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ORREY BOTANICAL CL 


GENERAL INDEX. 


(Illustrated articles are designated by an asterisk * before the page number.) 


Antidromy of Plants, 379. 
Archegoniates, the Classification of the, 124. 


Botanical Notes, 182, 233, 275, 331, 370, 470. 

Botany of the Southeastern alted Cotes Stndies 
in the, *43, “Bs. 

Brinton, Dr. J. Bernard, Biographical Sketch of 
(with portrait), 93. 

Bryology, Contributions to American, *62. 

Buxbaumia aphylla, so. 


Calamagrostis scopulorum, Jones, 517. 
Characeae, Japanese, 68. 
Characeous Plants of Maine, Contributions to the, 


ee: dh 
Citations, Rules for, 130. 


Contributions to American Bryology, *62, *447. 
Contribution to the Characeous. Plants of Maine, 
397- 
ConTRIBUTORS: 
len, T. F., 68. 


Bailey, W. W., 470. 

ate a Hendley, 1, 394. 
Beringer, rge M., 93. 

Bicknell, Eugene P., 211, 351. 
Bolley, H. L., Sec’y, 408. 

Boyer, Charles S., 171, 260. 

Britton, Elizabeth G., 36, 62, 447. 
Britton, N. L., 220, 460, 470. 
Canby, William M., 162. ' 


Hill, E. J., 71. 
Hollick, Arthur, 225, 460. 
elliffe, Smith Ely, 274. 


Miller, A. W 
Olle, dea rw 

e +» 390. 
Peck, Chas. 3 oe ee 485. 
Pollard, Charles Louis, 231, 513. 
Rich, Wm. P., sr. 


Rydberg, P. A., 306. 
Schneider, bet 189, 494- 
Setchell, Albert, 341, 424. 


Small, John K., 43, 74, 365, 399- 

Tracy, S. M., 174. 

Trelease, Wm., 329, 331- 

Underwood, Lucien M., 124. 

Vail, Anna Murray, 25, 228, 458, 500. 

Ward, Lester F., 308. 
Cryptogamic Notes from Long Island, 274. 
Cyanophyceae, Notes on some New Rngland, 424. 


Description of a new problematical Plant from 
the Lower Cretaceous of Arkansas, *387. 

Desmids of the United States, Some new and 
rare, #289. 

Diatomaceous Deposit at St. Augustine, Florida, 
a Fossil marine, 171. 

Diatomaceous Deposit from an Artesian Well at 
Wildwood, N. J., 260. 


Eaton, Daniel Cady (with portrait), 341. 
Emory’s Report, On the two editions of, 394. 


Family Nomenclature, 1, 77, 216. 

Flora of Richmond County, New York. 
tions and New Localities, 460. 

Florida Plants, Notes on Some, 141. 

Food Piants of the North American Indians, 98. 

Fossil marine Diatomaceous Deposit at St. 
gustine, Florida, A, 171. 

Fungi, New Species of, 198, 362, 434, 485- 

Fungi, New Species of Parasitic, 174. 


Genus Cenchrus in North America, The, 298. , 

Genus Cracca, A Revision of the North American 
Species of, 25. 

Genus Galactia in North America, A Study of, 


500. 

Genus Sanicula in the United States with Descrip- 
tions of two new Species, *351. 

Genus Scouleria, a Revision of the, with Descrip- 
tion of one new Species, *36. 

Genus, Zenobia, The, 231. 

Grasses, New or Noteworthy American, 419, 463. 


Addi- 


511. 
Hybrid Oaks from the Southern States, Some new, 
% . 
H a boreale and Related Species, 2tr. 


Index to Recent Literature Relating to American ; 
Botany, 52, 89, 134, 185, 235, 283, 337» 374+ 
415, 442, 478, 519- 

Japanese Characeae, 68. & 

Juncus scirpoides and its Immediate Relatives, 
302. 

Lichens of Newfoundland and Labrador, An En- 
umeration of the, 2: 


_ Lichens, Some Special Phylogenetic Adaptations 
in, 
Lichens, the Biological Status of, 189. 


iv 


Malpighiaceaeand Zygophyllaceae,A Preliminary 
List of North American Species of, 228. 
Missouri Botanical Garden, 329. 


Nelumbo Lutea, Preliminary Notes on, *266. 
New Fungi, 362. 
‘New or Noteworthy American Grasses, 419, 463, 


517: 

New or Noteworthy North American Phanero- 
gams, 220. 

New Plants from Idaho, 48. 

New Plants from the Cretaceous of Kansas, De- 
scriptions of Three, *225, 

New Species of Fungi, 198, 434, 485. 

New Species of Parasitic Fungi, 174. 

New Species of Physalis, 306. 

New Species of Ustilagineae and Uredineae, 57. 

Nomenclature Question, The, 233, 308. 

Notes on some Cyanophyceae of New England, 


424. : 
Notes on some Florida Plants, 141. 
Notes on some southern Cassias, * 513. 


Oklahoma Plants, Observations upon some, 390. 
On the Carpets of Opulaster malvacea, 271. 

On the two editions of Emory’s Report, 394. 
Opulaster malvacea, On the Carpels of, 271. 


Papers read before the Section of Botany, A. A. 
A. 5., Springfield Meeting, 413. 

Personal Nomenclature in the Myxomycetes, 431. 

Phanerogams, New or noteworthy North Ameri- 


can, 220. 
Physalis, New Species of, 306. 


Pignuts, The, 331. 

Preliminary Notes on Nelumbo lutea, * 266. 

Proceedings of the Botanical Club, A. A. A. 5., 
Springfield Meeting, 408. 

Proceedings of the Botanical Society of America, 
Springfield, Meeting, 414. 

Proceedings of the Club, 51, 87, 133, 183, 234, 281, 
440, 477, 518. 


Redfield, John H. (with portrait), 162. 
Reviews, 78, 276, 331, 370, 400, 472. 
Rules for Citation, 130. 


Some special phylogenetic Adaptations in Lichens, 


494. 
Sinrnilien:; Vegetable, 465. 
Studies in the Botany of the Southeastern United 
States, * 43, * 365. 
Study of the Genus Galactia in North America, 


A, 500. 
| Systematic Botany of North Ameriza, The, 180. 


Teratological Notes, * 396, * 399." 
Tradescantia Virginica var. i qi. 
Tumble Mustard, 370. 

Two New Mountain Plants, 273. 

Two Undescribed Species of Phyncheath: 458. 


Ustilagineae and Uredineae, New Species of, 57. 
Vegetable Spiralism, 465. 
Young, Herbert A., 51. 


GENERIC INDEX. 


Abies, 211. 
Abutilon, 383. 
Acacia, 56, 99, 391- 
Acer, 52, 340, 367, 384, 460, 482. 
Acerates, 392. i 
Achnanthes, 261, 274, 275- 
Acnida, 288. 
Acolium, 259. 
Aconitum, 412, 474. 
Acorus, 466. 
Actinidia, 279. 
Actinocyclus, 172. 
—— 53- 

iantum, gI, 349- 
Adoxa, ae seit 


Aecidium, 60, 61, 136, 237, 284, 363» 364, 392s 443: 


Aeschynomene, 143. 
Aesculus, 219, 384. 
Aethusa, 460, 
Agaricus, 203, 204, 286, 489. 
Ageratum, 136. 
Agoseris, 186. 
Agrimonia. 460. 
Agropyrum, 518. 
Agrostis, 138, 463, 464. 
Atra, 511. 
Akebia, 279 
Alectoria, 113, 242. 
Alliaria, 282. 
Allionia, 223, 224, 284. 
Allium, 53, 173- 
Allocarya, 186, 416. 
Alnus, 461. 
7. pa, 405. 

A, 3372 479- 
rg aging 395, 483- 


Anabaena, 427, °. 
inten 
Andromeda, 153, 231, 232; 437- 
Andropogon, 143, 145, 146, 437- 


a e, 412. 
ngioridium, 432, 433- 


» 460. 
Anthurium, 480 
_ Anthurus, 337. 
ne Anychia, 170. 
_ Apera, 405. 
_ Aphaerena, 137. 

en i ieee 


Apios, ror. 

Aplopappus, 186. 
Aporophallus, 23%. 
Aquifolium, 3. . 
Aquilegia, 91, 236, 460, 473- 
Arabis, 137, 475- 

Arachis, 136, 281, 445. 
Aralia, 225. 

Archilejeunea, 287. 
Arctium, 417. 
Arctostaphyllos, 120, 40:. 
Arcyria, 333- 

Ardisia, 483. 

Arenaria, 362. 

Arethusa, 445- 

Argemone, 392; 475- 
Argylia, 285. 

Arisaema, 106 

Aristida, 135; 143, 145- 
Aristolochia, 54. 

Armeria, 405. ~ 

Arnica, 57- 

Arthothelia, gt. 
Arthrodesmus, 296, 297, 416. 
Arthonia, 258. 

Arthronia, 91. 

Arthrospira, 430, 431, 482. 
Artotragus, 337- 

Arum, 467. 

Asarum, 170. 

Aschersonia, 436, 440. 
Asclepias, 134, 137, 1432 153+ 154, 238, 392- 
Asimina, 143, 474- 
Aspergillus, 210, 286. 
Aspicarpa, 228. 

Aspidum, 53, 137» 169, 179, 284, 322, 350 406, S2z- 
Aspilia, 337- 

Asplenium, 91, 179, 35°- 
Aster, 59, 138, 182, 273, 284, 375, 391- 
Asterella, 140. 

Asteridium, 176, 177, 288, 479- 
Asterina, 139, 2372 435- > 
Asterionella, 275- 
Asteroma, 237. 

Asterula, 82, 237- 
Astragalus, 54, 284, 338, 284. 
Atta, 413. 

Aulacodiscus, 261. 

Auliscus, 261. 

Azalea, 282, 437, 461. 
Azolla, 441, 462. 


Baccharis, 391, 392- 
Badhamia, 334- 
Baeomyces, 252- 
Baileya, 395- 
Balsamorrhiza, 110. 


Baptisia, 45, 238- 
ula, 442- 

Batemaria, 417. 

Battarrea, 208, 286. 


 Bejaria, 145. 


Berberis, 135, 474- 
Berchemia, 147- 


Berlandiera, 143. 
Betencourtia, sor. 
Betula, 92, 338, 417. 
Biatora, 252, 253, 254, 375. 
Biddulphia, 173, 261, 275. 
Bigelovia, 375. 
Blumenavia, 238. 
Bocconia, 383. 
Boehmeria, 140. 
Boerhaavia, 156. 
Borsduvalia, 375. 
Bolelia, 338. 

Boletinus, 206, 286. 
Boletus, 207, 286, 477. 
Bonapartea, 158. 
Boretta, 406. 
Botridium, 402. 
Botrychium, 170, 444. 
Brachiolejeunea, 287. 
Bradburya, 324. 
Brasenia, 474. 
Brassica, 224, 278, 413. 
Braya, 285. 
Brebissonia, 261. 
Breweria, 143, 144, 154, 155, 238. 
Brintonia, 338. 
Brissonia, 25. 
Brittonamra, 25. 
Brodiaea, 114. 

Bromus, 512. 
Brousonnetia, 461. 
Bruchia, 350. 
Brugmansia, 481. 
Bryophylium, 382, 384. 
Bryum, 442. 

Buda, 405. 

Buechnera, 393. 
Buellia, 256, 257, 258, 260, 
Buettneria, 24. 
Bumelia, 144, 392. 
Bursa, 407. 
Buthograptus, 56. 
Buxbaumia, 50, gl, 350. 
Byrsonima, 229. 


Cabomba, 138. 
Cactus, 116. 

Caeoma, 185, 210, 286. 
Caffea, 381, 384, 
Calamagrostis, 517. 
Calicum, 259, 493. 
Calla, 106. 
Calliprora, 416. 
Callithamnion, 52. 
Callithamnopsis, 56. 


Calymperes, 339, 442. 
Calycanthus, 24. 


Capnoides, 405, 406. 
gee : 
Capsella, 376, 407, 417. 
pine orc 147. 
Capsovia, 427. 
Cardamine, 1 


ma 147, 148 
Cardiospermum, 147, S 
-Carduus, 324, 446. : 


Carex, 58, 82, 220, 221, 222, 223, 267, 273, 284, 375, 


vi 


Carpinus, 377. 

Carum, 108, 109. 

Carya, 331. 

Cassandra, 231. 

Cassava, 438. 

Cassia, 219, 392, 513, 514, 515, 516. 
Cassiope, 231. 

Castalia, 147, 286, 407, 473. 
Castanopsis, 377. 

Catasetum, 520. 

Catopteris, 168. 

Caulerpa, 402. 

Cavendishia, 92. 

Ceanothus, 143, 186, 393. 
Celastrus, 468. 

Celtis, 147. 

Cenchrus, 298, 299, 300, 301, 416. 
Centaurea, 461. 

Centrosema, 324. 
Cephalanthus, 151, 392. 
Cerataulus, 261. 

Ceratiola, 144. 

Ceratiomyxa, 235, 283. 
Ceratomyces, 140, 
Ceratophyllum, 138. 
Cercocarpus, 135. 

Cercospora, 82, 178, 179, 288, 438, 479. 
Cereus, 116, 478, 482, 484, 522. 
Cetraria, 240, 241. 
Chaetocladus, 56. 

Chalara, 285. 

Chamaedaphne, 231. 
Chamaeraphis, 422. 
Chapmania, 143. 


Chara, 52, 68, 79; 71, 138, 398, 427» 43°, 444, 468. 


Chaunostoma, gz. 
Cheilanthes, 350. 
Chimaphila, 224, 225. 
Chloris, 423, 424, 481. 
Chlorogalum, 114. 
Chondrioderma, 332, 334. 
Chondrophora, 338. 
Chrysobalanus, 438. 
Chrysoma, 186. 
Chrysopogon, 58, 59. 
Chrysopsis, 441 
Chrysothamnus, 338, 375- 
Cicendia, 405. 
Creuta,:238, 

Cimicifuga, 135, 474- 
Cirsium, 324. 


Cladium, 6r. 

Cladonia, 191, 194, 250, 251. 
Cladop! 430. 

Clados 


Clematis, 278, 473, 474, 493. 
Cleome, 238, 475. 


Clitocybe, 199, 286, 
Clitoria, 392, 500. 
Cnicus, 110, 143. 
Closterium, 291, 207» 427- 
Cocconeis, 261, 275. 
Colmil, 25. 
rye oae » 500, 501. 
ollema, 192, 197, 198, 246. 

Colletotrichum, 135, 374, 4375 479- 
Collen? 186, 285, 520. 

ia. 286, 486. 
ie 
Sologania, soz. 
Colubrina, 53. 
Colus, 238. 


Commelina, 143, 160, 238 437- 
Completoria, 337- 
Conomitrium, 350. 
Conopholis, 234. 461. 
Convallaria, 384. 
Convolvulus, 154, 395, 461. 
Cooperia, 391. 

Coprinus, 205, 206, 286, 491. 
Cordia, 483. 

Corema, 169, 171. 

Coreopsis, 47, 48 92, 147, 481- 
Coronilla, 460, 

Coronopsis, 324- 

Coronopus, 407- 

Corticium, 210. 

Cortinarius, 203, 286. 
Corydalis, 405, 475- 

Corylus, 339, 416. 
Corynephorus, 407. 


Coscinodon, 277, 499, 447» 448» 449- 


Cosinodiscus, 173, 261, 262. 


Cosmarium, 238, 299, 292, 293» 294» 296, 297,298,416. Ephemerum, 67, 442. 
6 


Cosmocladium, 296, 298, 410- 
Cosmos, 521. 
Costaria, 139. 
Covillea, 229, 288. 
Ctenium, 423- 
Cucurbita, 100, 415. 
Cuscuta, 339, 417- 
Cycas, 384. 
Cycloloma, 392. 
Cycloteila, ie. 
Cymatopleura, 275- 
Cymbella, 262, 274- 
Cymopterus, 110. 
Cynanchum, 337- 
Cynodontium, 277. 
Cynoglossum, 461. 
Cynosciadium, 391- 
Cyperus, 44, 60, 115, 161. 
Cypressus, 376. 
Cypripedium, 462. 
Cyrtodeira, 336- 

‘ Cyrtopodium, 99, 519- 
Cystopleura, 262. 


Daimonelix, 83, 89, 186. 
Danthonia, 469. 
Daphnopsis, 521. 
: oe gt. 
rluca, 437, 479° 
ee ts 
Deanea, 415. 


Delphinium, 53, 375; 383» 474» 489- 


Desmanthus, 391, 392- 
Desmonema, 428. 
Diachea, 334. 
Dianthus, 219. 
Dicentra, 475- 
Dichomyces, 140. 


Dicranella, 411, 449» 45, 451, 452- 


Dicranodontium, 451 
Dicranum, 415, 45° 451- 
Dictydium, 332. 
Dictyophora, 238. 
Se omimeer 333- 
idymosphaeria, 439, 479- 
Dieffenbachia, tt od és 
Digitaria, 419, 42°. 
Dimerosporium, 175, 237, 288- 
Diodia, 151. 
Dioscorea, 412. 
Diospyros, 178, 339- 
Diplazium, gt- 
Diplodia, 177, 178, 288. 


Vii 


| Dothidella, 237. 

, Dryopteris, 322, 405, 462. 
| Drosera, 159. 

| Dysphinctium, 292- 


Eastwoodia, 53. 

| Eatonia, 511. 

Eccilia, 201, 286. 

| Echinocactus, 287. 
Echinopsis, 91. 

| Echinospermum, 324- 
Ectocarpus, 402- 

| Elaeagnus, 122. 

Elodea, 24, 138, 405» 406. 
| Elymus, 517, 518. 


Encyonema, 262, 274- 
Endocarpon, 196, 259- 
Enslenia, 238. 

| Entoloma, 200, 201, 286, 489. 
Entyloma, 57, 136- 


Ephestia, 416. 
Epidendrium, go. 
Epigaea, 236. 
Epilobium, 131, 138, 330- 
Epipactis, 405. 
| Epithemia, 275- 
| Equisetum, 377, 389, 468- 
Eragrostis, 287, 495- 
Eranthis, 475- 
Erianthus, 419, 481. 
Erigeron, 137, 186, 237, 238, 416. 
Eriodictyon, 482. 
Eriogonum, 143, 391. 
Eriophorum, 462. 
Eriophyllum, 416. 
Erodium, 468, 449- 
Ervum, 5°3, 5°7- 
Erycina, 9°. 
Eryngium, 147, 237- 
Erysimum, 285. 
Erythrina, 135, 437- 
Erythronium, 230, 415- 
Euasplenium, gt. 
Euastrum, 292, 297- 
Eucantharomyces, 149. 
Euglena, 402. 
Euglenopsis, 136. 
Eunotia, 262, 275- 
Euoaia, 174. 
Eupatorium, 134, 136, 337- 
Euphorbia, 393, 439- 
Euphrasia, 179- 
Eupodiscus, 173, 275- 
Evernia, 241- 
Excipulina, 209 , 286. 
Exoascus, 238, 339- 


Fagonia, 229. 
Fagus, 187, 287- 
Falcata, 121. 
Faxonia, 53- 
Festuca, 362- 
Fimbristylis. 145- 
Fissidens, 54, 442- 
Flammula, 202, 286, 489. 
Floerkea, 237 
Flukigeria, 55- 
Fomes, 362; 443- 
Forestiera, 53- 
Forsythia, 467- 
Fothergilla, 52t- » 
Fragaria, 219. 
Fragilaria, 275- 
Fraxinus, 153, 461- 
Frittellaria, 237- 
Froelichia, 392- 
Fucus, 402. 
Fuligo, 432, 433- 
Funarta, 469- 
Fusarium, 414- 


| Encalypta, 452, 453» 454) 455 455, 457+ 458. 


Vili 


Galactia, 391, 438, 500, 50T, 502, 503, 504, 505, 506, | Hyalodiscus, 173, 263. 


507, 508, 509, 510, 511. 
Galega, 25, 27, 30, 31, 32, 33, 34, 35. 
Galinsoga, 55. 

Galium, 477. 

alphimia, 228. 
Garberia, 152. 
Garovaglia, 442. 
Gaultheria, 120, 461. 
Gaura, 391, 392. 
Gaylussacia, 120. 

nista, 54. 
Geranium, 89, 469. 
Gerardia, 461, 393. 
Gilia, 338. 
Ginko, 219. 
Gloeosporium, 185. 
Gloeotrichia, 450. 
Glossopetalon, 1 37- 
Glyceria, 405, 406, 512. 
Glycine, 503. 
Glycyrrhiza, 108. 
Gnaphalium, 46r. 
Gomphonema, 262, 263, 274. 
Gomphrena, 288. 
Gonatozygon, 291, 297. 
Gonioptecis, 137. 
yera, 405, 406. 

Goniothecum, 173. 
Sordoatn, 438. 

rammatophora, 173, 275. 
Graphis, 258. ebts 
Grevillia, 435. 
Grimmia, 36, 37, 41, 435 448. 
Guajacum, 229. 
Gymnogramme, 337- 
Gymnostomum, 277. 
Gymnosporangium, 188, 
G YS, 405, 406. 


Habenaria, 53, 15 » 186, 462. 
Halenia, pr fag é .f 
Halicteres, 468. 
Hamamelis, 412. 
Haplomitrium, 402. 
Haplosporella, 82, 449, 479. 
Hariota, 139. : 

lia, 428. 
Hebeloma, 202, 286, 
Hedysarum, 508, 
Heimatomyces, 140, 


Um, 147. _ 
Helianthella, 142, 152. ; 
Helianthemum, 147, 148, 238. 
Helianthus, 100, IOT, 103, 393, 444, 479- 


Hendersonia, 178, 288, 436. 
a, 


| 


ydrocoleum, 429. 
Hydrolea, 154. 
Hydrosera, 26 35 374. 
Hydrothyria, 197, 496. 
Hygrobiella, 188. 
Hygrophorus, 200, 211, 286, 486, 487. 
Hymenophyllum, 350. 
Hymenophytum, 188. 
Hyophila, 442. 


Hypericum, 46, 148, 211, 212, 213, 214, 215, 283. 


Hypholoma, 204, 286, 490. 
Hypnum, 339. 
Hypocrella, 237. 

Hyptis, 89. 

Hystrix, 518, 


Tex, 3, 176, 177. 
Ilicium, <7, Ss, 
Impatiens, 383, 385, 441. 
Indigofera, 392. 
Inocybe, 488. 

Ipomoea, 461. 

Iris, 89, 178, 187, 375, 386, 416, 467. 
Isactis, 426, 

Isariopsis, 438, 479: 
Ischaemum, 175. 
Isoetes, g2. 

Isopyrum, 413. 
Isotachis, 287. 

Itayahya, 238. 
Ithyphallus, 238. 

Iva, 521. 

Ixophorus, 422, 423. 


Jacaratia, 92. 

Jamesoniella, 188. 

Janusia, 228, 

Jatropha, 143, 188, 393. 
Jeffersonia, 442. 

Joosia, 469. 

Jubaea, 522. 

Juglans, 483. 

Juncoides, 405, ‘ 
7 


3302 415, 416, 467. 
Jungermania. 
Juniperus, 118. 


Kallstroemia, 230. 

Kalmia, 377, 413, 461, 482. 
Kantia, 237. 

Koellia, 368. 

Kraunhia, 324. 

Ki istera, 135, 137, 149 324» 392- 


Laboutbenia, 140. 
Lachnoca 


ja, 176, 288, 


i aE: 
Larrea, 56, 229, 230, 288, 
Lastrea, 137, 406, 
Lathyrus, 139, 399- 
Laternea, 233. 


Lecanactis, 2:8, 


Lentibularia, 219. ‘ 
Lepidium, 53° 137; 272, 385. 


JUNCUS, 44, 45, 92, 139, 145, 281, 302, 303, 304, 30 


Lepidodendron, 470. 
Lepigonum, 325, 405. 
Lepiota, 198, 199, 286. 
Lepra, 198. 
Leptochloa, 175. 
Leptogium, 197, 246, 495- 
Leptoglossum, 210, 386. 
Leptonia, 201, 286. 
Leptorchis, 405. 
Leptotherium, 82. 
Lesquerella, 475. 
Leucobryum, 442. 
Lencodictyon, sor. 
Leucothoé, 146, 231, 232- 
Lewisia, 111. 
Liatris, 143, 152, 461. 
Ligusticum, 363. 
Lilium, 170. 
Limodorum, 145, 158. 
Limonium, 405, 406. 
Limosella, 519. 
Linanthus, 186, 375. 
Linum, 393- 
Liparis, 405. : 
Liquidambar, 467. 
Liriodendron, 54, 285, 376, 383, 415. 
Liriodendropsis, 415. 
Liriophyllum, 54. 
Lissocarpa, 521. 
Lithophragma, 338. 
Lithothamnion, 285. 
Litobrochia, gt. 
Litsia, 440. 
Lepargyrea, 324. 
Lobelia, 383, 484. 
Lophopappus, 55- 
Lophotocarpus, 287. 
Lotus, 469. 
Ludwigia, 392, 393- 
Lupinus, 108, 186, 325- 
Lunaria, 310. 
pees 4°. . 
ycogala, 333, 334- 
Lycoperdon, 107, 209, 286, 477- 
Lycopersicum, 47. 
Lygodesmia, 143- | 
Lygodium, 170, 481. 
Lyngbya, 285, 428, 429- 
Macrocarpaea, 483. 


Macromitrium, 442. 
Macrosporium, 493. 


9 
Magnolia, 146, 176, 278, 286, 439, 449, 474- 


Mallinoa, 136. 
Mallotium, 197, 496. 
Malpigia, 229. 
Malvaviscus, 446. 
Mammillaria, 116. 
Manihot, 136. 
Marasmius, 487. - 
Marattia, 135- 
eg 405, 406. 
arsilia, 161, 283, 393- 
Masdevallia, at ” 
Maxillaria, 417. 
Maytenus, 483. 
Medeola, 114. 
Medicago, 385, 460. 
Megapterum, 393- 
Melasmia, 209, 286. 
Meliola, 237. 
Melioloa, 434, 438. 
enamel 483. 
ogramma, 82, 440, 479- 
Melosira, 173, 263, 275- 
Melothria, 151. 


IX 


| Metzgeria, 136. 
| Meum, 405. 
| Mibora, 405. 
| Micrasterias, 292, 297- 
Microchaete, 427. 
Microcoleus, 427 
Microseris, 111, 338- 
Microsphaera, 52. 
Mikania, 446. 
Mimosa, 413. 
Mimulus, 90, 185, 237, 285- 
Mirabilis, 135 
Mitchella, 141. 
Mitella, 137 
Mnioides, 406. 
Mnium, 414. 
Mocinna, g2. 
Monilea, 135. 
Monilia, 238. 
Monniera, 46, 47, 92, 393- 
Monnina, 236, 479- 
Monoblepharis, 483. 
Montanoa, 136. 
Mormodes, 480. 
Morongia, 143. 
Mosquitoxylum, 483. 
Mongeotia, 445- 
Muhblenbergia, 135. 
Musa, 383. 
Musineon, 337. 
Mutinus, 238. 
Mycena, 199, 200, 286, 486. 
Mylittopsis, 377- 
Myosotis, 383- 
Myosurus, 474- 
Myriophyllum, 138. 
Myrsine, 436, 483. 


Naias, 138. 
Nasturtium, 405, 475- 
Naucoria, 486. 
Navicula, 173, 263, 264, 274- 
Neckeria, 405. 
Needhamia, 25. 
Neillia, 271. 
Nelumbium, 324, 384. 
Nelumbo, 102, 120, 266, 270, 271» 324» 376. 
Nemophila, 479, 519- 
_ Neogoezia, 519, 529. 
Neosyris, 375 
Nephroma, 245- 
Nephromium, 194, 497- 
Nicotiana, 104, 105, 383, 443- 
Nidulariopsis, 416, 481. 
Nitella, 68, 69, 79> 71, 85, 134, 398, 402- 
Nitzschia, 264, 275- 
Nolina, 158, 238- 
Nostoc, 197, 428, 429- 
Nostochopsis, 427. 
Notothylas, 138. 
Notylia, 417. 
Nummularia, 362. 
Nuphar, 325, 497- 
Nymphaea, 120, 147, 407, 460, 473- 


Obolaria, 413- 
Octoblepharum, 442. 
Odonia, 500. 

Odontidium, 265. 

Oenothera, 151, 339, 381 383- 
Oligonema, 333- 

Olpidium, 185- 

Omphalia, 200, 286. 

Onagra, 392, 460. 

Onoclea, 235, 283, 412- 


| Opulaster, 271, 272, 376, 460. 
Opuntia, 115, 116, 393» 417» 428- 


Orontium, 102, 106. 

Orthaea, 521. 

Orthotrichum, 52. 

Osbertia, 186. 

Osmunda, 288. 

Ostryra, 377. 

Oxalis, 45, 56, 139, 336, 338, 519. 
Oxycoccus, 407. 

Oxytropis, 405. 


Pachyma, 107. 
Paleohillia, 387, 388, 390. 
Palmella, 402. 

Panaeolus, 205, 286. 
Panax, 138. 

Panicularia, 405, 406, s12. 
Pannaria, 246, 375. 


Panicum, 137, 175, 301, 419, 420, 421, 422, 423,435, 


441, 462, 451. 
Parmelia, 191, 194, 242, 243, 244, 407. 
Parodelia, 2 37. 
Paronychia, 156. 
Parosela, 392, 393. 
Parrotia, oY fae 
Paspalum, 175, 420. 
Passiflora, 104, 392. 
Paullinia, 522. 
Paulownia, 467. 
Pavonia, 480. 
Pediastrum, 140. 
Peganum, 231. 
Pelargonium, 383, 469. 
Pelexia, 417. 


Peronospora, 364, 443. 
Sa 144, 156, 157, 238. 
usaria, 249. 
Pestalozzia, 177, 178 288, 
Petalostemon, 324. 


US, 99, 100, 460. 


X; 541 339, 480. 
ile 8 
n, 366. 
Phormidium, 427, 430°. 
Phyllachora, 210, 237, 449, 479. 
Phyllocactus, 188. 
Phyllospadix, 53. 


x 


Placodium, 246, 247. 

| Plagiogramma, 265. 

| Plantago, 46r. 

| Platanus, 226. 
Pleurosigma, 173, 265, 274. 

| Pleurotaenium, 290. 

| Pleurothallis, 90, 376, 417. 

| Plowrightia, 135. 

| Pluchea, 54, 174, 392. 

| Pluteolus, 203, 286, 
Pluteus, 488 

| Pneumaria, 407. 
Poa, 138, 465, 511, 517. 
Podosira, 173. 

| Podostigma, 145. 

| Polychidium, 197, 496. 
Polygala, 236, 479. 

| Polygonatum, 405. 


| Polygonum, 56, 139, 235, 280, 281, 383. 


| Polymnia, 136 
Polymyxus, 261, 265. 
| Polanisia, 144. 
| Polyplocium, 492. 
| Polyporus, 207, 286, 409, 418, 493. 
| Polypodium, 137, 146, 444, 520. 
| Polystichum, 405, 406. 
Populus, 131, 286, 462. 
| Porlieria, 229. 
Portulaca, 143. 
| Potamogeton, 142, 374, 393, 462. 
| Poterium, 405, 406. 
| Pottia, 442. 
Pottiariparia, 469. 
_ Primicorallina, 56. 
| Prionodon, 339. 
| Prochynanthes, 442. 
_ Prosopsis, 120, 121, 363, 385. 
| aad et 198, 402. 
| Protophyllum, 227. ‘ 
Pome, 238. 
| 519- 
| Psathyrella, 82, 490. 
| Pseudauliscus, 265. 


| Pteris, g1, 115, 210. 

| Prerocaulon, 143. 

| Pterospermites, 227. 
| Priloria, 87. 

| Puccinellia, 512, 512. 


| Puccinia, 58, 59, 60, 61, 136, 174, 185, 237, 284, 288, 


t 363> 435» 442, 443. 

_ Punctaria, 52. 

| Pycnanthemum, i 
Pyrenula, 191, 198, 260. 

| Pyrola, 461. 

| Pyrrocoma, 186, 

| Pyrus, 278. 

| Pythium, 235, 283. 


| 
| Quercus, 74, 75, 76, 89, 92, 118, 11g, 137, 139, 140, 
| 142, 146, 178, 330, 339, 365, 367, 377, 383, 392, 


396, 4375 443, 462, 519. 


Radiofilum, 140. 
| Railardella, 186. 
| Ramalina, 240. 

Ramisia, 521, 
| Ramularia, 57. 


| Raveoelt 63, 36 
| Ravenelia, 90, 132, 23 » 364, 443. 
| Rebusta, tt 3?» 237s 363, 364, 443. 


» 82. 
Rhaphoneis, 173, 265. 


| Prunus, 103, 132, 219, 237 278, 283, 286, 339, 442, 


xi 


Rhexia, 145, 150, 238. | Sisyrinchium, 375, 385- 
Rhizophidium, 52, 283. | Smilax, 113, 114, 144, 1465 179. 
Rhizosolenia, 174, 275- | Solanum, 112, 238, 279, 285, 412, 413, 481, 521. 
Rhododendron, 287. Solidago, 145, 147, 152, 182, 338, 368, 369, 393, 461, 
Rhoikosphenia, 274. | 519. 
Rhus, 54, 285, 338) 339, 392, 4175 443» 444, 482. | Solorina, 245, 497. 
Rhynchosia, 143, 149, 150, 238, 458, 459- | Sorosporium, 363, 443- 
Rhynchospora, 144. | Sparassis, 207, 208, 286. 
Rhynchostegium, 469. | Spathyema, 106. 
Ribes, 54, 91, 136, 137, 285, 338. | Specularia, 192,405, 406. 
Riccia, 402. | Spergularia, 324, 325, 405- 
Richardia, 15t, 382. | Spiesia, 138, 405, 406. 
Ricinus, 383, 469. Spiraea, 237. 
Rinodina, 249. | Spiranthes, 158, 380, 405, 406. 
Rivularia, 426, 427, 482. | Spirogyra, 185. 
Roripa, 186, 187, 405, 460. | Sphaeralcea, 364. 
Rosa, 122. Sphaerella, 82, 176, 288. 
Rosellinia, 439, 479- | Sphaerophorus, 259. 
Rubus, 150, 460, 471. | Sphagnum, 92, 344, 346, 442, 443- 
Rudbeckia, 139, 378- | Sporobolus, 55, 143, 463, 464, 405, 518. 
Rumex, 44, 92, 137, 237» 33° 367) 444» 480, 482. Sporotrichum, 237. 
Ruppia, 426. | Stachys, 177, 461. 
Russula, 477. | Statice, 405, 406. 
Staurastrum, 294, 295, 297, 298- 
Sabal, 146, 147. Stauroneis, 265, 274. 
Sabbatia, 393. Stemona, 3. 
Sagedia, 259. | Stellaria, 171, 324, 405. 
Sagittaria, 105, 106, 287, 330, 364. | Stenophyllus, 143, 161, 238. 
Salix, 90, 383, 385, 462, 482, 521. | Stenorrhynchus, 158. 
la, 54, 92, 135, 130, 138, 187, 188, 417,466, 481. | Stenosiphon, 393. 

Salvinia, 462. Stephanodoria, 186. 
Sambucus, 151, 444. | Stephanogonia, 174. 
Samolus, 137. | Stephonopyxis, 174. 

* Sanguinaria, 89, 287. Stereocaulon, 249. 
Sanguisorba, 405. Stereum, 82. 
Sanicula, 237, 35t» 352s 353» 354» 355» 356, 3579 358, | Sticta, 245, 495s 496, 497» 498, 499» 509. 

359, 360, 361, 442. Stictina, 495, 496, 497, 498. 

Saracha, 483. Stictodiscus, 174. 
Sassafras, 177, 225, 226, 283. Stigonema, 444. 
Saussurea, 350. Stillingia, 143, 392- 
Savastana, 405, 406. Stipa, 423, 49, 481. 
Saxifraga, 58, 137- Stipulicida, 148, 238. 
Scabiosa, 171. Stropharia, 204, 286, 489- 
Scaevola, 436. Streptanthus, 338. 
Scenedesmus, 427, 439. Surirella, 265, 275, 374- 
Schinus, 186. Sweetia, 500. 
Schistidium, 65, 66. Symploca, 428. 429, 439. 
Schizaea, 169, 441. | Synchytrium, 89. 

- Schizophyllum, 439, 479- | Synedra, 265, 275- 
Schizostega, 478. | Syntherisma, 420. 
Schizothrix, 429. | Syrrhopodon, 442. 
Schoenocaulon, 439- Systegium, 442. 
Scholecotrichum, 178, 288. : 

Schollera, 497 461. Tabellaria, 275- is 
+ 30s 37s 38, 39+ 49, 41, 425 43» 135- Talinum, 107, 39%. 
Scirpus, 115. Taphrina, 178, 286. 
Scirrhia, 435. Taraxacum, 375- 
Sclerotina, 135. Taxodium, 92. 
se er open 446. Tayloria, 442. 
Scutellaria, 54 Tecoma, 177, 178. 
Scytonema, 428. Tephrosia, 25, 27, 28, 29» 30, 31, 32, 33» 34» 35» 51¥- 
Secotium, 492. 4 Terpsinoe, 265. 
Securidaca, 479. Tessela, 275. 
Sedum, 287. Tetracyclus, 266. 
ers 259. ‘Tetraedron, 140. 
enipedium, 520. Tetraploa, 179, 288. 
Senebiera, 324, 407. Teucrium, 54, 147- 
Senecio, 137, 186, 337, 375, 376- Thalictrum, 53, 474- 
onium, 333- Thamnolia, 194, 252- 
Septoria, 192, 437, 479- Thelephora, 492. 
4 Sequoia, 186, 375. Theloschistes, 242. 
Sericocarpus, 461. Thelotrema, 249- 
Sesuvium, 391, 392- Thermopsis, 186. 
. Setaria, 161, 175, 422, 423- Thryallis, 228. 
herdia, 121, 122, 324. Tiarella, 58. 
Sida, 137. Tibouchina, 339- 
Sigmatella, 442. Tilia, 278, 460. 
Sieglingia, 365, 407, 482. Tillaea, 138. 
Silene, 57, 460. Tillandsia, 158. 
Silybum, 405. Tilmadoche, 432, 433- 
Sinapis, 224. Tipularia, 462. 
Sisymbrium, 370, 385, 443, 477 Tissa, 186, 324, 405. 


Tolypothrix, 428. 
Townsendia, 349. 
Trabutia, 237. 
Tradescantia, 71, 72, 73, 1375 143, 147, 462. 
Trematodon, 442. 
Trentepohlia, 188. 
Tribulus, 230, 23r. 
Tricamphora, 332. 
Triceratum, 174, 263, 266, 275. 
Trichia, 333. 
Tricho aena, 238. 
Tricholoma, 485. 
Trichomanes, 479. 
Trichostema, 54, 445. 
Trifolium, 53, 139, 186, 219, 409, 519. 
Triglochin, 375. | 
Trilisa, 145. 
Triodia, 407. 
Tripsacum, 420. 
Teena 

thelium, 259. 
Tsyblidium, 237. : y 

a, 43, 44, 207, 209, 442, 467. 

Tubsriat 202, 286, oh 
Tubulina, 333. 
Tussilago, 465. 
Tylostoma, 209, 286. 
Typha, 467. 


Udora, 24,374, 405. 406, 462. 
leomyces, 237. 
Ulex, 460. 
Ulmaria, 52. 
Ulmus, 147. 
Uncinula, 413. 
Unona, 446. 
Umbellularia, 340. 
Umbilicaria, 244, 245. 
Urceolaria, 249. 
tenon Nie 36 8 
©, 55» 2375 416, 435, 436, 437, 438, 479. 
Urochloa, 422, i al 
Uromyces, 57, 136, 237, 284, 436. 
241. 


snea, 
Ustilago, 57, 136, 175, 288, 237, 362, 443, 481. 
Utricularia, 52, 146, 155, 156, 478. 
Vaccinium, 54, 120, 153, 285. 

Valeriana, 112, 460. . 

Valsa, 210, 211, 286, 

Vi a, 142, 234, 469, 470. 


Xii 
bod 


| Vanilla, 417. 
| Vascoucellia, 92. 
| Vaucheria, 339. 
| Veratrum, 185, 384. 
| Verbascum, 383. 
| Verbena, 137. 
Verbesina, 136. 
| Vermicularia, 177, 288. 
| Vernonia, 136, 143. 
Veronica, 46r. 
Verrucaria, 198, 259. 
Vicia, 186, 522. 
Victoria, 147. 
Vilfa, 463, 464. 
Viola, 286, 313, 377,475, 484. 
Vitis, 104, 226, 237, 283, 330. 
Vittaria, 147. 
Volutella, 82, 337. 
Volvaria, 487, 488. 
Volvox, 402. 


Weingaertneria, 407. 
Weldenia, 186. : 
Whipplea, 364, 
Willoughbaeya, 16. 
Wistaria, 324. 
Wolffia, 462. 
Woodsia, 349. 
Wyethia, 110. 


Xanthidium, 295, 296, 297. 
Xanthium, 417. aided 
Xanthorrhiza, 474. 
Ximenia, 141, 144. 
Xolisma, 153. 

Xylaria, 237. 
Xylographa, 258. 

Xyris, 159, 160, 238. 


Yucca, 119, 158, 288, 330, 377, 392, 440. 


Zamia, 107. 
Zea, 99, 520, 
Sensula 231, 232, 286. 
Zephyranthes, 287. : a 
Zinnia, 395, 412. 
izania, 53. 
Zizia, 60. 
Zygodesmus, 82. 


Zygophyllum, 230. 


VoL. 22. JANUARY, 1895. No. 4. 


BULLETIN 


OF THE 


TORREY BOTANICAL CLUB. 


A MONTHLY JOURNAL OF BOTANY. 


EDITED BY 
‘NATHANIEL LORD BRITTON, 


* AND OTHER MEMBERS OF THE CLUB. 


CONTENTS: 


PAGE, PAGE, 

Family Nomenclature: }vAn Hendley Barn- | Deby gn 5 a ee ee ee a ae ele 

MOO SS LS ee tees FT New Plants from Idaho: Lomis F. Hender- : 

A Revision of the North American Species of | SOM, we ek ee ER Seen ee 48 

the Genus Cracca: Anna Murray Vail, 25| Burbaumia aphylla; Geo. GC. Kennedy, 50 

_ A Revision of the Genus Sceuleria with De- | Herbert A. Young: Wm. P. Rich, 2... 5% 

scription of one new Species: Efizabeth G. . PROceEDINGS OF THE CLUB... . ce 
detitem (Yate ate) os is ae 36 Inpex To recent LirERATURE RELATING 


Studies in the Botany of the Southeastern | ToAmeRICANBorany .-.-+... 52 
United States—Ill: F¥okn K. Smeadl (Plate | 


PUBLISHED FOR THE CLUB. 


“Tan Naw Ena Pennine House, 


REY BOTANICAL CLUB. 
OFFICERS FOR 1895. 


| President, 
HON. ADDISON BROWN. 


Vice Protienté, 
T. F. ALLEN, M. D. L, H. LIGHTHIPE. 


, Recording Secretary, Corresponding Secretary, 
HENRY H, RUSBY, M. D., JOHN K. SMALL, 
College of Pharmacy, New York City. Columbia College, New York City. 


Editor, een Oy hee ‘Treasurer, 
N L. BRITTON, Ph. D, HENRY OGDEN, 
Columbia College, New York oF sere 1 Pine Street, New York City, 


f 
i 


BULLETIN 


OF THE 
~ TORREY BOTANICAL CLUB 
Vol. 22. Lancaster, Pai Sintacs 15, 1895. No. 1. 
Family Nomenclature, i 


By JoHN HENDLEY BARNHART. 


Although for over a hundred years botanists have recognized 
certain natural groups of plants, variously called “orders” or 
“families,” the naming of these groups has been full of incon- 
sistencies, and subject entirely to the caprice of each writer. 
When we come to consider the fact that no author has ever con- 
sistently followed any rule in naming such groups (some have 
even called the same family by two or more different names in 
the same work) it seems strange that the present confusion is 
no greater than it is. 

In spite of the fact that there are no rules, there is a marked 
tendency toward the use of uniform terminations in the naming of 
all groups of codrdinate rank higher than genera. In the case of 
family names this tendency has shown itself by the extension of 
the use of the termination ‘“-aceae,” until this has become uni- 
versally recognized as a distinctive mark of family rank. Yet 
some of the usual names are improperly formed from the generic 
root upon which they are based, while quite a number of the 
families retain names which aresnot founded upon genera at all. 

This latter class requires special consideration. A generic 
name stands or falls with its typical species, and why should not a 
family name stand or fall with its typical genus? Such names as 
Umbelliferae, Cruciferae, Leguminosae, Labiatae, Gramineae, Com- 
_ positae, etc., are not named after genera. Nor do these names 


2 


express characteristics peculiar to the families to which they are 
applied. The Araliaceae are as truly umbelliferous as the Um- 
belliferae; the Capparidaceae, like the Cruciferae, have cruciform 
flowers; in the Dipsacaceae the flowers are “composite,” as in the 
Compositae. Not being founded upon genera, such names may 
be applied to very different groups, even though they have the 
correct termination -aceae. The name Lomentaceae has been 
used for a division of Leguminosae, and also for a group of Cruci- 
ferae, and these two applications of the name, while equally appro- 
priate, are at the same time alike improper. Custom, then, is the 
‘only excuse for the continued use of this class of names. But it 
has proven true in the case of generic and specific nomenclature, 
that custom must yield to inflexible law; and surely the same 
should hold in the case of family nomenclature. 

What, then, should this law be? The evident preponderance 
of opinion demands as its most important feature the uniform 
termination -aceae. Then it must be recognized that the family 
names must be properly formed from the roots of generic names. 
‘ Next in importance is the law of priority, so necessary for stable 
botanical nomenclature. As the history of family names is fol- 
lowed out practical questions arise from time to time, and for 
their decision new provisions suggest themselves, and in this way 
gradually have been formulated the following rules for family 
nomenclature, which it is the object of this sketch to propose, 


Rules for Family Nomenclature. 


Rule r, The name of each natural family shall consist of the 
root of the accepted name of a recognized genus belonging to 
that family, with the addition of the termination aceae. 

Rule 2. The name of each natural family shall be the oldest 
name published in accordance with Rule 1, for any group of 
plants, based upon the accepted name of any recognized genus 
belonging to that family. 

Rule 3. The family name aust be published in Latin, and in 
the plural number, though not necessarily in the nominative case. 

Rule 4. Authorities shall be cited for family names in the same 
manner as for generic names. If the original author of a family 


ee ‘name spells the root incorrectly, his name shall be cited in paren- 


3 


‘theses, foliowed by the proper citation of the authority who first 
spells the name correctly. 

Rule 5. In conformity with the accepted rules of generic and 
specific nomenclature, no family name shall be accepted on the 
autnority of any work published prior to the first edition of 
Linné’s Species Plantarum, in 1753. 

The words “order” and “ family” have usually been used in- 
terchangeably in botany, but judging from the present tendency it 
appears likely that in the future the name “order” will be re- 
stricted, as has long been the case in zodlogy, to groups of higher 
rank than families. For this reason the word family has been 
used in the proposed rules. 

It will be well, perhaps, to elucidate the various points touched 
upon by the rules, by giving a few illustrations. 

Rule 1. The family name should consist of a generic root with 
the termination -aceae. Thus the family founded on the genus , 

- Triuris (root: TRIURID) should be called 7riuridaceae, Lindl., not 
7nuraceae, Gardn., nor yet 7rzuriaceae, Miers. 

It should be based on the name of a recognized genus. 
‘ Palmaceae’”’ is not founded on a genus, and cannot stand. 
‘«« Aquifoliaceae”’ is founded upon the genus Agwifolium, which is 
not now usually recognized, but is considered a section of //ex. 

‘The author who thus disposes of Aguzfolium cannot consistently 
use the name Aquifoliaceae. 

It should be founded on the accepted name of a genus. The 
genus on which Lindley established his family Roxburghiaceae is 
still recognized, but its accepted name at present is the older one 
Stemona, and hence for the family must be accepted the newer 
name Stemonaceae. 

Rule 2. Over twenty-five names have been proposed ending in 
-aceae and founded upon genera now included in the family Lili- 
aceae. Of these the oldest which fulfills the conditions of Rule 1 
is the one just mentioned, which was proposed by Adanson in 
1763. 

Rule 3. In 1819 De Candolle published the new families which 

_ he called, in French, “ Fumariacées ”’ and Frankeniacées ” (Théor. 
Elém. 244), but they were not published in Latin for two years, 
when S. F. Gray published Frankeniaceae, and De Candolle him- 


+ 


self Fumariaceae. Again, Schlechtendal, in Linnaea, in 1826, re- ~~ 
ferred to the “ Melanthiaceen,” but the Latin name first appeared 


in 1830 in Lindley’s Natural System. 

Bartling, in 1830 (Ord. Nat. 144), proposed a group “ Cen- 
taureacea.” This form was probably intended for the neuter 
plural, but might be the singular of Centaureaceae. In any event 
it does not fulfil the requirements of this rule. However, in 1873, 
Pfeiffer, in his Nomenclator Botanicus, referring to Bartling, spells 
the name Centaureaceae, thus duly publishing it. 

In 1825 Nees von Esenbeck, making a list of the plants belong- 
ing to the family which he called « Gesneriées,” entitled the list 
« Generum familiae Gessneriacearum brevis expositio.” Although 
the name Gessneriaceae occurs nowhere else in the article, and in 
this instance only in the genitive case, this constitutes a true pub- 
lication of the name according to the rule under consideration. 

Rule 4. Citation. There are many illustrations of this rule in 
the accompanying list. 

Rule 5. While this rule brings family nomenclature into line 
with specific and generic naming, it is desirable for another 
reason. In 1759 Bernard de Jussieu laid the foundation of the 
Natural System, which is now so universally accepted. As 2 
nothing on the subject was published between 1753 and 1759,the 
latter date might have been taken as a starting-point but for the 
convenience of uniformity. 

The following list has been prepared in accordance with, and 
in illustration of, the proposed rules. The omissions may be 
many ; the errors, it is to be hoped, are few. Additions to this 
list and corrections of it will be gratefully received. 

It will be well to remember, however, that many of the refer- 
ences in published books are not to be trusted, for they refer to 


places where the group is described, it may be under a very differ- 
ent name. 


List of the Natural Families According to the Classification 
Adopted in Engler and Prantl’s Natuerlichen Pflan- 
zenfamilien. Corrected in accordance 
with the proposed rules, 


This list includes only the Spermatophyta or seed-producing 
plants; it includes only names which end in -aceae; it includes 
only names which are formed, directly or indirectly, from the | 


5) 


name of some genus. This last restriction throws out such 
names as: 


AMENTACEAE, GLUMACEAE, PALMACEAE, 
AMPELIDACEAE, GRAMINACEAE, PAPILIONACEAE, 
ANTHERACEAE, GRANATACEAE, PATMACEAE, 
ASPERFOLIACEAE, GROSSULACEAE, PISTILLACEAE, 
AURANTIACEAE, HIPPOCASTANACEAE, POMACEAE, 
BALSAMACEAE, HOLORACEAE, POTAMIACEAE, 
CAMPANACEAE, HypDROCARYACEAE, PYRENACEAE, 
CERACEAE, LABIACEAE, ROTACEAE, 
CEREACEAE, LABIATACEAE, SAPONACEAE, 
CHLAENACEAE, LENTICULACEAE, SARMENTACEAE, 
COMPOSITACEAE, LENTISCACEAE, ScITAMINACEAE, 
CONACEAE, LOMENTACEAE, SPATHACEAE, 
DRUPACEAE, LUPULACEAE, STELLACEAE, 
ERUCACEAE, NUCAMENTACEAE, STROBILACEAE, 
FICOIDACEAE, NUCULACEAE, SYNANTHERACEAE, 
FRUMENTACEAE, OLERACEAE, UMBELLACEAE, ETC. 


Names formed by prefixing Eu- to the generic root have been 
omitted also, as 


EUBUXACEAE, EULACTUCACEAE, EULOBELIACEAE, EUVERNONIACEAE. 


The signs used are as follows: 5 

* Signifies that the family name is founded on more or less 
than the root of the generic name. In some of these cases it is 
formed by the addition of “-eae” to generic roots ending in “ ac”’; 
e. g. Smilac-eae. Such names are perfectly correct for tribes, 
though improper for families. | 

+ Indicates that the generic root is wrongly spelled. 

t Marks family names formed from generic names which are 
not recognized by Engler and Prantl. 

§ After a citation means that it is accepted on some authority 
believed to be trustworthy, but the reference has not been verified 
by actual comparison with the original work. In any such case 
it is possible that the publication may not have been in accordance 
with the proposed rules. ; 

The families which, in this list, are numbered respectively 129- 
137, 153-159, 175-182, 208-210, 227-229, 234, 235, and 243-248 


6 


i 


(all inclusive) have not yet been treated by Engler and Prantl, and 
of course their exact limitations and the generic names which will 
be recognized are largely matters of conjecture. 


1. CYCADACEAE Lindl. Nat. Syst. Ed. 2, 312 (1836). 
CyYCADEACEAE* Reichb. Consp. 40 (1828). 

ZAMIACEAE Reichb. Handb. 139 (1837). 

2. CORDAITACEAE Engler; Engl. & Pr. Nat. Pfl. 2: part 1: 26 (1889). 

3. PINACEAE Lindl. Nat. Syst. Ed. 2, 313 (1830). 
DamMarACcEAg£t Link, Abh. Berl. Ak. f. 1827, 157 (1830). 
SALISBURIACEAE} Link, Handb. 2: 523 (1831). 
TAXACEAE Lindl. Nat. Syst. Ed. 2, 316 (1836). 
PINEACEAE* Horan. Tetract. Nat, 22 (1843). 
JUNIPERACEAE Horan, Tetract. Nat. 22 (1843). 
CUPRESSACEAE Walp. Ann. Bot. 3: 444 (1853). 
ABIETACEAE Walp. Ann. Bot. 3: 446 (1853). 
PopOCARPACEAE Walp. Ann. Bot. 3: 448 (1853). 
ABIETINACEAE*® KI. & Grcke. Bot. Erg. Wald. 31 (1862), 
TAXODIACEAE Schimp. Paleont. Veg. 2: 309 (1870). 
ARAUCARIACEAE Strasb. Conif, 25 (1872). 

Called ConIFERAE by Engler & Prantl. 

4. EPHEDRACEAE Dumort. Fl. Belg. 9 (1827).8 
FHOACEAE} Agardh, Aphor. 204 (1825). 

GNETACEAE Lindl. Bot. Reg. ¢. 2686 (1834).§ 

5. .TYPHACEAE J. St. Hil. Expos. Fam. 1: 60 (1805). 

6. PANDANACEAE Lindl. Nat. Syst. Ed. 2, 361 (1836). 

7. SPARGANIACEAE Agardh, Theor. Syst. Pl. 13 (1858). 

8 ZANNICHELLIACEAE Dumort. Anal. Fam. 61 (1829). 
ZANICHELLIACEAE} Dumort. Anal. Fam. 59 (1829). 
ZOSTERACEAE Dumort. Anal. Fam. 65, 66 (1829). 
RupPiACcEAE Horan. Tetract. Nat. 22 (1843). 
POTAMOGETO ACEAE Engl. Fuehr. Bresl. Bot. Gart. 18 (1886).§ 
POSIDONIACEAE Kerner, Pflanzenleb. 2: 644 (1891). 
CYMODOCEACEAE Kerner, Pflanzenleb. 2: 644 (1891). 

9. NAJADACEAE (Lindl.) Asch. Linnaea, 35: 160 (1867). 
NAIADACEAE Lindl. Nat, Syst. Ed. 2, 366 (1836). 

1o. APONOGETONACEAE Engler, Bot. Jahrb. 8: 261 (1887). 
APONOGETACEAE* Agardh, Theor. Syst. Pl. 44 (1858). 

11, SCHEUCHZERIACEAE Agardh, Theor. Syst. Pl. 44 (1858). 
JuNCAGINACEAEt Lindl. Nat. Syst. Ed. 2, 367 (1836). 
LILAEACEAE Hieron, Ber, Ges. Nat. Berl. 116 (1878).$ 

12, ALISMACEAE DC. FI. Franc. 3: 181 (1805). 

13. BUTOMACEAE §. F. Gray, Arr. Brit. Pl. 2: 217 (1821). 

14. TRIURIDACEAE Lindl. Veg. Kingd. 213 (1847), 
TRIURACEAE* Gardn. Trans. Linn. Soc. rg: 160 (1843). 
TRIURIACEAE* Miers, Trans. Linn. Soc, 21: 43 (1850). 


mm, 


a8 


T 


15. ELODEACEAE Dumort. Anal. Fam. 54 (1829). 
VALLISNERIACEAE Dumort. Anal. Fam. 54, 55 (1829). 
HyDROCHARACEAE® Lindl. Nat. Syst. Ed. 2, 335 (1836). 
HyDROCHARIDACEAF} Lindl, Veg. Kingd. 141 (1847). 
HyYDROCHARITACEAE (Lindl.) Asch, Linnaea, 35: 158 (1867). 
HALOPHILACEAE Kerner, Pflanzenleb. 2: 644 (1891). 
STRATIOTACEAE Kerner, Pflanzenleb, 2: 645 (1891). 
OTTELIACEAE Kerner, Pflanzenleb. 2: 645 (1891). 
THALASSOIDACEAE} Kerner, Pflanzenleb. 2: 645 (1891). 
BLYXEACEAE* Kerner, Pflanzenleb. 2: 645 (1891). 
HyYpDRILLACEAE Kerner, Pflanzenleb. 2: 645 (1891). 

16. POACEAE R. Brown, Flind. Voy. App. 2, 583 (1814). 
STIPACEAE HBK., Nov. Gen. 1: 121 (1815). 
AVENACEAE HBK. Noy. Gen. 1: 143 (1815). 
FesTucAceaE HBK. Nov. Gen. 1: 143 (1815). 
ARUNDINACEAE HBK. Nov. Gen. 1: 148 (1815). 
Horpeacear HBK. Nov. Gen. 1: 179 (1815). 
OLyRACEAE HBK. Noy. Gen, 1: 196 (1815). 
BAMBUSACEAE IIBK. Nov. Gen. 1: 199 (1815). 
BROMACEAE Dumort. Agrost. Belg. 82 (1823). 
PHLEACEAE Dumort. Agrost. Belg. 83 (1823). 
LotaAceaAE Dumort. Agrost. Belg. 95 (1823). 
MILIACEAE Dumort. Agrost, Belg. 135 (1823). 
SPARTINACEAE Link, Hort. Berol. 1: 46 (1827).§ 
EcHINARIACEAE Link, Hort. Berol. 1: 197 (1827).$ 
CHONDROSIACEAEt Link, Hort. Berol. 1: 269 (1827).§ 
PASPALACEAE Link, Hort. Berol. 1: 269 (1827).§ 
MELICACEAE Link, Hort. Berol. 1: 271 (1827).§ 
ZEACEAE Reichb, Consp. 55 (1828). 

TRIPSACEAE* Dumort. Anal. Fam. 64 (1829). 
ROTTBOELLIACEAE Kunth, Gram. 150 (1830).§ 
ANTHISTIRIACEAE Presl, Reliq. Haenk. 1: (1830).§ 
VILFACEAE} Trinius, hinnaea, ro: 302 (1835). 
SESLERIACEAE W. Koch, Synops 788 (1837). 
PAPPOPHOREACEAE® Parlat. Fl. Palerm. 1: 127 (1845).$ 
SESSLERIACEAE}+ Fries, Summ. Veg. Scand. 1: 80 (1846). 
AGROSTACEAE* Pfeiff. Nomencl. Bot. r: 85 (1873). 
Called GRAMINEAE by Engler and Prantl. 

17. CYPERACEAE J. St. Hil. Expos, Fam, 1: 62 (1805). 
ELYNACEAE Reichb. Consp. 55 (1828). 

SCLERIACEAE Reichb. Consp. 56 (1828). 
ScrrPAcEAE Kerner, Pflanzenleb. 2 : 654 (1891). 

18. ARECACEAE Reichb. Consp. 72 (1828). 

CorvPHACEAE Reichb. Consp. 73 (1828). 
NIPACEAE Brongn. Enum. Genr. 15 (1843). 

ae SABALACEAE Schimp. Paleont. Veg. 2: 486 (1871). 
PHOENICACEAE Schimp, Paleont. Veg. 2: 496 (1871). 


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8. 


BoRASSACEAE Schimp. Paleont. Veg. 2: 499 (1871). 
LEPIDOCARYACEAE Kerner, Pflanzenleb. 2: 649 (1891). 
CEROXYLONACEAE* Kerner, Pflanzenleb. 2: 649 (1891). 
PHYTELEPHANTACEAE Kerner, Pflanzenleb. 2: 649 (1891). 
Called PALMAE by Engler and Prantl. 
19. CYCLANTHACEAE Lindl. Nat. Syst. Ed. 2, 362 (1836). 
LupoviAcEAE Drude; Engl. & Pr. Nat. Pfl. 2: part 3, 93 (1889). 

20. ARACEAE Neck. Act. Ac. Theod. Palat. 2: 462 (1770). 
ORONTIACEAE R. Brown, Prodr. 1: 337 (1810). 
Pist1aAcEAE HBK, Nov. Gen. 1: 81 (1815). 

CALLACEAE Reichb. Consp. 44 (1828). 

ACORACEAE Lindl. Nat. Syst. Ed. 2, 365 (1836). 
CoLocaSsIACEAE Kerner, Pflanzenleb, 2: 646 (1891). 
PHILODENDRACEAE Kerner, Pflanzenleb, 2: 646 (1891). 
LASIACEAE Kerner, Pflanzenleb. 2: 646 (1891). 
MONSTERACEAE Kerner, Pflanzenleb. 2: 646 (1891). 
PoTHOIDACEAE* Kerner, Pftanzenleb, 2: 646 (1891). 

21. LEMNACEAE Dumort. Fl. Belg. 147 (1827).§ 

22, FLAGELLARIACEAE Agardh, Theor. Syst. Pl. 20 (1858). 

23. RESTIONACEAE Hieron.; Engl. & Pr. Nat. Pf. 2: part 4, 3 (1888). 
REsTIACEAE* R, Brown, Prodr. 1: 243 (1810), 

24. CENTROLEPIDACEAE Hieron. Abh. Nat. Ges. Halle, 205 (1873). 
DEVAUXIACEAE}} Dumort. Anal. Fam, 62, 63 (1829). 
DesvAuXIACEAEt Lindl. Nat. Syst. Ed. 2, 386 (1836). 

25. MAYACACEAE Walp. Ann. Bot. 3 : 662 (1853). 
MAYACEAE* Meisn. Pl. Vasc. Gen, 406, 407 (1842). 

26. XYRIDACEAE Lindl. Nat. Syst. Ed. 2, 388 (1836). 

27. ERIOCAULACEAE Lindl. Veg. Kingd. 122 (1847). 
ERIOCAULONACEAE* OK, Rev. Gen. 745 (1891). 

28. RAPATEACEAE Dumort. Anal. Fam. 60, 62 (1829). 

29. BROMELIACEAE J. St. Hil. Expos. Fam. 1: 122 (1805). 

30. COMMELINACEAE Reichb. Consp. 57 (1828). 
COMMELYNACEAE} Endl. Gen. 124 (1837). 

31. PONTEDERIACEAE Dumort. Anal. Fam. 59, 61 (1829). 
PONTEDERACEAE* Martius, Consp. 7 (1835). 
PONTEDERAEACEAE} OK. Rev. Gen. 718 (1891). 

32. PHILYDRACEAE Lindl. Nat. Syst. Ed. 2, 357 (1836). 
PHYLIDRACEAE}+ Horan. Tetract. Nat. 22 (1843). 

33- JUNCACEAE (Vent.) Dumort. Comm. Bot. 66 (1822), 
JONCACEAEt Vent. Tabl. 2: 150 (1799). 

34. STEMONACEAE Fr. & Sav. Enum. Pl. Jap. 2: 92 (1879). 
ROXBURGHIACEAE$ Wallich, Pl. As. Rar. 3: 50 (1832). 

35- LILIACEAE Adans. Fam. Pl. 2: 42 (1763). 

ALLIACEAE Batsch, Gen. PI. Jenens. 10, 30 (1786). 
TULIPACEAE Batsch, Gen, Pl. Jenens. 11, 30 (1786). 
SMILACEAE* Vent. Tabl. 2: 146 (1799). 
CoLcHICACEAE DC, FI. Frang. 3: 192 (1805). 


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9 


MELANTHACEAE® R, Brown, Prodr. 1: 272 (1810). 
ASPHODELIACEAE® S, F. Gray, Arr. Brit. Pl. 2: 174 (1821). 
PHYLESIACEAE+ Dumort. Anal. Fam. 53, 54 (1829). 
ASTELIACEAE Dumort. Anal. Fam. 59, 61 (1829). 
XANTHORHAEACEAE} Dumort. Anal. Fam. 60, 62 (1829). 
PHILESIACEAE Dumort. Anal. Fam. 97 (1879). 
CONVALLARIACEAE Link, Handb, 1: 184 (1829). 
DRACAENACEAE Link, Handb. 1: 187 (1829). 
MELANTHIACEAE Lindl. Nat, Syst, Ed. 1 (1830).§ 
TRILLIACEAE Lindl, Nat. Syst. Ed. 2, 347 (1836). 
GILLIESIACEAE Lindl, Nat. Syst. Ed. 2, 348 OP); 
KINGIACEAE Endl. Gen. 132 (1837). 

LAXMANNIACEAE Horan. Tetract. Nat, 23 (1843). 
FUNKIACEAE} Horan. Tetract. Nat. 23 (1843). 
ASPHODELACEAE Horan. Tetract. Nat. 23 (1843). 
HAWORTHIACEAE Horan. Tetract. Nat. 23 (1843). 
ASPARAGACEAE Horan. Tetract. Nat. 23 (1843). 
NARTHECIACEAE Fries, Summ. Veg. Scand. 1: 65 (1846). 
LAPAGERIACEAE Kunth; Walp. Ann. Bot. 3: 646 (1853). 
HERRERIACEAE Kunth; Walp. Ann. Bot. 3: 646 (1853). 
OPHIOPOGONACEAE Kunth; Walp. Ann. Bot. 3; 646 (1853). 
ASPIDISTRACEAE Kunth; Walp. Ann. Bot. 3: 646 (1853). 
UvuLARIACEAE Walp. Ann. Bot. Bot. 3: 650 (1853). 
PHORMIACEAE Agardh, Theor. Syst. Pl. 7 (1858). 

36. HAEMODORACEAE R. Brown, Prodr. 1: 299 (1810). 
HAEMADORACEAE} Reichb, Consp. 60 (1828). 
HeMoporaceEAE+ Dumort. Anal. Fam. 58 (1829). 
WACHENDORFIACEAE Dumort. Anal. Fam. 61 (1829). : 
HoeMoporaceEAFf A. Rich. Sert. Astrol. 80 (1834). 

37. LEUCOJACEAE Batsch, Gen. Pl. Jenens. 10, 30 (1786). 
ALSTROEMERIACEAE Dumort. Anal. Fam. 58 (1829). 
CAMPYNEMACEAE Dumort. Anal. Fam. 58 (1829). 
AGAVEACEAE Dumort. Anal. Fam. 58 (1829). 
AMARYLLIDACEAE Lindl. Nat. Syst. Ed. 2, 328 (1836). 
PANCRATIACEAE Horan. Tetract. Nat. 23 (1843). 
HyypoxipAceAe Lindl. Veg. Kingd. 154 (1847). 

38. VELLOZIACEAE Drude, Phanerog. 333 (1879). 

39. TACCACEAE Reichb. Consp, 44 (1828). 

40. TAMACEAE Gray, Arr. Brit. Pl. 2: 189 (1821). 
DroscorEaceak Lindl. Nat. Syst. Ed. 2, 359 (1836). 
Droscorrpaceae} Kl. & Greke. Bot. Erg. Wald. 42, 55 (1862). 
STENOMERIDACEAE Kerner, Pflanzenleb. 2: 666 (1891). 

41. IXIACEAE Ecklon, Verzeichn 18 (1827).§ 

MorAEACEAE Dumort. Anal. Fam. 58 (1829). 
IRIDACEAE Lindl. Nat. Syst. Ed. 2, 332 (1836). 
42. MUSACEAE J. St. Hil. Expos. Fam, 1: 151 (1805). 


10 


43. ALPINIACEAE Link. Enum. 1: 228 (1821). 
CurcuMACEAE Dumort. Anal. Fam. 20, 25 (1829). 
ZINGIBERACEAE Lindl. Nat. Syst. Ed. 2, 322 (1836). 
AMOMACEAE Horan. Tetract, Nat. 22 (1843). 

44. CANNACEAE Link, Enum. r: 1 (1821). 

45. MARANTACEAE Lindl. Nat. Syst. Ed. 1 (1830), 

46. BURMANNIACEAE Blume, Enum. Pl. Jav. 1: 27 (1830). 
THISMIACEAE Miquel, Fl. Ind. Bat. 3: 615 (1858) § 
ARACHNITACEAE Philippi, Cat. Pl. Vasc. Chil. 278 ( 1881). 

47- ORCHIDACEAE Lindl. Nat. Syst. Ed. 2, 336 (1836) 
VANILLACEAE Lindl. Nat. Syst. Ed. 2, 341 (1836). 
APOSTASIACEAE Lindl, Nat. Syst. Ed. 2, 342 (1836). 
LimoporaceaE Horan. Tetract. Nat. 22 (1843). 
NEOTTIACEAE Reichb. f. Poll. Orch. Gen. 9 (1852).§ 
ConntacEart Reichb, f. Bot. Zeit. 929 (1852). 
RODRIGUEZIACEAE Reichb. f. Bot. Zeit. 929 (1852). 
CHLORAEACEAE Reichb, f. Bot, Zeit. 1 (1853). 
CypRIPEDIACEAE KI. & Greke. Bot. Erg. Wald. 33, 38 (1862). 
OPHRYDACEAE Kerner, Pflanzenleb, 2: 661 (1891). 
EPIDENDRACEAE Kerner, Pflanzenleb. 2: 661 (1891), 
VANDACEAE Kerner, Pflanzenleb, 2: 661 (1891). 

48 SAURURACEAE Lindl. Nat. Syst. Ed. 2, 184 (1836). 

49. PIPERACEAE HBK. Nov. Gen. 1: 46 (1815). 

50. CHLORANTHACEAE Blume, Enum. Pl. Jav. r: 78 (1830). 

5'. LACISTEMACEAE Lindl. Nat. Syst. Ed. 2, 183 (1836). 

52, CASUARINACEAE Lindl. Veg. Kingd. 249 (1847). 
Casuaracear* Lindl. Nat, Syst. Ed. 2, 181 (1836). 

53- JUGLANDACEAE Lindl. Nat. Syst. Ed. 2, 180 (1836). 

54. MYRICACEAE Dumort, Anal. Fam. 95 (1829). 

55. LEITNERIACEAE Drude, Phanerog. 407 (1879). 

56. SALICACEAE Lindl. Nat. Syst. Ed. 2, 186 (1836). 

57- CORYLACEAE Mirbel, Elem. 2: 296 (1815). 

BeTULACEear Agardh, Aphor. 208 (1825), 

58. FAGACEAE Drude, Phanerog. 409 (1879). 
CasTANEACEAE Baill. Dict, Bot, 1: 650 (1884 ?), 

59. ULMACEAE Mirbel, Elem. 2:905 (1815). 
CELTIDACEAE Walp. Ann. Bot. 3: 394 (1853). 

60. ARTOCARPACEAE Horan. Tetract. Nat, 25 (1843). 
CANNABINACEAE Lindl, Veg. Kingd. 265 (1847). 
Moraceae Lindl, Veg. Kingd. 266 (1847). 
CANNABACEAE* A. Braun; Asch. Fl. Brand. 58 (1864).g 
Dorsrentaceag Kerner, Pflanzenleb. 2: 680 (1891). 
ConocePHALacear Kerner, Pflanzenleb. 2: 680 (1891). 

61. URTICACEAE Reichb, Consp. 83 (1828). 

PHENACEAE* Weddell, Ann, Sc. Nat. Ser. 4, 1: 175 (1854). 

62. PROTEACEAE J. St. Hil. Expos. Fam. 1: 185 (1805). 

_ Prrsoniaceac} Klotsch, Linnaea 20: 471 (1847). 


68. 


70. 


71. 


72. 


73- 


74- 


75- 
76. 


77: 
78. 


it 


. LORANTHACEAE D. Don. Prodr. Fl. Nepal. 142 (1825). 


VISCACEAE Miers, Ann. & Mag. N. H. (II.) 8: 179 (1851). 
MYZODENDRACEAE Hieron.; Engl. & Pr. Nat. Pfl. 3: part 1, 198 (1889). 


- SANTALACEAE R. Brown, Prodr, 1: 350 (1810). 


CanopiAceatt Presl. Epimel. Bot. 608 (1850). 


. GRUBBIACEAE Endl. Gen. 327 (1838). 
- OLACACEAE Lindl. Nat. Syst. Ed. 2, 32 (1836). 


OvaceaeE* Benth. Trans. Linn. 18: 677 (1841). 

SCHOEPFIACEAE Blume, Mus. Bot. Lugd. 1 : 175 (1850). 

APTANDRACEAE Miers, Ann. & Mag. N. H. Ser. 2, 7: 206 (1851). 

OLACINACEAE* KI], & Grcke. Bot. Erg. Wald. 151 (1862). 
CYNOMORIACEAE Lindl. Nat. Syst. Ed. 2, 394 (1836). 

BALANOPHORACEAE Lindl. Nat. Syst. Ed. 2, 525 (1836). 

LATHRAEOPHILACEAE Leand. de Sacram.; A. St. Hil. Ann. Sc. Nat. Ser 2. 

7: 32 (1837). 

LOPHOPHYTACEAE Horan. Tetract. Nat. 21 (1843). 

SARCOPHYTACEAE Kerner, Pflanzenleb. 2; 708 (1891). 

SCYBALIACEAE Kerner, Pflanzenleb. 2: 708 (1891). 


. ASARACEAE Link, Enum. 2: 1 (1822). 


ARISTOLOCHIACEAE Blume, Enum. Pl. Jav. 1: 81 (1830). 

APAMACEAE Kerner, Pflanzenleb. 2: 700 (1891). 
RAFFLESIACEAE Dumort. Anal. Fam. 13, 14 (1829). 

CyYTINACEAE Lindl. Nat. Syst. Ed. 2, 392 (1836). 

APODANTHACEAE Kerner, Pflanzenleb. 2: 700 (1891). 

HY DNORACEAE Graf zu Solms, Bot. Zeit. 66 (1874).§ 
POLYGONACEAE Lindl. Nat. Syst. Ed. 2, 211 (1836). 

ERIOGONACEAE Walp, Ann. Bot. 3: 297 (1853). 
CHENOPODIACEAE Dumort. Anal. Fam: 15, 17 (1829). 

CHENOPODEACEAE} Martius, Consp. 15 (1835). 

SALSOLACEAE Mogq.-Tand.; DC. Prodr. 13: part 2, 41 (1849). 

ATRIPLICACEAE Simonkai, Enum. Fl. Trans. 465 (1886). 
AMARANTACEAE (J. St. Hil) Martius, Nov. Act. Ac. Leop. 13: part I, 215 

(1826). 

AMARANTHACEAE} J. St. Hil. Expos. Fam. 1: 204 (1805). 
BATIDACEAE Dammer; Engl. & Pr. Nat. Pfl. 3: part Ia, 118 (1893). 
CYNOCRAMBACEAE Pouls.; Engl. & Pr, Nat. Pfl. 3: part 1a, 121 (1893). 

THELYGONACEAE} Caruel, Nuov. Giorn. Bot. It. 5: 170 (1873). 
BASELLACEAE Mog.-Tand. Chenop. x (1840). 

PETIVERIACEAE Link, Handb. 1: 392 (1829). 

RIvINIACEAE} Dumort. Anal. Fam. 17 (1829). 

PETIVERACEAE® Lindl. Nix. Pl. 16 (1833). 

PHYTOLACCACEAE Lindl. Nat. Syst. Ed. 2, 210 (1836). 


79. ALLIONIACEAE Reichb. Consp. 85 (1828). 


NycraGInaceAEt Lindl. Nat. Syst. Ed. 2, 213 (1836). 


So. TETRAGONIACEAE Reichb.; Moessl. Handb. 1: 52 (1827).§ 


MEseMBRYACEAE® Lindl. Nat. Syst. Ed. 2, 56 (1836). 
SESUVIACEAE Horan. Tetract Nat. 29 (1843). 


* 


12 


AIZOACEAE A. Braun; Asch. Fl. Brand. 60 (1864).§ ? 
MESEMBRIANTHEMACEAE Lowe, FI. Madeir. 306 (1868). 
MOLLUGINACEAE Rohrb.; Martius, Fl. Bras. 14; part 2, 228 (1872). 

81. PORTULACACEAE Reichb. Consp, 161 (1828). 
PoRTULACEAE* Juss. Gen. 312 (1789). 

82. ALSINACEAE Wahlenb. Fl. Suec. 2: Ixxiv (1824). 
CORRIGIOLACEAE Reichb.; Moess]. Handb. 1: 51 (1827).§ 
STELLARIACEAE Dunnort. FI. Belg. 106 (1827) § 
QUERIACEAE DC. Prodr. 3: 379 (1828). 
CARYOPHYLLACEAE} Reichb. Consp, 206 (1828). 
TELEPHIACEAE Link, Handb. 2: 45 (1831). 

. PARONYCHIACEAE Link, Handb. 2: 420 (1831). 
SILENACEAE Lindl, Nat, Syst. Ed. 2, 124 (1836). 
ILLECEBRACEAE Lindl, Nat. Syst. Ed. 2, 127 (1836). 
SCLERANTHACEAE Lind], Nat. Syst. Ed. 2, 213 (1836). 
MALACHIACEAEt C, Koch, Linnaea, 15: 709 (1841). | 
LOEFFLINGIACEAE} Fzl.; Walp. Repert. 1: 263 (1843). 

83. NYMPHAEACEAE DC. Propr. Méd. Ed. 2, 119 (1816). 
NetumBtaceAe} Lindl. Nat. Syst. Ed. 2, 13 (1836). 
CABOMBACEAE A. Gray, Ann. Lyc. N. Y. 4: 46 (1837). 
EuRYALACEAE Kerner, Pflanzenleb. 2: 699 (1891). 
NuPHARACEAE Kerner, Pflanzenleb. 2: 699 (1891). 
BARCLAYACEAE Kerner, Pflanzenleb, 2; 699 (1891). 

84. CERATOPHYLLACEAE A. Gray, Ann. Lyc. N. Y. 4: 41 (1837). 

85. MAGNOLIACEAE J. St. Hil. Expos. Fam. 2: 74 (1805). 
SCHIZANDRIACEAE*® G, Don, Gen. Syst. 1: 191 (1831). 
SCHIZANDRACEAE Martius, Consp. 39 (1835). 
WINTERACEAE} Lindl. Nat. Syst. Ed. 2, 17 (1836). 

86. LACTORIDACEAE Engler, Bot. Jahrb. 8: 53 (1887). 

87. TROCHODENDRACEAE Prantl; Engl. & Pr. Nat. Pfl. 3, se 2: 21 (1891)- 

88. ANONACEAE DC. Syst. 1: 463 (1818). 

ANNONACEAE} Link, Enum. 2? 87 (1822). 

89. MYRISTICACEAE Lindl. Nat. Syst. Ed. 2, 15 (1836). 

90. RANUNCULACEAE Juss. Gen. 231 (1789). 
PAEONIACEAE DC. Prodr. 1: 64 (1824). 

CALTHACEAE Presl, Fl. Sicul. 1: 20 (1826).§ 
POEONIACEAF} Presl, Fl. Sicul. 1: 26 (1826). 
HELLEBORACEAE Spach, Hist. Veg. Phan. 7: 285 (1839).8 
NIGELLACEAE Agardh, Theor. Syst. Pl. 76 (1858). 

91. LARDIZABALACEAE Lindl. Veg. Kingd. 303 (1847). 

92. PODOPHYLLACEAE DC, Prodr..1: 111 (1824). 
DiPHYLLEIACEAE Schultz, Nat. Syst. Pfl. 328 (1832). 
BERBERACEAE* Lindl. Nat. Syst. Ed. 2, 7 (1836) 
BERBERIDACEAE Torr. & Gr. Fl. N. Am. r: 49 (1838). 
NANDINACEAE Horan. Tetract. Nat. 30 (1843). 


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13 


93. MENISPERMACEAE DC, Prodr. 1: 95 (1824). 

94. CALYCANTHACEAE Lindl. Nat. Syst. Ed. 2, 159 (1836). 

95. MONIMIACEAE Dumort. Anal. Fam. 16 (1829). 
ATHEROSPERMACEAE Lindl, Nat. Syst. Ed. 2, 189 (1836). 

g€ CASSYTHACEAE Dumort. Anal. Fam. 16 (1829). 
LAURACEAE Lindl. Nat. Syst. Ed. 2, 200 (1836). 
CassYTACEAE} Horan. Tetract. Nat. 24 (1843). 
PERSEACEAE Horan. Tetract Nat. 25 (1843). 
LiTsEACEAE Benth. & Hook, Gen. Pl. 3: 149, 160 (1880). 

97, HERNANDIACEAE Dumort. Anal. Fam. 14, 16 (1829). 
ILLIGERACEAE Lindl. Nat. Syst. Ed. 2, 202 (1836). 
GYROCARPACEAE KI]. & Grcke. Bot. Erg, Wald. 151 (1862). 

98. PAPAVERACEAE B. Juss. Hort. Trian. (1759). 
FUMARIACEAE DC, Syst, 2: 104 (1821). 

99. BRASSICACEAE Lindl. Nat. Syst. Ed. 2, 58 (1836). 
Called CRUCIFERAE by Engler and Prantl. 


‘too. TOVARIACEAE Pax; Engl. & Pr. Nat. Pf. 3: part 2, 207 (1891). 


1o1. CAPPARIDACEAE Lindl. Nat. Syst. Ed. 2, 61 (1836). 
CLEOMEACEAE Horan. Tetract. Nat. 31 (1843). 

102, RESEDACEAE S. F, Gray, Arr. Brit. Pl. 2: 665 (1821). 
ASTEROCARPACEAF} Kerner, Pflanzenleb. 2: 688 (1891). 

103: MORINGACEAE Dumort. Anal. Fam. 43, 48 (1829). 

104. SARRACENTACEAE La Pylaie, Mem. Soc. Linn. Par. 6: 379 (1827).§ 

103. NEPENTITACEAE Lindl. Nat. Syst. Ed. 2, 204 (1836). 

106. DROSERACEAE §, F. Gray, Arr. Brit. Pl. 2: 664 (1821), 
DIONAEACEAE Lindl. Nat. Syst. Ed. 2, 14 (1836). 

DrionaceAe* Dumort. Bull. Ac. Brux. 4: 447 (1838). 

107. PODOSTEMACEAE Lindl. Nat. Syst, Ed. 2, 190 (1836). 
PHILOCRENACEAEf Bongard, Mem. Ac. St. Pet., Ser. 6, 1: 72 (1835).§ 
TRISTICHACEAE Kerner, Pflanzenleb. 2: 673 (1891). 
WEDDELLINACEAE Kerner, Pflanzenleb. 2: 673 (1891). 
HyYDROSTACHYDACEAE Kerner, Pflanzenleb. 2: 673 (1891). 

108. SEDACEAE Neck. Act. Ac. Theod. Palat. 2: 487 (1770). 
CRASSULACEAE DC, FI. Franc. 4: 382 (1805). 

109, CEPHALOTACEAE Lindl. Key (1835) § 

110. ESCALLONIACEAE Dumort. Anal. Fam. 35, 37 (1829). 
RIBESIACEAE* Reichb. Consp. 160 (1828). 

SAXIFRAGACEAE Dumort. Anal. Fam. 36, 38 (1829). 
HyYPRANGEACEAE Dumort. Anal. Fam. 36, 38 (1829). 
GROSSULARIACEAE{ Dumort. Anal. Fam. 37, 42 (1829). 
PARNASSIACEAE Dumort. Anal. Fam. 44, 49 (1829). 
BAUERACEAE Lindl. Nat. Syst. Ed. 1 (1830). 
BREXIACEAE Lindl Nat. Syst. Ed. 1 (1830). 
FRANCOACEAE A. Juss. Ann. Sc. Nat. 25: 9 (1832). 
PHILADELPHACEAE Lindl, Nat. Syst, Ed. 2, 47 (1836). 


Itt. 
112, 
113. 
114. 


115. 
116. 


117. 


118, CONNARACEAE R. Brown, Exp. Cong. App. 5, 431 (1818). 


14 


RoussAEACEAE} DC. Prodr. 7, part 2: 521 (1839). 
PoLyosMACEAE Blume, Mus. Bot. Lugd. 1: 258 (1850). 
IxXERBIACEAE Griseb. Grundr, Syst. Bot. 122 (1854). 


RovussEACEAE (DC.) Griseb. Grundr. Syst. Bot. 123 (1854). 


ITEACEAE Agardh, Theor. Syst. Pl. 151 (1858). 


CUNONIACEAE R. Brown, Flind. Voy. App. 3, 548 (1814). 
MYROTHAMNACEAE Niedenzu; Engl. & Pr. Nat. Pfl. 3: part 2a, 103 (1891) 


PITTOSPORACEAE Lindl. Nat. Syst. Ed. 2, 31 (1836). 
ALTINGIACEAE Hayne, Flora, 1: 172 (1830). 
AMBRACEAE® Reichb. Consp. 113 (1828). 
HAMAMELACEAE® Lindl. Nat. Syst. Ed. 2, 48 (1836). 
PARROTIACEAE Horan. Tetract. Nat. 28 (1843). 
HAMAMELIDACEAE Lindl. Veg. Kingd. 784 (1847). 
AMAMELIDACEAE Pfeiff. Nomencl. Bot. 1: 129 (1873). 
BRUNIACEAE R. Brown, Abel Journ. App. 374 (1818). 
PLATANACEAE Lindl. Nat. Syst. Ed. 2, 187 (1836), 
ROSACEAE B. Juss. Hort. Trian. (1759). 
AGRIMONIACEAE DC. Fl, Franc. 4: 448 (1805). 
FRAGARIACEAE Nest. Potent. 14 (1816). 
SPIRAEACEAE Dumort. Comm. Bot. 59 (1822). 
POTENTILLACEAE IIBK. Nov. Gen. 6: 215 (1823). 
SPIREACEAE + D. Don. Prodr. Fl. Nepal. 227 (1825). 
HAGENIACEAE Reichb. Consp. 145 (1828). 
AMYGDALACEAE Reichb, Consp. 177 (1828), 
CLIFFORTIACEAE Dumort. Anal. Fam. 18 (1829). 
CHRYSOBALANACEAE Lindl. Nat. Syst. Ed. 2, 158 (1836). 
SANGUISORBACEAE Lindl. Veg. Kingd. 561 (1847). 
NEILLIACEAE Miguel, Fl. Ind. Bat. 1: 390 (1855).S 
DRYADACEAE Frank; Leunis, Synops. Pf. 2: 160 (1885). 
POTERIACEAE Frank ; Leunis, Synops. Pf. 2: 173 (1885). 


119. CASSIACEAE Link, Handb. 2: 135 (1831). 


FaBACEAEt Reichb. Consp. 149 (1828). 

SOPHORACEAE Link, Handb. 2: 143 (1831). 

MiMmosacEAe Reichb. Fl. Exc. 437 (1832). 

VictacEaE C. Koch, Linnaea 12: 727 (1841). 
CAESALPINIACEAE KI. & Greke. Bot. Erg. Wald. 157 (1862). 
PHASEOLACEAE Pfeiff. Nomencl. Bot. 2: 668 (1874). 

Called LEGUMINOSAE by Engler and Prantl. 


120. GERANIACEAE J, St. Hil. Expos, Fam. 2: 51 (1805). 


VIVIANIACEAE Klotsch, Linnaea ro: 433 (1836). 
VIVIANACEAE} Agardh, Theor. Syst. Pl. 203 (1858). 


LEDOCARPACEAEt Kl. & Grcke. Bot. Erg. Wald. 121 (1862). 


"121, OXALIDACEAE Lindl. Nat. Syst. Ed. 2, 140 (1836). 
122, TROPAEOLACEAE Lindl. Veg. Kingd. 366 (1847). 


Sr sea 


ie Gia eta ee Oe AOE 


Zines 


Settee 


123. 
124. 
125. 


126. 


127, 


128. 


129. 


130. 


131. 


132. 


133. 


134. 


135. 


136. 


137. 


15 


LINACEAE Dumort. Comm. Bot. 61 (1822). 

HUGONIACEAE Arn.; Wight & Arn. Prodr. 1: 71 (1834). 
HUMIRIACEAE A. Juss; A. St. Hil. Fl. Bras. Mer. 2: 87 (1829).8 
ERYTHROXYLACEAE A, Rich. Pl. Vasc. Cub. 254 (1842).§ 
MALPIGHIACEAE Vent. Tabl. 3: 131 (1799). 

HIRAEACEAE Griseb,; Martius, Fl. Bras. 12: 3, 75 (1858). 
NITRARIACEAE Lindl. Nat. Syst. Ed. 1 (1830). 

GUAIACEAE* Reichb. Consp. 200 (1828). 

ZYGOPHYLLACEAE Lindl. Nat. Syst. Ed. 2, 133 (1836). 
CNEORACEAE Engler; Engl. & Pr. Nat. Pfl. 3: part 4, 93 (1890). 
RUTACEAE Juss. Gen, 296 (1789). 

PTELEACEAE Kunth, Ann. Sc. Nat. 2: 354 (1824). 

XANTHOXYLACEAE Lindl. Nat. Syst. Ed. 2, 135 (1836). 

ZANTHOXYLACEAE} Meisn. Pl. Vasc. Gen. 64 (1837). 

CORREACEAE Agardh, Theor. Syst. Pl. 229 (1858). 

CITRACEAE Drude, Phanerog. 391 (1879). 

BORONIACEAE Kerner, Pflanzenleb, 2: 676 (1891). 
SIMARUBACEAE DC. Bull. Soc. Philom. 2: 209 (1811). 

SURIANACEAE Lindl. Nat. Syst. Ed. 2, 142 (1836). 

SIMABACEAE* Horan. Tetract. Nat. 31 (1843). 
BURSERACEAE Kunth, Ann. Sc. Nat. 2: 346 (1824). 

BALSAMEACEAE Dumort. Anal, Fam. 36, 41 (1829). 

BURSERIACEAE* G, Don, Gen. Syst. 2: 79 (1832). 

AMYRIDACEAE Lindl. Nat. Syst. Ed. 2, 165 (1836). 
MELIACEAE Vent. Tabl. 3: 159 (1799). 

CEDRELACEAE A. Juss. Mem. Mus, 1g: 213, 247 (1830). 
TRIGONIACEAE Martius, Consp. 51 (1835). 
VOCHYSIACEAE Mart. & Zucc. Noy. Gen. 1: 123 (1824). 

ERISMACEAE Dumort. Anal, Fam. 41 (1829). 

VocuyacEAE* Lindl. Nat. Syst. Ed. 1 (1830). 
TREMANDRACEAE Dumort. Anal. Fam. 43 (1829). 
POLYGALACEAE Reichb. Consp. 120 (1828). 

KRAMERIACEAE Dumort. Anal. Fam. 20, 23 (1829). 
CHAILLETIACEAE DC. Prodr. 2: 57 (1825). 


138. EUPHORBIACEAE J. St, Hil. Expos. Fam. 276 (1805). 


RictnaceaE Nor.; Dup.-Thouars, Veg. Il. Afr. 28 (1807).§ 
HuraceaAg Dumort. Anal. Fam. 45 (1829). 

ScEPACEAE Lindl. Nat. Syst. Ed. 2, 171 (1836). 
TREWIACEAE Lindl. Nat. Syst. Ed. 2, 174 (1836). 
STILAGINACEAE Lindl. Nat. Syst. Ed. 2, 179 (1836). 
ANTIDESMACEAE Horan. Tetract. Nat. 25 (1843). 
BENNETTIACEAE} Schnizl. Icon. ¢. 77? (1843).§ 
BERTYACEAE Agardh, Theor. Syst. Pl. 190 (1858). 
PeraceaE Klotsch, Tricoce. 12 (1860) § 

ACALYPHACEAE Klotsch, Tricocc. 12 (1860).§ 


16 


PHYLLANTHACEAE Klotsch, Tricoce. 12 (1860).§ 


DAPHNIPHYLLACEAE Muell.-Arg.; DC. Prodr. 16: part 1, 1 (1869). 


TITHYMALACEAEt Kerner, Pflanzenieb. 2: 674 (1891). 

139. CALLITRICHACEAE Lindl. Nat. Syst. Ed. 2, 191 (1836). 
STELLARIACEAEt Mac M. Metasp. Minn. Val. 344 (1892). 

140. EMPETRACEAE Dumort. Fl. Belg. 106 (1827).§ 

141. CORIARIACEAE Dumort. Anal. Fam. 87 (1829). 

142, BUXACEAE Dumort. Comm, Bot. 54 (1822). 

143) LIMNANTHACEAE Lindl. Nat. Syst. Ed. 2, 142 (1836). 

144. SPONDIACEAE Kunth, Ann. Sc. Nat. 2: 362 (1824). 
TEREBINTACEAE}}{ Juss. Gen. 368 (1789). 
TEREBINTHACEAE} DC, FI. Frang. 4: 613 (1805). 
ANACARDIACEAE Lindl. Nat. Syst Ed. 1 (1830). 

145. CYRILLACEAE Lindl. Veg. Kingd. 445 (1847). 

146. ILICACEAE Lowe, Fl. Madeir. 2: 11 (1868). 
AQUIFOLIACEAEt DC. Prodr. 2: 11 (1825). 

147. CELASTRACEAE Lindl. Nat. Syst. Ed. 2, 119 (1836). 

148. HIPPOCRATEACEAE HBK. Nov. Gen. 5: 136 (1821). 

149. STACKHOUSIACEAE Lindl. Nat. Syst. Ed. 2, 118 (1836). 

150. ICACINACEAE Miers, Ann. & Mag. N. H. Ser. 2, 9: 218 (1852). 
PHYTOCRENACEAE Miers; Lindl. Veg. Kingd. Ed. 3, 271a (1853). 
BARRERIACEAEt Martius, Consp. 41 (1835). 

151. STAPHYLEACEAE DC. Prodr. 2: 2 (1825). 
STAPHYLACEAE*® Reichb, Consp. 200 (1828). 
OcHRANTHACEAEt Lindl. Nat. Syst. Ed. 2, 78 (1836). 

152. ACERACEAE J. St. Hil. Expos. Fam. 2: 15 (1805). 
ACERINACEAE* KI]. & Grcke. Bot. Erg. Wald. 124 (1862). 

153. AESCULACEAE Lindl; Orb. Dict. 1: 155 (1841). 

Called HippocasTANaCEAE by Engler and Prantl. 

154. SAPINDACEAE R. Brown, Exp. Congo, App. 5, 427 (1818). 
PAULLINIACEAE HBK. Nov. Gen. 5: 99 (1821). 
DoDoNAEACEAE HBK. Nov. Gen. 5: 130 (1821). 

155. MELIANTHACEAE Endl. Gen. Supp. 5: 80 (1850). 

156. IMPATIENTACEAE (nom. nov.). 

BALSAMINACEAE} Dumort. Anal. Fam. 46 (1829). 

157- SABIACEAE Blume, Mus. Bot. Lugd. 1: 368 (1851). 
MILLINGTONIACEAE} Wight & Arn. Prodr. 1: 115 (1834). 
WELLINGTONIACEAE+} Meisn, Pl. Vasc. Gen. Comm. 207 (1840). 

158. FRANGULACEAE DC. FI. Franc. 4: 619 (1805). 

RHAMNEACEAE* D; Don, Prodr. Fl. Nepal. 188 (1825). 
RHAMNACEAE Dumort. Fl. Belg. 102 (1827).§ 
GouaNIACEAE Reichb, Consp, 145 (1828). 

PHYLICACEAE Agardh, Theor. Syst. Pl. 186 (1858). 

159. LEEACEAE DC. Prodr. 1: 635 (1824). 

VITACEAE Lindl. Nat. Syst. Ed. 2, 30 (1836). 


ee ee ge ees 


17 


160, ARISTOTELIACAE Dumort. Anal. Fam. 37, 41 (1829). 
ELAEOCARPACEAE Lindl. Nat. Syst. Ed. 2, 97 (1836). 

TILIACEAE Juss, Gen. 289 (1789). 

SPARMANNIACEAE Agardh, Theor. Syst. Pl. 260 (1858). 

162, MALVACEAE Neck. Act. Ac. Theod. Palat. 2: 488 (1770). 
GOETHEACEAE Reichb. Consp. 204 (1828). 

SIDACEAE Dumort, Anal. Fam. 46 (1829). 
MALVAVISCACEAE Presl, Relig. Haenk. 2: 1, 135 (1831). 
GossYPIACEAE Kerner, Pflanzenleb. 2: 681 (1891). 

163. BOMBACACEAE Schum.; Engl. & Pr. Nat. Pfl. 3: part 6, 53 (1890). 
BoMBACEAE* HBK. Noy. Gen. 5: 294 (1821). 

164. BUETTNERIACEAE (R. Brown) HBK. Nov. Gen. 5: 309 (1821). 
BuTTNERIACEAEt R. Brown, Flind. Voy. App. 3, 540 (1814). 
STERCULIACEAE IIBK. Nov. Gen. 5: 310 (1821). 
HERMANNIACEAE HBK. Nov. Gen. 5: 312 (1821). 
DoMBEYACEAE HBK. Nov. Gen. 5: 313 (1821). 
ByTTNERIACEAE} DC. Prodr. 1: 481 (1824). 
BUETTNERACEAE* Trattin. Gen. Nov. (1825). 

TRIPHACEAE*} Reichb. Handb. 291 (1837).§ 

165. DILLENIACEAE R. Brown, Flind. Voy. App. 3, 541 (1814). 
DELIMACEAEt DC, Syst. r: 396, 397 (1818). 

166. EUCRYPHIACEAE Gay, Bot. Zeit. 6: 130 (1848). 

167. OCHNACEAE DC. Ann, Mus. 17: 410 (1811). 
OCHNEACEAE* D, Don, Prodr. Fl. Nepal. 224 (1825). 
SAUVAGESIACEAE Dumort. Anal. Fam. 44, 49, (1829). 

168. CARYOCARACEAE Szysz.; Engl. & Pr. Nat. Pfl. 3: part 6, 153 (1893). 
RHIZOBOLACEAE} Lindl. Nat. Syst. Ed. 2, 76 (1836). 

169. MARCGRAVIACEAE Choisy; DC. Prodr. t: 565 (1824). 
MARGRAVIACEAE} Dumort. Anal. Fam. 43 (1829). 
MARCGRAAVIACEAE} Lindl. Nat. Syst. Ed. 1 (1830). 
NORANTEACEAE Martius, Consp. 61 (1835). 

QUIINACEAE Engler; Martius, Fl. Bras. 12, 1: 477 (1888). 
QUIINEACEAE* Choisy, Descr. Gutt. Ind. 12 (——).§ 

171. SCHIZOCHLAENACEAE (nom. nov.). 

Called CHLAENACEAE by Engler and Prantl. 
172. THEACEAE DC. Prodr. 1: 529 (1824). 
TERNSTROEMIACEAEt R. Brown, Abel Journ. App. 378 (1818). 
LaPLaceaE*t DC, Prodr. 1: 526 (1824). 
TERNSTROMIACEAE}t Agardh, Cl. Pl. 18 (1825). 
CAMELLIACEAE Dumort. Anal. Fam. 43, 47 (1829). 
173. STACHYURACEAE Gilg; Engl. & Pr. Nat. Pf. 3: part 6, 192 (1893). 


161. 


~ 


° 


170, 


- 174. SYMPHONIACEAE Presl, Symb. Bot. 1: 71 (1832). 


C.ustaceakE Lindl. Nat. Syst. Ed. 2, 74 (1836). 
HyPericaceak Lindl. Nat. Syst. Ed. 2, 77 (1836). 
CAMBOGEACEAE*t Horan. Tetract. Nat. 32 (1843)- 


* 


18 


Called GUTTIFERAE by Engler and Prantl. 

175. SHOREACEAE Roxb.; Wall. Catal. ». 4405 (1832).§ 
DIPTERACEAE*® Lindl. Nat. Syst. Ed. 2, 98 (1836). 

’ LopurrAceaE Endl. Gen. 1014 (1840). 
DirrEROcARPACEAE Eichl. Bluethendiagr. 2: 262 (1878). 

176. ELATINACEAE Lindl. Nat. Syst. Ed. 2, 88 (1836). 

177. FOUQUIERACEAE DC. Prodr. 3: 349 (1828). 
FouQuIERIACEAE* Dumort. Anal. Fam 27 (1829). 
REAUMURIACEAE G. Don, Gen. Syst. 3: 155 (1834). 
TAMARICACEAE Lindl. Nat. Syst. Ed. 2, 126 (1836). 
TAMARISCACEAE+ Lowe, Fl. Madeir. 46 (1868) 

178. FRANKENIACEAE §, F. Gray, Arr, Brit. Pl. 2: 663 (1821). 

179. CISTACEAE Lindl. Nat. Syst. Ed. 2, 91 (1836). 

180. BIXACFAE Reichb. Consp. 190 (1828). 

181. CANELLACEAE Martius, Nov. Gen. 3: 163 (1829): 

182, VIOLACEAE DC, FI. Franc. 4: 801 (1805), 

LrongEaAceakt A. DC. Prodr. 8: 668 (1844). 
LEONIACEAE (A. DC.) Agardh, Theor. Syst. Pl. 142 (1858). 

183. SAMYDACEAE Dumort. Anal. Fam. 16, 18 (1829). 
PAROPSIACEAE Dumort. Anal. Fam. 37, 42 (1829). 
FLACURTIACEAE} Dumort. Anal, Fam. 44, 49 (1829). 
FLACOURTIACEAE (Dumort.) Lindl. Nat. Syst. Ed. t (1830). 
KIGGELARIACEAE Link, Handb. 2: 221 (1831). 
BLACKWELLIACEAEt Schultz, Nat. Syst. Pfl. 444 (1832). : 
PATRISIACEAE Martius, Consp. 58 (1835). 

HoMALIACEAE Lindl, Nat. Syst. Ed. 2, 55 (1836). 
PANGIACEAE Lindl. Nat. Syst. Ed. 2, 70 (1836). 
184. TURNERACEAE HBK. Nov. Gen. G: 123 (1823). 


185. MALESHERBIACEAE D. Don, Edinb. N. Phil. Journ. 2: 320, 321 (1827)- 


186. PASSIFLORACEAE Dumort. Anal. Fam, 37, 42 (1829). 
MopeEccacEAE} Agardh, Theor. Syst. Pl. 386 (1838). 

187, CARICACEAE Dumort. Anal. Fam, 37, 42 (1829). 
PapavAceAEt Blume, Batav. Cour. (1823). 

188. LOASACEAE Reichh. Consp. 160 (1828). 
CEVALLIACEAE Griseb, Grundr. Syst. Bot. 136 (1854). 

189. BEGONIACEAE R. Brown, Exp. Cong. App. 5, 464 (1818). 

190. DATISCACEAE Dumort. Anal. Fam. 13, 14 (1829). 

191, OPUNTIACEAE HBK. Nov. Gen. 6: 64 (1823). 
Cacraceart Lindl. Nat. Syst. Ed. 2, 53 (1836), 
PERESKIACEAE+ Salm-Dyck, Otto & Dietr. Gartenz. 61 (1840).8 
LEUCHTENBERGIACEAE Salm-Dyck, Otto’s Gartenz. 188 (1854) § 

192, GEISSOLOMACEAE Sonder, Linnza 23: 105 (1850). 

193 PENAEACEAE Sweet, Hort. Brit. 488 (1826). 

194. OLINIACEAE Presl, Abh. Boehm, Ges, Folge 5, 3: 467 (1845).§ 
OLINACEAE* Kl. & Grcke. Bot, aig Wald. 152 (1862). 


octal 


195. 


196. 
197. 


198. 


203. 


204. 


205. 


206. 


207. 


19 


DAPHNACEAE J. St. Hil. Expos. Fam, 1: 180 (1805). 
THYMELEACEAE} Reichb. Consp, 82 (1828). 
AQUILARIACEAE Dumort. Anal. Fam, 15, 18 (1829). 
THYMELACEAE* Lindl, Nat. Syst. Ed. 1 (1830).g 
THYMELAEACEAE (Reichb.) Reichb. Fl. Exc. 164 (1831). 

ELAEAGNACEAE Lindl. Nat. Syst. Ed. 2, 194 (1836). 

LYTHRACEAE Lindl. Nat. Syst. Ed. 2, 100 (1836). 
LYTHRARIACEAE* Dumort. Anal. Fam. 36, 39 (1829). 
AMMANNIACEAE Horan. Tetract. Nat. 29 (1843). 

* CUPHEACEAE Kerner, Pflanzenleb. 2: 698 (1891), 
LAGERSTROEMIACEAE Kerner, Pflanzenleb. 2: 698 (1891). 

HENSLOWIACEAE (Lindl.) Martius, Consp. 14 (1835). 
HENSLOVIACEAE} Lindl. Bot. Reg. 20: ¢. 7686 (1834). 
CRYPTERONIACEAE A. DC. Prodr. 16, part 2: 677 (1868). 
BLATTIACEAE Niedenzu; Engl. & Pr, Nat. Pfl. 3: part 7, 16 (1892). 


. PUNICACEAE Horan. Tetract, Nat. 30 (1843). 
. NAPOLEONACEAE Dumort. Anal. Fam. 28, 29 (1829). 


BELVISIACEAE} Lindl. Nat. Syst. Ed. 1 (1830). 
LECYTHIDACEAE Lindl. Nat. Syst. Ed. 2, 523 (1836). 
BARRINGTONIACEAE Lindl. Veg. Kingd. 754 (1847). 


. RHIZOPHORACEAE Lindl. Nat. Syst. Ed. 2, 40 (1836). 


MYRTACEAE R. Brown, Flind. Voy. App. 3, 546 (1814). 
MyRTEACEAE* Nees, Noy. Act. Leop. 11, 1: 113 (1823). 
CHAMAELAUCIACEAE Lindl. Veg. Kingd. 721 (1847). 
LEPTOSPERMACEAE Kerner, Pflanzenleb. 2: 691 (1891). 

TERMINALIACEAE J. St. Hil. Expos, Fam. 1: 178 (1805). 
COMBRETACEAE R. Brown, Prodr, 1: 351 (1810). 

BLAKEACEAE Reichb. Consp. 174 (1828). 

MELASTOMACEAE*® R. Brown, Exp. Cong. App. 5, 434 (1818). 
RHEXIACEAE Martius, Consp. 64 (1835). 

MEMECYLACEAE® Lindl. Nat. Syst. Ed. 2, 40 (1836). 

MourIRIACcEAE Gardn. Hook. Journ. Bot. 2: 22 (1840). 

MIconiAceakt C. Koch, Berl. Gartenz. 241 (1857).§ 
MELASTOMATACEAE Krasser; Engl. & Pr. Nat. Pfl. 3: part 7, 130 (1893). 
CHARIANTHACEAE Kerner, Pflanzenleb. 2: 697 (1891). 

EPILOBIACEAE DC, Prodr. 3: 35 (1828). 

ONAGRACEAE Dumort. Anal. Fam. 36, 39 (1829). 
FucuHstaceak Dumort. Anal. Fam. 39 (1829). 
CIRCAEACEAE Lindl. Nat. Syst. Ed. 1 (1830). 
JUssIEUACEAE Drude, Phanerog. 385 (1879). 
OENOTHERACEAE Drude, Phanerog. 385 (1879). 

TRAPACEAE Dumort. FI. Belg. 90 (1827).§ 

Called HypROCARYACEAE by Engler and Prantl. 

GUNNERACEAE Endl. Gen. 285 (1837). 

HA.oraGaceAE*} Horan. Tetract. Nat. 25 (1843). 
HALORRHAGIDACEAE KI. & Grcke. Bot. Erg. Wald. E51 (1852). 
Hrppuripacear Sag. & Schn. Fl. Carp. Cent. 2: 23, 468 (1891). 


Aig 


20 


208. HEDERACEAE Linn. Ord. Nat. (1764). 
ARALIACEAE Vent. Tabl. 3: 2 (1799). 
PANACEAE* Reichb. Consp 144 (1828). 
HELWINGIACEAE Morren & Dec. Bull. Ac. Brux. 169 (1836). 
Hetvincraceart Agardh, Theor. Syst. Pl. 310 (1858). 
209. AMMIACEAE Presl, Delic. Prag. 1 (1822).§ 
SILERACEAE Presl, Delic. Prag. 1 (1822).§ 
BotacEaE® Reichb.; Moessl. Handb. 1: 45 (1827). 
APIACEAE Lindl, Nat. Syst. Ed. 2, 21 (1836). 
Called UMBELLIFERAE by Engler and Prantl. 
210, NYSSACEAE Dumort. Anal. Fam. 13 (1829). 
CoRNACEAE Link, Handb. 2: 2 (1831). 
GARRYACEAE Lindl. Bot. Reg, 20: ¢. 7686 (1834). 
ALANGIACEAE Lindl, Nat. Syst. Ed. 2, 39 (1836). 
AUCUBACEAE Agardh, Theor. Syst. Pl. 303 (1858). 
air. CLETHRACEAE Klotzsch, Linnea, 24: 12 (1851). 
212, PIROLACEAE (Agardh) Drude; Engl. & Pr. Nat. Pfl. 4, part 1: 3 (1889). 
PyRoOLaceAk+ Agardh, Cl. Pl. 18 (1825). 
MonotrRopACceaE Lindl. Nat. Syst, Ed, 2, 219 (1836). 
PyROLEACEAE*+ Brongn. Enum, Gen. 72 (1843). 
Hyporiryaceart Kl. & Grcke. Bot. Erg. Wald. 99 (1862). 
213. LENNOACEAE Solms-Laub. Abh, Nat. Ges. Halle, rr: 174 (1870): 
214. ERICACEAE DC. FI. Frang. 3: 675 (1805). 
RHODORACEAEt Vent. Tabl. 2: 449 (1799). 
VACCINACEAE® Lindl, Nat. Syst. Ed. 2, 221 (1836). 
VACCINIACEAE Lindl. Veg. Kingd. 757 (1847). 
MENZIESIACEAE Klotzsch, Linnzea 24: 11 (1851). 
SIPHONANDRACEAE Klotzsch, Linnza, 24: I1, 13 (1851). 
ARBUTACEAE Kerner, Pflanzenleb, 2: 671 (1891). 
OxycoccacEAEt Kerner, Pflanzenleb. 2: 713 (1891). 
215. STYPHELIACEAE Reichb, Consp. 127 (1828). 
RICHEACEAE Reichb, Consp. 128 (1828). 
SPRENGELIACEAE Reichb, Consp. 128 (1828). 
LYSINEMACEAE Reichb. Consp. 128 (1828). 
EPACRIDACEAE Lindl, Nat. Syst. Ed. 2, 222 (1836). . 
216. DIAPENSIACEAE Link, Handb, 1: 595 (1829). : 
GALACEAE* DC, Prodr. 7, part 2: 776 (1839). 4 
217. ARDISIACEAE Juss. Ann. Mus. 15: 350 (1810). 
THEOPHRASTEACEAE* D, Don; Lindl. Bot. Reg. an: ¢. 7764 (1835): 
MYRSINACEAE Lindl. Nat. Syst. Ed. 2, 224 (1836). 
MyRSINEACEAE G. Don, Gen. Syst. 4: 7 (1837). 
AEGICERACEAE* A. DC, Prodr. 8: 141 (1844). 
218. PRIMULACEAE Vent. Tabl. 2: 285 (1799). 
LYSIMACHIACEAE Reichb. Consp, 127 (1828). 
ANDROSACEAE* Reichb. Consp, 128 (1828). 
HorToniaceak Reichb. Fl. Exc. 398 (1831). 
219. ARMERIACEAE racic Comm, Pee: 61 (1822). 


220, 
221. 


222. 
223. 


224. 


Pe 


226. 


227. 


228. 


229. 


230. 


231. 


23 2. 


233. 


21 


PLUMBAGINACEAE Lindl. Nat. Syst. Ed. 2, 269 (1836). 
STATICACEAE Trautv. Bull. Ac. Pet. 14: 250 (1856). 
BUMELIACEAE (nom. nov.) 
SAPOTACEAEt Reichb. Consp. 135 (1828). 
DIOSPYRACEAE Drude, Phanerog. 377 (1879). 
EBENACEAE} Vent. Tabl. 2: 443 (1779). 
SYMPLOCACEAE Miers; Lindl. Veg. Kingd. Ed. 3, 593 (1853). 
HALESIACEAE Link, Hanbd. 1: 667 (1829). 
STYRACEAE* Reichb.; Moessl. Hanbd. 1: xlii (1827).§ 
STYRACACEAE A. DC, Prodr. 8: 244 (1844). 
OLEACEAE Lindl. Nat. Syst. Ed. 1 (1830). 
LILACEAE Vent. Tabl. 2: 306 (1799). 
JASMINACEAE Lindl. Nat. Syst. Ed. 2, 308 (1836). 
BOLIVARIACEAE Griseb. Gentian. 20 (1836).§ 
JASMINEACEAE * G. Don, Gen. Syst. 4: 58 (1837). 
SYRINGACEAE Horan. Tetract. Nat. 27 (1843). 
SALVADORACEAE Lindl. Nat. Syst. Ed. 2, 269 (1836). 
AZIMACEAE Wight & Gardn. Calcutta Journ. 6: 52 (1845).§ 
SPIGELIACEAE Martius, Nov. Gen. 2, 2: 132 (1827). 
STR YCHNEACEAE * Blume, Bijdr. n. 16: 1018 (1826).$ 
LOGANIACEAE Dumort. Anal. Fam, 21, 26 (1829). 
-POTALIACEAE Dumort. Anal. Fam. 21, 26 (1829). 
STRYCHNACEAE Link, Handb. 1: 439 (1829). 
FAGRAEACEAE Meisn Pl. Vasc. Gen. 167 (1839). 
GENTIANACEAE Dumort. Anal, Fam. 20, 25 (1829). 
MENYANTHACEAE G, Don, Gen. Syst. 4: 167 (1837). 
ERYTHRAEACEAE Griseb. Gen. et Sp. Gent. 69 (1839)-$ 
CHIRONIACEAE Horan. Tetract. Nat. 27 (1843). 
EXACEAE * Benth. & Hook. Gen. Pl. 2: 800 (1876). 
APOCYNACEAE Lindl. Nat. Syst. Ed. 2, 299 (1836). 
Vincaceae Horan. Tetract. Nat. 27 (1843). 
STAPELIACEAE Reichb.; Moessl. Handb. 1: 40 (1827).§ 
ASCLEPIADACEAE Lindl. Nat. Syst. Ed. 2, 302 (1836). 
HoyaceaeE G. Don, Gen. Syst. 4: 107 (1837). 
CONVOLVULACEAE Vent. Tabl. 2: 394 (1799). 
DICHONDRACEAE Dumort. Anal. Fam. 20, 24 (1829). 
CuscuTAcEAE Dumort. Anal. Fam. 20, 25 (1829). 
PORANACEAE Agardh. Theor. Syst. Pl. 364 (1858). 
POLEMONIACEAE DC. FI. Franc. 3: 645 (1805). 
POLEMONACEAE * Vent. Tabl. 2: 398 (1799). 
CopeaceaE+ D. Don, Edinb. Phil. Journ. ro: 111 (1824). 
COBAEACEAE Dumort. Anal. Fam. 20 (1829). 
HYDROLEACEAE HBK. Nov. Gen. 3: 125 (1818). 
HyprovaEAceaEt Dumort. Anal. Fam. 20, 25 (1829). 
RomanzoviacEAE} Dumort. Anal. Fam. 26 (1829). 
HyYDROPHYLLACEAE Lindl. Nat. Syst. Ed. 2, 271 (1836). 
EHRETIACEAE Schrad. Diss. Asperif. 20 (1820).§ 


22 


CorDIACEAE Dumort. Anal. Fam, 20, 25 (1829). 
BorAGINACEAEF} Lindl. Nat. Syst. Ed. 2, 274 (1836). 
ONOSMACEAE Horan. Tetract. Nat. 28 (1843). 
BorRAGINACEAE (Lindl.) A. Gray, Man. Ed. 2, 319 (1856). 
234. VERBENACEAE J. St. Hil. Expos. Fam. 1: 245 (1805). 
VeERBACEAE® Link, Enum. 1: 174 (1821). 
STILBACEAE Lindl. Nat. Syst. Ed. 2, 280 (1836). 
PHRYMACEAE Schauer; DC. Prodr. 11: 520 (1847). 
DuRANTACEAE Agardh, Theor. Syst. Pl. 295 (1858). 
PETRAEACEAE Agardh, Theor, Syst. Pl, 364 (1858). 
235. LAMIACEAE Lindl. Nat. Syst. Ed. 2, 275 (1836). 
NeEPETACEAE Horan, Tetract. Nat. 28 (1843). 
SALVIACEAE Drude, Phanerog. 374 (1879). 
Called LABIATAE by Engler and Prantl. 
236. NOLANACEAE Dumort. Anal. Fam. 20, 24 (1829). 
237. SOLANACEAE Pers Ench. 1: 214 (1805). 
RETZIACEAE Bartl. Ord. Nat. 192 (1830) 
CrSTRACEAE Lindl. Nat. Syst. Ed. 2, 296 (1836). 
SCLEROPHYLACEAE*® Miers, Lond. Journ. Bot. 7: 57 (1848). 
ATROPACEAE Miers, Ann. & Mag. N. H. Ser, 2, 3: 163 (1849). 
238. RHINANTHACEAE J. St. Hil. Expos, Fam, 1: 227 (1805). 
MELAMPYRACEAE Dumort. Fl Belg. 32 (1827).§ 
LINDERNIACEAE Reichb. Consp. 123 (1828). > 
CAPRARIACEAE Reichb. Consp. 124 (1828). 
HALLERIACEAE Link, Handb, 1: 506 (1829). 
SCOPARIACEAE Link, Handb. 1: 822 (1829). 
ARAGOACEAE D, Don, Edinb, N. Phil. Journ. rg: 113 (1835). 


S1BTHORPIACEAE D, Don, Edinb. N. Phil. Journ. 1g: 114 (1835)- 


SELAGINACEAE Lindl. Nat. Syst. Ed, 2, 279 (1836). 
SCROPHULARIACEAE Lindl, Nat. Syst. Ed. 2, 288 (1836). 
SCROFULARIACEAE} Horan. Tetract. Nat. 27 (1843). 
VERONICACEAE Agardh, Theor. Syst. Pl. 392 (1858). 
ERINACEAE Pfeiff. Nomencl. Bot, 1: 1236 (1874). 

_ 239. PINGUICULACEAE Dumort. Anal. Fam. 19, 23 (1829). 

UTRICULARIACEAE Dumort. Anal. Fam. 19, 23 (1829). 
LENTIBULARIACEAE} Lindk Veg. Kingd. 686 (1847). 

240. OROBANCHACEAE Lindl. Nat. Syst. Ed. 2, 287 (1836). 
LATHRAEACEAE Walp. Ann. Bot. 3: 204 (1853). 

241. CYRTANDRACEAE Jack, Linn. Trans. 14, 1: 23 (1823). 
GESSNERIACEAE} Nees, Ann. Sc. Nat. 6: 295 (1825). 
GESNERIACEAE (Nees) Dumort Anal. Fam. 28, 30 (1829). 
GESNERACEAE* Lindl. Nat. Syst. Ed. 2, 283, 286 (1836). 

af RAMONDIACEAE Gren. & Godr, Fl. Franc. 2: 506 (1850.)$ 

242, COLUMELLIACEAE Lindl. Nat. Syst. Ed. 1 (1830). 


243. BIGNONIACEAE Pers. Ench. 2: 168 (1807) 


- CRESCENTIACEAE Dumort. Anal. Fam. 20, 24 (1829). 
: ~~ MARTYNIACEAE Link, Handb, 1: 504 


23 


PEDALIACEAE Lindl. Nat. Syst. Ed. 2, 281 (1836). 
SFSAMACEAE Drude, Phanerog. 373 (1879). 

245. GLOBULARIACEAE Dumort. Anal. Fam. 19, 21 (1829). 
GLOBULACEAE* Dumort. Comm, Bot. 55 (1822). 

246. ACANTHACEAE J. St, Hil. Expos. Fam. 1: 236 (1805). 

247. MYOPORACEAE Lindl. Nat. Syst. Ed. 2, 279 (1836). 
BonTIACEAE Horan. Tetract. Nat. 27 (1843). 

248. PLANTAGINACEAE Lindl. Nat. Syst. Ed. 2, 267 (1836). 

249. RUBIACEAE B. Juss. Hort. Trian. (1759). 

CoFFEACEAE Batsch, Tab. Affin. 233 (1802). 
CINCHONACEAE DC. Ann. Mus. 9: 217 (1807). 
GUETTARDACEAE DC. Ann. Mus. g: 217 (1807). 
GARDENIACEAE HBK. Nov. Gen. 3: 407 (1818). 
HAMELIACEAE HBK. Nov. Gen. 3: 413 (1818). 
PsYCHOTRIACEAE Cham. & Schlecht. Linnaea 4: 4 (1829). 
OPERCULARIACEAE Dumort, Anal. Fam, 29, 32 (1829). 
LyGopysopDEacEAE}t Bartl. Ord. Nat. 123, 207 (1830). 
LyGopDysIACEAE*}} Martius, Consp. 31 (1835). 
GALIACEAE Lindl. Nat. Syst. Ed. 2, 249 (1836). 
NAUCLEACEAE Meisn, Pl. Vasc. Gen. 157 (1838). 
MORINDACEAE Schimp. Paleont. Veg. 2: 874 (1872). 

250. VIBURNACEAE Dumort. Comm. Bot. 56 (1822). 
CAPRIFOLIACEAEt Vent. Tabl. 2: 593 (1799). 
LinneEACEAEt Dumort. Fl. Belg. 55 (1827).§ 
LINNAEACEAE Dumort. Anal. Fam. 33 (1829). 
LoNICERACEAE Drude, Phanerog. 370 (1879). 
SAMBUCACEAE Kerner, Pflanzenleb. 2: 711 (1891). 

251. ADOXACEAE Fritsch; Engl. & Pr. Nat. Pf. 4, part 4: 170 (1891). 

252. VALERIANACEAE Batsch, Tab. Affin. 227 (1802).§ 

253. MORINACEAE Dumort. Anal. Fam. 32 (1829). 
DipsaceaE* B, Juss. Hort. Trian. (1759). 

DipsacacEak Lindl. Veg. Kingd. 699 (1847). 

254. CUCURBITACEAE B. Juss. Hort. Trian. (1759). 
ZANONIACEAE Blume, Bijdr. 15: 936 (1826). 
ZANNONIACEAE Dumort. Anal. Fam. 103 (1829). 

255. CAMPANULACEAE Juss. Gen. 163 (1789). 
LoBELIACEAE Dumort. Comm. Bot. 57 (1822). 
SPHENOCLEACEAE Martius, Consp. 31 (1835). 
DELISSEACEAE Presl, Prodr. Mon. Lobel. 46 (1836). 
CyPHIACEAE A. DC. Prodr. 7, part 2: 497 (1839). 
NEMACLADACEAE Nutt. Amer. Phil. Trans. N. Ser. 8: 254 (1843). 
CYPHOCARPACEAE Miers, Lond. Journ. Bot. 7: 61 (1848). 

256. BRUNONIACEAE Reichb. Consp. 91 (1828). 
GOoDENIACEAE Dumort. Anal. Fam. 28, 30 (1829). 
SCAEVOLACEAE Lindl. Nat. Syst. Ed. 2, 242 (1836). 
GoODENOUGHIACEAE + Schum.; Martius, Fl. Bras. 3, part 3: 161 (1894). 

257. CANDOLLEACEAE Schoenl.; Engl. & Pr. Nat. Pfl. 4, part 5 : 79 (1889). 


> 


24 


STYLIDIACEAE ¢ Lindl. Nat. Syst. Ed, 2, 240 (1836). 


258. CALYCERACEAE Lirdl. Nat. Syst. Ed. 2, 251 (1836). 
259. CARDUACEAE Neck. Act. Ac. Theod. Palat. 2: 465 (1770). 


15. ELODEACEAE, L£lodea Michx. is known as Udora Nutt. 


CICHORACEAE * B, Juss. Hort. Trian. (1759). 
CHICORACEAE * + Neck. Act. Ac. Theod. Palat. 2: 463 (1770). 
GUNDELIACEAE DC. Ann. Mus. 16: 153 (1810). 
VERNONIACEAE HBK. Novy. Gen. 4: 39 (1820). 

INULACEAE Presl, Delic, Prag. 1 (1822).§ 

JACOBACEAE* ¢ Dumort. Fl. Belg. 65 (1827).§ 
GRINDELIACEAE Reichb. Consp. 107 (1828). 

PsIADIACEAE Reichb. Consp. 107 (1828). 

CALENDULACEAE Reichb. Consp, 112 (1828). 
AMBROSIACEAE Reichb. Consp, 112 (1828). 

IvaceaE Reichb. Consp. 112 (1828). 

CYNARACEAE Dumort. Anal. Fam. 32 (1829). 
ACARNACEAE ¢ Link, Handb, 1: 684 (1829). 
EuPAToRIACEAE Link, Handb. 1: 729 (1829). 
PARTHENIACEAE Link, Handb. 1: 816 (1829). 
HIERACEAE* D, Don, Edinb. N. Phil. Journ. 6: 306 (1829). 
TARAXACEAE* D. Don, Edinb, N. Phil. Journ. 6: 307 (1829). 
CICHORIACEAE Reichb, Fl. Exc. 248 (1831). 

MUTISIACEAE Lessing, Syn. Compos. 92 (1832). 
NASSAUVIACEAE Lessing, Syn. Compos. 396 (1832). 
ASTERACEAE Lindl. Nat. Syst. Ed. 2, 251, 253 (1836). 
NASSAVIACEAE} Endl. Gen. Suppl. 1: 1386 (1841). 
EVACEAE*® Schultz-Bip.; Walp. Repert. 2: 955 (1843). 
CassINIACEAE Schultz-Bip. Flora, 1: 129 (1852). 
CENTAUREACEAE Pfeiff. Nomencl. Bot. 1: 646 (1873). 
HELIANTHACEAE Pfeiff. Nomencl. Bot. 1: 1579 (1874). 
LacTucaceaE Drude, Phanerog. 369 (1879). 

Called ComposirTAaE by Engler and Prantl, 


NOTES ON A FEW OF THESE NAMES, 


Hence this family 


bocomes VALLISNERIACEAE Dumort. Anal. Fam. 54, 55 (1829). 


94. CALYCANTHACEAE. Calycanthus L. should be called Buettneria Duham. 


(see Kearney, Bull. Torr. Club, 21: 173). As this family contains but one 


genus, it must be called BUETTNERIACEAE (nom. nov.), 


164. BUETTNERIACEAE,, The name Auettneria Loefl. being no longer tenable, 


this family should be called STERCULIACEAE HBK. Nov. Gen. 5: 310 


(1821). 


25 


A Revision of the North American species of the genus Cracca. 


By ANNA MurRAyY VAIL. 


The genus Cracca was established by Linnaeus first in Fl. Zey. 
139-141 (1747) and then in Sp. Pl. 752 (1753), the genus being 
based on Cracca Virginiana. In the Sp. Pl. Ed. 2, 1062 (1763) 
it was Galega Virginiana, and after that the synonymy becomes 
more and more complicated and names for the genus appeared in 
quick succession among which are: Co/inil Adans. 1763; Need- 
hamia Scop. 1777; Brissonia Neck. 1790; Reinera Moench, 1802; 
and finally Zephrosia Pers. 1807, under which all the Linnaean 
species of Cracca have been described until 1891, when Kuntze in 
Rev. Gen. Pl. 173 transferred them all to the original generic title. 

Bentham in Oerst. Kjoeb. Vidensk. Meddel, 8 (1853) estab- 
lished a genus Cracca based on a West Indian species, Galega 
Caribaea, Jacq. Am. 212, ¢. 725 (1781). The six known spe- 
cies of which genus have been transferred by Kuntze, Rev. Gen. PI. 
164 to the genus Brittonamra. 

Bentham and Hooker in 1867 gave the number of the spe- 
cies of Tephrosia as 90, of which some 16 are ascribed to 
Africa and America. Taubert, in Engler & Prantl Nat. Pfl. part 
tor: 269 (1894), gives 120 as the total of species, ascribing 
few to America. The following revision is an attempt to 
clearly describe especially the rather difficult Southern State spe- 
cies. As far as is known twelve species are native within the 
boundaries of the United States and two C. purpurea and C. cinerea 
are cosmopolitan plants, the latter known in North America only 
from ballast ground in Alabama. The Mexican and _ tropical 
American species are as yet imperfectly known and are more 


numerous than it is supposed.* 
The genus is accepted as described by Bentham and Hooker, 


a 


under Zephrosia. 


* The following species appears to be undescribed : 


Cracca ScHortir n, sp. = 
Perennial from a somewhat woody base, more or less cinereous or silvery-strigdse 
throughout. Stems branching, angled, 3 dm. or more high, erect or decumbent. sti- 
pules 4-7 mm. long, subulate, persisting; petioles 1-25 cm. SS 4-7 es 
long, obovate-oblong in outline; leaflets 5-7, obovate or obovate-oblong, I~2.5 - 
long, 5-15 mm. wide, retuse, minutely apiculate, strigillose above, silvery or cinereo 


26 


ARTIFICIAL KEY TO THE NORTH AMERICAN SPECIES. 


Flowers few or crowded in oblong panicles or racemes at the summit of the simple or 
branched stems (somewhat elongated and spicate in C. devocarpa). 
Leaflets linear-oblong or elliptical, panicles sessile or short-peduncled. 
Stems erect, simple ; pubescence on the sessile, crowded panicle silvery ; legume 
cinéreous—pubescent-or villous . 2... ee a 1. C. Virginiana, 
Stems erect, branching ; pubescence on the sessile or short-peduncled, crowded 
panicle tawny; legume rusty or tawny, villous or tomentose. . 2. C. eucantha. 
Stem erect, sub-simple or branching; pubescence on the short-peduncled pani- 
cle appressed-cinereous; legume yellowish, glabrous. . . 3. C. dedocarpa. 
Leaflets obovate or sub-orbicular; panicles long-peduncled. 
Stems prostrate or ascending; legume yellowish, velvety pubescent. 
4. C. Lindheimeri. 
Flowers fascicled or more or less remote, forming a somewhat spicate long-peduncled 
inflorescence. 
Stems erect, pilose with mostly spreading rusty hairs; leaflets g-25 cuneate- 
EE OT  ae  Seae  A Surtees Kee TS PRL Ae 5. C. onobrychotdes. 
Stems decumbent or ascending, pilose with reflexed or spreading rusty hairs; 
leaflets 5-15, oval or oblong, or in the variety linear-oblong. , . 6. C. spicata. 
Flowers scattered, single or geminate, forming a slender, elongated, spicate inflor- 
escence, much exceeding the leaves. . . . ......2...-- 7. C, purpurea. 
Flowers single or geminate, scattered at the summit of slender ancipital peduncles, 
mostly exceeding the leaves (often shorter in C. ambigua); flowers white, turning 
reddish or purple. 
Petioles much longer than the leaflets. 
Stems erect, very slender, flexnous ; leaflets linear. . . 8. C. angustissima. 
Stems trailing or assurgent; peduncles and leaves erect; leaflets oblong or 
CHECALCODION DY oi5 475 es Soe i Re ee 9. C. ambigua. 
Petioles the length of or shorter than the leaflets (leaves sessile or nearly so in 
C. chrysophylle). 
Stems decumbent or assurgent, slender; leaflets mostly acute at each end, 
POHEKOCL Oe ee es re 10. C. hispidula. 
Stems assurgent, leaflets oblong-obovate. 11. C. Smallit. 
Stems prostrate, spreading. 
Leaflets 1-7 cuneate-obovate, silky-hirsute beneath. 
12. C. chrysophylla. 


Ne Me aes LR ae SAT ae kee he pe A eS tae “glee Aa eg 


PE Bee ee SE 


strigose and somewhat glaucous beneath, the terminal one usually considerably 
_ larger than the lateral ones; racemes 1 dm. or less long; flowers few, 6 or 7 mm. 
__ long, scattered along the slender ancipital peduncle; bracts minute, setaceous ; calyx- 
_ teeth setaceous, as long as the tube; corolla rosé-purple; vexillum minutely hirsute ; 
legume 3-5 cm. long, 4 mm. wide, straightish, strigose; seeds 5-6, oblong-ovoid, 
truncate at the ends, brownish. . 
_ Nearest to C. purpurea, from which it differs in the apparently constant broadly 
obovate leaflets. 
< U. S. Colombia, Cartajena, Schott, Leguminose No. 16; Ruatan Island, Bay of 
Honduras, G. F. Gaumer, No. 50, 1886. 


__ Types in Herb, Columbia College (Schott.) and Herb. U. S. Depart. Agric. 


27 


Leaflets 5—13-oblong or oblong-obovate, strigose beneath. 
13. C. Floridana, 


Flowers few, in short-peduncled somewhat spicate racemes, exceeding the leaves. 
Stems prostrate or ascending; leaflets, 13-17, oblong or linear-oblong, cinereous- 
PuOercenh ce i aes a ee 14. C. cinerea, 


I. CRACCA VIRGINIANA L, 

Cracca Virginiana L. Sp. Pl. 752 (1753). 

Galega Virginiana L. Sp. Pl. Ed. 2: 1062 (1763). 

Tephrosia Virginiana Pers. Syn. 2: 329 (1807). 

Perennial from a woody base and long, tough, ligneous root, 
villous-pubescent or canescent throughout. Stems growing in 
patches, simple, erect, 3-6 dm. high, angled and striate; leaves 
elliptical or linear-oblong in outline, 6-10 cm. or more long, sub- 
sessile; stipules caducous; leaflets 11-21, 1-3 cm. long, linear- 
oblong or elliptical, obtuse or acutish, apiculate, glabrous or nearly 
so above, silky-villous beneath, often becoming-glabrate with age; 
raceme terminal, oblong, sessile; pedicels 5-10 mm. long; calyx 
silky-villous, the teeth acuminate, cuspidate, mostly longer than 
the tube; corolla cream-colored, streaked with purple or pink; 
vexillum pubescent on the outer surface; legume 3-5 cm. long, 
4-5 mm. wide, straight or somewhat falcate, villous; seeds 4-8 
oblong-ovoid, rounded or truncate at the ends, brownish, mottled 
with black. ; 

Dry sandy soil, throughout the eastern half of the United 


States and Canada to North Mexico. 
Original locality: Virginia, Canada. 
Type in Herb. Linn. 

CRACCA VIRGINIANA HOLOSERICEA (Nutt). 

Tephrosia holosericea Nutt. Journ. Acad. Phila. 7: 105 (1834). 

Tephrosia Virginiana var. holosericea Torr. & Gray, Fl. N. Am.. 
I: 296 (1838 . 

Stems and racemes densely villous; leaflets sericeous on both 
sides, often sub-lanceolate and acute; legume densely villous or 
tomentose, 

Arkansas (Nuttall), Illinois (Pitcher), Wisconsin (Liders), 
Louisiana (Hale). 

Original locality: Plains of Arkansas. 


2. CRACCA LEUCANTHA (H. B. K.) Kuntze. 
Tephrosia leucantha H. B. K. Nov. Gen. 6: 460, 4 577 (182 3). 
Cracca leucantha Kuntze, Rev. Gen. Pl. 175 (1891). 


28 


Perennial from a stout root, cinereous or rusty villous-pubes- 
cent throughout. Stems 3-6 dm. or more high, erect, branching, 
angled and striate; leaves elliptical in outline, 1 dm. or more 
long; stipules setaceous, caducous; petioles 1-2 cm. long, leaflets 
19-25, 2-3 cm. long, oblong, obtuse, often retuse, apiculate, 
the terminal one often obovate, minutely pubescent above, silky, 
cinereous-pubescent or villous beneath; raceme oblong, sessile 
or short peduncled; pedicels 5-8 mm. long; bracts 5-8 mm. long, 
villous; flowers 1.5 cm. long; calyx villous, tawny or rusty, the 
teeth acuminate, shorter, or as long as the tube, corolla cream- 
colored(?), vexillum pubescent on the outer surface; legume 3-4 
cm. long, straight or somewhat falcate, spreading, rusty-villous or 
tomentose; seeds 6-9, oblong-ovoid, pale greenish, smooth. 


Very close to C. Virginiona, from which it differs in the branch- 
ing stems, more numerous leaflets, and the tawny pubescence of 
the inflorescence and legumes. 

Arizona, New Mexico, Mexico. 

Original locality: Near Guanajuato, South Mexico. 


3. CRACCA LEIOCARPA (A, Gray) Kuntze. 


Tephrosia lewcarpa A. Gray, Pl. Wright. 2: 36 (185 3). 
Cracca letocarpa Kuntze, Rev. Gen. Pl. 175 (1891). 


Perennial from a thick, lignescent root, appressed cinereous- 
pubescent. Stems 3 dm. or more high, many from the same root, 
sub-simple, angled and striate; stipules 4-6 mm. long, setaceous, 
persisting ; petioles t-3cm. long; leaves linear-oblong in outline ; 
leaflets 17-12 or more, linear-oblong, 2-4.5 cm. long, 5-7 
mm. wide, obtuse or acutish, apiculate, glabrous above, sericeous 
and cinereous beneath; racemes terminal and axillary, short- 
peduncled, exceeding the leaves; bracts caducous; pedicels 5-8 
mm. long; flowers 2 cm. or less long; calyx cinereous, the subulate 
teeth as long as the tube; corolla purplish; vexillum minutely 
pubescent ; legume 3-5 cm. long, 6 mm. wide, straight, yellowish, 
glabrous; seeds about 10, sub-orbicular or ovoid, brownish. 
Mature seeds not seen. 

New Mexico, Arizona, Mexico. 


Original locality: New Mexico, Wright (No. 965) and North 
Mexico. 


Authentic specimen in Herb. Columbia College. 


4. Cracca LINDHEIMERI (A. Gray) Kuntze. 
Tephrosia Lindheimeri A. Gray, Bost. Journ. Nat. Hist. 6: 172 
(1850). , ; 


29 


Cracca Lindheimeri Kuntze, Rev. Gen. Pl. 175 (1891). 


Perennial from a tuberous and ligneous root, cinereous-pubes- 
cent or sericeous throughout. Stems prostrate or ascending, rather 
stout, flexuous, branching, spreading, 6-12 dm. long, more or less 
angled ; leaves oblong, linear-oblong or obovate-oblong in outline; 
stipules somewhat persisting; leaflets 9-17, roundish-obovate, 
nearly sub-orbicular, or obovate-cuneate, 23 cm. long, apiculate, 
pubescent above, densely sericeous or velvety-pubescent beneath ; 
racemes loosely many-flowered; peduncle 1-2 dm. long; flowers 
1.5—2 cm. long ; pedicels and bracts 5-8 mm. long; calyx sericeous, 
the acuminate teeth about the length of the tube; corolla red- 
purple, vexillum pubescent; legume 3-4 cm. long, 5-9 mm. broad, 
undulate, densely velvety-pubescent, tawny or yellowish; seeds 
about 4, broad, nearly orbicular, brownish. 


Texas, New Mexico, North Mexico. 
Original locality, Muskit prairies, on the Liano, Lindheimer 


(No. 592). 
Type in Herb. Columbia College. 


5. CRACCA ONOBRYCHOIDES (Nutt.)- Kuntze. 


Tephrosia onobrychoides Nutt. Journ. Acad. Phila. 7: 104 (1834). 

Tephrosia angustifolia and T. multiflora Featherman, Bot. Rep. 
Lousiana Univ. 73 (1871). 

Cracca onobrychoides Kuntze, Rev. Gen. Pl. 175 (1891). 


Perennial from a somewhat woody base, more or less pilose 
with spreading rusty hairs throughout. Stems erect, somewhat 
stout, simple or branching, flexuous, striate and angled above ; 
stipules 8-12 mm. long, often persisting; petioles 1-3 cm. long; 
leaves oblong or linear-oblong in outline, 1-1.6 dm. long; leaflets 
13-25, 2-5 cm. long, cuneate-obovate or oblong, obtuse, 
often retuse, apiculate, pubescent or glabrate above, more or 
less closely silky-pilose beneath; raceme terminal or axillary, 
rusty-villous or rarely glabrate, 3-6 dm. long; bracts 5 mm. 
long, subulate, mostly caducous; pedicels slightly longer than 
the bracts; flowers I-1.§ cm. long, 2-several together on al- 
ternate nodes often the whole length of the peduncle; upper 
calyx-teeth shorter than the tube, the lower one subulate and 
longer: corolla white turning to pale scarlet; vexillum pubescent 
on the outer surface, green toward the middle; legume 3-5 cm. 
long, 5 mm. wide, linear, acute, somewhat falcate, spreading, 
puberulent; seeds 6-10, round-oblong. Mature fruit not seen. 


Arkansas, Indian Terr., Louisiana, Alabama, Texas. 
Original locality: In the plains of Arkansas. 


30 


6. Cracca spicata (Walt.) Kuntze. 
Galega spicata, Walt. Fl. Car. 188 (1788). 
Galega villosa Michx. Fl. Bor. Am. 2: 67 (1803). 
Tephrosia villosa Pers. Syn. 2: 329 (1807). 
Tephrosia paucifolia Nutt. Gen. 2: 119 (1818). 
Tephrosia hispida DC. Prodr. 2: 250 (1825). 
Galega paucifolia Curtis, Bost. Journ. Nat. Hist. 1: 122 (1837): 
Tephrosia spicata Torr. & Gray, Fl. N. Am. 1: 296 (1838). 


Tephrosia: mollissima Bertol. Bot. Misc. g: 10, z. 3. (Bot. Zeit. > 


g: 902) fide A. Gray, Am. Journ. Sci. (2) 14: 115 (1852). 
Cracca spicata Kuntze, Rev. Gen. Pl. 175 (1891). 


Perennial from a long, stout root, more or less rusty-hirsute, of 
villous throughout. Stems decumbent or ascending, simple or 
diffusely branching, flexuous, spreading, somewhat angled above, 
clothed with mostly spreading or reflexed rusty hairs and a shorter, 
retrose, appressed pubescence, often becoming glabrate with age 
below; stipules 5-10 mm. long, sometimes persisting ; leaves 5 
_cm.—1.5 dm. long, oblong or linear-oblong in outline, the upper 
ones subsessile, the lower short petioled (1-3 cm.); leaflets 5-15, 
1-2.5 cm. long, 7-12 mm. wide, oval, oblong or cuneate- 
oblong (the terminal ones often obovate), obtuse, mucronate, often 
reflexed, minutely pubescent, silky-villous or glabrous above, rusty- 
villous beneath, often reflexed; racemes terminal and axillary; 
peduncles ancipital, ;-3 dm. long: bracts 5-10 mm. long, subulate, 
mostly persisting; flowers I-1.5 cm. long, remote, few or several 
together at the summit of the peduncle; calyx-teeth subulate, 
hispid, slightly longer than the tube; corolla white, turning 
purple; vexillum pubescent; legumes 3-5 cm. long, 5-6 mm. 
wide, linear, acute, straightish ; seeds 8-12, ovoid, brown. 

Differing from C. onobrychoides in its more slender and de- 
cumbent habit, fewer flowers and less silky pubescence. Growing 
in low sandy pine lands mostly among grasses, notable for its 
ascending stems and erect peduncles. Depauperate specimens 
with the flowers in the axils of the upper leaves have been 


collected in Florida. Virginia to Florida, westward to Tennessee 
and Mississippi. 

Original locality not given. 

Type in Herb. Walter. 
CRACCA SPICATA FLEXUOSA (Chapm.) 


«Tephrosia flexuosa Chapm.; Torr. & Gray, Fl. N. Am. 1: 297 
(1838). : ; 


:% 


Kg a ccpcioiebaatbe Really De Si PROLAPSE OS 


31 


Lephrosia hispidula y Torr. & Gray, Fl. N. Am. 1: 297 (1838). 

Stems becoming glabrate; leaflets 2-7 pairs, linear, acute, 
emarginate, reflexed, the terminal one much elongated. 

Florida and Alabama. 

Original locality : Florida (Chapman). 

Type in Herb. Columbia College. 


7. CRACCA PURPUREA L, 


Cracca purpurea L. Sp. Pl. 752 (1753). 

Galega piscatoria Ait. Hort. Kew. 3 : 71 (17869). 

Tephrosia leptostachya DC. Prodr. 2: 251 (1825). 

Lephrosia adscendens Macfad. Fl. Jam. 257 (1837). 

Lephrosia tenella A. Gray, P|. Wright, 2: 36 (1853). 

Annual or perennial from a slender woody base, glabrate or 
strigillose. Stems 1-6 dm. high, erect or ascending, branching, 
spreading, strigillose on the angles, often glabrate or glabrous and 
glaucous, striate above; stipules setaceous, often persisting; peti- 
oles 1-3 cm. long; leaves 6 cm.—1 dm. long, oblong or linear-ob- 
long in outline; leaflets 7-17, 2-5 cm. long, linear, linear-ob- 
long or oblong-cuneate, obtuse, or acute at each end, apicu- 
Jate, glabrous above, strigillose or glabrate, lighter and often 
glaucous beneath ; racemes terminal and axillary, 1-2 dm. long ; 
peduncles ancipital; flowers 510 mm. long, very short pedicelled ; 
bracts setaceous ; calyx-teeth as long as the tube; corolla white, 
turning purple; vexillum minutely pubescent; legume 3-5 cm. 
long, 3-4 mm. wide, linear or slightly falcate, strigillose or gla- 
brate ; seeds 6-10, oblong brown. 

A very variable species with a long and complicated list of sy- 
nonyms. Known everywhere in tropical and subtropical coun- 
tries. Two distinct strains have been collected in the United 
States, the broader-obtuse leaved form occurs in East Florida, and 
the narrow acute-leaved form (C. tenella) in Texas, Arizona, etc. 

O. Kuntze (Rev. Gen. Pl. 173) has united c. purpurea and 
several other species to C. villosa L. The latter has a villous 
calyx with long, acuminate teeth and a short reflexed, tomentose 
legume and is not known from America. 

Florida, Texas, New Mexico, Arizona, Mexico. Also in East- 
ern Central America, Eastern South America to South Brazil, and 
in the West Indies. 

_ Original locality: Ceylon. 


32 


8. CRACCA ANGUSTISSIMA (Shuttlw.) Kuntze. 


Tephrosia angustissima Shuttleworth ; Chapm. Fl. 96 (1860). 
Cracca angustissima Kuntze Rev. Gen. Pl. 174 (1891). 


Perennial from a somewhat woody base, very slender, glabrate 
or minutely pubescent. Stems 3-6 dm. long, prostrate, diffusely 
branching, flexuous; stipules 3-7 mm. long, setaceous, persist- 
ing; petioles 3-9 cm. long; leaves linear-oblong in outline; 
leaflets 5-17, 1-3 cm. long, 2-3 mm. wide, linear, acute at 
each end, minutely pubescent beneath; racemes terminal or axil- 
lary ; peduncles 5-10 cm. long; flowers 8-12 mm. long, solitary 
or geminate; bracts setaceous, persisting; pedicels 7-10 mm. 
long; calyx-teeth subulate, about the length of the tube; vexillum 
pubescent; legume 3 cm. or more, 4 mm. wide, minutely hirsute, 


somewhat inflated at maturity; seeds 6-8, ovoid, truncate at the 
ends, black. ' 


Pine barrens, South Florida, near Eau Gallie, Indian River, 
A. H. Curtiss, N. Am. Pl. No. 584. 


Original locality: South Florida, Rugel. 


g. Cracca Ampicua (M. A. Curtis) Kuntze. 


Galega ambigua Curtis, Bost. Journ. Nat. Hist. 1: 121 (1837). 
Tephrosia hispidula pg. Torr. & Gray Fl. N. Am. 1: 297 (1838). 
Liphrosia ambigua Chapm. F1. 96 (1860). 

Cracca ambigua Kuntze, Rev. Gen. Pl. 174 (1891). 


Perennial from a woody base, and long, ligneous root, hirsute 
with short spreading, somewhat viscid hairs. Stems 3-4 dm. or 
more high, trailing or assurgent, often purplish, dichotomously 
branching, straggling, angled; stipules 5-7 mm. long, lanceolate, 
persisting; petioles 3-8 cm. long; leaves 7 cm—1.5 dm. long, 
remote, linear-oblong in outline; leaflets 7-11, linear-oblong to 
cuneate-oblong or obovate, 2-14 cm. long, 5-15 mm. wide, acutish 
or mostly obtuse, apiculate, coriaceous, glabrous and yellowish- 
green above, appressed hirsute with whitish hairs and the veins 
often turning reddish or purplish beneath; peduncles 1-1.5 cm. 
long, ancipital; flowers few, 10-12 mm. long; bracts 5 mm. long, 
persisting; calyx-teeth subulate, as long as the tube; vexillum 
pubescent; legume 4 cm. or more long, 4 mm. wide, straightish ; 
seeds 8-13, ovoid, brown, variegated with black. 


High pine lands, dry sandy soil. Trailing, with ascending 


and almost erect leaves, leaflets and peduncles, or stems ascending — | 


with a somewhat bushy habit. 
North Carolina and Florida, westward to Mississippi. 


etl 


33 


Original locality : Sandy woods near Wilmington, North Caro- 
lina. 
Type in Herb. Columbia College. 


10, CRACCA HISPIDULA (Michx.) Kuntze. 


Galega hispidula Michx, F1. Bor. Am. 2: 68 (1803). 

Tephrosia hispidula Pers. Syn. 2: 329 (1807). 

Lephrosia gracilis Nutt. Gen. 2: 119 (1818). 

Lephrosia elegans Nutt. Journ. Acad. Phila. 7; 105 (1834). 

Cracca hispidula Kuntze, Rev. Gen. Pl. 174 (1891). 

Perennial from a slender, woody base, minutely appressed- 
hispid or glabrate. Stems 1-3 dm. or more long, decumbent or 
assurgent, dichotomously branching, straggling, angled; stipules 
2-5 mm. long, subulate, sometimes persisting; leaves 3-6 cm. 
long, linear-oblong in outline, short-petioled, (5-10 mm.) the 
upper ones often sessile; leaflets 7-13, elliptical, oval-oblong, or 
linear-oblong, 8 mm.—2 cm. long, 4-8 mm. wide, acutish, some- 
times obtuse, apiculate, reflexed, glabrous above, appressed hirsute 
with whitish hairs and the veins often turning purplish beneath ; 
peduncles terminal and axillary, ancipital, 5-1o cm. long; flowers 
solitary or geminate, scattered at the summit of the peduncle, 10- 
12 mm. long; bracts 2-5 mm. long, setaceous; calyx-teeth acute 
as long as the tube; vexillum minutely pubescent; legume about 
4 cm. long, 4-6 mm. wide, straightish or slightly falcate, minutely 
hispid ; seeds 8-12, ovoid or nearly orbicular, brownish. 

Low pine lands, sandy soil, stems ascending, rarely prostrate 
or trailing, leaves and peduncles erect, leaflets reflexed, smaller 
and more acute than the other species in this section. 

Virginia and North Carolina to Florida, westward to Louisiana. 

Original localities: Virginia, Carolina and Georgia. 

Type in Herb. Michaux. 


11. CrRaccA SMALLII. 


Cracca intermedia Small, Bull. Torr. Club, 21: 303 (1894), not 
Tephrosia intermedia Graham; Hook. FI. Brit. Ind. 2: 112 (1879). 

Perennial from a woody base, pubescent throughout and some- 
what viscid. Stems 4-6 dm. long, branched from the base, spread- 
ing, assurgent, flexuous, more or less angled; stipules subulate, 
caducous; leaves oblong-obovate in outline,6-12cm. long; petioles 
I-cm. long; leaflets 3-11, oblong-obovate, I-3 cm. long, 6-14 mm. 
wide, glabrous and yellowish-green above, strigose with whitish 


34 


hairs and becoming somewhat purplish beneath, truncate at the 
apex, apiculate; peduncles 8 cm., 1-5 dm. long; bracts subulate, 5-7 
mm. long; flowers I cm. long, mostly solitary or geminate, remote; 
calyx-teeth lanceolate, acuminate; vexillum minutely pubescent; 
legume 3.5-4 cm. long, about 4 mm. wide, straightish, strigillose; 


seeds 5-9, oblong or oblong-ovoid, compressed, smooth, variegated 
with black. 


Dry and poor “ blackjack thickets.” Differing from C. chyy- 
sophylla in its assurgent habit, the greater number and shape of 
the leaflets, the smaller flowers and larger seeds, as well as the 
character of the pubescence on the under surface of the leaflets. 

Florida, Chapman, Curtiss; Georgia, Boykin. 

Original locality: near Jacksonville, Florida (Curtiss). 

Type in Herb. Columbia College. 


12. CRACCA CHRYSOPHYLLA (Pursh) Kuntze. 
Tephrosia chrysophylla Pursh, F\, Am. Sept. 489 (1814). 
Galega prostrata Nutt. Gen. 2: 120 (1818). 

Cracca chrysophylla Kuntze, Rev. Gen. Pl. 174 (1891). 


Perennial from a long ligneous root. Stems prostrate, 3 dm.- 
1 m. long, dichotomously branching, clothed with a close, short, 


somewhat viscid and spreading silvery or tawny pubescence; sti-_ 


pules subulate, caducous; leaves 3—5 cm. long, oval in outline, ses- 
sile or very short petioled ; leaflets 3-9, (rarely 1-foliolate,) 1-3 cm. 
long, 7 mm.-—2 cm. wide, cuneate-obovate, obtuse, often retuse, 
sometimes apiculate, coriaceous, glabrous and yellowish-green 
above, silky hirsute and somewhat lighter or rusty beneath, the ter- 
‘minal one often conspicuously larger than the lateral ones; pe- 
duncles 4-6 cm. long, axillary, ancipital; flowers 1—-1.5 cm. long, 
few; bracts 5 mm. long, subulate, persisting ; calyx-teeth acumin- 
ate,as long as the tube ; vexillum minutely pubescent ; legume 3-4 
cm. long, 5-7 mm. wide, minutely hispid, straightish, erect of 
spreading ; seeds 8—10, oblong, ovoid, or sub-orbicular; brownish or 
greenish, variegated with black. 


_ Pine lands. Truly prostrate, with prostrate leaves, widely 
spreading, forming broad mats. 

Georgia to Florida and westward. 

Original locality: In Georgia. 


CRACCA CHRYSOPHYLLA:‘CHAPMANNI n. var. 


A low prostrate slender plant with stems 2 dm. or more long; | 
aoe leaves 1-2 cm. long; leaflets 5-10 mm. long, oblong or obovate, 


35 


apiculate, glabrous above, appressed silky-hirsute beneath ; legume 
2 cm. long, 3 mm. wide, minutely hirsute, 4-7 seeded. 


St. Joseph’s, Florida, Chapman. 
Type in Herb. Columbia College. 


13. CRACCA FLORIDANA N. sp. 


Perennial from a short, somewhat creeping ligneous root. 
Stems prostrate, 2-6 dm. or more long, dichotomously branching, 
spreading, angled above, clothed with a short appressed or spreading 
often somewhat viscid pubescence ; stipules 3-5 mm. long, subulate, 
often persisting ; petioles 1-2.5 cm. long; leaves 4~10 cm. long, ob- 
long or rarely linear-oblong in outline; leaflets 9-13, oblong or 
oblong-obovate obtuse or truncate at the apex apiculate, 1.5—3 
cm. long, 5-12 mm. wide, glabrous and yellowish-green above, 
lighter, strigose with whitish hairs and the veins turning reddish 
beneath ; peduncles terminal and axillary, 7 cm._2 dm. long; bracts 
subulate; pedicels 5-8 mm. long; flowers solitary or geminate, 
I-1I.5 cm. long ; legume 3-4 cm. long, 4 mm. wide, erect, straight, 
strigillose ; seeds 6-10, oblong, ovoid, grayish or brownish varie- 
gated with black. ; 

Differing from C. chrysophylla in its more numerous, narrower 
leaflets and the pubescence of the lower surface, which is that of 
C. Smallit. \t is very close to the latter, from which it differs 
however, in its truly prostrate and spreading habit, narrower and 


more numerous leaflets. 

Central Florida, G. V. Nash, Nos. 494%, 1198, 1263, 1334, 
1552, 1615. Louisiana, New Orleans, Dr. Ingalls. 

Types in Herb. Columbia College. 


14. CRACCA CINEREA (L.) Morong. 
Galega cinerea L.. Amcen. Acad. 5: 403 (1759). 
Tephrosia cinerea Pers. Syn. 2: 528 (1807). 
Cracca cinerea Morong, Ann. N. Y. Acad. Sci. 7: 79 (1 892). 

_ Perennial from a stout, ligneous root and woody base. Stems 
prostrate, diffuse, or ascending, 3-6 dm. or more long, appressed 
cinerous-pubescent, or with somewhat spreading rusty” hairs 
above, becoming glabrate with age, angled and channelled above ; 
Stipules 3-8 mm. long, subulate, acuminate, persisting ; petioles 5 
mm. to 1.5 cm. long; leaves oblong in outline, 4-10 cm. long ; 
leaflets 16-17 oblong, linear-oblong, 2-5 cm. long, 4-8 mm. 
broad, the basal and terminal ones often obovate-oblong, obtuse 
or acutish at the apex, acute at the base, glabrous above, — 
_ 0us-strigose or pubescent beneath, becoming often glabrate wi 


36 


age; racemes. 7-8 cm. long; flowers 1 cm. long, geminate or in 
clusters, scattered; bracts subulate or setaceous, persisting ; calyx- 
teeth acuminate, as long as the tube; corolla purplish, vexillum 
pubescent; legume 3-4 cm. long, 4 mm. wide, spreading, ciner- 
eous-pubescent or glabrate, straight; seeds 6-9, ovoid, somewhat 
truncate at the ends, brown. 


Ballast ground, Mobile, Alabama (Ch. Mohr); Mexico and the 
West Indies, etc. 

Original locality: Jamaica. 

I am much indebted to Dr. N. L. Britton for his help and 
counsel in this study and for the use of the Herbarium of Colum- 
bia College. Mr. G. V. Nash has given me valuable assistance 
with copious field notes of the Florida species. 

Mr. Coville also very kindly loaned me the collection of the 
United States Department of Agriculture for examination. 


Contributions to American Bryology.—IX. 


ad sient S, BRITTON. 


A REVISION OF THE GENUS SCOULERIA WITH DESCRIP. 
TION OF ONE NEW SPECIES. 


(PLATE 227.) © 


The genus Scouleria was founded by Wm. Hooker in 1830, 
on specimens collected by Dr. Scouler at Observatory Inlet, de- 
scribed as S. aquatica, and subsequently distributed in Drum- 
mond’s Musci Americani as No. 63, collected in the Columbia 
and Portage Rivers. A few autograph duplicates of Dr. Scouler’s 
Specimens were also distributed in this country, and Dr. Torrey 
was fortunate i in possessing one of them, as well as a set of Drum- 
mond’ s Mosses. 

_ dn. 1851. GC. Mueller ' transferred Scomleria aquatica to Grimmia 
as G. Scouleri,and Lesquereux and James in the Manual 1884 
followed his example. Mitten in 1869 also subordinated the 
‘genus to Grimmia, describing one new species Grimmia patagonica 
(Journ. Linn. Soc. 12: 96, 1869), which Jaeger (Adumb. 1875) 

_ changed to Scouleria patagonica.. Since then the genus has been 


37 


maintained as valid: In 1889 Kindberg described in the BULLETIN 
S. aquatica var. nigrescens, which Mueller in 1890 raised to specific 
rank as S. Nevit. In Macoun’s Catalogue, 1892, another species 
was described by Kindberg S. MJuelleri, and in a recent number 
of Hedwigia Mueller described S. aguatica var catilliformis from 
Roll’s collections. This makes four species and one variety thus 
far described in the genus: 

Having had occasion to examine critically some specimens re- 
ceived from the Department of Agriculture, collected in the State 
of Washington by Leiberg and Sandberg, I found it necessary to 
see authentic specimens and original descriptions of all the species. 
These we have been fortunate in possessing either in the Torrey 
or Jaeger herbaria, where we found Lobb’s specimens of S. pata- 
" gonica, and the others have been sent us by Prof. Macoun, and by 

Dr. Watson in former years. 

As a result we have reached different conclusions from those 
of Miiller and Kindberg, and are of the opinion that S. aguatica is 
a very variable species, within certain narrow limits. That the 
forms which have been separated from it as S. Mevit, and S. Muel- 
/ert, intergrade with it is beyond question, and I have been able to 
prove that the characters which have been relied on to found spe- 
cific differences may all be found on one specimen, of either or 
any of the species distributed under the new names. There are 
some characters which seem to have been given too much weight, 
and others too little. I find that the tendency to differentiation of 
the cells bordering the leaves, runs through all the species, and 
reaches its maximum in the Patagonian specimens, by the forma- 
tion of a thick border just inside the margin, composed of paren- 
chyma cells on the upper surface of the leaf, with yellow prosen- 
chymatous cells on the lower surface like those composing the 
vein, giving the leaf the aspect of being triple-veined. In Wat- 
son’s specimens from Spokane Falls, which I have described as a 
new species, S. marginata (presumably those referred to in the 
Manual under G. Scouleri) I also find this character developed 
though in a less marked degree. The margins are bistromatic, or 

else the cells are larger and darker in color, though never as 
dense and dark, as in S. patagonica. — Specimens of S. aquatica 


var. nigrescens, also show an upward continuation of the basal 


38 


submarginal prosenchymatous cells, and this is more or less evi- 
dent in all the specimens, even Scouler’s of S. aguatica, though 
much less prominent in the young green leaves from the apex ot 
the stems than in the older leaves, where they become different 
jated in color, showing as yellow streaks, irregularly between the 
vein and margin, but always near the margin. 

Another peculiar character, which has been mentioned in S 
patagonica, but not in any other species, is the development of 
filamentous radicles on the vein at base. Mitten described them 
« Nervo obscuro inferne dorso radicellis vestito.’ This character 
is very prominent in some of our specimens, and in fact is hardly 
absent from any of them, the whole lower surface of the vein in 


some leaves being densely covered with scattered or tufted, seem , 


ingly glandular hairs. The serrations of the margins too, are very 
variable, and of no value for distinguishing the species. The 
young green leaves. are always more sharply and _ irregularly set- 
rate, generally also at the apex, but the older leaves on the same 
stems, are often entire at apex, and indistinctly serrate below; I 
have even seen leaves quite entire, on the same plants. The teeth 
are often black and swollen, though this is never a constant chat- 
acter. : 

The cucullate apex, for which Mueller has named the var. caéil- 
liformis, is due to the bending of the vein a short distance below 
its apex. This too seems to be a character of the younger leaves. 
They also vary in being serrate on the back near the apex, and the 
_ vein is sometimes much thickened, and prolonged to the summit of 
the leaf, as figured by Schwaegrichen, instead of ending below the 
apex as it usually does. The color and size of the 3-4 rows of 
marginal cells of the leaves also varies, and in some leaves the 
green quadrate cells of the margins are so sharply differentiated 
from the elongated narrower submarginal ones, that they form am 
undulate border next to a deep yellow fold on each side. 
ScouULERIA AQUATICA Hook. Bot. Mise. 1: 33, ¢. 78 (1830). 

A portion of the type specimens collected by Scouler are in 
our possession, and the following description was drawn from 
them, giving measurements which Hooker did not give. 

Plants 5-6 cm. long, stems flexuose, sparingly branched ; lower 


. s 2 : leaves abraded, sede eaten lanceolate, 4 mm. lene by 1.5 mm. 


i 
t 
7 
. 7 
¥ 
t 
; 


39 


wide, apex cucullate, rounded, entire or serrate, vein ending below 
it, not filamentous at base ; margins serrate only above the middle, 
not bordered, the cells only slightly larger and darker, upper cells 
irregular, 10-013 mm.; basal rectangular, rarely a few prosen- 
chymatous cells. were seen just inside the basal margin, mostly 
pale and rectanglar. Spores .o48-.050 mm., smooth, brown. 


On consulting the original description and plate, we find that 
Scouleria aquatica was originally described as black, and the type 
specimens are quite as dark as Macoun’s specimens of var. nigres- 
cens, thus invalidating the first and most conspicuous character of 
that variety. The leaves are described as dark green, the upper 
ones only as green. The border of the margin is indicated by 
submarginal lines in figures 2—3 of the original plate. 

The specimens distributed in our set and Prof. Macoun’s of 
Drummond’s Mosses No. 63 differ from Scouler’s and from each 
other slightly. Prof. Macoun has three plants, two are 12-13 cm. 
long, large, coarse, simple stems, and sterile, with the leaves long 
and broad, coarsely serrate and bordered with yellow, the vein 
strongly filamentous, the submarginal basal cells yellow and pro- 
senchymatous, and the lower margins undulate. The third plant 
in his set is a small branching, fertile one, with black abraded 
leaves, shorter and denser, often entire, with the margins yellow 
and thickened. Our specimens of Drummond's No. 63 are like 
these, the leaves being only 2-2.5 mm. long, and differing from 
Scouler’s in their blunt, flat, entire apex with the basal cells more 
distinctly prosenchymatous and yellow. They were cited in the 
original description of S. aguatica. 

It seems evident from the above that, as originally founded, 
this species was recognized as variable, for we have indicated three 
discrepancies in the original specimens and descriptions. The 
larger forms may be referred to the variety migrescens Kindb., 
which may be distinguished by the taller plants, darker and 
coarser than the type, with long simple stems, large black leaves, 
often blunt and entire at apex, the basal cells yellow or brown, 
prosenchymatous, with the vein often thickened and serrate at 


apex and filamentous at base. 


Scouterta Nevi Miiller, Bull. Torr. Bot. Club, 17: 273 (1890). 
The description given in Macoun’s catalogue for S. Nevit is 


more than half devoted to S. aguatica, and from it we gather that 


40 


the main difference is the width and ‘shape of the leaves, and the 
broad, rounded entire apex. We have seen all of Prof. Macoun’s 
specimens, and tried the following experiment; taking several 
stems from different ‘plants, we divided them into 5-6 sections 
each, and compared the leaves. In all cases we found that the 
upper green leaves at the tips of the branches were longer and 
narrower, more acuminate and more sharply serrate, with the apex 
also serrate and generally cucullate; the lower cells also were 
seldom differentiated, generally paler and oblong, not prosenchy- 
matous. The lower leaves on the same plants were shorter and 
broader, often entire and rounded at apex, the cells denser and 
darker, and often yellow with traces of prosenchymatous cells. 

We have not been able to find that any of the characters are 
constantly associated together, so that we cannot maintain S. Vev7 
as a species, even if the name had priority over the var. igrescens ; 
but there seems to be sufficient reason in maintaining the latter as 
a variety, as we have shown from the descriptions of Drummond's 
specimens and Macoun’s collections. 

We found one of Macoun’s specimens of “« S. Nevii” agreed 
with the specimens from Yale labelled var.-virescens (Bull. Torr. 
Bot. Club 16: 93, 1889), having the upper leaves of that brilliant 
emerald green color which is so striking in these specimens. Oa 
the younger and smaller plants the leaves are green. Large, 
coarse, old plants, with ragged leaves, are almost black. 


ScouLerta Mue.tert Kindb.; Macoun’s Catalogue, 6: 62 (1892). 
Macoun’s Canadian mosses, No. 558. 


The description calls for different specimens from those of No. 
558, which we have received from Prof. Macoun. Those sent us 
are undoubtedly referable to S. aquatica, with which they agree in 
every way. The poe reads the median basal cells “ linear, 
porose and aeroUs,, the apex entire rounded, and the margin 
“pale orange.” In our specimens the basal cells are rectangular, 
with a few very faint traces on some of the leaves of the yellow 
prosenchymatous cells referred to in the description. The apex 
is as often serrate as entire, and the marginal cells are green, in 


6-7 rows, and though larger and more distinct than the inner 
ones, are = not “pale orange.” 


41 


We have taken particular pains to see as many specimens of 
Macoun’s No. 588 of S. Muelleri,as possible, thinking that perhaps 
there might be a mixture of specimens, and perhaps our S. mar- 
gimata be found among the number. But Prof. Macoun assures 
us that the species is local, and all the specimens of this number 
were collected from the same place and grew on the same rock: 
This is very interesting, for the specimen in our set agrees with 
Scouler’s specimens of S. aguatica, the ones from Prof. Eaton’s set 
are referable to var. nigrescens, and these last of Macoun’s, he 
assures me, are. the very ones from which Kindberg named S. 
Muelleri. 


Key. 


Leaves bordered by slightly larger, thick-walled cells, in a single layer, green, 
yellow or black, peristome present, . ee SP EET Ny nee I. aquatica, 


Leaves bordered by larger, denser cells in a double layer, often prosenchymatous 
almost to apex, peristome absent,..........2.+--+- seeeesees 2. marginata, 


1. SCOULERIA AQUATICA Hook. 


Scouleria aquatica Hook. Bot. Misc. 1: 33, ¢ 28 (1830). 

Grimmia Scouleri Miill. Syn. Musc. Frond. 2: 654 (1851). 

Scouleria aquatica var. virescens Kindb. Bull. Torr. Club, 16: 93 
(1889). 

Scouleria Muetleri Kindb. Macoun’s Cat. part 6: 62 (1892). 

Scouleria aquatica var. catilliformis Mill. Hedwigia, 32: 207 


(1893). 

Plants dark green or black, gregarious, growing in tufts; stems 
rigid, simple or branching, 5-15 cm. long; leaves when old 
abraded, only the veins remaining, upper green, walls less thick- 
ened, more prominently serrate than the lower, which are often 
black, entire, bordered with yellow or black thick-walled cells; 
vein thick, ending below the apex, often arched and serrate on 
back above and filamentous below; apex rounded and flat or cu- 
cullate, serrate or entire; basal cells variable on the same mi 
on the upper leaves pale often entirely rectangular, on a soe 
Often yellow or brown, with streaks of prosenchymatous cells, ae 
inside the Margin, extending upward irregularly, the 2 
rectangular, often undulate. Capsules almost immerse i ai 
short seta, oblate-spheroidal, becoming more depressed after . 
dehiscence of the lid, which remains attached to the oo > 
and exserted, long after maturity; calyptra Oa es = 
Single, red, teeth 16, irregularly divided and broken, often falling 


42 


with the lid; spores large, smooth, .037-.059 mm. maturing in 
May and June to August and September. 


A variable species, growing on rocks in mountain streams,’ 


more or less local, but abundant. 

Original locality: Observatory Inlet, Scouler, 1829. Distrib- 
uted from the Columbia and Portage Rivers in Drummond's North 
American Mosses, No. 63. Also collected by Lyall in the Columbia 
River, by E. Hall in Oregon, Bolander in California, Leiberg in 
Traille River and Lake Pend d’Oreille, Idaho, and by Macoun in 
several localities in British Columbia and Vancouver Island. 


Ta. SCOULERIA AQUATICA NIGRESCENS Kindb. Bull. Torr. Club, 16: 
~ 94 (1889). 
Scouleria Nevii Miiller, Bull. Torr. Bot. Club, 17: 273 (1890). 


_ Plants coarse and rigid, in large dense black tufts ; stems 10- 
-15 cm. long, brittle, and clothed with the persistent veins of the 


leaves, branching above; leaves 2-3 mm., oblong lanceolate, blunt: 


and entire at the rounded apex, vein ending below it, often radicu- 
lose at base; margins serrate, basal cells rectangular next the vein, 
prosenchymatous near the margin, but extending upward only a 
short distance; some leaves simply hyaline at base with all cells 
rectangular. 

Original locality «On rocks in Nanaimo River, Vancouver 
Island. Also collected at Sicamous, B. C., in 1889, and Rogers 


Pass, Selkirk mountains, B. C., in 1890 and 1885. Distributed as 


No. 388 of Macoun’s Canadian mosses. 
These four specimens, presumably named by Kindberg, illus- 
trate the variation of the species, two being large, coarse plants, 
with larger leaves and the basal cells prosenchymatous, the other 


two small branching plants, with short leaves, and the basal cells 
scarcely prosenchymatous. 


2. SCOULERIA MARGINATA n. sp. Plate 227. 


_ Plants 3-4 cm. high, gregarious in dense black tufts; stems 
_Wiry and naked at base, branching and densely leafy above; 

leaves crowded, curled and twisted when dry, only the uppermost 
green, 2-3 mm. long, oblong lingulate, serrate above the middle, 
or obscurely serrulate near the base, teeth occasionally black and 
thickened ; apex blunt, entire or toothed, vein thick, ending below 
it, smooth on back; basal cells green rectangular, a narrow band, 
__ near the margin elongated, prosenchymatous, forming a dark dense 
__ border nearly to the apex of the leaf, saperposed by rounded small 


43 


cells; perichaetial leaves surrounding the capsules, ovate-lanceolate; 
capsules, small, broader than long, cupuliform when old; lid persist- 
ent on the columella, bordered with red; peristome none, mouth 
bordered ; spores .048-.054 mm., green with a minutely roughened 
coat, maturing in August and September. 


“Spokane Falls, Washington Territory, collected by Sereno 
Watson, September 24, 1870.” Presumably the same specimens 
are referred to in the manual under G. Scouleri, Miiller, as being 
abundant. Distributed with the plants collected on Clarence 
King’s Expedition on the Exploration of the 4oth Parallel. 

Since collected by Marshall A. Howe on rocks just above the 
water in the Sacramento River, Sims, Shasta county, Cal., August 
IO, 1894. 

Closely allied to S. Patagonica, but the marginal cells are less 
dense, often only one layer of cells, but larger and square in 
section. 


Description of Plate 227. 


Fig. 1. Plants natural size, 2. Capsule enlarged, lid on. 3. Capsule after de- 
hiscence of lid. 4-5. Outlines of leaves. 6. Basal cells of leaf. 7. Apex of leaf. 
8. Cells from the middle of the leaf, showing the elongated, submarginal cells. 9. 
Cross-sections of leaf, showing the thickened margins. 10. Cells from the upper sur- 
face of the leaf. 11. Spores. 


Studies in the Botany of the Southeastern United States,—III. 
By Joun K, SMALL, 


(PLATE 228.) 


TsvGa Carorintana Engelm. Coult. Bot. Gaz. 6: 223 (1881). 
This tree can now be added to the flora of Georgia. Formerly 
it was known only from the Carolinas and Virginia. There it 
grew at altitudes ranging from 2,100—5,000 feet. I found it in 
1893 growing on the southern ledges of the cafion at Tallulah 
Falls, thus extending its range many miles to the south and its 
altitude to 1,600 feet. It was most plentiful about 300-400 feet 
above the river and reached no great development on account of 
the scarcity of soil and the perpendicular position of its place of 
growth. As was the case in all the other localities where I saw 


: . 1 


44 


this species, 7suga Canadensis was present and rather the more 
plentiful of the two, thus serving to show the great contrast 
between these two hemlocks. 

Cyperus souarrRosus L. 


This most beautiful little species of Cyperus has been found 
for the firsttime on the North American continent at Jacksonville, 
Florida, by Mr. A. H. Curtiss. The Florida Specimens agree 
exactly with Wright’s Cuba collection, No. 3355 (distributed as 
C. aristatus), also with plants from the Antilles, Surinam (Schwein- 
itz), and Bailies’ Niger Expedition of 1857-9 as well as with East 
Indian specimens. 
Juxcus GrorGianus Coville.* 


RUMEX SPIRALIS n. sp. 


- Perennial, slender, glabrous, light-green, somewhat glauces 
cent. Rootstock woody, creeping, 1-2 dm. long; roots fibrous; 
stem erect, 8-9 dm. long, simple or sparingly branched above, 


oh ee acai 


~- * Juncus Georgianus Coville n. sp. 


: Perennial, densely tufted, 20 to 35 cm. high; stems erect, barely exceeding I mm. 
in diameter, striate when dry; leaves all radical; sheaths striate, stramineous, loose, 
minutely auriculate, commonly 2 to 4 cm, long, the innermost closely embracing the 
stem and sometimes reaching a length of 8 cm.; blades erect, some of them reaching 
at least the base of the inflorescence, transversely flattened, nodeless, striate on the 
back, I mm. or less in width, sometimes involute when dry, sharply acute at apex; 
inflorescence paniculate, about 5 to 10 cm. high, strict or only slightly spreading 5 
lowest involucral leaf foliose, not exceeding the panicle ; flowers rarely more than 25, 
usually not more than 10, inserted singly on the branches of the panicle, prophyllate; 
perianth 4 to 6 mm. long, its parts subulate-Janceolate, when young with a green mid- 
rib, usually reddish brown lateral stripes, and hyaline margins, when old stramineous ; 
stamens 6, one-half to two-thirds the length of the perianth, the anthers 1.5 to 2 mm. 
in Jength and several times longer than the filaments ; style and stigma long, the 
former often reaching 2, the latter 3, mm. in lengta; capsule about three-fourths a5 _ 
long as the perianth, narrowly oblong-lanceolate in outline, obtuse or broadly acute, 
mucronate, 3 celled ; seed about 0.4 to 0.5 mm. in length, oblong, reticulate, the areola 
linear and arranged transversely on the seed in about sixteen longitudinal rows. 
Type specimen in the U. S. National Herbarium, collected in May, 1869, 0D 
Stone Mountain, Georgia, by William M. Canby. 
a This plant is most nearly related to Funcus tenuis, but is easily distinguishable 
eo! from that species by its long radical leaves, its longer and brown-striped perianth, 
__ Rarrower capsules, and especially by its very long anthers. Contrasted with Ysscws 
_ tenuis, the long radical leaves, short stems, and large inflorescence of ¥. Georgianus 
give the plant a characteristic general appearance. Mr. Canby’s specimens were dis- 
_ tributed doubtfully identitied as ¥. semwis, and no botanist seems to have collected 
__ the plant since, until Mr, Small rediscovered it July 4, 1893, on the summit of Stone 
Mountain, at the altitude of 1686 feet. He collected it also on Little Stone Mountain, 
July 7, 1893, between 1,000 and 1,100 feet altitude. — : pans 
hae ey - FREDERIC VERNON COVILLE. 


> 


45 


leafy throughout, slightly flexuous, strongly channeled, woody 
below; leaves lanceolate or oblong-lanceolate, 6-13 cm. long, 
1.5—4.5 cm broad, acute or sometimes attenuate at the apex, the 
lower ones obtuse or truncate at the base, the upper acute or 
acuminate at the base, all rather long petioled, coriaccous, light 
green, undulate and crisped, neither prominently nor conspicu- 
ously nerved; petioles strict, 2-5 cm. long; orceae cylindric 
nearly one half as long as the internodes; inflorescence terminal, 
simply paniculate, naked; racemes (fruiting) 5-12 cm. long, 
dense, rather erect, the terminal one- usually about twice as long as 
the lateral ones ; calyx 2 mm. broad ; pedicels varying from 2-4 
mm. in length, jointed below the middle; wings broadly ovate 
cordate, broader than high, 1 cm. long, I-1.2 cm. broad, straw- 
colored, sometimes slightly constricted below the apex, conspicu- 
ously and prominently nerved, crenulate and undulate, each one 
bearing an oblong-ovoid callosity, the three wings strongly spirally 
twisted ; achene broadly oblong-ovoid, 3 mm. long, short-pointed, 
chestnut colored, its faces nearly flat, its angles conspicuously 
margined. Plate 228. 
Found growing in the mud on the margins of ponds near 
Kenedy, Carnes county, Texas, by Mr. A. A. Heller, collected in 
flower and fruit on May 26, 1894. The altitude of the station is 
about 400 feet. Ao. IDS! | 
Its nearest relative is Rumex altissimus, from which, however, 
it differs in having more characteristically lanceolate leaves, which 
are longer-petioled, crisped and the larger ones more or less trun - 
cate at the base instead of acuminate. The panicle of &. spiralis 
is more open, not leafy, and its racemes are denser and thicker. 
Wings twice to thrice as large as in X. altissimus invest the broadly 
oblong-ovoid achene. The former are broader than high and 
strikingly cordate, whereas those of &. a/tissimus are higher than 
broad, not strongly cordate and less prominently nerved. So far 
as observed three callosities are developed throughout. 


Baptista SereNAE M.A. Curtis, Amer. Journ. Sci. (I.) 7: 406 
(1845). ete 
The range of this species, heretofore confined to the upianre 
and foot-hills in South Carolina and Georgia, has now been ex: 
tended into the low country by its discovery by Miss samt: 
A. Taylor in the pine barrens about Summerville, South: Carolina. 


OxaLis rEcurva Ell. Bot. S. C. & Ga. 1: 526 (1821). 2 
- Since writing my paper on the above species, which was 


46 


published in the November Butters, the plant has been found 
by Prof. A. Ruth and Mr. T. H. Kearney, Jr., at Wolf Creek, 
Eastern Tennessee, and by myself at the Falls of the Yadkin Rive’ 
in Stanley county, middle North Carolina, where it grows in the 
sand in shady places at the bottom of the cafon. The lathes 
locality is within the range as formerly known, but the former 15 4 
little west of the range shown in the above cited paper. 
Hypericum Buckteyi M. A. Curtis. Amer. Journ. Sci. 44: 80 
(1843). 
When on the summit of the Thomas Bald, on the Georgia and 
North Carolina boundary, in 1893, I encountered a peculiar Hyper 


icum. This summit is nearly 5,300 feet above the sea level and 


is remarkable for its shape, which is almost knife-like, being but 
a few yards broad and three miles long by actual measurement. 
The top, which is composed of soil and outcropping gneiss, is free 
from timber, except a few scattered red oaks. On the gneiss out- 
crops this Hypericum forms dense cushions and mats, rising above 
the ground only two inches. Altitude has had a striking effect om 
the species there and at first sight one is not inclined to refer it to 
the above. The locality is in view of the original and later stations 
for H. Buckleyi and a comparison with all the material at hand 
shows these differences. Its leaves are at least one-half smaller 
than the usual and rather constant form. The seeds are one-third 
smaller and more curved, while the flower and all its parts, together 
with the capsule,are also one-third smaller than those from the 
neighboring territory. 

MONNIERA CRENULATA N. Sp. 

Perennial, bright green, very aromatic. Stem procumbent of 
decumbent, creeping, ascending at the ends, 3-6 dm. long, 
branched from the creeping nodes, more or less channeled, pilose 
with rather rigid, irregularly jointed hairs; leaves broadly of 
orbicular-ovate, 1.5-2.5 cm. long, 1.5~2.2 cm. broad, subcordate 
and amplexicaul, obtuse or slightly emarginate at the apex, re- 
_motely but distinctly crenulate, mostly eight-nerved, obscurely 
_ pilose-ciliate near the base and on the midrib beneath, glandular- 


punctate, exceeding the internodes, except on the lower part of 


the stem; pedicels I-1.5 cm. long, pilose; bractlets 1 cm. long, 
_ very similar to the leaves in shape, texture, etc., but eciliate ex- 


cept a tuft of hair at the apex; calyx segments lanceolate, ciliate, 
- nearly equalling the bractlets; corolla campanulate, 1-1.3 cm. 


- eemrsttaecsieteaeteene OED 
SRI EN SM I ME 


eee ee 


&. 


47 


long, slightly unsymmetrical, cleft for about one-third of its length ; 
style about equalling the distal pair of stamens; capsule ovoid- 
oblong. 


Found by Mr. A. H. Curtiss, growing in the bottom of ditches 
between Jacksonville and Trout Creek, Florida, on July 13th 1893. 
- The above described species is related to Monniera amplexi- 
caulis of Florida, but is easily separated from it by its much larger 
size, its broader leaves, the elongated pedicel and the larger flowers. 
In some respects it is closely related to J. lanigera of the tropics, 
but in the latter species the nerving of the leaves is pinnate and 
the intenodes longer than the leaves, while in JZ. crenulata the 
leaves are flabellinerved and longer than the intenodes. 


-LYCOPERSICUM ESCULENTUM Mill. 


Is spontaneous about the village of Stone Mountain, Georgia, 
and also at points along the Georgia Railroad. 


COREOPSIS LONGIFOLIA n. sp. 


Annual or perennial (?) from an enlarged and somewhat woody 
base, slender, glabrous, bright green. Stem erect, 7-10 dm. 
long, simple or sparingly branched at the summit, not angled but 
channeled by twelve or thirteen grooves, leafy on the lower half, 
naked above, ‘slightly flexuous; leaves linear-oblong, linear- 
lanceolate or linear, 7-10 cm. long, .2-1 cm. broad, acutish, 
acuminate at the base, long-petioled, reduced above to narrow and 
inconspicuous bracts; petioles 6-8 cm. long, very narrowly 
winged, enlarged at the base, in most cases forming a short sheath 
which clasps the stem; heads 1-6, about one hundred flowered, 
3-4 cm. broad; rays yellow, eight, 1.5-2 cm. long; spatulate 
or oblanceolate, three-cleft, the segments acutish or the middle one 
obtuse ; the outer involucral scales lanceolate, marked with a dark 
rib, the inner scales twice as long as the outer, oblong-elliptic, 
thin, acutish, tipped with brown; floral scales, linear, 6 cm. nt 
acutish ; flowers, 4 cm. long, style slightly exserted, two-c oh 
achene obovoid or spatulate in outline, black, the soe mostly 
fugacious, the wings pectinate, the segments equal or nearly so. 


Related to C. angustifolia, from which species it differs in its 
more slender build, the striking length of the leaves and the 
larger flowers. The achene is also larger and of a darker sar 
In C. angustifolia to the wings of the achene are cut in an eater 
manner and the awns persistent, while the wings in C. longifo a 
are regularly and evenly divided and the arms mostly oe ! 

The type was found in dry, grassy pine woods about Jackso 
ville, Florida, by Mr. A. H. Curtiss on October 16, 1893. 


48 


V COREOPSIS MAJOR LINEARIS N. var. 


Perennial by a slender creeping root-stock, slender, sparingly 
and inconspicuously pubescent. _ Stem erect, 3-5 dm. long, simple, 
conspicuously channeled ; leaves three-parted, the segments linear, 
3-11 cm. long, 1-3 mm. broad, acuminate at both ends ; heads 
solitary, 3-4-cm. broad ; outer involucral bracts 4 mm. long, oblong, 


obtuse; rays oblong obtuse, somewhat two-cleft; achene smaller 
than in the typical form. 


Although reluctant to describe varieties the above is so distinct 
from Coreopsis major that it would be hardly fair to pass it. It is 
much more distinct than the variety Oe/meri. The strongest 
characters are’ the very slender build and the strikingly narrow 
leaf-segments which average about 2 mm. in breadth. Their 
length is also greater than we find the ordinary forms of the type 

I noticed the plant at different localities in middle Georg}, 


and collected it on Little Stone Mountain, at the base of Stone — 


Mountain, and also in the Yellow River Valley, in Gwinnette 
county. ; 


Coreopsis Major Walt. Fl. Cor. 214 (1788). 

_ A peculiar state of this species has been coming to my notice 
for a year or two. It is the above with its leaves undivided, there 
being two opposite and entire leaves at each node in place of the 
normal three-parted ones. It seems to have been first collected 
by Dr. and Mrs. Britton, at Black Mountain Station, North Caro- 
lina. Later Mr. Heller secured it near Salisbury, N. C., and last 
season (1893) I came upon it at the western base of Stone 
Mountain, Georgia. Besides having this state we have an inter- 
mediate one in which the upper leaves are entire and the lowet 
ones three-parted in the usual manner. This latter was secured 
by Dr. and Mrs. Britton at Balcony Falls, Virginia (1885). 


New Plants from Idaho, 
By Louis F. HENDERSON. 


-PHACELIA. IDAHOENSIS, n. sp. 


_ A foot and a half to two feet high, from nearly glabrous at 
bottom to slightly villous-hirsute in the inflorescence, leafy to the 
top ; radical and lower cauline leaves about 5 inches long, on peti- 


’ 


< 


49 
oles from an inch and a half to two inches in length, pinnately 
parted or divided into broad or comparitively narrow inch-long, 
cleft divisions, dark above, light beneath, and delicately strigose ; 
middle cauline leaves short-petioled, upper sessile, all pinnately 
cleft or parted; short spikes crowded during anthesis in a naked 
spike-like thyrsus about 3 inches long by 34 inch thick, in fruit be- 
coming more elongated and open; flower buds violet blue, becom- 
white-blue on opening; flowers open-campanulate, cleft barely to 
the middle, and bearing the vertical appendages of P. sericea ; an-* 
thers oval; styles 2-cleft at apex, these with the stamens hardly 
longer than the corolla; capsule ovate, short-a€uminate, con- 
tained within the marcescent-persistent ,coxolla, 12+22-seeded; 
seeds oblong-oval, generally irregularly and strongly angled by 
pressure, acute at one end, less so at the other, longitudinally and 
rather deeply alveolate, the walls’ separating the alveolations thin 
and sharp. 

Common in moist, natural meadows of Craig Mountains, Nez 
Perces county, at about 3000 ft. alt. A single specimen in fruit 
was found on the St. Marie’s River, Kootenai county, proving 
that the plant must be well distributed in Northern Idaho. That 
the species is closely related to P. sericea A. Gray, is evident on close 
inspection. In aspect, however, it is very different, and the rela- 
tionship would hardly be suspected. It differs from this species in 
being nearly glabrous, erect, and 2-3 times as high; in its stamens 
and style being never more than half again as long as the corolla; 
in its thyrsus being much more slender and rather longer; finally 
in the deeper alveolations of its strongly angled seeds. It differs 
from the var. Lya//i in its taller stature, in its narrower and longer 
thyrsus, and, if the flower and seed characters are those of the 
type, in these also. As the writer has no good specimen of the 


variety, on the last points he is doubtful. 


CLAYTONIA ARENICOLA N. sp. 

Annual with delicate, fibrous roots, 2-6 inches high: radical 
leaves linear-spatulate, the broadest not over 2% lines wide (gen- 
erally about a line wide), I-2 inches long, tapering from near the 
obtuse apex into a delicate petiole ; cauline leaves a single pair, 
similar to the radical but shorter, opposite and distinct: racemes 
numerous and prolifically flowered, the flowers on pedicels, Yy-% 
inch long; petals pink-white, 3 lines long, emarginate ; seeds % 
line long, shining and resembling those of C. Siéirica but only 
half.as large. 4 


Dry, sandy banks along streams as well as dry pine woods, 


Ma Rat Carden 


50 


Idaho and Eastern Washington. This plant has been referred for 

me to C. spathulata var. tenuifolia Gray. I am convinced that 
this ought not to go into C. spathulata Dougl., for throughout this 
whole country the cauline leaves are never united but spatulate- 
linear. The flowers are also much larger and in much loosef } 
racemes than in this species. Prof. E. L. Greene has sent mea — 
species very near this, only differing in the shape of the cauline 
leaves, the leaves in his specimen being linear and slightly em 
larged at the base, while in this species they are invariably spai- — 
late-linear. 1 should not think this enough to found a species — 
upon and separate it. from his species, which he names C. Sypso 
philoides Fisch. & Meyer, were it not that Dr. Gray says (ProG 
Amer. Acad. 22: 282) that C. gypsophiloides F. & M. is the — 
same as the ¢ype of the species, viz.: C. spathulata of Douglas. — 
Relying upon this, I give this plant of mine the specific name of 
arenicola. If it proves that Dr. Gray is wrong and Prof. Greene — 
tight, this name of mine would probably sink into a synonym 
of C. gypsophiloides. 


UNIVERSITY OF IDAHO, 


Buxbaumia aphylla L, 


Buxbaumia aphylla is generally considered a rare moss, and a 
from its large and peculiar capsule it could not escape notice if it _ 
were at all common. My friends Messrs. Chas. E. and Edwin 
Faxon inform me that usually single plants, or at most two OF — 
three together, have rewarded their patient search. But this De : 
cember it has been very abundant, particularly in the Blue Hill 
region, and in one locality of less than 200 feet square I counted 
the following patches of it: Nine of about one inch square con 
taining from ten to fifteen plants each; one of three by two inches 
of seventy plants; one of two by four inches of eighty-one plants; — 
three patches each about as large as my hand crowded with plants, 
of which one contained two hundred and eighty specimens. I was 
reminded of the pictures of a Roman legion under its testudo 
shields marching to attack a walled town. The locality where 
‘these are growing was burned over in a wood fire eighteen months | 


51 


ago, and the soil is yet charred from its effects, and nine months 
later what underbrush had started was pretty effectually cut off by 
the Park Commissioners, so that the conditions under which these 
plants grew were different from any they would: have found there 
in the past fifteen years that I have known the place. I would be 
very glad to know if anyone has ever found this moss in such 
abundance. . Gro. G. KEnNepy. 


READVILLE, Mass. 


Herbert A. Young. 


The news of the death, at Toledo, Ohio, December 8, of Herbert 
A. Young, formerly of Revere, Mass., will be received with regret 
by his many friends in. this vicinity.. He early in life became 
interested in botany, and in 1882 published the “Flora of Oak 
Island,” a botanical station in the vicinity of Boston, familiar to 
botanists since the days of Jacob Bigelow. He later became in- 
terested in the sedges, grasses and mosses, and contributed largely © 
to these sections of the Flora of Middlesex County, Mass. 

He was a good scholar and a keen botanist, but in recent 
years the demands of his profession as a civil engineer, and later 
as an officer of the Mexican Central Railway, have prevented his 
giving much attention to his favorite study. He passed away at 
the early age of thirty-seven, but he had already accomplished a 
work that entitles him. to the esteem and remembrance of the 
botanists of Boston and vicinity. Ws. P. Ricu. 

Boston, December 26, 1894. 


Proceedings of the Club. 


TursDAY EvENING, DECEMBER IITH, 1894. 


The regular meeting of the Club was held in the lecture room 
of the new building of the College of Pharmacy, 115 West 68th 
street. The evening was very stormy. Vice-President Allen 
occupied the chair and there were forty-eight persons present. 

The Committee on Membership reported favorably upon the 


52 


nomination of Mis Harriet B. Elder, of 515 Lexington avenue, 
City, who was unanimously elected an ‘active member. 

Mr. Lighthipe reported the transfer of the Club’s herbarjum 
to its mew quarters in the College of Pharmacy building. 

The paper of the evening was then presented by Dr. Rusby, 
on “ Pharmaceutical Botany.” 


Index to recent Literature relating to American Botany, 


Allen, T. F. Note on Chara sejuncta A. Br. Bull. Torr. Bot. Club, 
21: 526. 24 D. 1894. 


Anderson, C. L. Some new and old Algae but recently recognized 
* on the California coast. Zoé, 4: 358-362. 7 fig. 1894. 
New species in Punctaria and Callithamnion, — 
Arechavaleta, J. Contribucion al Conocimiento de los Liquenes 
Uruguayos. Ann. Mus. Nac. Montevideo, 2: 173-186. 1594. 


Arechavaleta, J. Las gramineas Uruguayas (continuatine). Ann. 
Mus. Nac. Montevideo, 2: 93-171. figs. 1894. 


Armendariz, E. Apuntes acerca de una Contrahierba de Mexico. 
‘La Naturaleza (II.) 2: 380-382. 1894. 


Atkinson, G. F. Intelligence manifested by the Swarm-spores of 


Rhizophidium globosum (A. Br.) Schroeter. Bot. Gaz. 19: 503, 504: 
26 D. 1894. 


Atkinson, G. F. Microsphaera densissima (Schwein.) Peck. Bull. 
Torr. Bot. Club, 21: 528, 529. 24D. 1894). 


Bain, S. M. Notes on U:ricularia inflata Walt. Asa Gray Bull. 
32S s 4; Sv- Je. rhage. é 


Barnhart, J. H. Udmaria Uimaria(L.). Bull. Torr. Bot. Club, 21: 
49t. 23 N. 1894. 


Beal, W. J. The Sugar Maples of Central Michigan. Ann. Rep. | | 
Sec. State Board Agric. Mich. 33: [Reprint, pp. 8. figs. 18¢4.] 
Best, G. N. Orthotrichum gymnostomum Bruch. Bull. Torr. Club, 

Si: 527,528. 24 D. 189%. 
Records this species from Newfoundland, 

Blochman,I.M. Californian Herb lore.—V, Erythea,2: 162, 163- 
1 O. 1894. : 
Medicinal character of various plants. 


53 


Botanical Congress, Madison. Proceedings, 1893. 8vo. pp. 60. 
Madison, Wisconsin. Je. 1894. 

Brandegee, T. S. Additions to the Flora of the Cape Region of 
Baja California—II. Zoé, 4: 398-408. pl. 77. 12. Mr. 1894. 


' Faxonia, new genus of Compositae. New species and varieties in 7halictrum, 
Ranunculus, Colubrina, Carica, Forestiera and Adelia. 


Brandegee, T.S. Two undescribed Plants from the Coast Range. 
Zoé, 4: 397% 380. pl. 70. 12. Mr. 1894. 


Lastwoodia elegans, new genus and species of the Compositae, and Lepidium 
Faredi, 
Bray, W.L. See Uline, E. B. 
Clarke, C.B. On certain authentic Cyperaceae of Linnaeus. Journ. 
Linn. Soc. 30: 299-315. 6 O. 1894. 


References to many American species, 
Coville, F. V. The Wild Rice of Minnesota. Bot. Gaz. 19: 504-509. 
26 D. 1894. 
Davenport, G. E. Two new Ferns from New England, with some 
observations on Hybridity and Nomenclature. Bot. Gaz. 19: 492- 


497. 26D. 1894. 


Describes Aspidium cristatum <A. marginale and A. simulatum. 

Davy, J.B. Transcripts of some Descriptions of California Genera and 
Species.—II. III. IV. Erythea, 2: 148-153. 1 S.; 164-170. 1 O.; 
185-187. 1 N. 1894. 

Deane, W. Lemna Valaiviana Philippi. Bull. Torr. Bot. Club, 21: 
490. 24N. 1894. 

Dudley, W.R. Phyllospadix, its Systematic Characters and Distribu- 
tion. Zoé, 4: 381-385. ‘26 F. 1894. 
Fernald, M. L. Notes from the Gray Herbarium. Zoé, 4: 379, 380. 

26 F. 1894. 


New species and varieties in Habenaria, Allium and Trifolium. 
Fernald, M. L. See Robinson, B. L. 
Fernow, B.E. The Battle of the Forests. Nature, 50: 116. 
Greene, E. L. Corrections in Nomenclature.—V. Erythea, 2: 192- 
194. 1D. 1894. 
Greene, E. L. Novitates occidentales.—VIII. IX. Erythea, 2: 181- 


385. 1. N.; 189-192. 1 D. 1894. ; a 
itched aie species in Ranunculus, Delphinium, Phacelia, Amsinchia and 


Trifolium, 
Greenman, J. M. See Robinson, B. L. 


1894. 


54 


Guppy, H. B. On the Habits of Lemna minor, L. gibba and L. 
polyrrhiza. Journ. Linn. Soc. 30: 323-330. 6 O. 1894. 


Heald, F. De F. Contribution to the comparative Histology of Pul- 
vini and the resulting photeolitic Movements. Bot. Gaz. 19: 477-491: 
pl. 34. 26 D. 1894. 


Hill, E. J. Salsoli Kali Tragus. Bot. Gaz. 19: 506, 507. 26D. 
1894. : 
Notes early appearance of the Russian Thistle in [linois and Indiana. 

Hollick, A. Wing-like Appendages on the Petioles of Liriophyllum 
populoides Lesq. and Liriodendron alatum Newb., with Description of 


the Latter. Bull. Torr. Bot. Club, 21: 467-471. pl. 220, 222. 24 
N. 1894, 


Howe, M. A. Chapters in the early History of Hepaticology.—Il. 
Erythea, 2: 143-147. 15S. 1894. 

Jack, J. G. Notes on Trees and Shrubs. Gard. & For. 7: 294. 25 
Jy.; 306. 1 Au.; 315. 8 Au.; 326. 15 Au. 1894. 
Notes on various species of Vaccinium; Genista in Massachusetts; R2des. 

Jelliffe, S. E. Cryptogamic Notes from Long Island.—II.' Bull. 
Torr. Bot. Club, 21: 489. 24 .N. 1894. 
List of 13 Hepaticae. 

Jones, M. E. Contributions to Western Botany.—VI. Zoé, 4: .366- 
369. 1894. 
New species in Phlox and Astragalus. 


Jones, M. E. Systematic Botany. Zoé, 4: 374-379. 1894- 


Kearney, T.H., Jr. Some new Florida Plants. Bull. Torr. Bot. Club, 
21: 482-487. 24N. 1894. 


Describes Scutellaria integrifolia multiglandulosa, Trichostema suffrutescenss 
Pluchea foetida imbricata, Teucrium Nashii, Physalis arenicola, Aristolochia 
Nashii and Rhus Blodgettii, new species and varieties. 


Kerr, W. C. Survival of Storm-injured Leaves. Proc. Nat. Sci. Asst. 
Slt 52. 8 Dy 2894; 


Linn, A., and Simonton, J. S. Fissidens hyalinus in Pennsylvania. : 
Bull. Torr. Bot. Club, 21: 529. 24 D. 1894. 


Mocifio y Sesse. Flora Mexicana. La Naturaleza (II.) 2: App- 9- 
48. 1893; 49-88. 1894. 


Peck, C. H. Annual Report of the State Botanist of the State of New 
York. Rep. N. Y. State Mus. Nat. Hist. 47: [reprint pp. 48]. 1894- 
Peckholt, T. Brasilianische Nutz-und Heilpflanzen. Pharm. Rund. — 
12: 187-199. Au.; 240-242. O.; 285-287. D. 1894. 


55 


Pieters, A. J. The History of the Uredineae. Asa Gray Bull. 3: 8: 
8-10. Ja. 1895. 


Pound, R., F. E. Clements and others. Additions to the reported 
Flora of Nebraska made during 1893. Bot. Surv. Nebr. 3: 5-20. 
18 Je. 1894. 


List of 182 additions, including descriptions of 13 new species of Fungi. 
Robinson, B. L. Contributions from the Gray Herbarium of Harvard 

University, new series. —VI. Proc. Am. Acad. Arts and Sci. 29: 273- 

330. 23 My. 1894. 

Contains: I. The North American Alsineae ; Il. Descriptions of Mr. Pringle’s 
Mexican collections; I1I. Notes on the Genus Ga/insoga ; IV, Miscellaneous notes 
and new species. 


Robinson, B. L. and Greenman, J. M. Contributions from the 
Gray Herbarium of Harvard University, new series.—VII. Proc. 
Am. Acad. Arts and Sci. 29: 382-394. 29 Je. 1894, 

Further new and imperfectly known plants collected in Mexico by Mr. Pringle. 

Robinson, B. L. and Fernald, M. L. Contributions from the Gray 
Herbarium of Harvard University, new series.—VIII. Proc. Am. 
Acad. Arts and Sci. 30: 114-123. 27 Au. 1894. 
New plants collected by C. V. Hartman and C. E. Lloyd in northwestern Mexico. 

Rovirosa, J. N. Viaje 4 Teapa y 4 las Sierras que concurren 4 la 
Formacion de su Valle. La Naturaleza (II.) 2: 269-293. 1894. 
Contains an account of the flora of the region. 

Rusby, H. H. Two new Genera of Plants from Bolivia. Bull. Torr. 
Bot. Club, 21: 487-489. f/. 225, 226. 24N. 1894. ; 

Describes and figures Lophopappus (Compositae) with LZ. foliosus and Fluck- 
igeria (Gesneriaceae) with 7. Fritschii. 

Russell, H. L. The Fixation of free Nitrogen by Plants. Bot. Gaz. 
Ig: 284-293. 16 Jl. 1894. 

Saint-Lager, Dr. Traditional Interpretation of Linnaean Nomencla- 
ture. Erythea, 2: 194-201. 1 D. 1894. (Translation). 


Scribner, F. L. ° Lower California Grasses. Zoé, 4: 385-393- 26 


F, 1894. sag 
An enumeration of 69 grasses collected by Mr. T. S. Brandegee, in Lower al 


nia in 1893. Sporobolus expansus proposed as new. 


Scribner, F. L. Grasses of Tennessee. Part II. 


Exp. Sta. Bull. 7: 1-141. 787 figs. 1894. 
Descriptions and illustrations of all the grasses known to occur in the State. 


Univ. Tenn. Agric. 


56 


Small, J. K. Twospeciesof Oxadis. Bull. Torr. Bot. Club, 21: 471- 
475. pl. 222-223. 24N. 1894. 

Describes and figures O. recurva Ell. and: O. grandis n. sp. 

Small, J. K. Notes on some of the rarer Species df Polygonum. Bull. 
Torr. Bot. Club, 21: 476-482. p/. 22g. 24N. 1894. : 
Figures P. boreale, 

Stevens, F. Pistillody; staminody; teratelogy. Bull. Torr. Bot. 
Club, 21: 489, 490. 24N. 1894. 

Sturtevant, E. L. Notes on Maize (continued). Bull. Torr. Bot. 
Club, 21: 503-523. 24 D. 1894. 

Villada, M.M. La Goma Loca de Mexico—I. La Natuaraleza (II.) 
2: 383-385. pl. 78. 1894. 

Pictures of Larrea Mexicana Moric. and Acacia filicina Willd. 

Whitfield, R.P. On new Forms of Marine Algae from the Trenton 
Limestone, with Observations on Buthograptus laxus Hall. Bull, Am. 
Mus. Nat. Hist. 6: 351-358. A/. zz. 20 D. 1894. 


Callithamnopsis, Chaetocladus and Primicorailina are proposed new genera. 


SCOULERIA MARGINATA E. G. BRITTON. 


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Second=-Hand Books for Sale. 


The following botanical works, owned mainly by the late Mr. Justus F. Poggen- 


burg, are offered for sale by the Secretary of the Torrey Botanical Club at prices 
considerably below those ordinarily charged. Purchasers are requested to order by 
number, remitting the amount in advance to Ff.. H. RUSBY, 


96. 


97. 


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BULLETIN 
TORREY BOTANICAL CLUB. 


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A MONTHLY JOURNAL OF BOTANY. 


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AND OTHER MEMBERS OF THE CLUB. 


CONTENTS: 


PAGE, 
New Bpeciap of Ustilagineae and Uredineae: 
\ % B. Ellisand B. M. Everhart... . 

Contributions to American Bryology—IX. 1. 
__ The Systematic Position of Piyscomitrelia 
Patens; 2, On a Hybrid growing with 
Aphanorhegma serrata; 3.Ona eee 
pean Hybrid of Physcomitreila patens: 
Elisabeth G. Britton (Piates 229-231) _ 
Japanese Characeae—Il.: 7: F. Allen... 68 
Tradescantia Virginica var. villosa Wat- 


PAGE. 
ar ey ae Y, Heved .<: 2. o 
57 | Revrews.—The Life and writings of Rafin- — 
| esque; Annual Report of the Stafe Botamist — 
of New York; Additional Notes on the new — 
Fossil, Daimonhelix ; Through Glade and 
Mead; ‘The Characeae of America—Part2; _ 
List of Pteridophyta and Spermatophyta of : 
‘Northeastern North America... +. + 3 
Procespincs or THE CLUB. . . PEE 
: ‘Inpex. TO RECENT LrreRaTuRE RELATING © 
ro Ammrica® BOTANY ..-++.-+-- % 


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New Species of Ustilagineae and Uredineae. 


By J. B. ELtis AND B. M,. EVERHART. 


Ustiraco Wasuinctoniana E. & E. 
On leaves of some grass, State of Washington, spring of 1892 


(E. R. Lake). 

Sori linear, 1 mm.—2 cm. long, covered at first by the lead- 
colored epidermis, soon exposed and then mass of spores nearly 
black. Spores globose, 8-10 “ diam., olive-brown, minutely 
echinulate and filled with numerous small nuclei. The sori are 
sunk in the substance of the leaf, which is finally eaten away and 
perforated by them. 

Closely resembles Ustilago longissima (Sow.), but the spores are 


larger and minutely echinulate. 


Entytoma arnicauis E. & E. 
On leaves of Arnica cordifolia, Latah county, Idaho, July, 


1893 (C. V. Piper, No. 122). 

Spots amphigenous, deep rusty brown above, paler below, 
subangular, 2-4 mm. diam., with a pale yellow shaded border. 
Sori subepidermal. Spores spherical, hyaline at first, then pale 
brown, 10-12 » diam., with a smooth epispore 1%4-2 , thick. 
Conidia (Ramularia arnicalis E. & E. Proc. Acad. Nat. Sci. Phil. 
1891, 85) hypophyllous subcylindrical 15-20X2}4—-3 #, 2-3-nu- 
cleate, the upper end mostly a little curved, borne on subfascicu- 
late hyphae 12~-20X2 Yy-3 #. 


UrRomyces putcuerius E. & E. 
On leaves and petioles of Szlene ? sp. Lake Chelan, Wash. 


Aug. 1892 (Lake & Hall). 


58 


III. Sori scattered or circinate, mostly hypophyllous, small 
(7% mm.), naked, nearly black. Teleutospores obovate or ellipti- 
cal, becoming chestnut-brown, 22—28%18-—22 », epispore smooth, 
rather thick, moderately thickened at the rounded apex. Pedicels 
35-50 » long, stout, hyaline. There was no indication of any 
Aecidium on the specimen examined. 


UROMYCES CARICINA E. & E. 
On leaves or culms of Carex scoparia, Alcove, N. Y., July, 


1893 (C. L. Shear, No. 81). I. & III. 


_ Sori scattered, oblong, %-1 mm. long by about % mm. wide, 
covered by yellowish epidermis, not confluent. Uredospores 
sparingly mixed with the teleutospores in the same sori, ovate or 
elliptical, pale, tuberculo-echinulate, 20-22X14-16. Teleuto- 
spores obovate, pale below, rounded at the apex, or often with a 
distinct hyaline papilla, epispore smooth, strongly thickened and 
dark colored at the apex, 19-23+14-16. Pedicels yellowish, 
about as long as the spores. 

Uromyces Caricis Pk. has naked cinnamon-colored sori and 
larger spores. 


“ 


PUCCINIA TRIFOLIATA E, & E. 

On Jiarella trifoliata Seattle, Wash. June 1892 (Prof. C.V. Piper)- 

I. Aecidium on the slightly swollen stems and _ petioles. 
Erumpent, closed at first, soon open and cup-shaped, about 
mm. diam., thin, margin toothed. Spores ovate or subglobose, 
15-20 » diam., yellowish, epispore slightly echinulate, ampht- 
genous. 

Il. Uredo sori small, pale cinnamon color, y%-Y% mm. diam., 
naked above. Uredospores obovate or globose, 15—22 / in the 
longer diam., rather closely echinulate, hyaline, becoming pale 
brown. 

III. Teleutospores in larger (1 mm.) nearly black naked 
sori, elliptical, 22~-3515—20 », pale chestnut brown, rounded at 
each end, contents granular, scarcely constricted at the septum, 
epispore scarcely thickened at the apex, and when examined dry, 
under a high power, covered with a network of raised lines. 
Pedicels very short, almost wanting. : 

Differs from the other related species on Ziare//a and Sasifrag@ 
in the presence of Uredo and also in other particulars. 


-PUCCINIA SUBSTERILIS E. & E. ENA. F. 3141.] c. 
On Chrysopogon sp. Fort Collins, Colo., March, 1894 (C. F- 
Baker, No. 219). : 


Mostly hypophyllous. Sori (II. and III.) superficial, pulvinate 
elliptical, black-brown, 4-1 mm. long, naked. Uredospores 


59 


echinulate, globose or elliptical, 20-3018-22 “, brownish-black, 
epispore nearly equally thickened throughout ; pedicels slender 
hyaline, subpersistent, 20—30 long. Teleutospores (in the same 

_ Sori as the uredospores), oblong or clavate, pale, constricted at 
the septum, 22~30X12-15 «, epispore smooth, mostly not at all or 
only slightly thickened at the rounded or subtruncate apex. 


The uredospores are abundant and well developed, while the 
teleutospores are few in number and apparently not well matured. 

This is quite distinct from P. omnivora E. & E. and from P. 
Chrysopogt Barcl. 


Puccinia omnivora E. & E. [N. A. F. 3049.] 

On Chrysopogon nutans, leaves and stems, Newfield, N. J., 
autumn 1893. 

II. and III. Uredospore sori minute, narrow, 1-2 mm. long, 
at first covered, then rupturing the epidermis and discharging the 
yellow, globose or subelliptical, 18-22 “, subechinulate spores. 
III. Sori amphigenous, but mostly hypophyllous, oblong or 
% linear, 1 mm.—1 cm. long, \% mm. wide, at first covered, but soon 
bare and margined by the ruptured epidermis, nearly black. Te- 
leutospores elliptical to oblong, 22-40%12-16. K, scarcely con- 
stricted, mostly rounded at each end, but those in the center of 
the sori narrower and paler, and gradually attenuated into the 
Stout, 50-70X4-6 », yellowish-hyaline pedicel, upper cell darker, 
and in the shorter, elliptical spores mostly rounded at the apex 
without any distinct papilla, the narrower, paler spores with a dis- 
tinct yellowish-hyaline obtusely conical papilla, epispore smooth, 
distinctly thickened at the apex. 

Has the habit of P. graminis, but the spores are decidedly 


smaller, both the uredo and teleutospores. 


Puccinta macnoecia E. & E. : 
On leaves of Aster pulchellus, Mts. above Lake weet: deen: 


Aug. 1892 (Lake & Hall). 

III. Teleutospores slender-clavate, 40-50x14-16 Ms . 
almost hyaline below, epispore smooth, much thickened at t “5 ce 
and darker, slightly constricted, crowded in broad(2-3 me be he 
nearly black, hypophyllous sori, which are partly a <i cad 
epidermis. The upper surface of the leaf is marke as meh 
brown spots corresponding to the sori on the opposite side. as 
Spores are mostly regularly rounded at the apex, ot Cars a the 
_Obtusely pointed or even truncate. Pedicels about as long a 
spores. 

Differs from P. As¢eris in its large, dark sori. 


60 


PucciniA PHILIBERTIAE E. & E. 

(P. Gonolobi Rav. var. Philibertiae Pk. in M. E. Jones’ list of 
western plants, not described.) 

On leaves and follicles of Philbertia viridiflora ? Britton and 
Rusby, near Las Cruces, New Mexico, Oct., 1892 (E. O. Wooton, 


No. 43). : 
Amphigenous, but mostly hypophyllous. Sori hemispherical, 
dark chestnut brown, 13-34 mm. diam., superficial, mostly circin- 
ate around a compact central group of several confluent or con- 
nate sori; on the follicles the sori are densely crowded, covering 
the entire follicle, which is thus dwarfed and rendered abortive. 
Teleutospores elliptical or obovate, 20~-30y<15—20 #, scarcely con- 
stricted, rounded and obtuse at the apex, lower cell a little paler 
and often narrowed at the base, epispore smooth, scarcely thickened 
at the apex, contents of the cells granular. Pedicels slender, sub- 
hyaline, 55-65 “ long. Mesospores abundant, mostly smaller. 


Puccinia ZiZiaE E. & E. 
On leaves of Zizia cordata, Pullman, Wash., September, 1893 
(Prof. C. V. Piper, No. 164). 


III. Amphigenous but mostly epiphyllous. Sori small (4-M% 
mm.), not confluent, nearly black, soon naked; seated on smal 
(1-2 mm.), whitish, irregularly shaped spots, which are often con- 


fluent. Teleutospores obovate, elliptical or oblong-elliptical, “ 
22-30X15—20 #, only slightly constricted ; epispore smooth, rather 


thin or only slightly thickened at the rounded apex. Pedicels 
hyaline, about as long as the spores. 


Differs from P. dudlata, in its darker sori, smaller spores and, a5 & 


far as yet known, in the absence of any Aeccidium. 


PUCCINIA NIGROVELATA Ell. & Tracy. 
On Cyperus strigosus, Mississippi (Tracy). I] & III. 


Uredospores in short (1-2 mm.), oblong-elliptical sori, suf be 
rounded by the erect margin of the ruptured epidermis, ovate OF — 


elliptical, pale yellowish, aculeolate, 15-24 (mostly 15-20) 12-15 


u, Mass of spores cinnamon colored, | Teleutospores in flat sori, . 
I-3 mm. long, 34~—1 mm. broad, closely covered by the epidermis, 


which appears black by translucence, clavate-oblong, 35-55% 14-18 


“, broadly constricted, lower cell pale and narrowed to the pedicel, J 
upper cell broader and darker, epispore smooth, thickened at the 


apex, which is either rounded or obtusely pointed, with or without 


a hyaline papilla. Pedicels mostly shorter than the spores. The : 
teleutospore sori at length open by a longitudinal crack along 


the middle. 


61 


Puccinta Crapu EIl. & Tracy. 

On Cladium effusum, Ocean Springs, Miss., Aug., i889 (Prof. 
S.M. Tracy). II. & III. 

On the culms and peduncles of the cyme. -Sori small, ellip- 
tical, subconfluent, so as often to envelope and cover the peduncles 
for 2 or more cm. in extent; at first covered, then bérdered by the 
ruptured yellowish epidermis. Uredospores ovate or elliptical, 
22-25 X20 #, or subglobose, 18-20 #, at first hyaline, then deep red- 
brown (or ferruginous-yellow while lying in the sori) aculeate. 
Teleutospores in similar but much darker colored sori, clavate or 
oblong, 45-60X18—22 «, constricted, deep brown, lower cell nar- 
rower and paler, epispore smooth or slightly granular-roughened 
above, strongly thickened at the obtusely rounded apex, or with 
an oblique papilla, or sometimes subtruncate-flattened. Pedicels 
shorter than the spores, hyaline or slightly colored, stout. The 
uredospores are also pedicellate. 


PUccINIA GRANULISPORA Ell. & Galloway. 
On stems and leaves of Alum cernuum? Montana, 1890 


(Prof. F. D. Kelsey). ; 

II. and III. Sori linear, 4-1 cm. long, shorter on the leaves, 
¥% mm. wide, at first covered, then exposed by a longitudinal 
cleft in the epidermis, but only slightly prominent. Uredospores 
Subglobose, pale, faintly echinulate, 20-30X15~-20 #. pepe 
spores oblong, clavate or obovate, 45-60%20-25 /, smooth, wit 
granular contents, slightly constricted, upper cell subglobose re 
elliptical, darker, moderately thickened at the rounded or ae 
pointed or often truncate apex, lower cell paler, cuneate. Pedicels 
shorter than the spores, colored. Mesospores not abundant, 
shorter, obovate, 20-23 / long. 

Differs from P. Porri in its linear sori and larger teleutospores. 


AEcIpIum cytinpricum E. & E. 
On leaves of Houstonia angustifolia, Osborne, Kansas, June, 


Da hyll bseriate along each 
Spe ia? Aecidia hypophyllous, subseriate ¢ 

sae ithe AGEs white, atc about 1 mm. high, sini 3 
minutely subfimbriate-dentate. Spores orange-red, angular-g “di 
bose, smooth, 18-22 #. The upper side of the leaves is ange io 
less blackened and papillate from the projecting bases of the 
aecidia. . : = 

‘Differs from Accidium houstoniatum Schw. in its elongated, cy- 


lindrical aecidia. 


62 


Contributions to American Bryology—IX. 
By ELIZABETH G. BRITTON. 
(PLATES 229-231.) 
I. THe SysTEMATIC POSITION OF PHYSCOMITRELLA PATENS. 


It must have occurred to every student of the mosses of the 
United States that the foot-note* at the bottom of page 39 of 
Lesquereux and James’ Manual indicated a curious state of 
classification, and this feeling is increased when a study of the 
specimens and literature convinces him that the facts which have 
been taken for fixing arbitrary lines are still open to question and 
have been disputed by several well-known bryologists. The 
questions in doubt are these:—Has Physcomitrella patens a de- 
hiscent, differentiated, lid? and is it distinct generically from 
Aphanorhegma? We propose to answer both these questions in 
the affirmative. 


The history with regard to European specimens is best shown 
by the following citations :— 


PHYSCOMITRELLA PATENS (Hedw.) Br. & Sch. 
Phascum patens Hedw. Descr. 1: 28. pl. ro (1787). 
Physcomitrella patens Br. & Sch. Bryol. Eu. 1: p/. 3 (1849). 
Aphanorhegma patens Lindb. in Ofv. K. Vet. Akad. Forh. 580 
(1864). 


Limpricht (Rab. Kryptfl. 4: 174. 1886) and Braithwaite (Brit. 
Mosses, 2 : 127. 1890) maintain Physcomitrella as a genus and state 
that the capsule is indehiscent, or the lid not differentiated ; 
hence the former places it among the cleistocarpous mosses, and 
the latter, following Lindberg and Hampe, classes it with the 
Funariaceae. The latter is undoubtedly its most natural alliance. 
Lindberg claims to have seen the lid. Schimper in the last edition 
of the Synopsis Muscorum (1876) says that he has not seen it. 

“ De operculi vestigi a Clarissimo Lindberg laudato adhuc nil vidi.” 


*« Aphanorhegma serratum, Sulliv., differs from this spec.es (Physcomitrella 
patens) only in the regular dehiscence of the capsule, which divides in the middle and 
is therefore considered as operculate or stegocarpous, though no discoloration nor any 
kind of modification of the texture is observable on the line of disruption. But for 
this regular dehiscence Aphanorhegma should be described here merely as a variety 
_ _ of Physcomitrella patens. It is therefore a remarkable connecting link between the 


: Ephemereae : nd the Physcomitrieae, which resemble each other also in the areolation 
of the leaves.” 


r 63 


We have taken pains to see all the available American ma- 
terial of Physcomitrella patens in the Sullivant herbarium as well 
as in that of the Department of Agriculture and others. In Drum- 
mond’s Southern Mosses, No. 5, collected near St. Louis, the 
specimens are still in perfect condition for examination. Many 
of the capsules have not split, but several of those which have 
show with a low magnification a perfectly circular rim. Under a 
high magnification, after boiling and mounting, the walls are torn 
irregularly, but a distinct median line of separation, where the 
cells are elongated with longer transverse walls, was seen, even on 
the irregularly torn capsules. 

No. 30, Sull. & Lesq. Musci Bor. Am. Ed. 2 (1865), collected 
in Ohio, also are in fine condition. Few of the capsules have 
split, but after mounting in glycerine jelly, three capsules did split 
regularly in half, and although we have seen no differentiation of 
the cells along the suture, yet from its regularity, we suspect it is 
there. It is almost impossible to determine this after the dehis- 
cence of the capsules, as the walls are so thin, the cells so tender, 
that in boiling to get rid of the spores, the walls are injured be- 
yond possibility of examination. The stomata are basal, and 
Seem partly immersed. | 

In Sullivant’s own herbarium we found specimens from Colum- 
bus, Ohio, collected in 1852, several capsules of which had split 
regularly in half as seen by reflected light with a low power, 

Quite recently we have received from Fort Snelling, Minne- 
sota, specimens collected by Grace E. Sheldon on October I, 
1894, which show a differentiation of the cells around the middle 
of the capsule. Fig. 8 of our-plate was drawn from these speci- 
mens. These capsules are very fresh and young, the cells of the 
walls still contain chlorophyll, and are more distinct than any we 
have seen. Starting from the stomatose base, the longest diame- 
ter of the cells is vertical, they measure .013%.027 mm., gradually 
the cells become larger, still keeping their longest diameter verti- 
cal, and twice the length of the shortest, measuring .027.054. 
At the line of dehiscence the cells are transversely elongated, 
three times longer than wide, and measures .o16xby .054 mm. 

Taking the above observations only for what they are worth, 
it seems to me that we may at least give the question the benefit 


64 


of a doubt, and would advise any one having access to fresh 
material of this rare and interesting species, and who is familiar 
with modern laboratory methods, to do some careful bleaching 
and staining, and try to prove conclusively whether P. parens be- 
longs with the Cleistocarpous mosses or not. 

The stomata as figured by our artist do not agree with fig. 59 
of Limpricht’s Laubmoose (Rab. Kryptfl. 4: 158), but in all our 
specimens they seem to be more or less immersed, and it is only 
in deep focussing that the outlines appear distinct. 

There seems to be some uncertainty as to the position of the 
antheridia in Physcomitrella patens. Schimper in the Bryologia 
(¢. 3) figures them in basal buds, but in the Synopsis Muscorum 
(Ed. 2, 1876) he corrects himself, saying they are axillary to the 

“upper leaves. As in Aphanorhegma they have also globose-tipped 
paraphyses. 

Limpricht says they may be either naked and pseudo- 
lateral under the perichaetium or on a branch, seldom at the base 
of the vaginule ; the paraphyses may be thread-like or globose- 
tipped. We have not been able to verify any of these statements 
from fresh American material. 

The genus Aphanorhegma was founded by Sullivant in 1848 
(Gray’s Man. Ed. i.,647. 1848). Physcomitrella was not published 
till 1849 (Br. & Sch. Bryol. Eu. fasc. 42). In his search after ear- 
lier names, Lindberg seems to have allowed dates to have more 
weight than facts. From the accompanying plates and descrip- 
tions, it seems to me that we cannot consider Physcomitrella pa- 
tens and Aphanorhegma serrata as congeneric, much less is one 
“merely a variety of the other.” 


Description of Plate 229, Physcomitrella patens (Hedw.) 
Br. & Sch. 
1. Plants, natural size. 
2-3. Two enlarged, showing the immersed —— 
"4. Outlines of leaves. 
5. Half of the base of one leaf enlarged, showing cells. 
6. Apex enlarged. 
7. Capsule enlarged, showing the stomatose base, and irregular line of dehis- 
’ ence, the parenchymatic walls and large apical cells. 
8. Cells of the walls still further enlarged, showing the transverse elongation of 
ogg in the middle of the capsule, the ey ones above. 
+ .g. Apex of capsule enlarged. 


oS ae 


\ > ees Sapa ea i. 7 ( ed 2-— 
rooting at base of the innovations ; leaves crowded, 


63 


ro. Spore. 

11. Same much enlarged, snowing the spines, 

12. ‘Stoma, enlarged, showing the overlapping cells, 
13. Calyptra. 


Description of Plate 230, Aphanorhegma serrata Sull, 
- Plants, natural size. 
. Same, enlarged. 
. Outlines of leaves. 
- Cells at base of one side of leaf. 
- Cells of the apex. 
. Perichaetial leaves removed, showing the antheridia mixed with the arche- 
gonia in the upper ax:ls, 
6-7, Distinct antheridial clusters. 
8. One of the globose-tipped paraphyses. 
9. Leaves removed, showing the subapical innovations of the stems and the 


Aw Bw dn 


nearly sessile capsules, } wt ety 
To. Single capsule enlarged, showing median dehiscence and flaring rim. 


II. Spores, one enlarged, showing spines. 

12. Collenchyma cells below the mouth showing the thickened walls. 
13. Cells of lid. 

14. Two stomata enlarged. 

15. Calyptra. 


2. On A Hysprip GROWING WITH APHANORHEGMA SERRATA Sull. 


(PLATE 231.) . 


These specimens were distributed in Drummond’s mosses of 
the Southern States (1841) as No. 20, labelled as follows: 


Schistidium Serratum, Nov, Sp. eee 
‘e ids . . a. 
Foliis obovatis acuminatis sub apicum serratis, capsula hemisphaeric 


Near St. Louis, growing with Phascum serratum. 

In the set belonging to the Columbia College Herbarium, there 
is an unmistakable “hybrid between Aphanorhegma serrata and an 
unknown Physcomitrium, probably P. turbinatum, though, of course, 
the hybrid is not characteristic, as the archegonial om = 
Aphanorhegma, and the antheridial plant was not distributed an 
could not be as easily determined. oe 
Aphanorhegma po @ X Physcomitrium aes sagt $ (?). : 

Schistidium serratum Hook. & Wilson in Drummond's Sout 
€rn Mosses no. 20 (1841). 

Plants 3-5 mm. high, gregarious, bushy ; stems b 


s 


ranching and 
3 mm. long, 


oe re ee i serrate 
lanceolate or oblanceolate, from an oblong base, margins 


66 


above, marginal cells inflated with oblique walls; vein ending 
below the apex, or excurrent into the subulate tip. Sporophyte . 
of two kinds. The normal, immersed ones of A. serrata, splitting 
in half when mature, becoming broadly flaring, with several rows, 
810, of dark collenchymatic cells bordering the mouth and form- 
ing the whole of the lid; the spores rough, .024-.027 mm. The 
hybrid sporophytes exserted ; seta 2-6 mm. ; capsule turbinate, not 
splitting in the middle, but with a smaller apical lid, which is 
bordered by one row of denser cells and is composed of parenchy- 
matic cells, as well as the walls of the capsule ; mouth bordered by 
two rows of dense oblong cells, and a persistent well-differentiated 
annulus ; spores .016—.018 mm., immature. 


Type locality near St. Louis, Missouri; type specimen No. 20, 
Drummond’s Southern Mosses, in the Herbarium of Columbia 
College. 

I wrote to Kew for information concerning Hooker and Wilson's 
specimens, and Mr. Wright says they also have a mixture in Drum- 
mond’s No. 20, but he does not mention any exserted capsules. 
“Some of the plants have broad leaves, similar to those of 
Phascum patens,as is shown by Sullivant in Mem. Acad. n. ser 
3:60. @ rr (1848) and in Sullivant’s Icones, ¢. 57. Others have 
longer, narrower leaves; and an almost sessile capsule, with a 
regular operculum. On another specimen (13b, Mississippi, 
Drummond) Wilson has written ‘In this tuft is also found Phascum 
patens.” 

Drummond's specimens were supposed to be the ones on 
which Sullivant founded his genus Aphanorhegma, but it is clear, 
on consulting his herbarium, that his set of Drummond’s mosses 
could not have supplied the figures which are given in the Me- 
moirs of the American Academy (3: 60. 4 2. 1848), for he had 
only a few old ragged plants of No. 20 with deoperculate cap- 
sules. He distributed in the Musci Alleghanienses No. 198 
(1846) specimens from Virginia and Ohio, as Schistidium serratum, 
and in the original description of Aphanorhegma in Gray's Manual 
he makes no mention of Drummond's specimens, and gives the 
distribution “from New England to Ohio.” These must be the 
types of the genus, and the ones figured in the Icones Muse. ¢. 57: 


67 
Description of Plate 231, 

All the plants taken from Drummond’s Southern Mosses, No. 20. 

1. A plant of Aphanorhegma serrata bearing three normal capsules, old, im- 
mersed and empty, with a flaring mouth, broader than the depth of the capsule; the 
Spores measure .024~.027 mm., and are rough and brown. 

1a. Hybrid sporophyte, with exserted seta 6 mm. long, the capsule not splitting 
in the middle, but with a smaller lid, the mouth bordered by two rows of cells and 
an annulus, the spores imperfect, .016-.018 mm. 

2. A plant of Aphanorhegma serrata with one old, flaring, immersed capsule, 
spores .024—.027 mm.; to the left, on a separate branch, two younger capsules, 


immersed and immature. 
3. The lower plant is Aphanorhegma serrata with a single immersed flaring 


capsule, the spores .027-.029 mm. 

3a. Another immersed, flaring capsule with mature spores .o27 mm. in diameter. 

3b. Four other branches, all bearing exserted capsules, less flaring than the 
other two and bordered by only two rows of cells with a distinct annulus, the spores® 
imperfect and massed together in the sporesac. 

3d. was found in place on 3c., but in boiling it fell off. 
cells of the lid are quite different from that of Fig. 6. 

4. A simple, unbranched plant of Aphanorhegxa serrata, showing the shape 


just after dehiscence. 
5. Same, enlarged, showing the 8 rows of thick, collenchymatic cells bordering 


the mouth. 
6. Lid of same, enlarged; also composed of collenchyma. 


7. Spores spinosely roughened, .024—.027 mm. from Fig. 4. 
8. A normal plant of Aphanorhegma serrata with two capsules, one old and 


flaring, the spores .024-.027 mm., the other just splitting in half, with the calyptra 
still on. 
3. On A European Hysprip OF PHYSCOMITRELLA PATENS. 
There has been recorded in Europe another hybrid which 
makes a curious parallel between Aphanorhegma and Physcomitrella, 
and it seems to have puzzled European bryologists, for they do 
not agree in the interpretation of the facts. The synonymy is as 
follows ne 
Puyscomirretta Hamper Limpr. (Rab. Kryptfl. 4: 175. 1886). 
Physcomitrella patens X Physcomitrium sphacricum. 
Physcomutrella patens var. pedicellata Br. & Sch. Br. Eu. 4 37 
(1849). 
Ephemerum patens var. anomalum Hampe in sched. 
Aphanorhegma patens var. anomalum Hpe.; Lindb. in Ofv. K. 


Vet. Akad. Forh. 580 (1864). ae ae 
Physcomitreila patens var. anomala Hpe.; Milde in Bryol. Sil. 


_ I9t (1869). : 


It will be seen that the 


68 


Milde says of it that the capsules are long-stalked with unmis- 
takable indications of a lid. These specimens were collected at 
* Blankenburg in the Harz Mountains by Hampe. 

Schimper, Hampe, Lindberg, Braithwaite, Husnot, Sullivant 
and Lesquereux and James (our American authorities probably 
cited the European) all state that P. patens may have the cap- 
sules either immersed or exserted, and Boulay states that it is very 
variable and that the different forms may be found growing 
“ pell-mell” in the same locality. 

Limpricht, in a foot note to his description of P. Hampei, says 
that Lindberg claims to have seen this form in the axils of the 

leaves of P. patens but that he has never seen it. 

_ We havea slight addition to make from our own observations, 
for in the Jeger Herbarium we find authentic specimens of 
Hampe’s labelled: “ Ephemerum patens var. anomalum. Seta 
exserta. p. Blankenburg. Ex. herbar. E. Hampe, 1865.” These 
specimens show what Limpricht says he has not seen, two kinds 
of capsules on the same plant. 

One plant which we have mounted has two capsules; the 
lower one small, immersed, with thin walls and characteristic large 
apical cells of Physcomitrella patens. The other is exserted on a 
thick seta 2 mm. long, with regular distinct cells in the walls of 
the capsules and a differentiated lid. The exserted capsules are 
almost all either young or aborted, so that no spores have been 
seen, but one old capsule has been found with the lid off, showing 
the mouth bordered with an orange-colored rim of cells and a dis- 
tinct annulus. The immersed capsules are on the lower part of 
the plants, as in our hybrid; most of the capsules are older and 


have perfected their spores with measure .024~.027 mm. and are 
rough. 


Japanese Characeae—ll, 
By T. F. ALuen. 


11. Chara coronata Ziz.,collected in Province Ise, differs in no 
essential point from the forms of Nos, 2 and 3. 

12. Nitella Faponica Allen, from a new locality in Province 
Ise, namely Yamagami ; distributed with No. 14, under the ex- 
_ siccatae number 9. 


_* 


69 


13. NITELLA PULCHELLA sp. nov. 

Nitella polyarthrodactyla, monoica, gloeocarpa. 

Stems about 460 in diamater. Verticels consist of eight 
or nine leaves (with an occasional simple, undivided leaf, but 
not heterophyllous.) Leaves about 150 in diam., thrice di- 
vided, primary segment much longer, (2680) ; first node bears 
6-7 secondary segments, 68 diam., 600 long; second node 
bears 5 tertiary segments, 34 in diam., 175 to 200 long; oc- 
casionally one of these divisions is undivided like a simple ter- 
minal with three cells; the third node bears four to six (usually 
four) two-celled terminals, 25. diam., 170 to 240 long. The ter- 
minals are two (rarely 3) celled, the cells about equal in length; 
the terminal cell is not mucroniform, but terminates, rather ab- 
ruptly, in a sharp point (some collections more elongated and 
slender) ; other specimens, terminals very short, forma brachyteles. 
The fertile verticils are somewhat compact and borne upon pedi- 
cels arising from the stem within the primary verticils ; the entire 
fertile verticil becomes a globular, gelatinous mass. The leaves 
of the fertile verticil are usually scvice divided, the antheridia 
borne on the second node are about 225 in diam., and decidedly 
stipitate, the stipes 200 long and 54 indiam. The oogonia are 
borne on both nodes of the leaf, single; usually on the terminal 
node, sessile, the coronula minute. 

- The oospore, dark reddish, is 250 long, 200 broad, with 7-8 
prominent ridges, the surface is strongly reticulated, the reticulae 
5-12 in diam. : 

The relationship of this beautiful species is not clear, in some 
features it may be related to NV. ¢érichotoma A. Br., but it is clearly 


separated from it as well as from the subspecies Zeyhert and 


Lechleri. 2s 
It may be considered remotely similar to WV. gelatinosa A. Br., 


of Australia, where species of the polyarthrodactylae-gloeocarpae 
section most abound, but this species remains quite unique and 


individual. 
No. 14, Nitella Japonica, additional collections, more mature, 


from Yamagami, Province Ise, distributed as exsiccatae No. 9. 


No. 15, NITELLA SUBGLOMERATA JAPONICA var. NOV. 

Verticels consist of a large number of leaves, often 12 to 16, 
apparently irregularly in a double series, some eo some 
shorter, but all twice divided ; the first node bears t ei sea i 
and three secondary divisions, the second node also he t oe 
oogonia and three terminals; rarely one of the secon ay ivi 
sions does not form a node, but remains as a simple one-celled ter- 


70 


minal; the terminals of the second node, usually three in number, 
are one-celled and acuminate above the middle, terminating in a 
sharp point which is solid. . 


The oogonia are aggregated, three together, at each node of 
the leaf, not closely sessile, with a rather large and persistent 
coronula, the lower cells of which appear to be spreading, as in 
some sub-species of the polygiochin group, in all from 350 to 400 
long. The oospore is marked with 6 striae, 204 long, 180 to 190 
broad. The membrane of the spore is strongly reticulated, the 
reticulae averaging 5 “in diameter. This Japanese form differs 
mainly in the somewhat smaller spores, which are much more 
strongly reticulated; the antheridia also are smaller, 200 in diam. 
This species was collected in Mikawa, Seishin pond, and distrib- 
uted as No. 15 of my Japanese exsiccatae. 

No. 16. NITELLA SUBLUCENS sp. nov. 

Nitella diarthrodactyla, homoeophylla, monoica macrodactyla, 
subflabellata, gymnocarpa. Fertile verticils contracted into dense 
terminal or axillary heads, long overtopped by the sterile leaves; 
verticils very dissimilar. The long sterile leaves are surmounted 
by a crown of about four minute two-celled leaflets. Fertile 
leaves twice- rarely thrice-divided, terminal segments usually four, 
short, two-celled, the lower of the two cells inflated, theterminal 
cell a sharp mucro, oogonia clustered at the base of the fertile ver- 
ticils and at the first node of the leaves; oospore, 285 long and 
broad, globular, six or seven striate. The present specimens are 
too immature to determine the character of the membrane of the 
spore. Antheridia about 200 in diameter. | 

This species is closely allied to WV. translucens (Pers.) Ag; from 
which it differs by its smaller size, and especially the much 
smaller oospore (and its locality). 

This species was collected in Sagami, Kodsu, and distributed 
as No. 16 of my Japanese exsiccatae. 


17. Chara gymnopitys A. Br. var. “ alpha” A. Br. 

This form, though not exactly corresponding to gymnopitys 
genuina A, Br., seems to be on the border between that and var. 
(deta) duriuscula A. Br., both from Australia; and as A. Braun 
has left var. “ Alpha” blank, it may well be occupied by this 
variety. The stems are about 306 in diam., with small, conical, 
broad-pointed spines; leaves about 8, stipules 16, the cortex cells 
double, alternately large and small, often with additional, partially 


71 


developed cells, so that sections show about 20 cortex tubes; 
bracts at the leaf-nodes equally developed. all around, slender, 
long, acuminate-pointed, about 60 in diam. The oogonia seem to 
be distinctly stipitate, the oospores average 560 long by 365 to 390 
broad, with 8 (10-11) striae. The coronula is short, broad and 
square-shaped. Antheridia are rather rarely found, 293 in diam.; 
when present, conjoined. 

This species was collected in Mikawa, Tennu pond, and dis- 
tributed as No. 10 of my Japanese exsiccatae. 

No. 18, same species and variety as the last, collected in To- 
kio, Shinbashi pond, and distributed as No. 11 of my exsiccatae 
Japonicae. Nos. 19, 20 and 21, Nitella pulchella Allen, collected 
respectively in Chikubushima pond, Mikawa Tennu pond, and 
Tokio Shinbashi pond, and distributed as Nos. 13 and 14 in my 
exsiccatae Japonicae. 

As a matter of record it may be well to add that the following 
list of Exsiccatae Japonicae has been distributed as far as speci- 
mens would allow; of WV. sudblucens Allen only fourteen speci- 


mens could be sent out. 


CHARACEAE JAPONICAE EXSICCATAE. 
DISTRIBUTAE A T. F. ALLEN. 


I 
1 Chara fragilis Desv. 11 Chara gymnopitys A. Br. 
2 Nitella coronata Ziz. var “alpha” A. Br. 
3 Nitella coronata Ziz. 12 Nitella paucicostata Allen. 
4 Nitella Japonica Allen, 13. Nitella pulchella Allen. 
5 Nitella mucronata A. Br. 14 Nitella pulchella Allen. 
yar. tenuior A. Br. 15 Nitella subglomerata A. Br. : 
Nitella Japonica Allen. var. Japonica Allen. 
Nitella orientalis Allen. 16 Nitella sublucens Allen. 
Nitella pulchella Allen. 17 Nitella oligospira A. Br. 
Nitella Japonica Allen. 


CO ON a 


10 Chara gymnopitys A. Br. 
var. “alpha” A, Br. 


i 


Tradescantia Virginica var. villosa Watson. 


There are doubts whether this is a variety of 7radescantia Vir- 
ginica L. or should have specific rank. Forms are found which 
apparently connect the extremes and make it difficult to draw the 


' 


72 


line of separation. As the two are found growing in company at 
Forest Hill, in the southern border of Chicago, and where they 
are clearly distinct, I have had a good chance to observe them for 
several years in their native condition. I first met with the vari- 
ety in 1878, and have watched it more or less since. At that 
time it was not easy to refer this low or dwarf plant, broad leaved, 
green and early maturing, to 7vadescantia Virginica as described 
in the books, usually a much taller, smooth, narrow-leaved and 
glaucous plant, which kept on flowering throughout the summer. 
Still more than their different look, it was their early flowering 
and disappearance which particularly called attention to them. 
Rafinesque, in the species which he made out and gives in his 
_ “New Flora and Botany of North America,’ calls some “ vernal” 
and others “estival.”* This holds good between the two found 
here. Notes made in 1878 give the time of gathering as May 
30th. Those of 1880 state that hundreds were examined on June 
11th, and none were in flower, all having gone to seed. 7vades- 
caniia Virginica was then fairly in its season of bloom, only a few 
having passed that stage. May 12, 1894, a few of the low kind 
were found in flower; on the 21st they were in great abundance. 
At the latter date but three of the other form were found in flower 
after a long search. On June 19th half a dozen flowers of the 
low form were discovered after a search of an hour or more in 
spots where the plants were most numerous. The taller form was 
then in full bloom everywhere. The stems of the low form were 
mostly dead or dying, some lying flat on the ground. Their sea- 
son was virtually over. A number of plants of 7: Virginica were 
seen in flower August 16th. The last one observed was on Au-. 
gust 29th, except a single plant on October 2d, probably a case 
of flowering the second time. The broad leaved form behaves 
like a vernal plant, early in the season maturing its buds for the 
coming year, its aerial parts then disappearing, a process mostly 
completed by the last of June. . Some of the other form will have 
passed their floral season then, but others keep up the succession 
so that they are common or even abundant in July. I have as 
yet detected the low kind in but one locality, while the other 
grows in profusion wherever the conditions are suitable. 


*lic. p. 84 


—— 


ne Re OR reg 


73 


The two do not intergrade here, something which might be 
expected where they are so intimately associated in some places 
that both forms can be taken up with the same handful of earth, 
with their roots intermingling. Hence there is little difficulty in 
distinguishing each, their involucral leaves in most cases being at 
once decisive. Sometimes the typical form becomes low, and oc- 


casionally pilose, generally in poor soil or in the sand region, but 


it preserves the main characteristics of the plant, and rarely de- 
ceives one. 

As found here the following descriptions will give their points 
of similarity or difference: 


TRADESCANTIA VIRGINICA L. 

Plants glaucous, stems single or clustered, 1-5 feet high, gen- 
erally 114-3 feet, simple, or frequently branched, smooth. Leaves 
channeled, narrow, linear to linear-lanceolate, scarcely ciliate ex- 
cept at base, the sheaths and base of the leaves on the lower part 
of the stem sometimes pilose. Involucral leaves mostly shorter 

than the stem-leaves, generally much shorter, often abruptly con- 

tracted from an ovate base, which usually sheaths the umbel when 
in bud. Umbel simple, many flowered. Peduncles and calyx 
smooth. Flowers blue, varying to purplish blue. Roots coarsely 
fibrous, variable in color, whitish to yellow. 


Abundant in open woods, fields and borders of woods. 
May—August. 
T. Virernica L. var. vi.tosa Watson. 

Plants green, rarely glaucous, stems single or oftener clustered, 
2-15 in. high, mostly 5-10 in., simple or occasionally branched, 
sometimes flexuose, smooth or hairy. Leaves flattish, promi- 
nently nerved, broadly linear to linear lanceolate, ciliolate, hairy 
or roughish with short hairs, especially the upper, the lower be- 


_coming smooth or smoothish. Involucral leaves usually longer 


and broader than the stem leaves, %4-11% in. wide, gradually” 
tapering from the base, which scarcely or not at all sheaths the sim- 
ple, many flowered umbel. Peduncles and calyx villous. Flow- 


-€rs commonly purplish-blue, varying to blue. Roots coarsely 


fibrous, yellow, often deep yellow. : 
Woods or borders of woods, frequently in rather dense woods. 


May-—June.* 


* Torrey (Flora of the Northern and Middle Sections of the United States, 1824, 
335); ene: Bs T. Virginica as if he had this form in hand. He says pe is 
‘‘about a foot high,” the involucrum “large, 2-leaved,” « flowers algae h "2 
is given as the time of flowering. He adds, “ My specimensare from Lake Michgan. 

! By. OL. 


Cuicaco, ILL. 


‘ 


74 


Some new Hybrid Oaks from the Southern States. 
By JouHNn K. SMALL. 
(PLATES 232-235.) 


The following is a record of some observations on several 
interesting forms of Quercus growing in North Carolina and 
Georgia, together with a striking hybrid existing in two well- 
marked forms, found in Lake county, Florida, by Mr. Geo. V. 
Nash, during his collecting trip of last season. : 


Quercus PHELLOS X Q. DIGITATA. 


A large and stout tree with rough scaly bark, reaching a 
height of from twenty to thirty-five meters, and. having a trunk | 
diameter ranging from six to nine decimeters. Trunk forking 
several feet from the ground, the divisions thence branching, the 
branches rather erect and the branchlets straggling ; leaves oblong, 
obovate or oblanceolate, 5-20 cm. long, 2-10 cm. broad, mostly 
entire and undulate or somewhat crisped, or more or iess twO-  ~ 
_ lobed or three-lobed near the apex, acute or obtuse at the apex, 

acute, obtuse or subcordate at the base, the upper surface dark- 
green and glabrous, the lower brown and more or less tomentose 
with reddish brown, stellate hairs, especially about the midrib and 
principal nerves ; mature fruit not seen. (Plate 232.) 


Hills west of the Falls of the Yadkin River, North Carolina. 

In 1892 I found a small grove of peculiar looking oak trees in 
a very shallow depression in the foot-hills of the Falls Mountains, 
just west of the Falls of the Yadkin River, in Stanley county, 
North Carolina. Specimens were collected, but there was not 
time for a thorough investigation of the case. The specimens 
suggested a form of Q. Rudkini(Q. Phellos * Q. nigra), and some 
were distributed under that name. Each succeeding year I have > 
observed the trees and their surroundings, and am now confident 
that the parents are not those of QO. Rudkini, but Q. Phellos and 
Q. digitata, the two prevailing species of the immediate region. 
Q. mgra, one of the undoubted parents of Q. Rudkini, was not 
observed within several miles. The form of Q. digitata, which iS 
apparently one of the parents of the hybrid under considera- 
tion, is not that with long, falcate leaf-lobes, but one common 
through the pine woods in the middle country of the Southern — 
States. Its leaf is not as deeply lobed and has a more cuneate 


— 


75 


outline. As in the case of the following hybrid, this one pro- 
duces on the same branches leaves almost identical in shape with 
those of the parents. Many, however, are intermediate, in vari- 
ous degrees resembling one or the other of the parent forms. 
The texture is about intermediate, and the pubescence on the 
lower surface less than in Q. digitata and much more than exists 
in true QO. Phellos. 

The cup and acorn, although not mature, each exhibit charac- 
ters which suggest Q. digitata rather than Q. nigra. Most of the 
trees noticed had the peculiar habit of forking about three feet 
from the ground into two erect secondary trunks. This charac- 
ter, together with the striking irregularity in the shape of the 
foliage, makes the trees quite conspicuous among their associates. 


QUERCUS GEORGIANA X Q. NIGRA. 

A small tree of a dark-green color and a somewhat straggling 
mode of branching, ranging from three to ten meters in height and 
having a trunk diameter varying from ten to twenty-five centimeters 
clothed with a dark, smooth, glabrous bark, which becomes rough 


onthe trunk. The leaves are mostly obovate in outline, sometimes 


oblong, 4-20 cm. long, 2.5-15 cm. broad, 3-5-lobed, dark green 
and shining above, brown and dull beneath, glabrous on both sur- 
faces, except a small tuft of stellate hairs in the axils of the nerves 
on the lower side, obtuse or acute, equilateral or inequilateral at the 
base, the, sinuses either shallow or deep, sometimes penetrating 
almost to the midrib; the lobes rounded or square-oblong and 
slightly lobed at the ends, all ending in a slender apiculation; fruit 
12-15 mm. long, 13-16 mm. broad, short-stalked, peduncle 4-6 
mm. long; cup saucer-shaped or slightly turbinate, reddish, the 
scales triangular, the acorn depressed-globose, pubescent, some- 
times sparingly striped, ending in a short, abrupt apiculation. 
(Plate 233.) 

Northern slope of Stone Mountain, DeKalb county, Georgia. 

On the northern slope of Stone Mountain, at an altitude of 
about 1300 feet, there is a grove of stunted trees of various 
species of oaks, Quercus Georgiana and Q. nigra predominat- 
ing. I had been in the grove a number of times, but noticed 
nothing peculiar until January, 1894, when my attention was 
called to some odd shaped leaves on the ground and anomalous 
acorns on the branches overhead. This material suggested a hy- 
brid form. Last September I visited the same spot to secure 


fresh foliage from the trees and make some further observations. 


: 76 


Q. Georgiana and Q. nigra were the only species in the imme- 
diate vicinity, and the trees in question appear intermediate be- 
tween the two. In place of the graceful port of the preceding 
‘species there was a certain amount of the rugged habit of the 
latter. The texture of the foliage is intermediate, and leaf forms 
suggesting both species occur on the same branches. Mature 
fruit also has traces of the characters of that of both the parents. 
The accompanying plate gives some of the extreme leaf forms. 


Quercus CATESBAEI X Q. CINEREA. 


Form A, in which Q. cinerea predominates (No. 1586). 

A small tree with the habit of Q. civerea, ranging from two and 
one-half to four meters in height, with a trunk diameter ranging 
from six to twelve centimeters, branching about one meter from the 
ground; the branches somewhat spreading, clothed with a smooth, 
striate bark; the young shoots tomentose with dark-colored 
trichomes. The leaves are narrowly obovate or oblanceolate, some- 
times elliptic or lanceolate, 5-15 cm. long, 2-6 cm. broad, entire, 
undulate and more or less crisped or partially 3-7-lobed, either on 
‘one side or on both, acute or acuminate at both ends, short-petioled 
or sessile, the upper surface light and very bright green, the lower 
surface lighter but rather dull and more or less tomentose, the mid- 
rib and nerves white but not prominent above, prominent beneath, 
the lobes short-apiculate; cup hemispheric-turbinate, 13-14 mm. 
broad, 10 mm. high, nearly sessile. (Plate 234.) 


Growing in dry, sandy soil,in high pine lands on the road be- 
tween Umatilla and Lake Ella (about two miles from the latter 
place) in Lake county, Florida. 


Form B, in which Q. Catesbaei predominates (No. 1577)- 


_ Asmall tree, with much the habit of Q. Catesbaei, reaching # 
_ height of two or three meters and having a trunk diameter of six t0 
nine centimeters, the trunk branching from within 3 or 5 dm. of 
‘the ground, the branches more spreading than in the former. 
Branchlets conspicuously marked with white lenticels, the young 
twigs white-tomentose; leaves mostly oblong in outline, some 
times obovate, 6-15 cm. long, 3-10 cm. broad, usually 5—7-lobed, 
sometimes 2~-3-lobed or nearly entire, more or less irregular 
and inequilateral, acute or obtuse at the base, short-petioled, 
the lobes narrow, acute and apiculate by a long, sharp bristle, 
somewhat tomentose beneath, the nerves prominent and conspicu- 
ous on both surfaces. Flowers and fruit not seen. (Plate 235) 3 


_... Grows in dry, sandy soil in high pine lands, on the road beé- 
tween Umatilla and Lake Ella (about three-fourths of a mile from 
_ the latter place) in Lake county, Florida. : 


77 


Family Nomenclature. 


By Dr. V. HAvarp. 


Much credit is due Mr. Barnhart for taking up and elaborating 
this important problem which now forces itself upon the attention 
of botanists for a satisfactory solution. Uniformity and stability 
in family nomenclature are as desirable and necessary as in genera 
and species, and therefore such nomenclature must be built upon 
secure foundations and placed within the dominion of law. 

The advantages of a common ending for all family names are 
obvious and generally recognized, and the ending most habitually 
used is aceae. There seems to be no valid reason why it should 
not be applied under all circumstances; it does no violence to 
logic, grammar or analogy, and there should be no difficulty in 
Sacrificing custom and tradition to law and uniformity. 

To this extent I agree with Mr. Barnhart, and I also recognize 
the wisdom of not going behind the Species Plantarum of 1753. 

To his rule that the root of the family name shall always be 
that of the accepted name of a recognized genus belonging to 
that family, I am inclined to demur, and would instead offer the 
following propositions : 

1. The name of each natural family shall be the oldest name 
properly published, changing its termination into aceae if other- 
wise ended. 

2. The name shall be properly published, if in Latin, in the 
plural number, and accompanied with description clearly based 
upon one or more genera of said family. 

My reasons for dissenting are the following: 


1. Priority is the fundamental law of nomenclature and must 


dominate every other consideration. As the original author of a 


genus name is allowed entire freedom in his choice (within the 
limits of correct construction) and receives credit for it, so should 
the author of a family name. 
2. Genera are often separated only by minor wad secondaty char- 
acters, so that their limits are variously estimated, and their names 
accordingly liable to change with the discovery of new species. 
Should the family name share the risk of this instability? 


\ 


78 oe 


3. A family name constructed from a genus name logically 
implies that said genus is the most important, and that the char- 
acters of the family are, in the main, those of this genus, while in 
reality the genus may be one of the least important and materially 
differ in its structure from that of other genera, ¢. g., Cassiaceae, 
which is proposed for Leguminosaceae and Carduaceae for Com- 
positaceae. 

4. A family name should, so far as possible, describe a charac-' 
ter common to all or a great number of genera in the family, and 
this seems to have been the praiseworthy aim of several of the 
distinguished authors to whom we owe the existing nomenclature. 
The fact that plants of other families may have the same character 
does not matter; there can be no hard and fast line in classifica- 
tion. Forthis reason I prefer Cruciferaceae to Brassicaceae, Gram 
naceae to Poaceae, Contferaceae to Pinaceae, Umbelliferaceae to . 
Ammuaceae, etc., even though it is true that umbels are also found 
in Araliaceae, and cruciform flowers in Capparidaceae. 

5. Our main object being stable uniformity, the rule which 

will accomplish it with the least disturbance, and preserve the 
most of the familiar old names should be preferred ; if priority 1 
sufficient for the purpose we must reject ane more radical and sub- 
versive rule of Mr. Barnhart. 

Concerning tribal names I presume there is no question that 
they should invariably be formed from the root of a generic name 


with the addition of the termination eae, as is the usage of our 
best authors. 


Reviews. 


The 2 Life and Writings of Rafinesque siPreoared for the Filson 
Club and read at its Meeting, Monday, April 2, 1894. By Richard 
Ellsworth Call. 

Filson Club Publications, No. 10. 4to, broad margins, pp. 227- 
plates 5. John P. Morton & Co., Louisville, Ky., 1895. 

This splendid volume, creditable both from a literary and from 
an artistic standpoint, had its inception, so the preface informs 
oe, in an attempt to clear up certain matters connected with 
S ue synonymy of the Poiawides,, in which family of molluscs 


79 


Rafinesque was much interested and in which he did such valuable 
work. 

The Filson Club is an organization having for its object the 
collection and preservation of original matter connected with the 
history of the State of Kentucky, hence the publication of this 
volume as a recognition of « the first resident professor-naturalist 
Within the limits of the State.” 

Constantine Samuel Rafinesque [Schmaltz]* was born in Ga- 
lata, near Constantinople, October 22, 1783. His father was 
French, his mother of German parentage. His early life was 
spent in France and Italy, and his first essay, written in 1796, was 
in the form of a journal, describing a tour to Genoa. That he 
early had the instincts of a naturalist is manifest, for by the time 
he was fifteen years old he had made collections of plants, fishes, 
birds and shells, and apparently took but little interest in boyish 
Sports or games, or in association with other youths of his own 
age, 

His first visit to America was in 1802, and here he remained, 
mostly in the vicinity of Philadelphia, until 1805, when he re- 
turned to Italy. There he remained fora period of ten years, 
during which time he wrote many papers, dealing not only with 
Matters in connection with his surroundings, but also others based 
upon his observations and. experiences while in America. His 
domestic life was unhappy, however, and doubtless had much to 
do with his subsequent eccentric manners. In his description of 
Sicily he says: « She offers * * * a fruitful soil, a delightful 
climate, excellent productions, perfidious men, deceitful women.” 
In 1815 Rafinesque left Europe forever and set sail for America 
Once more. His arrival was dramatic in the extreme, and doubt- 
less still further heightened his pessimistic ideas. At midnight, 
November 2d, in a dense fog, the ship ran aground off Fisher's 
Island, in Long Island Sound, and all the results of his years of 
toil, mercantile and scientific, were lost. 

He finally came to New York and accepted a position as pri- 
vate tutor in a wealthy family. He soon resigned this position, 


* He added this, his mother’s name, for prudential reasons, on his return to ltaly, 
in order that he might pass for an American and thus avoid certain political compli- 
cations in which he feared he might become involved. 


80 


however, and went to Philadelphia, where he found several good 
friends and other persons, it seems, who were not, for we hear of him 
during his stay there as having been bankrupted in a business ad- 
venture and defrauded of all his savings by the falseness of a fel- 
low-countryman, to whom he had intrusted his affairs while on his 
exploring expeditions into what was then called the West, between 
the Alleghanies and.the. Mississippi. 

Through the influence of one of ‘his’ Philadelphia friends he 
secured an appointment to a professorship in the Transylvania 
University, at Lexington, Kentucky, upon the duties of which — 
position he entered in the fall of 1819. Here he passed seven 
busy years, during which time he made extensive collections, ¢S- 
pecially in conchology and botany, wrote and published many 
papers, attended to his duties at the University and acted as sec- 
retary of the Kentucky Institute—the first scientific society formed 
within the State. One of his great ambitions seems to have been 
to found or to assist in the foundation of a botanic garden at Lex- 
ington, and in 1823 he presented the matter before the State Leg- 
islature with such success that the Senate psssed a bill to that 
effect, but it failed of passage in the Assembly. He then under- 
took to push the scheme by means of private subscription and the 


_. formation of a joint stock company. An act of incorporation 


was secured, ground was purchased and planting was actually 
begun, but those upon whom he relied failed to meet their obliga- 
tions and the attempt was finally abandoned. This embittered 
and saddened him still further, and he says: « * * * this garden 
would have been my delight; I had traced the plan of it, with a 
retreat among the flowers, a greenhouse, museum and library; 
but I had to forsake it at last and make again my garden of the 
woods and mountains.” 

He does not seem to have been in touch with his associates in 
_ the University, who took but little interest in his scientific work, 
and he was doubtless impatient of their criticisms and indiffer- 
ence. Added to this, the students ridiculed him and finally, in 
June, 1825, he left Lexington and once more made his way t® 
Philadelphia. During the next fifteen years he seems to have 


lived in a hand-to-mouth manner, practicing medicine in his ow? 


: _ way, lecturing at the Franklin Institute and assisting in the estab- 


i 


- + Re, 
Ei. ane es 
(iibinies 


8] 


lishing of a savings. bank. This latter seems to have met with 
Success, but although it was in existence at the time of his death, 
his end came to him amidst surroundings of absolute poverty, 
alone, in a garret in a poor quarter of Philadelphia in 1840.* 

His estate consisted entirely of personal property—mostly 
books, unpublished manuscript and specimens of natural history. 
The latter had suffered much from his inability to properly care 
for them, and a large part of his plant collection was sold as waste 
Paper. Eight dray loads comprised the material which he left 
behind. Some of this is yet preserved in the National Museum; 
another portion was secured by the University of Pennsylvania 
through Mr, Isaac Burk, and many of his botanical specimens 
finally went to the Jardin des Plantes in Paris, and the Philadel- 
phia Academy of Natural Sciences. The final settlement of his 
estate left it indebted to the administrator in the sum of $14.43. 

This, in brief, is the outline of his life as given in the first sixty- 
nine pages of the volume. Accompanying this are two portraits, 
One taken from a painting in the collection of the Wisconsin His- 
torical Society, the date of which is uncertain, and another as he 
appeared in 1810 at the age of twenty-seven, besides which there 
is a photographic reproduction of one of his letters to DeCandolle, 
written in 1838. 

Following this is a bibliography, arranged in chronologic 
Sequence, enumerating 203 publications, containing 420 titles. It 
is almost impossible to conceive, in these days of specialization, 
the wide range of his mental activity of which these give evidence. 
Botany, Zodlogy, History, Social and Political Economy, Mete- 
orology, Geology, Poetry, Philosophy, Book Reviews, Jour- 
nals of Travel, Astronomy, Physics, Archaeology, Ethnology, 
Medicine and other subjects which might be classed as subdivi- 
sions of the above, all received his attention. 

There is also a chapter devoted to a list of publications in 
which Rafinesque or his works are mentioned by other authors 
and an appendix giving a copy of his will, which, in addition to 
the part describing as to how he wishes his property to be dis- 


* According to an article by Thos. Meehan, published in the Philadelphia Public. 
Ledger a few years ago, the exact date of which is unknown to me, Rafinesque is 


Said to have died September 18, 1842. 


Pees 


walle 
ee ae 


82 
posed, contains the clause, “I wish my body to be burnt rather 
than buried, as I do not want to contamine the earth by decay, 
nor be a cause of desease to other men. My ashes, if they can be: 
collected, I wish to be deposited in a Urn, to be kept with my 
collections.” 

Apparently his property and certainly his body, was not dis- 
posed of in accordance with either the letter or spirit of this docu- 
ment, and his biographer says that he “appears to have been de- 
spoiled of his rights in nomenclature while living; he was de- 

‘spoiled of his possessions when dead.” : 

Many of the most interesting details contained in the volume 
are necessarily omitted in this review, but attention should be 
called, at least, to the titled headings, “ Rafinesque’s Name in 
Nomenclature,” where he is commemorated both generically and 
specifically in botany and zoology ; “ Medals, Diplomas and Other 
Honors,” in which a list of these, conferred upon him by scientific 
societies at home and abroad is given; “ Rafinesque and Evolu- 
tion;” “ Rafinesque’s Literary Style;” “ Personal Appearance of 
Rafinesque,” etc. : : 

The volume is a model of its kind and isa fitting tribute to the 
versatile man whose life it memorializes, A. H. 


Annual Report of the State Botanist of New York. Charles H. 

Peck. pp. 48 (From the 47th Report of the New York State 
- Museum of Natural History, Albany, 1894). 

Mr. Peck’s report for the year 1893 contains a great deal of 
important and valuable information. It deals with the poisonous 
toadstools ; gives a list of plants added to the State Herbarium, 

| of which 40 species were not before represented, gives notes on 
species not before reported in any of his communications, there 
being 14 new Fungi here described in the genera Psathyrella, 
_—- Merulius, Stereum, Discosia, Haplosporella, Rhabdospora, Volutella, 
_ Cercospora, Peziza, Sphaerella, Clavaria, Leptothyrium, Zygodesmus, 
 Asterula and Melogramma, not all from New York State, how- 
_ ever; and under “ Remarks and Observations” we find a variety 
_ of interesting notes on distribution and on characters of numerous 
_ species. Dr. E. C. Howe contributes the description of Carev 
Peck n. sp., the C. Emmonsii elliptica Boott, considering it more 
nearly’ related to C. deflexa than to C. Emmonsii. NL. B. 


7 aS 
mene 
a 


83 


Science. This weekly journal of research and discovery, 
abandoned for some months has been revived under most favorable 
circumstances and bids fair to give American students a reputable 
and authoritative medium of rapid and regular publication. It is 
published under the codperation of the following Editorial Com- 
mittee: S. Newcomb, Mathematics; R. S. Woodward, Mechanics . 
E. C. Pickering, Astronomy; T. C. Mendenhall, Physics; R. H. 
Thurston, Engineering; Ira Remsen, Chemistry ; Joseph Le Conte 
Geology ; W. M. Davis, Physiography ; O. C. Marsh, Paleon- 
tology; W. K. Brooks, Invertebrate Zoology; C. Hart Merriam, 
Vertebrate Zodlogy; N. L. Britton, Botany ; Henry F. Osborn, 
General Biology; H. P. Bowditch, Physiology; J. S. Billings, 
Hygiene; J. McKeen Cattell, Psychology; Daniel G. Brinton, J. 
W. Powell, Anthropology. 

Volume I of the new series was begun January 4th, and the 
seven numbers so far issued are replete with matters of both 
general and special interest. In these days of extreme special- 
ization and enormous publication, the student tends to confine 
himself to his chosen sphere of thought, and to know as much 
aS possible about that little sphere, and it would seem in many 
cases as little as possible about everything else. This tendency 
has been especially noticable in America, and we may now hope 
that it will be in large measure corrected by the publication of 
the journal here noticed. 

Manuscript intended for publication and books or papers in- 
tended for review should be sent to Professor J. McKeen Cattell, 
Garrison on Hudson, N. Y. The subscription is fixed at $5.00 a 
year and should be sent to the Publisher of Science, 41 East 49th 
St., New York City. We bespeak a cordial support of the enter- 
prise by all American botanists. 

Additional Notes on the new Fossil, Daimonelix ; its Mode of 
Occurrence, its gross and minute Structure. E. H. Barbour. 
Univ. Studies, Univ. Neb. 2: 1-16. fl. 7-72. 

This exceedingly interesting contribution to the literature of 
problematic organisms is the second one on the subject by the 
author since the original ‘‘ Notice of new gigantic Fossils,” pub- 
lished in Science, 19: 99-100. jigs. I—3. The name has evidently 


‘seemed somewhat provincial to the author, so he begins the paper 


84 


with the modest excuse that it was adopted in deference to the 
ranchmen and early settlers who used to know the fossils as 
« Devil’s corkscrews,” or “ fossil twisters.” They have been ex- 
amined by botanists, geologists and paleontologists and have 
been considered as plants, animal burrows and “ accidents.” If 
they are to be classed with the latter, the author justly says: 

“Such accidents should be immortalized.” Sections examined 
under the microscope, however, show a structure which is cellular 
but not vascular, and the author concludes that they represent a 
new order of aquatic plants, “ resembling the red sea weeds more 
closely than anything else.” The illustrations are to be particu- 
larly commended, as they represent the location of the fossils in 
the field at a distance, single specimens close to, microscopic Sec 
tions and a general view of the collection as it appears in the Mu- 
seum of the University. A. H. 


American Algae. Century I. 1894. Josephine E. Tilden, Uni- 
versity of Minnesota, Minneapolis, Minn. 

The first distribution of this collection of fresh water algae 
from the upper Mississippi Valley has made its appearance. 
All the specimens, including some rare forms, are mounted upon 
good white card paper in a quarto volume. Labels, includ- 
ing descriptive details, are printed, and the material has evidently 
been determined and arranged with great care. The work as 4 
_ whole has only commendable features. It will certainly prove to 
be of great value to the few herbaria who may be fortunate enough 
to acquire it, as the edition is limited to twenty-five copies. 

3a 


Through Glade and Mead.—Under this attractive title Mr. 
Joseph Jackson presents a most pleasing account of the flora of 
of Worcester county, Mass. This large county, extending across 
the centre of the State, embraces a very diversified topography- 
We have the high peak of Mt. Wachusett, with a few plants ap- 
proaching the sub-alpine character; the rocky cliffs of West 
minster, the Purgatory Chasm near Uxbridge, and many fine 
__ lakes and ponds, swamps and forests. All this region Mr. Jack- 

_ son has made peculiarly his own. He takes the reader at once 
e into his confidence, and with charming talk by the way leads 


85 


him to his chosen haunts. He knows the poets well, and culls 
from them their sweetest thoughts. In these days, when so much 
of our botanical reading has an aggressive and controversial turn, 
it is refreshing to escape into the fens and forests. Every page 
in this book is odorous with wild flowers and tuneful with bees 
and birds. 

The volume is amply illustrated by photogravure impressions 
of actual Worcester county scenes, or of typical wild flowers. 
Thus the peat-meadow given opposite to page 196 is enough to 
fill a botanist’s soul with envy; and who would not love to wan- 
der through the path in the frontispiece ? 

The series of familiar papers is followed by a catalogue of the 
plants thus far found in the county, following the last edition of 
Gray’s Manual as to nomenclature. The author, who is eminently 
fair minded, says: “In the present state of botanical nomenclature, 
and for the purpose for which this catalogue is intended, I have 
not thought it wise or necessary to adopt proposed changes not 
yet generally accepted. The student in such matters can easily 
adapt himself to changing conditions.” 

The press-work and general composition of the book are very 
Satisfactory, indeed elegant. W. W. Balzey. 


Lhe Characeae of America. T.¥F. Allen. Part II. Fascicle 2. 
Issued December, 1894. Pages 9-17, plates 8, species 9, illus- 


trated. 
In this fascicle Dr. Allen has described and figured three new 


species of Nitella, N. Bastini, N. dilatata,and N. annularis, and 
six others are figured and described, \V. capitata, N. praclonga, 
NV. clavata, N. Macounii, N. axillaris,and N. Morongit.  E. G. B. 


List of Preridophyta and Spermatophyta growing without Culti- 
vation in Northeastern North America. Prepared by a Committee 
of the Botanical Ciub, American Association for the Advancement 
of Science, 8vo, pp. 377. New York, 1893-1894. 
_ This work, originally issued in signatures to members of the 
committee and such other botanists as expressed a desire for it in 
that form, is now sent complete, simultaneously with its appear- 
ance as Vol. 5, Memoirs of the Torrey Botanical Club. 

_ It had its first definite inception at the Rochester, N. Y., meet- 


86 


ing of the American Association for the Advancement of Science 
in 1892, after the botanists there present had been engaged in an 
earnest consideration of the principles of plant nomenclature for 
many years, a subject of discussion in which most American 
botanists had taken part. An interchange of views between those 
who were present showed that while there were many differences 
of opinion in regard to details all were practically agreed as to 
certain fundamental principles. A committee was, therefore, ap- 
pointed to prepare a draft of a code, which draft was subsequently 
presented, discussed, amended and finally adopted. This is now 
known as * The Rochester Code.” 

This committee was then continued with power to prepare and 
print a list of the plants embraced within the area covered by the 
sixth edition of Gray’s Manual of Botany, with the addition of the 
States of Kansas and Nebraska, and the Canadian Provinces from 
Manitoba to Newfoundland, in accordance with the principles 
enunciated in the code. The committee immediately went to 
work and were able to present the list, in manuscript form, almost 
complete, at the next meeting of the Club, at Madison, Wis., in 
1893. 

The committee was then increased by two members, and was 
authorized to proceed with the printing of the list. Several minor 
points not determined by the Club, such as capitalization, insertion 
or omission of commas and use of trinomials, were referred to the 
committee with power, decided by vote of the members, and the 
final result is the volume now before us. 

That it will meet with the approval of over three-fourths of the 
working botanists in America, we think there can be no question. 
The Rochester Code is and has been accepted by such a large 

‘Majority that it is safe to say no work of magnitude could hope 
for future success here if based upon different principles. Some of 
the minor rules adopted in the printing, however, are not yet 5° 
generaily accepted and may be expected to remain as matters of 
discussion for some time to come. Commas may be either omitted 
or retained between the specific or varietal name and the authority; 
capital letters may or may not be used tor specific or varietal 
_ names derived from persons or places, etc., and varieties may OF 
-May not be written as trinomials. These are minor matters. 


87 


What concerns us most nearly is that, unless the citations of 
date of publication for some of the names should have been 
wrongly determined, we now have established a permanent basis 
for our nomenclature, and an amount of condensed study with- 
in our grasp, for which all botanists owe the committee a 
deep debt of gratitude. The labor of this research and of the 
verifications which were necessary only the members of the 
committee can appreciate, and we are perfectly willing to forgive, 
in advance, any errors in this respect which may and probably 
will be discovered when the list comes to be put into practical use. 
Each species and variety is given a consecutive number, beginning 
with 1. Ophioglossum vulgatum L., and terminating with 4336. 
Ptiloria tenuifolia (Torr.) Raf., the sequence being in accordance 
with Engler and Prantl’s “ Natiirliche Pflanzenfamilien.” To the 
list is added an appendix containing references to typographical 
€rrors, wrong determinations, changes in synonymy and about 
fifty additional species. 

The committee to whose labors we are indebted consists of N. ~ 
L. Britton, John M. Coulter, H. H. Rusby, Wm. A. Kellerman, 
Fredk. V. Coville, L. M. Underwood, Lester F. Ward, Edward L. 
Greene and Wm. Trelease. A. H. 


Proceedings of the Club. 
Annual MEETING, TUESDAY EvENING, JANUARY. 8, 1895. 


The President in the chair and seventeen persons present. 

Mr. Leonard Barron and Rev. A. P. Ekman were elected ac- 
tive members. 

The following officers were elected for 1895: 

President, Hon. Addison Brown; Vice-Presidents, Dr. T. F. 
Allen, Rev. L. H. Lighthipe ; Recording Secretary, Dr. Henry H. 
Rusby; Corresponding Secretary, John K. Small; Editor, Dr. N. 
L. Britton; Treasurer, Henry Ogden; Associate Editors, Dr. Emily 
L. Gregory, Anna Murray Vail, Arthur Hollick, Dr. Byron D. 
Halsted, A. A. Heller; Curator, Josephine E. Rogers; Librarian, 
Effie A. Southworth. 

Under the reading of communications, a letter was read 
from Prof. Dr. A. Cogniaux accepting the honorary member- 


88 - 


ship which had been conferred upon him, with warm expressions 
of appreciation. A communication: was then read from Mr. 
Joseph Crawford announcing the death, on December 6th, of Dr. 
J. Bernard Brinton. The Secretary remarked briefly on the life 
and character of Dr. Brinton, and announced that an obituary 
notice was being prepared which would appear in the BULLETIN. 
A committee consisting of Dr. Britton and Dr. Rusby was then 
appointed to draft appropriate resolutions, and to present copies 
of the same to the family of the deceased. 

The death was also announced, by the President, of Prof. E. H. 

Day, which occurred in Algiers on January 4th. At the same time 
he remarked upon the high character of the deceased as a scien- 
tist, a teacher and a gentleman of refinement. Similar remarks 
were also made by Mrs. Britton and by Mr. Ogden, and a com- 
mittee consisting of Mr. Ogden and Mrs. Britton was appointed 
to draft suitable resolutions and to present the same to the family 
of the deceased and to the President of the Board of Education 
of New York City. 

The paper of the evening was then presented by Mr. A. A. 
Heller, entitled “Collecting in Western Texas,” illustrated by 
specimens of the flora. 


WEDNESDAY EVENING, JANUARY 30, 1895. 


The President in the chair and thirty persons present. 
Mrs. Annie M. Smith, of 78 Orange street, Brooklyn, was 
elected an active member. . 
On behalf of the Committee appointed to draft resolutions 
concerning the death of Prof. Day, a report was Ces by Mr. 
Ogden and a copy placed on file. 
_ ‘The announced paper of the evening, on “ Food Plants of the 
American Indians,” was then read by Dr. Valery Havard, and 
“proved to be one of the most interesting and valuable communi- 
cations presented to the club in recent years. It will be published 
in a subsequent issue of the BULLETIN. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 229. 


PHYSCOMITRELLA PATENS (HEDW.) BR. & SCH. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 230. 


APHANORHEGMA SERRATA SULLIV. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 231. 


a < 
tansregeeett es 


A HYBRID OF APHANORHEGMA SERRATA. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 232. 


l 


QUERCUS PHELLOS X QUERCUS DIGITATA. 


a 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 233. 


QUERCUS GEORGIANA X QUERCUS NIGRA. — 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 234. 


io 


QUERCUS CATESBAEI X QUERCUS CINEREA.—Form A. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 235. 


QUERCUS CATESBAEI & QUERCUS CINEREA.—Form B. 


a 


89 


Index to recent Literature relating to American Botany. 


Ayres, H.B. The Muskeag Spruce. Gard. & For. 7: 504. f. 80. 
19 D. 1894. 

Barbour, E. H. Additional Notes on the new Fossil, Daimonelix, 
its Mode of Occurrence, its gross and minute Structure. Univ. 
Studies, Univ. Neb. 2: 1-16. /. r-72. 1894. 


Cross sections under the microscope show these fossils to have been plants with a 
Structure that was cellular but not vascular. 


Barnhart, J. H. Family Nomenclature. Bull. Torr. Bot. Club, 22; 
I-24. 15 Ja. 1895. 

Bastin, E. S. Laboratory Exercises in Botany. 8vo. pp. 540. jigs. 
fl. 87. Philadelphia. 1895. 

Designed for the use of colleges and other schools where botany is taught by 
laboratory methods. ' 

- Bastin, E.S. Some further Observations on the Structure of San- 
guinaria Canadensis. Am. Journ. Pharm. 67: 4-9. f. 1-5. Ja. 
1895. 

Bastin, E.S. Structure of Jrés. Am. Journ. Pharm. 67: 78-83. 
f. 4-5. Ja. 1895. 

Bessey, C. E. The Russian Thistle in Nebraska. Agric. Sci. 8: 286. 
Je-S. 1894. . 

Briquet, J. Decades Mentharum novarum. Bull. Herb. Boiss. 2: 
691~709.: D. 1894. 

Descriptions of a large number of proposed new varieties in AZentha, some of them 

American. 

Briquet, J. Sur un singulier AHypts brésilien. Bull. Herb. Boiss. 2: 
715-719. D. 1894. 

Description and discussion of Hyftis Glaziovi?. 

Britten, J. Jane Colden and the Flora of New York. Journ. Bot. 33: 
12. Ja, 1895. 

Bush, B. F. Hybrid Oaks in Western Missouri. Gard. & For. 8: 


a2. . 2% Ja. 1895. 
Carleton, M. A. Uredineae Americanae exsiccatae. Bot. Gaz. 20: 


32. 18 Ja. 1895. 
Clendenin, I. Syachytrium on Geranium Carolinianum. Bot. Gaz. 


20: 29-31. fl. g. 18 Ja. 1895. 


S, Geranii n. Sp. 


90 


Coe, C. H. The so-called Florida Sea-Beans. Gard. & For. 7: 502- 
504. f. 79. 19 D. 1894. 

Coulter, J.M. Formulae for Life histories. Bot. Gaz. 20: 31. 18 Ja. 
1895. 


Coupin, H. The Thorns of Plants. Pop. Sci. Month. 46: 498-501 
figs. ¥. 1895. (Translated from La Nature.) 


Dietel, P. Die Gattung avenelia iene Hedwigia, 33: 
367-371. 20 D. 1894. 


Describes R. Farlowiana, R. Indigoferae Tranzschel and R, Mexicana Tranz- 
schel as new species. 


Ellis and Everhart. Analytical Key to Ellis and Everhart’s North 
American Pyrenomycetes. Pamphlet, pp. 11 [1894]. 


Ellis, J. B., arid Everhart, B. M. New Species of Fungi from 
various Localities. Proc. Acad. Nat. Sci. Phila. 1894: 322-386. 30 
N. 1894 (reprint). 


Fawcett, W. Two new Orchids from Jamaica. Journ. Bot. 33: 12 
Ja. 1895. 


Descriptions of Epidendrum tridentatum and Pleurothallis uncinata. ° 


Fawcett, W., Editor. . Notes on Castleton Gardens. Bull. Bot. De- 
part. Jamaica, 1: 161-200. D. 1894. 


- Fernald, M. L. Salix balsamifera. Gard. & For. 8: 28. 16 Ja. 

1895. ; 

Greene, E.L. Mimulus luteus and some of its Allies. Journ. Bot. 33: 
4-8. Ja. 1895. 


Henderson, L.F. New Plants from Idaho. Bull. Torr. Bot. Club, 
22: 48-50. 15 Ja. 1895. , 


Describes Phacelia Idahoensis and Claytonia arenicola. 


Hollick, A. Additions to the Local Flora. Proc. Nat. Sci. Assn. 
Staten Island, 4: 55. 12 Ja. 1895. 


Ja. 1895. 


A native of Central America and Cuba. ; 


ue ‘Hooker, i ea BR Cyrtopodium virescens. Curt. Bot. Mag. 51: p/. 7396. 


Ja. 1895. 
A native of Brazil, 


ae D. iets a5 


Hooker, J. D. Acacia spadicigera. Curt. Bot. Mag. 51: pl. 7395: 


Hooker, j- D. Brycina echinata, Curt. Bot. Mag. 50: /. 7389. 


sic 


" ae sila - a. 


91 


Jenman, G.S. Adiantum Capellis-veneris Gard. Chron. 16: 690. 
8 D. 1894. 
Note on the occurrence of the species in Jamaica. 

Jenman, G.S. Asplenium Harrisi (Luasplenium). Gardn. Chron. 
17: 69. 19 Ja. 1895. 
A new species from Blue Mountain Peak, Jamaica. 

Jenman, G. S. Asplenium (Diplasium) tenebrosiun n. sp. Gardn. 
Chron. 16: 690. 8D. 1894. 
A new species from Jamaica. 

Jenman, G.S. Prris (Litobrochia) regia. Gardn. Chron. 17: 39. 
12 Ja. 1895. 
A new species from Jamaica. 

Johnson, D. S. The Crystallization of Cellulose. Bot. Gaz. 20: 
16-22. 18 Ja. 1895. 

Keffer, C. A. Black Walnut in the West. Gard. & For. 8: 12. 9 
-Ja. 1895. 

Keffer, C. A. Green Ash in the West. Gard. & For. 8: 32. 23 Ja. 
1895. 

Kennedy, G.G. Buxbaumia aphylla L. Bull. Torr. Bot. Club, 22: 
59, 5%. 15 Ja. 1895. : 

Knowlton, F. H. Vanilla and its Cultivation. Pop. Sci. News, 29: 
I, 2. figs. Portrait. Ja. 1895. . 

Kurtz, F. Die Flora des Chilcatgebietes in Siidéstlichen Alaska. 
Engler’s Bot. Jahrb. 19: 328-431. 1894. 


Ribes laxiflorum var. inermis is described as new, and a long list of species 
given with localities, 


Kurtz, F. Die Flora der Tschuktschenhalbinsel. Engler’s Bot. Jahrb. 
IQ: 431-493- 1894. 
A long list of species. f 

Meehan, T. <Aguilegia Canadensis. Meehan’s Month. §: 21, 22. 
pl. 2. F. 1895. 

Meehan, T. Darlingtonia Californica. Meehan’s Month. 5: 1, 2. 
Bl. zr. Ja. 1895. : 

Meyer, R. Die interessanten Arten der Gattung Zchinopsis. Mon- 
atssch. Kakteenk. 4: 184-186. figs. 20 D. 1894. 

Miller, J. Arthroniae et Arthothelii species Wrightii in Insula Cuba 


lectae. Bull. Herb. Boiss. 2: 725-736- D. 1894. i 
Descriptions of several species in botlr genera, with critical notes on others. 


92 


Mohr, C. Die Wilder der Alluvial Region des Mississippi in den 
Staaten Louisiana, Mississippi und Arkansas. Pharm. Rund. 13: 14, 
15. Ja. 1895. 

Taxodium distichum is the species under consideration. 


Palmer, T.C. Jsvetes saccharata. Bot. Gaz. 20: 32. 18 Ja. 1895. 


Ramirez, J. La Mocinna heterophylla. Ann. Inst. Med. Nacional, 1: 

~ 205-211. pl. 2-5. Au. 1894. 
The author is trying to prove that the genus MJocinna will stand next to Carica 

and Yacaratia of the Passifloraceae and Vascoucellia DC. 

Rich, W. P. Herbert A. Young. Bull. Torr. Bot. Club, 22: 51. 15 
Ja. 1895. 
Notes the death of this botanist. 

Rodway, J. Nature’s Triumph. Pop. Sci. Month. 46: 456-465. F. 
1895. 


Describes the reversion of neglected plantations to a state of nature. 
Rothrock, J. T. River Birch. Forest Leaves, 4: 185. D. 1894. 
With two illustrations of Betula nigra in Pennsylvania. ; 


Sargent, C.S. Quercus Texana, Gard. & For. 7: 514-516. /. 82; 
&2. 19 D. 1894. 


Sargent, C. S., Editor. A monstrous form of the Black Spruce. 
Gard. & For. 8: 44.f. 7. 30 Ja. 1895. 

Illustration of Picea Mariana. : 

Selby, A.D. The Russian Thistle in Ohio. Journ. Columbus Hort. 

Soc. g: 127-132. p/..6, 7. S. 1894. 

Small, J. K. Studies in the Botany of the Southeastern United 

3 States.—III. Bull. Torr. Bot. Club, 22: 43-48. 15 Ja. 1395. 
Describes Funcus Georgianus Coville, Rumex spiralis, Monniera crenutata 

and Coreopsis longifolia as new species. 

Smith, J. D. Undescribed Plants from Guatemala and other Cen- 
tral American Republics.—XIV. Bot. Gaz. 21: 1-11. pl. J-3- 
18 Ja. 1395. 

Describes 13 new species with plates of Cupparis Pavlleane, Cavendishia callista 
and Chaunostoma mecistandrum, the latter a new genus (Labiatae). 

Vail, A.M. A Revision of the North American Species of the Genus 
Cracca. Bull. Torr. Bot. Club, 22: 25-36. 15 Ja. 1895. 
Description and synopsis of 14 species; C. Floridana new. 

Warnstorf, C. Characteristik und Ubersicht der nord-, mittel-, und 

stidamerikanischen Torfmoose nach dem heutigen Standpunkte der 

: Sphagnologie. Hedwigia, 33: 307-337. 20D. 1894. 

____ Synopsis and brief diagnoses of 85 species of Sphagnum. 


- 


— : - 
Se 


Contributions from the Herbarium of Columbia 


College. 


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and the Adjacent Region. By H. H. Rusby (1888),. . . . . 25 cents. 
Preliminary Notes on the North American Species of the Genus 7issa, 
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Flint, Grasses and Forage Plants. Ed. 4, 1859. 50 cts. 


-*# LIST OFEK-- 


Pieridophyta and Spermatophyta of Northeastern North America 


The subscriptions to the official list prepared by a Committee of the Botanical Club, 
A. A. A. S., call for 380 copies. We have had 400 copies made and bound up at the 
alopd of $1.00 for paper binding and $1.25 for cloth binding. There remain a few 
en not yet taken. After these are sold, the list can only be had in the form of Me- 
moirs Torrey Botanical Club, Vol. V., at the regular price, or $3.00. 


EDITORS, 
Torrey Botanical Club. 


THE ANNUAL CATALOGUE 


OF THE 


Vienna Botanical Exchange Society 


Will be issued in December. It will contain about 4,000 names of spe- 
cles, forms and hybrids for exchange or sale, some of which have never 
been offered in any similar publication. The number of new species 
offered is large. It may be obtained gratis by addressing the undersigned, 


Ts. DORF CER, 


Bureringe, 7, Vienna, Austria. 


JUST OUT. 
Gray’s Forest, Field and Garden Botany, 


Revised by Bailey, $1.30. Postage, 15 cents. 


Lister’s Mycetozoa, 70 plates, all American species included, $4.00. 
Postage, 18 cents. This is the latest monograph of the Myxomy- 


cetes yet produced. , 
Everything useful to Botanists. 


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I desire to sell the European part of my herbarium (75 fasc. 
phanerogamia and _ filices), containing considerably more than 
half the European flora; all species are in good condition and 
represented by abundant specimens from various localities. 


Dr. P. TAUBERT, 
S. W. 47 York Str. 58, Berlin, Germany. 


~ CHARACEA: OF AMERICA. 


The first fascicle of the Second Part of the Characez of America is - 
now ready. It contains descriptions of eight species of Vited/a, as fol- 
lows: LVitella opaca, Ag.; obtusa, Allen; montana, Allen; Blankin- 
shipii, Allen; Missourtensis, Allen; flexilis, Ag.; subglomerata, A. Br. ; 
glomeruiifera, A. Br., with fourteen full-page illustrations (eight litho- 
graphic plates and six photogravures). These fascicles will be issued 
from time to time as plates can be prepared ; price of each part $1, the 
actual cost, if the whole edition of 500 copies be sold. -Address 


A St eee 
10 East 36th St., N. Y.- 


American Algae. Century I. 1894. 


The first hundred of a carefully prepared series of American Algae is now offered 
for sale at the uniform price of ten dollars net ($10.00). The edition of Century I. is 
limited to twenty-five. The plants have been determined with great care and have 
been compared with such standard specimens as those of Wittrock and Nordstedt, Go- 
mont, etc. Century I. comprises only freshwater forms from the Upper Mississipp! 
Valley. The set is arranged in a handsome quarto volume, with printed labels and 
abundant material for critical study. Microscopic measurements and other details are 
incorporated in the labels. The first Century contains several rare varieties, as for ex- 
ample, Sphaeroplea, a new gigantic species of 7etraspora,a new calcareous Chaeto- 
phora and others. A specially fine series of Cladophora is included, comprising -fif 
teen different species and varieties. It is believed that this, the first set of freshwater 
Algae published in America, will prove of value to herbaria that may acquire it. 


Address orders to JOSEPHINE E. TILDEN, 
- University of Minnesota, Minneapolis, Minn. 
MEMOIRS 


Department of Botany of Columbia College. 


A new series of papers under this title was commenced in January, 1895. They 
will be printed in quarto and issued as matter becomes available. Vol. I., No. 1, to be 
issued in March, will contain «A Monograph of the North American Species of the 
Genus Polygonum,” by Mr. John K. Small, consisting of about 150 pages of text and 
86 plates. The price of this number is $6.00. The Memoirs will not be sent in eX 
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This journal has been published consecutively since 1870, beginning with four 
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MONTHLY JOURNAL OF 


THE TORREY BOTANICAL CLUB. 


OFFICERS FOR 1895. 


President, 
HON. ADDISON BROWN. 


Vice Presiaents, 


T. Fi. ALLEN, M. D. L, H. LIGHTHIPE. 
: Recording Secretary, Corresponding Secretary, 
ine HENRY H. RUSBY, M. D., JOHN K. SMALL, 
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ee " PHANEROGAMIA, tie ooh aes eer ha. et oa ii CRYPTOGAMIA, 
: ‘Pow: THOS, fe PORTER, Mee ahs ELIZABETH G. BRITTON, 
N. L mee Do ; I 


| 
| 


. 


IN OF THE 


SR el ea 


BULLETIN 


TORREY BOTANICAL CLUB 


Biographical Sketch of Dr. J. Bernard Brinton. 


[Wirt PorTRAIT]. 


The botanical community of Philadelphia has met with an 
almost irreparable loss in the very sudden death, on December 
6, 1894, of the distinguished scientist, Dr. J. Bernard Brinton, 
the founder of the Philadelphia Botanical Club. 

The many expressions of sympathy and high regard which 
have been received by his family from his fellow members of the 
Academy of Natural Sciences, the Torrey Botanical Club and 
numerous other scientists with whom he was engaged in active 
correspondence, prove conclusively that his premature death 
causes a vacancy which will be difficult to fill. 

He was preéminent in more than one respect ; most emphati- 
cally, however, in the happy faculty of imparting to others his own 
enthusiasm and love for the study of the Natural Sciences. He 
was noted for the accuracy of his observations in field excursions, 
in which he was generally recognized as the leader and guide, — 

His methods were always painstaking and careful, and in his 
aim to secure choice specimens no trouble, labor nor expense 
was too great. His botanical specimens were preserved by the 
most approved methods, mounted in the most artistic style and 
identified with the most scrupulous care. His mechanical ingen- 
uity was frequently exercised in preparing specimens of fruits, 
stems and other organs, so as to advantageously exhibit their in- 
ternal structure. In his desire for absolute accuracy he frequently 


sent difficult genera to monographers for scrutiny and revision. 


: ra 2 94 


This solicitude for accuracy made his herbarium exceedingly val- 
uable for reference and comparison to his unmerous friends, who 
were in the habit of congregating in his botanical workroom. 
This place, built as an addition to his home, was familiary known 
as ‘The Den.” Here he delighted in receiving his friends and 
exhibiting his scientific collections and numerous devices for faci- 
litating botanical studies. 

He was endowed with a marvellous memory for names and 
physical characteristics. This gift frequently enabled him to 
recognize specimens which he had not met with for many years. 
It was always a matter of gratification to him to surprise his sci- 
entific visitors with the demonstration of his mechanical skill as 
an amateur cabinet-maker. He personally constructed in the 
most skillful manner, his herbarium cases, tables, stands, micro- 
scopical cabinets, etc., with a degree of perfection rarely excelled 
by expert artisans. He also prepared, with that same mechanical 
skill, all his own collecting presses, which combined the several 
features of collecting portfolio and drying press. He was a micro- 
scopist of no ordinary ability, and took considerable interest in the 
application of this instrument in the investigation of vegetable his- 
tology. His collection of minerals in microscopic crystals has, 
' perhaps, never been excelled in beauty and in the neatness dis- 
played in their mounting. His dexterity in the dissection of bo- 
tanical specimens was frequently envied by his less expert com- 
panions. While so ardently devoted to nature in her various 
manifestations, Dr. Brinton did not overlook the advantages of 
linguistic attainments. In his earlier life, much of his time was 
devoted to the study of German, in which language he conversed 
fluently. He was also proficient in Latin and French. 

Physically, Dr. Brinton seemed to embody the highest ex- 
pression of perfect manhood. His commanding presence and 
graceful bearing stamped him at once asa leader, His powerful 
frame enabled him to endure and overcome great hardship and 
fatigue. 

_ Dr. J. Bernard Brinton was born near Waynesburg, Chester 
County, Penna., August 16, 1835. His parents belonged to the 
religious Society ‘of Friends. His early education was received 
at this place and subsequently at the High School in Philadelphia, 


95 


during the short residence of the family in that city, previous to 
removal to a farm in Maryland, in 1848. He began the study of 
medicine in 1857 and matriculated at the Jefferson Medical Col- 
lege, from which school he was graduated on March 25, 18509. 

During his college course, the attention of Prof. Samuel D. 
Gross was attracted to him by the assiduity displayed in his 
studies, and furthermore by the successful management of an 
aneurism case treated by digital compression. Asa result he was 
appointed Chief of the Surgical Clinic soon after graduation. He 
lectured on Practical Anatomy at the Philadelphia School of 
Anatomy and Operative Surgery, and also conducted a Quiz on 
Materia Medica. From his graduation to the breaking out of the 
Civil War he was an active practitioner of medicine, and in 1860 
was a delegate to the American Medical Association, held in New 
Haven, Conn. 

But the fire of patriotism proved too strong for the peaceful 
tenets of his fathers, and led him early in the war to apply for the 
position of assistant surgeon in the regular army. He success- 
fully passed the rigorous examination, and his commission was 
dated April 16, 1862, signed by the President, Abraham Lincoln, 
and Edwin M. Stanton, Secretary of War. 

On September 14, 1863, he was appointed Medical Purveyor 
to the Army of the Potomac, and he retained that position to the 
close of the war. During his entire army life he continued his 
botanical studies and collection of plants. At this time it was his 
good fortune to meet another officer equally interested in the study 
of the same science, Maj. Gen. G. K. Warren. A wayside flower 
served as a means of introducing these officers, and the occa- 
sion of that meeting was a favorite reminiscence of Dr. Brinton. 
The collections he made during the Virginia campaign were cap- 
tured by the Confederate, Col. Mosby, at Belle Plain, May 12, 
1864, and burned with the supply wagons. Dr. Brinton himself 
barely escaped capture. May 13, 1865, he was brevetted Captain 
and Major for gallant and meritorious services, and on Novembet 
16th, of the same year, he resigned from the army. His services 
to the Union were marked by his usual application and devotion 
to his sense of duty; and his report at the close of his term of office 

was considered a remarkably accurate record for one handling a 
vast amount of material under such turbulent conditions. 


96 


Returning to Philadelphia, he continued in the practice of 
medicine for a few years. Desiring more leisure time for the 
study of his chosen science, he abandoned medicine and engaged 
in various manufacturing pursuits. On October 29, 1878, he was 
elected a member of the Academy of Natural Sciences, and in 
the same year he connected himself with the Botanical Section of 
that institution. He was faithful in attendance and contributed 
numerous specimens, notes and verbal communications. He was 
an indefatigable collector and made numerous excursions in Penn- | 
sylvania and neighboring States. He made a special study of the 
peculiar flora of the Pine Barrens of New Jersey, in which depart- 
ment he was recognized as an authority. He acceptably filled 
‘ numefous positions of honor and trust in the Academy of Natural 
Sciences, and at the time of his death was a member of the Board 
of Councillors. During the session of the American Association 
for the Advancement of Science, in Philadelphia, in 1884, he was 
elected a member, and he acted as guide to an excursion of visit- 
ing botanists to the pine barren region of New Jersey. 

Only the ardent lovers of nature can understand his feelings on 
that occasion, when the main object was to show, Dr. Asa Gray 
and Mr. Caruthers, President of the Linnean Society, the se- 
cluded Schizea pusilla Pursh. Nor can the joy of those gentle- 
man be expressed when‘their eyes rested on that quaint fern form 
for the first time. 

He was elected to active membership in the Torry Botanical 
Club of New York, January, 1891. Although publishing but 
little on botanical subjects, he corresponded with most of the bo- 
_tanical authorities in America and made numerous exchanges- 
Perhaps his most important labor consisted in inducing the young 
to study botany, and his greatest pleasure seemed to be in impart- 
ing to others, either in the field or in his “den,” a portion of his 
rich store of knowledge. Chiefly with this object in view, he 
founded the Philadelphia Botanical Club, in December, 1892, of 
which he was the President from its organization until the time of 
his decease. 

The fundamental aim of the Club is to study the local flora and 
prepare an herbarium representing the plants found within a 

radius of fifty miles. gee: of the members gratefully remember 


97 


the aid they have received in the study of the science from Dr 
Brinton, whom they regard as their botanical preceptor. An in- 
. timate friend, Professor F. Lamson-Scribner, has forcibly expressed 
this sentiment in the following language : 

“Those who have been with Dr. Brinton in his botanical excur- 
sions, as I have, will say with me, that in the field he was a keen | 
observer and zealous collector, observing and collecting with an 
enthusiasm which was always contagious to his party.” 

« The results of these trips, which we enjoyed to the utmost, have 
enriched the herbaria of many scientific institutions and those of 
botanists in all parts of our country. His work and his conversa- 
tion rarely failed to excite the ambition of others, and I am confi- 
dent that many young men have received inspirations from their 
associations with Dr. Brinton, leading them to become better 
botanists or more earnest students.” 

Dr. Brinton was married on November 13, 1862, to Sallie W. 
Clemens, of Philadelphia. A married daughter and two sons sur- 
vive him. As a source of consolation, after the death of his wife, 
he engaged more earnestly in botanical studies. It is a peculiar 
coincidence that he had expressed an intention of retiring from 
the Presidency of the Botanical Club, and at the time of his de- 
cease had in preparation a farewell address. His last evening was 
spent at the home of a lifelong friend, and the following verses 
were discussed, which now seem to have fittingly foreshadowed 
the final sleep that should soon come to one of the participants: 


* Oever de stillen Straten, 
Geit klar de Glockenslag, 
God’ Nacht! Din Hart will slapen ; 
Un’ Morgen is ook een Dag. 


Noch eenmal lat uns spriken ; 
Goden Abend, gode Nacht! 
De Maand schient up de Daken 

Uns Herrgott, halt de Wacht,” 


A. W. MILLER, M. D., 

Gro. M. BERINGER, 

Jos. CRAWFORD, 
Committee. 


98 


Food Plants of the North American Indians. 


By Dr. V. HAVARD, U. S. ARMY. 


The maxim that “Necessity knows no law” is well exemplified 
in the diet of the North American Indians who, when driven by 
stress of hunger, eat whatever the animal and vegetable kingdoms 
bring within reach, so that it may be truly said of some tribes that 
they reject nothing which their teeth can chew or their stomachs 
digest, however tasteless, unclean and repulsive. 

A review, therefore, of all the Indian food plants would include 
hundreds of species and be as tedious as unprofitable. I shall 
confine myself, in this paper, to the most important; those formerly, 
or yet, habitually used. 

The subject naturally divides itself into two heads: Ist, plants 
- cultivated; 2d, plants growing wild; and of the cultivated plants 
we may consider separately those introduced and those which are 
indigenous. 

At the time of the discovery of America many tribes had 
already emerged from a wild nomadic life and, although still 
largely depending upon game and fish, were entering upon a par- 
tial sedentary agricultural state. So far as a chronic state of war- 
fare would permit, land was set apart for farming purposes and 
upon it was almost invariably planted the triad of vegetables: 
maize, pumpkin or squash, and beans. This primitive agriculture 
was mostly pursued on and east of the Mississippi; in the arid in- 
terior comparatively small areas were occupied by agricultural 
tribes, and these dwelt chiefly in New Mexico. and Arizona, and 
along the Missouri, Platte and Arkansas rivers. 

The vegetables just mentioned were introduced from the south, 
being indigenous to Mexico or South America where a compara- 
tive state of civilization had fostered their evolution, and soon 
_ found their way to the St. Lawrence river and from the Atlantic 
to the Rio Colorado of the West. Cartier found them at Mon- 
treal in 1535, Champlain among the Five Nations in 1603, 
Hudson along the river bearing his name in 1609, the English at 
Jamestown in 1607, De Soto in the Gulf States in 1539, Mar- 
_ quette, Hennepin and La Salle in the Mississippi States, Cabe¢a 


99 


de Vaca among the Pueblo Indians in 1528, and Alarcon on the 
Colorado in 1542. 

The Pacific Coast tribes did not cultivate the soil and depended 
entirely on the spontaneous products of land and water. It is re- 
markable that these tribes, although having more or less inter- 
course with the Missouri Indians on the east and the Pueblo Indians 
on the south, and always on the verge of starvation, failed to 
introduce and cultivate the maize or other vegetables. Lewis and 
Clarke in their memorable expedition across the continent, from 
the Missouri to the mouth of the Columbia, in 1804 and 1805, 
found plenty of maize and beans at the Mandan villages, about 60 
miles above the present town of Bismarck, N. D., but none north 
or west of that point. 

Maize (Zea Mays L.) is probably indigenous to the southern 
highlands of Mexico. Under the civilization of the Mayas, Na- 
huas and Aztecs this precious plant received special care and 
many varieties were developed. Humboldt states that the Aztecs 
began to cultivate it in 666 A. D. It rapidly spread northward 
and, it is estimated, in the year 1000 had already reached the 
eastern States. At the time of the landing of Columbus it was 
the principal and most common crop of all agricultural Indians, as 
itsoon became that of the white settlers. According to E. L. 
Sturtevant, one of our best authorities on the history of garden 
vegetables, the various agricultural species of maize: the flints, 
dents, softs, sweets and pops appear to be aboriginal forms, and 
we have no evidence that any new varieties have resulted from 
our three centuries or more of civilized cultivation. The very 
hardy form grown in Maine, and which must ripen its grain in 
three months from the day of planting, is certainly of Indian de- 
velopment, and so are the peculiar red and blue varieties culti- 
vated in New Mexico as well as on the upper Missouri. 

The common Kidney Bean (Phaseolus vulgaris Savi) is a South 
American plant; its culture under many varieties spread rapidly 
_ through Mexico and the States to Canada and was, next to Maize, 
the most important crop of farming Indians. The finding of seeds 
of this species by Professor Witmack in the prehistoric graves of 
Arizona, not only completed the demonstration of its American 
origin but likewise proved the antiquity of its culture in our own 


100 


country. It is also probable that the Lima Bean (P. /unatus L.), 
which is of Brazilian origin, was used by our Southern Indians. 

The third genus of introduced plants cultivated in pre-Colum- 
bian time is Cucurbita. The Common Pumpkin (C. Pepo L.) as 
well as the Squash (C. maxima Duch.) are natives of South or 
Central America and were grown coextensively with the preced- 
ing by North American Indians some of whose names, as ap- 
plied to certain varieties, have endured to this day. The China or 
Barbary Squash (C. moschata Duch.) was also thought by Asa 
Gray to be of American origin and, if so, was doubtless known 
to our Southern Indians. 

We come next to the consideration of native plants cultivated 
by the Indians. It may at once be stated that our gardens and 
orchards, unless we except the Jerusalem Artichoke and the Sun- 
flower, do not contain a single plant developed by the Indians 
from any of our wild species. In exculpation we may say that, so 
far as fruits were concerned, an abundant supply could always be 
relied upon, while as regards roots, tubers or bulbs, it seems 
very doubtful whether there are more than two or three sus- 
ceptible of marked improvement. It is only during the last 
49 or 50 years that we have ourselves mostly evolved our 
many orchard forms of native fruits, while we have not yet 
produced a single vegetable. The Indians certainly exhib- 
ited commendable promptness and industry, after the ad- 
vent of the whites, in introducing such fruits as were shown 
to be desirable. Thus the Peach brought into Mexico soon after 
the conquest was, according to the testimony of Du Pratz, found 
in general cultivation among the Indians of Louisiana when the 
French settled that province in 1698, and had become abundant 
in Georgia at the time of the settlement of the English in 1732. 
Wm. Bartram describes the carefully planted Orange groves of the 
Indians which he noticed in 1773. The early introduction and 
propagation of these two plants by the Indians led to the erroneous 
impression that they were of American origin. 

Of the native plants cultivated by Indians we have no very 
clear or precise information, but I believe the following list in- 
cludes most, if not all of them. The first place belongs to the 
Jerusalem Artichoke (Helianthus tuberosus L.). It produces many 


101 


edible tubers, sometimes 2 inches in diameter, in our day mostly 
used for the feeding of cattle, horses and pigs, but which were 
precious to the Indians on account of their hardiness and prolifi- 
cacy, retaining possession of the soil for many years. These 
tubers were mentioned by Champlain. in 1603 and brought to 
France by Lescarbot who, in 1612, describes them as being “as 
big as small turnips, excellent to eat, with the taste of artichoke 
but more agreeable, and multiplying in a wonderful way.” As 
the plant is native of the valleys of the Ohio and Mississippi and 
does not reach any part of Canada, it is evident that the Canadian 
and New England Indians who planted it must have obtained it 
from the tribes further south and west, so that we may infer a 
rather large area of cultivation. The Jerusalem Artichoke is, so 
far, the only contribution of North America, exclusive of Mexico, 
to the vegetable garden of the world, and it can be said to be an 
aboriginal contribution. Strange to note, it is now much more 
cultivated in the Old World than on this continent. 

The allied species H. doronicoides Lam., found from Ohio to 
Missouri, and for a while erroneously thought to be the original 
of the Jerusalem Artichoke, has similar tubers although of coarser 
texture; it is quite probable it was also cultivated. The “ Indian 
Potato” of the Assiniboine tribe, mentioned by Bourgeau, is the 
tuber of a formof H. giganteus. \., which ranges. from Minnesota 
to the Saskatchawan river: 

Apios Apios (L.) MacM. (A. tuberosa Mcench), the Ground-Nut, 
has edible round or pyriform tubers strung upon long subterranean 
shoots, varying in size from a nut to a hen’s egg; they contain a 
large proportion of starch and have, when cooked, somewhat the 
taste of potatoes, being wholesome and nutritious. Rafinesque 
tells us that this plant was formerly cultivated by the Indians, and 
still by the Creeks at the time of his writing, not only for the tubers 
but also for the seeds which, he thinks, are as good as peas. It 
must have been extensively used by the eastern and southern In- 
dians as all early explorers mention it and some were saved by it 
from starvation. The tubers were the “Penacs” of the ae 
the “Hopnis” of the eastern tribes, and apparently the “Openauk 
of Hariot, seen in Virginia in 1584 and described as “a kind of 
round root, some as big as walnuts, some far greater, found in 


102 


moist and marshy grounds, growing many together in ropes as 
_ though they were fastened with a string.’ Asa Gray said that 
had civilization started in America instead of Asia, “ our Ground- 
Nut would have been the first developed esculent tuber and would 
probably have held its place in the first rank along with potatoes. 
and sweet potatoes of later acquisition.” In this connection it 
should be said that native cultivation does not appear to have had 
any effect upon the size and quality of this tuber, and that experi- 
“ments by Vilmorin and others with a view to its improvement 
have not been successful, although hardly continued long enough 
to be conclusive. The tuber is of slow growth, requiring two or 
three years before reaching sufficient size to be useful, and its 
creeping, scattering habit renders the harvest laborious. 

Neiumbo lutea Pers., the Yellow Nelumbo or Water Chiquapin, 
is the finest of our water-lilies, occurring from the Great Lakes to 
the Gulf, and westward to Minnesota and Nebraska. In the 
northern Atlantic States it has only been found in the Delaware 
river below Philadelphia, in Swartswood Lake, N. J., and in the 
Connecticut valley, so far out of its range as to lead to the gen- 
eral supposition that the Indians brought and naturalized it in 
these local habitats. Although difficult of propagation the plant 
was certainly worthy of the effort. The spindle-shaped, often 
angled and furrowed tubers are 5 to 10 inches long and weigh 
from 2 to 8 ounces ; when baked they have, says Dr. G. Engel- 

mann, a pleasant, sweet and mealy taste much resembling that of 
the sweet potato, without anything reminding one of their growth 
in stagnant water. The boiled or baked seeds have the taste of 
chestnut and are highly nutritious, while the petioles and young 
leaves may be eaten“as~spinage. 

Orontium aquaticum L., or Golden Club, the « Tawkee” or 
“ Tawkin” of the Indians, is also an aquatic perennial with deep, 
bulbous rootstock and large pea-like seeds. Both roots and seeds 
were much prized by the Indians and, according to Peter Kalm, 
by some of the white colonists as well; the seeds specially being, 
after sufficient cooking, quite palatable. This plant grows in 
ponds along the coast from Massachusetts to Florida. It has also 
been found in isolated places further inland, near the site of Indian 
villages, where it is supposed to have been propagated by the 

natives; but this is a mere hypothesis. — 


103 


The following native plants were cultivated for their fruit. 

The Common Sunflower (#7. annuus L.) is an American plant 
which, under several forms, extends from the ‘arctic circle to the 
tropics and from the Missouri river to the Pacific. It was com- 
monly cultivated by the Indians, from Canada to Mexico, west 
and east of the Mississippi, being for many of them a staple article 
of food. The seeds were parched, ground and made-into a pala- 
table and nutritious bread said by Palmer and Douglas to be 
hardly inferior to corn bread. By expression they yield about 20 
per cent. of an excellent table oil which the Indians, more mindful 
of their appearance than of their diet, mostly used for anointing 
their hair and skin. The culture of this plant in Canada was 
noticed by Champlain and, a few years after, by Sagard; in that 
region the seeds must have been obtained from beyond the Missis- 
sippi and a little south, through the agency of intervening tribes. 
As in the case of H. svéerosus it is interesting to note that this 
native plant is much more extensively cultivated in the Old World 
than in the New. 

Prunus Americana Marsh, and P. nigra Ait., our two species 
of Wild Yellow or Red Plum, were, according to several authori- 
ties, planted by the New England and Canadian natives, and from 
the many forms discovered farther west it is not improbable that 
this culture extended to the Mississippi. Some forty-five horticul- 
tural kinds derived from them are described by Prof. Bailey, and it 
is not assuming too much to suppose that several of them are due 
to variations initiated by Indian industry. It is probable enough, 
however, that the native orchard was seldom regularly planted, 
but oftener the accidental result of seeds dropped in the vicinity 
of camping grounds and villages. 

Prunus angustifolia Marsh, the Chickasaw Plum of the South, 
is regarded by Prof. Sargent as native of the eastern slopes of the 
southern Rocky Mountains and of the plateaus extending thence 
to the Mississippi, and as having been introduced by the Indians 
into the southern Atlantic States where it soon became exten- 
sively naturalized. Clumps of it were found about all Indian vil- 
lages, and the variations thus started have doubtless developed 
into some of our seventeen horticultural forms. 

Of the cultivation of Prunus hortulana Bailey, the common 


104 


native species of the middle Mississippi region, we have no direct 
evidence; but as Plums were planted north and south of that 
region it is quite probable.that.the differentiation which has re- 
sulted in giving ‘us some twenty-six horticultural forms is also of 
Indian origin. 

At least one species of grape was cultivated for its fruit. V2tes 
Arizonica Eng. has been found growing in rows near Fort Whipple, 
Ariz., which may be accepted as conclusive evidence of its culture 
by the Pueblo Indians. It was unfortunate that the Grape thus 
selected should be described by Munson, the eminent viticulturist, 
as the least promising of all the species for improvement in fruit. 
Considering the ease with which Grapes are propagated, by seeds, 
cuttings or layers, it is likely enough that other species were 
grown, specially V. Laérusca, our northern Fox-Grape, although 
there appears to be no proof of it. 

It seems quite probable that the Pecan and one or more Hick- 
ories were more or less planted, as the abundance of nuts is often 
alluded to by the first explorers. Wm. Bartram, in the account 
of his travels through the South in 1773, mentions an Indian 
plantation of Shellbark Hickory. There is likewise some ground 
for the belief that our native Red Mulberry was cultivated, the fine 
quality and great quantity of the fruit being mentioned by De Soto 
and others. 

The last of this group of food plants, so far as I know, is Pass?- 
flora incarnata L.., the Maypop of the Southern States, which was 
cultivated by the Indians of Virginia. Says Captain John Smith: 
« They plant also maracocks, a wild fruit like a lemon, which in- 
creases infinitely’; elsewhere he refers to it as ‘‘pleasant”’ and 
“wholesome.” The fact is that it contains but little nutriment 
and does not appear susceptible of improvement. The plant itself, 
on account of its spreading perennial and deep roots, is a noxious 
weed, 

For the sake of completing the list of plants cultivated by abor- 
igines I may be allowed to mention one or two which, although 
not food plants, were propagated with full as much industry and 
success as the Maize or the Sunflower ; I mean species of Tobacco 
(Nicotiana). The Common Tobacco (NV. Zabacum L.), of South 

America, was not known to our Indians except perhaps as var. 


105 


undulata Sendtner, the Yaqui Tobacco grown by some of the 
natives of Arizona and California. The Tobacco cultivated by our 
Southern and Eastern Indians was the Mexican species VV. rustica 
L. now naturalized and occasionally found as a weed, while on the 
Missouri and farther west the native V. guadrivalvis Pursh was 
mostly used. The var. mu/tivalvis Gray of the latter, a form de- 
rived from cultivation, or perhaps a distinct species, has been used 
by the Pacific Indians from time immemorial and is said, by 
Douglas, to be the only vegetable which the natives of the Co- 
lumbia cultivated. 

I shall now proceed to enumerate the native plants which in 
their wild or uncultivated state furnish food to the Indians. 


Roots, TUBERS AND BuLss. 

Sagittaria latifolia Willd. (S. variabilis Eng.), the Common 
Arrow-head, is an extremely variable species, extending from the 
Atlantic to the Pacific and from British America to Mexico. It 
has tuberous oblong roots the size of a hen’s egg, but sometimes 
as large as a man’s fist, which either boiled or roasted were much 
consumed by the natives all over the land. It was the “ Katnis’” 
of the eastern tribes, the “ Wabesipinig” or Swan Potato of the 
Chippeways in Minnesota, and the “ Wappatoo” of the Columbia 
river Indians. It appears to have been, next to Camas, the most 
useful root of the Pacific slope natives; its name recurs almost on 
every page of certain chapters of Lewis and Clarke’s narrative from 
which I quote the following: “The most important spot is Wap- 
patoo Island, a large tract lying between the Multnomah and an 
arm of the Columbia. The chief wealth of this island is found in 
the numerous ponds of the interior which abound with the com- 
mon Arrow-head. The bulb, to which the Indians give the name 
of Wappatoo, is their great article of food and almost the staple 
article of commerce on the Columbia. It is never out of season, 
so that at all times of the year the valley is frequented by the 
neighboring Indians who come to gather it. It is collected 
by the women ; each takes a light canoe into a pond where the 
water is as high as the breast, and by means of her toes separates 
from the root the bulb which, on being freed from the mud, rises 
immediately to the surface of the water and is thrown into the 


106 


canoe. In this manner these patient females will remain in the 
water for several hours, even in the depth of winter.” 

A form with very large smooth leaves, about as wide as long, 
from Central to Southern California, thought by Parish to be the 
introduced S. Sinensis Sims, is cultivated by the Chinese in Cali- 
fornia for its tubers. 

We next come to the roots furnished by the Arum Family 
(AraAcgakE). All the plants of this order are impregnated with an 
intensely acrid and pungent principle. The pangs of hunger 
must indeed have been keen which drove the natives to experi- 
ment with them, but the happy discovery was made that drying 
and cooking dissipated this noxious acridity and that the roots 
contained a large proportion of nutritious starchy food. Avzsaema 
triphyllum (L.) Torr., the Indian Turnip of our woods, has a bulbous 
rounded or flattened root, I to 2 inches in diameter, which, ac- 
cording to G. H. Harris, is the “« Hopnis”’ of the Seneca Indians 
and once their habitual bread-root. Peltandra Virginica (L.) Kunth, 
the Arrow Arum, has a short, very thick, deep-seated rootstock, 
sometimes 6 inches in diameter and weighing 5 or 6 tbs., the 
«“ Tawho,” “ Tuckah” or “ Tuckaho”’ of the natives, and, accord- 
ing to Capt. J. Smith, the root chiefly used for food by the Vir- 
ginia Indians. “In one day,” says Peter Kalm, “a savage will 
gather enough for a week. . Unless carefully roasted it will 
prickle the throat extremely, but he so manages it in case of ne- 
cessity as to make bread of it.” He adds that hogs are very 
greedy of the roots and grow fat upon them. Bartram told him 
that the savages also boiled the spadix with the berries and de- 
voured them as a great dainty. According to Rafinesque the 
seeds may be used as a substitute for pepper. The other species, 
P. sagittaefolia (Michx.) Morong, has somewhat similar roots. 

The roots of Calla palustris or Water Arum, Orontium aquati- 
cum ot Golden Club, and even of Spathyema foetida (L.) Raf., our 
Polecat Weed, were also used but to a lesser extent. Colocasia 
antiquorum Schott and C. esculenta Schott were introduced into 
the Southern States at a very early date, but probably not before 
the advent of the whites. 

_ Imay mention here, not as a root, but as a root growth, the 
true “ Tuckahoe” or “Indian Bread” of the Southern States, 2 


107 


fungus, Pachyma cocos Fries (Lycoperdon solidum), springing from 
the roots of trees in sandy soil, of about the size of a man’s head, 
with scaly woody bark and white, homogeneous granular interior. 
It is, like other fungi, destitute of starch but contains about 80 
per cent. of carbo-hydrates almost entirely in the shape of insipid 
pectin whose nutritive value is still undecided but probably not 
' great. 

On the low grounds of Southern Florida grows a beautiful plant 
of the Cycapacar, Zamia integrifolia Willd., the « Coontie” of the 
Indians, which contains in the caudex and roots about 65 per 
cent. of an excellent starch sometimes called Florida arrow-root. 
This plant supplied the Seminoles with food during their long 
wars with the United States and is now more or less cultivated. 

Certain species of Claytonia, which we cultivate in our gardens 
for their delicate beauty, grow from deep edible bulbs whose crisp 
flesh and nutty flavor were much prized by the natives. I may 
mention C. l2rginica and C. Caroliniana of the eastern States and 
€. Caroliniana sessilifolia Torr. of the Rocky Mountains and west- 
ward. C. megarrhiza Parry has a large fleshy tap-root, but it is 
confined to the summits of the Rocky Mountains and seldom 
available. C. perfoliata Don, of the Pacific States, where it is 
known as “Spanish Lettuce,” is eaten as salad and cooked as 
greens. 

Another elegant little plant of the same Purslane Family is 
Talinum aurantiacum Eng., found everywhere in Texas west of 
the Pecos river and whose tuberous root is quite edible when 
cooked. 

If we now leave the eastern States and extend our investigation 
westward, beyond the Mississippi, the first plant deserving of at- 
tention is Psoralea esculenta Pursh, the “« Pomme de Prairie” or 
“ Pomme Blanche” of the Canadian explorers and voyageurs, the 
« Prairie Turnip” or “Prairie Potato” of the Americans, the 
“ Tipsinah ” of the Sioux and the “ Tahgu” of the Osage Indians. 
It is widely distributed, being most abundant on the dry table- 
lands of the Missouri region, from Montana through Dakota and 
Nebraska to Kansas. The tuber is irregularly elliptical or conical 
in shape, ranging in size from a large filbert to a hen’s egg, aver- 
aging 114 to 2 inches in length. On section it is seen to be com- 


cagriellew 


108 


. 


posed of a white granular substance easily pulverized into a light — 
starchy flour. Raw it has a very palatable farinaceous flavor 
entirely devoid of bitterness; cut into slices and dried it is easily 
kept and may be found to this day in all the tents of the Sioux 
Indians for whom it has always been a staple food. They gener- 
ally eat it cooked, and as they appreciate the advantages of a mixed 
pot-au-feu, boil it with tripe, fattened pup or venison. According 
to an analysis by Mr. Clifford Richardson (from material I sent 
him) the tuber contains 70 per cent. of starch, about 5 of a sugar 
new to chemistry, and 9 of nitrogenous matter under several 
forms. No serious and prolonged attempt has as yet been made to 
- develop this tuber by cultivation and selection; such attempt is 
certainly desirable and, if successful, would give us a toothsome, 
wholesome and nutritious vegetable, remarkable for its large pro- 
portion of starch and nitrogen, and peculiarly adapted to our arid 
regions. 

Five other species have, likewise, tuberous edible roots: P. 
hypogaea Nutt., a smaller plant than the preceding, with a some- 
what more southern range and probably often confounded and 
collected with it by the Indians; P. Californica Wats. (P. mephitica 
Wats.) and the allied, if distinct, P. castorea Wats., ranging from 
Colorado to California, and affording food to the Pah-Utes; P. 
canescens Michx. of the Southern States, remarkable for its very 
long subterranean stem ending with a large conical tuber, evi- 
dently too deeply buried to have been noticed by the natives; P. 
subacaulis T, & G., a rare and local- Tennessee species. 

Another member of the Leguminous family is our Wild Liquor- 
ice (Glycyrrhiza lepidota Nutt.), mostly noteworthy as a very bad 
weed in the west, everywhere too prevalent. The long slender 
-thizome has a faint sweetish taste of liquorice, and the Indians 
sometimes chew it for its supposed tonic and expectorant effects. 
Much more valuable to the natives is the Chenook. Liquorice, a 

Lupine (Lupinus littoralis Dougl.), of the coast of Oregon and 

_ Washington, with long tapering granular roots which, when roasted, 

yield a pleasant farinaceous food. 

- Our native Umbellifers yield a certain number of useful edible. 
_ roots. To my taste and within my experience, the most delicately 

_ flavored (in the raw state) is that of Carum Gairdneri B. & H., the 


109 


“Yamp” or “ Yampah” of the west, extending from the Rocky 
Mountains to California and Britis‘: Columbia, being particularly 
abundant on the Little Snake, or Yampah,river. The fusiform ‘or 
conical roots are in close clusters of 2-5, about an inch long anda 
half-inch thick, consisting, within the blackish skin, of a pure white, 
farinaceous substance. They have a delicious sweet nutty 
aromatic flavor, without any bitterness or astringency. Raw or 
cooked, they have always been highly valued by Indians, specially 
the Shoshones and tribes of the Snake river and tributaries, as well 
as by all explorers and settlers. Bears and pigs are very fond of 
them, often tearing up large tracts of ground in their search. 
Chemically, they are remarkable for the large proportion of sugar - 
and albuminoid matter they contain. There is little doubt that 
if susceptible of enlargement by culture, the Yamp would soon 
become a favorite in the vegetable garden. : 

Two other species, C. Kelloggit Gray, the “ Wild Anise” of 
Central California, and C. Oregonum Wats., have similar roots. 
# Probably still more important to the Indians, among the Um- 
bellifers, on account of its wide distribution and abundance, is the 
genus Peucedanum of which we have some 46 rather closely re- 
lated species. Ten, ranging from the Rocky Mountains to the 
northern Pacific Coast, have tuberous edible roots more or less 
used by the natives. The tubers are seldom. over an inch in 
diameter and almost entirely made up of an homogeneous white 
starchy substance, palatable when raw, with a faint celery-like 
flavor, but generally roasted or baked and pounded into flour 


from which nutritious and wholesome bread or cake is made. ~~ 


The best tuber is probably that of P. Candyi C. & R., the “ Chuk- 
lusa” of the Spokane Indians. It contains, according to an 
analysis by Prof. H. Trimble, 17 per cent. of starch, 11 of saccha- 
rose, 3 or 4 of albuminoids and 15 or 16 of mucilage. The tuber 
of P. eurycarpum C. & R., the “ Skelaps” of the Spokane Indians, 
has also been examined by Prof. Trimble, who found 35 per cent. 
of starch, about 10 of albuminoids, only 2 of saccharose and 3 or | 
4 of mucilage, a composition surprisingly different from that oF 
the preceding. The bulb of P. farinosum Geyer, the “ Tuhwha 
of the Spokanes, is equally good ; to it, as well as to other species, 
the name « Biscuit-root” has been given by the whites. 


110 


Other tuberous species worthy of mention are P. Geyert Wats., 
P. ambiguum T. & G., and P. Cous Wats., the Indian designation 
« Kous”’ applying likewise to other species. 

The allied genus Cymopterus contains also species with thick 
elongated or tuberous, more or less edible roots, much prized by 
the Indians of the Rocky Mountain region and the Great Basin. 
The best known, as esculents, are C. globosus Wats., C. glomeratus 
Raf., C. montanus T.& G. The latter has a fusiform parsnip-like 
root the size of a man’s finger, of a very agreeable taste, and a 
good vegetable in early spring when soft and tender. C. Fendlert 
Gray, the “ Chimaja” of the Mexicans, is impregnated with a 
pleasant anisate volatile oil, and used to flavor meats and make 
bitters in New Mexico. C. purpureus Wats. is used as a pot-herb 
to season soup and mush by Navajo Indians. 

The immense family of the Composirak presents very few plants 
for our consideration; the most noteworthy belong to the genus 
Cnicus or Thistle. Lewis and Clarke describe a Thistle on the 
Lower Columbia called “Shanataque” by the natives, with fusiform 
root a foot long and about the size of a man’s thumb which when 
baked becomes very palatable and “the sweetest vegetable eaten 
by the Indians.’’ Mr. Thomas Howell thinks the plant must be 
Cnicus edulis Gray, although the specific name of the latter refers 
only to the esculent*qualities of the young stems which are fleshy 
and tender; the value of the root appears to have been overlooked. 
Fremont speaks of another Thistle, not determined, found on the 
middle and lower part of Bear river (in Idaho and Utah), with 
root the size of a carrot and very agreeably flavored. I may also 
mention C. Drummondi Gray, whose edible roots are stated to 
have saved from starvation a lost explorer of the Yellowstone 
Park in 1870. 

The genus Balsamorrhiza Hook., of the Rocky Mountains and 
the Pacific States is characterized by thick and deep roots which 
exude a terebinthine balsam. They are edible, raw or cooked, 
after peeling off the thick bark which contains most of the bal- 
samic oleo-resin. Not only the roots but also the young stalks 
and the seeds were used for food by the Indians. 

The same remarks apply to the allied genus Wyethia Nutt., 
which has likewise fleshy roots and large albuminous seeds. 


Ill 


The last edible Composite to be mentioned is Microseris nutans 
Gray, with succulent almost transparent roots full of a bitterish 
milky juice, which are eaten raw by the Nez Percés Indians. 

The PortuLacaceak contain one species noted from the earii- 
est days as an important Indian food plant, Lewssta rediviva 
Pursh, the « Spatlum ” and « Chitah” of the natives, the “ Bitter- 
root’ of the whites, ranging from the Rocky Mountains to the 
Pacific, and giving its name to the Bitter-root mountains of Mon- 
tana. It is a handsome little plant and very remarkable for its 
wonderful vitality, having been revived and successfully planted 
after being kept months in herbarium. The roots, 3 or 4 or more, 
are curled and twisted and seldom thicker than a goose quill; their 
brownish-red bark is intensely bitter while the inner white and 
farinaceous part is quite palatable and nutritious, although having 
in the raw state a slight bitterish flavor. Boiled or otherwise 
cooked they are excellent food, extensively used by the Indians. 
An analysis by Prof. Trimble yielded about 15 per cent. of gum 
and mucilage, 3 or 4 of albuminoids and 8 or 9 of starch; no evi- 
dence of sugar could be obtained. The statement once made by 
a too credulous observer, and since often repeated by authors that 
this root “abounds in concentrated nutriment,” is certainly very 
much exaggerated. The other species, L. drachycalyx Eng. has 
similar roots. 

In the Matvacear we find but one genus with fleshy edible 
roots, Callirrhoe Nutt. One of the handsomest species, C. digitata 
Nutt. of the southern plains, has a fusiform root, in shape and size 
between a small turnip and a parsnip, said to be even more pleas- 
ant tasted than that of Psoralea and highly prized by the natives. 
C. pedata Gray, of the Texas prairies, and ‘other species have 
similar farinaceous roots. 

The showy genus Amoreuxia, of the BrxiNeag, is represented — 
in our country by A. Wrightii Gray, of southwest Texas and A. pal- 
matifida DC. (A. Schiedeana Planch.) of Arizona and Mexico. The 
roots of the former are greedily devoured by peccaries and other 
animals; those of the latter when roasted have the taste of : the 
Parsnip and carrot, and are eaten by the Papago and Pimo Indians 


as well as by the Mexicans under the name of “ Sayas.” 
- Turning to the mountains of New Mexico, Arizona and Texas, 


112 


we find two Potatoes, Solanum tuberosum boreale Gray and S. 
Jamesu Torr., whose tubers have always been among the chief 
articles of food of the Navajo Indians. The var. doreale is so far 
removed and cut off from its Andean type, by the absence of in- 
termediate Mexican and Central American forms, as to make their 
close relationship a very interesting fact in geographic botany. 
Its tubers differ only from those of the Common Potato by their 
smaller size, being from half to three-quarters of an inch in diam- 
eter, and by a peculiar aroma. The tubers of S. /amesii are still 
smaller, dark-colored and usually covered with minute tubercles. 
These plants are hardy and of easy growth, readily responding to 
cultivation, so that it seems strange that the Navajo or Pueblo 
Indians should not have planted and developed them. The tubers 
of var. doreale are readily improved in size, as shown by Prof. 
~ Bailey who has obtained some 4 inches long still retaining their 
distinguishing aroma. This variety of the Common Potato is be- 
yond a doubt the most promising native food plant of North 
America outside of Mexico, and was the one most worthy of the 
_ attention of the aborigines. 

In about the same region and extending north into British 
America is found the “ Kooyah” or “ Tobacco-root ” of the French 
(Valeriana edulis Nutt.), with large fusiform perpendicular root- 
stock dividing below into deep and thick branches. This root, 
black outside and bright yellow inside, is remarkable when fresh 
for its very repulsive odor and taste, resembling those of chewing 
tobacco, but when thoroughly cooked is sweet, palatable and nu- 
tritious. This plant is of great importance to the Indians of the 
Great Basin andthe Northwest. Fremont, on reaching Bear river, 
in northern Utah, wrote: “I ate here for the first time the 
Kooyah, the principal edible root among the Indians who inhabit 
the upper waters of the streams on the western side of the Rocky 
Mountains.” I doubt whether this plant is worth cultivating for 
its root, but its medicinal properties. might be profitably in- 
vestigated. 

The Lily Family (Littaceag) contributes quite generously to 
the vegetable diet of the Indians, as I shall now proceed to show. 
| Taking everything into account, abundance, size, taste and nu- 

_ tritiousness, the best of all native bulbs is doubtless that of the 


113 


Camas or Quamash (Camassia esculenta Lindl.),a showy plant 
ranging from the Rocky Mountains to California and British 
Columbia, sometimes so abundant on rich meadows as to tint 
them a uniform light blue color, suggesting, as expressed by an 
early explorer, “a lake of clear water.” The bulb is globular- 
ovoid, resembling a small onion; raw it has.a mucilaginous, rather 
insipid taste, but baked it acquires the flavor as well as the color 
of chestnut. Lewis and Clarke were probably the first white men 
to eat it, as told in their narrative: “ The Indians set before us a 
small piece of buffalo-meat, some dried salmon, berries and several ° 
kinds of roots. Among these last is one which is round and much 
like an onion in appearance and sweet to the taste; it is called 
Quamash and is eaten either in its natural state or boiled into a 
kind of soup or made into a cake called « pasheco.” After our 
long abstinence this was a sumptuous treat.” 

Camas has always been one of the chief articles of subsistence 
of all Indians in and west of the Rocky Mountains. They usually 
bake it in heated pits, sometimes mixed with a black hair-like 
lichen (Alectoria jubata) which grows in profusion on Larzx occt- 
dentalis, the result being a dark brown homogeneous soft mass 
which is fashioned into large cakes. 

The other four species of Camassia described, all closely inter- 
related, have the same kind of edible bulb, with the exception of 
C. Cusickii Wats., the largest and finest, whose bulb is said, perhaps 
without sufficient investigation, to be nauseous and very acrid. 

The bulbs of all the species of Adium, or Garlic, are more or 
less edible and nutritious in spite of the strong-scented volatile 
oil they contain; many references are made to the ‘Wild Leekes” 
and “Wild Onions” by the first explorers who were sometimes 
compelled to follow the example of the Indians and eat them to 
Sustain life; however, it was their abundance all over the land 
which gave them value rather than their quality. 

Several species of Siilax have thick knobby tuberous root- 
Stocks, which were utilized by our southern Indians. The most 
noted is S. Pseudo-China L., with extensively spreading and fas- 
Cicled roots containing a large proportion of starch readily obtained 
as a reddish sediment by washing in water, and formerly, accord- 
ing to Bartram, made into soup, bread or jelly. Later these roots 


114 


were used by the white settlers, mixed with molasses and parched 
corn or rice, to make a very wholesome and palatable beer. The 
rootstocks of S. Bona-nox L., S. glauca Walt., S. rotundifolia L. 
and S. Beyrichi Kunth (S. auriculeta Chap.) were indiscriminately 
collected and used with those of the last. 

Another eastern plant of this family, but of little consequence, 
is the Indian Cucumber-root (Medeola Virginiana \..) with hori- 
zontal yellowish rhizome, 1 to 2 inches long, having a sweetish 
cucumber-like taste and more medicinal than nutritious. ; 

The Star Tulips and Mariposa Lilies of the genus Calochortus 
so abundant and conspicuous in the Pacific States, have nearly all 
edible farinaceous bulbs. The best known, as food plant, is C. 
Nuttalla T. & G. which extends eastward to the Rocky Moun- 
tains and even the prairie region beyond; it is the “Sego” 
of the Indians and Mormons; the bulb is about the size of a wal- 
nut, very palatable and nutritious, and is still used not only by 
Indians but by hunters and miners as well. 

Brodiza Smith, a California genus, has likewise edible bulbs ; 
those of B. congesta Smith and B. capitata Benth., although small, 
are agreeably sweet and mucilaginous. 

Hesperocallis uudulata Gray, the Desert Day Lily, grows in the 
Colorado and Mohave deserts; it has a deep-buried ovate-globose 
bulb 1 to 4 inches in diameter, with firm palatable and refreshing 
flesh, quite welcome to Indians and explorers in the absence of 
other vegetables. 

Chlorogalum pomeridianum Kunth., of California, is better 
known as a soap and a fiber plant than as a food plant, but it is 
also sometimes ealled, and for good reason, “ Wild Potato.” The 
egg-shaped bulb is 1 to 3 inches in diameter and about 4 long, 
thickly covered with coarse brownish fibres resembling the coir of 
the cocoa-nut. A chemical investigation by Prof. Trimble 
showed 1.87 per cent. of saponin (or 6.95 in the absolutely dry 
bulb), as well as glucose, saccharose and mucilage. The large 
_ proportion of saponin accounts for the use of the bulb as a valued 
substitute for soap. Cooking eliminates all acrid and injurious 
_ substances, rendering the bulb good, wholesome food with much 
the taste of camas. — 

_ In the Sedge Family (CypEracea), we only find 2 or 3 food 


115 


plants of minor interest. The Great Bullrush ( Scerpus lacustris L.) 
is a widely distributed plant, ranging from the Atlantic to the 
. Pacific and from British America to the tropics. The stalks and 
leaves are commonly used by many tribes for mats and baskets ; 
the pollen, beaten off and collected on a cloth, is-sometimes made 
into cake. The thick fleshy rootstock of var. occidentalis Wats., 
the « Tule” of the Pacific States, is baked and eaten by hungry 
Indians. Gen. J. Bidwell describes a honey produced abundantly 
on a form of “tule” in Nevada, and eagerly gathered by the 
natives. 

Two species of Cyperus, the Chufa (C. esculentus L.) and the 
Nut-Grass (C. rotundus L.) are extremely noxious weeds in the 
eastern and southern States on account of their rapid propaga- 
tion by tuberiferous stolons and difficult extirpation. The Indians, 
however, looked upon them with favor because of the small edible 
tubers, specially those of Chufa, which are sweet and palatable, 
and even now occasionally planted as food for swine. 

Finally, it remains to mention one Fern, the Common Brake 
or Bracken (Preris aguilina L.), the most widely distributed of the 
order. The blackish rootstocks are eaten in parts of Europe and 
by some of our Pacific Indians. ‘They have a pungency,” say 
Lewis and Clarke, “which was disagreeable to us, though the 
natives eat them voraciously, and they seem to be very nutritious.” 


FRUITS, 


The Indians eat not only all the native fruits which we eat and 
have more or less improved, but also many others for which we 
have never cultivated a taste. It is only of these specially aborigi- 
nal fruits that Ivshallsspeak. 

In the Cactus Family the genus Opuntia, economically speak- 
ing, is probably the most important. Even the fruit of our little 
Prickly Pear (O. vulgaris Haw.) is not entirely worthless, but it is 
in the arid regions of the Southwest that we find a majority of our 
50 native species in their best development. The fruit has a pecu- 
liar and mucilaginous taste, sometimes pleasantly acid, but often 
insipid and mawkish. It contains little nutriment, but quenches 
thirst in the desert. O. Engelmanni Salm, ranging from the mouth 
of the Rio Grande to the Pacific, is the most noteworthy ; not 


116 


only do Mexicans and Indians eat its fruit with avidity, but the 
leavesas well when necessity requires. The leaves, or rather joints, 
of this and allied species are very juicy and an important fodder 
for cattle and sheep, being at once food and drink. 

The Mexican Prickly Pear or Tuna (O. 7uua Mill.), cultivated 
in Mexico from time immemorial, and whose fruit is found in all 
the markets of that country, was not known to our Indians; it 
was brought by the Spaniards into Florida and California where 
it is now naturalized. 

The seeds, not only of the fruit of Opuntia but of all edible 
species of the order, are often separated by the Indians, parched 
and pulverized and made into excellent gruel. 

Most remarkable is the Giant Cactus (Cereus giganteus Eng.) 
the Suhuara or Pitahaya of the Mexicans and the vegetable wonder 
of Arizona, a tree mostly without branches, the straight, grooved 
shaft 30 to 50 feet high. The fruit is 2 to 3 inches long, full of a 
rich crimson pulp of a fine flavor and a great dainty to the 
Apaches, Pimos and Papagos. From it they prepare a clear 
light-brown syrup used as a substitue for sugar, and a fermented 
liquor having the taste and smell of sour beer. 

Still larger, sweeter and finer is. the fruit of GC. Thurderi Eng., 
the Pitahaya Dulce, common in Sonora and Lower California but 
not yet observed in the United States. Half a dozen other arbo- 
rescent species of Cereus with edib‘e fruit have been described 
from Northern Mexico and Lower California. Of the low and 
cespitose species, by far the most interesting from our standpoint 
is the Straw Cactus (€. stramineus Eng.) of Western Texas. The 
ripe fruit is red, 1% inches long and 1 thick, with thin skin bear- 
ing but few spines and easily peeled off; the seeds are so fine as 
to be unnoticed; it is equal or superior, in quality and flavor, to 
the best strawberry. 

The genus Mammillaria or, as revised by Prof. Coulter, Cactus, 
contains many native species bearing red berries of excellent 
taste ; I have eaten these with great relish on the Upper Missouri 
from C. viviparus and in Western Texas from C. Heyderi and others, 
while in Southern California C. Goodrichii is said Bie Orcutt to 
yield a delicious strawberry-like fruit. 

I may close my remarks upon this order by a mention of the 


117 


Peyote (Anhalonium fissuratum Eng.) of the rocky highlands of 
Western Texas and Northern Mexico, a plant which, when chewed, 
is said to produce a sort of delirious intoxication and on that ac- 
count sometimes called « Dry Whisky.” Another species (A. 
Lewini) of the Lower Rio Grande and Mexico possesses the same 
remarkable properties. Both are well worthy of investigation. 

The seeds or “nuts” of many species of Pine are large and 
albuminous, forming, in several parts of the country, not only an 
important but almost an indispensable source of subsistence to the 
Indians. They are oily and often have a strong terebinthine or 
bitter taste when raw, but after being roasted they are not only 
nutritious but also pleasantly flavored. 

Our best known Nut Pines are the Single-leaved Pine (Pinus 
monophylla Torr.), so precious to the Indians of the Great Basin, 
and the Two-leaved Pine or Pifion (P. edulis Eng.), perhaps only a 
variety of the preceding, ranging from Colorado to Texas and 
Arizona. The wingless secds are elliptical or globose in outline . 
and half an inch in length. Speaking of their value to the In- 
dians Dr. Newberry says: ‘‘ They are treasured as their choicest 
delicacies, and a handful of pine-nuts is to an Indian child as much 
of a treat as are sugar-plums to our boys and girls. Some of 
the Pifton groves on the flanks of the Sierra de la Plata, in South- 
western Colorado, have evidently been visited periodically by the 
Pueblo Indians for ages, for fragments of their peculiar ornamented 
pottery cover the ground.” 

The Mexican Nut Pine (P. cembroides Zucc.), with leaves mostly 
in threes but with the same seeds, extends north into Arizona and 
Lower California. Also belonging to this group is P. Parryana 
Eng., of Southérn California and Lower California, with leaves 
mostly in fours. 

The Sugar Pine (P. Lambertiana Dougl.) of the Pacific slope, 
the most gigantic species of the genus, with cone sometimes a foot 
and a half long, has edible seeds a half-inch long.. The Gray-leaf or 
Digger Pine of California (P. Sabiniana Dougl.) has a shorter, 
thick, massive cone with black seeds, the largest in the genus, 
_ Nearly an inch long; these are collected in immense quantities by 
the Digger Indians for winter use, being not only a nutritious 
food but very digestible and specially suitable for delicate 


118 


stomachs. Nearly related to the preceding is P. Coulteri of the 
coast ranges south of San Francisco, with stout, long, strongly 
incurved cone-spines, and somewhat smaller but equally pala- 
table seeds. 

Another genus of Conifers, /wniperus, contains three species 
whose fruit deserves mention: //. occidentalis Hook., with its 
several varieties, extending from Texas to the Northern Pacific 
coast; /. Californica Carr. of Southern California, with a variety 
extending to Utah and Nevada; /. pachyphloca Torr. of Western 
Texas, New Mexico and Arizona, apparently merging into /. 
Mexicana Schl. of Northern Mexico. They all bear abundant 
globose greenish or copper-colored berries with a mealy, resinous 
and not very unpalatable taste. Those of the last-named species 
are the largest (often half inch in diameter) and best ; I have eaten 
them, not greedily, but without repugnance. Mexicans and In- 
dians consume large quantities of these berries and make them 
into abread which, Dr. Palmer says, is of “ chaffy and saw-dust 
consistency.” According to the same authority this bread con- 
tains the following constituents which would indicate unexpected 
nutritiousness: Water 14.34, protein compounds 5.69, starch 
17.87, sugar 10.66, 

Another fruit of much importance to the Indians is that from 
many of our Oaks. Acorns contain starch, fixed oil, citric acid and 
sugar, as well as astringent and bitter principles. They are some- 
times sweet enough to be eaten raw without preparation, but it is 
generally necessary to rid them of their bitter principle ; this is 
done by shelling and skinning, then pounding them into meal and 
washing thoroughly in water; the meal is then ready for boiling 
into mush or baking into*cake or bread. Not long’ago’l received 
specimens of Quercus Garryana from Fort Gaston, in Northern 
California, with the information that the acorns were still a com- 
mon article of food among the Hoopah and other Indians of that 
reservation. To remove the bitterness they place the meal in @ 
hole dug in wet sand, so that in gathering it up more or less sand 
is unavoidably mixed with it, enough to have a decided effect 
upon the teeth. My informant, a medical officer, tells me that he 
has seen an Indian 45 years old with the crowns of his otherwise 
healthy teeth half gone, while, in Indians 60 years old, itis not un- 
common to see all the teeth worn down, even with the gums. 


119 


The White Oaks have sweeter and more palatable fruit than 
the Black Oaks, and it is mostly from them that the Indians sup- 
ply themselves. In California the large conical fruit of Q. /obata 
Nee is considered best by the natives who collect enormous 
quantities of it for winter use. Further north, that of Q. Garryana 
Dougl. is held in the same esteem. All the Live Oaks yield 
Sweet palatable acorns, from the eastern Q. Virginiana Mill. to 
Q. undulata Torr., Q. oblongifolia Torr., Q. pungens Liebm., Q. 
Emoryi Torr., of the Rocky Mountain region, and QO. Engelmanni 
Greene, of Southern California. Our eastern Indians consumed 
large quantities of the acorns of Q. Virginiana, also obtaining 
from them a sweet oil much used in cooking; they, likewise, ate 
the acorns of QO. Michauxii Nutt. and Q. prinoides Willd. 

Of the Black Oaks the only one affording food to the Indians 
is Q. agrifolia Nee, the Coast Live Oak of California. 

_Of the Walnuts and Hickories our Indians knew how to take 
full advantage, and sometimes from the nuts obtained delicacies 
apparently unknown to us; thus Bartram states that the Creeks 
pound the nuts and cast them into boiling water which is then 
passed through a very fine strainer; the thicker oily part of the 
liquid thus preserved is called hickory milk; it is as sweet and 
rich as fresh cream, and an ingredient in most of their cookery, 
specially hominy and corn cakes. 

Our species of Yucca are not only handsome and ornamental, 
but the section Savcoyucca of our arid southwestern territory pro- 
duces fleshy, banana-like, fruits of agreeable taste, wholesome and 
nutritious. Unfortunately the fertilization of these plants, depend- 
ing largely, if not entirely, upon the agency of certain moths, is 
often imperfect, so that well developed fruit is scant. Y. daccata 
Torr. is the most widely distributed of our species, ranging under 
several forms from Southern Colorado to Texas, California and 
Mexico. The ovate or cylindric, more or less beaked fruit is 3 to 
5 inches long,with pulp about a half-inch thick over the large seeds. 
As birds and insects are very fond of this fruit and have the first 
chance at it, Indians and Mexicans collect it when still green and 
let it mature in their dwellings; they also eat it green after baking 
in hot ashes. The young flower buds, when about to expand, are 
roasted and a prized article of diet. I may also mention that 


120 


the leaves of this very useful plant yield strong, flexible textile 
fibres, while the caudex and root are rich in saponin and an ex- 
cellent substitute for soap.  Y. macrocarpa Coville has a stout 
arborescent trunk 10 to 30 feet high and 1 to 1% in diameter, and 
still larger fruit. Y. Schottid Eng., of Southern Arizona, also be- 
longs to this section, as well as Y. 7veculeana Carr., of Northern 
Mexico.. ae 

The NYMPHAEACEAE contain two plants whose seeds are, or 
were, highly prized by the natives, Nelumméo lutea already specially 
noticed for its roots, and Nymphaea polysepala (Eng.) of the north- 
_ern Pacific slope. The former is called Water Chinquapin, from 
the resemblance of its seeds in shape and taste to the Chinquapin 
chestnut of the South; they are eaten raw or cooked and said to 
be even more delicate food than the roots. The latter plant dif- 
fers from the eastern V. advena chiefly in having a larger number 
of sepals and a larger fruit ; the pod is often the size and form of 
an egg, filled with well flavored and nutritious seeds which con- 
stitute one of the most valuable winter stores of the Klamath 
Indians. 

The Ericaceae are rich in finely flavored fruits; the many 

species of Gaylussacia, Vaccinium and Gaultheria furnish the 

Indians with a notable proportion of their vegetable food. Two 
species of Arctostaphylos are likewise utilized by the natives of 
California, A. Manzanita Parry, the Common Manzanita of the 
coast range, and A. tomentosa Dongl.,the Hairy Manzanita of the 
western part of the State. The small apple-like fruit is decidedly 
acid before maturity, tasting somewhat like an agreeably tart apple 
and used for making a cooling drink in summer. When ripe and 
dried, it is pounded and made into cake or bread. 

The Mezquite (Prosopis juliflora DC.) is one of the most wide- 
spread of trees, ranging from the Atlantic to the Pacific, along the 
entire Mexican border, and from the Indian Territory, through 
Texas and Mexico, to South America. It thrives best on bottom 
lands, where it acquires somewhat the size and aspect of an apple- 
tree, but will grow almost anywhere, its long slender tap-roots 
dipping down to great depths in quest of moisture. On arid and 
fire-swept plains the spreading superficial roots absorb most of the 
nutriment, becoming thick and tortuous, while hardly any growth 
is Sranast — ground. 


121 


The Mezquite is invaluable to the Mexicans and Indians of our 
Southwestern territory, to whom it supplies food and fuel, and 
sometimes bad beer. The fruit is a yellow bean-like pod, 6 to 8 
inches long, filled around and between the seeds with a sweet and 
very palatable pulp; it contains more than half its weight of as- 
similable nutritive principles? of which the most important is sugar 
in the proportion of 25 to 30 per cent. Most herbivorous animals, 
specially the horse, mule and donkey, are fond of this pod and 
thrive upon it. : 

A second species deserves mention, P. pubescens Benth., the 
Screw-Bean or Tornillo, so well characterized by the screw-like 
fruit. It is abundant from western Texas to California and south- 
ward into Mexico. The pulp of the bean is even finer than that 
of the Mesquite, but is too scant to be of much importance. 

Another but very different plant of the Lecumnosae affording 
fruit to the Indians is Falcata comosa (L.) Kuntze (Amphicarpa 
monoica Ell.), the Hog Peanut, a slender, twining perennial, ranging 
from Canada to Florida and westward to Dakota. The rudimentary 
lower flowers, borne on filiform creeping branches, bury themselves 
into the ground where they mature usually only one large fleshy, 
obovate or pear-shaped seed. This subterranean seed is edible 
and nutritious. I have seen the Indians dig it up in the spring as 
far north as Bismarck, N. D. The seeds of the pods on the 
upper branches, are said to be as good as peas for the table. 

Along the banks of the upper Missouri and its many tribu- 
taries, grows the Bullberry (Shepherdia argentea Nutt.), a most 
abundant and ubiquitous shrub, sometimes forming miles of im- 
passable thickets. The pistillate plant becomes covered with a | 
profusion of small globose, nearly sessile, bright red berries, which 
contrast prettily with the bluish-white foliage; they are very acid 
and hardly edible until touched by frost in the early days of 
October, when they are sweetened and acquire a pleasant flavor. 
They have always been one of the staple foods of the Sioux and 
other Indians who eat them raw and stewed or mixed with other 
esculents. The whites use large quantities of them for making a 
delicious jelly, preferred by many to currant jelly. An analysis by 
Prof. Trimble gave the following constituents: water 71.28, nitro- 
genous substances 0.14, free acid (citric and malic) 2.45, total 


Sugar 5.47, mucilage and pectin 0.42. 


122 


The other species, Shepherdia Canadensis Nutt, the Soapberry 
of the northern States and British America, bears yellowish-red, 
sweetish-acidulous and bitter berries; these, according to Prof. 
Penhallow, contain 0.74 per cent. of saponin to which they owe 
their persistent bitter taste and their well known property of foam- 
ing when triturated in a little water and beaten up; the thick 
cream-like, strawberry-colored foam thus produced is a favorite 
dish of the natives and, if sweetened, quite palatable. The ber- 
ries are also preserved, dried or made into jam. 

Another plant of this family, with edible fruit, is Alacagnus 
argentea Pursh, the Silverberry of our northern middle States, and 
an attractive garden plant on account of its silvery-white foliage 
and the delicious fragrance of its flowers. The globose berry is 
dry and mealy and not at all appetizing. 

The fruit or hip of several of our wild Roses, after being touched 
by frost, is sweet and palatable; as it persists through most of the 
winter, when hardly anything else is available, it sometimes be- 
comes useful food to the natives as well as to birds and mammals. 
Rosa Nutkana Presl., the Nutka Rose, ranging from the northern 
Pacific coast to the Rocky Mountains, is the most showy of west- 
ern Roses, having the largest flower and fruit; the latter is juicy 
and pleasantly acidulous and an excellent antiscorbutic for the 
Indians of Alaska. 

Passing over the many native plants yielding edible seeds, 
specially of the Gramineae, I shall close with the notice of a few 
of those whose stem and foliage afford food to the Indians. In 
this connection the genus Agave is first to be mentioned; it is 
essentially Mexican, only a few species extending into our southern 
territory, and has always been of the greatest economic importance. 

The sap of A. Americana, a species sparingly naturalized, but not 
indigenous, north of the Rio Grande, is converted into a weak alco- 
holic beverage called “pulque,” the national drink of Mexico. 
Other species, baked, supply the “ mezcal de comer,” an excellent 
article of food, and also, by fermentation and distillation, a strong 
drink called “aguardiente mezcal.” The edible part consists of 
the thick juicy base of the leaves, the fleshy axis and central bud, 
together forming the “cabeza” or head which is slowly baked in 
a pit or oven. In the raw state no sugar can be discovered in 


123 


these plants, but only a citro-glucosid which by heat is converted 
into grape sugar and citric acid, so that, by cooking, the cabeza is 
rendered very sweet and pleasant to the taste. 

Our Indians were not slow in learning the value of Agave from 
the Mexican natives; from time immemorial they have utilized 
our indigenous species, and wherever these grow can be found the 
stone-lined pits in which the mezcal heads are, or were, baked for 
food. As our native species do not yield sap enough for the pro- 
duction of pulque, and the process of fermentation and distillation 
is too complicated for native art, it follows that our Indians never 
obtained any alcoholic drink from them. 

A. Parryt Eng. is the Mezcal of New Mexico and Northern 
Arizona, one of the staple foods of the Apaches. When properly 
Prepared it is saccharine, palatable and wholesome, mildly acid, 
laxative and antiscorbutic. 4. Palmen Eng. takes its place in 
southern Arizona. 4. Washzeni Eng. is the common Mezcal 
species of the mountains of western Texas, while A. deserti Eng. 
is that of southern California and adjoining deserts. 

On the mesas and foot-hills of western Texas and northern 
Mexico where Agave is rare, the observant Indians discovered a 
plant which takes its place and is equally useful, furnishing both 
food and drink. This is Dasylirion Texanum Scheele, the Bear- 
Grass of the Americans and Sotol of the Mexicans. When 
trimmed down, the head or edible portion consists entirely of the 
thick, €xpanded and imbricated bases of the leaves; it is refresh- 
ing and palatable even in the raw state, but of much better flavor 
when cooked. By fermentation and distillation, the Mexicans ob- 
tain from it a strong whisky called Sotol mezcal, of penetrating 
smell and peculiar taste, and the common alcoholic beverage of the 
frontier population. 

The LeNNoacear have several interesting food plants, the prin- 
cipal of which is Ammobroma Sonorae Torr., the Sand-food, a 
leafless parasite in the sand-hills of southern Arizona and Lower 
California. The long fleshy stem, creeping in the sand, is edible 
Taw and cooked, but specially palatable when boiled or roasted, 
the taste being variously described as that of a sweet potato or the 
heart of a cabbage. It is eagerly eaten by Cocopa, Papago and 
Yuma Indians, as well as by the Mexicans. It is a valuable 
Substitute for water in the desert. 


124 


The Classification of the Archegoniates. 


By Lucien M. UNDERWOOD. 


There has hitherto been much looseness in the codrdina- 
tion of the groups of plants above what have been regarded the 
natural orders and at the same time wide difference in usage in 
group names. It is equally apparent that in some respects 
the botanists have followed a different system of nomenclature 
from the zodlogists, and they have not even followed a uni- 
form or consistent system among themselves. While it is 
not possible to crowd a series of forms within the limits of a rigid 
classification, or accurately coordinate all natural groups of plants 
in equally related categories, it certainly is possible to follow cer- 
tain broad principles and maintain a uniformity of nomenclature 
for the higher groups as well as for genera and species. Leaving 
particulars for further discussion there ought to be no difficulty in 
securing the adoption of the following system of group names and 
sequences : = 


SuB-KINGDOM. 
Crass. 
ORDER. 
FAMILy. 
GENUS. 
SPECIES. 


This is in skeleton the exact usage of the zodlogists and con- 
forms more nearly with the greater portion of the more recent 


usage among those whose work is connected with cryptogamic “ee 


botany. The differences that have been most prominent have 
arisen among the phanerogamic botanists, and some of the differ- 
ences have even become almost hoary-headed from long usage- 
Indeed incipient baldness marks not a few of them, and perhaps 4 
proper ventilation of some of the remainder will cause this to be- 
come even more apparent. 

1. In place of the term sud-fingdom, so generally used by 
zoologists, the term series has been used. The use of the former 


125 


must eventually be followed by all who deal in taxonomy, if con- 
sistency is to be maintained. : 

2. There has been a standing confusion of the terms order and 
Jamily, The term “natural order” has a considerable antiquity, but 
has been confined Slargely to the Spermatophytes. While crypto- 
gamic botanists have by no means been uniform in their termino- 
logy, they have in the majority of instances clearly distinguished 
these two group names. In this they have followed the lead of 
the zodlogists, for there would be no opportunity in animal classi- 
fication to confound such distinct group names as the order Car- 
nivora and the Jamily Canidae. And yet the Ranunculaceae have 
been called indiscriminately a natural order (or simply an order) 
andafamily. On account of this general confusion among the plants 
which are popularly the best known, it will be all the more diffi- 
cult for a time to introduce a uniform and consistent system. 
And yet it seems to need no argument to show that there is no 
good reason for perpetuating this confusion in our terminology, 
and the stand taken on this point by the German systematists, and — 
quite consistently carried out in a number of the most elaborate 
floras that have ever appeared in any country, will go far toward 
establishing the correct terminology. 

3. There has also been a tendency to use the term cofort, much 
as we have used the term order in the proposed system. This 
Was introduced by Lindley, and its adoption by Bentham and 

Hooker has had the tendency to fix its use especially in those 
quarters where other emanations from the same source have had 
much weight. As the Spermatophytes are not all of the vegetable 
world and much less not all of the domain of living things, it be- 
comes a question when we broach the matter of uniformity of 
usage as to whether we shall follow a generally established prin- 
ciple or merely a localized usage. Since all zodlogists and 
the great majority of cryptogamists have already established 
the term order to include a group of families, it would seemingly 
be folly to attempt to adopt the term cohort in place of the term 
order. Here priority of use might well. be argued in addition to 
the axiom that a part is not more important than the whole. : 

The adoption of intermediate groups such as sub-classes, sub- 
Orders, sub-families and sub-genera can well be left to monog- 


126 


raphers of special groups, for their necessity will evidently vary in 
every group of plants. It would be a consummation devoutly to 
be wished if their use anywhere were restricted as much as possi- 
ble, for a too extended and minute subdivision is not only not de- 
‘manded by most groups of organic forms, but when introduced in 
excess tends more to confusion than otherwise and often obscures 
the real groupings of allied forms. 

So far as tniformity of termination is concerned it would seem 
that the termination-a/es could properly be adopted for orders and 
the termination-aceae for families where this would not interfere 
with long established names. In this connection uniformity would 
seem to be of more importance than priority, especially as the 
limitations of these larger groups have been so long a matter of 
uncertainty and variety of opinion. In the groups above orders, 
there would seem to be less necessity for uniformity of termination 
than in the orders and families themselves. 

In order to illustrate the matter of uniformity of terminology 

‘and as a contribution toward securing this uniformity, I present 
‘the following arrangement of the sub-kingdom known as Arche- 
goniata. It will be necessary as an introduction to state the limt- 
tation of that group as we understand it. In the present 
condition of our knowledge of plant forms, we can probably find 
no more satisfactory primary division of the plant world than the 
following, though of course we are still very far from a natural ar- 
rangement. It is practically the arrangement followed in the 
Engler-Prantl series* with minor modifications. 

Sub-kingdom MyCEtTozoa. 

Sub- kingdom THALLOPHYTA, 

Sub-kingdom ARCHEGONIATA. 

Sub-kingdom SPERMATOPHYTA. 

A few remarks may be to the point in explanation of this di- 
vision, which in its lower half is necessarily artificial and unsatis- 
factory. 

1. The slime moulds are evidently sufficiently distinct from 
other plants to warrant a separation in this way. That they have 
undoubted affinities with the animals no one can doubt, but it is 
equally clear that they have decided affinities with at least two 


* Die natirlichen Pflanzenfamilien, 2: 1, 1-2. 


127 


groups of plants, the bacteria and the moulds (Mucorini), that can- 
not be disregarded. While their structure and function causes 
them to stand somewhat midway between plant and animal, it 
would seem more consistent in a systematic arrangement to place 
them at the bottom of. the plant world. 

2. Until the inter-relations of the algae and fungi can be more 
definitely known and some suitable system devised that shall make 
it possible to unite allied forms on a rational basis, it will be best 
to maintain this unnatural and heterogenous group which we call 
Thallophyta. It is true, as has been stated, that the group is 
Practically undefinable, but that is always true of the last division 
of any series in which those forms are placed that do not conform 
to any known natural arrangement. The close alliance of some 
of the lower fungi and algae .absolutely precludes the idea of 
establishing these as primary groups, and of the various systems 
of subdivision that have been maintained there are none thet seem 
to be founded on real or natural relationships. Unsatisfactory as 
the group is, we see no present improvement that can replace it.. 

3. As has been often maintained, there is not a wide distinction 
between a simple Riccia and some of the green algae, and the 
group Archegoniata is not entirely distinct at its lower limit. 
This, however, is true of any great group when its outliers are all 
known, for evolution has not yet resulted in the destruction of all 
the connecting links. We have for the same reason excluded the 
Characeae from this group because of their apparently stronger 
affinities with the other green algae. For a similar reason we 
have included the Gymnosperms at the upper limit because their 
affinities when properly. understood seem to link them more 
strongly here than with the higher plants. 

With these limitations we present the following outline of the 
classification of the Archegoniata. 


Sub-kingdom ARCHEGONIATA. 
Class BRYOPHYTA. 
(Sub-class Hepaticae.) * 
Order Marchantiales. 
*I have placed these subgroups in parenthesis as representing group names that 


Modern research seems destined to retire toa merited oblivion. I have elsewhere 
discussed their abandonment more in detail. Cf Proc. A. A. A.S. 43: 259-274, 1894. 


128 


Family RICCIACEAE. 
Family MARCHANTIACEAE. 
Order Jungerianiales. 
Family METZGERIACEAE. 
Family }UNGERMANIACEAE. 
Order Anthocerotales. 
Family ANTHOCEROTACEAE. 
(Sub-class Musct.)* 
Order Sphagnales. 
Family SPHAGNACEAE. 
Order Andraeales. 
Family ANDRAEACEAE. 
Order Archidtales, 
Family ARCHIDIACEAE. 
Order Bryales. 
(I omit the somewhat numerous families of 
this order as unnecessarily extending this 
_ illustration.) 
Crass PTERIDOPHYTA. 
Order Filicales. 
Family OPHIOGLOSSACEAE. 
Family MARATTIACEAE. 
Family HYMENOPHYLLACEAE. 
Family OSMUNDACEAE. 
Family ScHIZAEACEAE. 
Family GLEICHENIACEAE. 
Family PoL_ypapiacear. 
Family CYATHEACEAE. 
Family MARSILEACEAE. 
Family SALvINIACEAE.* 
Order Equisetales. 
Family EQuisETACEAE. 
Family CALAMARIACEAE (fossil).+ 
Order Sphenophyllales. 
Family SPHENOPHYLLACEAE (fossil).+ 
Order Lycopodiales. 
Family LycopoprAcEae. 


_* See foot-note on preceding page. 


129 


Family Psttoraceae. 
Family LepipopENDRACEAE (fossil).+ 
Family S1GiLiartaceak (fossil).+ 
Family SELAGINELLACEAE. 
Family Isorracear.+ 
Crass GyMNOSPERMAE. 

Order Cycadales. 
Family Cycapacear. 

Order Cordaitales (fossil). 

Order Pinales. 
Family Pixnacear. (Cowiferae.) 

Order Guetales. 
Family GNETACEAE. 


The above classification is not proposed from the fact that it 
contains anything especially novel, but is merely an adaptation of 
the principle of uniformity in the terminology of the lower groups. 
So far as unlike things can be compared at all, the above families, 
as indicated, are groups equivalent with the so-called “ natural 
orders ”’ of the higher plants, e. g. Ranunculaceae, Liliaceae, Orcht- 
@aceae, etc.. The separation of the Gymnosperms entirely from 
the Spermatophytes will doubtless jostle with the ideas of those 
Who still sandwich them in between the Monocotyledons and 
Dicotyledons, unaware of the modification that has taken place in 
our ideas of homology in the past two decades, while they have 
been asleep and failed to note the march of progress over their heads. 
In some instances it may be desirable to introduce a few subdivi- 
sions, but in the above I have purposely left them out except in 
One instance, that the simplicity of the arrangement might be the 
More apparent. They should not appear except for sufficient 
and well founded cause. 


GREENCASTLE, INDIANA. 


* No lineal series of these families can represent their true affinities. 
t+ The position of many of these fossil forms is still problematical. 
} The position of this family is by no means permanently decided. 


130 


RULES FOR CITATION 


Adopted by the Madison Botanical Congress and Section G, A. A. A- Ss 


Writers and publishers of botanical matter are earnestly requested 
to adopt the forms here recommended. - Examples of various cita- 
tions illustrating the application of the rules in specific cases are 
given. Correspondence may be addressed to Secretary of the Com- 
mitlee on Bibliography, 1284 Massachusetts A ve., Cambridge, Mass. 


In each complete citation there should be given the following 
items: 

a. Author’s surname in full, followed by a comma. 

6. Exact title, verbatim, following the capitalization required 
by the usage of the language in which the title is written, but not 
necessarily the capitalization employed. 

c. Name of periodical or work, abbreviated in accordance with 
list of journals and catalogue of authors under recommendation I. 
a. b,* 

d. Series, if any, in Roman capitals. 

é. Volume number in bold face Arabic numerals, followed by 
colon. In case there is no volume number, the number of the 
part, heft, lieferung, or fascicle is to occupy this place but is to 
be printed in Arabic numerals of ordinary face. When a volume 
is composed of parts separately paged the number of the part 
shall be written as an index figure to the Volum2-number. Vol- 
umes in parts with continuous paging require no designation of 
parts. 

f. Page, in Arabic numerals of ordinary face. In case paging 
of the paper is in Roman numerals these should be used, pre‘er- 
ably small caps. Re-paging in reprints and separates is to be in- 
dicated by enclosing the numerals in parentheses. In case the 
original paging is unknown an em dash should occupy its place, 
the reprint paging being given in accordance with the foregoing 


*See Proc. Mad. Bot. Cong. 45. Je 1894. 2 


51 


Tile, ‘No individual or unique paging is to be cited under any 
circumstances. 

g. Figures, plates and exsiccate are to be printed in Italic 
Arabic numerals, the number designating the figure or plate to 
be preceded .by the abbreviations f. and pi., respectively, in Ital- 
Ics. d. following a page number may be used, when desired, to 
indicate description of a species. 

h. Exact date must be given if possible, written in the mode 
and with the abbreviations for months used by Library Bureau.* 
The year at least must be given. 

7. Punctuation. Except the comma following the author’s 
name, and the colon following the volume number all the items 
are to be separated by periods. If another citation follows in the 
Same line it is to be separated from the first by an endash. Spe- 
cific, generic and. varietal names are to be written and punctuated 
in the method used in the ‘‘List of Pteridophyta and Spermato- 
eg issued under the direction of the Botanical Club, A. A. 

J. If it is considered desirable to give other data than series 
number (if any), volume number, page and date, these should 
be added in brackets after the date. But useless or unnecessary 
data should be avoided. 

k. Citations of reviews, abstracts, and all such secondary refer- 
ences should be enclosed in parentheses. 


Examples. 


vy Lagerheim, G. von. Ueber das Vorkommen von Eu- 
ropzischen Uredineen auf der Hochebene von Quito. 
Bot. Centralb. 54:324~-331. 1893. 

2. Trelease, W. A revision of the American species of 

Epilobium occurring north of Mexico. Rept. Mo. Bot. 

Gard. 2:69-117. pi. z-g8. 22 Ap18ogl. 

3- Sargent, C. S., Editor. Populus monticola. Gard. 

ae and For. 7:313. f. 56. 8Ag 1894. 


*Those abbreviations are as follows: Ja, F, Mr, Ap. My, Je, Jl, Ag, S, O, N, 
D; i. e., the initial of the month followed by the first distinctive letter. 


152 


Dietel, P. Die Gattung Ravenelia. Hedw. 33: 22-48. 
' pl. 1-5. 30Ja. 49-69. 15Ap 1894. 
The foregoing are correct forms for catalogue by author. The 


following illustrate cases arising under the rules indicated by 
the letter preceding. 


El. and Everh. Pyren. 491. My 1892. 
Proc. Phil. Acad. 1894: 53-59. 1894. 


The year number, 1894, is the volume number, and not neces- 
sarily the year cf publication. E. g., 


Bessey, Am. Pomol. Soz. 1885: 42. 1886. 
Mez, C. Bromeliacece. III. Flora Brasiliensis 115: 
425-634. pl. Sr-r1rg. 1F 1894. 


Saccardo, P. A. Syll. Fung. 77:481. N 1890. 


j- Bull. Geol. and Nat. Hist. Surv. Minn. 9:39-42. 2Mr 


1894. 
Not 9?; nor 9 part 2; nor 1894 [part 2]. 


j- Linn. Sp. Plant. 62:125. 1852. [ed. Willd.] 


Gray, A. Man. Bot. 225. 1890. [6th ed.] 

Peck, C. F. Rep. N.Y. Mus. 47:— (18). N 1894. 

Ell. and Rverh. N. A. F. 7642. F1889. 

Rept. Mo, lsut. Gard. 2:98. d. pl. 28. 22 Apr89gt. 

Beringer, Am. Jour. Pharm. 66:220 My 1894.—Tu 
lasne, Ann. Sci. Nat. Bot. III. 7:85. d. pl. 2. f- 3- 
1847. 

Bailey, The Japanese plums in North America. Bull. 
Cornell Exp. Sta. 62:3-36. Ja1894._ [Illust.] 


The figures are not numbered. 


Ell. and Kell. Jour. Myc. 1:12. d@. Ja1885.—(Hedw. 
24:45 d. Je1885.) Peck, (Grev. 222111. Je1894.) 


133 


Proceedings of the Club. 
Tugspay EveninG, FEBRUARY 12TH, 1895. 


Vice President Allen in the chair and 29 persons present. 

Lieut. Wm. Lassiter, 1st Artillery, U. S. Army, was elected 
an active member. — 

A communication from the Secretary of the Council of the 
Scientific Alliance of New York, transmitting a draft of a pro- 
posed act for incorporating the Council, was read. The matter 
was discussed by the chairman, Mr. Van Brunt, and the Secretary. 
On motion and unanimously carried, it was 

Resolved, That this Society approves of the measure outlined 
in the proposed act of incorporation just presented by the Council 
of the Scientific Alliance of New Y ork, and, so far as this Society 
is concerned, hereby authorizes the said Council to procure the 
Passage of the said act by the Legislature and thereafter to carry 
the same into effect. : 

A communication from Professor Halsted, who was announced 
to speak on «“ Abnormalities in Plants due to Fungus Diseases,” 
was read, stating his inability to be present. 

The paper of Mr. Van Sickle, deferred from the last meeting 


was then presented, entitled. “:Notes. from my Herbarium.” The 
speaker exhibited and remarked upon a very large number of the 


rarer plants of northern New Jersey, representing numerous un- 
recorded localities and several species hitherto unknown to occur 
within the State. The communication was a very valuable addi- 
tion to our knowledge of the distribution of the local flora and 
will be published in a subsequent issue of the BULLETIN. 

Professor Britton read the announced paper of the evening, 
“The Family Ranunculaceae,” outlining his proposed treatment 
of the group in the “ Systematic Botany of North America.” 


WEpNEsDAY EVENING, FEBRUARY 27TH, 1895. 
Vice President Allen in the chair and about 165 persons 


present. 

Miss Beatrix Jones was elected an active member. 

The announced paper of the evening was then read by Le 
Cornelius Van Brunt, entitled « Wild Flowers in and about New 


134. 


York City.” The paper was illustrated by a large number of lan- 
tern slides made and colored from nature, the beauty of which 
elicited many expressions of admiration from the audience. The 
paper was discussed by the Secretary, who expressed the hope 
that those present would follow up their acquaintance with our 
wild flowers made from the lantern slides, by a personal acquaint- 
ance made in the field during the coming season, in connection 
with the excursions of the Cluband of the summer class in botany. 
Dr. Britton called attention to the scientific features of such ac- 
curate illustrations, by which he had noted that a rare species of 
Asclepias, A. decumbens, not previously reported from this locality, 
occurred in the vicinity of New York. He also mentioned the 
strong contrast between the lowland and mountain forms of Ee 
patorium purpureum as endorsing his view that they were distinct 
species. 

Dr. Britton called attention to two important publications, viz., 
1, American Algae, Century 1, 1894, by Miss Josephine E. Tilden, 
of the University of Minnesota; 2, Phycotheca boreali-Ameri- 
canae, fascicle 1, by Messrs. Collins, Holden and Setchell. 


Index to recent Literature relating to American Botany. 


Allen, T. F. Japanese Characeae—II. Bull. Torr. Bot, Club, 22: 
68-71. 26 F. 1895. 
Describes Nitella pulchella, N. subglomerata Faponica, and N, sublucens as 

new. 

Arthur, J. C. Black-knot and other Excrescences. Trans. Ind. Hort. 
Soc. 1894: 76-80. 1894. 


Arthur, J. C. Discrimination of Diseases without the Use of the Micro- 
scope. Amer. Florist, g: 646. 22 F. 1894. Also Ann, Rep. Am- 
Carnation Soc. 1894: 8-14. 1894. : 
Describes seven or eight fungous diseases of carnation and tells how to recognize” 

them aided only by a hand lens. 

Arthur, J. C. and Holway, E. W. D. Uredineae exsiccatae et 
icones. Fascicle I. Decorah. pp. 4. f/. 7. 31 packets of specimens. 
S. 1894. 


Contains 17 species of Eaten ay with drawings of the — on a uniform 
scale of magnification. 


135 


Bastin, E.S. Structure of Cimicifuga. Am. Journ. Pharm. 67: 121- 
128. f. 7-7. Mr. 1895. : 


Bates, J, M. Notes on the Trees of Northern Nebraska. Am. Nat. 
28: 1034-1036. D. 1894. 


Beach, S. A. Some Observations on the Life-history of Plowrightia 
morbosa (Schw.) Sacc. Ann. Rep. N. Y. Agr. Exp. Sta: (Geneva) 
1893: 686-688. 1894. - ruts 


An infection of young nursery stock from the conidia of 1892 (or possibly from 
ascospores of the following winter), produced an outbreak of the knots in June, 1893; 


Bessey, C. E. Notes on the Distribution of the Yellow Pine in Ne- 
braska. Gard. & For. 8: 102— -103. 13 Mr. 1895. 


Britton, E.G. Contributions te American Bryology—IX. A Revis- 
- ion of the Genus Scouleria with Description of one new Species. 
Bull. Torr. Bot. Club, 22: 36-43. A/. 227. 15 Ja. 1895. 


Describes S. marginata from Washington and California. 


Britton, E.G. Contributions to American Bryology—X. Bull. Torr. 
Bot. Club, 22: 62-68. AZ. 229-271. 26 F. 1895. 


Discussion of Physcomitrella patens and Aphanorhegma serrata and a hybrid of 
the latter with some Physcomitrium, with plates. 


Britton, N. L., and Kearney, T. H.,Jr. An Enumeration of the 
Plants collected by Dr. Timothy E. Wilcox and others in Southeastern 
Arizona. Trans. N. Y. Acad. Sci. 14: 21-44. Ja. 1895. 


New species in Aristida, Muhlenbergia, Mirabilis, Berberis, Cercocarpus, 

Lrythrina and Kuhnistera. 

Budd, J. L. The Russian Thistle in its natal Home. Bull. Iowa 
Agr: Exp. Sta. 26: 30-33. 1894. 

Campbell, D. H. Observations on the Development of Marattia Doug- 
fasii Baker. Ann. Bot. 8: 1-20. f/. 7-2. Mr. 1894. 


Campbell, D. H. The Origin of the sexual Organs of the Pteri- 
dophyta. Bot. Gaz. 20: 76-78. 16 F. 1895. 
Cheney, L. S. Leucoplasts. . Bot. Gaz. 20: 81. 16 F. 1895. 
Chester. F. D. Report of Mycologist. Sixth Rep. Del. Exp. Sta. 
18y3: 103-132. Ja. 1895. 
Gives treatment for Peach rot (Monilea) ; observations on rot of scarlet clover 
(Sclerotina T> rifoliorum), Colletotrichum in cow pea seed and ripe rot ( Cod/eto- 
trichum phomoides) of tomatoes. 


Collins, F. S., Holden, I. and Setchell, W. A. Phycotheca Bore- 
ali-Americana. Fascicle I. Malden, Mass. 1895. 


136 


Coulter, J. M. New or noteworthy Compositae from Guatemala. 
Bot. Gaz. 20: 41-53. p/. 5-6. 16 F. 1895. 
Describes Vernonia Luxensis, V. Heydeana, V. Shannonti, Ageratum rugosum, 

Eupatorium griseum, E. vernonioides, Willougbaeya globosa, Mallinoa (n. g:) 

corymbosa, Polymnia Quichensis, Montanoa Samalensis, Verbesina Donnell- 

Smithii, Peresiopsis (n. g.) Dennell-Smithit, with plates of the two new genera. 


Coville, F. V. Report of the Botanist U. S. Department of Agricul- 
trure for 1893. Rep. Secy. Agric. 1893: 235-244. 1894. 


Crandall, C.S. The Russian Thistle. Bull. Col. Agr. Exp. Sta. 28: 
pp. 18. £7. 6. S. 1894. 


Davis, B. M. Zuglenopsis ; a new Alga-like Organism. Ann. Bot. 8: 
377-390. pl. 19. D. 1894. 


Deane. W. Notes from my Herbarium—I. Bot. Gaz. 20: 12-!5- 
18 Jy. 1894. 


Dewey, L. H. The Russian Thistle. Cir. Div. of Bot. U. S. Dept. 
Agric. 3; pp. 8. f. 3. 1894. 


Eggleston, W. W. The Flora of Mt. Mansfield. Bot. Gaz. 20: 
72-75. - 16: F.1895. 


Ellis, J. B. and Everhart, B. M. New Species of Ustilagineae and 
Uredineae. Bull. Torr. Bot. Club, 22: 57-61. 26 F. 1895. 


Species of Ustilago, Entyloma, Uromyces, Puccinia and Aecidium. 

Foerste, A. F. Botanical Notes. Bot. Gaz. 20: 78-80. 16 F. 1895- 
Spreading of raspberry bushes and superposed buds and leaf scars. 2 

Ganong, W. F. An Outline of Phytobiology. Educational Review, 
St. John, New Brunswick. Je. 1894 [Reprint, pp. 15, St. John, 
1894]. 

Gepp, A. Additional Notes on Mr. W. R. Elliott’s Hepaticae. Journ. 
Bot. 33: 82-84. Mr. 1895. 
Notes on Lejeunea and Metzgeria. 

Guignard, L. Sur l’existence et la localisation de l’emulsine daus les 
plantes du genre Manthot. Bull. Soc. Bot. France, 4: 101-107. 1894- 

Guppy, H. B. On the Habits of Zenna minor, L. gibba and L. 
folyrhiga. Journ. Linn. Soc. Bot. 30: 323-330. 6 O. 1894. 

Halsted, B.D. Some fungous Diseases of Beats. Bull. N. J. Exp. 
Sta. 107: pp. 13.f. 5. 10 Ja. 1895. 

Handy, R. B. Peanuts: Culture and Uses. Farm. Bull. U. S. Dept 
Agric. 25: pp. 24, fig. 1895. 


137 


Harms, H. ~ Plantae Lehmannianae in Colombia et Ecuador collectae- 
Passifloraceae.  Beibl. Engler’s Bot Jahrb. 18: Hft.5.1-14. 21 Ag. 
1894. 


Havard, V. Family Nomenclature. Bull. Torr. Bot. Club, 22: 77, 
78. - 26 F. 1895. 

Hay, G. U. Report on Botany. Bull. Nat. Hist. Soc. New Bruns- 
wick, 12: 69, 70. 1894. 


Heller, A. A. Botanical Explorations in Southern Texas during the 
Season of 1894. Contrib. Herb. Franklin and Marshall College 1: 
1-116, pf. 9: 6 F. 1895. 

List of a-large number of species with synonymy, and critical notes, novelties 
being described in Panicum, Rumex, A mor pha, Kuhnistera, Sida, Samolus, Asciep- 
tas, Cressa, Verbena, Pentstemon and Hou:t. nia. 


Hieronymus, G. Plantae Lehmannianae-in Colombia et Ecuador 
collectae additis quibusdam ab aliis collectoribus ex iisdem regionibus : 
allatis determinatae et descriptae.-Compositae. Engler’s Bot. Jahrb. 
19: 43-75. 13 Ap. 1894. : 

Hill, E. J. Zradescantia Virginica var. villosa Watson. Bull. Torr. 
Bot. Club, 22: 71-73. 26 F. 1895. 

Hooker, J. D. Aphaerema eee Curt. Bot. Mag. 51: A/. 7398. 
F. 1895. 

Native of South Brazil. 

Howe, M.A. Chapters in the early History of Hepaticology.—HII., 
IV. Erythea, 3: 1-6, 25-30. 1 F. 2 Ja. 1895. 

Review of Columna, Caspar Bauhin and others. 

Howell, T. New Species of Pacific Coast Planis. Erythea, 3: 32-36. 
1 F. 1895. 


New species in Lepidium, Arabis, Cardaminée, Mitella, Saxifraga, Ribes, Ert- 
Seron, Senecto and Phacelia. 
Jenman, G.S. Aspidium (Lastrea) basiattenuatum. Gard, Chron. 
17%. 132. 2 F: 1896: 
A new species from Jamaica. 
Jenman, G. S. Polypodium ( Gontopterts) nigrescentium. 
Chron. 17: roo. Ja. 1895. 
A new species from Jamaica. 
Koehne, E. Glossopetalon eisnamtvacn n. sp. Gartenflora, 43: 


237-240. f. 52. 1 My. 1894. 
Koehne, E. Quercus Gambelit. Gartenflora, 44: 6-10: f: 7. 1 Ja. 


1895. 


Gard. 


138 


Knuth, P. E. Loew’s Bluthenbiolozische Floristik die mittleren und 
nérdlichen Europa sowie Grénlands. Biol. Centr. 15: 78. 15 Ja. 
1895. 

A brief review of Loew’s work. 

Lindau, G. Beitrage sur Argentinischen Flora. Beibl. Engler’s Bot. 
Jahrb. 19: Hft. 4.8. 28 D. 1894. 

Macoun, J. The Forests of Canada and their Distribution, with Notes 
on the more interesting Species. Trans. Royal Soc. Can. 12: Sec. 
4. 3-20. 1894. 

Macoun, J. M. Contributions to Canadian Botany—lI.-II. Can. 
Rec. Sci. 6: 23-27. Ja.; 76-88. Ap.; 141-153. N. 1894. 

Record of new localities and critical notes. New species and varieties by Trelease 
in Efilobium, in Spiesia by Britton, in Agrostis and Poa by Scribner. 
Macfarlane, J. M. Irrito-contractibility in Plants. Biological Lec- — 

tures, Wood’s Holl, 1893: 185-209. 1894. 

Meehan, T. Aster spectadilis. Meehan’s Month.: 5: 41, 42. AY 3: 
Mr. 1895. 


Mobius, M. Ueber einige an Wasserpflanzen beobachtete Reizer- 
scheinungen. Biol. Centr. 15: 1-14; 33-44. 1 Ja. and 15 Ja. 
1895. 


Discussion of irritable organ-positions in Ceratophylium, Naias, Ranuncu Us, 
Cabomba, Myriophyllum, Elodea, Hippuris, Callitriche and Chara. 


Mohr, C. Die Walder des Siidlichen Alabamas. Pharm. Rund. 13: 
30-33- FF. 1895. (Concluded. ) 

Mottier, D. M. Contributions to the Life-history of Votothy/as. 
Ann. Bot. 8: 391-402. p/. 20, 27. D. 1894. 

Nash, G. V. American Ginseng ; its Commercial History, Protection 
and Cultivation. Bull. U. $. Dept. Agric. Div. Botany, 16: PP- 
22. fig. 1895. 


Discusses Panax guinguefolium. 

Newcombe, F.C. The Cause and Conditions of lysigenous Cavity- 
formation. Ann. Bot. 8: 403-421. D, 1894. 

Nichols, M. A. Observations on the Pollination of some of the Com- 
positae. Proc. lowa Acad. Sci. 1: 100-103. 1894. 

Owen, M.L. Tillaca simplex. Bot. Gaz. 20: 80, 81. 16 F. 1895- 

Pammel, L. H. Botany of the Russian Thistle. Bull. Iowa. Agric. 
Coll. Exp. Sta. 26: 8-26. p77. 1894. | 

Pammel, L. H. Results of crossing Cucurbits.' Bull. Iowa Agric- 
Coll. Exp. Sta. 23: 906-917. pl. 2, f. 2. 1894. 


159 


Pammel, L. H. Some obnoxious Weeds of Iowa. Rep. Iowa State 
Agric. Soc. 1893: 437-442. 1894. 

Praeger, R.L. Juncus tenuis Willd. Journ. Bot. 33: 86. Mr. 1895. 
Allusion to its occurrence in Ireland, 

Reiche, K. Zur Kenntniss der Chilenischen Arten dur Gattung Oxa/is. 
Engler’s Bot. Jahrb. 18: 259-305. p/. 9. 1894. 

Ridgway, R. Additional Notes on the Native Trees of the Lower 
Wabash Valley. Proc. U. S. Nat. Mus. 17: 409- 421. 1894. 
Measurements given of many species. 

Robertson, C. The Philosophy of Flower Seasons and the phaeno- 
logical Relations of the entomophilous Flora and the anthophilous - 
Insect Fauna. Am. Nat. 29: 97-117. ~/. 8-10. F. 1895. 

Rolfe, R. A. New Orchids. Kew. Bull. 1894: 361, 363, 364, 392, 


393-395. 1894. 


Descriptions of a large number of Runhans species. 
Sargent, C. S. Notes on North American Oaks. Gard. & For. 8: 
92-93. f. 13-14. 6 Mr. 1895. 


Description and illustration of a new oak; Quercus Toumey?, from the hills of 


Southeastern Arizona. 

Sargent, C. S., Editor. The Forests of the Wabash Va alley. Gard. & 
For. 8: 101-102. f. 46-17. 13 Mr. 1895. 

Saunders, De A. A preliminary Paper on Costaria with Description 
of a new Species. Bot. Gaz. 20: 54-58. f/. 7. 16 F. 1895. 


C. reticulata from California. 
Schilling, A. J. Anatomisch-biologische Untersuchugen iiber die 
Schleimbildung der Wasserpflanzen. Flora, 78: 280-364. 4 Je. 1894. 
Schlichter, R. Britrige zur Kenntniss Siidamerikanischen Asclepia- 
daceen. , Beibl. Engler Bot. Jahrb. 18: Hft. 4. 1-37. 22 Je. 1894. 
Schumann, K. Aariota salicornioides. Monatss. Kakteen K 5: 
22-20. figs. F. 1895. 
Scribner, F. L. Giant Knotweed or Sachaline. Cir. Div. 
S. Dept. Agric. 5: pp. 4. f. 3. 1895. 


Discusses Polygonum Sach dinense and its value as a forage plant. 

Scribner, F. L. The Flat Pea (Lathyrus sylvestris Wagneri.) Cir. 
Div. Bot. U. S. Dept. Agric. 4: pp. 7./. 2. 1895. 

Small, J. K., and Vail, A. M. Report on the Botanical Exploration 
of Southwestern. Virginia during the Season of 1892. Mem. Torr. 
Club, 4: 93-201. p/. 75-82. 1893-1894. 


Enumeration of several hundred species and descriptions of the following as new : 
Trifolium Virginicum, Rudbeckia Brittonti and Asterina Leemingtae. 


Bot. U. 


140 


Small, J. K. Some new hybrid Oaks from the Southern States. Bull. 
- Torr. Bot. Club, 22: 74-76. p/. 272-275. 26 F. 1895. 


Smith, E. F. Length of Vessels in Plants. Science (II.) 1: 77- 
18 Ja. 1895. 


Some notes culled from Strasburger’s “Bau und Verrichtungen der Leitungsbahnen 
in den Pflanzen.” 


Spalding, V. M. The Traumailopic Curvature of Roots. Ann. Bot. 
423-450. pi. 22. D. 1894. 

Spruce, R. Hepaticae Elliottianae, insulis Antillanis St. Vincintii et 
Dominica a cler, W. R. Elliott annis 1891-1892, lectae. Journ. 
Linn. Soc. 30: 331-372. pl. 20-70. 5 F. 1895. 


Twenty-six new species are described. 


Stitzenberger, E. Notes on Western Lichens. Erythea, 3: 39-3? 
1 F. 1895. ' 


Stubbs, W.C. Ramie (Bochmeria nivea). Uses, History, Composi- 
tion, Cultivation, etc. Bull. La. Exp. Sta, 32: 1126-1146. 1895- 


Thaxter, R. Notes on Laboulbeniaceae with Descriptions of new 
Species. Proc. Am. Acad. 30: 467-481. 1895. 
New species in Laboulbenia, Heimatomyces, Dichomyces, Eucantharouy¢ a 
(new genus), and Ceratomyces. 


Tilden, J. E. American Algae—Century I. 1894. 
Exsiccata of 100 species. 

Underwood, L. M. Notes on our Hepaticae—III. The Distribution 
of the North American Marchantiaceae. Bot. Gaz. 20: 59-7!- 16 
F. 1895. - 
Describes Asterella Pringlei from Mexico, and A. Austini and A. Wr ight 

from Cuba. 

Urban, I. Additamenta ad cognitionem florae Indiae occidentalis—H. 
Myrtaceae. Engler’s Bot. Jahrb. 19: 562-576. 28 D. 1894. 

Waite, M. B. The Pollination of Pear Flowers. Bull. Div. Veg: 
Path. U. S. Dept. Agric. 5: pp. 110. pl. 72. f. 5. 1894. 

Wallace, R. W.  Calochortus Plummerae. Garden 47: 8- 2: F. 
1895. 

West, Me and West, G. S. New American Algae. Journ. Bot. 33° 
52. F s808. : 


A new variety of Pediastrum duplex Meyen, and new species of Tetraedron and 
of Radiofilam, 


West, W. Some recently published Desmidieae. Journ. Bot. 33° 
65-70. Mr. 1895. 


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TORREY BOTANICAL CLUB. 


Notes on some Florida Plants, 


By Gro. V. NASH. 


The section of Florida visited by me, and where I spent some 
five or six months collecting the flora, had never before been sys- 
tematically explored botanically. One or two parties had col- 
lected for a few weeks in the winter season, but no one had spent 
any great length of time, and so it was practically a virgin field 
for the botanist. Its flora is most interesting, both in new and 
rare forms. The northern and tropical vegetation seem to overlap 
here. Ximenia Americana, a common West Indian shrub, and 
Mitchella repens, our common partridge berry, which has a far 
northern range, were equally common. This is a fair example of 
the extreme diversity of the forms. 

Eustis, a beautiful little town in the high pine land country, was 
the centre of my operations. This is situated on a lake of the 
Same name, some six miles long and three miles wide, connected 
with Lakes Griffin and Harris by the Ocklawaha River. It is 
about 25 miles from the Gulf, an equal distance from the Atlantic, 
and some 160 miles south of Jacksonville. I confined my opera- 
tions to a radius of 12 miles of this place. 

Lake county, a small section of which I explored, is situated in 
Central Peninsular Florida, right in the heart of the lake region. 
It seemed strange at first to find in a country where the soil is prac- 


tically nothing but sand, such a superabundance of lakes. They 


are everywhere. Five or six can be seen at once from the tops of 


142 


- 


some of the slight elevations in the high pine land. They vary 
in size from only a few hundred feet to several miles. There are 
two groups of them: first, the large ones, which are really widen- 
ings of the Ocklawaha River, and second, the small, clear-water 
lakes. These latter are never very large, and the water contained 
in them is as clear as crystal. These are particularly numerous 
in the high pine land. They have no apparent inlet or outlet, and 
as a rule occur in deep depressions. How it is that the water re- 
mains so perfectly clear and limpid is hard to explain. Compared 
with our northern lakes, they contain little vegetation. /otamo- 
getons, Vallisneria, and other forms so common in the lakes of 
the North, are entirely absent, and are to be found only in the 
Ocklawaha River and the large lakes alluded to above. 

Another peculiarity about these clear-water lakes is their varia- 
tion in elevation. The case of Lakes Alfred and Irma will illus- 
trate this feature. They lie a little to the southeast of Eustis, and 
are only separated by a steep hillside some 500 or 600 feet long, 
Lake Imra being at the top of the hill. The difference in eleva- 
tion is about 65 feet. One or two small streams, rising in this 
hillside, flow into Lake Alfred. But in spite of this its waters 
gradually lowered during the entire summer, and when I left in 
September the lake was nearly dry. The waters of Lake Irma, 
on the contrary, remained at the same level, although it is consid- 
erably higher and would be expected to drain into Lake Alfred. 

As stated before, the flora of this section is very rich, and this 
fact is due to the exceeding variation in the physical features of 
the surrounding country. There are five well-marked areas, the 
flora of each one being different, and having certain plants pecu- 
liar to itself. These might be designated as: High Pine Land, 
“ Scrub,” Low Pine Land, “ Bayheads’’ and « Hammocks.” 

Of these the high pine land is the greatest in extent. The 
tall timber is composed entirely of the long-leaved or yellow pine, 
Pinus palustris. The trees have perfectly straight trunks, rising 
to a height of 50 to 75 feet, the branches all being borne near the 
top, leaving the trunks entirely naked. The two other prevailing 
trees are Quercus Catesbaei and Q. cinerea, the shining bright 
green deeply cut leaves of the former making a strong contrast to 
the narrow entire and grayish-green foliage of Q. cinerea. The 


145 


Leguminosae are well represented, and in fact the members of 
this family predominate among the herbaceous plants. Chap- 
mania Floridana, Aeschynomene hispidula, Morongia angustata, 
Rhynchostia cinerea, Crotalaria Purshii, Cracca ambigua and C. chry- 
Sophylla are the ones met with everywhere. Among the more 
frequent of the Compositae are Berlandiera subacaulis, Pterocaulon 
undulatum, Cuicus spinosissinus Elliottii, Lygodesmia aphylla, Ver-" 
nonia angustifolia, Hieracium megacephalon, Helianthella grandi- 
Jlora and Liatris secunda. Among other plants characteristic of 
this region are: Tvadescantia rosea, Asclepias amplexicaulis, 
Asimina pygmaca, Commelina angustifolia, Polanisia tenuifolia, 
Breweria angustifolia, Stilingia sylvatica, Croton argyranthemus, 
Jatropha stimulosa, Eriogonum longifolium and E. tomentosum, Por- 
tulaca pilosa, Ceanothus microphyllus and Pirigueta Caroliniana. The 
grasses are represented by few species, but these occur in great 
numbers. Aristida stricta, Sporobolus juncea, Andropogon argy- 
vaeus and A. longiberbis are the commonest. There is only one 
member of the Cyperaceae that occurs in any quantity and that is 
Stenophyllus Waret. 

The high pine land is subject to annual fires. The cattle 
growers burn off the old grass, so that the roots will start up and 
give a crop of young, succulent shoots. It is wonderful what a 
variety of features protect the plants against these destructive 
fires. Pinus palustris has a very thick bark at the base of the 
trunk, and the thickening is very noticeable in the shape of a sud- 
den swelling just above the ground. All the herbaceous plants 
have large underground parts, tubers or thick roots, which 
store up nutriment and protect them. Commelina angustifolia 
has an immense fascicle of fleshy horizontally spreading roots, 
which anchor the plant deep in the ground and keep its 
soft, tender parts out of harm’s way. Tradescantia rosea has a 
similar arrangement. The provision for the preservation of Hfe/t- 
anthella grandiflora from extinction is even more noticeable. 
Buried in the ground, 6 or 8 inches deep, is an oblong tuber; 
the stem arising from this, instead of pushing abruptly upwards, 
continues horizontally, sometimes for a distance of 12 or 18 inches, 
and then rises obliquely to the surface; no matter how much the 
fire may damage the aerial portions, the plant has a supply of nu- 


144 


triment, enough to enable it to start up again atonce. The above 
are only a few instances of the many means which insure the 
preservation of the different species. 

The “scrub” flora is entirely different from that of the high 
pine land, hardly a single plant being common to both; in fact 
these two floras are natural enemies and appear to be constantly 
fighting each other. The only large tree growing in this region 
is Pinus clausa, which is markedly different from P. palustris, the 
pine of the high pine land. The latter is a tall, straight tree, 
with the branches restricted entirely to the top. P. causa, oh the 
contrary, is branched almost from the very base and does not at- 
tain a height of over 30 feet. Its needles are very fine, and it has 
received the name of the “spruce pine.” The great part of the 
growth in the “ scrub” is made up of scrub oaks, which are so var- 
iable that no two of them seem to be alike, and how many species 
there may be would be hard to tell. | Only a number of years of 
close study in the field could determine this. Cevatiola, with its 
heath-like leaves, occurs here very abundantly. Persea humilis, a 
most beautiful little shrub, makes itself very conspicuous by its 
bright brown silky pubescence, which is noticeable a long way off. 
Bumelia lanuginosa is very common, as is Ximenia Americana. 
Smilax Beyrichii climbs all over the scrub oaks, and is the only 
one of this genus observed growing in the “scrub.” One of the 
most beautiful plants of this part of Florida is Breweria grandt- 
flora, with its large, bright blue flowers. The stems, with their 
_ peculiar gray-green erect leaves, spread out in all directions over 
the white sand. I remember one place, several acres in extent, 
where this plant reached its perfection, the stems sometimes eight 
or nine feet long, fairly covering the ground in all directions, with 
hundreds of the flowers in sight. It is remarkable that a plant of 
such luxuriance and beauty can thrive in an area so devoid of 
moisture as the “scrub,” and yet it selects and confines itself ex- 
clusively to this region. Only one species of Cyperaceae is to be 
found here. Thisis Rynchospora dodecandra, and is quite common. 
Grasses there are absolutely none. For some reason this family 
is unable to obtain a foothold. Occasionally some member of the 
Andropogoneae tries it, but soon dies, not being able to stand the 
opposition it meets with for more than a few months. , 


145 


I spoke above of the antagonism of these two floras. This is 
so marked that there is no mistaking it. Wherever they come to- 
gether the line of division is very distinct. A bare space of pure | 
white sand usually separates the two. On one side you will see 
the tall Pinus palustris as far as the eye can reach, and on the 
other the diffusely branched P. clausa of much lower stature. 
You may look in vain in the “scrub” for plants occuring in abun- 
dance just over the line in the high pine land, and vice versa. 
The strip of bare white sand dividing the two is neutral ground, 
and each seems to jealously guard against the other's gaining a 
foothold there. The soil of these two sections was apparently 
originally the same pure white sand. That in the high pine land 
is now darker in color, being probably due to the charcoal de- 
posited there by the annual fires. This seems to be the only dif- 
ference. As fires are of rare occurrence in the “scrub,” the 
plants have made no provision against it, and so when a fire does go 
through it causes great havoc, almost entirely killing the pines 
and oaks. It is fortunate that fires are of such rare occurrence. 

The flora of the low pine land region is not so distinct. It 
grades into that of the high pine land. Each has its separate and 
peculiar plants, but the line of demarcation is not so evident. 
Pinus serotina and P. heterophylla are the prevailing trees. Some 
of the plants which are peculiar to and characteristic of this region 

re: ° Podostigma pedicellata, Trilisa odoratissima, Solidago Chap- 
mant, Bejaria racemosa, Cracca hispidula and C. spicata, Eryngium 
yuccaefolium synchaetum, Rhexia ciliosa and R. serrulata, Bletia 
verecunda, Limodorum parviflorum and Juncus marginatus pine- 
torum. The commonest member of the Cyperaceae is Fimbristylis 
puberula, which occurs in great abundance throughout the low 
pine land. Aristida stricta also occurs here, as well as in the high 
pine land. Andropogon Floridanus is quite common and.a very 
showy member of that genus. = 

The low pine land occupies an intermediate position between 
the high pine land and the flatwoods. The latter is not repre- 
sented near Eustis, and so I did not get a chance to do any col- 
lecting among that flora, which differs in a marked degree from 
that of any other section. 

The large swamps, lying generally along the low pine land, 


146 


‘have a peculiar flora and one quite interesting. These are locally 
known as ‘“‘bayheads,” so called, I presume, from the large number 
of bay trees, Magnolia Virginiana, that occur in them. The shrubs 
most prominent are fiers mitida and Leucothoe racemosa. 
Gordonia Lasianthus, with its large white showy flowers, occurs in 
quantity along the margins. It ranges in height from ten to thirty 
feet, and when in full bloom isa very pretty sight. The plant 
most common, and which attracts the eye above all others, is the 
ever prevailing Swlax laurifolia. It climbs and clambers over all 
the shrub; and bushes, and makes the “ bayheads” almost im- 
penetrable. Here and there open places will be found where 
sphagnum grows in quantity, and in such spots one is almost 
sure to find Utricularia fibrosa, with its large showy yellow flowers. 
And in the very wet places Peltandra sagittacfolia makes itself 
conspicuous by its showy ivory-white spathes. Amdropogon 
brachystachys occurs in the vicinity of “‘ bayheads” and I found it 
no where else. 

The soil of all the above, excepting the “ bayheads,” is pure 
sand, overlying a stratum of clay, the depth of sand varying from 
a few feet in the low pine land to as much as fifty feet in the high 
pine land. In places the clay comes to the top, and the character 
of the vegetation entirely changes. This forms what is locally 
known as ‘hammock land.” It isa heavy clay soil, and is gen- 
erally subjected to continuous inundation during the rainy season, 
which lasts from about June to September. Quercus virens, the 
live oak, is the principal tree, and often attains a great height. The 
most conspicuous object to the eye of a northerner is certainly the 
Cabbage Palmetto, Sabal Palmetto. It rises toa height of 50 to 75 
feet, overtopping all the other vegetation. These trees are very 
plentiful and can be found in all stages of growth, from the young 
plant only a few feet high to the tall ones just alluded to. Until 
they attain a considerable size they retain the bases of the petioles 
of the old leaves. This gives the young palms a different appear- 
ance from the one they assume later when they shed these, expos- 
ing the slim symmetrical trunk to view. So unlike do they seem 
that at first sight it is hardly possible to believe that they can be 
the same in different stages of growth. In some sections the 
Golden Fern, Polypodium aureum, as well as the Old Man’s Beard, 


hee 147 


Vittaria lineata, take possession of the trunks of the young palmet- 
tos, thrusting their roots down between the sheaths and the trunk. 
A palmetto thus adorned is certainly a beautiful object. 

The blue palmetto, Sadal Adansonit, also occurs in the “ ham- 
mocks.” It never attains a height of more than three or four feet. 
Among the other plants restricted to the hammock lands are: Zra- 
descantia pilosa, Coreopsis Leavenworth, Solidago Leavenworthii, 
Flelenium nudiflorum, Teucrium Nashii, Berchemia scandens, Cardio- 

Spermum microcarpum, Eryngium Baldwinii, Ulmus Floridana and 
U. alata, Celtis Mississippiensis and Capsicum baccatum. 

Owing to this exceeding variation in the physical conditions 
of the country explored, I was enabled to secure a large number of 
species, some 800, several of them being new to science and 
others additions to the flora of the United States. The range of sev- 
eral plants was extended much further south. I also secured a 
number of rare things hitherto poorly represented in herbaria. 
The following are some of those of peculiar interest : 

1153, 1180. CASTALIA RENIFORMIS (Walt). [Mymphaea rentformis 

Walt.) 

This was found growing commonly in Lake Ella. There 
seem to be two forms of it, one growing along the shore, the 
other in water 10 to 20 feet deep. The two merged into each 
other, and a careful comparison showed them to be the same. 
The rootstocks of the shore form were secured and there was no 
indication of any tubers. The leaves varied from 6 to 8 inches in 
diameter in the shore form, to nearly 2 feet in the one growing in 
deep water. The edges of the latter were distinctly turned up, 
much as in the Victoria Amazonica and occurred in large masses, 
making it difficult to propel a boat through them. The flowers 
were long and entirely without odor, with the exception of a faint 
Suggestion of that given off by apples. 

815. Eler1anrHemum Nasui Britton, n. sp. 

Diffusely branched from a thick woody, horizontal root, the 
branches decumbent or ascending, slender, terete, densely stellate- 
tomentose even when old, leafy, 2-4 dm. long. Leaves oblong or 
linear-oblong, densely stellate-canescent on both sides, ste at 
both ends or the lower obtuse at the apex, I1.5-3 cm. long, 


3-6 mm. wide, short-petioled, the margins somewhat revolute ; 
flowers all alike in terminal leafy-bracted thyrsi; pedicels 2-5 mm. 


148 


long, divergent and ascending; outer sepals subulate ; inner sepals 
oval-oblong, very obtuse, firm, concave, 3-4 mm. long; petals _ 
yellow, broadly cuneate, slightly eroded, 5 mm. long; stamens 
about 15, shorter than the petals; ovary globose, puberulent. 


Nearest to 7. arenicola Chapm., but very different in its thyr- 
soid, not corymbose, inflorescense, and obtuse inner sepals. It 
occurs in the “scrub.” 


14. STIPULICIDA FILIFORMIS n. sp. 


A diffusely branched glabrous annual, of very slender habit, 
1-2 dm. high. Radical leaves orbicular, acute, about 4 mm. 1 
diameter, on slender petioles 5-7 mm. long; bracts triangular- 
subulate, about 1 mm. long; inflorescence capitate, 1~2-flowered ; 
flowers about 1.5 mm. Jong, sessile or slightly stalked; sepals 5, 

scarious-margined, oval, obtuse, the inner 1.5 mm. long, I mm. 
broad, outer about two-thirds as long ; petals 5, oblong, 2 mm. long, 
-75 mm. broad, entire or somewhat eroded at apex, minutely 3-5- 
toothed on each side near the base; stamens 3, opposite inner 
sepals; filaments linear, .75 mm. long, .25 mm. broad; anthers 
oval, .5 mm. long, .3 mm. wide; capsule globose, obscurely 3- 
angled; seeds more or less triangular in outline, flattened, less 
than .5 mm. long. 


Differs from S. sefacea in being much more slender; in the 1 
florescence, which is composed of fewer and sessile flowers, and 
in the shorter bracts. ; 

One of the few plants that grow in both the “scrub” and high 
pine land. It prefers the latter, and attains a much more luxuriant 


growth there. Begins to flower in March and continues through- 
out the summer. 


755. Hypericum aspalathoides Willd. 


This has been considered as a variety of //. fascicularts. The 
two occur very commonly around Eustis and are clearly distinct 
species. The latter is found invariably around the lake shores 
and grows from 4 to 12 feet high, sometimes forming a small ee 
with a trunk 2 to 3 inches in diameter. The leaves are long and it 
flowers about the middle of April, maturing its fruit early in hae 
gust. H. aspalathoides never occurs along the lake shores, but in 
the low pine land. It is a small undershrub, 1 to 3 feet high, has 
much shorter leaves, does not flower until five or six weeks later, 
and matures its fruit sometime in September. 


149 


673. Hibiscus incanus Wendl. 

This has been put*in with A. Moscheutos but it certainly ap- 
pears distinct from that plant. Its flowers are always pure white 
with a deep purple centre. The leaves are softly pubescent on 
both sides, and are very seldom lobed at the apex, and when they 
are the lobing it is very slight. 


378. Cardiospermun microcarpum Kunth. 

Of frequent occurrence in the “ hammock lands,” climbing up 
the bushes and small shrubs. It appears to be new to the flora of 
the United States. It occurs in the West Indies, and so its turn- 
ing up in Florida is not a matter of great surprise. It has much 
smaller fruit than C. Halicacabum L., which is very common in 
South Florida. , 


261. Amorpha virgata Small. 
Mr. Small found this on Stone Mountain, Georgia, and it has 
now been found in Florida, several hundred miles further south. 


1523.” KuHNISTERA Fray (Chapm.) (Fetalostemon Feayt Chapm.). 
This occurs very commonly in both the “scrub” and high 

Pine land. It forms large clumps 3 to 4 feet across. The flowers 

are a bright lavender. A white-flowered form was also found 


(No. 1524). 


1336. RHYNCHOSIA CINEREA N. sp. 

Perennial. Stems numerous, trailing, branching, 6-10 dm. 
long, 3-angled, striate, clothed with appressed cinereous pubes- 
cence; stipules brown, ovate, acuminate, 3-4 mm. long; leaves 
from 3-5 cm. long, 3-foliolate, terminal leaflet largest; petiole 
11-20 mm. long, rachis 5-9 mm; leaflets appressed-pubescent 
above and on the veins beneath, somewhat rugose; lateral ones 
unequally ovate or orbicular-ovate, 13-23 mm. long, 9-18 mm, 
wide ; terminal leaflet larger, orbicular-ovate to depressed-orbicu- 
lar, 15-26 mm. long, 13-30 mm. wide; petiolules about I mm., 
covered with a dense cinereous, appressed pubescence; flower- 
clusters about as long as petiole ; peduncle 3-5-flowered ; pedicels 
slender, 1-3 cm. long ; calyx somewhat 2-lipped ; lower lip 3-parted, 
divisions lanceolate, acuminate, middle one 13 mm. long, a little 
exceeding the lateral ones; upper lip 2-cleft, teeth acuminate; 
Ovary pubescent; pod oblong, obliquely acute at apex, 16 mm. 
long, 8 mm. broad, appressed-pubescent, 2-seeded ; seed orbicular, 
flattened, 5 mm. in diameter, mottled brown. 

Nearest to R. somentosa Ell., which has a climbing habit, larger 


150 


leaflets, a yellowish brown spreading pubescence, and the young 
pod densely pubescent, almost villous. In &. cinerea the pod is 
sparingly appressed-pubescent. Grows only in the high pine land 
region. 

70. Rubus cunetfolius Pursh.? 

There is some doubt as to this being this species, but more ma- 
terial is needed to decide the matter. It grows strictly erect and 
is armed with most aggressive spines. The leaves are much 
larger and the pubescence seems to differ. There is a specimen in 
the Columbia College Herbarium collected by Mr. A. H. Curtiss, 
No. 791, which appears to be the same thing. 


1142.’ CRATAEGUS FLAVA INTEGRA Nn. var. 


A small tree 4-5 metres high with spreading branches. Bark 
light gray; on new shoots reddish-brown ; leaves obovate, more or 
less undulate especially at the apex, glandular on the margin, acu- 
minate at base and narrowed into a slender glandular petiole, 
10-13 mm. long. Fruit (not fully ripe) globose, on villous ass 
cels, 5-15 mm. long. 

. Collected in vicinity of Lake Ella in old fields, July 2. - The 


fruit was green though apparently full grown. The flowers were 
not seen. : 


948. Dresera capillaris Poir. 

This plant occurs in great abundance along the shores of the 
clear-water lakes, and seems to be confined to them. It never 
grows more than 2 or 3 inches high. There is another form 
which is also quite common, but it is much taller. 


1218. RHEXIA FLoRIDANA n. sp. 


Whole plant of a dark olive green, diffusely branched from a 
woody base, 2-4 dm. high, branches ascending. Pubescence 
glandular-hirsute, spreading; leaves linear, 1.5-4 cm. long, I-3 
mm. wide, smooth on both sides, one-nerved, remotely and 
sharply spinulose-serrulate, acute, sessile, narrowed at base; 
flowers short-pediceled; calyx-tube cylindric-campanulate, spar- 
ingly glandular-hirsute, about 1 cm. long; lobes triangular to 
ovate-lanceolate, about 2 mm. long, I mm. broad at base; petals 
purple, mucronate, mucro about 1 mm. long; fruiting calyx 13 
mm. long, neck 6 mm. long, 3 mm. in diameter, globose portion 
about 6.5 mm. in diameter ; seeds snail-shaped, .6—.75 mm. long, 
irregularly ridged. 


Very distinct and well marked. Its dark color and very nar- 


151 


row leaves readily distinguish it from all the other forms of this 
genus, 
Found growing in quantity in wet clay soil on east bank of 
canal leading from Hicks’ Prairie, near Eustis. Collected early 
in July. 
515. Ginothera sinuata L. (?) 
The stems of this plant are prostrate and 3 or 4 feet long, 
with leaves nearly entire or but slightly toothed. It appears quite 
different from the true plant, which is very common about Eustis. 


865. Melothria pendula L. (?) 

There are two forms of this occurring commonly. One, the 
ordinary plant, climbs over the bushes and tall weeds, and has 
thin sharply angled leaves with a broad open sinus at base. The 
other is always prostrate, and has thicker and more obtusely lobed 
leaves, with the sinus at base almost closed. 


377. Sambucus Canadensis L. 

This often attains a height of 10 to 15 feet, with a trunk 6 to 8 
inches in diameter. Its leaves are very dark green and shining 
above, with long acumination. 


892. Cephalanthus occidentalis L. 

In the cypress swamps this often reaches a height of 30 feet, 
- with a trunk 5 to6inchesin diameter. The leaves are also corres- 
pondingly large, some of those measured being over 17 inches 
long. 

730. Richardia Brasiliensis Gomez. 

This appears to be new to the United States. It is evidently 
introduced, as it occurred only in one place along the railroad 
track, but it appeared at home and well established. 

729, 1346. Diodia hirsuta Pursh. 

Why this plant should have been put in with D. Virginiana is 
hard to understand. The mistake would certainly not have been 
made had the two been seen growing. They are entirely differ- 
ent. The latter is a glabrous plant with large thick leaves. D. 
hirsuta is very hairy with much smaller leaves and larger fruit. 
It occurs in dry sandy soil or in open swamps. In the former 

situation the stems are prostrate, forming mats on the ground. 


s 


In the latter they are only decumbent at base, being erect for the 
last 10 or 12 inches. 


1183. Garberia fruticosa A. Gray. 


Only a few specimens of this very rare shrub were secured. 
The plant seemed to be very common, but the flowers were 
scarce. It probably blooms much later. The shrub is some 4 or 
5 feet high. 


691. Solidago Leavenworth T. & G. 


This rare golden-rod is by no means common. It occurred in 
only one or two places. 


1225. Helianthclla grandiflora A. Gravy. 


The tubers of this showy composite exude a copious resinous 
matter. 


1711. Liatris secunda Ell. 


This is the L. pauciflora of the Synoptical Flora where the above 
name is cited as a synonym. It is clearly not the plant Pursh 
had in view; he divides the genus into two sections, tuberous 
and non-tuberous. The first contains the species now placed in 
Liatris, the second is composed of a number of plants, part of 
which are now put in Carphephcrus, the others in 7ivilisa. L. 
paucifiora occurs in this latter group. The description given of 
it is clearly not that of a Zzatris, as now understood. 


390.” HIERACIUM MEGACEPHALON nN. sp. 


Annual, whole plant generally of a purplish hue, ste,,s 3-5 
dm. high, channeled, strongly pilose especially at the base. Radi- 
cal leaves broadly oblanceolate, 8-12 cm. long, 2-3 cm. wide, den- 
ticulate, pilose, particularly on the upper surface and the midrib 
beneath; cauline leaves oblong to oblong-ovate, decreasing in 
size towards the inflorescense, lower 4-6 cm. long, 1.5-2 cm. 
broad, truncate at base, sessile; inflorescence corymbose-panicu- 
late, densely glandular-pubescent; involucre 8-10 mm. long; 
achenes fusiform. 

Grows only in the high pine land. Flowers from March—May. 

This seems to belong near H. Gronovii, and the achene 
certainly places it near that species. Its inflorescence, large 
heads, and dense glandular-pubescence clearly separate it from 
that plant. Moreover, H. Gronovii does not begin to flower in 


153 


Eustis until late in August, whereas this plant flowers early in the 
spring. 
700. “XOLISMA FRUTICOSA (Michx.) 

(Andromeda ferruginea fruticosa Michx.) This certainly is 
specifically distinct from Xolisma ferruginea, which grows from 
6-12 feet high, has light green leaves which are very much rolled 
in on the margins, and flowers early in March. X. fruticosa never 
attains a height of more than 6 feet, has a much stricter habit, 
very dark leaves which are not revolute, and does not flower until 
two months later. This difference is quite marked in herbarium 
specimens, but in the field there can be no mistaking it. 


573. Vaccinium stamineum L.? 

This occurs in very dry sandy soil, and has much smaller 
leaves than usual. They are thick, somewhat rugose and of a 
light green color. The true plant occurs in similar situations, but 
is much larger and thinner leaved. 


941, 1698. Fraxinus epiptera, Michx. 

-In the herbarium of Columbia College there are two things 
placed under /. platycarpa. One of these has fruit over half an 
inch wide, elliptical in outline, gradually narrowed at each end. 
The other has linear-oblong fruit” This latter I found in quan- 
tity in the cypress swamps and it is evidently distinct from the 
broad-fruited form. Its leaves are thicker and entire, or occasion- 
ally with very obscure serrauion. It grows from 20-50 feet high 
and fits very well the description of the F. efiptera of Michx.,and I 
have called it that. The broad-fruited one is F. platycarpa. 


796. Asclepias Feayi Chapm. 
This was obtained in but one place, in the low pine land. Its 

pure white flowers, large in proportion to the size of the plant, 

make it very conspicuous. 

419. Asclepias decumbens L. 

This is very common in the high pine land country. Although 
it is hard to detect any difference between the flowers of this 
species and A. suderosa, in the fruit the characters seem to be 
good. The latter has a pod about 3 inches long and 34 of an inch 
wide, while in A. decumbens the pod is considerably longer than 
A 


iAim 


: ne x Th con tk, afforeS hacks ( bans CAL AAA. > eee cat 


154 


this and hardly 4 an inch broad. The relative difference between 
length and breadth is very marked. 


1092. “ASCLEPIAS ACERATOIDES Nn. sp. 


-Perennial. Stems decumbent or erect, dark purple, simple or 
branched, terete, striate and puberulent; leaves smooth on both 
sides, those of the stem nearly orbicular or very broadly oblong, 
about 5 cm. long, 3.5-4 cm. broad, emarginate, rounded at base, 
on petioles about 8 mm. long; those of the branches oblong, 3-3-5 
cm. long, 1-2 cm. broad, apiculate, more or less acute at base, on 
petioles 2-4 mm. long; umbels borne in the axils of the upper 
leaves, 20-50-flowered; peduncles puberulent, 2-3 cm. long; 
pedicels slender, puberulent, about 1.5 cm. long; sepals greenish, 
lanceolate, reflexed, acute, 2.5 cm. long; petals greenish, ob- 
long-lanceolate, reflexed, obtuse, 6 mm. long, 2 mm. wide; lobes 
of the crown white with a greenish keel, flattened, semi-lanceo- 
late, 5 mm. long, acute at each end, short-stalked; horn protrud- 
ing from the middle of the lobe, horizontal, resting on the column, 
tip more or less ascending ; column sessile, 2 mm. high ; pod erect 
or ascending, puberulent, lanceolate, obtusely acuminate, about 8 
cm. long, 1.5 cm. wide, on a twisted and reflexed thickened 
pedicel. 

This. plant very much resembles an Acerates. It grows ex- 
clusively in the “scrub,” flowering late in June and through July. 
1715. Hydrolea corymbosa Fl. 

Occurs in swamps and is very scarce. I searched for it 
carefully, but failed to secure more than a few specimens. It 
grows in the tall grass and might be easily overlooked. 

609. Convolvulus repens L? : 

This differs from the ordinary plant in having pure white 
flowers, the lobes of the leaves decidedly divaricate-spreading, the 


pubescence stronger, and in a distinct climbing habit. It may be 
only an extreme form. 


Zz 
770.” BREWERIA VILLOSA Nn. sp. 


_ Whole plant villous-pubescent. Stems several from a peren- 
nial root, simple or much branched, trailing; leaves oblong to 
oblong-ovate, 2.5-7 cm. long, 7-20 mm. broad, obtuse, apiculate, 
rounded at base, on petioles 2-8 mm. long; peduncles equalling 
or exceeding the leaves, 1-7-flowered; bracts shorter than the 
pedicels; sepals oblong to ovate-lanceolate, 8-11 mm. long, 3-4 
mm. broad, acuminate, villous; corolla 1.5-2 cm. long, white; 
filaments filiform-subulate, two-thirds as long as corolla, adnate 
and villous for two-fifths their length, lower part of free portion 


155 


pubescent; anthers 1.5 mm. long; ovary villous, 2-celled, cells 2- 
ovuled ; styles 2, adhering for one-third their length; capsules on 
erect pedicels, ovate, villous at apex, 2-celled, cells 1-seeded; 
seeds yellowish. brown, ovate, compressed, flat on one side, 5 mm. 
long, 3 mm. broad. 


Found in low lying oak land in the vicinity of water. Near- 
est to B. aquatica but readily distinguished by its larger white 
corolla, villous filaments and adhering styles. 

Nos. 771 and 1508 appear to be the same. They have larger 
and thinner leaves and are not so villous. They grew in more 
shaded situations, and this probably accounts for the variation 
in the leaves and pubescence. In No. 770 the peduncles almost 
invariably exceed the leaves and are many flowered. In Nos. 771 
and 1508 the leaves and peduncles are nearly equal, and the 
latter often only 1-flowered. : 


O71," BREWERIA ANGUSTIFOLIA Nn. sp. 


Stems numerous, from a perennial root, trailing, slender, 8-10 
dm. long, more or less depressed-pubescent. Leaves narrowly 
linear, 2.5—7 cm. long, 1.5—3.5 mm. broad, glabrous or pubescent, 
acute, sessile or on petioles not exceeding 2 mm. in length; pe- 
duncles slender, 1-flowered, about equaling the leaves, appressed- 
pubescent; bracts shorter than the pedicels; sepals elliptical to 
oblong, acute, smooth, ciliate, 7-10 mm. long, 2.5-3 mm. broad ; 
corolla white, about 2 cm. long; filaments villous with jointed 
hairs, about one-half as long as coralla, adnate for one-third their 
length ; styles 2, three-fourths as long as coralla, adnate for one- 
third their length ; ovary villous at apex; capsules 8 mm. long, on 
recurved pedicels, oval, acute, with a tuft of hairs at the apex, 2- 
celled, cells 1-seeded. 

Grows only in the high pine land region, where it is very 
abundant. It has been confused with B. Aumistrata and B. Pick- 
eringit, from both of which it is evidently distinct. It differs from 
the former in its larger flowers, very narrow leaves, invariably 
I-flowered peduncles, and 2-seeded capsule. From the latter it 
can be separated by its much shorter bracts, villous filaments, 
styles adnate for a much shorter distance, and like the stamens, 
included. 

1299. Utricularia resupinata B. D. Greene. 

Occurs very plentifully along the shores of several of the 

clear-water lakes. A few specimens of it were secured by Dr. 


156 


Garber and Dr. Porter was the first to note the extension of its 
range into-Florida. It seems to have been reported from no in- 
termediate stations. 


248. Utriculana oligosperma St. Hil. 

This is the largest of its genus occurring in Florida. It is found 
in both the rivers and lakes, and is quite common. It reaches 
its perfection in Lake Ella, where the stems often attain a length of 
6 to 8 feet and the branches spread 2 to 3 feet. The growing end 
is of a beautiful pink color, the middle portion a dark green, and the 
decaying end a rich brown. It is entirely floating, forming a 
beautiful object spread out in the water, and looks like a large 
sea-weed. Its flowers are an inch in diameter and yellow. It is 
found in South America, and the extension of the range to cen- 
tral Florida, where it is evidently indigenous, is remarkable. On 
a specimen of a Utricularia in the Columbia College Herba- 
rium, collected by Dr. Chapman, is a note to the effect’that the 
plant appeared in the bay at Apalachicola in 1842, but entirely 
disappeared in a short time. This specimen proves to be this 
same thing. 

974. Boerhaavia decumbens Vahi. 

This plant was very plentiful, much more common than the 
other species growing with it, B. erecta. It always occurred in 
cultivated ground or its vicinity and may be introduced. I can- 
not find that it has been reported from the United States before. 
1185. Paronychia herniariodes (Michx.) Nutt. 

This rare plant was found growing quite freely in one locality 
near Tavares. It’occurred in dry sandy soil just west of the rail- 
road bridge crossing the Ocklawaha River. It had been pre- 
viously known only from Georgia and North Carolina. 

435. Persea pubescens (Pursh) Sargent. 

Dr. Chapman published this as a variety of P. Carolinensis. It 
has several well-marked characters which seem sufficient to jus- 
tify its elevation to specific rank. In P. Borbonia (L.) Spreng. 
(P. Carolinensis Nees) the leaves and inflorescence are glabrous, the 
peduncles equaling or shorter than the petioles, and fruit fully a 
half inch indiameter. In P. pudescens the Jeaves and inflorescence 
are strongly pubescent, the peduncles generally much exceed 


157 


the petioles, and the fruit is but little more than half as large, of a 
light blue color. 


574.’ PERSEA HUMILIS n. sp. 

A compact shrub, 2-3 metres high, with branchlets, young 
leaves, and whole inflorescence covered with a dense bright brown 
sericeous pubescence. Branches dark slate-color; leaves from el- 
liptical to oblong, 4.5-8.5 cm. long, 2~3 cm. wide, more or less 
revolute on the margins, acuminate at both ends, obtuse at apex, 
yellowish green, smooth and slightly shining above, blackish and 
pubescent beneath, the midrib very prominent; petioles from 1-2 
cm. long; inflorescence capitate ; peduncles stout, 4-7 mm. long, 
generally 3-flowered; flowers about 6 mm. long; sepals erect, obtuse, 
outer ones oval, 2 mm. long, shorter than the stamens, inner oblong, 
5 mm. long, exceeding the stamens; fruit purplish black with a 
bloom, globose, 15 mm. in diameter, on peduncles about 15 mm. 
long ; seed globose, 11 mm. in diameter. 

This seems to be nearest to P. Borbonia(L.) Spreng. It differs 
in its smaller size, shorter, stouter and fewer-flowered peduncles 
which elongate in fruit; and especially in the dense brown seri- 
ceous pubescence of the young leaves and branchlets. This latter 
feature makes the shrub very conspicuous, and seems to be unusual 
in the genus. This, moreover, flowers some five weeks later than 
P. Borbonia. ; 

Occurs exclusively in the “scrub,” where it is very common. 
Collected at Eustis in flower in May. Fruit kindly collected on 
November 8th at the same place and sent to me by Mr. W. T. 
Swingle. 


1397a.’CROTONOPSIS SPINOSA N. sp. 

An erect much branched annual, 4-7 dm. high, clothed with 
stellate-pubescence. Branches erect; leaves linear to linear-ob- 
long, 1.5-3 cm. long, 2-5 mm. broad, acute at both ends, on 
petioles 1-2 mm. long; fruit spiny; seeds ovoid to ellipsoid, 
only slightly flattened, 2 mm. long, 1.5 mm. broad, minutely 
pitted. : 

Differs from C. dinearis in having a more strict habit, smaller and 
much less flattened seeds, narrower leaves, and fruit covered with 
spines. 

Collected by Mr. W. T. Swingle at Dunellon, July 24. 


1700. Habenaria blephariglottis (Willd.) Torr. 
Evidently an abnormal form of this plant. Its parts are nearly 


158 


twice as large as usual. Some of the flowers are slightly tinged 
with orange. 


6. LimoporuM PARVIFLORUM (Lindl.) (Calopogon parviflorus Lindl.) 
Quite common in the low pine land. It grows from 6 to 12 
inches high. Its flowers are deep purple, much resembling those 
of L. tuberosum of the north, which is also common, but flowers 
much later. L. parviflorum was in full bloom early in March. 


557,” STENORRHYNCHUS Jaiscana (Watson.) (Spzranthes Jaliscana 

Watson.) 

Finding this plant in Florida was a surprise. It-was previously 
known only from Mexico. 

857, 1703. Tillandsia juncea Le Conte. 

This has been placed with 7? setacea Sw. by Mr. Baker, in 
his monograph of the Bromeliaceae. The two are very common 
where I collected and are clearly distinct. 7: se¢acea has erect 
setaceous leaves, very narrow at the base while in 7: juncea the 
leaves are much stouter, with a much broader base, and are 
conspicuously recurved, The specimens in the Columbia College 
herbarium collected by Dr. Chapman and so named are'evidently 
another species, the 7: Balbisiana R. & S. Le Conte says [Ann. 
Lyc. N. Y. 2: 131 (1828)] that his species very much resembles the 
figure of Bonapartea juncea in Flora Peruviana, p/. 262. The 
plant referred to above certainly does bear a strong resemblance 
to this figure. | 
1562. Yucca filamentosa L. var. 

There are two forms of Y. filamentosa occurring in this region. 
One flowers in May, has conspicuously twisted leaves which gener- 
ally are strongly recurved and rather short, and matures its fruit in 
July.. The other is taller, has whiter flowers in a larger panicle, 
longer and more erect leaves which are narrower in proportion, 
and does not flower until August. They grow in similar sit- 
uations. With these differences it hardly seems possible that 
they can be the same. 

459. “Nouina BRITTONIANA n. sp. 


Roots a cluster of long spreading fleshy fibres. Radical 
leaves 40-70, recurved and prostrate, forming a dense mass around 
the base of the plant, linear, 1-2 metres long, s-1o mm. wide, 
strongly scabrous on the margins, attentuated into a long slender 


159 


point; cauline leaves setaceous, 1-3 dm. long, about I cm. broad 
at base; stem arising from a bulbous base, 1-2 metres high; in- 
florescence paniculate, ovoid to oblong in outline, 3-4 dm. long ; 
flowers in clusters of 2 or 3 on erect pedicels 3-5 mm. long ; peri- 
. anth segments elliptical to elliptical-ovate, 4 mm. loag, 2 mm. 
wide, glandular at apex; filaments flat, broadly subulate, 2 mm. 
long, .5 mm. broad; anthers 1.5 mm. long ; cells oblong, .75 mm. 
broad; ovary 3-celled, orbicular-ovoid, truncate, somewhat trique- 
trous; stigmas sessile; capsule orbicular, on slender recurved 
pedicels about I cm. in length, 8-9 mm. long, 10 mm. broad, 
cordate, retuse, triquetrous, 2 of the cells 1-seeded, the other gen- 
erally empty. 

Grows in dry, sandy soil in the high pine land region. Some- 
times five or six plants occur together, and then it makes a very 
showy appearance in flower. 

This plant is named in honor of Dr. N. L. Britton who has 
given me much help and advice in working up my collections, and 


to whom I wish to extend my thanks for his many kindnesses. 
1389. Xyris platylepis Chapm. . 

This is found in the low pine land region and occurs with either 
white or yellow flowers (No. 1377.), the former by far the more 
frequent. 


954. Ayris torta Sm. 

This is the commonest species, inhabiting the low pine land. 
It invariably has white flowers, a fact which does not seem to have 
been noted before. 


1584. Xyris SMALLIANA n. sp. . 

Plant smooth, glabrous and shining throughout. Leaves about 
one-half as long as scape, linear, 3.5-6 dm. long, 3-9 mm. wide, 
striate, the lower half purple; sheath from 2.5—4 dm. long, striate, 
with a blade from 1-10 cm. long; scape erect, scarcely striate, 
7-10 dm. tall, nearly terete, 1-edged throughout, flattened and 2- 
edged just below the head, one edge being much more prominent, 
heads ellipsoidal, about 2 cm. long, 1 cm. in diameter when ma- 
ture, ovoid in the early flowering state; scales orbicular-obovate, 
entire, 5-6 mm. long, 3-5 mm. wide; lateral sepals conspicuously 
exserted, 8 mm. long, 1 mm. wide, curved, wing gradually broad- 
€ning to the top where it is from one-half to two-thirds as wide as 
the sepal, lacerate-fimbriate for the upper third; petals yellow; 
capsule obovoid, obscurely triquetrous, placentae parietal; seeds 
from cylindric-oblong to cylindric-obovoid, more or less curved, 
pointed at both ends, striate, about .6 mm. long, .2 mm. wide. 


160 + 


Growing in shallow water at Lake Ella and in very wet soil 
along edge of cypress swamp north of Trout Lake, near Eustis. 
No. 539. X. zorta and X. fimbriata are the only other species in the 
United States with exserted sepals, the former with white petals. 


a3 
1525. COMMELINA SWINGLEANA N. Sp. 


Stems 1-3 dm. high, smooth, sparingly branched, prostrate or 
ascending, from a compact fascicle of descending fleshy roots 3-5 
cm. long; sheaths inflated, 6-9 mm. long; leaves linear to linear- 
lanceolate, 4-6 cm. long, 5~7 mm. wide, glabrous or slightly pu- 
bescent; peduncles about 5 mm. long, pubescent; spathe more or 
less pubescent, 2 cm. long, I cm. broad, falcate-acuminate, 
rounded posteriorly ; the two large petals deep blue, reniform, 12 
mm. long, 17 mm. broad, claw 3-4 mm. long; capsule depressed- 
globose, 5 mm. long. 4 mm. broad, obscurely 3-lobed, 3-celled,. 
cells 1-seeded; seed broadly oval in outline, somewhat com- 
pressed, flat on one side, smooth, dark-brown. 


The stems of this plant are generally prostrate, about 1 dm. 
long ; occasionally one is found with longer and ascending stems, 
but they are very rare. Collected in the high pine land region at 
Umatilla early in August. 

There are two specimens in the Cofuiiibis College herbarium 
which evidently belong here, one collected by Dr. vce ec and 
the other by Mr. Blodgett at Key West. 

Near to C. hamipila Sauvalle, of the West Indies. That is dif- 
fusely branched; has a smaller seed, gray in color, narrower and 
longer leaves, longer sheaths, not inflated, and an ovoid capsule. ~ 

Named in honor of Mr.- W. T. Swingle, Director of the Sub- 
Tropical Laboratory at Eustis, to whom I wish to extend my 
thanks for the many favors shown me. His knowledge of the 
country and flora, which were freely imparted, enabled me to make 
much more valuable and extensive collections than would have 
otherwise been possible. 


1015. Peltandra sagittaefolia (Michx.) Morong. 

Occurs in the bayheads, generally among sphagnum. It is by 
no Means common. 
148. Lachnocaulon Beyrichianum Sporleder. 

This is another rare plant, but very common around Eustis, 


along the shores of the clear water lakes. It grows in small tufts, 
generally among tall grass. 


161 


1295. Lachnocaulon Beyrichianum Sporleder ? 
Appears to be the same as No. 148, but the heads are much 
larger than the original description calls for. 


1407. Cyperus leucolepis Carey. 

This rare plant was previously only known from a single col-_ 
lection. I found it growing quite freely in one place at Eustis. 
Its silvery white scales make it very conspicuous among the tall 
grass. 


“ STENOPHYLLUS FLORIDANUS Britton, n. sp. 

Similar to S. capillaris, culms filiform, erect, 10 cm. to 20 cm. 
tall, much exceeding the thread-like leaves. Sheaths ciliate; 
spikelets linear, acute, several-flowered, rich brown, 4 mm. to 8 
mm. long, I mm. wide, sessile and densely capitate in 4s to 10s at 
the summit; scales ovate-lanceolate, keeled, about I mm. long, 
with a minute slightly spreading tip; achene obovoid, 3-angled, 
light-colored, slightly longer than thick, the angles rounded, the 
faces somewhat concave, the whole surface cellular-reticulated, its 
cells oblong; style glabrous, 3-cleft for about one-fourth its length. 


Dry sandy soil, high pine land. 


566. Setaria flava Kunth. 
This appears to be new to the United States. 


1382. Setaria Ventenati Kunth. 
Also an addition to the flora of the United States. 


831. Marsilia vestita H. & G. 
This was found growing in abundance at Orange Bend, the 


same place from which Prof. Underwood reported it. The plant 
occurs along the track on both sides of the depot for about one- 
quarter ofa mile. Itis confined to that limited area so far as I could 
find out. Its occurrence at such a distance from its ordinary range — 
and its limitation to this small section point very strongly to its 


being introduced. 


162 


John H, Redfield. 
By Wm. M. CAnsy. 


(WITH PORTRAIT.) 


On the banks of the beautiful Connecticut and near the center 
of the State of the same name is to be found the place anciently 
and still called Middletown; and, in accordance with a custom 
nowhere so common as in New England, of retaining for offshoots 
from the original settlements the name of the mother town with a 
prefix or a suffix, the little hamlet, a few miles up the river, was 
of old called by the somewhat picturesque name of “Middletown 
Upper Houses,” now, alas! changed to the unmeaning one of 
Cromwell. Here,on July 10, 1815, the subject of this sketch was 
born. He could claim John and Priscilla Aiden among his an- 
cestors and was in every way of pure New England blood. Many 
of his family had been sea captains, a vocation nowhere repre- 
sented by more honorable, hardy and vigorous men than on our 
northern coast. His father, William C. Redfield, at this time a 
country storekeeper in humble circumstances, was a man of enter- 
prising character and of an unusually inquiring and vigorous mind. 
The son only knew his mother as an invalid and she died when 
he was but four years old; and although his father married after- 
ward, he was again bereaved; so that his son owed much of his 
good bringing-up to a widowed relative who came to take charge 
of the household and who, according to the custom of those days, 
did not stint the lessons to be derived from the “ New England 
Primer” and the “Shorter Catechism.” Other lessons, more 
pleasant perhaps, came to him early from his father and served to 
stimulate his inherited scientific tastes. 

The following pleasant account is of one of these which oc- 
curred when he was six years old during a long ride taken soon 
after the storm long known as the “Great September Gale.” * 
“My father’s habits of close observation led him to watch the 
fallen trees and the effects of that destructive wind. At Middle- 
town the wind had been from the southeast and the trees lay 


* See Oliver Wendell Holmes’ poem “The September Gale.” 


163 


with their heads northwestward; but on reaching Berkshire he 
was surprised to see that they lay in an opposite direction and he 
repeatedly called my attention to the fact. In conversing -with 
the residents of that region as to the time these trees were pros- 
‘trated he was still more astonished to learn that the wind, which © 
at 9 p. M. had been from the southeast at Middletown, had been 
at Stockbridge from the northwest precisely at the same hour. 
* * * * It did not appear to him possible that two winds of 
such violence should be blowing against each other at the dis- 
tance of only seventy miles. The only explanation of this para- 
doxical phenomenon was one which he was then led to accept 
hypothetically, but which he afterwards confirmed by years of ob- 
servation and innumerable facts.” It was thus that the elder Red- 
field was led to the theory of the rotary as well as progressive 
movement of storms which procured him so much note as a 
meteorologist. 

Our friend’s first public education came from the district 
school, which his father had taken great pains to have above the 
usual standard. In addition, there were the “spelling classes” 
and ‘friendly associations,” and a small circulating library— 
agencies which he acknowledged to have been helps to him in his 
aspirations for knowledge, as they have been to many others. Of 
the effects of the “spdllling class” exercises he says: “I am fool- 
ish enough to believe that those winter evening battles were more 
useful and creditable than some of the athletic contests which in 
these days are doing so much to brutalize young men, and which, 
by their attendant betting leading to the worst results of gam- 
bling, are tending to make old and thoughtful men raise the 
question whether colleges are not becoming institutions to be 
avoided.” 

Of books at this time there were but few, but all he could get 
he read with avidity. Like every one else, he was fascinated with 
Bunyan’s Pilgrim’s Progress, both as to the story and the quaint 
old prints. He writes: “That of Apollyon’s battle with Christian 
so excited my imagination that when, being a little older, I was 
sent to the wood pile in the fast darkening twilight of a winter after- 
noon to bring in the evening supply of wood, I never felt alto- 
gether secure from that dreadful demon until the last armful was 


164 


fairly in;’ which leads one to wish that these Apollyons would 
always scare the bad boys and never the good ones. 

One other book, forgotten now perhaps, gave him the bent 
towards botany, which afterwards so much occupied him. This 
was Thornton’s Grammar of Botany. 

About this time steam navigation was occupying the father’s 
mind and after some efforts in that way on the Connecticut his 
attention was turned to the Hudson. He was thus frequently in 
New York for long periods while the son’s education was con- 
tinued for a year and a half at Stamford. Finally, in 1824, the 
family was removed to New York. The boy was now sent 
to the High School where, under the influence and _ instruc- 
tion of one of the teachers, a Mr. Barnes, he was instructed 
in mineralogy and had many a pleasant ramble in the 
country in his company. His school education was _ finally 
completed by a short course which he provided for himself 
at a private school, but between these two periods he at- 
tended the chemical lecture course of Dr. Torrey, an associa- 
tion which must have had great influence in his-pursuits in 
after life. His first business occupation was in a dry goods store, 
where he continued long enough to acquire a thorough detesta- 
tion of it. He then assisted his father in his steam transportation 
ventures and this occupied his business hours for many years. It 
is of more interest to us to know that his love of science continued 
and was intensified when, in 1836, he became a member of the New 
York Lyceum of Natural History, of which Dr. Asa Gray was 
then the Librarian and Superintendent. Here was commenced 
that friendship which was destined to be close and lasting. It was 
at this time that he acquired a taste for conchology, in which he 
made much progress and which resulted in a number of papers on 
this subject published in the Annals of the Lyceum. He thor- 
oughly explored the country in the vicinity, over land much of 
which is now probably closely built upon, and in every way which 
the time at his command and his means permitted strove to 
advance the scientific interests of himself and his associates. As 
early as 1846 he became a member of the Academy of Natural 
Sciences, of Philadelphia. In 1843 he made a very happy mar- 
riage and this, perhaps, was the eventual cause of his removal to 


165 


Philadelphia in 1861, where he long held a prominent position in 
the extensive and well-known car wheel works of A. Whitney & 
Sons, with the members of which his marriage connected him. 
His allegiance was necessarily transferred from the Lyceum to the 
Academy, of which he soon became a life member, and was gradu- 
ally advanced to many of its most important and laborious offices. 
Thus, in 1870, he became a member of its Council and was also 
made Conservator of its Botanical Section, the latter a most im- 
portant office as it placed the various and very important herbaria 
in his charge. He was Corresponding Secretary of the Concho- 
logical Section in 1879, and after having been long a member of 
its Publication Committee was made its Chairman in 1891. It 
will thus be seen how important his services were to this institu- 
tion and how great the esteem in which his good sense and active 
exertions as well as his wise and thoughtful counsel were held by 
his associates. But beyond all this, and especially after his retire- 
ment from business cares in 1885, he accomplished a great work 
which no one else connected with the Academy had time to do and 
for which, indeed, no one was better fitted than he. When he 
took charge he found four distinct herbaria as follows: that of Dr. 
C. W. Short; that of Schweinitz, composed principally of Fungi, 
very many of them types; the General Herbarium, and the North 
American Herbarium, the latter of which is of the utmost value, 
not only because of its size and completeness, but also because it 
contains a large number of the type specimens of Nuttall, Pursh 
and others of early botanists of this country. The specimens in 
these were loose in sheets of paper, very often those of more than 
one collector huddled in together, with the labels but loosely 
attached to the specimens. With great care and good judgment, 
and an indefatigable energy, he brought order out of this con- 
fusion, so that at last he had got the greater and more-valuable 
parts of the herbaria arranged and mounted and properly cata- 
logued. Nor did his benefations end with this, for he purchased 
all valuable sets of plants and bestowed them upon the Academy. 
The tender and appreciative minute adopted by it and hereafter 
appended is but a fitting testimony to his usefulness and unselfish 


devotion. 
Mr. Redfield lived for many years in one of the pleasantest 


166 


parts of Philadelphia and quite close to the Academy. He made 
occasional botanical excursions, of which notable ones were to the 
mountains of North Carolina in company with Dr. Gray and other 
botanists. There could not have been a more delightfully cheer- 
ful and obliging travelling companion. The writer well remem- 
bers that on one occasion when at Linville Falls, in what was 
then the wildest and least frequented part of the country, Mr. Red- 
field “ turned up missing,” to the serious concern of the rest.of the 
party. After considerable search he was found sitting on a mossy 
bank, writing up his diary with the utmost serenity, cheerfully an- 
swering anxious inquiries by saying, “Oh, I knew you would 
come for me.” In later years his summers were spent on Mt. 
Desert Island. The excellent catalogue of its flora lately pub- 
lished by Mr. Rand and himself attest his industry while there. 

It is impossible to speak too highly of Mr. Redfield’s personal 
character. Honorable, sincere, courteous, cheerful, always ready 
to do a kind act or to say a gracious word, he displayed that true 
nobility of character which comes of right principle faithfully ad- 
hered to, yet without a trace of aceticismn or austerity. 

Mr. Kand writes: « He was always being good and doing good. 
I have letters lamenting his death from young botanists, whose 
names even he may not have remembered or known, all telling the 
same story,—‘ he was so good to us, so kind in his interest and 
help, so courteous to us in ourignorance.’”” The Rev. Dr. Dickey 
said of him: “I have touched many good lives and found pleasure 
and example in close intercourse with many, * * * * but I have 
never touched a smoother life than this. * * * It was not the 
quietness of silence—it was like the soothing murmur of a mount- 
ain brook ; there was a beauty and fragrance like the beauty and 
fragrance of wild flowers, in this simple yet vigorous life.” 

And ‘so one cannot wonder that he won sincere and lasting 
affection and left a bright example of a right-living, true-hearted 
and attractive gentleman. Once, indeed, the serenity and happi- 
ness of his old age was broken by the stroke of a severe bereave- 
ment; but it only the better showed the strength of his character. 


_ “And thé more 
Fate tried his bastions, she but forced a door 
Leading to sweeter manhood and more sound.” 


167 


After some weeks of failing health he died on the twenty- 
seventh of Febrauary last, in the eightieth year of his age. 

A beautiful western grass, the sigs fees flexuosa commemo- 
rates his name and services. 

Appended are testimonials of learned societies and a bibli- 
ography. 

FROM THE ACADEMY OF NATURAL SCIENCES. 

The Academy of Natural Sciences of Philadelphia has heard 
with deep sorrow the announcement of the death of John H. Red- 
field, who, in his unselfish devotion to its interests, has long been 
one of its most active benefactors. 

Always an earnest student of nature his last years of deserved 
freedom from business engagements were devoted to his favorite 
studies in connection with the Academy, and to the arrangement 
and care of the Herbarium. 

The steady growth and admirable condition of the botanical 
collection constituted an enduring memorial of his industry and 
zeal. | : 
As Chairman of the Publication Committee and Member of 
the Council the same fidelity and discretion characterized the dis- 
charge of his duties. 

He was a man of strong but tender character; firm in his sup- 
port of the right, but tolerant of all honest difference of opinion ; 
cheerful, gentle, modest and cultured. Time to him was one of 
his most precious possessions, yet he was ever gladly at the service 
of those requiring advice or assistance. 

He was an earnest student, a wise counsellor and a steadfast 
friend. His encouragement and loving sympathy endeared him 
to his associates, who felt for him a’ personal affection which 
enables them to appreciate the irreparable loss sustained by his 
family, to whom they would offer their heartfelt sympathy. 


FROM THE NEW YORK ACADEMY OF SCIENCES. 
The Academy has learned with sorrow of the death of Mr. 
John H. Redfield, at his home, in cinta sae on February lav 


1895. 
Mr. Redfield was one of the earliest members of the Lyceum 


of Natural History, having been elected in 1836. During his 


168 


years of residence in New York he was most active in furthering 
the work of the Lyceum, a frequent contributor to its proceedings 
and the author of several conchological papers which were printed 
in its Annals. In connexion with his father, Mr. W. C. Redfield, 
he published, in Vol. IV. of the Annals, the first description of 
fossil fishes from the Mesozoic rocks of America, proposing the 
name of the genus Catopfterus and its type species C. gracilis, be- 
sides some others, for specimens from the Triassic beds at Dur- 
ham, Conn. He was thus the pioneer in this important branch 
in American palaeontology. He held the office of Recording 
Secretary of the Lyceum in the years 1887-8, and of Correspond- 
ing Secretary for the entire period from 1839 to 1860. After 
his removal to Philadelphia he did not lose his interest in the Ly- 
ceum, but continued his relations with it as a Corresponding Mem- 
ber, not only through the whole period of its existence under the 
old name, but also when the organization was changed and en- 
larged into the Academy and down to the time of his death. 
When the memorial volume was published, in 1887, Mr. Redfield 
furnished a large amount of most valuable data and reminiscences, 
which are embodied and acknowledged at many points in the book. 

Although personally known to but few of our present mem- 
bers, many have known of his great work in connexion with the 
Academy of Natural Sciences in Philadelphia, and by reason of 
this, and his early prominence in our Society, he has had our pro- 
found respect and grateful esteem. It is, therefore, 


Resolved, That it is the sense of the Academy that in the death 
of Mr. John H. Redfield, American science has lost a critical and 
enthusiastic student, a liberal patron and a devoted friend; and the 
Academy a co-laborer who greatly aided in its early period of 
organization, as an officer and a scientific investigator, and who 
was almost the last to connect’ its present membership with the 
generation of its founders and pioneers. 


FROM THE TORREY BOTANICAL CLUB. 


Mr. John H. Redfield, a highly esteemed active member of the 
Club since the time of its organization, the last but one of its 
original incorporators, a frequent contributor to our publications, 
the Conservator of the Botanical Section of the Philadelphia 
Academy of Natural Sciences, and well-known to the botanical 


169 


world as an author and editor, died at his home in Philadelphia, 
February 27th, 1895. Therefore it is 
Resolved, That the following record be made in our minutes: 

As a scientific co-laborer, we found in Mr. Redfield an enthusiastic 
lover of nature and of knowledge for its own sake, an energetic 
and persistent worker in the field and in the closet to the very end 
of a long and memorable life, and a gentleman of ripe culture and 
pleasant manners, always generous and helpful to others, and, 
though firm in his convictions as to what was right, ever modest 
and courteous in the expression of them. 

Resolved, That in his death we mourn the loss of a valued as- 
sociate and an ardent and faithful friend. 

Resolved, That this action be printed in our proceedings and a 
copy thereof transmitted to the family of the deceased. 


List of Scientific Papers and Notices by John H. Redfield. 


1. Fossil Fishes of Connecticut and Massachusetts, with a Notice of an unde- 
scribed Genus. Ann, N. Y. Lyc. Nat. Hist. 4: 35. pZ 2. 1837. 

2. Descriptions of some new Species of Shells. Ann, N. Y. Lyc. Nat. Hist. 4: 
163. p22. 1846. 

3. On the distinctive Characters of Cyfraea reticulata of Martyn and Cyfrae 
histrio of Menschen. Ann. N. Y. Lyc. Nat. Hist. 4: 417. 1. 7. 1847. 

4. Descriptions of new Species of ABullia and Marginella, with Notes upon 
S. B. Sowerby, Jr.’s Monograph of the latter Genus. Ann, N. Y. Lyc. Nat. Hist. 4: 
491. pir. 1848. e 

5. Description of new Species of Margined/7, with Notes upon sundry Species of 
Marginella and Cypraea. Ann. N.Y, Lyc. Nat. Hist. 5: 224. 1852. 

6. Descriptions of new Species of Helidicae. Ann. N. Y. Lyc. Nat. Hist. 6: 
14. M. 1853. 

7. Descriptions of new Species of Shells. Ann. N. Y. Lyc. Nat{Hist. 6: 
130. 7 Al. Ap. 1854. 

8. Descriptions of two new Species of North American Helicidae. Ann. N.Y, 
Nat. Hist. 6: 170. D. 1850. 

9. Description of a new Species of Mfarginella. Proc. Acad. Nat. Sci. Phila. 
1860: 174. My. 1860. 

Io. Letter to His Excell. Rawson H. Rawson, Governor of the Bahama Islands, 
with a Chart of the Bahama Hurricane of Oct., 1866. In Gov. Rawson’s Report in 
Blue Book for 1866, 2,3. Mr. 1853. 

11. Note on the first Discovery of Schisaea pusilla. Proc. Acad. Nat. Sci. 
Phila. 1869: 13. Ap. 1869. 

12. Search for Corema Conradii in Monmouth County, N. J. Proc. Acad. Acad. 
Nat. Sci. Phila. 1869: gt. My. 1869; Amer, Nat. 3: 327, Au. 1869. 

13. Notes upon the Monograph of the genus Marginella in Reeve’s Concholo- 
-gia Iconica. Tryon’s Amer, Journ. Conch. 5: $8 7 pZ QO. 1869. 

14. New Locality of Aspidium aculeatum in Stony Clove, Catskill Mountains. 


Amer. Nat. 3: 495. N. 1867. | 


170 


15. Observations on Marginellidae, introductory to a catalogue of the known re- 
cent and fossil species, Tryon’s Amer. Journ. Conch. 6: 2. Jl. 1870. 

16. Are certain Species of Botrychium epiphytic? Proc. Acad. Nat. Sci. Phila. 
1870: 91. Au. 1870. 

17. Rectification of the Synonomy of certain Species of A/arginelia, Tryon’s 
Amer. Journ. Conch. 6: 172. 0. 1870, 

18. Catalogue of the known Species, recent or fossil, of the Family Marginelli- 
dae. Tryon’s Amer. Journ. Conch. 6: App, 215. Q. 1870. 

19. Tetramerism in Lilium auratum Lindl, Bull. Torr. Bot. Club, 2: 32. 
Au, 1871. 

20. Oaks and Mistletoe. Bull. Torr. Bot. Club, 4:13. Ap. 1873. 

21. Fertilization of Asarum Canadense, Bull. Torr. Bot. Club, 4: 21. Je. 
1873. 

22. Dr. Torrey and Torrey’s Peak. Bull. Torr. Bot. Club, 5: 18. Ap. 1874. 

24. On Rd rcimiane ebenoides. Proc, Acad, Nat. Sci. Phila. 1874: 154. D. 
1874. 

25. Geographical Distribution of Ferns of North America, Bull. Torr. Bot. 
Club,6: 1. Ja. 1875. 

26. Notes upon Anychia dichotoma. Bull. Torr. Bot. Club, 6: 61. N. 1875. 

27. Note upon Dr, Torrey’s first Trip to the New Jersey Pines, prefixed to a let- 
ter of his dated July 9, 1818. Bull. Torr. Bot. Club, 6: 82. Mr. 1876. 

28. Notice of the Botanical Correspondence of Zacharias Collins, in Possession 
of the Academy of Natural Sciences of Philadelphia. Proc. Acad. Nat. Sci. Phila. 
1876: 81. Jl. 1876. : 

29. Southern Localities of Lygodium palmatum. Bull. Torr. Bot. Club, 6: 232. 
My. 1878. . 
30. Obituary notice of Robert H. Brownne. Bull. Torr. Bot. Club,6: 291. F. 
1879. : 

31. Aspidium aculeatum in Pennsylvania. Bull. Torr. Bot. Club, 6: 291. 
F. 1879. _ 

32. Aspidium aculeatum at Bushnellsyille Clove in Catskill Mountains. Bull. 
Torr. Bot. Club,6: 331. Au. 1879. 

_» 33. Notes of a Botanical Excursion into North Carolina. Bull. Torr. Bot, Club, 
6: 331. Au. 1879. 

34. Dissent from Mr. Meehan’s Views upon the Timber-line of ne Mountains. 
Proc. Acad. Nat, Sci. Phila. 1880: 345. N. 1880. 

35. Herbarium of the Academy of Natural Sciences of Philadelphia, Bull. Torr. 
Bot. Club, 8: 42, Ap. 1881. 

36. The Muhlenberg Herbarium, ° Bull. Torr, Bot, Club, 8: 80. Jl. 1881. 

37. Aspidium Lonchitis Swz. in Colorado. Bull, Torr. Bot. Club, 8; 105. S. 
1881, ; 

38. Occurrence of Hieracium aurantiacum in the Catskill Mountains. | Bull. 
Torr. Bot. Club, 8: 112, O. 1881; Proc. Phila. Acad. Nat. Sci., 1881: 429.. D. 
1881. e 

39. Biographical Sketch of Dr. William Baldwin. Bot, Gaz. 8: 233. Je. 1883, 

40. Note upon the Date of a Letter from Dr. Torrey. to.Amos Eaton. Bot. Gaz 
8: 317. O. 1883. i 


171 


41. Corema Ccnradii and its Localities. Bull. Torr. Bot. Club, 11: 97. S. 1884. 

42. Obituary Notice of John Williamson. Bull. Torr, Bot. Club, 11: 104. 
S. 1884. 

43. Further Notes upon Corema Conradti. Bull. Torr. Bot. Club, 12: 93. 
S. 1885. 

44. Insular Vegetation; Flora of Great Duck Island, Maine. Bull. Torr. Bot. 
Club, 12: 103. O. 1885. 

45. On the Flora of Martha’s Vineyard and Nantucket. Proc, Acad. Nat. Sci. 
Phila. 1885: 378. D. 1885. 

46. Still further Notes upon Corema Conradii. Bull. Torr. Bot. Club, 13: 220, 

N. 1886. 

47. Euphrasia officinalis on the Coast of Maine. Bull. Torr. Bot. Club, 232. 
D. 1886, 

48. On Insular Floras.' Bull. Torr. Bot. Club,.13: 245. D. 1886. 

49. Rediscovery of Corema Conradii in Monmouth County, N. J. Bull. Torr. 
Bot. Club, 16: 192. Jl. 1889; Proc. Acad. Nat. Sci. Phila. 1889: 135. Jl. 1889. 

50. Pinus Banksiana with Corema Conradii on Schoodic Peninsula, Coast of 
Maine. Bull. Torr. Bot. Club, 16: 295. N. 1889: Proc. Acad. Nat. Sci. Phila, 
1889: 344. N. 1880. 

51. Stellaria hemifusa on the Coastof Maine. Bull. Torr. Bot. Club, 17: 38. 
F. 1890. 

52. Notice of the Occurrence of Scadiosa australis near Pittsfield. Mass, Bull. 
Torr. Bot. Club, 19: 341. N. 1892. : 

53. Obituary Notice of Isaac C. Martindale. Bull. Torr. Bot. Club, 20: 98. 
1893. 
54. Preliminary Catalogue of the Plants growing on Mt. Desert and adjacent 
Islands. By Edward L. Rand and John H. Redfield. Cambridge. 1894. 


A rosaity marine Diatomaceous Deposit at St. Augustine, Florida. 


By CHARLES S, Boyer. 


In-1886 an artesian well was sunk at the Ponce de Leon Hotel, 
at St. Augustine, Florida. Samples of earth from different depths 
were sent to Mr. Lewis Woolman, who proposes to publish the 
results of his investigations into the geological character of the 
different strata. A layer of bluish clay at a depth of between 85 
and 135 feet was found to contain diatoms, spicules, foraminifera 
and a few polycistinae. Unfortunately, the material was very 
small in amount, and the diatoms occurred in but two layers at 
the depths of 90 and 120 feet, so that the list furnished below, 
although exhaustive of the material obtained, appears to but indi- 
cate the existence of a richer bed which, it is hoped, may be 


172 


brought to light by another well-boring or the discovery of some 
outcrop. 

It may be briefly stated, according to Mr. Woolman’s conclu- 
sions, that the diatomaceous clay bed immediately overlies an 
Eocene deposit and is beneath a Pleistocene. As will be noticed, 
the diatoms correspond, to a great extent, to those of the great 
Miocene beds of the Atlantic coast, such as the Richmond, Not- 
tingham and Atlantic City deposits; the presence of one form, at 
least, if not two, appears to indicate a correspondence with the 
Barbadoes deposit, which is now claimed to be Pliocene, while 
other forms are still extant. It is, therefore, difficult to determine 
to what period the Florida deposit belongs. 

The following are the forms :— 

Actinocyclus Ehrenbergtt Ralfs, several varities. 

A, ellipticus Grun., rare. 

A. interpunctatus Ralfs. 

A. Ralfsu (Wm. 5m.) Ralfs. 

A, subtilis (Greg.) Ralfs, var. 

Actinoptychus Grundleri A. S. 

A, undulatus Ralfs, several vars. 

_ A. vulgaris Schum., several vars. 

Aulacodiscus Crux Ehr., rare. : 

A. mammosus (var. extans?) Grev., very rare. Diam. 0. 152 mm. 
Surface, central portion, flat to about one-half of the radius, outer 
edge distinct but somwhat turned inward at the inflations, giving 
a slightly quadrangular outline. Central space in one specimen 
round, in the other irregular. Width of bullae adout one-fourth the 
radius. In Greville’s figure the width appears to be about one- 
half. The processes are also smaller in the Florida form. Not 
being able to secure a specimen of Greville’s species for compari- 
son, I am unable to determine the differences further. Mr. Adre. 
Le Tourneur, of Paris, from an examination of a photopraph sent 
him, thinks it may be the variety extans (Grev.) Rattr. This 
beautiful form is quite rare, only two nearly perfect valves and 
three fragments having been noticed. It has not been found, here- 
tofore, we believe, in any other of the continental deposits, and its 


presence appears to link the Florida diatoms with those of Barba- 
does. 


“ans 


A. Mollert Grun. 

A. Rogersit (Bail.) A. S., common at 90 ft. 

The deposit presents all the forms figured by Schmidt, with a 
number of intermediate variations. 

Biddulphia Tuomeyi (Bail.) Roper. Several variations of this 
form approach 2. e/egantula Grev., and others are much smaller 
than the type. 

Coscinodiscus apiculatus Ehr. 

C. marginatus Ehr. 

C. emphalanthus Ehr. 

C. perforatus Ehr. 

C. radiatus Ehr. 

C. vobustus Grev. 

The forms of Coscinodisct noticed furnish an interesting study 
of variations, especially in those which approach apzculatus and 
marginatus. 

Craspedodiscus coscinodiscus Ehr., 120 ft. 

Lupodiscus radiatus Bail. A form with five ocelli occurs rarely 
at the depth of go ft. 

Goniothecum odontella Ehr., 90 ft. 

Grammatophora maxima Grun. var.? rare. 

Fyalodiscus laevis Ehr. 

FT. subtéis Bail. The markings in the greater number of 
specimens examined are much coarser than in the type form. In 
some valves which present a greenish color when dry, numerous 
spines are seen at regular intervals, and the zone between umbil- 
icus and margin resembles that of 7. maximus, P. Petit (Diato- 
mee’s de l’le Campbell, f/. z. fig. 7). Other forms without spines 
correspond closely to the var. Japonica of Castracane (Rep. on 
Diat. coll. by H. M. S. Challenger, p/. 78. fig. 4). 

Melosira sulcata (Ehr.) K., abundant. 

Navicula Lyra Ehr., very rare, 90 ft. 

NV. praetexta Ehr., very rare, 90 ft. 

Pleurosigma affine, var. fossilis Grun. 

Podoswa Montagnei Kiitz.? , very rare, 90 ft. This form is 
doubtful as the rim is wider and the general appearance more. 
robust than in any. of the figures published. 

Khaphoneis gemmifera Ehr. 


174 


Rhizosolenia sp. ? 

Stephanogonia (Mastogonia) actinoptychus Ehr., rare. 

Stephanopyxis appendiculata Ehr. 

S. corona (Ehr.) Grun. 

S. turris (Grev.) Ralfs, rare. 

Stictodiscus Trunanii Witt. ?, very rare, 90 ft. This form differs 
from that figured by Truan and Witt. (Die Diat. der Polycyst. von 
Jeremie in Hayti, p/. 4. figs. 23 & 24), inasmuch as the rim is less 
definite and the outline slightly irregular. Only one imperfect 
specimen was found. It appears to form another link between 
the continental and insular deposits. 

Triceratium condecorum Ehr., rare, 90 ft. 

T. Kainit E. A. Schultze, rare,-120 ft. 

T. semicirculare Brightw.— Euoata Brightwellii Ralfs. 

T. spinosum (Ehr.) Bail. 


New Species of Parasitic Fungi. 
By S. M. Tracy anp F; S. Earte. 


[Type specimens in the herbaria of the authors, of the U. S. Department of Agri- 
culture, of Rutgers, Harvard and Columbia Colleges. ] 


PUCCINIA NOTABILIS n. sp. III. Amphigenous; sori black, con- 
fluent, forming small hemispherical or irregular masses on the 
bracts and petioles or involving the larger stems, forming fusiform 
black gall-like swellings two or three times their diameter and 
3-4 cm. in length; teleutospores uniformly oval and obtusely 
rounded, slightly constricted, epispore smooth, thickened at the 
apex, 55-60 by 30-33 ; pedicel hyaline or slightly tinted, very 
long and flexuous, 225-275 ps. 

On Arrow-wood (Pluchea borealis ?) Rio Penasco, New Mexico, 
January, 1895. 

Puccinia Paspati n. sp. II. Usually hypophyllous, sometimes 
amphigenous; sori linear, sometimes confluent, dark brown; 
uredospores globose or obovate, very abundantly and sharply 
echinulate, brown, 24 by 25-30 w. III. Sori linear, darker than 

te uredo sori, usually on the leaf sheaths; teleutospores irregular, 
35 by 27 to 30 by 35 yp, usually oval, much constricted, with the 
slender nearly hyaline pedicel attached obliquely to one side of 
the lower end, often orbicular with the septum vertical and the 


175 


pedicel attached either to the septum or near it, and the epispore 
of both cells distinctly thickened at the apex. 

On Paspalum virgatum, New Orleans,.La., November, 1894. 

UstiLaco Crus-Gatii n. sp. Involving the panicles and upper 
nodes, making the panicles abortive and forming pustules some- 
times I cm. in diameter at the nodes; spore masses powdery, 
black, at first covered by a gray membrane; spores oval or sub- 
globose, fuscous, sharply echinulate, 9-10 by 11-13 p. 

On Panicum Crus-galli, Salt Lake City, Utah, August, 1887. 

This seems to approach U. AMaydes, but the spores, though 
equally echinulate, are uniformly larger and more elongated. 

UstILaGo. TONGLINENSIS n. sp. Spore masses infesting the 
Ovaries and causing the glumes to open widely at maturity, black ; 
spores globose, dark brown, sharply and thickly echinulate, 9-11 y. 

On Jschaemum ciliare, Tonglin, Singapore. H. N. Ridley. 

UsTILAGO ORNATA n. sp. Infesting the ovaries. Spore masses 
black, pulverulent, 20-30 mm. in diameter; spores subglobose, 
very thickly beset with strong and prominent echinulations which 
give the spore a burr-like appearance, 12-12 by 15 yp. Panicles 
bearing affected ovaries always remain partially enclosed within 
the sheath of the upper leaf. 

On Leptochloa mucronata, Starkville, Miss., November, 1894. 


UstTILaGo PERTUSA n. sp. Infesting the ovaries ; spore masses 
hard and compact, black, finally pulvinate; spores small, globose, | 
€pispore covered with prominent irregular verrucose projections, 
5-7 fe 

On Setaria macrochaeta, Queensland. F. M. Bailey, 1890. 

Ustiraco pustutata n. sp. Infesting the ovaries, or forming 
rounded bullate swellings which often surround the entire stem 
and branches of the panicle, or form irregular distortions on the 
leaves and sheaths; spore masses dark brown, long covered by a 
stramineous membrane which is a distortion of the tissues of the 
host, 25—75 mm. in diameter; spores dark brown or fuscous, sub- 
globose or oval, cell-wall very thin, slightly echinulate, 7.5 to 9 y., 
or 7 by 9 ps. 2 

On Panicum proliferum, Starkville, Miss., October, 1894. 

Dimerosporium MacnoriaE n. sp. Epiphyllous, on small in- 
_ definite areas ; mycelium of dark brown irregular branching and 
Septate hyphae; conidia clavate, dark colored, 4-5-septate, 7-8 
by 45-55 #; perithecia depressed-hemispherical, black, usually in 
clusters of 5 to 10, astomous, 50-100; asci numerous, broadly 
- oval or obovate, 8:spored, 35-40 by 45-50, paraphysate; para- 
physes thread-like, colored towards the tips; sporidia biseriate, at 


176 


first hyaline, dark fuliginous when mature, obovate, I-septate, 
constricted, 9-10 by 20-23 p. 

On living leaves of Magnolia Virginiana, Ocean Springs, Miss., 
May, 1894. 

AstTeripiIuM ILricit n. sp. Hypophyllous; mycelium none; 
perithecia black, scattered, orbicular, aplanate, membranaceous, 
cellular, not radiant, fragile, wrinkled, 200-400 #4; asci numerous, 
suborbicular, 8-spored, 30~40 ; sporidia subpyriform, often curved, 
granular, colorless, at first uniseptate, becoming 3-septate at ma- 


so. turnty. 


On living leaves of [/icium Floridanum, Ocean Springs and 
Biloxi, Miss., March, 1889. 


LagsrapiA ILiiciicoLa n. sp. Amphigenous, occupying large 
irregular brownish subarid definitely limited areas, usuaily in- 
volving the apical half of the leaf; perithecia very numerous, scat- 
tered, erumpent, more abundant on the upper surface, black, 
membranaceous, suborbicular or lenticular, obscurely ostiolate, 
finally collapsing; asci without paraphyses, clavate, stipitate, 
thickened at the apex, 40-50 by 10-12 »; sporidia ovoid or fusi- 
form, continuous, hyaline, granular, about 15 by 5 yp. 

On living leaves of ///icitum Floridanum, Ocean Springs, Miss., 
March, 1892. 

_ SPHAERELLA ANDROMEDAE n. sp. Hypophyllous; spots none ; 
perithecia abundant, scattered, often covering the entire lower 
surface of the leaf, black, erumpent, ostiolate, at length collapsing ; 
asci obovate, somewhat thickened at the apex; sporidia oval, 
hyaline, uniseptate, cells about equal; 7-8 by 2.5-3 4. 

On living leaves of Pieris nitida, Ocean Springs, Miss., March, 
1888. 


LEMBOSIA ANGUSTIFORMIS n. sp. Epiphyllous, on raised brown 
irregularly stellate blisters; mycelium scant; perithecia black, long 
and narrow, often flexuous, seldom branched, 60-80 by 175-300 /4; 
subiculum reduced to a few short flexuous slightly fuscous 
branching threads; asci broadly oval, about 15 by 18 4; sporidia 
obovate, somewhat unequally uniseptate, constricted, at first hya- 
line, becoming light fuliginous, 8-10 by 4-5 p. 
On flex coriacea, Ocean Springs, Miss., May, 1894; Biloxi, 
Miss., July, 1894. 


This differs widely from the following in gross appearance on 


the leaf, in the narrower perithecia, less conspicuous subiculum 
and smaller asci and sporidia. 


LEMBOSIA PRINOIDES n. sp. Epiphyllous, on orbicular pallid 


177 


spots; mycelium scant, brown; perithecia scattered, black, sub- 
carbonaceous, fimbriate-margined, elliptical, subacute, often fork- 
ing, 200-350 by 120-150; subiculum of dark brown irregular 
nodular usually continuous and branching brittle threads; asci 
ovate, 8-spored, 30-35 by 15-18; sporidia elliptical, unequally | 
uniseptate, somewhat constricted, subhyaline, becoming fuligi- 
nous, 10-15 by 4-5 p. 

On Mex coriacea, Biloxi, Miss., July, 1893. 

Lemposia Iniiciicora n. sp. Epiphyllous, on large light 
brown orbicular or irregular areas; perithecia numerous, super- 
ficial, carbonaceous, usually linear and strict, occasionally trian- 
gularly stellate, 100 by 300-400; subiculum of light brown 
flexuous transparent continuous variously branching and anas- 
tomosing threads; asci very numerous, oval or ovate, 8-spored, 
25-30 by 12y; sporidia oval, uniseptate, slightly constricted, 
ends obtusely rounded, hyaline, at length slightly colored, 8-10 by 
by 3-4 p. 

On LMicium Floridanum with Asteridium (llicit, Ocean Springs, 
Miss., March, 1889. 

VERMICULARIA STACHYDIS n. sp. Perithecia scattered, sub- 
superficial; setae somewhat floccose and nodular, septate, oliva- 
ceous, paler towards the tips, which are obtuse and slightly en- 
larged ; conidia falcate, attenuate at each end but without evident 
basidia, guttate, at length faintly 4-5-septate, 35-40 by 3-4 p. 

On dead stems of Stachys affinis, Starkville, Miss., October, 


1893. 
This differs from other described species in the larger, at length 
several-septate conidia, and the weak rather light colored setae 


with swollen tips. 

DIPLODIA MINUTA nN. sp. Scattered over indeterminate whitened 
areas; perithecia erumpent, small, 90-120 yp, membranaceous, 
smooth, ostiolate; sporules minute, oval, uniseptate, not constricted, 
light yellow, 6-8 by 3-4 p. oe 

On living stems of Zecoma radicans, with Pestaloszia breviaris- 
fata, Starkville, Miss., March, 1895. : 

DieLtopia SaAssAFRAS n. sp. Perithecia very numerous over 
large areas, black, hemispherical, erumpent, finally opening by an 
irregular fissure; sporules numerous, ovat=, fuscous or nearly 
black, with a very distinct hyaline septum near the smaller end, 
13-14 by 5-6 p. : 

On living twigs and branches of Sassafras, Starkville, Miss., 


April, 1894. 


178 


HENDERSONIA TAPHRINICOLA n. sp. Epiphyllous, on white 
bordered spots. Perithecia scattered, black, erumpent, at length 
collapsing; conidia truncate, cylindrical, fuliginous, 2-septate, 
constricted at the septa, 12-14 by 4-5 p. 

On old whitened blisters of Zaphrina on Quercus Virginiana, 
Ocean Springs, Miss., February, 1887. 


PesraLozziaA CLIFTONIAE n. sp. Epiphyllous, on orbicular 
arid brown-bordered spots. Acervuli scattered, bursting through 
the dried epidermis; conidia obovate, sometimes curved, 4-septate, 
septa often oblique, three medial cells fuliginous, the upper two 
dark and opaque, the lower one paler, basal cell colorless, small, 
short, acute, abruptly contracted to the short stipe; apical cell 
reduced to a short colorless apiculus bearing the three widely 
divirgent setae, 16-18 by 8 yw; setae 12-14 p. 

On living leaves of Cliftonia ligustrina, Ocean Springs, Miss., 
November, 1893. 


PESTALOZZIA BREVIARISTATA n. sp. Acervuli scattered over in- 
definite whitened areas, black, at length collapsing ; conidia curved, 
elliptical, 5—septate, apical and basal cells colorless, medial cells 
fuscous, septa often diagonal, 25-27 by 7-8 »; stipe hyaline, half 
the length of the conidium, somewhat swollen at the base, often 
deciduous; arista single, strongly oblique, thickened, scarcely 
one-fourth the length of the conidium. 

On living stems of Tecoma radicans with Diplodia munuta, 
Starkville, Miss., March, 1895. 


SCOLECOTRICHUM PUNCTULATUM n. sp. Amphigenous; spots 
indefinite ; hyphae in small caespitose clusters, irregularly flexu- 
ous and nodular, olivaceous, 2~3-septate, 60-70 by 5-6 p; conidia 
oval or oblong, ends obtusely rounded, fuscous, epispore distinctl 
granulose or punctulate, typically uniseptate but often continuous, 
and occasionally 3-septate, 15-20 by 6-8 p. 

On /ris pabularia, Starkville, Miss., January, 1894. 

CERCOSPORA FLEXUOSA n. sp.. Forming large indefinite effused 
patches on the under side of the leaf. Hyphae ferruginous, ir- 
regularly flexuous throughout, sometimes branched, many-septate, 
denticulate, 75-150 by 4-5 »; conidia obclavate, fuscous, 2-6, 
3-4-septate, not constricted, 20-30 by 4-5 p. 

On leaves of Diospyros Virginiana, Biloxi, Miss., July, 1892, and 
Ocean Springs, Miss., October, 1880. 


This differs from C. Dzospyri(Thum.) Cke. in its septate hyphae 


and much longer spores, and from other Cercosporas reported on 
the same host in the absence of definite spots. 


179 


CERCOSPORA GRAMINICOLA n. sp. Spots none; amphigenous 
and most abundant on the midvein; hyphae caespitose, straight 
or somewhat flexuous, fuscous, 2—3-septate, 75-100 by 4-5 ; 
conidia clavate, continuous when young but finally becoming 1—3- 
septate, fuscous, 35-40 by 7.5-10 yp... 

On languishing leaves of Phleum pratense, Starkville, Miss., 
November, 1894. 

Crercospora Hiniscr n. sp. Hypophyllous, on large indeter- 
minate areas; hyphae fuliginous, clustered, somewhat itregular, 
once or twice septate near the base, 25-40 4; conidia obclavate, 
somewhat curved, hyaline, guttulate, at length faintly 3~-5-septate, 
40-60 by 3-4 pu. 

On living leaves of Azbiscus esculentus, New Orleans, La., No- 


vember, 1894. 

CERCOSPORA MARITIMA n. sp. Amphigenous; not forming defi- 
nite spots, but densely effused over considerable dark-colored 
areas ; hyphae fasciculate, mostly straight, olive brown, 5~6—sep- 
tate, often swollen at the septa, 40-100 by 4-6 ys: conidia paler, 
elongated, straight or slightly curved, mostly uniseptate, the upper 
cell shorter and broader than the lower, 50-60 by 5-7 yz. 

On Croton maritimum, Horn Island, Miss., March, 1892. 

CrRcospora MISSISSIPPIENSIS n. sp. Amphigenous, but more 
abundant on the under side of the leaf. Spots small, brownish, 
irregular, usually bounded by the veins, surrounded by an irregu- 
lar raised and darker border having a pallid outer margin; hyphae 
divergent, irregularly floccose, nodular, usually uniseptate above 
the somewhat bulbous base, 60-70 by 5 4; conidia fuscous, nar- 
rowly clavate, tapering from the rather obtuse apex to the attenu- 
ated filiform nearly hyaline base, 4-8 or more septate, 75-150 by 
4-5 p 
On Smilax glauca, and S. rotundifolia, Starkville, Miss., No- 
vember, 1893. 

TETRAPLOA DIVERGENS n. sp. In black irregularly linear masses; 
conidia broadly oval, dark-fuliginous, 12-14 by 8-9, quarter- 
_ nately divided, each division usually 2-septate, and terminated by 
an obtuse semi-transparent continuous divergent horn-like pro- 
jection, 4-5 win length. 

On living or languishing leaves of Panicum agrostidiforme, 


Starkville, Miss., October, 1894. 


180 


The Systematic: Botany of North America. 


The Board of Editors of the “Systematic Botany of North 
America” announce the following arrangements for the mono- 
graphing of groups thus far decided on. Other assignments will 
be reported as made. It is requested that collectors communicate 
material for study to the monographers, even of the commoner 
and well-known species, so that the geographical distribution may 
be presented as accurately as possible. 


Myxomycetes: Mr. O, F. Cook, Huntington, N. Y, (at present abroad). 

Scuizomycetes: Prof. H. L. Russell, University of Wisconsin, Madison, Wis. 

CHLOROPHYCEAE: In charge of Prof. Geo. L. Atkinson, Cornell University. 

PHycoMyYCETES: Prof. Byron D. Halsted, Rutgers College, New Brunswick, N. J. 

SACCHAROMYCETES: Dr. J. Christian Bay, State Bcard of Health, Des Moines, Iowa. 

TAPHRINEAE: Prof. Atkinson. 

HELVELLINEAE: Prof. Lucien M. Underwood, Greencastle, Ind. 

PEZIZINEAE, PHACIDINEAE: Mr. Ellis J. Durand, Cornell University. 

FUNGI IMPERFECTI: Prof. Byron D. Halsted, Rutgers College, and Mr. J. B. Ellis, 
Newfield, N. J, ° 

USsTILAGINEAE: Prof, Halsted. 

UREDINEAE: Prof. J. C. Arthur, La Fayette, Ind. 

POLYPORINEAE: Prof. Underwood. 

BOLETINEAE, AGARICINEAE: Prof. Chas. H. Peck, State Museum, Albany, N. Y. 

GASTEROMYCETES: Mr. A. P. Morgan, Preston, Ohio. 

HEPATICAE: Prof, Underwood. 

Musci—Acrocarpi-: Mrs. N. L. Britton, Columbia College, New York City; //euro- 

_ carpi Prof. C. R. Barnes, University of Wisconsin; Sphagnaceae: Mr. John 

K. Small, Columbia College, New York City. 

PTERIDOPHYTA: Prof. Underwood. 

TYPHACEAE, SPARGANIACEAE, NAIDACEAE, JUNCAGINACEAE: Manuscript of the late 
Dr. Thos. Morong. 

ALISMACEAE: Mr, Jared G, Smith, U. S, Department of Agriculture. 

HyYDROCHARITACEAE: Manuscript of Dr. Morong, 

GRAMINEAE: In charge of Prof. F, Lamson-Scribner, U.S. Department of Agriculture. 


CYPERACEAE: Prof. L. H. Bailey, Cornell University, and Prof. N. L. Britton, Col- 
umbia College. 


ARACEAE: Manuscript of Dr, Morong. 

LEMNACEAE: Mr. E. P. Sheldon, University of Minnesota, Minneapolis, Minn. 
ERIOCAULACEAE: Manuscript of Dr. Morong, 

COMMELINACEAE: Mr. J. N. Rose, U.S, Department of Agriculture. 
JuNncAcEAE: Mr. F. V. Coville, U. S. Department of Agriculture. 

LILIACEAE: Mr J. N, Rose. 

SMILACEAE: Manuscript of Dr. Morong. 


181 


DioscorEACEAE: Prof. A. S, Hitchcock, Kansas Agricultural College, Man- 
hattan, Kansas, 

SAURURACEAE, PIPERACEAE, CASUARINACEAE: Mr. T. H. Kearney, Jr., U. S. De- 
partment of Agriculture. 

JUGLANDACEAE: Prof. N. L. Britton. 

MYRICACEAE: Prof. Britton. 

LEITNERIACEAE: Prof. Wm. Trelease, Missouri Botanical Garden. 

SALICACEAE: Mr. M. S. Bebb, Rockford, Ill. (Saéix). 

BETULACEAE, FAGACEAE, ULMACEAE, MorACEAE; Mr. Geo. B. Sudworth, U. S. 
Department of Agriculture. 

URTICACEAE, LORANTHACEAE, SANTALACEAE, OLACACEAE, ARISTOLOCHIACEAE: 
Mr, T. H. Kearney, Jr. 

POLYGONACEAE: Mr. John K. Small, Columbia College. 

CHENOPODIACEAE: Mr. Willis L. Jepson, University of California, Berkeley, Cal. 

AMARANTHACEAE: Messrs. E. B. Uline and W. L. Bray, Lake Forest University» 
Lake Forest, Ill. ‘ 

PHYTOLACCACEAE: Prof. A. S. Hitchcock, Kansas Agricultural College. 

PORTULACACEAE: Mr. Jepson. 

NyYMPHEACEAE: Prof. Chas. A. Davis, Alma College, Alma, Mich. 

RANUNCULACEAE: Prof. N. L. Britton. 

MENISPERMACEAE: Prof. A. S. Hitchcock. 

CALYCANTHACEAE: Mr, T, H, Kearney, Jr. 

GERANIACEAE, OXALIDACEAE, LINACEAE: Prof. Wm. Trelease. 

MALPIGHIACEAE, ZYGOPHYLLACEAE, RUTACEAE: Miss Anna M. Vail, Columbia 
College, New York City. 

SIMARUBACEAE, BURSERACEAE, MELIACEAE: Mr. Geo, B. Sudworth. 

POLYGALACEAE: Dr. Wm. E, Wheelock, Columbia College: 

CALLITRICHACEAE: Studies of Dr. Morong. 

MALVACEAE: Mr. J. Burtt Davy, University of California. 

GUTTIFERAE, HypERICACEAE: President John M. Coulter, Lake Forest University. 

CISTACEAE: Prof. Britton. 

EMPETRACEAE: Prof. Hitchcock. 

LIMNANTHACEAE, AQUIFOLIACEAE, CELASTRACEAE, HIPPOCRATACEAE, STAPHYLEA- 
CEAE, ACERACEAE, HIPPOCASTANACEAE, SAPINDACEAE, BALSAMINACEAE! 
Prof. Trelease. 

RHAMNACEAE: Mr. Willis L. Jepson. 

ViTaAcEAE: Prof. L. H. Bailey. 

LoasacreaE: Dr Wm. E. Wheelock. 

CACTACEAE: President Coulter. 

THYMELEACEAE, ELEAGNACEAE: Prof. Hitchcock, 

HALORACEAE: Studies of Dr. Morong. 

ARALIACEAE: Prof, Hitchcock. 

UMBELLIFERAE; Pres. Coulter and Mr. Kose. 

CORNACEAE: Pres. Coulter. 

CLETHRACEAE, PYROLACEAE, MONOTROPACEAE, LENNOACEAE, ERICACEAE, DIAPEN- 
SIACEAE: Mr. F. V. Coville, U. S. Department of Agriculture. 


182 


MYRSINACEAE, PRIMULACEAE, PLUMBAGINACEAE,. SAPOTACEAE, EBENACEAE, SYM- 
PLOCACEAE, STYRACACEAE, OLEACEAE, LOGANIACEAE: Mr. John K. Small, 
Columbia College. 

GENTIANACEAE: Mr. Chas, L. Pollard, U. S. Department of Agriculture. 

BORAGINACEAE: Prof, Edward L. Greene, University of California, 

VERBENACEAE: Prof, A. S. Hitchcock. . 

SOLANACEAE, SCROPULARIACEAE: President Coulter. 

CICHORIACEAE, COMPOSITAE: Prof. Greene (Aster and Solidago by Prof. Thos. C. 
Porter, Lafayette College, Easton, Penn.). 


Botanical Notes. 


A Redfield Memorial Herbarium Fund. It has been decided 
that no better monument to the memory of John H. Redfield 
could be erected than to arrange for completing and caring for the 
work he loved, and to which he gave freely so many years of his 
life—namely, the Herbarium of the Academy of Natural Sciences. 
- Mainly through his disinterested labors, it stands to-day scarcely 
second to any in the United States, containing, besides many un- 
named, over thirty-five thousand named species of flowering plants 
and ferns, the half of which have been verified and fastened down. 

No one can probably be found to give the years of time he so 
freely gave. In order to carry on the work, and add to the col- 
lection, as exploring expeditions afford the opportunity, it has 
been proposed to establish a fund for its maintainance. 

Mr. Redfield’s will provides that his herbarium, minerals, 
shells and scientific works shall be sold to help the Herbarium— 
thus furnishing a nucleus for the proposed fund. It is in mind to 
raise $20,000, but the interest of any sum that may be contributed 
can at once be made available. Statements will be furnished from 
time to time to contributors, keeping them informed of the pro- 
gress of the contributions. Checks may be made payable to the 
order of Thomas Meehan, Director, or Stewardson Brown, Treas- 
urer, and mailed to either at the Academy of Natural Sciences, 
Nineteenth and Race streets, Philadelphia. 

Announcement of the next Meeting of the American Microscopical 
Society. The next meeting of the American Microscopical Society 
will be held at Cornell University, in Ithaca, N. Y., August 21, 22 
and 23, 1895, that is the week previous to the meeting of the 


183 


American Association for the Advancement of Science, which is 
to be held in Springfield, Mass. 

The unsurpased beauty of the location of the University, and 
the richness of both its terrestrial and aquatic fauna and flora, 
make this an ideal place for holding the meeting. It is equally 
attractive to the student of natural history and to those who love 
beautiful scenery. 

The University buildings, which will be at the disposal of the 
' Society, are especially adapted for the formal presentation of 
papers, blackboard illustrations, hanging of diagrams, etc., as well 
as for any demonstration that authors may desire. to make. The 
armory is very conveniently located, both for the University and 
for the city, and a soiree there can hardly fail to be a great success. 

Besides the attraction of papers and demonstrations by mem- 
bers, nearly all the opticians have expressed not only a willingness 
but a desire to be present and make an exhibit of their micro- 
scopes and microscopical apparatus, thereby affording the mem- 
bers an opportunity to see all the new and standard apparatus. 

A special feature of the coming meeting will be the setting 
apart of one or more sessions for the reading of papers on methods 
and the demonstration of special or new methods. The chairman 
of the local committee, Professor W. W. Rowlee, or the President, 
will be glad to receive requests from those who desire to have 
some specially difficult method or structure elucidated, and an 
effort will be made to get some member particularly expert in such 
subject to demonstrate it before the Society. 


Proceedings of the Club. 


Turspay Eventnc, Marcu 12th, 1895. 
Vice-President Lighthipe in the Chair and 24 persons present. 
The following were elected active members: Mr. Claude Crit- 
tenden, Mrs. Wm. Starr Dana, Mr. Eugene H. Van Nest, Mr. 
James P. Gardner, Mr. S. Cook. 

The Committee appointed to draw up resolutions on the death 
of Dr. J. Bernard Brinton presented a report which was accepted, 
and a copy of it placed on file. 


184 


The Instruction Committee reported progress, and submitted 
the prospectus of the summer class for 1895. The report was 
accepted. 

The Chair appointed the following Field Committee for 1895: 
Chairman, Mr. Small; Mrs. Britton; Mr. Tyler; Mr. Van Sickle; 
Dr. Rusby. 

The announced papers of the evening were then presented 
as follows: 

George V. Nash, “Remarks on some new and little-known 
Plants of Central Florida.” Illustrated by specimens. (Published 
in this issue of the BULLETIN.) 

Elizabeth G, Britton, “Some Notes on the Genus J/nium.” 
(To be published in a subsequent issue of the BULLETIN.) 

A. Cogniaux, “ Descriptions of new Melastomaceae from Bo- 
livia,’ communicated by Dr. Rusby. (lo be published in a sub- 
sequent issue of the BULLETIN.) 


WEDNESDAY EVENING, MARCH 27TH, 1895. 


Dr. Britton in the Chair and 34 persons present. 

Mr. Eugene Smith, Miss Bliss, Miss Augusta Bliss and Miss 
Edith Parish were elected active members. 

Dr. Britton announced the death of Mr. John H. Redfield, 

and remarked upon his life and labors. The Secretary also spoke 

upon the subject. Mr. Small moved that the Chair appoint a 
committee of three to draft suitable resolutions. Upon the adop- 
tion of the motion, a committee was appointed, consisting of Mr. 
Small and Professors Porter and Rusby. 

Dr. Albert Schneider read his announced paper on “ The Bio- 
logical Status of the Lichens,” maintaining that they are entitled 
to recognition as an independent group, and not to be classed as 
a division of either Algae or Fungi. It will be published in a 
subsequent issue of the BuLLETIN. 


185 


Index to recent Literature relating to American Botany.* 


Alwood, W. B. Ripe Rot or Bitter Rot of Apples.. Bull. Va. Agric. 
& Mech. Coll. (II.) 3: 59-82. p/. 2. My. 1894. 
Description of Gloeosporium fructigenum Berk, with a bibliography. 
Bailey, L. H. Experimental Evolution amongst Plants. Am. Nat. 
29: 318-325. Ap. 1895. 
' Argument that species have repeatedly been evolved by cultivation. 
Bastin, E. S. Structure of Veratrum viride. Am. Journ. Pharm. 
67: 196-203. f. r-6.:. Ap. 1895. 
Brandegee, T. S. Mimulus Clevelandit. Gard, & For. 8: 134. f. 
20. 3 Ap. 1895. 
A new species from Southern California. : 
Britton, N. L. Publication by Signatures. Erythea, 3: 50-52. 1 
Mr. 1895. 
With notes on the subject by E. L. Greene. 
Clinton, G..P. Relationship of Caeoma nitens and Puccinia Peckiana. 
Bot. Gaz. 20: 116, 117. 16 Mr. 1895. 


Cogniaux, A. Orchidaceae.—II. Flora Bras. 117: 158-318. £d. 
35-75-15 Ja. 1895. 

Durand, E. J. Sporangial Trichomes in certain Ferns. Bull, Torr. 
Bot. Club, 21: 408. 1894. 


Durand, E. J. The Development of O/idium, one of the Chytridia- 
ceae. Bull. Torr. Bot. Club, 21: 410. 1894, 
Describes briefly the development of O/pidium entophytum, A. Braun, in the 
cells of Spirogyra. 
Gibson, W.H. Welcomes of the Flowers. Harper’s Mag. 1894: 551 


—556. figs. Mr. 1894. 


Discusses relation of insects to flowers, wild and cultivated. 

Golden, K. E. Movements of Gases in Rhizomes. Proc. Am. Assoc. 
Ady. Sci. 43: [reprint, pp. 10]. 1894. 

Graves, J. A., Chairman. The Pteridophyta of North America, 
north of Mexico. Linnaean Fern Bull. 9: pp. 23. Binghampton, 
1895. 


A numbered list of species and varieties. 


* It is requested that omissions from this list be communicated to the editors. 


186 


Greene, E. L. Ceanothus leucodermis, n. sp. Kew Bulletin, 97: 15. 
Ja. 1895... 


Description of a new species from California, found in Kew Herbarium; with 
discussion of C. d aricatus. 
Greene, E. L. Corrections in Nomenclature-—VI. Erythea, 3: 36. 
1 F. 1895. 


Greene, E. L. Novitates occidentales.—XI. Erythea, 3: 44~-49. 

1 Mr. 1895. 

New species of Ranunculus, Roripa, Tissa, Trifolium, Raillardclla, Crepis, 
Allocarya, Collinsia and Habenaria. 

Greene, E. L. Novitates occidentales.—X. Erythea, 3: 17-24. 

1 F. 1895. 

New species in Vicia, Lupinus, Trifolium, Thermopsis, Ranunculus, Erigeron, 
Pyrrocoma, Senecio, Agoseris, Phacelia and Linanthus. 

Greene, E. L. Observations on the Compositae.—VIII. Erythea, 3: 

6-15. 2 Jl. 1895. 

Discusses Aplopappus and Chrysoma with several transfers of species. Establishes 
the genera Stephanodoria, Petradoria and Osbertia for previously described species. 
Greene, E. L. Some Species of Dodecatheon. Erythea, 3: 37-4°- 

1 Mr. 1895. 

Describes D. radicatum, D. alpinum, D. tetrandrum Suksdorf and D, conjugens. 
Hansen, G. Flora of the Seguoia Region collected in the Counties of 

Amador, Calaveras and Alpine, State of California. Pamph. pp. 14- 

1 Ja. 1895. 

A list of several hundred species. 

Hasse, H. E. Lichens of the Vicinity of Los Angeles.—I. Erythea, 

3: 41-44. 1 Mr. 1895. 

Havard, V. Food Plants of the North American Indians. Bull. 

Torr. Bot. Club, 22: 98-123. 27 Mr. 1895. 

Hooker, J. D. Schinus dependens. Curt. Bot. Mag. 51: pé. 7400. 

Mr. 1895. 

Native of South America. 

_ Hooker, J. D. Weltenia candida. Curt. Bot. Mag. 51: pl. 7405. 
Mr. 1895. 


Native of Mexico and Guatemala. 

Kelsey, F. D. Some Field Notes. Bot. Gaz. 20: 117-118. 16 Mr. 
1895. 

Kenyon, F. C. In the Region of the new Fossil, Daemonelix. 
Am. Nat. 29: 213-227. pl. rz. figs. Mr. 1895. 


From the mode of growth the analogies of the organism are thought to be “ among 
some of the lower plants and grasses,” 


187 


Knoblauch, E. Die Nomenclature der Gattungen und Arten. Bot. 
Centralb. 61: 1-6. Ja. 1895. 

Lagerheim, G. de. See Patouillard, W. 

Lemmon, J. G. A new Calochortus. Erythea, 3: 49-50. 1 Mr. 
1895. 


C. codlinus from Central California. 

Linsbauer, L. Beitrage zur Vergleichenden Anatomie der Capri- 
foliaceen. Verh. Zool. Bot. Gesel. Wien, 45: 43-68. A/. 6. 1895. 

Meehan, T. /r’s cuprea. Meehan’s Month. 5: 61-62. p/. g. Ap. 
1895. 

Miller, A. W., Beringer, G. M., and Crawford, J., Committee. 
Biographical Sketch of Dr. J. Bernard Brinton. Bull. Torr. Bot. 
Club, 22: 93-97. Portrait. 27 Mr. 1895. 

Muller, J. Graphideae Eckfeldtianae in Louisiana et Florida lectae, 
additis observationibus in Graphideas Calkinsianas ejusdem regionis. 
Bull. Herb. Boiss. 3: 41-50. F. 1895. 

Pammel, L.H. Botany of the Russian Thistle. Bull. Iowa Agric. 
Coll. Exper. Sta. 26: 8-25. Af. 8 1894. 

Pammel, L.H. Further Notes on Cladosporium carpophilum. Proc. 
Towa Acad. Sci. 1: 92, 93. 1894. 

Pammel, L. H. Powdery Mildew of the Apple. Proc. Iowa Acad. 
Sci. 1: 92. 1894. 

Patouillard, W. et Lagerheim, G. de. Champignons de I’Equa- 
teur. Bull. Herb. Boiss. 3: 53-74. p/. 2. F. 1895. 


Descriptions of fifty new species. 

Robertson, C. Flowers and Insects.—XIII. Bot. Gaz. 20: 104-110. 
16 Mr. 1895. 

Robertson, C. The Philosophy of Flower Seasons and the Phaeno- 
logical Relations of the Entomophilons Flora and the Anthophilous 
Insect Fauna. Am. Nat. 29: 97-117. p/. 8. F. 1895. 

Robinson, B. L. On the ‘List of Pteridophyta and Spermatophyta 
of Northeastern America,’’ prepared by the Nomenclature Committee 
of the Botanical Club. Bot. Gaz. 20: 97—103. 16 Mr. 1895. 

Sargent, C. S., Editor. The Cherokee Rose. Gard, & For. 8: 114. 
J. 18. 20 Mr. 1895. 

Sargent, C. S., Editor. The Winter Aspect of Trees. Gard. & 


& For. 8: 121. f. 79. Mr. 1895. 
With illustration of the Beech-tree. 


188 


Schumann, K. Phyllocactus Thomasianus. Monats. Kakteenk. 5: 

6. pl. 20 Ja. 1895. 

Stephani, F. Hepaticarum species novae.—VII. Hedwigia, 34: 43- 

Qo." 15. F. 2895. 

New species of Heréerta, Hy grobiella, Hymenophytum and Famesoniella includ- 
ing Herberta dura from Straits of Magellan, 1. pumila from Argentina, and Fames 
_ ontella Leiboldiana from Mexico. 

Stevens, W.C. Apparatus for Physiological Botany. Bot. Gaz. 20: 

89-96. pi. 9-12. 16 Mr. 1895. 


Terry, W. A. Diatoms of the Connecticut Shore.—VII. Am. Mo. 
Mic. Journ. 16: 41-47. 1895. 

Underwood, L. M. The Classification of the hes ree aes Bull. 
Torr. Bot. Club, 22: 124-129. 27 Mr. 1895. 


Underwood, L. M. The Relation of the Red Cedar to our Orchards. 
Trans. Indiana Hort. Soc. 1894: 81-84. 1895. 
Popular account of Gymsnosporangium macropus and G. globosunt. 

Walker, Ernest. How Plants use Spines and Prickles. ‘Trans. 
Indiana Hort. Soc. 1894: 86-92. 1895. 


Wildeman, E. de. TZvrentepohita Pittieri. La Notarisia, 9: 6-7- 
1894. 
A new species from the leaves of various plants in the forests of Costa Rica. 

Wiley, H.W. Sweet Cassava ;: its Culture, Properties and Uses. Bull. 
Div. Chemistry, U. S. Dept. Agric. 44: pp. 16. f I. pl, 2. 1894. 
Discussion of Fatropha Manihot. 

Wilson, J. The Russian Thistle—(Salsola Kah Tragus). Bull. lowa 
Agric. Ex. Sta. 26: 3-7. 1894. : 

Wooten, E. D. New Mexico Weeds. Bull. N. Mex. Agric. Exp. Sta. 
rte 90. 7. a5. Oe. 1864. 

Wright, J. S. A Guide to the Organic Drugs of the United States 
Pharmacopoeia. r2mo., pp. 118. Eli Lilly & Co. 1895. 


Gives brief accounts of drugs and the species yielding them, and a conspectus of 
the natural orders of plants, 


Contributions from the Herbarium of Columbia College. 


No. 4. 
No. 5. 
No. 6. 


« 


[The numbers omitted from this list are out of print. ] 
A List of Plants Collected by Miss Mary B. Croft at San Diego, Texas. By 


N. L. Britton and H. H. Rusby (1887),: .° 3 2-42 es 25 cents. 
New or Noteworthy North American Phanerogams. By N. L, Britton 
CIOBS SS ee ee ee ek ee oe ee 25 cents, 


An Enumeration of the Plants Collected by Dr. H. H. Rusby in South 
America, 1886-1887. By N. L. Britton. (Twenty-three parts published ; 
not yet completed.) 

The Genus Hcoria of Rafinesque. By N. L. Britton (1888),  . 25 cents. 

A List of Plants Collected by Dr. E. A. Mearns at Fort Verde and in the 
Mogollon and San Francisco Mountains, Arizona, 1884-1888. By N,. L. 
Britton. 

The General Floral Characters of the San Francisco and Mogollon Mountains 


and the Adjacent Region. By H. H. Rusby (1888),. . . . . 25 cents. 
Preliminary Notes on the North American Species of the Genus 7issa, 
Adans.: By N.-L. Britton (1880), 66 Gee . . . 25 Cents, 
New or Noteworthy North American Phanerogams, II. By N. L. Britton 
C1SSQ) eo ee ee Ear a ee eee 25 cents. 
A Descriptive List of Species of the Genus Heuchera. By Wm. E. Wheel- 
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Sai A et ES ae ee ae Oe a ig a i Se ee, ce eS Re a es Be 


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Notes onthe North American Species of Eriocaulee. By Thos. Morong 
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New or Noteworthy North American Phanerogams, V. By N. L. Britton 
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Review of the North American Species of the Genus Xyris. By Heinrich 
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Note on a Collection of Tertiary Fossil Plants from Potosi, Bolivia. By N. 
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Ranunculus repens and its Eastern North American Allies. By N. L. 
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A Preliminary List of American Species of Polygonum. By John K. Small 
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PROF. N. L, BRITTON, 


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The following botanical works, owned mainly by the late Mr. Justus F; Poggen- 


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(CONTENTS: 


PAGE, ; 
The Biological Status of Lichens: Albert | Acetate in ite st bea 
SERRE oe oI EO ag Sencwe Manes Sot FETT 
New Species of Fungi: Ghee. Pack os 198| laceae: Anna Murray Vail ...... 2: 
Hypericum boreale and related Species: The Genus Zenobia Don: Charles Bouts’ 
ect e Beat See ate eae pear: Din egy bate ee 

urther Remarks on Family Nomenclature Borantcat Norss.—Nomenclatural + es 233 
te eo wis ye ee See 
Peer rer et pneicn Pham 


taceous (Dakota | Group) of Kansas: Be 
reas — 230s 237) Ha rte Testes 


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The Biological Status of Lichens. 


By ALBERT SCHNEIDER. 


For reasons to be enumerated, our present knowledge of 
lichens is very imperfect. Lack of attention is not the cause, as 
the voluminous literature on the subject will testify. The refer- 
ences, authentic and otherwise, number many thousands. It 
would be an endless task to bring together all the monographs, 
treatises, and especially the “ fragments,’ referring to lichens. 
Lichenologists of ante-Schwendenerian time supposed that the 
- question of the true nature of lichens and their position in the 
vegetable kingdom was permanently settled. Nothing was left 
for them to do but to issue “fragments” describing presumably 
new species and varieties. Collectors set to work in widely dis- 
tributed and circumscribed areas to add their mite to the heap of 
confusion. We all know that the ultimate aim of science is to 
systematize; but no system can be formed from unknown mate- 
rial, whatever it may be. A scientist's first duty then is to study 
(as far as possible) his material before attempting to classify it. 
_ This careful studying of material is what the mass of lichenolo- 
gists have heretofore failed to do. It is not my intention to enter 
into a historical review of lichenology, as that has already been 
thoroughly done by Krempelhuber and others. With the above 
introduction I shall now attempt to make somewhat clearer the 
_ Present status of general lichenology. 


1906 


The epoch-making researches of de Bary, Schwendener, Bor- 
net and others have conclusively demonstrated the dual nature of | 
the lichen structure ; that is, it consists of a colorless hyphal por- 
tion homologous with certain filamentous fungi; and a green 
celled portion homologous with certain low forms of algae. The 
specific algal homologues have been pretty accurately studied out. 
Concerning the specific homologues of the fungal portion our 
knowledge is less accurate. But in regard to both symbionts 
we are certain of their originally independent ancestral 
forms. This theory of Bornet and Schwendener has from the 
_ very first met with strong opposition from nearly all lichenologists 
(taxonomists, so called). Even a considerable number of phys- 
iologists and morphologists misconstrued evident facts with un- 
scientific perversity, notably Itzigsohn, Famintzin and Baranetzky. 
Even to this very day there are a number of lichenographers who 
persist in ignoring or directly opposing Schwendener’s theory. 
This is simply additional evidence of the correctness of the state- 
ment “ None are so blind as those who do not wish to see.” 

It would be useless to repeat the arguments based upon actual 
experimentation which conclusively prove the correctness of 
Schwendener’s theory. There is, however, a question which 
Schwendener and his immediate followers have almost unani- 
mously answered wrongly and that is the question of the true 
position of lichens in the vegetable kingdom. During the ante- 
Schwendenerian time, beginning with the earliest periods, most 
lichenologists looked upon lichens as autonomous structures, 
though this conclusion had no scientific basis founded on mor- 
phology and physiology. Their characteristic distribution and 
marked macroscopic appearance were thought sufficient to make 
'them a distinct group. Schwendener assumed that lichens re- 
sembled certain groups of fungi, both in structure and in their 
manner of growth, and should therefore be classed under fungi as 
ascolichenes and basidiolichenes. It is much to be regretted 
that Schwendener did not see his mistake in time to avoid con- 
fusion and unnecessary and unwarranted opposition to his theory. 
I will frankly admit that I formerly thought it most expedient to 
classify lichens as modified fungi. - But having since made a 
special study of lichen morphology, I now consider such a pro- 


191 


cedure both unscientific and impracticable. This idea is not 
original with me, nor do I stand as its only advocate. Many of 
the most recent scientific writers, notably Reinke, strongly up- 
hold this view. I shall briefly consider lichens from the stand- 
points of morphology and physiology to show that they can only 
be treated as autonomous structures having a phylogeny of their 
own. I shall not consider it worth while to enter into word quib- 
bling as to the meaning of “autoaomy.” If such were my desire 
I might well deduce good arguments to show that angiosperms 
are not autonomons, that they are simply modified gymnosperms, 
that gymnosperms are modified pteridophytes, pteridophytes 
modified mosses, etc., finally having it reduced to a primal cell; 
this cell could still be reduced to organic matter, and that to in- 
organic matter, etc. I shall consider as autonomous any compre- 
hensive group of organisms having morphological and physiologi- 
cal characteristics differing from those of any other comprehensive 
group of organisms ; meaning by comprehensive group any col- 
lection of allied individuals comprising natural species, genera and 
families. I will first show why lichens cannot be classed as fungi. 
The method of reproduction in lichens is wholly different from 
that of fungi. It is true that lichen spores have a strong morpho- 
logical resemblance to those of their probable homologues among 
the fungi. Functionally they differ widely. They cannot be con- 
sidered specific reproductive organs of lichens as they formerly 
were of fungi, because they are not capable of developing into a 
complete mature lichen or- even a fungus. They will indeed de- 
velop a mycelial network which will however not produce spores 
unless associated with its symbiotic alga. They can only be 
looked upon as degenerate reproductive organs of their fungal an- 
cestors. This degeneracy is the more marked as we ascend the - 
scale of lichen development. Taking one of the lower types, as 
exemplified in Pyrenula, we find spore organs having almost typical, 
fungal characters; that is, apothecia are prominent, paraphyses 
distinct and numerous, spore wall rather thick and colored, spores 
distinctly septate. These are the usual characters of fungal spore 
Organs. Considering some of the higher types as exemplified in 
Parmelia and Cladonia, we find apothecia few or wholly wanting ; 
when apothecia are present the asci are few in number, spore wall 


192 


comparatively delicate and colorless, spores non-septate. There 
is no doubt that lichen spores still play a part in lichen reproduc- 
tion. Zhis can however only take place when the spore with the 
specific algal symbionts are placed in a suitable environment. That 
is, spores and algae must fall upon a spot where the algae can 
maintain existence independently until such time as the spores 
shall develop a mycelial network with haustoria with which to sur- 
round the algae, thus forming the beginning of a new lichen thallus. 
Should, for example, the spores with the requisite algae fall upon 
a dry rock the algae would die, and if the spores should subse- 
quently develop there would be no algae with which to form a 
lichen. From this it is evident that lichen spores must be very 
unreliable as assistants in lichen reproduction. From the very 
nature of things, lichen spores are not true reproductive organs of 
lichens, hence their tendency to degenerate. 

The question whether lichen spores are sexual or asexual 
products is still unsettled. The observations of Stahl in’the case 
of Collema microphyllum have not yet been verified. If his obser- 
vations prove to be correct, then we may assume that lichen spores 
are sexual products. I am, however, strongly inclined to believe 
that Stahl’s observations were probably erroneous. From numer- 
ous examinations of so-called “spermagonia,” I believe them to 
be parasitic fungi, of which the “spermatia” are the spores. 
From a rather hasty comparative study it seems probable that 
their homologues are to be found in Septoria or allied genera. 
For example, Septoria Speculariae presents the general morpho- 
logical appearance of spermagonia. The fact that we readily re- 
cognize Sep/oria as parasitic fungi lies only in the nature of things. 
In case of Septoria the morphological and physiological contrast 
between host and parasite is great, while in the case of spermago- 
nia this contrast is only slight. No one would ever think of re- 
cognizing Septoria Speculariae as the male reproductive organs of 
Specularia perfoliata, upon which it lives. Such a suggestion 
would certainly be highly ridiculous. Then why should sperma- 
gonia of lichens be recognized as male reproductive organs, espe- 
cially since no one has demonstrated that they play such a func- 
tion? According to Wiesner, spermatia do develop a mycelium 
which finally produces spermagonia. From this the true nature 


195 


of spermagonia may safely be conjectured. To say the least, it 
is certainly unreasonable to assume that spermatia will at one 
time function as non-sexual spores, and at another time as male 
sexual organs. Further investigations will reveal the true nature 
of things. The sooner this is accomplished the better, as many 
lichenologists have already made the deplorable mistake of consid- 
ering spermagonia as important characters in lichen classification. 
To classify plants according to the characteristics of the parasites 
found upon them would certainly be a questionable procedure in 
modern taxonomy. 

Other characteristics which distinguish lichens from fungi are 
the presence of various chemical compounds, notably lichenin, 
which is never found in fungi. 

Characteristics which distinguish lichens from fungi also distin- 
guish them from algae. There is certainly less similarity between 
an alga and a lichen than there is between a fungus and a lichen, 
though several attempts had been made to classify them as algae. 
In general it may be stated that lichens resemble algae only in so 
far as the algal symbiont resembles algae. The differences will be 
brought out in the discussion of those characters which separate 
lichens from both algae and fungi. For convenience sake I wil! 
separate these characters into morphological and physiological. 
These are the characters which fully establish the autonomy of 
lichens. 


MORPHOLOGICAL. 


Lichens, macroscopically considered, have such a peculiar ap- 
pearance that the most superficial observer is naturally led to 
suppose that they form a group by themselves. They are found 
in places where neither alga nor fungus can exist alone. Especially 
peculiar is their ability to resist low temperatures. Freezing only 
checks their growth. A temperature of —40 ° C. does not kill them. 
Such crude observations are however not sufficient to establish 
their individualism. 

The lichen thallus is of special interest to the morphologist 
since this structure is typically lichenological. It always consists 
(Of the hyphal and algal symbionts. The algal symbiont is usually 


More centrally located, being surrounded by the hyphae of the 


194 


fungal symbiont.. Three types of jichen thallus may be recognized, 
namely, the crustaceous, the foliaceous and the fruticulose. The 
crustaceous type is the most rudimentary and cannot be said to 
to have even a dorsiventral structure, though one would naturally 
expect this from the nature of things. The lower surface differs 
only in having more numerous extended hyphal filaments to en- 
able it to adhere more firmly to the substratum as well as to take 
up soluble food materials. The second type already indicates a 
considerable advance in the evolution of the lichen thallus. It is 
typically dorsiventral. Dorsal and ventral layers are semi-cortical 
in structure ; that is, the hyphal cells are closely united and have 
only few air passages. Between these two layers is a layer 
of loosely interwoven hyphal tissue in which are imbedded the 
algae. From the lower surface extend the rhizoids. On the 
_ upper are found the apothecia (with exceptions, example, Vepro- 
muum) and soredia, besides the so-called “ spermagonia” and oc- 
casionally accidental fungal and algal parasites. The third type 
(as exemplified by the vertical thallus of Cladonia and Zhamnolia) 
shows a typical radial structure. Numerous examples showing 
the gradual gradation from the dorsiventral to the radial type can 
be found. In the radial type there is an outer semi-cortical layer, 
which usually differs from that of the dorsiventral cortical layer 
in that itis more compact. The fungal cell walls have become 
somewhat gelatinized and adhere very closely. Next to this layer, 
on the inside, is the layer of loosely interwoven hyphae containing ~ 
the algae. The third and innermost layer consists of longitu- 
dinal closely united hyphae. Sometimes this thallus is hollow in 
the center, sometimes solid, containing a central core of closely 
united longitudinal hyphae. 

Soredia are also typical lichen structures. They are very 
numerous in the higher forms of lichens (example, Parmelia sore- 
diata), and are found on the dorsal surface of the thallus, more 
frequently near the margin. Each soredium isin reality a miniature 
thallus. It is usually spherical in form, the outer layer consisting 
_ of closely united hyphal cells; the central portion consists of algal 
cells and loosely interwoven hyphal filaments. Soredia contain 
all the elements necessary for the development of a new lichen. 

From what has been stated above, apothecia can not be looked 


4 195 


upon as typical lichen structures, yet their morphology is of great 
importance in the consideration of lichen evolution and classifica- 
tion. Some of the changes in apothecia indicating a probable 
higher or lower stage of development have. already been referred 
to. AsI intend to consider these changes more particularly in 
a future paper on lichen classification, I shall at present omit 
further discussions. I shall now briefly consider the physiological 
characters which distinguish lichens from both fungi and algae. 


PHYSIOLOGICAL. 


In their method of growth lichens stand alone, The two 
symbionts form a microcosmos which is enabled to perform the 
necessary life functions which were originally inherent in both, 
and in addition they have acquired new characteristics during their 
phylogeny as lichens, which unmistakably stamp them as autono- 
mous structures. . As a unit they can now exist where neither sym- 
biont could exist alone. In spite-of this intimate mutualism, it 
is not at all likely that the fungal symbiont is w/ol/y dependent 
upon the algal symbiont for its food supply. For example, a 
lichen spore may develop to a considerable extent as a sapro- 
phyte upon: decaying wood, humus, and other dead organic mat- 
ter; nor is it at all likely that a.lichen can develop upon purely 
inorganic matter, as, for example, pure quartz crystals. Of course, 
the spore, with the requisite algae or a soredium, has bound up 
within itself a certain amount of extra food material, which enables» 
development to begin in the absence of all organic matter. The 
mycelial network then forms a structure for collecting within its 
meshes organic substances, carried to it by air and water currents ; 
_ this allows growth to continue. No amount of food supply will, 
however, allow the fungal symbiont to mature without its algal 
_Symbiont, excepting perhaps the lowest forms. Thus we see that 
mutualism of fungus and alga is necessary to form a lichen. 
The fungal symbiont, considered by itself, still retains its ancestral 
function as a saprophyte ; in addition it has acquired the habits Or. 
_ 4 semi-obligative parasite upon its algal symbiont. The algal 
_ Symbiont, which has the function of chlorophyll-bearing plants in 
general, that of assimilating carbon, must be looked upon asa 
| facultative parasite, since it can exist and mature independently of 


196 


its fungal symbiont. This has been repeatedly demonstrated ex- 
perimentally. Considered as a unit, the fungal portion of the 
lichen supplies the algal with water, the necessary mineral sub- 
stances, N., O. and H., from the underlying substratum and air. 
The algal symbiont as a result of this unusual supply of food ma- 
terials, forms an extra amount of carbon and nitrogenous com- 
pounds, which is assimilated by the fungal symbiont. Thus it 
is seen that the benefit derived from this association is mutual. 
The term “mutualism,’ proposed by Tubeuf, is very appropriate 
and may well supplant the equally correct but more complex ex- 
pression “ mutualistic symbiosis,’ proposed by Frank. 

Reproduction by means of soredia stands without a parallel 
in the vegetable kingdom. They are of course asexual, and are 
formed in the algal zone of the lichen thallus by the symbiotic 
association of algae and hyphae. They may be designated as 
mutualistic brood buds. They are really vegetative reproductive 
organs, and on that account the objection may be raised that they 
are not true reproductive organs. I shall not here enter into a 
discussion on the ultimate difinition of reproductive organs. The 
fact remains that soredia alone contain the necessary elements for 
JSorming a new lichen. There is no doubt that the great majority 
of lichens are reproduced from soredia; in fact, this is the only 
means of reproduction in some species. The outer semi-cortical 
hyphal layer of soredia enclosing the algae maintains a sufficient 
‘degree of moisture to enable them to lie dormant for a long 
period of time, or until conditions are suitable for their develop- 
ment. They are certainly far more reliable than spores (associa- 
ted with the necessary algae) as reproductive organs. In fact, as 
Reinke has indicated, lichen reproduction by the aid of spores is 
in most respects similar to reproduction by means of soredia. In 
Endocarpon pusillum, for example, some of the algae are ejected 
and with the spore adhere to the mucilaginous spore wall, thus 
forming a sort of soredium in which the spore represents the 
fungal element. If the algae are not ejected with the spores, the 
chances for developing a new lichen must indeed be slight, for 
reasons already given. 

In my opinion sufficient reasons have been given why lichens 
cannot be classed with fungi, much less with algae, and must 


197 


therefore be considered as an independent group. Based upon 
morphological and physiological considerations this group would 
naturally belong midway between fungi and algae. 

In conclusion I shall offer a few suggestions on the probable 
origin and phylogeny of lichens. There is little doubt that vari- 
ous subdivisions of lichens indicate a polyphyletic origin. Of 
this polyphylogeny either one or all of several forms may have 
- occurred. For example, in ascolichenes, a certain fungal type may | 
have (during its phylogenyas a lichen symbiont) become so modi- 
fied by its symbiosis with a given algal type, as to enable it to as- 
sociate with other algal species; or it may be that the same algal 
type became adapted to one or even several fungal types. As to 
what the conditions actually were we are at present scarcely able 
to say. Of one thing we are, however, certain and that is that a 
lichen is the result of the mutualistic association of a fungal and 
an algal type. Though in general I agree with Reinke as to the 
origin of lichens, yet I am not so ready to assume (theoretically) 
that Co//ema represents the oldest lichen type. Cod/ema is the re- 
sult of the symbiotic association of the alga Mostoc with some 
fungus whose ancestral type is not definitely known. The mass 
of the lichen structure consists of the alga. As compared with 
some other lichens the following are some of the reasons why 
Collema does not represent the lichen prototype: I. The alga 
has undergone considerable change by way of adapting itself to 
new environments. Originally it was no doubt accustomed to a 
high degree of moisture (as is its present homologue, Nostoc com- 
mune), while in its present form as a lichen it is able to exist on 
tree trunks, rocks, etc., as most other lichens. 2. Its thallus 
shows a considerable degree of differentiation, as exemplified in 
the closely allied genera Hydrothyrio, Polychidium, Leptogium and 
Mallotium. 3. Spores have probably become considerably degen- 
erated as indicated by their thin colorless walls, and in many 
Cases indistinct septae. As a rule apothecia are few, though there 
are exceptions to this. 4. Soredia, though not numerous, are more 
frequently present than in many other lichen forms. 

The above are the main reasons why Collema is perhaps not 
the prototype of lichens. In my opinion the true prototype of 
oe lichens is perhaps to be found in those structures which were for- 


198 


merly recognized as pseudo-lichenes. They may be observed on 
nearly every tree trunk, on fences, rocks, pavements, etc.; in fact, 
anywhere where the lower forms of algae (especially Protococcus 
viridis) can exist. Examination of these algae will find them 
usually associated with fungal hyphae, sometimes forming ex- 
tended thin thallus-like layers. The structure formerly recognized 
as Lepra viridis is an excellent example. Though apothecia are 
never found, yet I am inclined to believe that in so-called Lepra 
we find the beginnings of a future lichen. At least there are 
many lichens which show an inferior structure as compared with 
Collema, and for that reason are ‘perhaps nearer the prototype. 
As an example we may mention Pyrenula. Verrucaria perhaps 
represents a degenerate type rather than a lower type of an as- 
cending series, as is indicated by.a rudimentary thalline structure 
associated with rudimentary or degenerate apothecia, spores and 
paraphyses. Whether a given lichen represents a low type of an 
ascending series or a degenerate higher form is in many cases dif- 
ficult to decide; also the question as to the relative phylogenetic 
ages of various lichen groups. There is perhaps little doubt that 
basidiolichenes have hada much’ shorter phylogenetic history than 
~ascolichenes. There are many problems in lichenology which 
must be left to the conscientious morphologists and physiologists 
to solve. In fact, we know so little of the life history of individual 
lichens that the time for final specific arrangement has not yet 
come. We, however, know sufficient of lichens as a whole to 
give them a proper position in the vegetable kingdom which is 
in reality the first step toward establishing a lichen system. 
Their proper position I have attempted to indicate in this paper. 


New Species of Fungi. 


By CuHas. H, Peck. 


LeEPIOTA FULVopIScA. Pileus thin, convex or nearly plane, ob- 
tuse or umbonate, viscid. when moist, white, with the disk or umbo 
fulvous or tawny-brown; lamellae narrow, close, free, white; stem 
slender, flexuous, viscid, hollow, white or whitish, the base abruptly 
bulbous, the annulus thin, membranous, pure white; spores ovate- 
elliptical, .0003 to .0004 in. long, .00016 to .0002 in. broad, usu- 


199 


ally containing a shining nucleus and furnished with a slight 
apiculus at one end. 

Pileus 1 to 1.5 in. broad; stem 2 to 3 in. long, I to 1.5 line . 
thick. 

Plant fragile, growing among fallen leaves in woods. Pasa- 
dena, California. January. Prof. A. J. McClatchie. 

From Z. ilinita Fr. this species is separated by its tawny disk 
of the pileus, its membranous annulus, the bulbous base of the 
stem and the different shape of the spores. 

CiirocyBe pusILLA. Pileus at first hemispherical or convex, | 
then nearly plane, obtuse, sometimes with a very small umbo, dry, 
pruinose, grayish, flesh thin, whitish; lamellae narrow, close, ad- 
nate or decurrent, white; stem short, solid, pruinose, grayish ; 
spores subglobose or very broadly elliptical, .00016 to .0002 in. 
long. ‘ 

Pileus 3 to 8 lines broad ; stem about 6 lines long, scarcely 1 
line thick. 

Densely gregarious or subcaespitose. Onmanure. Pasadena. 
February. MeClatchie. . | 

: Apparently belonging to the tribe Disciformes and related to 
C. Bresadolae Schulz, but an exceedingly small plant for that tribe. 

CoLLyBIA ALBoGRISEA. Pileus fleshy, thin, convex or nearly 
plane, often somewhat irregular on the margin, glabrous, whitish 
or grayish, flesh white; lamellae broad, distant, adnate, a or 
whitish, the interspaces often venose; stem nearly equal, ho 7 
sometimes twisted, whitish or grayish, the lower part covered wit 
a dense whitish, grayish or yellowish tomentum, the upper ees 
naked or merely white-pruinose ; spores broadly elliptical, iene 
00024 in. long, .00016 broad, generally containing a single shin 
ing nucleus. 

Pileus about 1 in. broad; stem 1.5 to 2 in. long, I to 2 lines 
thick. 
Plant often caespitose and then the pileus irregular from mu- 
tual pressure. Among fallen leaves. Pasadena. January. Mc- 
Clatchie. 

The species belongs to the tribe Vestipedes. : 

MYcENA ELEGANTULA. Pileus membranous, conical or cam- 
‘ panulate, sulcate-striate, brown or purplish-brown ; lamellae distant, 


ae adnate with a decurrent tooth, whitish or pallid with the edge 


p ; : 
_Purplish-brown, the interspaces (in the dried specimens) aoirea wee 
transversely wrinkled; stem slender, hollow, glabrous, s 


200 


with a loose grayish fibrillose tomentum at the base ; spores oblong- 
elliptical, .00035 to .0005 in. long, .00016 to .0002 broad, usually 
containing one or two small nuclei. 
Pileus, 4 to 10 lines broad; stem I to 2 in. long, .5 a line thick. 
Gregarious or caespitose. Among fallen leaves under trees. 
Pasadena. December. McClatchie. 


This plant may be distinguished from the closely related JZ. 
atromarginata Fr. by its smaller size and the purplish tint to the. 


edge of the lamellae, and from JZ. purpurcofusca Pk. by its differ- 
ently shaped, longer spores. 


OMPHALIA SEMIVESTIPES. Pileus very thin, deeply convex or 
subcampanulate, glabrous, grayish-brown in the dried state, paler 
when fresh; lamellae rather broad, distant, arcuate, decurrent, 
white or whitish; stem hollow, white or whitish, the upper half 
glabrous, the lower half clothed with a white mycelioid tomentum ; 
spores elliptical, .0002 to .00024 in. long, .00012 to .00016 broad. 
Pileus 6 to 12 lines broad; stem about 1 in. long, 1 to 2 lines 
thick. 


Growing on much decayed wood. Newfoundland. May. 
Rev. A. C. Waghorne. 

This plant is apparently related to such species as O. setipes 
Fr. and O. grisea Fr. and belongs to the tribe Mycenariae. _It is 
easily recognized by the peculiar character of the stem. In the 
dried examples the upper glabrous part of the stem is shriveled 


and longitudinally striate, but the lower tomentose part is plump 
and even as in the fresh plant. 


HyGropuorus ELEGANTULUS. Pileus convex or nearly plane, 
glabrous, viscid, grayish-yellow or slightly tawny, flesh white ; 
lamellae distant, slightly decurrent, white; stem equal, solid, slightly 
floccose-squamulose at the top, elsewhere glabrous, glutinous, 
white or whitish, sometimes abruptly pointed at the base; spores 
elliptical, .0004 in. long, .0002 to .00024 broad. 

Pileus 1 to 2 in. broad; stem 2 to 3 in. long, 3 to 4 lines thick. 

Woods. Maryland. November. T. Taylor. 


The species belongs to the tribe Limacium, and is related to 


- H. discoideus Fr., from which it may be separated by its solid stem 
and larger spotes. : : ; 


ENTOLOMA FERRUGINANS.  Pileus fleshy, convex, obtuse or um- 
bonate, often irregular, hygrophanous, glabrous, shining, dark 
fuliginous or broccoli-brown, flesh whitish, fibrous and colored at 


201 


the surface; lamellae 4 to 7 lines broad, adnexed, easily splitting 
transversely, grayish-salmon, becoming clay-color; stem solid, 
glabrous, yellowish or cream-color, blunt at the base or sometimes 
attenuated and radicating ; spores subglobose, irregular or angular, 
-0003 to .0004 in. long. 

Pileus 2 to 6 in. broad; stems 3 to 4 in. long, 4 to 8 lines or 
more in thickness. 

Under oak trees. Pasadena. February. McClatchie. 

This plant is related to E. rhodopolium, but is separated from it 
by its darker color and solid stem. According to Prof. McClatchie, 
the fresh plants have a strong odor resembling that of ferric chlo- 
ride, and chemical tests showed the presence of iron. It is pro- 
nounced edible by him. 

LepToniA EDULIS. Pileus thin, convex or centrally depressed, 
with or without an umbo, velvety, dark-gray; lamellae rather 
broad, subventricose, adnexed ; moderately close, at first whitish 
or light drab, becoming flesh-color ; stem slender, hollow, colored 
_ like the pileus, often with an abundant white mycelioid tomentum 
at the base; spores subglobose, angular, apiculate at one end, 
0003 to .0004 in. long, containing a single large nucleus. 

Pileus 6 to 18 lines broad; stem 12 to 18 lines long, .5 to I 
line thick. 

Among grass and. weeds. Pasadena. _.January. | 

According to Prof. McClatchie, this plant when fresh has a 
nutty flavor and is edible. The velvety appearance of the pileus 
has disappeared from the dried specimens. In some, the margin 
of the pileus is striate, but in the fresh plant the margin is said to 
be even. 

Eccinia nicricans. Pileus thin, convex, umbilicate or cen- 
trally depressed, subzonate, unpolished, grayish-black ; lamellae 
broad, distant, decurrent, light-drab or brownish, becoming tinged 
with flesh-color; stem short, hollow, grayish-black, commonly 
with an abundant white mycelium ; spores angular, .0004 in. long, 
nearly as broad, containing a single large nucleus. : 

Pileus 6 to 18 lines broad; stem about 1 in. long, .5 to 1 line 
thick. 

_ Grassy ground. Pasadena. January. 

Prof. McClatchie’s notes say that this plant has the odor and 
flavor of butternuts and that it is delicious when cooked. Also 
that when fresh the pileus is tomentose and the margin even, but 
these characters are not clearly shown in the dried specimens. 


202 


PHOLIOTA ANOMALA. Pileus at first hemispherical or subconi- 
cal, then convex, glabrous, hygrophanous, broccoli-brown when 
moist, pale-yellow or cream-color when dry; lamellae adnate or 
slightly decurrent, subarcuate, pale becoming brownish-ferruginous, 
often white on the edge; stem cavernous or hollow with irregular 
transverse partitions, sometimes containing a cottony tomentum, 
whitish, with a slight evanescent annulus; spores elliptical, .0003 
to .0004 in. long, .00016 to .0002 broad. 

Pileus 8 to 18 lines broad; stem 1.5 to 2.5 in. long, 1 to 3 lines 
thick. 

Sticks and leaves lying on the ground. Pasadena. January. 
McClatchie. 

The species belongs to the tribe Truncigenae, section Hygro- 
phanae. It is well marked by its fugacious annulus, subdecurrent 
lamellae and peculiar cavernous stem. 

HEBELOMA FOEDATUM. Pileus fleshy, convex becoming plane 

or centrally depressed, glabrous, very viscid or glutinous, reddish 
_ cinnamon, flesh yellowish-white; lamellae subventricose, emar- 
ginate with a decurrent tooth, cinnamon-color, becoming mummy- 
brown; stem solid, equal or slightly thickened at the base, fibril- 


lose, paler than the pileus; spores broadly elliptical, .00024 to 
.0003 in. long, .00016 to .0002 broad. 


Pileus 1.5 to 3 in. broad; stem 1.5 to 2.5 in. long, 2 to 4 lines 
thick. 

Streets of Pasadena. December. McClatchie. 

The species is apparently related to H. firmum, H. testaceum 
and 7. glutinosum, from all of which it is separated by its small 
spores. Its viscid pileus causes dirt to adhere to it in such quan- 
tity as to give the plant a very defiled, unattractive appearance. 

FLAMMULA ANOMALA. Pileus deeply umbilicate or infundibuli- 
form, often irregular, glabrous, whitish; lamellae narrow, close, 


decurrent, pale-ferruginous ; stem short, irregular, whitish ; spores 
globose, brownish-ferruginous, .00024 in. broad. 


Pileus about I in. broad; stem 6 to 12 lines long. 

Plant commonly caespitose. Ground. Trexlertown, Pennsyl- 
vania. October. Dr. William Herbst. 

A whitish umbilicate pileus is unusual among species of Flam- 
mula. This plant appears to belong to the tribe Gymnotae. 

TUBARIA PALLESCENS. Pileus fleshy but thin, convex or nearly 


plane, sometimes slightly depressed in the center, glabrous, hy- 
grophanous, brick-red when moist, yellowish or cream color when 


2058 


dry; lamellae broad, adnate or slightly decurrent, tawny-buff, 
becoming brownish-ferruginous; stem slender, hollow, yellowish, 
with a white mycelium at the base; spores elliptical, .0003 in. 
long, .00016 broad. 

Pileus 5 to 10 lines broad; stem 12 to 18 lines long, .5 to 1 
line thick. 

Sticks and leaves under trees. Pasadena. January. McClatchie, 

When young, slight vestiges of a veil are visible, connecting 
the incurved margin of the pileus with the stem. 


PLUTEOLUs LUTEUS. Pileus thin, at first subovate, then convex 
or subcampanulate, glabrous, viscid, slightly striate on the margin, 
yellow ; lamellae numerous, close, free or but slightly adnexed, yel- 
lowish becoming ferruginous; stem slender, hollow, slightly 
thickened toward the base, striate at the top and there sprinkled 
with mealy particles, yellowish; spores elliptical, .0004 to .0005 
in. long, .00024 to .0003 broad. : 

Pileus 6 to 12 lines broad; stem 1.5 to 2.5 in. long, I to 2 
lines thick. Plant very fragile, gregarious. Under trees. Pasa- 
dena. December. McClatchie. 

The yellow color and viscid pileus are prominent characters of 3 
this species. In some of the dried specimens the lamellae appear 
free, in others slightly adnexed, but because of the viscid pileus I 
have referred the plant to the genus Pluteolus. 


CorTINARIUS viRGATUS. Pileus thick, fleshy, hemispherical or 
convex, obtuse or subumbonate, slightly viscid, ochraceous tinged 
with olive-buff, conspicuously virgate with reddish fibrils, flesh 
dingy-white; lamellae subdistant, adnexed, at first subcinnamon, 
then ochraceous-russet ; stem short, stout, solid, enlarged and 
fibrillose at the base, pale-ochraceous; spores subglobose or 
broadly elliptical, .00024 to .0003 in. long, .0002 to .00024 broad. 

Pileus 3 to 4 in. broad; stems about 2 in. long, 8 to 12 lines 
thick. 

Undér oak trees. Pasadena. February. McClatchie. 

This species is well marked by its stout habit and by the red- 
dish fibrils of the pileus. . 

AGaricus Cauirornicus. Pileus at first subconical, becoming 
convex, minutely silky or fibrillose, whitish, tinged with purple or 
brownish-purple on the disk, flesh whitish; lamellae close, free, 
Pink becoming. purplish, then blackisk-brown; stem rather long, 
Solid or stuffed, equal or tapering upward, distinctly and rather 
abruptly narrowed above the entire externally silky annulus, pallid 


204 


or brownish; spores broadly elliptical, .0002 to .00025 in. long, 
.00016 to .0002 broad. 

Pileus 1 to 3 in. broad; stem 1.5 to 3 in. long, 2 to 4 lines 
thick. 

Under oak trees. Pasadena. January. McClatchie. 

This fungus is similar in size, shape and habitat to A. hemor- 
vhoidarius, but it is unlike that species in color, in the adornment of 
the pileus and in its color not changing where bruised or broken. 


STROPHARIA BILAMELLATA. Pileus fleshy, convex, even, whitish 
or yellowish, flesh pure white; lamellae close, adnate, purplish- 
brown when mature; stem short, solid, white, with a well-developed 
pure white annulus which is striately lamellate on the upper sur- 
face; spores elliptical, purplish brown, .0004 in. long, .0002 to 
to .00024 broad. 


Pileus 1 to 2 in. broad; stem about 1 in. long, 3 to 4 lines 
thick. 

Streets of Pasadena. January. McClatchie. 

This fungus is remarkable for the lamellated upper surface of 
the rather thick membranous annulus. These lamellae are uneven 
on the edge and in some cases they appear to extend upward on 
the stem till they meet the true lamellae. The plant is said by its 
discoverer to be edible. The color of the young lamellae is not 
shown by the examples. 


HypHotoma tonerres. Pileus thin, campanulate, even or ob- 
scurely striate on the margin, fibrillose becoming glabrous, hygro- 
phanous, yellowish-brown when moist, brown or isabelline-brown 
when dry, the margin appendiculate with the very white floccose 
fugacious veil; lamellae narrow, close, adnate, white or whitish, 
becoming nearly black, often whitish on the edge; stem slender, | 
long, hollow, striate at the top, white, with a white mycelioid 


earth at the base; spores elliptical, 005 in. long, .0003 
road, 


Pileus 1 to 1.5 in. broad; stem 2 to § in. long, 1 to 2.5 lines 
thick. 


Plant fragile, growing among fallen leaves in very wet weather. 
Pasadena. September. McClatchie. 


The disk of the pileus is so thin and the stem so completely 
hollow to the apex that in the dried specimens there is a depres- 
sion or umbilicus in the center of the pileus. 


205 


PANAEOLUS INTERMEDIUS. Pileus campanulate or convex, even, 
glabrous, moist or hygrophanous, grayish-brown ; lamellae ascend- 
ing or subarcuate, subdistant, adnate, black* when mature ; stem 
slender, often elongated, hollow, grayish-brown, white-pruinose at 
the top; spores oblong-elliptical, .0005 to .0006 in. long, .00025 to 
.0003 broad. 


Pileus 6 to 12 lines broad; stem 2 to 4 in. long, .5 to 1 line 
thick. : 

Rich soil along gutters or in cafions. Pasadena. January. 
McClatchie. 


_ The margin of the pileus does not extend beyond the lamellae, 
and this character with the slender stem suggests the genus 
Psathyrella, but because of the absence of striae on the pileus it 
seems best to refer the plant to the genus Panaeolus. 

PANAEOLUS DIGREssUS. Pileus hemispherical or convex, gla- 
brous, bay-red; lamellae very broad, plane, distant, adnate, pur- 
_plish black with a white edge; stem short, floccose-fibrillose 
toward the base, striate at the apex, hollow, a little paler than the 
pileus ; spores broadly elliptical, .o005 to..0006 in long, .00035 to 
.0004 broad. 

Pileus 4 to 6 lines broad; stem about 1 in. long, 1 line thick. 
On dung. Pasadena. July. McClatchie. 3 

This plant also diverges from the generic character in its 
lamellae extending quite to the margin of the pileus, and in its 
unpolished stem. 

Coprinus caLyprratus. Pileus when mature adorned with a 
few grayish floccose scales and crowned with a persistent stellately 
Split membranous dingy-yellow or subtawny calyptra, radiate 

‘Striate to the disk, grayish-flocculent along the ridges of the striae, 
blackish ; lamellae free, dark lead color becoming black; stem 
equal, hollow, white, becoming blackish in drying except at the 
base, neither annulate nor -distinctly volvate; spores elliptical, 
black, .0006 to .o008 in. long, .00045 to .0005 broad. 


Pileus about 2 in. broad; stem 3 to 4 in. long, 2-to 3 lines 
thick. Open cultivated ground. Rockport, Kansas. August. 


E. Bartholomew. 

This species is well marked by the persistent membranous 
calyptra that adheres to the summit of the pileus. Its margin is 
split into four to six broad rays. The change of color in thestem 
is similar to that ascribed to the stem of C. sterguilinus Fr., but our 
Plant differs from that in its calyptra and in the absence of an an- 


206 


nulus and volva at the base of the stem. Only mature specimens . 
were seen, consequently the characters of the young plant remain 
unknown and the description to that extent is defective. 


Coprinus Jonesul. Pileus submembranous, campanulate be- 
coming broadly convex or expanded and split or revolute on the 
margin, very blunt or truncate at the apex when young, everywhere 
covered with tawny-gray or pale-cervine floccose scales which 
wholly or partly disappear with age revealing the striate surface 
beneath ; lamellae crowded, linear, free, at first white or whitish, 
becoming black ; stem equal or slightly tapering upward, minutely 
floccose, hollow, white; spores black, broadly elliptical, .0003 to 
00035 in. long, .00025 broad, with an apiculus at one end. 


Pileus 1 to 2 in. broad ; stem 2 to 3 in. long, 2 to 3 lines thick. 

Plant fragile, sometimes caespitose. In a cellar. Vermont. 
April. Prof. L. R. Jones. 

The species is closely related to C. fimetarius, of which it might 
easily be considered a variety, but it is easily distinguished by the 
truncate apex of the young pileus, the differently colored scales and 

the smaller spores. C. soboliferus Fr. has the pileus truncated at 
the apex, but it is a very different species. 


+ 


CopRINUS APICULATUS. Pileus membranous, campanulate or 
deeply convex, acute 01 apiculate, furfuraceous, plicate-striate to 
the disk, grayish; lamellae few, subdistant, reaching the stem, 
black ; stem filiform, glabrous, white; spores elliptical, black, .0003 
in. long, .00016 broad. 


Pileus about 3 lines broad; stem I to 1.5 in. long, scarcely 
_ half a line thick. Lewiston, Pennsylvania. Mrs. E. B. Noyes. 


BOLETINUS BOREALIS. Pileus fleshy, convex, obtuse or subum- 
bonate, brownish yellow, obscurely and somewhat reticulately 
streaked with reddish-brown lines; pores large, angular, unequal, 
slightly decurrent, brownish-yellow ; stem short, equal or slightly 
tapering upward, brownish-yellow with a whitish mycelioid to- 
mentum at the base ; spores oblong, .0004 to .0005 in. long .00016 
to .0002 broad. 
Pileus I to 2 in. broad; stem about 1 in. long. 
Sandy soil. Capstan Island, Labrador. October. Waghorne. 
The markings of the pileus appear as if due to the drying of 
a glutinous substance. The radiating lamellae and the transverse 


partitions of the interspaces are very plainly shown. Described 
from two dried specimens. 


207 


BoLetus INFLExuS. Pileus convex, glabrous, viscid, yellow, 
often red or reddish on the disk, the margin thin, inflexed, con- 
cealing the marginal tubes, flesh whitish, not changing color 
where wounded ; tubes rather long, adnate, yellowish, becoming 
dingy-yellow with age, the mouths small, dotted with reddish 
glandules ; stem rather slender, exannulate, solid, viscid, dotted 
with livid-yellow glandules; spores yellowish, .0004 to .0005 in. 
long, .00016 to .0002 broad. 

Pileus about 1 in. broad; stem about 2 in. long, 2 to 4 lines 
thick. 

Open woods. Trexlertown. September. Herbst. 

This Boletus belongs to the tribe Viscipelles. It is remarkable 
for and easily recognized by the inflexed margin of the pileus, 
which imitates to some extent the appendiculate veil of Boletus 
versipellis. It sometimes grows in tufts. The paper in which 
fresh specimens were wrapped was stained yellow. Boletus 
Braunii Bres. has an inflexed margin, but that is a much larger 
plant with a yellowish-brown pileus, a fibrillose stem and much 
smaller spores. 

PoLyporus ANcEPs. Effuso-reflexed or resupinate, inseparable 
from the matrix, firm, subcorky but flexible, white; pileus narrow, 
about 6 lines broad, laterally elongated or confluent, minutely 
downy, sometimes. rugosely pitted; pores minute, subrotund, 
commonly 2 to 3 lines long, the dissepiments obtuse; mycelium 
white, permeating the bark and wood. 

Dead trunk of hemlock, 7suga Canadensis. Stony Brook, 
Massachusetts. October and November. Prof. E. A. Burt. 

The plants are commonly resupinate, but sometimes reflexed, 
forming a narrow pileus about half an inch broad but extending 
laterally for several inches. They are suggestive of the first year’s 
growth of P. connatus Fr., but they do not revive the next year, 
and they have a different habitat. Though differing somewhat in 
texture they are apparently related to such species as P. semisupt- 
nus and P. semipiteatus, and with them they serve to connect the 
genus Polyporus with the genus Poria. 

Sparassis Herpsti.—Plants much branched, forming tufts 4 to 
5 in. high and 5 to 6 in. broad, whitish, inclining to creamy-yel- 
low, tough, moist, the branches numerous, thin, flattened, con- 
Crescent, dilated above and spatulate or fan-shaped, often some- 
what longitudinally curved or wavy, mostly uniformly colored, 
Tarely with a few indistinct, nearly concolorous, transverse zones 


208 


near the broad entire apices; spores subglobose or broadly ellip- 
tical, 0002 to .00025 in. long, .00016 to .0002 broad. 


Trexlertown. August. Herbst. 

The species is evidently closely allied to S. spathulata Schw., 
but differs especially in its paler color, with no rufescent hues, in its 
much more branching habit and in the absence of any distinct 
zones. 

BATTARREA ATTENUATA. Exoperidium unknown; endoperidium 
2 in. or more in breadth, the basal part hard, thick, even and con- 
cave beneath, convex above and somewhat coarsely reticulate by 
the bounding walls of broad shallow pits; stem 8 to 10 in. long, 
gradually attenuated toward the base, hard, almost woody, solid, 
rough except at the top with rather coarse spreading or reflexed 
scales, brown externally, rusty-brown within; spores globose, 
ferruginous, .0003 in. broad; threads of the capillitium destitute 
of spiral thickenings. 

Plants commonly growing in tufts of 3 to 5 individuals. 

Dry sandy soil. Nevada. Collected by C. W. Irish; com- 
municated by Dr. Thomas Taylor. 

The single dried specimen from which, with notes kindly com- 
municated by the collector, the above description was derived, 
was past maturity and destitute of any volva or exoperidium. 
The upper part of the endoperidium, which is apparently mem- 
branous, had nearly all disappeared, and but a mere remnant of 
the spores and capillitium remained. It is very unsatisfactory to 
attempt the description of a species from such imperfect data, yet 
the characters seen are so peculiar and distinct that I have been 
willing to strain a point in order to make this remarkable plant 
known. It does not agree rigidly with the characters ascribed to 
the genus Battarrea, differing apparently in the solid stem, the 
absence of spiral thickenings in the capillitium threads and in the 
coarsely pitted subreticulate hymenial substratum, so that it might 
easily be taken as the type of a distinct genus. 

The dried specimen has a strong, unpleasant odor, indicating 
its relationship to the Phalloideae. From the notes of Mr. Irish 
we learn that the long stem, which is about half an inch thick at 
the base and one and a half at the top, is almost wholly buried in 
the soil, and that the plants appear above the surface only in 
seasons after heavy snow falls, whose gradual melting has moist- 
ened the earth deeply. 


209 


TYLOSTOMA SEMISULCATUM. Peridium subglobose, usually a little 
longer than broad, 6 to 8 lines broad, Q lines long, glabrous above, 
ferruginous-tomentose on the lower half ; ostiolum entire, stem 
equal, about 2 in. long, even and glabrous or but slightly furfur- 
aceous on the upper part, the lower part longitudinally sulcate, © 

‘whitish; spores ferruginous, globose, .00016 to .0002 in. broad; 
threads of the capillitium colorless, not septate. 


Sandy soil. Nevada. Collected by C. W. Irish; communi- 
cated by T. Taylor. ares 

This species is separated from 7. mammosum Fr. by its peri- 
dium, which is tomentose on the lower half and not depressed, and 
by its stem, which is distinctly furrowed’in its lower half. 


Lycoperpon BEtui. Peridium about 2 in. broad, subglobose, 
sessile, grayish or yellowish-brown, exterior peridium continuous, 
adorned with numerous persistent hard angular irregular or sub- 
stellate warts which are smaller toward the base, separable at ma- 
turity from the inner peridium and falling away in flakes or patches; 
the inner peridium thinner, paler, glabrous; capillitium composed 
of long slender interwoven, slightly colored: and occasionally 
branched filaments, a little broader than the diameter of the spores; 
_ Spores globose, olive-brown, .00024 to .00028 in. broad. 


Rocky ground. Digges Island, Hudson’s Bay. August. Col- 
lected by R. Bell; communicated by Prof. J. Macoun. 

This puff-ball is peculiar in its continuous but warty exterior 
peridium, which is less friable than that of Z. separans, The peri- 
dium appears to rupture irregularly, as in Calvatia. It is also well 
marked by the dark olive-brown spore mass. 


Excipurina opscura. Perithecia cupulate, 14 to ¥ line broad, 
Sessile, black, the margin at first incurved, then erect, thin ; hyme- 
nium subgelatinous; spores subfiliform, curved, plurinucleate or 
obscurely pluriseptate, hyaline, .oo12 to .0018 in. long, .00016 

road, supported on short sporophores. 

Bark of hemlock trees, Tsuga Canadensis. Newfoundland. 


Waghorne. 


_ MEtrasmia imitans. Perithecia hypophyllous, membranous, 
variable in size and shape, commonly subelliptical or oblong, some~ 


What confluent in nerve-following lines, rugosely uneven, black, 


_ pening irregularly ; spores subcylindrical, straight or nearly so, 

_ Colorless, sometimes obscurely plurinucleate, .0008 to .oor2 in, 
long, .00016'to .0002 broad, oozing out and forming pallid globu~ 
‘ar or irregular masses or short thick tendrils. 


210 


Lower surface of living fronds of Freris aguilina. California. 
Prof. M. A. Howe. 

This fungus imitates Phydlachora pteridis in habitat and general 
appearance, and might easily be supposed to be the conidial state 
of it. The lines of the perithecia are more narrow and less prom- 
inent. The masses of discharged spores are rather large and very 
numerous and partly conceal the black perithecia beneath them. 


CAEOMA ABERRANS. Pustules suborbicular, slightly elevated, 
.5 to 1 line broad, at first covered by a whitish membrane which 
finally disappears revealing the mass of orange-yellow spores; 

spores subglobose or elliptical, smooth or nearly so, .0007 to .0009 

* in. long. 

Bark of living alder. Newfoundland. May. Waghorne. 

This fungus is peculiar in its habitat. The spores do not easily 
separate from each other, but possibly this is due in some measure 
to immaturity. 


ASPERGILLUS SUBGRISEUS. Grayish; sterile hyphae creeping; 
fertile erect, continuous, simple, .0003 in. thick, terminating above 
in an inflated subglobose vesicle .0012 to .0016 in. broad; sterig- 
mata none or obsolete; spores globose, .00016 in. broad. 

On Corticium amorphum. Newfoundland. Waghorne. 

This species is separated from the related A. griseus by its 


larger spores, continuous fertile hyphae and by the absence of 
distinct sterigmata. 


LerroGLossum LATuM. Club 3 to 6 lines long, nearly or quite 
as broad, soft when moist, rather fragile when dry, compressed 
and somewhat irregular, black; stem about as long as the club, 
black ; asci clavate-cylindrical, .0045 to .005 in. long, about .0005 
broad, 8-spored; spores crowded in the ascus, oblong or cylindri- 
cal, straight or slightly curved, continuous, obtuse, colorless, .OO! 
to .0o16 in. long, .0002 to .00024 broad; paraphyses colored, 
thickened at the top and sometimes recurved. 

Sandy soil. Labrador. September. Waghorne. 

A species very peculiar in its broad compressed club. The 
colored paraphyses are conspicuous under the microscope. They 
project slightly above the surface of the receptacle and give it a 
soft, almost velvety appearance. : 


VaLsa BREVIS. Pustules numerous, rather prominent, peri- 
thecia 10 to 20 or more in a pustule, nestling in the inner bark; 
ostiola even, black, barely emerging from and dotting or oblitera- 


211 


ting the orbicular or elliptical erumpent subpulverulent disk; asci 

-0008 to .oo! in. long; spores allantoid .00024 to .0003 in. long. 
Bark of balsam fir, Adies dalsamea. Labrador. Waghorne. 
This fungus is allied to but differs from Valsa Friesii in its 

larger pustules, more numerous perithecia and shorter asci and 


spores. The color of the disk is grayish or grayish-green. 


HyGropHorus NiGRIDIus. Pileus fleshy, convex, obtuse or 
subumbonate, glabrous, grayish-brown or black-brown, often a 
little darker in the center, flesh white; lamellae distant, decurrent, 
white; stem rather slender, solid, brownish, white at the top; 
spores elliptical, .0004 to .0005 in. long, .00024 to .0003 broad. 


Pileus I to 2 in. broad; stem 1 to 2 in. long, 2 to 4 lines thick. 

Gregarious or rarely two or three united at the base. Pine and 
fir tree woods. Prince Edwards Island. October and November. 
J. MacSwain. 

This fungus differs from A. caprinus Fr. in its smaller size, 
glabrous pileus and larger spores. A description of it was pub- 
lished in the Country Gentleman of November 29, 1894, but one 
is here given that it may be more readily accessible to mycologists. 

In many of the foregoing descriptions I have been obliged to 
give the colors indicated by the dried specimens. It is not likely 
therefore that they will in all cases agree rigidly with the colors of 
the fresh plant. 


| Hypericum boreale (Britton) and related Species. 


By EvucENE P. BICKNELL, 


At York Harbor, Maine, Hypericum Canadense L. abounds, 
Presenting itself in varying forms; Hypericum majus (Gray) Britton 
frequently grows with it, either strongly typical or with aberrant 
tendency; Hypericum mutilum L. is common also. 

More characteristic of the region, however, than any of these 
is a small species, which grows in abundance in open situations, 

about the muddy borders of pools or in moist sandy soil, and 
_ Shows some interesting lines of variation. Some fragmentary 
specimens, collected in 1888, were pronounced by Dr. Britton to 


212 
be the little known Hypericum Canadense var. minimum,* a plant 
which had never been well understood, but which, on account of 
its narrow leaves, had been associated unhesitatingly with /7. 
Canadense. Now, however, the study of a full series of specimens, 
collected at York Harbor in August, shows clearly that, though 
in one of its forms the plant certainly bears a strong general re- 
semblance to forms of H. Canadense, its real affinity is with 7. 
mutium, to which, indeed, it is nearly allied. Certain noteworthy 
differences, however, forbid its direct reference to that species, 
and give it an identity which is unmistakable when once the plant 
is understood. These differences, moreover, prove to be constant 
over a wide area; at least there is nothing to discredit them in 
specimens which show the plant to have an extensive distribution, 
ranging along the coast and islands of Maine into Nova Scotia, 
and westward through Canada. That it also extends southward 
in the mountains is attested by a specimen from Luzerne county, 
Pennsylvania, and one from ah elevated station in northern New 
Jersey. . 

It is to be noted of this New Jersey specimen that it shows : a 
near approach to muzi/um and may, perhaps, be taken as presump- 
tive evidence that the two plants intergrade. If such be the fact, 

however, it is, after all, only a part of the complementary fact that 
elsewhere the same plants have reached a condition of complete 
separation. The sub-species of one region has become a full 
species elsewhere in its range. 

In cases such as this of complete differentiation in one region, 
how are we to deal with intergradation in another? It is practicable 
to recognize a plant at the same time as a variety and as a species 

according as to whether it occurs in one or in another part of its 
habitat? It can only be said that if this condition of things 
exists in nature we cannot refuse to recognize it, and shall have to 
adjust our methods to it as best we may. But while we cannot 
dissect the facts of nature too delicately for certain purposes of 
botanical study, for other, more general purposes, we can often 
reach better practical results with duller tools. For general pur- 


*Dr. Britton has since shown that the name minimum of Choisy is not properly 


applicable to this plant, and has therefore designated it ve ‘pericum, Canadense vax. 
boreale. Bull. Torr. Club, 18: 365. 1891. 


213 


poses, therefore, what shall be our practice in cases similar to the 
one before us? The older idea would appear to have been; Some- 
where a variety, everywhere a variety ; with equal, as it seems to 
me with better reason, it may be held, Somewhere a species, every- 
where a species; that is to say, everywhere where individuals 
of such a regional species are found upholding the characters 
which distinguish their type in the region of its perfect em- 
ancipation. Let zutermediates be frankly accepted as inter- 
mediates, not feared or avoided as elements of confusion or as 
being necessarily prejudicial to this or that otherwise unexcep- 
tionable “ species”; let them be sought for and studied ; in time we | 
may come to learn from them, in one case perhaps, that they re- 
present weakening links between diverging forms; in another, 
that they afford indices to old relationships; in yet another, it 
may be that they mark the first steps in a union of two allied 
Species into a broader single one. 

Upon these. views the plant here defined is named with specific 
rank as Hypericum boreale (Britton). The conceptions of other 
botanists may require them to write Wpericum mutilum boréale, or, 
if even more conservative, Hypericum mutilum var. boreale. In 
any case, the fact of nature sought to be expressed is the same. 

‘Hypericum soreate (Britton), From 1-18 in. high, mostly 
3-8 in.; stem obscurely quadrangular to terete, upright from an 
assurgent, or reclined and rooting base, rarely erect from the root, 
simple or cymose-branched. Main stem-leaves from elliptic- 
linear to narrowly oblong and elliptic-oblong, or one or two 
pairs in the cyme sometimes slightly broadened to a sub-clasping 
base, those of the lower part of the stem much reduced and 
crowded, oval to short-oblong, at the very base of the stem some- 
times thickened and transformed into small lanceolate overlapping 
Scales. The small form of the plant which has been known as 
#1. Canadense var. minimum, has a simple stem, leafy-cymulose at 
the top, bearing many pairs of nearly linear or linear-oblong leaves, 
sometimes no larger than 3/’-4/x 1”. Another small form has 
fewer short-oblong obtusely rounded, small leaves, mostly nar- 
Towed to the base, but the pair which subtends the cymule some- 
times broadened and sub-clasping. Larger plants may be cymose- 
branched, even from the base, with narrowly oblong os 
' elliptic leaves reaching an extreme size of 16”x 3" or 4’. 


Cymes leafy-bracted throughout, varying from very simple = 
_ few-flowered to compound and_ contractedly — 
_ Bracts foliaceous throughout, oblong to linear, obtuse. Sepals 


214 


linear to oblong-linear, obtuse. Flowers about 2%" broad. Cap- 
sules 2-214” long by 1-114” broad, oblong, rounded or abruptly ~ 
contracted to the apiculate apex, membranaceous and strongly 
cross-wrinkled, the exposed portion deep purplish in color almost 
from the first, and conspicuously exceeding the sepals. Seeds rela- 
tively large, .03 inch long, linear, 3-5 times longer than broad, pale, 
strongly longitudinally furrowed and minutely cross-lined, under 
the microscope suggesting a miniature ear of corn.* 


As compared with doreale, mutilum, though having sometimes 
the assurgent base of the stem, is commonly erect from the root, 
the stem distinctly quadrangular, the stem-leaves, which reach an 
extreme size of 14’x'7’’, are throughout larger and much broader, 
and vary in shape from ovate-orbicular to ovate-oblong, often 
gradually narrowed from the broadly-dilated base to the obtuse 
tip, sometimes oblong, but always with a broad sub-clasping base; 
5-7-nerved, thin and much reticulated instead of thicker and often 
3-nerved, with the reticulation sparse, obscure or wanting; the 
cymes more compound and delicately dichotomous, with minute - 
and setaceous instead of leafy bracts; the capsule smaller, even 
only half the size(1’-2”” x 14-1’), mostly ovoid- or oblong-conical, 
often but little exceeding the sepals, commonly greenish or, in age, 
greenish-purple, instead of early deep purple; the seeds smaller, 
oblong, only 2-3 times longer than broad, yellowish-brown and 
shining instead of dull whitish, very minutely cross-wrinkled, not 
furrowed, but sometimes indistinctly striate. 

The seeds of doreate, while differing strikingly from those of 
mutilum, resemble somewhat those of Canadense, which, though 


* Besijes the York Harbor series, specimens of Hypericum boreale have been 
examined, as follows: 

Somesville, Mount Desert Island, Maine, Aug. 25, 1890, Edward L. Rand. 

Prince Edward’s Island, Aug. 2, 1888, John Macoun. 

St. John’:, Newfoundland, Aug. 6, 1894, B. L. Robinson and H. Schrenk. 

Notre Dame du Lac, Termiscouta Co., Canada, Aug. 13, 1887, John I. Northrop. 

« Vermont.” 

Lily Lake, Luzerne Co., Pa, Aug. 16, 1889, John K. Small. 

Morris Pond, New Jersey, Sept. 11, 1890, N. L, Britton. 

This last is the apparently intermediate speci.ren already referred to. It is the 
only example seen in which there are any minute bracts in the cymes. 

Hooker’s description of A. guinguenervium of Walter, in Fl. Bor. Am. 1: 
110, clearly refers to HH. doreale, as here defined, and as clearly excludes true 7. 
mutilum, of which guinguenervium is a synonym. The habitat of the plant is given 
as “Canada to Lake Winipeg. Dr. Richardson. Frequent in upper Canada.” 


215 


shorter, are of the same pale color, and are evidently striate— 
striate-angled, however, rather than furrowed. The seeds of majus 
prove to differ from those of Caxadense in respect of size, color and 
Surface character, much as those of mutilum differ from those of 
boreaie. The broad-based leaves of majus and mutilum here comes 
_ to mind and suggests a sort of parallelism between Canadense and 
majus on the one hand and éoreale and mutilum on the other. 

It may be noted that sayus has the smallest and narrowest 
seeds of the entire series; doreade, the smallest plant of all, having 
the largest seeds. 

The relationship between Canadense and doreale needs no ex- 
tented comment. It is true that from the leaves alone some 
forms of each plant would scarcely be regarded as different, but 
the character of the inflorescence of each is perfectly destinctive : 
Canadense will be found to differ constantly in its more strict and 
compound, minutely bracted cymes, with more or less tapering cap- 
sules and calyx lobes. 

It has just been said that forms of Canadense cannot by the 
leaves alone be separated from forms of doreale. This is equally 
true of majus. All three species assume in some of their states a 
narrowly. oblong form of leaf which is practically the same in all. 
In specimens of majus, however, which exhibit this form of feaf, 
the other distinctive characters remain unimpaired: it maintains 
the large pod, long tapering sepals, and characteristic seeds. Not 
So, however, Canadense. In the oblong-leaved form collected at 
York Harbor, there is a general difference also in the inflor- 
€scence and capsules; the cyme is less strict and more delicately 
branched, even becoming somewhat diffuse, and the short-pedi- 
celled capsules are more numerous and crowded, and only about 
half the usual size, 1-2” instead of 22" long; the seeds also 
take on a darker color, and seem to approach those of mutilum 
Indeed, some examples of this form have much similarity = 
less branched forms of mutilum, and actually seem to be impli- 
cated somehow with that species. In fact, in the case of some 
immature specimens, it can aaeoy be said to which species they 
belong. 

Compared with fully typical Canadense, this form reer . 
striking contrast, and in studying the group I have found it con- 
venient to label it var. parvicarpum. 


Ox: 


Further Remarks on Family Nomenclature. 


By Dr. V. HAvarRpD. 


It is generally admitted that the animal and vegetable king- 
doms cannot be classified on the same lines and divided into 
groups of the same value. Species and genera are nearly identi- 
cal in the two kingdoms, but as we rise above genera the similar- 
ity is less apparent and the higher we go the more doubtful be- 
comes the analogy between the groups, so that near the summit 
of the scale no comparison is possible. This is due to the great 
dissimilarity in the nature of the organisms, the infinitely more 
numerous and varied functions of animals and the greater diff- 
culty of ascertaining their natural relationship. From these con- 
siderations it follows that the application to both kingdoms of one 
uniform and identical nomenclature is impracticable; but, were it 
otherwise, the desirability of different and distinctive endings 
would still be manifest. 

-Zoologists appear to have reached a satisfactory uniformity in 
the terminations of the names of their principal groups. As 
early as 1845, the Association of American Naturalists, convened 
at New Haven, adopted the following proposition: 

. “ It is recommended that names of families should end uni- 
formly in zdae and sub-families in dae.” : 

These endings were then in general use; they were confirmed 
at the meeting of the A. A.A. S., in. 1877, and are now invaria- 
bly accepted... The name of the family or sub-family is a/ways that 
of one of its genera with the above endings. It is to be noticed, 
however, that animal families usually contain but few genera, not 
rarely only 2 or 3, often only 3 to 5,and seldom more than 10 or 
15. Perhaps they might be more correctly assimilated to botani- 
cal tribes. The term ¢rile appears to be very seldom used by 
zoologists, the name of the group intermediate between the fam- 
ily and genus being mostly swd-family in inae. — 

_ A cause of confusion, as pointed out by Mr. L. M. Underwood in 
the March BuLLeETIN, is the variable conception of order, being con- 
sidered by phanerogamic botanists a synonym of family, while, by 
zodlogists and the majority of cryptogamists, it is applied to the 


217 


next higher group above family. It seems to me that’a desirable 
step in the direction of uniformity would be taken if all botanists 
would, in this regard, conform to the usage of zodlogists; for, al- 
though we need not strive after similarity of nomenclature in the 
two kingdoms, yet we should avoid, so far as practicable, the use 
of the same appellations with a distinctly differeut meaning. 

The termination in aceae is the one now generally accepted by 
botanists for family names. It is strictly in accordance with good 
Latin and otherwise unobjectionable. The vexed question is, 
can it and must it be applied in all cases? A. de Candolle in his 
“Laws of Botanical Nomenclature” (1867) establishes three ex- 
ceptions: 

Ist. When the genus from which is derived the family name 
ends in zx or 2s (genitive icis, idis or iscis) the termination zeae, 
‘deze or ineae is admitted: SALICINEAE, BERBERIDEAE, TAMARIS- 
CINEAE. 

2d. When the genus from which the family name is derived has 
a name of unusual length, and there is no tribe founded upon it, 
the termination in eae is admitted: DipTEROCARPEAE from Dipter- 
ocarpus. 

3d. For a few large families, named long ago, and now every- 
where known under their irregular names, the latter are main- 
tained: CruciFERAE, LEGUMINOSAE, GUTTIFERAE, UMBELLIFERAE, 
Compositar, Lapiatar, CUPULIFERAE, CONIFERAE, PALMAE, GRAM- 
INEAE. 

These exceptions, when formulated, expressed the best con- 
Servative usage of the day, but it is apparent they have lost much 
of their strength and I doubt whether their illustrious author 
would now insist upon the first two. The first has long since 
been disregarded ; thus in Gray’s Manual, as well as in Pflanzen- 
familien and Baillon’s Histoire des Plantes, we have SALICACEAE, 
BERBERIDACEAE, PLANTAGINACEAE, ORCHIDACEAE, AMARYLLIDA- 
CEAE, etc? The length of the name is a point upon which we 
need not dwell; so long as a name is correctly constructed eed 
number of syllables cannot be of much moment; thus in Baillon’s. 
Histoire we have DiprEROCARPACEES and CENTROLEPIDACEES, and 
in Pflanzenfamilien, PoraMOGETONACEAE, HyDROCHARITACEAE, etc. 
It is therefore obvious that whenever the name of a family is de- 


218 


rived from that of a genus the termination in aceae is applicable 
without exception. 

The third exception presents serious difficulties. Here the 
family name is descriptive, that is, not based upon a genus but 
derived from some structural characteristic. Can the ending aceae 
be applied to it? This ending (from akos or acus) is an adjective- 
suffix, and therefore only applicable to a substantive, ¢. ¢.: rosa, 
vosaceus ; lilium, liliaceus, &c., the e being inserted for reasons of 
enphony. It cannot be affixed to adjectives (like compositus, conif- 
erus, leguminosus, gramineus, &c.), and such names as COMPOSITA- 
CEAE, CONIFERACAE, &c., with two adjective terminations, are 
obviously inadmissible. But it happens that all family descriptive 

“names (CoMPOSITAE excepted) are based upon a substantive (conus, 
umbella, crux, legumen, labium, cupula, gramen, &c.), 30 that we can 
replace the ordinary and variable adjective-ending by the uniform 
aceae and correctly form such family names as CONACEAE, UMBEL- 
LACEAE, LEGUMINACEAE, practically equivalent to CONIFERAE, Um-- 
BELLIFERAE, LEGUMINOSAE. The principle was recognized by 
Linnaeus who adopted such descriptive names as AMENTACEAE, 
PAPILIONACEAE, SPATHACEAE, &c. 

But admitting, as we may, that CoNnACEAE and LEGUMINACEAE 
are as correctly constructed as RosaceakE and LILIACEAE, it is still 
true that, although with the same ending, they convey an entirely 
different meaning; the former are plants wth cones, dearing le- 
gumes, while the latter are plants “ke the Rose, similar to the 
Lily; the former imply the possession of a certain character, the 
latter imply similitude to atype genus. Clearly this is not admis- 
sible. The uniformity we are striving for must be not only in 
outward form but likewise in the mode of construction and in- 
ward meaning. 

We are thus forced to the conclusion that the ending aceae is 
inapplicable to the names under consideration, and that no change 
in their construction is possible or advisable. What is then to be 
done? Two alternatives offer themselves: Leave them alone, as 
they have stood for several generations of botanists, or abolish 
and replace them by the first published generic names as proposed 
by Mr. Barnhart. The latter course will commend itself to all who 
think uniformity essential, and is urged upon us by the example 


a 


219 


of zodlogists, but such a radical change will not be easily ac- 
quiesced in. In the case of zodlogists, the family usually contain- 
ing only few genera, it is comparatively easy to adopt the name 
of one of them for that of the family, and, as a rule, the characters 
of the adopted genus will not as widely differ from those of the 
other genera as, say, Fragaria does from Prunus, Cassia from Tri- 
folium or Pinus from Ginkgo, so that such name is readily accept- 
able as representative of the family. It happens that the botanical 
families in question are among the largest of the vegetable world 
and it seems difficult to recognize CASSIACEAE as the representa- 
tive of all Lecuminosag, and AmMiaceae as that of all UmBe.ir- 
ERAE. The difficulty would be lessened if botanists, instead of 
extending the boundaries of families as has been the tendency in 
recent times, endeavored to restrict them to the lowest admissible 
limits, those, for instance, of several of our well defined suborders. 

I am not prepared to make a choice between the two alterna- 
tives. Perhaps a majority of botanists would now prefer that the 
old familiar names be maintained. Be that as it may, a decision 
Will soon impose itself and, judging from the present trend of 
thought, at least in this country, it is safe to predict that long and 
honorable usage will eventually be sacrificed to the claims of 
uniformity and stability. 

If we admit the law that the family name must in all cases be 
based upon that of a recognized genus, we still have difficulties to 
surmount; what is a “recognized” genus? The comparative 
instability of genera is well known; authors do not agree upon 
their value and limitations; should the family name be exposed 
to this insecurity? The danger of possible changes from that 
Source is very much mitigated by A. De Candolle’s law: 

“ An old genus name which has become the name of a section 
Or species can be maintained as the radical of a family name: 
LENTIBULARIACEAE from Lentidularia, HipPocASTANACEAE from 
Aesculus Hippocastanum, CARYOPHYLLACEAE from Dianthus Cary- 
ophyllus, etc.” 

If this law can be accepted by nomenclature reformers, their 
path will be made much smoother and we shall be the more readily 
inclined to follow them thereon. 


220 


New or noteworthy North American Phanerogams.—IX. 


By N. L. BRITTON. 


“Carex Baivevi Britton, n. sp. 

Carex tentaculata var. gracilis Boott, Il. 94. 1860. Not C. gra- 
cults Rom: 

Carex lurida var. gracilis Bailey, ee Torr. Club,1: 11. 1889. 

After observing this plant over a wide range of territory for 
several seasons, I have concluded to propose it as a species. I 
obtained crucial proof of its distinctness from C. lurida last 
autumn by finding the two growing side by side near Lake Placid, 
N.Y. I think the following description will readily identify it. 
It affords me much pleasure to associate the name of my friend, 
Prof. L. H. Bailey, with this very beautiful sedge. 

Glabrous, culms erect or reclining, very slender, minutely 
scabrous above, 1°-2° long. Leaves slightly scabrous, elongated, 
1’’-2”” wide, the upper and the similar bracts exceeding the culm ; 
staminate spike solitary, short-peduncled ; pistillate spikes 1-3, 
narrowly cylindric, very densely many -flowered, all erect or as- 
cending, 9’’-2’ long, about 4” in diameter, the upper sessile, the 
lower more or less stalked; perigynia inflated, ovoid, 214’’—3” 
long, ascending, abruptly contracted into the subulate sootica 
beak, prominently several-nerved, the lower about equalling, the 


upper longer than the linear-subulate ciliate-scabrous scale ; 
stigmas 3. 


In bogs, Vermont to Pennsylvania, Virginia and Tennessee 
along the mountains. 

Another species has recently been distributed as C. Badleyt 
from the Herbarium of Harvard University, but no description of 
it has appeared, and 1 am informed by Dr. B. L. Robinson that it 
is probably C. Raeana Boott, in which conclusion I concur. 

_ Carex Asa-Grayi hispidula (A. Gray) Bailey, Bull. Torr. Club, 
20: 427. 1803, 

Carex Grayi var. hispidula A. Gray; Bailey, Mem. Torr. Club, 

: 54. 1889. 

The occurrence of trichomes on the perygynia in Carex, while 
a very strong character in some groups, is evidently a poor one in 
others. In the species under consideration it is so variable that 
it does not seem to me to warrant the separation of the proposed 


221 
variety, especially as this does not appear to have other characters 
nor any well marked distribution.as compared with the glabrous 
plant. 


CAREX WALTERIANA Bailey, Bull. Torr. Club, 20: 429. 1893. 

This is another species whose perigynia vary from glabrous 
to quite densely pubescent. Professor Bailey has proposed (loc. 
cit.) a variety dvevis for those with glabrous perigynia. I find the 
pubescence exceedingly variable ; it is true,as he remarks, that the 
northern plants tend to have no pubescence, while the southern 
ones have a great deal, but I have New Jersey specimens in which 
the perigynia are hairy at the base and South Carolina specimens 
whose perigynia are very nearly glabrous. Under these circum- 
stances I see no desirability of separating the northern plant as a 
Variety. : 


CAREX BULLATA Schk. A form of this species with solitary 
long-stalked spreading’ pistillate spikes is collected by Mr. Light- 
hipe at Sand Hills, Middlesex Co., N. J. It was erroneously 
recorded in my Catalogue of Plants found in New Jersey as C. 
Olneyi, which, after an examination of several authentic specimens, 
I refer with confidence to C. monile. 


Carex Hartu Dewey Am. Journ. Sci. (IL.) qr: 226. 1866. 

Carex retrorsa var. Hartii A. Gray, Man. Ed. 5, 600. 1867. 

Examination of a considerable suite of specimens convinces me 
that this is a species distinct from C. retrorsa. 1 append a descrip- 
tion: 

Glabrous, culms very slender, smooth or very slightly scabrous 
above, erect or reclining, 114°-2%° long. _Leaves elongated, 
rough on the margins and lower side of the midvein, or. 3 ite, 
the upper and the similar bracts much overtopping the culm ; 
staminate spikes 1 or 2, the lower sometimes pistillate at the base, 
borne on astalk 1%4’-1’ long; pistillate spikes 2-4, scattered, rather 
loosely many-flowered, the upper sessile, the lower slender-stalked, 
1’-2’ long, about y’ thick, all erect or ascending; perigynia in- 
flated, ovoid-conic, spreading or the lower somewhat reflexed, prom- 
inently few-nerved, about 3/’ long, gradually tapering into the long 
2-toothed beak, 2-3 times as long as the lanceolate acute or acum- 
ate scale; stigmas 3. 

In marshes, Ontario to central New York (and Pennsylvania ?) 


_ West to Michigan. 


222 


CAREX ATRATIFORMIS Britton. 

Carex ovata Rudge, Trans. Linn. Soc. 7: 96. p/. 9. 1804. Not 
Burm. 

Carex atrata var. ovata Boott, Ill. 114. 1892. 

This plant appears to me constantly different from the Euro- 
pean C. atrata; I characterize it is as follows: 

Glabrous, culms very slender, erect, sharp-angled, scabrous 
above, 8’—2° tall, leafy only below. Leaves smooth or but 
slightly scabrous, 1/’—114"’ wide, rarely over 6’ long, much shorter 
than the culm; spikes 2-5, dense, oblong or oblong-cylindric, 4’/’— 
12” long, about 214” in diameter, the terminal one usually stami- 
nate at the base and sessile or nearly so, the others filiform- 
stalked and drooping when mature; lower bracts %4’-114’ long, 
very narrow, the upper ones subulate; perigynia flattened, ovate 
or nearly orbicular, puncticulate, ascending, about 1/’ long, tipped 
with a very short minutely 2-toothed beak; scales reddish-brown, 
oblong, obtuse or subacute, slightly narrower than and about 
equalling the perigynia; stigmas 3. 

Newfoundland to the mountains of New England, west to the 
Northwest Territory. . : 

~ CAREX STRICTA XEROCARPA (S. H. Wright). 

Carex xerocarpa S. H. Wright, Am. Journ. Sci. (II.) 42: 334- 
1866. 

This appears to me to be a very well marked variety, if not a 
species distinct from C. stricta Lam. The pistillate spikes are 
almost filiform, loosely flowered and about 1” in diameter. Be- 
sides original specimens collected in central New York by the 
describer of the species, I have it from Illinois, collected by J. 
Wolf. : 

Carex Haypen Dewey, Am. Journ. Sci. (II.) 18: 103. 1854. 
Carex aperta Carey in A. Gray, Man. 547. 1848. Not Boott. 
Carex stricta var. decora Bailey, Coult. Bot. Gaz. 13: 85. 1888. 
Professor Bailey has shown that the plant referred to C. aperta 

Boott, in the earlier editions of Gray’s Manual, is not the same as 

the species of the Northwest to which the name was originally 

applied. I am confident that it is specifically distinct from C. 

stricta. 1 refer it to C. Haydeni with hesitation, although exam- 

ination of an immature authentic specimen indicates that this name 


may be correctly applied to it. The species may be characterized 
as follows: 


223 


Glabrous, similar to C. s¢ricta but smaller, culm slender, scab- 
rous above, seldom over 2° high. Leaves 1/114” wide, rough- 
margined, shorter than or sometimes overtopping the culm, their 
sheaths slightly or not at all fibrillose; lower bract foliaceous, about 
equalling the culm; staminate spikes linear-cylindric, 6-15” long, 
about 2’ in diameter, erect or somewhat spreading, all sessile 
or nearly so, sometimes with a few staminate flowers at the sum- 
mit; perigynia orbicular, obtuse, about 4" broad, faintly 2-4- 
~nerved, minutely beaked, the orifice entire; scales lanceolate, 
purplish, spreading, very acute, about twice as long as the peri- 
gynia; stigmas 2. 

In swamps, New Brunswick to Western Ontario, south to 
Rhode Island, Pennsylvania, Wisconsin and Nebraska. 


CAREX CosTaTA Schwein. Ann. Lyc.N. Y. 1: 67. 1824. Not Presl, 

1819. : 

Carex virescens var. costata Dewey, Am. Journ. Sci. 9: 260. 
1825. 

My observations on this plant lead me to believe it constantly 
different from C. virescens Muh]. As the name applied to it by 

«, U 
Schweinitz has been used before, I propose for it"CAREX COSTEL- 
LATA and give its characters as follows: 

Similar to C. virescens but taller and more spreading, culms 
slender, 1°_2 1%Z° long. Leaves 114’’-2” wide, pubescent, espe- 
cially on the sheaths, shorter than the culm, the upper one and the 
similar lower bract sometimes overtopping the spikes ; spikes 2=5; 
narrowly cylindric, many-flowered, rather loose, ¥4’-1/% long, 
1%” in diameter, erect or slightly spreading, the terminal one 
Staminate below, the lower one commonly filiform-stalked; peri- 
gynia oblong, densely pubescent, narrowed at each end, strongly 
Several-ribbed, 1’ long, rather more than yn" thick, beakless, the 
Orifice entire; scales ovate, scarious-margined, acuminate or 
Cuspidate, shorter than the perigynia; stigmas 3. 

In woods, Maine and Ontario to North Carolina, chiefly along 


the mountains. 

“ ALLIONIA BusuHI n. sp. Low, glabrous, somewhat fleshy; stem 
Nearly white, diffusely branched, about 8’ high, the branches 
slender, widely divergent; leaves narrowly linear, sessile, 1-3’ long, 
1’’_114” wide, blunt, their width almost uniform from base to 
pex; involucres clustered at the ends of the branches, at first 
campanulate and longer than the flowers, at length rotate ane be- 
_€oming 10” broad, membranaceous, pubescent, finely reticulate- 
_ Veined, their 3 short lobes semi-circular, rounded, the mid-veins 
_ prominent. 


224 


In dry ground. Jackson county, Missouri. August, 1893. 
B. F. Bush. ee 

Resembles 4. Bodin (Holzinger) Morong, which is much 
- more slender, its involucres short-pedicelled in the axils, the lobes 
ovate-oblong and acute. : 


sae 
RANUNCULUS ALLEGHENIENSIS n. sp. 


Similar in aspect to R. adortivus and R. micranthus, glabrous, 
stem widely branched, 1°-2° tall. Radical leaves reniform or sub- 
orbicular, 6-2’ wide, long-petioled, crenate or some of. them 
lobed, the teeth and lobes subacute; stem leaves sessile or the 
lower petioled, divided nearly or quite to the base into linear acute 
entire toothed or cleft segments; flowers about 2” broad; petals 
oblong, glandular, not exceeding the calyx; head of fruit sub-glo- 
bose or oblong, 2” in diameter; receptacle linear, about 2’ long, 
pubescent ; achenes slightly compressed and margined, tipped with 
subulate hooked or recurved styles of about one-half their length. 


Mountains of Virginia and North Carolina. April-May. 


Brassica JUNCEA (L.) Cosson, Bull. Soc. Bot. France, 6: 609- 
4859... 

Sinapis guncea L. Sp. Pl. 668. 1753. 

Annual, glabrous, somewhat succulent, stem erect, usually 
stout, 2°-4° tall. Lower leaves runcinate-pinnatifid and dentate, 
long-petioled, 4’-6/ long, the uppermost sessile or nearly so, lance- 
olate or linear, commonly entire, much smaller; fruiting racemes 
sometimes 1° long; pods erect or nearly so on slender ascending 
pedicels, not appressed to the axis, 1-2’ long, rather more than 1’ 


wide, the conic-subulate beak one-fourth to one-third the length 
of the body. 


_ This plant has been sent to me from a aumber of points dur- 
ing the past three years and I have collected it twice myself. It 
seems to be fairly naturalized in some regions. As represented in 
the specimens examined, it is readily distinguished from &. Siza- 
pistrum Boiss. by the total absence of the hispid pubescence of that 
species and by its erect longer and subulate-beaked pods. 

In waste places, Southern New York and Pennsylvania to 
Michigan and Virginia. Adventive or naturalized from Asia. 
Also introduced into the West Indies and South America. 


_ Cuimapnica Pursh. Additional proof of the non-publication 
of Pseva Raf., as an older name for this genus is to be found in 
Rafinesque’s review of “ A Manual of Botany for the Northern 


225 


States, compiled by the Editor of Richard’s Botanical Dictionary,” 
Albany, 1817. The author of this work is reported to have been 
Prof. Amos Eaton. Rafinesque’s review is printed in the “ Ameri- 


«an Monthly Magazine,” 1: 426-430, September, 1817, where 


among his criticisms he remarks, « He (Eaton) has not adopted the 
good genera Pte Si a . ete, 


Descriptions of new Leaves from the Cretaceous (Dakota Group) 
of Kansas.* 


By ArtTHuUR HOLLICK. 
(PLATES 236, 237.) 


During the past year one of the students f at Columbia Col- 
lege was engaged under my direction in overhauling and nam- 
ing the Dakota Group material in the Geological Museum, with 
instructions to put aside all specimens which could not be satis- 
factorily identified. I take pleasure in saying that the specimens 
now under consideration were the only ones, except a few frag- 
ments not capable of being satisfactory determined, which he 
found necessary to thus separate; also to state that they appar- 
ently represent three species and one variety new to the horizon, - 
and to give him credit for having recognized them as posessing 
characters different from those of any published plates or descrip- 
tions with which they could be compared. All are from the 
vicinity of Fort Harker, Kans. 


SASSAFRAS (ARALIOPSIS) Lesq. 


This subdivision of the genus Sassafras was made by Lesque- 
reux to contain a number of leaves which might be classed with 
either Sassafras or Aralia. Their systematic position is yet prob- 
lematic, but they are included under the former genus in his post- 
humous Flora of the Dakota Group, edited ae F, H. Knowlton. 
(Monog. xvii. U.S. G.S., 1891.) 


* Read by title at the meeting of the New York Academy of Sciences, February 


u, 1895. j 
" +Mr. Chas. R. Pollard, now Assistant Curator of the National Herbarium. 


296 


SASSAFRAS (ARALIOPSIS) DISSECTUM SYMMETRICUM N. var, 


(Plate 236.) 


Differs from S. (A.) dissectum Lesq. Cret. & Tert. Fl. 57; FI. 
Dak. Gr. 101. p/. rg. f. 7, in its symmetrical branching, especially 
that of the lateral primaries, which start from the base of the leaf 
exactly opposite to one another and fork at an equal distance 
above ; also in the fact that the blade of the leaf is not decurrent 
along the petiole, but ends at the point where the lateral primaries 
branch from the midrib. 

I was at first inclined to describe this as a new species, but the 
imperfect condition of the upper portion of the specimen seemed 
to render this inadvisable, and its substantial agreement in essen- 
tial particulars with S. (A.) dissectum decided me to class it asa 
variety of that species. 


Cissires HEEr. 


This genus was founded by Heer, to include leaves presumably 
allied to Czsszs, but subsequently made by other authors to in- 


clude leaves having more or less resemblance to Vitis, Platanus, 
Sassafras, etc. 


CISSITES PLATANOIDEA Nn. sp. 
(Plate 237. f. 2.) 


Leaf symmetrical, 23¢ in. long by 23 in. broad, sub-orbicular 
to fan-shaped in outline, abruptly decurrent at base, obscurely 
3-lobed; margin undulate or obscurely dentate; nervation 3-pal- 
mate, craspedodrome; midrib abruptly thickened below the point 
where the lateral primaries branch off, also to a lesser extent be- 
low the point where the upper secondaries branch off; secondaries 
clustered together in two pairs above the middle of the midrib, 
the upper pair extending to the margins, the lower pair merging 
gradually into the tertiary nervation, of which it may perhaps be 
considered to form a part; the latter forming polygonal meshes, 
well defined; lateral primaries branched mostly from below, ob- 
scurely from above near the extremities and abruptly thickened 
below the point where the first secondaries branch off. 


This leaf is suggestive of species which have been described 
under the genera Sassafras (S. obtusum Lesq., etc.,) Platanus (P. 
Heeri Lesq., P. obtusiloba Lesq., etc.), Cissites (C. ingens Lesq., etc.), 


227 


and Parrotia (P. Canfieldi Lesq.), but is clearly distinct from any of 
these, and rather than erect a new genus I have decided to class 
it with C7sszfes and to indicate other characteristics in the specific 
name. 


CISSITES ACUTILOBA Nn. sp. 
(Plate 237. f. 3.) 


Leaf 234 in. long by 234 in. broad, sub-orbicular in outline, 
three lobed, lobes acute; nervation 3-palmate ; lateral primaries 
long, almost equalling the midrib, somewhat incurved, margin en- 
tire. 

Differs from C. Harkerianus Lesq., with which it is closely 
allied, in its more rounded outline, longer lateral primaries and 


acute lobes and apex. 


PROTOPHYLLUM Lesq. 


This genus was founded by Lesquereux to include certain 
leaves of an apparently synthetic type, some of which had been 
described under the genera Credneria and Pterospermites. The 
systematic position of the genus cannot yet be said to be defini- 
tely determined, although in Fl. Dak. Gr. /. c. it is classed in the 
Sterculiaceae. Whatever its systematic position may be there is 
no doubt that our species belongs to the genus as defined. — 


PROTOPHYLLUM QUERCIFORME nN. Sp. 


- 


(Plate 237, f. 1.) 


Leaf 3% in. long by almost 3% broad at middle, rhombic- 
vate in outline, rounded above, more or less abruptly narrowed 
from middle to base, slightly acuminate at apex; margin undulate- 
dentate; nervation craspedodrome; lower secondaries relatively 
Slender, crowded together, branching from the midrib at an obtuse 
angle; median ones stronger, more distant, branching from the 
midrib at a more acute angle, forked two-three times; upper ones 
again slender and branching as before at a more obtuse angle; 
tertiary nervation uniform, slightly curved outward, simple, fine, 


_ and at right angles to the secondaries throughout. 

This leaf somewhat resembles P. Haydenii Lesq., but differs 
in its smaller size, narrowed base and. more rounded apex. The 
Superficial appearance suggests one of our broad leaved oaks, as 
I have indicated in the specific name. 


228 


In conclusion I wish to acknowledge my indebtedness to Dr. 
F. H. Knowlton of the United States Geological Survey for criti- 
cisms and references in nomenclature and synonomy. 


A Preliminary List of the North American Species of Malpighiaceae 
. and Zygophyllaceae. 


By ANNA Murray VAIL. 


MALPIGHIACEAE. 
1. JANUSIA A. Juss. Monog. Malp. 349. p/. 27. 1843. 

JANuSIA GRACILIS A. Gray, Pl. Wright. 1: 37. 1852. 

Distr. Arizona, New Mexico, Texas and Mexico. 

2. ASPICARPA Rich. in Mem. Mus. Par. 2: 398. fl. 7. 1815. 
1. ASPICARPA HYSSOPIFOLIA A. Gray, Bost. Journ. Nat. Hist. 6: 

167. 1850. 

Distr. Téxas, New Mexico and North Mexico. 

2. ASPICARPA LONGIPES A. Gray, Pl. Wright. 1: 37. 1852. 

Distr. Texas, New Mexico, southern Arizona and North 
Mexico, 

Very close to A. humilis (Benth.), from which it differs mainly 
in its trailing, and decumbent habit and somewhat larger leaves.* 
3. THRYALLIS L. Sp. Pl. Ed. 2, 554. 1763..Not Thryallis 
) Mart. Nov. Gen. 3: 77. pd. 230, 2317. 1829. 

THRYALLIS ANGUSTIFOLIA (Benth.) Kuntze, Rev. Gen. Pl. 89. 1891. 
 Galphimia angustifolia Benth. Bot. Sulph. 9. 1844. 

_ Galphimia linifolia A. Gray, Bost. Journ. Nat. Hist. 6: 166. 
1850. 

Distr. Western Texas to New Mexico, Lower California and 
Mexico. 

_ Very variable. The broader leaved form is 
THRYALLIS ANGUSTIFOLIA OBLONGIFOLIA, (A. Gray.) 

Galphimia linifolia 3 oblongifolia A. Gray, Pl. Wright. 1: 36 

1852. It is found with the type and may possibly be a species. 


* Aspicarpa Hartwegiana A, Juss. Arch. Mus, Par. 3: 598. 1843, is a synonym 
of Gaudichaudia humilis Benth. Pl. Hartw. 6. 1839, fde Kew Index. 


229 . 


4. MALPIGIA -L. Sp. -Pla25.: 1783. 


MALPiGIA GLABRA L. Sp. Pl. 425. 1753. 
Malpigia nitida Mill. Gardn. Dict. Ed. 8: No. 5. 1768. 
Distr. Texas, Mexico, West Indies and South America. 

5. BYRSONIMA Rich. & Juss. Ann. Mus. Par, 18: 481. 1811. 
Byrsonima Lucipa (Sw.) H. B. K. Nov. Gen. 5: 147. 1821. 
Malphigia lucida Sw. F\.. Ind. Occ. 852. 1800. 

Distr. South Florida and West Indies. 


ZYGOPHYLLACEAE. 
1. FAGONIA L. Sp. Pl. 386. 1753. 


* Faconta CAuiFornica Benth. Bot. Sulph. to. 1844. 
Distr. California, North and South Mexico. 


2. GUAJACUM L. Sp. Pl. 381. 1753. 


Guajacum Sanctum L. Sp. Pl. 382.. 1753. 

Guajacum verticale Ortega, Dec. 8: 93. 1800. 

Guajacum Sanctum var. parvifolium Nutt. Sylva, 3: 17. eas 
Distr. South Florida, Bahamas, San Domingo, Porto Rico, etc. 


3. PORLIERIA Ruiz & Pav. Prod. 55. p/. 9: 1794. 


PoRLIERIA ANGUSTIFOLIA (Engelm.) A. Gray, Pl. Wright. 1: 28. 
1852. . 

Guajacum angustifolium Engelm. Wisliz. Rep. 29. 1848. 

Distr. Texas and North Mexico. 


4. COVILLEA.+ 


Larrea Cav. in Ann. Hist. Nat. 2: 119. pf. 18, 19. 1800. Not 
Larrea Orteg. Hort. Matr. Dec. 15. pl. 2. 1797. 


COVILLEA DIVARICATA (Cav.). 
Larrea divarica‘a Cav. Ann. Hist. Nat. 2: 122. 1800. 


* FAGONIA CALIFORNICA GLUTINOSA Nf. var. 

Somewhat stouter, the leaflets larger; the terminal one sub-rhomboid, the whole 
plant beset with sub-sessile, aromatic, gold-colored glands. 

Sonora, Mexico, Pringle (1884); Los Angeles Bay, Lower California, Palmer 
No, 546 (1887) ; Sta. Rosalia Island, Palmer, Lower California (1890). 

The above name appears on 4 sheet of Mr. Pringle’s collection in Herb, Columbia 
College, and I have not been able to find any printed reference to it. 

+ Named in honor of Frederick Vernon Coville. 


230 


& 


Zygophyllum tridentatum DC, Prodr. 1: 706, 1824. 

Larrea Mexicana Moric. Pl. Nouv. Am. 71. £48. 1833-1846. 

Larrea glutinosa Engelm. Wisliz. Rep. 9. 1848. 

Distr. Southern Utah, Nevada, Arizona, Texas, New Mexico, 
Mexico and in the deserts of Chili. 

Careful examination does not reveal any characters that war- 
rant keeping the Mexican species distinct from the South American. 
The principal differences were found in the stamineal scale. The 
latter is a very variable organ, and three.or four variations were 
found, not only on the same plant, but in the same flower.* 


5. JRIBULUS-L, Sp. Pi.:386..°-1753, 

I. TRIBULUS CISTOIDES L. Sp. Pl. 387. 1753. 
Distr. Florida, Texas, Mexico, Lower California, West 
Indies and tropical America. 

2. TRIBULUS TERRESTRIS L. Sp. Pl. 387. 1753. 
Distr. Ballast Grounds, Hunter’s Point, Long Island, N. Y.; 
South Carolina, Mexico and Brazil. A native of Southern 
Europe, East Indies, etc. 


6. KALLSTROEMIA Scop. Introd. 212. 1777. 
I. KALLSTROEMIA CaLirornica (S. Wats.) 
Tribulus Californicus S. Wats. Proc. Amer. Acad. 11: 125. 1876. 
‘Distr. Arizona, North Mexico and Lower California. 
2. KALLSTROEMIA GRANDIFLORA Torrey; A. Gray, Pl. Wright. 1: 28. 
1852. 


Kalistroemia grandiflora var. detonsa A. Gray, PI. Wright. 1: 
28. 1852. 


Tribulus grandifiorus Benth. & Hook. Gen. Pl. 1: 264. 
1862-67. 


Distr. Texas, Arizona, New Mexico, California, North and 
South Mexico, Guatemala. 
3. KALLSTROEMIA Maxima (L.) T. & G. Fl. N. A. x: 213. 1838. 
Tribulus maximus L. Sp. Pl. 386. 1753. 
Tribulus trijugatus Nutt. Gen. 1: 297, 1318, 


* The other species are Covillea nitida (Cav.) and Covillea. cuneifolia (Cav-)» 
both from South America,and described under Larrea in Cav. Icon. 6: 40-41. p% 
559, 500. 1801. 


231 


Tribulus decolor Macfadyen, FI. Jamaic. 186. 1837. 
Distr. Kansas, Arizona, Texas, New Mexico, Florida, Mexico, 
and common in tropical and sub-tropical America to Brazil. 


7..PEGANUM L. Sp. Pl. 444. 1753. 


Prcanum Mexicanum A. Gray, Pl. Wright. 1: 30. 1852. 2: 
106. 1853. 
Distr. New Mexico and Mexico. 


The Genus Zenobia Don, 


In 1834 David Don published « A New Arrangement of the 
Ericaceae,”* in which he separated a number of species from the 
large Linnaean genus Andromeda, creating at the same time sev- 
eral new genera in which the detached species were included. Of 
these additions, Cassandra (now Chamaedaphne), Casstope and 
Leucothoe have long been recognized as distinct; but Zenodia, 
although it was taken up by Bentham and Hooker,} has been 
considered by Gray and later botanists only a subgenus of Andro- 
meda. 1 here propose to restore it to its original rank, thus pre- 
serving the arrangement of Don and of Hooker. 


ZENOBIA Don. 


Calyx free, 5-lobed; corolla campanulate with 5 rounded 
lobes; stamens 10, on short basally dilated filaments; anther- 
cells elongated, 2-awned; stigma simple; capsule depressed-glo- 
bose, 5-angled, loculicidally 5-valved, many-seeded ; seeds oval, 
with a spongy testa and fleshy albumen. Smooth or glaucous 
shrubs with somewhat coriaceous strongly reticulated leaves, and 
white flowers disposed in axillary fascicles. 


ZENOBIA CASSINEFOLIA (Vent.). 
Andromeda cassinefolia Vent. Hort. Cels. 1: 00. 1800. 
Andromeda nitida Sims, Bot. Mag. fl. 970. 1803. Not Bartr. 
Andromeda speciosa Michx. Fl. Bor. Am. 1: 256. 1803. 


Fruticose, glabrous throughout; leaves coriaceous, oblong- 
Ovate, acuminate, irregularly serrate, the earlier obtuse, the later 
acute, length 5 cm., width 2.5-to 3 cm.: flowers in naked umbelli- 
————— 


* Edinb, N. Phil. Journ. 17: 158, 1834. 
+ Genera Plantarum, 2: 587. 1873. 


232 


form fascicles; corolla not deeply 5-lobed, 6 to 8 mm. long; calyx 
spreading, its lobes acutely triangular. 


North Carolina to Florida. 


ZENOBIA PULVERULENTA (Willd). | 
Andromeda pulverulenta Willd. Sp. Pl. 2: 610. 1799. Bartr. 


Trav. N. & S. Car. Georg. fl. 474. 1799, without synonymy or 
description. . ; 

Andromeda speciosa var. pulverulenta Michx. Fl. Bor. Am. 1: 
256. 1803. 

Fruticose, glabrous and prominently glaucous throughout; 
leaves subcoriaceous, elliptical, entire or obscurely serrulate, 
slightly cuspidate, acute when young, becoming obtuse; length 
5.5 to 6 cm., width 2.5 to 3 cm.; calyx somewhat campanulate, 
its lobes closely appressed to the corolla, and with the peduncle 
glauco-pulverulent; corolla with lobes more acute than in cassine- 


Solia.. 
Same range as the last, but less common. 
This species was figured by Bartram,* who evidently considered 
it worthy of something more than varietal rank, although he did 
not describe it. Michaux placed it under his Andromeda speciosa, 
and his example was followed by Pursh,+ who remarks in a note 
on speciosa: “1 certainly must epincide with Michaux’s idea of A. 
_ pulverulenta being only a variety, as I very frequently have seen 
intermediate varieties and even had the trace of bothon one plant.” 
While it is true that Z. pulverulenta exhibits a variable degree 
of glaucosity, there are other characters which, on careful examin- 
tion, serve to separate it from cassinefolia. The calyx in the one 
case is campanulate, inthe other spreading, with smaller lobes, the 
leaves are elliptical, and not oblong-ovate; and the peduncles in 
pulverulenta are always prominently glauco-pulverulent, giving the 
plant a very distinctive appearance. It is also possible that the 
two species may hybridize, as they occur throughout the same 


range, and this would explain the various forms which Pursh 
claims to have seen on the same plant. 


CuHaries Louis PoLLarD. 


* Trav. N. & S. Car. and Georg, 1. c. 
+ FL Am. Sept. 2: 294. 1814. 


233 


Botanical Notes. 


Nomenclatural—_A_ document has recently been issued from the 
herbarium of Harvard University, accompanied by a letter signed 
by the curator of that establishment, objecting to the system of 
nomenclature adopted almost unanimously by the botanists of the. 
American Association for the Advancement of Science at meetings 
held in Rochester, N. Y., 1892; Madison, Wis., 1893, and Brook- 
lyn, N. Y., 1894, and recommending certain rules as substitutes. 
It seems desirable that this document be brought-before our botan- 
ists for their kindly consideration. 

The principles, or it may be better expressed, lack of princi- 
ples, which are here favored have, as a matter of fact, been most 
seriously considered by the American Association botanists for 
more than five years, and every one of them has been carefully 
weighed and found wanting in its application to a stable system 
of nomenclature. The proposed rules have, indeed, been framed 
to support what has recently been fittingly termed the “discredited” 
plan of nomenclature. Still it may be assumed that they will 
receive a certain amount of support from plant collectors and 
physiologists, at least until more books shall have been issued 
based on the rules which have been proven by long trial in other 
branches of biological science to give nomenclature the stability 
which is so necessary. The way for these has been paved by that 
magnificent work, the “Silva of North America,’ the most ex- 
haustive and elegant botanical publication yet produced in our 
country ; by several State and local floras; and by other papers 
too numerous to mention. The want of easily accessible descrip- 
tive floras, written on this very practical system approved by the. 
botanists of the American Association, is the only circumstance 
which retards its still wider adoption. There is every reason to 
believe that these will soon be supplied. 


254 


Proceedings of the Club. 


Turespay Eveninc, APRIL 9TH, 1894. 


Vice President Lighthope presided and there were 27 persons 
present. 

The following persons were elected active members: Miss 
Alice M. Isaacs, Prof. E. S. Burgess, Miss Francis M. Chapin, Mr. 
John Dallas, Mr. Carl D. Schaeffer, Miss Adelaide Porter, Miss S. 
LeB. Drumm, Mr. Wm. C. Alpers, Miss Helen Parish, Miss Mabel 
Choate and Mrs. Hamilton Kean. 

Mr. Small, for the committee appointed to draft resolutions 
upon the death of Mr. Redfield, presented a report and submitted 
resolutions. The report was accepted and the resolutions adopted. 
[The resolutions were printed with Mr. Canby’s biographical 
sketch of Mr. Redfield in the April BuLLETIN. | 

The announced paper of the evening, entitled “ Vallsneria 
Spirals,” was then read by Miss Effie A. Southworth. The paper 
consisted of an exhaustive review of the literature of this subject, 
some contradictory statements by different authors being con- 
trasted, followed by an account of the work which had been done 
by Miss Southworth looking toward an explanation of such dis- 


crepancies, and the contribution of additional facts to our knowl- 
edge of this plant. 


WepbnesDAy Eveninc, Aprit 24TH, 1895. 


Vice-President Allen in the chair and 29 persons present. 

Mr. John G. Block and Mr. John H. Stottler were elected 
active members. 

Dr. Britton exhibited specimens of last year’s growth of Conopho- 
lis Americana, found by Mr. Stottler on Staten Island, where it was 
apparently parasitical on an oak. He stated that several hundred 
plants were growing in this locality, comprising a patch about 10 
feet square. Mr. Lighthipe reported two collections of this plant 
in the vicinity of Woodbridge, N. J., and was able to state that in 
one of the places there were only a few isolated plants. Dr. 
Rusby had collected the plant a number of times, but had always 


235 


found but a few plants growing in one locality. He had also 
noticed this peculiarity in the case cf C. Mexicana. Mr. Van 
Sickle had found a patch of about 50 specimens growing upon the 
top of the Palisades. 

Dr. Britton called attention the 6th volume of Hough’s Wood 
Sections, which was now ready for distribution and to the publica- 
tion of Mr. Small’s monograph on Polygonum and exhibited copies 
of both. 

The announced paper of the evening was then read by Mr. H. 
A. Siebrecht, entitled «Some Interesting Orchids of Cultivation.” 
The paper was handsomely illustrated with living specimens in 
bloom. The paper dealt with the history of the earliest collec- 
tions and collectors of orchids, and-traced the development of 
their cultivation up to the present time. Interesting facts concern- 
ing their native habits and the best methods of growing the dif- 
ferent classes were presented, and a discussion concerning their 
special habits of growth and fertilization was introduced. 


Index to recent Literature relating to American Botany. 


Atkinson, G.F. Additional Note on the Fungi of Blowing Rock, N. C. 
Journ. Elisha Mitchell Sci. Soc. 10: 78. 1894. 

Atkinson, G. F. Additions to the Erysipheae of Alabama. Journ. 
Elisha Mitchell Sci. Soc. 10: 74-76. 1894. 

Atkinson, G. F. Comparative Study of the Structure and Function of 
the Sporangia of Ferns in the Dispersion of Spores. Proc. A. A. A. 
S. 42: 253, 254. 1894. 

Atkinson, G. F. Photography asan Instrument for Recording the ma- 
croscopic Characters of Microdrganisms in artificial Cultures. Proc. 
A.A. A.S. 42: 255. 1894. 

Atkinson, G. F. Preliminary Note on the Relation between the sterile 
and fertile Leaves of Onoclea. Bot. Gaz. 19: 374, 375: 1894. 

Atkinson, G. F. Preliminary Note on the Swarm-spores of Pythium 
and Ceratiomyxa. Bot: Gaz. 19: 375-378. 1894. 

Atkinson, G. F. Some Septoriae from Alabama. Journ. Elisha Mitchell 
Sci. Soc. 10: 76-78. 1894. 

Atkinson, G. F. Symbiosis in the Roots of the Ophioglosseae. Proc. 

. A. A.A. S. 42: 254,255. 1894. 


236. 


Atkinson, G, F, The Study of the Biology of Ferns by the Collodian 
Method. 8vo. pp. 134. f. 763. London and N. Y., McMillan & Co. 
1894. 

Bailey, L. H. The Recent Apple Failures. Bull. N. Y. (Cornell) 
' Exp. Sta. 84: pp. 34-f. 9. pi I. Ja. 1895. 

Treats largely of apple scab, with colored plate. 

Bailey, W. W. George Hunt. Bot. Gaz. 20: 176, 177. 20 Ap. 
1895. 

Bastin, E. S. Structure of Zfigaea repens. Am. Journ. Pharm. 67: 
231-2306. f. I-37. My. 1895. 

Bolley, H. L. Distribution of Weed Seeds by Winter Winds. Bull. 
N. Dak. Exp. Sta. 17: 102-105. Mr. 1895. 


Bolley, H. L. Effect of Seed Exchange upon Culture of Wheat. 
_ Bull. N. Dak. Exp. Sta. 172 85-102. Mr. 1895. 


Blodgett, F. H. On the Development of the Bulb of the Adder’s- 
- tongue. Bot. Gaz. 20: 172-175. fl. 7g. 20 Ap. 1895. 


Boyer, C.S. A fossil marine Diatomaceous Deposit at St. Augustine, 
Florida. Bull. Torr. Bot. Club, 22: 171-174. 18 Ap. 1895. 


Britton, N. L. The Systematic Botany of North America. Bot. Gaz. 
20: 177-179. 20 Ap. 1895. 

. Announcement of assignments of groups to monographers. 

Canby, W.M. John H. Redfield. Bull. Torr. Bot. Club, 22: 162- 
171. 18 Ap. 1895. ‘ 
Sketch of life, with bibliography and portrait. 


Chodat, R. Polygalaceae novae vel parum cognitae. Bull. Herb. 
Boiss. 3: 121-135. Mr. 1895. 
Five new species of Polyga/a and fourteen of Monnina from Central oe South 
America are described. 
Coville, F. V. A Reply to Dr. Robinson’s Criticism of the ‘‘ List of 


Pteridophyta and Spermatophyta of Northeastern America.” Bot. 
Gaz. 20: 162-167. 20 Ap. 1895. 


Coville, F. V. Report of the Botanist U. S. Department of Agricul- 
ture for 1893. Rep. Sec. Agric. 1893: 235-244. 1894. 

Deane, W. Notes from my Herbarium. II. Bot. Gaz. 20: 150-154- 
20 Ap. 1895. 

Eastwood, A. Two Species of Aquilegia from the anes Sonoran 
Zone of Colorado and Utah. Proc. Cal. Acad. Sci. II. 4: 559-552. 
pl. 18, 79. 19 Mr. 1895. 


Aguilegia micrantha n. sp. and A, ecalcarata Eastwood are illustrated and de- 
scribed. 


237 


Forbes, S. A. Experiments with the Muscardine Disease of the Chinch 
Bug, etc. Bull. Ill. Exp. Sta. 38: 25-86. p/. 8. Mr. 1895. 
Sporotrichum was cultivated upon chinch bugs, and also equally as successfully 

upon a mixture of cornmeal and beef broth. 

Gardner, F. D. Corn Experiments. Bull. Ill. Exp. Sta. 37: pp. 24. 
F. 1895. 

Cross-fertilization—Effect of removing tassals. 

Greene, E. L. Novitates occidentales.—XII. Erythea. 3: 62-68. 

t Ap, 1895. 


New species in /loerkea, Calyptridium, Rumex, Spiraea, Eryngium, Sanicula, 
Erigeron, Cladothamnus, Phacelia, Cryptanthe, Mimulus, Fritillaria. 


Greene, E. L. Phytographic Notes and Amendments.—I. Erythea, 
3: 53-57. 1 Ap. 1895. ; 
Hennings, P. Fungi Goyazenses.. Hedwigia, 34: 88-112. 8 Ap. 


1895. 
Describes Fungi collected by Dr. E. Ule in Brazil. New species in Ustilago, 


Uromyces, Puccinia, Cronartium,Ravenelia, Uredo, Aecidium, Asterina, Asterula, 

Asteroma, Dimerosporium, Parodiella, Meliola, Hypocrella, Uleomyces (n. g.), 

Ophioceras, Ti abutia, Xylaria, Phyllachora, Dothidella and Tryblidium. 

Keffer, C. A. The Banksian Pine in the Nebraska Sand Hills. Gard. 
& For. 8: 152. 1 My. 1895. 

Kellerman, W. A. and Werner, W. C. Catalogue of Ohio Plants. 
Geology of Ohio, 7: 56-406. 1895. 


A list of plants of the state, citing distribution and known localities for the rarer 

Species, prefaced by a bibliography. 

Kindberg, N. C. Check-list of European and North American Momes: 

~ Can. Rec. Sci. 6: 17-23; 72-76. 1894. [Also reprinted.] 

Knobel, E. A Guide to find the Names of all wild-growing Trees 
and Shrubs of New England by their Leaves. Pamph. pp. 41. 1894. 

Knobel, E. Ferns and Evergreens of New England. Pamph. //. zz. 
1895. 

Koehne, E. [Critical notes on certain shrubs and trees, many of 
them American.] Mittheil. Deutsch. Dendr. Gesell. 3 : [reprint, pp. 
7-] 1894. 

Lodeman, E. G. Black Knot of Plums and Cherries. Bull. 
(Cornell) Exp. Sta. 81: pp. 24./.6. N. 1894. 

Gives bibliography, manner which the fungus enters the host, and method of suc- 
cessful treatment, 

Lodeman, E.G. Some Grape Troubles of Western New York. Bull. 

N. Y. (Cornell) Exp. Sta. 76: 410-454.f. 6. N. 1894. 

‘Deals largely with fungous diseases, with figures and treatment. 


NY. 


238 


Macoun, J. M. Contributions to Canadian Botany.—III. Can. Rec. 
Sci. 6: 198-210. 1895. [Contr. Herb. Geol: Surv. Canada, III. ] 

_ Record of numerous new localities and of several species additional to the Canadian 

flora. 

Meehan, T. Saptisia tinctoria. Meehan’s Month. 5: 81, 82. pl. 5. 
My. 1895. 


Meehan. T. John H. Redfield. Bot. Gaz. 20: 175, 176. 20 Ap. 1895. 


Moller, A. Brasilische Pilzblumen. Bot. Mitth. aus den T ropen, 7: 
pp. 152. pl. & 1895. 
Description and illustration of fungi in the genera Protudera, Clathrus, Colus, La- 


ternea, Blumenavia, Aporophallus, Mutinus, Ltajahya, Ithyphallus and Dictyo- 
phora. 


Nash, G. V. Notes on some Florida Plants. Bull. Torr. Bot. Club, 
22: 141-161, 18 Ap. 1895. 
New species in Helianthemum, Stipulicida, Rhynchosia, Rhexia, Hieractum, 
Asclepias, Breweria, Persea, Crotonopsis, Nolina, Xyris, Commelina and Steno- 


phyllus. 

Oliver, D. TZyricholaena Monachyron. “ook. Icon. Pl. 24: pl. 2374- 
Ap. 1895. 
A native of Cape Verde Islands introduced on St. Vincent. 

Pammel, L. H. Potato Scab and its Preventive. Bull. Iowa Exp. 
Sta, 27: 120-130. f. 3. 1895. 

Pammel, L.H. The pte ai of Fungi. Gard. & For. 7; 248 
249. 1894. 

_ Brief notes on Zxoascus deformans and Monilia fructigena. 

Pammel, L. H. Vergiftung durch Wasserschierling (Cicuta virosa 
var. maculata.) Pharm. Rund. 13: 102-103. f. 7-¢ My. 1895. 
Parish, S. B. Further Additions to the Flora of Southern California. 

. Erythea, 3: 58-62. 1 Ap. 1895. 


Robertson, C. Flowers and Insects.—_XIV. Bot. Gaz. 20: 139-1.49- 
20 Ap. 1895. 


Schmidle, W. Einige. Algen aus Denver, Colorado. Hedwigia, 34: 
84, 85. fig. 8 Ap. 1895. 
Describes Cosmarium mesochondrion new. 
Schneck, J- Observations on the Spider-flower. Bot. Gaz. 20: 168- 
170. fig. 20Ap. 1895. _ 
Notes on Cleome spinosa. : ? 
Shull, G. H. Observations on Ansdenia albida. Bot. Gaz. 20: 17° 
172. pl. 13. 20 Ap. 1895. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 236 


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Fic, 1. PROTOPHYLLUM QUERCIFORME HOLLICK. 


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LOBA HOLLI 


Fic. 2. CISSITES PLATANO 


Fic. 3. CISSITES ACUTI 


' 


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CONTENTS: 
An Enumeration of the Lichens of New- — [Becca Note ata 


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TORREY BOTANICAL CLUB. 
Vol. 22. Lancaster, Pa., June 25, 1895. s No. 6. 


An Enumeration of the Lichens of Newfoundland and Labrador, 


By JouHn W. ECKFELDT. 


The purpose of this paper is to present as full a list of all 
authenticated lichens that are to be found in the much unex- 
plored regions of Newfoundland and Labrador, as has been af- 
forded during a number of years’ sojourn in these lands. The 
entire field of exploration has been the work of the untiring Rev. 
A. C. Waghorne, who has been located there through parochial 
work, and during such residence has endeavored to secure all such 
plants as it has been possible for him to obtain. The labor has 
been attended with many difficulties; the inconveniences for trav- 
eling, the character of the climate, and the ruggedness of the 
country have hindered and delayed him in his efforts. The gen- 
eral character of these plants may be regarded as sub-arctic; 
many, however, are strictly more boreal in distribution, while 
others are of more southern climes. The determination of these 
plants has been attended with some difficulty, owing to the great 
tendency to variation, depending in part on the nature of the cli- 
mate, and the character of the substratum. The entire work has 
been subject to my supervision, and a number of the more diffi- 
cult species, as well as some new types (particularly in the Leca- 
norei and Lecidacei), have been under the kind observation of Dr. 
F. Arnold, of Munich, and Dr. J. Hulting, of Norrkoping. A few 
species are regarded as new and these will be described. The list 
embraces a large number of types common to both hemispheres. 


240 


Most of the lichens of Newfoundland and Labrador were collected 
from the following localities: Trinity Bay, on the southeastern 
coast; others from Notre Dame Bay, and a few from White Bay, 
further north from Placentia Bay to the Labrador. On the La- 
brador the collection was principally made in the Straits of Belle 
Isle, then along the coast from Battle Harbor north to Seal Is- 
lands, and thence about Sandwich and Gross waters to Hamilton 
Inlet. 
RAMALINA Ach. 
R. calicaris (L.) Fr. 
Var. canaliculata Fr. 
Var. farinacea Schaer. 
R. intermedia Del. 
Thallus whitish, pale straw color and glaucescent, erect, dicho- 
tomously branched, divided and the apices attenuated and divided 
with frequent soredia on the edges, the cortical layer more fila- 


mentous.. Apothecia terminal, subtended by elongated forked ex- 
tremities of the lacinia, pale yellow. Spores hyaline, ellipsoid 


uniseptate. 
R. rigida (Pers.) Tuck. 
R. pusilla Prev. 
Var. geniculata Tuck. 
R. minuscula Ny\. ( Alectoria minuscula Ny\.) 


Thallus densely entangled, small and quite scabrous, apothecia 


not seen in our specimens. A plant closely allied to R. pusilla 
Prev. from which it distinctly differs. 


R. polymorpha Ach. 
R. pollinaria Ach. 
R. pollinariella Ny\. 


Thallus small and finely divided, mostly sorediferous, erect, 


pale straw color. Plants generally sterile. R. minuscula. f. pol- 
finariella Ny. 


CETRARIA Ach. 
 C. muricata Ach. 


_ Thallus smaller than in the following species, more densely 
intricate and caespitose, compactly ramulose, terete to sub- 


terete, and frequently compressed, smooth, shortly branched, 
the branches becoming spinulose at the apices. 


C. aculeata Ach. 
C. Richardsonit Tuck. 


241 


C. arctica Hook. 

C. oaontella Ach. 

C. Fendlert Tuck. 

C. commixta Nyl. 

Thallus clear bright brown to fuscescent, smooth, irregularly 
laciniate, divided; laciniae erect, much entangled, plane, the mar- 
gins not elevated, apices obliquely furcate, uniform in color, be- 
neath paler; apothecia reddish brown, disk rugose, margin granu- 
lose ; spores simple, hyaline, oblong, — mic. 

C. Fahlunensts (L.) Schr. 

C. Istandica (L.) Ach. 

Var. crispa Ach. 
Var. Delisaei Borr. (C. htascens Th. Fr.) 

C. ciliaris Ach. 

C. lacunosa Ach. 

C. nivalis Bell. 

C. aureus Tuck. 

C. glauca (L.) Ach. 

Var. fusca Fl. Thallus brown and dark. 

C. juniperina L. 

Var. Pinastri Ach. 
Var. serrestris Schr. 
C. saepincola Ehrh. 
C. saepincola var. chlorophylla Wahl. 


Evernia Ach. 
E. prunastri (L.) Ach. 


Usnea Ach. 
U. barbata (1..) Fr. 


Var. ceratina Schr. 
Var. jirta Fr. 
F. strigosa Ach. 
Var. florida Fr. 
F. glabrescens Nyl. 
Var. dasypoga Fr. 
Var. plicata Fr. 
U. longissima Ach. 
U. trichodea Ach. 
U. cavernosa Tuck. 


242 


ALEcToRIA Ach. 


A. divergens (Ach.) Nyl. 
A. jubata L. 
Var. implexa Fr. 
Var. chalybetformis Ach. 
Var. bicolor Fr. 
Var. prolixa Ach. 

A. capillaris (Ach.) Nyl. (A. judbata var. cana Ach.) 

Thallus pale, hoary to yellowish-rufescent, much elongated 
and frequently branched ; branches flaccid, densely entangled and 
somewhat compressed, angular, lacunose beneath; apothecia un- 
known. , 

A. ochroleuca (Ehrh.) Nyl. 

Var. vigida Fr. 
Var. osteina Nyl. 
Var. nigricans Ach. (A. nigricans Ny}.) 
Var. circinata Fr. 
~ Var. sarmentosa Ny\l. 


THELOSCHISTES Norm. 


L chrysopthalmus (L.) Norm. 
T. parietinus (L.) Norm. 
Var. aurcola Ach. 


Thallus brighter yellow to orange, rounded, lobate, sinuately 
incised, thicker, crenate, the margins erect, granulate; apothecia 
sub-crenate. 


T. lychneus Ny). 
T. polycarpus Ehrh. 
Var. pygmaeus Fr. 
_T. concolor Dicks. (Xanthoria candelaria Ny\.) 


PARMELIA Ach. 
P. aurulenta Tuck. 


P. tiliacea (Hoff.) Flk. 
Var. sublaevigata Ny). 
P. cetrata Ach. 
P. Borreri Turn. 
Var. rudecta Tuck. 
P. saxatilis (L.) Fr. 


243 


P. saxatilis var. panniformis (Ach.) Schr. 

P. saxatilis var. sulcata Ny. 

P. saxatilis {. furfurascens Schr. Thallus very furfuraceous. 

P. saxatilis var. omphaloides Fr. 

P. fraudans Ny}. 

This has been regarded as a subspecies of P. saxatilis and fre- 
quently taken for that species, but has distinctions worthy of 
specific rank. 

P. physodes (U..) Ach. 

Var. vittata (L.) Ach. 
Var. enteromorpha Tuck. 
Var. odscurata Ach. 

P. encausta (Sm.) Nyl. 

P. encausta var. alpicola Ny). 

P. colpodes (Ach.) Nyl. 

P. olivacea (L.) Ach. 

P. olivacea var. aspidota Ach. 

P. olivacea var. panniformis Nyl. 

P. olivacea var. sorediata (Ach.) Nyl. 

P. lanata (L.) Wallr. 

P. stygia (L.) Ach. 

P. conspersa (Ehrh.) Ach. 

P. leucochlora Tuck. 

P. centrifuga (L.) Ach. 

P. incurva (Pers.) Fr. 

P. ambigua (Wulf.) Ach. 

Var. albescens Wahl. (P. hyperopta Ach.). 


Puyscia DC. 


P. hypoleuca (Mull.) Tuck. 

P. comosa Esch. (P. galactophylla Willd). 
P. leucomela (L.) Michx. 

P. ciliaris (L.) D. C. 

P. ciliaris var. crinalis Schr. 

P. aquila var. detonsa Tuck. 

P. pulverulenta (Schreb.) Nyl. 

P. pulverulenta var. leucoleiptes Tuck. 

P. pulverulenta £. muscigena Auct. 


244 


P. stellaris L. 
P. stellaris {. adscendens Fr. 
P. astrotdea (Fr.) Nyl. 
P. tribacia (Ach.) Tuck. (P. erosa Bort.) 
P. hispida Schreb. (P. tenella Ach.) 
P. crispa (Bers.) Nyl. 
P. caesia (Esch.) Nyl. 
P. obscura (Ehsh.) Ny). 
Var. endochrysea Ny. 


Umpsiuicaria Hoff. 
U. rugifera Nyl. 


U. cylindrica (L.) Del. 

U. proboscidea (L..) Sten. 

U. arctica Ach. (U. proboscidea v. arctica (Ach.) Tuck.) 

U. anthracina (Wulf.) Schr. 

U. polyphylla (L.) Hoff. 

U. flocculosa Hoff. 

U. hyperborea Hoff. 

U. erosa (Web.) Hoff. 

U. Muhlenbergu (Ach.) Tuck. 

U. hirsuta (Ach.) Sten. 

Var. grisea Th. Fr. 

U. vellea (L.) Nyl. 

U. Pennsylvanica Hoff. 

U. spodochroa Ehrh. 

Thallus varying from moderately large and irregular to small 
and orbicular, thin, flattish and spreading, with an irregular 
waving crenate margin in the smaller forms, pale ashy to light 
purplish brown in older specimens, paler at the centre and mealy; 
beneath brown and minutely granulose and thickly covered 
with irregular stout brownish fibrils; apothecia very numerous, 
sessile and scattered towards the border, elevated, stout and robust, 
orbicular, disk prominent, rounded and conspicuously papuloid, 
surrounded by a thick, even or fissured margin, smooth, black; 
spores ellipsoid, simple, 3 mic. 

The occurrence of this Scandinavian lichen in this country is 
rather remarkable. 

U. pustulata (L.) Hoff. 

Var. papulosa Tuck. 


245 


U. LapraporensE Hulting n. var. 


Thallus thinner than the preceeding, small, from ash color be- 
coming olivaceous or brown, rugulose, papulose, papules scattered, 
remotely prominent, beneath finely granular, foveolate, foveoli 
small, contracted, sanguineous within; apothecia reddish-brown, 
orbicular with a well raised margin I-2 mm. broad; spores large, 
orbicular $2 mic. Onrocks associated with the preceding variety. 
Labrador. 


Sticta Schreb. 


S. amplissima (Scop.) Mass. 

S. herbacea (Huds.) Ach. 

S. pulmonaria (L.) Ach. 

S. guerizans (Michx.) Ach. 

S. limbata (Sm.) Ach.; evidently quite new to this region. 
S. aurata (Sm.) Ach. 

S. scrobiculata (Scop.) Man. 


NEPHROMA Ach. 


N. arcticum (L..) Fr. 
LV. expallidum Ny). 
NV. laevigatum Ach. 
Var. parile Ny). 
N. Lusitanicum Schr. 


PELTIGERA Willd. 


P. venosa (L.) Hoff. 
P. apthosa (L.) Hoff. 
P. polydactyla (Neck.) Hoff. 
P. rufescens (Neck.) Hoff. 
P. pulverulenta (Tayl.) Nyl. (P. scabrosa Th. Fr.) 
P. matacea (Ach.) Fr. 
P. canina (L.) Hoff. 
Var. spuria Ach. 
Var. sorediata Schr. 
Var. spongiosa Tuck. 


SoOLORINA ACH. 


S. crocea (L.) Ach. 
S. saccata (L.) Ach. 


246 


PuysMA Mass. 


P. luridum Mont. A single specimen found. 


PANNARIA Del. 


P. hypnorum (Hoff.) Koerb. 

P. lanuginosum (Ach.) Koerb. 

P. rubiginosa (Thunb.) Del. 

P. leucosticta Tuck. 

P. brunnea (Sw.) Mass. (P. pestzoides Sw.) 
P. microphylla (Sw.) Del. 

P. lepidiota Th. Fr. 

P. tryptophyllum (Ach.) Man. 

P. carnosa Dicks. 


P. WAGHORNEI n. sp. 


Thallus thickened, congested and irregular, fissured, conglom- 
erate, coarsely granulate, rugulose, from reddish brown to pur- 
plish and seated upon a thin limiting hypothallus; apothecia scat- 
tered but frequently conjoined, small, rounded and erect, and at 
length flat, with a prominent margin, from pale reddish to 


rufescent ; spores ellipsoid, simple, aa mic. 


On turfy earth. Evidently a distinct and well marked species, 
and not closely related to any other. 


P. plumbea (Leight.) Del. 


CoLtemaA Hoff. 
P. melaenum Ach. 
P. flaccidum Ach. 


Leprocium Fr. 


L. lacerum (Sw.) Fr. 
L. tremelloides (L.) Fr. 


Var. cyanescens Ach. (Collema cyanescens Schaer.) 


PLacoptum DC, 
P. elegans (Link.) DC. 
P. murorum (Hoff.) DC, 


F. minitatum Hoff. Thallus fulvescent, punctate; apothecia 
bright. 


P. cirrochroum (Ach.}; Hepp. 


247 


P. auranhacum (Leight.) N. H. 
P. cerinum (Hedw.) N. H. 
Var. pyraceum Ny)l. 


F. pyrithronum Ach. Thallus gray to brownish, more or less 
rugulose; apothecia small, sunken and concave, pale green to buff- 
colored. Spores eee mic, 


P. crenulatum Wahl. 
P. cvitellinum (Ehrh.) N. H. 
Var. aurellum Ach. 


LecaAnora Ach. 


L. rubina (Vill.) Ach. 
L. muralis (Schrb.) Schr. 
Var. Garovagli Anzi. 
Var. saxicola Scur. 
L. pallida (Schrb.) Schr. . 
L. sordida (Pers.) Th. Fr. 
L. atra Huds. 
L. dispersa (Pers.) Nyl. 
L. badia (Pers.) Ach. 
L. Hageni Ach. (L. umbrina Mass.) 
F. sostericola Nyl. 
L. subfusca (L.) Ach. 
Var. hypnorum (Wulf.) Ach. 
Var. argentata Ach. (L. campestris Schr.) 
Var. atrynea Ach. 
Var. coilocarpa Ach. 
Var. distans Ach. ( L. chlarona Ach.). 
F. rugosa Pers. 
L. symmictera Nyl. 
Closely allied to Z. varia v. symmicta Ach., but differs as to 
the reaction. K—C. orange. 
L. atrosulphurea (Wahl.) Ach. 


L. OcHRACEORUBESCENS Arnold n. sp. 

Thallus scattered, elevated, roundish or irregular, greenish or 
yellow, granular, granules at length becoming tortulose and dis- 
tinct ; apothecia prominent, elevated, biatorine, sessile and finally 
conglomerate, proper margin depressed or quite wanting, brown- 
ish green to olivaceous with a delicate bloom, spores ellipsoid 


248 


hyaline, °—tmic. On sandstone. This species is closely re- 
lated to L. Swartsit Ach. 
L. varia (Ehrh.) Nyl. 
Var. symmicta Ach. 
Var. intricata Nyl. 
Var. saepincola Fr. 
Var. polytropa Ehrh. 
F. dlusoria Ach. 
Thallus evanescent, thin, apothecia dispersed, infrequent, large, 


disk plane, frequently convex, pale yellow, margin frequently oblit- 
erated. 


L. ventosa (L.) Ach. 
L. elatina Ach. 
Var. ochrophoea Tuck. 
Var. minor Tuck. (Haematomma Crsmonica Belt). 
L. pallescens (L.) Schr. 
Var. rvosella Tuck. 
L. tartarea (L.) Ach. 
F. leprosa Ny}. 
F. telephoroides Th. M. Fr. 
F. grandinosa Ach. 
F. macrocarpa Th. Fr. 
L. cenisia Ach. 
L. cinerea (L.) Somf. 
Var. laevata Fr. 
L. calcarea (.) Somf. 


_ L. cCINEREO-RUFESCENS (Ach.) Eck. 


Thallus dark ashy white, paler towards the border, verrucose 
to areolate, rimose; hypothallus black; apothecia sunken in the 
thallus, convex, disk dark brown, bordered by an entire thalline « 
margin ; spores hyaline, ellipsoid, simple, “mic. On rocks. 

L. CAESIO-CINEREA (Nyl.) Eck. 

Thallus ashy white to darker, thick, verrucose, areolate, broken; 
apothecia sunken in the areoles, concave, terminating in a white 
thalline border which finally becomes more or less prominent, 
spores hyaline, oblong, simple, <3 mic. On rocks. 

L. gibbosa Nyl. : 

L. fuscata (Schr.) Th. Fr. 

Var. rifescens Th. Fr. 


249 


L. glaucocarpa (Wallr.) Ach. 
F. distans Arn. 
L. privigna (Ach.) Nyl. 


RINODINA Mass. 


R. sophodes (Ach.) Nyl. 
R. turfacea (Wall.) Nyl. 


Prertusaria DC. 

P. bryantha (Ach.) Nyl. 
P. velata (Turn.) Nyl. 
P. panyrga (Ach.) Th. Fr. 
P. multipuncta (Turn.) Nyl. 
P. dactylina (Ach.) Nyl- 
P. rhodocarpa Koerb. 
P. communts D. C.. 

F. rupestris D.C. 
P. letoplaca (Ach.) Schr. 
Plaevigata (Nyl.) Arn. (P. letoplaca var. laevigata Th. Fr.) 
P. subobducens Nyl.(P. glomerata Schr.) 
P. protuberans Smf. 
P. globularis Ach. 


URCEOLARIA Ach. 


U. scruposa (L..) Nyl. 
Var. dryophila Esch. 


THELOTREMA Ach. 


T. lepadinum Ach. 
7. subtile Tuck. 
STEREOCAULON Schr. 
S. coralloides Fr. 
S. paschale (L.) Fr. 
S. denudatum F\. 
S. condensatum Hoff. 
S. tomentosum Th. Fr. 
S. alpinum (Th. Fr.) Laur. (5S. comentosum var. alpinum Th. Fr.) 
S. pileatum Ach. 
S. nanodes Tuck. 


250 


PILoPHORUs Th. Fr. 


P. cereolus Ach. 
Var. fibula Tuck. 


CLaponia Tuck. 
C. symphicarpa Fr. 


Var. epiphylla (Ach.) Nyl. 
C. cariosa (Ach.) Spreng. 
C. decorticata Fik. 
C. pyxidata (L.) Fr. 
Var. simplex Hoff. 
F. costata F}, 
Var. pocillum Ach. 
C. fimbriata (L.) Fr. 
Var. tubaeformis Fr. 
Var. radiata Fr. 
C. degenerans F', 
C. gracilis (L.) Nyl. 
Var. verticillata Hoff. 
Var. verticillata f. phyllophora Eth. 
Var. hybrida Schr. 
Var. elongata Fr. 
F. macroceras Tuck. 
F. chordalis Tuck. 
F. amaura F\, 
F. aspera FI. 
C. cornuta (L.) Fr. 
C. turgida (Ehr.) Hoff, 
Var. conspicua (Ach.) Nyl. 
C. papillaria (Esch.) Hoff. 
C. cenotea (Act.) Schs, 
Var. furcellata Fr. 
C. squamosa Hoff. 
‘F. turfacea Rehm. 
F. ventricosa Fr. 
F. densicollis Hoff. 
F. asperella Fk. 
C. furcata (Huds.) Fr, 
Var. crispata Fl, 


251 


Var. racemosa F\. » 
Var. subulata F\. 
Var. sguamulosa Schr. 
Var. adspersa F1. 
C. rangiferina (L.) Hoff. 
C. syévatica L. 
Var. alpestris L. 
C. Botrytes (Hag.) Hoff. 
C. carneola Fr. 
Var. cyanipes Somf. 
C. amaurocraea (F1.) Schr. 
C. uncinalts (L.) Fr. 
Var. adunca Auct. 
Var. Caroliniana Tuck. 
Var. turgescens Fr. 
C. Boryi Tuck. (C. lacunosa Bory.) 
C. chlorophea L. 
F. semplex Hoff. 


C. MULTIBRACHIATA (FI.) Eck. 

Thallus ashy brown, squamulose, mealy ; podetia erect, slender, 
mostly smooth with repeated proliferations or branches, pervious ; 
ant is somewhat allied to C. Floerk- 


spores curved,” mic. This pl 
3 nsidered by 


eana Fr., from which it is not easily separated, but co 
Dr. Arnold as distinct. 
C. Floerkeana Fr. 
C. cristatella Tuck. - 
Var. ochrocarpia Tuck. 
C. pulchella Schw. 
C. macilenta (Esch.) Hoff. 
F. clavata Ach. 
C. digitata F\. 
F. drachytes Ach. 
F. monstrosa Ach. 
C. deformis (L.) Hoff. 
C. coccifera L. (C. cornucopioudes (L.) Fr.) 
F, pleurotica Ach. 
C. bellidiflora (Ach.) Schr. 
Var. Hookeri Nyl. 


252 


THAMNOLIA Ach. 
T. vermicularis (Sw.) Schr. 


Baromyces Fee: 
B. aeruginosus Scop. 
B, byssoides (1.) Schr. 
B. roseus Pers. 


Var. fungoides (Sw.) Ach. (Sphyridium fungiformis Scop.) 


Biatora Fr. 

B. granulosa (Esch.) Poetch. 

Var. escharoides Esch. Thallus very minutely granulose, yellow 
or ashy white ; apothecia confluent, diffiform, strongly convex and 
immarginate, dark brown to reddish chestnut; spores simple, = 
mic. On clay soil. 

: B. coarctata Sm. 
Var. elascista Ach. 
Thallus white or greyish, very thin, leprous and effuse, apothecia 


sessile, quite innate, concave or plane with a weil defined promi- 
nent white coarctate margin. Spores similar. 


B. flexuosa Fr. 

B. russula (Ach.) Mont. 

B. cinnabarina (Somf.) Fr. 

B. vernalis (L.) Fr. 

Var. helvola Koerb. 

B. sanguineo-aira (Fr.) Tuck. Spores about "=" mic. 

b. Berengeriana Ny. 

This plant is allied to B. sanguneo-atra Fr. but may be readily 
distinguished by the absence of violet or bluish granules found 
among the paraphyses in that species. Spores ae mic. 

B. rufo-fusca Anzi. 

B. turgidula Fr. 

B, rwulosa (Ach.) Fr. 

B. ARCUATULA (Arn.) Eck. 

Thallus flat and thickened, areolate, rimulose, olivaceous to 

fuscescent; apothecia brownish-black to very black, margin ele- 


vated concolorous and shining, epithecium brown, spores hyaline, 
simple, 2—4-guttate, somewhat curved and obtuse at the apex, 


“=;' mic. A species closely related to B. rivulosa (Ach.) Fr. On 
rocks. 


ii 
ee 


253 


B, SCRUPULOSA n. sp. 


Thallus of thickened coalesing granules, areolate, rimulose, 
pale olive to yellowish brown, areoles adnate, contiguous, mingling 
into a continuous uneaven crust; apothecia scattered, diffiform, 
erect, black, plane and somewhat depressed, surrounded by a tor- 
tulous erect concolorous margin; spores simple, broadly ellipsoid, 
hyaline, = mic. A well marked species which seems to approach 


4. Kochiana Hepp. 


B. ALBO-FUSCESCENS (Nyl.) Eck. 


Thallus thinly diffused and even over the surface, plane and 
slightly granular or furfuraceous, or at length quite smooth and 
glaucous; epithecium pale; apothecia very sparse, rounded and 
elevated, at length flat; disk brown and prominent, reddish, mar- 


gin elevated. Spores ellipsoid, pointed, i mic. On rocks. 


B. mollis Wahl. 

B. Nylanderi Anzi. 

B. uliginosa (Schr.) Fr. 

B. varians Ach. ‘B. exigua Chaub.) 

B. mixta Fr. (B. tricolor Ny).) 

B. Laureri Hepp. 

L. Heeria Hepp. Parasitic on Solortna saccata. 

B. STEREOCAULORUM (Th. Fr.) Eck. (Biaéorima Lich. Miquelon, 
Arnold.) Thallus mostly wanting; apothecia very small, black, 
plane; epithecium brown, hymenium colorless to bluish, para- 
physes loosely disposed; spores hyaline, bilocular, 8 in thekes, a8 
mic. Parasitic on Stercocaulon alpinum. 

B. sphaeroides Dick. 

B. hypnophila Turn. 

B. artyta Ach. 

B. rubella Esch. 

B. rupEstrIs (Sch.) Eck. Thallus cinerascent, thin and closely 
adnate, apothecia flat, innate, smooth, rufescent with a raised mar- 
gin around the sunken disk. Spores immature. On rocks. 

B. muscorum Sw. 

B, umbrina Ach. 

B. opscuraTa (Sm.) Eck. Thallus composed of minute pale 
ashy to white rugose granules; apothecia minute, rounded and 
flat, dark brown to black, smooth and shining with a receding 
Margin, very numerous. Spores broadly ellipsoid 4-locular, ea 
mic. On dead moss. 


254 


B. PALLIDA (Arn.) Eck. 


Thallus thin, chinky and minutely granulose, sparsely scattered 
over a thin but even surface, yellowish brown to fuscous ; apothecia 
small, rounded, black, prominent, slightly flattish or convex, with a 
concolorous thalline margin; hypothallus mostly wanting. 
Spores fusiform, 6-locular, pee mic. On smooth barks. 

B. CAESIO-RUFA (Ach.) Eck. 

Thallus thin, white, scattered irregularly, of rounded and closely 
adnate granules, but forming an even uniform crust; apothecia 
sessile, minute, quite black, turgid, and becoming flat, with a per- 
sistent regular elevated margin; spores mostly 4-locular, ue mic. 
On smooth barks. 


HETEROTHECIUM Flot. 

fT. grossum Pers, 
HI sanguinarium (L.) Flot. 

Var. endorhoda Th. Fr. 

Var. effusa Ach. 
H, alpina Ach. 
fT, porphyrites Tuck. 
H. leucoxanthum (Spreng.) Mass. 
HT, pezizoideum (Ach.) Flot. 


Lecipea Ach. 

L. panaeola (Ach.) Fr. 
L. platycarpa Ach. 

Var. superba Ach. 
L enteroleuca (Ach.) Fr. 

Var. diasemoides Ny). 
L fusco-atra L. 
L fuscescens Somf. 
L. polycarpa Fr. 

Var. declinans Nyl. Thallus mostly crustaceous; hypothe- 
cium fuscous. 

Var. sudblactea Somf. 


L. CRENULATA (Dicks.) Eck. 


Thallus thin, evanescent, cinerascent to pale yellowish grey > 
apothecia black, plane and simple, naked, with a raised flexuous 


crenulate proper margin; spores ellipsoid, hyaline, simple, + mc- 
On rocks. ; 


255 


L crustulata Ach. 

Thallus pale to brownish or often grey, adnate, thin, tartareous 
effuse, subrimulose to areolate; apothecia numerous, dispersed, 
sessile and simple, black with a distinct thin margin; hypothecium 
swollen, nearly brown; spores 8 in the thekes, simple, oblong, * 
mic. On rocks. 


L albocoerulescens (Wulf.) Schr. 
Var. flavocoerulescens Schr. 

Var. cinercoatra Ach. 

. variegata Fr. (L. pantherina Th. Fr.) 
. macrocarpa D.C. 

. lessellata F 1. (L. cyanea Th. Fr.) 
. lapicida (Ach.) Fr. 

. Leucothallina Ach. 

. empetrea Nyl. 

. tenebrosa F\. 

L. latypea Ach. 


L. DISTANS n. sp. 

Thallus thin, evanescent, but at length becoming minutely 
granulose and frequently continuous over a scurfy surface; apo- 
thecia mostly subsessile, elevated, large, flat, black with a depressed 
disk surrounded by a conspicuous granulose and crenate margin ; 
disk covered by a delicate evanescent thin bloom; hypothecium 


pale ; spores long, ellipsoid, simple, =, mic. On rocks. Labrador 


L. contigua var. metospora Ny}. 
Thallus thin, ashy white, areolate, rimulose; apothecia black, 
plane, with a proper margin, disk somewhat pruinose, spores sim- 


ple, hyaline, 2° mic. On rocks. 


L. Kochiana Hepp. 

Thallus pale ashy brown to fuscescent, rimose, areolate, 
broken, areoles sunken, plane, bordered by a black margin; apothe- 
cia innate, black, immarginate, flexuous or slightly angulose, diffi- 
form; disk plane, polished and smooth; spores 8 in the thekes, 


oblong, simple, =" mic. On rocks. 


L. dispansa Nyl. (L. expansa Ny.) 

Thallus furfuraceous, rimulose, thin, black; apothecia very 
numerous, minute, sessile, scattered, black, plane, the margin very 
smooth; hypothecium thin, fuscous; epithecium hyaline; spores 
about 8 in the thekes, simple, ellipsoid, “~~ mic. On rocks. 

L. auriculata Th. Fr. 

L. sylvicola (Flot.) Nyl. 


» id 
7-8 


HANNAN 


256 


L. CONFERENDA (Nyl.) Eck. 


Thallus verruculose, granulate, obscurely cinerascent ; apothecia 
very small, black, convex and glaucous; spores oblong, simple, 
hyaline, ay mic. On rocks. 


BuEtuia De Not. 


B. capitulata Th. Fr. var. erudbescens (Flot.) Eck. 


Thallus of thin smooth flat discrete granules, becoming at 
length areolate; areoles irregular and evanescent at the outer mar- 
gin; apothecia scattered, more or less sessile, small and very 
black, flat and bordered by a well raised margin; spores ellipsoid, 


bilocular, fuscescent, oy mic. On smooth barks. 


B. punctiFrormis (Hoff.) Eck. 


Thallus very thin, scurfy, minutely granular, grey or whitish, 
becoming at length nearly obsolete or with a delicate coloration ; 
apothecia small, flat, granular and the disk quite rugose, brownish- 
black to very black, with a raised stout thick granular margin ; 
_ spores bilocular, +? mic. On smooth barks. 

B. albo-atra (Hoff.) Th. Fr. 
B. parasema (Ach.) Fr. 

B. disciformis Fr. 

B. spunia Schr. 

B. lepidastra Tuck. 

B. myriocarpa D. C. 


B. aTRATA (Sm.) Eck. 


Thallus blackish to ashy brown, areolate, broken, areoles plane, 
small, smooth, convex, hypothallus black; apothecia springing 
from the hypothallus, usually innate, small, appressed, margin thin, 
entire ; spores 8 in the thekes, bilocular,fuscous, 2-7 mic. On rocks. 


B. colludens Ny}. ne 
B. ExcENTRICA (Nyl.) Eck. 


Thallus brownish, white or gray, thin, sparingly more or less 
broken; apothecia pale, black, sessile, plane with a thickened ob- 
tuse margin. Spores we mic., the cells murilocular. 

B petraea (Flot.) Koerb. 

Var. grandis Fk. 
Var. Montagnei Tuck. (B. atro-alba Fr.) 

F. empetraca Nyl. 


257 


B. conropsoipEuM (Hepp.) Eck. 

Thallus consisting of scattered white irregular flat scales, which 
are diffused over a black hypothallus; apothecia arising from the 
hypothallus, sparingly scattered, black, flat and somewhat promi- 
nent, with a raised rugose margin; disk roughened and finely pa- 
pillose; spores ellipsoid, 4-locular, sat mic. On rocks. 


B. geographica (F1.) Tuck. 
B. pertusaricola Willey. 

B. parmeliarium Som. 

B. saxatilis (Sch.) Koerb. 
B. scabrosa (Ach.) Koerb. 


B, opscurata (Ach. Koerb.) Eck. 

Thallus pale to fuscesent or rufofuscescent, thin, finally areo- 
late, continuous, areoles plane, simple; apothecia black, naked, 
plane, sessile, innate or strongly adnate with a thicker obtuse pale 
margin; spores 8 in the thekes, ellipsoid, murilocular, ss mic. 
On rocks. pies 


B, r1vuraris (Flot.) Eck. ‘ 

Thallus pale olive brown, areolate, diffiform, areoles flat upon a 
black hypothallus; apothecia large, flat, sunken in the areoles, black 
with a prominent raised waving margin; spores broadly ellipsoid, 
bilocular, fuscous, os mic. On rocks. 


B. Copetanp1 (Koerb.) Eck. 

Thallus composed of large irregular coarse whitish granules, 
becoming areolate, dispersed, seated upon a black hypothallus; 
apothecia large, black and prominent, more or less finely turbu- 
culate, but becoming flattish and concave, with a persistent mar- 
gin; spores ellipsoid, bilocular, fuscescent, aa mic. On rocks. 


B. badioatra (F 1.) Schr. 


B, concreta (Koerb.) Eck. 

Thallus minutely granulose, closely adnate, pale ashy to 
whitish, bordered by a flexvous hypothalline line; apothecia ses- 
sile, minute and innate, immarginate with a thickened brown 
border; spores ellipsoid, murilocular, 4a mic. On rocks. 


B, cALcaREA (Weis) Eck. 

Thallus determinate, white, roundish, of mealy or tartareous 
granules, areolate, rimulose towards the centre, configurate at the 
border ; apothecia large, black, innate, plane, bluish, pruinose, con- 
vex, surrounded by a white thalline margin, proper margin dark, 
persistent; spores ellipsoid to oblong, 4-locular, = mic. On 
rocks. 


258 


B. RUBESCENS (Arn.) Eck. 


Thallus very thin, white, uneven and broken up into areole- 
like flat scales; apothecia scattered, sessile, small, from flat to con- 
cave, with a prominent raised margin but frequently wanting, from 
reddish-brown to black; spores broadly ellipsoid, bilocular, = 
mic. On smooth barks. 


LecaANnactis (Esch. Kbr.) Tuck. 


L. abtetina Ach. 
L. premnea Ach. 


OPEGRAPHA (Humb.) Ach. 
O. zonata Kbr. 


Thallus rufous to ferruginous, thin, subtartareous, smoothish, 
covered by nnmerous yellow or pale whitish soredia, marked at 
the border by conspicuous thick erect dark lines forming the 
hypothallus ; apothecia fuscous to black, scattered, sparse and ses- 
sile; spores hyaline, contracted, fusciform, pointed, °° mic. On 
rocks. Sie 


O. atra Ach. 
_O. vulgata Ach. 


XYLOGRAPHA Fr. 
X. opegraphella Nyl. 
X. parallela Fr. 
GrRaAPHIS Ach. 

G. scripta (L.) Ach. 

Var. recta Schr. 

Var. varia Ach. 

Var. limitata Schr, 

Var. pulverulenta Pers. 

F. radiata Leight. 

F. flexuosa Leight. 

F. typographa Willd. 


ARTHONIA Ach. 
A. radiata Pers. 


Var. Swartzoidea Nyl. 
A. spectabilis F\. 
A. punctiformis Ach. 
A. patellulata Nyl. 
A. raunidia Ny). 


259 


SPHAEROPHORUS Pers. 
S. fragilis (L.) Pers. 
a corallordes Pers. (S. globiferus (L.) D.C.) 


Acotium Fee. 
A, tigillare (Ach.) DLN. 


Caticrum Ach. 
C. subtile Fr. 


C. turbinatum Pers. 
ENDOCARPON Hedw. 


£. mintatum (L.) Schr. 
Var. complicatum Schr. 
Var. fulvofuscum Tuck. 
£.. cinereum Pers. 


SEGESTRIA Fr. 


S. majusculum Ny1. 


TRYPETHELIUM Ach. 


T. virens Tuck. 


SaGEpIA Mass. Kbr. 


S. Cestrensts Tuck. 
S. oxyspora (Nyl.) Tuck. 


VERRUCARIA Pers. 


V. maura Wahlb. 

Thallus dark reddish or jet black, tumid, polished, areolae 
smooth covered with minute points; apothecia immersed and 
enclosed in the thallus, scattered, roundish ; epithecium conspicu- 
ous, poriform ; perithecium spreading at the base, internally black; 


12-16 


spores oblong, simple, 7 mic. On rocks. 


V. epigaea Pers. 

V. theliodes Smf. 

V. bryophila Lonnr. (Polyblastia). 

V. rimosicola Leight. 

Thallus wanting; apothecia black, minute, sessile, rounded, 
shining and polished; perithecium entire, black; epithecium pori- 


on 
‘ « 
*, 


260 


form; spores 8 in. the thekes, brown, oblong to linear, 4-locular, 
constricted, 5-2 mic. Parasitic on Buellia excentricum. 


PYRENULA Ach. 
P. punctiformis Ach. 
P. glabrata Ach. 
P. lactea Mass: 
P. thelaena (Ach.) Tuck. 
P. nitidella Mull. 


A Diatomaceous Deposit from an Artesian Well at Wildwood, N. J. 


By CHARLES S. BOYER. 


An artesian well sunk recently at Wildwood, N. J., exhibits at 
certain depths diatomaceous deposits of unusual interest. Mr. 
Woolman, whose researches into the geology of artesian borings 
are well known, has sent me samples of earths with the request that 
I enumerate the diatoms found therein. The accompanying list 
includes specimens occurring in a bed from 78 to 180 feet deep, of 
which Mr. Woolman remarks in the annual report of the Geolog- 
ical Survey of New Jersey for 1893 (page 401) that “a correspond- 
ing clay bed does not exist beneath the beaches to the northward, 
if we may judge by the specimens of borings furnished from the 
various wells or by the records where no specimens were ob- 
tained. It was probably in some way associated with the deposits 
of the Delaware River delta in a somewhat recently past geologi- 
cal age, and before the present peninsula of Cape May was formed.” 

It is not the purpose of the present paper to give a list of the 
forms obtained from the entire series of the well boring which has 
reached the depth of 1245 feet, passing through beds apparently 
identical with the Miocene deposits noticed at Atlantic City and 
which have already been described, but as the fresh water forms 
are very numerous and as there are associated with them certain 
marine and brackish water species not heretofore noticed in North 
America, the enumeration of all the diatoms thus far seen at the 
depths of from 78 to 180 feet, is here given. 


261 


The occurrence of Polymyxus coronalis is of peculiar interest, 
as it has been hitherto considered as growing exclusively at the 
mouths of the Para and the Amazon. It is not a mere accidental 
form, but is abundant in the deposit in all its numerous variations. 
Several years ago I found one specimen in material from one of 
the Atlantic City wells, but supposed at the time that it might 
have been a waif cast upon the shore. As Polymyxus now 
flourishes only in warmer waters the conclusion must be reached 
that the Delaware River delta formed the deposit under conditions 
quite different from those existing at the present time. It is at 
any rate now extinct along the North Atlantic coast. 

The nomenclature of the following list is based upon DeToni’s 
Sylloge Algarum, except in a few cases. 

Achnanthes Hudsonis Grun. 

A. inflata (Kiitz.) Grun. 

A. subsessilis Ehr. 

Actinocyclus Ehrenbergit Ralfs. 

A, subtilis Greg. 

Actinoptychus heliopelta Grun. forma minor. 

Only one specimen has been noticed. 

A, undulatus Ehr. 

A. vulgaris Schum. 

Amphora ovalis (Bréb.) Kiitz. var. gracilis (Ehr.) V. H. 

Aulacodiscus Argus Ehr. 

Auliscus pruinosus Bail. 

Biddulphia Rhombus Wm. Sm. 

Brebissonia Boecku (Kiitz.) Grun. 

I have noticed this form in material from the Saguenay River, 
Can. 

Campylodiscus echenets Ehr. 

The habitat of this species is usually given as marine or brack- 
ish water. Van Heurck is the only authority, as far as I have 
noticed, who states that it occurs in fresh water. I have found 


several specimens in mud taken from a supply reservoir in Phila- 
delphia, the water of which is drawn from the Schuylkill River. 


Cerataulus laevis Roper. 
Cocconets placentula Ehr. 
Coscinodiscus Argus Ehr. 
C. excentricus Ehr. 


262 


C. fasciculatus A. Schm. 

C. marginatus Ehr. 

C. minor Ehr. 

C. nitidulus Greg. 

C. Oculus-Iridis En. 

C. radiatus Ehr. 

C. subtilis Ehr. 

Cyclotella Kitzingiana Thw. 
Cymbella affinis Kiitz. 

C. cistula (Hempr.) Kirchn. 
C. cuspidata Kiitz. 
C. cymbiformis Ehr. 

C. Ehrenbergu Kitz. 

C. gastroides Kiitz. 

C. lanceolata (Ehr.) Kirchn. 
C. tumida (Breb.) V. H. 
Lncyonema ventricosum Kiitz. 


Cystopleura (Epithemia) Argus (Ehr.) Kunze. 


An abnormal form with flexuose outline has been noticed. 


C. gibba (Ehr.) Kunze. 

C. gibberula (Ehr?) Kunze. 

C. Musculus (Kiitz.) Kunze. 

C. Musculus constricta (Bréb.) V. H. 
C. Sorex (Kiitz.) Kunze. 

C. Zebra (Ehr.) Kunze. 

Eunotia Arcus Ehr. 

£:, diodon Ehr. 

E. impressa Ehr. 

£. major (W. Sm.) Rab. 

E. parallela Ehr. 

E. pectinalis (Dillw ?) Rab. 

E. praerupta Ehr. 

E. robusta Ralfs. 

E. tetraodon Ebr. 

E. triodon Ehr. 

Gomphonema acuminatum coronatum Ehr. 
G. Augur Ehr. 

G. capitatum Ralfs. 


263 


G. geminatum (Lyngb.) Ag. 
G. gracile Ehr. 

G. lanceolatum Kitz. 

G. olivaceum (Lyngb.) Kiitz. 
G. turgidum Ehr. 

G. Vibrio Ehr. 

Fyalodiscus stelliger Bail. 
fT, subtilis Bail. 


HyprosERA (TERPSINOE ?) NovAE-CAESAREAE Boyer n. sp. 

At first sight this species might appear to be a variety of Hy- 
arosera (Triceratium) trifoliata Cleve, but a close comparison will 
show that the two are distinct. 

Triangular, sides concave, angles broad at the base, equally ai- 
vided into three projections, surface sparsely punctate. In Cleve’s 
species the angles are cuneate and the middle lobe or tooth is nar- 
rower than the two outer, while the punctae are quite evident except 
at the center of the valve. There is a marked difference in gen- 
eral appearance, the New Jersey form being much less robust than 
that from New Zealand. It is quite common in the deposit and 
is of interest, as there is no form similar to it except that from 
New Zealand with the exception of two specimens noticed by 
Mr. Heinrich Reis in clay found near Cold Spring, Long Island, 
which discovery, however, was antedated by that made in the 
Wildwood material. I have not seen Mr. Reis’ specimens, which 
he identifies as Zriceratum trifoliatum and do not know, therefore, 
whether the Wildwood and Cold Spring forms are identical. 


Melosiva granulata (Ehr.) Ralfs. 
Navicula acrosphaeria Rab. 
N. affinis Ehr. 

NV. Americana Ehr. 

NV. bicapitata Lagerst. 

N. Bombus (Ehr.) Kiitz. 
NV. Brebissonit Kiitz. 

NV. columnaris Ehr. 

NV. Crabro (Ehr.) Kiitz. 

N. Dactylus Ehr. 

N. Dariana A. Schm. 

N. decurrens Kitz. 

NV. distans (W. Sm.) Ralfs. 
NV. elliptica Kiitz. 


264 


N. Fischeri A. Schm. 

N. formosa Greg. 

N. gibba Ebr. 

N. Hitcheocku Ehr. 

N. humerosa Breb. 

N. Iridis amphirhyncus Ehr. 
N. Kamorthensis Grun. 

N. latissima Greg. 

N. Lewisiana Grev. 

N. Liber linearis (Grun.) V. H. 
N. limosa Kitz. 

N. Lyra Ehr. 

NV. major Kiitz. 

N. mesolepta Ehr. 

LV. mesolepta nodosa Ehr. 
LV. mesostyla Ebr. 

N. nobilis (Ehr.) Kiitz. 

XN. pachyptera Kitz. 

NV. permagna (Bail.) Edw. 
NV. peregrina (Ehr.?) Kiitz. 
N. placentula (Ehr.) Kitz. 
NV. polyonca Breb. 

NV. pusilla W. Sm. 

NV. rhomboides Ehr. 

NV. sphaerophora Kiitz. 

N. Smithi Breb. 

NV. stauroptera parva Grun. 
NV. tabellaria Kitz. 

N. termitina Ehr. 

N. trinodis inflata Schultze. 
NV. viridis (Nitzsch) Kiitz. 
Nitzschia Campeachiana Grun. 
NV. cirumsuta (Bail.) Grun. 
NV. granulata Grun. 

NV. salinarum Grun. 

N. scataris (Ehr. ?) W. Sm. 
NV. Sigma (Kiitz), W. Sm. 
XV. tryblonella maxima Grun. 


265 


Odontidium mutabile genuinum Grun. 

Flagiogramma tessellatum Grev., rare. 

Pleurosigma eximium (Thw.) Grun. and Cl. 

P. Sctotoense Sulliv. 

P. Spencerit (Quek.) W. Sm. 

Polymyxus coronalis L. W. Bail. 

Pseudauliscus radiatus Bail. 

Pseudocunotia fiexuosa (Breb.) Grun. 

Rhaphoneis amplaceros Ehr. 

R. Belgica Grun. 

R. gemmifera Ehr. 

R. Rhombus Ehr. 

Stauronets acuta W. Sm. 

S. gracilis Ehr. 

S. Phoenicenteron Ehr. 

Surirella angusta Kiitz. 

S. diseriata (Ehr.) Breb. 

S. cruciata A. Schm. 

S. crumena Bréb. 

S. elegans Ehr. 

S. Febigerti Lewis. 

S. oblonga Ehr. 

S. ovalis Bréb. 

S. ovalis ovata (Kiitz) V. H. 

S. splendida Ehr. 

S. striatula Turp. 

S. fenera Greg. 
SURIRELLA WooLMANIANA Peticolas, n. sp. 

Mr. Peticolas, who discovered several specimens, remarks that 
it differs from S. Gemma, of which I had at first considered it a 
variety, in “ size, outline, areolation and the hyaline centre.” 

Valves broadly obovate, rounded at both ends ; costae slightly 
twisted, not reaching the median line, thus forming a pseudoraphe 
broadening at the centre. Costae and intercostate striae much 
more robust than in .S. Gemma. It somewhat approaches certain 
varieties of S. striatula. 

Synedra delicatissina W. Sm. 

S. investiens W. Sm. 

Terpsinoe Americana (Bail.) Ralfs. 


266 


Tetracyclus emarginatus (Ehr.) W. Sm. 
T. lacustris Ralfs. 

Tnceratium alternans Bail. 

T. favus Ehr. 

T. sculptum Shadb. 


Of the above about 80 are exclusively fresh water forms, 47 ex- 
clusively marine, while the others are found in fresh water and 
brackish or in brackish and marine. 


Preliminary Notes on Nelumbo lutea. 


By Benjy. HERITAGE. 
(PLATE 23%.) 


In August, 1890, being desirous of investigating the root 
growth of Nelumbo lutea, the late Isaac Burk, and self visited the 
mill-pond at Sharptown, Salem County, N. J., for that purpose. 

There were about eight acres thickly covered with the species, 
which presented a most gorgeous sight. 

We procured a boat, and after considerable research, we found 
a place at which the water was but a few inches in depth, over a 
very soft mud. He managed the boat, and seating ourself on its 
bottom, with an arm stripped to the shoulder we selected one of 
the large leaves which had a flower stalk in close proximity. 

These we traced with the hand, down into the ooze, to their 
common point of attachment, which brought the arm up to the 
elbow in the mud. There we found a very dense cluster of fibrous 
roots which were loosened, but this did not release the plant; as 
there was a horizontal portion, the size of a finger extending from 
it in opposite directions. The truth then dawned upon us that 
this majestic plant is stoloniferous! 

We traced this horizontal stolon or stem until we accidentally 
broke it, before reaching its termination in either direction. Its 
course was quite direct, and lay at a uniform depth of about one 
foot beneath the surface of the mud. 

At short distances we encountered very many other similar 
subterranean vines or stolons—portions of other plants perhaps 


267 


lying mostly in the same plane and crossing the pathway of our 
plant at various angles. 

Such as were above ours had to be severed in order to liberate 
the one we were in quest of. At varying distances we found 
other clusters of roots, from each of which a single, large, peltate. 
leaf rose about two feet above the surface of the water, while in 
the axil of several of them there was a flower stalk as tall. At 
many of the nodes we found a branch, each of which we traced 
until it, like the main stem, was accidentally broken, except in two 
instances, in which we secured the growing points. Upon finding 
our plants disconnected with the earth at all points, after about 
four hours’ assiduous labor, we carefully took it on board, rowed 
ashore, spread it out upon the grass, and with the miller’s “ten- 
foot pole” accurately measured it. The main stem was forty- 
seven feet long, and the combined length of the branches forty- 
three; in all ninety feet! It was to us a revelation, and exceed- 
ingly interesting, but it would have been even more so had we 
succeeded in unearthing the exézre plant which, as it lay upon the 
ground before us, proved itself to be an aquatic ve of gigantic 
proportions. 

The internodes were smooth, dull-white and of a uniform size 
throughout the main stem; they were about half an inch in diam- 
eter and were furnished with seven large air passages arranged in 
a circle equi-distant from a small central one and the epidermis. 
They were plentifully supplied with spiral tissue which could be 
drawn out a quarter of a yard before complete separation took 
place. These internodes varied in length from two feet, which 
was the shortest one in the main stem to five feet three inches, the 
longest in our specimen. ‘Those of’the branches were much 
shorter and their diameter less. Zong internodes insured the sep- 
aration of the leaves so that they could fully develop without 
coming in contact with each other, and perhaps the nature of the 
soil favored their growth, in confirmation of which we have seen 
Carex vestita Willd., growing in compact soil with stolons only a 
few inches long, whereas in an open porous soil they sometimes 
exceed two feet in length between the plants. 

The diameter of the nodes was considerably greater than that 
of the internodes and it was from these alone the roots originated. 


268 


Later examination has shown the roots to be arranged in six- 
usually-circular clusters placed side by side and extending com- 
pletely around the node, those growing from the upper side of 
the node also follow the general law of root growth, 7. ¢., descend. 

In those examined the number in each cluster ranged from 14 
to 21, averaging 110 roots at a node. 

The branches occurred regularly at several consecutive nodes, 
but irregularly at others. They sometimes extended to the right 
and to the left alternately like those of a Cucurbitaceous vine; at 
_ an estimated angle with the main stem of about 60° upon an 
average. 

During the course of our investigation, the queries arose 
whether those long internodes are perennial or survive but a 
single season? Whether the plantlet at each node has a perma- 
nent character or otherwise; and, if the former prove true, 
whether each node becomes the centre from which new growth 
radiates the following year? 

_ We contemplated a further inquiry the succeeding spring, hop- 
ing to solve those questions; but, during the interval, our friend 


was stricken with paralysis and intapacitated for additional re- . 


search, and the matter rested. 

In November last and again a month later, in company with 
Chas D. Lippincott, we visited the locality to inquire into the 
winter status of the plant, as well as to determine more fully its 
method of growth; and perchance to throw some light upon the 
above questions. The results thus far obtained are here sub- 
mitted. 

We secured many specimens containing buds, but in every 
instance they were at or near the end of a stem or branch, and 
consisted of one or generally two tuber-like enlargements of the 
stem following each other consecutively. The internodes of these 
thickened stems had failed to develop longitudinally more than a 
few inches, while transversely their diameter was greatly in- 
creased. The general structure of the stem was maintianed in the 
tubers, and the leaf buds were invariably found at their ends—the 
nodes—which had undergone but a slight modification. The tubers 
obtained varied from three to eleven inches in length, and very 
closely resembled a banana in appearance, both in regard to shape 


+ 


| 


269 


and proportions; except that the color was creamy-white, marked 
with purplish dots. They were generally somewhat flattened on 

the side from which the lateral bud grew at the next lower node, 
which might indicate that the growth was directed to the develop- 
ment of the bud at that point rather than into that portion of the 
tuber in line with and beyond it. 

Their texture is crisp, and it cuts with that peculiar grating 
which we hear and feel while cutting a raw potato with a thick 
knife; and which is doubtless owing, as Prof. Halsted has shown 
with relation to the winter buds of trees, to the presence of large 
quantities of starch stored during the growing season, for the rapid 
development of the embryo leaves in early spring, before they are 
capable of assimilating their nourishment. 

There are at least two kinds of buds shown at the tubers; 
First, lateral leaf buds, developed at the nodes; and secondly, ter- 
minal buds, which contain an embryo vine. They are not uniform 
in size; the largest of the former measured five and one-half 
inches in length, and consisted of a single petiole surmounted by 
an involute blade two inches in diameter; the whole completely 
enveloped by a whitish, brittle succulent sheath, the edges of 
which overlap. 

We obtained some specimens in which growth had extended 
just beyond the full capacity of the sheath, and it had been torn 
completely asunder below the blade, both portions having turned 
black. 

The severed part would doubtless have continued to envelop 
the blade, and it would have been carried up through the mud by 
the lengthening petiole—as the calyptra of Musci is borne up on 
the apex of the capsule—and when its office of protection shall 
have been fulfilled the unfolding leaf will cast it aside. 

_The terminal buds are shorter and thicker than the others, and 
include a portion of the future vine. An internode and node, to- 
gether with its young leaf, are all well formed; also a growing 
point beyond, and, like the leaf buds, they are protected by a 
similar sheath. 

_ We also obtained the black shrunken shells of old tubers, from 
which all the nutriment had been drawn, which, with contiguous 
portions of the vine, were lifeless. 


270 


The roots at the nodes, so far as observation has thus far ex- 
tended, all die in the fall; but we found much vitality still existing 
in the nodes. Portions of the internodes were discolored, had 
lost their rigidity, and were dead; such might have grown early 
in the season, and were the first to die. Other portions appeared 
full of life, and their cells, as well as those of the nodes, were filled 
with starch grains; but not one of the many nodes examined ex- 
hibited a bud, except as above stated, those at or near the ex- 
tremity of the vines. This absence of buds, and the dead roots, 
suggest the early dissolution of all parts of the plant except those 
last formed. Although other portions of this wonderful plant 
may survive the winter, yet if this prove true it is presumed that 
after growth re-commences in the buds of the tubers, the starch 
in the vines behind them will be utilized, after which they will 
probably die. Let investigation be directed to this, as well as to 
- other points of interest. 

The living buds are at the ends of the tubers. Back of the 
tubers we have. a long vine in which starch is stored, and which 
contains no buds, and show no signs of further growth. From 
this it appears that both the. vines and tubers after being fully de- 
veloped serve merely to store nutritive materials, and to transmit 
them to the growing points. 

We now behold this noble plant not only as a true aquatic, 
subterranean vine, but it is metamorphosed into a migratory vine. 
Why migratory ? Simply because the next year’s buds are so far 
removed from the location of those of the last. Just how far a 
plant removes its situation in a single season can only be deter- 
mined by locating one in early spring, tracing it throughout its 
entire length in the fall, and noting the position of the next year’s 
buds. Our specimen furnished ocular demonstration that this re- 
moval must have been more than forty-seven feet! Is this method 
of growth found in any other genus than Nelumbo? we ask 
for information. The annual character of the growth from each 
particular node is a most wise provision. If all those plantlets 
survived the winter, and became the centre from which new growth 
proceeded, there soon be an overcrowding ; and none would have 


room to develop naturally when deterioration and extinction would 
eventually ensue. 


271 


We are here furnished with another illustration, among many 
seen on every side around us, of unmistakable evidences of per- 
fected thought in their creation, each being peculiarly adapted to 
the position it holds in the divine economy. 

I am indebted to Dr. Ida A. Keller for the accompanying 
drawings and explanation of figures. 

MICKLETON, N, J., 1894. 


Explanation of Plate 23r. 


I, Diagram of vine of Nelumbo /utea Pers. 

St, St—main stem 47 feet long. 

X—unknown dist. to tubers with living buds—T. 

xy— «old tuber shells—O. T. 

II, End of stem ¥ of natural size. 
St.—unthickened stem. 
St/—two tubers terminating the stem. 
Sh.—remaining portions of sheaths (black). 
B—terminal bud which will develop into a stem. 
B/,—axillary bud (sheath which covered it removed). 
L. B/.—leaf bud, subtending terminal bud. It has a sheath of its own, which 
has not been removed, is distinct from the sheath marked Sh, The former is 
not black, the latter is. 
L. B’.—leaf bud, upper portion exposed by removal of sheath. 
R—points from which roots will emerge, these arranged in groups as indicated 
in the drawing. 
R/—dead roots showing their articulations. 
P—petiole of the last leaf of the season (now dead). 
III.—Cross section of vine, tiatural size. 

A—air passage. ° 
IV. Cross section of tuber, natural size (shows the flattening). 
vi id * involute leaf bud enlarged. 

A—air passage, 

VI. Starch grains. The largest represented (’) being .0492><.0328 mm. 


On the Carpels of Opulaster malyacea (Greene). 


This species was originally described by Prof. Greene in Pitt. 
2: 30, 1889, as Weillia malvacea, from specimens collected by the 
author himself at the north shore of Lake Pend d’Oreille, in 
Idaho. In the diagnosis of the species it is referred to a section 
of the genus characterized by indehiscent carpels. 


272 


Recently I had occasion to investigate the fruit characters of 
this Opulaster and find that the carpels do rupture along well-de- 
fined sutures, although it takes place tardily. The fruit of the 
species is usually 2-carpellary, very rarely 3-carpellary. Each carpel 
is from 1—3-ovuled, developing as many or fewer seeds; the usual 
number is I or 2. At maturity’the carpel is nearly triangular, 
much flattened, not at all inflated, and when well developed 
it measures about 4.5 mm. in length, and 3.5 mm. in width across 
the broadest portion. The seeds are oblong, somewhat com- 
pressed, nearly 2 mm. long and 1.3 mm. wide, grayish in color, 
with a firm polished testa and extremely bitter taste. The two 
carpels seen together bear a great deal of resemblance to the 
silicle of a Lepidium as Prof. Greene notes. 

The carpels are connate except at the divergent apices. They 
usually separate along a lateral line at maturity. Occasionally 
this line of demarcation does not form and they remain united. 
The pedicels commonly fall away from the axis of inflorescence 
in autumn or early winter by the development of a constriction at 
their bases, carrying the unopened fruit with them. The dehis- 
cence of the carpels does not usually occur until after they reach the 
ground, but if the pedicels are not detached from the axis of the 
corymb in the fall, which sometimes happens, the carpels will 
rupture in the spring while retaining their original position and 
enfolded by the persistent calyx. 

The dehiscence of the carpels takes place along two sutures, a 
ventral and a dorsal. In the majority of the cases that have come 
under my observation the inner or ventral suture opens first, the 
fissure beginning near the apex or at the point of juncture of the 
carpels, and is complete from base to summit. The outer or dorsal 
suture opens immediately after the rupture of the inner, but the 
line of dehiscence extends across the top of the carpel only. 
Owing to this circumstance the carpel is never two-valved in de- 
hisence, apparently a new character in our North American 
Opulasters, and therefore, clearly, to be considered as follicular. 
When the carpels are so firmly united that no lateral separation 
between them takes place the inner suture opens through both 
simultaneously. Joun B. LEIBERG. 

Hope, IDAHO, March gth, 1895. 


273 


Two new mountain Plants. 


By MERRITT LYNDON FFRNALD, 


“AsTER HENDERSONI n sp. 

Stem slender, 3 or 4 feet high, cinereous-pubescent, almost la- 
nate, except toward the glabrate base, branching above the mid- 
dle, the densely cienerous branches bearing single terminal heads, 
or themselves divided into naked or scarcely leafy branchlets; 
leaves thin, cinereous on both surfaces, especially on the mid-rib, 
or becoming glabrate above, the entire margins often ciliate; the 
upper cauline oblong or oblong-lanceolate, 2 or 3 inches long, 
with auricled clasping bases, the lower oblanceolate, conspicuously 
narrowed above the clasping bases, 4-6 inches long; leaves on 
the branchlets few, scarcely an inch long, often so few as to give 
the appearance of almost naked peduncles; heads large, an 
inch or two across, 4-6 lines high; involucre of two or three 
loose rows of cinereous linear-attenuate bracts, mostly herbaceous, 
but the inner sometimes scarious below, rarely with one two nar- 
row foliaceous bracts; the fifty or more blue rays %-3¢ inch 
long, a line wide. 

Collected by Prof. L. F. Henderson (No. 2798) in rich moist 
meadows along the St. Maries River, Kootenai county, Idaho, 
August 5, 1894. 

Closely related to puniceus and Cusichit. The involucre is like 
the former, but the plant is more slender, with no trace of the stiff 
pubescence and harsh serrate leaves ‘of that species, while the 
lower cauline leaves are contracted above the clasping bases as in 
Cusickii. Hendersoni has the same pubescence as Cusickii var. 
Lyalli Gray, but this latter plant has much broader leaves, and the 
outer bracts of the involucre are very broad and foliaceous, and, 
according to Prof. Henderson, it grows in more open and drier 


bottoms than the plant here described. 


CAREX SCABRATA X CRINITA n. hyb. 

Either stout or slender, the leaves and culm harsh as in sca- 
brata; spikes 5~7, mostly androgynous, varying from % to 1% 
inches in length, the upper nearly sessile, the lower on peduncles 
an inch or more long, slightly spreading; scales lanceolate or 
ovate-lanceolate, with brown scarious margins, and strong green 
mid-veins sometimes continued into rough awns 2-3 times as long 
as the perigynia, sometimes not equalling the perigynia; peri- 
gynia broadly ovate, with the few nerves either obscure or well 


. 


274 


marked, some of them puncticulate, others slightly puberulent; 
the beak short and entire. 


Collected by Dr. Geo. G. Kennedy in a damp hollow near the 
Crawford Bridle Path, on Mt. Clinton, N. H., August 20, 1891. 


Cryptogamic Notes from Long Island. III. 
By SMITH ELy JELLIFFE, M. D, 


DIATOMACEAE. 
Amphora ovalis Kutz. 
Cymbella Cistula Hempr. 
Encyonema ventricosum Kutz. 
Stauroneis acuta W. Sm. 
Stauronets anceps Ehrb. 
Stauroneis gracilis Ehrb. 
Stauronets Phoenicenteron Ehrb, 
Navicula Americana Ehrb, 
Navicula ambigua Ehrb. 
Navicula flamma A. Sch. var. 
Navicula cuspidata Kutz. 
Navicula dilatata Ehrb. 
Navicula firma Kutz, 
Navicula laevissima Kutz, 
Navicula gibba (Kutz.) Ehrb. 
Navicula Hitchcockti Ehrb, 
Navicula apis Kutz. 
Navicula lata Breb. 
Navicula Lyra Ehrb. 
Navicula major Kutz, 
Navicula marina Ralfs. 
Navicula radiosa Kutz. 
Navicula rhyncocephala Kutz. 
Navicula varians Greg. 
Navicula viridis Kutz. 
Pleurosigma angulatum W., Sm. 
Pleurosigma Balticum W. Sm. 
Pleurosigma elongatum W. Sm. 
Pleurosigma Spencerti W.Sm, 
Amphiprora ornata Bailey. 
Gomphonema acuminatum Ehrb. 
Gomphonema capitatum Ehrb. 
Gomphonema constrictum Ehrb. 
Rhotkosphenia curvata (Kutz.) Grun. 
Achnanthes brevipes Ag. 


275 


Achnanthes longipes Ag. 
Achnanthes subsessilis Erb. 
Cocconets Pedicularis Ehrb. 
Cocconets scutellum Ehrb. 
LEpithemia gibba (Ehrb.) Kutz. 
Epithemia turgida (Ehrb.) Kutz. 
Lunotit lunaris (Ehrb.) Grun. ‘ 
LEunotia tridentula Ehrb. 
Eunotia major (W. Sm.) Rab. 
Synedra affinis Kutz. 

Synedra fulgens (Kutz.) W.S. 
Synedra pulchella Kutz. 

Synedra longissima Kutz. 
Synedra lanceolata Kutz. 
Synedra Ulna (Nitzsch.) Ehrb. 
Fragilaria capucina Desmaz. 
Asterionella formosa Hass. 
Meridion circulare Ag. 

Tessela interrupta Ehrb. ? 
Tabellaria fenestrata Kutz. 
Tabellaria flocculosa (Roth.) Kutz. 
Grammatophora marina Kutz. 
Rhabdonema Adriaticum Kutz. 

_ Cymatopleura elliptica (Breb.) W. Sm. 
Cymatopleura Solea (Breb.) W. Sm. 
Surirella elegans Ehrb. 

Nitzschia fasciculata Grun. 
Niteschia gracilis Hautzsch. 
Nitzschia scalaris (Ehrb.) W. Sm. 
Niteschia sigmoidea (Ehrb.) W. Sm, 
Niteschia tabellaria Grun. 
Melosira Borrerti Grev. 

Melosira granulata (Ehrb.) Ralfs. 
Melosira varians Ag. 

Biddulphia laevis Erb. 
Biddulphia pulchella Gray. 
Rhizosolenia gracilis H. L. S. 
Triceratium alternans Bailey. 
Eupodiscus radiatus Bailey. 


Botanical Notes. 


Hough's American Woods. Part VI. of this valuable series of 
thin sections of North American woods, together with the 458 
press to accompany Parts IV. and V., have recently been 


276 


tributed to subscribers. The six parts now issued contain tangen- 
tial, radial and transverse slices of 150 pieces of trees, and form an 
nvaluable part of the equipment of any botanical museum. Cross 
sections of these woods mounted as lantern slides are also pre- 
pared by Mr. Hough. Descriptive matter and specimens of the 
work may be obtained from him by addressing R. B. Hough, 
Lowville, N. Y. 

Photographs of Fungi. We have received from Mr. C. G. 
Lloyd, of Cincinnati, the first set of a series of photographs and © 
photogravures of the larger fungi, prepared by him for distribu- 
_ tion among a limited number of students. The execution of these 
photographs is excellent and they cannot fail to be of great aid to 
mycologists. The species are determined by Mr. A. P. Morgan, 


of Preston, Ohio. Mr. Lloyd informs us that the sets are not for 
sale. 


Reviews, 


Guide to the Study of common Plants. An Introduction to Bo- 
tany. V.M. Spaulding. 2nd edition, pp. 287. Boston, D. C. 
Heath & Co. 1895. 

The second edition differs from the first mainly in the addition 
of a glossary and index of botanical terms, an index of plant 
names, and a chapter on fungi. 

The text has undergone some alterations and a number of ad- 
ditions have been made to the reviews and summaries at the ends 
of chapters. The chapter on Fruits has been completed by a few 
pages on “ The Organs of Flowerless Plants.” 

The chapters on ferns and club mosses have been simplified by 
the omission of the anatomy and developmental history, although 
the latter reappears in a review and summary of the Vascular 
Cryptogams. Many additions have been made to the “ special 


studies,” particularly in the portion of the book “devoted to the 
families of flowering plants. E. A. S: 


Ai preliminary List of the Mosses of Minnesota, John M. Hol- | 
zinger. (Minn. Bot. Studies, Bull. No. 9, March 5th, 1895). 

This list includes 156 species, arranged according to Renauld 
and Cardot’s list, with the addition of one new variety, Amblyodon 
dealbatus var. Americanus, R. & C. ined., which has been distrib- 


277 


uted as No. 180 of their Musci. Am. bor. Exsicc., with description 
and notes in their observations. These specimens should be com- 
pared with JZ, acount’ Aust. which has been referred to Amdélyo- 
dan dealbatus in the Manual. Gymnostomum curvirostrum var. 
scabrum is also an addition to the flora, not having previously 
been reported. We find several typographical errors, one of 
which is guilty of making a synomym for Coscinodon Raui. We 
have two sets of specimens from the Bluffs at Winona, and have 
not found C. Wright? among them, but C. Rau does occur, and 
the fact that M. Cardot has determined it as C. Renauldi, adds 
another point to my argument that the species are identical. 

The citation of authorities for Cynodontium Wahlenbergit 
(Brid.) Hartm. is incorrectly given, and Physcomitrium pyriforme 
is listed on Cardot’s determination. We have not yet seen an 
American specimen which agreed with this species, and the ones 
sent to us by M. Cardot bearing this name are not excepted. 
Physcomitrella patens (Hedw.) Br. & Sch. should be added as hav- 
ing been collected by Miss Sheldon at Fort Snelling, Minn., Octo- 
ber, 1894. BGS. 

On new Species of Cretaceous Plants from Vancouver [sland, 
J.W. Dawson. Trans. Roy. Soc Canada, sec. 4: 53-72. pl. 5-14. 
1893. 

In this paper are included descriptions of twenty-eight new 
species, some of which are, however, certainly referable to pre- 
viously described species. The figures are poor and some of the 
material upon which new species are founded is too fragmen- 
tary to be satisfactory. In deference, doubtless, to certain critics 
of palaeobotany the author finds it advisable to use these words: 
“T think it proper to say that I cannot be expected to pledge my- 
self for the accuracy of the generic names attached to mere leaves. 
When the fruit shall be found connected with them, they may re- 
quire very different reference. At present they merely stand as 
forms of certain types characteristic of a certain geological age, 
and admitting of more or less accurate comparison with modern 
plants,” which sentence he sums up quite concisely all that the 
palaeobotanists claims for his determinations. A chapter on the 
value of fossil plants as indices of climate in the past completes 
the paper. _ we. 1. 


278 


Field, Forest and Garden Botany, by Asa Gray, revised and ex- 
tended by L. H. Bailey. 

This once popular little manual, first published in 1869, had 
never been revised nor re-edited by its author, and, in conseqence, 
had, in time, lost much of its value and become somewhat obso- 
lete. Prof.C.R. Barnes began a much needed revision which was 
later taken up and has just been successfully carried through by 
Prof. Bailey. Considering the nature of the task it is doubtful 
whether anybody else was as well equipped for it. 

The reviser states that his first aim was to preserve, as far as 
possible, the method of the original, attempting “ nothing more 
than to bring it down to date.’ The work, however, has been 
more thorough and extensive than those words would imply ; not 
only were the nomenclature and definition corrected, but the an- 
alytical keys and grouping of species into sections have been re- 
arranged and much new matter added, so that the original of 374 
pages has been increased to 503 pages. The added matter con- 
sists of no less than 82 genera and 533 species, the total number 
of species being 3203, of which 1784 are indigenous and 1419 
extra-limital (in cultivation). To give a few examples taken at 
random: Clematis has now 17 described species instead of II in 
the original ; Magnolia, 8 species and hybrids instead of 3 ; Bras- 
sica, 8 instead of 5; Zilia, 6 instead of 3; Prunus, 29 instead of 19; 
Pyrus, 16 instead of 10; Canna, 11 instead of 5; Palms, 17 instead 
of 4, &c., and the quality has kept pace with the quantity. The 
typography is also very much improved, being even better than 
that of Gray’s Manual, and, indeed, about as perfect as one could 
desire. 

This book is intended to remain, as before, a companion to the 
Manual, and the nomenclature and definition are made to conform 
strictly with it; whatever shortcomings, therefore, the Manual is 
guilty of in these respects are shared by its companion work. A 
very welcome innovation is the citation of the authority after each 
name; this, in our day, has become a necessity in a work with any 
pretention to exactness. 

As would be expected, special attention was devoted to culti- 
vated plants, whether ornamental or useful, native or introduced ; 
in this respect the revision is remarkably complete, including even 


279 


such comparatively recent introductions as Akedia guinata, Acti- 
nidia polygama, Solanum muricatum, etc. It is not only the best, 
but, I may say, the only work descriptive of our domesticated 
flora. It will be heartily welcome by the many who have neither 
time nor facilities for the pure study of systematic botany, but 
who, nevertheless, love flowers and takes an intelligent interest in 
the wealth of decorative and useful plants which surround them on 
on ail sides. VA, 


Phlanzenkrankheiten durch kryptogamen Parasiten verursacht 
Karl Freiherr von Tubeuf. Berlin, 1895. 

This book is a condensed treatise on plant “parasitism” or 
“symbiosis” in its widest sense. Unfortunately the term plant- 
disease (Pflanzenkrankheit) is not defined, but the reader is led to 
assume that any change produced in a plant due to a “symbiotic” 
or “parasitic” relation with another plant is considered a plant 
disease. 
The attempt is made to treat the physiological and morpho- 
logical relations of parasitism from a comparative standpoint. 
Physiological and anatomical details are not entered into as that 
would be both impossible and unnecessary because of the copious 
citations of special authors whom the specialist interested may 
consult. All the authorites quoted are recent and standard. With 
few exceptions the figures are excellent. The numerous photo- 
graphs are especially interesting. 

The student of general botany will no doubt find Part I. the 
most interesting. Here the author has very briefly outlined his 
conception of parasitism in the wider sense. Here he also shows 
his greatest originality. Especially interesting are his distinctions 
between Mutualism (“Mutualismus”) and Nutricism (‘ Nutri- 
cismus”). By the former is meant that form of parasitism in which 
the two symbionts mutually benefit each other. As the most im- 
portant examples are to be mentioned the lichens. Here two or- 
ganisms, an alga and a fungus live together for mutual benefit. 
He emphasizes the fact that this symbiosis has so changed the 
life history of the two organisms that they combine to form a mew 
organism. This union of two originally distinct organisms to 
form an independent organism he designates “individualism.” By 
nutricism is meant that form of parasitism in which one of the 


280 


symbionts is not benefited because no assistance is required. As 
an example may be mentioned the mycorhiza of certain trees and 
the mycodomatiae (rhizobia) of Leguminosae. Here the author 
supposes that the infecting fungus works in the interest of the 
plant infected without receiving any benefit in return. 


In Part I. are also given various approved methods and means 
for guarding against infection of pernicious parasites and how to 
to destroy them after infection has taken place. 


Part II., which is by far the most extensive, treats of “ phyto- 
pathogenic fungi and algae.” It is essentially a systematic de- 
scription of the infecting fungi and algae based upon the most re- 
cent investigations and conclusions. 


Taken all in all, this work is certainly a most valuable contri- 
bution to the science of botany. It will prove of great benefit to 
all botanists. The condensed retrospect of the subject will suffice 
for the general student, while the citations will be found very useful 
by the special student. 

Language and style are scientifically clear and simple. Only a 
a few of the figures are borrowed. ALBERT SCHNEIDER. 


A Monograph of the North American Species of the Genus Poly- 
gonum, by John Kunkel Small. This work, a folio of 183 pages 
(not counting the plates), was issued on the 23d of April, 1895, as 


the first volume of Memoirs from the Department of Botany of 
Columbia College. 


In the introduction, after discussing the position of the family 

to which it has given a name, the relation of the genus Polygonum 
to its allies is taken up and considered. Then follow chapters on 
the geographical distribution of the species, the general morphol- 
ogy of the genus, the anatomy of the stem and general anatomy. 


Natural and distinct as the genus appears, it admits of division 
into well-marked sub-genera, nine of which contain representa- 
tives in the New World, viz.: Bistorta, Aconogonon, Persicaria, 
Amblyogon, Tovaria, Avicularia, Duravia, Tiniaria and Echino- 
caulon. These, as described and figured by Dr. Small, including 
a fossil species from the Miocene Tertiary of Colorado, number 71. 
Each of them is illustrated by an uncolored outline-drawing on a 
separate sheet, so admirably executed as to make the salient charac- 


281 


ters clear to the eye, and the descriptions on the opposite pages 
are models for their terse and accurate use of botanical termi- 
nology. 

The new species, 9 in all and the varieties 8, form a large 
addition to the genus. 

The latter part of the volume is devoted to the comparative 
anatomy of the stems of certain species, accompanied by 15 ex- 
cellent plates, which exhibit such diversities of internal structure 
as to afford essential aid in support of sub-generic and even specific 
distinctions. 

To withhold recognition, so fairly earned and so worthy of 
acknowledgment, from a work like this would be wrong. It is 
highly creditable both to the author and to the college whose 
ample resources and friendly help enabled him to produce it. 
The fruit, not of a hasty and rapid incursion, but of persistent, 
patient and thorough study of the field which it covers, the con- 
tribution it has made to science is solid and will endure. What 
Dr. Engelmann did for /uxcus has now been done for Polygonum, 
and more of just this kind of labor is needed for the better eluci- 
dation of our North American Flora. eG. P: 


Proceedings of the Club. 


Turspay Eventnc, May 14TH, 1895. 


The President in the chair and 22 persons present. 

Mr. A. Ruth and Mr. W. A. Bastedo were elected active 
members. 

The editor announced the publication of the third part of Vol. 
4 of the Memorrs, and stated that the fourth part would contain a 
paper by Mrs. Pettit on Arachis hypogaea. 

Mr. Lighthipe reported excellent progress in the rearrange- 
ment of the herbarium. 

The announced papers of the evening were then presented. 

1. By Mr. George V. Nash, “‘ The North American Species of 
the Genus Cenchrus,” illustrated by specimens. 

2. By Mrs. Anna S. Pettit, « Notes from a Study of the Pea- 
nut, Arachis hypogaea.” 


282 


Dr. N. L. Britton exhibited a series of lantern-slides of cross- 
sections of North American Woods, prepared by Mr. R. B. Hough. 


WEDNESDAY EVENING, May 29TH, 1895. 


Vice-President Lighthipe in the chair and 32 persons present. 

Miss Rose T. Roux, Mr. James A. Graves and Mr. Frederick 
King Vreeland were elected active members. 

Mr. Barnhart reported the occurrence of Azalea lutea L. (A. 
calendulacea Michx.) in an apparently native state near Tarrytown, 
N. Y., and remarked on the distribution of this spccies, which is 
not recorded in recent writings from further northeast than the 
mountains of Pennsylvania. Dr. Britton called the attention of 
the Club to the recorded occurrence of a yellow-flowered Azalea 
in southern New York by Governor Cadwallader Colden about 
the middle of the last century, and that Colden’s account of it had 
been made a part of the citations by Linnaeus of his Azalea lutea 
in 1753. Linnaeus later abandoned the name /ztea on account of 
having received specimens of the pink-flowered species which he 
described as A. nudifiora in 1762, supposing them to be the same. 
Mr. Barnhart’s discovery is an interesting confirmation of the posi- 
tion taken in the recently issued “List of Northeastern North 
American Plants” that the name A. /utea L. must sekgeeeas A. calen- 
dulacea Michx. 

Mr. Barnhart also reported that A/iaria Alliaria, found by him 
last year at Hastings, N. Y., had this year spread over a consider- 
able area and was well established. 

The announced paper of the evening was read by Mr. A. A. 
Tyler on ‘Stipules in the Family Rosaceae,” illustrated by speci- 
mens and drawings. It was discussed by Dr. Schneider and Dr. 
Britton. 


The Club then adjourned to the second Tuesday evening in 
October. 


283 


Index to recent Literature relating to American Botany, 


Arthur, J. C. Joseph Schroeter. Bot. Gaz. 20: 230-232. 20 My. 
1895. 


Atkinson, G. F. Intelligence manifested by the Swarm Spores of 
khisophidium globosum. Bot. Gaz. 19: 503, 504. 1894. 


Atkinson, G. F. On the Swarm Spores of Pythium and Ceratiomyxa. 
(Abstract.) Proc. Am. Assn. Adv. Sci. 43: 290, 291. Mr. 1895. 


Atkinson, G. F. Relations between the Functions of the Vegetative 
and Reproductive Leaves of Onoclea. (Abstract.) Proc. Am. Assn. 
Adv. Sci. 43: 290. Mr. 1895. 


Atkinson, G. F. The Exoascaceae of Stone Fruits. Gard. & For. 7: 
463, 464. 1894. 

Atkinson, G. F. The Extent of the Annulus and the Function of the 
different parts of the Sporangium of Ferns in the Dispersion of 
Spores. Int. Journ. Micros. and Nat. Sci. Ap. 1894. (Reprinted 
from Bull. Torr. Bot. Club, N. £893). 

Barnes, C. R. Vitality of Warsitia quadrifolia. Bot. Gaz. 20; 229. 
20 My. 1895. 

Bastin, E.S. Structure of Sassafras. Am. Journ. Pharm. 67: 312- 
318. 7. 7g. Je. 1895. 

Benedict, A. L. Tabular Review of Organography prepared for the 
Use of Classes in Botany. Pamph. pp. 21. [Buffalo] 1895. 

Bessey, C. E. Botany of the Plums and Cherries. Ann. Rept. Ne- 
braska State Hort. Soc. 1895: 163-178. 1895. 

Descriptions and illustrations of various species of Prunus, mostly North Ameri- 
can, 

Bessey, C. E. The Botany of the Grape. Ann. Rept. Nebraska 
State Hort. Soc. 1895: 7-26. f. 72. 1895. 

Descriptions and figures of various species of Vitis, mostly North American. 

Bicknell, E. P. Hypericum boreale (Britton) and related Species. 
Bull. Torr. Bot. Club, 22: 211-215. 15 My. 1895. 

Blight, R. The Origin of the Orange. Rural Californian, 18: 224- 
226. My. 1895. (Reprinted from N. Y. Evening Post.) 

Historical resumé, 

Britton, E. G. A. Hybrid Among the Mosses. (Abstract.) Proc. 

Am. Assn. Adv. Sci. 43: 292. Mr. 1895. 


284 


Britton, N. L. New and noteworthy North American Phanerogams.— 
IX. Bull. Torr. Bot. Club, 22: 220-225. 15 My. 1895. 
New species in Carex, Allionia and Ranunculus, 
_ Britton, N. L. Notes on the Primary Foliage and the Leaf-scars of 
Pinus rigida. (Abstract.) Proc. Am. Assn, Ady. Sci. 43: 293- 
Mr. 1895. 


Budd, E. M. Laws of Floral Colors. Rep. Iowa State Hort. Soc. 
28: 53-56. 1894. 

Bush, B. F. Notes on the Mound Flora of Atchison county, Mis- 
souri. Ann. Rep. Mo. Bot. Gard. 6: 121-134. 1895. 


Card, F. W. Two wild Vegetables of Merit. Gard. & For. 8: 223. 
5 Je. 1895. 
The two plants in question are Lactuca Canadensis and Astragalus crassicarpus. 

Chamberlain, C. J. The Embryo-sac of Aster Novae-Angliae. Bot. 
Gaz. 20: 205-212. p/. 15, 16. 20 My. 1895. 

Chalmot, G. de. The Influence of Nitrates on Germinating Seeds. 
Agric. Sci. 8: 463-465. Mr. 1895. 

Cook, A. J. Parthenogenesis among Plants. Rural Californian, 18: 
237-238. My. 1895. 
Discusses effect of pollen on development of fruits. 

Coulter, J. M. Some Affinities Among Cactaceae. (Abstract.) Proc. 
Am. Assn. Ady. Sci. 43: 286-287. Mr. 1895. 


Crozier, A. A. What is Common Millet? Agric. Sci. 8: 449. Mr. 
1895. 


Davenport, G.E. <Asfidium simulatum Dav. Bot. Gaz. 20: 229, 
230. 20 My. 1895. 


Dietel, P. New North American Uredineae. Erythea, 3: 77-82. 1 
My. 1895. 
New species in Aecidium, Uromyces and Pu:cinia, 


Franceschi, F. Agaves in Southern California. Gard. & For. 8: 
228. 5 Je. 1895. 


Galloway, B. T. Some Factors influencing the Health of Plants 
under Glass. The Florists’ Exchange, 468-469. Ap. 1895. 


Ganong, W.F. Present Problems in the Anatomy, Morphology and 
Biology of the Cactaceae. Bot. Gaz. 20: 129-138. 20 Ap. 1895: 
213-221. 20 My. 1895. 


Greene, E.L. Corrections in Nomenclature.—VII. Erythea, 3: 75, 
76. 1 My. 1895. 


285 


Greene, E. L. Novitates occidentales.—XIII. Erythea, 3: 69-73. 
t My. 1895. 

New species in Ranunculus, Braya, Erysimum, Ribes, Mentzelia, Dodecatheon, 

Solanum, Mimulus and Crepis. 

Greene, E. L. Problems in modern Botany. Bull. Catholic Univ. 
I: 200-212. Ap. 1895. 

Halsted, B. D. Notes upon a Root Rot of Beets. (Abstract.) Proc. 
Am: Assn. Adv. Sci. 43: 293. Mr. 1895. 

Halsted, B. D. Notes upon Chalara paradoxa. (Abstract.) Proc. 
Am. Assn. Adv. Sci. 43: 293. Mr. 1895. 

Halsted, B. D. Notes upon Poisonous Plants. Gard. & For. 8: 1 
My. 1895. 

Hariot, M. P. Algues du Golfe de California. Journ. de Bot. g: 167- 
170. 1 My. 1895. 

New species are described in Lithothamnion and Lyngbya. 

Havard, V. Further Remarks on Family Nomenclature. Bull. Torr. 
Bot. Club, 22: 216-219. 15 My. 1895. 

Havard, V. When is Rhus Zoxicodendron most active? Gard. & 
For. 8: 203. 22 My. 1895. 

- Hollick, A. A new fossil Zzriodendron from the Laramie at Walsen- 
berg, Colo. and its Significance. (Abstract.) Proc. Am. Assn. Adv. 
Sci. 43: 225. Mr. 1895. 

Hollick, A. Descriptions of new Leaves from the Cretaceous (Dakota 
Group) of Kansas. Bull. Torr. Bot. Club, 22: 225-228. pl. 236, 
237. 15 My. 1895. 

Hooker, J. D. <Argylia canescens. Curt. Bot. Mag. 51: pl. 7474. 
My. 1895. 

A native of Chili. 

Hooker, J. D. Vaccinium erythrocarpum. Curt. Bot. Mag. 51: fv. 
7413. My. 1895. 

Hue, l’Abbé. Lichens de Californie. Journ. de Bot. g: 108-113. 16 
Mr. 1895. 

Enumeration of a small collection made by M. Diguet in 1894 in the Sierra de la 

Laguna at an altitude of 1800 m. 

Jack, J. G. Some unusual androgynous Flower-clusters. Gard. & 
For. 8: 222. f. 33. 5 Je. 1895. 

Keener, A. E. -Collinsia bicolor. Bot. Gaz.20: 232. 20 My. 1895. 

Keffer, C. A. Bull Pine in the West. Gard. & For. 8: 163. 24 


Ap. 1895. 


286 


Keffer, C. A. Conifers in the West. Gard. & For. 8: 182. 8 My. 
1895. 

Lloyd, F.E. A new Violet. Ervthea, 3: 74. 1 My. 1895. 
Viola Macloskeyi from Oregon. 

- Maxwell, C. F. Some morphological Relationships of the Cactaceae. 
Trans. Texas Acad. Sci. 1: 29-31. f. 7-5. 1895. 

Meehan, T. <Amorpha canescens, Meehans’ Month. 5: 101-102. 
Je. 1895. 

Newcombe, F. C. Regulatory Growth of Mechanical Tissue. (Ab- 
stract.) Proc. Am. Assn. Adv. Sci. 43: 287, 288. Mr. 1895. 

Nicholson, G. The Magnolias. Gardn. Chron. 17: 515, 516. 27 
Ap. 1895. 

Pammel, L. H. Diseases of Foliage and Fruit. Rep. Iowa State 
Hort. Soc. 28: 467-474. 1894. 

Pammel, L..H. Diseases of the Cherry. Rep. Iowa State Hort. 
Soc. 28: 248-253. 1894. 

Pammel, L. H. Is Parsnip Poisonous? Gard. & For. 8: 228. 5 
Je. 1895. 


Pammel, L. H. Results obtained in crossing Cucurbits. Rep. lowa 
State Hort. Soc. 28: 320-322. 1894. 


Patterson, Mrs. F. W. Species of Zaphrina parasitic on Populus. 
(Abstract.) Proc. Am. Assn. Ady. Sci. 43: 293, 294. Mr. 1895- 


Peck, C. H. New Species of Fungi. Bull. Torr. Bot. Club, 22: 

198-211. 15 My. 1895. 

New species in Lefiota, Clitocybe, Collybia, Mycena, Omphalia, Hygrophorus, 
Entoloma, Leptonia, Eccilia, Pholiota, Hebeloma, Flammula, Tubaria, Pluteolus, 
Cortinarius, Agaricus, Stropharia, Hypholoma, Panaeolus, Coprinus, Boletinus, 
Boletus, Polyporus, Sparassis, Battarrea, Tylostoma, Lycoperdon, Excipulina, 
Melasmia, Caeoma, Aspergillus, Leptoglossum and Vaisa. 


Philippi, R. A. Plantas nuevas Chilenas. Ann. Univ. Chil. 54-57: 
1893-1894. 
Enumeration and description of a large number of new species. 

Pollard, C.L. The genus Zenobia Don. Bull. Torr. Bot. Club, 22: 
231, 232. 15 My. 1895. 

Rolfe, R. A. New Orchids. Kew Bull. 98. F. 1895. 
Three new species are described from South America. : 

Rose, J. W. A blue Water-Lily from Mexico. Gard. & For. 8: 205- 
jf. 31. 22 My. 1895. 
Illustrating a species supposed to be Castalia elegans (Hook.) Greene. 


287 


Rothrock, J. T. The Beech (Fagus ferruginea Ait.). Forest Leaves 
5? 40-41. Je. 1895, 

With two illustrations. 

Sargent, C.S. An Arizona Agave. Gard. & For. 8: 184. f. 28. 8 
My. 1894. 

Mlustration of Agave Huachucacensis. 

Sargent, C. S., Editor. Rhododendron Vaseyt. Gard. & For. 8: 
214. 29 My. 1895. 

Sargent, C.S., Editor. Sanguinaria Canadensis. Gard. & For. 8: 
214. f. 92. 29 My. 1895. 

Schneider, A. The biological Status of Lichens. Bull. Torr. Bot. 
Club, 22: 189-198. 15 My. 1895. _ 

Smith, E. F. The Watermelon Disease of the South. (Abstract. ) 
Proc. Am. Assn. Ady. Sci. 43: 289, 290. Mr. 1895. 

Smith, J. G. Notes and Observations on new or little known 
Species. Am. Rep. Mo. Gard. 6: 113-119. f/. 48-56. 25 Ap. 
1895. 

_ New species are described in Sedum, Zephyranthes, Sagittaria and Eragrostis. 

Smith, J. G. Revision of the North American Species of Sag7tfaria 
and Lophotocarpus. Am. Rep. Mo. Bot. Gard. 6: 27-64. pl. 7-29. 
24. My. 1894. 

Sprenger, C. Beitrag zur Kentnis der Agave Americana. Monats. 
Kakteenk. 5: 51. 20 Ap. 1895. 

Stephani, F. Hepaticarum species novae.—VII. (Schluss). Hed- 
wigia, 34: 49-65. 8 Ap. 1895. 


Describes new species in /sotachis, Fungermania, Kantia, Archilejeunea and 
Brachiolejeunea mostly from South America. 


Stone, G. E. Plant Diseases. Rep. Mass. Agric. Coll. 1894. [reprint.] 


Sturgis, W. C. Report of Mycologist. Rep. Conn. Agr. Exp. Sta. 
17: 72-111. f/. 2. 1894. 
Treats of apple scab, grape mildew, quince rot, celery bli ght, tobacco rots, etc., with 

remedies, 

Thomas, M. B., and Dudley, W. R. A Laboratory Manual of Plant 
Histology. 8vo. pp. 300. Crawfordsville, 1894. 

Tight, W. G. Notes from the Botanical Laboratory. Bull. Sci. Lab. 
Denison Univ. 8: part 2, 7-12. f. 7-9. F. 1894. 
Notes on Phallus Daemonum and on some floral monstrosities. 

Toumey, J. W. £chinocactus Wislizent and some related Species. 
Gard. & For. 8: 154. f. 2g. 1 My. 1895. 


288 


Tracy, S. M., and Earle, F. S. New Species of parasitic Fungi. 

Bull. Torr. Bot. Club, 22: 174-179. 18 Ap. 1895. 

New species in Puccinia, Ustilago, Dimerosporium, Asteridium, Laestadia, 
Sphaerella, Lembosia, Vermicularia, Diplodia, Hendersonia, Pestalozsia, Scoleco- 
trichum, Cercospora and Tetraploa, mostly from Mississippi. 

Uline, E. B., and Bray, W.L. Synopsis of North American Ama- 

ranthaceae.—II. Bot. Gaz. 20: 155-161. 20 Ap. 1895. 

The genera Acnida and Gomphrena. Describes A. tamariscina prostrata as 
new. 

Underwood, L. M. The Evolution of the Hepaticae. Proc. Am. 

Assn. Adv. Sci. 43: 259-274. J/lust. Mr. 1895. 


Vail, A. M. A preliminary List of the North American Species of 
Malpighiaceae and Zygophyllaceae. Bull. Torr. Bot. Club, 22: 228- 
231. 15 My. 1895. 

Establishes the genus Covz//ea in place of Larrea Cay. (nom. praeoc). 

Webber, H. J. Studies on the Dissemination and Leaf Reflexion of 
Yucca aloifolia and other Species. Ann. Rep. Mo. Bot. Gard. 6: 
QI-112. pt. 45-47. 1895. 

Weed, C. M. The Cultivation of Specimens for Biological Study. 
Pamph. pp. 22.7. 77. 1895. 

Wilder, Burt G. Evidence as to the former Existence of large Trees 
on Nantucket Island. Proc. Am. Assn. Ady. Sci. 43: 294. Mr. 1895. 


Zenetti, P. Das Leitungs system im Stamm von Osmunda regalis L. 
und dessen Uebergang in den Blattstiel. Bot. Zeit. 53: 53-78, A/. 2. 
16 Ap. 1895. 


BULLETIN OF THE TORREY al CLUB. aa, 238 


Lari Ths bs : 4 
Cie 


NELUMBO LUTEA. 


Contributions from the Herbarium of Columbia College. 


No. 4. 
No. 5. 
No. 6, 


No. 7. 
No. 9. 


[The numbers omitted from this list are out of print. J 
A List of Plants Collected by Miss Mary B. Croft at San Diego, Texas. By 


N. L, Britton and H. H. Rusby (1887), . 2... 0), ۤ cents. 
New or Noteworthy North American Phanerogams. By N. L. Britton 
CERES): ery oe ee a es eee _ , 25 cents, 


An Enumeration of the Plants Collected by Dr. H. H. Rusby in South 
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The Genus /icorta of Rafinesque. By N. L. Britton (1888),  . 25 cents. 

A List of Plants Collected by Dr. E. A. Mearns at Fort Verde and in the 
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Preliminary Notes on the North American Species of the Genus Zissa, 
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A Descriptive List of Species of the Genus Heuchera. By Wm. E. Wheel- 
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A Preliminary List of the Species of the Genus A/eibomia occurring in the 
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TORREY BOTANICAL CLUB. 
Vol. 22. Lancaster, Pa, July 31, onic ‘ ‘idan ts v. 


Some new and rare Desmids of the United States.—II, 


By L. N. JoHNsON. 
(PLATES 232, 233.) 


Since the publication of the first paper of this series! I have 
examined a number of collections of desmid-bearing material 
from various parts of the country. The results of this examina- 
tion have been to extend very greatly the known range of many 
species, and to add more than a score to the number reported 
from the United States by previous writers. Many species are 
still undetermined and held for further study. Material is also 
being accumulated which will illustrate the great variability of 
many species, but this subject will be discussed at another time. 

The present paper is based on collections made in New Hamp- 
shire, Connecticut, New York, North Carolina, Florida, Louisiana, 
Michigan, Kansas and California. From some of these States 
Material representing several localities has been examined. I 
wish to acknowledge my indebtedness to the friends who have 
responded so kindly to my request for material, and to Prof. Nord- 
stedt, who has aided me greatly by his criticism and advice. 

I have thought it best to make one change in the manner of 
reporting additions to our flora. After every such species is given 
a number in parenthesis. This refers to a numbered specimen in 

my herbarium. Thus if any species is incorrectly determined it 


1 Bulletin of Torrey Botanical Club, 21: 285. Je. 1894. 


290 


will be possible to see the particular specimen on which the re- 
port was based. 

The limits of genera among desmids are so unsettled that 
hardly any two writers agree in all points. Especially is this true 
of the genus Cosmarium. No attempt has been made to settle 
the points in dispute. In nearly every case the specimens have 
been compared with the original figuresand descriptions. Where 
the variation from the type is slight varietal names have not been 
used, as it seems useless to encumber our literature with a long 
list of varieties, many of which represent simply forms of a vari- 
able species. 

In the enumeration of species only the more noteworthy ones 
from the various localities are mentioned, although careful record 
has been made of ail species observed, and specimens have been 
preserved. Species new to the United Statcs are marked with an 
asterisk. 

Pleurotaenium nodosum (Bail.) Lund. Near Tallahassee, Fla. 
Scarce. Surface of cell smooth; ends ornamented with a circle 
of conical-rounded granules. 

Docidium verrucosum (Bail.) Ralfs. Frequent in various col- 
lections from Florida. Walle’s figures do not give a correct idea 
of this beautiful species. The elevations are nearly square, except 
those of the terminal circle, which are elongated. The apex of 
the semi-cell bears a crown of rounded granules. The Florida 
specimens resemble the figures of Joshua’s D. tesse//atum*, and it 
seems to me that the latter must be regarded as a synonym of D. 
verrucosum, 

D. coronulatum Griin. Occasional at Meredith, N. H. Sev- 
eral cells were often found joined in series. Diam=—=25-35 ps 

This is apparently the same species figured by Wolle under 
this name. Griinow’s original measurements are larger, and the 
cells as figured by him were slightly different in shape. 

Penium annulare West. Baton Rouge, La. Frequent. Found 
by W. West in material from Maine, and described by him’. The 


2 Joshua, W. Burmese Desmidieae, with descriptions of new species occurring in 
the neighborhood of Rangoon. Journ. Linn. Soc. 21 : 650. pl, 25. f. 15. Je. 1886. 

3 West, W. The Freshwater Algae of Maine. Repr. from Journal of Botany 
1894: (2) pl. 775. f. 5.6. D. 1891. 


291 


present discovery extends its range more than a thousand miles. 
My specimens agree with his description, except that the undula- 
tions are less numerous and the cells rather shorter. Diam.=20 p; 
length=110 ». Fig. 1. 

* P. Cylindrus (Ehrb.) Breb. var. Silesiacum Kirch. (532). Wake 
Forest, N.C. Scarce. Agrees well with the description. Diam. 
=14 #; length=42 p. Fig. 2. 

Closterium maculatum Hast. Bridgeport,Conn. Rare. Nearly 
twice as large as Hastings’ measurements, but otherwise agreeing 
with his description. Striae of minute granules and interrupted 
irregularly. Diam.=50 yp. Prof. Nordstedt suggests that this 
Species may possibly be a form of CZ. arcolatum Wood. Fig. 7. 

Cl. angustatum Kiitz., var. clavatum Hast. Bridgeport, Conn. 
Scarce. Diam.=25 ». This seems well marked, but only a few 
Specimens were found. Fig. 6. 

* Cl. abruptum West. (395). Holderness, N. H. Frequent. ° 
Variable in length. Occasionally a specimen is longer than 
West’s measurements. Diam=12 »; length=84-200 p. Fig. 5. 

* Cl. Cynthia De Not. (537). Wake Forest, N. C. Frequent 
and typical. Striae very fine and numerous. Diam =12-14 p; 
length=100-120 p. Fig. 3. 

Cl. tumidum nov. nom. (416). (C2. Cornu 3 of Ralfs.t; Cl. 
cornu of Wille’, but not of Ehrenberg®). Burlington, Kansas. Fre- 
quent. Diam. usually 12 »; length= 116-120 p#. Occasionally a 
specimen reaches a diameter of 18-20 yp, representing Wille’s 
forma major®, West found the latter in Maine. Fig. 4. 

Gonatozygon aculeatum Hast. Meredith, N. H. Occasional. 
Seems to be a well-marked species. Diam.=16 yp; length 300 p. 
Aculeae 6-8y long. Fig. 9. 

G. asperum (Breb.) Cleve. (460). Tallahassee, Fla. Rare. West 
reports it from Maine. The single specimen found was about 
twice as long as the type, but otherwise the same. Diam.—6 p; 
length=260 p». Fig. 8. 


4 Ralfs, J. British Desmids. 176° p/. 30. f 64 and é, 1848. : : 

5 Wille,N. Ferskvandsalger fra Novaja Semlya. Ofversigt af Kongl. Veten- 
skaps. Akademiens Forhandlingar, 1879, //. 24. f, 80,87. 1879. 

6 Ehrenberg, C. G. Die Infusionsthierchen als volkommene Organismen. 94. 


B16. f. 5. 1838. 


292 


Micrasterias Nordstedtiana Wolle. Meredith, N. H. Rare. 
Diam —170 p. These specimens differ from Wolle’s description 
and figures in that the lobes terminate in two spines instead of 
three or more. The end lobe is not so emarginate, and the projec- 
tion on the sides of the neck is larger and nearly cylindrical, end- 
ing in two spines. Fig. 14. 

M. Mahabuleshwarsensis Hobs. Meredith,N. H. Rare. In 
same material as the preceding. Rays more slender than in 
Wolle’s figure. 

* Euastrum Sibiricum Boldt (5€8). Duval county, Florida. 
Agrees well with Boldt’s description and figuyes. Diam=18y5 
length=2on; thick.=11y. It is interesting to find this Siberian 
species in Florida. Probably it will be found to have a wide dis- 
tribution, escaping notice because of its small size. Fig. 13. 

Dysphinctium viride (Corda) De Toni. (Cosmarium Cordanum 
(Breb.) Wolle.; Desmids of U.S. 1892 edition). Reported from 
Nova Scotia, and by West from Massachusetts. I find it common 
in material from Bridgeport, Conn., and Staten Island and Ithaca,. 
N.Y. Diam.26p. 

Cosmarium amoenum Breb. Frequent in collections from Flo- 
rida, and from Meredith, N. H. Typical. 

C. cyclicum Lund. Meredith, N. H. The specimens come 
close to var. sudtruncatum Hansg. 

C. elegantissimum Lund. Duval county, Fla., and Meredith, N. 
H. The specimens resemble Lundell’s figure 7, though the gran- 
ules are most commonly only faintly emarginate. Wolle’s figures 
are singularly incorrect in showing a deep linear sinus, though he 
describes it correctly in this particular. Usually about seven 
longitudinal rows of granules are visible at onee in front view- 
Diam.=22y; length=soy. Fig. 27. 

C. Minneapolitanum Hansg.(C. protuberans Lund. var. granite 
latum WWolle) (489). Ithaca, N. Y. Rather larger than Wolle’s 
specimens (diam.= 34-36 2), but similar in form. 

C. margaritatum (Lund.) Roy and Bissett (388). Meredith, N. 
H. Rare. The drawing does not show the granules well. They 
should be much larger and closer together. The position of the 


7 Lundell, P.M. De Desmidiaceis quae in Suaecia inventae sunt. 53- p13 f- 
20, 3571, 


293 


hyaline punctations is difficult to make out. They seem to be be- 
tween the granules in the longitudinal rows. Diam.=72 y; length 
=90n. Fig. 32. 

C. pseudoprotuberans Kirch. New Baltimore, Mich. Rare. 
Typical. Diam=32y; length=36; isth—1op. West re- 
ports this from Maine. Fig. 24. 

* C. costatum Nordst. (522.) Wake Forest, N.C. Scarce. 
Diam.= 32 #; length=42 ; thick.=21 y; isth—=12y. This bears 
some resemblance to C. sub-speciosum var. validius Nordst., but is 
much smailer and with less numerous and larger crenae on the 
margin. Fig. 31. 

* C. Gotlandicum Wittr. Burlington, Kans. Occasional. 
Diam.= 28 ; length= 36; isth—=9oy. This may be the same as 
C. rectangulare Grun. (in Rab. Fl. Eur. Alg.),and in that case the 
latter name should prevail by the right of priority, but it seems to 
me that the identity of the two is not positively proven. This is 
reported as occurring in Nebraska’. Fig. 26. 

* C. granatum Breb. var. sub-granatum Nordst. (341.) Ann 
Arbor, Mich. Occasional. Diam.—25 p; length=36y; isth— 
Sp. Fig. 16. 

* C. Paulense (Borgesen). (C. polymorphum Nordst. subsp. 
Paulense Borgesen). 277. Duval Co., Fla. Scarce. At the sug- 
gestion of Prof. Nordstedt I raise this to specific rank, as it seems 
to differ sufficiently from C. polymorphum Nordst. Diam.= 36 p; 
length=50 y; isth—10y. Slightly smaller than Borgesen’s speci 
mens and differing slightly in the position of the granules, but 
seemingly not distinct. In very clear specimens one can see that 
the wall is hyaline punctate between the five large granules on the 
front. Fig. 17. : 

*C. pericymatium Nordst. (445). Ithaca, N. Y. Occasional. 
Differing from Nordstedt’s figures’ in its deeper constriction in 
side view, but otherwise the same. Diam.=28y; length=5op; 
thick=26y; isth—24y. Fig. 28. 

*C. porrectum Nordst. (360.) Burlington, Kans. Frequent. 


‘8 Contr. from Bot. Dept. of Univ. of Nebraska. New Series III. 46. 14 Je. 1892. 
9 Nordstedt,O. Desmidieae Arctoae. Ofvers. K. Vet. Akad, Forh. 1875: pi. 


7. f. 26. 1875, 


294 


Slightly smaller than the original specimens from Brazil. Diam. 
=50y; length=50y; isth=13y. Fig. 30. 

* C. quadrum Lund. forma. (359.) Plymouth, N. H. Seems 
to be near var. minus Nordst. Semi-cells decidedly reniform at 
base; sometimes slightly retuse at the apex. Diam.—40 4; 
length=44 yp; isth—=14 yp. Fig. 22. 

* C. Raciborskii Lagerh. (365.) Meredith, N. H. Very rare. 
The membrane seems faintly granulate in the upper part of the 
semi-cell, but as the granules were not distinct they were not 
shown in the drawing. This agrees with the type in shape and 
surface, but differs slightly in the inner part of the sinus. Diam. 
=50y; length=44 y; isth—=21y. Fig.20. 

* C. subspectosum var. validius Nordst. (433.) Whitmore Lake, 
Michigan. Occasional. Diam.=48 »; length=62 y; isth—14/- 
Differs from the original description in having seven vertical rows 
of granules on the inflation and in the fact that they are in more 
or less definite horizontal rows also. This hardly seems sufficient 
ground for separating it as a new variety. Fig. 33. 

*C. synostegos Schaarschm. (437). Bridgeport, Conn., and 
Sisson, Cal. Common but very minute. The specimens from 
Bridgeport average 10 » in diameter, the others 8 » with length= 
11, thick=5 ». The angles in the Californian specimens do not 
appear cuspidate. In most cases the sinus gapes little, if at all- 
This may prove to be a distinct species. Fig. 29. 

*C. tetragonum Naeg. var. Lundellii Cke. (512). Ithaca, N. Y- 
Occasional. Diam.=24y; length=42; th.=16y. Fig. 21. 

*C. Turpini Breb. var. podolicum Gutw. (346). Baton Rouge, | 
La. Slightly smaller than Gutwinski’s measurements, but other- 
wise the same. Diam. 4o #; length=46p; th.=264 isth.= 
124. Fig. 18. 

Staurastrum brachiatum Ralfs. Duval Co., Fla. Rare. 

St. brevispinumBreb. Staten Island, N. Y., and Sisson, Cal. 
Frequent. 

St. commutatum Kiitz. (349). Duval Co., Fla. Rare. 

Sz. grande Bulnh. Bridgeport, Conn., and Meredith, N. H. 
Diam.=68-78 p. 

St. irthedrale Wolle. Frequent in collections from Florida, 
also from Staten Island and Ithaca, N. Y. Diam.=28—30 /- 


295 


Apex of semi-cell sometimes shows three slight elevations corre- 
sponding to the three lobes. 

ST. PROTRACTUM nov. nom. (.S¢. grallatorium forma Wolle") 
(428). This is certainly not a form of St. grallatorium. WVolle 
gives no note on it, but merely figures it. It may be described as 
follows: Length slightly less than breadth. Sinus shallow, linear, 
gaping widely in the outer half. Semicells nearly square with 
basal angles rounded and armed with two minute spines. Sides 
nearly parallel. Near the apex each is produced into a stout ser- 
rate ray tipped with three spines. Apex of semi-cell truncate, 
projecting above the rays and bearing several serrations. End 
view triangular, sides concave, angles drawn out into rays. Sur- 
face marked by several rows of serrations. Around the centre of 
the semi-cell (in end view) six larger bifid or trifid projections. 
Diam.=50-52 p; length=44 ». Whitmore Lake, Mich. Fig. 35. 

* St. leve Ralfs. (450). Whitmore Lake, Mich. Frequent. 
Diam.—14y; length=18y. Fig. 34. 

De Toni cites among the localities for this species “ Americae 
borealis (Wolle),"” but I cannot find that Wolle ever reported it, 
and believe this must be a mistake. 

* St. Reinschit Roy (348). Duval county, Fla. This does not 
agree exactly with Roy’s description, but it comes so near it that 
I place it here provisionally, The spines are short but distinct. 
It is smallerthan the type. Diam.—24,; length—20p. Fig. 15 

* Xanthidium antilopaeum Kiitz. forma _Javanicum Nordst. (324). 
Ann Arbor, Mich. Diam.—52p; length=52 4; thick.— 30 p; isth. 
=1I2y. The position and number of spines on the end of the 
semi-cell is variable, even in the same individual, but they show a 
tendency to arrangement in a single series. Fig. Io. 

X. concinnum Arch. var. Boldtianum West. (422). (Arthrodes- 
mus hexagonus forma Boldt.) Duval county, Fla. Occasional. 
Very minute. Diam.—12y; length—14y; thick—6y; isth.— 
4y. Nearly like Boldt.’s Fig. 17, but with sharper angles. In 
front view this looks like an Arthrodesmus, but in vertical view it 


10 Wolle, F. Desmids of the United States. 7.57. f 20.27. 1892. : 

"De Toni, J. B. Sylloge Chlorophycearum omnium hvcusque cognitarum. 
1227. 1889. 2 

12 Boldt, R. Bidrag till Kannedomen om Sibiriens Chlorophyllophyceer. Ofvers. 
K. Vet. Akad. Forh. 1885: 109. 74 5. 7.77. 1885. 


296 


shows a marked papilla on each side. It seems to be truly a 
Xanthidium. 

X, Tylerianum West. (X. antilopaeum Kitz. var. truncatum 
Hast. Wollein Desmids of the U.S. 1892 edition.) Ithaca, N.Y. 
Commonand typical. Although first described by Hastings, West's 
name must take precedence, since Hastings published the species 
merely in a local newspaper. 

The following species are believed to be new : 


ARTHRODESMUS MICHIGANENSIS n. sp. Fig. 12. 

Size medium. Length about one-third greater than breadth. 
Sinus gaping, broadly rounded, semi-cells being connected by a 
short neck. Semi-cells sub-elliptical, with the apex of the cell 
drawn out into a broad truncated lobe, and each end armed with 
a long slightly recurved spine. In vertical view rhombic elliptical 
with a long spine at each end. Side view circular. Diam.=24- 
30 #; length= 36-40 ; thick=16-18 yp; isth—=7-9 p. Whitmore 
Lake, Mich. Frequent. Type in herb. Johnson, 427. 
COSMOCLADIUM TUMIDUM n. sp. Fig. 23. 


Minute. Length and breadth nearly equal. Cells usually 
joined in fours by a hyaline, mostly unbranched filament. Great- 
est diameter of cell placed at right angles to the connecting fila- 
ments. Sinus deep, linear; semi-cells hexagonal elliptic, with 
apex flattened. In vertical view elliptical with slight but distinct 
median inflation. $ide view nearly circular. Diam=8 yz; length 
=9p; thick=5; isth—=2y. Whitmore Lake, Mich. Type in 
herb. Johnson, 455. 


This resembles C. subramosum Schmidle, but differs from it in 
its shorter cell, and in the median inflation of the semi-cell. j 


_ Ido not feel perfectly sure that Cosmocladium should be sepa- 
rated from Cosmarium. 


Cosmarium Paciricum n. sp. Fig. 25. 


_ Small. Length about one-fourth greater than the breadth. 
Sinus strongly gaping. Isthmus narrow. Semi-cells sub-ellipti- 
cal. Sides curving regularly from the base to above the middle, 
then converging by a sharp angle to form the rounded end. End 
view elliptical, with no inflation. Side view nearly circular. A 
single pyrenoid in each semi-cell. Membrane punctate. Diam. 
=25-28 ; length= 32-34 4; thick=15-17; isth=6y. Sisson, 
Siskiyou county, California. Altitude 3000 feet. M. A. Howe 
coll. Type in herb. Johnson, 402. 


297 


This resembles Staurastrum bientanum Rabh. var. connectens 
Boldt, but is much smaller, and lacks the short spines near the 
base of the semi-cells. It seems to be a true Cosmarium. It re- 
sembles C. zxflatum Wolle, differing chiefly in length of cell and 
curvature of the sides. Wolle’s figures are not aiways accurate, 
and it may be that this is a form of his species. 


COSMARIUM DISPERSUM n. sp. Fig. 109. 

Size,medium. Length and breadth about equal. Sinus deep, 
narrowly linear. Semi-cells semi-circular, with the apex some- 
times slightly flattened. Margin gently undulate with about 
eighteen elevations, often less distinct toward the apex. End view 
elliptical ; side view nearly circular. Two pyrenoids in each semi- 
cell. Membrane finely punctate. Diam.=40-44 4; length=4o p; 
thick.=20 1; isth=11y. Found in material from Baton Rouge, 
La., and from Meredith, N. H., at about the same time. Type 
(from Louisiana) in herb. Johnson, 347. 

BOTANICAL LABORATORY, UNIV. OF MICHIGAN. 


Description of Plates 232 and 233. 
(Reduced one-fourth in Photographing.) 
a=front view, vertical view, c—side view of cell. 


. Penium annulare West’ 560. 

F. cylindrus (Ehrb.) Breb., var. Silesiacum Kirch. 750. 
. Closterium Cynthia De Not. X 560. 

. C. tumidum nov. nom. < 560. 

. C. abruptum West « 560. 

C. angustatum Kiitz., var. clavatum Hast. 320. 

C. maculatum Hast. < 320 (a semi-cell). 

. Gonatozygon asperum (Breb.) Cleve. X 320, @ 759. 

. G. aculeatum Hast. < 320. 

. Xanthidium antilopaeum Kiitz. forma Favanicum Nordst. 560. 
. X. concinnum Arch., var. Boldtianum West. % 759. 
Arthrodesmus Michiganensis, n. sp. X< 560. 

- Luastrum Sibiricum Boldt. 750. 

. Micrasterias Nordstediana Wolle. 320. 

15. Staurastum Reinschit Roy. < 750. 

. Cosmarium granatum Breb., var. subgranatum Nordst. % 560. 
C. Paulense (Borgesen). < 759. 

C. Turpini Breb., var. podolicum Gutw. > 750. 

C. dispersum n. sp. X 750. 

C. Raciborskii Lagerh. > 750. 

. C. tetragonum Naeg., var. Lundellit Cke. X 750. 

C. guadrum Lund, forma. > 750. 


Pw hb 


Cor an 


ttt ttt 
Pw by HO 


2 eee « y 
Se ee 


298 


23. Cosmocladium tumidun n. sp. XX 750. 

24. Cosmarium pseudoprotuberans Kirch. X< 750. 
25. C. Pacificum n. sp. 750. 

26. C. Gotlandicum Wittr. X 750. 

27. C. elegantissimum Lund. > 750. 

28. C. pericymatium Nordst. X 750. 

29. C. synostegos Schaarschm. X 750. 

30. C. porrectum Nordst. 560. 

31. C. costatum Nordst. X 750. 
32. C. margaritatum (Lund.) Roy and Bissett. 560. 
33. C. subspectosum var. validius Nordst. < 750. 
34. Stavzrastrum leve Ralfs. % 750. 

35. S. protractum nov. nom. 560. 


The Genus Cenchrus in North America. 


By Gero. V. NASH, 


Cenchrus is a small genus of grasses containing about a dozen 
species, found principally in tropical and sub-tropical regions. 
In the United States it extends into the temperate zone. One 
species, C. tribuloides, the common bur-grass, occurs frequently in 
sandy soil along river banks, and has a very wide distribution, €x- 
tending from Massachusetts to Ontario, Minnesota and Nebraska, 
south to Mexico. It is also found in South America and in Cuba. 
The other species occurring in the United States are confined to 
the southern parts of the country, one, C. echinatus, extending as 
far north as North Carolina. 

The genus in the United States is readily divided into two 
well-marked groups, the first containing those forms in which the 
involucre consists of two spine-bearing valves, more or less con- 
cealing and enclosing the 2-6 spikelets, as illustrated in the com- 
mon C. tribuloides ; the second has only one species, the involucre 
consisting of 2-4 rows of erect bristles, the outer shorter than the 
inner, the single spikelet merely surrounded by the bristles and 
not enclosed. The single species belonging to this group is C. 
myosuroides, which ranges from Georgia and Florida to Texas, 


and south through Mexico to South America, where it is very 
common. 


- 


299 


Key to the Species. 


Involucre consisting of two spine-bearing valves, enclosing the 2-6 spikelets. 
Involucre armed at base. 
With shorter, generally reflexed spines; pedicels smooth; involucre 2-flow- 


ered, 
Involucre globose, pubescent. 1, C. tribuloides. 
Involucre ovate, smooth, 2. C. gracillimus. 


With erect barbed bristles; pedicels villous; involucre 4-6 flowered. 
3. C. echinatus. 


Involucre naked at base. 4. C. incertus. 
Involucre consisting of 2-4 rows of erect slender spines and bristles, at the base of the 
single spikelet. 5. C. myosuroides. 


I. CENCHRUS TRIBULOIDES L. Sp. Pl. 1050. 1753. 


C. Carolinianus Walt. Fl. Car. 79. 1788. Teste Pursh. 

Stems erect or ascending, generally decumbent at base, robust, 
2-6 dm, long, branching freely, sheaths generally strongly inflated, 
smooth, striate, compressed, the lower ones overlapping, the upper 
shorter than internodes, 3-8 cm. long; leaves 6-12 cm. long, 4-8 
mm. wide, smooth or scabrous, flat or slightly complanate ; spikes 
3-6 cm. long, generally partially included in upper sheaths, densely 
6-20-flowered; internodes of scabrous rachis 3-5 mm. long; invo- 
lucre 2-flowered, globose, pubescent to villous, yellowish to light 
green in color, smooth at base, spines 3-4 mm. long, slightly to 
Strongly ciliate at base, acutely barbulate at apex; spikelets 6-7 
mm. long, generally not exserted beyond the involucre; Ist glume 
hyaline, triangular, acute, about one-half as long as spikelet, spar- 
ingly scabrous, strongly I-nerved; 2d glume of firm texture, 
ovate, obtuse, about three-fourths as long as spikelet, scabrous, 
faintly 5-nerved at base, strongly so at apex; 3d glume firm, 
ovate, somewhat acute, about equaling spikelet, scabrous, 5-nerved; 
4th glume chartaceous, broadly ovate, acuminate, 5-7 mm. long, 
scabrous at apex, 5-nerved ; palet of first flower about as long as 
glume, 2-nerved, strongly scabrous between the nerves especially 
toward the apex, empty or enclosing 3 stamens; palet of per- 
fect flower chartaceous, broadly ovate, slightly shorter than glume, 
faintly 2-nerved, scabrous toward the apex; seed oblong-obovate, 
3 mm. long, 2—2.5 mm. broad, compressed. 


Extensively distributed in the United States ranging from 
Massachusetts to Ontario, Minnesota and Nebraska, south to Flor- 
ida and Mexico. Extremely variable, but its robust habit, gener- 
ally strongly inflated sheaths and light colored globose involucres 
will distinguish it from its nearest relatives. 


300 


“2, CENCHRUS GRACILLIMUS 0. sp. 


Stems ascending or erect, 3-6 dm. high, simple or occasionally 
sparingly branched above, slender; sheaths smooth, striate, the 
lower overlapping, the upper shorter than internodes, 6-7 cm. 
long, but slightly inflated; leaves complanate, 5-8 cm. long, 
about 2 mm. broad, acuminate, scabrous on margins and mid- 
nerve; spike finally long-exserted, 3-5 cm. long, loosely 3-6 
flowered, internodes of scabrous rachis about I cm. long; in- 
volucre 2-flowered, smooth, ovate, purplish, deeply furrowed and 
smooth at base; spines purple, 5-6 mm. long, 1 mm. broad at 
base where they are sometimes ciliate, acutely barbulate at the 
apex; spikelets about 8 mm. long, exserted beyond involucre 
for one-third to one-half their length; Ist glume triangular, hya- 
line, acute, one-half as long as spikelet, strongly 1-nerved, slightly 
scabrous; 2d glume firm, ovate, acute, 7-nerved, scabrous ; 3d 
glume of firm texture, ovate-lanceolate, acuminate, slightly shorter 
than spikelet, 5-nerved, scabrous especially toward apex; 4th 
glume chartaceous, 7~8 mm. long, ovate-lanceolate, long-acumi- 
nate, strongly 5-nerved at apex, scabrous for upper half; palet of 
first flower equaling glume, strongly scabrous, 2-nerved, empty or 
enclosing three stamens; palet of perfect flower ovate, long-acu- 
minate, chartaceous, a little shorter than glume, 2-nerved, strongly 


scabrous toward apex; seed oblong, 2.5 mm. long, I.5 mm. 
broad. 


Florida, occurring in the high pine land, together with C. ¢7i- 
ulotdes and C. echinatus, flowering a little earlier than the former 
and later than the latter. It is readily distinguishable from either, 
especially when growing in the field, by its long slender purple 
spines, smooth involucre, very slender habit and closely compla- 


nate leaves. Begins to flower early in April. My Nos. 188 and 
288, collection of 1894.. 


3. CENCHRUS ECHINATUs L. Sp. Pl. 1050. 1753. 


‘ Stems creeping, erect, 3-6 dm. long, branching ; sheaths smooth, 
striate, inflated, lower ones overlapping, upper shorter than inter- 
nodes, 8-12 cm. long; leaves 1-4 dm. long, 5-15 mm. wide, smooth 
or scabrous, flat; spikes 4-12 cm. long, finally more or less €X- 
serted, very densely 20-50 flowered, internodes of the scabrous 
rachis 2-3 mm. long; involucre 4-6 flowered, globose, green to 
purplish, more or less pubescent, villous at base; spines 3-4 mm. 
long, barbulate at apex, more or less ciliate at base ; bristles at base 
of involucre numerous, slender, distinctly barbed for their whole 
length; spikelets 6-7 mm. long, more or less exserted from in- 
volucre; Ist glume hyaline, lanceolate, obtuse or acute, one-half 
as long as spikelet, strongly 1-nerved, sparingly scabrous at apex; 


‘ 


301 


2d glume firm, ovate, obtuse, three-fourths as long as spikelet, 
5-nerved, scabrous; 3d glume firm, ovate-lanceolate, acuminate, 
slightly exceeded by fourth glume, 5-nerved, scabrous; 4th glume 
chartaceous, ovate-lanceolate, acuminate, 6-7 mm. long, 5-nerved, 
scabrous at apex; palet of first flower about as long as glume, 2- 
nerved, strongly scabrous ; palet of perfect flower as long as glume, 
chartaceous, ovate-lanceolate, faint!y 2-nerved; seed obovate to 
elliptical, 2.5 mm. long, 1.5 mm. wide, compressed. 

This plant is well marked by its broad flat leaves, and the 
slender barbed bristles surrounding the base of the involucre, 
which contains double the number of spikelets found in the other 
species. 

North Carolina to Texas, along the coast. 


4. CENCHRUS INCERTUS M. A. Curtis, Bost. Journ. Nat. Hist. 1: 


135) 1837. 

Stems erect or decumbent at base when young, finally creep- 
ing, 3-6 dm. long, smooth; sheaths smooth, inflated, compressed, 
striate, lower ones more or less overlapping, upper shorter than 
internodes, 5—9 cm. long; leaves smooth, flat, 4-15 cm. long, 3-5 
mm. wide; spike generally exserted, 3-6 cm. long, 8-20-flowered, 
internodes of the scabrous rachis 2-5 mm. long; involucre 2- 
flowered, pubescent, ovoid, smooth at base; spines about 3 mm. 
long, very broad at base, ciliate; spikelets 5 mm. long, more or 
less exserted from involucre; 1st glume hyaline, triangular, acute, 
strongly 1-nerved, one-half as long as spikelet; 2d glume of firm 
texture, ovate, obtuse, about three-fourths as long as spikelet, 5—7- 
nerved, scabrous; 3d glume ovate, acute, 5-nerved, somewhat 
shorter than fourth glume, scabrous; 4th glume chartaceous, 
broadly ovate, acute, about 5 mm. long, strongly 5-nerved and 
scabrous at apex; palet of first flower about equaling glume, 
strongly scabrous, 2-nerved, empty or enclosing three stamens; 
palet of perfect flower chartaceous, ovate, acute, about equaling 
glume, scabrous at apex; seed oval, 2 mm. long, 1.5 mm. broad. 

Ranges from South Carolina to Florida and Texas, The 
smaller involucre, naked at base, with fewer and shorter spines, 
readily distinguish this plant from C. triéuloides which it some- 


what resembles. 


5. CencHrus myosuroives H. B. K. Nov. Gen. 1: 115. 1815. 


Panicum cenchrotaes EN. Bot. S. C. and Ga. 1: 111. 181 7. 

Perennial from a running rootstock. Stems 6-14 dm. high, 
simple, or branched above, terete, smooth; sheaths somewhat in- 
flated, 7-14 cm. long, striate; leaves smooth, 1-3 dm. long, 3-8 


302 


mm. wide; spikes included in upper sheath, or more or less ex- 
serted, densely many-flowered, 5—20 cm. long, internodes of the 
scabrous rachis 2-5 mm. long; involucre consisting of I-2 rows 
of barbed spines as long as the spikelet, subtended by 1-2 rows of 
barbed bristles one-fourth to one-half as long; spikelet about 5 
mm. long, resting on the involucre and equaling its spines; 
glumes all more less short-pointed by the excurrent midnerve; 
ist glume hyaline, triangular, acute, about one-half as long as 
spikelet, I-nerved; 2d glume firm, ovate, acute, about three- 
fourths as long as spikelet, 5-nerved, sparingly scabrous; 3d 
glume as long as the fourth, firm, ovate, acute, strongly 5-nerved 
at apex; 4th glume chartaceous, ovate, acute, 5 mm. long, 
5-nerved at apex; palet of first flower from one-quarter to one-half 
as long as glume; palet of perfect flower equaling glume, strongly 
2-nerved and scabrous at apex. 

Ranges from Georgia (Elliott) to South Florida, western 
Texas and Mexico. 


Juncus scirpoides and its immediate Relatives. 


By FREDERICK VERNON COVILLE. 


In the year 1868 Dr. Engelmann grouped under /uncus scir- 
poides several closely related plants which, from the data now at 
hand, seem to require separation into four species. In 1880 Dr. 
Franz Buchenau described one of these as J. Engelmanni, and in 
1890 another as /. crassifolius. 1 am now convinced that still an- 
other should be separated specifically under the name of /. mega- 
cephalus, published sixty years ago by M. A. Curtis, of Wilming- 
ton, North Carolina. 

After examining the earlier literature of these plants it has 
been found necessary to change the current names of three species, 
in two cases substituting older names, and in one case supplying 
a newname. The first requirement for a proper disposition of 
the many published names was the positive identification of 
Michaux’s /. polycephalus, published in 1803, which, at different 
times, has been attached by various authors to nearly a dozen 
different plants, and has therefore fallen into disrepute and conse- 
quent disuse. With our present knowledge of the group, however, 
the name is capable of positive identification. The current dis- 


503 


position of /. polycephalus has been to treat it as a synonym of /. 
scirpoides Lam., a species published in 1789; but upon examina- 
tion of Michaux’s work it is found that he described two varieties 
of /. polycephalus, variety crassifolius and variety tenutfolius, re- 
spectively, giving to the former the diagnosis: « Major, foliis 
multo crassioribus et conspicue compressis.” It is clear that this 
variety, which is the type form of /. folycephalus, is one of the 
two plants known at the present time as /. Enge/manni Buch. and 
J. crasstifolius Buch. Since the latter species, however, is a plant 
of the Arkansas-Texas region there can be no doubt that 
Michaux’s /. polycephalus, accredited to Georgia and the Caro- 
linas, is the species now known as /. Engelmanni, which is abundant 
in these States, and it is necessary, therefore, to so treat it. 
Michaux’s second variety, Zenuifolius, was diagnosed simply as 
“foliis subfiliformibus.” Dr. Buchenau, examining a flower from 
Michaux’s specimen! of this variety, has pronounced it true Juncus 
Scirpottes. 
Key to the Species. 

Capsule with a true beak, the valves in dehiscence united above by the beak, their 


margins usually involute. . 

Uppermost cauline leaf with an almost obsolete blade rarely exceeding I or 2 cm. 
in length. 1. F. megacephalus, 
Uppermost cauline leaf with a normal blade, few to several cm. in length. 

Stamens as long as the inner perianth parts, the anthers exserted between 
them ; leaf-blades slender and terete. 2. F. scirpotdcs. 
Stamens considerably shorter than the inner perianth parts, their anthers in- 
cluded; leaf-blades usually stout and laterally compressed or even gladiate. 
3. F. polycephalus. 
Capsule without a true beak, the valves in dehiscence flat and separating throughout, 


their apices spreading or rarely loosely attached. 4. F. validus. 


I. JUNCUS MEGACEPHALUS M. A. Curtis.* 


Funcus megacephalus M. A, Curtis, Bost. Journ. Nat. Hist. 1: 
092, '1835. 


1 Buchenau, Monog. Junc. 325. 1890. 
* After examining a large number of herbarium specimens and observing both 
plants in the field for several years, I am convinced that Torrey’s Funcus nodosus 
megacephalus does not intergrade with ¥. zodosus proper. In view of the earlier 
Funcus megacephalus of Curtis, Torrey’s plant is named as follows: 
Juncus Torreyi nom. nov. 
Funcus nodosus var. megacephalus Torr. Fl. N. Y. 2: 326. 1843 


Funcus megacephalus Wood, Classbook Bot. Ed. 2: 724. 1861. Not Curtis. 


304 


Juncus scirpoides var. echinatus Engelm. Trans. St. Louis Acad. 
2: 468. 1868. 

Juncus scirpoides Carolinianus Coville, Mem. Torr. Bot. Club, 
5: 107. 1894. 

Plant 30 to 100 cm. high; blade of the uppermost cauline 
leaf only in very vigorous specimens exceeding I or 2 cm. in 
length ; fruiting heads 8 to 12 mm. in diameter; perianth reddish 
brown above; stamens one-half to two-thirds the length of the 


perianth, the anthers included; capsule equaling the perianth, its 
valves united above in dehiscence. 


Specimens have been examined only from the southern coast 
region from North Carolina to Florida, although the plant was 


accredited by Dr. Engelmann with a range as far north as Mary- 
land. 


2. JUNCUS SCIRPOIDES Lam. 


Juncus scirpoides Lam, Encycl. 3: 267. 1789. 

Juncus polycephalus tenuifolius Michx. Fl. Bor. Amer. 1: 193. 
1803. 

Juncus echinatus Muh\, Descr. Uber. Gram. 207. 1817." 

Juncus echinatus “ Muhl.;”’ Ell. Bot. S. C. & Georg. 1: 410. 
18172 

Juncus macrostemon J.Gay ; Laharpe Monog. Junc. 140. 1825. 

Juncus scirpoides var. macrostemon Engelm. Trans. St. Louis 
Acad. 2: 467. 1868. Includes the forms macrostylus and 
brachystylus. 

Juncus scirpoides genuinus Buch. Monog. Junc. 323. 1890. 


Plant 20-70 cm. high; leaf-blades terete, 2 mm. or less in 
thickness, with perfect septa; fruiting heads 8-11 mm. in diame- 
ter; perianth 2.5-3.5 mm. long, green throughout ; stamens equal- 
ing the inner perianth parts, the short anthers exserted ; capsule 
exceeding the perianth, its valves united above in dehiscence. 

A species of the coastal plain from New Jersey and eastern 


Pennsylvania southward through the Atlantic States to Florida 
and westward to Texas. 


1Buchenau (Monog. Junc. 323) has recorded his critical examination of flowers 
from Muhlenberg’s specimen, 
2 There is nothing to indicate that Elliott distinguished ¥. megacephalus from 


7. scirpoides, He undoubtedly included both, if indeed he ever saw the former, 
under F. echinatus. 


305 


3. JUNCUS POLYCEPHALUS Michx, 


Juncus polycephalus Michx. Fl]. Bor. Amer. 1: 192. 1803. 

Juncus polycephalus crassifolius Michx. Fl]. Bor. Amer. 1: 193. 
1803. 

Juncus nodosus polycephalus Pers. Syn. Pl. 1: 384. 1805. 

Juncus echinatus Muhl. Cat. 36. 1813.* 

Juncus scirpoides var. polycephalus forma minor Engelm. Trans. 
St, Louis Acad. 2: 468. 1868. 

Juncus Engelmanni Buch. Krit. Verz. Junc. 67. 1880. 

Plant robust, about 1 meter high; leaf-blades in most cases 
gladiate, 3-8 mm. broad and with incomplete septa, sometimes 
merely compressed, narrower and with complete septa; fruiting 
heads 8-12 mm. in diameter; stamens one-half to three-fourths as 
long as the petianth, the anthers included; valves of the capsule 
united above in dehiscence and usually with conspicuously involute 
margins; the beak well defined, commonly 2 mm. in length and 
not splitting at maturity. 

From North Carolina to Florida and westward through the 
Gulf States to Texas. This species, on account of its gladiate 
leaves, has been placed by Dr. Buchenau in the group with 
Juncus xiphioides and J. oxymeris, but its true relationship is un- 


questionably with /. scirpoides, as indicated by its flowers and fruit. 


JUNCUS VALIDUS nom. nov. 


Juncus scirpoides polycephalus forma major. Engelm. Trans. St. 
Louis Acad. 2: 468, 1888. 

Juncus crasstfolius Buch. Monog. Junc. 326. 1890. Not /. 
polycephalus crassifolius Michx. 1803. 

Plant 40 to 100 cm. high, stout and stiff; leaf-blades com- 
pressed, but never gladiate, 2 to 4 mm. wide, the septa complete ; 
fruiting heads 12 to 15 mm. in diameter; perianth 4 to 5 mm. 
long; stamens about one-half as long as the perianth, the anthers 
included; capsule exceeding the perianth, its valves flat, separat- 
ing throughout in dehiscence or sometimes slightly united above; 
no well defined solid beak present. : 

A plant of the southern prairie region, extending from Arkan- 
sas to Indian Territory, Oklahoma, and Texas on the west, and 
_to Mississippi on the east. 


* Muhlenberg based his name upon the ¥. folycephalus of Michaux, evidently 
not distinguishing the two varieties which Michaux included therein. 


306 


New Species of Physalis. 


By P. A. RYDBERG. 


Ina recent study of the North American species of Physalis the 
following new species have been determined. Since the material 
is widely scattered in herbaria, I deem it best to publish the de- 
scriptions pending the completion of the final monograph. 


” PHYSALIS SUBULATA N. sp. 


Fruiting calyx sharply 5-angled, more or less acuminate, calyx- 
lobes (at flowering time) lanceolate or acuminate, as long as the 
tube or longer; plant more or Jess villous or viscid-pubescent, 
erect, dichotomously branched, 2-4 decimetres high, stem angular 
and striate; leaves round-ovate, somewhat oblique at the base, 
generally coarsely dentate ; pedicels shorter than the small corolla, 
which is 2-3 millimetres in diameter ; calyx-lobes shorter than the 
corolla; fruiting calyx sharply angled and purple-veined, heart- 
shaped in section. 

This is intermediate between ?. Barbadensis and the South 
Mexican P. nicandrioides Schlecht. From the former it differs in 
the more glandular pubescence, and the long acumination of the 
calyx-lobes; from the latter in its smaller rounder leaves, 
in its calyx-lobes, which are shorter than the corolla, and in the 
fruiting calyx, which is smaller and not of a firm texture.* 

P. subulata has not yet been within the United States, but 
" comes near to its border. 


Mexico. State of Chihuahua: C. G. Pringle, No. 1344, 1887 


(type). It is in the following herbaria: Columbia College, Uni- 
versity of Minnesota and Professor Greene. 


Si 
PHYSALIS COMATA nN. sp. 


Perennial, erect, about 0.5 metre high; pubescence fine and 
short, that on the calyx, pedicels and upper branches mixed with 
long white flat jointed hairs ; like P. heterophylla Nees (P. Virginiana 
Gray, not L.), but leaves smaller, blade not over 0.5 decimetre long, 
rounded, ovate, scarcely at all cordate at the base, about 5 centi- 


* Another related Mexican species is P. hirsuta Mart. & Gal., not Dunal. It dif- 
fers from P, subu/ata in its larger, less veiny fruiting calyx, from P. nicandrioides by 
the calyx, which is not of a firm texture, and from both by its subentire leaves. It 
most resembles P. pubescens, but differs in its subulate calyx-tips. 


507 


metres long, thin, somewhat repand-dentate or nearly entire ; peti- 
oles as long as.the leaves; corolla greenish-yellow with brown 
centre; fruiting calyx as in P. hederaefolia Gray, but of thinner 
texture. 
The validity of this as a species may be doubted. It is inter- 
mediate between P. heterophylla and P. hederaefolia and might be 

placed as a variety of either with about equal right. It, perhaps, 
most resembles the latter, but differs in the thinner texture of the 
leaves and the fruiting calyx, in the larger flower and in the long 
white hairs. 

The following localities are recorded : 

Nebraska: P. A. Rydberg, No. 269, 1891 (type). 

Kansas: E. Bartholomew, No. 2, 1892; E. A. Popenoe, 1875. 

Texes: G. Jermy, No. 618, 1890. 


“PHYSALIS VERSICOLOR Nn. sp. 


Finely pubescent, in age glabrate; stem from a stout perennial 
root, much branched, at first erect but later spreading, slender, ob- 
tusely angled; lower leaves reniform-cordate, the upper ovate, 
all more or less oblique at the base, sinuately toothed, 2-4 centi- 
metres long, on slender petioles which are generally a little longer 
than the blade; pedicels slender, about the length of the petioles ; 
calyx-lobes taiangular-ovate, shorter than the tube; flowers about 
I centimetre wide, yellow or drab with brown spots in the centre, 
turning bluish in drying; fruiting calyx thin, ovoid-cylindrical, 
reticulate, decidedly 10-angled, 2.5-3.5 centimetres long, gener- 


ally open at the mouth. 
The specimens collected within the United States are more 


erect, have larger leaves and fruiting calyces than the Mexican. 
Rare within the United States. , 
New Mexico: C. Wright, 1851 (Mo. Bot. Gard. herb.). 
Arizona: Treadwell, 1879 (Cal. Acad. Sci.). 
Mexico, Guaymas: Ed. Palmer, 621 and 622. 1887. 


“ PHYSALIS VERSICOLOR MICROPHYLLA N. var. 


Like the species, but leaves only about 1 centimetre long, del- 
toid, coarsely toothed, peduncles about twice the length of the 
leaves; fruiting calyx nearly spherical, 1.5 centimetres long, 
tinged with purple. 

Mexico, Guaymas: Ed. Palmer, No. 94, 1887 (herbaria of iB 
Donnell Smith, of Columbia College, Professor Greene, etc). 


308 


’ PHYSALIS MACROPHYSA N. sp. 


Perennial; root somewhat thick and fleshy ; stem erect, 0.5-1 
metre high, comparatively slender, angled, perfectly smooth, or 
the upper parts sparingly pubescent with very short hairs ; leaves 
large, thin, 4-8 centimetres long, 2-5 centimetres wide, the lower 
obtuse, the upper acute or acuminate, on slender petioles 2-4 
centimetres long; pedicels 1-1.5 centimetres long, erect, in 
fruit reflexed; calyx smooth, lobes ovate-triangular or broadly 
lanceolate, generally a little shorter than the tube; corolla yellow 
with a dark centre, about 2 centimetres in diameter; anthers gen- 
erally yellow, sometimes tinged with purple; fruiting calyx large, 
3-4 centimetres long, 2.5-3 centimetres in diameter, pyramidal 
to ovoid-conical, indistinctly 10-angled, deeply sunken at the base; 
berry small, in the centre of the calyx. 


This is nearly related to P. longifolia and P. Philadelphica, but 
differs from both by its very large and inflated fruiting calyx 
and its broader leaves. The following specimens have been ex- 
amined : 

Arkansas: A. E. Heacox, 1889. 

Kansas: E. A. Popenoe, No. 68, 1875. 

Texas: Lindheimer, 1828; A. A. Heller, No. 1756. 1894. 

North Carolina (?): Small and Heller, No. 389. 1891.* 

Ohio (?): T. H. Horseford,* 1879; C. W. Short,* garden (?)- 


UNIVERSITY OF NEBRASKA. 


The Nomenclature Question, 


By Lester F. Warp. 


The German who was asked why he called his boy Hans re- 
plied: 

“ Pecaus it vas hees name.” 

The story is an old one, but no one has ever questioned the 
conclusiveness of the reply. It is the same answer that must be 
made to the question why a botanical name should be changed 
to make it conform to the law of priority. When a child is 
christened the name he receives is the one that he is supposed to 
have during life. A man with several aliases is always an object 


* These specimens lack fruit and may belong to the preceding, but the leaves 
most resemble those of P. macrophysa. 


309 


of suspicion. Is there any reason why the first name: that is 
given to a plant or animal should not always be its name as well 
as in the case of a human being? It is true that there is this dif- 
ference, that the poor plant or animal has no choice at any time, 
while the child after it becomes a man or woman might have 
something to say if an outsider should attempt to impose a differ- 
ent name. Still it does not seem that the principle is fundament- 
ally changed by this circumstance. If a lost child were to be 
found and named and brought up by the finder, and he should 
afterwards learn who his parents were and what his name was, he 
would very likely insist on being called by that first name. I re- 
member that one of my boyhood playmates was called ‘ Ed. 
Wheelock,’ but even when I knew him he was aware that Whee- 
lock was not his name, but that of the person who had adopted him, 
and afterward, having lost him for many years from view, on meet- 
ing him again, it was Mr. Edgar Currier with whom I had to do. 

Now it seems to me that all we are trying to do is to find out 
what the name of a plant is. It has happened in so many cases 
that plants have strayed, as it were, from home, been lost, adopted 
by strange persons, and given different names, lost for a time 
again and again found and renamed, and so on, that for us who 
now know them it is an exceedingly difficult matter to trace their 
history back and find out who they are. All this is due to the 
well known vicissitudes of all modern branches of natural history, 
especially of botany. In this general search for the true parentage 
and the true names of plants there should certainly be no differ- 
ence of opinion on the main question and all should admit that 
what is wanted is to ascertain the real name, for all synonyms are 
simply aliases, and the only real name is the first name. 

Nothing can certainly be more confusing than the existence of 
a large number of different names for the same thing, and it is no 
wonder that a movement was set on foot near the beginning of the 
present century, to endeavor to trace up the true history and find 
the true names of plants. It is a significant fact that this move- 
ment was initiated by a botanist, the great Augustin Pyrame de 
Candolle, in 1813, in his “ Théorie élémentaire de la botanique,” 
from which I translate ihe following paragraphs : 

Page 228: “In order that a nomenclature become universal it 


310 


must be fixed, and the fixity of that of natural history is founded 
onthis . .. principle . . . that the first one who discovers an 
object, or who records it in the catalogue of science, has the right 
to give it a name, and that this name must be necessarily ac- 
cepted, unless it already belongs to another object or transgresses 
the essential rules of nomenclature.” 

Page 241: “It may be said in general that any name which 
does not involve a contradiction with the plant, and especially 
which does not belong to any other species, is sufficiently good 
to be preserved. The impropriety of a specific name or the pos- 
sibility of finding more suitable ones is not sufficient to authorize 
a change.” 

Page 250, conclusion: “All this scaffolding of botanical 
nomenclature would crumble at its base and inevitably fall if the 
great majority of naturalists did not recognize the principle of 
which I have spoken, viz., the necessity of accepting the name 
given by the discoverer of a plant whenever that name is con- 
formable to the rules. A name cannot be changed because it has 
little meaning ; for on the same principle the second could be sup- 
pressed if a third better one was found, and the third if a fourth 
should present itself, etc.; thenceforward there would be no longer 
any fixity in nomenclature, or rather, there would be no longer 
any scientific nomenclature. The author himself who has first es- 
tablished a name has no more right than any one else to change 
it for the simple cause of impropriety. Priority, on the contrary, 
is a fixed, positive limitation, which admits of nothing arbitrary oF 
partial ; the most ancient name must therefore be always < admitted.” 

De Candolle, it is true, made five exceptions to this universal 
rule, some of which would not now be regarded as valid, such, for 
example, as his exception according to which the name Lunaria 
annua* might be changed because the plant is not an annual; but 
it has not been pretended that de Candolle fully grasped the im- 
portance of the movement, but only that the movement is itself in 
the nature of an evolution to which de Candolle, even that early, 
gave the initial impetus. 

The English mind did not become fully aroused to the subject 
until nearly thirty years later, but the movement in that country 


ET ean 


* This name is accepted in the « Kew Index.” [ED.] 


311 


was much broader and embraced not merely botany but all 
branches of natural history. | 

Such was the “Stricklandian code,’ adopted in 1842 and 
otherwise known as the “Rules of the British Association.” 
Among the great names connected with this Stricklandian code 
are those of Mr. Charles Darwin and Professor Henslow. In 
1860 this code was reénacted with only a few changes, all looking 
to greater success in attaining the same object. Mr. Darwin still 
served on the committee, likewise Mr. A. R. Wallace, Mr. P. L. 
Clayton, Professor Balfour, Professor Huxley, and among botan- 
ists proper Dr. J. D. Hooker and Mr. George Bentham. AA still 
further revision of the same was made in 1865, and this code now 
stands, but, strangely, has been supposed to be applicable only to 
zodlogy, although its provisions were equally applicable to the 
vegetable kingdom. In the preface of this code occurs this 
sentence: 

‘No one person can subsequently claim an authority equal to 
that possessed by the person who is the first to define a new genus 
or describe a new species.”’ 

In 1867 Alphonse De Candolle presented to the Inter- 
national Botanical Congress, held at Paris, a system of laws of 
nomenclature, upon which he had been long engaged and which 
with very few changes was adopted by that Congress. No one 
certainly could have felt more forcibly the evil effect of the multi- 
plication of plant names than the author of the Prodromus, and in 
the introduction to these rules he says “ in the four volumes of the 
Prodromus published from 1824 to 1830 the proportion of ad- 
mitted genera to synonyms was approximately 100 to 55; that is 
to say, there were at that time about half as many synonyms as 
admitted genera. In the Genera Plantarum of Bentham and 
Hooker, fascicles 1 and 2, published from 1862 to 1865, which 
comprise about the same series of families, I have found in mak- 
ing the same approximate calculation 117 synonyms for 100 ad- 
mitted genera. Therefore, the proportion of generic synonyms 
Must have doubled in 36 years.” This Candollean code was 
based, like the Stricklandian, on the law of priority and Article 
15 of that code is as follows : 

“Each natural group of plants can bear in the science only 


312 


one valid designation, namely, the most ancient one adopted by 
Linnaeus, or given after him on condition that it be conformable 
to the essential rules of nomenclature.” 

It was a noticeable fact that the Botanical Congress of Paris, 
which adopted these rules, was not attended by the English 
botanists, and Mr. W. B. Hemsley in an article in “« Nature” for 
December 24, 1891, says of this Congress: 

“In 1867 a Botanical Congress was held in Paris, to which 
botanists of all countries had been invited, and the most important 
subject discussed was botanical nomenclature. Mr. A. de Can- 
dolle had drawn up a most carefully considered code of rules to 
govern botanists in their writings; and this code was submitted 
to the assemblage of botanists, each rule being formulated and 
modified as the majority deemed wise. Finally the whole was 
printed and circulated. The fundamental principle of these laws 
was priority of publication with adeguate descriptions, and unfor- 
tunately it was made retrospective without any sufficiently defined 
statute of limitations. For reasons of their own the Kew botan- 
ists took no part in the proceedings of this Congress ; whether 
wisely or not it would be difficult to determine and fruitless to 
discuss.” 

It would be fruitless for me to discuss the reasons which have led 
the botanists of Kew to manifest so little sympathy with the gener al 
movement in favor of reform in nomenclature. Most of these 
reasons are well known to readers, but, as has already been 
said, this indifference was not due to any lack of appreciation of 
the importance of this reform, or of general sympathy with it, 
on the part of the two great leading systematists of England, Dr. 
Hooker and Mr. Bentham, who, as we have seen, both signed the 
Stricklandian code. In fact, no systematist has ever squarely ap- 
proached the question and given it due attention without arriving 
at substantially the same conclusion. Dr. Asa Gray in his Struc- 
tural Botany, page 348, says: “ For each plant or group there can 
be only one valid name and that always the most ancient if it is 
tenable, consequently no new name should be given to an old 
plant or group except for necessity. That a name may be bet- 
tered is no valid reason for changing it.” And on this principle 
it is worthy of note that against his convictions he maintained 


513 


our common blue violet under the name Viola palmata L., var. 
cucullata Gray, because he admitted the necessity of taking up an 
older name if the plant should be given specific rank, saying: 

“ Viola cucullata Ait. ought to have been referred, as an entire- 
leaved variety, to the Linnaean Viola palmata. I am the more 
constrained to do so now by the fact that the name cucullata 
would have to give way to the much earlier published V. odligua 
Hill, well figured and unmistakable in his Hortus Kewensis.”"* 

Dr. David Starr Jordan, President of Leland Stanford Univer- 
sity and a well-known ichthyologist has said, “ There are only 
two ways of naming plants or animals, either to give them their 
oldest names or to give them any names you please.”+ Not- 
withstanding the general agreement among zodlogists to the 
principle of the Stricklandian code it was found difficult to en- 
force these principles unanimously, and in 1876 the question came 
up afresh at the Buffalo meeting of the American Association for 
the Advancement of Science, and a new and slightly modified 
code which had been drawn by Professor Dall was adopted one 
year later.t It deals largely with the multitudinous details of 
zological science and makes no concessions, but holds the gen- 
eral law of priority as the basis of all sound nomenclature, which 
is there reénacted and amplified. This code is now, I believe, 
almost unanimously enforced by zodlogists within the United 
States. 

It does not, however, seem to have covered the case of orni- 
thology, and the ornithologists were still in the worst possible con- 
dition in relation to the multiplication of names. At last, losing, 
as it would seem, all patience with the system in vogue, they met, 
and bya unanimous action of the American Ornithologists’ Union 
the most stringent code of nomenclature was adopted that has 
ever been proposed. This is known as the code of nomenclature 
of the American Ornithologists’ Union, published in 1886. The 
ornithologists had the advantage of perfect unanimity, which is 
one of the most important conditions to making any code a com- 
plete success. The condition of affairs that prevailed before this 


*Asa Gray, Botanical Gazette, 11: 254. 1886. 
+Botanical Gazette, 20: 163. 1895. 
tProc. A. A. A. S., 26: 1877. Appendix. 


314 


list (1886) is well shown by comparing the lists that preceded it, 


that is, the lists of North American birds published by Baird in 
1859, Coues in 1874, Ridgway in 1881 and Coues in 1882. By 
taking the first fifty genera given in the American Ornithologists’ 
Union check-list it is found that in only five cases has the generic 
name remained the same from 1859 to 1886. That is, 45 of the 
50 generic names (90 per cent.) have been unstable. Since the 
American Ornithologists’ Union list xot one of these fifty names 
has changed. The accompanying tabular statement will show 
more clearly than words the changes in these fifty genera. This 
complete list embraces 322 genera and about one thousand spe- 
cies and sub-species. In the ten years that have elapsed since its 
publication it has been found necessary to change only three gen- 
era, one sub-genus, three species and one sub-species by action of 
the law of priority. (See opposite table.) 

This truly astonishing result must certainly be highly gratify- 
ing to the ornithologists, and the question arises whether botan- 
ists can bring about any such result in their department. A feel- 
ing in favor of such a movement has been growing stronger and 
stronger for a number of years, and has at last taken shape in the 


appointment of a committee of the Botanical Club of the Amer- 


ican Association for the Advancement of Science at Rochester, in 
1892, who proposed a set of rules of botanical nomenclature with 
which all are now familiar. The Club also authorized the. publi- 
cation, as an earnest of what the enforcement of these rules might 
be expected to accomplish, of a list of the plants of northeastern 
North America. This list is also too familiar to require com- 
ment. Of it Professor Bessey has said: «‘ This book is the sign 
that the day of ‘ authority ’ as such, is ended, and the day of law’ 
has begun.* All that it seems necessary to say is that there seems 
to be a misapprehension on the part of some botanists as to the 
method by which this list was prepared, it being imagined by a 
few persons that the particular individuals who had most to do 
with it were in some way personally responsible for the result. It 
should be known to all that they were merely the instruments in 
the hands of a large committee, and that every question was sub- 
mitted to all the members of that committee, even when not in 


—$——$—— 


* American Naturalist, 29: 350. 1895. 


Podiceps. 
Podiceps. 
Podilymbus. 
Colymbus. 
Mormon. 
Mormon. 
Cerorhina. 
Ptychorhamphus. 
Ombria. 
Phaleris. 
Brachyramphus. 
Brachyramphus. 
Uria. 

Uria. 

Alca. 

Alca. 

Mergulus. 
Stercorarius. 
Stercorarius. 
Pagophila. 
Rissa. 

Blasipus. 
Rhodostethia. 
Xema. 

Sterna. 

Sterna. 
Hydrochelidon. 
Sterna. 
Rhynchops. 
Diomedea. 
Diomedea. 
Diomedea. 
Procellaria. 
Procellaria. 
Puffinus 
Procellaria. 


Daption. 


Thalassidroma. 
Thalassidroma. 
Thalassidroma. 
Fregetta. 
Phaéton. 

Sula. 

Plotus. 
Graculus. 


COUES RIDGWAY. CouEs. A. O. U. CHEcx- 
1874. 1881. 1882. ake a 
| 
Podiceps. Aechmophorus. | Aechmophorus. Aechmophorus. 
Podiceps. Podiceps. Podiceps. Colymbus. 
Podilymbus. Podilymbus. Podilymbus. Podilymbus. 
Colymbus. Colymbus. Colymbus. Urinator. 
Fratercula. Lunda. Fratercula. Lunda. 
Fratercula. Fratercula. | Fratercula. Fratercula. 
Ceratorhina, Ceratorhina. | Ceratorhina. Cerorhinca. 
Ptychorhamphus. Ptycorhamphus. | Ptychorhamphus. Ptychorhamphus, 
Phaleris. Phaleris. Simorhynchus. Cyclorrhynchus. 
Simorhynchus. Simorhynchus. ' Simorhynchus. Simorhynchus, 
Synthliborhamphus. Synthliborhamphus. | Synthliborhamphus. Synthliboramphus. 
Brachyrhamphus, Brachyrhamphus. | Brachyrhamphus. Brachyrhamphus. 
Uria. Uria. | Uria. Cepphus. 
Lomvia. Lomvia. | Lomvia. Uria. 
Utamania. Utamania. , Utamania. Alca. 
Alca. Alca. | Alea. Plautus. 
Mergulus. Alle. | Alle, Alle. 
Stercorarius. Megalestris. Stercorarius. Megalestris. 
Stercorarius. Stercorarius. Stercorarius. Stercorarius. 
Larus. Pagophila. | Pagoph la. Gavia. 
Larus. Rissa. | Rissa. Rissa. 
Blasipus. Larus. | Larus. Larus. 
Rhodostethia. Rhodostethia, Rhodostethia. _Rhodostethia. 
Xema. Xema. Xema. Xema. 
Sterna, Sterna. Sterna. Gelochelidon. 
Sterna. Sterna. Sterna. Sterna. 
Hydrochelidon. Hydrochelidon. Hydrochelidon. Hydrochelidon. 
Anous. Anous. Anous. Anous. 
Rhynchops. Rhynchops. Rhynchops. Rhynchops. 
Diomedea. Diomedea. Diomedea. Diomedea 
Diomedea. Thalassogeron. 
Diomedea. Diomedea. ~ Phoebetria. Phoebetria. 
Fulmarus. Ossifraga. | Ossifraga. Ossifraga. 
Fulmarus. Prioculla. - | Priocilla. Fulmarus. 
Puffinus. | Priofinus. Priofinus. Puffinus. 
Aestrelata. Oestrelata. | O<cstrelata. Aestrelata. 
Oestrelata. | Oecestrelata. Bulweria. 
Daption. Daption. | Daption. Daption. 
Halocyptena. Halocyptena, | Halocyptena. Halocyptena. 
Procellaria. Procellaria. | Procellaria. Procellaria. 
Oceanodroma. Oceanodroma. | Oceanodroma. Oceanodroma. 
Oceanites. Oceanites. | Oceanites Oceanites. 
Fregetta. Fregetta. | Fregetta. Cymodroma. 
Phaéthon. Phaéthon. _ Phaéthon. Phaéthon. 
Sula. Sula. Sula. ne 
Plotus. Plotus. | Plotus. Anhinga. 
Graculus. Phalacrocorax. | Phalacrocorax Phalacrocorax. 


515 


session, by sending out circulars, and that the deliberate vote of 
each member was taken in each case and the questions settled, 
where not unanimous, by a majority vote. For my own part I 
confess that I voted with the minority on a number of minor 
questions, but always with the feeling which I observed to prevail, 
not only among the members of the committee, but apparently 
throughout the Botanical Club itself, that minor questions were to 
be ignored in the presence of the great necessity for the adoption 
of rules to which all would subscribe. This list, prepared under 
many disadvantages, is, of course, imperfect in many respects and 
contains a few features which are especially irritating to those who 
attempt to use it. I have never known a botanist who was not 
irritated at the changing of names, yet we have all been obliged 
during the entire course of our studies to submit to wholesale 
changes of names at periodical intervals. This is no new thing, 
as any one may learn by reading the preface to the sixth edition 
of Prof. Amos Eaton’s Manual of Botany, published in 1833. He 
says: 

“It may be asked, why I do not follow De Candolle, servilely, 
since so many distinguished botanists have borne testimony to 
his great merit? Perhaps no one is a more devoted admirer of 
his discriminating talents, great learning and untiring assiduity 
than myself. But he imposes on his readers the labor of learning 
a multitude of new names without even a shadow of pretence, 
* * % * As far as I have any influence I pledge it here that 
the embarrassing innovations of De Candolle and others are of no 
possible use to the science of Botany. All new discoveries, how- 
ever (which are not a few), should be adopted; and they are 
adopted in this edition. And the mecessary new names and new 
nomenclature are also adopted and fully explained.” (Italicized 
as in the original). 

No one can doubt Professor Eaton’s high motive in giving 
utterance to what he considered so conservative an expression of 
his views regarding the changes in classification and nomenclature 
made by De Candolle, and doubtless he considered De Candolle’s 
researches as ill-advised and ephemeral as do some of our esti- 
mable contemporaries the Association code of to-day, yet I am 
constrained to look upon their protests as belonging to no differ- 
ent category than Professor Eaton’s. 


¢ 


316 


The botanical world has submitted to frequent changes like those 
we have known in the past fifty years with very little remonstrance 
compared to the great annoyance which they produce. In this work 
a new set of changes is thrust upon us, some of them very great 
and calculated to appeal strongly to our sense of veneration for 
the older names which we have so long known, and it is not to 
be wondered at that those who do not understand that there is 
any difference between this movement and the long series of 
changes that have been introduced in times gone by, in the differ- 
ent editions of our manuals and the new botanical works that 
have appeared, should strongly resent this last proposition to com- 
pel us to memorize a new set of names. In America the princi- 
pal reasons for submitting as tranquilly as botanists have done to 
the changes that have been imposed has chiefly been the great re- 
spect in which all American botanists have held the authors of 
these books. In the case of Dr. Asa Gray that respect amounted 
in avery large number of cases to something more—to a real 
sentiment of personal affection; but this condition of things no 
longer exists. The argument at best was an unsound one, but 
one which was nevertheless effective. At the present time botan- 
ists must be convinced that any wholesale changes that are to be 
introduced in the botanical nomenclature of America are made 
for good reasons. 

But, on the other hand, there is now far greater necessity for 
the adoption of some fundamental rules of nomenclature than 
have heretofore existed. Formerly there was one high seat from 
which the botanical decrees emanated, and there was far less dan- 
ger that unreasonable things would be done by one or two per- 
sous than by many. At the present time there are large numbers 
of botanical centres, and if matters are to be left to the individual 
judgment of publishing botanists, there will be no comparing the 
confusion that is in store for us with that which we have had in 
the past. Heretofore we have only had the differences which one 
man or one class of men in close codperation thought best to in- 
troduce at different periods in their own work. Now we shall 
not only have the changes that each individual is likely to make 
at different dates, but as many differences as there are different 
sources from which our works are to emanate. It is difficult 


GRAY, MANUAL, GRAY, MANUAL, 
ED:: 1. Ep: 2 


GRAY, MANUAL, 
ED. 4. 


GRAY, MANUAL, 
ED. 5. 


Lappa major Gaertn.|Lappa Major Gaertn. 


Nabalus Fraseri DC. |Nabalus Fraseri DC. 


Diplopappus amygda-|Diplopappus amygda- Diplopappus amygda- 


linus T, & G. linus T, & G. 


Solidago serotina Ait.|Solidago serotina Ait. 


me gigantea Ait.) « gigantea Ait.) gigantea Ait. 


Lappa major Gaertn. 
Nabalus Fraseri DC. 


linus T, & G. 
Solidago serotina Ait, 


Lappa officinalis All. 
Nabalus Fraseri DC. 


Diplopappus amygda- 
linus T & G. 
Solidago serotina Ait, 


MK gigantea Ait. 


Gray, SYNOPT. 
FL. N. Am. 


Arctium Lappa L. 
Prenanthes serpenta- 
ria Pursh. 
Aster umbellatus Mill. 
var. latifolius G. 
Solidago serotina Ait. 
var. gigantea G. 
Solidago serotina Ait. 


GRAY, MANUAL, — 
Ep. 6. 


Arctium Lappa L. 
Prenanthes serpenta- 

ria Pursh. 
Aster umbellatus Mill. © 

var, latifolius G. 
Solidago serotina Ait. 
var. gigantea G. 
Solidago serotina At: 


Smilacina bifolia Ker. Smilacina bifolia Ker, Smilacina bifolia Ker, 


| var. Canadensis G. 
| 
| 


var. Canadensis G, 


Smilacina bifolia Ker, 
var. Canadensis G, 


Maianthenum Cana- 
dense Desf. 


Gray, MANUAL, Gray, MANUAL, 
ED. 1. Ep. 2. 


Gray, MANuaL, 
ED. 4. 


Pulsatilla patens Mill.|Pulsatilla Nuttalliana/Pulsatilla Nuttalliana 
. Gray. 
Ranunculus Purshii/Ranunculus Purshii Ranunculus Purshii 


Rich.| — Rich. 


Ranunculus aquatilis Ranunculus aquatilis Ranunculus aquatilis 
L. L. 


var. divaricatus Gr. 
Viola Muhlenbergii Viola Muhlenbergii 

Torr.| Torr, 
- Elodea Virginica Nut. /Elodea Virginica Nut. 


Lechea thymifolia Lechea thymifolia 
Pursh. 


Gray, MANUAL, 
ED. 5. 


Watson, BIBLIOG. 
INDEX. 


GRAY, MANUAL, 
Ep. 6. 


Gray. 

Rich. 

L. 

var. divaricatus Gr. 
Viola Muhlenbergii 
Torr. 
Elodea Virginica Nut. 
Lechea thymifolia 


Anemone patens L. 
var. Nuttalliana Gr. 
Ranunculus multifidus 
Pursh. 
var. terrestris Gray. 
Ranunculus divarica- 
tus Schrank, 


Viola canina L. 
var. sylvestris Regel. 


Elodes Virginica Nut. 
L echea hymiglia 


Pursh. 
Spergularia media 
Presl. 


Nuphar luteum Smith.| 


var, macrocarpa Gr.) 


Anemone patens L. 
var. Nuttalliana Gr, 
Ranunculus multifidus 
Pursh. 
Ranunculus aquatilis 
L. 
var. stagnatilis D.C. 
Viola canina L, 
var. sylvestris Regel. 
Elodes Virginica Nut. 
Lechea_ thymifolia 


als 


See ed ' 


Anemone patens L. 
var. Nuttalliana Gr. 
Ranunculus multifidus 


var. — Gray. 
Ranunculus circinatus 
: Sibth. 


Viola canina L. 
var. Muhlen 


Elodes 


317 


under such circumstances to imagine what the condition of things 
would be were this to go on for several generations. If this ist 
was as complete and general as that of the Ornithologists’ Union 
adopted in 1886, there is no reason to suppose that the result in 
botany may not be practically the same as it has proved to be in 
ornithology, and that with the publication of this one last set of 
changes,'which would be simply a serious attempt to actually find 
what the true names of our plants are, the long continued process 
of bandying these plants about from one name to another must 
cease and each plant would have at last found its true and per- 
manent resting place. 

To illustrate in botany as has been done in ornithology we 
may take several of the editions of Gray’s Manual, Sereno Wat- 
son’s Bibliographical Index and Gray’s Synoptical Flora, and 
make a few comparisons to show the fluctuations that species of 
American plants have undergone. (See opposite table.) 

These are only a few samples taken almost at random of the 
extensive changes that were made at the different dates given. 
To mention my own personal experience, I began with the fourth 
edition of Gray’s Manual only a short time before the appearance 
of the fifth, yet long enough for me to have wasted many precious 
hours in memorizing names destined to be changed. And then 
came the Bibliographical Index for the Polypetalae, introducing 
large numbers of other changes, followed by the Synoptical Flora, 
Carrying the work into the Gamopetalae. The sixth edition of 
Gray’s Manual edited by Mr. Watson often differs from any of 
the preceding, showing that the general work of wholesale alter- 
ation was still going on. Many botanists supposed, as I did at 
first myself, that all this was necessary and often the authors 
stated that the reasons for their changes were because the names 
formerly published were not the original names, thus directly ap- 
pealing to the law of priority and defending themselves under this 
law, but a general glance at the whole affair shows there never 
was really any systematic attempt to base these changes upon any 
permanent and consistent scientific principles, but that to a large 
extent it was left to the individual judgment of the author at the 
particular time at which he was writing. The utter chaos into 
which this system has thrown the science of botany is the real 
cause of a movement for a stable nomenclature. 


318 


But it would seem that notwithstanding the general spirit of 
harmony that prevailed in the Botanical Club, and especially in the 
large committee that it appointed, the work that has thus far been 
done does not receive the unanimous approval of the working 
botanists of the country. <A circular has recently been sent out 
bearing the signatures of a considerable number of men whom the 
science of botany justly honors, which is, in fact, in the nature of 
a protest against the movement. In urging the “ postponement 
of any radical measures of reform” these gentlemen seem to ad- 
mit the possibility of reform and perhaps the need of it, but, 
after a careful reading of this paper, I am obliged to conclude that 
it is in the main the result of the temporary irritation, already 
mentioned, which any new attempt to change the names of our 
plants is certain to produce. Of course, there are other causes 
arising out of the respective claims of rival universities, etc., etc. 
Especially is the argumentum ad verecundiam very prominent, and 
I might almost say justly so, since I yield to none in the profound 
respect which is generally shared for the great and good Dr. 
Gray, and for the unrivalled work in systematic botany that has 
been done at Harvard University. But still I am disposed not to 
permit mere sentiment to stand in the way of the settlement of so 
momentous a question as the one now before the botanical world, 
and I must say frankly, with all due respect for the eminent names 
that are appended to this circular, that I do not regard their gen- 
eral argument as a sound one, and I look upon the circular as 
little more than an appeal to botanists to preserve the status quo. 
In other words, it seems to be the product of that natural conser- 
vatism which always goes hand in hand with the spirit of progress 
and has its true function of preventing rash actions and hasty 
revolutions. With this spirit of order I fully sympathize, but at 
the same time I believe that the time has come for the completion 
of the reform movement which has merely been arrested, although 
gradually gaining ground since the date of the Stricklandian and 
of the Candollean codes. I do not regard the present movement 
in any sense revolutionary. It is merely an attempt on the part 
of botanists to secure a uniform system which has not thus far 
been actually put in practice, except to a limited extent. It is 
now proposed to practice what has been preached. 


319 


Before attempting to discuss any of the so-called principles 
laid down in this circular, it may be premised that the advance 
movement should be regarded as essentially one of disinterested — 
principles which only has to oppose what is really prejudice, but 
may be called by the milder names of sentiment and conservatism. 
The botanists who approve of these rules have just as much to 
lose as those who oppose them, and the difference is that they are 
willing to make this sacrifice, not for their own sakes, but for the 
sake of the future of botany. Their work is therefore a labor of 
love. It is opposed to their personal interest, and they represent 
the class of botanists who are willing for the sake of the future, 
in which they will have no part, to make a great personal sacri- 
fice. Very few of the older ones will ever be able to rid them- 
selves of the older names with which they are now familiar. Only 
the very young workers can hope that this action will redound to 
their personal advantage. Those who oppose this movement, if 
there be any (and I have no doubt there are) who really see that 
it might be the last time that serious changes would have to be 
made in botanical names, would seem to do so purely from a per- 
sonal disinclination to incur the annoyance of accustoming them- 
Selves to a new set of names. It must be admitted that this mo- 
tive is not as high as we might ‘hope botanists generally to be 
actuated by. It is probable that some of the signers of this cir- 
cular think that no stable nomenclature is possible. It is for the 
benefit of such that I have introduced the argument showing the 
action of the Ornithologists’ Union, and surely no one can deny 
that it is equally applicable to botany. In so far as any personal 
rivalry or rivalry between different institutions is concerned I take 
no interest in it, and arguments of that nature are not only un- 
worthy of an answer, but really admit of no answer. 

As regards the familiarity of names and their sanctity on that 
account, much more is made of it than it deserves. For exam- 
ple, I have been obliged to familiarize myself with no less than 
four different sets of botanical names in the course of my own 
work, . The first name I learned for a common plant was felt by 
me to be sacred, and it seemed a sacrilege to substitute another, 
but when convinced that it was necessary I submitted, and soon 
the second name became equally sacred with the first, and so on 


320 


to the end. Now this is the case with us all in learning the names 
_of our familiar plants. The particular name that we learn for a 
a plant is all that makes it sacred, and the rising generation of 
botanists, who will have only before them the actual first name or 
the real name of the plant, instead of some false synonym that 
occurs in the present books, will look upon that name with the 
same veneration as we did upon the false one, and the names that 
we have learned to cherish will be to them nothing but worthless 
synonyms. In their case this will be true, whereas in ours we 
were simply cherishing the names that did not properly belong to 
the objects to which they were applied. 

I have said that the new movement is not only not revolutionary, 
but is simply in the nature of an evolution which has long been 
going on. On the contrary, it might be maintained that the so- 
called principles embodied in this circular, which are alleged to be 
an expression of conservative views, are really, on the contrary, 
revolutionary in their character. The following are the principal 
codes which have been proposed by responsible organizations for 
the guidance of writers in zodlogy and botany : 

De Candolle’s Théorie élémentaire de la botanique, 1813. 

British Association for the Advancement of Science, 1842. 

Association of American Geologists and Naturalists, 1845. 

International Botanical Congress, Genoa, 1865. 

American Association for the Advancement of Science, 1877- 

Societe Zodlogique Internationale, 1882. 

American Ornithologists’ Union, 1883-85. 

International Botanical Congress, Genoa, 1892. 

Botanical Club, American Association for the Advancement of 
Science, 1892-93. 

The codes adopted by these associations show a steady ad- 
vance from the idea of giving genera and species names to suit 
individual taste toward the idea of giving them strictly their old- 
est names. And the history of nomenclature shows an advance 
in stability and uniformity corresponding exactly with the thor- 
oughness with which these codes have been carried out. The 
_ circular to which reference has been made proposes a new de- 
parture in nomenclature, revolutionary in its character and, judg- 


ing from the history of the science, capable of producing most 
chaotic results. 


321 


It may be said to embody four rules or principles which are 
opposed to those adopted by the Botanical Club of the American 
Association. These, which may for convenience be designated 
the Harvard rules, afford a good opportunity for the comparison 
of the two codes, which every botanist should make for himself 
deliberately and judicially. It seems eminently desirable that 
those who have not yet given thought to the subject should un- 
derstand the full significance of the problem with which the As- 
sociation botanists have been dealing. The first of them, that 
relative to ordinal names, it is unnecessary to discuss from the 
standpoint of the new nomenclature, for no official action on this 
matter has as yet been taken by the botanical club committee. 

The Harvard rules are promulgated after a distinct statement 
of belief* that no stability in nomenclature is possible, and that 
the decision as to what names shall be used for genera is to be 
left to the judgment of individual botanists and not decided by 
the principle of priority.+ 

These rules represent the system which for many decades 
botanists all over the world have been trying to escape, a system 
which renders the nomenclature of a book thirty or forty years 
old largely unintelligible, except to the systematist, and which 
gives every promise of repeating its own history. It is preémi- 
nently a /azssez faire system, and the most that can be claimed for 
it is that it has served « fairly well.” .If at the beginning of the 
present century botanists had adopted a system based on priority, 
how great would be our obligation to them! Instead of a hun- 
dred years of heterogeneous and largely unrecognizable names, we 
should have had a botanical literature in which a plant would al- 
ways have had the same name, and ready intelligibility of this lit- 
erature would be possible to every reader. The botanists of the 
next century will not, it is hoped, have such a hundred years of 
constant change to look back upon as we in our time have had. 

Some botanists are prone to pin their faith to the arbitrary 
authority of a standard book, and are holding up that truly mag- 
nificent work, the Index Kewensis, now nearly completed, as the 
safe and only guide in nomenclature. But history shows that the 


* Robinson, Botanical Gazette, 20: 103. Ap. 1895. 
+ Harvard Rules, No. 2 (May, 1895). 


322 


influence of such a work is only temporary at best. Where now 
is the authority of Steudel’s Nomenclator, of Pfeiffer’s Nomencla- 
tor, and to what extent have they contributed to uniformity in 
plant names? Both these works have filled an important place in 
the literature of botany, but for the simple reason that they did 
not bring forward the oldest name as the valid one they have 
added little to the stability of our nomenclature. It is greatly to 
be regretted that at the time when the Index Kewensis was in 
preparation the demand for a stable nomenclature had not yet be- 
come sufficiently strong to lead its authors to add principle to 
prestige and thus insure its permanency as a nomenclator in addi- 
tion to its inestimable value as an index. 

In the prelude to the Harvard rules reference is made to the 
calling, at an early date, of an International Botanical Congress, 
presumably for the purpose of ‘settling’ the nomenclature ques- 
tion. It may not be out of place here to urge upon every one 
who may be a delegate to that Congress, or who may aid in the 
selection of a delegate, the careful consideration of the fact that no 
law is stronger than the authority that makes it, and that no au- 
thority is stronger in the end than the principle upon which it 
rests. 

A specialist in fungi recently made the admission in conver- 
sation, not only that the objections to the new code did not apply 
in the case of the lower cryptogams, but that the actual applica- 
tion of the code itself would be desirable. The reason given for 
this was that these orders are not popularly known and hence 
their nomenclature has not become established by usage. . There 
is the same tacit admission in the language of the Harvard circular: 

‘These rules [the Harvard rules] are designed to apply only 
to phaenogams and vascular cryptogams.” 

In all the lower orders of plants, then, we are to be guided by 
the law of priority; but as soon as the Pteridophyta are reached, 
principle is cast aside in favor of sentiment, and because Swartz’ 
name of Asfidium happens to be in common use among fern- 
gatherers, we are enjoined from taking up the perfectly valid de- 
signation Dryopteris given years previously by Adanson ! 

Relative to international action it is a matter of gratification to 
note that recent legislative change has been in conformity with the 


* 


323 


American Association principles. The adoption of the first edi- 
tion of Linnaeus’ Species Plantarum, 1753, as the starting point of 
our system of nomenclature was carried through the Genoa Con- 
gress immediately after its adoption at Rochester in 1892. The 
Austro-German botanists in a meeting held last September 
adopted another fundamental principle of the American code, the 
retention of the oldest specific name, under whatever genus pub- 
lished, a principle already put in practice in the later numbers of 
Engler and Prantl’s Natiirlichen Pflanzenfamilien. 

There are some botanists who hold that the Association rules, 
although sound, should not be made retroactive. Even in the 
Harvard rules, three of the four Association rules are expressly ad- 
mitted to be desirable in future practice. It should be evident to 
every thoughtful person that if these principles are not made 
retroactive the desired reform will in no sense be secured. A 
gardener might with equal wisdom propose to improve a weedy 
garden simply by preventing the introduction of any more weeds. 
In the view of the reformers it is necessary to remove the old 
weeds as well as to keep out the new. 

In some respects nomenclatural reform will escape in botany 
the difficulties that formerly beset it in zodlogy. We have the 
gratifying assurance that we are not trying an experiment, that 
the plan is not a merely theoretical one, and that its complete 
success will unquestionably be attained in botany as it has already 
been attained in zodlogy. Furthermore, we are able to do ina 
few years, in one stroke, as it were, what zoodlogists, feeling their 
Way over new ground, were many years in accomplishing. All 
botanists dislike changes in names, and the sooner they can be 
properly made the better. 

There are doubtless some botanists who believe that by gen- 
eral agreement any set of names may be made permanent—that, 
for example, an International Congress may decide arbitrarily that 
certain generic or certain specific names are to be considered the 
Proper ones, regardless of any principle. If this were feasible it 
would be an easy solution of the question, but those who have 
confidence in such a solution surely cannot have taken into con- 
sideration the fact that naturalists and other scientists usually 
have very little respect for mere authority and very great respect 


324 


for principle. This sentiment is constantly growing stronger, and 
there is every reason for believing not only that a large percentage 
of botanists would refuse to be influenced by an arbitrary agree- 
ment of this kind, but that the ever-growing younger element 
would within a few years absolutely reject it. 

One of the statements made under the second head in this 
circular deserves special eonsideration. It is as follows: “ While 
the scope of this rule is left to the discretion of writers, it is 
urged that generic nomenclature should not at present depart far 
from that of the three important works, Bentham and Hooker's 
Genera Plantarum, Baillon’s Histoire des Plantes, and Engler and 
Prantl’s Natiirlichen Pflanzenfamilien, from which for some time to 
come our most complete and accurate information as to generic 
limits and affinities is to be derived.” Nothing could better illus- 
trate the present chaos of botanical nomenclature than a compar- 
ison of these three great works. It would be interesting to make 
such a comparison throughout their entire extent, but of course 
space will not permit of this. We may, however, refer to a few 
cases, taken principally at random, to show how widely these 
alleged standard authorities differ in the case of generic names. 


| 


BENTH. & HOOK, ENGL. & PRANTL, | BAILLON, HIST. 
Stellaria L. Alsine L. | Stellaria L. 
Wistaria Nutt. Kraunhia Raf. Wistaria Nutt. 

- Petalostemon Michx. Kuhnistera Lam, | Petalostemon Michx. 
Centrosema DC.* Bradburya Raf. | Centrosema DC.* 
Shepherdia Nutt. Lepargyraea Raf. | Shepherdia Nutt. 
Spergularia Pers. Tissa Adans, ‘Lissa Adans, 
Senebiera. Coronopus Gaertn. | Coronopus Hall. 
Nelumbium Juss. Nelumbo Adans, | Nelumbo Adans. 
Cirsium DC, Cirsium Scop. |  Carduus L. 
Echinospermum Sw. Lappula Moench. | Lappula Moench. 


Any one who has the patience to carry out such a comparison 
to a much greater extent will readily see how little effort has been 
made by the great systematists named to arrive at the original 
names of genera, and so long as authorities are privileged to adopt 


* Bradburya Raf. is included by Benth. & Hook. and Baillon in a list of “ genera 
dubia.” The plants in question are placed under Centrosema DC. 

It will be seen that all but one of these names cited from Engler and Prantl ac- 
cord with those accepted by the Botanical Club’s committee. 


325 


the names that best please them, there never will be any end to 
these wide variations. 

Here again, as in the more general case above mentioned, it 
seems to me that the present circular is based wholly upon senti- 
ment. The time has gone by when it was regarded as the im- 
portant consideration to give special credit to the person who 
becomes the author of a name. It is no longer a question of 
credit, but a question of practical utility. The namer of a plant 
has done nothing more than his duty and while his name should 
of course stand as the author of that name, botanists are not called 
upon to violate the rules of nomenclature for the sole purpose of 
doing him some special honor. The argument that the more im- 
portant service is that of fixing a species under its proper genus, 
and that therefore the binomial combination should bear the name 
of the one who established it falls to the ground, and it becomes 
of far greater importance that the original namer of the species, or 
even of the variety, if there be such, be accredited with that name, 
no matter how many vicissitudes it may subsequently undergo. 
The rule that the namer of the combination may append his nam® 
for the author of the combination, although the last term of that 
combination may have been named long before by another, seems 
to be very vicious from a number of points of view. In the first 
place, if the question of justice were worth considering, it would 
certainly be a gross injustice to the original namer for another 
botanist to usurp his rights and take credit for his name; but this 
is not the chief objection. When I see the names Nuphar kal- 
miana Pursh and Nuphar kalmianum Ait., I at once assume that 
the last name refers to a different plant from the first and that the 
words “non Pursh” or “ not of Pursh” are understood; and when 
I see written Spergularia media Presl. var. macrocarpa Gray in one 
book and Lepigonum medium Fries var. macrocarpum Wats., | am 
entirely at a loss who the true author for the variety macrocarpa 
is. What seems to be most needed is some clue to the history of 
these names, and the particular name should always bear the au- 
thority of the one who first wrote it. 

But I would like to say here that this whole matter of quoting 
authority is one of the worst evils of botanical writing. Any one 
who has the least respect for style must be infinitely annoyed by 


396 


the necessity of tacking on one or two abbreviations at the end of 
a name in order to give his reader an idea of what he is talking 
about. It makes an ugly cacophony that should not be tolerated 
any longer than is absolutely necessary. Now, as I understand 
it, one of the chief objects of this whole movement is, not to lum- 
ber up botanical writing with more things of this kind, but to get 
rid at the earliest possible moment of the whole of it. The orni- 
thologists in adopting one set of names for all birds, the same 
name always meaning the same bird, and all agreeing that that 
shall be the case, have already reached the point at which they 
can write popular articles about birds and omit the authority, thus 
lending smoothness and grace as well as clearness to their discus- 
sions. This should be one of the great aims of botany. Botanists 
ought to draw up a list, international in its scope and based ona 
thorough application of the principle of priority, of all the plants 
known to the world, and all agree that this list should henceforth 
and forever be adhered to as the authoritative list of all known 
plants. This once done and subscribed to by all, it would no 
longer be necessary in any mention that botanists wish to make 
of any plant known to science to append the abbreviations of the 
_ various persons who have had to do with naming it. This surely 
would be a consummation devoutly to be wished. Of course, in 
all subsequent names the authority must be given as heretofore, 
and supplementary lists could from time to time be prepared to 
embody the results of current research. 

But it may be said, and is said by some as a matter of fact, 
that advocates of the new rules of nomenclature do not adhere to 
the law of priority, that it has always been necessary to fix a limit 
or earliest date back of which it is not permissible to go. It 
seems superfluous to argue this question because the reasons are 
so thoroughly well known to all, but it may be said in general 
that in going back to Linnaeus, the founder of binominal nomen- 
clature, and to the particular work of his which is regarded as con- 
taining the most complete expression of his law of binomials, we 
are practically going back, as in the case of the individual, to the 
birth or first christening of a genus or species. We may go back 
in language to the time when there were no common nouns and all 
nouns were proper names. We are told that some rude languages 


327 


are still in that condition. Now in ante-Linnaean days there were 
no true names of plants, certainly not systematic names. All 
such names may be regarded as trivial or common names, and 
at the time when Latin was generally written and largely spoken 
they were scarcely more than vernacular names. Moreover, bot- 
anists commenced describing them before they commenced nam- 
ing them, and the binominal nomenclature is a direct descendant 
of a system of describing plants. It resulted from the dropping 
more and more of the adjective terms contained in the character 
until at last the description consisted of only two names, that of 
the genus and that of the species. In later times trinominals 
came more or less into vogue, and as the science advanced it be- 
came apparent that the name had little to do with the description, 
so that although up to this day many or most specific names have 
a greater or less descriptive value, still large numbers of them pus- 
sess no such value whatever, and the combination has become 
simply a name or symbol by which the plant may be known. 
Elementary as these remarks are, it is upon such facts that is 
based the reason for fixing some specific standard for the origin of 
systematic nomenclature, and to all who clearly understand the 
fact this reason is wholly conclusive. 

Since I have commenced the study of fossil plants I have 
found the same difficulty, and although the science scarcely dates 
back of the beginning of the present century the nomenclature is 
in a condition of great confusion. M. R. Zeiller in 1877 encoun-- 
tered this difficulty and expressed himself on the subject in the 
following clear and trenchant language :* 

“The unfortunate confusion that results from these successive 
changes in the name of one and the same object has taken place 
the same and in a much greater degree in other older branches of 
natural history, and to-day the necessity for some remedy is recog- 
nized. The only equitable and rational basis that can be adopted 
is the one that was proposed in 1813 by [A. P.] De Candolle in 
his Théorie elémentaire de la botanique, in the chapter on Phy- 
tography, viz., the maintenance throughout all changes in the 


*R. Zeiller, Ingénieur en chef au corps national des mines. Explication de le 
carte geologique de la France, Tome IV., 2me Partie. Végetaux fossiles du terrain 


houiller, 1879, Pi 5: 


328 


genus of the oldest specific name, or, more generally, the absolute 
principle of the right of priority. In order that the nomenclature 
may be invariable and universally accepted it must rest upon fixed 
principles whose application is not subject in any way to arbitrary 
judgment; thus we should adhere without exception in the case 
of each genus or species as De Candolle had established it, to the 
name first in date, even where this name has been recognized as 
improper and in contradiction with such and such characters of 
the object or group to which it is applied. Generic or specific 
names are in fact only designations and not definitions, and if it is 
admitted that they may be changed because improper, the door is 
opened to arbitrary action, each author interpreting differently the 
propriety or impropriety of a name.” 

I have been engaged for over fourteen years, as time would 
permit, upon the bibliography and synonymy of fossil plants, and 
hope ultimately to publish a complete catalogue of all the names 
that have ever been given to the extinct vegetation of the globe, 
fully exposing the confusion referred to and adhering strictly in 
the final revision to the law of priority. 

Inasmuch as the representative character of the Botanical Club 
of the American Association has been called in question, it is of 
interest to know that some of the opponents of the Rochester 
movement formerly thought otherwise, as is seen by the following 
extract from the Proceedings of the Washington Botanical Club 
of May 7, 1892, which were published in the Botanical Gazette 
for June, 1892. These resolutions were duly signed by each mem- 
ber of the committee and are preserved in the minutes of the Club. 

“Ata meeting of the Botanical Club of Washington, held 
April 23, 1892, a committee was appointed to consider and report 
upon the questions of botanical congress and nomenclature. Ata 
special meeting called May 7, this committee presented the fol- 
lowing report, which was unanimously adopted by the Club: 


* Your committee, appointed to consider the questions of a botanical congress 
and botanical nou:enclature, held a meeting on the second of May and prepared the 
following resolutions : 

“Resolved, That, while favoring the final settlement of disputed questions by 
means of an international congress, we do not regard the present as an opportune 
time, but we recommend the reference of the question of plant nomenclature first to a 
representative body of American botanists. 


329 


“We suggest the consideration, by such body, of the following questions, among 
others: the law of priority; an initial date for genera; an initial date for species ; 
the principle once a synonym always a synonym; what constitutes publication; the 
form of tribal and ordinal names; the method of citing authorities; capitalization. 

“We recognize the Botanical Club of the A. A. A, S. asa representative body 
of American botanists and commend to that body for discussion and disposal the sub. 
ject of nomenclature as set forth in these resolutions,” 

Respectfully submitted, 
LesTer F, Warp, 
GEO, VASEY, 
F. Hi. Know tron, 
B. T. GALLoway, 
ERWIN F. SMITH, 
Gro, B. SupworTtH, 
FREDERICK V. COVILLE, 
Committee. 


Missouri Botanical Garden,” 


The attention of botanists is called to the facilities afforded for 
research at the Missouri Botanical Garden. In establishing and 
endowing the Garden, its founder, Henry Shaw, desired not only 
to afford the general public pleasure and information concerning 
decorative plants and their best use, and to provide for beginners 
the means of obtaining good training in botany and horticulture, 
but also to provide facilities for advanced research in botany and 
cognate sciences. For this purpose additions are being made 
constantly to the number of species cultivated in the grounds and 
planthouses and to the library and herbarium, and, as rapidly as 
it can be utilized, it is proposed to secure apparatus for work in 
vegetable physiology, etc., the policy being to secure a good gen- 
€ral equipment in all lines of pure and applied botany, and to 
make this equipment as complete as possible for any special sub- 
ject on which original work is undertaken by competent students. 

A very large number of species, both native and exotic, 
and of horticulturists’ varieties, are cultivated in the Garden 
and Arboretum and the adjoining park, and the native flora 
easily accessible from St. Louis is large and varied. The her- 

[* This article has recently been issued as a leaflet. We take pleasure in placing 
it more permanently on record and in calling the attention of botanists to the valuable 
facilities for research afforded.—Ens. ] 


330 


barium, which includes nearly 250,000 specimens, is fairly 
representative of the vegetable life of Europe and the United 
States, and also contains a great many specimens from less 
accessible regions. It is especially rich in material illustra- 
tive of Cuscuta, Quercus, Coniferae, Vitis, Juncus, Agave, Yucca, 
Sagittaria, Epilobium, Rumex, Rhamnaceae and other groups 
monographed by the late Dr. Engelmann or by attachés of the 
Garden. The herbarium is supplemented by a large collection of 
woods, including veneer transparencies and slides for the micro- 
scope. The library, containing about 8,000 volumes and 10,000 
pamphlets, includes most of the standard periodicals and proceed- 
ings of learned bodies, a good collection of morphological and 
physiological works, nearly 500 carefully selected botanical vol- 
umes published before the period of Linnaeus, an unusually large 
number of monographs of groups of cryptogams and flowering 
plants, and the entire manuscript notes and sketches representing 
the painstaking work of Engelmann. 

The great variety of living plants represented in the Garden, 
and the large herbarium, including the collections of Bernhardi 
and Engelmann, render the Garden facilities exceptionally good 
for research in systematic botany, in which direction the library 
also is especially strong. The living collections and library like- 
wise afford unusual opportunity for morphological, anatomical and 
physiological studies, while the planthouse facilities for experi- 
mental work are steadily increasing. The E. Lewis Sturtevant 
Prelinnean library, in connection with the opportunity afforded 
for the cultivation of vegetables and other useful plants, is favor- 
able also for the study of cultivated plants and the modifications 
they have undergone. 

These facilities are freely placed at the disposal of professors 
of botany and other persons competent to carry on research work. 
of value in botany or horticulture, subject only to such simple 
restrictions as are necessary to protect the property of the Garden 
from injury or loss. Persons who wish to make use of them are 
invited to correspond with the undersigned, outlining with as 
much detail as possible the work they desire to do at the Garden, 
and giving timely notice so that provision may be made for the 
study of special subjects. Those who have not published the re- 


331 


sults of original work are requested to state their preparation for 
’ the investigation they propose to undertake. 

Under the rules of Washington University, persons entitled to 
candidacy in that institution for the Master's or Doctor’s degree 
may elect botanical research work asa principal study for such 
degrees, if they can devote the requisite time to resident study. 

WILLIAM TRELEASE, 


Director. 
St. Louts, Mo., May 8, 1895. 


Botanical Notes. 


Lhe Pignuts. There is some question as to the exact distribu- 
tion of the Common Pignut (Carya porcina or Hicoria glabra) and 
the related Carya or Hicoria microcarpa, and the undersigned will 
be grateful for herbarium specimens and especially nuts with their 
husks, representing both. In the recently published seventh vol- 
ume of Professor Sargent’s Silvaa, the range of g/aéra is given as 
southern Maine to southern Ontario, through southern Michigan 
to southeastern Nebraska, southward to the shores of the Indian 
River and Peace Creek in Florida, and to southern Alabama and 
Mississippi, through Missouri and Arkansas to eastern Kansas and 
the Indian Territory, and to the valley of the Nueces River in 
Texas. H. microcarpa (treated in the Silva as a variety of g/aéra, 
under the varietal name odorata) is said to occur in eastern Mass- 
achusetts, Connecticut, eastern and central New York, eastern 
Pennsylvania, Delaware, the District of Columbia, central Michi- 


gan, southern Indiana and Illinois and Missouri. 
WILLIAM TRELEASE. 


Reviews, 


A Monograph of the Mycetozoa. Arthur Lister, F. L. S. 
London. Printed by order of the Trustees of the British Museum. 
Rostafinski’s monograph of the Mycetozoa appeared in 1875, 
and with the appendix described about 230 species. Massee’s 
“Monograph of the Myxogastres” was published in 1892; it 


332 


enumetates 430 species. Lister’s volume comes to us dated 1894; 
it furnishes descriptions of 174 species “taken from specimens I 
have personally examined,” and a list is appended at the end of 
each genus of “species not met with in the quoted collections ;” 
this comprises about 100 species. 

The basis of work in all three of these monographs is the 
same, viz.: the types of the species of Rostafinski’s monograph in the 
collections at Kew, in the British Museum and at Strassburg. Of 
course the continued accession of specimens to these herbaria has 
augmented the material of each succeeding author, and important 
contributions appear to have been made to Mr. Lister's resources 
by several parties on this side the water. 

Rostafinski, by reason of his original method, which involved a 
complete reorganization of the classification, necessarily reduced 
numerous species of the old writers to the condition of synonyms. 
Massee in general accepted the species as established by Rosta- 
finski, while at the same time making numerous additions to this 
list from descriptions and material derived from American sources 
and elsewhere. Lister, however, while professing to follow in the 
main the arrangement of orders and genera given by Ros- 
tafinski, opens up afresh the question of species and proceeds to 
make a thorough and complete revision of the works of his prede- 
cessors. Upwards of ezghty of the species of Rostafinski’s mono- 
graph have disappeared in synonomy, united with other species, 
becoming reduced two, three and four or more into one. This 
involves at the same time the abolition of five of the Rostafinskian 
genera. 

The manner in which species and genera are disposed of is 
sometimes remarkable. For example, it is suggested that the 
single species of the genus Crateriachea is but a variety of P/ysa- 
rum cineream Batsch, and “a careful examination of the type 
specimen of Heterodictyon mirabile Rost. leads to the conclusion 
that it is a form of Dictydium umbilicatum Schrad.” On page 
89 eight Rostafinskian species are excluded from the genus C/on- 
drioderma, four of them being unceremoniously dumped into 
Inichamphora pezizoidea Jungh; the single species of this genus 
is composed of five species of Rostafinski and one of Massee. 

Of the numerous American species of Berkeley and Curtis, all 


333 


but two or three subside into synonomy. Peck and Rex come off 
a little better, a paltry half dozen or so being graciously allowed 
to remain to each. It is George Massee, however, who has to 
bear the brunt of the crush and to suffer the most acutely. Out 
of upwards of fifty new species named and described by him, only 
a single one is allowed to live, with the exception of two or three 
which are permitted to linger along until Lister can lay his hands 
on the types. This, too, it will be observed, is in addition to the 
Rostafinskian squeeze, the sum of which leaves Massee’s volume 
in a sad state of collapse. ‘ 

If the details of Lister’s book are accurate and reliable, and the 
work is to be accepted as authoritative, then Massee’s volume is a 
tissue of mistakes and blunders and a monument of ignorance— 
and vice versa. There is scarcely an agreement in spore measure- 
ments under any of the species, and the discrepancies are often im- 
mense. For example, the spores of Lamproderma irideum Massee 
are given in one volume as II-I5 y, in the other as 6.5-8 ». The 
measurements of sporangia exhibit the same diversity. Species 
appear under one genus in the one volume and under an entirely 
different genus in the other. Didymium flavicomum of Massee be- 
comes Physarum Berkeleyi of Lister. The difference in treatment 
of the elegant genus, 777chia, by the two is something appalling ; 
the number of species described by Massee is c/zrty, while Lister 
recognizes but ¢ez. Massee’s single genus, Heterotrichia, the only 
one he ventured to establish, is incorporated in Arcyria ferruginea 
Sauter. Two of Massee’s species of Lycogala are excluded from 
the Mycetozoa! And Zudulina spumarioidea Cooke & Massee is 
declared to be nothing but the common fungus Sepedonium chrys- 
ospermum k.! The number of species that are occasionally got 
together sometimes rivals the synonomy of Rostafinski, and pos- 
sibly a righteous retribution has now fallely on him for having 
made such havoc with the species of the old writers. For ex- 
ample, in Physarum compressum A. & S.are dumped four of Rosta- 
finski’s species, two of Berkeley’s, three of Massee’s and one each 
belonging to Balfour, Phillips and McBride. Olgonema nitens 
Libert is made a dumping place for eight different species, and he 
seems to have seriously considered whether he shouldn’t dump the 
whole into Ziichia affinis De B. 


834 


Mr. Lister has made a few attempts at novelties himself, which 
in the main are unfortunate. It is doubtful if Physarum murinum 
Lister is anything different from Fhysarum puichripes Peck, as he 
has it. Physarum calidris Lister, on his own showing should be 
called Physarum pusillum B. & C., and the pretext for not doing 
so is perfectly flimsy. At any rate Physarum nodulosum Cke. & 
Balf. has priority; it was published as Badhamia nodulosa in the 
Journal of Mycology in 1889. If Hemiarcyria stipitata Mass. is 
to be absorbed in H. c/avaza Pers., with more reason should H. 
intorta Lister be included in the same species. The definition of 
his new order Margaritaceae is illogical; in fact, the order is un- 
necessary and the genera may easily be distributed elsewhere. 
The theory of the tubules in Lycogala is questionable; but if true 
the tubules are not a capillitium traversing the zzterior of the spo- 
rangia, but only “air-spaces’’ between them, in which case Lycogala 
goes to Reticulariaceae. The placing of Chondrioderma and Dia- 
chaea in Physaraceae is awkward indeed, and willreceive the approval 
ofnoone. Hemiarcyria isa good Friesian designation much older 
than Hemitrichia. Something is said in the introduction concern- 
ing the “laws of botanical nomenclature,” but it will be seen 
that the nomenclature remains a purely personal one, and that re- 
vision of the generic and specific names in the Myxomycetes is 
still a thing greatly needed. 

We are of the opinion that Mr. Lister has shown too little re- 
spect for the labors of his predecessors and has exhibited a great 
want of consideration for the views and opinions of his contempo- 
raries. We have a lurking suspicion that he is influenced by 
some strong personal bias greatly to the discredit of many of his 
statements ; possibly it is only a yearning for notoriety. And we 
are inclined to look upon him as a narrow specialist, disqualified, 
by reason of his limited studies, for forming a proper judgment 
concerning genera and species. AF Me: 

Untersuchungen uber die Starkekorner. By Dr. Arthur Meyer: 
With 99 cuts in the text and 9g plates. Jena, 1895. 

This treatise will no doubt be welcomed by all scientific 
botanists. The author has made an earnest, conscientious effort 
to clear up some of the mysteries concerning the life history of 

the individual starch-grain. He admits that some of his conclu- 


* 


335 


sions in regard to structure, imbibition, stratification and growth 
are yet theoretical. Our knowledge concerning the chemistry of 
starch is especially imperfect. It would be impossible to attempt 
to give a full summary of the author's investigations and results. 
The following are perhaps the most important conclusions: 

1. The starch-grain consists of amylose, which separates into 
a-amylose and #-amylose, amylodextrin, dextrin, isomaltose, and 
maltose. Of these substances amylose is perhaps the true starch- 
substance ; the others are the result of processes of inversion. 

2. Starch-grains are sphaero-crystals of amylose and amylo- 
dextrin. This is evident from their optical behavior. 

3. Amylose is split up by the starch-ferment diastase as fol- 
lows: with the aid of water, the amylose molecule is converted 
into two or more molecules of amylodextrin; this is converted 
into dextrin and isomaltose ; a further process of splitting up con- 
verts dextrin into maltose, while isomaltose may also revert into 
maltose. 

4. The starch-grain is porous. The pores are scarcely per- 
ceptible with the highest magnifying powers. The author makes 
a special attack upon Biitschli’s « Wabenstructur” theory. Among 
other things he says, ‘Es ist héchst interessant zu sehen wie ein 
So vortrefflicher Beobachter (Biitschli) die Schaum und Waben- 
structur mit welcher er sich eingehend beschaftigt, in die Ob- 
jecte hinsieht,” which, I think, expresses it rightly. 

The following are some of the more important conclusions in 
regard to the biology of the starch-grain. 

1. Starch-grains occur exclusively in the chromatophores. 
They may occur in any kind of chromatophore. The starch-grain 
of angiosperms originates and grows from the beginning until its 
final solution within a chromatophore. 

2. Every starch-grain is entirely enclosed by the substance of 
the chromatophore. The author points out the difficulty of de- 
monstrating the presence of the chromatophore substance. 

3. The form of the starch-grain is more or less dependent 
upon the form of the chromatophore. The chromatophore sub- 
Stance does not always form an even layer over the starch-grain. 
The thickest portion of the chromatophore always lies in contact 
with the thickest stratifications of the starch-grain. 


336 


4. Several starch-grains may develop in one and the same 
chromatophore, 

3. Solution of the starch-grain within the cell is due to the 
ferment diastase. 

6. Stratification of starch-grains is still not well understood, 
it is perhaps due to a periodicity of growth (deposition of new 
layers), and to the action of diastase. 

Part IV. comprises monographs on the biology of the starch- 
grains of Adoxa moschatellina, Hordeum distichum, Dieffenbachia 
Seguine, Pellionia Daveauana, Hyacinthus orientalis, Oxalis Origiest 
and Cyrtodeira cupreata. 

The treatise, which comprises 318 large octavo pages, is written 
in clear scientific style. Some of the cuts are poor. Taken asa 
whole, it is certainly the standard work on starch. It shows a 
great advance made since Nageli’s memorable communications on 
the same subject. ALBERT SCHNEIDER. 


Catalogue of Ohio Plants. WW. A. Kellerman and William C. 
Werner (Geology of Ohio, 7: Part 2, 56-406. 1895). 

Since: the publication of Dr. Beardslee’s “Catalogue of the 
Plants of Ohio” in 1874, a great amount of botanical exploration 
and critical study of the flora of the State has been accomplished, 
no less than 110 published papers and references to the plants of 
the area, during that period, being cited in the work here noticed. 

It was therefore highly desirable that these records should be 
' brought together and incorporated with the unpublished results of 
the recent work of Professor Kellerman, his students and associates 
in the region. The duty has been discharged in a thorough and 
painstaking manner, as evidenced by the fine volume which lies 
before us, and we tender its authors the cordial congratulations of 
American botanists upon its completion. 

The chapter on bibliography cites the titles of 132 papers and 
references, all but, two or three of which have been examined. 
The arrangement of the families is that of Engler and Prantl, but 
in a reverse sequence, beginning with the Compositae and ending 
with the Myxomycetes. The nomenclature is based on the prin- 

_ciples adopted by the Botanical Club of the American Association 
for the Advancement of Science. N. L. B. 


337 


Index to recent Literature relating to American Botany. 


Alwood, W.B. Ripe-rot or Bitter-rot of Apples. Bull. Va. Exp. 
Sta. 4o: 56-82. f/. 2. My, 1894. 

Atkinson, G. F. Damping Off. Bull. N. Y. (Cornell) Exp. Sta. 
94: 230-272. f. 55. My. 1895. 

Treats at length of Artotragus Debaryanus (Hesse), A. intermedius (De Bary), 

Completoria complens Lohde and Volutella leucotricha Atkin. 

Baker, J.G. LZupatorium clibadoides, Aspilia Glaziovi, Senecio arctit- 
folius. Kew Bull. ror: 105, 106. My. 1895. 

New species from Brazil. 

Britton, N. L., and Vail, A. M. An Enumeration of the Plants col- 
lected by M. E. Penard in Colorado during the Summer of 1892. 
Bull. Herb. Boiss. 3: 197-221. My. 1895. 

Brown, N. E. Cynanchum formosum. Kew Bull, tor: 112. My. 
1895. 

Native of Peru and Ecuador. 

Burt, E. A. A North American Azthurus. Mem. Bost. Soc. Nat. 
Hist. 3: 487-505. fl. 49, 50. 1894. 

A. borealis, from East Galway, N. Y., described as new. 

Card. F. W. Garden Herbariums. Gard. and For. 8: 242. 149 Je. 
1895. 

Christ, H. Fougéres nouvelles ou peu connus. Bull. Soc. Bot. Belg. 


33: part 2, 92, 95. 27 Je. 1895. . 
Gymnogramme Eggersii from Cuba (Eggers, 4882a), and Alsophila monosticha 


from British Guiana, 
Christ, H. Un cas d’androgynie dans le genre Pinus. 


Bot. Belg. 33: part 2, 88-92. 27 Je. 1895. 
P. Cubensis Griseb. from Bluffton, S. C. 


Clark, H. L. Studies in Plant Development. Folio, Chicago, 1895. 
Blanks for recording studies in comparative morphology. 

Corbett, L. C. Squashes. Bull. S. Dak. Exp. Sta. 42: 77-92. Fa: 
Fb. 1895. 


Demonstrates the desirability of artificial pollination in fiel@ culture. 

Coulter, J. M. The Botanical Work of the Government. Bot. Gaz. 
20: 264-268. 17 Je. 1895. 

Coulter, J. M., and Rose, J. N. Musineon of Rafinesque. Bot. Gaz. 
20: 258-260. 17 Je. 1895. 


Enumeration of the species of Musineon; M. alpinum is described as new. 


Bull. Soc. 


308 


Coville, F. V. Directions for collecting Specimens and Information 
illustrating Aboriginal Uses of Plants. Bull. U. S. Nat. Mus. 39: 
1-8. 1885. 


Dietel, P. Uber Quellungserscheinungen an den Telenutosporenstielen 
von Uredineen. Pringsheim’s Jahrbiicher, 26: 49-81. A/. 4. 1894. 
Refers to a number of North American species. 

Eggleston, W. W. Astragalus Blaket. Bot. Gaz. 20: 271. 17 
Je. 1895. 

A new species from New England. 

Greene, E. L. Novitates occidentales.—XIV. Eryth. 3: 98-101. 1 
Je. 1805. 

Describes new spacies in Astragalus, Psoralea, Streptanthus, Erigeron, Micro- 
servis, Hieracium and Bolelia. 

Greene, E. L. Observations on the Compositae—IX. Eryth. 3: 89-98. 
I Je. 1895. 

Establishes the genus Arintonia for Solidago discoidea; enumerates the species of 

Chondrophora and Chrysothamnus; describes C. elegens as new. 

Greene, E. L. Phytographic Notes and Amendments.—II. Eryth. 3: 
Io2-105. 1 Je. 1895. 

Describes new species in Lithophragma and Gilia. 

Harrison, W.H. The Poison Ivy. Gard. and For. 8: 268. 3 ji. 
1895. 

Harshberger, J. W. When is Rhus Toxicodendron most active ? 
Gard. & For. 8: 239. 12 Je. 1895. 


Hillman, F.H. Early Flora of the Truckee Valley. Bull. Nevada 
Exp. Sta. 24: pp. 96. N. 1894. 
Account of the commoner flowering plants occurring within the Truckee Valley by 

midsummer. 

Hooker, J. D. Peraphylium ramosissinum, Curt. Bot. Mag. f/. 
7420. Je. 1895. 


Hooker, J. D, Ribes bracteosum. Curt. Bot. Mag. f/. 7419. ji. 
1895. 


Jack, J.G. Hybrid Birches. Gard. and For. 8: 243, 244. f. JO. 19 
Je. 1895. 


Illustrations are given of Betula pumila X< lenta, 


MacDougal, D. T. Experimental Plant Physiology. 8vo, pp- 88. 
Jigs. New York, 1895. 


Meehan, T. Ocxadis violacea, Meehan’s Month. 5: 121-122. f/ 
7. Jy. 1895. 


339 


Nichols, M. A. Abnormal Fruiting of Vaucheria, Bot. Gaz. 20: 
269-270. pl. 27. 17 Je. 1895. 

Putnam, B. L. Poison Ivy. Gard. and For. 8: 249. 19 Je. 1895. 

Reiche, K. Die Vegetations-Verhiltnisse am Unterlaufe des Rio 
Maule, Chile. Engler’s Bot. Jahrb. 21: 1-54. 
General account of the flora and enumeration of 664 species. 

Renauld, F., and Cardot, J. Musci exotici novi vel minus cogniti,— 
VI. Bull. Soc. Bot. Belg. 33: Part 2, rog. 1895. 


Includes descriptions of Calymperes Nicaraguense from Nicaragua, Prionodon 
Haitensis from Haiti, and Hypnum Baréeyi from Bolivia. 


Robinson, B. L. The Nomenclature Question. Bot. Gaz. 20: 261- 
263. 17 Je. 1895. 

Rose, J. N. Notes upon Corylus rostrata and C. Californica, Gard. 
and For. 8: 263. 3 Jl. 1895. 

Sargent, C.S., Editor. The Live Oak at Drayton Manor. Gard. & 
For. 8: 233. f. 35. 12 Je. 1895. 


Illustrating Quercus Virginiana in South Carolina. 

Sargent, C. S., Editor. The New York Botanical Garden. Gard. & 
For. 8: 261. 3 Jl. 1895. 

Sargent, C.S., Editor. The Persimmon. Gard. & For. 8: 262. 
J. 38. 3 Ji. 1895. 

Stapf, O. Zrlouchina metodon n. sp. Kew Bull. 100: 104. Ap. 1895. 
Native of Brazil. 

Stewart, F.C. Witches’ Brooms on Cherry trees. Gard. and For. 


8: 269. 3 Jl. 1895. 


Note concerning the occurrence of Axoascus Cerast on Prunus Avium and P. 

Cerasus in America. 

Strasburger, E. The Development of Botany in Germany during the 
Nineteenth Century. Bot. Gaz. 20: 193-204. 20 My.; 249-257. 
17 Je. 1895. 

Ten Bosch, J. Does Poison Ivy discriminate? Gard. & For. 8: 
239. 12 Je. 1895. 

Tricker, W. Mexican Water-lilies. Gard. & For. 8: 237. 12 Je. 
1895. 

Waugh, F. A. Cnotheras on the Prairies. Gard. & For. 8: 253. 
26 Je. 1895. 

Waugh, F. A. Phlox divaricata on the Plains. 
239. 12 Je. 1895. 


Gard. & For. 8: 


340 


Wetherill, H. E. List of Plants obtained on the Peary Auxiliary Ex- 
pedition of 1894. Bull. Geogr. Club, Philadelphia, 5: [reprint pp. 


Io]. 

Wood, A. H. The Flow of Maple Sap. Bull. N. H. Exp. Sta. 24: 
pp. 8. Feb. 1895. 

Wright, F. W. Oregonian forms of Umbellularia. Eryth. 3: 97- 
1 Je. 1895. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 232. 


eR Stree he es SS 


ese Tr aad a) shigicesan. fee a 
wings bak Comid* 
Oe Laat 


NEW AND RARE DESMIDS. 


acQ? O°9°Ooq) ye “poho: OO 
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209 02020209 620° +0°O? OQ 
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NEW AND RARE DESMIDS. 


Contributions from the Herbarium of Columbia College. 


No. 4. 


[ The numbers omitted from this list are out of print. ] 


VotumE I. 
A List of Plants Collected by Miss Mary B. Croft at San Diego, Texas. By 
N.-L, Britton ond TH. BH. Rusby (1887), op sk ee 25 cents. 
New or Noteworthy North American Phanerogams. By N. L. Britton 
CESBS eS re ee ee es . . 25 cents, 
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A List of Plants Collected by Dr. E. A. Mearns at Fort Verde and in the 
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Preliminary Notes on the North American Species of the Genus 7issa, 
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a el Oe ee 


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eat Pe ei. dean eat Weeks Vier MR RG, SOARES rset 


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AUGUST, 1895. 


_ BULLETIN 


OF THE - 


*,. 


THE TORREY BOTANICAL CLUB. 


OFFICERS FOR 1895. 


President, 
HON. ADDISON BROWN. 


Vice Presiaent:, 


T. F. ALLEN, M. D. L. H. LIGHTHIPE. 
Recording Secretary, Corresponding Secretary, 
HENRY H. RUSBY, M. D., JOHN K. SMALL, 
College of Pharmacy, New York City. Columbia College, New York City. 
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__ EFFIE A. SOUTHWORTH, Rev. L. H. LIGHTHIPE. 
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BULLETIN 


TORREY BOTANICAL CLUB. 


Daniel Cady Eaton, 
1834-1895. 


By WILLIAM ALBERT SETCHELL. 
(With portrait.) 


The death of Prof. Daniel C. Eaton, at his home, in New 
Haven, on June 30, 1895, removes from among us the last link in- 
timately connecting the systematic botany of the present with 
that of the past. During the first half of the present century the 
most influential writer upon, and teacher of, botany in this country 
was Amos Eaton, Senior Professor of the Rennselaer Institute at 
Troy, N. Y., and grandfather of the subject of our sketch. His 
Manual was the inspiration and guide of our earlier botanists, and 
continued to be until supplanted by the works of Torrey and 
Gray. The botanists earlier than and contemporaneous with 
' Amos Eaton were writers of Floras, but Amos Eaton, himself, 
was a teacher of great ability and awakened, even among the 
members of the New York Legislature, so it is said, a deep and 
widespread interest in natural history. From him John Torrey 
learned the rudiments of botany, and was able to broaden and 
deepen the knowledge and interest in botanical things of Asa 
Gray, whose first knowledge of the subject came from Amos 
Eaton’s text-books, Of Gray’s influence upon the botany, not 
only of this country, but of the world, it is needless to speak, so 
present is it with all of us. Of Gray’s associates and pupils two 
were more intimately thrown together than any others, and these 


342 


were Daniel C. Eaton and Sereno Watson. With them we may 
say, perhaps, that the direct line of succession ends. Others 
there are, no less illustrious, but they have departed more widely 
than these two from the direct line of work, and the mantle, while 
it has fallen upon most worthy shoulders, yet lacks the accumu- 
lated traditions which were a precious heritage in themselves. 

Daniel C. Eaton, as he was called to distinguish him from his 
cousin, D. Cady Eaton, was the son of Amos B. Eaton and Eliza- 
beth Selden, and was born at Fort Gratiot, Mich.,on February 
12, 1834. His father was an officer in the Regular Army of the 
United States, had served with distinction in the Seminole and 
Mexican Wars, had risen to the office of Brigadier General, and 
was entrusted with the very important work of supervising the 
Commissary Department as Quartermaster General during the 
Civil War. Although he never had any special scientific educa- 
tion, as had a sister and a brother who became teachers in these 
lines, yet he possessed a keen interest in natural history and 
helped to direct his son’s education whenever the roving life of an 
army officer allowed him to do so. Mrs. General Eaton was a 
sister of Samuel K. and Henry R. Sheldon, two distinguished 
jurists of Rochester, N. Y. 

Daniel C. Eaton’s early training was obtained in different places. 
For a time the family resided in New York City, and during the 
Mexican War the mother and children remained at Rochester. 
Later he attended the Rennselaer Institute for a short time and 
General Russell’s Military School in New Haven, while prepat- 
ing for college. His final preparation was made with a private 
tutor, and he entered Yale in the autumn of 1853. 

During his college course he distinguished himself particularly 
in Latin, and he retained during the rest of his life an especial 
fondness for this language and its literature. But his chief inter- 
est lay in botanical pursuits, and this distinguishing trait is 4 
prominent feature in the recollections of his classmates with 
whom he was a great favorite. He never had a doubt as to his 
future vocation, and his ambition was to become the professor of 
his favorite subject in his alma mater, the institution where his 
grandfather had received a part of his own botanical and chemi- 
cal education under Professors Silliman and Ives. 


545 


a 


During his undergraduate days he derived much pleasure and 
profit from his correspondence with Torrey, Gray and. Sullivant, 
to whom he sent his specimens and to whom he applied for 
counsel. His herbarum still contains many of these specimens 
with notes from these distinguished authorities. Such was his 
advancement even then that in his junior year he published a short 
paper in “ Silliman’s Journal” on three new ferns from California, 
his first contribution to science upona group to which he devoted 
later his chief attention. 

He obtained the degree of B. A. from Yale College in 1857, 
and spent the three succeeding years in the Lawrence Scientific 
School of Harvard University in special botanical studies, under 
the direction of Professor Asa Gray. He devoted himself partic- 
ularly to the study of the ferns and produced several papers. He 
enumerated and described the new species of ferns from the col- 
lections made by Charles Wright of the Rodgers Exploring Expe- 
dition in Cuba and Japan, for Torrey’s Flora of the Mexican 
Boundary Survey and for Chapman’s Flora of the Southern United 
States. Finally, in 1860, he presented to the faculty of Harvard 
University, as his thesis for the purpose of obtaining the degree 
of Bachelor of Science, his “ Filices Wrightianae et Fendlerianae.” 

During the Civil War his botanical studies were interrupted and 
he held the position of clerk and inspector of stores in the Com- 
missary Department of the United States Army in New York City. 
He, however, had an opportunity of associating more intimately 
with Professor Torrey and of increasing his store of botanical 
tradition. 

After the war, in 1864, he was elected to the chair of botany 
in Yale College, which had been established by some of his friends, 
and his duties were assigned to the Sheffield Scientific School, 
with which he continued to be principally identified. A few years 
later he was appointed University Professor, and continued to give 
instruction in both departments until the end of his life. He took 
up his residence in New Haven in the fall of 1864, and in 1866 he 
married Caroline, daughter of Treadwell Ketcham, of New Haven. 
Mrs. Eaton, a son, and a daughter survive him. 

Professor Eaton made two trips to Europe, one early in 1866 
and another in 1887. As the later trip was taken for the benefit 


344 


3 


of his health, he visited little and indulged in practically no botan- 
ical study. In 1866 he visited the botanists and botanical collec- 
tions at Florence, at Geneva, at Paris, and in London at the 
British Museum and the gardens at Kew. At the latter place he 
spent two weeks in the careful study of ferns, working assiduously 
among the collections, every courtesy and every assistance being 
rendered him by Sir Joseph Hooker, then Director of the Royal 
Gardens. 

He took many collecting trips, especially into New Jersey, 
with Gray, Canby and others, to the White Mountains of New 
Hampshire, and a portion of the last summer of his life was spent 
at Shelburne, N. H., with Professors Farlow and Penhallow in col- 
lecting the Sphagna of that region. During the summer of 1869 
he spent a month botanizing among the mountains of Utah, as the 
guest of Clarence King, who was in charge of the geological sur- 
veys of the fortieth parallel. 

Even in his undergraduate life his preference for the cryptoga- 
mous plants was marked, and he will be remembered chiefly for 
the work he did among them. His knowledge of the phaenoga- 
mous species, nevertheless, was very extensive and exact. He 
preferred, however, to entrust the work of publishing upon these 
plants to Professor Gray. His attainments in this line are shown, 
however, in the masterly way in which he has treated the Com- 
positae of King’s Expedition, and the additional material and other 
aid afforded by him in the preparation of the whole of the botany 
of that expedition is gracefully and effectively set forth in Wat- 

son’s general introduction. Besides this work his published ob- 
' servations on the flowering plants are few, although he retained 
much interest in them even until the last. 

His chief work from the beginning (1856) until about 1883 
lay among the ferns and their immediate allies. His first several 
papers were devoted to enumerations of various collections of 
ferns and descriptions of new species. He contributed the ac- 
counts of the Vascular Cryptogams to Torrey’s Botany of the 
United States and Mexican Boundary Survey, to Chapman’s Flora 
of the Southern United States, to Gray’s Manual of Botany of 
the Northern United States (both to the 5th and the 6th editions), 
to Gray’s Field, Forest and Garden Botany, to the Botany of 


345 


King’s Survey, to that of Wheeler’s Survey, and to the Botany of 
California. 

In 1873 he began a series of notices of “ New or Little 
Known Ferns of the United States,” directly leading up to his 
great work on ‘‘ The Ferns of North Amcrica.” This classic and 
and well-known work consists of two royal octavo volumes con- 
taining 683 pages and illustrated by 81 colored plates, after draw- 
ings by Emerton and Faxon, representing 149 species, all that 
were known at the time from North America north of Mexico. 
They were issued in 1879 and 1880, but the notes on ferns in the 
BULLETIN of the Torrey Botanical Club were continued actively 
until 1883, after which there were very few articles from his pen on 
this subject. 

He was for many years much interested in Algz and spent a 


- considerable time upon them. He spent a portion of one summer 


with the United States Fish Commission at Noank, engaged in 
the study of this group. He published very little upon them, 
however, limiting himself to a list of Eastport Algz, another of 
those collected by Edward Palmer in Florida and the Bahama 
Islands, and directions for mounting and preserving specimens. 
He was associated with Professor W. G. Farlow and Dr. C. E. 
Anderson in issuing the Algae Americae-Borealis Exsiccatae. 

The later years of Professor Eaton’s life were devoted to the 
Study of the mosses and liverworts. His interest in these plants 
Was well advanced even in his undergraduate days. He collected 
much and carried on a considerable correspondence with W. S. 
Sullivant in regard to the determination of his specimens. His 
knowledge of the New Haven Moss and Hepatic Flora was very — 
extensive, and he contributed the account of these plants to the 
Catalogue of Plants growing within 30 miles of New Haven, pub- 
Published by the Berzelius Society of the Sheffield Scientific 
School in 1878. His other publications on these groups are con- 
fined to lists of Patagonian species and a few notes on new or rare 
Species of the United States. 

He spent a great deal of time in the study of the Hawaiian 
Species, both of Mosses and Hepatics, and had almost completed 
the determination of the species of the various collections in his 
Possession. He spoke to the writer only a few weeks before his 


346 


death of issuing a list with notes and descriptions of new species. 
His collection of Polynesian Mosses is very large and complete. 

For-a number of years he had been collecting and studying 
the various species of the genus Sphagnum. In 1893 he issued a 
list of the North American species and a prospectus of a proposed 
distribution, undertaken in connection with Mr. Edwin Faxon, to 
be called “Sphagna Boreali-Americana Exsiccata.” He spent 
much time in collecting, preparing and determining sets, and was 
very particular about the excellence and homogeneity of his speci- 
mens. Many of the sets are practically ready for distribution, 
and it is to be hoped that these, at least, may be issued at some 
future time. 

Professor Eaton prepared the botanical definitions for Web- 
ster’s International Dictionary, and contributed many reviews of 
botanical books to the various periodicals. During the last four 
or five years he has contributed the botanical reviews to the ‘‘ Na- 
tion”’ and to the “ New York Evening Post,” touching upon cur- 
rent botanical changes and opinions in the graceful and discerning 
way characteristic of him. 

Professor Eaton was one of the original members of the Tor- 
rey Botanical Club, and remained an active member for a number 
of years. Later he became a corresponding member. He was a 
regular and active contributor to the BuLierin, especially during 
the earlier years of its existence. 

Besides his botanical interests Professor Eaton had many 
others. He was an enthusiastic lover of athletic sports, of archery, 
of baseball and football, and of fishing and hunting. He was an 
ardent student of anthropology and genealogy. He was a mem- 
ber of several genealogical societies and made a considerable study 
of the genealogies of the Eaton and Selden families, and published 
the results in several papers. He was also Governor of the Con- 
necticut Society of the Colonial Wars for some years. 

In politics Professor Eaton was a Republican until 1884, but 
from that time took an active part in the independent movement 
in Connecticut. 

His interest in the classics, both of the Latin and of the 
Greek, was very great, and he much deplored the present lack of 
facility and even use of proper Latin, both in the choice of generic 


347 


and specific names and in descriptions. He strongly advocated 
the publication of a Latin description in the case of a new species. 
He was very impatient of the use of barbarisms in Latin nomen- . 
clature. 

As a man, Professor Eaton was possessed of a most pleasant 
personality, winning the esteem and love of all who had the privi- 
lege of acquaintanceship with him. He was generous to the ex- 
treme and counted neither time nor trouble when performing 
any act of friendship. : 

As a botanist he was careful to an extraordinary degree. No 
work of his was ever slovenly or hastily done, and he had little 
sympathy with work of that kind on the part of others. His in- 
struments were most carefully kept in order, his microscopic 
preparations most neatly prepared, finished and labelled, and all 
the results of his study compared and worked over and over 
again. His extraordinary severity in these matters led him to 
publish much less than he otherwise might well have done. 

Professor Eaton was very conservative in regard to changes in 
scientific methods and views. He was loth to part with what he 
considered good until he was absolutely convinced that he might 
obtain something better. But when the better had really been 
demonstrated he lost no time in changing either views or methods. 
His attitude toward. the various proposed changes in the rules for 
governing the nomenclature is a good case in point. While 
anxious that there should be no blind adherence to rules already 
established, and perfectly ready to accept such changes as might 
lessen such confusion as already really existed, he viewed retroac- 
tive measures overturning hosts of names already long and firmly 
established in the literature as productive of great immediate con-_ 
fusion without giving by any means a certain promise of surer 
criteria upon which to establish stability in the future; in fact as 
giving no greater certainty, if as great, as that given by the rules 
already existing and followed for years by the best workers. 

Asa teacher he was kindly and inspiring, not suited to manage 
large classes of unwilling students, such as often fell to his lot, 
where much sternness and rigor was needed to compel the 
wavering attention and to force the stubborn mind to effort, but 
especially fitted to encourage and to train those desirous of pur- 


348 


suing either some especial line or even of obtaining a general bo- 
tanical education. To these he tendered the privileges of his 
library and collections, which had to be supported by himself 
alone, and opened to them not only all the material resources at 
his command, but unlocked for them all the treasures of his own 
experience, the results of the patient study of many years. 
Professor Eaton’s religious views were shown rather in the 
earnestness and simplicity of his life than by any profession of 
faith. Where he could not thoroughly understand and believe he 
was contented to hope, and his last months, even during intense 
suffering, were hopeful and peaceful, and he left this life calm in 
his trust in the good to come. He was a communicant of the 
Episcopal Church, and attendance upon divine worship was to 
him a pleasure and an inspiration rather than a duty. At his 
particular request, his funeral services were of the nature of simple 


religious consolation to his relatives and friends, with an entire ab- 
sence of official ceremony. 


Bibliography, 
1856. On three new Ferns from California and Oregon. Amer. Journ. Sci. (II.) 
2a 138. 


1858. Characters of some new Filices, from Japan and adjacent regions, collected 
by Charles Wright in the North Pacific Exploring Expedition under Captain Jehn 
Rodgers, Proc. Amer. Acad. Arts and Sci. 4: 110, 111. 

1859. Enumeration of Ferns collected by Mr. Charles Wright in Eastern Cuba 
in 1856-7. Amer. Journ, Sci. (II.) 27: 197-201. 

Equisetacex, Filices, etc. U.S. and Mexican Boundary Surv. f Botany by John 
Torrey.] 233-236. 

1860. Filices (Ferns). [In: Chapman’s Flora of the Southern U. S., 1st Ed., 
- 8, New York.] 585-599. 

[Reprinted without change in the 2d Edition, 1885. ] : 

Filices Wrightianae et Fendlerianae, nempe Wrightianae Cube nses et Fendlerianae 
Venezuelanae (nonnullis Panamensibus, etc.,ex coll. A, Schott et S. Hayes interjectis) 


* The basis for the present list was taken from the “ Bibliographies of the present 
officers of Yale University,” edited by Prof. Irving Fisher (New Haven, 1893). The 
list of works there given was written out by Prof, Eaton. This has been followed 
the main, but the greater part of the references have been compared and some changes 
and additions made. The writer is very much indebted to Mrs. Eaton, to Mr. George 
F. Eaton and to Prof. W. H. Brewer for much help, both in connection with the de 
tails of the bibliography and of the sketch of the life of Prof. Eaton. 


549 


enumeratae novaeque descriptae: dissertatio inauguralis, quam in auditorio botanico 
Universitatis Harvardianae ad gradum baccalaurealem in scientiis legitime obtinendum 
die XIV. Jul. MDCCCLX., habuit Daniel C. Eaton, A.M. Mem. Acad, Amer, 
Scient. et Artium. (II.) 8: 193-218. 

1861. Statistics of the Class of 1857 of Yale College, collected by Daniel C. 
Eaton, Class Secretary. New York, 8°, pp. 93. 

1865. On the Genus Woodsia. Canadian Naturalist, (n. s.) 2: 89-92, 

Two new Ferns of California. [In: Botan, Contrib. of Asa Gray.] Proc. Amer. 
Acad. of Arts and Sci., 6: 555, 556. 

1867. Filices (Ferns), {In: Manual of the Botany of the North United States, 
ete., by Asa Gray, 5th Ed., 8°.] 655-672. pl. 25-79. 

1868. Acrogens [In: Field, Forest and Garden Botany, etc., by Asa Gray. New 
York, 8°.] 359-374. 

1869. Ein neues Adiantum von Cuba (Adiantum sericeum.) Botan. Zeitung, 
27: 361-362. 

1870. Biographical Record of the Class of 1857 of Yale College during twelve 
years from graduation with an account of the decennial meeting of the class. New 
Haven, 8°, pp. 54. 

1871. Compositae, Equisetaceae and Filices. Exploration Fortieth Parallel, 5: 
136-208. f/. 15-20; 395-397. pl. go; 422-425. 

1872, Plain Directions for collecting Algae. New Haven, 8°, pp. 4. 

1873. Characters of new Ferns from Mexico. Proc. Amer. Acad. Arts and Sci., 
8: 618-619, ; 

List of marine Algae collected near Eastport, Maine, in August and September, 
1872, in connection with the work of the U. S. Fish Commission under Prof. S. F. 
Baird. Trans. Conn. Acad. Arts and Sci., 2: part 2, 343-350. 

1873-1883. New or little known Ferns of the U.S, Bull. Torrey Botan. Club, 
4: 11-12, 1873; 4: 18-19, 1873; 6: 33, 1875; 6: 71-72, 1876; 6: 263-265, 1878; 
6: 306-307, 1879; 6: 360-361, 1879; 7: 62-64, 1880; 8; 4-5, 1881; 8: 99-100, 
1881; 8: 111, 1881; 9: 49-50, 1882; 10: 26-29, 1883; 10: 101-102, 1883. 

1874. Townsendia Parryi and 7. condensata. [In: Botanical observations in 
Western Wyoming by Dr. C. C. Parry.] Amer. Naturalist, 8: 212, 213. 

1875. A List of the marine Algae collected by Dr. Edward Palmer on the coast 
of Florida and at Nassau, Bahama Islands, March-August, 1874. New Haven, June, 
8°, pp. 6. 

1877. A Moss new to the U. S. (Lilotrichum undulatum Beauv.) Botan. 
Gazette, 2: 98, 

Calvin Selden, of Lyme, and his children. (An address delivered at a meeting of 
the Selden Family at Fenwick Grove, Saybrook, Conn, Aug. 22.) New Haven, 8°, 
Pp. 15. 

1877-1889, Joint editor of Farlow, Anderson and Eaton’s Alge America- 
Borealis Lxsiccate, Fasc. 1-5, Boston. 

1878, Ferns of the Southwest. An account of the ferns which have been col- 
lected in so much of the territory of the United States of America as is west of the 
Tosth degree of W. Longitude, and south of the goth degree of N. Latitude. U.S. 
Geog. Surveys West of the One Hundredth Meridian. First Lieut. G. M. Wheeler, 
Corps of Engineers, U. S. Army, in charge, 6: 299-340, f/. 30. 


390 


« Introduction” and « Anogens.” [In: A Catalogue of the flowering Plants and 
higher Cryptogams growing without cultivation within thirty miles of Yale College. 
Published by the Berzelius Society. New Haven, 8°.] V.-VII.; 61-71. 

Hybrids and hybridism. Rept. Sec. Conn. Board Agric., pp. 19. 

Conomitrium Fulianum at Hamden, Conn. Bull. Torrey Botan. Club, 6: 244. 

1878 and 1880, Fendler’s Ferns of Trinidad. Botanical Gazette, 3: 89-91; 5: 
I0g—L1O (121-122). 

1879. A new Hawaiian fern (Hymenophyllum Baldwinii). Bull. Torrey 
Botan. Club, 6: 293. 

A new Hawaiin fern (Aspidium Boydie). Ibid. 361, 362. 

1879-1880, The Ferns of North America. Colored figures and descriptions, 
with synonymy and geographical distribution of the ferns (including the Ophioglos- 
saceae) of the United States of America and the British North American Possessions. 
The drawings by J. H. Emerton and C, E, Faxon. 2 vols., 4°, 1: XIV.+-35% pl. 
I-45; 2: XXXI.+ 285,, p/. 46-Sz. : 

1880. Vascular Acrogens. [In: Botany of California, by Gray, Brewer and 
Watson, 4°, Cambridge, Mass.] 2: 329-352. 

Systematic Fern-list : a classified list of the known ferns of the U. S., with the 
geographical range of the species. New Haven, 8°, pp. 12. 

1881. Anew American Cynaroid Composite (Saussurea Americana). Botan, 
Gazette, 6: 283. 

1882. Vegetable Fibres in an Oriole’s Nest. Bull. Torrey Botan Club, 9: 57- 

1883. Plants new to the Connecticut Flora. Ibid. 10: 102. 

1884. Tea, Coffee and Chocolate : their nature and their effects, West. Farmer's 
Almanac, 36-38. 

Another Florida Fern ( Phegopteris tetragona Mett). Bull. Torrey Botan. Club, 
II: 67. 

Teratology in the Ox-eye Daisy. Ibid, 11: 67. 

The Dedham Eatons from 1635 to the fifth generation. New Haven, 8°, pp- 8. 

1887. Asplenium rhizophyllum var. Biscaynianum, Bull. Torrey Botan. 
Club, 14: 97. 

1888, The Family of Nathaniel Eaton, of Cambridge, Mass, (read April 22 
1884). Papers of the New Haven Colony Histor. Soc. 4: 185-192. 

Report of the Secretary. [In: Rept. Eaton Family Assoc. New Haven. ] 1-6. 

The family of John Eaton, of Dedham. Ibid. 11, 12. 

1890. Vascular Acrogens or Pteridophytes. [In: Manual of Botany of the 
Northern United States by Asa Gray; 6th Ed., by Sereno Watson and John M. Coul- 
ter. New York and Chicago, 8°.) 675-701. /. 76-27. 

An undescribed Heuchera from Montana (H. Williamsii), Botan. Gazette. 
15: 62. 

A new Moss of the genus Bruchia. Bull. Torrey Botan. Club, 17: 100, 101. p/. 407+ 

On Buxbaumia indusiata Bridel. Ibid, 126, 127. 

A new Fern ( Chetlanthes Brandegei), Ubid. 215. pl. 104. 

A new Fern from Lower California (Asplenium blepharodes). Zoe, 1: 197+ 

Botanical definitions in Webster’s Internat. Dict., Springfield. 

1891. Report of the Secretary and report on the Eatons of Dover, England, 
[In: Rept. Eaton Family Assoc. 1890. New Haven.) 1-14. 


301 


1892. List of Ferns from Southern Patagonia. List of Mosses from Fuegia and 
Patagonia. Contrib. U. S. National Herbarium, 1: 1 38-139. 

1893. A Check-list of North American Sphagna, arranged mostly in accordance 
with the writings of Dr. Carl Warnstorf. New Haven, 8°, pp. Io. 

Sphagna Boreali-Americana Exsiccata. [ Prospectus, with directions for collect- 
ing and preparing specimens.] New Haven, 8°, pp. 3. 

Reviews. Amer. Journ. Science (III.), 7: 520-522, 1874; 15: 75, 1878; 17: 
487-488, 1879; 18: 76-77, 1879; 21: 330-332, 1881; 22: 158-160, 1881; 24: 
156, 1882; 39: 240-243, 1890. The Nation, 51: 488, 1890; 52: 36-37, 1891; 52: 
326, 1891; 52: 466, 1891; 53: 55» 1891; 53: 130-131, 1891; 53: 320-321, 1891; 
53: 336, 1891; 54: 193, 1892; 54: 197, 1892; 54: 267, 1892; 55: 114, 1892; 
55+ 360, 1892; 55: 482, 1892; 57: 271, 1893: 57: 287, 1893; 57: 373, 1893; 
57: 474-475, 1893; 58: 155, 1894; 59: 146-147, 1894; 59: 176, 1894; 59; 
195~196, 1894; 59: 216, 1894; 59: 234, 1894; 59: 407, 1894; 60: 12, 1895; 60: 
147, 1895; 60: 258, 1895; 60: 306, 1895. 


The Genus Sanicula in the eastern United States, with Descriptions 
of two new species. 


By EvuGENE P. BICKNELL. 
(PLATES 241-245.) 


It has been generally received as a settled fact of our geogra- 
phical botany that the genus Sanicula had but two representatives 
in the flora of eastern North America. These plants, S. Mary- 
landica and S. Canadensis, though coming down to us as well- 
accredited species from the time of Linnaeus, have proved a recur- 
ting source of confusion to our botanists and, as now appears, 
have never been rightly understood. In 1824 Dr. Torrey reduced 
Canadensis to a variety of Marylandica (F\. U.S. 302), and in 1838 
Torrey & Gray (Fl. N. Am. 1: 602) discredited the plant altogether 
naming it as a synonym of the latter species. A few years later, 
however, Dr. Torrey took occasion to restore the plant to its 
Original status, remarking that he had become persuaded that the 
two species were quite distinct (Fl. State of N. Y. 1: 265); not- 
withstanding this, recent authorities have reverted to Dr. Torrey’s 
€arlier view. In the “Review of North American Umbelliferae, 
Coulter & Rose” (1888), S. Canadensis finds recognition only as a 
Slightly differentiated form of S. Marylandica, a disposal of the 
Plant which is followed in the sixth edition of Gray’s “ Manual” 


352 


(1893). Dr. Britton, however, in his ‘Catalogue of Plants found 
in New Jersey” (1889), again accords both plants full specific 
rank. That this understanding of the problem is the only one 
which the facts allow does not admit of any question whatever. 
The most zealous reductionist need have no trouble in assuring 
himself of this, provided only that ordinary eyesight and an open 
mind be not wanting to his equipment. Indeed, as species present 
themselves, these two plants must be regarded as not even very 
closely related within their common genus. The trouble all along 
has been that we have unwittingly been seeking to force four dis- 
tinct species into the limits defined for two. As an inevitable re- 
sult these two, though wholly distinct, have been seen under such 
confused outlines as to make it appear quite impossible to assign 
any constant characters to either. It thus becomes necessary to 
diagnose them anew in connection with the description of their 
long over-looked congeners. 

It has not been ventured to define two new eastern species of 
this difficult genus without considering very carefully their rela- 
tionships with the species already known, at the same time keep- 
ing in mind the harmfulness of distinctions too finely drawn. The 
danger of fallacious distinctions, however, proves not to press at 
all closely in the present case. A critical study from herbarium 
specimens of the four plants here presented, which has supple- 
mented an intimate acquaintance with three of them in the field, 
enables me without any reservation to subscribe them all as 
authentic and well-defined species. 

Sanicula Marylandica LL. 


. Commonly two or three feet high—one-and-a-half to four feet; 
stem single, or two or three together from the same rootstock, tet 
minating in a general umbel which is usually but 14 to 4 the length 
of the entire plant; there is frequently also a short, imperfectly um- 
bellate lateral branch. Leaves thickish, dull bluish green, two Of 
three on the stem between the basal leaf and the involucre; basal 
leaves several, on long erect petioles, only one of them cauline, the 
others rising separately from the rootstock; free stem-leaves near ly 
or quite sessile, or the lowermost short-petioled ; involucral leaves 
usually much reduced, sometimes under an inch long, cleft or 
parted, not divided into separate leaflets; larger leaves five-divided, 
appearing seven-divided from the deep partition of the basal leaf- 
lets; upper leaves five-divided, or appearing so. The leaves ge 


353 


erally are large, spreading from three or four to eleven inches, the 
leaflets reaching a size of six inches by three; leaflets variable in 
shape and marginal pattern; cuneate-obovate, to oblanceolate, or 
sometimes elliptic, obtuse or acute, irregularly doubly serrate, ser- 
rate or dentate-serrate, with mucronately-pointed teeth, usually 
more or less incised-lobed towards the apex. Branches of umbel 
rather rigid, usually sharply ascending, normally three—two to 
four—and sub-equal, two to eight inches long; ultimate rays three, 
an inch long—3’”-18/’—(abnormally longer and bearing a pair of 
minute bracts) sometimes divaricate; rarely the umbel may be 
unevenly three times compound; secondary involucres of firm foli- 
aceous bracts, either cut-lobed or merely serrate, often minute, 
rarely reaching a length of one inch. Sterile flowers numerous, 
with the perfect flowers, or in separate peduncled capitate clus- 
ters; pedicels 2” long; calyx 1”, cleft nearly to the base into 
narrowly lanceolate attenuate-cuspidate lobes; petals oblanceo- 
late, equalling or slightiy exceeding the sepals, greenish-white, as 
are the anthers; filaments exserted, 2” long; umbellets at anthesis 
5-7” wide. Fruits large, 3-6 in each umbellet, somewhat ovoid, 
sessile, spreading, the body 2” long and nearly as broad or, 
through the bristles, 3-5” wide, 3-314" high; bristles numer- 
ous, crowded, arranged in no regular order, very straight to the 
minutely hooked apex, prominently corky-bulbous below, rudi- 
mentary at base of carpel, above becoming 2” long, ascending, 
the uppermost erect, parallel with and about equalling the erect 
calyx-lobes; styles long, at first erect-spreading but early recurv- 
ing from the top of the closed calyx sometimes quite outside of 
the bristles. Transverse section of seed somewhat oblong, seed- 
face plane or slightly concave, dorsal surface more or less fluted 
or grooved for the partial accommodation of the five large oil- 
tubes ; frequently a slight separation of the wall of the pericarp 
from the middle of the seed-face leaves a cavity having the ap- 
pearance of an additional oil-tube ; pericarp corky-thickened, pale 
brown; commissural scar broad, elliptic. Root perennial, of 
very coarse fibres fasciculate from a short knotted contractedly- 
branched rootstock, which, when cut, has a distinct terebinthine 
odor. 


Newfoundland and Canada, southward to the mountains of 
Georgia, west to the Rocky Mountains. (Plate 241.) 

A specimen of a Sanicula which, for the present, must be re- 
garded as an aberrant form of this species was gathered on Look- 
out Mountain, Tennessee, June 21, 1894. The plant is small and 
has two very unequal branches, one capped by a single umbellet, 
the other bearing an umbel of four divaricate rays; the leaves are 
trifoliate, with obovate nearly simply-serrate leaflets, the lateral 


304 


ones deeply cleft ; the three stem leaves are slender-petioled, even 
the involucral leaves are short-petioled. The immature fruit when 
crushed exhales an odor much like that of the herbage of the 
carrot, and very different from that of true Jarylandica. The 
styles are shorter than normally, and not recurved, and with the 
stylopodium are perfectly developed in all the infertile flowers, 
which are further noteworthy from their rigidly spreading sepals. 

I have observed no other instance of styliferous sterile flowers — 
in any of our eastern Sawicudae, though an examination of speci- 
mens of Sanicula Europaca show that they occur normally in that 
species. 

So far as an inference may be drawn from this single specimen, 
we may suspect the existence in the Southern Alleghenies of still 
another form of Sanicula meriting definite recognition. 

The plate of S. Marylandica in “The Flora of the State of 
New York” well illustrates the confusion that has prevailed with 
regard to this species. The large leaf in outline, together with 
the single bristle, the mature carpel and the fruit section, are of 
S. Marylandica; the figure of the upper part of a fruiting plant, 


and the detached umbellet, are good representations of S. gregaria 
as here distinguished. 


SANICULA GREGARIA N. sp. 


~ Stems rather weak, often clustered, from one to three feet high, 
commonly two to two-and-a-half feet, naked to the first branch or 
with a single petioled leaf, above dividing into a freely branched 
three or four times compound general umbel which takes up US- 
ually 14 to % the length of the plant. Basal leaves usually 
more numerous than in Marylandica, sometimes nearly as large 
and as long-petioled, only one of them truly cauline, the others 
rising from the rootstock near the base of the stem. Leaves thin, 
bright green, digitately five-divided, the divisions all petiolulate, or 
the lateral pairs slightly united at base; leaflets varying from 
cuneate-obovate, through rhombic, to elliptic and lanceolate, 
sharply doubly-serrate with bristle-tipped teeth, often regularly 
serrate-lobed along the margins, sharply incised-lobed above, 
acutely-pointed ; involucral leaves scarcely reduced, trifoliate, the 
petiolulate leaflets commonly lanceolate, often somewhat curved or 
sub-falcate ; involucres of the second, and even of the third, series 
conspicuously foliaceous, the former three-divided, and sometimes 
spreading four or five inches, the latter merely lobed. Branches 
of general umbel, with their divisions, commonly in sub-equal 


305 


series of three, sometimes of two, slender, straight, ascending, the 
ultimate rays 6’’-15’’ long or, occasionally, much longer in effort 
to branch again for an umbellate division of the fourth series, 
which may be perfectly realized. Frequently the primary 
division of the stem is only sub-umbellate owing to the displace- 
ment of one of the branches; on many plants there is a well- 
developed lateral flowering branch. Staminate flowers with the 
fertile, or in separate heads peduncled in the forks of the stem or 
main branches; pedicles slender, 1-24” long, four or five times 
times the length of the minute campanulate calyx, which is cleft 
little more than half way down into somewhat triangular-ovate 
rather obtuse lobes; flowering umbellets small, about 3/4” in dia- 
meter; petals obovate, heart-shaped, fully twice the length of the 
sepals, yellowish-green, filaments exserted, anthers bright yellow. 
Fruits very small, 3-5 in each umbellet, broader than long, and 
somewhat obovate in outline, measuring through the bristles 114’”— 
2” high, by 2-214” wide, evidently pedicelled, pedicels not over 
1” long, papillose with rudimentary bristles; bristles of the fruit very 
small and weak, rudimentary at base of pericarp, gradually length- 
ened above, but not exceeding 1”, and commonly half that size, 
not crowded, arranged in eight or ten rows which may be evident 
or obscure, curved at tip into a more open hook than in Mary- 
landica ; though sometimes slightly depressed-dilated at base, they 
commonly rise abruptly from the nearly even surface of the ma- 
ture carpel, which is of a dark or blackish color; fruit reflexed at 
maturity; styles abruptly spreading and recurved among the 
bristles, sometimes extending more than half way around the fruit. 
Seed rounded-oblong in cross-section, dorsal surface even, not at 
all sulcate or angled, commissural face plane; pericarp thin and 
membranous, closely investing the seed ; oil-tubes five, very small ; 
commissural scar linear. Rootstock much as in Marylandica, but 
not so stout and contractcd, the fibres less fasciculate, much more 
slender and fibrillose, and of a blackish instead of brownish color. 
The substance of the rootstock hasa fainter and altogether differ- 
ent odor. (Plate 242.) 


This plant when once known is in no danger of being con- 
fused longer with either Marylandica or Canadensis. It is clearly 
a perfectly distinct species, not even to be regarded as inter- 
mediate between the other two, nor as related to them by any 
nearer ties than those of acommon genus. Its nearest affinity is with 
Marylandica, with which it has been mainly confounded, notwith- 
standing that the general aspect of the two is strikingly dissimilar. 
To compare the superficiary characters, S. gregaria is a less robust 
plant, more widely branched and more leafy above, but with less 


356 


leafy stem, quite missing the general appearance presented by 
Marylandica of a tall strict plant stiffly umbellate at the top of a 
simple stem. The general umbel of gregaria is more compound 
and much more slenderly branched and foliaceous; the leaves are 
thin and bright green, in contrast with the firm and dull or bluish- 
green leaves of Marylandica; furthermore the leaves of gregaria 
are never seven-divided, nor are the basal leaflets parted or deeply 
cleft, as seems always to be the case with both Marylandica and 
Canadensis; the leaves, too, are more pointed, and more sharply 
incised and serrate-lobed than in Marylandica, and never assume 
the narrowly oblanceolate form with long tapering base so often 
seen in that species. In the early spring the plant forms dense 
clusters of leaves which may number forty or more rising from 
one compound rootstock, At flowering time the bright yellow an- 
thers and yellowish-green petals of gregaria give the plant a char- 
acteristic appearance and distinguish it conspicuously from our 
other species. The flowers, moreover, have a slight fragrance 
faintly suggestive of those of the spice bush. 

This plant seems to have escaped the observation of every one 
of our botanists except Dr. Darlington, who, in his “Flora Ces- 
trica,” makes an unmistakable allusion to it. Speaking of S. Mary- 
landica, he says, “It also presents a variety, with dullish yellow 
flowers.” 

As already remarked, a good figure of this species appears in 
“The Flora of the State of New York,” in the plate of S. Mavy- 
landica. 

S. gregaria usually grows in close leafy often extensive com- 
munities, in damp low woods and thickets; S. Marylandica is much 
less gregarious, scattering itself in loose colonies through rich, often 
hilly or rocky woods; S. Canadensis often grows in a still more 
scattered way, and seems to prefer a simpler more earthy soil than 
Marylandica. Sometimes, however, in damp thickety spots in 
somewhat rocky woods all three plants may be found near 
together. 

S. gregaria is the first of the three to flower in the spring: 
coming into bloom at New York from the second to the fourth 
week of May; S. Marylandica follows a week or two later, more 
or less; S. Canadensis is much later, not flowering before the 
third or fourth week in June, in some seasons not until July. 


357 


S. gregaria is perhaps the most common Sanicula near New 
York City, and is described from specimens collected in Van 
Cortlandt Park. In the Herbarium of the Torrey Club is a speci- 
men labeled « New Durham, N. J., June 6, 1862, W. H. L. 
[eggett].”” The specimens contained in the Columbia College 
Herbarium are as follows: « Virginia, Asa Gray, 1840,” also 
bearing the stamp of the Meisner Herbarium; “Southwestern 
Virginia, slopes of White Top Mountain, 2600-5000 feet, col- 
lected May 28, 1892, by N. L. and E. G. Britton and A. M. Vail;” 
“Southeastern Virginia, Greenville County, June 19, 1893, A. A. 
Heller;” « Fort Riley, Kansas, May 27, 1892, E. E. Gayle;” 
“Arkansas, Dr. Pitcher ;” « Northeastern Nebraska, June 15, 1893, 
Fred Clements.” 


SANICULA CANADENSIS L. 


~ Usually lower and more slender than S. Marylandica, but some- 
times even stouter and taller, one to over four feet high. Stem 
always single, widely branched above, and readily putting forth 
alternate axillary branches which, sometimes beginning at the 
base of the stem, may number six or more up to the terminal 
fork ; branches ascending, naked below, their upper divisions often 
widely spreading, even horizontal, the whole forming an open 
panicle which may spread to a breadth of two feet. General plan 
of branching dichotomously or paniculately-umbellate, often ap- 
pearing widely dichotomous throughout from the constant 
Suppression of a third branch in each ascending series ; the um- 
bellate character of the branching is thus much less obvious than 
in our other species, realizing itself definitely only in the distal 
ramifications or in the terminal umbels; above the involucres the 
branching varies from definitely dichotomous and two or three 
times compound, to indefinitely decompound, with somewhat 
fasciculate umbellets; the fruit-bearing rays are only I ie 2 ' 
long, much shorter than in any other one of the eastern species. 
Basal leaves from two or three to six or more, erect on long 
petioles, all strictly cauline ; free stem leaves commonly four to 
eight on petioles of gradually decreasing length above, but not 
becoming obsolete even in the involucral leaves, or indeed, in the 
Secondary involucres; leaves three-divided appearing pentafid 
from division ofthe lateral leaflets, or the upper stem leaves, more 
rarely the lower, sometimes simply trifoliate ; leaflets dull green, 
cuneate-obovate, oblong or elliptic, often oblique, with less pro- 
nounced tendency to narrowly obverse development than is shown 
by Marylandica, and never, apparently, assuming the oblanceolate 
forms so frequent in that species; also, as a rule, less coarsely ser- 


358 


rate, and much smaller, though occasionally reaching a breadth of 
six inches, with leaflets over 314” 14’; involucral leaves more 
or less reduced, even sub-bracteal, three-divided, those of the 
succeeding series very small and bract-like, divided or cut- 
lobed. Umbellets very small at anthesis, few-flowered; sterile 
flowers with the perfect ones, never in separate heads, few, often 
only one or two in each umbellet, 1%” long, on pedicels not over 
1’’ long, often less; calyx deeply parted, sepals narrowly lanceo- 
late, acute or cuspidate, exceeding the minute white petals ; an- 
thers white, little exserted. Fruit subglobose, three together = 
one plane, the lateral spreading or slightly reflexed, very small, 1 > 
1%” long and broad, or, measured through the bristles, about 2 
long by 2’-3’’ wide; short-pedicelled, more distinctly so when 
young ; pedicels 4””-'%4”” long, wrinkled, striate ; bristles numer- 
ous, dilated below, somewhat regularly arranged in longitudinal 
rows, well developed to the very base of the carpels, of more unl- 
form length throughout than in Marylandica,and more Syren 
rarely exceeding 1’ in length, the uppermost closely parallel 
with and equaling, or slightly exceeding, the erect calyx-lobes ; 
styles small and difficult to observe, shorter than the calyx-lobes; 
mature fruit dull brown; commissural scar linear. Seed more 
rounded in cross-section than in Marylandica, the dorsal surface 
more prominently sulcate-fluted, the face convex, sometimes 
sharply so, and bevelled off on either side to permit a more interior 
position of the commissural oil-tubes than is seen in Marylan 
Root consisting of rigid horizontal fibres, which taper slenderly 
from a strong woody base. 
Massachusets to Florida and Texas, west to Kansas and Ne- 
braska. (Plate 243.) 
The genus Sanicula is set down as consisting of “ perennial 
herbs,” yet this plant, in the vicinity of New York at least, 1S 
not perennial but apparently biennial. Unlike Marylandica and 
Sregaria, the mature plant is readily puiled up from the soil, and 
in the autumn, while these species are still green, is to be found 
completely dead throughout, with the roots beginning to decay. 
This species stands sharply apart from both Marylandica and 
gregaria, not only in the characters of leaves, flowers and fruit, 
but in fundamental differences in the root and the plan of branch- 
ing. It is indeed hard to understand how such obviously distinct 
species could ever have been confused. 
The species shows two main lines of variation which, in their 
extreme manifestations, present plants of somewhat dissimilar o 
pect. On the one hand the result is a slender plant with sma 


359 


fruit and leaves, the latter with finely-cut marginal pattern, the 
branching widely dichotomous and confined mainly to the upper 
part of the stem; the opposite form is coarser and stouter, with 
ascending or even-suberect, often simpler, branches and larger 
fruit; the leaves are much larger with coarser serration, and more 
trifoliate tendency, those of the upper stem sometimes perfectly 
trifoliate. An extreme example of this form in the Columbia Col- 
lege Herbarium, labelled «« Massachusetts ” in Dr. Torrey’s hand- 
writing, is the only specimen seen in which the character of 
short styles does not hold strictly true; in this specimen, which 
is in the flowering stage, the styles are longer than the sepals, and 
somewhat spreading, though not recurved. It is to be said, how- 
ever, that this particular plant is pretty certainly abnormal; the 
branching is unusually coarse, and the rays and pedicels some- 
what thickened, apparently indicating a tendency to fasciation. 


SANICULA TRIFOLIATA n. sp. 


About two feet high; branches several, alternate, ascending, 
or the lower often longer and suberect, simple, terminating either 
in the flowering umbel or in two frequently widely-spread- 
ing umbel-bearing rays, with a shorter simple ray between ; rays 
of flowering umbels three to five, about an inch long; involucres 
and involucels of small serrate leafy bracts. Cauline leaves petioled, 
large, five to seven inches broad, conspicuously trifoliate, the 
broad leaflets petiolulate, ovate, elliptic-ovate or rhombic, coarsely 
doubly serrate with rather open, almost spinescently-mucronate 
teeth, or even incised-lobed, acute, the lateral leaflets mostly not 
at all cleft, except sometimes in the lower leaves; basal stem- 
leaves long-petioled, smaller than the main cauline leaves, the 
leaflets even slightly united at base, often obtuse, the lateral pair 
Narrowly cleft on the lower side, but not sufficiently so to destroy 
the trifoliate character of the leaf. Sterile flowers few, 14” or less 
long, on slender pedicels about 2’ long ; sepals lanceolate, acumi- 
nate-cuspidate with slightly incurved points. Fruits sessile, three 
to five together, somewhat reflexed at maturity, ellipsoid or 
globose-oblong, large, becoming 3%” high to tip of the erect 
Sepals, the greatest spread of the bristles under 3’’; prickles 
Stout, swollen below, not crowded, small and reflexed at the 
base of the carpel, above becoming 114” long, and spread- 
ing or ascending; sepals on mature fruit 1” long, united at 
base, appressed, echinate with sharp, slightly incurved points, 
raised on a slight prolongation at the apex of the fruit, and 
forming a conspicuous beak-like projection quite distinct from the 


360 


surrounding prickles and usually a little exceeding them; styles 
short, nearly erect, strictly included; commissural scar elliptic, 
occupying most of the face of the mericarp. Seed somewhat cres- 
centic in transverse section, the dorsal surface even, the face con- 
cave, sometimes almost sulcate; oil-tubes of two kinds, one set 
consisting of a pair of large vittae latero-commissural in the peri- 
carp; the other set a series of minute ducts lining the inner face 
of the pericarp, especially along the commissural concavity, where 
they may number as many as twenty. In the dried fruit the 
hardened secretion of the large tubes is nearly colorless, that of 
the minute ducts of areddish-amber color. Root somewhat similar 
to that of S. Caxadensis and likewise, apparently, not perennial. 
(Plate 244.) 

While in its general aspect this plant stands out clearly enough 
from our other species, the crucial test of fruit characters shows it 
to be indeed profoundly different. In fact, its fruit, in the 
numerous minute oil-tubes, supplemented by a large outer pair 
having, apparently, a different secretion, presents characters hith- 
erto unreported in the the genus so far as I have been able to dis- 
cover. 

I find three specimens of this plant in the Columbia College 
Herbarium, and one in the Herbarium of the Torrey Club, now at 
the College of Pharmacy in New York. Of the former, one is €S- 
pecially interesting as having passed through Dr. Torrey’s hands 
and bearing his note: “Intermediate between S. Marylandica and 
S. Canadensis,” it is labeled simply «‘ Ohio.” The other specimens 
are all named S. Canadensis ; one is No. 835 of the Geological and 
Natural History Survey of Canada, and is labeled “ dry, rich woods, 
Ontario, Amherstburg, Macoun, 5, 10, 1892;” another was col- 
lected by Dr. Britton at Indianapolis, Aug. 25, 1890. The speci- 
men in the local Herbarium is of a complete fruiting plant and 
bears witness to Mr. Leggett’s careful methods in botanical 
work; it is labeled “ Canaan, Conn., ’68, W. H. L.” The label, 
however, is not in Mr. Leggett’s handwriting, and, as it was doubt- 
less copied and affixed after his death, is perhaps open to doubt 
as applying to the identical plant which now bears it. The locali- 
ties of Ontario, Ohio and Indiana suggest a more exclusively 
central distribution than limits our other species, though the plant 
may well occur further east. I feel almost certain of having myself 
seen it in Alleghany County, New York, in 1891, in a flora 
strongly characteristic of the northern Alleghenies. 


361 


In this connection it is interesting to note that when, in 1824, 
Dr. Torrey reduced S. Canadensis to a variety of S. Marylandica 
he evidently based his conclusion upon the examination of a spec- 
imen of the very plant here described as S. *#ifoliata. This is 
made pretty clear by his description of his var. Canadensis, which 
was drawn up, by the way, from a Specimen collected at Litch- 
field, Connecticut. As already remarked, Dr. Torrey afterwards 
revised his opinions in regard to Canadensis and a plate of the 
species occurs in his flora of the State of New York. 


Descriptive Key to the eastern Species of Sanicula. 


Styles long, recurved ; branching definitely uméellate throughout; sterile flowers 
often in sepirate heads; perennial. 

a. Fruit large, 3’’ or more high to tip of sepals, ovoid, sessile, spreading ; pericarp 
corky-thickened ; bristles stout, bulbous at base, becoming 2/’ long, but not exceeding 
calyx-tubes ; sterile calyx 1// long, deeply parted; sepals lanceolate, cuspidate-acute ; 
general umbel twice compound, ¥% to ¥ the length of entire plant; stem-leaves two or 
three; larger leaves 7-divided; involucral leaves cleft; petals and anthers greenish-white, 
the petals oblanceolate, equalling the sepals; commissural scar elliptic; seed furrowed 
dorsally ; oil tubes large. Marylandica. 

6, Fruit very small, under 11%4’/ somewhat obovoid, short-pedicelled, reflexed ; 
pericarp membranaceous; bristles weak, not bulbous xt base, 14// to 1// long, but exceed- 
ing calyx-lobes, sterile calyx 34’! or less, cleft only half way down, sepals ovate, obtuse; 
general umbel three times compound, & to % the length of plant; stem naked or with 
a single leaf; larger leaves 5-divided ; involucre of distinct leaflets; anthers deep yel- 
low; petals yellowish-green, obovate, twice the length of sepals; commissural scar 
linear ; seed not furrowed, oil tubes very small. gregaria. 
Styles short, included; branching mostly alternate and dichotomous, sterile flowers 

not in separate heads; biennial at least as toc. 

_ ¢ Fruit small, under 2//, subglobose, short-pedicelled, scarcely reflexed ; bristles 
weak, under 1// long, dilated below; sepals on mature fruit inconspicuous among the 
longer erect bristles; pedicels of sterile flowers 1/’ or less; calyx deeply parted ; 
larger leaves mostly 5-divided; branching more or less decompound; commissural 
Scar narrow; oil tubes five, large; seed furrowed on dorsal surface ; seed-face convex. 

Canadensis. 

@, Fruit large, becoming 3 14’, ellipsoid, sessile, reflexed ; bristles stout, becoming 
1¥4/’ long; sepals on mature fruit forming a conspicuous beak-like projection free from 
the somewhat spreading bristles; pedicels of sterile flowers 2/’; leaves large, conspicu- 
ously trifoliate ; branching once or twice compound ; commissural scar broad ; oil tubes 
of two kinds, a pair of large ones and numerous small ones; dorsal surface of seed 
not furrowed; seed face concave. trifoliata, 
Plate 245 is a conventional representation illustrating the plan of branching and 


foliar arrangement of each of the four eastern species of the genus. 


362 


New Fungi, mostly Uredineae and Ustilagineae from various Lo- 
calities, and a new Fomes from Alaska. 


By J. B. ELtis AnD B, M. EVERHART. 


Fomes Tincrorius E. & E. 

(On hemlock trees or logs ?) 

Admiralty Island, Alaska. Collected by James G. Swan, no. 
20, 851. Comm. Frederick V. Coville, Botanist, U.S. Dept. Agr. 

Pileus dimidiate, sessile, subunguliform, convex below, slaty 
brown, witha few elevated concentric zones, the surface of which 
is more or less cracked, substance flocose-fibrous, deep red, some- 
what friable, superficial layer indurated, slate color, margin ob- 
tuse, 6-7 cm. long and broad, by 4-5 cm. thick. Pores large, I-2 
cm. long, 1%-2 mm. thick, same color as the inner substance of 
the pileus, filled with grumous matter and spores. Spores sub- 
globose or short elliptical, red, 5-6 x 31%4-4 4p. 

Resembles somewhat F. /ateritius Cke., but pores much larger 
and not stratose. When ground up it looks like madder, and 13° 
used for dyeing. 


UstiraGo ARENARIAE E. & E. 
In the inflorescence of Avenaria congesta Nutt., North Park, 


Colo., July, 1894. Alt. go00 ft. (Prof. C. S. Crandall, no. 119-) 

Spore mass purplish-black. Spores oblong-elliptical, opaque, 
14-17X 8-10, with a minute hyaline appendage at the base. 
The spores resemble the sporidia of Musmmularia Bulliardi Tul. 
Usti-aco Murrorpiana E. & E. 


Infesting and destroying the panicles of a species of Festuca 
_ while still enclosed in the sheath. Near Boise City, Idaho, June, 
1892. (A. I. Mulford.) , 


Mass of spores nearly black. Spores when moistened glo- 
bose or nearly so, 10-14 diam., epispore minutely roughened; 
when dry many of them become oblong or short-cylindrical, 10- 
14x 6-8 p. 

UsTILaGo MONILIFERA E, & E. 


In ovaries of Heteropogon contortus. Tucson, Arizona, May, 
1893. (Prof. J. W. Toumey, no. 2.) 


Mass of spores tobacco brown. Spores concatenate, lying 1” 
parallel chains, subglobose, subcubical or subangular, hyaline at 
first, then brown, 8-12y in the longer diameter, and very ™ 
nutely echinulate when highly magnified. 


363 


Sorosporium Souipacinis E. & E. 

Proc. Acad. Nat. Sci. Phil. February, 1893: 156. 

This is the same as S. cuneatum Schofield, in the 2d Ed. of 
Webber’s Appendix to the Cat. of Flora of Neb., published June, 
1892. 


Puccinta Licusticr E. & E. 

On leaves of Ligusticum scopulorum Gray. Sangre de Christo 
Mts., Colo. Alt. 10,000 ft. July, 1888. (Rev. C. H. Demetrio, 
no, 201.) 

III. Sori hypophyllous, minute, chestnut-colored, erumpent and 
surrounded by the ruptured epidermis, densely crowded in subor- 
bicular clusters 11%4-2 mm. diam. on small, pallid spots mostly 
near the margin of the leaf. Teleutospores, elliptical or oblong- 
elliptical, obtusely rounded at both ends, pale brown, scarcely or 
only slightly constricted, 22-30 x 15-20 yp, epispore slightly 
roughened but not distinctly thickened above. 


Puccinia NESAEAE (Ger.). 
Aecidium Nesaeae Ger. Bull. Torr. Bot. Club, 6: 47. 
On leaves of Nesaea verticillata near Concordia, Missouri. 


(Rev. C. H. Demetrio, no. 145.) 

III. Sori orbicular, minute, bordered by the upturned epidermis, 
collected in dense clusters or also scattered singly, usually clus- 
tered on the tubercular-thickened parts of the leaf previously oc- 
cupied by the Aeczdium. ee 

Teleutospores oblong, clavate oblong or oblong-elliptical, 
deeply constricted, strongly thickened at the apex, usually with a 
hyaline papilla, pale yellowish brown, 30-45 x 12-15 p. Pedicels 
Stout, subhyaline, about as long as the spore. 


Ravenetia Arizonica E. & E. 
On living leaves of Prosopis julifora. Tucson, Arizona, Aug., 


1894. (Prof. J. W. Toumey, no. 37.) 

Amphigenous; sori erumpent, soon naked, small, black; heads 
not compact in the sori, orbicular, 75-85 » diam., deep chestnut 
browh, hemispherical, spiny, marginal spores about 20 “18—25) in. 

number, inner spores about as many more, 18-22 x 7-8 #; number 
of spores in a cross section through the center of the head 7-9. 
Cystidia ovate-globose, swelling out so as to be visible around the 
Margin of the head viewed from above. Stipe short, straight, con- 
Sisting of only a few hyphae. The short, nearly hyaline spines 
are distributed over the entire surface of the head, about as in &. 
echinata D. & L., from which this differs in the larger orbicular 


364 


heads, containing a much greater number of spores. 4. Holwan 
Dietel on the same host, has no spines. Uredospores in the same 
sori with the teleutospores, obovate or elliptical, rough, 23-30 
15-18 », pale yellowish-brown. 


DOASSANSIA AFFINIS Ell. & Dearness. 


On Sagittaria variabilis. \ondon, Canada, July, 1895. (Dear- 
ness, 2269.) 


Sori hypophyllous, pustuliform, globose or elliptical, 200-300 p 
in the longer diameter, dark-colored and collapsing above, grega- 
rious in elongated groups, the part of the leaf occupied being at 
first yellowish. Spores globose or elliptical, 8-1o y in the longer 
diameter, epispore comparatively thin. 

Differs from D. Sagittariae on the same host, in its larger sori 
and smaller less angular spores with thinner epispore. From D. 
obscura Setchell it differs in habit and character of the sori; J. 
opaca Setchell differs in its larger spores as well as in some other 
respects. 


AECIDIUM SPHAERALCEAE E. & E. 


On leaves of Sphaeralcea angustifolia. Las Cruces, New Mexico. 
June, 1895. (Prof. T. D. A. Cockerell.) 
Hypophyllous, densely cespitose in clusters 2-8 mm, diam. 
Aecidia deep orange color, cylindrical, closed at first, then open, 
margin toothed, becoming entire, cups about % mm. high, 350- 
400 » broad. Spores subglobose or elliptical, smooth, 15-20 # 19 
the longer diameter, orange-yellow. Spermogonia on yellowish 
spots on the upper side of the leaf. 
Differs from A. Callirhoes E. & K. in its cylindrical aecidia 
and deep orange color. 


PERONOSPORA WHIPPLEAE E, & E. 


On leaves of Whipplea modesta, Ukiah, Mendocino Co., Cal., 
May, 1894 (W. C. Blasdale), 


Mycelium hypophyllous, effused, dirty gray. Fertile hyphae 
250-350 » high, 3-4 times dichotomously branched, the ultimate 
divisions simple or bifid and usually bearing on one side a short, 
straight, lateral branchlet 6-8 7 long. Conidia short-elliptical, 
brownish, smooth, 18-22x 12-15 ». Ovdspores not seen. 


This appears to be distinct from P. ribicola Schrtr., the only 
other species recorded on Saxifragaceae. 


365 


Studies in the Botany of the Southeastern United States.—IV, 
By JoHn K. SMALL. 


(PLATE 246.) 


SIEGLINGIA CHAPMANTI nN. sp. 

Wiry, slender, glabrous (except the tops of sheaths and parts 
of the inflorescence), bright-green, perennial by a horizontal root- 
stock. Culms solitary or two or three together, erect, strict, 9-15 
dm. tall, mostly purple about the nodes; lower leaves rather 
numerous, nearly erect, 4-6 dm. long, the upper few, divaricate, 
somewhat shorter, all firm, flat when young, soon involute and al- 
most filiform, 7—11-ribbed, smooth and glabrous; lower sheaths 
about 1 dm. long, upper ones often 2 dm. long, all %—% shorter 
than the internodes; ligule a short fringe of rigid villous hairs, 
above which, on the upper surface of the leaf, is a tuft of longer 
villous hairs; panicle averaging about 2 dm. high, viscid above, 
broadly ovoid, its branches rigid, filiform, divaricate (not drooping 
at the ends), the nodes tufted with bunches of silvery-villous more 
or less viscid hairs ; spikelets very slender-pedicelled, rather few, 
7-8 mm. long, tinged with purple, almost linear (when dry oblong) 
5-flowered; empty glumes lanceolate, one-nerved, the lower one 
34 longer than the upper; flowering glumes oblong-elliptic, 3- 
nerved, 3-pointed by the excurrent nerves which are villous for 
\% their length; palet 2-nerved, scabrous on the two nerves, 
slightly curved. 

A species of southern distribution separable from Steglingia 
seslerioides by its more slender and wiry habit, the very slender and 
diverging branches of the panicle and the usually conspicuous 
tufts of viscid hairs at the nodes throughout the panicle. 

I found it last season growing in sand on the slopes of Cur- 
rahee Mountain, near Toccoa, Georgia, and along theYellow River, 
Gwinnett County, in the same State. Its range may be given 
as follows: Georgia, as just cited, to Texas: J. Reverchon ; Bige- 
low, Camp No. 4, between Ft. Smith and the Rio Grande, south 
to Florida: Chapman, Curtiss (3454) Duval County. 

Quercus GeorciAna M. A. Curtis; Chapm. Fl. S. States, 422 
(1860). Although heretofore supposed to be confined to a single 
Station and to exist only in shrub form, Q. Georgiana is now 
known to have a considerable range and also to reach a develop- 
ment which allows it to be classed as a tree. 


366 


Stone Mountain, Georgia, is the original locality of this 
species. There it is more plentiful than at any of the new sta- 
tions, but seldom attains the form of a tree. During the last two 
years I have found this interesting oak at four new places, and at 
all of these it grows plentifully and the tree form predominates 
instead of the shrub. 

Along the Yellow River, about six or seven miles east of Stone 
Mountain, there are several large groves on the sides of the cafions 
where the granite rocks outcrop. Six or seven miles east of the 
Yellow River there occurs another outcrop of granite, and here 
two fine trees stand and bear fruit in great abundance. Travel- 
ling between the last mentioned station and the Oconee Mountain, 
a little to the south of east, I came upon another granite outcrop 
just west of the Oconee River. There the species again appears, 
and it is interesting to note that whenever found it is confined to 
granite outcrops. A fifth, or fourth, new locality is Little Stone 
Mountain, situated about nine miles south of Stone Mountain. 
There the species reaches its greatest development in size, meas- 
urements showing a height varying from twenty to thirty feet 
and a maximum trunk diameter of fourteen inches. The follow- 
ing map will give an idea of the geographic distribution of this oak. 


GWINNETT 


NXE Gatres S117 


Granite outcrop & 
Granite oulcrop 


WALTON 


Stone / & 


DE KALB 


Little Stone (te. 


eT 


PHORADENDRON FLAVESCENS (Pursh) Nutt.; A. Gray, Man. Ed. 2, 
383. 1856. 


This plant is quite plentiful in the vicinity of Stone Moun- 


367 


tain, Georgia. At all the places where I found it in that region 
its host was Quercus Georgiana, although there were numerous 
other deciduous-leaved trees growing with the latter species. 


‘RUMEX FASCICULARIS n. sp. 

Perennial by a cluster of fusiform tuberous roots, glabrous, of 
a dull, rather dark or olive-green color. Roots in clusters of from 
3-5, 5-10 cm. long; stem lax and weak, 5-6 dm. long, de- 
cumbent and ascending, strongly grooved, abruptly thickened at 
the base and slightly thickened at the nodes; internodes 1-6 cm. 
long; leaves mostly oblong, occasionally a few oblong-ovate or 
Ovate, 7-17 cm. long, 4-7 cm. broad, of much the same size 
throughout and clustered at the shortened end of the stem near 
the inflorescence, acute or obtuse, somewhat undulate and crisped, 
mostly truncate or cordate at the base, sometimes obtuse, thick 
(drying very thin), petioled; petioles stout, 3-7 cm. long, strongly 
dilated at their bases; ocreae thin and brittle, fugacious; panicle 
12-16 cm. long, rather dense in fruit; racemes ascending, 1-5 cm. 
long; pedicels slender, narrowly clavate, winged at the summit, 
deflexed in fruit, 1-2 cm. long, articulated at the base; wings of 
the calyx broadly deltoid, 4 mm. long, 5 mm. broad, undulate, 
strongly nerved, each bearing an ovoid rugose callosity: style- 
segments reflexed on the angles of the achene; achene broadly 
Pyramidal-ovoid, triquetrous, 2 mm. long, chestnut-colored, 
smooth and shining, the faces concave, the angles slightly mar- 
gined. Plate 246. 

A peculiar and striking species on account of its clustered 
leaves. The shape of the latter is different from that of any other 
North American member of the genus. By its inflorescence, 
wings and achene it is related to R. Floridanus and R. verticillatus, 
but the achene is broader and shorter than that of either of those 
two species. The stem is unable to support the weight of the 
leaves and inflorescence and bends over, endeavoring to rise again 
at the end. 

The specimens were collected by Mr. Geo. V. Nash, on his re- 
cent excursion to Florida, in cypress swamps, on the marshy shore 


of Lake Harris, near Eldorado, in the central part of the peninsula. 


“ACER LEUCODERME nN. sp. 
Acer Flovidanum acuminatum Trel. 

A shrub, or small tree reaching a height of eight meters and 
a trunk diameter of from one to five dm., clothed with a smooth 
white bark. Trunk very short, sometimes almost wanting; 
branches (secondary trunks), two to eight together, erect or as- 


368 


cending, conspicuous by their white bark ; branchlets clothed with 
a gray or reddish bark; leaves depressed-orbicular (2. é., broader 
than high), or rarely orbicular in outline, mostly three-lobed, 
sometimes imperfectly five-lobed, 4-9 cm. in diameter, cordate or 
truncate, petioled, with a rather open and shallow sinus, dark green, 
glabrous and marked with light nerves above, greenish, tinged 
with red, prominently nerved and very velvety (to the touch) be- 
neath, the lobes acute or acuminate (the 4th or 5th when present 
obtuse), each (or the terminal one only) bearing two obtuse teeth ; 
petiole slender, reddish, 3-6 cm. long; flowers not seen; wings 
of the samaras oblong-spatulate, 1-2 cm. long, red, conspicuous, 
parallel or nearly so (more or less spreading when the fruits sepa- 
rate at maturity); seed oblong, its covering prominently veined. 


A very handsome maple, characteristic on account of its habit 
of branching near the base into from several to many second- 
ary trunks, and the white bark. The bright green color of its 
foliage and the usually bright red fruit render it conspicuous, 
and while the velvety pubescence on the lower surfaces of the 
leaves is not prominent it is remarkably soft and dense to the 
touch. ; 

As far as I have observed, the tree is confined to the bottoms 
of two rocky cafions, that of the Yadkin River, in Stanley county, 
North Carolina, and especially that of the Yellow River, in Guitt- 
nett county, Georgia. Dr. Trelease has reported the tree from 


further south, where it doubtless occurs, but I do not know the 
character of the localities. 


KOELLIA VERTICILLATA (Michx.) Kuntze, Rev. Gen. Pl, 520. 1891. 
Pycnanthemum Torreyi Benth. Lab. Gen. & Sp. 329. 1834: 
This species of Koellia has never been recorded as growing 

further south than Southwestern Virginia,* and consequently it has 

not been credited to the Southern Flora. In August, 1893, I 

found the plant growing in Northern Georgia, in Rabun county, 

near Estotoah Falls. It was quite plentiful in the valleys and 
ravines, at about 2000 feet altitude and inhabited localities much 
like those in which it was found in Southwestern Virginia. 


/ SOLIDAGO YADKINENSIs (Porter). 
Solidago Boottii var. Yadkinensis Porter, Bull. Torr. Club. 1892: 
Perennial, slender, wand-like, glabrous and of an olive-green 
* Mem. Torr. Club, 4: 146. 


369 


color throughout. Rootstock long, chaffy, horizontal ; stem 5—15 
dm. long, erect, simple below the inflorescence ; basal leaves tufted, 
lanceolate or linear-lanceolete, the blade 10-30 cm. long, acute at 
_ the apex, acuminate at the base, the petiole 0-15 cm. long, winged ; 
cauline leaves lanceolate, linear-lanceolate or linear-oblong, 3-15 
cm. long, erect and appressed, acute at the apex, sessile, the lower 
ones like the basal, serrate with a few distant appressed or spread- 
ing teeth ; inflorescence consisting of a simple terminal secund ra- 
ceme or thyrse; heads campanulate, stalked, 3-4 mm. high, 25-35 
flowered ; involucral bracts in four to five series, oblong or linear- 
oblong, 1-2 mm. long, ciliate, obtuse, with a dark-green midrib 
and tip ; corolla slightly longer than the pappus, more or less pubes- 
cent; rays yellow, oblong-spatulate, entire, 2—3-apiculate ; achene 
columnar, obtuse at the base, 10-14-ribbed, pubescent with a few 
Spreading hairs. 

This species was first described as a variety of Solidago Boottit 
by Prof. Porter, from collections made by Mr. Heller and myself 
in middle North Carolina in 1891. On first meeting with the 
plant it seemed to me a good species, and field observations on it 
each succeeding season have convinced me of its specific validity. 
It grows only in rather open meadows scattered through the 
pine woods. The first specimens seen were of the simple type 
(z.é, in which the inflorescence consisted of a simple terminal 
raceme) and not over six dm. tall. They were found near Gold 
Hill, N.C. Last season I found this type near the base of Dunn’s 
Mountain, in the vicinity of Salisbury, N.C. However, a more 
robust and branched form is the more common state, and this oc- 
curs at many localities in middle North Carolina. Up to last 
Season Solidago Vadkinensis was not known to grow outside the 
last mentioned region, but in September (1894) I met the plant 
growing luxuriently at two stations in middle Georgia; the one a 
botanically prolific meadow near Loganville, Walton county, and 
the other meadows near the base of Little Stone Mountain, De- 
Kalb county. The Georgia specimens are a little more robust 
than those from North Carolina, but otherwise they are almost 
identical. The variation that does exist is due to the less exposed 


conditions under which the Georgia plants grew. 


370 


Botanical Notes. 


Tumble mustard. A species of mustard that promises to be 
one of the most formidable tumbleweeds yet introduced in the 
United States has been found by Mr. J. H. Sandberg and Mr. J. 
M. Holzinger, well established as a roadside weed in the side 
streets of Minneapolis. It is Sisymdrium altissinum L. This 
species was collected on ballast ground at Philadelphia in 1878, 
and in 1885 it was found near Castle Mountain on the western 
boundary of Alberta. During the past five years it has become a 
troublesome weed in the vicinity of Indian Head, Assiniboia. It 
is there known by the very appropriate name of “tumble mustard.” 
It has also been found at Ottawa, Canada. 

The tumble mustard may be distinguished from the other 
mustards by its slender siliques, 3 to 5 inches long, and by its 
characteristic tumbleweed habit, making a rather dense bush-like 
plant at maturity. If discovered elsewhere than in the five places 
mentioned, the undersigned will be grateful for reports of the lo- 
calities. In case of doubt as to determination franks will be sent 
for mailing and the specimens will be determined at the National 
Herbarium. Lysrer H. DEWEY. 

WasuincTon, D. C., August 14, 1895. 


Reviews. 


A Manual for the Study of Insects. John Henry Comstock 
and Anna Botsford Comstock. 8vo., pp. 7o1. Ithaca, N. Y- 
Comstock Publishing Company. 1895. 

The entomological sky has become much cleared, so far as 
the student or beginner is concerned, by the publication of this 
Manual for the Study of Insects. There should be no difficulty 
now in determining in nearly every case the native insects as far 
down the scale of classification as the families to which they 
scientifically belong. This is made possible by numerous figures 
and tables of classification for each order, coupled with a general 
clearness of statement that does not bristle with an array of 
scientific terms. The book is essentially about insects and not 


371 


their describers, for no authorities are given. This matter is no 
doubt left to the check lists and other more special works. 

The first chapter deals with classification and nomenclature; 
the second with crabs, spiders, centipedes and other near relatives 
of the true insects, and the remaining nineteen with the six leg- 
ged insects or Hexapoda, each chapter being devoted to a sepa- 
rate order. This number of orders, as Prof. Comstock says, 
slightly exceeds that commonly adopted. The difference, as will 
be seen by the reader, results from dividing the Platyptera of 
recent classifications into four separate orders. Those adopted in 
the work are the following: 


RTORNORA ee Bristle-tails, Spring-tails. 
PPUMMERIDA 5 33+ 4s | May-flies. 

2s a hi re Sia Dragon-flies, 

PERCOPTERAS 2) 2.8 4" . . . Stone-flies. 

NSORPTER A C8 st Peck tee White-ants. 

WORRODENTIA® | .' 5 2 oy ges Procids and Book-lice. 
WEATLOPHAGAT Eo Sore oe Bird-lice. 

EUPLEXOPTERA®, 22 8 Earwigs. 

ORTHOPTERA. . . . . . . . Cockroaches, Crickets, Grasshoppers, etc. 
NNN Fe ary Thrips. 

RIERA Bugs, Lice, Alphids and others. 
WIROROPTEBA oo 5 SS Aphis-licris, Ant-licris and others. 
MOP TERA a Scorpion-flies, etc. 

SeiOHOPTERA 5 Caddice-flies. 

PRPIDPIBRA (ss se ee Moths and Butterflies. 

PRR Flies 

DEPHONAPTERA 2 Sy, Fleas 

CA MNTERA Go a Beetles. 

ELYMENOPTERA 6 ok SS Bees, Wasps, Ants and others. 


It is not supposed that this sequence is altogether a natural 
One, but the best that can be expressed in a linear series, and 
indeed, it does not make any difference whether, for instance, the 
Lepidoptera immediately precede or follow the Diptera. 

The greatest space is given to the chapter on Lepidoptera, 
which occupies 222 pages, and it is chiefly in this much collected 
order that changes of classification are noticed. They are based 
upon the venation of the wings; the character of the antenne, 
Vestiture, etc., not holding so prominent a position as in other sys- 
tems. This classification seems to be very good so far as clear- 


372 


ness is concerned, and is easily comprehended by the general 
reader, but perhaps specialists will not take to it so kindly. 

In the chapter on Coleoptera the tiger beetles are mentioned 
first and the Scolytidae last. In some recent classifications running 
from the lowest to the highest this system is reversed. 

The book is finely illustrated, containing 797 figures and six 
plates, the frontispiece (Plate I.) being colored. On page 154 is 
a picture of four grotesque leaf-hoppers sitting in a row on a grass 
blade, who rival in absurdity of expression the famous Brownies. 
Some of the illustrations include the plants on which the insects 
feed, and the food plants are also often mentioned in the text. 
Entomology as a rule does not form a conspicuous part in books 
on botany, but the botany of an entomology is a more important 
matter, many of the insects taking their names from the plants 
upon which they feed. The illustrations of the wing-veins are 
particularly numerous, and it is these, and the tables for determin- 
ing the families and higher groups, that contribute largely to the 
clearness of the book. W. T. Davis. 

Studies in Plant Deackimen Henry L. Clark. Chicago. 
1895. 

One is at a loss to know just what to say on examining this 
outline of plant analysis, for such it is, neither more nor less. 
Judging from its mechanical construction and arrangement one 
would conclude that it was intended to serve as an aid in teaching 
young children the first principles of systematic arrangement, but 
the terminology, which is that of Vines (see his recent text-book 
on botany), will at once condemn it for such a purpose. The 
terminology used would lead one to suppose that it was intended 
for the use of advanced students and specialists in botany. Any 
one sufficiently advanced to comprehend the meaning and applica- 
tion of such terms as gametophyte, apogamy, germ-plasm, body- 
plasm, microspore, macrospore, etc., etc., certainly does not require 
an outline in the study of plant types. It is detrimental to teach 
students that such outline-study is botany. It is true, the author 
recommends a “ constant use’’ of Vines’ Text Book of Botany in 
connection with this outline work, which is good as far as it goes. 
It is, however, wholly wrong to teach students that they can be- 
come systematists without having first studied morphology and 


373 


physiology. This prevailing idea has already done an inestima- 
ble amount of harm to the advance of botany. No student « may 
quickly be taught the small amount of technique (of the compound 
microscope) necessary to do outline work.” It requires years of 
patient toil to learn how to use thecompound microscope. Any per- 
son can look down the tube of a compound microscope, but whether 
said person knows and comprehends what he sees, or whether he 
sees anything at all, is another question. My advice is that the 
student should lay aside the «‘ Studies in Plant Development” un- 
til he has mastered the rudiments of plant morphology and physi- 
ology by years of patient work in some well equipped laboratory 
and by consulting not only Vines but all other standard authori- 
ties on plant physiology and morphology. After this is accom- 
plished the “Studies” may be permanently stored away in the 
garret along with other similar analyses. 

As far as the make-up of the “ Studies”’ is concerned it lacks 
nothing. Binding, paper and print are excellent. There is space 
for the “study” of fifty types. A short list of modern (Vines) 
botanical terms with etymology and meaning is added. In his 
classification we are pleased to note that lichens are recognized as 
adistinct class. It may be that the “ Studies” could be made use of 
by advanced students in our universities, provided it was thought 
advisable to take the time for making the records. Of what use 
these records might be afterwards is more than we can understand 
at present because there are already printed records of all the 
known plant species. 

It is with regret that we make this adverse criticism, but we 
firmly believe that any mistaken tendency should be checked 
early. The mistaken idea of what constitutes plant study has 
already gained such a strong footing that we must put forth every 
effort to check its progress. Let us sincerely hope that pernicious 
“plant analyzing” may never be substituted for Botany in our 
higher institutions of learning. Let us hope that in the future the 
author will expend his time and talent on some work which will 
Prove a real benefit to the advance of botanical science. _ 

ALBERT SCHNEIDER. 


374 


Index to recent Literature relating to American Botany. 


Atkinson, G. F. Some Observations on the Development of Cod/eto- 
trichum Lindemuthianum in artificial Cultures. Bot. Gaz. 20: 305-311. 
pl. 22. 15 Jl. 1895. 


Avé-Lallemant, Prof. Briefe aus Argentinen. Mitteilungen d. 
Geogr. Gesellsch. u. d. Naturhist. Mus. i. Liibeck, 7: 53-91. 1895. 


Contains a few notes on the flora. 


Bailey, L. H. The Plant Individual in the Light of Evolution. Sci- 
ence (p. 8.)) 1: 281-292. 15 Mr. 1895. 


Bates, J. M. A new Astragalus. Am. Nat. 29: 670, 671. Jl. 1895- 


A, lotiflorus Hook. var. Nebraskensis, from Brown County, Nebraska. 


Beal, W. J. Notes from a Botanic Garden.—I. Gard. & For. 8: 
20%.:. 31 J1, 1898. 
Notes from Agricultural College, Michigan. 


Beal, W. J. Teaching Botany one Topic at a Time, illustrated by 


suitable Materials at any Season of the Year. Science (n. s.), I: 355: 
29 Mr. 1895.. 


Bennett, A. Notes on the Pofamogetons of the Herbarium Boissier. 
Bull. Herb. Boiss. 3: 249-260. Je. 1895. 


Memoranda ona number of American species. 


Bessey, C.E. Some Features of the native Vegetation of Nebraska. 
Am. Nat. 29: 486,487. My. 1895. 


Bessey, C. E. The Division of Agrostology. Am. Nat. 29: 487) 
488. My. 1895. 


Boyer, C. S. A Diatomaceous Deposit from an Artesian Well at 
Wildwood, N. J. Bull. Torr. Bot. Club, 22: 260-266. 25 Je 
1895. 


Describes new species in Hydrosera and Surirella. 


Britton, N.L. Daniel Cady Eaton. Science (n. s.), 2: 57, 58- 19 
Je. 1895. 


Britton, N.L. The generic Name of the Water Weed. Science (0- 
S.), 2: 5. 5. Jl. 1895. 
Proposes Phtlotria Raf. 1818, for Udora Nutt. 1818, but later. 

Bush, B.F. A list of the Trees, Shrubs and Vines of Missouri. Rep- 
State Hort. Soc. 37: 353-393. Je. 189s. Reprinted. 


375 


Clarke, C. B. On certain authentic Cyperaceae of Linnaeus. Journ. 
Linn. Soc. London, 30: 299-315. 6 O. 1894. 


Contains numerous notes on American species. 


Curtis, C. C. Interesting Features of well-known Plants of New York 
Harbor. Journ. N. Y. Micros. Soc. 11: 63-73. pd 50, 57. Ji. 


1895. 
Davy, J.B. Plants hitherto undescribed.—I. Erythea, 3: 116-118. 
Describes new species in Lupinus, Senecio and Triglochin. 


Didrichsen, A. Om Tornene hos Aura crepitans. (Sur les épines de 
! Hura crepitans), Bot. Tidskr. 19: 189-200. 1895. 


Eastwood, A. On heteromorphic Organs of Seguota sempervirens. 
Proc. Cal. Acad. (II.) 5: 170-176. pl. 15-18. 13 My. 1895. 

Eckfeldt, J. W. An Enumeration of the Lichens of Newfoundland 
and Labrador. Bull. Torr. Bot. Club, 22: 239-259. 25 Je. 1895. 
New species in Pannaria, Lecanora, Biatora and Lecidea, 

Fernald, M. L. A red-seeded Dandelion in New England. Bot. Gaz. 
20: 323,324. 15 Je. 1895. 
Note on 7araxacum erythrospermum. 

Fernald, M. L. Supplement to the Portland pone of Maine 
Plants. Proc. Portland Soc. Nat. Hist. 2: 73-76. 1895. 

Fernald, M. L. Two new mountain Plants. Bull. Torr. Bot. Club, 


22: 273,274. 25 Je. 1895. : 

Describes Aster Hendersoni and Carex scabrata X crinita new hybrid, 

Ganong, W. F aLboratory Teaching of large Classes in Botany, 
Science (n. s.), 1; 230-234. 1 Mr. 1895. 

Gerard, J. N. American /rises. Gard. & For. 8: 286. 17 Jl. 1895. 

Goodale, G. L. The New York Botanic Garden. Science (n. s.), 2 
1,2. 5 Jl. 1895. 

Greene, E. L. Novitates Occidentales—XV. Erythea, 3: 118-121. 
I Jl. 1895. : 
Describes new species in De/phinium, Botsduvalia, Aster, Linanthus and Sisy- 

rinchium, 

Greene, E. L. Observations on the Compositae—X. Erythea. 3: 
YO7—115,-- 1 Jl. 1895. 

Enumerates the species of Cérysothamnus ; establishes the genus Neosyris for 

Sigelovia hypoleuca and B, fuliginea. 

Harshberger, J. The Origin of our vernal Flora. Science (n. s.), 


Ti 92-98. 25 Ja. 1895. 


376 


Heritage, B. Preliminary Notes on Welwmbo /utea. Bull. Torr. Bot. 
Club, 22: 265-271: .p/. 2377. 25 Je. 1865. 


Hill, E. J. The Saguenay Region—I.-III. Gard. & For. 8: 182- 
183; 193; 213. My. 1895. 

Holm, T. On the Validity of some fossil Species of Liriodendron. 
Bot. Gaz. 20: 312-316. p/. 23. 15 Jl. 1895. 


Hooker, 1; D. “Senecio Hualtata. Curt. Bot. Mag. fl. 7422. ji. 


1895. 
Native of Chili and Argentaria. 


Jelliffe, S. E. Cryptogamic Notes from Long Island.—III. Bull. 
- Torr: Bot.Club, 22: 274~275. “25 Jé. 1895. 


Kellerman, W.A. Ohio Forest Trees. Pamph. pp. 16. Columbus. 
1895. 


Keys to and descriptions of the native species. 


Kellerman, W. A. Poisoning by Shepherd’s-purse. _ Bot. Gaz. 20: 
325, 326. 15 Jl. 1895. 

Kellerman, W. A. Spring Flora of Ohio. 8vo. pp. 128. ‘Colum- 
bus, 1895. 


A concise descriptive flora. . 


Knowlton, F. H. Use of the initial Capital in specific Names of 
Plants. Science (n.s.), 1: 423, 424. 19 Ap. 1895. 

Kranzlin, F. Eine neue Pleurothailis-Art. Bull. Herb. Boiss. 3! 
359, 360. Jl. 1895. 


P. Autraniana n., sp. 


Leiberg, J. B. On the Carpels of Dtecdaiie malvacea (Greene) 
Bull. Torr. Bot. Club, 22: 271,272. .25 Je, 1895. 


Lemmon, J. G. Handbook of West American Cone-Bearers. Ed. 3: 
Pp- 104. Jigs. Oakland. 1895. 


Masters, M.T. The Guadeloupe Cypress. Gard. Chron, (III) 18 
62. f. 9. 20 Jl. 1895. 

- Description and illustration of Cypressus macrocarpa Guadeloupensis (S- Wats.) 

Mast. 

Minot, c 5 The fundamental Difference between Animals and 
. Plants. Science (n. s.), 1: 311, 312, 22 Mr. 1895. 

Mottier, M. D. Contributions to the Embryology of the Ranuncula- 


ceae.» Bot. Gaz. 20: 241-248. pl. 17-20. 17 is ate 15 
Jl. 1895. 


377 


Pammel, L. H. Conifers in Iowa. Gard. & For. 8: 279. 10 Jl. 
1895. : 
Patouillard, N. Aplittopsis, nouveau geure d’Hyménomycetes 
hétérobasidiés. Journ. de Bot. g: 245-247. 16 Jl. 1895. 

: Description of the genus, and of 1/7. Langloisit from Louisiana. 

Pollard, C. L. Viola sagittata Hicksit, var. nov. Bot. Gaz. 20: 326. 
15 Jl. 1895. 

Robertson, C. Harshberger on the Origin of our vernal Flora. —Sci- 
efice (N.:s.), £:.371~375. 5 Ap. 1895. 

Sargent, C.S.  Carpinus cordata. Gard. & For. 8: 294. f. ¢J. 
24 Jl. 1895. 

Sargent, C. S., Editor. A northern Forest. Gard. & For. 8: 282. 
J gO. - 17.31. 1805; 

Sargent, C.S., Editor. <Xadmia latifolia var. myrtifolia. Gard. & 
For. 6:..315. f. gg. 7 Aus 1895. 

Sargent, C. S., Editor. The Tree Yuccas in the United States. Gard. 
m@ For. 8: 301. f. 2. 31 Jl. 1898: 


Illustrating Y. macrocarpa. 
Smith, E.F. The Botanical Club Check List. Pamph. pp. 16. No 
date. 

Smith, Y.D. Undescribed plants from Guatemala and other Central 
American Republics. —XV. Bot. Gaz. 20: 281-295. 15 Jl. 1895. 
Tracy, W. A. The simultaneous Origin of similar (or identical) — 

Varieties from different Stock. Am. Nat. 29: 485, 486. My. 1895. 
Toumey, J. W. Notes on the Tree-Flora of the Chiricahua Moun- 

tains—I.-II. Gard. & For. 8: 12, 13; 22. figs. Ja. 1895. 
Trelease, W. The Missouri Botanical Garden. Am. Nat. 29; 668- 

670. Jl. 1895; Science, n. s. 1: 716, 717. 28 Je. 1895. 
Trimble, H. On the tanning Properties of the Bark of three North 

American Trees. Gard. & For. 8: 293. 24 Jl. 1895. 

Notes on the tannin of Castanopsis chrysophylla, Quercus densiflora and Ostrya 
Virginiana, 

Underwood, L. M. An interesting Zguisefum. Bot. Gaz. 20: 320. 
(figs. 15 Jl. 1895. 

Abnormal form of £. hyematle. 
Vail, A. M. The June Flora of a Long Island Swamp. Gard. & 

For. 8: 282. 17 Jl. 1895. 


378 


Vance, L. J. The Future of the Long-leaf Pine Belt. Gard. & For. 8: 
278. 10 Jl. 1895. ; 


Ward, L. F. The Mesozoic Flora of Portugal compared with that of 
the United States. Science, n. s., 1: 337-346. 29 Nov. 1895. 


Williams, T. A. Notes on Mexican Lichens—I. Am. Nat. 29: 
480-485. My. 1895. 


Wolley-Dod, C. Cone-flowers. Garden, 47: 418-419. pl. roré. 
15 Je. 1895. 


Descriptions of the cultivated Rudbeckiae, with illustrations of R. maxima and 
R. pinnata. 


Surplas Books for Sale from the Library of Dr. M. C. Cooke. 


146 Junction Road, London N., England. 


Balfour, J. B. Botany of Socotra, 4°. 100 plates. Edinburgh. 
Hedwig. Theoria generationis and Hist. Nat. Muscorum. 4°. 
Lamarck and DeCandolle. Flore Francaise. 6 vols., 8°. 
Steudel. Nomenclator Botanicus. 8°, board. 
Linnaeus. Flora Suecica. First Edition. 
Leers. Flora Herbornensis. 16 pl., 8° 
Acharius. Synopsis Lichenum. 
Marsilius. Dissert: de generatione Fungorum. 
Retz, Observationes Botanicae. Folio. 
Vaillant. Botanicon Parisiense. Folio. 
Queckett. Histological Catalogue. 4°. 2 vols., cloth. 
Luerssen. Medico-Pharmaceutik Botanik. Vol. 1, Kryptogamen, 8°, paper. 
Lindley, Dr. Flora Medica. 1 vol., cloth. 
Waring, Dr. Pharmacopaeia of {ndia. 8°, cloth. 
_ Fluckiger, Dr., and Hanbury, D. Pharmacographia. 8°. 
Hanbury, D. Notes on Chinese Materia Medica. 
Hanbury, D. Science Papers. 1 vol., 8° 
Waring, Dr. Bibliotheca Therapeutica. 2 vols., 8°. 
Christison, Dr. On Poisons. 8°, 1 vol. 
Allen and Thomson. Expedition to the Niger. 8°, 2 vols. 
Farlow, W. G. Marine Algae of New England. 8°, paper. 
Toni and Levi. Flora Algologica Venezia. Parts I. and III. 
Toni and Levi. L’ Algarum Zanardini. 8°. 
Underwood. Catalogue of N. Am. Hepaticae. 8°. 
Tuckerman, Synopsis of Lichens of New England. 
Bescherelle. Prodromus Bryologiae Mexicanae. 8°. : 
_, Harvey, Dr. Phycologia Britannica. 4 vols. in 2 half morocco, gilt tops. Splen- 
did copy, first issue. te 
: rs English Botany, 2d Edition, Vol. XIL, only containing Algae. 8°, 
cloth. 
Goebel, K. Morphologische und Biologische Studien. 8°, 15 plates. 
Queckett, J. Lectures on Histology. 2 vols., 8°, cloth. 
Queckett, J. Treatise on the Microscope. 8°, cloth. 
Humboldt. Flora Friburgensis. 4°, board. 
Ardissone. Enum. della Alghe della Marca di Ancona. 4° 
De Notaris. Epilogo della Briologia Italica, Roy. 8°, half calf. 
Wahlenberg. Flora Suecica. 2 vols. 8°. 
Bauhin. Theatrum Botanicum. Folio. 
Meyer. Primitiae Florae Essequeboensis. 4°, calf. 
Sullivant. Musci and Hepaticae of U.S. 8° : 
Kickx, J. Flore cryptogamique des Flandres. 2 vols. in 1, Half calf. 1867, 
Roth, A. G. Catalecta Botanica. 3 vols., 8°, half calf. 1806. 
Schrader, H. Spicilegium Florae Germanicae. 8°. 1794. 
Haller, A. Flora Jenensis. 12°. 1745. 
Weigel. Flora Pomerano Rugica. 12°. 1769. 
Olhaff. Elenchus Plantarum. 12°. 1656. 
Sprengel, C. Florae Halensis. §°. 1806 and 2 suppl. 
Schumacher. Plantae Saellandiae. 2 vols., 1801. : 
Schlechtendal. Flora Berolinensis, Part 2, Cryptogamia. 1824. 
Wulff, J.C. Flora Borussica, 12° 1765. 
Schrank. Florae Salisburgensis. 12. 1792. _ 
Weinmann. Enum: Stirpium in Agro Petropolitano. 8°. 1837. 
Martius. Flora Erlangensis Crypt. 8, 6 pl., 1817. 


‘ 


} 


Nuovo Giornale Botanico Italiano, 1872 to 1891. 20 years complete, 18 in 9%, 
calf, rest in parts. ! 

Michelia. Ed. P. A. Saccardo, 1879 to 1882 in 2 vols., half calf (all published). 

Hedwigia (Ed. Rabenhorst, etc.), 1852 to 1891 (Vol. II. imperfect) together in 
15 vols., cloth). 

Flora (Regensburg), vols. 34 to 47, 14 vols., cloth, : 

Notarisia, Commentarium Phycologiae, 1886-1891. Parts 1 to 25 (wanting part 
21). 

) La Nuova Notarisia. 1890-1891. 7 parts, 
Revue Mycologique (Roumeguere). 1888-1891. 13 vols. in 6, cloth. 
Rabenhorst’s Algae Europaea Aquae dulces. 3 parts in 2 vols., half calf, com- 
plete, 1864. 

Monthly Microscopical Journal, (Ed, Dr. Lawson.) 18 vols., cloth. 1869-1877. 

Grevillea (Ed. M, C, Cooke), Complete set in parts, 20 vols, 1872-1892. 
Complete sets out of print. 

Hassalls, Dr. British Fresh Water Algae. 2 vols., cloth, 

Smith, W. British Diatomaceae. 2 vols., cloth. 

Ralfs, J. British Desmidiae. 1 vol., half calf. 

Bornet et Thuret. Notes Algologiques, part 2 only, folio, plates. 

Flora of Herefordshire. Mosses by Aug. Ley. Fungi by M. C. Cooke, 1 vol. 
8, cloth. 1889. 

Hooker, W. J.,and Baker, J. G. Synopsis Filicum. Colored plates, 8°, cloth. 
1868. 

DeBary, A. Morphologie und Biologie der Pilze. Mycetozoen und Bacterien. 
8°, uncut. 1884. 

Natural History Review. Vols. I. to VI., 1854 to 1859 and 1861 to 1865, II 
vols. (wanting Vol. VIL.). 

American Naturalist, 1867 to 1886 (wanting 1876), in 17 vols., half calf, 2 vols 
in parts. Also 1887-8 complete; 1889 (wanting 266, 270, 274, 275); 1890 complete; 
1891 (wanting 298, 300). 

American Entomologist, and American Botanist and Entomologist. 1868-1879. 
2 vols. in one, half calf. 

Ray (John). Historia Plantarum. 3 vols., folio. ; 

Mitten, W. Musci Indiae Orientalis, 8°, with Griffith’s Muscologia Itineris AS 
ora pe Griffith’s Notulae ad plantas Asiaticas, part 2. Cryptogamia in 1 vol., 8°, 

calf. 

Buxbaum J.C. Plantarum minus cognitarum. Complete in 2 vols, 4°. 

. __ Girod Chautraus. Recherches chimiques et micro, sur les Conferves. Fol. 36 pl. 
half calf. 1802. 


Gerarde’s Herbal. Old folio, wanting title and last page of index, otherwisé 
good condition. 


Botanical Tracts, chiefly Cryptogamia, many vols, requiring separate list. 
Greville’s Scottish Cryptogamic Flora. 6 vols., half morocco, gilt. Colored 


plates, 
Many other works, specially on Fungi, not enumerated here. 
In applying for details of price, etc., quote the headings here written. 


Lonpon, July, 1895. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 239. 


NEW AND RARE DESMIDS. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 240. 


-O G28. 


BQ0 


oe Oxy 


020° BLO:0 


NEW AND RARE DESMIDS. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 241. 


SANICULA MARYLANDICA L. 


Co ee ~— 


BULLETIN OF THE TORREY BOTANICAL CLUB—.PLATE 242. 


SANICULA GREGARIA BICKNELL. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 243. 


SANICULA CANADENSIS L. 


— 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 244. 


SANICULA TRIFOLIATA BICKNELL. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 245. 


ie 
"_ 

Le. 
<4 5 Nx 


BRANCHING OF EASTERN SANICULAS. (DIAGRAMMATIC,) 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 246. 


f ) UN RY . 

ANAS Nie \ SS \ 
BD = CaN 

ESR JX 


oe 


.y 
SARIS 
SS Se 
SE NESS 
ny NS » a 1; 
Shy oe Le, 
S . 


> Sk 
rae BN 
y ANS 


RUMEX FASCICULARIS SMALL. 


— 


Contributions from the Herbarium of Columbia College. 


No. 37. 
No. 38. 


No. 39. 


No. 40. 
No, 41. 


[The numbers omitted from this list are out of print, ] 


VoLuME I. 
A List of Plants Collected by Miss Mary B. Croft at San Diego, Texas. By 
N.'L, Britton and H..H Rusby (18$7)5 0. ok oe wea 25 cents. 


New or Noteworthy North American Phanerogams. By N. L. Britton 
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An Enumeration of the Plants Collected by Dr. H. H. Rusby in South 
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A List of Plants Collected by Dr. E. A. Mearns at Fort Verde and in the 
ss See and San Francisco Mountains, Arizona, 1884-1888. By N. L. 
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The General Floral Characters of the San Francisco and Mogollon Mountains 


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Preliminary Notes on the North American Species of the Genus 7?ssa, 
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A Descriptive List of Species of the Genus Heuchera. By Wm. E. Wheel- 
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CSCO E ie hea: weet oh & no oe ee enemas core 25 cents. 
Notes onthe North American Species of Eriocaulee. By Thos. Morong 
RUNGE, os chy Seg ie ee at Gy + cal ot . . . 25 cents. 
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ee he ee ee ee ee 


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La apie mie eee Ua pees ets? 


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VoLUME IV. 


No. 76. The Biological Status of Lichens. By Albert Schneider (1895), . 25 cents. 
No. 77. New or Noteworthy North American Phanerogams, IX. By N, L. Britton 
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PROF. N. L. BRITTON, 


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Pteridophyta and Spermatophyta of Northeastern North America 


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TORREY BOTANICAL CLUB. 


Antidromy of Plants.* 


By GEORGE MACLOSKIE, 


During the summer of 1893 I made observations on Indian 
corn, which were published in abstract in the Princeton College 
Bulletin of November, 1893. It was then shown that, if we judge 
from the modes of overlapping of the leaf margins of Maize and 
of other Gramineae, there must be two kinds of plants of every 
species of the order; the one kind or “caste” has its lowest foliage 
leaf (the leaf next above the pileolus in the embryo) with the 
right margin of its sheath overlapping the left margin, “ dextrally 
infolded” as I term it; and the other caste has the left margin 
Overlapping the right, “ sinistrally infolded.” The leaves at sub- 
sequent nodes are alternately sinistral and dextral. In order to 
aScertain the origin of this duplicity I read Van Tieghem’s re- 
_ Searches on the Cotyledons of the Gramineae (Ann. des Sci. 
re Naturelles, 1872), which stated that the leaves of the plumule 
within the seed are alternately enfolded on each other, but failed 
_ to indicate the direction of the enfolding of the first leaf The 
_ dissection of one or two seeds revealed a dextral initial folding; 
, end soon other grains were found with sinistral folding. Thus 


it became manifest that as there are two castes of maize-plant, so 
“* 3 —.. 
* Read before Section G., A. A. A. S., Springfield meeting, September, 1895. 
+ He states that the direction of the evolution of the leaves is determinate, but 
_ there is variation as to the first green leaf, and he had not been able to discover its 
Cause, 


380 


there are two castes of grains, the one being the “ anudrom” of 
the other, the leaf-folding starting diversely, and the leaves of suc- 
cessive nodes, running contrariwise in the plumule of one grain 
as compared with that of another grain. 

The next problem was to orientate the grains of each caste in 
the ear of Maize. The ear consists of columns each containing 4 
pair of rows of grains; we may designate the row opposite our 
right hand dextral and the other row (opposite our left) as sinis- 
tral. It was soon made out that in the particular ear examined, 
the grains of the dextral row were all with dextral embryos, and 
those of the sinistral row had sinistral embryos. Whether this 
law would apply to all the ears on one plant, or whether the order 
would be inverted between the ears arising from successive nodes, 
or between the ears of different plants, is yet to be determined. 
On examining the very young ear of maize I found the grains of 
the paired rows of each column orientated close to each other, 
almost face to face, the young styles running up together, and a 
gap between the adjoining two-rowed columns. 

From this discovery the inference was obvious that the seeds 
of corn differ from each other antidromically, according to the side 
of the placenta or axis from which they arise; that their embryos 
vary in consequenee, and determine the caste of the future plant. 
Whilst it was easy to see that the same rule will include all the 
Gramineae, I hazarded the suggestion that it may be found in 
some measure to dominate other orders of plants. The dis- 
covery as to the corn .disposes of Sachs’s crowning argument 
against phyllotaxy, which he supposed could have no significancy 
as to Gramineae, on account of their distichous leaves. 

Early in August, 1895, 1 was struck by the graceful inflores- 
cence of Ladies’ Tresses (Spiranthes praecox Watson, S. £ ramined 
var. Walteri Gray). Its spiral rows of pure white flowers are anti- 
dromic as between different individual plants; about half the 
specimens have dextral spirals (viz., turning in the direction of the 
thread of a common screw), and the same number have sinistral 
spirals. Another interesting point was that the phyllotaxy, or 
arrangement of the distichous leaves in a primitive spiral, of each 
of the plants of Sprranthes follows the order of the inflorescence, 
dextrorse phyllotaxy invariably accompanying the dextrorse an- 


381 


thotaxy, and conversely.* The numerous seeds of these plants 
are too small to be easily compared together ; but the phyllotaxy 
or even the inflorescence serves as our best guide when other 
indications fail. A similar remark will apply to Oenothera bt- 
eunis, whose inflorescence curves to the right or left in the same 
direction as the leaves; though not always so easily determined 
as in Ladies’ Tresses. In some cases the seeds form a ready 
guide. For example, the Lima-bean is found on examination to 
have its cotyledons right and left, and two foliage leaves respec- 
tively dorsal and ventral (towards the dorsal and ventrai sutures 
of the pod). Now I found that the right margin of the dorsal 
leaf of the plumule of one seed overlaps the right margin of the 
ventral leaf, but the left margin of the dorsal leaf is overlapped by 
the left margin of the ventral leaf; and in another seed these 
relations are reversed. I also found that the seeds growing on 
one valve of the pod, being Nos. 1, 3,5, were all similar; whilst 
the seeds on the other valve, being Nos. 2, 4, were reversed. 
Thus the seeds on one margin of a carpel are found to be antidrom- 
ically related to those on the opposite margin. (I have not ex- 
amined whether successive pods from the same plant reverse these 
characters or not.) The pea showed its antidromy best when 
germinating, the emerging plumules of different seeds appearing 
under careful orientation to come up with opposing twists. The 
large plumule of the Almond seed can be seen by dissection to 
have two modes of torsion in different seeds. The akenes of Caf 
Sea exhibit antidromy, an interesting point in this plant as the op- 
Posite leaves render the determination otherwise difficult. The 
deeply enfolded endosperm of the coffee-bean is seen on a cross-sec- 
tion of different seeds to be in opposite directions; a mark which 
is a sure indication of similar diversity in the minute embryo. 
Even from the outside the two kinds of akenes are easily distin- 
Suishable, the figure of one reversing that of the other like one’s’ 
tight and left hand, thus proving that the two mericarps are rela- 
tively antidromic; and confirming the evidence already derived 


from the corn and Lima-bean, as to the origin of this character. 


* sep vaeeaa mane eee ee 
oo + Dextral phyllotaxy has the primary spiral traced by the insertion of the leaves 
directed like the thread of a common screw. With leaves overlapping at the margins, 


: dextral overlapping may produce sinistral phyllotaxy. 


382 


The case of Banksia of Proteaceae further illustrates the subject. 
Judging from the figure in Engler and Prantl’s Pflanzenfamilien, 
(3: part I, 152) it has a pair of flowers, situated back to back, with 
bractlets, stamens, ovary and seeds, all antidromic; in this case 
the diversity of the seeds seems to be anticipated by the structure 
of the ower. The flowers on the same branch of Althaea have 
their petals twisted in contrary directions. 

In connection with the explanation of antidromy as depending 
on the origin of seeds along the margins of a bilateral organ, it 
would be interesting to examine the few cases in which seeds are 
represented as terminal on the floral axis, a view which may be 
confirmed or refuted by this law. It is likewise interesting to note 
that in specimens of Bryophyllum calycinum (kindly furnished me 
by Prof. S. T. Maynard, of Amherst Agricultural College), I was 
able in this opposite-leaved flower to make out the marginal buds 
on the leaves to be antidromous. The Calla lily (Richardia) has 
both the alabaster spathe and the arrangement of the akenes on 
the spadix antidromic as between plants which grow from the 
same rootstock. The Iris also when growing by bifurcation of the 
rootstalk gives antidromic plants; how they are when propa- 
gated by lateral branching I cannot say. Rushes growing to- 
gether in clumps are antidromic as between the individuals 
united at the base of the rootstalk (I do not find it so in Carex or 
grasses growing in tufts; all of the same tuft seem to me to be 
homodromic, but this subject requires more careful examination 
than I have made). 

Phyllotaxy is a particular outcome of antidromy, and in very 
many cases is the readiest evidence of the antidromic organiza- 
tion of the plant. But whilst antidromy is a primitive character, 
influencing the general morphology of the plant, yet each part - 
the evidence, and most frequently the order of the leaves, the in- 
florescence and the perianth, may be modified by secondary 
changes, which send us back to the mother-seed and its germina 
tion as the only remaining proof of the primitive character. Twin- 
ing of stems, contortion of perianth, accumulations of flowers 1? 
complex ramifications, leaves becoming opposite, or being spread 
out to the sunlight, and even difficulties of orientation of seeds 
disguise the truth and explain why it has so long remained 4 


383 


secret. The distortions of phyllotaxy misled the early students 
on the subject, so as to nearly banish the term and the theme 
from botanical science. Old books spoke vaguely of some plants 
being homodromous and others heterodromous, without attempting 
to find any law to explain these apparent irregularities. We now 
See its significancy and find its anomalies all reduced to order, and 
we must welcome it back to its rightful place. So far as I have 
been able to find, all plants are homodromic within the indi- 
vidual, and heterodromic as between different individuals of the 
Same species. Apparent exceptions to these rules are no more 
than apparent, and if anybody will set himself to look up the evi- 
dence, it will soon be so overwhelming, and nothing against it, as 
to render his work monotonous. Go into the nearest orchard 
and you will find two kinds of every species of fruit tree, two 
kinds of every shrub, two kinds of common flowers, having the pri- 
Mary spirals of the leaf-insertions dextral in one set and sinistral 
in the other set. In case of plants with opposite leaves this evi- 
dence will fail you, for you can make out two crossing primary 
spirals in the same branch; in other cases you are baffled by the 
leaves assuming new positions for the sake of the light, though 
Sometimes even in these plants you may find the primitive traits 
in branches not exposed to the sunlight. 

With a few weeks’ observation I have found double phyllotaxy, 
4S a mark of antidromy in the following plants representing the 
More important orders of Phaenogams: De/phinium (Ranuncula- 
Ceae), Liriodendron (Magnoliaceae), Bocconia (Papaveraceae), mus- 
tard (Cruciferae), Aéutilon and Hibiscus (Malvaceae), Pelargonium 
and Jimpatiens (Geraniaceae), bean and pea (Leguminosae), apple, 
Pear, peach (Rosaceae), Ocnothera (Onagraceae), carrot (Umbelli- 
ferae), sunflower and other Compositae, Lodelia (Lobelieae), Myosotis 
(Borraginaceae), Verbascum (Scrophulariaceae), tobacco (Solana- 
ceae), Polygonum (Polygonaceae), Ricinus (Euphorbiaceae), Salix 
(Saliaceae), Quercus (Cupuliferae),and among monocotyledons, lily 
(Liliaceae), Musa, Ladies’ Tresses (Orchidaceae), species of Aroi- 
deae, Iridaceae, Juncaceae, Cyperaceae, Gramineae. To the above 
we have to add from the perianth and arrangement of stamens 
Ny mphaeaceae ; and by the courtesy of Mr. Everett H. Barney, 


of Springfield, Mass., I was furnished from the Forest Park Ponds 


384 


with seeds of Nelumbium which on being opened, showed the 
embryos rich in chlorophyll, with their leaves in one seed folded 
round in the inverse order of those in another seed; besides this 
the insertion of the stamens in the flower of different individuals of 
Nelumbium form antidromic spirals. A special case among the 
Liliaceae is Convallaria majalis, whose two leaves are at the usual 
angle among the Monocotyledones of 120°, and as one of the 
leaves forms a sheath around the other, it is instructive to observe 
how in a bed of Lily-of-the-Valley, all the plants being regarded 
under the same orientation, the inner leaf in half the plants bends 
over 120° to the right, and in the other half to the left. Arthur 
K. Harrison of Lebanon Springs, N. Y., informs me that before he 
heard of my work he had taken note of the double phyllotaxy of 
Veratrum viride. ; 

With a little care we can make out the general law as applying 
to the Gymnosperms; both the phyllotaxy and the spirals of the 
cones of Coniferae show it; and it may be expected to manifest its 
presence in the relative position of the cotyledon, during germina- 
tion; also in the embryo and its suspensor in Cycas. (See figures 
in Engler & Prantl, 2: 1. 17.) 

I have not yet tried the opposite-leaved orders, further than 
the case of Caffea, and that of Bryophyllum, as insufficiently 
tested by the marginal buds on the leaves; if this last observa- 
tion be verified, it may reinforce the old doctrine that ovules are 
the homologues of such marginal buds. Acer platanoides shows 
it by the antidromic folding of the cotyledons of the seeds of ad- 
joining carpels; also Aesculus by the contrary curvature of its 
radicles and by its plumules. Nor have I tried the twiners 
and climbers, saving that I have partially succeeded with 
Morning-glory (Convolvulaceae). Its embryo resembles that of 
some Cruciferae in having the radicle folded “ incumbently’’ upo” 
the cotyledons ; and again (unlike the Cruciferae) its orientation 
is changed so that the radicle lies next the floral axis and the 
cotyledons lie towards the periphery. Now if we reduce the 
orientation back to that of Cruciferae, we shall find a very close 
correspondence of the embryonic structures in the two orders; 
and the same antidromic difference will be found in Convolvu-. 
laceae that is readily shown by the phyllotaxy of Cruciferae. 


385 


The embryo of Lepidium Virginicum has its cotyledons twisted 
So as to be erroneously described in the books as accumbent; 
something similar occurs in Sesymorium officinale. In these cases 
the adjoining seeds have the embryos twisted in contrary direc- 
tions, so that the apparent anomalies are explained as cases of 
Pronounced antidromy. Whilst the pericarp of akenes follows 
the torsion of the enclosed seed, capsular pericarps seem to 
follow the phyllotaxy of the mother plant; thus the pods of 
mesquit (Prosopis) are similarly twisted in the same plant, as are 
those of Medicago, those of balsam (Impatiens fulva and I. bal- 
samina) spring open with a right or left twist in harmony with the 
dextral or sinistral phyllotaxy of the bearing plant. 

Amongst the peculiar cases that occurred was that of Saéix 
Babylonica, of which only the female plants are known; and con- 
Sequently there is no reproduction by seed, and we expected to 
find no antidromy. Specimens growing about Princeton appear 
to be sinistral in their phyllotaxy, and as tradition derives 
them from the tree growing by Napoleon’s grave at St. Helena 
the inference arises that the St. Helena willow also was sinistral. 
This inference is confirmed by a young tree in Forest Park, at 
Springfield, which Mr. Barney knows to have come from St. He- 
lena. But other specimens of the same species with dextrat phyl- 
lotaxy are common. Thus we learn that whilst all representatives 
of the male line of this species have probably perished, there are 
at least two independent branches of the female line perpetuated 
by cuttings, 

Another interesting case is that of Canna. The leaves of all 
the Specimens which I have found at Northampton, Mass., and at 
Princeton, are when young spirally folded, right flap uppermost, 
and when expanded have a slightly dextral phyllotaxy. I am 
told that this is usually propagated from bulbs; but that it is 
sometimes grown from seeds; if this last statement be true we 
should expect individuals of both castes; but I have been unable 
to find any except the one caste (even a second variety bearing 
reddish leaves is by coincidence of the same cast as the common 
One). Doubtless the other caste occurs through the country. 

The scape or flower-stalk of Canva may cast light on the 
a Structure of Gramineae. Whilst all the foliage leaves of the Canna 


Re ne 


386 


(so far as I have seen it) are dextral, the flower-stalk is provided 
with a succession of sheathing bracts, the edges folding over each 
other in reversed order at every succeeding node. This is ex- 
actly as in Maize, being a special kind of phyllotaxy, which de- 
pends not on a primary spiral, but on a reciprocating overlapping 
of the margin of sheathing organs; also a double leaf between 
each branch and the mother-axis is present as described by Van 
Tieghem in Maize. Thus it appears that the whole corn-plant, 
culm, leaves and flowers is the counterpart of the flower-stalk of 
canna and of its bracts and flowers, but having no representative 
of its foliage-leaves. According to this view one part of the re- 
duction of the Gramineae is the non-development of proper foli- 
age leaves, and by way of compensation the excessive development 
of the bracts with a green lamina for assimilation. It is of special 
advantage to gregarious plants to have their assimilating organs 
lifted up to the air and sunshine. * (No notice is taken of the pecu- 
liar alternancy of leaves of gramineae by Pax, Vines or the other 
recent writers on Morphology). 

Among the results of this account may be mentioned the 
extension of unity of primitive structure thus shown to exist 
among all the Phaenogams; a unity that may yet be found to 
include some of the Cryptogams; also the relative simplicity of 
the Monocotyledones, which show few secondary distortions, 
though they are often reduced. A new problem of heredity 1s 
started, running differently through two sides of the carpel; yet 
each seed transmits both castes, one to appear forthwith in its im- 
mediate offspring, and the other to appear ultimately in a moiety 
of its successors. The objection that the discontinuity betwee? 
carpel and ovule negatives the possibility of such transmission of 
characters is of no weight; the discontinuity is only apparent, for 
characters of secondary acquirement are carried across the g4P» 
and @ fortiori we may expect such a primitive law of organization 
as antidromy to be inherited. This law is also useful to suggest 
discovery. It suggested to me at the outset diversities betwee" 
the stalks and inflorescence of Iris that had escaped Arnold Do- 
del in his study and illustration of /ris Sibirica, which was the work — 
of some years; it has also opened problems about the significancy 
of opposite leaves, the real direction of leaf-traces in stem-struc” 


387 


ture (hitherto assumed to be entirely symmetrical) and other sub- 
jects bearing on vegetable anatomy, and the difficulties which 
brought discredit on phyllotaxy all vanish. Perhaps it may fur- 
nish a solution of the problem why wooden poles split in anti- 
dromic spirals, for which phenomenon some people have sug- 
gested the stress of wind on the living tree. 

My work has been necessarily hurried; and I shall be glad 
if others will verify or amend it, and help to fill the many lacunae 
which I am compelled to leave unsupplied. ~ 


PRINCETON COLLEGE, Sept. 7, 1895. 


Description of a new problematical Plant from the Lower Creta- 
ceous of Arkansas. 


By F. H. KNow ton. 


PALEOHILLIA ARKANSANA gen. et. sp. nov. 


Stems hollow, . 5-.75 cm. in diameter, several centimetres long, 
roken; wall two or three layers of cells thick; cells of epider- 
mis of two kinds: 3-5 longitudinal rows of elongated, thin-walled 
Cells that are two or three times longer than wide, alternating with 
broad bands of shorter and more irregular cells; stomata numer- 
©us, confined to the broad bands of irregular cells, arranged in 
three rows, two next to the rows of elongated cells with a row of 
distant ones between; stomata with apparently 4-6, usually 5, 
guardian cells. 

The material upon which this description is based was collected 
by Prof. R. T, Hill, of the United States Geological Survey, dur- 
Ing the season of 1888, while engaged under the auspices of the 
Arkansas Geological Survey in making a general investigation of 
the geology of southwestern Arkansas. It came from a gulch on 
One of the smaller branches of the Muddy Fork of Little River, 
about six miles northeast of Center Point, Howard county. The 
deposits containing these fossils were referred by Prof. Hill to the 
Trinity Division of the Lower Cretaceous. The beds are described 
aS consisting of basal ferruginous sands, succeeded by firm white 
°r yellow sand often filled with small concretions of iron pyrites, 
and mixed with clay. This clay is in sufficient quantity to bind 
the sandy material together “so that in drying it often becomes 


388 


almost as hard as burnt brick.” It was in this clayey material 
that these plant-stems were found. 4 
The stems are very abundant and scattered in all directions 
through the mass, They appear to have been much rolled about 
and broken up, it being difficult to find a piece three centimetres 
in length. They were originally hollow, but most of ae are 
now pressed perfectly flat, although an occasional one is found 
that was filled with clay when probably in a fresh state and conse- 
quently retains nearly its cylindrical form. When liberated by 
the crumbling of the sandy clay, the stems are very dark brown, 
almost black, in color and perfectly opaque. By boiling them for 
a few moments in dilute acid, a large part of the coloring matter 
was discharged and the cellular structure could then be made out. 


Fig. 1.—Epidermis of Paleohillia Arkansana X< go. 
a, a, bands of long, narrow cells. a 
b, b, bands of short irregular cells, with three longitudinal © 
stomata, 4 
¢, c, stou ata with four and six guardian cells respectively. 
Fig. 2.—Stoma with four guardian cells X 300. 
Fig. 3.—Stoma with five guardian cells xX 300. 


389 


This curious plant, as already stated, was hollow, with the walls 
consisting of three or four, or possibly more, layers of cells. The 
epidermal cells are markedly separable into two kinds: first, narrow 
bands of three or four rows of thin-walled cells which are two to 
four times longer than broad, separated by broad areas or bands 
ten or twelve cells wide, of short irregularly quadrangular cells. 
In figure 1 a, a, represents the bands of narrow cells, and b, b, the 
short irregular cells. Under the microscope the two kinds of cells 
divide the surface longitudinally in very clearly mark parallel lines. 

The stomata are the most remarkable feature about this plant. 
As may be seen by the drawing, the guardian cells are quite irreg- 
ular in shape and appear to vary in number from four to six, the 
most frequent number being five.* The opening is in all cases 
large and in some instances appears even larger than either of its 
guardian cells. It is difficult to see how so large an orifice could 
be completely closed by them. 

The stomata, it will be observed, are confined to the broad 
bands of irregular cells, and are also arranged in longitudinal rows, 
those next to the bands of long cells being numerous, while in the 
central row they are scattered, there being only about one-third as 
Many as in the others. The orifice between the guardian cells is 
approximately circular. The guardian cells are themselves ar- 
Fanged in a more or less regularly circular manner, their shape de- 
Pending on the number present. When there are only four they 
are rectangular with rounded outer sides; when there are five or 
More they are keystone-shaped, or broadly wedge-shaped. 

Iam at the present time unable to suggest any satisfactory re- 
lationship, either living or fossil, for this anomalous plant. The 
first thought when seeing the numerous fragmentary stems 
throughout the clay mass, and finding that they are hollow, is that 
they belong to Aguisetum. But on examining large numbers of 
fragments, not a single joint with the characteristic teeth could be 
detected, and moreover the stomata are entirely different. It is of 


Course well known that the stomata of living Aguise¢um have two 
; atk as pidermal 


*In figure 1 the guardian cells appear to be in the same plane as the e 
“ig They are of course below them, but the lines of the overlying epidermal cells 
have not been drawn in, so that there may be less confusion in following their eee 

* Figures 2 and 3, much enlarged views of single stomata, make the real condi 


tion plain, 


390 


pairs of guardian cells, but they are not all in the same horizontal 
plane, one pair, the “subsidiary cells,” of Strasburger being below 
the other. The guardian cells are usually ornamented by silicified 
ridges radiating from the orifice, not a trace of which can be found 
in the fossil under discussion. 

Some of the Bryophytes, notably the liverworts, have stomata 
with more than two guardian cells, but they are loosely cellular 
plants quite unlike this hollow-stemmed plant. 

In the absence of more specific information as to its affinities, 
I have decided to describe this plant under a tentative name, Te 
cognizing the fact that this provisional name may be changed at 
any time provided the fossil can be more definitely placed. I 
had at first given it the name of //i//ia, but as there is a genus of 
this name in Rubiaceae, I have called it Paleohillia, It com- 


memorates the collector and is not intended to imply relationship 
with the Rubiaceous genus. 


Observations upon some Oklahoma Plants. 


By Epcar. W. OLIVE. 


| The botany of Oklahoma is exceedingly interesting, because 
this territory is a borderland region between the Gray's Manual 
and Western Texas Manual regions. Until about five years 48° 
the plants of this district were but little known to botanists, and 
the results of recent collections disclose a flora rich in interesting 
forms. Especially valuable is a “ List of Plants collected by . 
S. Sheldon and M. A. Carleton in the Indian Territory in 1891, 
published as a contribution from the National Herbarium in 189? 
The months of July and August, 1893, were spent in and about 
Payne county, in the very northeast of Oklahoma, about 9° miles 
south of the Kansas line through the Cherokee strip, and about 
150 miles west of Arkansas. This is in lat. 97° W. and is bul? 
few miles south of the parallel bounding on the north Tennessee 
and North Carolina, so that the collections were made just south 


391 


of the line of the extreme southwestern limit of Gray’s Man. 6th 
Ed. About 175 species of Phanerogams and Pteridophytes were 
collected, about 60 of them new to Messrs. Sheldon and Carleton’s 
list, most of these, however, the commoner plants, and 12 of 
which are not reported in Gray’s Manual. Of these 12, 3 are not 
included in Dr. Coulter’s Martual of the Texas Flora, nor 8 of 
them in his Botany of the Rocky Mountain region. 

These 12 plants are Zalinum calycinum Engelm., found abund- 
antly on the red sandstone rocks outcroping in ravines and along 
the Cimarron River; Desmanthus Jamesti T. & G., very abundant 
on dry prairies; Galactia mollis Michx., in sand along the river 
banks; Acacia Jiliculoides (Cav.) Trel., abundant in the sandy 
woods; Gaura villosa Torr., showing gradations into forms ; 
Sesuvium Portulacastrum L., in sand along the saline banks of 
the river; Crnoscitadium pinnatum DC., but one plant collected 
by a roadside; Aster patens Ait. var. gracilis Hook., the va- 
riety not in Gray’s Manual, very abundant in rich sandy ground 
Near the river; Baccharis glutinosa Pers., the fertile plant conspic- 
Yous by its very long and white pappus along the sandy river 
banks ; Lriogonum longifolium Nutt., on dry prairies; Aphanoste- 
phus ramosissimus DC., found in abundance in the sand of rich 
river bottoms; Cooperia Drummondii Herb., near Stillwater on rich 
Prairies. 

The flora of Oklahoma is very similar to that of southern 
Kansas. The climatic and geologic conditions are very similar in 
both regions, and this fact, combined with the fact of the prox- 
imity of the Cimarron and Arkansas rivers flowing southward 
_ through the territory from Kansas, tends to make the floras alike. 
Outcropping “red-beds” (whence probably the name Oklahoma— 
“home of the red earth”) occur in both regions associated gener- 
ally with a very sandy soil. Rich, sandy land, well timbered, oc- 
curs along the rivers and creeks, while back some distance on the 
uplands, prairies supplant the forests. The farther west one goes, 
the observer can readily notice the dwarfing of the trees as the 
Tegions of less rainfall are passed through, and many plants show 
Stadation into dwarfed forms. Most of the plants of this whole 
: district, moreover, present the characteristics of the plants of dry 
Sterile regions, viz., thickened leaves and epidermis, sunken sto- 


392 


mata, absence or narrowness of leaves, or an unusual amount of 
wooliness or hairiness. 

There are quite a number of common oaks in some portions 
of this eastern part, Spanish oak, Post oak, but most abundant in 
the upland reduced forests is Quercus nigra L., the dwarfed, 
gnarled “ black jack.” There are some hickories, black walnuts, 
cottonwoods, and elms along the river and creeks, the elms bearing 
abundantly large bunches of mistletoe. 

A few observations as to the occurrence and habitat of some 
plants may be interesting. In the rich sandy land along the river 
bottoms the commonest shrubs are Cephalanthus occidentalis L., 
Stillingia sylvatica L., Rhus copallina L. On Stillingia was found 
an Aecidium which has not been yet reported on this host, as far 
as can be determined. Somewhat abundant in similar places were 
Argemone platyceras Link and Otto, Callirhoe involucrata (Nutt.) A. 
Gray, Parosela enneandra (Nutt.) Britton, /roelichia Floridana 
(Nutt.) Moq., /udigofera leptosepala Nutt., Aphanostephus ramosisst- 
mus DC. In wet, salty sand near the river were Pluchea campho- 
vata (L.) DC., Sesuvtum Portulacastrum L., and in the dry sand, 
Cycloloma atriplicifolium (Spreng.) Coulter, Baccharis glutinosa 
Pers., and Parosela /anata (Spreng.) Britton. The latter is reported 
in Gray’s Man. to have “3-4 pairs” of leaflets, while 6-7 pairs were 
usually present on the specimens collected. On the high bluffs of 
the river Yucca glauca Nutt. was not infrequent. 

In the woods which extend back from the river bottoms tw° 
or three miles are Cassia Chamaecrista \.., C. nictitans L., or “ sen- 
sitive plant,” Desmanthus Jamesii T. & G., Clitoria Mariana L, 
Gaura villosa Torr., Onagra biennis (L.), Scop., grandiflora Lindl., 
a beautiful passion-flower, Passiflora incarnata L., Lacinaria squat 
rosa (L.) Hill, Chrysopsis villosa Nutt., in many of its variable 
forms, Asclepias verticillata L., and Acerates angustifolia (Nutt.) 
_ Dec.; also low shrubs of Busmelia lanuginosa (Mx.) Pers. and sev" 
eral species of plum. A perhaps noteworthy point was the occur- 
rence of Ludwigia alternifolia L. in sandy but perfectly a t4- 
vines. Gray’s Manual reports the habitat of this as “ swamps.” 

The whole prairie region is characterized by an abundance of 
plants belonging to the orders Leguminosae and Compositae. Par- 
ticularly abundant on the prairies are Kuhnistera multiflora (N utt.) 


393 


Heller, K. purpurea (Vent.) MacM., Amorpha canescens Pursh, 
Parosela aurea (Nutt.) Britt., Solidago Missouriensis Nutt., Helian- 
thus mollis Lam., Hieracium longipillum Torr. The fact is signifi- 
cant that of the 175 species collected, 33 were Leguminosae and 
32 were Compositae. Sadbdbatia angularis (1..) Pursh, S. campes- 
tris Nutt. and Buechnera Americana L. give bright colors to the 
Prairies during June and July; Linum sulcatum Riddell, Ceanothus 
Americanus L.., Jatropha stimulosa Michx., Euphorbia corollata L., E. 
petaloidea Eng., E. marginata Pursh, occur on the richer prairies, 
while Megapterium Missouriensis (Sims) Spach, Houstonta anguste- 
Solia Mx., Stenosiphon linifolium (Nutt.) Britton, Opuntia poly- 
acantha Haw., and Gerardia densiora Benth., are found on dry 
sterile prairies. A very severe case of poisoning was incurred from 
collecting Euphorbia corollata. 

A very paradise for a collector of aquatic vegetation is a large 
Pond near Perkins, Oklahoma. Several Sagittarias, Nelwmdo lutea 
Pers,, Potamogeton lonchites Tuckerm., the latter growing “rarely 
in ponds” (Gray’s Man.), P. diverstfolius Raf., are most abundant 
throughout, while near the edges Heteranthera limosa (Sw.) Willd., 
Ludwigia glandulosa Walt., Monniera rotundifolia Mx., and Mar- 
silia vestita Hook. & Grev. grow rank. Some of the specimens of 
this Marsilia growing both in and along the banks of the pond 
have petioles 7-8’ long, the type being reported in the Manuals 
1-4’. Dr. Underwood, however, pronounces this undoubtedly 
MM. vestita, 

As suggested above, the special interest of this region lies in 
the fact of the meeting of two floras and the sometimes abrupt, 

-S€nerally gradual, transition of one into the other. The flora can- 
not be studied comprehensively except by an extended period of 
field work and by carefully noting all the environmental conditions. 
The farther west one goes into the territory the more sandy and 
desert the regions become; and such are the variations from some of 
the more eastern forms that many are classed as varieties, no 
doubt the result of a change in habitat. According to Mr. 
Coville’s suggestions in his “ Botany of Death Valley Expedition,” 
the shrubs and trees and on the prairies the perennials should 
age be noted to determine the characteristic plants of the 
ora, 


394 


Grateful acknowledgments are due to Dr. John M. Coulter and 
Professor E. B. Uline for kindly determining some of the species 
and checking most of the list of collections. 


WABASH COLLEGE, CRAWFORDSVILLE, IND. 


On the two Editions of Emory’s Report, 1848, 


For some time past I have been aware of the fact that there 
were two editions of Lieut. W. H. Emory’s “ Notes of a Military 
Reconnoissance,” both bearing the same date (1848); but I sup- 
posed they were identical, except that one had appended to it the 
reports of Abert and Cooke and the journal of Johnston. But I 
now possess copies of both of these books, and find that they dif- 
fer in various details, which are of importance on account of Prof. 
Torrey’s reports on the botany of the expedition. 

Both editions are Executive Documents of the 30th Congress, 
Ist session, and they are numbered 7 and 41, respectively. No. 7 
was issued for the Senate. It contains nothing but Emory’s re- 
port, with its seven appendices, and consists of 416 pages. Each 
age has[* ]in the upper external corner. No. 41 was issued 
for the House. It contains the reports of Emory, Abert and 
Cooke, and Johnston’s journal, and consists of 614 pages. “ Ex. 
Joc, No. 41," appears in the middle of the top of each page, with 
the exception of pp. 145-158, in the botanical portion, which have 
[7] in the upper external corner. This is evidently a tyP? 
graphical error, and not a mistake in binding, for the pages af 
not the same as pp. 145-158 in No. 7. 

In both of these volumes Professor Torrey’s report on Emory’s 
plants occupies pp. 135-159, constituting Appendix No. 2. My 
copy of No. 41 bears the following manuscript note at the top of 
page 135. “ This appendix is full of gross typographical errors— 
the printer having refused to send me proof sheets according to 
agreement. J. Torrey.” The same remark would apply with 
equal force to No. 7. Both books show evidence that they were 
published either carelessly or in very great haste. The principal 


differences between the two documents in the botanical portions 
are as follows: 


395 


I. In No. 7 Prof. Torrey’s general report occupies pp. 13-5 
155; page 156 contains an “ Explanation of the Plates; and on 
PP. 157-159 is a letter from Dr. Geo. Engelmann describing (from 
drawings only) the Cactaceae of the Expedition, in fine print. In 
No. 41 Prof. Torrey’s general report occupies pp. 135-155 (but 
the matter on the various pages is differently distributed); Dr. 
Engelmann’s letter extends from page 155 to page 159; and chere 
#s no “« Explanation of the Plates.” 

2. In No. 41 three species are entirely omitted which are 
given in No. 7, viz.: (a) Zinnia grandiflora Nutt., (6) Convolvulus 
Nuttallii,and (c) Alternanthera? (Endotheca)lanuginosa. Of these (a) 
and (c) are described, and (4) and (c) are published as new names. 
A person having reference to these in “ Emory’s Mil. Rec.” and 
looking them up in No. 41, would be puzzled at their absence, 
and conclude that the references were incorrect. 

3. The fourteen plates in No. 7 were lithographed by E. Weber 
& Co., Baltimore. Those in No. 4! are similar, but were litho- 
8taphed by C. B. Graham, Washington. The latter are somewhat 
better (or less bad), from an artistic standpoint, but there seem to 
be no differences of importance—save that in No. 41 plate VI. is 
labeled Baileya multiflora instead of B. multiradiata. 

4. In both documents Appendix No, 6, occupying pp. 386-— 
414, consists of Lieut. Abert’s “ report of such objects of natural 
history as came under my observation while I was attached to the 
topographical party * * * during the journey from Fort Leaven- 
Worth to Bent’s Fort. The plants which were collected were sub- 
mitted to the inspection of Dr. Torrey, to whom I am indebted 
for their names.” Prof. Torrey’s list, to be found on pp. 406-414, 
differs somewhat in the two books, but this is of little importance, 
for it is a mere catalogue of names, without descriptions. 

I hope that this note, published in the BULLETIN, may help to 
Prevent trouble and confusion in the future. 

Joun HEenDLEY BARNHART. 


Tarrytown, N: Y., April 20, 1895. 


396 


Teratological Notes. 


By Francis E, Lioyb. 
PLATE 247. 


The many germinating acorns which were to be seen during 
the past winter in western Oregon afforded a good opportunity 


teratological interest have been noticed. 

One of these is an acorn which, after having pushed its radi- 
cle out some distance, had met with a misfortune. Presumably a 
snail or other depredator, judging the young plant a tender mor- 
sel, had bitten into it just at the juncture of one of the petioles 
and the hypocotyl (Fig. I). The mouthful included the little plu- 
mule, so that further development of any or all of the remaining 
parts would have been in vain. The hypocotyl, nevertheless, con- 
tinued to grow, the wound healing. Furthermore, the detached 
petiole healed and produced an incipient rootlet of its own. 

In order to determine experimentally the power of the detached 
petiole to produce rootlets, three partly germinated acorns were 
chosen and a petiole and the hypocotyl in each were separated by 
cutting. Ina few days the scars healed over, and in two of the 
acorns rootlets were formed, two in one (Fig. Ila) and four in the 
second. The third failed to grow. 

Another acorn was found, from the ruptured apex of which 
three lengthened petioles had followed the protruding radicle. 
Two of these were more slender than the third, and were closely 
approximated at their bases. These the plumule had spread apart, 
pushing its way up between them. The relative positions of the 
three petioles can be understood from the diagram (Fig 1Va) of 
a transverse section through the base of the plumule. It is evi- 
dent that the two cotyledons with the approximated petioles are 
the equivalent of one in the normal seed. The earlier leaves ar© 
reduced to mere subulate bracts, and in their number and arrange- 
ment vary greatly. In these I observed no difference, except that 
they are more crowded in one region of the young stem, and this 
probably had no relation to the number of cotyledons. 


397 


A number of acorns have been found with two fertilized and 
developed ovules. The presence of the supernumerary seed is 
betrayed by the unsymmetrical shape of the acorn. The rightful 
occupant—if might makes right—is usually well developed, and 
pushes out its radicle earlier than does the intruder, which is cor- 
Tespondingly smaller and flattened and twisted out of shape. 
Occasionally, when the supernumerary seed is large, if its position 
is favorable it gets its radicle out of the ruptured apex first. At 
all events it makes a brave effort to reach soil and sunlight. A 
few acorns have been found in which the two plantlets had de- 
veloped into two well-formed seedlings. Acorns containing more 
than one seed have all been found under young trees. In no case 
have I found such under aged trees. 


BIOLOGICAL LABORATORY, PACIFIC UNIVERSITY. 


Explanation of Plate 247. 


Fic. 1.—«¢,” cotyledon ; « f,” foliar appendages; “p,” petiole; “ pl,” plumule ; 
“h,” hypocotyl. The dotted lines indicate the portion excised. 

Fic. 2.—The seédling after the wound had healed and the second root-tip had 
formed. R—original radicle; R,—the rootlet of the cut petiole. 

Fic. 2a.—A seed, the petiole of which was cut. After healing the separated 
Petiole (p) produced two rootlets. The plumule continued to develop as seen. 

Fic. 3.—Shows the end view of the seedling with three cotyledons, the petioles 
of which are shown in situ. The plumule (pl) projects through the smaller, paired 
Petioles p, and p.. 

Fic. 4.—The same with the husk removed, and the three cotyledons outspread. 
The cotyledons c, and c, belong to the petioles p, and p, (in Fig. 3) respectively. 

Fic. 4a.—Transverse section through the base of the plumule of Fig. 4. 

_ Fic. 5.—An acorn with the husk removed showing the supernumerary seed in 
Situ. Its hypocotyl (h) had grown, the radicle partially decayed and thrown off two 
new rootlets (a and b). 


Contribution to the Characeous Plants of Maine—t. 


By Fy Lu. FEARVEY. 


So far as we know but little attention has been given to the 
Plants of this order by collectors in Maine; the species named 
in Redfield & Rand’s Catalogue of the Plants of Mt. Desert being 
the Only record we know of regarding them. We have recently 
been considering the few species found in the Blake Herbarium of 


398 


the Maine State College, and, adding our own observations, bring 
the record to date. 

Research will no doubt bring to light many more species. We 
are under obligation to Dr. Allen for professional courtesies. We 
will be pleased to learn of other species occurring in the State, 
or to know of Maine collectors of these interesting plants. 


ORDER CHARACEAE. 


Famity I. NITELLAE. 

1. Nitella opaca Ag. 

Blake Herb. Aroostook River (Blake), Sept., 1878. 

Redfield & Rand’s Cat. Plants of Mt. Desert, p. 227. 

Common in small streams about Orono (Harvey). 
2. NV. flexilis Ag. 

Western Maine (Kate Furbish.) 

Rand & Redfield’s Cat. Mt. Desert, p. 227. 

Miss Furbish’s plants show the monoecious fruiting and the 
abruptly acuminate leaves. The plants from Mt. Desert are 
doubtfully placed here by the editors of that catalogue. 


Famity II. CHARACEAE, 


3. Chara coronata A. Br. 
Aroostook R., Me. (Blake) Sept. 1878. 
4. C. contraria A. Br. 
Caribou, Me., Sept., 1878. 
The stream not mentioned in Blake’s notes. 
5. C. fragilis Desv. 
Very common in various streams in Penobscot Co. (Harvey-) 
In Sunkhoze Stream, occurring on gravelly and rocky bottoms 
for a distance of ten miles, the patches sometimes covering the 
bottom of the stream for rods. 


399 


Teratological Notes, 


1. On the Stipules of Lathyrus poly- 
phyllus. Dr. Masters* only mentions 
additional stipules in the case of Sa- 
4x fragilis. On a specimen of La- 
thyrus polyphyllus collected in California 
by Kellogg and Harford in 1868-1869, 
the lowest pair of leaflets is replaced by 
two stipules identical with those that 
subtend the rachis. The leaflets are 
normally alternate, but the additional 
stipules are opposite, as are the lower 
ones. About one-half of the leaves of 
the specimen are thus modified. 


== ) 


2. Polyphylly of the andrcecium. 
__3. Polyphylly of the gyncecium. 


*Veg, Teratol, 387. 


- 
QZ 

eer 
eZ 


2. On Polyphylly of 
the Flower of Dirca 
palustris. A specimen 
y of this species from 

one of the European 
botanic gardens shows 
this abnormal state in 
three ways. 

1. Polyphylly of the 


calyx. 


400 


Many instances of the multiplication of parts in the different 
whorls of a flowerhave been recorded. This takes place in 
different ways, and one or more whorls may be affected. In 
the present case there seems to be a union of two flowers, mak- 
ing. a duplication of each whorl, or polyphylly of the calyx, 
the andrcecium and the gyncecium, The undulation-like teeth of 
the margin of the calyx are increased from four to eight, the 
stamens from eight to sixteen and the pistil from one to two. 
The change dccurs in all the flowers on the specimen and 
modifies the shape of the calyx from “tubular-funnelform” to 
campanulate. 

Joun K. SMALL. 


Reviews. 


Systematische Phylogenie der Protisten und Pflanzen. Exnst 
Haeckel. Ist Portion. Berlin. 1894. 

As the author states in his preface, he has been engaged for 
the past thirty years upon questions pertaining to the subject- 
matter of this volume. All are acquainted with the “ History of 
Creation” and have enjoyed its generalizations, and the present vol- 
ume offers a rare treat to those who would advance a step and en- 
ter into generalizations based upon more specific and detailed 
data. 

He opens with a chapter upon Phylogenie, defining it, and 
giving Paleontology, Ontogony and Morphology as the sources 
for the working-out of the problems. He rapidly and skillfully 
sums up the positive and negative in the geological record; dis- 
cusses the value of Ontogeny and outlines the principles of 
Morphology. <A brief review of the methods is followed by # 
summary of the geological systems and the first chapter, including 
thirty-two pages, closes with a critical review of the monophyletic 
and polyphyletic theories of Phylogeny. 

The second chapter, fifty pages, deals with the phylogeny of 
the Protista, where the beginnings of life are taken up. 
Haeckel’s familiar Monera, “Structureless organisms without OF 
gans,” again does service as a starting point from which the pil 
tista arise, in which class he groups all those organisms “ which 


401 


do not build tissues;” calling those of a solitary turn of mind 
“ Monobionten” and the more social “Coenobionten.” F ollowing 
the definition of the Protista, a paragraph or two is devoted to the 
differences in the plant and animal kingdoms. Here much stress is 
laid upon the synthetic character of the one and the analytic char- 
acter of the other class of organisms. The plants are rechristened 
“Plasmodomen”’ and the animals “ Plasmophagen” or plasma 
builders and plasma destroyers. 

The Protista are then divided into three main groups, the 
atypical Protista; the plant type protista or Protophyta and the 
animal type protista or the Protozoa. In the first of these are in- 
cluded those organisms that show no particular affinity to either — 
Stroup, on account of an indifferent growth or because they may 
be at times either animal or vegetable in their functions, and are 
to be divided purely upon artificial grounds according to what 
seems to the author to be the predominant physiological activity. 
We note that the botanists gain the Peridineae and the zoologists 
the Bacteria, which latter are said to be lacking a nucleus which, 
_ if we are not sadly mistaken, is somewhat “ behind the times.” 

The typical Protophyta include those organisms in which 

I. The cell-wall is surrounded by a special membrane which 
is entirely closed or has only a few openings; this membrane con- 
Sisting of cellulose. 

2. The cell-body is either entirely motionless throughout its 
life history, or, at rare intervals and in small degrees, moves about 
by means of flagellae, as in the swarm spores. 

3. The cell is always colored, generally green or gold, and 
Contains chromoplasts which generally contain chlorophyll, also 
at times diatomin, haemochromatin, etc. 

4. The metabolism is entirely vegetable ; the cell is “ plasmo- 
domen” and assimilates CO,; it takes up no formed organic matter 
and possesses no mouth opening. 

In this group are included (1) Algarien, (2) Algetten. 

The pages following are too solid for abstracting, but are of 
interest with much philosophic thought; the phylogeny of the soul 
of the Protista forming an interesting extravaganza on pages 75, 
76. The third chapter deals with the systematic phylogeny of the 
Protophyta; in this the starting point is with those atypical Pro- 


402 


tista that have a “vegetable tendency.” Those without a cell 
nucleus, including many of the Cyanophyceae, under the name of 
Chromaceae begin the family tree. We can give here only the 
main lines of ascent, as the work is an abstract itself, and to further 
abstract it would out-Herod Herod. Through the Chromaceae the 
Algariae are reached in Palmella, from which Diatoms and Des- 
mids diverge and represent radiating extremes of development in 
their respective lines. The Algetten, the second main class, in- 
cludes many forms hitherto reckoned under the care of the zodlo- ° 
gists; many of the flagellate infusoria for instance. It includes all 
those single cell forms that have zoospores and flagellae move- 
ments: Luglena, Protococcus, Peridinium, Volvox, Botrydium and 
Caulerpa are included in order. 

The fourth chapter treats of the Protozoa, already reviewed by 
the zodlogists in “Science.” 

The fifth chapter takes up the general morphology of the 
Metaphytes or those plants which are many-celled and tissue 
builders, including (a) Thallophyta, (b) Diaphyta, (c) Antho- 
phyta. This chapter we consider the finest in the book and can 
be read by all botanists to some advantage independent of their 
interests, morphological, systematic or physiological. Chapters 
six, seven and eight take up the three classes respectively. 
The Thallophytes ascending through the Chlorophyceae, Proto- 
coccus to Confervales and branching in several directions; the 
Florideae, the highest branch, through Bangia ; Fucus, through 
Ectocarpus and Laminaria. The Characeae a special branch 
through Nitella, The Ulvaceae to the form similar to Riccia and 
from them to chapter seven on the Diaphytes. Among the Thal- 
lophytes the fungi are represented as on offshoot from the Chloro- 
phyceae, which is hardly tenable after Professor Farlow’s well- 
founded suggestions upon this group, and the lichens are regarded 
_ in the Schwendenerian sense. 

The Diaphytes include the Hepatics and Mosses. These are 
made to arise through a primitive Riccia, from thence through 
Pellia to the foliose forms and to Haplomitrium when a true axis 
is first reached and forni this, through Sphagnums and Andreaceaé 
or their common ancestor to the true mosses. Marchantiales and 
Anthoceratales are made blind alleys of growth. 


403 


The Pteridophyta are made to rise from the same ancestral 
Riccia-like form as the “ Diaphytes,” to a hypothetical “ Archi- 
pterides,” thence through the Hymenophyllaceae, when the stem di- 
vides into three lines, the Filices, the Equisetaceae and Lycopodi- 
aceae, which last make the way for the Anthophyta. 

These, in the closing chapter, through the Cycads and some 
forms similar to the fossil Noeggerathia split into a forest of stems. 
The Gymnosperms representing a growth through the Arau- 
carieae, while through some forms resembling the Australian 
“ Casaurineae” the Angiosperms were developed. 

The critic would need to be a specialist in many branches, and 
could write a book equalling, in size, the original, if all of the in- 

. teresting points touched upon were to be discussed, but to give an 
opinion of the work in a word, we can heartily say that a good 
and mighty work has been accomplished. 

Upon points of classification varying points of view must 
always be borne in mind. Possibly the author has made a mis- 
take in a fundamental point, upon which he devotes much space 
and thought to elucidate for having cautioned others not to con- 
found Homology, form: with Analogy, function: he deliberately 
makes his most sweeping classes and distinctions upon a purely 
Physiological, hence functional basis. 

The numerous points of difference which would interest 
Workers in special fields can not be discussed here, but the work 


is to be recommended to all readers as one of more than common 
interest. Situ Evy JELLIFFE. 


Volume I., of Anton Kerner von Marilaun’s Pfhlanzenleben, 
translated by Prof. F. W. Oliver as Ze Natural History of Plants, 
has recently been issued from the press of Henry Holt & Co. 
This work will be warmly received by all lovers of nature. In 
a Style simple, attractive and still thoroughly scientific, Professor 
von Marilaun considers in this first volume: The Living Principle 
in Plants, Absorption of Nutriment, Conduction of Food, Forma- 
tion of Organic Matter from the Absorbed Inorganic Food, Meta- 
bolism and Transport of Materials, Growth and Construction of 
Plants and Plant-forms as Completed Structures. 

Under there captions are discussed in a most charming man- 


404 


ner hundreds of interesting biological questions that have been 
practically excluded from the attention of very many people 
through the technicality of authors. Now, indeed, for the first 
time, are thrown open wide the gates to the natural history of 
plants and all may enter and enjoy the wide range of the whole 
field. The above mentioned seemingly unattractive subjects are 
invested with a charm that must arouse the enthusiasm of all. 
Thus under the rather unattractive title of Absorption of Nutri- 
ment, is discussed, among many other subjects, the adjustment of 
leaves to the welfare of the root; plants with traps and pitfalls to 
ensnare animals. So also under conduction of food a chapter is 
given to all those wonderful adaptations whereby the plant may 
suffer no inconvenience from untoward external conditions that » 
would interfere in any way with its life current; and then again 
are taken up those adjustments of branches and that exquisite 
moulding of leaf forms and their relation to the branches where- 
by the best interests of the plant are subserved, and in this con- 
nection follows a discussion of the protection of leaves against 
the attacks of animals. The work is in a sense exhaustive, 
copiously illustrated, the wood engravings especially being ¢*~ 
cellent, and the translator reflects most happily the spirit of the 
author. Cc: C. Curt. 


The London Catalogue of British Plants. The recent publica- 
_ tion of the ninth edition of the list of higher plants growing natu- 
rally in Great Britain and Ireland (the Characeae are appended), 
affords opportunity for ascertaining just how widely British bota- 
nists differ from our recently issued “List of Pteridophytes and 
Spermatophytes” in the nomenclature of genera. From the 
statements of some writers one might be lead to infer that the dis- 
agreements would be very numerous, but it appears that this 1S 
not the case. 

The total number of genera listed in the English publication, 
exclusive of the Characeae, is 538. The «List of Pteridophyt# 
and Spermatophyta growing without cultivation in eastern North 
America” contains 970. Allowing for differences in generic limi- 
tations, the London Catalogue sometimes uniting genera which 
the “Check-list” holds distinct, and vice versa, there are about 


405 


440 genera in common, all but 18 of which bear the same names. 


I have indicated these in the following table: 


CHECK-LIST. 


Capnoides Adans. 1 703. 
Roripa Scop. 1760, 
Alsine L. 175 3, 

lissa Adans. 1 763. 
Spiesia Neck. 1 790. 
Sanguisorba L.. 1 75%: 
Stlybum Gaertn. 1 788. 
Legousia Durand, 1782. 
Limonium Adans. 1 763. 
Statice L.. 1753: 

Udora Nutt. 1818. 


Leptorchis Thouars, 1808. 


Gyrostachys Pers. 1807. 
Peramium Salish. 1812. 
Juncoides Adans. 1763. . 


LONDON CATALOGUE. 


Neckeria Scop. 1777 (1). 
Nasturtium L. 1735 (2). 
Stellaria L. (3). 

Buda Adans. 1763 (4). 
Oxytropis DC. 1802 (5). 
Fotertum L. (6). 

Mariana Hill, 1762 (7). 
Specularia Heist. 1748 (8). 
Statice L. (9). 

Armeria L. (10). 

Elodea Michx. 1803 (11). 
Liparis L. C. Richard, 1818 (12). 
Spiranthes L.C. Richard, 1818(13). 
Goodyera R. Br. 1813 (14). 
Luzula DC. 1805 (15). 


Hierochloe 8.G.Gmelin, 1747. (16). 
Glycerta R. Br. 1810 (17). 
Polystichum Roth, 1794 (18). 


Savastana Schrank, 1789. 
Panicularia Fabric. 1763. 
Dryopteris Adans. 1 763. 


Notes. 


1. Both lists thus reject Corydalis Vent. 1803. While the British botanists were 
taking up another name, it seems strange that they could not accept Capmotdes, which 
18 quite as well defined as any of the rest of Adanson’s genera, and of these they have 

: admitted a considerable number : A/eum, Arctostaphylos, Cicendia, Epipactis, Poly- 
Senatum, Mibora, Apera, for example. 

2. The Linnaen use of Vasturtium is confined to the first edition of his « Sys- 
tema,” published in 1735. He did not use it subsequently and it was only again taken 
up by R. Brown in 1812, Meanwhile Roripa had been published and well defined. 
The American rejection of Vasturtium is thus based on the Rochester agreement, 
adopted by the Genoa congress, and more recently approved by the Austro-German 

tanists, to begin generic nomenclature at 1753. 

3- Here, again the “starting-point” for generic names comes into consideration, 
and 4/sin. has precedence of place in the first edition of the “Species Plantarum,” both 
genera being recognized in that work. It appears possible to me that both. genera 
will again be recognized, Alsine for A. media and its relatives and Ste//aria for 


Such types as S. graminea. : 
4. Both lists thus reject Spergularia Presl, and Lepigonum Wahl. Tissa has 


406 


precedence of place in Adanson’s “ Familles des Plantes” and is accepted by Baillon 
in his “ Histoire” and by Pax in Engler-Prantl’s « Natiirliche Pflanzenfamilien.” 

5. Necker’s name for this genus is satisfactorily identified by the description; I 
have been unable to discover any reason under any set of principles which really 
warrants its rejection. The authors of the London Catalogue have admitted oretta 
proposed by him, Why not Sfzesia ? 

6. Poterium L. is considered, and in my opinion correctly, by European authors, 
as monotypic, including only P. sfinosum L. of southern Europe. : 

7. This is a case in which the London Catalogue is right, and the American list 
wrong, under all rules, and we should be grateful for the correction. The citations 
are MARIANA Hill, Veg. Syst. 4: 19. 1762. Mariana Marianum (L.) Hill, Hort. 
Kew. 61. 1769. Hill’s use of duplicate binomials is the earliest which has come to 
my attention; he proposed a number of others in the same publication. 

8. Speculavia Heist. was taken up by no subsequent author until employed by AL 
phonse DeCandolle in 1830. The 1753 “starting point” excludes it. Meanwhile 
Legouzia had been published and the genus defined, : 

9-10. In adopting Statice L, for S. Armeria L, and its allies, and Limo- 
nium Adans. for Statice Limonium and its congeners, the American list returns 
to the use of Statice in the “Species Plantarum” of 1753, in which the genus Ar- 
meria was not taken up. The use of these generic names, as adopted in the London 
Catalogue, dates only from their publication by Willdenow, 1809. 

11. Elodea was regarded by the American committee as a homonym of Elodes 
Adans, 1763. I have recently discussed this point (Science, n. S., 2: 5- 1895), 
showing that PAz/otria Raf., has a few months’ priority over Udora. It will be 
noted that both lists reject Anvacharis. 

12, The validity of publication of Du Petit Thouars’ genera of Orchidaceae has 
been questioned, but also ably defended. I have not had access to his paper. 

13. There can be no doubt as to Persoon’s intention in proposing the name 
Gyrostachys ( Gyrostachis in original), as he refers to Ophrys spiralis L., to illus- 
trate it; this species is included in Spiranthes by recent authors. 4 

14. There can be no possibility of mistaking Salisbury’s meaning in the publica 
tion of Feramium, for he bases it on one of the same species cited by R. Brown under 
Goodyera a year later. It is true that Salisbury published no description of the 
the genus; but many other widely accepted genera rest on just this form of pub- 
lication, 

15. My remarks under Capmoides apply equally well to this. Perhaps the com 
pilers of the London Catalogue objected to the termination odes, but they have good 
authority for its use in the publication by Benthan and Hooker (Genera, 2: 301) of 
Mniodes A. Gray, and this is also accepted in the Kew Index. 

16, Gmelin’s name antedates the first edition of « Species Plantarum” and was 
not again used until after the publication of Sevastana. 

_ 17. I have not seen the original publication of Fabricius, but Famicularta and 
Glyceria are cited as equivalent by Kuntze. 

18, Both lists thus reject Aspidium Sw. 1800, It should be added that the Lon 

don Catalogue recognizes Polystichum Roth, and Lastraca Presi, as genera. 


The percentage of difference in the two lists is 4.09 per cent. 


407 


The changes of generic names in the ninth edition of the Lon- 
don Catalogue, from those used in the eighth edition, published 
in 1886, and which agree with the American catalogue, are as 
follows: 

Nymphaea 1.., 1753, for Nuphar Sibth. & Smith, 1808. 

Castalia Salisb., 1805, for Nymphaea J. E. Smith, 1806. 

Bursa Weber, 1780, for Capsella Medic., 1792. 

Coronopus Gaertn., 1791, for Senebiera DC., 1799. 

Schollera Roth, 1788, for Oxycoccus Pers., 1805. 

Pueumania Hill, 1762, for Mertensia Roth, 1797.* 

Homatocenchrus Mieg., 1768, for Leersia Sw., 1788. 

Weingaertneria Bernh., 1800, for Corynephorus Beauv., 1812. 

Steglingia Bernh., 1800, for Ziiodia Beauv., 1812. 

It would appear, from a consideration of these comparisons, that 
the British botanists were, after all that has been said, gradually 
coming to recognize the validity of the principle of priority, in at 
least generic nomenclature. There also appears to be no good 
feason why this process should not continue, so that by the time 
of the issue of the tenth edition of their useful list the percentage 
of difference might be reduced to 2 per cent. or perhaps less. 

For, if the recognition of this principle had been complete, and 
the 1753 « starting-point” had been accepted, not more than six 
of the genera would have borne different names. Noi, S : 

Among Rhode Island Wild Flowers. By W. Whitman Bailey. 
Providence, R. I. Preston and Rounds. 1895. Cloth. 16 mo. 
Pp. 105. 

Under the above title, Prof. Bailey has grouped a number of 
chapters which we may well designate as bits of floral poetry, 
Written in prose. The author does not attempt to give a list of 
Rhode Island plants, but to call attention to the fact that the 
flowers of the State are of surpassing beauty and interest. Rhode 
Island lies on the line dividing our northern and southern floras, 
and hence contains many interesting forms. Does 

After an introduction, we have a scheme of plant distribution, 
showing the wide variation of plant life, in plants of the salt water, 
the salt marsh, the river, the sea beach, the bog, the forest and the 


* The American publication maintains Pneumaria and Mertensia as distinct 
enera, 


408 


like. Nearly half the book is taken up with a description of ule 
Favored Spots, and this constitutes the chief charm. A few choice 
plants are mentioned in each case, while a delightful picture of 
the locality is presented, enticing the botanist to visit the spot 
and search for the treasures growing there. Especially interest- 
ing is the chapter on Wild Flowers of Block Island, where the 
presence or the absence of certain forms is suggestive of specula- 
tion as to the distribution of plants. The weeds of the State 
claim a separate discussion. Interesting to a botanist they are, as 
Prof. Bailey says, “A flower garden is lovely ; but for vital, every- 
day, continual interest, a weed-grown yard is far-and-away more 
fascinating.” 

A list of Rhode Island trees is appended. We welcome the 
little book, for it cannot fail to fill its readers with a love for our 
wild flowers in general. WALTER DEANE. 


Proceedings of the Botanical Club, A. A. A. S., Springfield Meet- 
ing, August 29th to September 2d, 1899. 


The meetings were held in the room assigned to Section “G, 
in the State Street Baptist Church. 


Tuurspay Morninc, AUGUST 29TH. 


In the absence of the President, Prof. D. H. Campbell, and of 
the Secretary, Prof. F.C. Newcombe, the meetings of the Club 
were placed in organization by Prof. Geo. F. Atkinson. Hon. 
David F. Day was made Chairman pro tem. and Prof. H. L. Bolley, 
Secretary. 

On motion-of Professor Atkinson, those having papers to pre 
sent were requested to hand titles of the same to the Secretary 
upon the day preceding that upon which it was wished the pape 
should be read. 


The meeting adjourned at 11:30 to meet at 9 A. M., Friday, 
August 30th. 
Fripay Morninc, Aucusr 30TH. 


The Club met as ordered, with President D. H. Campbell ae 
the chair. In order to facilitate the reading of papers, the titles 0 


409 


which for the first time in the history of the Club now appeared 
Printed in the regular daily program of the A. A. A. S., the 
reading of the minutes of the previous meetings was dispensed 
with. 

The first paper presented was on “Crimson Clover Hair-balls,” 
by Mr. F. V. Coville. These balls, composed of the hairs of the 
Crimson Clover, Trifolium incarnatum, had been found in the 
stomach of ahorse. Mr. Coville exhibited specimens, also mounted 
slides showing their composition. 

Professor Byron D. Halsted reported the results of field ex- 
Periments with beans. He had found that 25 per cent. of plants 
stown on soil previously occupied by beans were affected by spot, 
Whereas when grown on new soil only 6 per cent. were diseased. 

Mrs, Elizabeth G. Britton reported corrections upon descrip- 
tions of Coscinodon. 

Mr. O. F. Cook remarked upon “A peculiar Habit of a Li- 
berian Species of Polyporus,’ and exhibited specimens showing 
various degrees of prolification, one pileus arising from another 
upon very extended delicate stalks, due perhaps, to the extreme 
Moisture of their environment. 

An apparatus for the bacteriological sampling of well-water 
was described and illustrated by Professor H. L. Bolley, the merits 
of which were facilities afforded for sterilization in toto, and in 
general accuracy of work afforded without contamination by air 
and water. 

Mr. C. L. Pollard described the methods of work in the Na- 
tional Herbarium. The colored labels in use to designate type 
Specimens were of especial interest, because of the new range offered 
for convenience of reference. 

Passing to order of unfinished business Dr. Trelease called for 
the report of the committee appointed at the Rochester meeting 
to prepare and print a check list of the plants of northeastern 
North America. Dr. N. L. Britton, as chairman of the committee, 
Submitted the appended report: 


_ “The committee reports that it has completed the task as- 
Signed it by the Club at its Rochester and Madison meetings, by 
Preparing, to the best of its ability, a list of plants in accordance 
With instructions received at those meetings. The committee 


410 


herewith presents a printed copy of such list, which has been pre- 
pared and published without expense to the Club. 
« For the committee, 
“«N. L. BRITTON, 
“ Chairman.” 

Mr. O. F. Cook, seconded by Dr. F. H. Knowlton, moved the 
acceptance of the report. After some discussion as to the scope of 
the term “acceptance” as here moved, an adjournment was taken 
until afternoon without action being taken upon the motion. 


Fripay AFTERNOON, AUGUST 30TH. 


Following the regular session of Section “G” the Club, upon 
further discussion, adopted the motion of Mr. Cook to accept the 
report. 

On motion of Prof. L. H. Bailey the Club then proceeded to 
the discussion opened in the morning by passing the regular pro- 
gram. 

On motion of Mr. F. V. Coville, seconded by Prof. E. L. 
Greene, and carried, it was resolved that the meeting proceed to 
a discussion of the principles on which the list was based. 

Dr. B. L. Robinson then alluded to certain generic names which 
he thought had been inconsistently employed in the list. He also 
discussed the admission of specific names first published as syn 
nyms; the practice of admitting such names was defended by 
Prof. Greene, who maintained that the practice of “ taking up of 
synonyms” as used by the committee was a principle established 
by Gray. 

Professor N. L. Britton also maintained that the principles 
adopted by the Club at the Rochester meeting required the admis- 
sion of such synomyms as those cited by Dr. Robinson. 

After much rambling discussion the following resolution 
offered by Professor Britton, and seconded by Professor L. H. 
Bailey, was adopted : 

Resolved, That in view of the opinions which have been eX- 
pressed at home and abroad on principles of nomenclature, during 


the progress of the work of the committee, the matter be referre 


to the committee for consideration and report at the next meeting 
of the Club. 


Prof. Britton also introduced the following resolution : 


411 


Resolved, That the committee be increased to eleven members 
by the additions of Dr. B. L. Robinson and Dr. C. S. Sargent. 


At this point Dr. B. L. Robinson stated that because of the 
radical difference of opinion existing between himself and the ma- 
jority of the present committee upon certain vital points, it was 
plain to him that he must decline to serve upon the committee. 
In compliance with these wishes, the Club reluctantly accepted Dr. 
Robinson’s withdrawal, and upon motion, Professor L. H. Bailey’s 
name was substituted in the resolution, and the same adopted as 
amended. The Club then adjourned to meet at the same place at 
9 A. M., Monday, September 2d. 


Monpay MorninG, SEPTEMBER 2D. 


Prof. N. L. Britton, Dr. W. H. Seaman and Mr. Walter Deane 
were appointed a committee to nominate officers for the next 
meeting, 

The report of Treasurer F. C. Newcombe, showing the bal- 
ance in hand, $6.57, was read and accepted, as here appended: 


ANN ARBOR, MICH., August 26, 1895. 


BOTANICAL CLUB, A. A. A. 5S. 
Report of the Treasurer for the year ending August 26, 1895. 


1894, RECEIPTS. 
August 17, Voluntary contributions from members present at Brooklyn... . $7.32 
1895. DISBURSEMENTS. 
August 26. Express charges on records, Ann Arbor to Springfield. . . . . 75 
Balance ow hand, 9). 6. ee 6.57 


The first paper was read by Mrs. Elizabeth G. Britton, entitled 
“Some Notes on Dicranella heteromala and allied Species.” 

Prof. J. C. Arthur described a new form of clinostat, and re- 
marked on its advantages over similar machines previously con- 
structed, its great superiority being multiple arms for holding 
Plants, allowing of checks upon tests made. 

A paper by Mr. A. B. Seymour describing the Mary A. Gil- 
breth collection illustrating the dissemination of seeds, now the 
Property of Radcliffe College, was read by Mrs. Flora W. Pat- 
terson. 


412 


Judge David F. Day described the dissemination of the seeds 
of Zinnia by means of the persistent ray-flowers. 

Mr. Walter Deane mentioned the expulsion of the seeds from 
the capsules of the Witch-hazel, Hamamels Virginica, stating that 
he had observed them strike a pane of glass fourteen feet away with 
almost force enough to crack it. 

Judge Day spoke also on the desirability of further observa- 
tions on climbing plants, referring to his observations on the genus 
Dioscorea, some species of which twine in one direction, others in 
another. He mentioned Aconitum uncinatum as a twining plant, 
and had observed a secondary peduncle in Anemone Virginiana 
twining around the primary one. 

The following papers were read by title during the meetings: 

« Notes upon Pig-nut Hickories,” by William Trelease. 

« Experiments with Lime as a preventive of Club-root,” by B. 
D. Halsted. 

“Notes on the alkaline Reaction of the vascular Juices of 
Plants,” by Erwin F. Smith. 

“Continuation of Experiments upon the Relation between the 
fertile and sterile Leaves of Onoclea,’ by George F. Atkinson. 

“A Hybrid between an Egg Plant and Tomato Plant,” by P. 
H. Rolfs. : 

‘‘A Method of using Formalin Gelatine as a Mounting Me- 
dium,” by A. F. Woods. 

The committee appointed to nominate officers submitted the 
following names and they were unanimously elected : 

President, Frederick V. Coville, Washington, D. C. 

Vice-President, Conway MacMillan, Minneapolis, Minn. 

Secretary and Treasurer, J. F. Cowell, Buffalo, N. Y. 

The Secretary was requested to append to the minutes for 
future reference, a list of persons who have been officers of the 
Club, since its formation. : 

The Club then adjourned to meet as usual during the meeting 
of the Association in 1896, 

Fifty-three botanists were registered during the different ses- 
sions. 

H. L. Bo.tey, 
Secretary pro tem. 


413 


Titles of Papers read before the Section of Botany, A. A. A. S., 
Springfield Meeting. 


The following were presented either in full, in abstract or by 
title. A summary of the proceedings may be found in the Sep- 
tember issue of the « Botanical Gazette.” 

A Leaf Rot of Cabbage. By H. L. Russett. 

The Southern Tomato Blight. By Erwin F. Suita. 

Observations on the Development of Uncinula spiralis. By 
B. T. Gattoway, 

The Effect of Sudden Changes of Turgor and of Temperature 
on Growth. By Ropney H. True. 

Recording Apparatus for the Study of Transpiration of Plants. 
By Atzert F. Woops. 

Pressure, Normal Work and Surplus Energy in Growing 
Plants. By Grorcr M. Hourerty. 

Notes on the Ninth Edition of the London Catalogue of Brit- 
ish Plants. By N. L. Brirron. 

Obolaria Virginica L. A Morphological and Anatomical Study. 
By Turo. How. 

Botany of Yakutat Bay, Alaska. By Freperick V. CoviLte. 

Variation after Birth. By L. H. BatLey. 

Rejuvenation and Heredity. By Cas. S. Minor. 

The Distinction Between Animals’ and Plants. By J. C. 
ARTHUR. 

Fungous Gardens in the Nests of an Ant, Ata tardigrada 
Buckl., near Washington, D.C. By WatrTer T. SWINGLE. 

Poisoning by Broad-leaved Laurel, Kalmia latifolia. By FReD- 
ERICK V. Covitte. 

The Physiology of Jsopyrum biternatum L. D. T. Mac- 
Doveat. 

The Transmission of Stimuli-effects in Mimosa pudicaL. By 
D. T. MacDoveat. 

Personal Nomenclature in the Myxomycetes. By O. F, Coox. 

A New California Liverwort. By Doucras H. CAMPBELL. 

The Number of Spore Mother Cells in the Sporangia of Ferns. 
By Witus L. JEPson. 


414 


The Constancy of the Bacterial Flora of Fore Milk. By inf 
L. BOLLEY. 

The Watermelon Wilt and other Wilt Diseases due to Fusarium. 
By Erwin F. Situ. 

Antidromy in Plants. By GrorGe MACLOSKIE. 


Proceedings of the Botanical Society of America, Springfield Meet- 
ing, Aug. 26th and 27th, 1895, 


The botanists conducted their sessions at the meeting of the 
Botanical Society of America with Prof. Wm. Trelease, President, 
in the chair, and Prof. C. R. Barnes, Secretary. They elected 
Prof. C. E, Bessey, President; Prof. W. P. Wilson, Vice-President; 
Prof. C. R. Barnes, Secretary, and Mr. Arthur Hollick, Treasurer, 
for the ensuing year. They also elected Dr. A. W. Chapman an 
honorary member; and Mr. M. S. Bebb, Prof. W. R. Dudiey, Prof. 
D. P. Penhallow and Prof. W. A. Setchell as active members. It 
was decided to deposit such books and pamphlets as may be re- 
ceived for the Society, in the Library of the Missouri Botanical 
Garden. 

Most of the papers presented were brief summaries of larger 
works and enterprises, on which the authors ‘are engaged. 

_ Mrs. N. L, Britton read a paper on ‘Some Notes on a Revision 
of the genus Mnium,” in which an attempt was made to show how 
nearly the American specimens correspond with European types 
and descriptions. 

Dr. N. L. Britton gave a brief account of the history of the 
founding of the New York Botanical Garden, describing the site 
in Bronx Park and outlining the plans of the board of managets- 

Prof. L. H. Bailey presented the following resolution which 
was passed unanimously : 

« Resolved, That the Botanical Society of America express its 
thanks to Dr. N. L. Brinton for his account of the condition an 
progress of the movement for a botanical garden in the City of 
New York, and congratulate the people of that city on the pros” 
pect of its rapid development; and, furthermore, that the Society 
commend the board of managers of the garden and its board 0 


_ scientific directors for their wisdom in securing a broad foundation 
and an assurance of liberal management.” 


415 


Prof. George F. Atkinson presented a paper as a contribution 
to the knowledge of North American phycophilous fungi. 

Mr. Arthur Hollick exhibited a series of plates illustrating the 
fossil leaves of Liriodendropsis, contrasting them with the stages of 
development of the leaves in Lirtodendron, showing that there is a 
parallelism between the shapes of the earliest rudimentary leaves 
of the living species with that of the development of the fossil 
ones. 

Prof. Roland Thaxter exhibited the plates of a monograph on 
the Laboulbeniaceae which he is preparing, explaining the struc- 
ture of these minute fungi, which grow on living insects, and de- 
Scribing several new genera and species. He also made a few 
remarks on the phycomycetous fungi, and showed diagrams of 
one new genus and species. 

Mr. F. V. Coville gave a brief abstract of his revision of the 
genus Funcus, subdividing it into subgenera, and recognizing 
seventy-six species in North America. 

Prof. Chas. R. Barnes made remarks on a revision of the genus 
Dicranum which he is preparing, with the assistance of Mr. Rod- 
ney H. True, recognizing thirty-three North American species, 
and explaining his reasons for rejecting several recently described. 


Index to recent Literature relating to American Botany. 
Anderson, A. P. The Grand Period of Growth in a Fruit of Cucur- 
bita Pepo, determined by Weight. Bull. Geol. Nat. Hist. Surv. Minn. 
9! 238-279. pl. rz7-20. 5 Mr. 1895. 
Bessey, C. E. A Protest against the ‘‘ Rochester Rules.” Am. Nat. 
29: 666-668. Jl. 1895. 


Opposes the Harvard herbarium recommendations. 

Bessey, C. E. Progress of the Botanical Survey of Nebraska. Am. 
Nat. 29: 580-582. Je. 1895. 

Bolander, H. N. Anew Erythronium, Eryth. 3: 127. 15 Au. 1895. 
Erythronium Fohnsonii from southern Oregon. 


Coulter, J. M. and Rose, J. N. Deanea, a new Genus of Umbelli- 
ferae from Mexico. Bot. Gaz. 20: 372. f/ 27- 15 Au. 1895. 


416 


Coville, F. V. Juncus scirpoides and its immediate Relatives. Bull. 
Torr. Bot. Club, 22: 302-305. 31 Jl. 1895. 


Davenport, G. E. Daniel Cady Eaton. Bot. Gaz. 20: 366-369. 
pl. 26 A. 15 Au. 1895. 


Greene, E. L. Novitates occidentales.—XVI. Eryth. 3: 123-126. 
15 Au. 1895. 


Describes new species in Clarkia, Eriophyllum, Erigeron, Allocarya and Callt- 
préora. 


Holm, T. A Study of some anatomical Characters of North American 
Gramineae.—IV. Bot. Gaz. 20: 362-365. p/. 26. 15 Au. 1895. 


Describes the leaf-structure of Leersia. 


Holzinger, I. M. A Preliminary List of the Mosses of Minnesota. 
Bull. Geol. Nat. Hist. Surv. Minn. 9: 280-294. 5 Mr. 1895. 


Jack, J. G. Another Herbarium Pest. Gard. & For. 8: 323-324-/- 


45. 14 Au. 1895. 
Hlustrating and describing Zphestia interpunctella. 


Jack, J.G. The Hazels. Gard. & For. 8: 344-346. f. 8. 28 Au. 
1895. 
Notes on Corylus rostrata. 
Johnson, L. N. Some new and rare Desmids of the United States. 
—II. Bull. Torr. Bot. Club, 22: 289-298. p/. 2379, 240. 31 Jl. 1895- 
Describes new species in Arthrodesmus, Cosmocladium and Cosmarium. 
Kellerman, W. A. On Plant Names. Journ. Columbus Hort. Soc. 
10: 7-10. Mr. 1895. 
Maintains the necessity of adherence to priority. 
La Mance, L. S. Jris hexagona. Gard. & For. 8: 329. 14 Au. 
1895. 
Recording the discovery of /ris hexagona La Mance Gerard, in Benton County, 
Arkansas, 
Lotsy, J. P. Some Euphorbiaceae from Guatemala. Bot. Gaz. 20: 
348-355. pl. 24, 25. 15 Au. 1895. 
Describes a number of new species. 
Magnus, P. Die Teleutosporen der Uredo Aspidiotus Peck. Ber. 
Deutsch. Bot. Gesell. 13: 285-288. f/. 27. 25 Jl. 1895. 
Mez, C. Einige Bemerkungen iiber Midudariopsis. Ber. Deutsch. 
Bot. Gesell. 13: 236-239. 25 Jl. 1895. 


Nash, G. V. The Genus Cenchrus in North America. Bull. Tort- 
Bot. Club, 22: 298-302. 31 Jl. 1895. 


417 


Pammel, L. H. Some troublesome Iowa Weeds. Rep. Iowa Agric. 

Soc. 1894: 523-539. 1895. 

Figures of Salsola Kali var. Tragus, Arctium Lappa, Xanthium Canadense, 
Ambrosia trifida, Hordeum jubatum, Capsella Bursa-pastoris,Cuscuta Epithymum, 
with accounts of these weeds, 

Pammel, L. H. The Geographical Distribution of Plants. Rep. 

Towa State Hort. Soc. 29: 324-433. 1895. 

Penhallow, D. P. Rhus Poisoning. Gard. & For. 8: 359. 45S. .- 

1895. 

Robinson, B. L. The Nomenclature Question. Bot. Gaz. 20: 370- 

371. 15 Au. 1895. 

Rolfe, R. A. New Orchids. Kew Bull. 104: 191-195. Au. 1895. 


Tropical American species in Pleurothallis, Batemaria, Maxillaria, Notylia and 

LPelexia, 

Rolfe, R. A. Vanillas of Commerce. Kew Bull. 104: 169-178. 
Au. 1895. ae 
An historic and descriptive account of the species yielding aromatic fruits. Two 

are described as new. 

Rydberg, P. A. New Species of Pfysalis. Bull. Torr. Bot. Club, 
22: 306-308. 31 Jl. 1895. 

Pca fe. Bue aa Beobachtungen und kritische Bemerk- 
ungen iiber die Exoascaceae. Ber. Deutsch. Bot. Gesell. 13: 265- 
280. p/. 27, 25 Jl, 1895. 

References to many American species. : 

Sargent, C. S. The American White Birches. Gard & For. 8: 355. 
J. 50. 48. 1895. 

Note on Betula populifolia and B. papyrifera. oo 

Setchell, W. A. On the Classification and Geographical Distribution 
of the Donenene Trans. Conn, Acad. Arts and Sci. g: Part 2, 
333-375. 1895. 

Sheldon, E. Pp. Compilation of Records of some Minnesota ayia 
ing Plants. Bull. Geol. Nat. Hist. Surv. Minn. 9: 223-227. 5 Mr. 
1895. 

Smith, J. B. Why certain Hickories died. Gard. & For. 8: 352- 
353-7. 49. 48. 1895. 
Tilden, J. E. List of fresh-water Algae collected in eee eg 

1894. Bull. Geol. Nat. Hist. Surv. Minn. 9: 228-237. 5 Mr. 1895. 

Toumey, J. W. Opuntia fulgida, Gard. & For. 8: 324-326. f. 46. 

14 Au. 1895. 


418 


Toumey, J. W. Vegetal Dissemination in the Genus Opuntia. Bot. 
Gaz. 20: 356-361. 15 Au. 1895. 


Uline, E. B., and Bray, W. L. Synopsis of North American Ama- 
ranthaceae—Ill. Bot. Gaz. 20: 337-344. 15 Au. 1895. 

Ward, F. L. The Nomenclature Question. Bull. Torr. Bot. Club, 
22: 308-329. 31 Jl. 1895. 

Wilcox, E. M., Chairman. Report of the Committee on Botany. 
Journ. Columbus Hort. Soc. 10: 5. Mr. 1895. 


Record of species of Folyforus near Columbus, Ohio. 


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TERATOLOGY IN QUERCUS GARRYANA. 


Surplus Books for Sale from the Library of Dr. M. C. Cooke. 


146 Junction Road, London N., England. 


Balfour, J. B. Botany of Socotra. 4°. 100 plates. Edinburgh. 
Hedwig. Theoria generationis and Hist. Nat. Muscorum. 4°. 
Lamarck and DeCandolle. Flore Francaise. 6 vols., 8°. 
® Steudel. Nomenclator Botanicus. 8°, board. 
! Linnaeus. Flora Suecica. First Edition. 
Leers. Flora Herbornensis. 16 pl., 8°. 
Acharius. Synopsis Lichenum. 
Marsilius. Dissert: de generatione Fungorum. 
Retz. Observationes Botanicae. Folio. 
Vaillant. Botanicon Parisiense. Folio. 
Queckett. Histological Catalogue. 4°. 2 vols., cloth. 
Luerssen, Medico-Pharmaceutik Botanik. Vol. 1, Kryptogamen, 8°, paper. 
Lindley, Dr. Flora Medica. 1 vol., cloth. 
Waring, Dr. Pharmacopaeia of India. 8°, cloth. 
Fluckiger, Dr., and Hanbury, D, Pharmacographia. 8°. 
Hanbury, D. Notes on Chinese Materia Medica. 
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Christison, Dr. On Poisons. 8°, 1 vol. 
Allen and Thomson. Expedition to the Niger. 8°, 2 vols. 
Farlow, W.G. Marine Algae of New England. 8°, paper, 
Toni and Levi. Flora Algologica Venezia. Parts I. and III. 
Toni and Levi. L’ Algarum Zanardini. 8°. 
Underwood. Catalogue of N. Am. Hepaticae. 8°. 
Tuckerman. Synopsis of Lichens of New England. 
Bescherelle. Prodromus Bryologiae Mexicanae. 8°. 4 
__ Harvey, Dr. Phycologia Britannica. 4 vols. in 2 half morocco, gilt tops. Splen 
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Queckett, J. Lectures on Histology. 2 vols., 8°, cloth. 
Queckett, J. Treatise on the Microscope. 8°, cloth. 
Humboldt. Flora Friburgensis. 4°, board. : 
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De Notaris. Epilogo della Briologia Italica, Roy. 8°, half calf. 
Wahlenberg. Flora Suecica. 2 vols. 8°. 
Bauhin, Theatrum Botanicum. Folio. 
Meyer, Primitiae Florae Essequeboensis. 4°, calf. 
Sullivant. Musci and Hepaticae of U.S. 8°. ; 
Kickx, J. Flore cryptogamique des Flandres. 2 vols. in 1, Half calf. 1867. 
Roth, A.G. Catalecta Botanica. 3 vols., 8°, half calf. 1806. 
Schrader, H. Spicilegium Florae Germanicae. 8°. 1794. 
Haller, A. Flora Jenensis. 12°. 1745. 
Weigel. Flora Pomerano Rugica, _ 12°, 1769. 
Olhaff. Elenchus Plantarum. 12°. 1656. 
Sprengel, C. Florae Halensis. 8°. 1806 and 2 suppl. 
umacher, Plantae Saellandiae. 2 vols., 1801. : 
Schlechtendal. Flora Berolinensis. Part 2, Cryptogamia. 1824. 
Wulff, J.C. Flora Borussica, 12° 1765. 
Schrank. Florae Salisburgensis. 12, 1792. | 
einmann, Enum: Stirpium in Agro Petropolitano. 8°, 1837. 
Martius. Flora Erlangensis Crypt. 8, 6 pl., 1817. 


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calf, rest in parts. : 
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Hedwigia (Ed. Rabenhorst, etc.), 1852 to 1891 (Vol. II. imperfect) together in 
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21). 
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Revue Mycologique (Roumeguere). 1888-1891. 13 vols. in 6, cloth. 
Rabenhorst’s Algae Europaea Aquae dulces. 3 parts in 2 vols., half calf, com- 
plete, 1864. 
Monthly Microscopical Journal. (Ed. Dr. Lawson.) 18 vols., cloth. 1869-1 877. 
Grevillea (Ed. M. C. Cooke). Complete set in parts, 20 vols., 1872-1892. 
Complete sets out of print. : 
Hassalls, Dr. British Fresh Water Algae. 2 vols., cloth. 
Smith, W. British Diatomaceae. 2 vols., cloth. 
Ralfs, J. British Desmidiae. 1 vol., half calf. ' 
Bornet et Thuret. Notes Algologiques, part 2 only, folio, plates. 
Flora of Herefordshire. Mosses by Aug. Ley. Fungi by M. C. Cooke. 1 vol. 
8°, cloth. 1889. 
PO plies W. J.,and Baker, J.G. Synopsis Filicum. Colored plates, 8°, cloth. 
I 


DeBary, A. Morphologie und Biologie der Pilze. Mycetozoen und Bacterien. 
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vols, (wanting Vol. VII.). 

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in parts. Also 1887-8 complete; 1889 (wanting 266, 270, 274, 275); 1890 complete; 
1891 (wanting 298, 300). 

American Entomologist, and American Botanist and Entomologist. 1868-1870. 
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Ray (John). Historia Plantarum. 3 vols., folio. Ren: 

_Mitten, W. Musci Indiae Orientalis. 8°, with Griffith’s Muscologia Itineris As- 
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Buxbaum J.C. Plantarum minus cognitarum. Complete in 2 vols, 4°. 

Girod Chautraus. Recherches chimiques et micro. sur les Conferves. Fol. 36 pl. 
half calf. 1802, i 

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VoLuME I. 
A List of Plants Collected by Miss Mary B. Croft at San Diego, Texas. By 
N. L., Brittonaml HH, Rasby (6887); «oo es | 25 cents. 
New or Noteworthy North American Phanerogams. By N. L. Britton 
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A List of Plants Collected by Dr. E. A. Mearns at Fort Verde and in the 
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and the Adjacent Region. By H. H. Rusby (1888),. . . . . 25 cents. 
Preliminary Notes on the North American Species of the Genus 7éssa, 
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Sco dont. Hae eR ia ne rs Las 5S Oem 
Ranunculus repens and its Eastern North American Alhes. By N. L. 
PW Pi a er ee 
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hg it Cone wee FO ee 
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72893) ee CG re eee eC Sore, 


Miss Anna Murray Vail (1892-1893), «s+ tt ttt ee $1.50 
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Contributions to American Bryology, VII. A revision of the Genus Phys- 
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WAS A SEE RE Se oon ee le Cee 


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New or noteworthy American Grasses.—I. 


By Gero. V, NAsH. 


_ ERIANTHUS COMPACTUS 7. sp. Culm erect, 4°—8° tall, stout, 
its nodes barbed, its summit and the axis of the panicle densely 
pubescent with appressed long rigid silky hairs. Sheaths glabrous 
or pubescent at the apex; leaves scabrous above, sparingly ap- 
Pressed-pubescent beneath, 6’-2° long, 3/’-6” wide, long-acumi- 
nate, narrowed toward the base; panicle narrowly oblong, 4'-6/ 
In length, about 114’ wide, branches erect, 1/-2’ long; spikelets 
Crowded, the internodes of the rachis about 1’ long; outer 
Scales of the spikelet about 21%” long, exceeding the pedicels 
and about equalling the basal hairs, lanceolate, acuminate, pubes- 
cent with long spreading hairs; inner scales shorter, the awn 
5,-10” long, straight, scabrous. 

In moist or wet soil, New Jersey to North Carolina and Ten- 
nessee. The compact panicle composed of short erect branches, 
the short internodes of the rachis, and the crowded small spike- 
lets readily distinguish this plant from the other straight-awned 

Species, 

SYNTHERISMA Walt. Fl. Car. 76 (1788). : 
[Digétaria Scop. Fl. Carn. Ed. 2, 1: 52.1772. Not Heist. 
1763]. 

_ The Digitaria section of Panicum is certainly sufficiently dis- 
tinct to take generic rank and it has been so regarded by many 
authors. Panicum as now generally understood comprises too 
‘Many forms. It would seem preferable to restrict the genus to the 


Eupanieae. 


iis: hela ia Bag Oi 


420 


The name Digitaria was originally applied by Heister (Adans. 
Fam. Pl. 2: 38. 1763),to the Zripsacum ot Linnaeus, and ante- 
dates by nine years the Digitaria of Scopoli (FI. Carn.'Ed. 2,1: §2. 
1772), who applied it to a group of plants that have since been 
considered by many as only a section of Panicum. This name 1s 
therefore a synonym of Zripsacum and not available. The oldest 
available name appears to be Syxtherisma of Walter. A partial 
sysonymy of the three species occurring in the north-eastern 
United States would be as follows: 


1. SYNTHERISMA SANGUINALIS (L.). 

Panicum sanguinale L. Sp. Pl. 57. 1753. 

Digitaria sanguinalis Scop. Fl. Carn. Ed. 2, 1: 52. 1772. 

Syntherisma praecox Walt. Fl. Car. 76. 1788. 

Paspalum sanguinale Lam. Tabl. Encycl. 1: 176. 1791. 

2. SYNTHERISMA LINEARIS (Krock.). 

Panicum lineare Krock. FI. Sil. 1: 95. 1787. 

Syntherisma serotina Walt. Fl. Car. 76, 1788. 

Digittana humifusa Pers. Syn. 1: 85. 1805. 

Syntherisma glabra Schrad. Fl. Germ. 1: 163. 1806. 

Panicum glabrum Gaud. Agrost. 1: 22. 1811. 

3. SYNTHERISMA FILIFORMIs (L.). 

Panicum filiforme L. Sp. Pl. 57. 1753. 

Paspalum filiforme Sw. Prodr. 22. 1788. 

Syntherisma villosa Walt. Fl. Car. 74. 1788. 

Digitaria filiformis Muh\. Gram. 131. 1817. 

Panicum PorTERIANUM nom. n. 

Panicum latifolium Walt. F\. Car. 73. 1788. Not Linneus, 
1753. 

Panicum Walteri Poir. in Lam. Encycl. Suppl. 4: 282. 1816. 
Not Pursh, 1814. 

Panicum latifolium var. molle Vasey, Bull. Bot. Div. U. 5. 
Dept. of Agric. 8: 33. 18809. 

Search has failed to disclose an available published name Aer 
this plant. The Linnaean name belongs to a tropical Specie®: 
The P. Walteri of Poiret is antedated by two years by Pursh's 
name which was applied to an entirely different plant, the F- Crus- 
galli vax. hispidum of Torrey. The varietal name of Vasey has 


421 


already been used. It is necessary, therefore, to give this well- 
known plant a name, and the above is proposed in honor of Prof. 
Thos. C. Porter. 


Panicum ScRIBNERIANUM nom. n. 

Panicum Scopartum S. Wats. in A. Gray, Man. Ed. 6, 632. 
1890. Not Lam. 

Panicum scoparium var. minor Scribn. Bull Univ. Tenn. 7: 48. 
1894. Not P. capillare var minor Muhl. 1817. 

This plant is certainly not the P. scoparium of Lamarck. 
That is found only in the southern mountains and is a tall large- 
leaved grass with a few-flowered panicle. There seems to be no 
available name published for this plant, and so the above is pro- 
posed, in honor of Prof. F. L. Scribner, who was the first to indi- 
Cate its difference from P. scoparium Lam. 


“Panicum minus (Muhl.). 

Panicum diffusum Pursh, Fl. Am. Sept, 1 : 68. 1814. 

Panicum capillare var. minus Muhl. Gram. 124. 1817. 

Panicum capillare var. sylvaticum Torr. Fl. U.S. 149. 1824. 

Panicum Philadelphicum Bernh; Nees. Fl. Bras. 198. 1829. 

This plant is readily distinguished from P. capillare, of which 
it has been considered a variety by its more slender habit, and 
smaller spikelets in pairs at the extremities of the ultimate divi- 
Sions of the smaller and less branched panicle. 

PANICUM BOREALE n.sp. Culmsat first erect and simple, 1°—2° 
tall, later sometimes decumbent and somewhat branched, smooth 
and glabrous. Sheaths shorter than the internodes, smooth and 
glabrous, ciliate; ligule short, ciliate; leaves 3’—5’ long, yy'-¥4! 
Wide, erect, truncate or rounded at the sparsely ciliate base, acumi- 
hate; panicle 2/—4’ in length, ovate, branches 1/~2’ long, spread- 
Ing or ascending; spikelets 1’ long, about equalling the pedicels, 
ellipsoid, pubescent; first scale ovate; obtusish, about one-third the 
length of the spikelet; second and third ovate, 7-nerved, pubes- 
cent, equalling the fourth; fourth oval, chartaceous, acute, slightly 


exceeding 3/” in length. 

Moist soil, Newfoundland and Ontario to New York, Vermont 
and Minnesota. This plant was first noted by the writer in 1893 
in the Catskill Mts., near Cario, N. Y. The smaller forms some- 
what resemble P. laxiflorum, but the glabrous sheaths and shorter. 
leaves, and the different shaped spikelets readily distinguish it 


422 


from that species. No. 222, Robinson and Schrenk, 1894, and 
Macoun’s plant, collected at Windsor, N.S., June 29, 1883, be- 
long here. 

PANICUM SPHAGNICOLUM n. sp. Culms 2°—3° long, at first 
simple, later repeatedly dichotomously branched, the branches 
very divergent. Primary sheaths hirsute, those of the branches 
glabrous; ligule a ring of short hairs; leaves smooth and glabrous, 
the primary 2/-3% long, 2’’-5’” wide, those of the branches é 
long or less, %4/’-114” wide, appressed ; primary panicle about 2 
in length, the branches 1%4’-1’ long, spreading; secondary pan- 
icles about 14’ long, raceme-like; spikelets 11/’” long, few and 
appressed, pubescent; first scale ovate, acute, I—3-nerved, about 
half the length of the spikelet; second scale broadly ovate, 
7-nerved, somewhat shorter than the 7—9-nerved third one, the 
fourth chartaceous, smooth, elliptic, acute, slightly exceeding 1 
in length and equalling the third. 


The late and much branched state was collected by the writer 
this summer in a sphagnum bog at Lake City, Florida, and will 
be distributed as No. 2500. The main stem rises through the 
sphagnum and then divides dichotomously, the branches spread- 
ing. out over the surface of the moss, the ends being erect. The 
early and simple form was found by A. W. Bitting in a cypress 
swamp at the same place, on April 2, 1892. 


IXOPHORUS Schlecht. Linnaea, 31: 420. 1861-62. 


[SerariA Beauv. Agrost. 113. 1812. Not Ach. 1798.] 


Otto Kuntze (Rev. Gen. Pl. 767) has referred these grasses to 
the genus Chamaeraphis R. Br., and this disposition of them has 
been accepted by several writers. But Chamaeraphis 1s a tropical 
genus in which the spikelet and bristle fall attached, while in the 
plants in question the spikelet articulates above the bristles, hence 
‘leaving them persistent when it falls. Feeling confident that the 
two groups are generically distinct, I have sought for a published 
name for the latter, and find /xophorus of Schlechtendahl to be the 
earliest one available. It is based on Uvochloa uniseta Presl, which 
is Se¢aria uniseta Fourn. The forms occurring in the northeastern 
United States are as follows: 


1.“ IXOPHORUS VERTICILLATUs (L.). 
Panicum verticillatum L. Sp. Pl. Ed. 2, 82. 1762. 
Setaria verticillata Beauv. Agrost. 51. 1812. 


423 


2. IxoPHORUS GLAUCUS {Lay 
Panicum glaucum L., op. PL 56: 1753. 
Sefaria glauca Beauv. Agrost. 51. 1812. 


3. Ixopnorus Iraricus (L.). 
Panicum Ttalicum L. Sp, Plog6: 1753. 
Setaria Italica R. & S. syst. 47 493, 1817. 


4. IXOPHORUS ViRIDIS uy 
Fanicum viride L. Sp. Pl, Ed. 2, 835 170. 
Setaria viridis Beauv. Agrost. 51. 1812. 

“STIPA AVENACIOIDES n. sp. Culms 2°-4° tall, simple, erect, slen- 
der, smooth or puberulent below the lower nodes. Sheaths about 
half the length of the long internodes, smooth and glabrous; ligule 
obtuse or acute, I’” long or less; leaves involute-setaceous, smooth 
and glabrous, the basal about 1° long, those of the culm eed 
long, the uppermost often nearly wanting; panicle simple, 4-10 
in length, finally long-exserted, the scabrous branches 1’—3/ long, 
the lower in pairs; spikelets few ; lower halves of the empty scales 
Purple, 5-nerved below, acuminate, the lower 8’”—10” in length, the 
upper slightly longer; flowering scale with a ring of short brown 

airs at the apex, brown, otherwise glabrous, 7’’—8’ long, includ- 
ing the callus which is 3/’-4’’ long, and covered with appressed 
brown silky hairs increasing in length toward the top; awn 3'-4 
long, pubescent below, twisted for about half its length, tightly so 
at the base, twice bent. 
Collected by the writer in 1895 in the pine lands near Cassia, 


Lake Co., Florida, No. 2051. 


Camputosus CHapapensis Trin. Spec. Gram. P/. 30}. 

Ctenium Chapadense Doell, Mart. Fl. Bras. 2: Part 3, 7 3. 1878. 

This plant was found growing plentifully by the writer this 
Summer in the flat woods at Lake City, Florida, no. 2212. A. H. 
Curtiss collected the same in East Florida in 1875; his no. 4058 
also appears to be this plant. It was also secured by Fredholm 
in Duval Co., Fla., in 1893, no. 313. : 

It differs from C. aromatica (Walt.) Scribn. (Ctenium —"* 
canum Spreng.) in its more slender habit and narrower spikes ; 
smaller spikelets divergent from the rachis, not oe the 
Scales narrower and very acute; the awns more delicate and much 
longer. 


“Cutoris neciecta n. sp. Culms 2°-4° tall, erect, much oe 
Pressed, smooth and glabrous. Sheaths glaucous, compressed, 


424 


those at the base of the culm crowded and equitant; ligule a ring 
of short hairs ; leaves smooth and glabrous, glaucous, 4’-15’ long, 
2-4” wide, abruptly acute, the uppermost culm leaf short, and 
usually remote from the infloresence, or often wanting ; spikes 
4-6, 3'-5’ long, erect or slightly spreading ; spikelets divergent to 
the triangular rachis, which is scabrous onthe angles ; empty scales 
of the spikelet 1-nerved, scabrous on the keel, the first ovate, 
acute, about two-thirds as longas the second; second 2-toothed, 
the teeth obtuse and erose at the apex, oblong, 11%4’’—2” long, 1n- 
cluding the scabrous awn which is 14” long; third scale about 
144” long including the awn which is 14” in length, brown, ovate, 
3-nerved, the nerves pubescent with long ascending hairs; fourth 
scale empty, 1” long, elliptic in outline, cucullate, remote from the 
third, bearing an awn about 14” long just below the apex; seed 


34’’ long, narrowly oval in outline, triangular, translucent, yellow- 
ish streaked with purple. 


Resembles C. Floridana Vasey; that species differs in having 
only 1 or 2 spikes; larger spikelets having 5 scales and 2 flowers, 
the upper one staminate; second empty scale broad and rounded 
at the erose apex; the hairs on the nerves of the third scale 
shorter; the fourth scale obovate and obliquely truncate at the 
apex. 

Collected by the writer near Orange Bend, Lake Co., Florida, 
during the past summer, in low pine lands, no. 2149. It was also 
secured by A. H. Curtiss at Jacksonville in 1875 ; his number 3445 
is the same. Fredholm obtained it along the banks of the St. 
John’s River in Duval Co., Florida, in 1893, no. 319. 


Notes on some Cyanophyceae of New England. 


By WILLIAM ALBERT SETCHELL. 


The-Cyanophyceae or Blue-Green Algae of the United States 
are little represented in collections and exsiccatae, and their occut- 
rence and distribution is in great need of more careful study. For 
many years there has been a lack of good monographs relating to 
the group, and the redescribing under many different names of 
the same form made the matter of the identification both of gen- 
era and of species so difficult that there has been little temptation 


to the ordinary student to pay any attention at all to any of the 
forms. 


425 


Recently, however, there have been published two monographs, 
the results of the patient working over of abundant materials by 
men of such experience and insight that the filamentous species of 
the Cyanophyceae are now arranged and defined in a most satisfac- 
tory fashion. 

The heterocysted forms, multiplying by means of hormogonia, 
are given in the “ Revision des Nostocacées Heterocystées” by 
Bornet and Flahault (Ann. Sci. Nat. (VII.)3: 323-381, 1886; 
4: 343-373, 1886; 5: 51-129, 1887; 7: 177-262, 1888), and the 
homocysted forms with the same kind of reproduction are given 
in the “ Monographie des Oscillariées” by Gomont (Ibid. VII. 
15: 263-368, p/. 6-74, 1892; 16: 91-264, pl. 1-8, 1892.) 

In both of these works a considerable number of species are 
credited to America, chiefly from specimens collected by Farlow, 
Collins, Holden, Wolle and the writer, but many more are to be 
expected and the writer hopes to be able to supply at some future 
time a full list of all found within our boundaries. 

The marine forms have been much more studied than those of 
the fresh waters, and have been mentioned in the papers of Far- 
low and Collins. The account of the species of the fresh waters 
found in Wolle (Fresh Water Algae of the United States, Bethle- 
ham, 1887) is, however, so confused and inexact as to allow no de- 
pendence to be placed upon it. Consequently we must, for the 
Present, accept as a summary of the knowledge of our species the 
information contained in the two monographs mentioned above 
and proceed to work out from them and with their aid. 

The following species, occurring for the most part in New 
England, seem to deserve mention. 

Calothrix Juliana (Menegh.) B. & F., known only in this coun- 
try from Mill Brook, Shelburne, N. H., where it was collected by 
Prof. W. G. Farlow, has occurred to the writer in Massopoag 
Brook, Sharon, Mass., and in Trading Cove Brook, Norwich, Conn. 
tt grew upon stones in small tufts consisting of a few erect paral- 
"lel filaments and could easily be distinguished by the touch, even 
©n stones thickly covered with diatoms and other forms. It is 
fasily distinguished from any other of our species by the absence 
of heterocysts. It grows upon wood as well as stones in the 


Quinebaug River, in Lisbon, Conn. 


426 


C. Braunii B. & F., credited by Bornet and Flahault to Ger- 
many and Denmark, was first found in this country by the writer, 
growing upon stones in a small brook at Sharon, Mass. Later it 
was found near Bridgeport, Conn., by Mr. Isaac Holden, growing 
upon wood and stones. It also forms extended patches on rounded 
stones in a small rivulet at Cataumet, Mass. It is of rapid growth, 
forming patches which are at first small and orbicular, but later 
enlarging and confluent into broad expansions of indefinite ex- 
tent and shape, and often becoming decidedly verrucose on the 
surface. In color the patches vary from dark green to nearly 
black. The filaments are short and parallel and the structure of 
the patch resembles that of /sactis plana, but, of course, lacks the 
common enclosing matrix of that form. 

RIvuLaRIA BorNETIANA n. sp. This curious species was found 
growing on the stems and leaves of Ruppia maritima, forming 
solid spherical thalli varying from somewhat less than a milli- 
meter to something over a centimeter in diameter, according to 
age. Frequently a row of closely-placed spheres coalesce into a 
cylindrical mass 5-8 centimeters long. The thalli are of fairly 
firm consistency and vary in color from a deep bluish-black when 
young to a light olive green or pale yellow when old. They are 
not at all incrusted with lime. The filaments radiate in a regular 
fashion from the center, becoming, however, decidedly flexuous in 
the older thalli. They vary in diameter from 8-20 yz, and separate 
readily from one another on crushing. The sheath is wide and con- 
conspicuous, colorless to deep brown in color, and very much 
lamellose and ocreate above. It varies in width below from 2-4 /” 
sometimes becoming as wide as 20 ». The trichome is usually about 
4 » in diameter, but occasionally reaches 16». It is usually very 
much torulose when young, but very little so when older. The cells 
are from one-halfto five times as long as broad. The cell limits are 
distinct in younger but very obscure in the older specimens, and 
the cell contents are light blue and provided with a few scattered 
granules. The heterocysts are basal, from depressed globular to 
ellipsoidal in shape, and from 6-8 pin diameter. The terminal 
hairs are long and slender in the younger plants, but are almost 
wanting in the older ones. 

For several years this species has been found in abundance !? 


427 


shallow water ahout the edges of Watch Hill Pond, in southwest- 
ern Rhode Island. The pond is separated from the open Atlantic 
Ocean by only a narrow line of sand dunes, through and over 
which the autumnal and winter storms often dash the surf. Con- 
Sequently the water of the pond has a distinctly saltish taste. 
The Rivularia is associated with Anabaena variabilis, Phormidium 
Valderianum, Microcoleus chthonoplastes, Chara aspera, Chara crin- 
aa and various species of Closterium, Scenedesmus, etc. 

This species approaches &. iasolettiana in structure, but dif- 
fers from it decidedly in habit, and it was thought at first it might 
Perhaps be an epiphytic form of that species. But the globular 
shape of the thalli is exceedingly constant, and although carefully 
sought for, typical R. Aiasolettiana was not found anywhere in the 
Same vicinity. It seems best therefore to regard it as a distinct 
Species, and the writer takes great pleasure in dedicating it to Dr. 
Edouard Bornet, of Paris, who has done so much toward furthering 
our knowledge of this group of plants. 

R. Biasolettiana Menegh. occurs in fairly typical form in per- 
fectly fresh water at Gardner's Lake, in eastern Connecticut. It 
forms small flattened thalli on stones along the shores of the lake. 

Capsosira Brebissonit Kuetz., occurring in Sweden, France and 
Germany, grows also in this country upon a large rock on the 
eastern side of Round Pond at Lantern Hill, near Mystic, Conn- 
It forms small hemispherical black thalli from 1-2 mm. in diam. 
eter. They are crushed with some difficulty, but then show the 
Stigonema-like filaments with lateral heterocysts very plainly. It 
©ccurs sparingly also upon submerged dead wood. 

Nostochopsts lobatus Wood, found near Philadelphia by Wood 
and in Vermont by Faxon, has occurred to the writer in some 
abundance at Mt. Carmel, about seven miles north of New Haven. 
The thalli grew upon stones in more or less rapid water in a brook 
just west of the “ head’ of the mountain. 

Microchaete tenera Thuret, not hitherto credited to this coun- 
tty, grows in various localities in Connecticut. Specimens collected 
hear Norwich and examined by Dr. Bornet, have cylindrical 
Spores in a series, exactly corresponding to those described for 
M. diplosiphon Gomont. This species much mixed with various 
Selatinous algae occurs upon dripping rocks near Norwich and 
New Haven, Conn. 


428 


Scytonema crispum (Ag.) B. & F. This species, known as 
Scytonema cincinnatum or Lyngbya cincinnata, although not credited 
to North America by Bornet and Flahault, is probably not un- 
common in the Eastern United States. Wolle had it from Penn- 
sylvania or New Jersey, as is evidenced by a specimen in the 
writer's possession. Mr. Isaac Holden has found it near Lanes- 
ville, Conn.; Mr. W. J. V. Osterhout finds it in abundance near 
Providence, R. I.; Mr. Wesley R. Coe, in Florida; Dr. J. E. 
Humphrey, in Jamaica, W. I., and the writer about New Haven, 
Conn. The filaments vary very much. Sometimes both branches 
and heterocysts are rare and the species looks very much like a 

Lyngbya, very often the scanty branches occur single and adjacent 
toa heterocyst and it resembles greatly a 7Zolypothrix, while the 
geminate branches midway between two heterocysts, characteristic 
of Scytonema are generally found only after long and careful search. 

Scytonema Javanicum (Kuetz) Bornet, one of the species with 
erect branches agglutinated into Symploca-like fascicles and pre- 
viously credited chiefly to the tropics (Java, Brazil and Guyana), 
but also found in France, according to Bornet and Flahault, reap- 
pears in this country in small quantity, growing on the trunks of 
trees in Middlesex Fells, Melrose, Mass. : 

Hassallia byssoidea Hassall is a very variable species and is 
credited to South Carolina only in this country by Bornet and 
Flahault. A form of it, not exactly typical, grows on rocks at the 
water’s edge along the Quinebaug River in Lisbon, Conn. It 
forms small tufts nearly black in color. It is characterized, a5 4 
form, by its regular, erect branches given off at an acute angle. 

Desmonema Wrangelii (Ag.) B. & F. is a curious genus of the 
Tribe Scytonemaceae, characterized by having several filaments 
inclosed in a common sheath and also by having basal heterocy Sts 
It is very abundant in Roaring Brook, a mountain rivulet 10 
Cheshire, near New Haven, Conn. It grows upon stones and 
rocks in the swiftest currents and forms slippery, bluish black ex- 
pansions. On being detached from the substratum it falls apart 
into small penicillate tufts. It is present during the month of 
June at least. 

Nostoc parmelioides Kuetz. is one of the most readily recog” 
nized species of the genus. The frond is erect and attached at 


429 


one point. It is generally strongly flattened and orbicular, with 
edges more or less strongly crenate. Within, the central filaments 
radiate from the attached portion, but the cortical filaments are 
convolute and twisted. It is the WV. cristarum of Bailey and the 
NV. alpinum ot Wood and Wolle. Bailey found it in New York, and 
Wood and Wolle in Pennsylvania. The writer has it from near 
Harrisburg, Penn., collected by Harry M. Kelley, and has found 
it in abundance in brooks at Mt. Carmel, Conn. 

Anabaena variabilis Kuetz., which may be distinguished from 
any other species of the genus occurring in this country by its 
oval, seriate spores remote from the heterocysts, occurred to the 
Writer in the autumn of 1894 in the brackish pond near Watch 
Hill, R. I., mentioned above. There was a great abundance of it, 
forming a dark greenish-brown, gelatinous layer on the floating 
leaves of Ruppia. The species is new to North America. 


Schizothrix lardacea (Ces.) Gomont. Of the twenty-seven 
Species of this homocysted genus given by Gomont in his mono- 
graph, only four are credited to North America north of Mexico. 
S. lardacea was found in some abundance on dripping vertical 
faces of trap at East Rock, New Haven, Conn. It formed masses 
of a dirty brown color. It has previously been found only in 
Germany, France and Italy. 

Sch. fr agilts (Kuetz.) Gomont was found at Brookfield, Conn., 
forming reddish crusts on stones kept moist by the spray from a 
Waterfall. The red color was due to a unicellular organism asso- 
ciated with it. Hitherto it has been credited only to Switzerland. 

flydrocoleum homoeotrichum Kuetz. The species of the genus 
ydrocoleum are distinguished by their caespitose habit and calyp- 
trate terminal cell. No one of the four fresh water species is cred- 
ited to North America. A. homocotrichum, however, was found 
by the writer growing in small short tufts on the posterior ends of 
shells of living fresh water mussels (Axodonta) in Trading Cove 
Brook, N orwich, Conn. 

Symploca muralis Kuetz., collected in New England by Farlow 
8Tows also on moist soil about the public pump in East Falmouth, 
Mass., and what appears to be the same species occurs abundantly 
On flower pots in greenhouses in New Haven, Conn. 

Lyngbya versicolor (Wartm.) Gomont, not credited to North 


430 


America in the “ Monograph,” has been found by Dr. Humphrey 
in Jamaica, W. I. (cf. Phyc. Bor. Am., No. 54), and by the writer 
at Salisbury, Conn., and in two localities near New Haven, Conn. 
(cf: ibid., No. 54). It grows attached at first, but later floats free. 

L. Lagerheimii (Moeb.) Gomont. A very slender species dis- 
tinguished by being fairly regularly twisted into a loose spiral, 
previously found only in Brazil, has reappeared in Southern New 
England. The species grows in brackish water in a small pool 
near Norwich, Conn., and in Little Pond, Falmouth, Mass. In 
each case it was growing upon Cladophora expansa and in one 
case was associated with Scenedesmus caudatus and various des- 
mids, while in the other it grew with Chara crinita and Glocotrichia 
Pisum. It also grows somewhere near New Haven, as consider- 
able quantities of it appeared in an aquarium in the Biological 
Laboratory of the Sheffield Scientific School. 

Phormidium Valderianum (Delp.) Gomont. This is another in- 
teresting species found in Watch Hill Pond, R. I. It grows in 
fresh water in Europe and forms expanded, lamellose strata on 
rocks and plants. In this locality, it forms a verdigris-green gela- 
tinous cylindrical covering to the elongated stems of Ruppia mart 
tama. It is common to find along the shore of the pond Ruppia 
stems partially dried and then the Phormidium layer is papery 19 
texture. 

Arthrospira Jenneri Stiz. is not credited to this country by 
Gomont. It is a large form and unmistakable. It has been found 
by Mr. Isaac Holden in some abundance near Schenectady, N.Y. 

ARTHROSPIRA GOMONTIANA n.sp._ For several years Mr. Isaac 
Holden and the writer have found in the vicinity of Bridgeport, 
Conn., scanty material of an Arthrospira which appeared to be 
different from any previously described. In June, 1895, howevet, 
a considerable quantity was found and sufficient material was Pre 
served for future distribution and study. 

This species makes its home in a pool of fresh water near 
Factory Pond in the northern part of Bridgeport, and floats ase 
the surface in verdigris-green patches of comparatively small size, 
resembling very much the so-called « Wasser bliite” of Anabaena 
flos-aquae, Gloiotrichia Pisum, etc. The floating masses are made 
up of small flocculent bits, each of which contains a considerable 


431 


number of the trichomes of the Arthrospira held together by a 
common jelly. 

The diameter of the trichome is from 2.5-3 4. It is regularly 
twisted into a rather lax spiral, the distance between the turns 
being 16-18 m, and the width of the turn about 6 yw. Caduceus 
forms are not uncommon. The cells of the trichome vary from 
4-5 » in length; the cell divisions in vigorous trichomes are in- 
distinct with few granules and with the cell contents light bluish 
green by transmitted light. In the majority of the cells large 
vacuoles are present. 

This form is decidedly smaller than either A. Jenneri or A. 
Platensis and with a laxer spiral. From A. mintaza it differs both 
in habitat and habit, and in the fact that the extremities are not 
at all capitate. As the species appears to be new, the writer de- 
Sires to dedicate it to M. Maurice Gomont, of Paris, as a token of 
his esteem and respect. 

The writer wishes also to express his deep obligation to Dr. 
Bornet and M. Gomont for assistance in determining the above- 
mentioned forms, as well as others, without which no certainty 
Would have been possible. 


Personal Nomenclature in the Myxomycetes. 
By O. F. Cook. 

As has been said, there are two ways of naming plants, ac- 
cording to the system of priority, or according to the personal 
System, which is no system at all. The alleged object of both 
methods is to secure uniformity of nomenclature, and opinion upon 
their relative merits depends on whether uniformity with the past 
is looked upon as more important than uniformity with the future. 
That the personal system does not always secure uniformity even 
With the immediate past is evidenced by two recent books, both 
Purporting to be monographs of the same group of organisms. 
They are based, practically, on the same collections and are pub- 
lished Only two years apart. Massee’s “ Monograph of the Myxogas- 
“res” recognizes 41 genera. Lister’s “ Mycetozoa”’ describes 38, but 
13 of these are different from those of Massee. Massee has 430 
Species, of which nearly one-third are not recognized as valid by 
Lister, while another third have their names changed, so that of 


432 


the 430 binomial specific names in Massee’s book just 160 are to 
be found in Lister’s. The student who would compare or make 
use of both works must spend much time in more or less fruitless 
attempts at cross-references, for the indexes are defective. 

No charge of adhesion to the laws of priority can be made 
effective against either Massee or Lister, any more than against 
their great predecessor Rostafinski. The indications are that all 
three have believed that variety in names would add a pleasant 
synonymical spice to the study, with the advantage of discourag- 
ing beginners not accustomed to much seasoning of this kind. 

Thus to judge from the past the personal system will not give 
us uniformity, for the recent works differ in nomenclature as much 
or more than the older. A most radical change, however, must 
follow any attempt at an application of the law of priority, for it 
will remove nearly all the names used for the last twenty years, Or 
since the publication of the works of DeBary and Rostafinski. 
Nearly all the Rostafinskian generic names must be replaced by 
others now disused for half a century or longer. If, however, 
these sweeping changes will bring us at once into uniformity with 
all the future, the wisdom of making them is evident—the soonef 
the better. Apparently, however, the application of the rules 
through which uniformity is expected for the higher plants does 
not give the same promise in the case of the Myxomycetes. A 
large number of old generic names exist, and their authors in- 
cluded under them forms now recognized as belonging to several 
different families, for the older writers had absolutely no idea of 
genera as we understand them. The classification in this group 
is by no means settled, and the principle that a genus is a nega 
tive conception merely, and consists of what may be left under a 
generic name after subsequent genera are withdrawn, will bring 
about in the Myxomycetes the result that with changes of view on 
the relative importance of characters, changes in nomenclature 
will be necessary. To illustrate by a simple case, the genus 


Physarum was described by Persoon (1 797) on four species, aS 
follows: 


Physarum columbinum=Lamproderma. 


“eé 


bivalve=Angioridium, 


“ . | —Tilmadoche, ween 


433 


Thus all the original species of Physarum have at one time or 
another been removed from the genus, and we are brought to the 
alternative of dropping the old name or of preserving it by slip- 
ping it around over the different generic groups, representatives 
of several of which were included under most of the old names. 
If we adopt Lister’s characterization of Fuligo we must abandon 
the genus Physarum in the sense in which recent authorities have 
used the name, for, as Lister himself states, the generic character 
of Fuligo depends on the presence or absence of a bell-jar over 
the ripening plasmodium. Physarum would then either be 
dropped or passed back to what is now called Lamproderma, a 
genus of a distinct family. What would then be the confusion if 
the next writer should find characters on which Fuligo, Tilma- 
doche and Angioridium might be held distinct ? 

That there will be immediate uniformity of view on the classi- 
fication of the Myxomycetes is not to be expected, and if we are 
to have an arbitrary system of rules by which the law of priority 
is to be applied in nomenclature it seems that one might be made 
which would at least fix a generic name to a certain natural group 
which varying views of classification could change only in size or 
Supposed importance. To follow a principle which may necessi- 
tate that a name be passed back and forth between different fami- 
lies or natural orders is to create confusion by law. To drop 
names based on species which represented new generic types in- 
volves a principle so manifestly unjust that continued acceptance 
of it is not to be expected. 

One method by which both these difficulties could be forever 
avoided is to agree that unless the author of a genus designates a 
Specific type in connection with his original description, the first 
Species referred to the genus may be looked upon as the genéric 
type, the genus to stand only if its type is found distinct from pre- 
Viously named genera. In the Myxomycetes this principle would 
Probably necessitate fewer changes than either of the others. 
Such a simple method of settling nomenclatorial problems may be 
thought to smack of empiricism. Its utility and practicality can- 
Not, however, be doubted, and it seems to be in general and grow- 
ing favor among zodlogists. The only serious objection to its 
8eneral application seems to be that a large number of names 


454 


would be changed. Whether the subsequent stability of generic 
names would not entirely overbalance this objection is a question 
worthy of careful consideration, for if botanists are to follow arbi- 
trary rules for the sake of uniformity in nomenclature, it is not 
probable that they will be satisfied until a system is framed which 
shall insure the uniformity. 


New Species of Fungi. 


By J. B. ELLIs AND B. M. EVERHART. 


I. SANDWICH ISLAND FUNGI. 


Collected by Mr. A. A. Heller, in the Island of Oahu, during 
the summer of 1895. 


MELIOLA Sanpicensis E. & E. 

Mycelium amphigenous, but mostly hypophyllous, forming 
suborbicular black patches 2-4 mm. diam. with a well defined 
outline. Mycelium abundant, with opposite spreading branches 
6-7 thick. Capitate hyphopodia abundant, opposite, small (12- 
15 X 6-7 p), short-stipitate, the upper cell ovoid or subglobose ; 
mucronate hyphopodia very scarce, ampulliform with a short, 
straight neck, found occasionally standing opposite a capitate hy- 
phopodium. Bristles abundant, both on the mycelium and sut- 
rounding the lower part of the perithecia, 300-500 yp long, 8-10 /4 
thick, generally with slight swellings at intervals, translucent- 
brown at first, finally almost opake, septate at intervals of about 
40 p, gradually attenuated above toa point. Perithecia scattered on 
the mycelium, subglobose, of coarse, cellular structure, papillose- 
roughened, ;50-200 yz diam. subostiolate, finally collapsing. As- 
ci clavate-elliptical, 50 X 25-30 yp, sessile, 2-3-4-( mostly 2- ) 
spored. Sporidia oblong-cylindiical, 4-septate and constricted at 
the septa, rounded at the ends, 33-40 X 12-15 p, brown. 


The bristles are so abundant that the patches of mycelium Te 
semble dense tufts of black hair, almost entirely hiding the pet 
ithecia. This character, with the greater length of bristles and 
the comparative absence of mucronate hyphopodia, with the ed. 
spored asci, distinguish this from M. malacotricha Speg., MV. amphi- 
tricha Fr., and the other species having similar sporidia. 


On leaves of some plant of the Family Rubiacez. No. 2369- 


455 


ASTERINA GLOBIFERA E. & E. 


Epiphyllous. Perithecia scattered, ovate, becoming more or 
less depressed and collapsed, small, 100-200 p diam., astomous, 
membranaceous, overrun and surrounded by an obscure mycelium, 
consisting of yellowish-hyaline prostrate, septate, branching 
threads and erect or ascending moniliform, brown threads. Peri- 
thecia at first filled with yellowish-brown, globose cells, 6-9 7 
diam, among which, later on appear the obovate or subglobose 
asci 20-30 x 15-20 y containing 8, elliptical hyaline spordia 15— 
20 < 8-10 » composed of two globose cells with a constriction 
between them. 

On Grevillia sp. No. 1946. 


SCIRRHIA LOPHODERMIOIDES E. & E. 

Stromata narrow-elliptical or linear, 44-2 mm., long, or by con- 
fluence 4-1 cm., long, raising and splitting the epidermis, black. 
Ascigerous cells globose, 150-200 p diam., lying in a single series. 
Ostiola inconspicuous. Asci oblong, sessile, 80-90% 20-22 p, 8- 
Spored. Sporidia crowded-biseriate, cylindrical, curved at the lower 
end (with a septum where the curve begins)? greenish-hyaline, 
35-40x7-8 uw. Differs from S. striaeformis, Niessl., in its longer, 
cylindrical sporidia. 

_ On dead culms of some grass. No. 2368. 


Puccinia Oanuensis E. & E. 

II. & III. Sori mostly hypophyllus, erumpent, naked, orange- 
yellow, pulverulent, subconfluent, not margined by epidermis. 
Uredospores obovate, orange-yellow, coarsely and strongly echinu- 
late, 22-30 X 20-22 p. ce 

Teleutospores mixed with the uredospores, obovate-elliptical, 
smooth, yellow-brown, slightly constricted, darker at the apex 
and slightly thickened, but without any distinct papilla, 27-40 
* 15-20 pv. Pedicles shorter than the spores. 

Remarkable for its almost permanently yellow sori, which 
only become a little darker after the leaves become dead and dry. 
P. Windsoriae Schw., has spores on longer pedicels, more dis- 
tinctly thickened above and scarcely constricted. 

On leaves of some unknown grass (resembling Panicum or 
ffolcus). No. 1976. 

Urepo verara E, & E. 

Sori amphigenous but mostly epi hyllous, pluvinate, scat- 
_ tered, Gate 141 mm. diam., Ae pen Te by a reddish- 
Purple discoloration, covered by the epidermis which is finally 
ruptured in the center or cracks across, but still remains covering 
_ Me sori. Spores elliptical, pale, with a thick epispore, faintly 
_ aculeolate, 22-30 «x 18-20 p. Paraphyses sparingly developed. 


436 


Differs from U. euphorbicola B. & C. and U. tordillensis Speg. in 
its larger spores and does not seem referable to any of the eu- 
phorbicolous species of Uromyces hitherto described. 

On Luphorbia cordata. No. 2027. 


PHYLLOSTICTA SCAEVOLA E. & E. 


Spots orbicular, light brown, 2-3 mm. diam. Perithecia 
hypophyllous, innate-superficial, black, 60-80 y diam., pierced 
above. Sporules clavate-ob‘ong, 2-3-nucleate, hyaline, 10-12 x 
214-3 pp. : 

The perithecia in the specimens examined were mostly sterile. 

On leaves of Scaevela Chamissoniana. No. 2124. 


HENDERSONIA NITIDA E. & E. 


Perithecia epiphyllows, scattered, innate-prominent, globose, 
90-110 y diam., black and shining, perforated above, about one- 
half buried in the substance of the leaf. Sporules oblong, I—-3- 
septate, not constricted, hyaline at first, then pale brown, 12-15 
x 3%-4% yp, on slender basidia shorter than the sporules. 
On living leaves of Myrsine sp. No. 2305. 


ASCHERSONIA MARGINATA E. & E. 


Stromata amphigenous, carnose, sessile, adnate, hemispher ical 
or strongly convex, mostly narrowly marginate, 2—3 mm. diam, 
nearly black outside (yellow when fresh)? light-yellow within and 
of sclerotoid structure. Perithecia (ascigerous cells) sunk in eid 
stroma in groups or clusters, of 4—8, minute (75—I10 yz) irregular in 
shape from mutual pressure. Ostiola slightly prominent, soon pet 
forated. Sporules fusoid, 2~-3-nucleate, hyaline, 5-7x1% acute 
at each end. Basidia slender, longer than the sporules. 

Differs from A Tahitenis Mont., and A turbinata Berk., in its 
clustered perithecia and smaller sporules. 


On living leaves of Psidium No. 1945. 


II. FLORIDA FUNGI. 


Collected by Mr. Geo. V. Nash, in Florida, during the sum 
mer of 1895. 


ASTERIDIUM DOTHIDEOIDEs E. & E. 


Perithecia epiphyllous, scattered or loosely grouped, and often 
3-4 confluent, flattened-convex, 300 » diam., or elongated an 
hhysteriiform, 500 X 200 y, fringed around the base with a pros 
trate radiating sparingly anastomosing mycelium, of brown sub- 
continuous threads about 5 y thick with obovate or globose one” 
celled hyphopodia about 7 » diam. at distant intervals. As¢l 


437 


obovate, sessile, 30-40 y 18-21 yp. Sporidia crowded, oblong- 
elliptical, about 20 x 7-8 », often slightly curved, obtuse, yellow- 
ish-hyaline and mostly uniseptate at first, finally brown and bisep- 
tate, the septa hyaline. 

Very different from A. depidigena E. & M. on the same host. 


On leaves of Andromeda Jerruginea. No. 1939. 


SEPTORIA QUERCICOLA Sacc. Mich., 1: 174; Syll. 3: $05. 

_ Spots orbicular, 1-2 mm. diam., dirty-white with a dark mar- 
gin, or ferruginous, larger and irregular. Perithecia amphigenous 

ut more distinct above; on the white spots, mostly only one in 
the center of the spot; on the larger ferruginous spots, more 
Numerous and _ scattered. Sporules cylindrical, curved,, sub- 
attenuate at the ends, 3-septate, 25-40 X 3, w. Var. cinerceaE. & 

+» On leaves of Quercus cinerea, differs only in its longer (40~- 
60), 3-6-septate sporidia. 

On Quercus sp. No. 2091. 


Dartuca arcuata E. & E. 

Perithecia globose, 75-100 » diam. Sporules arcuate-fusoid, 
hyaline, 3-septate, not constricted, 14-162 Ya-3 pe. 

Distinguished from the other species of this genus by its 
arcuate sporules. 

Parasitic on Uredo, on leaves of Andropogon? No. 1837. 


CoLterorricnum CoMMELINAE. 
Spots dark brown, suborbicular, 2-3 mm. diam., sub-indefi- 
Nite, paler and more obscure below. Acervuliabout 100 # diam., 
tinged with dark brown, obscurely septate hairs 4-5 p thick 
below and tapering above, 60-75 » long. Conidia oblong, hyaline 
Continuous, 12-164-5 p, a little narrower at one end. 


On leaves of Commelina angustifolia. No. 1798. 


Cotterorricnum Eryturinak E. & E. 
_ Spots orbicular, rust-color, with the margin darker, y% cm. 
lam. Acervuli epiphyllous, erumpent-superficial, 150-200 4, 
diam. loosely clothed with simple, brown, sparingly septate hairs 
70-100 X 3-4 p. Conidia oblong—cylindrical, hyaline, continuous, 
14-16 X 3%, fhe 
Differs from C. Commelinae in its larger spots and narrower 
Sporules, 
On leaves of Exythrina herbacea. No. 2123. 


Cotterorricuum Azae& E. & E. ab cette 
Spots dark rusty-brown, orbicular, 3-8 mm. diam., definite, wi 


438 


the margin concolorous. Acervuli mostly epiphyllous, erumpent, 
120-” diam., sparingly clothed with slender, black, slightly 
curved, septate bristles 50-70 X 3-3%p. Conidia oblong, 8-13 
X3-3% yp 
Sporules shorter than in either of the two preceding species. 
On leaves of Asalea viscosa. No. 1991. 


Crrcospora CAssaAvaE E, & E. 

Spots orbicular, light-rusty brown, 3-5 mm. diam. definite, with 
a narrow, darker margin. Hyphae cespitose, continuous, slightly 
toothed above, yellowish-hyaline, 30-40 x 4. Conidia cylindri- 
cal or clavate-cylindrical, 3-5-septate, hyaline, 20-50 y% 5-6 p. 
Amphigenous. 

On leaves of Cassava. No. 1950. 


CERCOSPORA CHRYSOBALANI E. & E. 

Spots deep red-brown, definite, mostly irregular in shape, 1-6 
mm. diam. Hyphae amphigenous; cespitose, brownish, continu- 
ous, slightly narrowed and subundulate above, 20-30x31%2-4 /- 
Conidia, slightly narrowed above, subhyaline, 3-8-septate, 30-70" 
X3 p- 

On leaves of Chrysobalanus oblongifolius. Nos. 1949 and 
1793. : 


Gercospora GALACTIAE E, & E. 

Amphigenous. Spotsnone. Hyphae densely tufted, 100-110" 
<6 p, septate, dark-brown, subgeniculate, and subundulate above; 
the tufts of hyphae forming loose floccose patches 4-114 mm. 
across, scattered over the entire surface of the leaf. Conidia ob- 
clavate cylindrical, slightly colored, 2-8-septate, 30-70 ~ 5-6 p. 

Very distinct from C. flagellifera Atk., on Galactia pilosa. 

On Galactia Nuttallii. No. 1987. 

IsARIOPSIS PENICILLATA E, & E. 

__ Stipe about 1% mm. high, composed of simple, pale yellow 
threads closely compacted at base and 10-15 yp thick, relaxed 
above and spreading in a brush-like manner, the fusoid yellowish- 
hyaline 22-25 5 y (obscurely 1~3-septate)? conidia term'n 
on the fibers composing the stipe. ; 

Parasitic on mycelium of Meliola on leaves of Gordonia lasian- 
thus. No. 1956. 


Urebo ScHOENOCAULI E. & E. 


Sori erumpent, mostly elliptical, %4-34X\% mm., at first COV- 
ered by the epidermis, soon exposed, yellow. Spores globose 


439 


Is p diam., oftener obovate or elliptical, 19-22 14-16 p, slightly 
aculeate. 


On Schoenocaulon gracilis, No. 2015. 


Ill. MEXICAN FUNGI. 


Collected in the vicinity of Monterey, Mexico, by Dr. B. F. G. 
Egeling. 


SCHIZOPHYLLUM EGELINGIANUM FE. & E. 


Pilei_imbricated, reniform, margin entire or lobed, 1% cm. 
long, 2 cm. wide, white-tomentose, the margin cleft, white-pilose, 
depressed behind around the short lateral stem, zoneless; flesh 
about I mm. thick, white, carnose-coriaceous. Lamellae pale 
flesh-color, unequal, with a row of very short ones around the 
Margin, narrow. Spores colorless, globose-elliptical, about 
3 X 21% yw, borne on clavate-cylindrical basidia, 18-20 X 3% yp. 

The pilei are borne on a white, subcoriaceous subiculum 
€rumpent through the epidermis in patches 3-4 mm. across, often 
becoming confluent for 1-8 cm. in extent and nearly surrounding 
the stem. 


On dead, hollow stem of Magnolia Mexicana, June, 1895. 


Roseciinta pourosa E. & E. : 
Perithecia hemispheric-prominent, 134-2 mm, diam., covered 
y a thin, ashen-gray coating, which also overspreads the surface 
of the wood, and may be only accidental. Ostiolum papilliform. 
Asci_ cylindrical, 150-200 X 13-16 yw, (paraphysate) ?, 8-spored. 
Sporidia broad oblong-fusoid, brown, finally nearly opake, but 
When young the extremities are paler, 25-35 12-15 pv. 
On dead wood, June, 1895. 


Dipymospuarrra SPHAEROPHORA E. & E. 
: Perithecia scattered, buried, 1 mm. diam., finally more or less 
distinctly collapsing, leaving the erumpent apex and broad papilli 
°rm ostiolum prominent in the center of the depression. Asci 
Cylindrical, stipitate, paraphysate, 8-spored, p. sp. 100-1I10X 10— 
12 Sporidia uniseriate, oblong, uniseptate, dark brown, con- 
_ Stricted, more or less distinctly roughened, 16-20X6-8 yz. with a 
_ “epressed-globose, yellowish-hyaline appendage at each end. 
Differs from D. appendiculosa Speg. in its larger perithecia, 
Smaller asci and sporidia, and the different character of the append- 


ages, 


On Agave sp. 


440 


PHYLLACHORA? YuccaE E. & E. 

Stromata gregarious, elliptical or oblong, %-1%4  % mm., sub- 
seriate, sunk in the substance of the leaf, and covered above by 
the blackened epidermis. Ascigerous cells 80-90 » diam., finally 
confluent. Ascioblong—cylindrical, narrowed above, aparaphysate, 
50-60 X 7-8 yw, filled with granular matter and oil-globules. 
Sporidia not yet formed. 


On Yucca angustifolia. June, 1895. 


HAPLOSPORELLA MexIcAna E. & E. 


Stromata numerous, purplish-black, erumpent, subseriate, often 
confluent for I or more cm. and 2—3 mm. broad, entirely covered 
at first, soon visible through cracks in the ruptured epidermis, of 
soft, carnose texture, flat-pulvinate, the mycelium blackening the 
inner substance of the stem. Perithecia (ascigerous cells)? closely 
packed, 190-230 yp. diam., becoming subconfluent. Sporules obo- 
vate at first, and hyaline, finally brown and mostly elliptical, 15- 
2211-13 y., on stout basidia mostly shorter than the sporules. 

On account of the carnose stroma this should perhaps be 
referred to the genus Aschersonia in the Fam. Nectrioideae, but 
the sporules and the dark color would make it Hap/osporella. 

On dead stem of Magnolia Mexicana. June, 1895. 
MeEvocraAmMa Ecetinoeu E. & E. 


_ Stroma subcuticular, 1-2 mm. diam., ovate, the apex erum- 
pent, substance white, granular and finally crumbling. Perithecia 
(14-20), peripherical, ovate, 200 w high, 150 yw broad, black, im- 
mersed in the surface of the stroma which finally crumbles away, 
leaving them partially free, narrowed above into an acute ostlo- 
lum, which is finally porforated and obtuse. Asci clavate-cylin- 
drical, short-stipitate, paraphysate, p. sp. 75-85 12 Sporidia 
biseriate, cylindrical, obtusely rounded at the ends, 5-septate 
and more or less constricted at the septa, one cell near the mid- 
dle generally swollen, yellow-brown, becoming opake, 20- 
23 X 6 ps. 


On dead leaves of Agave. May, 1893. 


Proceedings of the Club. 


TuEsDAY EvENING, OcToBer 8TH, 1895. 


The President in the chair and 27 persons present. 
Prof. Arthur M, Edwards, M. D., Mrs. C. Rice and Mr. Charles 
Ericson were elected active members. 


441 


A communication was received from the Secretary of the Coun- 
cil of the Scientific Alliance, announcing the opinion of the Coun- 
cil that it would be advantageous for the secretaries of the socie- 
ties forming the Alliance to transmit abstracts of the proceedings 
for publication in Science. Upon motion of Dr. Allen, it was 
unanimously resolved that the Club concur in the above opinion. 

Dr. ‘I’. F. Allen reported a visit to the islands lying near Lands 
End, and spoke of the peculiarly equable climate, and its effects 
upon the flora and productions. He had found Hooker's British 
Flora an unsatisfactory book for field work. 

Mr. Lighthipe spoke of his observations in the pine barrens, 
especially with reference to Schizaea and Chrysopsts. 

Dr. Small exhibited a map upon the blackboard, indicating his 
travels in Georgia in securing a forestry exhibit for the Atlanta 
€xposition. He also briefly described the form of the exhibit, 
which was in duplicate. He promised a farther report at a subse- 
quent meeting. 

Dr. Britton announced collecting Panicum colonum L., a tropi- 
cal species related to P. Crus-galli, in Virginia, and the discovery of 
a large patch of P. verrucosum on Staten Island by Mr. Tyler. He 
also announced that the organization and actual commencement 
of work on the New York Botanical Garden had taken piace. 

Brief remarks were also made by Miss Ingersoll and Messrs. 
Tyler, Van Brunt and Van Sickle. Mr. Van Sickle referred par- 
ticularly to the occurrence of the Russian Thistle in northern 
Jersey and of Azolla Caroliniana at Passaic. 

The President reported upon his observations upon the western 
slope of the Catskills, a few miles from the locality of the old 
Mountain House, speaking particularly of a white flowered form 
of Impatiens aurea. He brought a number of sets of some of his 
More interesting collections which were distributed among the 
members. 


442 


Index to recent Literature relating to American Botany.” 


Andrews, F. M. Development of the Embryo-sac of /effersonta 
diphylla, Bot. Gaz. 20: 423-425. p/. 28. 25 S. 1895. 

Arthur, J. C. Development of Vegetable Physiology. Bot. Gaz. 20: 

. 381-402. 25S. 1895. 

Aubert, A. B. Liste partielle des Diatomées d’Orono, Maine, U. S. A. 
Le Diatomiste, 2: 150-151. Mr. 1895. 


Aubert, A. B. Liste partielle des Diatomées de Somesville, Seal Har- 
bor, et Northeast Harbor, Etat du Maine, U.S. A. Le Diatomiste, 
2: 140-143. D. 1894. 

Baker, S.G. Prochynanthes Bulliana, Curt. Bot. Mag. 51: p/. 7427- 
Au. 1895. 


Native of Mexico. 


Barnhart, J. H. Onthetwo Editions of Emory’s Report, 1848. Bull. 
Torr. Bot. Club, 22: 394, 395. 30S. 1895. 


Bastin, E. S. Structure of our Cherry Barks. Am. Journ. Pharm. 
G7: 435-452: f. r—74. S. 1895. 

Bastin, E.S. Structure of our Hemlock Barks. Am. Journ. Pharm. 
67: 356-362. f. 7-5. Jl. 1895. 

Bicknell, E. P. The Genus Sanicu/a in the eastern United States, 
with Descriptions of two new Species. Bull. Torr. Bot. Club, 22: 
351-361. pl. 247-245. 31 Au. 1895. 

Blasdale, W.C. Observations on Puccinia mirabilissima. Erythea, 
3: 131-135. pl. z. 3S. 1895. 

Brotherus, V. F. Beitrige zur Kenntniss der brasilianischen Moos- 
flora. Hedwigia, 34: 117-131. 19 Je. 1895. 

Describes new species in Ephemerum, Systegium, Trematoden, Leucobryum, Oc- 


toblepharum, Fissidens, Syrrhopodon, Calymperes, Pottia, Hyophila, Bar bula, 


Macromitrium, Tayloria, Physcomitrium, Bryum, Garovaglia, Sigmatella and 
Sphagnum. 


oe es 
* At the Springfield meeting of the American Association for the Advancement - 
Science, the Section of Botany adopted the following limitations of the contents of this 
Bibliography : i 
1, That all bacteriological, horticultural and agricultural titles be omitted, but 
any case of doubt the title is to be included. 
2. That all references to exsiccatae be excluded. 
3. That all references to reviews be excluded. 
The Bibliography Committee and the Editors of the BULLETIN would renew their 
request that omissions from this record be communicated. 


443 


Burrill, T. J. Rhws Poisoning. Gard. & For. 8: 368. 11S. 1895. 


Bush, B. F. Quercus Phellos X rubra in Missouri. Gard. & For. 
8: 375. 18S. 1896. 


Cheney, L. S. Sphagna of the Upper Wisconsin Valley. Trans. 
Wisc. Acad. Sci. Arts & Lett. 10: 66-68. 1894-95. 


Cheney, L. S. Parasitic Fungi of the Wisconsin Valley. Trans. 
Wisc. Acad. Sci. Arts & Lett. 10: 69. 1894—95. 


Cheney, L. S. Hepaticae of the Wisconsin Valley. Trans. Wisc. 
_ Acad. Sci. Arts & Lett. 10: 70-72. 1894-95. 


Cleve, P. T. Les Diatomées de l’Equateur. Le Diatomiste, 2: 99- 
103. Je. 1894. 

Cleve, P. T. Sur quelques Especes nouvelles on peu connues. Le 
Diatomiste, 2: 143-147. D. 1894. 

Descriptions of several new American diatoms. 

Comes, O. Sulla sistemazione botanica delle specie e delle razze del 
genere Wicotiana. Atti Ist. Incor. Napoii (IV.) 8: [reprint, pp. 
32]. 1895. 

Coville, F. V. Dr. Robinson and Homonyms. Bot. Gaz. 20: 320- 
322. 15 Jl. 1895. 

Coville, F. V. The Nomenclature Question ; theoretical objections to 
a stable nomenclature. Bot. Gaz. 20: 428, 429. 25S. 1895. 

Davis, W. T. Botanical Notes. Proc. Nat. Sci. Assoc. Staten Island, 
4: 83. 148. 1895. 

Davy, J. B. Transcripts of some Descriptions of Californian Genera 
and Species.—V. Erythea, 3: 136-138. 3S. 1895. 

Deane, W. Notes from my Herbarium.—III. Bot. Gaz. 20: 345- 
348. fig. 15 Au. 1895. 

Dewey, L. H. Laphamia ciliata sp. nov. Bot. Gaz. 20: 425. 25 
S. 1895. 

Dewey, L. H. Tumble Mustard. Bull. Torr. Bot. Club, 22: 370. 
31 Au. 1895. 


Note on the distribution of Sisymébrium altissimum. 

Ellis, J. B., and Everhart, B. M. New Fungi, mostly Uredineae 
and Ustilagineae from various Localities, and a new /omes from 
Alaska. Bull. Torr. Bot. Club, 22: 362-364. 31 Au. 1895. : 
Describes new species in /omes, Ustilago, Sorosporium, Puccinia, Ravenelia, 


Doassansia, Aecidium and Peronospora. 


444 


Engelhardt,.H. .Uber neue Tertiairpflanzen Stid-Amerikas. Abh. 
Senckenb. Naturforsch. Gesellsch. 19: 1-47. p/. 1-9. 1895. 


Ganong, W.F. Botanical Nomenclature and Non-systematists. Bot. 
Gaz. 20: 317-320. 15 Jl. 1895. 


Gerard, W. R. Origin of the name Saméucus. Gard. & For. 8: 
368. ..11S, 1895. 


Grevillius, A. S. Ueber Mykorrhizen bei den Gattung Botrychium, 
nebest einigen Bemerkungen ueber das auftreten von Wurzelsprossen 
bei B. Virginianum Sw. Flora, 80: 445-453. 1895. 


Harvey, F. L. Contribution to the Characeous Plants of Maine. 
Bull. Torr. Bot. Club, 22: 397, 398. 30S. 1895. 


Hieronymus, G. Bemerkungen iiber einige Arten der Gattung S/go- 
nema. Hedwigia, 34: 154-172. 19 Je. 1895. 
New species from South America and notes and redescriptions of North American 

species. 

Hollick, A. Identification of fossil Leaves. Bot. Gaz. 20: 332- 15 
Jl. 1895. 


Hooker, J. D. Helianthus debilis. Curt. Bot. Mag. 51: p/. 7432+ 
S. 1895. 


Hooker, J.D. Rumex hymenosepalus. Curt. Bot. Mag. 51: f/. 7433- 
S. 1895. 

Jenman, G. 8. Polypodium (Phegopteris) Trinidadensis. Gard. 
Chron. 18: 235. 31 Au. 1895. 


A new species from Trinidad. 

- Knowlton, F. H. Description of a new problematical Plant from the 
Lower Cretaceous of Arkansas. Bull. Torr. Bot. Club, 22: 387-399: 
Je F-3. 30S. 1895. 

Lloyd, F. E. Teratological Notes. Bull. Torr. Bot. Club, 22: 399 
397. pl. 247. 30S. 1895. 

Lindau, G. Acanthaceae Americanae. Bull. Herb. Boiss. 3: 361- 
3723 479-493. Au. S. 1895. | 
Descriptions of many new species from South and Central America. 

Lodeman, E.G. Poisoning from Rhus. Gard. & For. 8: 398- ? 
O. 1895. 

Lueders, H. F. The Vegetation of the Town Prairie du Sac. Trans. 
Wisc. Acad. Sci. Arts & Lett. 10: 510-524. f/. 77. 1894-95- 


Macloskie, G. Antidromy of Plants. Bull. Torr. Bot. Club, 22: 
279-387. 30S. 1895. 


445 


MacDougal, D. T. Irritability and Movement in Plants. Pop. Sci. 
Month. 1895: [reprint pp. 10]. Je. 1895. 

McClatchie, A. J. Flora of Pasadena and Vicinity. Reid’s History 
of Pasadena (Cal. ) 605-649. 1895. Reprint. 

A list of 1056 species and varieties, from Myxomycetes to Compositae, 

Meehan, T. TZvrichostema dichotomum. Meehan’s Month. § 2. 261. 
Pl.9. S. 1895. 

Meehan, T. Arethusa bulbosa. Meehan’s Month. ge. 1 oe oe 
Au. 1895. 

Micheli, M. Contributions a la Flore du Paraguay. Pipéracées par 
C. de Candolle. Mem. Soc. Phys. et d’Hist. Nat. de Genéve, 32. 
no. 2, 1-10. pl. 52-54. 1894-95. 

New species and varieties described and figured are Peperomia psilostachya, Piper 
cinerascens, P. amplecteus aromaticum, P. fulvescens C. DC. from Morong and 
Britton’s Enumeration, p. 214, is also figured. 

Millspaugh, C. F. Decapitalization. Bot. Gaz. 20: 429. 25 S. 
1895. 

Mobius, M. Ueber einige brasilianische Algen. Hedwigia, 34: 173- 
176. 19 Je. 1895. 177-180. pl. 2. 1 Au. 1895. 

Describes and figures Mougeotia Uleana. 

Muller, J. Lichenes exotici.—IV. Hedwigia, 34: 139-145. 19 Je. 
1895. 

Describes several species from both North and South America. 

Muller, J. Lichenes Ernstiani a cl. Prof. Dr. Ernst prope Caracas 
lecti quos enumerat. Hedwigia, 34: 146-153. 19 Je. 1895. 

Olive, E. W. Observations upon some Oklahoma Plants. Bull. Torr. 
Bot. Club, 22: 390-394. 30S. 1895. 

Pettit, A.S. Arachis hypogaea L. Mem. Torr. Bot. Club, 4! 275~ 
296. p/. 83-85. 5 Je. 1895. 

Philippi, R. A. Plantas nuevas Chilenas. Ann. Univ. Chil. 87-90. 
1895. 

Descriptions of many new species of Sympetalae. 

Robinson, B. L., and Greenman, J. M. On the Flora of the Gala- 
pagos Islands as shown by the Collections of Dr. G. Baur. Am. 
Journ. Sci. (III.) 50: 135-149. 1 Au. 1895. 

Robinson, B. L., and Greenman, J. M. New and noteworthy 
Plants, chiefly from Oaxaca, collected by Messrs. C. G. Pringle, L. 
C. Smith and E. W. Nelson. Am. Journ. Sci. (III.) 50: 150-168. 
1 Au. 1895. 


446 


Robinson, B. L., and Greenman, J. M. A synoptic Revision of the 
Genus Zamourouxia. Am. Journ. Sci. (III) 50: 169-174. 1 Au. 
1895. 


Robinson, B. L., and Greenman, J. M. Miscellaneous new Species. 
Am. Journ. Sci. (III.) 50: 175, 176. 1 Au. 1895. 


Unona Panamensis, U. bibracteata, Malvaviscus Pringlei E. G. Baker and Lap 
hamia Toumeyi. 


Rodrigues, J. B. Hortus Fluminensis on Breve Noticia sobre as 
Plantas Cultivadas no Jardim Botanico do Rio de Janeiro. 4to. pp. 
307. pl. I-23. Rio de Janeiro. 1895. 

Rothrock, J. T. Our Pennsylvania Forests. Journ. Frankl. Inst. 140: 
105-117. Au. 1895. 

Rothrock, J. T. The Sugar Maple. Forest Leaves, 5: 56-58. Au. 
1895. , 

Rothrock, J. T. The Locust Tree. Forest Leaves, 5: 72,73. O- 
1895. 

Rowlee, W. W. The Aeration of Organs and Tissues in AZihania 
and other Phanerogams. Proc. Am. Micros. Soc. 15: 143-166. A/. 
I-6, Mr. 1894. 

Rusby, H. H. An Enumeration of the Plants collected in Bolivia by 
Miguel Bang, with descriptions of new genera and species. Part 2. 
Mem. Torr. Bot. Club, 4: 203-274. 27 Ap. 1895. 

Describes 97 new species of Spermatophytes. 

Rydberg, P. A. Flora of the Sand Hills of Nebraska. Contr. Nat. 
Herb. 3: 133-203. pd. z. 14S. 1895. 

Annotated catalogue of species. Carduus Plattensis is described as new and 


figured, 


Sargent, C.S.  Litsia geniculata. Gard. & For. 8: 374./- 52: 18 
S. 1895. 


Sargent, C.S. Cladrastis. Gard. & For. 8: 372. 18S. 1895. 


Sargent, C. S., Editor. Agave Utahensis. Gard. & For. 8: 384- 
S53: 25 S. 1895. 


Saunders, C. F. Scolopendrium Scolopendrium. innaean Fern 
Bull. 12: 1, 2. .O. 1895. 


Schumann, K. Redutia minuscula. Monats. Kakteenk. 5: 102- ji. 
1895. 


Setchell, W. A. Daniel Cady Eaton, 1834-1895. Bull. Torr. Bot. 
Club, 22: 341-351. 31 Au. 1895 (with portrait). 


Surplas Books for Sale from the Library of Dr. M. ¢. Cooke, 


146 Junction Road, London N., England. 


Balfour, J. B. Botany of Socotra. 4°. 100 plates. Edinburgh. 

Hedwig. Theoria generationis and Hist, Nat. Muscorum. 4°. 

Lamarck and DeCandolle. Flore Francaise. 6 vols., 8°. 

Steudel. Nomenclator Botanicus. 8°, board. 

Linnaeus. Flora Suecica. First Edition. 

Leers. Flora Herbornensis. 16 pl., 8°. 

Acharius. Synopsis Lichenum. 

Marsilius. Dissert: de generatione Fungorum. 

Vaillant. Botanicon Parisiense. Folio. 

Queckett. Histological Catalogue. 4°. 2 vols., cloth. 

Luerssen. Medico-Pharmaceutik Botanik. Vol. 1, Kryptogamen, 8°, paper. 
Lindley, Dr. Flora Medica. 1 vol., cloth. : 
Waring, Dr. Pharmacopaeia of [ndia. 8°, cloth. 

Fluckiger, Dr., and Hanbury, D. Pharmacographia. 8°. 

Hanbury, D. Notes on Chinese Materia Medica. 

Hanbury, D. Science Papers. 1 vol., 8°. 

Waring, Dr. Bibliotheca Therapeutica. 2 vols., 8°. 

Christison, Dr. On Poisons. 8°, 1 vol. 

Allen and Thomson. Expedition to the Niger. 8°, 2 vols. 

Farlow, W. G. Marine Algae of New England. 8°, paper, 

Toni and Levi. Flora Algologica Venezia. Parts I. and ik; 

Underwood. Catalogue of N. Am. Hepaticae. 8°. 

Tuckerman. Synopsis of Lichens of New England. 
Bescherelle. Prodromus Bryologiae Mexicanae. 8°. FS a 

Harvey, Dr. Phycologia Britannica. 4 vols. in 2 half morocco, gilt tops. Splen 


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Sowerby’s English Botany, 2d Edition, Vol. XII., only containing Algae. 8° 
Queckett, J. Lectures on Histology. 2 vols., 8°, cloth. 

Queckett, J. Treatise on the Microscope. 8°, cloth. 

Ardissone. Enum, della Alghe della Marca di Ancona. 4° 

De Notaris. Epilogo della Briologia Italica. Roy. 8°, half calf. 

Wahlenberg. Flora Suecica, 2 vols. 8°. 

Bauhin. Theatrum Botanicum. Folio. 

Meyer. Primitiae Florae Essequeboensis. 4°, calf. 

Sullivant. Musci and Hepaticae of U.S. 8°. 

Roth, A. G. Catalecta Botanica. 3 vols., 8°, half calf. 1806. 

Schrader, H. Spicilegium Florae Germanicae. 8%. 1794. 

Haller, A. Flora Jenensis. 12°. 1745. 

Weigel. Flora Pomerano Rugica. 12°. 1769. 

Olhaff. Elenchus Plantarum. 12°. 1656. 

Sprengel, C. Florae Halensis. 8°. 1806and 2 suppl. 

Schlechtendal. Flora Berolinensis. Part 2, Cryptogamia. 1 824. 

Schrank. Florae Salisburgensis. 12. 1792. 
Martius. Flora Erlangensis Crypt. 8, 6 pl., 1817. 
Nuovo Giornale Botanico Italiano, 1872 to 1891. 


n 2 vols., half calf (all published). 
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Michelia. Ed. P. A. Saccardo, 1879 to 18827 
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Flora (Regensburg), vols. 34 to 47. 14 vols., cloth. 


Notarisia, Commentarium Phycologiae, 1886-1891. Parts 1 to 25 (wanting part 


La Nuova Notarisia. 1890-1891. 7 parts. 

Revue Mycologique (Roumeguere). 1888-1891. 13 vols. in 6, cloth. 

Rabenhorst’s Algae Europaea Aquae dulces. 3 parts in 2 vols., half calf, com- 
plete, 1864. 

Monthly Microscopical Journal. (Ed. Dr. Lawson.) 18 vols., cloth. 1869-1877. 

Grevillea (Ed. M. C. Cooke). Complete set in parts, 20 vols, 1872-1892. 
Complete sets out of print, 

Hassalls, Dr. British Fresh Water Algae. 2 vols., cloth. 

Smith, W. British Diatomaceae. 2 vols., cloth. 

Ralfs, J. British Desmidiae. 1 vol., half calf. 

Bornet et Thuret. Notes Algologiques, part 2 only, folio, plates. 

Flora of Herefordshire. Mosses by Aug. Ley. Fungi by M. C. Cooke. 1 vol., 
8°, cloth. 1889. 

Hooker, W. J.,and Baker, J.G. Synopsis Filicum. Colored plates, 8°, cloth. 
1868. 

DeBary, A. Morphologie und Biologie der Pilze. Mycetozoen und Bacterien. 
8°, uncut. 1884. 

Natural History Review. Vols. I. to VI., 1854 to 1859 and 1861 to 1865, 11 
vols, (wanting Vol. VII.). 

American Naturalist, 1867 to 1886 (wanting 1876), in 17 vols., half calf, 2 vols 
in parts. Also 1887-8 complete; 1889 (wanting 266, 270, 274, 275); 1890 complete; 
1891 (wanting 298, 300). 

American Entomologist, and American Botanist and Entomologist. 1868-1870. 
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Ray (John). Historia Plantarum. 3 vols., folio. ; 

Mitten, W. Musci Indiae Orientalis. 8°, with Griffith’s Muscologia Itineris As- 
ao Griffith’s Notulae ad plantas Asiaticas, part 2. Cryptogamia in 1 vol., 8° 

calf, 

Buxbaum J.C. Plantarum minus cognitarum. Complete in 2 vols, 4°. 

Girod Chautraus. Recherches chimiques et micro. sur les Conferves. Fol. 36 pl. 
half calf. 1802. : 

Gerarde’s Herbal. Old folio, wanting title and last page of index, otherwise 
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Lonpon, July, 1895. 


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VoLuME I. 
A List of Plants Collected by Miss Mary B. Croft at San Diego, Texas. By 
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New or Noteworthy North American Phanerogams. By N. L. Britton 
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A List of Plants Collected by Dr. E. A. Mearns at Fort Verde and in the 
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The General Floral Characters of the San Francisco and Mogollon Mountains 
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Preliminary Notes on the North American Species of the Genus 7issa, 
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Contribution to American Bryology—XI. 


By EvizAsetH G, Britton. 
I, COSCIN ODON RAUI AND COSCINODON RENAULDI. 
(PLATE 248.) 


Owing to the fact that the upper part of the leaf in Coscinodon 
is Colorless, it has happened that in the first descriptions of both 
C. Wrightii and C. Raut, the vein has been described as ending 
below the apex. In describing C. Wrightii Sullivant said of the 
leaf that it was « costate half way’ (Mosses U. S. 38. p/. 4. 1856). 
In the Icones (71. pl. 45. 1864), he corrected this mistake, figuring 
the vein extending into and forming the awn. 

Austin made the same mistake about C. Raui (Bull. Torr. Bot. 
_ Club, 6: 46. 1875), describing the leaf with the vein ending below 
the apex, « costa valida sub apice finienti,” yet in the type speci- 
; mens preserved in his herbarium at Columbia College, the vein is 
Clearly €xcurrent, forming a terete, white awn, most clearly seen in 

Old abraded leaves when the blade is torn away. He recognized its 
_ true character in C Wrightit, for he says of it “costa extending into 
the much longer, more terete and more scabrous hair point;’’ yet 
he failed to see it in his own species. This is the more remarkable, 
4s he had in his herbarium an original sketch of the leaves by E. 
A. Rau, on the margin of which Mr. Rau had written, “ Leaves 
throughout chlorophyllose, except the long excurrent costa or tip.” 
In the Manual (L. & J. Mosses of N. Am. 155, 1884) this mis- 


448 


take is repeated and in describing C. Renauldi (Bot. Gaz. 15; 41. 
1890) M. Cardot cites this as one of the points of difference 
between these two species, the other being a difference in the teeth. 
These are described in the Manual as “ entire, split or merely per- 
forated here and there on the line of division, erect when moist, 
open when dry.” This isa translation from Austin’s original Latin 
description, but it does not agree with all his specimens. The 
teeth are very fragile in his species, and are seldom seen entire 
and unbfoken, except on a freshly opened capsule. They are long 
and slender, perforate at base, three to four parted above, and 
densely papillose; later they divide into slender divisions, and on 
old capsules have disappeared altogether, leaving only the annulus. 
As compared with the figures of C. Renauldi, they differ only in 
_ not being truncate when perfect. From C. Wrightii, as figured in 
the Icones, these teeth are much longer, more acuminate, the 
divisions narrower, more slender and regular, the perforations 
almost continuous above, and splitting into three to four slender 
divisions to each tooth. The annulus also is not like the mosaic 
of flat cells figured by Sullivant, there are only two rows of cells 
instead of three, and the last row has elongated vesicular cells. In 
the description of C. Renauldi, the annulus is not mentioned, nor 
is it figured in the plate. We have not seen a capsule of this 
species, but in a recent letter M. Cardot states that the annulus is 
present, but that he has not been able to determine whether it is 
persistent or fugacious as all the capsules which he has examined 
were too young. In the plate the teeth are figured as truncate. 
M. Cardot says that they were taken from freshly opened capsules, 
but we are of the opinion that if he were to search in the apex of 
the lid he would find the remnants of those teeth, as we have 
found in C. Rawi, that the tips often fall with the lid, and it was 
with difficulty that we secured a mount sufficiently perfect for our 
drawing. 

In the description of C. Renauldi, M. Cardot states that he has 
not seen authentic specimens of C. Raui, Aust., but this is 4 mis- 
take, for the specimens I sent him were from Austin’s herbarium, 
labelled Austin by “Grimmia Raui, Colorado, Mrs. Roy,” and 
were probably from the same collection as those in the herbarium 
of E. A. Rau, collected by Brandegee sent by Mr. Rau. These 


449 


are the specimens cited by Cardot under C Renauldi, and seem to 
prove conclusively that the two species are the same. 

The specimens collected by J. M. Holzinger on exposed sand 
bluffs at Winona, Minnesota, have been determined by M. Cardot 
as C. Renauldi, and by me as C. Rauz. We also have specimens 
collected by Mrs. T. A. Williams at Rapid City, South Dakota, 
which we have referred to this species. : 

The plate of C. Raui is taken from type specimens in Austin’s 
herbarium sent to him by E. A. Rau, collected by Brandegee. 


Description of Plate 248. 

I. Plants natural size; 2. one enlarged; 3. antheridial branch; 4. archegonia 
branch with the calyptra partially exserted; 5, 6,7. outlines of leaves; 8. antheridia 
and bracts; 9. basal half of leaf ; Io. apex, showing long rough awn; I1. calyptra; 
12. capsule with lid; 13. capsule without lid; 14. old capsule with fragments of 
teeth; 15. two teeth; 16. annulus $17. Spores. 


2. DICRANELLA HETEROMALLA AND ITS 
VARIETIES. 
(PLATE 249.) 

According to European authorities this is a very variable 
Species, several of its more striking varieties having received 
Names. The typical form seems to be according to Schimper 
and Limpricht, the one having the pedicels erect but more or less 
sinuous, Braithwaite, however, states that the pedicels may be 
curved, and Boulay says that owing to the variability of the flexion 
Of the pedicel at its summit the capsule assumes very diverse atti- 
tudes. 

Several of the European varieties have been recognized and 
distributed in American exsiccatae; but we have been surprised 
to find that even in this common species there still remain some 
Points to be cleared up, and possibly two allied species to be 
€rased from our lists. 

The common lowland form ranging along the Atlantic plain 
from Newfoundland to Florida, west to Dakota, which grows 
abundantly along sandy roads in dense cushions at the roots of 
trees and on dry banks, seems to be typical, having the seta erect, 
— flexuose and glossy yellow, agreeing with European specimens 
- and exsiccatae, It was distributed in Drummond’s Mosses, 2d 


450 


Ed. No. 54, in Sulliv. & Lesq. Musci Bor. Am. No. 67, and Aus- 
tin Musci App. No. 79. 


DicRANELLA HETEROMALLA ORTHOCARPA (Hedw.). 

Dicranum orthocarpum Hedw. Sp. Musc. 131. p/. 30. 1801. 

Dicranella Fitageraldi Ren. & Card. Bot. Gaz. 13: 197. pl. 13. 
1888. 

This variety is not recognized as distinct by Limpricht, but is 
given as a synonym of the species, into which it merges insensi- 
bly, having the same range. The type locality was at Lancaster, 
Pennsylvania, but we have not seen the type specimens. The 
capsule, as the name implies, is not only erect but straight, in the 
most depauperate forms being so small and black as to suggest 
Ditrichum tortile. This is true of the specimens distributed in 
Drummond’s Southern Mosses as No. 53, and in the original 
specimens of D. Fitzgerald, from Florida, as described and figured 
in the Gazette. M. Cardot has sent us four of these typical speci- 
mens ; all are old and deoperculate, and represent, in our opinion, 
a very depauperate state of this variety. Quite recently we have 
received No. 156 of Ren. & Cardot Musci Am. Sept. collected by 
J. M. Holzinger at Rock Creek, near Washington, D.C. These 
specimens are in fine condition, and contradict the original de- 
scription of D. Fitzgeraldi in several points, the old capsules when 
deoperculate being contracted below the mouth, which is slightly 
oblique ; the walls also are often sulcate, so that we have no hesi- 
tation in saying that these are D. heteromalla var. orthocarpa. 
They agree with Sull. & Lesq. Musci Bor. Am. No. 68, and Aus- 
tin Musci Ap. No. 80. 

The variety ixterrupta Schimp. has also been recognized and 
distributed in American exsiccatae, and the var. séricta occurs 
abundantly on decomposing sandstone rocks in the Dells of the 
Wisconsin. Recently I have made the acquaintance of what may 
be called the mountain form of this species ( Plate 249 ) for which 
we have not yet discovered a name, but which we think has been 
figuring in the Manual as D. curvata. It first puzzled me by 
growing in the dense mats of Campylopus Virginicus Aust., raising 
false hopes that it might be the fruit of this species, which has 
thus far only been collected sterile. The pedicels were strongly 
recurved, and the capsules when fresh, ovoid and smooth; but 


451 


when dried they changed to the characteristic cinnamon color of 
Dicranella heteromalla, with its twisted mouth and sulcate walls, 
and the pedicels became erect. These specimens grew at an 
elevation of 5678 ft. on the summit of White Top, Virginia, on 
rocky ledges in shade, and were in young fruiting stage on June 
26, 1892. Later in the same year, September 20th, I collected a 
few small plants in the trail up Mt. Marcy, in the Adirondack 
Mountains, New York. These also were small plants, with plump, 
ovoid, green capsules and yellow curved seta. But when dried 
the pedicels became erect, the capsules turned yellow, and showed 
unmistakably that they were only Dicranella heteromalla, It was 
also found at other points in the vicinity, usually in shade under 
or on the roots of trees in steep, sloping paths, often in very damp, 
black soil. Along the roadside, near the Lodge, the pedicels’ were 
seldom recurved, though the plants were small. In 1894 the form 
with recurved pedicels was very abundant both on Mt. Marcy and 
Mt. McIntyre, and formed large patches at considerable elevation 
on both these mountains. I had also collected it near Stowe, 
Vermont, in 1884, and been puzzled by the straightening of the 
pedicel in drying. In 1893 it was found growing with Dicrano- 
dontium longirostre on shady sandstone ledges of the Wisconsin 
: Dells, and it surprised me to learn from Dr. Barnes that it was 
the common form of Dicranella heteromalla in that vicinity. It 
has recently been sent to me by D. A. Burnett, collected in the 
Mountains of Pennsylvania on sandstone rocks at Bradford, asso- 
ciated with Dicranodontium longirostre, and it was collected in 
Similar habitat and the same association by Mr. D. A. Hopkins in 
West Virginia. We also have specimens collected by F. L. Har- 
vey in Maine, by Edwin Faxon on the bridle path at 3,500 feet 
elevation and on the summit of Mt. Lafayette ; also from the Lake of 
the Clouds on Mt. Washington. Pringle collected it on Mt. Mans- 
field, Vermont, and A. C. Waghorne in Newfoundland. I find 
three specimens collected by Austin in 1872 in the White Moun- 
tains, which had been sent to T. P. James for determination; ac- 
companying them is an autograph slip from James stating that 
they “ must be a Dicranum, but I do not make it subulatum.” 
Now it is a curious fact that Dicranella curvata, though re- 
Ported from two stations in North America, does not occur in any 


452 


North American herbarium, as far as we can determine. We 
have searched diligently for it both in the field and in several col- 
lections, and have come to the conclusion that this mountain form 
of D. heteromalla with curved pedicels has been mistaken for it. 

It is recorded in the Manual from two localities only: 

“On sandstone rocks, Lancaster, Pennsylvania (T. C. Porter), 
and from the White Mountains (James). Very rare.” 

Dr. Porter very kindly sent us a bit of his specimen, and it 
does not match European specimens of D. curvata. It seems ref- 
erable to depauperate specimens of D. heteromalla var. orthocarpa. 
The White Mountain specimens cited have not been found in the 
herbarium of T. P. James. Macoun does not record it from any 
locality in Canada, nor have we any other record of its having 
been collected by anyone else. 


3. NOTES ON THE GENUS LEERSIA, HEDW. 


Leersia Hedw. Fund. Musc. 2: 88. 1782. 
ENcALypTa Schreb. Gen. 2: 759. 1791. 

This genus is much in need of revision, but we have not been 
able to take the time to do more than set right two or three gla- 
ring mistakes which have come to our notice, in studying the 
Eastern species, and to call attention to collectors in our Western 
and Northern States to the importance of securing good and 
abundant specimens. 

There are five species known to occur in the eastern part of 
the continent, L. extinctoria (E. vulgaris) L. laciniata (E. ciliata) L. 
rhabdocarpa, L. procera and L. contorta (E. streptocarpa). One of 
the Eastern species LZ. rhabdocarpa has been found in Quebec, 
but not yet credited to the United States, it is hoped that it will 
be found in the mountains of New England if attention is called 
to the genus. 

Of the additions in Macoun’s catalogue, £. subspathulata, E. 
leiomitra, E. leiocarpa and E, cucullata are from British Columbia, 
or Athabasca, and although Prof. Macoun has generously furnished 
us with all his collection of this genus, and allowed us to keep 
duplicates, yet we have not felt warranted in taking the time . 
present to compare the specimens carefully. We have thought rf 


453 


very important, however, to correct Kindberg’s confusing remarks 
regarding £. Macounii Aust. and E. ciliata Hedw. 

Encatypta Macouni Aust. Bot. Gaz. 2: 97-1877. hai BF 
Man. 182, 1884. Macoun’s Cat. 6: 94-95 only in part (1892). 

The type of this species is preserved in Austin’s Herbarium 
at Columbia College. The specimens are abundant though im- 
mature. They have been compared with the original description, 
Which is copied in Lesquereux and James Manual. All the state- 
ments but one have been verified; the seta is described as “ min- 
utely papillose, rather densely so above the middle.” They have 
been examined with a magnification of 330 diameters, and out of 
a dozen pedicels on none of them has been found a trace of 
roughness. 

Austin compared it to Z. affinis Hedw. (EZ. apophysata, N. & H.). 
We have also made the same comparison with No. 816 of Raben- 
horst’s Bryotheca Europaea, and find that Austin was right. The 
calyptra agrees exactly in having pale irregular papillae to the 
fringed base, though we should call it rather minutely scabrous 
than « densely papillose.” But the leaf characters are also very 
plain, the margins being strongly revolute, the vein ending below 
the apex, and very rough on the back with double papillae; the 
basal cells are clear with the short transverse walls, yellow and 
thickened into ridges and also papillose. This is true also of £. 
affinis, but the vein in that species is excurrent into an awn. The 
Peristome is described by Austin as “single, the teeth of medium 
length, very narrow and filiform, red, more or less split into two 
€qual segments, nodulose and granulose.” This agrees with the 
figures of £. apophysata in the Bryologia Europaea, but Lim- 
Pricht says the peristome is double and has a short basal mem- 
brane, one-third the length of the teeth, which he figures as 247, 
Page 118. We have been able to detect this in No. 816 of Rab. 
Byoth. Eu. even when the peristome is old, but not in Austin’s 
specimens. This would add another distinction between £. apophy- 
Sata and E. Macounit, though they are evidently closely allied. 

The type locality is «On rocks, Stewart's Lake Mountain, B. 
C., collected by Macoun, June 21, 1875. ue 

On consulting Macoun’s Catalogue, I find this locality is not 
given the first place as it should be, and that the specimens sent 


454 


by me from Mount Mackay and Kakabeka Falls, named F£. celzata, 
duplicates of which I have in my herbarium, are also referred to 
this species, I have re-examined them lately, and still insist that 
the name I originally gave them is correct. Number 133 of Ma- 
coun’s Canadian Mosses was distributed as /. MJacouniz, collected 
“on rocks at Wellington Mine,” Vancouver Island, by Macoun. 
These specimens in our set prove to be Excalypta vulgaris vat. 
pilifera. If these are the specimens on which Kindberg bases 
his remarks, it is not surprising that he could not find the charac- 
ters indicated by Austin. 


Under the heading of £. ciliata he says (Macoun Cat. 6: 94): 


“Encalypta ciliata Hedw. and £. Macounii Aust. are very 
difficult to distinguish apart. The descriptions of the best authors 
are also not consistent.” He then quotes from them to show that 
the position of the teeth does not agree in all the descriptions. 
From this he says: “It is probable that the authors are confound- 
ing both species, also occurring in Europe. F#. ciliata is princi- 
pally found in the lower mountain districts. /. Macounii seems 
to be an alpine species also collected by Kindberg in the Nor- 
wegian Alps and considered a new species, &. dorealis Kindb. 
Laubm. Schwed. & Norweg., but exactly agreeing with the orig!- 
nal specimens of &. Macounii sent by Prof. Macoun.” 


It is interesting to know what Norwegian bryologists think 
about this, and we would refer to the remarks by Chr. Kaurin in 
the Bot. Centbl. 41: 358 (1890). He states that in the contribu- 
tion referred to by Kindberg the specimens which were called £x- 


calypta Macounii are found to differ from the portion of the type 
which I sent him. 


_ To continue the quotation from Macoun’s Catalogue, page 95+ 
“The description made by Austin, cited by Lesq. & James, 15, 
however, not completely exact; ‘calyptra densely papillose, pedi- 
cel reddish, papillose, the leaves muticous ;’ such characters are 
not to be found, the calyptra and the yellow pedicel are nearly as 
smooth as in the true &. ci/iata, to which the descriptions of the 
peristome by Schimper and Braithwaite probably belong. The 
descriptions by Lesq. & James and Boulay could partly be re- 
ferred to £, Macouni, although all authors agree that in the 
description of £. ciliata,‘ without a distinct collum;’ £. Macount 
has a distinct collum and the margins of the leaves distinctly Te 
flexed. I possess no specimens of the true £. ciliata from No 
America. It may not occur there.” 


Now it seems evident from all this that Macoun has sent Kind- 


455 


berg a specimen of Excalypta closely allied to £. ciliata, wrongly 
named £. Macounii, and from this false premise he has arrived at 
the above conclusions. Whether all the specimens cited in Ma- 
coun’s Catalogue as E. Macounii were sent to Kindberg is not 
clear, but probably this is not the case. It is likely that only a 
few of them were, and that the rest have been transferred without 
examination by Prof. Macoun in making up the list. 

We note one that was not transferred, for No. 132 Canadian 
Mosses is cited under £&. ciliata, with the label locality reading: 
“Crevices of rocks, common from Ottawa westward.” These 
specimens have been examined and compared with the types of 
£. Macounii, They are quite distinct in the excurrent vein, plane 
Margins, smooth calyptra, and the mouth bordered by 5-6 rows of 
hexagonal cells projecting above the base of the teeth. We have 
also examined all the other specimens in Austin’s herbarium sent 
to him by Macoun and named by Austin £. cata. These in- 
clude specimens collected at Stewart’s Lake, date and locality the 
same as type of 4. AMZacounit, but distinctly FE. ciata; also those 
from Lake Athabasca, August 29, 1875; Cascades, May 17, 1875 ; 
and Hastings county, August, 1874. All of these agree with the 
characters of E. ciliata as described by Limpricht, even to the size 
of the spores, but in many cases it is difficult to distinguish the 
Preperistome as described by him. The deep projecting border 
of the mouth is quite distinct, as well as the scattered stomata, 
with the surrounding cells not differentiated, but long and thick- 
ened longitudinally, The neck also is short, but always distinct, 
and sometimes stomatose, though usually the stomata are above 
the base of the sporesac. 

ENcatypra ciLiaTa (Hedw.) Hoffm. 

We find in the Jaeger Herbarium all the exsiccatae cited by 
Limpricht, and have critically compared No. 19 of his Bryotheca 
Silesiaca, with our specimens of Macoun’s Canadian Mosses No. 
132. and Sull. & Lesq. Musci bor. Am. No. 165, Ed. 2. We find 
they agree in all the characters described, with a certain amount 
of variation in the length of the awn, and the vein which is also 
Sometimes serrulate on the back for a short distance below the 
apex of the leaf; the margins are more or less undulate and 
slightly revolute below, erose papillose above, and the basal cells 


456 


are clear and smooth, with the short transverse walls often brown 
but not projecting. The seta varies somewhat in length, but is 
smooth and yellow, often red at the junction with the capsule; 
there is a distinct neck measuring .2 mm., that is the sporesac 
does not reach the base of the capsule, the stomata are scattered 
above its base, and the cells of the walls are long and thickened, but 
not radiating around the stomata; those of the mouth are shorter 
and hexagonal in 3-6 rows, and project above the base of the 
orange-red teeth, the last rows falling in fragments with the lid. 
The beak of the lid is shorter than the capsule, 1-1.5 mm. and 
the calyptra is fringed at base and generally entirely smooth, 
though occasionally scabrous at apex. The peristome is erect 
when dry, and strongly incurved when moist; the teeth are com- 
posed of 5—7 joints, and are paler and smoother at apex. Lim- 
pricht describes a “ preperistome of 32 isolated, smaller, brownish- 
red plates half the length of the teeth, falling off, or occasionally 
lacking.” We have had great difficulty in distinguishing these 
plates, both in our specimens and the Europeon ones, but find the 
teeth are irregularly papillose, that is some joints will be and 
others not, especially the upper ones, which are generally lighter 
colored, and presume that is only under very favorable circum- 
stances, on fresh young teeth, that the preperistome can be seen. 
The Bryologia Europaea indicates this in figures 14 and 15, as 
well as a certain amount of irregularity in the teeth which we have 
also observed. The spores are quite alike in American and 
European specimens, in size and the peculiar lines due to shrink- 
ing which give them the aspect of a rose-cut diamond with a flat 
central facet and six radiating around it. 

If there are two species mixed under Encalypta ciliata, as 
Kindberg seem to think, we have not yet been able to detect it in 
American specimens, as compared with authenticated European 
ones. It has been distributed as Drummond's No. 50 PP- 
“ Rocks and banks along the Mountains,” mixed with £. rhabde- 
carpa in Sull. & Lesq. Musci Bor. Am. Ed. 2, No. 165, in Austin’s 
Musci App. No. 174 (1870) and Macoun’s Canadian Mosses No. 
132. It ranges in the Eastern States from the mountains of New 
York and New England to Illinois, Wisconsin and Minnesota, 
Ontario, Quebec, New Brunswick, and north to Greenland. In the 


457 


Rocky Mountains from New Mexico to Idaho, also in California, 
Oregon and Washington, through British Columbia, to Behring 
sea. 

Var, microstoma (Bals. & DeNot.) Sch. Br. Eu. Index, p. 7. 
1855. LEncalypta microstoma Bals. & DeNot. Pugill. No. 18. 1836. 

We received from the Department of Agriculture specimens 
collected by Wolf and Rothrock in Colorado, 1873, which agree 
with the description given by Limpricht of this variety in the 
short seta, small capsule, with a small mouth, short teeth, lacking 
on some of the capsules, and the cells of the walls broader and 
much thickened, with very distinct stomata. The spores, how- 
‘ver, seem to be smooth as in the species. 


ENcALypPTa LONGIPES Mitt. Journ. Linn. Soc. 8: 29. 1865. 

We quote the original description: 

Dioica ? caule brevi ramoso, foliis patentis concavis ambitulate ellipticis acutis 
Paulo supra basin angustatis nervo percurrento obtuse carinatis, margine minute eroso, 
cellulis basi infima oblongis hyalinis inde viridibus mox abbreviatis quadrato-rotunda- 
tis papillosis, perichaetialibus parvis vaginulam vix superantibus latissime ovatis acutis 
theca in pedunculo longissimo rubro ovato-cylindracea basi apophysata, operculo subu- 
lato subaequilongo-peristomo dentibus angustis elongatis, calyptra basi nuda apice 
laevi.” 


“Seta an inch and a half long, slightly flexuose. Capsule too 


immature to show if it is furrowed.” 
Lesquereux and James, in their Manual, on page 183, say ina 


foot note: 

“Upon examination of the specimens of £. /ongipes Mitt. in all 

of Drummond’s sets the characters have been found identical with 

ose of LZ. procera; the plants monoecious ; the calyptra papil- 
lose, its borders emarginate, erose or fimbriate at base; perichae- 
tal leaves piliferous ; capsule spirally striate,etc. The differences 
that appear in the characters indicated by Mitten result from the 
Unripeness of the specimens he had for examination.” 

We were permitted by Dr. Watson to examine the type of 
this species at the time that Mitten’s types were still at Cam- 
bridge, and we have compared them with the original description 
and with the above remarks. We can corroborate Mitten’s state- 
ments, with additional evidence that this is not the same species 
as £. procera. 

The calyptra is smooth throughout, and so is the vein of the 
Narrow leaf, which, moreover, as the description says, has a 


Pere Ort a ee 


458 


minutely erose margin, formed of projecting cells which appear 
smooth, not densely papillose as in E. procera. The upper cells 
of the leaf, in fact, are not papillose but mamillate, projecting on 
both surfaces of the apex of the leaf, but slightly. Not only is 
the peristome also described as simple, but a drawing is preserved 
with the type, showing a single row of teeth, with 2-3 appendicu- 
late basal segments. Kindberg in Macoun’s Catalogue says that 
he too has seen authentic specimens of Drummond’s in the col- 
lection of Dr. C. Mueller, and states that the “costa is very rough, 
the calyptra regularly laciniate, very rough and subspinulose 
above.” 

It is evident from these remarks, however, that the specimens 
could not have been authentic, for this is a direct contradiction of 
the original description, verified by a reéxamination of the type 
specimen. 

We do not like to venture an opinion without careful com- 
parisons, especially as this species is so meagrely represented in 
the type, which is also immature, so that it is rather unsatisfactory 
for purposes of comparison; yet it seems to be very closely re- 
lated to £. leiomitra Kindb., belonging to the group of £. rhabdo- 
carpa Schwaeg., from which it differs presumably in its smooth 
capsule (?), although the original description states that the speci- 
mens are too young to show this character. 


Two undescribed Species of Rhynchosia. 


By ANNA MuRRAY VAIL, 


/ RHYNCHOSIA MICHAUXII. 


Perennial. Stems prostrate 6-9 dm. or more long, twining above, 
angled, channelled, clothed, especially on the angles, with a short 
matted pubescence; stipules 2-5 mm. long, ovate-lanceolate, spread- 
ing, persistent ; petioles angled, striate, pubescent, 2-4 cm. long, 
leaves 2.5-5 cm. long, depressed orbicular, much dilated, broader 
than long, obtuse, sometimes obscurely mucronulate, rugose an 
minutely hirsute above, sparingly resinous-dotted, reticulated and 
softly hirsute beneath; the upper ones rarely 3-foliolate with 
obtuse obliquely sub-orbicular lateral leaflets; racemes 1-several 
flowered; peduncles 1~2.5 cm. long, angled; calyx I-1.5 cm. 
long, pubescent, resinous-dotted, the oblong-lanceolate acuminate 


459 


lobes conspicuously foliaceous; corolla pale yellow; vexillum 
minutely pubescent above ; legume about 1.4 cm. long, 6 mm. 
broad, obliquely acute, seed 2-5 mm. in diameter, red-brown. 
March-September. 

Dry pine barrens, Florida. 

Charlotte Harbor, Blodgett; Tampa, A. H. Curtiss, No. 659; 
DeLand, Hulst ; Sanford, Lake Co., G. V. Nash, No. 2314, in 


Herb. Columbia College. 

A species with the leaf-form of R. simphicifolia, but the habit 
of R. menispermoidea. It has passed under the above names and 
also under that of R. tomentosa var. volubilis, but is very distinct 
from all forms of the type of the latter species. The lateral leaflets 
of R. tomentosa are always acutish, those of R.Michauxi being ob- 
tuse and rounded. The leaves are largerthan those of A. menis- 
permoidea, less conspicuously veined, and apparently never cordate, 
the flowers more numerous, the calyx larger and more distinctly 


foliaceous. 


“ RHYNCHOSIA TORREYI. © 


Rhynchosia latifolia 8 Torr. & Gray, Fl. N. Am. 1: 285. 1838. 

Perennial. Stems slender, branched, prostrate, spreading and — 
and possibly twining, angled, obscurely striate, minutely pubes- 
Cent; stipules 4-6 mm. long, lanceolate, slender, ec a 
petioles angled, channelled, minutely pubescent, 3-5 cm. ong 
leaves 6-10 cm. long, 3-foliolate ; leaflets 2.5-3.5 cm. long, orbi- 
Cular-obovate or oblong-obovate, obtuse with a short abrupt 
acumination, narrowed at the base, minutely and sparingly pubes- 
Cent on both surfaces, or nearly glabrous, beset with scattered 
Tesinous dots beneath, the terminal ones larger, the others inequi- 
lateral ; racemes 5~8 cm. long, slender, the flowers scattered ; 
flowers about 1 cm. long; calyx 8-10 mm. long, pubescent, resi- 
nous-dotted, the upper lobe 2-toothed, the middle lower Nise 
slightly the longest; vexillum nearly orbicular, 8 mm. long, pu a 
cent along the top on the outer surface, with a very small crescen 
Shaped callosity above the claw. Legume not seen. 

OrIGINAL Locatity. Sand Hill, Texas, Leavenworth. Type 


in Herb. Columbia College. 


460 


Flora of Richmond Co., N.Y. Additions and new Localities, 1891-1899. 


APPENDIX, No. 7. 


Ranunculus Pennsylvanicus L. New Dorp, A. A. Tyler. 
Aguilegia Canadensis L.. Tottenville and Richmond Valley, 
Wm. T. Davis and G. H. Pepper. 

Nymphaea rubrodisca (Morong) Greene. Bull’s Head, J. V. 
Leng. 

Roripa sylvestris (L.) Bess. Sailors’ Snug Harbor, Dr. F. Hol- 
lick, 

Silene nutans L. Arrochar, W. C. Kerr. 

Silene vulgaris (Moench) Garcke. Tottenville, W. T. Davis. 

Tilia Americana L. Willow Brook and New Springville, W. 
T. Davis. 


Acer Saccharum Marsh. Moravian Cemetery and New Spring- 
ville, Wm. T. Davis. 


Acer platanoides L. New Brighton and Todt Hill, Wm. T. 
Davis. 

Acer Negundo L, Port Richmond. 

Medicago sativa L. New Brighton, W. T. Davis; Princes’ Bay. 

Coronilla varia L. Tottenville, W. T. Davis. 

Ulex Europacus L. Ward’s Hill, Tompkinsville, Dr. F. Hollick. 
(A single plant which has persisted for several years). 

Amorpha fruticosa L. Egbertville, W. T. Davis. 

Phaseolus polystachyus (L.) B.S.P. Egbertville, W. T. Davis. 

Opulaster opulifolius (L.) Kuntze. Todt Hill, W. T. Davis; 
Willow Brook (some undoubtedly seedlings). 

Rubus odoratus L. Todt Hill, W. T. Davis. 

Crataegus coccinea L. Karle’s Neck, W. T. Davis. 

Crataegus punctata Jacq. West New Brighton. 

Agrimonia mollis (T. & G.) Britton, and Agrimonia striata 
Michx., replace A. Eupatoria L. in our catalogue. 

Valeriana officinalis L., Gifford’s Lane (Replaces V. sylvatica 
Banks, in our catalogue). 

Onagra Oakesiana (Gray) Britton. New Dorp. 

Ethusa Cynapium L. Streets of New Brighton, W. T. Davis. 

Anthriscus vulgaris (L.) Hoffm. New Dorp. 


461 


Philadelphus coronarius L. Todt Hill, W. T. Davis (apparently 
established from old garden waste). 

Solidago Elliottii T. & G. Garrettsons and New Dorp. 

Solidago patula Muhl. Garrettsons and Richmond, W. T. 
Davis. 

Gnaphalium purpureum L. Egbertville. 

Lusstlago Farfara L. Garrettsons. 

Liatris spicata (L.) Nutt. Mariners’ Harbor, W. T. Davis. 

Sericocarpus linifolius (L.) B.S.P. Watchogue, W. T. Davis. 

Centaurea nigra L. Moravian Cemetery. 

Onopordon Acanthium L. Sailors’ Snug Harbor, Dr. F. Hollick. 

Schollera macrocarpa (Ait.) Britton. Tottenville, W. T. Davis ; 
New Dorp, Mrs. N. L. Britton; Kreischerville. 

Azalea viscosa rosea Hollick. Arlington, W. T. Davis. 

Gaultheria procumbens L.  Giffords. 

Kalmia angustifolia L. Bogardus’ Corners. 

Pyrola secunda L. Bogardus’ Corners. 

fraxinus viridis Michx. f. Clifton and New Dorp. 

Cynoglossum officinale L. Arlington, W. T. Davis. 

Convolvulus Sepium repens (L.) Gray. Oakwood. 

Ipomoea pandurata (L.) Meyer. Mariners’ Harbor, W. T. 
Davis. 

Gerardia purpurea paupercula Gray. New Dorp. 

Veronwa Anagallis-aquaticaL. New Brighton, Dr. F. Hollick. 

Pentstemon Digitalis (Sweet) Nutt. New Dorp, Mrs. N. L. 
Britton; West New Brighton, T. C. Leng. 

Soechys cordata Ridd. Eltingville, W. T. Davis. 

Mentha gentilis L. Egbertville, A. A. Tyler. 

Mentha sativa L. New Dorp. 

Mentha citrata Ehrh. Richmond Valley. 

Conopholis Americana (L. F.) Wallr. Todt Hill, Mr. Stottler. 

Plantago aristata Michx. St. George, in recently filled-in 
ground, 

Brousonnetia papyrifera Vent. West New Brighton, Mariners’ 
Harbor and Richmond, W. T. Davis. 

Alnus glutinosa Willd. Egbertville, W. T. Davis (A number 
of trees, thoroughly naturalized and apparently spreading). 

Alnus incana (L.) Willd. Grant City. 


462 


Quercus Britton Davis. Watchogue. 

Salix fragilis latifoka And. Todt Hill. 

Populus heterophylla L. Huguenot, W. T. Davis. 

Tipularia unifolia (Muhl.) B.S. P.  Tottenville. 

Calopogon pulchellus R. Br. Watchogue, C. W. Leng. 

Hlabenaria blephariglottis (Willd.) Torr. Watchogue, W. T. 
Davis. 

Cypripedium acaule Ait. An albino form. Mrs. Heylyn. 

Tradescantia Virginica L. Arlington, W. T. Davis. 

Wolffia Columbiana Karst. Old Town Pond, Thos. Craig. 

Potamogeton Spirillus Tuckm. Court House, Mrs. N. L. 
Britton. 

Udora Canadensis Michx. Clove Lake, W. T. Davis. 

Pinus echinata Mill. Linoleumville and not uncommon along 
the south side, W. T. Davis. Arrochar and Four Corners. 

Eriophorum Virginicum L. Tottenville, W. T. Davis; Gifford’s 
Mr. Twiggs. 

Panicum pubescens Lam. Richmond Valley. 

Panicum microcarpon Muhl.  Tottenville. 

Panicum commutatum Schultes. Richmond Valley. 

Panicum verrucosum Muhl. New Dorp, A. A. Tyler. 

Carex tenera Dewey. New Dorp, A.A. Tyler. 

Carex muricata L. New Dorp. 

Carex Muhlenbergiti Schk. Abundant at Richmond Valley. 

Carex tribuloides Wahl. Tottenville and Grant City; A. A- 
Tyler. 


Dryopteris cristata (L.) Gray. Oakwood, Mrs. N. L. Britton. 
Mariners’ Harber. 


Azolla Caroliniana Willd. Clove Valley, Thos. Craig. (Intro- 
duced some years ago by Mr. Samuel Henshaw.) 
Salvinia natans(L.) All. Silver Lake and in a small pond on 
Ocean Terrace, Thos. Craig. (Probably introduced.) 
ArtTHUR HOLLICcK, 
N. L. BRITTON. 


463 


New or Noteworthy American Grasses,—II. 
By Gero. V. Nasu. 


SPOROBOLUS ASPER (Michx.) Kunth. Enum. 1: 210. 18 33. 
Agrostis aspera Michx. Fl. Bor. Am. 1: $2. 160% 

Vilfa Drummondii Trin. Mem. Acad. St. Petersb. (V1.).5¢ 
Nat. 4: Botanique, 106. 1845. 

Sporobolus asper var. Drummondti Vasey, Contr. U. S. Natl. 
Herb. 3: 60. 1892. 

This grass and the next seem to have been confused. The 
plant named S. asper by Dr. Vasey is the Agrostis longifolia Torr., 
and not the 4, aspera of Michaux, who apparently had in mind, 
So far as can be determined from his short description, the form 
with a long palet. The plant of Michaux, as here understood, has 
a long-acuminate, sometimes almost awned, palet, which usually 
much exceeds the scale; the outer scales narrow and acute; the 
culm and panicle, which is generally exserted, quite slender. 

Grows from Delaware to Illinois and Missouri, south to Florida 
and Texas, 

Vilfa Drummondit Trin. may eventually be separated as dis- 
tinct, but at present it seems preferable to refer it to this species, 
intermediate forms being very numerous and apparently connect- 
ing the two, 

SPOROBOLUS LONGIFOLIUS (Torr.) Wood, Class-book, 775. 1861. 
Agrostis involuta Muhl. Gram. 72. 1817. Not Poir. 1810. 
Agrostis longifolia Torr. Fl. go. 1824. 

Vilfa Hockeri Trin. Mem. Acad. St. Petersb. (VI.) Sc. Nat. 4: 
Botanique, 106. 1845. 

Sporobolus asper Vasey, Contr. U.S, Natl. Herb. 3: 59. 1892. 
Not Kunth. 1833. 

This is usually a more robust plant than the preceding. The 
Panicle js generally more or less included in the upper sheath, 
SOMmetimes merely protruding from the sheath fissure. The spike- 
lets are wider, with broad, obtuse scales and palet, the latter about 


€qualling the third scale. 
Occurs from Maine to Illinois and Kansas, south to Long Is- 


land, Tennessee, Mississippi and Texas. 


464 


SPOROBOLUS VAGINAEFLORUs (Torr.) Wood, Classbook, 775. 1861. 


Vilfa vaginacflora Torr. A. Gray, Gram. & Cyp. No. 3. 1834. 

Sporobolus minor Vasey; Wats. & Coult. in A. Gray, Man. Ed, 
6, 646. 1890. Not Kunth 1833. 

Sporobolus filiculmis L. H. Dewey, Contr. U.S. Natl. Herb. 2: 
519. 1894. 

The type of this plant, as indicated in A. Gray, Gram. & Cyp., 
is the form with long and narrow spikelets, the scales narrow and 
acuminate. Dr. Vasey took up and named as S. vaginacflorus the 
form with shorter and broad spikelets, the scales merely acute. 
This plant is considered below. 


SPOROBOLUS NEGLECTUS nom. nov. 


Sporobolus vaginacflorus Vasey, Wats. & Coult. in A. Gray, 
Man. Ed. 6, 645. 1890. Not Wood. 1861. 


This is the plant that has long passed as Vilfa vaginaefiora 
Torr., but it is not the same, as indicated under the preceding 
species. Owing to this mistaken identity this plant appears to be 
. without a name, and so the one given above is proposed. The 
broader and shorter spikelets, with the scales broad and merely 
acute, will readily separate this plant from the S. vaginacflorus 
Wood. 

In the vicinity of New York it is much less common than S. 
vaginacflorus, which grows plentifully in many localities. 


Dry sandy soil, Massachusetts to Kentucky, Tennessee and 
Kansas. 


SPOROBOLUS BREVIFOLIUS (Nutt.) Scribn. Mem, Torr. Bot. Club. 
5: 39. Inpart. 1894. 
Agrostis brevifolia Nutt. Gen. 1: 44. 1818. 


There seems to be much confusion in regard to Sporobolus de- 
pauperatus and S. cuspidatus. Forms of what appears to be a 
good species have been variously referred to either of the above. 
They agree well with the description of Agrostis brevifolia Nutt., 
and they are here referred to that species. The leaves are short, 
-1/-2' long; the empty scales obtuse or merely abruptly acute, less 


than one-half as long as the acuminate and sometimes short-cuspi- 
date flowering scale. 


465 


Grows from Anticosti Island and Maine to British Columbia, 
south in the mountains to New Mexico and California. 

S. depauperatus (Torr.) Scribn. is a plant of the extreme north- 
west, occurring in Washington and Oregon, possibly extending 
southward in the mountains. The culms are short and decum- 
bent, the internodes very short, usually 14’ long or less, the nodes 
often swollen; the empty scales broad, thin, white, delicate in 
texture, obtuse or acutish, more than one-half the length of the 
broad flowering scale, sometimes about equalling it. 

In S. cuspidatus (Torr.) Wood the empty scales are acuminate 
and short-awned, more than one-half as long as the acuminate 
and awn-pointed narrow flowering scale. The plant is usually 
taller and has much longer leaves than either of the two species 
above mentioned, and ranges from Manitoba to the Northwest 
Territory, Missouri and Kansas. 


ERAGROSJIS TRICHODES (Nutt.). 

Poa trichodes Nutt. Trans. Am. Phil. Soc. 5: 146. 1833-37. 

Eragrostis Geyeri Steud. Syn. Gram. 272. 1855. 

Eragrostis tenuis A. Gray, Man. Ed. 5, 632. 1867. Not 
Steud. 1855, nor Poe tenuis Ell. 1817. 

Nuttall’s name for this plant is the oldest, and is here taken 
up. This grass extends from Illinois westward and southwest- 
ward, and is not known to occur in South Carolina or Georgia, the 
Tegion covered by Elliott’s Botany. It could hardly, therefore, 
be the same as the Poa éenuis of that work, which, according to a 
Specimen of that plant in the Columbia College Herbarium, 
labeled as coming from Elliott, is apparently the same as 
the grass now known as &ragrostis capillaris Nees, and 
which well accords with the description given by Elliott of his 
Poa tenuis. 

“Poa BUCKLEYANA nom. nov. 
Poa tenuifolia Buckley, Proc. Acad. Phila. 1862: 96. 1862. 


Not A. Rich. 1851. b 1 
~ No other name for this plant can be found, and so the above is. 


given itin honor of Dr. S. B. Buckley, who first published a de- 
Scription of it. 


466 


Vegetable Spiralism. 


By GEORGE MACLOSKIE. 


My paper on Antidromy* attempted to show that every pheno- 
gamic plant produces two kinds of seeds, having their embryos 
turning in opposite directions, according to the side of the car- 
pellary leaf on which they originated ; also that the forthcoming 
plants have a primitive twist in opposite directions, showing itself 
in the histology of the stem, the dextrorse or sinistrorse phyllotaxy 
and anthotaxy, and in some cases in the structure of the carpels. 
The statement that the grains of Maize produce dextrorse or sinis- 
trorse plants, according to the orthostichy of the ear in which they 
were produced, depended on hasty dissection; and although it 
seemed at first to be confirmed by the seedlings raised from the 
grains, I now find that there is no apparent constancy in this matter, 
My last experiment included nineteen seedlings gréwn from 
twenty grains taken in order from one of the paired rows in an ear 
(the row opposite my left hand) giving Nos. 1, 2, 3, 6, 8,9, 10, 12, 
15, 17, 20 with sinistrally overlapping leaves (Sc. the Ist leaf 
above the pileolus having its left margin external), and Nos. 4, 5, 
7, 11, 13, 14, 18, 19 with dextrally overlapping leaves (No. 16 
being abortive): thus giving eleven of one kind against eight of 
the other kind. In the grains from another homologous row, this 
proportion was nearly reversed. It is therefore necessary for the 
present to regard the beautiful symmetry of the ear of Maize 
(more beautiful in its early development than in its maturity) as 4 
rearrangement of members at first as promiscuously arranged as 
are the staminate flowers of its panicle.+ 

Besides cases previously cited we have significant examples of 
antidromy in the seeds of Salsola (figured in Engler and Prantl, 
3: 1a, 84. Y, Z), in the spirals of the Screw-pines, in the 
florets on the large head of the Sunflower and other large Com- 
positae, which concur with the phyllotaxy, and even in the sword- 
like leaves of Acorus Calamus, those of one plant being dextrorsely 


*BULLETIN, 22: 379. Correct: erratum, p- 380, three lines from foot, “ dis- 
tichous,” so as to read * tristichous.” 


+I am obliged to the eminent agrostologist, Prof. W. J. Beal, for friendly criticism 
on this point. 


467 


twisted, and of another plant sinistrorsely. All the long leaves of 
all plants of Lypha (both species) have a sinistral twist, in this 
showing no antidromy; but the mode of overlapping of the mar- 
gins of their leaves near the nodes, which is the same for all the 
leaves of one plant, is contrary as between different plants. If 
this is a case of genuine antidromy, it is an argument against the 
view that the ovules of Typha arise terminally on the floral axis, 
a view which on other grounds has been doubted. 

In the somewhat decussate leaves of Paulownia, and of the 
Shaded branches of forsythia, we can trace a spiral twist by fol- 
lowing any one of the leaf orthostichies (they twist all in one di- 
rection about Princeton, where the plants are not propagated from 
seeds), But the branches of /orsytiia when exposed to the sun- 
light lose their decussation and have all their leaves in two hori- 
zontal rows, thus demonstrating the controlling influence of the 
light. I believe that in a similar way many of our plants, as Elm, 
Beech and Morning-glory, have sacrificed their primitive phyllo- 
taxy to the allurements of sunshine. 

The Coniferae furnish an example showing how growth may 
Supersede or even reverse the primitive order of parts. The young 
cones of Picea, Tsuga and other trees have the same spiral caste 
as the phyllotaxy, when we estimate their spirality by taking the 
longer of the two dominant curves as we do for the phyllotaxy. 
But after the cones open to give exit to the seeds, there comes a 
displacement of the scales, producing a false spirality in the con- 
trary direction; thus the same tree, or even the two sides of a 
half-opened cone may present a quasi-antidromy. This may pos- 
Sibly be the explanation of published observations as to Coni- 
fers having antidromic cones on the same tree.* 

The cases of Arum, Jris and Juncus were previously referred 
to as giving antidromic plants produced not by seeds but by divi- 
Sion of the same rootstalk. A more curious case of diversity 
Within the same plant is that of Bilsted (Liguidambar Styracifiua), 
Which is cited in books as having opposite spirals in stem and 
branches. Every branch of this tree is true to its own phyllotaxy, 
with a 2 divergence, dextrorse or sinistrorse for each, and this per- 
Sisting through the annual innovations, whose crowded scale-scars 
Verreepe sane 


* American Naturalist, August, 1873. 


468 


are true to their phyllotaxy. In the horizontal members even the 
ridges of cork partially conform to the phyllotaxy of the particu- 
lar member by a slight flexure to one side or the other. But the 
branches fromthe stem, or the branches from a branch do not 
necessarily conform to the phyllotaxy of the part from which they 
diverge; some of the daughter-branches may be homodromic, 
others heterodromic, as compared with the mother-branch, nor 
have I been able to discover any law in the case. 

Perhaps these instances may favor the view that what we have 
in antidromy is not so much a special kind of heredity as an im- 
pulse depending on physical or nutritional causes, giving a bias 
to the young embryo or the young bud, which when once received 
is maintained. But whatever be the explanation offered, the facts 
are too important to be overlooked. 

Sachs in his Astory of Botany treats Piylataxy as an ex- 
ploded error, and gracefully celebrates its obsequies with the part- 
ing note that though wrong, it was useful in its day, adding “ we 
would as little wish to omit it from our literature as modern as- 
tronomy would wish to see the old theory of epicycles disappear 
from its history.’ But what he and others condemned was an 
artificial or idealistic law imposed on plants in mathematical dra- 
pery; the twists that will naturally result from pressure upon 
young parts or from peculiarities of direction and of amount of 
nutriment, and from the modifying influence of light or climbing 
habits, are conceptions that were unknown to the fathers of 
mathematical phyllotaxy. From inattention to these considera- 
tions our botanists often miss what ought to be plain enough. 
Thus among the cryptogams Sachs gives us spores of Eguisefum 
with a wrong spirality for their elaters (I confess my own sin 
here). Dodel-part gives odgonium of Chara wrong twisted ; Engler 
and Prantl seem to require amendment as to Evodium (3: 4- 2) 
and as to Flalicteres (3: 6.93); and such instances may be multi- 
plied. 

Homodromic spiralism is not infrequent in phenogamic as 
well as in cryptogamic plants; as the dextrorse twining of some 
Leguminosae, of Convolvulaceae (including the Dodders) and of 
Celastrus ; and other species are sinistrorse. Whilst it was shown 
in the paper on Antidromy that the mode of bursting of the car- 


469 


pels of Balsam is antidromic, in harmony with the phyllotaxy, we 
find that the carpellary “beaks” of Geranium, Pelargonium and 
Erodium, of the same natural order with Balsam, are all dextrorsely 
twisted. In these cases all the individuals of a species, or even of 
a suborder or order of plants, have the same kind of spiralism. 
But the leaves of all these are antidromic. 

As a contrast to this we sometimes find a quasi-antidromy 
within the same plant, or even inthe same carpel. Thus Azdiscus 
has the corollas of flowers on opposite sides of the same branch 
slightly contorted in contrary directions. The pod of the Lotus 
corniculatus and other Leguminosae bursts open through the forci- 
ble curving of its valves into antidromic spirals. Thesame occurs 
in Foosia of Rubiaceae (figure in Engler and Prantl, 4: 4. 46), 
and in the opening with a spring of the cocci of Ricinus. (A 
branch of this in fruit if left over night on a table may next morn- 
ing have all its cocci opened and its seeds scattered about). 
Doubtless this is the dehiscing mechanism of the sandbox fruit of 
fTura, 

The awns of Gramineae usually have secondary twists of func- 
tional significancy. In Axthoranthum a brown dextrally twisted 
base is surmounted by a pale straight seta ; in Danthonia, Stipa, etc., 
the base is a brown ribbon dextrally twisted, and this is sur- 
Mounted by a rigid sinistrorse style. On the application of water 
the basal ribbon straightens out, causing the style to screw its way 
into the soil, into the wool of a sheep, or into the clothing and 
skin of man (witness Captain Cook’s Crew in Australia in 1770). 
This kind of double twist may be termed didromic (as suggested by 
my colleague, Prof. Brackett). It is found in the sete of some 
Mosses (as figured by Sullivant, /cones muscorum, supplement), for 
example, Munaria Americana, Pottiariparia, Rhynchostegium; or 
Some species of the mosses have the upper part of the seta dex- 
trorsely, and others have it sinistrorsely twisted; but we do not 
See the significance of these peculiarities. 

Charles Darwin showed that in some instances this didromic 
Spirality is a physical necessity, as where tendrils must be 
Shortened and yet their extremities are not to be rotated. This 
is the sort of spiral made by the cord-like scape of Vadlisnenia 
_ Spiratis, so pulling down the fertilized carpel without having to 


470 


turn it round; and I find that in some plants of this species the 
upper section is dextrorsely spiral, and the lower section sinis- 
‘trorsely, whilst in other plants these relations are reversed. Thus 
we have didromy within each plant of Valisneria and antidromy 
between neighboring plants. 

It would be dangerous at this stage to attempt generalizations 
as to the cyptogams. I may be permitted, however, to call atten- 
tion to a few points. Inthe Atlases of Vegetable Palaeontology, 
by Schimper, Zeiler, Lesquereux and others, some of the figures 
of Carboniferous Acrogens, as Lepidodendron, Ulodendron, cones 
of Lepidostrobus, indicate a trend to one side which would indicate 
antidromy if others be found trending to the other side. The 
photograph of Lepidodendron lycopodioides in Plate LXX. of 
Zeiler's Atlas, does give a contrary spiral to that in Schimper’s 
plates, but Zeiler’s photograph may have been reversed in the 
process. The segments of apical cells of Hepaticae and Pterido- 
phytes seem to indicate in some cases a direct or ‘clock-wise’ 
order of appearance, and in other cases a reverse order ; but I do 
not know whether they may not vary in the same individual plant, 
or whether, on the other hand, they may not be homodromic in 
the totality of a species. I have been unable to find any variation 
of phyllotaxy in Lycopodium and allied forms, the great bilateral 
symmetry of the leafy axis obscuring the traces that might exist. 
But in two specimens of the tree-fern Alsophila the common 
phyllotaxy of 2 can be made out at the apex of the stem; in both 
our Princeton specimens the spirality is dextrorse (that is, after 
the course of the thread of a common screw). If anybody can 
produce a specimen with sinistrorse phyllotaxy, he will thereby 


furnish the lacking evidentiary fact, and complete the proof of 
the antidromy of the ferns. 


Botanical Notes. 


Two new botanical Serials. One of our esteemed contempo- 
raries has recently expressed some irritation on learning of the 
founding of a new serial publication. The number of opportuni- 
ties for the publication of botanical papers is indeed great, but the 
supply of matter evidently exceeds the space provided, for we 


471 


very seldom learn that any journal has suspended printing, 
and most of them have had such pressure brought upon their 
Space that they have been obliged to increase their number of 
pages. The “ Journal of Botany” is indeed an exception to the 
general rule, having continued about the same annual dimensions 
for a long time. 

The new channels of publication are both German. The Ber- 
lin Botanical Garden began in January to issue its “ Notizblatt des 
Kéniglichen botanischen Gartens und Museums zu Berlin,” pro- 
posing to distribute it at irregular intervals as matter becomes avail- 
able. The first number bears date January 2d, the second June 
5th. They contain much interesting information on species 
grown in the Garden, on additions to the Museum, and numerous 
descriptions of new species in the Herbarium contributed by 
Engler, Urban, Gurke, Schumann and Gilg. 

_ The “Allgemeine botanische Zeitschrift, fiir Systematik, 
Floristik, Pflanzengeographie, etc.,’ was also commenced in Janu- 
ary and is published monthly at Karlsruhe, under the editorship 
of A. Kneucker. It isa general botanical journal, containing be- 
Sides original communications, reviews, notes on literature and on 
; institutions, societies, exsiccatae and explorations. iM. ia 


Compound Leaves in Rubus odoratus. The leaves of this species 
are always described as simple. A student of mine, Mr. Millett 
Thomson, has just shown me, from a plant in his garden, that the 
leaf with its long petiole first falls, leaving a short stub or main peti- 
ole behind, which is itself later deciduous. This, it will be recalled, is 
Something like what happens in the Japanese Ampelopsis, except 
that there the leaf is sessile on a long petiole, which falls after the 


blade itself has separated. Both of these plants then have unifolio- 
late leaves, W. WuitMAN BAILEY. 


Sisymbrium altissimum Linn. in Minnesota. Mr, L. H. Dewey 
has called attention, in a recent number of the BULLETIN, to the 
Occurrence of Sisymbrium altissinum Linn. in the side streets of 
Minneapolis, 

This plant has been previously reported from Minnesota locali- 
ties. The Minneapolis Daily Tribune of September 22, 1894, 
Contains an account of my discovery of this species near Minne- 


472 


apolis. Since the first introduction of a few straggling specimens, 

the plant has spread so as to become a nuisance in the elevator 

districts on the outskirts of Minneapolis and St. Paul. Specimens 

have been found also in several other localities in Hennepin, Ram- 

sey and Dakota counties. EpmunD P. SHELDON. 
MInneEaApoLis, Minn., Nov. 12, 1895. 


I found this plant in considerable quantities along the railroad 
at Port Arthur, Lake Superior, in September, 1889. We also 
have a specimen of it collected at Danville, Quebec, in 1894. 

N. L. Britton. 


Reviews, 


Synoptical Flora of North America: Vol. 1. Part I. Fascicle 
I. Polypetalae from the Ranunculaceae to the Frankeniaceae. 
By Asa Gray, LL. D., and Sereno Watson, Ph. D., continued and 
edited by Benjamin Lincoln Robinson, Ph. D. (Issued October 
10, 1895.) 

The second volume of this work was published some years since, 
in two parts, the first part appearing in the year 1884, while 
the second was issued in 1878. In 1886a revised edition of these 
two parts with an appendix of additions and corrections was issued 
by the Smithsonian Institution. The first fascicle of part I., volume 
I., has now been issued from the University press at Cambridge 
after a lapse of eleven years. Dr. Robinson is to be congratulated 
on the completion of the pages under consideration and we are 
promised a second fascicle in the near future. The bulk of the 
matter was written by Dr. Gray; Dr. Watson subsequently 
took up the work and continued it until his death; since then Dr. 
Robinson has completed parts left unfinished by the other authors, 
besides editing their manuscripts by bringing together facts of 
recent discovery, mostly in the form of foot-notes. 

The pages contain a vast amount of useful and valuable informa- 
tion and the book will be welcomed by everyone. Together with 
the many good qualities of this issue appear the same faults that 
characterize the second volume. After a general key to the 
families, the Ranunculaceae are taken up, and other families fol- 


473 


low on the old Candollean sequence to Frankeniaceae inclusive. 
It is to be regretted that so great a work must be continued on a 
system of classification which has proved itself inadequate. The 
want of system in the matter of nomenclature is also to be regretted, 
for here, as in the former parts, sentiment is the predominating 
guide in place of rule. 
The generic limits as a rule are well taken; some genera, how- 
ever, are too composite according to modern ideas. The interpre- 
tation of species is in many cases not as good as our present 
knowledge would permit. In Clematis, C. Addisonii is taken up in 
its Proper place, but C. viornioides, a very distinct form, if not a 
good species, is merely hinted at. Clematis ovata is reduced to 
C. ochroleuca, a position not warranted by the abundant specimens 
of recent collection. The author, following Coulter, has reduced 
Clematis Scottii to a variety of C. Douglasti; to be consistent why 
fot reduce C. Catesbiana to a variety or state of C. Virginiana? In 
Ranunculus we find R. Allegheniensis Britton inserted as a species, 
while R. micranthus Nutt. stands as a variety of R. adbortivus L. 
In Aguilegia, A. saximontana Rydberg is published for the first 
time and well distinguished from the related A. érevistyla Hook. 
Very scant courtesy is paid to A. Canadensts flavifiora, but almost 

. immediately following we find A. caerulea var. albiflora A. Gray, 
described as a new variety (while there are at least two published 

| available specific names for it),and its characters as given separate 
it from the type in only just the same degree that differentiates 
flaviflora from Canadensis. We can only assume that the editor 
holds a color variety proposed by Gray to be valid, while rejecting 
One maintained by others. On page 76, Nymphaca odorata var. 
minor appears in the old stereotyped form. Why should this 
State be called a variety when it is the original of the species 
odorata? Ifa variety must be maintained it should be done in a 
Consistent way. Little recognition is given an apparently dis- 
tinct species of Castalia from Florida and adjoining territory- 
The editor says ‘this form has been confidently identified with 
N. rentformis Walt. by the collector, . . adisposition in no sense 
Warranted by the brief and wholly dubious characterization of 
Walter’s species.” If there were nothing but Walter’s description 
to Support Mr, Nash’s position, it certainly would be weak, but 


474 


careful field observations, morphological characters separating the 
form from related species, and the fact that the plant is very 
common in Walter’s region, furnish pretty strong evidence in favor 
of its correct identification. So other cases might be cited. The 
geographic ranges of many species as given are too restricted, as 
a few appended examples will show, and no data bearing on alti- 
tudinal distribution are furnished. 

Clematis Viorna. Common east of the mountains as far south 
as middle Georgia. 

Clematis Addisonii. Also in the Cumberland Mountains, 
Tennessee. 

Clematis Douglasit Scotti. Also in Montana and Dakota. 

Clematis verticillata. Occurs in southwestern Virginia. 

Clematis alpina tenuiloba. Also in Colorado. 

Thalictrum cortaceum, Also in the Cumberland Mountains, 
Tennessee. 

Myosurus minimus. Occurs in southeastern Virginia. 

Ranunculus recurvatus. Also in Montana. 

Ranunculus abortivus micranthus. Common south to south- 
ern Pennsylvania. 

Ranunculus septentrionalis. West to Nebraska. 

Caltha palustris. Also west to Nebraska. 

Aconitum Columbianum. East to the Black Hills, South 
Dakota. 

Cimicifuga racemosa cordifoha. Occurs in the mountains of 
Virginia. 
_ Xanthorrhiza apiifolia. Ranges east of the mountains and as — 
far south as middle Georgia. 

Delphinium tricorne. Occurs as far west as Nebraska. 

Delphinium exaltatum. Also west to Nebraska. 

Magnolia tripetala. Grows as far south as middle Georgia. 

Asimina triloba. Occurs east to New Jersey and west to 
Nebraska. 

Asimina angustifolia. Rather common as far north as middle 
Georgia. foe 

Brasenia Schreberi. Also south to Georgia and Florida. 

_Menispermum Canadense. Occurs west to Nebraska. 

Berberis repens. Ranges east to the Black Hills, South Dakota. 


475 


Argemone alba. Grows in southwestern South Dakota. 

Dicentra Canadensis. Occurs in Nebraska. 

Corydalis curvisiligua. East to Louisiana and Florida, also 
north to Nebraska. 

Corydalis aurea occidentalis. Common in N ebraska, 

Arabis dentata. Occurs west to Nebraska. 

Arabis laevigata. Also south to middle Georgia and west to 
the Black Hills, South Dakota. 

Arabis Holboellii. Ranges east to Nebraska. 

Lesquerella Ludoviciana arenosa. Occurs in the Black Hills, 
South Dakota. 

Physaria didymocarpa. Grows in Nebraska. 

Nasturtium sessiliflorum. Ranges west to Nebraska. 

Cristatella Jamesii. Common in western and central Nebraska. 

Cleome lutea. East to Nebraska. 

Cleomella angustifolia. Also in Colorado and Nebraska. 

Viola palustris Occurs in the Black Hills, South Dakota. 

More or less inconsistency in the treatment of families is ap- 
parent; compare, for an example, the Ranunculaceae with the 
Cruciferae; in the former the genera are treated as collectives, 
while in the latter they appear as segregates and are naturally 
much clearer. We are pleased to note that the Cruciferae is the 
best described family in the fascicle. 

The book has very few typographical errors, but the often 
Copious foot-notes spoil the appearance of the pages. The matter 
Contained in these notes might better have been incorporated in 
the text. A curious statement occurs on page 42, where we are 
told that Eranthis hyemalis is “a relict of former cultivation.” On 
Pape 191, Table Mountain is given as the locality for Hudsonta 
montana. Table Rock is meant, as Table Mountain is not in 
North Carolina, but in South Carolina and isa very different place. 
Dr, Gray confused these two names many years ago and botanists 
seem to have been unable to get them right since. 

J. K. Smart. 

Our Edible Toadstools and Mushrooms and How to Distinguish 
Them is the title of a volume recently issued by the well-known 
firm, Harper & Brothers, New York. The author is W. Hamil- 
_ton Gibson, who has for a long time given special attention to the 


476 


subject here set forth. Previous American booklets, reports, etc., 
bearing on this subject have been less extensive, complete and 
satisfactory, yet they have been, as it were, skirmish lines that 
have made this.one possible. It contains 337 pages, and, accord- 
ing to the title page, 30 colored plates and 57 other illustrations 
The full-page plates are 38. An “Introduction” of 40 pages is 
no less interesting and important than the body of the work, 
which is devoted to the descriptions and illustrations of about 30 
edible and a half dozen or more poisonous or suspected species. 
The descriptions are given in a general or popular style, followed 
in each case by a more condensed and systematized one, but in 
both all unnecessary technical terms are avoided, inasmuch as the 
work is especially designed for those who simply desire a better 
acquaintance with our most common and easily-recognized escu- 
lent species, so that they may feel safe in collecting and using 
them for food. Great care has been taken in placing safeguards 
around the genus Amanita that there might be no danger of an 
invasion of its dangerous precincts. Indeed, as a concession to 
the ignorance or carelessnes of such as might otherwise make 
serious mistakes, this line of safety has confessedly been drawn so 
rigidly as to exclude from use even some well-known edible species. 
Nevertheless, the author has done well in showing the worthless- 
ness of many popular rules and tests for discriminating between 
the good and the bad, and in insisting upon a sufficient knowledge 
for the recognition of each species to be eaten, save in a few ex- 
ceptional genera. 

The work is enriched by a list of the edible fungi of North Car- 
olina, taken from the catalogue of Rev. M. A. Curtis, by a history 
of asuccessful use of atropine, the antidote of amanitine, in a case 
of dangerous mushroom poisoning, by a series of thirty-four re- 
cipes for preparing, cooking or preserving mushrooms, and by 
three pages of bibliography pertaining to the subject. Marginal 
indices and a general index at the end of the volume facilitate 
reference to any subject. 

In a work so full of good things and so generally accurate and 
reliable as this is, one feels like uttering only words of praise and 
commendation and passing in silence the very few mistakes that 
might by a critical mind be taken as blemishes. Happily those 


477 


that have met my notice are so few and of such a character as not 
to interfere at all seriously with the design and usefulness of the 
work, and perhaps would scarcely be noticed by any one except a 
close student of mycology, Such a one might say there was some 
mistake in the rather sweeping assertion in the description of 
Russula virescens, that “a faint fluting of the edge” is “a peculiar- 
ity of all the Russula ;” and also that there must be some error in 
describing the spores of Boletus alveolatus in one place as “ rose- 
colored” and in another as “yellowish-brown.” There is also 
quite a discrepancy between the characters assigned in this work 
to Clavaria formosa and Lycoperdon saccatum and those found in 
European works, so that I suspect some misconception of these 
species, but I do not see how these mistakes can lead the my- 
cophogist into any serious danger. 

The publishers have evidently done their part toward making 
the book attractive. Plates, press work and paper are all good. 

We believe that this very useful and creditable volume will be 
the means of giving a renewed and stronger impulse to American 
mycology, and that those who are fortunate enough to obtain it 
will soon be demanding the means of obtaining a knowledge of 
other species which will attract their notice and which are no less 
desirable and available for food than those here described. 

CHARLES H. PEcK. 


Proceedings of the Club. 


WEDNESDAY EVENING, OCTOBER 30TH, 1894. 


The President in the chair and 34 persons present. 
The committee on admissions reported favorably on the nomi- 


Nations of Miss Annie E. Hamilton, Mrs. Archibald D. Russell, 
Dr. F. C. Stewart and Mrs. Theron G. Strong, and they were 
elected active members, the Secretary being directed to cast an 
affirmative ballot. 
The following announced papers were then presented : 
“Notes on the Morphology of the Leaves in Galium,” illus- 


trated by specimens and drawings, by Mr. A. A. Tyler. 


478 


«“ Remarks on the Luminous Moss, Schistostega osmundacea,” 
illustrated by specimens and drawings, by Elizabeth G. Britton. 

« Some adaptive structural Features in the Lichens,” illustrated 
by specimens and drawings, by Dr. Albert Schneider. (To be 
published in the December BULLETIN.) 


TuEsDAY EVENING, NOVEMBER 12TH, 1895. 


Vice-President Lighthipe in the chair and 37 persons present. 

Dr. Britton nominated for active membership Mr. B. Heritage, 
of Mickleton, Gloucester county, N. J. 

Prof. Emily L. Gregory presented a paper on “ Theories of the 
Origin and Nature of the Starch Grain,” giving the history of the 
subject from the work of Naegeli to that of Meyer, who has re- 
cently submitted evidence that the substance of the starch grain 
is truly crystalline and not organized, so that all our theories of 
the growth of organized substance based upon the starch grain as 
a type fall, and we must begin to study the subject de ovo. 

The Secretary exhibited an ear of corn showing some perfect 
kernels of sugar corn distributed among those of yellow flint, as 
well as many kernels of an intermediate character. 

Mr. E. S. Miller remarked upon the blooming of certain Cac- 
taceae in cultivation, notably Cereus Thurberi, the plant being 
only a foot or two high and the flowers very small. 


Index to recent Literature relating to American Botany. 
Arthur, J.C. Development of Vegetable Physiology. Science (II.) 
2: 360-373. 20S. 1895. 


Arthur, J. C. Deviation in Development due to the Use of unripe 
Seeds. Am. Nat. 29: 804-815. S. 1895. 


Arthur, J.C. Potato Scab and its Prevention. Bull. Ind. Exp. Sta. 
56: pp. 11. Au. 1895. 


Bain, S. M. Notes on Utricularia inflata. Asa Gray Bull. 3: 4 
Ja. 1895. 


Barnes, C. R. The Botanical Society of America. Science (Il.) 2: 
448, 449. 40. 1895. 


Beardslee, H.C. Occurrence of Glacial Plants in Ohio. Ann. Rep” 
Ohio Acad. Sci. 3: 17. 1895. 


|. see 


479 


Bessey, C. E. Ninth Annual Report of the Botanist of the Nebraska 
State Board of Agriculture. Ann. Rep. Neb. Bd. Agric. 1895: 107— 
131. 27 Je. 1895. 

Bolley, H. L. Proceedings of the Botanical Club, A. A. A. S. 
Springfield meeting, August 29th to September 2d, 1895. Science 
Prt.) 2: 445-448, 4 O. 1895. 

Britten, J. American Nomenclature. Journ. of Bot. 33: 212-216. 
ae Sos. 281-282. 5. 1895. 

Burnham, S. H. The Ericacex in California. Asa Gray Bull. 3: 
18-20. Ap. 1895. 

Chodat,R. Polygalaceae novae vel parum cognitae.—IV. Bull. Herb. 
Boiss. 3: 538-549. O. 1895. 


New South and Central American species in Monnina, Securidaca and Polygala, 
Christ, H. Zrichomanes orbiculare n. sp. Beibl. Engl. Bot. Jahrb. 
1g: 25. 21 Au. 1894. 


A Brazilian species, 

Coe, C.H. The Floating Gardens of Mexico. Gard & For. 8: 432. 
30 O. 1895. 

Cook, O. F. Personal Nomenclature in the Myxomycetes. Bull. Torr. 
Bot. Club, 22: 431-434. 31 O. 1895. 

Cooley, G. E. On the Reserve Cellulose of the seeds of Liliaceae 
and of some related Orders. Mem. Bost. Soc. Nat. Hist. 5: 1-30. p/. 
Z~6. Jl. 1895. 

Davidson, A. Botanical Excursion to Antelope Valley. Erythea, 3: 
£53-158. ‘1 N. 1896. 

Dewey, L. H. Weeds, and how to kill them. U. S. Dept. Agric. 
Farmers’ Bull. 28: pp. 31.f. ZZ. 1895. 

Dietel, P. Zur Kenntiss der Gattung Uredinopsis Magnus. 


Deutsch. Bot. Gesellsch. 13: 326-332. 28 Au. 1895. 
Uredinopsis Preridis Diet. & Holw. n. sp., from California. 


Dodge, C.K. Helianthus tuberosus in Eastern Mic 
Bull. 3: 17. Ap. 1995. 

Eastwood, A. Observations on the Habits of Vemophila. 
3: 151-155. pl. 37. 1 N. 1895. 

Ellis, J. B., and Everhart, B. M. New Species of Fungi. Bull. 


Torr. Bot. Club, 22: 434-440. 31 O. 1895. ae 
New species from Sandwich Islands, Florida and Mexico, in Asteridium, Septoria, 


Ber. 


higan. Asa Gray 


Erythea, 


Dar luca, Colletotrichum, Cercospora, Isariopsis, Uredo, Schizophyllum, Rosel. 
a Ania, Didymosphaeria, Phyllachora, Haplosporelia and Melogramma, 


480 a. 
Engler, A. Ueber die Entwicklung des Canaigre. Notizb. bot. Gart. 
Berlin, r: 37, 38.- 5 Je. 1895. 


Note on Rumex hymenosepalus. 


Galloway, B. T. The Springfield Meeting of the American Associa- 
tion for the Advancement of Science, Section G., Botany. Science 
(II.) 2: 402-406. 27S. 1895. 


Galloway, B. T., and Woods, A. F. Water as a Factor in the 


Growth of Plants. Yearbook U. S. Dept. Agric. 1894: 165-176. /. 72- 
5: “Au, 1895. 


Gilg, E. Ueber die Bliithenverhaltnisse der Gentianaceengattungen 
° Hockinia Gardn. und Halenia Borckh. Ber. Deutsch. Bot. Gesellsch. 
13: 114-126. p/. 17. 25 Ap. 1895. 


South American forms considered, 


Gray, A., Watson, S., and Robinson, B. L. Synoptical Flora of 


North America. Large 8vo. 1: part 1: fascicler. 10 O. 1895. 
Ranunculaceae to Frankeniaceae, 


Gurke. Pavonia Schwackei. n. sp. Notizb. bot. Gart. Berlin, 1: 15, 
19. .2 Ja. 1895. 
Native of Brazil. 


Hegelmaier, F. Systematische Uebersicht der Lemnaceen. Engl. 
Bot. Jahrb. 21: 268-305. 6 Au. 1895. 


Hennings, P. Fungi Goyazenses. Hedwigia, 34: 88-112. 8 Ap. 
113-116. 19 Je. 1895. 


Describes sundry new species from Brazil collecied by E. Ule. 


Higgins, J. Hibernating forms of Phlox. Asa Gray Bull. 3: 35-37- 
~ O. 1895. 


Hill, E. J. Notes on Western New York Woodlands—I.—II. Gard. & 
For. 8: 342. 28 Au. 382-385. 25 S. 1895. 

Hitchcock, A. S. Note on Buffalo-gaass. Bot Gaz 
O. 1895. 


Hooker, J.D. Anthurium Gustavi Curt. Bot 
O. 1895. 


Native of New Grenada. 


“90: 464. 17 


» Mage 5rs: Pee 7437- 


Hooker, J. D. Mormodes Rolfeanum. Curt. Bot. Mag. 51: p/. 7436: 
O. 1895. 


Native of Peru. 


Huth, E. Monographie der Gattung Dedphinium. _— Bot. Jahrb. 
20: 322-499. p/. 6-8. 9 Ap. 1895. 


- 481 


Kellerman, W. A. The Nomenclature Question: Some points to be 
emphacized in the discussion. Bot. Gaz. 20: 468-470. 170.1895. 

Lagerheim, G. Monographie der Ecuadorianischer Arten der Gat- 
tung Brugmansia Pers. Engl. Bot. Jahrb. 20: 665~668. A/. rz. 28 
Je. 1895. 

MacDougal, D. T. Botanic Gardens. Minnesota Mag. 2: Part 1, 
pp. 16. [Reprint]. 

MacMillan, C. Current Problems in Plant Morphology—II. The 
Influence of Spray and Rain on the Forms of Leaves. Science (II. ) 
2: 481,482. 11 O. 1895. 

MacMillan, C. Current Problems in Plant Morphology—I. The 
Question of Pteridophyte Phylogeny. Science (II.) 2: 183,184. 16 
Au. 1895. 

Meehan, T. Coreopsis grandifora. Meehans’ Month. 5: 201. 2. 
qI. N. 1895. 

Meehan, T.  Lygodium palmatum. Meehans’ Month. 5: 181. f/. 70. 
O. 1895. 

Meehan, T. On the Derivation of Linnaean specific Names. Bot. 
Gaz. 20: 461, 462. 17 O. 1895. 

Moffatt, W. L. The Russian Thistle in Chicago. Asa Gray Bull. 3: 
12,13. Ap. 1895. 

Muller, F. Die Untergattung Widulariopsis Mez. Ber. Deutsch Bot. 
Gesell. 13: 155-164. AZ. 25. 26 Ap. 1895. 

Nash, G. V. New or noteworthy American Grasses—I. Bull. Torr. 
Bot. Club, 22: 419-424. 31 O. 1895. 


New species in Erianthus, Panicum, Stipa and Chloris. 

Newcombe, F.C. The regulatory Formation of mechanical Tissue. 
Bot. Gaz. 20: 441-448. 17 O. 1895. 

Norton, J.B. S. Ustlago Reiliana on Corn. Bot. Gaz. 20: 463. 
17 O. 1895. 

Pammel, L. H. Poisoning from Cowbane. Bull. Ia. Exp. Sta. 28: 
215—228. f. 7. S. 1895. 

Pammel, L. H. Two noxious Weeds. Bull. Ia. Exp. Sta. 28: 199— 
214.7. 8 S. 1895. 


Prickly Lettuce (Zactuca Scariola) and Buffalo-Bur (Solanum rostratum) are 
considered. 


Pieters, A.J. The History of the Uredineae. Asa Gray Bull. 3: 8- 
to, Ja. 1895. 


482 


Pollard, C. L. Nomenclature at the Springfield Meeting. Erythea, 
3: 158-161. 1 N. 1895. 

‘Putnam, B. L. A day-blooming Cereus grandifiorus. Bot. Gaz. 20: 
462,463. 17 O. 1895. 

Rimbach, A. Jahresperiode tropisch-andiner Zwiebelpflanzen. Ber. 
Deutsch. Bot. Gesell. 13: 88-93. 25 Ap. 1895. 

Ritter, F. W. EZriodictyon glutinosum. Am. Journ. Pharm. 67: 565- 
572. f. Ta. N, 1895. 

Rompel, J. Drei Carpelle bei einer Umbellifere (Cryptotaenia Cana- 
densis). Odecsterr. Bot. Zeitschr. 45: 334-337. S. 1895. 

Sargent, C. S. Kalmia cuneata. Gard. & For. 8: 434. f/f. 60. 
30 O. 1895. 

Sargent, C. S. Rhus Michauxit. Gard. & For. 8: 404. f. 55- 9 
O. 1895. 

Sargent, C. S., Editor. Pinus ponderosa. Gard. & For. 8: 392. 
S: $4. ‘2 O. 1895. ; 

Schinn, C. H. Observations in California Forests. Gard. & For. 8: 
402. 9 O. 1805. 

Scribner, F. L. Agrostology in the Department of Agriculture. 
Science (Il.) 2: 151, 152. g Au. 1895. 

Searing, A. H. The Flora of Long Pond. Proc, Rochester Acad. 
Sci. 2: 297-300. 1805. 

Seemen, O. von. Fiinf neue Weidenarten in dem Herbar des KGénig- 
lichen Botanischen Museums zu Berlin. Beibl. Engl. bot. Jahrb. 21: 


6-11. 6 Au. 1895. 
New species of American willows. 


Setchell, W. A. Notes on some Cyanophyceae of New England. 
Bull. Torr. Bot. Club, 22: 424~431. 

New species in Revularia and Arthrospira. 

Small, J. K. Studies in the Botany of the southeastern United States. 
—IV. Bull. Torr. Bot. Club, 22: 365-369. p/. 2g6. 31 Au. 1895. 
Describes new species in Sieglingia, Rumex and Acer, 

Small, J. K. Teratological Notes. Bull. Torr. Bot. Club, 22: 399, 
400. figs. 30S. 1895. 

Summers, E. California’s Flora. Asa Gray Bull. 3: 40-43- O. 
1895. 

Swingle, W. T. The Grain Smuts; their causes and prevention. 
Year-book U.S. Dept. Agric. 1894: 409-420. f. 92-99. Au. 1895. 


483 


Tchouproff, O. Quelques notes sur I’anatomie systématique des 
Acanthacées. Bull. Herb. Boiss. 3: 550-560. O. 1895. 

Thaxter, R. Newor peculiar aquatic Fungi.—I. Bot. Gaz. 20: 433- 
440. Pf. 29. 17 O. 1895. : 

Two species of A/onodlepharis. 

Tilden, J. E. A Contribution to the Bibliography of American Algae 
Bull. Geol. Nat. Hist. Survey Minnesota, 9: 295-419. [Minn. Bot. 
Stud. 1]. 26 Au. 1895. 

Tracy, S. M. and Earle, F.S. Mississippi Fungi. Bull. Miss. Exp. 
Sta. 34: pp. 44. My. 1895. 


Enumerates 353 species collected, of which 55 are new. 

Trelease, W. The Pignuts. Bull. Torr. Bot. Club, 22: 331. 31 
Jl. 1895. Bot. Gaz. 20: 373. 15 Au. 1895. 

True, R. H. On the Influence of sudden Changes of Turgor and of 

‘ Temperature on Growth. Ann. Bot. Q: 365-402. 5S. 1895. 

Uline, E. B. and Bray, W. L. Synopsis of North American Amar- 
anthaceae—IV. Bot. Gaz. 20: 449-453. 17 O. 1895. 


Alternanthera Kerberi, from Mexico, described as new. 
Urban, J. Diagnosen westindischer Arten. Notizb. bot. Gart. Berlin, 
I: 78-80. 5 Je. 1885. 


New Jamaican species in Maytenus, Myrsine, Ardisia, Macrocarpaea, Cordia 
and Saracha, and Mosquitoxylum, new genus of Asclepiadaceae. 


Urban, J. Jacques Samuel Blanchet. Beibl. Engl. Bot. Jahrb. 21: 
I-5. 6 Au. 1895. 

Urban, I. Juglans Jamaicensis C. DC. Notizb. bot. Gart. Berlin, 1: 
29-31. figs. 2 Ja. 1895. 

Urban, I. Ueber die Sabiaceengattung MJeliosma. Ber. Deutsch. 
Bot. Gesell. 13: 211, 222. f/. 19. 26 Je. 1895. 

Ward, L. F. Fossil Plants. Johnson’s Univ. Cycl. 6: 639-645. 
1895. 

Waters, C. E. An Analytical Key for our local Ferns based on the 
Stipes. Johns Hopkins Univ. Circ. 14; 74, 75- Je. 1875. [Also 


reprinted. 
Includes 35 species growing in the vicinity of Baltimore, Maryland. 


Webber, H. J. Fertilization of Soil as affecting the Orange in Health 
and Disease. Year-book U.S. Dept. Agric. 1894: 191-202. /. 7 8-19. 
Au. 1895. 

White, T. G. Notes on the Orchidaceae of Mt. Desert, Maine. Asa 
Gray Bull. 3: 44-46. O. 1895. 


484 


Wiegand, K. McK. The Structure of the Fruit in the Order Ranun- 
culaceae. Proc. Am. Micros. Soc. 1894: 69-100. p/. 7-8. 1894. 


Williams, T. A. Native Trees and Shrubs of South Dakota. Bull. 
S. Dak. Exp. Sta. 43: pp. 32. My. 1895. 
Briefly describes 117 species. 
Wilson, F. Variations in Viola. Asa Gray Bull. 3: 13. Ap. 1895. 
Wolf, T. Die Galapagos-Inseln. Verhandl. d. Gesellsch. f. Erdk. z. - 
Berlin, 22: 246-265. p/. 37. 1895. 
Contains references to the flora. 
Zahlbruckner, A. Lobeliaceae Brasilienses. Medd. Nat. Foren. 
Kobenh. 1895: 67-71. 1895. 
Enumeration of the species collected by Dr. Glaziou; Lobelia Glazioviana is de- 
scribed as new. 
Zeissold, H. Kakteen Neuheiten. Monats. Kakteenk. 5: 69-75 ; 
86-90. My. Je. 1895. 


Illustrations are given of Cereus Eruca, C. Cochal, Pilocereus Sargentianus and 
Mamillaria Halei, 


Contributions from the Herbarium of Columbia College. 


No. on. 
No. 39. 


No, 40. 


No. 4. 


{The numbers omitted from this list are out of print. ] 


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TORREY BOTANICAL CLUB. 


New Species of Fungi. 


By Cuas. H. PEcK. 


AMANITOPSIS VELOSA. 


Pileus at first subglobose, then campanulate or nearly plane, 
generally bearing patches of the remains of the whitish felty or 
tomentose volva, elsewhere glabrous, becoming sulcate-striate on 
the margin, buff or orange-buff, flesh compact, white; lamellae 
close, reaching the stem, subventricose, pale cream color; stem 
firm, at first tomentose and attenuated at the top, then nearly 
equal, stuffed, white or whitish, closely sheathed at the base by 
the thick volva; spores globose, .0004 to .0005 in. broad. 

Pileus 2 to 4 in. broad ; stem 3 to 4 in. long, 3 to 4 lines thick. 


Under oak trees. Pasadena, California. April. A. J. Mc- 
Clatchie. 

This fungus is closely related to Amanifopsts vaginata, from 
which it may be separated by the more adherent remains of the 
thicker volva which sometimes cover the whole surface of the 
pileus, and by the thicker lamellae which are somewhat adnate to 


the stem and terminate with a decurrent tooth. 


TRICHOLOMA SEMIVESTITUM. 


Pileus thin, expanded, centrally depressed or subumbilicate, 
glabrous, blackish-brown, the margin decurved or involute ; lamel- 
_lae close, emarginate, often dentate or eroded on the edge, whitish 
tinged with blue; stem short, solid, usually slightly thickened at 
the base, the lower part clothed with a tawny-brown tomentum, 
the upper part brown; spores broadly elliptical or subglobose, — 
uninucleate, .00016 to .0002 in. long, .00016 broad. 


* 486 


Pileus 6 to 12 lines broad; stem 8 to 12 lines long, 2 to 3 lines 
thick. 

“On old grass roots in sandy prairie pasture,’ Rooks County, 
Kansas. July. E. Bartholomew. 

The small size, dark-brown pileus and semitomentose stem are 
the prominent characters in this species. 


COLLYBIA MICROSPORA. 


Pileus thin, convex, subumbonate, glabrous, white; lamellae 
broad, subdistant, nearly plane, white, changing to brown in dry- 
ing; stem slender, hollow, glabrous, white, becoming brown in 


drying; spores minute, broadly elliptical, .ooo16 in. long, .00O012 
broad. 


Pileus about 6 lines broad; stem about I in. long, 1 line thick. 

Wet ground under bushes. Rooks county, Kansas. July. 
Bartholomew. 

This plant, in the dried state, might easily be taken for some 
species of Vaucoria, since the lamellae assume a rusty-brown hue. 
This change of color in the lamellae and stem is a peculiar feature 


of this small species which the notes of the collector record as 
“ pure white throughout.” 


MYCENA CAESIA. 

Pileus submembranous, campanulate, striate, glabrous, grayish- 
brown, blackish or blackish-brown in the center; lamellae thin, 
subventricose, grayish or bluish-gray; stem slender, hollow, glab- 
rous, grayish-brown; spores subelliptical, slightly apiculate at one 
end, .0003 to .0004 in. long, .00016 to .0002 broad. 

_ Pileus 4 to 6 lines broad ;.stem I to 2 in. long,.5 to 1 line thick. 

Among sphagnum. Newfoundland. September. Rev. A. C. 
Waghorne. 

I have seen dried specimens only and am not certain that the 
bluish-gray hue of the lamellae is so conspicuously present in the 
fresh plant. The margin of the pileus is sometimes tinged with 
yellowish-brown. 


HyYGROPHORUS SPHAEROSPORUS. 


Pileus fleshy and thick in the center, subobconic, convex, ob- 
tuse or slightly umbonate, whitish, inclining to reddish-brown, the » 
_ margin incurved, flesh firm, white; lamellae rather broad, subdis- 

_ tant, adnate or slightly decurrent, white; stems tufted, flexuous, 


487 


solid, glabrous, often slightly thickened at the base, colored like 
the pileus ; spores globose, .00024 to .0003 in. broad. 

Pileus 6 to 12 lines broad; stem 1 to 2 in. long, 2 to 3 lines 
thick. 

Iowa. October. Communicated by C. Mcllvaine. 

The fresh plant is said to have no decided odor, but when 
partly dried it emits a slight but rather unpleasant odor. It be- 
longs apparently to the Section Camarophyllus, and is related to 
flygrophorus Peckit. 


MARASMIUS SUBTOMENTOSUS. 

Pileus thin, at first subcampanulate, then convex or nearly 
plane, even or slightly striate on the margin, minutely tomentose- 
pubescent, grayish or reddish gray; lamellae broad, subdistant, 
free or but slightly adnexed, ventricose, colored like the pileus ; 
stem equal or slightly swollen toward the base, inserted, velvety- 
tomentose, grayish or grayish-brown, white within; spores .0004 
to .0005 in. long, .0002 to .00025 broad. 

Pileus 5 to 9 lines broad; stem about 1 in. long, 1 line 
thick. 

Abundant on roots of grasses and other plants in sandy soil. 
Kansas. July. Bartholomew. 

The minute tomentum of both pileus and stem which gives 
them a grayish hue, the inserted stem and the free subdistant 


lamellae mark the species. 


MARASMIUS BADIUS. 


Pileus thin, convex, even, glabrous, bay-brown or reddish- 
brown; lamellae narrow, subdistant, adnate, whitish ; stem gla- 
brous, hollow, blackish-brown; spores broadly elliptical, .oo02 in. 


long, .ooo12 broad. 
Pileus 3 to 6 lines broad; stem about 1 in. long, 1 line thick. 


Decaying sticks lying on wet ground. Kansas. July. Bar- 


tholomew. 
In the dried specimens the stem is of a darker hue than the 


Pileus. This gives a peculiar appearance to the plant and makes 


the species easily recognizable. 


VOLVARIA STRIATULA. 


Pileus thin, convex or nearly plane, minutely silky, striate on 
the margin and somewhat reticulate when dry, white; lamellae 


5 


488 


narrow, free, white, becoming flesh color; stem short, glabrous, 
white, with the cup-like remains of the membranous volva at the 


base; spores subglobose, uninucleate, .0003 in. long, nearly as 
broad. 


Pileus 6 to 9 lines broad; stem about 1 in. long, .5 to I line 
thick. 

Wet ground under weeds. Kansas. July. Bartholomew. 

This species is allied to V. parvula, from which it is separated 
by the striate margin of the pileus and the larger and nearly 
globose spores. In the dried specimens there are transverse ridges 
or wrinkles between the marginal striations which give a reticu- 
late appearance. 


PLUTEUS STERCORARIUS. 


Pileus very thin, nearly plane, glabrous, pure white: lamellae 
rather broad, close, free, flesh color; stem equal, solid, glabrous, 
white ; spores large, even, commonly uninucleate, .0005 to .0006 
in. long, .0003 broad. 


Pileus about 2.5 in. broad; stem 3 in. long, 3 lines thick. 

Manure heaps. Kansas, July. Bartholomew. 

The habitat of this species is peculiar. Most of the species of 
this genus grow on decaying wood and have smaller spores. 


INOCYBE RADIATA. 


Pileus thin, convex or subcampanulate, distinctly umbonate, 
silky-fibrillose, slightly rimulose, distantly radiately wrinkled when 
dry, yellowish-brown, the umbo commonly blackish-brown ;_ la- 
mellae rather broad, close, emarginate, brownish becoming tawny- 
cinnamon when old, whitish on the edge; stem equal, solid, sub- 

glabrous, a little paler than the pileus ; spores subovate, slightly 
nodose or angular, .0004 to .0005 in. long, .0002 to .00025 broad. 


Pileus about 1 in, broad ; stem I to 2 in. long, 1 to 2 lines 
thick. 

In open grassy ground. Massachusetts. Aug. and Sept. 
W. D. Jackson. 

The species belongs to the Rimosae. In general appearance 
it is similar to /. fuscodisca, I. infida and I. agglutinata, but the 
radiating ridges of the dried pileus and especially the peculiar 
spores easily separate it from any of these. Its spores are some- 
what variable. Some are nearly even, others have one or two 
nodes, and many of them are irregular or angular like the spores 


489 


of species of Enxtoloma. Some are nearly elliptical in outline, but 
generally they are narrowed toward one end. They are mostly 
‘one or two-nucleate. The radiations of the pileus are not notice- 
able in the fresh plant. 


FLAMMULA DECURRENS. 


Pileus thin, umbilicate centrally depressed or funnelform, 
Moist, minutely floccose-squamulose, pale yellow or cream color ; 
lamellae subdistant, strongly decurrent, pale yellow becoming 
ochraceous, the interspaces sometimes veiny ; stem equal or taper- 
ing downward, minutely downy, stuffed with a cottony pith, 
‘colored like the pileus; spores elliptical, .0003 to .00035 in. long, 
-00016 broad. 

Pileus about 1 in. broad; stem 6 to 10 lines long, 1 to 2 thick. 

Wet ground in shade of bushes. Kansas. July. Bartholo- 


mew. . 

The species is easily known by its color being a uniform rich 
‘creamy or sulphury hue in the fresh plant, and by its strongly de- 
current lamellae which become ochraceous-yellow in the dried 


State. 
CREPIDOTUS CINNABARINUS. 

Pileus thin, sessile, resupinate or reflexed, minutely tomentose 
Or pulverulent, cinnabar-red; lamellae rather broad, subdistant, 
Minutely reddish-flocculent on the edge, brownish-tawny in the 
‘dried plant; spores broadly elliptical, .0003 in. long, .00024 to 
-00028 broad. 

Pileus 3 to 4 lines broad. 


Decaying wood. Michigan. Sept. L. N. Johnson. 


STROPHARIA C/ESIFOLIA. 


Pileus convex, glabrous, white or whitish, sometimes brownish 
in the center; lamellae close, rounded or emarginate behind, light 
blue, becoming dingy bluish-brown ; stem equal or slightly thick- 
ened at the base, solid, glabrous, white or whitish, the annulus 
white; spores subelliptical, .0004 to .0005 in. long, .00024 to 
-0003 broad. 

Pileus 1 to 2 in. broad; stem 1 to 1.5 long, 2 to 3 lines thick. 

Low sandy pastures. Kansas. July. Bartholomew. — 

The collector of this plant remarks that it is much like the 
common mushroom, Agaricus campester, except that its lamella 
have a fine light blue color instead of pink. This is an unusual 


490 


and very distinctive character. In the dried specimens the 
lamellae have assumed a dingy grayish-blue hue, inclining to 
brown. 

HyYPHOLOMA CUTIFRACTA 

Pileus thin, campanulate or expanded, sometimes faintly striate 
on the margin, grayish-buff, the disk sometimes darker, the cuticle 
commonly irregularly rimose ; lamellae thin, close, adnate, at first 
white or whitish, then rosy-brown; stem long, slender, hollow, 
glabrous, white; spores elliptical, brown, .0003 in. long, .0002 
broad. 

Pileus I to 2 in. broad; stem 3 to 4 in. long, I to 2 lines thick.. 

About stumps of Lombardy poplar. Kansas. July.  Bar- 
tholomew. 

This species is closely related to H. zncertum, but differs in its 
long slender stem and in its cuticle cracking in an irregular man- 
ner and revealing the thin white flesh beneath. No remains of 
the veil are visible in the dried specimens. Sometimes the mar- 
gin of the pileus is deeply split. 


PSATHYRELLA LEUCOSTIGMA. 


Pileus submembranous, campanulate, striate, bluish-white whem 
fresh, changing to sepia-brown when dried, the apex remaining 
whitish ; lamellae close, lead color when young, becoming black 
with age, whitish on the edge; stem slender, flexuous, hollow, 
white ; spores black, elliptical, .0005 to .0006 in. long, .0003. 
broad. 

Pileus 4 to 6 lines broad; stem 1 to 1.5 in. long, about 1 line: 
thick. 

Wet ground under trees. Kansas. July. Bartholomew. 

This pretty little species is well marked in the dried state by 


the white apical spot on the pileus. The plant is probably hygro- 
phanous in the fresh state. esa 


PSATHYRELLA BARTHOLOMAEI. 


; Pileus thin, subconical or convex, glabrous, striate on the mar- 
gin, pale brown; lamellae close, nearly plane, adnate, brownish 
becoming black; stem slender, flexuous, hollow, adorned with a 
few grayish fibrils, pale brown; spores elliptical, .0004 to .0005 in- 
long, .0002 to .00025 broad. 


Pileus 6 to 12 lines broad; stem 1 to 1.5 in. long, scarcely 
line thick. Re sh 


491 


“ Rich ground in shade of trees.” Kansas. July. Bartholo- | 
mew. 


COPRINUS EBULBOSUS. 


__ Pileus thin, campanulate, variegated by the cuticle breaking 
into broad superficial persistent whitish scales, the surface beneath 
the cuticle somewhat striate, grayish-brown, the margin at length 
revolute, lacerated; lamellae narrow, thin, crowded, free, slate 
color becoming black; stem equal, hollow, white; spores ellipti- 
cal, .0003 to .0004 in. long, .0002 broad. 


Pileus 2 to 3 in. broad; stem 3 to 6 in. long, 2 to 3 lines 
thick. | 

Caespitose at the base of cottonwood stumps. Kansas. July. 
Bartholomew. 

This plant resembles C. picaceous very closely. New York 
specimens were formerly referred to it as variety edalbosus, but 
having now received it from various widely separated localities 
and finding that it maintains its distinctive characters with con- 

__ Stancy, it seems best to consider it a good species. Its peculiar 
characters are the absence of a bulbous base to the stem and its 
smaller spores. It also sometimes grows in large tufts. “ About 
fifty grew in a solid clump, all united at the base.” 


COPRINUS LANIGER. 


Pileus thin, conical or campanulate, covered when young with 
numerous tawny tomentose or floccose scales which partly or 
wholly disappear with age, sulcate-striate nearly to the apex, 
Pallid, tawny or grayish-ochraceous; lamellae crowded, at first 
whitish, then brownish-black; stem slightly thickened at the 
base, minutely downy or pruinose, hollow, white; spores oblong- 
elliptical, commonly uninucleate, .0003 to .0004 in. long, .00016 
broad. 

Pileus 6 to 12 lines broad; stem about 1 in. long, 1 to 2 lines 
thick. 

- Caespitose at the base of cottonwood stumps. Kansas. July. 
Bartholomew. 

The species resembles C. micaceus, from which it is distin- 

: guishable by the floccose-squamose coating of the young pileus 

and by its more narrow spores. Mr. Bartholomew remarks that 


“it is of slow growth, taking three or four days for development.” 


492. 


TTHELEPHORA SUBUNDULATA. 


Pileus thin, subcoriaceous, centrally depressed, plicate-undulate 
onthe margin, subcinereous or grayish-brown; hymenium slightly 
uneven, paler than the pileus, grayish or grayish-yellow; stem 
firm, solid, rarely branched, colored like the pileus; spores broadly 
elliptical, .0003 in. long, .0002 broad. 


Pileus about 6 lines broad; stem 8 to 10 lines long, scarcely I 
line thick. 

Ground. Delaware. July. A. Commons. 

This species appears to be related to, but much smaller than 
T. undulata. It differs from it in having a glabrous hymenium 
and larger spores. The stem is not polished, but tothe naked eye 
appears to be priunose-pubescent. Sometimes the margin of the 
pileus is more than wavy, its folds overlapping. 


SECOTIUM DECIPIENS. 


Peridium subglobose or depressed globose, its surface rupturing 
into rather broad, loose or appressed scales, cream-colored, the in- 
ferior part at first closely pressed to the stem and sometimes sep- 
arating from the upper part and forming a kind of annulus, some- 
times splitting longitudinally and gradually falling away, leaving 
the upper part in the form of a pileus; the glebe lamelliform, but 
variously united and anastomosing, forming irregular and some- 
what labyrinthiform cells, free from the percurrent stem, almost or 
quite black; stem stout, commonly tapering upward, abruptly nar- 
rowed as it enters the peridium, firm, solid, externally colored like 
the peridium, internally tinged with yellow or rhubarb color; 

spores globose or subglobose, even, black or brownish-black, 
0002 to .00028 in. long. 

Peridium 1 to 4 in. broad; stem 3 to 6 in. long, 1 in. or more 
thick at the base, 5 to 10 lines where it enters the peridium. 

Streets and lawns. Pasadena, California. April. McClatchie. 

The copious blackish spores are an unusual feature in the genus 
Secotium. The lamelliform septa sometimes rupture in such a 
way as to form coarse aculeiform processes as in the*genus Poly- 
plocium, and thus the plant might easily be referred to that genus. 
But [ find no filamentous capillitium in the specimens before me, 
and no evidence of a volva, and have therefore referred them to 
the genus Secotium, between which and Polyplocium this species 

forms a connecting link, 


er 


~ 


493 


MACROSPORIUM AMARANTHI. 


Hyphae short, .0008 to .0012 in. long, septate, slightly nodose 
at the top, forming minute punctiform blackish tufts; spores cla- 
vate oblong-clavate or subfusiform, 3- to 8-septate, one to three 
of the cells with longitudinal septa, .oo12 to .0024 in. long, 
exclusive of the slender pedicel which is commonly shorter than 
the spore. 

Dead spots of leaves of Amaranthus retroflexus. Kansas. 


August. Bartholomew. 
The fungus appears to the naked eye to form brownish 


patches on the spots. 


MACROSPORIUM CLEMATIDIS. 


Spots small, 2 to 3 lines broad, suborbicular, whitish, gray or 
brownish, often with a more or less distinct brown border; 
hyphae amphigenous, caespitose or thinly effused, colored, septate, 
flexuous, .002 to 004 in. long, .00025 to .0003 broad; spores 
obovate or subclavate, 3- to 4-septate, commonly constricted at 
the septa, .oo1 to .0016 in. long, .0005 broad, the pedicel nearly 
as long as the spore. 

Living or languishing leaves of Clematis Fremonit. 


September. Bartholomew. 


Kansas. 


HEYDENIA FUNGICOLA. 


Plant scarcely half aline high, black; receptacle at first ellip- 
tical, becoming hemispherical truncate or disciform above and 
dusted with the spores, about .007 in. broad ; stem subcylindrical ; 
spores catenulate, globose or subglobose, colored, .00016 to 0002 
in. long. . 

On old specimens of Polyporus adietinus green with incrusting 


alge. Maine. Sept. F. L. Harvey. © oe 
This was found associated with Calicium tigillare which it 


_ Closely resembles. 


494 


Some special phylogenetic Adaptations in Lichens.—, 


By ALBERT SCHNEIDER. 


In a previous paper* I endeavored to show why lichens should 
be considered as a distinct class of plants. It is my purpose in 
this paper to discuss briefly some of the special adaptive features 
which these plants have acquired since their phylogeny as lichens. 
I shall preface this discussion with a few introductory statements. 

Everywhere we can observe a balanced relationship between 
living organisms and their environment. All structures, no mat- 
ter where they may occur, or what form they may assume, serve 
a definite purpose and_ perform a definite function. In many in- 
stances the structural conformations are so “rudimentary” and 
“imperfect” that we are unable to recognize their physiological 
significance. It is however unscientific to say that such structures 
perform no function, because we are unable to recognize a func- 
tion. It were much more consistent to admit our ignorance and 
await further investigations, which may clear up some of the diffi- 
culties. J make these preliminary statements with special refer- 
ence to lichens, because many of their structural adaptations have 
as yet not been satisfactorily explained. I also wish to call at- 
tention to the importance and special biologic significance of the 
tendency among modern scientific botanists to point out the in- 
terrelation of structure‘and function. There is no function with- 
out structure: without structure there could be no function. 

From this it becomes evident that morphological and physio- 
logical investigations must go hand in hand. If we neglect one 
for the other we fall into a dangerous error, dangerous to the pro- 
gress of biological science. The time is fortunately slowly pass- 
ing away when morphology (usually external morphology) alone 
constitutes the science of life. The herbalists who go into wild 
ecstacies over having “ discovered” a “new species” are slowly 
giving way to the scientists who are solving or endeavoring to 
solve some difficult life-problem. The fact that many scientific 
efforts are fruitless in direct results is not a reason for becoming 


* The Biological Status of Lichens, BULLETIN, 22: 189-198. 1895. 


495 


discouraged ; much less a reason why old fashioned “ empiricists ” 
should gloat over these failures and make strenuous efforts to per- 
petuate the old regime. 

The knowledge gained from a purely morphological study of 
lichens, is in itself of little practical value if we can not find the 
correlated physiological interpretation. The scientific study of 
lichens is of very recent origin. Our knowledge of the physiology 
of these plants is especially deficient. Sufficient is, however, 
known in a general way, to enable us to give fairly accurate 
hypothetical physiological interpretations of most of the anatom- 
ical structures. 

It is perfectly consistent to suppose that the lichen, during 
phylogenetic history, has undergone great adaptive changes. This 
statement is axiomatic and requires no further elucidation. From 
the nature of things it is also evident that the thallus has under- 
gone the greatest adaptive changes. 

As has been known for some time, the thallus in particular per- 
forms the function of assimilation, due to the presence of the sym- 
biotic algae. Within recent years, Jumelle * has made a special 
Study of chlorophyllian assimilation in lichens. The special 
structural adaptations which we are about to discuss are due to 
the fact that the fungal symbiont, which is incapable of assimi- 
lating CO,, must make suitable concessions to the position and ar- 
rangement to the algal symbiont. This we find to be the case, 
In this paper I shall devote myself to foliaceous thalli and shall 
_ briefly consider some of the typical structures met with in such 
thalli; namely, the epidermal layer, the upper cortical layer, the 
algal (“gonidial” ) layer, the medullary layer, the lower corti- 
cal layer and the rhizoids. A typical epidermal layer is not 
Present in all foliaceous lichens. It is, however, well devel- 
Oped in Svcta and Stictina. It consists of hyphal cells, two 
or three layers in thickness. The cells are placed horizontal'y 
and are very closely crowded, thus forming a protection against 
excessive evaporation. It serves a function similar to eS ce 
dermis in higher plants. In the genus Lepéogium the epider- 


mis consists of a single layer of cells presenting a close resem- 


ia See pena : le de Bo 
*Jumalle H. Recherches Physiologiques sur les Lichens. Revue generale de Bo. 


_— tanique, 4: 49-64, 103-121, 159-175) 220-231, 259-272, 305-320. 1892 


496 


blance to the cuticle of higher plants. We find a similar cuticular 
covering in the related genera, Mallotium, Hydrothyria and Poly- 
chidium. This structure not only serves the function of an epi- 
dermal layer, that is, prevents excessive evaporation, but also a 
mechanical function, giving greater firmness to the thallus. In 
other words, the single layer of cells in the above genera of the 
Collemaceae serves a function similar to the many-layered epider- 
mis and cortical structure in S¢tcfa, Stictina and other genera. 

The epidermal layer is generally colored, usually a yellowish 
brown or dark brown. This is no doubt for the purpose of tem- 
pering the influence of sunlight. We may safely assume that the 
epidermal layer serves three functions. Its primary function is to 
prevent the excessive evaporation of moisture; its secondary func- 
tions are to reduce the injurious effect that direct isolation would 
have upon the symbiotic algae and to aid in protecting the under- 
lying tissues as well as to give additional firmness to the thallus.* 
Below the epidermis is found the cortical layer, which is usually of 
considerable thickness in the majority of foliaceous lichens. I will 
not dwell upon its anatomical structure, as that is sufficiently well 
known. Its primary function is mechanical. It also serves a 
function similar to that of the epidermis; it prevents the evapora- . 
tion of moisture. 

According to the principles of mechanics and for the purpose 
of protection the cortical layer should occur near the upper surface 
of the thallus. But for physiological reasons the algal layer 
should also be near the upper surface, that the algae may be acted 
upon by the sunlight and that they may more readily come in_ 
contact with the CO, of the atmosphere and that they may readily 
give up the O which is liberated as the result of assimilation. We 
actually find such an adaptive relationship between the algal 
layer and the cortical layer. In many of the foliaceous lichens 
we find that the algae extend almost to the upper surface of cer- 
tain circumscribed areas; that is the cortical layer is not of 
uniform thickness. Such an arrangement enables the algae 
to take up CO, from the atmosphere in exchange for the O 


*It should be borne in mind that there is no living structure whose function is 


purely mechanical or purely physiological. «Dead ” structures may have only a me- 
chanical function. 


497 


liberated. This could, however, not take place, if the epidermis 
and the upper and lower cortical layers were unbroken or without 
intercellular spaces. I shall now discuss some of the structural 
adaptations to meet this requirement. 

On the closer examination of a thin vertical section of the 
thallus, it is found that in many of the foliaceous lichens numerous 
intercellular canals (intercellular spaces) pass from the algal layer 
through the cortical and epidermal layers. These canals, which 
resemble the stomata in their physiological function, facilitate the 
interchange of the gases resulting from the activities of chloro- 
phyllian assimilation. The canals (breathing pores) are much 
branched and occur most frequently in the thin areas of the upper 
cortical layer. Asa rule, they do not pass to the exterior in a 
vertical direction. In the epidermal layer they take almost a hort- 
zontal course; this is because the cells of the epidermis are elon- 
gated horizontally. 

In the dry state these canals are practically closed, thus reducing 
the loss of moisture to a minimum. In the moist state they en- 
large considerably, thus enabling assimilation, which is dependent 
upon the presence of CO, and sunlight, to take place. These 
breathing pores are especially numerous in Mephromium, Solorina 
and Parmelia. 

Other lichen genera do not have the breathing pores just 
Mentioned. For example they are not noticable in the genera 
Sticta and Stictina. In these two genera the epidermal layer is 
distinct, as has been stated. The upper cortical layer is of uniform 
thickness and consists of very compact, rather small hyphal cells. 
The lower cortical layer is comparatively thin and its cells are less 
closely united; yet they are sufficiently compact that no inter- 
cellular spaces can be detected. The question now arises, how are 
the enclosed algae * supplied with the necessary atmosphere? Inmy 


opinion this is done by means of the so-called cyphellae which 


Occur on the lower surface of the thallus. These structures have 


long been known to lichenologists. Haller (1776) was perhaps 


the first author who called special attention to them. He de- 


nf! have purposely substituted the term “algae” for the term “ gonidia.” Go 


nidia and related terms as “ gonimia,” “ gonidimia,” etc., are meaningless in modern 
lichenology, and should therefore be rejected. 


498 ~ 


scribed them as ‘white circular depressions.” This was about 
all that was known concerning these structures at that time. 
Acharius was the first to introduce the term cyphellae, which term 
has been retained up to the present. Since these organs are little 
known, it will be well to describe them more in detail. 

The cyphellae occur almost exclusively in the genera Sécta 
and Stictina and are primarily breaks in the continuity of the 
lower cortical layer. The hyphae of the medullary layer immedi- 
ately about the opening in the cortical layer divide more fre- 
quently, producing a dense net-work of hyphae or evena semicor- 
tical structure. As a rule, the cells of this secondary cyphellar 
formation extend in a direction at right angles to the outer sur- 
face; that is, they assume a suitable position for the conduction 
of food-substances. The margin of the primary cortical layer (the 
cortical layer of the thallus) is turned more or less outward. As 
a rule the algae of the algal layer immediately over the cyphellar 
opening multiply more rapidly, this causing them to accumulate 
at these areas. 

Two kinds of cyphellae may be recognized. Form 1, as 
seen in Svictina damaccornis. The cyphellar depression is usually 
circular, concave inward, the margin of the cortex forming an 
outer constriction. The broken-down cortex of the thallus is 
replaced by the secondary semicortical formation of the medullary 
hyphae. Its outer surface is usually smooth, devoid of rhizoids, 
and generally of a paler color than the primary cortical tissue. 
They are irregularly distributed over the elevated portions of the 
lower surface of the thallus; none ever occur in the grooves. 
They appear first in the older portions of the thallus. In form 
they are quite constant, in size they decrease toward the younger 
portion of the thallus. : 

Form 2, as seen in the majority of Sé#ctas and Stictinas. It is 
much more common and differs from the former in the absence of 
the formation of a secondary semicortical cyphellar covering. 
Instead of a depression as in the former case, there is usually a 
protrusion of the densely interwoven network of medullary 
hyphae. Their form is usually less regular. Their position 
and arrangement is the same as in the first form. 

Both forms of cyphellae are sufficiently large to be seen by 


499 


the naked eye. No cyphellae occur near the actively growing tip 
of the thallus. 

Acharius applied the term cyphellae to the form first described. 
Those of the second form he considered to be soredia. Ny- 
lander retained the name cyphellae for the first form. The second 
form he designated as psuedocyphellae. There is no morphological 
or physiological reason why the latter should be designated as 
“false.” Indeed it were more consistent to designate the first 
form as false since they are of less frequent occurrence. Stitzen- 
berger* retains Nylander’s distinction into true and false cyphellae 
and further subdivides them as to color into white and yellow. 
This subdivision seems to be valueless, since their color depends 
upon the color of the medullary hyphae. Stitzenberger considers 
_ them of special importance in his classification of the Stictei. I 
am, however, opposed to adopting as primary, morphological 
characters whose physiology is unexplained. 

Having thus briefly treated of the morphology of cyphellae we 
will now refer to their probable physiological significance. As 
already stated, they very likely serve to admit air into the interior 
of the thallus. Some of the older lichenologists looked upon them 
as vegetative propagative organs, similar to the soredia. They 
are not soredia, since they usually contain no algae. That the 
second form may accidentally contain algae is possible, since we 
find occasional algae throughout the medullary layer. It is, how- 
€ver, unreasonable to suppose that they would normally contain 
algae, since their position is not suitable for the development of the 
Symbiotic algae. From the nature of the cortical layers and the 
epidermis in the majority of Stictei, it is reasonable to assume 
that the cyphellae serve a function similar to that of the stomata 
of ordinary foliage leaves. That is, the cyphellae and breathing 
Pores of lichens are functionally similar. If it is scientific to make 
any comparison of the morphology of lichens and phanerogams it 
may be stated that the cyphellae are analogous to lenticels.* The 
tissue which closes the opening in the lower cortical layer is the 
result of a special cambial or meristematic activity. 


beste 


*Stitzenberger, E. Die Gruebchenflechten (Stictei) und ihre geographische Ver- 


breitung. Flora, 81: 88-1 50. 1895. : 
> © This anal ne was pointed out by Dr. Smith Ely Jelliffe at a recent meeting of the 


: -Togrey Botanical Club (November 5th). 


500 


There is still another adaptive structural feature in the thallus 
of many lichens, especially in the Stictei, which is of undoubted 
physiological importance. These lichens present a striking ap- 
pearance by the numerous ridges and depressions on the upper 
surface of the thallus. Morphologically and physiologically the 
ridges are analagous to the vascular system in the true foliage 
leaves. On microscopic examination it is found that the majority 
of the medullary hyphae of the ridges extend parallel to the direc- 
tion of the ridges. In this region the medullary layer is also con- 
siderably thickened. It is a structure specialized to conduct the 
products of assimilation. This wavy outline of the thallus also 
increases surface expansion, whereby assimilation is increased. 
Assimilation is most active in the depressed portions of the thallus, 
which also corresponds to the position of the cyphellae on the 
lower surface. Such a wavy arrangement of structures also serves 
a mechanical function; according to the principles of mechanics 
it affords a better support to the frequently very large thallus. 


A Study of the Genus Galactia in North America. 


By ANNA MuRRAY VAIL. 


The genus Ga/actia was established by Patrick Browne in Nat. 
Hist. Jamaica, 298. p/. 32. f. 2. 1756. It was based on the plant 
figured in the illustration, a species with the stamens of Galactia 
and the showy scarlet flowers of Co//aea, which forms a connect- 
ing link between these two sections. Linnzus reduced the genus 
to Clitoria, describing the Jamaica plant under the name of C7itoria 
Galactia** Michaux in Fl. Bor. Am. 2: 61, revived the genus, 
with two North American species. 

The following genera have been reduced to Galactia: 

Odonta Bertol. Lucub. Herb. 35. 1822. 

Sweetta DC. Mem. Leg. 358. 1823. 


* The synonymy of this species is as follows: 
Gavactia Gatactia (L.). 

Clitoria Galactia L. Sp. Pl. Ed. 2, 1026. 1763. 
Galactia pendula Pers. Syn. 2: 302. 1807. 
Ittusr. Lindl. Bot. Reg. fi. 269. 1818. 


501 


Collaea DC. Ann. Sci. Nat. (1.) 4: 96. 1825. 

Betencourtia St. Hil. Voy. 25 376.. 3833. 

Leucodictyon Dalz. in Hook. Kew Journ. Bot. 2: 264. 1850. 

fleterocarpaea Scheele, Linnaea, 21: 467. 1850. 

The genus is accepted as described by Taubert in Engler and 
Prantl, Nat. Pflantzenf. 3: part 3, 369, 1894, where some fifty 
species are credited to the temperate and tropical regions of both 
hemispheres. 


Stems erect, decumbent or prostrate, zo¢ twining. 
Leaves simple; flowers 1-2, sub-sessile in the axils of the leaves. 
1. G. heterophylla. 
Leaves 3-foliolate, sometimes simple; flowers 1-several in short-peduncled or sub- 


sessile, axillary racemes or fascicles. 2. G. erecta, 
Leaves pinnately 3-foliolate or digitately 5-foliolate, with the terminal leafle 
petioled ; flowers 1-4, peduncled. 3. G. Grayz. 
Leaves 3-foliolate. a 
Stems erect, branched; leaflets oblong-linear; flowers few, approximated 
near the top of the rhachis. 4. G. brachypoda. 


Stems prostrate; leaflets oval or oblong-oval; flowers numerous, showy, scat- 
tered often the whole length of the rhachis, or in var. dongeracemosa 
mostly near the top. 5. G. Floridana. 

Stems ‘wining, prostrate or climbing on bushes. 
Leaflets 3. 
Racemes 2-7, usually fascicled, in the axils of the leaves. 

Racemes many-flowered, single or branched, as long as or longer than 
the short-petioled leaves ; leaflets round-oblong or sub-orbicular, densely 
canescent. 6. G. fasciculata, 

Racemes simple, 5-18 cm. long, flexuous and twisted ; leaves long- 
petioled ; leaflets oblong or oblong-orbicular, canescent. 

y. G, canescens. © 
Racemes solitary or with a smaller, or rarely with 2 accessory ones. 

Racemes as long as the leaves, few-flowered, or the upper flowers soli- 
tary in the axils; leaflets broad, oval, thinnish. 8. G. 7exana. 

Racemes as long as or somewhat longer than the leaves; leaflets glab- 


rate or nearly so; leaves and rigid racemes erect on the prostrate or 
g. G. regularis. 


climbing stem. 
Racemes usually much exceeding the leaves or in some forms of G. 
volubilis shorter than the leaves. 
Leaflets ovate-oblong, more or less pilose on both surfaces, rarely 
nearly glabrate ; in var. ¢¢ermediéa linear or linear-oblong. 
10, G. voludilis. 
Leaflets oval, villous; peduncles stout, hirsute, much elongated : 
flowers bright rose-purple. aL G. mollis. 
Leaflets ovate-oblong, coriaceous, bright yellow-green ; legume hir 
sute, the broad suture glabrous at maturity. 12. G. Cudensis. 


502 


Leaflets oblong-linear, more or less silvery canescent. 
13. G. Wrightit. 
Leaflets 7-9, elliptical-oblong ; flowers white, tinged with red. 
14. G. Elliottit. 


1. Galactia heterophylla (Gillies). 


Procumbent from a ligneous somewhat fusiform creeping 
rhizome; stems sub-simple, flexuous, 1-3 dm. long, minutely re- 
trorse-hirsute, becoming glabrous, obscurely angled above ; stipules 
2-3 mm. long, subulate or setaceous; petioles 4-6 mm. long; 
leaves few, the basal ones 2-4 cm. long, oblong, the others 3-9 cm. 
or more long, oblong-lanceolate or linear, acutish at each end, 
coriaceous, glabrous above, the reticulated veins confluent with the 
conspicuous marginal nerve and minutely strigose beneath ; flow- 
ers 1-16 cm. long, rose-purple, on 1-3 mm. long pedicels or I-—3, 
minutely pedunculate in the axils of the upper leaves; bracteoles 
minute, setaceous; calyx 6-7 mm. long, campanulate, strigose- 
hirsute, the acuminate teeth somewhat scarious-margined, the 
broadly ovate-lanceolate upper one minutely 2-toothed, the others 
more slender, the middle one the narrowest and longest ; vexillum 
with a conspicious indentation at the apex, minutely biappen- 
diculate at the base, abruptly tapering to the filiform claw; anthers 
oblong-ovoid; legume 2.5-3.5 cm. long, 4-5 mm. wide, coria- 
ceous, acuminate, retrorsely hirsute and puberulous, with a some- 
what raised suture. Mature seeds not seen. 


Original locality, El Rio Quarto, west side of Las Pampas, 
Argentine Republic. 


Type, Gillies in Herb. Arnott, fide Torrey & Gray. 


Eastern Texas, Central America. Uruguay and Argentine 
Republic. 


Synonymy: Cologania heterophylla Gillies ; Hook. & Arn. in Hook. 
Bot. Misc. 3: 181. 1833. ; 


Galactia marginalis Benth. Ann. Wien. Mus. 2: 126. 1838. 


2. Galactia erecta (Walt). 


Erect from a slender ligneous somewhat fusiform rhizome, 
‘stems often several from the same root, 2-3 dm. or more high 
somewhat flexuous, glabrous or nearly so, angled above ;_ stipules 
2-3 mm. long, subulate; leaves few, usually remote; petioles 2- 
5 cm. long, slender; leaflets 2.5—4 cm. long, often reflexed on the 
rhachis, oblong, oblong-linear or linear, obtuse, rarely acutish, 
coriaceous, glabrous, glaucous and the veins reticulated beneath, 
_ the terminal one subsessile; flowers 8-10 mm. long, purplish, 2- 

5 in obscurely peduncled, axillary racemes or fascicles; pedicels 


505 


1-2 mm. long; bracteoles setaceous, 1-2 mm. long; calyx turn- 
ing brown, 5-6 mm. long, hirsute, scarious-margined, the lower 
teeth slender, acuminate, the upper one broader, minutely 2— 
toothed; vexillum round-ovate, retuse; legume linear-oblong, 
about 2 cm. long, hirsute; seeds about 6, June-August. 


Original locality, “ Carolinas.” 
Dry pine barrens, North Carolina to Florida, Alabama and 


Louisiana. 


Synonymy: Evvum erectum Walt. Fl. Car. 187. 1788. 
Glycine stricta Hook. & Arn. in Hook. Comp. Bot. Mag. 1: 22, 


1835. 
Galactia sessiliflora Torr. & Gray, Fl.N. Am. 1: 288; 687. 1838. 


3. Galactia Grayi. 


Decumbent, canescent-sericeous throughout ; stems slender, 
several from a large, thick, ligneous root, retrorsely strigose, 3-6 
dm. or more long, simple or branched; stipules minute, subulate ; 
petioles 1-2.5 cm. long, canescent-strigose; leaflets 7-16 mm. 
long, oblong, sub-cuneate or nearly linear, obtuse, retuse, mucron- 
ulate or acutish, emarginate, when 5-foliolate, the terminal leaflet 
short petioluled; the accessory ones affixed to the lateral pair; . 
Sessile or nearly so; racemes 1—4-flowered; peduncles 1-2.5 cm. 


long; bracteoles minute ; pedicels 5-7 mm. long; flowers 1.5-1.8 
cm. long; calyx 8-10 mm. long, the teeth slender, acuminate, the 


slightly broader, upper one minutely 2-toothed ; vexillum yellowish, 
oblong-ovate, retuse, acute at the base, the other petals rose-purple, 
the keel conspicuously longer than the narrow wings; anthers 
oblong ; legume about 8 cm. long, 5 mm. wide, phe eee 
acute; seeds 3-4 mm. long, oblong, apparently truncate at the 


ends. 
Original locality, on the Llano River, Texas. 


Type, Lindheimer No. 591 in Herb. Columbia College. 
Texas, Llano River; San Diego (Miss Croft) ; Pena and Che- 
nate Mts. (Nealley). , 
Synonymy: Galactia heterophylla A. Gray, Bost. Journ. Nat. Hist. 
6: 171. 1850. Not G. heterophylla (Gillies.) ° 


4. Galactia brachypoda Torr. & Gray, Fl. N. Am. 1: 288. 


1838. 
Erect, stems slender, branched, at length decumbent, some- 


: led and chan- 
_ what puberulous, or nearly glabrous, slightly ang : A 
Saale A pein 4-6 dm. long; stipules 2-4 mm. long ; leaflets 2 


504 


35 cm. long, oblong or linear-oblong, thinnish, obtuse or emar- 
ginate, glabrous or minutely appressed-hirsute above, paler, some- 
what strigose or glabrate beneath, the lateral ones very short 
petioluled or nearly sessile; flowering racemes usually shorter 
than the leaves, at length elongated ; flowers rose- purple or purple, 
8-10 mm. long, sessile or very short pedicelled; bracteoles 
minute ; calyx 5 mm. long, clothed with spreading, yellowish hairs, 
the lower lobes acutish, the middle one somewhat the longest, the 
broader upper one, minutely 2-toothed ; vexillum 7-8 mm. long, 
round-ovate, apparently not indented at the apex and paler in. 
color than the other petals, the filiform claw 1 mm. long; anthers 
very small, round-oblong. Legume notseen. July-August. 


Original locality, Middle Florida. 
Type, Chapmanin Herb. Columbia College. 
Dry pine barrens, Middle Florida. 


5: Galactia Floridana Torr & Gray, Fl. N. Am. 1: 228. 1838. 


Prostrate, the whole plant hoary-pubescent ; stems 2dm.—I m. or 
more long, simple or branched, the upper part often densely and 
retrorsely white-tomentose ; stipules 2 mm. long, setaceous; in- 
ternodes usually shorter than the leaves; petioles 1-2.5 cm. long, 

-angled; leaflets 1.5-4.5 cm. long, oval or oblong-oval, obtuse, 
rarely acutish, usually mucronulate, thick and coriaceous, reticulated 
and paler beneath, the basal ones often one-foliolate; racemes 4-9 
cm. long, rarely longer than the leaves, usually many-flowered ; 
flowers 1.2-1.5 mm. long, showy, rose-purple, approximated often 
the whole length of the rhachis; bracts minute, setaceous ; brac- 
teoles close to the base of the calyx; calyx 7-8 mm. long, the 
teeth scarious-margined, long-acuminate, the upper one entirely or 
obscurely 2-toothed, the middle lower one considerably longer; 
vexillum obovate, rounded above, acute at the base, the claw 
broad and flat; anthers linear-oblong, acute at each end; legume 
4-5 cm. long, 4-5 mm. wide, linear, acuminate, tomentose. Ma- 
ture seeds not seen. June-August. 

Original locality, Tampa Bay, Florida. 


Type in Herb. Columbia College. Florida.. 


5¢. Galactia Floridana microphylla Chapm Fl. S. States, 
108. 1884. 
Smaller; petioles 5-8 mm. long; leaflets 1-2 cm. long, acutish 


or obtuse, mucronulate, the few flowers almost sessile in axillary 
clusters. Flowers and legumes ndt seen. 


Original locality, Florida. 


Type in Herb. Columbia College. 
_ Florida. 


505 


54. Galactia Floridana longeracemosa np. var. : 


Stouter, the leaves and racemes erect on the prostrate stems; 
leaflets 2.5-4.5 cm. long, 2-3 cm. wide, oval-oblong, conspicu- 
ously retuse, glabrate above, somewhat strigose-pubescent be- 
neath; flowers 1.2-1.5 cm. long, bright rose-purple, 2-4 together 
at regular intervals above the middle of the rhachis; calyx becom- 
ing glabrate. Legume not seen. 


Florida. J. H. Simpson. 1889. 


6. Galactia fasciculata n. sp. 


Prostrate or climbing high on bushes; stems I-2 m. or more 
long, terete, clothed with a close fine retrorse-canescent tomen- 
_ tum ; stipules minute, subulate; internodes longer than the leaves, 
4-6 cm. or more; petioles 5-10 mm, long; leaflets round-oblong 
or sub-orbicular, 1.5—3 cm. long, 1.5-2.5 cm. broad, retuse; finely 
appressed silky-pubescent above, appressed silky-villous and paler 
beneath; racemes several, fascicled in the axils of the leaves, form- 
ing conspicuous clusters at the slightly enlarged nodes; flowers 
pale lilac-purple, 1.5 cm. long, solitary or two together, scattered 
along nearly the whole length of the somewhat flexuous rhachis; 
bracts minute; bracteoles 1 mm. long, subulate, acuminate, close 
to the base of the calyx; calyx 8 mm. long, ‘clothed with short, 
white, silky hairs, the lobes acuminate, the upper one apparently 
entire, the middle lower one the longest; vexillum 1.5 cm. long, 
oblong-obovate, tapering to the base of the claw, obtuse or acutish 
at the apex, the wings very slender; authers linear-oblong, obtuse. 
Legume not seen. 

Conspicuous for its roundish leaflets, and its numerous fascicled 


Often branched and flexuous racemes. 
' Type, Nash, no. 2480 in Herb. Columbia College, from 


Tampa, Florida. 


7. Galactia canescens (Scheele) Benth. Ann. Wien. Mus. 2: 126. 
1838. 


Prostrate, creeping on the ground, canescent throughout, many 
of the racemes becoming subterranean and bearing globular Ir 
seeded pods: stems several from the same slender, ligneous root, 
branched at the base; stipules 1-2 mm. long, subulate; inter- 
nodes 1-15 cm. long; petioles 3-7 cm. long; leaflets 2-4 cm. 

long, 1.5—-3 cm. wide, broadly ovate or oval, retuse, thick and cori- 
aceous, slightly hirsute above, silky canescent-pubescent, much 
whiter and the veins reticulated beneath; peduncles slender, 
_ elongated, few-flowered; bracts and bracteoles minute, subulate ; 


506 


calyx silky-hirsute, the teeth acuminate; corolla rose-purple ? ; 
vexillum obovate; subterranean pods 8 mm. long and broad, re- 
ticulated, emarginate, appressed-hirsute, the seeds 6 mm. long, 4 
mm. wide, ovoid-elliptical, slightly rounded at the ends, light 
brown, the other pods 2~3 cm. long, 6-7 mm. wide, linear-oblong, 
acute, tapering to the base, densely pubescent or tomentose, the 
seeds elliptical-oblong, mature ones not seen. 


Original locality, Texas. 
Type, Lindheimer, No. 370. 
Sandy ground, Texas. 


Synonymy: Heterocarpaea Texana Scheele, Linnaea 21: 468. 1850- 


8. Galactia Texana (Scheele) A. Gray, Bost. Journ. Nat. Hist. 
GO: f70: . 15s. 


Procumbent from a ligneous root; stems slender, long, con- 
spicuously twining, retrorse-pubescent or hirsute ; stipules minute, 
caducous; petioles 1-4 cm. long, slender ; leaflets 2.5-5 cm. long, 
2-3.5 cm. wide, oval, retuse, mucronulate, minutely appressed- 
cinereous-puberulous or glabrate above, appressed sericeous or 
sparingly villous beneath, thin, becoming slightly coriaceous; ra- 
cemes few-flowered, much shorter than the leaves; flowers not 
seen; calyx 5 mm. long, the teeth attenuate; legume slightly fal- 
cate, 4-5 cm. long, 5-6 mm. wide, acute at each end, appressed- 


hirsute ; seeds 5-7, 5 mm. long, elliptical, truncate at the ends, 
light brown. 


Original locality, near New Braunfels, Texas. 
Type, Lindheimer. Confined to Western Texas. 
Synonymy: Ladlab Texanus Scheele, Linnaea, 21: 467. 1848. 


9- Galactia regularis (L.) B. S. P. Prel. Cat. N. Y. 14. 1888. 


Prostrate and climbing on bushes, from a stout, ligneous root; 
stems 4 dm—1 m. long or more, terete, minutely retrorse-pubes- 
cent, becoming glabrate; internodes about the length of the 
leaves; stipules minute, subulate; petioles 1-3 cm. long; leaflets 
2-4.5 cm. long, rather rigid, elliptical-oblong, or ovate-oblong, ob- 
tuse, rarely acutish, often retuse and apiculate, emarginate, gla- 
brous above, somewhat appressed-hirsute or pilose and lighter be- 
neath ; racemes 3-9 cm. long, usually rigid, and, with the leaves 
erect on the prostrate stem, the smaller accessory ones only occa- 
sional; flowers violet-purple, showy, often scattered the whole 
length of the rhachis; bracts small; bracteoles subulate, 1-2 mm. 
long; calyx 6-9 mm. long; glabrate or with a few scattered hairs, 
conspicuously acuminate in bud, the teeth slender, obscurely sca- 


507 


rious-margined, the upper broader one entire; vexillum obtuse or 
acutish, tapering to the flat claw, with an obscure, brownish spot 
below the middle; anthers linear-oblong, acute at each end; le- 
gume 3.5 cm. long, 4-5 mm. wide, linear, acutish at each end, ap- 
pressed-hirsute, becoming glabrate; seeds about 3 mm. long, ob- 
long-orbicular, brown. June—August. 

Original locality, Virginia. 

Type, in Herb. Mus, Brit. 

Pine barrens and sandy woods, Southern New York to Florida 
and Mississippi. 
Synonymy: Dolichos regularis L. Sp. Pl. 726. 1753. 

Frvum volubile Walt. F\. Car. 187. 1788. 

Galactia glabella Michx. Fl. Bor. Am. 2: 62. 1803. 

Galactia Purshit Desv. Ann. Sci. Nat. 9: 413. 1826. 


Ittusrrations: Hill, Veg. Syst. 21: pl. rg. 1772. 
A slender, smaller leaved form of this species has been col- 
lected at Aiken, S. C., by H. W. Ravenel. 


to. * Galactia volubilis (L.) Britt. Mem. Torr. Club, 5: 208. 
1894. 


Prostrate and climbing on bushes, from a slender, ligneous 
root; stems 4 dm.—I m. or more long, retrorsely and finely hir- 
sute; stipules minute, subulate; petioles 1-5 cm. long; leaflets 
2-5 cm. long, ovate-oblong or rarely lanceolate-oblong, obtuse, 
often retuse, mucronulate, rarely acutish, glabrous or somewhat 
pilose above, lighter, minutely pubescent or pilose beneath; ra- 
cemes 2 or 3 cm.—2.4 dm. long, often slender and filiform ; flowers 8— 
10 mm. long, lilac or pink-purple; bracts and bracteoles minute; 
calyx 6-8 mm. long, attenuate in bud, glabrate or somewhat pilose, 


* This species is based on the Dillenius plate cited below and is most variable 
and difficult to define, The type specimen is evidently the larger leaved, long- 
racemed form that occurs principally from North Carolina to Florida on the coast, 
and has been confounded with G. mo//is Michaux. The more northern plant has usually 
shorter racemes, the upper part of which often falls off before maturity of the cuit, 
which gives the appearance of very short, sometimes sessile racemes. a has in its 
more southern limits been confounded with G. tenuiflora (Willd.) Wight and Arn. 
An original type specimen of Wight Nos. 874 and 875 from Coleman; East er 
Peninsula preserved in Herb. Columbia College differs materially from the Nort 
American plant. G. ¢enuiflora occurs in South America and is i in Index 
Kewensis to G. dubia DC. Prod. 2: 238. Galactia Macreet, the type speci- 
men of which is preserved in Herb. Columbia College, is merely a very slender fili- 


form-racemed variation. 


508 


the lobes nearly equal, slender; vexillum obovate, rounded above, 
biappendiculate at the base, claw slender; legume 3-5 cm. long, 
4-5 mm. wide, acute at each end, linear or linear-oblong ; seeds 
6-9, when mature nearly as wide, truncate at each end, brown, 
with black markings. May—August. 


Original locality, “America Septentrionali.” 
Long Island to Florida, west to Pennsylvania, Tennessee and 
Texas. 


Synonymy: HHedysarum volubile L. Sp. Pl. 750. 1753. 
Galactia mollis Nutt, Gen. 2:117. 1818. Not Michaux. 
Galactia pilosa Ell. Bot. S. C. & Ga. 2: 238. 1824. Not 
Nuttall. : 
Galactia Macreei M. A. Curtis, Bost Journ. Nat. Hist. 1: 120. 
1837. 
Galactia villosa Eat. & Wright, N. Am. Bot. 248. 1840. Not 
Wight & Arnott. 


Ittustrations: Dill. Elth. 173. £2. BIT TIO. NFS, 
Hill, Veg. Syst. 21: pl. g3.fi 4. 1772. 


10a, Galactia volubilis Mississippiensis n. var. 


Leaflets oval or oval-oblong, pilose on both surfaces, lighter 
beneath, mostly retuse; racemes slender, varying in length; 


flowers slightly smaller, pale lilac; legume 3-4 cm. long. Other- 
wise as in the type. 


_ Missouri, Arkansas, Indian Territory and Louisiana. 
An oval, pilose-leaved form that in Missouri seems very dis- 


tinct, but specimens from other localities seem to intergrade with 
the broader leaved form of the type. 


1ob. Galactia volubilis intermedia. 


Very slender; leaflets linear or linear-oblong ; racemes fili- 


form, the flowers on the upper portion often sessile in the axils of 
the leaves; legume 2-3 cm. long. 


Florida to Louisiana. : 


Synonymy: Galactia pilosa + angustifolia Torr. & Gray, Fl. N. Am. 
z: 287. 1838. Not G. angustifolia Kunth. 
A very slender form, intermediate between the type and G. 
angustifolia Kunth, or what passes as such. 


509 


11. Galactia mollis Michx. Fl. Bor. Am. 2: 61. 1803. 


Prostrate or climbing, villous and somewhat hoary through- 
out ; stems several from the same slender root, simple or branched; 
the racemes and petioles mostly erect on the decumbent stems ; 
stipules subulate I-2 mm. long ; petioles 2, rarely 5 cm. long ; 
leaflets 2.5-5 cm. long, oval or oblong, obtuse or emarginate at 
both ends, sometimes retuse, the upper ones often acutish, bright, 
light green, scabrous and villous above, villous and paler beneath, 
the young ones densely tomentose, the lateral ones usually 
rounded at each end, the terminal one more tapering at the base; 
racemes solitary I-2 dm. or more long; flowers acute in the bud, 
bright red or rose-purple, short pedicelled, approximated near 
the summit of the short peduncle, bracts setaceous, minute, 
bracteoles subulate, 1.5 mm. long; calyx about 6 mm. long, 
densely villous, the broader upper tooth entire, nearly equalling the 
more slender lower ones ; vexillum obovate, rounded above, taper- 
ing.to the obscurely biappendiculate base, keel-petals paler in 
color and considerably smaller than the slender wings; legume 
2.5-3.5 cm. long, 4.5-5.5 mm. wide, linear-oblong, acutish,densely 
tomentose; seeds ovoid, acutish, brown. July—August. 

Original locality, ‘ Carolina inferiore.” 


Type, in Herb. Michaux. ; 
Dry, sandy pine lands and swamps, North Carolina to Florida. 


Synonymy: Galactia pilosa Nutt. Gen. 2: 116. 1818. Not EI- 


liott. 
A very slender form of the above species with narrower leaflets, 


which are obscurely scabrous or glabrate above and less villous 
beneath, occurs in Florida. i 


12. Galactia Cubensis H. B. K. Nov. Gen. 6: 429. 1823. 


Climbing high on shrubs and trees; stems terete, retrose-pu- 
bescent, becoming often nearly glabrate ; stipules 2-3 mm. long, 
subulate; petioles 1-3 cm. long; stipels 2 mm. long, setaceous, 
2mm. long, often persistent; leaflets 2. 5-8 cm. long, oblong, ob- 
: long-oval or elliptical, obtuse and emarginate at both ends, thick, 
rigid and coriaceous, glabrous and finely reticulated above, pubes- 
cent, paler and reticulated beneath; racemes mostly longer than 

€ leaves, 5 cm.—1.4 dm. long; flowers 10-12 mm. long, pale 
purple, sessile or nearly so, solitary or two, opposite at — in- 
tervals often the whole length of the rhachis; calyx 5-6 mm. long, 


the lobes attenuate, pubescent ; vexillum obovate see hg ea 
long, 5-6 mm. or more wide, acutish, hirsute, at length gta ; 
& 1 labrous; seeds 6-10, 3-4 mm. 


€ conspicuous, raised sutures g 


510 


long, oblong, ovoid, brown, mottled with black. Very variable. 
March—December. 


Original locality, near Havana, Cuba. 
Keys, Florida, West Indies; also in Central America. 


Synonymy: Galactia spiciformis T. & G. Fl. N. Am. 1: 287. 1838. 


13. Galactia Wrightii A. Gray, Pl. Wright. 1: 44. 1852. 


Sub-erect or decumbent; stems several from the same lignes- 
cent root, branched from the base, cinereous with a fine, white, 
appressed, retrose pubescence; stipules 1-2 mm. long, subulate 
or setaceous ; petioles 1-4 cm. long; leaflets oblong or oblong- 
linear, 2-5 cm. long, obtuse, mucronulate, less canescent above 
than beneath, rarely becoming glabrate above; racemes many- 
flowered, 5-15 cm. long, the lower ones longer than the upper 
ones; flowers purplish, about 1 cm. long, sessile or nearly so, 2-3, 
fascicled at mostly regular intervals along the upper portion of the 
rhachis ; calyx 5-7 mm. long, sericeous, the lobes acute, the mid- 
dle lower one attenuate and longer; vexillum 7-10 mm. long, 
obovate, retuse, tapering to the biappendiculate base ; legume 3-4 
cm. long, 4 mm. wide, sericeous, acute at each end, 4-7 seeded. 
Mature seeds not seen. July—October. 


Original locality, Hills near Limpia, Texas. 
Type, Wright No. 111. 1849. 
Extreme western Texas and adjacent New Mexico and Mexico. 


Synonymy: Galactia tephrodes A. Gray, Pl. Wright. 2: 34. 1853- 


. 


a 14. Galactia Elliottii Nutt. Gen. 2: 117. 1818. 


Prostrate and climbing high on brushes; stems branched, 1-3 
m. long, from a long, branched, lignescent rhizome, angled, 
channelled, striate and somewhat retrorse-hirsute ; stipules minute, 
subulate; petioles 2-5 cm. long; leaflets 2-3 cm. long, elliptical- 
oblong, obtuse, somewhat retuse, mucronulate, coriaceous, 
glabrous and shining above, pubescent beneath; racemes longer 
than the leaves, 1-3 dm.; flowers 10-12 mm. long, white, tinged 
with red, 2 or 3 together, fascicled at regular intervals along the 
upper part of the rhachis; bracts and bracteoles minute; calyx 
about 7 mm. long, appressed-hirsute, the lobes slender, attenuate, 
vexillum obovate, obtuse or acutish, obscurely biappendiculate 
at the base; legume 3-4 cm. or more long, 1 cm. wide, oblong, 
acutish, emarginate, tomentose; seeds 3-5, about 5 mm. long, 
ovoid, blackish, smooth and shiny. 


Original locality, South Carolina. 


: 520 ee 


511 


Dry soil and pine lands, South Carolina to Florida, near the 
coast, 
SYNONYMY: Gadlactia pinnata Muhl. Cat. 65.1813. Not Persoon, 
1807, 

Lephrosia Elhotti Benth. Ann. Mus. Wien 2: 127. 1838. 


14a. Galactia Elliotti Leavenworthii Torr. & Gray. Fl. N. 
Am. 1: 687. 1838. 
Silky pubescent or villous throughout. 
Type in Herb. Columbia College. 
Florida, Leavenworth ; Lake Co., Nash. 


New or noteworthy American Grasses.—lI], 
By Geo, V. Nasu. 
EATONIA NITIDA (Spreng.). 


Aira nitida Spreng. Fl. Hal. Mant. 1: 32. 1807. 
Aira Pennsylvanica Spreng. Mem. Acad. St. Petersb. 2: 299. 


pl. 7. 1810. 


Aira mollis Muhl. Gram. 82. 1817. 

Eatonia Dudleyi Vasey, Coult. Bot. Gaz. 11: 116. 1886. 

Sprengel in his description of Azra nitida describes the leaves 
as 1’ long and pubescent at the base. These are characters which 
are peculiar to the plant known as Eatonia Dudleyi Vasey, which 
is very common in Pennsylvania, the region from which Sprengel’s 
plant came. The Azra Pennsylvanica Spreng. is certainly the 
LEatonia Dudleyi Vasey. The figure cited above well represents 
this species. 

The more prominent characters which appear to separate this. 
plant from the Eatonia Pennsylvanica(DC.) A. Gray, are: The almost 
constant pubescence of the sheaths, at least the lower ones; the 
slender culm; the short, often pubescent leaves; and the broader 
first scale, which is about one-third the width of the second. 


Puccrneciia arrorpes (Nutt.) Wats. & Coult. in A. Gray, Man. 


Ed. 6, 668. 1890. 
Poa airoides Nutt. Gen. 1: 68. 1818. 


512 


Glyceria distans var. airoides Vasey, Cat. Grasses U.S. 87. 
1885. 

Panicularia distans airoides Scribn. Mem. Torr. Club, 5: 54- 
1894. 

This plant appears to be specifically distinct from the P. ads- 
tans Parl. The taller and more slender culms, the panicle- 
branches dividing nearer the extremities, the fewer and less 
crowded spikelets, and the comparative length of the empty and 
flowering scales serve well to distinguish it. In P. distans, a coast 
plant, the second empty scale is less than one-half as long as the 
flowering ones ; while in P. airoides, a plant of the western interior, 
it exceeds one-half the length of the flowering scales. 

” PUCCINELLIA ANGUSTATA (R. Br.). 

Poa augustata R. Br. App. Parry’s Voy. 287. 1824. 

Glyceria angustata Fries, Mant. 3 : 176. 1842. 

Panicularia angustata Scribn. Mem. Torr. Club, 5+ 54. 1894. 


~ Bromus Porrert (Coulter). 

Bromus Kalmii var. Porteri Coulter, Man. Rocky Mt. Region, 
425. 1885. 

Culms 1 14°-3° tall, erect, simple, pubescent below the nodes. 
Sheaths shorter than the internodes, glabrous or sometimes softly 
pubescent; ligule 14” long, truncate; leaves 1/’—3/’ wide, rough, 
those of the culm 4’-9’ long, the basal narrow and about one- 
half of the length of the culm; panicle 3/—6’ in length, its branches 
drooping and flexuous, at least when old, the nodes of the axis 
pubescent; spikelets 5-10-flowered, 9/’-15” long, on slender 
flexuous pedicels; empty scales pubescent, the first narrower than 
the second, both 3-nerved ; flowering scales 5’’-6”’ long, obtuse, 
5-7 nerved, densely pubescent with long silky hairs; awn 1/-2” 
long. 


In dry soil, South Dakota to Montana, south to western Ne- 
braska, New Mexico and Arizona. 

This plant is readily distinguished from B. Kalmii by its 3- 
nerved second scale and longer flowering scales. In B. Kalmit 


the second scale is 5~7-nerved and the flowering scale about 4” 
long. 


513 


Notes on Some Southern Cassias, 


By CHARLES Louts PoLtaRD,. 
(PLATES 250-252.) 


The Chamaecristoid Cassias have always been recognized as a 
group of great perplexity within a genus of otherwise clear delimi- 
tations. Even Bentham, whose careful monographic work is still 
our standard authority on this genus, admits that his treatment of 
the Chamaecristae verae is far from satisfactory.* The difficulty 
is due to the close relationship which the species exhibit, and to 
the consequent fact that specific distinctions throughout the group 
are hardly obvious in a superficial examination. I am convinced, 
however, that differences in the periods of flowering and fruiting, 
as well as in geographical range, will afford characters by which 
our American species may be satisfactorily separated. The recent 
collections of Messrs. A. H. Curtiss and G. V. Nash, in Florida, 
have contributed much toward a clearer comprehension of this es- 
sentially Southern group. 

The Chamaecristae verae fall naturally into two divisions: 
those with small or minute, and those with relatively large flow- 
ers; and the general tendency among American botanists has been 
to refer all plants belonging to the first division to C. wictitans, 
while all large-flowering forms were unhesitatingly classed with C 
Chamaccrista. Muhlenberg was probably the first to recognize 
that the South contained more than two species and he accordingly 
named the hirsute plant which grows in dry fields in Florida 
Cassia aspera.+ This was reduced by Torrey and Grayf, but 
has been quite generally recognized in recent years as a perfectly 
valid species. Twenty years before this Michaux had described 
what he took to be a new Cassia, under the name of C: fasciculata,|| 
but this was referred by Bentham, who had access to Michaux’s 
type in Paris, to C. Chamaecrista, and it probably belongs there. 

After a critical study of numerous specimens I have reached 
the conclusion that the true C. nictitans of the North does not ex- 
tend into Florida; a theory that seems to be confirmed by nu- 
Merous field observations kindly communicated to me by Mr. 


*Trans, Linn. Soc. London, 27: 536. 1871. + Ell. Bot. S. Car. & Georg. 1: 474. 
1821. + Torr & Gray. Fl. N. Am. 396. 1838. | Fl. Bor. Am. 1: 262. 1803. 


514 


Nash. It isreplaced by C. aspera, a plant of dry barren fields and 
by a multipinnate, rather tall growing plant, hitherto unnamed, dis- 
tributed as zcftfans, but certainly not referable to that species. 

C. Chamaecrista grows in. Florida, assuming an erect, bushy 
habit, and often attains a height of from two to three feet. It is 
associated with another undescribed species, of low growth and 
divaricately much branched. The technical distinctions between 
these two new species will be discussed in the subjoined descrip- 
tions; but the differences shown in the flowering period are worthy 
of special note. 

Cassia nicticans flowers in August throughout the North and 
somewhat earlier south of the Virginia line. Its Florida congener 
is barely in flower by the middle of August, while flowering speci- 
mens collected by Mr. Curtiss bear date of September 15, at which 
time mictitans has withered foliage and thoroughly mature pods. 
C. Chamaecrista flowers from May 1 to 15,in Florida,and through- 
out June at the North; the new allied species was collected by 
Mr. Nash on September 5, with no pods as yet formed. 

These data are remarkable and quite conclusive. It is natural 
to find a plant of northern ‘range blooming a week or even a 
month earlier in a more southerly latitude ; but itis not to be sup- 
posed that a plant whose flowering period extends through May 
and June should assume an autumnal form in the South, especially 
when we bear in mind the fact that all these Cassias are annuals, 
and hence can bloom normally but once in a season. It is of 
course barely possible that C. Chamaecrista ripens its fruit in June 
and presents well developed plants two or three feet in height in 

late August; in case careful field observations should prove this to 
be a fact, we shall have a most remarkable case of dimorphism in 
a single species. 

The following key presents the salient characters of the Chamae- 
cristae verae in North America. C. cinerea Cham. and Schlecht., a 
species of Texano-Mexican distribution simulates this group, but 
belongs in the series Dimidiatae. 


KEY TO THE SPECIES. 
Flowers large (exceeding 1 cm.). 
Leaves glabrous. 
Plants tall, erect. C. Chamaecrista, 
Plant low, diffusely branched. C. depressa, 


EE EE ES ek ae eS at eae tee ee ee ee 


515 


Leaves pubescent. 
Leaflets linear falcate, appressed. C. procumbens. 
Leaflets elliptical, spreading. C. mississippiensis, 


Flowers small (not exceeding 3 cm.). 
Plants strongly hirsute, 


Leaves 8—-12-foliolate. C. simpsont. 
Leaves 20-24-foliolate. C. aspera. 

Plants glabrate or pubescent. 
Leaflets narrow, 20-25. C. muttipinnata, 
Leaflets broad, 8-14. C. nictitans. 


CASSIA MULTIPINNATA 0. sp. 

. Slender, tall, erect, more or less branched; stem slightly woody 
at base, pubescent; leaves lanceolate-elliptical in outline, 20-25- 
foliolate, the rhachis obscurely strigose-pubescent; leaflets 10 mm. 
long, 114 mm. wide, glabrous, narrowly linear, cuspidate, the mid- 
vein approximating the outer margin; petiolar gland minute, de- 
pressed-cupuliform, substipitate ; stipules long-acuminate, striate, 
foliaceous; flowers scattered, sessile, the corolla quite irregular, 
its small yellow petals not much exceeding the narrow, scarious- 
edged sepals; legume linear, compressed, pubescent or even hir- 
sute, dorsally and ventrally dehiscent, elastic, 2 to 2% cm. long 
_ Seeds 4 to 6, oblique. (Plate 250.) 

Near Jacksonville. A. H. Curtiss, North American Plants. 
No. 512, distributed as C. nictitans and C. nictitans var. aspera; 
Second distribution, No. 5157, September 15, and October 27, 
1894. ‘Tallahassee. G. V. Nash, No. 2403, August 12, 1895. 

This is separable from victitans by the narrowly linear, very 
numerous leaflets, the more hairy pod, and the late period of flow- 
ering. The irregular corolla, noticeable in some other members 
of this group of Cassia, but not often commented on, is here very 
prominent, one petal greatly exceeding the others. 


CASSIA MULTIPINNATA NASHII n. var. 
Differs from the erect type in being of low, diffusely spreading 


habit. 
Collected in low pine woods, River Junction, Gadsden Co., 


Florida, by Mr. Nash, September 5, 1895 (no. 2577). 


CASSIA DEPRESSA Nn. Sp. 
Low and spreading, but not prostrate, divaricately much 


516 


branched; stems obscurely strigose-pubescent; leaflets 6 to 10 
pairs, linear-elliptical, glabrous, 9 mm. long, 2 to 3 mm. broad, 
with the excentric midrib of C. Chamaecrista ; petiolar gland small, 
circular, sessile, depressed; flowers solitary on filiform mostly 
supra-axillary peduncles; petals large, obovate, much exceeding 
the sepals, 11%4 cm. long, 1% cm. broad; sepals linear, scarious, 
strongly ciliate; legume not observed. (Plate 251.) 

Low pine woods, River Junction, Gadsden_Co., Florida. G. 
V. Nash, September 5, 1895 (no. 2571). 

Potosi, Missouri. F. Pech. 

The difference between this and C. Chamaecrista lies mainly in 
its aspect, but the leaflets are smaller and less numerous and the 
flowers are relatively much larger. The flowering period, more- 
over, is from two to three months later than in that species. 

Cassia Mississippiensts Pollard, Bull. Torr. Club, 21: 219. 
1894. 

This plant has not yet been collected, so far as I am aware, 
outside of the State from which it was first described. Professor 
S. M. Tracy, of the Mississippi Agricultural Experiment Station, 
wrote me that he had four acres covered with a dense growth of 
the species ; from which J infer that it is locally as troublesome a 
weed as C. nictitans. 

Cassia Simpsoni Pollard, Bull. Torr. Club, 21: 221. 1894. 

Apparently a subtropical type, confined in the United States 
to the Keys of Florida, but probably growing in Cuba. It may 
_ be considered, in fact, the West Indian representative of the 
Texano-Mexican species C. procumbens L. The originalspecimens, 
collected by Mr. J. H. Simpson, on No Name Key, are in the 
possession of the National Herbarium. I have detected it in no 
other collection thus far examined. 


Explanation of Plate 252. 
Fig. 1. Cassia nictitans L. 
Fig. 2. Cassia aspera Muhl. 
Fig. 3. Cassia multipinnata Pollard, n. sp. 
Fig. 4. Cassia Simpsoni Pollard. 
Fig. 5. Cassia Chamaecrista L. 
Fig. 6. Cassia depressa Pollard, n. sp. 
Fig. 7. Cassia Mississippiensis Pollard. 
Fig. 8. Cassia procumbens L. 


ol7 


Calamagrostis scopulorum Jones.* 


Last spring, when Mr. Jones was engaged in naming his col- 
lection of Utah plants at the National Herbarium, his specimens 
of Calamagrostis were submitted to me for determination, as I am 
engaged in revising the North American species of that genus. I 
mentioned to Mr. Jones that the species in question was doubtless 
an undescribed one, and requested that the specimens be left with 
me for further study. Mr. Jones told me that he would have to 
take the specimens away then to catalogue them, but gave me the 
impression that I should have an opportunity to examine them 
later on. Since then I have not seen them, and specimens of 
this, as well as of Mr. Jones’ two other new species of grasses, ap- 
pear to be missing from the set deposited in the National Herbar- 
ium. I think it well to call attention to this as an instance of a 
Species of discourtesy (to use no harsher word) too often suffered 
by students of special groups. 

Calamagrostis scopulorum is a pédlectie distinct species of 
rather perplexing relationship. It is mo¢ closely allied to C. sy/- 
vatica, being distinguished from all species of that group by its 
slender, straight awn not exceeding the flowering-glume. In its 
spikelets it resembles C. A/eutica, but is widely different in its 
vegetative organs, habit and appearance. At first glance it ap- 
pears very close to C. Cusickit, but is readily distinguished by the 
shorter and more delicate awn and longer callus-hairs. The same 
species was collected by Mr. Jones at Alta, Utah, in 1879 (no. 
1145). Some expressions used in the description are rather pecu- 
liar, such as “ spikelets appressed,” “ outer ” and “inner rays ” of the 
panicle, lower empty glume and palet “simply acute,” etc. It is 
not usual, in describing grasses, to mention the characters of the 
palet before describing the flowering- glume. 

In passing, it may be worth while to mention that, as Lamarck 
published a Poa festucoides, that name is not available for Mr. 
Jones’ species, for botanists of every school have practically agreed 
that, from now on, at any rate, the law of homonyms shall be ob- 
served, even if it be not made retroactive. An Elymus with the 


 *ME, Jones, Contributions to Western Botany ; Proc. Calif. Acad. (IL.) 5: 722, 
1895. 


518 

“habit of Sporobolus airoides, but culms very different ;”’ ‘ spike- 
lets single at each joint;” “loose spike like Agropyrum glaucum, 
which it much resembles ;’’ and “ most of the technical characters 
of Hystrix, but manifestly allied to Elymus condensatus and may 
prove to be only a form of it,” may well be called a “ remarkable 
plant.” T. H. Kearney, JR. 


Proceedings of the Club. 
WEDNESDAY EVENING, Nov. 27, 1985. 


The President occupied the chair and there were about 50 per- 
sons present. Benjamin Heritage and Jos. A. Hays were 
elected active members. 

The announced paper of the evening was then read by Dr. 
Rusby, entitled, “ The Poisonous Plants of the Vicinity of New 
York,” illustrated by lantern slides, mostly colored, and loaned by 
Mr. Van Brunt for the occasion. The paper was discussed by the 
President and Messrs. Small, Barnhart, Stewart and others. It is 


printed in full in the « Alumni Journal of the New York College 
of Pharmacy,” December, 1895. . 


Wepnespay Evenine, Dec. 11, 1895. 


Vice-President Lighthipe in the chair, and 52 persons present. 

The Secretary called attention to the necessity for a new sup- 
ply of copies of the Constitution and By-Laws and suggested that 
a revision of this publication be made, to include a short history 
of the Club. He referred also to the need of other stationery to 
facilitate the work of the officers of the Club, and offered a resolu- 
tion that a committee to consist of the Editor, the Treasurer and 
the Secretary be appointed with the power to prepare a revised edi- 
tion of the Constitution and By-Laws and such other stationery as 
is required. The resolution was unanimously adopted. 

A communication from Mr. Wm. H. McDonald, of 2297 First 
avenue, was read, tendering a number of seeds for use in the nur- 
sery of the New York Botanical Garden. 

The announced lecture of the evening, by Dr. Smith Ely 

Jelliffe, entitled, « Common Edible and Poisonous Fungi,” illus- 
_ trated by colored lantern slides, was then delivered. 


519 


Index to recent Literature relating to American Botany. 


Allen, C. L. Late-flowering Golden-rods. Gard. & For. 8: 458. 
13 N. 1895. 
Bastin, E. S. Some further Observations on Cherry barks. Am. 

Journ. Pharm. 67: 595-599. f. 7-3. D. 1895. 
Bequirot, A. Sulla presenza in Italia della Oxalis violacea L. Bull. 
Soc. Bot. Ital. 6: 110. O. 1895. 


Bigelow, C. M. Study of Glands in the Hop-tree. Proc. Iowa 
Acad. Sci. 2: 138-140. A/. ro. 1895. 
Glands of Prelea trifoliata. 

Bioletti, F. T. Notes on the Genus Vemofhila, Erythea, 3: 139-142. 
pl. 2. O. 1895. 


Broadway, W. E. Cyrtopodium Woodford. Gardn. Chron. 18: 
Org 93 N. 189k. 


Bush, B. F. A new Hybrid Oak in the Indian Territory. Gard. & 
For. 8: 463. 20 N. 1895. 

Crosier, A. A. The common Names of the Clovers. Bull. Mich. 
Exp. Sta. 125: 14-28. Je. 1895. 

Davy, J. B. Pacific Slope Plants in English Gardens. Erythea, 3: 
143-147, .O. 1895. 

Elfert, W. Morphologie und Anatomie der Zimosella aquatica. 
Inaug. Diss. pp. 44. Berlin. No date. 

Fink, B. Lichens collected by Dr. C. C. Parry in Wisconsin and 
Minnesota in 1848. Proc. Iowa Acad. Sci. 2: 137. 1895. 
List of twenty-three species. 

Folsom, J. N. Autumnal Changes in Leaves—I.-II]. Gard. & For. 
8: 383-384, 392-393. 25 S.-2 O. 1895. 

Gibson, W.H. Our edible Toadstools and Mushrooms. New York, 
PP. 337- PA 38. 1895. 

Gilbreth, M. E. Dissemination of Plants chiefly ae their Seeds. 
Pamphlet, pp. 15. 65S. 1895. 

Gregory, E. L. Elements of Plant Anatomy. pp. oe eer 64. 
Boston, 1895. 

Hemsley, W. B. NVeogoesia gracilipes. Hook. Icon. Pi) age DL. 
2424. N. 1895. 


520 


Hemsley, W. B. Neogoezia minor. Hook. Icon. Pl. 25: pl. 2425. 
N, 1895. 


Hieronymus, G. Plantae Lehmannianae in Guatemala, Costa Rica, 
Colombia et Ecuador collectae, additis quibusdam ab aliis collectori- 
bus ex iisdem regionibus necnon e Venezuela et Peruvia allatis, quos 
determinavit et descripsit adjurantibus aliis auctoribus. Beibl. Engl. 
Bot. Jahrb. 20: 1-72. 9 Ap. 1895. 


Hieronymus, G. Plantae Steubelianae oe quas descripsit adju- 
rantibus aliis auctoribus. Engl. Bot. Jahrb. 21: 306-368. 6 Au. 
1895. 

New species from Colombia, Bolivia, Ecuador and Peru. 

Hooker, J.D. Amasonia erecta var. latebracteata. Curt. Bot. Mag. 
51: pl. 7445. N. 1895. 

Hooker, J.D. Catasetum Lemosii. Curt. Bot. Mag. 51: pl. 7444- 
N. 1895. 


Hooker, J. D. Selenipedium Sargentianum., Curt. Bot. Mag. 51: pl. 
7446. N. 1895. 


Hoskins, T. H. An Indian on Indian Corn. Gard. & For. 8: 23 
J. 3-¢. 16 Ja. 1895. 


Hoskins, T. H. The Germination of Nuts. Gard. & For. 8: 448. 
6N. 1895. 


James, J. F. Remarks on a “Catalogue of Ohio Plants’’ by Keller- 
man and Werner. Journ. Cinn. Soc. Nat. Hist. 18: 46-57. Ap.- 
Jl. 1895. 


Jenman, G. S. Polypodium (Eupolypodium) xiphopteroidacfolium. 
Gardn. Chron. 18: 612. 23 N. 1895. 


A new species from Cuba. 


Kearney, T. H., Jr. Notes on Grasses and Forage Plants of the 


Southeastern States. Bull. U. S. Dept. Agric. Div. Agrost. 1: pP- 
28. f. 6. 1895. 


Kranzlin, F. Masdevallia Forgetiana. Gardn. Chron. 18: 484- 
26 Q. 1895. 
A new species from northern Brazil. 

Kuntze, O. Les Besoins de la Nomenclature Botanique. Reprint 
from Monde des Plantes. 1895. 

Massalongo, C. Interno ad una nuova varieta di Coll/insia bicolor 
Benth. Bull. Soc. Bot. Ital. 1895: 222-224. N. 1895. 


Collinsia bicolor incisiflora n. var. 


521 


Meehan, T. Asfidium marginale. Meehan’s Month, B+ eat. Of, 72. 
D. 1895. 


Millspaugh, C. F. Contribution to the Flora of Yucatan. Contr. 
Field Columbian Mus. Bot. 1: 1-56. f/ 7-¢. Au. 1895. 


Morgan, A. P. New North American Fungi. Journ. Cinn. Soc. Nat. 
Hist. 18: 36-45. /. 7-3. Ap.-Jl. 1895. 

Nemnich, H. Ueber den anatomischen Bau der Achse und die Ent- 
wicklungsgeschichte der Gefassbiindel bei den Amarantaceen. Inaug. 
Diss. pp. 38. f. 7. Erlangen, 1894. 

Oliver, D. Daphnopsis Beta. Wook. Icon. Pl. 25: p/. 2409. N. 
1895. 

Oliver, D. Lissocarpa Benthami. Hook. Icon. Pl. 25: pl. 2473. 
N. 1895. 

Oliver, D. Orthaea cordata. Hook. Icon. Pl. 25: pl. 2gr2. N. 
1895. 

Oliver, D. Ramisia Brasiliensis. Wook. Icon. Pl. 25: pi. 2404. ° 
N. 1895. 

Pammel, L. H. Diseases of Plants at Ames, 1894. Proc. Iowa 
Acad. Sci. 2: 201-208. 1895. 

Pammel, L. H. Distribution of some Weeds in the United States, 
especially /va xanthiifolia, Lactuca Scariola, Solanum Carolinense 
and Solanum rostratum. Proc. Jowa Acad. Sci. 2: 103-127. 1895. 

Pammel, L. H., and Beach, A. M. Pollination of Cucurbits. Proc. 
Iowa Acad. Sci. 2: 146-152. f/. 21-14. 1895. 

Plank, E. N. Botanical Notes from Texas—XIX-XXV. Gard. & 
For. 7: 222, 253, 283, 313, 342, 6 Je., 29 Au. 1894. 8: 72, 193. 

_F. My. 189s. 

Pringle, C. G. Notes of Mexican Travel.—X. Gard. & For. 8: 
272-273. f. 399. 10 Jl. 1895. 

With illustration of Lamourouxia Pringlet. 

Rose, J. N. A yellow-flowered Cosmos. Gard. & For. 8: 484. /. 66. 
4D. 1895. 

Note on Cosmos sulphureus Cav. 

Sargent, C. S. Fothergilla Gardeni. Gard. & For. 8: 446. f. 62. 
6N. 1895. 

Sargent, C. S. The Names of some North American Tree Willows. 


Gard. & For. 8: 463. 20 N. 1895. 


522 


Sargent, C.S., Editor. James Buckman and Plant Variation. Gard. 
& For. 8: 442, 6N. 1895. 


Schumann, K. Cereus Greggii. Monatss. Kakteenk. 5: 149-153. 
20 O. 1895. - 

Schenck, R. Botanisch-pharmacognostische Untersuchungen der 
Quamacai cipo. Inaug. Diss. pp. 22. 7.4. Erlangen, 1894. 
Examination of Paullinia thalictrifolia from Brazil. 

Scribner, F. L. Hairy Vetch, Sand Vetch or Russian Vetch (Vicia 
villosa). U.S. Dept. Agric. Div. Agrost. Circular 2: pp. 4. fig: 
1895. - 

Sirrine, E. Structure of the Seed-coats of Polygonaceae. Proc. Iowa 
Acad. Sci. 2: 128-135. p/. 7-9. 1895. 

Smith, J.D. Enumeratio Plantarum Guatemalensium necnon Salvador- 
ensium, Hondurensium, Nicaraguensium, Costaricensium. 4: PpPp- 
183. 1895. 

Van Tieghem, P. Observations sur la structure et de la déhiscence 

- des antherés des Loranthacées, suivies de remarques sur la structure et 
la déhiscence de l’anthére en général. Bull. Soc. Bot. France, 34: 
363-368. 1895. 

Van Tieghem, P. Sur le groupement des especés en genres dans la 
tribu des Psittacanthées de la famille des Loranthacées. Bull. Soc. 
Bot. France, 34: 343-362. 1895. 


Definitions of sixteen new genera, 


Watson, W. Jubaea spfectabilis. Gardn. Chron 18: 518. f. 90. 2 
N. 1895. 


BULLETIN OF THE TORREY BOTANICAL CLUB—.PLATE 250. 


Ui 
er ap 


CASSIA MULTIPINNATA POLLARD. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 251. 


CASSIA DEPRESSA POLLARD. 


BULLETIN OF THE TORREY BOTANICAL CLUB.—PLATE 252. 


4 
Lapal 
B 
0 
Z; 
fs 
x 
o 
5 
n 
te 
6) 
Nn 
a 
4 
- 


Contributions from the Herbarium of Columbia College. 


No. 4 
No. 

No. 6 
NG. 9 
No. 9 


[ The numbers omitted from this list are out of print. ] 


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