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AGRIC',"  OTfTf* 


AGRIC. 
'    LIBRARY 

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DEPARTMENT  OE  AGRICULTURE, 

NEW  SOUTH   WALES. 


SCIENCE  BULLETIN, 
No.  3. 


September,  1911, 


March  Flies 


WALTER  W.  EROGGATT,  E.L.5., 

Government  Entomologist. 


Workers  in  the  respective  branches  of  Economic  Science  covered 

by  this  series  of  Science  Bulletins  will  receive  such  of  them  as  may 

be  of  use  in  their  special  branches  of  study  upon  application  to  the 

Under  Secretary,  Department  of  Agriculture,  Sydney. 


t  1586 


SYDNEY;    WILLIAM  APPLCGATC  GULLICK,  GOVERNMENT  PRINTER.— 1911, 


No.  of  Copies  Issued,  1,000. 


DEPARTMENT    OF  AGRICULTURE, 
NEW  SOUTH  WALES. 


SCIENCE  BULLETIN,  No.  3. 


MARCH    FLIES. 


BY 

WALTER  W.  FROGGATT,  F.L.S., 

Government  Entomologist. 


t  1586 


SYDNEY:   WILLIAM   APPLEGATE  GULLICKt  GOVERNMENT   PRINTER. 

1911. 


DEPARTMENT  OF  AGRICULTURE, 

NEW    SOUTH    WALES. 


SCIENCE   BULLETIN,  No.  3. 


MARCH    FLIES. 


WALTER  W.  FROGGATT,  F.L.S.,  Government  Entomologist. 

THE  family  Tabanidke  is  one  of  the  most  important  groups  of  the  flies  from 
an  economic  point  of  view,  in  habits,  range,  and  numbers.  In  the  latest 
list  of  Diptera,  about  1,800  species  have  been  described,  of  which  the 
typical  genus  Tabanus  contains  half,  or  900  species,  from  all  parts  of  the 
world,  and  in  this  number  are  included  forty-eight  species  found  in 
Australia. 

These  biting  flies  are  commonly  known  in  the  Australian  bush  as  "  March 
Flies,"  under  the  impression  that  they  appear  in  March;  but  though  usually 
most  abundant  towards  the  end  of  summer,  they  can  be  found  in  suitable 
localities  all  through  the  warmer  months  of  the  year.  In  England  their 
popular  name  is  "  Breeze  Flies,"  on  account  of  their  loud  hum  when,  flying; 
or  "  Gad  Flies,"  on  account  of  their  blood-sucking  propensities,  and  the 
way  they  annoy  horses  and  cattle  in  their  resting-places.  In  America  they 
are  known  as  "  Green  Heads,"  from  the  usual  deep  green  tint  of  their  very 
large  eyes;  and  in  some  districts  simply  as  "  Horse  Flies." 

These  flies  are  well  known  in  the  bush,  not  onh^  from  the  savage  way  in 
which  they  fasten  on  one's  horse  to  suck  up  blood,  but  also  from  the  per- 
sistent manner  in  which  they  settle  upon  the  hands  and  neck  of  the  traveller, 
and  allow  themselves  to  be  killed  in  their  eagerness  to  draw  blood. 

The  typical  March  Fly  is  of  a  uniform  greyish  brown,  or  dull-yellow 
tint,  marked  with  brown;  the  abdominal  segments  barred  or  blotched  with 
darker  tints.  The  head  is  large,  the  greater  portion  taken  up  with  the  large 
eyes,  which  in  the  males  almost  meet  at  the  front  of  the  head.  '  The 
thorax  is  well  defined,  as  broad  as  the  body,  and  fitting  close  against  the 
head  along  the  front  margin.  The  wings  are  large,  strongly  veined,  and 
adapted  for  sustained  flight;  the  legs  stout;  and  the  abdomen  of  a  uniform 
width  from  the  hind  margin  of  the  thorax  to  the  rounded  tip  of  the  abdomen, 
and  somewhat  flattened  on  the  dorsal  surface. 

The  male  March  Flies  live  upon  the  sap  of  plants,  the  honey-dew  exuded 
by  homopterous  insects,  aphids,  &c. ;  and  they  sometimes '  attack  aphids  and 
other  soft-bodied  insects  and  suck  up  their  blood ;  but  it  is  only  the  female 
flies  that  have  the  true  blood-sucking  instinct.  Like  mosquitoes "  and  other 
blood-sucking  flies,  it  is  probable  that  the r  females,  in ^  defrft  of 'blood, 
adopt  the  habits  of  the  males,1  and  live  upon  vegetable  s'dp. 


SCIENCE  BULLETIN    No.  3. 


The  females  deposit  their  spindle-shaped  brown  or  black  eggs  in  masses 
upon  the  foliage  of  plants  growing  on  flat,  swampy  land,  or  round  the  edges 
of  lagoons  and  water-holes.  According  to  Professor  J.  S.  Hine,  several 
species  in  the  United  States  hatch  out  in  nine  days.  The  larvae  live  in  wet 
soil,  under  rotten  logs,  or  stones,  and  sometimes  even  in  the  water,  where 
they  feed  upon  all  kinds  of  small  creatures.  Professor  Hine  reared  them 
very  easily  in  jars  of  moist  earth,  supplied  with  chopped-up  worms.  After 
pupating,  they  remained  in  the  dormant  state  till  the  following  spring,  so 
that  their  life  history  occupied  about  eleven  months. 

The  larvae  are  elongate,  eleven -jointed  maggots,  each  segment  encircled 
with  a  row  of  fleshy  tubercles,  very  well  defined  on  the  ventral  surface. 

Nothing  is  known  regarding  the  life-history  of  any  of  our  Australian 
species,  and  there  is  a  wide  field  for  investigation  in  studying  the  earlier 
stages  of  our  biting  flies. 

In  other  parts  of  the  world  a  great  deal  of  attention  has  been  given  to  the 
range  and  life-histories  of  Tabanid  flies,  since  it  has  been  proved  that  several 
species  transmit,  when  biting,  the  blood  parasites  of  man  and  domesticated 
animals,  causing  some  of  the  most  deadly  tropical  diseases.  In  most  of  the 
tropical  agricultural  experiment  stations  the  entomologists  are  devoting 
much  of  their  time  to  the  study  of  these  flies.  At  the  Agricultural  Kesearch 
Laboratories  at  Pusa,  India,  Mr.  ITowlett  was  specially  engaged  to  work 
on  these  flies;  and  at  Cairo,  the  Soudan,  and  other  stations  in  tropical 
Africa,  work  is  being  carried  out  to  find  the  distribution  of  these  insects. 
In  the  laboratories  of  the  Tropical  School  of  Medicine  at  Liverpool, 
Mr.  Newstead  is  tabulating  the  collections  received  from  all  parts  of  the 
world. 

Mr.  Austen,  of  the  British  Museum,  has  issued  two  works,  one  dealing 
with  the  "  Tse-tse  Flies  of  Africa,"  and  the  second  with  the  biting  flies  of 
Great  Britain.  Miss  Ormerod  had  previously,  in  her  "  Annual  Reports  on 
Injurious  Insects,"  devoted  many  pages  to  them.  All  over  the  world  col- 
lectors are  on  the  lookout  for  flies  belonging  to  this  family,  on  account  of 
the  important  role  they  play  in  economic  entomology. 

Through  the  publication  of  Part  III  of  Dr.  Kertesz's  "  Catalogus  Dip- 
terorum,"  1908,  we  are  now  enabled  to  obtain  some  idea  of  the  number  of 
species  that  have  been  described,  and  the  countries  in  which  they  are  found. 
It  is  very  unfortunate  that  so  many  of  the  Australian  species  have  been 
described  with  no  other  locality  attached  except  "  New  Holland "  or 
"  Australia."  It  is  therefore  only  by  collecting  specimens,  with  authentic 
locality  labels  attached,  and  by  sending  duplicates  to  specialists  where  the 
original  type  specimens  are  kept  and  having  them  determined,  that  we  can 
obtain  any  idea  of  the  range  of  our  indigenous  species  of  biting  flies. 

The  writer  proposes  to  give  some  notes  on  the  species  known  to  him,  some 
of  which  are  figured  in  the  accompanying  plate,  and  a  list  of  species 
described  from  Australia. 


The  genus  Pangonia  contains  a  number  of  large,  handsome  flies,  found 
in  open  forest  country,  often  resting  upon  the  trunks  of  trees.  The  mem- 
bers of  this  genus  have  a  world-wide  range,  and  about  twenty  species  have 


MARCH  FLIES. 


been  described  from  Australia.  A  species  has  been  recorded  from  New 
Caledonia  as  being  the  cause  of  an  outbreak  of  malignant  pustules  on 
cattle.  They  differ  from  the  typical  Tabanus  in  having  ocelli,  and  the 
third  joint  of  the  antennae  elongate,  instead  of  being  compressed. 

Pangonia  guttata,  Donov. 

(Figs.  8  and  7.) 

This  large  and  handsome  species  was  originally  described  and  figured  by 
Donovan  in  his  now  rare  and  costly  work,  "Insects  of  New  Holland,  1805." 
It  has  been  described  and  noticed  a  good  many  times  since;  by  Guerin  in 
the  "Zoologie  du  Voyage  de  la  Coquille,"  published  in  1830;  it  was  again 
figured  by  Macquart  seven  years  later;  and  it  is  the  insect  shown  as  No.  2 
on  Plate  XXVIII  in  "Australian  Insects,  1907."  It  has  a  wide  range 
along  the  eastern  coast  of  Australia,  but  I  have  seen  no  specimens  from  the 
western  coast.  Kertesz  also  records  it  from  Asia. 

It  is  a  large  fly,  measuring  1|  inches  from  the  tip  of  the  wings  to  the  front 
of  the  head,  and  is  broad  in  proportion.  The  general  colour  is  black  to  dark- 
chocolate  brown  011  the  upper  surface  of  the  thorax;  the  wings  clouded  with 
black  at  the  base,  running  out  into  light  brown  towards  the  tips.  The  hind 
margin  of  the  head  is  silvery  white;  the  centre  of  the  thorax  marked  with 
two  parallel  white  stripes;  a  tuft  of  black  hairs  on  either  side,  the  sides  of 
the  thorax  further  ornamented  with  fine  silky-white  hairs  that  form  an 
irregular  band,  with  two  tufts  011  the  hind  margin.  Both  the  upper  and 
under  surfaces  of  the  abdomen  are  marked  with  tufts  of  white  hairs,  the 
cuter  margins  fringed  with  tufts  of  white  and  black  hairs,  the  latter  also 
fringing  the  hind  margin  of  the  apical  segments  of  the  abdomen. 

Pangonia  auriflua,  Donov. 

(Fig.  4.) 

This  handsome  fly  also  has  a  wide  range  along  the  eastern  coast,  and  was 
among  the  insects  figured  by  Donovan.  Kertesz  has  placed  it  in  Rondano's 
genus  Diatomineura. 

It  measures  under  three-quarters  of  an  inch  from  the  front  of  the 
head  to  the  tip  of  wings,  and  is  of  the  usual  stout  form,  with 
rather  short,  rounded  abdomen.  The  general  colour  of  the  eyes,  upper 
surface  of  the  thorax,  and  abdomen  is  black;  the  face,  hind  margin  of 
head,  and  under  surface  of  thorax  and  abdomen  clothed  with  silvery 
white  hairs.  The  centre  of  the  dorsal  surface  of  the  thorax  is  marked  with 
pale,  whitish  parallel  bars,  with  a  broad  band  of  golden  yellow  hairs  right 
round  the  sides  and  hind  margin;  a  parallel  row  of  three  tufts  down  the 
centre  of  the  abdomen,  with  the  apical  segments  densely  clothed  with  bright 
golden  hairs.  The  wings  are  semi-transparent,  with  black  nervures. 


The  genus  Tabanus  contains  our  common  and  most  typical  March  Flies. 
It  embraces  an  immense  number  of  species,  distinguished  from  the  other 
genera  of  the  family  in  having  the  eyes  bare  (not  pubescent),  and  having 
no  raised  tubercle  for  the  ocelli. 


SCIENCE  BULLETIN,  No.  3. 


Tabanus  abstersus,  Walk. 

This  is  one  of  the  commonest  and  most  widely  distributed  species  in 
Australia,  specimens  have  been  received  from  Cardwell,  North  Queens- 
land; Clarence  River  and  Bega,  New  South  Wales. 

It  measures  slightly  over  half  an  inch  in  length  from  the  front  of  the  head 
to  the  tips  of  the  wings.  It  is  of  the  usual  stout  form,  with  the  abdomen  some- 
what flattened.  The  general  colour  is  dark-brown,  shading  into  grey  tints; 
the  wings  semi-opaque,  with  the  nervures,  base,  and  centre  of  the  wings 
clouded  with  brown.  The  under  surface  of  the  head  and  thorax  is  clothed 
with  fine,  grey,  downy  hairs.  The  abdominal  segments  are  edged  along  the 
hind  margin  with  a  fine  white  band,  well-defined  011  the  first  four  segments, 
and  forming  an  angular  blotch  in  the  centre,  sometimes,  though  not  so 
distinctly,  expanded  on  the  sides. 

,  ,    Tabanus  brevidentatus,  Macq. 

(Fig.  1.) 

This  is  the  common,  small,  greyish-brown  March  Fly,  found  in  the  scrub 
or  bush  around  Sydney,  which  comes  round  trying  to  settle  upon  one's  face 
or  hands,  to  bite  if  undisturbed.  They  are  most  plentiful  about  shaded 
watercourses  and  valleys  in  the  Hawkesbury  country,  and  have  a  wide  range 
along  the  coast. 

It  measures  under  three-quarters  of  an  inch  in  length  from  the  front  of 
the  head  to  the  tips  of  the  wings.  The  general  colour  is  greyish-brown; 
face  and  under  surface  of  the  head  and  thorax  clothed  with  silvery- white 
hairs.  The  centre  of  the  upper  surface  of  the  thorax  is  marbled  with 
darker  brown ;  and  the  central  portions  of  the  abdominal  segments  lighter 
than  the  outer  edges,  with  a  delicate  white  line  along  the  extreme  hind 
margin  of  each  segment.  The  wings  are  semi-transparent,  with  brown 
nervures. 

Tabanus  sanguinarius,  Bigot. 

(Fig.  2.) 

This  is  one  of  our  largest  species,  measuring  up  to  an  inch  in  length 
from  the  front  of  the  head  to  the  tips  of  the  wings.  The  eyes  are  dark 
brown,  with  rich,  silvery  tints ;  the  face  dull  white.  General  colour,  yellowish 
brown;  semi-opaque  wings,  slightly  clouded  with  dull  yellow  along  the  front 
margins.  The  abdomen  is  somewhat  flattened  and  rounded  at  the  tip. 
This  March  Fly  is  common  in  the  scrubs  of  the  Tweed  River  and  the 
northern  parts  of  New  South  Wales,  and  ranges  into  Southern  Queensland. 

Tabanus  ardens,  Wiedm. 

(Fig.  3.) 

This  fly  was  originally  described  from  Java,  but  probably  has  a  wide 
range  over  the  M&lay  Archipelago,  as  it  js  very  common  in  the1  open  forest 
country  of  the  Solomon  Islands. 


MARCH  FLIES. 


It  measures  slightly  over  half  an  inch  in  length  from  the  front  of  the 
head  to  the  tips  of  the  wings.  The  eyes  are  dark  and  shining;  face  dull 
yellow;  general  colour  brownish  yellow,  with  the  upper  surface  of  the  last 
two  segments  clouded  in  the  centre  with  brown.  The  sides  and  under 
surface  of  thorax  and  abdomen  are  clothed  with  yellow  hairs,  which  also 
form  a  fringe  along  the  outer  edge  of  the  abdominal  segments.  Wings 
semi-opaque,  clouded  with  brown;  nervures  brown. 

Tabanus  edentulus,  Macq. 

(Fig.  5.) 

In  size  and  general  colouration  this  species  might  be  easily  taken  for 
Tabanus  brevidentatus ;  but  it  can  be  easily  distinguished  by  the  more 
irregular  or  lobed  form  of  the  thorax,  which  is  of  a  uniform  greyish-brown 
tint,  not  showing  any  darker  black  or  brownish  markings  on  the  dorsal 
surface,  while  the  apical  bands  on  the  hind  margins  of  the  abdominal 
segments  are  more  defined  and  lighter  coloured. 

This  species  is  common  on  the  highlands  of  the  southern  parts  of  New 
South  Wales,  and  probably  ranges  into  eastern  Victoria.  Specimens  in  our 
collections  were  obtained  at  Binda,  near  Bega,  and  others  from  the  slopes 
of  Mount  Kosciusko. 

Tabanus  gregarius,  Erich. 

(Fig.  9.) 

This  distinctive  species  is  just  under  three-quarters  of  an  inch  in  length 
from  the  front  of  the  head  to  the  tips  of  the  wings.  The  head  and  thorax 
are  lead-coloured;  antennae  yellow  at  base,  black  at  the  tips;  legs  brownish 
on  thighs,  with  tibiae  and  tarsi  yellow ;  wings  hyaline,  with  the  front  margin 
very  lightly  clouded  with  yellow.  The  under  surface  of  the  head  and 
thorax  is  clothed  with  long  silvery  hairs;  the  abdomen  with  fine  golden 
hairs  on  upper  and  under  surface.  The  abdomen  is  dull  yellow,  with  a 
broad  stripe  of  blackish  brown  down  the  centre  of  the  back,  spreading  out 
towards  the  apex. 

This  species  was  originally  described  by  Erichson,  from  Tasmania,  but 
ranges  along  the  eastern  coast  of  Australia  well  into  Queensland. 

Tabanus  jacksonii,  Macq. 

(Fig.  6.) 

This  large  species  is  common  on  the  slopes  of  Mount  Kosciusko,  and 
differs  from  all  the  other  greyish-brown  species  in  its  larger  size,  uniform 
lead-coloured  thorax,  and  the  abdominal  segments  being  mottled  with  yellow 
and  brown  down  the  centre,  forming  fine  transverse  bands  fringed  with 
golden  hairs  along  the  apical  segments. 

The  face  is  grey,  the  antennae  and  legs  yellow.  The  under  surface  of  the 
head  and  thorax  is  densely  clothed  with  long,  white,  downy  hairs ;  the  under 
surface  of  the  abdomen  with  more  scattered  hairs.  Wings  semi- opaque, 
slightly  clouded,  and  blotched  in  the  centre. 


SCIENCE  BULLETIN,  No.  3. 


Tabanus  Regis-Georgii,  Maeq. 

This  is  one  of  the  common  species,  found  from  the  Tweed  River,  New 
South  Wales,  to  Cardwell,  North  Queensland. 

It  measures  slightly  over  half  an  inch  in  length  from  the  front  of  the  head 
to  the  tips  of  the  wings.  It  is  of  a  uniform  light-brown  tint,  marbled  with 
grey  on  the  thorax,  with  the  hind  margins  of  the  abdominal  segments  edged 
with  greyish  white;  the  wings  transparent,  with  the  front  (subcostal)  area 
shaded  with  brown. 

Tabanus  nigro-pictus,  Macq. 

This  is  a  species  found  about  the  Richmond  River,  New  South  Wales. 
It  is  a  large  fly,  measuring  nearly  an  inch  in  length;  of  a  uniform  reddish- 
brown  tint;  the  upper  surface  clothed  with  fine  golden  hairs,  the  under 
surface  with  grey  hairs.  The  eyes  are  lead  colour;  and  the  dorsal  surface 
of  the  thorax  is  mottled  with  black  showing  through  the  golden  pubescence- 
It  resembles  Tabanus  sanguinarius  in  size  and  general  colouration,  but 
differs  in  having  darker  markings  upon  the  thorax. 

Tabanus  concolor,  Walker. 

This  species  has  a  wide  range  from  Eastern  Victoria  to  the  Tweed 
Hiver,  New  South  Wales,  and  probably  Southern  Queensland.  We  have 
specimens  from  Mount  Kosciusko. 

It  measures  just  under  an  inch  in  length  from  the  front  of  the  head  to  the 
tips  of  the  wings,  and  is  of  a  uniform  dark-lead  colour.  There  are  a  few 
•cattered,  short,  white  hairs  on  the  sides  of  the  thorax  and  along  the  apical 
edge  of  the  abdominal  segments.  The  under  surface  of  the  head  and 
thorax  is  clothed  with  silvery  long  hairs,  thickest  below  the  mouth  parts. 
The  wings  have  brown  nervures,  and  are  slightly  clouded,  darkest  towards 
the  shoulders. 


LIST  OP  THE  AUSTRALIAN  SPECIES  OF  THE  BITING  FLIES  OF  THE  FAMILY 
Tabanidce,  DESCRIBED  IN  Dr.  C.  KERTESZ'S  CATALOGUE  OF  THE  DIPTERA 
OF  THE  WORLD,  VOL.  III.,  BUDAPEST,  1908. 

Family : — TABANID.E. 

Sub-family :— Pangoniinre. 

Genus  :— APOCAMPTA  (Schiner,  1867). 

1.  Apocampta  subcana,  Walker,  List  Dip.  Brit.  Museum,   1848  ;    Ricardo, 
Annal.  Mag.  Nat.  Hist.  ser.  7,  viii,  287,  1901 ;  and  ser.  7,  xiv,  357,  1904. 
Walker  described  it  under  the  Genus  Chrysops. 

Diachlorus  melas,  Bigot,  1892,  and  Apocampta  nigra,  Schiner,  1868, 

are  the  same  insect. 

AUSTRALIA. 


MARCH  FLIES.  9 


Genus  :— PELECORRHYNCHUS  (Macquart,  1850). 

2.  Pelecorrhynchus  eristaloides,  Walker,  I.e.,  p.  193,  1848. 

Walker  described   this  species  in  the  Genus  Silvius,  1848,  and  later, 
in  the  Supplementary  Catalogue,  1854,  in  the  Genus  Dasybasis. 

AUSTRALIA  AND  TASMANIA. 

3.  P.fusconiger,  Walker,  I.e.,  p.  192,  1848. 

He  dealt  with  this  species  as  with  the  previous  one,  placing  it  in  the 

Genus  Silvius. 

PORT  STEPHENS,  N.S.W.,  AUSTRALIA. 

4.  P.   maculipennis,  Macq.,   Exotic  Diptera,   1850.     Suppl.   4,  28,   Tab.  ii; 

Fig.  6. 

Macquart  figured  this  fly,  and  it  is  again  figured  in  the  Eugenies  Resa, 
1868,  by  Thomas,  who  placed  it  in  the  Genus  Ccenopnyga. 

AUSTRALIA  AND  TASMANIA 

5.  P.  personatus,  Walker,  l.c.,  p.  192,  and  Suppl.  1,  267,  1848. 

Walker  dealt  with  this  species  as  with  Nos.  2  and  3. 
Schiner  renamed  it  P.  ornatus  from  New  Zealand,  1868. 

AUSTRALIA  AND  NEW  ZEALAND. 

Genus:— SCIONS  (Walker,  1850). 

6.  Scione  singularis,  Macq.,  I.e.,  p.  192,  Supp.  1,  27,  Tab.  iii,  Fig.  2,  1845. 

Macquart  figured  this  species  under  the  Genus  Pangonia. 

AUSTRALIA. 

Genus :— PANGONIA  (Latreille,  1802). 

7.  Pangonia  binotata,  Latr.,  Encyclop.  Method,  viii,  703,  1811. 

AUSTRALIA. 

8.  P.  conjungens,  Walk.,  List  Dipt.  B.  Mus.  140,  1848.     Ricardo  Annal.  and 

Mag.     N.  Hist,  v,  114,  1900. 

WESTERN  AUSTRALIA. 

9.  P.  dorsalis,  Macq.  Dipt.   Exot.  100,  1838.     Walker,  List  Dip.  B.  Mus., 

Supp.  1,  144,  1854.     Ricardo,  l.c,  p.  114,  1900. 

AUSTRALIA. 

10.  P.  fulviventris,  Macq.,  I.e.,  1838.     Walk, l.c.,  1854.     Ricardo,  I.e.,  p.  112, 

1900.  AUSTRALIA. 

11.  P.  maculiventris,  Westw.     London  and   Edinburgh,  Philos.  Magaz.,  <fec., 

vi,  449,  1835.     Isis,  ii,  87,  1838.     Ricardo,  I.e.,  p.  114,  1900. 

AUSTRALIA. 

12.  P.  roei,  King's  Narrative,  Survey  Coasts  of  Australia,  ii,  467,  185, 1827. 

Wied,  Aussereurop.  Zweifl.,  Ins.  11,  647,  72,  1830.     Walk.,  l.c ,  p.  145, 
1848,  and  Supp.  1,  145,  136,  1854.     Ricardo,  l.c.,  p.  114,  1900. 

AUSTRALIA. 

13.  P.  Walkeri,  Newm.,  Trans.  Ent,  Soc.,  London,  n.  ser.  iv,  56,  1,  1857. 

AUSTRALIA. 


10  SCIENCE  BULLETIN,  No.  3. 

Genus  :— EREPHOPSTS  (Ricardo,  1900). 

14.  Erephopsis   albinota,    Macq.,    I.e.,    Supp.,  5,   20,    61,    1855    (Pangonia). 

Ricardo,  I.e.,  p.  112,  1900.  AUSTRALIA. 

15.  E.  aureohirta,  Ricardo,  I.e.,  p.  v,  116,  Tab.  i,  Fig.  10,  1900. 

AUSTRALIA. 

16.  E.  contigua,  Walk.,  I.e.,  p.  138,  1848.     Ricardo,  ser.  v,  112-116,  1900. 

AUSTRALIA. 

17.  E.  dilatata,  Jaennicke,  Abhandb.  Senckinberg,  Naturforsch,  Ges.,  vi,  328, 

20,  1867.     Ricardo,  I.e.,  p.  v,  112,  1900. 

AUSTRALIA. 

18.  E.  divisa,  Walk.,   Insecta   Saunders,   Dip.  i,  17,   1850.     Ricardo,  I.e.,  p. 

112-117,  1900.  AUSTRALIA. 

19.  E.  gemina,  Walk.,  List.   Dip.   B.  Mus.   v,    138,   1848.     Ricardo,   I.e.,  p. 

112-117,  1900. 

WESTERN  AUSTRALIA. 

20.  E.  guttata,  Don.,  Ins.  N.  Holland,  1805.     Tab.  and  figure.     Latr.  Ency. 

Method,  viii,  703,  5,  1811.  Wied.  Aussereurop.  Zweifl.,  Ins.  i,  194,  124, 
1828.  Guer.  Voy.  de  Coquille,  Zool.  ii,  2,  289,  1830.  Walk.,  I.e.,  p. 
i,  141,  1848,  and  Supp.,  1854.  Ricardo,  I.e.,  p.  112-114,  1900. 

Syn:  margaritifera,  Wied,  I.e.,  p,  1828.  Macq.  Ann.  Soc.  Ent. 
France,  vi,  429,  1837  :  Dip.  Exot.,  Supp.  3,  8,  1847.  Williston, 
Kansas  Univ.,  Quart,  iii,  191,  1895. 

EAST  COAST,  N.  S.  WALES,  AUSTRALIA. 

21.  E.  Jacksonii,  Macq.,  Dip.   Exot.,  i,  102,  10,  1838.     Walk.,  I.e.,  p.  1848  ; 

Insecta  Saunders,  Dip.  i,  14,  1850;  List  Dip.,  Supp.  v,  143,  126,  1854. 
Ricardo,  I.e.,  p.  112-117,  1900. 

NEW  SOUTH  WALES,  AND  SWAN  RIVER,  WESTERN  AUSTRALIA. 

22.  E.  limUnervis,  Macq.,    I.e.,    p.  21,    62,    1855.     Ricardo,    I.e.,    p.   v,  112, 

1900.  AUSTRALIA. 

23.  E.  macroporum,  Macq.,  I.e.,  Tab.  xv,  Fig  2,  1838.  Walk.,  I.e.,  p.  v,  Supp. 

i,  143,  129,  1854.     Ricardo,  I.e.,  p.  112-117,  1900. 

KANGAROO  ISLAND,  SOUTH  AUSTRALIA. 

24.  E.  maculipennis,  Macq.,   I.e.,  Supp.  4,   20,   45,  1850.     Schiner,  Novara 

Reise,  Dip.  99,  46,  1868.     Ricardo,  I.e.,  p.  106,  1900. 

AUSTRALIA. 

25.  E.  Media,  Walk.,  I.e.,  p.  42,  1848.     Ricardo,  Z.c.,  p!  112,  1900. 

26.  E.  quadrimacula,  Walk,  I.e.,  p.  1,  138,  1848.     Ricardo,   I.e.,  p.  112-116, 

1900.  AUSTRALIA. 

27.  E.  submacula,  Walk.,  I.e..  p.  142,  1848.     Insecta  Saunders,  i,   13,  1850. 

Ricardo,  I.e.,  p.  112-115, 1900.  AUSTRALIA. 

28.  E.  tricolor,  Walk.,  I.e.,  p.  139,  1848.     Ricardo,  I.e.,  p.  112-117,  1900. 

WESTERN  AUSTRALIA 


MARCH  FLIES.  11 


Genus  :— DIATOMINEURA  (Rondani,  1863). 

29.  Diatomineura  auriflua,  Don,  Ins.  N.  Holland,  1805  ;    figured  under  the 

name  of  Pangonia.     Wied.  Aussereurop,  Zweifl.   Ins.   194,   125,  1828. 
Walk.,  I.e.,  Supp.  i,  142,  122,  1854.     Ricardo,  I.e.,  p.  112-119,  1900. 

AUSTRALIA. 

30.  D.  brevirostris,  Macq.,   Dipt.  Exot.,  Supp.  4,  22,  48,  1850.     Ricardo,  I.e., 

p.  113,  1900. 

AUSTRALIA. 

31.  D.  carnea,  Bigot,  Mem.  Soc.  Zool.  France,  v,  619,  1892.     Ricardo,  I.e., 

p.  113,  1900. 

AUSTRALIA. 

32.  D.  clavata,  Macq.,  I.e.,   p.  101,  1838.     Walk.,    I.e.,    Supp.    1,    143,  125, 

1854.     Ricardo,  I.e.,  p.  113,  1900. 

AUSTRALIA. 

33.  D.  concolor,  Walk.,  Insecta,   Saunders,   Dip.  1,   14,  1850.     Ricardo,  I.e., 

p.  113-118,  1900. 

AUSTRALIA, 

34.  D.  constans,  Walk.,  I.e.,  p.  1,  15,  1850.     Ricardo,  I.e.,  p.  113-118,  1900. 

AUSTRALIA. 

35.  D.  dorsomaculata ,  Macq.,    I.e.,  Supp.  4,  21,    47,   1850.     Ricardo,  I.e.,  p. 

113,  1900.  TASMANIA. 

36.  D.gagitina,  Bigot,  I.e.,  p.  620,  1892.     Ricardo,  I.e.,  p.  113,  1900. 

AUSTRALIA. 

37.  D.  gemella,  Walk.,  List.   Dip.  B.M.  1,   139,  1848.     Ricardo,   I.e.,  p.  118, 

1900. 

WESTERN  AUSTRALIA. 

38.  D.  jacksonienis,  Guer.,  Voy.  de  la  Coquille,  Zool.  ii,  2,  288.     Tab.  xx, 

Fig.  3,  1832.     Ricardo,  l.c.,  p.  118,  1900. 

AUSTRALIA. 

39.  D.  minima,  Ricardo,  Ann.  Mag.  Nat.  Hist.,  Ser.   7,    119,  Tab.  1,  Figs. 

4-5,  1900.  AUSTRALIA, 

40.  D.patula,  Walk.,  I.e.,  p.  144,  1848.     Ricardo,  I.e.,  p.  120,  1900. 

AUSTRALIA. 

41.  D.plana,  Walk.,  I.e.,  p.  144,  1848.     Ricardo,  I.e.,  p.  118,  1900. 

AUSTRALIA. 

42.  D.  ruficornis,  Macq.,  Dip.  Exot.,  Supp.  1,  25,  31,  Tab.  iii,  Fig.  3,  1845. 

Walk.,  I.e.,  p.  128,  1854.     Ricardo,  I.e.,  p.  113,  1900. 

TASMANIA. 

43.  D.  mb-appendicidata,  Macq.,  I.e.,  p.  19,  Tab.  ii,  Fig.  2,  1850.     Ricardo, 

Lc.,  p.  113,  1900. 

AUSTRALIA* 


12  SCIENCE  BULLETIN,  No.  3. 

44.  D.  testacea,  Macq.,   I.e.,  p.   99,   1838.     Walk.,  I.e.,  p.    145,   135,    1854. 

Ricardo,  I.e.,  p.    113,   1900.      Hunter,  Trans.  Am.   Ent,   Soc.  Philad., 
xxvii,  134,  1901. 

AUSTRALIA. 

45.  D.  testaceomaculaia,  Macq.,  I.e.,  p.  21,  46,   1850.     Ricardo,  I.e.,  p.  113, 

1900.  AUSTRALIA. 

46.  D.  violacea,  Macq.,  I.e.,  p.  22,  49,  Tab.  ii,  Fig.  3,  1850.     Ricardo,  I.e., 

p.  v,  113,  1900. 

TASMANIA. 

Genus  :— CORIZONEURA  (Ricardo,  1900). 

Nearly  all  these  species  are  originally  described  in  the  Genus  Pangonia  or 
Tabanus. 

47.  Corizoneura  angusta,  Bigot,   Mem.  Soc.   Zool.,    France,  v,   617,    1892. 
Ricardo,  I.e.,  p.  114,  1900. 

AUSTRALIA. 

48.  C.  anthracina,  Macq.,  I.e.,  p.  23,  50,  Tab.  ii,  Fig.  3,  1850.     Ricardo,  I.e., 

p.  113,  1900. 

TASMANIA. 

49.  C.  brevipalpis,  Macq.,  I.e.,  p.  38,  1847.     Walker,  I.e.,  p.  144,  134,  1854. 
Ricardo,  I.e.,  p.  113,  1900. 

AUSTRALIA. 

50.  C.  chrysophila,  Walk.,  I.e.,  p.   155,  1848.      Ricardo,  I.e.,  p.   120,   1900. 
Jaennicke,  Abha.  Senckeriberg,  Nat.  Ges.  vi,  327,  19,  Tab.  xliii,  Fig.  3, 
1867.  AUSTRALIA. 

51.  C.  dives,  Macq.,  I.e.,  p.  30,  Tab.  iii,  Fig.  3,   1845.     Walker,  I.e.,  p.    124, 

1854.     Ricardo,  I.e.,  p.  113,  1900. 

TASMANIA. 

52.  C.  erratica,  Walk.,  I.e.,  p.  189,  1848.     Ricardo,  I.e.,  p.  121,  1900. 

AUSTRALIA. 

53.  C.  fulva,  Macq.,  I.e.,  p.  43,  1850.     Ricardo,  I.e.,  p.  120,  1900. 

AUSTRALIA. 

54.  C.  nigricornis,  Bigot,  I.e.,  p.  614,  1892.     Ricardo,  I.e.,  p.  114,  1900. 

AUSTRALIA. 

55.  C.  nigrosignata,  Thomas,  Eugenics  Resa,  Dipt.,  451, 12,  1868.     Rt3ardo, 

l.c.,  p.  113,  1900. 

AUSTRALIA. 

56.  C.  rubiginosa,  Bigot,  I.e.,  p.  617,  1892.     Ricardo,  I.e.,  p.  114,  1900. 

AUSTRALIA. 

57.  C.  rufovittata,  Macq.,  I.e.,  p.  19,  42,  Tab.  ii,  Fig.  1,  1850.     Ricardo,  l.c.y 
p.  113,  1900.  TASMANIA, 

58.  C.  trichocera,  Bigot,  I.e.,  v,  616,  1892.     Ricardo,  I.e.,  p.  114,  1900. 


MARCH  FLIES.  13 


Genus  :—  SILVIUS  (Meigen,  1820). 

59.  sihius  marginatus,  Walk.,   List  Dipt.  B.  Mus.  i,  189,  1848.     Ricardo, 

Annal.  and  Mag.  Nat.  Hist.,  ser.  7,  viii,  296,  1901. 

AUSTRALIA. 

60.  S.    nitescens,    Walk.,    Trans.     Entom.     Soc.,    London,   n.    ser.    iv,    124, 

1857.     Ricardo,  Annal.  and  Mag.  Nat.  Hist.,  ser.  7,  viii,  296,  1901. 

AUSTRALIA. 

61.  S.  Silvester,  Bergr.,  Stettin.  Entom.  Zeitg.  Iv,  71,  1894. 

AUSTRALIA. 


:—  CHRYSOPS  (Meigen,  1800). 

62.  Chrysops  testaceus,  Macq.,  Dipt.   Exot.  Supp.  4,  38,  17,  1850.     Ricardo, 
Annal.  and  Mag.  Nat.  Hist.,  ser.  7,  ix,  371,  1902. 

TASMANIA. 

Sub-family  :  —  Tabaninae. 
Genus  :—  DASYBASIS  (Macquart,  1846). 

63.  Dasybasis  appendiculata,   Macq.,   Dipt.   Exot.,  Supp.    2,   25,   1,  Tab.  i 

Fig.  1,  1846.     Walker,  List  Dipt.  B.  Mus.,  v,  Supp.  i,   266,  1,   1854 
Ricardo,  Annal.  and  Mag.  Nat.  Hist.,  ser.  7,  xiv,  350,  1902. 

AUSTRALIA 

Genus  :—  TABANUS  (Linn.,  1761). 

64.  Tabanus  abstersus,  Walk.,  Insecta  Saunders,  Dipt,  i,  58,  1850.     Schiner, 

Novara  Reise,  Dipt.,  85,  14,  1868. 

AUSTRALIA. 

65.  T.  antecedens,  Walk.,  List  Dipt.   B.   Mus.,  i,   178,  1848.     Supp.  i,  253, 

440,1854. 

TASMANIA  AND  AUSTRALIA. 

66.  T.  avidus,  Bigot,  Mem.  Soc.  Zool.,  France,  v,  673,  1892.     Described  in 
the  Genus  Atylolus. 

AUSTRALIA. 

67.  T.  basalis,  Walk.,  I.e.,  p.  i,  182,  1848.  ' 

AUSTRALIA. 

P8.  T.  bifascialus,   Macq.,  Suit,  a  Buffon,  i,   201,  17,  1834.     Walk.,  I.e.,  v, 
Supp.  i,  254,  447,  1854. 

AUSTRALIA. 

69.  T.  brevidentatus,  Macq.,  Dipt.  Exot.,  Supp.  v,  28,  118,  1855. 

TASMANIA,  QUEENSLAND,  AND  NEW  SOUTH  WALES. 

70.  T.  brevior,  Walk.,  List  Dipt.  B.  Mus.  i,  188,  1848. 

PORT  ESSINGTON,  NORTHERN  AUSTRALIA. 

71.  T.  brevitta,  Walk.,  I.e.,  p.  i,  157,  1848. 

AUSTRALIA. 


14  SCIENCE  BULLETIN,  No.  3. 

72.  T.  cinerascens,  King,  Narrative,  Survey  Coasts  of  Australia,  ii,  467,  184, 

1827.     Wied.,  Aussereurop.  Zweifl.  Ins.,  ii,  647,  71,  1830.     Walk.,  I.e., 
p.  i,  150,  1848,  v,  Supp.  i,  253,  443,  1854. 

AUSTRALIA 

73.  T.  cinereus,  Walk.,  I.e.,  p.  i,  167,  1848. 

WESTERN  AUSTRALIA. 

74.  T.  claripennis,  Big.,   Mem.    Soc.  Zool.,  France,  v,  675,  1892.     Described 
in  the  Genus  Atylotus. 

AUSTRALIA. 

75.  T.  concolor,  Walk.,  I.e.,  p.  179,  1848. 

EAST  COAST,  NEW  SOUTH  WALES. 

76.  T.  constans,  Walk.,  I.e.,  p.  186,  1848. 

AUSTRALIA. 

77.  T.  cyaneoviridus,  Macq.,  Dipt.  Exot.  Supp.  iv,  31,   101,  1850. 

TASMANIA. 

78.  T.  eyaneus,  Macq.,  I.e.,  Supp.  iv,  30,  99,  Tab.  ii,  Fig.  7,  1850. 

TASMANIA. 

79.  T.  diminutus,  Walk.,  I.e.,  p.  i,  183,  1848. 

PORT  ESSINGTON,  NORTHERN  AUSTRALIA. 

80.  T.  dorsolimaculatus,  Macq.,  I.e.,  Supp.  iv,  29,  97,  1850. 

AUSTRALIA. 

81.  T.  edentulus,  Macq.,  I.e.,  Supp.  i,  34,  68,  Tab.  iii,  Fig.  13,  1845.    Walk., 

List.  Dipt.  B.  Mus.  v,   Supp.  i,   251,  432,  1854.     Originally  described 
from  Tasmania. 

MT.  KOSCIUSKO,  NEW  SOUTH  WALES. 

82.  T.  exulans,  Erichson,  Archev.  f.  Naturgesch.,  viii,  270,  247,  1842.  Walk., 

I.e.,  p.  252,  435,  1854. 

AUSTRALIA  AND  TASMANIA. 

83.  T.  fraterculus,  Macq.,  Dipt.  Exot.,  Supp.  iv,  30,  98,  1850..V 

TASMANIA. 

84.  T.funebris,  Macq.,  I.e.,  Supp.  i,  33,  67,  Tab.  iii,   Fig..  12,  1845.     Walk., 
I.e.,  v,  Supp.  i,  254,  446,  1854. 

AUSTRALIA. 

85.  T.gentilis,  Erich.,  Arch.  f.  Naturgesch.,  viii,  1,  271,  249,  1842.     Walk., 

I.e.,  p.  254,  445,  1854. 

TASMANIA. 

86.  T.  gregarius,  Erich.,  I.e.,  p.  viii,  1,  271,  248,  1842.     Walk.,  I.e.,  Supp.  i, 

252,436,  1854. 

BRISBANE,  QUEENSLAND,  AND  TASMANIA. 

87.  T.  hyperythreus,  Bigot,  Mem.  Soc.   Zool.,   France,  v,  674,  1892.     De- 
scribed in  the  Genus  Atylotus.  >  < •  .1 

AUSTRALIA. 


MARCH  FLIES.  15 


88.  T.  imperfectus,  Walk.,  List.  Dipt.  B.  Mus.,  i,  179,  1848. 

AUSTRALIA. 

89.  T.  leueophilus,  Walk.,  I.e.,  p.  i,  1'54,  1848. 

AUSTRALIA. 

90..  T.  limbatinervis,  Macq.,  Dipt.  Exot.,  Supp.  ii,  16,   93,  1847;    and  Su'pp. 
iv,  29,  96,  1850.     Walk.,  List,  Dipt.  B.  Mus.  v,  Supp.  i,  254,  444,  1854. 

TASMANIA  AND  AUSTRALIA. 

91.  T.lunulatus,  Bigot,  Mem.  Soc.  Zool.,  France,  v,  688,  1892. 

AUSTRALIA. 

92.  T.  maeropthalmus,  Schiner,  Novara  Reise,  Dipt,  82,  7,  1868. 

AUSTRALIA. 

93.  T.  microdonta,  Macq.,  Dipt.  Exot,  Supp.  ii,  17,  94,  1847.     Walk.,  List. 

Dipt.  B.  Mus.,  v,  Supp.  i;  252,  438,  1854. 

TASMANIA. 

94.  T.  nigriventris,  Macq.,  I.e.,  Supp.  i,  34,  69,   1845.      Walk.,  I.e.,  Supp.  i, 

252,  437,  1854. 

SYDNEY,  NEW  SOUTH  WALES. 

95.  T.  nigropictus,   Macq.,   I.e.,  Supp.    v,    24,    111,    1855.      Van  d.  Walp., 

Catal.  Dipt.,  South  Asia,  60,  1896. 

INDIA,  AND  RICHMOND  RIVER,  NEW  SOUTH  WALES. 

96.  T.  pallipennis,  Macq.,  I.e.,  Supp.  i,  32,  65,  1854.     Walk.,  I.e.,  v,  Supp.  i, 

252,  439,  1854. 

AUSTRALIA. 

97.  T.  postpone™,  Walk.,  I.e.,  i,  179,  1848. 

AUSTRALIA. 

98.  T.  prcepositus,  Walk.,  I.e.,  i,  158,  1848. 

PORT  ESSINGTON,  NORTHERN  AUSTRALIA. 

99.  T.  propinquus,  Macq.,  Dipt.  Exot.,  Supp.  v,  27,  117,  1855, 

AUSTRALIA. 

100.  T.  pusillus,  Macq.,  I.e.,  p.  29,  120,  1855. 

AUSTRALIA. 

101.  T.  R&gis-Georgii,  Macq.,  I.e.,  i,  IA,   132,   21,    1838.     Walk.,  List.  Dipt. 
B.  Mus.,  i,  178,  1844 ;  Supp.  i,  251,  434,  1854. 

KING  GEORGE'S  SOUND,  WESTERN  AUSTRALIA;  NORTH  QUEENSLAND  TO 
TWEED  RIVER,  NEW  SOUTH  WALES. 

102.  T.  remo^s,,Walk.,  I.e.,  i,  177,  1848. 

AUSTRALIA. 

103.  T.  ruftfrons,  Macq.,  I.e.,  Supp..v,  28,  119,  1855. 

AUSTRALIA. 

104.  T.  rufinotatus,  Bigot,  Mem.  Soc.  Zool.,  France,  v,  673,  1892. 

AUSTRALIA. 


16  SCIENCE  BULLETIN,  No.  3. 


105.  T.  fanguinarius,  Bigot,  Mem.  Soc.  Zool.,  France,  v,  675,  1892. 

SOUTHERN  QUEENSLAND. 

106.  T.  sequent,  Walk.,  I.e.,  i,  178,  1848. 

PORT  ESSINGTON,  NORTHERN  AUSTRALIA. 

107.  T.  sidneyensis,  Macq.,  I.e.,  Supp.  i,  66,   1845.     Walk.,   I.e.,  v,   Supp.  i, 
253,  442,  1854. 

SYDNEY,  NEW  SOUTH  WALES. 

108.  T.  similis,  Macq.,  I.e.,  Supp.  iv,  31,  100,  1850, 

TASMANIA. 

109.  T.  vetustus,  Walk.,  List.  Dipt.  B.  Mus.  i,  179,  1848. 

SWAN  RIVER,  WESTERN  AUSTRALIA. 


Sydney :  William  Applegate  Gullick,  Government  Printer. — 191L 


k  / 


MARCH    FLIES. 

1.    Tabanus  brevidentatus.  2.    Tabanus  sanyuinarius. 

4.   Pangonia  auriflua.  5.   Tabanus  edentulus. 

7>  Foot  of  Pangonia  guttata.      8.  Pangonia  gutlata. 

[All  enlarged. 


3.    Tabanus  ardtns. 
6.   Tabanus  jacksomi. 
9.   Tabanus  gregarius 


7  DAT  USE 

RETURN  TO 

AGRICULTURE  UBRAR' 
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This  publication  is  due  on  the  LAST  DATE 
and  HOUR  stamped  below. 


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General  Library    . 
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