pidulous on the ribs; pappus purple or purplish, 
rarely green. 

In moist soil, Massachusetts to Pennsylvania, 
North Carolina, West Virginia, Mississippi and Mis- 
Sg Erroneously recorded west to Minnesota. July— 

ept. 


A 
7h) 
NY 


3. Vernonia glauca (L.) Britton. Broad- mal 
ees YY Ne 
leaved Iron-weed. Fig. 4142. VAN, 
Serratula glauca L. Sp. Pl. 818. 1753. 
Vernonia noveboracensis var. latifolia A. Gray, 
Syn. Fl. 1: Part 2, 89. 1884. 
Vernonia glauca Britton, Mem. Torr. Club 5: 


BII. 1894. 


Slender, glabrous or finely puberulent, 2°-5° 
high. Leaves thin, the lower broadly oval or 
slightly obovate, sharply serrate, acute or acu- 
minate, 4-7’ long, 1’-23’ wide, the upper nar- 
rower and more finely toothed; inflorescence 
loosely branched; heads slender-peduncled, 10- 
20-flowered; involucre campanulate, 3’-4” 
broad; bracts ovate, with filiform tips, ap- 
pressed; achenes minutely hispidulous; pappus 
yellowish. 


In woods, Pennsylvania and Maryland to 
Georgia and Alabama. Southern plants pre- 
viously referred to this species prove to be dis- 
tinct. Aug.—Sept. 


COMPOSITAE. Vor. III. 


4. Vernonia marginata (Torr.) Raf. 
James’ Iron-weed. Fig. 4143. 


Vernonia altissima var. marginata Torr. Ann. 
Lyc: N- Y222 210. (1827 

eons marginata Raf. Atl. Journ. 1: 146. 
1832. 

Vernonia Jamesii T. & G. Fl. N. A. 2: 58. 1841. 

Cacalia marginata Kuntze, Rev. Gen. Pl. 968. 
1891. 


Glabrous or very nearly so, 1°-3° high. 
Leaves linear or linear-lanceolate, minutely 
denticulate, 1-nerved, firm, punctate, 2-5’ long, 
13-3” wide, acuminate; inflorescence rather 
loose; heads slender-peduncled; involucre 
campanulate or turbinate, 15-30-flowered, 4’— 
6” broad; bracts ovate or oval, acute, mucro- 
nate or obtusish, purplish, somewhat pubes- 
cent, appressed; achenes nearly glabrous, or 

. somewhat pubescent; pappus brownish. 


Prairies, Nebraska and Kansas to Texas and 
New Mexico. Autumn. 


Ly ox, OP 


i NA 
yas MN). |! 
Wy we 


5. Vernonia altissima Nutt. 
Iron-weed. Fig. 4144. 
Vernonia altissima Nutt. Gen. 2: 134. 1818. 


Vernonia maxima Small, Bull. Torr. Club 27: 
280. 1900. 


Glabrous or nearly so, 5°-10° high. Leaves 
thin, lanceolate, sometimes broadly so, usu- 
ally long-acuminate, finely serrate, 4-12’ 
long, 2’-12’ wide, glabrous on both surfaces, 
or puberulent beneath; inflorescence at 
length loosely branched and open; heads 
short-peduncled or some of them sessile; 
involucre campanulate or turbinate, 2’-3” 
broad, 15-30-flowered; bracts obtuse or 
mucronate, more or less ciliate, appressed; 
achenes slightly hispidulous; pappus pur- 
plish. 

In moist soil, New York to Florida, Illinois, 
Michigan, Missouri, Kentucky and Louisiana. 
July-Sept. Included in our first edition in V. 
gigantea (Walt.) Britton, of the Southern 
States. te . , 

6. Vernonia fasciculata Michx. 


Western Iron-weed. Fig. 4145. 


Vernonia fasciculata Michx. Fl. Bor. Am. 2: 
94. 1803. 


Cacalia fasciculata Kuntze, Rev. Gen. Pl. 970. 
1891. 


Glabrous, or puberulent above, 2°-6° high. 
Leaves firm, lanceolate or linear-lanceolate, 
long-acuminate, 3-6’ long, 2”—4” wide, gla- 
brous or nearly so on both surfaces; inflor- 
escence usually compact; heads short-pedun- 
cled, or some of them sessile; involucre 
campanulate, 2’-3” broad, 20-30-flowered ; 
bracts all appressed, ovate or oval, acute, 
ciliate, or sometimes pubescent; achenes gla- 
brous, or a little pubescent; pappus purple. 


In moist soil or on prairies, Ohio to Minne- 
sota, Nebraska, Kansas and Oklahoma. South- 
ern plants formerly referred to this species 
prove to be distinct. July—Sept. 


Vernonia corymbosa Schwein., ranging 
from Manitoba to western Nebraska, has 
broader leaves but is otherwise similar. 


GENUS I. THISTLE FAMILY. 353 


7. Vernonia Baldwinii Torr. Baldwin’s 
Iron-weed. Fig. 4146. 

V. Baldwinii Torr. Ann, Lyc. N. Y. 2: 211. 1827. 

Cacalia Baldwinii Kuntze, Rev. Gen. Pl. 969. 1891. 


Vernonia interior Small, Bull. Torr. Club 27: 279. 
1900. 


V. interior Baldwinii Mack. & Bush, Fl. Jackson Co. 

190. 1903. 

Stout, 2°-54° high, finely and densely tomen 
tose-pubescent. Leaves lanceolate or oblong- 
lanceolate, acuminate or acute at the apex, sharply 
serrate, 4-8’ long, 2’-2’ wide, scabrate above, 
densely tomentulose beneath; heads stout-pedun- 
cled, 15-30-flowered; involucre hemispheric, 3’— 
4” broad; bracts ovate, the acute tips recurved 
or spreading; pappus purple. 


In dry soil, Iowa to Missouri, Nebraska, Kansas 
and Texas. July—Sept. 


8. Vernonia misstrica Raf. Drummond’s Iron-weed. Fig. 4147. 


V. missurica Raf. Herb. Raf. 28. 1833. 

Vernonia altissima var. grandiflora A. Gray, 
Syne Ble ne Part 2; 90.) 51664. 

Vernonia Drummondii Shuttlw.; Werner, 
Journ. Cine. Soc. Nat. Hist. 16: 171. 
1894. 

V. illinoensis Gleason, Bull. N. Y. Bot. 
Gards4'> 21a.) 2906: 


Stout, densely tomentose, 3°-5° high. 
Leaves lanceolate to narrowly oblong, 
acuminate, finely serrate, 3’-6’ long, 
3’-1}’ wide, scabrous above, densely pu- 
bescent beneath; inflorescence usually 
compact; heads short-peduncled; invo- 
lucre hemispheric or short-cylindric, 2’- 
6” in diameter, 30-60-flowered; bracts 
purplish, appressed, ovate, obtuse, acute, 
or mucronate, more or less floccose- 
pubescent or ciliate; achenes glabrous 
or a little pubescent; pappus purplish to 
tawny. 


Prairies, Ontario and Ohio to Illinois, 
Kentucky, Missouri and Texas. Autumn. 


2. ELEPHANTOPUS [Vaill.] L. Sp. Pl. 814. 1753. 


Perennial rigid pubescent herbs, with alternate or basal, simple pinnately-veined leaves, 
and in our species glomerate bracted heads of blue or purple flowers in branching corymbs. 
Heads discoid, 2-5-flowered. Involucre compressed, oblong, 1ts chaffy bracts imbricated in 
about 2 series, the 4 outer bracts shorter. Bracts of the glomerules large, foliaceous. Recep- 
tacle small, naked. Corolla nearly regular, 5-lobed, but a little deeper cleft on the inner 
side. Achenes 10-ribbed, truncate. Pappus of rigid persistent awn-like scales or bristles in 
I or 2rows. [Greek, Elepihant’s-foot.] 


About 14 species, natives of tropical or warm regions. Besides the following, another occurs 
in the southern United States. Type species: Elephantopus scaber L. 


Stem and branches leafy. 1. E. carolinianus. 
Stem scapiform, naked, or with 1 or 2 leaves. 
Leaves oblong or oblanceolate, 9”—2’ wide; heads 4” long. 2. E. nudatus. 
Leaves ovate, oval, or obovate, 2’-4’ wide; heads 6” long. 3. E. tomentosus. 


23 


354 COMPOSITAE. Vo. III. 


1. Elephantopus carolinianus Willd. 


Carolina Elephant’s-foot. 
Fig. 4148. 
Elephantopus carolinianus Willd. Sp. -Pl. 3: 

2390. 1804. 

Erect, hirsute-pubescent, or glabrate 
above, corymbosely branched above, 1°-3° 
high. Leaves oval, ovate, or obovate, thin, 
the lower rather abruptly narrowed into 
margined petioles, obtuse, crenate-dentate, 
3-8’ long, 2’-4’ wide, the upper smaller, 
narrower and sessile; glomerules, includ- 
ing the bracts, nearly 1’ broad; scales of 
the pappus lanceolate-subulate, gradually 
narrowed into a long slender awn. 

In dry woods, southern New Jersey and 


Pennsylvania to Florida, Illinois, Kansas and 
Texas. Aug.—Sept. 


2. Elephantopus nudatus A. Gray. 

Smoothish Elephant’s-foot. 
Fig. 4149. 

Elephantopus nudatus A. Gray, Proc. Am. 

Acad. 15: 47. 1880. 

Erect, rather stout, appressed-pubescent, 
or glabrate, 1°-2° high. Leaves oblong or 
oblanceolate, obtuse at the apex, gradually 
narrowed at the base, crenate or repand, 
2’-10’ long, 9’’—2’ wide, the basal ones usually 
much larger than those of the stem and 
branches, or these usually few and bract- 
like; glomerules 6’—-9” broad; heads about 
4” long; scales of the pappus ovate-trian- 
gular, abruptly narrowed into the awn. 


In sandy woods, Delaware and Maryland to 
Florida, west to Arkansas and Louisiana. 
Aug.—Sept. 
<\) 


Wy} 
whys RY 


ye 


3. Elephantopus tomentosus L. 
Woolly Elephant’s-foot. Tobacco- 
weed. Fig. 4150. 


Elephantopus tomentosus L. Sp. Pl. 814. 1753. 


‘4 ovate, oval, oblong, or obovate, obtuse, ses- 
sile, narrowed at the base, usually silky- 
pubescent beneath, crenate-dentate, 4-0’ long, 
2’-4’ wide; glomerules 1’-12’ broad; heads 
about 6” long; scales of the pappus triangu- 
lar-subulate, gradually narrowed into the 
awn. 

In moist soil, Virginia to Florida, west to 


Kentucky, Arkansas and Louisiana. Called 
also devil’s-grandmother. Aug.—Sept. 


GENUS 3. THISTLE, FAMILY. 355 


3. SCLEROLEPIS Cass. Bull. Soc. Philom. 1816: 198. 1816. 


Slender aquatic herbs, with linear entire verticillate leaves, and solitary (rarely 2-4) dis- 
coid peduncled terminal heads of small purplish flowers. Involucre broadly campanulate or 
hemispheric. Bracts imbricated in about 2 series. Receptacle conic, naked. Corolla regular, 
its tube short, its limb campanulate, 5-lobed. Anthers obtuse at the base. Style-branches 
slender, obtuse. Pappus of 5 broad cartilaginous obtuse scales. Achenes 5-angled. [Greek, 
hard scale, referring to the pappus. | 

A monotypic genus of eastern North America. 


1. Sclerolepis uniflora ( Walt.) Porter. 
Sclerolepis. Fig. 4151. 


Aethulia uniflora Walt. Fl. Car. 195. 1788. 
Sparganophorus verticillatus Michx. Fl. Bor. Am, 2: 98. 

1803. 

Eplerciebis verticillata Cass. Dict. 48: 155. 1827. 
meets uniflora Porter, Mem. Torr. Club 5: 311. 

1894. 

Perennial; stem simple, decumbent at the base, 
erect or ascending, glabrous or slightly pubescent, 
1°-2° long, leafy. - Leaves sessile, verticillate in 
4’s—6’s, linear, I-nerved, 4-12” long, 47-5” wwide, or 
the submerged ones filiform: head about 5” broad; 
bracts of the involucre linear-oblong, acutish, usually 
puberulent. 

In shallow ponds and streams, New Hampshire to 
Florida. July—Sept. 


4. EUPATORIUM [Tourn.] L. Sp. Pl. 
836. 1753. 


Erect, mostly branching, perennial herbs, with opposite or verticillate, or sometimes alter- 
nate, often punctate leaves, and in our species cymose-paniculate discoid heads of white, blue 
or purple flowers. Involucre oblong, ovoid, campanulate, or hemispheric, the bracts imbri- 
cated in 2-several series. Receptacle flat, convex, or conic, naked. Corolla regular, its tube 
slender, its limb 5-lobed or 5-toothed. , Anthers obtuse and entire at the base, appendiculate 
at the apex. Style-branches elongated, flattened, or thickened above, stigmatic at the base. 
Achenes 5-angled, truncate. Pappus of numerous capillary usually scabrous bristles arranged 
in I row. [Named for Mithridates Eupator, 1. e., of a noble father.] 

Over 500 species, mostly of warm or tropical regions. Besides the following, some 35 others 


occur in the southern and western parts of North America. Type species: Eupatorium cannd- 
binum L. 


* Leaves aes pinnatifid into filiform segments. 1. E. capillifolium. 
'** Leaves petioled, verticillate in 3’s—6’s, or the upper opposite; involucral pede in several series. 
Leaves thick, rugose, pubescent ; jdorescenee depressed. 

Leaves ovate, acute. E. maculatum. 
Leaves lanceolate to ovate-lanceolate, acuminate. E. Bruneri. 
Leaves thin, nearly glabrous; inflorescence pyramidal. 4. E. purpureum. 

*** Leaves opposite (rarely in 3’s), or the uppermost alternate. 
{ Involucral bracts imbricated in 2 or more series, the outer shorter. 
I. Leaves, at least the lower, slender-petioled. 5. E. serotinum. 
2. Leaves all sessile, short-petioled or connate-perfoliate. 
a. Leaves not clasping nor connate-perfoliate. 
§ Leaves narrowed at the base. 
Bracts of the falacee acute or cuspidate, scarious-tipped, white. 


wh 


Leaves linear-lanceolate, sparingly toothed, 2”—6” wide. | 6. E. leucolepis. 
Leaves oblong or lanceolate, coarsely toothed, 14’ wide. 7. E. album. 
Bracts of the involucre obtuse, not scarious, or but slightly so. 
Leaves linear, crowded, usually entire, obtuse. 8. E. hyssopifolium. 
Leaves lanceolate, oval, or oblong, usually dentate. 
Leaves linear-lanceolate, acute, dentate. . 9. E. Torreyanum. 
Leaves oblong to oval, sharply dentate, obtusish or acute. 10. E. semiserratum. 
Leaves lanceolate, sparingly dentate, long-acuminate. 11. E. altissimum. 
§§ Leaves rounded, obtuse or truncate at the base. 
Plant glabrous; leaves lanceolate, long-acuminate. © 12. E. sessilifolium. 
Plants pubescent; leaves ovate or oblong, acute or obtuse. 
Leaves ovate-oblong, rounded or narrowed at the base, usually obtuse. 13. E. verbenaefolium, 
Leaves broadly ovate, crenate-dentate, mostly truncate at the base, obtusish. 14. E. rotundifolium. 
Leaves ovate, dentate, acute. 15. E. pubescens. 
b. Leaves clasping or connate-perfoliate at the base. 
Leaves connate-perfoliate ; involucral bracts acute. 16. E. perfoliatum. 
Leaves merely clasping ; involucral bracts obtuse. 17. E.resinosum. 


+t Involucral bracts in 1 or 2 series, all equal or nearly so. 


356 COMPOSITAE. Vo. IIT. 


Receptacle flat. 
Flowers white ; leaves ovate. 


Leaves thin, 2-5’ long, sharply dentate, acuminate. 18. E. urticaefolium. 
Leaves firm, 1’/—2’ long, obtusely dentate, acute or obtusish. 19. E. aromaticum. 
Flowers pink to purple; leaves deltoid-ovate. 20. E. incarnatum. 
Receptacle conic; flowers blue or violet; leaves petioled. 21. E. coelestinum. 


1. Eupatorium capillifolium (Lam.) Small. 
Dog-fennel. Hog-weed. Fig. 4152. 


Artemisia capillifolia Lam. Encycl. 1: 267. 1783. 
Eupatorium foeniculcides Walt. Fl. Car. 199. 1788 
E. foeniculaceum Wild. Sp. Pl. 3: 1750. 1804. 

E. capillifolium Small, Mem. Torr. Club 5: 311. 1894. 


Erect, paniculately much branched, with the as- 
pect of an Artemisia, the stem finely pubescent, 
4°-10° high. Leaves crowded, glabrous or nearly 
so, alternate, pinnatifid into filiform segments, the 
lower petioled, the upper sessile; heads very nu- 
merous, about 13%” high, short-pedicelled, race- 
mose-paniculate, 3-6-flowered; bracts of the invo- 
lucre in about 2 series, linear, cuspidate, narrowly 
scarious-margined, glabrous; flowers greenish- 
white. 


In fields, Virginia to Florida. In ballast, at Phila- 
delphia. Also inthe West Indies. Sept. 


2. Eupatorium maculatum L. Spotted 
Joe-Pye Weed. Fig. 4153. 

E. maculatum L. Amoen. Acad. 4: 288. 1755. 

Eupatorium purpureum var. maculatum Darl. FI. 

Cest. 453. 1837. 

Eupatorium maculatum amoeénum Britton, Mem. 

Torr. Club 5: 312. 1894. 

Similar to the two following species, sca- 
brous or pubescent, often densely so, 2°-6 
high. Stem usually striate, often rough and 
spotted with purple; leaves thick, ovate or 
ovate-lanceolate, coarsely dentate, verticillate 
in 3’s—5’s, or the upper ones opposite; inflores- 
cence depressed, cymose-paniculate; pedicels 
and outer scales of the involucre pubescent; 
flowers pink_or purple. 

In moist soil, Newfoundland to New York, 
Kentucky, British Columbia, Kansas and New 
Mexico. Spotted boneset. Perhaps to be re- 
garded as a race of E. purpureum. Aug.-Sept. 


3. Eupatorium Brutneri A. Gray. Bru- 
ner’s Trumpet-weed. Fig. 4154. 


Eupatorium Bruneri A. Gray, Syn. Fl.1*7: 96. 1884. 
Eupatorium Rydbergi Britton, Manual 921. 1901. 


Stem tall, pubescent, often densely so, at 
least above. Leaves verticillate in 3’s—5’s, rather 
slender-petioled, lanceolate, serrate, acuminate 
at the apex, narrowed at the base, scabrous 
above, finely densely pubescent and reticulate- 
veined beneath, 4’-6’ long, #’-2’ wide; inflores- 
cence depressed or subpyramidal; outer bracts 
of the cylindric involucre pubescent; flowers 
pink or purple. 


In moist soil, South Dakota to Wyoming, Ne- 
braska and Colorado. Apparently erroneously 
recorded from Iowa. July—Sept. 


GENUS 4. THISTLE FAMILY. iw 4 


4. Eupatorium purpureum L. Joe-Pye or Trumpet-weed. Gravel-root. Tall 
or Purple Boneset. Fig. 4155. 
Eupatorium purpureum L., Sp. Pl. 838. 1753. 


E. falcatum Michx. Fl. Bor. Am. 2: 99. 1803. SOW VA iy 
Eupatorium purpureum var. angustifolium T. & G. FI. a th A YY 

N. A. 2: 82. 1841. EEG y 
Eupatorium purpureum falcatum Britton, Mem. Torr. = i Vi VARY 

@lubs5)) 382: euso04: i We F| | Aw 


SE 


i 


i) 
( 


Glabrous or sparingly pubescent, simple or 
branched at the summit, 3°-10° high. Stem green 
or purple, terete or striate, usually smooth; leaves 
thin, verticillate in 3’s—6’s, ovate, oval, ovate-lan- 
ceolate or narrowly lanceolate, petioled, acumi- 
nate, serrate, 4’-12’ long, 6’-3’ wide, glabrous or 
slightly pubescent along the veins on the lower 
surface; inflorescence usually elongated; heads 
very numerous; involucre cylindric, its bracts 
pink, oblong, obtuse, imbricated in 4 or 5 series, 
the outer shorter; flowers pink or purple, occa- 
sionally white. 

In moist soil, New Brunswick to Manitoba, Flor- 
ida and Texas. Kidney-root. Skunk-weed. Indian 
gravel-root. Marsh-milk weed. Nigger-weed. Quill- 
wort. Motherwort. King- or queen-of-the-meadow. 
Aug.—Sept. 

Eupatorium trifoliatum L. has the teeth of the 
leaves bluntly apiculate, but otherwise closely re- 
sembles E. purpureum and may not be specifically 
distinct. 


7 5. Eupatorium serétinum Michx. Late- 
flowering Thoroughwort. Fig. 4156. 


E. serotinum Michx. Fl. Bor. Am. 2: 100. 1803. 


Much branched, finely and densely pubescent, 
or glabrate below, 4°-8° high. Leaves all slen- 
der-petioled, lanceolate or ovate-lanceolate, acu- 
minate, sharply serrate, 3-6’ long, 2-2’ wide, 
5-nerved at the base, the lower opposite, the 
upper alternate; heads very numerous, the in- 
florescence broadly cymose; heads 7-15-flow- 
ered, 2-3” high; involucre campanulate, its 
bracts pubescent, linear-oblong, obtuse or trun- 
cate, imbricated in 2 or 3 series, the outer 
shorter; flowers white. 


N, 


Z 


6. Eupatorium leucélepis T.&G. White- 
bracted Thoroughwort. Justice-weed. 


Bigs 4157; 

E. leucolepis T.& G. Fl. N. A. 2: 84. 1841. 

Slender, puberulent, branched above, 1°-2 
high. Leaves opposite, sessile, linear, oblong 
or oblong-lanceolate, glaucous green, rough on 
both sides, thick, blunt-pointed, sparingly ser- 
rate, or the upper entire, 1-3’ long, 2’-5” wide, 
obscurely 3-nerved and narrowed at the base; 
inflorescence cymose-paniculate; heads 3’-4” 
high, about 5-flowered; bracts of the narrow 
involucre imbricated in about 3 series, white, 
lanceolate, acute, densely canescent, the outer 
shorter; flowers white. 

In moist places, Massachusetts and Long Island 
to Florida, Georgia and Louisiana. Aug.—Sept. 


° 


358 COMPOSITAE. Vo. III. 


SRY AE sa 
SVEN Yvale y 


i 
< - Ni: t) 
: A 5 8 Yy 


7. Eupatorium album L. White Thor- 
oughwort. Fig. 4158. 


Eupatorium album L. Mant. 111. 1767. 
Eupatorium glandulosum Michx. Fl. Bor. Am. 2: 
98. 1803. 
Eupatorium album subvenosum A. Gray, Syn. FI. 
: 1: Part 2,98. 1884. 


Pubescent with spreading hairs, branched 
above, 1°-3° high. Leaves opposite, sessile or 
nearly so, oblong or oblong-lanceolate, obtuse, 
obtusish or the upper acute at the apex, nar- 
rowed at the base, coarsely or rather finely ser- 
rate, 1-4’ long, 4-1’ wide, rather thick, mi- 
nutely scabrous above, more or less pubescent 
beneath ; inflorescence cymose-paniculate ; heads 
numerous, 4-5” long, 5-7-flowered; involucre 
narrow, its bracts bright white, linear, cuspi- 
date, imbricated in 3-4 series, the outer short 
and usually pubescent, the inner much longer, 
glabrous and shining; flowers white. 

In sandy soil, Long Island to Florida, Ten- 
nessee, Arkansas and Louisiana. Ascends to 3000 
ft. in Virginia. Aug.—Sept. 


8. Eupatorium hyssopifolium L. Hyssop- 
leaved Thoroughwort. Fig. 4159. 
Eupatorium hyssopifolium L. Sp. Pl. 836. 1753. 

E. linearifolium Walt. Fl. Car. 199. 1788. = 
Roughish-puberulent, densely corymbosely 
branched above, bushy, 1°-2° high. . Leaves linear, 
opposite, and fascicled in the axils of the stem, 
or on short axillary branches, entire or very 
nearly so, 2-2’ long, 1-2” wide, firm, obtuse at 
the apex, narrowed at the base; inflorescence 
densely cymose-paniculate; heads 3’-4” long, 
about 5-flowered; involucre campanulate, its bracts 
linear or linear-oblong, obtuse or truncate, some- 
time apiculate, puberulent, imbricated in about 3 

series, the outer shorter; flowers white. 

In dry fields, Massachusetts to Florida and Texas. 
Justice-weed. Leaf-margins usually revolute. Aug.— 
Sept. A plant from the coast of Maryland with 
very narrow leaves, closely approaches Eupatorium 
lecheaefolium Greene, from Florida. 


9g. Eupatorium Torreyanum Short. Tor- 
rey’s Thoroughwort. Fig. 4160. 


Eupatorium Torreyanum Short, 2nd Suppl. Cat. Pl. 
Ky. 5. 1836. 

Eupatorium hyssopifolium var. laciniatum A. Gray, 
Syn. Fl. 1: Part 2, 98. 1884. 


Rootstock tuberous-thickened; stem usually 
densely puberulent, mostly branched above, 13°- 
3° high. Leaves opposite, or sometimes in 3's, 
commonly with short leafy branches in their 
axils, often drooping, lanceolate, acute at the 
apex, narrowed at the base, sessile, 1-3’ long, 
2”-6” wide, usually distinctly 3-nerved, distantly 
serrate, the upper and those of the branches 
much smaller, linear, entire; inflorescence mostly 
loose; heads about 3” high, generally 5-flow- 
-ered; bracts of the campanulate involucre linear- 
oblong, obtuse, pubescent, or puberulent, imbri- 
cated in about 3 series, the outer shorter; flow- 
ers white. 

In dry soil, Pennsylvania to North Carolina, 
Kentucky, Florida and Texas. July—Sept. 


GENUS 4. THISTEE 


10. Eupatorium semiserratum DC. 
Small-flowered Thoroughwort. 
Fig. 4161. 


E, semiserratum DC. Prodr. 5: 177. 1836. 


Pubescent or puberulent, loosely branched 
above, 2°-3° high. Leaves rather thin, short- 
petioled, oblong-lanceolate to spatulate, acute 
or obtusish at the apex, narrowed at the base, 
sharply serrate, at least above the middle, 
2’-4’ long, 5-12” wide, 3-nerved, -usually 
with short branches in their axils; inflores- 
cence cymose-paniculate; heads 2’—3” high, 
about 5-flowered; involucre campanulate, 
its bracts linear-oblong, obtuse, canescent, 
imbricated in 2 or 3 series, the outer shorter, 
flowers white. 


In dry soil, Virginia to Florida, Missouri, 
Arkansas and Texas. Aug.—Sept. 


12. Eupatorium sessilifolium L. Up- 


land or Bastard Boneset. Fig. 4163. 
Eupatorium sessilifolium L. Sp. Pl. 837. 1753. 

Glabrous throughout, or pubescent above, 
branched near the summit, 2°-6° high. Leaves 
opposite, or the lower rarely in 3’s, closely ses- 
sile, lanceolate or ovate-lanceolate, long-acumi- 
nate, thin, sharply serrate, 3’-6’ long, 4’-12’ 
wide, inflorescence cymose-paniculate; heads 
4-5” high, about 5-flowered; involucre cam- 
panulate, its bracts linear-oblong, imbricated in 
about 3 series, the inner obtuse, the outer acut- 
ish, shorter; flowers white. 

In dry woods, Vermont to Massachusetts, Penn- 
sylvania, Georgia, Alabama and Missouri. Aug— 
Oct. A related plant, described under the name 
Eupatorium sessilifolium Brittonianum Porter, 
has leaves ovate or oblong-ovate, sparingly and 
finely serrate, firm, rounded at the base, acute, the 
upper 17-2’ long, 6”-9” wide, pinnately veined, 


dark green; cymes dense, their branches puberulent. 


NS \ 


FAMILY. 


Sl Wie, aD 
yds $ 
ay Vg Va» Vy y / 


VAISS 


x 


11. Eupatorium altissimum L. Tall 
Thoroughwort. Fig. 4162. 


— 


Eupatorium altissimum L. Sp. Pl. 837. 1753. 

Densely and finely pubescent, corymbosely 
much-branched above, 4°-8° high. Leaves 
lanceolate, acuminate at the apex, tapering 
below into a short petiole, roughish, rather 
thick, sparingly dentate above the middle, 
or some of them entire, strongly 3-ribbed, 
2’-5/ long, 5’—12” wide; inflorescence dense- 
ly cymose-paniculate; heads about 5-flow- 
ered, 3-4” high; involucre campanulate, 
its bracts oblong, obtuse or truncate, densely 
pubescent, imbricated in about 3 series, the 
outer shorter; flowers white. 


: In dry open places, Pennsylvania to North 
Carolina, Alabama, Illinois, Minnesota, Ne- 

braska and Texas. Sept.—Oct. 

Q RA 


GAOsh 
YW 


‘ 


It is known only from Budd’s Lake, N. J. 


360 COMPOSITAE. Vor 


13. Eupatorium verbenaefolium Michx. Rough or Vervain Thoroughwort. 
Fig. 4164. 


Eupatorium pilosum Walt. Fl. Car. 199. 1788. ? 
Eupatorium verbenaefolium Michx. Fl. Bor. 

Am. 2: 98. 1803. 

E. teucriifolium Willd. Sp. Pl. 3: 1753. 1804. 
E. verbenaefolium Saundersi Porter; Britton, 

Man. 923. 1901. 

Rough-pubescent, slender, 3°-8° high, 
branched at the summit. Leaves opposite, 
or occasionally in 3’s, ovate-oblong, usually 
obtuse or blunt-pointed, closely sessile or 
rarely short-petioled, rounded or narrowed 
at the base, crenate-dentate, or incised, 
2’-4’ long, 2’-1’ wide, the upper pairs dis- 
tant and small; inflorescence cymose- 
paniculate; heads 5-flowered, about 3” high; 
involucre campanulate, its bracts linear- 
lanceolate, acute, imbricated in about 3 
series, densely pubescent, the outer shorter; 
flowers white. 

In moist soil, Massachusetts to Pennsylva- 
nia, West Virginia, Florida and Louisiana, 
mostly near the coast. Called also wild hoar- 
hound. July—Sept. 

¥y WO 


wy Mp) WA Yj 
OAs 


SEW 7's 


14. Eupatorium rotundifélium L. 
Round-leaved Thoroughwort. 
Wild Hoarhound. Fig. 4165. 


E. rotundifolium L. Sp. Pl. 837. 1753. 


Pubescent, branched at the summit, 1°-3° 
high. Leaves opposite, sessile, broadly ovate, 
often as wide as long, acutish or obtuse, trun- 
cate to subcordate at the base, coarsely dentate- 
crenate, 1-2’ long, ascending; inflorescence 
cymose-paniculate; heads about 5-flowered, 
2”-3” high; involucre campanulate, its bracts 
linear-oblong, acutish, densely pubescent, im- 
bricated in about 3 series, the outer shorter; 
flowers white. 


In dry soil, Rhode Island to Pennsylvania, 
Florida, Kentucky, Arkansas and Texas. Erro- 
neously reported from Canada. July—Sept. 


15. Eupatorium pubéscens Muhl. Hairy 
Thoroughwort. Fig. 4166. 


E. pubescens Muhl.; Willd. Sp. Pl. 3: 1755. 1804. 


Eupatorium rotundifolium var. ovatum Torr.; DC. 
Prodr. 5: 178. 1836. 


Puberulent or pubescent, branched above, 
usually taller than the preceding speciés. Leaves 
opposite, ovate, sessile, often twice as long as 
wide, acute at the apex, rounded or subtruncate 
at the base, coarsely dentate with acute or acut- 
ish teeth, or the lower incised; inflorescence 
cymose-paniculate, sometimes 10’ broad; heads 
5-8-flowered, 2’-3” high; involucre campanu- 
late, its linear-oblong bracts in 2 or 3 series, 
the outer shorter; flowers white. 


In dry soil, Maine to Pennsylvania, South Caro- 
lina, West Virginia and Tennessee. Recorded 
from Florida. Ascends to 3000 ft. in Virginia. 
July—Sept. 


GENUS 4. 


16. Eupatorium perfoliatum L. 
Common Thoroughwort. Bone- 
set. Indian Sage. Fig. 4167. 

E. perfoliatum L. Sp. Pl. 838. 1753. 
Eupatorium truncatum Muhl.; Willd. Sp. 

ID eye ern ntsKeye 
Eupatorium perfoliatum truncatum A. Gray, 

Syn. Fl. 1: Part-2, 99. 1804. 

Pubescent, stout, branched above, 2°-5° 
high. Leaves opposite, or rarely in 3’s, 
connate-perfoliate, or the upper, rarely 
all, truncate and separated at the base, 
divaricate, lanceolate, long-acuminate 
with a slender apex, finely crenate-ser- 
rate, rugose and pubescent beneath, 4-8’ 
long, 1’-13’ wide; heads crowded, 10-16- 
flowered, 2-3” high; involucre cam- 
panulate, its bracts lanceolate, acutish, 
in 2 or 3 series, pubescent, the outer 
shorter; flowers white, rarely blue. 

In wet places, Nova Scotia and New 
Brunswick to Manitoba, Florida, Nebraska 
and Texas. Called also ague-weed, cross- 
wort, wild sage, thorough-wax, thorough- 
grow, thorough-stem. July—Sept. 


s 
Set 


ary) NT 
a NTT 4 
Ny And NW 
Wawa: 


Qs" 
\ ‘ 


/\s 


ZN 


E. urticaefolium Reichard, Syst. ait 


719. 
1780. 

E. ageratoides L. f. Suppl. 355.. 1781. 

E. boreale Greene, Rhodora 3: 83. 1901. 


Glabrous or villous and somewhat vis- 
cid, much branched, 1°-4° high. Leaves 
opposite, ovate, thin, acuminate at apex, 
rounded, truncate or cordate at the base, 
or abruptly narrowed into the slender 
petiole, coarsely and usually sharply den- 
tate-serrate, sometimes crenate, 3-6 
long, 1-3’ wide, 3-nerved and veiny; 
petioles 3-23’ long; inflorescence rather 
loose, ample; heads 10-30-flowered; re- 
ceptacle flat; involucre narrowly cam- 
panulate, about 2” high, its bracts linear, 
acute or acuminate, in I or 2 series, 
equal or nearly so; flowers bright white. 


In rich woods, New Brunswick to Flor- 
ida, Ontario, Nebraska and Louisiana. In- 
dian sanicle. Richweed. Stevia (Wis.). 
White sanicle. Deerwortboneset. July—Nov. 


THISTLE FAMILY. 


h WN Ny 
AOE, 
vy 
LZ 


Vv 
Vy 
‘s\ 
Wee 
oa = 
cS ane 
J FG aa 
y Z 


\ Nf Ma 
AN Ny ) 
Wie 


; vy, 


\ 


A plant described as Eupatorium perfoliatum 
cunedtum Engelm., with leaves smaller, narrowed 
and separated at the base, from Arkansas and Mis- 
souri,.is probably a hybrid with E£. serotinum. 


17. Eupatorium resindsum Torr. Resin 
Boneset. Fig. 4168. 
E. resinosum Torr. DC. Prodr. 5: 176. 


Slender, finely pubescent and resinous, 
high, branched at the summit, the inflorescence 
comparatively small, 3-4’ broad. Leaves op- 
posite, closely sessile, clasping, or slightly con- 
nate-perfoliate at the base, linear-lanceolate, 
long-acuminate, sharply serrate, 3-6’ long, 
3-6” wide, roughish above, canescent beneath; 
heads 10-15-flowered; involucre campanulate, 
about 2” high, its bracts oblong, obtuse, in 2 
or 3 series, the outer shorter; flowers white. 


In wet pine-barrens of New Jersey. Aug.—Sept. 


1836. 


2° 3° 


18. Eupatorium urticaefolium Reichard. 
White Snake-root. Fig. 4169. 


362 COMPOSITAE. Vor. III. 


y =| 19. Eupatorium aromaticum L. Smaller 


White Snake-root. Fig. 4170. 


Eupatorium aromaticum L. Sp. Pl. 8309. 1753) 


Puberulent or glabrate, slender, branched at 
the summit, 1°-2° high. Leaves opposite, peti- 
oled, firm, obtuse or acutish at the apex, round- 
ed, cordate or sometimes narrowed at the base, 
crenate-dentate, 12-3’ long, 9-18” wide, 3- 
nerved and veiny; petioles 2’-8” long; in- 
florescence usually compact; heads 10-25-flow- 
ered; receptacle flat; involucre campanulate, 
12-2” high, its bracts linear, generally obtuse, 
In I or 2 series; flowers white. 


In dry soil, Massachusetts to Florida, Pennsyl- 
vania, West Virginia, Tennessee and Mississippi. 
Blooms Tather later than the preceding species 
where the two grow together, near New York. 
Called also poolroot, poolwort, and wild hoar- 
hound. Aug.—Oct. 


Pink Thoroughwort. Fig. 4171. 
E. incarnatum Walt. Fl. Car. 200. 1788. 


Minutely pubescent, 2°-4° high, some- 
times reclining, often diffusely branched. 
Leaves opposite, deltoid-ovate, #}-2}’ 
long, long-pointed, rather coarsely blunt- 
serrate or crenate-serrate, truncate or 
cordate at the base, the petioles slender, 
often as long as the blades; heads about 
20-flowered; involucres campanulate, 
about 2” high, its bracts few, narrowly 
linear, slightly ribbed when dry, acute; 
corolla pink or pale purple. 

In woods and sandy soil, Virginia to Mis- 
souri, Florida and Mexico. Aug.—Oct. 

SY eH BAY 
y ‘ 


‘ (NV 9 Ve at ral y 
Git ig 5 ‘ SOc 
) S AWA, 
) 


WA ia LP SigiTAe the Vy, 
— e A ae 
WS Vo Wik By La 

Y 


is if 
WRU aa 
WY 


21. Eupatorium coelestinum L. Mist- 
flower. Blue Boneset. Fig. 4172. 


Eupatorium coelestinum L. Sp. Pl. 838. 1753. 
Conoclinium coelestinum DC. Prodr.5: 135. 1836. 


Pubescent or puberulent, branched, 1°-3° 
high. Leaves opposite, petioled, ovate, obtuse 
or acute at the apex, truncate at the base, or 
abruptly contracted into the petiole, crenate- 
dentate, 13’-3’ long, 9-18” wide; inflorescence 
cymose-corymbose, rather compact; heads 2’— 
3” high; involucre broadly campanulate, its 
bracts linear+lanceolate, acuminate, equal or 
nearly so, in I or 2 series; receptacle conic; 
flowers blue or violet. 

In moist soil, New Jersey to Florida, Illinois, 


Michigan, Kansas, Arkansas and Texas. Also in 
Cuba. Aug.—Oct. 


5. MIKANIA Willd. Sp. Pl. 3: 1742. 1804. 


[WiLtLuGHBAEA Neck. Elem. 1: 82. Hyponym. 1790.] 
Herbaceous twining vines, or some tropical species erect shrubs, with opposite, petioled 
leaves, and discoid, mostly cymose-paniculate heads of whitish flowers. Heads 4-flowered. 
Involucre oblong, of 4 slightly unequal narrow bracts. Receptacle small, naked. Corolla 


GENUS 5. THISTLE FAMILY. 363 


regular, its tube slender, the limb campanulate, 5-cleft. Anthers entire and obtuse at the 

base. Style-branches elongated, acutish. Achenes truncate, 5-angled. Pappus of numerous 

capillary bristles. [In honor of Joseph Gottfried Mikan, 1743-1814, professor at Prague.] 
About 150 species, natives of America. me 

Besides the following, two others occur in . 


the southern United States. Type species: 
Mikania hastata (L.) Willd. 


1. Mikania scandens (L.) Willd. 
Climbing Hempweed or Boneset. 


Fig. 4173. 
Eupatorium scandens L. Sp. Pl. 836. 1753. 
M. scandens Willd. Sp. Pl. 3: 1743. 1804. 
Willoughbya scandens Kuntze, Rev. Gen. 

PISs71. 1891. 

Glabrous or nearly so, twining over 
bushes, 5°-15° long. Leaves ovate or 
hastate, deeply cordate at the base with © 
the lobes rounded or truncate, acumi- 
nate at the apex, repand or obtusely 
dentate, 2-4’ long, 1-2’ wide; petioles 
slender, shorter than the blades; heads 
in compound clusters borne at the ends 
of the branches; involucre about 3” 
long, its bracts acuminate or apiculate; 
flowers white or pink; achenes resinous. 

In swamps and moist soil, Maine to west- 
ern Ontario, Florida and Texas. West 
Indies and South America. July—Sept. 


6. COLEOSANTHUS Cass. Dict. 10: 36. 1817. 
[BricKELLIA Ell. Bot. S. C. & Ga. 2: 290. 1824.] 

Herbs or shrubs, with opposite or alternate leaves, and discoid heads of white yellowish 
or pink flowers, in panicles or cymes, or rarely solitary. Involucre campanulate or oblong, 
its bracts striate, imbricated in several series, the exterior ones successively smaller. Recep- 
tacle flat or convex, naked. Corolla regular. 5-lobed. Anthers obtuse at the base. Style- 
branches long, obtuse. Achenes Io-striate or 10-ribbed. Pappus a single row of numerous 
rough or serrate bristles. [Greek, sheath-flower. | 


A genus of about 70 species. Besides the following, some 45 others occur in the southern and 
western United States. Type species: Coleosanthus Cavanillésii Cass. 


1. Coleosanthus grandiflorus (Hook.) Kuntze. Large-flowered Thoroughwort. 


EN » SQ WHEN) | EE Ea ge Hook. Fl. Bor. . 
SSS a y > UZAY <== Bache Granda Nutt. Trans. Am. 
Wi SS AY Gate dean tle geintiiorns Gdns Rey. 
} A, Co ee Gen Pittonia 4: 238. 


19Ol. 


Erect, glabrous or puberulent, much 
branched, 2°-3° high. Leaves slender- 
petioled, deltoid-ovate, cordate at the 
base, acuminate at the apex, coarsely 
crenate-dentate, 2-4’ long, 1-2’ wide; 
petioles shorter than the leaves; in- 
florescence cymose-paniculate; heads 
short-peduncled, 6-7” long; pedun- 
cles pubescent; involucre campanu- 
late, 30-45-flowered, the outer bracts 
ovate, pubescent, usually abruptly 
acuminate or subulate-tipped, the in- 
ner linear, glabrous, striate, obtuse 
or acute; pappus bristles scabrous. 


Montana to Wyoming, Missouri, New 
Mexico, Washington and Arizona. Aug.— 
Sept. Tassel-flower. 


304 COMPOSITAE. VoL. III. 


7. KUHNIA L. Sp. Pl. Ed. 2, 1662. 1763. 


Perennial herbs, with alternate punctate resinous-dotted leaves, and discoid heads of 
white or purplish flowers in terminal cymose corymbs. Involucre turbinate-campanulate, 
its bracts striate, imbricated in several series, the outer shorter. Corolla regular, the tube 
slender, the limb 5-lobed. Anthers obtuse and entire at the base, nearly or quite separate. 
Style-branches slender, obtusish. Achenes Io-striate. Pappus a single row of numerous 
gery plumose bristles. [Named for Dr. Adam Kuhn, of Philadelphia, a pupil of Linnaeus.] 

About 8 species, native of North America and Mexico. Type species: Kuhnia eupatorioides L. 
Puberulent ; leaves sparingly dentate, or entire; heads loosely clustered, 4”-5” high. 


1. K. eupatorioides. 
Pubescent or tomentulose; leaves sharply serrate; heads densely clustered, 6”—-8” high. 


2. K. glutinosa. 


1. Kuhnia eupatorioides L. False 
Boneset. Fig. 4175. 


Kuhnia eupatorioides L. Sp. Pl. Ed. 2, 1662. 1763. 


Eupatorium alternifolium Ard. Spec. Bot. 2: 40. 
pl. 20. 1764. 
Erect, puberulent and resinous, 1°-3° high, 
branched above. Leaves lanceolate or linear- 
lanceolate, acute or obtusish at the apex, nar- 


i H i cled, 4-5” high, loosely clustered; outer bracts 
jy of the involucre lanceolate or ovate-lanceo- 
| late, acuminate, the inner much longer, linear, 
| cuspidate; pappus tawny, or sometimes nearly 
| white. 

In dry soil, New Jersey to Georgia, Ohio, Min- 


nesota, West Virginia and Texas. Ascends to 
3300 ft. in West Virginia. Aug.—Sept. 


Kuhnia glutinosa Ell. Bot.S. C. & Ga. 2: 292. 1821-24. BY i \\ ty \ 
Kuhnia suaveolens Fresen. Ind. Sem. Francf. 1838. i\ Why Wi V4 WY ee 
Kuhnia eupatorioides var. corymbulosa T. & G. Fl. N. it . Wek We VF fits 
A207 Si) WSATe A rN z “ALL 
Ny 


y 
Wy ZZ 
A yay! EZZ. Z 


Stouter and often taller than the preceding spe- WN / 
cies, corymbosely or paniculately branched, pubes- ur A 1/ 
cent or tomentulose, somewhat viscid. Leaves all ff NW 
sessile, lanceolate to ovate-lanceolate, usually §& \ 
sharply serrate with distinct teeth, veiny, 1-3’ 
long, 3-10” wide, or those of the branches linear- 
lanceolate and entire; heads numerous, 6-8” 
high, densely clustered in the cymes, their pedun- 
cles mostly short; inner bracts of the involucre 
lanceolate, acuminate; pappus tawny or brown. 

In dry soil, Illinois to North Dakota, Colorado, 


Alabama and Texas. Perhaps a race of the preceding 
species. Aug.—Oct. 


Kuhnia Hitchcéckii A. Nelson, a little known 
species of Kansas, differs by having very small linear 
leaves, at least on the upper part of the plant. 


8. LACINARIA Hill, Veg. Syst. 4: 49. pl. 40. 1762. 
[Liatris Schreb. Gen. Pl. 542. 1791.] 

Erect perennial herbs, usually from a globular tuber, simple or little branched, with alter- 
nate, entire, narrow I-5-nerved leaves, and spicate or racemose discoid heads of rose-purple 
or white flowers. Involucre oblong, ovoid or subhemispheric, its bracts imbricated in several 
series, the outer shorter. Receptacle flat, or slightly convex, naked. Corolla regular, its 


GENus 8. THISTLE FAMILY. 365 


tube slender, its limb 5-lobed or 5-cleft. Amnthers obtuse at the base. Style-branches elon- 
gated, obtuse or flattened at the apex. Achenes Io-ribbed, slender, tapering to the base. 
Pappus of 1 or 2 series of slender barbellate or plumose bristles. [Latin, fringed, from the 
appearance of the heads.] 

About 35 species, natives of eastern and central North America, known as Blazing Star, or 
Button Snakeroot from the globular tubers. Type species: Serratula squarrosa L. 

* Bracts of the involucre acute, acuminate or mucronate. 

Involucre cylindric, or turbinate, 15—60-flowered, its base rounded. 


Bracts with lanceolate spreading rigid tips. 1. L. squarrosa. 

Bracts mucronate, closely appressed. 2. L. cylindrica. 
Involucre oblong, or narrowly campanulate, 3—6-flowered. 

Inner bracts with prolonged petaloid tips. 3. L. elegans. 


Bracts all acute, mucronate or acuminate. 
Bracts appressed ; pappus-bristles very plumose. 


Leaves 1”—2” wide; spike usually leafy below. 4. L. punctata, 
Leaves less than 1” wide; spike mostly naked. 5. L. acidota. 
Tips of the bracts spreading; pappus-bristles barbellate. 6. L. pycnostachya. 
** Bracts of the involucre rounded, obtuse or acutish. 
Involucre hemispheric, %’—1’ broad, 15—45-flowered ; heads peduncled. 7. L. scariosa. 
Involucre oblong, 2”—4” broad, 5—15-flowered. 
Bracts obtuse, rounded. 
Involucre rounded at base ; bracts usually not punctate ; heads mostly sessile. 
. L. spicata. 


Involucre narrowed at base; bracts usually punctate; heads peduncled. 
Leaves, even the lower, narrowly linear; involucre narrowly obovoid, 5”—6” high. 
g. L. graminifolia. 
Lower leaves linear-oblong; involucre broadly obovoid, 6”—7” high. 1o. L. pilosa. 
Bracts acutish, punctate. 11. L. Smallit. 


1. Lacinaria squarrosa (L.) Hill. Scaly 
Blazing Star. Colic-root. Fig. 4177. 


Serratula squarrosa L. Sp. Pl. 818. 1753. 

Lacinaria squarrosa Hill, Hort. Kew. 70. 1769. 

Liatris squarrosa Willd. Sp. Pl. 3: 1634. 1804. 

Liatris intermedia Lindl. Bot. Reg. pl. 948. 1825. 
Lacinaria squarrosa intermedia Porter, Mem. Torr. Club 

5: 314. 18094. 
pants squarrosa var. intermedia DC. Prodr. 5: 129. 

1836. 

Usually stout, 4°-2° high, pubescent or glabrous. 
Leaves narrowly linear, rigid, sparingly punctate, 
3-6’ long, 1’’-23”" wide; heads sessile or short-pedun- 
cled, 15-60-flowered, usually few, or sometimes soli- 
tary. 3’-13’ long, 4”-8” thick; bracts of the involucre 
imbricated in 5-7 series, lanceolate, rigid, acuminate, 
glabrous or pubescent, their tips more or less spread- 
ing when old; flowers purple; pappus very plumose. 

In dry soil, western Ontario to Pennsylvania, Virginia, 
Florida, South Dakota, Nebraska and Texas. Called also 
rattlesnake-master. Races differ in pubescence and in 


WY 


\) 


2. Lacinaria cylindracea (Michx.) 
Kuntze. Cylindric Blazing Star. 
Fig. 4178. 


Liatris cylindracea Michx. Fl. Bor. Am. 2: 93. 
1803. 

Liatris graminifolia Willd. Sp. Pl. 3: 1636. 1804. 

Lacinaria cylindracea Kuntze, Rev. Gen. Pl. 340. 
I8o1. 


Glabrous or nearly so, stout, 1°-13° high, 
sometimes branched above. Leaves narrowly 
linear, rigid, scarcely punctate, 3-7’ long, 1-2” 
wide; heads several or numerous (rarely soli- 
tary), peduncled, or the lower sessile, turbi- 
nate-cylindric, 4-1’ high, 4-6” thick, 15-60- 
flowered; bracts of the involucre imbricated in 
5 or 6 series, broadly oval, appressed, abruptly 
acuminate at the apex; flowers purple; pappus 
very plumose. 


In dry soil, western Ontario to Minnesota, south 
to Illinois and Missouri. July—Sept. 


366 COMPOSITAE. Vou. Te 


3. Lacinaria élegans (Walt.) Kuntze. 
Handsome Blazing Star. Fig. 4179. 


Stoepelina elegans Walt. Fl. Car. 202. 1788. 
Liatris elegans Willd. Sp. Pl. 3: 1635. 1804. 
Lacinaria elegans Kuntze, Rev. Gen. Pl. 349. 1891. 


Densely and finely pubescent, rarely glabrate, 
2°-3° high. Leaves linear, very punctate, 1-5’ 
long, 1’-3” wide, the upper much smaller than 
the lower and soon reflexed; heads 4-5-flowered, 
6-7” high, narrowly campanulate, very numerous 
in a dense spike or raceme sometimes a foot long; 
bracts of the involucre in 2 or 3 series, the inner 
ones linear, dilated above into oblong or lanceo- 
late acuminate rose-colored petaloid tips, or some- 
times white; pappus very plumose; flowers purple. 


In dry soil, Virginia to Florida, Alabama, Arkan- 
sas and Texas. Aug.—Oct. 


4. Lacinaria punctata (Hook.) Kuntze. 
Dotted Button-Snakeroot. Fig. 4180. 


Liatris punctata Hook. Fl. Bor. Am. 1: 306. pl. 55. 1833. 
Lacinaria punctata Kuntze, Rev. Gen. Pl. 349. r8o1. 


Glabrous, or sparingly pubescent, 6-30’ high; root- 
stock stout, branching, or globose. Leaves linear, 
rigid, very punctate, 2-6’ long, about 1” wide, or 
the lower 2”, the upper gradually shorter, all erect 
or ascending ; heads 3-6-flowered, 6’’-8” long, sessile, 
crowded into a dense spike; spike commonly leafy 
below; involucre narrowly campanulate, acute or 
acutish at the base, its bracts oblong, cuspidate or 
acuminate, often ciliate on the margins, sometimes 
woolly, imbricated in 4 or 5 series; flowers purple; 
pappus very plumose. 

In dry soil, Minnesota to Manitoba, Saskatchewan, 


Montana, Missouri, Texas, New Mexico and Sonora. 
Recorded from Ohio. Aug.—Oct. 


5. Lacinaria acidéta (Engelm. & Gray) Kuntze. 
Slender Button-Snakeroot. Fig. 4181. 


Liatris acidota Engelm. & Gray, Bost. Journ. Nat. Hist. 5: 
218. 1847. 


Lacinaria acidota Kuntze, Rev. Gen. Pl. 349. 1891. 


Similar to the preceding species, but usually glabrous 
throughout, the rootstock globose, or elongating. Stem 
2° high, or less, slender. Leaves narrowly linear, 3-3” 
wide, or the lower wider, slightly punctate, 1-4’ long, 
the upper gradually shorter; spike slender, naked, or 
sparingly leafy at the base, 4’-10’ long; heads 3-5-flow- 
ered, 6-9” long; involucre narrowly oblong-campanulate, 
its bracts oblong to ovate-lanceolate, more or less punc- 
tate, abruptly or gradually acuminate; glabrous or 
sparingly ciliate; flowers purple; pappus plumose. 


Prairies, Kansas to Arkansas and Texas. Aug.—Oct. 


Genus 8. THISTLE FAMILY. 


6. Lacinaria pyenostachya (Michx.) Kuntze. 
‘Prairie or Hairy Button-Snakeroot. 
Fig. 4182. 

Liatris pycnostachya Michx. Fl. Bor. Am. 2: 91. 1803. 
Lacinaria pycnostachya Kuntze, Rev. Gen. Pl. 349. 1891. 

Hirsute-pubescent above, usually glabrous below, 2°-5° 
high, very leafy. Lower leaves linear-lanceolate, nar- 
rowed below the middle into a slender margined petiole, 
acuminate at the apex, often 1° long and 2’ wide, the 
upper much smaller, linear-subulate, rigid, punctate; 
spikes very dense, 6-18’ long; heads 4-6” long, 3- 
6-flowered; involucre oblong or cylindric, its bracts 
oblong, pubescent and ciliate with acute, spreading, 
colored tips; flowers purple; pappus barbellate or 
roughened, scarcely plumose. 

On prairies, Indiana to Minnesota, Nebraska, Kentucky, 
Louisiana and Texas. Aug.—Sept. 


Zn Lez 
SS 


z- 


a a 


PAN 
Orme 


Al 


We 


Va! 


ALES 
= 


7. Lacinaria scaridsa (L.) Hill. Large. 
Button-Snakeroot. Fig. 4183. 


Serratula scariosa L. Sp. Pl. 818. 1753. 
Lacinaria scaricosa Hill, Hort. Kew. 70. 1769. 
Liatris scariosa Willd. Sp. Pl. 3: 1635. 1804. 
Liatris squarrulosa Michx. Fl. Bor. Am.2: 92. 1803. 


Finely pubescent, at least above, 1°-6° high. 
Lower leaves oblanceolate, spatulate or oblorg- 
lanceolate, narrowed into a margined petiole, 
acute or obtusish at the apex, often 1° long 
and 13’ wide; upper leaves linear or linear- 
lanceolate, acute, or sometimes very obtuse, 
much smaller, all densely punctate; heads hemi- 
spheric, 2-1’ broad, 15-45-flowered, on stout 
peduncles 2”-2’ long, or sometimes sessile; 
bracts of the involucre imbricated in 5 or 6 
series, spatulate-linear, oblanceolate or obovate, 
rounded at the apex, appressed, their tips dry 
and scarious, often colored; flowers bluish 
purple; pappus barbellate. 


In dry soil, Maine to Florida, western Ontario, 
Manitoba, Nebraska and Texas. Consists of many 
races, differing in leaf-form and size of hedds. 
Called also blue blazing-star, gray feather, rattle- 
snake-master, devil’s-bite. Aug.—Sept. 


8. Lacinaria spicata (L.) Kuntze. 
Dense Button-Snakeroot. Gay 
Feather. Devil’s Bit. Fig. 4184. 


Serrulata spicata L. Sp. Pl. 819. 1753. 

Liatris spicata Willd. Sp. Pl. 3: 1636. 1804. 
?Liatris pumila Lodd. Bot. Cab. pl. 147. 1821. 
L. spicata Kuntze, Rev. Gen. Pl. 349. 1891. 


Glabrous or nearly so, 2°-6° high. Lower 
leaves linear-lanceolate or linear-oblong, 
usually blunt-pointed, sometimes 1° long 
and 5” wide, the upper linear or even subu- 
late, somewhat or obscurely punctate; spike 
generally dense, 4’-15’ long; heads short- 
oblong or cylindric, 5-13-flowered, 2’-4” 
broad, mostly sessile; involucre subcam- 
panulate, rounded or obtuse at the base, 
its bracts appressed, oblong, obtuse and 
scarious-margined at the apex, obscurely 
punctate, imbricated in 4-6*series; flowers 
blue-purple, occasionally white; pappus 
roughened or barbellate. 

In moist soil, Massachusetts to Florida, On- 
tario, Wisconsin, Kentucky, Louisiana and Ari- 
zona. Called also rough- or backache-root, 
throat-wort, prairie-pine, colic-root. Aug.—Oct. 3 

Lacinaria kansana Britton, of Kansas, differs in having a short, very dense spike, a more leafy 
stem, the involucral bracts purple and the pappus plumose. 


368 COMPOSITAE. Vou. III. 


g. Lacinaria graminifolia ( Walt.) 
Kuntze. Loose-flowered Button- 
Snakeroot. Fig. 4185. 


Anonymos graminifolius Walt. Fl. Car. 197. 
1788. 

Liatris graminifolia Pursh, Fl. Am. Sept. 
2: 508. 1814. Not Willd. 1804. 

L. graminif. Kuntze, Rev. Gen. Pl.349. 1891. 

Liatris graminifolia var. dubia A.. Gray, 
Man. Ed. 2, 185. 1856. 


Glabrous, or sparingly pubescent, 1°-3° 


Ay, high. Leaves similar to those of the pre- 
ip a ceding species, but more conspicuously 
WAGE ae 
ge punctate, usually somewhat ciliate, at 
WZ Wy, least near the base and acute or acutish 
WZ 


at the apex; heads spicate, racemose or 
sometimes panicled, mostly peduncled; 
involucre obovoid, narrowed or acute at 
the base, 2’-4” broad, its bracts dis- 
tinctly punctate, appressed, rounded, 
more or less scarious-margined, thick; 
flowers purple; pappus _barbellate; 
achenes hairy. 

In dry soil, New Jersey to Florida. Fine- 
leaved blazing-star. Aug.—Sept. 


to. Lacinaria pilosa ( Ait.) Heller. Moun- 
tain Button-Snakeroot. Fig. 4186. 


Serratula pilosa Ait. Hort. Kew. 3: 138. 1780. 
L. pilosa Heller, Muhlenbergia 1: 6. 1900. 


Glabrous, except the ciliate leaf-bases and 
involucral bracts, 5’/-3%° tall, simple. Leaves 
rather numerous, the basal often fully half as 
long as the stem, with broadly linear or linear- 
oblong blades, the upper stem leaves much 
smaller, with linear blades, all more or less 
ciliate near the base; involucres turbinate, 23”—3” 
broad, sessile or short-peduncled, the bracts 
purple, the outer ovate or oval, the middle ones 
with oval or ovate blades and short claws, the 
inner linear or linear-spatulate, all obtuse and 
rather broadly margined; pappus_ barbellate; 
achenes 23-3” long, hairy. 


In dry or rocky soil, Virginia and West Virginia 
to Georgia and Alabama. July—Sept. 


Ii. Lacinaria Smallii Britton. Small’s 
Button-Snakeroot. Fig. 4187. 


Lacinaria Smallii Britton, Manual 927. 1901. 


Similar to L. graminifolia. Stem very slender, 
1°-2° high. Leaves scattered, linear, 2”—5” 
wide, acute, pubescent on the midvein beneath, 
spreading, very punctate; heads scattered, ses- 
sile in the axils of the upper small leaves, the 
uppermost leaves not larger than the involucral 
bracts; involucre oblong-campanulate, about 
8-flowered, about 5” high, its innermost bracts 
linear, acute or acutish, the outermost obtusish, 
all punctate; flowers purple; pappus plumose. 


Iron Mountain, Smyth Co., Virginia. Aug. 


Lacinaria Hélleri (Porter) Heller, of the North 
Carolina mountains, with narrower involucral 
bracts and erect or ascending leaves, is recorded 
from Virginia. 


GENUS 9. - “FTHISTEE, FAMILY. 369 


9. TRILISA Cass. Dict. Sci. Nat. 552310, 1828; 


Erect perennial herbs, with fibrous roots and alternate simple leaves; those of the stem 
sessile or clasping, the basal ones narrowed into petioles. Heads small, discoid, of 5-10 
purple, or white flowers, in terminal thyrsoid or corymbose panicles. Involucre campanulate, 
its bracts imbricated in 2 or 3 series, the outer scarcely shorter than the inner. Receptacle 
flat, naked. Corolla regular, its limb 5-lobed. Anthers obtuse and entire at the base. Achenes 
nearly terete, 10-ribbed. Style-branches slender, obtuse. Pappus of I or 2 series of elon- 
gated, barbed bristles. [Anagram of Luatris.] 


Two knows species, natives of the southeastern United States. Type species: Trilisia odora- 
tissima (Walt.) Cass. 


Stem glabrous ; heads corymbose-paniculate. 1, 7. odoratissima. 
Stem viscid-pubescent ; heads thyrsoid-paniculate. 2. T. paniculata. 


1. Trilisa odoratissima ( Walt.) Cass. 
Vanilla-leaf. Vanilla-plant. Dog’s- 
tongue. Carolina Vanilla. 

Fig. 4188. 
eres odoratissima Walt. Fl. Car. 198. 
Poa odoratissina Michx. Fl. Bor. Am. 2: 

93. 1803. \ 
Trilisa odoratissima Cass. Dict. Sci. Nat. 55: 

310. 1828. 

Rather stout, glabrous, 2°-3° high. Leaves 
thick, entire, or sometimes dentate, the 
lower oblanceolate, oblong or spatulate, 
obtuse, 4-10’ long, 1-13’ wide, those of the 
stem gradually smaller, oblong, ovate or 
oval, the uppermost bract-like; heads 
corymbose-paniculate, about 3” high; bracts 
of the involucre oblong, obtusish; achenes 
glandular-pubescent. 


In pine-barrens, North Carolina to Florida 
and Louisiana. Recorded from Virginia, and 
to be looked for in the southeastern part of that 
state. Deer’s- or hound’s-togue. Aug.—Sept. 


yyy 2. Trilisa paniculata ( Walt.) Cass. 
LAR Hairy Trilisa. Fig. 4189. 


\ (i W x Mig 
NYY \p Nit, Anonymos paniculatus Walt. Fl. Car. 198. 1788. 
Saul Uy V pe Liatris paniculata Michx. Fl. Bor. Am, 2: 93. 


x iN ft 1803. ¢ i 
x hye Trilisa paniculata Cass. Dict. Sci. Nat. 55: 310. 
= U2 
t 1828. 


Stem viscid-pubescent, 1°-3° high. Leaves 
\/ entire, the basal ones lanceolate or narrowly 
oblong, acute or obtusish, 3-10’ long, 3’—13’ 
wide, those of the stem much smaller, lan- 
ceolate or oblong-lanceolate ; heads thyrsoid- 
paniculate, about 3” high; bracts of the 
involucre oblong, obtusish; achenes finely 
pubescent. 


In pine-barrens, Virginia to Florida. Sept— 
Oct. 


10. GUTIERREZIA Lag. Gen. & 
Sp. Nov. 30. 1816. 


Erect branching, usually glabrous and often glutinous herbs, or shrubs, with linear entire 
alternate leaves, and small heads of both tubular and radiate yellow flowers, corymbosely 
paniculate in our species. Radiate flowers few, pistillate. Disk-flowers regular, mostly 


24 


379 COMPOSITAE. : Vor. III. 


perfect, or some of them only staminate, the corolla-limb 5-lobed. Involucre ovoid, or 
narrowly campanulate, its bracts coriaceous, imbricated in few series. Receptacle flat, convex 
or conic, commonly foveolate. Anthers obtuse and entire at the base. Style-branches narrow, 
flattened, their appendages slender. Achenes terete, ribbed or 5-angled. Pappus of several 
scales, those of the ray-flowers shorter than or equalling those of the disk. [Named from 
Gutierrez, a noble Spanish family.] ' 

About 25 species, natives of western North America, Mexico and western South America. 


Besides the following, several others occur in the western United States. Type species: Gutierrezia 
linearifolia Lag. 


1. Gutierrezia Sarothrae (Pursh) Britton and Rusby. Broom-weed. Fig. 4190. 
Sora 4 4 7%: 
py SY Solidago Sarothrae Pursh, Fl. Am. Sept. 540. 1814. 
: Viog 2 Gutierrezia Euthamiae T.&G. Fl. N. A. 2: 193. 1841. 
Gutierrezia Sarothrae Britton & Rusby, Trans. N. Y. 
Acad. Sci. 7: 10. 1887. 


Glabrous or minutely pubescent, bushy, woody 
at the base, 4°-14° high, the branches rigid, as- 
cending. Leaves linear, 2-13’ long, 4’—-1” wide, 
acute; heads oblong, 13”-2” high, few-flowered, 
usually in clusters of 2-5 at the ends of the 
numerous branches; rays I-6, scarcely 1” long; 
scales of the pappus linear-oblong, subulate or 


acuminate; achenes pubescent. 


In dry or rocky soil, Manitoba and Minnesota to 
western Nebraska, Kansas, Texas and Sonora, west to 
Alberta and. Utah. Adventive at Rochester, N. Y. 
Far western plants formerly referred to this species 
prove to be distinct. Rabbit-brush. Aug.—Sept. 


Gutierrezia linearis Rydb., of the Rocky Mountain 
region, ranging eastward into Kansas and Nebraska, 
differs in being more woody and has somewhat wider 
leaves. 


11. AMPHIACHYRIS [DC.] Nutt. Trans. Am. Phil. Soc. IT. 72 313emee 


Erect, much branched, glabrous herbs, with small alternate entire leaves, and very 
numerous small heads of both tubular and radiate-yellow flowers, solitary or clustered at 
the ends of the branches. Involucre ovoid or hemispheric, its bracts coriaceous, imbricated 
in few series. Receptacle naked. Ray-flowers pistillate. Disk-flowers perfect, but sterile, 
or staminate. Pappus of the ray-flowers obsolete or coroniform, that of the disk-flowers of 
5-20 subulate scales or bristles somewhat united at the base. Achenes pubescent. [Greek, 
chaff-around. | 


Two known species, natives of the south-central United States, the following typical. 


1. Amphiachyris dracunculoides (DC.) 
Nutt. Amphiachyris. Fig. 4191. 
Brachyris dracunculoides DC. Mem. Soc. Phys. Gen. 

7 bart 2e2O5 spit Toso. 

Amphiachyris dracunculoides Nutt. Trans. Am. Phil. 

Soc. (LT) gears exSar- 

Annual, slender, much branched, 6’-18’ high, the 
branches ascending. Leaves linear, 6-18” long, 
1-2” wide, acutish, the uppermost almost fili- 
form; heads solitary at the ends of short branches, 
10-30-flowered, about 2” high; involucre hemi- 
spheric, its bracts oval, obtuse; rays 5-10, about 
as long as the involucre; disk-flowers mostly 
staminate, their ovaries abortive, their pappus of 
5-8 subulate aristate scales, united into a short 
cup at the base. 

In dry soil, Missouri and Kansas to Texas and 


New Mexico. Found adventive at Easton, Penna. 
Sept.—Oct. 


GENUS 12. THISTLE FAMILY. aa 


12. GRINDELIA Willd. Gesell. Nat. Fr. Berl. Mag. 1: 260. 1807. 


Perennial or biennial herbs, sometimes woody at the base, with alternate sessile or clasp- 
ing leaves, usually spinulose-dentate, and rather large heads of both discoid and radiate 
yellow flowers, solitary at the ends of the branches (rays rarely wanting). Involucre hemi- 
spheric or depressed, its bracts imbricated in several or many series, usually subulate-tipped. 
Receptacle flat or convex, naked, foveolate. “Ray-flowers fertile. Disk-flowers perfect, or 
sometimes only staminate. Anthers obtuse and entire at the base. Style-branches narrow, 
flattened, their appendages linear or lanceolate. Achenes short, thick, sometimes compressed, 
glabrous, 4-5-ribbed. Pappus of 2-8 soon deciduous awns or bristles. [Named for Prof. H. 
Grindel, of Riga, 1776-1836. | 

About 30 species, natives of western North America, Peru and Chile. Besides the following, 


some 15 others occur in the western and southwestern parts of North America. Known as Gum- 
plant or Tar-weed. Type species: Grindelia inuloides Willd. 


Leaves spatulate or oblong, obtuse or obtusish; achenes truncate; bracts squarrose. 
: ; 1. G. squarrosa. 
Leaves linear or linear-oblong, acute; achenes 1-2-toothed; bracts not squarrose. 2. G. lanceolata. 


1. Grindelia squarrésa (Pursh) Dunal. Broad-leaved Gum-plant. Fig. 4192. 
Donia squarrosa Pursh, Fl. Am. Sept.559. 1814. Wi 


Grindelia squarrosa Dunal in DC. Prodr. 5: 
BEG S30. 

G. grandiflora Hook. Bot. Mag. pl. 4628. 1852. 

Grindelia nuda Wood. Bot. Gaz. 3: 50. 1878. 

Grindelia squarrosa nuda A. Gray, Syn. FI. 1°: 
118. 1884. 


AY LE 


SS 


Glabrous, erect or ascending, branched, 
10’-2° high. Leaves oblong or oblong- 
spatulate, obtuse, more or less clasping at 
the base, sharply spinulose-dentate, some- 
times laciniate, 3’-14’ long, 3”-0” wide; 
heads 10’-15” broad, very glutinous; bracts 
of the involucre linear-lanceolate, subulate- 
tipped, strongly squarrose; achenes trun- 
cate, those of the outer flowers usually 
thicker than those of the inner; rays up to 
1’ long or wanting; pappus of 2 or 3 awns. 

In dry soil, Illinois and Minnesota to Mani- 
toba, Missouri, Texas, Arizona and Mexico. 


Adventive in southern New Jersey, Pennsyl- 
vania and New York. June—Sept. 


AY i 


2. Grindelia lanceolata Nutt. Nar- 
row-leaved Gum-plant. Fig. 4193. 


Grindelia lanceolata Nutt. Journ. Acad. Phil. 
T7154 


Slender, erect, glabrous, branched, about 
2° high. Leaves lanceolate or linear, acute 
at the apex, sessile or clasping at the base, 
spinulose-dentate, laciniate, or the upper 
entire, 1-2’ long, 2”-5” wide; heads nearly 
as large as those of the preceding species; 
bracts of the involucre linear-subulate, the 
inner erect, the outer spreading; achenes 
1-2-toothed; pappus of I or 2 awns. 


In dry soil, Tennessee to Missouri, Kansas, 
Louisiana and Texas. July—Sept. 


3i2 COMPOSITAE. Vot. III. 


13. HETEROTHECA Cass. Dict. Sci. Nat. 21: 130. 1821. 


Erect, hirsute or pubescent, branching herbs, with alternate, mostly dentate leaves, and 
rather large heads of both discoid and radiate yellow flowers, generally solitary at the ends 
of the branches. Involucre hemispheric or broadly campanulate, its bracts imbricated in 
several series, the outer shorter. Receptacle flat, alveolate. Ray-flowers pistillate. Disk- 
flowers perfect, or some of them only staminate. Style-branches flat, their appendages 
lanceolate or triangular. Achenes pubescent, obtuse, those of the ray-flowers thick, those 
of the disk-flowers flattened. Pappus of the ray-flowers obsolete or of a few caducous bristles, 
that of the disk-flowers of an inner row of numerous capillary rough bristles, and an outer 
row of shorter stouter bristles or scales. [Greek, different-case, from the dissimilar achenes.] 


Five or 6 species, natives of the southern United States and Mexico, the following typical. 


1. Heterotheca subaxillaris (Lam.) Britton & Rusby. Heterotheca. Fig. 4194. 


Inula subaxillaris Lam. Encycl. 3: 259. 1799. 


Heterotheca Lamarckii Cass. Dict. Sci. Nat. 21: 131. 
1821. 


Heterotheca subaxillaris Britton & Rusby, Trans. N. Y. 
Acad. Sci. 7: 10. 1887. 


Biennial or sometimes annual, 1°-3° high. Basal 
and lower leaves petioled, ovate or oblong, 2’-3’ 
long, the upper ones oblong, sessile or clasping, 
smaller, all acutish or obtuse, dentate; heads rather 
numerous, 6-9” broad; involucre nearly hemispheric, 
3°-5” high, its bracts linear, or slightly dilated above, 
the inner with scarious margins; rays 10-25; inner 
bristles of the pappus of the disk-flowers about 2” 
long. 


In dry soil, Delaware to Florida, Louisiana, Kansas, 
vin ips and Mexico. In ballast, at Philadelphia. July— 
ept. 


14. CHRYSOPSIS [Nutt.] Ell. Bot. S. C. & Ga. 2: 333. 1824. 
[DreLocon Raf. Amer. Month. Mag. 2: 268. 1818. Not Poiret, 1811.] 


Perennial or biennial, rarely annual, branching herbs, with alternate sessile entire leaves, 
or the basal ones dentate, and large many-flowered heads of both tubular and radiate yellow 
flowers (rays wanting in some western species), loosely corymbose, or solitary at the ends 
of the branches. Involucre campanulate to hemispheric, its bracts narrow, imbricated in 
several series, the outer shorter. Receptacle usually flat, more or less foveolate. Ray-flowers 
pistillate. Disk-flowers mostly all perfect. Pappus double in both the disk- and ray-flowers, 
the inner of numerous rough capillary bristles, the outer of smaller or minute scales or 
bristles. Achenes flattened, oblong-linear or obovate, pubescent. Style-branches narrow, 
somewhat flattened, their appendages linear or subulate. [Greek, of golden aspect.] 


About 20 species, natives of North America and Mexico. Besides the following, about 8 others 
occur in the southern and western United States. Type species: Chrysopsis gossypina (Michx.) EIl. 


Leaves elongated-linear, entire, parallel-veined ; achenes linear; involucre campanulate. 


Plants 1°-3° high, silvery-pubescent ; leaves grass-like, 3-12’ long. 1. C. graminifolia. 
Plants 4’-10’ high, woolly-pubescent ; leaves rigid, 1—4’ long. 2. C. falcata. ’ 
Leaves oblong, lanceolate, or linear, pinnately veined; achenes obovate, or oval; involucre hemi- 
spheric. ; 
Plant densely woolly-pubescent. 3. G. gossypina. 


Plants hirsute, or villous-pubescent. f E : 
Heads numerous, corymbose-paniculate ; pubescence of long deciduous hae ae species. 
. mariana, 
Heads fewer, corymbose, or terminating the branches ; pubescence persistent ; western species. 
Villous-pubescent, hirsute or hispid ; perennials. 
Villous-pubescent and canescent with appressed hairs. 
Hirsute or hispid-pubescent. 
Leaves linear, acutish. 
Leaves mostly spatulate, obtuse. 
Pilose-pubescent with soft spreading hairs; annual. 


C. villosa. 


. C. stenophylla. 
. C. hispida. 
. C. pilesa. 


OID 1 


GENUS I4. THISTLE FAMILY. 373 


1. Chrysopsis graminifolia (Michx.) Ell. 
Grass-leaved Golden Aster. Fig. 4195. 


Inula graminifolia Michx. Fl. Bor. Am. 2: 122. 1803. 
Chrysopsis graminifolia Ell. Bot. S. C. & Ga. 2: 334. 
1824. 


Slender, corymbosely branched above, very 
silvery-pubescent, 1°-3° high. Leaves linear, 
soft, grass-like, 3-5-nerved, shining, the basal 
ones 4'-12’ long, 2-5’ wide, the upper much 
smaller, and the uppermost subulate and erect; 
heads several or numerous, about 2’ broad, soli- 
tary at the ends of the branches; involucre cam- 
panulate, its bracts glabrate; achenes linear- 
fusiform. 

In dry soil, Delaware to Florida, Ohio, Kentucky, 


Arkansas, Texas and Mexico. Great Bahama Island. 
Silver-grass. Scurvy-grass. Silk-grass. Aug.—Oct. 


WA 2. Chrysopsis falcata (Pursh) Ell. Sickle- 
P leaved Golden Aster. Fig. 4196. 


Inula falcata Pursh, Fl. Am. Sept. 532. 1814. 
Chrysopsis falcata Ell. Bot. S. C. & Ga. 2: 336. 1824. 


Corymbosely branched above, rather stiff, 4’-12’ 
high, leafy to the top, very woolly-pubescent, at 
least when young, or becoming glabrate. Leaves 
linear, rigid, spreading, sometimes curved, 17-4’ 
long, 1-3” wide, obscurely parallel-nerved; heads 
rather few, corymbose, 3’-5” broad, terminating 
the branches; involucre campanulate, its bracts 
slightly pubescent; achenes linear. 


In sandy soil, eastern Massachusetts to New Jersey. 
Ground gold-flower. July—Aug. 


We 


3. Chrysopsis gossypina 
(Michx.) Ell. Cottony Golden 
Aster. Fig. 4197. 


Erigeron pilosum Walt. Fl. Car. 206. 
1788. 

Inula gossypina Michx. Fl. Bor. Am. 2: 
122. 1803. 

I. gossypina Nutt. Gen. 2: 150. 1818. 

C. gossypina Ell. Bot. S. C. & Ga. 2: 337. 
1824. 

Chrysopsis pilosa Britton, Mem. Torr. 
Club 5: 316. 1894. Not Nutt. 


Stout, 1°-2° high, branched above, 
densely woolly-pubescent all over. 
Leaves spatulate or oblong, obtuse, 
the lower and basal ones 17-3’ long, 
3-5” wide, the uppermost much 
smaller; heads usually nearly 1’ 
broad, terminating the branches, 
bright yellow; involucre hemispheric, 
its bracts densely pubescent when 
young, becoming glabrate; achenes 
obovate. 


y 


In pine-barrens, Virginia to Florida 
and Alabama. Autumn. 


374 COMPOSITAE. Vot. III. 


4. Chrysopsis mariana (L.) 
Ell. Maryland Golden 
Aster. Fig. 4198. 
sore: mariana L. Sp. Pl. Ed. 2, 1240. 

Lz ° 
if =e ee Nutt. Gen. 2: 151. 18718. 

Ee see a Ell, Bot: S; Gi& Gatiz-ts35- 

1824. 

Stout, 1°-23° high, loosely villous- 
pubescent with silky hairs when 
young, at length nearly glabrous, 
corymbosely branched at the sum- 
mit. Upper leaves oblong or lanceo- 
late, acutish or obtuse, sessile, 1/2’ 
long, the lower oblanceolate or spatu- 
late and narrowed into a petiole, 
generally obtuse, 2-4’ long and 
sometimes I’ wide; heads commonly 
numerous, 9’—12” broad, on glandu- 
lar peduncles; involucre hemispheric, 
its bracts glandular, acute, viscid- 
pubescent; achenes obovate. 

In dry soil, southern New York and 
Pennsylvania to Tennessee, Florida and 
Louisiana. Aug.—Sept. Golden-star. 


\ ) 
\ 


z rit 
5. Chrysopsis villdsa (Pursh) Nutt. 
Hairy Golden Aster. Fig. 4199. 
Amellus villosus Pursh, Fl. Am. Sept.564. 1814. 


> 


Inula villosa Nutt. Gen. 2: 151. 1818. 

C. villosa Nutt. Trans. Am. Phil. Soc. II. 7: 
316. 1841. 

C. foliosa Nutt. loc. cit. 316. 1841. 

he camporum Greene, Pittonia 3: 88. 

1897. 

Stem villous or strigose-pubescent, 1°-2° 
high. Leaves oblong, lanceolate, or oblan- 
ceolate, obtuse or acutish, 1-2’ long, 2’-5” 
wide, the upper sessile, the lower narrowed 
into a petiole, pale, persistently canescent 
with appressed hairs; heads rather few, 1’ 
broad or more, terminating the short 
branches; rays oblong-linear, golden yellow ; 
involucre hemispheric, its bracts 4”—5” high, 
linear-subulate, pubescent and often ciliate; 
achenes obovate, 3-5-nerved. 

In dry soil, Minnesota and Illinois to Ala- 
bama, Manitoba, British Columbia, Nebraska 
and New Mexico. July—-Aug. Rosin-wood. 


wy’ 


YF « 
() \) i 


6. Chrysopsis sténophylla (A. Gray) 
Greene. Stiff-leaved Golden 
Aster. Fig. 4200. 


Chrysopsis villosa var. stenophylla A. Gray, 
Syn. Fl. 1: Part 2, 123. 1884. 

C. stenophylla Greene, Erythea 2: 96. 1894. 

C. angustifolia Rydb. Bull. Torr. Club 37: 128. 
IQI0. 


Low, slender, hirsute or rough-pubescent, 
6-10’ high. Leaves linear or slightly broad- 
ened above, densely canescent and ciliate, 
acutish, 9-15” long, 1-2” wide, the mar- 
gins revolute in drying; involucre hemi- 
spheric or broadly campanulate, its bracts 
pubescent or the outer densely ciliate; heads 
few, 6’-10” broad. 

In dry soil, Missouri and Nebraska to Arkan- 
sas and Texas. Aug.—Sept. 


GENUS 14. THISTLE FAMILY. 375 


7. Chrysopsis hispida (Hook.) Nutt. Hispid 
Golden Aster. Fig. 4201. 


Diplopappus hispidus Hook. Fl. Bor. Am. 2: 22. 1834. 
Chrysopsis hispidus Nutt. Trans. Am. Phil. Soc. (II.) 7: 

316. 1841. 

Chrysopsis villosa var. hispida A. Gray, Syn. Fl. 1: Part 
2ye2 3. SOs: 

Lower than C. villosa, stem rarely over 1° high, with 
spreading, sparse or copious, hirsute or hispid pu- 
bescence, sometimes viscid. Leaves spatulate to 
oblong, entire, spreading, 9-18” long, obtuse at the 
apex, narrowed at the base, often into petioles half 
as long as the blade or more; heads smaller, often 
more numerous; involucre not over 4” high, its 
bracts lanceolate, hirsute; achenes 3-5-nerved. 

In dry soil, Manitoba to Idaho, Kansas, Texas and 
Arizona. Consists of several races, differing in size, 
pubescence and leaf-form. July—Sept. . 

vv 
S\\\ | WY 


AS 


8. Chrysopsis pilosa Nutt. Nuttall’s Golden 
Aster. Fig. 4202: 


Clr pspe sis pilosa Nutt. Journ. Acad. Nat. Sci. Phila. 7: 
pelo s4e 
C. Nuttallii Britton, Mem. Torr. Club 5: 316. 1894. 
Pilose-pubescent with soft hairs and minutely glan- 
dular, 1°-2° high, branched above. Leaves oblong- 
lanceolate, mostly acute, 1-23’ long, 2’-5” wide, 
entire, or the lower serrate or even incised; heads 
few, 8-12” broad, terminating the branches; invo- 
lucre hemispheric, its bracts glandular-viscid; achenes 
obovate, 10-nerved. 
In dry soil, Missouri and Kansas to Louisiana, Arkan- 
sas and Texas. July—Sept. 


15. CHONDROPHORA Raf. New FI. N. 
Rnd Os 1836: 
[BicELow1A DC. Mem. Comp. pl. 5. 1833. Not Bigelovia Spreng. 1821.] 

Perennial stiff herbs, with alternate entire leaves, and corymbose-paniculate discoid 
heads of yellow flowers. Involucre oblong or campanulate, its bracts rigid and glutinous, 
imbricated in several series. Receptacle flat, generally foveolate, naked. Corolla regular, 
tubular, the limb 5-cleft. Anthers obtuse at the base. Style-branches flattened, their 
appendages lanceolate. Achenes oblong, somewhat compressed, 1I-2-ribbed on each side. 
Pappus of I or 2 series of numerous capillary unequal bristles. [Greek, cartilage-bearing. ] 

Two species, native of eastern North America, the following typical. 


1. Chondrophora nudata (Michx.) Britton. Rayless Golden-rod. Fig. 4203. 
Chrysocoma nudata Michx. Fl. Bor.Am.2: 101. 1803. ‘ 4 \ 1 Wag p 
Bigelowia nudata DC. Prodr. 5: 329. -1836. 
C. nudata Britton, Mem. Torr. Club 5: 317. 1894. 


10° 


Glabrous, erect, simple, 1°-23° high. Lower 
and basal leaves spatulate, obtuse, or obtusish, 
3-6’ long, 3-6” wide, attenuate into a margined 
petiole; stem leaves distant, small, linear or subu- 
late; heads numerous, 2”-3” high, crowded in a 
compound terminal corymbose cyme; involucre 
narrowly campanulate, acute at the base, its bracts 
coriaceous, appressed, linear-oblong, imbricated 
in 3 or 4 series, the outer much smaller than the 
inner; achenes_ short-turbinate; pappus-bristles 
rigid; edges of the depressions in the receptacle 
prolonged into subulate teeth. 

In moist pine-barrens, New Jersey (?) to Florida 
and Texas. Aug.—Oct. 

Chondrophora virgata (Nutt.) Greene, with nar- 
rowly linear basal leaves, or some of them linear- 
spatulate, though originally cited by Nuttall as from 
New Jersey, is not definitely known from north of 
North Carolina. 


376 COMPOSITAE. Vor. III. 


16. CHRYSOTHAMNUS Nutt. Trans. Am. Phil. Soc. (II) 7: 323. 1840. 


Low shrubs, with equably leafy branches, hard wood, linear leaves, and discoid heads 
of yellow perfect flowers. Heads narrow, 5-7-flowered. Involucre oblong to narrowly 
campanulate, its bracts more or less keeled, thin, or papery, impricated in several series, 
often so as to form 5 vertical rows. Corolla 5-toothed, Anthers obtuse at the base. Style- 
branches exserted, their appendages subulate to filiform. Achenes narrow, mostly pubescent. 
Pappus of copious capillary roughened bristles. [Greek, golden-bush.] 


About 30 species, natives of western North America. Type species: Chrysothamnus pumilus Nutt. 


Heads about 6” high; involucral bracts obtuse or mucronulate. 1. C. graveolens. 
Heads 7”-10” high; involucral bracts subulate-tipped. 2. C. Howardi. 


1. Chrysothamnus gravéolens( Nutt. ) 
Greene. Fetid Rayless Golden- 
rod. Fig. 4204: 


Chrysocoma graveolens Nutt. Gen. 2: 136. 
1818. 

Bigelovia graveolens A. Gray, Proc. Am. Acad. 
8: 644. 1873. 

Chrysothamnus graveolens Greene, Erythea 3:- 
108. 1895. 


Sparingly tomentose above, or glabrate, 
much branched, very leafy, 1°-4$° high, 
odorous. Leaves linear, 1-3’ long, 1’—2” 
wide; heads 5’-6” high, very numerous and 
crowded in terminal compound corymbose 
cymes; rays none; involucre narrowly cam- 
panulate, acute at the base, its bracts oblong 
or linear-oblong, acute or acutish, imbri- 
cated in about 4 series; achenes linear; 
pappus-bristles soft, copious. 

In sterile, especially alkaline soil, Montana 
to western Nebraska, Utah and New Mexico. 
Included in our first edition in the C. nauseosa 
(Pursh) Britton, a densely tomentose far 
western species. Rabbit-brush. Aug.—Oct. 


2. Chrysothamnus Howardi (Parry) 
Greene. Howard’s Rayless Golden- 
rod. Fig. 4205. 


Linosyris Howardi Parry; A. Gray, Proc. Am. 
Acad. 6: 541. 1865. 


Bigelovia Howardi A. Gray, Proc. Am. Acad. 8: 
641. 1873. 


Chrysothamnus Howardi Greene, Erythea 3: 113. 
1895. 


Similar to the preceding species, white- 
tomentose when young, glabrous or nearly so 
when old, tufted, much branched, 6—10’ high. 
Leaves narrowly linear, entire, 1-2’ long, 
about 1” wide; heads 7’-10” long, narrow, 
5-flowered, more or less glomerate, usually 
surpassed by the upper leaves; rays none; 
involucre slightly arachnoid-pubescent, its 
bracts lanceolate, thin, acuminate or subulate- 
tipped; achenes nearly linear, pubescent. 


In dry soil, western Nebraska to Wyoming, 
Colorado, Utah and New Mexico. July—Sept. 


GENUS 17. THISTLE FAMILY. 377 


£7: OONOPSIS [Nutt.] Greene, Pittonia 3: 45. 1896. 


Shrubs or herbs, the following species glabrous, mostly tufted, with woody roots, the 
stems leafy to the top. Leaves entire, linear to lanceolate. Heads cymose at the end of 
the stem or branches. Involucre campanulate to ovoid, its bracts flat, imbricated in several 
series, herbaceous, cuspidate, appressed and erect, or the outer spreading. Disk-flowers with 
a nearly cylindric 5-toothed corolla. Stamens and style included or scarcely exserted. Ray- 
flowers present or wanting, pistillate. Style-appendages ovate to subulate. Achenes glabrous, 
or somewhat pubescent. Pappus-bristles coarse, rigid. [Greek, resembling an egg, referring 
to the ovoid involucre. ] 


Four known species, natives of central North America. Type species: Oonopsis multicaulis 
(Nutt.) Greene. 


1. Oonopsis Engelmanni (A. Gray) Greene. Engelmann’s Oonopsis. Fig. 4200. 


Bigelovia Engelmanni A. Gray, Proc. Am. Acad. 11: Nias | y 
Fen Lo70. SN \ | Y 
SW tei Y, 

Oonopsis Engelmanni Greene, Pittonia 3: 45. 1896. WA | WV, 


Perennial by a deep woody root, glabrous 
throughout; stems stiff, about 8’ high, densely 
leafy. Leaves narrowly linear, sessile, 9-2’ long, 
less than 1” wide, brittle when dry; heads clus- 
tered, or sometimes solitary at the ends of the 
branches, 3’ wide or less, sessile among the upper 
leaves; involucre oblong-campanulate, its bracts 
in about 4 series, oblong to spatulate, short- 
acuminate or mucronate, appressed; ray-flowers 
none; disk-flowers about as long as the rather 
rigid capillary pappus-bristles; achenes linear- 
oblong, narrowed at the base, many-striate. 


aes Kansas, Nebraska and Colorado. Sept.— 
Gti 


18. PRIONOPSIS Nutt. Trans. Am. Phil. Soc. (II) 7: 329. 1841. 


A glabrous annual or biennial herb, leafy to the top, with sessile spinulose-dentate leaves, 
and large heads of yellow radiate and tubular flowers. Involucre broadly hemispheric, its 
bracts imbricated in several series, lanceolate, acuminate, the outer more or less spreading. 
Receptacle naked. Disk-flowers perfect, their corollas 5-toothed. Ray-flowers very numer- 
ous, pistillate. Achenes glabrous, those of the ray-flowers broader than those of the disk; 
pappus of a few deciduous, rigid, unequal bristles, the outer very short. [Greek, resembling 
a saw, referring to the leaf-margins. ] 


A monotypic genus of south-central United States. 


1. Prionopsis ciliata Nutt. Prionopsis. 
Fig. 4207. 
Donia ciliata Nutt. Journ. Acad. Nat. Sci. Phila. 2: 118. 
1821. 
Aplopappus ciliatus DC. Prodr. 5: 346. 1836. 
Prionopsis ciliata Nutt. Trans. Am. Phil. Soc. (II.) 7: 
329. 1841. 


oOo ~O 


Stem erect, stout, branched, very leafy, 2°-5° high. 
Leaves sessile, oval or the lower obovate, obtuse, con- 
spicuously veined, 1’-3’ long, 4’-13’ wide, sharply ser- 
rate with bristle-pointed teeth; heads few, clustered, 
stalked or nearly sessile, 1’-13’ broad; involucre de- 
pressed-hemispheric, its bracts glabrous; achenes of 
the ray-flowers ellipsoid, those of the disk-flowers 
oblong, the central sterile; pappus-bristles rigid, the 
inner ones rough or ciliate. 


On hillsides and river-banks, Missouri and Kansas to 
Texas. Aug.—Sept. 


378 COMPOSITAE. Vou. III. 


19. SIDERANTHUS Fraser, Cat. 1813; Sweet, Hort. Brit. 227. 1826. 
[ErtocarpuM Nutt. Trans. Am. Phil. Soc. (11) 7: 320. 1841.] 


Perennial or annual herbs or shrubs with alternate spinulose-dentate or lobed leaves and 
many-flowered heads of tubular or of both tubular and radiate yellow flowers (heads rarely 
without rays). Involucre hemispheric to campanulate, its bracts imbricated in several series, 
the outer ones gradually smaller. Receptacle flat or convex, generally foveolate, naked. 
Ray-flowers fertile. Disk-flowers usually perfect. Anthers obtuse and entire at the. base. 
Style-branches flattened, their appendages short, lanceolate. Achenes oblong or obovoid, 
obtuse, white-tomentose, or canescent, usually 8-10-nerved. Pappus of 1-3 series of numer- 
ous capillary persistent more or less unequal bristles. [Greek, iron-flower.] 


About 15 species, natives of America. Besides the following, about to others occur in the 
western parts of the United States. Type species: Sideranthus spinulosus (Nutt.) Sweet. 


Rays none; leaves dentate. - 1. S. grindelioides. 
Rays present. 
Leaves dentate; annual. 2. S. annuus. 
Leaves pinnatifid ; perennial. 3. S. spinulosus. 


1. Sideranthus grindelioides ( Nutt.) Britton. 
Rayless Sideranthus. Fig. 4208. 


NEY, 


y 


Eriocarpum grindelioides Nutt. Trans. Am. Phil. Soc. 
Cle) ayer GAT 


Aplopappus Nuttallii T. & G. Fl. N. A. 2: 240. 1842. 


Sideranthus grindelioides Britton, Manual 932. 1901. 


Perennial by a deep woody root, finely pubescent; 
stems tufted, simple, erect, 4-12’ high. Leaves ob- 
long-lanceolate to spatulate, sessile, or the lower peti- 
oled, 4’-1’ long, firm, acute or obtusish, spinulose- 
dentate; heads several or solitary, terminating the 


stem or branches; peduncles 1’ long, or less; invo- 
\ lucre campanulate, its bracts linear, acute, puberulent, 
y ~ their tips somewhat spreading, the outer shorter than 


the inner; achenes densely silky tomentose. 


In dry soil, South Dakota to Assiniboia, Nebraska, 
New Mexico and Arizona. July—Aug. 


2. Sideranthus annuus Rydb. Viscid 
Sideranthus. Fig. 4200. 


we 
Sideranthus annuus Rydb. Bull. Torr. Club 31: SN 
653. 1904. = 


Aplopappus rubiginosus A. Gray, Syn. Fl. 17: 130. \ A 
1884. Not T. & G. AN Bz. 
\ WZ 
Viscid, glandular-pubescent, erect, annual, x 


branched near the summit, 1°-3° high. Leaves 
sessile, or the lowest narrowed into short 
petioles, oblong, lanceolate, or oblanceolate, 
conspicuously dentate with distant awn-point- 
ed teeth, acute or obtusish at the apex, 14’-2}’ 
long, 2”-6” wide; heads several, cymose- 
paniculate, 8-15” broad; involucre hemi- 
spheric, its bracts linear-subulate with spread- 
ing tips; rays large; pappus bristles rigid, very 
unequal; achenes villous-canescent, turbinate, 
not compressed. 

On plains and in canons, Nebraska, Kansas 


and Colorado. Erroneously referred’ in our first 
edition to Eriocarpum rubiginosum. Aug —Sept. 


GENUS Io. THISTLE FAMILY. 


3. Sideranthus spinulosus ( Nutt.) Sweet. NN WK Ke 
Cut-leaved Sideranthus. Sapo. ae 
Fig. 4210. WN 


Amellus spinulosus Pursh, Fl. Am. Sept. 2: 564. 
1814. 

Sideranthus spinulosus Sweet, Hort. Brit. 227. 
1826. 

Aplopappus spinulosus DC. Prodr. 5: 347. 1836. 

Eriocarpum spinulosum Greene, Erythea 2: 108. 


1894. 
S. glaberrimus Rydb. Bull. Torr. Club 27: 621. 


1900. 


Canescent or glabrate, much branched at the 
base, perennial by thick woody roots, 615’ 
high. Leaves pinnatifid, sessile, linear to ovate 
in outline, 2’-14’ long, 1-22” wide, the lobes 
with bristle-pointed teeth; heads several or nu- 
merous (rarely solitary), 6-12” broad; invo- 
lucre hemispheric, its bracts linear, acute, ap- 
pressed; rays narrow; achenes pubescent, nar- 
rowed below; pappus soft and capillary. 


In dry soil, Minnesota and North Dakota to 
Saskatchewan, Alberta, Colorado, Nebraska, 
Texas and Mexico. March-Sept. 


20. STENOTUS Nutt. Trans. Am. Phil. Soc. (Cl) 75 3345. TOs. 


Low undershrubs, with coriaceous narrow entire evergreen leaves, scapose or leafy stems, 
and rather large heads of both radiate and tubular yellow flowers. Involucre mostly hemi- 
spheric, its bracts imbricated in several series, appressed, ovate to lanceolate. Receptacle 
alveolate. Disk-flowers perfect, their corollas tubular, usually somewhat enlarged upward, 
deeply 5-toothed. Ray-flowers fertile’ Anthers obtuse at the base. Appendages of the 
style-branches short, lanceolate. Achenes white-villous. Pappus of soft white capillary bris- 
tles. [Greek, narrow, referring to the leaves. | 


About 18 species, natives of western North America. Type species: Stenotus acaulis Nutt. 


I. Stenotus armerioides Nutt. Narrow-leaved 
Stenotus. Fig. 4211. 


Stenotus armerioides Nutt. Trans. Am. Phil. Soc. (II.) 

FES ahtodits 
Aplopappus armerioides A. Gray, Syn. Fl. 1: Part. 2, 

P32. Leod. 

Perennial, tufted from a branched woody caudex, 
glabrous throughout; flowering stems slender, naked 
above, or quite leafless, 4-8’ high. Basal leaves nu- 
merous, narrowly spatulate or linear, acute or acutish, 
firm, 1-3’ long, 1”-2” wide, entire, narrowed below; 
stem leaves usually 1-3, sessile, linear, sometimes 
none; head commonly solitary, about I’ broad; in- 
volucre campanulate, 4’-6” high, its bracts broadly 
oval, green, appressed, obtuse or retuse, scarious- 
margined, or the inner ovate-oblong and acutish; 
rays 8-10; achenes canescent or villous; pappus bris- 
tles soft, white. 


In dry, mostly rocky soil, western Nebraska to Wyo- 
ming, Utah and New Mexico. June—July. 


21. ISOPAPPUS T. &G. Fl. N. A. 2: 230. 1841. 


Rough-hairy annual or biennial herbs, loosely paniculately branched, with alternate linear 
to lanceolate, I-nerved, entire or somewhat toothed leaves, and small slender-peduncled heads 
of radiate and tubular yellow flowers. Involucre campanulate-cylindric, its appressed lan- 
ceolate or subulate bracts in 2 or 3 series. Receptacle alveolate. Ray-flowers 5-12, pistil- 
late. Disk-flowers 10-20, perfect. Anthers not sagittate. Style-appendages narrow, hirsute. 
Achenes terete, narrowed below, silky-villous. Pappus a single series of rough capillary 
bristles, nearly equal in length. [Greek, equal-pappus. ] 


Two known species, natives of the southern United States, the following typical. 


380 COMPOSITAE. Vor. ITI. 


1. Isopappus divaricatus (Nutt.) T. & G. 
Isopappus. Fig. 4212. 
fe [nula divaricata Nutt. Gen. 2: 152. 1818. 


ae ls a divaricatus A. Gray, Syn. Fl. 1: Part 2, 130. 
1884. 


Isopappus divaricatus T.& G. Fl. N. A. 2: 239. 1841. 


Annual or biennial, erect, paniculately much 
branched, slender, rough-pubescent or glandular, 
1°-3° high. Leaves linear, linear-lanceolate, or the 
ja. \owest linear-spatulate, acute or cuspidate, dentate 

with distant teeth, or sometimes entire, 1’-3’ long, 
1-3” wide, the uppermost much smaller, subulate 
or bract-like; heads numerous, 3’—5” broad; invo- 
lucre campanulate, its bracts linear-lanceolate, subu- 
late-tipped, pubescent, peduncles very slender, or 
filiform. 


In dry soil, Kansas to Texas, east to Georgia and Flor- 
ida. Aug.—Oct. 


22. SOLIDAGO L. Sp. Pl. 878. 1753. 


Perennial erect herbs, sometimes woody at the base, simple, or little branched, with alter- 
nate simple toothed or entire leaves, and small heads of both tubular and radiate, yellow or 
rarely white flowers, in terminal or axillary panicles, thyrsi, or cymose-corymbose or capitate 
clusters. Involucre oblong or narrowly campanulate, its bracts imbricated in several series, 
the outer successively shorter. Receptacle small, flat, or somewhat convex, generally alveo- 
late. Ray-flowers in one series, pistillate. Disk-flowers mostly all perfect, their corollas 
tubular or narrowly campanulate, 5-cleft or 5-lobed. Anthers obtuse and entire at the base. 
Style-branches flattened, their appendages lanceolate. Achenes terete or angled, usually 
ribbed. Pappus of numerous capillary rough nearly equal bristles in 1 or 2 series. [Greek. 
to make whole.] Golden-rod; also called yellow-top, yellow-weed or flower-of-gold. 


About 125 species, mostly of North America, 2 or 3 in Europe, a few in Mexico and South 
America. Type species: Solidago Virgaurea L. 


A. Tips of the involucral bracts, or some of them spreading or recurved. 


Leaves glabrous or glabrate, 4’—10’ long. 1. S. squarrosa. 
Leaves rough-ciliate, 1’—2’ long. 2. S. petiolaris. 


B. Tips of the involucral bracts all erect and appressed. 


* Heads in axillary clusters, or also in a terminal spike-like, sometimes branched thyrsus. 


1. Heads 2”—3” high, chiefly in axillary clusters; achenes pubescent. 


Stem and branches terete; leaves lanceolate to oblong. 3. S. caesia. 
Stem and branches grooved or angled. 
Leaves broadly oval, contracted into margined petioles. 4. S. flexicaulis. 
Leaves lanceolate, sessile. 5. S. Curtisit. 


2. Heads 2”—3” high, chiefly in a terminal spike-like thyrsus ; achenes glabrous, or nearly so. 


Rays white; stem pubescent. 6. S. bicolor. 
Rays yellow; stem densely pubescent. 7. S. hispida. 
Rays yellow ; stem glabrous, or sparingly pubescent. 
Leaves thick, not acuminate, dentate, or the upper entire. 8. S. erec'a. 
Leaves thin, acuminate, sharply serrate. 9g. S. monticola. 


3. Heads 5”—6” high ; bracts elongated, acute ; leaves ovate. 10. S. macrophylla. 
** Heads in a terminal simple or branched thyrsus, not at all secund on its branches, or scarcely so. 


Plant rough-pubescent or scabrous; leaves sessile, entire; bracts acute. 11. S. Lindheimeriana. 
Plants glabrous, puberulent, or sparingly pubescent above. 
Low arctic-alpine species, seldom over 10’ high. 
Heads with 30 flowers or more. 12. S. Cutleri. 
Heads with fewer than 30 flowers. 13. S. multiradiata. 
Taller species, not arctic-alpine. 
Bracts of the involucre linear-subulate, very acute; stem puberulent. 14. S. puberula. 
Bracts of the involucre obtuse or merely acutish ; stem glabrous, or sparingly pubescent above. 
Glabrous throughout; upper leaves very small, appressed. 15. 9. stricta. 
At least the involucre or peduncles pubescent or puberulent. 


GENUS 22. THISTLE FAMILY. 381 


Bog species; inflorescence wand-like. 16. S. uliginosa. 
Upland species, the inflorescence various. 
Heads very short-peduncled. 
Leaves thick, firm in texture, little toothed or entire; tall species. 
Lower leaves ovate to broadly oval, serrate. 17. S. speciosa. 
Leaves lanceolate to oblong-lanceolate, entire. 18. S. rigidiuscula, 
Leaves thin in texture, at least the lower serrate ; low species. 
19. S. Randii. 
Heads distinctly slender-peduncled. 


Basal leaves narrowly oblanceolate, 4” wide or less. 20. S. racemosa. 
Basal leaves broadly oblanceolate or obovate, 6’—18” wide. 
Heads 4” high; basal leaves sharply serrate. 2t. S. Gillmani. 
Heads 3” high; leaves nearly entire. 22. S. sciaphila. 


*** Heads in a terminal, usually large panicle, secund on its spreading or recurved branches. 
: = Plant maritime; leaves thick, fleshy, entire. 23. S. sempervirens. 
ti Plants not maritime ; leaves not fleshy. 
1. Leaves pinnately-veined, not triple-nerved. 
(a) Leaves all entire, thin and glabrous. 24. S. odora. 
(b) Leaves, at least the lower, more or less dentate or serrate. 
{ Leaves linear or linear-oblong, 1’—2' long, scabrous. 25. S. tortifolia. 
tt Leaves broader, lanceolate, oblong, or ovate, 2’—10’ long. 


£ Stem densely pubescent; leaves more or less so. 


Leaves not rugose, sparingly dentate or entire. 26. S. fistulosa. 
Leaves rugose-veiny beneath, sharply serrate. 27. S. rugosa. 


££ Stem glabrous, or merely puberulent above. 
Leaves very rough on the upper surface, serrulate. 28. S. patulu. 


Leaves smooth, or minutely roughened on the upper surface. 
Racemes few, widely divergent, very slender. 


Lower leaves oblong, coarsely serrate, thin. 29. S. ulmifolia. 

Lower leaves ovate or lanceolate, rather finely serrate, firm. 30. S. Boottii. 
Racemes numerous, spreading, recurved or ascending. 

Leaves all oblong or oblong-lanceolate, sessile. 31. S. Elliottit. 


At least the lower leaves petioled, lanceolate or ovate-lanceolate. 
Leaves firm, ovate-lanceolate or oblong-lanceolate; heads about 2” high; racemes short; 
rays several. 32. S. neglecta. 
Leaves firm, narrowly lanceolate; heads about 2” high; racemes few, short; rays 1-5. 
33. S. uniligulata. 
Leaves firm, lanceolate or oval-lanceolate; heads 11%4”-2%” high; racemes numerous, 


slender. 34. S. juncea. 
Leaves thin, the lower broadly ovate, short-acuminate; heads 2%4”-3%” high; racemes 
numerous. 35. S. arguta. 


2. Leaves triple-nerved, 7. e., with a pair of lateral veins much stronger than the others. 


Heads small, the involucre only 1%” high or less; stem glabrous, or pubescent. 36. S. canadensis. 
Heads larger, the involucre 1%4”—3” high. 
Stem glabrous. 


Leaves, and involucral bracts thin. 37. S. serotina. 
Leaves, and involucral bracts firm, somewhat rigid. 
Leaves linear-lanceolate ; achenes glabrous. 38. S. glaberrima. 
Leaves oblong-lanceolate; achenes silky-pubescent. 39. S. Shortit. 
Basal leaves oblanceolate, upper bract-like. 40. S. Ga’tingeri. 
Stem pubescent or scabrous. 
Leaves lanceolate, sharply serrate or entire, rough above. 41. S. altissima. 


Leaves oblanceolate, spatulate, oblong, or ovate, the lower crenate. ; 
Minutely rough-pubescent, grayish; lower leaves oblanceolate; heads 2”—3” high. 
42. S. nemoralts. 


Canescent and pale; leaves oblong or ovate; heads 3” high. 43. S. mollis. 
Very scabrous, green, not grayish, nor canescent. 44. S. radula. - 
Leaves broadly ovate-oval, sharply serrate, finely pubescent. 45. S. Drummondit. 


**** Heads in a terminal, corymbiform, sometimes thyrsoid cyme, forming a flat-topped inflorescence. 


(Genus OLIGONEURON Small.) 


Leaves ovate, oblong, or oval, mostly rough on both sides. 46. S. rigida. 
Leaves lanceolate, linear, oblong, or oblanceolate, glabrous or nearly so. 
Lower leaves oblong-lanceolate, serrulate. 47. S. ohioensis. 
Leaves all lanceolate or linear, entire. 
Stout; leaves lanceolate, the basal 8’—12’ long. 48. S. Riddellii. 


Slender ; leaves linear, the basal 4’—5’ long. 49. S. Houghtoni. 


352 COMPOSITAE. Vor. III. 


1. Solidago squarrésa Muhl. Stout 
Ragged Golden-rod. Fig. 4213. 
Solidago squarrosa Muhl. Cat. 76. 1813. 


Stem stout, simple, or rarely branched above, 
glabrous or puberulent, 2°-5° high. Upper 
leaves oblong, acute, entire or nearly so, ses- 
sile; lower and basal leaves obovate, oval, or 
broadly spatulate, acute or obtuse, 4-10’ long, 
1-3’ wide, sharply dentate, often narrowed 
into-a margined petiole, all glabrous, or some- 
times slightly pubescent; heads 15-25-flow- 
ered, 4-5’ high, numerous in a terminal nar- 
row often leafy thyrsus sometimes 12’ in 
length; rays 10-16, showy, 2-3” long; tips of 
the involucral bracts green, acute or obtuse, 
rarely some of them erect, all usually strongly 
recurved, giving the heads a ragged appear- 
ance; achenes glabrous. ; 

In rocky soil, New Brunswick to Ontario, North 
Carolina and Ohio. Ascends to 2000 ft. in the 


Catskills. Lower branches of the inflorescence 
sometimes elongated. Aug.—Oct. 


2. Solidago petiolaris Ait. Downy Ragged 
Golden-rod. Fig. 4214. 


Solidago petiolaris Ait. Hort. Kew. 3: 216. 1789. 
Solidago Wardii Britton, Man. 935. 1901. 

Stem rather slender, pubescent or puberulent, 1°-3° 
high, simple, or branched above. Leaves sessile, or 
very short-petioled, oblong to oval, acute, obtuse or 
mucronate, entire or dentate, 2-3’ long, rough or ciliate 
on the margins, often silvery-pubescent; heads 3’—4” 
high, in a terminal narrow more or less compound 
thyrsus; involucral bracts pubescent, with green acute 
to acuminate tips, the outer spreading, the inner ap- 
pressed; achenes glabrous or nearly so. 

In dry soil, Illinois to Kansas and Texas, east to North 


Carolina and Florida. Races differ in pubescence and in 
leaf-form and leaf-serration. Sept.—Oct. 


3. Solidago caésia L. Blue-stemmed 
or Wreath Golden-rod. Fig. 4215. 


Solidago caesia L. Sp. Pl. 879. 1753. 

S. gracilis Poir. in Lam. Encycl. 8: 476. 1808. 
S. axillaris Pursh, Fl. Am. Sept. 542. 1814. 

S. caesia axillaris A. Gray, Proc. Am. Acad. 17: 

189. 1882. 

Stem glabrous, slender, often glaucous, 
usually bluish or purple, branched or sim- 
ple, terete, 1°-3° high. Leaves lanceolate or 
oblong-lanceolate, sessile, acuminate at the 
apex, narrowed at the base, glabrous, sharply 
serrate, 2’-5’ long, 3-15” wide; heads 2”-3” 
high, in axillary clusters or racemes, or oc- 
casionally with some in a short terminal 
thyrsus; bracts of the involucre obtuse, ap- 
pressed; achenes pubescent. 

Woods and thickets, Nova Scotia to Ontario, 
Minnesota, Florida, Arkansas and Texas. Con- 


sists of several slightly differing races. Wood- 
land golden-rod. Aug.—Oct. 


GENUS 22. THISTLE FAMILY. 


4. Solidago flexicaulis L. Zig-zag or 
Broad-leaved Golden-rod. Fig. 4216. 


Solidago flexicaulis L. Sp. Pl. 879. 1753. 
Solidago la ifolia L: loc. cit. 1753. 


Stem glabrous, angled, usually simple, zig-zag, 
1°-3° high. Leaves thin, ovate, acuminate at the 
apex, abruptly narrowed at the base into mar- 
gined petioles, somewhat pubescent, or glabrous 
beneath, sharply serrate, 2’-7’ long, 1-4’ wide, the 
uppermost sometimes lanceolate and entire or 
nearly so; heads about 3” high, in short axillary 
racemose clusters, and rarely also in a narrow 
terminal thyrsus; bracts of the involucre obtuse 
to acutish, appressed; achenes hirsute-pubescent. 


In rich woods, Nova Scotia and New Brunswick to 
Georgia, Tennessee, Minnesota and Missouri. As- 
cends to 2300 ft. in the Catskills. July—Sept. 


5. Solidago Curtisii T. & G. Curtis’ 
Golden-rod. Fig. 4217. 
Solidago Curtisii T. & G. Fl. N. A. 2: 200. 1841. 


Stem glabrous or sparingly pubescent, sim- 
ple or branched, slender, 13°-3° high, angled 
and grooved. Leaves thin, sessile, elongated- 
lanceolate or sometimes broader above the 
middle, long-acuminate, narrowed below into 
an entire base, sharply serrate, 3-6’ long, 
4-12” wide, glabrous or nearly so; heads 
2”-3” high, in rather loose axillary clusters 
and sometimes also in a narrow terminal 
thyrsus; bracts of the involucre few, obtuse. 

In mountain woods, Virginia and West Vir- 
ginia to Kentucky and Georgia. Aug.—Sept. 

Solidago pibens M. A. Curtis, of nearly the 
same range, differs in being quite densely pubes- 
cent. 


6. Solidago bicolor L. White or 
Pale Golden-rod. Silver-rod. 
Fig. 4218. 

Solidago bicolor L. Mant. 114. 1767. 


1 Oy, NN \ 
BN EN SNR 


Stem rather stout, hirsute-pubescent, 
or nearly glabrous, 6—4° high, simple or 
branched. Basal and lower leaves obo- SRY ; om 6S 
vate or broadly oblong, mostly obtuse, Tp, ENOAY, ae A HG Lk) SQ Wee 
2’-4' long, 1-2’ wide, narrowed into j bp i ¥ 
long margined petioles, dentate or cre- 
nate-dentate, more or less pubescent; 
upper leaves smaller and narrower, ob- 
long or sometimes lanceolate, obtusish 
or acute, sessile or nearly so, often en- 
tire; heads 2-3” high, crowded in a 
terminal narrow thyrsus 2’-7’ long, and 
sometimes also clustered in the upper 
axils; rays white; bracts of the invo- 


q 
See é. - SI iy 
lucre whitish, obtuse, the midvein broad- i 


=: 
ened above; achenes glabrous. v 
: : Aes 
In dry soil, Prince Edward Island to aK 
Georgia, west to Ontario, Minnesota and Yi} Vox 
Tennessee. Ascends to 6300 ft. in North pL 
Carolina. Belly-ache-weed. Silver-weed. VS 


July—Sept. 


_ COMPOSITAE. Vor. III. 


7. Solidago hispida Muhl. Hairy 
Golden-rod. Fig. 42109. 


S. hispida Muhl.; Willd. Sp. Pl. 3: 2063. 1804. 

Solidago hirsuta Nutt. Journ. Phil. Acad. 7: 
103. 1834. 

Solidago bicolor concolor T. & G. Fl. N. A. 2: 
197. 1841. 


_Stout, stem densely pubescent or hirsute, 
simple or sometimes branched, 13°-3° high. 
Lower leaves oval, acute, or obtuse, peti- 
oled, pubescent on both sides, usually den- 
tate, 2’-5’ long, 1’-2’ wide; upper leaves 
oblong, sessile, acute, dentate or entire, 
smaller, sessile, heads about 3” high, crowd- 
ed in a dense narrow terminal thyrsus and 
also often in racemose clusters in the upper 
axils; rays yellow; involucral bracts yel- 
lowish, obtuse, the midvein narrow; achenes 
with a few appressed hairs, or glabrous. 

In dry soil, Newfoundland to western On- 
tario, Manitoba, New Jersey, Pennsylvania, 
Georgia and Missouri. Ascends to 2000 ft. in 
the Catskills. Aug.—Oct. 


VW 


8. Solidago erécta Pursh. Slender 
Golden-rod. Fig. 4220. 


Solidago erecta Pursh, Fl. Am. Sept. 542. 
1814. 

Solidago speciosa angustata T. & G. Fl. N. A. 
2: 205. 1841. 


Stem slender, glabrous, or puberulent 
above, 2°-3° high, simple or rarely branch- 
ed. Leaves firm, nearly glabrous on both 
sides, ciliolate on the margins, the lower 
and basal ones broadly oblong or oval, 
obtuse or obtusish, crenate-dentate, the 
upper lanceolate or oblong-lanceolate, acute, 
usually quite entire; heads 2’-3” high ina 
very narrow terminal thyrsus, rarely also 
with a few clustered in the upper axils; 
bracts of the involucre obtuse; rays light 
yellow; achenes glabrous. 


In dry soil, southeastern New York, New 
Jersey and Pennsylvania to Kentucky, Geor- 
gia and Alabama. Aug.—Sept. 


g. Solidago monticola T. & G. 
Mountain Golden-rod. 
Fig. 4221. 


Solidago Curtisti var. monticola T. & G. FI. 
N. A. 2: 200. 1841. 

Solidago monticola T. & G.; Chapm. FI. S. 
States 209. 1860. 


Slender, glabrous or nearly so, 1°-3° 
high. Stem leaves ovate-oblong, or ob- 
long-lanceolate, thin, acuminate at the 
apex, narrowed at the base, sharply and 
sparingly serrate, or the upper entire, 
1-6’ long, 4”-13’ wide, the upper sessile, 
the lower petioled; basal leaves broadly 
oblong, obtuse, with slender petioles; 
heads about 2” high; in a terminal spike- 
like, simple or branched thyrsus; bracts 
of the involucre acutish or obtuse; 
achenes glabrous. 


In mountain woods, Pennsylvania and 
Maryland to Georgia and Alabama. 


GENUS 22. THISTLE FAMILY. 335 


10. Solidago macrophylla Pursh. Large- 
leaved Golden-rod. Fig. 4222. 


Solidago macrophylla Pursh, Fl. Am. Sept. 542. 1814. 
Solidago thyrsoidea E. Meyer, Pl. Lab. 63. 1830. 


Stem striate, glabrous or sparingly pubescent, 
stout, 6’-4° high. Leaves thin, ovate, acuminate, 


WYtz-4 


Ye 


by 


or the uppermost lanceolate, entire, sessile; heads 
4-6” high, in a terminal compact or loose thyrsus 
and usually also in axillary clusters; bracts of 
the involucre linear, acute; rays 8-10, linear-ob- 
long, conspicuous; achenes glabrous or nearly so. 


In rocky woods, Catskill Mountains, N. Y., and 
Greylock Mt., Mass., to Newfoundland, Labrador, 
Hudson Bay and Lake Superior. Ascends to 4000 ft. 
in the Adirondacks. July—Sept. 


Solidago calcicola Fernald, a related plant found 
in Maine and Quebec, has smaller heads, 3”—4” high, 
and pubescent achenes. 


or the basal ones obtuse, sharply serrate, glabrous WA iN 

or sparingly pubescent beneath, 3-5’ long, 1-23’ A\\\ any) / 

wide, abruptly contracted into margined petioles, QW) NY) WZ 
=" Ve 


= 
= 


\ 


——~F 
AN 


$e 11. Solidago Lindheimeriana Scheele. 


Lindheimer’s Golden-rod. Fig. 4223. 


Solidago Lindheimeriana Scheele, Linnaea 21: 599. 
1848. 


Scabrous, simple, 13°-3° high, leafy, rather 
stout and rigid. Leaves oblong, oblong-lanceo- 
late or oval, acute or acutish at the apex, nar- 
rowed or rounded at the base, all entire, thick- 
ish, rough on both surfaces, sessile or the lowest 
petioled; heads about 3” high, in a terminal, 
often short thyrsus; bracts of the involucre 
acute or the outer obtuse, puberulent; achenes 
nearly glabrous. 


Southern Kansas to Texas and northern Mexico. 
Aug.—Nov. 


Solidago Bigelovii A. Gray, another southwestern 
species, which is rougher than this, with oval or ob- 
long leaves obtuse or obtusish at both ends, is re- 

5 \} ported from Kansas, probably erroneously. 


. 


12. Solidago Citleri Fernald. Cutler’s Al- 
pine Golden-rod. Fig. 4224. 
Solidago Virgaurea alpina Bigel. Fl. Bost. Ed. 2, 307. 


1824. 
Solidago Cutleri Fernald, Rhodora 10: 87. 1908. 


Glabrous, or somewhat pubescent; stems simple, 
often tufted, 3-12’ high, ascending, or erect, angu- 
lar. Basal leaves obovate, or broadly spatulate, ser- 
rate with low sharp or blunt teeth, at least above the 
middle, obtuse, or acute, 24’ long, narrowed into 
petioles; stem leaves few, oblanceolate, spatulate, 
or oblong, sessile, or the lower petioled, mostly dis- 
tant; inflorescence a short raceme or thyrsus, and 
often with clusters of heads in the axils of the 
leaves; heads 3-4” high, over 30-flowered; bracts 
of the involucre obtuse to acute; achenes hirsute. 


Alpine summits of the mountains of Maine, New 
Hampshire, Vermont and northern New York, mostly 
above timber line. Referred, in our first edition, to the 
European S. alpestris Waldst. & Kit., which it resembles. 
Aug.-Sept. 


25 


COMPOSITAE. Vo. III. 
Vy 


We) ZZ o A AS e 
WR 13. Solidago multiradiata Ait. Northern 


UZ 
bp : 

(Wz Golden-rod. Fig. 4225. 

Solidago multiradiata Ait. Hort. Kew. 3: 218. 1780. 


Stem glabrous or somewhat pubescent above, 
rather slender, 6-15’ high. Leaves firm, glabrous 
or very nearly so, the basal and lower ones spatu- 
late or oblanceolate, entire, or sparingly serrate, 
obtuse, finely reticulate-veined, 3-5’ long, 3-9” 
wide, the upper smaller, narrower, sessile, _ en- 
tire; heads about 4” high, usually few in a termi- 
nal rather compact, corymbose cyme, but the in- 
florescence sometimes elongated and thyrsoid; 
bracts of the involucre thin, linear-lanceolate, 
acute or acutish; glabrous; rays 8-15, prominent, 
linear, achenes pubescent. : 


Labrador and Hudson Bay to British Columbia, the 
Rocky Mountains and Colorado. July—Aug. 


Solidago decimbens Greene, of the Rocky Moun- 
tains, with broader involucral bracts is recorded from 
Mt. Albert, Quebec. 


14. Solidago pubérula Nutt. Downy 
Golden-rod. Fig. 4226. 
Solidago puberula Nutt. Gen. 2: 162. 1818. 


Minutely puberulent, or glabrous, usu- 
ally simple, rather slender, 13°-3° high, 
leafy. Stem leaves oblong-lanceolate, acute, 

sparingly serrate or entire, I’”—-2’ long, ses- 

sile, or the lower petioled, basal leaves and 
sometimes the’ lowest ones of the stem 
spatulate, obtuse, often sharply serrate, 2-4’ 
long, narrowed into margined petioles; 
heads about 24” high, in a terminal, often 
leafy thyrsus, the branches of which are 
spreading or ascending; bracts of the invo- 
lucre subulate, very acute; achenes gla- 
brous; heads rarely a little secund. 


In sandy soil, Prince Edward Island to h 
Florida and Mississippi, near the coast and on vy 
sandstone rocks in the Appalachian mountain SQ \\!/4 
system, west to Tennessee. Minaret-golden- Vy 

A 


rod. Aug.—Sept. \ 
15. Solidago stricta Ait. Wand-like or 
Willow-leaf Golden-rod. Fig. 4227. 


Solidago s:ricta Ait. Hort. Kew. 3: 216. 17809. 
S. virgata Michx. Fl. Bor. Am. 2: 117. 1803. 


Glabrous throughout, slender, erect, simple, 
2°-8° high. Basal and lowest stem leaves ob- 
long, or somewhat spatulate, with few lateral 
veins, obtuse, entire, or very sparingly dentate, 
3-8’ long, 3’-1’ wide, narrowed into long peti- 
oles; upper stem leaves abruptly smaller, nar- 
rowly oblong, spatulate or linear, appressed, 
the uppermost very small and bract-like; heads 
about 3” high, in a dense simple, or some- 
times branched, naked thyrsus; bracts of the 
involucre oblong, obtuse, or the inner acutish; 
achenes glabrous, or sparingly pubescent. 


In wet sandy pine-barrens, New Jersey to Flor- 
ida and Louisiana. Also in western Cuba. Aug.— 
Oct. 


GENUS 22. THISTLE FAMILY. 


16. Solidago uligindsa Nutt. Bog or 
Swamp Golden-rod. Fig. 4228. 


Solidago uliginosa Nutt. Journ. Phil. Acad. 7: 101. 
1834. 


Stem glabrous, rather stout, simple, 2°-4° | 
high, the branches of the inflorescence more 
or less pubescent. Leaves oblong-lanceolate or 
lanceolate, glabrous, firm, more or less cilio- 
late or scabrous on the margins, few-veined, 
acute or acuminate, the lower and basal ones 
4-9’ long, 4’-12’ wide, more or less serrate and 
narrowed into petioles, the upper smaller, 
mostly sessile and entire; heads 2’-3” high, in 
a terminal oblong dense thyrsus, its branches 
appressed; bracts of the involucre linear-ob- 
long, obtuse; achenes glabrous. 


In swamps and bogs, Newfoundland to New 
Jersey, North Carolina, western Ontario, Minne- 
sota and Wisconsin. Aug.—Sept. 


17. Solidago speciésa Nutt. Showy or 
Noble Golden-rod. Fig. 42209. 


Solidago speciosa Nutt. Gen. 2: 160. 1818. 


Stem stout, glabrous below, often rough 
above,, usually simple, 3°-7° high. Leaves 
glabrous, firm, the lower and basal ovate, or 
broadly oval, 4-10’ long, 1-4’ wide, dentate 
or crenate, pinnately veined, acute or obtuse 
at the apex, long-petioled; upper leaves 
smaller, oblong or oval, acute at each end, 
crenate-dentate, or entire, sessile or short- 
petioled, rough-margined; heads 3’-4” high, 
in a large terminal thyrsus, the branches of 
which are ascending and often leafy; bracts 
of the involucre oblong, very obtuse; achenes 
glabrous or nearly so. 


In rich soil, Massachusetts to North Carolina, 
west to Minnesota, Tennessee and Arkansas. Ap- 
parently erroneously recorded from Canada and 
Nova Scotia. Aug.—Oct. 


18. Solidago rigidiiscula (T. & G.) 
Porter. Slender Showy Golden- 
rod. Fig. 4230. 


Solidago speciosa rigidiuscula T. & G. Fl. N. A. 

p 21205. To4t.. 
Solidago speciosa pallida Porter, Bull. Torr. 
Club 19: 130. 1892. 

Solidago rigidiuscula Porter,,.Mem. Torr. Club 
5: 319. 1894. 

S. pallida Rydb. Bull. Torr. Club 33: 153. 1906. 


Stem rather slender, usually glabrous be- 
low, rough-pubescent above, simple, 2°-4° 
high. Leaves lanceolate to ovate-lanceo- 
late, entire, or the basal ones sometimes 
crenate, strongly ciliolate on the margins, 
1-5’ long, 3’—-12” wide, the upper sessile, 
the lower sometimes narrowed into peti- 
oles; heads similar to those of the pre- 
ceding species; thyrsus generally narrow, 
dense, simple or branched. 


In dry soil, mostly on prairies, Ohio to Ala- 
bama, Ontario, Minnesota, Colorado, Louisi- 
ana and Texas. Aug.—Oct. 


COMPOSITAE. Vor. ITT. 


1g. Solidago Randii (Porter) Britton. Rand’s 
Golden-rod. Fig. 4231. 
Sees Virgaurea Randii Porter, Bull. Torr. Club 20: 208. 
1893. 


Solidago Virgaurea Redfieldii Porter, Bull. Torr. Club 20: 
209. 1893. 

Solidago Virgaura monticola Porter, Bull. Torr. Club 20: 
209. 1893. ‘ 

Solidago Virgaurea Deanei Porter, Mem. Torr. Club 5: 320. 


1894. 

Suldage Randii Britton, Manual 937. 1901. 

Somewhat pubescent, at least above, often glutinous; 
stem usually simple, rather stout, 5—2° high. Basal 
leaves oblanceolate, broadly spatulate, or obovate, 3’-8’ 
long, I’ wide, or less, obtuse or acute, mostly dentate, 
narrowed into margined petioles; stem leaves few, ses- 
sile, or the lower petioled, oblong-lanceolate to spatu- 
late, acute; heads 3-4” high, in a dense or interrupted, 
rarely branched thyrsus and often in axillary clusters; 
bracts of the involucre obtuse or acute; achenes more 
or less pubescent. 

In dry, mostly rocky situations, Maine, New Hampshire, 


Vermont and northern New York to Michigan and on high mountains in Virginia. Referred, in our 
first edition, as by previous authors, to the Old World Solidago Virgaurea L. Aug.—Sept. 


20. Solidago racemosa Greene. 


River-bank 


Golden-rod. Fig. 4232. 
S. racemosa Greene, Pittonia 3: 160. 1897. 


Nearly glabrous, sometimes glutinous; stems sim- 
ple, usually somewhat glutinous, 6-18’ high. Lower 
and basal leaves oblanceolate, obtuse or acutish, den- 
tate, or crenate, 13-4’ long, 23-4” wide, narrowed 
into slightly margined petioles; stem leaves sessile, 
lanceolate, oblong or linear, numerous, crenate or 
entire, mostly acute, smaller; heads about 3” high, 
distinctly peduncled, in a terminal simple or branch- 
ed thyrsus; bracts of the involucre linear-oblong, 
obtuse, or the inner acutish; achenes striate, pubes- 
cent. 

On rocky river-banks, Newfoundland to northern New 
York, Vermont and Virginia. Described, in our first 
edition, under the name S. Purshii Porter, which proves 
to be untenable. July—Sept. 

Solidago chrysdélepis Fernald, of Quebec, has bright 
yellow acute involucral bracts. 


21. Solidago Gillmani (A. Gray) Steele. 
Gillman’s Golden-rod. Fig. 4233. 


S. humilis Gillmani A. Gray, Proc..Am. Acad. 17: 


I9t. 1882. 

S. Virgaurea Gillmani Porter, Bull. Torr. Club 20: 
209. 1893. 

S. Gillmani Steele, Contr. Nat. Herb. 13: 367. 
IQII. 

S. racemosa Gillmani Fernald, Rhodora 10: 91. 
1908. 


Glabrous, except the puberulent inflores- 
cence; stem erect or reclining, rather stout, 
sometimes 3° long. Lower and basal leaves 
spatulate or oblanceolate, dentate, 312’ long, 
narrowed into long narrowly margined peti- 
oles; upper stem leaves lanceolate to linear- 
lanceolate, gradually smaller; inflorescence 
narrowly thyrsoid-paniculate, sometimes 16’ 
long; heads distinctly peduncled, about 4” 
high; bracts of involucre oblong, scarcely 
glutinous; rays 6-10, deep yellow, 2” long; 
achenes sparingly pubescent. 


Sandy shores of Lakes Michigan and Superior. 
Aug.—Sept. 


GENUS 22. THISTLE FAMILY. j 389 


22. Solidago sciaphila Steele. Shadowy 
Golden-rod. Fig. 4234. 


)) 
WSO 
WIG 


AAS 


S. sciaphila Steele, Contr. Nat. Herb. 13: 371. 1911. 


Glabrous, except the ciliate leaf-margins and pu- 
berulent inflorescence, 2°-34° high, light green. 
Leaves thin in texture, sparingly faintly veined, the 
lower spatulate to broadly oblanceolate, obtuse, 3-5’ 
long, narrowed into long petioles, the upper oblong 
to oblong-lanceolate, sessile, smaller; heads race- 
mose on the slender branches of the narrow thyrsus, 
on peduncles as long as the involucre or longer; 
involucre campanulate, about 3” long, its bracts 
linear-oblong, obtuse; rays about 5, light yellow, 
short. 


Shaded cliffs of the Wisconsin River, Sauk County, 
Wisconsin. Aug.—Sept. 


23. Solidago sempérvirens L. Sea-side 
Golden-rod. Fig. 4235. 

Solidago sempervirens L. Sp. Pl. 878. 1753. 

S. angustifolia Ell. Bot. S. C. & Ga. 2: 388. 1824. 

Not Mill. 1768. 

Stem stout, leafy, usually simple, 2°-8° high, 
glabrous or slightly puberulent above. Leaves 
thick, fleshy, entire, with 2-5 pairs of lateral 
veins, the lower and basal ones oblong, spatu- 
late or lanceolate, mostly obtuse, sometimes 1° 
long, narrowed into long petioles; upper leaves 
sessile, lanceolate to oblong-lanceolate, acute; 
heads 3’—5” high, in secund racemes of a large 
terminal often leafy panicle; rays 8-10, showy; 
bracts of the involucre lanceolate, acute. 

On salt marshes, sea-beaches, along tidal rivers 
and in sandy soil near the sea, Nova Scotia and 
New Brunswick to Florida and Mexico. Also in 


Bermuda. Salt-marsh or beach golden-rod. Aug— 
ec. 


24. Solidago odora Ait. Sweet or Anise- e 
scented Golden-rod. Fig. 4236. 


Solidago odora Ait. Hort. Kew. 3: 214. 1780. ’ 
Solidago odora inodora A. Gray, Man. Ed. 5, 244. iy 
1867. RS) BW 


Slender; stem simple, glabrous, or minutely 2G ‘SR 
pubescent above, 2°-4° high. Leaves usually 8 Mi LMS 
punctate, anise-scented when bruised, or some- aN, PY YN SE 
times inodorous, lanceolate, quite entire, acute ap ae 
or acuminate, 2-4’ long, 3-8” wide, sessile, 
or the lowermost petioled; heads 2’-23” high, 
secund on the spreading racemes of the termi- 
nal, usually ample panicle; rays 3 or 4, 2-3” 
long; bracts of the involucre oblong-lanceo- 
late, acute, the inner much longer than the 
outer. 


In dry soil, Nova Scotia (according to Som- 
mers); New Hampshire to Florida, New York, 
Kentucky, Missouri and Texas. Blue mountain- 
tea. True golden-rod. July—Sept. 


$ 
Wwe AA 
VRRA a 

WS 


A 


N 
a 


uv 
DYN 
Petes 


Fra 
SV SAR 
KS SS 


Pine 
Barren Golden-rod. Fig. 4238. 
Solidago fistulosa Mill. Gard. Dict. Ed. 8, No 


26. Solidago fistulosa Mill. 


19. 1768. 
Solidago pilosa Walt. Fl. Car. 207. 
Mill. 1768. 


Stem rather stout, simple or branched 
above, 3°-7° high, hirsute. Leaves numer- 
ous, sessile, ovate-oblong, oblong-lanceolate, 
or sometimes lanceolate, thick, rough or 
hirsute on the margins and midrib beneath, 
the upper small, obtuse or obtusish, entire, 
the lower sparingly serrate, obtuse or acute, 
1-4’ long with a broad basé; heads about 
2” high, secund on the spreading or recury- 
ing branches of ethe panicle; rays 7-10, 
small; bracts of the involucre, at least the 
outer, acute. 


1788. Not 


In moist pine-barrens, New Jersey to Florida 
Aug.—Oct. 


and Louisiana. 


an, 
ay 


i 
AR 
. 


; / a [ly WA Wp: F)) 
AY \y Aa Apr ie IZ 
UE, 
Ie aw Si 
5 \" RAT, y 
Ws 


7 


COMPOSITAE. 


25. Solidago tortifolia Ell. 


mostly near the coast. 


AL SS 


Vo. III. 


Twisted-leaf 
Golden-rod. Fig. 4237. 


Solidago retrorsa Pursh, Fl. Am. Sept. 539. 1814. 
Not Michx. 1803. 
S. tortifolia Ell. Bot. S. C. & Ga. 2: 377. 1824. 


Stem slender, rough-pubescent or puberulent, 
2°-3° high, simple. 
oblong, often twisted, scabrous, sessile, acute, 
1’-2’ long, 13’-3 
with a distinct midrib, the lower serrate, the 
upper entire; heads about 2” high, secund on the 
usually recurved branches of the terminal pani- 
cle; rays 3-5, short; bracts of the involucre 
linear, obtuse or obtusish. 


Leaves linear or linear- 


47 


wide, obscurely veined but 


In dry sandy soil, Virginia to Florida and Texas, 
Autumn. 


4 Adee 
GE 


eS AN), 
Wy LL 
BERL 2 


Wrinkle- 


27. Solidago rugosa Mill. 
leaved, Tall Hairy, or Pyramid 


Golden-rod. Bitter-weed. 
Fig. 4239. 


S. rugosa Mill. Gard. Dict. Ed. 8, No. 25. 
Solidago aspera Ait. Hort. Kew. 3: 212. 


1768. 
17809. 


Stem hirsute or scabrous, rarely glabrate, 
usually stout, 1°-73° high, simple, or 
branched at the summit. Leaves more or 
less pubescent or scabrous, oval, oblong- 
lanceolate, or ovate-lanceolate, acute or 
acuminate, rarely obtusish at the apex, nar- 
rowed or obtuse at the base, rugosely 
veined on the lower surface, serrate, I’-4’ 
long, 4-18’ wide, sessile, or the lowest 
sometimes tapering into petioles; heads 
13-2” high, secund on the spreading or 
recurving, often leafy branches of the usu- 
ally large and compound panicle; bracts of 
the involucre linear, obtuse or obtusish. 


Usually in dry soil, in fields and-along road- 
sides, Newfoundland to western Ontario, south 
to Florida and Texas. Consists of many races, 
differing in leaf-form, leaf-thickness and in 
pubescence. Supposed to hybridize with S. 
sempervirens L. Dyer’s-weed. July—Nov. 


GENUS 22. THISTLE FAMILY. 


28. Solidago patula Muhl. Rough- 
leaved or Spreading Golden-rod. 
Fig. 4240. 


Solidago patula Muhl.; Willd. Sp. Pl. 3: 2059. 
1804. ; 


Stem stout, rather rigid, usually simple, 
glabrous, or sometimes puberulent above, 
2°-7° high. Leaves thick, glabrous beneath, 
exceedingly rough above, pinnately veined, 
the lower and basal ones very large, 3’-16° 
long, 12-5’ wide, oval or elliptic, narrowed 
into margined petioles, the upper smaller, 
oval or oblong, sessile, acute, finely serrate 
or the uppermost entire; heads 2’’—33” high, 
secund on the widely spreading and recuryv- 
ing branches of the loose panicle; rays 
small; bracts of the involucre linear-ob- 
long, obtuse. 


In swamps, Maine and Ontario to Minnesota, 
south to Georgia, Missouri and Texas. As- 
cends to 5000 ft. in North Carolina. Stem 
strongly angled, at least below. Aug.—Oct. 


29. Solidago ulmifolia Muhl. Elm- 
leaved Golden-rod. Fig. 4241. 
Solidago ulmifolia Muhl.; Willd. Sp. Pl. 3: 

2060. 1804. 

Stem slender, glabrous, or puberulent at 
the summit, 2°-4° high, simple, or branched 
above, the arched branches puberulent or 
pubescent. Leaves thin, oblong to elliptic- 
lanceolate, acute or acuminate at the apex 
and base, coarsely and sharply serrate, pin- 
nately veined, glabrous or sparingly pubes- 
cent, the lower and basal ones wider, 3-5’ 
long, 1-13” wide, narrowed into margined 
petioles, the upper smaller, sessile; heads 
2”’-3” high, secund on the usually few and 
elongated, usually leafy branches of the 
panicle; rays few, small, deep yellow; 
bracts of the involucre oblong-lanceolate, 
obtusish. 

In woods and copses, Nova Scotia to Geor- 
gia, west to Minnesota, Missouri and Texas. 
July—Sept. 


Solidago microphylla Engelm., ranging from 
Kansas to Texas, appears to be a race of this 
species with acutish inner bracts of the involucre. 


30. Solidago Boottii Hook. Boott’s 
Golden-rod. Fig. 4242. 
S. Boottii Hook. Comp. Bot. Mag. 1: 97. 1835. 


Stem glabrous, or puberulent above, slender, 
usually branched near the summit, 2°-5° high. 
Leaves firm, pinnately and finely reticulate- 
veined, the lower and basal ones ovate or 
oblong-lanceolate, narrowed into margined, 
sometimes ciliate petioles, acuminate at the 
apex, sharply and rather coarsely serrate, 3/6’ 
long, the upper smaller, entire, or finely ser- 
rate, sessile; heads 2’—3” high, secund on the 
elongated, spreading or recurving branches of 
the usually ample panicle; rays few, small; 
bracts of the involucre linear-oblong, obtuse; 
achenes pubescent. 


In dry woods, Virginia to Florida and Texas. 
Recorded from Missouri. Ascends to 3000 ft. in 
Virginia. July—Sept. 


392 COMPOSITAE. Vor. III. 


31. Solidago Ellidttii T. & G. Elliott’s 
Golden-rod. Fig. 4243. 


Solidago Elliottti T. & G. Fl. N. A. 2: 218. 1841. 
Solidago elliptica Ell. Bot. S. C. & Ga. 2: 376. 
1824. Not Ait. 1789. 


Stem glabrous, or minutely puberulent 
above, stout, 3°-6° high, simple, or branched 
at the inflorescence. Leaves firm, oblong or 
oblong-lanceolate, rarely ovate-oblong, acute 
or acuminate, sessile by a broad base, or 
sometimes narrowed below, finely serrate, 
crenate-serrate, or the upper entire, rough 
on the margins, pinnately veined, glabrous on 
both sides, or puberulent on the veins be- 
neath, 17-5’ long, 4’—12” wide; heads about ° 
3” high, more or less secund on the short, 
_ spreading or recurving branches of the nar- 
row panicle; bracts of the involucre linear- 
oblong, obtuse; rays 6-12, short; achenes pu- 
bescent. 

In swamps, Nova Scotia (?), Massachusetts 


to North Carolina and Georgia, mainly near the 
coast. Sept.—Oct. 


32. Solidago neglécta T. & G. Swamp : 
Golden-rod. Fig. 4244. 
Solidago neglecta T. & G. Fl. N. A. 2: 213. 1841. 


Stem glabrous, or slightly rough above, simple, 
rather stout, 2°-4° high. Leaves firm, the basal 
and lower ones lanceolate or oblong-lanceolate, 
large, sometimes 12’ long, acute or acutish, closely 
serrate or serrulate, tapering into margined peti- 
oles, rough on the margins; upper leaves smaller, 
lanceolate, acute, -sessile, serrate or nearly entire; 
heads about 23” high, more or less secund on the 
short branches of the thyrsoid panicle; rays 3-8, 
small; bracts of the involucre thin, linear-oblong, 
obtuse; achenes glabrous, or nearly so. 


Var 
S 


rw 
\h aN 
Yor > 


In swamps and bogs, Maine to Vermont, Michigan, 
Maryland, Illinois and Wisconsin. Recorded north 
to New Brunswick. Forms with the heads little 
secund resemble S. uliginosa. Pyramid-golden-rod. 
Aug.—Sept. 


33. Solidago uniligulata (DC.) Porter. 
Few-rayed Golden-rod. Fig. 4245. 


Bigelovia (?) uniligulata DC. Prodr. 5: 329. 1836. 

Solidago linoides T. & G. Fl. N. A. 2: 216. 1841. 
Not Soland. 

Solidago neglecta var. linoides A. Gray, Syn. Fl. 1: 
Part 2) 154.5 18843 

S. uniligulata Porter, Mem. Torr. Club 5: 320. 1894. 


Stem simple, slender, 13°-23° high, glabrous, or 
slightly pubescent above. Leaves firm, obscurely 
pinnately veined, lanceolate or oblong-lanceolate, 
finely and sharply serrate, acute or acuminate, the 
lower long-petioled, 4-0’ long, 4-9” wide, the 
upper sessile, the uppermost very small and erect; 

/ heads about 2” high, densely secund on the short 
spreading or recurving branches of the small 
naked panicle; rays I-4; bracts of the involucre 
firm, linear-oblong, obtuse; achenes glabrous. 


In bogs and swamps, Newfoundland to New York, 
New Jersey, Ontario and Illinois. Aug.—Sept. 


GENUS 22. THISTLE FAMILY. 393 


34. Solidago juncea Ait. Early or Sharp- 
toothed Golden-rod. Fig. 4246. 


Solidago juncea Ait. Hort. Kéw. 3: 213. 17809. 

S. arguta scabrella T. & G. Fl. N. A. 2: 214. 1841. 

Solidago juncea scabrella A. Gray, Syn. Fl. 2: Part 2, 
155. 1884. 

Solidago juncea ramosa Porter & Britton, Bull. Torr. 
Club 18: 368. 1891. : 
Stem glabrous, or very nearly so throughout, 

rigid, rather stout, simple, or branched at the in- 

florescence, 13°-4° high. Leaves firm, glabrous, 
sometimes rough, lanceolate or oval-lanceolate, 
acute or acuminate, serrate, serrulate, or nearly 
entire, the lower large, sometimes 12’ long and 

2’ wide, long-petioled, the upper smaller, sessile; 

heads 13”-2” high, secund on the recurved or 

sometimes nearly erect branches of the usually 

ample spreading panicle; rays 7-12, small; bracts & 

of the involucre oblong or ovate-oblong, obtuse %& mF 

or acute; achenes glabrous or sparingly pubescent. a Oy 
In dry or rocky soil, New Brunswick to Hudson wn ! 

Bay, Saskatchewan, North Carolina and Missouri. YY 

One of the earliest flowering species. Yellow top. N 

Plume or pyramid-golden-rod. June—Nov. 


35. Solidago arguta Ait. Cut-leaved 
Golden-rod. Fig. 4247. 

Solidago arguta Ait. Hort. Kew. 3: 213. 1789. 

S. Muhlenbergii T. & G. Fl. N. A. 2: 214. 1841. 

S. Vaseyi Heller, Muhlenbergia 1: 7. 1900. 

S. Harrisii Steele, Contr. Nat. Herb. 13: 369. 

IQII. 

Stem simple, rather stout, glabrous, or 
sparingly pubescent above, 2°-4° high. Leaves 
thin, pinnately veined, the lower and basal 
ones broadly oyate or oval, short-acuminate, 
3-16’ long, 1-5’ wide, narrowed into mar- 
gined petioles, or subcordate, sharply and 
coarsely serrate; upper leaves sessile, ovate 
to oblong, acute or acuminate, more or less 
serrate, smaller; heads 23’-3” high, secund 
on the lateral racemose branches of the termi- 
nal, often leafy panicle; rays 5-7, large; 
bracts of the involucre oblong, obtuse; achenes 
glabrous or nearly so. 

In rich woods, Maine to Ontario, Virginia and 


Tennessee. Ascends to 2700 ft. in the Adiron- 
dacks. July—Oct. 


) 


36. Solidago canadénsis L. Canada or Chaat 
Rock Golden-rod. Fig. 4248. 2 Vi ee 
BANK, 7 AN “oe o. 


Solidago canadensis L. Sp. Pl. 878. 1753. 

Solidago rupestris Raf. Ann. Nat. 14. 1820. 

S. canadensis glabrata Porter, Bull. Torr. Club 21: 
310. 18094. 


Stem slender, glabrous, or pubescent above, 
1°-5° high. Leaves thin, triple-nerved, linear- 
lanceolate, 2’-5’ long, 3’—5’’ wide, entire, or ser- 
rate with somewhat appressed teeth, acuminate at 
the apex, narrowed at the base, sessile, or the 
lowest petioled, glabrous, or pubescent; heads 
small, 14” high or less, secund on the spreading 
branches of the often large panicle; rays 4-6, 
short; bracts of the involucre thin, linear, acutish; 
achenes small, nearly glabrous. 


Hillsides, thickets and banks of streams, New- 
foundland to Virginia, Ontario, Saskatchewan, Ten- 
nessee and South Dakota. Aug.—Oct. 


Solidago gilvocanéscens Rydb. differs in being 
canescent; it ranges from Illinois to Manitoba, Sas- 
katchewan, Nebraska and Utah. 


COMPOSITAE: Vor. III. 


37. Solidago serétina Ait. Late Golden- 
rod. Fig. 4249. 
Solidago serotina Ait. Hort. Kew. 2) 210. a7Soe 
S. gigantea Ait. Hort. Kew. 3: 211. 17809. 
me gee Pitcheri Nutt. Journ. Acad. Phil. 7: 1or. 
1834. 


S. serotina gigantea A. Gray, Proc. Am. Acad. 17: 
180. 1882. . 


Stem stout, 3°-8° high, glabrous, sometimes 
glaucous. Leaves lanceolate or oblong-lanceo- 
late, thin, triple-nerved, sharply serrate, or nearly 
entire, sessile, or the lowest petioled, glabrous on 
both sides or pubescent beneath, more or less 
rough-margined, 3’-6’ long, 2’—-15”" wide, acumi- 
nate at the apex, narrowed at the base; heads 
23’-3” high, crowded on the spreading or re- 
/ curving branches of the usually large and often 
eh cans wiied are on puberulent ; 
ly racts of the involucre oblong, thin, obtuse; rays 
\ Wy 7-15, rather large; achenes finely pubescent. 
m0 \ i In moist soil, New Brunswick to British Columbia 
t | \ and Oregon, south to Georgia, Texas and Utah. As- 
a cends to 2300 ft. in Virginia. Aug.—Oct. 


\) 


iy Kd 
we 


38. Solidago glabérrima Martens. Mis- 
souri Golden-rod. Fig. 4250. 


Solidago glaberrima Martens, Bull. Acad. Brux. 8: 68. 
1841, 
S. moritura Steele, Contr. Nat. Herb. 13: 370. 1911. 


Stem glabrous, rather slender, 3°-5° high. 
_Leaves firm or thick, those of the stem linear- 
lanceolate to linear, sessile, acuminate at the apex, 
narrowed at the base, 2-4’ long, rough-margined, 
triple-nerved, entire, or sparingly serrate with 
low sharp teeth, the basal and lowest ones mostly 
larger, spatulate, petioled; heads 2’—3” high, se- 
cund on the spreading or recurving branches of 
the short and broad, usually naked panicle; bracts 
of the involucre oblong, greenish-tipped, obtuse, 
or the inner acute, thick; rays 6-13, short; achenes 
nearly glabrous. 


On dry prairies, Manitoba and Minnesota to Ten- 
nessee, Alberta, Washington, Missouri and Texas. 
Autumn. Referred, in our first edition, to S. mis- 
souriensis Nutt. of the far west. 


we, 
Ke 
\ 


39. Solidago Shortii T. & G. Short’s 
Golden-rod. Fig. 4251. 


Solidago Shortt T. & G. Fl. N. Alo. 2am 


4 


Stem slender, glabrous below, minutely pu- 
bescent above, 2°-4° high. Leaves firm, ob- 
long-lanceolate, sessile (the lowest petioled?), 
triple-nerved, acute or acuminate at the apex, 
glabrous on both sides, the larger 2’-4’ long, 
5-6” wide, sharply serrate with rather small 
and distant teeth, the upper gradually smaller, 
sparingly serrate, or entire; heads about 3” 
high, secund on the usually recurved branches 
of the commonly large puberulent panicle; in- 
volucre narrowly campanulate, its bracts linear- 
oblong, obtuse; rays 5-7, rather small; achenes 
silky-pubescent. 


On rocks at the Falls of the Ohio river. Er- 
roneously recorded from northwestern Arkansas. 
July—Aug. 


GENUS 22. THISTLE FAMILY. 395 


40. Solidago Gattingeri Chapm. Gattin- 
ger’s Golden-rod. Fig. 4252. 


Solidago Gattingeri Chapm.; A. Gray, Syn. Fl. 1: 
Parie2) 50. ood 


Stem slender, 2°-3° high, branched at the in- 
florescence, glabrous throughout. Leaves firm, 
glabrous beneath, rough above, ciliolate, the lower 
and basal ones oblanceolate or spatulate, acutish, 
3-6’ long, 6-10” wide, serrate with low distant 
teeth, narrowed into margined petioles; upper 
leaves abruptly smaller, linear-oblong or oblan- 
ceolate, bract-like, entire, sessile; heads 2’-23” 
high, somewhat secund on the spreading, often 
very slender and elongated branches of the pani- 
cle; bracts of the involucre oblong, very obtuse; 
rays 6-10; achenes puberulent, or glabrous below. 


In dry soil, Tennessee and Missouri. July—Aug. 
A plant similar to this, but with larger serrate stem- 
leaves, occurs in central Nebraska. 


41. Solidago altissima L. Tall, High, or 
Double Golden-rod. Fig. 4253. 


Solidago altissima L. Sp. Pl. 878. 1753. 

S. procera Ait. Hort. Kew. 3: 211. 1789. 

Solidago canadensis procera 1.& G. Fl. N.A.2: 224. 1841. 
Solidago canadensis scabra T.& G. Fl. N. A. 2: 224. 1841. 


Stem stout, pubescent, or hirsute nearly through- 
out, 2°-8° high. Leaves lanceolate, triple-nerved, 
acute at each end, roughish above and pubescent 
beneath, the lower ones sharply serrate and petioled, 
3-6’ long, 4-12” wide, the upper smaller, often en- 
tire, sessile; heads 13”-23” high, usually numerous, 
secund on the spreading or recurving branches of 
the usually large panicle; bracts of the involucre 
linear, obtuse or acutish; rays 9-15; achenes gla- 
brous or somewhat pubescent. 


Usually in dry soil, Maine to Ontario, Nebraska, Geor- 
gia and Texas. Confused, in our first edition, with 5S. 
canadensis L. Yellow weed. Aug.—Nov. 


42. Solidago nemoralis Ait. Gray, Field, or Dwarf Golden-rod. Dyer’s Weed. 
Fig. 4254. 


Solidago nemoralis Ait. Hort. Kew. 3: 213. 1789. 
Solidago nxemoralis arenicola Burgess; Britton & Brown, 
Ill. Fl. 3: 344. 1808. 


Slender, ashy-gray, 6’-2° high, erect, depressed or 
sometimes prostrate, finely and densely pubescent. 
Leaves thick, roughish, the basal and lower ones of 
the stem oblanceolate or spatulate, petioled, obscurely 
triple-nerved, obtuse or acutish, crenate-dentate, 3-6’ 
long, 4’-12” wide; upper leaves gradually smaller, 
oblanceolate or linear-oblong, acute or acutish, en- 
tire; heads 2-3” high, secund on the spreading or 
recurving branches of the terminal, usually one-sided 
panicle; bracts of the involucre linear-oblong; rays 
5-9; achenes pubescent. 


In dry soil, Nova Scotia and Quebec to Saskatchewan, 
Florida, Texas and Arizona. July—Nov. 


Solidago pulchérrima A. Nelson (S. longipetiolata 
Mack. & Bush), of Central North America, ranging east- 
ward into Wisconsin and Missouri, differs mainly by 
somewhat larger heads, and is here regarded as one of 
the many races of S. nemordalis Ait. 


COMPOSITAE. Vor sti 


43. Solidago mollis Bartl. Velvety or Ground 
Golden-rod. Fig. 4255. 


Solidago mollis Bartl. Ind. Sem. Goett. 5. 1836. 
Solidago incana T. & G. Fl. N. A. 2: 221. 1841. 


Solidago nemoralis var. incana A. Gray, Proc. Am, Acad. 

17's 197. L882. 

Stem rigid, stout, low, canescent or slightly scabrous, 
6-12’ high. Leaves pale, canescent or rough, entire or 
dentate, strongly 3-nerved, oblong, ovate or oblanceo- 
late, the lower petioled, 2’-3’ long, 3-12” wide, very 
obtuse, the upper sessile, smaller; heads 2’-3” high, 
somewhat or scarcely secund on the short branches of 
the erect, scarcely one-sided, dense thyrsoid panicle; 
bracts of the involucre oblong, obtuse or acutish; rays 
5-9; achenes pubescent. 


Dry plains, Minnesota to Manitoba, Assiniboia, Kansas, 
Texas and Mexico. July—Sept. 


44. Solidago radula Nutt. Western Rough 
Golden-rod. Fig. 4256. 


Solidago radula Nutt. Journ. Acad, Phila. 7: 102. 1834. 


Stem rough-pubescent, rather slender, 1°-3° high. 
Leaves thick, green, rough on both sides, the lower 
oblanceolate or spatulate, dentate-crenate, obtuse or 
acutish, petioled, 3-8’ long, 6-20” wide, obscurely 
3-nerved, the upper smaller, oblanceolate, entire or 
nearly so, sessile, distinctly 3-nerved, the uppermost 
very small; heads 2-3” high, secund on the short, 
usually somewhat recurved branches of the dense, 
often one-sided panicle; bracts of the involucre ob- 
long or linear-oblong, obtuse; rays 3-7, very short; 
achenes minutely pubescent. 


In dry soil, Illinois to Missouri, Louisiana and Texas. 
Aug.—Sept. 


oN 
= 


45. Solidago Drumméndii T. & G. 
Drummond’s Golden-rod. Fig. 4257. 


Solidago Drummondii T. & G. Fl. N. A. 2: 217. 
1841. 


ifn. Stem rather slender, 1°-3° high, finely soft- 


pubescent. Leaves rather thin, broadly ovate 
or oval, glabrous or nearly so above, finely, 
but sometimes sparingly, pubescent, or rough- 
‘ish beneath, sharply serrate, acute at the apex, 
narrowed at the base, 3-nerved and pinnately 
~ veined, petioled, or the upper sessile, the larger 
3-4’ long, 13’-2’ wide; heads 2’—23” high, 
secund on the usually few, spreading or re- 
curving branches of the panicle; bract-like 
upper leaves obtuse, or acute, entire; rays 4-6, 
conspicuous; bracts of the involucre oblong- 
lanceolate, obtuse; achenes pubescent. 


In rocky soil, Illinois to Louisiana and Mis- 
souri. Sept.—Oct. 


GENUS 22. 


46. Solidago rigida L. Stiff or Hard- 
leaved Golden-rod. Fig. 4258. 

Solidago rigida L. Sp. Pl. 880. 1753. 

Oligoneuron canescens Rydb. Bull. Torr. Club 31: 

652. 1905. 

Stem stout, simple, or branched above, 
densely and finely rough-pubescent, hoary, 1°- 
5° high. ‘Leaves thick, flat, rigid, ovate to 
oblong, pinnately veined, often obtuse, rough 
on both sides, or smoothish beneath, the upper 
sessile, clasping, and rounded or sometimes 
narrowed at the base, I’—2’ long, mostly entire; 
lower and basal leaves long-petioled, some- 
times 1° long and 3’ wide, entire or serrulate; 
heads 4-5” high, many-flowered, in a terminal 
dense, compound corymb, the clusters some- 
times slightly secund; involucre broadly cam- 
panulate, its bracts oblong, obtuse, the outer 
pubescent; rays 6-10, large; achenes glabrous, 
1o-15-nerved. 

In dry sandy, gravelly or rocky soil, Ontario to 
Saskatchewan, Massachusetts, Georgia, Texas and 
Colorado. Aug.—Oct. 

Solidago corymbosa Ell., of the Southern States, 
differing by being iess rough and with slightly 


smaller heads, probably a race of this species, 


48. Solidago Riddéllii 
Golden-rod. Fig. 4260. 


Solidago Riddellii Frank; Riddell, Syn. Fl. West. 
States 57. 1835. 


Stem stout, glabrous, or slightly pubescent . 


above, 1°-3° high. Leaves numerous, thick, 
glabrous on both sides, entire, acute at each 
end, the lower and basal ones long-petioled, 
elongated, lanecolate, somewhat triple-nerved 
and conduplicate, often 1° long, 4-10” wide, 
the upper smaller, similar, sessile and clasping 
at the base, conduplicate, somewhat falcate; 
heads 3’-4” high, 20-30-flowered, very numer- 
ous in a dense corymb; involucre oblong-cam- 
panulate, its bracts broadly oblong, obtuse; 
rays 7-9, narrow; achenes 5-nerved, glabrous. 

On moist prairies, Ontario to Minnesota, Ohio 
and Missouri. Also at Fortress Monroe, Va. 
supposed hybrid with S. rigida L. occurs in Iowa. 
Aug.—Sept. 


THISTLE FAMILY. 


sin. 


Ua 


MES Aig 
a 


t frizgz 


47. Solidago ohioénsis Riddell. Ohio 
Golden-rod. Fig. 4259. 
Solidago ohioensis Riddell, Syn. Fl. West. 


1835. 


Very smooth throughout; stem rather 
slender, simple, 2°-3° high. Leaves firm, 
pinnately veined, flat, the basal and lower 
ones elongated-lanceolate or oblong-lanceo- 
late, obtuse, long-petioled, serrulate toward 
the end, or entire, often 1° long; upper 
leaves sessile, lanceolate, entire, gradually 
smaller and those of the inflorescence bract- 
like; heads 23-3” high, numerous in a 
terminal compound corymb, 15-25-flowered ; 
rays 6-9, small; bracts of the narrowly cam- 
panulate involucre oblong, very obtuse, gla- 
brous; achenes glabrous, 5-nerved. 


States 57. 


_ In moist soil, western New York and south- 
ern Ontario to Illinois, Michigan and Wiscon- 


Aug.—Sept. 
Ny 5 
; 7 os 


eG, 


COMPOSITAE. Vor; Li 


) 49. Solidago Houghtonii T. & G. 
Houghton’s Golden-rod. Fig. 4261. 


Solidago Houghton T. & G.; A. Gray, Man. 
Zila eA 


Stem slender, glabrous below, sparingly 
pubescent above, 1°-2° high. Leaves linear, 
the basal and lower ones petioled, 4’—5’ 
long, 2-4” wide, 3-nerved, entire, acute at 
each end, the upper smaller, sessile, slightly 
conduplicate, otherwise similar, the upper- 
most small and bract-like; heads about 3” 
high, few, in a small corymbose cyme, 20- 
30-flowered; involucre broadly campanu- 
late, its bracts oblong, obtuse; achenes gla- 
brous, 4-5-nerved. 


In swamps, north shores of Lakes Michigan 
and Huron, and in Genesee Co., N. Y. Re- 
corded from Lake Superior. Autumn. 


2. 


23. EUTHAMIA Nutt. Gen. 2: 162. 1818. 


Erect, paniculately-branched herbs, perennial by long rootstocks, with linear or linear- 
lanceolate, entire, sessile, 1-5-nerved punctate leaves, and very numerous small heads of both 
tubular and radiate yellow flowers; clustered in the large corymbose, convex or nearly flat- 
topped inflorescence. Bracts of the involucre obtuse, imbricated in several series, appressed, 
somewhat glutinous. Receptacle flattish, fimbrillate, or pilose. Ray-flowers pistillate, usually 
more numerous than the disk-flowers, the rays small. Disk-flowers perfect. Anthers obtuse 
at the base. Style-branches with lanceolate appendages. Achenes top-shaped or oblong, 
villous-pubescent. [Greek, referring to the clustered heads. | 

About 10 species, natives of North America. Type species: Euthamia graminifolia (L.) Nutt. 


Leaves distinctly 3-5-ribbed ; heads 20~30-flowered. 
Involucre 2”—21%4” high, the bracts yellowish. : 1. E. graminifolia. 
Involucre less than 2” high, the bracts with appressed green tips. 2. E. floribunda. 
Leaves 1-ribbed, or with a pair of indistinct lateral nerves; heads rarely more than 20-flowered. 
Involucre 21%4”-3” high ; southwestern species. 


Leaves 2”—4” wide; involucre scarcely viscid. 3. E. leptocephala. 
Leaves 17-214” wide; involucre very viscid. 4. E. gymnospermoides. 
Involucre 2” high, or less. ged 
Leaves 1”-214” wide; involucre campanulate. — 5. E. tenuifolia, 
Leaves less than 1” wide; involucre subcylindric. 6. E. minor. 
1. Euthamia graminifolia (L.) Nutt. stig, BON 5 
ZN fab 


Bushy, Fragrant, or Flat-topped Golden- any 
rod. Fig. 4262. = Ay 


Chrysocoma graminifolia L. Sp. Pl. 841. 1753. \ 
Solidago lanceolata L. Mant. 114. 1767. X \ 
S. graminifolia Salisb. Prodr. 109. 1796. ’ 
E. graminifolia Nutt. Gen. 2: 162. 1818. 
E. Nuttallii Greene, Pittonia 5: 73. 1902. 
E. camporum Greene, loc. cit. 74. 1902. 


Stem paniculately much branched, or rarely 
simple, glabrous or roughish-pubescent, 2°-4° 
high. Leaves numerous, linear-lanceolate, 
acuminate or acute at each end, 17-5’ long, 
2’’-4”" wide, 3-5-nerved, minutely rough-pubes- 
cent on the margins and nerves of the lower 
surface; resinous dots few; heads 2”—23” high, 
sessile in capitate clusters arranged in a flat- 
topped compound corymb; involucre ovoid- 
campanulate to subcylindric, its yellowish 
bracts oblong or oblong-lanceolate, slightly 
viscid; rays 12-20; disk-flowers 8-12. 

In moist soil, fields and roadsides, New Bruns- 
wick to Saskatchewan, Alberta, Florida, Nebraska 
and Wyoming. Fragrant. July—Sept. 


GENUS 23. THISTLE FAMILY. 399 


Ma 

SV “y me 

SH ee NV? 
Sw Wee WY Wp 
Ww ZN cos 
SING u)/ Wz 


2. Euthamia floribunda Greene. Small-headed 
Bushy Golden-rod. Fig. 4263. 


WZ We SS 

E. floribunda Greene, Pittonia 5: 74. 1902. N YA y. WA y, N \ Wy 
a XN " &N \ 1 Wi 
Solidago polycephala Fernald, Rhodora 10: 93. 1908. gS KE ANC fs 
Z r AY ff Se 

5 i, . 2 i) Sy JA\/| 
Finely roughish-pubescent, at least above, panicu- \ Y Ne 
lately branched, 2°-3° high. Leaves linear-lanceo- aN 


late, the larger 2’-3’ long, 23’-3’’ wide, 3-nerved, 
those of the branches much smaller, spreading or 
deflexed; heads numerous, small, 13’-2” high, ses- 
sile or very nearly so in small corymbed clusters; 
involucre turbinate, its glutinous bracts puberulent, 
their triangular-lanceolate green tips appressed. 


Fields and borders of marshes, southern New Jersey, 
and recorded from eastern Pennsylvania. Aug—Oct. 


3. Euthamia leptocéphala (T. & G.) Greene. 
Western Bushy Golden-rod. Fig. 4264. 


Solidago leptocephala T. & G. Fl. N. A. 2: 226. 1841. 


Bulhowes leptocephala Greene, Mem. Torr. Club 5: 321. 
1894. 

Stem smooth, 14°-23° high, branched above. Leaves 
linear-lanceolate, acuminate or acute at each end, 
I-nerved, or with a pair of indistinct lateral nerves, 
rough-margined, those of the stem usually 2’-3’ long, 
2”’-4” wide; heads about 3” high, rather narrow, ses- 
sile in the clusters of the flat-topped inflorescence; 
bracts of the subturbinate involucre linear-oblong, 
scarcely viscid; disk-flowers 3 or 4; ray-flowers 7-10. 


In moist soil, Missouri to Louisianaand Texas. Aug.— 
Oct. 


4. Euthamia gymnospermoides 
Greene. Viscid Bushy Golden- 
rod. Fig. 4265. 

Euthamia gymnospermoides Greene, Pittonia 

Sigo. LOO. 


Solidago gymnospermoides Fernald, Rhodora 
10: 93. 1908. 


Usually branched from the base or from 
below the middle, glabrous, resinous, 13°-2° 
high, the branches strict, ascending. Leaves 
narrowly linear, 1-2” wide, 3’ long or less, 
light green, I-nerved, or the larger 3-nerved; 
heads numerous, sessile in the clusters of 
the broad nearly flat-topped inflorescence; 
involucre turbinate, about 24” high, its 
bracts linear-oblong, blunt, very viscid; 
disk-flowers 4-6; ray-flowers about 12. 

Prairies, Nebraska to Kansas, Texas and 


Louisiana. .Aug.—Oct. Confused, in our first 
edition, with the preceding species. 


400 COMPOSITAE. Vor. III. 


5. Euthamia tenuifolia (Pursh) Greene. Slender Fragrant Golden-rod. 
Quobsque-weed. Fig. 4266. 
ra : ?Erigeron carolinianum L. Sp. Pl. 863. 1753. 


Solidago tenuifolia Pursh, Fl. Am. Sept. 540. 
1814. 


E. tenuifolia Greene, Pittonia 5: 77. 1902. 
E. remota Greene, loc. cit. 78. 1902. 


?S. Moseleyi Fernald, Rhodora 10: 93. 1908. 


Glabrous and somewhat resinous, seldom 
over Iz° high, branched above. Leaves nar- 
rowly linear, entire, acuminate, sessile, nar- 
rowed at the base, I-nerved or with an 
additional pair of faint lateral nerves, 1-3’ 
long, 1-2” wide, punctate, often with 
smaller ones clustered in the axils, the resi- 
nous dots minute; heads about 13” high, 
very numerous and crowded in the dense 
nearly flat corymb; involucre oblong-cam- 
panulate, its bracts oblong; rays 6-12; disk- 
flowers 4-6. 


\\i 


Ys 
mt: iit olay 
J Ware 


In dry sandy soil, eastern Massachusetts to 

Illinois, Wisconsin, Florida and Louisiana. Re- 

/ ferred, in our first edition, to Euthamia caro- 

y 1 liniana (L.) Greene, but the identity of 

: by a Eur oe carolinianum L. is doubtful. Aug.— 
ct. 


6. Euthamia minor (Michx.) Greene. 
Narrow-leaved Bushy Golden-rod. 
Fig. 4267. 


Solidago lanceolata minor Michx. Fl. Bor. Am. 2: 
TIO. §LS03. 


Euthamia minor Greene, Pittonia 5: 78. 1902. 
Solidago minor Fernald, Rhodora 10: 93. 1908. 


Glabrous, bushy-branched above, 3° high or 
less. Leaves very narrowly linear, I-nerved, the 
larger about 23’ long, not over 1” wide, often 
with tufts of smaller ones in the axils, the upper 
much smaller, often not more than #4’ wide; 
heads very numerous, short-stalked, or sessile; 
involucre cylindraceous, about 2” high, its yel- 
lowish oblong bracts appressed, viscid; ray-flow- 
ers about Io. 


In dry sandy soil, Virginia to Florida and Missis- 
sippi. Sept.—Oct. 


24. BRACHYCHAETA T. & G. Fl. N. A. 2: 194. 1841. 


An erect, perennial herb, with the aspect of a golden-rod. Leaves alternate, the lower 
and basal ones large, cordate, long-petioled, the upper ovate, short-petioled or sessile. Heads 
composed of both tubular and radiate flowers, sessile, in a terminal narrow spike-like thyrsus. 
Involucre narrowly campanulate, its bracts coriaceous, imbricated in few series, the outer 
successively smaller. Receptacle small, naked. Rays small, yellow, pistillate. Disk-flowers 
perfect, their corollas tubular, somewhat expanded above, 5-cleft. Anthers obtuse and entire 
at the base. Style-branches flattened, their appendages lanceolate. Achenes 8~10-ribbed. 
Pappus a single row of scale-like bristles, shorter than the achene. [Greek, short-bristle, 
referring to the pappus.] 


A monotypic genus of eastern North America. 


GENUS 24. THISTLE FAMILY. 401 


1. Brachychaeta sphacelata (Raf.) Brit- 
ton. False Golden-rod. Fig. 4268. 


Solidago sphacelata Raf. Ann. Nat. 14. 1820. Zi 
S. cordata Short, Trans. Journ. Med. 7: 599. 1834. < 
Brachychaeta cordata T. & G. Fl. N. A.2: 194. 1841. 

B. sphacela.a Britton; Kearney, Bull. Torr. Club 20: 

484. 1893. 

Stem erect, pubescent, simple or branched above, 
2°-4° high. Basal and lower leaves broadly ovate, 
cordate or truncate at the base, acute at the apex, 
pinnately veined, sharply serrate, 3-6’ long, the 
slender petioles 3-9’ long, stem leaves gradually 
smaller and shorter-petioled, the uppermost very 
small and sessile; heads about 23” high, racemose- 
secund or densely clustered on the short branches 
of the narrow elongated terminal thyrsus; bracts 
of the involucre oblong or linear-oblong, obtuse 
or acutish; rays and disk-flowers each about 5. 

In dry woods, Virginia to Indiana, western Ken- 
tucky, North Carolina, Georgia and Alabama. Aug.— 


Sept. H Wi 
25. APHANOSTEPHUS DC. Prodr. 5: 310. 1836 


Erect or ascending canescent branching herbs, with alternate leaves, and rather large 
heads of both tubular and radiate flowers, solitary at the ends of the branches. Involucre 
hemispheric, its bracts lanceolate or linear, scarious-margined, imbricated in a few series, the 
outer smaller. Receptacle convex or conic, naked. Ray-flowers pistillate, white or purplish. 
Disk-flowers perfect, yellow, their corollas tubular, the limb expanded above, 5-dentate. 
Anthers obtuse and entire at the base. Style-branches flattened, their appendages short, 
obtuse. Achenes many-ribbed. Pappus a short dentate crown. [Greek, faint-crown.] 


About 5 species, natives of the southwestern United States and Mexico. Type species: 
Aphanostephus ramosissimus DC. 


1. Aphanostephus skirrébasis (DC.) Tre- 
\ 


We ZN We lease. Aphanostephus. Fig. 4269. 
ge 


WH 
7 


Keerlia skirrobasis DC. Prodr. 5: 310. 1836. 
Leucopsidium arkansanum DC. Prodr. 6: 43. 1837. 


Aphanostephus arkansanus A. Gray, Pl. Wright. 1: 
Q3y Lose: 

Aphanostephus skirrobasis Trelease; Coville & Bran- 
ner, Rep. Geol. Sury. Ark. 1884: Part 4, 191. 1891. 
Erect, or diffusely branched, densely canescent, 

6’-2° high. Basal and lower leaves spatulate, ob- 

tuse, I—4’ long, somewhat dentate, laciniate or 
entire, narrowed into margined petioles; upper 
leaves lanceolate, oblong or oblanceolate, obtuse 

or acute, mostly sessile, smaller; heads 8-12” 

broad, 3-5” high; rays numerous, narrow, entire; 

achenes ribbed and angled; pappus a lobed or 
dentate crown. 


In dry soil, Kansas to Texas and Chihuahua, east 
to Florida. May—Aug. 


26. BELLIS [Tourn.] L. Sp. Pl. 886. 1753. 


Tufted herbs, with branching or scapose stems, alternate or basal leaves, and rather large 
heads of both tubular and radiate flowers, solitary at the ends of the branches, or of the 
monocephalous scape. Involucre hemispheric or broadly campanulate, its bracts herbaceous, 
imbricated in I or 2 series, nearly equal. Receptacle convex or conic, naked. Ray-flowers 
white or pink, pistillate. Disk-flowers yellow, perfect, their corollas tubular, the limb 4-5-toothed. 
Anthers obtuse and entire at the base. Style-branches flattened, their appendages short, tri- 
angular. Achenes flattened, obovate, nerved near the margins. Pappus none, or a ring of 
minute bristles. [Latin, pretty.] 

About 9 species, natives of the northern hemisphere. Only the following are known to occur 
in the United States, but 2 others are found in Mexico. Type species: Bellis perennis L. 


Stem branched, 6’—15’ high; involucral bracts acute. 1. B.integrifolia. 
Scapes monocephalous, 1’-7’ high; involucral bracts obtuse. 2. B. perennis. 


26 


COMPOSITAE. Vou iim 


1. Bellis integrifolia Michx. Western Daisy. 
Fig. 4270. 
Bellis integrifolia Michx. Fl. Bor. Am. 2: 131. 1803. 


Slender, diffusely branched, pubescent, 6-15’ high. Leaves 
thin, entire, obtuse, the lower and basal ones spatulate, 1-3’ 
long, narrowed into margined petioles, the upper smaller, 
oblong, oblanceolate or linear; heads 6-15” broad; bracts 
of the involucre acute or acuminate, scarious-margined, 
glabrous or nearly so; rays usually violet, oblong-linear ; 
peduncles terminating the branches, 2-7’ long. 


In moist soil, Kentucky and Tennessee to Arkansas and 
Texas. May-July. 


2. Bellis perénnis L. European or Garden 
Daisy. Marguerite. Fig. 4271. 
Bellis perennis L. Sp. Pl. 886. 1753. 


Perennial, tufted. Leaves all basal, obovate, ob- 
tuse, slightly dentate, 1-2’ long, narrowed into mar- 
gined petioles, pubescent and ciliate; scapes naked, 
1-7’ high, usually several from the same root, pu- 
bescent; heads 6’—12” broad; rays numerous, linear, 
white, pink, or purple; bracts of the involucre ob- 
long, obtuse, usually purple. 


In waste places, or occasionally spontaneous on lawns, 
southern New York and eastern Pennsylvania to Nova 
Scotia and Ontario. Fugitive from Europe. Native also 
of Asia. Naturalized in California and British Colum- 
bia. Herb-margaret. Ewe-ormay-gowan. Childing-daisy. 
Bone- or bruise-wort. Bone-flower. Hen-and-chickens. 
Ban-wort. Bennert. March daisy. Bairn-wort. April—Nov. 


27. TOWNSENDIA Hook. Fl. Bor. Am. 2: 16. 1834. 


Tufted scapose or branching herbs, with alternate, entire, linear or spatulate leaves, and 
large heads of both tubular and radiate flowers. Involucre hemispheric or broadly campanu- 
late; bracts imbricated in several series, the outer shorter. Receptacle nearly flat, naked or 
fimbrillate. Ray-flowers pink or white, pistillate. Disk-flowers tubular, mostly perfect, their 
corollas regular, 5-lobed. Anthers obtuse and entire at the base. Style-branches flattened, 
their appendages lanceolate. Achenes of the disk-flowers compressed, those of the rays com- 
monly 3-angled. Pappus a single series of rigid bristles or short scales. [Named for David 


Townsend, botanist, of Philadelphia. | 


About 25 species, natives of western North Amerjca. Type species: Townsendia sericea Hook. 


Branching from the base; heads terminal. 1. T. grandiflora. 
Acaulescent, or nearly so; heads sessile among the leaves. 2. T. exscapa. 


4 


1. Townsendia grandifldra Nutt. Large- 
flowered Townsendia. Fig. 4272. 


Townsendia grandiflora Nutt. Trans. Am. Phil. Soe. (II) 
7: 306. 1841. 


Perennial from a long woody root, branching at 
the base and sometimes also above, pubescent, or 
at length glabrate, 2’-8’ high. Leaves linear or linear- 
spatulate, 1-3’ long, 13’-3” wide, canescent heads 
1’-13’ broad, solitary at the ends of the branches; in- 
volucre hemispheric, its bracts scarious-margined, 
lanceolate, conspicuously acuminate; rays violet or 
purple; pappus of the ray-flowers a crown of short 
scales, that of the disk-flowers of rigid bristles longer 


hairs. 


In dry soil, South Dakota to western Nebraska, Wyo- 
ming, Texas and New Mexico. May-Aug. 


than the achene, which is pubescent with 2-toothed 


GENUS 27. THISTLE FAMILY. 403 


2. Townsendia exscapa (Richards) Porter. 
Silky or Low Townsendia. Fig. 4273. 


Aster (?) exscapus Richards. App. Frank. Journ.32. 1823. 
a a sericea Hook. Fl. Bor. Am. 2: 16. pl. r19. 

I . 

Townsendia exscapa Porter, Mem. Torr. Club 5: 321. 

1594. 

Acaulescent, or nearly so, from a deep woody root, 
tufted, 1-3’ high. Leaves all clustered at the base, 
narrowly linear or slightly spatulate, 1-2’ long, 1’—23” 
wide; heads closely sessile, 1-12’ broad, equalled or 
surpassed by the leaves; involucre broadly campanu- 
late, its bracts lanceolate, acute, the inner scarious- 
margined; rays white or purplish; pappus of both 
ray- and disk-flowers a row of rigid bristles, those of 
the rays shorter and smaller. 

In dry soil, Saskatchewan to Montana, Nebraska, 
ea and New Mexico. Often silky-pubescent. April— 

uly. 

Townsendia intermedia Rydb., ranging from Ne- 
braska and Colorado to Texas, appears to be a race of 
this species with leaves wider than in the type. 


28. CHAETOPAPPA DC. Prodr. 530%. Tage: 


Annual slender much branched herbs, with small alternate entire leaves, and small long- 
peduncled heads of both tubular and white or purple radiate flowers. Involucre narrowly 
campanulate, its bracts lanceolate, acute or acuminate, imbricated in few series, the outer 
slightly shorter. Receptacle small, naked. Ray-flowers in 1 row, pistillate. Disk-flowers 
perfect, or the central ones staminate, their corollas 5-lobed. Anthers obtuse and entire at 
the base. Style-branches narrow, flattened, their appendages short, obtuse. Achenes nearly 
terete, fusiform, or linear, 5-ribbed. Pappus usually of 5 rigid awn-like scabrous bristles, 
alternating with as many short scales or more.. [Greek, bristle-pappus. ] 


Two known species, natives of the southwestern United States, the following typical. The genus 
Distasis DC. (Chaetopappa modesta A. Gray) is here regarded as distinct. 


se 1. Chaetopappa asteroides DC. Chaetopappa. 
Mi Fig. 4274. 


Chaetopappa asteroides DC. Prodr. 5: 301. 1836. 


Pubescent, erect, 2-12’ high, the branches filiform. 
Lower and basal leaves spatulate, obtuse, 4-1’ long, 
petioled, the upper linear, much smaller and bract- 
like, sessile; heads about 3” broad; involucre about 
2” high, its bracts scarious-margined, glabrous or 
nearly so; rays 5-I2, raised on slender tubes; cen- 
tral sterile flowers usually awnless; shorter scales 
of the pappus of the fertile flowers hyaline, some- 
times lacerate. 


In dry soil, Missouri to Texas and northern Mexico. 
April—July. 


29. BOLTONIA L’Her. Sert. Angl. 16. 1788. 


Perennial glabrous branching herbs, with striate or angled stems, alternate, entire, sessile 
or clasping leaves, and numerous rather large heads of both tubular and radiate flowers, 
paniculate, or solitary at the ends of the branches. Involucre hemispheric or broadly cam- 
panulate, its bracts scarious-margined, imbricated in few series, the outer slightly shorter. 
Receptacle convex or conic, foveolate. Ray-flowers pistillate. Disk-flowers perfect, their 
corollas elongated, 5-lobed. Anthers obtuse and entire at the base. Style-branches flattened, 
their appendages short, lanceolate. Achenes flattened, obovate, their margins thickened or 
narrowly winged, glabrous or nearly so. Pappus a series of short scales, usually with 2- 
slender rigid bristles. [Named for James Bolton, an English botanist of the 18th century.] 


As here limited, the genus consists of the 4 following species, with perhaps a fifth in Oregon. 
Type species: Boltonia glastifoha (Hill) L’Her. 


404 COMPOSITAE. Vou. IIL. 


Disk about 2” broad; leaves linear. 1. B. diffusa. 
Disk 3”—6” broad; leaves lanceolate to oblanceolate. 
Leaves narrowed at the base, sessile, not decurrent on the stem. 


Involucre-bracts lanceolate, acute. 2. B. asteroides. 
Involucre-bracts spatulate, obtuse, or mucronate. 3. B.latisquama. 
Stem leaves, and sometimes those of the branches decurrent, sagittate. 4. B. decurrens. 


I. Boltonia diffusa Ell. Panicled Boltonia. 
Fig. 4275. 


Boltonia diffusa Ell. Bot. S. C. & Ga. 2: 400. 1824. 


Paniculately much branched, 2°-7° high, the branches 
very slender or filiform. Leaves linear, or the lower 
linear-lanceolate, acutish, the larger 1-2’ long, 13-2” 
wide, those of the branches very small and subulate; 
heads about 2” high; disk about 2” broad; rays usually 
white, 1-2” long; involucre broadly campanulate, its 
bracts oblong or oblong-lanceolate, acutish or obtuse; 
achenes obovate, narrowly winged; pappus of several 
short scales and 2 subulate bristles shorter than the 
achene. 


In dry soil, southern Illinois to Texas, east to South Caro- 
lina and Florida. Aug.—Oct. 


2. Boltonia asteroides (L.) L’Her. Aster- 
like Boltonia. Fig. 4276. 


Matricaria asteroides L. Mant. 116. 1767. 

Matricaria glastifolia Hill, Hort. Kew. 19: pl. 3. 1769. 
Boltonia glastifolia L’Her. Sert. Angl. 16. 1788. 
Boltonia asteroides L’Her. Sert. Angl. 16. 1788. 


Rather stout, 2°-8° high, somewhat cymosely @) 
paniculate. Leaves lanceolate, to oblanceolate, ses- SS 
sile, 2’-5’ long, 3-12” wide, the upper linear-lanceo- 
late, smaller; heads 2”-4” high; disk 3-6” wide; 
rays white, pink or purple, 3’-6” long; involucre 
hemispheric, its bracts lanceolate or oblong-lanceo- 
late, acute or acuminate; pappus of setose scales, 
with or without 2-4 slender bristles nearly as long 
as the obovate or oval achene. 


In moist soil, Connecticut to Florida, west to Minne- 
sota, Nebraska and Louisiana. July—Sept. 


3. Boltonia latisquama A. Gray. Broad- 
scaled Boltonia. Fig. 4277. 


Boltonia latisquama A. Gray, Am. Journ. Sci. (II) 
23) 238) 1S62: 


Similar to the preceding species and perhaps a 
race of it. Leaves lanceolate, acute, sessile; heads 
rather larger; rays violet-blue; bracts of the in- 
volucre oblong-spatulate, obtuse or mucronate; 
pappus of numerous small short broad scales and 
2 long bristles. 


Western Missouri, Kansas, Arkansas and Okla- 
homa. Autumn. 


GENUS 20. THISTLE FAMILY. 495 


4. Boltonia dectrrens (T. & G.) Wood. 
Clasping-leaved Boltonia. Fig. 4278. 


Bolionia glastifolia var. (?) decurrens T. & G. Fl. N. 
Ae 2) oo noAms 


Boltonia decurrens Wood, Bot. & Flor. 166. 1870. 


Boltonia asteroides var. decurrens Engelm.; A. Gray, 
Syn. Fl. 1: Part 2, 166. 1884. 


Stout, 3°-6° high, branched above. Leaves oblong- 
lanceolate or elongated-lanceolate, mucronate at 
the apex, those of the stem decurrent and sagittate 
at the base, 3-6’ long, 6-8” wide, those of the 
branches smaller and merely sessile or some of them 
also decurrent; heads 23’—3” high; involucre hemi- 
spheric; rays about 3” long, violet or purple; pap- 
pus of several or numerous short scales and 2 very 
slender bristles. 


In wet prairies, Illinois and Missouri. Aug.—Sept. 
Perhaps a race of B. asteroides. 


30. SERICOCARPUS Nees, Gen. & Sp. Ast. 148. 1833. 


Erect perennial herbs, with alternate leaves, and middle-sized heads of both tubular and 
radiate flowers, in terminal cymose panicles. Involucre ovoid, oblong, or campanulate, its 
bracts coriaceous, with herbaceous or squarrose tips, imbricated in several series, the outer 
shorter. Receptacle small, foveolate. Ray-flowers white, pistillate. Disk-flowers mostly 
perfect, their corollas tubular, narrow, yellowish or purplish, 5-lobed. Anthers obtuse and 
entire at the base. Style-branches with lanceolate-subulate appendages. Achenes slightly 
_ compressed, linear-oblong, I-nerved on each side, pubescent. Pappus of numerous capillary 
scabrous bristles, the outer usually shorter. [Greek, silky fruit, referring to the achenes. | 


Four known species, natives of North America. Besides the following, another occurs on the 
northwestern coast of North America. Type species: Sericocarpus solidagineus (Michx.) Nees. 


Leaves entire, linear, spatulate, or obovate, rigid. 


Glabrous, or nearly so; leaves linear or linear-spatulate. 1. S. linifolius. 
Puberulent or scabrous ; leaves obovate. 2. S. bifoliatus. 
Leaves dentate, oblong, or obovate, thin. 3. S. asteroides. 


1. Sericocarpus linifolius (L.) B.S.P. Narrow-leaved White-topped Aster. 
Fig. 42709. 


Conyza linifolia L. Sp. Pl. 861. 1753. 
Aster solidagineus Michx. Fl. Bor. Am. 2: 108, 


aN 1803. 

} Sericocarpus solidagineus Nees, Gen. & Sp. Ast. 
149. 1832. 

Fer Ceara Ms linifolius B.S.P. Prel. Cat. N. Y. 26. 
I ° 


Glabrous or very nearly so throughout; stem 
rather slender, striate, 1°-23° high. Leaves 
linear or linear-spatulate, spreading, faintly 
3-nerved, thick, entire, obtuse at the apex, nar- 
rowed at the base, 1-2’ long, 13’’-3” wide, ses- 
sile, or the lowest on short margined petioles, 
their margins scabrous; heads about 3’-4” 
high, clustered in 2’s-6’s at the ends of the 
cymose branches; involucre oblong-campanu- 
late, 2-33” long, its bracts oblong, obtuse, the 
outer with somewhat spreading or reflexed 
green tips, the inner scarious and often lacer- 
ate or ciliate at the apex; rays 4 or 5, about 
5” long; pappus white. 


In dry, usually sandy soil, Maine to Ohio, Geor- 
gia and Louisiana. Erroneously recorded from 
Canada. June—Sept. 


_ 406 COMPOSITAE. Vor. III. 


2. Sericocarpus bifoliatus ( Walt.) Porter. 
Rough White-topped or Silk-seed- 
Aster. Fig. 4280. 


Conyza bifoliata Walt. Fl. Car. 204. 1788. 

Aster tortifolius Michx, Fl. Bor. Am. 2: 109. 1803. 
S. tortifolius Nees, Gen. & Sp. Ast. 151. 1832. 

S. bifoliatus Porter, Mem. Torr. Club 5: 322. 1894. 


Densely puberulent or scabrous, about 2° high, 
the stem terete, or slightly angled. Leaves ses- 
sile, obscurely veined, obovate or broadly spatu- 
late, thick, obtuse, ascending or erect by a twist 
at the base, 2’-1’ long, 4-8” wide, entire, mu- 
cronulate, densely appressed-puberulent on both 
sides, the upper mostly oblong, much smaller; 
heads 4”-5” high, cymose-paniculate; involucre 
narrowly campanulate, its bracts oblong or the 
inner linear, pubescent, their tips somewhat 
spreading; rays short; pappus white. 


S LYE 
: AN ie a; a 
R <i J ; 
In dry soil, Virginia to Florida and Louisiana, &Q NES \ ; gO py 
mainly near the coast. Rattlesnake-master. July—Sept. SQ Ay Vv, \\) vm ah 


3. Sericocarpus asteroides (L.) B.S.P. 
Toothed White-topped Aster. Fig 4281. 


Conyza asteroides L. Sp. Pl. 861. 1753. 
S. conyzoides Nees, Gen. & Sp. Ast. 150. 1832. 
S. asteroides B.S.P. Prel. Cat. N. Y. 26. 1888. 
Stem pubescent, or glabrate, slightly angled, 
1°-2° high. Leaves thin, pubescent or glabrous, 
ciliate, pinnately veined and faintly 3-nerved, the 
basal and lower ones obovate or spatulate, den- 
tate or rarely entire, 2’-4’ long, 1-14’ wide, nar- 
rowed into margined petioles; upper leaves ob- 
long or oblong-lanceolate, entire or dentate, acute 
or obtuse, smaller; heads 5-6” high, densely clus- 
tered; involucre campanulate, its bracts oblong, 
ciliate or pubescent, the outer with green reflexed 
tips; pappus brown or white. 
In dry woods, Maine to Florida, west to Ohio, 
Kentucky and Alabama. July—Sept. 


31. ASTER L. Sp. Pl. 872. 1753.* 


Perennial or rarely annual, mostly branching herbs, with alternate leaves, and corymbose 
or paniculate (rarely racemose or solitary) heads of both tubular and radiate flowers. Invo- 
lucre hemispheric, campanulate or turbinate, its bracts various, imbricated in several series, 
the exterior ones usually smaller and shorter. Receptacle flat or convex, generally foveolate. 
Ray-flowers white, pink, purple, blue, or violet (very’ rarely yellowish), pistillate. Disk- 
flowers tubular, perfect, their corollas 5-lobed, usually yellow and changing to red, brown, 
or purple. Anthers obtuse and entire at the base. Style-branches flattened, their appendages 
subulate, lanceolate or ovate, acute. Pappus-bristles slender, numerous, scabrous or denticu- 
late, usually in 1 series, sometimes in 2 series. Achenes mostly flattened and nerved. [ Greek, 
star. ] 

A genus of not less than 250 species, most abundant in North America, where, in addition to 
the following, many others occur beyond our limits. A large number of the species apparently con- 
sist of many slightly differing races, and hybridism is also suspected. Known as Asters or Star- 
worts. Type species: Aster amellus L. 

A. Basal and lower leaves, or some of them, cordate and slender-petioled. (See No. 50.) 

1. None of the stem leaves cordate-clasping. 
* Rays white, violet, or rose. 
§ Rays white, or rarely rose, usually 2-toothed ; plants not glandular. 
+ Involucre ovoid, campanulate or turbinate, its bracts mostly obtuse or rounded; basal leaves few 
and small, or commonly none (except in No. 5). 
(a) Leaves membranous or thin, smooth, or nearly so. 

Heads short-peduncled, 9” broad or less, the disk turning crimson ; leaves acute or short-acuminate. 

1. A. carmesinus. 


* Text prepared for our first edition with the assistance of Prof. Epwarp S. Burcess, 
here somewhat revised. 


GENUS 31. THISTLE FAMILY. 407 


Heads long-peduncled, 9” broad or more, the disk turning brown or reddish brown; leaves long- 
acuminate. 
Heads 1’ broad or more; leaves of the branches large, long, lanceolate, acuminate. 
2. A. tenebrosus. 


Heads 9’—12” broad; leaves of the branches small, obtuse, or acute. 3. A. divaricatus. 
(b) Leaves thick, firm, rough. 

Heads 6”-8” high; inflorescence forking. 4. A. furcatus. 
Heads 4”—5” high ; inflorescence paniculate or glomerate. ; 

Leaves acute, or short-acuminate, pilose beneath ; inflorescence glomerate. 5. A. glomeratus. 

Leaves long-acuminate, not pilose beneath ; inflorescence open-paniculate. 6. A. Claytoni. 

ti Involucre cylindric, its bracts tapering to an obtuse apex; basal leaves large, tufted. 

Bracts of the involucre pale, scarious, usually without herbaceous tips. 7. A. curvescens. 
Bracts of the involucre broader, with herbaceous tips. 8. A. Schreberi. 


§§ Rays violet, usually 3-toothed ; plants glandular. 
7 Predominant glands large, capitate; leaves thick, coarse, heavy. 
Sinus broad ; glands chiefly confined to the inflorescence; plant usually harsh. 9. A. macrophyllus. 
Sinus narrow; glands abundant on the leaves and stem ; growing plant clammy. 10. A. roscidus. 
77 Predominant glands minute, scarcely capitate ; leaves usually thin. 


(a) Inflorescence rather regular, flat, or convex-topped ; plants usually less than 214° tall. 
Sinus broad, shallow. 
Broader leaves orbicular-cordate, their teeth and the inflorescence-leaves inconspicuous. 
11. A. tanthinus. 
Broader leaves reniform, sharply incised ; some inflorescencesleaves conspicuous. 
12. A. violaris. 


Sinus rather deep and narrow ; broader leaves ovate-cordate, sharply serrate. 13. A. multiformas. 
(b) Inflorescence very irregular, paniculate-corymbose; plants often 4°-5° high; broader leaves 
large, cordate, acute. 14. A. nobilis. 


** Rays blue or purple; plants not glandular. 


7 Bracts of the involucre spreading or recurved ; rays 30-45. 15. A. anomalus. 
77 Bracts of the involucre appressed, or erect ; rays 8—20. 
(a) Leaves all entire, or nearly so, thick, or firm. 


Leaves nearly or quite glabrous above. 16. A. Shortit. 
Leaves rough-puberulent on both sides, the upper bract-like. 17. A. azureus. 


(b) Leaves nearly all sharply serrate, thin. 
Heads 2”-3” high, numerous; bracts obtuse or obtusish. 


Leaves rough ; petioles not wing-margined ; bracts appressed. 18. A. cordifolius. 

Leaves smooth, or nearly so; petioles, or some of them, wing-margined. 19. A. Lowrieanus. 
Heads 4”-5” high, usually few; bracts acute or acuminate. 20. A. Lindleyanus. 
Heads 3”—5” high, numerous; bracts acute or acuminate. 

Stem densely and finely pubescent. 21. A. Drummondit. 

Stem glabrous or nearly so; bract-tips spreading. 22. A. sagittifolius. 


2. Stem leaves, vr some of them, cordate-clasping ; plant rough when dry. 
23. A. undulatus. 


B. No cordate and petioled leaves; those of the stem, or some of them, with more or less cordate 
or auricled clasping bases (only slightly auricled in A. tardiflorus, and sometimes in A. /aevis). 
1. Stem rough, or hirsute-pubescent. 

* Leaves entire, oblong, linear, or lanceolate. 

§ Heads 12’ broad; leaves sessile, strongly cordate-clasping. 

7 Stem rough ; leaves oblong to lanceolate ; involucre turbinate. 


Leaves thick, firm, very rough, oblong to oval. 24. A. patens. 
Leaves thin, roughish, oblong-lanceolate. 25. A. phlogifolius. 
Tf? Stem hirsute; leaves lanceolate; involucre hemispheric; bracts viscid. 
26. A. novae-angliae. 
§§ Heads %’-1’ broad; leaves but slightly clasping. 


Involucre hemispheric, its bracts glandular. 27. A. oblongifolius. 
Involucral bracts hispid or ciliate. . 

Leaves lanceolate to oblong-lanceolate, 4”—6” wide. 28. A. nebraskensis. 

Leaves linear to linear-lanceolate, 2”-3” wide. 29. A. amethystinus. 

** Leaves, at least the lower, serrate. 
Stem usually pilose ; bracts very glandular. 30. A. modestus. 
Stem hispid-pubescent ; bracts glabrous, or ciliate. 31. A. puniceus. 
2. Stem glabrous, or only sparingly pubescent above. 
* Leaves sharply serrate. 

§ Leaves tapering to the base. 
Leaves narrowed to the base, the lower into winged petioles. 32. A. tardiflorus. 
Leaves scarcely or gradually narrowed to the base. 31. A. puniceus.. 


§§ Leaves abruptly contracted into margined petioles, often enlarged near the base. 
33. A. prenanthoides. 
§§§ Leaves usually strongly cordate-clasping ; bracts green-tipped. 34. A. laevis. 
** Leaves entire, or very nearly so. 
§ Involucre campanulate, its bracts appressed, green-tipped. . 
t Bracts of the involucre with rhomboid green tips. 

Stem leaves oblong, lanceolate, or oval-lanceolate. 34. A. laevis. 
Stem leaves elongated-lanceolate. 35. A. concinnus. 
tt Bracts of the involucre linear, the tips narrower, lanceolate; stem leaves linear or narrowly 
lanceolate. 36. A. purpuratus. 

§§ Involucre hemispheric. 
+ Bracts of the involucre narrow, not foliaceous. 


408 COMPOSITAE. Vor. III. 


(a) Bracts in several series, unequal. 


Bracts linear-subulate ; leaves narrowly linear. 37. A. junceus. 
Bracts lanceolate, linear, or spatulate; leaves lanceolate to linear. 
Western dry soil plant, 1°—2° tall; leaves 1’—3’ long. 38. A. adscendens. 
Eastern swamp plant, 2°-5° tall; leaves 2’—6’ long. 39. A. novi-belgii. 


(b) Bracts in only 1 or2series ; leaves linear to lanceolate. 40. A. longifolius. 
tft Bracts of the involucre green, foliaceous; western. 
Leaves lanceolate, firm ; heads few. 41. A. foliaceus. 
_ Leaves oblong-lanceolate, thin; heads many. 42. A. phyllodes. 
C. Leaves sessile or petioled, not at all clasping, or scarcely so. 
1. Leaves silky, silvery or canescent on both sides, entire. 
Heads corymbose-paniculate ; bracts oblong ; achenes glabrous. 43. A. sericeus. 
Heads in a narrow raceme; bracts linear; achenes silky. 44. A. concolor. 
2. Leaves neither silvery, silky nor canescent, entire or toothed. 
* Bracts of the involucre with herbaceous tips. 
7 Bracts thin or coriaceous, relatively small. 
+ Leaves bristly-ciliate, linear, rigid; western species. 45. A. Fendleri. 
t£ Leaves not bristly-ciliate. 
© Tips of the involucral bracts spreading (little spreading in A. Radula, erect or spreading in 
A. Hervey) ; heads large and showy ; rays violet to purple. 
Leaves linear to linear-oblong, rigid, obtuse, entire ; heads 2’ broad. 46. A. grandiflorus. 
Leaves lanceolate to oblong, the lower sparingly dentate. 
Basal leaves with margined petioles. 
Involucre hemispheric to campanulate; heads 1’ or more. 


Bracts of the involucre glandular. 47. A. speciabilis. 

Bracts of the involucre ciliate, or glabrous. 48. A. surculosus. 
Involucre turbinate; heads 6’—9” broad. 49. A. gracilis. 
Basal leaves with unmargined petioles. 51. A. Herveyi. 


Leaves oblong-lanceolate, sharply serrate, rugose, the basal usually wanting. 50. A. Radula. 
©° Involucral bracts all appressed (except in A. multiflorus and A. commutatus, small-headed species). 
A Involucre top-shaped ; rays violet. 52. A. turbinellus. 
AA Involucre hemispheric to campanulate; rays mostly white, sometimes purple. 
Heads unilaterally racemose. : 
Stem leaves oval, oblong, or lanceolate, serrate, or chiefly so. 


Stem pubescent or glabrate. ; 53. A. lateriflorus. 
Stem villous ; leaves narrowly lanceolate, thin. 54. A. hirsuticaulis. 
Stem leaves linear-lanceolate to linear, nearly entire; stem glabrate. 55. A. vimineus. 


Heads not unilaterally racemose, mostly paniculate. j 
Involucral bracts spatulate, mostly ciliate, somewhat spreading, at least the outer obtuse; plants 
roughish-puberulent. 


Heads 3”-4” broad, numerous; rays 10-20. 56. A. multiflorus. 

Heads 5”—8” broad, relatively few ; rays 20-30. 57. A. commutatus. 
Involucral bracts appressed, acute. 

Heads solitary at the ends of very small-leaved branchlets. 58. A. dumosus. 


Heads paniculate. 
Stem leaves lanceolate, serrate or entire. 
Heads 8”—10” broad. 
Plants glabrous, or sparingly pubescent above. 

Leaves firm, roughish or rough; rays often purplish; involucral bracts acute. 
59. A. salicifolius. 

Leaves thin, smoothish; rays chiefly white; involucral bracts acuminate. 
60. A. paniculatus. 


Plant puberulent all over. 61. A. missouriensis. 
Heads 6”-8” broad; stem leaves narrowly lanceolate. 62. A. Tradescanti. 
Stem leaves linear-lanceolate to subulate, mostly entire. 
Heads scattered, 6”—9” broad ; upper leaves linear. 63. A. Faxoni. 


Heads numerous, 4”—7” broad; upper leaves subulate. 
Involucre subhemispheric, 21%4”—3” high. 
Rays usually white; heads 4”—7” broad. 


Paniculately branched, bushy. 64. A. ericoides. 
Simple, or with slender ascending branches. 65. A. Pringlei. 
Rays purple; heads 8’—12” broad. 66. A. Priceae. 
Involucre top-shoped, 214” high or less. 
Plant pilose-pubescent, 114°-2° high. 67. A. parviceps. 
Plant glabrous, very slender, 1° high or less. 68. A. depauperatus. 


77 Bracts stiff, relatively large; leaves narrow, rigid, entire. [Genus HELEAStRuM DC.] 
; 69. A. paludosus. 
** Bracts of the involucre without herbaceous tips. 


Bracts linear-subulate, acuminate. 


Leaves firm, 3’ long or less, entire or sparingly serrate. 70. A.nemoralis. 
Leaves thin, 6’ long or less, sharply serrate. 71. A. acuminatus. 
Bracts oblong or oblong-lanceolate, obtuse or acutish; leaves narrow, entire. [Genus UNamia 
Greene. ] 72. A. ptarmicoides. 


D. Leaves fleshy, narrow, entire; plants of salt marshes or saline soil (No. 74 sometimes in 
non-saline situations). 
Perennial ; heads 6”—12” broad; involucral bracts lanceolate, acuminate. 73. A. tenuifolius. 
Annuals; head 3”—5” broad; involucral bracts linear-subulate. [Genus Trrpot1um Nees.] 
Involucre campanulate; disk-flowers more numerous than the rays; rays about 2” long. 
74. A. exilis. 
Involucre cylindraceous ; disk-flowers fewer than the very short rays. 75. A. subulatus. 


GENUS 31. THISTLE FAMILY. 409 


Stems erect, delicate, closely tufted, 1°-2° high, 
glabrous, reddish brown, terete. Leaves all petioled, 
glabrate, very thin, but firm and crisp, the lower and 
basal ones oval, rounded, or with a small deep and 
rounded sinus at the base, bluntly acute or short- 
acuminate at the apex, crenate-serrate, the upper 
ones sometimes ovate-lanceolate, the uppermost short- 
elliptic; petioles slender, the uppermost sometimes 
winged; inflorescence 5’ broad, or less, usually of 
about 5 convex glomerules, each often of 10-15 short- 
peduncled heads, its branches spreading, 3’ long, or 
less; rays chiefly 6, white; disk at first golden yellow, 
finally deep purplish crimson; florets broadly bell- 
shaped; outer bracts obtuse, ciliate, pale, with a green 
tip; achenes glabrous. 


On shaded rocks, near Yonkers, N. Y. Peculiar in its 
~ dense glomerules subtended by large short-elliptic leaves, 
but probably a race of A. divaricatus L. September. 


wl! Y Sy AW 
wg SS Ys 
PN aS 


\ 


2. Aster tenebrosus Burgess. Long-leaved 
Wood Aster. Fig. 4283. 


Stems solitary or scattered, glabrate, striate, 
about 3° high. Leaves very thin and smooth, 
slender-petioled, broadly oblong, coarsely toothed 
with remote acuminate teeth, abruptly long- 
acuminate at the apex, the basal sinus broad, 
rounded, shallow, except in the lowest ones; 
leaves of the inflorescence lanceolate, subentire, 
sessile, sometimes 4’ long; inflorescence broadly 
corymbose, heads about 4” high, often 12’ broad; 
rays usually 9-12; disk pale yellow, becoming 
purplish brown, the florets funnelform with a 
long slender tube; outer bracts chiefly elongated- 
triangular, acute, green, the others linear, obtus- 
ish, the green tip lance-linear; achenes generally 
glabrous. 

In moist dark woodlands, New York to Virginia. 
Peculiar in its large dark leaves with coarser teeth 
than in the next species. Aug.—Sept. 


3. Aster divaricatus L. White Wood sly g XY ANN WZ 
Aster. Fig. 4284. ; NNEC SAR 
aa “~{ PO NGS ee Za \ AM fj) i 
Aster divaricatus L. Sp. Pl. 873. 1753. GSS x () ee) 
Aster corymbosus Ait. Hort. Kew. 3: 207. 1789. = LANG 
Stems tufted, assurgent, flexuous, brittle, IBS 


terete, 12°-23° high, glabrate at maturity. SK 
Leaves thin, smoothish, slender-petioled, ovate- Ca 
lanceolate, dentate with sharp teeth, or the 

small basal ones coarsely serrate, acute to 

acuminate, the basal sinus broad or narrow; 

leaves of the inflorescence small, ovate and W 
acute to orbicular; corymb broad, flattish, re- 

peatedly forked, the slender branches long, 

divergent; heads 9’-12” broad; rays chiefly 

6-9, linear, white; disk turning brown; bracts 

of the involucre broad, ciliate, the rounded 

tip with an inconspicuous green spot. \ 

In open woodlands and thickets, in rather dry — Hh) 
soil, Quebee to Manitoba, Georgia and Tennessee. \ i) 
Here regarded as consisting of many slightly dif- } 
fering races, a number of which have been con- \y 
sidered species and varieties. Sept.—Oct. “ 


Aster viridis Nees, remarkable for its coarse rough basal leaves, and large oval rhomboid rameal 
ones, occasionally from New York and Pennsylvania to Virginia, may prove to be a hybrid between 
the preceding and A. macrophyllus L. 


410 COMPOSITAE. Vor.-IIT. 


4. Aster furcatus Burgess. Forking Aster. 
Fig. 4285. 


Aster furcatus Burgess; Britt. & Brown, Ill. Fl. 3: 358. 
1898. 


Stem leafy, 13° high, or less, loosely forked above. 
Leaves hispid above, hispidulous beneath, firm, sa- 
liently cut-toothed, the lower ovate, short-petioled, 
with a small or shallow sinus, the upper sessile, with 
broad laciniate winged bases, often 5’ long by 23’ 
wide, the uppermost elliptic-oblong, often 3%’ long; 
teeth long and low, sharp; heads few (5-20), slender- 
peduncled; involucre turbinate to campanulate, with 
a truncate or rounded base; rays 3-toothed; disk 
turning brown, the florets funnelform with rather 
broad lobes; pappus long, straight; achenes pubes- 
cent, subangular, not constricted at the summit. 


In woods, especially on shaded cliffs, Indiana, Illinois 
and Missouri. Aug.—Oct. 


= 


EMSS 
POs US = 
a. ‘ ? Z 
PANS Es xs 

(Tr 
yy ; AP 


\ 


5. Aster glomeratus (Nees) Bernh. Bern- 
hardi’s Aster. Fig. 4286. 


Eurybia glomerata Nees, Gen. & Sp. Ast. 139. 1832. 
Aster glomeratus Bernh.; Burgess in Britt. & Brown, 
fil. Fl. 3: 358. 1808. 


Loosely clustered, dull green. Leaves not 
large, mostly short-pilose beneath, thickish, 
rough above; basal leaves present, these and the 
lower stem leaves cordate with a deep, narrow 
sinus, the teeth sharp, rather close and small; 
petioles slender, ciliate; upper leaves much 
smaller, ovate, truncate with a short broadly 
winged base, or the uppermost ovate to lanceo- 
late, sessile, entire; inflorescence compact, of 
many glomerate clusters, round-topped; heads 
about 4” high; bracts pubescent, obtuse, green, 
the inner twice as long as the outer; rays about 
6, cream-white, short, soon deciduous; disk 
turning brown. 

In moist thickets or swamps, especially in ravines, 
Maine to New York and Virginia. July. 


QW, Bh OSS 
f 


“1 


>> 
fit 


a, er: Oey 6. Aster Claytoni Burgess. Clayton’s 
TASS Aster. Fig. 4287. 


Y Aster Claytoni Burgess; Britt. & Brown, Ill. Fl. 3: 
eS 358. 1808. 


— TONS oe AL Pe 

A Similar to A. divaricatus, stems red, tough. 

RE iy Leaves chiefly ovate-lanceolate, not large, rough, 
Keg» thick, slender-petioled, coarsely serrate, pale, or 


AE 
\.S% dull, the apex incurved-acuminate, the upper 
¥ spreading or deflexed, sessile by a broad base, 
lanceolate-triangular, serrulate; inflorescence high, 
Vp with percurrent axis, the long suberect branches 
y Ja each bearing a small umbelliform cluster of heads; 
VL peduncles filiform, as long as the heads, 3’—-4” 
long; bracts pale; rays short, narrow, chiefly 6, 
Ig snow-white; disks at first golden-yellow, finally 
sienna-brown;,. florets about 20, achenes densely 


\ short-hairy. 


In sunny or slightly shaded rocky places, Maine to 
New York and the mountains of Virginia. Sept. 


GENUS 31. THISTLE FAMILY. 411 


7. Aster curvéscens Burgess. Dome-topped 
Aster. Fig. 4288. 


Aster curvescens Burgess; Britt. & Brown, Ill. Fl. 3: 
359. 1808. 


Dark green, chiefly glabrous; rootstocks often 10’ 
long; stem smooth, striate, delicate, 13°-33° high. 
Basal leaves tufted, conspicuous, these and the lowest 
stem leaves with a broad sinus tapering into a 
petiole I-2 times as long as the blade, abruptly in- 
curved-acuminate; middle leaves ovate, short-peti- 
oled, rounded at the base, the upper lanceolate, slen- 
derly acuminate, often falcate; leaves firm, smooth- 
ish, the teeth broad, curved; inflorescence mostly 
convex, 3’—5’ broad, its short filiform naked branches 
widely ascending; heads 4’—-5” high; lower bracts 
short, obtuse, the others longer, nearly uniform, 
scarious, shining, linear, often acute, usually gla- 
brous; rays about 8, cream-white, about 5” long; 
disk becoming purple-brown; pappus early redden- 
ing; achenes slender, glabrous. 


In loose moist shaded soil, New England and New 
York to Virginia. Aug.—Sept. 


8. Aster Schréberi Nees. Schreber’s Aster. 
Fig. 4289. 


Aster Schreberi Nees, Syn. Ast. 16. 1818. 


Stem stout, 2°-3° high, with long internodes. 
Basal leaves often in extensive colonies, thin, dull 
green, firm, rough above, with scattered slender ap- 
pressed bristles, pubescent beneath on the veins, 
reniform-cordate or cordate-triangular, often 7’ long 
by 5° wide, the basal sinus when well developed 
rectangular, 2’ across and 1’ deep; upper leaves 
ovate-oblong to lanceolate, with a short broad basal 
"wing, or sessile; petioles of the lower leaves long, 
conspicuously ciliate when young; inflorescence de- 
compound, flattish, or irregularly convex, 6’-12’ 
broad; heads abiut 5” high; bracts greenish, mostly 
obtuse, ribs and midrib dark green, ciliate; rays 
usually ro. ) 


In borders of woods, and along fence rows in partial 
shade, New York to Michigan and Virginia. July—Aug. 


g. Aster macrophyllus L. Large-leaved 
Aster. Fig. 4290. 


Aster macrophyllus L. Sp. Pl. Ed. 2, 1232. 1763. 


Rough; rootstocks long, thick; stem reddish, 
angular, 2°-3° high. Basal leaves forming large 
colonies, 3 or 4 to each stem, broad, cordate with 
a large irregular sinus, rough above, harsh, thick, 
the teeth broad, curved, pointed, the petioles long, 
narrow; upper stem leaves oblong with short 
broadly winged petioles, the uppermost sessile, 
acute; inflorescence strigose and glandular, broadly 
corymbose, irregular; heads 5’-6” high; peduncles 
rigid, thickish; rays about 16, 5-7” long, chiefly 
lavender, sometimes violet, rarely pale; bracts con- 
spicuously green-tipped, the lower acute, the inner 
oblong, obtuse; disk turning reddish brown; florets 
short-lobed. x 


In moderately dry soil, in shaded places, Canada to 
Minnesota and North Carolina. Here regarded as con- 
sisting of numerous slightly differing races, perhaps in- 
cluding the five following described as speties. Aug. 


412 COMPOSITAE. Vor. LL. 


1o. Aster roscidus Burgess. Dewy-leaf 
Aster. Fig. 4291. 


Aster roscidus Burgess; Britt. & Brown, Ill. Fl. 3: 
360. 1808. e 
Clammy-hairy, odorous, copiously glandular 

when young, somewhat so at maturity; stem 3° 
high, or less. Basal leaves in close colonies, co- 
riaceous, the earlier ones cordate-quadrate, low- 
serrate, the sinus deep, narrow, the later, or 
winter leaves, elliptic, long-petioled, often pros- 
trate, often 5’ long; stem leaves chiefly orbicular 
and not cordate, with short broadly winged 
petioles, rarely slender-petioled; inflorescence 
convex, sometimes irregular; involucre hemi- 
spheric, its bracts chiefly with rounded ciliate 
tips; rays 14-16, broad, clear violet; disks at first 
golden yellow, soon turning red; pappus long, 
white, copious. 


In slight shade and rich cleared woodlands, Maine 
to Pennsylvania and Michigan. Aug.—Sept. 


11. Aster ianthinus Burgess. Violet Wood 
Aster. Fig. 4292. 


Aster ianthinus Burgess; Britt. & Brown, Ill. Fl. 3: 360. 
18908. 


Glandular, dark green, slightly strigose-pubescent ; 
stem erect, or decumbent, 2°-3° tall. Leaves thin- 
nish, rough, the lower and basal ones orbicular to 
oblong, 5’ long, or less, abruptly acuminate, low- 
serrate or crenate; the sinus broad, open, shallow, 
upper leaves sessile by a narrowed base, crenate- 
serrate: inflorescence open, nearly naked, peduncles 
slender, divergent; heads large; rays 10-13, long, 
very deep violet or sometimes pale, 4”-6” long; 
bracts green-tipped, little pubescent. 


On shaded banks and along woodland paths, Maine 
to Lake Erie and West Virginia. July—Oct. 


12. Aster violaris Burgess. Violet-leaf Aster. 
Fig. 4293. 
Het oielarts Burgess; Britt. & Brown, Ill. Fl. 3: 361. 

1898. 

Caudex thick, fleshy; plant glabrate, bluish green, 
minutely glandular; stem slender, erect, or assur- 
gent, 2° high, or less. Basal and lower leaves broadly 
reniform, abruptly acuminate or apiculate, often 3’ 
long by 4’ wide, their slender petioles 6-8’ long, the 
sinus very broad and shallow; middle stem leaves 
similar, not cordate; the upper numerous, long- 
elliptic, chiefly with narrowed bases, all thin, firm, 
rough above; inflorescence leafy, small, loose, rather 
narrow and high, paniculate-corymbose, nearly level- 
topped, its slender branches with nearly opposite, 
oblong leaves; heads 6” high, or more; rays 12-15, 
pale violet, narrow. 

In shaded moist places, sometimes in leaf-mold among 


ped New York from the Hudson to Lake Erie. Sept— - 
ct. 


GENUS 31. THISTLE FAMILY. 


13. Aster multiformis Burgess. Various- 
leaved Aster. . Fig. 4294. 


Aster multiformis Burgess; Britt. & Brown, Ill. FI. 
3: 361. 1808. 


Deep green, minutely glandular; stem erect, 
slender, 1°-2° high, angular-striate in drying. 
Basal leaves usually 2, large, thick, cordate- 
oblong, often accompanied by later smaller ob- 
long ones; stem leaves very thin, sharply ser- 
rate, rough above, minutely puberulent beneath, 
the lower ovate, acuminate, usually with a nar- 
row sinus, the upper oval to ovate-lanceolate, 
petioled, the uppermost elliptic-lanceolate, ser- 
rulate, sessile or nearly so; inflorescence small, 
its branches upwardly curved; heads about 7” 
high; rays about 13, rounded and retuse at the 
apex; bracts green, glands few, almost hidden 
by the minutely strigose pubescence of the pe- 
duncles. 


In moist shaded places, Maine to western New 
York, Pennsylvania and Maryland. July—Aug. 


14. Aster nobilis Burgess. 


413 


Stately Aster 


Fig. 4295. 


Aster nobilis Burgess; Britt. & Brown, Ill. Fl. 3: 361. 
18908. 


Tall, minutely glandular above; stem shining, 
bright green, 4°-5° high. Leaves thin, but firm, 
smooth in growth, roughened in drying, mi- 
nutely puberulent beneath, dark green, basal and 
lower leaves large, the blade often 9’ long by 6’ 
wide, about as long as the stout petiole, sharply 
toothed, the sinus deep, broad, or the lobes over- 
lapping; stem leaves similar, the upper oblong- 
lanceolate, sessile; inflorescence irregularly cy- 
mose-paniculate, with small subulate recurved 
leaves; bracts long, acute, green; heads 6” high, 
or less; rays 13-15, violet-blue or pale violet; 
disk-flowers not numerous, their corollas funnel- 
form with a long capillary tube. 


In leaf-mold, Lake Champlain to Lake Erie. Aug. 


15. Aster andmalus Engelm. Many- 
rayed Aster.. Fig. 4206. 


Aster anomalus Engelm.; T. & G. Fl. N. A. 2: 503. 
1843. 


Stem rough, rather stout, branched above, 
1°-3° high. Leaves thin, the lower and basal 
ones deeply cordate, ovate, or ovate-lanceolate, 
entire or slightly repand, rough-pubescent on 
both surfaces, acute or acuminate at the apex, 
3-4 long, 1-2’ wide, on slender naked peti- 
oles; upper leaves short-petioled or sessile, 
lanceolate, oblong, or linear, much smaller; 
heads few, 4-6” high, 12”-15” broad; recep- 
tacle hemispheric, its bracts lanceolate, acute, 
or acuminate, hirsute, imbricated in several 
series, their foliaceous tips spreading or re- 
flexed; rays 30-45, 5’-6” long, bright violet- 
blue; pappus whitish. 


On limestone cliffs, Illinois to Missouri and 
Arkansas. Sept. 


> 


414 COMPOSITAE. Vor. IIT. 


i 16. Aster Shortii Hook. Short’s Aster. 


SNe ANY YPN Wi? Aster Shortii Hook. Fl. Bor. Am. 2: 9. 1834. 
Ae Stem roughish or smooth, slender, panicu- 
<a lately branched above, 2°-4° high. Leaves 
Pr, thick, glabrous or nearly so above, finely and 
IN sparingly pubescent beneath, the lower and 


basal ones ovate or ovate-lanceolate, cordate 
at the base, acute or acuminate at the apex, 
slightly dentate; or entire, 2’-6’ long, 17-2’ 
wide, borne on slender naked petioles; upper 
leaves lanceolate, entire, sessile or short-peti- 
oled, not cordate, those of the branches small 
and bract-like; heads numerous, 5’-7” high; 
involucre broadly campanulate, its bracts lin- 
ear, acute, puberulent, imbricated in several 
series, their green tips appressed; rays 10-15, 
linear, violet-blue, 5’-6” long; pappus tawny. 
= On banks and along edges of. woods, western 

Pennsylvania to Virginia, Georgia, Illinois, Wis- 
consin and Tennessee. Sept.—Oct. 


17. Aster azureus Lindl. Sky-blue Aster. 
Fig. 4208. 


Aster azureus Lindl.; Hook. Comp. Bot. Mag. 1: 
Qos) eLosi5: 

A. capillaceus Burgess; Small, Fl. S.E. U. S. 1215. 
1903. 


Stem slender, stiff, rough, branched above, 
1°-4° high. Leaves thick, usually all entire, sca- 
brous on both sides, the lower and basal ones 
cordate, ovate, ovate-lanceolate, or lanceolate, 
acute, acuminate, or obtusish, 2’-6’ long, with 
slender, naked, often pubescent petioles; upper 
leaves short-petioled or sessile, lanceolate or 
linear, those of the branches reduced to small 
appressed bracts; heads numerous, 4-5” high; 
involucre turbinate, its bracts glabrous, linear- 
oblong, abruptly acute, imbricated in several 
series, their green tips appressed; rays 10-20, 
bright blue, 3-4” long; pappus tawny. 


On prairies and along borders of woods, Ontario 
and western New York to Georgia, Minnesota, Kan- 
sas, Alabama and Texas. Aug.—Oct. 


18. Aster cordifdlius L. Common 
Blue Wood Aster. Fig. 4299. 


Aster cordifolius L. Sp. Pl. 875. 1753. 
Aster choralis Steele, Contr. Nat. Herb. 10: 
37 2.- LOUD 


Stem glabrous or nearly so, rarely pu- 
bescent, much branched, bushy, 1°-5° high. 
‘\ TS ASE Leaves thin, rough, more or less pubescent 
Za aa with scattered hairs above and on the veins 

7, beneath, sharply serrate, acuminate, the 
lower and basal ones slender-petioled, 
broadly ovate-cordate, 2’-5’ long, the upper 
short-petioled or sessile, ovate or lanceo- 
late, smaller; petioles scarcely margined; 
heads usually very numerous, small, 2-3” 
high, 6-9” broad, handsome; involucre 
turbinate to cylindric, its bracts oblong- 
linear, obtuse or obtusish, green-tipped, ap- 
pressed; rays 10-20, 3-4” long, violet or 
blue, sometimes pale, rarely white. 


Woods and thickets, Nova Scotia and New 
Brunswick to Ontario, Minnesota, Georgia and 
Missouri. Consists of many races, differing 
mainly in leaf-form and inflorescence. Sept.— 
Dec. Tongue. Bee-weed. 


AN 
WP 
NY / 


GENUS 31. THISTLE FAMILY. 415 


1g. Aster Lowrieanus Porter. Lowrie’s \\ 
or Fall Aster. Fig. 4300. 


Aster cordifolius var. laevigatus Porter, Bull. 
Torr. Club 16: 67. 1889. Not A. laevigatus 
Lam. 1783. 

Aster Lowrieanus Porter, Bull. Torr. Club 21: 
121. 1894. 


Glabrous, or very nearly so throughout; 
stem branched, 1°-4° high. Leaves thickish, 
firm, a little succulent, the basal slender-peti- 
oled, ovate to ovate-lanceolate, mostly cordate, 
acute or obtusish, serrate, or sometimes in- 
cised, 2-6’ long, those of the stem ovate to 
oblong, often cordate, contracted into winged 
petioles, the uppermost lanceolate; heads usu- 
ally not very numerous, 23’—-3” high, loosely 
panicled; involucre turbinate, its bracts obtuse 
or obtusish, appressed; rays light blue, 3’—4” 
long, but variable in length. 


In woods, Connecticut and southern New York 
to Pennsylvania, Ontario, North Carolina and 
Kentucky. Races differ in leaf-form and serration. 
Sept-Oct. Bee-weed. Blue-devil. 


20. Aster Lindleyanus T. & G. Lindley’s Aster. Fig. 4301. 


As er Lindleyanus T. & G. Fl. N. A. 2: 122. 1841. 

A, Wilsoni Rydb. Bull. Torr. Club 37: 138. 1910. 

Aster Lindleyanus eximius Burgess; Britt. & Brown, 
Ill. Fl. 3: 364. 1898. 


Stem usually stout, glabrous, or sometimes pu- 
bescent, 1°-6° high, branched above. Leaves 
rather thick, glabrous, or slightly pubescent, espe- 
cially on the veins, the lower and basal ones cor- 
date at the base, sharply serrate, ovate, acute or 
acuminate, 2-4’ long, with slender naked peti- 
oles; upper leaves ovate, ovate-lanceolate, or lan- 
ceolate, less serrate, or entire, sessile, or with 
margined petioles, those of the branches lanceo- 
late or linear-lanceolate, smaller; heads usually 
not numerous, 4-5” high; involucre broadly 
turbinate or nearly hemispheric, its bracts linear- 
lanceolate, acute, rather loosely imbricated, gla- 
brous, or nearly so, their tips green; rays 10-20, 
blue or violet, 3’-5” long; pappus nearly white. 

In open places, Labrador to Mackenzie, Alberta, 


Maine, New York, Michigan and Montana. Aug.— 
Oct. 


21. Aster Drumméndii Lindl. Drummond’s 
Aster, Vig. 4302. 
Aster Drummondii Lindl. in Hook. Comp. Bot. Mag. 1: 


97. 1835. i 
Aster hirtellus Lindl. in DC. Prodr. 5: 233. 1836. 


Stem usually stout, finely and densely canescent, 
branched above, 2°-5° high. Leaves mostly thin, 
ovate or ovate-lanceolate, acuminate, rough above, 
canescent beneath, the lower and basal ones cordate, 
with slender naked petioles, sharply toothed, 2-4’ 
long, the upper cordate or rounded at the base, usu- 
ally on margined petioles, those of the branches 
sessile and entire or nearly so, much smaller; heads 
high, rather numerous on the racemose 
branches; involucre turbinate, its bracts linear, 
slightly pubescent, acute or acuminate, their green 
tips appressed; rays 8-15, blue, 3-4” long; pappus 
whitish. 

In dry soil, borders of woods and on prairies, Ohio 


to Minnesota, Kentucky, Arkansas and Texas. Perhaps 
not specifically distinct from the following. Sept.—Oct. 


416 COMPOSITAE. Vor. IIT. 


22. Aster sagittifolius Willd. Arrow- 
leaved Aster. Fig. 4303. 


Aster sagittifolius Willd. Sp. Pl. 3: 2035. 1804. 


Stem stout, or slender, strict, glabrous, or spar- 
ingly pubescent above, 2°-5° high, paniculately 
branched at the inflorescence, the branches as- 
cending. Leaves thin, slightly roughened, or gla- 
brous above, glabrate or pubescent beneath, the 
lower and basal ones cordate or sagittate, ovate- 
lanceolate, sharply serrate, acuminate, 3-6’ long, 
with slender naked or narrowly margined peti- 
oles; upper leaves lanceolate, sessile, or on short 
and usually margined petioles, serrate or entire, 
those of the branches very small; heads 23’—4” 
high, 8-10” broad, numerous, crowded, race- 
mose; involucre turbinate, its bracts linear-subu- 
late, glabrous or nearly so, their tips green and 
slightly spreading; rays 10-15, light blue or pur- 
plish, 3-4” long; pappus whitish. 

In dry soil, New Brunswick to Ontario, North Da- 
kota, New Jersey, Georgia and Missouri. Aug.—Oct. 

Aster Saundersii Burgess, of the Dakotas and 
Iowa, differs by a corymbose inflorescence. 


23. Aster undulatus L. Wavy-leaf 
Aster. Fig. 4304. 
Aster undulatus L. Sp. Pl. 875. 1753. 

Stem stiff, very rough and pubescent, divari- 
cately branched above, 1°-33° high. Leaves 
usually thick, rough on both sides, pubescent 
beneath, dentate, undulate or entire, acute or 
acuminate, the lowest and basal ones ovate, 
cordate, 2’-6’ long, with naked or margined 
petioles; middle ones ovate, lanceolate or ob- 
long, with margined petioles dilated and clasp- 
ing at the base, the upper sessile or clasping, 
those of the branches small and subulate; 
heads numerous, racemose and often secund 
on the spreading branches, about 4” high, 
8-10” broad; involucre broadly turbinate, its 
bracts linear-oblong, pubescent, acute or acut- 
ish, their green tips appressed; rays 8-15, pale 
blue to violet, 3-5” long; pappus whitish. 

In dry soil, New Brunswick and Ontario to 
Minnesota, Florida, Alabama, Louisiana and 
Arkansas. Various-leaved aster. Races differ in 
leaf-form and inflorescence. Sept.—Oct. 


J~ 
24. Aster patens Ait. Late Purple 
Aster. Purple Daisy. Fig. 4305. 
Aster patens Ait. Hort. Kew. 3: 201. 17809. 
A. patens gracilis Hook, Comp. Bot. Mag. 1: 97. 

1835. 

Stem slender, rough, 1°-3° high, diver- 
gently branched. Leaves ovate-oblong to 
oblong-lanceolate, rough or pubescent, thick 
and somewhat rigid, strongly cordate or 
auriculate-clasping at the broad base, entire, 
acute, or the lowest obtuse, 1-3’ long, those 
of the branches much smallér and bract-like, 
the margins rough-ciliate; heads 1’ broad or 
more, solitary at the ends of the branches; 
involucre broadly turbinate, its bracts linear- 
oblong, finely pubescent or scabrous and 
somewhat glandular, imbricated in several 
series, their green acute tips spreading; rays 
20-30, purplish-blue, or deep violet, 4’-6” 
long; pappus tawny; achenes pubescent. 

In dry, open places, Maine to northern New 
York, Minnesota, Florida, Louisiana and Texas. 
Reported from Canada. Races differ in leaf- 
form and pubescence. Aug.—Oct. 


GENUS 31. THISTLE FAMILY. 


25. Aster phlogifolius Muhl. Thin-leaved 
Purple Aster. Fig. 4306. 


A. phlogifolius Muhl.; Willd. Sp. Pl. 3: 2034. 1804. 


Aster patens var. phlogifolius Nees, Gen. & Sp. Ast. 
49. 1832. 


Similar to the preceding species, usually taller. 
Leaves larger, lanceolate to oblong-lanceolate, 
entire, thin, or membranous, acuminate at the 
apex, strongly auriculate-clasping at the base, 
roughish above, pubescent beneath, usually nar- 
rowed below the middle, sometimes 6’ long; heads 
usually numerous, I’-2’ broad, panicled, or some- 
what racemose on the branches; bracts of the 
involucre lanceolate, glabrate, rather loose, with 
herbaceous tips; rays numerous, purple-blue. 


In woods and thickets, New York to Ohio, North 
Carolina and Tennessee. Perhaps a sylvan race of 
the preceding species. Aug.—Sept. 


Yj 
WY 


rado. 


27. Aster oblongifolius Nutt. Aromatic 
Aster. Fig. 4308. 


Aster oblongifolius Nutt. Gen. 2: 156. 1818. 

Aster oblongifolius var. rigidulus A. Gray, Syn. Fl. 1: 
Part 2, 179. 1884. 

Aster Kumleinii Fries; Rydb. Fl. Colo. 354. 1906. 


Stem much branched, hirsute-pubescent, 1°-23° 
high, the branches divaricate or ascending. Leaves 
crowded, oblong, or oblong-lanceolate, sessile by 
a broad, partly clasping base, usually rigid, en- 
tire, acute or mucronulate at the apex, rough or 
hispidulous on both sides, rough-margined, those 
of the stem 17-2’ long, 2’-4” wide, those of the 
branches gradually smaller; heads corymbose, 
nearly 1’ broad; involucre hemispheric, its bracts 
much imbricated, glandular, aromatic, linear or 
linear-oblong, the acute green tips spreading; rays 
20-30, violet-purple, rarely rose-pink, 3’—5” long; 
pappus light brown; achenes canescent. 

On prairies and bluffs, central Pennsylvania to 
Minnesota, North Dakota, Nebraska, Colorado, Vir- 


ginia, Tennessee and Texas. Races differ in leaf- 
form and pubescence. Plant odorous. Aug.—Oct. 


27 


26. Aster novae-angliae L. New England 


Aster novae-angliae L. Sp. Pl. 875. 
A, roseus Desf. Cat. Hort. Paris, Ed. 3, 4o1. 


Stem hispid pubescent, corymbosely 
branched above, 2°-8° high, very leafy. Leaves 
lanceolate, entire, rather thin, acute, pubescent, 
2’-5’ long, 6’-12” wide, clasping the stem by an 
auriculate or broadly cordate base; heads nu- 
merous, 1-2’ broad, clustered at the ends of the 
branches; involucre hemispheric, its bracts linear- 
subulate, somewhat unequal, green, spreading, 
pubescent and more or less glandular, viscid; 
violet-purple, 
rarely pink or red, or white; achenes pubes- 
cent; pappus reddish-white. 


rays 40-50, 


In fields and along swamps, Quebec to Saskatche- 
wan, South Carolina, Alabama, Kansas and Colo- 
One of the most beautiful of the genus. 
Aug.—Oct. 


418 COMPOSITAE. Vo. III. 


28. Aster nebraskénsis Britton. Nebraska 
Aster. Fig. 4309. 


Aster nebraskensis Britton, in Britt. & Brown, Ill. Fl. 
Bis G75 eLOOe: 


Stem strictly erect, slender, stiff, rough to the 
base, simple, or with a few short nearly erect 
branches, very leafy, 12°-22° tall. Leaves thick, 
rather rigid, ascending, lanceolate to oblong- 
lanceolate, entire, sessile by a subcordate base, 
acute or acuminate at the apex, 1-3’ long, 4-6” 
wide, very rough on both sides, the midvein 
prominent beneath, the lateral veins obscure; 
heads few, terminating short leafy branchlets, 
1-17’ broad; involucre broadly campanulate or 
hemispheric, about 3” high, its bracts green, ob- 
long, acute, imbricated in several series, the outer 
quite foliaceous; rays purple, about 6” long. 


Lake shores, central Nebraska. Sept. Lower and 
basal leaves not seen. 


29. Aster amethystinus Nutt. Amethyst 
Aster, Fig. 4310: 


Aster amethystinus Nutt. Trans. Am. Phil. Soc. (II) 7: 
294. 1841. VEZ 


. ; Xf 5 | : : 
Resembles Aster novae-angliae, but is often taller, Ze Ne 
TOergS 


sometimes 5° high. Leaves often crowded, linear- Sx 
lanceolate, entire, rough or hispidulous on both sides, 
partly clasping, though sometimes slightly so, at the QQ WN¢@ 
sessile base, acute at the apex, those of the stem 7@™ AN 
1-2’ long, 2’-3”” wide; heads rather numerous, race- : 
mose or corymbose, 2’-1’ broad; involucre broadly 
turbinate, its bracts much imbricated, linear, hispid, 
not glandular, the acutish green tips spreading; rays 
20-30, blue or violet, about 3” long; pappus brown; 
achenes canescent. 


I 


ann 
RTOS ena SP 


In moist soil, Vermont and Massachusetts to New ‘ 
York, Pennsylvania, Illinois, lowa and Nebraska. Sept.— 
Oct. Specimens have intermediate characters between 
Aster novae-angliae L. and Aster multiflorus L., and 
hybridism is suspected. 


Wks 
ean ae ayy aan he eee a 


30. Aster modéstus Lindl. Great Northern 
Aster. Fig. 4311. 
Aster unalaschensis var. major Hook. Fl. Bor. Am. 2: 7. 
ee wiieas Lindl. ; Hook. loc. cit. 8. 1834. 
Aster majus Porter, Mem. Torr. Club 5: 325. 1804. 
Stem stout, leafy to the summit, usually densely 
pilose-pubescent with many-celled hairs, rarely gla- 
brate, branched above, 4°-6° high. Leaves mem- 
branous, lanceolate, partly clasping by a narrowed 
base, acuminate at the apex, sharply serrate with 
low, distant teeth, dark green and slightly pubes- 
cent above, villous-pubescent on the veins beneath, 
3-5’ long, 5-10” wide; heads mostly solitary at 
the ends of short branches, 1%’ broad; involucre 
hemispheric, its bracts little imbricated, green, 
linear-subulate, densely glandular; rays 35-45, pur- 
ple to violet, 5’-7”" long; achenes appressed-pubes- 
cent; pappus tawny. 


In moist soil, western Ontario to Minnesota, Oregon 
and British Columbia. Sept.—Oct. 


GENUS 3I. THISTLE FAMILY. 419 


31. Aster puniceus L. Red-stalk or 
Purple-stem Aster. Fig. 4312. 


Aster puniceus L. Sp. Pl. 875. 1753. 


Stem usually stout, reddish, corymbosely or 
racemosely branched above, hispid with rigid 
hairs to glabrous, 3°-8° high. Leaves lanceolate 
to oblong-lanceolate, acuminate, sessile and clasp- 
ing by a broad or narrowed base, sharply serrate, 
or entire, usually very rough above, pubescent on 
the midrib or glabrous beneath, 3’-6’ long, 4’-12’ 
wide; heads generally numerous, 1’-12’ broad; 
involucre nearly hemispheric, its bracts linear or 
oblong, attenuate, imbricated in about 2 series, 
glabrous or ciliate, green, loose, spreading, nearly 
equal, sometimes broadened; rays 20-40, violet- 
purple or pale (rarely white), 5’-7” long, showy; 
pappus nearly white; achenes pubescent” 


a \ i 
YS 


In swamps, Newfoundland to Ontario, Manitoba, 
Minnesota, Georgia, Tennessee, Ohio and Michigan. 
Races differ in pubescence, leaf-form and leaf-serra- 
tion. Early purple aster. Swan-weed. Cocash. Meadow- 
scabish. July—Nov. 


EWAWZ 
Wn 


32. Aster tardiflorus L. Northeastern 
Aster. (hig airs: 


Aster tardiflorus L. Sp. Pl. Ed. 2, 1231. 1763. 
Aster patulus Lam. Encycl. 1: 308. 1783. 


Stem glabrous, slightly pubescent, or villous, co- 
rymbosely branched near the summit, 1°-3° high. 
Leaves lanceolate, oblong-lanceolate, or ovate-lan- 
ceolate, serrate with low teeth, or some of them 
entire, acuminate at the apex, narrowed into a 
slightly clasping base, or the lower into winged 
petioles, glabrous or nearly so on both sides, rough- 
ish-margined, 3’-6’ long, 4”-10” wide; heads about 
1’ broad, not very numerous, involucre hemispheric, 
its bracts often 6” long, acute, somewhat unequal; 
rays 20-30, violet; pappus nearly white; achenes 
pubescent. 


Along streams, New Brunswick to Pennsylvania. 
Aug.—Oct. 


33. Aster prenanthoides Muhl. Crooked- 
stem Aster. Fig. 4314. 
A. prenanthoides Muhl.; Willd. Sp. Pl. 3: 2046. 1804. 


Aster prenanthoides porrectifolius Porter, Mem. Torr. 
Club 5: 326. 1894. 


Stem glabrous, or pubescent in. lines above, 
flexuous, much branched, 1°-2° high. Leaves 
thin, gblong to ovate-lanceolate, or lanceolate, 
sharply and coarsely serrate, scabrous above, gla- 
brous or nearly so beneath, 3’-8’ long, 9’-18” 
wide, acuminate at the apex, abruptly narrowed 
below into a broad margined entire petiole, the 
base auriculate-clasping ; heads usually numerous, 
1’ broad or more; involucre hemispheric, its bracts 
linear, acute, green, spreading, imbricated in 3 or 
4 series, the outer shorter; rays 20-30, violet, 
4-6” long; pappus tawny; achenes pubescent. 


_In moist soil, Massachusetts to Minnesota, Vir- 
ginia, Kentucky and Iowa. Aug.—Oct. 


Aster schistésus Steele, of West Virginia, is in- 5 : ) 
termediate in characters between this species and A. Lowrieanus Porter, and may be a hybrid. 


420 . COMPOSITAE. Vor. IIT. 


34. Aster laévis L. Smooth Aster. 
Fig. 4315. . 
Aster iaevis L. Sp. Pl. 876. 1753. 
Aster laevis ambplifolius Porter, Mem. Torr. Club 5: 
324. 1804. : 


Aster laevis potomacensis Burgess; Britt. & Brown, 
Ill. Fl. 3: 369. 1898. 


Stem usually stout, glabrous, often glaucous, 
2°-4° high, branched or simple. Leaves thick, en- 
tire, or serrate, glabrous, slightly rough-margined, 
the upper all sessile and usually cordate-clasping, 
lanceolate, oblong-lanceolate, oblanceolate or 
ovate, acute or obtusish, 17-4’ long, 4-2’ wide, 
the basal and lower gradually narrowed into 
winged petioles, those of the branches often small 
and bract-like; heads usually numerous, about 1’ 
broad; involucre campanulate, its bracts rigid, 
acute, appressed, green-tipped, imbricated in sev- 
eral series; rays I5-30, blue or violet; pappus 
tawny; achenes glabrous or nearly so. 


Usually in dry soil, Maine to Ontario, Virginia, 
Alabama, Louisiana, Saskatchewan, Missouri and 
Colorado. Races differ in leaf-form. Sept.—Oct. 


35. Aster concinnus Willd. Narrow-leaved \\ 
Smooth Aster. Fig. 4316. r= 


(f 


Aster concinnus Willd. Enum. 884. 1809. 


Similar to narrow-leaved forms of Aster laevis, 
and perhaps a race of that species, glabrous, or 
sparingly pubescent above, not glaucous; stem 
paniculately branched, 1°-3° high. Leaves light 
green, lanceolate to linear, entire, or sometimes ser- 
rulate, the upper sessile, somewhat clasping, 17-3’ 
long, the lower and basal ones spatulate, or oblong, 
narrowed into margined petioles, sometimes coarsely 
toothed; heads usually numerous, about 10” broad; 
bracts of the involucre with rhomboid acute herba- 
ceous tips; rays violet to purple. 


Woodlands, Connecticut to Pennsylvania, Virginia, 
North Carolina, Missouri and Arkansas. Sept.—Oct. 


36. Aster purpuratus Nees. Southern 
Smooth Aster. Fig. 4317. 


Aster virgatus Ell. Bot. S. C. & Ga. 2: 353. 1824. Not 
Moench, 1802. 
A, purpuralus Nees, Gen. & Sp. Ast. 118. 1832. 


Stem slender, glabrous, simple, or branched 
above, 13°-3° high, the branches sometimes pu- 
berulent. Leaves firm, glabrous, dark green, en- 
tire, the upper sessile and clasping at the base, 
elongated-lanceolate or linear-lanceolate, acumi- 
nate, 2’-6’ long, 2-4” wide, the lower and basal 
ones petioled, oblong-lanceolate, obtusish, those 
of the branches very small; heads rather few, 
loosely paniculate, 8’-12” broad; involucre cam- 
panulate to turbinate, its bracts coriaceous, linear, 
appressed, green-tipped, acute, imbricated in sev- 
eral series, the outer shorter; rays 5-10, blue or 
violet, 3-5” long, pappus tawny; achenes glabrous. 


Virginia and West Virginia to Georgia and Texas. 
Aug.—Sept. : 


GENUS 31. THISTLE FAMILY. 421 


37. Aster janceus Ait. Rush Aster. . 
Fig. 4318. 
Aster junceus Ait. Hort. Kew. 3: 204. 17809. 
Aster longulus Sheldon, Bull. Geol. Surv. Minn. 9: 18, 
pl. 2. 1894. 
Aster junciformis Rydb. Bull. Torr. Club 37: 142. 1910. 
Stem very slender, glabrous, or pubescent above, 
simple or little branched, 1°-3° high. Leaves firm, 
glabrous, roughish-margined, narrowly linear, entire 
or sometimes with a few distant teeth, acute or 
acuminate at the apex, sessile by a broad clasping 
and often slightly cordate base, 3’-6’ long, 13’-4” 
wide; heads paniculate, rather distant, about 1’ 
broad; involucre hemispheric, about 3” high, its 
bracts glabrous, linear-subulate, very acute, imbri- 
cated in 3 or 4 series, the outer shorter; rays violet 
to white, 4’-5”" long; pappus pale. 


In swamps and bogs, Nova Scotia to British Columbia, 
New Jersey, Ohio, Wisconsin and Colorado. July—Sept. 


38. Aster adscéndens Lindl. Western 
Aster. Fig. 4319. 
A. adscendens Lindl.; Hook. Fl. Bor. Am. 2: 8. 1834. 


Stem slender, rigid, glabrous, or sparingly 
hirsute-pubescent, branched or simple, 62° high. 
Leaves firm; entire, rough-margined, sometimes 
ciliolate, those of the stem linear-lanceolate or 
linear-oblong, acute or obtusish, 1-3’ long, 2”—5” 
wide, sessile by a more or less clasping base; 
basal leaves spatulate, narrowed into short peti- 
oles; heads not numerous, about 1’ broad; invo- 
lucre hemispheric, its bracts imbricated in 3-5 
series, oblong-linear or spatulate, their tips obtuse 
or obtusish, slightly spreading, the inner often 
mucronulate; pappus nearly white; achenes pu- 
bescent. 

On prairies and moist banks, western Nebraska to 


Wyoming, Montana, Assiniboia, Colorado, New Mex- 
ico and Nevada. July—Sept. 


39. Aster novi-bélgii L. New York Aster. Fig. 4320. 


; = WY 
Aster novi-belgii L. Sp. Pl. 877. 1753. \\ e ee 
Aster novi-belgii elodes A. Gray, Syn. Fl. 17: 190. 1884. WS yz NA lia 
Aster novi-belgii litoreus A. Gray, loc. cit. 189. 1884. ES ie NA AQ NW 
ANS cy WY : 


Aster novi-belgii atlanticus Burgess; Britt. & Brown, 
Ill. Fl. 3: 370. 1808. 
Aster novi-belgii Brittonii Burgess, loc. cit. 371. 1898. 


Stem slender, usually much branched, glabrous, or 
slightly pubescent above, 1°-3° high. Leaves lanceo- 
late, oblong-lanceolate, or linear-lanceolate, firm, 
often somewhat fleshy, entire, or slightly serrate. 
glabrous, or very nearly so, acuminate at the apex, 
narrowed, sessile and more or less clasping at the 
base, 2-6’ long, 3-8” wide, the lowest petioled; 
heads corymbose-paniculate, usually numerous, 10’— 
15” broad; involucre hemispheric to campanulate, 
its bracts linear, acute, or obtusish, green, somewhat 
spreading, in 3-5 series, the outer shorter; rays 15- 
25, violet, 4’-5” long; pappus whitish; achenes gla- 
brous or nearly so. 


422 COMPOSITAE. Vor. IIT. 


In swamps, Newfoundland to Maine and Georgia, mainly near the coast. Races differ in leaf- 
form and in the involucrai bracts. Aug.—Oct. 


Givi a 40. Aster longifolius Lam. Long-leaved 
WE Aster. Fig. 4321. 
VS 


Aster longifolius Lam. Encycl. 1: 306. 1783. 


Aster longifolius villicaulis A. Gray, Syn. Fl. 1: Part 2. 
189. 1884. 


Stem glabrous, or pubescent, leafy, paniculately 
branched, 1°-3° high. Leaves lanceolate to linear- 
lanceolate, entire, or nearly so, acuminate at the 
apex, narrowed into a sessile clasping usually slightly 
cordate base, 3-8’ long, 2”-6” wide; heads rather 
numerous, about I’ broad; involucre hemispheric, 
4-5” high, its bracts glabrous, narrow, green, acute, 
imbricated in few series, nearly equal; rays numer- 
ous, 23’—-7” long, violet or pale purple; pappus pale. 


In swamps and moist ground, Labrador to Saskatche- 
wan, northern New England, Ontario and Montana. 
Summer. 


41. Aster foliaceus Lindl. Leafy-bracted 
Aster. Fig. 4322. 
Aster foliaceus Lindl, in DC. Prodr. 5: 228. 1835. 


Stem usually stout, sparingly pubescent in lines, 
2°-3° high, branched above, the branches ascend- 
ing. Leaves lanceolate, acute at the apex, entire, or 
with a few small distant teeth, glabrous on both 
sides, very rough-margined, 2-4’ long, 3-1’ wide, 
the upper clasping at the base, the lowest petioled; 
heads few, 1’ broad or more; involucre hemispheric, 
its bracts green, foliaceous, oblong, the inner nar- 
rower and acute; rays about 30, violet, 4-5” long; 
pappus nearly white; achenes pubescent. 


Mountains of Quebec; Oregon to Alaska. July—Sept. 
The figure of this species in our first edition was erro- 
neously stated to have been drawn from specimens col- 
lected in western Nebraska. 


42. Aster phyllodes Rydb. Large-bracted 
Aster. Fig. 4323. 


Aster phyllodes Rydb. Bull. Torr. Club 37: 145. 1910. 


Stem rather slender, somewhat branched, loosely 
pubescent, 2°-3° high. Leaves oblong-lanceolate, 
entire, or sparingly low-dentate, the larger 4’ 
long or less, 4—1’ wide, acute or acutish at the 
apex, narrowed to a subcordate and slightly 
clasping base, rough-margined, glabrous on both 
sides; heads leafy-paniculate, about 1’ broad; in- 
volucre hemispheric, its bracts narrowly oblong, 
foliaceous, acute; rays purplish; achenes pubes- 
cent. 


Wet grounds, western Nebraska and eastern Colo- 
rado. Aug.—Sept. Included in A. foliaceus Lindl. in 
our first edition. 


GENUS 3l. THISTLE FAMILY. 423 


43. Aster sericeus Vent. Western Silvery or 
Silky Aster. Fig. 4324. 

Aster sericeus Vent. Hort. Cels, pl. 33. 1800. 

Aster argenteus Michx. Fl. Bor. Am. 2: 111. 1803. 


Stem slender, paniculately or corymbosely branched, 
stiff, glabrous, leafy, 1°-2° high. Stem leaves sessile, 
with a broad base, oblong, entire, mucronate, 3’—12’ 
long, 2’-5”’ wide, erect or ascending, with a dense 
silvery-white silky pubescence on both sides; basal 
and lowest leaves oblanceolate, narrowed into mar- 
gined petioles; heads numerous, about 13’ broad; in- 
volucre turbinate, its bracts oblong, or the inner 
lanceolate, canescent, imbricated in 3 or 4 series, their 
tips green, acute, spreading; rays I5-25, violet-blue, 

“8” long; pappus tawny; achenes glabrous. 


In dry open soil, Ilinois to Minnesota, Manitoba, 
South Dakota, Tennessee, Missouri and Texas. Aug.— 
Sept. 


SN 2 44. Aster concolor L. Eastern Silvery 
CANN . : 
WM WwW Y) Aster. Fig. 4325. 
es Aster concolor L. Sp. Pl. Ed. 2, 1228. 1763. 
aS 
WN Stem slender, glabrous, or pubescent above, 


<—— 


== 


= Tt 


, 


= 


1°-23° high, leafy, simple, or with few erect 
branches. Leaves oblong or linear-oblong, 
finely and densely canescent on both sides, or 
the lower glabrate, sessile, obtuse or mucro- 
nate, 12’-2’ long; heads numerous in an elon- 
gated narrow raceme resembling Lacinaria; 
involucre broadly turbinate, its bracts linear 
or linear-oblong, appressed, canescent, imbri- 
cated in 4 or 5 series, their tips green, acute, 
the outer shorter; rays 10-15, lilac, 3’—4” long; 
pappus tawny; achenes villous. 


SS 


>» sah 
i. 


SN 


pS 


Ke 


Nw 


a 


= N AD In dry sandy soil, eastern Massachusetts and 
A 4 rR D Rhode Island to Florida, Tennessee and Louisiana, 
CT Y / if f J JX ed near the coast. Lilac-flowered aster. Aug.— 
Ws / : ct. 
NG Wavy 
45. Aster Féndleri A. Gray. Fendler’s <a DY VR 
i] Pogue 


exstet, | «Fig. 41326, 


ae Fendleri A. Gray, Mem. Am. Acad. (II) 4: 66. 
1849. 


Aster Nuttallii var. Fendleri A. Gray, Pac. R. R. Rep. 
At o7. Toso: 


Stems several or solitary from thick woody 
roots, rigid, hirsute, 6-12’ high. Leaves linear, 
rigid, I-nerved, acute or acuminate, 8’-15” long, 
1-13” wide, glabrous on both sides, but the mar- 
gins bristly-ciliate; heads usually few and race- 
mose, 3-1’ broad; involucre turbinate, its bracts 
glandular, linear-oblong, imbricated in about 4 
series, the inner acute, the outer shorter and ob- 
tuse; rays 10-15, violet, 3-5” long. 


In dry soil on the plains, Nebraska and Kansas to 
Colorado and New Mexico. Aug.—Sept. 


to 


4 COMPOSITAE. | Vot. III, 


4 


46. Aster grandiflorus L. Large- 
flowered Aster. Fig. 4327. 


Aster grandiflorus ILS SpE IY tee Gis HGS) 


Stem rather stiff, divaricately much branched, 
hispid with short hairs, 1°-23° high. Leaves 
oblong, linear, or somewhat spatulate, rigid, 
sessile by a broad, sometimes slightly clasping 
base, reflexed, entire, obtusish, hispid, the 
larger 2’ long and 4” wide, those of the branches 
very numerous, 2-5” long; heads about 2’ 
broad, terminating the branches; involucre 
hemispheric, its bracts very squarrose and 
foliaceous, imbricated in 5-7 series, linear, or 
linear-oblong, glandular, the outer obtusish, 
the inner acute; rays very numerous, deep 
violet, nearly 1’ long, 13” wide; pappus brown- 
ish; achenes ribbed, canescent. 


In dry soil, Virginia, east of the mountains, to 
Florida. Sept. 


47. Aster spectabilis Ait. Low showy 
Aster. Seaside Purple Aster. 
Fig. 4328. 
Aster spectabilis Ait. Hort. Kew. 3: 209. 1789. 


Stem stiff, simple, or corymbosely branched 
above, puberulent, or rough below, more or 
less glandular above, 1°-2° high. Leaves firm, 
thickish, the basal and lower ones oval, acute 
or acutish, 3-5’ long, 1-13’ wide, sparingly 
dentate with low teeth, narrowed at the base 
into slender petioles; upper leaves sessile, en- 
tire or very nearly so, acute, linear-oblong; 
heads several or numerous, about 13’ broad, 
corymbose, very showy; involucre nearly hemi- 
spheric, its bracts linear-oblong or slightly 
spatulate, glandular, viscid, imbricated in about 
5 series, their green obtusish tips spreading ; 
rays 15-30, bright violet, 6’—-10” long; pappus 
whitish; achenes slightly pubescent. 


In dry sandy soil, Massachusetts to Delaware, 
mostly near the coast. Aug.—Oct. 


WV 48. Aster surcul6sus Michx. Creeping 
WZ Aster. Fig. 4320. 

Toe te g. 43 9 

al Aster surculosus Michx. Fl. Bor. Am. 2: 112. 1803. 


Stem slender, from elongated-filiform root- 
stocks, minutely scabrous-pubescent, 10-18" 
high, corymbosely branched above. Leaves 

' firm, lanceolate or linear, the lower petioled, 
2’-3/ long, 4”-8” wide, rough-margined, slightly 
scabrous above, sparingly dentate, the upper 
narrower, sessile, entire; heads few, or some- 
times solitary, about 15” broad; involucre tur- 
binate-hemispheric, its bracts coriaceous, im- 
bricated in about 5 series, ciliate, but scarcely 
glandular, their green tips spreading; rays 15- 
30, violet; pappus whitish; achenes nearly gla- 
brous. 


_ In sandy or gravelly soil, Kentucky, North Caro- 
lina and Georgia. Erroneously reported from New 
Jersey. Sept.—Oct. 


GENUS 31. 


49. Aster gracilis Nutt. Slender or 
Tuber Aster. Fig. 4330. 
Aster gracilis Nutt. Gen. 2: i158. 1818. 


Stem slender, finely puberulent and sca- 
brous, corymbosely branched above, 1°-12 
high. Leaves minutely scabrous, the basal 
and lower ones oval, acute or obtusish, 2-3’ 
long, 4-8” wide, dentate, narrowed into 
slender petioles; upper leaves linear, linear- 
oblong, or slightly oblanceolate, acute, en- 
tire, sessile or a little clasping; heads usu- 
ally numerous, 6’-10” broad; involucre 
narrowly turbinate, its bracts coriaceous, 
glabrous or very nearly so, imbricated in 
about 5 series, their tips green and spread- 
ing, obtusish; rays 9-15, violet, 3-43” long; 
pappus nearly white; achenes minutely pu- 
bescent. 

In dry sandy soil, New Jersey to Kentucky, 


Tennessee and South Carolina. Rootstock 
tuberous-thickened. July—Sept. 


Nai 
awk 
SY Wi 


51. Aster Hérveyi A. Gray. 
Astera’ Pigeass2: 


Aster Herveyi A. Gray, Man. Ed. 5, 229. 1867. 


THISTLE FAMILY. 


Hervey’s 


425 


oa 


en 


cz 


p 


File-blade Aster. Fig. 4331. 


Aster Radula Ait. Hort. Kew. 3: 210. 1780. 

Aster bifiorus Michx. Fl. Bor. Am. 2: 114. 1803. 

Aster strictus Pursh, Fl. Am. Sept. 556. 1814. 

Aster Radula var. strictus A. Gray, Syn. Fl. 1: Part 
2,176. 1884. 

Aster Radula biflorus Porter, Mem. Torr. Club 5: 
326. 1894. 


Stem glabrous, or puberulent above, slender, 
corymbosely branched near the summit, or sim- 
ple, 4’-2° high. Leaves sessile, rough above, 


‘more or less pubescent beneath, lanceolate to 


oblong-lanceolate, acute, sharply serrate, strong- 
ly pinnately veined, 2’-3’ long, 3’-12” wide; 
heads several, or sometimes numerous, I-12’ 
broad; involucre hemispheric, its bracts oblong 
or oblong-spatulate, coriaceous, appressed-pu- 
bescent, conspicuously ciliolate, their green ob- 
tuse or acutish tips very little spreading; rays 
20-30, violet, 4-6” long; achenes glabrous, 
striate; pappus nearly white. 

In swamps, Newfoundland to Delaware, Penn- 
sylvania, Maryland and West Virginia. July—Sept. 


Stem roughish, at least above, slender, simple 
or corymbosely branched, rarely paniculate, 1°-3° 
high, the branches glandular-puberulent. Leaves 
firm, rough above, pubescent on the veins beneath, 
the basal and lower ones on slender naked peti- 
oles, ovate, dentate with low usually distant teeth, 
acute at the apex, narrowed, rounded or rarely 
cordate at the base, 2’-6’ long, 1-3’ wide; upper 
leaves sessile, or narrowed into winged petioles, 
smaller, entire or nearly so; heads 1-13’ broad; 
involucre turbinate or campanulate, its bracts ap- 
pressed, or sometimes spreading, densely glandu- 
lar, oblong or spatulate, obtuse or mucronulate; 
rays 15-25, violet, 5’—7’”’ long; achenes minutely 
pubescent, striate; pappus nearly white. 


In dry soil, eastern Massachusetts, Rhode Island, 
Connecticut and Long Island. Aug.—Oct. 


COMPOSITAE. Vor. III. 


53. Aster iaterifldrus (L.) Britton. Starved 
Aster. Calico Aster. Fig. 4334. 
Solidago lateriflora L. Sp. Pl. 879. 1753. 


52. Aster turbinéllus Lindl. Prairie 
Aster, “Bigs 4333: 


A, turbinellus Lindl. Comp. Bot. Mag. 1: 98 1835. 


Stem slender, paniculately branched, glabrous 
below, puberulent above, 2°-3° high. Leaves firm, 
lanceolate, or oblong-lanceolate, entire, ciliate, 
acute or acuminate, 2-3’ long, the lower and 
basal ones petioled, the upper sessile, those of 
the branches much smaller; heads about 1’ broad, 
mostly solitary at the ends of the branches; in- 
volucre turbinate, its bracts oblong, coriaceous, 
obtuse, appressed, imbricated in 5 or 6 series, 
their tips green only at the apex; rays 10-20, 
3-5” long, violet; pappus tawny; achenes finely 
pubescent. - 


In dry soil, especially on prairies, Illinois to Mis- 
souri, Nebraska, Kansas, Louisiana and Arkansas. 
Sept.—Oct. 


Aster diffusus Ait. Hort. Kew. 3: 205. 1780. << WS 
Aster miser Nutt. Gen. 2: 158. 1818. Y) 
A, lateriflorus Britton, Trans. N. Y. Acad. Sci.g: 10. 1889. / aS 


Stem puberulent, or nearly glabrous, slender, diver- 
gently branched, 1°-5° high. Basal leaves ovate, slen- 
der-petioled; stem leaves broadly lanceolate or oblong- 
lanceolate, mostly acuminate, serrate, 2-5’ long, 6’—12” 
wide, those of the branches smaller, oblong or linear- 
oblong; heads 3-5” broad, racemosely unilateral on 
the branches, short-peduncled or sessile, usually nu- 
merous and crowded; involucre turbinate, its bracts 
linear-oblong, obtuse or acutish, imbricated in about 4 
series, their short green tips appressed or slightly 
spreading ; rays numerous, short, white or pale purple; 
disk-flowers purple; pappus white; achenes minutely 
pubescent. 

In dry or moist soil, Nova Scotia to western Ontario, 
south to North Carolina, Louisiana and Texas. Consists 
of many races, differing in leaf-form, inflorescence and pu- 
bescence. Called in Maryland rosemary. Aug.—Oct. 


54. Aster hirsuticatlis Lindl. Hairy- 
stemmed Aster. Fig. 4335. 


A. hirsuticaulis Lindl.; DC. Prodr. 5: 242. 1836. 


. Aster lateriflorus hirsuticaulis Porter, Mem. Torr. 
Club 5: 324. 1894. 


Stem slender, erect, 12°-3” high, pubescent, 
often nearly or quite to the base, the usually 
short branches spreading or ascending. Leaves 
thin, glabrous above, usually pubescent on the 
midvein beneath, serrate with a few appressed 
teeth, or entire, linear-lanceolate to lanceolate, 
sometimes 6’ long, 2’—7” wide, sessile, or the 
basal ones spatulate and petioled; heads more 
or less unilateral on the branches, densely or 
loosely clustered, often also solitary or few in 
the lower axils; bracts of the involucre in 3 
or 4 series, linear-lanceolate, acuminate or 
acute; rays white, about 2” long. 


In woods and thickets, New Brunswick to Penn- 
sylvania, Kentucky and Michigan. Perhaps a 
race of the preceding species. Aug.—Oct. Wise- 
weed. Old-fieldsweet. Farewell-summer. Old-Vir- 
ginia. Stickweed. White-devil. Nail-rod. 


GENUS 3I. THISTLE FAMILY. 427 
55. Aster vimineus Lam. Small White Aster. Fig. 43306. 

Aster vimineus Lam. Encycl. 1: 306. 1783. i) MIG 
Aster foliolosus Ait. Hort. Kew. 3: 203. 1789. NUN a 
Aster Tradescanti T. & G. Fl. N. A. 2: 129. 1841. Not evn ae 

17530 ‘ WW WW) WP 
Aster vimineus foliolosus A. Gray, Syn. Fl. 1: Part 2, a Sy 

186, 1884, ay We VY 

Glabrous or nearly so throughout; stem slender, NZ “US XY NW 
divergently branched, 2°-5° high. Stem leaves linear- Wg y WE 
lanceolate, entire, or with a few low teeth, 3’-5’ wy \\ a 
long, 2’-4”" wide, acuminate at the apex, narrowed SOS \Z 
to a sessile base, those of the branches much smaller ; , 
heads very numerous, small, 3-5” broad, 2’-3” \ / Sauer 
high, generally densely racemose-secund, sometimes 4 fp Awe oY" 
subpaniculate, short-peduncled; involucre broadly / — 


turbinate, its bracts linear, acute or acutish, imbri- 
cated in about 3 series, green-tipped, appressed; 
rays numerous, about 2” long, white to purplish; 
pappus white; achenes minutely pubescent. 


A WYNY iy 


In moist soil, Ontario to Florida, west to Minnesota WO, 
and Arkansas. Aug.—Sept. , CEL fp. 
Le SS 
Aster racemdésus EIl., with smaller heads, the leaves ; SS 


all very small, of the Southeastern States, apparently 
occurs in extreme southern Illinois. 


56. Aster multiflorus Ait. Dense-flowered 


Aster. White Wreath Aster. Fig. 4337. 
Aster multiflorus Ait. Hort. Kew. 3: 203. 1789. 
A. multiflorus stricticaulis T.& G. Fl. N. A.2: 125. 1841. 
A. multiflorus exiguus Fernald, Rhodora 1: 187. 1899. 
A, exiguus Rydb. Bull. Torr. Club 28: 505. i901. 
A, polycephalus Rydb. Bull. Torr. Club 33: 153. 1906. 


Stem strict, much branched and bushy, rough- 
pubescent or scabrous, 1°-7° high, the branches as- 
cending or spreading. Leaves rigid, linear or linear- 
oblong, entire, mostly obtuse, sessile or slightly 
clasping at the base, rough and ciliate, those of the 
stem 47-12’ long, those of the branches very small 
and crowded; heads 3’—-4” broad, densely crowded, 
nearly sessile, sometimes slightly secund on the 
branches; involucre 2”-3” high, its bracts coriaceous, 
mostly ciliate or pubescent, in 3 or 4 series, their 
short green tips obtuse or mucronate, spreading; 
rays 10-20, white, 13”-2” long; pappus brownish 
white; achenes puberulent. 

In dry open places, Maine and Ontario to Alberta, 


Georgia, Texas and Arizona. Consists of many slightly 
differing races. Fall-flower. Aug.—Nov. 


57. Aster commutatus (T. & G.) A. Gray. 


White Prairie Aster. Fig. 4338. 
Aster ramulosus var. incanopilosus Lindl. in DC. Prodr. 5: 
243. 1836. 
Aster multiflorus var. commutatus T. & G. Fl. N. A. 2: 125. 
1841. 
Aster commutatus A. Gray, Syn. Fl. 1: Part 2, 185. 1884. 
A. incanopilosus Sheldon, Bull. Torr. Club 20: 286. 1893. 


Similar to the preceding species, except in the inflores- 
cence, the stem rough-pubescent, or sometimes nearly 
glabrous, 12°-23° high, with ascending or divergent 
branches. Leaves linear or linear-oblong, obtuse, entire, 
sessile ot slightly clasping at the base, those of the stem 
1’ s long; heads larger than those of A. multiflorus, 
6’-8” broad; involucre 3-4” high, its ciliate bracts 
squarrose-tipped and sometimes foliaceous; rays 20-30, 
about 3” long. 


On prairies and along rivers, Minnesota to Saskatchewan, 
Nebraska, Texas and New Mexico. Aug.—Oct. 


428 COMPOSITAE. Vor. III. 


58. Aster dumosus L. Bushy Aster. Rice-button Aster. Fig. 4339. 


Aster dumosus L. Sp. Pl. 873. 1753. 
Aster coridifolius Michx.; Willd. Sp. Pl. 3: 2028. 1804. 
As er dumosus strictior T. & G. Fl. N. A. 2: 128. 1841. 
Aster dumosus coridifolius T.& G. Fl.N.A.2:128. 1841. 
Glabrous or nearly so throughout, rather stiff and 
viscid, usually paniculately branched, 1°-3° high. 
Leaves firm, those of the stem linear or linear-lan- 
ceolate, entire, acute, or obtusish, 1-3’ long, 13-3” 
wide, roughish-margined, often reflexed, those of 
the branches very numerous, small and bract-like, 
the basal ones spatulate, dentate; heads 4’-7” broad, 
terminating the slender minutely leafy branches and 
branchlets, usually numerous; involucre broadly 
campanulate, its bracts linear-subulate, obtuse or 
acutish, appressed, imbricated in about 4 series, 
green-tipped ; rays 15-30, blue to pale violet or white, 
2’-4” long; pappus white; achenes pubescent. 
Sandy soil, Massachusetts to western New York, 
Ontario, Florida, Louisiana and Missouri. Aug—Oct. 
Aster Gravesii Burgess, known only from Water- 
ford, Conn., has oblong-lanceolate leaves, acuminate at 
both ends, the larger 8” wide, the rays bright purple. 


/ 


SG 
\ 


NNW 
59. Aster salicifolius Lam. Willow Aster. SS Ps ~Z 
Fig. 4340. SNE 
Aster salicifolius Lam. Encycl. 1: 306. 1783. f ) SS 
Aster subasper Lindl. Comp. Bot. Mag. 1: 97. 1835 \ ( 
Aster stenophyllus Lindl. DC. Prodr. 5: 242. 1836 


— 


Stem rather slender, paniculately much branched, 
usually very leafy, 2°-5° high, glabrous, roughish or 
somewhat pubescent above. Leaves firm, lanceolate ZA 
or linear-lanceolate, roughish or rough-margined, 
acute or acuminate at the apex, narrowed and sessile 
or slightly clasping at the base, entire or sparingly 
dentate with low teeth, glabrous or nearly so, 2’—4’ 
long, 2”-6” wide, the lowest sometimes petioled, 
those of the branches gradually smaller; heads nu- 
merous, 8-12” broad; involucre 3-4” high, its 
bracts linear-oblong, appressed, imbricated in 4 or 5 
series, their green tips acute or obtusish; rays nu- 
merous, violet, or violet-purple, or sometimes white, 
3-4” long; pappus white; achenes pubescent. 

In moist soil, Maine to Massachusetts, Florida, On- 
tario, Assiniboia, Texas and Colorado. Aug.—Oct. 

Ny Fi e ~ . 
WAV, 66. Aster paniculatus Lam. Tall White 
WA as or Panicled Aster. Fig. 4341. 
SSS\ \V/ 'Z Aster paniculatus Lam. Encycl. 1: 306. 1783. 
Aster bellidiforus Willd. Enum. 886. 1809. 
Aster tenuifolius var. bellidiforus T. & G. Fl. N. A. 

PSNR tev lie 

Stem glabrous or nearly so, or sometimes pu- 
bescent, paniculately much branched, 2°-8° high. 
Leaves lanceolate to oblong-lanceolate or linear, 
acuminate at the apex, narrowed to a sessile or 
slightly clasping base, glabrous, usually thin, 
roughish-margined, those of the stem sparingly 
serrate, or sometimes entire, 3’-6’ long, 3-6” 
wide, the upper and those of the branches grad- 
ually smaller; heads numerous, 8’—10” broad; in- 
volucre 3’-4” high, its bracts narrowly linear- 
lanceolate, acuminate, appressed, green-tipped, 
imbricated in 4 or 5 series; rays numerous, white, 
or faintly tinged with violet, 3’-4” long; pappus 
white or nearly so; achenes minutely pubescent. 

In moist soil, New Brunswick to western Ontario 
and Montana, south to New Jersey, Virginia, Ken- 
tucky, Louisiana and Missouri. Consists of many 
races, differing in leaf-form, leaf-serration, size of 
heads, color of rays, and pubescence. Aug.—Oct. 


ee 
Ts <4 


—s 
= 


GENUS 31. THISTLE FAMILY. 429 


61. Aster missouriénsis Britton. Missouri 
AStetes Whe. 4342! 


Aster missouriensis Britton, in Britt. & Brown, Ill. Fl. 3: 
378. 1898. 


Stem densely puberulent or pubescent, at least 
above, much branched, 2° high or more. Leaves thin, 
oblong-lanceolate to oblanceolate, acute or acumi- 
nate at the apex, sharply serrate above the middle, 
gradually tapering to an entire sessile or slightly 
clasping base, or the lower petioled, puberulent above, 
finely pubescent beneath, the larger 3’-4’ long, the 
upper much smaller, entire; heads 6”’-8” broad, pan- 
icled, short-peduncled, or terminating short leafy 
branchlets, sometimes somewhat secund; involucre 
2-3” high, its linear acute bracts well imbricated, 
ciliate or pubescent; rays white. 


In moist soil, Kansas, Missouri and Iowa. Sept—Oct. 


62. Aster Tradescanti L. Tradescant’s 
Aster. Michaelmas Daisy. Fig. 4343. 
Aster Tradescanti L. Sp. Pl. 876. 1753. 


Stem slender, paniculately branched, 2°-5° high, 
the branches usually ascending and often pubes- 
cent in lines. Stem leaves linear-lanceolate or 
lanceolate, acuminate at the apex, narrowed to a 
sessile base, 3-6’ long, 13-6” wide, glabrous or 
nearly so on both sides, commonly thin, sharply 
serrate in the middle with low teeth, or sometimes 
entire; heads very numerous, racemose but not 
secund on the branches, 5-8” broad; involucre 
hemispheric to broadly turbinate, 2-3” high, its 
bracts linear, acute, appressed, green-tipped, im- 
bricated in 4 or 5 series; rays white or nearly so, 
numerous, 2’-3” long; pappus white; achenes 
minutely pubescent. 

In fields and swamps, Maine to Virginia, Ontario, 
Illinois and Missouri. Aug.—Oct. 

Aster saxatilis (Fernald) Blanchard, of rocky situ- 

_.ations in New England, appears to be a low race of 
this species, rather than of A. vimineus. 


4 1S é 


63. Aster Faxoni Porter. Faxon’s Aster. 
Fig. 4344. 


Aster polyphyllus Willd. Enum. 888. 1809. Not Moench, 
1802. 


Aster Faxoni Porter, Mem. Torr. Club 5: 323. 1894. 
A, ericoides Randi Britton, in Britt. & Brown, Ill. Fl. 3: 
379. 18098. 


Glabrous throughout; stem paniculately or corym- 
bosely branched; rather stout, 3°-5° high. Stem 
leaves lanceolate or linear-lanceolate, acute or acumi- 
nate, narrowed to'a sessile base, or the lower into 
margined petioles, entire or nearly so, firm, 2’—5’ 
long, 2’-4” wide, those of the branches gradually 
smaller; basal leaves oblong to spatulate, obtuse, 
dentate, or entire; heads not very numerous, 6-9” 
broad; involucre hemispheric, nearly 4” high, its 
bracts linear-lanceolate, acute or subulate, green- 
tipped or green on the back, imbricated in about 3 
series, the outer shorter; rays bright white, rarely 
purplish, 3’-4” long, numerous; pappus_ white; 
achenes minutely pubescent. 


On moist cliffs, Maine and Vermont to Pennsylvania, 
Wisconsin and North Carolina. Aug.—Sept. 


430 COMPOSITAE. Vou. LIT. 


Gi A 64. Aster ericoides L. White Heath Aster. 
OA Sz Frost-weed Aster. Fig. 4345. 


Aster ericoides L. Sp. Pl. 875. 1753. 
Aster villosus Michx. Fl. Bor. Am. 2: 113. 1803. Not 


Thunb. 1800, 

Aster ericoides var. villosus T. & G. Fl. N. A. 2: 124. 
1841. 

Aster ericoides pilosus Porter, Mem. Torr. Club 5: 
323. 1894. 


Stem glabrous, villous, or hirsute, paniculately 
branched, usually bushy, 1°-3° high, the branches 
racemose, and the branchlets often somewhat se- 
cund. Leaves firm or rigid, the basal ones spatu- 
late, obtuse, dentate, narrowed into margined peti- 
oles, glabrous or ciliate; stem leaves narrowly 
linear to linear-lanceolate, acute, entire, 1-3’ long, 
1-3” wide, those of the branches linear-subulate 
numerous; heads usually very numerous, 4-7’ 
broad; involucre campanulate to hemispheric, its 
bracts coriaceous, lanceolate or linear-lanceolate, 
abruptly acute or acuminate, green-tipped, imbri- 
cated in about 3 series; rays 15-25, white, or tinged 
with rose; pappus white; achenes finely pubescent. 

i In dry soil, Maine to Ontario, Florida, Minnesota 
and Missouri. Frost-weed. Michaelmas daisy. Farewell-summer. White rosemary. Dog-fennel. 
Mare’s-tail. Scrub-bush. Steel-weed. Sept—Dec. 

A densely villous, broad-leaved relative or race of this species, ranging from Ohio to North 


Carolina and Michigan, is known as Aster ericoides platyphyllus T. & G 
This species apparently hybridizes with A. paniculatus Lam. where the two grow together. 


2 
fd 


65. Aster Pringlei (A. Gray) Britton. 
Pringle’s Aster. Fig. 4346. 
Aster ericoides var. Pringlei A. Gray, Syn. Fl. 1: 

Part 2, 184. 1884. 

Aster Pringlei Britton, in Britt. & Brown, Ill. Fl. 3: 

379. 1898. 

Stem very slender, glabrous, simple, or with 
few or numerous slender ascending branches, not 
bushy, 6’-2° high. Basal leaves lanceolate, oblong 
or oblanceolate, 2’-6’ long, 2”-6” wide, entire, or 
slightly toothed, ciliate and sometimes a little 
pubescent, at least on the slender petioles which 
are often as long as the blades; stem leaves nar- 
rowly linear, those of the branches small and sub- 
ulate; heads as large as those of A. ericoides, or 
commonly smaller, usually fewer, solitary at the 
ends of the branches and branchlets; bracts of 
the involucre with short green tips; rays white. 

On banks, especially in rocky places, Massachusetts 
and Vermont to Wisconsin. Aug.—Oct. 


Ww (4 
NY) Wy 
N ) 
AW Wt 
Cin 66. Aster Priceae Britton. Miss Price’s Aster. 


Fig. 4347. 
Aster Priceae Britton, Manual 960. gor. 


Stem pubescent, widely branched, 1°-23° high. Basal 
leaves oblanceolate, obtuse or acutish, entire, petioled, 
1-3’ long, the petioles ciliate, broad; stem leaves linear- 
lanceolate, sessile, ciliate, acuminate, ?’—2’ long, those 
of the branches similar but smaller; involucre nearly 
AW; hemispheric, about 3” high, its bracts linear, the outer 
QW \ gradually acuminate, green, the inner a little broader. 
abruptly acuminate, acute or obtusish; heads about I” 
broad; rays bright purple or pink. 


In dry soil, Kentucky and North Carolina. October. 


Aster kentuckiénsis Britton, also of Kentucky and North 
Carolina, differs in being glabrous, its involucral bracts all 
gradually acuminate. 


GENUS 31. THISTLE FAMILY. 431 


67. Aster parviceps (Burgess) Mackenzie & 


= WA PS 
Bush. Small-headed Aster. Fig. 4348. y ay ae 
Aster ericoides parviceps Burgess; Britt. & Brown, Ill. : 
Hla 37050 eos: as WA} 
« ! =) 


Ly Wz. 
Aster parviceps Mackenzie & Bush, Fl. Jackson Co. 196. * 


1902, 


A. depauperatus parviceps Fernald, Rhodora 10: 94. 
1908. 


Pilose, sometimes slightly so, much branched above, 
1°-23° high, the branches ascending. Basal leaves 
spatulate; stem leaves linear to linear-lanceolate, 
1-3’ long, often with short leafy branches in their 
axils, those of the branches very small; heads many, 
paniculate; involucre about 2” high, turbinate, its 
linear-subulate bracts imbricated in several series; 
rays 10-12, white, about 2” long. 


Dry prairies and open woodlands, Illinois, Iowa and 
Missouri. Aug.—Sept. 


68. Aster depauperatus (Porter) Fernald. 
Serpentine Aster. Fig. 4340. 


Aster ericoides pusillus A. Gray, Syn. Fl. 17: 184. 1884. 
Not A. pusillus Horn. 

Aster ericoides depauperatus Porter, Mem. Torr. Club 
Bi 323.) 1804. 

Aster depauperatus Fernald, Rhodora 10: 94. 1908. 

Aster parviceps pusillus Fernald, Rhodora 11: 59. 1909. 


Glabrous, slender, widely branched, 4’-15’ high. 
Basal leaves spatulate or oblanceolate, obtusish, 3’—13’ 
long, narrowed into petioles; stem leaves linear to 
linear-subulate, 1’ long or less, 2’”-1”’ wide, those of 
the branches minute; heads many, terminating short 
branchlets; involucre turbinate, about 2” high, its 
bracts linear-subulate, acute, rays white, about 2” 
long. 


On serpentine barrens, southern Pennsylvania and 
West Virginia. Aug.—Oct. 


69. Aster paluddosus Ait. Southern 
Swamp Aster. Fig. 4350. 


Aster paludosus Ait. Hort. Kew. 3: 310. 1780. 
Heleastrum paludosum DC. Prodr. 5: 264. 1836. 


Stem roughish, or rough-pubescent, slen- iy g\ 
der, simple, or somewhat branched above, 
1°-23° high. Leaves linear, entire, glabrous, 
but margins rough or ciliate, rigid, 26’ long, 
2-4” wide, mostly t-nerved, acute, the lower 
part commonly sheathing the stem; heads 
few or several, racemose or paniculate, 14-2’ 
broad; involucre broadly campanulate or 
hemispheric, its bracts imbricated in about 5 
series, gp aan the outer lanceolate, 
acute, the inner oblong or spatulate; rays ‘Gs 
20-30, deep violet, 5-7” long, pappus tawny; ———— 

a 
2 


achenes 8-10-nerved, glabrous, or nearly so. 


In swamps, Kansas and Missouri to Texas, east 
to North Carolina and Florida. Aug.—Oct. p 


432 COMPOSITAE. Vor. III. 


7o. Aster nemoralis Ait. Bog Aster. 


Fig. 4351. 
_ Aster nemoralis Ait. Hort. Kew. 3: 198. 1780. 
» Aster nemoralis Blakei Porter, Bull. Torr. Club 21: 

Sr. ereod. 

Stem puberulent, slender, simple, or, corym- 
bosely branched above, 62° high. Leaves ses- 
sile, oblong-lanceolate or linear-oblong, acute at 
each end, pubescent or puberulent on both sides, 
dentate or entire, 1-3’ long, 13”—10” wide, mar- 
gins often revolute; heads several, or solitary, 
1-13’ broad, the peduncles slender; involucre 
broadly obconic to hemispheric, its bracts ap- 
pressed, linear-subulate, acute or acuminate, im- 
bricated in about 3 series; rays 15-25, light 
violet-purple to rose-pink; achenes glandular- 
pubescent; pappus white. 

In sandy bogs, New Jersey to northern New 
York, Ontario, Newfoundland and Hudson Bay. 
Races differ in leaf-form and serration. Aug.—Sept. 


x. Whorled 
or Mountain Aster. Fig. 4352. 
Aster divaricatus Lam. Encycl. 1: 305. 1783. Not L. 
1753. 
Aster acuminatus Michx. Fl. Bor. Am. 2: 109. 1803. 
Stem pubescent or puberulent, zigzag, corym- 
bosely branched, often leafless below, 1°-3° high. 
Leaves thin, broadly oblong, acuminate at the 
apex, narrowed to a somewhat cuneate sessile 
base, sharply and coarsely dentate, pinnately vein- 
ed, glabrous or pubescent above, pubescent at 
least on the veins beneath, 3-6’ long, 4’-12’ wide, 
often approximate above, and appearing whorled; 
heads several or numerous, 1-13’ broad; invo- 
lucre nearly hemispheric, its bracts subulate- 
linear, acuminate, the outer much shorter; rays 
12-18, narrow, 6-8” long, white or purplish; 
pappus soft, fine, nearly white; achenes pubescent. 
Moist woods, Labrador to Ontario, western New 
York, and in the mountains to Tennessee and Geor- 
gia. July—Oct. 


72. Aster ptarmicoides (Nees) T.& G. Upland White Aster. Fig. 4353. 

Mi Chrysopsis alba Nutt. Gen.2:152. 1818. Not 
A. albus Willd. 

Doellingeria ptarmicoides Nees, Gen. & Sp. 
Ast. 183. 1832. 

A. ptarmicoides T. & G. Fl. N. A. 2: 160. 


1841. : 
Unamia alba Rydb. Bull. Torr. Club 37: 
146. 1910. 


Stems tufted, slender, rigid, usually 
rough above, corymbosely branched near 
the summit, 1°-2° high. Leaves linear- 
lanceolate, I-3-ribbed, entire, or with a 
few distant teeth, firm, shining, rough- 
margined or ciliate, sometimes scabrous, 
acute, narrowed to a sessile base, or the 
lower petioled, the lowest and basal ones 
3-6’ long, 2”-4” wide, the upper smaller, 
those of the branches linear-subulate; 
heads 8-12” broad; involucre nearly 
hemispheric, 2-3” high, its bracts lin- 
ear-oblong, obtuse, or the outer acutish, 
appressed, nearly glabrous, green, 1m- 
bricated in about 4 series; rays 10-20, 
white, 3-4” long; pappus white; achenes 
glabrous. 

; In dry or rocky soil, Massachusetts, Ver- 
mont and Ontario to Saskatchewan, Illinois, Missouri and Colorado. July—Sept. 


GENUS 31. THISTLE FAMILY. 433 


Aster lutéscens (Lindl.) T. & G. is a very interesting race with light yellow rays, known from 
Illinois, Wisconsin and Saskatchewan. Yellow rays are otherwise almost or quite unknown in the 
genus Aster as here limited. 


73. Aster tenuifolius L. Perennial Salt- 
marsh Aster. Fig. 4354. 


Aster tenuifolius L. Sp. Pl. 873. 1753. 
Aster flexuosus Nutt. Gen. 2: 154. 1818. 


Perennial, glabrous and fleshy; stem flexuous, 
striate, at least when dry, sparingly and loosely 
branched, 1°-2° high. Stem leaves linear, entire, 
acute, sessile or partly clasping at the base, the 
lowest lanceolate-linear, 2’-6’ long, 2”-3” wide, 
those of the branches minute, bract-like, ap- 
pressed; heads rather few, 6-12” broad, termi- 
nating the branches; involucre turbinate, about 
4” high, its bracts lanceolate, acuminate or mu- 
cronate, glabrous, green on the back or tip, ap- 
pressed, imbricated in about 5 series, the outer 
shorter; rays numerous, longer than the pappus, 
pale purple or nearly white; pappus tawny; 
achenes hispid-pubescent, 5-nerved. 


In salt marshes, coast of Massachusetts to Flor- 
ida. Aug.—Oct. 


74. Aster exilis Ell. Slim Aster. Fig. 4355. 
Aster exilis Ell. Bot. S. C. & Ga. 2: 344. 1824. 
Aster divaricatus T.& G. Fl. N. A. 2: 163. 1841. Not L. 1753. 


Annual, glabrous, fleshy; stem slender, usually much 
branched, the branches usually divergent. Leaves linear 
to linear-lanceolate, 1-4’ long, 1’—23” wide, entire, sessile, 
acute or acuminate, or the lowest narrowly oblong, 3’’-4” 
wide and petioled, those of the branches subulate; heads 
numerous, panicled, about 5” broad; involucre campanu- 
late, about 3” high, its bracts linear-subulate, appressed, 
imbricated in 3 or 4 series; rays purplish, about 2” long, 
mostly fewer than the disk-flowers, longer than the pappus ; 
achenes somewhat pubescent. 


In moist or wet soil, especially in saline situations, Kansas to 
Texas, South Carolina and Florida. Bahamas; Cuba. Aug.—Oct. 


75. Aster subulatus Michx. Annual Salt- 
marsh Aster. Fig. 4356. 


Aster subulatus Michx. Fl. Bor. Am. 2: 111. 1803. 
Asier linifolius T, & G. Fl. N. A. 2: 162. 1841. Not 
L753: 


Annual, glabrous and fleshy; stem paniculately 
branched, flexuous above, 1°-6° high, slightly an- ™ 
gled, sometimes 1’ in diameter at the base, but 
usually smaller. Stem leaves linear-lanceolate, 
acute, entire, sessile by a broad or slightly clasping 
base, 2’-10’ long, 1-8” wide, those of the branches 
very small and subulate; heads numerous, 3-5” 
broad; involucre campanulate, or at length hemi- 
spheric, 2’-3” high, its bracts linear-subulate, 
green, imbricated in 3 or 4 series, the outer shorter; 
rays 20-30, purplish, scarcely exceeding the nearly 
white pappus, more numerous than the disk- 
flowers; achenes compressed, minutely pubescent. 


In salt marshes, coast of New Brunswick to Flor- 
ida. Also on salt lands Onondaga Lake, N. Y. Aug.— 
Nov. 

28 


434 COMPOSITAE. VoL. III. 


32. LEUCELENE Greene, Pittonia 3: 147. 1896. 


Low perennial herbs, with much branched leafy stems, sessile, rather rigid, narrow, 
entire leaves and small heads of both tubular and radiate white flowers, solitary at the ends 
of the numerous slender branchlets, involucre turbinate, its bracts well imbricated. Disk- 
flowers perfect, their corollas white, tubular-funnelform, 5-toothed. Ray-flowers numerous, 
white, or drying red to rose, pistillate. Style appendages acutish. Achenes elongated, flat- 
tened, hispidulous. Pappus a single series of slender rough white bristles. [Greek, referring 
to the white disk.] 


Two or three species, natives of the central and southwestern States and Mexico, the follow- 
ing typical. 


Wx, Leucelene ericoides (Torr.) Greene. Rose 
Heath Aster. Fig. 4357. 


Inula (?) ericoides Torr. Ann. Lyc. N. Y. 2: 212. 1828, 
Aster ericaefolius Rothrock, Bot. Gaz. 2: 70. 1877. 
Leucelene ericoides Greene, Pittonia 3: 148. 18096. 


Stems tufted from deep woody roots, corymbosely 
much branched, 3’-12’ high, hispid or scabrous, the 
branches erect or diffuse. Leaves hispid-ciliate, erect, 
or slightly spreading, obtusish or mucronulate, the 
lower and basal ones spatulate, 3-6” long, tapering 
into short petioles, the upper sessile, linear or linear- 
spatulate; heads terminating the branches, 5-8” 
broad; involucre broadly turbinate, its bracts lanceo- 
late, appressed, scarious-margined, imbricated in 3 
or 4 series; rays I2-15, white to rose, 2’—4” long. 


In dry soil, western Nebraska to Kansas, Texas and 
New Mexico. May—Aug. 


33. BRACHYACTIS Ledeb. Fl. Ross. 2: 495. 1846. 


Annual, nearly glabrous, somewhat fleshy herbs, with narrow chiefly entire leaves, and 
small racemose or racemose-paniculate heads of tubular, or also radiate purplish flowers. 
Involucre campanulate. Central flowers of the head few, perfect, their narrow corollas 
4-5-toothed; outer flowers pistillate, usually in 2 series or more, and more numerous than 
the perfect ones; style-appendages lanceolate; rays very short, or none. Achenes 2-3-nerved, 
slender, appressed-pubescent. Pappus a single series of nearly white bristles. [Greek, short 
rays. | 

About 5 species, natives of western North America and northern Asia. Type species: Brachy- 
actis ciliata Ledeb. 


1. Brachyactis angista (Lindl.) Britton. Rayless Aster. Fig. 4358. 


ee angustum Lindl.; Hook. Fl. Bor. Am. 2: 15. 
1834. 

Aster angustus T.& G. Fl. N. A. 2: 162. 1841. ; 
Brachyactis angusta Britton, in Britt. & Brown; Ill. FI. 

3: 383. 1808. : 

Stem usually sparsely pubescent, at least above, 
racemosely or rarely paniculately branched, 6-24’ 
high, striate, at least when dry. Leaves linear, fleshy, 
ciliate on the margins, acutish, entire, sessile by a 
rather broad base, the basal (when present) spatu- 
late; heads 4-6” broad, racemose on the ascending 
branches, or terminating them; involucre campanu- 
late or nearly hemispheric, 2’—3” high, its bracts 
linear or linear-oblong, somewhat foliaceous, green, 
acute or acutish, imbricated in 2 or 3 series, glabrous 
or slightly ciliate, nearly equal; rays none, or rudi- 
mentary; pappus soft and copious. 

In wet saline soil, or sometimes in waste places, Min- 
nesota to Saskatchewan, Utah and Colorado, and along 
the St. Lawrence River in Quebec. Found also about 
Chicago. July—Sept. 

Brachyactis frondésa (Nutt.) A. Gray, of the Rocky 
Mountain region, differing by bluntly pointed leaves and 
oblong or oblanceolate involucral bracts, has been found 
on Prince Edward Island. 


GENUS 34. THISTLE FAMILY. 435 


34. MACHAERANTHERA Nees, Gen. & Sp. Ast. 224. 1832. 


Annual, biennial or perennial branched herbs, with leafy stems, alternate, mostly serrate 
or pinnatifid leaves, the teeth or lobes usually bristle-tipped, and large heads of both tubular 
and radiate flowers. Involucre of numerous series of imbricated canescent or glandular 
bracts with herbaceous or foliaceous spreading or appressed tips. Receptacle alveolate, the 
alveoli usually toothed or lacerate. Ray-flowers numerous, violet to red or purple, pistillate. 
Disk-flowers perfect, their corollas tubular, 5-lobed, yellow, changing to red or brown; 
anthers exserted, appendaged at the tip, rounded at the base; style-appendages subulate to 
lanceolate. Achenes turbinate, narrowed below, pubescent. Pappus of numrous stiff, rough 
unequal bristles. [Greek, sickle-anther. | 

About 15 species, natives of western North America. Type species: Machaeranthera tanaceti- 
folia (H.B.K.) Nees. 


Annual or biennial ; leaves pinnatifid. 1. M. tanacetifolia. 
Perennial or biennial ; leaves sharply serrate. 2. M. sessiliflora. 


1. Machaeranthera tanacetifolia (H.B.K.) 
Nees. Tansy Aster. Dagger-flower. 
Fig. 4359. 
Aster tanacetifolius H.B.K. Nov. Gen. Sp. 4: 95. 1820. 
M. tanacetifolia Nees, Gen. & Sp. Ast. 225. 1832. 


Annual or biennial; stem glandular-pubescent, 
often viscid, densely leafy, much branched and 
bushy, 1°-2° high. Leaves sessile or short-petioled, 
pubescent, the lowest 17-3’ long, 2-3-pinnatifid, their . 
lobes linear or oblong, acute or mucronate, the up- 
per pinnatifid, those of the branches sometimes en- 
tire; heads numerous, corymbose-paniculate, 1’—2’ 
broad; involucre hemispheric, 4”-6” high, its bracts 
linear, glandular, imbricated in 5-7 series, their green 
tips very squarrose; rays 15-25, violet-purple, 5-8” 
long, pappus copious, tawny; achenes villous. 

In dry soil, South Dakota to Nebraska, Texas, Mex- 
ico, Montana and California. June—Aug. 


VAN, 


Sy, 


2. Machaeranthera sessiliflora (Nutt. ) 
Greene. Viscid Aster. Fig. 4360. 
Dieteria sessilifora Nutt. Trans. Am. Phil. Soc. Tao Oe 

1840. 

M. sessiliflora Greene, Pittonia 3: 60. 1896. 

Stem usually stout, finely rough-pubescent or ca- 
nescent, branched, and viscid-glandular above, 1°-2° 
high. Leaves lanceolate, linear, or the lowest spatu- 
late, sessile, somewhat viscid, sharply incised-dentate, 
the larger 17-3’ long, the teeth bristle-tipped; heads 
numerous, racemose, or corymbose above, 17-13’ 
broad, the lower often nearly sessile; involucre broad- 
ly turbinate or hemispheric, 4’-6” high, its bracts 
acute, imbricated in 6-10 series, their tips strongly 
squarrose; rays numerous, violet, 4”-6” long; pap- 
pus copious; achenes narrow, appressed-pubescent. 


In dry soil, central and western Nebraska and Colo- 
rado. July—Oct. 


A Kansas plant differs from this species by having 
acute appressed tips to the involucral bracts. 


35. ERIGERON L. Sp. Pl. 863. 1753. 


Branching or scapose herbs, with alternate or basal leaves, and corymbose, paniculate or 
solitary, peduncled heads of both tubular and radiate (rarely all tubular) flowers. Involucre 
hemispheric, cylindric or campanulate, its bracts narrow, nearly equal, mostly imbricated in 
but 1 or 2 series. Receptacle nearly flat, usually naked. Ray-flowers, in our species, white, 
violet or purple, pistillate. Disk-flowers yellow, tubular, perfect, their corollas mostly 5-lobed. 
Anthers obtuse and entire at the base. Style-branches more or less flattened, their appendages 
short, mostly rounded or obtuse. Achenes flattened, usually 2-nerved. Pappus-bristles fragile, 
slender, scabrous or denticulate, in I series, or often an additional outer short series. [ Greek, 
early-old, alluding to the early hoary pappus.] 


436 COMPOSITAE. | Vou. IIL. 


A genus of 130 species or more, of wide geographic distribution, most abundant in the New 
World. In addition to the following, many others occur in the southern and western parts of North 
America. Type species: Erigeron acris L. 

* Rays long, narrow, usually equalling or longer than the diameter of the disk. 
7 Stem 110’ high, simple, leafy ; head solitary ; involucre woolly. 1. E. uniflorus. 
TT Stem 4’-3° high, leafy, usually branched ; involucre hirsute or glabrous. 
1. Roots perennial, thick and woody. 
Heads 1’-2’ broad ; leaves lanceolate, ovate, oblong, or spatulate. 
Rays violet or purple; stem leaves ovate, lanceolate, or oblong. 
Stem leaves linear-lanceolate, the upper much smaller, 


2. E. asper. 

Stem leaves ovate-lanceolate, the upper little smaller. 3. E. subtrinervis. 

Rays mostly white ; stem leaves linear or linear-oblong. 4. E. caespitosus. 
Heads %’-1’ broad; leaves linear. 

Plants hirsute or canescent ; pappus double; western species. 

Stem hirsute ; achenes pubescent ; flowers white. 5. E. pumilus. 

Stem appressed-canescent ; achenes glabrous ; flowers purple or white. 6. E. canus. 
Plant nearly glabrous; pappus simple; northern. 7. E. hyssopifolius. 

2. Perennial by decumbent, rooting stems or stolons. 8. E. flagellaris. 


3- Roots annual or biennial, fibrous; plants often perennial by offsets. 
Heads 1’-1%’ broad, few; stem simple; eastern. 9. E. pulchellus. 
Heads ¥%’—1’ broad, numerous; stem branched. 
Rays 100-150, narrow, mostly purple or violet. 


Pappus simple; plant erect, corymbosely branched. 10. E. philadelphicus. 

Pappus double; plant diffusely branched, western. 11. E. divergens. 
Rays much less numerous, purplish or white. : 

Plants 6’—12’ high, diffuse, western ; pappus simple. 12. E. Bellidiastrum. 


Plants 6’—3° high, erect, branched ; pappus double. 
Pappus of the ray-flowers and disk-flowers unlike. 


Stem leaves lanceolate, nearly all sharply serrate. 13. E. annuus. 
Stem leaves linear-lanceolate or oblong, nearly all entire. 14. E. ramosus. 
Pappus of the ray-flowers and disk-flowers alike; plant hirsute. 15. E. tenuis. 


ttt Stem leafless or nearly so ; heads %4’ broad, corymbose. 16. E. vernus. 
** Rays inconspicuous or short; a row of tubular pistillate flowers inside ee row of rays. 
17. &. acrts. 


1. Erigeron uniflorus L. Arctic Erigeron. Fig. 4361. 
Erigeron uniflorus L. Sp. Pl. 864. 1753. 


Perennial by short branching rootstocks; stems slender, 
single or,tufted, more or less pubescent, simple, erect, I-10) 
high. Basal leaves petioled, spatulate, obtuse, entire, 1’-2 
long; stem leaves sessile, lanceolate or linear-lanceolate, en- 
tire, acute or obtuse; head solitary, peduncled, 3’-1’ broad; 
rays about 100, purple or purplish, 2’-4” long; involucre 
hemispheric, its bracts linear-lanceolate, acute, copiously 
woolly; pappus simple. 

Labrador and Arctic America to Alaska, south in the Rocky 
Mountains to Colorado and in the Sierra Nevada to California. 
Also in Europe. Summer. 


2. Erigeron asper Nutt. Rough Erigeron. 
Fig. 4362. 


Erigeron asper Nutt. Gen. 2: 147. 1818. 
Erigeron glabellus Nutt. loc. cit. 1818. 


Perennial by a woody root; stem simple, or 
branched above, more or less pubescent, sometimes 
hirsute, 6-24’ high. Leaves glabrous, pubescent or 
ciliate, entire, the basal ones spatulate, obtuse, 2’-4’ 
long, 3-1’ wide, narrowed into margined petioles; 
stem leaves oblong-lanceolate or linear-lanceolate, 
obtuse or acute, the upper smaller; heads several or 
solitary, slender-peduncled, 1’-2’ broad; involucre 
hemispheric, its bracts linear, acute, hirsute or pu- 
bescent; rays 100-150, very narrow, violet, purple, 
or nearly white, 4’-7” long; pappus double, the 
outer row of bristles much shorter than the inner. 


In dry soil, Minnesota to Wisconsin, Nebraska, Manitoba, Utah and New Mexico. Races 
differ in pubescence. June—Sept. 


GENUS 35. THISTLE FAMILY. 437 


W 


3. Erigeron subtrinérvis Rydberg. Three- 
nerved Fleabane. Fig. 4363. 


Erigeron glabellus var. mollis A. Gray, Proc. Acad. 
Phila. 1863: 64. 1864. Not &. mollis D. Don. 


Erigeron subtrinervis Rydberg, Mem. Torr. Club 5: 
328. 18094. 


Similar to the preceding species, perennial by 
a woody root, finely pubescent all over; stems 
leafy to the inflorescence. Leaves entire, thin, the 
basal and lower ones oblanceolate to oblong, ob- 
tuse or acute, petioled, the upper lanceolate or 
ovate-lanceolate, sessile or somewhat clasping, 
acute, rather distinctly 3-nerved; heads 17-13’ 
broad, corymbose, or rarely solitary; involucre 
hemispheric, hirsute; rays numerous, blue to pink; 
pappus double, the outer bristles very short. 


In dry soil, South Dakota to Wyoming, Nebraska, 
Utah and New Mexico. July—Sept. 


WZ , 4. Erigeron caespitosus Nutt. Tufted 
wr Wy Erigeron. Fig. 436 
Sk WS LZ get : §- 4304. 
Zit a Diplopappus canescens Hook. Fl. Bor. Am. 2: 21. 
\ v 1834. Not E. canescens Willd. 1804. 
} it 7 Erigeron caespitosus Nutt. Trans. Am. Phil. Soc. (II) 
8 = 3207.) [Od d. 

Perennial by a deep root; stems tufted, canes- 
cent, simple, or branched above, 6-12’ high. 
Leaves canescent or pubescent, entire, the lower 
and basal ones petioled, narrowly oblanceolate 
or spatulate, obtuse or acutish, 13’ long; stem 
leaves linear or linear-oblong, acute or obtuse, 
sessile, the upper gradually shorter, heads soli- 
tary or several, short-peduncled, 1-13’ broad; in- 
volucre hemispheric, its bracts lanceolate or linear- 
oblong, acute, canescent; rays 40-60, 3-6” long, 
white or pinkish; pappus double, the outer series 
of bristles very short. 

In dry soil, Manitoba to Yukon, Nebraska (accord- 


ing to Webber), British Columbia and Colorado. 
June—Aug. 


5. Erigeron pumilus Nutt. Low Erigeron. 
Daisy. Fig. 4365. 


Erigeron pumilis Nutt. Gen. 2: 147. 1818. 


Perennial by a deep root; stems tufted, hirsute, slen- 
der, simple, or branched, 4’-10’ high. Leaves entire, 
hirsute, the lower and basal ones narrowly spatulate or 
linear, petioled, obtuse or acutish, 1-4’ long, 1-2” wide; 
stem leaves linear, sessile, ’—-23’ long, acute; heads soli- 
tary or several, 6-10” broad, short-peduncled; invo- 
lucre hemispheric, its bracts linear, acute, hirsute; rays 
50-80, white, 3’-4’” long, at length deflexed; pappus 
double, the outer row of bristles short and more or less 
intermixed with the inner; achenes pubescent. 


Dry plains, North Dakota to western Nebraska, Kansas, 
British Columbia, Colorado and Utah. May-Sept. 


COMPOSITAE. Vor. III. 


6. Erigeron canus A. Gray. Hoary Erigeron. 
Fig. 4366. 


Erigeron canus A. Gray, Mem. Am. Acad. (II) 4: 67. 1849. 


Perennial by a deep woody root, resembling the pre- 
ceding species; stems slender, erect, tufted, appressed- 
canescent, 6’—10’ high, simple, or branched above. Leaves 
narrow, entire, canescent, the basal and lower ones nar- 
rowly spatulate, petioled, 2-4’ long, the upper linear, 
sessile, acute, gradually smaller; heads solitary, or 2-4, 
peduncled, 6’-8” broad; involucre hemispheric, its bracts 
linear, acute, densely canescent; rays 40-50, purple or 
white, 2-3” long; achenes glabrous, 8-10-nerved; pap- 
pus double, the outer row of bristles rather conspicuous. 


In dry soil, South Dakota to western Nebraska, Wyoming, 


Colorado and New Mexico. June—Aug. 


7. Erigeron hyssopifolius Michx. Hyssop- 
leaved Erigeron. Fig. 4367. 


ere hyssopifolius Michx. Fl. Bor. Am. 2: 123. 
Aster premiere Wa Pursh, Fl. Am. Sept. 545. 1814. 

Perennial by slender rootstocks; stems tufted 
or single, very slender, simple or branched, gla- 
brous or very nearly so, 4-15’ high. Leaves nar- 
row, thin, the basal and lower ones oblong or 
spatulate, short-petioled, 1-12’ long, 14’-2” wide, 
the upper linear or linear-oblong, acute, usually 
numerous; heads solitary or several, slender- 
peduncled, 5”-8” broad; peduncles appressed- 
pubescent; involucre nearly cylindric at flower- 
ing time, its bracts linear-lanceolate, sparingly 
pubescent; rays 12-30, white or purplish, 3’-6” 
long; pappus simple. 


On moist cliffs, Newfoundland to northern. Ver- 
mont, Mackenzie and Lake Superior. July—Aug. 


WW 


S\ Wi 8. Erigeron flagellaris A. Gray. Run- 


ning Fleabane. Fig. 4368. 


Y 


\ Erigeron flagellaris A. Gray, Mem. Am. Acad. (11) 
\ 4: 68. 1849. 


Appressed-pubescent, sometimes densely so, 
perennial by decumbent rooting stems or sto- 
lons; root slender; stem slender, branched, 
the branches elongated. Leaves entire, the 
basal and lower ones spatulate or oblong, ob- 
tuse or acute, 1-2’ long, narrowed into long 
petioles, the upper sessile, linear or linear- 
spatulate, much smaller; peduncles solitary, 
elongated; heads about 1’ broad and ?’ high; 
involucre hemispheric, its narrow bracts pu- 
bescent; rays very numerous, white to pink; 
pappus double, the outer series of subulate 
bristles. 


In moist soil, South Dakota to Wyoming, Utah, 
western Texas and New Mexico. May-July. 


GENUS 35. THISTLE FAMILY. 439 


g. Erigeron pulchéllus Michx. Robin’s 
or Poor Robin’s Plantain. Fig. 4369. 


E. pulchellus Michx. Fl. Bor. Am. 2: 124. 1803. 
E. bellidifolius Muhl.; Willd. Sp. Pl. 3: 1958. 1804. 
Perennial by stolons and offsets, villous- 
pubescent; stems simple, slender, 10-24’ high. - 
Basal leaves tufted, spatulate or obovate, 
somewhat cuneate at the base, narrowed into 
short margined petioles, obtuse at the apex, 
1-3’ long, 2’-2’ wide, dentate or serrate; stem 
leaves sessile, partly clasping, oblong, lanceo- 
late or ovate, mostly acute, entire, or spar- 


ingly serrate; heads 1-6, slender-peduncled, i 
1-12’ broad; involucre depressed-hemispheric, 2. WY 
its bracts linear, acuminate, villous; rays nu- Su 29X 
merous, violet or purplish, 4’—7” long; achenes AN ja 
nearly BOETOE DaUDUs simple. . \) 

On hills and banks, Maine to Ontario and Min- 24 AS 


nesota, Kansas, Florida and Louisiana. Recorded Zi \N 
from Quebec and Nova Scotia. Rose-petty. EU f)\X 
Robert’s-plantain. .Blue spring-daisy. April—June. / 


a 
NWA 
WINS 

WYNZs 

=A) | 


to. Erigeron philadélphicus L. Philadel- 
phia Fleabane. Skevish. Fig. 4370. 
Erigeron philadelphicus L. Sp. Pl. 863. 1753. 


Perennial by stolons and offsets, soft-pubescent 
or sometimes nearly glabrous; stems slender, 
mostly branched above, 1°-3° high. Basal and 
lower leaves spatulate or obovate, obtuse, dentate, 
1-3’ long, narrowed into short petioles; upper 
stem leaves clasping and often cordate at the 
base, obtuse or acute, dentate or entire; heads 
several or numerous, corymbose-paniculate, 5’”— 
12” broad, slender-peduncled; peduncles thick- 
ened at the summit; involucre depressed-hemi- 
spheric, its bracts linear, usually scarious-mar- 
gined; rays 100-150, 2’-4” long, light rose-purple 
to pinkish; pappus simple; achenes puberulent. 


In fields and woods, Labrador to British Columbia, 
Florida and California, but locally rare. Races differ 
in leaf-form and pubescence. Buds drooping. Sweet 
scabious. Daisy-fleabane. April—Aug. 


11. Erigeron divérgens T. & G. Spreading 
Fleabane. Fig. 4371. 
Erigeron divergens T.& G. Fl. N. A. 2: 175. 1841. 


Annual or biennial, diffusely branched, 6-15’ high, 
densely cinereous-pubescent or hirsute. Basal and 
lower leaves spatulate or oblanceolate, acute or ob- 
tuse, mostly petioled, entire, dentate or lobed, 1-2’ 
long, 2’-5”" wide, the upper sessile, linear or nar- 
rowly spatulate, usually acute, gradually smaller; 
heads slender-peduncled, 8-12” broad, usually nu- 
merous; involucre hemispheric, about 2” high, its 
bracts linear, acute, hirsute or canescent; rays about 
100, purplish, violet or nearly white, 2’-4” long; 
pappus double, the shorter outer row of bristles 
subulate. 


In moist soil, Montana to Nebraska, Texas, Mexico, 
Washington and California. April—Sept. 


COMPOSITAE. Vot. ITT. 


12. Erigeron Bellidiastrum Nutt. Western 
Daisy Fleabane. Fig. 4372. 


Erigeron Bellidiastrum Nutt. Trans. Am. Phil. Soc. (II) 7: 
307. 1841. 


Annual, much branched, 6-15’ high, cinereous-pubes- 
cent throughout. Leaves entire, linear-spatulate, the 
lower and basal petioled, 1’-12’ long, the upper sessile 
and smaller; heads several or numerous, 5-8” broad; 
involucre hemispheric, about 2” high, its bracts linear, 
hirsute, acute; rays 30-65, purplish, about 2” long; 
pappus a single series of capillary bristles. 


In moist soil, South Dakota to Nebraska, Kansas, Texas, 
Wyoming and Arizona. May-July. 


| 


13. Erigeron Annuus (L.) Pers. Sweet Scabious. White-top. Fig. 4373. 


Aster annuus L. Sp. Pl. 875. 1753- 
Erigeron annuus Pers. Syn. 2: 431. 1807. 


Annual, sparingly pubescent with spreading hairs ; 
stem erect, corymbosely branched, 1°-4° high. Leaves 
thin, the lower and basal ones ovate or ovate-lan- 
ceolate, mostly obtuse, petioled, usually coarsely 
dentate, 2’-6’ long, 1-3’ wide, the upper sessile or 
short-petioled, lanceolate, oblong, or linear-lanceo— 
late, acute or acuminate, mostly sharply dentate in 
the middle, those of the branches narrower and 
often entire; heads rather numerous, 5-7” broad, 
mostly short-peduncled; bracts of the hemispheric 
involucre somewhat hispid; rays 40-70, linear, 
white, or commonly tinged with purple, 2’-4” long; 
pappus double, the inner a series of slender fragile 
deciduous bristles, often wanting in the ray-flowers, 
the outer a persistent series of short, partly united, 
slender scales. 


ran 


In fields, Nova Scotia to Manitoba, Georgia, Kentucky and Missouri. Naturalized in Bermuda 


and in continental Europe. Daisy-fleabane. Lace-buttons. May—Nov. 

NN! - & = 

Ny) 14. Erigeron ramosus (Walt.) B.S.P. 
Daisy Fleabane. Fig. 4374. 


Doronicum ramosum Walt. Fl. Car. 205. 1788. 
E. strigosus Muhl.; Willd. Sp. Pl. 3: 1956. 1804. 
’ Stenactis Beyrichii F. & M. Index Sem. Hort. Petrop. 
5: 1838. 
Erigeron ramosus B.S.P. Prel. Cat. N. Y. 27. 1888. 
Erigeron ramosus Beyrichii Smith & Pond, Bot. Surv. 
Neb. 2: 11. 1893. 


Resembles the preceding species, but is usually 
lower and the pubescence more appressed. Stem 
leaves linear-oblong or linear-lanceolate, nearly 
all of them entire; basal and lowest spatulate or 
oblong, usually serrate; bracts of the invclucre 
glabrous or nearly so; pappus similar to that of 
the preceding; rays white, or sometimes purplish, 
occasionally minute or wanting. 

In fields, Nova Scotia to British Columbia, Florida, 


Louisiana, Texas and California. Naturalized in 
Europe. White-top. May-—Nov. 


GENUS 35. THISTLE FAMILY. 


15. Erigeron tenuis T. & G. Slender Rough 
Fleabane. Fig. 4375. 


Erigeron tenuis T. & G. Fl. N. A. 2: 175. 1841. 


Annual or biennial, branched from the base and some- 
times also above, strigose-pubescent; stems slender, erect 
or ascending, 1° high or less. Basai leaves obovate to 
spatulate, 1-23’ long, 3’-6” wide, usually toothed; stem 
leaves linear or linear-oblong, toothed or entire; heads 
several or solitary, slender-peduncled, 7’-10” broad; in- 
volucre 2-3” high, its linear bracts glabrous or spar- 
ingly pubescent; rays white or purplish, numerous; 
pappus of ray-flowers and disk-flowers alike, of few 
long bristles and short small scales. 


Moist prairies and plains, Missouri to Texas and Louis- 
iana. April-May. 


WY 
AST, 


16. Erigeron vérnus (L.) T. & G. Early 


Fleabane. Fig. 4376. 
Aster vernus L. Sp. Pl. 876. 1753. 
E. nudicaulis Michx. Fl. Bor. Am. 2: 124. 1803. 
Erigeron vernus T. & G. Fl. N. A. 2: 176. 1841. 
Perennial by stolons and offsets; stem slender, 
3 
4 


simple or branched above, glabrous, or the 
branches pubescent, 1°-23° high. Leaves mainly 
in a basal rosette, glabrous, obovate, oval or spat- 
ulate, obtuse, repand-denticulate or entire, 2-4’ 
long, narrowed into margined petioles; stem 
leaves mostly reduced to subulate-lanceolate 
scales, the lowest sometimes spatulate or oblong; 
heads not numerous, corymbose, peduncled, about 
5” broad; involucre hemispheric, its bracts linear- 
subulate; rays 20-30, white or pink, 2’-3” long; 


é 
oh pappus simple; achenes usually 4-nerved. 
P AN eccei In marshes and moist soil, Virginia to Florida and 
Nx Louisiana. April-May. 


17. Erigeron acris L. Blue or Bitter Flea- 
bane. Fig. 4377. 


Erigeron acris L. Sp. Pl. 863. 1753. 

Erigeron Droebachianus O. F. Mueller, Fl. Dan. pl. 874. 
1782. 

Erigeron acris Droebachianus Blytt, Norg. Fl. 1: 562. 
1861. ’ 

Erigeron acris debilis A. Gray, Syn. Fl. 1: Part 2, 220. 
1884. 


Biennial or perennial; stem hirsute-pubescent or 
glabrate, slender, simple, or branched above, 6’-2° 
high. Leaves pubescent or glabrous, entire, the 
basal and lower ones spatulate, mostly obtuse, 1’-3’ 
long, petioled, those of the stem mostly oblong or 
oblanceolate, obtuse or acutish, sessile, shorter; 
heads several or numerous, racemose or paniculate, 
peduncled, 5’-6” broad; involucre hemispheric, its 
bracts linear, hirsute to glabrous; rays numerous, 
purple, equalling or slightly exceeding the brownish 
pappus; tubular pistillate flowers filiform, numer- 
ous; pappus simple or nearly so, copious. 


Labrador to Alaska, Maine, Ontario, south in the 
Rocky Mountains to Colorado and Utah. Also in Eu- 
rope and Asia. Races differ in size, pubescence and 
length of rays. July—Aug. 


WES 


Sen) 
: i 


442 COMPOSITAE. Vo. III. 


36. LEPTILON Raf. Am. Month. Mag. 2: 268. 1818. 
[CaENnotus Raf. Fl. Tell. 2: 50. 1836.] 


Annual or biennial herbs, with small racemose thyrsoid or panicled heads of white 
flowers, the rays small, usually shorter than the diameter of the disk, or none. Involucre 
mostly campanulate, its narrow bracts in 2 or 3 series. Receptacle naked. Ray-flowers 
pistillate; disk-flowers perfect, their corollas usually 4-lobed or 4-toothed, the anthers obtuse 
at the base; style-branches somewhat flattened, their appendages short. Achenes flattened. 
Pappus of numerous simple fragile bristles in 1 series. [Greek, referring to the small heads.] 


About 20 species, natives of America and Asia. Besides the following, 2 or 3 others occur in 
the southwestern United States and one is naturalized from Asia on the southern Atlantic and 
Pacific coast. Type species: Leptilon divaricatum (Michx.) Raf. 


1. Leptilon canadénse (L.) Britton. Horse-weed. Canada Fleabane. 
Fig. 4378. 
Erigeron canadensis L. Sp. Pl. 863. 1753- 


Leptilon canadense Britton, in Britt. & Brown, IIl. 
BIS3eh3or-) “E898: 


Stem hispid-pubescent or glabrate, 3’-10° high, 
the larger plants paniculately much branched. 
Leaves usually pubescent or ciliate, the basal and 
lower spatulate, petioled, incised, dentate or en- 
tire, 1-4’ long, obtuse or acutish, those of the 
stem linear and mainly entire; heads usually very 
numerous; about 2” broad; involucre campanu- 
late, 1-13” high, its bracts linear, acute, gla- 
brate, the outer shorter; rays numerous, white, 
shorter than the pappus and mostly shorter than 
their tubes. 


\\\ 

In fields and waste places, a common weed through- 
out North America except the extreme north. Widely 
distributed as a weed in the Old World, the West 
Indies and South America. Butter-weed. Prideweed. 
Fireweed. Blood-staunch. Cow’s-, mare’s- or colt’s- 
tail. Hogweed. Bitter-weed. June—Nov. 


2. Leptilon divaricatum (Michx.) Raf. Low 
Horse-weed. Purple Horse-weed. Dwarf 


Fleabane. Fig. 4379. 
Erigeron divaricatus Michx. Fl. Bor. Am. 2: 123. 1803. 
Leptilon divaricatum Raf. Am. Month. Mag. 2: 268. 1818. 


Stem diffusely much branched, 3’-12’ high, pubescent 
or hirsute. Leaves all linear or subulate, entire, 4-12” 
long, about 2” wide, the uppermost minute; heads nu- 
merous, about 2” broad; involucre campanulate, 1” 
high, its bracts linear, acute, pubescent, the outer 
shorter; rays purplish, shorter than their tubes. 


In sandy soil, especially along rivers, Indiana to Minne- 
sota, Tennessee, Louisiana, Nebraska and Texas. June—Oct. 


GENUS 37. THISTLE FAMILY. 443 


37. DOELLINGERIA Nees, Gen. & Sp. Ast. 177. 1832. 


Perennial leafy herbs, the lower leaves reduced to scales or sheaths, the upper large, 
mostly entire, acute or acuminate in our species. Basal leaves none. Heads corymbose, 
usually numerous; involucre campanulate to hemispheric, its bracts 1mbricated in several 
series, appressed, thin, sometimes scarious-margined, their tips not herbaceous nor foliaceous. 
Receptacle foveolate. Ray-flowers white, pistillate, not very numerous. Disk-flowers per- 
fect, the corolla with a slender tube abruptly expanded into a campanulate 5-lobed limb, 
white to greenish in our species. Anthers obtuse at the base; style-appendages ovate to subu- 
late (rarely obtuse). Achenes obovoid, glabrous, or pubescent. Pappus double, the outer 
series of numerous short bristles or scales, the inner series of long capillary bristles, some 
or all of which have thickened tips. [In honor of Th. Dollinger, botanical explorer.] 


About 5 species, natives of eastern North America. Type species: Doellingeria umbellata 
.(Mill.) Nees. 


Leaves lanceolate to ovate; heads mostly numerous. 


Leaves lanceolate to oblong-lanceolate, acuminate. vty QD), umbellata. 
Leaves ovate to ovate-lanceolate, acute. 2D! humilis. 
Leaves, at least the lower, obovate ; heads commonly few. 3. D.infirma. 


1. Doellingeria umbellata (Mill.) Nees. Tall Flat-top White Aster. Fig. 4380. 


Aster umbellatus Mill. Gard. Dict. Ed. 8, No. 22. 1768. 
Diplopappus umbellatus Hook. Fl. Bor. Am. 2:22. 1834. 
D. umbellata Nees, Gen. & Sp. Ast. 178. 1832. 

Aster umbellatus var. pubens A. Gray, Syn. Fl. 1: Part 

ZO 7 loots 
D. pubens Rydb. Bull. Torr. Club 37: 147. 1910. 

Stem glabrous or pubescent above, striate, corym- 
bosely branched at the summit, 1°-8° high. Leaves 
lanceolate to oblong-lanceolate, ascending, glabrous 
above, usually pubescent at least on the veins be- 
neath, membranous, acuminate at the apex, nar- 
rowed at the base into short petioles, or the upper- 
most sessile, hispid-margined, those of the stem 
5-6’ long, 3’-1’ wide; heads numerous, 6’—10” broad, 
in terminal compound corymbs; involucre broadly 
campanulate or hemispheric, 13-2” high, its bracts 
lanceolate, usually pubescent or ciliate, acutish or 
obtuse, imbricated in 3 or 4 series, the outer shorter ; 
rays 10-15, white; style-appendages ovate, acute; 
pappus nearly white; achenes nerved, slightly pu- 
bescent. 


In moist soil, Newfoundland to Georgia, Saskatche- 
wan, Iowa and Michigan. July—Oct. 


2. Doellingeria htmilis ( Willd.) Britton. 
Broad-leaved Flat-top White Aster. 
Fig. 4381. 


Aster humilis Willd. Sp. Pl. 3: 2038. 1804. 

D. amygdalina Nees, Gen. & Sp. Ast. 179. 1832. 

Aster umbellatus var. latifolius A. Gray, Syn. FI. 1: 
Part 2, 197. 1884. 

Doellingeria humilis Britton, in Britt. & Brown, Ill. FI. 
3: 392. 1808. 


Similar to the preceding species, usually lower, 
seldom over 4° high; stem striate, corymbosely 
branched above, glabrous, or somewhat pubescent. 
Leaves ovate to ovate-lanceolate, rather firm, ses- 
sile, or the lower very short-petioled, acute or short- 
acuminate at the apex, narrowed, or sometimes 
rounded at the base, rough-margined, those of the 
stem 1-3’ long, 4’-2’ wide, inflorescence as in D. 
umbellatus; pappus dirty white; achenes somewhat 
pubescent. 


; 4 In moist soil, eastern Massachusetts to New Jersey, 
Pennsylvania, Florida and Texas. Perhaps a broad-leaved race of the preceding species. July—Sept. 


444 COMPOSITAE. | Vou. IIT. 


3. Doellingeria infirma (Michx.) 
Greene. Cornel-leaved Aster. 
Fig. 4382. 
Aster infirmus Michx. Fl. Bor. Am. 2: 109. 1803. 
Diplopappus cornifolius Less.; Darl. Fl. Cestr. 
474. 1837. 
D. infirma Greene, Pittonia 3: 52. 1896. 

Stem slender, glabrous, or roughish above, 
sparingly branched at the summit, terete, 14°-3° 
high. Leaves entire, hispid-margined, glabrous 
above, sparingly hispid on the veins beneath, 
the lower obovate, small, obtuse, usually ses- 
sile, the upper larger, oblong-lanceolate, acute, 
2’-5’ long, 1-12’ wide; heads few, about 1 
broad, in a divergently branched terminal co- 
rymbose cluster; involucre broadly campanu- 
late, 2-3” high, its bracts oblong-lanceolate, 
obtuse, sparingly pubescent, imbricated in about 
4 series, the outer much shorter; rays 8-15, 
white; style-appendages subulate; pappus 
tawny; achenes nerved, glabrous. 

\\, In dry, usually rocky soil, Massachusetts to 
Se New York, Georgia, Tennessee and Alabama. 
Aug.—Sept. 


38. IONACTIS Greene, Pittonia 3: 245. 1897. 


Low, mostly. branching, perennial herbs with numerous narrow, entire, I-nerved leaves, 
and rather large and showy heads of tubular and radiate flowers terminating the stem and 
branches. Basal leaves none. Involucral bracts coriaceous, imbricated in several series, 
appressed, their tips not herbaceous. Ray-flowers normally violet, pistillate. Disk-flowers 
perfect, the corolla with a campanulate limb. Achenes villous. Pappus double, the inner 
series of long capillary bristles, the outer much shorter. [Greek, violet rays.] 

Three known species, natives of North America, the following typical. 


Aster linartifolius L. Sp. Pl. 874. 1753. ( f y, 

Diplo linariifolius Hook. Fl. Bor. Am. 2: 21. 1834. Ni VR 
iplopappus linartifolius Hoo or. Am 21. 1834 a aS wy 
Stems tufted, stiff, puberulent or scabrous, very leafy, WW ry WN 

corymbosely branched above, 62° high. Leaves linear ALIN 

usually ciliolate on the margins, mucronulate, 9”-18” 

long, 1-2” wide, sessile, those of the branches much 

nating the branchlets, about 1’ broad; involucre broadly 

turbinate, its bracts linear-lanceolate, keeled, green on 

inner obtuse, the outer usually acute; rays 10-15, violet, 

rarely white, 4’-5” long, entire, or their tips dentate, or 

outer ones setose; achenes silky. 
In dry or rocky soil, Maine to Florida, west to Minne- 

land, New Brunswick and Quebec, apparently erroneously. 

Sandpaper- or pine-starwort. July—Oct. 

Dioecious shrubs or herbs, with alternate leaves, and paniculate or corymbose heads of 
tubular flowers. Involucre campanulate in our species, its bracts imbricated in several series, 
flowers slender, that of the staminate tubular, 5-lobed. Anthers obtuse and entire at the 
base. Style-branches narrow or subulate, those of the fertile flowers smooth, exserted, those 
or less compressed, ribbed. Pappus of the fertile flowers copious, capillary, that of the 
sterile flowers short. [Named for Bacchus; originally applied to some different shrubs. ] 
18 others occur in the southern and western United States. Type species: Baccharis ivifolia L. 
Shrubs ; pappus of fertile flowers in 1 or 2 series. 

I 
Leaves deltoid-obovate, or oblong, the lower coarsely dentate. 2. B. halimifolia. 
Leaves linear or linear-lanceolate, mostly acute, sparingly dentate. 3. B. neglecta. 
4 


1. Ionactis linariifélius (L.) Greene. Stiff or Savory-leaved Aster. Fig. 4383. 
Tonactis linartifolius Greene, Pittonia 3: 245. 1897. cy 
or spatulate, spreading, I-nerved, rigid, entire, rough, 
smaller and somewhat appressed; heads several, termi- 
the back, appressed, imbricated in 4 or 5 series, the 
even laciniate; pappus bristles in 2 series, tawny, the 
sota, Mississippi and Texas. Recorded from Newfound- 
39. BACCHARIS L. Sp. Pl, 860 1753. 
the outer shorter. Receptacle flat, naked, commonly foveolate. Corolla of the pistillate 
of the sterile flowers rudimentary, tipped with an ovate pubescent appendage. Achenes more 
About 300 species, all American, most abundant in South America. Besides the following, some 
Leaves oblong, or lance-oblong, mostly obtuse, sparingly dentate. . B. salicina. 
Herbaceous, from a woody base; pappus of fertile flowers in several series. . B. Wrightit. 


GENUS 30. THISTLE FAMILY 445 


1. Baccharis salicina T. & G. Willow Bac- 
charis. Fig. 4384. 


Baccharis salicina T. & G. Fl. N. A. 2: 258. 1841. 


A glabrous glutinous much-branched shrub, 3°-6° 
high, the branches ascending. Leaves firm, oblong- 
lanceolate or somewhat oblanceolate, more or less 
conspicuously 3-nerved, mostly obtuse at the apex, 
narrowed into a cuneate subsessile base, 1’-12’ long, 
2”-6” wide, sparingly repand-dentate, or entire; 
heads in peduncled clusters of 1-7, the involucre of 
both sterile and fertile ones campanulate, 23-3” 
high, its bracts ovate or ovate-lanceolate, acute or 
subacute; pappus usually but a single series of nearly 
white capillary bristles. 


Western Kansas and eastern Colorado to Texas and 
New Mexico. May-July. 


2. Baccharis halimifolia L. Groundsel-tree or 
-bush. Pencil-tree.. Fig. 4385. 
Baccharis halimifolia L. Sp. Pl. 860. 1753. 


A branching glabrous shrub, 3°-10° high, the branch- 
lets angled, sometimes minutely scurfy. Leaves thick, 
those of the stem and larger branches obovate or del- 
toid-obovate, obtuse, petioled, coarsely angular-dentate, 
1-3’ long, 4’-2’ wide, those of the branchlets oblanceo- 
late, short-petioled or sessile, entire, or few-toothed 
toward the apex; heads in peduncled clusters of 1-5, 
those of the sterile plant nearly globose when young, 
the bracts of the involucre oblong-ovate, obtuse, gluti- 
nous, appressed, the inner ones of the pistillate heads 
lanceolate, acute or acutish; fertile pappus bright white, 
3-4” long, of 1-2 series of capillary bristles, much ex- 
ceeding the involucre. 

Along salt marshes and tidal rivers, extending beyond 
saline influence, Massachusetts to Florida and Texas. Ba- 
hamas; Cuba. The white pappus is very conspicuous in 
<a Cotton-seed tree. Ploughman’s-spikenard. Sept.— 

OV. 

Baccharis glomerulifldra Pers., which has larger heads 
glomerate in the axils of the upper leaves, is doubtfully re- 
ported from southern Virginia, but occurs along the coast 
from North Carolina to Florida, and in Bermuda. 


Wan ' ND: me y 
a % ~ , NY in) \ Ny A \ Y Le 
3. Baccharis neglécta Britton. Linear- WAN eZ 
; : \ WO hy x GP 
leaved Baccharis. Fig. 4386. WA 


= 
= 


Baccharis neglecta Britton, in Britt. & Brown, Ill. Fl. 
3: 304. 1808. 


A much-branched, glabrous or slightly glutinous 
shrub, 3° high or more, the branches paniculate, 
slender, ascending. Leaves narrowly linear to 
linear-lanceolate, faintly 3-nerved, acute, or the 
lower subobtuse at the apex, gradually attenuate 
into a nearly sessile base, 1’—3’ long, 1-3” wide, 
entire, or remotely dentate or denticulate, green 
in drying; heads in short-peduncled clusters; in- 
volucre of both kinds of heads .campanulate, 2” 
high, its outer bracts ovate, acute or somewhat 
obtuse, the inner lanceolate, acuminate; pappus 
of the fertile flowers a single series of capillary 
dull-white bristles. 


Nebraska to Texas and North Mexico. July—Sept. 


; COMPOSITAE. Vo. ITT. 
Y 
A ‘ Raecmies . 
We v= 4. Baccharis Wrightii A. Gray. Wright's 
5A Baccharis. Fig. 4387. 
Baccharis Wrightwi A. Gray, Pl. Wright. 1: 101. 1852. 


y, Herbaceous from a thick woody base, much 
\ branched, glabrous, not glutinous, 1°-3° high, the 
/ branches straight, nearly erect, slender, striate. 
i \ Y Leaves linear, sessile, I-nerved, entire, 3-12” long, 

\ 4’"-1” wide; heads solitary at the ends of the 
J branches, 5-6” broad; involucre of the sterile 


heads hemispheric, about 3” high, that of the fertile 
) ) ones somewhat campanulate and longer; bracts of 
both involucres lanceolate, acuminate, with scarious 
margins and a green back; pappus of the fertile 


flowers of several series of tawny or purplish capil- 
lary bristles. 


Western Kansas and Colorado to Texas, Arizona and 
Chihuahua. April—July. 


40: .FILAGO Loeilj; Lo Sp: Pl.9277" 1753: 
[Evax Gaertn. Fr. & Sem. 2: 303. pl. 165. f. 3. 1791.] 

White-woolly annual herbs, with alternate entire leaves, and small discoid clustered 
heads, usually subtended by leafy bracts. Bracts of the involucre few and scarious. Recep- 
tacle convex or elongated, chaffy, each chaffy scale subtending an achene. Outer flowers 
of the heads in several series, pistillate, fertile, their corollas filiform, minutly 2-4-dentate. 
Central flowers few, perfect, mainly sterile, their corollas tubular, 4-5-toothed. Anthers 
sagittate at the base, the auricles acuminate. Achenes compressed or terete. Pappus none. 
[Latin filum, a thread.] 


About 12 species, natives of temperate or warm regions of both the New World and the Old. 
In addition to the following, 3 others occur in the western and southwestern United States. Type 


species: Filago pygmaea L. Biz 
1. Filago prolifera (Nutt.) Britton. Filago. a yr 
Fig. 4388. CIENT, 
Evax prolifera Nutt.; DC. Prodr. 5: 459. 1836. ence \ 2 Wag 
Diaperia prolifera Nutt. Trans. Am. Phil. Soc. (II) J: Avis EY ARR 
338. 1841. N\ SY 
Filago prolifera Britton, Mem. Torr. Club 5: 329. 1894. \\ iA 
Stem simple, or branched at the base, very leafy, L SES 
2’-6’ high. Leaves spatulate, obtuse, sessile, ascend- Mas 
ing or appressed, 4’-8” long, 1’—23” wide; heads in PEK 
a sessile leafy-bracted cluster, usually subtended by I \ 
or several slender, nearly leafless branches, each ter- \) 
minated by a similar cluster, or these again prolifer- Nii y XK { 
ous; heads oblong or fusiform; receptacle convex; \\ XQ Sv 
chaff of the central sterile flowers woolly-tipped, that \ WA SAN 
of the fertile flowers scarious, mostly glabrous. S = 3 
In dry soil, Texas to western Kansas and South Da- XS we 5 \\ 
kota, west to Colorado and New Mexico. April—July. = 


41. GIFOLA Cass. Bull. Soc. Philom. 1819: 143. 1819. 
[Firaco L. Gen. Pl. Ed. 5. 1758. Not Sp. Pl. 927. 1753.] 

White-woolly herbs, closely resembling those of the preceding genus, with alternate, 
entire leaves, and small discoid glomerate heads, often subtended by leafy bracts, the clusters 
proliferous in our species. Involucre small, its bracts scarious, imbricated in several series, 
the outer usually tomentose. Receptacle subulate, cylindric or obconic, chaffy, each chaffy 
scale near its base enclosing an achene. Outermost flowers pistillate, fertile, with filiform 
corollas and no pappus, or the pappus a few rudimentary bristles; inner flowers also pistillate 
and fertile, but with a pappus of capillary scabrous bristles; central,(uppermost) flowers few, 
perfect, their corollas tubular, their pappus capillary and scabrous. Anthers sagittate at the 
base. Achenes terete or slightly compressed. [Anagram of Filago.] 


About 10 species, natives of warm and temperate regions, the following typical. In addition to 
the following, 3 Californian species are probably to be referred to this genus. 


GENUS 4I. THISTLE FAMILY. 


1. Gifola germanica (L.) Dumort. Cudweed. Cotton 
Rose. Herb Impius. Fig. 4389. 


Gnaphalium germanicum L. Sp. Pl. 857. 1753. 
Filago germanica L. Sp. Pl. Ed. 2, 1311. 1763. 
Gifola germanica Dumort. Fl. Belg. 68. 1827. 


Annual, erect, cottony, 4-18’ high, simple, or branched at 
the base, very leafy. Leaves sessile, lanceolate, linear, or 
slightly spatulate, erect or ascending, obtuse or acutish, 3-12” 
long; stem terminated by a sessile dense cluster of heads, 
usually subtended by several leafy branches terminated by 
similar clusters and these often again proliferous; heads 12-30 
in each cluster, many-flowered; involucre ovoid, light yellow, 
its bracts mainly acute. 

In dry fields, southern New York and New Jersey to Pennsylva- 


nia, West Virginia and North Carolina. Old names downweed, 
hoarwort, owl’s-crown, chafeweed, childing cudweed. May-Sept. 


42. PLUCHEA Cass. Bull. Soc. Philom. LOU. 2a LO 7 


Pubescent or glabrous herbs, or some tropical species shrubby, with alternate dentate 
leaves, and small heads of tubular flowers in terminal corymbose cymes. Involucre ovoid, 
campanulate, or nearly hemispheric, its bracts appressed, herbaceous, imbricated in several 
series. Receptacle flat, naked. Outer flowers of the head pistillate, their corollas filiform, 
3-cleft or dentate at the apex. Central flowers perfect, but mainly sterile, their corollas 5-cleft. 
Anthers sagittate at the base, the auricles caudate. Style of the perfect flowers 2-cleft or 
undivided. Achenes 4-5-angled. Pappus a single series of capillary scabrous bristles. [Named 
for the Abbé N. A. Pluche, of Paris.] 


: About 35 species, widely distributed in warm and temperate regions. In addition to the follow- 
ing, 2 or 3 other indigenous species occurs in the southern United States, and two introduced ones 
have been found in waste places in Florida. Type species: Conyza marilandica Michx. 


Perennial; leaves sessile, cordate, or clasping at the base. 1. P. foetida. 
Annual; leaves, at least those of the stem, petioled. 
Leaves short-petioled; heads about 3” high; involucral bracts densely puberulent. 
2. P.camphorata. 
Leaves slender-petioled ; heads 2”—2%” high; involucral bracts granulose, ciliate. 
3. P. petiolata. 


D 1. Pluchea foétida (L.) DC. Viscid Marsh 


Wf Fleabane. Fig. 4390. 


AW Prin RS, Ly Baccharis foetida L. Sp. Pl. 861. 1753. 
RY, Ny SUA 
SWV4 \ Ve ye < . 
SV \ ILS Baccharis viscosa Walt. Fl. Car. 202. 1788. 
= QS —_ 
. = Pluchea bifrons DC. Prodr. 5: 451. 1836. 


Pluchea foetida DC. Prodr. 5: 452. 1836. 


} Root perennial; stem simple or sparingly branch- 

1) ed at the summit, puberulent and slightly viscid, 

VY 12°-3° high. Leaves oblong, ovate or ovate-lan- 

s ceolate, closely sessile and more or less cordate- 

clasping at the base, obtuse or acute at the apex, 

sharply denticulate, pubescent or puberulent, 2-4’ 

VA long, 4-14’ wide, reticulate-veiny; clusters of 

t heads sessile, or stalked, compact, leafy-bracted ; 

VW, involucre 24-3” high, its bracts lanceolate, acute, 
gS, viscid-puberulent. 


In swamps, southern New Jersey to Florida and 
Texas, mainly near the coast. Also in the West In- 
dies. Foetid marsh-fleabane. July—Sept. 


Ale 


448 COMPOSITAE. Vor. III. 


2. Pluchea camphorata (L.) DC. Spicy or 
Salt-marsh Fleabane. Fig. 4391. 


Erigeron camphoratum L. Sp. Pl. Ed. 2, 1212. 1763. 
Conyza marilandica Michx. Fl. Bor. Am. 2: 126. 1803. 
Pluchea camphorata DC. Prodr. 5: 451. 1836. 


Root annual; stem usually branched, finely viscid- 
puberulent, or nearly glabrous, 2°-3° high, somewhat 
channeled. Leaves ovate, oblong or lanceolate, pu- 
berulent or glabrous, acute or acuminate at the 
apex, narrowed at the base, short-petioled, or the 
upper sessile but not clasping, 3’-8’ long, 1’—3’ wide, 
serrate or denticulate, not conspicuously reticulate- 
veined; heads about 3” high, rarely leafy-bracted, 
mostly in naked open corymbiform cymes; bracts 
of the involucre ovate-lanceolate, or lanceolate, 
acute, puberulent; flowers purplish; achenes pubes- 
cent. 

In salt marshes, coast of Massachusetts to Florida, 


Texas and Mexico. Called also ploughman’s-wort. 
Aug.—Oct. 


3. Pluchea petiolata Cass. Inland Marsh Flea- 
bane. Fig. 4392. 


Pluchea petiolata Cass. Dict. Sci. Nat. 42: 2. 1826. 
Pluchea foetida DC. Prodr. 5: 452. 1830. 


Similar to the preceding species, but glabrate, darker 
green, usually taller, 23°-4° high, the stem stout, rather 
strongly channelled. Leaves ovate-lanceolate to oval, 
thin, 4-10’ long, 13’-3’ wide, mostly acuminate at the 
apex, cuneate-narrowed at the base, long-petioled, ir- 
regularly serrate; petioles of the larger leaves 8-12” 
long; heads 2”-23” high, in terminal and often also 
axillary clusters; bracts of the involucre granular, cil- 
iate; achenes short-pubescent. 


In moist soil, often in woods, Maryland to Florida, Illi- 
nois, Missouri and Oklahoma. Aug.—Oct. 


, 


43. ANTENNARIA Gaertn. Fruct. & Sem. 2: AIO. pl.. 167.. 170% 


Perennial woolly dioecious or polygamo-dioecious herbs, with alternate and basal leaves, 
and small discoid many-flowered heads usually capitate, glomerate or corymbose. Involucre 
oblong, ovoid or campanulate, its bracts scarious, imbricated in several series, the outer 
shorter, usually woolly. Receptacle convex, or nearly flat, foveolate, not chaffy. Staminate 
flowers with a truncate or minutely dentate corolla, usually undivided style and scanty pappus 
of club-shaped smooth or minutely barbed bristles. Pistillate or perfect flowers with tubular 
mostly 5-toothed corollas, 2-cleft style, and copious pappus of capillary naked bristles, slightly 
united at the base, the style often crimson. Achenes oblong, terete, or slightly compressed. 
[Latin, in allusion to the fancied resemblance of the sterile pappus to insect antennae. ] 

About 50 species, natives of the north temperate zone and southern South America. In addi- 
tion to the following, many others occur in the western parts of North America. The patches of 
fertile and sterile plants are usually quite distinct, and the sterile plants of some species are much 
less abundant than the pistillate. Perfect achenes are, however, abundant, parthenogenesis being 
a known feature of this genus. Type species: Antennaria dioica (L.) Gaertn. 


GENUS 43. THISTLE FAMILY; 449 


* Pappus-bristles of sterile flowers with club-shaped or dilated tips. 
+ Plant not stoloniferous; basal leaves oblanceolate. 1. A. carpathica. 
77 Plants stoloniferous, growing in patches. 


Involucral bracts of fertile heads dark brownish green, lanceolate, acute or acuminate; plant 
1-4’ high. ; 2. A. alpina. 
Involucral bracts all light green, or light brown, with white or pinkish scarious tips. 
Basal leaves and those of the ends of stolons bright green and glabrous above from the first. 
Basal leaves small, spatulate, with only 1 prominent nerve. 3. A. canadensis. 
Basal leaves large, obovate, distinctly 3-nerved. 4. A. Parlinii. 
Basal leaves and those of the ends of stolons dull green, arachnoid at first, becoming glabrous. 
Basal leaves large, mostly 2’-5’ long, mostly 10” wide or wider, oblong to obovate, 
3-5-nerved. 
Stolons leafy only at the ends, scaly-bracted ; head solitary. 5. A. solitaria. 
Stolons leafy throughout ; heads corymbose or subcapitate. 6. A. plantaginifolia. 
Basal leaves small, 2’ long or less, spatulate to oblanceolate or obovate, mostly 1-nerved. 
Basal leaves persistently appressed silvery-silky on both sides. 7. A. microphylla. 


Basal leaves dull, the upper surface more or less persistently arachnoid. 
Basal leaves abruptly apiculate; stolons leafy throughout. 8. A. neodioica. 
Basal leaves obtuse or acute. 
Stolons leafy throughout ; western species. 


Basal leaves spatulate. g. A. aprica. 
Basai leaves obovate. 10. A. campestris. 
Stolons leafy only at the ends. 11. A. neglecta. 
** Pappus-bristles of sterile flowers not dilated, barbellate; plant about 1’ high, tufted; 
head solitary. 12. A. dimorpha. 


1. Antennaria carpathica (Wahl.) Hook. Carpathian 
Everlasting. Fig. 4393. 


Gnaphalium carpathicum Wahl. Fl. Carp. 258. pl. 3. 1814. 
Antennaria carpathica Hook. FI. Bor. Am. 1: 329. 1833. 


Plant floccose-woolly throughout, not stoloniferous; stem 
2’-10' high, simple. Basal leaves oblanceolate or oblong, ob- 
tuse or acutish at the apex, narrowed at the base into short 
petioles, 1-2’ long, 2-4’ wide; stem leaves linear, acute or 
acutish, erect, the upper gradually smaller; heads in a termi- 
nal subcapitate cluster, or rarely solitary, 23”-3” broad; invo- 
lucre 2-3” high, woolly at the base, its bracts brownish purple, 
the inner ones of the fertile heads mostly acutish, those of the 


sterile heads mainly obtuse. 


Europe and northern Asia. May-—Sept. 


In dry soil, Labrador and Anticosti to Hudson Bay. Also in 


2. Antennaria alpina (L.) Gaertn. Alpine Everlasting. Fig. 4394. 


Gnaphalium alpinum L. Sp. Pl. 856. 1753. 


seats 
LES Antennaria alpina Gaertn. Fr. & Sem. 2: 410. 1791. 
eS ta ae acorn Nutt. Trans. Am. Phil. Soc. (1]) 7: 406. 
_—_ 1841. 
y A. angustata Greene, Pittonia 3: 284. 1808. 
\\ Surculose by short stolons; stems floccose-woolly, 
C 1-4’ high. Basal leaves usually numerous, tufted, 
\ spatulate or linear-oblong, obtuse, silvery-woolly on 
\ both sides, or glabrate and green above, 4-12” long; 
NN) stem leaves linear and small, few, sessile, woolly; 
ee y heads in a terminal capitate, or seldom somewhat 


\\ | | y iN corymbose cluster, rarely solitary, 2-23” broad; in- 

\ \) volucre about 23” high, the bracts of the fertile heads 

WW dark brownish-green, the inner ones acute or acumi- 

\ nate, those of the sterile heads lighter, brownish, 
broader and obtuse; achenes glandular. 


; wae Labrador and Arctic America to Alaska and British 
NS Columbia. Western plants previously referred to this 
species prove to be distinct. April—Aug. 


29 


COMPOSITAE. Vor. IIT.’ 


3. Antennaria canadénsis Greene. Cana- 
dian Cat’s-foot. Fig. 4395. 
Antennaria canadensis Greene, Pittonia 3: 275. 1898. 


Stems 8-12’ high, slender. Basal leaves and 
those of the ends of stolons spatulate to oblan- 
ceolate, obtuse or apiculate, 1-13’ long, 6” wide 
or less, gradually tapering from above the middle 
to a long narrow base, I-nerved or with two faint 
lateral nerves, bright green and glabrous above, 
lanate beneath; stolons short, leafy, assurgent; 
stem-leaves linear-lanceolate, distant, about 1” 
wide; inflorescence capitate to racemose-corym- 
bose;’ fertile involucre about 43” high, its outer 
bracts oblong, obtuse, the inner lanceolate, acute 
to attenuate; bracts of the staminate involucre 
white-tipped. 

In dry soil, Newfoundland to Connecticut, Mani- 
toba and Michigan. May-July. 

The Old Worid Antennaria dioica (L.) Gaertn., 
with stem leaves close together and rose-colored in- 
volucral bracts, is recorded as long ago found at 
Providence, R 


4. Antennaria Parlinii Fernald. Parlin’s aie NY 
Cat’s-foot. Fig. 4396. Q 
Antennaria Parlinii Fernald, Gard. & For. 10: 284. < \ 
1897. a 
A. arnoglossa Greene, Pittonia 3: 318. 1808. ZG 
A, Parlinii arnoglossa Fernald, Proc. Bost. Soc. Nat. 
Hist. 28: 243. 1898. 
A. propinqua Greene, Pittonia 4: 83. 1899. 


More or less glandular-pubescent; stems of 
fertile plant 1°-13° tall. Leaves bright green 
and devoid of tomentum on the upper surface 
from the time of unfolding, or very slightly floc- 
cose when very young, the basal ones obovate or 
spatulate to elliptic, obtuse or acutish, gradually 
contracted into a narrow base about as long as 
the expanded part, 2’-33’ long, #’/-14’ wide; stem- 


leaves lanceolate or the lower narrowly oblong; 
heads corymbose; involucre 33’—5” high, its bracts 
———s 
a 


all lanceolate-acuminate or the outer ones linear- 
oblong and obtusish. 


Fields, hillsides and woodlands, Maine to Ontario, 
Virginia and Iowa. May-July. 


2: 
S 


7 5. Antennaria solitaria Rydb. Single- 
headed Cat’s-foot. Fig. 4397. 


Antennaria plantaginifolia monocephala T. & G. FI. 
IN. Az .2: 43. = 1843- 


Antennaria monocephala Greene, Pittonia 3: 176. 
1896. Not DC. 1836. 


Antennaria solitaria Rydb. Bull. Torr. Club 24: 304. 
1897. 


Stem slender, weak, floccose-woolly, 2’-10' 
long, bearing a solitary head. Basal leaves obo- 
vate to oblong-obovate or broadly spatulate, 33’ 
long or less, 8’-16’ wide, obtuse or apiculate, 
densely floccose beneath, loosely floccose, becom- 
ing glabrate above, 3-5-nerved; stem-leaves 
linear, few and distant; stolons procumbent, leafy 
at the ends; involucre 4”-6” high, its linear 
white-tipped bracts very woolly. 


Woodlands, Pennsylvania to Georgia, Ohio, Ala- 
bama and Louisiana. March—May. 


Geauseag. THISTLE FAMILY. 451 


6. Antennaria plantaginifolia (L.) 
Richards. Plantain-leaf Everlasting. 


Fig. 4398. 
Gnaphalium plantaginifolium L.Sp. Pl. 850. 1753. 
Antennaria plantaginifolia Richards. App. Frank. 

Journ Pda-zso8 te23" 

Floccose-woolly, stoloniferous, forming broad 
patches; flowering stems of fertile plant 4-20 
high, slender or stout, sometimes with glandu- 
lar hairs. Basal leaves obovate, spatulate, or 
broadly oval, obtuse or acutish, distinctly 3- 
ribbed, petioled, dull dark green and arachnoid 
above, silvery beneath, 13’-3’ long, 5/18 
wide; stem-leaves sessile, oblong or lanceolate, 
the upper usually small and distant; heads in 
corymbose or often subcapitate clusters, 4”-5” 
broad; involucre 3’—43” high, its bracts green- 
ish-white, linear to lanceolate, acute or acutish; 
achenes minutely glandular; sterile plant 
smaller, 3-8’ high; basal leaves somewhat 
smaller; heads smaller, 3-4” broad; bracts 
oblong, obtuse. 

In dry soil, especially in open woods, Quebec to 
Florida, Minnesota, Nebraska and Texas. Spring- 
or early everlasting. White plantain. Pussy-toes. 
Ladies’-tobacco. Dog-toes. Four-toes. Love’s-test. Indian- or woman’s-tobacco. Poverty-weed. Pearly 
mouse-ear everlasting. Consists of many races differing in size, leaf-form, leaf-size, size of heads 
and shape of involucral bracts; these have been variously regarded by authors as species and as 
varieties. April-June. 


7- Antennaria microphylla Rydb. Small-leaved 
Cat’s-foot. Fig. 4399. 

A. microphylla Rydb. Bull. Torr. Club 24: 303. 1897. 

?A. parvifolia Nutt. Trans. Am. Phil. Soc. (II) 7: 406. 184r. 


Stem slender, 8’-12’ high; stolons short, not over 23’ 
long. Basal leaves and those of the ends of the stolons 
spatulate, obtuse, or apiculate, 2’-8” long, 1-2” wide, 
narrowed from above the middle; stem-leaves linear- 
oblong, or the leaves linear-spatulate, often longer than 
the basal ones; heads corymbose, rather numerous; 
involucre 23”-34” high; bracts of the fertile heads 
linear-oblong, acute or acutish, those of the sterile ob- 
tuse. 


Dry plains and hills, Saskatchewan to Nebraska, British 
Columbia and New Mexico. July—Aug. 


8. Antennaria neodioica Greene. Smaller 
Cat’s-foot. Fig. 4400. 
Antennaria neodioica Greene, Pittonia 3: 184. 1897. 
A, alsinoides Greene, Pittonia 4: 83. 1899. 
A. rupicola Fernald, Rhodora 1: 74. 1899. 
Floccose-woolly, with numerous stolons which 
are leafy throughout; stem of fertile plants slen- 
der, about 1° high. Basal leaves about 1’ long, 
3-5” wide, broadly obovate to spatulate, 1-nerved, 
or indistinctly 3-nerved, white-tomentose beneath, 
becoming glabrate above, usually narrowed into 
distinct petioles; stem-leaves linear, acute; heads 
loosely corymbose, 3-4” broad; outermost bracts 
‘of the involucre obtuse, the rest lanceolate, acute, 
or acuminate, all greenish or brownish below, 
with white scarious tips; achenes obtusely 4-an- 
gled, granular-papillose; sterile plant lower, 3’-8’ 
high; heads more densely clustered, the bracts of 
the involucre oblong, obtuse. 


In dry places, Newfoundland to Virginia, Quebec, Michigan and South Dakota. April-July. 


452 COMPOSITAE. Vor. III. 


pi, g. Antennaria aprica Greene. Rocky Moun- 

Sy tain Cudweed. Fig. 4401. 

TI en Antennaria aprica Greene, Pittonia 3: 282. 1808. 

7 TON 

Zt Floccose-woolly or canescent, surculose, forming 
broad patches; flowering stems 2-12’ high. Basal 
leaves spatulate or obovate, obtuse, narrowed into 
short petioles, white-canescent on both sides, 4’—-15” 
long, 1-4” wide; stem-leaves linear, sessile; heads 


3-4” broad in a terminal capitate or corymbose 
cluster; involucre 2’-3” high, the bracts of the fer- 

by tile heads oblong, white or pink, all obtuse, or the 
inner ones acute; those of the sterile heads oval or 
elliptical, obtuse; achenes glabrous, or minutely 
glandular. 


In dry soil, South Dakota to Nebraska, Alberta, Utah 
and New Mexico. June-Sept. Erroneously referred, in 
SSN our first edition, as by previous authors, to the Old 
World Antennaria dioica (L.) Gaertn. 
SWS ) M( pg 
IN 
SS 


to. Antennaria campéstris Rydberg. 
Prairie Cat’s-foot. Fig. 4402. 


Antennaria campestris Rydberg, Bull. Torr. Club 

24: 304. 1897. 

Stolons short, leafy; flowering stems of both 
fertile and sterile plants 2-6’ high. Basal leaves 
obovate-cuneate, without a distinct petiole, 
white-tomentose beneath, glabrate above, I- 
nerved, or indistinctly 3-nerved; stem-leaves 
small, linear; heads 3’—-4” broad in subcapi- 
tate clusters; bracts of the fertile head¢ lan- 
ceolate, greenish below, brownish at the mid- 
dle, the apex white, acute or acuminate; 
bracts of sterile heads elliptic, obtuse. 


On dry prairies, Nebraska and Kansas to Sas- 
katchewan. May-June. 


11. Antennaria neglécta Greene. Field Cat’s- 
foot. Fig. 4403. 


Antennaria neglecta Greene, Pittonia 3: 173. 1897. 

An ennaria petaloidea Fernald, Rhodora 1: 73. 1899. 
A. neglecta simplex Peck, Bull. N. Y. State Mus. 67: Bot. 
629335 1903 

Stoloniferous, the stolons long and slender, bear- 
ing small leaves, except at the ends, where they are 
normally developed. Basal leaves oblanceolate or 
cuneate-spatulate, gradually tapering to a_ sessile 
base, without a distinct petiole, white-tomentose be- 


neath, glabrate above, I-nerved; stem-leaves linear; 
(/ | fertile plant nearly 1° high; heads 3’-4” broad, 


corymbose or sometimes only I or 2; bracts brown- 
ish, with white tips, lanceolate, acute; sterile plant 


KX — N j 4-8 high, the heads densely clustered, the bracts 
NSS oblong, obtuse. 


In fields and pastures, Maine to New York, Virginia 
and Wisconsin. April-June. 


CY) 


GENUS 43. THISTLE FAMILY. 453 


12. Antennaria dimorpha (Nutt.) T. & G. 
Low Everlasting. Fig. 4404. 
Gnaphalium dimorphum Nutt. Trans. Am. Phil. Soc. 

CLD) 7-sAossee no T- 

A. dimorpha T.& G. Fl. N. A. 2: 431. 1843. 

Tufted from a thick woody often branched cau- 
dex, 1-13’ high. Leaves all in a basal cluster, spatu- 
late, white-canescent or tomentose on both sides, 
obtuse or acutish, 3-1’ long, 1-2” wide, narrowed 
into short petioles; heads of staminate flowers 
about 3” broad and high, solitary and sessile 
among the leaves, or raised on a very short spar- 
ingly leafy stem, with obtuse or obtusish brown- 
ish involucral bracts, those of pistillate flowers 
longer, their inner bracts linear-lanceolate, acumi- 
nate; pappus of the staminate flowers slender, 
scarcely thickened, but barbellate at the tips, that 
of the pistillate fowers of fine and smooth bristles. 

Dry soil, Nebraska to Utah and California, north 
to Montana and British Columbia. April-June. 


44. ANAPHALIS DC. Prodr. 6: 271. 1837. 


Perennial white-tomentose or woolly herbs, with leafy erect stems, in our species alter- 
nate entire leaves, and small corymbose discoid heads of dioecious flowers. Involucre oblong 
to campanulate, its bracts scarious, imbricated in several series, mostly white, the outer 
shorter. Receptacle mostly convex, not chaffy. Staminate flowers with a slender or filiform 
corolla, an undivided style, and a pappus of slender bristles, not thickened at the summit, or 
scarcely so; anthers tailed at the base. Pistillate flowers with a tubular 5-toothed corolla, 
2-cleft style, and a pappus of capillary bristles. [Greek name of some similar plant.] 

About 35 species, natives of the north temperate zone. Besides the following, 2 or 3 other 
species occur in western North America. Type species: Anaphalis nubigena (Wall.) DC. 


1. Anaphalis margaritacea (L.) Benth. & 
Hook. Pearly or Large-flowered Ever- 
lasting. Fig. 4405. 

Gnaphalium margaritaceum L. Sp. Pl. 850. 1753. 
Antennaria margaritacea Hook. Fl. Bor. Am. 1: 329. 1833. 
A. margaritacea Benth. & Hook. Gen. Pl. 2: 303. 1873. 

Stem floccose-woolly, corymbosely branched at the 
summit, leafy, 1°-3° high. Leaves linear-lanceolate, 
narrowed to a sessile base, revolute, green, but mostly 
more or less pubescent above, woolly beneath, 3’-5’ 
long, 2-4” wide, the lowest shorter, spatulate, usually 
obtuse; corymb compound, 2-8’ broad; heads very nu- 
merous, short-peduncled or sessile, about 3” high, 4” 
broad when expanded; involucre campanulate, its 
bracts ovate-lanceolate, obtuse, finely striate, pearly 
white, mostly glabrous; pappus-bristles of the fertile 
flowers distinct and falling away separately. 

Dry soil, Newfoundland to Alaska, Pennsylvania, Kansas, 
Oregon and northern Asia. Recorded south to North Caro- 
lina. Adventive in Europe. Silver-leaf. Life-everlasting. 
Moonshine. Cotton-weed. None-so-pretty. Lady-never-fade. 
Indian-posy. Ladies’-tobacco. Poverty-weed. Silver-button. 
July—Sept. 

Anaphalis occidentalis (Greene) Heller, occurring from Alaska to California and in New- 
foundland and Quebec, differs by its mostly broader leaves being green and glabrous. 


45. GNAPHALIUM I. Sp. Pl. 850. 1753. 


Woolly erect or diffusely branched herbs, with alternate leaves, and discoid heads of 
pistillate and perfect flowers arranged in corymbs, spikes, racemes, or capitate. Receptacle 
flat, concex or conic, not chaffy, usually foveolate. Pistillate flowers in several series, their 
corollas filiform, minutely dentate or 3-4-lobed. Central flowers perfect, tubular, few, their 
corollas 5-toothed or 5-lobed. Anthers sagittate at the base, the auricles tailed. Achenes 
oblong or obovate, terete or slightly compressed, not ribbed. Pappus a single series of cap- 
illary bristles, sometimes thickened above. [Greek, referring to the wool.] 

About 120 species, widely distributed. Type species: Gnaphalium luteo-album L. 


1. Pappus-bristles distinct. 
‘Tall, erect ; inflorescence corymbose, or paniculate. 
Leaves sessile; plant not viscid. 
Leaves sessile; plant glandular-viscid. 
Leaves decurrent; plant glandular-viscid. 


SS 
AN Zam 


hi fi 


. G. obtusifolium. 
. G. Helleri. 
. G. decurrens. 


WON He 


454 COMPOSITAE. 7 Vor. IT. 


Low, diffuse; inflorescence mostly capitate ; pappus-bristles distinct. 


Floccose-woolly ; involucral bracts yellowish, or white. 4. G. palustre. 
Appressed-woolly ; involucral bracts becoming dark brown. 5. G.uliginosum. 
Tufted low mountain herbs ; heads few; bracts brown; pappus-bristles distinct. 6. G. supinum, 


2. Slender, simple; heads spicate; pappus-bristles united at base. 
Leaves linear or lanceolate-spatulate, acute; heads about 3” high; northeastern. 
Bracts dark brown; stem leaves lanceolate-spatulate. 7. G. norvegicum. 
Bracts brownish tipped ; stem leaves linear. 8. G. sylvaticum. 
Leaves spatulate, obtuse or obtusish ; heads 2”—214” high; eastern and southern. 9. G. purpureum. 
WAR Me . Seed 
WIE. 1. Gnaphalium obtusifolium L. Sweet or 
INE) SW White Balsam. Sweet or Fragrant Life 
LA Everlasting. Fig. 4406. 
aS Gnaphalium obtusifolium L. Sp. Pl. 851. 1753. 
= G. polycephalum Michx. Fl. Bor. Am. 2: 127. 1803. 
SS Annual or winter-annual, fragrant; stem erect, 
‘i simple; or branched above, tomentose, 1°-3° high. 
Leaves lanceolate or linear-lanceolate, sessile, 
acute or acutish, or the lower obtuse at the apex, 
narrowed at the base, densely white-woolly be- 
neath, glabrate and commonly dark green above, 
ea 1-3’ long, 2’-4” wide, the margins undulate; heads 
in numerous corymbose or paniculate clusters of 
I-5, about 3” high; bracts of the involucre white, 


or tinged with brown, oblong, thin and scarious, 

obtuse, the outer woolly at the base; pappus-bris- 

\ : tles distinct, separately deciduous; achenes glabrous. 
LY Z] In dry, mostly open places, Nova Scotia to Florida, 


Manitoba, Kansas and Texas. Jamaica. Poverty-, 
LP  chafe-or balsam-weed. Old-field balsam. Indian-posy. 
ye SS Feather-weed. Fussy-gussy. Rabbit-tobacco. Life-of- 
ORR man. Moonshine. Leaves of rosettes oblong. Aug.—Sept. 


2. Gnaphalium Hélleri Britton. Heller’s 
Everlasting. Fig. 4407. 

G. Helleri Britton, Bull. Torr. Club 20: 280. 1893. 
G. polycephalum Helleri Fernald, Rhodora 10: 94. 1908. 

Similar to the preceding species, corymbosely or 
somewhat paniculately branched above, 13°-2° high, 
the stem and branches densely glandular-pubescent, 
not tomentose. Leaves oblong-lanceolate, sessile, 
acuminate at both ends, green and hispidulous above, 
white-tomentose beneath, the larger about 2’ long 
and 5” wide, the uppermost much smaller and nar- 
rower; heads very numerous, corymbose or corym- 
bose-paniculate, sessile or short-peduncled in the 
clusters, about 23” broad; involucre oblong, or be- 
coming campanulate, 3” high, its bracts bright white, 
tomentose, the outer oblong, the inner linear-oblong, 
all obtuse; pappus-bristles distinct; achenes glabrous. 

In fields and woods, New York and New Jersey to 
Virginia, Kentucky and Georgia. Sept.—Oct. 


3. Gnaphalium dectrrens Ives. Clammy Everlasting. 
Winged Cudweed. Fig. 4408. 


Gnaphalium decurrens Ives, Am. Journ. Sci. 1: 380. pl. r. 1819. 


Annual or biennial, similar to the two preceding species, fra- 
grant; stem very leafy, glandular-viscid, corymbosely branched 
above, 2°-3° high. Leaves lanceolate or broadly linear, acutish 
at the apex, densely white-woolly beneath, glabrate or loosely 
woolly above, sessile and decurrent on the stem at the base, 
1-3’ long, 2-3” wide, or the lowest shorter and slightly spatu- 
late; heads in several or numerous corymbose glomerules of 
2-6, about 3” high; bracts of the involucre yellowish-white or 
brownish, ovate, acutish or the inner lanceolate and acute, the 
outer woolly at base; pappus-bristles distinct; achenes glabrous. 


In open, moist or dry places, Nova Scotia to Pennsylvania, West 
Virginia ?, Ontario, British Columbia, Michigan, south in the Rocky 
Mountains to Arizona. Sweet balsam. Balsam-weed. July—Sept. 


GENUS 45. THISTLE FAMILY. 


.4. Gnaphalium palistre Nutt. Western Marsh 
Cudweed. Fig. 4400. 


Gnaphalium palustre Nutt. Trans. Am. Phil. Soc. (II) 7: 
403. 1841. 


Annual; diffusely branched from the base, densely 
but loosely floccose-woolly all over, 2-8 high. Leaves 
sessile, oblong, linear-oblong, or the lower spatulate, 
obtuse or acutish, 6-12” long, 14’’-3” wide; heads 
about 2” high, several together in leafy-bracted glom- 
erules; involucre more or less woolly, its bracts linear 
or lanceolate, acute, white or pale yellow; pappus- 
bristles distinct, separately deciduous. 


In moist wet soil, Saskatchewan to Montana, western 
Nebraska and New Mexico, west to British Columbia and 
California. May—Aug. 


5. Gnaphalium uligindsum L. Low or 
Marsh Cudweed. Wartwort. Mouse- 
ear. Fig. 4410. 


Gnaphalium uliginosum L. Sp. Pl. 856. 1753. 


Annual; diffusely branched from the base, or 
the stems sometimes erect or ascending, ap- 
pressed-woolly all over, 2’-8’ high. Leaves sessile, 
spatulate-linear, linear, or the lower oblanceolate 
or spatulate and narrowed into petioles, all ob- 
tuse or obtusish, generally mucronulate, 1’—14’ 
long; heads about 2” high, numerous in dense 
leafy-bracted terminal glomerules; bracts of the 
involucre oblong or oblong-lanceolate, brown, the 
outer obtuse or obtusish and more or less woolly, 
the inner acute; pappus-bristles distinct, sepa- 
rately deciduous. 

In damp soil, Newfoundland to Virginia, west to 


“western Ontario, Saskatchewan, British Columbia and 
Oregon, and Indiana. Also in Europe. July—Sept. 


6. Gnaphalium’supinum L. Dwarf Cudweed. 
Fig. 4411. 


Gnaphalium supinum L. Syst. Ed. 2, 234. 1767. 


Perennial, white~woolly, much tufted; stems sim- 
ple, 17-33’ high. Leaves mainly basal, linear, acute, 
‘narrowed at the base, sessile, 6’—-12” long, 1’-2” 
wide; heads few or several, capitate or short-spicate, 
about 3” high; flowers yellowish; bracts of the invo- 
lucre brown, glabrous, lanceolate or oblong-lanceo- 
late, acute; pappus-bristles distinct, separately de- 


ciduous. 


Alpine summit of the White Mountains of New Hamp- 
shire, and of Mt. Katahdin, Maine; Labrador and Green- 
land, and on high mountains in Europe and Asia. Called 
also mountain-cudweed. July—Aug. 


COMPOSITAE. Vor iit 


7. Gnaphalium norvégicum Gunner. Nor- 
wegian Cudweed. Fig. 4412. 


G. norvegicum Gunner, Fl. Norveg. 2: 105. 1772. 


Perennial; stem simple, 6-18’ high. Leaves lan- 
ceolate to spatulate, elongated, acute, narrowed 
at the base, woolly on both sides, or green and 
glabrate above, 3-6’ long, 2’-5” wide, the lower 
and basal ones petioled; heads about 3” high, 
numerous in a more or less leafy spike, the lowest 
often distant, solitary or glomerate in the upper 
axils; bracts of the involucre ovate-oblong, dark 
brown, or brown-tipped, glabrous or slightly 

woolly, obtuse; pappus-bristles united at the base, 
LD aN falling away in a ring; achenes hispidulous. 


INN : 
SW , 
SS > 


Mt. Albert, Gaspé, Quebec, north to Greenland and 
Arctic America. Also in Europe. July—Aug. 


tare’ 


wt V4 
Why 


8. Gnaphalium sylvaticum L. Wood 
Cudweed. Chafweed. Owl’s Crown. 
Golden Motherwort. Fig. 4413. 


Gnaphalium sylvaticum L. Sp. Pl. 856. 1753. 


Perennial; stem slender, simple, 6-18’ high. 
Leaves linear, acute, 1-2’ long, 1-23” wide, 
or the lowest linear-spatulate, woolly beneath, 
glabrous or glabrate above; heads about 3” 
high, numerous in a more or less leafy spike, 
or the lowest solitary or glomerate in the 
upper axils; bracts of the involucre linear- 
oblong, obtuse, mostly glabrous, yellowish or 
greenish with a brown spot at or just below 
the apex; pappus-bristles united at the base; 


achenes hispidulous. \\ DB 
New Brunswick and Cape Breton Island to 

Quebec and northern Maine and New Hampshire. | 

Widely distributed in Europe and northern Asia. 

June-Aug. 


g. Gnaphalium purpureum L. Purplish 
Cudweed. Fig. 4414. 


Gnaphalium purpureum L. Sp. Pl. 854. 1753. 


Annual or biennial, simple and erect or branch- 
ed from the base and the branches ascending, 2’—2° 
high. Leaves spatulate, or the uppermost linear, 
mostly obtuse, mucronulate, woolly beneath, usu- 
ally green and glabrous or nearly so above when 
old, sessile, or the lowest narrowed into petioles, 
1-2’ long, 2”-6” wide; heads 2’-23” high in a 
terminal, sometimes leafy, often interrupted spike, 
or the lowest ones distant and axillary; bracts of 
the involucre yellowish brown or purplish, lan- 
ceolate-oblong, acute or acutish, the outer woolly 
at the base; pappus-bristles united below; achenes 
roughish. 


In dry sandy soil, eastern Maine to Florida, Penn- 
sylvania, West Virginia, Kentucky, Kansas and Texas. 
Bermuda; Jamaica; Mexico. Far western plants for- 
merely referred to this species prove to be distinct. 
May-Sept. 


Genus 46. | THISTLE FAMILY. 457 


46. ADENOCAULON Hook. Bot. Misc. 1: 19. 1830. 


Perennial herbs, with broad alternate slender-petioled leaves, woolly beneath, and pani- 
cled small 5-10-flowered heads of tubular flowers. Involucre campanulate, composed of a 
few herbaceous bracts. Receptacle nearly flat, naked. Corollas all tubular, 4-5-lobed. Mar- 
ginal flowers pistillate, fertile. Central flowers perfect, sterile, the style undivided; anthers 
slightly sagittate at the base. Pappus none. Achenes obovoid or clavate, very obtuse, faintly 
nerved, glandular above, longer than the bracts of the involucre. [Greek, gland-stem.] 


Two species, natives of North America, Japan and the Himalayas. Only the following typical 
one is known in North America. 


1. Adenocaulon bicolor Hook. Adenocaulon. 
Fig. 4415. 


Adenocaulon bicolor Hook. Bot. Misc. 1: 19. pl. 15. 1830. 


Stem floccose-woolly, or at length glabrous, 1°-3° 
high, leafless and mostly paniculately branched above. 
Leaves all basal or nearly so, deltoid-ovate, obtuse or 
acute at the apex, deeply cordate at the base, coarsely 
repand-toothed or lobed, thin, green and glabrous above, 
densely and persistently white-woolly beneath, 2’-6’ 
long and broad, with slender narrowly margined peti- 
oles; heads numerous, very slender-peduncled, small; 
bracts of the involucre 4 or 5, ovate to lanceolate, re- 
flexed in fruit, at length deciduous; achenes 3’-4” 
long, 4” thick, the upper part beset with nail-shaped 
glands. 


In moist woods, northern Michigan and Lake Superior to 
British Columbia, Montana and California. May-July. 


47. INULA L. Sp. Pl. 881. 1753. 


Perennial, mostly tomentose or woolly herbs, with alternate and basal leaves, and large 
heads of both tubular and radiate yellow flowers. Involucre hemispheric or campanulate, 
its bracts imbricated in several series, the outer often foliaceous. Receptacle flat or convex, 
areolate or foveolate, not chaffy. Ray-flowers pistillate, their ligules 3-toothed. Disk-flowers 
perfect, their corollas tubular, 5-toothed. Anthers sagittate at the base, the auricles caudate. 
Style-branches of the disk-flowers linear, obtuse. Achenes 4-5-ribbed; pappus of capillary 
rough bristles in our species. [The ancient Latin name.] 

About 90 species, natives of Europe, Asia and Africa, the following typical. 


1. Inula Helénium L. Elecampane. 
Horseheal. Fig. 4416. 


Inula Helenium L. Sp. Pl. 881. 1753. 


Stems tufted from large thick roots, simple 
or rarely somewhat branched, 2°-6° high, 
densely pubescent above. Leaves large, broadly 
oblong, rough above, densely pubescent be- 
neath, denticulate, the basal ones acute at each 
end, long-petioled, 10’-20’ long, 4’-8’ wide; 
stem leaves sessile, or cordate-clasping at the 
base, acute at the apex, smaller; heads solitary 
or few, terminal, stout-peduncled, 2’-4’ broad; 
involucre hemispheric, nearly 1’ high, its outer 
bracts ovate, foliaceous, pubescent: rays nu- 
merous, linear; achenes glabrous, 4-sided. 


Along roadsides and in fields, Nova Scotia to 
Ontario and Minnesota, south to North Carolina 
and Missouri. Naturalized from Europe. Native 
also in Asia. Scabwort. Horse-elder. Yellow star- 
wort. Elfdock. Elfwort. Wild sunflower. 


458 COMPOSITAE. Vot. III. 


48. POLYMNIA L. Sp. Pl. 926. 1753. 


Perennial herbs (some tropical species woody), with opposite membranous lobed or 
angled leaves, or the lower alternate, and mostly large corymbose-paniculate heads of both 
tubular and radiate yellow or whitish flowers, or rays sometimes wanting. Involucre hemi- 
spheric or broader, of about 5 large outer bracts, and more numerous smaller inner ones. 
Receptacle chaffy. Ray-flowers pistillate, fertile, subtended by the inner involucral bracts, 
the ligules elongated, minute, or none. Disk-flowers subtended by the chaffy scales of the 
receptacle, perfect, sterile, their corollas tubular, 5-toothed. Anthers 2-toothed at the base. 
Pappus none. Achenes thick, short, turgid, glabrous. [From the Muse Polhymnia.] 

About 10 species, natives of America. Only the following are known in North America. Type 
species: Polymnia canadensis L. 

Rays commonly 6” long or more, yellow; achenes strongly striate. 1. P. Uvedalia. 
Rays commonly minute or up to 6” long, whitish, or none; achenes 3-ribbed. 2. P. canadensis. 


1. Polymnia Uvedalia L. Yellow or Large- 
flowered Leaf-cup. Fig. 4417. 
Polymnia Uvedalia L. Sp. Pl. Ed. 2, 1303. 1763. 


Rough-pubescent, stout, branched, 3°-10° high. 
Leaves broadly ovate or deltoid, 3-nerved, abruptly 
contracted above the base, minutely ciliate, more 
or less pubescent on both sides, angulate-lobed, the 
lower often 1° long and broad, petioled, the upper 
sessile, somewhat clasping; heads few in terminal 
clusters, peduncled, 14-3’ broad; rays I0-15, com- 
monly 6-12” long, linear-oblong, bright yellow, 
3-toothed or entire; exterior bracts of the cup-like 
involucre ovate-oblong, obtuse, ciliate, 4’—10” long; 
achenes slightly oblique and laterally compressed, 
strongly striate, nearly 3” long. 


In rich woods, New York to Indiana, Florida, Mis- 
souri, Oklahoma and Texas. Bermuda. Yellow bears- 
foot. July—Aug. 


2. Polymnia canadénsis L. Small-flowered 
Leaf-cup. Fig. 4418. 

Polymnia canadensis L. Sp. Pl. 926. 1753. 

Polymnia canadensis radiata A. Gray, Syn. Fl. N. A. 

Tevbant 2 2goussteod: 

P. radiata Small, Fl. S.E. U. S. 1239. 1903. 

Rather slender, viscid-pubescent, at, least above, 
simple or branched, 2°-5° high. Leaves deltoid- 
ovate to hastate, usually very thin, all petioled, 
deeply angulate-lobed and the lobes dentate, or 
the lower lyrate-pinnatifid, 4-10’ long, the upper- 
most sometimes ovate and entire or merely den- 
ticulate; heads few in terminal clusters, short- 
peduncled or sessile, 4”-6” broad; outer bracts of 
the involucre ovate to lanceolate, obtuse or acut- 
ish, 2-3” long; rays small, minute or none, or 
sometimes up to 6” long and 3-lobed, whitish or 
yellowish; achenes 3-angled, obovoid, obcom- 
pressed, 3-ribbed, not striate. 


In damp, rich shaded places, Vermont and Ontario i 
to Minnesota, Pennsylvania, North Carolina, Tennessee and Arkansas. June—Sept. 


49. MELAMPODIUM L. Sp. Pl. 921. 1753. 

Herbs, some species woody, with opposite entire or dentate leaves, and terminal peduncled 
heads of both tubular and radiate, white or yellow flowers in our species. Involucre hemi- 
spheric, its bracts in 2 series, the 4 or 5 outer ones broad, often connate at the base, the inner 
hooded, embracing or permanently surrounding the pistillate fertile ray-flowers. Receptacle 
convex or conic, chaffy. Ray-flowers in I series, the rays spreading, 2-3-lobed or entire. 
Disk-flowers perfect, sterile, their corollas with a narrowly campanulate 5-toothed limb, the 
anthers entire at the base, the style undivided. Achenes obovoid, more or less incurved. 
Pappus none. [Greek, black-foot, of doubtful significance, but the stem bases of the typical 


species are dark-colored. ] 


GENUS 49. THISTLE FAMILY. 459 


About 35 species, natives of the warmer parts of America. Besides the following, 2 or 3 others 
occur in the southwestern United States. Type species: Melampodium americanum L. 


1. Melampodium leucanthum T. & G. 
Plains Melampodium. Fig. 4419. 


Melampodium leucanthum T. & G. Fl. N. A. 2: 271. 
1842. 


Perennial, woody at the base, branched, canes- 
cent, 4’-12’ high, the branches slender. Leaves 
linear, lanceolate, or the lower spatulate, sessile, 


TIS EE 


entire, or nearly so, canescent, 1-2’ long, 13’’-3 
wide, obtuse or obtusish at the apex; heads 3’—3’ 
broad, terminating the branches; peduncles slen- 
der, 1-3’ long; outer bracts of the involucre ovate 
or oval, obtuse, united below; rays 5-9, cuneate- 
oblong, white, 2-3-lobed, firm in texture, veiny, 
persistent; inner bracts turbinate or terete, hood- 
ed, muricate, the hood wider than the body. 

In dry soil, Kansas to Colorado, Arizona, Texas 
and Mexico. June—Oct. Not distinguished, in our 
first edition, from M. cinereum DC. of Texas and 
northern Mexico. 


50. ACANTHOSPERMUM Schrank, Pl. Rar. Hort. Monac. pl. 53. 1819. 


Annual rather coarse herbs, with pubescent foliage and erect or creeping stems, opposite, 
broad, often leathery, toothed leaves, and radiate but inconspicuous heads, axillary to leaf- 
like bracts. Involucre double, an outer one of flat herbaceous bracts, and an inner one of 
several smaller bracts which become bur-like and fall away enclosing an achene at maturity. 
Receptacle concave or convex. Ray-flowers few, in I series, the rays very small, yellowish, 
concave or hooded. Disk-flowers perfect, sterile. Anthers entire at the base. Achenes 
broadest above the middle, slightly curved. Pappus wanting. [Greek, thorn-seed, from the 
prickly, bur-like fruits. ] 


About 3 species, natives of tropical America. Type species: Acanthospermum brasilium Schrank. 


Ly 


1. Acanthospermum australe (Loefl.) Kuntze. 
Spiny-bur. Fig. 4420. 
Melampodium australe Loefl. Iter. Hisp. 268. 1758. 
A. xanthoides DC. Prodr. 5: 521. 1836. 
A, australe Kuntze, Rev. Gen. Pl. 303. 1891. 


Stems branching at the base, the branches prostrate or 
creeping, 3°-2° long. Leaves ovate, oval or rhombic, 4’—1’ 
long, usually acute, serrate-dentate above the middle, cu- 
neate at the base; peduncles shorter than the subtending 
leaves; involucre campanulate, the bracts broadly ovate, 
obtuse, 1”-14” long, uniformly prickly; disk-flowers with 
corollas 1” long, puberulent; mature inner bracts of the 
involucre forming a starfish-like bur, each lobe 4-5” long, 
densely beset with uniform blunt weak prickles. 


In waste places and dry soil, Virginia to Florida and Louisi- 
ana. June—Oct. Widely distributed as a weed in tropical re- 
gions. 


51. SILPHIUM L. Sp. Pl. 919. 1753. 


Tall perennial herbs, with resinous juice, opposite whorled or alternate leaves, and large 
corymbose or paniculate (rarely solitary) peduncled heads of both tubular and radiate yellow 
flowers. Involucre hemispheric or campanulate, its bracts imbricated in few series. Recep- 
tacle flat or nearly so, chaffy, the chaff subtending the disk-flowers. Ray-flowers in 2 or 3 


460 COMPOSITAE. Vor. III. 


series, pistillate, fertile, the ligules numerous, linear. Disk-flowers perfect but sterile, their 
corollas tubular, 5-toothed, the style undivided. Anthers minutely 2-toothed or entire at the 
base. Achenes broad, dorsally flattened, 2-winged, notched at the apex. Pappus none, or of 
2 awns confluent with the wings of the achene. [Greek, from the resinous juice.] 


About 12 species, natives of North America, known as Rosin-weed or Rosin-plant. Type species: 
Silphium Asteriscus L. 


Stem leafy, the leaves opposite, alternate, or verticillate. 


Leaves, or their petiole-bases, connate-perfoliate ; stem square. 1. S. perfoliatum. 
Leaves not connate-perfoliate, sessile or petioled. 
Leaves ‘opposite, or the uppermost alternate; cauline sessile. 2. S. integrifolium. 
Leaves, or some of them, verticillate in 3’s or 4’s, petioled. 3. S. trifoliatum. 
Most or all of the leaves alternate, entire or dentate. 4. S. Asteriscus. 
Leaves all alternate, pinnatifid or bipinnatifid, large. 5. S. laciniatum, 
Stem leafless or nearly so, scaly above; leaves basal, large. 
Leaves sharply serrate to pinnatifid ; achenes obovate. 6. S. terebinthinaceum. 
Leaves coarsely dentate; achenes suborbicular. 7. S. reniforme. 


1. Silphium perfoliatum L. Cup- 
plant. Indian-cup. Fig. 4421. 


S. perfoliatum L. Sp. Pl. Ed. 2, 1301. 1763. 


Stem square, glabrous, or rarely some- 
what hispid, branched above, or sometimes 
simple, 4°-8° high. Leaves ovate or deltoid- 
ovate, opposite, the upper connate-perfol- 
iate, the lower abruptly contracted into 
margined petioles, all thin, usually scabrous 
on both sides, or pubescent beneath, coarse- 
ly angulate-dentate, or the upper entire, the 
larger 6-12’ long, 4’-8’ wide; heads com- 
monly numerous, 2-3’ broad; rays 20-30, 
about 1’ long and 2” wide; involucre de- 
pressed-hemispheric, its outer bracts broad, 
ovate, ciliolate, spreading or erect; achenes 
obovate, emarginate, sometimes 2-toothed. 


In moist soil, southern Ontario to Minne- 
sota, South Dakota, New Jersey, Georgia, Ne- 
braska and Louisiana. Naturalized near New 
York City, and elsewhere escaped from culti- 
vation. Called also ragged-cup. July—Sept. 


2. Silphium integrifolium Michx. Entire- 
leaved Rosin-wood. Fig. 4422. 
S. integrifolium Michx. Fl. Bor. Am. 2: 146. 1803. 


S. speciosum Nutt. Trans. Am. Phil. Soc. (II) 7: 341. 
1841. 


Stem glabrous, rough or sometimes hirsute, co- 
rymbosely branched above, 2°-5° high. Leaves ovate 
to ovate-lanceolate, acute or acuminate, entire, den- 
ticulate or remotely dentate, rough above, pubescent 
or glabrous beneath, those of the stem all closely 
sessile, often half-clasping but not connate-perfol- 
iate at the rounded base, 3’-5’ long, 1’-2’ wide; 
heads usually numerous, 1-2’ broad; involucre 
nearly hemispheric, its outer bracts ovate or ovate- 
lanceolate, acute, spreading, ciliolate or pubescent; 
rays 15-25; achenes oval or obovate, 4’-5” long, 
deeply emarginate. 


On prairies, Ohio to Minnesota, south to Louisiana, 
Nebraska, Arkansas and Texas. Aug.—Sept. 


GENUS 5I. THAIS OLE 


3. Silphium trifoliatum L. Whorled 
Rosin-weed. Fig. 4423. 
Silphium trifoliatum L. Sp. Pl. 920. 1753. 


Stem glabrous, sometimes glaucous, corym- 
bosely branched at the summit, 4°-7° high. 
Leaves lanceolate or oblong-lanceolate, the 
middle ones almost always whorled in 3’s or 
4's, acuminate at the apex, narrowed at the 
base and usually somewhat petioled, rough 
or roughish above, pubescent or nearly gla- 
brous beneath, entire or denticulate, 3-7’ long, 
3’-13’ wide; heads several or numerous, 13’—2’ 
broad; involucre hemispheric, its outer bracts 
ovate or oval, acute or obtuse, glabrous or 
slightly pubescent, ciliolate; rays 15-20; achenes 
oval-or obovate, narrowly winged, emarginate, 
sharply 2-toothed. 


In woods, Pennsylvania to Ohio, Ontario, Vir- 
ginia and Alabama. July—Oct. 


4. Silphium Asteriscus L. Starry Rosin- 
weed. Fig. 4424. 
Silphium Asteriscus L. Sp. Pl. 920. 1753. 


Stem hispid-pubescent, simple or branched above, 
2°-4° high, usually purple. Leaves nearly all 
alternate, ovate, ovate-oblong, or lanceolate, acute 
or obtusish, sessile, somewhat clasping, or the 
lower narrowed into short petioles, dentate, or 
the upper entire, 2’-5’ long, 4-1’ wide; heads 
commonly few, I’-2’ broad; rays 12-15; invo- 
lucre hemispheric, its bracts mostly hispid, ovate 
to oblong, acute or obtuse, squarrose; achenes 
oval or obovate, narrowly winged, 2-toothed. 


In dry soil, Maryland to Tennessee and Missouri, 
south to Florida and Louisiana. June—Sept. 


5. Silphium laciniatum L. Compass- 
plant. Pilot-weed. Fig. 4425. 


Silphium laciniatum L. Sp. Pl. 919. 1753. 


Rough or hispid, very resinous; stem 6°-12° 
high; basal leaves pinnatifid or bipinnatifid, 
long-petioled, 1° long or more, the lobes ob- 
long or lanceolate; stem leaves alternate, ver- 
tical, their edges tending to point north and 
south, sessile, or the lower short-petioled, the 
upper cordate-clasping at the base, gradually 
smaller and less divided; heads several or 
numerous, sessile or short-peduncled, 2’—5’ 
broad, the peduncles bracted at the base; rays 
20-30, 1’—2’ long; involucre nearly hemispheric, 
its bracts large, rigid, lanceolate or ovate, very 
squarrose; achenes oval, about 6” long, the 
wing broader above than below, notched at 
the apex, awnless. 


On prairies, Ohio to South Dakota, south to 
Alabama, Louisianaand Texas. Turpentine-weed, 
polar-plant, rosin-weed. July—Sept. 


I 


COMPOSITAE. Vor. IIT. 


6. Silphium terebinthinaceum Jacq. 
Prairie Dock. Prairie Burdock. 


Fig. 4426. 
any A or : Silphium terebinthinaceum Jacq. Hort. Vind. 
PL NEY E 1: pl. 43. 1770. 
va . ; ; S. pinnatifidum Ell. Bot. S. C. & Ga. 2: 462. 


1824. 


Silphium terebinthinaceum pinnatifidum A. 
Gray, Man. 220. 1848. 


Stem glabrous or nearly so, branched 
and scaly above, 4°-10° high. Leaves all 


fee Nas Wig . basal or nearly so, coriaceous, ovate, mostly 
wet. Tyo eS long-petioled, acute at the apex, cordate 
ee Seo ae at the base, rough on both sides, often 12° 

Gsee oan . long and 6’ wide, sharply dentate, laciniate 


or pinnatifid; heads numerous, 13’—3’ broad, 
borne on glabrous peduncles; involucre 
eG C ora Ree hemispheric, its bracts ovate-oblong, erect, 
: Se eee glabrous or minutely pubescent; rays 12-20; 
ae Sa: ne ae achenes obovate, narrowly winged, slightly 
2-toothed and emarginate at the apex. 


On prairies and in dry woods, southern On- 
tario and Ohio to Minnesota, south to Georgia, 
Iowa and Louisiana. Rosin-plant. July—Sept. 


7. Silphium reniforme Raf. Kidney- 
leaved Silphium. Fig. 4427. 


Silphium reniforme Raf.; Nutt. Trans. Am. 
Phil. Soc. (II) 7: 342. 1840. 


S. compositum reniforme T. & G. Fl. N. A. 2: 

276. 1842. 

Stem scape-like, 3°-6° tall, glabrous or 
nearly so, corymbosely branched above. 
Leaves mostly basal, broadly ovate to reni- 
form, 4-15’ long, obtuse or abruptly point- 
ed, coarsely dentate, deeply cordate at the 
base, long-petioled; heads numerous, about 
1’ broad, the peduncles glabrous; involucre 
campanulate, the bracts ovate or broadly 
oblong to narrowly oblong or oblong-spatu- 
late, obtuse; rays several, yellow; achenes 
suborbicular or orbicular-obovate, 34” long, 
the wings prolonged enough to form a shal- 
low apical sinus. 


In dry or stony soil, mountains of Virginia 
and North Carolina. July—Aug. 


Silphium compositum Michx., a related 
southern species with pedately parted leaves, 
is reported as observed in southern Virginia. 


52. CHRYSOGONUM L. Sp. Pl. 920. 1753. 


Pubescent perennial herbs, with opposite and basal petioled leaves, and slender-peduncled- 
axillary and terminal, rather large heads of both tubular and radiate, yellow flowers. Invo- 
lucre hemispheric, its bracts in 2 series of 5, the outer large, obovate or spatulate, foliaceous, 
the inner oval, firm, each subtending a pistillate ray-flower. Receptacle chaffy, each scale 
subtending and partly enclosing a perfect but sterile tubular Mower with a 5-toothed corolla. 
Anthers nearly entire at the base. Achenes obovate, compressed, their margins acute, not 
winged, I-nerved on the back, I-2-ribbed on the inner side. Pappus a short half-cup-shaped 
crown. [Greek, golden-knee. ] 


A monotypic genus of eastern North America. 


GENUS 52. THISTLE FAMILY. 463 


1. Chrysogonum virginianum L. Chryso- 
gonum. Fig. 4428. - 


Chrysogonum virginianum L. Sp. Pl. 920. 1753. 


Chrysogonum virginianum dentatum A. Gray, Bot. 
Gaz. 72 3h esse: 


Perennial by rootstocks or runners, pubescent 
or hirsute throughout, branched from the base, 
or at first acaulescent, 3-12’ high. Leaves ovate or 
oblong, obtuse or acutish at the apex, the upper 
sometimes subcordate at the base, dentate or 
crenate-dentate, rather thin, 1-3’ long, 3’-2’ wide, 
the basal ones with petioles as long as the blade 
or longer, those of the upper one shorter; pedun- 
cles 1-4’ long; heads 1-13’ broad; outer bracts 
of the involucre obtuse or acute; rays about 5, 
4-7” long. 


In dry soil, southern Pennsylvania to Florida and 
Alabama. April—July. 


53. BERLANDIERA DC. Prodr. 5: 517. 1836. 


Perennial canescent or pubescent herbs, with alternate leaves and rather large, peduncled 
solitary or corymbose heads of both tubular and radiate yellow-flowers. Involucre depressed- 
hemispheric, its bracts imbricated in about 3 series, the outermost small, mostly oblong, the 
second series broader, oval or obovate, the inner membranous, similar, reticulated when mature, 
subtending the ray-flowers and exceeding the disk. Receptacle nearly flat, chaffy, the chaff 
subtending the disk-flowers. Ray-flowers 5-12, pistillate, fertile. Disk-flowers perfect, tubular, 
sterile, their corollas 5-toothed. Anthers entire, or minutely 2-toothed at the base. Style of 
the tubular flowers undivided, hirsute. Achenes obovate, compressed, not winged, 1-ribbed 
on the inner side, the pappus obsolete, early deciduous or of 2 caducous awns. [Named 
after J. L. Berlandier, a Swiss botanical collector in Texas and Mexico.] 


About 8 species, natives of the southern United States and Mexico. Type species: Berlandiera 
texana DC. 


Stem leafy ; leaves ovate to oblong, crenate. ; 1. B. texana. 
Plant acaulescent, or nearly so; leaves lyrate-pinnatifid. 2. B.lyrata. 


1. Berlandiera texana DC. Texan 
Berlandiera. Fig. 4420. 


Berlandiera texana DC. Prodr. 5: 517. 1836. 


Hirsute-pubescent throughout; stem erect, 
branched above, or simple, 2°-3° high, leafy. 
Leaves ovate, or the basal oblong, crenate, 
acutish or obtuse at the apex, rounded or 
cordate at the base, 2’-4’ long, 1-2’ wide, the 
upper sessile, the lower petioled; heads few 
or several, 1-14’ broad, in a terminal corym- 
bose-cymose cluster; peduncles 3-13’ long; 
inner bracts of the involucre twice as large 
as the outer. 


In dry soil, Missouri and Kansas to Arkansas 
and Louisiana. July—Aug. 


464 COMPOSITAE. Vor. II. 


2. Berlandiera lyrata Benth. Lyre-leaved 
Berlandiera. Fig. 4430. 


Silphium Nuttallianum Torr. Ann. Lyc. N. Y. 2: 216. 
Name only. 1827. 


Berlandiera lyrata Benth. Pl. Hartw. 17. 1839. 


Finely whitish-canescent, acaulescent or  short- 
stemmed; scapes or peduncles slender, 3-8 long, 
bearing a solitary head, or rarely 2. Leaves lyrate- 
pinnatifid, obtuse, petioled, the terminal segment 
usually larger than the lateral ones, the lower ones 
very small, all obtuse, mostly crenate, sometimes 
becoming green and glabrate above; head about 1’ 
broad; inner bracts of the involucre much broader 
than the outer, orbicular, or wider than long; achenes 
obovate, keeled on the inner face. 


In dry soil, Kansas to Texas, Arizona and Mexico. 


54. ENGELMANNIA T. & G. Fl. N. A. 2: 283. 1841. 


Perennial hirsute herbs, with alternate pinnatifid leaves, and corymbose slender-peduncled 
rather large heads of both tubular and radiate yellow flowers. Involucre hemispheric, its 
bracts imbricated in 2 or 3 series, the outer linear, loose, hirsute, ciliate, the inner oval or 
obovate, concave, appressed, subtending the ray-flowers. Receptacle flat, chaffy, the chaff 
subtending and partly enclosing the disk-flowers. Rays 8-10, pistillate, fertile. Disk-flowers 
about as many, tubular, perfect, sterile, the corolla 5-toothed. Anthers minutely 2-dentate 
at the base. Style of the tubular flowers undivided. Achenes obovate, compressed, not 
winged, I-ribbed on each face. Pappus a persistent irregularly cleft crown. [Named for 
Dr. Geo. Engelmann, 1809-1884, botanist, of St. Louis.] 


A monotypic genus of the south-central United States. 


1. Engelmannia pinnatifida T. & G. Engel- 
mannia. Fig. 4431. 


E. pinnatifida T. & G. Fl. N. A. 2: 283. 1841. 


Stem usually branched above, 1°-3° high. Basal 
leaves slender-petioled, 4’-8’ long, their lobes lanceo- 
late or oblong, dentate or entire, obtuse or acutish; 
upper leaves smaller, sessile, less divided, the upper- 
most sometimes entire, or with a pair of basal lobes; 
heads usually numerous, about 1’ broad; peduncles 
1-5’ long; outer bracts of the involucre somewhat 
in 2 series, the first linear, the second broadened at 
the base. 

In dry soil, Kansas to Colorado, Louisiana, Arizona 


and North Mexico. Reported as found along railroads 
in western Missouri. May—Aug. 


55. PARTHENIUM L. Sp. Pl. 988. 1753. 


Perennial, mostly pubescent or canescent herbs, or shrubs, with alternate leaves, and 
small corymbose or paniculate heads of both tubular and radiate white or yellow flowers. 
Involucre broadly campanulate or hemispheric, its bracts imbricated in 2 or 3 series, obtuse, 
appressed, nearly equal. Receptacle convex or conic, chaffy, the chaff membranous, sur- 
rounding the disk-flowers. Ray-flowers about 5, pistillate, fertile, their ligules short, broad, 
2-toothed or obcordate. Disk-flowers perfect, sterile, their corollas 5-toothed, the style undi- 
vided. Anthers entire at the base. Achenes compressed, keeled on the inner face, margined, 
bearing the persistent rays on the summit. Pappus of 2-3 scales or awns. [Greek, virgin.] 


About 12 species, natives of North America, Mexico, the West Indies and northern South America. 
Besides the following, some 3 others occur in the southern and southwestern United States. Type 
species: Parthenium Hysterophorus L. 


GENUS 55. THISTLE FAMILY. 


Leaves 1—2-pinnatifid ; annual weed. , 
Leaves crenate-dentate, or somewhat lyrate; perennials. 


Stem glabrous, or pubescent or puberulent above; rootstock 


tuberous-thickened. 
Stem hirsute or villous. ; ; 
Stem leaves auricled, clasping at base; rootstock thick. 


Stem leaves sessile, not auricled; rootstock long, slender. 


1. Parthenium Hysteréphorus L. Santa Maria. 
Fig. 4432. 
P. Hysterophorus L. Sp. Pl. 988. 1753. 


Annual, strigose-pubescent or somewhat villous, usu- 
ally much branched, 1°-23° high. Leaves ovate to ob- 
long in outline, I-2-pinnately parted into linear or lan- 
ceolate toothed or pinnatifid segments, thin and flaccid; 
heads numerous, 22”—3” broad; involucre saucer-shaped, 
its bracts concave, the outer ones rhombic, the inner 
broader; ray-flowers few; rays reniform, white, about 


465 


1. P. Hysterophorus. 


2. P.integrifolium. 


3. P. auriculatum. 
4. P. hispidum. 
ER, 


3” wide; achenes obovate, about 24” long. 


Waste and cultivated grounds, southern Pennsylvania to 
Illinois, Missouri, Florida and Texas. 


America. July—Sept. 


Throughout tropical 


2. Parthenium integrifolium L. Amer- 
ican Fever-few. Prairie Dock. 


Fig. 4433. 
P. integrifolium L. Sp. Pl. 988. 1753. 

Stem stout, striate, finely pubescent with 
short hairs, or glabrous below, corymbosely 
branched above, 1°-4° high. Rootstocks tuber- 
ous thickened; leaves firm, ovate or ovate- 
oblong, acute or acuminate, crenate-dentate 
or somewhat lyrate at the base, hispidulous 
and roughish on both sides, the lower and 
basal ones petioled, often 12’ long and 5’ wide, 
the upper smaller, sessile; heads numerous 
in a dense terminal corymb; involucre nearly 
hemispheric, about 3” high, its bracts firm, 
the outer oblong, densely appressed-pubes- 
cent, the inner broader, glabrous, or ciliate 


In dry soil, Maryland to Minnesota, south to 
Georgia, Missouri and Arkansas. Cutting-almond. 
Wild quinine. May-Sept. 


3. Parthenium auriculatum Britton. Auri- 
cled Parthenium. Fig. 4434. 


P. auriculatum Britton, in Britt. & Brown, Ill. Fl. 3: 
521. 18098. 


Rootstock an oval erect tuber twice as long as 
thick ; stem villous-pubescent, 13°—23° high. Leaves 
rough above, villous, especially on the veins be- 
neath, oval, ovate or oblong, irregularly crenate- 
dentate, some or all of them laciniate or pinnatifid 
at the base, the basal and lower slender-petioled 
with petiole as long as the blade, or longer, the 
upper with a sessile clasping auricled base, or 
with margined clasping petioles; inflorescence 
densely corymbose, its branches villous-tomen- 
tose; bracts of the involucre densely canescent. 


Mountains of Virginia. Y 


30 


on’the margins; rays white or whitish. 


a pgy 


COMPOSITAE. Vor. ITI. 


4. Parthenium hispidum Raf. Creeping 
or Hairy Parthenium. Fig. 4435. 


Parthenium hispidum Raf. New Fl. N. A. 2: 35. 
1836. 


P. repens Eggert, Cat. Pl. St. Louis 16. 1891. 


Similar to the two preceding species, but 
lower, seldom over 2° high. Rootstocks slen- 
der, forming runners; stem pilose or hispid 
with spreading hairs; leaves hispid on both 
sides, irregularly crenate, sometimes lyrate at 
the base, the teeth rounded and obtuse; heads 
fewer, slightly larger, in a small loose corymb; 

. outer bracts of the involucre proportionately 
broader. 


Barren rocky soil, Missouri to Kansas and 
Texas. April—July. 


56. CRASSINA Scepin, Sched. Acido Veg. 42. 1758. 
[Zinnia L. Syst. Ed. 10, 1221. 1759.] 

Annual or perennal herbs, some species shrubby, with opposite, entire, or sparingly ser- 
rate, mostly narrow and sessile leaves, and large or middle-sized heads of both tubular and 
radiate flowers. Ray-flowers pistillate, yellow, or variegated, persistent on the achene. Disk- 
flowers perfect, fertile; corolla cylindraceous, its lobes villous. Involucre campanulate to 
nearly cylindric, its bracts obtuse, dry, firm, appressed, imbricated in 3 series or more, the 
outer gradually shorter. Receptacle conic or cylindric, chaffy, the chaff subtending and 
enwrapping the disk-flowers. Style-branches elongated, not appendages. Achenes of the 
ray-flowers somewhat 3-angled, those of the disk flattened. Pappus of few awns or teeth. 
[In honor of Paul Crassus, an Italian botanist of the sixteenth century.] 


About 12 species, natives of the United States and Mexico. Type species: Chrysogonum peru- 
vianum L, 


1. Crassina grandiflora ( Nutt.) Kuntze. 
Prairie Zinnia. Fig. 4436. 


Zinnia grandiflora Nutt. Trans. Am. Phil. Soe. (II) 
7: 348. 1841. 


Crassina grandiflora Kuntze, Rev. Gen. Pl. 331. 1891. 


Perennial, woody at the base, tufted, much 
branched, rough, 4’-6’ high. Leaves rather rigid, 
linear to linear-lanceolate, entire, 6’-15” long, 
12” wide, or less, connate at the base, acute or 
acutish, crowded; heads numerous, peduncled, 
terminating the branches, 10’-18” broad; rays 4 
or 5, broad, yellow, rounded, or emarginate, their 
achenes with a pappus of 2 or 4 awns; involucre 
campanulate-cylindric, 3’—4” high; style-branches 
of the disk flowers subulate. 


In dry soil, Kansas and Colorado to Texas, Mexico 
and Arizona. June—Sept. 


57. HELIOPSIS Pers. Syn. 2: 473. 1807. 


Perennial herbs (a tropical species annual), with opposite petioled 3-ribbed leaves, and 
large peduncled terminal and axillary heads of tubular and radiate yellow flowers. Invo- 
lucre hemispheric or broadly campanulate, its bracts oblong or lanceolate, imbricated in 2 
or 3 series. Receptacle convex or conic, chaffy, the chaff enveloping the disk-flowers. Ray- 
flowers pistillate, fertile, the rays spreading, the tube very short, commonly persistent on 
the achene. Disk-flowers perfect, the tube short, the limb elongated, 5-toothed. Anthers 


GENUS 57. THISTLE FAMILY. 467 


entire, or minutely 2-toothed at the base. Style-branches tipped with small hirsute append- 
ages. Achenes thick, obtusely 3-4-angled, the summit truncate. Pappus none, or of 2-4 
teeth, or a coroniform border. [Greek, sun-like.] : 


About 6 species, natives of America. Besides the following, two others occur in the southern 
and southwestern United States. Type species‘! Heliopsis laevis Pers. 


Leaves mostly smooth, or nearly so; pappus none, or of 2-4 stout teeth. 1. H. helianthoides. 
Leaves rough; pappus crown-like, or of 1-3 sharp teeth. 2. H. scabra. 


1. Heliopsis helianthdides (L.) Sweet. 
Ox-eye. False Sunflower. Fig. 4437. 
Buphthalmum helianthoides L. Sp. Pl. 904. 1753. 

Heliopsis iaevis Pers. Syn. 2: 473. 1807. 


Heliopsis helianthoides Sweet, Hort. Brit. 487. 
1827. 


Stem glabrous, branched above, 3°-5° high. 
Leaves opposite, or rarely in 3’s, ovate or 
ovate-lanceolate, rather thin, acuminate at the 
apex, usually abruptly narrowed at the base, 
sharply and nearly equally dentate, smooth on 
both sides, or roughish, 3’-6’ long, 1’-23’ wide; 
heads long-peduncled, somewhat corymbose, 
13-23’ broad; rays 9-12” long, persistent, or 
at length decaying away from the achenes; 
bracts of the involucre oblong or linear-oblong, 
obtuse or acutish, the outer commonly longer 
than the inner; achenes glabrous, the summit 
truncate; pappus none, or of 2-4 short teeth. 


In open places, Ontario to New York, Florida, 
North Dakota, Illinois and Tennessee. July—Sept. 


\\g@Z 

(/ trata gy 
Nis Za 

L$. |. =_=a\ 


Zs 


=i 
2 
2. Heliopsis scabra Dunal. Rough Ox-eye. 


Fig. 4438. 


Heliopsis scabra Dunal, Mem. Mus. Paris 5: 56. pl. 4. 
1819. 

Heliopsis laevis var. scabra T. & G. Fl. N. A. 2: 303. 
1842. 


Wo " L 


Similar to the preceding species, but stem 
rough, at least above, simple or branched, 2°-4° 
high. Leaves ovate or ovate-lanceolate, acute or 
sometimes acuminate, sharply dentate, rough on 
both sides, firm, 2-5’ long, 14’-2’ wide, abruptly 
narrowed at the base, short-petioled; heads few, 
or sometimes solitary, long-peduncled, 2’—23’ 
broad; rays usually 1’ long, or more; bracts of 
the involucre canescent, oblong or linear-oblong; 
achenes pubescent on the margins when young; 
pappus a short laciniate crown, or I-3 sharp teeth. 

Usually in dry soil, Maine to New York, New Jer- 
sey, Manitoba, British Columbia, Arkansas and New 
Mexico. June-Sept. False sunflower. 


58. VERBESINA L. Sp. Pl. gor. 1753. 


Erect or diffuse branching pubescent or hirsute herbs, with opposite leaves, and small 
peduncled terminal and axillary heads of tubular and radiate whitish flowers. Involucre 
hemispheric or broadly campanulate, its bracts imbricated in about 2 series, nearly equal, or 
the outer longer. Receptacle flat or convex, chaffy, the chaff awn-like, subtending the achenes. 
Ray-flowers pistillate, fertile. Disk-flowers perfect, mostly fertile, their corollas tubular, 
4-toothed or rarely 5-toothed. Anthers entire or minutely 2-toothed at the base. Style- 
branches of the disk-flowers with obtuse or triangular tips. Achenes thick, those of the rays 
3-sided or 4-sided, those of the disk compressed. Pappus none, or of a few short teeth. 
[Name changed from Verbena.] 


About 4 species, mostly of tropical distribution, the following typical. 


468 COMPOSITAE. Vo. III. 


1. Verbesina alba L. Yerbadetajo. Fig. 4439. 
Verbesina alba L. Sp. Pl. 902. 1753. 

Eclipta erecta L. Mant. 2: 286. 1771. 

Eclipta procumbens Michx. Fl. Bor. Am. 2: 129. 1803. 
Eclipta alba Hassk. Pl. Jav. Rar. 528. 1848. 


Annual, rough with appressed pubescence, erect or 
diffuse, 63° high. Leaves lanceolate, oblong-lanceo- 
giate or linear-lanceolate, acute or acuminate, denticulate 

or entire, narrowed to a sessile base, or the lower peti- 
/ oled, 1-5’ long, 2’-10” wide; heads commonly numer- 

UY FS ous, 3-6” broad, nearly sessile, or slender-peduncled ; 

Yaa rays short, nearly white; anthers brown; achenes 
¥ 4-toothed, or at length truncate. 

Along streams, and in waste places, Massachusetts to IIli- 

nois, Nebraska, Florida, Texas and Mexico. Naturalized 


from the south in its northeastern range and widely dis- 
tributed in warm regions as a weed. July—Oct. 


59. TETRAGONOTHECA (Dill.) L. Sp. Pl. 903. 1753. 


Erect perennial mostly branched herbs, with opposite, sessile or connate-perfoliate, broad 
dentate leaves, and large peduncled heads of tubular and radiate yellow flowers. Involucre 
depressed-hemispheric, its principal bracts 4, large and foliaceous, inserted in I series; inner 
bracts 6-15, small, subtending the pistillate ray-flowers. Receptacle conic, chaffy, the chaff 
concave, enwrapping the perfect fertile disk-flowers, the corollas of which are slender and 
5-toothed. Anthers entire or minutely 2-toothed at the base. Style-branches of the disk- 
flowers hispid, tipped with elongated appendages. Achenes thick, 4-sided, truncate at the 
summit. Pappus none, or of several short scales. [Greek, 4-angled-case, referring to the 
involucre. | : 


Four known species, natives of the southern United States and northern Mexico, the follow- 
ing typical. 


1. Tetragonotheca helianthoides L. 
Tetragonotheca. Fig. 4440. 


Tetragonotheca helianthoides L. Sp. Pl. 903. 1753. 


Viscidly pubescent; stem branched or sim- 
ple, 1°-23° high Leaves ovate, ovate-oblong, 
or somewhat rhomboid, thin, coarsely and un- 
equally dentate, pinnately veined, acute at the 
apex, narrowed at the sessile or somewliat 
clasping base, 2’-6’ long, 17-3’ wide; heads 
usually few, 14’-3’ broad; involucre 4-angled 
in the bud, its principal bracts broadly ovate, 
acute; rays 6-10, strongly parallel-nerved, 2-3- 
toothed; corolla-tube villous below; achenes 
4-sided, or nearly terete; pappus none. 


In dry soil, Virginia to Tennessee, Florida and 
Alabama. May-June. Sometimes flowering again 
in autumn. 


GENUS 60. THISTLE FAMILY. 469 


60. SPILANTHES Jacq. Stirp. Am. 214. pl. 126. 1763. 


Annual or perennial branching herbs, with opposite, usually toothed leaves and rather 
small, long-peduncled discoid and radiate heads, terminal, or in the upper axils, or rays 
wanting in some species. Involucre campanulate, its bracts in about 2 series, herbaceous, 
loosely appressed. Receptacle convex or elongated, chaffy, its chaff embracing the disk- 
achenes and at length falling away with them. Ray-flowers yellow, or white, pistillate, some- 
times wanting. Disk-flowers yellow, perfect, their corollas tubular with an expanded 4-5-cleft 
limb. Anthers truncate at the base. Style-branches of the disk-flowers long, sometimes 
penicillate at the summit. Ray-achenes 3-sided, or compressed, those of the disk-flowers. 
compressed, margined. Pappus of 1-3 awns, or more. [Greek, spot- or stain-flower, not 
significant. | 

About 30 species, natives of warm and tropical regions. Type species: Spilanthes urens Jacq. 


1. Spilanthes répens ( Walt.) Michx. 
Spilanthes. - Fig. 4441. 


Anthemis repens Walt. Fl. Car. 211. 1788. 
Spilanthes repens Michx. Fl. Bor. Am. 2: 131. 1803. 
S. americana repens A. H. Moore, Proc. Am. Acad. 

42: 547. 1907. I 

Perennial, usually rooting at the lower nodes; 
stem slender, simple or branched, spreading or 
ascending, 82° long, pubescent, or nearly gla- 
brous. Leaves ovate to lanceolate, petioled, acute 
or acuminate at the apex, or the lower obtuse, 
coarsely toothed, or nearly entire, 1’-3’ long; 
heads long-peduncled, solitary at the end of the 
stem and branches, 6’—10” broad; bracts of the 
involucre oblong to oblong-lanceolate, obtuse or 
acute; rays 8-12, yellow; receptacle narrowly 
conic; achenes oblong, most of them roughened 
when mature and hispidulous; pappus of I or 2 
very short awns, or none. 


In moist or wet soil, Missouri to Texas, east to 
South Carolina and Florida. June—Sept. 


61. RUDBECKIA L. Sp. Pl. 906. 1753. 


Perennial or biennial (rarely annual), mostly rigid, usually rough or hispid herbs, with 
alternate undivided lobed or pinnatifid leaves, and large long-peduncled heads of tubular 
(mostly purple) and radiate (yellow) flowers. Involucre hemispheric, its bracts imbricated 
in 2-4 series. Receptacle conic or convex, with chaffy concave scales subtending or envelop- 
ing the disk-flowers. Ray-flowers neutral, the rays entire or toothed. Disk-flowers perfect, 
fertile, their corollas 5-lobed. Anthers entire or minutely 2-mucronate at the base. Style- 
branches tipped with hirsute appendages. Achenes 4-angled, obtuse or truncate at the apex. 
Pappus coroniform, sometimes of 2-4 short teeth, or none. [In honor of Claus Rudbeck, 
1630-1702, Swedish anatomist and botanist.] 


About 30 species, natives of North America and Mexico. In addition to the following, some 20 
others occur in the southern and western United States. Type species: Rudbeckia hirta L. 


Disk globose or ovoid and purple or dark brown in fruit ; lower leaves entire or lobed. 
Lower leaves deeply 3-lobed or 3-divided. 
Plant more or less hirsute; leaves thin; chaff awned. 1. R. triloba. 
Plant scabrous ; leaves thick; chaff blunt, pubescent at apex. 2. R. subtomentosa. 
Leaves neither 3-lobed nor 3-divided. 
Plants hispid ; style-branches subulate. 
Stem leaves lanceolate to oblong; involucre shorter than the rays. 3. R. hiria. 
Stem leaves oval to obovate; involucral bracts foliaceous, nearly as long as the rays. 
4. R. Brittonit. 
Plants pubescent or glabrate; style-branches obtuse. 
Chaff merely ciliate. 
Leaves denticulate or entire; rays 9”—12” long. 
Basal leaves narrowed at base. 5 
Basal leaves cordate at base. 6. R. umbrosa. 
Leaves dentate or laciniate ; rays about 18” long. 7. R. speciosa. 
8 


R. fulgida. 
R 
R 
Chaff canescent. . R. grandiflora. 
R 
R 


Disk elongated or cylindric in fruit, yellowish or gray. 
Leaves very thick, shallowly toothed. 


: 5 had : : 9. R. maxima. 
Leaves thin, pinnately divided or pinnatifid. 10. 


. laciniata. 


470 COMPOSITAE. Vor. III. 


1. Rudbeckia triloba L. Thin-leaved 
Cone-flower. Fig. -4442. 
Rudbeckia triloba L. Sp. Pl. 907. 1753. 

Stem somewhat pubescent and rough, rarely 
glabrate, branched, 2°-5° high. Leaves thin, 
rough on both sides, bright green, the basal 
and lower ones petioled, some or all of them 
3-lobed or 3-parted, the lobes lanceolate or ob- 
long, acuminate, sharply serrate; upper leaves 
ovate, ovate-lanceolate, or lanceolate, acumi- 
nate or acute, narrowed to a sessile base or 
into short margined petioles, serrate or entire, 
2’-4' long, 3’-1’ wide; heads nearly 2’ broad, 
corymbed; bracts of the involucre linear, 
acute; pubescent, soon reflexed; rays 8-12, 
yellow, or the base orange or brownish-purple; ° 
disk dark purple, ovoid, about 6” broad; chaff 
of the receptacle awn-pointed; pappus a mi- 
nute crown. 

In moist soil, New Jersey to Georgia, west to 
Michigan, Minnesota, Missouri, Kansas and Louisi- 
ana. Sometimes escaped from gardens to road- 


“fr 
- “A, 
, 
as 
easy 
LEELA 


Vs 


2. Rudbeckia subtomentdsa Pursh. Sweet 
Cone-flower. Fig. 4443. 
Rudbeckia subtomentosa Pursh, Fl. Am. Sept. 575. 1814. 


Densely and finely cinereous-pubescent and scabrous; 
stem branched above, 2°-6° high. Leaves thick, some 
or all of the lower ones deeply 3-lobed or 3-parted, 
petioled, 3-5’ long, the lobes oblong or lanceolate, acute 
or acuminate, dentate; upper leaves, or some of them, 
lanceolate or ovate, acuminate, sessile or nearly so; 
heads numerous, 2’-3’ broad; rays 15-20, yellow, or 
with a darker base; disc subglobose, rounded, purple 
or brown, 6’-8” broad; bracts of the involucre linear- 
lanceolate, acuminate, squarrose, sweet-scented; chaff 
of the receptacle linear, obtuse or obtusish, pubescent, 
or somewhat glandular at the apex; pappus a short 
crenate crown. 


On prairies and along rivers, Illinois to Louisiana, Kan- 
sas and Texas. July—Sept. 


*y 


3. Rudbeckia hirta L. Black Eyed Susan. 
Yellow Daisy. Fig. 4444. 
Rudbeckia hirta L. Sp. Pl. 907. 1753. 

Hirsute or hispid throughout, biennial or some- 
times annual; stems simple or sparingly branched, 
often tufted, 1°-3° high. Leaves thick, sparingly 
serrate with low teeth, or entire, lanceolate or ob- 
long, the lower and basal ones petioled, mostly ob- 
tuse, 3-5-nerved, 2-7’ long, 4’-2’ wide, the upper 
sessile, narrower, acute or acutish; heads commonly 
few or solitary, 2-4’ broad; rays 10-20, orange or 
orange-yellow, rarely darker at the base; bracts of 
the involucre very hirsute, spreading or reflexed, 
much shorter than the rays; disk globose-ovoid, 
purple-brown; chaff of the receptacle linear, acute 
or acutish, hirsute at the apex; style-tips acute; 
pappus none. 

Prairies and plains, Ontario to Manitoba, Florida, 
Colorado and Texas. Widely distributed in the east as 
a weed, north to Quebec. Races differ in pubescence 
and in length and color of the rays. Nigger- or darkey- 

head. Nigger- or poor-land daisy. Golden-jerusalem. Yel- 
low ox-eye-daisy. English bull’s-eye. Brown daisy or betty. Brown-eyed susan. May-—Sept. 
Rudbeckia monticola Small, of the southern Alleghanies, with broader, ovate, acute or acumi- 
nate stem leaves, is recorded as extending northward into Pennsylvania. 


GENUS OI. THISTLE FAMILY. 471 


4. Rudbeckia Brittonii Small. Britton’s 
Cone-flower. Fig. 4445. 


R. Brittonii Small, Mem. Torr. Club 4: 130. 1894. 


Stem stout, hispid, erect, 13°-22° high, simple, 
grooved, leafy, at least below. Leaves serrate 
or crenate-serrate, strigose-pubescent, the basal 
ones ovate to ovate-lanceolate, 3-4’ long, obtuse, 
long-petioled; stem leaves obovate to oval, often 
with a lateral lobe, the petioles wing-margined ; 
uppermost leaves often ovate-lanceolate, sessile, 
cordate; bracts of the involucre foliaceous, often 
1’ long or more; head 2’-3’ broad; rays about 12, 
2-lobed; outer chaff oblanceolate, the inner linear, 
acute, purple-tipped, fringed with jointed hairs; 
style-tips slender, acute. 


In woods, Pennsylvania to Virginia and Tennessee. 
May-July. 


5- Rudbeckia filgida Ait. Orange or Bril- 
liant Cone-flower. Fig. 4446. 
Rudbeckia fulgida Ait. Hort. Kew. 3: 251. 17809. 
R. spathulata Michx. Fl. Bor. Am. 2: 144. 1803. 
Rudbeckia missouriensis Engelm.; Boynton & Beadle, 
Biltmore Bot. Studies 1: 17. 1901. 
R, palustris Eggert ; Boynton & Beadle, loc. cit. 16. 1901. 
Perennial; stem hirsute, or strigose-pubescent, 
slender, sparingly branched or simple, 1°-3° high. 
Leaves entire or sparingly serrate with distant teeth, 
more or less hirsute or pubescent on both sides, the 
basal and lower ones oblong or spatulate, obtuse, 
2’-4' long, 3-nerved, narrowed into margined petioles, 
the upper lanceolate, oblong-lanceolate, ovate or obo- 
vate, sessile, or slightly clasping at the base; heads 
few, 1’-13’ broad; bracts of the involucre oblong or 
lanceolate, 3-8” long; rays 8-15, linear, bright yel- 
low or with an orange base; disk globose or globose- 
ovoid, brown-purple, 5-7” broad; chaff of the re- 
ceptacle linear-oblong, glabrous, or ciliate at the 
summit; pappus a minute crown. 


In dry soil, New Jersey and Pennsylvania to Florida, 
west to Missouri and Texas. Consists of races differing 
in pubescence and leaf-form. Aug.—Oct. 


6. Rudbeckia umbrosa Boynton & Beadle. 
‘ Woodland Cone-flower. Fig. 4447. 


Rudbeckia umbrosa Boynton & Beadle, Biltmore Bot. 
Studies 1: 16. 1901. 


Pubescent, perennial; stems 14°-33° tall, striate, 
simple or branched. Leaves thin, the basal and 
lower cauline ones ovate, 2’—43’ long, coarsely ser- 
rate, rounded, truncate or cordate at the base, acute 
at the apex, the upper stem leaves diminishing in 
size, short-petioled or subsessile, narrower and less 
toothed than the lower; heads mostly several, showy ; 
rays 8-12, yellow or orange-yellow, 7’-10” long; 
bracts of the involucre oblong to linear-oblong, 
5-74” long, pointed; disk somewhat depressed, 5’— 
74” wide, dark purple; chaff broad, densely ciliate 
at the apex; pappus coroniform. 


In moist soil and woodlands, Kentucky, Tennessee and 
northwestern Georgia. Aug.—Sept. 


472 COMPOSITAE. Vor. III. 


7. Rudbeckia specidsa Wenderoth. Showy 
Cone-flower. Fig. 4448. 

Rudbeckia aspera Pers. Syn. 2: 477. 1807? 
R. speciosa Wendler. Ind. Sem. Hort. Marb. 1828. 

Perennial, more or less hirsute or hispid; stem 
branched above, 1°-4° high. Leaves firm, slender- 
petioled, 2-5’ long, 1-2’ wide, dentate with low teeth, 
acute or sometimes acuminate, 3-5-nerved; stem 
leaves sessile or partly clasping, or narrowed into 
broad margined petioles, laciniate or serrate, lanceo- 
late to ovate, acuminate, often 6’ long, the uppermost 
smaller and sométimes entire; heads several, 2’-3' 
broad; bracts of the involucre linear-lanceolate, 
acute; rays 12-20, 1-13’ long, bright yellow, usually 
orange at the base; disk depressed-globose, 5’—-8” 
broad, brown-purple; chaff of the receptacle obtusish 
or acute, ciliate or naked; pappus a short crown. 

In moist soil, New Jersey to Michigan, south to Ala- 
bama and Arkansas. Aug.—Oct. 

Rudbeckia Sullivantii Boynton & Beadle has been 
separated from R. speciosa on account of its broader 
leaves, larger disk-flowers and larger achenes. 


f 


V 


8. Rudbeckia grandiflora Gmelin. ee 
Large-flowered Cone-flower. iN La 


Ns 
RU Fe" 


Fig. 4449. 
Rudbeckia grandiflora Gmelin; DC. Prodr. 5: 

556. 1836. 

Perennial; stem 13°-3° tall, scabrous or 
hispid throughout. Leaves mainly on the 
lower part of the stem, very rough on both 
surfaces, ovate-elliptic to lanceolate, 23’-6’ 
long, acute or acuminate at the apex, cu- 
neate or more abruptly contracted at the 
base, shallowly serrate or denticulate, the 
petioles of the lower cauline and basal 
leaves as long as the blades or longer; 
heads large, solitary or few, showy; rays 
several, yellow, 17-13’ long, ieee: 
bracts of the involucre linear, acuminate ; 
disk ovoid or ovoid-globose, 3’-1’ thick; 
chaff obtuse, canescent; pappus conspicuous, 
crenate or toothed. 

On dry prairies, Oklahoma to Louisiana and 
Texas; introduced into Missouri. June—Aug. 


g. Rudbeckia maxima Nutt. Great Cone- 
flower. Fig. 4450. 


ay, SS Zine Rudbeckia maxima Nutt. Trans. Am. Phil. Soc. (II) 


ZS 


7 20354 OA. 


Perennial; stem 3°-0° tall, simple or branched 
above, smooth, glaucous. Leaves oblong, oval, 
or ovate, or pandurate, 23’-8’ long, mostly ob- 
tuse, undulate, repand-denticulate or entire, the 
upper sessile and partly clasping; heads large, 
showy; rays several, yellow, 7-20” long; bracts 
of the involucres linear or linear-lanceolate, 
acute, short; disk cylindric to conic-cylindric, 

1’—23’ long; chaff abruptly short-pointed, pubes- 
ee at the summit; pappus conspicuous, den- 
ticulate, accentuated at the angles. 


In moist soil, Missouri to Louisiana and Texas. 
June—Aug. 


GENUS OI. THISTLE FAMILY. 473 


10. Rudbeckia laciniata L. Tall, or Green- 
headed Cone-flower. Fig. 4451. 
Rudbeckia laciniata L, Sp. Pl. 906. 1753. 


Perennial; stem much branched, glabrous, or nearly 
so, 3°-12° high. Leaves rather thin, minutely pubes- 
cent on the margins and upper surface, broad, the 
basal and lower ones long-petioled, often 1° wide, 
pinnately 3-7-divided, the segments variously toothed 
and lobed; stem leaves shorter-petioled, 3-5-parted 
or divided, the uppermost much smaller, 3-lobed, 
dentate or entire; heads several or numerous, 23’—4’ 
broad; rays 6-10, bright yellow, drooping; bracts of 
the involucre unequal; chaff of the receptacle trun- 
cate and canescent at the apex; disk greenish-yellow, 
at length oblong and twice as long as thick or longer; 
pappus a short crown. 


In moist thickets, Quebec to Manitoba, Idaho, Colo- 
rado, Florida and Arizona. Thimble-weed. A double- 
flowered form in cultivation is called golden-glow. July— 
Sept. 

A southern mountain race, lower, often only 1° high, 
with smaller heads, has been described as Rudbeckia 
laciniata humilis A. Gray. 


62. DRACOPIS Cass. Dict. Sci. Nat. 25 27355 1525) 46" 400, | 1827: 


An annual caulescent herb, with smooth and glaucous foliage, and alternate broad thickish. 
entire or slightly serrate, I-ribbed, clasping leaves. Heads radiate, showy. Involucre flat, 
many-flowered, of few narrow, somewhat foliaceous bracts. Receptacle slender, with early 
deciduous chaffy scales. Ray-flowers few, neutral, the rays yellow or often brownish-purple 
at the base. Disk-flowers perfect, fertile, brownish, their corollas 5-lobed. Style-branches 
with small pubescent appendages. Achenes terete or nearly so, not angled, striate and 
minutely transversely wrinkled. Pappus wanting. [Greek, dragon-like, referring to the 
appendaged style-branches. ] 


pic genus of the southeastern United States. 


1. Dracopis amplexicailis (Vahl) Cass. 
Clasping-leaved Cone-flower. Fig. 4452. 


Rudbeckia amplexicaulis Vahl, Act. Havn. 2: 29. pi. 4. 
1783. 
Dracopis amplexicaulis Cass.; DC. Prodr. 5: 558. 1836. 


Annual; glabrous throughout, somewhat glaucous; 
stem branched, grooved, 1°-2° high, the branches 
ascending. Leaves entire or sparingly toothed, 1-ribbed, 
. reticulate-veined, the lower oblong to spatulate, ses- 
sile, the upper ovate, ovate-oblong, or lanceolate, 
acute, cordate-clasping ; heads solitary at the ends of 
the branches, long-peduncled, about 2’ broad; bracts 
of the involucre few, lanceolate, acuminate; rays 
yellow, or sometimes brown at the base; disk ovoid- 
oblong, often becoming 1’ high; achenes not angled, 
obliquely attached to the elongated receptacle; chaff 
at length deciduous. 


In wet soil, Missouri to Oklahoma, Louisiana and 
Texas. June-Aug. 


63. RATIBIDA Raf. Am. Month. Mag. 2: 268. 1818. 


[Lépacuys Raf Journ, Phys. 89: 100. 1819.] 


Perennial herbs, with alternate pinnately divided or parted leaves, and long-peduncled 
terminal heads of tubular and radiate flowers, the disk-flowers gray or yellow, becoming 
brown, the rays yellow, or with brown bases, drooping or spreading. Involucral bracts in 
2 or 3 series. Disk globose, oblong or cylindric. Receptacle columnar to subulate, the con- 
cave chaff subtending or enveloping the disk-flowers, truncate, the tips inflexed, canescent. 
Ray-flowers neutral. Disk-flowers perfect, fertile, their corollas with scarcely any tube. 


474 


COMPOSITAE. Vor. III. 


Achenes short, flattened, sharp-margined, or winged, at length deciduous with the chaff. 


Pappus with 1 or 2 teeth, or none. 


[Name unexplained. | 


About 4 species, natives of North America. Type species: Rudbeckia columnaris Sims. 


Style-tips lanceolate-subulate ; leaf-segments lanceolate; rays 1/3’ long. 1. R. pinnata. 
Style-tips short, blunt ; leaf-segments linear, rays 3”-15” long. 
Disk cylindric, at length 1’ long or more; rays mostly as long, or longer. 2. R.columnaris. 
Disk globose to short-oblong, about 14’ high; rays mostly short. 3. R. Tagetes. 


1. Ratibida pinnata ( Vent.) Barnhart. Gray-headed Cone-flower. Fig. 4453. 


Rudbeckia pinnata Vent. Hort. Cels. pl. 77. 1800. 

Lepachys pinnata T. & G. Fl. N. A. 2: 314. 1842. 

Ratibida pinnata Barnhart, Bull. Torr. Club 24: 410. 
1897. 


Rough and strigose-pubescent throughout; stem 
branched or simple, 3°-5° high. Leaves pin- 
nately 3-7-divided, the basal ones sometimes 10’ 
long, petioled, the segments lanceolate, dentate, 
cleft or entire, acute or acuminate; upper leaves 
sessile or nearly so, the uppermost commonly 
small and entire; bracts of the involucre linear 
or linear-oblong, short, reflexed; rays 4-10, yel- 
low, 1-3’ long, 3-0” wide, drooping; style-tips 
lance-subulate; disk oblong, gray or becoming 
brown, rounded, at length twice as long as thick; 
chaff of the receptacle canescent at the summit; 
achenes compressed, acutely margined, the inner 
margin produced into a short tooth. 

On dry prairies, Ontario and western New York to 
Florida, South Dakota, Nebraska, Kansas and Louisi- 


ana. Adventive eastward to Massachusetts. June- 
Sept. 


2. Ratibida columnaris (Sims) D. Don. Long-headed or Prairie Cone-flower. 


Rudbeckia columnaris Sims, Bot. Mag. pl. 1601. 1813. 
Ratibida columnaris D. Don; Sweet, Brit. Fl. Gard. 2: 


361. 1838. 


Lepachys columnaris T. & G. Fl. N. A. 2: 
Lepachys columnaris var. pulcherrima T. 


1842. 


Strigose-pubescent and scabrous; stem slender, 
usually branched, 1°-23° high. Leaves thick, pin- 
nately divided into linear or linear-oblong, acute or 
obtuse, entire dentate or cleft segments, the cauline 
short-petioled or sessile, 2-4’ long, the basal ones 
sometimes oblong, obtuse and undivided, slender- 
petioled; bracts of the involucre short, linear-lan- 
ceolate or subulate, reflexed; rays 4-10, yellow, 
brown at the base, or brown all over, 4’-15” long, 
drooping; disk gray, elongated-conic or cylindric, 
blunt, at length 3 or 4 times as long as thick; chaff 
of the receptacle canescent at the apex; achenes 
scarious-margined or narrowly winged on the inner 
side; pappus of I or 2 subulate teeth usually with 


several short intermediate scales. 


On dry prairies, Minnesota to Assiniboia, British Co- 
lumbia, Montana, Nebraska, Texas, Mexico and Ari- 
zona. Also in Tennessee. Brush. May—Aug. 


3 
& 


Fig. 4454. 


1842. 


iis} 
G. loc. cit. 


GENUS 63. THISTLE FAMILY. 


3. Ratibida Tagétes (James) Barnhart. 
Short-rayed Cone-flower. Fig. 4455. 


Rudbeckia Tagetes James in Long’s Exp. 2: 68. 1823. 


Lepachys Tagetes A. Gray, Pac. R. R. Rep. 4: 103. 
1856. 


Hanbide Tagetes Barnhart, Bull. Torr. Club 24: 100. 
1897. 

Rough-canescent; stem 1°-13° high, usually 
much branched, leafy. Leaves firm, pinnately 
divided into 3-7 narrowly linear, mostly entire 
segments; peduncles terminal, 4’-2’ long; heads 
1’ broad, or less; bracts of the receptacle narrow, 
deflexed; rays few, mostly shorter than the glo- 
bose to short-oval disk; style-tips obtuse; achenes 
scarious-margined; pappus of I or 2 subulate 
deciduous teeth, with no short intermediate teeth. 

On dry plains and rocky hills, Kansas to Texas, 


Colorado, Chihuahua, New Mexico and Arizona. 
July—Sept. 


64. ECHINACEA Moench, Meth. sot. 1704. 


[BraunerIA Neck. Elem. 1: 17. Hyponym. 1790.] 

Perennial erect branched or simple herbs, with thick black roots, thick rough alternate 
or opposite, 3-5-nerved entire or dentate, undivided leaves, and large long-peduncled heads of 
tubular and radiate flowers, the rays purple, purplish, crimson or yellow, the disk green or 
purple, at length ovoid or conic. Involucre depressed-hemispheric, its bracts lanceolate, 
spreading or appressed, imbricated in 2-4 series. Receptacle conic, chaffy, the chaff carinate 
and cuspidate. Ray-flowers neutral, or with a rudimentary pistil. Disk-flowers perfect, the 
corolla cylindric, 5-toothed. Achenes 4-sided, obpyramidal, thick. Pappus a short dentate 
crown. [Greek, referring to the spiny chaff of the receptacle. ] 


Four species, natives of eastern and central North America. Type species: Echinacea purpurea 
(L.) Moench. 


Rays purple, crimson, pink or white. 


Leaves broad, ovate to ovate-lanceolate, often toothed. 1. E. purpurea. 
Leaves narrow, linear to lanceolate, entire. 
Rays about 1’ long, spreading. 2. E. angustifolia. 
Rays 1%’—-3’ long, drooping. 3. E. pallida. 
Rays bright yellow, drooping. 4. E. paradoxa. 


1. Echinacea purpurea (L.) Moench. 
Purple Cone-flower. -Black Sampson. 


Fig. 4456. 
Rudbeckia purpurea L. Sp. Pl. 907. 1753. 
Echinacea purpurea Moench, Meth. 591. 1794. 


Brauneria purpurea Britton, Mem. Torr. Club 5: 334. 
1894. 


Stem glabrous, or sparingly hispid, usually 
stout, 2°-5° high. Lower and basal leaves slen- 
der-petioled, ovate, mostly 5-nerved, acute or 
acuminate at the apex, abruptly narrowed or 
rarely cordate at the base, commonly sharply 
dentate, 3’-8’ long, 17-3’ wide; petioles mostly 
winged at the summit; upper leaves lanceolate 
or ovate-lanceolate, 3-nerved, sessile or nearly 
so, often entire; rays 12-20, purple, crimson, or 
rarely pale, 14’-3’ long, spreading or drooping. 


In moist, rich soil, Pennsylvania to Alabama, 
Georgia, Michigan, Kentucky, Louisiana and Arkan- 
sas. Called also Red sunflower. July—Oct. 


3. Echinacea pallida (Nutt.) Britton. 
Pale Purple Cone-flower. Fig. 4458. 


Rudbeckia pallida Nutt. Journ. Acad. Phila. 7: 


77. 1834. } 
Brauneria pallida Britton, Mem. Torr. Club 5: 


333. 1804. 

Similar to the preceding species, but often 
taller, sometimes 3° high. Leaves elongated- 
lanceolate or linear-lanceolate, entire; rays 
narrow, linear, elongated, drooping, 12’-3’ 
long, 14”-3’" wide, rose-purple or nearly 
white. 


In dry soil on prairies, Illinois to Michigan, 
May-July. 


Alabama and Texas. 


COMPOSITAE. 


Vou lle 


2. Echinacea angustifolia DC. Narrow- 
leaved Purple Cone-flower. Fig. 4457. 


Echinacea angustifolia DC. Prodr. 5: 554. 1836. 
Brauneria angustifolia Heller, Muhlenbergia 1: 

5. 1900. 

Stem hispid or hirsute, slender, often sim- 
ple, 1°-2° high. Leaves lanceolate, oblong- 
lanceolate, or linear-lanceolate, hirsute, acute 
and about equally narrowed at each end, 
strongly 3-nerved and sometimes with an ad- 
ditional pair of marginal less distinct nerves, 
entire, 3-8’ long, 4”-12” wide, the lower and 
basal ones slender-petioled, the upper short- 
petioled or sessile; heads and flowers similar 
to those of the preceding species, but the rays 
usually shorter, spreading. 


In dry soil, especially on prairies, Minnesota to 
Saskatchewan, Nebraska and Texas. Confused 


with the following species in our first edition. 
Comb. 


June—Oct. 


MWG/7 
ANAS HE 
Zaid 


a 
A 


4. Echinacea paradoxa (Norton) Brit- 

ton. Bush’s Cone-flower. Fig. 4459. 
Brauneria atrorubens Boynton & Beadle, Biltmore 

Bot. Stud. 1: 11. 1901. Not Rudbeckia atro- 

rubens Nutt. 

Brauneria paradoxa Norton, Trans. Acad. St. 

Louis 12: 40. 1902. 

Stem glabrous to somewhat hispid, 1°—23° 
high. Leaves smooth or sparingly rough- 
hairy, lanceolate to linear-lanceolate, the lower 
ones petioled, sometimes 1° long, 2-1’ wide, 
3-5-nerved, the upper smaller and nearly ses- 
sile; involucre about 1’ high and broad, the 
disk-flowers brown; rays bright yellow, droop- 
ing or somewhat spreading, 12’—22’ long. 

Prairies and barren soil, Missouri to Texas. 
June. 


GENUS 65. THISTLE FAMILY. 477 


65. BORRICHIA Adans. Fam. Pl. 2: 130. 1763. 


Fleshy, more or less canescent, branching shrubs of the sea-coast, with opposite entire 
or denticulate, cuneate oblong spatulate or obovate, I-3-nerved leaves, and terminal large 
long-peduncled heads of both tubular and radiate yellow flowers. Involucre hemispheric, its 
bracts slightly unequal, imbricated in 2 or 3 series, the inner ones coriaceous. Receptacle 
convex, chaffy, the chaff rigid, concave, subtending or enwrapping the disk-flowers. Ray- 
flowers pistillate, fertile. Disk-flowers perfect, the corolla tubular, 5-toothed, the style-branches 
elongated, hispid. Anthers dark-colored, entire at the base, or minutely sagittate. Achenes 
of the ray-flowers 3-sided, those of the disk-flowers 4-sided. Pappus a short dentate crown. 
{Named for Olaf Borrick, a Danish botanist. | p 


About 5 species, natives of America. In addition to the 
following typical one, another occursin South Florida. 


1. Borrichia frutéscens (L.) DC. Sea Ox-eye. 
Fig. 4460. 


Buphthalmum frutescens L. Sp. Pl. 903. 1753. 
Borrichia frutescens DC. Prodr. 5: 488. 1836. 


Finely canescent, even when old; stems terete, spar- 
ingly branched, 1°-4° high. Leaves mostly erect or 
ascending, lanceolate, spatulate or obovate, obtuse or 
acutish and mucronulate at the apex, fleshy, tapering 
to the sessile base, somewhat connate, 1’-3’ long, 2’’-7” 
wide; heads solitary or few, about 1’ broad; rays 15-25, 
rather short; exterior bracts of the involucre ovate and 
somewhat spreading, the inner ones and the chaff of 
the receptacle cuspidate. 


Sea-coast, Virginia to Florida and Texas. Also on the 
coasts of Mexico and in Bermuda. April—Oct. 


66. HELIANTHUS [Vaill.] L. Sp. Pl. 904. 1753. 


Erect, annual or perennial, mostly branched herbs, with opposite or alternate, simple 
leaves, and large peduncled corymbose or solitary heads of both tubular and radiate flowers, 
the rays. yellow, the disk yellow, brown, or purple. Involucre hemispheric, or depressed, its 
bracts imbricated in several series. Receptacle flat, convex or conic, chaffy, the chaff sub- 
tending the disk-flowers. Ray-flowers neutral (in our species), the rays spreading, mostly 
entire. Disk-flowers perfect, fertile, the corolla tubular, the tube short, the limb 5-lobed. 
Anthers entire, or minutely 2-toothed at the base. Style-branches tipped with hirsute append- 
ages. Achenes thick, oblong or obovate, compressed, or somewhat 4-angled. Pappus of 
2 scales or awns, or sometimes with 2-4 additional shorter ones, deciduous. [Greek, sun- 
flower. | 

About 70 species, natives of the New World. Besides the following, about 30 others occur in 
the southern and western parts of North America, and hybrids may exist. Type species: Helianthus 


annuus L. 
A. Annual species; disk brown or purple; heads large. 
_ Leaves dentate; bracts ovate to ovate-lanceolate, strongly ciliate. 1. H. annuus. 
Leaves mostly entire; bracts lanceolate to ovate-lanceolate, canescent, sometimes ciliate. 
2. H. petiolaris. 
B. Perennial species. 
* Disk purple or purple-brown. 
Leaves narrowly linear or the lower broader, many of them alternate. 


Stem rough ; leaves linear or linear-lanceolate. as angustifolius. 
Stem smooth; leaves elongated-lanceolate. 4. orgyalis. 
Leaves lanceolate to ovate, mainly opposite. 
Leaves hispid, rather thin, abruptly contracted into winged petioles. 5. H. atrorubens. 
Leaves firm in texture, gradually narrowed into petioles. 
Leaves lanceolate; bracts acute or obtusish. 6. 


Leaves rhombic-ovate to rhombic-lanceolate; bracts acute or acuminate. 


Fe subrhomboideus. 


** Disk yellow or yellowish; receptacle convex or conic. 
¢ Leaves nearly all basal or near the base ; upper bract-like. 8. 
tt Stem leafy ; leaves alternate or opposite. 
1. Leaves prevailingly lanceolate and 3-8 times as long as wide. 
a. Leaves glabrous on both sides ; heads 1’—1%’ broad. 9. H. laevigatus. 
b. Leaves scabrous, at least on the upper surface. 
Heads 1’-1%4’ broad; leaves thin; stem glabrous. 10. H. microcephalus. 
Heads 11%4’—3%’ broad; leaves firm, many of them often alternate. 
Stem scabrous, scabrate or hispid; leaves sessile or nearly so. 
Leaves lanceolate, scabrous above, hirsute beneath, flat. 11. H. giganteus. 


EP 
H. 
H. scaberrimus. 
(eh 
H. 


occidentalis. 


COMPOSITAE. Vo. III. 


Leaves very scabrous on both sides. 
Leaves elongated-lanceolate, conduplicate, pinnately-veined. 


Heads numerous; leaves mostly alternate. 12. H. Maximiliani. 
Heads only 1 or 2; leaves, all but the upper, opposite. 13. H. Dalyi. 
Leaves lanceolate to ovate-lanceolate, flat, 3-nerved. 14. H. subtuberosus. 
Stem glabrous; leaves sessile by a truncate base. 17. H. divaricatus. 
Stem glabrous; leaves petioled. 
Leaves lanceolate, firm, canescent and pale beneath. 15. H. grosse-serratus. 
Leaves linear-lanceolate, thin, green on both sides. 16. H. Kellermani. 
2. Leaves prevailingly ovate, ovate-lanceolate, or oblong. 
a. Leaves sessile, or very nearly so. 
Stem glabrous ; leaves divaricate. 17. H. divaricatus. 
Stem hirsute or hispid; leaves ascending. 
Leaves cordate-clasping at the base. 18. H. mollis. 
Leaves narrowed from below the middle. 19. H. doronicoides. 
b. Leaves manifestly petioled. 
Stem puberulent or glabrous. ; 
Leaves membranous or thin, slender-petioled, sharply serrate. 20. H. decapetalus. 
Leaves firmer, shorter-petioled, less serrate or entire. 
Bracts of the involucre much longer than the disk. 21. H. tracheliifolius. 
Bracts of the involucre about equalling the disk. 22. H. strumosus. 
Stem hirsute, hispid, or scabrous. 
Leaves rounded or truncate at the base, short-petioled. 23. H. hirsutus. 
Leaves, at least the upper, narrowed at the base. 3 
Bracts of the involucre ovate-lanceolate, appressed. 24. H.lae iflorus. 
Bracts of the involucre lanceolate-acuminate, spreading. 
Leaves villous-pubescent beneath. 25. H. tomentosus. 
Leaves scabrous or puberulent beneath. 26. H. tuberosus. 


2. Helianthus petiolaris Nutt. Prairie Sun- 
flower. Fig. 4462. 


H., petiolaris Nutt. Journ. Acad. Phila. 2: 115. 1821. 
Hi. aridus Rydb. Bull. Torr. Club 32: 127. 1905. 


Annual, similar to the preceding species, but smaller 


1. Helianthus annuus L. Common Sunflower. 
Fig. 4461. 


Helianthus annuus L. Sp. Pl. 904. 1753. 
Helianthus lenticularis Dougl. Bot. Reg. pl. 1265. 18209. 


Stem hispid or scabrous, stout, branched above, 3°-6° 
high, or in cultivated races sometimes 15° high. Leaves 
all but the lower alternate, broadly ovate, petioled, 
3-nerved, dentate or denticulate, acute at the apex, 
rough on both sides, sometimes pubescent beneath, the 
lower cordate at the base, 3-12’ long; heads in the wild 
plant 3-6’ broad; disk dark purple or brown, 10-2’ 
broad; involucre depressed, its bracts ovate to ovate- 
lanceolate, usually long-acuminate or aristate, hispid- 
ciliate; chaff of the flat receptacle 3-cleft; achenes 
obovate-oblong, appressed-pubescent, or nearly glabrous. 


On prairies, etc., Minnesota to North Dakota, Idaho, Mis- 
souri, Texas and California. Recorded north to Saskatche- 
wan. Much larger in cultivation; an occasional escape in 
the east. Gold. Golden. Larea-bell. Comb-flower. Its flowers 
yield honey and a yellow dye; its leaves fodder; its seeds, 
an oil and food; and its stalks a textile fibre. July—Sept. 


and with smaller heads; stem strigose-hispid or hir- 
sute, 1°-3° high. Leaves all but the lowest alternate, 
petioled, oblong, ovate, or ovate-lanceolate, rough on 
both sides, usually paler beneath than above, sometimes 
canescent beneath, 1-3’ long, entire, or denticulate, ob- 
tuse or acutish at the apex, mostly narrowed at the 
base; heads 13’-3’ broad; disk brown, mostly less than 
10” broad; involucre depressed-hemispheric, its bracts 
lanceolate or oblong-lanceolate, densely canescent, some- 
times hispid-ciliate, acute or short-acuminate; achenes 
villous-pubescent, at least when young. 


On dry prairies, Minnesota to Saskatchewan, Oregon, 
Iowa, Missouri, Texas and California. Found rarely in 
waste places farther east. Races differ in leaf-form, size 
and pubescence. June—Sept. 


GENUS 66. THISTLE FAMILY. 


3- Helianthus angustifolius L. Narrow-leaved 
or Swamp Sunflower. Fig. 4463. 
Helianthus angustifolius L. Sp. Pl. 906. 1753. 


Perennial by slender rootstocks ; stems branched above. 
or simple, slender, rough or roughish above, often hir- 
sutes below, 2°-7° high. Leaves firm, entire, sessile, 
linear, slightly scabrous, rarely somewhat canescent be- 
neath, 2-7’ long, 2”-3” wide, the margins revolute when 
dry, the upper ones all alternate, the lower opposite; 
heads usually few, sometimes solitary, 2-3’ broad; in- 
volucre hemispheric, its bracts linear-lanceolate, acute 
or acuminate, scarcely squarrose, pubescent; receptacle 
slightly convex; disk purple; chaff entire or 3-toothed ; 
rays 12-20; achenes truncate, glabrous; pappus usually 
of 2 short awns. 


In swamps, Long Island, N. Y., to Florida, Kentucky and 


ZA, 


Texas, mainly near the coast. 


5. Helianthus atrorubens L. Hairy Wood Sunflower. Purple-disk Sunflower. 


Aug.—Oct. 


4. Helianthus orgyalis DC. Linear-leaved 
Sunflower. Fig. 4464. 
Hi, giganteus var. crinitus Nutt. Gen. 2: 177. 1818? 
Helianthus orgyalis DC. Prodr. 5: 586. 1836. 
Perennial by slender rootstocks; stems glabrous, 
branched near the summit, very leafy to the top, 
6°-10° high. Leaves sessile, entire, linear or nearly 
filiform, or the lowest lanceolate, remotely dentate 
and short-petioled, rough with mucronate-tipped 
papillae, especially on the lower surface, acuminate, 
4-16’ long, 1-4” wide, the upper all alternate and 
I-nerved, the lower commonly opposite; heads nu- 
merous, about 2’ broad, terminating slender branches; 
involucre nearly hemispheric, its bracts linear-subu- 
late to lanceolate, acuminate, squarrose, ciliate; disk 
purple or brown; receptacle convex, its chaff entire, 
or toothed, slightly ciliate; rays 10-20; achenes ob- 
long-obovate, glabrous, 23’-3” long, 2-4-awned. 


On dry plains, Missouri and Nebraska to Colorado 
and Texas. Sept.—Oct. 


Fig. 4465. 


Helianthus atrorubens L. Sp. Pl. 906. 1753. 


Perennial; stems hirsute below, often minutely pu- 
bescent above, branched at the summit, 2°-5° high. 
Leaves hirsute on both sides, or canescent beneath, 
mostly thin, ovate or ovate-lanceolate, acutish, con- 
tracted near the base into margined petioles, sometimes 
subcordate, dentate or crenate-dentate, 4-10’ long, 1-4’ 
wide, the lower opposite, the upper few, distant, small, 
mainly alternate; heads not numerous, slender-pedun- 
cled, about 2’ broad; involucre hemispheric, its bracts 
oblong to obovate, obtuse, ciliolate, appressed; disk 
purple; recptacle convex, its chaff acute, entire, or 
3-toothed; rays 10-20; achenes obovate, truncate, finely 
pubescent, about 2” long; pappus usually of 2 lanceo- 
late awns. 


In dry woods, Virginia to Florida, west to Ohio, Mis- 
souri, Arkansas and Louisiana. Aug.—Oct. 


48o COMPOSITAE. Vor. ill. 


6. Helianthus scabérrimus Ell. Stiff Sun- 
flower. Fig. 4466. 


H. scaberrimus Ell. Bot. S. C. & Ga. 2: 423. 1824. 
Hi. rigidus Desf. Cat. Hort. Paris, Ed. 3, 184. 1829. 
Perennial; stems simple or little branched, hispid 
or scabrate, 1°-8° high. Leaves thick, coriaceous, 
serrate or serrulate, very scabrous on both sides, 2-7’ 
long, 2’-2’ wide, acute at the apex, narrowed at the 
base, the lower ovate or ovate-oblong, petioled, the 
upper lanceolate or oblong-lanceolate, sessile or short- 
petioled, all opposite, or the uppermost bract-like and 
alternate; heads solitary or few, 2’-3’ broad; invo- 
lucre hemispheric, its bracts ovate, acute or obtusish, 
ciliate, appressed; disk purple or brown; receptacle 
convex, its chaff obtuse; rays 15-25, light yellow; 
achenes more or less pubescent, oblong-obovate; 
pappus of 2 broad scales or of 2-4 stout awns. 
Prairies, Illinois to Minnesota, lowa, Kansas, Georgia 
and Texas. Aug.—Sept. 


7. Helianthus subrhomboideus Rydb. Sit 
Rhombic-leaved Sunflower. Fig. 4467. 


Helianthus subrhomboideus Rydb. Mem. N. Y. Bot. 
Gard. I: 419. 1900. 


Stem simple, sparingly hirsute, usually tinged 
with red. Leaves opposite, firm, very scabrous, 
3-nerved, slightly serrate, the basal ones broad- 
ly ovate to obovate-spatulate, those of the 
stem rhomboid-ovate to rhomboid-lanceolate, 
short-petioled, 2-4’ long, the uppermost very 
small; heads 1-3, 1’-1%’ in diameter; bracts i 
of the involucre oblong, acutish, densely white- } 
ciliate; disk purple. ; 


| 
YY YY 
SSS 


re 


Plains, Manitoba and Saskatchewan to South 
Dakota, Nebraska, Montana and New Mexico. 
Adventive, New Jersey to New Hampshire. July— 
Sept. Included in H. scaberrimus in our first 
edition. 


8. Helianthus occidentalis Riddell. Few-leaved 
Sunflower. Fig. 4468. 


H. occidentalis Ridd. Suppl. Cat. Ohio Pl. 13. 1836. 

H. illinoensis Gleason, Ohio Nat. 5: 214. 1904. 

H, occidentalis illinoensis Gates, Bull. Torr. Club 37: 81. 
1910. 


Perennial; stems appressed-pubescent or sometimes 
nearly glabrous, slender, mostly simple, 1°-3° high. 
Leaves mostly basal, or below the middle of the stem, 
firm, ovate or oblong-lanceolate, obtuse or obtusish at 
the apex, narrowed at the base, 3-5-nerved, serrulate 
or entire, scabrous above, pubescent beneath, with 
slender petioles about as long as the blades; stem usu- 
ally bearing I or 2 pairs of small distant leaves; heads 
several or solitary, 13’-23’ broad; involucre hemi- 
spheric, its bracts lanceolate or ovate-lanceolate, acute 
or acuminate, generally ciliate, appressed; receptacle 
convex, its chaff acute; disk yellow; rays 12-15; 
achenes truncate and pubescent at the summit; pappus 
of 2 lanceolate-subulate awns. 

In dry soil, Ohio to Minnesota, south to Florida and Missouri. Aug.—Sept. 

Helianthus Dowellianus M. A. Curtis, which differs in being stouter, the stem leafy, the leaves 
merely puberulent, and ranges from the District of Columbia to Georgia, appears to be a race 
of this species. 


GENus 66. THISTLE PAMILY. 


g. Helianthus laevigatus T. & G. Smooth A 
Sunflower. Fig. 44609. Xa 


Helianthus laevigatus T. & G. Fl. N. A. 2: 330. 1842. 


Stems slender, from a perennial root, simple or 
little branched, glabrous, or somewhat glaucous, 
2°-6° high. Leaves nearly all opposite, firm, gla- 
brous, lanceolate, short-petioled, or the upper sessile, 
serrulate or entire, pale beneath, acuminate, nar- 
nowed at the base, 3-6’ long, 4’-12’ wide, the mar- 
gins sometimes roughish-ciliate; heads few or soli- 
tary, 1-12’ broad; involucre campanulate, its bracts 
lanceolate or ovate-lanceolaté, firm, ciliolate, the 
tips of the outer ones spreading; rays 5-10; disk 
yellow; chaff linear; achenes slightly pubescent at 
the summit; pappus of 2 lanceolate or ovate owns, 
with or without 2 intermediate scales. 


In dry soil, mountains of Virginia, West Virginia 
and North Carolina. Aug.—Oct. 


10. Helianthus microcéphalus T. & G. 
Small Wood Sunflower. Fig. 4470. 
Helian‘hus parviflorus Bernh.; Spreng. Syst. 3: 617. 

1826. Not H.B.K. 1820. 
H. microcephdus T.& G. Fl. N. A. 2: 329. 1842. 

Stems slender, glabrous, branched above, or 
rarely simple, 3°-6° high. Leaves thin or thin- 
nish, petioled, most of them opposite, lanceolate, 
or the lower ovate-lanceolate, rough above, canes- 
cent or puberulent beneath, long-acuminate at the 
apex, narrowed at the base, serrulate, or the 
lower serrate, 3-7’ long, 3’-13’ wide; heads com- 
monly several or numerous, 17-1?’ wide, borne 
on slender, sometimes roughish peduncles; invo- 
lucre campanulate, 4’—5” broad, its bracts lanceo- 
late or ovate, acute or acuminate, ciliolate, the 
tips of the outer ones spreading; chaff of the 
receptacle oblong, entire, or 3-toothed; rays 5-10; 
disk yellow; achenes nearly glabrous; pappus 
usually of 2 subulate awns. 

In moist woods and along streams, Pennsylvania to 
ere west to Ohio, Missouri and Louisiana. July— 

ept. 


11. Helianthus gigantéeus L. Tall, Giant or 
Wild Sunflower. Fig. 4471. 
Helianthus giganteus L. Sp. Pl. 905. 1753. 


Perennial by fleshy roots and creeping rootstocks ; 
stems hispid or scabrous, at least above, branched near 
the summit, or simple, 3°-12° high. Leaves sessile or 
short-petioled, firm, lanceolate, very rough above, 
rough-pubescent beneath, serrate or denticulate, acumi- 
nate at the apex, narrowed at the base, many or all of 
the upper ones alternate but sometimes all opposite, 
2’-6’ long, 4-1’ wide; heads usually several, mostly 
long-peduncled, 14’-23’ broad; involucre hemispheric, 
its bracts squarrose, lanceolate-subulate, hirsute or 
ciliate, commonly as long as the diameter of the yel- 
lowish disk; chaff of the receptacle oblong-linear, 
acute; achenes oblong, glabrous; rays 10-20; pappus 
of 2 subulate awns. 

In swamps and wet meadows, Maine and Ontario to Sas- 
katchewan, Florida, Louisiana and Colorado. Stem com- 
monly purple. Aug.—Oct. 

Helianthus ambiguus (A. Gray) Britton, differing in 
having all but the uppermost leaves opposite and rounded 
at the base, found on Long Island, New York, and recorded 
from Georgia, appears to be a hybrid, with this species one of its parents. 


31 


482 COMPOSITAE. Vot. ILI. 


12. Helianthus Maximiliani Schrad. Maxi- 
milian’s Sunflower. Fig. 4472. 


Helianthus Maximiliani Schrad. Ind. Sem. Hort. Goett. 
1835. 


Perennial by fleshy roots and thickened rootstocks; 
stems stout, scabrous or hispid below, 2°-12° high. 
Leaves sessile or short-petioled, long-lanceolate, fold- 
ing in drying, alternate or the lower opposite, very 
rough on both sides, rigid, acuminate or acute at 
both ends, denticulate or entire, 3-7’ long, 3-13’ 
wide; heads few or numerous, 2-3’ broad on stout 
densely rough-pubescent peduncles; involucre hemi- 
spheric, its bracts lanceolate, acuminate, squarrose, 
densely strigose-pubescent, often 9” long; disk yel- 
lowish; rays 15-30; chaff linear, acute, pubescent 
above; achenes linear-oblong, glabrous or nearly so; 
pappus commonly of 2 lanceolate awns. 


On dry prairies, Minnesota and Manitoba to Sas- 
katchewan, Missouri, Nebraska and Texas. Locally 
adventive eastward. Aug.—Oct. 


13. Helianthus Dalyi Britton. Judge 
Daly’s Sunflower. Fig. 4473. 


Helianthus Dalyi Britton, Journ. N.-Y. Bot. 
Gard. 2: 89. 1901. 


Perennial by a fusiform tuber 17-13’ 
long; stem slender, simple, roughish, ap- 
pressed-pubescent, about 2° high. Leaves 
firm, conduplicate, drooping, finely rough- 
pubescent on both sides, narrowly lanceo- 
late, acuminate at both ends, distantly ser- 
rate with low teeth, the larger about 4’ 
long and 7” wide, all opposite or the up- 
permost alternate; heads usually solitary 
and long-peduncled, rarely 2, about 32’ 
broad; involucral bracts narrowly lanceo- 
late with long subulate tips; rays acute; 
pappus 2 subulate awns; chaff of the re- 
ceptacle pubescent at the acute apex. 


Ona dry bank, Sag Harbor, New York. Sept. 


14. Helianthus subtuberdosus Bourgeau. 
Indian Potato. Fig. 4474. 


H. giganteus subtuberosus (Bourgeau) Britton, in 
Britt. & Brown, Ill. Fl. 3:°425. 1898. 

H. subtuberosus Bourgeau; Britton, Manual 993. 
1901. 


Fleshy roots, thick and edible. Leaves thick, 
more or less serrate, all or all but the upper dis- 
tinctly petioled, sometimes all opposite, 23’-5’ 
long, acuminate at the apex, mostly narrowed at 
the base, the petioles ciliate; heads not very nu- 
merous; bracts of the involucre lanceolate, acumi- 
nate, densely pubescent and white-ciliate, about 
8” long; rays 1-1?’ long. 


Dry soil, Michigan and Minnesota to Saskatche- 
wan, Montana and Wyoming. Aug.—Sept. 


Helianthus Rydbergi Britton, of western Ne- 
braska, differs by broader, ovate-lanceolate leaves, 
rather abruptly narrowed at the base. 


GENUs 66. THISTLE FAMILY. 483 


15. Helianthus grdésse-serratus Martens. 
Saw-tooth Sunflower. Fig. 4475. 


Helianthus grosse-serratus Martens, Sel. Sem. Hort. 
Loven. 1839. 


Perennial by fleshy roots and slender rootstocks ; 
stems glabrous, glaucous, branched above, 6°-10° 
high, the branches usually strigose-pubescent. Leaves 
long-lanceolate, slender-petioled, the upper alternate, 
the lower opposite, long-acuminate, narrowed at the 
base, sharply serrate, or merely denticulate, the up- 
permost entire, rough above, densely puberulent or 
canescent beneath, 4’-8’ long, 4’-1’ wide; heads sev- 
eral or numerous, 14’-3’ broad; involucre hemi- 
spheric, its bracts squarrose, narrowly lanceolate, 
acuminate, hirsute; chaff linear-oblong, pubescent at 
the summit, acute, often 3-toothed; disk yellowish; 
rays 10-20, deep yellow; achenes nearly glabrous; 
pappus of 2 lanceolate awns. 


In dry soil, Maine to Long Island, Pennsylvania, On- 
tario, South Dakota, Missouri, Kansasand Texas. Aug.— 
Oct. 


16. Helianthus Kellermani Britton 
Kellerman’s Sunflower. Fig. 4476. 


Helianthus Kellermani Britton, Manual 994. 1901. 


Stem 6°-10° high, very smooth, much branch- 
ed above, the branches slender. Leaves nar- 
rowly elongated-lanceolate to linear-lanceolate, 
drooping, rather thin, distantly serrate with 
very small teeth, long-acuminate at the apex, 
attenuate at the base into short petioles or the 
upper sessile, scabrate and sparingly pubes- 
cent on both surfaces, pinnately veined, the 
lower about 8’ long and 7” wide; branches of 
the inflorescence pubescent; bracts of the in- 
volucre linear-lanceolate, about 7” long, and 1” 
wide or less at the base, ciliate, long-acumi- 
nate; rays golden-yellow, 1-12’ long; chaff of 
the receptacle linear. 


Dry soil, vicinity of Columbus, Ohio. Aug.— 
Sept. 


17. Helianthus divaricatus L. Rough or 
Woodland Sunflower. Fig. 4477. 
Helianthus divaricatus L. Sp. Pl. 906. 1753. 


Perennial by slender rootstocks; stem glabrous 
throughout, or pubescent at the summit, slender, 
2°-7° high. Leaves usually all opposite, divaricate, 
lanceolate or ovate-lanceolate, firm, dentate or den- 
ticulate, 3-nerved, rough above, pubescent beneath, 
sessile or nearly so by a truncate base, tapering 
gradually to the long-acuminate apex, 3’-8’ long, 
+’-13’ wide; heads few or solitary, about 2’ broad, 
borne on strigose-pubescent peduncles; involucre 
hemispheric, its bracts lanceolate or ovate-lanceo- 
late, strigose or hirsute, the outer ones spreading; 
disk yellow; rays 8-15; chaff of the receptacle 
apiculate; achenes glabrous; pappus of 2 short 
subulate awns. 


In dry woodlands, Maine and Ontario to Manitoba, 
Nebraska, Florida and Louisiana. July—Sept. 


484 COMPOSITAE. Vor. III. 


18. Helianthus mollis Lam. Hairy Sun- 
flower. Fig, 4478. 
«Helianthus mollis Lam. Encycl. 3: 85. 17809. 


Perennial; stem stout, simple or sparingly’ branched 
above, densely hirsute, 2°-4° high. Leaves ‘ovate or 
, ovate-lanceolate, closely sessile and somewhat clasp- 
ing by a broad cordate base, pinnately veined, 3- 
nerved above the base, all opposite, or the upper 
alternate, ascending, acute or acuminate, scabrous or 
cinereous-pubescent above, densely and finely pubes- 
cent beneath, serrulate, 2-5’ long, 1-23’ wide; heads 
solitary or few, 2’-3’ broad; involucre hemispheric, 
its bracts lanceolate, acuminate, densely villous-ca- 
nescent, somewhat spreading; disk yellow; rays I5- 
25; chaff canescent at’ the summit; mature achenes 
nearly glabrous; pappus of 2 lanceolate scales. 


In dry barren soil, Massachusetts to New Jersey, Geor- 
gia, Iowa, Missouri, Kansas and Texas. Aug.—Sept. 


a= 


1g. Helianthus doronicoides Lam. Oblong- 
leaved Sunflower. Fig. 4479. 
Helianthus doronicoides Lam. Encycl. 3: 84. 1789. 


Perennial by slender rootstocks; stems stout, rough, We NSS 


or finely rough-pubescent, branched above, 3°-7° high. 
Leaves oblong, ovate-oblong, or ovate, thick, ascend- 
ing, serrate or serrulate, acute or acuminate at the 
apex, narrowed at or below the middle to a sessile or 
slightly clasping base, rough on both sides, or finely 
pubescent beneath, 3-nerved, 4’-8’ long, 1’-22’ wide; 
heads commonly numerous, 23-4’ broad; involucre 
hemispheric, its bracts lanceolate, acuminate, pubescent 
or hirsute, somewhat spreading; disk yellow; rays 12- 
20, 4-6” wide; achenes glabrous; pappus of 2 subu- 
late awns. 


A 


In dry soil, Ohio to Missouri and Arkansas. Recorded 
from Michigan. Aug.—Sept. ; 


20. Helianthus decapétalus L. Thin-leaved or Wild Sunflower. Fig. 4480. 


Hleianthus decapetalus L. Sp. Pl. 905. 1753. 


Perennial by branched, sometimes thickened root- 
stocks; stem slender, glabrous or nearly so, branched 
above, 1°-5° high; branches puberulent. Leaves 
thin or membranous, 3-8’ long, 1-3’ wide, ovate or 
ovate-lanceolate, the lower all opposite and slender- 
petioled, the upper commonly alternate, all usually 
sharply serrate, roughish above, finely but often 
sparingly pubescent beneath, acuminate, the rounded 
or truncate base decurrent on the petiole; heads nu- 
merous, 2-3’ broad; involucre hemispheric, its bracts 
linear-lanceolate, acuminate, hirsute, long-ciliate, 
spreading, often longer than the yellow disk; rays 
8-15, light yellow; chaff entire or 3-toothed, pubes- 
cent at the apex; achenes glabrous; pappus of 2 
subulate awns. 


In moist woods and along streams, Quebec to Michi- 
gan, Georgia, Tennessee and Missouri. Aug.—Sept. 


Helianthus scrophulariaefOlius Britton, from near 
Woodlawn, New York, differing by laciniate-serrate 
leaves, is probably a race of this species. 


GENUS 66. THISTLE FAMILY. 485 


21. Helianthus tracheliifolius Mill. Throatwort 
Sunflower. Fig. 4481. 


Helianthus tracheliifolius Mill. Gard. Dict. Ed. 8, No. 7. 
1768. 


Similar to the following species, but the stem usually 
roughish-pubescent above. Leaves short-petioled, ovate- 
lanceolate, or lanceolate, 3-nerved, green both sides, but 
darker above, generally rougher on the upper surface, 
the lower ones sharply serrate; branches and peduncles 
scabrous; heads several, 23’-32’ broad; bracts of the 
hemispheric involucre linear-lanceolate, long-acuminate, 
ciliate and puberulent, longer than the yellow disk, 
sometimes foliaceous, and 3 times its length. 


In dry soil, Connecticut to Pennsylvania, North Carolina, 
Ohio, Minnesota and Arkansas. Aug.—Sept. 


22. Helianthus strumosus L. Pale-leaved 
Wood Sunflower. Fig. 4482. 


Helianthus strumosus L. Sp. Pl. 905. 1753. 
H. mollis Willd. Sp. Pl. 3: 2240. 1804. Not Lam. 1789. 
Hi. macrophylius Willd. Hort. Berol. pl. 70. 1806. 


Perennial by branched, sometimes tuberous-thick- 
ened rootstocks; stem glabrous below, sometimes 
glaucous, 3°-7° high, branched above, the branches 
usually pubescent. Leaves short-petioled, ovate or 
ovate-lanceolate, rarely lanceolate, not membranous, 
rough above, pale and somewhat puberulent or ca- 
nescent beneath, serrate, serrulate, or nearly entire, 
acuminate, contracted much below the middle and 
decurrent on the petiole, 3-nerved above the base, 
3-8’ long, 17-23’ wide, mostly opposite, the upper 
often alternate; heads commonly several, 23-4’ 
broad; involucre hemispheric, its bracts lanceolate 
or ovate-lanceolate, acuminate, ciliate, equalling or 
a little longer than the diameter of the yellow disk; 
rays 5-15; chaff pubescent; achenes nearly glabrous. 


In dry woods and on banks, Maine and Ontario to 
Minnesota, Georgia, Tennessee and Arkansas. July— 
Sept. Races differ in leaf-form and texture. A hybrid 
with H. decapetalus has been described. 


23. Helianthus hirsttus Raf. Stiff-haired 
Sunflower. Fig. 4483. 


Helianthus hirsutus Raf. Ann. Nat. 14. 1820. 
Helianthus hirsutus trachyphyllus T. & G. Fl. N. A. 2: 
329. 1842. 


Perennial; stem densely hirsute, usually branched 
above, stout, rigid, 2°-4° high. Leaves ovate-lanceo- 
late or the upper lanceolate, rounded, truncate or 
subcordate at the base, acuminate at the apex, thick, 
very rough above, rough-pubescent beneath, 3-nerved, 
serrate or serrulate, 3-6’ long, 4-2’ wide, short- 
petioled, or the uppermost sessile, nearly all oppo- 
site, the petioles of the lower ones 4’—}’ long; heads 
usually several, 2-33’ broad; involucre hemispheric, 
its bracts lanceolate, or ovate-lanceolate, acuminate, 
ciliate, mostly puberulent, erect or somewhat spread- 
ing, equalling or shorter than the yellow disk; rays 
12-15; chaff obtusish, pubescent at the summit; 
achenes oval, rounded at the summit, glabrous; 
pappus of I or 2 subulate awns. 


In dry soil, Pennsylvania and Ohio to Wisconsin, Ne- 
braska, West Virginia, Georgia and Texas. July—Oct. 


486 COMPOSITAE. Vor. III. 


24. Helianthus laetiflorus Pers. Showy Sun- 
flower. Fig. 4484. 
Helianthus laetiflorus Pers. Syn. 2: 476. 1807. 

Perennial; stem scabrous or hispid, leafy, 4°-8° high. 
Leaves oval-lanceolate or ovate-lanceolate, short-peti- 
oled, 3-nerved, rough on both sides, narrrowed at the 
base, acute or acuminate at the apex, serrate or serru- 
late, 4/-10’ long, 4’-13’ wide, the upper often alter- 
ynate; heads usually several, 2-4’ broad, mostly short- 
peduncled; bracts of the hemispheric involucre ovate- 
lanceolate, or oblong-lanceolate, imbricated in only 2 or 
3 series, ciliate, otherwise nearly glabrous, appressed or 
but little spreading, shorter than or equalling the yellow 
disk; rays 15-25, showy; chaff of the receptacle entire, 
or sometimes 3-toothed. 

On prairies and barrens, Pennsylvania to Minnesota. Re- 
corded as adventive in Massachusetts. Aug.—Sept. 


> 


25. Helianthus tomentosus Michx. Woolly 
Sunflower. Fig. 4485. 

AH. tomentosus Michx. Fl. Bor. Am. 2: 141. 1803. 

Perennial; stem stout, hirsute or hispid, especially 
above, branched, 4°-10° high. Leaves rather thin, 
ovate, or the lower oblong, mostly alternate, 3-ribbed 
above the base, gradually or abruptly contracted into 
margined petioles, rough above, softly villous-pubes- 
cent beneath, sparingly serrate, the lower often 1° 
long and 4’ wide; heads commonly several or nu- 
merous, 3-4’ broad; involucre hemispheric, its bracts 
imbricated in many series, linear-lanceolate, long- 
acuminate, squarrose, densely hirsute and ciliate, 
usually longer than the broad yellowish disk; chaff 
of the receptacle and lobes of the disk corolla pubes- 
cent; pappus of 2 subulate awns. 

In dry soil, Virginia to Georgia and Alabama. Re- 
ported from Illinois, probably erroneously. Aug.—Oct. 


26. Helianthus tuberosus L. Jerusalem Arti- 
choke. Earth Apple. Fig. 4486. 


Helianthus tuberosus L. Sp. Pl. 905. 1753. 
Helianthus tuberosus subcanescens A. Gray, Syn. FI. 1: 
Part 2, 280. 1884. 


Perennial by fleshy thickened rootstocks, bearing 
tubers; stems hirsute or pubescent, branched above, 
6°-12° high. Leaves ovate or ovate-oblong, rarely 
ovate-lanceolate, firm, 3-nerved near the base, nar- 
rowed, or the lower rounded, truncate or subcordate 
at the base, acuminate at the apex, rather long- 
petioled, scabrous above, finely pubescent or canes- 
cent beneath, serrate, 4-8’ long, 13’-3’ wide, the up- 
per alternate, the lower opposite; heafs several or 
numerous, 2’-33’ broad; involucre hemispheric, its 
bracts lanceolate, acuminate, hirsute or ciliate, squar- 
rose; disk yellow; rays 12-20; chaff of the receptacle 
acute and pubescent at the summit; achenes pubescent. 


In moist soil, Nova Scotia and Ontario to Manitoba, 
Georgia and Arkansas and Kansas. Often occurs along 
roadsides in the east, a relic of cultivation by the abo- 
rigines. Now extensively grown for its edible tubers. 
Canada potato. Girasole. Topinambour. Sept.—Oct. 


mS - 
67. RIDAN Adans. Fam. Pl. 2: 130. 1763. 
[ActrinoMERIsS Nutt. Gen. 2: 181. 1818.] 
Rough-pubescent, rather coarse, herbs with alternate or opposite, simple, more or less 
toothed leaves, usually decurrent on the stem and branches, and corymbose, radiate or discoid 


GENUS 67. THISTLE FAMILY. 487 


heads. Involucre small, flattish, of few spreading or recurved narrow bracts. Receptacle 
convex or conic, becoming globose, chaffy. Ray-flowers neutral, yellow or white, sometimes 
wanting. Disk-flowers perfect, fertile, embraced by the chaff. Anthers entire at the base. 
Style-branches of the disk with acute appendages. Achenes flattened, more or less winged, 
spreading in all directions on the globose receptacle. Pappus of 2 or 3 finally spreading 
smooth awns, sometimes with 2-3 smaller awns or scales. [Name unexplained.] 


Two species, of eastern North America, the following 
typical. 


1. Ridan alternifolius (L.) Britton. Wing- 
stem. Yellowlron weed. Fig. 4487. 
Coreopsis alternifolia L. Sp. Pl. 909. 1753. 
Aciinomeris squarrosa Nutt. Gen. 2: 181. 1818. 
Actinomeris alternifolia DC. Prodr. 5: 575. 1836. 
Verbesina alternifolia Britton; Kearney, Bull. Torr. Club 

20: 485. 1893. 

Perennial; stem puberulent or glabrous, narrowly 
winged, or wingless, simple, or branched near the 
summit, leafy, 4°-9° high. Leaves oblong-lanceolate 
or lanceolate, acuminate, serrate, serrulate or entire, 
rough or roughish on both sides, gradually narrowed 
to the sessile base, or short-petioled, 4-12’ long, 
wide, alternate, or the lower opposite and 
slender-petioled; heads numerous, 17-2’ broad, co- 
rymbose-paniculate; rays 2-10, yellow; involucre of 
few lanceolate, at length deflexed bracts; disk glo- 
bose, yellow; achenes broadly winged or nearly wing- 
less, sparingly pubescent; pappus 2 divergent awns. 

In rich soil, New Jersey to Iowa, Kansas, Florida and 
Louisiana. Winged ironweed. Aug.—Sept. 


68. PHAETHUSA Gaertn. Fr. & Sem. 2: 425. 1701. 


Perennial, pubescent or scabrous herbs (some tropical species shrubby), with alternate 
or opposite dentate leaves often decurrent on the stem and branches, and corymbose or soli- 
tary heads of both tubular and radiate yellow or white flowers, or the rays sometimes want- 
ing. Involucre campanulate or hemispheric, its bracts imbricated in few series. Receptacle 
convex or conic, chaffy, the chaff embracing the disk-flowers. Ray-flowers pistillate or neu- 
tral. Disk-flowers perfect, mostly fertile, their corollas with an expanded 5-lobed limb, 
usually longer than the tube. Style-branches of the disk-flowers with acute papillose append- 
ages. Achenes flattened, or those of the ray-flowers 3-sided. Pappus of 2 (1-3) subulate 
awns, sometimes with 2 or 3 intermediate scales. [The daughter of Helios.] 

A large genus, mainly natives of the New World. Besides the following, about 6 others occur 
in the southern and southwestern United States. Type species: Phaethusa americana Gaertn. 
Involucre campanulate, 2”—3” broad ; heads small, numerous. 


Leaves alternate ; rays white. 1. P. virginica. 
Leaves opposite ; rays yellow. 2. P. occidentalis. 
Invoiucre hemispheric, 7”—12” broad; heads few, large. 3. P. helianthoides. 


1. Phaethusa virginica (L.) Britton. Small 
White or Virginia Crownbeard. Fig. 4488. 


Verbesina virginica L. Sp. Pl. 901. 1753. 


Perennial; stem densely puberulent, terete or winged, 
simple or branched, 3°-6° high. Leaves usually thin, 
alternate, ovate, roughish above, puberulent, canes- 
cent or glabrate beneath, acute or acuminate at the 
apex, 4-10’ long, 1’-3’ wide, contracted at the base 
into winged petioles, the uppermost sessile, lanceo- 
late, smaller, often entire; heads corymbose-panicu- 
late at the ends of the stem and branches, numerous, 
‘6-10 broad; involucre oblong-campanulate, 2’-3” 
broad, its bracts narrowly lanceolate, erect, obtuse, 
pubescent; rays 3-5, obovate, white, pistillate; achenes 
minutely pubescent, winged or wingless; pappus of 
2 slender awns, or sometimes none. 


In dry soil, Pennsylvania to Virginia, Illinois, Mis- 
souri, Kansas, Florida and Texas. Aug.—Sept. 


488 COMPOSITAE. Vor. III. 


‘ 


2. Phaethusa occidentalis (L.) Britton. Small Yellow Crownbeard. Fig. 4489. 


Siegesbeckia occidentalis L. Sp. Pl. 900. 1753. 
Verbesina occidentalis Walt. Fl. Car. 213. 1788. 
Phaethusa americana Gaertn. Fr. & Sem. 2: 425. 1791. 
V. Siegesbeckia Michx. Fl. Bor. Am. 2: 134. 1803. 


Perennial; stem glabrous, or puberulent above, 
usually much branched, narrowly 4-winged, 3°-7° 
high, the branches also winged and pubescent. 
Leaves thin, ovate, or the upper oblong, opposite, 
minutely rough-pubescent on both sides, or gla- © 
brate, acuminate at the apex, narrowed or con- 
tracted below into slender margined or naked 
petiole, serrate, 4’-10’ long, 1’-33’ wide; heads 
numerous, 6-12” broad, corymbose at the ends 
of the stem and branches; involucre oblong-cam- 
panulate, 2”-3” broad, its bracts lanceolate, ob- 
tuse, erect, or the tips slightly spreading, pubes- 
cent; rays 1-5, yellow, usually pistillate, rarely 
none; achenes wingless; pappus of 2 slender, at 
length divergent awns. 


In dry thickets and on hillsides, Maryland and southern Pennsylvania to Illinois, Florida, 
Alabama and Texas. Aug.—Oct. 


3. Phaethusa helianthoides ( Michx.) Britton. 
Sunflower Crownbeard. Fig. 4490. 


entesie helian‘hoides Michx. Fl. Bor. Am. 2: 135. 
Bernconane helianthoides Nutt. Gen. 2: 181. 1818. 

Perennial; stem hispid or hirsute, 4-winged, usu- 
ally simple, 2°-4° high. Leaves ovate or oval, ses- 
sile, acute, acuminate or obtuse at the apex, nar- 
rowed at the base, serrate or serrulate, rough or 
appressed-hispid above, densely pubescent or canes- 
cent beneath, 2-4’ long, 1’-13’ wide, all alternate, 
or the lower opposite; heads solitary or few, 2’-3’ 
broad; involucre hemispheric, about 2’ high, its 
bracts lanceolate, acutish, canescent, appressed; rays 
8-15, pistillate or neutral, linear-oblong, yellow; 
achenes scabrous or pubescent, broadly winged; 
pappus of 2 subulate awns. 


On dry prairies and in thickets, Ohio to Georgia, west 
to Iowa, Missouri and Texas. June—July. 


69. XIMENESIA Cav. Icones 2: 60. 1703. 


Pubescent caulescent herbs, with alternate or sometimes opposite, simple, toothed or 
somewhat laciniate leaves, and solitary or few, radiate, showy, peduncled heads. Involucre 
rather flat, of narrow spreading bracts. Receptacle convex, chaffy. Ray-flowers pistillate, 
fertile, numerous, the rays yellow. Disk-flowers numerous, perfect, fertile. Anthers some- 
what sagittate at the base. Style-branches with slender pubescent appendages. Achenes 
flat, winged. Pappus of short and straight awns. [In honor of Joseph Ximenes, a Spanish 
apothecary and botanist. ] 


About 4 species, natives of America, the following typical. 


GENUS 69. THISTLE FAMILY. 489 


1. Ximenesia encelioides Cav. Golden Crownbeard. Fig. 4491. 


Ximenesia encelioides Cav. Icon. 2: 60. pl. 178. 1793. 
Verbesina encelioides A. Gray, Syn. Fl. 1: Part 2, 288. 
1884. 


Annual; stem densely puberulent, much branched, 
1°-2° high. Leaves deltoid-ovate or deltoid-lanceo- 
late, thin, 2’-4’ long, acuminate, acute or blunt at the, 
apex, coarsely dentate, or even laciniate, green and \\ 
minutely pubescent above, pale and densely canescent 
beneath, all alternate, or the lowest opposite, nar- 
rowed at the base into naked or wing-margined peti- 
oles, which are often provided with dilated append- 
ages at the base; heads several or numerous, 1I’-2’ 
broad; involucre hemispheric, about 2’ high, its bracts 
lanceolate, canescent; rays 12-15, bright golden yel- 
low, 3-toothed; achenes of the disk-like flowers obo- 
vate, winged, pubescent, their pappus of 2 subulate 
awns, those of the ray-flowers rugose, thickened, 
often wingless. 


In moist soil, Kansas to Texas, Arizona and Mexico, 
and occasional in waste grounds farther east. Also in 
Florida and widely distributed in warm regions as a 
weed. Summer. 


70. COREOPSIS L. Sp: Pl007:- 1753; 


Anual or perennial, mostly erect herbs, with opposite leaves, or the upper alternate, and 
large long-peduncled heads of both tubular and radiate flowers, the rays yellow, or brown 
at the base, or brown throughout, or pink. Involucre usually hemispheric, its bracts in 2 
distinct series, all united at the base, those of the outer series commonly narrower and shorter 
than the inner. Receptacle flat or slightly convex, chaffy, the chaff flat or concave. Ray- 
flowers neutral. Disk-flowers perfect, fertile, their corollas with slender tube and broader 
5-toothed limb. Anthers mostly entire at the base. Style-tips truncate or subulate. Achenes 
flat, orbicular to oblong, winged or wingless. Pappus of 2 short teeth, or a mere coroniform 
border, or none. [Greek, bug-like, referring-to the achenes.] 


About 55 species, natives of America, South Africa and Australia, known as Tickseed. In 
addition to the following, some 20 others occur in the southern and western United States. Type 
species: Coreopsis lanceolata L. 


Ts ae Urs ree or cuspidate; outer involucral bracts about as long as the inner (except in 
. tripteris). 
Leaves simple, or pinnately lobed; achenes often with a callus on the incurved inner side. 
Achenes with thin flat broad wings. 
Leaves mostly near the base of the stem, the heads long-peduncled. 
Glabrous, or sparingly hairy below. 1. C. lanceolata. 
Pubescent or hirsute. 2. C. crassifolia. 
Stem leafy to near the shorter-peduncled heads. 
Pubescent or hirsute; leaves entire or with a few lateral lobes. 3. C. pubescens. 
Glabrous or very nearly so; leaves 1-2-pinnately parted into narrow segments. 
4. C. grandiflora. 
Achenes with thick, involute narrow wings. 5. C. auricula’a. 


Leaves palmately 3-cleft or divided; achenes without callus. 
Leaves petioled, 3-divided into lanceolate segments, or the upper entire. 6. C. tripteris. 
Leaves sessile. 


Leaves rigid, deeply 3-lobed above the base. 7. C. palmata. 
Leaves divided to the base. 
Leaf-segments entire; disk-flowers yellow. 8. C. major. 
Leaf-segments 1—2-parted. 
Disk-flowers yellow; leaf-segments linear-filiform. 9. C. verticillata. 
Disk-flowers purple-brown ; leaf-segments 1”—3” wide. 10. C. delphintfolia. 


2. Style-tips truncate or obtuse; outer involucral bracts much shorter than the inner. [Genus 
CALLiopsis Reichb. ] 


Achenes broadly winged ; annual. 11. C. cardaminefolia. 
Achenes wingless. : : 
Rays yellow with brownish bases ; annual with pinnately divided leaves. 12. C. tinctoria. 


Rays rose-pink, rarely white; perennial with linear entire leaves. 13. C. rosea. 


490 COMPOSITAE. Vot. IIT. 


1. Coreopsis lanceolata L. Lance-leaved 
Tickseed. Fig. 4492. 


Coreopsis lanceolata L. Sp. Pl. 908. 1753. 
Coreopsis lanceolata var. angustifolia T. & G. Fl. N. 
An 2 a BAAnenOAG: 


Perennial; stem slender, glabrous, or sparingly 
hispid near the base, 1°-2° high. Leaves glabrous, 
the lower and basal ones slender-petioled, spatu- 
late or narrowly oblong, very obtuse, 2’-6’ long, 
entire, or with 1-3 lateral obtuse entire lobes; 
stem leaves few, sessile or nearly so, lanceolate 
or oblong, obtuse or acutish, usually quite entire; 
heads few or solitary on elongated slender pedun- 
cles, 13-23’ broad, showy; peduncles often 12’ 
long; involucre depressed-hemispheric, its bracts 
glabrous or ciliate, lanceolate or ovate-lanceolate, 
the outer narrower than the inner, but nearly as 
long; rays 6-10, bright yellow, cuneate, 3-7-lobed; 
achenes oblong, broadly winged, about 1” long; 
pappus of 2 short teeth. 

In dry or moist soil, Ontario to Virginia, Michigan, 


Illinois, Florida, Louisiana and Missouri. Escaped 
from cultivation eastward. May—Aug. 


2. Coreopsis crassifolia Ait. Thick-leaved 
or Hairy Tickseed. Fig. 4493. 


Coreopsis crassifolia Ait. Hort. Kew. 3: 253. 1789. 
Coreopsis lanceolata var. villosa Michx. Fl: Bor. Am. 
2a Fete TOO Zs 


Perennial; stem hirsute or villous-pubescent 
below, ascending, little branched, 8-20’ high. 
Leaves rather thick, hirsute or pubescent, all 
entire, the lower and basal ones petioled, oblong 
to obovate-spatulate, mostly very obtuse, 1’-3’ 
long, 4’-10” wide; stem leaves few, short-peti- 
oled or sessile, obtuse or acutish, narrower; heads 
few, 1-2’ broad, borne on slender, puberulent pe- 
duncles often I° long, similar to those of C. lan- 
ceolata, as are the rays, involucral bracts and 
achenes. 


Dry soil, Illinois and Missouri to Kansas, Louisi- 
ana, east to South Carolina and Florida. May—Aug. 


<— 
Ts 


Wi—J =z. = 3. Coreopsis pubéscens Ell. Star Tick- 
seed. Fig. 4494. 
C. pubescens Ell. Bot. S. C. & Ga. 2: 441. 1824, 


Perennial; stem pubescent or puberulent, leafy, 
little branched or simple, erect, 2°-4° high. Leaves 
firm in texture, pubescent or glabrous, entire, or 
, some of them 3-5-lobed or divided, the basal and 
lower ones slender-petioled, obovate-oval, obtuse, 
the upper short-petioled or sessile, broadly lan- 
ceolate or oblong, acute or acutish, 2-3’ long; 
heads few, I-13’ broad, long-peduncled; invo- 
lucre depressed-hemispheric, glabrous, star-like, 
its lanceolate obtuse or acute outer bracts nearly 
as long as but much narrower than the ovate 
inner ones; rays 8-10, yellow, cuneate, lobed at 
the apex; achenes similar to those of the preced- 
ing species or broader. 


In dry woods, Virginia to Illinois and Missouri, 
south to Florida and Louisiana. June-Aug. 


GENUS 70. THISTLE FAMILY. 491 


4. Coreopsis grandiflora Hogg. Large- 
flowered Tickseed. Fig. 4495. 


Coreopsis grandifora Hogg; Sweet, Brit. Fl. Gard. 
2: pl. 175. 1825-27. 


Perennial; stem glabrous, usually branched 
above, 1°-3° high. Leaves, or most of them, 
I-2-pinnately parted, or the lower entire and 
slender-petioled; segments of the lower stem 
leaves oblong, obtuse, the terminal one larger 
than the lateral; segments of most of the stem 
leaves linear or even filiform, petioles ciliate; 
heads commonly several, 1’—2’ broad, long-pedun- 
cled; outer bracts of the involucre lanceolate, 
narrower and mostly somewhat shorter than the 
oval or ovate-lanceolate inner ones; rays 6-10, 
yellow, lobed; achenes oblong, broadly winged 
when mature, the projections on the inner face 
usually large; pappus of 2 short scales. 


In moist soil, Kansas and Missouri to Texas, east 
to Georgia. May—Aug. 


5. Coreopsis auriculata L. Running or 
Lobed Tickseed. Fig. 4496. 


Coreopsis auriculata L. Sp. Pl. 908. 1753. 


Perennial; stoloniferous; stems weak, very slen- 
der, decumbent or ascending, 6-15’ long, little 
branched, or simple, pubescent or hirsute, at least 
below, mostly glabrous above. Leaves thin, en- 
tire or pinnately 3-5-lobed or 3-5-divided, the 
terminal segment entire and much larger than the 
lateral ones, the lower and basal more or less pu- 
bescent, broadly oblong or nearly orbicular, with 
slender pubescent petioles; stem leaves few, peti- 
oled or sessile, mostly obtuse; heads 1-13’ broad, 
slender-peduncled; outer bracts of the involucre 
oblong to lanceolate, narrower than the mostly 
ovate and acute inner ones; rays 6-I0, cuneate, 
about 4-toothed, yellow; achenes oval, narrowly 
winged, the wings involute and thick. 


In woods, Virginia to Illinois, Kentucky, Florida 
and Louisiana. May—Aug. 


| 
a 


%, A | 
mX: 4] 
\\ j 


6. Coreopsis tripteris L. Tall Tickseed. 
Fig. 4497. 
Coreopsis tripteris L. Sp. Pl. 908 1753. 


Perennial; stem glabrous, much branched above, 
4°-8° high. Leaves petioled, glabrous, or very 
nearly so, firm, the lower all divided into lanceo- 
late, entire, acute, rough-margined segments, which 
are 2-5’ long, }+’-1’ wide and pinnately veined; 
uppermost leaves lanceolate, entire; heads numer- 
ous, slender-peduncled, 17-13’ broad; outer bracts 
of the involucre linear, obtusish, much narrower 
and somewhat shorter than the ovate-oval or ovate- 
lanceolate, acute inner ones; rays 6-10, yellow, ob- 
tuse, entire; achenes oblong to obovate, narrowly 
winged, emarginate; pappus none. 


In moist woods and thickets, southern Ontario to 
Pennsylvania, Wisconsin, Arkansas, Virginia, Florida 
and Louisiana. July—Oct. 


492 _ COMPOSITAE. Vo. III. 


heads 


Texas. 


Coreopsis major Walt. Fl. Car. 214. 1788. 
Coreopsis senifolia Michx. Fl. Bor. Am. 2: 138. 1803. 
Coreopsis Oemleri Ell. Bot. S. C. & Ga. 2: 435. 1824. 


Coreopsis stella a Nutt. Journ. Acad. Phil.7: 76. 1834. 


Coreopsis senifolia var. stellata T. & G. Fl. N. A. 2: 

342. 1842. 

Perennial; stem pubescent or glabrous, branch- 
ed above, 2°-3° high. Leaves sessile, more or 
less pubescent, or glabrous, divided to the base 
into 3, lanceolate, linear, ovate-lanceolate or ob- 
long, acute, entire segments 2’-4’ long, 2’-12” 
wide, which appear as if in verticils of 6; upper 
and lower leaves (rarely all of them), undivided 
and entire; heads several or numerous, slender- 
peduncled, 1-2’ broad; bracts of the hemispheric 
involucre all united at the base, the outer ones 
linear-oblong, obtuse, equalling or shorter than 
the broader inner ones, all pubescent; rays 6-10, 
yellow, oblong, entire; disk yellow; achenes ob- 
long to elliptic, winged, 2’-2” long; pappus of 2 
short deciduous teeth. 

In dry sandy woods, Virginia to Kentucky, Florida 
and Alabama. July—Aug. Consists of several races, 
differing in pubescence and in shape and width of the 


leaf-segments. 
AN wh 
NG: 


we 
wy 


teeth. 


7. Coreopsis palmata Nutt. Stiff Tick- 


seed. Fig. 4408. 


Coreopsis palmata Nutt. Gen. 2: 180. 1818. 


Perennial; stems rigid, glabrous, simple, or 
little branched, very leafy, 1°-3° high. Leaves 
sessile, 2’-3’ long, palmately deeply 3-lobed at or 
below the middle, or the uppermost entire, thick, 
rigid, the lobes linear-oblong, obtusish, entire, 
or with 1-3 lateral lobes, their margins rough; 
few or solitary, short-peduncled, 17-2’ 
broad; involucre hemispheric, its bracts some- 
what united at the base, those of the outer series 
narrower and nearly as long as the inner ones; 
rays 6-10, bright yellow, oblong or obovate, most- 
ly 3-toothed; achenes oblong, narrowly winged, 
slightly incurved; pappus of 2 short tips, or none. 

On dry prairies and in thickets, Indiana to Mis- 
souri, Louisiana, Minnesota, Manitoba, Nebraska and 
June-July. 


8. Coreopsis major Walt. Wood or Greater 
Tickseed. Fig. 4499. 


all 
2 


AS g. Coreopsis verticillata L. Whorled 


Tickseed. Fig. 4500. 


J Coreopsis verticillata L. Sp. Pl. 907. 1753. 


Perennial; stem stiff, much branched, slender, 
leafy, 1°-2° high. Leaves sessile, glabrous, 2-3- 
ternately dissected into linear-filiform entire seg- 
ments; heads numerous, 1-12’ broad; involucre 
hemispheric, or short-cylindric in fruit, glabrous, 
its outer bracts linear, obtuse, commonly some- 
what shorter and much narrower than the ovate- 
oblong inner ones; rays 6-10, yellow, spatulate- 
oblong, obtuse; disk dull yellow; achenes oblong, 
narrowly winged, 2” long; pappus of 2 short 


In dry soil, Maryland to South Carolina, Kentucky, | 
Nebraska and Arkansas. Apparently erroneously re- 
corded from farther north. June—Sept. 


GENUS 70. THISTLE FAMILY. 493 


10. Coreopsis delphinifolia Lam. Lark- 
spur Tickseed. Fig. 4501. 


Coreopsis delphinifolia Lam. Encycl. 2: 108. 1786. 


Perennial ; stem glabrous, branched above, rather 
slender, 1°-3° high. Leaves sessile, 1-2-ternately 
partly into linear or linear-lanceolate segments, 
which are 17-2’ long, 1-3” wide; heads several 
or numerous, 12-2’ broad; involucre hemispheric, 
its bracts glabrous; the outer linear-oblong, ob- 
tuse, shorter than or equalling the ovate-oblong 
inner ones; rays 6-10, yellow, entire; disk brown; 
achenes oblong to oval, narrowed at the base, 
narrowly winged; pappus of 2 short teeth. 


In dry woods, Virginia (according to Torrey and 
Gray), North Carolina to Georgia and Alabama. 
Aug.—Sept. 


11. Coreopsis cardaminefolia (DC.) T. & G. 
Cress-leaved Tickseed. Fig. 4502. 


Calliopsis cardaminefolia DC. Prodr. 5: 568. 1836. 
C. cardaminefolia T. & G. Fl. N. A. 2: 346. 1842. 


Annual; stem glabrous, branched, 1°-23° high. 
Basal leaves petioled, 24’ long, 1-2-pinnately parted 
into oblong or oval obtuse segments, the petioles 
sometimes slightly ciliate; stem leaves distant, ses- 
sile, or nearly so, pinnately parted into linear seg- 
ments, or the uppermost entire; heads 8-12” broad, 
slender-peduncled; involucre hemispheric, its inner 
bracts brown, ovate or ovate-lanceolate, obtuse or 
obtusish, scarious-margined, much longer than the 
lanceolate obtusish outer ones; rays 4-8, yellow with 
a brown base, 3-toothed; achenes oval, 1-14” long, 
winged, smooth, or slightly papillose; pappus of 2 
minute awns, or none. 


In moist soil, Kansas to New Mexico, Louisiana and 
northern Mexico. May-—Oct. 


Coreopsis Atkinsoniana Dougl., a northwestern 
species, with linear leaf-segments and very narrowly 
winged achenes, ranges eastward into North Dakota. 


12. Coreopsis tinctoria Nutt. Golden Co- 


F Sy = anf : \ WBE 
reopsis. Garden Tickseed. Fig. 4503. ANY 4 


C. tinctoria Nutt. Journ. Acad. Phila. 2: 114. 1821. 


Annual; stem glabrous, branched, 1°-34° high. 
Leaves I-2-pinnately divided into linear, obtusish, 
mostly entire segments, or the uppermost linear and 
entire, the lower petioled; heads slender-peduncled, 
10-12” broad, or in cultivation much broader; invo- 
lucre hemispheric, its inner bracts brown, ovate or 
oblong, obtuse or acute, scarious-margined, 3-5 times 
as long as the obtuse outer ones; rays 6-10, cuneate, 
yellow with a brown base or brown all over; achenes 
linear or linear-oblong, about 1” long, wingless; pap- 
pus a mere border, or none. 


In moist soil, Minnesota to Alberta, Nebraska, Louisi- 
ana and Arizona. Escaped from gardens to roadsides 
and waste places eastward. Wild flax. Nuttall’s-weed. 
May-Sept. 


494 COMPOSITAE. Vo. III. 


13. Coreopsis rosea Nutt. Small Rose or Pink Tickseed. Fig. 4504. 
Coreopsis rosea Nutt. Gen. 2: 179. 1818. 


Perennial by slender rootstocks; stems at length 
much branched, slender, strict, glabrous, 6-24’ high. 
Leaves opposite, linear, glabrous, entire, obscurely I- 
nerved, 1’-22’ long, 2”-1” wide, sessile, or the lower 
petioled; heads slender-peduncled, several or numer- 
ous, 6’-12” broad; disk yellow; rays 4-8, pink or rose- 
colored (occasionally white), oblong to obovate, slight- 
ly 3-toothed or sometimes entire; inner bracts of the 
hemispheric involucre ovate-oblong, acutish or obtuse, 
glabrous, much longer than the lanceolate outer ones; 
achenes oblong or linear-oblong, thin, not winged, 
nearly straight, slightly ribbed on the inner face; pap- 
pus a very short truncate crown. 


In open swamps, eastern Massachusetts to Georgia, near 
the coast. July—Aug. 


71. BIDENS L. Sp. Pl. 831. 1753. 


Annual or perenniai herbs, with opposite serrate lobed divided or dissected leaves, or the 
uppermost alternate, and mostly large heads of both tubular and radiate flowers, or the rays 
none, or rudimentary. Involucre campanulate or hemispheric, its bracts in 2 series, distinct, 
or slightly united at the base; the outer often foliaceous and much larger than the inner. 
Receptacle flat or nearly so, chaffy, the chaff subtending the disk-flowers. Rays, when present, 
neutral, mostly entire, yellow in our species. Disk-flowers perfect, fertile, their corollas tubu- 
lar, 5-toothed. Anthers entire, or minutely sagittate at the base. Style-branches with short 
or subulate tips. Achenes flat, or quadrangular, cuneate, oblong or linear, the outer ones 
often shorter than the inner. Pappus of 2-6 teeth or subulate awns, upwardly or down- 
wardly barbed or hispid. [Latin, two-toothed, referring to the achenes.] 


About 75 species of wide geographic distribution. Besides the following, about 12 others occur 
in the southern and southwestern United States. Type species: Bidens tripartita L. 


Leaves lanceolate, serrate, undivided, rarely 3—5-lobed or incised. 
Rays present, large and conspicuous. 


Rays large, longer than the involucral bracts. 1. B. laevis. 

Rays short, rarely as long as the involucral bracts. ’ 2. B.cernua. 
Rays rudimentary, or none. 

Heads nodding after flowering. 2. B.cernua. 


Heads persistently erect. 
Pappus awns downwardly barbed. 
Involucral bracts not foliaceous; stem purple; flowers orange. 3. B. connata. 
Involucral bracts foliaceous; stem straw-color; flowers greenish yellow. 
4. B.comosa. 
Pappus awns upwardly barbed; involucre narrow. 5. B. bidentoides. 
Leaves, some or all of them, pinnately 1-3-parted or dissected. 
Rays rudimentary, or none, or very short. 
Achenes flat; leaves, some or all of them, 1—3-divided. 
Outer involucral bracts 4-8; achenes black or nearly black. 


Leaves membranous; heads 2”—3” high; awns short. 6. B. discoidea. 
Leaves not membranous; heads 5”—7” high; awns long. 7. B. frondosa. 
Outer involucral bracts 10-16; achenes brown. 8. B. vulgata. 
Achenes linear ; leaves dissected. g. B. bipinnata. 
Rays large and conspicuous. 
Achenes sparingly pubescent, not ciliate; pappus of 2 short teeth. 10. B. coronata. 
Achenes ciliate ; pappus 2—4 subulate teeth or awns. 
Achenes cuneate, or linear-cuneate. 11. B. trichosperma. 
Achenes obovate, very flat. 
Bracts of the involucre glabrous, or ciliate, short. 12. B. aristosa. 
Outer bracts densely hispid, much longer than the inner. 13. B.involucrata. 


GENUS 7I. THISTLE FAMILY. 495 


1. Bidens laévis (L.) B.S.P. Larger or Smooth Bur-Marigold. Brook Sun- 
flower. Fig. 4505. 


Helianthus laevis L. Sp. Pl. 906. 1753. 

Bidens chrysanthemoides Michx. Fl. Bor. Am. 2: 
136. 1803. 

Bidens laevis B.S.P. Prel. Cat. N. Y. 29. 1888. 

Bidens lugens Greene, Pittonia 4: 254. 1901. 


Annual; glabrous throughout; stems branch- 
ed, erect or ascending, 1°-3° high. Leaves 
sessile, lanceolate, evenly serrate or serrulate, 
acuminate at the apex, narrowed to the some- 
times connate-perfoliate base, 3-8’ long, 2’-1’ 
wide; heads numerous, short-peduncled, erect 
in flower, often declined in fruit, 1-23’ broad; 
rays very showy, golden yellow; involucre 
hemispheric, its outer bracts linear-oblong or 
spatulate, equalling or exceeding the broader, 
ovate or oblong, membranous inner ones, 
shorter than the rays; rays 8-10, obovate- 
oblong, obtuse; achenes cuneate, truncate, 2’— 
4” long, retrorsely hispid on the margins; 
pappus of 2-4 (usually 2), rigid downwardly 
barbed awns, shorter than the achene. 


In swamps and wet meadows, Massachusetts to Illinois, Kansas, Florida, Louisiana and Mexico. 
Northern records of this species apply to Bidens cernua. Aug —Nov. 


Bidens élegans Greene, of southeastern Virginia, appears to be a narrow-rayed race of this 
species. 


2. Bidens cérnua L.. Smaller or Nodding Bur-Marigold. Fig. 4506. 


Bidens cernua L. Sp. Pl. 832. 1753. 
Coreopsis Bidens L. loc. cit. 908. 1753. 

Annual; stems glabrous, or hispid, usually erect, 
branched, 3’-3° high. Leaves sessile and commonly 
somewhat connate-perfoliate at the base, lanceolate 
to oblong-lanceolate, usually coarsely and sharply 
serrate, glabrous, acuminate, 3-6’ long, 3’-1’ wide; 
heads numerous, globose, short-peduncled, 4’-1' 
broad, nodding after or during flowering; rays 
6-10, short (3”-6”), or none; involucre depressed- 
hemispheric, its outer bracts commonly ciliate, 
often large, foliaceous and much exceeding the 
broad, yellowish-margined membranous inner ones + 
achenes cuneate, 2” long, retrorsely hispid on the 
margins; pappus of 2-4 (usually 4), downwardly 
barbed awns, about half as long as the achene. 

In wet soil, Nova Scotia to Hudson Bay and British 
Columbia, North Carolina, Missouri and California. 
Also in Europe and Asia. Consists of many races, 


differingin size. Water-agrimony. Double-tooth. Pitch- 
forks. July—Oct. 


Bidens Eatoni Fernald, known only from brackish soil, along the Merrimac River, Mass., has 
narrower heads and smaller achenes with awns either upwardly or downwardly barbed. 


A plant, with all the leaves pinnately divided, growing in the vicinity of Minneapolis, Minn., 
described as Bidens connata pinnata S. Wats., may be a hybrid with B. aristosa. 


496 COMPOSITAE Vor. III. 


3. Bidens connata Muhl. Purple-stemmed 
Swamp Beggar-ticks. Fig. 4507. 
B. connata Muhl.; Willd. Sp. Pl. 3: 1718. 1804. 


Annual; glabrous throughout; stem erect, usu- 
ally branched, 68° high, purple. Leaves peti- 
oled, lanceolate or oblong-lanceolate, sharply and 
coarsely serrate, thin, 2-5’ long, #/-1’ wide, apex 
acuminate, base tapering, the uppermost some- 
times sessile, nearly entire and acutish, the lower 
sometimes with a pair of basal lobes, decurrent 
on the petiole; heads several or numerous, pe- 
duncled, 4-13’ broad; involucre campanulate or 
hemispheric, the outer bracts somewhat exceed- 
ing the ovate-oblong, inner ones; rays none, or 
I-5 and inconspicuous; disk-flowers orange; 
achenes cuneate or obovate, hairy and tubercled, 
or nearly glabrous, keeled, or angled, 2’-—3” long, 
the margins with either erect or retrorse hairs, 
or both, the 2-4 pappus awns downwardly barbed, 
half as long as the achene. 


In swamps or moist soil, Rhode Island to Ontario, 
Minnesota, Delaware, Georgia, Kentucky, Missouri 
and Nebraska. Introduced into Europe. Cuckold. 
Harvest-lice. Pitchforks. Aug.—Oct. 


4. Bidens comosa (A. Gray) Wiegand. 
Leafy-bracted Tickseed. Fig. 4508 


. connata var. comosa A. Gray, Man. Ed. 5, 261. 1867. 
. comosa Wiegand, Bull. Torr. Club 24: 436. 1897. 

. riparia Greene, Pittonia 4: 261. 1901. 

. acuta (Wiegand) Britton, Man. 1001. 1901. 


Bowy 


Annual, glabrous; stem erect, branched, 6—43° 
high, straw-colored. Leaves short-petioled, or sub- 
sessile, lanceolate to elliptic-lanceolate, coarsely ser- 
rate with mostly smaller teeth than in B. connata, 
tapering to each end, the petioles broadly margined ; 
heads several or numerous, 2-13” broad; outer 
bracts of the involucre linear, spatulate or lanceo- 
late, foliaceous, erect or spreading, often toothed, 
2-4 times as long as the head; rays none; corollas 
mostly 4-lobed, pale greenish yellow; stamens and 
style included; achenes larger, 32’—53” long, evenly 
cuneate, very flat; pappus awns commonly 3, down- 
wardly barbed, somewhat shorter than the achene. 


In wet soil, Massachusetts to Illinois, North Dakota, 
New Jersey, West Virginia, Kentucky and Kansas. 
Leaves thicker and paler than in B. connata. Aug.—Oct. 


) 
! 


i 
N" i 


XN i \\ 
S\ 


5. Bidens bidentoides (Nutt.) Britton. 
Swamp Beggar-ticks. Fig. 4509. 
Diodonta bidentoides Nutt. Trans. Am. Phil. Soc. 

(II) 7: 367. 1841. 
Coreopsis bidentoides T. & G. Fl. N. A. 2: 339. 1842. 
B. bidentoides Britton, Bull. Torr. Club 20: 281. 1893. 


!) 


7 


Closely resembles B. connata, glabrous through- 
out; stem branched, 1°-4° high. Leaves similar, 
lanceolate, sharply serrate, petioled, or the upper 
sessile and entire, acuminate at the apex, nar- 
nowed at the base; involucre narrowly or be- 
coming somewhat broadly campanulate, its outer 
bracts linear, foliaceous, not ciliate, usually much 
exceeding the oblong inner ones; rays none, or 
rarely present and very short; achenes linear- 
cuneate, 3-5” long, their sides and the 2 slender 
pappus awns (rarely with 2 short intermediate 
awns) upwardly barbed or hispid. 

Muddy shores of the Delaware River and Bay in 


Pennsylvania, New Jersey, Delaware and Maryland. 
Aug.—Oct. 


GENUS 7I. THISTLE FAMILY. 


6. Bidens discoidea (T. & G.) Britton. 
Small Beggar-ticks. Fig. 4510. 


Coreopsis discoidea T. & G. Fl. N. A. 2: 339. 1842. 
B. discoidea Britton, Bull. Torr. Club 20: 281. 1893. 


Annual, glabrous, slender, branching, erect, 2’-6° 
high. Leaves membranous, very slender-petioled, 
all the lower ones divided into 3 lanceolate or 
oblong-lanceolate, dentate, acuminate segments 
which are 17-3’ long; uppermost leaves commonly 
rhombic-lanceolate and undivided; heads usually 
numerous, slender-peduncled, 2-4” broad and 
about as high; involucre broadly campanulate or 
hemispheric, its outer bracts mostly 4, usually 
foliaceous and obtuse, usually much surpassing 
the inner ones; rays apparently always wanting; 
achenes flat, narrowly cuneate, upwardly strigose, 
about 2” long; pappus of 2 short, upwardly his- 
pid, rarely downwardly barbed awns. 


In swamps and wet places, Massachusetts to Vir- 
ginia, Ohio, Michigan, Louisiana and Texas. July— 


LG 7. Bidens frondésa L. Beggar-ticks. 


Stick-tight. Fig. 4511. 


Z- 
— 
\ TS 


Bidens frondosa L. Sp. Pl. 832. 1753. 
B. melanocarpa Wiegand, Bull. Torr. Club 26: 4o5. 


= 
Y 


y 1899. 

y J | Annual; stem erect, branched, glabrous, or 
T| nearly so, often purplish, 2°-3° high. Leaves thin, 

\ Y GZ but not membranous, slender-petioled, pinnately 
C77 3-5-divided or the uppermost undivided, the seg- 


ments lanceolate or oblong-lanceolate, sharply 
serrate, acuminate at the apex, narrowed at the 
base, usually slightly pubescent beneath, stalked, 
2’-4’ long, 4-1’ wide; heads usually numerous, 
long-peduncled, about 6” high, 5’—10” broad; in- 
volucre campanulate, becoming hemispheric, its 
outer bracts 4-8, more or less foliaceous, often 
much exceeding the ovate-lanceolate, scarious- 
margined inner ones; rays none or rudimentary 
and inconspicuous; disk-corollas orange; achenes 
flat, narrowly cuneate, nearly black, 3’’-5’’ long, 
ciliate, the two slender awns downwardly barbed, 
or sometimes upwardly hispid. 


In moist soil, often a weed in fields, Nova Scotia to Florida, British Columbia, Texas, Colorado 
and California. Introduced as a weed into southern Europe. Rayless marigold. Beggar-lice. 
Devil’s-pitchfork. Stick-seed. Common _ bur-marigold. 

Old-ladies clothes-pins. Cuckles. July—Oct. 


AWN 


8. Bidens vulgata Greene. Tall Beggar-ticks. 
Fig. 4512. 
Bidens vulgata Greene, Pittonia 4: 72. 1 


: 899. 
B. eeTers puberula Wiegand, Bull. Torr. Club 26: 408. 
1899. 


Taller, sometimes 9° high, glabrous or nearly so, 
or crisp-pubescent above. Leaves pinnately 3-5- 
divided, the veins straight and prominent; heads 
larger, 7’-12” broad, stout-peduncled; outer invo- 
lucral bracts linear to linear-spatulate, ciliate, the 
inner mostly ovate or narrowly triangular, pubescent 
at apex; ray-flowers usually present, small, yellow; 
achenes very flat, 3-43” long, 2’—24” wide, brown 
or greenish brown, the margins downwardly barbed 
above, upwardly hairy below; awns 2, half as long 
as the achene or more, downwardly barbed. 


In moist soil, Quebec to British Columbia, New York, 
North Carolina, Missouri, Colorado and California. 
Aug.Sept. Included in the preceding species in our 
first edition. 


32 


498 COMPOSITAE. Vor. LI. 


g. Bidens bipinnata L. Spanish Needles. 
Cuckolds. Fig. 4513. 


Bidens bipinnata L. Sp. Pl. 832. 1753. 
Annual; stem quadrangular, erect, freely branch- 


ed, rather slender, 1°-5° high. Leaves thin, acumi- 
nate, petioled, 1-3-pinnately dissected into ovate or 
oblong, toothed or lobed segments, the lower often 
8’ long; heads usually numerous, long-peduncled, 
2’-4” broad; involucre narrow, its outer bracts 
linear, ascending, nearly as long as the broader erect 
inner ones; rays 3-4, yellow, short, sometimes none; 
achenes linear, 4-angled, slightly pubescent, narrowed 
upward into a beak, 5’-9” long, the outer ones com- 
monly shorter and thicker than the inner; pappus 
of 2-4 (usually 4), downwardly barbed, slightly un- 
equal, spreading awns, much shorter than the achene. 


In various situations, often a weed in cultivated fields, 
Rhode Island to Florida, Ohio, Nebraska, Kansas and 
Arizona. Introduced as a weed into southern Europe 
and Asia. July—Oct. 


to. Bidens coronata (L.) Fisch. Southern 
Tickseed-Sunflower. Fig. 4514. 


Coreopsis coronata L. Sp. Pl. Ed. 2, 1281. 1763. 
Coreopsis aurea Ait. Hort. Kew. 3: 252. 1780. 
Bidens coronata Fisch.; Steudel, Nom. Ed. 2, 202. 1840. 


Annual, glabrous or nearly so throughout; stem 
branched, 1°-3° high. Lower leaves petioled, 3’-5’ 
long, 3-divided, the terminal segment lanceolate, acute 
or acuminate, serrate, much larger than the serrate 
or entire lateral ones; upper leaves much smaller, 
3-parted, 3-lobed or undivided, sessile or short- 
petioled, entire or serrate; leaves rarely all undi- 
vided; heads numerous, slender-peduncled, 1-2’ 
broad; involucre hemispheric, its outer bracts linear- 
oblong, obtuse, equalling or slightly exceeding the 
broader inner ones; rays 6-10, obtuse; achenes 
broadly cuneate, slightly pubescent, 1-2” long; pap- 
pus of 2 chaffy blunt divergent somewhat laciniate 
teeth, rarely with 2 shorter intermediate ones. 


In wet places, Virginia to Florida and Alabama. July- 
Sept. 


11. Bidens trichospérma (Michx.) Britton. 
Tall Tickseed-Sunflower. Fig. 4545. 

C. trichosperma Michx. Fl. Bor. Am. 2: 139. 1803. 
B. trichosperma Britton, Bull. Torr. Club 20: 281. 1893. 
Coreopsis trichosperma var. tenuiloba A. Gray, Syn. Fl. 

i 2) Part 2,205.) 184: 
Bidens trichosperma tenuiloba Britton, Bull. Torr. Club 

20: 281. 1893. 

Annual or biennial, glabrous; stem tall, obscurely 
quadrangular, much branched, 2°-5° high. Lower 
leaves petioled, 4’-8’ long, pinnately divided into 5-7 
lanceolate or linear, acuminate, sharply serrate, in- 
cised or nearly entire, sessile or short-stalked seg- 
ments; upper leaves sessile or nearly so, 3-lobed, 
_3-divided, or entire and linear-lanceolate; heads nu- 
merous, 13-23’ broad, long-peduncled; involucre 
hemispheric, its outer bracts linear or spatulate, 
sometimes ciliate, about the length of the broader 
inner ones; rays 6-19, obtuse, golden yellow, 7-12” 
long; achenes oblong-cuneate or the inner ones nar- 
rower, hispid-pubescent and ciliate, 2-4” long; pap- 
pus of 2 short erect or divergent, upwardly hispid 
teeth or short awns. 

In swamps and wet meadows, Massachusetts to Georgia; Illinois to Michigan and Kentucky. 

Recorded from Ontario and Minnesota. Aug.—Oct. 


GENUS 7I. THISTLE FAMILY. 499 


12. Bidens arist6sa (Michx.) Britton. Western Tickseed-Sunflower. 
Fig. 4516, 


Coreopsis aristosa Michx. Fl. Bor. Am. 2: 140. 1803. 
C. aristata Muhl.; Willd. Sp. Pl. 3: 2253. 1804. 
B. aristosa Britton, Bull. Torr. Club 20: 281. 1893. 


Annual or biennial; stem much branched, 1°-3° 
high. Leaves thin, slender-petioled, pubescent 
beneath, the lower ones pinnately 5-7-divided, 
3-6’ long, the segments lanceolate, serrate, in- 
cised or pinnatifid, acuminate, narrowed at the 
base; upper leaves less divided, lanceolate, or 
merely lobed, sessile or short-petioled; heads nu- 
merous, slender-peduncled, 1-2’ broad; outer 
bracts of the hemispheric involucre 8-10, linear 
or spatulate, usually ciliate, not surpassing the 
inner; rays 6-9, obtuse; achenes very flat, oblan- 
ceolate or obovate, upwardly ciliate and strigose- 
pubescent; pappus of 2, rarely 4, slender upward- 
ly or downwardly barbed awns, sometimes nearly 
as long as the achene, rarely wanting. 


In swamps and wet prairies, Ohio to Minnesota, 
south to Louisiana and Missouri; southeastern Penn- 
sylvania and Delaware. -Aug.—Oct. 


13. Bidens involucrata (Nutt.) Britton. 
Long-bracted Tickseed-Sunflower. 


Fig. 4517. 


Coreopsis involucrata Nutt. Journ. Phil. Acad. 7: 74. 
1834. , 


Bidens involucrata Britton, Bull. Torr. Club 20: 281. 
1893. 


Similar to the two preceding species, minutely 
pubescent, 1°-3° high, much branched. Segments 
of the leaves narrower, linear-lanceolate, incised 
or pinnatifid, long-acuminate; heads numerous, 
1-2’ broad, on slender usually hispid peduncles; 
outer bracts of the hemispheric involucre 10-20, 
linear-lanceolate, acuminate, densely hispid and 
ciliate, much exceeding the inner ones; rays 
orange at the base; achenes flat, ciliate and stri- 
gose; pappus of 2 short teeth. 


In swamps, Illinois to Kansas, Arkansas and Texas; 
southeastern Pennsylvania and Delaware. July—Sept. 


72. MEGALODONTA Greene, Pittonia 43:270:., 1901. 


Perennial aquatic herbs, with opposite or whorled leaves, those of the submersed ones 
filiformly dissected, those of the emersed ones serrate to laciniate, and solitary peduncled 
showy heads with both tubular and radiate flowers. Involucre hemispheric, its bracts in 2 
series, rather broad, distinct or nearly so, the outer ones smaller than the inner, often lax. 
Receptacle chaffy. Rays neutral, yellow. Disk-flowers perfect, fertile, their corollas tubular, 
3-toothed. Anthers minutely sagittate at the base. Style-branches. with subulate tips. 
Achenes nearly terete, truncate at both ends. Pappus of 3-6 long-subulate awns which are 
retrorsely barbed, except near the base. [Greek, large-toothed, referring to the pappus awns.] 


Two known species, the following typical, the other occurring in Washington State. 


500 COMPOSITAE. Vou. III. 


1. Megalodonta Béckii (Torr.) Greene. 
Water Marigold. Fig. 4518. 


Bidens Beckii Torr.; Spreng. Neue Entdeck. 2: 
1350, . LO2zE- 


M. Beckii Greene, Pittonia 4: 271. 1901. 


M. nudata Greene, loc. cit. 1901. 


Stems simple, or little branched, 2°-8° long. 
Submersed leaves sessile, 1-2’ long, repeat- 
edly divided into numerous capillary segments ; 
emersed leaves few, sessile, opposite, or some- 
times in 3’s, lanceolate or oblong, acute, serrate 
or laciniate, 3-13’ long; heads solitary or few, 
short-peduncled, 1-13’ broad; involucre hemi- 
spheric, its bracts oval or oblong, obtusish, 
glabrous, the outer somewhat shorter than the 
inner; rays 6-10, obovate, or oblong, notched, 


SY | Lar golden yellow; achenes nearly terete, 5-7” 
YU, SS long; pappus of 3-6 slender awns, downwardly 
barbed above, smooth below, divergent, 6’—12” 
3 long. 
Ss) | In ponds and streams, Quebec to New Jersey, 


west to Manitoba and Missouri. Aug.—Sept. 


73. THELESPERMA Less. Linnaea 6: 511. 1831. 


Glabrous annual or perennial herbs, with opposite linear and undivided, or finely dis- 
sected leaves, and long-peduncled heads of both tubular and radiate flowers, or the rays 
wanting. Involucre hemispheric or campanulate, of 2 distinct series of bracts, the outer 
short, narrow and somewhat spreading, the inner united nearly to or beyond the middle into 
a cup, their tips scarious-margined. Receptacle flat, chaffy, the 2-nerved broad white scarious 
chaff subtending the disk-flowers and achenes. Ray-flowers, when present, neutral, the rays 
yellow, entire or toothed. Disk-flowers perfect, fertile, their corolla with a slender tube and 
5-toothed limb. Anthers obtuse and entire at the base. Style-tips acute. Achenes, some or 
all of them, papillose on the back, oblong or linear, slightly compressed or terete, wingless. 
Pappus of 2 retrorsely hispid awns or scales, or sometimes none. [Greek, nipple-seed.] 

About 10 species, natives of the south central United States, Mexico and southern South Amer 
ica. Type species: Theiesperma scabiousoides Less. 


Rays large; pappus-awns shorter than the width of the achene. 


Leaves not rigid, their segments filiform-linear ; annual or biennial. 1. T. trifidum. - 
Leaves rigid, their segments linear; perennial. : : _2. T. intermedium. 
Rays inconspicuous, or none; awns longer than the width of the achene; perennial. . 
3. T. gracile. 
1. Thelesperma trifidum (Poir.) Brit- 
ton. Fine-leaved Thelesperma. =) 
Fig. 4519. SS 


Coreopsis trifida Poir. in Lam. Encycl. Suppl. 2: 
2535, “LOL 
Thelesperma filifolium A. Gray, Kew. Journ. Bot. 


Lis 252.5 1640; 

Thelesperma trifidum Britton, Trans. N. Y. Acad. 

Sci. 9: 182. 1890. 

Annual or biennial; stem branched, 1°-3° 
high. Leaves numerous, not rigid, 13’-2’ long, 
bipinnately divided into filiform or linear- 
filiform segments; heads several or numerous, 
12-15” broad; outer bracts of the involucre 
about 8, subulate-linear, equalling or more 
than half as long as the inner, which are 
united not higher than the middle; rays 6-10, 
somewhat spatulate, 3-lobed; disk purple or 
brown; achenes linear-oblong, straight, or 
slightly curved, the outer ones strongly papil- 
lose; awns of the pappus not longer than the 
width of the summit of the achene. 

In dry soil, South Dakota, Missouri and Ne- 
braska to Colorado, Texas, New Mexico and north- 
ern Mexico. June—Aug. 


Sy 


GENUS 73. THISTLE FAMILY. 501 


2. Thelesperma intermédium Rydb. Zn WY Wy U), 
Stiff Thelesperma. Fig. 4520. WAVZAN\G By Ry 
WIZ = y= Wig Ne 
Thelesperma intermedium Rydb. Bull. Torr. Club Lif} Y 


27: 631. 1900. 


Perennial from a deep woody root and slen- 
der rootstocks; stem rigid, usually much 
branched, 1°-14° high. Leaves usually numer- 
ous, 13-2’ long, bipinnately divided into entire, 
rigid, linear segments, but less compound than 
those of the preceding species; outer bracts of 
the involucre. lanceolate-subulate, usually much 
shorter than the inner ones, which are united 
to about the middle: rays and achenes similar 
to those of the preceding. 


In dry soil, on plains, Nebraska and Wyoming 
to Colorado and New Mexico. In our first edition 
included in T. ambiguum A. Gray, of the South- 
west. June—Aug. 


3. Thelesperma gracile (Torr.) A. Gray. 
Rayless Thelesperma. Fig. 4521. 


Bidens gracilis Torr. Ann. Lyc. N. Y. 2: Ass Tee 
T. gracile A. Gray, Kew. Journ. Bot. 1: 252. 1849. 
Perennial from a deep root; stem rigid, branched, 
1°-3° high, the branches nearly erect. Leaves rigid, 
erect or ascending, 23’ long, pinnately or bipin- 
nately divided into linear segments, or the upper 
linear and entire; heads 6-10” broad; rays usually 
none, sometimes present and 2-3” long; outer bracts 
of the involucre 4-6, oblong or ovate, mostly ob- 
tuse, very much shorter than the inner ones, which 
are united to the middle or beyond; disk yellow or 
brownish; outer achenes slightly papillose; pappus 
awns longer than the width of the summit of the 
achene. 


On dry plains, Nebraska and Wyoming to Texas, 
northern Mexico and Arizona. May—Aug. 


74. GALINSOGA Cav. Icon. 3: 41. 1794. 


Annual branching herbs, with opposite, mostly petioled, dentate or entire leaves, and 
small peduncled heads of both tubular and radiate flowers, terminal and in the upper axils. 
Involucre hemispheric or broadly campanulate, its bracts in 2 series, ovate, obtuse, mem- 
branous, striate, nearly equal, or the outer shorter. Receptacle conic or elongated, its thin 
chaff subtending the disk-flowers. Ray-flowers white or red, pistillate, fertile, the rays 4 or 
5, short. Disk-flowers yellow, perfect, the corolla 5-toothed. Anthers minutely sagittate at 
the base. Style-branches tipped with acute appendages. Achenes angled, or the outer ones 
flat. Pappus of the disk-flowers of several short laciniate or fimbriate scales, that of the 
ray-flowers of several or few short slender bristles, or none. [Named in honor of M. M. 
Galinsoga, superintendent of the Botanic Gardens at Madrid.] 


About 5 species, natives of tropical and warm temperate America, the following typical. 


502 COMPOSITAE. Vor. III. 


1. Galinsoga parviflora Cav. Galinsoga. Fig. 4522. 


Galinsoga parviflora Cav. Icon. 3: 41. pl. 281. 1794. 
Galinsoga parvifiora hispida DC. Prodr. 5: 677. 1836. 

Slightly appressed-pubescent or hirsute, 1°-3° high. 
Leaves thin, ovate or deltoid-ovate, 3-nerved, 1-3’ long, 
acute at the apex, mostly obtuse at the base, dentate, 
the lower slender-petioled, the upper short-petioled or 
sessile, and sometimes nearly or quite entire; heads 
usually numerous, 2’-3’” broad, slender-peduncled; 
bracts of the involucre glabrous or nearly so, the outer 
shorter; pappus of the disk-flowers 4-16, oblong to 
spatulate, fimbriate obtusish or bristle-tipped scales, 
somewhat shorter than or equalling the finely pubescent 
obpyramidal achene. 

In door-yards and waste places, Maine to Ontario, Oregon, 
North Carolina, Missouri, Arizona, California and Mexico. 
Bermuda; Jamaica. Naturalized from tropical America. 
Introduced into Europe as a weed. June—Nov. 

Galinsoga caracasana (DC.) Sch. Bip., similar to this 
species but with reddish rays, and the pappus of the disk- 
flowers only half as long as the achene, also tropical Amer- 
ica, has been found in waste grounds in New Jersey and 
Maryland. 


a 
75. ENDORIMA Raf. Am. Month. Mag. 4: 195. 1819. 
[Batpurna Nutt. Gen. 2: 175. 1818. Not Baldwinia Raf. F. 1818.] 

Perennial, caulescent, simple or branched, erect herbs, with alternate entire narrow punctate 
leaves, and large terminal heads of both tubular and radiate, yellow flowers, or those of the 
disk purple. Involucre hemispheric, its small bracts imbricated in several series, appressed, 
or with spreading tips, the outer shorter. Receptacle convex, deeply honey-combed, chaffy, 
the persistent chaff coriaceous or cartilaginous, laterally united, subtending the disk-flowers. 
Rays large, neutral, toothed. Disk-flowers perfect, fertile, the corolla 5-toothed. Anthers 
sagittate at the base. Style-branches with truncate subulate tips. Achenes turbinate, silky- 
villous. Pappus of 7-12 scarious nearly equal scales. [Greek, perhaps referring to the 
immersion of the-achenes in the honeycombed receptacle. ] 


Two known species, natives of the southwestern United States. Type species: Balduina uni- 
flora Nutt. . 


1. Endorima uniflora (Nutt.) Barnhart. One- 
headed Actinospermum. Fig. 4523. 
Balduina uniflora Nutt. Gen. 2: 175. 1818. 
sae haha unifiorum Barnhart, Bull. Torr. Club 24: 411. 
1697. 
E. uniflora Barnhart; Small, Fl. SE. U. S. 1283. 1903. 

Stem stout, puberulent, simple, or with a few erect 
branches, 1°-3° high. Leaves thick, spatulate-linear or the 
upper linear, sessile, erect or ascending, 1-2" long, the 
lower 2-3” wide; heads long-peduncled, solitary, 2-23’ 
broad; bracts of the involucre ovate, acuminate, thick, their 
tips at length spreading; rays 20-30, cuneate, 3-4-toothed 
at the truncate apex; disk 8’-12” broad; chaff of the re- 
ceptacle cuneate, truncate, very cartilaginous, more or less 
united laterally, the summit eroded; achenes obconic; pap- 
pus of 7-9 oblong scales about as long as the achene. 

In wet pine-barrens, Virginia (according to Torrey and Gray) ; 
North Carolina to Florida and Louisiana. July—Sept. 


76. MARSHALLIA Schreb.; Gmelin, Syst. 1208. 1791. 


Perennial, often tufted, simple or branched herbs, with basal or alternate, entire leaves, 
and large long-peduncled discoid heads of purple, pink or white, glandular-pubescent flowers. 
Involucre hemispheric or broadly campanulate, its bracts in I or 2 series, herbaceous, narrow, 
nearly equal. Receptacle convex or at length conic, chaffy, the scales narrow, rigid, distinct. 
Rays none. Flowers all perfect and fertile, their corollas with a deeply 5-lobed or 5-parted 
campanulate limb and a slender tube. Anthers minutely sagittate at the base. Style-branches 
long, truncate. Achenes turbinate, 5-ribbed and 5-angled. Pappus of 5 or 6 acute or acumi- 
nate, ovate or lanceolate-deltoid, nearly entire scales. [Named for Humphrey Marshall, of 
Pennsylvania, botanical author. ] 

About 6 species, natives of the central United States. Type species: Marshallia Schreberi Gmel. 


= 


GENUS 76. THISTLE FAMILY. 593 


Leaves ovate, oval, or ovate-lanceolate, 3-nerved. 1. M. trinervia. 
Leaves linear, lanceolate, or the basal spatulate, or obovate. 
Chaff of the receptacle linear ; leaves linear; western. 2. M.caespi osa. 
Chatt of the receptacle broader ; leaves obovate to lanceolate; eastern. 3. M. grandiflora. 
j 4 


leaved Marshallia. Fig. 4524. Wa 

Athanasia trinervia Walt. Fl. Car. 201. 1788. “ai / 
Marshallia Schreberi Gmelin, Syst. 1208. 1791. 
Marshallia latifolia Pursh, Fl. Am. Sept. 519. 1814. | / 
Marshallia trinervia Porter, Mem. Torr. Club 5: 337. 1894. f, 

Stem simple, or little branched, leafy to or beyond the 
middle, 1°-2° high. Leaves thin, those of the stem 
ovate, oval or ovate-lanceolate, 3-nerved, acute or acumi- : 
nate at the apex, narrowed to a sessile base, 2’-3’ long, NN 
9-18” wide; heads 2-1’ broad, corolla purplish; bracts 
of the involucre linear-lanceolate, acute, rigid; chaff of 
the receptacle subulate-filiform; pappus scales lanceo- 
late-acuminate from a triangular base; achenes gla- 
brous when mature. 


In dry soil, Virginia to Alabama and Mississippi. May— 5 
June. ; 


. Marshallia trinérvia (Walt.) Porter. Broad- (WKY, 
I arshallia trinérvia ( Walt.) Porter oa Wy 
\\ 


2. Marshallia caespit6sa Nutt. Narrow-leaved 
Marshallia. Fig. 4525. 
Marshallia caespitosa Nutt.; DC. Prodr. 5: 680. 1836. 


Stems usually tufted and simple, sometimes sparingly 
branched, leafy either only near the base or to beyond 
the middle, 8-15’ high. Leaves thick, mostly basal, 
faintly 3-nerved, linear or linear-spatulate, obtuse, some- 
times 4’ long and 3” wide, the upper ones linear, acutish, 
shorter; head about 1’ broad, borne on a peduncle often 
10’ long; corollas pale rose or white; bracts of the, invo- 
lucre linear-lanceolate, acute or acutish; chaff of the 
receptacle linear, or slightly dilated above; achenes vil- 
lous on the angles; scales of the pappus ovate, acutish, 
equalling or longer than the achene. 


Prairies and hills, Missouri and Kansas to Texas. May- 
June. 


3. Marshallia grandiflora Beadle & 
Boynton. Large-flowered Marshallia. 
Fig. 4526. 


Marshallia grandiflora Beadle & Boynton, Bilt- 
more Bot. ‘Stud. 1: 7. 1901. 


Stem simple, 1°-2° high, leafy to above the 
middle. Lower and basal leaves obovate to 
oblong-lanceolate, tapering into petioles often 
as long as the blade, obtuse or obtusish; 
upper leaves lanceolate, sessile, or more or 
less clasping; florets slightly larger than those 
of M. trinervia, 7-10” long; achenes larger, 
2”’-2%” long, pubescent. 


In moist soil, Pennsylvania to West Virginia 
and North Carolina. July—Aug. 


Marshallia obovata (Walt.) Beadle & Boyn- 
ton, a lower plant of the Southern States, with 
obovate or spatulate leaves mainly basal, is re- 
corded as extending northward to southwestern 
Pennsylvania. 


504 COMPOSITAE. Vout 


77. PSILOSTROPHE DC. Prodr. 7: 261. 1838. 
[RippeLtia Nutt. Trans. Am. Phil. Soc. (II) 7: 271. 1841.] 


Branched annual or perennial woolly herbs, often nearly glabrous when old, with alter- 
nate leaves, and middle-sized heads of both tubular and radiate yellow or orange flowers, 
corymbose, or clustered at the ends of the branches. Involucre cylindraceous, its bracts 4-10 
in I series, narrow, equal, densely white-woolly, separate, but erect and connivent, commonly 
with I-4 scarious ones within, and occasionally a narrow outer one. Rays broad, becoming 
papery and whitish, persistent, 5-7-nerved, 2-3-toothed, pistillate. Receptacle small, naked. 
Disk-flowers perfect, fertile, their corollas with a short proper tube and elongated cylindra- 
ceous limb, 5-toothed, the teeth glandular-bearded. Anthers obtuse and entire at the base. 
Style-branches of the disk-flowers capitellate. Achenes linear, striate. Pappus of 4-6 nerve- 
less acute scales, glabrous or villous. [Greek, referring to the naked receptacle. ] 


About 7 species, natives of the southwestern United States and northern Mexico. Type species: 
Psilostrophe gnaphalodes DC. 


1. Psilostrophe villosa Rydb. Plains Psilo- 
strophe. Fig. 4527. 
P. villosa Rydb.; Britton, Manual 1006. t1go1. 


Perennial, branched, 6’-2° high, loosely white- 
woolly. Basal and lower leaves spatulate, entire, den- 
tate or rarely pinnatifid, mostly obtuse, 2’-4’ long; 
upper leaves sessile, or nearly so, smaller, linear to 
spatulate, usually entire; heads several together in 
the clusters, 4-6” broad, short-peduncled; rays few, 
lemon-yellow, commonly as wide as long, with 2 or 
3 broad teeth or lobes at the summit; achenes gla- 
brous, or sparingly pubestent; pappus scales linear- 
lanceolate to oblong-lanceolate, glabrous, shorter 
than the disk-corollas. 


In dry sandy soil, Kansas to Texas and Arizona. In- 
cluded, in our first edition, in P. Tagetinae (Nutt.) 
Kuntze, which has much larger yellow rays. June—Sept. 


78. FLAVERIA Juss.; Gmelin, Syst. 1269. 1791. 


Glabrous or minutely puberulent, light-green, mostly annual herbs, with opposite sessile 
entire or serrate leaves, and small 1-several-flowered, usually sessile, oblong and densely 
cymose-capitate heads of tubular, or both tubular and radiate yellow or yellowish flowers. 
Involucre of 2-5 narrow, nearly equal, appressed bracts, sometimes with 1 or 2 additional 
small exterior ones. Receptacle small, naked or setose. Ray-flower commonly only 1, pistil- 
late, fertile, sometimes wanting. Disk-flowers 1-15, perfect, fertile, their corollas 5-toothed. 
Anthers entire at the base. Style-branches of the disk-flowers truncate. Achenes oblong or 
linear-oblong, 8-10-ribbed. Pappus none. [Latin, flavus, yellow, from its dyeing properties. ] 


About 7 species, natives of the warmer parts of America. In addition to the following, 3 others 
occur in the southern United States. Type species: Flaveria chilensis Gmelin. 


1. Flaveria campestris Johnston. Plains Fla- 
veria. Fig. 4528. 


Flaveria campestris Johnston, Proc. Am. Acad. 39: 287. 

1903. 

Annual, glabrous, erect, 1°-2° high, little branched. 
Leaves linear or lanceolate, serrulate or entire, 3-nerved, 
acuminate or acute at the apex, sessile by a broad and 
somewhat clasping base, 1’—23’ long, 2”—-4” wide; heads | 
about 3” high, closely sessile in terminal glomerules or 
these pedunculate from the upper axils; involucre of 3 
oblong-lanceolate bracts, 2-5-flowered; ray equalling or 
longer than the breadth of the disk; achenes linear, gla- 
brous, about 1%” long. 

In alkaline soil, Missouri to Colorado, Texas and Mexico. 


Aug.-Oct. In our first edition included in the Mexican F. 
angustifolia (Cav.) Pers. 


GENUS 79. THISTLE FAMILY. 505 


79. HYMENOPAPPUS byters Michx: FI. Bor. Am), 2¢ 103... 1803. 


Perennial or biennial, erect herbs, with angled stems, alternate or basal, mostly pinnatifid 
or dissected leaves, and corymbose or solitary, small or rather large discoid heads, of white 
or yellow flowers. Involucre hemispheric or broadly campanulate, its bracts 6-12 in I or 2 
series, nearly equal, mostly appressed, colored, petal-like, the margins and apices scarious. 
Receptacle small, naked. Rays none. Disk-flowers all perfect and fertile, their corollas with 
slender tubes and reflexed or spreading campanulate 5-lobed limbs, the lobes ovate. Anthers 
entire at the base. Style-branches with short conic appendages. Achenes obovoid or obpyra- 
midal, 4-5-angled, the faces usually prominently I-3-nerved. Pappus of 10-20 thin obtuse 
scales, sometimes very short or none. [Greek, membrane-pappus. ] 

About 7 species, natives of southern and central North America and Mexico. Type species: 
Hymenopappus scabiosaéus L’ Her. 


Bracts of the involucre broadly ovate or oval, bright white. 1. H. carolinensis. 
Bracts obovate to oblong, green or with white tips. 
Heads numerous, 4”—6” broad; biennials. 


Achenes puberulent; corolla white. 2. H.corymbosus. 
Achenes densely villous. 
Plant glabrate, or loosely woolly; corolla dull white. 3. H. tenuifolius. 
Plant densely white-woolly ; corolla yellow. 4. H. flavescens. 
Heads few, 6”-12” broad; corolla yellow; perennial. 5. H. filifolius. 


1. Hymenopappus carolinénsis (Lam.) Porter. White-bracted Hymenopappus. 
Fig. 4529. 


Rothia carolinensis Lam. Journ. Hist. Nat. 1: 16. pl. 
Tt. O 2 
ib onebbak scabiosaeus L’Her.; Michx. Fl. Bor. 
Am. 2: 104. 1803. 
Hymenopappus carolinensis Porter, Mem. Torr. Club 
5 335: 15942 
Biennial; stem woolly-pubescent or glabrate, 
leafy below, corymbosely branched and nearly 
naked above, 2°-3° high. Basal and lower leaves 
petioled, 4-6’ long, I-2-pinnately parted or deeply 
pinnatifid into linear or oblong, obtuse or ob- 
tusish lobes, more or less white-tomentose be- 
neath, green and glabrate above; upper leaves 
few, smaller, sessile, less ee es cagr 
monly numerous, corymbose, 6-10” broad; 
bracts of involucre oblong, ovate or oval, some- ZA 
times slightly obovate, thin, bright white, pu- 
berulent or glabrate; corolla-lobes about as long 
as the throat, white; achenes puberulent or pu- 
bescent; pappus of very small nerveless scales, \\ 
shorter than the width of the top of the achene. 


[ANI 


NG 


Z—_ 
SSSA TA 


In dry sandy soil, Illinois to Texas, east to South \ 
Carolina and Florida. March—June. 
SD 


(ir XY Yess 2. Hymenopappus corymbosus T. & G. 
) sry \ Corymbed or Smooth White Hymeno- 
V4 AM pappus, Fig. 4530. 
,) 
Y WA Hymenopappus corymbosus T. & G. Fl. N. A. 2: 372. 
fl" 1842. 
f Biennial; stem glabrous, or nearly so, corymbosely 
yy branched and nearly naked above, 1°-2° high. Lower 
and basal leaves petioled, 1-2-pinnately parted into 
\ linear or nearly filiform, acute or acutish, glabrous 
lobes, or somewhat tomentose beneath; upper leaves 
(WN few, much smaller and less divided, or the upper- 
most reduced to linear scales; heads corymbose, nu- 
merous, 4-6” broad; bracts of the involucre obo- 
vate to oblong, puberulent, their tips greenish white; 
corolla white, its lobes about as long as the throat; 


(|| achenes puberulent ; pappus scales small, nerveless, 
in shorter than the width of the top of the achene. 
Z On dry prairies, Missouri and Nebraska to Texas. 


Summer. 


' 


506 COMPOSITAE. Vor. III. 


3. Hymenopappus tenuifolius Pursh. 
I) Woolly White Hymenopappus. 
Pig: 4531. 


Hymenopappus tenuifolius Pursh, Am. Sept.742. 1814. 


\ 


Biennial; stem lightly tomentose, or at length 
glabrate, 1°-2° high, slender, leafy below, corym- 
bosely branched and nearly naked above. Lower 
and basal leaves petioled, 1-3-pinnately parted 
into linear or filiform lobes, woolly pubescent be- 
neath, at least when young; upper leaves much 
smaller and less compound; heads numerous, co- 
rymbose, 4-6” broad; bracts of the involucre 
obovate-oblong, usually densely tomentose; co- 
rolla white, its lobes slightly shorter than the 
throat; achenes densely villous-pubescent; pappus 
of several oblong to ovate, ribbed or nerved scales, 
which are about as long as the width of the top 
of the achene or shorter. 


On dry prairies, South Dakota to Nebraska, Kansas 
and Texas. June—Sept. 


4. Hymenopappus flavéscens A. Gray. 
Woolly Yellow Hymenopappus. 
Fig. 4532. 


Hymenopappus flavescens A. Gray, Mem. Am. Acad. 
(II) 4:97. 1840. 


f 
\ pe. 
i A igs 
(= an 

oN a, yay 
fo oes SAPS h} 
A Wie 

pA TES, 25 

SES 

be hen 

2 a3 


vanke 


Biennial; stem densely white-woolly, at least 
when young, 1°-23° high, leafy, branched above. 
Leaves I-3-pinnately parted or divided into linear 
segments; heads numerous, usually larger than 
those of the preceding species; involucral bracts 
obovate to ovate with greenish white margins; 
corolla yellow or yellowish, the lobes about equal- 
ling the throat, achenes short-villous; pappus scales 
spatulate, shorter than the slender corolla-tube. 


In sandy soil, Kansas to Texas, Arizona and north- 
ern Mexico. 


5. Hymenopappus filifolius Hook. Low 
Tufted Hymenopappus. Fig. 4533. 


Hymenopappus filifolius Hook. Fl. Bor. Am.1: 317. 1833+ 


Perennial from a deep woody root; stems usually 
tufted, woolly when young, sometimes glabrate when 
old, densely leafy toward the base, usually naked or 
nearly so and sparingly branched above, 6-18’ high. 
Leaves tomentose when young, the lower and basal 
ones petioled, I-3-pinnately parted or pinnatifid into 
narrowly linear, somewhat rigid lobes; heads com- 
monly few, 6-12” broad; bracts of the involucre 
obovate-oblong, usually densely woolly, their tips 
whitish; corolla yellow or yellowish, its lobes much 
shorter than the throat; achenes densely villous; 
pappus scales costate, short. 


On prairies and in dry rocky soil, Saskatchewan to 
oe Dakota, Montana, Nebraska and Colorado. June- 
ept. 


GENUS 8o. THISTLE FAMILY. 507 


80. OTHAKE Raf. New FI. N. A. 4:73. 1836. 


Erect rough, glandular or cinereous, branching annual herbs, with alternate, mostly entire 
leaves, or the lower opposite, and corymbose or panicled heads of tubular or both tubular 
and radiate pink or purple flowers. Involucre campanulate or obconic, its bracts in I or 2 
series, narrow, herbaceous, nearly equal, or with a few exterior shorter ones, appressed, 
usually colored. Receptacles small, flat, naked. Ray-flowers, when present, pistillate, fertile, 
the rays 3-cleft. Disk-flowers perfect, fertile, their corollas with slender tubes and deeply 
5-parted campanulate limbs. Anthers entire or emarginate at the base. Style-branches fili- 
form, acutish, glandular-pubescent throughout. Achenes linear or narrowly obpyramidal, 
quadrangular. Pappus of 6-12 lanceolate strongly costate scales, that of the outer achenes 
often much shorter. [Greek, warty apex, referring to the callous-tipped leaves of some 
species. | 

About 6 species, natives of the south-central United States and Mexico; in our first edition 
referred to the genus Polypteris Nutt. Type species: Othake tenuifolium Raf. 


Rays purple, deeply 3-lobed ; leaves lanceolate. 1. O. sphacelatum. 
Rays none; leaves linear. 2. O. callosum. 


1. Othake sphacelatum ( Nutt.) Rydb. Hooker’s __,, 
Othake. Fig. 4534. & 

Stevia sphacelata Nutt.; Torr. Ann. Lyc.N. Y.2: 214. 1827. 
Palafoxia Hookeriana T. & G. Fl. N. A. 2: 368. 1842. 
peers Hookeriana A. Gray, Proc. Am, Acad. 19: 31. 

1883. 
O. sphacelatum Rydb. Bull. Torr. Club 37: 331. Igto. 

Annual; stem rather stout, glandular-pubescent and 
viscid above, 1°-3° high. Leaves lanceolate, entire, 
acute or acuminate, narrowed at the base, rough on both 
sides, the upper alternate, the lower opposite and slen- 
der-petioled, 2-4’ long, 3-5” wide; bracts of the invo- 
lucre 10-16, linear-lanceolate or spatulate, glandular- 
hispid, the inner with purplish tips; ray-flowers 8-10; 
rays rose-purple, deeply 3-cleft, sometimes small, or 
none; achenes about 4” long and 4” thick; pappus scales 
of the disk-flowers 6-8, lanceolate, awned, more than 
half the length of the achene, those of the ray-flowers 
as many, spatulate, obtuse, shorter. 

In dry soil, Nebraska to Colorado, Texas and Mexico. 
July—Sept. 


2. Othake callosum (Nutt.) Bush. Rayless 
Othake. Fig. 4535. 


S'evia callosa Nutt. Journ. Acad. Phila. 2: 121. 1821. 
O. tenuifolium Raf. New Fl. N. A. 4: 74. 1836. 

¢ Poiypteris callosa A. Gray, Proc. Am. Acad.19: 30. 1883. 
O. callosum Bush, Trans. Acad. Sci. 14; 174. 1904. 


Annual, glandular, at least above; stem slender, 
paniculately branched, 1°-2° high. Leaves linear, 
or linear-lanceolate, 1-23’ long, 1-23” wide, entire, 
short-petioled, mostly alternate; heads discoid; 
bracts of the top-shaped involucre 8-10, linear or 
narrowly oblong, herbaceous, pubescent, about + 
long, their tips reddish; corollas purple, deeply 5- 
parted; achenes narrowly obpyramidal, pubescent or 
glabrous, nearly as long as the involucre; pappus 
scales obovate or nearly orbicular, rounded, or re- 
tuse, or sometimes minute, or none. 


In dry soil, Missouri to Texas and New Mexico. 
June- Oct. 


81. PICRADENIOPSIS Rydb.; Britton, Man. roo8. 190T. 


Herbs more or less woolly, with opposite leaves, and small corymbose heads, of both 
tubular and radiate yellow flowers. Involucre campanulate or hemispheric, its bracts in I 
or 2 series, herbaceous, obtuse, appressed, nearly equal. Receptacle small, nearly flat, naked, 
foveolate. Ray-flowers in I series, pistillate, fertile. Disk-flowers perfect, fertile, their corollas 


508 COMPOSITAE. Vor. IIT. 


with campanulate or cylindric, 5-cleft limb. Anthers entire or emarginate at the base. Style- 

tips mostly truncate and obtuse. Achenes quadrangular, linear or oblong. Pappus of several 

scales, obtuse or truncate and scarious at the apex. [Named for its resemblance to Picradenia.] 
Two known species, natives of western North America, the following typical. 


1. Picradeniopsis oppositifolia ( Nutt.) 
Rydb. Picradeniopsis. Fig. 4536. 
Trichophyllum oppositifolium Nutt. Gen.2: 167. 1818. 


Bahia oppositifolium Nutt.; T. & G. Fl. N. A. 2: 376. 
1842. 


P. oppeositifolia Rydb.; Britton, Manual 1008. i901. 


Perennial, herbaceous; stem densely cinereous, 
much branched, 4-12’ high, very leafy. Leaves 
opposite, or the uppermost alternate, 4/—12’ long, 
palmately 2-5-parted into linear, obtuse or ob- 
tusish, entire segments, finely cinereous on both 
sides; heads short-peduncled, 6-9” broad; invo- 
lucre campanulate, or becoming hemispheric, its 
bracts oblong, obtuse, densely tomentose; rays 
5-7, short; achenes linear-oblong, glandular-pu- 
bescent; pappus of 4-8 spatulate to lanceolate 
scales with thickened bases. 


On plains, especially in alkaline soil, South Da- 
kota to Montana, Nebraska, Texas, New Mexico. 
June—Sept. 


82. TETRANEURIS Greene, Pittonia 3: 265. 1808. 
[AcTINELLA Nutt. (1818), not Pers. (1807), nor Actinea Juss. (1803).] 

Branched or scapose, villous-pubescent or glabrous, bitter and aromatic punctate herbs, 
with alternate or basal, often punctate leaves, and small or rather large, peduncled heads of 
both tubular and radiate, yellow flowers, or rays rarely wanting. Involucre hemispheric, 
campanulate or depressed, its bracts imbricated in 2-3 series, appressed. Receptacle convex 
or conic, naked. Ray-flowers pistillate and fertile, the rays 3-toothed, 4-nerved. Disk-flowers 
perfect, fertile, their corollas with 4-5-toothed limbs. Anthers entire or minutely sagittate 
at the base. Style-branches truncate and penicillate at the summit. Achenes turbinate, 5—10- 
ribbed or angled, villous or pubescent. Pappus of 5-12 thin aristate, acuminate or truncate 
scales. [Greek, four-nerved.] 

About 18 species, natives of western North America and Mexico. Besides the following, some 
12 others occur in the western and southwestern parts of the United States. Type species: Te ra- 
neuris acaulis (Pursh) Greene. 


Stem leafy, branching; stem leaves linear ; annual or biennial. 1. T. linearifolia. 
Stems tufted, simple, scapose ; leaves basal; perennials. 
Leaves narrowly linear; branches of the caudex slender. 2. T. stenophylla. 
Leaves broader, linear to spatulate; branches of the caudex short and thick. c 
Bracts of the involucre acutish. 3. T. acaulis. 
Bracts of the involucre obtuse, rounded. 4. T. herbacea. 


1. Tetraneuris linearifolia (Hook.) Greene. 
Fine-leaved Tetraneuris. Fig. 4537. 


Hymenoxys linearifolia Hook. Icon. pl. 146. 1837. 
Actinella linearifolia T. & G. Fl. N. A. 2: 383. 1842. 
T. linearifolia Greene, Pittonia 3: 369. 1898. 


Annual or perhaps biennial; stem usually diffusely 
branched, finely hirsute, or glabrous, or woolly at 
the base, slender, 6-15’ high. Stem leaves narrowly 
linear, sessile, 3-13’ long, 3’”-1” wide; basal leaves 
spatulate, often villous, much broader, obtuse, nar- 
rowed into margined petioles; heads numerous, 
slender-peduncled, 6”-8” broad; involucre broadly 
campanulate, its bracts oblong, obtuse, pubescent, 
distinct to the base, imbricated in about 2 series; 
receptacle conic; rays 6-10, oblong; achenes pubes- 
cent; pappus of 5 or 6 ovate awned scales. 


In dry soil, Kansas to Louisiana, Texas and New 
Mexico. May-—Sept. 


7 
j 
{ 


Genus 82. THISTLE FAMILY. 509 


\ 


2. Tetraneuris stenophylla Rydb. Narrow- 
leaved Tetraneuris. Fig. 4538. 


Tetraneuris stenophylla Rydb. Bull. Torr. Club 33: 155. 
1906, 


Perennial with a branched caudex, the branches 
often 33’ long, the upper portion covered with the 
scarious bases of old leaves. Bases of the leaves 
dilated, sparingly silky-villous; blades narrowly lin- 
ear, 1-1} long, about 1” wide, glabrous, conspicu- 
ously punctate; scape 2-4’ high, minutely strigose; 
involucre 5’—-7” wide; bracts oval or oblong-acutish ; 
rays about 5” long and 2” wide. 


In dry soil, Kansas to Colorado and Mexico. Formerly 
confused with 7. scaposa (DC.) Greene. 


Tetraneuris fastigiata Greene, of Kansas, differs by 
the leaf-bases and involucre being more densely pubes- 
cent; it is known only from imperfect specimens, and 
may not be distinct from T. stenophylla, over which it 


YY Jp 
Hj SS 3. Tetraneuris acatlis (Pursh) Greene. 
iis Stemless Tetraneuris. Fig. 4539. 


Gaillardia acaulis Pursh, Fl. Am. Sept. 743. 1814. 

T. acaulis Greene, Pittonia 3: 265. 1808. 

Actinella acaulis Nutt. Gen. 2: 173. 1818. 
Picradenia acaulis Britton, Mem. Torr. Club 5: 3309. 


1894. 
T. simplex A. Nelson, Bot. Gaz. 28: 127. 1899. 

Perennial with thick roots and a stout branched 
caudex; scapes tufted, rather stout, or slender, 
densely silky or tomentose, 2’-8’ high. Leaves all 
borne on the ends of the branches of the caudex, 
linear-spatulate, entire, obtuse or obtusish, 1-3’ 
long, 12-3’ wide, densely silky or villous; heads 
9-18” broad; involucre hemispheric, its bracts 
densely villous, acutish; rays 10-15; pappus of 5 
or 6, ovate or oblong, awned scales. 

In dry or rocky soil, North Dakota to Assiniboia, 
Montana, Nebraska and New Mexico. May—Aug. 

Tetraneuris scaposa (DC.) Greene, admitted into 
our first edition as from Nebraska, is not definitely 
known north of Texas. 


pt ee 
ee ae 


4. Tetraneuris herbacea Greene. Eastern Tetra- vy 
neuris. Fig. 4540. 5 

T. herbacea Greene, Pittonia 3: 268. 1808. 

Actinia herbacea Robinson, Rhodora 10: 68. 1908. 

Perennial by a stout thick caudex; scape stout, villous- 
pubescent, especially above, 6-8’ high. Leaves linear- 
spatulate, slightly fleshy, bluntish, strongly punctate, spar- 
ingly loosely long-hairy, at least toward the base, 2-3’ long, 
about 3” wide; heads nearly 2’ broad; involucral bracts 
oblong, obtuse and rounded at the apex, densely villous; 
rays about 15; pappus scales ovate-oblong, obtuse, acute or 
apiculate. 

Southern Ontario, Ohio and Illinois. May—June. 


83. HYMENOXYS (Cass. Dict. Sci. 
Nat. 55: 278. 1828. 
[PicrapENIA Hook. Fl. Bor. Am. 1: 317. 1833.] 
Pubescent herbs with branching stems and alternate entire or dissected leaves, the blades 


or divisions narrow, usually linear or filiform, and relatively small, peduncled, radiate or 
discoid yellow heads. Involucre turbinate to campanulate, its bracts in 2 series, appressed, 


58) COMPOSITAE. Vor. IIT. 


rather broad, the outer connate at base. Receptacle flat, convex or conic. Ray-flowers fertile, 
the rays short and broad, 3-lobed. Disk-flowers perfect, fertile, with 5 short lobes. Anthers 


ENS os. notched at the base, with rounded auricles. Style- 
mes KN 
yA NU 
Ne 


Za MS branches truncate and penicillate. Achenes turbi- 
ri aS nate, pubescent. Pappus of 5-8 acuminate or aristate 


hyaline scales. [Greek, referring to the thin and 
pointed pappus scales. ] 
Type species: Hymenopappus anthemoides Juss. 


1. Hymenoxys odorata DC. Limonillo. 


Fig. 4541. 
Hymenoxys odorata DC. Prodr. 5: 661. 1836. 
es odora‘a A. Gray, Mem. Am. Acad. (II) 4: ror. 
1840. 
Picradenia odorata Britton, in Britt. & Brown, Ill. Fl. 3: 
449. 1808. 
pg multiflora Buckl. Proc. Acad. Phila. 1861: 459. 
I Ze 
Hi. multiflora Rydb. Bull. Torr. Club 33: 157. 1906. 
Annual; stem much branched, puberulent, spar- 
ingly hirsute or glabrous, 1°-2° high, leafy. Leaves 
1-2’ long, 1-3-parted into filiform, entire, somewhat 
pubescent segments about 4” wide; heads commonly 
numerous, 6-10” broad; involucre campanulate, pu- 
berulent, its outer bracts 6-9, lanceolate, keeled, 
acute, united at. the base; rays 7-10, cuneate. 
In dry soil, Kansas and Colorado to Texas, Mexico 
and southern California. April—July. 


84. HELENIUM L. Sp. Pl. 886. 1753. 


Erect, mostly branching herbs, with alternate, mainly decurrent, punctate bitter entire or 
dentate leaves, and large peduncled heads of both tubular and radiate, yellow or brownish- 
yellow flowers, or rays sometimes wanting. Involucre broad and short, its bracts in I or 2 
series, linear or subulate, reflexed or spreading. Receptacle convex, subglobose or oblong, 
naked. Ray-flowers .pistillate and fertile, or neutral, the rays cuneate, 3-5-lobed. Disk- 
flowers perfect, fertile, their corollas 4-5-toothed, the teeth glandular-pubescent. Anthers 
2-toothed or sagittate at the base. Style-branches of the disk-flowers dilated and truncate at 
the apex. Achenes turbinate, ribbed. Pappus of 5-8 entire, dentate or incised, acuminate 
or aristate scales. [The Greek name of some plant, from Helenus or Helena.] 


About 24 species, natives of North and Central America. In addition to the following, some 18 
others occur in the southern and southwestern United States. Type species: Helenium autumnale L. 


Leaves oblong-lanceolate or ovate-lanceolate, dentate; rays fertile; disk yellow. 1. H. autumnale. 
Leaves lanceolate or linear-lanceolate, mainly entire; rays neutral; disk purple. 2. H. nudiflorum. 
Leaves all linear-filiform, entire; rays fertile. 3. H. tenuifolium. 


1. Helenium autumnale L. False or 
Swamp Sunflower. Fig. 4542. 


Helenium autumnale L. Sp. Pl. 886. 1753. 

Helenium pubescens Ait. Hort. Kew. 3: 287. 1789. 

Helenium autumnale pubescens Britton, Mem. 
Torr. Club 5: 339. 1804. 


Perennial; stem puberulent or glabrous, 
rather stout, narrowly winged by the decur- 
rent bases of the leaves, corymbosely branched 
above, 2°-6° high. Leaves firm, oblong, lan- 
ceolate or ovate-lanceolate, acuminate or acute 
at apex, narrowed to the sessile base, pinnately 
few-veined, 2’-5’ long, #/-2’ wide, dentate, den- 
ticulate or entire, puberulent, glabrous or pu- 
bescent, bright green; heads numerous, 1-2’ 
broad, borne on long puberulent peduncles; 
bracts of the flattish involucre densely canes- 
cent; rays 10-18, drooping, bright yellow, 
equalling or longer than the globose yellow 
disk, pistillate and fertile, 3-cleft; achenes pu- 
bescent on the angles; pappus scales ovate. 

In swamps and wet meadows, Quebec to Flor- 
ida, Manitoba, Oregon, Nevada and Arizona. Yel- 
low-star. Ox-eye. Sneezeweed. Ascendsto 2600 ft. 
in Virginia. Aug.—Oct. 


GENUS 84. THISTLE FAMILY. 511 


2. Helenium nudifldrum Nutt. Purple- 
head Sneezeweed. Fig. 4543. 
Helenium nudiforum Nutt. Trans. Am. Phil. Soc. 

CD) 7s 384. © rear. 

Benabods brachypoda T. & G. Fl. N. A. 2: 388. 

1842. 

Perennial; stem mostly slender, puberulent at 
least above, corymbosely branched near the sum- 
mit, 1°-3° high, narrowly winged by the decur- 
rent leaf-bases. Stem leaves lanceolate or linear- 
lanceolate, entire or sparingly denticulate, acute 
or obtusish at the apex, 12-3’ long, 2-6” wide, 
sessile; basal and lower leaves spatulate, obtuse, 
more or less dentate, tapering into margined 
petioles; heads several or numerous, 1-12’ broad, 
on slender or short-puberulent peduncles; rays 
10-15 (sometimes wanting), drooping, yellow, 
yellow with a brown base, or brown throughout, 
3-toothed, neutral, or with rudimentary pistils, 
sterile, equalling or exceeding the brown or 
purple globose disk; pappus scales ovate, aristate. 

In moist soil, Missouri and Illinois to Texas, east 
to North Carolina and Florida. Also locally natural- 
ized from Pennsylvania to Connecticut. June—Oct. 


3. Helenium tenuifolium Nutt. [ine- 
leaved Sneezeweed. Fig. 4544. 


H. tenuifolium Nutt. Journ. Phil. Acad. 7: 66. 1834. 


Annual; glabrous or minutely pubescent above ; 
stem slender, very leafy and usually much 
branched, 8’-24’ high. Leaves all linear-filiform, 
entire, sessile, often fascicled, 4’-13’ long, 3” or 
less wide; heads several or numerous, corym- 
bose, 9-15” broad, borne on slender or filiform 
peduncles; bracts of the involucre few, linear or 
subulate, sometimes pubescent, soon reflexed; 
rays 4-8, fertile, 3-4-toothed, at length drooping, 
longer than the globose disk; achenes villous; 
pappus scales ovate, tipped with slender awns. 


In moist soil, southeastern Virginia to Florida, 
Missouri, Kansas and Texas. Naturalized in waste 
places, northward to Massachusetts, and in Cuba and 
Santo Domingo. Aug.—Oct. 


85. GAILLARDIA Foug. Mem. Acad. Sci. Paris 1786: 5. pl. 1, 2. 1788. 


Branching or scapose, more or less pubescent herbs, with alternate or basal leaves, and 
large peduncled heads of both tubular and radiate flowers, or rays wanting. Involucre 
depressed-hemispheric, or flatter, its bracts imbricated in 2 or 3 series, their tips spreading or 
reflexed. Receptacle convex or globose, bristly, fimbrillate or nearly naked. Rays cuneate, 
yellow, purple or parti-colored, neutral or rarely pistillate, 3-toothed or 3-lobed. Disk-flowers 
perfect, fertile, their corollas with slender tubes and 5-toothed limbs, the teeth pubescent with 
jointed hairs. Anthers minutely sagittate or auricled at the base. Style-branches tipped 
with filiform or short appendages. Achenes turbinate, 5-ribbed, densely villous, at least at 
the base. Pappus of 6-12, I-nerved awned scales, longer than the achene. [Named for M. 
Gaillard de Marentonneau, a French botanist. ] 


About 1s species, natives of the south-central part of the United States, and Mexico, 1 in 
southern South America. Type species: Gaillardia pulchella Foug. Called in Texas blanket-flower. 
Stem leafy ; style-tips with filiform hispid appendages. 

Fimbrillae of the receptacle obsolete, or short. 1. G. lutea. 
Fimbrillae subulate or bristle-like, mostly longer than the achenes. 


Rays yellow; fimbrillae exceeding the achenes. 2. G. aristata. 
Rays purple, or red at base; fimbrillae about equalling the achenes. 3. G. pulchella. 
Leaves basal; style-tips with short naked appendages ; rays none, or few. 4. G. suavis. 


COMPOSITAE. — Vot. III. 


1. Gaillardia lutea Greene. Yellow Gaillardia. 
Fig. 4545. 


Gaillardia lutea Greene, Pittonia 5: 57. 1902. 


Stem roughish-puberulent or cinereous, usually branched, 
13°-2° high, the branches straight, nearly erect. Stem leaves 
sessile, oblong-lanceolate, serrate, roughish-puberulent, acute 
at the apex, narrowed to the base, 17-2’ long, 2”-5”” wide; 
heads about 2’ broad, peduncled; bracts of the involucre about 
equalling the yellow disk; rays 8-12, yellow; style-tips with 
filiform hispid appendages; achenes villous at the base, or to 
beyond the middle; fimbrillae of the receptacle short or none; 
awns of the pappus slender. 


In dry woods, Missouri to Texas. July-Sept. Included, in our 
first edition, in G. /anceolata Michx., of the Southern States. 


2. Gaillardia aristata Pursh. Great- 
flowered Gaillardia. Fig. 4546. 
Gaillardia aristata Pursh, Fl. Am. Sept. 573. 1814. — 


Perennial; stem simple, or little branched, 
hirsute or densely pubescent with jointed hairs, 
1°-3° high. Leaves firm, densely and finely 
pubescent, the lower and basal ones _petioled, 
oblong or spatulate, laciniate, pinnatifid or en- 
tire, mostly obtuse, 2’-5’ long; upper leaves ses- 
sile, lanceolate, or oblong, or slightly spatulate, 
smaller, entire or dentate, rarely pinnatifid; 
heads 12-4’ broad, long-peduncled; bracts of 
the involucre lanceolate, acuminate, hirsute; 
rays 10-18, yellow; style-tips with filiform ap- 
pendages; fimbrillae of the receptacle mostly 
longer than the achenes, which are villous at 
least at the base. 


On plains and prairies, Minnesota to Saskatche- 
wan, British Columbia, Colorado, New Mexico and 
Oregon. Adventive eastward. Leaves sometimes 
all basal. May—Sept. 


3. Gaillardia pulchélla Foug. Showy 
Gaillardia. Fig. 4547. 


Gaillardia pulchella Foug. Mem. Acad. Sci. Paris 
1786: 5. 1786. 


Annual; diffusely branched at the base, the 
branches ascending, 6-15’ high, or larger in 
cultivation, more or less hirsute or pubescent 
with jointed hairs. Leaves lanceolate, oblong, 
or the lower spatulate, 1-3’ long, entire, den- 
tate or sinuate-pinnatifid, all but the lowest 
sessile; heads 17-3’ broad, long-peduncled, 
bracts of the involucre lanceolate, acuminate, 
hirsute or pubescent; rays 10-20, red or purple 
at the base, yellow toward the apex; style-tips 
with filiform hispid appendages; fimbrillae of 
the receptacle equalling or scarcely longer than 
the achenes, which are more or less villous, 
or glabrous. 


In dry soil, Nebraska and Missouri to Louisi- 
ana, Mexico and Arizona. May-—Sept. 


Genus 85. THISTLE FAMILY. 513 


4. Gaillardia suavis (A. Gray) Britt. & 
Rusby. Rayless Gaillardia. Fig. 4548. 


Agassizia suavis A. Gray, Proc. Am. Acad. 1: 49. 1846. 

Gaillardia simplex Scheele, Linnaea 22: 160. 1849. : 

Gaillardia suavis Britt. & Rusby, Trans. N. Y. Acad. Sci. 
Wat) L887 


Annual or biennial. Leaves in a basal tuft, or a 
few near the base of the slender pubescent scape, 
spatulate or obovate in outline, 2-6’ long, pinnatifid, 
dentate, or some of them entire; scape 1°-2° high, 
monocephalous; head about 1’ broad with the odor 
of heliotrope, globose in fruit; rays none, or short 
and pistillate, or a few of them longer and neutral; 
bracts of the involucre oblong or lanceolate, spar- 
ingly pubescent; fimbrillae of the receptacle obso- 
lete; style-appendages short, naked; achenes densely 
villous; pappus scales broad, their awns very slender. 


In dry rocky soil, Kansas to Texas. April-June. “4 } 


86. BOEBERA Willd. Sp. Pl. 3: 2125. 1804. 


Erect or diffuse, branching, annual, or perennial, strong-scented, more or less glandular 
herbs, with opposite, mostly finely dissected leaves, and small peduncled heads of both tubular 
and radiate yellow flowers. Involucre campanulate or nearly hemispheric, its bracts in I 
series, united into a cup, with small additional outer ones. Receptacle flat, pubescent. Ray- 
flowers pistillate, the rays short. Disk-flowers perfect, their corollas 5-toothed. Anthers 
entire or minutely 2-toothed at the base. Style-branches of the disk-flowers hirsute, apicu- 
late. Achenes narrowly obpyramidal, 3-5-angled, striate. Pappus of about 10 scales, parted 
to beyond the middle into numerous capillary, bristle-like segments. [In honor of J. von 
Boeber, a Russian botanist, died 1820, ] 


About 3 species, natives of the central United States and of Mexico, the following typical. 


1. Boebera papposa ( Vent.) Rydb. Fetid Mari- 
gold. False Dog-fennel. Fig. 4549. 

Tagetes papposa Vent. Hort. Cels. pl. 36. 1800. 

Bocebera chrysanthemoides Willd. Sp. Pl. 3: 2125. 1804. 

Dysodia chrysanthemoides Lag. Gen. et Sp. Nov. 29. 1816. 

D. papposa Hitche. Trans. St. Louis Acad. 5: 503. 1891. 

B. papposa Rydb.; Britton, Manual 1tor2. 1901. 

Annual, very leafy, glabrous or finely pubescent, 
gland-dotted, much branched, 6-18’ high, the branches 
diffuse or erect. Leaves sessile, or short-petioled, +’—12’ 
long, pinnately parted into linear or slightly spatulate, 
sharply serrate or incised: segments; heads numerous, 
short-peduncled, 3-5” broad; involucre campanulate, 
of 8-10 appressed, oblong, obtuse, green or purplish, 
glabrous or ciliate bracts, with several narrow shorter 
outer ones; rays few, not longer than the width of the 
disk; receptacle and achenes pubescent. 

Along streams and roadsides, Ohio to Minnesota, Mon- 
tana, Louisiana, Mexico and Arizona. Occasionally found 


as a weed in waste places in the Eastern and Middle States, 
and in Ontario. Prairie-dogweed. July—Oct. 


87. THYMOPHYLLA Lag. Gen. et Sp. Nov. 25. 1816. 


[HyMENATHERUM Cass. Bull. Soc. Philom. 1817: 12. 1817.] 

Annual or perennial herbs, some species low undershrubs, with gland-dotted foliage and 
involucre, alternate or opposite leaves, and small heads of both tubular and radiate, mostly 
yellow flowers. Involucre campanulate, its principal bracts united into a cup, sometimes 
with smaller outer ones. Receptacle naked, or fimbrillate, not chaffy. Ray-flowers pistillate, 
fertile. Disk-flowers perfect, fertile. Style-branches truncate or blunt. Achenes striate. 
Pappus of several or numerous scales or bristles. |Greek, thyme-leaf, not applicable to the 
following species. ] 


About 15 species, natives of America. Besides the following, some 4 others occur in the west- 
ern parts of the United States. Type species: Thymophylla setifolia Lag. 


38) 


514 COMPOSITAE. Vor. III. 


K ; VG: Sse 


1. Thymophylla aurea (A. Gray) Greene. 
Thyme-leaf. Fig. 4550. 


pei pe aurea A. Gray, Mem. Am. Acad. (II) 4: 91. 
1849. 
Hymena herum aureum A. Gray, Proc. Am. Acad. 19: 
42. 1883. 
T. aurea Greene; Britt. & Brown, Ill. Fl. 3: 453. 1808. 
Annual, glabrous, 4-12’ high, much branched; the 
leaves and involucre with large oval oil-glands. Leaves 
alternate, or the lower opposite, sessile or nearly so, 
very deeply parted into 5-9 linear-filiform, mostly 
entire, blunt segments; heads numerous, corymbose, 
6-10” broad, terminating the branches; involucre 
about 3” high, its bracts acute; rays about 12, 23-3” 
long; pappus of 6-8 erose truncate scales, somewhat 
longer than the thickness of the achene. 
Kansas and Colorado to Texas and New Mexico. 
June—Sept. 


88. PECTIS L. Syst. Nat. Ed. to, 1221) 
1759. 

Annual or perennial, diffuse prostrate or erect, mostly glabrous herbs, gland-dotted and 
strong-scented, with opposite narrow sometimes ciliate leaves, and small usually cymose 
heads of both tubular and radiate yellow flowers. Involucre cylindric, oblong or campanu- 
late, its bracts in 1 series, narrow, keeled, distinct. Receptacle small, naked. Ray-flowers 
pistillate, the rays small, entire or 3-lobed. Disk-flowers perfect, their corollas with expanded, 
somewhat irregularly 5-cleft limbs. Anthers entire at the base. Style-branches of the disk- 
flowers very short, obtuse. Achenes linear, slightly angled, striate. Pappus of several or 
numerous scales, slender bristles or awns, sometimes with a few outer smaller additional 
ones. [Latin, pecten, comb, referring to the pappus.] 


About 75 species, natives of the warmer parts of America. Besides the following, about 10 
others occur in the southern and western parts of the United States. Type species: Pectis ciliaris L. 


1. Pectis angustifolia Torr. Lemon-scented 
Pectis: © Pigs4s5t: 

Pectis angustifolia Torr. Ann. Lye. N. Y. 2: 214. 1827. 

Annual, much branched, 4-12’ high, the branches 
diffuse or ascending. Leaves narrowly linear, sessile, 
obtusish, 3/-2’ long, 1” wide or less, often ciliate with 
a few bristles near the base; heads several or numer- 
ous, short-peduncled, about 3” broad; involucre short- 
cylindric or narrowly campanulate, its bracts about 8, 
linear, acutish, partly enclosing the outer achenes; rays 
few, 3-toothed, or entire; pappus a crown of 4-6 some- 
what united short scales, with or without 2 slender 
short awns. 

In dry soil, Nebraska and Colorado to Mexico and Ari- 
zona. Plant with the odor of lemons. May-—Oct. 


89. ACHILLEA [Vaill.] L. Sp. Pl. 898. 1753. 


Herbs, mostly perennial, with erect leafy stems, finely 
dissected, pinnatifid or serrate alternate leaves, and small 
heads of both tubular and radiate flowers, corymbose at 
the ends of the stem and branches. Involucre obovoid, or campanulate, its bracts appressed, 
imbricated in few series, the outer shorter. Receptacle nearly flat or convex, chaffy, the mem- 
branous chaff subtending the disk-flowers. Ray-flowers pistillate, fertile, the rays white or 
pink. Disk-flowers perfect, fertile, their corollas yellow, 5-lobed. Anthers obtuse and entire 
at the base. Style-branches of the disk-flowers truncate. Achenes oblong or obovate, slightly 
compressed. Pappus none. [Named for Achilles.] 

About 75 species, natives of the northern hemisphere, mostly of the Old World. Besides the 
following, another, or perhaps 2 others, occur in northwestern North America. Type species: 
Achillea santolina L. 

Involucre broadly campanulate; leaves serrate. 1. A. Ptarmica. 
Involucre ovoid to cylindric; leaves finely dissected. 
Involucral bracts greenish-yellow; rays small. 
Plant loosely woolly or nearly glabrous; inflorescence flat-topped. 2. A. Millefolium. 
Plant densely woolly ; inflorescence convex. 3. A. lanulosa. 
Involucral bracts black-margined ; rays large. 4. A. borealis. 


GENUS 80. THISTLE FAMILY. 


1. Achillea Ptarmica L. Sneezewort. White 
Tansy. Sneezewort-Yarrow. Fig. 4552. 


Achillea Ptarmica L. Sp. Pl. 898. 1753. 


Perennial from horizontal or creeping rootstocks ; 


stem glabrous, or slightly pubescent, nearly or quite 
simple, 1°-2° high. Leaves linear or linear-lanceolate, 
sessile and slightly clasping at the base, acute at the 
apex, regularly and closely serrate, sometimes pubes- 
cent on the veins beneath, 1-23’ long, 13’-3” wide; 
heads not very numerous, 5’—9” broad; peduncles pu- 
berulent; involucre broadly campanulate, its bracts 
ovate-oblong, obtuse or obtusish, slightly tomentose; 
rays 5-15, white, rather large. 


In moist soil, Newfoundland, New Brunswick and Quebec 
‘to Massachusetts and Michigan. Naturalized from Europe. 
Native also of northern Asia. Goose-tongue. Wild, bastard- 
er european pellitory. Fair-maid-of-France. Sneezewort- 
tansy. July—Sept. 
CRESS 
2 isararhangeraneste Nc 
SR ER PAR 


eX \ | |/ 


Ny 


2. Achillea Millefolium L. Yarrow. 


Milfoil. Fig. 4553. 


Achillea Millefolium L. Sp. Pl. 899. 1753. 
Perennial from horizontal rootstocks; flow- 


3. Achillea lanulosa Nutt. 


Woolly Yar- 


ering stems pubescent, or nearly glabrous, sim- 
ple, or corymbosely branched above, 1°-2° 
high. Basal leaves, and those of the numerous 
short sterile shoots, mostly petioled, sometimes 
10’ long and 3’ wide, those of the stem sessile, 
all narrowly oblong or lanceolate in outline 
and finely dissected into narrow pinnatifid seg- 
ments, tomentose, pubescent or nearly glabrous; 
heads numerous, 2’—3” broad, in terminal com- 
pound dense, somewhat convex or nearly flat- 
topped corymbs; involucre ovoid-cylindric, its 
bracts oblong, obtusish, pubescent; rays 4-6, 
white, or often pink or purple, less than 2” 
broad. 

In various situations throughout eastern North 
America, often occurring as a naturalized weed. 
Native also of Europe and Asia. Old names, san- 
guinary, thousand-leaf, nosebleed, old-man’s-pep- 
per, soldier’s-woundwort, gordaldo. June—Nov. 


a 


oh 


row. Fig. 4554. 


Achillea lanulosa Nutt. Journ. Acad. Phila. 7: 36. 
1824. 


Similar to the preceding species, perennial by 
rootstocks, 1°-24° high, densely  silky-woolly 
nearly all over. Leaves deeply bipinnatifid into 
narrow lobes and segments, those of the stem 
mostly sessile; inflorescence convex, 2’—4’ broad; 
involucre oblong-cylindric, its bracts greenish- 
Ag with brownish margins; rays 1’-23” broad, 
white. 


In dry soil, Quebec and Ontario to Michigan, 
Yukon, south to Oklahoma, Mexico and California. 
June-Sept. Locally naturalized eastward. 


Achillea ligiistica All., differing from A. Millefo- 
lium by being stouter with loosely corymbose heads, 
native of Europe, has been found in cultivated ground 
near Tannersville, New York. 


COMPOSITAE. Vor. III. 


4. Achillea borealis Bongard. Northern Yar- 
row. Fig. 4555. 


Achillea borealis Bongard, Veg. Sitch. 149. 1831. 


More or less silky-woolly; stem erect, 16’ high or 
less. Leaves deeply bipinnatifid into narrow crowded 
lobes and segments, those of the stem few, sessiie 
or nearly so, the ultimate divisions very small; co- 
rymb dense, strongly convex, 23’ broad, or less; in- 
volucre about 3” high, its bracts with broad black or 
blackish margins; rays 10-20, white or pink, broadly 
oblong or suborbicular, often 23” broad. 


In wet places, on hillsides and rocks, Newfoundland 
to Quebec and Alaska. Sumimer. Rocky Mountain 
pias referred to this species appear to be distinct 
rom it. 


go. ANTHEMIS [Micheli] L. Sp. Pl. 893. 1753. 


Annual or perennial herbs, with pinnatifid or dissected, alternate leaves, and usually large 
peduncled heads of both tubular and radiate flowers, terminating the branches, or heads 
rarely rayless. Involucre hemispheric, its bracts imbricated in several series, scarious-margined, 
appressed, the outer shorter. Receptacle convex, conic or oblong, chaffy at least toward the 
summit, the chaff subtending the disk-flowers. Ray-flowers pistillate and fertile, or neutral, 
the tube terete or 2-winged, the ray white or yellow, entire or 2-3-toothed. Disk-flowers 
perfect, fertile, yellow, their corollas with 5-cleft limbs. Anthers obtuse and entire at the 
base. Style-branches of the disk-flowers truncate. Achenes oblong, angled, ribbed or striate. 
Pappus none, or a short coroniform border. [Greek name of Camomile. ] 

About 60 species, natives of Europe, Asia and Africa. Type species: Anthemis maritima L. 
Rays white. 


Rays neutral; plant glabrous, or nearly so, fetid. 1. A. Cotula. 
Rays pistillate ; plants pubescent. 
Annual ; chaff of the receptacle acute. 2. A. arvensis. 
Perennial ; chaff of the receptacle obtuse. 3. A. nobilis. 
Rays yellow; plant pubescent, or tomentose. 4. A. tinctoria. 


1. Anthemis Cétula L. Mayweed. Dog’s or Fetid Camomile. 
Dillweed. Fig. 4556. 


Anthemis Cotula L. Sp. Pl. 894. 1753. 
Maruta Co'ula DC. Prodr. 6: 13.- 1837. 


Annual, glabrous, or sometimes pubescent 
above, glandular and with a fetid odor and 
acrid taste, much branched, 1°-2° high. Leaves 
mostly sessile, 1-2’ long, finely 1-3-pinnately 
dissected into narrow, or almost filiform, acute 
lobes; heads commonly numerous, about 1’ 
broad; bracts of the involucre oblong, obtuse 
or obtusish, usually somewhat tomentose; rays 
10-18, white, at length reflexed, neutral, or 
rarely with abortive pistils, mostly 3-toothed; 
receptacle convex, becoming oblong, its chaff 
bristly, subtending the central flowers; achenes 
10-ribbed, rugose or glandular-tuberculate; 
pappus none. 

In fields, waste places and along roadsides, all 
over North America except the extreme north. 
Naturalized from Europe, and widely distributed 
as a weed in Asia, Africa and Australia. Other 
names are mather, dog- or hog’s-fennel, dog-finkle, 
morgan. Dog-daisy. Pig-sty-daisy. Maise. Chig- 
ger-weed. Balders. June—Nov. 


GENUS 90. THISTLE FAMILY. 517 


2. Anthemis arvénsis L. Corn or Field Camomile. Fig. 4557. 
Anthemis arvensis L. Sp. Pl. 894. 1753. > 


Annual or sometimes biennial, not fetid; stem 
finely pubescent, usually much branched, about 
1° high, the branches decumbent or ascending. 
Leaves sessile, 1-3’ long, I-2-pinnately parted 
into linear or lanceolate acute lobes, less divided 
than those of the preceding species and with 
broader segments; heads commonly numerous, 
1-12’ broad; bracts of the involucre oblong, 
obtuse, usually somewhat pubescent, with broad 
scarious margins; rays 10-18, white, pistillate, 
spreading, mostly 2-toothed; chaff of the obtuse 
receptacle lanceolate, acute or acuminate; achenes 


oblong, obtusely 4-angled; pappus a mere border. 
In fields and waste places, Nova Scotia to Virginia, 


west to Michigan, Missouri, and on the Pacific coast. 
Naturalized from Europe. May—Aug. 


3. Anthemis nobilis L. Garden, Scotch, 
White or Low Camomile. Fig. 4558. 


Anthemis nobilis L. Sp. Pl. 894. 1753. 


Perennial, pubescent, aromatic, much branched, 
6-18’ high, the branches procumbent. Leaves 
numerous, 1I’-2’ long, finely and compactly dis- 
sected into nearly filiform lobes and segments; 
heads about 1’ broad; bracts of the involucre ob- 
tuse, pubescent, their scarious margins broad; 
rays 12-18, white, spreading, pistillate, 2-3-toothed ; 
chaff of the conic receptacle broad, membranous, 
obtuse; achenes oblong, obtusely 3-angled; pappus 
none. 


Sparingly escaped from gardens, Rhode Island to 
Delaware, Michigan and Wyoming. Adventive from 
Europe. June—Aug. 


4. Anthemis tinctoéria L. Yellow or Ox- 
eye Camomile. Fig. 4559. 


Anthemis tinctoria L. Sp. Pl. 896. 1753. 


Perennial, pubescent or tomentose; stem erect, 
branched, 1°-3° high, with nearly erect branches. 
Leaves sessile, 1-3’ long, pinnately divided, the 
oblong segments pinnatifid into narrow acute 
lobes; heads few or several, 1-14’ broad; bracts 
of the involucre oblong, obtuse, densely tomen- 
tose; rays 20-30, pistillate, usually 2-toothed, 
bright yellow or sometimes paler; chaff of the‘ 
nearly hemispheric receptacle lanceolate, acumi- 
nate, rather rigid; achenes 4-angled, somewhat 
compressed; pappus a crown-like border. 

In fields and waste places, New Brunswick to New 


Jersey, and locally escaped from gardens. Adventive 
from Europe. Native also of Asia. June—Sept. 


Anthemis atirea (L.) DC., a species with small ray- 
less heads, was found many years ago near St. Louis, 
Missouri. 


518 COMPOSITAE. Vor. IIT. 


91. CHRYSANTHEMUM [Tourn.] L. Sp. Pl. 888. 1753. 


Perennial or annual, mostly erect and branching herbs, with alternate, dentate, incised 
or dissected leaves, and large, usually long-peduncled heads of both tubular and radiate 
flowers, or rays rarely wanting. Involucre hemispheric or depressed, its bracts appressed, 
imbricated in several series, the outer shorter. Receptacle flat, convex or hemispheric, naked. . 
Ray-flowers pistillate, fertile, the rays white, yellow or rose-colored, entire or toothed. Disk- 
flowers perfect, fertile, their corollas with terete or 2-winged tubes and 4-5-cleft limbs. 
Anthers obtuse and entire at the base. Style-branches of the disk-flowers truncate, penicil- 
late. Achenes angled or terete, 5-10-ribbed, those of the ray-flowers commonly 3-angled. 
Pappus none, or a scaly cup. [Greek, golden-flower. ] 

About 100 species, of wide geographic distribution in the northern hemisphere. Besides the 


following, 2 others occur in northwestern arctic America. Type species: Chrysanthemum coro- 
narium L. 


Heads large, few or solitary at the ends of the stem or branches. 
Rays white. 
Stem leaves linear-spatulate, pinnately incised ; weed. 
Stem leaves cuneate-spatulate, toothed or lobed above; arctic. 
Rays yellow. 
Heads numerous, small, corymbose; plants ecsaped from gardens, 
Leaves pinnatifid, the segments incised. 
Leaves oblong, serrate. 


. C. Leucanthemum. 
. C. arcticum, 
. C. segetum. 


. C. Parthenium. 
. C. Balsamita, 


on WNH 


1. Chrysanthemum Leucanthemum L. 
White-weed. White, Field or Ox-eye 
Daisy. Fig. 4560. 


C. Leucanthemum L. Sp. Pl. 888. 1753. 
Leucanthemum vulgare Lam. FI. Fr.2: 137. 1778. 


Perennial; stems glabrous, or sparingly pu- 
berulent, simple or little branched, 1°-3° high, 
often tufted, the branches nearly erect. Basal 
leaves obovate, oblong, or spatulate, coarsely 
dentate, incised or pinnatifid, narrowed into 
long slender petioles; stem leaves mostly sessile 
and partly clasping, 1-3’ long, linear-spatulate 
or linear, pinnately incised or toothed, the up- 
permost very small and nearly entire; heads 
few or solitary, 1-2’ broad, on long naked pe- 
duncles; rays 20-30, white, spreading, slightly 
2-3-toothed; bracts of the involucre oblong- 
lanceolate, obtuse, mostly glabrous, with scari- 
ous margins and a brown line within the mar- 
gins; pappus none. 

In pastures, meadows and waste places, com- 
mon throughout our area as a weed, but less 
abundant in the south and west. Bermuda. Na- 
turalized from Europe. Native also of Asia. 
Other English names are dog-, bull-, butter-, big-, 
midsummer-, moon-, horse-, poorland- or maudlin-daisy ; 
dutch morgan, moon-flower, moon-penny, great white ox- 
eye, poverty-weed, white man’s-weed, herb margaret ; bull’s- 
eye daisy. Sheriff-pink. Dog-blow. Rays rarely short and 
tubular. Disk bright yellow. May-—Nov. 


2. Chrysanthemum 4arcticum L. Arctic Daisy. 
Fig. 4561. 


Chrysanthemum arcticum L. Sp. Pl. 889. 1753. 
Leucanthemum arcticum DC. Prodr. 6: 45. 1837. 


Similar to the preceding species, but somewhat fleshy, 
lower, seldom over 13° high. Leaves cuneate-spatulate, 
13’-3’ long, crenate or cleft at the apex, narrowed into 
a long tapering entire base, or the lower into slender 
petioles, slightly clasping at the base, the uppermost 
few, small, linear and nearly entire; heads solitary or 
few, long-peduncled, 1’-2" broad; rays 20-30, white; 
bracts of the involucre oblong, obtuse, brown, or with 
broad brown scarious margins, usually pubescent; 
pappus none. 


Coast of Hudson Bay to Alaska. Also in arctic Europe 
and Asia. Summer. 


GENUS QI. THISTLE FAMILY. 519 


3. Chrysanthemum ségetum L. Yellow Ox- \ My IN 


eye. Corn Marigold. Fig. 4562. | \ () 
Chrysanthemum segetum L, Sp. Pl. 889. 1753. ve 
Annual, glabrous, 12° high or less. Leaves oblong Uf 
to oblanceolate, the upper auriculate-clasping, the pe if 


lower petioled, dentate, incised, or nearly entire, Be SS 
long or less; heads about 13’ broad; involucral bracts a Dy 
obtuse, scarious; rays obovate, yellow, emarginate ; Yi Vi} e 
pappus a mere margin. Y 

Waste grounds, New York, New Jersey, and in ballast | Wes VA 
about the seaports. Adventive from Europe. Va ae MI 

GED 
ue FY Bae 

Chrysanthemum coronarium L., also European, with | ] \\ 

yellow rays and bipinnatifid leaves, has been found in Ta (| 


Ontario. Z . ( 
-* 


4. Chrysanthemum Parthénium (L.) 
Pers. Common Feverfew. Feather- 
few. Fig. 4563. 


Matricaria Parthenium L. Sp. Pl. 890. 1753. 
C. Parthenium Pers. Syn. 2: 462. 1807. 


Perennial; stem puberulent or glabrate, much 
branched, 1°-23° high. Leaves thin, the lower 
often 6’ long, petioled, or the upper sessile, 
pinnately parted into ovate or oblong, pinna- 
tifid or incised segments; heads numerous, co- 
rymbose, slender-peduncled, 6’-10” broad; 
bracts of the depressed involucre lanceolate, 
rather rigid, keeled, pubescent, acute or acutish; 
rays 10-20, white, oval or obovate, spreading, 
mostly toothed, long-persistent; pappus a short 
toothed crown. 


In waste places, New Brunswick and Ontario to 
New Jersey, Ohio, and in California, mostly es- 
caped from gardens. Naturalized or adventive 
from Europe. Called also pellitory, wild camo- 
mile. Rays variable in length. Summer. 


ED 
RANE 
oS NY, {Zs 
YS 

4 


5. Chrysanthemum Balsamita L. Cost- 
mary. Mint Geranium. Fig. 4564. 


C. Balsamita L. Sp. Pl. Ed. 2, 1252. 1763. 
Pyrethrum Balsamita Willd. Sp. Pl. 3: 2153. 1804. 


Perennial, puberulent or canescent; stem 
much branched, 2°-4° high. Leaves oblong, 
obtuse, crenate-dentate, 1-2’ long, those of the 
stem mostly sessile, and often with a pair of 
lateral lobes at the base; heads numerous, co- 
rymbose, slender-peduncled, 5-8” broad, or 
when rayless only 3” broad; bracts of the in- 
volucre narrow, obtuse, pubescent; rays 10-15, 
white, spreading; pappus a short crown. 


Sparingly escaped from gardens, Ohio to Mich- 
igan, Ontario and Nova Scotia. Native of the Old 
World. Other English names are cost, alecost, ale- 
coast. Summer. 


520 COMPOSITAE. Vot. III. 


92. MATRICARIA L. Sp. Pl. 890. 1753. 


Annual or perennial, mostly erect herbs, similar to some species of the preceding genus, 
with alternate leaves, dissected into filiform or narrowly linear segments and lobes, and 
peduncled heads of both tubular and radiate flowers, or rays wanting in some species. Invo- 
lucre hemispheric, its bracts appressed, imbricated in few series, the outer shorter. Recep- 
tacle conic, elongated or hemispheric, naked. Rays, when present, white, pistillate and fertile. 
Disk-flowers yellow, perfect, fertile, their corollas 4—-5-toothed. Anthers obtuse and entire 
at the base. Style-branches of the disk-flowers truncate, penicillate. Achenes 3-5-ribbed. 
Pappus none, or a coroniform border. [Latin, matrix, from its medicinal virtues. | 

About 20 species, natives of the northern hemisphere and South Africa. The following are the 
only ones known to occur in North America. Type species: Matricaria inodora L. 

Rays present, white. 
Achenes obpyramidal, strongly 3-ribbed. 
Plant tall, much branched ; bracts of the involucre green. . M. inodora. 
Plant low, nearly simple, arctic; bracts dark brown or black. . grandiflora. 


x. 
2. M 
Achenes nearly terete, oblong, faintly 3—5-ribbed. 3. M. Chamomiilla. 
Rays none; achenes oblong, faintly nerved. 4. M. matricarioides. 


1. Matricaria inodora L. Scentless Camo- 
mile. Corn Mayweed. Fig. 4565. 


Matricaria inodora L. Fl. Suec. Ed. 2, 297. 1755. 


Chrysanthemum inodorum L. Sp. Pl. Ed. 2, 1253. 
1763. 


Annual; stem usually much branched, glabrous, 
or very nearly so throughout, 1°-2° high. Leaves 
numerous, sessile, 2-3-pinnately dissected into fili- 
form lobes, the rachis somewhat dilated at the 
base; heads several or numerous, terminating the 
branches, 2’-12’ broad; bracts of the involucre 
lanceolate-oblong, obtuse, green with brown scari- 
ous margins; rays 20-30, white, spreading; recep- 
tacle hemispheric or ovoid; achenes obpyramidal 
with three prominent ribs; pappus a short entire 
or 4-toothed crown. 


In fields and waste places, Newfoundland to New 
Jersey, Pennsylvania and Michigan. Naturalized or 
adventive from Europe. June—Sept. 


2. Matricaria grandiflora (Hook.) Britton. 
Arctic Camomile. Fig. 4566. 


Chrysanthemum grandiflorum Hock. in Parry’s 2d Voy. 398. 
1825. 


Pyrethrum inodorum var. nanum Hook. Fl. Bor. Am. 1: 
3205 11533: 


M. grandiflora Britton, Mem. Torr. Club 5: 340. 1894. 


Perennial; stem usually simple and monocephalous, 
glabrous, 4-12’ high. Leaves sessile, or the lowest 
short-petioled, 1I-—2-pinnately dissected, 1’-22’ long; 
head not very long-peduncled, 17-2’ broad; bracts of 
the involucre ovate or ovate-oblong, obtuse, glabrous, 
brown or nearly black, or with broad, brown, scarious 
margins; rays 15-35, bright white, slightly 3-5-toothed 
at the summit; receptacle hemispheric when mature. 


Coast of Hudson Bay to Alaska. Reported from Lake 
Huron. Summer. 


GENUS 92. THISTLE FAMILY. 521 


3. Matricaria Chamomilla L. Wild or 
German Camomile. Fig. 4567. 


Matricaria Chamomilla L. Sp. Pl. 891. 1753. 


Annual, glabrous, much branched, 1°-2° high. 
Leaves aromatic, finely 2-3-pinnately dissected 
into numerous linear lobes; heads numerous, 
8-12” broad, slender-peduncled at the ends of 
the branches; bracts of the involucre oblong, 
obtuse, green, or with brownish margins; rays 
10-20, white, spreading; receptacle ovoid, be- 
coming conic and hollow; achenes nearly oblong, 
or somewhat obovoid, faintly 3-5-ribbed; pappus 
none. 


In waste places and on ballast, southern New York 
to Pennsylvania. Adventive or fugitive from Eu- 
rope. Horse-gowan. Summer, 


4. Matricaria matricarioides (lLess.) 
Porter. Rayless Camomile.. Wild 
Marigold. 4568. 


Santolina suaveolens Pursh, Fl. Am. Sept. 520. 
1814. Not M. suaveolens L. 1755. 

Artemisia matricarioides Less. Linnaea 6: 210. 
1831. 

Matricaria discoidea DC. Prodr. 6: 50. 1837. 

Matricaria matricaricides Porter, Mem. Torr. 
Club 5: 341. 1894. 

M. suaveolens Buchenau, Fl. Nord. Tief. 496. 
1894. 


Annual, glabrous; stem very leafy, at length 
much branched, 6-18’ high. Leaves 2-3-pin- 
nately dissected into linear acute lobes; heads 
numerous, 3-4” broad, peduncled; bracts of 
the involucre oval or oblong, green, with 
broad white scarious margins, much shorter 
than the ovoid yellow disk; rays none; recep- 
tacle conic; achenes oblong, slightly angular, 
faintly nerved; pappus an obscure crown, 
sometimes produced into 2 coriaceous oblique 
auricles. 

In waste places, in ballast and along railroads, 
Missouri to Massachusetts and Maine. Adven- 
tive from the Pacific coast. Naturalized as a 
weed in northern Europe. May-—Aug. 


93. TANACETUM [Tourn.] L. Sp. Pl. 843. 1753. 


Erect, strongly aromatic herbs, our species perennials, with alternate, 1-3-pinnately dis- 
sected or divided leaves, and numerous small corymbose heads of tubular flowers, or with 
rays sometimes present and imperfectly developed. Involucre hemispheric, depressed, or cam- 
panulate, its bracts appressed, imbricated in several series. Receptacle flat or convex, naked. 
Marginal flowers pistillate, fertile, their corollas 2-5-toothed or lobed, sometimes produced 
into short rays. Disk-flowers perfect, fertile, their corollas 5-toothed. Anthers obtuse and 
entire at the base, their tips broad. Style-branches truncate and penicillate at the summit. 
Achenes 5-angled or 5-ribbed, truncate or obtuse. Pappus none, or a short crown. [From 
tanasie, old French for tansy; Greek, athanasia, immortality. ] 


: About 30 species, natives of the northern hemisphere. Besides the following, another occurs 
in California. Type species: Tanacetum vulgare L. 


Glabrous, or nearly so; heads numerous, 3”—5” broad. 1. T. vulgare. 
Villous-pubescent ; heads few, 6”—8” broad. 2. T. huronense. 


COMPOSITAE. Vor. III. 


1. Tanacetum vulgare L. Tansy. 


Fig. 4569. 
Tanacetum vulgare L. Sp. Pl. 844. 1753. ° 
| EI vulgare crispum DC, Prodr. 6: 128. 

1837. 

Stem stout, usually simple up to the inflo- 
rescence, glabrous, or sparingly pubescent, 
13°-3° high. Leaves pinnately divided into 
linear-oblong, pinnatifid or incised, often crisp- 
ed segments, the lobes acute, usually serrate; 
lower segments of the leaves often smaller 
than the others; basal leaves often 1° long; 
heads commonly numerous, 3-5” broad, 
rather short-peduncled; involucre depressed- 
hemispheric, its bracts oblong-lanceolate, ob- 
tuse, or the outer acute, slightly pubescent or 
ciliate; receptacle flat; flowers yellow; mar- 
ginal corollas with short oblique 3-toothed 
limbs; pappus a short crown. 

Along roadsides, mostly escaped from gardens, 
Nova Scotia and Ontario to Minnesota, Oregon, 
Nevada, North Carolina and Missouri. Natural- 
ized or adventive from Europe. Bitter-buttons. 
Hindheal. Ginger-plant. July—Sept. 


2. Tanacetum huronénse Nutt. Lake 
Huron Tansy. Fig. 4570. 

Tanacetum huronense Nutt. Gen. 2: 141. 1818. 

Villous-pubescent throughout, at least when 
young, less so when mature, 1°-2° high. Leaves 
2-pinnately divided, the lobes dentate or incised, 
acute, the lower segments commonly smaller than 
the others; heads 1-8, 6-8” broad, on very stout 
pubescent peduncles; involucre depressed-hemi- 
spheric; marginal flowers with 3-5-lobed limbs, 
often expanded into short rays; pappus a short 
crown. 

In moist soil, especially along streams or lakes, 


New Brunswick to Hudson Bay, Maine, Lake Su- 
perior, Alaska and Oregon. July—Sept. 


94. ARTEMISIA [Tourn.] L. Sp. Pl. 
845. 1753. 


Odorous herbs or shrubs, often canescent or 
tomentose, with alternate leaves, and small pendu- 
lous or erect, discoid racemose spicate glomerate or paniculate heads of greenish or yellowish 
flowers. Involucre ovoid, oblong, or broadly hemispheric, its bracts imbricated in few 
series, the outer gradually shorter. Receptacle flat, convex or hemispheric, naked or pubes- 
cent, not chaffy. Central flowers perfect, sometimes sterile, with abortive ovaries and undi- 
vided style, sometimes perfect and fertile, with truncate style-branches; marginal flowers 
usually pistillate and fertile, their corollas 2~-3-toothed, or flowers all perfect and fertile in 
some species. Anthers obtuse and entire at the base, often tipped with subulate appendages. 
Achenes obovoid or oblong, 2-ribbed or striate, rounded at the summit, usually bearing a 
large epigynous disk. Pappus none. [Named for Artemisia, wife of Mausolus.] 

About 225 species, natives of the northern hemisphere and southern South America. Besides 
the following, some 40 others occur in the western parts of North America. Type species: Arte- 
misia vulgaris L, 

* Marginal flowers pistillate; central flowers perfect, sterile. 
a. Biennial or perennial herbs. 


Leaves pinnately dissected into narrowly linear lobes. 
Heads very numerous, 1” broad; leaves mostly glabrous. 1. A. caudata. 
Heads 2” broad, in narrow panicles; leaves silky-pubescent. 
Heads few; involucre brown, mostly pubescent. 2 
Heads numerous; involucre green, mostly glabrous. 3 
Leaves linear, the lower sometimes 3-cleft or pinnately divided. , 
Leaves glabrous. 4. A. dracunculoides. 
5 
6. 


Why 


Jk 
ea 
oe GD 
frtides G 


. A. borealis. 
. A. canadensis. 


Leaves finely and densely pubescent. . A. glauca. 
b. Shrubby, silvery-canescent ; heads small and numerous. A, filifolia. 


GENUS 94. THISTLE FAMILY. 523 


** Marginal flowers pistillate; central flowers perfect, fertile. 
a. Receptacle villous-pubescent. 
Leaf-segments linear-filiform, short; native. 7. A. frigida. 
Leaf-segments oblong, or linear-oblong ; introduced. . A. Absinthium. 
b. Receptacle glabrous, or sparingly pubescent. 
Leaves dissected, glabrous or pubescent, green, not tomentose. 


Heads about 2” broad, numerous in panicled racemes; perennial. 9. A. Abrotanum. 
Heads about 1” broad, paniculate or spicate; annuals. 
Leaves finely 2—3-pinnately divided ; heads paniculate. 10. A. annua. 
Leaves pinnately divided ; segments pinnatifid ; heads in leafy spikes. 11. A. biennis. 


Leaves densely white-canescent or tomentose, at least beneath. 
Leaves pinnatifid or dissected. 


Heads 3-4” broad, racemose-glomerate ; sea-beach plant. 12. A. Stellariana. 
Heads 1”-2” broad, spicate-paniculate or racemose. 
Leaves deeply pinnatifid, the segments mostly incised. 13. A. vulgaris. 
Leaves finely dissected into short linear lobes. 14. A. Pontica. 
Leaves pinnately parted into 5-7 narrow entire segments. 15. A. kansana. 
Leaves lanceolate or linear, serrate or entire, not pinnatifid. 
Leaves lanceolate, sharply serrate, glabrous above. 16. A. serrata. 
Leaves linear, oblong, lanceolate or obovate, entire or lobed. 
Leaves at length glabrous above. 
Leaves linear, elongated, all entire. 17. A. longifolia. 
Leaves various, at least the lower pinnately lobed or toothed. 
Involucre densely woolly ; leaf-lobes broad. 18. A. ludoviciana. 
Involucre loosely woolly ; leaf-lobes linear. 19. A. mexicana. 
Leaves shorter, oblong or lanceolate, tomentose both sides. 20. A. gnaphalodes. 
Leaves cuneate, %’ long, 3-toothed at the apex. 21. A. Bigelovii. 
*** Flowers all perfect and fertile; far western species. 
Leaves cuneate, 3-toothed or 3-lobed. 22. A. tridentata. 
Leaves linear, entire. 23. A. cana. 


1. Artemisia caudata Michx. Tall or Wild 
Wormwood. Fig. 4571. 
Artemisia caudata Michx. Fl. Bor. Am, 2: 129. 1803. 


Root biennial (sometimes perennial?) ; stems slender, 
glabrous, tufted, strict, very leafy, 2°-6° high, at length 
paniculately branched, the branches glabrous, or rarely 
slightly pubescent, nearly erect. Lower and basal leaves 
and those of sterile shoots slender-petioled, sometimes 
a little pubescent, 3-6’ long, 2-3-pinnately divided into 
narrowly linear, acute lobes, about 4” wide; upper leaves 
sessile or nearly so, pinnately divided, or the uppermost 
entire and short; heads about 1” broad, very short- 
peduncled, very numerous in a large somewhat leafy 
panicle, mostly nodding; bracts of the ovoid-campanu- 
late involucre ovate, or the inner elliptic, glabrous; re- 
ceptacle hemispheric, naked; central flowers sterile. 

In dry sandy soil, abundant on sea-beaches, from Quebec 
to Florida, west to Ontario, Indiana, Manitoba, south to 
Nebraska and Texas. July—Sept. 


Vf 


2. Artemisia borealis Pall. Northern Worm- 
wood. Fig. 4572. 

Artemisia borealis Pall. Iter. 129. pl. hh, f. 2. 1771. 

Artemisia groenlandica Wormsk. Fl. Dan. pl. 7585. 1818. 


Perennial, 5’-15’ high, densely silky-pubescent all 
over, resembling small forms of the following spe- 
cies. Leaves less divided, the basal and lower ones 
petioled, 1’-23’ long, the upper sessile, linear and 
entire or merely 3-parted; heads about 2” broad in 
a dense terminal rarely branched thyrsus; involucre 
nearly hemispheric, its bracts brown or brownish, 
pilose-pubescent or nearly glabrous; receptacle con- 
vex, naked; disk-flowers sterile. 


Quebec to Greenland, west through arctic America to 
Alaska, south in the Rocky Mountains to Colorado. Also 
in northern Asia. Apparently erroneously recorded from 
Maine. July—Aug. 


524 COMPOSITAE. Vor. IIT. 


3. Artemisia canadénsis Michx. Canada Worm- 
wood. Fig. 4573. 


Artemisia canadensis Michx. Fl. Bor. Am. 2: 129. 1803. 


Root perennial (or sometimes biennial); stem pubes- 
cent or glabrous, strict, simple or branched, 1°-2° high, 
the branches appressed and erect. Leaves usually pubes- 
cent, but sometimes sparingly so, the basal and lower ones 
petioled, 2’-3’ long, 2-pinnately divided into linear, acute 
lobes which are shorter and broader than those of Arte- 
misia caudata; upper leaves sessile, less divided; heads 
short-peduncled, about 2” broad, commonly numerous in 
a narrow virgate panicle, mostly spreading or erect, in 
small forms the panicle reduced to a nearly or quite sim- 
ple terminal raceme; involucre ovoid, its bracts ovate or 
oval, green, glabrous or pubescent; receptacle hemispheric; 
central flowers sterile. 


In rocky soil, Newfoundland to Hudson Bay, Maine, Ver- 
mont, west along the Great Lakes to Minnesota and Manitoba 
and to the Canadian Pacific coast. Sea- or wild-wormwood. 
July—Aug. 

Artemisia Forwoddii S. Wats., a taller plant of the Rocky 
Mountain region, with somewhat smaller heads, ranges east- 
ward into Nebraska. 


4. Artemisia dracunculoides Pursh. Linear- 
leaved Wormwood. Fig. 4574. 


A, dracunculoides Pursh, Fl. Am. Sept. 742. 1814. 


Perennial, glabrous; stem somewhat woody, usu- 
ally much branched, 2°-4° high, the branches nearly 
erect. Leaves linear, 1-33’ long, 1-2” wide, acute, 
entire, or the lower and basal ones sometimes 3-cleft 
or even more divided; heads very numerous, 1”—13” 
broad, nodding, very short-peduncled, racemose- 
paniculate; involucre nearly hemispheric, its bracts 
ovate or oblong, green, scarious-margined; recep- 
tacle hemispheric, naked; central flowers sterile. 


Dry plains and prairies, Manitoba to British Colum- 
bia, Illinois, Missouri, Nebraska, Texas, Chihuahua, 
New Mexico and California. July—Nov. 


wood. Fig. 4575. 


A. glauca Pall.; Willd. Sp .Pl. 3: 1831. 1804. 
Ariemisia dracuncuioides var. incana T. & G. Fl. N. 
A. 2: 416. 1843. 


ee N 

U tee . 

ey \ aR \ 
we “ay j [ 4) 


U dt 2 
\ Nid 
AG® 


i 
Wes 


Perennial, similar to the preceding species; 
stems strict, leafy, usually simple or little branch- 
ed, 1°-2° high, pubescent, tomentose or canescent, 
or glabrous below. Leaves linear, 4’-23’ long, 
about 1” wide, entire, finely and densely pubes- 
cent, obtuse or obtusish, or the lower or some- 
times nearly all of them 3-cleft into linear lobes, 
1-13’ long; panicle narrow, branched, its branches 
nearly erect; heads drooping, sessile, very numer- 
ous, scarcely more than‘13” long; involucre hemi- 
spheric, its bracts scarious-margined, obtuse; re- 
3 ceptacle naked; central flowers sterile. 


Minnesota to North Dakota, Manitoba and Sas- 
katchewan. June—Sept. 


E i 
: wn 
rf ee 


GENUS 94. THISTLE FAMILY. 


6. Artemisia filifélia Torr. Silvery Worm- 
wood. Fig. 4576. 


Artemisia filifolia Torr. Ann. Lye. N. Y. 2: 211. 1827. 


Shrubby, finely silvery-canescent throughout; stem 
branched, 1°-3° high, the rigid branches nearly erect. 
Leaves 1’-2’ long, nearly all 3-parted into filiform 
entire segments less than 3” wide, or the uppermost 
undivided; heads exceedingly numerous, about 2” 
broad, racemose-paniculate, very short-peduncled, 
3-5-flowered; involucre oblong, its bracts densely 
canescent; receptacle small, naked or slightly fim- 
brillate; central 1-3 flowers sterile. 


On dry plains, Nebraska to Utah, Wyoming, Nevada, 
Texas, Mexico and Arizona. Wormwood-sage. July- 
Oct. 


7. Artemisia frigida Willd. Pasture Sage- 
Brush. Wormwood Sage. Fig. 4577. 

Artemisia frigida Willd. Sp. Pl. 3: 1838. 1804. 

Perennial, woody at the base, densely silky-canes- 
cent all over; stem branched or simple, 10-20’ high. 
Leaves 37-13’ long, ternately or 5-nately divided 
into numerous short acutish mostly entire lobes 
less than 4” wide, the lower and basal ones petioied, 
and often with a pair of entire or 3-cleft divisions 
near the base of the petiole, the upper sessile and 
less divided; heads rather numerous, racemose or 
racemose-paniculate, short-peduncled, nodding, about 
2” broad; involucre hemispheric, its bracts oblong, 
canescent or tomentose; receptacle villous-pubes- 
cent; central flowers fertile. 


On dry plains and in rocky soil, Minnesota to Sas- 
katchewan, Yukon, Idaho, Nebraska, Texas and Ari- 
zona. Wild sage. July—Oct. 


8. Artemisia Absinthium L. Common Worm- 
wood. Absinth. Fig. 4578. 


Artemisia Absinthium L. Sp. Pl. 848. 1753. 


Shrubby, finely canescent; stem much branched, 
2°-4° high. Leaves 2’-5’ long, I-3-pinnately divided 
into numerous linear to obovate, obtuse lobes, the 
lower long-petioled, the upper short-petioled or ses- 
sile, the uppermost commonly linear and entire; heads 
numerous, yellow, racemose-paniculate, drooping, 
short-peduncled, 2’—23” broad; involucre hemispheric, 
its outer bracts linear, the inner much broader, scari- 
ous-margined; receptacle pilose-pubescent; central 
flowers fertile, the marginal ones pistillate, fertile or 
sterile. 

In waste places, Newfotindland and Hudson Bay to 
Massachusetts, Pennsylvania, North Carolina, western 
Ontario, New York, North Dakota and Montana. Natu- 
ralized or adventive from Europe, mostly escaped from 


gardens. Old English names, madderwort, mugwort, 
mingwort, warmot. Boys’-love. July—Oct. 


526 COMPOSITAE. Vor. ITI. 


g. Artemisia Abrétanum L. Southernwood. Fig. 4579. 
Artemisia Abrotanum L. Sp. Pl. 845. 1753. 


Perennial, somewhat shrubby; stem puberulent or 
glabrous, much branched, 2°-4° high, the branches 
short, erect or ascending. Leaves glabrous or some- 
what pubescent, 1-3’ long, I-3-pinnately parted into 
linear obtuse entire lobes about 2” wide, or the upper- 
most linear and entire, the lowest petioled; heads sey- 
eral-flowered, yellow, very numerous, nodding, race- 
mose-paniculate, 2’—24” broad; involucre nearly hemi- 
spheric, pubescent, its outer bracts lanceolate, acute, 
the inner ones obovate; receptacle glabrous; central 
flowers fertile. 


In waste places, Massachusetts to western New York, 
southern Ontario, and Nebraska. Adventive from con- 
tinental Europe. Old English names, lad’s-love, boys’-love, 
slovenwood, old-man, sweet benjamin. 


Artemisia procera Willd., a similar species, but with 
glabrous involucre, is recorded as escaped from gardens at 
Buffalo, N. Y. 


; w 
10. Artemisia A4nnua L. Annual Wormwood. 
Fig. 4580. 
Artemisia annua L. Sp. Pl. 847. 1753. 


Annual, glabrous throughout, much branched, 2°-5° 
high. Leaves 2’-6’ long, finely 2+3-pinnately dissected 
into very narrow short, obtuse lobes, the lower and 
basal ones slender-petioled, the upper sessile and less 
divided, but none of them entire; heads very nu- 
merous, about 1” broad, drooping, borne on very 
slender peduncles of about their own length.or less; 
involucre hemispheric, glabrous, its bracts few, ovate 
to oblong; receptacle glabrous; flowers commonly all 
fertile. 


In waste places, Ontario to New Hampshire, Virginia, 
West Virginia, Tennessee, Kansas and Arkansas, a bad 
weed in some places. Adventive or naturalized from 
Asia. Summer. 


11. Artemisia biénnis Willd. Biennial Worm- 
wood. Fig. 4581. 


Artemisia biennis Willd. Phytogr. 11. 1794. 


Annual or biennial, glabrous throughout; stem very 
leafy, usually branched, 1°-4° high, the branches nearly 
erect. Leaves 1’-3’ long, I-2-pinnately divided into 
linear or linear-oblong, acutish, serrate or incised 
lobes, the lowest petioled, the uppermost less divided 
or rarely quite entire; heads about 13” broad, not 
drooping, sessile and exceedingly numerous in axillary 
glomerules which are crowded, forming a compound 
spicate inflorescence, the subtending leaves much ex- 
ceeding the clusters; involucre nearly hemispheric, its 
bracts green, scarious-margined; receptacle naked; 
) central flowers fertile. 


Native from Tennessee to Nebraska, Manitoba, British 
Columbia and California, now widely distributed as a weed 
from Manitoba to Nova Scotia, south to Missouri, Ken- 
tucky and Delaware. Aug.—Oct. 


GENUS 94. THISTLE FAMILY. 527 


12. Artemisia Stelleriana Bess. Beach 
Wormwood. Fig. 4582. 


Artemisia Stelleriana Bess. Abrot. 79. pl. 5. 1829. 


Perennial, densely white-tomentose ; stem branch- 
ed, 1°-23° high, bushy, the branches ascending. 
Leaves obovate to spatulate, 1-4’ long, pinnatifid 
into oblong, obtuse, entire or few-toothed lobes, 
the lower petioled, the upper sessile, all densely 
tomentose beneath, but becoming green and gla- 
brous above when old; heads racemose-spicate or 
racemose-glomerate, 3-4” broad, not drooping; 
involucre oblong-campanulate, its bracts tomen- 
tose, lanceolate or oblong-lanceolate; receptacle 
naked; central flowers fertile. 


Sandy sea-beaches, Quebec to New Jersey; Oneida 
Lake, N. Y. Cultivated in gardens along the coast. 
Native of northeastern Asia. Occurs also on the 
coast of Sweden. Foliage similar to that of the dusty 
miller, Cineraria maritima L. July—Aug. 


13. Artemisia vulgaris L. Common Mug- 
wort. Fig. 4583. 


Artemisia vulgaris L. Sp. Pl. 848. 1753. 


Perennial; stem glabrous or nearly so, much 
branched, 1°-32° high. Leaves 1-43’ long, deeply 
pinnatifid, into linear, oblong or somewhat spatu- 
late, pinnatifid, toothed or entire lobes, densely 
white-tomentose beneath, dark green and glabrous 


a 


AN 
EY oR 
5S Wy Te 


SS above, the lower petioled and often with I or 2 
EWon|| pairs of small lateral divisions at or near the 
AWN Vi \ y base of the petiole, the upper sessile, the upper- 


most sometimes linear and entire; heads numer- 
ous, erect, about 2” broad, in panicled, simple or 
compound epikes: involucre oblong-campanulate, 
its bracts oblong, obtusish, scarious-margined, 
tomentose or glabrous; receptacle naked; central 
flowers fertile. 


In waste places, Nova Scotia to Ontario, Michigan, 
New Jersey, Pennsylvania and Georgia. Naturalized 
from Europe. Native also of Asia. Reported as 
native of arctic America. Motherwort. Fellon-herb. Sailor’s-tobacco. Wormwood. Bulwand. 
Green ginger. July—Oct. 

Artemisia elatidr (T. & G.) Rydb., a northwestern 
species, with elongated acuminate leaf-lobes, ranges 
eastward to Hudson Bay. 


14. Artemisia pontica L. Roman or Hun- 
garian Wormwood. Fig. 4584. 


Artemisia pontica L. Sp. Pl. 847. 1753. 


Perennial; stem branched, glabrous or canescent, 
1°-3° high. Leaves 14’-23’ long, 2-3-pinnately dis- 
sected into short narrow lobes less than 1” wide, 
canescent on both sides, or tomentose beneath, the 
lower petioled and the petioles somewhat clasping 
or auricled at the base, the upper mostly linear and 
entire; heads numerous, 1-2” broad, drooping, 
slender-peduncled; involucre hemispheric, canes- 
cent, its bracts oblong or obovate, obtuse, the outer 
short, lanceolate; receptacle glabrous; central flow- 
ers fertile. 


Waste grounds, Massachusetts, New Jersey, Pennsyl- 
vania, Ohio and Colorado. Fugitive or adventive from 
central Europe. July—Aug. 


COMPOSITAE. Vo. III. 


15. Artemisia kansana Britton. Kansas Mug- 
wort. Fig. 4585. 


eg Corns A. Wood, Trans. Kans. Acad. Sci. 5: 51. 
1876 


A. kansana Britton, in Britt. & Brown, Ill. Fl. 3: 466. 1808. 


Densely white-woolly all over; stem erect, much 
branched, the branches strict, bearing very numerous 
small heads forming a narrow dense panicle. Leaves 
numerous, crowded, the lower pinnately divided into 
3-7 narrowly linear revolute-margined segments 3” 
wide or less, greenish above; upper leaves mostly nar- 
rowly linear and entire; heads oblong-oval, sessile, or 
very short-peduncled, erect, or somewhat spreading, 
14” long; involucre very woolly, its bracts ovate-lan- 
3 ceolate to oblong-lanceolate, acute; receptacle naked. 
4 Plains, Kansas to Colorado and New Mexico. Intro- 
duced in Missouri. July—Seot. 


16. Artemisia serrata Nutt. Saw-leaf 
Mugwort. Fig. 4586. 
Artemisia serraia Nutt. Gen. 2: 142. 1818. 


Perennial; stem stout, tomentose or be- 
coming glabrous, much branched, 5°-10° high. 
Leaves lanceolate, 2’-6' long, 3-12’ wide, 
densely white-tomentose beneath, dark green 
and glabrous above, acuminate at the apex, 
narrowed to a sessile base, or the lowest peti- 
oled, sharply serrate or incised, or the upper 
entire; heads very numerous, greenish, erect, 
about 13” broad, sessile or short-peduncled 
in panicled spikes or racemes; involucre ca- 
nescent, its bracts oblong, or the outer ones 
lanceolate; receptacle naked; central flowers 
fertile. 


ES 


| Wy 


Prairies, Illinois to Minnesota and Dakota. 
Introduced on the Mohawk River, near Sche- 


17. Artemisia longifolia Nutt. Long- 
leaved Mugwort. Fig. 4587. 


Artemisia longifolia Nutt. Gen. 2: 142. 1818. 


Perennial; stem densely white-tomentose, 
branched, 2°-5° high. Leaves linear or linear- 
lanceolate, elongated, entire, 2’-5’ long, 17-5” 
wide, acuminate, tapering to a sessile base, or 
the lower petioled, densely white-tomentose 
on both sides, or becoming green and glabrate 
above; heads numerous, erect, spicate-panicu- 
late, about 2” broad; involucre tomentose, its 
bracts narrowly oblong; receptacle naked; 
central flowers fertile. 


In dry rocky soil, western Nebraska to Minne- 
ae Idaho, Oregon and Saskatchewan. Aug.- 
ept. 


GENUS 94. THISTLE FAMILY. 529 


18. Artemisia ludoviciana Nutt. Dark- 
leaved Mugwort. Fig. 4588. 


Artemisia lndoviciana Nutt. Gen. 2: 143. 1818. 


Perennial, 2°-4° high; stem woolly, branched 
above. Leaves linear to obovate, 3’ long or less, 
white-woolly beneath, at length dark green and 
glabrous, or very nearly so above, the base com- 
monly narrowly cuneate, at least the lower ones 
pinnately lobed or toothed, their lobes lanceolate, 
acuminate; upper leaves often linear and entire; 
heads numerous, spicate-paniculate, 1-12” broad; 
involucre oblong, tomentose; receptacle naked; 
central flowers fertile. 


In dry soil, Missouri to Texas, Wyoming, Colorado 
and Arizona. Aug.—Nov. 


1g. Artemisia mexicana Willd. Mexican Mug- 
wort. Fig. 4580. 


Artemisia mexicana Willd.; Spreng. Syst. 3: 490. 1826. 


Perennial, less densely pubescent than A. ludoviciana, 
2°-3%° high, often branched; stem finely pubescent and 
ultimately often floccose. Leaves ovate or orbicular in 
outline on the lower part of the stem, 2’-33’ long, 
densely white-tomentulose beneath, green above, the 
lobes of the lower and the blades of the upper entire 
ones linear to narrowly linear or nearly so; heads small 
and numerous, usually inclined or nodding; involucre 
campanulate, loosely woolly, the pubescence sparse. 


On prairies, hillsides and barrens, Missouri to Texas, 
Arkansas and Mexico. Sept.—Oct. 


20. Artemisia gnaphalddes Nutt. Prairie 
or Western Sage. Cud-weed 
Mugwort. Fig. 4590. 

Artemisia gnaphalodes Nutt. Gen. 2: 143. 1818. 


Artemisia ludoviciana var. gnaphalodes T. & G. FI. 
N. A. 2: 420. 1843. 


—aae 
WS 


HP 3 
Ve 


S 


<i 


Perennial; stem white-tomentose, usually 
much branched, 1°-4° high. Leaves lanceolate 
or oblong, 1’-3’ long, 2”-6” wide, entire, or 
the lower somewhat toothed, or rarely few- 
lobed, white-tomentose on both sides, acute or 
acuminate, sessile or the lower narrowed into 
short petioles ; heads numerous, spicate-panicu- 
late, about 14” broad; involucre oblong, to- 
mentose; receptacle naked; central flowers 
fertile. 


On prairies, plains, and dry banks, western On- 
tario and Illinois to Alberta, Missouri, Texas and 
Mexico. Locally established in waste grounds 
from New Hampshire to Delaware. Far western 
plants formerly referred to this species, which 
consists of many races, are, apparently, distinct. 


34 


COMPOSITAE. Vor, Ti 


21. Artemisia Bigelovii A. Gray. Bigelow’s 
Sage-Bush. Fig. 4591. . 
gape Bigelovii A. Gray, Pac. R. R. Rep. 4: 110. 

1856. 

Perennial, shrubby, silvery-canescent throughout, 
8-15’ high, much branched, the branches erect. Leaves 
narrowly cuneate, or oblong, obtuse, truncate, or 
3-5-toothed at the apex, 5-0” long, about 1” wide; 
heads very numerous, about 1” broad, densely glom- 
erate-spicate in a narrow virgate panicle, 2-5-flow- 
ered, I or 2 of the marginal ones pistillate, the others 
perfect and fertile; involucre short-oblong, canescent 
or tomentose, its bracts obtuse; receptacle naked. 


Kansas (according te Smyth) ; Colorado to Texas and 
Arizona. Aug.—Oct. 


22. Artemisia tridentata Nutt. Common 
Sage-bush. Sage-brush. Sage-wood. 
Mountain Sage. Fig. 4592. 
Artemisia tridentata Nutt. Trans. Am. Phil. Soc. 

(11) 7: 398. 1841. 

Shrubby, silvery-canescent ; stem much branch- 
ed, 1°-12° high. Leaves-narrowly cuneate, 3”— 
12’ long, 1-3” wide, sessile, 3-7-toothed at the 
truncate apex; heads very numerous, 5-8-flow- 
ered, about 14” broad, sessile, or very nearly so, 
in large dense panicles; involucre oblong, to- 
mentose, its inner bracts oblong, the outer short, 
ovate, all obtuse or obtusish; receptacle naked; 
flowers all perfect and fertile. 

On dry plains and in rocky soil, western Ne- 
braska to Colorado, Utah and California, north to 
Montana and British Columbia. July—Sept. 


23. Artemisia cana Pursh. Hoary Sage- 
bush. Fig. 4593. 


Artemisia cana Pursh, Fl. Am. Sept. 521. 1814. 


Shrubby, densely white-canescent ; stem much 
branched, 1°-2%° high. Leaves linear, linear- 
oblong or narrowly lanceolate, sessile, acute » 
at both ends, 1-2’ long, 14’-3” wide, usually 
quite entire, rarely with 2 or 3 acute teeth or 
lobes; heads numerous, about 14” broad, glom- 
erate or sometimes solitary in the axils of 
the leaves, or crowded into a naked thyrsus 
at the summit, 5-9-flowered; involucre ob- 
long, canescent, its inner bracts oblong or lan- 
i ceolate, obtuse, usually with 1-3 shorter outer 
| ones; receptacle naked; flowers all perfect and 
fertile. 


Plains, Nebraska and Colorado to North Da-— 
kota, Montana and Saskatchewan. July—Sept. 


95. TUSSILAGO [Tourn.] L. Sp. Pl. 865. 1753. 


An acaulescent herb, more or less white-tomentose, with slender perennial rootstocks, 
broad basal cordate, dentate or lobed, long-petioled leaves, and large solitary, monoecious 


GENUS 95. . THISTLE FAMILY. 531 


heads of both tubular and radiate yellow flowers at the summit of a scaly scape, appearing 
before the leaves of the season. Involucre campanulate to cylindric, its principal bracts in 
a single series, equal, with or without a few shorter outer ones. Receptacle flat, naked. 
Ray-flowers in several series, pistillate, fertile. Disk-flowers perfect, sterile, the corolla 
5-cleft, the style undivided and obtuse, lobed. Achenes of the ray-flowers linear, 5—10-ribbed. 
Pappus copious, of numerous slender roughish a a 
bristles, that of the sterile flowers shorter than 2 ZR 
that of the fertile. [Latin, tussis, cough, for % RW 
which the plant was a reputed remedy. | 3 

A monotypic genus of northern Europe and Asia. 


7 


CP iaete, 
peat 


ie 
ier 
War iet 


1. Tussilago Farfara L. Coltsfoot. 
Coughwort. Fig. 4594. 
Tussilago Farfara L. Sp. Pl. 865. 1753. 


Scape slender, 3-18’ high, bearing a solitary 
large head at the summit. Leaves nearly orbicu- 
lar, or broadly ovate-reniform, angulately lobed 
and dentate, 3-7’ broad, green and glabrous above, 
persistently white-tomentose beneath; head about 
1’ broad; involucre campanulate; rays bright yel- 
low, numerous, linear. 

In moist soil, on banks and roadsides, Nova Scotia 
and New Brunswick to New Jersey, Pennsylvania 
and Minnesota. Naturalized from Europe. Horse- 
foot. Horse-hoof. Dove-dock. Sow-foot. Colt-herb. 
Hoofs. Cleats. Ass’s-foot. Bull’s-foot. Foal-foot. 
a Clay-weed. Butter-bur. Dummy-weed. April-— 
une. 


96. PETASITES [Tourn.] Mill. Gard. Dict. Abr. Ed. 4. 1754. 


Herbs with perennial thick horizontal rootstocks, broad, basal, petioled leaves, and scaly 
scapes bearing racemose or corymbose heads of tubular or both tubular and radiate, white 
or purplish, often dioecious or subdioecious flowers. Involucre campanulate to cylindric, 
its bracts in I series, equal. Receptacle flat, or nearly so, not chaffy. Corolla of pistillate 
flowers very slender, 2-5-toothed, truncate or sometimes with a ray, marginal, or composing 
most of the head; perfect but sterile flowers with a tubular 5-cleft corolla, the style undi- 
vided. Anthers entire or minutely sagittate at the base. Fertile achenes linear, the pappus 
of numerous capillary rough or barbellate bristles. [Greek, a broad-rimmed hat, referring 
to the broad leaves of these plants. ] 


About 20 species, north temperate and subarctic. Type species: Tussilago Petasites L. 


Flowers whitish, the pistillate radiate; natives; northern. 
Leaves orbicular, 7—11-cleft nearly to the base. 1. P. palmata. 
Leaves deltoid-reniform, sinuate-lobed and toothed. 2. P. trigonophylla. 
Leaves deltoid-ovate, repand-denticulate. 3. P. sagittata. 
Flowers all rayless, purple; introduced. 4. P. Petasites. 


1. Petasites palmata (Ait.) A. Gray. Pal- 
mate-leaf Sweet Coltsfoot. Fig. 4595. 


Tussilago palmata Ait. Hort. Kew. 2: 188. pl. 2. 1780. 
Nardosmia palmata Hook. Fl. Bor. Am. 1: 308. 1833. 
P.palmata A. Gray, in Brew. & Wats. Bot. Cal.1: 407. 1876. 


Scape very scaly, stout, 6-24’ high. Leaves nearly 
orbicular in outline, 3-12’ broad, deeply 7-11-cleft 
to much beyond the middle, green and glabrous 
above, densely white-tomentose beneath, at least 
when young, sometimes becoming glabrate, the lobes 
oblong or obovate, acute, often somewhat cuneate, 
sharply dentate or incised; heads mostly dioecious, 
corymbose or racemose-corymbose, numerous, 4-6” 
broad; flowers nearly white, fragrant, the marginal 
ones of the pistillate heads radiate. 


In swamps and along streams, Newfoundland to Mas- 
sachusetts, New York, Wisconsin, Minnesota and Alberta. 
Far western plants, formerly included in this species, 
prove to be distinct. April-June. Butter-bur. 


532 _ COMPOSITAE. Vor. III. 


2. Petasites trigonophylla Greene. 
Arctic Sweet Coltsfoot. Fig. 4596. 


BERS trigonophylla Greene, Leaflets 1: 180. 
~ 1906. 


Scape very scaly, 3’-10’ high. Leaves 
deltoid-reniform to ovate-orbicular in out- 
line, 2-6’ long, irregularly lobed, green and 
glabrous above, persistently white-tomen- 
tose beneath, the lobes few-toothed; heads 
corymbose, the inflorescence about 4’ long; 
involucre campanulate; flowers nearly 
white, the marginal ones of the pistillate 
heads radiate. 


Wet grounds, Quebec, Minnesota and Sas- 
katchewan. June—Aug. 


Petasites frigida (L.) Fries, admitted, in 
our first edition, as recorded from Lake Win- 
nipeg, is a high boreal species, not known to 
occur within our area. 


3. Petasites sagittata (Pursh) A. Gray. 
Arrow-leaf Sweet Coltsfoot. Bitter- 


bur. Fig. 4597. 


Tussilago sagittata Pursh, Fl. Am. Sept. 332. 1814. 

Nardosmia sagittata Hook. Fl. Bor. Am.1: 307. 1833. 

Petasites sagittata A. Gray, in Brew. & Wats. Bot. 
Cal: 2407, 21076. 


Scape and racemose-corymbose inflorescence 
similar to those of the two preceding species. 
Leaves deltoid-ovate to reniform-ovate, persist- 
ently white-tomentose beneath, glabrous or nearly 
so above, 4’-10’ long, their margins sinuate-den- 
ticulate, neither cleft nor lobed; involucre cam- 
panulate; flowers nearly white, the marginal ones 
of the pistillate heads radiate. 

In wet grounds, Labrador to Hudson Bay, Manitoba 


and Minnesota, west to British Columbia, south in the 
Rocky Mountains to Colorado. May-June. 


4. Petasites Petasites (L.) Karst. | 
Butter-bur. Butterfly-dock. 


Fig. 45098. 


Tussilago Petasites L. Sp. Pl. 866. 1753. 
Petasites officinalis Moench. Meth. 568. 1794. 
Petasites vulgaris Desf. Fl. Atlant. 2: 270. 1798. 
P. Petasites Karst. Deutsch. Fl. 1062. 1880-83. 


Scape very scaly, 6-15’ high. Leaves orbicu- 
lar or hastate-reniform, often 12’ broad when 
mature, rounded or pointed at the apex, repand- 
denticulate all around, persistently white-to- 
mentose beneath, green and mostly glabrous 
above; heads 4-6” broad, mostly dioecious, 
in a dense raceme, the staminate ones smaller 
than the pistillate; flowers pink-purple, fra- 
grant, none of them radiate. 


In cultivated and waste ground, eastern Penn- 
sylvania and Massachusetts. Naturalized from 
Europe. Native also of northern Asia. Batter- 
or flea-dock. Bog- or poison-rhubarb. Eldin. 
Gallon. Umbrella-leaves. Pestilence-wort. Ox- 
wort. April. 


eS SS ee” | a. ee 


GENUS 97. THISTLE FAMILY. 533 


97. ARNICA L. Sp. Pl. 884. 1753. 


Erect, simple or little branched herbs, with opposite leaves, or the upper rarely alternate, 
and large, long-peduncled heads of both tubular and radiate, yellow flowers, or rays wanting 
in some species. Involucre turbinate or campanulate, its bracts in I or 2 series, narrow, 
nearly equal. Receptacle flat, naked, fimbrillate or villous. Ray-flowers pistillate, fertile, the 
rays spreading, entire, or 2-3-toothed. Disk-flowers perfect, fertile, the corolla 5-lobed, the 
style with slender branches. Anthers entire or minutely 2-auriculate at the base. Achenes 
linear, 5-10-ribbed, more or less pubescent. Pappus a single series of rough or barbellate, 
rigid, slender bristles. [Derivation uncertain, perhaps from Ptarmica.] F 


About 45 species, natives of the northern hemisphere. Besides the following, many others 
occur in the western parts of North America. Type species: Arnica montana L 


Basal leaves ovate or oval, sessile; southern. ] 1. A. acaulis. 
Basal leaves oblong, lanceolate, or cordate-ovate, petioled. na 
Basal leaves cordate-ovate. 2. A. cordifolia. 


Basal leaves not cordate, tapering to the petiole. 
Leaves dentate. 
Pappus brownish, plumose. 
Pappus white, barbellate. 
Leaves entire or nearly so. 


. mollis. 
. chionopappa. 


1. Arnica acatlis (Walt.) B.S.P. 
Leopard’s-bane. Fig. 4599. 


Doronicum acaule Walt. Fl. Car. 205. 1788. 
Arnica Claytoni Pursh, Fl. Am. Sept. 527. 1814. 
Arnica nudicaulis Nutt. Gen. 2: 164. 1818. 
Arnica acaulis B.S.P. Prel. Cat. N. Y. 30. 1888. 


Glandular-hirsute; stem 1°-3° high, bearing 
several slender-peduncled heads at the sum- 
mit. Basal leaves tufted, ovate or oval, obtuse, 
narrowed to a sessile base, denticulate or en- 
tire, 2-5’ long, 13’-3’ wide; stem leaves I-3 
pairs, and some alternate, very small ones 
above; heads 112’ broad; bracts of the invo- 
lucre linear-lanceolate, acute or acutish; rays 
12-15, commonly 3-toothed at the truncate 
apex; achenes pubescent when young, glabrous 
or nearly so when mature. 


In low woods, Delaware and southern Pennsyl- 
vania to Florida. April-May. 


2. Arnica cordifolia Hook. Heart- 
leaf Arnica. Fig. 4600. 


Arnica cordifolia Hook. Fl. Bor. Am. 1: 331. 
1833. 


Villous or pubescent; stem simple or 
sparingly branched, glandular above, 1°-2° 
high. Basal and lower leaves ovate to 
nearly orbicular, obtuse or acute, deeply 
cordate at the base, dentate, 173% long, 
with slender sometimes margined petioles; 
stem leaves I-3 pairs, ovate to oblong, ses- 
sile or short-petioled, much smaller; heads 
1-8, 2’-3’ broad; bracts of the involucre 
acute or acuminate, villous, 6-10” long; 
rays 12-16, toothed at the apex; achenes 
hirsute-pubescent, or glabrous at the base; 
pappus barbellate, white. 


Lake Superior to North Dakota, Yukon, 
Montana, New Mexico and California. Re- 
corded from western Nebraska. May-July. 


534 COMPOSITAE. Vo. III. 


3. Arnica mollis Hook. Hairy Arnica. 
Fig. 4601. 
Arnica mollis Hook. Fl. Bor. Am. 1: 331. 1833. 


Villous-pubescent; stem simple, or little 
branched, 1°-22° high, bearing 1-6 heads at 
the summit. Leaves oblong or oblong-lanceo- 
late, dentate or entire, acute or obtuse, 2-5’ 
long, 3-9” wide, the lower and basal ones 
narrowed into petioles, the upper sessile, and 
usually somewhat connate by a broad or nar- 
rowed base, those of the stem 3-5 pairs, usu- 
ally with some alternate small ones on the 
branches; heads 17-2’ broad; bracts of the in- 
volucre acute; rays I0-15, 3-toothed; achenes 
hirsute-pubescent; pappus yellow-brown, plu- 
mose. 


Mountains of northern New York, New Hamp- 
shire and Maine to New Brunswick, west to Lake 
Superior, British Columbia and California, south 
in the Rocky Mountains to Utah and Colorado. 
Included, in our first edition, in the northwestern 
A. Chamissonis Less. 


4. Arnica chionopappa Fernald. White-plumed 
Arnica. Fig. 4602. 
Arnica chionopappa Fernald, Rhodora 7: 148. 1905. 


Stem 15’ high or less, villous to the base. Leaves thin in 
texture, the lower and basal ones ovate to lanceolate, acute, 
narrowed at the base, sharply dentate with few teeth, peti- 
oled, 23’-42’ long, the upper few pairs narrowly lanceolate, 
entire, sessile; heads solitary or few, about 12’ broad; invo- 
lucre villous, 4’—5”’ high, its bracts linear-lanceolate, acumi- 
nate; rays I0-I5; pappus bright white, barbellate. 


On wet cliffs, Quebec and New Brunswick. June-July. 


Arnica gaspensis Fernald, from ledges in Gaspé County, Que- 
bec, has creamy-white pappus, the involucre and peduncles 
glandular-pubescent. 


5. Arnica alpina (L.) Olin & Laden. 
Mountain Tobacco. Arctic Leopard’s- 
bane. Arctic Arnica. Fig. 4603. 


Arnica montana var. alpina L. Sp. Pl. 884. 1753. 
Arnica alpina Olin & Laden, Diss. 11. 1799. 

4, A. angustifolia Vahl, Fl. Dan. pl. 1524. 1814. 
A. plantaginea Pursh, Fl. Am. Sept. 527. 1814. 
A. Sornborgeri Fernald, Rhodora 7: 147. 1905. 


Stem slender, 6-15’ high, villous or pubes- 
cent, and glandular or glabrous below; stem 
simple, usually bearing but a single head, but 
sometimes with 1-3 additional ones from the 
axils of the upper leaves. Leaves lanceolate, 
linear-oblong, or the lowest spatulate, thickish, 
entire or denticulate, 3-nerved, the basal ones 
petioled, those of the stem I-4 pairs, sessile 
or short-petioled, scarcely connate, the upper 
pair usually much smaller than the lower ones; 
heads about 2’ broad; rays 10-15, 3-toothed; 
achenes hirsute; pappus brownish. 


Labrador to Greenland and the Arctic Sea, 
Also in northern Europe. Races differ in pubes- 
cence. Rocky Mountain plants formerly referred 
to this species appear to be distinct. May—Sept. 


GENUS 98. THISTLE FAMILY. 535 


98. HAPLOESTHES A. Gray, Mem. Am. Acad. (II) 4: 109. 1849. 


Perennial caulescent, partly woody and partly fleshy plants, with opposite narrow entire 
leaves, the lower connate-sheathing, and corymbose heterogamous radiate heads. Invo- 
lucre of few broad many-nerved bracts. Receptacle flat or slightly convex, naked. Ray- 
flowers pistillate, fertile, the rays yellow, spreading or recurved. Disk-flowers perfect, fertile. 
Anthers obtuse and entire at the base. Style-branches of the disk-flowers capitate-truncate. 
Achenes narrow, 10-ribbed. Pappus of 1 series of slender scabrous bristles. [Greek, simple 
garment, the involucre composed of few bracts. ] 


A monotypic genus of the south-central United States and Mexico. 


UAT Ny Ne! 
arn sind f 
avy IN\ 


WW 


e” 
—_ 
VAL 
ys 
~ 


1. Haploesthes Greggii A. Gray. Gregg’s 
Haploesthes. Fig. 4604. 


a 


Haploesthes Greggii A. Gray, Mem. Am. Acad. (II) 
4: 109. 18409. 


Stems usually branched at the base, the branches 
1°-2° tall, glabrous, striate, corymbose above. 
Leaves fleshy, the lower ones connate and sheath- 
ing the stem, narrowly linear or linear-filiform, 
#’-2’ long, entire; heads short-peduncled, few to- 
gether in cymes; involucres 13’—2” high, the bracts 
oval to orbicular, rounded at the apex, thin-mar- 
gined; rays yellow, 1’—2” long; achenes 1” long. 


In saline and gypsum soil, Kansas and southern 
Colorado to Texas and Mexico. April—Sept. 


99. ERECHTITES Raf. Fl. Ludov. 65. 1817. 


Erect, usually branching herbs, with alternate leaves, and (in our species) rather large 
discoid many-flowered heads of whitish flowers, corymbose-paniculate at the ends of the 
stem and branches. Involucre cylindric, swollen at the base, its principal bracts in 1 series, 
linear, with or without some much smaller outer ones. Receptacle concave, naked. Marginal 
flowers in 2-several series, pistillate, fertile, their corollas filiform, the limb 2-4-toothed. 
Central flowers perfect, fertile; corolla narrowly tubular, the limb 4-5-toothed, the style- 
branches elongated, truncate or obtuse at the summit. Anthers obtuse and entire at the base. 
Achenes linear-oblong, angled or striate. Pappus of copious capillary soft smooth white 
bristles. [Ancient name of some groundsel.] 


About 12 species, natives of America and Australasia. The following typical one is the only 
species known to occur in North America. 


COMPOSITAE. Vo. III. 


1. Erechtites hieracif6lia (L.) Raf. 
Fire-weed. Fig. 4605. 
Senecio hieracifolius L. Sp. Pl. 866. 1753. 

E. prealta Raf. Fl. Ludov. 65. 1817. 


Erechtites hieracifolia Raf. DC. Prodr. 6: 

294. 1837. 

Annual, glabrous, or somewhat hir- 
sute; stem striate, succulent, usually 
branched, 1°-8° high. Leaves thin, lan- 
ceolate or ovate-lanceolate, dentate and 
often deeply incised, 2’-8’ long, the 
upper sessile or auriculate-clasping, 
mostly acuminate, the lower usually nar- 
rowed into petioles; heads 6’—10” long, 
about 3” in diameter, the involucre con- 
spicuously swollen at the base before 
flowering, its bracts numerous, striate, 
green, with narrow scarious margins; 
pappus bright white. 

In woodlands, thickets and waste places, 
very abundant after fires, Newfoundland to 
Florida, Ontario, Saskatchewan, Nebraska 
and Texas. Also in Mexico, the West In- 


dies and South America. Pilewort. July- 
Sept. 


100. MESADENIA Raf. New 
FL N. A..4278: Saeae 


Tall perennial mostly glabrous herbs, with alternate petioled leaves and numerous, rather 
small, corymbose, discoid heads of white, yellowish or pinkish flowers, all tubular and perfect. 
Sap milky (at least in some species). Involucre cylindric or nearly so, its principal bracts 
s, in I series, equal, usually with a few short outer ones. Receptacle flat, not chaffy, with a 
fleshy projection in the center. Corollas with somewhat spreading 5-cleft limbs, the lobes 
usually with a mid-nerve. Style-branches conic or obtuse at the apex. Achenes oblong, 
glabrous. Pappus of copious white scabrous bristles. [Greek, referring to the central pro- 
jection of the receptacle.] 

About 30 species, natives of North and Central America. Besides the following, 4 others occur 


in the southern and southwestern parts of the United States. Type species: Mesadenia atriplici- 
folia (L.) Raf. 


\ 
Leaves thin, reniform or fan-shaped, lobed, or angulate-dentate. 


Leaves green both sides, angulate-dentate. Toe reniformis. 
Leaves glaucous beneath, green above, angulate-lobed. 2. M. atriplicifolia. 
Leaves thick, green both sides, ovate or oval, entire, or repand. 3. M. tuberosa. 


1. Mesadenia renifoérmis ( Muhl.) 
Raf. Great Indian Plantain. 
Wild Collard. Fig. 4606. 


Cacalia reniformis Muhl.; Willd. Sp. Pl. 3: 
1753. 1804. : 

Mesadeniareniformis Raf. New F1.4: 79. 1836. 

M. rotundifolia Raf. New Fl. 4: 79. 1836. 


Glabrous; stem angled and grooved, 4°- 
10° high. Leaves thin, green both sides, 
coarsely angulate-dentate with mucronate- 
pointed teeth, the basal and lower reniform, 
long-petioled, sometimes 2° wide, the upper 
ovate or fan-shaped, mostly cuneate at the 
base, the uppermost small and oblong; heads 
numerous, mostly 5-flowered, about 2” 
broad, in large compound corymbs; invo- 
lucre 3-4” high, its bracts linear-oblong, 
obtuse or acutish, scarious-margined, with 
or without I-3 minute outer ones. 


In woods, New Jersey and Pennsylvania to 
Minnesota, south to North Carolina and Ten- 
nessee. July—Sept. 


GENUS 100. THISTLE FAMILY. 537 


2. Mesadenia atriplicifolia (L.) Raf. 


Cacalia atriplicifolia L. Sp. Pl. 835. 1753. 


Senecio atriplicifolius Hook. Fl. Bor. Am. 1: 
Bae. 1833. 


Mesadenia atriplicifolia Raf. New FI. 4: 79. 
1836. 5 
Stem terete, glabrous and glaucous, 

3°-6° high. Leaves thin, angulate-lobed, 

palmately veined, glaucous beneath, the 
lower and basal ones slender-petioled, 
sometimes 6’ wide, the upper reniform, 
fan-shaped, or triangular with a nearly 
truncate base, the uppermost commonly 
small, lanceolate or oblong and entire; 
heads very numerous, about 12” broad, 
in large, loose compound corymbs; invo- 
lucre 3-4” high, its bracts linear-oblong, 

scarious-margined, with or without 1-3 

minute outer ones. 


In woods, New Jersey to Indiana, Minne- 
sota, south to Florida, Tennessee, Missovri 
and Kansas. Recorded from Ontario. Called 
also wild caraway. July—Sept. 


3. Mesadenia tuberésa ( Nutt.) Britton. 
Tuberous Indian Plantain. Fig. 4608. 


Cacalia tuberosa Nutt. Gen. 2: 138. 1818. 

Mesadenia plantaginea Raf. New Fl. 4: 79. 1836. 

Senecio Nuttallii Sch. Bip. Flora 27: 499. 1845. 

M. tuberosa Britton in Britt. & Brown, Ill. Fl. 3: 
474. 1808. 


Glabrous and green throughout; stem angled, 
stout, 2°-6° high. Leaves thick, strongly 5-0- 
nerved, the lower and basal ones oval, ovate, 
or ovate-lanceolate, obtuse or acutish, usually 
quite entire, but sometimes repand, long-peti- 
oled, narrowed at the base, or rarely subcordate, 
4-8 long, 1-3 wide; upper leaves ovate to 
oblong or cuneate-obovate, sessile or short-peti- 
oled, much smaller, sometimes toothed toward 
the apex; heads very numerous in a compound 
corymb, about 2” broad, mostly 5-flowered; 
involucre 3’-4” high, its bracts linear-oblong, 
obtuse or obtusish, scarious-margined. 


On wet prairies and in marshes, Ohio and west- 
ern Ontario to Minnesota, Kansas, Alabama, Louisi- 
ana and Texas. June—Aug. 


ror. SYNOSMA Raf.; Loud. Gard. Mag. 8: 247. 1832. 


A perennial leafy herb, glabrous or very nearly so, with triangular or hastate, alternate 
leaves, the lower petioled, the upper sessile, and several or numerous, corymbose or corym- 
bose-paniculate, discoid heads of white or pinkish flowers. Involucre nearly cylindric, its 
principal bracts 12-15, linear, acute, usually with some subulate outer ones. Receptacle flat, 
naked. Flowers perfect; corolla 5-lobed. Style-branches not appendaged. Pappus of very 
numerous white soft capillary bristles. [Greek, perhaps signifying a fragrant composite. ] 


A monotypic genus of eastern North America. 


539 COMPOSITAE. Vor. Tie 


1. Synosma suavéolens (L.) Raf. Sweet-scented Indian Plantain. Fig. 4609. 


\ 


‘ 


Cacalia suaveolens L. Sp. Pl. 835. 1753. 
AN 
Wi 


VE vi | 
Dr, WZ Senecio suaveolens Ell. Bot. S.C. & Ga. 2: 328. 1821-24. 
Uys Synosma suaveolens Raf.; Loud. Gard. Mag. 8: 247. 


1832. 


Glabrous or very nearly so throughout; stem stri- 
ate, 3°-5° high, leafy to the inflorescence. Leaves 
triangular-lanceolate or hastate, sharply and irregu- 
larly serrate, acuminate, 4’-10’ long, 2’-6’ wide at 
the base, the auricles sometimes with I or 2 lobes 
on the lower side; petioles margined, or those of 
the basal leaves naked and slender; uppermost leaves 
sometimes merely lanceolate and sessile; heads 2’—3” 
broad in a usually large and compound corymb; in- 
volucre 4’-6” high, its principal bracts linear, acute; 
heads 20-30-flowered. 


In woods, Rhode Island to Pennsylvania, New Jersey, 
Illinois, Minnesota, Florida, West Virginia and Ken- 
tucky. Called also wild caraway. Aug.—Oct. 


102. SENECIO [Tourn.] L. Sp. Pl. 866. 1753. 


Annual or perennial herbs (some tropical species shrubby or even arborescent), with 
alternate or basal leaves, and solitary corymbose or paniculate many-flowered heads, of both 
tubular and radiate, or only tubular flowers, in our species yellow. Involucre cylindric or 
campanulate, its principal bracts in I series, distinct, or united at the base, usually with some 
shorter outer ones. Receptacle flat or slightly convex, mostly naked, often honeycombed. 
Rays, when present, pistillate, fertile. Disk-flowers perfect, fertile, their corollas tubular, 
the limb 5-toothed or 5-lobed. Anthers obtuse and entire at the base, or rarely slightly sagit- 
tate. Style-branches of the disk-flowers usually recurving or spreading. Achenes terete, 
or those of the marginal flowers somewhat compressed, 5-10-ribbed, papillose or canescent, 
at least after wetting, and then usually emitting a pair of spiral threads. Pappus of numerous 
slender or capillary, smooth or rough, mostly white bristles. [Latin, senex, an old man, 
‘referring to the hoary character of some species, or to the white pappus. | 


An immense genus of probably at least 1200 species, of very wide geographic distribution. In 
addition to the following, many others occur in the southern and western parts of North America. 
Our species known as Groundsel, Ragwort, or Squaw-weed. Type species: Senecio vulgaris L. 


A. Annual or biennial species, with stems leafy throughout. 
Rays none, or very short and inconspicuous ; introduced. 


The short outer involucral bracts black-tipped ; rays none. 1. S. vulgaris. 
The short outer involucral bracts not black-tipped, sometimes none; rays very short. 
Plant sparingly pubescent. 2. S. sylvaticus. 
Plant densely viscid-pubescent. 3. S. viscosus. 
Rays large and conspicuous; native species. 
Leaves pinnately divided; heads 2”—3” high. 4. S. glabellus. 
Leaves sinuate-dentate or the lower entire; heads 47-5” high. 5. S. palustris. 


B. Perennial species, mostly with rootstocks. 


a. Stems woody at base; leaves linear, or pinnatifid with linear lobes; western species. 
Leaves linear, entire or serrate, thin. 6. S. spartioides. 
Leaves deeply pinnatifid into linear lobes, firm. 7, S. Riddellii. 


b. Stems wholly herbaceous; leaves various. 
* Heads very large; involucre 7”-10” high; boreal species. 


Stem stout, leafy above; heads several, 1%4’—2’ broad. 8. S. Pseudo-Arnica. 
Stem slender ; upper leaves few and small; head mostly solitary, about 1’broad. 9. 5S. frigidus. 


** Heads smaller; involucre 4”-7” high. 


+ Leafy up to the inflorescence; stem leaves 2~-3-pinnatifid; introduced species. 
10. S. Jacobaea. : 
++ Leaves mostly borne on the lower part of the stem, the upper ones much smaller; native species. 


+ Leaves and stems more or less persistently woolly or tomentose. 
Low species, seldom over 1° high, with small oval-oblong to spatulate basal leaves. . 
Basal leaves angulate-dentate, oval. 11. S. antennariitfolius. 
Basal leaves entire or sparingly toothed, oblong to spatulate. 12. S.canus. 


GENUS I02. THISTLE FAMILY. 539 


Taller, up to 2%4° high; basal leaves ovate to oblong-lanceolate. 

Densely persistently tomentose ; stem-leaves mostly merely dentate. 13. S. tomentosus. 

Loosely tomentose, becoming glabrate; stem leaves mostly pinnatifid. 14. S. plattensis. 

tt Plants glabrous, or nearly so, at least when mature, the stem sometimes tomentose at the base. 

Basal leaves or some of them deeply cordate. 

Basal leaves lanceolate or oblong-lanceolate. 15. S. Robbinsii. 

Basal leaves orbicular to ovate. 16. S. aureus. 
None of the leaves cordate. 

At least the stem-leaves lobed, laciniate, or pinnatifid; pubescence, if any, woolly; basal leaves 

dentate or crenate. 


Head rayless ; boreal species. 17. S. discoideus. 
Heads radiate. : 
Basal leaves obovate to suborbicular. 18. S. obovatus. 


Basal leaves oblong to spatulate. 
Basal leaves cblong to lanceolate, dentate or crenate. 


Involucre 4” high; basal leaves sharply serrate. 19. S. Crawfordii. 
Involucre 2%4”-3%” high; basal leaves mostly crenate. 
Heads few; basal leaves mostly short. 20. S. pauperculus. 
Heads very numerous; basal leaves long. 21. S. Smallii. 
Basal leaves linear-cuneate, entire, or few-toothed at the apex. 22. S. densus. ; 
All leaves entire or very nearly so; pubescence, if any, of crisp hairs. 23. S. integerrimus. 


1. Senecio vulgaris L. Common Ground- 
sel. Fig. 4610. 


Senecio vulgaris L. Sp. Pl. 867. 1753. 


Annual, puberulent or glabrate; stem hollow, 
usually much branched, 6-15’ high. Leaves pin- 
natifid, 2-6’ long, the lower spatulate in outline, 
petioled, obtuse, the upper sessile or clasping at 
the base, more deeply lobed or incised, their seg- 
ments oblong, dentate; heads several or numer- 
ous in the corymbs, nearly 3” broad, 4-6” high; 
bracts of the involucre linear, with few or sev- 
eral subulate black-tipped outer ones; rays none; 
achenes slightly canescent; pappus white. 


In cultivated ground and waste places, Newfound- 
land to Hudson Bay, North Carolina, Minnesota, 
Michigan, and west to the Pacific Coast. Bermuda. 
Naturalized from Europe. Other names are grinsel, 
simson, birdseed, chickenweed. April—Oct. 


2. Senecio sylvaticus L. Wood Groundsel. 
Fig. 4611. 


Senecio sylvaticus L. Sp. Pl. 868. 1753. 


Annual, glabrous or puberulent; stem usually much branch- 
ed, 1°-23° high, leafy. Leaves pinnatifid, oblong or lanceo- 
late in outline, the segments oblong or spatulate, obtuse, 
dentate, lobed or entire, or the uppermost leaves linear and 
merely dentate; heads several or numerous in the corymbs, 
slender-peduncled, about 2” broad, 3’-4” high; involucre 
usually quite naked and swollen at the base; rays very short 
and recurved; achenes canescent; pappus white. 


In waste places Newfoundland to Nova Scotia, Quebee and 
Maine. Also on the coasts of California and British Columbia. 
Naturalized or adventive from Europe. April—Sept. 


COMPOSITAE. Vor. ilT. 


3. Senecio viscdsus L. Fetid or Viscous Groundsel. 
Fig. 4612. 
Senecio viscosus L. Sp. Pl. 868. 1753. 


Annual, viscid-pubescent, strong-scented; stem usually 
much branched, 1°-2° high. Leaves 1-2-pinnatifid, 14’-3’ 
long, oblong or somewhat spatulate in outline, the segments 
oblong or cuneate, dentate or incised; lower leaves peti- 
oled; heads few in the corymbs, 3-4” broad, mostly slen- 
der-peduncled; involucre nearly cylindric, 4’-5” high, its 
bracts linear, acute, with 1-3 shorter outer ones; rays com- 
monly about 20, very short, recurved and inconspicuous; 
achenes glabrous; pappus bright white, about one-third 
longer than the involucre. 


In waste places and on ballast near the coast, Nova_ Scotia, 
New Brunswick and Quebec to North Carolina. July—Sept. 


4. Senecio glabéllus Poir. Butterweed. Cress-leaved Groundsel. Fig. 4613. 
S. lyratus Michx. Fl. Bor. Am. 2: 120. 1803. Not L. 1753. 
S. glabellus Poir. in Lam. Encycl. 7: 102. 1806, 

Senecio lobatus Pers. Syn. 2: 436. 1807. 


Annual, glabrous throughout, or slightly woolly 
when young, fleshy and tender; stem hollow, sim- 
ple or branched, 1°-3° high. Leaves 2’-10’ long, 
pinnately divided, the segments orbicular, oblong, 
obovate or cuneate, obtuse, sinuate-dentate, entire or 
lobed, the terminal segment usually larger than the 
others; lower and basal leaves slender-petioled; 
heads numerous, 7’-10” broad, slender-peduncled 
in terminal corymbs; involucre nearly cylindric, 23” 
high, its bracts linear, acute, usually with no small 
outer ones; rays 6-12; achenes minutely hispidu- 
lous on some of the angles; pappus white, somewhat 
longer than the involucre. 


In swamps, North Carolina to Kentucky, Illinois, Mis- 
souri, Arkansas, Florida, New Mexico and Mexico. 
April-Sept. 


5. Senecio palustris (L.) Hook. Marsh Fleawort. Pale Ragwort. Marsh 
Groundsel. Fig. 4614. 


Cineraria palustris L. Sp. Pl. Ed. 2, 243. 1763. 
Senecio palustris Hook. Fl. Bor. Am. 1: 334. 1833. 


Annual or biennial, pubescent or glabrate; stem 
stout, simple, hollow, 6-24’ high. Leaves lanceo- 
late, oblong or spatulate, entire, dentate, or lacin- 
iate, acute or obtuse, 2-7’ long, 3-15’ wide, or 
the upper linear-lanceolate and small, those of 
the stem sessile and somewhat auriculate-clasping, 
the basal petioled; heads numerous, 6-12” broad, 
mostly short-peduncled in a large, rather dense, 
terminal corymb; involucre cylindric, becoming 
campanulate, 3-4” high, its bracts linear, acute, 
more or less pubescent, with no shorter outer 
ones; rays I5-20 or more, pale-yellow; achenes 
glabrous; pappus white, elongated, at length twice 
the length of the involucre. 


In swamps, Iowa and Wisconsin to Manitoba and 
arctic America, west to Alaska. Reported from Lab- 
rador. Also in Greenland, northern Europe and Asia. 
June-Aug. 


GENUS 102. ANSUUSMNES, VA Oe 541 


6. Senecio spartioides T. & G. Broom-like 
Senecio. Fig. 4615. 
Senecio spartioides T.& G. Fl. N. A. 2: 438. 1843. 


U; 
x a = 


Woody at the base, usually branched, sometimes 
shrubby, glabrous or nearly so, leafy, 1°-6° high. 
Leaves sessile, or the lowest petioled, 1-3’ long, 
linear, entire, or more or less serrate, not lobed; 
heads corymbose at the ends of the branches, 4’-1’ 
broad, slender-peduncled; involucre cylindric or 
becoming campanulate, 4”-5” high, its -bracts 
linear, acute or acuminate, usually with some \ 
subulate exterior ones; rays 8-15; achenes ca- 
nescent; pappus bright white. 


Plains, in dry soil, Nebraska to Texas, Wyoming 
and Arizona. June-Sept. This and the following 
species were included in the description oi the far 
western S. Douglasii DC. in our first edition. 


7. Senecio Riddellii T. & G. Riddell’s 
Senecio. Fig. 4616. 


S. Riddellui T. & G. FI. N. A. 2: 444. 1843. 
S. Fremontii (T. & G.) Rydb.; Britton, Manual 1028. 
1901. Not S. Fremontii T. & G. 


Woody at the base, usually branched, some- 
times shrubby, glabrous or nearly so, leafy, 1°-6° 
high. Leaves sessile, or the lowest petioled, thick, 
14’-34’ long, pinnately parted into 3-9 linear or 
filiform, entire segments, or the upper entire; 
heads corymbose at the ends of the branches, 
5-10” broad, slender-peduncled; involucre cylin- 
dric or becoming campanulate, 5-8” high, its 
bracts linear, acute or acuminate, usually with 
some subulate exterior ones; rays 8-15; achenes 
canescent; pappus white. 


Plains, in dry soil, Nebraska to Texas and Mexico. 
June-Sept. 


8. Senecio Psetido-Arnica Less. Sea-beach 
Senecio. Fig. 4617. 


Arnica maritima L. Sp. Pl. 884. 1753. Not S. mari- 
timus L. 
Senecio Pseudo-Arnica Less. Linnaea 6: 240. 1831. 


Perennial, somewhat fleshy; stem stout, mostly 
simple, very leafy, 63° high. Leaves oblong-obo- 
vate, lanceolate, or the lower spatulate, acute or 
obtuse at the apex, 4-8’ long, 4’-2’ wide, densely 
tomentose beneath, at least when young, repand- 
dentate or denticulate, narrowed to a sessile and 
partly clasping base, or the lowest into margined 
petioles; heads solitary, or several (2-7) and co- 
rymbose, stout-peduncled, 14’-2’ broad, 8-10” 
high; involucre broadly campanulate, its bracts 
lanceolate, acuminate, mostly tomentose, common- 
ly with several subulate spreading ones at the base; 
rays 12-25, linear, 3-toothed, conspicuous; disk- 
corollas 5-lobed; achenes glabrous; pappus dull. 


On sea-beaches and rocks near the sea, Maine, New 
Brunswick and the lower St. Lawrence to Labrador 
and the Arctic Sea. Also in Alaska. July—Aug. 


542 COMPOSITAE. Vot. III. 


g. Senecio frigidus Less. Arctic Senecio. 
Fig. 4618. 


Senecio frigidus Less. Linnaea 6: 239. 1831. 


Perennial, more or less tomentose, or becoming 
glabrous when old; stem slender, 6’-12’ high, bearing 
a solitary head (rarely 2 or 3) 3’-1’ broad. Basal 
and lower leaves spatulate or obovate, 1’-2’ long, peti- 
oled, obtuse, repand-dentate or entire; stem leaves 
oblong to linear-lanceolate, obtuse or acute, sessile, 
mostly entire, smaller; involucre broadly campanu- 
late, about 7” high, its bracts lanceolate, acute, with 
no exterior smaller ones; rays 10-16, 6’—10” long, 
3-toothed, linear-oblong, or cuneate at the base; 
achenes glabrous or sparingly pubescent; pappus 
white. 


Labrador and arctic America to Alaska. Also in north- 
eastern Asia. Summer. 


10. Senecio Jacobaéa L. Tansy Ragwort. 
Staggerwort. Fig. 4619. 


Senecio Jacobaea L. Sp. Pl. 870. 1753. 


les Pak B 


Perennial by short thick rootstocks, somewhat 
woolly, or glabrous; stems stout, simple, or 
branched above, 2°-4° high, very leafy. Stem 
leaves 2-3-pinnatifid, 2-8’ long, the lower peti- 
oled, the upper sessile, the lobes oblong-cuneate, 
dentate or incised; basal leaves lyrate-pinna- 
tifid; heads very numerous, 6’—-8” broad, short- 
peduncled in large compact corymbs; involucre — 
narrowly campanulate, about 23” high, its bracts 
linear-lanceolate, acute, green, or tipped with 
black, usually with a few subulate outer ones; 
rays 12-15; achenes of the disk-flowers pubes- 
cent, those of the rays glabrous; pappus white. 

In waste places, Newfoundland and Nova Scotia to 
Maine and Ontario, and in ballast about New York 
and Philadelphia. Adventive from Europe. Stave- 
wort. Cankerweed. Kettle-dock. St. James’-wort. 
Felonweed. Fairies’-horse. Ragweed. Saracen’s-com- 
pass. July—Sept. 


tz. Senecio antennariifdlius Britton. Cat’s- 
paw Ragwort. Fig. 4620. 


Senecio antennariifolius Britton, in Britt. & Brown, Ill. 
Fl. 3: 478. 1808. 


Perennial, tufted in mostly large clumps; stems 
slender, 8-18’ high, loosely white-woolly. Leaves 
nearly all basal, commonly numerous, oval to spatu- 
late, angulately few-toothed or entire, mostly obtuse, 
narrowed into a petiole as long as the blade or longer, 
densely white-tomentose beneath, green and finally 
glabrous above, 1-23’ long; stem leaves small, spatu- 
late, laciniate, or the upper narrowly linear and en- 
tire; heads several, corymbose, slender-peduncled, 
rathed less than 1’ broad; rays golden-yellow, showy; 
involucre 3” high, white-woolly; achenes glandular- 
pubescent. 


. 


Stony hillsides, mountains of Virginia and West Vir- 
ginia. May—June. 


a ee ee a 


GENUS 102. THISTLE FAMILY. 543 


12. Senecio canus Hook. Silvery Ground- 
sel. Fig. 4621. 


S. canus Hook. Fl. Bor. Am. 1: 333. p/. 116. 1833. 


S. Purshianus Nutt. Trans. Am. Phil. Soc. (II) 7: 
Ame. TOA tT. 


Perennial, densely and persistently white- 
tomentose to the inflorescence; stems slender, 
usually tufted, 6-18’ high. Basal and lower 
leaves spatulate or oval, entire, or rarely some- 
what repand, very obtuse, 1-2’ long, narrowed 
into petioles; upper leaves oblong or spatulate, 
obtuse or acute, mostly sessile, smaller, entire 
or dentate; heads several or numerous, 8’—10” 
broad, usually slender-peduncled; involucre 
campanulate, or at first short-cylindric, about 
5” high, its bracts linear-lanceolate, acute, spar- 
ingly tomentose, or glabrate, usually with no 
exterior smaller ones; rays 8-12; achenes gla- 
brous, at least below; pappus white. 


In dry soil, Manitoba to North Dakota, Nebraska, 
west to British Columbia and California. Recorded 
from Minnesota. May—Aug. 


13. Senecio tomentdosus Michx. Woolly 
Ragweed. Ashwort. Fig. 4622. 
S. tomentosus Michx. Fl. Bor. Am. 2: 119. 1803. 


Perennial, more or less densely and persistently 
tomentose or woolly-canescent; stems rather stout, 
solitary, or sometimes tufted, 1°-23° high. Basal 
and lower leaves ovate-lanceolate, oblong or rarely 
spatulate, long-petioled, erect, very obtuse, 2’-6’ 
long, 2-2’ wide, crenate-dentate, narrowed or 
truncate at the base; stem leaves few and distant, 
small, linear-lanceolate or spatulate, crenate or 
rarely laciniate; heads several or numerous, co- 
rymbose, mostly long-peduncled, 8’-10” broad; 
_involucre cylindric, or narrowly campanulate, its 
bracts linear-lanceolate, acute or acuminate, at 
length glabrate, with or without a few small outer 
ones; rays I0-15; achenes hispidulous, at least 
on the angles; pappus white. 


In moist soil, southern New Jersey to Florida, 
Louisiana, Texas and Arkansas. April-June. 


14. Senecio platténsis Nutt. Prairie Rag- 
wort. Fig. 4623. 


SA pcre Nutt. Trans. Am. Phil. Soc. (IT) 7: 413. 
1841. 


Perennial, similar to the preceding species, usu- 
ally smaller-leaved, lower and less tomentose, or 
becoming glabrate in age, seldom over 14° high. 
Basal leaves oval, ovate or oblong, some or all of 
them often more or less pinnatifid, with the termi- 
nal segment much larger than the lateral ones, 
crenulate or dentate, long-petioled; stem leaves 
mostly smaller than the basal ones, usually pin- 
natifid; heads several or numerous, compactly or 
loosely corymbose, conspicuously radiate. 


Indiana and Illinois to Ontario, North Dakota, Colo- 
rado, Missouri and Texas. April—June. 


Senecio pseudotomentésus Mackenzie & Bush, of 
Missouri, differs in having the basal leaves mostly 
merely dentate. 


544 , COMPOSITAE. Vor. III. 


“15. Senecio Robbinsii Oakes. Robbins’ 
Squaw-weed. Fig. 4624. 


Senecio Robbinsii Oakes; Rusby, Bull. Torr. Club 20: 
19. 1893. 


Perennial, glabrous or very nearly so through- 
out; stems slender, 1°-23° high. Basal leaves 
long-petioled, lanceolate, obtuse or acute at the 
apex, cordate, subcordate or truncate at the base, 
usually thin, 2-4’ long, 4’-12’ wide, sharply den- 
tate; stem leaves mostly pinnatifid or lobed, at 
least below the middle; heads several or numer- 
ous, slender-peduncled in an open corymb, 8-10” 
broad; rays 6-12; achenes glabrous or pubescent; 
pappus white. 


_In swamps and mountain meadows, Nova Scotia to 
New Hampshire, Vermont and New York. June— 
ept. 


16. Senecio atreus L. Golden Ragwort. 
Life-root. Swamp Squaw-weed. 
Fig. 4625. 

Senecio aureus L. Sp. Pl. 870. 1753. 


Senecio pauciflorus Pursh, Fl. Am. Sept. 529. 1814. 
Senecio gracilis Pursh, Fl. Am. Sept. 529. 1814. 


Perennial, glabrous or very nearly so through- 
out; stems rather slender, solitary or tufted, 
6’-23° high. Basal leaves cordate-ovate or 
cordate-orbicular or reniform, crenate-dentate, 
very obtuse and rounded, often purplish, 1-6’ 
long, with long slender petioles; lower stem 
leaves lanceolate or oblong, usually laciniate, 
pinnatifid or lyrate, the uppermost small, ses- 
sile, somewhat auriculate and clasping; heads 
usually several, 8’—10” broad, 4-5” high, slen- 
der-peduncled in an open corymb; rays 8-12, 
golden-yellow ; achenes glabrous; pappus white. 


In swamps and wet meadows, Newfoundland to 
Florida; Ontario, Michigan, Missouri and Texas. 
Grundy-swallow. False-valerian. Root strong- 
scented. Races differ in size of plant, size of 
leaves and number of heads. May-July. 


Senecio pseudaiireus Rydb. (S. semi-cordatus Mac- 
kenzie & Bush), of the Rocky Mountain region, differing 
by elliptic or broadly oblong basal leaves, ranges east- 
ward into North Dakota and Missouri. 


17. Senecio discdideus (Hook) Britton. 
Northern Squaw-weed. Fig. 4626. 


Senecio aureus var. discoideus Hook. Fl. Bor. Am, 1: 

333. 1833. 

Senecio discoideus Britton, in Britt. & Brown, Ill. FI. 

3: 479. 18098. 

Perennial, glabrous except for small tufts of wool 
in the axils of the lower leaves; stem rather stout, 
1°-23° tall; basal leaves oval to ovate, obtuse, thin, 
sharply dentate, abruptly narrowed into petioles 
longer than the blade; stem leaves few, small, more 
or less laciniate; heads few or several, slender- 
peduncled, corymbose; principal bracts of the invo- 
lucre narrowly linear, 3-5” long, the short outer 
ones few or none; rays none; achenes glabrous. 


In moist places, Labrador to Yukon, Quebec, Michi- 
gan, Wyoming and British Columbia. June—Aug. 


GENUS 102. THISTLE FAMILY. 545 


18. Senecio obovatus Muhl. Round-leaf 
Squaw-weed. Fig. 4627. 
Senecio obovatus Muhl.; Willd. Sp. Pl. 3: 1999. 1804. 
Senecio Elliottii T.& G. Fl. N. A. 2: 443. 1843. 
S. aureus var. obovatus T. & G. loc. cit. 442. 1843. 
S. rotundus (Britton) Small, Fl. SE. U. S. 1304. 1903. 

Perennial; stems glabrous, or a little woolly at 
the base, 9-24’ high. Leaves glabrous, rather thick, 
the basal ones obovate with a cuneate base, subor- 
bicular or broadly spatulate, very obtuse and round- 
ed at the apex, 17-33’ long, 2-2’ wide, crenate- 
dentate, often purplish; stem leaves commonly few 
and sessile, spatulate to oblong, often incised or 
pinnatifid; heads several, corymbose, 6-8” broad, 
about 3” high, slender-peduncled; involucre nearly 
cylindric, its principal bracts linear-lanceolate, 2’—3” 
long, acute, usually with 1-3 small exterior ones; 
rays 8-12, usually conspicuous, sometimes fewer 
and short; achenes glabrous; pappus white. 

In moist soil on banks and in woods, Maine and Ver- 
mont to Florida, Ohio, Michigan, Alabama and Texas. 
Races differ in size, leaf-form, number and size or rays. 
Apparently erroneously recorded from Nova Scotia and 
Ontario. April-June. 


Ig. Senecio Crawfordii Britton. Crawford’s 
Squaw-weed. Fig. 4628. 


Senecio Crawfordii Britton, Torreya 1: 21. 1901. 
S. Balsamitae var. Crawfordi Greenman, Rhodora 10: 69. 
1908. 


Perennial, glabrous, or with sparse woolly pubescence 
below; stem slender, about 16’ high. Leaves thick, firm, 
the basal ones erect, the larger 8-10’ long, the blades 
oval, oblong, or some of them narrowly obovate, mostly 
not more than one-half as long as the slender petioles, 
sharply and nearly equally serrate from the apex to the 
entire cuneate base; stem leaves lanceolate or narrower, 
mostly acuminate, incised-serrate, clasping, the upper 
sessile, the lower petioled, the uppermost very small; 
heads 3-7; peduncles slender, bracted, rarely forked; 
involucre 4” high, its bracts linear-lanceolate, acumi- 
nate, shorter than the white barbellate pappus; rays 
4-5” long; achenes linear, striate. 


Wet meadows, southeastern Pennsylvania. May-June. 


20. Senecio pauperculus Michx. Balsam 
Groundsel. Fig. 4629. 
ce teeaeg pauperculus Michx. Fl. Bor. Am. 2: 120. 

1803. 

Senecio Balsamitae Muhl.; Willd. Sp. Pl. 1999. 1804. 
Senecio aureus var. Balsamitae T. & G. Fl. N. A. 2: 

442. 1843. 

Perennial, often tufted; stems slender, 14’-20’ 
high, woolly at the base and in the axils of the 
lower leaves, or essentially glabrous. Basal leaves 
slender-petioled, oblong, rarely slightly spatulate, 
very obtuse, narrowed at the base, mostly thick, 
crenate, or rarely dentate, often purplish, 1-5’ 
long, 3-6” wide, their petioles and sometimes 
their lower surfaces persistently tomentose or 
woolly, or glabrous throughout; lower stem leaves 
petioled, laciniate or pinnatifid, the upper sessile, 
very small; heads few or several, slender-pedun- 
cled, 6’—10” broad; involucre about 3” high; rays 
8-12; achenes hispidulous or glabrous; pappus 
white. 

In dry or rocky soil, Newfoundland to North Caro- 
lina, Ontario, British Columbia, Alabama, Tennessee 
and Nebraska. May-July. 


35 


COMPOSITAE. Vor. III. 


21. Senecio Smallii Britton. Small’s Squaw- 
weed. Fig 4630. 


S. aureus var. angustifolius Britton, Mem. Torr. Club 
2:39. 1890. Not S. angustifolius Willd. 1804. 


S. Smallii Britton, Mem. Torr. Club 4: 132. 1893. 


Similar to the preceding species but taller, grow- 
ing in large clumps; stem 12°-23° high, slender, 
densely and persistently floccose-woolly at the base 
and in the lower axils, or finally glabrate. Basal 
leaves elongated-oblong or linear-oblong, obtuse or 
acute, long-petioled, crenate-dentate, 3/-6’ long, 
3-12” wide, at first tomentose, at length nearly 
glabrous; stem leaves several, deeply pinnatifid, or 
the lower lyrate, the uppermost very small; heads 
very numerous, 4”-5” broad, about 23” high, slen- 
der-peduncled, forming large corymbs; rays 8-10; 
achenes hispidulous; pappus white. 


In meadows and thickets, southeastern Pennsylva- 
nia to Florida and Alabama. May-June. . 


22. Senecio densus Greene. Western Squaw- 
weed. Fig. 4631. 


yy) 


WWD 
yy 
y 


Senecio aureus var. compactus A. Gray, Syn. Fl. 1: Part 2, 
391. 1884. 

Senecio compactus Rydberg, Mem. Torr. Club 5: 342. 1893. 
Not T. Kirk. 

Senecio densus Greene, Pittonia 4: 226. 1900. 


Perennial; stem usually tufted, low, rather stout, 
6-12’ high, woolly at the base and in the lower axils, 
or glabrous. Basal leaves linear-cuneate, entire or 
3-toothed at the apex, 1’-3’ long, 2’-3” wide, thick, 
slender-petioled, the petioles commonly woolly-mar- 
gined; lower stem leaves often much larger and broader, 
usually laciniate or pinnatifid, but sometimes similar to 
the basal, the uppermost very small and sessile; heads 
several, 8-10” broad, short-peduncled in a compact co- 
rymb; rays 10-15; achenes hispidulous; pappus white. 


On dry plains, Manitoba to Nebraska, Colorado and Texas. 
May-June. 


YZ 
ZY 


23. Senecio integérrimus Nutt. Entire- 
leaved Groundsel. Fig. 4632. 
Senecio integerrimus Nutt. Gen. 2: 165. 1818. 


Perennial, more or less pubescent when young, 
glabrous or nearly so when old; stem stout, 1°-4° 
high. Leaves entire, or sparingly denticulate, 
somewhat fleshy, the lower and basal ones oval 
or oblong, obtuse or obtusish at the apex, 3-8’ 
long, 1’-13’ wide, petioled, the upper linear or 
lanceolate, acute, the uppermost very small; heads 
numerous, corymbose, long-peduncled, 6’—10” 
broad; involucre nearly cylindric, *4’—-5” high, its 
principal bracts linear, acuminate, green, usually 
with a few subulate outer ones; rays 8-12, linear- 
oblong; achenes nearly glabrous; pappus white. 


Iowa and Minnesota to Manitoba, Saskatchewan 
and Wyoming. June—July. 


Senecio ligens Richards., of northwestern North 
America, admitted in our first edition, is not defi- 
nitely known to occur within our area. 


GENUS 103. THISTLE FAMILY. 547 


103. ARCTIUM L. Sp. Pl. 816. 1753. 


Large coarse branching, rough or canescent, mostly biennial herbs, with broad alternate 
petioled leaves, and rather large heads of purple or white perfect tubular flowers, racemose, 
corymbose or paniculate at the ends of the branches. Involucre subglobose, its bracts rigid, 
lanceolate, tipped with spreading or erect hooked bristles, imbricated in many series. Recep- 
tacle flat, densely bristly. Corollas tubular with 5-cleft limbs. Filaments glabrous. Anthers 
sagittate at the base. Achenes oblong, somewhat compressed and 3-angled, ribbed, truncate. 
Pappus of numerous short serrulate scales. [Greek, bear, from the rough involucre. | 


About 6 species, natives of Europe and Asia, readily distributed, their burs adhering to animals. 
Type species: Arctium Lappa L. 


Bracts of the involucre densely cottony ; heads corymbose. 1. A. ijomentosum. 
Bracts of the involucre glabrous, or slightly woolly. 
Involucre 1’ broad or more; inner bracts equalling or exceeding the flowers. 2. A. Lappa. 
Involucre’6”—9” broad; inner bracts not exceeding the flowers. 3. A. minus, 


1. Arctium tomentosum (Lam.) Schk. 
Woolly or Cottony Burdock. Fig. 4633. 


Lappa tomentosa Lam. Encycl. 1: 377. 1783. 
Arctium tomentosum Schk. Bot. Handb. 3: 49. 1803. 


- Arctium Lappa var. tomentosum A. Gray, Syn. Fl. 1: 
Part 2, 397. 1884. 


Similar to the following species; heads 8-10” 
broad, corymbose at the ends of the branches, mostly 
long-peduncled; bracts of the involucre densely cot- 
tony, the inner ones erect and somewhat shorter 
than the flowers. 


In waste places, Massachusetts to southern New York. 
Adventive from Europe. July—Aug. 


2. Arctium Lappa L. Great Bur, Burdock or Clotbur. Fig. 4634. 


Arctium Lappa L. Sp. Pl. 816. 1753. 
Lappa major Gaertn. Fruct. & Sem. 2: 379. pl. 162. 1802. 


Stem much branched, 4°-9° high. Leaves thin, broadly 
Ovate, pale and tomentose-canescent beneath, obtuse, 
entire, repand or dentate, mostly cordate, the lower 
often 18’ long; petioles solid, deeply furrowed; heads 
clustered or subcorymbose, sometimes long-peduncled, 
1-14’ broad; bracts of the involucre glabrous or nearly 
so, their spines all spreading, the inner ones equalling 
or exceeding the flowers; corolla-tube longer than the 
limb. 


In waste places, New Brunswick and Ontario to southern 
New York, and locally in the interior. Not nearly as com- 
mon as the next species in the Middle States. Naturalized 
from Europe. Other names are cockle-bur, cockle-button, 
cuckold-dock, hurr-bur, stick-button, hardock, bardane, beg- 
gar’s-buttons. July—Oct. 


548 COMPOSITAE. Vor. III. 


3. Arctium minus Schk. Common Burdock. 
Fig. 4635. 


Arctium minus Schk. Bot. Handb. 3: 49. 1803. 
Lappa minor DC. Fl. Fran. 4: 77. 1805. 


Arctium Lappa var. minus A. Gray, Syn. Fl. 1: Part 2, 
397. 1884. 


Smaller than the preceding species, seldom over 
5° high. Leaves similar, the lower deeply cordate; 
petioles hollow, not deeply furrowed; heads numer- 
ous, racemose on the branches, short-peduncled or 
sessile, 6-9” broad; bracts of the involucre glabrous 
or slightly cottony, the spines of the outer ones 
spreading, those of the inner erect and shorter than 
the flowers; corolla-tube about as long as the limb. 


In waste places, common nearly throughout our area, 
extending west to Colorado. Naturalized from Europe. 
Called also cuckoo-button. Leaves rarely laciniate or 
pinnatifid. July—Nov. 


x Meee 
104. CIRSIUM [Tourn.] Mill. Gard. Dict. Abr. Ed. 4. 1754. 


Erect, branching or simple, prickly herbs, some species acaulescent, with alternate or 
basal, sinuate-dentate, lobed or pinnatifid, usually very spiny leaves, sometimes decurrent, and 
large, many-flowered, solitary or clustered, discoid heads of purple, yellow or white, tubular, 
perfect and fertile, or rarely dioecious flowers. Involucre ovoid or globose, its bracts prickle- 
tipped or unarmed, imbricated in many series. Receptacle flat or convex, bristly. Corolla- 
tube slender, the limb deeply 5-cleft. Filaments pilose, or rarely glabrous. Anthers sagit- 
tate at the base. Style-branches short or elongated, obtuse. Achenes obovate or oblong, 
compressed or obtusely 4-angled, glabrous, smooth or ribbed. Pappus of several series of 
slender, plumose bristles, connate at the base. [Greek, referring to the use of the thistle as 
a remedy for swollen veins.] 


Over 200 species, widely distributed in the northern hemisphere. Besides the following, some 
50 others occur in the southern and western parts of North America and many hybrids have been 
described. Type species: Carduus heterophyllus L. 


+ Outer involucral bracts, or all of them, strongly prickly-pointed. 
1. Leaves glabrous or hispid above, tomentose beneath. 


All the bracts of the involucre tipped with prickles ; naturalized weed. 1. C. lanceolatum. 
Outer bracts prickle-tipped, the inner merely acuminate ; native species. 
Branches leafy up to the heads; involucral bracts firm or rigid. 


Leaves undivided, lobed or dentate, rarely pinnatifid. 2. C. altissimum. 
Leaves deeply pinnatifid into lanceolate or linear segments. 3. C. discolor. 
Heads naked-peduncled, 1’ high: involucral bracts thin. A 4. C. virginianum. 
2. Leaves tomentose on both sides, or becoming glabrous above; western. 
Leaves pinnately parted; segments linear, entire or lobed. 5. C. Pitcheri. 


Leaves pinnatifid into triangular or lanceolate dentate segments. 
Outer bracts with spines less than one-half their length. 
Leaf-lobes triangular ; flowers pink or purple. 
Leaf-lobes linear-lanceolate to oblong. 


fon 


. C.undulatum. 


Flowers yellow or cream-color. 7. C. plattense. 
Flowers purple. : : 8. C. Flodmani. 
Outer bracts with spines of nearly or quite their length. 9g. C. ochrocentrum. 
Leaves entire or undulate; outer pappus-bristles barbellate. 10. C. nebraskense. 
3. Leaves green both sides, somewhat pubescent beneath. 
Leaf-lobes acute; bracts of the involucre faintly nerved ; roots solid. TreGe odoratum. 
Leaf-lobes blunt ; bracts with prominent glutinous midnerve ; root hollow. 12. C. Milli. 


++ Bracts of the involucre not at all prickly-pointed, or scarcely so. 
Heads large, few, 1-4’ broad; flowers all perfect and fertile. 


Heads involucrate by the upper very spiny leaves; flowers usually yellow. 13. C. horridulum. 

Heads peduncled, naked, or with 1 or 2 bracts at the base; flowers purple. 14. C. muticum. 
Heads small, numerous, 1’ or less broad. 

Heads partly dioecious ; leaves not decurrent. 15. C. arvense. 

Heads not dioecious; leaf-bases decurrent. 16. C. palustre. 


GENUS 104. THISTLE FAMILY. 


A fe 
WAVE 
Si Salah 
SUES 


= 


1. Cirsium lanceolatum (L.) Hill. Common 
Bur or Spear Thistle. Fig. 4636. 
Carduus lanceolatus L. Sp. Pl. 821. 1753. 
Cirsium lanceolatum Hill, Herb. Brit. 1: 80. 1769. 
Cnicus lanceolatus Willd. Prodr. Fl. Berol. 259. 1787. 
Biennial; stem stout, branched, more or less to- 
mentose, 3°-5° high, leafy to the heads. Leaves dark 
green, lanceolate, acuminate, deeply pinnatifid, 3-6’ W' 
long, or the lowest larger, decurrent on the stem and 
branches, the lobes triangular-lanceolate, tipped with 
stout prickles, the margins and decurrent bases 
bristly, the upper surface strigose-pubescent or his- 
pid, the lower brown-tomentose and midnerve pilose, 
especially when young; heads mostly solitary at the 
ends of the branches, 14’-2’ broad, 14’-2’ high; bracts 3 
of the involucre cottony, narrowly lanceolate, acumi- ; MN 
nate, all tipped with slender, erect or ascending \\ 


prickles; flowers dark purple. = 
In fields and waste places, Newfoundland to Georgia, CRO 
Minnesota, Nebraska, Oregon and California. Natural- : 


“Al 


g 


eA 
ae 
RA\ 
cel 4 
AL 
NU Lf ys 


Se 
=<, Si 
i 


i 
i i Mf 53 
: iM | 


rn|— 


ized from Europe. Native also of Asia. Plume-, bank- 
or horse-thistle, bell-, bird-, blue-, button-, boar-, bull- or 
roadside-thistle. July—Nov. 


2. Cirsium altissimum (L.) Spreng. Tall or Roadside Thistle. Fig. 4637. 
Carduuse altissimus L. Sp. Pl. 824. 1753. 
Cnicus altissimus Willd. Sp. Pl. 3: 1671. 1804. 
Cirsium altissimum Spreng. Syst. 3: 373. 1826. 

Biennial or perennial; roots often thickened; 
stem pubescent or tomentose, stout, branched, leafy 
to the heads, 3°-10° high. Leaves ovate-oblong or 
oblong-lanceolate, sessile or slightly clasping, spar- 
ingly pubescent above, densely white-tomentose be- 
neath, scarcely or not at all decurrent, acute, spinu- 
lose-margined, entire, dentate with bristle-pointed 
teeth or lobed, sometimes pinnatifid into oblong or 
triangular-lanceolate segments, the lowest some- 
times 8’ long, narrowed into margined petioles, the 
uppermost linear or lanceolate, much smaller; 
heads about 2’ broad, 12’-2’ high, mostly solitary at 
the ends of the branches; outer bracts of the invo- 
lucre ovate or ovate-lanceolate, firm with a dark, 
wightly glandular spot or band on the beak, tipped 
with short prickles, the inner linear-lanceolate, 
acuminate, unarmed; flowers light purple. 

In fields and thickets, Massachusetts to Ohio, Min- 
nesota, Florida, Nebraska and Texas. Aug.—Sept. 

Cirsium iowénse Pammel, with slightly larger heads 
and longer-tipped inner involucral bracts, appears to 
be a northwestern race of this species. 


3. Cirsium discolor (Muhl.) Spreng. Field \0 ly 
Thistle. Fig. 4638. WN ANY 
>|) 


\ 
Cnicus discolor Muhl.; Willd. Sp. Pl. 3: 1670. 1804 W AN / 
ae . Sp. Pl. 3: 1670. ‘ Bs\\\ | 


Carduus discolor Nutt. Gen. 2: 130. 1818. SS. WwW 
Cirsium discolor Spreng. Syst. 3: 373. 1826. é AN \ 


Similar to the preceding species, but lower and \\ 
more leafy, seldom over 7° high. Leaves deeply pin- \ 
natifid into linear, linear-lanceolate or falcate, prickly ~2— 
toothed segments, white tomentose beneath, sessile, 
the basal ones sometimes 12’ long; heads 13-2’ broad, 
about 14’ high, usually involucrate by the upper 
leaves, mostly solitary at the ends of the branches; NY 
outer bracts of the involucre coriaceous, ovate, SN 
slightly woolly, tipped with slender bristles, which 
are longer than those of the preceding species; inner 
bracts lanceolate, acuminate, unarmed; flowers light 
purple or pink, rarely white. 


In fields and along roadsides, New Brunswick to On- ff 
tario, Georgia, Minnesota, Nebraska and Missouri. July— 
Nov. 


55° < COMPOSITAE. Vor. IT. 


——, 


lei 4. Cirsium virginianum (L.) Michx. Vir- 
ginia Thistle. Fig. 4639. 
Carduus virginianus L. Sp. Pl. 824. 1753. 


y 
WZ 
ZGEZ 


SSaN K Cirsium virginianum Michx. Fl. Bor. Am.2:90. 1803. 
Gx Cnicus virginianus Pursh, Fl. Am. Sept. 506. 1814. 
Y] WY Biennial; stem slender, naked or scaly above, 


SS - y | pubescent or somewhat tomentose, simple or 

branched, 2°-33° high. Leaves oblong, oblong- 
lanceolate, or the lowest slightly spatulate, ses- 
sile, or somewhat clasping, not decurrent, acute 
or acutish, spinulose-margined, entire, lobed or 
pinnatifid into triangular-lanceolate lobes, the 
lower sometimes 8’ long and 2’ wide, narrowed 
into margined petioles, all pubescent or glabrate 
above, and densely white-tomentose beneath; 
heads long-peduncled, 1-13’ broad, about 1’ high; 
outer bracts of the involucre not coriaceous, lan- 
ceolate or ovate-lanceolate, tipped with weak short 
bristles, the inner ones very narrow and merely 


f} Wy 
Vj aN Wy acuminate; flowers purple. 
Ii | ff py) In dry woods aed thickets, Virginia to Kentucky, 
\ 7 cs w Ohio, Florida and Texas. April—Sept. 


5. Cirsium Pitcheri (Torr.) T. & G. Pitcher’s Thistle. Fig. 4640. 


Cnicus Pitcheri Torr.; Eaton, Man. Ed. 5, 180. 1829. ~~ 
Cirsium Pitcheri T. & G. Fl. N. A. 2: 456. 1843. 
Carduus Pitcheri Porter, Mem. Torr. Club 5: 345. 1804. 


Biennial, persistently white-tomentose throughout ; 
stem stout, leafy up to the heads, usually branched, 
1°-2° high. Leaves sessile, partly clasping or slightly Y Uy 
decurrent, pinnately divided into narrowly linear, Y) 
entire lobed or pinnatifid, acute sparingly prickly seg- \ HH} 
ments, 2”—-3” wide, with revolute margins; basal \ WG 
leaves often 12’ long; heads solitary or several and \ j Yj 
racemose-spicate at the ends of the branches, about xaN | eo 
12 broad; outer bracts of the involucre ovate-lan- 
ceolate, sparingly pubescent and tomentose-ciliate, 
glutinous on the back, tipped with short spreading 
bristles, the inner narrowly lanceolate, acuminate or 
sometimes tipped with weak prickles; flowers cream 
color. 


Shores of Lakes Michigan, Huron and Superior. June— 


6. Cirsium undulatum ( Nutt.) Spreng. 
Wavy-leaved Thistle. Fig. 4641. 


Carduus undulatus Nutt. Gen. 2: 130. 1818. 

Cirsium undulatum Spreng. Syst. 3: 374. 1826. 

Cnicus undulatus A. Gray, Proc. Am. Acad. 10: 42. 1874. 

Cnicus undulatus var. megacephalus A. Gray, Proc. Am. 
Acad. 10: 42. 1874. 

Carduus undulatus megacephalus Porter, Mem. Torr. 
Club 5: 345. 1894. 


Biennial, persistently and densely white-tomentose 
throughout, or the upper surfaces of the leaves at 
length green and glabrous; stem stout, leafy, usually 
branched, 1°-3° high. Leaves lanceolate or oblong- 
lanceolate in outline, acute, sessile or decurrent, or 
the lowest petioled, undulate, lobed or pinnatifid, the 
lobes dentate, triangular, often very prickly; basal 
leaves often 8’ long; heads 13’—3’ broad, and nearly 
as high, solitary at the ends of the branches; outer 
bracts of the involucre ovate or ovate-lanceolate, 
firm, glutinous on the back, tipped with short spread- 
ing prickles, the inner ones lanceolate, acuminate; 
flowers purple or pink. 


On plains and prairies, Lake Huron to Assiniboia, Alberta, Kansas, New Mexico and Arizona. 
June-Sept. 


ee Oe ee eee ee ee ee eee ee ke Pee, core fe ee ee ee 


et SPA Sea Ot 


whee 


pe Mate oboe ee 


i 


ea Ted be 3) 


=n 
ok 


(Jes “a! ir 


ae 
— 


GENUS I04. THISTLE FAMILY. 551 


7. Cirsium platténse (Rydb.) Britton. 
Prairie Thistle. Fig. 4642. 


Carduus plattensis Rydberg, Contr. Nat. Herb. 3: 167. 
pl. 2. 1895. $ 


Perennial or biennial, the root thick and deep; 
stem stout, simple, or little branched, 12°-22° tall, 
densely white-felted. Leaves deeply pinnatifid, 
white-tomentose beneath, green, loosely tomen- 
tose, or glabrate above, the lower 5-7’ long, the 
lobes lanceolate to oblong, acute, prickly tipped 
and margined; upper leaves smaller and less di- 
vided; heads few, about 2’ high and broad; outer 
bracts of the involucre lanceolate to ovate-lanceo- 
late, firm, dark, tipped with a short weak spread- 
ing prickle, the inner linear-lanceolate, unarmed, 
tipped with a scarious reflexed erose appendage; 
corolla yellow, its lobes linear; pappus of outer 
flowers merely barbellate. 


Sand hills, Nebraska, Colorado and South Dakota. 
May-July. 


8. Cirsium Fl6dmani (Rydb.) Britton. Flod- 
man’s Thistle. Fig. 4643. 


Carduus Flodmani Rydb. Mem. N. Y. Bot. Gard. 1: 451. 
1900. 


Stem rather slender, 13°-3° tall, loosely white-cottony, 
usually more or less branched. Leaves deeply pinnatifid 
into linear-oblong or lanceolate, acute or acuminate, 
toothed or entire segments, floccose and green above, 
densely white-cottony beneath, the lower 6’ long or less; 
heads 14’-2’ broad; involucre campanulate, its linear 
bracts tipped with yellow prickles; flowers reddish- 
purple to rose. 


Meadows and river bottoms, Iowa and North Dakota to 
Saskatchewan, Nebraska and Colorado. Has been referred 
to the western C. canescens. July—Sept. 


g. Cirsium ochrocéntrum A. Gray. 
Yellow-spined Thistle. Fig. 4644. 


Cirsium ochrocentrum A. Gray, Mem. Am. Acad. 
Bowron » Lod 0. 
Cnicus ochrocentrus A. Gray, Proc. Am. Acad. 19: 


57- 1883. 
Carduus ochrocentrus Greene, Proc. Phil. Acad. 
1892: 336. 1893. 


Similar to Cirsium undulatum, but commonly 
taller and more leafy, often 6° high, equally 
white-tomentose. Leaves oblong-lanceolate in 
outline, usually very deeply pinnatifid into tri- 
angular-lanceolate, serrate or entire segments, 
armed with numerous long yellow prickles; 
lower leaves often 6-8’ long; heads about 2’ 
broad, 13’-2’ high, solitary at the ends of the 
branches; outer bracts of the involucre lan- 
ceolate; tipped with stout yellow prickles of 
nearly or quite their own length, the inner nar- 
rowly lanceolate, long-acuminate; flowers pur- 
ple (rarely white?). 

On plains, Nebraska to Texas, Nevada and Ari- 
zona. May-—Sept. 


ap? 


MMi 
A WO AYA 
vee 
GCA 
SN 
AN Wa 
\ 


COMPOSITAE. Vot. III. 


10. Cirsium nebraskénse Britton. Nebraska 
Thistle. Fig. 4645. 


Carduus nebraskensis Britton, in Britt. & Brown, Ill. Fl. 
3: 487. 1898. 


Stem densely white-woolly, apparently over 1° high. 
Leaves linear-oblong to lanceolate, white-woolly be- 
neath, green and sparingly loosely woolly above, ir- 
regularly slightly toothed or entire, the upper 3-6’ 
long, #’-1’ wide, the margins prickly; heads solitary, 
or few, short-peduncled, about 13’ high; outer bracts 
of the involucre lanceolate, prickle-tipped, the inner 
narrower with a reflexed acute scarious appendage; 
pappus bristles of inner flowers plumose, of the outer 
barbellate. 


Western Nebraska and Wyoming. Summer, 


Pasture Thistle. Fragrant Thistle. 
Fig. 4646. 


Cnicus odoratus Muhl. Cat. 70. 1813. 

Carduus pumilus Nutt. Gen. 2: 130. 1818. 

Cnicus pumilus Torr. Compend. 282. 1826. 

Carduus odoratus Porter, Mem. Torr. Club 5: 345. 
1894. 


Biennial, more or less villous-pubescent; stem 
stout, simple or branched, leafy, 1°-3° high; roots 
thick, branched, solid; stem leaves green both sides, 
sessile and clasping, oblong or oblong-lanceolate in 
outline, acute, 3-7’ long, 1-2’ wide, pinnatifid into 
triangular, acute, dentate, prickly lobes. Basal 
leaves petioled; heads solitary, terminal, 2’—3’ 
broad, about 2’ high, often involucrate by the 
upper leaves; outer bracts lanceolate or ovate- 
lanceolate, with a slight glutinous strip on the 
back, glabrous or sparingly tomentose, tipped with 
slender prickles, the inner narrow, long-acuminate; 
flowers purple, rarely white, fragrant; tips of the 
pappus bristles usually spatulate. 


1 My Y 
wy 
wy zz 
AG — 


{ 


In fields, Maine to Pennsylvania, Delaware and 


West Virginia. July—Sept. 
\Salah 
WS MM Wy 
WY) Ge 
Si VA 
<= Ni g 


ti] WZ 


12. Cirsium Hillii (Canby) Fernald. 
Hill’s Thistle. Fig. 4647. 


Cnicus Hillii Canby, Gard. & For. 4: 101. 1891. 
Carduus Hillii Porter, Mem. Torr. Club 5: 344. 1894. 
Cirsium Hillii Fernald, Rhodora 10: 95. 1908. 


Perennial, low, villous-pubescent or somewhat 
woolly; stem leafy, simple or branched, 1°-2° 
high; root perpendicular, fusiform, slender and 
hollow above, enlarged below, 8-12’ long; leaves 
green both sides, mostly obtuse, lobed or pin- 
natifid, the lobes mostly broad and rounded, den- 
tate, spinulose or with some rather stout prickles, 
the upper oblong, sessile and clasping, the lower 
spatulate-oblong, narrowed at the base or the 
lowest ones petioled and 6’-& long; heads 23° 
broad, about 2’ high; outer bracts of the invo- 
lucre ovate-lanceolate, tipped with short bristles, 
conspicuously glutinous on the back, the inner 
narrowly lanceolate, long-acuminate; flowers pur- 
ple; pappus bristles slender-pointed or some of 
them slightly spatulate. 


In fields, western Ontario to Minnesota, south to 
Pennsylvania, Illinois and Iowa. June-July. 


j 
math oe Odes) 


a 


Pe ee eS Le 


a a ee ee a a Tr 


GENUS 104. THISTLE FAMILY. 553 


13. Cirsium horridulum Michx. Yellow Thistle. 
Fig. 4648. 


Carduus spinosissimus Walt. Fl. Car. 194. 1788. Not Cir- 
sium _spinosissimum (L.) Scop. 

Cirsium horridulum Michx. Fl. Bor. Am. 2: 90. 1803. 

Cnicus horridulus Pursh, Fl. Am. Sept. 507. 1814. 


Biennial or perennial, somewhat woolly when young, 
but becoming glabrate; stem branched, leafy, 2°-5° 
high. Leaves green both sides, lanceolate or oblong in 
outline, sessile and clasping or the basal ones short- 
petioled and somewhat spatulate, pinnatifid into trian- 
gular or broader, spinulose-margined and prickle-tipped, 
entire or dentate lobes; heads involucrate by the upper 
leaves, 2’-4’ broad, 14’-22’ high; bracts of the involucre 
narrowly lanceolate, roughish and ciliate, long-acumi- 
nate, unarmed; flowers pale yellow, yellowish, or occa- 
sionally purple. 


__In moist or dry sandy soil, Maine to Pennsylvania, Flor- 
ida and Texas. Abundant along the edges of salt-meadows 
= New York and New Jersey. May—Aug., or earlier in the 

outh. 


14. Cirsium muticum Michx. Swamp 
Thistle. Fig. 4649. 
Cirsium muticum Michx. Fl. Bor. Am. 2: 89. 1803. 
Carduus muticus Pers. Syn. 2: 386. 1807. 
Cnicus muticus Pursh, Fl. Am. Sept. 506. 1814. 
Carduus muticus subpinnatifidus Britton, in Britt. 
& Brown, Ill. Fl. 3: 489. 1808. 
Grice muticus alpicola Fernald, Ott. Nat. 1905: 
I 
Biennial; stem woolly or villous when young, 
becoming glabrate, slender, striate, leafy, panicu- 
lately branched above, 3°-8° high. Leaves dense- 
ly white-tomentose beneath when young, some- 
times becoming glabrous on both sides, deeply 
pinnatifid into lanceolate or oblong, entire, lobed 
or dentate, spiny segments usually tipped with 
slender prickles, or sometimes merely lobed; 
basal leaves petioled, 4-8’ long, those of the 
f stem sessile and smaller; heads about 13’ broad 
| and high, solitary, terminal, naked-peduncled, 
Dy or with a few small bract-like leaves near the 
7 \ base; outer bracts viscid, appressed, more or 
less cottony, ovate or ovate-lanceolate, the in- 
ner linear-lanceolate, acute, all unarmed; flowers purple. 
In swamps and moist soil, Newfoundland to Florida, Saskatchewan and Texas. July—Oct. 


15. Cirsium arvénse (L.) Scop. Canada Thistle. Creeping Thistle. Fig. 4650. 


Serratula arvensis L. Sp. Pl. 820. 1753. 

Cirsium arvense Scop. Fl. Carn. Ed. 2, 2: 126. 1772. 

Carduus arvensis Robs. Brit. Fl. 163. 1777. 

Cnicus arvensis Hoffm. Deutsch. Fl. Ed. 2, 1: Part. 2, 
130. 1804. 


Perennial by horizontal rootstocks, forming patches, 
nearly glabrous, or the leaves sometimes woolly be- 
neath; stems striate, 1°-3° high, branched above. 
Leaves sessile, slightly clasping, but not decurrent, 
lanceolate or oblong-lanceolate, deeply pinnatifid into 
very prickly, lobed or dentate segments, or sometimes 
nearly or quite entire; basal leaves sometimes peti- 
oled, 5’-8’ long; heads numerous, corymbose, dioe- 
cious, I’ broad or less, nearly 1’ high, purple or white, 
staminate heads globose, corollas projecting; pistil- 
late heads oblong-campanulate, corollas shorter, the 
long pappus conspicuous; outer bracts ovate or ovate- 
lanceolate, appressed, tipped with short prickly points, 
inner bracts of the pistillate heads linear, elongated. 

Tn fields and waste places, Newfoundland to Virginia, 
British Columbia, Nebraska and Utah. In many places 
a pernicious weed. Races differ in leaf-form and in 
pubescence. Naturalized from Europe. Way- or cursed 
thistle. Corn-, hard- or prickly-thistle. June—Sept. 


554 COMPOSITAE. Vor. III, 


16. Cirsium palustre (L.) Scop. Marsh Thistle. 
Fig. 4651. 


Carduus palustris L. Sp. Pl. 822. 1753. Y 
Cirsium palustre Scop. Fl. Carn. Ed. 2, 2: 128. 1772, : i 


Annual or biennial; stem little branched, 4°-5° high, 
loosely floccose or glabrate and covered by the decur- 
rent prickly margins of the leaves. Leaves pinnatifid, 
the lower often 6-8’ long, linear-oblong in outline, the 
segments lobed, loosely floccose beneath, spinulose; 
heads usually many, rather less than 1’ broad, densely 
clustered, short-peduncled, the involucre ovoid, its bracts 
with very short, prickly tips. 


f ee | wos 


Woodlands, East Andover, New Hampshire, recorded as 
thoroughly naturalized. Native of Europe and northern 
Asia. Summer. 


Cirsium canum (L.) Bieb., with larger, long-peduncled 
heads, the decurrent leaf-bases merely ciliate, is recorded as 
established in Massachusetts. Adventive from Europe. 


ee Te Ee ie See 


105. CARDUUS [Vaill.] L. Sp. Pl. 820. 1753. 


Herbs resembling Cirsium in habit, usually annual or biennial, the leaves decurrent on 
the stem and branches as spiny wings, the heads often nodding. Involucre ovoid to globose, 
many-flowered, its bracts narrow, in many series. Receptacle copiously bristly, flat or convex. . 
Corolla-tube slender, the limb deeply 5-cleft. Filaments papillose-pubescent. Anthers sagit- 
tate at the base and with slender auricular appendages. Style-branches obtuse. Achenes 
mostly obovoid, sometimes angled or ribbed, glabrous. Pappus of many naked or merely 
roughened bristles. [Ancient Latin name of these plants. | 


About 80 species, natives of the Old World. Type species: Carduus nutans L. 


Heads solitary at end of stem or branches, nodding. 1. C. nutans. 
Heads usually several, crowded at ends of winged branches. 2. C. crispus: 


1. Carduus nitansL. Musk Thistle. Plumeless Thistle. Fig. 4652. 


Carduus nutans L. Sp. Pl. 821. 1753. 


Biennial, branched, sparingly tomentose, 2°-3° 
high. Leaves lanceolate in outline, deeply pinnatifid, 
acuminate, 3-6’ long, the lobes triangular, very 
prickly; heads long-peduncled, solitary at the end 
of the stem or branches, 12’-23’ broad, nodding, pur- 
ple, rarely white, fragrant; involucre hemispheric, its 


OC a ee ee ee ee ee 


. . . i 
bracts in many series, lanceolate, long-acuminate, the ; 
prominent mid-nerve prolonged into a prickle, or the re 
° . , - 
inner nerveless and awned; pappus bristles 10’—-1’ 
long, white, very minutely barbed. a 

* 

In waste places, District of Columbia, Pennsylvania ‘ 
and New Jersey to New Brunswick, and in ballast about 'y 
the seaports. Naturalized or adventive from Europe. § 
Native also of Asia. Bank- or buck-thistle. Queen Ann’s- H) { 
thistle. July—Oct. fi 1 


GENUS 105. THISTLE FAMILY. 


2. Carduus crispus L. Curled Thistle. Welted 
Thistle. Fig. 4653. 
Carduus crispus L. Sp. Pl. 821. 1753. 

Biennial, somewhat tomentose; stem much branch- 
ed, densely prickly, 2°-4° high. Leaves lanceolate 
in outline, with undulate and ciliate-spiny margins, 
all sinuate-pinnatifid into broad, 3-lobed, toothed 
segments, the teeth prickle-pointed; heads several, 
usually crowded at the ends of the winged branches, 
1’ broad or less, purple or white, sessile or short- 
peduncled, or some of them rarely solitary and 
slender-peduncled; involucre ovoid, its bracts very 
numerous, linear, the outer prickle-tipped and rigid, 
the inner thinner and merely acuminate. 

In fields and waste places, New Brunswick, Quebec, 
Nova Scotia and Pennsylvania, and in ballast about the 
seaports. Adventive from Europe. Native also of Asia. \ 
July—Sept. 

Carduus acanthoides L., which resembles this species, 
but has larger hemispheric involucres, their outer bracts 
not rigid, has been collected in ballast on waste grounds 
about Atlantic seaports, and several other species of 
Carduus have been found in ballast deposits. 


106. MARIANA Hill, Veg. Syst. 4: 19. 1762. 
[Sirysum Vaill.; Adans. Fam. Pl. 2: 116. 1763.] 


Annual or biennial, simple or branched, nearly glabrous herbs, with large alternate clasp- 
ing, sinuate-lobed or pinnatifid, white-blotched leaves, and large discoid heads of purple 
tubular flowers, solitary at the end of the stem or branches. Involucre broad, subglobose, 
its bracts rigid, imbricated in many series, the lower ones fimbriate-spinulose at the broad 
triangular summit, the middle ones similar but armed with huge spreading or recurved spines, 
the inner lanceolate, acuminate. Receptacle flat, densely bristly. Corolla-tube slender, the 
limb expanded and deeply 5-cleft. Filaments monadelphous below, glabrous. Anthers sag- 
ittate at the base. Style nearly entire. Achenes obovate-oblong, compressed, glabrous, sur- 
mounted by a papillose ring. Pappus bristles in several series, flattish, barbellate or scabrous. 
[St. Mary’s thistle. ] 


A montypic genus of the Mediterranean region. 


1. Mariana mariana (L.) Hill. Milk 
Thistle. Fig. 4654. 


Carduus marianus L. Sp. Pl. 823. 1753. 

Mariana mariana Hill, Hort. Kew. 61. 17609, 

Silybum marianum Gaertn. Fruct. & Sem. 2: 378. 
1802. 


Stem striate, glabrous or slightly woolly, little 
branched, 2°-4° high. Leaves oblong-lanceolate, 


YX 
Ss prickly, strongly clasping, the lower often 12’ 
a] & long and 6’ wide, the upper much smaller, scarce- 


7 ly lobed, acute; heads about 23’ broad; spines of 
Vi 


WOAE SNE 


if 

UN 

K SS 

Eee aN Z ‘ the middle involucral bracts often 13’ long; pap- 
poZa pus bristles white, barbellate. 

Vg LAS Escaped from gardens near Kensington, Ontario 

~ waste grounds about the eastern seaports, south to 


uw (T. Walker, according to Macoun), in ballast and 
Si ] Bs N Alabama, and on the Pacific Coast from British Co- 


( 


| A\ \ lumbia to southern California, where it is naturalized. 


i 


(é 
107. ONOPORDON [YVaill.] L. Sp. Pl. 827.1753. 

Coarse, branching or rarely acaulescent, tomentose herbs, with stout stems winged by 
the decurrent bases of the alternate dentate or pinnatifid, prickly leaves, and large discoid 
heads of purple violet or white flowers, mostly solitary at the ends of the branches. Invo- 
lucre nearly globular, its bracts imbricated in many series, all tipped with long spines in our 
species, the inner narrower than the outer. Receptacle flat, fleshy, honeycombed, not bristly. 


556 COMPOSITAE. Vor. II. 


Corolla-tube slender, the limb expanded and deeply 5-cleft. Filaments pilose. Anthers sagit- 
tate at the base. Achenes obovate or oblong, 4-angled or compressed, smooth or corrugated. 
Pappus bristles in several series, filiform, barbellate or plumose, united at the base. [Greek, 
Asses’ thistle, the ancient name. ] 

About 12 species, natives of the Old World, the following typical. 


1. Onopordon Acanthium L. Cotton 
Thistle. Scotch Thistle. Fig. 4655. 


Onopordon Acanthium L. Sp. Pl. 827. 1753. 
{ 

Biennial, white-tomentose all over; stem usually 
much branched, leafy, 3°-0° high. Leaves oblong, 
lobed and dentate, acute, very spiny, the lower 
often 12’ long; heads 14’-2’ broad, about 13’ high, 
solitary at the ends of the branches; outer bracts 
of the involucre ovate or oblong, minutely serru- 
late, tipped with long stout spreading spines; 
flowers pale purple; achenes slightly corrugated; 
pappus bristles brownish, longer than the achene. 


In waste places, Nova Scotia and Ontario to New 
Jersey, Pennsylvania and Michigan. Naturalized 
from Europe. Native also of Asia. Argentine. Asses’, 
oat or down-thistle. Queen Mary’s-, silver- or musk- 
thistle. July—Sept. 


108. CENTAUREA L. Sp. Pl. 909. 1753. 


Perennial or annual herbs, with alternate entire dentate incised or pinnatifid leaves, and 
large or middle-sized heads, of tubular purple violet white or rarely yellow flowers. Invo- 
lucre ovoid or globose, its bracts imbricated in many series, appressed, fimbrillate, or dentate. 
Receptacle flat, densely bristly. Marginal flowers usually neutral and larger than the central 
ones, which are perfect and fertile, or flowers all perfect and fertile in some species. Corolla- 
tube slender, the limb regular or oblique, 5-cleft or 5-lobed, the segments sometimes appearing 
like rays. Anthers sagittate at the base. Style-branches short, somewhat connate, obtuse. 
Achenes oblong or obovoid, compressed or obtusely 4-angled, usually smooth and shining, 
obliquely or laterally attached to the receptacle, surmounted by a disk with an elevated mar- 
gin. Pappus of several series of bristles or scales, rarely none. [Greek, of the Centaurs, 
who were said to use it in healing. ] 


About 350 species, mostly natives of the Old World. Type species: Centaurea Centaurium L. 


Bracts of the involucre lacerate or fimbriate, not spiny. _ : 
Heads 2’ broad or less; achenes laterally attached ; introduced species. 


Annual ; leaves entire. 1. C. Cyanus. 
Perennials or biennials, or C. maculosa annual. 
Bracts of the involucre laciniate or entire. 2. (Gaiiacea: 


Bracts of the involucre, or their tips, pectinate-fringed. } 
Lower bracts of the involucre pectinate-fringed to below the middle. F 
Leaves entire or merely dentate. 3. C. nigra. 
Leaves pinnatifid. : 4. C. Scabiosa. 
Lower bracts of the involucre pectinate-fringed only at the tips. ; F 
Leaves entire, toothed, or the lower lyrate. 5. C. vochinensis. 
All but the upper leaves pinnatifid into linear segments. 6. C. maculosa. 
Heads 2’—4’ broad; achenes obliquely attached ; native western species. 7. C. americana. 
Bracts of the involucre tipped with stout spines. 
Flowers purple; stem wingless. 8 
Flowers yellow; stem winged by the decurrent leaf-bases. : ; 
Spines slender, purplish, 5” long or less, branched below. 9. C. melitensis. 
Spines stout, yellow, 6”-10” long, with smaller ones at the base. 10. C. solstitialis. 


. C. Calcitrapa. 


GENUS 108. THISTLEE-PAMILY. Sh 


1. Centaurea Cyanus L. Blue-bottle. 
Corn Blue-bottle. Corn-flower. 
Fig. 4656. 
Centaurea Cyanus L. Sp. Pl. 911. 1753. 


Annual, woolly, at least when young; stem 
leafy, slender, branched, 1°-23° high, the branches 
ascending. Leaves linear or linear-lanceolate, 
mucronate, 3-6’ long, the basal and lower ones 
mostly remotely dentate, the upper, or sometimes 
all of them, entire; heads 17-13’ broad, on long 
naked peduncles; involucre campanulate, its bracts 
greenish-yellow, or the inner with darker tips 
and margins, appressed, fimbriate with scarious 
teeth; flowers blue, purplish, pink or white, the 
marginal ones neutral with large radiant corolla- 
limbs; achenes slightly compressed, or 4-angled; 
pappus bristles unequal, nearly as long as the 
achene. 


In waste places, escaped from gardens, and in bal- 
last, Quebec to Ontario, western New York, Nebraska 
and Virginia. Witches’-bells or -thimbles, corn- 
centaury, corn-bottle or -binks. Brushes. Hurt-sickle. 
Blue-bonnets. Blaver. Blue poppy. Bachelor’s-but- 
tons. Bluecaps. Barbeau. French pink. July—Sept. 


2. Centaurea Jacea L. Brown or Rayed 
Knapweed. Fig. 4657. 
Centaurea Jacea L. Sp. Pl. 914. 1753. 


Perennial, 2° high or less. Leaves entire or 
denticulate, rarely lobed; heads 1-13’ broad; in- 
volucre globular to ovoid, its bracts closely im- 
bricated, brown or with brown backs, the outer 
ones pale brown with fimbriate appendages, the 
middle ones lacerate, the inner entire or nearly 
so, dark brown; marginal flowers neutral with 
enlarged radiant corolla-limbs; achenes obscurely 
4-sided; pappus none, or a minute crown. 

In waste places, northern New York, Vermont and 
Massachusetts, and in ballast about eastern seaports. 
Also in British Columbia. Fugitive from Europe 
June-Sept. 


3. Centaurea nigra L. Black Knapweed. 
Horse-knops. Hardheads. Centaury. 
Fig. 4658. 


Cen aurea nigra L. Sp. Pl. 911. 1753. 

Perennial, scabrous or pubescent; stem stiff, 
branched, 1°-2° high. Lower and basal leaves 
spatulate or oblong, acutish, entire, denticulate, \S 
dentate or lobed, not pinnatifid, 3-6’ long, nar- 
rowed into long petioles; upper leaves oblong or 
lanceolate, sessile, or partly clasping, entire or 
nearly so; heads rarely 1’ broad, bracted by the 
small, uppermost leaves; involucre globose, its 
bracts lanceolate or ovate-lanceolate, closely im- 
bricated, tipped with brown fimbriate appendages, 
or the uppermost merely lacerate; flowers rose- 
purple, all perfect, the marginal ones usually not 
at all enlarged, or sometimes radiant; achenes 
slightly 4-sided; pappus none, or a ring of minute 
scales. 

In waste places and fields, Newfoundland to On- 
tario, New Jersey and Pennsylvania. Naturalized 
from Europe. Among many other English names are 
iron-head or -weed, club-weed, matfelon, hurt-sickle, 
tassel, horse-knobs; crop-, knob-, bole- or button- 
weed; loggerheads, lady’s-cushion, blue-tops, hard- 
weed, bullweed, sweeps, bachelor’s-buttons. July- 
Sept. 


558 COMPOSITAE. 


5. Centaurea vochinensis Bernh. Tyrol Knap- 
weed. Fig. 4660. 


C. voechinensis Bernh.; Reichenb. Icon. Fl. Germ. 15: 15. 
1853. 


Perennial, roughish, branched, 2° high orless. Leaves 
firm in texture, the basal and lower ones dentate or 
lyrate, 3-5’ long, the upper lanceolate to oblong-lanceo- 
late, few-toothed or entire; heads about 12’ broad; in- 
volucre ovoid-cylindric, about 8” high, its faintly nerved 
bracts pectinate at the brownish tip, the lower short, 
ovate, the upper elongated; flowers rose-purple, the 
margined ones radiant. 


Waste grounds, Ontario to Massachusetts and southern 
New York. Naturalized from Europe. Aug.—Oct. 


J 


C. Scabiosa L. Sp. Pl. 913. 


6. Centaurea maculosa Lam. 


C. maculosa Lam. Encycl. 1: 669. 


branches ascending. 


Vor. III. 


4. Centaurea Scabiosa L. Scabious Knapweed. 


Greater Centaury. Fig. 4659. 


Slightly pubescent or villous, perennial; stem simple 
or branched, about 2° high. Leaves all pinnatifid, the 
lower and basal ones petioled, often 6’ long, the upper 
sessile and much smaller; heads about 2’ broad, on 
bracted peduncles 1-4’ long; involucre ovoid, its bracts 
all pectinate to or below the middle, and blackish-mar- 
gined, the outer ovate, the inner oblong; corolla purple, 
the outer ones enlarged and neutral; pappus of stiff 
bristles, about as long as the achene. 


Waste grounds and fields, Quebec and Ontario to Ohio. 
Naturalized from Europe. 


Spotted Knap- 
weed. Fig. 4661. 


83. 


Annual or biennial, loosely floccose-pubescent or 
glabrate, usually much branched, 2°-3° high, the stiff 
Leaves pinnatifid into linear 
segments, or the upper linear and entire, the lower 
up to 3’ long; heads peduncled, about 10” broad; 
involucre ovoid, its ribbed bracts pectinate only at 
the black tip, the inner ones longer than the outer 
and entire or merely erose; corollas white to purple, 
the marginal ones radiant. 


Waste grounds, Massachusetts to New Jersey and 


Pennsylvania. July—Aug. 


GENUS 108. THISTLE: FAMILY. 559 


7. Centaurea americana Nutt. American 
Star Thistle. Fig. 4662. 


C. americana Nutt. Journ. Phila. Acad. 2: 117. 1821. 


Plectocephaius americanus Don, in Sweet, Brit. FI. 
Gard. (11) pl. 51. 1831. 


Annual, roughish; stem stout, simple, or little 
branched, 2°-6° high. Leaves entire or denticu- 
late, the lower and basal ones spatulate or oblong, 
2’-5’ long, narrowed into petioles, the upper ob- 
long-lanceolate, sessile, mucronate; heads solitary 
_at the much thickened ends of the leafy stem or 
branches, very showy, 2-4 broad; involucre 
nearly hemispheric, its bracts ovate or lanceolate 
with conspicuously pectinate appendages; flowers 
pink or purple, the marginal ones with enlarged 
and radiant corolla-limbs; achenes somewhat 
compressed, obliquely attached at the base; pap- 
pus of copious unequal bristles longer than the 
achene. 


Dry plains, Missouri and Arkansas to Louisiana, 
Mexico and Arizona. May-—Aug. 


8. Centaurea Calcitrapa L. Star Thistle. 
Fig. 4663. 
Centaurea Calcitrapa L. Sp. Pl. 917. 1753. 


Annual, pubescent or glabrous, green; stem 
much branched, not winged, 1°-13° high. Leaves 
I-2-pinnatifid into oblong-lanceolate to linear, 
serrulate-spinulose, dentate or entire mostly acute 
lobes, the upper sessile and slightly clasping, the 
lower and basal short-petioled, 4’-7’ long, the 

_ uppermost somewhat involucrate at the bases of 
the sessile heads which are about 1’ broad; invo- 
lucre ovoid, its outer bracts ovate-oblong, tipped 
with stout, spreading, yellowish spines which are 
simple, or commonly with 2-6 bristles at the base; 
flowers purple, none of them radiant; achenes 
compressed or obscurely 4-sided; pappus none. 


In waste places and ballast, southern New York 
and New Jersey to Virginia. Also from British Co- 
lumbia to California. Adventive or naturalized 
from Europe. Called also caltrops, maize- or mouse- 
thorn. Knop-weed. June—Oct. 


g. Centaurea meliténsis L. Rayless Winged 
Centaury. Fig. 4664.° 


Centaurea melitensis L. Sp. Pl. 917. 1753. 


Annual, 1°-4° high, grayish-pubescent, much branched, 
the stem and branches narrowly winged by the decur- 
rent leaf-bases. Basal leaves lyrate, their lobes obtuse; 
stem leaves few-lobed or entire, the upper ones 1’ long 
or less; heads sessile or nearly so; involucre about 4’ 
thick, its principal bracts tipped by a slender purplish 
divergent spine 5” long or less, which is often branched 
below and with smaller spines at its base; flowers yel- 
low, none of them radiant; pappus scales unequal. 


Waste and cultivated grounds, Georgia to Missouri, Ari- 
zona, California and Oregon, and in ballast about the 
Atlantic seaports. Naturalized or adventive from Europe. 
Widely naturalized in South America. April—Sept. 


COMPOSITAE. . ’ Mor. Di 


10. Centaurea solstitialis L. Barnaby’s 
Thistle. Fig. 4665. 


Centaurea solstitialis L. Sp, PL on7.se75 3s 


Annual, cottony-pubescent, branched, 1°-2° high, 
the stem and branches winged by the decurrent leaf- 
bases. Basal leaves pinnatifid, often 6’ long; stem 
leaves lanceolate to linear, mostly entire, the upper 
2’-1’ long; involucre ovoid-globose, about 4’ thick, 
its principal bracts tipped by a stout, spreading or 
reflexed yellow spine, 6’-10” long, with several 
much smaller ones at its base; flowers yellow, none 
radiant. 


Waste and cultivated grounds, Massachusetts to New 
York, Pennsylvania, Ontario and Utah, and in Cali- 
fornia. Adventive from Europe. July—Sept. 


ro9. CNICUS [Tour.]-L. Sp: P8265 -75e- 


An annual herb, with alternate pinnatifid or sinuate-dentate leaves, the lobes or teeth 
spiny, and large sessile heads of yellow tubular flowers, solitary at the ends of the branches, 
subtended by the upper leaves. Bracts of the involucre imbricated in several series, the 
outer ovate, the inner lanceolate, tipped by long pinnately branched spines. Receptacle flat, 
bristly. Achenes terete, striate, laterally attached, the horny margin 10-toothed at the sum- 
mit; pappus of 2 series of awns, the inner fimbriate, the outer longer, naked; anther-append- 
ages elongated, united to their tips. [Latin name of Safflower, early applied to thistles. | 

A monotypic genus of the Old World. 


1. Cnicus benedictus L. Blessed Thistle. Our Lady’s Thistle. Fig. 4666. 


Cnicus benedictus L. Sp. Pl. 826. 1753. 

Centaurea benedic.a L. Sp. Pl. Ed. 2, 1296. NY 

T7O3. 

Hirsute or pubescent, much branched, 
. seldom over 23° high. Leaves oblong-lan- 2 
ceolate in outline, rather thin, reticulate- 
veined, 3-6’ long, the upper clasping, the 
basal and lower ones narrowed at the base 
and petioled; heads about 2’ broad, sub- aN 
tended by several large lanceolate to ovate- 
lanceolate leaves; bristles of the receptacle \ 


soft, long; outer awns of the pappus alter- \ 
nating with the inner. \ 


In waste places, Nova Scotia and New 
Brunswick to Maryland, Pennsylvania, Ala- 
bama and Michigan, and on the Pacific Coast. \Y Z 
Adventive from southern Europe. Holy- or <1 \ : <zs 
bitter thistle. Sweet-sultan. St. Benedict’s- YA SS fj 
thistle. May—Aug. a 


Echinops sphaerocéphalus L., a tall spinose 
plant, with large clusters of 1-flowered invo- 
lucres, the flowers white or bluish, occasionally 
escapes from cultivation into waste grounds. 
It is native of Europe. 


SUMMARY. 
Families. Genera. Species Figured. 
Benn ain OTA TIL AIS eRe PO IGE catacl cs A chert separ: II 31 130 
SPERMATOPHYTA 
Gyomiespenmae. "52. cu. 1s cee eae ate cko.. 2 10 28 
Angiospermae 
Monocotyledones, | 2... a--2nie 2 = = 32 251 1261 
Dicotyledones 
Ghoripetatdees . cus reese ois ts 103 526 1775 
Gamopetalde’ 2.21 2 yetee orien .- 46 4II 1472 


PROTA! | SPs hetero Eee S Ol oye Sle cole crhers visas 194 1229 4666 


* 
rT? ees eo 


GENERAL INDEX OF LATIN GENERA AND SPECIES 


[Classes, Families and Tribes in SMALL CAPITALS; genera in heavy face; synonyms in italics. 
Heavy face figures indicate the volume; other figures, the page. Varietal synonyms are indented. 


Abama 


(Narthecium) 


americana 
Abies 
balsamea 
canadensis 
caroliniana 
Fraseri 
Mariana 
nigra 
Picea 
Tsuga 
Abolboda 
Abronia 
californica 
fragrans 
micrantha 
Abutilon 
Abutilon 
avicennae 
Theophrasti 
Acacia 
angustissima 
brachyloba 
filicina 
filiculoides 
Acalypha 
caroliniana 
gracilens 
ostryaefolia 
virginica 


Acamptoclados 


sessilispicus 


I 487 


L230 
231 


ACANTHACEAE 3 208,240 
Acanthospermum 3459 


australe 459 
brasilium 459 
xanthoides 459 
ACERACEAE 2 494 | 
Acer 2 494 
barbatum 496 
dasycarpum 404 
carolinianum 495 
Drummondii 496 
eriocarpum 494 
glabrum 497 
interior 498 
montanum 407 
Negundo 498 
pennsylvanicum 497 
platanoides 494 
Pseudo-Platanus 494 
rubrum 495 
tridens 495 
saccharinum 494,496 
spicatum 497 
stenocarpum 495 
strictum 497 
tripartitum 497 
Acerates 3 34 
angustifolia 34 
auriculata 35 
floridana 35 | 


Acerates 
lanuginosa 
longifolia 
viridiflora 

Achillea 
borealis 
lanulosa 
ligustica 
millefolium 
Ptarmica 
santolina 

Achroanthes 


(Microstylis) 


monophylla 
unifolia 
Achyranthes 
dichotoma 
lanuginosa 
Acnida 
cannabina 
floridana 
rusocarpa 
tamariscina 


“ concatenata 


“ brostrata 

“ subnuda 

tubérculata 
Aconitum 


noveboracense 


paniculatum 
reclinatum 
uncinatum 
Acorus 
Calamus 
Acrostichum 
alpinum 
areolatum 


hyperboreum 


ilvense 
Marantae 
platyneuros 


polypodioides 


Thelypteris 
Actaea 
alba 
arguta 
eburnea 
neglecta 
racemosa 
rubra 
“ dissecta 
spicata 
Actinella 
acaulis 
lifiearifolia 
odorata 
Actinomeris 
alternifolia 


helianthoides 


squarrosa 
see Ridan 


| Actinospermum 


(Baldwinia) 
uniflorum 


L570: 


3 508 


487 | 


3 486 
3 502 


Acuan 2 331 
(Desmanthus) 
illinoensis 332 
leptoloba 232 

Adelia 2 728 
(Forestiera) 
acuminata 728 

Adenocaulon 3 457 
bicolor 457 

Adiantum 1 ayn 
Capillus-Veneris 31 
pedatum 31 
“ aleuticum 31 

Adicea I 636 
(Pilea) 
pumila 637 

Adlumia 2 143 
cirrhosa 143 
fungosa 143 

Adnaria 2 695 

Adonis 2 121 
annua 121 
autumnalis 121 

Adopogon 3 306 
(Krigia) 
carolinianum 307 
Dandelion 308 
occidentale 308 
virginicum 307 

Adorium 2 643 
divaricatum 643 
tenuifolium 644 

ADOXACEAE 3 283 

Adoxa 283 
Moschatellina 283 

Aegilops 
aromatica I 225 

Aegopodium 2 653-4 
Podagraria 654 

Aeschynomene 2 392 
aspera 392 
hispida 393 
Sesban 376 
virginica 393 

AESCULACEAE 2 498 

Aesculus 498 
arguta 499 
austrina 500 
flava 499 
“ burpurascens 499 
glabra 499 
Hippocastanum 498 
hybrida 499 
lutea 499 
octandra 499 
Pavia 500 

Aethulia 
uniflora 3 355 

Aethusa 2 645 
Cynapium 645 

Afzelia 3 205 | 
(Seymeria) 
macrophylla 205 

Agalinis 3 208 


Agalinis 
aspefa 3 209 
Besseyana 211 
fasciculata 210 
Gattingeri 213 
heterophylla 209 
linifolia 209 
maritima 211 
palustris 208 
parvifolia 212 
paupercula 210 
purpurea 210 
Skinneriana 212 
setacea 212 
tenuifolia 211 
Agassizia 
SUAVIS 3 513 
Agastache Brn 
(Lophanthus) 
anethiodora 112 
nepetoides III 
scrophulariaefoliar12 
Agave 
virginica I 534 
AGGREGATAE 3 347 
Agoseris 3 323 
(Troximon) 
cuspidata 3 323-4 
glauca 3/323 
parviflora 323 
Agrimonia 2 266 
Brittoniana 268 
Eupatoria 266 
“ hirsuta 266 
* mollis 267 
gryposepala 266 
hirsuta 266 
microcarpa 267 
mollis 267 
parviflora 267 
parviflora 268 
pumila 267 
rostellata 267 
striata 268 
Agropyron I 283 
biflorum 285 
cristatum 283 
caninum 285 
dasystachyum 284 
Gmelini 289 
novae-angliae 285 
pseudorepens 283 
pungens 284 
repens 283 
Richardsonii 285 
Smithii 284 
spicatum 284 
subvillosum 284 
tenerum 285 
tetrastachys 284 
unilaterale 285 
violaceum 285 
Agrostemma 2 61 
coronaria 7 


562 


Agrostemma 
Githago 2 61 
AGROSTIDEAE I 108-9 
Agrostis I 202 
airoides 199 
alba 202-3 
“ aristata 203 
algida 193 
altissima 206 
“ laxa 207 
antecedens 207 
arachnoides 204 
aspera 196 
asperifolia 204 
borealis 205 
brevifolia 188 
canina 205 
clandestina 195 
coarctata 203 
compressa I 200 
cruciata 225 
cryptandra I 199 
diffusa 186 
elata 206 
Elliottiana 204 
filiformis 185 
geminata 205 
hyemalis 207 
“ geminata 205 
intermedia 206 
juncea 198 
latifolia 202 
longifolia 196 
maritima 203 
mexicana 185 
novae-angliae 207 
oreophila 207 
paludosa 205 
pauciflora 186 
perennans 206 
pseudo-intermedia206 
racemosa I 186 
rubra 205 
“ americana 205 
rupestris 204-5 
scabra 207 
Scribneriana 206 
serotina 198 
Spica-venti 213 
sobolifera 185 
Schweinitzii 206 
sylvatica 186 
tenuiflora 186 
Torreyana 200 
virginica 197 
vulgaris 203 
Ailanthus 2 445 
glandulosa 446 
Aira I 237 
aquatica 245 
atropurpurea 216 
caespitosa 215 
caryophyllea 214 
coerulea 237 
cristata 245 
flexuosa 216 
melicoides 262 
nitida 244 
obtusata 244 
pallens 244 
praecox 215 
purpurea 235 
spicata S17, 
subspicata 217 
AIZOACEAE 2rsAr 
AJUGEAE 3 99 
Ajuga 100 
genevensis IOI 
reptans 101 
Albizzia iz aot 
julibrissin 331 
Alchemilla 2 263 


Alchemilla 
alpina 2 264. 
arvensis 264 
pratensis 264 
vulgaris 264 
Aletris Toor 
aurea 511 
farinosa 511 
Alisma Paces 
arcuatum 94 
brevipes 04 
cordifolia 96 
Geyeri 94 
parviflorum 94 
Plantago 94 
Plantago-aquatica 93 
rostratum 95 
tenellum 95 
subcordatum 04 
subulata 104 
superbum 94 
ALISMACEAE I 93 
Alliaria 2 169 
(Sisymbrium in 
part) 
Alliaria 170 
Allionia 2 31 
(Oxybaphus) 
albida 31 
Bodini 31 
bracteata Daa 
Bushii 31 
comata 32 
Carletoni 32 
chersophila 31 
glabra ai 
hirsuta 32 
lanceolata 31 
linearis ae 
nyctaginea 32 
ovata 32 
pilosa 32 
violacea 31 
Allium I 407 
alleghaniense 498 
canadense 499 
carinatum 499 
cernuum 498 
Helleri 500 
mutabile 500 
Nuttallii 500 
ornithogaloides 501 
reticulatum 500 
sativum 497 
Schoenoprasum 498 
sibiricum 4907 
stellatum 498 
striatum 501 
tricoccum 497 
vineale 499 
Allocarya 379 
(Eritrichium 
in part) 79 
lithocarya 79 
scopulorum 79 
Alnus I 612 
Alnobetula 612 
Alnus 613 
crispa 612 
glutinosa 613 
incana 613 
japonica 614 
maritima 614 
Mitchelliana 612 
mollis 612 
noveboracensis 613 
rugosa 613 
serrulata 613 
viridis 612 
vulgaris 613 
Alopecurus I 191 
agrestis 192 


Alopecurus 
alpinus I 103 
aristulatus 192 
geniculatus 192 
Monspeliensis 201 
myosuroides 192 
pratensis TOTS 
ALSINACEAE 2 4I 
Alsine 2 41 
(Stellaria) 
aquatica 2 
borealis 46 
“ alpestris 46 
crassifolia 46 
fontinalis 46 
glauca 44 
graminea 45 
Holostea 44 
humifusa 42 
longifolia 45 
longipes 45 
“ Edwardsii 45 
media 43 
Michauxii 55 
pubera 43 
tennesseensis 44 
tennesseensis 44 
uliginosa 43 
Alsine 2 52 
Althaea 2 513 
cannabina 514 
officinalis 514 
rosea 514 
ALTINGIACEAE 2 235 
Alyssum 2 153 
alyssoides 154 
arcticum 156 
calycinum 154 
incanum 153 
miaritimum 153 
montanum 154 
AMARANTHACEAE 2 1 
Amaranthus 21 
albus 3 
blitoides 3 
caudatus nae 
crispus 5 
chlorostachys 2 
deflexus 4 
graecizans 3 
hybridus 2 
hypochondriachus 2 
lividus 4 
Palmeri 2 
paniculatus 2 
pumilus 5 
retroflexus 22 
spinosus 3 
tamariscinus 6 
Torreyi 4 
AMARYLLIDACEAE I 531 
Amaryllis 
Atamasco 1.532 
Amblogyne 24 
Torreyi 4 
AMBROSIACEAE 3 338 
Ambrosia 3 340 
acanthicarpa 342 
artemisiaefolia 341 
bidentata 341 
elatior 341 
integrifolia 341 
maritima 340 
psilostachya 342 
tomentosa 3 343 
trifida 341 
“ integrifolia 341 
Amelanchier 2 291 
alnifolia 203 
Bartramiana 293 
Botryapium 292 
canadensis 292 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Vot. ITI. 
Amelanchier 
canadensis 2 292 
oblongifolia 292 
humilis 292 
intermedia 292 
laevis 292 
nantucketensis 292 
oligocarpa 293 
rotundifolia 293 
sanguinea 2903 
spicata 292 
stolonifera 292 
Amellus 
spinulosus 3 379 
villosus 374 
Amianthium I 489 
leimanthoides 493 
muscaetoxicum 489 
Nuttallii 492 
Ammannia 2577 
auriculata 2 578 
coccinea 578 
humilis 578-9 
Koehnei 578 
latifolia 577 
pedicellata 578 
ramosior 579 
Wrightii 578 
AM MIACEAE 2 619 
Ammi 
capillaceum 2656-7 
Majus * 657 
Ammedenia 258 
(Arenaria in part) 
peploides 258 
Ammophila I 212 
arenaria 212 
arundinacea 212 


Ammoselinum 2 652 
(Apium in part) 


Butleri 653 
Popei 652 
Amorpha 2 365 
angustifolia 365 
canescens 366 
fruticosa 365 
microphylla 366 
nana 366 
Ampelanus 3 36 
albidus 36 
Ampelopsis 2 509-10 
arborea 510 
cordata 509 
heterophylla 510 
quinquefolia SII 
Veitchu Err 
Amphiachyris 3 370 
dracunculoides 370 
Amphicarpa 2 418 
monoica 418 
Pitcheri 418 
Amphicarpon I 163 
Amphicarpon 164 
Purshti 164 
Amphilophis T1209 
saccharoides 119 
Amsinckia 3 84 
intermedia 84 
lycopsioides 84 
Amsonia 3 20 
Amsonia 20 
salicifolia 20 
Tabernaemontana 20 
AMYGDALACEAE 2 322 
Amygdalus 2 330 
persica 330 
Anacampseros 
triphylla 2 20 
ANACARDIACEAE 2 480 
Anacharis 
canadensis I 105 
Nuttall 105 


Vo. III. 
Anagallis 2 716 
arvensis 716 
“ coerulea 716 
Ananthrix 
decumbens B33 
Anaphalis 3 453 
margaritacea 453 
nubigena 453 
occidentalis 453 
Anastrophus I 125 
compressus 125 
furcatus 125 
paspaloides 125 
Anchistea I 24 
virginica 24 
Andrachne 2 453 
phyllanthoides 453 
Telephioides 453 
Andromeda 2 689 
arborea 692 
axillaris 687 
baccata 696 
calyculata 687 
Catesbaei 688 
coerulea 685 
ferruginea 691 
floribunda 690 
glaucophylla 689 
hypnoides 686 
ligustrina 691 
mariana 691 
nitida 690 
Polifolia 6890 
racemosa 6890 
recurva 688 
spinulosa 688 
taxifolia 685 
tetragona 686 
see Leucothoé 
Pieris 
Andropogon 1 ads 
alopecuroides mes} 
ambiguus 225-6 
argenteus 117 
argyraeus 117 
avenaceum 120 
Belvisii 117 
Cabinisii L107. 
chrysocomus 118 
corymbosus 116 
dissitiflorus 116 
divaricatus vie 
Elliottii TI 7, 
furcatus 118 
geminatus 118 
glaucus 119 
glomeratus 116 
halepensis 121 
Hallii 118 
littoralis 115 
macrourum 116 
mississippiensis 117 
nutans 120 
paucipilus 119 
provincialis 118 
saccharoides 119 
scoparium 115 
ternarius 117 
tetrastachyus 116 
Torreyanus 119 
vaginatus 116 
virginicus 116 
ANDROPOGONEAE I 107-8 
Androsace 2 709 
diffusa 709 
maxima 709 
occidentalis 709 
Androstephium 1 sor 
coeruleum 501 
wiolaceum 501 
Anemone 297 


canadensis 99 


Anemone 
caroliniana 2 98 
coronaria 97 
cylindrica 99 
decapetala 98 
dichotoma 99 
globosa 98 
Hepatica IOI 
hudsoniana 98 
lancifolia 100 
multifida 98 
nemorosa 100 
nudicaulis 106 
Nuttalliana 102 
parviflora 98 
patens 102 
pennsylvanica 99 
Pulsatilla 102 
quinquefolia 100 
Richardsonii 100 
riparia 99 
thalictroides 102 
trifolia 100 
virginiana 99 
ANEMONEAE 2 84 
Anemonella 2 102 
thalictroides 102 
Anemopsis I 577 
Anethum 2 633 
Foeniculum 643 
graveolens 634 
Angelica 2 636 
Archangelica 2 646 
atropurpurea 637 
Curtisii 635 
hirsuta 637 
triquinata 637 
villosa 637 
ANGIOSPERMAE I 68 
Anisostichus 3 236 
capreolata 237 
Annona 
triloba 2 83 
ANNONACEAE 2 83 
Anogra 2 598 
(Oenothera in 
part) 
albicaulis 598 
coronopifolia 598 
Douglasiana 598 
latifolia 599 
Nuttallii 599 
Nuttalliana 599 
pallida 599 
Anonymos 
aquatica 3 19 
bracteata 2 394 
carolinensis 3 89 
cassioides 3 205 
graminifolius 3 368 
odoratissima 3 369 
paniculatus 369 
petiolata 2 731 
rotundifolia 347 
Anoplanthus 3 233 
fasciculatus 234 
uniflorus 233 
Antennaria 3 448 
alpina 449 
alsinoides 451 
angustata 449 
aprica 452 
arnoglossa 450 
campestris 452 
canadensis 450 
carpathica 449 
dimorpha A52.3 
dioica 3 448, 450 
labradorica 449 
margaritacea 452 
microphylla 451 
monocephala 450 


Antennaria 
neglecta 3 452 
simplex 452 
neodioica 451 
Parlinii 450 
parvifolia A451 
petaloidea 450 
plantaginifolia 450-1 
propinqua 450 
rupicola 451 
solitaria 450 
Anthemis 3 516 
arvensis 517 
aurea 517 
Cotula 516 
maritima 516 
nobilis a7, 
repens 469 
tinctoria 517 
Anthoxanthum 1 171 
giganteum I 113 
odoratum rie fit 
Puelii 171 
Anthriscus 2 629 
Anthriscus 629 
Cerefolium 629 
sylvestris 630 
vulgaris 629 
Anthyllis 2 359 
vulneraria 360 
Anticlea I 491 
chlorantha 491 
elegans 491 
sibirica 4QI 
Antiphylla 2 215 
oppositifolia 216 
Anthirrhinum 3 178 
canadense 177 
Cymbalaria 175 
Elatine 176 
Linaria 177 
majus 178 
minus 176 
Orontium 179 
repens 178 
spurium 2507/5 
tenellum 189 
Anychia 2 29 
argyrocoma 27 
canadensis 30 
capillacea 30 
dichotoma 29, 30 
divaricata 2 
polygonoides 29 
Anychiastrum 2 29 
montanum 29 
riparium 29 
Apalanthe 
Schweinitait 105 
Apargia 3 300 
autumnalis 310 
incana 310 
nudicaulis 310 
hispida 310 
Apera I 213 
spica-venti 213 
Aphanes 2 264 
arvensis 264 
australis 264 
Aphanostephus 3 401 
arkansanus 401 
skirrobasis 401 
Aphora 
humilis 2 457 
mercurialina 456 
Aphyllon 3 233 
fasciculatum 234 
“luteum 234 
ludovicianum 235 
uniflorum 233 
Apiastrum 2 652 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Apiastrum 
see Spermolepis 
2 652 
patens 652 
Abpios 
Abios 2 418 
Priceana 418 
tuberosa 418 
Apium 2 642 
Ammi 660 
divaricatum 651 
echinatum 652 
graveolens 660 
leptophylium 660 
patens 652 
Petroselinum 642 
Popei 652 
Aplectrum I 573 
hyemale 574 
Shortii 574 
spicatum 574 
Abplopappus B37. 
armerioides 379 
ciliatus 377 
divaricatus 380 
Nuttallii 378 
rubiginosus 378 
spinulosus 379 
APOCYNACEAE 3 19 
Apocynum 3 21 
album 22 
androsaemifolium 21 
cannabinum 22-3 
“ glaberrimum 22 
“ pubescens 23 
divergens 21 
hypericifolium 22 
medium 21 
Milleri 22 
nemorale 22 
pubescens 21 
sibiricum 22 
speciosum 21 
urceolifer 22 
Apogon 
humilis 3 306 
Aquilegia 2 92 
brevistyla 93 
canadensis 92 
coccinea 2 
flaviflora 92 
vulgaris 92-3 
Arabidopsis 2 175 
novae-angliae 176 
Thaliana 176 
Arabis 2 178 
alpina 178, 180 
arsenicola 179 
brachycarpa 182 
bulbosa 186 
canadensis 182 
Collensii 183 
confinis 182 
dentata 180 
Douglassti 186 
Drummondii 182 
glabra 181 
hirsuta 181 
Holboellii 2 183 
humifusa 179 
laevigata 181 
* laciniata 181 
“ Burkei 181 
ludoviciana 179 
lyrata 179 
* occidentalis 179 
ovata 181 
patens 180 
perfoliata 181 
petraea 176 
retrofracta 183 
rhomboidea 186 


564 


Arabis 
serotina 2 181 
Thaliana 175 
virginica 179 
viridis 181 
ARACEAE I 441 
Aralia 2 617-18 
hispida 618 
nudicaulis 618 
“ elongata 618 
“ proliféra 618 
racemosa 617 
spinosa 617 
Arbutus 
alpina 6904 
Uva-Ursi 693 
Arceuthobium 
minutum I 638 
pusillum 638 
Archangelica 2 636 
atropurpurea 637 
Gmelini 646 
perigrina 646 
Archemora 2 637 
Arctagrostis I 201 
latifolia 201 
Arctium 3 547 
Lappa 547 
minus 548 
tomentosum 547 
Arctostaphylos 2 693-4 
alpina 694 
Uva-Ursi 2 603 
Arctous 2 603 
alpina 604 
Arenaria 2 52 
arctica 54 
biflora 55 
canadensis 60 
caroliniana 55 
ciliata 53 
humifusa 53 
Fendleri 54 
groenlandica 57 
Hookeri 253 
lateriflora 57 
leptoclados 53 
litorea 56 
macrophylla 58 
Michauxii 55 
patula 56 
peploides 58 
pinetorum 53 
Pitcheri 5 
propinqua 54 
rubra 59, 60 
Sajanensis 55 
serpyllifolia 2 
squarrosa 55 
stricta 55 
texana 2 56 
thymifolia 42 
verna 54 
Arethusa I 562 
bulbosa 562 
divaricata 560 
ophioglossoides 560 
spicata 8371,4 
trianthophora 562 
verticillata 561 
seePogonia 1 467-8 
Argemone 2 138 |. 
alba 139 
albiflora 139 
hispida 139 
intermedia 139 
mexicana 138-9 
platyceras 139 
Argentina 2 258 
Anserina 258 
Babcockiana 258 
littoralis 258 


Argentina 
vulgaris 2 258 
Argyroihamnia 
humilis 2 457 
mercurialina 456 
Aria 2 291 
Arisaema I 442 
atrorubens 442 
Dracontium 443 
pusillum 442 
Stewardsonii 443 
triphyllum 442 
Aristida I 178 
adscensionis 178 
basiramea 179 
Curtissii 179 
desmantha 183 
dichotoma 178 
* Curtissti 179 
dispersa 180 
divaricata 183 
fasciculata 180 
Fendleriana 180 
gracilis 181 
“ depauperata 181 
intermedia 182 
lanata 182 
lanosa 182 
longiseta 181 
oligantha 181 
purpurascens 182 
ramosissima 179 
“ uniaristata 179 
tuberculosa 183 
Wrightii 180 
ARISTOLOCHIACEAE 
I 641 
Aristolochia I 645 
Clematitis 645 
hastata 645 
macrophylla 646 
Nashii 645 
rotunda 645 
Serpentaria 645 
Sipho 646 
tomentosa 646 
Armeria 2 218 
vulgaris 218 
Armoracia 2 162 
Armoracia 163 
rusticana 263 
Arnica 2.543 
acaulis 533 
alpina 534 
angustifolia 534 
Chamissonis 534 
chionopappa 3 534 
Claytoni 533 
cordifolia 533 
gaspensis 534 
maritima 541 
mollis 534 
montana 533 
nudicaulis 533 
plantaginea 534 
Sornbergeri 534 
Arnoseris 3 308 
minima 309 
pusiila 309 
Aronia 2 290 
(Pyrus in part) 
alnifolia 293 
arbutifolia 291 
atropurpurea 291 
melanocarpa 291 
nigra 291 
Arrhenatherum 1 220 
avenaceum I 220 
elatius 220 
Artemisia 2) Be 
Abrotanum 526 
Absinthium 52 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Artemisia 
annua 3 526 
biennis 526 
Bigelovii 530 
borealis 523 
cana 530 
canadensis 524 
capillifolia 356 
Carruthii 528 
caudata 3 523 
dracunculoides 524 
““incana 524 
elatior 527 
filifolia 525 
Forwoodsi 524 
frigida 525 
glauca 524 
gnaphalodes 529 
groenlandica 523 
kansana 528 
longifolia 528 
Ludoviciana 529 
matricarioides 521 
mexicana 529 
pontica 3 527 
procera 526 
serrata 528 
Stelleriana 527 
tridentata 530 
vulgaris 522. 527 
Arum I 442-3 
Dracontium I 442 
nepenthoides 442 
triphyllum 442 
virginicum 444 
Aruncus 2 247 
(Spiraea in part) 
Aruncus 2 247 
sylvester 247 
Arundinaria I 205 
macrosperma 205 
tecta 295 
Arundo 
arenaria 212 
brevipilis 212 
cinnoides 211 
festucacea 262 
Phragmites 232 
tecta 295 
seeCalamagrostis 207 
Asarum I 642 
acuminatum 642 
arifolium 643, 4 
canadense 642 
europaeum 642 
grandiflorum 643 
heterophyllum 644 
macranthum 643 
Memmingeri 644 
reflexum 643 
“ ambiguum 643 
Shuttleworthii 643 
virginicum 644 
ASCLEPIADACEAE 3 23 
Asclepias 24 
amoena 26 
amplexicaulis 28 
arenaria 30 
auriculata a6 
Bicknellii 28 
brachystephana 31 
Cornutt 30 
decumbens aE 
Douglasii 30 
exaltata 28-9 
floridana 35 
galioides A 3 
incarnata 26 
intermedia 28 
Jamesii 27 
kansana 30 
lanceolata 25 


Vot. III. 
Asclepias 
lanuginosa S035 
latifolia 25. 
Meadii 28 
nigra 36 
obtusifolia 27-8 
ovalifolia 31 
perennis 31 
paupercula 25 
phytolaccoides 29 
pulchra 27 
pumila 32 
purpurascens 26 
quadrifolia 29 
rubra 3 26 
speciosa 30 
stenophylla 34 
Sullivantii 27 
syriaca 24, 29-30 
tuberosa 25 
variegata 29 
verticillata 32 
viridiflora 34 
viridis 33 
Asclepiodora 3133 
decumbens yn 
viridis 33 
Ascyrum 2 528 
Crux-Andreae 
2 528,535 
hypericoides 2 528 
stans 528 
Asimina 2 83 
triloba 83 
Asparagus Bente) 
officinalis 514 
Asperugo 3 84 
procumbens 84 
Asperula 3 266 
arvensis 267 
galioides 3 267 
odorata 267 
Aspidium I 16-22 
see Dryopteris 
americanum 22 
Asplenium I 25-30 
acrostichoides 30 
angustif oliunm 28 
Bradleyi 30 
ebeneum 27 
ebenoides 26 
Filix-foemina 30 
fontanum 29 
montanum 29 
parvulum 27 
pinnatifidum 27 
platyneuron 27 
pycnocarpon 28 
resiliens 27 
rhizophylla 26 
Ruta-muraria 29 
Sclopendrium 25 
thelypteroides 30 
Trichomanes 26-8 
viride 28 
Asprella 
HAystrix I 294 
virginica 168 
Aspris I 214 
caryophyllea 214 
praecox 215 
Aster 3 406 
amellus 406 
acuminatus 432 
adscendens 421 
amethystinus 418 
angustus 434 
annuus 440 
anomalus 413 
argenteus 423 
azureus 414 
bellidiflorus 428 


Vor iit. 


Aster 


wv 


biflorus 3 
capillaceus 
carmesinus 
choralis 
Claytoni 3 
commutatus 
concinnus 
concolor 
cordifolius 
“ laevigatus 
coridifolius 
corymbosus 3 
curvescens 
depauperatus 
“ parviceps 
diffusus 
divaricatus 

409, 410, 432, 
Drummondii 
dumosus 
“ coridifolius 
“ strictior 
ericaefolius 
ericoides 
“ depauperatus 
“ parviceps 
“ pilosus 
“ platyphyllus 
“ Pringlet 
“ busillus 
“ Randi 
“ villosus 
exscapus 3 
exilis 
Faxoni 
Fendleri 
flexuosus 
foliaceus 
foliolosus 
furcatus 
glomeratus 
gracilis 
graminifolius 
grandiflorus 
Gravesii 
Herveyi 
hirsuticaulis 
hirtellus 
humilis 3 
ianthinus 
incanopilosus 
infirmis 
junceus 
junciformis 
kentuckiensis 
Kumleini 
laevis 
“amplifolius 3 
“ botomacensis 


‘ lateriflorus 


Lindleyanus 

“ eximius 
linariifolius 
linifolius 
longifolius 

“ villicaulis 3 
longulus 
Lowrieanus 415, 
lutescens 
macrophyllus 
MajuUs 3 
miser 
missouriensis 
modestus 
multiflorus 418, 
“ commutatus 

“ exiguus 

“ stricticaulis 
multiformis 
nebraskensis 
nemoralis 


Aster 
nemoralis 3 432 
“ Blake 432 
nobilis 413 
novae-angliae 417 
novi-belgii 421 
“atlanticus 421 
“ Brittoni 421 
“ elodes 421 
“ litoreus 421 
NuttalliiFendleri 423 
oblongifolius 417 
“rigidulus 417 
paludosus 431 
paniculatus 428, 430 
parviceps 431 
patens 416 
“ gracilis 416 
“ phlogifolius 417 
patulus A419 
phlogifolius 417 
phyllodes 422 
polycephalus 3 427 
polyphyllus 429 
prenanthoides 419 
“ borrectifolius 419 
Priceae 430 
Pringlei 430 
ptarmicoides 432 
pusillus "431 
puniceus 419 
purpuratus 420 
racemosus 427 
Radula 425 
“ biflorus 425 
“ strictus 425 
ramulosus var. 427 
roscidus 412 
roseus 417 
sagittifolius 416 
salicifolius 428 
“ subasper 428 
Saundersii 416 
saxatilis 429 
schistosus 419 
Schreberi AII 
sericeus 423 
Shortii 414 
solidagineus 405 
spectabilis 424 
stenophyllus 428 
strictus 425 
subasper 428 
subulatus 433 
surculosus 424 
tardiflorus 419 
tanacetifolius 435 
tenebrosus 409 
tenuifolius 433 
tenuifolius 428 
“ bellidiflorus 428 
tortifolius 406 
Tradescanti 429 
Tradescanti 427 
turbinellus 426 
umbellatus 443 
“ latifolius 443 
“ pubens 443 
unalaschensis 418 
“ major 418 
undulatus 3 416 
vernus 441 
villosus 430 
vimineus 427-9 
“ foliolosus 427 
violaris 412 
virgatus 420 
viridis 409 
Wilsoni 415 
ASTEREAE 3 344-5 
Astilbe 2 215 
biternata 215 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Astilbe 
decandra 2 215 
rivularis 215 
Astragalus 2 378-90 
aboriginorum 2 384 
adsurgens 379 
alpinus 382 
bisulcatus 380 
Blakei 382 
caespitosus 386 
campestris 390 
canadensis 379 
carnosus 377 
carolinianus 2 379 
caryocarpus 377. 
Cooperi 385 
crassicarpus 377 
distortus 382 
Drummondii 381 
elatiocarpus 383 
elegans 384 
elongatus 385 
eucosmos 384 
filifolius 386 
flexuosus 385 
frigidus amer, 2 385 
gracilis 384 
hyalinus 388 
Hypoglottis 380 
Jesupi 382 
Kentrophyta 387 
lotiflorus 383 
melanocarpus 383 
mexicanus 377 
microlobus 384 
missouriensis 383 
mollissimus 379 
multiflorus 386 
ne braskensis 383 
neglectus 385 
oroboides 384 
Orobrychis 378 
parviflorus 384 
pectinatus 380 
pictus filifolius 386 
plattensis 2 377 
racemosus 381 
Robbinsii 381-2 
“ Jesupi 382 
sericoleucus 388 
Shortianus 383 
striatus 379 
tenellus 386 
tennesseensis 378 
trichocalyx 377 
triphyllus 387 
Atamosco I 532 
Atamasco 532 
Athamanta 
chinensis 2 636 
Athanasia 
trinervia 3 503 
Atheropogon I 227 
apludoides 227 
curtipendulus 228 
oligostachyus 227 
Athyrium I 30 
cyclosorum 30 
Filix-foeminea 30 
pycnocarpon 28 
thelypteroides 30 
Atragene 2 126 
(Clematis in part) 
americana 126 
Atriplex 2 17 
arenaria 19 
argentea 18 
canescens 19 
expansa 18 
hastata 18 
hortensis 17 
littoralis 18 


565 


Atriplex 
Nuttallii 2 19 
patula 18 
rosea 18 
volutans 18 
Atropa 
physalodes 3 155 
AVENEAE I 108-9 
Avena I 218 
elatior 220 
fatua 218 
flavescens 217 
Hookeri 219 
mollis 217 
palustris 217 
pennsylvanica 217 
pubescens 219 
pratensis americana 
219 
purpurascens 219 
sativa 218 
Smithii 219 
spicata 220 
sterilis 218 
striata 216, 219 
Torreyi 219 
Axonopus 
furcatus Pes 
Axyris 2 20 
amarantoides 20 
Azalea 2 677-80 
(Rhododendron 
in part) 
arborescens 679 
calendulacea 678 
canescens 678 
indica 677 
lapponica 680 
lutea 678 
nudiflora 678 
pilosa 682 
procumbens 683 
viscosa 679 
Azolla 1 38 
caroliniana 38 
filiculoides I 38 
Baccharis 3 444 
foetida 447 
glomeruliflora 445 
halimifolia 445 
ivifolia 3 444 
neglecta 445 
salicina 445 
VISCOSA 447 
Wrightii 446 
Bacopa 
Monniera 192 
rotundifolia 192 
Bahia - 
oppositifolia 3 508 
Balduina 3 502 
uniflora 502 
Baldwinia 502 
Ballota 3°122 
nigra 123 
BALSAMINACEAE 2 440 
BAMBUSEAE I 108,111 
Baptisia 2 344 
alba 346 
australis 344 
bracteata 345 
leucantha 346 
leucophaea 344-5 
mollis 343 
tinctoria 345 
villosa 345 
Barbarea 2 176 
Barbarea 177 
praecox 177 
stricta 177 
vulgaris 176-7 


566 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Barbarea 
v. arcuata Z A177 
verna 177 
Barkhausia 
grandiflora 3,325 
Bartonia 3 16 
albicaulis 2 566 
decapetala 567-8 
iodandra ZaL7 
Janceolata Br7 
nuda 2 567 
ornata 2 568 
paniculata E77, 
tenella 16 
verna 17 
virginica 16 
Bartonia 2 566 
Bartsia 3 218 
acuminata 215 
alpina 219 
coccinea 214 
Odontites 219 
viscosa 219 
Bassia 
hirsuta 2 16 
Batis 
vermiculata 2 23 
Batodendron 2 698 
andrachnaeforme 698 
arboreum 698 
Batrachium 2 115 
(Ranunculus 
in part) 
circinatum 2 116 
divaricatum 116 
hederaceum 115, 6 
longirostre 116 
trichophyllum 116 
Batschia 
canescens 3 89 
carolinensis 89 
Beckmannia I 228 
erucaeformis 228 
“ uniflora 228 
Behen 2 64 
vulgaris 64 
Belamcanda I 541 
chinensis 542 
Bellis 3 401 
integrifolia 402 
perennis 401-2 
Benthamidia 2 664 
Benzoin 2 135 
(Lindera) 
aestivale 135 
Benzoin 2 135 
melissaefolium 136 
BERBERIDACEAE 2 126 
Berberis 2 127 
Aquifolium 128 
canadensis 127 
repens 128 
sinensis I2 
vulgaris 127 
Berchemia 2 501 
scandens 502 
volubilis 502 
Bergia 2 539 
capensis 239 
texana 539 
Berlandiera 3 463 
lyrata 464 
texana 463 
Berteroa 2 153 
(Alyssum in part) 
incana 153 
mutabilis 153 
Berula 2 65 
angustifolia 655 
erecta 655 
Besseya 
rubra 3 199 


Betonica 3 128 
(Stachys in part) 
officinalis 128 

BETULACEAE I 605 

Betula I 607 
alba 608, 9 
alleghanensis 610 
Alnobetula 612 
alnus 613 
coerulea 608 
cordifolia 609 
fontinalis 610 
glandulifera 611 
glandulosa 611 
Jackii 609 
lenta 609 
lutea I 610 
Michauxii 612 
microphylla 610 
nana 612 
nigra 609 
occidentalis 610 
papyracea 609 
papyrifera I 609 
“ minor 609 
pandula 608 
populifera 608 
pumila 611 
Sandbergi 611 

Bicuculla 2 142 
(Dicentra) 
canadensis 142 
Cucullaria 141 
eximia 143 

Bidens 3 494-9 
acuta 3 496 
aristosa 495-6, 499 
Becki 500 
bidentoides 3 496 
bipinnata 498 
cernua 495 
chrysanthemoides 495 
comosa 49 
connata 495-6 
“ pinnata 495 
coronata 3 498 
discoidea 497 
Eatoni 3 495 
elegans 495 
frondosa 497 
“ puberula 497 
gracilis 501 
involucrata 499 
laevis 3 495 
lugens 495 
melanocarpa 3 497 
riparia 496 
trichosperma 498 
“ tenuiloba 498 
tripartita 494 
vulgata 497 

Bifora 2 647 
americana 647 
radians 647 

Bigelovia 
Engelmanni 2377; 
graveolens 376 
Howardi 376 
nudata 375 
uniligulata F302 

BIGNONIACEAE 3 236 

Bignonia 237 
capreolata Dar 
Catalpa 238 
crucigera 237 
radicans 237 
sempervirens 2 730 
tomentosa 3 189 

Bistorta I 671 
vivipata 671 

Blechnum 
virgimicum I 24 


Blephariglottis 1 556 
albiflora 556-7 
blephariglottis me 
ciliaris 557 
cristata 557 
grandiflora 558 
lacera 558 
leucophaea 558 
peramoena 559 
psycodes 559 

Blephilia 3 134 
ciliata 134-5 
hirsuta 135 
nepetoides 135 

Bletia 
aphylla L577. 

Blitum 2 15 
Bonus-Henricus 14 
capitatum 15 
chenopodioides 17 
glaucum pa) 
Nuttallianum 17 
rubrum 13 

Bocconia 
cordata 2 140 

Boebera ey Tie) 
chrysanthemoides 513 
papposa 513 

Boehmeria I 637 
cylindrica 637 
ramiflora 637 
scabra 637 

Boltonia 3 403 
asteroides 404 
decurrens 405 
diffusa 404 
glastifolia 403-4 
latisquama 404 

Bonamia 
see Stylisma 3 41 

Boottia 2 262 

BoRAGINACEAE 2072 

Borago 92 
officinalis 92 

Borrichia 3.477 
frutescens 477 

Botrychium I 2-6 
dissectum 5 
gracile 6 
lanceolatum 6 
Lunaria 3 
matricariae 5 
matricariaefolium 4 
matricarioides 5 
neglectum 4 
obliquum 5 
occidentale 6 
onondagense 4 
rutaceum 5 
rutaefolium 5 
silaifolium 6 
simplex 3 
tenebrosum 4 
ternatum vars. a) 
virginianum 6 

Bouteloua I 227 
curtipendula 228 
hirsuta 227 
oligostachya 227 
racemosa 228 

Boykinia 
aconitifolia 2 223 
humilis 579 

Brachiaria 
digitarioides 1 137 
obtusa 138 

Brachyactis 3 434 
(Aster in part) 
angusta 434 
ciliata 434 
frondosa 434 

Brachychaeta 3 400 


Vo. II1. 

Brachychaeta 

cordata 3 401 

sphacelata 401 
Brachyelytrum 1 189 

aristatum 190 

“ Engelmanni 190 

erectum 190 
Brachylobus 

hispidus 2 161 
Brachyris 

dracunculoides 3 370 
Brachystemon 


see Koellia 3 141-5 


Bradburya 2 417 
virginiana 417 
Bradleia 2 374,418 
frutescens 374 
Bramia 3 192 
indica 192 
Monniera 192 
rotundifolia 192 
Brasenia 2 76 
peltata 76 
purpurea 76 
Schreberi 76 
BRASSICACEAE 2 146 
Brassica 2 g92 
alba I9I 
arvensis 192 
campestris 193 
Eruca 192 
japonica 194 
juncea 193 
Napus 194 
nigra 193 
oleracea 192-4 
orientalis 175 
perfoliata 175 
Rapa 193 
Sinapistrum 192 
Brauneria 3475 
see Echinacea 475-6 
atrorubens 476 
angustifolia 3 476 
pallida 476 
paradoxa 476 
purpurea 475 
Braya 
humilis 2 176 
Breweria 
see Stylisma 3 4! 
Brickellia 
grandiflora 3 363 
Briza I 250 
canadensis 263 
Eragrostis 238 
media 250 
minor 251 
Brizopyrum__. 
see Distichlis 1 249 
BROMELIACEAE I 456 
Bromus I 274 
arvensis 279 
asper 275 
breviaristatus 280 
brizaeformis 280 
ciliatus 276 
commutatus 278 
erectus 276 
giganteus 273 
hordeaceus 278 
incanus 276 
inermis 277 
Kalmii 277 
madritensis 275 
maximus 275 
mollis 278 
patulus 279 
Porteri 277 
purgans 276 
“ latiglumis 276 


Vou. III. 


Bromus 
racemosus 1 278 
tubens 275 
Schraderi 280 
secalinus 274, 278 
squarrosus 279 
sterilis 275 
tectorum 274 
unioloides 280 
Broussonetia 
papyrifera I 632 
Brunella 
see Prunella 3 115 
Brunnichia I 677 
cirrhosa 677 
Bryanthus 
taxifolius ~ 2 685 
Buchloé 
dactyloides Teak 
Buchnera 3 205 
americana 205 
canadensis 97 
Buda 2 59 
borealis 60 
marina 59 
minor 59 
rubra 60 
Buettneria 2 132 
fertilis 132 
florida 132 
Bulbilis I 231 
(Buchloé) 
dactyloides 231 
Bulliarda 
aquatica 2 206 
Bumelia 2 719 
lanuginosa 720 
lycioides 719 
retusa 719 
Bunias 
Cakile 2 195 
edentula 196 
Buphthalmum 
frutescens 3.477 
helianthoides 467 
Bupleurum - 2 639 
Odontites 639 
rigidum 639 
rotundifolium 2 634 
BURMANNIACEAE I 546 
Burmannia I 547 
biflora - 547 
disticha 547 
Bursa 257 
(Capsella) 
Bursa-pastoris 158 
pastoris 158 
Burshia 
humilis 2 615 
Butneria 2132 
Butomus 
umbellatus I 104 
BUXACEAE 2 480 
CABOMBACEAE 2575 
Cabomba 2 76 
aquatica 76 
caroliniana 76 
Cacalia 


seeMesadenia 3 536-7 
Vernonia 3350-3 


arkansana 351 
atriplicifolia 537 
fasciculata 352 
reniformis 536 
suaveolens 538 
tuberosa Pers 
CACTACEAE 2 568 
Cactus 2 570 
ferox 573 
fragilis 573 
humifusus 572 


Cactus 
mamillaris 2570 
MISSOUFIENSIS 570 
Opuntia 571 
viviparus 571 
Caenotus 3 442 
CAESALPINIACEAE 
2 334-40 
Caesalpinia 338 
Falcaria 338 
Jamesii 338 
Cakile 2 195 
americana 196 
edentula 196 
Calamagrostis 1 207 
arenaria 212 
brevipilis zi 
breviseta 208 
“ debilis 208 
canadensis 209 
“ acuminata 209 
cinoides 211 
confinis 210 
gigantea pra 
hyperborea 211 
“ americana 211 
inexpansa 210 
lacustris 208 
labradorica 211 
Langsdorfli 209 
longifolia 212 
Macouniana 210 
neglécta 210 
“ borealis 210 
nemoralis 209 
Nuttalliana 205 
perplexa 209 
Pickeringii 208 
Porteri 208 
robusta 211 
stricta 210 
sylvatica 208 
Calamintha 3 138 
Clinopodium 138 
glabella Nuttallit 139 
officinalis 139 
Nuttallii ; 139 
see Clinopodium 
137-9 
Calamovilfa q I 212 
brevipilis Zia 
gigantea 213 
longifolia 212 
“ magna 212 
Calceolaria 2 564 
(Solea) 
verticillata 564 
Calla I 444 
palustris 444 
Callicarpa 3 99 
americana 99 
purpurea 99 
Calligonum 
canescens 2 19 
Calliopsis 


cardaminefolia 3 493 


Callirhoé 2 516 
alceoides 516 
Bushii 517 
digitata 2 517 
involucrata 518 
triangulata 517 


CALLITRICHACEAE 2 477 


Callitriche 2477 
Austini 477 
autumnalis 478,579 
bifida 478 
deflexa 477 
heterophylla 478 
palustris 477-8 
terrestre 477 
verna 478 


Callitriche 
vernalis 
Calluna 
vulgaris 
Calochortus I 
elegans 
Gunnisonii 
Nuttallii 
Calophanes 
oblongifolia 
Calopogon I 
pulchellum 
Caltha 
flabellifolia 
natans 
palustris 
radicans 
CALYCANTHACEAE 2 
Calycanthus 
fertilis 
floridus 
glaucus 
laevigatus 
Calycocarpum 2 
Lyoni 
Calymenia 
pilosa 
Calypso z 
borealis 
bulbosa 
Calystegia 
sepium 3 
“ pubescens 
spithamaea 
Camassia I 
Fraseri 
Camelina 2 
microcarpa 
sativa 
sylvestris 
Cammarum 2 
hyemale 
Campanula 3 
americana 
aparinoides 
biflora 
divaricata 
fiexuosa 
glomerata 
latifolia 3 
linifolia var. 
medium 
patula 
perfoliata 
rapunculoides 
rotundifolia 
“ Langsdorfiana 
“ velutina 
Speculum 
Trachelium 3 
uliginosa 
uniflora 
CAMPANULACEAE 3 
Campsis 
radicans 3 
Camptosorus I 
rhizophyllus 
Campulosus I 
(Ctenium) 
aromaticus 
Campylocera 
leptocarpa 
CAN NABINACEAE 
Cannabis I 
sativa 
Cantua 
aggregata 3 
longiflora 
pungens 
Capnoides 
(Corydalis) 
aureum 


pn 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


85 
132 
132 
132 
132 
132 
132 


567 
Capnoides © 
a. occidentale 2 145 
campestre 145 
crystallinum 2 146 
curvisiliquum 145 
flavulum 144 
micranthum 144 
montanum 145 
sempervirens 144 
CAPPARIDACEAE 2 196 
Capraria 
gratioloides 3 196 
multifida IQI 
CAPRIFOLIACEAE 267 
Caprifolium 
Douglassii 279 
gratum 278 
Capriola T ‘222 
(Cynodon) 
Dactylon 222 
Capsella 
Bursa-pastoris 2 158 
elliptica 158 
Carara 2 166 
coronopus 167 
didymus 167 
Cardamine 2 183 
arenicola 185 
bellidifolia 186 
bulbosa 186 
Clematitis 185 
diphylla 188 
Douglasii 186 
flexuosa 184° 
heterophylla 188 
hirsuta : 184 
“ sylvatica 185 
laciniata 187 
ludoviciana 179 
maxima 188 
parviflora 185 
purpurea 186 
pennsylvanica 184 
“ Brittoniana 184 
pratensis 183-4 
rhomboidea 186 
rotundifolia 187 
spathulata 179 
uniflora 189 
virginica 179, 185 


seeDentaria 2 187-8 
Cardiospermum 2 so01 


Halicacabum 501 
CARDUACEAE 3 347 
Carduus | 3 554 

see Cirsium 548-53 

acanthoides 555 

crispus 555 

marianus 55 

nutans 553-4 

pumilus 552 
Carex I 352 

abacta 431 

abbreviata 415 

abdita I 394 

abscondita 400 

acutiformis 420 

adusta 386 

aenea 387 

aestivaliformis 409 

aestivalis 409 

agglomerata 368 

aggregata 368 

alata I 384 

alba setifolia 396 

albicans 3902 

albolutescens 385 

“ cumulata 385 

albursina 401 

alopecoidea 368 

alpina 417 

altocaulis 399 


568 


Carex 
ambusta I 432 
amphibola 406 
amphigena 373 
annectens 369 
aperta 421 
anceps 402 
“ patulifolia 402 
aquatilis 422 
arcta 375 
arctata 410 
“ Faxont 410 
arida 381 
arenaria 363 
argyrantha 387 
aristata 428 
Asa-Grayi 439 
assiniboinensis 411 
atherodes 428 
atlantica 378 
atrata ovata 418 
atratiformis 418 
atrofusca 412 
aurea 397 
“ celsa 396 
austrina 366 
australis 366 
Backana 388 
Backti 388 
Baileyi I 436 
Barrattii 417 
Bebbii 382 
Bellardi 351 
Bella-villa 440 
Bicknellii 383 
bicolor 396 
Bigelovit 421 
bipartita 351 
blanda 402 
Boottiana 395 
bromoides 376 
brunnescens 375 
“ gracilior 375 
bullata 435 
“ Greenei 435 
Bushii 414 
Buxbaumii 419 
caespitosa 422 
camporum 362 
canescens 373-5 
“ disjuncta 375 
“ bolystachya 375 
“ subloliacea 375 
“ vulgaris 375 
capillaris 411 
“ elongata 411 
capitata 361 
Careyana 400 
caroliniana 414 
caryophyllea 391 
castanea 4II 
cephalantha 378 
cephaloidea 367 
cephalophora 1 366 
“ angustifolia 367 
“ maxima 368 
cherokeensis 412 
chordorhiza 362 
Collinsii 430 
colorata 398 
communis 391 
“ Wheeleri 391 
comosa 438 
compacta 434 
complanata 414 
concinna 395 
concolor 421 
conjuncta 369 
conoidea 405 
contigua 365 
Cooleyi 437 
costellata 413 


Carex 

Crawei I 383, 404 
Crawfordii 380 
“ vigens 380 
crinita 416, 425 
“ minor 425 
“ simulans 425 
cristata 381 
cristatella 381 
crus-corvi 371 
cryptocarpa 424 
curta brunnescens 375 
cuspidata 423 
Davisii 408 
debilis I 410 
“ prolixa 410 
“ bubera 410 
“ Rudgei 410 
decomposita arr 
deflexa 393 
“ Deanei 393 
“ Farwellit 393 
“ media 393 
delicatula 377 
Deweyana 376 
diandra 370 
digitalis 401 
“ copulata 401 
disperma 372 
Douglasii 363 
Drummondiana 390 
durifolia 388 
eburnea 396 
echinata 365, 378 
Eggertii 440 
elachycarpa B77, 
Eleocharis 362 
Emmonsii 392 
Emoryi 423 
exilis 376 
extensa I 429 
Farwellii 393 
festucacea 383 
filifolia 390 
filiformis 427 
“ lanuginosa 427 
“ latifolia 427 
flacca 415 
flaccosperma 407 
flava 429-30 
“ cyperoides 429 
“ graminis 430 
“vectirostra 430 
“ wiridula 429 
flaxilis AII 
flexuosa 410 
floridana 393 
foenea 387 
“ ferruginea 384 
“ sabulonum 385 
“ sparsiflora 386 
“« xerantica 385 
folliculata 431 
formosa 408 
Frankii 438 
Fraseri 441 
Fraseriana 441 
fulva 429 
fulvescens 429 
furcata 438 
fusca 419 
gigantea I 441 
glabra 409 
glareosa 373,4 
glauca 415 
glaucescens 420 
glaucodea 406 
Goodenowii 422 
gracillima 407 
“humilis 407 
grandis 441 
granularis I 404 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Carex 

g. recta I 404 
“ Shrineri 403 
gravida 367 
Grayi 439 
“ hispidula 439 
Greenii 435 
grisea 406 
“ angustifolia 406 
“ mutica 406 
“ rigida 406 
gynandra 425 
gynocrates 361 
Haleana 403 
Halei 371, 440 
Halleri 417 
Harperi 388 
Hartii 435 
Hassei 396 
Haydeni I 421 
Heleonastes 374 
heliophila 391 
helvola 373 
heterostachya 404 
- hirsuta 414 
hirta 428 
hirtifolia 394, 407 
Hitehcockiana 405 
Hoppneri 423 
hormathodes 384 
Houghtonii 427 
Howei 377 
hyperborea 421 
hystricina 437 
“ Dudleyi 437 
impressa 426 
incomperta 378 
incurva 361 
interior 377 
“ capillacea 377 
intumescens 439 
“ Fernaldii 439 
irrigua 417 
Jamesii 388 
Jamesii 423 
Joori 419 
katahdinensis 405 
Kneiskerni 410 
Lachenalii 373 
lacustris 425 
laeviconica 428 
lagopina 373 
lagopodioides 380 
lanuginosa I 427 
“ kansana 427 
lasiocarpa 427 
laxiculmis 401 
laxiflora 402 
“ divaricata 403 
“ gracillima 402 
“ latifolia 401 
“mutica 407 
“ latifolia 401 
“ leptonervia 402 
“ patulifolia 402 
* styloflexa 403 
“-varians 402 
Leavenworthii 367 
Leersii 378 
lenticularis 422 
lepidocarpa 430 
leporina 385 
leptalea 388 
limosa 416 
littoralis 417 
livida 397 
longirostris 412 
louisianica A40 
lupuliformis 440 
lupulina 440 
“bedunculata 440 
“ polystachya 440 


Vo. III. 
Carex 

lurida I 436 
“ exundans 436 
“ flaccida 436 
macloviana 383 
macrochaeta 416 
macrokolea 419 
magellanica 417 
mainensis 432 
marcida 362 
maritima 424 
Meadii 398 
mediterranea 366 
membranacea 434 
membranopacta 434 
mesochorea 366 
Michauxii 430 
Michauxiana 431 
microglochin 389 
miliacea 407 
miliaris 431-2 
“ aurea 432 
“ major 432 
mirabilis 382 
“ berlonga 382 
misandra TA ns 
monile 433 
“ monstrosa 433 
Muhlenbergii 365-6 
“ austrinus 366 
“ enervis 365 
muricata 365 


“ cephaloidea 367 
muskingumensis 381 


nardina 360 
nebraskensis 423 
nigro-marginata 393 
normalis 382 
norvegica 374 
novae-angliae 392 
oblita 409 
Oederi 429 
“ pumila 429 
Olneyi 435 
oligocarpa 404 
oligosperma 436 
oronensis 379 
ovalis 385 
ovata 418 
oxylepis 408 
pallescens 415 
paludosa 420 
panicea 397 
Parryana 418 
pauciflora 389 
paupercula 417 
“ irrigna I 417 
“ pallens 417 
pedicellata 391 
“ Wheeleri 391 
Peckii 392 
pedunculata 395 
pennsylvanica 391 
picta 395 
pinguis 386 
plantaginea 399 
platyphylla 400 
podocarpa 416 
polygama 419 
polystachya 375, 440 
polymorpha 399 
polytrichoides 388 
Porteri 425 
praecox 391 
prairea 379 
prasina ~ 407 
pratensis 386 
praticola 386 
projecta 381 
pseudo-cyperus 437 
““ americana 438 


pseudo-helvola 374 


Vor. Err. 
Carex 
ptychocarpa I 400 
pubescens 394 
pulla 432 
pulicaris Bee 
Raeana 433 | 
rariflora I 416 
recta 424 
recurva 412 
Redowskyana 361 
retrocurva 401 
retroflexa 364 
tetrosa PeAS5 
Richardsoni 396 
rhomalea 432 
rigida 421 
riparia 425-6 
rosea 365 
“radiata 365 
roseoides 379 
Rossii 392 
rostrata 431, 434 
rotundata 434 
rupestris 390 
sabulosa 385 
salina 423 
saltuensis 399 
Sartwellii 363 
saxatilis 432 
“ miliaris 432 
“rhomalea 432 
scabrata I 416 
scabrior 370 
Schweinitzii 437 
scirpoides 377 
“ capillacea 377 
scirpoidea 377,390 
scoparia 380 
“ condensa 380 
SCOrsa 379 
setacea 370 
setifolia 396 
Shortiana 419 
Shriveri 403 
sicaeformis 371 
siccata 364 
silicea 385 
Smithri 414 
sparganioides 368 
sparsiflora 387 
Sprengelii 412 
squarrosa 438-9 
stellulata 378 
stenolepis 438 
stenophylla 362 
sterilis 377 
sterilis 378 
Steudelii 388 
stipata Tei 
straminea 382 
“ brevior 383 
“ Crawei 383 
“ invisa 384 
“ mirabilis 82 
“moniliformis 385 
striata 426 
“ brevis 426 
striatula 403 
stricta 420-1 
“ angustata 420 
* decora 421 
styloflexa 403 
stylosa 418 
suberecta 384 
subspathacea 423 
subulata 430 
subulosa 385 
Sullivantii 407 
supina 389 
Swanii 413 
sychnocephala 379 
tenella 372 


Carex 
tenera 
© Richii 
tentaculata 
“ gracilis 
tenuiflora 
tenuis 
teretiuscula 
“ prairea 
“ ramosa 
tetanica 
“ Canbyi 
“ Carteri 
“ Meadii 
“ Woodiu 
texensis 
tonsa 
Torreyana 
Torreyi 
torta 
tribuloides 
“ Bebbii 
“ cristata 


I 382 
384 
436 
436 
372 
410 
370 
370 
$70 

I 398 

398 

398 

398 

398 

364 

394 

415 

415 

421 

380 

382 

381 


408, 


“moniliformis 381 
“ vreducta 381 
triceps 414 
“ hirsuta 414 
 Smuthii 414 
trichocarpa 428 
“ aristata 428 
“imberbis 428 
“ Deweyi 428 
“ turbinata 428 
trisperma 372 
“ Billingsii 372 
Tuckermani 435 
typhina 439 
typhinoides 439 
umbellata I 394 
“ tonsa 394 
“ vicina 394 
utriculata 434 
ursina 373 
ustulata 412 
V ahlii 417 
vaginata 399 
* altocaulis 399 
varia 392 
“ colorata 392 
Vaseyi 433 
venusta minor 409 
verna 391 
verrucosa 419 
vesicaria 432-3 
vestita 426 
“ Kennedyi 426 
virescens 413 
“ costata 413 
* Swanit 413 
viridula 429 
vulgaris 422 
“ strictiformis 422 
vulpina 369 
vulpinoidea 369 
Walteriana 426 
Willdenovii 387 
xanthocarpa 369 
“Vv. annectens 369 
xanthophysa 431 
xerantica 386 
xerocarpa 420 
Carpinus I 606 
Betulus 606 
caroliniana 606 
virginiana 606 
Carum 2 658 
Carui 659 
Carya I 579 
alba 581 
amara 580 
aquatica 581 


nucrocarpa 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Carum 
olivaeformis 1 580 
porcina 583 
sulcata 582 
tomentosa 582 
see Hicoria 1 580-3 
CARYOPHYLLACEAE 2 61 
Cassandra 2 686 
calyculata 687 
Cassia 2 335 
Chamaecrista 337 
“robusta 337 
fasciculata 337 
fistula 335 
marylandica 336 
Medsgeri 336 
nictitans 337 
obtusifolia B35 
occidentalis 336 
Tora 335 
Cassiope 2 685 
hypnoides 686 
tetragona 636 
Castalia 2 79 
(Nymphaea) 
Leibergii 80 
odorata 79 
“ minor 79 
“ rosea 79 
pudica 79 
pygmaea 80 
tetragona 80 
tuberosa 80 
Castanea I 615 
dentata 615 
pumila 616 
vescaamericana 615 
vulgaris 615 
Castilleja 3 214 
acuminata 215 
affinis minor 215 
coccinea 214 
fissifolia 214 
indivisa 215 
minor 215 
pallida 215 
Septentrionalis 215 
sessiliflora 216 
Catabrosa I 245 
aquatica 245 
Catalpa 3 238 
bignonioides 238 
Catalpa 238 
cordifolia 238 
speciosa 238 
Cathartolinum 2 436 
Berlandieri 439 
catharticum 438 
floridanum 438 
medium 438 
rigidum 439 
striatum 437 
sulcatum 439 
virginianum 437 
Caucalis 2 626 
Anthriscus 626 
nodosa 626 
Caulinia 
flexilis I 90 
guadalupensis 90 
Caulophyllum 2 128 
thalictroides 128 
Ceanothus 2 504 
americanus 504 
ovalis 505 
ovatus 505 
“ pubescens 505 
Cebatha 2 131 
(Cocculus) 
carolina 131 
Cedronella 
cordata 3 113 


569 


CELASTRACEAE 2 490 
Celastrus 2 492 
bullatus 493 
scandens 493 
Celeri 2 659-60 
graveolens 660 
Celosia 
argentea 26 
paniculata 8 
Celtis I 628 
australis 628 
canina 629 
crassifolia 629 
georgiana 630 
Mississippiensis 629 
occidentalis 629 
pumila 629 
reticulata 630 
Smallii 629 
Cenchrus I 166 
carolinianus 167 
echinatus 166 
macrocephalus 167 
racemosus I21 
tribuloides 167 
Centaurea 3 556 
americana 559 
benedicta 560 
Calcitrapa 559 
Cyanus 557 
Jacea 557 
maculata 3.558 
melitensis 559 
nigra 557 
scabiosa 558 
solstitialis 560 
vochinensis 558 
Centaurium 3° 
calycosum 3 
Centaurium 2 
exaltatum 3 
pulchellum 2 
spicatum 2 
texense 3 
Centaurella 
paniculata 17 
verna 3 17 
Centella 2 651 
(Hydrocotyle 
in part) 
asiatica 651 
repanda 651 
villosa 651 
Centrosema 
virginianum 2 417 
Centunculus 2 716 
minimus 717 
Cephalanthus 3 255 
occidentalis 255 
Cerastium 2 47 
alpinum 49 
aquaticum 2 
arvense 49 
“oblongifolium 49 
“villosum 49 
brachypodum 48 
“compactum 48 
cerastioides 50 
Fischerianum 49 
glomeratum 47 
longipedunculatum 48 
nutans 48 
oblongifolium 49 
semidecandrum 47 
trigynum 50 
triviale 48 
velutinum 49 
viscosum 47 
vulgatum 47,8 
Cerasus 2 322 
Besseyt 2 327 
Mahaleb 328 


570 GENERAL INDEX OF LATIN GENERA AND SPECIES. Voi. III. 
Cerasus Chamaesaracha Chiogenes 2 703 | Cicuta 
pennsylvanica 2 328 sordida 3 163 hispidula 704 bulbifera 2 658 
pumila 2 326 | Chamaesyce 2 462 japonica 704 maculata 658 
Ceratochloa albicaulis 2 466 serpyllifolia 704 virosa 657 
breviaristata 1 280 Fendleri 465 | Chionanthus 2 728 “ maculata 658 
CERATOPHYLLACEAE Geyeri 463 virginica 728 | Cimicifuga 2 90 
AG fs glyptosperma 466 | Chironia americana 92 
Ceratophyllum 75 humistrata 467 see Sabbatia 3 5-6 cordifolia QI 
demursum 75 lata 465 | CHLORIDEAE I 108-9 foetida - 90 
Cercis 2 334 maculata 467 | Chloris I 225 palmata 103 
canadensis 335 maritima 462 curtipendula 228 racemosa QI 
Cercocarpus 2 274 petaloidea 464 elegans 225 “ dissecta QI 
parviflorus 274 “ Nuttallii 464 monostachya 224 | Cineraria 
Cerefolium 2 62 polygonifolia 463 verticillata 225 maritima Sa5e7, 
Cerefolium 620 Preslii 468 | Chondrilla 3 314 palustris 540 
Ceresia Rafinesqui 467 juncea 314 | Cinna I 202 
fluitans I 126 serpens 464 | Chondrophora 3 375 arundinacea 202 
Cereus serpyllifolia 465 (Bigelovia in part) glomerata 116 
caespitosus 2 569 stictospora 466 Nauseosa 37 lateralis 116 
viridiflorus 569 zygophylloides 464 nudata 375 latifolia 202 
“ tubulosus 569 | Cheilanthes re Be virgata 375 pendula 202 
Chaenorrhinum 3 176 alabamensis 34 | Chondrosea 2 223 | Circaea 2 610 
minus 176 dealbata 35 Aizoon GAR} alpina 2 611 
Chaerophyllum 2 629 Feei 34 pyramidalis . 223 intermedia 61L 
procumbens 62 gracilis 34 | CHORIPETALAE Tey lutetiana 611 
“ Shortii 62 lanosa 34 | Chrosperma 1 489 | Cirsium 3 548 
reflexum 630 anuginosa 34 (Amianthium) altissimum 549 
sativum 629 micropteris 33 muscaetoxicum 489 arvense 553 
sylvestre 630 tomentosa 35 | Chrysanthemum 3518 canescens 551 
Teinturieri 630 vestita 34 arcticum 518 canum 554 
texanum 630 | Cheiranthus Balsamita 519 discolor 549 
Chaetochloa I 164 cheiranthoides 2 172 coronarium 518 Flodmani 551 
(Setaria) Cheirinia 2 172 grandiflorum 520 iowense 549 
glauca 165 aspera 173 inodorum 520 heterophyllus 548 
imberbis 165 cheiranthoides 172 Leucanthemum 518 Hillii 552 
italica 166 inconspicua * 172 Parthenium 519 horridulum 553 
magna 166 repanda 173 segetum 519 lanceolatum 549. 
occidentalis 165 | Chelidonium 2 141 | Chrysaspis 2 353 muticum 553 
versicolor 165 diphyllum 140 | Chrysocoma nebraskense 3 552 
verticillata 164 Glaucium 140 gigantea ochrocentrum 551 
viridis 165 majus 141 graminifolia 3 398 odoratum 55 
Chaetopappa 3 403 | Chelone 3 180 nudata 375 Pitcheri 55 
asteroides 403 Digitalis 184 | Chrysogonum 3 462 palustre 3554 
modesta 403 glabra 181 peruvianum 466 plattense 551 
Chamaecistus 2 683 hirsuta 182 virginianum 463 undulatum 550 
(Loiseleuria) 683 Lyoni 181 “ dentatum 463 virginianum 550 
procumbens 683 obliqua 181 | Chrysopogon 420 | Cissus 2 Octo 
serpyllifolia 683 Pentstemon 184 avenaceus 120 Ampelopsis 509 
Chamaecrista 2 337 | CHENOPODIACEAE 2 8 | Chrysopsis 3,372 incisa 510 
depressa 337 | Chenopodium 29 alba 432 stans 510 
fasciculata 337 album EO; angustifolia 374 vitaginea 510 
nictitans Ber altissimum 2 camporum 374 | CISTACEAE 2 539 
Chamaecyparis 1 65 ambrosioides 14 falcata 373 | Cistus 
sphaeroidea 65 anthelminticum 14 foliosa 374 canadensis 540 
thyoides 65- Berlandieri 10 gossypina 20S Ledon 676 
Chamaedaphne 2 686 Bonus-Henricus 14 graminifolia 373 | Citrullus 3 2901 
(Cassandra) Boscianum 12 hispida 375 Citrullus 291 
calyculata 687 Botrys 14 mariana 374 | Cladium 347 
Chamaelirium 1 488 capitatum 15 Nuttallii 375 mariscoides 348 
carolinianum 489 Fremontii 12 pilosa 375 effusum — 348 
luteum 489 glaucum 10 pilosa 373 jamaicense 348 
obovale 489 humile 13 stenophylla 374 | Cladothrix 27 
Chamaenerion 2 589 hybridum 13 villosa 374 lanuginosa 7 
(Epilobium in part) incanum 10 “hispida | 375 | Cladrastis 2 343 
angustifolium 590 leptophyllum 11 | Chrysosplenium 2 229 fragrans 343 
latifolium 590 “ subglabrum II alternifolium 230 lutea 343 
Chamaepericlymenum maritimum 21 americanum 230 tinctoria 343 
2 664 multifidum 15 iowense 230 | Claytonia 2 37 
canadense 664 murale 13 oppositifolium 229-30 caroliniana 38 
suecicum 665 oblongifolium 11 | Chrysothamnus 3 376 Chamissoi 38 
Chamaeraphis paganum 10 (Bigelovia in part) lanceolata 38 
glauca I 165 polyspermum II graveolens 376 perfoliata 39 
ttalica 166 Scoparia 16 Howardi 376 virginica EV, 
verticillata 164 rubrum O,13 nauseosa 3 376 | CLEMATIDEAE 2 84 
viridis 165 urbicum 12 pumilus 376 | Clematis 2 121-2 
Chamaerhodos 2 263 viride 10 | Cicendia Addisonii 123 
altaica 263 Vulvaria II exaltata 2S Catesbyana 122 
erecta Nuttallii 263 | Chimaphila 2 672 | CICHORIACEAE 3 304 crispa 123 
Nuttallii 263 corymbosa 672 | Cichorium 305 cylindrica 123 
Chamaesaracha 3 163 maculata 672 Intybus 305 Douglassii 126 
conioides 163 umbellata 672 _ “ divaricatum 305 Fremontti 125 
Coronopus 163 | Chimonanthus 2 132! Cicuta 2 657 glaucophylla 124 


Vo. III. 


Clematis 
hirsutissima 2 102 
ligusticifolia 122 
missouriensis 122 
ochroleuca 125 
ovata 125 
Pitcheri 123 
Scottii 126 
sericea 125 
Simsit 123 
versicolor 124 
verticillaris 126 
virginiana 122 
Viorna I22,4 
viornoides 123 
vitalba 121 
Cleome 2 196 
dodecandra 199 
gynandra 196 
integrifolia 197 
lutea 197 
pinnata 2 169 
pungens 197 
serrulata 197 
spinosa 197 
Cleomella 2 10908 
angustifolia 198 
mexicana 198 
CLETHRACEAE 2 666 
Clethra 2 667 
acuminata 667 
alnifolia 667 
Clinopodium 3 138 
(ine. Calamintha) 
Acinos 3 139 
Calamintha 139 
glabrum 139 
glabellum 139 
incanum 139 
Nepeta 138 
vulgare 138 
Clintonia 1 idl 
borealis 514 
ciliata 515 
unbellata 515 
umbellulata 515 
Clitoria 2 416 
mariana 417 
Ternatea 416 
virginiana 417 
Clypeola 
alyssoides 2 151 
maritima 153 
Cnicus. 3 560 
seeCirsium 3 554 
benedictus 560 
horridulus 553 
lanceolatus 549 
pumilus 552 
Cnidoscolus 2 462 
humosus 462 
stimulosus 462 
Cobresia tT) 451 
Cocculus 2 131 
carolinus 131 
Cochlearia 2 163 
aquatica 164 
Armoracia 163 
Coronopus 167 
danica 163 
oblongifolia 163 
officinalis 163 
Coeloglossum I 552 
bracteatum 552 
viride 552 
Coelopleurum 2 645 
actaeifolium 646 
Gmelini 646 
Coelorachis Terry 
cylindrica 112 
rugosa 112 
Cogswellia 2 631 


Cogswellia 
daucifolia +2 632 
foeniculacea 632 
nudicaulis 631 
orientalis 631 
villosa 632 
Coix 
dactyloides Tes 
Coloesanthus 3 363 
(Brickellia) 
Cavanillesii 363 
grandiflorus 363 
umbellatus 363 
Collinsia 3 188 
bicolor 188 
parviflora 189 
tenella 189 
verna 188 
violacea 188 
Collinsonia 3 953 
canadensis 153 
Collomia 3 63 
linearis 64 
mucrantha 59 
see Gilia 59-64 
Colpodium 
latifolium I 201 
Colutea 
arborescens 2376 
Comandra I 639 
livida 639 
pallida 639 
Richardsiana 640 
umbellata 639-40 
Comaropsis 
fragarioides 2 269 
Comarum 2 258 
palustre 258 
COMMELINACEAE I 457 
Commelina 457 
agraria 457 
caroliniana 457 
communis 457-8 
crispa 458 
dubia 464 
erecta 459 
hirtella 459 
longifolia 459 
nudiflora 457 
virginica 458 
Willdenovii 458 


COMPOSITAE 3 347-560 


Comptonia I 585 
asplenifolia 586 
peregrina 586 

CONIFERAE I 55 

Conioselinum 2 635 
canadense 636 
chinense 636 
pumilum 636 
tataricum 635 

Conium 2 653 
maculatum 653 

Conobea 3 191 
aquatica IOI 
multifida I9I 

Conoclinium 
coelestinum 3 362 

Conopholis 3 235 
americana 235 

Conostylis 
americana I 535 

Conringia 2 174 
orientalis 175 
perfoliata 175 

CONVALLARIACEAE I 513 

Convallaria I 522 
biflora 521 
bifolia 516 
commutata 521 
majalis 22 
majuscula 522 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Convallaria 
Polygonatum 1 521 
racemosa 5s 
stellata 516 
trifolia 516 

CONVOLVULACEAE 3 40 

Convolvulus 3 45 
aquaticus 40, I 
arvensis 47 
camporum 47 
carolinus 44 
fraternifolius 46 
hunustratus AI 
incanus 47 
interior 46 
japonicus 46 
panduratus 43 
Pickeringii 41 
purpureus 45 
repens 46 
sepium 45-6 
“ americanus 46 
“fraternifiorus 46 
“repens 46 
spithamaeus 47 

Conyza 
asteroides 3 406 
bifoliata 406 
linifolia 3 405 
marylandica 3 447-8 

Cooperia I 532 
Drummondii 533 

Coprosmanthus 
herbaceus TA527, 
tamnifolius 527 

Coptis 2 88 
trifolia 88 

Corallorhiza I 574 
Corallorhiza 574 
innata 574 
Macraei 576 
maculata B75 
multiflora 575 
Neottia 574 
ochroleuca 576 
odonthorhiza 565 
striata 76 
trifida 574 
Wisteriana 575 

Corema 2 479 
Conradii 479 

Coreopsis 3 489 
alternifolia 487 
aristata 499 
aristosa 499 
Atkinsoniana 3 493 
aurea 498 
auriculata 491 
Bidens 495 
bidentoides 496 
cardaminefolia 493 
coronata 3 4908 
crassifolia 490 
delphinifolia 493 
discoidea 497 
grandiflora 491 
lanceolata 489, 90 
“ angustifolia 490 
“ uillosa 490 
involucrata 499 
major 3 402 
“ Oemleri 492 
palmata 492 
pubescens 490 
rosea 494 
senifolia 492 
stellata 492 
tinctoria 493 
tripteris 491 
verticillata 492 
trichosperma 408 | 
“ tenuiloba 408 


571 
Coreopsis 
trifida 3 500 
see Bidens 
: 3 494-500 
Coriandrum 2 646-7 
sativum 647 
Corispermum 2 20 
hyssopifolium 21 
microcarpum 21 
nitidum 21 
CoRNACEAE 2 660 
Cornella 2 664 
canadensis 664 
florida 664 
suecica 665 
Cornucopiae 
altissima I 206 
hiemalis 207 
perennans 206 
Cornus 2 660 
alba 662 
alternifolia 663 
Amomum 661 
asperifolia 662 
Baileyi 662 
canadensis 664 
candidissima 663 
circinata 661 
Drummondii 662 
fastigiata 663 
femina 663 
florida 664 
Mas 660 
obliqua 661 
paniculata 663 
Purpusi 661 
Priceae 661 
rugosa 661 
sericea 661 
stolonifera 662 
stricta 663 
suecica 665 
unalaskensis 664 
Coronilla 2 391 
valentina 391 
varia 392 
Coronopus 2 166 
Coronopus 2 167 
didymus 167 
CORRIGIOLACEAE 2 26 
Corydalis 2 143-6 
canadensis 142 
glauca 144 
see Capnoides 
2 143-6 
Corylus I 607 
americana 607 
avellana 607 
rostrata 607 
Coryphantha 2 570 
missouriensis 570 
similis 571 
vivipara 571 
Cotinus 2 484 
(Rhus in part) 
americanus 485 
cotinoides 485 
Cotinus 484 
Cotoneaster 2 321 
Pyracantha 322 
Cracca 2 372 
(Tephrosia) 
hispidula 373 
spicata 373 
villosa 372 
virginiana 72 
“ holosericea 372 
Crantzia 
lineata 2 648 
Crassina 3 466 
(Zinnia) 
grandiflora 466 


572 GENERAL INDEX OF LATIN GENERA AND SPECIES. Vot. III. 

CRASSULACEAE 2 205 | Crataegus Crataegus Cucubalus 

Crataegus 2 294-321 intricata 2 307,9 spicata 2 292 Behen 2 64 
acerifolia 2 319 irrasa 304 stolonifera 312 latifolius 64 
albicans 318 “ divergens 304 Stonei 308 NIVeUS 63 
alnorum 311 Jackii 2 306 straminea 309 stellatus 62 
mara 309 Jasperensis 298 submollis 319 | Cucumis 

1 6 Jesupi 312 suborbiculata 301 perennis 3 2901 

pee ceesaey 3 Jonesae 2 300 succulenta 301 | CUCURBITACEAE 290 
aptifolia 320 Kelloggii 315 Tatnalliana 2 318 | Cucurbita 3 291 
apposita 309 lanuginosa 316 tenuifolia 315 foetidissima 291 
arborescens 307 laurentiana 304 tiliaefolia 319 Lagenaria 291 
Arnoldiana 318 leiophylla Bs tomentosa 302, 319 Pepo 291 
ascendens 310 Lettermani 301 “ punctata 300 perennis 291 
aspera 312 lucida 207 uniflora 320 | Cunila 3 145 
asperifolia 303 lucorum 304 Vailiae 303 glabella 139 
beata 2 314 Macauleyae 20305 villipes 315 Mariana 146 
Beckwithae 313 macracantha 301 viridis 307 origanoides 146 
bedfordensis 314 macrosperma 311 “mtida 307 pulegioides 136 
berberidifolia 297 maineana 313 | Crepidium Cuphea 
Bicknellii 305 Margaretta 300 glaucum 3 326 petiolata 2 582 
Biltmoreana 307 Marshallii 320 | Crepis Bases VISCOSIiSSiIMa 582 
Bissellii 309 medioxima 314 biennis 327 | Cupressus 
Blanchardi 304 michiganensis 302 capillaris 327 disticha I 64 
blandita 310 microcarpa 320 glauca 326 imbricaria 64 
Boyntoni 309 Missouriensis 2 303 intermedia 327 thyoides 65 
bracteata 312 modesta 307 nudicaulis 310 | CUSCUTACEAE 3 48 
Brainerdi 303 mollis 319 occidentalis 328 | Cuscuta 48 
brevispina 321 monogyna 319 perplexans 3 326 arvensis 49 
Brownii 300 neofluvialis 302 polymorpha 327 Cephalanthi 50 
Buckleyi 309 nitida 307 pulchra 325 chlorocarpa 49 
Calpodendron 2 302 nitidula 301 riparia 3 326 compacta 52 
canadensis 318 oblongifolia 316 runcinata 326 Coryli 50 
Canbyi 299 occidentalis 301 tectorum 325, 326 cuspidata 51 
campestris 303 opulens 313 VITENS 307, decora 50 
Chapmani 302 ovata 306 | Cristaria densiflora 48 
chrysocarpa 306 Oxyacantha 294, 319 coccinea 2 519 Epilinum 48 
coccinea 307, 317 padifolia 2 308 | Cristatella 2 198 Epithymum 49 
“ mollis 319 pallens 309 erosa 198 europaea 48-9 
* oligandra 314 Palmeri 298 Jamesii 198 glomerata 52 
coccinioides 317 pausiaca 299 | Crocanthemum 2 539 Gronovii 51 
collina 301 parvifolia 320 canadense 540 indecora 50 
coloradensis 301 Peckii 308 carolinianum 539 inflexa 50 
compta 314 pedicellata 317 corymbosum 541 obtusiflora 49 
cordata 321 Pennypackeri 299 majus 540 paradoxa 52 
crudelis 310 peoriensis 299 Crotalaria 2 346 Polygonorum 49 
Crus-galli 297, 302 pertomentosa 302 alba 346 pulcherrima 50 
cuneiformis 299 Phaenopyrum 321 latifolia- 346 rostrata 51 
deltoides 253103 pisifera 302 ovalis 347 tenuiflora 50 
denaria 298 polita 318 Purshii 347 Trifolit 49 
Dewingii 301 polybracteata 309 rotundifolia 347 vulgivaga 51 
dilatata 317°| populifolia 308, 315 sagittalis 347 | Cuthbertia I 459 
disjuncta 314 populnea 312 “ ovalis 347 graminea 459 
disperma 299 porrecta 299 | Croton 2 454 rosea 459 
dispessa 316 Porteri 315 capitatus 454 | Cyclachaena 
Doddsii 306 premorsa 307 glandulosus 454 xanthiifolia 3 341 
Douglasii 321 Pringlei 317 Lindheimerianus 455 | Cyclanthera 3 292 
duracina 313 pruinosa ams monanthogynus 2 455 dissecta 203 
Edsoni 2 311 punctata 300 texensis 455 pedata 292 
effera 314 “ brevispina 321 Tiglium 2 454 | Cycloloma 2 16 
Eggertii 317 Pyracantha 2 322| Crotonopsis 2 456 atriplicifolium 16 
Egglestoni 303 pyriformis 316 linearis 456 platyphyllum 16 
elliptica 299 Reverchoni 298 | CRUCIFERAE 2 146| Cymbalaria 2 175 
Elwangeriana 317 roanensis 310 | Crunocallis 2 38 Cymbalaria 175 
Engelmannii 207 Robbinsiana efits) Chamissonis 38 | Cymbia 3 308 
exclusa 317 rotundata 306 | Crypsis occidentalis 308 
fecunda 2 2099 rotundifolia 306 schoenoides i 190| Cymbidium I 574-5 
Fernaldi 304 “ Bicknellii 305 squarrosa 231 hyemale 574 
fillipes 313 rubrifolia 298 | Cryptantha 3°79 odontorhizon 575 
flabellata 310 rubrisepala 298 (Krynitzkia in part) pulchellum 563 
flava 300, 310 rugosa 313 crassisepala 80 | Cymophyllus I 441 
flexispina 310 rustica 2 313 Fendleri 80 Fraseri 441 
fluviatilis 310 Saundersiana 301 glomerata 79 | Cymopterus 2 644 
foetida 309 scabrida 303 | Cryptogramma 1 32 acaulis 644 
fortunata 309 schizophylla 298 acrostichoides a2 glomeratus 644 
Gattingeri 2 314 Schuettii 303 Stelleri 32 montanus 645 
gemmosa 302 sequaxr Siane: Cryptotaenia Cynanchum 3 36 
glandulosa 301 sheridana 306 canadensis 2 630 (Vincetoxicum) 
globosa 302 silvicola 313 | Ctenium I 224 acutum 3 36 
grandis 209 Smithit 320 americanum 225 carolinense 38 
Grayana 3II sordida 301 | Cubelium 2 563 hirtum 38 
Holmesiana 2 315 speciosa 317 (Solea) nigrum 36 
Holmesii 315 spathulata 320 concolor 564 obliquum 38 


Vor. ITI. 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Cynanchum 
suberosum 30.37 
Vincetoxicum 36 

CYNAREAE 3 350 

Cynoctonum 2 730 
(Mitreola) 

Mitreola 731 
pbetiolatum 731 
sessilifolium 730 

Cynodon 
Dactylon I 222 

Cynoglossum 375 
boreale Wi 
glomeratum 81 
Morisoni 78 
officinale 75-6 
pilosum 78 
virginianum 76 

Cynomarathrum 2 632 
Nuttallii 633 

Cynosciadium 2 646 
pinnatum 646 

Cynosurus I 251 
aegyptius 229 
coracanus 228 
cristatus 251-2 
indicus 229 
virgatus 229 

Cynoxylon 2 664 
floridum 664 

Cynthia 3 307 
(Adopogon) 
Dandelion 308 
falcata 3 307 
virginica 307 

CYPERACEAE I 295 

Cyperus ‘I 297 
acuminatus 302 
aristatus 300 
arundinaceus 340 
Baldwinti I 310 
Bushii 302 
cayennensis 309 
calcaratus 302 
compressus 300 
cylindricus 308 
dentatus 303 
“ctenostachys 303 
diandrus 298 
“ castaneus 298 
“ elongatus 298 
dipsaciformis 307 
echinatus 310 
Engelmanni 306 
erythrorrhizos 305 
esculentus 297, 304 
“ angustispicatus 304 
ferax 306 
filicinus 299 
filiculmis 309 
“ macilentus 309 
flarus 309 
flavescens 298 
flavicomus 300 
fuscus 303 
Gatesit 2909 
globulosus 310 
Grayi 309 
Halei 305 
Hallii I 304 
Haspan 303 
Houghtoni 301 
Hydra 304 
hystricinus 308 
inflexus 300 
Iria 301 
lancastriensis 307 
macilentus 309 
Michauxianus 305 
microdontus 299 
Nuttallit 299 
ovularis 308 


Cyperus 
paniculatus I 299 
phymatodes 304 
pseudovegetus 302 
refractus 307 
tetrofractus 307 
rivularis 298 
rotundus 304 
sabulosus 300 
Schweinitzii 301 
spathaceus 340 
speciosus 305 
strigosus 306 
“ capitatus 306 
“ compositus 306 
“ robustior 306 
Torreyi 308 
Cy PRIPEDIEAE 547, 
Cypripedium I 548 
acaule 550 
album 549 
arietinum 548 
bulbosum 573 
Calceolus 548 
candidum 549 
hirsutum 549 
parviflorum 550 
passerinum 549 
pubescens 550 
reginae 549 
spectabile 549 
CY RILLACEAE 2 485 
Cyrilla 485 
racemiflora 485 
Cyrtorhyncha 2 117 
(Ranunculus in part) 
ranunculina 117 
Cystopteris Tein: 
bulbifera 15 
fragilis 15 
montana 15 
Cytherea I 572 
bulbosa 573 
Cytisus 2 350 
rhombifolius 344 
scoparius 350 
Dactylis I 251 
glomerata 251 
maritima 224 
patens 223 
stricta 224 


see Spartina 1 223-4 
Dactyloctenium 1 22« 


aegyptium 229 
aegyptiacum 229 
Dalea 2 366 
alopecuroides 267 
enneandra 267 
lanata 267 
lanuginosa 367 
laxiflora 367 
obovatifolia 366 
parviflora 2 384 
see Parosela 2 366-8 
Petalostemon 

2 368-71 

Dalibarda 2 282 
fragarioides 269 
repens 282 
violaeoides 282 
Danthonia I 220 
Alleni 221 
compressa 221 
epilis 221 
glabra 221 
intermedia 222 
sericea 221 
spicata 220 
Daphne 2574 
Gnidium 574 
Mezereum 574 


Darbya I 641 
unbellulata 641 
Darlingtonia 29331 
Dasiphora 2 262 
fruticosa 262 
riparia 262 
Dasystephana 3 10 
(Gentiana in part) 
affinis Ch Gari 
Andrewsii 12 
flavida 12 
Grayi 13 
latifolia II 
linearis 12 
parvifolia igi 
Porphyrio 14 
puberula II 
rigida II 
saponaria 12 
villosa - 13 
Dasystoma 3 205 
(Gerardia in part) 
aurea 206 
calycosa 3 208 
Drummondii 207 
flava 206 
grandiflora 207 
laevigata 207 
pectinata 206 
pedicularia 206 | 
pubescens 206 
quercifolia 208 
“integrifolia 207 
serrata 207 
virginica 208 
Datisca 
hirta 2 481 
Datura 3 169 
Metel 170 
Stramonium 169 
Tatula 169 
Daucus 2 625 
Carota 625 
divaricatus 651 
pusillus 625 
Decamerium 2 6905 
Decodon 2 579 
(Nesaea) 
aquaticus 580 
verticillatus 580 
Decumaria 2 232 
barbara 233 
Delphinium 2 93 
Ajacis 93-4 
albescens 95 
azureum 95 
carolinianum 95 
Consolida 93 
exaltatum 94 
Nelsoni 95 
Nortonianum 95 
Penardi 95 
Treleasei 904 
tricorne 96 
urceolatum 04 
virescens 95 
Dendrium 2 682 
(Leiophyllum) 
buxifolium 682 
Dendropogon I -456 
usneoides 456 
Dennstaedtia 2 14 
punctilobula 14 
Dentaria 2 187 
anomala 188 
diphylla 188 
furcata 187 
heterophylla 188 
incisifolia 188 
laciniata 187-8 
maxima 188 
pentaphyllos 187 


bra 
Dentidia 
nankinensis 3 154 
Deringa 2 630 
(Cryptotaenia) 
canadensis 630 
Deschampsia I 215 
atropurpurea 216 
caespitosa 215 
flexuosa 216 
Descurainia MT te fat 
Hartwegiana 171 
incisa 171 
pinnata T7r 
Sophia 170 
Desmanthus 2 331 
brachylobus 332 
leptolobus 332 
Desmodium 
see Meibomia 
‘ 394-402 
acuminatum 2 396 


canadense hirsut. 398 


“ longifolium 399 
ciliare 402 
cuspidatum 3909 
humifusum 307 
lineatum 396 
Deveuxia 
see Calamagrostis 
I 207 
Macouniana 210 
Dianthera 3 242 
americana 242- 
humilis 243 
lanceolata 243 
ovata 243 
Dianthus 2 73 
Armeria 74 
barbatus 75 
caryophyllus 73 
deltoides 74 
prolifera 74 
Saxifraga 2 
Diapedium 243 
(Dicliptera) 244 
brachiatum 244 
DIAPENSIACEAE 2 705 
Diapensia 2 705-6 
barbulata 706 
lapponica 706 
Diaperia 
prolifera 3 446 
Diarina I 247 
Diarrhena 
americana I 247 
Dicentra 


see Bicuculla 2 141-3 


Dichondra 3 339 
carolinensis 40 
evolvulacea 40 
repens 39, 40 

DICHONDRACEAE 3 39 

Dichromena I 340 
colorata 341 
latifolia 341 
leucocephala 340 

Dicksonia I 14 

Dichrophyllum 2 468 
marginatum 469 

Dicksonia 
pilosiuscula Ter. 
punctilobula 14 

Dicliptera 3 244 
brachiata 244 

Diclytra 
seeBicuculla 2 141-3 

DICOTYLEDONES I 577 

Didiplis 2 579 
diandra 579 
linearis 579 

Dielytra 
Cucullaria 2 142 


574 GENERAL INDEX OF LATIN GENERA AND SPECIES. Vorsuiie 
Diervilla 3 283 | Ditaxis Dracopis Echinocactus 
Diervilla 283 Mercurialina 2 456 amplexicaulis 3 473 Simpsoni 2 570 
Lonicera 3 283 | Dodecatheon 2 717 | DROSERACEAE 2 202 | Echinocereus 2 568 
trifida 283 brachycera 717 | Drosera 2 203 caespitosus 569 
Dieteria Hugeri 717 americana 203 viridiflorus 568-9 
sessiliflora 3 435 Meadia 717 anglica 204 | Echinochloa Theos 
Digitalis 3 204 “ Frenchii 717 brevifolia 204 colona 134 
purpurea 204 | Doellingeria 3 443 filiformis 204 Crus-galli 133 
Digitaria Pere, (Diplopap pus) intermedia 203-4 Walteri 134 
filiformis 122 amygdalina 443 linearis 204 | Echinocystis 3 2092 
humifusa 123 humilis 443 longifolia 203-4 lobata 292 
paspaloides 133 infirma 444 rotundifolia | 203-4 | Echinodorus I 95 
Sanguinalis 123 ptarmicoides 432 “ comosa - 203 cordifolius 96 
Dioclea umbellata 443 tenuifolia 204 “ lanceolatus 96 
multiflora 2 420 pubens 443 | DRUPACEAE 2 320 parvulus 95 
Diodia 3 256] Dolicholus 2 421 | Dryas 2 273 radicans 96 
teres 256 erectus 22 chamaedrifolia 273 rostratus 96 
virginiana 257 latifolius 421 Drummondii 274 tenellus 95 
Diodonta minimus 421 integrifolia 274 | Echinopanax 2 619 
bidentoides 3 496 simplicifolius 422 octopetala 273 horridum 619 
DIOSCOREACEAE I 535 tomentosus 421 tenella 274 | Echinops 
Dioscorea I 535 | Dolichos 2 417 | Drymocallis 2 262 sphaerocephalus : 
paniculata 535 Catjang 425 agrimonioides 263 3 560 
Sativa 535 Lablab 417 rubricaulis 263 | Echinospermum 3 77 
villosa 535 luteolus 424 | Dryopteris 1 17, deflexrum am. 379 
Diospyros 2 720 polystachyus 423 acrostichoides 16 Lappula Hei 
Lotus 720 regularis 420 aculeata 17 Redowskii 78 
virginiana 720 sinensis 425 Boottii 22 see Lappula 77-8 
Diotis Dondia 2 23 Braunii 17 | Echites 
lanata 2 20 (Suaeda) Clintoniana 20 difformis 3 23 
Diphylleia 2 128 americana 23 cristata 19 | Echium 3 93 
cymosa 129 calciformis 2 24 dilatata 21 italicum 93 
Diplachne I 236 depressa 2 Dryopteris 23 vulgare 94 
acuminata 237 linearis 23 Filix-mas 21 | Eclipta 
fascicularis 236 maritima 24 fragrans 19 alba 468 
maritima 237 | Donia Goldiana 20 erecta 468 
procumbens 237 ciliata RES 77 hexagonoptera 23 procumbens 468 
rigida 231 squarrosa 371 intermedia I 21 | ELAEAGNACEAE 2enrir 
Diplogon 3 372 | Doronicum Linneana 23 | Elaeagnus 2 575 
Diplopappus 3 443 acaule 3 533 Lonchitis 16 angustifolia 575 
see Doellingeria 443 ramosum 440 marginalis 20 argentea 576 
canescens 437 | Doxantha noveboracensis 18 canadensis 576 
cornifolius 444 capreolata 3 337 Phegopteris 23 | ELATINACEAE 2 537 
hispidus 375 | Draba 2 148 pittsfordensis 22| Elatine 3 538 
linariifolius 444 alpina 152 Robertiana 2 americana 538 
umbellatus 443 androsacea 149 simulata 19 brachysperma 538 
Diplotaxis 2 194 arabisans 150 spinulosa Bi Hydropiper 538 
muralis 194 “ orthocarpa 150 Thelypteris 18 triandra 538 
tenuifolia 194 aurea 152 | Duchesnea 2 259 | Elatinoides 3.175 
DIPSACEAE 3 288 brachycarpa rife (Fragaria in part) (Linaria in part) 
Dipsacus 3 288 caroliniana 149 indica 259 see Kickxia 175 
fullonum 289 confusa 150-1 | Dulichium I 340 Elatine 176 
“sativus 289 corymbosa 149 arundinaceum 340 spuria 175 
laciniatus 289 crassifolia 152 spathaceum 340 | Eleocharis I 310 
sylvestris 289 cuneifolia 149 | Dupatya I 455 acicularis 315 
Dipteracanthus fladnizensis 149 (Paepalanthus) acuminata 318 
biflorus 240 hirta 150 flavidula 455 albida 317 
ciliosus var. parvi- hispidula 149 | Dupontia I 261 atropurpurea 312 
florus 242 incana 150-1 | - Cooleyi 262 capitata 313 
micranthus 241 megasperma 151 Fisheri 261 compressa 318 
Dirca 2 574 micrantha 149 | Dyschoriste 3 240 diandra 313 
occidentalis 574 nemorosa 152 depressa 240 dispar 313 
palustris 575 nivalis 150 oblongifolia 240 Engelmanni 314 
Discanthera pycnosperma 151 | Dysodia 3 512 equisetoides 311 
dissecta 3 293 ramosissima 151 chrysanthemoides 513 flaccida 312 
Discopleura 2 656 stylaris I51 papposa 513 glaucescens 314 
capillacea 657 verna 148 intermedia 318 
Nuttallii 657 | Dracaena Eatonia “ Habereri 318 
Disporum Tes Ly, borealis I 514 see Sphenopholis interstineta 311 
(Prosartes) umbellulata 515 I 243 macrostachya 314 
lanuginosum 517 | Dracocephalum 3 116 Dudleyi "244 Macounii 318 
pullum 517 cordatum 113 glabra 244 melanocarpa 317 
trachycarpum 518 denticulatum 117 pennsylvanica 244 microcarpa 316 
Distegia intermedium TL7 pubescens 244 monticola 314 
involucrata 3 282 Moldavica 114-5 robusta 244 mutata 311 
Distichlis I 249 Nuttallii 117 | EBENACEAE 2 720 nitida 318 
maritima 249-50 parviflorum 114, 7 | Echinacea 3.475 obtusa 313 
spicata 250 speciosum 116 angustifolia 476 “ jejuna 313 
stricta 250 virginianum 116 pallida 476 ochreata 312 
Ditaxis 2 456 | Dracontium paradoxa 476 olivacea 312 
fasciculata 456 foetidum I 445 purpurea 475 ovata z 353 
humilis 457 | Dracopis 3 473 | Echinocactus 2 569 “Engelmannt 314 


Vor. ITI. 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Eleocharis 
palustris I 314 
“ vigens 314 
pauciflorus 327 
Pygmaea 327 
gquadrangulata 311 
Robbinsii Hiei 
rostellata 319 
simplex 316 
Smallii 315 
tenuis 318 
Torreyana 316 
tortilis 316 
tricostata 317 
tuberculosa 316 
Wolf 315 
Eleogenus 
ochreatus 312 
Elephantella 3 222 
groenlandica 222-3 
Elephantopus 3 353 
carolinianus 354 
( nudatus 354 
S scaber 353 
tomentosus 354 
Eleusine I 228 
aegyptia 229 
indica 229 
mucronata 230 
Ellisia 
microcalyx 3 67 
Nyctelea 67 
Elodea_ I 104 
see Philotria 104 
canadensis 105 
latifolia 105 
Elodes 2 537 
campanulata 537 
petiolata 537 
virginica 537 
Elsholtzia 2) iy! 
cristata 154 
Patrinii 154 
Elymus I 288 
arenarius 288, 90 
australis 292 
arkansanus 204 
brachystachys 293 
canadensis 293 
“ glabriflorus 292 
“ intermedius 292 
condensatus 290 
curvatus 290 
’ diversiglumis 294 
elymoides 288 
glabriflorus 292 
glaucifolius 293 
glaucus 2890 
halophilus 291 
hirsutiglumis 292 
Hystris 294 
jejunus 291 
Macounii 289 
robustus 293 
sibiricus 289 
“ americanus 289 
Sitanion 288 
striatus 293 
© Ballit 293 
“ villosus 293 
virginicus 291 
“ Jejunus 291 
“ submuticus 290 
vulpinus 289 
Elyna T Ser 
Bellardi 351 
spicata 351 
EMPETRACEAE 2 478 
Empetrum 2 479 
Conradii 479 
nigrum 479 
Endorima 3 502 


Endorima 
uniflora 3 502 
Enemion 2 89 
biternatum 89 
Engelmannia 3 464 
pinnatifida 464 
Enslenia 
albida 3 36 
Entrema 
arenicola 2 179 
Epibaterium 2 131 
carolinum 131 
pendulum Tis 
EPIDENDREAE I 548 
Epifagus 3 236 
americana 236 
Epigaea 2 692 
repens 692 
Epilobium 2 590 
adenocaulon 593 
alpinum 591 
“ majus 594 
anagallidifolium 591 
angustifolium 589-90 
coloratum 593 
densum 592 
glandulosum 593 
hirsutum 590, I 
Hornemanni 594 
lactiflorum 591 
latifolium 590 
lineare 592 
molle 592 
oliganthum 592 
palustre 592 
“ lineare 592 
paniculatum 593 
spicatum 590 
strictum 592 
Epipactis I 563 
convallarioides 567 
decipiens 570 
latifolia 563 
pubescens 570 
repens ophioides 569 
tesselata 569 
viridiflora 563 
Epiphegus 3 236 
virginiana 236 
EQUISETACEAE I 38 
Equisetum I 39 
arvense 39 
fluviatile 41, 39 
hyemale 41 
laevigatum 42 
limosum 41 
littorale 40 
palustre 40 
pratense 39 
robustum 41 
scirpoides 42 
sylvaticum 40 
variegatum 42 
Eragrostis I 238 
campestris * 241 
capillaris 239 
capitata 243 
caroliniana 240 
curtipedicellata 242 
Eragrostis 240 
erythrogona 239 
Frankii 239 
hirsuta I 239 
hypnoides 243 
major 241 
minor 240 
oxylepis 242 
pectinacea 241 
“ spectabilis 241 
pilosa 240 
poaeoides 240 
“‘megastachya 241 


575 
Eragrostis Eriocoma_ , 
Purshii I 240 cuspidata I 174 
tefracta 241 | Eriogonum I 647 
reptans 243 alatum 648 
secundiflora 242 Alleni 649 
sessilispica 231 annuum 651 
Weigeltiana 243 brevicaule 651 
tenuis 239, 42 campanulatum 651 
trichodes 242 cernuum 652 
Eranthis 2 87-8 corymbosum 651 
hyemalis 88 effusum 652 
Erechtites 30535 flavum 649 
hieracifolia 536 gnaphaloides 650 
prealta 536 Gordoni 652 
Erianthus Tels Jamesii 649 
alopecuroides 113 lachnogynum 650 
brevibarbis 114 Lindheimerianum 
compactus 114 I 651 
contortus 114 longifolium 648 
divaricatus 113 micranthum 651 
japonicus Tins: microthecum 652 
saccharoides 114 multiceps 650 
ERICACEAE 2 675 parvifiorum 650 
Erica 2 604 pauciflorum 650 
cinerea 694 sericeum 649 
Tetralix 694 texanum 648 
vulgaris 694 tomentosum 648 
Erigenia 2 653 | Erioneuron easy 
bulbosa 653 pilosum 235 
Erigeron 3 435 | Eriophorum I 322 
acris 436, 41 alpinum 32 
“ debilis 441 angustifolium 325 
“ Droebachianus 441 callithrix 324 
annuus 440 capitatum 32 
asper 436 Chamissonis 323 
Bellidiastrum 440 “ albidum 32 
bellidifolius 439 cyperinum 337 
caespitosus 437 gracile 32 
camphoratum 448 hudsonianum 32 
canadensis 442 opacum 324 
canus 438] paucinervium 325 
carolinianum 400 polystachyon 325 
divaricatus 442 “ latifolium 325 
divergens 3 439 russeolum 323 
Droebachianus 441 Scheuchzeri 323 
flagellaris 3 438 tenellum 325 
glabellus 436 triquetrum 324 
* mollis 437 vaginatum 322-4 
hyssopifolius 438 virginicum 326 
nudicaulis 441 * album 326 
philadelphicus 439 viridicarinatum 325 
pilosum 373 | Eritrichium 
pulchellus 439 californicum 
pumilus 3 437 “ subglochidiatum 
ramosus 440 3 79 
“ Beyrichit 440 crassisepalum 80 
strigosus 440 glomeratum 81 
subtrinervis 437 “ humile 81 
tenuis 3 441 fulvocanescens 82 
uniflorus 436 Jamesii 81 
vernus 441 | Erodium 2 430 
Eriocarpum 3 378 cicutarium 430 
(Aplopappus in part) crassifolium 430 
grindelioides 378 moschatum 430 
rubiginosum 378 | Erophila 
spinulosum 379 vulgaris 2 148 
ERIOCAULACEAE 1 453 | Eruca 2 192 
Eriocaulon I 453 Eruca 192 
anceps 455-6 sativa 192 
articulatum 454 | Ervum 2 412 
compressum 454 hirsutum 411 
decangulare 455 Lens 412 
flavidulum 455 multifiorum 386 
gnaphalodes 454 tetraspermum 411 
Parkeri 454 | Eryngium 2 621 
septangulare 454 aquaticum 622 
umbellatum 455 integrif olium 623 
Eriochloa I 12 Leavenworthii 62 
polystachya 124 maritimum 621 
punctata 124 prostratum 623 
Eriocoma 1173] + virginianum 622 


576 


Eryngium 
yuccaefolium 2 622 
Erysimum 2e073 
Alliaria 170 
arkansanum 173 
asperum 173 
Barbarea 177 
cheiranthoides 172 
inconspicuum 172 
lanceolatum 173 
officinale 174 
orientale 175 
parviflorum 172 
perfoliatum 175 
pinnatum ATTA 
praecox ify 
repandum 173 
syrticolum 172 
virnum 177 
Erythraea wal gal BS 
. see Centaurium I 
calycosa 2 
Centaurium 2 
Douglasii 2 
exaltata 2 
pulchella 2 
spicata 2 
ramosissima 2 
texensis 2 
Erythronium 1 505-6 
albidum 506 
americanum 506 
angustatum 506 
bracteatum 506 
Dens-canis 506 
mesachoreum 507 
propullans 507 
Eubotrys 2 688 
racemosa 689 
recurva 688 
Eulalia 
japonica Tiers? 
Eulophus 2 654 
americanus 654 
Euonymus 2 490 
americanus 491 
atropurpureus 491 
europaeus 490, 2 
obovatus 491 
Eu PATORIEAE 3 347 
Eupatorium 35355 
ageratoides 361 
album 358 
* subvenosum 358 
alternifolium 364 
altissimum 359 
aromaticum 362 
boreale 361 
Bruneri 356 
cannabinum 355 
capillifolium 356 
coelestinum 362 
falcatum 357 
foeniculaceum 356 
foeniculoides 35 
glandulosum 358 
grandiflorum 363 
hyssopifolium 358 
“ laciniatum 358 
incarnatum 362 
lecheaefolium 358 
leucolepis 357 
linearifolium 358 
maculatum 3 356 
“ amoenum 356 
perfoliatum 361 
“ cuneatum 361 
“ truncatum 361 
pilosum 360 
pubescens 360 
purpureum 356-7 
“angustifolium 357 


Eupatorium 

bp. falcatum 3 357 
resinosum 361 
rotundifolium 360 
“ ovatum 360 
Rydbergi 356 
scandens 363 
semiserratum 359 
serotinum 357 
sessilifolium 359 
“Brittonianum 359 
Torreyanum 358 
teucriifolium 360 
trifoliatum 357 
urticaefolium 361 
verbeniefolium 360 
“ Saundersii 360 
EUPHORBIACEAE 2 452 
Euphorbia 2 463 

see Chamaesyce 
2 462-8 


“ Dichrophyllum 469 
“Tithymalopsis 469 


“Tithymalus 471 
albicaulis 466 
arenaria 464 
arundelana 476 
corollata 469 
cuphosperma 476 
dendroides 471 
dentata 476 
Fendleri 465 
Geyeri 463 
glyptosperma 464 
heterophylla 477 
hexagona 468 
hirsuta 2 467 
humustrata 467 
hypericifolia 467-8 
lata 465 
maculata 467 
marginata 469 
marylandica 476 
nicaeensis 474 
ohiotica 475 
petaloidea 464 
polygonifolia 463 
Preslit 468 
Rafinesquii 467 
serpens 464 
serpyllifolia 465 
zygophylloides 464 
Euphrasia 2) 217, 
americana 218 
arctica 27, 
canadensis 218 
hirtella 217 
latifolia 217 
Oakesii 218 
Odontites 219 
officinalis 218, 217 
Randii 218 
“ Farlowti 218 
Williamsii 218 
Euploca 
convolvulacea 3 75 
Eurotia 2 19 
lanata 20 
Eurybia 
glomerata 3 410 
Eustoma 37 
silenifolium 7 
Russellianum 8 
Eustylis I 541 
Euthamia 3 398 


(Solidago in part) 


camporum 398 
caroliniana 400 
floribunda 399 
graminifolia 398 
gymnospermoides 

3 399 


| Euthamia 
leptocephala 3 399 
minor 500 
Moseleyi 400 
Nuttallii 398 
remota 400 
tenuifolia 400 
Eutoca 3 69 
Franklinii 69 
Euxolus 2 4-5 
crispus PINS 
deflexus 4 
lividus 4 
pumilus 5 
ol 
prolifera 3 446 
Evolvulus 3 41 
argenteus 42 
nummularius 41 
pilos:'s 42 
FABACEAE 2 341 
FAGACEAE I 614 
Fagopyrum I 671 
esculentum 672 
Fagopyrum 672 
tataricum 671-2 
Fagus I 614 
americana 615 


Castanea dentata 615 


ferruginea 615 
grandifolia 615 
pumila 616 
sylvatica . 614 
Falcata 2 418 
(Amphicarpa) 
caroliniana 419 
comosa 419 
Pitcheri 419 
Fatsia 
horrida 2 619 
Fedia 3 287-8 


see Valerianella 288 


Fagopyrum 287 
patellaria 288 
umbilicata 288 
Ferula 
canadensis 2 647 
foeniculacea 632 
villosa 637 
Festuca I 268 
altaica 273 
brachyphylla 271 
brevifolia 271 
capillata 271 
confinis 27 
decumbens 1233 
diandra 247 
elatior 272 
fascicularis 236 
fluitans 266 
gigantea 273 
Kingii 273 
Myuros 269 
nutans 272 
“ palustris 272 
occidentalis 270 
octoflora 269 
ovina 269, 71 
“ brevifolia 271 
“ capillata 271 
* duriuscula 271 
pratensis 272 
rubra 270 
scabrella 273 
sciurea 270 
Shortii 272 
spicata 272 
tenella 269 
unioloides 280 
Watsoni 278 
FESTUCEAE I 102-13 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Vor. ITI. 
Ficaria 2 117 
Ficaria 117 
ranunculoides LLY, 
verna 117 
Ficus Carica I 632 
Filago 3 446 
germanica 446-7 
prolifera 3 446 
pygmaea 446 
FILICES ba 
Filix I 14 
bulbifera 15 
fragilis 15 
montana 15 
Filipendula 2 248 
denudata 249 
rubra 249 
Ulmaria 249 
Fimbristylis I 319 
acuminata 320 
autumnalis 322 
Baldwiniana 321 
capillaris 319 
castanea 320 
congesta 321 
Frankii 322 
geminata 322 
“ brachyactis 322 
interior 320 
laxa 321 
spadicea 320 
puberula 321 
Vahlii 321 
Fissipes I 550 
acaule 550 
Flaveria 3 504 
angustifolia 504 
campestris 504 
chilensis 504 
Floerkea 2 441 
occidentalis 441 
proserpinacoides 441 
Foeniculum 2 642 
Foeniculum 643 
vulgare 643 
Forestiera 2 728 
acuminata 728 
Forsteronia 
difformis 3°23 
Fothergilla 2 234 
alnifolia 234 
carolina 234i a 
Gardeni 234 
Fragaria 2 259— 
americana 261 
australis 260 
canadensis 259 
Grayana 260 
indica 259 
multicipita 259 
terra-novae 260 
vesca 260 
virginiana 260 
“ illinoensis 260 
Frangula 
caroliniana 2 503 
Franseria 3 342 
acanthicarpa 342 
discolor 343 
Hookeriana 342 
tomentosa 343 
Frasera 3 14 
carolinensis 15 
Fraxinus 2 724 
americana 725 
biltmoreana 725 
campestris 726 
caroliniana 727 
Darlingtonii 725 
excelsior 724 
lanceolata 726 
Michauxir 726 


Vo. III. GENERAL INDEX OF LATIN GENERA AND SPECIES. 577 
Fraxinus Galium Gentiana Geranium 
nigra 2 727 kamtschaticum 3 261 clausa zy ne tenuifolia 3 211 
pennsylvanica 726 labradoricum 264 crinita 9 “ asperula 212 
profunda 726 lanceolatum 260 decora II “ macrophylla 211 
platycarpa 727 latifolium 262 detonsa 9 tuberosa 208 
pubescens 726 Littellit 261 Elliottii II “ viridis 212 
quadrangulata 727 Mollugo 257, 63 flavida 12 see Dasystoma 206-7 
sambucifolia 727 palustre 265 Grayi 13 Agalinis 208-14 
viridis 726 parisiense 258 linearis 13 | Geum 2 269 
Fritillaria I 505 parviflorum 265 “ lanceolata 13 agrimonioides 263 
alba 508 pilosum 260 “ latifolia 13 album 270 
atropurpurea 505 “ puncticulosum 260 lutea 8 “ flavum 270 
pyrenaica 505 septentrionale 261 nesophila_ , 9 canadense 270-1 
Froelichia 27 spurium 259 ochroleuca 13 carolinianum 270 
campestris 27 sylvaticum 263 Porphyrio 14 ciliatum 273 
floridana DG tinctorium 263-4 procera 9 flavum 270 
gracilis 258 “ filifolium 263 propinqua 10 macrophyllum 271-2 
Fuirena Tsay “labradoricum 264 puberula II Peckii 272 
hispida 338 Torreyi 260 pulchella 22 pulchrum 272 
simplex 338 tricorne 259 purpurea 14 radiatum 272 
squarrosa 338 trifidum 264 quinquefolia 10 “ Peckii 272 
“ aristulata 338 “ bifolium 265 * occidentalis 10 rivale 262 
* pumila 338 “ latifolium 263 quinque flora 10 strictum 271 
umbellata 337 “ pusillum 264 ramosissima 2 triflorum 273 
FUMARIACEAE 2 141 triflorum 262 rigida II urbanum 269, 70 
Fumaria 2 146 Vaillantii 259 rubricaulis 13 vernum 271 
Cucullaria 42 verum 258 Saponaria 12 virginianum 270 
eximia 143 yvirgatum 260 “ linearis 13 | Gifola 3 446 
fungosa 143 Wirtgeni 258 scaberrima II germanica 447 
officinalis 146 | Galpinsia 2 606 serrata 9 | Gilia 3 59 
parviflora 146 (Oenothera in part) spicata Bee acerosa 2 
sempervirens 143-4 Hartwegi 606-7 ventricosa 8 aggregata 60 
interior 607 villosa 13 caespitosa 2 
Gaertneria 3 342 lavendulaefolia 2607 | Geoprumnon 2 376 congesta 61 
(Franseria) GAMOPETALAE 2 666 crassicarpum 377 coronopifolia 60 
acanthicarpa 342 | Gaultheria 2 692 mexicanum 277 iberidifolia 61 
discolor 343 procumbens 693 plattense 377 inconspicua 60 
Grayi 343 | Gaura 2 608 tennesseense 378 laciniata 59 
tenuifolia 342 biennis 608-9 | GERANIACEAE 2 425 linearis 64 
tomentosa 343 coccinea 609 | Geranium 2 426 longiflora 60 
Gaillardia 2 Gua filipes 609 Bicknellii 428 minima 64 
acaulis 509 glabra 609 carolinianum 428 pinnatifida 60 
aristata pe linifolia 610 cicutarium 430 pumila 61 
lanceolata 512 marginata 609 columbinum 427 pungens 
lutea 512 Michauxii 609 dissectum 429 “ caespitosa 62 
pulchella iiltin 3 parviflora 608 maculatum 427 rigidula acerosa 62 
simplex 513 Pitcheri 609 molle 429 rubra 60 
suavis 513 sinuata 609 pratense 2 426 spicata 61 
Galactia 2 419], villosa 610 pusillum 429 tricolor 61 
glabella 420 | Gaurella 2 604 pyrenaicium 427 | Gillenia 
mollis 420 canescens 605 Robertianum 426 stipulacea 2 248 
pilosa 420 guttulata 605 rotundifolium 428 trifoliata 248 
regularis 420 | Gaylussacia 2 605 sibiricum 427 | Ginkgo 
‘volubilis 420 baccata 696 sylvaticum 426 biloba 167 
Galax 2 706 brachycera 696 | Gerardia 3 208 | Ginseng 
aphylla 707 buxifolia 695 aspera 209 quinquefolium 2 618 
Galega dumosa 696 auriculata Bu2rs trifolium 619 
see Cracca 372-3 “ hirtella 696 Besseyana 211 | Gisopteris 
Galeopsis 3,119 frondosa 605 crustata 209 palmata I 9 
Ladanum 119 resinosa 696 decemloba 212| Glabraria 2 134 
Tetrahit 119 ursina 6905 densiflora 214 geniculata 135 
Galeorchis I 551 | Gelsemium 2 720 fasiculata 210 tersa 135 
spectabilis 551 nitidum 730 flava 206 | Glandularia 
Galinsoga 3 501 sempervirens 730 Gattingeri 213 bipinnatifida 397 
caracasana 502 | Gemmingia I 541 grandiflora 207 carolinensis 97 
parviflora 502 (Belamcanda) 541 “integriuscula 207 | Glaucium 2 141 
“ hispida 502 (Pardanthus) 541 “ serrata 207 flavum 141 
Galium Bu2s7. chinensis 542 heterophylla 209 Glaucium 141 
anglicum 258 | Genista 2 349 intermedia 210 luteum I4I 
Aparine 259 tinctoria 350 laevigata 207 | Glaux 2 715 
arkansanum 262 | GENTIANACEAE 302 linifolia 209 maritima 716 
asprellum 265 | Gentiana 3 8 Holmiana 212] Glecoma 3 113 
bermudense 266 acuta 39 macrophylla 205, 11 hederacea 114 
boreale 261 affinis II maritima 211 | Gleditsia 2 338 
circaezans 260-1 alba 12 parvifolia 212 aquatica 339 
“ glabellum 261 Amarellaacuta 39 paupercula 210 monosperma 339 
“lanceolatum 260 Andrewsti 12 pedicularia 3 206 triacanthos 338 
“ montanum 261 angustifolia 14 purpurea 208, 10 | Glyceria I 262 
Claytoni 3 264 asclepiadea 10 racemulosa 210 seePanicularia 263-6 
concinnum 265 calycina Sits quercifolia 208 Puccinellia 267 
erectum 263 Catesbaei 12 setacea 212 | Glycine 2 417 
hispidulum 266 Centaurium t;.2 Skinneriana 212 Apios 418 


578 


Glycine 
bracteata 2 419 
comosa 419 
frutescens 374 
Galactia 419 
monoica 419 
Priceana 418 
tomentosa 421 


umbellata 424 


Glycyrrhiza 2 391 
echinata 391 
lepidota 391 


Gnaphalium 3.453 


alpinum 449 
carpathicum 449 
decurrens 454 
dimorphum 453 
germanicum 447 
Helleri 454 
luteo-album 453 
margaritaceum 453 
norvegicum 456 
obtusifolium 454 
palustre 455 
plantaginifolium 451 
polycephalum 454 
purpureum 456 
supinum 455 
sylvaticum 456 
uliginosum 455 
see Antennaria 
3 449-53 
Gomphrena 
globosa 28 
interrupta 7 
Gonolobus an36 
laevis 36 
macrophyllus 37 
suberosus 3 


7 

see Vincetoxicum 37 
Gonopyrum 

americanum 077: 

Goodyera I 569 

seePeramium 569-70 


Gordonia 2 527 
Lasianthus 527 
GRAMINEAE I 107-295 
Graphephorum 1 262 
festucaceum 262 
flexuosum 235-6 
melicoideum 262 
Gratiola 3 104 
acuminata 193 
aurea 195 
dubia 196 
Monniera 192 
officinalis 194 
pilosa 196 
sphaerocarpa 194 
virginiana 194 
viscosa 195 
Grindelia 3 371 
grandiflora 371 
inuloides 371 
lanceolata 371 
nuda 371 
squarrosa 371 
“nuda 371 


GROSSULARIACEAE 2 236 


Grossularia 2 239 
cynosbati 239 
hirtella 241 
Grossularia 239 
missouriensis 240 
oxyacanthoides 240 
reclinata 241 
rotundifolia 241 
setosa 240 

Guilandina 
dioica 2 340 

Gutierrezia 3 369 


Euthamiae 370 


Gutierrezia 
linearis 3 370 
linearifolia 370 
Sarothrae 370 
Gymnadenia 1) i 
conopsea 553 
Gymnadeniopsis 1 552 
clavellata 553 
integra 553 
nivea 553 
Gymnandra 
Bullii 3 198 
rubra 199 
Gymnocladus 2 339 
canadensis 340 
dioicus 340 
Gymnopogon Tees 
ambiguus 225-6 
brevifolius 226 
racemosus 226 
GYMNOSPERMAE~ I 55 
Gymnostichum 
Aystrix I 294 
Gypsophila 2 71-2 
muralis 71 
paniculata 72 
repens 71 
Gyrostachys I 563 
(Spiranthes) 563 
cernua 565 
gracilis 566 
linearis 565 
_ ochroleuca 565 
odorata 565 
parviflora 565 
praecox 566 
Romanzoffiana 564 
simplex 566 
stricta 564 
Gyrotheca Sei 
(Lachnanthes) 
capitata 531 
tinctoria 531 
Habenaria if Be) 
(ine. Orchis in part) 
alba 557 
blephariglottis 1 557 
bracteata 552 
Canbyi 557 
ciliaris 557 
clavellata 553 
cristata 557 
dilatata 554 
fimbriata 558 
flava 552 
grandiflora 558 
holopetala 557 
Hookeriana 556 
hyperborea 554 
integra 553 
lacera 558 
leucophaea 558 
macrophylla 555 
nivea 553 
oblongifolia 556 
obtusata 556 
orbicula'a 555 
peramoena 559 
psycodes 559 
tridentata 553 
unalaskensis 555 
virescens 552 
viridis 552 
HAEMODORACEAE I 530 
Halenia gens 
Brentoniana Bars 
deflexa 15 
heterantha 15 
Halerpestes 2 117 
Cymbalaria 117 
Halesia 2 722 
carolina 722 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Halesia 
tetraptera 2 722 
Halimium 2 539 
canadense 540 
majus 540 
HALORAGIDACEAE 2 612 
HAMAMELIDACEAE 
2 234 
Hamamelis 2 234 
virginiana 235 
“ carolina 234 
Hamiltonia I 641 
oleifera I 641 
Haploesthes Bases 
Greggii 535 
Harperella 2 659 
nodosa 659 
vivipara 659 
Harrimanella . 2 686 
hypnoides 686 
Stelleriana 686 
Hartmannia 2 602 


(Oenothera in part) 


faux-gaura 603 
speciosa 603 
Hedeoma 3 135 
ciliata 136 
Drummondii 136 
glabra 139 
hirta 136 
hispida 136 
longiflora 136 
pulegioides 136 
Hedera , 
quinquefolia 2 511 
Hedyosmos 
origanoides 3 146 
Hedyotis ‘ 
see Houstonia 
3 251-4 
Hedysarum 2 392 
acuminatum 396 
alpinum amer. 392 
americanum 392 
boreale * 392 
canadense 401 
ciliare 402 
coronarium 392 
lineatum 396 
Mackenzii 392 
marylandicum 400-2 
nudiflorum 3905 
pedunculatum 364 
rigidum 401 
virginicum 393 
volubile 420 
see Meibomia 
2 396-402 
“Lespedeza 403-8 
Heleastrum 
paludosum 3 431 
HELEBOREAE 2 84 
HELENIEAE 3°349 
Helenium 3 510 
autumnale 510 
“ pubescens 510 
montanum 510 
nudiflorum 511 
pubescens 510 
tenuifolium init 
Heleochloa I 190 
(Crypsis) 
’ alopecuroides 190 
schoenoides 190 
Helianthemum 2 539 
canadense 540 
corymbosum 541 
georgianum 540 
majus 540 
propinquum 540 
HELIANTHEAE 3 349 
Helianthium I 95 


Vot. IIT. 

Helianthium 
parvulum I 95 
tenellum 95 
Helianthus 3.477 
ambiguus 481 
angustifolius 479 
annuus 477-8 
aridus 478 
atrorubens 479 
Dalyi 482 
decapetalus 484-5 
divaricatus 483 
doronicoides 484 
Dowellianus 480 
giganteus 481 
“ crinitus 479 
subtuberosus 482 
grosse-serratus 483 
hirsutus 3 485 


“ trachyphyllus 
Kellermani 3 483 


illinoiensis 480 
laetiflorus 486 
laevigatus 481 
laevis 495 
lenticularis 478 
macrophyllus 485 
Maximiliani 482 
microcephalus 481 
mollis 3 484-5 
Nuttallii 484 
occidentalis 480 
orgyalis 479 
parviflorus 481 
petiolaris 478 
rigidus 480 
Rydbergi 3 482 
scaberrimus 480 
scrophulariaefolius 

3 484 
strumosus 485 
“macrophyllus 485 


subrhomboideus 480 


subtuberosus 482 
tomentosus 486 
tracheliifolius 485 
tuberosus 486 
“ subcanescens 486 
Heliopsis 3 466 
helianthoides 467 
laevis 467 
scabra | 467 
Heliotropium 3 73 
convolvulaceum 75 
curassavicum 74 
europaeum og 
indicum 75 
spathulatum 74 
tenellum 74 
Helleborus 2 87 
hyemalis 88 
niger 87 
orientalis 87 
trifolius 88 
viridis 87 
Helmintha 
echioides 3 Bua 
Helonias 1 488 
asphodeloides 488 
bullata 488 
graminea 490 
Helopus I 124 
Hemerocallis I 496 
flava 496 
fulva 496 
Hemianthus 
micranthemoides 
: 3 197 
Hemicarpa i369 
aristulata 340 
Drummondii 339 
Isolepis 339 


Vor. III. GENERAL INDEX OF LATIN GENERA AND SPECIES. 579 
Hemicarpa Hicoria Hopea Hydrocotyle 
intermedia I 340 glabra I 583 tinctoria ayo rotundifolia 2 650 
micrantha 339 “ hirsuta 583 | HorDEAE I 108, rr umbellata 649 
occidentalis 340 “ odorata 583 | Hordeum I 286 “ ambigua 649 
subsquarrosa 339 * villosa 584 jubatum 287 verticillata 649 
Hendecandra laciniosa 582 murinum 287 vulgaris 648 
texensis 2 455 microcarpa 583 nodosum 286 | Hydrolea 
Hepatica 2 I01 minima 580 Pammelii 287 affinis 3 71 
acuta 101 ovata 581 pratense 286 caroliniana 72 
acutiloba 101 pallida 582 pusillum 286 see Nama 3 71-2 
Hepatica 101 Pecan 580 vulgare 286 | Hydropeltis 
triloba IOI sulcata 582 | Hosackia 2 359 purpurea 2 76 
Heracleum 2 635 villosa 584 americana 359 | HyDROPHYLLACEAE 
lanatum 635 | Hieracium 3 328 bicolor 359 3 65. 
Spondylium 635 alleghaniense 331 Purshiana 359 | Hydrophyllum 3 65, 
Herpestis 192 aurantiacum 334 | Hottonia 2 709 appendiculatum 66 
amplexicaulis 193 barbatum 330 diffusa ' 709 canadense 66 
Monniera 192 canadense 329 inflata 710 macrophyllum 66 
nigrescens 193 florentinum 332 | Houstonia 3 250 patens 66 
obovata 193 floribundum 333 angustifolia 254 virginicum 65 
rotundifolia 192 Greenii 332 calycosa 252 | Hydrotrida 3 193 
Hesperaster Gronovii 331 ciliolata 25a caroliniana 193 
Sstrictus 2 567 longipilum 330 coerulea 251 | Hymenatherum 
Hesperis 2175 marianum 331 lanceolata R252 aureum 3 514 
pinnatifida 178 molle 329 Linnaei minor 252 | Hymenocallis 1.536 
matronalis 175 murorum 328-9 longifolia 253 littoralis ‘533 
Heteranthera I 463 paniculatum 330 minima 252 occidentalis 533. 
dubia 464 Pilosella 333 minor 252 | Hymenolobus 
graminea 464 “peleterianum 333 patens 252 divaricatus 2 158 
limosa 464 praealtum 332 purpurea 252-3 | Hymenopappus 3 505 
peduncularis 464 pratense 333 “ calycosa 252 anthemoides 510 
reniformis 463 runcinatum 326 “ ciliolata 253 carolinensis 505 
Heterotheca Bs72 scabriusculum 3 330 “ longifolia 253 corymbosus 505 
Lamarckii 372 scabrum Sarr “ pubescens 253 filifolius 3 506 
subaxillaris 372 spathulatum 329 “ tenuifolia 254 flavescens 506 
Heuchera 2 224 umbellatum 330 serpyllifolia 251 scabiosaeus 505 
americana 225-6 venosum 332 tenuifolia 254 tenuifolius 506 
crinita 226 vulgatum 3 329 | Hudsonia 2 541 | HyMENOPHYLLACEAE 
Curtisii 227 | Hierochloé ericoides 541 rs 
hirsuticaulis 227 see Savastana tomentosa 542 | Hymenoxys 3 500 
hispida 227 I 171-3 “ intermedia 542 linearifolia 509 
lancipetala 226 borealis 172| Humulus | I 633 multiflora 510 
longiflora 228 | HipPpOCASTANACEAE Lupulus” 633 odorata 510 
macrorhiza 226 I 498 japonicus 633 | Hyoscyamus 3 168 
parviflora 225 | Hippophoé Hutchinsia 2 158 niger 169 
puberula 225 argentea 20577, petraea 158 | Hyoseris 
pubescens 228 | Hipposelinum 2 634 procumbens 158 caroliniana 306 
Richardsomi 227! Levisticum 635 | Hyacinthus minima 307 
roseola 227 | Hippuris 2 612 botryoides I 510 virginica 306 
Rugelii 225 maritima 613 racemosus 511 | Hypaelytrum 
villosa 226 tetraphylla 613 | Hydactica 2 220 argenteum I 339 
Hexaphoma vulgaris 612 foliosa 220 | HyPERICACEAE 2 527 
petiolaris 2 221 | Hoffmanseggia 2 338 petiolaris 221 | Hypericum 2 528 
Tiexalectris I 576 Falcaria 338 stellaris 221 adpressum 531 
aphyllus 577 Jamesii 338 | Hydrangea 2230, = angulosum 533 
spicata 577 stricta 338 arborescens 230 Ascyron 529 
squamosus 577 | Holcus I 120 “ kanawhana 231 aureum 531 
Hexastylis I 643 alpinus 173 cinerea 231 Bissellii 532 
arifolia 644 halepensis 121 radiata 231 boreale 534 
Memmingeri 644 lanatus 214 vulgaris 231 campanulatum 537 
Ruthii 644 laxus 248 HyDRANGEACEAE 2 230 canadense 534-6 
Shuttleworthii 643 odoratus 172 | Hydrastis 2 84 cistifolium 532 
virginica 644 sorghum 120 canadensis 85 corymbosum 533 
Hibiscus 2 523 striatus I 163 carolinensis 103 densiflorum 530 
incanus 524 | Holosteum 2 50 | HypROCHARITACEAE dolabriforme 532 
lasiocarpus 525 umbellatum 50 I 106 Drummondii 536 
militaris 525 | Homalobus 2 386 | Hydrocharis ellipticum 532 
Moscheutos 524 (Astragalus in part) cordifolia tT 107, galioides 531 
oculiroseus 524 aboriginum 384 Spongia 107 gentianoides 536 
opulifolius 524 caespitosus 386 | Hydrocotyle 2 648 graveolens 534 
palustris 524 montanus 387 ambigua 649 gymnanthum 535 
syriacus 526 multiflorus 386 americana 650 Kalmianum 530 
Trionum 523, 5 Stipitatus 386 asiatica 651 Lasianthus 527 
virginicus 523-5 tenellus 386 australis 649 maculatum 533 
Hicoria 1 579 | Homalocenchrus 1 168 Canbyi 649 majus 535 
alba 582 (Leersia) 168 chinensis 648 mutilum 535 
aquatica 581 lenticularis 169 interrupta 649 perforatum 529, 33 
borealis 583 oryzoides 169 lineata 648 petiolatum 537 
carolinae-septen- virginicus 168 natans 650 prolificum 530 
trionalis 581 | Honkenya 258 ranunculoides 650 pseudomaculatum 534 
cordiformis 580 peploides 58 repanda 651 punctatum 533 


580 


Hypericum 
pyramidatum 2 529 
Sarothra 536 
sphaerocarpum 532 
subpetiolatum 533 
virgatum 533 
“ ovalifolium 533 
virginicum 537 


Hypochaeris 3 309 


glabra 309 
radicata 309 
Hypopitys 2 674 
americana 675 
Hypopitys 675 
lanuginosa 675 
multiflora americana 
675 
Hypoporum 
verticillatum 1 351 
Hypoxis I 534 
erectum 534 
hirsuta 534 
-Hyssopus 3 140 
anisatus Mebee 
nepetoides IIL 
officinalis 140 
see Agastache 111-2 
Hystrix I 294 
(Asprella) 
Hystrix 294 
Ibidium I 563 
Beckii 566 
cernuum 565 
gracile 566 
incurvum 565 
ovale 565 
plantagineum 564 
praecox 566 
strictum 564 
vernale 565 
Tlex 2 486 
Aquifolium 486 
bron¥ensis 489 
Cassine 487 
coriacea 488 
Dahoon 487 
decidua 488 
fastigiata 489 
glabra 487 
laevigata 489 
lucida 488 
mollis 488 
montana 488 
monticola 488 
myrtifolia 487 
opaca 486 
verticillata 489 
“ padifolia 489 
“ tenuifolia 489 
vomitoria 487 
tliamna 2n522 
remota 522 
TLICACEAE 486 
Ilicioides 2 490 
mucronata 490 
Tllecebrum 
Paronychia 2 26 
Ilysanthes 3 195-6 
attenuata 196 
dubia 196 
gratioloides 196 
“ curtipedicellata 196 
riparia 196 
Impatiens 2 440 
aurea 441 
Balsamina 440 
biflora 440 
fulva 440 
noli-tangere 440 
Nortoni - “440 


Impatiens 
pallida 2 441 
Imperatoria 2 638 
Ostruthium 638 
Indigofera 2 371 
leptosepala 372 
tinctoria 371 
Inula 3.457 
divaricata 380 
ericoides 434 
falcata 373 
gossypina Bria 
graminifolia 3773 
Helenium 457 
mariana 374 
subaxillaris 72 
villosa 374 
INULEAE 3 348 
Iodanthus 2 178 
hesperidoides 178 
pinnatifidus 178 
Ionactis 3 444 
linariifolius 444 
Jonidium 
lineare 2 564 
polygalaefolium 564 
Ionoxalis 2 431 
violacea 431 
Toxylon r6g2 
Ipomoea 3 43 
carolina 44 
coccinea 43 
commutata 44 
hederacea 45 
hederaefolia 43 
lacunosa 44 
leptophylla 44 
Nyctelea 67 
pandurata 43 
pes-tigrinis 43 
purpurea 45 
Quamoclit 42 
trichocarpa 44 
Iresine 28 
celesioides 8 
paniculata 8 
TRIDACEAE x 536 
Trio 2 174 
Iris I 536 
aphylla 539 
carolina 539 
caroliniana i7/ 
cristata 540 
cuprea 539 
Duerinckii 539 
foliosa 538 
fulva 539 
georgiana 537 
germanica 536-9 
gracilis 539 
hexagona 537 
Hookeri 538 
lacustris 540 
missouriensis 538 
orientalis 540 
prismatica 539 
Pseudacorus 540 
setosacanadensis 538 
verna 541 
versicolor 537 
virginica 537,9 
Isanthus 3 104 


(Trichostema in part) 


brachiatus 104 
coeruleus 104 
Isnardia 2 585 
(Ludwigia in part) 
palustris 585 
ISOETACEAE r50 
Isoetes TSO 
Braunii 51 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Isoetes 
Butleri I 54 
canadensis 53 
Dodgei 53 
Eatoni 53 
echinospora 51 
“ Boottii 51 
muricata 51 
robusta 51 
Engelmanni 53 
“ fontana 53 
“ gracilis 53 
valida 53 
foveolata 52 
“ pleurospora 52 
Gravesii 54 
Harveyi 5I 
heterospora 50 
hieroglyphica 51 
lacustris 50 
macraspora 50 
melanopoda 54 
riparia 52 
saccharata 52 
Tuckermani 51 
“ borealis il 
Isopappus 379 
divaricatus 380 
Isopyrum 2 88 
biternatum 89 
thalictroides 88 
trifolium 88 
Isotria 560 
affinis 561 
verticillata 561 
Itea 233 
virginica 233 
ITEACEAE 233 
Iva 3 338 
annua 338, 40 
axillaris 339 
ciliata 340 
frutescens 339 
imbricata 339 
oraria 339 
xanthiifolia 340 
Ixia 
acuta 541 
chinensis 542 
Ixophorus I 164-5 
seeChaetochloa 1164 
Jacksonia 
trachysperma 199 
Jasione 299 
montana 299 
Jatropha 
stimulosa 462 
Jeffersonia 129 
Bartonis 129 
binata 129 
diphylla 129 
JUGLANDACEAE 578 
Juglans 578 
alba 582 
cinerea 579 
nigra 579 
regia 578 
tomentosa 582 
see Hicoria 1 580-3 
JUNCACEAE I 465 
Juncoides I 482 
(Luzula) 
arcticum 484 
bulbosum 485 
campestre 484-5 
carolinae 483 
confusa 484 
hyperboreum 484 
nemorosum 483 
nivale 484 


Vor. IIT. 

Juncoides 
parviflorum 1 483 
pilosum 483 
spicatum 484 
Juncus I 465 
acuminatus 481-2 
“robustus 482 
acutus 465 
alpinus 478 
“insignis 478 
aristulatus 473 
articulatus 477 
asper 479 
balticus 468 
biglumis 475 
brachycarpus 479 
brachycephalus 480 
brevicaudatus. 481 
bufonius 469 
bulbosus 477 
caesariensis 479 
campestris 485 
canadensis 480 
“ coarctatus 481 
castaneus 475 
compressus 470 
conglomeratus 467 
debilis 481 
dichotomus 472 
“ platyphyllus 470 
diffusissimus 482 
Dudleyi 470 
effusus 467 
Engelmanni 479 
filiformis 467 
Gerardi 223,470 
Greenei 472 
gymnocarpus 468 
interior 471 
Leersii I 467 
longistylis 474 
maritimus 469 
marginatus 473 
“ biflorus 473 


“ paucicapitatus 473 
“ setosus 474 
megacephalus 480 
militaris 477 
monostichus 470 
nemorosus 483 
nodatus 482 
_ nodosus I 478 
“megacephalus 478 
oronensis 472 
parviflorus 483 
pelocarpus 476 
“ subtilis 476 
pilosus 482-3 
polycephalus 479 
repens 474 
Richardsonianus 478 
robustus 482 
Roemerianus 468 
scirpoides 480 
secundus 471 
setaceus 473 
setosus 474 
Smithii 468 
spicatus 484 
stygius 475 
“ americanus 475 
subtilis 476 
tenuis 470-1 
Torreyi 478 
trifidus 469 
triglumis 476 
validus 479 
Vaseyi 471 
Juniperus I 65 
communis 65,6 
“ alpina 6 


Vor. IIT. GENERAL INDEX OF LATIN GENERA AND SPECIES. S8I 
Juniperus Koellia Lacinaria Laurus 
c. depressa I 66 pilosa 3 142 spicata 3 367 aestivalis 2 135, 
horizontalis 67 pycnanthemoides 144 squarrosa 365 Benzoin 135. 
nana 66 verticillata 143 “intermedia 365 Borbonia 133) 
prostrata 67 virginiana 142 | Lactuca Seay. carolinensis 134 
Sabina procumbens 67 | Koelreuteria 2 501 acuminata 321 geniculata 135 
scopulorum 66 paniculata 501 canadensis 320 melissaefolia 136 
sibirica 66 | Koenigia I 647 “ montana 320 Sassafras 134 
virginiana 66 islandica 647 campestris 320 | Lavauxia 2 603 
Jussiaea 2 589} Koniga 2553 elongata 319-20 (Oenothera in part) 
brachycarpa 587 (Alyssum in part) floridana 321 brachycarpa 604 
decurrens 589 maritima 153 hirsuta 319 triloba 603-4 
diffusa 589 | Korycarpus I 247 integrifolia 320 Watsonii 604 
repens 589 (Diarrhena) ludoviciana 319| Leavenworthia 2 189 
Justicia arundinaceus 247 Morssii 320 aurea 189 
brachiata 3 244 diandrus 247 pulchella 320 Michauxii 189 
chinensis 244 | Kosteletzkya 2 523 sagittifolia 320 torulosa 189 
althaeifolia 523 saligna 318 uniflora 189 
Kallstroemia 2 442 virginica 523 sanguinea 319 | Lechea 2 542 
hirsutissima 443 | KRAMERIACEAE 2 340 sativa 317 intermedia 544 
intermedia 443 | Krameria 2 340 Scariola 318 juniperina 545 
maxima 443 Ixine 340 Serriola 318 Leggettii 544 
Kalmia 2 683-4 lanceolata 340 spicata 321 major 540, 3 
angustifolia 683 secundiflora 340 “ integrifolia 321 maritima 543 
carolina 684 | Kraunhia 2 373 Steelei 319 “interior 543 
glauca 684 (Wisteria) 373 villosa 321 minor 542 
hirsuta 685 frutescens 374 virosa 318 moniliformis 544 
latifolia 683-4 macrostachys 374 | LAMIACEAE 3 99 novae-caesareae 542 
“ myrtifolia 684 | Krigia 3 306 | Lamium Br25 racemosa 543 
polifolia 684 amplexicaulis 307 album 122 stricta 545 
Kalmiella 2 685 caroliniana 308 amplexicaule I21 tenuifolia 544 
hirsuta 685 Dandelion 308 hispidulum 118 thymifolia 542 
Keerlia occidentalis 308 maculatum 122 Torreyi 544 
skirrobasis 3 401 oppositifolia 306 purpureum 121 villosa 543 
Kentrophyta 2 387 virginica 307 | Laportea Lecticula 3 230 
montana 387 | Krynitzkia 3 79 canadensis I 636 resupinata 230 
viridis 387 crassisepala 80 | Lappa 3 547] Ledum 2 676 
Kickxia 3 175 Fendleri 80 major 547 buxifolium 682 
Elatine 176 glomerata 81 minor 548 decumbens 677 
spuria 175 Jamesii 81 tomentosa 547 groenlandicum 677 
Knautia sericea 81 | Lappago I I21 latifolium 677 
arvensis 3 290 | Kuhnia 3 364 see Nazia TU2 7 palustre 677 
Kneiffia 2 599 eupatorioides 364 racemosa ae “ decumbens 677 
Alleni 2 600 “corymbulosa 364 Lappula 3 77 Leersia I 168-9 
fruticosa 602 glutinosa 364 (Echinospermum) see Homalocenchrus 
“ pilosella 602 Hitchcockii 364 americana 79 I 168 
glauca 599, 602 suaveolens 364 deflexa 79 | Legouzia 3 207 
linearis 600-1 | Kuhnistera 2 368 echinata 77 see Specularia 3 207 
linifolia 600 candida 369 floribunda 78 | LEGUMINOSAE 2 330, 41 
longipedicellata 600 compacta 369 Lappula 77 | Leiophyllum 
Sumstinet 602 foliosa 377i Myosotis 77 buxifolium 2 682 
pumila 601 oligophylla 369 Redowskii 78 | LEITNERIACEAE 1 586 
riparia 601 multiflora 370 texana 78 | Leitneria 586 
Kobresia Laser pinnata 368 virginiana 78 floridana 586 
(ine. Elyna) purpurea 379 | Lapsana 3 306] LEMNACEAE I 446 
Bellardi 351 tenuifolia 370 capillaris 327 | Lemna I 447 
bipartita 351 | Kyllinga I 296 communis 306 angolensis 447 
caricina 351 cayennensis 309 | Larix I 60 cyclostasa 447 
elachycarpa 377. maculata 339 americana 60 gibba 448 
scirpina 351 ovularis 30 laricina 60 hyalina 448 
Kochia 2 16 monocephala 2096 Larix 60 minima 447 
americana 16 pumila 296 | Lathyrus 2 412 minor 448 
atriplicifolia 16 decaphyllus 414 “ cyclostasa 447 
Scoparia 16 | LABIATAE 3 99 glaucifolius 415 paucicostata 448 
Koeleria I 245 | Lachnanthes incanus 415 perpusilla 448 
cristata 245 tinctoria List latifolius 416 “ trinervis 448 
gracilis 245 | Lachnocaulon I 455 linearis 409 polyrhiza 446 
nitida 245 anceps 456 maritimus 413 trisulca 447 
pennsylvanica 244 Michauxii 456 myrtifolius 414 valdiviana 447 
Koellia 3 141 | Lacinaria 3 364 “macranthus 414] Lemotrys 
(inc. Pycnanthemum (Liatris) ochroleucus 415 hyacinthina I 509 
and Brachystemum) acidota 366 ornatus 415 | LENTIBULARICEAE 3 225 
albescens 143 cylindracea 365 palustris 414 | Leontice 
aristata 144 elegans 366 “ linearifolius 414 thalictroides 2 128 
capitata 141-2 graminifolia 368 polymorphus 414 | Leontodon 93r5 
clinopodioides 143 Helleri 368 pratensis 415 autumnale 310 
flexuosa 142 kansana 368 pusillus 416 carolinianum 324 
hyssopifolia 143 pilosa 368 sativus 412 erythrospermum 315 
incana 144 punctata 366 tuberosus 416 hastile 310 
leptodon 142 pycnostachya 367 venosus 413 hirtum 310 
montana 145 scariosa 367 | LAURACEAE 2 133 hispidum 310 
mutica 145 Smallii 368 | Laurus 2 133 latilobum 315 


582 


Leontodon 
nudicaule 3 310 
Taraxacum 315 
Leonurus 3 119 
Cardiaca 119-20 
Marrubiastrum 120 
sibiricus 120 
Lepachys 3 473 
columnaris 474 
“pbulcherrima 474 
pinnata 474 
Tagetes 475 
Lepargyraea 257.6 
(Shepherdia) 
argentea 576-7 
canadensis 576 
rotundifolia 576 


Lepidanthus 


phyllanthoides 2 453 
Lepidium 2 164 
apetalum 166 
campestre 164 
densiflorum 166 
didymum 167 
Draba 165 
intermedium 166 
latifolium 164 
medium 166 
neglectum 166 
procumbens 158 
ramosissimum 166 
ruderale . 165 
sativum 166 
virginicum 165 
Leptamnium 236 
(Epiphegus) 
virginianum 236 
Leptandra 3 204 


(Veronica in part) 


virginica 204 
Leptanthus 
gramineus I 464 
ovalis 464 
see Heteranthera 463 
Leptasea 2 216-7 
aizoides 216-7 
Hirculus 216 
tricuspidata 217 
Leptilon 3 442 
(Erigeron in part) 
canadense 442 
divaricatum 442 
Leptocaulis 
divaricatus 2 651 
echinatus 652 
patens 652 
Leptochloa I 229 
attenuata 230 
filiformis 230 
mucronata 230 
Leptodactylon 3 62 
caespitosum 62 
californicum 62 
pungens 62 
Leptoglottis 2, 333 
Leptoloma I 124 
cognatum 124 
Leptopoda - 
brachypoda S058 
Leptorchis 
see Liparis 1 571-2 
Leptoseris 
sonchoides 3 314 
Lepturus I 282 
filiformis 282 
paniculatus 226 
repens 282 
Lespedeza 2 402 
acuticarpa 405 
angustifolia 407 
Bicknellii 407 
Brittonii 404 


Lespedeza 
capitata 2 407 
“ sericea 407 
frutescens 406 
hirta 407 
“ oblongifolia 407 
leptostachya 408 
longifolia 407 
Manniana 404 
neglecta 405 
Nuttallii 405 
prairea 404 
procumbens 403 
repens 403 
reticulata 406 
sessiliflora 402 
simulata 406 
striata 408 
Stuvei 405 
“ intermedia 406 
velutina 407 
violacea 404 
“ angustifolia 406 
virginica 406 
Lesquerella 2 154 
(Vesicaria in part) 
arctica 156 
“ Purshii 156 
argentea 155 
globosa I55 
gracilis 155 
Lescurii 154 
ludoviciana 155 
ovalifolia 156 
spathulata 154 
Leucanthemum 
arcticum 3 518 
vulgare 518 
Leucelene 434 
(Aster in part) 
ericoides 434 
Leucocrinum I 496 
montanum 496 
Leucophysalis 3 162 
grandiflora 163 
Leucopsidium 
arkansanum 3 4otI 
Leucothoé 2 687 
axillaris 687 
Catesbaei 688 
elongata 689 
racemosa 689 
recurva 688 
spinulosa 688 
Levisticum 2 647 
Levisticum 635 
officinale 635 
Liatris 3 304 
see Lacinaria 3 364 
cylindrica 365 
graminifolia 368 
“ dubia 368 
intermedia 366 
odoratissima 369 
paniculata 369 
pumila 367 
spicata 367 
“montana 367 
squarrosa 365 
squarrulosa 367 
LIGULIFLORAE 3 304 
Ligusticum 2 647 
actaeifolium 646 
canadense 647 
Levisticum 635 
Scoticum 647-8 
Ligustrum 2 729 
vulgare 729 
Lilaeopsis 2 648 
(Crantzia) 
lineata 648 
LILIACEAE I 495 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Lilium I 502 
andinum 502 
canadense 503 
candidum 502 
carolinianum 504 
Catesbaei 503 
Grayi 503 
lanceolatum 502 
philadelphicum 502 
superbum 503, 4 
tigrinum 504 
umbellatum 502 

LIMNANTHACEAE 2 441 

Limnanthemum 

- see Nymphoides 
3 18-9 

Limnia 2 39 
perfoliata 39 
sibirica 39 

Limnobium I 107 

Spongia 107 

Limnorchis I 554 
dilatata 554 
fragrans 554 
graminifolia 554 
huronensis 554 
hyperborea 554 
media 554 
major 554 

Limnocharis 
Spongia 14 S07) 

Limodorum I 562 
(Calopogon) 
praecox 566 
tuberosum 562-3 
unifolium Giz 

Limonium 2 718 
(Statice) 
carolinianum 718 

Limosella 3 197 
aquatica 198 
australis 198 
calycina 192 
tenutfolia 198 

LINACEAE 2 435 

Linaria 176 
canadensis 177 
Cymbalaria 175 
Elatina 176 
genistaefolia 177 
Linaria 177 
minor 176 
repens 178 
spuria 175 
striata 178 
supina 177 
vulgaris 177 

Lindera 2 135-6 
Benzoin 135 
melissaefolia 136 

Lindernia 
attenuata 197 

Linnaea 3 275 
americana 276 
borealis 276 

Linosyris 
Howardi 3 376 

Linum 2 435 
arkansanum 439 
Berlandieri 439 
Boottii 439 
catharticum 438 
diffusum 437 
floridanum 438 
grandiflorum 436 
humile 436 
intercursum 438 
Lewisii 436 
medium 438 
perenne 436 
rigidum 439 
simplex 439 


Vou. IIL. 
Linum 
striatum 2 437 
sulcatum 439 
usitatissimum 436 
virgimianum 438 
Liparis I 571 
liliifolia 572 
Loeselii 572 
Lipocarpha I 339 
maculata 339 
Lippia | 3 97 
americana 97 
cuneifolia 98 
lanceolata 98 
nodiflora 98 
Liquidambar 2 235 
asplenifolia I 586 
peregrina 586 
Styraciflua 2 (23% 
Liquivitia 
Jlepidota 2 391 
Liriodendron 2 83 
Tulipifera 83 
Lisianthus 
glaucifolius 38 
Russellianus 8 
Listera I 567 
auriculata 568 
australis 568 
convallarioides 567 
cordata 568 
reniformis 567 
Smallii 567 
Lithospermum 3 87 
angustifolinm 89 
arvense 87 
canescens 89 
carolinianum 89 
carolinense 89 
hirtum 89 
latifolium 88 
liniarifolium 89 
lycopsioides 84 
officinale 7,8 
pilosum 88 
tenellum 74 
Torreyi 88 
virginianum go 
Litsea 
geniculata 2 135 
Littorella 3 250 
lacustris 250 
uniflora 250 
‘| LOASACEAE 2 565 
LOBELIACEAE 3 299 
Lobelia 3 299 
amoena 301 
“ glandulifera 301 
Canbyi 304 
cardinalis 300 
Dortmanna 299, 300 
elongata 301 
glandulosa 302 
gracilis 304 
~ inflata 303 
Kalmii 303 
leptostachys 303 
Nuttallii 304 
paludosa 300 
puberula 302 
spicata 302 
“ hirtella 302 
“ parviflora 302 
syphilitica gor 
“ludoviciana gol 
Lobularia 2 153 
Loeflingia 2 60 
hispanica 60 
texana 60 
LOGANIACEAE 2 729 
Loiseleuria 
procumbens 2 683 


Vor III. GENERAL INDEX OF LATIN GENERA AND SPECIES- . 583 
Lolium 1 281 | Lupinus Lysias Malus 
italicum 282 argenteus 2 348 orbiculata E555 Malus 2 289-90 
multiflorum 282 argophyllus 348 | Lysiella I 556 Soulardi 289 
perenne 281 decumbens 348 obtusata 556 sylvestris 290 
temulentum 282 ornatus glabratus 348 | Lysimachia 2 710 | MALVACEAE 2513 
Lomatium perennis 348 ciliata 713 | Malva 2 514 
orientale 2 631 “ occidentalis 348 hybrida 714 Alcea 516 
villosum 632 plattensis 348 lanceolata 714] ° caroliniana 523 
Lonicera 3 277 pusillus 349 longifolia 714 coccinea 519 
altissima 281 | Luzula Nummularia 712 crispa 515 
canadensis 281 see Juncoides 1 482-5 producta TLE hederacea 521 
Caprifolium 277,8] Lychnis ~ 2 68 punctata 711 involucrata 518 
ciliata 281 affinis 70, 68 quadriflora 714 moschata 515 
coerulea 281 alba 68 quadrifolia 7[ii parviflora 514 
Diervilla 283 alpina 62 radicans 713 rotundifolia 515 
dioica 279 apetala 70 stricta 712 sylvestris 514 
Douglasii 279 chalcedonica 68-9 terrestris 712 triangulata 517 
flava 280 coronaria 70 thyrsiflora 715 verticillata Eee; 
glauca 279 dioica 69 vulgaris 711 | Malvastrum 2 519 
glaucescens 279 diurna 69 see Steironema 712-4 angustum 519 
grata 278 Drummondii 70 | LYTHRACEAE 2 577 coccineum 519 
hirsuta 278 Flos-cuculi 69 | Lythrum 2 580 | Malveopsis 
involucrata 282 Githago 61 alatum 581 hispida 519 
japonica 280 vespertina 68 Hyssopifolia 580 | Mamillaria 
marylandica 2 730] Lycium 3 168 lanceolatum 581 MiSSOUrIENSIS 570 
Morrowi 3 281 afreum 168 lineare 581 Nuttallii 2 570 
oblongifolia 281 Barbarum 168 Parsonsia 582 similis 571 
parviflora 279 halimifolium 168 petiolatum 582 sulcata 570 
sempervirens 280 vulgare 168 Salicaria 580-1 vivipara 571 
“ hirsutula 280 | Lycopersicon 3 167 verticillatum 580 see Coryphantha 
Sullivantii 279 esculentum 168 virgatum 581 2 570 
Symphoricarpos 276 Lycopersicon 168 Vulneraria 581 | Manfreda I 533 
-tatarica 282 | LYCOPODIACEAE I 42 virginica 534 
villosa 281 | Lycopodium I 43 | Machaeranthera 3 435 | Manisuris I 112 
Xylosteum 282 adpressum 45 sessiliflora 435 cylindrica 112 
Lophanthus Sy ihm alpinum 46 tanacetifolia 435 rugosa 112 
see Agastache alopecuroides 45 | Macleya “ Chapmani 112 
anisatus Dik, 2 annotinum 46 cordata 2 140 | MARANTACEAE I 546 
Lophiocarpus 1 96-7 apodum 49 | Maclura Mariana 3555 
Lophiola I 534 carolinianum 47 aurantiaca I 632 mariana 555 
americana 535 chamaecyparissus 48 pomifera 632 | Mariscus I 347 
aurea 535 clavatum 43,7 | Macounastrum 1 647 cylindricus 308 
Lophotocarpus 1 96-7 complanatum 47,8 islandicum 647 echinatus 310 
Sagittaria in part) dendroideum 45 | Macrocalyx 3 67 flavus 309 
calycinus y 97 inundatum 44 (Ellisia) jamaicensis 348 
depauperatus 97 “ Bigeloviit 44 Nyctelea 67 mariscoides 348 
maximus 97 lucidulum 44 | MAGNOLIACEAE 2 80| Marrubium 20010 
spathulatus 98 obscurum 45 | Magnolia 2 81 vulgare I10 
spongiosus 97 porophilum 44 acuminata 82 | Marshallia 3 502 
LorANTHACEAE I 638 pungens 45 auriculata 81 caespitosa 503 
Lorinseria I 24 rupestre 49 Fraseri 81 grandiflora 503 
areolata 25 sabinaefolium 47 glauca 82 latifolia 503 
Lotus 2 358 selaginoides 49 macrophylla 81 obovata 503 
“americanus 359 Selago 43 tripetala 82 Schreberi 502-3 
corniculatus 359 sitchense 46 umbrella 82 trinervia 503 
. Sericeus 359 tristachyum 48 virginiana 82 MARSILEACEAE x36 
Lowellia Lycopsis 3 93 | Mahonia 2 127] Marsilea I 36 
aurea 3 514 arvensis 93 repens 128 mucronata 37 
Ludwigia 2 586 virginica 87 | Maianthemum 1 516 natans 38 
alata 587 | Lycopus 3 146 canadense 517 quadrifolia 37 
alternifolia 586-8 americanus 148 | Mairania 2 693 vestita 37 
“ linearifolia 588 asper 148 alpina 694 | MARTYNIACEAE 3 239 
arcuata 585 communis 147 | MALACEAE 2 286] Martynia 3 239 
cylindrica 587 europaeus 146-8 | Malachodendron annua 239 
decurrens 589 integrifolius 147 ovatum | 2°527 louisiana 239 
glandulosa 587 lucidus 148 | Malacothrix 3 314 proboscidea 239 
hirtella 588 membranaceus 147 californica 314 | Maruta 
linearis 588 rubellus 147 sonchoides 314 Cotula 3 516 
palustris 585 sessilifolius 147 | Malapoenna 134-5 | Matourea 
polycarpa 587 sinuatus 148 geniculata 135 nigrescens 3 193 
rudis 586 virginicus 146 | Malaxis I 570]/ Matricaria 3 520 
sphaerocarpa 586 | Lygodesmia 3 322 monophylla 571 asteroides 404 
Ludwigiantha 2 585 juncea 322 paludosa 571 Chamomilla 521 
(Ludwigia in part) rostrata 322 spicata 570 discoidea 521 
arcuata 2 585 | Lygodium 19 unifolia 571 glastifolia 404 
brevipes 586 palmatum 9} Malus 2 288 grandiflora 520 
Lunaria 2 190 scandens 9 angustifoha 288 inodora 520 
annua 190 |} Lyonia baccata 289 matricarioides 521 
biennis 191 ligustrina 2 6901 coronaria 288 Parthenium 519 
rediviva 190 | Lysias 1 S55 glaucescens 289 | __ suaveolens 521 
Lupinus 2 347 bifolia 555 ioensis 289 | Matteuccia _ I ir 
albus 347 Hookeriana 556 lancifolia 288 Struthiopteris II 


584 “GENERAL INDEX OF LATIN GENERA AND SPECIES. Vor. III. 
MaAyYACACEAE 1 450 | Melanthium Mercurialis 2 460 | Mimosa 
Mayaca I 450 latifolium I 49 annua 460 filiculoides 2633 
Aubleti_ 450 “ longipedicellatum perennis 460 illinoensis 332 
fluviatilis” 450 494 | Merimea Intsia 334 
Michauxii 450 muscaetoxicum 489 texana 2 539 microphylla 334 
MAYDEAE I 107-8 parviflorum 495 | Meriolix 2 607 nilotica 330 
Mazus © racemosum 487 (Oenothera in part) | Mimulus 3 189 
japonicus I 191 virginicum 493 intermedia 608 alatus 190 
Mecardonia 3 193 | MELASTOMACEAE 2 582 serrulata 608 Geyeri IQI 
acuminata 193 | Melica 1 246 | Mertensia 3 82 glabratus IQI 
ovata _ 193 altissima 246 lanceolata 83 guttatus 190 
Meconopsis ciliata 246 linearis 83 Jamesii IQI 
diphylla 2 140 mutica 246 maritima 82 Langsdorffii 190 
Medeola Les22 “ diffusa 246 paniculata 83 moschatus IQI 
virginiana 523 “ glabra 246 pulmonarioides 82 ringens 189, 90 
Medicago 2 350 “ parviflora 247 virginica 83 | Miscanthus I 112 
arabica 352 nitens 246 | Mesadenia 3 536 sinensis 112 
denticulata 351 parviflora 247 (Cacalia in part) Mitchella 38255 
falcata 351 Porteri 247 atriplicifolia 537 repens 255 
hispida 351 Smithii 219 plantaginia 537 | Mitella 2 228 
lupulina 351 | Melilotus 2 352 reniformis 536 diphylla 228-9 
maculata 352 alba 352 rotundifolia 536 intermedia 229 
polymorpha 352 altissima 353 tuberosa 537 nuda 229 
sativa 350-1 indica 353 | Mespilus oppositifolia 229 
virginica 406 officinalis 352-3 Amelanchier 2 297) Mitreola 2 730 
Meehania Gh aity) vulgaris 352-3 apiifolia 320 petiolata 731 
cordata 113 | Melissa Bu137, arbutifolia 290-1 | Modiola 2 522 
Megalodonta 3 499 Acinos 139 calpodendron 302 caroliniana 523 
Beckii 500 Clinopodium 138 canadensis 293 multifida 523 
nudata 500 Nepeta 138 caroliniana 310 | Moehringia ysis f75 
Megapterium 2 605 officinalis 137 cordata 311 lateriflora 57 
(Oenothera in pulegioides 136 cornifolia 000 macrophylla 58 
part) see Clinopodium 138 Cotoneaster 321 muscosa 57 
Fremontii 605 | Melosmon 3 103 cuneiformis 299 | Moenchia 
missouriense 605 laciniatum 103 elliptica 299 erecta 250 
Nuttallianum 605 | Melothria 3 2901 flabellata 310 | Mohrodendron 2 722 
oklahomense 606 pendula 292 Phaenopyrum 321 carolinum 722 
Meibomia 2 394-402] MENISPERMACEAE 2 130 pruinosa 315 | Moldavica 3 114 
(Desmodium) Menispermum 2131 pyracantha 322 Moldavica 115 
arenicola 396 canadense 131 rotundifolia 306 parviflora 114 
bracteosa 399 carolinum 131 | Micrampelis 3 291 | Molinia I 237 
canadensis 401 Lyoni 130 (Echinocystis) coerulea 2371 
canescens 398 | Mentha 3 148 lobata 291 | Mollugo 2 35 
“ hirsuta 398 alopecuroides 151 | Micranthemum 3 197 verticillata 35 
Dillenii 400 aquatica ay micranthemoides 107 | Momordica : 
floridana 400 arvensis 152 Nuttallii 197 echinata 3 292 
glabella 397 borealis 152 orbiculatum 197 | Monarda 3 131 
grandiflora 396 canadensis 152 | Micranthes 25207) aristata 134 
illinoensis 401 “ glabrata 152 caroliniana 219 Bradburiana 133 
laevigata 399 Cardiaca 152 Geum 220 ciliata 135 
longifolia 399 citrata 150 micranthidifolia 219 citriodora 134 
marylandica 402 crispa I51 nivalis 218 clinopodia 132 
Michauxii 397 gentilis 153 pennsylvanica 219 didyma 131 
nudiflora 395 longifolia 150 semipubescens 217 dispersa 134 
obtusa 402 Patrinit 154 texana 218 fistulosa 131-2 
ochroleuca 307 piperita 149 virginiensis 218 “ media 132 
paniculata 309 rotundifolia 150 | Microsteris 359 “rubra 132 
“ Chapmani 399 Sativa 153 gracilis 59 hirsuta 135 
“ bubens 399 spicata _ 149 micrantha 59 media 132 
pauciflora 396 sylvestris 150 | Microstylis I 570 mollis 133 
rhombifolia 400 viridis 149 monophylla 571 pectinata 134 
rigida 4o1 | Mentzelia 2 566 ophioglossoides 571 punctata 133 
rotundifolia 397 albicaulis 566 | Mikania 3 362 scabra 133 
sessilifolia 398 aspera 566 hastata 363 tenuiaristata 134 
stricta 398 aurea 566 scandens 363 | Monardella 
viridiflora 400 decapetala 568 | Milium T0738 montana 3 145 
Melampodium 3 458 laevicaulis 568 amphicarpon 164 | Moneses 2 671 
americanum 459 nuda 567 ciliatum 164 grandiflora 671 
australe 459 oligosperma 566 compressum 125 uniflora 671 
cinereum 459 ornata 568 effusum 173 | Monniera 2 192 
leucanthum 459 | MENYANTHACEAE 3 17 punctatum 124 acuminata 193 
Melampyrum 3 224| Menyanthes 3 17 pungens 174 amplexicaulis 193 
americanum 224 nymphaeoides racemosum 175 caroliniana 193 
arvense 224 2 76; 3 19|.Milla Monniera 192 
latifolium 224 trachysperma 3 19 | Millegrana rotundifolia 192 
lineare 224 trifoliata 18 Radiola 2 440 | MonocoTyLEDONES I 68 
MELANTHACEAE I 485 | Menziesia 2 681-2 coerulea 1 501 | Monolepis 217 
Melanthium I 493 ferruginea 682 | Milleria chenopodioides 17 
canadense 517 glabella 682 angustifolia 3 504 Nuttalliana a7, 
glaucum 491 globularis 682 | MIMOSACEAE 2 330 trifida 17 
hybridum 494 pilosa 682 | Mimosa MoNOPETALAE 2 666 
laetum 489 taxifolia 685 | angustissima 2 331 | MONOTROPACEAE 2 673 


Vor. III. 

Monotropa 2 674 
Hypopitys 675 
lanuginosa 675 
procera 673 
uniflora 674 

Monotropsis 2 673 
(Schweinitzia) 
odorata 674 

Montia 2 38 
Chamissonis 38 
fontana 39 
perfoliata 39 

Morongia 2338 
(Schrankia) 
angustata 334 
microphylla 334 
uncinata 333 

MoRACEAE I 630 

Morus 630 
alba 631 
nigra 631 
bapyrifera 632 
rubra 631 

Muhlenbergia 1 184 
ambigua 187 
brevifolia 188 
capillaris 189 
comata 187 
cuspidata 188 
diffusa 184 
erecta 190 
foliosa 185 
glabrifiora 189 
glomerata © 186 
gracillima 188 
mexicana 185 
palustris 185 
pungens 189 
Tacemosa 186 
Schreberi 184 
“ palustris 185 
simplex 187 
sobolifera 185 
sylvatica 186 
tenuiflora 186 
umbrosa 186 
Willdenovii 186 

Mulgedium 
acuminatum 3 321 
floridanum 321 
leucopaeum Bye 

Munroa Te235 
squarrosa 232 

Muscadinia 
rotundifolia 2 509 

Muscari I 510 
botryoides 510 
racemosum SII 

Muscaria 2 222 
caespitosa 222 

Musenium 
see Musineon 2 643 

Musineon 2 643 
divaricatum 643 
Hookeri 644 
tenuifolium 644 
trachyspermum 643 

Myagrum 2 168 
argenteum aids 
paniculatum 159 
perfoliatum 168 
sativum 157 

Myosotis 3 85 
arvensis 86 
collina 86 
Lappula 77 
laxa 85 
macrosperma 87 
micrantha 86 
palustris 85 
scorpio‘des 85-6 
suffruticosa 81 


Myosotis 
verna 3 87 
virginiana 78 
virginica 87 
versicolor 86 
Myosurus 2 103 
minimus 103 
Shortii 103 
MyYRICACEAE I 584 
Myrica I 584 
asplenifolia 586 
carolinensis 585 
cerifera 585 
Gale 584 
Myriophyllum 2 614 
alterniflorum 615 
ambiguum 615 
Farwellii 616 
heterophyllum 616 
humile 615 
pectinatum 614 
pinnatum 616 
proserpinacoides 616 
scabratum 616 
spicatum 614 
tenellum 615 
verticillatum 614 
Myriopteris 
gracilis iy yz 
Myrrhis 
Claytoni 2 627 
longistylis 628 
Myzorrhiza’ 
ludoviciana 2235 
Nabalus ey ciyl 


(Prenanthes in part) 


albus 335 
altissimus 335 
asper 337 
Boottii 337 
crepidineus 338 
cylindricus 336 
integrifolius Bas 
nanus 336 
racemosus 337 
“ binnatifidus 337 
serpentarius 335 
trifoliolatus 334, 6 
virgatus 33 
NATIADACEAE I 80 
Naias I 89 
flexilis 90 
gracilis 89 
gracillima 90 
guadalupensis go 
indica feke) 
major 89 
marina 89 
recurvata 89 
robusta 90 
Nama A070 
(Hydrolea) 
affinis 71 
ovata 2 
quadrivalvis 72 
zeylanica 71 
Napaea 2 518 
dioica 518 
hermaphrodita 520 
Narcissus T 532 
poeticus 532 
pseudo-narcissus 532 
Nardosmia 
palmata 3637 
sagittata 532 
Nardus I 281 
stricta 281 
Narthecium 
americanum 1 487 
glutinosum 486 
ossifragum 487 


Narthecium 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


pubens I 487 | 
Nasmythia 

articulata I 454 
Nasturtium 2 159 


see Radicula 2 159-62 


lacustre 2 164 
natans 164 
officinale 162 
terrestre 161 
Naumbergia 2714 
(Lysimachia in part) 
guttata 715 
thyrsiflora 715 
Navarretia 3 64 
(Gilia in part) 
involucrata 64 
minima 64 
Nazia P02 
(Tragus) 
racemosa 121 
Neckeria 2 143 
Ne-eragrostis 
hypnoides 243 
W eigeltiana 243 
Negundo 
aceroides 2 408 
Negundo 498 
Neillia 
opulifolia 2 244 
Nelumbium 
luieum 2 7r7, 
speciosum 77 
NELUMBONACEAE 2 76 
Nelumbo 2 76 
jamaicensis 77 
lutea 77 
Nelumbo 77 
nucifera 77 
Nemastylis I 541 
acuta 541 
coelestina 541 
gemmiflora 541 
Nemopanthus 2 490 
canadensis 490 
fascicularis 490 
mucronata 490 
Nemophila 3 67 
microcalyx 67 
phacelioides 67 
Neobeckia 2 163 
aquatica 164 
Neopieris 2 690 
mariana 601 
nitida 690 
Neottia I 564, 6 
pubescens 570 
see Ibidium 
NEOTTIEAE I 547 
Nepeta 2113 
Cataria 113 
Glechoma 114 
. hederacea 114 
Nephrodium 
acrostichoides 1 16 
lanosum 34 
punctilobulum 14 
Nesaea 
verticillata 2 580 
Neslia 2 158 
paniculata 159 
Nestronia I 641 
(Darbya) 
umbellula 641 
Nicandra 3 155 
Nicotiana 3 170 
axillaris 171 
longiflora 171 
rustica 170 
tabacum 170 
Niobe I 496 
coerulea 496 


Niobe 
japonica I 496 
Norta 2 174 
altissima 174 
Irio 174 
Notaphoebe 
Borbonia 133 
Nothocalais 
cuspidata 3 324 
Nothoholcus I 214 
lanatus 214 
Notholaena I 35 
dealbata 35 
nivea 35 
Nothoscordum 1 sor 
bivalve 501 
pulchellum 501 
striatum 501 
Nuphar 278 
see Nymphaea 2 78-9 
Nuttallia 2 516 
digitata 516 
involucrata 518 
Nuttallia 2 566 
decapetala 568 
nuda 5, 
stricta. j; 567 
NYCTAGINACEAE 2 30 
Nyctelea 3 67 
Nyctelea 67 
NYMPHAEACEAE om rhe, 
Nymphaea 2 78 
advena 7 
fraterna 78 
Kalmiana 78 
lutea Kalmiana 78 
microphylla 7 
Nelumbo 77 
odorata 79 
“ rosea 79 
minor 79 
rubrodisca 78 
sagittaefolia 79 
tetragona 80 
tuberosa 80 
Nymphoides 3 18 
(Limnanthemum) 
aquaticum 19 
flava 18 
lacunosum 18 
nymphaeoides 19 
Nyssa 2 665 
aquatica 666 
biflora 666 
multiflora 665 
sylvatica 665, 6 
uniflora 666 
Oakesia 2 479 
puberula I 519 
sesstlifolia 519 
Oakesiella I 518 
Obolaria 3.15 
caroliniana 193 
virginica 16 
Oceanorus I 492 
leimanthoides 493 
Ocimum 
frutescens a 8 
Odontites 3 219 
(Bartsia in part) 
Odontites 219 
Odostemon 2 127 
aquifolium 128 
Oenanthe 
ambigua 2 638 
filiformis 638 
teretifolia 638 
Oenoplea 2 501 
Oenothera 2 504 
albicaulis 598-90 
argillicola 596 


586 


Oenothera 
biennis 2 594-5 
brachycarpa 604 
caespitosa 603 
canescens 605 
canovirens 596 
chrysantha 601 
coronopifolia 598 
cruciata 594 
Fremontii 606 
fruticosa 602 
“ humifusa 600 
glauca 602 
grandiflora 595 
guttulata 605 
humifusa 597, 600 
laciniata 597 
Lamarckiana 595 
latifolia 599 
linifolia 600 
linearis 601 
lavendulaefolia 607 
longipedicellaita 600 
macrocarpa 605 
minima 597 
MisSOUrIENSiS 605 
muricata 595 
Nuttallit 599 
Oakesiana 596 
pilosella 602 
pallida 599 
pinnatifida 598 
pratensis 601 
pumila 601 
rhombipetala 597 
serrulata 608 
scapigera 603 
sinuata 597 
speciosa 603 
strigosa 596 
triloba 604 
“ parviflora 604 
see Anogra 598 
Kneiffia 59 
Oldenlandia 3 254 
angustifolia 254 
coerulea 251 
corymbosa 254 
glomerata 254 
minima 252 
purpurea 253 
uniflora Y 254 
see Houstonia 250-4 
OLEACEAE 2I72 
Oligoneuron 
canescens 3 397 
ONAGRACEAE 2 584 
Onagra 2 594-6 
see Oenothera 
biennis 594 
cruciata 594 
Oakesiana 596 
Onoclea TKO 
sensibilis II 
Struthiopteris II 
Ononis 
repens 2355 
Onopordon 3 555 
Acanthium 556 
Onosmodium 3 90 
carolinianum gI 
hispidum coe) 
hispidissimum gI 
molle gt 
occidentale gI 
subsetosum 90 
sylvestre gt 
virginianum go 
Onychium 
densum iss 
Oonopsis 39877 


(Bigelovia in part) 


Oonopsis 
Engelmanni 3 377 
multicaulis 377 
OPHIOGLOSSACEAE I I 
Ophioglossum ror 
arenarium 2 
Engelmanni 2 
vulgatum 2 
Ophiorhiza 
Mitreola 2 731 
Ophrys I 567 
auriculata 568 
australis 568 
cernua 565 
convallarioides 567 
Corallorhiza 574 
cordata 568 
liliifolia 572 
Loeselii 572 
monophyllos 571 
ovata 567 
Smallii 567 
spiralis 563 
Oplopanax 
horridum 2 619 
Opulaster 2 243 
(Physocarpa) 
(Neillia) 
australis 244 
intermedius 244 
opulifolius 244 
Opuntia 2 571 
arborescens 573 
camanchica 572 
fragilis 573 
humifusa 572 
macrorhiza 572 
mesacantha 572 
missouriensis 573 
Opuntia 571 
polyacantha 573 
Rafinesquii 572 
tortispina 572 
vulgaris 571 
ORCHIDACEAE I 547 
ORCHIDEAE I 547 
Orchis I 550 
blephariglottis 557 
bracteata 552 
ciliaris Bey, 
clavellata 553 
cristata 557 
dilatata 554 
fimbriata 558-9 
fissa 558 
flava 552 
grandiflora 558 
hyperborea 554 
huronensis 554 
integra 553 
lacera 558 
leucophaea 558 
militaris 551 
nivea 553 
obtusata 556 
orbiculata 555 
psycodes 559 
rotundifolia 551 
spectabilis 551 
tridentata 553 
Virescens 552 
Oreocarya 3 80 
(Eritrichium and 
Krinitzkia in part) 
fulvocanescens 82 
glomerata 81 
sericea 81 
suffruticosa 80-1 
Origanum 3 140 
flexuosum 142 
vulgare 140 
Ornithogalum 1 509 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Ornithogalum 
arabicum I 509 
bivalve 501 
hirsutum 534 
nutans 510 
umbellatum 509 

OROBANCHACEAE 3 233 

Orobanche 234 
americana 235 
fasciculata 224 
ludoviciana 235 
major 234 
minor 235 
purpurea 234 
ramosa 234 
uniflora 233 
virgimiana 236 

Orontium I 445 
aquaticum 445 

Orophaca 2 387 
(Astragalus in part) 
argophylla 388 
caespitosa 387 
sericea 388 

Orthocarpus 3 216 
luteus 216 

ORYZEAE I 108 

Oryzopsis I 174 
asperifolia 174-5 
canadensis 174 
cuspidata 174 
juncea 174 
melanocarpa 175 
membranacea 174 
micrantha 175 
pungens 174 
racemosa 175 

Osmorrhiza 2 627-8 
brevistylis 627 
Claytoni 62 
divaricata 628 
longistylis 628 
obtusa 628 
see Washingtonia 

627-8 

OSMUNDACEAE rae 

Osmunda 17 
cinnamomea 7 
Claytoniana 8 
interrupta 8 
lanceolata 6 
Lunaria 3 
matricariae 5 
regalis 7 
Struthiopteris II 
virginiana 6 

Ostrya I 606 
Ostrya 606 
virginiana 606 
virginica 606 

Othake 3 507 
callosum 507 
sphacelatum 507 
tenuifolium 507 

Otophylla 3 213 
auriculata 213 
densiflora 214 

OXALIDACEAE 2 430 

Oxalis 2 430 
Acetosella 431 
Brittoniae 434 
Bushii 433 
corniculata 432 
cymosa 435 
filipes 433 
grandis 435 
Priceae 434 
recurva 435 
repens 432 
rufa 433 
stricta 432 
violacea 431 


Voz. IIT. 
Oxybaphus 253% 
see Allionia 2 31 
albidus I 
angustifolius 31 
hirsutus 32 
Oxycoccus 2 704 
erythrocarpus 705 
macrocarpus 705 
Oxycoccus 704 
palustris 704 
Oxydendrum 2 691 
arboreum 692 
Oxygraphis 2 157 
Cymbalaria 117 
“ alpina 117 
Oxypolis 2 637 
(Tiedemannia) 
filiformis 638 
longifolius 638 
rigidus 638 
Oxyria I 659 
digyna 659 
reniformis 659 
Oxytropis 2 388 
arctica 3890 
“ inflata 389 
Belli 3901 
campestris 390 
“ coerulea 390 
inflata 389 
“ johanensis 390 
Lamberti 390 
montana 388 
multiceps 389 
podocarpa 389 
sericea 390 
splendens 390 
Pachiloma 2 633 
Pachylophus 603 
(Oenothera in part) 
caespitosa 603 
Nuttallii 603 
Pachypodium 
integrifolium 2 169 
Pachysandra 480 
procumbens 480 
Pachystima 2 492 
Canbyi 492 
Myrsinites 492 
Padus 2 328 
demissa 329 
melanocarpa 329 
nana 329 
virginiana 329 
Paepalanthus 
flavidulus I 455 
Palafoxia 
Hookeriana 3 507 
Panax 2 618 
(Aralia in part) 
horridum 619 
quinquefolium 618 
trifolium 619 
Pancratium 
carolinianum 1 533 
occidentale 533 
PANICEAE I 108 
Panicularia I 262 
(Glyceria) 
acutiflora 266 
americana 265 
angustata 267 
borealis 266 
brachyphylla 266 
canadensis 263 
distans 268 
“ airoides 268 
elongata 264 
fluitans 266 
grandis 265 
laxa 263 


Voz. IIT. GENEKAL INDEX OF LATIN GENERA AND SPECIES. 587 
Panicularia Panicum Panicum Paspalum 
nervata I 264 hispidum 1134 virgatum I 141 dilatatum L432 
obtusa 264 huachucae 153 “ breviramosum 141 dissectum 127 
pallida 265 © silvicola 153 “ cubense 141 distichum 133 
septentrionalis 265 imberbe 165 * obtusum 141 Elliottii 125 
Torreyana 264 implicatum 154 viride 165 floridanum 132 
Panicum pamnesyl involutum 143 viscidum 159 “ glabratum 132 
aciculare 146 Ischaemum E28 Walteri 134, 162 fluitans 126 
aculeatum 160 italicum 166 Werneri 144 laeve 130 
Addisonii 156 Joorii 161 Wilcoxianum 159 “ pilosum 130 
agrostoides 142 languidum 153 Wrightianum 152 laeviglume 131 
albemarlense 154 lanuginosum 155 xalapense 146 longipedunculatum 
albomarginatum 148 latifolium 162 xanthophysum ~_—158 I 127 
amaroides 141 laxiflorum 146 yadkinense 149 membranaceum 127 
- amarulum 141 leucothrix I 152 | PAPAVERACEAE 2 136 Michauxianum 133 
amarum I4I Liebergii 157 | Papaver 2 136 mucronatum 126 
“ minor I4I Lindheimeri 152 alpinum 138 Muhlenbergii 129 
anceps 142 lineare 123 Argemone 138 ovatum 122 
angustifolium 146 linearifolium 144 dubium 137 paniculatum 126 
annulum 150 longifolium 143 nudicaule 138 baspaloides 125 
Ashei I6I longipedunculatum radicatum 138 platycaule 125 
atlanticum 153 145 Rhoeas 137 plenipilum 130 
auburne 155 lucidum 149 somniferum 136-7 praelongum 130 
autumnale 124 macrocarpon 162] PAPILIONACEAE 2 341 prostratum 128 
barbipulvinatum 139 malacophyllum 160] Papyrius I 632 psammophilum 128 
barbulatum 149-50 mattamuskeetense papyrifera 632 pubescens 129 
Bicknellii 145 151 | Pardanthus I 541 pubifiorum 131 
boreale 150 meridionale 154 chinensis 542 “ glabrum 131 
Boscii 162 microcarpon 148 | Parietaria I 637 purpurascens 131 
Brittoni 148 microcarpon 150 officinalis 637 remotum glabrum 131 
Bushii 147 miliaceum 134, 40 pennsylvanica 638 sanguinale 123 
calliphyllum 158 minus 140 | PARNASSIACEAE 2 211 setaceum 129 
capillare 139 mutabile 161 | Parnassia 2 212 stramineum 127 
“ campestre 139 Nashianum S77, asarifolia 214 supinum 130 
“ flexile 140 nervosum 161 caroliniana 212 tristachyon 125 
“ minor 140 neuranthum 146 grandifolia 212 virgatum 131 
* sylvaticum 140 nitidum 150-2 Kotzebuei 213 Walterianum 127 
carinatum 137 nudum 124 palustris 212-3 | Paspalus 
ciliatum 145 obtusum 138 parviflora 213 Boscianus T 3 
clandestinum 162 octonodum 151 | Paronychia 2 26 furcatus 12 
Clutei Gtk oligosanthes 158 argyrocoma 27 macrospermus 132 
coerulescens 149 oricola 154 depressa 28 | PASSIFLORACEAE 2 564 
cognatum 124 Owenae’ 155 diffusa 28 | Passiflora 565 
colonum 134 patulum 157 dichotoma 28 incarnata 565 
columbianum 156 pauciflorum 158 Jamesii 27-8 lutea 565 
“ thinium 156 paucipilum 151 sessiliflora 28 | Pastinaca 2 634 
Commonsianum 156 pedunculatum 162 Wardi 27 sativa 634 
commutatum 161 perlongum I 144] Parosela 2 366-8 | Paulownia 3 189 
condensum 142 philadelphicum 140 (Dalea) imperialis 189 
consanguineum 146-7 polyanthes 148 aurea 368 tomentosa 189 
Crus-galli 133-4 Porterianum 162 Dalea 367 | Pectis 514 
eryptanthum 160 praecocius 154 enneandra 367 angustifolia 514 
Curtisii 137 proliferum 138 Jamesii 368 ciliaris 514 
Dactylon I 222 psammophilum 156 lanata 367 | Pedicularis 3 219 
debile 138 pseudo-pubescens 153 nana 368 auriculata 221 
decoloratum 162 pubescens 153,9 obovatifolia 366 canadensis 221 
depauperatum 143 pubifolium 162 | Parsonsia 2 582 capitata 222 
dichotomiflorum 138 Ravenelii 159 (Cuphea) euphrasioides 220 
dichotomum 149 roanokense 149 petiolata 582 flammea 222 
“ elatum 149 rostratum 142 | Parthenium 3 464 Furbishiae 222 
diffusum 140 sanguinale 123 auriculatum 465 groenlandica 223 
digitarioides 137 scabriusculum 160 hispidum 466 lanceolata 221 
dissectum 127 scoparioides 153 Hysterophorus: 464-5 lapponica 220 
divergens 124 scoparium 159 integrifolium 465 palustris 221 
Eatoni 152 scoperinum 158, 9 repens | 466 parviflora 221 
elongatum 143 “ Liebergi 157 | Parthenocissus 2 510-1 pedicellata 220 
ensifolium 148 Scribnerianum 158 quinquefolia 511 sylvatica 220 
Enslini 144 serotinum 122 tricuspidata 511 Wlassoviana 221 
filiculme 154 sphaerocarpon 147 vitacea 511 | Pediocactus 2 560 
filiforme 122 spretum 152 | Paspalum L125 Simpsoni 570 
fimbriatum 123 stipitatum 143 angustifolium 130 | Pellaea r.93 
flexile 140 striatum 163 arundinaceum 132 atropurpurea 33 
Gattingeri 139 strictum 143 australe 130 densa 33 
geniculatum 138 strigosum 145 Boscianum rar glabella 33 
gibbum 163 subvillosum 154 Bushii 128 gracilis 32 
glabrum 123 tennesseense 155 ciliatifolium 128 | Peltandra I 443 
glaucum 165 tenue 1 148 “ brevifolium 127 alba 443 
gravius 149 tsugetorum 157 circulare 131 sagittaefolia 444 
Helleri 158 verrucosum 138 compressum 125 undulata 444 
hemitomon 137 verticillatum 164 dasyphyllum 130 virginica 444 
hians 163 villosum 147 debile 130 | PENTHORACEAE 2 2II 
hirtellum 134 villosissimum 153 difforme 132 | Penthorum 2 211 


588 


Penthorum 
sedoides 2 211 
Pentstemon 3 182 
acuminatus 186 
albidus 184 
angustifolius 187 
calycosus 185 
canescens 183 
Cobaea 186 
coeruleus 187 
cristatus 183 
Digitalis 184 
erianthera 183 
glaber 187 
gracilis 185 
grandiflorus 186 
Haydeni 187 
hirsutus 182 
laevigatus 183-4 
pallidus 183 
Pentstemon 184 
pubescens 182 
tubiflorus 185 
Peplis 
americana 2 538 
diandra 579 
Pepo 3 2901 
foetidissima 291 
Peramium I 569 
(Goodyera) 
decipiens 570 
Menziesii 570 
ophioides 569 
pubescens 570 
repens 569 
tessellatum 569 
Perilla 3153 
frutescens 154 
“ nankinensis 154 
ocimoides 154 
“ crispa 154 
Perivloca 
graeca 3 39 
Peritoma 
serrulata 2 197 
Persea 2 133 
Borbonia 133 
carolinensis 133 
“ palustris 133 
gratissima 133 
Persea 133 
pubescens 133 
Persicaria I 665 
amphibia 666 
Careyi 668 
Hydropiper 670 
hydropiperoides 669 
lapathifolia 666 
longistyla 667 
Muhlenbergii 666 
omissa 667 
opelousana 669 
orientalis 670 
pennsylvanica 667 
Persicaria 668 
persicarioides 668 
portoricensis 667 
punctata 670 
robustior 670 
setacea 669 
tomentosa 667 
Perularia I 551 
flava 552 
Petalostemum 2 368 
compactum 369 
candidum 368, 9 
foliosum 371 
gracile 369 
macrostachuus 369 
multiflorum 370 
oligophyllum 369 
purpureum 370 


Petalostemum 
tenuifolium 2 370 
villosum 371 
violaceum 370 

Petasites By pe 
(Nardosmia) 
frigida Bu532 
officinalis 532 
palmata Rie 
Petasites 532 
sagittata 532 
trigonophylla 531 
vulgaris 532 

Petrorhagia 272 
saxifraga 2 

Petroselinum 2 642 
hortense 642 
sativum 642 
Petroselinum 642 

Petunia Ber. 
axillaris 171 
nyctaginiflora 171 
parviflora 172 
violacea 172 


Peucedanum 


daucifolium 2 632 
graveolens 633 
Kingti 633 
sativum 634 
verticillatum 657 
Phaca 2 385-6 
aborigina 384 
americana 385 
argophylla 388 
astragalina 382 
baetica 385 
bisulcata 380 
Bodini 386 
caespitosa 387 
elatiocarpa 383 
elegans 384 
elongata 385 
flexuosus 385 
frigida ameri- 
cana 385 
longifolia 386 
latiflora 383 
neglecta 385 
parviflora 384 
pectinata 380 
Robbinsii 381 
sericea 388 
villosa 379 
Phacelia 68 
bipinnatifida 69 
Covillei ‘70 
dubia 69 
fimbriata 71 
Franklinii 69 
heterophylla 68 
hirsuta 70 
integrifolia 68 
leucophylla 68 
parviflora 69 
Purshii 70 
secunda 68 
Phaethusa 3 487 
americana 487 
helianthoides 488 
occidentalis 488 
virginica 487 
Phaiosperma 2 633 
PHALARIDEAE I 108 
Phalaris I 169 
americana 170 
arundinacea 169, 70 
“ picta 170 
canariensis 170 
caroliniana 170 
erucaeformis 228 
intermedia 170 
orysoides 168-9 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Pharbitis 
hederacea 3°45 
purpurea 45 
Pharnaceum 
maritiumum 2 34 
PHASEOLACEAE Zsa 
Phaseolus 2 422 
angulosus 423 
diversifolius 423 
helvolus 423-4 
leiospermus 424 
perennis 423 
polystachyus 423 
umbellatus 424 
vulgaris 422 
see Strophostyles 
423-4 
Phegopteris 
calcarea I 24 
Dryopteris ze 
hexagonoptera 23 
Phegopteris 23 
polypodioides 23 
Robertiana 34 
Phelipaea 
lutea 3 234 
Phellopteris 2 644 
montanus 645 
Philadelphus Daa 
coronarius 232 
grandiflorus 232 
inodorus 232 
Philozera 3 510 
multiflora 510 
Philotria I 104 
angustifolia 105 
canadensis 105 
minor 106 
Nuttallii 105 
Phippsia I 193 
algida 193 
Phileum I 190 
alpinum IQI 
pratense I9O0-I 
schoenoides 190 
Phlomis 3 118 
fruticosa 118 
tuberosa 118 
Phlox 353 
amoena 55 
amplifolia 54 
argillacea 55 
bifida 56 
Brittonii 58 
bryoides 58 
carolina 54 
divaricata 56 
Douglasii 59 
“ andicola 59 
“ longifolia 59 
glaberrima 53,5 
Hoodii 58 
Kelseyi 57 
longifolia 59 
maculata 54 
“ candida 54 
ovata 54 
paniculata 53 
pilosa 55 
reptans 56 
Stellaria 57 
stolonifera 56 
suaveolens 54 
subulata 57 
Phoradendron 1 639 
californicum 639 
flavescens 639 
Phragmites I 232 
COMMUNIS 232 
Phragmites 232 
PHRYMACEAE 3 244 
Phryma 244 


Vor. III. 


Phryma 
Leptostachya 3 245 
Phyla 3 91 
Phyllanthus 2 453 
carolinensis 453 
Niruri 453 
obovatus 453 
Phyllitis rere 
Scolopendrium 25 
Phyllodoce 2 685 
coerulea 685 
Phymosia 2 521 
abutiloides 521 
cuspidata 522 
remota 522 
Physalis 27555 
aequata 158 
Alkekengi 156, 162 
angulata 158 
barbadensis 157 
“ obscura 157 
comata 161 
grandiflora 163 
hederaefolia 161 
heterophylla 161 
ixocarpa 158 
Lagascae 157 
lanceifolia 158 
lanceolata 160 
“ laevigata 159 
“ hirta 160 
lobata 162 
longifolia 159 
macrophysa 159 
missouriensis 157 
obscura 157 
pendula 158 
pennsylvanica 160-2 
philadelphica 159 
peruviana 161 
pruinosa 157 
pubescens 156 
pumila 160 
rotundata 161 
subglabrata 159 
virginiana 160-1 
“ intermedia 160 
viscosa 161-2 
Physalodes g0rgs 
(Nicandra) 
peruvianum 155 
physalodes 155 
Physaria 2 156 
brassicoides 156 
didymocarpa 156 
Physocarpus 2 244 
intermedius 244 
MissOUriensis 244 
Physostegia 3 116 
denticulata TE7. 
digitalis 116 
intermedia 117 
parviflora 117. 
virginiana 116 
PHYTOLACCACEAE 2 25 
Phytolacca 2 25 
americana 26 
decandra 26 
Picea I 60 
(Abies in part) 
alba 61 
australis 61 
brevifolia 61 
canadensis 61 
mariana 61 
nigra 61 
rubens 61 
rubra 61 
Picradenia 3 510 
acaulis 509 
odorata 510 
Picradeniopsis 507 


Vor. III. 


Picradeniopsis 
oppositifolia 3 508 
Picris 3 311 
asplenioides Roar 
echioides 311 
hieracioides 311 
Picrococcus 2 697 
Pieris 2 689 
(Andromeda in part) 
floribunda 690 
formosa 690 
mariana 691 
nitida 690 
Pilea 636 
muscosa 636 
pumila 637 
Pilosella 175 
novae-angliae 176 
spathulata 332 
Pimpinella 654 
integerrima 639 
magna 655 
saxifraga 655 
PINACEAE Tass 
Pinguicula 3.225 
acutifolia 225 
alpina 226 
vulgaris 225-6 
Pinus I 56 
Abies 60 
alba 61 
australis 35) 
balsamea 63 
Banksiana 58 
canadensis 62 
divaricata 58 
echinata 58 
Fraseri 63 
inops 58 
laricina 60 
mitis 58 
nigra 61 
palustris 57 
pendula 60 
Picea 63 
ponderosa 57 
pungens 59 
resinosa 57 
rigida 59 
rubra 61 
scopulorum 57 
serotina 60 
Strobus 56 
sylvestris 56, 60 
“ divaricata 58 
Taeda 59 
virginiana 58 
Piperia 554 
unalaskensis 555 
Pisum 
maritimum 413 
Planera 628 
aquatica 628 
Planodes 
virginicum 179 
PLANTAGINACEAE 245 
Plantago 245 
arenaria 249 
aristata 248 
asiatica 245 
borealis 247 
cordata 247 
decipiens 247 
elongata 249 
eriopoda 247 
glabra 247 
gnaphalioides 248 
halophila 246 
heterophylla 249 
lanceolata 246 
major 245 
maritima 247 


Plantago 
media 3 246 
patagonica 
“ aristata 248 
“ gnaphalioides 248 
Purshii 248 
pusilla 249 
rhodosperma 248 
Rugelii 246 
sparsiflora 247 
spinulosa 248 
uniflora 250 
virginica 248 
PLATANACEAE 2 242 
Platanus 2 242 
occidentalis 242 
orientalis 242 
Platanthera 
holopetala I 557 
rotundifolia 551 
see Habenaria 552-9 
Plectocephalus 
americanus Cy ists) 
Pleiotaenia 2 633 
Nuttallii 633 
Pleurogyna By 1A 
(Swertia) ' 
rotata I4 
Pleurophragma 
integrifolia 2 169 
Pleuropogon I 247 
Sabinii 247-8 
Pleuropterus I 675 
cordatus 675 
Zuccarinii 676 
Pluchea 3 447 
bifrons 447 
camphorata 448 
foetida 447 
petiolata 448 
PLUMBAGINACEAE 2 717 
Pneumaria 3 82 
(Mertensia in part) 
maritima 82 
Poa I 252 
abbreviata 253 
airoides 268 
alpina 254 
alsodes 257 
andina 260 
angustata 267 
annua 253 
aquatica 262 
“ americana 265 
arachnifera 259 
arida 260 
autumnalis 257 
brachyphylla 258 
brevifolia 258 
Buckleyana 260 
caesia 254 
“ strictior 256 
capillaris 239 
capitata 243 
caroliniana 240 
cenisia 258 
Chapmaniana 253 
compressa I 259 
confusa 261 
cristata 253 
crocata 256 
debilis 255 
distans 267-8 
elongata 264 
eminens I 259 
Eragrostis 240 
fasciculata 258 
flava 233 
flexuosa 257 
glauca 254 
glumaris 259 
hirsuta 239 


Poa 
hypnoides 
interrupta 
Kingii 
laevigata 
laevis 
laxa 
maritima 
memoralis 
nervata 
nitida 
obtusa 
pectinacea 
pilosa 
pratensis 


“ angustifolia 


* pratericola 


pseudopratensis 1 


refracta 
seslerioides 
serotina 
sylvestris 
tenuifolia 
tenuis 
-Torreyana 
trichodes 
triflora 
trivialis 
uniflora 
Wolfii 


Weigeltiana 


Podalyria 
australis 
bracteata 
mollis 

Podophyllum 
diphyllum 
peltatum 


PoDOSTEMACEAE 


Podostemum 


ceratophyllum 


Pogonia 
affinis 


divaricata 


ophioglossoides 


pendula 


trianthophora 


verticillata 
Poinsettia 
dentata 


cuphosperma 


pulcherrima 


heterophylla 


Polanisia 
graveolens 


trachysperma 
POLEMONIACEAE 


Polemonium 
coeruleum 
dubium 
Nyctelea 
reptans 


van Bruntiae 


Polycodium 


melanocarpum 


stamineum 
POLYGALACEAE 
Polygala 
acutifolia 
alba 
ambigua 
brevifolia 
corymbosa 
cruciata 
Curtissii 
cymosa 
fastigiata 
incarnata 
lutea 
mariana 
Nuttallii 
paucifolia 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


589 


Polygala 
polygama 2 451 
“ abortiva 451 
ramosa 447 
sanguinea 449, 50 
Senega 451 
“ latifolia 451 
“ tenuifolia 451 
Torrey 448 
uniflora 452 
verticillata 488 
“ ambigua 449 
viridescens 449 
vulgaris 446 
POLYGONACEAE I 646 
Polygonatum To o20 
biflorum 521 
commutatum 521 
giganteum 521 
Polygonella I 676 
americana 677 
articulata 676 
ericoides 677 
parvifolia 676 
Polygonum I 659 
acre 670 
amphibium 666 
arifolium 673 
articulatum 676 
atlanticum 663 
aviculare 659, 61 
“ angustissimum 662 
Bellardi 663 
buxiforme 662 
camporum 063 
Careyi 668 
cilinode 674 
“ erectum 674 
Convolvulus 674 
cristatum 675 
cuspidatum 676 
densiflorum 667 
Douglasii 664 
dumetorum 675 
emersum 666 
erectum 663 
exsertum I 661 
Fagopyrum 672 
Fowleri 660 
glaucum 660 
Hartwrightti 666 
Hydropiper 670 
hydropiperoides 669 
** Macouni 669 
incarnatum 666 
lapathifolium 666 
“incanum 666 
“ nodosum 666 
leptocarpum 661 
littorale 662 
longistylum 667 
maritimum 660 
mite 669 
monspeliense 661 
Muhlenbergit 666 
neglectum 662 
opelousanum 669 
orientale 670 
pennsylvanicum 667 
Persicaria 665, 6 
persicarioides 668 
portoricense 667 
prolificum 662 
provinciale 662 
punctatum 670 
“leptostachyum 670 
“ robustius 670 
Rayi 660 
ramosissimum I 663 
“ atlanticum 663 
* prolificum 662 
Sagittatwm 673 


590 GENERAL INDEX OF LATIN GENERA AND SPECIES. VoL EE: 
Polygonum Populus Potamogeton Primula 
scandens I 674 occidentalis I 591 trichoides r 81 mistassinica 2 708 
setaceum 669 Sargentii 591 Tuckermani 81 veris 708 
tataricum 672 tremuloides 590 varians 78 | Prinos 
tenue 664 | Porteranthus 2 248 Vaseyi 83 see Ilex 2 486 
tomentosum 667 (Gillenia) “ latifolius 83 | Prionopsis 3 377 
triangulum 664 stipulatus 248 Zizi 79 (Aplopappus) 
virginianum 665 trifoliatus 248 zosteraefolius ‘81 ciliata 377 
viviparum 671 | PoRTULACACEAE 2 35 | Potentilla 2 249 | Prosartes 
Zuccarinit 676 | Portulaca 2 39 Anserina 258 lanuginosa ry yay 
Polymnia 3 458 grandiflora 41 argentea 253 trachycarpa 518 
canadensis 458 neglecta 40 arguta 263 | Proserpinaca 2 613 
“ radiata 458 oleracea 39, 40 bipinnatifida 257 intermedia 613 
Uvedalia 458 pilosa 40 canadensis 251 palustris 613 
Polyotus 3 34 retusa 40 “ pumila 251 pectinacea 613 
angustifolius 34 triangularis 35 canescens 254 pectinata 613 
PoLYPODIACEAE I 10 | Portuna 2 689 caroliniana 251 | Prosopis 2.332 
Polypodium x35 floribunda 690 collina 253 glandulosa 333 
bulbiferum 14,5 | Potamogeton I 74 effusa 256 juliflora 333 
calcareum 24 alpinus 7 emarginata 255 | Prunella 3 iis 
cristatum 19 americanus 77 frigida 255 (Brunella) 
dilatatum 21 amplifolius 76 fruticosa 262 laciniata 115 
Dryopteris 23 “ ovalifolius 76 inclinata 254 vulgaris 115 
Filix-foemina 30 angustifolius 79 Hippiana 256 | Prunus 2 322 
Filix-mas 17, 21 “ methyensis 79 hirsuta 253 alleghaniensis 324% 
fontanum 29 bipleuroides 80 intermedia 254 americana 323 
fragile 15 Claytonii 77 labradorica 253 angustifolia 324 
fragrans 19 compressus 81-3 leucocarpa 252 Avium 327 
hexagonopterum 23 confervoides 81 leucophylla 256 Besseyi 327 
incanum 36 crispus 81 littoralis 257 cerasifera 325 
intermedium 22 dimorphus 86 maculata 254 Cerasus 327 
Lonchitis 16 diversifolius 85 millegrana 252 Chicasa 324 
marginale 20 epihydrus 77 minima 255 cuneata 326 
montanum 15 “ vars. 85 monspeliensis 253-4 domestica 2 322, 326 
noveboracense 18 Faxoni 78 multifida 2 256 gracilis 325 
obtusum 14 filiformis I 86 nana 255 Gravesii 325 
Phegopteris 23| “ Macounii 86 nemoralis 250 hortulana 324 
polypodioides 36 “ occidentalis 86 Nicolletii 252 “ Mineri 324 
Robertianum 24 flabellatus 87 nivea 255 insititia 326 
spinulosum 21 fluitans 77 norvegica 253 Mahaleb 328 
vulgare 35,6 foliosus 82 palustris 258 maritima 325 
Polypogon I 201 Friesii 83 paradoxa 2y2 mollis 323 
monspeliensis 201 gemmiparus 85 pectinata 257 nana 329 
Polypremum REF ZT gramineus 78 pennsylvanica 257 nigra 323 
procumbens 731 heterophyllus 78 “ strigosa 257 pennsylvanica 328 
Polypteris 3 507 “ vars. 78 pentandra 253 pumila 326 
callosa 507 Hillii 82 pilosa 254 serotina 329 
Hookeriana 507 hybridus 85 procumbens 250  Smallii 329 
Polystichum I 16 illinoensis 79 procumbens 261 sphaerocarpa 325 
acrostichoides 16 interior 86 pumila 251 virginiana 329 
“ Schweinitzit 16 interruptus 87 Ranunculus 254 Watsoni 324 
Braunii 17 lateralis 84 recta 254 | Psedera 2 510 
Lonchitis 16 lonchites 77 reptans 250 | Pseudacacia 
scopulinum 17 “ noveboracensis 77 rivalis 252-3 odorata 2 375 
Polytaenia 2 632 “ connecticutensis 79 Robbinsiana 255 | Pseudotaenidia 2 630 
Nuttallit 633 lucens 79 rubens 254 montana 631 
POMACEAE 2 286 major 83 salisbrugensis 254 | Psilocarya I 346 
Pomaria marinus 86 simplex 251 nitens 347 
glandulosa 2 338 mysticus 8o strigosa 257 rynchosporoides 347 
PONTEDERIACEAE I 462 natans 74,5 sulphurea 254 scirpoides 346-7 
Pontederia 462 niagarensis 82 supina 252] Psilostrophe 3 504 
cordata 463 nove boracensis 77 tridentata ., 262 (Riddellia) 
“ angustifolia 463 Nuttallii 77 Vahliana 255 gnaphalodes 504 
lancifolia 463 Oakesianus 76 | Poteridium 2 265 Tagetinae 504 
limosa 464 obtusifolius 82 annuum 265 villosa 504 
Populus I 587 parormitanus 84 | Poterium 2 265 | Psoralea 2 360 
acuminata 589 pauciflorus 82 annuum 265 argophylla 363 
alba 587 pectinatus I 87 canadense 265 aurea 368 
angulata 590 perfoliatus 80 Sanguisorba 266 bituminosa 360 
angustifolia 588 “ lanceolatus 80 | Prasium collina 362 
balsamifera 588 “ Richardsonii 80 purpureum 3107 cryptocarpa 363 
candicans 588 pinnatum 2 616 | Prenanthes 3 334 cuspidata 363 
carolinensis 590 praelongis I 80 see Nabalus 3 335-8 Dalea 367 
coloradensis 589 pulcher 76 juncea 322 digitata 362 
deltoides 590 pusillus 84 mainensis 337 esculenta 363 
“ occidentalis 591 “ vars. 83-6 pauciflora 312 floribunda 361 
dilatata 590 Robbinsii 87 runcinata 312 hypogaea 364 
grandidentata 5890 rufescens 77 tenuifolia 312 imcana 362 
heterophylla 589 rutilus 83 | PRIMULACEAE 2 707 lanceolata 361 
italica 590 spathulaeformis 78) Primula 2 708 linearifolia 362 
monilifera 590 Spirillus 86 egaliksensis 709 longifolia 386 
nigra 590 “ curvifolius 86 farinosa 708 macrorhiza 363 


Vou, III. GENERAL INDEX OF LATIN GENERA AND SPECIES. sgl 
Psoralea Pyrrhopappus Quercus Ranunculus 
melilotoides 2 364 carolinianus 3 324 tridentata I 622 obtusiusculus 2 107 
micrantha 361 scaposus Ben triloba 619, 17 Ovalis 109 
obtusiloba 361 | Pyrularia I 640 velutina 619 parviflorus 114 
Onobrychis 365 oleifera 641 virens 625 parvulus 114 
pedunculata 364 pubera 641 virginiana 625 pedatifidus 108 
stipulata 364 | Pyrus 2 287 | Queria 2 30 pennsylvanicus 112 
tenuiflora 361 arbutifolia 290 canadensis 30 Philonotis 114 
Ptelea 2 444 “ atropurpurea 291 capillacea 30 Purshii 105 
tomentosa 445 americana 287 | Quinaria 2 510 pusillus 106 
trifoliata 445 angustifolia 288 | Quincula 3 162 pygmaeus 108 
“ mollis 445 baccata 289 (Physalis in part) recurvatus III 
Pteridium Tan Botryapium 292 lobata 162 repens [12,3 
aquilinum 32 communis 288 reptans 107 
pseudocaudatum 32 coronaria 288 | Radicula 2 159| . rhomboideus 109 
caudatum 31 10ensis 289 aquatica 164 Sabini 108 
PTERIDOPHYTA gt Malus 288-90 curvisiliqua 162 sceleratus III 
Pteris melanocarpa 291 hispida 161 Septentrionalis 113 
alabamensis re Syst nigra 291 Nasturtium-aquat- sicaeformis 113 
aquilina 32 sanguinea 203 icum 162 trichophyllus 116 
atropurpurea 33 see Sorbus 287 obtusa 160 | Raphanus 2 194 
Stelleri Re Malus 288 palustris 161 Raphanistrum 195 
Pterospora 2 673 | Pyxidanthera 2 706 sessiliflora 162 sativus 194-5 
Andromedea 673 barbulata 706 sinuata 160 | Rapistrum 
Ptilimnium 2 656 2 sphaerocarpa 161 rugosum 2 159 
(Discopleura) Quamasia I 508 sylvestris 160 | Ratibida 3 473 
capillaceum 657 (Scilla) ~ - Radiola (Lepachys) 
missouriense 657 esculenta 509 linoides 2 440 columnaris 474 
Nuttallii 657 hyacinthina 509 | Raimannia 2 596 pinnata 474 
Ptiloria 3 312| Quamoclit 3 42 coronopifolia 598 Tagetes 475 
(Stephanomeria) coccinea 42, 3 humifusa 597 | Razoumofskya_ 1 638 
pauciflora 312 Quamoclit 42 laciniata 597 (Arceuthobium) 
ramosa 312 vulgaris 42 rhombipetala 597 pusilla 638 
tenuifolia 312 | Quercus I 616 | Rajania ovata 1 677 | Redfieldia I 235 
Puccinellia I 267 acuminata 624 | RANUNCULACEAE 2 84 flexuosa 236 
(Glyceria in part) alba 622 | Ranunculus 2 104 | Relbunium 
airoides 268 Alexanderi 625 abortivus 110 bermudense 3 206 
angustata 267 aquatica 621 “ encyclus 110 | Renaelmia 
Borreri 268 bicolor 623 * Harveyi 109 usneoides I 456 
distans 267 borealis 618 “ micranthus 110 | RESEDACEAE 2 199 
fasciculata 268 Brittoni 620 Alleni 109g | Reseda 2 200 
maritima 267-8 Catesbaei 616 acris III-2 alba 201 
Pueraria coccinea 619, 18 affinis 108 lutea 200 
Thunbergiana 2 420 “ tinctoria 619 alismaefolius 107 Luteola 200 
Pulmonaria digitata 617,19 allegheniensis 110 odorata 200 
see Mertensia 3 83 elipsoidalis 618 ambigens 107 | RHAMNACEAE 2 501 
maritima 82 falcata 619 aquatilis 116 | Rhamnus 2 502 
Pulsatilla 2 102 “pagodaefolia 620 “ caespitosus 116 alnifolia 503 
(Anemone in heterophylla 621 “ capillaceus 116 caroliniana 503 
part) ilicifolia I 620 “ divaricatus 116 cathartica 502 
hirsutissima 102 imbricaria 622 arvensis II5 Frangula 503 
patens 102 laurifolia 621 auricomus 104 lanceolata 503 
Pycnanthemum 3 141 Leana 622 bulbosus 112 | Rhexia 2 582 
see Koellia I41—5 lyrata 623 cardiophyllus 108 aristosa 583 
lanceolatum 142 macrocarpa 622-3 circinatus 116 ciliosa 584 
linifolium 141 margaretta 622 Clintoni 113 latifolia 583 
Monardella 132 marylandica 620-2 Cymbalaria 117 mariana 583 
Torreyi 143 Michauxii 624 delphinifolius 105, 17 petiolata 584 
Tullia 144 minor 622 “ terrestris 105 virginica 000 
Pycraeus Muhlenbergii 624, 5 eremogenes 111 | Rhinanthus 3 223 
sabulosus I 300 nana 620 fascicularis II4 Crista-galli 223 
Pyrethrum nigra I 621 Ficaria Du major 223 
Balsamita CY Tai) “ digitata 619 filiformis 107 minor 223 
inodorumnanum 520 “ tridentata 622 Flammula 107 virginicus 208 
PYROLACEAE 2 067 obtusiloba 622 “veptans 107 | Rhodiola 2 206 
Pyrola 2 668 olivaeformis 623 “ intermedius 107 rosea 206-7 
americana 668 pagodaefolia 620 Harveyi 109 | RHODODENDREAE 2 676 
asarifolia 670 palustris 617-22 hederaceus 116| Rhododendron 2 680 
chlorantha 669-70 Phellos 621 hispidus 113 arborescens 679 
elliptica 669 platanoides 623 hispidus 112 calendulaceum 678 
grandiflora 668 prinoides 625 hyperboreus 2 105 canadense 680 
incarnata 669 “ rufescens 625 lacustris 105 canescens 678 
maculata 672 Prinus 622-4 lapponicus 106 catawbiense 681 
minor 670 Robur 616 limosus 105 ferrugineum 680 
oxypetala 670 rubra I 617 Macounii 112 lapponicum 680 
rotundifolia 668 “ nana 620 micranthus 110 maximum 681 
“ pumila 668 “ runcinata 617 missouriensis 105 nudiflorum 678 
secunda 671 Rudkini 620 multifidus 105 Rhodora 680 
uliginosa 669 Schneckii 618 “vepens 105 viscosum 679 
“ pumila 671 stellata 622 muricatus 115 | Rhodora 2 679 
umobellata 672 texana 618 nivalis 107 canadensis 680 
uniflora 671 tinctoria 619 oblongifolius 106 | Rhombolytrum 1 238 


592 GENERAL INDEX OF LATIN GENERA AND SPECIES. Vo. IIT. 
Rhombolytrum Roripa Rubus Rynchospora 
albescens I 238 americana 2 164 triflorus 2 278 axillaris I 344 
Rhus 2 481 Armoracia 163 trivialis 281 “microcephala 344 
aromatica 482-3 | ROSACEAE 2 242 villosus 280 capillacea 343 
bipinnata 482 | Rosa 2 282 “ frondosus 279 “ laeviseta 343 
canadensis 482 acicularis 283 “ humifusus 280 cephalantha 344 
copallina 481 blanda 283 “ montanus 280 compressa 345 
coriaria 481 bracteata 286 | Rudbeckia 3 469 corniculata 342 
cotinoides 485 canina 284 amplexicaulis 473 cymosa 345 
glabra 482 carolina 285 angustifolia 476 etuberculata 331 
“borealis 482 centifolia 282 aspera 472 filifolia 344 
hirta 481 cinnamomea 286 atrorubens 476 fusca 344 
microcarpa 484 eglanteria 286 Brittonii — 471 glomerata 344 
quercifolia 484 Engelmanni 283 columnaris 474 gracilenta 345 
radicans 484 Fendleri 284 fulgida 471 inexpansa 346 
Toxicodendron 484 gallica 286 grandiflora 472 Knieskernii 343 
trilobata 483 humilis 285 hirta 469-70 macrostachya 342 
typhina 481 lucida 285 laciniata 473 nitens 347 
venenata 483 micrantha 286 “ humilis 473 oligantha 342 
Vernix 483 nitida 285 maxima 472 pallida 342 
Rhynchosia 2 421 parviflora 285 MiSSOUrIENSIS 471 rariflora 344 
erecta 422 pratincola 284 monticola 470 scirpoides 347 
latifolia 421 rubiginosa 286 pallida 476 Smallii 345 
reniformis 422 Sayi 283 palustris 471 Torreyana 346 
simplicifolia 422 setigera 283 pinnata 474 ‘ 
tomentosa 421 spinosissima 284 purpurea 475 | Sabbatia 33 
“ monophylla 422 virginiana 285 spathulata 471 angularis 5 
“ volubilis 421 Woodsii 284 speciosa 472 “ angustifolia 5 
Rhynchospora Rotala 579 subtomentosa 470 brachiata 5 
see Rynchospora ramosior 579 Sullivantii 472 calycina 5 
I 341 verticillaris 579 Tagetes 475 calycosa 5 
Ribes 2 236 | Rothia triloba 470 campanulata 7 
americanum 238 carolinensis 505 umbrosa 471 campestris 6 
cereum 238 | Rottboellia 111 | Ruellia 3 240 chloroides 7 
Cynosbati 239 corrugata 112 biflora 240 dodecandra 7 
“ glabratum 239 cylindrica Ti2 ciliosa 241 Elliottii 6 
floridum 238 filiformis 282 “ parviflora 242 gracilis 7 
glandulosum 238 rugosa I12 “ ambigua 242 lanceolata 4 
gracile 240 | Roubieva 215 oblongifolia 240 paniculata 4,6 
Grossularia 241 multifida 15 parviflora 242 stellaris 6 
hirtellum 241 | Rubacer 2 275 pedunculata 242 | Sacciolepis I 163 
hudsonianum 237 | RUBIACEAE 3 250 strepens 241 gibba 163 
huronense 241 | Rubus _2 275 “ cleistantha 241 striata 163 
inebrians 238 acaulis 278 tuberosa 241 | Saecharum TRUE 
lacustre 236 alleghaniensis 2804 Rumex I 653 japonicum 113 
MiSSOUrIENSIS 240 americanus 278 Acetosa 654 polydactylon 113 
nigrum 237 arcticus 277 Acetosella 653 | Sagina 2 51 
odoratum 239 argutus 280 altissimus 656 apetala Bt 
oxyacanthoides 240 Baileyanus 280 Brittannica 656 decumbens 51 
“ calcicola 241 canadensis 279 conglomeratus 657 © Smithii 51 
“ lacustre 236 “ roribaccus 281 crispus 657 fontinalis 46 
pennsylvanicum 238 Chamaemorus 276 digynus 659 Linnaei 51 
prostratum 238 columbianus 276 elongatus 657 nivalis 51 
reclinatum 241 cuneifolius 278 Engelmanni 654 nodosa 52 
rotundifolium 241 Dalibarda 282 hastatulus 654 procumbens 51 
rubrum 236, 7 Ensleni 281 Hydrolapathum saginoides 51 
“ subglandulosum 237 frondosus 279 “americanum 656 subulata 51 
sSaxosum 241 fruticosus 275 maritimus 659 virginica 3 16 
setosum 240 hispidus 281 mexicanus 655 | Sagittaria I 98 
triste 237 * suberecta 279 obtusifolius 657, 8 ambigua IOI 
Uva-crispa 241 idaeus 277 occidentalis 657 arifolia 99 
vulgare 237 laciniatus 280 orbiculatus ” 656 “ stricta 99 
Ricinophyllum 2 619 Millspaughii 279 pallidus 655 brevirostra 100 
Ricinus 2 460 montanus 280 Patientia 653, 6 calycina 96-7 
communis 461 neglectus 2A Gf persicarioides 659 “ spongiosa 97 
Ridan 3 486 nigricans 279 pulcher 658 cristata 102 
alternifolia 487 nigrobaccus 280 salicifolius 655 cuneata 99 
Riddellia nutkanus 276 sanguineus 658 Eatonii 103 
Tagetinae 3 504 obovalis 281 venosus 654 Engelmanniana 99 
Ripidium occidentalis 277 verticillatus 655 falcata 101 
japonicum DiLn3 odoratus 276 | Ruppia 1 88 graminea 103 
Robertiella 2 425 parviflorus 276 curvicarpa 88 heterophylla 102 
Robertiana 426 parvifolius 278 lacustris 88 lancifolia 101 
Robertsonia 2 219 phoenicolasius 277 maritima 88 latifolia 100 
micranthidifolia 219 procumbens 281 occidentalis 83 longiloba IOI 
Robinia 2 374 Randii 280 | RUTACEAE 2 443 longirostra 99 
hispida 375 rubrisetus 281 | Ruta 445 lorata 104 
pseudacacia 375 saxatilis graveolens 445 natans gracillima 104 
viscosa 375 “ americanus 278 | Rynchospora I 341 platyphylla 103 
Roripa 2 159 “ canadensis 278 alba 343 pubescens 100 
see Radicula 2159-62 setosus 279 “ macra 343 pusilla 104 
(Nasturtium) strigosus 277. aurea 341 radicans 96 


Vo. III. 


Sagittaria 
rigida 
sagittifolia 
subulata 
teres 
variabilis 
“ diversifolia 
* gracilis 

SALICACEAE 

Salicornia 
ambigua 
Bigelovii 
europaea 
“ prostrata 
herbacea 
mucronata 
rubra 
virginica 

Salix 
acutidens 
adenophylla 
alba 
amygdaloides 
anglorum 
angustata 
arctica 
argyrocarpa 
atra 
babylonica 
balsanufera 
Barclayi 
Bebbiana 
brachycarpa 
Brown 
calcicola 
callicarpaea 
candida 
chlorolepis 
chlorophylla 
coactilis 
cordata 
cordifolia 
Cutleri - 
desertorum 
discolor 
eriocephala 
exigua 
falcata 
Fernaldii 
fluviatilis 

“ exigua 
fragilis 
fuscescens 
glauca 
gracilis 
groenlandica 
glaucophylla 
herbacea 
humilis 
interior 
labradorica 
laurentiana 
latiuscula 
liniarifolia 
longifolia 
longipes 
lucida 

lutea 
luteosericea 
Mackenziana 
Macounii 
MissoUriensis 
myrtillifolia 
myrtilloides 
nigra 

“ falcata 

“ Wardi 
obtusata 
orbicularis 
pedicellaris 
pellita 
pentandra 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 593 
Salix Santolina Scandix 
I 102 perrostrata I 599 suaveolens gecior. Pecten-Veneris 2 627 
99 petiolaris 598, 9 | SAPINDACEAE 2 500 procumbens 6290 
104 phylicifolia 600 | Sapindus 2 500] Schedonnardus 1 226 
102 prinoides 599 acuminatus 500 Paniculatus 226 
100 purpurea 597 Drummondii 500 texanus 226 
100 pyrifolia 596 marginatus 500 | SCHEUCHZERIACEAE I QI 
99 reticulata 602 Saponaria 500 | Scheuchzeria I 93 
I 587 rostrata 599 | Saponaria 272 palustris 93 
Ch Din sericea 598 officinalis 73| Schizachyrium 1 115 
22 serissima 594 Vaccaria Gis) littorale 115 
22 squamata 600 | SAPOTACEAE 2719 scoparium 115 
21 stricta 604 | Sarcobatus 2 22| ScHIZAEACEAE 19 
21 subsericea 598 vermicularis 23 | Schizaea 19 
21 syrticola 597 | Sarothamnus dichotoma 9 
22 tristis I 600 scoparius 2 350 pusilla 9 
21-2 Uva-ursi 601 | Sarothra 2 536| Schizonotus 2 247 
22 vacciniformis 605 Drummondii 536 sorbifolius 247 
I 591 vestita 603 gentianoides 536) Schoenus 
596 viminalis 597 | SARRACENIACEAE 2 201 albus I 343 
597 vitellina | 595 | Sarracenia 2 201 coloratus 341 
591, 5 Waghornei 604 flava 202 corniculatus 342 
593 Wardi 593 heterophylla 202 effusus 348 
605 Wheeleri 595 purpurea 201-2 Mariscus 347 
596 | Salomonia I 521 | Sassafras 2 134 mariscoides 348 
04, 5 biflora 521 officinale 134 rufus 332 
603 commutata 521 variifolium 134| see Rynchospora 
605 | Salpingia 2 606 Sassafras 134 I 341-6 
595 | Salsola 2 24| Satureia 3 137 | Schmaltzia 2 482 
596 caroliniana 2 hortensis 137 aromatica 482 
603 depressa 2 nepeta 138 crenata 482 
599 Kali 25 origanoides 146 trilobata 483 
604 “ rosacea 25 Thymus virg. 142 | Schollera I 463 
605 “ tenuifolia 25 virginiana 142 graminea 464 
602 linearis 23 virginica 142] Schrankia 
604 pestifer 25 | SAURURACEAE 1577 aculeata 2 333 
I 598 salsaamericana 23 | Saururus E577. angustata 334 
602 Soda 24 cernuus 578 uncinata 333 
600 Tragus 25 | Savastana I 171 | Schwalbea 3 216 
598 | Salvia 3 128 (Hierocloé) americana 217 
596 azurea grandiflora 129 alpina 173 | Schweinitzia 2 673 
604 lanceolata 130 hirta 171 caroliniana 674 
601 lancifolia 130 Nashii 172 odorata 67 
604 lyrata 129 odorata 172 | Scilla 
599, 600 officinalis 128 pauciflora 272 esculenta I 509 
599 Pitcheri 129 | SAXIFRAGACEAE 2 214 Fraseri 509 
594| pratensis 129 | Saxifraga 2 221 | Scirpus I 326 
592 reflexa 130 aconitifolia 22 acicularis 315 
603 Sclarea 131 aizoides 216-7 acuminatus 318 
595 urticifolia 130 Aizoon 22 americanus 330 
5904 Verbenaca 130 autumnalis 217 “longispicatus 330 
594 verticillata 130 caespitosa 222 atrocinetus 337 
602 | SALVINIACEAE 137, caroliniana 219 atropurpureus 312 
605 | Salvinia L337 cernua 222 atrovirens 335 
599 natans 38 comosa 220 autumnalis 322 
605 | Sambucus 3 268 erosa 219 caespitosus 328 
596 aanadedare 268 pengloss mee californicus 332 
or . size orbesit campestris 333 
600 pair ciare rh a Geum 220 Canbyi 331 
595 “ dissecta 268 granulata gat capillaris 319 
605 Grayana 219 capitatus 313 
599 racemosa 268 Hirculus 216 castaneus 320 
603 Samolus 2 710 leucanthemifolia Clintoni 328 
595 floribundus 710 220-1 cylindricus 331 
595 Valerandi 710 Michauxii 221 cyperinus 337 
593 americanus 710 micranthidifolia 219 debilis 329 
I 593 | Sanguinaria 2 140 muscoides 222 divaricatus 336 
596 canadensis 140 nivalis 218 equisetoides 311 
594 | Sanguisorba 2 264 oppositifolia 216 Eriophorum 337 
596 annua 265 pennsylvanica 219 etuberculatus 331 
605 canadensis 265 rivularis 222 Fernaldi 33 
596 minor 266 stellaris 221 flaccidus 312 
601 officinalis 265 Sullivantii 224 fluviatilis 333 
602 Sanguisorba 266 texana 218 georgianus Se 
592 | Sanicula 2 62 tricuspidata 217 Hallii 328 
592 canadensis 62 virginiensis 218 heterochaetus 332 
593 europaea 623 | Scabiosa 3 280 hudsonianus 323 
601 gregaria 62 arvensis 290 interior 333 
602 marylandica 62 australis 290 intermed us 318 
602 Smallhii 62 succisa 290 interstinctus 31t 
598 trifoliata 624 | Scandix 2 626 lacustris 326, 331 
594 | SANTALACEAE I 639 Cerefolium 629 lineatus 337 


yee 


an 


594 GENERAL INDEX OF LATIN GENERA AND SPECIES. Vor. III. 
Scirpus Scrophularia Senecio Sibbaldiopsis 2 261 
Longit ¥ 337 nodosa 3 179 Douglassii 3 541 tridentata 262 
maritimus occidentalis 180 Elliottii 545 | Sicyos 3 293 
“ cylindricus 331 | Scutellaria 3 105 Fremontit 541 angulatus 293 
“ fAuviatilis 333 ambigua 108 frigidus 542 lobatus 292 
“ macrostachyus 333 Brittonii 108 glabellus 540 | Sida 2 519 
micranthus 339 Bushii 109 gracilis 544 Abutilon 521 
microcarpus Tesss campestris 108 hieracifolius 536 alnifolia 520 
mucronatus 331 canescens 106 integerrimus 546 alceoides 516 
mutatus 311 cordifolia 107 Jacobaea 542 Elliottii 520 
nanus 327 Churchilliana 109 lobatus 540 hederacea 521 
nitens 347 Drummondii 108 lugens 546 hermaphrodita 520 
novae-angliae 334 galericulata 109 lyratus 540 hispida 519 
obtusus 313 hirsuta 107 Nuttallii 537 Napaea 520 
occidentalis 332 hyssopifolia 107 obovatus 545 spinosa 520 
Olneyi 330 incana 106 palustris 540 stellata 522 
“ contortus 330 integrifolia 107 pauciflorus 544 | Sideranthus 3 378 
ovatus 313 laevigata 106 pauperculus 545 (Eriocarpum) 
pallidus 335 lateriflora 106, 9 plattensis 543 annuus 378 
paludosus 333 nervosa 110 Pseudo-arnica 541 glaberrimus 379 
palustris 310, 314 ovalifolia 107 pseudoaureus 544 grindelioides 378 
pauciflorus 327 parvula 108 pseudotomentosus spinulosus 378,9 
Peckii 336 “ mollis 108 3 543 | Sideritis 3 110 
pedicellatus I 337 peregrina 105 Purshianus 543 hirsuta III 
planifolius 328 pilosa 107 Riddellii 541 romana III 
polyphyllus 336 resinosa 108 Robbinsii 544 | Sideroxylon 2-719 
puberulus 321 saxatilis 109 rotundus 545 lanuginosum 720 
pungens 330 serrata 106 Smallii 546 | Siegesbeckia 3 488 
quadrangulatus 311 teucriifolia 110 spartioides 541 occidentalis 488 
retrofractus 307 versicolor 107 suaveolens 538 | Sieglingia I 232 
robustus 333 Wrightii 108 sylvaticus 539 albescens 238 
rosteliatus 319 | Sedum 2 207 tomentosus 543 decumbens 233 
rubrotinctus 335 acre 208 viscosus 540 elongata 234 
rufus 332 Fabaria 207 vulgaris 538-9 pilosa. 235 
simplex 316 Nevii 210 | Serapias I 563 purpurea 235 
Smithii 329 Nuttallianum 208 Helleborine 563 seslerioides 233 
spadiceus 320 pulchellum 209 viridiflora 563 stricta 234 
stenophyllus 319 purpureum 207 | Sericocarpus 3 405 | Sieversia 2 272 
subterminalis 329 reflexum 209 asteroides 406 anemonoides 272 
“ terrestris 329 Rhodiola 207 bifoliatus 406 ciliata 273 
supinus Hallit 329 roseum 207 conyzoides 406 Peckii 272 
sylvaticus 334 Sieboldi 208 linifolius 405 Tadiata 272 
“ digynus 335 sparsiflorum 208 solidagineus 405 | Silene 2 62 
tenuis 318 stenopetalum 209 tortifolius 406 acaulis 63 
Torreyi 330 stoloniferum 210 |. Serinia 3 306 alba 63 
tortilis 316 telephioides 208 (Apogon) anglica 62, 67 
tuberculosus 316 Telephium 207 oppositifolia 306 antirrhina 66 
Vahlii 321 ternatum 210 | Serpicula I 105 “ divaricata 66 
validus 331 Torreyi 208 angustifolia 105 Armeria 66 
Wolf 315 triphyllum 207 occidentalis 105 caroliniana 65 
Scleranthus 2 30] SELAGINELLACEAE I 48 verticillata 105 conica 66 
annuus 30 | Selaginella 1 48 | Serratula dichotoma 67 
Scleria I 348 apus 49 see Lacinaria 3 365 Drummondi 70 
ciliata 350 rupestris 49 Vernonia 350-1 gallica 67 
flagellum-nigrorum selaginoides 49 arvensis 553 inflata 64 
348 spinosa 49 | Sesban 2 376 latifolia 64 
laxa 350 | Selenia 2 189 (Sesbania) Menziesii 68 
oligantha 349 aurea 190 macrocarpa 376 Nivea 63 
pauciflora 350 | Selinum Sesbania 2 376 noctiflora 67 
“ Elliottii 350 acaule 2 644} Seseli nocturna 67 
reticularis 349 canadense 636 divaricatum 2 643 nutans 64 
“ obscura . 349 | Sempervivum 2 210} Sesleria ovata 63 
“ pubescens 350 tectorum 211 dactyloides Te2gs pennsylvanica 65 
setacea 350 | Senebiera Sesuvium 2 34 racemosa 67 
Torreyana 350 Coronopus 2 167 maritimum 34 regia 65 
triglomerata 349 didyma 167 pentandrum 34 rotundifolia 65 
* gracilis 349 | SENECIONEAE 3 350 Portulacastrum 34 stellata 63 
verticillata 351 | Senecio 3-538 sessile 34 virginica 64 
Sclerolepis 3 355 antennariifolius 542 | Setaria I 164| vulgaris 64 
uniflora 355 atriplicifolius 537 see Chaetochloa 164] Silphium 3 459 
verticillata 355 aureus 544 | Setiscapella 3 230 Asteriscus 460-1 
Scolochloa I 261 “angustifolius 546 cleistogama 231 compositum 462 
festvcacea 262 “ Balsamitae 545 subulata 231 integrifolium 460 
Scolopendrium “ compactus 546 | Seymeria laciniatum 461 
Scolopendrium E25 || * discoideus 544 macrophylla 3 205 Nuttallianum 464 
vulgar 25 “ obovatus 545 | Shepherdia perfoliatum 460 
SCROPHUBARIACEAE Balsamitae 545 argentea 2 576 reniforme 462 
‘ 3 172 “ Crawfordit 545 canadensis 576 speciosum 460 
Scrophulariq 3:79] canus 543 | Sherardia 3 266 terebinthinaceum 462 
leporella 180 Crawfordii 545 arvensis 266 “pinnatifidum 462 
marylandica 179 densus 546 | Sibbaldia 2 261 trifoliatum 461 
neglecta 180 discoideus 544 procumbens 261 | Silybum 3555 


Vor. III. 
Silybum 
marianum 30555 
SIMARUBACEAE 2-445 
Sinapis 2 191 
alba IgI 
arvensis 192 
juncea 193 
nigra 193 
Sison 2 630 
canadense 630 
Sisymbrium 2 162 
Alliaria 170 
altissimum 174 
amphibium 159 
“ palustre 161 
canescens 171 
dentatum 140 
humifusum 179 
Hartwegianum 171 
humvile 176 
incisum 171 
Trio 174 
leiocarpum 174 
murale 194 
Nasturtium- 
aquaticum 162 
officinale 174 
pannonicum 174 
Sinapistrum 174 
Sophia 170 
strictissimum 174 
sylvestre 160 
tenuifolium 194 
Thalianum 176 
sophia | 170 
Sisyrinchium I 542 
albidum 543 
anceps 545 
angustifolium 543 
apiculatum 545 
arenicola 544 
atlanticum 545 
Bermudiana 542 
campestre 543-4 
“ kansanum 543 
Farwellii 544 
flaviflorum 543 
gramineum 545 
graminoides 545 
hastile 543 
intermedium 544 
montanum 543 
mucronatum 544 
Pringlei 543 
septentrionale 543 
strictum 545 
versicolor 543 
Sitanion I 288 
brevifolium 288 
elymoides 288 
longifolium 288 
Sitilias 3 324. 
(Pyrrhopappus) 
caroliniana 324 
grandiflora 325 
Sium 2 656 
angustifolium 655 
Carsoni 656 
cicutaefolium 656 
erectum 655 
latifolium 656 
lineare 656 
longifolinm 638 
rigidum 637-8 
SMILACEAE I 526 
Smilacina rose 
bifolia 517 
see Vagnera 1 515-6 
Smilax I 526 
aspera 527 
Bona-nox 529 
caduca 528 


Smilax 
ecirrhata I 528 
glauca 528 
hastata 529 
herbacea 527 
hispida 529 
lanceolata 530 
laurifolia 530 
Pseudo-China 529 
pulverulenta 527 
quadrangularis 528 
rotundifolia 528 
spinulosa 528 
tamnifolia 527 
tamnoides 529 
Walteri 530 


Smyrnium 


atropurpureum 2 639 


aureum 641 
barbinode 640 
cordatum 642 
integerrimum 640 
nudicaule 637 
SOLANACEAE 3.154 
Solanum 3 164 
carolinense 165 | 
citrullifolium 166 
conioides 163 
Coronopus 163-4 
Dulcamara 167 
elaeagnifolium 165 
heterandrum 166 
heterodoxum 166 
Lycopersicon 168 
Melongena 167 
nigrum 164 
rostratum 166 
sisymbriifolium 167 
Torreyi 166 
triflorum 165, 
triquetrum 167 
villosum 165 
Solea 2 563, 4 
concolor 564 
Solidago 3 380 
alpestris 385 
altissima 395 
angustifolia 389 
arguta 393 
“ scabrella 393 
aspera 390 
axillaris 382 
bicolor 383 
“ concolor 384 
Bigelovii 385 
Boottii 391 
caesia 382 
“ axillaris 382 
calcicola 385 
canadensis 393 
“ glabrata 393 
“ procera 395 
“ scabra 395 
chrysolepis 388 
cordata 401 
corymbosa 397 
Curtisii 383 
“ monticola 384 
Cutleri 385 
decumbens 386 
Drummondii 396 
Elliottii 3 392 
elliptica 392 
erecta 384 
fistulosa 390 
flexicaulis 383 
Gillmani 388 
gilvocanescens 393 
Gattingeri © 395 
Gillmani 388 
glaberima 394 


graminifolia 398 


Solidago 
gigantea 3 394 
gracilis 382 
Harrisii 393 
gymnospermoides 399 
hirsuta 384 
hispida 384 
Houghtonii 398 
humilis Gillmani 388 
incana 396 
juncea 393 
“ramosa 393 
“ scabrella 393 
lanceolata 398 
“ minor 400 
lateriflora 426 
latifolia 383 
leptocephala 399 
Lindheimerana 385 
linoides 392 
longipetiolata 395 
macrophylla 385 
microphylla 391 
minor 400 
missouriensis 394 
mollis 396 
monticola 387 
moritura 394 
Moseleyi 400 
Muhlenbergii 393 
multiradiata 386 
neglecta 392 
nemoralis 395 
“ arenicola 395 
“ incana 396 
odora 389 
“ inodora 389 
ohioensis 397 
pallida 387 
patula 391 
petiolaris 382 
pilosa 390 
procera 395 
Pitcheri 394 
polycephala 399 
pubens 383 
puberula 386 
pulcherima 395 
Purshii 388 
Randii 388 
racemosa 388 
“ Gillmani 388 
retrorsa 390 
Riddellii 397 
rigida 397 
rigidiuscula 387 
rugosa 390 
rupestris 393 
Sarothrae 370 
sciaphila 389 
sempervirens 3890 
serotina 394 
“ gigantea 394 
Shortii 304 
sphacelata 401 
speciosa 387 
“ angustata 384 
“ pallida 387 
“ rigidiuscula 387 
squarrosa 382 
stricta 386 
tenuifolia 400 
thyrsoidea 385 
tortifolia 390 
uliginosa 387 
ulmifolia 391 
uniligulata 392 
Vaseyi 393 
virgata 386 
virgaurea 380 
“ alpina 385 
“ Deanei 388 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 595 


Solidago 
v. Gillmani 3 388 
“ monticola 388 
© Randii 388: 
“ Redfieldii 388. 
“ Wardii 382 
Sonchus 3 316. 
acuminatus 321 
arvensis 316 
asper 317 
floridanus 321 
ludovicianus 319 
oleraceus 316-7 
pulchellus 320 
spicatus 321 
Sophia 2 170 
(Sisymbrium in part) 
brachycarpa 171 
Hartwegiana 171 
incisa EE 
intermedia 171 
pinnata 171 
Sophia 170 
Sophora 2 342 
alba 344 
alopecuroides 342 
australis 344 
sericea 342 
tinctoria 345 
villosa 345 
Sophronanthe’= g 105 
hispida 195 
pilosa 196 
Sorbaria 
sorbifolia 2 247 
Sorbus 2 287 
americana 287 
aucuparia 287 
domestica 287 
microcarpa 287 
scopulina 287 
sitchensis 287 
subvestita 287 
sambucifolia 287 
Sorghastrum I II9 
avenaceum 119 
Linnaeanum 120 
nutans 120 
Sorghum I 120 
avenaceum 120 
Halepense 120 
Linnaeanum 120 
nutans 120 
SPARGANIACEAE I 69 
Sparganium I 69 
acaule 72 
americanum 70 
androcladum 70 
“ fluctuans 73 
angustifolium 73-4 
chlorocarpum 71 
diversifolium 72 
erectum 69, 71 
eurycarpum 70 
fluctuans 73 
hyperboreum 73 
lucidum 71 
minimum y 


multipedunculatum 


natans 

cas pd 
angustifolium 

 submuticum 


NNNN SJ 
OwWwrly 


simplex 
“ acaule 71 
* androcladum 70 
“ Nuttallit 70 
Sparganophorus 3 355 
verticillatus 355 
Spartina I 232 
alterniflora 224 
caespitosa 223 


596 


Spartina 
cynosuroides 
glabra 
gracilis 
juncea 
Michauxiana 
patens 
polystachya 
Schreberi 
stricta 

“ maritima 

Spartium 
scoparium 

Spathyema 
foetida 

Spatularia 
petiolaris 

Specularia 
biflora 
leptocarpa 
perfoliata 

Spergula 
arvensis 
decumbens 
nodosa 
saginoides 
sativa 

Spergularia 
media 
rubra 
salina 

Spermacoce 
diodina 
glabra 
tenuior 


SPERMATOPHYTA 


Spermolepis 
divaricatus 
echinatus 
patens 

Sphaeralcea 
acerifolia 
cuspidata 
remota 
stellata 

Sphaerocarpus 

Sphenopholis 
nitida 
obtusata 
pallens 

Spiesia 


see Oxytropis 


Spigelia 
Anthelmia 
marylandica 

Spilanthes 
americana 
repens 
urens 

Spiraea 
alba 
Aruncus 


I 223 


HN 


“ hermaphrodita 


betulifolia 
“ corymbosa 
callosa 


chamaedrifolia 


corymbosa 
japonica 
latifolia 
lobata 
opulifolia 
prunifolia 
rubra 
salicifolia 
“ lanceolata 
“ latifolia 
sorbifolia 
tomentosa 
stipulata 
trifoliata 
Ulmaria 


224 
224 
223 
222 
223 
223 
222 
224 
224 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Spiraea Staphylea 2 493 
virginiana 2 246 Bolanderi 493 
see Ulmaria 249 pinnata 493 

Spiranthes I 563 trifolia 493 
see Ibidium 564 | Statice 2 718 
Beckii 566 Armeria 718 
cernua 565 caroliniana 718 
decipiens 570 Limonium 718 
gracilis 565 | Steinchisma SH nls) 
graminea praecox 566 hians 163 
lucida 564 | Steironema 2712 
neglecta 565 ciliatum 5 fie} 
odorata 565 intermedium 713 
ovalis 565 lanceolatum 714 
parviflora 565 longifolium 714 
plantaginea 564 quadriflorum 714 
Romansofiana 464 radicans 713 
simplex 566 tonsa 713 
unalaskensis 555 Stellaria 2 41 
vernalis 565 see Alsine 2 41 

Spirodela I 446 biflora 55 
(Lemna in part) cerastioides 50 
polyrhiza 446 Friesiana 45 

Sporobolus — I 194 groenlandica 57 
(ine, Vilfa) Stenactis 
airoides 199 Beyrichii 3 440 
angustus 197 | Stenandrium 
5 ee es tuberosum 3 208 
asperifolius 200 Stenanthium See 
Eee Sir 196 angustifolium 490 
brevifolius 188 Sfanuneum agO 
canovirens 195 robustum 490 
clandestinus 195 | Stenophragma 2 176 
compressus 200 Thaliana 176 
confusus 197 | Stenophyllus I 319 
cryptandrus 199 (Fimbristylis in part) 
cuspidatus 188 capillaris 319 
Drummondii 196 | Stenosiphon 2 610 
ejuncidus 198 linifolium 610 
gracilis 198 virgatus 610 
heterolepis 199 | Stenotus 3 379 
indicus 197 (Aplopappus in part) 
junceus 198 acaulis 379 
longifolius 196 armerioides 379 
minor 194 | Stephanomeria 
neglectus 195 minor 3/322 
pilosus 196 runcinata 312 
Richardsonis 188 | Stevia 
Serotinus 198 callosa 507 
simplex 187 sphacelata 507 
texanus 200 | Stewartia 2 526 
Torreyanus 200 Malachodendron 
uniflorus 198 2 526 
vaginaeflorus 194 pentagyna 527 
virginicus 197 virginica 526 

STACHYEAE 3 99 | Stillingia 2 461 

Stachys 3 123 salicifolia 461 
ambigua 12 sylvatica 461 
annua 127 | Stipa I 176 
arenicola 125 avenacea Ly, 
arvensis 127 barbata L777 
aspera 126 bicolor 177 
atlantica 124 canadensis 176 
Betonica 128 capillaris 189 
cincinnatensis 126 comata 177 
cordata To juncea 176 
germanica 123,8 Macounii 176 
glabra 126 membranacea 173 
hyssopifolia 12 pennata 176 
hispida 126 Richardsonii 176 
latidens 125 spartea 176-7 
Nuttallii 126 virginica 177 
palustris 125 viridula 176 
““ aspera 126 | Stoepelina 3 366 
“ cordata 127 elegans 366 
salvioides 127 | Stomoisia 3 231 
tenuifolia 126 comata 232 

Stanleya 2 169 juncea 232 
pinnata 169 virgatula 232 
pinnatifida 169 | Streptopus I 520 

STAPHYLEACEAE 2 4931 amplexifolius 520 


Vot. III. 
Streptopus 
lanuginosus 1517, 
longipes 520 
oreopolus 520 
roseus 520 
Strophostyles 2 423 
(Phaseolus in part) 
angulosa 423 
helvola 423 
missouriensis 423 
pauciflora 424 
peduncularis 424 
umbellata 424 
Struthiopteris 
germanica puis 
Stuartia 
see Stewartia 2 526 
Sturmia Z S71 
paludosa eae 
Stylipus 2 271 
vernus 271 
Stylisma 3 40 
aquatica 41 
humistrata 41 
Pickeringii SAT 
Stylophorum 2 140 
diphyllum 140 
Stylosanthes 2 393 
biflora 393 
“hispidissima 393 
elatior 393 
procumbens 393 
racemosa 365 
riparia 304 
STYRACACEAE 2. 72h 
Styrax 2 722 
americana 723 
grandifolia 723 
officinalis 722 
pulverulenta 723 
Suaeda 2 23 
americana 23 
depressa 24 
maritima 24 
Richii 24 
Subularia 2 159 
aquatica 159° 
“Succisa 3 290 
australis 290 
Succisa 290 
Sullivantia 2 223 
H apemany 22 
ohionis 224 
Sullivantii 224 
Svida 2 661 
see Cornus 661 
interior 662 
Priceae 661 
Swertia 
difformis 36 
deflexa 15 
rotata 14 
SY MPETALAE 2 666 
Symphoricarpos 3 276 
occidentalis 277 
orbiculatus 277 
paucifiorus 276 
racemosus 276 
Symphoricarpos 277 
vulgaris 277. 
Symphytum 3 92 
asperimum 92 
officinale 92 
tuberosum 92 
Symplocarpus I 444 
foetidus 445 
SYMPLOCACEAE 2 721 
Symplocos _ 721 
martinicensis 721 
tinctoria 721 
Synandra 3 118 
grandiflora 118 


WoL LL. 


Synandra 
hispidula 3 118 
Syndesmon 2 102 
(Anemonella) 
thalictroides 102 
Syngonanthus§ 1 455 
flavidulus 455 
Synosma 3 537 
(Cacalia in part) 
suaveolens 538 
Syntheris 198 
Bullii 198 
Houghtoniana 198 
treniformis 198 
tubra 199 
Syntherisma I 121 
(Digitaria ; 


Panicum, in part) 
filiforme 


122 
fimbriatum 123 
humifusum 123 
Ischaemum 123 
linearis 123 
marginatum 123 
praecox TIE 3 
sanguinale 123 
serotinum 122 
villosum 122 
Syringa 2 724 
vulgaris 724 
Tabernaemontana 
Amisonia 3 26 
Taenidea 2 640 
integerrima 640 
Tagetes 
papposa 5) is} 
Talinum 2 35 
calycinum 37 
parviflorum 36 
rugospermum 36 
teretifolium 36 
Tanacetum 2) Gain 
huronense 522 
vulgare 521-2 
“ crispum 522 
Taraxacum 315 
seeLeontodon 315 
Dens-leonis 315 
erythrospermum 316 
latilobum 315 
officinale Bit 
Taraxacum 315 
“alpinum 315 
TAXACEAE 167 
Taxodium 163 
ascendens 64 
distichum 64 
imbricarium 64 
Taxus I 67 
baccata 67 
brevifolia 67 
canadensis 67 
minor 67 
Tecoma CMe 
radicans 237 
stans 237 
Tephrosia 
seeCracca 2 372-3 
Tetragonanthus 3 15 
(Halenia) 
deflexus 15 
Tetragonia 
expansa 2 34 
Tetragonotheca 3 468 
helianthoides 468 
Tetraneuris 3 508 
acaulis 509 
fastigiata 509 
herbacea 509 
linearifolia 508 
scaposa 509 


Tetraneuris 
simplex 3 500 
stenophyllus 509 
Tetranthera 
geniculata 2 135 
Teucrium 3 101 
boreale 103 
botrys 104 
canadense 102 
fruticans IOI 
laciniatum 103 
littorale 102 
occidentale 103 
Scorodonia 103 
virginicum 102 
Thalesia 2 233 
(Aphyllon) 
fasciculata 234 
“ lutea 234 
uniflora 233 
Thalia I 546 
dealbata 546 
geniculata 546 
Thalictrum 2 118 
alpinum 118 
anemonoides 102 
campestre 120 
caulophylloides 119 
clavatum 118 
confine 120 
coriaceum 119 
Cornuti 121 
dasycarpum 120 
dioicum 120 
“ coriaceum 119 
foetidum 118 
occidentale 120 
polygamum 121 
purpurascens 120 
“ ceriferum 119 
revolutum I19 
venulosum 120 
Thapsia 
trifoliata 2 639 
Thaspium 2 639 
atropurpureum 639 
aureum 639 
“ apertum 641 
barbinode 640 
“angustifolium 640 
pinnatifidum 640 
trifoliatum 639 
“ apterum 642 
Walteri 640 
THEACEAE 2 526 
Thelesperma 3 500 
ambiguum PSO 
filifolium 500 
gracile 501 
. intermedium 501 
trifidum 500 
Thelypodium 2 169 
integrifolium 169 
pinnatifidum 178 
Thermia 
rhombifolia 2 344 
Thermopsis 2 343 
lanceolata 343 
mollis, 343 
rhombifolia 344 
Therofon 2 223 
(Boykinia) 
aconitifolium 223 
napelloides 223 
Thesium 
umbellatum I 640 
Thlaspi 2 167 
arvense 167-8 
bursa-pastoris 158 
campestre 164 
perfoliatum 168 
Thuja I 64 


Thuja 
occidentalis 
THY MELEACEAE 
Thymophylla 
aurea 
setifolia 
Thymus 
Acinos 
Serpyllum 
vulgaris 
Tiarella 
biternata 
cordifolia 
Tiedemannia 
filiformis 
rigida 
teretifolia 
TILIACEAE 
Tilia 
alba 
‘americana 
canadensis 
europaea 
glabra 
heterophylla 
leptophylla 
Michauxii 
pubescens 
Tillaea 
aquatica 
simplex 
Tillaeastrum 
aquaticum 
Vaillantii 
Tillandsia 
usneoides 
Tiniaria 
cilinodis 
Convolvulus 
cristata 
dumetorum 
scandens 
Tipularia 
discolor 
unifolia 
Tissa 
(Spergularia) 
canadensis 
marina 
rubra 
salina 
Tithymalopsis 
arundelana 
corollata 
Ipecacuanhae 
marylandica 
Tithymalus 
arkansanus 
commutatus 
Cyparissias 
Darlingtonii 
Esula 
falcatus 
Helioscopia 
Lathyrus 
lucidus 
missouriensis 
obtusatus 
Peplus 
platyphyllus 
robustus 
Tofieldia 
glutinosa 
palustris 
racemosa 
Tordylium 
Anthriscus 
nodosum 
Torilis 
Anthriscus 
nodosa 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


I 65 
2 574 
Si SUS) 

514 

513 
3 141 

139 


597 
Tomentilla 
reptans 2 250 
Torreya 
grandiflora 3.118 
Touterea 2 567 
stricta 567 
Tovara I 664 
virginiana 665 
Townsendia 3 402 
exscapa 403 
grandiflora 402 
intermedia 403; 
sericea 402-3 
Toxicodendron 2 483 
crenatum 482 
pinnatum 483 
pubescens 484 
tadicans 484 
Rydbergii 484 
Toxicodendron 484 
vernix 483 
vulgare _. 484 
Toxicoscordion 1 4o1 
gramineum 492 
Nuttallii 492 
Toxylon I 63L 
(Maclura) 
pomiferum 632 
Tracaulon I 672 
arifolium 673 
sagittatum 673 


Trachelospermum 3 23 
(Echistes in part) 
difforme 


23 

jasminoides 23 
Trachynotia I 223 
cynosuroides 223 
polystachya 223 
Tradescantia I 460 
bracteata 460 
brevicaulis 460 

“ villosa 460 
flexuosa 462 
montana 462 
occidentalis 461 
pilosa 462 
reflexa 461 
rosea 459 
virginiana 460-1 
Tragia 2 458 
cordata 460 
innocua 459 
macrocarpa 460 
nepetaefolia 459 
ramosa 459 
stylaris 459 
urens 459 
urticaefolia 459 
volubilis 458 
Tragopogon 3530s 
Dandelion 308 
porrifolius 313 
pratensis 313 
virginicum 307-8 
Tragus I 121 
TRAPACEAE 2 61I 
Trapa 611 
natans 612 
Trautvetteria 2 103 
applanata 103 
carolinensis 103 
palmata 103 
Triadenum 2 537 
longifolium 537 
petiolatum 537 
virginicum 537 
Triantha I 486 
glutinosa 486 
racemosa 487 
Tribulus 2 442 
terrestris 442 


| Trichelostylis I 


598 


Trichelostylis 
geminata I-322 
‘Trichomanes ris 
Boschianum 8 
crispum 8 
radicans 8 
Trichophyllum 
oppositifolium 3 508 


‘Trichostema 3 104° 
brachiatum 104 
dichotomum 104-5 
lineare 105 

Tricuspis T2323 
elongata 234 
purpurea 235 
sesslerioides 233 
stricta 234 

Tridens I 233 
elongata 234 
flava 233 
stricta 234 

‘Trientalis 2 715 
americana 715 

__ europaea 715 

Trifolium 2 353 
agrarium 354 
arvense 355 
aureum 354 
Beckwithii 356 
biflorum 393 
carolinianum 358 
dubium 354 
erectum 422 
hybridum 357 
incarnatum 355 
medium 356 
Melilotus 352-3 
MUNUS 354 
pratense 353,5 
procumbens 354 
reflexum 357 
Tepens 358 
simplicifolium 422 
stoloniferum 357 
virginicum 356 

Triglochin I oI 
elata 92 
maritima 92 
palustris gI-2 
striata 92 
triandra 92 

Trigonella 2 359 
americana 359 

Trilisa 3 369 
(Liatris) 
odoratissima 369 
paniculata 369 

TRILLIACEAE ER 

Trillium I 523 
cernuum 523, 6 
declinatum 525 
erectum 525 
erythrocarpum 526 
grandiflorum 525 
nivale 524 
recurvatum 524 
rhomboideum 

“ grandiflorum 525 
sessile ines 
undulatum 526 
viride 524 
viridescens 524 

Triodia 

albescens I 238 
decumbens 233 

Triosteum 3 274 
angustifolium 275 
aurantiacum 275 
perfoliatum 275 

Triplasis I 235 
americana 235 
purpurea 235 


Triphora I 561 
pendula 562 
trianthophora 562 

Tripolium 
angustum 3 434 

Tripsacum L 00 
cylindricum 112 
dactyloides III 
monostachyum III 

Tripterella I 547 
coerulea T1547, 

Tripteridium 2532 
micranthum 33 

Tripterocalyx 2 33 
micranthus We 
macranthus BR 

Trisetum I 216 
flavescens oy 
palustre 217 
pennsylvanicum 217 
pratense 27, 
purpurascens 219 
spicatum Pty) 
subspicatum ai, 

Triticum 
see Agropyron 

I 283-5 

Trollius 2 86 
americanus 87 
europaeus 86 
laxus 87 
“ albiflorus 87 

Troximon B6e23 
see Agoseris 3 323 
cuspidatum 324 
glaucum B23 
marginatum 324 
parviflorum 323 

Tsuga I 62 
canadensis 62 
caroliniana 62 
Sieboldi 62 

Tulipa nt OE 
sylvestris 505 

Tullia 3 144 


pycnanthemoides 144 


Tunica 20713 
Saxifraga 72 
Turritis 
see Arabis 2 178 
stricta 182 
Tussilago 3 530 
Farfara 531 
palmata 531 
Petasites 532 
sagittata 532 
TYPHACEAE I 68 
Typha I 68 
angustifolia 69 
latifolia 68 
Udora 
verticillata I 106 
Ulex 2 349 
europaeus 349 
ULMACEAE I 625 
Ulmaria 2 248 
(Spiraea in part) 
palustris 249 
rubra 248 
Ulmaria 249 
Ulmus I 626 
alata 627 
americana 626 
campestris 626 
fulva 627 
pubescens 627 
racemosa 626 
serotina 627 
Thomasi 626 


UMBELLIFERAE 2 620 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Unania 
“ alba 3 432 
Uncinia I 389 
muicroglochin 389 
Unifolium I 516 
(Maianthemum) 
(Smilacina in part) 
canadense 517 
Uniola I 248 
gracilis 248 
laxa 248 
latifolia 249 
paniculata 248-9 
spicata . 250 
stricta 250 
Urachne I 174 
asperifolia 174-5 
micrantha 17 
racemosa 175 
Uralepis 
elongata I 234 
pilosa 235 
URTICACEAE I 634 
Urtica 634 
canadensis 636 
capitata 637 
chamaedryoides 636 
cylindrica 637 
divaricata 636 
dioica 634, 5 
gracilis 635 
Lyallii 635 
pumila 637 
urens 635 
Urticastrum I 636 
(Laportea) 
divaricatum 636 
Utricularia 3 226 
biflora 227 
ceratophylla 230 
clandestina 229 
cleistogama 231 
cornuta 231, 2 
fibrosa 228 
fornicata 227 
geminiscapa 229 
gibba 227 
Greenei 230 
inflata 230 
“ minor 230 
intermedia 228 
juncea 232 
longirostris 227 
macrorhiza 229 
macrorhyncha 227 
minor 228 
personata 232 
pumila 227 
purpurea 226 
radiata 229 
resupinata 230 
saceata 226 
setacea 231 
simplex BED 
striata 228 
subulata 231 
virgatula 232 
vulgaris 227 
“americana, 229 
Uva-Ursi 2 693 
Uva-Ursi 693 
Uvularia I 518 
amplexifolia 520 
grandiflora 519 
nitida 519 
perfoliata 518 
puberula 519 
sessilifolia 519 
Vaccaria 2 73 
(Saponaria in part) 
Vaccaria 73 


Vor. ITT, 

Vaccaria 
vulgaris 2 73 
VACCINIACEAE 2 694 
Vaccinium 2 698 
album 3 281 
amoenum 2 700 
angustifolium 7O1 
arboreum 698 
atrococcum 702 
brachycerum 696 
buxifolium 6096 
caesariense 701 
caespitosum 699 
canadense 701 
Constablei 700 
corymbosum 700 
“ atrococcum 702 
disomorphum 702 
Dobbin 701 
dumosum 696 
Elliottii 703 
erythrocarpon 702 
frondosum 695 
hirtellum 696 
hispidulum 704 
ligustrinum 691 
macrocarpon 705 
membranaceum 699 
mucronatum 490 
myrtilloides 699 
Myrtillus 2 608 
myrsinites 703 
nigrum 702 
nubigenum 700 
ovalifolium 700 
Oxycoccus 704 
“intermedium 704 
pallidum 700 
pennsylvanicum 701 
“angustifolium 701 
“ nigrum 702 
resinosum 696 
simulatum 700 
stamineum 607 
tenellum 703 
uliginosum 699 
vacillans 702 
virgatum 793 
Vitis-Idaea 6907 
“© minus 607 


seeGaylussacia 605-6 
Oxycoccus 704-5 


Vagnera 5 tei ely- 
(ine. Smilacina 
in part) 
amplexicaulis 515 
racemosa 515 
stellata 516 
trifolia 516 
VALERIANACEAE 3 284 
Valeriana 3 284 
dioica 285 
edulis 285 
Locusta 286 
“ olitoria 286 
“ radiata 287 
officinalis 286 
pauciflora 284 
pyrenaica 284 
septentrionalis 285 
sylvatica 285 
uliginosa " e2zas 
Valerianella 3 286 
chenopodifolia 287 
Locusta 286 
longiflora 288 
olitoria 286 
radiata 287 
stenocarpa 287 
Woodsiana 288 


VALLISNERIACEAE I 104 
Vallisneria I 106 


Vor, III. 
Vallisneria 
spiralis I 106 
Vaseya I 184 
comata 187 
Veratrum I 494 
album 494 
angustifolium 490 
luteum 489 
parviflorum 495 
viride 494 
Woodii 495 
Verbascum 3.173 
Blattaria 174 
Lychnitis 174 
phlomoides 174 
Thapsus 173 
virgatum 174 
VERBENACEAE 3 94 
Verbena 3 94 
ambrosiaefolia 97 
angustifolia 96 
Aubletia 97 
bipinnatifida 97 
bracteosa 95,6 
canadensis 96, 7 
carolinensis 95 
Drummondii 97 
hastata 95 
nodiflora 98 
officinalis 94,5 
paniculata 95 
pinnatifida 95 
rigens 96 
riparia 95 
stricta 95, 6 
urticifolia 95 
Verbesina 3 467 
alba 468 
alternifolia 487 
encelioides 489 
helianthoides 488 
occidentalis 488 
Siegesbeckia 488 
virginica 487 
VERNONIEAE 3 347 
Vernonia 3 350 
altissima 352 
’ “ grandiflora 353 
“ marginata Boe 
arkansana 351 
Baldwinii 353 
corymbosa 352 
crinita 351 
Drummondii 353 
fasciculata 352 
glauca 351 
illinoensis Bog 
interior 353 
Jamesii 352 
marginata 352 
maxima 352 
missurica 353 
noveboracensis 351 
“latifolia 351 
“ tomentosa 351 
Veronica 3 199 
agrestis 203 
alpina 201 
americana 200 
Anagallis-aquatica 
200 
arvensis 202 
Beccabunga 200 
Buxbaumii 203 
byzantina 203 
Chamaedrys 201 
hederaefolia 203 
humifusa 202 
longifolia 201 
officinalis 199, 201 
peregrina 202 
scutellata 200 


Veronica 
serpyllifolia 3 202 
Teucrium 201 
Tournefortii 203 
virginica 204 
Wormskioldii 201 
Vesicaria 2 155 
argentea 155 
arctica 156 
didymocarpa 156 
globosa 155 
gracilis 155 
Shortii 155 
Vesiculina 3 226 
purpurea 226 
saccata 226 
Viburnum 3 269 
acerifolium 270 
alnifolium 269 
americanum 270 
cassinoides 272 
Demetrionis 272 
ferrugineum 274 
dentatum 271 
“ pubescens 271 
Lantana 274 
lantanoides 269 
Lentago 273 
molle 272 
nudum 273 
“ Claytoni 273 
obovatum 274 
Opulus 270 
-“ eradiatum 270 
pauciflorum 270 
prunifolium 273 
“ferrugineum 274 
* globosum 273 
pubescens 27 
rufidulum 274 
rufotomentosum 274 
scabrellum 271 
semitomentosum 271 
Tinus 269 
trilobum 270 
venosum 272 
“ Canbyi 272 
Vicia 2 408 
americana 409 
angustifolia 412 
caroliniana 410 
Cracca 409 
hirsuta All 
linearis 409 
ludoviciana 410 
micrantha 410 
Mitchelli 4Il 
narbonensis 412 
oregana 409 
parviflora 410 
sativa 408, 11 
“ angustifolia 412 
Sepium 412 
sparsifolia 409 
tetrasperma 411 
truncata 409 
villosa 409 
Vigna 2 42 
Catjang 425 
repens 425 
sinensis 425 
Vilfa 
see Sporobolus 
I 194-200 
Villarsia 
aquatica 3 19 
lacunosa 18 
Vinca 3 20 
major 20 
‘minor | 20 
Vincetoxicum 3 37 


(Gonolobus) 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


Vincetoxicum 
Baldwinianum 3 39 
carolinense 38 
gonocarpos 37 
hirsutum 38 
nigrum 36 
obliquum 38 
Shortii 39 
suberosum 37, 

VIOLACEAE 2.545 

Viola 2 545 
adunca 561 
affinis 551 
amoena 556 
arenaria 561 
arvensis 563 
atlantica 554 
blanda 555-0 


“ palustriformis 556 


“ renifolia 55s 
Brittoniana 553-4 
canadensis 560 
Calceolaria 564 
canina 562 
“ adunca 561 
“ puberula 561 
chinensis 558 
concolor 564 
congener 548 
conjugens 552 
conspersa 560 
cucullata 552 
cuspidata 549 
delphinifolia 554 
dentata 552 
domestica 548 
Egglestonei 547 
emarginata 2e553 
eriocarpa 559 
falcata 548 
fimbriatula 552 
hastata 558 
hirsuta 558 
hirsutula 550 
incognita 556 
“ Forbesii 556 
labradorica 561 
lanceolata 557 
lancifolia 558 
latiuscula 549 
longipes 561 
missouriensis 549 
Muhlenbergii 560 
“ minor 561 
multicaulis 562 
nephrophylla 551 
novae-angliae 550 
Nuttallii 558 
obliqua 548 
odorata 2558 
ovata 552 
pallens 556 
palmata 547 
palustris 555 
papilionacea 548 
pectinata 553 
pedata 547 
“ bicolor 547 
“ lineariloba 547 
pedatifida 554 
pratincola 548 
primulifolia ay, 
pubescens 559 
punctata 561 
Rafinesquii 562 
renifolia 555 
retusa 55! 
rostrata 562 
rotundifolia 556-7 
rugulosa 559 
Rydbergii 559 
Sagittata 553 


599 


Viola 


Ss. em L 
*: argiata 2 553 
ovata 


52 
Selkirkii Ete 
septemloba ; 548, 54 
Septentrionalis 550 
sororia 549 
Stoneana 548 
striata 2 560 
subvestita 561 
tenella 562 
tricolor 563 
triloba 548 
“ dilatata 548 
vagula 551 
venustula 551 
verticillata 564 
viarum 552 
villosa 550 
vittata 557. 
Walteri 562 
Viorna 2 122 
Addisonii 123 
crispa 123 
cylindrica 123 
flaccida 124 
Fremontii 125 
glaucophylla 124 
ochroleuca 125 
ovata 125 
Pitcheri 123 
Ridgwayi 124 
Scottii 126 
Simsti 123 
urnigera 122 
versicolor 124 
Viorna 124 
Virgilia 
lutea 2 343 
Viscaria 2 62 
alpina 2 
vulgaris 2 
Viscum 2 712 
flavescens I 639 
terrestre 2 712 
VITACEAE 2 505 
Vitis 2 505 
aestivalis 506 
“ canescens 507 
“ bicolor 507 
“ cinerea 507 
arborea 510 
Baileyana 508 
bicolor 507 
bipinnata 510 
cinerea 507 
cordifolia 508 
“riparia 507 
incisa 510 
indivisa 509 
Labrusca 506 
palmata 508 
quinquefolia 511 
riparia 507 
rotundifolia 509 
rubra 508 
rupestris 508 
vinifera 505 
virginiana 508 
vulpina 507 
Vitis-Idea 2 607 
Vitis-Idea 607 
Vieckia 
see Agastache 3 111 
anisata 112 
Volvulus 3 45 
spithamaeus 47 
Waldsteinia 2 268 
Doniana 269 
fragarioides 2690 
geoides 268 


600 


Waldsteinia 
parviflora 
Washingtonia 

(Osmorrhiza) 
Claytoni 
divaricata 
longistylis 
obtusa 
Weigela 
Willughbaea 
(Mikania) 
scandens 
Windsoria 
stricta 
pallida 
Wisteria 
frutescens 
macrostachys 
speciosa 
see Kraunhia 
Withania 
sordida 
Wolffia 
brasiliana 
columbiana 
floridana 
gladiata 
papulifera 
punctata 
W olffiella 
floridana 
gladiata 
oblonga 
Woodsia 
alpina 
Cathcartiana 
glabella 
hyperborea 
ilvensis 
obtusa 
oregana 
scopulina 


GENERAL INDEX OF LATIN GENERA AND SPECIES. 


2 269 
2 627 


627 
628 
628 
628 
3 283 
3 362 


W oodwardia 
angustifolia ries 
areolata 25 
virginica 24 

Wulfenia 
Houghtoniana 3 198 
rubra 199 

Xanthium 343 
americanum 346 
canadense 346 
commune 346 
echinatum 345 
fruticosum 342 
glabratum 346 
glanduliferum 345 
inflexum 345 
Macounii 345 
macrocarpum 
“ glabratum 346 
maculatum 345 
orientale 346 
oviforme 345 
pennsylvanicum 346 
speciosum 344 
spinosum 344 
strumarium 343, 6 

Xanthorrhiza 2 89 
apiifolia 89 
simplicissima 89 

Xanthoxalis 2 432-5 
Brittoniae 434 
Bushii 433 
corniculata 431-2 
cymosa 435 
filipes 433 
grandis 435 
interior 434 
Priceae 434 
recurva 434 
rufa 433 
stricta 432 


Xanthoxrylum 2 443 
Xerophyllum I 487 
asphodeloides 488 
setifolium 488 
Ximenesia 3 488 
encelioides 489 
Xolisma 2 6901 
(Andromeda in 
part 
foliosiflora 691 
ligustrina 691 
Xylosteum 3 282 
involucratum 282 
oblongifolium 281 
XYRIDACEAE I 450 
Xyris I 450 
arenicola 453 
caroliniana 451 
communis 452 
Congdoni 452 
difformis 452 
elata 452 
fimbriata 453 
flexuosa 451 
Jupacai 451 
macrocephala 452 
montana 451 
pusilla 451 
torta 451-2 
Yucca 58 ait 
aloifolia 512 
arkansana 512 
angustifolia Ruz 
baccata 512 
filamentosa Br 
glauca 512 
ZANICHELLIACEAE I 74 
Zanichellia 1 88 
intermedia 89 
palustris 89 


Vor. III. 
Zanthoxylum 2 443 
americanum 444 
carolinianum 444 
Clava-Herculis 444 
Zapania 
cuneifolia 3 98 
Zephyranthes 
Atamasco ad 5 
Zinnia 3 466 
grandiflora 466 
Zizania I 168 
. aquatica 168 
miliacea 167 
palustris 168 
Zizaniopsis I 167 
miliacea 167 
Zizia 2 641 
aurea 641 
Bebbii 641 
cordata 642 
integerrima 640 
pinnatifida 640 
Zornia 2 304 
bracteata 394 
tetraphylla 394 
ZOSTERACEAE I 90 
Zostera QI 
marina gi 
ZOYSIEAE I 107-8 
Zygadenus I 490 
elegans 491 
chloranthus 491 
glaberrimus 491 
glaucus 491 
gramineus 492 
helmanthoides 493 
intermedius 491 
Nuttallii 492 
VENeNOSUS 492 
ZYGOPHYLLACEAE 2 442 
Zygophyllidium 2 468 
hexagonum 468 


ENGLISH INDEX, INCLUDING POPULAR 
PLANT NAMES. 


[The heavy face figures 1, 2, 3 indicate the volume; those following them, the page. 


popular names, not printed in the text, are referred to the proper plant by 
the number of the Illustration or Figure (fig.) in the Index.] 


Aaron’s Beard 3 175 
Aaron’s Rod 
2 207; 3 173, 388 
Abele L597; 
Abel-tree er 
Abronia 33 
Abscess-root, Am. 3 65 
Absinth Bas25 
Acacia 2 330 
Bastard, False 375 
Illinois 332 
Prairie 331 
Rose 375 
Three-thorned 339 
ACANTHUS FAMILY 
3 239 
Ache 2 660 
Aconite 96-7 
Winter 88 
Aconite Saxifrage 223 
Actinomeris 3 487 
Adam-and-Eve 1 574 
Adam’s-cup 2 201 
-flannel 3 173, 289 
-needle 
eee a2 O27 
-rod 20173 
Adder’s-fern Tez, 36 
-flower 2 69 
-meat 44 
-mouth a yf 
-Pogonia 560 
Adder’s-spear ra 
Adder’s-spit 32 
ADDER’S TONGUE 
FAMILY 1 2 
Adder’s-tongue 
I 2,506; 3 332 
Engelmann’s r2 
Midland 507 
Minnesota 507 
Sand 2 
White 506 
Yellow 506 


Adder’s-violet 3 


Adders-wort 94 
Adonis-flower 2 121 
African-rose 137 
Agave 534 
Agrimony 2 266-8 
Britton’s 268 
Water 3 495 
-bark 2 445 
Ague-grass oe iit 
-root Sir 
-tree 2 134 
-weed 3 10, 361 
AILANTHUS FAMILY 
2 445 
Ailanthus 445-6 


Airif 3 259 
Aise-weed 2 654 
Aiten I 66 
Alams 590 
Albany-hemp 636 
-beech-drops 2 673 
Alder 612-4 
American 613 
Berry 2 504 
Black 
I 613; 2 489, 504 
Common I 613 
Dwarf 2 503 
European I 613 
False 2 489 
Green ZT 67253 
Hazel TE 6ms 
Hoary 613 
Mountain 612 
Seaside 614 
Smooth 613 
Speckled 613 
Spiked 2 667 
Spotted 235 
Striped 489 
Tag r O10 4 
White 2 480, 667 
Witch 2 233, 691 
Alecoast, Aleost 3 519 
Alehoof 114 
Alexanders 2 641 
Golden 640-1 
Heart-leaved 641 
Purple 639 
Alfalfa Bei 
Alfilaria 430 
Alfilarilla 430 
Alkali-grass I 250 
Alkanet 3 87 
Bastard 87 
Alkekengi 162 
All-bone 2 44 
Alleghany Fringe 143 
Aloe 32 
Alleghany Vine 143 
Alleluia 350 
Allgood 14 
All-heal 3 125, 286 


Alligator-tree 2 235 


-bonnets 79 
-pear 133 
Allison, Sweet 153 
White 180 
Allocarya 8°79 
Allseed 2 440 
Allspice, Carolina 132 
Wild 135 
Almond, cutting 3 465 
Earth 304 
Aloe, False 534 


Aloe-root, 511 
Alpine Bistort 671 
Alsike 2 356 
Althaea,Shrubby 526 
Alum-bloom 426 
Alum-root 226, 426 
Common 226 
Alyssum 153 
Hoary 153 
Small 154 
Sweet 153 
Yellow 154 
AMARANTH FAMILY 2 I 
Amaranth 2 
Coast 5 
Crisp-leaved 5 
Dwar 5 
Globe 8 
Green 2 f 1660 
Low 4 
Palmer’s 2 
Prostrate 3 
Purplish 4 
Red 3 
Spiny 3 
Spleen 2 
Thorny 3 
Torrey’s 4 
AMARYLLIS FAMILY 153 
Amber 2-533 
Ambrose 14 
Ambrosia, Tall 3 341 
Ammania 2575 
Ampelopsis 509-10 
Amy-root 22, 81 
Anchusa, Amer. f 3540 
Andromeda 689 
Lyon’s 691 
Privet 6901 
Anemone, Canada 2 97 
Carolina 98 
Cut-leaved 98 
False-rue 84, 89 
Long-fruited 98 
Meadow-rue 102 
Mountain 100 
Northern 98 
Prairie 102 
Richardson's 100 
Round-leaved 99 
-headed 99 

Rue 102 
Small-flowered 98 
Star 2715 
Tall 99 
Wind-flower 162 
Wood 100 
Angel-eyes 3 251 
Angelica 2 635-7 
Hairy 637 


A few 

Angelica 
High 2 637 
Sea-coast 646 
Afigelica-tree 2 647 
Angelico 647 
Angel’s-eyes 251 
Angleberries 415 
Angle-pod 373 
Anise, Sweet 2 627 
Anise-root 628 

Appalachian Tea 

48; 3 372 
APPLE FAMILY 2 280 
Apple 289 
Buffalo 377 
Crab 289 

Devil’s 
2 94,130; 3 169 
Earth 3 486 
Hog 2 180 
Honeysuckle 678 
Indian 130 
Love 3 167 
Mad 169 
May 2 130 
Mock 33292 
Osage I 632 
Peru 3 169 
Prairie 2 363 
Swamp 678 
Thorn 3 169 
White 2 418 
Wild Balsam 3 292 
Apple of Peru 3 155 
Apple of Sodom 165 
Apple-pie(smell) 2 591 
Apple-root 2 469 
April-fools 2 102 
Arbor-vitae I 65 
Arbutus 2 258 
Trailing 2 692 
Archangel 2 637 
Black 3 123 
Green 146-8 
Red or Sweet 121 
White 122 
Arctagrostis I 201 
Arethusa 562 
Argentina 2 258 
Argentill 2 264 
Arnica 3 315, 533-4 
Arnica-bud 3 310 
Aristida I 179-81 
Plains 182 
Arrow arum I 444 
Arrow-bean 2 492 


ARROW-GRASS FAMILY 

I or 
Arrow-grass I 92 
Arrow-head 98-102, 182 


602 


Arrow-leaf 2 98, 100 
ARROW-ROOT FAMILY 
I 546 
Arrow-wood 
2 504,664; 3 270-2 


Indian 2 441, 664 
AruM FAMILy I. 443 
Arum I 444 

Floating 445 

Poison 444 

Water 444 
Arsenic, Wild 2 672 
Artichoke, Jerusalem 

3 4 
Asarabacca I 642-3 
Asarum 642 
Ash 2 725-8 

Basket 727 

Biltmore 725 

Bitter 2 491 

Black 2 4098, 726-7 

Blue 2 726-7 

Brown 727 

Cane 2 725 

Carolina 727 

Darlington’s 725 

Flowering 728 

Green 2 726 

Hoop tI 629; 2 727 

Maple 2 498 

Michaux’s 726 

Mountain 2 287 

Ox 719 

Poison 2 483, 728 

Pop, or Poppy 72 

Prickly 2 444 

Pumpkin 726 

Red 726 

Rim I 629 

River 2 726 

Sea 2 444 

Southern Prickly 444 

Swamp 726 

Water. 2 445 

Water 2 498, 726-7 

White 2 725,8 

Yellow 2 343,444 
Ash-weed 2 654 

-wort 3 543 
Asp, Mountain I 590 

Quaking 590 

White f 1440 
Asparagus I 514 
Aspen, American 

587, 590 

Great, White 587 

Large-toothed 589 
Asphodel I 486-7 

Bog 487 

False 486-7 

Lancashire 487 

Scottish I 486 
Ass-foot ah ele 
Aster 3 406-35 

Amethyst 418 

Aromatic 417 

Arrow-leaved 416 

Bee-weed 415 

Bernhardi’s 3 410 

Blue-Devil 415 

Blue Wood 3 414 

Bog 432 

Broad-leaved 442 

Bushy 428 

Calico 426 

Clayton’s 410 

Cornel 3 444 

Creeping 424 

Crimson-disk 409 

Crooked-stem 419 

Dense-flowered 427 

Dewy-leaf 412 


Aster 


Dome-topped 3 411 


Drummond’s 415 
Early Purple 3 419 
Eastern Silvery 423 
Fall 415 
Faxon’s 429 
Fendler’s 423 
File-blade 425 
Flat-topped 3 443 
Forking 410 
Frost-weed 430 
Golden B73 
grass leaved 373 
Great Northern 418 
Hardy 417 
_Hairy-stemmed 426 
Hervey’s 42 
Heath, Rose 434 
White 430 
Large-flowered 3 424 
-bracted 422 
-leaved 4il 
Late Purple 416 
Leafy-bracted 422 
Lilac-flowered 423 
Lindley’s 415 
Long-leaved 422 
Lowrie’s 415 
Low Rough 425 
Low Showy 3 424 
Many-rayed 413 
Missouri 429 
Mountain 432 
Nail-rod 2 
Nebraska 418 
New England 417 
New York 421 
Northeastern 3 419 
Panicled 428 
Prairie 426 
Price’s 430 
Pringle’s 430 
Purple 419 
Early 3 419 
Late 416 
Seaside 424 
Thin-leaved 417 
Purple-stem 419 
Rand’s 429 
Rayless 434 
Red-stalk 419 
Rice-button 3 428 
Rose Heath 434 
Rosemary 426 
Rough 406, 25 
Rush 421 
Salt-marsh, Annual 
433 
Perennial 433 
Savory-leaved 444 
Schreber’s 411 
Seaside, Purple 3 424 
Serpentine 3431 
Short’s A414 
Showy, Low 424 
Silkseed 3 406 
Silky,Western 423 
Silver-grass 373 
Silvery, Western 
3 423 
Eastern 423 
Sky-blue 414 
Slender 425 
Slim 433 
Small-headed 3 431 
Small White 427 
Smooth 420 
Narrow-leaved 420 
Southern 420 
Starved 426 
Stately 413 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Aster 
Stiff 3 444 
Swamp, Southern 431 
Tall White 3428 
Tansy 435 
Thin-leaved Purple 


417 
Tradescant’s 429 
Tuber 425 


Various-leaved 413,6 


Violet-leaf 3 412 
Violet Wood 412 
Viscid 435 
Wavy-leaved 416 
Western 421 
White 
Flat-top 3 443 
Small 427 
Tall 428 
Upland 432 
-Devil 3 426 
White Heath 430 
White Prairie 427 


White-topped 405-6 


White Wood 3 404 
White Wreath 427 
Whorled 432 
Willow 428 
Wood 409 
Common Blue 414 
Long-leaved 409 
Violet 412 
White 409 
Asterworts 3.347 
Asthma-weed 303 
Astilbe 2 215 
Atamasco Lily I 532 
Atriplex 2 17-19 
Aunt Jericho 3 637 


Autumnal Hawkbit 


3 310 

Avens 2 270-2 
Cream-colored 2 270 
Drooping 272 
Drummond’s Mt. 274 
Early Water 271 
Entire-leaved 274 


Large-leaved 2 


Long-plumed 273 
Mountain 274 
Purple 2 272 
Rough 270 
Spring 270 
Water 272 
White 270 
White Mt. 273 
Yellow 2 271 


Yellow Mountain 
Awlwort, Water 2 


Axeseed 392 
Axewort 392 
Axweed 654 
Axyris 2 20 
Aye-green 2 211 
Azalea 2 683 
Alpine 683 
Clammy 679 
Flame 678 
Hoary 2 678 
Mountain 678 
Pink, Purple, 678 
Smooth 679 
Trailing 683 
Tree, White, 679 
Yellow 678 
Azolla 1 38 


Babies’-breath 
TSO 272651203 
-feet, -toes, -slip- 
pers 2 452 
Baccharis 3 445-6 


Vo. III. 


Bachelor’s-buttons 
269, 111; 3 305 


wild 2447; 3557 
-breeches 2 1A2 
Backache-brake 1 308 
-root Weil 77 
Back-wort 92 
Bacon-weed 2) 5G 
Badger 102 
Baked-apple, -berry, 
2 276 
Bahia 3 508 
Bairnwort 402 
Bald Cypress I 64 
Baldmoney 39 
Bald Rush I 347 
Balders 3.516 
Balewort 2 131 
Ball Mustard - 2 159 
Ballogan 3 306 
Balloon Vine 2 150% 
Balm, Basil 2 iaeg 
Bee, Amer. 1307 
Calamint 139 
Field 137 
Fragrant 131 
Garden 137 
Lemon 137 
Low 131 
Moldavian 115 
Ox, Horse- 153 
e 131 
Balm-leaf 2137 
Balm-mint 3 137,151 
Balm-of-Gilead 1 588-9 
-Fir I 63 
Balmony 3 181 
Balsam 
I 63; 2 440-1, 474 
Canada tog 
He 61 
Mountain 63 
Old-field 3 454 
She- I 63 
Sweet, White, 3 454 
Wild 2 440-1 
Balsam Apple 3 292 
Balsam Fir, Fraser’s 
I 63 
Balsam-flowers 2 353 
Balsam-Poplar 1 588 
-weed 3 454 
Bamboo-Vine I 530 
Redberry 530 
Bamboo Brier I 528 
Banana, False 2 83 
Baneberry 2 90 
Bank-cress 195 
Bannal 2 350 
Banwort 3 402 
Barbeau 557 
Barbary-fig 21575 
BARBERRY FAMILY 2 126 
Barberry 2 127 
Holly-leaf 2 128 
Bardane 3 547 
Bardock BA 
Bergman’s cabbage 
2 193 
Barley 1 286-7 
Mouse, Way, Wild 
287 
Bartonia 2 6n7 
Bartsia, Alpine 3 219 
Red 219 
Base Vervain 201 
Basil 3 137-42 
cow 2 73 
field, stone 3 137-8 


wild 3 137-8, 144-6 
Basil-balm 3 132,06 
-thyme 3 138-9 


VoL. III. ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 603 
Basil-weed 3 137 | Beard-grass Belia, High 3 301 | Bindweed 
Bass, Bast, 6. AGI Smooth 1 184 | Belene 3 168 Knot I 674 
Bass-wood 1 83, 511-3 White-flowered 184] Bellbind 46| Low 3 47 
white 2 512] Bearded-joint I 150 | Bellbine 47 Short-stalked 46 
yellow 512 | Bear’s-bilberry 2 693 | BELL FLrowrer FAMILY Small 47 
Bastard-Alkanet 3 87 | Bear’s-foot 2 264 3 203 Trailing 46 
-Boneset 3 35,39 yellow 3 458 | Bellflower 3 204-7 Upright 47 
-Cress 2 164 -grape 2 693 American Red 3 42] Brrcw FAMILY I 605- 13 
-Elm I 629 -moss 1513; f 1279 Arctic 295 | Birch I 605 
-Gentian 39 -pawroot 1210 Bedstraw 296 Alpine 612 
-Hellebore I 563 -thread T5153 Clustered 296 American I 608 
-Hemp 3 119] Beaumont-root 3 204 Creeping 295 Black 609-10 
-Horehound 123 | Beaver-lily 2 78 Clasping 298 Blue 608 
-Jasmine 3 168 -poison 2 658 European 295 Bolean 609 
-Pennyroyal 3 105 -root 2 78 Marsh 295-6 Bog 2 503 
-Pimpernel 2 717 -tree 82 Nettle-leaved 3 295 Broom 608 
-Rocket 2 192 -wood I 629 Panicled 297 Canoe I 609 
-Toad-flax I 640 |} Bedeguar 2 284, 6 Round-leaved 295 Cherry 609-10 
Bast-tree 2 512} Bed-flower 3 258 Slender 296 Dwarf I 611-12 
Bath-flower I 525 | Bed’s-foot 3 138-9 Tall 297 Glandular 611 
Batter-dock 175; 3532 | Bed-straw 3 258-67 | Bellis 3 402 Gray 608, ro 
Battlefield-flower 2 563 Arkansas 3 262] Bells 2 92 Low 611 
‘Bay D Ya sy Asperula 267 | Bell-tree 2.722 Mahogany 609 
Dwarf 2 574 Clayton’s 264] Bell-ware I gI Mountain 609 
Holly 527 Coast 266 | Bellwort I 518-9 Old Field 608 
Loblolly 527 Corn 259 Mealy 518 Paper 609 
North Carolina 81 Fragrant 262]! Belly-ache weed 3 383 Pin or Poverty 608 
Red 133 Great Hedge 263] Benjamin, Red 525, 6 Red 6090 
Rose 2 590, 680 Hairy 260 Sweet 3 526 River 609 
Swamp 2 134 Lady’s 258 White I 526 Running 2 704 
Sweet 82, 133 Our 258 | Benjamins I 525-6 Sandberg’s 611 
Tan 2 527 Marsh 3 263-4] Benjamin bush 2 135 Scrub I 616 
White by aie} Northern 261 | Bennels T2232 Silver 609-10 
BAYBERRY FAMILY I 584 Pointed 265 | Bennert 2655; 3 402 Spice 609 
Bayberry I 585 Purple 262} Bennet Herb 2 270-1 Swamp 610 
Tallow 585 Rough 265 | Bennet Pimpernel Sweet 609-10 
Baybush 584 Shining 265 2 631 Water 609, Io 
Bay Galls 2-133 Small 264 -Weed I 192 Western Red 610 
Beach-pea 2 413 Southwestern 260] Bent-grass 1 203-7, 213 White 608-9 
-plum 326 Sweet-scented 262 Sea. I 363 Yellow, SS 610 
Bead-ruby TS 7, Three-flowered 262] Benzoin-gum 251355) Bird-brier 2 284 
Beak-sedge I 341-6 Wall / 3 258] Berbine 3°95 -bell 3 335 
Beaked Rush 1 342-7 White 263 | Bergamot 3 132-3 -seed 11703 3 245,539 
Beam Tree 2 290 Wood « 3 263 -herb, or -mint 3 150 -weed I 661 
Bean, Blackeyed 425 Yellow 258 Praitie: _ 3 134 -vetch 2 409 
Buffalo 377 | Bee-balm By USS 7/ Purple, Wild, 132 Bird-on-the-wing 2 452 
China 2 425|| _ -tree 2 512| Bergia, Texas 2 539] In-the-bush 2 130 
Hyacinth 2 417| BeecH FaMiLy 1 614] Berry-alder 2 504] Bird’s-bread 2 208 
Indian 3 238| Beech, American 615 -tree 2 241 -eggs 2 64 
Magoty-boy 237 Blue I 606 | Besom 2 350, 604 -eye 2 51, 121,359, 
Sacred 277. Dutch 587 | Beth-flower,-root, 1 525 716; 3 201-3 
Smoking 3 238 Red, White 615 | Betony 3 128 Red 2 60, 426 
Water 2 77 Water 1606; 2242 Head Bge2t White 2 43-4 
Wild Kidney 423 | Beech-drops 3 236 Paul’s 3 148, 201-2] Bird’s-foot Trefoil 359 
Bean (Wild) Albany 2 673 Wil 2 273) Bird’s-nest 
2 418, 23-4 Carolina 674 Wood 3 128, 146, 221 2 625, 634, 674 
Bean-tree 3 238 False 675 | Bibernel 2 266 Giant 673 
Bean-trefoil 3 18] Beech-fern 1 23 | Biddy’s Eyes 2 563 Yellow 2 675 
Bean-vine 2 423 -wheat 672 | Bigbloom 2 81 | Bird’s-nest-plant 2 625 
-weed I 674; 3 226| Beefsteak plant Bilberry 2 292; 3 273 Bird’s-pepper 2 165 
-berry 2 488, 705 3 154, 221 Bog 2 699 | Bird’s-tongue 
Hed 2572-3, 693-4 | Beef-suct tree 2577 Burren 2 693 I 661; 2716 
Bear-bind 1674; 3 47 Bee’s-nest-plant 625 Dwarf 699 | Birth-root I 525 
- e 699 | Brrruwort FAMILY 
grass Bee-plant Great ; IRT 
I 67, 512-3; 2542 Rocke Ne 2 197 Oval-leaved 700 I 641 
COE pee qene~ 512 aed 2 697 ee wart I G45 
-tongue I 514 : a 270 pright 45 
erred Beet, wild 2 ee Thin-leaved 699 | Biscuit-leaves 528 
I 113-9, 129, 201 Beetlebung 2 5 Bilsted 235 | Biscuits 2 202 
-tongue 3 182-7 | Beetle-weed 2 7°7| Bindweed 1674; 342 -6 | Bishop-weed 2 654 
cobea 3 186 | Beeweed 3 414-5 Black I 674 Mock 2 657 
Foxglove 184 | Beewort I 446 Blackbird I 674 | Bishop’s-cap 2 229 
Funnelform 185 | Beggars’-lice Blue 3 167 -elder 654 
Gray 183 2401; 378,259,497 Bracted 3 46-7 -wig 180 
Hairy 182 -buttons 3 547 Climbing I 674 -wort 3 128 
Hayden’s 187 -needles 2 627 Corn or Ivy 674 | Bistort, Alpine 1 671 
Large-flowered 186] Beggar-ticks 266, False 3 42 | Bite-tongue 670 
Large Smooth 187 396; 3 78, ie 7 Fringed Black 1 674 | Bitter-ash 2 491 
Long sepaled 185 Swamp 3 496 Great 46 -bloom 3 18 
Pale-blue 183, 7 | Beggar-weed I 661 Hedge 3 46 -bush I 62 
Sharp-leaved 186 | Behen 2 64 Hoary 3 47 -clover 3 18 
Slender 185 | Belia, Low 3 303 Ivy I 674 -buttons SB E22 


604 


Bitter-cress 2 184-5 
Bitter-flower, Great 
2 85 
Bitter-grass Teele 
-herb 3 181 
Bitter-nut I 580 
Water 581 
Bitter-plant ier 
Bitter-root B02 
Bitter-sweet 
20267.) 20672 
Climbing 2 


493 
False or Shrubby 493 
Bitter-trefoil 18 
-weed 3 341, 390, 442 
-worm 18 
Black Alder 
I 613; 2 480, 504 
68 


Black-a-moor I 
Blackberry 278-81 
Bailey’s 2 280 
Bristly 279 
Creeping 281 
Dwarf red 278 
High Bush 280 
Hispid 281 
Knee-high 278 
Leafy-flowered 279 
Low 278 
Low Bush 281 
Low Running 281 
Millspaugh’s 279 
Mountain 280 
RunningSwamp 281 
Sand 278 
Tall 280 
Blackberry lily 1 542 
Black-bur 2277 
Black-cap 
I 68, 484; 2 277 
-caps 485 
Dogwood 2 504 
Black-eyed Susan 525 
Black-flower I 493 
Black-grass 2oOsicnt 
Black-head Grass 485 
Black-gum 2 665 
-haw 2 720; 3 233 
-heart I 668 
-jacks 3 246 
-laurel 2 52 
-lime-tree 2 512 
-linn 82 
-moss I 456 
-nonesuch 2 35a 
-root 3 204 
-sampson 475 
-sanicle 2 623 
Black-seed 2 351 
-snake-root 2 624 
-snap 2 696 
Black-walnut I 579 
Blackweed I 7, 341 
Black-wheat ge 223 
Black-wort 92 
Bladder-campion 2 64 
-ketmia 2e52c 
BLADDERNUT FAMILY 
2 493 
Bladder-nut 493 
Bladder-pod 2 154-6 


BLADDERWORT FAMILY 


3 225 

Bladderwort 3 226-32 
Closed Beer 
Fairy-wand 232 
Fibrous 228 
Flat-leaved 228 
Greater 229 
Hidden-fruited 229 
Horned 232 
Humped 227 
Lesser 228 


Bladderwort 
Pin BARE 
Purple 226 
Reclined 230 
Reversed 231 
Rush 232 
Swollen 227, 9, 30 
Tiny 3 231 
Two-flowered 227 
Zig-zag 231 

Blanket- fea 3 173 
-flower coat 

Blaver 2 13735557 


Blazing Star 
I 489, 511; LS 7 


| Bleaberry 
Blecdine- heart, Wild 
2 143 
Blephilia 3 135-6 
Blind-eyes 2 137. 
Blinks 2 39 
Blister-flowers 2 112 
-plant nig 
-weed I12 
-wort 2 Lie 
Blite Py ty 
Sea 23-4 
Strawberry 23 0S 
Blood-leaf 28 
-root 2 140 
-staunch ~ 3 442 
Bloodstrange 1/203 
BrLoopwort FAMILY 
I 530 
Bloodwort 
1658; 2.266; 3:2 
Mouse 3 333 
Striped 332 
Bloody Warrior 2 214 
-noses 1304 
Bloomfell 2 359 
Blooming Sally 2 590 
Bloomy-down 275 
Blowball 3 315 
Blue-ash 2 724 
Blue Beech I 606 
Blue Bell 


i SiO) 2 tes eesios 
Blue Bells 3 83, 200, 95 
of Scotland 2)205 
Blue-bonnet iis) 
=bottle £ 5105 3/557 


-buttons 290 
-caps 290, 557 
-cups 3 290 
-curls 2.105, L05 
-eyes 7 
-flag I 537-9 
-ginsing 2 128 
-jessamine 2eTe3 
-tops 3 557 
Blueberry 2 128, 698 
Black 702 
Bog 2 699 
Canada 701 
Dwarf 701 
Early 703 
Elliott’s black 702 
High-bush 700 
Low 700 
Low black 702 
Low-bush 701 
Mountain 700 
New Jersey 2 701 
Pale 700 
Small black 703 
Southern black 703 
Sugar 701 
Swamp 700 
Tall 700 
Blueberry-root 2 128 
Blue buttons “3 292 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Blue Devils 3 94,415 
Blue eye 3 201 
Blue-eyed-Babies 3 251 

seiaes PSAs 5 5 5 25% 


-san 1544 
-Mary 1543; 3188 
Blue-flag I 537-9 
Slender I 541 
Blue-hearts 3 205 


Blue-joint, see Grass 
Blue Mountain Tea 389 
Blue-pipe 2 724 

-sailors 3 305 
Blue-stem,see Grass 94 


Blue- tangles 2 605 
-thistle 3 94 
-weed 94 

Bluets 3 251-2 
Clustered 254 

Bob’s-root 2 364 

Bog-asphodel I 487 

Bog-bean 3 18 

Bogberry 704 

Bog-bilberry 2 699 
-birch 2 503 
-hop 2) 207, -130rS 
-leaves 2 207 

’ -myrtle 3 18 
-nut 3 18 
-onion I 7,442 

Bog Rush 348, 467 
(Many Junci so 

called) 
False 344 
Water 348 

Bog-strawberry 2 258 

Bog-violet 3 226 

Bogwort 704 

Bokhara-clover 2 352 

Bolder or Boulder 331 

Bole-wort 2 657 
-weed 3 557 

Bolgan-leaves 3 306 

Boltonia 404-5 

Bonaparte’s Crown 

2 474 

Boneflower 3 402 

Boneset 392 
Bastard 359 
Blue 362 
Climbing 3 363 
Deerwort 361 
False 364 
Prairie 3 364 
Purple 357 
Resin 361 
Spotted 356 
Tall 257 
Upland 359 

Bonewort 3 402 

Bongay 2 408 

Bonnets 2078 
Alligator 79 
-blue S55 

Bonny Rabbits 3 178 

Boor- or Bore-tree 

3 268 

Boots 2 85 

BoraGE FAMILY 3°72 

Borage 3°93 


Bottle Brush 
I 39,40; 2 294, 612 


Bottle-grass 74 siete 
Bottle-rush 204 
Bouncing-bet 2 73 
Bountry 3 268 
Bowel-hive 2 264 
Bowman’s-root 


2 248, 469; 3 204 


Bow-wood TeOs2 
Box FAMILY 2 480 
Box Bees 


Vor. IIT. 
Box, Flowering 2 607 
Mountain 693 


Wild Running 3 


2 
Box-berry 2 693; 3 ae 
-elder 2 408 
-thorn 3 168 
Box-wood 2 292, 664 
False 664 
Boys’ Love 3 525-6 
Boysand Girls 2 142 


Brake, Bracken 1 7, 32-3 


Backache 30 
Buckhorn 7 
Cane- 295 
Hog 32 
Knotty 25 

Braken, Lady 32 
Osmond 7 
Rock 36 
Royal 7. 
Stone 36 
Swamp 7 
Sweet 21 
Winter 33 

Bramble, Mt. 2 276 
Arctic ae 
Horse 2 
Strawberry-leav ea 

278 

Brank I 672 
Bread-and-butter 

I 528; 3 177 

Bread-root I 7, 264 
Indian 2 363 
Missouri 363 

Breakstone 25a 

Breast-weed I 578 

Breeches-flower 2 142 

Breeches, Kitten 142 
Little-boy’s 142 

Brere-rose 2 284 

Breweria 3 41 

Brick-timber 2 490 

Bride’s Laces I 170 

Brideweed 3) Ey, 

Bridewort 2 249 

Brier, Bramble 2 284 
Bird or Hip 284 
Sensitive 25338 

Brier-berry 2 278 

Brier Rose 284 

Bright-eyes 3 252 

Brinton Root 3 204 

Broad-leaf 3 245 

Brome I 273-80 
Briza-like 280 
Field 279 
Rye 278 
Short-awned 280 
Soft 278 
Wild 280 

Brook-bean 3 18 
-flower 3 65 
-grass T DDG 


Brooklime 2 162; 3 200 
Brook-mint 3 150 


Brook Sunflower 495 
Brookweed 2.716 
Broom 2 350 
Base or Dyer’s 350 
Clover 345 
Green, Scotch, 350 
Indigo 345 
Sweet 392. 
Thorn or Prickly 349 
Yellow 345 
Broom-brush 2 530 
-corn, Wild 232 
-cypress 2 16 


Broom Crowberry 2 479 
Broom-grass 118 


Vot. III. ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 605 
BROOM-RAPE FAMILY Bull-brier I 529 | Burseed Button- : 

3 233 | Bull-dogs 3 178 | Bursting-heart es Small Bis 68 

Broom-rape 3 233-5 -flower 2 85 | Bush-Clover 2 402 Syren Beggars a 
Naked, Pale, 233 -nettle 3 165 Britton’s 4oad Ruiton thee 3 pee 
Broom-sedge 1 116,182 -nut I 582 Creeping 403 | Button-weed Age 
-weed 3 37 -thistle 3 552 Hairy 407 2521; 3 288, 556-7 
Brotherwort 3 141 | Bullock’s-eye Py Pin Intermediate 406 | Button-wood +e $ 
Brown Betty 470 | Bull’s-eye 3 518 Japan 408 -Shrub 3 ate 
-dragon 442 English 470 Mann’s 404) Buczies 3 548 

Brown-eyed Susan Bull’s-foot 3 531 Narrow-leaved 407 ? 
-3 470 | Bullpates eis Nuttall’s 405 | Cabbage 2 193 
Brownwort 3 115 | Bullpoll 215 Prairie 408 Bargeman’s 193 
Bruise-wort Bull-rattle 2 64,8 Round-headed 407 Clubfoot 445 
2 73,141; 3 402| Bull-segg 168} Sharp-fruited 405|/ Meadow 445 
Brunella 3 115 | Bullweed 3 557 Slender 406 Sjerraik 445 
Brushes 474, 557 | Bullwort 2 657 Stuve’s 405 Swamp 445 
Bubby-bush Bulrush 1 330-7] ‘Trailing Aes)” “Water ee 
Perec {N27 7 Bog I 331 Wand-like 406 | Cabomba 2 76 
BucK-BEAN FAMILY Leafy 336 | Bush-Honeysuckle Cactus FAMILY 2 568 
3 17-8 Pole or Pool 1 331 3 283 Cactus 569-72 
Buck-bean 318 | Bulwand By eg) Tartarian 282 Plains 2 569 
Buckberry 2 697 Am.-great 331 -pea 2 3 Prickly-pear 571 
Seedy 691 | Bumblebee-root 1 525 Butcher’s Prick-tree Russian 25 
Buckbush 3 277) Bumelia 2 719 2 492, 504 Tree 2 573 
BucCKEYE FaMILy 2 498} Bunchberry 2 664 | Butter-bur 3 531-2] Cadlock 573, 193, 5 
Buckeye 498-500 | Bunch-evergreen 1 45| Butter-and-Eggs 3 177] Calabazilla 3 291 
Big, Fetid, 2 499 | BUNCH-FLOWER Butter-cresses 2 111-2] Calais, false 324 
Large 499 FAMILY I 485} Buttercup 2 104-17 | Calamint 135-45, 152 
Little 500] Bunch-flower I 493-4 see Crowfoot Field, Lesser 135, 41 
Ohio 2 499 Black 493 2 109-17 Low, Slender 3 139 
Purplish 499 Crisped 494 Allen’s 109 Water 152 
Red see Bunch-grass, see Grass Arctic 105 | Calamint-balm 139, 41 
Southern 500 | Bunch-Pink 275 Bristly 112| Calathian violet 3 12 
Sweet, Shrubby, ae -Plums 2 664 Bulbous 112 | Calfkill I 214 
Western 499 | Bunk 2 653; 3 305 Bundle-rooted 114 | Calf-snout 3 178 
Buckhorn Brake 1 7] Bur 3 547 Creeping 113 | Calico bush 2 684 
Buck’s-beard 3 313 Buffalo 3 166 Early 114 -wood 2 722 
Buck’s-grass I 47 Button, Clot, Figwort 117 | Calla, Wild 444 
Bucks-horn 3 346, 547 Hairy 114 | CALTROP FAMILY 2 442 
147; 2167; 3 247 Canadian 3 76, 346 Harvey’s 109 | Caltrop, Greater 2 443 
BucK-THORN FAMILY Cockle 547 Heart-leaved 108 irsute 443 
2 501 Common ie Hispid 113 Land 442 
Buckthorn 2 502-3 Creeping I 47 Lapland 106 Water 2 612 
Alder 2 503-4 Ditch, Sea, Sheep, Macoun’s 112] Caltrops 3 559 
Alder leaved 2 503 3 346 Marsh 113 | Calypso I 575 
Carolina 2 503,719 Great 3 547 Meadow 2 85,111 | Camass, Death 492 
Lance-leaved 2 603 Hurr 3 547 Missouri 2 106 Eastern 509 
Purging 501 Prickly I 615 Northern 108 Nuttall’s 492 
Southern 719 Sand T1267 Nuttall’s 117 | Camel’s-foot 550 
Woolly 720 Small Sheep Bae, Pigmy 108 | Cammock 2 533; 3516 
BuCKWHEAT FAMILY Spiny 459 Pursh’s 105 | Camomile 2 121 
I 646 | Bur-chervil 2 629 Running 2 251 Arctic 520 
Buckwheat I 672 -clover 2 352 Snow 107 Corn or Field 517 
Climbing false 674 -heart 2 352 Spotted-leaf 117 Dog’sorFetid 516 
Crested 674 | Bur-REED FAMILY I 69 Swamp 113 German 3) 52% 
Hedge 675 | Bur-reed I 69 Tall III GardenorLow 517 
Rough tartary 672 Branching 70 Tufted E14 Rayless 3 521 
Buffalo-berry 2 576-7 Floating 73 | Butter Daisy I1I-3 Red 2 121 
-apple, -bean 2 377 Green-fruited 71 | Butter-flowers 2 112 Scentless 3 520 
=pur ~ 3 166 Nuttall’s 70 Great 85 Scotch or White 517 
-nut I 641 Small 74 ore print 2 521 Wild 3 519, 21 
-pea 237 7. Simple-stemmed 73 -ros III YelloworOx-eye 517 
-weed 3 341 Stemless 72 Butterfly- dock 3 532] Campion 2 63-4 
Bugbane, Amer. 2 92] Bur-weed 3 344-6 -banners 2 142 Bladder 64 
False 103 | Burdock 3.34 -pea 2 417 Meadow 69 
Bugle 3 101 Prairie 3 462, 547-8 -weed 3 25 Moss 63 
Bitter 148 | Burhead 3 96] Butternut I 579 Red 6¢ 
Brown 101 Creeping 96 | Butterweed 2 521; Red Alpine 2 62 
Erect 101 Upright 96 3 320, 442, 540 Rose 71 
Bugle-weed 3 146-7 | BuRMANNIA FAMILY Butterwort 3 225-6 Snowy 63 
Bitter 3 148 1 546 | Button-ball 2 242 Starry 63 
Sweet 146 | Bur-marigold 3 495-7 -bush 3 255 Western White 63 
Bugle-wort 3 146] Burnet, Am. Gt. 2 265 | Button-Snakeroot _ White 68 
Bugloss Stier -salad 2 266 2 622; 3 366-8 | Camp-root 2 271 
Sea 62 Garden 266 Dense 3 367 | Canada-balsam I 63 
Small 93 | Burnet-rose 716 Dotted 366 -pest 3 8 
Small Wild 84 | Burning-bush 491 Hairy 367 -potato 3 486 
Viper’s 94 | Burnt-weed 2 590 Large 367 -root 3° 25 
Bugseed 2 21 | Burnut, Ground 2 442 Loose-flowered 368 -snake-root I 642 
Bugweed 21 | Burn-wood Bark 2 485 Mountain 3 365 -tea_ 2 693 
Bullace 2 326] Burren Myrtle 2 604 Prairie 367 | Canadian Bur 3 76 
Bull-berry 2 5761 Burton I 584 Slender 366 | Canary Grass I 170 


606 


Canary Grass 


Southern I 170 
Wild 170 
Canary-vine 2 143 
Canby’s Mountain 
Lover 2 492 
Cancer-drops 3 236 
-jalap 2 26 
-root 3 233-4 
-weed 129 
-wort Rei, 
Candleberry I aae 


Candles, Swamp 2 712 


Candle-tree 3 238 
Candlewick 1 68; 3173 
Can-dock 2578 
Cane I 205 
Giant 295 
» Maiden 121 
Saitch 295 
Sedge 446 
Switch 205 
Cane-brake 205 
Can-hoop 2 489 


Canker-blooms 2 284 
-lettuce 
-root 2 88, 718; 3 ie 


-rose 2 284 
-weed 3 335, 542 
-wort 315 
Canna-down I 324 
Canoe-wood 2 83 


Canterbury-bells 


3 295-6 
Cape Gooseberry 3 161 
Dwart 156 
CAPER FAMILY 2 196 
Caper, Wild 471 
Caper-bush 471 
Capers 2 85 
Capmint 3 139 
Capon’s-tail 2 93 
-grass 269 
Caraway 2 659 
Wild 3 537-8 
Carberry 2 241 
Cardinal-flower 3 300 
Blue 301 
Careless 252 
Carizzo 212 
Carnation-grass 
I 397,415, 28 
Carpenter’s-herb 
2 yLoraats 
-square 3 179 
-weed 115 
Carpet water 2 230 
Carpet-grass DA25 


CARPET WEED FAMILY 


2 34/ 


Carpet-weed 35 
Carrion-flower 527 
CARROT FAMILY 2 619 
Carrot, Wild 2 625 

American 625 
Carrot-weed 3 341 
Carvies 2 659 
Case-weed 158 
Cashes 2 653 
Cassandra, Dwarf 687 
Cassena 489 

Evergreen 489 
Cassia, Dwarf 2 337 
Cassiope 2 686 
Castor-bean 461 
Castor-oil Plant 461 
Catalpa — 3 238 

Hardy 238 

Western 238 
Catawba 3 238-9 

Tree 238 
Cat-berry 2 490 


Catbrier I 528 
Catchfly, Corn 2 66 
Dover 2 64 
English 67 
Forked 67 
Garden 66 
Limewort 66 
Lobel’s 2 66 
Night-flowering 67 
Nodding 2 64 
Nottingham 64 
Round-leaved 65 
Royal 65 
Sleepy 66 


Small-flowered 67 


Striate 66 
Sweet William 2 66 
Catch-weed 3 84, 259 
Cat-gut 2372 
Cat-locks 324 


Cat-mint, Catnep 3 113 


Cat-o’nine-tails I 68 
Cat-peas 2 409 
CAT-TAIL FAMILY 1 68 
Cat-tail, -flag 1 68-9 
-tree 2 492 
-whin 2 284 
-whistles I 40 
Cat’s-clover 2 350 
-cradles 3 246 
-ear 3 309, 331 
-eye 3 201-3, 93 
-faces 2 563 
-foot 3 114, 450-2 
-milk 2 473 
-paws 3 452 
-tail E 30502) 612 
Blue 394 
Cedar I 65-7 
Carolina 66 
False White 65 
Feather-leaf 65 
Ground 542 
Post 65 
Indian 607 
Red I 67 
Shrubby Red 67 
Southern White 65 
Swamp 65 
White 65 


Celandine 2 117, 21, 41 


Greater 2 141 
Brook 440 
Lesser prea 7, 
Wild 440-1 


Celandine Poppy 2 140 
Celery, Wild 
I 106; 2660 


Water 1106; 2111 
Celery-seed I 658 
Cenizo 2 19 
Centaury 251; 3558-9 

American 3 1-3 

Buckley’s 2g 

Corn a) 

European 2.557 

Ground 2 450 

Tall Bs 

Texan a3 

Western Ries 

Winged 559 
Cercocarpus 2 274 
Cereus 2 569-71 
Chaerophyllum 629 
Chadlock 2 192 
Chafe-weed 

p 447, 454, 456 
Chafi-seed Bi 207 
Chaff-weed 25717, 
Chain-fern I 24 

Netted 25 


Chair-maker’s rush 330 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES, 


Chamaesaracha 3 163-4 
Charlock 2 191-2; 3 195 


Cheat Pane, 
see Grass 
Pine 250 
Checkerberry 
2 603; 3 255 
Cheese-rennet 3 258 
Cheeses, Cheese-flower 
2 515 
Doll, Dutch, Fairy 
515 
Cheese-bowl 25587; 
Cheese-cake 2 514 
Cheese-flower 2 514 
Chenile 3 168 
Cherry, Appalach. 2 326 
Bird 328 
Bessey’s 2327, 
Black-merry 327 
Cabinet 329 
Choke 329 
Western 329 
Crab 2 327 
Dwarf 326 
Egriot 327 
Fire 328 
Gean 327 
Ground 3 156-63 
Indian 2 292,503 
Mahaleb 328 
ay 291 
Mazard 327 
Mer Be, 
Perfumed 328 
Pigeon or Pin 328 
Red 327 
Rum 2 329 
Sand 2 326 
Sour 327 
Sweet 327 
Western Sand Bey, 
Whiskey 329 
Wild 3 160, 327,9 
Rocky Mt. 329 
Wild Black 2 329 
Wild Red 328 
Winter 2501; 3 162 
Cherry-pie 2 591 
Cherry-tomato 3 168 
-wood 270 
Chervil, Bur 2 629 
Garden 2 629 
Hemlock 626 
Needle 627 
Rough 626 
Spreading 629 
Sweet 627 


Teinturier’s 2 630 
Wild 
Chess, see Grass 
Chestnut, Am. I 615 


Water 2 612 
Chicken-berry 
2 693; 3 255 
-fighters 2 547 
Chicken’s-toes 2 21 
Chickenweed 
2 43; 3 539 
Chickweed 2 41 
Allbone 2 44 
Alpine 49 
Barren 49 
Blinking 39 
Clammy 48 
Common 43 
Field 49 
Forked 29, 30 
Germander 3 203 
Gravel 2 30 
Great 2 43 
Indian 235 


Vor. III. 


Chickweed, Ivy 3 203 
Jagged 50 
Larger Mouse-ear 48 


Low 42 
Marsh 43 
Meadow 49 
Mouse-ear 2 42-7 
Small 47 
Spring 47 
Nodding 48 
Red 2 716 
Sea 58 
Short-stalked 48 
Silver 27 
Slender-forked 30 
Star 2 43 
Star-flowered 705 
Starwort "50 
Tennessee 44 
Water 2 39,42 
Water, Mouse-ear 


2 40 
-Wintergreen’ 2 715 
CuHicory FAMILY 3 304 
Chicory 3 305 
Chicot 2 340 
Chigger-weed 3 516 
Childing Cudweed 447 
Daisy 3 402 
-Pink 274 
-Sweet William 74 
Children’s-bane 658 
Chimney-pinks ° 2 73 
-sweeps 3 246, 485 
China-root, Amer. or 
False Ta520 
China-tree, Wild 2 500 
Chinks 693 
Chinquapin I 616 
Chittam-wood 2 485 


Chittim-wood 719 
Chive garlic I 497 
Chives: 497 
Chloris I 225 


Chocolate, Indian 2 272 
Chocolate-flower 

-root 2 270,2 
Choke-berry 2 290-1 


-cherry 329 
aoe "Serangle 3 36 
290 
Chelke-pear 2 290-1 
-vetch 3 234 
Chowley 2 425 
Chris-root 2 87 
Christmas Evergreen 
I 49 
-flower 2 88 
-rose 2 87 
Christmas-green 1 48 


Christopher, Herb 
17; 290; 3128 


Church-brooms 289 
Churnstaft 2 473 
Cicely, Fool’s 645 
Rough 626 
Sweet 627-8 
Cicely-root 627 


Cinnamon-wood 2 134 


Cigar-tree 3 238 
Indian 238 
Cinquefoil 2 250-7 
Arctic 255 
Branched 256 
Bushy 252 
Coast 257 
Common 251 
Cut-leaved 256 
Diffuse 252 
Downy 254 
Five-stamened 253 
Glandular 253 


ae 


Vo. III. 

Cinquefoil 
Hoary 2 253 
Nicollet’s 252 
Northern 254 
Marsh 258 
Plains 257 
Prairie 257 
Purple 258 
Robbins’ 255 
Rough 253 
Rough-fruited 254 
Shrubby 262 
Silvery 253 
Snowy 2B5 
Tall 263 
Three-toothed 262 
Wood 250 
Woolly 256 

Cissus 2 510 

Citronella 20053 

Civet I 408 

Clabber-spoon 2 417 


2 679 
Clammy-weed 2 199 
Clamoun 684 
Clapwort 3 235 
Clary, Wild Clary 
3 130-1 
Vervain 130 
Claver-grass 3 259 
Claver 2 359 
Claytonia 2 37 
Clayweed 3 531 
Cleareye ey eh 
Clearweed I 637 
Cleats Assit 
Cleavers 3 259 
Dyer’s 264 
Yellow 258 
see Bedstraw 3 259 
Cleever-wort 3 259 
Clematis 2 122-6 
Addison Brown’s 123 
Curl-flowered 2 123 
Dwarf 2125 
Fremont’s 2 125 
Marsh 123 
Mountain 126 
Scott’s 126 
Silky 125 
Sim’s 123 
Whorl-leaved 126 
Cleome 2 197 
Prickly 2 1907 
Cleomella 2 198 
Cliff-brake I 32-3 
Climath 2 484 


Climbing Bittersweet 


bet ey, 2 403 
Climbing False Buck- 


wheat I 674 
CLIMBING FERN 
: FAMILY I9 
Climbing-Fern 9 
-fumitory 20043 
-hempweed 3 363 
-sailor [75 
Cling-rascal 259 
Clintonia I 514-5 
Clock 3 246 
Cloistered Heart 3 12 
Clot-bur 3 344-5, 547 


piny 344 
Clothier’s Brush 3 289 


Clotweed 344 

Cloud-berry 2 276, 80 

Clove-strip 2 589 

Clover 2 351-8 
see Bush-clover 

2 402-8 

Alsatian 357 


Clover, Alsike 2 357 
Beckwith’s 35 
Bitter a, 
Bokhara 352 
Brazilian Bem 
Broad-leaved 355 
Buffalo 357 
Bur 2aane 
Burgundy 2) BS 
Bush 403-8 
California 25352 
Carnation 2a3 cc 
Cabul 352 
Calf 355 
Carolina 358 
Cat’s 359 
Chilian 351 
Cow 356 
Crimson 2 355 
Dutch 358 
Dusty 407 
French 355 
Giant 356 
Hare’s-foot 355 
Hart’s 353 
Heart 352 
Honeysuckle 355,8 
Hop 2 351-4 
Horned 351 
Honey 352 
Italian 355 
Japan 408 
King’s 353 
Lady’s 2 431 
Least Hop 2 354 
Low Hop 354 
Mammoth 356 
Marsh 3 18 
Meadow 355 
Old-field 355 
Pea-vine 356 
Pin 2 430 
Plaster 353 
Prostrate Mt. 356 
Prairie 2 SE ok 
Purple 355 
Pussy 355 
Rabbit-foot 355 
Red —~ 355 
Running Buffalo 357 
Sleeping 2 431 
SmallerHop 2 354 
Snail 351 
Spotted 352 
Stone 355 
Swedish 357 
Sweet, White 2 352 

Yellow 353 
Tree 352 
White - 358) 
Wild 408 
Winter 3 255 
Yellow 29354 
Zig-zag 356 

Clover-broom 345 

Clown's All-heal 3 125 
-heal 125 
-woundwort By ir25 


Citus-Moss FAMILY 


I 47-8 

Club-moss I 43-7 
Alaskan 46 
Alpine 46 
Box 44 
Carolina 47 
Cedar-like 47 
Chapman’s 45 
Dwarf I 49 
Fox-tail 45 
Interrupted 46 
Lloyd’s or Rock 44 
Marsh 44 


ENGLISH INDEX, INCLUDING POPULAR PLANT 


Club-moss, Prickly 1 49 


Savin-leaved 46 
Shining 44 
Stiff 46 
Tree-like 45 
Upright 43 
Club-rush I 327-34 
Chairmaker’s 330 
Dwarf 327 
Hedge-hog 308 
Red 332 
River 333 
Sea 333 
Wood I 334 
Club-weed 39557, 
Cluckies 2 92 
Clump-head grass 337 
Clump-foot I 445 
Cluster-berries 679 
Coakum 2 26 
Cocash 3 419 
-weed 3 544 
Cock’s-head 
2 380; 3 425 
Cockle 2 61,73 
Cockle-bur 
2 266; 3 346,547 
ea 3 346 
Cockle-button 3 547 
Cocks 246 
Cockscomb, Red 22 
Yellow 2°23 
Cocksfoot I 251; 2 93 
Cock-sorrel I 654 
Cockspur, see Thorn 
2 207 
Coco-grass 304 
Cod-head 3 181 
Codlings and cream 
2 591 
Coe-grass 469 
Coffee, Magdad 2 336 
Negro 336 
Wild 275 
Coffee-bean, Am. 2 340 
Coffee-nut 2 340 
-plant 2 585 
-senna 2 336 


-tree, Kentucky 2 
-weed 3 305, 335-6 
8 


Cohosh 9-90 
Black go-I 
Blue go, 128 
Red 90 
White 90 


Cojote, Chili aheon 
Coleseed 2 193 


Colewort, Hare’s 3 317 
Colic-root uy Gus ice 
642; 3 365-7 
-weed 2 142-4, 264 
-wortt 2 264; 3 217 
Collard 1 445; 3 536 
Collinsia 3 188 
Collinson’s Flower 153 
Collomia 3 63-4 
Colt-herb 531 
Coltsfoot 
2 85,707; 3 531-2 
False I 642 
Heart I 642 
Sweet 3 531-2 
Colt’s-tail 442 
Columbine 2 92-3 
Feathered 2 120 
Columbo, Amer. 3 15 
Coma 2 719 
Comandra I 640 
Comb ( Dak.) 476 
Comb-flower 478 | 
Comfrey 3 92 | 
Middle ror | 


NAMES. 607 


Comfrey, Rough 92 
Wild Nase 


Compass-plant 461 
Saracen’s 542 
Cone-flower 542 
Brilliant 471 
Britton’s 3 471 
Bush’s 476 
Ciasping-leaved 473 
Gray-headed 474 
Great 472 
Green-headed 473 
Large-flowered 472 
Long-headed 474 
Orange 471 
Prairie 3 474 
Purple 475-6 
Short-rayed 475 
Showy 472 
Sweet 470 
Tall 473 
Thin-leaved 470 
Woodland 471 
Congo-root 2 364 
CONIFERS I 55-67 
Conobea 3 191 
Conquer-John I 521 
Conquerors 2 498 
Consound 3 67, 92 
Consumption-weed 
2 668 
Continental-tea 2 676 
-weed 3177 
Convulsion-weed or 
-root 2 674 
Coolweed I 637 
Coolwort 2 22 
Cooper I 553 
Copal-balsam 2 235 
Copalm 235 
Copper-leaf 2 668 
Coral-and-Pearl 2 90 
Coral-berry B27, 
Coral Evergreen I 47 
Coral-root 1 574-7 
Crested 57 
Small, Late, 575 
Wister’s 575 
Cord-grass I 223-4 
Fresh-water 223 


Coreopsis, Golden 3493 
Coriander 2 264, 
Cork-elm I 
Corkscrew-plant 1 
CorK-woop FAMILY 


I 586 
Cork-wood 586 
Corn, Haver I 218 
Saracen’s 672 
Squirrel, Turkey 
2 142 
Wild I 514 
Corn-campion 2 61 
-bind I 674; 3 47 
-binks 3 577 
-blue-bottle 577 
-bottle 577 
-centaury 577 
-cockle 2 OI 
-flag I 540 
-flower 3 57Z 
-grass I 162 
-gromwell 3 87 
-heath I 672 
-kale 3 192 
-lily 3 47 
-Mayweed 520 
-poppy 2 137 
-pink 610 
-root 140 
-rose 61 
-salad 3 286-8 


608 


Corn-spurry 2 59 
Saracen’s I 672 
Cornel 2 661-5 


Alternate-leaved 663 
Bailey’s 662 


Blueberry 661 
Dwarf 664 
Lapland 665 
Low 2 664 


Northern Dwarf 665 


Panicled 663 
Price’s 661 
Red-osier 2 661-2 
Rough-leaved 662 
Round-leaved 661 
Silky 2 661 
Stiff 663 
Waxberry 2 662 
White 663-4 
Cornelian Tree 2 664 
Cornstalk-weed I 79 
Coronilla 21302 
Copper-leaf 2 668 
Corpse-plant 2 624 
Corydalis 2 144-6 
Golden 2 145 
Pale 2 144 
Plains 2 145 
Yellow 144 
Cost 3 519 
Costmary 519 
Cottagers 204 
Cotton, Wild 3 22,30 
Cotton-grass I 322-26 
Hare’s-tail I 324 
Cotton-gum 2 666 
-rose 3 447 


-rush, -grass I 322-6 


-seed tree 3 445 
-tree I 590 
-weed 2 453, 521 
Cottonwood 
I 588-91; 2 512 
Big I 590 
Black 588-9 
Narrow-leaved 588 
River 589 
Swamp 589 
Western 591 
Willow 588 
Yellow I 590 
Couch-grass 203, 283 
Black 192 
False 283 
Cough-root I 526 
Coughweed 3535 
Coughwort 531 
Cowbane 2 638 
Spotted 658 
Cow-basil 2 73 
Cowbell 2 64 
Cowberry 
2 258,607; 3 255 
Cow-cress 2 164 
-grass 1 661; 2355-6 
-herb 2 73 
-itch 3237 
-lily 2 85 
-parsnip 2 635 
-plant 681 
-quake I 256; 259 
-rattle 2 68 
-tongue reel 
-wheat 3 224 
Cowslip 
1517; 251, 114, 440 
American 2 85,717 
Spring 2 85 
Virginia 3 83 
Cow’s-lungwort 3 173 
-tail 442 
-wort 221 


Cowthwort 3 120 
Crab Apple 2 288-9 
American 289 
Fragrant 289 
Iowa 289 
Siberian 289 
Southern 289 
Sweet-scented 289 
Western 289 
Wild 289 
Crab Tree 2 288 
Wild 288 
Crab-grass I 661 
Little 1122; f 280 
Sprouting 138 
see Grass 
Crab-stock 290 
Cracker-berry 2 664 
Crackers 696 
Crain 2 117 
Crake-berry 2 479 
-needles 627 
Cramberry 2 704 
Cramblingrocket 2 200 
Cramp-bark 3 40, 220 
Cranberry, European 
2 704 
High-Bush 3 270 
Hog 2 479, 693 
Marsh 704-5 
Mountain 2 693,7 
peek 607 
Small 704 
Upland, Wild 607 


Cranberry-tree 
2 243; 3 270 


Crane-willow 3 255 
Crane’s-berry 2 704 
Crane’s-bill 2 427-9 
Bicknell’s 428 
Carolina 428 
Culver-foot 429 
Cut-leaved 429 
Dove’s-foot 429 
Long-stalked 427 
Pigeon-foot 429 
Round-leaved 428 
Siberian 2 427 
Small-flowered 429 
Spotted 426 
Starlights 429 
Wild 426 
Crap I 672 
Crape-myrtle 2 580 
Crashes 2 162 
Craw-berry 704 
Crawley-root 575 
Crazy-Bet 2 85 
Crazy-weed 2 379 
Colorado 2 390 
Stemless 390 
Woolly 379 
Cree-turnip 2 363 
Creek-grass 177 
Creek-stuft 223 
Low 224 
Upland 223 
Creek-thatch I 223 
Creeper 2 511; 3 46 
Japanese 54 
Creeping-bur I 47 


-Charlie 2 208; 3114 


-hemlock I 67 
-Jack 2 208 
-Jennie I 47-8; 

2 209; 3 292 
-Juniper I 67 
-Mitchella 3 255 


Creeping Snowberry 


2 704 


Cress, see Peppergrass 
2 165 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Alpine 2 186 
American Ge 
Bank 177, 
Bastard 2 164,8 
Belle-Isle 577. 
Bermuda 197 /7/ 
Bitter 184-5 
Brown 2 162 
Bulbous 2 186 
Cow 164 


Dock (succory) 3 306 


Early Winter 2177 
Field 164 
Garden 166 
Golden 166 
Hoary 165 
Lamb’s 2 184 
Land 177, 184 
Mouse-ear 2 176 
Penny 168 
Purple 2 185 
River 164 
Rock 2 179-83 
Rocket 177 
Round-fruited 161 
Sessile-flowered 162 
Spring 2 186 
Swine’s 2 167 
Thale 176 
Tower 181 
Town 2 166 
Wall 176 
Wart 2 167 
Water 161, 185-7, 192 
Winter 177 
Yellow 160-1, 177 

(see Watercress) 
2 160-3 

Crested dwarf Iris 
I 540 
Crinkle-root 2 188 
Cristatella, James’ 198 
Crocus (N. C.) 2 692 
Prairie, Wild 2 102 
Croneberry 704 

Crop-weed B55 7. 


Cross, Herb-of-the 3 95 
Cross-of-Jerusalem 


2 69 

Cross, Knight’s 69 
Maltese 69 
Crosstoes 2 359 
Cross-vine Su227, 
Crossweed 2 194 


Crosswort 2711; 3 

Croton 2 454-5 

Lindheimer’s 2 455 

Crouper-bush 3 255 
CROWBERRY FAMILY 

2 478 

Soy 479, 693,704 


Black 479 
Brown 479 
Plymouth 479 
Crow-corn nay aie 
Crowd-weed 2 192 
Crow-flower 2 69, 192 


CrowFroot FAMILY 2 84 


Crowfoot 1 45, 48, 85, 
99, 104-17, 187; 

2 426 
Biting III 
Bristly II2 
Celery-leaved 2 111 
Corn II5 
Cursed III 
Ditch III 
Dwarf 106 
Hooked iIt 
Ivy-leaved 116 
Kidney-leaved io 
Marsh RET 


Vot. III. 
Crowfoot 

Mountain 2 107,10 
Musk 3 283 
Prairie 2 109 
Rock 110 
Rough TEE 
Rough-fruited 108 
Seaside 117 
Small- flowereds II4 
Smooth IIo 
Spiny-fruited 115 
Water 2 105, 116 
Wood 3 283 


see Buttercup 104-17 


Crowfoot-grass 123 

Crownbeard 3 487-8 
Golden 489 
Small Yellow 488 
Sunflower 488 
Virginia 487 
White 487 


Crown-of-the-field 2 oth 


Crow-needles 627 
Crow’s-nest 2 625 
Crow-pea 412, 79 
-poison I 489 
Crowtoes 2 187, 359 
Crow-victuals 3) 114 
Crystal-wort 2 101 
Cuckle-bur 3 346 
Cuckles 497 
Cuckolds 496-8 
-dock 547 
Cuckoo-buds 2 112 
-button 3 548 
-flower 
2 68, 69, 184, 431 
-gilliflower 69 
-grass I 485 
-spit 2 184 
Cuckoos 2 69 
Cuckoo’s-meat 2 431 
Cucuma, Ohio 85 
Cucumber, Bur 3 293 
Creeping 292 
Star 293 


Wild 2100; 3 292-3 
Cucumber-root, Indian 


I 523 

-tree 2 81 
Cudweed 3 447,453 
Childing 447 
Dwarf 3.455 
Low 455 
Marsh 455 
Mountain 452, 5 
Norwegian 3 456 
Prairie 3 529 
Purplish 456 
Western Marsh 455 
Winged 454 
Wood 456 
Culver-foot 2 429 


-wort 2 93 
Culver’s Physic 3 204 
-root 204 
Cup, Ragged 460 
Cuphea, Clammy 2 582 


Cupid’s-delight 2 563 
Cupid’s-flower 3 42 
Cup-plant 3 460 
Cupseed 2 130 
Curdwort 3 208 
Cure-all 2 262 
Curlew-berry 479 
Curlock 2 192, 195 
Curly-grass ae) 
Curly Heads 2 125 
Currant 2 237-8 


American Red 2 237 
lack 237 
Buffalo 239 


Vo. IIT. 


Currant, Clove 2 239 


Daisy 3 437,518 
Arctic SUE T 
Big, or Bull 518 
Blue 3 305, 416-7 
Blue Spring 3 439 
Brown 470 
Bull’s-eye 518 


Butter 2 111-3; 3 518 


Childing 402 
Dog 516, 8, 20 
European 402 
Field 3518 
Garden, March 402 
Great-white 518 
Horse 3 518 
Irish 315 
Michaelmas 3 430 
(and Asters gener- 
ally) 
Midsummer 3 518 
Moon, or Maudlin Bee 
Nigger 
Ox-eye, Pages 
470, 518 
Pig-sty 516 
Purple 3 416 
Red 3 336 
Western 402 
White Epes 
Yellow 470 
Daffodil, a I 531 
Ox-ey 518 
Dafty- town dilly 1 525 
Dakota Potato 2 418 
Dalibarda 2 282 
Damask Violet 175 
Dame’s-gilliflower 175 
-rocket, -violet 175 
Dandelion Qeangs oe 
Blue 3 305 
Dog 3 310 
Dwarf 3 307,8 
False 3 307, 323-5 
Fall or Dog 310 
High 329 
Red-seeded 3 316 
Danes’ Blood 296 
Dangleberry 2 695-7 
Daphne 2 574 
Darkey-head 3 470 
Darnel, see Grass 
Darbya I 641 
. Date Plum 2720 
Day-berry 2 241 
Day-flower I 457-9 
-lily I 496 
Dead-Men’s Bones 177 


Fetid 238 
Flowering 239 
Garden Red 237 
Golden 2 239 
Indian 3 277 
Missouri 239 
Northern Black 236 
Red 2 237 
Skunk 238 
White-flowered 238 
Wild Black 238 
Currant-leaf 229 
Currant-tree 292 
Currantworts 236 
Cushion Pink 2 63 
Ladies’ 718 
CuSTARD-APPLE 
FAMILY 2 83 
Custard-apple 283 
Cut-heal 3 286 | 
Cut-paper I 632 
Cutting Almond 3 465 
Cyclanthera 293 
Cycloloma 2 16 
Cyperus I 297-310 
Awne 300 
Baldwin’s 310 
Bristly 308 
Brown 303 
Bush’s 302 
Cayenne Fi 309 
Coarse 306 
Coast 299 
Elegant 300 
Englemann’s 306 
Flat 300 
Globose 308 
Gray’s 309 
Hale’s 305 
Hall’s 304 
Houghton’s 301 
Lancaster 307 
Ow 298 
Marsh 302 
Michaux’s 305 
Nuttall’s 299 
Panicled 299 
Pine-barren 308 
Red-rooted 305 
Reflexed 307 
Rough 307 
Round-root 304 
Schweinitz’s 301 
Sheathed 303 
Shining 298 
Short-pointed 302 
Slender 309 
Straw-colored 306 
Toothed 303 
Yellow I 298, 301 
Cynthia 3 307 
Cypress 1 60,4; 2474 
ald I 64 
Belvedere 2s 


Black, Red, Swamp, 


Pond, Virginia or 
White I 64 
Broom 2 16 
Heath 46 
Juniper 60 
Standing 3 60 
Summer 2 16 
Cypress-moss I 46 
Cypress-vine 3 42 
Cypress-grass I 298 


CYRILLA FAMILY 2 485 


Daddy-nut 2 512 

Dagger-flower 3 435 

Daggers 540 

Daisies (Ohio) aad 
any Aster 


Dead-nettle 
Be gro yr 22,1025 


Hemp 3 119 
Henbit 121 
Death Camass I 492 
Deerberry 
2 693,73; 3 255 
Seedy 2 702 
Deer-food 2 76 
-grass 2 583 
-hair 328 
Deer-laurel 3 681 
-vine 276 
-wood I 607 
Deer’s-tongue 
1 506; 3 369 


Deil’s-spoons 1 
Devils, Blue 
Devil’s-apple 


2 130; 3 169 
-bit 1 489; 3 367 
-bite 1 494; 3 367 
-claws 2 115 
-club 2 619 


ENGLISH INDEX, INCLUDING POPULAR PLANT 


Devil’s-darning-needle 


122, 627 
-fig 2 139 
-flax 3 177 
-fingers I 590; 2 359 
-flower 2 69; 3 177 
-grandmother 3 354 
“grass I 133; 3 314 
"stip 2 35 
~gut 3°50 
-head-in-a-bush 
2 525 
-hair Dai 
-hop-vine I 528 
-iron-weed 3 320 
-milk 2 473-5 
-paint-brush 
j 3 329, 334 
-pitchforks 497 
-plague 2 625 
-rattle-box 2 64 
-root 3 235 
-shoestrings 2372 
-tether I 674 
-tongue 2571-2 
-trumpet 3 169 
-walking-stick 2 446 
-weed + 3 320 
Dewberry 
2 278-81, 790 
Dew-cup 2 264 
Dewdrop 282 
Dew-plant 2 203-4 
De Witt’s Snakeroot 
3 336 
Dewtry 3 169 
DIAPENSIA FAMILY 
2 705 
Dicksonia, Hairy 14 
Dill 2 634, 43 
Dillweed 3 516 
Dindle 316 
Dish-mustard 2 168 
Disporum I 517-8 
Ditaxis 2 456-7 
Ditch-bur 3 345 
-fern Le7, 
-grass r 88 
Dittany, Amer. 3 146 
Mountain 146 
Dock I 655-9 
Batter I 658; 3 532 
Bitter 658 
Bloody 658 
Blunt-leaved 658 
’ Broad-leaved 1 658 
Butter I 658 
Butterfly 3 532 
Can 2°78 
Clustered I 657 
Cuckold 3 547 
Curled I 657 
Dove 3 531 
Elf 3 457 
Fiddle I 658 
Flea 3 532 
Garden-patience 
I 656 
Golden I 659 
Great Water 656 
Kettle 3 542 
Mullen 173 
Narrow 1 657 
Pale I 655-6 
Passion 656 
Patience I 656 
Peach-leaved 656 
Prairie 3 462,5 
Red-veined I 657 
Sharp 654 
Smaller Green I 657 
Sour 653-7,9 


NAMES. 609 

Dock, Swamp ‘it 6 
Tall 6G 
Tubercled 655 
Veined 654 
Velvet 3 173 


Water I 445, 656 


Western 657 - 
White 655 
Willow-leaved 655 
Winged 654 
Yellow 657; £1608 
Dock-cress, Succory 
: 3 306 
Dockmakie 270 
DopvEeR FamMity 3 48 
Dodder 3 48-52 
American 2 
Beaked 3 51 
Button-bush 50 
Clover 49 
Compact 52 
Cuspidate 51 
Field 49 
Flax 48 
Glomerate 52 
Gronovius’ 51 
Hazel 50 
Lesser 49 
Love-vine 49, 51 
Lucerne 49 
Pretty 3 50 
Smart-weed 49 
Thyme 49 
DocBaNnE FAMILY 3 19 
Dogbane 
Bitter 3-25 
Clasping-leaved 3 22 
Intermediate BNoy 
St. Johns 3 22 
Spreading 3 21 
Velvet Rvs 
Dogberry 
I 544; 2 239, 87, 90 
-blow 3 518 
-bramble 2 289 
-tree 2 291, 662 
Dog-bur 3 53-4, 76 
-daisy 3 516 
-dandelion 3 310 
Dog-fennel 
3 56, 430, 513, 6 
False 513 
Dog-finkle 3 516 
-grass I 283, 382 
-laurel 2 688 
-lily 278 
-mint 3 138 
-nettle 3 119 
-plum I 515 
-poison 2 645 
-rose 284 
-thorn 2 284 
-toes 3 45! 
Doggers 170 
Dog’s-dinner 2 542 
-finger 3 204 
-mouth 3 178 
-parsley 2 630 
-rib 3 240 
-tansy 2 258 
-tongue 75-6, 309 
Dogs and Cats’, (2355 
Dog’s- tooth Violet I 506 
Yellow 506 
White 506 
Dog-weed 3 513 
DoGwoop FAMILY 2 660 
Dogwood 2 328, 503, 
661-3; 3 167 


(see Cornel 2 661-5) 
Alder-leaved 661 
American 664 


610 


Dogwood, Black 2 504 


Blue 663 
European 2 492 
False 2 497 
Florida 664 


Flowering 2 292, 664 


Green Osier 663 
Poison 2 483 
Pond 3 265 
Purple 2 663 
Red Osier 2 661-2 
Striped 497 


. Swamp 2 445, 83, 661 
White 2 663; 3 270 
White-fruited 2 663 

Dollar-leaf 2 397, 668 

Doob 222 

Door-grass,-weed I 661 


Double-claw 3 239 
-leaf I 568 
-tooth 3 495 

Dove-dock Sys5or 

Dove’s-foot 2 429 

Down-hill-of-Life 712 

Down-Ling I 325 


Down-weed 3 447 
Draba, see Whitlow- 


grass 2 159-61 
Dragon 
Brown I 442 
Female or Water 
285, 444 


Dragon-head 3 115-6 


Dragon-root 443 
Dragon’s blood 2 426 
-claw I 575 
-mouth I 562; 3 178 
-tongue 2 672 
Drake I 218 
Drawke 282 
Drew gI 


Drooping Avens 2 bee 
Starwort 489 
Drop-flower 3 334 
-seed 1 184-9, 177-200 
Dropsy-plant 2) cys 
Dropwort, Water 2 638 


Western 248 
Drumhead 448 
Drunk 282 
Drunkards 2 85, 693 
Dryas 273 
Duck-acorn 77 
Duckretten I 494 
Ducks I 549-50 

-foot 2 264 

-meat 448 
DuCKWEED FAMILY 

I 446 
Duckweed I 446-9 

Gibbous 448 

Greater 446 

Ivy-leaved 447 

Least 447 

Lesser 448 

Minute 448 

Pointed 449 

Star 447 

Valdivia 447 
Dummy-weed 531 
Dupontia 261 
Dusele 164 
Dusty-clover 407 

-husband 


Dusty Miller 
Dutch-beech 


NYNNHWDN NW HW 
aS 
° 
N 


Cheeses 514 
Flax 157 
Pink 200 
Dutchman’s-breeches 
; 2 142 
-pipe I 646; 2 674 


Dutchman’s-rush 


I 41 
Dutch-morgan 3 518 
Dye-leaves 2 487, 721 
Dyer’s-broom Beare 
-cleavers 3 263-4 
-Green-weed 2 350 
-mignonette 200 
-rocket 200 
-weed 2 350; 
3 200, 391, 5 
-whin 2 350 
Dye-weed 350 
Dyschoriste 3 240 
Dysentery-root, -weed 
3 78 
Ear-jewel 2 44 
Earning-grass 3 226 
Earth-almond 304 
-apple 3 486 
-club 235 
-gall I 494; 3 2 
-smoke 2 146 
Easter-bell 2 44 
-flower 102 
-lily 1532 
Eatonia 243-4 
Esony FAMILY 2.720 
Eclipta 3 468 
Eel-grass I gi, 106 
Green 26 
-pondweed TOT 
Egg-plant 2 276 
Eggs-and-Bacon 177 
Eglantine 2 286 
Egriot 327, 
Elder 2 268-70 
American 268 
Bishop’s 2 654 
Box 498 
Common 268 
Dog 2 654 
Dwarf 2 618, 54 
Horse 3 457 
Marsh 270, 339-40 
Mountain 268 
Poison 2 483 
Prickly 2 617 
Red 2 270 
Red-berried 268 
Rose 270 
Sweet 268 
‘Water 270 
Wild 2 618 
Elder-berry 268 
-blow 268 
Eldin 3 532 
Elecampane 457 
Election-posies 3 214 
Elephant’s-foot 3 354 
-trunk 239 


Eleven-o’clock-Lady 


I 506 
Elfdock 3 457 
Elfwort 3 457 
Elk-bark 281 
-nut I 641 
Elk-tree 2 692 
-wood 2 82 
Eller I 613 
ELM FAaMILy I 625 
Elm I 626-8 
American 626 
Bastard 629 
Cliff 626 
Cork 626-7 
Corky White 626 
False 629 
Hickory 626 
Indian 627 


Moose or Red 1 aoe | 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Elm, Racemed_ 1 626 
Rock 626-7 
Slippery I 627 
Sweet 627 
Swamp 626 
Water I 626-8 
White 626 
Winged 627 
Witch 627 

Elsholtzia ~ Bers 4! 

Emetic-weed 3 303 


Enchanter’s-Night- 
shade 2 611 

Enchanter’s-plant 3 95 

English Bull’s-eye 


: 3 470 
English Mercury 2 14 
Enslen’s Vine 3 ye 
Equisetum 

Smooth, Sedge- like 
42 
Variegated 42 
Erigeron 3 435 
Arctic 436 
Hoary 438 
Hyssop-leaved 438 
Low 437 
Rough 436 
Three-nerved ~ 437 
Tufted 439 
White-top 440 


Eriogonum I 647-52 


Allen’s 649 
Annual 651 
Branched 650 
Crisp-leaved 651 
Effuse 652 
Few-flowered 650 
James’ 649 
Long-leaved 648 
Long-rooted 650 
Narrow-leaved 651 
Nodding 652 
Slender 652 
Winged 648 
Yellow 649 
Ernfern Te 32 
Eryngo 2 622-3 
Water 622 
Virginia 622 
Erysimum 2 172 
Eubotrys, Mountain 688 
Eulophus 2 654 


Eurotia, American 2 20 


Eustoma 37-8 
Evans’-root 2 272 
Evening Lychnis 2 68 


EvENING-PRIMROSE 


FAMILY 2 584 
Evening-Primrose 

594-9 

Common 2 595 

Cut-leaved 598 
Dwarf or Golden- 

flowered 2 601 

Gray-leaved 2 599 


Narrow-leaved 596 


Northern 595 
Nuttall’s 599 
Oakes’ 2 596 
Prairie 598 
Rhombic 597 
Seaside 597 
Small-flowered 594 
White Shrubby 599 
White-stemmed 599 
Evening Trumpet- 
Flower 2 730 
Everfern f 83 
Evergreen 2 207 
Christmas I 49 
Trailing I 44 


Vo. IIT. 
Everlasting 2 207,18 
Ipine 3 449 
Carpathian 449 
Clammy 454 
Heller’s 454 
Early 451 
Large-flowered 453 
Life 453,4 
Fragrant, Sweet, 
454 
Low 453 
Mountain 452 
Mouse-ear 451 
Pearly 451, 3 


Plantain-leaf 3 451 
Spring 451 
Everlasting-grass 1 124 


-pea 2 413,4 

Sea-side 2 413 
Eve’s Darning-needle 

I 513 

Eve’s-thread I 496 

Evolvulus 3 217-8 

Ewe-gowan 3 402 


Eye-balm, or -root 2 85 


Eyebright 

2674; 3251, 716 
Spotted 2 467 
Eyeseed 3 130 
Eyewort 3 218 
Fabes or Fapes 2 241 

Fair-Maid of France 
515 
Fairy Bells 3 204 
-cap 204 
-cheeses 2V505 
-circles I 66 
-creepers 2 143 
-cup 229 
-fingers, -glove 3 204 
-grass I 250 
-smoke 2 674 
-thimbles, weed 3 204 
Fairies’ Horse 542 
Faitour’s-grass 2 473 
Fall Dandelion 3 310 
-flower 427 
-meadow-rue 2 121 
False-alder 2 489 
-aloe I 534 
-banana 2 83 
-Beech-drops 2 675 
-bindweed 3 43 
-bittersweet 2 493 
-boneset 364 
-box, boxwood 664 
-buckwheat I 674-5 
-bugbane 2 103 
-Calais 3 324 
-Choke-dog 36 
-Colt’s-foot I 642 
-Couch-grass 283 
-Dandelion 3 324-5 
-Dog-fennel 3 513 
-Dragon-head 3 116 
-elm I 629 
-flax 2 164,157; 3515 
-foxglove 3 206-8 


-garlic, yellow 1 501 
-Goat’s-beard 2 215 


-Golden-rod 3 401 
-grape 2 510 
-gromwell 3 90-1 
-heather 2 542 
-hellebore I 495 
False Indigo 2 344 
ue 344 
Dwarf 2 366 
Fragrant 2 366 
White 346 
Yellowish 2 345 


Vor, IIT. 

False Johnswort 2 536 
-lettuce gescr 
-lily-of-the-valley 

517, 
-loosestrife 2 585,7 
-lupine 2 344 
-mahogany 233 

False-mallow 2 519-20 
-mallows 521 

Fatse MERMAID 

FAMILY 2 441 

False-mermaid 441 
-Mitrewort 224 
-mustard 199 
-nettle Tos7 
-Oat 207, 
-Pennyroyal 3 104 
-Pimpernel 196-7 
-Rue-anemone 2 89 
-Sanicle 229 
-Sarsaparilla 618 


Solomon’s-seal 1 516 
-sunflower 3 467, 510 


-Sweet-flag I 540 
-sycamore 2 242 
Unicorn-plant 1 489 
-valerian 3 544 
-Varnish-tree 2 446 
-wheat I 283 
-wintergreen 2 668 
Fame-flower 2 36 
Fancy 563 


Farewell Summer 
3 426, 430 
2 698 


Farkleberry 

Fat Hen 2 10, 14-18 

Fayberry 2 241 

Feaberry 241 

Featherfew 3 519 

Feather-fleece 
Mountain I 490 


Featherfoil,Am. 2 710 
Feather-Geranium 2 14 


-Bunch-grass I 176 
-grass 177,214, 220 
-leaf (Cedar) 65 
-weed ° 3 454 
Felon-grass 2 638 
-herb 3 333, 527 
-weed 542 
=WOret, 2) 6383.3 137 
Felwort 39,14 
Feltwort B8173 
Fenberry 2 704 
Fennel 643 
Dog 3 35-6, 
430, 513-16 

Hog’s 2 638; 3 516 
Water 477 


FERN FAMILIES I 1-34 
Fern 


Adder’s I 2,36 
-spear 2 
-spit 32 
-tongue Te 

Alpine, Rough 16 

Backache, -brake 30 

Basket 21 

Bear’s-pawroot 21 

Beech Ti23 
Broad 23 
Common 23 
Hexagon 23 

Long 23 

Bladder, Brittle 1 15 

Bottle Tons 

Boulder- 14 

Brake, Bracken 

I 32-3 

Lady 32 
Sweet, Knotty 21 
Bristle 18 


Fern, Brittle Ts 
Buckhorn (Brake) 71 


Bulblet 15 
Chain (Netted) 1 25 
Virginia 24 
Christmas 16 
-shield 16 
Cinnamon Taz 
Clayton’s 8 
Cliff-brake 32-3 
Clayton’s 33 
Oregon 33 
Climbing 9 
Clinton’s 20 
Common Wood = 22 
Crested 19 
Creeping I 9 
Curly-grass 9 
Cystopteris 15 
Dicksonia, Hairy 14 
Dickson’s 14 
Ditch 7 
Dugger-grass ae 
Eagle 32 
Ernfern 32 
Everfern £83 
Female I 30 
Fiddle-heads 7 
Filmy 8 
Fine-haired 14 
Flower-cup 12 
Flowering- 18 
Clayton’s 8 
Interrupted 8 
Goldie’s 20 
Golden Palescian 36 
Golden Locks 36 
Grape I 3-6 
see Fern 
Hair f 72 
Hare’s-foot 18 
Hartford 9 
Hart’s Tongue 25 
Hay-scented 14 
Hog-brake 32 
Holly 16 
Braun’s 17 
Indian’s Dream 33 
King’s 7 
Knotty-brake 21 
Lady I 30 
Lady Bracken 32 
Lady’s-hair 31 
Lip, Alabama 34 
Clothed 34 
Hairy 34 
Webby 35 
Woolly 35 
Lock-hair an 
Maidenhair Tsai 
Black, “ True,” 31 
English 28 
Golden 36 
White 29 
Maidenhair Spleen- 
wort I 28 
Male 1/30 
Male Shield a 
Marsh 
Meadow 584- te 
Moon i3 
Moonwort T 5,6 
Hemlock- ieaved 6 
Moss rt 36 
Mountain 15 
New York 18 
Oak I 23 
Scented 24 
Osmond-the-Water- 
man ios 
Ostrich Tre 
Polypody I 36 


ENGLISH INDEX, INCLUDING POPULAR PLANT 


Fern, Polypody, Gol- 
den, Common 1 36 
Male 36 
Pale Mountain 


23 
Hoary, Scaly, 

ree, 36 

Tender Three- 
branched 23 
Prickly-toothed 21 
Broad 21 
Quill TS 
Rattlesnake I 6 
Regal 7 
Resurrection 36 
Rock 72 
Rockbrake 1 32-3, 36 
American 32 
Rough Alpine I 16 
Royal I 7 
Royal-Osmond 7 
Rue 29 
Sea-weed 25 
Sensitive i Gui 


Serpent’s tongue 2 


Shield (or Wood) 
I 18-22 
American 22 
Boott’s 22 
Broad Prickley- 
toothed 1? 215 
Narrow Prickley- 
toothed ea: 
Crested 19 
Dodge’s 19 
Eaton’s 17 
Fragrant 19 
Marginal 20 
ale E20 
Massachusetts f 39 
Marsh 18 
Prickly 21 
Prickly-toothed 21 
Spinulose 21 
Spreading 21 
Shield-roots | 21 
Shrubby 586 
Snake yas 
Snake’s-tongue 2 
Spleenwort 1 26-30 
Bradley’s 30 
Common Wall 28 
Dwarf I 28-9 
Ebony Tey, 
Green 28 
Little Ebony 27 
Maiden-hair 1 28 
Mountain 29 
Narrow-leaved 28 
Pinnatifid 27 
Rock (Smooth) 29 
Scott’s 26 
Silvery 30 
Small 2 
Swamp 28 
Wall Rue 2 
Stone, Stone-rue 29 
Stonebrake 36 
Sun 23 
Swamp 18 
Sweet I 36, 586 
Sweet-brake 21 
Tentwort 29 
Tree 7 
Venus’-hair t 32 
Walking 26 
Walking-leaf 26 
Wall I 28 
Wall Rue I 29 
Water 7 
Water-wort 28 
Wilson’s 15 | 
Windsor 9 | 


NAMES. 611 


Fern, Winter Brake 1 33 


Wood 19, 36 
Evergreen I 20 
Goldie’s 20 

Woodsia I 12-14 
Alpine 12 
Blunt-lobed 14 
Northern 12 
Oblong I I2 
Oregon 13 
Rays 12 
Rocky Mt. Cig: 
Rusty Ta 
Smooth 13 

Fern-bush I 586 
Fern-gale 584-6 
Fescue 261, 269-73 
Festoon-pine 48-9 
Fetid-shrub 
2 83, 269- 
Fetter-bush 2 ae 
Fetticus 3 286 
Fever-bush 2 135; 489 
-cup 232 
Rec ceiey 2 266; . 519: 
American 3 465 
Common 519 
Fever-plant 2 585 
Fever-root 3 275 

SOwiE 2 493 

-weed 3 206; 95-6 

-wort 3 275 

Fiddle-grass 2 591 
Field-balm I 139 
-pine 2 541 
-sorrel I 653 
Fig, Indian 257i 
Fighting-cocks 548 
Ficwort FAMILY 3 172 
Figwort, Hare 180 
Maryland 179 
Western 3 180 
Filago 3 446 
Filerie 2 430 
Filbert I 607 
Firmy Fern FAMILY 
18 
Finger-berry 2 280 
-grass Tae 
Finkel 2 643; 3 516 
Fiorin I 203 
Fir I 5, 63 

Amer. Silver 63 

Balm-of-Gilead 63 

Balsam 63 

-moss unas 

Fir-pine or -tree 63 
Fir-rope 2 675 
Fire-balls 2 69 

-grass 2 264 

-leaves 3 246 

-Pink 2 64 

-top 2 590 

-weed 2590; 3169, 

246, 319, 442, 536 
Fish-mouth 3 181 
-poison 2 500 
-wood 491 
Fit-root plant 2 674 
Five-finger 2 251-8 

Dwarf 2 251 

Marsh 258 

Mountain 262 

Five-Sisters 711 
Flag I 446, 537-40 

Corn 540 

3lue I 537-9 

False Sweet 540 

Myrtle 446 

Narrow 539 

Red-brown I 539 

Slender Blue 541 


612 


Flag, Sweet I 446 
Sword, Yellow 540 
Water 537 
Yellow Water 540 

Flag-lily 537 

Flagons 540 

Flagroot 446 
Poison I 539 

Flamy 2 563 

Flannel, Adam’s 3 173 
-leaf 173 

Flat-top 3 354 

FLax FAMILY 2 435 

Flax 2 437-40 
Berlandier’s 439 
Cathartic 438 
Cultivated 435 
Devil’s 25177, 
Dwarf, Fairy 2 438 
Dutch 157 
False 2 157-64 


Florida Yellow 2 438 
Grooved Yellow 439 


Large-flowered 439 
Lewis’ Wild 437 
Mountain 438, 451 
Prairie 436 
Purging 2 438 
Ridged Yellow 437 
Slender 437 
Spurge 2 574 


5 
Stiff Yellow 2 438 
Wild 3177; 2436-7 


Yellow 2 438-9 
Flax-tail I 68,170 

-weed BeL77 
Fleabane 


Bitter, or Blue 3 441 


Canada 442 
Daisy 437, 439, 440 

Western 3 440 
Dwarf 442 
Early 441 
Foetid 447 
Marsh 447-8 
Philadelphia 439 
Running 438 
Small 416 
Spreading 439 
Three-nerved 437 


Fleaweed, Horse 2 345 
Fleawort 3 258 

Marsh 540 
Fleur-de-lis I 536,539 
Flixweed 2 194 


Floating Heart 3 18-9 
Floramor 22 
Flora’s Paint Brush 


334 
Florida Moss I 456 
Flower-de-luce 536 


Flower-gentle Baz 
Flower-of-an-Hour 


2 525 
-of-Gold (any 
Solidago) 3 380-98 


Flowering-rush 451 
Fluellin 3 75-6, 201 
Female 175 
Sharp-pointed 175 
Flux-root 25 


-weed 2 170; 3 104 
Fly-honeysuckle 


3 281-3 

Garden 282 
Fly-poison I 489 
-trap 2 202; 321 
Foal-foot a) ak 
Foam-flower 2 224 
Fog-fruit 3 98 
Folk’s-glove 204 
Fool-hay 139, 207 


Fool’s Parsley 2 645 
Forefather’s-cup, or 


-pitcher 2 202 
Forget-me-not 
3 85-7, 201 
Smaller 25 
White 3 80-2 
Wild 2e50 
Fothergilla 2 234 
Four-0’cLock FAMILY 
2 30 
Four-o’clock 595 
Four-toes 451,595 
Fox-berry 2 693; 3 285 
-feet I 43 
-geranium 2 426 
Foxglove 
2 202; 3 204, 237 
False 3 206-8 
Lousewort 221-9 
Mullen 3 205 
Purple 204 
Yellow 206 
Fox-grape 2 506-8 
-plum 2 693 
-tail I 47 
see Grass 
French-grass 2 365 
-Pinks 2 66; 3 557 
Dwarf 2 66 
Fringe 
Alleghany 2 143 
American 728 
Mountain 2 143 
White 728 
Wood 2 142 
Fringe-Cup 239 
Fringe-tree’ 2728 
American 728 
Fringed-Gentian 250 


Fringed-Orchis 1 558-9 


Greenish 558 
Pink 559 
Ragged 550, 558 
Fritillaria I 505 
Frog-fruit 3 98 
-grass. I 469; 2) 21 
-leaf 2 76 
-plant 2 207 
-wort 2102 
FrRoG’s-BIT FAMILY 
I 106 
Frog’s-bit I 107 


-bladder, mouth 2 207 
Frost-blite 200 
Frost-flowers (any 


Aster) 
-weed 
2 540-1; 3 430 
any Aster 
Frostwort 2 540 


Frost, It-brings-the-, 
(Indian name of 
the Aster) 

Fuller’s-herb 273 

-teasel 3 2890 
Fumitory, Bulbous 283 
Climbing 2 143 
Hedge 146 

Furze 2 349 

Fustic-tree 2 343 

Fuzzy-guzzy 3 454 

Gadrise 3 270 

Gaertneria 3 343 

Gagroot 303 

Gaillardia 511-13 


Gaiter-tree 

2 492; 3 270, 662 
Galax, Galaxy 2 709 
Gale, Fern — 584-6 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Gale, Sweet, or 


Scotch I 584 
Galingale, or Galangal 
Edibl — 
ible 304 
Lank 306 
Galinsoga 3 501-2 
Gallberry 2 487 
Gall-of-the-Earth 
3 10, 
Gallon 3 ae 
Gall-weed 3 10 
-wort 3 L777. 
Gama sO tia 
Garb-willow I 595 
Garden-gate 2 563 
-plague 2 654 
Gardener’s-eye Bri 
-delight ; 71 
Garget 2 26 
Garlic I 499 
Chive 497 
Crow, Field 499 
False 501 
Hedge 2 170 
Keeled 499 
Meadow I 499 
Rush 498 
Wild I 499 
Y ellow-false 501 


Garlic-mustard 2 170 
-wort 170 
Garnet-berry 22a. 
Gaskins 2e2Knte ey, 
Gatten 2 492; 3 270 
Gatteridge 2 492 
Gaura 2 608-10 
Gay-feather 3 367-8 
Gay-wings 2 452 
Gean 327 
Gem-fruit 2 224 
Genson gu275 
GENTIAN FAMILY 3 I 
Gentian 3 8-14 
Barrel 2) 48) 
Bastard 39 
Blind Te 
Blue 3 12,104 
Bottle 3 12 
Closed 12 
Downy B0rE 
Elliott’s II 
Five-flowered 3 10 
Four-parted 3 10 
Fringed 39 
Horse 3.275 
Marsh Quire e 
Narrow-leaved 3 13 
Northern 39 


Oblong-leaved II 


One-flowered 14 
Red-stemmed 3 13 
Rough 3 12 
Small 3 14 


“Smaller Fringed 2 9 


Snake 3 335 
Soapwort 3 12 
Spurred 15 
Stiff 10 
Straw-colored 13 
Striped 3nus 
Swollen 38 
White 3 275 
Yellow 15 
Yellowish Bone 


Gentleman’sCane 1,670 
GERANIUM FAMILY 


2 425 

Geranium 2 426-9 
See Crane’s-bill 

Feather 2 14 

Fox, Wild 2 426 


Voz. III. 
Geranium 

Meadow 2 427 
Mint 3 579 
Mountain 426 
Turnpike 2 14 
Wood 2 429 
Gerardia 3 208-14 
Auricled 213 
Bessey’s 211 
Bushy 206 
Cut-leaved 214 
Fascicled 210 
Flax-leaved 209 
Gattinger’s 213 
Large Purple 210 
Prairie 209 
Rough Purple 209 
Salt-marsh 211 
Sea-side 2Ir 
Skinner’s 212 
Slender 211 
Small-flowered 210 
Ten-lobed 212 


Thread-leaved 212 


Cut-leaved 103-4 
Hairy 103 
Wood 103 
Germander Chickweed 
3 203 

Ghost-flower 2 674 


Giant Hyssop 3 111-12 


Gibbals 247 
Gifola 3 446-7 
Gilia 3 59-62 
Low 61 
Scarlet 60 
White 60 
Small 60 
Spicate 61 
Tufted 62 


Gill, Gill-ale 3 114 
Gill-go-by-the-Ground 


3°11 
Gill-over-the-Ground 


3 114 

Gilliflower 
Cuckoo 2 69 
Dame’s 2 196 
Marsh 2 69 


Mock 72 
Night-scented 2 175 
Queen’s, Rogue’s 175 
Sea 718 
Winter 2 175 
Gilliflower-grass 415,39 


Gill-run-over 3 114 
Gilt-cups 2 112 
Ginger I 642-3 
2 208-9; 3 531 

Green 3 527 
Indian I 642 


Southern Wild 1 644 


Wild 1 225, 642, 646 
Ginger-berry 
I 642; 2 693 
Ginger-plant Zune2 
-root : 531 
GINSENG FAMILY 2 616 
Ginseng 618 
Blue 2 128 
Dwarf 618 
Horse 2 275 
Gipsy-flower 3 76 
-herb 148 
-rose 290 
-weed 201 
-wort 3 147 
Gipsy-Combs 3 289 
Girasole 486 
Glasswort 2 21-2 
Jointed 21 


Vo. III. 
Glasswort 
Prickly 2 25 
Glen-pepper 2 164 
-weed 164 
Glidewort 3 119 
Globe-flower 
_ 2 87; 3 255 
Spreading 2 87 
wamp 87 
Globe-mallow 522 
Maple-leaved 522 
Sharp-fruited 2 522 
Gloden 3 478 
Glory-less 283 
Goat’s-beard 2 247 
Dwarf 3 308 
False 2 215 
Virginia 3 307 
Yellow, Purple 313 
Goat-foot 2 654 
Goat’s-rue 372-3 
God’s-eye 3 201 
Goggles 2 85, 241 
Gold 347 
Gold-and-silver-plant 
2 191 
Gold-balls 2 113 
Gold-chain 2 208 
_ Gold-cups 112 
-flower 373 
-knops III-I113 
-thread 2 88 
-weed 2 115 


Gold-of-pleasure 2 157 
Golden Alexanders 


2 639-41 

Golden Aster 3 373-5 
Cottony 373 
Grass-leaved 373 
Hairy 374 
Hispid B75 
Maryland 374 
Nuttall’s a5 
Sickle-leaved 373 
Stiff-leaved 374 
Golden-bush 376 
-club 445 
-coreopsis 3 493 
-corydalis 2 145 
-cup 117 
-currant 2 239 
-flower 3 518, 539 
Ground Bbs73 
-glow 3 473 
-guineas 2 117 
-Jerusalem 3 470-1 
-knops 2 012 
-meadow-parsnip 640 
-motherwort 3 456 
-moss 2 208 
-oak 3 208 
-osier I 584 
Goldenpert 3 195 
- Golden Ragwort 544 


Golden Rod 3 380-398 


Alpine 385 
Anise-scented 389 
Beach 389 
Blue-stemmed 3 382 

Mountain 389 
Bog 387 
Broad-leaved 383 
Boott’s 3901 
Bushy 398-400 
Canada 393 
Cutler’s 385 
Curtis’ 383 
Cut-leaved 393 
Double 395 
Downy 3 386 


Downy Ragged 
Drummond's 


Golden Rod 
Dwarf 3 395 
Early 393 
Elliott’s ae 


Elm-leaved 
European (Rand’ Se 


388 
False 401 
Few-rayed 3 392 
Field 395 
Flat-top 398 
Fragrant 3 398 
Gattinger’s 305 
Gilman’s 388 
Gray 395 
Ground 396 
Hairy 3 384 
Hard-leaved 397 
High . 395 
Houghton’s 3 398 
Large-leaved 385 
Late 3 394 
Lindheimer’s 385 
Minaret 386 
Missouri 3 384 
Mountain 384 
Noble 387 
Northern, Ohio 
286, 397 
Pale 383 
Pine Barren 390 
Plume 393 
Pyramid 390, 392-4 
Ragged 3 382 
Rand’s 388 
Rayless 375-6 
Fetid 376 
Riddell’s 397 
River-bank 388 
Rock 393 
Rough-leaved 391 
Salt-marsh 3890 
Sea-side 389 
Shadowy 389 
Sharp-toothed 393 
Showy 3 387 
Short’s 394 
Slender 3 384 
-fragrant 400 
-showy 387 
Small- headed 399 
Spreading 391 
Stiff 397 
Stout Ragged 382 
Swamp 3 387,392 
Sweet 389 
Tall 395 
Tall Hairy 390 
True 389 
Twisted-leaf 3 390 
Velvety 396 
Viscid 309 
Wand-like 386 
Western Rough 396 
Bushy 399 
White 383 
Willow-leaf 386 
Woodland 382 
Wreath 382 
Wrinkle-leaved 390 
Yellow-top 380-393 
Zig-zag 883 
Golden- pasiiraud 2 230 
-seal T Suse) 205 


-star, Maryland 3 374 
-trefoil 2 101 
-willow-herb 711 
Good-bye-summer (pur- 
ple asters, N.C.) 

Good King Henry 2 14 

Good Morning Spring 
2 37 


ENGLISH INDEX, INCLUDING POPULAR PLANT. NAMES. 


Gools 2 85 
Good Friday Grass 485 
GOOSEBERRY FAMILY 


2 236 

Gooseberry 697-8 
Bristly 2 240 
Cape el Tie, Bile 


Eastern Wild 2 241 


Garden 241 
Hawthorn 240 
Illinois 240 
Low wild 241 
Missouri 2 240 
Northern 240 
Prickly Wild 2 239 
Slender 240 
Smooth 240-1 
Swamp 2 236 
Wild 239 
Gooseberry-pie 2 591 


GOOSEBERRY FAMILY 


28 
Goosefoot 295 
Bosc’s 12 
City 12 
Cut-leaved 15 
Feather Geranium 14 
Fremont’s 12 
Good King Henry 14 
Jerusalem Oak 14 
Many-seeded II 
Maple-leaved 13 
Mealy 2 10 
Mercury 14 
Narrow-leaved II 
Nettle-leaved 13 
Mexican Tea 15 
Oak-leaved bao) 
Perennial 14 
Red 13 
Sea 23 
- Stinking 2 10 
Turnpike Geranium 
2 14 
Upright 12 
White 2 10 


Goose-grass I 220, 661; 
2 258; 3 259, 428 
Great 3 84 


Lesser 259 
Goose-tansy 2 258 
-tongue 3 137; 515 
Gopher-wood 2 343 
-berry 696 
Gordoldo 2.515 
Gorse 2 349 
Gosling 102 
Gosling-grass 3 259 
-weed 259 
Gosmore 309 
Go-to-bed-at-noon 313 


GourD FAMILY 3 
Gourd, Missouri 


290 
291 


Gout-weed, -wort 2 654 
Gowan, Ewe 3 402 
Horse 521 
Ling 328 
May 402 
Meadow 2 8&5 


Milk or Witch 3 315 

Open or Water 2 85 
Yellow 

2 IlI-33 

Grama, see Grass 


3 315 


GRAPE FAMILY 2 505 

Grape 
Arroyo 2 507 
Ashy 507 
Bear’s 693 | 
Blue 507 
Bull 507 | 
Bullace 509 | 


613 


Grape, Catawba 2 506 


Chicken 508 
Concord 506 
Downy 507 
False 511 
Fox 2 506, 508-9 
Frost 507-8 
Isabella 2 506 
Missouri 508 
Mountain 508 
Muscadine 509 
Oregon 2 128 
Plum 506 
Possum 508 
Rocky Mountain 
2 128 
Riverside 507 
Salt 2 25 
Sand 508 
Scuppernong 509 
ea aor 
English 21 
Small 2 506 
Southern Fox 509 
Sugar 508 
Summer 506 
Sweet-scented 507 
Winter 2 507-8 
Grape-fern, Little 1 3 
Cut-leaved 5 
Eaton’s 4 
Hitchcock’s 3 
Lance-leaved 6 
Leathery 6 
Rattlesnake 6 
Ternate 5 
Virginia 6 
Wood’s 4 
Grape-flower 510-1 
-hyacinth 1 510-11 
-root 2 128 
Grapewort 2 90 
GRASS FAMILY 
I 107-295 
Grass [of the Andes] 
220 
Ague 511 
Alkali 250 
Aristida I 179-80 
Arrow I 92, 182 
Bahama 222 
Barley I 286-7 
Little 286-7 
Squirrel-tail 287 
Wall 287 
Barnacle gI 
Barn I 133 
Barnyard 133 
Beach 212, 249 
Bear 1167,512; a oie 
Beard 1 113-9,7 76 , 201 
Annual 201 
Broad-leav Ap 
I 226 
Broom 115 
Bushy 116 
Forked 118 
Indian 116 
-joint I 150 
Naked 126 
Purple 180 
Sea-coast 115 
Short-leaved 226 
Silvery 117 
Torrey’s 119 
Triple-awned 180 
Woolly 113 
Beekman’s 228 
Bengal f 394 
Bennet-weed I 192 
Bent I 203, 2:3 
Black 141 


614 


Grass,Bent,BrownI 205 


Creeping 203 
Dense-flow’d I 203 
Dog 205 
Elliott’s 204 
Marsh 203 
Narrow 290 
New England 207 
urple 213 
Red 205 
Reed 209, 211-i2 
Rhode Island 206 
Rock 204 
Rough-leaved 204 
Sea I 363 
Sea-shore 212 
Silky 213 
Spider 204 
Tall 206 
Twin 205 
Upland 206 
Way 287 
White 203 
Wind s 213 
Bermuda 222 
Big Blue-stem 118 
Bird 255, 661 
-seed 170 
Bitter 511 
Black 2 351; 
I 275,470 
-bent 141 
-couch I 192 
Black-head 485 
Black-oat I 177 
-quitch 203 
Black-seed 197 
Blow-out 
177, 236, 242 
Blubber I 278 
Blue I 397, 543 
Colorado 283 
English 259, 81 
Kentucky 256 
Oregon 260 
Texas 259 
Blue-eyed 
I 251, 543-5 
Blue-joint 
I 115, 118, 209, 283 
Big 115 
Northern 209 
Blue-stem 115, 209 
Big 118 
Bushy 116, 120 
Little Tens 
Bonnet 203 
Bottle 2 165, 355 
Bottle-brush 294 
Bottle-rush 294 
Bowel-hive 2 264 
Bride’s-laces I 169 


Bristle, Green 165 


Glaucous 165 
Rough 164 
Brome I 274-280 
Awnless 280 
Briza-like I 280 
Corn 279 
Downy 274 
Field 279 
Fringed 276 
Hairy 275-6 
Hungarian 277 
Rescue 280 
Smooth-rye 218 
Short-awned 280 
Upright 276 
Wild 280 
Brook 116 
Broom 115, 118 


Grass, Broom-corn, 
Wild Le232 
Buck 43 
Buftalo I 227n 2a 
Bull 223, 278 
Bull-poll 215 
Bullpates 215 
Bunch I 15, 174, 

177, 199, 290 
Buffalo 373 
Early 244 
Feather 176 
Greater Tez 
Bur 167 
Small 167 
Burdock 121 
Button 220 
Burden’s 203 
Calf-kill 214 
Canary 170 
Reed 170 
Southern 170 
Wild 170 
Candy 241 
Cane fa222 
Maiden 12h, 137 
Switch 2905 
Cane-brake 2905 
Capon’s-tail 269 
Carnation 
I 397, 415, 428, 560 
Carpet xs 
Catch-fly 169 
Catstail 190-1 
Alpine IgI 
Meadow IgI 
Rush 190 
Causeway 253 
Cheat I 276, 282 
Pine 2 59 
Chess IL 275, 253 
Compact 275 
Field I 279 
Kalm’s 277 
Soft 278 
Southern 280 
Swamp 276 
Wild 277 
Wood 276 
Claver 3 259 
Clump-head £9337. 
Cock 278 
Cocksfoot 251 
Cockspur Ee123 
Coe 469 
Coco 304 
Corn 162, 213 
Cord- I 223-4 
Fresh-water 223 
Slender 224 
Cotton- I 322-6 
Hare’s-tail 1 324 
Couch 203, 283 
Black 192 
False I 283 
Cow 1661; 2355-6 
Cow-quake I 256 
Crab Tees 
2 21, 229, 661 
Diffuse I 124 
Flat 125 
Fringed, Large 123 
Little ae 
Small 123 
Sprouting 138 
Texas 226 
Creek I 281 
Creek-stuff 223-4 
-thatch 223 
Crested-hair 245 
Crop 229 
Croup 266 | 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Grass 


Crowfoot I 123, 229 


Cuckoo 485 
Citba I2I 
Curly b ame) 
Cut, Rice 169 
Cypress 298 
Dare 281 
Darnel I 281 
Bearded, Poison, 
282 
Red 281 
Dart 214 
Deer 2 583 
Deer-tongue 162 
Dennet I 293 
Devil’s 3 133, 314 
Dew I 203, 251 


Burden’s, Monkey’s 
orSummer I 203 


Ditch I 88 
Dithering 250 
Dodder I 250 
Dog 283, 382 
Dog-bent 205 
Dog-wheat 285 


Dog’s-tail 1 229-252 

Crested 252 
Dog’s-tooth 222, 285 
Doob 222 
Door 661 
Dover E272 
Dragge, Drawke 282 
Drake I 218 


Drop-seed 

184-9, 197-200 
Drunk 282 
Duck 256 
Dudder 31 
Earning 3 226 
Eaton’s 244 
Eel I gI 
Green 2 116 
Egyptian I 229 
-millet 121 
English Blue 1 259 
Ever I 281 
Evergreen 220, 272 
Everlasting I 124 
Fairy 250 
Faitour’s 2 Sag 
False I 169 
False Buffalo 232 
False Oat 217 
False Red-top I 256 
Feather 1 177, 214, 
230, 286 
Macoun’s I 176 
Richardson’s 176 
Feather-bunch 176 
Sedge 119 
Felon 2 638 
Fescue I 269-73 
Great 273 
Fiddle 2 590 
Hard L271 
Prickle I 262 

Tall or Meadow 
272 
Fine John I 205 
Fine-top I 203 
Finger I 122-3 
Hairy 123 
Late-flowering 122 
Slender 122 
Smooth 123 
Finger-comb 229 
Fiorin 203 
Fire 2 264 
Flote 1 192; 2 265-6 
Fly-away I 207 
Fool-hay 207 


Grass, Fowl 


Vor. III. 


I 264 
Fowl-meadow 


I 207, 256, 264 


Fox 22 
Fox-tail I oe 
Alpine 193 
Branching 225 
Bristly 164 
Brown 164 
Floating 192 
Giant 166 
Green 165 
Marsh 192 
Meadow 193 
Perennial 165 
Slender 192 
Short-awned 192 
Water I 192 
Yellow 165 
French 2 365 
Frisky Meadow 1 272 
Frog 2 21, 469 
Furze Top I 205 
Gallow I 634 
Gaia aks 
illiflower 97, 41 
Good Friday =F 7 


Goose I 229, 267, 428, 
661; 2258; 3 84, 


259 
Grama 1227 
Black 227 
Blue 227 
Common 227 
Side Oats 228 
Tall 228 
Grape-vine 138 
Great Goose 3 84 
Green I 256 
Hair I 214-6 
Crested 245 
Early 215 
Long-awned 1 189 
Mountain 216 
Rough 207 
Silvery 214 
Tufted 215 
Water I 245 
Wavy 216 
Wood 216 
Hair-Dropseed 199 
Hard 251, 282 
Hassock I 214 
Haver 275,278 
-corn 218 
Heather I-233 
Hedgehog 167 
Marsh 430 
Hendon Bent 252 
Herd’s IQI, 203 
Hever 218, 220 
Hirse I 140 
Holy 172 
Alpina 173 
Arctic 172 
Hooded 278 
Hungarian 166 
Hunger 192 
Indian Ll. 120,237. 
Indian-Beard 116 
Indian-Doob 222 
Indian Rice I 168 
-millet 174-5 
Tron I 391 
Ivray 282 
Jockey 250 
Johnson 1 121, 280 
Joint 133 
Bearded 150 
Pitted 112 
Wrinkled 1i2 + 
June 220, 256 


Vor. IIT. 
Grass, June 
Prairie I 245 
Kentucky-blue 256 
Knob 3 153, 245 
Knot Ti 533 
2 30, 83, 660 
-root I 185 
Koeler’s Gy eye 
Lace 1230 
Lady 170 
Ladies’-hair 250 
* -laces 170 
Lavender 237 
Leghorn Straw 252 
Lemon 225 
Leptochloa 230 |. 
Lob or Lop 278 
London Lace 170 
Louisiana 125 
Love I 239-43 
Clustered 242 
Frank’s 239 
Hair-like 242 
Hairy creeping 243 
Low 240 
Meadow 241 
Purple 241 
Pursh’s I 240 


Short-stalked 242 
Small tufted 240 
Smooth creeping 


243 

Stout I 239 
Strong-scented 241 
Tiny 239 
Lyme 290 
Maiden-cane' 121 
-hair = 2250 
Manna 263-6 
Poland 266 
Manna-croup 266 
Maram 212 
Marl 2 355-6 
Marram-Sea 290 
Marsh 223-4 


Smooth Salt, 
Tall I 223-4 
Marsh-hedgehog 430 
8 


Mat 281 
-reed 281 
-weed 212 
ay 253 
Meadow 
253, 255-7, 264 
-fescue 272 
-spear 264 
Melic 237, 246-7 
Merlin’s 50 
Mesquite I 227-8 
Bristly 227 
Early 231 
Hairy 227 
Honey. 21333 
Prairie 333 
Side-oats 228 
Velvet 214 
Vine 138 
Milk 3 286 
Millet, see Mille 286 
Molinia 19237 
Purple 237 
Money 3 223 
Monkey’s I 203 
Moor 


2 203, 233, 237, 487 
Mountain Rice 174-5 


Mouse I 214 
Mouse-tail 1 192, 269 
Munro’s 232 
Myrtle 446 
Nard 281 


Natural 1255; 2351 


Grass, Needle 1 177,180 


Needle and Thread 
177 
Neale 282 
Nimble Will 1 184 
Nonesuch, White 281 
Nut I 304 
Oat,.see Oat-grass 
I 120, 218-22 


Black WF) 
Downy 217 
False 217 
Marsh 217 
Golden I 217 
Poor 218 
Purple 219 
Tall 217 
Wild 120, 176, 249 
Yellow 217 
Oats 218 
Sea-side 249 
Old White-top 214 
Old Witch 139 
Onion I 220 
Orange 2 536 
Orchard I 251 
Painted 170 
Panic, see Panic 
I 137-63 
Pearl 220, 250 
Penny 2) 225) 
Pepper I 543 
Phippsia 193 
Pigeon 
I 121,165; 3 95 
Pin 2 430 
Pink 1 241, 391, 415 
Pitchfork I 129 
Plume I 87, 113-4 
Japanese © 004 
Pond I 283 
Pony 210 
Poor oat 218 
Porcupine 177 
Poverty 1 178, 318 
2 355, 470, 479, 
541-2 
Long-awned 1 183 
Southern 194 
Prairie I 195-6, 
199, 231, 240 
-Chloris I 225 
-June I 245 
-Rush 188 
Prickle 121 
Prim 171 
Purple 


I 209; 2 352,358 
Quake, Quaking 
I 250-1, 289 


Tall I 263 
Quick Quack 283 
Quickens 283 
Quitch 283 

Black 203 
Rancheria I 290 
Randall 272 
Range 1 138 
Rat-tail IQI 
Rattlesnake 263 
Ray, Red 1115, 281 
Redfield’s 236 
Red-stem 115 
Red-top 203 

Bunch I 260 

False 256, 260 

Northern 204 

Tall 233 

Wild 141 
Reed I 170, 208-13 


Common, Bog 232 
Dutch, Pole 232 


Grass, 


Reed 
Indian I 202 
Mat I 281 | 
Salt 223 
Southern 170 
Sweet, Wood 202 
Reed-bent 212 
Reed-canary 170 
-meadow 265 
Rescue 280 
Rib 3 246 
Ribbon I 170 
Rice 174-5 
Rice-cut 169 
Ripple 3 246 
Rot tI 214; 3 226 
Rush I 188, 194-9 
Rush-cat’s-tail 1 190 
Russia 266 
Rye 281 
Awned I 282 
Giant 290 
Italian 282 
Perennial 281 
Western 290 
Wild 288-94 
Ryle, Sturdy 1 282 
St. Mary’s 120 
Salem 214 
Salt 199, 223, 230 
Clustered 236 
Fine-top I 199 
Rush 223 
Salt-marsh 223 
Salt-meadow 223-4 
Sand 1 235, 250, 676 
Big I 212 
Sand-bur 167 
-dropseed 199 
Sand-reed 212 
Sand-spur 167 
Satin 184-6 
Saw I 348 
Scorpion 3 85-6 
Scratch I 673 
Scutch 222 
Cane : 295 
Scurvy 2 177; 3 373 
Sea 1 88; 
2. 21h pe 718 
English 2 21 
-Marram I 290 
-meadow 267 
Sea-oats 249 
-reed, -sand 212 
-spur 267 
Sedge- I 119, 446 
Seneca 172 
Serpent 671 
Sesame 111 
Shakers 250 
Shave 41 
Shear 283 
Sheep’s-fescue I 271 
Shelly I 283 
Shining 2 120 
Shore 3 250 
Shore-husk I 190 
Side-oats 228 
Sickle I 425,673 
Silk I 177, 207 
Su37s 
Silky I 174 
Silver 3.373 
Simpson’s 137 
Six-weeks 253 
Slender I 230 
Slough 282, 28 
Small Cane 2 
I 


Smut 
Snake 2 44; 3 


om) 
wm™ Uw 


co 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 615 


Grass, Soft I 214 
Meadow, Woolly 
I 214 


Soldier's Feather 191 
Sour, see Sour 


I 653- 

Southern Reed “ie 
Sow 2 167 
Snake 2 44; 3 85 
Sparrow I 514 
- Spart 22 
Spear 253-61 
Alpine 254 
Arctic 258 
Buckley’s 260 
Bunch 260 
Flexuous 257 


Large-flowered 259 


Meadow I 264 
Mountain 254 
Prairie 260 
Sea 267 
Short-leaved 258 
Smooth I 260 
Southern 240 
ylvan 257 
Tufted 261 
Weak 255 
Wolf’s 257 
Spike I 248-9 
Marsh 250 
Spiked 92 
Spires 170, 212, 232 
Spring 171 
Spurt 333 
Squirrel-tail 287 
Squitch 259, 283 
St. Mary’s 121 
Stagger L532 


Star I 251, 463-4, 
511, 534; 3 267 


Star-eyed I 543 
Steep 3 226 
Stroil I 283 
Sturdy 283 
Swamp 197 
Sweet 91, 19 
266, 268, as 
Sweet-vernal 171 
Swine’s I 661 
Switch I 141 
Sword 170 
Syrian I2I 
Tare 282 
Tassel I 88 
Terrell 291 
Texas Blue 259 
Thatch I4I 
Thin I 206 
Tall 206 
Three-awned 1 178 
Three-toothed 234 
Three-fork 223 
Tickle 139 
Timothy 191 
California 170 
Mountain 19I 
Rush-like 190 
White 214 
Wild 186 
Tine 2 409 
Toad I 4690 
Tongue 2 43, 165-6 
Toothache I 225 
Triple-awned 179-83 
Forked 179 
Long-awned 181 
Sea-beach 183 
Slender 81 
Western 183 
Woolly 182 


Tumble-weed 1 139 


616 


Grass, Turin f£ 686 
Turkey 3 259 
Turkey-foot 118 
Turtle I gI 
Tussocks 203 
Twin 206 
Twin-spike 224 
Twitch 283 

Black 271 
Butter 220 
Onion 220 
Water 203 
Two-penny 2 712 
Umbrella I 337-8 
Union 248 
Vanilla r 172 
Nodding 172 
Velvet 214 
-mesquite 214 
Vernal-grass 
Long-awned 171 
Sweet 171 
Vine-mesquite 138 
Wag-wanton 250 
Wart . 2 473 
Water I 245; 2 162 
Large I 132 
Water Oats 168 
Twitch 203 
Way 661 
Way-bent 287 
Well 2 162 
Wheat I 284-5 
Bearded 285 
Coast I 284 
Dog 285 
False 283 
Western 284 
Whip 349 
White I 168 
Whites 214 
White Top 203, 214 
White-rush 223 
Whitlow 2 147-52 
Whorl, Water I 245 
Wickens 283 
Widgeon I gI 
Wild-canary 1 170 
-ginger 225 
-Oat I 120, 218-22 
-Rye 288-94 
-Timothy 186 
Willow I 666 
Wind 213 
Windlestraw 213-8 
Windmill een 
Winter 409 
Wire a Saute 
138, 184, 198, 
Be2teeoO;, | 220, 
259, 470, 661, 
f 262 
Wire-bent 1 280-1 
Witch 139, 280 
Barbed 139 
Gattinger’s 139 
Old 139 
Spreading 138 
Wood, Wiry 140 
Wobsqua I 141 
Wood 120, 185 
Meadow- 256 
Purple 262 
Wool 337 
Wrack I 81,91 
Yard I 229 
Yellow 487 
Yellow-eyed 451-3 
Yellow-top 210 
Yorkshire-fog 214 


Grass-flower 1543; 237 
Grass-of-the-Andes 220 


Grass-of-Parnassus 


2 212-4 
Grass-pink 1563; 274 
Grass-poly 2 580 
-weed I gI 
-wrack gi 
Gravel-plant 2 692 
Gravel-root 3) 357 
-weed 283 
Graveyard-weed 2 474 
Gray-beard tree 728 
-feather 367 
Graymile 3 88 
Greasewood 2/23 
Greek Valerian 3 63 
American 63 
Creeping 63 
Green-berry 2 693 
Green-brier I 528 
Fiddle-shaped 529 
Green-dragon 443 
-ginger 3 527 
-osier 2 663 
-sauce I 654 
-sorrel 654 
-weed, or-wood 2 350 
Grig 694 
Grim-the-Collier 3 334 
Grinsel 539 
Grip, Grip-grass I 259 
Gromwell, Amer. 3 88 
Corn 87 
False 9o-I 
Woolly 88 
Ground-berry 2 693 
-burnut 442 
-cedar 1 48; 2 542 
-centaury 2 450 
Ground-Cherry 3 156 
Barbadoes 157 
Clammy 161 
Cut-leaved 158 
Hillside 161 
Lance-leaved 158 
Large-bladder 159 
Large White 163 
Long-leaved 159 
Low 160 
Low Hairy 156 
Mexican 158 
Missouri 158 
Peruvian 161 
Prairie 160 
Purple-flowered 162 
Round-leaved 161 
Smooth 159 
Stellate 3 162 
Tall Hairy mor7 
Virginia 160 
White-flowered 163 
Ground-Fir i ifren 
-hele Brecon 
-hemlock I 67 
-holly 267 
-ivy 3 114 
-laurel 2 692 
-lemon 2 130 
-lily I 526 
-moss I 306 
Ground-nut 2 542 
Dwarf 618 
Ground-pea 2 418 
Ground-pine 1 45, 47-8 
2 536,586; 3 102 
Festoon 47 
Ground-plum 2) B77, 
-raspberry 2 85 
Ground-squirrel-pea 
2 129 
Ground-vine 3 276 
Groundsel 439-46 
Balsam 545 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Groundsel 


Common 3 539 
Cress-leaved 540 


Entire-leaved 3 546 
Fetid 540 
Marsh 540 
Silvery 543 
Viscous 540 
Wood 539 
Groundsel-bush 3 445 
-tree 445 
Grouseberry 2 693 
Grundy-Swallow 3 544 
Guelder-rose, Wild 270 
Maple-leaved 270 
Gum, Black 2 665-6 
Cotton 2 666 
Red 235 
Sour 2 665-6 
Spruce-tree I 61 
Star-leaved 2 235 
Sweet 235 
Tupelo 666 
Water 2 666 
White 7) Bas 
Gum-Elastic 720 


Gum-plant 3 67, 92, 371 
2 6 


Gum-tree 65 
Spruce a 61 
Yellow 2 665 

Gun-bright I AI 

Gutierrezia 3 370 

Gutter-tree 2 662 

Gutweed 3 316 

Gymnopogon I 225-6 

Gypsophyll 2 71-2 

Hackberry I 629-30 

Hackmatack 1 60, 66 

Hacmack 60 

Hagweed 2 350 

Hail-, Hair-,weed 3 49 

Hair-fern f 62 


Hair Grass 
I 189, 207, 214-6 
Hairhoof, Sweet 3 267 


Hairhound 123 
Hairif 259 
Hammer-sedge 1 428 
Hammerwort I 638 
Handsome Harry 2 583 
Harbinger of Spring 653 
Hardhack 1 607; 2 262 
Purple 2 245 
Spice 2 245 
Hardheads 3.557 
Hardock 547 
Hardweed 557 
Harebell 3 295 
Arctic 2904 
Hare’s-beard 3.173 
-ear 2 175, 639 
Hare’s-foot Fern 1 8 
Hare’s-tail 324 
Hartford-fern IQ 
Hart’s-eye 2 634 
Hartshorn-bush 197, 
-plant 2 102 
Hart’s-thorn 2 502 
-tongue I 25 
Harvest-bells S) ate 
-lice 3 496 
-lily 3 46 
Hathorne 2 319 
Haver-corn I 218 
Haw (see Thorn) 
2 297-321 
Barberry-leaved 
2 207 
Biltmore 307 
Black- 2720; 3273-4 
Dotted 2 300 


Vor. III. 

Haw, Downy 2 319 
Parsley 320 
Pear 302 
Poplar-leaved 308 
Possum 2488; 3273 
Red = 2-297, 302, 7, 
17, 21 

Scarlet 317 
Shawnee 3278 


Small-fruited 2 320 


Summer 310 
Tree 2 307,19 
Yellow 310 
Hawkberry 2 327 


Hawkbit 3 310, 328, 332 
Hawk’s-beard 3 325-8 


Gray 28 
Hawkweed i 
3 311, 328-32 
Canada 329 
Early 332 
Field 333 
Golden Mouse-ear 
334 
Green’s 332 
Gronovius’ 331 
Hairy 331 
Long-bearded 330 
Maryland Bae 
Mouse-ear 333 
Narrow-leaved 330 
Orange 334 
Panicled 330 
Rough 331 
Tawny 334 
Vein-leaf 332 
Wall 329 


Hawthorn 2 297, 317-9 


English 319 
Haw-tree 319 
Hay-fever weed 3 341 
Hayhofe 3 114 
Haymaids 114 
Hay-plant 267 
Hazel, Black I 607 

Snapping 2; 285 

Witch 235 
Hazel-nut I 607 

Beaked 607 
He-huckleberry 2 485 
Headache 2 137-8 

-plant 2 102 
Head-flower Riess 

-Betony 221 
Headsman 3 246 
Heal-all I 514,555 

‘ 3 T15s179 

High 2 221 
Heal-bite 2 154 

-dog 154 
Healing-blade 

2 201, 245 

-herb 3 92, 246 
Heart-clover 2 352 

-leaf I 643-4; 2 352 
Heart-of-the-earth 

325 
Heart-pea 2 501 

-seed 501 

-trefoil 2 352 
Hearts 2 431 
Heartsease 

I 666,8; 2 563 
Heart’s-pansy 2 563 
Heartweed I 668 
Heartwort 2 353 
HEATH FAMILY 675 


Heath 2 479, 542; 3 295 


American 541 
Blackberried 2 479 
Corn I 672 


Cross-leaved 2 694 


VoL. II]. ENGLISH INDEX, INCLUDING POPULAR PLANT 
Heath, Mountain 2 685 | Henbane, Black 3 169] Hippo, Indian 2 248 
Scotch 694 ellow 162 | Hippophae 2 576 
Heath-bells 3 295 | Henbit 2 121-3 | Hirse I 140 
Heathberry 2 479 Small 203 | Hive-vine 
Heath-cypress I 46| Henbit Dead-nettle 2 392,397; 3 255 
-grass 233 I 121-3 | Hoarhound, Black 12 
Heather 2 694 | Hen-plant 3 245-6 Base 126 
Beach, False ae Hens 2 560 Bastard 123 
Monox 479 | Hep- or Hip-tree 2 284 Common IIo 
Heavenward Tree 446 | Herb-bane 3 235 Fetid 123 
Hedge-bells 3 47 -barbara 2 177 Marsh 148 
-burs, Wild 3 259 -bennet 2 270-1 Water 147-8 
-Dead-nettle I21 -christopher 17s White IIO 
-garlic 2 170 2 90, 249; 3 128 Wild 360-2 
Hedgehog Cereus -gerard 2 654 | Hoarwort 3 447 
568, 629 -grace 3 95,192 | Hobble-bush 269 
-thistle 570 -holy 3 95 | Hoffmanseggia 2 338 
Hedge-hyssep 3 192-6 -impious 3 447 | Hog-apple I 130 
Hedge-lily 3 46 -ivy 2 167 -bean 3 169 
Hedge-maids 114 -john 53 -bed I 48 
Hedge-mustard 2 174 -margaret 3 402,518 -bite 3 314 
Fine-leaved 170 -mercury 2 460 -cranberry 2 479 
Hedge-nettle 3 124-7 | Herb-of-the-Cross 395 | Hog Peanut 2 419 
-parsley 2 526! Herb-robert 2 426 -physic 3 300 
Hedge-peak 284 -sherard 3 266 | Hog’s-bean 3 169 
Hedge-plant I 632 -sophia 2 170 -fennel 2 638; 3 516 
-strawberry 2 260 -twopence 2 212 -potatoe I 492 
-taper 3 173 -trinity 2 101, 563 | Hogweed 
-thorn 2 319 -wicopy 101; 2 590 2 626; 3 341, 356 
-weed 2 174 -william 2 657 | Hogwort 2 454 
Hedysarum 2 392) Hercules’ Club Hollard I 613 
Heliotrope 3 ie 2 444, 617 | Hollow-root 3 283 
Bindweed Heron’s-bill 2 430] Hotty Famity 2 486 
Garden, Hardy 2 8% Heuchera 2 225-8| Holly, American 486 
Indian 75 | Hexastylis I 643-4 Dahoon 2 487 
Seaside 74 | Hickory I 580-4 Deciduous 488 
Slender 74 Big Shag-bark 1 582 Emetic 487 
Summer 2 a Bitter 580| Ground 2 672 
Wild Bitter-nut I 580 Large-leaved 2 488 
Hellebore 1 489; 2 & Bitter Pig-nut 580 Meadow 488 
American White Black 582 Mountain 488 
I 4904 Brown,or Broom 583 Swamp 2 488 
Bastard 563 | Bullnut 582 | White 486 
Big 494 Fragrant I 582 Wild 2 490 
False I 494-5 Hardbark 582 | Holly-bay 2-527 
Green 2 87 King-nut I 582 | Hollyhock 514 
Swamp I 494 Mocker-nut 582 Sea 524 
Winter 2 88 Northern 583 | Holy-hay 2 351 
Wood's False 1 495 Pale 582 -her 3.95 
Helleborine I 563 Pecan I 580 -rose, Marsh 2 689 
Hellroot 3 235 Pig 580 | Homewort 2e20% 
Hellweed 2 115 Pig-nut I 583 | Honesty 2 191 
Helmet-flower 3 105 Red 582, 583 | Honewort 2 630 
-pod 2 129 Redheart 581 | Honey 339, 352 
Hemlock | 62 Scurfy 584 Fenecballs 3 255 
Bulb-bearing 2 658 Shag-bark 1 581 | Honey-blobs 2 241 
Carolina 62 Southern 581 | Honey-bloom 3 21 
Creeping 67 Shell-bark I 581 -locust 2 339 
Ground 67 Big 582 -lotus 352 
Lesser, or Small 645 Thick, or Western -mesquite 2 333 
oison 2 653 I 582 -plant 3-137 
Southern I 62 Small-fruited 1 583 -shucks 2 339 
_. Spotted 2 658 Soft-shell 580 -talks 2 358 
Water 658 Swamp 580-1 | HonEYSUCKLE FAMILY 
Hemlock-Chervil 2 626 Sweet 581 3 267 
Hemlock-spruce 62 Upland 581 | Honeysuckle 2 92, 359 
Hemp FAMILY I 633 Water ne Mates 3 278-83 
Hemp I 634; 2 5,6 White 581-3 Bush 3 282-3 
Albany 636 White-heart 1 582 Chinese 280 
American 2 521 | Hickory Poplar 2 83 Cinnamon 2 679 
Bastard 3 119] Hieble 2 618 Clammy 2 679 
Indian 2 521 | High-belia 3 301 Coral 3 280 
Nettle 3 119 | High or Hig Taper Crimson 279 
Water Bas 6 2.273 Douglas’ 3 279 
Wild 3 119, 341) High-watershrub 3 339 Early 2 678 
Hemp Dead-nettle 119 | Hillberry 2 693 Fly 3 281-2 
Hemp-nettle, Red 119 | Hills-of-snow OR Glaucous 3 279 
Common 119 | Hillwort 3 I4! Ground 2 359 
Hempweed, Climbing Hindheal 2 14; 3 522 Hairy 3 278 
3 363 | Hip-brier,-rose 2 284-6 Italian 270 
Hen-and-chickens -tree 2 284-5 Japanese 280 
2 211; 3 402! Hippo, Wild 2 469 Garden Fly 282 


NAMES. 617 
Honeysuckle 
Perfoliate 3 278 
Purple 2 678 
Scarlet Trumpet 
3 280 
Small Yellow 3 279 
Smooth 2 679 
Sullivant’s 3 279 
Swamp 2 678-9 
Tartarian Bush 3 282 
Trumpet 3 280 
White 2 679 
Wild 2 609-10, 678 
Yellow 2 679 
279, 280 
Honeysuckle ‘Apple 
2 678 
-clover 3 565, 568 
Honey-sweet 249 
Hoodwort 106, 531 
Hoofs 3 245 
Hook-heal 3 115 
-weed I15 
Hoop-ash 1 629; 2 72 
Hoopkoop-plant 2 408 
Hoopwood 2 489 
Hop or Hops I 633 
Bog 2 18 
Japanese I 633 
Wild 2 122; 3 128 
Hop-clover 2 354 
-hornbeam I 606 
Hop-tree 2 445 
-trefoil 2 354 
-vine, Devil’s 1 528 


Hornbeam, Am. 


I 606, 665 
Swamp 2 665 
Horn-bine 665-6 
-pine, -pipe, 665 
Horned Clover 351 
Horned Rush i 342 
Horn-weed 7 
Hornwort FAMILY 
2 75 
Hornwort 75 
Horse-balm 3 153 
Horse-blobs 2 85 
-bramble 2 284 
Horse-brier I 528 
-cane 3 341 
Horse-chestnut 2 498 
American 498 
Horse-elder 3 457 
-foot 532 
Horse-fleaweed 2 345 
Horsefly-weed 2 345 
Horse-gentian 3 275 
-ginseng 275 
-gold 2 I1I-12, 
103, 105 
Horse-gowan 3 521 
Horse-heal 457 
-hoof 531 
-knobs, -knop 557 
Horse-laurel 2 681 
-lily 278 
Horse-mint 3 131-3, 
136, 150 
European 150 
Ohio 135 
Stone 146 
Sweet 146 
Water 151 
Horse-nettle 3 165 
Horse-pipes I 39-40 
-radish 2 163 
-savin 1 66 
| Horse-sorrel I 653 
Horse-sugar 2 721 
HorSE-TAILFAMILY 138 
| Horse-tail I 39-41 


618 


Horse-tail 
Cornfield I 39 
Field 39 
Marsh 40 
Meadow 39 
Rough 4I 
Shore I 40 
Swamp 40 
Thicket 39 
Water 41 
Wood 40 

Horse-thyme I 138 


Horse-weed 3153, 341 


Low, Purple 3 442 
Hosh-kawn Tsr2 
Hound-bene 
Hound’s-berry 110 


Hound’s-tongue 
3 76, 369 
House-leek 2 208-9, 211 


Houstonia 3 251-4 
Hove 3 114 
HvucCKLEBERRY FAMILY 
2 694 
Huckleberry 2 695- 7 
Black 696 
Blue 702 
Box 696 
Bush 696 
Dwarf 696 
He 2 485 
High-bush 696 
Southern Black 700 
Squaw 697 
Strawberry 701 
Tree 2 698 
Huckleberry Lily 1 502 
Hudsonia 2 541 
Hunger-flower 150 
Hunger-weed 200s 
Hungry-vine I 528 
Huntsman’s Cup 2 202 
Hurr-bur 3 547 
Hurt-sickle 557 
Huskroot 1g gi 
Hutchinsia 2 158 
Hutton-weed 3 289 
Hyacinth I 510-1 
-bean 2rAL7, 
Grape I 510-1 
Starch- 511 
Wild 1 509; 2 142 
HyDRANGEA FAMILY 
2 230 
Hydrangea 2 230 
Ashy 231 
Snowy 2a 
Hoe 231 
Hyeble 2 618 
eeieencallia E533 
Hymenopappus 3505-6 
Hyssop 3 140, 192 
Anise as 
Garden 140 
Giant 3 I1I-2 
Hedge 192-6 
Prairie 3 142 
Water 193 
Wild 395 
Hyssop Hedge-nettle 
Buz 
Loosestrife 2 580 
Iceland-moss 2 210 
-poppy 2 138 
Ice-leaf By ais) 
-plant, Amer. 2 674 
Tllinois-nut I 580 
Impudent-lawyer 3 177 
Indian-apple 2 130 
-arrow 491 


-arrow-wood 


Indian-balm I 525 
-bark 2 82 
-bean 3 238 
-beard-grass I 116 
-black-drink 2 489 


-boys and girls 2 142 


-bread-root 363-4 
-cedar I 607 
-cherry 2 292, 503 
-chickweed a 28 
-chief 25707, 
-chocolate 2 292 


-cucumber-root I 523 


-cup 2 201; 3 460 
-currant 277 
-doob 1 BOE 
-dye 2 85 
-elm I 627 
-fig 2 570 
-fog 2 209 
-ginger I 642 
-grass I 120 


-gravel-root 3 357 
Indian-hemp 2 521; 33 


Black B22 
White 3 26 
Indian-Hippo 2 248 
-ice-root 2 85 
-lettuce. 668 
-mallow 2 520 
-millet 5 ae ay 7 
-moccasin I 550 
-mozamize 2 287 
-paint 215; 388 
Red 2 140 
Yellow 85 
Indian-physic 2 181, 248 
-pine 2 159 
-pink 2 452, 640; 
3.42, 214-5 
INDIAN-PIPE FAMILY 
2 673 
Indian-pipe 674 
-pitcher 2 201 
-plantain 3 536-8 
-poke I 494-5 
-posy 3 453, 4, 82 
-potato 2 418 
-puccoon 3 89 
-red-root Pa5gk 
-root 2627, 
-sage 3 361 
-shamrock I 525 
-shoe I 550 
-soap-plant 2 500 
-strawberry 2s 
-thistle 3 289 
-tobacco 3 170,451 
-turmeric 2 85 
Indian-turnip I 442 
Brown's 443 
Three-leaved 442 


Indian-wickup 2 590 
Indians, Red 3 214 
Indian’s Dream 5 RGYs) 
Indian’s Plume 3 131 
Indigo, Wild 2 344-6 


False, Bastard, 2 365 
Blue False 344 
Dwarf False 366 
Fragrant False 2 366 


Prairie 2 346 
White False 2 345 
Indigo-broom 2 345 
-plant 372 
-weed 2 345 
Infant’s-breath 263 
Inkberry 2 26, 487 
Shining 2 488 
Ink-root 2 718 
Innocence 3 .188, 251 
Insane-root 169 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Todanthus 2 178 
Ipecac,Am. 2 248,470 
False 248 
Milk 469 
Spurge, White 2 470 
Wild 27 eso 
Wood 3 275 
Iresine 28 
Ir1s FAMILY T536 
Iris I 540-1 
Crested 540 
Dwarf 540-1 
Spring 541 
Irish-daisy Bus05 
-mahogany I 613 
Iron-head 3 557 
-weed 95, 351-3, 487 
Devil’s 3 220,557 

Tron-wood 
I 606; 2 485 


Southern 2 435,719 


Ironwort 3 119, 110-1 
Isabella-wood 2033 
Isopappus 3 380 
Itch-weed 494 
Itea 2233 


Ivory Plum 2 693, 704 


Ivray 282 
Ivy, American ; Sia 
Big-leaved 684 
Climbing 2 484 
Coliseum 3-175 
Five-fingered 2 511 
Five-leaf ire 
Ground 3 114 
Kenilworth 175 
Poison 2 484 
Three-leaved 484 
Spoonwood 2 683 
Ivy-berry 693 
-bush 2 684 
-chickweed 3 203 
-flower 2 101 
-weed 20175 
Jack-by-the-hedge2 170 
Jack-in-the-pulpit1 442 
-in-the-bush 2e070 
-in-trousers 2 92 
Pecks 442 
Jack-straws 3 246 
Jackson-vine 3 168 
Jacob’s-ladder 1 527; 


2 493; 3 63,177 


American e277 
Jacob’s-staff 3.173 
-sword 540 
Jalap, Cancer 2 26 
Wild 3 43 
Jamestown-weed 3 169 
lily 169 


Japanese-creeper 2 511 
Jasmine, Am. 3 43, 168 


Bastard 3 168 
Carolina 2 730 
Red 3 42 
Jaundice-berry or -tree 
2 127 

Javril, Sweet 628 
Jenny 2.712 
Jenny Wren 2 426 
Jersey Tea 2 504 
Jerusalem-oak 2 14 
-artichoke 3 486 
-sage 118 
-star 313 
Jessamine 3 168 
Bastard 168 
Blue 25123 
Red 3 42 
Water 194 
Yellow 730 


Vot. III. 
Jesuit’s-bark 3 339 
-Water-nut 2 612 
JEWEL-WEED FAMILY 
2 440 
Jewel-weed 2 440-1 
Speckled 440 
Jewels, Speckled 440 
Jew’s-harp-plant 1 526 
Jew’s Ear 3 168 
Jimson-weed 3 169 
Joan Silver-pin 2 137 
Job’s-tears TAS iGs 
i 3 90 
Joe Pye Weed 3 357 
Spotted 356 
Johnny-jump 2 717 
Johnny-jumper 2 563 
Johnny-jump-up 
2 547, 63 
-smokers 273 
Johnswort I 533 
False 2 536 
Joint-grass : er 
-vetch 


393 
-weed I 676-7; - 612 
Jointed-charlock 2 195 


Joseph’s-flower 3 313 
Jove’s-flower 273 
-fruit 2 136,.720 
Joy, American 2 511 
Joy-leaf 35335 
Juba’s-bush 8 
Judas-tree, Am. 334-5 
Red 235 
June-berry 291-2 
-flower 560 
-plum 292 
Jungle-rice I 134 
Juniper 1, 5-6 
Creeping 67 
Low, Red 66 
Juniper-bush 66 
-tree 629 
Juno’s-tears 395 
Jupiter’s-beard 2 211 
Jupiter’s-staft 2.173 
Justice-weed 3 357-8 
Jute, American 2 521 


Kale,Corn orField 192 


Kalmia 2 684 
Kedlock 2 191-2 
Kelp 2 28 
Kelpwort 2 25 
Kemps, Kempseed 
3 ae 
Sea 247 
Kentucky Mahogany 
2 41 
-moss 2 41 
Kerlock 2 191-2 
Ketmia, Bladder 2 525 
Kettle-dock 3 542 
Kicking-colt 2 440 
-horse 440 


Kidney-bean, Wild 
2 


428 

Kidney-bean Tree 374 
Kidney-root 3 357 
-wort 2 225 
Kill-cow 318 
-kid 2 683 
-wart 2 141 
-wood 581 
King-cups 2 8c, 172 
-devil 3 332 
-nut I 581-2 


-of-the-meadow 3357 


King’s-clover 2 352-3 
-crown 353 
-cure 672 
-cure-all 2 595 


VoL. III. ENGLISH 
King’s-fern Ta 
Kinnikinnic 2 661, 93 
Kino-root, Am. 2 426 
Kiss-me 2 563 
Kiss-me-Dick 2 474 
Kisses 2 563 
Knap 2 355 
-bottle 2 64 
Knapweed - 3 557-9 
Knawel 2 30 
Knight-cross 2 69 
Knit-back 3 92 
Knob-, or -knot-grass 
eS 3) 5 Sel 5S 
Bird’s I 661 
Coast 660, 663 
German 2 30 
Japanese I 676 
Knob-root geric3 
Knotweed 1 153, 663-5 
Biting 670 
Bushy 663 
Shore 662 
Spotted 668 
Virginia 665 
Knout-berry 2 276 
Kochia 20 
Koeleria 245 
Konigia, Seaside 2 153 
Korycarpus I 247 
Kosteletzkya 2 523 


KRAMERIA FAMILY 340 
Krameria 340 
Kraut-weed 2 192, 195 


Kudzu-vine 420 
Kyllinga I 296 
_ Labrador 2 617 
Labrador Tea 677 
Lace-button 3 440 
-flower 2 625 
Lad’s Love 3 526 
Ladies’-eardrop 2 440 
-cushion 3 557,718 
-mantle 2 264 
-pocket 2 440 
Ladies’-slipper 1 548- 


50; 2 440, 452 


Downy I 550 
Pink or Purple 550 
Ram’s-head 548 
Stemless 550 
Yellow 550 
Lady’s-smock 2 184 
-sorrel 3 gare 
-tobacco 
Ladies’ Tresses 1 neat ae 
Drooping 565 
Lady 
Eleven-o’clock 2 574 
Laurel I 506 
Ten-o’clock 506 
.Lady-by-the-gate 2 72 
Lady-fern I 30 


-finger, -glove 3 204 


-laurel 574 
-thimbles 3 winks 
Lady-never-fade 3 453 
-belt 2 249 
Lady’s-clover 431 
-comb 627 
-cushion 
2719 333) 557 
-finger 2 359-60, 415 
-delight 2 563 
-hair I 250 
-laces 170 
-lint 2 44 
-mantle 2 264 
-milk 555 
-needlework 2 626 
-nightcap 3 46 


INDEX, INCLUDING POPULAR PLANT NAMES. 


Lady’s-purse 2 158 
-shoes 2 93 
-shoes-and-stockings 

2 
-thimble 3 aoe 
-thumb I 668 

Lake-iris 540 

Lakeweed 670 

Lamb-kill 2 680, 683 
-lily I 486 
-mint 3 149 
-sucklings 2358 

Lambs 2 498 

Lamb’s-cress zea 
-foot 3 245 
-lettuce 3 246, 286-7 
-quarters 

Pes 25 210, 1S 
‘-succory 3 308 
-tails Ay, 
-tongue 

T5060) 93) 152, 246 

Land-cress 2 177, 184 

Langdebeef 2) Syrae 

Lap-love 47 

Larch, American 1 60 
Black or Red 60 

Larea-bell 3 478 

Larkspur 2 94-6 
Azure or Blue 95 
Nelson’s 95 
Prairie 95 
Rocket 2 94 
Trelease’s 904 

LAUREL FAMILY 2 133 

Laurel 2 680-1 
American 684 
Bay 82 
Bee 681 
Big, Big-leaf 681 

lack 2 527 
Deer 681 
Dog 688 
Dwarf 2 683 
Florida 2k 
Great 2 681 
Ground 2 692 
Hairy 685 
Horse 2 681 
Lady 2 574 
Mountain 681, 684 
Pale 684 
Sheep 683 
Small ‘ 684 
Spurge 574 
Swamp 82, 527, 684 
White 82 
Wood 2 684 

Laurel Magnolia 2 82 

Lavender 2 718 

Lavender-thrift 718 

Lawyers 2 284 
Sea, Seaside 718 

Lead-plant 2 366 

Leaf-cup 3 458 

Leather-bush TAG AS 
-flower (see Cle- 

matis) 2 123-6 
-leaf 2 687 

Leather-wood 575 
Southern 485 

Leavenworthia 189 

Leaver-wood 575 

Ledum-oil 2 676 

Leechwort 3 246 

Leek, Wild I 497 
Sour 653 
Three-seeded 497 

Leeks 2 207 

Leitneria I 586 


Lemon, Wild 2 
Ground 


130 
130 


Lemon-balm 


3 137 
-lily I 496 
-lobelia 3.137 
-monarda 3 134 
-walnut I 579 

Lentil 2 412 
Water I 448 
Leopard-flower 1 542 


Leopard’s-bane 3 533-4 


Leptochloa I 229 
Lesquerella 2 154 
Lettuce 3 318-20 
Arrow-leaved 3 320 
lue 320-1 
Canker 2 668 
False 3 320-1 
Florida Biaen 
Hare's 317 
Hairy Wood 319 
Indian 2 668 
Lamb’s 237-8 
Liverwort 2 668 
Prickly 3 318 
Red Wood 319 
Spanish 2 39 
Steele’s Wild 319 
Tall 3 320 
Tall Blue 321 
Tall White 335 
Western 319 
White 3 318, 35-7 
Wild 2 668 
Willow 3 318 
Wood 3 319 
Leucocrinum I 496 
Leucothoé 2 687-9 
Lever-wood I 607 
Liberty-tea Qa 


Licorice, Amer. 
2 391,419 
2 618 


Wild 
Torrey’s 3 260,1 
Licorice-root 2 3901 


Life-everlasting 3 453 
Fragrant or Sweet 


454 

Life-lasting f 4406 
Life-of-man 2 207, 287, 
é 617; 3 283,454 
Life-root 3 544 
Lilac 2 724 
Summer 2175 
Lilaeopsis 2 648 
Lity FAMILY I 495 
Lily I 502 
Asa-Gray’s I 503 
Atamasco 532 
Beaver 2 78 
Blackberry I 542 
Butterfly I 508 
Canada I 503 
Carolina 504 
Clinton’s 514 
Conval 22 
Corn 3°47 
Cow 2 78, 85 
Day I 496 
Dog, Horse 278 
Dwarf Tiger 1 542 
Easter, Fairy, 532 
Field 503 
Flag, Liver, 537 
Flame 502 
Frog 2 78 
Glade 502 
Great Yellow 2 77 
Ground I 526 
Gunebo 2 568 
Harvest 3 46 
Hedge 46 
House 2 78 
Huckleberry 1 503 


Lily, Jamestown 3 169 
Lamb I 486 
Lemon 496 
Little Water 76 

iver I 
Mariposa 238 
ay I 522 
Meadow 503 
Nodding 503-4 
Northern 514 
Philadelphia 502 
Plantain 496 
Pond 2 78-80 
Prairie 1 533; 2 568 
R ed af 502 
Rock 2 92, 102 
Sand I 496 
Southern Red 1 503 
Snake 537 
Spider 461 
Spring 506 
Swamp 532, 78 
Straw 519 
Tiger I 502,4 
Dwarf 542 
Wild 502, 4 
Toad 279 
Trinity I 52 
Trout 506 
Turk’s-cap 
1 50477 
-head I 504 
Water I 444; 319 
Western Red 1 502 
White 525 
White Water 2 79 
Wild Orange 502 
Wild Yellow’ 1 503 
Wood I 502-3, 
522; 2 670 
Yellow I 506 
Day I 496 
Pond 2 78 
Lily-bind 3 46 
LILY-OF-THE-VALLEY 
FAMILY T5r3 
Lily-of-the-valley 
517-22 
False DeSt7 
Wild 514-17, 599 

Lime Tree 2 83,512 
Black 14, 512 

Limewort-catchfly 2 66 

Limonillo 3 510 

Lin 2 512 

Linaria 3.177 

Lind, White 512 


LINDEN FAMILY 2 


Linden, American 512 
Ling 2 674 
Wire 479 
Ling-berry 607 
-gowans 3 333 


inn : 
Black, Yellow, 2 82 


Southern 2 512 
White 512 
Linseed 2 436 
Lint 437 
Fairy 2 438 
Lady’s 44 
Lint-bells 2 436 
Lin-tree 512 
Lion’s-ear 3 120 
-foot 2 64; 3 335-6 
-mouth 3 198, 204 
-heart 3 116-17, 335-6 
-snap 3 178 
-tail 49, 315 
Lip-fern I 34 
Lipocarpha 339 
Liquidamber 2 235 


620 


Liquorice 2 391 

See Licorice 419 

Little-good 2 473 
Little-boy’s Breeches 

2 UAT 

Little-pollom 452 


-washerwoman 3 251 
Littlewale 88 


Live-forever 2 207-8 
Wild 208 
Live-long 2 207 
Liver-berry I 520 
-leaf 2 101 
-lily I 537 
-moss 2 101 
Liverwort, Noble 2 t1o1 
Heart IOI 
Three-leaf IOI 
LIZARD’S-TAIL FAMILY 
1577 
Lizard’s-tail 578 
LoasA FAMILY 2 565 


LoBELIA FAMILY 3 299 


Lobelia 3 299-304 
Bladder-pod 303 
Brook 303 
Canby’s 304 
Dewny 302 
Glandular 302 
Great 301 
Kalm’s 303 
Long-leaved 301 
Nuttall’s 304 


Pale-spiked 302 
Red, Southern 300, 1 
Spiked 303 
Swamp, Water, 300 
Loco-vetch, Colorado 


2 390 

Loco-weed 2 347 
Palibine’s 391 
Stemless 390 
Woolly 379 


Locust, Black 2 339,375 
Bristly,or Moss 2 375 


Clammy 375 
Green 375 
Honey 339, 375 
Pea-flower 375 
Post, or White 375 
Red-flowering 375 
River 2 365 
Rose-lowering 375 
Swamp 2 339 
Sweet, Water, 339 
Thorn 339 
White 375 


Yellow 2 343,375 
Locust-tree 375 
Loeflingia, Texan 2 60 


LoGANIA FAMILY 729 
Loggerheads Bush 
London Lace I 170 
-pride 2 73-5 
-tuft 75 
Long-beard I 456 
-moss 456 
Long-purples 2 586 
Longshucks I 59 
LoOSESTRIFE FAMILY 
2577 
Loosestrife 580-5 
Bastard 585 
Bulb-bearing 2 712 
Creeping 712 
False 2 583, 587 
Fringed 2703 
Golden 711 
Hyssop 580 
Lance-leaved 714 
Linear-leaved 714 
Purple 581 


Loosestrife 
Southern 2 713 
Spiked 581 
Spotted 711 
Swamp 580 
Trailing 713 
Tufted 714 
Whorled 2 711 
Yellow 711 
Wing-angled 581 
Lophiola I 435 
Lophotocarpus 1 97-8 
LopsEED FAMILY 3 244 
Lopseed 3 244 
Lord-and-ladies f114 
Lotus, American 2 77 
Honey 2eaGe 
Indian 77 
Lotus-tree 2 720 
Lousewort 3 220-2 
Marsh,Swamp = 221 
Louseberry 2 492 
3 206, 221-5 
Lovage, Scotch 2 635 
Love-apple 3 168 
-grass I 239 
-entangled 2 208 
-in-a-chain 209 
-in-winter 672 
-in-idleness 563 
Loveman 3 259 
Love-me 85 
Lover’s Pride I 668 
Love roses 3 270 
Love-vine 2122; 352 
Love's test 3 451 
Low-belia 3 303 
Lucerne 2 351 
Ludwigia 586-9 
Ludwigiantha 585-6 
Lungwort 3 82-3 
Bullocks 173 
Cow’s 25173 
French, Golden, 329 
Sea 82 
Smooth 82 
Tree 83 
Lupine 2 348-9 
False 2 344 
Low 349 
Perennial 2 348 
Silvery 348 
Wild 348 
Lustwort 2 203 
Lychnis, Arctic 27,0 
Evening 2 68 
Nodding 70 
Scarlet 69 
Lygodesmia 3 322 
Lyre-tree 2 83 
Macounastrum I 647 
Mad-apple 3 169 
Mapper FAMILY 3 250 
Madder 263 
Blue Field 266 
Wild 263 
Mad-dog Weed 
I 94; 3 106 
Madderwort 3 250, 525 
Madnep 2 634 
Madweed 3 106 
Mad Woman’s Milk 
2 473 
Madwort 2 153, 157 
German 3 84 
MAGNOLIA FAMILY 2 80 


Magnolia, Fraser’s 81 


Ear-leaved 81 
Great-leaved 81 
Large, Long-leaved 

} 81 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Magnolia, Laurel 2 82 
Mountain 82 
Small, Swamp, 82 

Magotty Boy Bean 2 337 

Mahaleb 328 

Mahogany, Irish 1 613 


False 25133 
Kentucky 340 
Mountain I 609 


Mahonia, Trailing 2 128 
Maidenhair 
I 28, 29, 31,36 


Black or True’ I 31 
-berry 2 704 
-cane Y 1235927 
-tears 2 64 

Maid’s-hair 3 258 
Maize 3 516 

Thorn 2 554 
Malice 2505 
MALLow FAaMILy 2 513 
Mallow 

Blue 25505 

Bristly-fruited 523 

Common 514-5 

Country 514-5 

Curled 515 

Dwarf 515 

European 515 

False 519-20 

Glade 518 

Globe 521-2 

High 514 

Indian 2 520-1 

Low 515 

Marsh 514 

Musk 515 

Poppy 516-7 

‘Prairie 519 

Rose 524-5 

Running ue 

Swamp 524 

Venice 525 

Vervain 515 

Virginia 520 

Water 524 

Whorled 515 

Mallows, Blue 25515 
False 520 
Indian 520 

Maltese Cross 2 69 

Mandrake, Wild 

2 130, 611 
Manna-grass I 263-6 
Manna, Poland 266 


Man-of-the-earth 3 43 
Man-root 3 43-4 
Man’s Motherwort 


2 461 
MAPLE FAMILY 2 494 
Maple 494 
Ash-leaved 498 
Bark 497 
Bird’s-eye 496 
Black 496 
Black Sugar 496 
Carolina 495 
Creek 494 
Curled 496 
Cut-leaved 498 
Drummond’s 496 
Dwarf 497 
Goose-foot 497 
Hard 495-6 
Low 497 
Moose 497 
Mountain 497 
Narrowfruit 495 
Northern 494 
Norway 2 494-8 
Red I 639; 2 4905 
Red-river 2 49 


Vor. III. 

Maple, River 2 49 
Rock BS 
Rocky-Mountain 497 
Scarlet 495 
Shoe-peg 495 
Shrubby 497 
Silver 494 
Silver-leaf 494 
oft 2 494, 5,7 
Striped 497 
Sugar 496-8 
Swamp 494, 5,7 
Sweet 496 
Sycamore 494 
Water 495-7 
White 494-5 
Marble-flower 2 137, 
March 2 660 
Mardling 448 
Mare’s Tail Are 


2 612-3 ; 3 430, 442 
Margaret, Herb 3 402 


Marguerite 402 
Marigold 3 519 
Bur 495, 7 
Fetid 3 513 
Marsh 2 85-6 
Rayless 3 497 
Water 500 
Wild 521 
Mariposa Lily I 508 
Marjorum, Pot 3 140 
Wild 140 
Markery 214 
Markry 484 
Mark-weed 484 
Marl-gress 256 
Marram i252 
-sea-grass 290 
Marrube 3 110-1 
Marsh-beetle I 68 
-berry 2 704 
-clover cea) 
-elder 3 339-40 
-fleawort 540 
-five-finger 2 258 
-fleabane 3 447-8 
Fetid 447 
Spicy 448 
-gentian 12-3 
-gilliflower 2 69 
-grass I 223 
-hoarhound 3 148 
-holy-rose 2 689 
-lousewort 3 221 
-mallow isi 
-marigold 2 85-6 
-milkwort 448 
-parsley 660 
-pennywort 649 
-pestle 168 
-pink 3 6-7 
-root 2 718 
-rosemary 2 689,718 
-samphire 21 
-tea 677 
-trefoil 3 18 
-turnip I 442 
-valerian 3 285 
-weed I 40 
Marshallia 3 502-3 
Marshlocks, Purple 
2 258 
Marshwort 704 
MarSILEA FAMILY I 36 
Marsilea 37 
Martinoe 3 239 
Marvel 110 
Masterwort 
2 635-8, 654-7 
Imperial 638 
Matfelon 3 557 


Vot. III. ENGLISH 
Mather 3 516 
Matrimony-plant or 
-vine 2 191 
Matse 3 168, 516 
Matweed, Sea I 212 
Maul 2 514 
Maw-seed 137 
May 2 319 
May-apple 2 130 
-blob 85 
-blossoms I 522 
-bush 2 319 
-cherry 218, 91 
Mayflower’ 2 37,98, 
100-2, 84, 218, 678, 
692 
May-gowan 3 402 
May-grass E 253 
May-lily 522 
-pear 2 292 
-pops 565 
-rose 3 270 
-star 2 715 
-wings 452 
Mayaca FAMILY I 450 
Mayaca 450 
Mayweed 3 516 
Corn 520 
Mazard 2 327 
Mead-sweet 249 
MEADOW-BEAUTY 
FAMILY 2 50. 
Meadow-beauty 583-4 
Meadow-bouts 2 85 
-buttercups 85 
-cabbage I 445 
-cup 2 202 
-fern or -bur 
I 584, 586 
-gowan 2 85 
ee I 253-7 
Annua 253 
Arctic 267 
Common 256 
Dwarf 253 
Flat-stemmed 259 
Fowl 255-6, 264 
Frisky I 272 
Grove 257 
Prairie 255 
Reed 265 
Round-stalked 255 
Rough-stalked 255 
Sea 267-8 
Short-stalked 239 


Slender, or Spread- 


ing 268 
Torrey’s 268 
Water 265 
Wavy 254 
Wood 256 

Meadow-nuts 2 258 
-Meadow-parsnip 2 639 
Early, Golden, 641 
Meadow-pink 
1 558; 2 74 
-queen 2 249 
Meadow-rue 118-21 
Arctic, Dwarf 118 
Cohosh 119 
Early 121 
Fall 121 
Mountain 118 
Slender 118 
Tall or Purplish 120 
Veiny 120 
Waxy 119 
Wite tower 2 102 


Meadow-scabish 3 419 
Meadow-sweet 

2 245, 248-9 

American 245 


INDEX, INCLUDING POPULAR PLANT 


Meadow-sweet 


Birch-leaved 2 246 
Meadow-wort 2 249 
Meakin 2 614 
Meal-berry 2 693 

-plum 693 
Mealy Starwort I 511 
Mealy Tree 3 271 
Mecha Meck 3 43 
Mechoacanna 3 43 
Medaddy-bush 3 281 
Medic 2 351-2 

Black or Hop 351 

Purple 351 

Toothed 25355 

Spotted 352 
Medlar 2 292 
Meehania 3 113 
Meeting-houses 2 92 


Melampodium 3 458-9 


Melanthium I 493 
Melilot, White, 
Yellow 25353 
Melilot-trefoil 22351 
Melmot-berries I 66 
Mentzelia 2 566-8 
Menziesia 2 682 
Mercury, Black 484 
English 2 14 
Herb 460 
Scotch 3 204 
Three-seeded 2 457 
Mercury-weed 45 
2Mermaid-weed 2 613 


Merry, Black Merry 


: ii se 327 
Mesquite, Prai-ie 
(see Grass) “ 333 
Mexican-poppy 2 128 
-rose 41 
-tea 14 
Mexico-seed 2 461 
MEZEREUM FAMILY 
2 574 
Mezereon 574 
American 25575 
Micranthemum 3 197 
Midsummer-men 2 207 


MIGNONETTE FAMILY 


2 199 

Mignonette 2 200-1 
Mile 2 660 
Milfoil Z) cms 
Hooded 3 226 
Water 2 116,615-16; 

3 228 

Milk-gowan 3, 3n5 
-grass 286 
-ipecac 2 469 
-maid 184 
-pea 420 
-purslane 467-9 
Milk-vetch 2 377-87 
Alpine 382 
Arctic 385 
Ascending 379 
Bent 382 
Blake’s 382 
Canadian 379 
Carolina 379 
Cooper’s 385 
Drummond's 381 
Flexile 385 
Hoary 388 
Indian 384 
Long-leaved 386 


Loose-flowered 386 
Low 383 
Missouri 383 
Narrow-leaved 380 
Platte 2.377 
Pretty 384 


Milk-vetch 
Prickly 2 387 
Purple 380 
Racemose 381 
Robbins’ 381 
Sessile-flowered 387 
Short’s 383 
Silvery 388 
Slender 384 
Tennessee 378 
Tufted 386 
Two-grooved 380 


MILKWEED FAMILY 


3 23-35 
Milkweed 2 467,460; 


3 25-9; 317,321,335 


Auricled 236 
Bedstraw 32 
Blunt-leaved 28 
Broad-leaved 27 
Common 30 
Creeping 3 25 
Decumbent 3133 
Dwarf 31 
Few-flowered 3 25 
Florida 35 
Four-leaved 29 
Green 34 
Hairy 27 
Intermediate 28 
Low 32 
Marsh 357 
Mead’s 3 28 
Narrow-leaved 34 
Oblong-leaved 33 
Oval-leaved 31 
Poke 29 
Purple 26 
Red 26 
Running 3 38 
Sand 30 
Short-crowned Bin 
Showy 30 
Sullivant’s 27 
Swamp 26 
Tall 29 
Thin-leaved 31 
Trumpet 319 
Wandering By2r 
White 29 
Whorled 32 
Woolly 35 
Yellow 3 25 
MiLKwort FAMILY 
2 446 
Milkwort, Bitter 2 451 
Cross-leaved 2 448 
Curtiss’ 450 
Dwarf 452 
Field 2 449 
Fringed 452 
Loose-spiked 449 
Low Pine-barren 447 
Marsh 448 
Maryland 450 
Nuttall’s 450 
Orange 447 
Pink 2 449, 451 
Purple 2 449 
Racemed 451 
Sea 716 
Short-leaved 448 
Tall Pine-barren 447 
White 2 451 
Whorled 448 
Yellow 447 
Milky Tassel 4 307 
Millet i 121 
Broom-corn I 140 
Brown 140 
Cat-tail 166 
Dotted I 124 


NAMES. 621 


Millet, Egyptian 1 121 
Evergreen 121; £ 276 


German, Golden 166 
Hungarian 166 
Indian 174-5 
Italian 166 
Seaside 133 
Tall 173 
White 2 352 
Wild 165,174 
Yellow 2 353 
Mill-mountain 2 438 
Mimosa Family 2 330 
Mimosa 332 
Mingwort 3.525 
Minnie Bush 2 682 
Mint FaMity 3 99 
Mint 
American Wild 3 152 
Apple 150 
Balm 3.35,(age 
Bergamot 3 150 
Brandy 149 
Brook 150 
Brown 149 
Cat 113 
Common 149 
Corn 152 
Creeping Whorled 
152 
Crisped-leaved 151 
Cross I51 
Curled I51 
Dog 138 
Downy Whorled 152 
European 150 
Field 152 
Fish 150-1 
Garden 149 
Horse 3 131, 134-5, 
146, 150 
European 150 
Ohio 135 
Lamb 3 149 
Mackerel 149 
Marsh Whorled 152 
Mountain 
3 131, 138, 142-5 
Our Lady’s 3 149 
Patugonia 150 
Peppet 3 149 
Round-leaved 150 
Small-leaved 152 
Spear Sings 152 
Squaw 3754 
Stone 146 
Water 150-1 
Whorled 153 
Wild 3 150 
Woolly 151 


Missionary-weed 3 334 
Mire-blobs 2 85 
Mist 72 
Mist-flower 3 362 
MISTLETOE FAMILY 

I 638-9 


Mistletoe 1 638-9, 711 
Missey-moosey 2 287 
Mitchella 3 255 
Mitrewort 2 229, 731 
False 2 224 
Moccasin Flower I 550 
Yellow 550 
Mock-apple 3 2902 
-Bishop’s-weed 2 657 
-gilliflower 2 73 
Mock-orange 2 231-2; 
3 292, 719, 723 
Mocker-nut I 582 
Modesty 2 525, 639 
Moehringia 2 57-8 
Mohawk-weed I 518 


622 


Moldavian Bald 3 115 
Mole-plant, -tree 2 471 


Molinia 14237 
Monarda, Brad. 
3 131, 133-4 
Money-grass 89223 
-plant 2 191 
Moneywort 20712 
Prairie 714 


Monkey-flower 3 190-1 
Monkey-nut Tree 2 512 
Monkey’s Face 2 56 
Monk’s-head 3 315 
Monkshood 2 96-7,142 

Trailing 97 
Monk’s Rhubarb 1 656 


Monniera 3 192-3 
Monolepis 7A ai) 
Mouth-flower 2°17 
Moon-fern ig 2 
Moon-flower 318,518 


Moon-fruit Pine 1 44 
Moon-penny 3 518 
MooNnSEED FAMILY 


2 130 
Moonseed 2 131 
Red-berry 2013 
Moonshine 3 400, 453-4 
Moonwort 13 
Cut-leaved 5 
Hemlock-leaved 6 
Underwood's 4 
Moor 2 694 
Moor-berry 704 
-grass 1 487; 2 203 
Moorwort 2 689 
Moose-bush, -berry - 
3 269 
Moose-Elm I 627 
-misse 2 287 
-wood 497,575 
Morass-weed 275 
Morel 3 164 
Petty 2 617; 3 164 
Morgan 3 516 
Dutch 518 
Morgeline 3 203 
Mormon-weed 2 52% 
MoRNING-GLORY 
FAMILY 3 40 
Morning-glory. 3 44-5 
Bush, ,/ 44 
Dwari - 47 
Ivy-le*ved 3 45 
Poet 3 44 
Red 43 
White 44 
White Star 44 
Morocco, Red 2 Wee 
Mortification-root 
2 514 
MoSCHATEL FAMILY 
' 3 283 
Moschatel 3 283 
Moss, Black I 456 
Club I 43-7 
Cypress I 46 
Dwarf Club 49 
Fir I 43 
Floating 38 
Florida I 456 
Flowering 
2 209,706; 3 57 
Golden 2 208 
Ground I 306 
Hanging 456 
Iceland 210 
Irish 2 474 
Kentucky 2 41 
Long I 456 


Mountain I 49; 2 209 


Pixie 706 
Prickly Club I 49 


Moss, Rock 2 209 
Rose 2 41,519 
Running, Snake I 49 


Spanish 456 
Stag Horn I 49 
Tree 143; 2474 
Wall 2 208 
Moss-berry 2 704 
-bush 686 
-campion 2 63 
-crop I 324, 326 
-locust 2 375 
-milion (melon) 704 
-phlox 3 58 
-pink 3 58, 63 
-plant 2 686 
-rose 2 519 


Mossy Stonecrop 2 208 
Mother-of-thousands 


3 175 
-of-thyme 3139, 141 
-of-wheat 3 203 
Mother’s-heart 2 158 
Motherwort 
3 120, 357, 527 
Golden 3 456 
Hoarhound 120 
Man’s 2 461 
Siberian 3 120 
Mountain Ash 2 287 
Eider-leaved 287 


Mountain Avens, see 
Avens 

-bramble 2 276 
-fringe 143 
-joy 3 140 
-laurel 2 681, 684 
-lover 492 
-mahogany I 609 
-moss 208 
Mountain-mint 3 231 
Awned 144 
Basil 143 
Hairy 142 
Hoary © 144 
Hyssop 143 
Narrow-leaved 142 
Short-toothed 145 
Southern 144 
Thin-leaved 145 
Torrey’s 143 
Virginia 142 
White-leaved 145 
Mountaim-rice I 174-5 
-nut 3 80 
-snow 2 469 
-sweet 504 
-tea 2 693 
Mouse-ear 2 48, 101; 
3 85, 128, 455 
Spring 2 47 
Virginia 3 78 
Water 42 
-chickweed 47-8 
Mouse-ears 2 Lor 


Mouse-ear Cress 2 176 
Mouse-milk 473 
Mouse-tail 

I 192, 269; 2 103 


Little 2 103 
Mouse-thorn 3 559 
Mouth-root 2 88 
Moxie-berry 704 
Muckweed 

Curly tS1 532.550 
Mud-flower 3 197 

-plantain I 463-4 

-purslane 2 538 


-weed, or -wort 3 198 


Mugget, Mugwet, 267 
Mugwort 3 525-9 
Common 527 


eee 


Mugwort 
Cudweed 3 529 
Dark-leaved 529 
Kansas 528 
Long-leaved 528 
Mexican 529 
Prairie 529 
Saw-leaf 528 
Western 529 
Muhlenbergia I 184 
MULBERRY FAMILY 630 
Mulberry 1 631; 2276 
Bermuda 3 99 
French 99 
Otaheite I 632 
Paper 632 
Mullen 173-4 
Moth Reis 
Sage-leaf 3 118 
Mullen-pink 2 61, 71 
Musk 2. oom 
Wild 2 430 
Musk-crowfoot 3 283 
-flower IQ! 
-plant .2 515; 3 I91 
-root 3 283 
Muskrat-weed 2 121 
Musquash-root 2 658 
-poison 2 658 
-weed 2 T2% 
Musineon 2 643-4 
MusTArD FAMILY 2 146 
Mustard 2 174 
Ball 2 ™59 
Black 2 193, 195 
California 2 174 
Corn 192 
Dién 168 
‘alse 199 
Garlic 169 
Hedge 169,170,174 
Indian 2 193 
Mithridate 164 
Orange 173 
Poor Man’s - 170 
Tansy 171 
Tower 181 
Treacle 172,175 
White I9I 
Wild 192, 195 
Wormseed 172 
Myagrum 2 157, 168 
My Lady’s Belt 2 249 
Myriad-leaf 2 614 
Myrtle 20 
Bog 1 584;3 18 
Burren 2 693 
Burton I 584 
Candleberry 585 
Crape 2 580 
Dutch, Moor 1 584 
Running 3 20 
San 2 682 
Sweet I 446 
Wax 585 
Myrtle-flag 446 
-grass, -sedge, 446 


Mysterious Plant 2 574 


Naias I 89-90 
Nail-rod 426 
Nailwort 2 28, 148, 158 
Naked-weed 3 314 
Nama 3 71-2 
Nanny-berry 3)273 
-bush 273 
-plum 273 
Nap-at-noon 
I 509; 3 313 
Nape 2 193 
Napoleons 2-355 
Natural-grass 2 351 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Vox, III. 


Nature’s-mistake 2 664 


Navarretia 3 64 
Navelwort 2 614, 649 
Navew, Wild 2 193 


Necklace-poplar 1 590 


Necklace-weed 2 90 
Neckweed 
I 634; 3 202 
Needle-chervil 2 627 
-and-thread i503 
Neale 282 
Negro-vine 3 38 
Nelumbo, Amer 2 77 
Nemastylis I 541 
Nemophila 3 67 
Nep yea ie) 
Nerve-root 1 549-50 
Water 3 26, 38 
Neslia 2 159 
Nest-root 2 674 
Nestronia I 641 
NETTLE FAMILY I 634 
Nettle I 635-7 
Bee 3 119, 122 
Biind 119-22 
Bull 165 
Burning I 635 
Canada 636 
Day 3 121-2 
Dead, Dog, Dumb 
I1g—22 
Deaf I2I 
Dwarf I 635 
False 637 
Flowering 3 119 
French 121 
Great I 635 
Hemp 3 119, 
Horse 165 
Sand 2 462 
Slender I 635 
Small 635 
Spurge 2 462 
Stinging 
I 6353; 3 92,119 
Stingless I 367 
Tall Wild I 635 
Texas 3 166 
Weak I 636 
Wood 636 
Nettle-potato 2 461 
Nettle-tree I 629 
Networt 570 
New Jersey Tea 2 504 
Nickar-tree 340 
Nigger-head 
I 528; 3 470 
Nigger-weed 357 
Nightshade 3 164-7 
American 226 
Beaked 3 166 
Bindweed 2 611 
Bitter 3 167 
Black 164 
Climbing 3 167 
Cut-leaved 3 165 
Deadly 164 
Enchanter’s 2 611 
Fetid 3 168 
Garden 164 
Melon-leaved 166 
Prickly 166 
Silver-leaved 165 
Three-leaved 1 523 
Torrey’s 166 
Viscid 167 
Woody 167 
Nimble Kate 293 
Nimble-weed 2 100 
Ninebark 2 244 
Ninety-knot I 661 
Nipple-wort 3 306 


Vor. III.. 
Nipplewort 

Dwarf 3 309 
Noble-pine 2 672 
Nit-weed 2 536 
Noah’s Ark I 550 
Nondo 2 647 


None-so-pretty 
2 66,563; 3 453 


Nonesuch 2 69, 351 
Black @ 351 
White I 281 

Noon-flower By Eine, 

Noontide 313 

Nosebleed 

15253 3 214, 515 
Notholaena 735 
uns CU AAAI 
‘Nurse-garden 290 
_Nut-grass 1 304, 306 
a -rus I 349-51 

Nuttallia 2 567 
4 Nuttall’s-weed 3 493 

Nyctelea 3 67 

Nymph, paste 2 79 

_ Oak I 617-2: 
Barren Po 
Bartram ' 621 

. Basket 624 

Bear 620 

. Bitter 620 

Fd -bush 620 

ack 1 617, 618, 619 

| Dwarf 620 
| {/ iad jack 620, 621 
ei -)serab ie 


6: 
\ “2 486, 620 
I 620, 622 


a4 


Re ¢ 
2, AOC 624 
Rock Chestnut 624 
oy Rou 1, Rough White 


Sand Jack 621 


carlet 619 
_ Schneck’s 618 
Serub 620, 623-4 
_ Serub-chestnut 625 


620, 622% 


+ \, 6215629" 
_ ™.~ 622 | Old Ladies’ Clothes- 


| 
aa 
| 


622 
* ‘ Running White rer 


Oak, Shingle I 622 
hrub 624 
Spanish 617, 619 
Spanish-water 619 
Spotted 619, 621 
Stave 622 
Swamp 617,621, 623 
-chestnut — 625 
-spanish 617 
-white 623 
-post 623 
Sweet 585 
Tan-bark 624 
Turkey 619, 622 
Water 621 
Water White 623 
White I 622 
Mossy-cup 623 
Willow 621 
Yellow 618, 624 
-chestnut 624 
Yellow-bark 619 
Oak-fern I23 
Oat I 120, 217-19, 
518-19 
False 217 
Haver, or Poor 
218, 220 
Hooker’s 219 
Pearl 220 
Purple 219 
Sea, or Seaside 249 
Smith’s 219 
Wild 218 
Oat-grass I 177,218 
Black 177 
Common 220 
Downy 217 
Evergreen 220 
False, or Golden 
217, 220 
Fall. x 287,570, 641 
Tennessee 220 
Wild 120, 220-2 
| Oats, Wild 1 211,218 
Obedient-plant 3 116 
Oceanorus I 493 
Ocymum 144 
Ohio Cucuma 2 85 
Oil-nut I 579 
Oil-plant 2 461 
-seed, Siberian 157 
Olcott-root oven SSS 
Oid Goose 550 
Old Maid’s-bonnets 
2 348 
-pink 66, 73 
-root 617 
_Oldenlandia 3 254 


Old Field-sweet 3 426 


pins 3 497 
Old Man 3 526 
Old Man’s-bear 2 728 
-flannel B 173 
-night-caps 426 
-pepper 3 515 
~root 2 617 
-virginia 3 426 
Qleander, Wild 2 580 
OREASTER posers 
2 575 
C wi 2 576-7 
W 576-7 
O1 ths x 2 724 
Olive, Spurge 574 
Olive-tree. 76 
Bell, or Wild 722 
eben I 629; 
2 603; 3 2 
One-blade : $17 
One-leaf 


Onion, Wild r498-c0o 


ENGLISH. INDEX, INCLUDING POPULAR PLANT 


Onion 
Alleghany I 498 
Bog 442 
Nodding 498 
Prairie 498 
Shore 498 
Ooler I 613 
Oonopsis ZEN i/ 
Open Gowan 2 85 
Opium, Wild 3 320 
Opossum-tree 2 235 
Opuntia 573 
Orache 2 18 
Orange, Spring 723 
Mock 719 
Wild 2 444, 617 
-blossom TS 25 
Orange-flower Tree 
2 232 
Orange-grass 536 
-mustard 173 
Orange-root 2 85 
Climbing 493 
OrcHID FAMILY I 547 
Orchis 551-9 
Bog I 554, 556 
Brocted Green 552 
Crane-fly 575 
Crested Yellow 557 
Dwarf 556 
Feather-leaved 557 
Fen 572 
Flaming 559 
Fringeless Purple 
559 
Gay 551 
Great Purple 559 
Greenish 552 
Green Wood 552 
-fringed I 558 
Green Rein 552 
Hooker’s 556 
Heal-all 555 
Leafy Green 554 
Long-bracted 552 
Meadow Pink 550 
Northern Bog 556 
-green 554 
small 55 
-whitt 554 
One-leaved 556 
Pink Fringed 559 
Prairie White- 
fringed 558 
Preacher-in-the- 
Pulpit I 552 
Purple 551 
Purple-fringed 558 
Early 558 
Smaller 559 
Ragged 550, 558 
-fringed 558 
Rein Sse) 
Green 552 
Round-leaved 555 
Small 551 
Showy 551 
Small 4 PaleGreen 552 
-two-leaved 556 
Southern White 552 
Yellow I 552- 
Spring 55 r 


Tall White Bog 554 


Tattered-fringe 558 
Three-toothed 553 
Tubercled 552 
Two-leaved 
Large 555 
Small 556 
White-fringed 557 


Jestern-greenish 
558 


Yellow-fringed 557 


NAMES. 623 
Orchis, Yellow 1 552 
Oregon-grape 2 128 
-tobacco 3 285 
Oreocarya 3 81-2 
Organs, Organy 
3 110, 140 
ORPINE FAMILY 2 205 


Orpine, American 


207-8 
Garden 207 
Orthocarpus, Yellow 
3 216 
Oryzopsis fr 174 
Osage 632 
Osage-orange 632 
-apple 632 
Osier 595-7 
White 2 689 
Ostrich-fern I it 
Oswego Tea B03 5 
Othake 507 
Our Lady’s-mint 149 
-bedstraw 254 
-thistle 560 
Owler I 613 
Owl’sCrown 3 447,456 
Oxadoddy 204 
Oxalis 2 430-2 
Ox-balm 20053 
Ox-eye 3510, 518, 467 
Great-white 3 as 


Sea 
Ox- a 3.470, si8 
Oxford-weed 
Ox-tongue, Bristly 


3 311 
Ox-wort 3 532 
Oxypolis 2 638 
Oxytrope 2 3890 
Arctic 3890 
Showy 390 
Silvery 390 
Tufted 389 
Yellow 390 
Oyster-plant 3 82, 313 
-root 315 
Paddock-pipes 
LE40; AT 5 +2652 
Padelion 
2° aGA\*) 3-214 
Paint-brush 2 530 
Paint-root T5391 
ie Cup 3 214-16 
Pal 215 
Poe Christi 2 461 
Palmillo 512 
Palsy-wort 2 85 
Panicum, Panic-grass 
I 138 
American 15 
Ashe’s 161 
Barbed 150 
Beaked 142 
Bicknell’s 145 
Bitter 141 
Blunt 138 
Bog 142 
Bosc’s 162 
Broad-leaved 164 
Bush’s 147 
Clute’s 151 
Commons 156 
Dense 142 
Dwarf 144 
Early-branching 154 
Eaton's 152 
Eight-jointed 151 
Few-flowered 158 
Flat-stemmed 142 
Forked 149 
Gaping 163 
Gibbons 163 


624 


Panicum 

Grisebach’s I 146 
Hairy 153 
Hemlock 157 
Kunth’s 147 
Lax-flowered 146 
Lieberg’s Tila) 
Lindheimer’s 152 
Long-leaved 143 
Long-stalked 144 
Loose 133 
Low, Stiff 156 

Low, White-haired 
144 
Matting 154 
Narrow 000 
Narrow-leaved 146 
Nash’s 157 
Northern 150 
Owen’s 155 
Porter’s 000 
Purple I51 
Ravenel’s 159 
Red-top 142 
Ringed 150 
Rough-hairy 145 
Roughish 152 
Round-fruited I 147 
Scribner's 158 
Slender 158 
-stemmed 154 
Small-fruited 148 
Small-leaved 148 
Soft-leaved 160 
Spotted-sheath 149 
Spreading 138 
Starved 143 
Stiff-hairy 153 
Tall-flat 143 
-fringed 161 
-rough 160 
-swamp 160 
Tennessee 155 
Variable 161 
Velvety I 159 
Warty 138 
Werner’s 144 
White-edged 148 
White-haired 153 
Wilcox’s 159 
Wiry 140 
Wood 140 
Woolly 155 
Pansy 2 547, 562-3 

Pawpaw, North Arn. 
2 83 
Paper-mulberry 1 639 
Papoose-root 2 128 
Paradise-plant 574 


Paraguay (false) 3 272 


Parilla, Yellow 2 131 
Parosela 2 367-8 
Golden 368 
Low 368 
Pink 367 
Slender 367 
Woolly 367 
Parsley 2 632-3 
Beaked 629, 630 
Carrot-leaved 632 
Common 641 
Dog 630, 633, 645 
False, Fools 2 645 
Garden 641 
Hairy 632 
Hemlock 636 
Horse 634 
Marsh 2 660 
Poison 653 
Sand 652 
Sea 648 
Spotted 653 


White-flowered 631 


Parsley, Wild 2 641 
Parsley-breakstone 
2 264 
-piert 30, 264 
-vlix 264 
Parsnip 2 625, 634 
Wild 2 634, 658 
Cow 635 
Meadow 641 
Water 655-6 
Wild 2 634, 658 
Parthenium 
Auricled 3 465 
Creeping 446 


airy 466 
Partridge-berry 2 693, 
794; 3 255 

698 


Mountain 
Partridge-pea 2 337 
-vine 3 255 
Paspalum I 126-133 
Bosc’s 131 
Bush’s 128 
Ciliate-leaved 128 
Field 130 
Florida 132 
Joint-grass 133 
LeConte’s 132 
Long-stalked 127 
Many-haired 130 
Muhlenberg’s 129 
Prostrate 128 
Pubescent 129 


Round-flowered I 131 


Slender 129 
Smooth-scaled 131 
Straw-colored 127 
Tall 132 
Walter’s 127 
Water 126 
Yellow-haired 130 
Pasque-flower 2 102 
PassioN FLOWER 
FAMILY 2 564 
Passion Flower 565 
Passions I 656 
Passion-vine 2 565 


Patience,Garden I 656 
Paulownia 3 189 
Paul’s Betony 


3 201-2, 204, 148 
Pauson 2 140 
PEA FAMILY 2 341 
Pea, Beach 413 

Bird-egg 386 
Buffalo 377, 409 
Bush 343-4 
Butterfly 417 
Canada 409 
Cat 409 
Cow 425 
Craw, or Mouse 415 
Crow 412, 479 
Everlasting 
2 413-14, 416 
Ground-squirrel 129 
Heart 501 
Hoary 372 
Meadow 415 
Milk 420 
Myrtle-leaved 
Marsh 414 
Partridge 2 337 
Potatoes 418 
Rabbit 372 
Sea-side, Sea 413 
-everlasting 413 
Scurfy 300-1 
Sensitive 
Squirrel, Ground 
129, 418 
Trailing 2 418 
Turkey 142, 372 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES, 


ery 2 413 
i 347, 41 
Wild, Sweet oy 
Yellow 344 
Pea-nut 394 
Hog, or Wild 419 
Pitcher’s 419 
Pea-tree 2 376 
Pea-vine oe .. 450 
PEACH FAMILY 322 
Peach 2 330 
Peachwort I 668 
ear 2 287, 292 
Choke 288 
Juice 292 
May 292 
Prickly 571-2 
Swamp Sugar 292 
Wild 292 
-indian 292 
Pearl-grass I 250 
Pearls of Spain {its 
Pearl-plant 3 87-8 
Pearlwort 2 51-2 
Knotted 52 
Pearly Everlasting 
3 451 
Peatweed 2 580 
Pebble-vetch 2 411 
Pecan I 580 
Bitter 581 
Pectis 2) uy 
Pedicularis 3 220-3 
Peg-wood 2 492 
Pellas 515 
Pelican-flower 1 645 
Pellitory 


1 638; 3515-519 
Bastard, Wild or 
European 3 515, 519 
Pellitory of Spain 2 638 
Pencil-flower 393-4 
Pencil-tree 


-wood I 66 
Penny-cress ° 2 168 
Penny-flower IgI 

-hedge 170 
Penny-john 533 
Penny-grass a) 223 
Penny Mountain 141 
Pennyroyal 

3 136, 152, 142 

American 136 

Bastard 104, 108 

Drummond’s 136 

False 104 

Mock, Rough 136 

Wild 152 
Penny-post 2 650 
Pennywort 3 15,175 

Marsh 648-50 
Pepper, Poor Man’s 

2 208 

Birds 165 

Glen 164 

Old-man’s 3 515 

Water I 669 


Wild 2 504, 526,574 
Pepper and Salt 2 653 


Pepper-bush 667 
Sweet 667 
Water 691 

Pepper-crop 2 i 

Pepper-grass I 545; 
oe 2 164, 166 
English 2 164 
Garden 166 
Golden 166 
Narrow-leaved 165 
Roadside 165 
Town 166 
Wild 165-6 

Pepperidge 2 665 


Voz. III. 
Pepper-bush 2 127 
Peppermint 3 1490 
Pepper-plant 
I 670; 2 158 
Pepper-root 2 187-8 
-turnip I 442 
-vine 2 510 
-wood 444 
Pepperwort I 37 
European 37 
Hairy 37 
(see Peppergrass) 
2 164 
Perilla 3 154 
Periwinkle 20 
Small 20 


Persian-berry 2 504 
Persicaria, persicary 


ue I 666 
Biting 670 
Bristly 669 
Carey’s 668 
Common 668 
Dense-flowered 667 
Dock-leaved 666 
Lady’s Thumb 668 
Long-styled 667 
Opelousas 669 
Pale I 666 
Pennsylvania 667 


Slender Pink f 1632 
Southwestern 668 


Swamp 666 
Water 666 
Persimmon 2 720 
Pestilence-wort 3 532 
Peters-Staft 173 
Petty Morel 2 617 
Petunia 3.475 
Peucedanum 2 632 
Pewterwort Ne: 
Phacelia . 68 
Coville’s 70 
Crenate-leaved 065 
Franklin’s 69 
Fringed 71 
Hairy 70 
Loose-flowerel 69 
Mountain 7 
Pursh’s 70 
Silky j 68 


Small-floweted 69 
Pheasant’s-ey: 


Phippsia I 193 


Phleum, Mointain 191 
PuHitox FAM Ly 3./52 
Phlox 3453 
Britton’s 58 
Chickweed 57 
Cleft 56 
Crawling 56 
De ny 55 
Dougias’ 59 
Garden 53 
Hairy 35 
Hood’s “<5 
Kelsey’s Ss 
Large-leaved 54 
Moss 58 
Mountain . 54 
Prairie 54 
Smooth 55 
Wild Blue . 56 
Wood's 927 
Yellow 2 173 
Phthisic-weed 2 585 
Phyllanthus 2 453 
Physalis 3 156 


Physic, see Indian 

Picac 2 469 
Pickaway-anise 2 445 
Pick-cheese 2 514 


> % 


lo? 


Vot. III. 


PICKEREL WEED 
FAMILY 
Pickerel Weed 

© 463% 2 106 


I 462 


Pickpocket 2 158 
Pick-purse 2 59,158 
Pick-tree 2 617 
Pickle-plant 2 21 
Picradenia 3 508-10 
Picris 3,305 
Picry 2 484 
Be aarkar 2 521 
Pie-print 2 521 
Pigeon-berry 2 26, 
278, 292, 618, 
663; 3 255 
-foot 2 429 
-grass 3 95, 165 
-root 2 38 
-tree 2 617 
Pigmy-weed 2 206 
Pig-nut I 583 
Bitter 580 
‘Little, Small 583 
Woolly 584 
Pig-potato 2 638 
-root I 543 
Pigsty-daisy 3 516 
- Pig-tail 3 259 
Pigweed 
2 2, 10, 13, 40, 590. 
Rough 2 2 
Slender 
Winged 2 16 
Pilewort ..2 112,117; 
3 179, 536 
Pilot-weed 3 461 
Pimentary 137 
Pimpernel, Bennet 
2 631, 655 
Bastard 2 717 
Blue 3 106 


False 2717; 3196-7 
Mountain 2 631 
Red, or Scarlet 2 716 


Sea 58 
Virginia 631 
Water 2 710; 3 200 
Yellow 2 640 
Pimpernelle 2 266 
Pin-ball ©. Aor 
Pin-clover 2 430 
-grass 430 
Pincushion 3 290, 451 
-flower 290 
-shrub 2 492 
-tree 2 270 
PINE FAMILY 4 asi 
Pine I 56-63 


Banks’ or Black 58 
Black-norway 59 


Bastard 59 
- Blister 63 
Bull I 57,58 
Canadian I 57 
Candlewood 59 
Carolina 58 
Cat 61 
Cedar 58 
Cornstalk 59 
Deal 56 
Fat By 
Festoon 47-8 
Field 2 541 
Fir I 63 
Florida 57 
Foxtail 59 
Frankincense 59 
Gambier Parry’s 57 
Georgia 2 ye) 
-pitch,-yellow 57 
Tray I 58 


Pine, Ground 1 47, 48; 
25536" S102 
Hard I 57,59 
Hart oa 
Hickory 59 
Hudson Bay 58 
Indian 59 
ack 58 
Jersey, Labrador, 
I 58 
Loblolly 59 
Long-leaved BF, 


-pitch, -yellow 57 


Long-straw 57, 59 
Long-shucks 59 
Moon-fruit I 44 
Nigger 58 
Noble 2 672 
Northern 1S 
Northern Scrub 1 58 
Norway 57 
Black 58 
Old-field 59 
Pitch 57, 58-9 
Southern 57 
Western 57 
Pond 60 
Prairie 57 
Prickly 59 
Prince’s 2 672 
Red T5771 
River 58 
Rock 57,58 
Rosemary 59 
Running I 47 
ap 59, 675 
Scotch o 
Sweet 00 
Scrub 58 
Shore 58 
Short-leaved 58-9 
Shortschat Taso 
Short Shucks 1 58 
Slash 58, 59 
Soft TSG 
Southern Tes7, 
-hard or-heart 57 
-mountain 59 
-pitch 57 
-yellow wy) 
Spiral 45 
Spruce 
56, 62, 58, 61-2 
Swamp 59 


Table-Mountain 59 


Texas Yellow EVA 
Torch 59 
Turpentine ty 
Virginia 159; £139 
Western-pitch 57 
Weymouth 56 
White 56 
Yellow I 57,59 
Yellow-norway 59 
Yellow-pitch 57 
Yew 61 
PineE-APPLE FAMILY 
45 
Pine-broom 57, 
Pine-barren Beauty 
2 706 
Pine-cheat 2 59 
Pine-drops 2 673 
-sap 2 674 
Pine-tulip 672 
-weed 2 536 
Pink FAMILY 2 61 
Pink, Bearded I 563 
Boston 273 
Bunch 74 
Carolina 730 
Childing 2 74 | 
Chimney 2 73 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Pink, Corn 2 61 
Cushion 63 
Deptford 74 
Drummond’s 2 70 
Dutch 2 200 
Election 3 678 
Fire 2 64 
French 

266,75; 3559 
Grass 2 ‘74,563 


Ground 2 73; 3 57 


Hedge 257385 
42,214 
Indian 2 64, 69, 452, 
: 7303, 3.42 
Maiden 274 
Marsh 3 6-7 
Meadow Tes5s- 
2 69, 74, 679 
Menzies 2 68 
Mice 2 65 
Moss 63; 3 57 
Mountain 
_ 2 692; 3 57 
Mullein 2 61, 75 
Old Maid’s 2 61,73 
Proliferous 74 
River 2 678 
Rock Fi Be 
Rose 35 
Saxifrage 372 
Sea 2 64; 3 6 
Sheriff 3 518 
Swamp) 1 488, 563; 
2 678 
Wax 2 41 
Wild I 562; 
2 OS SiAis 3 357 
Winter 2 692 
Pink-bloom AE 
-grass I 241, 415, 391 
-needles 2 430, 625 
Pink-purse 59 
Pink- Se g 730 
-sir 2 331 
Pinks, ‘Old Maid’s 66 
Wax 4I 
Pinkster-flower 678 
Pink-weed 1 661, 668 
Pinweed 2 430, 542-5 
Pipe-plant 674 
Pipe-privets 724 
Piperidge-bush 2 127 
Spotted 672 
Piperia I 555 
Pipe-stem 2 690 
Pipe-tree 724 
Pipe-vine I 646, 724 
Woolly I 646 
PreEworRT FAMILY 453 
Pipewort 454-6 
Hairy 456 
Yellow 455 
Pipsissewa 2 672 
Spotted 672 
Piskies, Pixie 2 44, 706 
Pismire f 4560 
Pissabed 2 112 
Pistachis 235 
PITCHER-PLANT 
FAMILY 2 207 
Pitcher-plant 2 207 
Pitchforks 3 495-7 


PLANE-TREE FAMILY 


2 242 
Plane-tree 242 
Planer-tree I 628 


PLANTAIN FAMILY 
3 245 
Plantain 3 245-9 
Buck, Buckhorn 246 
Common 3 245 
Dwarf 248 


625 


Plantain 

Dooryard 24 
* English : ae 
Greater 245 
Heart-leaved 247 
Hoary 246 
Indian 3 536-8 
Large-bracted 248 
Lance-leaved 246 
Long 246 
Many-seeded 249 
Mud I 463 
Narrow 246 
Net-leaf 570 
one 3 246 
etty 9 
Poor Robin’s = 
3 332,439 
Pursh’s ae 
Rattlesnake 1 569-70 
Ripple 3 246 
Robert’s 3 439 
Robin’s 439 
Rose 439 
Round-leaf 245 
Rugel’s 3 246 
Sand 249 
Saline 247 
Sea, or Seaside 247 
Slender 249 
Snake 3 246, 322, 332 
Spotted 570 
Sweet-scented A 475 
Water 1 98; 3 247 
Wayside 245 
White I 569; 
3 248, 451 


White Dwarf 3 248 
Plantain Shore-weed 


3 250 
Pleurisy-root 5 
Pleurogyne 3 14 
Pleuropogon I 247 
Ploughman’s-wort 
3 448 
Prum FAMILY 2 325-6 
Plum 322 
Beach 2 325 
Bullace 2 326 
Bunch 664 
Canada 2 323 
Chickasaw 2 324 
Date 720 
Dog 1515 
Earth 2 376 
Garden 326 
Wild 2 324 
Goose 2 323 
Grave’s Beach 2 325 
Ground 377 
Hog 2 323-4 
Horse 2 323 
Ivory 2 693 
Larger Ground 377 
Low 325 
Native 2 323 
Porter’s 2 324 
Red 2 323 
Sand 2 325 
Seeded 720 
Thorn 297, 302, 317 
Wild 2 323-4 
Wild Goose 324 
Wild Red 323 
Wild Yellow 323 
Winter 720 
Plum-bog 2 278 
-granite 323 
Plum-grape 2 506 
PruMBAGO Fam, 2 718 
Plume-grass L. ¥t3 
Pocan-bush 2 26 


626 


Pogonia I 560-1 
Adder’s-mouth 560 
Nodding 562 
Whorled 561 

Poison-ash 2 483 
-arum I 444 
-berry 2 90; 3 167 
-dogwood 2 483 
-elder 483 
-flag I 537 
-flagroot 537,539 
-flower 3 167 
-hemlock 2 653 
-ivy 483 
-oak 483-4 
-parsley 2 653 
-rhubarb 3 470 
-snakeweed 2 653 
-sumac, ‘* 2 483 
-tobacco 3 169 
-tree, or-wood 2 483 
-vine 2 483 

Poke-root I 494 

POKE-WEED FAMILY 

2 25 

Poke, Pokeweed 26 
Indian I 494-5 

Polar-plant 3 461 

Pole-cat Weed 445 

Pole, or Pool-rush 331 

Polk-weed 445 

Pollom, Red 2 693 

Polly Mountain 3 139 

Polypody 35 
See Fern 

Polypremum 2 731 

Polypteris 3 507 

Polytaenia 2 633 

Pomegranate 323 

Pomme Blanche 2 363 

Pond-bush 25135 
-dogwood B0255 
-grass 1 $7 
-spice 2 135 

Pond-lily 2 78-80 
Arrow-leaved 79 
Red-disked 78 
White 9-80 
Sweet-scented 79 
Yellow 78 

PONDWEED FAMILY I 74 

Pondweed I 75-87 
Alga-like 81 
Blunt-leaved 82 
Choke 105 
Clasping-leaved 80 
Common Floating 75 
Curled-leaved 81 
Curly 81 
Eel-grass 1 81 
Faxon’s 78 
Fennel-leaved 87 
Filiform 86 
Fries’ 83 
Grass-wrack 81 
Hill’s I 82 
Horned 89 
Tllinois 79 
Inland 86 
Interrupted 87 
Large-leaved 76 
Leafy 82 
Long-leaved Tif 
Mystic Pond 80 
Northern 77 
Nuttall’s ae, 
Oakes’ 76 
Opposite-leaved 1 84 
Rafinesque’s 85 
Robbins’ 87 
Shining 79 
Slender 83 


Pondweed 
Small r 84 
Spatulate-leaved 78 
Spiral 86 
Spotted 76 
Tassel 88 
Thread-like 85 
Various-leaved 78 
Vasey’s 83 
White-stemmed 80 
ZAZS 79 

Pond Spice 2137 

Pool-blossom 3 18 


Pool-root,-wort 3 362 
Poor Annie I 494 
Poor-land Weed 3 256 


Jan’s Leaf 2 211 

Poor Man’s-mustard 
2 170 
-pepper 2 164, 208 
-rhubarb 2 120 
-soap 2 245 
-weather-glass 2 716 

Poor-Robin 

2 69; 3 259 

Poor Robin’s-plantain 
3 285, 388 
Pop-dock, or -glove 204 
Pop-weed 3 204, 220 
Poplar I 587-591 
American 590 
Aspen 587 
Balsam I 588 
Berrybearing 590 
Black I 590 
-Italian 590 
Blue 2 83 
Cat-foot 590 
Carolina 588, 590 
Downy I 589 
Hickory 2 83 
Lombardy I 590 
Necklace 590 
Old English 590 
Ontario 588 
River 590 
Rough-bark 588 

Silver-leaf 

I 587; 2 512 
Swamp 589 
Trembling I 590 
Tulip 2 83 
Water 590 
White I 587, 5890, 
590; 2 83 
Willow 590 
Yellow 2 83 
Poppy FAMILY 2 136 
Poppy 2 64, 136-9 
Arctic 138 
Blue 3557 
Celandine 140 
Corn, Field, Red, 137 
Frothy 2 64. 
Garden, Opium, 137 
Iceland 138 
Leafy-white 139 
Long-fruited 138 
Mexican 139 
Pale 138 
Plume 140 
Prickly 139 
Rouse fruited 138 
Sea 141 
Smaoii-tmured 137 
Spattling 64 
Thorn 139 


White Prickly 2 

Yellow 

Yellow Horned 141 
Poppy-mallow 2 418-19 
Pops 3 204 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Portulaca 2 39-41 
Showy 41 
Possum-haw 
2 488; 3 273 
Possum-wood 720 
Potato FAMILY 3 154 
Potato, Canada 486 
Cree 253603 
Dakota 2 418 
Hog’s I 492 
Indian 2 418; 3 482 
Nettle 2 461 
Pig 418 
Prairie 2 363 
Prickly 3 166 
Wild 237, 
Wild Sweet 3 43 
Potato-pea 2 418 
-vine, Wild 2043 
Potereum 2 se 
Plains 


Pot-herb, White 3 28 


Poukenel 627 
Poverty oH Tsk 
Poverty-grass 541 
Poverty-weed 2 503 


3 451, 453, 454, 518 
-plant 542 


Powder-horn 2 48 
Prairie-apple 363 
-acacia 331 
-anemone 102 
-bergamot 3 134 
-burdock 3 462 
Prairie-clover 
2 369-371 
Hairy 371 
Purple, or Violet 370 
Silk 370 
donk 462 
Prairie-dog Weed 
3 513, 195-6 


Prairie-grass 
I 195 —6, 199, 231 


-fire 3 214 
-grub 2 445 
-hyssop 3 142 
-indigo 2 346 
-lily I 5335 2 568 
-mesquite 2) 333 
-mimosa 2 332 
-money-wort 714 
-pine 3 367 
-potato 2 363 
-rocket 2 172-3 
-rose 283 
-sage 302 
-senna 336 
-smoke 2 102, 273 
-tea 2 455 
-turnip 363 
-weed 262 
-zinnia . 3 466 
Preacher-in-the-pulpit 

1442 sea 000 

Pretty Nancy 2 
Pricket 208 
Prickly-ash 444, 617 
Yellow 444 
Prickly-back 3 289 
Prickly-bur _ I 615 
Prickly-thistl® 3 553 
-elder 2 617 
See ae 2 571-2 
-pop 138 
Prick Reade 2 208-9 
-timber 2 492 

Prick-tree, Butcher’s 
2 492 
Prickwood 492 
Pride-of-Ohio 2717 
Pride-weed 3 442 


Vox. TIT: 


Priest’s Crown 315 
Prim 2 729 
PRIMROSE FAMILY 707 
Primrose 2 286, 708 
Bird’s-eye 2 708 
Dwarf Canadian 708 
Fremont’s 2 606 
Greenland 709 
Hartweg’s 607 
Lavendar-leaved 607 
Mealy 708 
Missouri. 605 
Mistassini 2 708 
Oblong-leaved 607 
Oklahoma 606 
Scapose 602 
Short-podded 604 
howy 603 
Spotted 2 605 
Three-lobed 604 
Tooth-leaved 608 
Tree 595 
Primrose-willow 2 589 
Primwort 729 
Prince’s-feather 22 
Prince’s-pine 2 672 
Print 729 
Prinopsis 3377. 
Privet 2 691, 729 
Procession aw 
2 449 
Pry f 1033 
Psoralea. 2 360 
Digitate 362 
Few-flowered 361 
Lance-leaved 361 
Large-bracted 363 
Large-stipuled 364 
Many-flowered 361 
Narrow-leaved 362 
Nebraska 362 
Sainfoin 365 
Silver-leaf 363 
Small-flowered 360 
Ptiloria 3 312 
Puccoon, Gmelin’s 3 89 
Hairy, Hoary 3 89 
Indian 89 
Red 2 140 
White 140 
Yellow 84; 3 89 
Puccoon-root 2 140 
Puck’s-foot 2 130 
Puff-ball 501 3/3) .315 
Puke-weed 3 308 
Pulsatilla Amer. 2 102 
Pumpkin, Wild 3 291 
Purging-root 2 469 
Purple-grass, or Wort 
2 358 
-rocket 2 590 
Purplewort 2 258 


PURSLANE FAMILY 2 35 


Purslane, Pussley 40 
Black 467 
French 41 
Garden 41 
Hairy 40 
Marsh 2 585 
Milk 467 
Mud 537 
Notched 40 
Sea 34 
Spotted 467 
Water 579 
Western 2 40 
White 469 

Pursley, Milk 469 
Black-spotted 467 

Purvain 3°95 

Pussies, Pussy cats 

2 355 


Vou. lll. ENGLISH 
Pussy-toes 3 451 
Putty-root I 574 
Pyracanth 2 322 
Pyramid-flower, or 
-plant Bets 
Pyxie 2 706 
Quafodil I 493 
Quaker-bonnets 
2 348; 3 251 
-ladies 3 251 
Quaker-lady 
2 245; 3 251 
Quarter-vine 37.237 


Queen Anne’s-lace 


2 625 
Queen-of-the-meadow 
245; 3 357 
Queen-of-the-prairie 
2 249 
Queen-weed 634 
Queen’s-delight 2 461 
-gillyflower 175 
Queen’s-root 2 461 
Quercitron I 619 
Quick T2832) 3119 
Quick-beam 2 287 


Quick-in-the-hand 
2 


Quickset 2 
Quicksilver-weed 2 


Quick-thorn 319 
Quill-fern I 18 
QuILLwort FAMILY 
50 
Quillwort 46-8 
Black-based 54 
Braun’s 51 
Butler’s 54 
Dodge’s 53 
Eaton’s Be) 
Engelmann’s 53 
Grave’s 54 
Lake 50 
Pitted 52 
Sugary 52 
Tuckerman’s 51 
Warty 51 
Quince-star 2 321 
Quinine, Wild 3 465 
Quinine-flower 36 
Quinine Tree 2 445 
Quinsy-berry 2 237-8 
Quitch 3 204 
Quitch-grass r283 
Quital 3 204 
Quiver-leaf I 590 
Quobsque-weed 1 400 


Rabbit-berry Zs77, 


-brush 3 370, 376 
-flower 177,204 
-meat 121 
_ -pea 2372 
-root 618 
-tobacco 454 
Rabbit’s-mouth 178 
Raccoon-berry 2 130 
Radical-weed 3.165 
Radish, Wild 2 195 
Garden 195 
Horse 163 
Ragged- -cup 3 460 
-Jack 2 69 
-Robin 69 
-sailor I 670 
RAGWEED FAMILY 3 338 
Ragweed 3 341-2 
Woolly 543 
Ragwort 540 | 
Cat’s-paw 542 
Golden 544 |. 


INDEX, INCLUDING POPULAR PLANT 


Ragwort, Prairie 3 543 
Tansy 542 
Rainbow-weed 2 581 
Raisin, Wild 2) ays) 
Raisin-tree 2 237 
Rampion, Large 2 595 
Ramps I 497 
Ram’s-claws 2 113 
-head re al 
-tongue 3 246 
Rancheria-grass 1 290 
Rancid Sey: 
Ranstead M77, 
Rantipole 2/625 
Rape 2 193-5 
Summer 2 193 
Rapper-dandies 2 693 
Raspberry 2 276-7 
Arctic, Cuthbert 277 
Black, Black-cap 277 
Canadian 276 
Carolina Zig) 
Flowering 276 
Purple 276 
Gladstone 276 
Gregg 277 
Ground 2 85 
Hansell : 277 
Hilborn 277 
Mountain 276 


Purple, Wild 277 


Rose-flowering 276 
Running 278 
Virginia 276 
White-flowering 276 
Wild Red 277 
Ratsbane I 570; 2 672 
Rat-stripper 2 492 
Rat-tail 3 246 
Rattan-vine 2 502 
Rattle 3 320-3 
Bull 2 64, 68 
Cow 68 
penny, Yellow 3 223 
Red 223 


Rattle-bags 264; 3 223 
Rattle-box 


2 34733 223, 588 
-bush 2 344-5 
-nut 2 77,91 
-root gI 

Rattler-tree I 587 


Rattle-snake-bite 2 221 


-fern TG 
Rattle- Seales herb 2 90 
-flag 2 622 
-leaf T1570 
-master I 534; 
2 622; 3 406, 367, 
365 

-plantain I 569-70 
-creeping-root 569 
-root ES Zoi 
2 91, 451; i 335-8 
-weed 5703 
2 fe 622 
Rattle-top 2 gI 
Heart-leaved oI 
Mountain 2 92 
Rattle-weed 2 QI 
Canadian 379 
Red-bay®™ 2 133 
-berry 2 618, 622 
-benjamin I 525 
-Betty 3 300 

Red-berry, Swamp 
2 704 
-Bird’s-eye 2 60, 426 
-bud 2 335 
-brush 661-2 
-camomile 121 
-cedar I 66-7 


Red-cedar 
Rocky Mt. I 66 
Shrubby 67 
-gum 2 235 
-Indian-paint 140 
-Indians 3 214 
-ink-plant 2 26 
Red: knees I 670 
-morocco 2 T2T 
-pollom 693 
-puccoon 661 
-rattle 3 221 
-robin 2 69, 426 
-rod 69, 270 
-root T5338 OSA 
2 2, 140, 270, 718 
Carolina I 531 
Indian 531 
Red-rot 2 203 
-Sally 258 
Red- Bhenke I 654 
2 420, 666, 668, ene 
Red-top I 203 
see Grass 
Red-weed I 653, 668; 
2926,,137 
-willow 2 661 
Redberry 2 90, 618 
-bamboo I 530 
-snakeroot 2 90 
Redfieldia © 235 
Reed 295 
see Grass 
Reed-bent 18 aie 
Reed-grass I 207-11 
Bog 210 
Common 232 
Giant 213 
Indian 232 
Labrador Aaa 
Lake 208 
Langsdorf’s I 209 


Long-leaved 212 
Macoun’s, Narrow 


210 
Northern, Nuttall’s 
211 
Pickering’s 208 
Porter’s 2 208 
Salt 223 
Sea I 212 
Short-haired 213 
Southern 179 
Wood, or Sweet 
202, 209 


Reed-Mace, Great 1 68 
Lesser 69 
Resurrection-plant I 49 
Rheumatism-root 1535 ; 
2120, 612)8)22 


-weed S425 
-wood 21 
Rhineberry 2 502 
Rhododendron 681 
Rhubarb 
Bog, or Poison 3 532 
Monk’s 656 
Poor Man’s 2 120 
Rhodora 680 
Rhynchosia 2 421-2 
Rib-grass, -wort 
3 246-8 
Rice, Indian 1 168 
Canada 168 
False, White, 168 
Jungle I 134 
Water, Wild 168 
Rice’s-cousin I 169 
Rice-cut-grass 169 
Rich-leaf 3 153 
Rich-weed 1 637; 291; 


3 153, 341, 361 


NAMES. 627 
Rignum glrss: 
Rim-ash I 629 
Ripple-grass 3 246 
River-bush 255 
RIVER-WEED FAMILY 
2 205 
River-weed 205 
Road-weed 000 
Roanoke-bell 3 83 
Robert’s-plantain 439 
Robin, Robins 2 69 
Robin-run-away 
2825) 2011s. 
Robin’s-plantain 
3 332, 439 
Rock-bells 2 92 
Rock-berry 693 
-brake’ I. 32, 36 
Rock-cress 2 176 
Arctic 179 
Low, Northern 179 
Lyre- leaved 179 
Virginia 179 
Rock-lily 2 92 
-plant 208 
ROcCK-ROSE FAMILY 
2 539 
Rock-rose 540 
Rock-weed 
2 4265 3 267 
Rocket 
Bastard 2 192 
Crambling 200 
Dame’s 175 
Dyer’s 200 
False 178 
Garden 192 
Italian 200 
Prairie 172-3 
Small-flowered 172 
Purple 2 178,590 
Sand 194 
Sea, American 196 
Scrambling 174 
Sweet 175 
Wall 194 
Winter, or Wound 
177 
Yellow 


2 200, 177, 711 
Rocket-cress 2 179, 183 
Rocky Mountain- 

grape 2 128 


Rogation-flower 449 
Rogue’s-gilliflower 175 
Roman-plant 2 14 
-willow 72 
-wormwood 
2 144; 3 34! 
Romeria 2 20 
Roosters 547 


Rooster-heads 2717 


Rope-bark 575 
Rope-wind 3 45 
Rosa-solis 
1 487; 2 203 
Rose FAMILY 2 242 
Rose 283 
African 137 
Arkansas 284 
Bramble 284 
Brere, Brier 284 
Burnet 716 
Canker 137; 284 
Christmas 87 
Cinnamon 286 
Climbing 283 
Common 
-wild 285 
Corn 2 61, 137 
Cotton 3 447 


628 


Rose, Dog 2 284 
Dwari-wild 285 
Early-wild 283 
Egyptian, Gipsy 290 

_Evergreen 286 
Hip 284-6 
Kitchen 286 
Low 285 
Mallow 2 524 
Maple-leaf, Gacider 

3 270 

Marsh-holy 2 689 
Meadow 283 
Mexican 2 41 
Michigan 283 
Noble 2076 
Northeastern 2 283 
Pale 283 
Pasture 2 85 
Prairie, Prickly 283 
Rock = 2. 530, 539-40 
Rosin 533 
Scotch 284 
Sensitive 2 333 
Shining 285 
Smooth 283 
Snowdon 207 
Sun 539-40 
Swamp, Sweetbrier 

285 

Wild 283-5 
Wind 138 
Woods’ 284 

Rose-acacia 2 375 

Rose-bay 590, 681 
Ground 589 
Lapland 680 
Mountain 681 
Wild, or Dwarf 681 

Rose-blush 283 

Rose-campion 2 70 

Rose-mallow 524-5 

Rosemary 
Marsh 2 689, 718 
Maryland 3 426 
White 430 
Wild 2 689 

Rose-moss 41 

Rose-noble 3°95 


Rose-of-Plymouth 6 


Rose-of-Sharon 536 
Rose-petty 3 439 
Rose-pink see 

-pogonia I 560 
Rose-willow 2 661 


Rosewort, Roseroot 207 
Rosin-plant 


3 462; z 207 

ose 533 
ace: White a 57 
-weed 3 460-1, 374 
-wood 375 
Rosita 3 80 
Rosy-bush 2 245 
Rotala 579 


Rot-grass 1214; 3 226 


Roubieva 215 
Rough-root 3 367 
-weed 125 | 
Round-dock 2 514 
Round-heart 639 
-root 304 
-rush I 467 
-wood, -tree 2 287 
Roving-sailor 3 175 
Rowan-berry 2 287 
Rowan-tree 2 287 
American 287 


og 3 270 
Royal-bracken I7 
ROYAL-FERN FAMILY 

17 


Royal-fern 1 7 | Rush, Slender 1 470 
Royal Osmond 7 Small- headed 480 
Rue Anemone = 2 102 Spart 477 
False 2 89 Spike 311-19 
Meadow 118-20 Staff 467 
RueE FAMILY 2 443 Stout 482 
Rue ; 2 445 Sweet 446 
Ruellia 3 241-2 Thread 467 
Long-tubed 241 Three-flowered 476 
Short-tubed 241 Three-square 330 
Runch 2 192 Toad 469 
Running-pine £47 Torrey’s 478 
-moss I 47 Tule 331 
Ruppia r 88 Twig 348 
RusH FAMILY I 465 Two-flowered 475 
Rush 467-485 Vasey’s 471 
Aglet-head 314 Water 467 
Awl-leaved 473 Water-bog 348 
-petaled 474 Weak 481 
Bald I 347 White 223 
Baltic 468 Wood I 483-5 
Bass 331 Yard 470 
Bayonet 477 Yellow-flowering 451 
Beaked I 342-7 (see Bulrush) 330-7 
Black 331 (see Clubrush) 
Black-grass 470 327-34 
Bog I 348, 467 | Rush-garlic 497 
Bolles Boulder 331 -nut 304 |- 
Bottle 294 | Russia-grass 266 
Brown-fruited 476 | Russian thistle 2 25 
Bulbous 477 | Rutland Beauty 3 46 
Bulrush I 326-327 | Rye, Wild, 
Canada 480 I 281, 288-94 
Candle 467 | Rye-grass, see Grass 
_Chair-maker’s 330 
Chestnut 475 | Sabbatia 3 3-7 
Club 327-9 Branching 2-4 
Clustered Alpine 475 Coast 5 
Common 467 Elliott’s 6 
Cotton 323-6 Lance-leaved 4 
Cotton-grass 323-6 Narrow-leaved 5 
Creeping 474, 476 Prairie 6 
Diffuse 482 Square-stemmed 5 
Dudley’s 470 | Sabino-tree I 64 
Dutch 41 | SACRED-BEAN FAMILY 
False Bog I 344 2 76 
Forked 472 | Sacred-bean 2 76 
Glomerate 462 | Saddle-tree 2 83 
Grass-leaved 473 -leaf 83 
Large 473 | Safflower 3 560 
Greene’s 472 | Sage 
Hare’s-tail 324 Indian 3 361 
Hard 467 Jerusalem 118 
Highland 469 Lance-leaved 130 
Horned I 342 Lyre-leaved 129 
Inland 471 Meadow 129 
Jointed 477 Mountain 530 
Knotted 478 Nettle-leaved 130 
Knotty-leaved 481 Pitcher’s 129 
Long-styled 474 Tall 129 
Maine 472 Western 3 529 
Many-headed 479 White 2 20 
Mat 331 Wild 3129, 130, 525 
Moor 475 Wood 3 101 
Narrow-panicled 481 Wormwood Bu525 
New Jersey 479 | Sage-brush 2 19; 
Nut 349-51 3 525-30 
Pennsylvania 468 Pasture 3 525 
Pin 467 | Sage-bush 530 
Pith 467 | Sage-of-Bethlehem 149 
Pole, or Pool 331 | Sage-leaf 3 118 
Richardson’s 1 478 | Sage-willow 
Roemer’s 468 I 598; 2 581 
Round 467 -wood 3 530 
Scirpus-like 480 | Sagittaria I 101-4 
Scouring 41 | Sailor, Climbing 3 175 
Sea 469 | Sailor’s-knot 2 426 
Sea-club 335 -tobacco 3 527 
Secund 471 | Sainfoin 2) 3553 400 
Sedge 446 | St. Andrew’s Cross 528 
Sharp-fruited 481 | St. Anthony’s Turnip 
Short-fruited 479 2 112 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Vo. III. 


St. Bennett’s herb 2 
St. George’s herb 3 
St. James’-weed 
-wort 
St. JoHn’s Wort 
FAMILY 2 
St. John’swort 2 


653 
286 
158 
3 542 


527 
530 


Bedstraw 531 
Bushy 530 
Canadian 536 
Clasping-leaved 535 
Common 533 
Copper-colored 533 
Corymbed 533 
Creeping 531 
Dense-flowered 530 
Drummond's 536 
Dwarf 535 
Elliptic-leaved 532 
False 536 


Giant, Great 2 
Kalm’s 
Large-spotted 


Larger Canadian 535 
Larger Marsh 537 
Marsh 537 
Mountain 534 
Northern 534 
Pale 532 
Round-podded 532 
Shrubby 2 530 
Slender 535 
Spotted 533-4 
Small-flowered 535 
Straggling 532 
Virgate 533 
St. Joseph’s Wand 
3 186 


St. Peter’s Wort 2 = 
Salad Tree 
Salfern Stone-seed 3 3° 


Sally-bloom 2 590 
Sally, Creeping 712 
Wandering 712 
Salmon-berry Py A) 
Saloop 2 134 
Salsify, Meadow 3 313 
Salt-grass I 250 
Rush 223 
Salt-grape 2 25 
Salt-meadow Grass 
ryan 
Salt-of-Lemons 2 431 


Salt-rheum weed 3 181 


Saltweed 1469; 218 
Saltwort 2 21,25 

Black 2 716 
SALVINIA FAMILY I 37 
Salvinia 8 


a} 

Samphire, Marsh 2 21 
Jamaica 477 
Sampson’s Snake-root 
2 3645 Simzyas 

2 


Sanctuary 3 
Sand-bur 5 tan oy) 
3 166, 342 
-cherry 2. 205 
-grass T2128 
2 80, 250, ees 

Sand-myrtle 2 

-nettle 2 ye: 
-parsley 2 652 
-reed I 212 
-spine 2 461 
-spur 167 
-spurry 2 59, 60 
Northern fe) 
Salt-marsh 59 
-star Lsedge] 363 
-vine 3°36 

SANDALWOOD FAMILY 
I 639 


Vo. ITI. 
Sandweed 2 59 
Sandwort 2 52-60 
Arctic 54 
Beach 56 
Bed 59-60 
Blunt-leaved 257 
Fendler’s 54 
Fringed 53) 
Hooker’s 53 
Large-leaved 58 
Mountain 757 
Pine-barren 55 
Pitcher’s 56 
Purple 60 
Rock BG 
Sea-beach 58 
Seaside 59 
Showy 57 
Siberian 55 
Slender 53 
Texan 56 
Thyme-leaved 52 
Vernal 54 
Sang 618 
Sanghara-nut 2 612 
Sangree-root, or 
Sangrel I 645 
Sanguinary 3515 
Sanicle 2 624 
American 2 226 
Black 623 
False 2 229 
Great 2 264 
Indian, White, 3 361 
Yorkshire 3 226 
Santa Maria 3 465 
SAPODILLA FAMILY 
2 719 
Saracen’s corn, or 
wheat I 672 
-compass 3 542 
Sarah T5260 


Sardian Nut 615 
Sarsaparilla 1 528-30; 


2 506-7 
Big I 646 
Bristly I 529, 618 
False 618 
Rough 2 618 
Texas, or Yellow 131 
Virginia 2 618 


Sassafras, Swamp 2 82 


Sassafras-tree 134 
Satin 2 191 
Satin-flower 2 43, 190-1 
-grass I 184-6 
Satin-pod 2 190 
-walnut 2235 
Sauce-alone 2 170 
Savin I 66 
Horse 66 
Red 66 
Savory, Summer 3 137 
Saw-grass I 348 
Saw-wort 3 367 

SAXIFRAGE FAMILY 
2 214 


Saxifrage 2 216-23, 230 


Aconite 223 
Alpine-brook 222 
Bulbous 222 
Burnet 655 
Clustered Alpine 8 
Drooping 222 
Early 218 
Foliose 220 
Golden 230 
Gray’s 219 
Iowa 230 
Kidney-leaved 220 
Lettuce 219 
Livelong 223 


Saxifrage 
Michaux’s 2 221 
Mountain 216 
Nodding 222 
Pennsylvania 219 
Purple 216 
Sengreen 217 
Spring 218 
Starry 221 
Swamp 219 
Texan 218 
Three-toothed 217 
Tufted 222 
Yellow Marsh 216 


Yellow Mountain 217 


Scabby-head 2 626 
Scabious 3 290 
Sweet 439-40 
Scabish 2 595, 602; 
3 419 

Scabwort 3 457 


Scaldweed 2 Bu 
Scammony, Wild 3 43 


German 3 46 
Scarb-tree 2 290 
Scarlet-berry 3 167 

-lightning 2 69 

-lychnis 69 
Schedonnardus 1 226 
Scheuchzeria 93 
Schizaea 9 
Schizonotus 2 247 
Sclerolepis 2 AEE 
Scolochloa I 262 
Scoke 2 26 
Scorpion-grass 3 85-7 

Early 87 

Field 86 

Marsh, Mouse-ear, 85 

Spring 3 86-7 

Yellow and Blue 86 
Scotch-cap 2277, 


Scotch-mercury 3 204 
Scouring Rush 1 41-2 
Scrambling Rocket 
2 174 
Scratch-grass 
I 675; 3 259 


-weed 3 259 
Screw-auger I 565 
Screw-stem 3 16 
Scrofula-plant 

2 540; 3 179 

-root I 508 

-weed 570 
Scrub-oak, see Oak 

-bush 3 430 
Scurfy-pea 2 360, 361 
Scurvish 2 505 
Scurvy-senvie 2 193 


Scurvy-grass 
2 163,177; 3373 


Danis 2 163 
-weed 163 
Sea-ash 2 444 
-beach ry aie 
Southern 141 
Smaller 141 
-bent 363 
Sea-blite 2 23 
Annual 2 
Low 24 
Western 2 
Sea-bugloss 3 82 
-burdock 345-6 
Sea-chickweed 2 58 
-gilliflower 718 
-goosefoot 2 23 
Sea-grape 22 
English a2 
Sea-grass Tr ss)o1- 
2 21,718 


\ Sea-grass 

English 2 21 
-hay I QI 
ahallyhoek 2 524 
-kemps 247 
-lavender 2 718 
-lovage 648 
-lungwort 3 82 
-Mat-weed Tare 
-milkwort 2 716 
-Ox-eye 3 477 
-parsley 2 648 
-pea 413 
-pimpernel 58 
-pink 264,718; 36 
-purslane 2 34,58 
-reed I 212 
-rocket 2 196 
-Sand-grass Lee t2 
-Sand-reed 1 Be 
-sedge 446 
-thrift Py As, 7 fils) 
-trifoly 2 716 
-wrack I gI 
Sealwort reser 
Seaver-wood 2575 
SEDGE FAMILY I 205 
Sedge I 352-441 
Alpine - 417 


Loose-flowered 416 
Arctic Hare’s-foot 


373 
Assiniboia AIL 
Aroostook Baa 
Awl-fruited 371 
Awned 428 
Back’s 388 
Bailey’s 436 
Barratt’s 417 
Beak I 341-6 
Beaked 434 
Bear 373 
Bebb’s 382 
Bent 403 
Bicknell’s 383 
Bigelow’s 421 
Black 418 
Black-edged 393 
Bladder 439 
Blunt Broom 380 
Bog 417 
Boott’s 395 
Bristle-leaved 396 
Bristle-stalked 388 
Bristly 438 
Bristly-spiked 370 
Broad-leaved 400 
Broad-winged 384 
Brome-like 376 
Broom 115, 182 
Brown 419 
Browned 386 
Brownish 375 
Bur-reed 368 
Bush’s 414 
Button 435 
Capitate 361 
Carey’s 400 
Carnation-grass 
397, 415, 428 
Carolina 414 
Cat-tail 439 
Cherokee 412 
Chestnut 411 


Clustered 
I 362, 373-5 


Field 362 
Coast 376 
Collins’ 430 
Common 22 

Great 25 

Lesser 420 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


629 


Sedge, Crawe’s 1 304 
Crawford's 380 
Creek 224 
Creeping t) 362 
Crested 381 
Curved 361 
Cuspidate 424 
Cyperus-like 437 
Cypress-swamp 419 
Dark Green 409 

Brown 412 
Davis’ 408 
Deep-green 394 
Dense Long-beak 379 
Dewey’s 376 
Douglas’ 363 
Drooping 407 
Drooping Wood 410 
Dry-spiked 364 
Eastern 378 
Emmons’ 392 
FalseSummer 409 
False Uncinia 389 
Falkland Island 383 
Farwell’s 393 
Fernald’s Hay 387 
Fescue 383 
Few-flowered 3890 
Few-fruited 304 
Few-seeded 436 
Fibrous-rooted I 391 
Field 405 
Fox 369 
Fox-tail 368 
Fragile 434 
Frank’s 438 
Fraser’s 441 
Fringed 25 
Glaucescent 406 
Glomerate 368 
Golden-fruited I 397 
Goodenough’s 22 
Graceful 407 
Grass-like 397 
Gray 406 
Gray’s 439 
Greater Prickly 365 
Green 429 
Greenish-white 385 
Hairy-fruited 428 
Hairy 428 
Hair-like 411 
Hammer 428 
Handsome 408 
Hare’s-foot 385 
Hart Wright’s 426 
Hasse’s 396 
Hay 387 
Hayden's 421 
Heath- 415 
Heavy 367 
Hidden-fruited 42 
Hillside 364 
Hirsute 414 
Hitchcock’s 405 
Hoary 375 
Hop 440 
Hop-like 440 
Hoppner’s 423 
Houghton’s 427 
Howe's 377 
Hudson Bay 374 
Inflated 433 
Inland 377 
Involute-leaved 362 
James’ 388 
Lake-bank 425 
Large 441 
Large-panicled 371 
Larger Straw 382 
Leavenworth’'s 367 
Lenticular 422 


630 


Sedge 
LesserCommonti1 420 
Lesser Panicled 370 


Lesser Prickly 365 
Little Prickly 378 
Livid 397 
Long 431 
Long-beaked 412 
Long-bracted 429 
Long-stalked 395 
Loose-flowered 402 
Loose-flowered- 
alpine 416 
Low Northern 395 
Louisiana 440 
Magellan 417 
Maine 432 
Marsh 420 


Marsh Straw 1 384 
386, 404 

Mead’s 397 
’ Moosehead Lake 432 


Midland 366 
Mt. Katahdin 405 
Mud 416 
Muhlenberg’s 365 
Muskingum 381 
Myrtle 446 
Nard I 360 
Narrow-leaved 406 
Nebraska 423 
Necklace 381, 433 
New England 392 
Nodding 425 
Northeastern 431 
Northern 393 
Bog 361 

Novthern Clustered 
373, 375 

Northern Meadow 

386 
Norway 374 
Orono 379 
Oval-headed 328 
Pale 415 
Parry’s 418 
Pennsylvania 391 
Pink-leaf 397 
Plantain-leaved 399 
Pointed Broom 380 
Porcupine 437 
Prairie 370 
Prairie Straw 384 
Prickly I 365 
Greater 365 
Prickly-bog 378 
Pubescent 3904 
Purplish-tinged 398 
Rae's 433 
Raven’s-foot 371 
Reflexed 364 
Retrorse 435 
Ribbed 413 
Richardson’s 396 
Rock 390 
Rough 416 
Bou: fruited 434 
Russet 432 
Sallow 436 
Salt-marsh 423 
Sand 363 
Sartwell’s 363 
Schweinitz’s 437 
Scirpus-like 390 
ea I 363 
Seabeach 385 
Seaside 424 
Sharp-scaled 408 
Sheathed 399 
Short-leaved 413 
Short’s 419 
Silvery 375 


'UHIiUwttwOwt= iii... ene a ed 


Sedge, Slender 1 427 
Slender-stalked 410 


Slender Wood gor 
Sniddle (Sedge) 420 
Soft Fox 369 
Soft-leaved 372 
Southern 366 
Glaucous 420 
Sparse-flowered 372 
Spreading 401 
Squarrose 438 
Star, or Stare 422 
Shriver’s 403 
Stellate 365 
Straw 382 
Striate 403 
Summer 409 
Swamp 420 
Swan's 413 
Sweet 446; £1119 
Tawny 429 
Texas 364 
Thicket 400 
Thin-fruited 407 
Thin-leaved I 367 
Thread-leaved 390 
Three-fruited 372 
Torsets, or Turrets 
422 
Torrey’s 415 
Tuckerman’s 435 
Tufted 422 
Tussock 420 
Twisted 421 
Two-edged 402 
Umbel-like 394 
Variable 399 | 
Variegated 418 
Velvet 426 
Vernal 391 
Walter’s 426 
Water 422 
Weak Arctic 389 
Weak Clustered 374 
Stellate 379 
White-bear 401 
White-edged 410 
White-scaled 386 
-tinged 392 
Whitish 375 
Willdenow’s 1 387 
Winter-grass 
(Sedge) 409 
Woodland 402 
Wood’ Ss 398 
Woolly 427 
Yellow 430 
Small 430 
Yellow-fruited 369 
Yellowish 431 
Sedge-cane,-rush 446 
See-bright ares 
Seed-box 2 588 
SELAGINELLA FAMILY 
1 48 
Selaginella 
Creeping, Low 1 49 
Rock 49 
Selenia 2 190 
Self-heal Ph ii 
Seneca Snakeroot2 451 
Senecio, Arctic 3 542 
Broom-like 541 
Riddell’s 541 
Sea-beach 541 
Senega-root 2 451 
Sengreen 2, 212 
-saxifrage 2 216 
Senhalanac 2 482 
SENNA FAMILY 2 334 
Senna, American 336 
Coffee 336 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Senna, Low 2 335 
Prairie 337 
Sickle 235 


Wild, Medsger’s 336 


Sensitive-brier 333-4 
-fern I II 
-Joint-vetch 2 393 
-pea 337 
-plant 2 333-4, 

337, 393 
Bastard 2 393 
-rose 333 

Senvie 2 191-2 

Serpent-grass I 671 

Serinia 3 306 

Serpentary I 645 

Serpent’s-tongue cee 


Service-berry 2 292-3 
Service-tree 2 287, 292 


American 287 
Sesame-grass_—-—‘I:-=‘Iil 
Sesban 2 376 
Setwell 3 286 
Seven-barks 2 28% 

-sisters 2 473-5 
Shad-bush 2 292 

-flower 2 148, 692 
Shag-bark I 581-2 

Big 582 

Little 583 

Southern 581 
Shakers 250 
Shame-face 2 426 

-vine 2 333 
Shamrock T5255 

2 351,358, 431 

True (?) 2 354 

Water 3 18 
Shave-grass Weed I 41 
Shave-weed 41 
Shavings 728 
Shawnee-wood 3 23 

-haw 20273 
Sheath-flower 3 303 
Sheep-berry Beers 

-bine 347 

-bur 345 
Sheep-bur, Small 3 77 
Sheep-foot 2 359 
Sheep-laurel 683 

-lice 3 76 
Sheep-noses 2 260 
Sheep-poison 2 683 
Sheep-root, -rot 3 226 
Sheep-sorrel THOSS)s 

2 683-4 

Poison 2 432 
Sheep’s-bib 3 299 
Sheep’s-gowan 2 358 
Sheepweed 2 73,521; 

226 
Shell-bark I 581-2 

Big, Thick 582 

Western 582 
Sheli-flower 3 181 
Shepherd’s Clock 3 716 

-club BE173 

-delight 2 716 

-needle 2 627 

-purse, -bag, -pouch 

158 

-stafi 3 289 


-Weather-glass 2 716 


Sherard, Herb 3 266 
Sheriff-pink 518 
Shield-fern I 18-22 

-root 21 


Shining-grass 2 120,440 
Shin-leaf 2 669-71 
Shin-wood I 67, 
Shittim-wood 2 722,720 
Shoe-make 2 482 


Vor. III. 


Shoes-and-Stockings, 


Lady’s 2 359 
Shoe-strings ‘2 366 
Devil’s 2 372 
Shoo Fly 2 345 
Shooting Star 2 717 


Shore-grass, -weed 


3 250 
Short-husk, Bearded 


I 189 

Short-shucks I 58 

~Shotbush 2 618 
Shovel-weed 158 


Shrub Yellow-root 89 
Shrubby Althaea 2 526 


-fern 586 
-trefoil 2 445 
Shunis 2 648 
Sibbaldia 2 261 


Siberian Oilseed 2 157 


Sickle-grass 1 425, 673 
Sickle-pod 2 182 
Sickle-senna 335 
-weed I 670 
-wort 3 101, 115 
Sida 2 520-1 
Side-saddle Flower 202 
Sieglingia I 234-8 
Silk, Virginia 3 30 
Silk-grass 1177,513; 
Bal: 

Silk-plant 246 
-tree 2 331 
Silk-weed 3 30 
Rose, or Swamp 3 26 
-seed 3 406 
Silky-grass T 74 
Silphium 3 462 
Silver-beard 117 
Silver-bell tree 2 722 
Silver-berry 576 
— 453 
-chain 2) 375 
-chickweed 27 
-feather 2 258 
-fir, American 1 63 
-grass 373 
Silverhead a 27 
Silver-leaf 2 81,577, 


: 245,440; 3 453 
Silver-pin, Joan 2 136 


Silver-rod 2 245, 
440-1; 3 383 

-weed 2 121, 440-1; 

3 383 

Simon’s- -weed 3 179 
Simpler’ s-j oy 95 
Simpson’s-grass I 137 
Simson, Simpson 539 
Single-delight 2 671 
Sinkfield 251 


Siris, pink 331 
Sisymbrium, Tall 2 
Sitfast 
Skedge, Skedgwith 
Skedlock 
Skegs 

Water, Yellow 1 
Skeleton-weed 3 314 
Skevish 
Skewer-wood 2 492 


Skiver-wood 492 
Skrew-stem “* 3 16 
Skull-cap 105 
Blue 105 
Britton’s 108 
Bush’s 109 
Downy be! eS 
European 
Hairy, Heartieaee 
107 
Hyssop 107 


_ 


Vor. IIL. 

Skull-cap 
Large-flowered 3 107 
Larger 107 
Little 108 
Mad-dog 106 
Marsh 109 
Resinous 108 
Rock 109 
Showy 106 
Side-flowering 106 
Small 108 
Veined 110 

Skunk-bush 2 483 
-cabbage 

I 445; 2 202 

Skunk-currant 238 
-weed 445,455 

Sleeleleaf 2 682 

Sleeping-beauty 431 

Sleepy Dick I 509 

Stink-weed 

2 580; 3 300 

Slippers 2 441 

Slipper-root I 550 
-weed 2 440 

Slippery-elm I 627 

Sloe 3 273 
Alleghany 324 

Slovenwood 3 526 

Slunkweed 357 

Smallage 2 660 

Small-pox plant 202 

Smartweed I 670 
Dotted, or Water 670 

Smelling-stick 2 134 

Smick-smock 184 

Smiddy-leaves 14 

SmILAx FAMILY : 526 

Smilax 27-8 


Smoke-tree, Wild 2 485 


Smoking-bean 3 238 
Snaffles 221 
Snag-tree 2 665 
Snake-berry 2 90; 
3 167, 255 

-bite 1 526; 2 140 
-flower 2 44, 68; 

3 94, 122 

-gentian 3.335 
-grass 2 44; 3 85 
-head 3 181 
-leaf I 506 
-lily 537 
-milk 2 469 
-moss I 47 
Snake-mouth I 560 
Whorled 561 
Snake-pipes I 39, 40 
Snake-root I 506; 
2 90, 623-4; 3544 
Black T1OAss 


2 91, 623, 684 
Button 2 622, ee 8 


Canada I 642 
Clustered 2 624 
Coltsfoot I 642 
Corn 622 
De Witt’s- a) 336 
Evergreen 452 
Heart I 642 
Heart-leaved 2 gI 
Large-fruited 624 
Poison 2 653 
Redberry feke) 
Sampson Ped aie 
Samson’s- 2 364 
Seneca 451 
Short-styled 624 
Southern I 642 
Thermon 2 63 
Vermont I 642 
Virginia I 645 
6 


Snake-root 


White 3 361-2; 

I 642 

White-berry 2 90 
Wild Gy atl 
Snake-head 181 
Snake-grass 85 


Snake’s-tongue T2 
Snake-weed 1 644, 670; 


2 658 

Black I 644 
Poison 2 653 
Saugrel I 645 
Snapberry 39277 
Snap-dragon 293,440; 
3 176-9 

Corn 3 179 
Snap-Jack 2 44 
Snappers 2 44, 64 
Snap-weed 135, 440-1 
Snap-wood Ea 


Sneezeweed 3 510-11 


Sneezewort 515 
-tansy,-yarrow 515 
Sniddle 20 
Snow-ball, Little 
3 270, 255 
Wild 2 504 
Snowberry 3 276 
Creeping 2 704 
Snow-blossom 2 728 
-drift I 280 
Snowdon-rose 2 207 
Snow-drift 153 
Snowdrop 722 
Yellow I 506 
Snowdrop-berry 3 276 
Snowdrop Tree 2 722 
Snowdrops 2 100 
Snowflake 275 
Snowflower tree 728 
Snow-on-the-mountain 
2 469 
Snowy-campion 63 


-Hydrangea 2 


Soap, Poor-man’s- 245 
Indian 500 
SOAPBERRY FAMILY 
2 500 
Soap-berry 500 
-plant, Indian 500 
-root 273 
Soap-weed L512 
Soapwort 3 12; 2 73 
-gentian 73 
Soapwort, White 69 
Soldier-buttons 85 
Soldiers 2 60, 284, 
581; 3 78 
Soldier’s-cap 2 142 
-plume I 559 
-woundwort 3 515 
Solentine 2 440 


Solomon’s-seal 
I 515-17, 521, 556 


Dwart 1 52 
False 516 
Great, or Giant 521 
Hairy 521 
Small, or Zigzag 515 
Smooth 521 
Star-flowered 516 
Three-leaved 516 
Two-leaved 517 
Soopoo-lalia 2 576 
Sophora, Silky 342 
Sorrel I 653-4 
Cock, English 654 
Cow 653 
Engelmann’s €54 


Field, Gentleman's 


ENGLISH INDEX, INCLUDING POPULAR PLANT 


Sorrel, Green, or 
Meadow 
_ Horse 
Ladies’ 
Meadow 
Mountain 
Red 53 
Sheep 1 653; 2 431-2 
Toad’s 653; 2432 
Wood 653-4; asst 
Sorrell-tree 2 691 
Sour-,orSow-berry 70 
Sour-bush 3 a 


I 654 

653, 656 

2 432 

I 654° 

653, 659 
6 


Sour-dock 1 653-4,659 
-grass 653-4 
Sour-grass, Lady. 2 432 
is ae 2 665-6 
-leek 1653 
-trefoil,-trifoly 2 431 
-wood 691 
Southernwood 3 526 
Sow-bane 2073 
-berry 704 
-foot 3531 
-grass 2 167 
Sow-thistle 3 316-17 
-tit 2 260, 280 
Spangles 504 
Spanish-bayonet I 512 
-dagger 512 
Spanish-lettuce 2 39 
Spanish needles 3 498 
Sparkleberry 2 698 
Sparrow-grass I 514 
-tongue I 661 
Spart 477 
Spatter-dock 2 78 
Spattling-poppy 2 64 
Spear-grass 
Te25es20n, 207 
Alpine 254 
Arctic 258 
Buckley’s 260 
Bunch I 260 
Chapman's 253 
Flexuous 257 
Glaucous 254 
Large-flowered 259 
Lew 253 
Meadow 267-8 
_ Mountain 254 
_~ Northern 256 
Prairie 260 
Sea 267 
Short-leaved 258 
Smooth 260 
Southern 240 
Sylvan 257 
Tufted 261 
Weak 255 
Wolt’s 258 
Spearmint 3 149,153 
Spearwort 2 106-7 
Speckled-jewels 2 440 
Speedwell 
Alpine 3 201 
Buxbaum’s- 203 
Byzantine 203 
Common 201 
Corn 202 
Field 203 
Garden 203 
Germander 3 201 
Ivy-leaved 203 
Marsh 200 
Procumbent 203 
Purslane 202 
Skull-cap 200 
Tall 3 204 


Thyme-leaved 3 202 
Upland 201 
Wall 202 | 


NAMES. 631 
Speedwell 
Water 3 200 
Wormskold’s 201 
Speerhawk 3 328 
Sperage - 514 
Spermolepis 2 651-2 
Spice 
Pond 2 135 
Spice-berry 693 
-bush 
2 132, 135-6, 617 
-hardhack 2 245 
-wood 2 135 
Spider-flower 2 197 
“lily ce 
SPIDERWORT FAMILY 
I 457 
Spiderwort 459-62 
Spignet 2 617 
Spike-grass 
I 92, 236, 248-9 
Spikenard 1515; 2617 
American 2617 
False I515 
Ploughman’s 
3 83, 445 
Small 2 618 
Wild I 515 
Spikenard-tree 2 617 
Spike-rush I 311-19 
Aglet-headed 314 
Angled 311 
Beaked 319 
Black-fruited 217, 
Blunt 313 
Bright Green chee 
Capitate 313 
Creeping 314 
Engelmann’s 314 
Flat-stemmed 318 
Knotted 311 
Large-tubercled 316 
Least 315 
Matted I 318 
Needle 315 
Ovoid 313 
Pale 312, 314 
Purple 312 
Robbins 311 
Slender 318 
Small’s 315 
Three-ribbed 317 
Torrey’s 316 
Twisted 316 
White 317 
Wolf's 315 
Spilanthes 3 469 
Spinach; French 2 13 
Strawberry 2 15 
Wild 2 10,14 
Spindle-tree 2 491-2 
Spingel 2 643 
Spink 2 74, 184 
Spiny-bur 459 
Spiraea 2 246-9 
Spiral-pine 1145 
Spires I 170, 212, 232 
Spirit-weed I 531 
Spleen-amaranth 2 22 
Spleenwort I 26 
See Fern 
Spleenwort-bush 1 586 
Spool-wood 609, 684 
Spoonflower, Arrow- 
leaved 444 
Spoonhunt 2 684 
Spoonhutch 2 681 
Spoonwort 163 
Spoon-wood 2 512 
Spoonwood-ivy 683 
Spotted-alder 2 235 
-eyebright 467 


632 


Spring Beauty 
2 37-8, 101 


White-leaved 38 
Spring-cowslip 2 85 
-lily I 506 
Springwort 2 471 
Spruce I 60-1 
Black 61 
Blue 61 
Cat 61 
Double 61, 63 
Gum-tree 61 
Hemlock 62 
Red 2a (6irn 
Single 61, 63 
Skunk 61 
Swamp I 61 
White I 61 
Spruce-gum tree I 61 
Spruce-pine I 61 


SPURGE FAMILY 2 452 


Spurge 2 463-74 
Alleghany Mt. 2 480 
Angled 468 
Anne Arundel 2 470 
Arkansas 2 473 
Black 2 467 
Blooming 469 
Blotched 467 
Blunt-leaved 472 
Broad-leaved 472 
Caper 2 eA 
Cypress 474 
Darlington’s 474 
Fendler’s 465 
Flowering 469 
Garden 474 
Hairy 2 467 

Spreading 2 467 
Hoary 465 
Geyer’s 463 
Ipecac 2 470 
Knotweed 463 
Large Spotted 468 
Leafy 473 
Maryland 2 470 
Myrtle 2EAGE 
Narrow-seeded 466 
Nicaean 474 
Petty 475 
Prairie 2 464 
Quack salver’s 474 
Reticulate-seeded 

472 
Ridge-seeded 466 
Round-leaved 464 

Spreading 464 
Rocky Mountain 475 
Seaside 463 
Shore 2 463 
Spotted 2 467 
Sun. 2 473 
Thyme-leaved 465 
Tinted 475 
Toothed 476 
Upright Spotted 468 
Variegated 469 
Various-leaved 477 
Wart 473 
Warted 2 472 
Warty 2 476 
White- Hemeaele 463 
White-margined 469 

-stemmed 466 

Spurge-flax 574 
-ipecac 470 
-laurel 2nErA 

Spurge Nettle 2 462 

Spurge Olive 574 

Spurry 2152550 
Corn 59 
Knotted 52 


Spurry, Sand 259 
Spurt-grass tesgs 
Spurtwort 3 266 
Squash-berry 270 
Squatmore 2 141 
Squaw-berry 697 
-bush 2 661-2; 3 270 
-drops 36233 
-flower meses 
-huckleberry 2 697 
-mint 3 136 
-plum 255 
-root T2681: 32385 
-V ine 3 255 
Squaw-weed 3 544-6 
Crawford’s 545 
Northern 544 
Robbins’ 544 
Round-leaf 545 
Small’s 546 
Swamp 544 
Western 546 
Squirrel-corn DeTAZ 
-cup IO 
-ear I 569 
-pea, Ground 2 129 
-shoes I 550 
-tail 287 
STAFF-TREE FAMILY 
2 490 
Staff-tree 493 
-vine 493 
Stag-bush 3 273 
Stag’s-head I 504 
Stagger-bush 2 691 
-grass I 532 
-weed 2 143 
-wort 3 542 
Staghorn 2 481 
Staghorn-moss I 47 
Stammer-wort 3 341 
Stanleya 2 169 
Starchwort 442 
-hyacinth Tse 
Star-bloom 730 
-chickweed 2 43 
-eyed grass 
I 543; 2 715 
Star-flower I 509, 534, 


543; 244, 251 
Star-grass, see Grass 
Star-leaved Gum 2 235 
Star-of-Bethlehem 


I 509-10; 2534; 
3251 

Star-quince 2327 
-root pee arig 
Stare I 363 
Starlights 2 429 
Starry Campion 63 
Starveacre 2 115 
Star-violet 3 252-4 
Starwort, Bog 2 43 
Drooping I 489 
Glaucous 2 44 
Greater 2 44 
Lesser 44 
Mealy eat 
Mountain 2157 
Pine 444 
Sandpaper 444 
Water 2477 
Yellow 3 457 


Starworts 3 406-35, 444 


Stavewort 3 542 
Stedfast 2 461 
Steel-weed 3 430 
Steep-grass 3 226 
Steeple-bush 2 245 
Stenanthium I 490 
Stenophyllus 319 
Stenosiphon -2 610 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Stenotus 3 379 
Stepmother 2 563 
Stevia 3 361 
Stewartia 2 526-7 
Stick-a-back 3 259 
Stick-button 547 
Stickle-back 259 
Stickseed 2 266; 
_. .3 77-9, 487, 497 
Stick-tight 3 77-8, 497 
-weed 2260" 

_ 3 341, 426 

Old Virginia 3 426 
Stinging-bush 2 462 
-nettle I 633 
Stipa I 176-7 
Stitchwort, Bog 2 43 
Fleshy 46 
Greater 2 44 
Lesser 45 
Long-leaved 45 
Long-stalked 45 
Marsh 43 
Northern 46 
Swamp 43 
Water 46 
Stonecrop, Biting 2 208 
Crooked Yellow 209 
Ditch 210 
Mossy 208 
Narrow- petaled 209 
Nevius’ 210 
Nuttall’s 208 
Reflexed 2 209 
ee ona ed 210 
Torr 208 
Vereen Wild 210 
Stone-mint 3 146 
-root 153 
-rue Tt 29 
-seed, Salfern 3 87 
-weed I 661 
STORAX FAMILY 2 721 
Storax 723 
Stork’s-bill 430 


Stramonium 3156, 169 
Strangle-tare 


2 411; 3.235 
Strangle-weed 3 48 
Straw-bell I 518 


Strawberry 2 259-261 


American-wood- 261 
Barren 251, 253, 269 
Bog 258 
Dry 269 
European-wood- 260 
Gray’s 260 
Hedge 260 
Indian 215, 259 
Mock 259 
Mountain 259 


Northern-wild 2 259 
Scarlet, Virginia, 


259, 260 
Southern dry- 269 
Wild 251 
Yellow 259 
Strawberry-blite 2 15 
-bush tree 132, 491 | 
-spinach 200s 
STRAWBERRY-SHRUB 
FAMILY 2) 132 
Strawberry-shrub 
2 132; 3 39 
-tassel 2 449 
-tomato 
3 157-8, 161-2 
-tree 2 491 
Straw-lilies I 519 
Stub-wort 2 431 
Stud-flower t 488 
Styptic-weed 22236 


Vo. III. 
Succory, Gum 
3 314, 306 
Dwarf Hog’s, or 4 
Swine’s 309 
Lamb 309 
Wild 305 
Succory Dock-cress 
306 
Suck-bottle Saree 
Suckles 2 355 
Suckling, Yellow 354 
Lam 358 
Sugar-berry 
I 5203 2288 
Sugar Maple 2 496 
-pear 291-2 
Sugar-plums 355, 292 
Sugar-tree 496 
Sullivantia 224 
Sultan, Sweet 3 560 
Sumac FaMILy 2 480 
Sumac, Amer. 481 
Black 481 
Chinese 446 
Climbing 484 
Common, Dwarf 481 
Elder-leaved 287 
Fragrant 482 
Hairy 481 
Ill-scented 483 
Mountain 481 
Pennsylvania 2 482 
Poison 483 
Scarlet, Sleek 482 
Smooth 481-2 
Staghorn 481 
Swamp 483 
Sweet-scented 482 
Trailing 484 
Upland 481-2 
Velvet 481 
Virginia 2 481 
White 482 
Summer-cypress 2 16 
-lilac 175 
-savory 2.137 
-snowflake I 509 
SUNDEW FAMILY 2 202 
Sundew 203-4 
Sun-dial 348 
Sundrops 2 600-2 
Sunflower 716 
Brook 3 495 
Common 47 
False 467, 510 
Few-leaved 480 
Garden 478 
Giant 481 
Hairy 484 
Hoe Wood- 479 
Judge Daly’ s- 482 
Kellerman’s- 483 
Linear-leaved 479 
Maximilian’s- 482 
Narrow-leaved 479 
Oblong-leaved 484 
Pale-leaf wood- 485 
Prairie 478 
Purple-disk We 
Red 3 475 
Rhombic-leaved 480 
Rough 483 
Saw-tooth 483 
Small wood 481 
Smooth 481 
Showy 486 
Stiff 480 
Stiff-haired 485 
Swamp 479, 510 
Tall 481 
Throatwort 485 
Thin-leaved 484 


VoL. III. ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 633 
Sunflower Sweet-root, Tea, Continental 2 677 Thistle 
Thick-seed -rush I 446 Labrador 677 Musk 3 554, 556 
3 491-8, 498-9 | = -scabious 3 439 Liberty 711 Nebraska 3 552 
Wild 3 457, 478, -scented-shrub 2 132 Marsh 677 Oat 556 
481, 484 -sedge I 446 Mexican 2ELA Our Lady’s 560 
Woodland 483 -shrub 25132 Mountain 693 Pasture 552 
Wooly 486 -slumber 140 New Jersey - oe Pitcher’s 550 
Sun-plant 2eAT -Sultan 560 Oswego Plume 549 
-rose 539 -Susan 2 66 Paraguay ae Plumeless 554 
-weed 473 -tree 496 272 Prairie 551 
Supple-Jack 502 -walnut I 581 Prairie 2 455 Prickly 553 
Susan Sweet-william 2 69, 75; South Sea 2 489 Queen Ann’s) 3 554 
Black-eyed 3 470 3 42, 556 Swamp 677 Queen Mary’s ~ 556 
Brown-eyed 470 Barbadoes 3 42 Walpole 2 504 Roadside 3549 
Suterberry 2 444 Childing 2 74 ; Russian 2 
Wild 2 366 25 
Swallow-wort I4I Mock 66 Teab = baat St. Benedict’s 3 * 560 
Black 3 36 Old-field AgaG ot a #2 Z| Scotch, Silver, ~ 556 
range 3 25 i 273; 3 54 Sow, Swine, 16a5 
Silky 30 | Sweet-willow 1 584 | lea-plant omer Spear, Tall, 3 uae 
Swamp-apple 2 678 | Sweet-wilson 2 218| Tear-thumb I 673 Star 559 
-berry 2 278] Swine-cress 167 | TEASEL FaMILyY 3 288 American 559 
-brake I 7 | Swine’s-bane 2 13| Teasel,Common 289 Swamp, Way, 553 
-cabbage 445 | Swine-grass I 661 Card, Draper’s, 289 Virgin Mary’s 3 555 
-candles 2 712! Swinies Beatz Fuller’s, Wild, 289 Virginia 550 
-fern 18 | Switch-grass I 141 } Teil, Tile 2 512 Water 289 
-globeflower 2 87 | Sword-flag 540 | Ten-o’clock-lady 1 509 Wavy-leaved 550 
-hornbeam 2 665-6 -grass 170, 330 | Tench-weed is ay 553 
-lily I 578! Sycamore Tentwort 29 Welted 555 
-pink 488 I 628; 2 242 | Tetter-wort 2 141 Yellow 2 139; 3 553 
-redberry 563 False 2 242]! Thale-cress 2075 -spined 551 
-robin I 444; 2 609] Synandra 3 118 | Thalia I 546 | Thorn 2 poate 
-spruce I 61 | Syndaw 2 264; 3 170| Thaspium 2 639 See Haw 297-321 
-tea 2 677 | Synthyris 3 198 | Thatch I 141 Alleghany 2 309 
-tupelo 666 Bull’s 198 Creek I 223-4 Arnold’s 318 
-wood 2 575; 3 255 Western 199 | Thick-stamen, Asa Gray’s 311 
Swan-weed 3 419 | Syrian-tobacco 3 170 American 2 480 Ashe’s 300 
Sweat-root 65 | Syringa 2 231-2 | Thimble-berry Beadle’s Yellow 309 
Sweat-weed 2 514 2 276-7, 280 Beckwith’s (Miss) 
Sweating-weed 2 525 | Tacamahac I 588 -flower 3 115 313 
Sweeps 1 485; 3 557| Tag-alder 611 | Thimble-weed Bicknell’s 305 
Sweet-allison 2 153] Talinum 2 36-7 3 99, 379, 473 Black 39273; 
-alyssum 153 | Tallow-bayberry 1 585 | Thimbles 3 204, 205 2 302, 326 
Sweet-bay 2 82, 633 -root 575 Fairy 204 Blanchard’s 304 
-bean 339 -shrub 585 Witches 3 204 Bosc’s 310 
-Benjamin 3 526] Tamarack I 60| THISTLE FAMILY 3 347 Boynton’s 309 
-berry 273 | Tan-bay 2 527 | Thistle 3 549-56 Brainerd’s 303 
-Betties 2 132] Tangleberry, Blue 695 Argentine 556 Brown’s 300 
-birch 1610; 2704] Tangle-foot,-legs 3 269 Asses’ 555-6 Buck 326 
-Betty PA fay ee, -tail 2 208 Pank 3 549, 554 Bush 316 
Sweet-brier 2 2854 Tank 634 Barnaby’s 560 Canadian 318 
-broom 392 | Tansy Bu522 Bell, Bird, Button, 549 Canby’s 299 
-bubby Zines Dog’s 2 258 Bitter 560 Caughuawaga 301 
-bush I 586 Goose 2 258 Blessed 3 560 Chapman’s-hill 301 
-cane 446 Lake Huron 3 522 Blue 94, 349 Christ’s 322 
Sweet-cicely 2 627-9 Sneezewort, or Boar 549 Cockspur 207 
Hairy 627 White Bs 05 Bull 549, 552 Common 302 
Sweet-clover 2 352-3 Wild 2 258; 3 341 Buck 554 Dog 284 
-fern 1 586 | Tansy-mustard 2 171 Briery 2 622 Douglas’ 321 
-ferry 586 | Tansy-ragwort 3 522 Bur 3 549 Downy 319 
Sweet-flag I 446 | TAPE-GRASS FAMILY Canada, Corn, 39553 Dunbar’s 314 
False 540 I 104 Card 289 Dwarf 320 
Sweet-gale 584 | Tape-grass 106 Common, Bur 3 549 Edson’s 311 
Canada 586 | Taper hedges Cotton, Down, 556 Eggert’s 317 
Sweet-grags I 91,197, Hig-high 8.173 Creeping, Cursed, Egyptian 322 
46; 3 267] Tar-fitch, Yellow 2 415 553 Emerson's 319 
Sweet-gum 2 235 | Tar-weed 582; 3 371 Curled 555 Evergreen 322 
-hairhoof Bu2o7 || hare I 2623.2) AinI—12 Down 556 Fernald’s 304 
-hay 2 249 | Tare-grass I 282 Field 3 549 Fire 322 
Sweetheart 208 -vetch 415 Flodman’s 3551 Fretz’s 313 
Sweethearts 3 259 | Tarrify 2 172 Flowering 2 139 Fruitful 299 
Sweet-jarvil 2 627-8 | Tassel, Milky 31.317, Fragrant 3.552 Gattinger’s 314 
Sweet-johns 2°75 -flower, Red 370 Fuller’s 289 Globose 302 
SWEET-LEAF FAMILY White 369 Hard 30S Grove 304 
2 721 | Tassel-grass, Sea- 1 88 Hare’s 3.307, Gruber’s 312 
Sweet-leaf 721 | Tassel-pondweed 1 88 Hedgehog 2 570 Haw 2 297, 317,319 
pMany 2 131393 137 -weed 3 341 Hill’s 3 552 Heart-leaved 321 
-myrtle I 446 | Tawkin 445 Holy 3 555, 560 Hedge 321 
-oak 585 | Tea FAMILY 2 526 Horse 3 318, 549 Jack’s 306 
-pea, Wild 2 372 | Tea,Appalachian 2 487 Indian 3 2890 Jesup’s 312 
-pepper-bush 2 667 Blue-mountain 3 389 Jamaica 2 139 Jones’ 300 
-potato, Wild 3 43 Canada 9, 693 Marsh 3 554 Kellogg’s 315 
-rocket 2078 Carolina 2 487 Milk 3 316-17,555 Lange's 307 


634 


Thorn 
Large-fruited 2 300 


Long-spined 301 
Macauley’s 305 
Maine’s 313 
Maize 3 569 


Marshall’s 2 299 
Martha’s Vineyard 


2 298 
May 319 
Missouri . 314 
Mouse 3: 5559, 314 
Mrs. Ashe’s 2 300 
Narrow-leaved 320 
Newcastle 297 
New-river 302 
Oakes’ 305 
Oblong-leaved 316 
Ovate-leaved 306 
Palmer’s 298 
Padus-leaved 308 
Parsley-leaved 320 
ear 302 
Pick’s 308 
Pin 2907 
Poplar-leaved 308 
Prairie 303 
Pringle’s 317 
Red-fruited 319 
Reverchon’s 298 
Roan Mountain 310 
Rough-leaved Bu 
Round-leaved 306 
Scarlet 317 
Shining 307 
Small-fruited 320 
Southern 307 
Stone’s 308 
Tatnall’s 2 318 
Thin-leaved 315 
Tree 307 
Vail’s (Miss) 303 
Variable Zhe 
Virginia 321 
Washington 321 
Waxy-fruited 315 
White 
300, 302, 317, 319 
Woolly 316 


Thorn-apple 2 297, 302, 
317; 3169, 170 


-bush 2 297, 317 
Re Sa7, 
-locust 339 
gel 297, 302, 317 
-poppy 139 
Thorough-grow 3 361 
-stem 361 
Thorough-wax 2 639; 
3 361 
Thoroughwort 2 639; 
3 357-61 

Common, Hairy 
360, 361 
Hyssop-leaved 358 
Large-flowered 3 363 
Late-flowering 357 
Pink 362 
Rough 360 
Rouds leaved 360 
Small-flowered 359 
Tall 359 
Torrey’s 358 
White 358 
White-bracted 357 
Vervain 360 
Thousand-leaf 3 515 
Thread-foot 2 205 
-and-needle I 513 
Three-birds 562 
Three-leaved Ivy 2 484 
-nightshade I 523 


Three-seeded Mercury 


2 357-8 
Three-square I 330 
Thrift 2 718 
American 717 
European 718 
Lavender 2 717 
Sea 718 
Throat-root 
2 270; 3 204 
Throat-wort 
2 272; 3 367 
Through-grow 3 361 
Thrumwort I 94 
Thunder-flower 
2 44, 68, 137 
-plant De ier 
Thyme 3 113, 139,141 
Basil 139 
Creeping I4I 
Field, Horse 138 
Mother of 141 
Mountain 139 
Shepherd’s 141 
Virginia 142 
Wild 141 
Thyme-leaf 3.514 
Tickle-grass I 139 
Tickle-weed 494 
Tickseed 2 395-402; 
3 490-4 
Cress-leaved 3 493 
Garden 493 
Greater 492 
Hairy 490 
Large-flowered 491 
Larkspur 493 
Lance-leaved 490 
‘Leafy-bracted 496 
Lobed, Running, 491 
Pink, Rose, 494 
Star 490 
Stiff 492 
Tall 491 
Thick-leaved 490 


Whorled, Wood, 492 
Tickseed Sunflower 


; f 3 491-8 
Tick-trefoil 2 395-403 


Canadian 399, 401 
Cream-flowered 397 
Dillen’s 400 
Few-flowered 396 
Hairy, Small-leaved 
2 402 
Hoary 398 
Illinois 401 


Large-bracted 399 
Naked-flowered 395 


Panicled 399 
Point-leaved 396 
Prostrate 397 
Rhomb-leaved 2 400 
Rigid, Showy, 401 
Round-leaved 397 
Sand 2 396 
Sessile-leaved 2 398 
Smooth 399 
Smooth Small- 
leaved 402 
Stiff 398 
Trailing 397 
Velvet-leaved 400 
Tick-weed 3 136 


Tiger-lily 1 502-4, 542 


Tiger’s-mouth 3 178 
Timothy I 191 
Wild 186 
see Grass 
Tine-grass 2 409 
Tine-tare 411 
-weed 411 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


Tinker’s-weed 3 275 
Tipsin, Tipsinna 2 363 
Tiresome-weed I g1 
Tisswood 245335 fee 
Tithymal 473 

Titi, Red, White 2 485 


Toad-flax 
I 640; 3 175-7 
Bastard I 640 
Blue R717 7. 
Broom-leaved 177 
Ivy-leaved is 
Pale-blue 178 


Round-leaved 3 


Sharp-pointed 176 
Wild 177 
Yellow 177 
Toad-grass I 469 
-lily 2 79 
-pipe, Tadpipe 1 39 
-root 2 90 
Toad’s mouth 3 178 
Toadspit 448 
Toad’s tail I 47 
Tobacco 3.170 
Indian 170, 451, 
303, 170, 451 

Ladies 3 453 
Long-flowered 7 
Mountain 534 
Oregon 285 
Poison 169 
Rabbit 454 
Real 170 
Sailor’s- 527 


Syrian, Wild 303, 170 


Woman’s 451 
Tobacco-wood 2 235 
-root 3 285 
-weed 354 
Tofieldia I 487 
Tom Thumb 2 415 
Tomatillo 3 158 
Tomato 168 
Cherry 168 
Husk 161 
Strawberry 


3 157; 161-2, 414 


Tongue, Devil’s 2 572 
Tongue-grass 
2 43, 165-6 
Wild 164 
Toothache-grass I 225 
-tree 2 444, 617 
Tooth-cup 579 
Toothroot 187 
Toothwort 158 
Purple-flowered 
187-8 
Toper’s plant - 266 
Topinambour 3 486 
Torches 173 
Tormentil, Trailing 
2 250 
Torrets I 422 
Tory-weed 3 76 
Touch-and-heal 2 533 
Touch-me-not 
184, 440-1 
Wild 440 
Tow-cok 425 
Tower-cress 181 
Tower-mustard 181 
Townsendia 3 402-3 
Toywort 2 158 
Tragia 459-60 
Traveller’s-joy 122 
-delight 418 
Treacle-mustard 
172,175 
Treaclewort 168 


VoL. III. 


Tread-softly 
3 165; 2 462 
Treasure-of-love 2 208 


Tree-beard I 456 
Tree-fern 17 
-moss LAs 
Tree-of-heaven 2 446 
Trefoil 
(see Clover) 2402-8 
Bean 3 18 
Bird’s-foot 2 359 
Bitter 3 18 
Black, Great 2 351 
Golden 2 I01 
Heart 352 
Hop, Least 354 
Marsh 3 18 
Melilot 2 351 
Shamrock 354 
Shrubby 445 
Sour 431 
Spanish 2 351 
Water 3 18 
White 2 358 
Wild, Yellow 354 
Triantha I 486 
Trick-madam 2 209 
Trickle 188 
Trifoly, Sea 715 
Sour 431 
Trilisa 3 369 
Trillium 
Red, Purple, I 525 
Trinity 461 
Herb- 3 IOI 
Trinity-violet 2 563 
Trip-madam 209 
Trip-toe 3 269 
Troll-flower 2 87 
Trompillo - 3 166 
Trout-flower 506 
-lily 506 
True-love I 525 
Trumpet, Devil’s 3 169 
Trumpet-ash 237 
TRUMPET-CREEPER 
FAMILY 3 236 
Trumpet-creeper 237 
Trumpet-flower 237-8 
Evening 2 730 
Trumpet-honeysuckle 
3 280 
Scarlet 280 
Trumpet-leaf 2 202 
-milkweed 3 319 
Trumpet-vine 237 
-weed 356-7, 319 
Trumpets 2 202; 3 319 


Yellow 2 202 


Tube rose, Wild 1 566 
Tuber-root 3°25 
Tule 1335 
Tulip-tree 2 83 
Tulip, Wild 505 
Tulip poplar 83 
Tumble-weed I 139; 


2 3, 16, 21, 99, 361 


Tunhoof 3st 
Tunica 272 
Tupelo 1 639; 2 665 
Large 2 666 
Southern 666 
Swamp, Water 666 
Turin-grass I 686 
Turkey-beard 488 
-berry 3°277 
-claw L575 
-corn 2 142-3 
-grass 3 259 
-oak I 619, 622 
-pea 2 142, 653 
-pod 175 


Vo. III. 


Turkey-troop I 670 


Turmeric, Indian 


2 85, 140 
Turmeric-root 85 
Turnip 193 

Cree, Dakota, 363 
Marsh 442 
Pepper 442 
Prairie 363 
St. Anthony’s 2 112 
Wild 442 
Turnpike-geranium 14 
Turnsole 473; 3 75 


Turpentine-weed 3 461 


Turrets TAs 
Turtle-bloom 3 181 
-grass I gI 
Turtle-head 3 181-2 
- Tussocks I 203 
Tway-blade 567-8, 572 
Twi-foil 568 
Twigwithy aay 
Twin-berry Bee55 
-flower 3 276 
-grass I 206 
-leaf 2 129 
-sisters 3 276 
Two-eyed berries 
255, 276 
Twisted stalk 
I 520, 566 
Two-lips 550 
Two-penny grass 2 712 
Ulim 349 
Umbrella-leaf 129 
-grass 338 
-leaves 3 532 
Umbrella-tree 
2 81-2, 663 
Fraser’s 81 
Silver-leaf 81 
Umbrella-wort 
31-2, 663 
Uncinia I 389 
UNICORN-PLANT 
FAMILY 3 239 
Unicorn-plant 239 
False I 489 
Unicorn-root, or -horn 
489, 511 
Universe-vine 2 693 
Unlucky-tree I 58 
VALERIAN FAMILY 
Si 3 284 
Valerian 3 63, 284-6 
American 1 548,550; 


3 63, 284-5 
Cat’s, Common 3 286 


Creeping, Great 63 
Edible 3 285 
False 544 
Garden, or Great 286 
Greek 65 
Marsh 285 
Northern 285 
Swamp 285 
Wild 1550; 3 284-6 
Vandal-root 286 
Vanilla, Carolina 369 
Vanilla-grass ee 
Vanilla, Nodding 172 
Vanilla-leaf, -plant 
: 3 369 
Varnish-tree, False 
2 446 
Vegetable-hair 1 456 
-oyster 3 313 
-satyr I 552 
Velvet-leaf 2 521 
-plant 173 


Velvet-weed 2 521 
Velvets 547 
Venus’-bath 3 289 
-buskin I 548 
-comb 2 627 
-hair E31 
-cup 289 
-looking-glass 298 
-pride 251, 253-4 
-sock I 548 
Veratrum 494-5 
Verbena 3 94-7 
VERVAIN FAMILY 94 
Vervain 95-73; 3130 
American 95 
Base 201 
Blue 95 
European, False 95 
Hoary 96 
Mallow 2 515 
Prostrate 3 96 
Vetch, American 2 409 
Bird, Blue 409 
Bush 412 
Carolina 410 
Choke 3 234 
Common 2 4II 
Cow 409 
Hairy 411 
Kidney 360 
Milk 377-88 


Narrow-leaved, 
American 2 409 


Louisiana, Pale 410 
Pebble 411 
Purple 409 
Sensitive Joint- 393 
Slender 411 
Smaller Common 412 
Small-flowered 410 
Spring, Tare 411 
Tufted 409 
Wild 359 
Vetchling 
Cream-colored 2 415 
Low 416 
Marsh 414 
Pale 415 
Prairie 414 
Showy 415 
Yellow 415 
Viburnum 3 273-4 
Naked, Sweet 273 
Vincetoxicum 3 37-9 
Vine, Wild 2 506 
Alleghany 143 
Canary, Cypress 143 


Enslen’ S, or Sand 


3 36 

Pepper 2 510 
Wild 2 506 
Wild potato 3 43 
Vinegar-tree 2 482 


VIOLET FAMILY 545-64 


Violet 545-64 
Adder’s I 570 
Alpine 2 561 
Arrow-leaved 553 
Beaked 562 
Bird’s-foot 547 
Blue 548-9 
Bog 551; 3 226 
Calathian BT 
Canada 2 560 
Canker 562 
Chicken-fighters 547 
Coast 554 
Common-blue 548 
Crowfoot 547 
Cut-leaved 553 


Damask, Dame’s- 


ENGLISH INDEX, INCLUDING POPULAR PLANT 


Violet, Dog, 
American 2 560 
Dog’s-tooth I 506 
White, Yellow 506 
Early Blue 
2 553, 560, 547 
English 558 
Field 561-3 
Fringed 552 
Garden 563 
Great-spurred 555 
Green 564 
Hairy 559, 550 
Hairy Yeliow 559 
Halberd-leaved 
Yellow 558 
Hand-leaf 547 
Hens 2 560 
Hood-leaf 547 
Hooded Blue 548 
Hooked 561 
Horse, Horse-shoe, 
547 
Kidney-leaved 555 
Lance-leaved 257 
Larkspur 554 
Le Conte’s- 2 551 
Long-spurred 562 
Long-stem purple 548 
March 558 
Marsh 555 
Marsh Blue 552 
Meadow Blue 548 
Missouri 2 549 
New England 550 
Nodding 564 
Northern 550 
Nuttall’s 558 
Ovate-leaved 552 
Pale 560 
Pansy 2 562- 
Plains 552 
Prairie 554 
Yellow 558 


Primrose-leaved 557 


Prostrate Blue 562 
Rattlesnake’s- 

I 506): 2 552 
Round-leaved 2 557 
Rydberg’s- 559 
Sand = 547, 552, 561 
Selkirk’s- 555 
Smoothish Yellow 

559 

Southern Wood _ 550 
Southern Coast 554 
Snake 547 
Spear-leaved 558 
Spade-leaf 553 
Stepmother 563 
Striped 560 
Star 3 252-4 
Stone’s 548 
Sweet 558 
American 560, 556 


Sweet-white 2 55 


Northern 556 
Large-leaved 556 
Three-lobed 548 
Triangle-leaved 553 
Trinity 563 
Velvets 547 
Water 557 
Western blue 551 
Wood 547-9 
Whorl-leaf 564 
Wooly blue 549 
Yard 292 
Yellow 2 557 
Viper’s-bugloss 3°94 
-herb, -grass 04 


2 175 | Virginia-creeper 2 511 


NAMES. 635 


Virginia-silk 

ViRGINIA WILLOW 
FAMILY 2 223 

Virginia Willow 233 

Virgin’s Bower 

122, 374 
126 


3 30 


Purple 


Wafer Ash 
Wahoo 


Wait-a-bit 
Wake-robin 
I 442, 523-6 
Dwarf White 524 
Early,Green 1 524 


445 

I 626-7; 
2 491, 512 
I 528 


Ill-scented 529 
Nodding 526 
Painted 526 
Prairie 524 
Purple 525 
Showy $24 
Virginia 


Walking-fern,-leaf 1 ape 

Wall-cress 2 175 
see Rock-cress 

Wall-flower, Western 


2: 0735.39.25 
Wallink 3 200 
Wall-moss 2 208 
Wall-pepper 208 
-rue Tz 
-rocket 2 194 
WaLnuT FAMILY 
I 578-81 
Walnut 579 
Black, Lemon 579 
Satin 2 235 
Sweet 581 
White 579 
Walpole-tea 504 


Wandering-jenny 2712 


Wandering-jew 3175 
-milkweed Bhat 
-Sally 2 712 

Wankapin 77 

Warlock 193-5 

Warmot Ce 

Wart-cress 2 167 
-grass 473-5 

. -lesser 167 
-weed 473 
-wort 14I, 167, 473; 

3 455 

Watches 2 202 

Water-agrimony 3 495 
-archer I 99 
-arum 444 
-awlwort 2 159 
-bean 77 
-beech 1606; 2242 
-blinks 2 39 
-blobs 85 
-cabbage 79 
-calamint 3 152 
-caltrop 2 612 
-can 78 
-carpet 230 
-celery I 106; 2 I11 
-chestnut 2 612 
-chickweed 2 478 
-chinkapin 77 

Water-cress 2 186, 197 
American 186 


Creeping Yellow 160 
Lake 164 
Marsh, Yellow 161 
Mountain 186 
Round-leaved 186 
True 162 


Water-crowfoot 
White, stiff 2 


636 


Water-crowfoot 
Yellow 105, 648-50 | 

Water-cup 202 
-dock 445 
-dragon 1444; 285 
-dropwort 2 638 
-elm I 626-8 
-eryngo 2 622 
-feather 616, 710 
-fennel 478 
-fern 1G, 

Water-flag 537 
Yellow 540 


Water-gladiole 3 300 
-goggles, -gowan 2 85 


-grass Tera 
2 162, 649 
Large reas 
gum 2 666 
Fremloel 658 
-hemp 2 
-hickory 581 
-hoarhound 3 147-8 
-hyssop 193 
-jessamine 194 
WaATER-LEAF FAMILY 
3 65 
Water-leaf 26; 3 65-6 
-lentils I 448 
WATER-LILY FAMILY 
277 
Water-lily 1444; 319 
Great Yellow NG 
Little 76 
Sweet-scented 79 
Tuberous 80 
White 79-80 
Small 80 
Water-lily-tree 81 


Water Marigold 3 500 

WATER-MILFOIL 
FAMILY 2 612-15 

Water-milfoil 


116, 614-16 

Hooded 3 229 

Water-melon 291 

-navelwort 2 614, 649 

-nerve-root 3 26 
WATER-NUT FAMILY 


2 611-12 
Water-nut 277 
Jesuit’s 2 613 
Swimming 2 612 
Water-nymph 2 79 
-oats I 168 
Water-Parsnip 2 655 
Carson’s 656 
Creeping 655 
Cut-leaved 2 655 
Hemlock 656 
Lesser 655 
Narrow-leaved 655 
Water-pennywort 
2 648-50 
-pepper I 670 
-pimpernel 2 710 
WaTER PLANTAIN 
FAMILY I 93 
Water-plantain 
I 94; 3 247 
Dwarf 95 
-poplar I 590 
-purslane 2 579, 585 
-rice 68 
-rush bf ee 
-seg, - <egs 540 
_ -shamrock 3 18 
-shield 2 75-6 
W ATER-SHIELD 
FAMILY 23 Gi) 


-spike (any Potamo- 
geton) I 74-87 


; Water 
-speedwell 3 200 
-stargrass I 464 
WATER STARWORT 
FAMILY 2EAT7 
Water-starwort 477-8 
-target 76 
-thyme I 105 
-torch I 68 
-trefoil 18 
-tupelo 2 666 
-violet 710 
-weed I 105-6 
Nuttall’s 105 
-willow 3 243 
WaTERWORT FAMILY 
2 537 
Waterwort 2 538 
Water-yarrow 710 


Waxberry 1585; 2 276 


Wax-bush 2 582 
-dolls 2 146 
Wax-myrtle I 585 
Wax-pinks 2 41 
-plant 2 66 
Wax-weed 2 582 
Wax-work 3 493 
Roxbury 2 49 
Way-bread 3 245 
Wayfaring Tree 
269, 274 
Way-grass I 661 
Waythorn 2 502 
Weathercock 2 440 
Weather-glass 2 716 


Welcome-to-our-House 


2 474 
Weld 2 200 
Well-grass 2 162 
Western Dropwort 
2 248 
Wheat 
Black 224 
Cow 3 224 
Wheat-grass I 284-5 


Wheat, Saracen’s I 672 
Whin 2 349 
Cat 284 
Dyer’s 350 
Whip-grass I 349 
Whipporwill 550 
-boots 2 202 
-shoe I 540-50; 
2 202 

Whip-tongue Beas 


Whistle-wood 2 497,512 

WuiteE-ALDER FAMILY 

2 666 

White-alder 2 667 
-topped Aster 

3 405-6 

White Allison 2 180 

-apple 418 


-arrow-arum I 444 
White-ash Herb 2 654 
White-bark I 587 

-bay 2 82, 133 

-beads 2 90 

-Ben 2 64 

-Benjamin I 526 

-berry 2 90 

-flow 148 

-bush 2 667, 691 

-cankerweed 3 335 

-cap 2 245 

-coolwort 2 224 

-daisy 2) Bins! 

-Devil 426 

-fringe 2 728 

-gum 235 

-hearts 2 142 

-hellebore,Am. I 494 


White-heart Hickory 


I 582 
White Man’s-foot 
3 245, 246 
-weed 518 
White-oak I 622-3 
Box I 622 
Mossy-cup 623 
Rough 622 
Running 625 
Swamp 623 
Water 623 
White-osier 2 688 
-robin 2 68 
-root 3 25 
-rosin-tree 157. 
-sage 2 20 
-soapwort 2 69 
-thorn 300 
-top 3 440 
-walnut I 579 
-weed 3 518 
-wood 
2 83, 511-12, 691 
Whites I 214 
Whitlow-grass 2 148-52 
Alpine Lge 
Arctic 149-50 
Branching 151 
Canescent I51 
Carolina 149 
Golden Ts2 
Hoary 150 
Rock-cress 150 
Short-fruited 151 
Twisted 150 
Vernal 148 
Wedge-leaved 149 
Wood 152 
Whitlow-wort 2 27-8 
Whitten-tree 3 270 
Whorl-leaf 2 564 
Whorlywort 204 
Whortleberry Bears 
2 693 
Bog 699 
Blue 6905 
Great 700 
Red 2 697 
Squaw 697 
Wickakee ; 3 214 
Wick, Wicken 
I 2833. 2-319 
Wicke . 206or 


Wickup 2 5127573, 592 


Indian 590 
Wicky 2 683 
Wicopy 2 575,590 

Herb 590 
Widow's Cress 2 209 
Wild-pink T 562 

-brier 2 284 

-hippo 2 469 

Tube-rose 565 
Wilding-tree 290 
Wilgers I 597 
WILLOW FAMILY 1 587 
Willow 591-605 

Almond-leaf 593 

Arctic 604 

Ausier 597 

Autumn 594 

Balsam I 596 

Barclay’s 603 

Basket I 597 

Bay 2 590 

Bay-leaved I 594 

Beaked I 599 

Bearberry 601 

Bebb’s 599 

Bitter 597 

Black 592-3 


ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 


eee ———_————— 


Vor, ie 


Willow, Black 


Scythe-leaf 1 5092 


Blooming 2 590 
Blunt-leaved 1 601 
Bog I 599, 602 
Brittle 594 
Broad-leaved 596 
Brown’s 605 
Bush 600 
Button 3 255 
Common 595 
European 595 
Crack gue 
Crane 255 
Dark Long-leaf 3 599 
Desert 604 
Diamond 596 
Drooping 595 
Duck 595 
Dwarf 601 
Dwarf-gray 600 
Flowering 2 590 
French 2 590 
. Furry T5071 
Garb 595 
Glaucous 599 
Glossy 593 
Golden Osier : 
595; £1459 
Green-scaled 602 
Ground I 604, 666 
Hairy 603 
Heart-leaved 596 
Herb-like 601 
Hoary 598 
Huntingdon 595 
Livid 599 
Long-leaf 595 
Low-bush 600 
Missouri 596 
Narrow-leaf 595 
Net-veined 602 
Northern 605 
Osier I 595,597 
Peach-leaved 593 
Persian 2 590 
Prairie I 600, 604 
Primrose 2 5890 
Purple I 597 
Pussy 592 
Red 1 595; 2 661 
Red-wood I 594 
Ring 595 
River-bank 595 
Roman 2 724 
Rose I 597; 2 661 
Sage I 598, 600; 
2 581 

Sandbar I 505 
Satiny 598 
Scythe-leaved 592 
Shining 593 
Shrub 595 
Silky 598 
Silver 599, 603 
Slender 594, 599 
Snap 594 
Stag’s head 594 
Swamp 592, 594 
Sweet 584 
Tea-leaved 600 
Twig Withy 597 
Varnished 594 
Velvet-osier 597 
Virginia 2 233 
Waghorne’s I 604 
Ward's I 593 
Water 3 243 
Weeping I 595 
Whipcord 597 
White I 595 
Wilger’s 597 


Vot. II. ENGLISH INDEX, INCLUDING POPULAR PLANT NAMES. 637 
Willow Wintergreen Woodbine 2122,511; | Yarrow 
Wrinkle-leaf 602 False 26 3 46, 278-9 Water 2 710 
Yellow 1459 Flowering 452 American 3 278 | Yaupon 487 
Willow-grass I 666 Greenish-flowered Carolina Wild 2 730 | Yaw-root 462 
Willow Herb 2 591-5 669 Fragrant 3 278 | Yellow-bark I 619 
Alpine 2 591 Larger 668 Rough 278 -ash 2 343 
Bay 590 Lesser 670 Small 279 -balm 711 
Broad-leaved 2 590 Liver-leaf 2 670 | Wood-brooms 3 289 -bells I 506 
Downy 2 592 One-flowered 671 -cinquefoil 2 250 -cress, hispid 
French 590 One-sided 671 -cress, Yellow 161 2 160-1 
Golden 711 Pear-leaved 668 -flower 100 -eye 85 
Great 590 Round-leaved 668 | Wood-lily 1 502, 522; -dock, see Dock 1 657 
Great Hairy 591 Serrated 670 2 601,670 | YELLOW-EYED GRASS 
Hooded 3 109 Sharp-petaled 670 -nep 2 657 FAMILY I 450 
Hornemann’s 2 593 Shin-leaf 669-70 -Tip, -rowel 3 267 | Yellow-eyed Grass 
Linear-leaved 592 Spicy 693 | Woodland-beauty 2 262 I 451-3 
Marsh 592 Spotted 672 | Woodruff, Sweet 3 267] Yellow-flower 2 192 
Milk 580-1 Spring 693 | Wood-rush I 483-5 -gum-tree 665 
Night 595 | Winter-pink 2 692 | Woop SorreL FAMILY -ladies’-slipper 1 550 
Northern 593 -plum 720 2 431-5 -moccasin-flower 550 
Panicled 593 -rocket 177 | Wood-sorrel 432-5 -pine 58 
Purple 581 -sweet 3 140 Purple 431 -puccoon 2 85 
Purple-leaved 2 593 -weed 2433 3 203 Yellow 433-5 -rocket, see Rocket 
Purple-veined 593 | Wire-grass 1470,661 | Wood-sour 2 127, 431 711 
Pimpernel 597 | Wire-ling 2 479 | Wood-sower 431 -rod 3 380-908 
Soft 592 -weed I 661 | Wood-wax 350 -root 88-9 
Spiked 590 | Wise-weed 3 426 | Woodsia,seeFern 1 12 -shrub 89 
Swamp 580, 592 | Wistaria 2 373-4 | Wool-grass 337 | Yellow-seed 2 164 
Yellow 711 | Witch-alder 2 234 | Woolen 3 173 -skegs I 540 
Willow-Oak I 621 -gowan 3 315 | Woolly Pipe-vine 1 646 -snow-drop I 506 
Willow-weed -grass, see Grass World’s Wonder 2 73 -star-grass 534 
I 666-8; 2 581 -hobble, -hopple, Worm-grass 2 730 -star 3 510 
-wood 2 581 269-70 | Worm-seed 172 -starwort 3 457 
Willow-wort 2 711 | Witcu Haze. Famrity | Wormweed 199 -suckling 2 354 
Wind 3.47 2 234 Wormwood 3 523-7 -tops 3 380-98, 393 
Wind-berry 2 697 | Witch Havel 235 Annual 526 | Yellow-weed (any 
Wind-flower hed Gea Mics Beach 527 Golden Rod) 2 112, 
102, 100, 158, 122 Witches’-bell 7,49 Biennial 526 200; 3 380-98, 386, 
Red 98 ee Bees Canada 524 395 
Wind-root 3 25 3 295,557 Common 525] Yellow-wood 1 632; 
-rose 2 137 coon 2 87 Hungarian 527 2 343, 485, 721 
Windles 3 246 -milk 612 Linear-leaved 524 American 2 343 
Windlestraw 1 213,218| -money-bags 207} Northern 3.523] Kentucky 264 
Wine-berry -pouches 158 Roman 2 144; 3 527 Prickly rae 
2 237, 241, 277, 697 -thimbles 3 204, 557 Sea 3 341,524 West Indian 444 
-tree 287 | Withe-rod 3 271-2 Silky 524 | Yellow-wort, Parsley- 
Wing-seed 445 Naked 273 Silvery 525 leaved 2 80 
-stem 3 487 | Withe-wood 271, 273 Tall 523: | -Valines oe < 
Wink-a-peep 2 716 | With-wind 47 Wild 529-4 be ace, 3 
Winterberry 487-9 | Withy-tv" I 597 | Wormwood-sage 3 525 VE . Fa. saa 3 af 
Winter-bloom 235 | Woad, W 2 200 Wound-rocket 2 177 Yew ae 3 4 
Winter-brake I 33 | Woad-waxca 350 | Woundwort 3 124-7 ew, oe . rs 
Winter-cherry Wobsqua-grass I 141 Clown’s 125 Dwar 7 
501; 3 162} Wode-whistle 2 653 Corn or Field 127 European 67 
-clover 3 255 | Wolfber - 3 277 Downy 12 _ Oregon 67 
-cress 2 177 | Wolffia I 449 Soldiers 515 | Yorkshire-fog 214 
-fat 20 | Wolffiella 1 449 | Wrack I gl -sanicle 3 226 
Winter-grass I 409 | Wolf-foot 3 145-6 | Wren’s-flower 2 426] Youth-wort 2 203 
-Gilliflower 2 175 | Wolf-peach 167 | Wulfenia 3 198-9 
WINTERGREEN FAMILY | Wolf’s-bane 2 88, 96-7 | Wymote 2 514 | Zannichellia 1 88-9 
667 Trailing 07 Zeus’-wheat 2 720 
Wintergreen 668-72 | Wolf’s-claws I 47 | Yam FamIty I 535 | Zinnia, Prairie 3 466 
Bitter 672 | Wolf’s-milk 2 473 | Yam-root, Wild 535 | Zizaniopsis I 167 
Bog 669 | Woman’s-tobacco 451 | Yankapin 2 77 | Zizia, Bebb’s 2 641 
Chickweed 715 | Wood Betony 3 128,146 | Yarr 259 | Zornia 394 
Creeping 693 | Woodbind 46 | Yarrow 3 575-6 | Zygadenus I 491 


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