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Full text of "A list of the fishes of Pennsylvania"








FOR THE PEOPLE 

FOR EDVCATION 

FOR SCIENCE 






LIBRARY 

OF 

THE AMERICAN MUSEUM 

OF 

NATURAL HISTORY 





ON PERMANENT LOAN | 

TO THE DEPARTMENT i 

OF ICHTHYOLOGY I 

AMERICAN MUSEUM 1 

OF NATURAL HISTORY | 



Vol. 32, pp. 49-74 May 20, 1919 

PROCEEDINGS 

OF THE . 

BIOLOGICAL SOCIETY OF WASHINGTON ^'^^O^i^' 

A LIST OF THE FISHES OF PENNSYLVANIA. 6 

BY HENRY W. FOWLER. i 



The present paper is a summary of the data accumulated 
during the past twenty years, presented in condensed form, so 
that the distribution of each species may be traced so far as 
present details permit. For this reason they are arranged ac- 
cording to the various hydrographic basins with only the 
counties mentioned, additional records, where noteworthy, being 
supplied in parentheses. The work is therefore intended as a 
slight contribution to the distribution of our local fishes. 

Like many departments of natural history the founding of 
the binomial system by Carl von Linne in 1758 first establishes 
several fishes from Philadelphia. Alexander Wilson contributes 
the first notice of shad and alewife in the article on ichthyology 
in Ree's Encyclopaedia, to which he secured an assistant editor- 
ship in 1806. His article was published about 1812. Charles 
Alexandre Le Sueur is the first to carefully study the fishes of 
this State, much of his material doubtless having been secured 
near Philadelphia. He is credited with sixteen of our species, 
while eight other names he proposes are synonyms. Con- 
stantine Samuel Rafinesque described many of our species in 
his Ichtlyologia Ohiensis. The localities given are seldom 
definite and usually would apply to the entire Ohio basin. He 
has described twenty-three of our species, besides fourteen 
synonyms. 

Several of our species are also described by Achille Valen- 
ciennes, in colaboration with Baron Cuvier, in the great Histoire 
Naturelle des Poissons. Samuel Stehman Haldeman studied the 
fauna of the lower Susquehanna, though his contributions to 
ichthyology are rather incidental. He was signally unfortunate 

1 Published by permission of the Academy of Natural Sciences of Philadelphia. 
14— Phoc. Biol. Soc. Wash., Vol. 32, 1919, (49) 



50 Proceedings of the Biological Society of Washington. 

in that the four species he described were all anticipated. 
Charles Girard was a frequent contributor to North American 
ichthyology, and though he described two species from our 
limits both are synonyms. Spencer Fullerton Baird published 
a few notes on Pennsylvania fishes and the only one he described 
added another synonym to our well known fall-fish. 

The serious study of Pennsylvania fishes, however, begins 
with Edward Drinker Cope in his elaborate memoir the "Synop- 
sis of the Cyprinidae of Pennsylvania. "^ Later he attempted 
"The Fishes of Pennsylvania, "^ intended as a partly popular 
descriptive catalogue, apparently modeled from Jordan's 
Manual of Vertebrates. This work, likely valuable at the time 
of its publication, is chiefly useful for the notes on habits, etc. 
Cope described fourteen valid species and five synonyms. In 
this connection mention should be made of Jacob Stauffer, of 
Lancaster, who discovered two interesting fishes in his region, 
which were described by Cope. Stauffer's only contribution 
appears as his list of the fishes of Lancaster County.' This 
work is often faulty in the obscurity of determinations of many 
species. For instance, our common white catfish is described no 
less than three times. Tarleton Hoffman Bean gives a descrip- 
tive account^ somewhat like Cope's. It is similarly marred by 
the inclusion of a number of species native to regions adjacent 
to Pennsylvania, but never actually taken in the State limits. 

Among living writers Evermann and Bollman^ publish an 
important account of Monongahela River fishes in 1885. « In 
1902 Ross gives an account of twenty-five Center County 
fishes,^ largely of an economic nature. His Minytrema melanops, 
Leuciscus margarita and Hybopsis dissimilis are important addi- 
tions, and listed without comment or details of satisfactory com- 
parison they may be dropped. 

Below follows a list of my own papers: 

iTr. Amer. Philos. Soc, XIII, 1869, pp. 35-399, PI. 10-13. 

2Rep. State. Comm. Fisher., 1879-80 (1881), pp. 59-145, figs. 1-44. 

3 In Mombert's Hist. Lancaster County, Pa. 1868. 

4The Fishes of Pennsylvania. Rep. State Comm. Fisher., 1889-91 (1892), pp. 1-149, 
PIb. 1-35. 

SBoUman died of fever contracted in Georgia swamps, 1889. 

6Ann. N. Y. Acad. Sci., Ill, 1883-5 (1886), pp. 335-340. 

^Studies of Fishes of Central Pennsylvania. Rep. State Comm. Fisher., 1902, pp. 111- 
124. 



Fowler — A List of the Fishes of Pennsylvania. 51 

1905. Exoglossum in the Delaware. < Science, XXI, June 30, 1905, p. 
994. 

1906. Notes on Pennsylvania Fishes. < Amer. Nat., XL, 1906, pp. 
595-6. 

Pimephales in the Susquehanna. < L. c, p. 743. 

Fishes in Winter. < Forest and Stream, LXIX, December 28, 

1906, p. 1022. 

1907. Records of Pennsylvania Fishes. < Amer. Nat., XLI, January, 

1907, pp. 5-21, fig. 1. 

1908. Fishes of Bristol. < Bucks Coimty Gazette, December 25, 1908. 
(Popular newspaper account, with list.) 

1909. A Synopsis of the Cyprinidse of Pennsylvania. < Proc. Acad. Nat. 
Sci. Phila., 1908 (Feb. 2, 1909), pp. 517-553, PI. 27. 

1910. Note on some Pennsylvania Fishes. < Science, XXXI, March 4, 
1910, pp. 345-6. (List of some Indiana County species.) 

Note on the distribution of some Pennsylvania Fishes. < L. c.,,; 

XXXII, December 23, 1910, pp. 926-7. (Few species from Altoona and 
Easton.) 

1912. Records of Fishes for the Middle Atlantic States and Virginia. < 
Proc. Acad. Nat. Sci. Phila., 1912, pp. 34-59, 2 figs. (Pennsylvania, 
pp. 41-50.) 

Some Features of Ornamentation in Fresh-water Fishes. < Amer. 

Nat., XLVI, 1912, pp. 470-6, figs. 1-21. (On spawning cyprinoids 
and catostomoids.) 

Hermaphrodite Shad in the Delaware. < Science, XXXVI, July 5, 

1912, pp. 18-19. 

1914. Some Local Fish-eating Birds. < Cassinia, 1913 (March, 1914), 
pp. 6-16. 

Fishes in Polluted Waters. < Copeia, April 15, 1914, No. 5. (List 

of species in the Schuylkill tidal.) 

Fishes of the Poquessing Creek. < Aquarium Notes and News, I, 

No. 4, April, 1914, pp. 35-6. 

Description of a new Blenny from New Jersey, with notes on other 

fishes from the Middle Atlantic States. < Proc. Acad. Nat. Sci. Phila., 
1914 (June 24), pp. 342-358, 1 fig. (Pennsylvania, pp. 346-353.) 

Hadropterus peltatus in the Delaware. < Science, XL, December 

25, 1914, pp. 939-940. 

1915. An Annotated List of the Cold-blooded Vertebrates of Delaware 
County, Pennsylvania. < Proc. Del. Co. Inst. Science, VII, No. 2, 
June, 1915, pp. 1-13. (Fishes, pp. 1-8.) 

Fishes of the Neshaminy Creek Tidal Region. < Aquar. Notes and 

News, II, No. 6, June, 1915, pp. 49-51. 

1916. The River Fishes of Upper Philadelphia. < L. c. III, No. 4, April 
1916, pp. 28-30. 

The Fishes of Pennypack Creek in Upper Philadelphia. < L. c, 

III, No. 8, October 1916, pp. 60-64. 

1917. Fishes found in Bucks County, Pennsylvania, during 1915 and 1916. 
< L. c, IV, No, 2, February, 1917, pp. 11-15, 



52 Proceedings of the Biological Society of Washington. 

Notes on the Spawning-habits of our Lampreys. < L. c, IV, No. 

4, April, 1917, pp. 28-32. 

Notes on Fishes from New Jersey, Pennsylvania and Maryland. < 

Proc. Acad. Nat. Sci. Phila., 1917 (April 24), pp. 108-126. (Pennsyl- 
vania, pp. 115-122.) 

Fish Life of Mill Creek, a Delaware tributary. < Bristol Daily 

Courier, X, April 23, 1917. (Popular newspaper article.) 

Spawning Habits of Local Fishes, No. 1. < Aquar. Notes and News, 

IV, No. 5, May, 1917, pp. 37-40, 1 fig. 

Spawning Habits of Local Fishes. No. 2. < L. c, IV, No. 7, 

September, 1917, pp. 52-56. 

1918. Fishes from the Middle Atlantic States and Virginia. < Occas. 
Papers Mus. Zool. Univ. Michigan, No. 56, May 6, 1918, pp. 1-19, Pis. 
1-2. (Pennsylvania, pp. 8-14.) 

The Fishes of Perry County, Pennsylvania. < Copeia, November 

♦ 21, 1918, No. 63, pp. 89-91. 

PETROMYZONID^. 
Petromyzon marinus Linnseus. Lamprey. 
In the Delaware basin (Douglasville) Berks, (Neshaminy Falls and 
George School) Bucks, (Stock Grange, French Creek?) Chester, North- 
ampton, Philadelphia, Pike and Wayne Coimties. 

Susquehanna basin (Athens and Sayre) Bradford, (Emporium) Cameron, 
(Harrisburg) Dauphin, (Marietta, McCall's Ferry, Peach Bottom, Pequea, 
Safe Harbor) Lancaster, (Newton Hamilton) Mifflin, (MarysvUle) Perry 
and (Peach Bottom, York Furnace) York Counties. 

Ichthyomyzon concolor (Kirtland). Silvery Lamprey. 
(Conemaugh River formerly and Two Lick Creek) Indiana and McKean 
Counties. Lake Erie? 

Entosphenus aepypterus (Abbott). Brook Lamprey. 
Delaware basin in (Red Clay Creek near Kennett Square and White 
Clay Creek near Leonard) Chester County. 
Elk basin, (Elk View?) Chester County. 
Susquehanna basin in Cameron County. 

Ohio basin (Pittsburg) Allegheny, Indiana and (Seven Bridges) Potter 
Coimties. 

ACIPENSERID^. 

Acipenser sturio Linnseus. Sturgeon. 

Delaware basin in Bucks, Delaware, Northampton and Philadelphia 

Counties, now rare Susquehanna basin (reported at Coxtown in 1844*) 

Dauphin and (Fite's Eddy, Marietta, McCall's Ferry, Peach Bottom and 

Safe Harbor) Lancaster Counties. 

Acipenser fulvescens Rafinesque. Lake Sturgeon. 
Ohio basin (Pittsburgh formerly) Allegheny, (Foxburg and mouth of 

iForest and Stream, XXXIV, March 20, 1890, p. 171. 



Fowler — A List of the Fishes of Pennsylvania. 53 

Clarion River) Clarion, (Conemaugh and Kiskiminitas Rivers formerly) 
Indiana and (Tedionte and Warren) Warren Counties. 
Erie basin, (Massassauga Point) Erie County. 

Acipenser brevirostrum Le Sueur. Short-nosed Sturgeon. 
Delaware basin in Bucks and Philadelphia Counties. Le Sueur's type' 
is stUl in the Academy and agrees with several examples I examined in the 
field. 

Scaphirhynchus platorynchus (Rafinesque). Shovel-nosed Sturgeon. 
Though the late Dr. Bean mentions^ that "in the large tributaries of the 
Ohio, in western Pennsylvania, the species is very common," the original 
account by Rafinesque^ contains our only definite record. 

POLYODONTID^. 
Polyodon spathula (Walbaum). Paddle Fish. 
Ohio basin in Allegh6ny, (Foxburg and Clarion River formerly) Clarion, 
(Conemaugh River formerly at Blairsville) Indiana, McKean and (Tedionte 
and Warren) Warren Counties. 

LEPISOSTEID^. 
Lepisosteus osseus (Linnaeus). Long-nosed Gar-pike. 
Delaware basin in Bucks, (Chester) Delaware, Monroe and Philadelphia 
Counties. • 

Susquehanna basin (Marietta, Safe Harbor) Lancaster and (Peach 
Bottom) York Counties. 

Ohio basin (Foxburg) Clarion, (Conneaut Lake) Crawford, (Conemaugh 
River formerly) Indiana and Warren Counties. 
Erie basin in Erie County. 

Lepisosteus platostomus (Rafinesque). Short-nosed Gar-pike. 
Known from Rafinesque's original account* which says "as far as Pitts- 
burgh in the Allegheny River." Reported to me at Erie. 

AMIATID.E. 
Amiatus calvus (Linnaeus). Bowfin. 
A "dog fish" reported at Allegheny City^ is the only record I have for 
the Ohio basin, though I found it abundant at Erie. Stauffer mentions an 
example from the Susquehanna at Safe Harbor,^ likely an introduction? 

HIODONTIDiE. 
Hiodon alveoides (Rafinesque). Gold-eye, 
Known from Cope's Beaver and Youghiogheny River examples which 
I reported.'^ 

iProo. Acad. Nat. Sci. Phila., 1910, p. 604, PI. 38, fig. 1. 

2Rep. Com. Fisher. Pa., 1889-91 (1892), p. 5. 

3Ich. Ohien., 1820, p. 80. Seldom reaching as high as Pittsburgh in the Ohio River. 

4lch. Ohien., 1820, p. 72. 

6Forest and Stream, VIII, May 24, 1877, p. 243. 

6Amer. Nat., XIII, 1879, p. 525. 

7Proc. Acad. Nat. Sci. Phila., 1911, p. 204. 



54 Proceedings of the Biological Society of Washington. 

Hiodon tergisus Le Sueur. Moon-eye. 

Originally described from the Ohio at Pittsburgh,' though no recent 
records given. I found it at Erie. 

DOROSOMIDiE. 
Dorosoma cepedianum (Le Sueur). Mud-shad. 
Only recorded from the tidal of the Delaware basin, (Andalusia) Bucks, 
Chester, Delaware and (Bridesburg, Tacony) Philadelphia Counties. Le 
Sueur originally^ obtained it in the "market of Philadelphia." 

CLUPEIDiE. 
Pomolobus chrysochloris Rafinesque. Inland Alewife. 
Known from Rafinesque's original account,^ which gives it from the Ohio 
River and also says "it seldom goes as far as Pittsburgh." "Herring" re- 
ported formerly in (the Conemaugh River) Indiana County were likely this 
species. 

Pomolobus mediocris (Mitchill). Fall Herring. 
I have no local material though include this species provisionally, or 
only on reports of its occurrence in the Delaware River tidal of Bucks 
County at Tullytown. In late October, 1906, a school of about fifty large 
examples were reported at this point. When swimming at the surface they 
produced quite a disturbance, somewhat like a school of menhaden. About 
eight were taken on hooks baited with our common river killifishes {Fim- 
dulus), which were fastened to the hook by the upper jaw. When opened 
the herring were found more or less gorged with killifish. They took the 
bait with a rush and greatly interested the local anglers who called them 
"jacks" or "skipjacks." They remained until November. A large herring 
taken at the same place in December, 1912, was doubtless the same species. 

Pomolobus pseudoharengus (Wilson). Alewife. 

Delaware basin, (New Hope) Bucks, Delaware and Philadelphia Counties. 

Susquehanna basin, (Harrisburg) Dauphin, (reported at Clarke's Ferry 
dam in North Branch'*) Lackawanna, (Bainbridge, Fite's Eddy, Marietta, 
McCaU's Ferry, Pequea, Safe Harbor) Lancaster, (Marysville) Perry and 
(McCall's Ferry, Peach Bottom and York Furnace) York Counties. 

Pomolobus aestivalis (Mitchill). Summer Herring. 

Only found in the Delaware basin in Bucks, Delaware and Philadelphia 
Counties. 

lAs Hiodon tergisus Le Sueur, Journ. Acad. Nat. Sci. Phila., I, pt. 2, 1818, p. 366, and 
H. clodalus Le Sueur, 1. c, p. 367. Rafinesque next names it Hyodon vernalis in Ich. Ohien., 
1820, p. 43. 

2A8 Megalops cepediana in 1. c., p. 361. 

^Rafinesque, 1. c, p. 38. 

^Forest and Stream, LXVII. November 3, 1906, p. 701. 



Fowler — A List of the Fishes of Pennsylvania. 55 

Alosa sapidissima (Wilson). Shad. 

Delaware basin (Andalusia, Biles Creek, Dunk's Ferry, New Hope, 
Scott's Creek, Uhlertown, Yardley) Bucks, (Long Ford, Ming and mouth 
of French Creeks, Perkiomen, Phoenixville all formerly) Chester, Dela- 
ware, (Delaware Water Gap) Monroe, (Easton formerly) Northampton, 
(Manayunk, Port Providence, Valley Forge all formerly) Montgomery, 
Philadelphia, (Bushkill, Conashaugh, Delaware, Egypt Mills, Matamoras, 
Milford, Shohola) Pike and (Damascus, Hancock, Millanville) Wayne 
Counties. 

Susquehanna basin (Browntown, Sayre, Towanda) Bradford, (Beach 
Haven, Berwick, Bloomfield) Columbia, (Harrisburg) Dauphin, (Bain- 
bridge, Columbia, Creswell, Fite's Eddy, Marietta, Peach Bottom, Pequea, 
Safe Harbor, Washington Borough) Lancaster, (Hemlock Creek, Nanti- 
coke, Pittston, Shickhinny, Wilkes Barre all formerly) Luzerne, (Long 
Level, Northumberland) Northumberland, (Duncannon, Newport) Perry, 
(Keelersburg, Meshoppen, Tunckhannock) Wyoming and (East Prospect, 
Long Level, York Furnace) York Counties. 

Brevoortia tyrannus (Latrobe). Menhaden. 
Valenciennes early mentions it as abundant in the markets of New York 
and Philadelphia, and later Cope says it ascends the Delaware for a short 
distance in our limits. I have it from the New Jersey shore at Washington 
Park in August, 1908, and since then small examples have been found in 
Bucks (far as Dunk's Ferry) and Philadelphia Counties. 

SALMONIDiE. 
Coregonus clupeaformis (Mitchill). Whitefish. 
Abundant at Erie. 

Leucichthys sisco (Jordan). Sisco. 
Known from Erie examples reported by Jordan and Evermann.^ 

Leucichthys artedi (Le Sueur). Lake Erie Herring. 
Abundant at Erie. 

Cristivomer namaycush (Walbaum). Lake Trout. 
Erie County. Introduced in the Delaware and Susquehanna basins. 

Salvelinus fontinalis (Mitchill!) . Brook Trout. 

Delaware basin in Berks, Bucks, Carbon, Chester, Lackawanna, Leba- 
non, Lehigh, Montgomery, Monroe, Northampton, Philadelphia, Pike, 
Schuylkill and Wayne Counties. 

Susquehanna basin in Berks, Blair, Bradford, Cameron, Center, Clear- 
field, Clinton, Columbia, Cumberland, Dauphin, Elk, Fulton, Huntingdon, 
Juniata, Lackawanna, Lancaster, Lebanon, Luzerne, Lycoming, Mifflin, 
Perry, Potter, Schuylkill, Somerset, SulUvan, Susquehanna, Wayne, Wyo- 
ming and York Counties. 

iBuU. Bur. Fisher., XXIX, 1909, p. 12, fig. 6 (type) and PI. 2, as L. sisco huronius. 



56 Proceedings of the Biological Society of Washington. 

Potomac basin in Adams, Franklin, Fulton and Somerset Counties. 
Genesee basin in Potter County. 

Ohio basin in Erie, Fayette, Forrest, Indiana, McKean, Lawrence, 
Potter, Somerset, Warren and Westmoreland Counties. 
Erie basin in Erie County. 

ARGENTINIID^. 
Osmerus mordax (Mitchill). Smelt. 
Delaware tidal in Bucks, Delaware and Philadelphia Counties. Besides 
the types of 0. sergenti Norris' many examples in the markets examined, 
but the latter largely imported. 

ICTALURID^. 

Ictalurus furcatus (Valenciennes). Chuckle-head Cat. 

Reported from the Monongahela River by Evermann and Bollman.^ 

Ictalurus punctatus (Rafinesque). Spotted Cat. 

Ohio basin in Allegheny, Beaver, Clarion, (Blairsville formerly) Indiana 
and Westmoreland Counties. 

Erie basin in Erie County. Possibly the alleged A. nigricans (Le Sueur) 
is not distinct from the present species. The late Dr. Bean says "Erie re- 
ceives its supply of catfish from fishermen who operate in the lake from 
Erie to Elk Creek, with set-lines during the summer months." A number 
of catfish in the markets and fisheries at Erie were examined and all of the 
fork-tailed forms were the spotted cat. All had the predorsal bony bridge 
complete and but few were without spots. 

Ameiurus catus (Linnseus). White Cat. 
Delaware basin in Bucks, (Brandywine Creek) Chester, Delaware and 
(League Island) Philadelphia Counties. 
Susqueharma basin in Lancaster and (Peach Bottom) York Counties. 

Ameiurus natalis (Le Sueur). Yellow Cat. 
Only met with by me at Erie. Not found east of the Alleghanies in our 
limits, though in the Ohio basin early recorded as far as Pittsburgh. ^ 

Ameiurus nebulosus (Le Sueur). Common Cat. 

Delaware basin in Berks, Bucks, Carbon, Chester, Delaware, Lebanon, 
Monroe, Montgomery, Northampton, Pike and Wayne Counties. 

Susquehanna basin in Bedford, Berks, Blair, Bradford, Cambria, Center, 
Chester, Clearfield, Clinton, Cumberland, Fulton, Juniata, Lackawanna, 
Lancaster, Lebanon, Luzerne, Lycoming, Mifflin, Montour, Northumber- 
land, Perry, Snyder, Somerset, Sullivan, Susquehanna, Tioga, Wayne, 
Wyoming and York Counties. 

iProc. Acad. Nat. Sci. Phila., 1868, p. 93. Schuylkill below Fairmount dam. Brandy- 
wine Creek. 

iAnn. N. Y. Acad. Sci., Ill, 1883-5 (1886), p. 336. 
iPimelodus cupreus Rafinesque, Ich. Ohien., 1820, p. 65. 



Fowler — A List of the Fishes of Pennsylvania. 57 

Northeast River basin in Chester County. 

Potomac basin in Franklin, Fulton and Somerset Counties. 

Genesee basin in Potter County. 

Ohio basin in Armstrong, Beaver, Cambria, Crawford, Indiana, Jeffer- 
son, Lawrence, McKean, Somerset, Venango, Warren and Westmoreland 
Counties. 

Erie basin in Erie County. 

Ameiurus melas (Rafinesque). Black Cat. 
Abundant and frequently marketed at Erie. 

Ameiurus nigrilabris (Cope). Blind Cat. 
Only known from the types,' procured in the Conestoga Creek, tributary 
to the Susquehanna. Originally placed in a separate genus on account of 
the rudimentary eyes, though otherwise very close to A. nebulosus. 

Leptops olivaris (Rafinesque). Mud Cat. 
Known from Cope's material I reported from the Youghiogheny River.^ 

Noturus flavus Rafinesque. Stone Cat. 
Reported from Pigeon Creek by Evermann and Bollman* and I have 
it from the Youghiogheny River and Indiana County. 

Schilbeodes gyrinus (Mitchill). Tadpole Cat. 
Many examples from the Delaware basin in Bucks. Lehigh and Phila- 
delphia Counties, and the Genesee basin in Potter County. 

Schilbeodes insignis (Richardson). Margined Cat. 

Delaware basin in Bucks, (Jordan Creek near Helfrich's Spring) Lehigh, 
Montgomery, Monroe, Northampton, Philadelphia, Pike and Wayne 
Counties. 

Elk basin in Chester County. 

Susqueharma basin in Blair, Bradford, Cameron, Center, Cumberland, 
(Columbia, Fite's Eddy, Marietta) Lancaster, Sullivan and( Peach Bottom, 
Sowego Creek) York Counties. 

Schilbeodes miurus (Jordan). Brindled Cat. 
Recorded by McConnell* from Indiana County. I have been unable 
to examine the material on which this record is based. 

CYPRINID^. 
Campostoma anomalum (Rafinesque). Stone Roller. 
Susquehanna basin in Columbia County. 

Ohio basin in Allegheny, Beaver, Indiana, Lawrence and McKean 
Counties. 

^Gronias nigrilabris Cope, Proc. Acad. Nat. Sci. Phila., XVI, 1864, p. 231. 

2Amer. Nat., XLI, 1907, p. 14. 

3 Ann. N. Y. Acad. Sci., Ill, 1883-5 (1886), p. 335. 

4Rep. Com. Fisher. Pa., 1905, p. 179. 



58 Proceedings of the Biological Society of Washington. 

Chrosoitius erythrogaster (Rafinesque). Red-bellied Dace. 
Only known from Cope's material obtained in the Kiskiminitas River, 
which I reported recently. ^ 

Chrosomus erythrogaster eos (Cope). Eastern Red-bellied Dace. 
Known only from Cope's types,^ obtained in Meshoppen Creek, Susque- 
hanna basin, in September, 1861, and of them but three now extant. 

Hybognathus nuchalis Agassiz. Silvery Minnow. 
Reported from the Ohio basin by Cope,^ though I have no material. 

Hybognathus nuchalis regius (Girard). Eastern Silvery Minnow. 
Only in the Delaware tidal in Bucks, Delaware and Philadelphia Counties. 

Hybognathus nuchalis argyritis (Girard). Blunt-jawed Silvery Minnow. 
Known from Cope's material from the Kiskiminitas I recently noticed.* 

Pimephales promelas Rafinesque. Black-headed Minnow. 
I met with it in the Ohio basin of McKean County. 

Pimephales notatus (Rafinesque). Blunt-nosed Minnow. 

Delaware basin in Bucks, Lehigh and Montgomery Counties. 

Susquehanna basin in Berks, Blair, Lebanon, Dauphin, Snyder and York 
Counties. 

Ohio basin in Allegheny, Clarion, Indiana, McKean and Westmoreland 
Counties. 

Semotilus bullaris (Rafinesque). Fall Fish. 

Delaware basin in Berks, Bucks, Chester, Delaware, Lehigh, Monroe, 
Montgomery, Northampton, Philadelphia, Pike and Wayne Counties. 

Susquehanna basin in Berks, Bradford, Cambria, Chester, Clinton, 
Cumberland, Huntingdon, Lancaster, Lycoming, Mifflin, Montour, 
Potter, Somerset, Wayne and York Counties. 

Potomac basin in Franklin and Somerset Counties. 

Semotilus atromaculatus (Mitchill). Creek Chub. 

Delaware basin in Berks, Bucks, Chester, Delaware, Lehigh, Monroe, 
Montgomery, Northampton, Philadelphia, Pike and Wayne Coimties. 

North East basin in Chester County. 

Potomac basin in Franklin and Somerset Counties. 

Susquehanna basin in Berks, Blair, Cameron, Chester, Clinton, Colum- 
bia, Cumberland, Dauphin, Huntingdon, Lackawanna, Lancaster, Lebanon, 

iProc. Acad. Nat. Sci. Phila., 1908 (1909), p. .519. 
2C. eos Cope, Proc. Acad. Nat. Sci. Phila., 1861, p. 523. 
3Rep. Com. Fisher. Pa., 1879-80 (1881). p. 91. 
4Proo. Acad. Nat. Sci. Phila., 1908 (1909), p. 521. 
5L. c, p. 522. Port Allegany. 



Fowler — A List of the Fishes of Pennsylvania. 59 

Lycoming, Mifflin, Montour, Perry, Potter, Snyder, Somerset, Sullivan, 
Wayne and York Counties. 

Genesee basin in Potter County. 

Ohio basin in Allegheny, Armstrong, Cambria, Clarion, Fayette, Indiana, 
Jefferson, McKean, Potter, Somerset, Warren and Westmoreland Counties. 

Leuciscus margarita (Cope). Pearl Dace. 
One of Cope's six examples, originally obtained in the Conestoga near 
Lancaster,' remains in the Academy. Ross records this species from Center 
County,^ though I have been unable to examine his material. 

Leuciscus carletoni Kendall. Carleton's Dace. 
A single example I obtained in high color July 23, 1899, in McKean 
County at Cole Grove, is interesting as a new addition to the fauna. ^ It 
was captured in a small mountain-stream, the surface of which was largely 
covered with petroleum, so that the fish was not discovered until taken 
out of the water in a net. Associated were a number of other small fishes 
as creek chubs, silver-fins and cut-lips. Though I wrongly identified the 
present example as the pearl dace, as indicated below, it agrees in every 
way with the present species. 

Leuciscus vandoisulus Valenciennes. Rosy-sided Dace. 
Delaware, Elk and North East basins in Chester County. 
Susquehanna basin in Cameron, Chester and Lancaster Counties. 

Leuciscus elongatus (Kirtland). Slender Dace. 
Ohio basin in Crawford, Indiana and McKean Counties. 

Abramis crysoleucas (Mitchill). Roach. 

Delaware basin in Berks, Bucks, Chester, Delaware, Lebanon, Lehigh, 
Monroe, Montgomery, Northampton, Philadelphia and Pike Counties. 

Susquehanna basin in Cameron, Lackawanna, Lancaster, Mifflin, Sulli- 
van and York Counties. 

Ohio basin in Clarion and Elk Counties. 

Erie basin in Erie County. 

Ceratichthys vigilax Baird and Girard. Cliola Minnow. 
Recorded from the Monongahela by Evermann and BoUman.* 

Notropis bifrenatus (Cope). Bridled Minnow. 
Delaware basin in Berks, Bucks, Chester, Delaware, Lehigh, Mont- 
gomery, Northampton and Philadelphia Counties. 
Susquehanna basin in York and Lancaster Counties. 

IClinostomus margarita Cope, Tr. Amer. Philos. Soc, (2) XIII, 1866, p. 377, fig. (teeth), 
PI. 13, fig. 1. 

2Rep. Com. Fisher. Pa., 1902, p. 114. 

3L. margarita (non Cope) Fowler, Science, XXXI, March 4, 1910, p. 346. 

4 Ann. N. Y. Acad. Sci., Ill, 1883-5 (1886), p. 336. 



60 Proceedings of the Biological Society of Washington. 

Notropis deliciosus (Girard). Straw-colored Minnow. 
Recorded from the Monongahela by Evermann and Bollman/ and I 
have it from Indiana County. 

Notropis procne (Cope). Swallow Minnow. 

Delaware basin in Berks, Bucks, Chester, Delaware, Montgomery and 
Philadelphia Counties. 

Elk and North East basins in Chester County. 

Susquehanna basin in Berks, Columbia, Dauphin, Lancaster, Lebanon, 
Snyder and York Counties. 

Notropis boops Gilbert. Big-eyed Minnow. 
I obtained it at Foxburg in Clarion County.* 

Notropis keimi Fowler. Allegheny Minnow. 
Only the types obtained in the Allegheny basin.* 

Notropis hudsonius (CHnton). Spawn-eater. 
Ohio basin in the Monongahela River and McKean County. Erie basin 
in Erie County. 

Notropis hudsonius amarus (Girard). Eastern Spawn-eater. 

Delaware basin in Bucks, Chester, Delaware, Lehigh, Montgomery, 
Philadelphia and Pike Counties. 

Susquehanna basin in Center, Columbia, Dauphin, Lancaster, Lebanon, 
Snyder and York Counties. 

Notropis whipplii (Girard). Silver-fin. 
Ohio basin in the Kiskiminitas and Youghiogheny Rivers, and McKean 
County. Erie basin in Erie County. 

Notropis whipplii analostanus (Girard). Eastern Silver-fin. 

Delaware basin, Berks, Bucks, Carbon, Chester, Delaware, Lehigh, 
Montgomery, Philadelphia and Pike Counties. 

Elk basin in Chester County. 

Susquehanna basin in Berks, Blair, Columbia, Dauphin, Huntingdon, 
Lancaster, Lebanon, Mifflin, Perry, Snyder, Susquehanna, Wyoming and 
York Counties. 

Notropis cornutus (Mitchill). Red-fin. 

Delaware Basin in Berks, Carbon, Chester, Delaware, Lehigh, Monroe, 
Montgomery, Northampton, Philadelphia and Pike Counties. 

Elk basin in Chester County. 

Susquehanna basin in Bedford, Berks, Cameron, Chester, Columbia, 
Cumberland, Center, Dauphin, Lancaster, Lebanon, Luzerne, Lycoming, 
Perry, Sullivan, Snyder, Susquehanna, Wyoming and York Counties. 

iL. 0. 

sScience, XXXI, March 4, 1910. p. 338. 

3Proc. Acad. Nat. Soi. Phila., 1908 (1909), p. 533, PI. 27. 



Fowler — A List of the Fishes of Pennsylvania. 61 

Genesee basin in Potter County. 

Ohio basin in Clarion, Indiana, Lawrence, McKean and Warren Counties. 

Erie basin in Erie County. 

Notropis chalybaeus (Cope). Iron-colored Minnow. 
Originally described from the Schuylkill near Conshohocken' and I 
obtained it in the Delaware basin in Bucks, Montgomery and Northampton 
Counties. 

Notropis jejunus (Jordan). Hungry Minnow. 
Reported from the Monongahela by Evermann and BoUman.^ 

Notropis atherinoides Rafinesque. Emerald Minnow. 
The Monongahela by Evermann and BoUman, and I have it from the 
Beaver River, Indiana and Erie Counties. 

Notropis photogenis (Cope). Arrow Minnow. 
Two of the types from the Youghiogheny River^ and an example from 
Indiana County examined. 

Notropis photogenis amoenus (Abbott). Attractive Minnow. 

Delaware basin in Bucks, Monroe, Montgomery and Philadelphia 
Counties. 

Susquehanna basin in Center, Columbia, Dauphin, Lancaster, Lebanon, 
Perry, Snyder and York Counties. 

Notropis dilectus rubrifrons (Cope). Red-faced Minnow. 
The types^ and Indiana County examples. 

Ericymba buccata Cope. Scalloped Minnow. 
The types^ and Indiana County material examined. Both this and the 
preceding recorded from the Monongahela by Evermann and Bollman. 

Rhinichthys cataractae (Valenciennes). Long-nosed Dace. 

Delaware basin in Bucks, Carbon, Chester, (Hosensack) Lehigh and 
Northampton Counties. 

Susquehanna basin in Blair, Cameron, Center, Columbia, Cumberland, 
Dauphin, Lancaster, Lebanon, Luzerne, Perry and Snyder Counties. 

Elk basin in Chester County. 

Ohio basin in Fayette and Indiana Counties. 

Rhinichthys atronasus (Mitchill). Black-nosed Dace. 
Delaware basin in Bucks, Carbon, Chester, Delaware. Lehigh, Monroe, 
Montgomery, Northampton, Philadelphia and Pike Counties. 
Elk and North East basins in Chester County. 

iHybopsis chalybcBus Cope, Trans. Amer. Philos. Soc, (2), XIII, 1869, p. 383. 

2Ann. N. Y. Acad. Sci., Ill, 1883-5 (1886), p. 338. 

3Squaluis photogenis Cope, Proc. Acad. Nat. Sci. Phila., 1864, p. 280. 

iAlbtirnus rubrifrons Cope, 1. c, 1865, p. 85. 

8L. c, p. 88. 



62 Proceedings of the Biological Society of Washington. 

Susquehanna basin in Berks, Blair, Cameron, Columbia, Chester, Center, 
Cumberland, Dauphin, Huntingdon, Lackawanna, Lancaster, Lebanon, 
Luzerne, Mifflin, Perry, Potter, Snyder, Sullivan and York Counties. 

Potomac basin in Fulton County. 

Genesee basin in Potter County. 

Ohio basin in Beaver, Fayette, Indiana, McKean, Potter, Somerset, 
Warren and Westmoreland Counties. 

Rhinichthys bowersi Goldsborough and Clark,' from the Ohio basin of 
West Virginia appears only a local variety. A number of specimens from 
many of the above listed districts show even greater ranges of variation, 
with many details in extremes of other characters in combination. The 
fact of the lesser row of teeth, for instance, being absent or consisting only 
of a single tooth is a very variable condition. 

Hybopsis dissimilis (Kirtland). Spotted Chub. 
Cope's material from the Monongahela and Youghiogheny Rivers ex- 
amined. 

Hybopsis storerianus (Kirtland). Storer's Chub. 
Evermann and Bollman report it from the Monongahela.^ 

Hybopsis kentuckiensis (Rafinesque). Horned Chub. 

Elk basin in Chester County. 

Susquehanna basin in Berks, Cameron, Center, Dauphin, Lancaster, 
Lebanon, Perry, Snyder, Susquehanna and York Counties. 

Ohio basin in Beaver, Fayette, Indiana, Lawrence, McKean, Warren 
and Westmoreland Counties. 

No definite record has been given for the Delaware, where it likely does 
not occur. Possibly H. dissimilis Ross is the present species. 

Exoglossum maxillingua (Le Sueur). Cut-lips. 

Delaware basin in Berks, Bucks, Chester, Delaware and Montgomery 
Counties. 

Elk basin in Chester County. 

Susquehanna basin in Berks, Blair, Cameron, Center, Columbia, Cum- 
berland, Dauphin, Juniata, Lancaster, Lebanon, Luzerne, Snyder, Sullivan 
and York Counties. 

Ohio basin in McKean County. 

CATOSTOMID.E. 

Cycleptus elongatus (Le Sueur). Black-horse. 

Mentioned by Rafinesque in the Oliio as far as Pittsburgh, ^ and Cope 

reports it from the Allegheny. An example obtained by Cope from the 

Kiskiminitas River is doubtless partly the basis of his record and the only 

one from our limits I examined. 

iBull. Bur. Fisher., XXVII, 1907 (March 14, 1908), p. 36, fig. 
2Ann. N. Y. Acad. Sci., Ill, 1883-5 (1886), p. 339. 
sichth. Ohien., 1820, p. 60. 



Fowler — A List of the Fishes of Pennsylvania. 63 

Amblodon bubalus Rafinesque. Small-mouth Buffalo. 
Reported by Rafinesque, who says it comes as far as Pittsburgh. ' No 
other records have been given, though doubtless it may have been found 
in the Allegheny, at least until recently. 

Carpiodes carpio Rafinesque. Carp Sucker. 
Known from two of Cope's examples I reported^ from the Beaver River. 

Carpiodes thompsoni Agassiz. Lake Carp Sucker. 
Found at Erie by the writer 1912-1914. 

Carpiodes cyprinus (Le Sueur). Eastern Carp Sucker. 
Found at several localities in the Susquehanna basin in Lancaster and 
York Counties. Not certainly known from the Delaware in our limits. 

Carpiodes difformis Cope. Snub-nose Carp Sucker. 
A small example obtained by Cope in the Youghiogheny River reported 
by the writer.^ 

Carpiodes cutisanserinus Cope. Rough-nose Carp Sucker. 
Only known from the type taken in the Kiskiminitas River.* 

Carpiodes velifer (Rafinesque). Quill-back. 
Besides Rafinesque's original record from the Ohio River to Pittsburgh.* 
Evermann and Bollman obtained it in the Monongahela and I have two 
examples obtained in the Youghiogheny by Cope. 

Catostomus catostomus (Forster). Long-nose Sucker. 
Jordan records it from the Youghiogheny River/ though this material 
I have not seen. 

Catostomus commersonnii (Lac^pede). Common Sucker. 

Delaware basin in Berks, Bucks, Carbon, Chester, Delaware, Lehigh, 
Monroe, Montgomery, Northampton, Philadelphia, Pike and Wayne 
Counties. 

North East basin in Chester County. 

Susquehanna basin in Bedford. Berks, Blair, Bradford, Cameron, Center, 
Chester, CUnton, Columbia, Cumberland, Dauphin, Fulton, Juniata, Lack- 
awanna, Lancaster, Lebanon, Luzerne, Lycoming, Mifflin, Perry, Snydery 
Susquehanna, Wyoming and York Counties. 

Potomac basin in Franklin and Fulton Counties. 

Genesee basin in Potter County. 

Ohio basin in Armstrong, Beaver, Cambria, Clarion, Fayette, Forest, 

iL. c, p. 55. 

2Proc. Acad. Nat. Sci. Phila., 1913, p. 46. 

aProc. Acad. Nat. Sci. Phila., 1913, p. 46. 

4Proc. Amer. Philos. Soc, XI, 1870, p. 481. 

5Ich. Ohien., 1820, p. 56. 

6Bu]l. U. S. Nat. Mus., No. 12, 1878, p. 175. 



64 Proceedings of the Biological Society of Washington. 

Indiana, McKean, Somerset, Venango, Warren and Westmoreland 
Counties. 

Erie basin in Erie County. 

Catostomus nigricans Le Sueur. Black Sucker. 

Delaware basin in Lehigh County, likely introduced? 

Elk basin in Chester County. 

Susquehanna basin in Berks, Blair, Cameron, Chester, Dauphin, Lan- 
caster, Lebanon, Mifflin, Perry, Snyder and York Counties. 

Ohio basin in Beaver, Clarion, Indiana, Lawrence, Warren and West- 
moreland Counties. 

Erimyzon sucetta oblongus (Mitchill). Chub Sucker. 

Delaware basin in Berks, Bucks, Chester, Delaware, Lehigh, Mont- 
gomery, Northampton and Philadelphia Counties. 

Susquehanna basin in Blair, Cameron, Center, Columbia, Cumberland, 
Dauphin, Lancaster, Mifflin, Sullivan and York Counties. 

Erie basin in Erie County. 

Moxostoma anisurum (Rafinesque). White-nose Red-horse. 

The original account by Rafinesque' mentions it from the Ohio and large 

streams far as Pittsburgh. Cope later describes it from the Youghiogheny 

River, 2 and these types, together with Beaver River material, I have 

examined. McConnell has reported it from Jamestown in Mercer County.^ 

Moxostoma aureolum Le Sueur. Red-horse. 
Ohio basin in Beaver, Indiana and Westmoreland Counties. 
Erie basin in Erie County. 

Moxostoma macrolepidotum (Le Sueur). Eastern Red-horse. 
Delaware basin in Philadelphia County, where rare. 
Susquehanna basin in Cumberland, Lancaster and York Counties. 

Moxostoma breviceps (Cope). Long-tailed Red-horse. 
The type from the Youghiogheny and Erie examples examined. 

Moxostoma duquesnii (Le Sueur). Fine-scaled Red-horse. 
Le Sueur originally had it from the Ohio at Pittsburgh.^ Cope obtained 
an example in the Youghiogheny and one in the Beaver River in 1880. 
These and a small example from Indiana County constitute my material. 

Placopharynx carinatus Cope. Big-toothed Sucker. 
Giinther records it from the Youghiogheny^ and I have Cope's Beaver 
River material. 

ilch. Ohien., 1820, p. 54. 

iPiychostomus velatus Cope, Proc. Amer. Philos. Soc, XI, 1870, p. 470. 

3Rep. Com. Fisher. Pa., 1905 p. 178. 

*Catostomus duquesnii Le Sueur, Journ. Acad. Nat. Sci. Phila., I, 1817, p. 105. 

6A8 C. duquesnii (non Le Sueur) Giinther, Cat. Fish. Brit. Mus., VII, 1868, p. 18. 



Fowler — A List of the Fishes of Pennsylvania. 65 

ANGUILLID^. 
Anguilla rostrata (Le Sueur). Eel. 

Delaware basin in Berks, Bucks, Carbon, Chester, Delaware, Lehigh, 
Monroe, Montgomery, Northampton, Philadelphia, Pike and Wayne 
Counties. 

Susquehanna basin in Bedford, Berks, Blair, Bradford, Cameron, Clear- 
field, Clinton, Huntingdon, Lackawanna, Lancaster, Luzerne, Lycoming, 
Mifflin, Montour, Potter, Perry, Sullivan, Susquehanna, Wyoming and 
York Counties. 

Potomac basin in Somerset County. 

Ohio basin in Allegheny, Cambria, Clarion, Indiana, Potter, Somerset 
and Warren Counties. 

Genesee basin in Potter County. 

Erie basin in Erie County. 

ESOCID^. 
Esox americanus (Gmelin). Banded Pickerel. 
Delaware basin in Berks, Bucks, Carbon, Chester, Delaware, Lehigh, 
Monroe, Montgomery, Northampton, Philadelphia and Pike Counties. 

Susquehanna basin in Berks, Blair, Chester, Columbia, Cumberland, 
Elk, Fulton, Huntingdon, Lancaster, Luzerne, Lycoming, Montour, North- 
umberland, Perry and York Counties. 

Esox vermiculatus Valencieimes. Western Pickerel. 
Ohio basin in Armstrong, Clarion, Crawford, Elk, Indiana, McKean, 
Mercer and Venango Counties. 

Esox tridecemlineatus Mitchill. Chain Pickerel. 

Delaware basin in Bucks, Carbon, Lehigh, Monroe, Philadelphia, Pike 
and Wayne Counties. 

Susquehanna basin in Bradford, Center, Clearfield, Clinton, Cumber- 
land, Lackawanna, Lebanon, Luzerne, Sullivan, Susquehanna, and Wyom- 
ing Counties. Introduced in Lake Erie. 

Esox lucius Linnaeus. Pike. 
Ohio basin in Clarion, Indiana, Mercer, McKean, Warren and West- 
moreland Counties. 

Erie basin in Erie County. 

Esox masquinongy Mitchill. Muskallunge. 
Ohio basin in Beaver, Clarion, Crawford and Warren Counties. 
Erie basin in Erie County. Introduced in the Delaware basin of Wayne 
County. 

UMBRID^. 
Umbra limi (Kirtland). Western Mud Minnow. 
Known from my examples obtained at Meadeville in Crawford County.* 

iProc. Acad. Nat. Soi. Phila., 1912, p. 47. 



66 Proceedings of the Biological Society of Washington. 

Umbra pygmaea (De Kay). Eastern Mud Minnow. 
Delaware basin in Bucks, Delaware and Philadelphia Counties. Shu- 
feldt recently describes U. pygmcBa bilineata as a new form, from the 
tributaries of Chesapeake Bay, which I have discussed elsewhere.' 

POECILIID^. 
Fundulus heteroclitus macrolepidotus (Walbaum). Mummichog. 
Delaware basin in Bucks. Delaware and Philadelphia Counties. 

Fundulus diaphanus (Le Sueur). Barred Killifish. 

Delaware basin in Bucks, Berks, Chester, Delaware, Lehigh, Mont- 
gomery, Northampton and Philadelphia Counties. 

Susquehanna basin in Berks, Center, Columbia, Dauphin, Lancaster, 
Lebanon, Snyder and York Counties. 

Fundulus diaphanus menona (Jordan and Copeland). Menona Killifish. 
My examples from the Allegheny River near Warren and at Erie. 

BELONIDiE. 
Strongylura marina (Walbaum) Green Gar. 
Delaware basin in Bucks, Delaware and Philadelphia Counties. 
Susquehanna basin in Lancaster and York Counties. 

Strongylura acus (Lac6p6de). Houndfish. 
A single example in the Academy obtained by E. D. Cope in the lower 
Susquehanna, within our limits, is an addition to the State's fauna. No 
exact locality is given. The specimen is 667 mm. long and still in good 
preservation. Comparison with Massachusetts and Italian material reveal 
all its characters in harmony. As the species enters our limits by way of 
Chesapeake Bay it may occur casually in any of the other large rivers 
tributary. I have, however, not certainly secured it in the Delaware. 

ATHERINID^. 
Labidesthes sicculus (Cope). Brook Silversides. 
My material from the Youghiogheny River and Erie. 

PERCOPSIDiE. 

Percopsis omiscomaycus (Walbaum). Trout Perch. 

I have this interesting fish from the Jordan Creek in Lehigh County, the 

only locality in the Delaware basin. ^ In the Ohio basin it is recorded from 

the Monongahela by Evermann and Bollman,^ and French Creek above 

Frenchtown, by McConnell.^ 

iCopeia, No. 50, Oct. 26, 1917, pp. 94-96. 
aProc. Acad. Nat. Sci. Phila., 1917, p. 119. 
3Ann. N. Y. Acad. Sci., Ill, 1883-5 (1886), p. 339. 
4Rep. Com. Fisher. Pa., 1905, p. 178. 



Fowler — A List of the Fishes of Pennsylvania. 67 

GASTEROSTEID.^. 
Eucalia inconstans (Kirtland). Brook Stickleback. 
I obtained it in the Allegheny at Foxburg. 

Gasterosteus aculeatus Linnaeus. Two-spined Stickleback. 
I have no local material, though it has been recorded from the Delaware 
at Philadelphia by Dr. C. C. Abbott.' It must be rare or extremely local 
in our limits, as I never met with it away from the coastal region, or far 
from salt water. To the contrary the late Dr. Bean says "in Pennsylvania 
Mr. Seal has found this fish abundant in pools and ditches along the Dela- 
ware," a statement^ which in every way agrees with my knowledge of 
Apeltes quadracus. 

Apeltes quadracus (Mitchill). Three-spined Stickleback. 
Delaware basin in Bucks, Delaware, Lehigh, Northampton and Phila- 
delphia Counties. Jordan and Evermann state "male almost black; 
ventralswith the membrane red in spring," though of the countless spawn- 
ing examples I have examined never a black male has been seen, and most 
all the males at every season of the year had red ventrals. 

APHREDODERID^. 
Aphredoderus sayanus (Gilliams). Pirate Perch. 
Delaware basin in Bucks, Delaware and Philadelphia Counties. 

MICROPTERIDiE. 
Pomoxis annularis Rafinesque. Crappie. 
Reported from the Monongahela and my material from the Kiskiminitas 
River. Likely introduced in the Delaware and Susquehanna basins, 
where frequently met with. 

Pomoxis sparoides (Lac^pede). Calico Bass. 
Delaware basin in Bucks, Delaware, Monroe, Montgomery, Northamp- 
ton, Philadelphia and Pike Counties. 

Susquehanna basin in Lancaster and York Counties. 
Ohio basin in Clarion County 
Erie basin in Erie County. 

Ambloplites rupestris (Rafinesque). Red-eyed Bass. 

Ohio basin in Beaver, Clarion, Fayette, Indiana, Lawrence, McKean, 
Warren and Westmoreland Counties. 

Genesee basin in Potter County. 

Erie basin in Crawford and Erie Counties. Introduced in the Delaware 
and Susquehanna basins. 

iGasterosteus biaculeatus Abbott, Geol. N. J., 1868, p. 814. 
2Rep. Com. Fisher. Pa., 1889-91 (1892), p. 98. 
3Bull. U. S. Nat. Mus., No. 47, T, 1896, p. 752. 



68 Proceedings of the Biological Society of Washington. 

Enneacanthus obesus (Girard). Sphagnum Sunfish. 
Only taken in the Delaware basin at Philadelphia, though abundant 
outside our limits in the Coastal Plain. 

Enneacanthus gloriosus (Holbrook). Blue-spotted Sunfish. 
Delaware basin in Bucks, Delaware and Philadelphia Counties. 
Susquehanna basin in Lancaster and Lebanon Counties. 

Mesogonistuis chaetodon (Baird). Banded Sunfish. 
Delaware basin in Bucks and Philadelphia Counties. 

Lepomis cyanellus Rafinesque. Green Sunfish. 
No definite record ever given. I include it as the late Eugene Smith 
informed me he secured several examples from the Monongahela River at 
McKeesport, over twenty years ago 

Lepomis auritus (Linnajus). Red-bellied Sunfish. 

Delaware basin in Berks, Bucks, Chester, Delaware, Lehigh, Mont- 
gomery. Northampton, Philadelphia, Pike and Wayne Counties. 

Elk basin in Chester County. 

Susquehanna basin in Berks, Center, Dauphin, Lancaster, Lebanon, 
Snyder, Sullivan and York Counties. 

Lepomis megalotis (Rafinesque). Long-eared Sunfish. 
I have it from the Kiskiminitas River, where it was obtained by Cope. 

Lepomis macrochirus Rafinesque. Chain-side Sunfish. 
Type of L. nephelus Cope, from the Kiskiminitas River, examined. 

Lepomis incisor Valenciennes. Blue Sunfish. 
Besides Cope's type of L. ardesiacus from the Kiskiminitas I have it 
from the Delaware basin in Bucks, Lehigh, Pike and Philadelphia Counties. 
Ohio basin in Warren County. 
Erie basin in Erie County. 

Pomotis gibbosus (Linnaeus). Common Sunfish. 

Delaware basin in Berks, Bucks, Carbon, Chester, Delaware, Lehigh, 
Monroe, Montgomery, Northampton, Philadelphia, Pike and Wayne 
Counties. 

Susquehanna basin in Berks, Blair, Bradford, Cambria, Clinton, Center, 
Columbia, Dauphin, Franklin, Fulton, Huntingdon, Lackawanna, Lan- 
caster, Lebanon, Luzerne, Mifflin, Montour, Northumberland, Perry, 
Sullivan, Susquehanna, Wyoming and York Counties. 

Potomac basin in Fulton County. 

Ohio basin in McKean and Indiana Counties. 

Erie basin in Erie County. 

Micropterus dolomieu Lac^pMe. Small-mouth Bass. 
Ohio basin in Allegheny, Armstrong, Beaver, Butler, Cambria, Clarion, 



Fowler — A List of the Fishes of Pennsylvania. 69 

Clearfield, Crawford, Erie, Fayette, Forest, Indiana, Lawrence, McKean, 
Mercer, Somerset, Venango and Warren Counties. 

Erie basin in Crawford and Erie Counties. Introduced in Delaware, 
Susquehanna and Potomac basins. 

Micropterus salmoides (Lac6pede). Large-mouth Bass. 
Ohio basin in Crawford, Indiana, McKean and Warren Counties. 
Erie basin in Erie County. Introduced in Delaware and Potomac basins. 

PERCID^. 
Stizostedion vitreum (Mitchill). Pike Perch. 
Genesee basin in Potter County. 

Ohio basin in Allegheny, Beaver, Butler, Cambria, Clarion, Crawford, 
Forest, Indiana, Lawrence, McKean, Warren and Westmoreland Counties. 
Erie basin in Erie County. Introduced in the Delaware and Susque- 
hanna basins. 

Stizostedion canadense (Griffiths). Sanger. 
Ohio basin in Indiana and Warren Counties, and Beaver and Youghio- 
gheny Rivers. 

Erie basin at Erie. 

Perca flavescens (Mitchill). Yellow Perch. 

Delaware basin in Bucks, Carbon, Delaware, Lehigh, Monroe, Mont- 
gomery, Pike, Philadelphia and Wayne Counties. 

Susquehanna basin in Bradford, Lackawanna, Lancaster, Luzerne, 
Mifflin, Wyoming and York Counties. 

Erie basin in Erie County. Introduced in Ohio basin of Crawford 
County. 

Percina caprodes (Rafinesque). Log Perch. 

Susquehanna basin in Lancaster County. 

Ohio basin in Crawford County. Reported from the Allegheny and 
Monongahela and I have Cope's Kiskiminitas and Youghiogheny River 
material. 

Erie basin in Erie County. 

Haldeman described it as Perca nebulosa from the Susquehanna.' At the 
same time he also described Perca minima from the same stream, though 
not the young as he suggests, but simply a tessellated darter. He also 
gave the manuscript name Percina himaculata to Storer for still another 
example from the Susquehanna several years later, ^ this insuring the 
generic name for the log perches. The types of the first two nominal 
forms I have examined. Cope overlooks' the occurrence of the log perch 
in the Susquehanna, as he only mentions it from Lake Erie and the Alle- 
gheny River. 

iJourn. Acad. Nat. Sci. Phila., 1842, p. 330. 
2Proc. Boston Soc. Nat. Hist., 1844, p. 157. 
3Rep. Com. Fisher. Pa., 1879-80 (1881), p. 124. 



70 Proceedings of the Biological Society of Washington. 

Hadropterus phoxocephalus (Nelson). Slender-headed Darter. 
Reported from the Monongahela by Evermann and Bollman.* 

Hadropterus macrocephalus (Cope). Long-headed Darter. 
Cope's types of Etheostoma macrocephalum from the Youghiogheny ex- 
amined. The late Dr. Bean reports it from French Creek^ and Foxburg.^ 

Hadropterus peltatus (Cope). Shielded Darter. 
Delaware basin in Montgomery County. 
Susquehanna basin in Berks, Dauphin, Lancaster and Snyder Counties. 

Hadropterus aspro (Jordan). Black-sided Darter. 
A number of examples from Indiana County, some I wrongly identified 
with H. macrocephaltcs.* 

Etheostoma blennioides Rafinesque. Green-sided Darter. 
Reported from the Kiskiminitas and Monongahela Rivers, and my 
material from the Beaver River. 

Boleosoma nigrum (Rafinesque). Johnny Darter. 
Ohio basin in Clarion, Indiana and McKean Counties. 

Boleosoma nigrum olmstedi (Storer). Tessellated Darter. 

Delaware basin in Berks, Carbon, Chester, Delaware, Lehigh, Monroe, 
Montgomery, Northampton, Pliiladelphia and Pike Counties. 

Susquehanna basin in Berks, Blair, Cameron, Chester, Cumberland, 
Huntingdon, Columbia, Dauphin, Lancaster, Luzerne, Mifflin, Perry, 
Snyder, SuUivan and York Counties. 

Elk basin in Chester County. 

Ammocrypta pellucida (Putnam). Sand Darter. 
Reported from the Youghiogheny and the Monongahela. I have it 
from Erie. 

Poecilichthys variatus (Kirtland). Variegated Darter. 
Described as Hadropterus tessellatus by Jordan from the Allegheny at 
Foxburg.^ Evermann and Bollman report it from the Monongahela,* and 
McConnell mentions it from French Creek above Franklin.' The specimen 
mentioned by Jordan and Evermann from Easton is likely with wrong 
locality.* 

lAnn. N. Y. Acad. Sci., Ill, 1883-5 (1886). p. 339. 

2Rep. Com. Fisher. Pa., 1889-91 (1892), p. 122. 

3Bull. U. S. Nat. Mu8., No. 27, 1884, p. 466. 

^Science, XXXI, March 4, 1910, p. 346; Proc. Acad. Nat. Sci. Phila., 1912, p. 49. 

5Bull. U. S. Nat. Mus., No. 10, 1877, p. 7. 

6Ann. N. Y. Acad. Sci., Ill, 1883-5 (1886), p. 339. 

7Rep. Com. Fisher. Pa., 1905, p. 178. 

8BulI. U. S. Nat. Mus., No. 47, 1, 1896, p. 1069. 



Fowler — A List of the Fishes of Pennsylvania. 71 

Poecilichthys zonalis Cope. Zoned Darter. 
Recorded from the Monongahela by Evermann and Bollman.i 

Poecilichthys maculatus (Kirtland). Trout Darter. 
McConnell records it from the Shemango River at Jamestown. ^ 

Poecilichthys coeruleus (Storer). Blue Darter. 
Recorded from the Monongahela by Evermann and BoUman, and I have 
it from the Kiskiminitas. 

Poecilichthys flabellaris (Rafinesque). Fantail Darter. 
Ohio basin in Allegheny, Indiana, McKean and Westmoreland Counties. 
Susquehanna basin in Perry County. 

Boleichthys fusiformis (Girard). Lowland Darter. 
Only met with in the Delaware tidal region near Bristol, Bucks County. 

SERRANID^. 
Roccus chrysops (Rafinesque). White Bass. 
Genesee basin in Potter County and Erie basin at Erie. Ohio basin in 
Clarion, Indiana, Mercer and Warren Counties. Introduced in the Dela- 
ware basin. 

Roccus lineatus (Bloch). Striped Bass. 
Delaware basin in Bucks, Chester, Delaware, Montgomery, Philadelphia 
and Pike Counties. 
Susquehanna basin in Dauphin, Lancaster, Luzerne and York Counties. 

Morone americana (Gmelin). White Perch. 
Delaware basin in Bucks, Carbon, Delaware, Montgomery, Northamp- 
ton and Philadelphia Coimties. 

Susquehanna basin in Lancaster and York Counties. 

SCI^NID^. 
Aplodinotus grunniens Rafinesque. Fresh-water Drum. 
Ohio basin in the Allegheny and Monongahela, and Indiana County. 
Erie basin in Erie County. 

COTTIDiE. 
Cottus ictalops (Rafinesque). Western Sculpin. 

Genesee basin in Potter County. 

Ohio basin in Allegheny, Clarion, Crawford, Indiana, McKean, Somerset 
and Westmoreland Counties. 

Cottus gracilis Heckel. Sculpin. 
Delaware basin in Berks, Bucks, Carbon, Chester, Lehigh, Montgomery 
and Northampton Counties. 

lAnn. Acad. Nat. Sci., Ill, 1883-5 (1886). p. 339. 
2Rep. Com. Fisher. Pa., 1905, p. 178. 



72 Proceedings of the Biological Society of Washington. 

Elk basin in Chester County. 

Susquehanna basin in Berks. Center, Columbia, Cumberland, Dauphin, 
Lancaster, Luzerne, Huntingdon, Perry and Snyder Counties. 

SOLEIDiE. 
Achirus fasciatus Lacepede. American Sole. 
Delaware basin in Bucks and Philadelphia Counties. 

GADID.E. 
Lota maculosa (Le Sueur). Burbot. 
Found at Erie. Introduced in the Susquehanna and Delaware. 



Besides the species already indicated as introduced are several others 
and several which may be considered accidental Such are: 

Carcharias taurus Rafinesque. Sand Shark. 
Once recorded in the region of the Delaware opposite Philadelphia. 

Eulamia milberti MuUer and Henle. Brown Shark. 
Once found in the Delaware opposite lower Philadelphia. 

Raja ocellata Mitchill. Big Spotted Skate. 
Once in the Delaware opposite upper Philadelphia. 

Oncorhynchus tschawytscha (Walbaum). Quinnat Salmon. 
Introduced in the Delaware and Susquehanna. 

Salmo salar IJnnaeus. Salmon. 
Introduced in the Delaware. 

Salmo salar sebago (Girard). Landlocked Salmon. 
Introduced in Luzerne, Monroe, Pike and Wayne Counties. 

Salmo fario Linnaeus. Brown Trout. 
Introduced in Blair, Huntingdon, Luzerne, Monroe, Philadelphia, Pike 
and Wayne Counties. 

Salmo clarkii Richardson. Columbia River Trout. 

Introduced in Center and Susquehanna Counties. 

Salmo irideus Gibbons. Rainbow Trout. 
Introduced in Center, Clearfield, Cumberland, Fayette, Franklin, Mont- 
gomery, Pike, Wayne and Wyoming Counties. 

Thymallus tricolor Cope. Michigan Grayling. 
Introduced. 



Fowler — A List of the Fishes of Pennsylvania. 73 

Cyprinus carpio Linnaeus. Carp. 

Introduced. Delaware basin in Berks, Bucks, Carbon, Chester, Lehigh, 
Montgomery, Northampton and Philadelphia Counties. 

Susquehanna basin in Adams, Blair, Bradford, Center, Clearfield, 
Columbia, Huntingdon, Cumberland, Lackawanna, Lancaster, Lebanon, 
Lycoming, Luzerne, Mifflin, Perry, Snyder, Susquehanna, Union and York 
Counties. 

Ohio basin in Allegheny, Armstrong, Butler, Crawford, Greene, Indiana, 
Jefferson, Lawrence, Mercer, Somerset, Venango, Washington, Warren and 
Westmoreland Counties. 

Erie basin in Erie County. 

Carassius auratus (Lirmseus). Goldfish. 
Introduced. Delaware basin in Bucks, Lehigh and Philadelphia Counties, 

Scardinius erythrophthalmus (Linnaeus). 
Introduced about Philadelphia. 

Pomatomus saltatrix (Linnaeus). Blue Fish, 
Accidental in the Delaware at Philadelphia. 

Chaenobryttus gulosus (Cuvier). Warmouth. 
Introduced in the Delaware and Susquehanna. 

Morone interrupta Gill. Yellow Bass, 
Introduced in the Delaware. 



.^^'