occasional papers of the
Farlow.
Herbarium of cryptogamic botany
No. 17 August, 1982 Harvard University, Cambridge, Massachusetts
Donalc H. Pfister A Nomenclatural Revision of F.J. Seaver’s North American
Cup-Fungi (Operculates)
Edited by: Donald H. Pfister
Carolyn S. Hesterberg
ISSN: 0090-8754
occasional papers of the
Farlow
HerbariuM of cryptogamic botany
No. 1.
No. 2
No. 3
No. 4.
No. 5.
No. 6
No. 7.
No. 8.
No. 9
No. 10.
Sylvia A. Earle: Hummbrella, a New Red Alga of Uncertain Taxonomic
Position from the Juan Fernandez Islands (June 1969).
. |. Mackenzie Lamb: Stereocaulon arenarium (Sav.) M. Lamb, a
Hitherto Overlooked Boreal-Arctic Lichen (June 1972).
. Sylvia A. Earle and Joyce Redemsky Young: Siphonoclathrus, a
New Genus of Chlorophyta (Siphonales: Codiaceae) from Panama
(July 1972).
|. Mackenzie Lamb, William A. Weber, H. Martin Jahns, Siegfried
Huneck: Calathaspis, a New Genus of the Lichen Family Cladoniaceae
(July 1972).
|. Mackenzie Lamb: Stereocaulon sterile (Sav.) M. Lamb and Stereo-
caulon groenlandicum (Dahl) M. Lamb, Two More Hitherto Over-
looked Lichen Species (March 1973).
_ 1. Mackenzie Lamb: Further Observations on Verrucaria serpuloides
M. Lamb, the Only Known Permanently Submerged Marine Lichen
(April 1973).
Bruce H. Tiffney and Elso S. Barghoorn: The Fossil Record of the
Fungi (June 1974).
Donald H. Pfister: The Genus Acervus (Ascomycetes, Pezizales).
|. An Emendation. II. The Apothecial Ontogeny of Acervus flavidus
with Comments on A. epispartius (May 1975).
. Donald H. Pfister: A Synopsis of the Genus Pulvinula. A New
Combination in the Genus Gymnomyces. Norton G. Miller: Studies
on North American Quaternary Bryophyte Subfossils. |. ANew Moss
Assemblage from the Two Creeks Forest Bed of Wisconsin (July
1976).
Emmanuel Sérusiaux: Some Foliicolous Lichens from the Farlow
Herbarium (August 1976).
Continued on back cover
A NOMENCLATURAL REVISION OF F.J. SEAVER'S
NORTH AMERICAN CUP-FUNGI (OPERCULATES)
DONALD H. PFISTER
The North American Cup-fungi (Qperculates) (Seaver
1928) has been a standard reference for more than 50 years.
It was revised by Seaver in 1942 and has stood as an author-
itative source since that time. No modern work on the Pez-
izales fails to cite it and, though there are some mono-
graphs for genera and some regional floras, one invariably
still turns to this book when one is identifying Pezizales.
In the years since Seaver's work, the Pezizales have
been reworked at the generic level and important cytologi-
cal, histochemical, and developmental work has been done on
the group. Several major monographs of genera have been
written as well as checklists and regional reports. It is
not surprising that, when using Seaver's treatment, one is
often able to identify a species without having any idea of
where the species belongs in modern generic systems.
Seaver's treatment was a highly artificial one. I have set
about annotating Seaver's nomenclature.
Modern currently accepted names for his concepts or
Names are provided in the list which follows. These are
opinions. Certainly one can turn to Seaver's list of
synonyms and find other opinions. I have tried to follow a
conservative pathway and have listed binomials which for the
most part are now in common usage. In a few cases some
rearrangement is proposed but generic concepts are _ those
which are found in Eckblad!s (1968) revision, Rifai's (1968)
treatment of the Austral-Asian Pezizales, Dennis's (1978)
recent revision, or Korf's keys (1972, 1973a).
I have not tried to provide all of the synonyms but
rather to fit Seaver's taxa into recognizable modern genera.
This has been easier in some groups than in others as_ the
listing which follows will amply show. Some genera have
been well worked; others are woefully understudied. Still
it is hoped that those using the book will find this annota-
tion helpful.
The annotation is made up of three parts. Part I lists
the name used by Seaver, the page number in North American
Cup-fungi (Operculates) revised edition, and the equivalent
name where known. Comments on some of the names are made in
numbered notes referred to in the revised list. Those
Seaver names which are underlined were listed as doubtful by
Seaver. The Seaver names marked with a "+" represent names
which for sundry reasons were not possible to verify in
North America. Part II is an index to the revised names as
listed in Part I. The page numbers refer to Seaver's book.
With each generic name, reference to recent monographic work
is given. Part III is a bibliography. Most recent treat-
ments containing references to North American taxa are
listed.
I wish to thank several curators who have seen to it
that I had specimens for study. These are Clark T. Rogerson
(NY), David Farr (BPI), John H. Haines (NYS), and Richard P.
Korf (CUP). My debt to Prof. Korf is a profound one. He
reviewed this manuscript, made suggestions, and gave
encouragement. He provided many of the clues which lead to
the solution of some of the tangles. He also got me started
on discomycetes as a graduate student. This research was
supported in part by National Science Foundation grant DEB
8023018 to the author.
Page
43
44
45
46
46
47
48
49
50
51
6
a2
53
56
58
58
59
PART I
NAME USED BY SEAVER
Sphaerospora trechispora
S. flavovirens
S. perplexa
S. brunnea
S. hinnulea
S. monilifera
Pseudoplectania
vogesiaca
P. nigrella
P. fulgens
Sphaerosoma hesperium
Boudiera areolata
B. echinulata
B. marginata
Lamprospora crouani
L. laetirubra
L. dictydiola
L. areolata
REVISED NAME
Scutellinia trechispora
(Berk. & Broome) Lambotte
Scabropezia flavovirens
(Fuckel) Dissing & Pfister
Lamprospora fide Denison
(1960)
Trichophaea brunnea (Alb.
& Schw.) Batra in L. R.
and S. T. Batra
T. confusa (Cooke) Berthet
T. hinnulea (Berk. &
Broome) Batra in L. R.
and S. T. Batra
T. monilifera (Berk. &
Curtis) Pfister
Pseudoplectania melaena
(Fr...) Sacc.. (1)
P. nigrella (Pers.)
Fuckel
Caloscypha fulgens (Pers.)
Boud.
Ruhlandiella berolinensis
P. Hennings
Boudiera areolata Cooke &
Phill.
B. echinulata (Seaver)
Seaver
Ascobolus nodulosporus
Brum.
Lamprospora miniata De
Not.
"L. laetirubra (Cooke)
Lagarde"
L. dictydiola Boud.
L. areolata Seaver
60
60
61
62
63
63
64
65
66
66
67
67
68
69
69
70
70
70
71
iz
72
ascoboloides
annulata
spinulosa
crec'hqueraultii
macrantha
brevispinosa
tuberculata
maireana
. wrightii
tuberculatella
amethystina
carbonaria
. haemastigma
constellatio
wisconsinensis
gemmea
discoidea
salmonicolor
trachycarpa
detonia
nigrans
L. ascoboloides Seaver
L. annulata Seaver
L. spinulosa Seaver
L. crec'hqueraultii
(Crouan & Crouan) Boud.
L. macracantha (Boud. )
Seaver
L. brevispinosa Seaver
L. tuberculata Seaver
L. maireana Seaver
L. wrightii (Berk. & Curtis
in Berk. & Br.) Seaver
L. tuberculatella Seaver
?Pulparia amethystina
(Quel. ) Donadini
Pulvinula carbonaria
(Fuckel) Boud.
P. convexella (P. Karst.)
Pfister
P. convexella (P. Karst.)
Pfister
P. laeterubra (Rehm)
Pfister
P. archeri (Berk. & J. D.
Hooker) Rifai
P. discoidea (P. Henn. &
E. Nym.) Batra
P. salmonicolor (Seaver)
Pfister
Plicaria trachycarpa
(Curr. ) Fuckel
Scabropezia flavovirens
(Fuckel) Dissing & Pfister
Plicaria nigrans (Morgan)
Pfister (2)
73
74
74
73
75
76
77
78
a2
80
82
83
83
84
85
85
86
86
87
87
58
89
89
90
L.
Tes
leiocarpa
planchonis
lobata
exasperata
polytrichina
Pithya pithya
P.
Pe
lacunosa
cupressi
Ascodesmis microscopica
A,
porcina
Ascobolus stercorarius
immersus
glaber
winteri
americanus
leveillei
viridulus
candidus
magnificus
carbonarius
geophilus
subglobosus
albinus
xylophilus
Plicaria endocarpoides
(Berk.) Rifai
Pulparia planchonis (Dun.
ex Boud.) Korf, J. K.
Rogers, Pfister
Lamprospora lobata (Berk.
& Curtis) Seaver
Plicaria recurva (Berk. )
Rifai
Lamprospora polytrichina
(Rehm) Seaver
Pithya vulgaris Fuckel
P,. vulgaris Fuckel
P. cupressina (Batch)
Fuckel (3)
Ascodesmis sphaerospora
Obrist (4)
A. porcina Seaver
Ascobolus furfuraceus
(Pers.) W. J. Hooke
A. immersus (Pers.) Pers.
Saccobolus glaber (Pers.)
Lambotte
A. elegans J. Klein
A. amoenus Oudemans
(5)
A. crenulatus P. Karst.
A. aglaosporus Heimerl
A. scatigenus (Berk.)
Brumm,
A. carbonarius P. Karst.
A. geophilus Seaver
A. subglobosus Seaver
A. albinus
A. xylophilus Seaver
90
A. striisporus
A. epimyces
atrofuscus
viridis
purpurascens
brunneus
cubensis
vinosus
.
major
[> [> [> I> I> I> I>
I>
conglomeratus
A. trifolii
A. pusillus
Saccobolus kerverni
S. portoricensis
S. violascens
S. depauperatus
S. globuliferellus
S. obscurus
Aleuria aurantia
A. wisconsinensis
A. cestrica
A. bicucullata
A. rhenana
A. foliicola Berk. &
Broome
A. epimyces (Cooke)
Seaver
A. carbinarius P. Karst.
(6)
(7)
(8)
A. cubensis Berk. &Curtis
(9)
A. scatigenus (Berk.)
Brumm,
Angelina rufescens (Schw.)
Inby fide Seaver (not a
member of the Pezizales).
Pseudopeziza trifolii
(not a member of the
Pezizales)
A. pusillus Boud. (10)
S. glaber (Pers.)
Lambotte
S. portoricensis Seaver
S. versicolor (P. Karst.)
P., Karst.
S. depauperatus (Berk. &
Broome) E. C. Hansen
S. globuliferellus Seaver
(rd)
Aleuria aurantia (Pers.)
Fuckel
A. aurantia
A. cestrica (Ellis &
Everh.) Seaver
A. bicucullata Boud.
A. rhenana Fuckel
100
101
102
103
104
104
106
106
107
107
108
108
Log
109
Lo
LEZ
Je
Lis
Hl
A. rutilans
Aleurina atrovinosa
A. aquehongensis
Melastiza charteri [sic]
M. asperrima
Melastiza pennsylvanica
Psilopezia nummularia
P. hydrophila
P. deligata
P. trachyspora
P. flavida
P. aquatica
P. mirabilis
Pyronema omphalodes
P. leucobasis
Ascophanus Holmskjoldii
A. isabellinus
A. sarcobius
A. bermudensis
Leucoscypha rutilans
(Fr.) Dennis & Rifai
Peziza atrovinosa Gerard
(12)
Melastiza chateri (W. G.
Smith) Boud.
Scutellinia asperrima
(Ell. & Everh. ex Seaver)
Le Gal
S. pennsylvanica (Seaver)
Denison
(13)
Pachyella hydrophila
(Peck ex Sacc.) Pfister
Psilopezia deligata
(Peck) Seaver
Pachyella adnata (Berk.
& Curtis) Pfister
Acervus flavidus (Berk.
& Curtis) Pfister
Miladina lechithina
(Cooke) Svrtek
Aleurodiscus, a Basidio-
mycete
Pyronema omphalodes
(Bull.) Fuckel
Psilopezia deligata
(Peck) Seaver
Thecotheus? cinereus
(Crouan & Crouan) Chenan-
tais
T. cinereus (Crouan &
Crouan) Chenantais
Iodophanus sarcobius
(Boud.) Kimbr.
I. bermudensis (Seaver)
Kimbr.
114
114
115
bm Be)
116
A le
cat
118
118
bah
120
120
120
a2.
121
124
124
124
A. vicinus
A. lacteus
A. argenteus
A, aurora
A. carneus
A. granulatus
A. granuliformis
A. ochraceus
A. gallinaceus
A. cinereus
A. tetraonalis
A. lilacinus Cooket
A. cinerellus (Karst. )
Speg.t
A. microsporus
A. cesatii
A. glaucellus
Humarina axillaris
H. ithacaensis
H. pallens
(14)
Coprotus lacteus (Cooke &
Phill.) Kimbr.,
Luck-Allen, and Cain
Coprotus argenteus (Curr.)
Kimbr., Luck-Allen, and
Cain
C. aurora (Crouan &
Crouan) Kimbr.,
Luck-Allen and Cain
Iodophanus carneus (Pers.)
Korf
Peziza humosoides in syn.
= immature Iodophanus
Coprobia granulata (Bull.)
Boud.
Coprotus granuliformis
(Crouan & Crouan) Kimbr.
C. ochraceus (Crouan &
Crouan) Larsen
(15)
Thecotheus cinereus
(Crouan & Crouan)
Chenantais
(16)
(17)
(18)
Coprotus glaucellus (Rehm)
Kimbr.
Octospora axillaris
(Nees) Moser
(19)
Thecotheus pallens (Boud.)
Kimbr. (20)
125
fa)
127
i127
L27
128
128
129
130
130
i3t
13}
pas
132
132
Lao
135
134
134
134
135
136
clausat
testacea
orthotricha
Glenenteit”
rubens
convexulat
lilacina
leucoloma
araneosa’
semiimmersa
ochroleucat
rufat
wisconsinensis
nectriodes
deligata
glumarumt
peckii
H. phyllogena
tetraspora
permuda |sic]
ollaris*
- aggregata
Iodophanus testaceus
(Moug. in Fr.) Korf
Octospora orthotricha
(Cooke & Ell.) Khare &
Tewari
Octospora sp.
O. rubens (Boud.) Moser
in Gams
Octospora sp.
(21)
Octospora leucoloma
Hedwig and 0. Humosa (Fr.)
Dennis included in
synonymy
Geopyxis carbonaria
O. semiimmersa (Karst.)
Khare & Tewari
Iodophanus testaceus
(Moug. in Fr.) Korf (22)
I. testaceus (23)
Octospora peckii (House)
Khare & Tewari
(24)
Byssonectria tetraspora
(Fuckel) Korf
B. fusispora (Berk.)
Rogerson and Korf in Korf
Byssonectria aggregata
(Berk. & Broome) Rogerson
& Korf
10
136
137
137
138
138
139
139
140
140
140
140
140
140
140
140
140
141
141
141
141
141
142
143
H. fusispora
H. coccinea
H. cookeina
H. gerardi
H. purpurea
H. hepatica
H. trachyderma*
Humaria bella
[x
- scatigena
salmonicolort
olivatrat
[a |
+
fuscocarpa
vitigenat
subcrenulatat
.
Im [me fm |x
secretat
groenlandica*
iat ie
microspora
H. xanthomela americana
Peziza elachroa
Humaria subgranulata
Pseudombrophila deerata
Streptotheca crouani
S. woolhopensis
B. fusispora (Berk. )
Rogerson and Korf in Korf
Octospora coccinea
(Crouan & Crouan) Brumm,
Hymenoscyphus sp.
Peziza gerardi Cooke and
Gerard
(25)
Fimaria hepatica (Batsch)
Brumm,
Nanoscypha bella (Berk.
& Curtis) Pfister
Ascobolus scatigenus
(Berk. & Curtis) Brumm.
an inoperculate disco-
mycete
Acervus epispartius
(Berk. & Curtis) Pfister
Peziza elachroa Berk. &
Curtis
Iodophanus subgranulatus
(Berk, & Curtis) Pfister
Pseudombrophila deerata
(Karst.) Seaver
Ascozonus crouani (Renny)
Schroet.
A. woolhopensis (Berk. &
Broome) Boud,.
143
144
145
145
146
147
147
148
149
150
151
152
152
ios
154
155
156
159
160
Ss.
obscura
Ryparobius sexdecimi-
sporus
Re
R.
R.
R.
crustaceus
hyalinellus
polysporust
monascus
niveus
Thecotheus pelletieri
Sepultaria arenicola
Sis
S.
Se
S.
longii
arenosa
aurantia
pellita
Pseudopithyella
minuscula
Perrotia flammea
Lasiobolus equinus
L.
ruber
Patella scutellata
Ps
pilisetat
11
Thelebolus obscurus
(Seaver) Eckblad
Coprotus sexdecimisporus
(Crouan & Crouan) Kimbr.
Thelebolus crustaceus
(Fuckel) Kimbr.
Thelebolus sp.
Thelebolus sp.
Thelebolus stercoreus
(Tode) Fr.
Coprotus niveus (Fuckel)
Kimbrough, Luck-Allen and
Cain
Thecotheus pelletieri
(Crouan & Crouan) Boud.
Geopora sepulta (Fr.)
Korf & Burdsall
G. longii (Seaver)
Burdsall & Korf
G. arenosa (Fuckel) Ahmad
Scutellinia aurantia
(Clements) Waraitch (26)
Geopora pellita (Cooke &
Peck) T. Schumacher
Pseudopithyella minuscula
(Boud. & Torrend) Seaver
An inoperculate disco-
mycete
Lasiobolus equinus (Mil11.)
P. ReEst.
L. ruber (Quel.) Sacc.
Scutellinia scutellata
(L) Lambotte
12
160
161
162
162
163
163
164
165
165
166
166
167
167
168
169
270
171
re es
172
173
cubensis
umbrorum
lusatiae
miniatat
albocincta
ovilla
hetieri
ricciophila
puniceat
gilva
setosa
sequoiae
melaloma
crucipila
stercorea
theleboloides
coprinaria
maculosa*
pulcherrima
raripila
Scutellinia cubensis
(Berk. & Curtis)
Gamundi
S. texensis (Berk. &
Curtis) Le Gal
S. umbrorum (Fr.)
Lambotte
S. lusatiae (Cooke) 0.
Kuntze
Leucoscypha vivida (Nyl.)
Dennis
L. ovilla (Peck) Harmaja
L. hetieri (Boud.) Rifai
(27)
Tricharina gilva (Boud.)
Eckblad
Scutellinia setosa (Nees)
Kuntze
(28)
Anthracobia melaloma
(Albertini & Schw.) Boud.
Cheilymenia crucipila
(Cooke & Phill.) Le Gal
in Denison
C. ciliata (Bull.) Maas
C. theleboloides (Alber-
tini & Schw.) Boud.
C. coprinaria (Cooke)
Boud. (30)
C. pulcherrima (Crouan &
Crouan) Boud.
C. raripila (Phill.)
Dennis
173. ~P. fimetariat
174 P. paludosa
174 tuberculata
175 P. albida
176 P. gregaria
177 Ps irregularist
1 WY av P. abundans
178 P. erinaceus
178 P. albospadicea
179 P. pygmaea
17 Lachnea vitellina
179 L. crispata
180 Neottiella spraguei
180 Lachnea lividat
180 Neottiella albotecta
180 Lachnea hirta
180 Neottiella sericeovillosat
180 Lachnea fissilist
180 Neottiella callichroa
180 Neottiopezis macrosporat
EOL Wynnea gigantea
181 W. americana
i3
Trichophaea paludosa Boud.
T. tuberculata (Seaver)
Kanouse
Humaria hemispherica
(Wigg.) Fuckel
Trichophaea gregaria
(Rehm) Boud.
T. abundans (P. Karst.)
Boud.
Scutellinia erinaceus
(Schw.) Kuntze
Trichophaea albospadicea
(Grev.) Boud. -- a species
complex
Trichophaea pygmaea
(Clements) Waraitch (31)
Cheilymenia vitellina
(Pers.) Dennis
Phillipsia crispata (Berk.
& Curtis) Le Gal
Peziza spraguei Berk. &
Curtis
a lichen
Scutellinia hirta (Schum. )
Lamb.
Octospora callichroa
(Boud.) Arpin
Wynnea gigantea Berk. &
Curtis
W. americana Thaxter
14
182 Phillipsia domingensis Phillipsia domingensis
Berk.
183. P. gigantea P. domingensis
183 P. chardoniana P. domingensis
184 P. dochmia Aurophora dochmia (Berk.
& Curtis) Rifai
185 Scodellina leporina Otidea leporina (Fr.)
Fuckel
185 S. auricula Wynnella silvicola (Berk.
in Sacc.) Nannf.
186 S. grandis Otidea grandis (Pers. )
Rehm
+
187 Otidea obtecta
187 Cookeina tricholoma Cookeina tricholoma
(Mont.) Kuntze
188 C. sulcipes C. sulcipes (Berk.)
Kuntze
189 C. tetraspora Nanoscypha tetraspora
(Seaver) Denison
190 C. colensoi C. venezuelae (Berk. &
Curtis) Le Gal (Denison
1967)
191 Plectania coccinea Sarcoscypha coccinea
(Scop.) Sacc.
192 P. floccosa Microstoma floccosum
(Schw.) Raitviir
193. +P. occidentalis Sarcoscypha occidentalis
(Schw.) Sacc.
193 P. hiemalis Microstoma protractum
(Fr.) Kanouse
194 Sarcoscypha roseotincta*
195 Bulgaria globosa Sarcosoma globosum (Fr.)
Casp. in Rehm
196 B. rufa Galiella rufa (Schw.)
Nannf. & Korf
197 B. melastoma Plectania melastoma (Sow.)
Fuckel
204
205
205
206
207
208
208
209
210
7!
a.
211
212
Peziza aurantiopsis
Urnula craterium
U.
U.
geaster
terrestrist
Paxina acetabulum
P.
Py
P..
Macropodia schweinitziit
macropus
platypodia
leucomelas
sulcata
barlae
hispida
subc lavipes
dupainii
corium
nigrella
semitosta
fusicarpa
olivaceat
Plectania rimosa Peckt
Geopyxis cupularis
15
Wolfina aurantiopsis
(Ellis) Eckblad
Urnula craterium (Schw.)
Fr.
Chorioactis geaster
(Peck) Eckblad
Helvella acetabulum (L.)
Quel.
H. macropus (Pers.) P.
Karst.
H. queletii Bres.
H. leucomelas (Pers.)
Nannf.
H. solitaris (P. Karst.)
P, Karst,
H. acetabulum (L.) Quel.
H. villosa (Hedw.)
Dissing & Nannf.
H. macropus (Pers.) P.
Karst.
H. queleti Bres.
H. corium (Weberb. )
Massee
Plectania nannfeldtii
Korf
Jafnea semitosta (Berk. &
Curtis) Korf
J. fusicarpa (Gerard)
Korf
Tarzetta cupularis (L.)
Korf and J. K. Rogers
16
213
213
214
214
214
214
214
y i He
215
216
216
217
218
218
221
222
222
223
224
224
yaa)
Pa ae
226
op)
Ia |Q [a [a
bronca
- catinus
vulcanalis
nebulosa*
nebulosoidest
cinerascenst
verruculosa
Hicksiit
Rhizina inflata
R. spongiosa
Discina ancilis
D.
D.
D.
apiculata
convoluta
leucoxantha
Peziza badia
. brunneoatra
spissa
succosa
pustulata
emileia
melaleuca
melaleucoides
violacea
T. bronca (Peck) Korf &
J. K. Rogers
T. catinus (Holmsk.) Korf
and J. K. Rogers
Geopyxis vulcanalis
(Peck) Sacc.
Helvella acetabulum (L.)
Quel.
Rhizina undulata Fr.
Galiella spongiosa (Berk.
& Curtis) Pfister
Gyromitra perlata (Fr.)
Harmaja
Peziza apiculata Cooke
Gyromitra leucoxantha
(Bres.) Harmaja
G. leucoxantha (Bres.)
Harmaja
Peziza badio-confusa
Korf (32)
P. brunneoatra Desmaz.
P. spissa Berk.
P. succosa Berk.
P. petersii Berk.
P. emileia Cooke
Gyromitra melaleuca
(Bres.) Donadini
G. melaleucoides (Seaver)
Pfister
Peziza praetervisa Bres.
(33)
va
re |
P. griseorosea
P. proteana
P. concentrica
P. abietina
P. venosa
. domiciliana
- repanda
P
2
P. vesiculosa
P. fimeti
P
- sylvestris
P. clypeata
P. caeruleomaculata
Humaria lacteocinereat
Peziza brunneovinosa
‘ chrysopelat
+
roseolilacina
caliganst
irrorata
[ho [rd [to |
|td
repandoides
P. irregularis*
P. convoluta
Sarcosphaera coronaria
Morchella crassipes
M. esculenta
Li
P. griseorosea Gerard
P. proteana (Boud.)
Seaver
P. concentrica Seaver
Otidea propinquata
(Karst.) Harmaja
Disciotis venosa (Pers.)
Boud,
Peziza domiciliana Cooke
P, repanda Pers.
P. vesiculosa Bull.
P. fimeti (Fuckel) Seaver
P. sylvestris (Boud.)
Sacc.. -& Trett.
Pachyella clypeata (Schw.)
Le Gal
Peziza caeruleomaculata
Rehm (34)
Peziza michelii (Boud.)
Dennis
P. irrorata Berk. &
Curtis
P. repandoides (Rehm)
Sacc. & Trott.
P. vesiculosa Bull.
Sarcosphaera coronaria
(Jacq.) Schroet. in Cohn
Morcella crassipes (Vent.)
Pers,
M. esculenta (L.) Pers.
18
239
240
240
241
242
243
244
245
245
246
247
247
248
249
249
250
250
251
251
252
253
253
254
254
255
M. conica
M. deliciosa
M. angusticeps
M. hybrida
Durandiomyces phillipsii
Verpa conica
V. bohemica
Y. atroalbat
V. perpusillat
Elvela mitra
E. crispa
E. palustris
- atra
- elastica
E
E
E. klotzschiana
E. adhaerens
E
- californica
ica
- umbraculiformis
- infula
. +
brevissima
. albipes
SH HH mH
. caroliniana
E. underwoodii
E. sphaerospora
Underwoodia columnaris
M. conica Pers.
M. deliciosa Fr.
M. angusticeps Peck
M. semilibera DC.
Peziza proteana forma
sparassoides (Boud. ) Korf
Verpa conica (Mull.)
Swartz
V. bohemica (Kromb.)
Schroet in Cohn
Helvella lacunosa Afzel.
H. crispa (Scop.) Fr.
H. lacunosa Afzel
H. atra Oed.
H. elastica Bull.
H. elastica Bull.
H. elastica Bull.
Gyromitra californica
(Phill.) Raitviir
G. californica (Phill.)
Raitviir
G. infula (Schaeff.) Quel.
Helvella leucopus Pers.
Gyromitra caroliniana
(Fr.) Fr.
G. brunnea Underw.
Pseudorhizina sphaero-
spora (Peck) Pouz.
Underwoodia columnaris
Peck
304
306
306
307
SLL
314
315
318
a
325
326
332
x Ps
333
337
19
New taxa listed from the revised section
Boudiera walkerae
Lamprospora sphagicola
P. pyrophila
Cubonia bulbifera
Humarina waterstonii
Patella contradicta
P. michiganensis
Sarcoscypha imperialis
Wolfina aurantiopsis
Paxina compressa
P. recurva
Peziza pseudoclypeata
P. waltersii
Sarcosphaera ammophila
Daleomyces phillipsii
Boudiera walkerae Seaver
Lamprospora sphagicola
Seaver
?Pulvinula archeri (Berk.
in Hook.) Rifai
Nanoscypha waterstonii
(Seaver) Pfister (35)
Trichophaea contradicta
(Seaver) H. J. Larsen
Cheilymenia coprinaria
(Cooke) Boud.
Sowerbyella imperialis
(Peck) Korf
Wolfina aurantiopsis
(Ellis) Eckblad
Helvella compressa
(Snyder) Weber
Gyromitra melaleucoides
(Seaver) Pfister
Pachyella adnata (Berk.
& Curtis) Pfister
Peziza waltersii Seaver
Peziza ammophila Dur. &
Lév.
Peziza proteana forma
sparassoides (Boud.) Korf
20
NOTES REFERRED TO IN PART I
(1) This species is still widely known as Pseudoplectania
vogesiaca. Though the name Peziza vogesiaca was proposed in
1818, Fries in 1822 did not use it but rather proposed Pez-
iza melaena and listed P. yogesiaca as a synonym. Under the
ICBN, Fries's name should be used. Saccardo and Boudier
treated the species in the genus Melascypha.
(2) Plicaria nigrans (Morgan) Pfister comb. nov. = Peziza
nigrans Morgan, J. Cinn. Soc. Nat. Hist. 18: 43. 1895. The
Plicarias in North America have never been completely worked
out. Lamprospora nigrans is a distinctive Plicaria. The
asci and hymenium blue faintly, the paraphyses are agglu-
tinated, the spores are heavily warted and are 7-8 um.
(3) Pithya cupressina and P. vulgaris have been considered
identical by many recent authors but Denison (1972) thought
them distinct.
(4) Ascodesmis microscopica (Crouan & Crouan) Seaver was
based on material with sphaerical spores which Obrist (1961)
renamed A. sphaerospora.
(5) According to van Brummelen's (1967) study of type speci-
mens, Ascobolus leveillei is correctly named A. boudieri
Quél, but it is known only from Europe. It is not clear to
which species Seaver was referring.
(6) Ascobolus viridis, according to van Brummelen (1967),
has not been recorded from North America. This was also
Seaver's decision when he redescribed the species as A. geo-
hilous, an accepted species.
(7) This may refer to a misprint of "furfuraceus."
(8) According to van Brummelen's (1967) study of the type
specimen, this is a species of Rhyparobius. There is no re-
port of the species from North America.
(9) No North American collections are definitely known under
this name.
(10) van Brummelen (1967) has studied the Dodge specimen re-
ferred to by Seaver and finds it to be Ascobolus pusillus.
21
It is the only collection of this species which van Brum-
melen reported from North America.
(11) Seaver reported the name from Clements! Crypto. Form.
Colo. no. 301 but excluded it because no specimens could be
found. Van Brummelen (1967) accepts the species but lists
no North American collections.
(12) This and several other species are very like Peziza
badia. The apothecia are dark colored and the spores are
reticulate. The detailed study necessary to properly place
taxa in this complex has not yet been done.
(13) Psilopezia nummularia is known from North America but
the description Seaver gives seems to refer to Pachyella ba-
bingtonii, a name listed by Seaver among the synonyms but
considered distinct by Pfister (1973a; 1973b).
(14) The specimen upon which Seaver based his conclusions
does not seem to have been recently examined. Van Brummelen
(1967) suggests that this is close to Ascophanus granuli-
formis (= Coprotus granuliformis).
(15) The type collection was examined but no ascocarps could
be found.
(16) Type material was examined. The species is very close
to Peziza fimeti; it differs in the habitat, partridge or
grouse dung, and the ascospore size. The ascospores are
16-18 um in length, a range which is slightly larger than
that of Peziza fimeti.
(17) Type material has not been seen. Van Brummelen (1967)
states that this is a very common species of "Ascophanus."
(18) Van Brummelen (1967) says this is Ascophanus micros-
porus (Berk. & Broome) E. C. Hansen.
(19) Rifai (1968) briefly discussed this taxon, which has
been variously placed in the Pezizales, he writes, "Neot-
tiella crozalsiana, Humaria ithacaensis and Peziza ricciae
differ from the other species of Leucoscypha in their small
(usually less than 1.3 jim diam.) apothecia, in the poorly
developed medullary excipulum and in the _ possession of
vegetative mycelium of a mildew, which is probably due_ to
their peculiar habitat, growing directly on hepatics rather
than among mosses." Rifai suggests a separate genus. North
22
American materials need to be critically examined. I do not
agree with the inclusion of this species in OQctospora as
done by Khare (1975).
(20) Kimbrough (1969) stated that Thecotheus pallens was not
known in North America but he had overlooked Seaver's report
of it under the genus Humarina. The specimen in NY which
was examined agrees with Boudier's original description.
The large spores which reach a length of more than 40 um and
unusual habitat, on soil among mosses, makes this a distinc-
tive species. The habitat is unusual for a Thecotheus which
has been known to occur primarily on dung and occasionally
on water soaked wood; T. phycophilus Pfister is similar (see
Pfister 1981).
(21) Type material has been examined but is fragmentary. No
conclusive placement could be made.
(22) I have examined Seaver's type material and found
apothecia of Iodophanus testaceus.
(23) See Pfister (1979c)
(24) This is a member of the Sarcoscyphaceae, perhaps be-
longing to an undescribed genus. Khare and Tewari (1978)
placed it in Octospora, a position with which I cannot
agree.
(25) Material from the type locality shows asci which are
faintly J+. Little material is available but based on
Seaver's slide and description, this is most likely a small
species of Peziga.
(26) I have not examined the material of Clements! type
which was apparently not examined by Waraitch (1977) when he
combined Clements! name in Scutellina.
(27) See note 21. Seaver's name is a later synonym being
predated by Peziza ricciae Crouan and Crouan.
(28) Denison (1959) could not properly place this species.
(29) There is a difference of opinion; see Maas Geestranus
(1969).
(30) Cheilymenia dalmeniensis placed by Seaver in synonymy
has been recognized as a distinct species.
Zo
(31) Material has been examined but resolution of generic
placement remains. It differs from Trichophaea.
(32) Both Peziza badia and P. badio-confusa exist in North
America. Elliot & Kaufert (1974) discuss these and cite
various collections.
(33) Peziza praetervisa has lightly ornamented spores as
described by Seaver for P. violacea. The spores of P.
violacea are completely smooth. Though P. violacea exists
in North America, it is infrequently collected. Most col-
lections under that name are P. praetervisa.
(34) See Pfister (1979b)
(35) Khare & Tawari (1978) included this species in Octo-
spora, a view with which I cannot agree.
24
PART IL
Morchellaceae H. macropus p. 203, 206
pissiocia H. queletii p. 203, 207
Te aaa 929 H. solitaris p. 204
ene _ H. villosa
Morchella Pseudorhizina
Hy ae Beaee Re. De aay P. sphaerospora p. 254
M. conica p. 239
M. crassipes p. 23/7 Rhizina
M. deliciosa p. 240 R. undulata p. 215
ey — ea Underwoodia
a Ps U. columnaris
Verpa
V. bohemica p. 244 iba
V. conica p. 243 W. silvicola p. 185
oo oo eee Ascobolaceae
Ascobolus ref. Van
Brummelen (1967)
A. aglaosporus p. 86
A. albinus p. 89
A. amoenus p. 85
A. carbonarius p. 87,
92
Gyromitra ref. Kempton &
Wells (1973) McKnight (1969,
1971, 1973) and Pfister
(1980).
G. brunnea p. 254
G. californica p. 251
carolinae p. 253
G.
G. anfula.- oy 251 A. crenulatus p. 86
A. cubensis p. 92
G. leucoxantha p. 218
A. elegans p. 84
G. melaleuca p. 225
G. melaleucoides 225 soe
. ee , A. foliicola p. 90
326
G. pevlata p. 216 A. furfuraceus p. 82
: A. geophilus p. 88
Helvella ref. Weber (1972, A. immersus p. 83
1975) and Dissing (1966) A. nodulosporus p. 53
Dissing & Lange (1967) Kemp- A. pusillus p. 92
ton & Wells (1970). A. scatigluses p. 87,
H. acetabulum p. 202, 205, 92, 140
214 A. subglobosus p. 89
H. atra p. 248 A. xylophilus p. 90
ee aa ca Saccobolus ref. Van Brum-
H. corium p. 208
melen (1967)
H. crispa p. 247
; S. depauperatus p. 95
H. elastica p. 249, 250
S. @leber O. 835.93
H. lacunosa p. 246, 247
S. globuliferellus
H. leucomelas p. 204 95
H. leucopus p. 253 Ps
S. obscurus p. 96
S. portoricensis p. 94
S. versicolor p. 94
Pezizaceae
Boudiera ref. Dissing
& Schumacher (1979)
B. areolata p. 52
B. echinulata p. 52
B. walkerae p. 304
1979)
Iodophanus ref. Kimbrough,
Luck-Allen, and Cain (1969)
I. bermudensis p. 113
I. carneus p. 115
I. sarcobius p. 113
I
I
« Bestdeeue p,. 125, 132
. subgranulatus p. 141
Pachyella ref. Pfister (1973b)
Fr. sonate ps Ls, 332
P. babingtonii p. 106
P. clypeata p. 233
P. hydrophila p. 106
Peziza
P. ammophila p.
P. apiculata p. 217
atrovinosa p. 101
badio-confusa p. 101
brunneoatra p. 222
caeruleomaculata p.
concentrica p. 228
domiciliana p. 230
elachroa p. 141
emileia p. 224
fimeti p. 232
gerardii p. 138
grislorosea p.
irrorata p. 235
michelii p. 234
petersii p. 224
praetervisa p.
P. proteana p. 227
var. sparassoides
Hs 262,..33)
P. repanda p.
333
234
227
guowitir sew
226
eal
25
P. repandoides p. 235
P, spissa p. 222
P. spraguei p. 180
P., succosa p. 223
P. sylvestris p. 233
P. vesiculosa p. 231
P. waltersii p. 332
Plicaria
P. endocarpoides p. 73
P, nigrans p. 72
P. recurva’ pb. 75
P, trachycarpa p. 71
Ruhlandiella ref. Dissing
and Korf (1980)
R. berolinensis p. 51
Sarcosphaera
S. crassa p. 235
Scabropezia ref. Dissing
and Pfister (1981)
S. flavovirens p. 44,
72
Thecotheus ref. Kimbrough
(1969)
T. cinereus p. 112, 118
T. pelleturi p. 148
T. pallens p. 124
Acervus ref. Pfister
(1975a)
A. epispartius p. 141
A. flavidus p. 108
Aleuria
A. aurantia p. 97, 98
A. bicucullata p. 99
A. cestrica p. 98
A. rhenana p. 99
Anthracobia
A. melaloma p. 167
Ascodesmis ref. Obrist
(1961)
A. porcina
A. sphaerospora
26
Ascozonus
Seaver treated two species:
A. crouani p. 142
A. woolhopensis p. 143
Byssonectria ref. Dennis &
Itzerott (1973) under the
name Inermisia.
B. aggregata p. 136
B. fusispora p. 134, 136
B. tetraspora p. 134
Caloscypha
C. fulgens p. 50
Cheilymenia ref. Denison
(1964)
C. ciliata p. 169
C. coprinaria p. 171, 315
C. crucipila p. 168
C. pulcherrima p. 172
C. raripila p. 173
C. theleboloides p. 170
C. vitellina p. 179
Coprobia
C. granulata p. 116
Coprotus ref. Kimbrough,
Luck-Allen and Cain (1972)
C. argenteus p,. 114
- aurora p. 115
- glaucellus p. 121
granuliformis p.
- lacteus p. 114
- niveus p. 147
- ochraceus pp.
C. sexdecmisporus
117
AAQAQAAANOD
117
p. 144
Fimaria ref. Van Brummelen
(1962)
F. hepatica (p. 139 only)
Geopora ref. Burdsall (1968)
G. arenosa p. 151
G. longii p. 150
G. pellita p. 152
G. sepulta p. 149
Geopyxis
G. vulcanalis p. 214
Humaria
H. hemispherica p.
Jafnea ref. Korf (1960)
J. fusicarpa p. 210
J. semitosta p. 209
Lamprospora
- annulata p. 60
- areolata p. 59
L
L
L. ascoboloides p. 60
L. brevispinosa p. 63
L
. crec'hqueraultii
p. 62
L. dictyiola p. 58
. laetirubra p. 58
lobata p. 74
Macrantha p. 63
maireana p. 65
miniata p. 56
sphagnicola p.
spinulosa p. 61
tuberculata p. 64
tuberculatella p.
L. wrightii p. 66
CPHerr rr ee er
Lasiobolus ref. Bezerra
& Kimbrough (1975)
L. equinus p. 155
L. ruber p. 156
Leucoscypha
L. albocincta p.
L. hetieri p. 164
L. ovilla p. 163
L. rutilans p. 100
163
Melastiza ref. Lassueur
(1980)
M. chateri p. 103
Miladina ref. Pfister &
Korf (1974)
M. lechithina p. 108
Octospora ref. Khare &
polytrichina p. 75
306
175
66
Tewari (1978), Dennis and
Itzerott (1973).
O. axillaris p. 124
O. callichroa p. 180
O. coccinea p. 137
O. leucoloma p. 129
QO. orthotricha p. 127
O. peckii p. 133
O. rubens p. 127
O
semi-immersa p. 130
Otidea ref. Kanouse (1950)
O. grandis p. 186
O. leporina p. 185
O. propinquata p. 228
Pseudombrophila
P, deerata p.
141
Psilopezia ref. Pfister
(1973a)
P. deligata p. 107, 110
Pulparia ref. Donadini (1976)
P. amethystina p. 67
P. planchonis p. 74
Pulvinula ref. Pfister (1975b)
Po areneri 2. 70
P. carbonaria p. 67
P. convexella p. 68
P. discoidea p. 70
P., laeterubra p. 69
P. slamonicolor p. 71
Pyronema ref. Moore & Korf
(1963)
P. omphalodes p. 109
Scutellinia ref. Denison (1959)
S. asperrima p. 104
aurantia p. 152
cubensis p. 160
erinaceous p. 178
hirta p. 180
lusatiae p. 162
pennsylvanica p.
scutellata p. 159
setosa p. 166
texensis p. 160
trechispora p. 43
umbrorum p. 161
104
AWMnNNNNNNNAHANN
.
27
Sowerbyella
S. imperialis p. 318
Tarzetta
iy Dronca ps Z2L5
T. catinus p.. 213
T. cupularis p. 212
Thelebolus
T. obscurus p. 143
T. stercoreus p. 147
Tricharia
T. gilva p. 166
Trichophaea ref. Kanouse
(1958)
T. abundans
p.- 177
T. brunnea p. 46
T. contradicta p. 314
T. gregaria p. 176
T. hinnulea p. 46
T. monilifera p. 47
T. paludosa p. 174
T. pygmaea p. 179
T. tuberculata p. 174
T. woolhopeia p. 178
Sarcosomataceae
Chorioactis
C. geaster p. 200
Galiella
G. rufa p. 196
G. spongiosa p. 216
Plectania ref. Korf (19562),
Paden & Tylutki (1969)
P. melastoma p. 197
P. nannfeldtii p. 208
Pseudoplectania
P. melaena p. 48
P. nigrella p. 49
Sarcosoma
S. globosum p. 195
Urnula ref. Kempton & Wells
(1974)
U. craterium p. 199
28
Wolfina
W. aurantiopsis p. 198,
$21
Sarcoscyphaceae
Aurophora
A. dochmia p. 184
Cookeina ref. Denison (1967)
C. colensoi p. 190
C. sulcipes p. 188
C. tricholoma p. 187
C. venezuelae p. 190
Microstoma ref. Kanouse
(1949)
M. floccosum p. 193
M. protractum p. 193
Nanoscypha ref. Denison
(1972)
N. bella p. 140
N. tetraspora p. 189
N. watersonii p. 311
Phillipsia ref. Denison
(1969)
P. crispata p. 179
P, domingensis p. 182, 183
Pithya ref. Denison (1972)
P. cupressina p. 78
P. vulgaris p. 76, 77
Pseudopithyella
P. minuscula p. 153
Sarcoscypha ref. Denison
(1972)
S. coccinea p. 191
S. occidentalis p. 193
Wynnea ref. Pfister (1979a)
W. americana p. 181
W. gigantea p. 181
29
PART III
Bezerra, J. L. and J. W. Kimbrough. 1975. The genus
Lasiobolus (Pezizales, Ascomycetes). Can. J. Bot. 53:
1206-1229.
Brummelen, J. van. 1962. Studies on Discomycetes. II. On
four species of Fimaria. Persoonia 2: 321-330.
- 1967. A world monograph of the genera Ascobolus and
Saccobolus (Ascomycetes, Pezizales). Persoonia
(Suppl.) 1: 1-260.
Burdsall, H. H., Jr. 1968. A revision of the genus Hydno-
cystis (Tuberales) and of the hypogeous species of Geo-
pora (Pezizales). Mycologia 60: 496-525.
Cain, R. F. and J. W. Kimbrough. 1969. Coprobolus, a new
genus of the tribe Thelebolus (Pezizales). Can. J.
Bot. 47: 1911-1914.
Denison, W. C. 1959. Some species of the genus Scutel-
linia. Mycologia 51: 605-635.
__- 1964. The genus Cheilymenia in North America.
Mycologia 56: 306-317.
__+ 1965. Central American Pezizales. I. A new genus of
the Sarcoscyphaceae. Mycologia 57: 649-656.
- 1967. Central American Pezizales. II. The genus
Cookeina. Mycologia 59: 306-317.
- 1969. Central American Pezizales. III. The genus
Phillipsia. Mycologia 61. 289-304.
- 1972. Central American Pezizales. IV. The genera
Sarcoscypha, Pithya, and Nanoscypha. Mycologia 64:
609-623.
Dennis, R. W. G. 1978. British Ascomycetes. Revised Edi-
tion. J. Cramer. 585 pp.
and H. Itzerott. 1973. Octospora and Inermisig in
Western Europe. Kew Bull. 28: 5-23.
Dissing, H. 1966. The genus Helvella in Europe with spe-
cial emphasis on the species found in Norden. Dansk
Bot. Arkiv 25(1): 1-172.
and R. P. Korf. 1980. Preliminary studies in the
genera Ruhlandiella, Sphaerosoma, and Sphaerozone
(order Pezizales). Mycotaxon 12: 287-306.
and M. Lange. 1967. Notes on the genus Helvella in
North America. Mycologia 59: 349-360.
and D. H. Pfister. 1981. Scabropezia, a new genus of
Pezizaceae (Pezizales). Nord. J. Bot. 1: 102-108.
and T. Schumacher. 1979. Preliminary studies in the
genus Boudiera, taxonomy and ecology. Norw. J. Bot.
26: 99-109.
30
Donadini, J.-C. 1976. Le Genre Pulparia Karsten en France.
Rev. Mycol. (France) 40: 255-272.
Eckblad, F. ~E. 1968. The genera of the operculate
discomycetes, a re-evaluation of their taxonomy, phy lo-
geny, and nomenclature. Nytt Mag. Bot. 15: 1-191.
Elliot, M. E. and M. Kaufert. 1974. Peziza badia and Pez-
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