580,5
FB
V,2
1900-1916
cop, 2
Inc.
-Op, K. FIELD COLUMBIAN MUSEUM
PUBLICATION 126
BOTANICAL SERIES VOL. II, No. 6.
NEW OR NOTEWORTHY
SPERMATOPHYTES
FROM
MEXICO, CENTRAL AMERICA
AND THE WEST INDIES
BY
JESSE MORE GREENMAN, PH. D.
Assistant Curator, Department of Botany.
CHARLES FREDERICK MILLSPAUGH, M. D.
Curator, Department of Botany.
CHICAGO, U. S. A.
December, 1907.
FIELD COLUMBIAN MUSEUM
PUBLICATION 126
BOTANICAL SERIES VOL. II, No. 6
NEW OR NOTEWORTHY
SPERMATOPHYTES
FROM
MEXICO, CENTRAL AMERICA
AND THE WEST INDIES
BY
JESSE MORE GREENMAN, PH. D.
Assistant Curator, Department of Botany.
CHARLES FREDERICK MILLSPAUGH, M. D.
Curator, Department of Botany.
CHICAGO, U. S. A.
December, 1907.
New or Noteworthy Spermatophytes from Mexico,
Central America, and the West Indies.
By J. M. GREENMAN.
The diagnoses and notes here presented are the results of critical
study in the determination of several recent collections of plants
from Mexico, Central America and the West Indies, particularly
those of Mr. Edward A. Goldman, Professor Cassiano Conzatti, Dr.
George F. Gaumer, Professors C. R. Barnes, G. J. Chamberlain and
W. J. G. Land, Dr. Charles F. Millspaugh, Dr. J. N. Rose and assist-
ants, Sr. Dr. Fernando Altamirano, Professor W. A. Kellerman,
the late Dr. -G. M. Emrick, Mr. H. A. Van Hermann, and several
others including the writer. The material of certain groups, especially
in the genus Senecio collected by Mr. C. G. Pringle, has been gener-
ously submitted to me for identification by Professor B. L. Robinson.
The new species here proposed in this genus are preliminary to a
forthcoming monograph of the North American Senecios.
CYPERUS OCHRACEUS Vahl, Enum. ii. 325 (1805).
Specimens agreeing well with the original description and with
West Indian representatives of this species were collected at
Laguna, near the City of Vera Cruz, Mexico, 22 January, 1906,
J. M. Greenman, no. 30 (hb. Field Mus.). -This species seems
not to have been noted by Hemsley in the Biologia Centrali-
Americana.
Hechtia macrophylla Greenman, sp. nov.
Leaves about i m. in length, 3.5 cm. broad just above the base,
gradually tapering to the apex, glabrous above, lepidote-cinere-
ous beneath; margins spinose; spines 3 cm. or less apart, up-
wardly curved, 6 mm. or less in length, usually bearing a tuft of
persistent white floccose tomentum in the axils: inflorescence
paniculate, about 4 dm. long, 1.5 to 2 dm. broad, lepidote-
tomentulose; ultimate branches 2 to 12 cm. long, spicate, more
or less loosely but evenly flowered throughout their entire length ;
floral bracts ovate, acute, 4 mm. long: staminate flowers sessile,
about 5 mm. long in anthesis, spreading or reflexed; sepals
247
248 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
broadly ovate, 2.5 mm. long, acute; petals elliptic or elliptic-
obovate, about 4 mm. long, strongly concave, free or slightly
united at the base; ovary rudimentary: pistillate flowers and
fruit unknown. MEXICO. State of Vera Cruz: Carrizal, 12 to
14 May, 1901, E. A. Goldman, no. 712 (lib. U. S. Nat. Mus. ;
fragment and photograph in hb. Field Mus.).
The species here described is apparently nearest related to
Hechtia Schottii Baker, and H. texensis Watson; from the for-
mer it differs in having longer leaves, more profusely branched
inflorescence, and somewhat smaller floral bracts; from the
latter it is readily separated by the longer leaves, the presence
of conspicuous tufts of tomentum in the upper axils of the
leaf-spines, and by the smaller and more scattered flowers.
HECHTIA SCHOTTII Baker, in Hemsl. Biol. Cent. -Am. Bot. iii.
318 (1884) & Handb. Bromel. 139 (1889); Mez in DC., Monogr.
Phan. ix. 548 (1896).
In the herbarium of the Field Museum there is a specimen,
collected by Schott in Yucatan, which is unmistakably refer-
able to Hechtia. The label accompanying the plant bears no
number, and likewise no time of collection, but it bears the
data "Maxeana." The specimen consists of leaves, an inflores-
cence of staminate flowers, and a portion of a panicle bearing
mature fruit. The characters exhibited by all these parts
agree well with Baker's description, hence the plant is confi-
dently referred to the above species; and, moreover, it probably
represents a part of the same collection on which the species was
founded. With this species are also identified specimens collect-
ed at Xcholac, Yucatan, Dr. Geo. F. Gaunter, no. 578 (hb.
Field Mus.).
TILLANDSIA BALBISIANA Schult. f. in Roem. & Schult. Syst. vii.
1212 (1830); Mez in DC. Monogr. Phan. ix. 709 (1896). T.
setaeea, Millsp. Field Col. Mus. Bot. Ser. i. 356 (1898), not Sw.
To this species are referred the following. MEXICO. State
of Yucatan: Merida, n July, 1865, Dr. A. Schott, nos. 842,
842a in part (hb. Field Mus.); Izamal, 21 February, 1906,
J. M. Greenman, no. 403 (hb. Field Mus.).
TILLANDSIA BRACHYCAULOS Schlecht. Linnaea, xviii. 422 (1844);
Morr. Belg. Hort. 1878, 185, t. n; Baker, Handb. Bromel.
201 (1889); Mez in DC. Monogr. Phan. ix. 732 (1896); Millsp.
Field Col. Mus. Bot. Ser. i. 356 (1898).
Fruiting specimens of this species were collected by the writer
at Izamal, Yucatan, in February of 1906. These agree in habit
and foliar characters with flowering specimens which were
secured in the same locality by Dr. A. Schott and also by Dr.
Geo. F. Gaumer. The species is widely distributed, occurring
from Mexico to South America, and although well known from
flowering specimens, the fruit seems not to have been hitherto
described, hence the following characterization is here given:
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 249
Mature capsules large, 3.5 to 4 cm. long, subcylindrical or obtusely
triangular, sliort-acuminate at the apex; valves dorsally pale-
stramineous, glabrous, i-nerved, recurved and somewhat
spirally twisted ; exocarp readily separating from the endocarp ;
seeds including the coma about 3 cm. long. MEXICO. State
of Yucatan: Izamal, 21 February, 1906, /. M. Greenman, no.
404 (hb. Field Mus.).
TRADESCANTIA FLORIDANA Watson, Proc. Am. Acad. xvii. 381 (1882).
Tradescantella floridana Small, Fl. Southeastern U. S. 238
(1903)-
Dr. Sereno Watson very clearly defined the above species
and pointed out the characters by which it is readily distinguished
from T. gracilis HBK. to which it was referred by C. B. Clarke
in DC. Monogr. Phaner. iii. 297 (1881). The examination of
a considerable number of specimens from Florida and elsewhere
shows that Dr. Watson's species retains the distinctive characters
originally ascribed to it without any evidence, at least as far
as yet observed by the writer, of intergradation with the South
American species. It seems best therefore to regard T. flori-
dana Watson as well worthy of specific rank. The following
specimens are identical in every detail with Dr. Watson's species.
MEXICO. State of Yucatan: Izamal, Dr. Geo. F. Gaumer, no.
573 (hb. Field Mus.); Chichankanab, Dr. Geo. F. Gaunter, no.
1855 (hb. Field Mus.). This species has not been recorded
hitherto from Yucatan.
SMILAX MOLLIS Humb. & Bonpl. in Willd. Sp. PI. iv. 785 (1805);
A. DC. in DC. Monogr. Phaner. i. 67 (1878); Hemsl. Biol. Cent.-
Am. Bot. iii. 365 (1884).
Mature fruiting specimens of this species were collected on
old sand dunes along the shore, north of the City of Vera Cruz,
24 January, 1906, /. M. Greenman, no. 116 (hb. Field Mus.,
and hb. Kew). The mature fruit in the fresh state is bright
red.
I am indebted to Lieut. -Col. David Prain, Director of the
Royal Botanic Gardens, Kew, for the identification of this
plant.
POUZOLZIA PRINGLEI Greenm. Proc. Am. Acad. xxxiii. 476 (1898).
This species, hitherto known only by Mr. Pringle's no. 6736
from Tomellin Canyon, has been recollected at El Parion, Dis-
trict of Etla, Oaxaca, Mexico, altitude 1,400 m., 2 September,
1906, C. Conzatti no. 1551 (hb. Field Mus.). While Senor Con-
zatti's specimens present no additional characters, yet the collec-
tion records a second station towards mapping the distribution
of the species.
PSITTACANTHUS AURicuLATus, Oliver, ace. to Eichl., in Mart. Fl.
Bras. v. II, 25 (1866). Loranthus auriculatus D. Oliver in
Kjoeb. Vidensk. Meddel. 1864, p. 174.
250 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
To this well marked species are referred specimens collected at
Alturas de Ejutla, Oaxaca, Mexico, altitude 1,300 m., 13 Decem-
ber, 1907, C. Conzatti, no. 1641 (hb. Field Mus.).
PHORADENDRON MUCRONATUM Krug & Urban in Engl. Bot. Jahrb.
xxiv. 34 (1897). P. -ftavescens Millsp. Field Col. Mus. Bot. Ser.
i. 294 (1896) in part, not Nutt.
MEXICO. State of Yucatan: near Izamal, Dr. Geo. F. Gau-
nter, no. 561 in part (hb. Field Mus.). Dr. Gaumer's specimens
correspond in every essential detail with the descriptions of this
species, and with material in the herbarium of the Field Museum
from the West Indies and from South America. This species
seems not to have been reported hitherto from Mexico or Central
America.
PHORADENDRON QUADRANGULARE Krug & Urban in Engl. Bot.
Jahrb. xxiv. 35 (1897) & Urban, Symb. Antil. iv. 207 (1905).
Fruiting specimens of this species were collected near the
coast north of the City of Vera Cruz, Mexico, 24 January, 1906,
/. M. Greenman, no. 120 (hb. Field Mus.).
Phoradendron vernicosum Greenman, sp. nov.
Glabrous throughout: younger parts more or less vernicose:
stems and branches terete; ultimate branchlets compressed
at the nodes: leaves lanceolate-oblong to ovate-elliptic, often
slightly oblique or subfalcate, 2 to 7 cm. long, i to 2.7 cm. broad,
obtuse or rounded at the apex, entire, narrowed below to a
subpetiolate base, 3~5-nerved: spikes sessile or essentially so,
i to 3 (rarely 5) in the leaf-axils, i to 2 cm. long; segments 2 to
5 (usually 4), 5 mm. or less in length, 6- 12 -flowered in the
staminate spike, 2-flowered in the pistillate spike; perianth
3-merous: berry ovate-oblong, about 5 mm. long, not con-
tracted below the calyx-limb, more or less glaucous; endocarp
distinct, ovate-oblong, 4 mm. long, 2 mm. wide, abruptly
acuminate. P. ftavescens, Millsp. Field Col. Mus. Bot. Ser.
i. 294 (1896) in part, not Nutt. MEXICO. State of Yucatan:
Izamal, 22 February, 1906, /. M. Greenman, no. 440 (hb. Field
Mus.), type; Silam, June, 1895, Dr. Geo. F. Gaumer, no. 876
(hb. Field Mus.); Chichankanab, Dr. Geo. F. Gaunter, nos. 1850,
201 1 (hb. Field Mus.).
The vernicose character of the young stem and leaves, the
short axillary inflorescences, the two-flowered segments of the
fertile spike and the distinctly acuminate endocarp well char-
acterize this species. The nearest affinity of P. vernicosum is
with P. Wattii Krug & Urban, from which it differs in having
relatively shorter and broader leaves, the fruit not constricted
below the limb of the calyx, and a smaller and distinctly acu-
minate instead of acute endocarp.
MILLSPAUGHIA ANTiGONOiDES Rob. in Engl. Bot. Jahrb. xxxvi. Beibl.
80: 14 (1905).
In addition to the specimens cited in the original publication
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 251
of this very interesting genus the following collections in the
herbarium of the Field Museum represent further the above
species. MEXICO. State of Yucatan: Merida, April, 1865,
Dr. A. Schott, no. 217; Colonia San Cosme, 20 February, 1906,
J. M. Greenman, no. 348; Izamal, collection of 1888, Dr. Geo. F.
Gaumer, without number; Izamal, Dr. Geo. F. Gaunter, nos.
3001, 3002, 3004; Puerto Morelos, 12 to 31 March, 1901, E. A.
Goldman, no. 626 (hb. U. S. Nat. Mus. ; fragment in hb. Field
Mus.).
Guatteria Gaumeri Greenman, sp. nov. Tree, 10 to 15 m. high:
stem and branches covered with a gray bark ; ultimate branchlets
glabrous or sparingly strigulose-puberulent : leaves alternate,
petiolate, lanceolate to elliptic-lanceolate, 5 to 15 cm. long, 2
to 2.5 cm. broad, usually short-acuminate and obtuse, rarely
retuse at the apex, entire, glabrous on both surfaces or in the
very early stages slightly pubescent with a few scattered appressed
hairs, soon glabrate and rather strongly reticulate-nerved;
petioles stoutish, 3 to 10 mm. long, canaliculate, often turning
blackish in the dried state: inflorescence terminal or lateral;
peduncles thickish, i to 3 cm. in length, jointed, sparingly pubes-
cent with appressed tawny hairs, bracteate at the base and
usually bearing a single ovate acute or acutish ciliate fer-
rugineous-pubescent bract below the middle: sepals subrotund,
3 to 5 mm. high, usually broader than long, ciliate and sparingly
pubescent to glabrous: petals large, oblong-ovate to some-
what obovate, 2 to 4.3 cm. long, 1.2 to 3 cm. broad, thick and
leathery: berries numerous, elliptic-obovoid, about i cm. long,
7 to 8 mm. in diameter, minutely verrucose, glabrous: stipes
slender, 1.5 cm. or less in length; torus somewhat depressed-
globose. MEXICO. State of Yucatan: vicinity of Izamal,
specimens communicated February, May, June, and July, 1906,
Dr. Geo. F. Gaumer (hb. Field Mus., catalogue nos. 189976-189-
978, 189160, 189161). In general appearance the species here
proposed resembles G. dolichopoda Donn. Sm., but it differs in
the less acuminate and blunt leaves, character of the pubescence,
subrotund sepals, larger petals, shorter peduncles and stipes.
G. Gaumeri is rich throughout all its parts in oil-glands, and
when crushed it produces a pleasant aromatic odor. Dr. Gaumer
in whose honor the species is named states that the plant is known
about Izamal under the name of "Elemuy," and that from it
is obtained one of the most valuable medicines used in Yucatan.
TRISTICHA HYPNOIDES Spreng. Syst. Veg. iv. pt. 2, 10 (1827); DC.
Prodr. xvii. 44 (1873) ; Hemsl. Biol. Cent. -Am. Bot. iii. 39 (1882).
Specimens well representing this species were found growing
on stones under water near Cordoba, State of Vera Cruz, Mexico,
25 January, 1906, /. M. Greenman, no. 124 (hb. Field Mus.).
This interesting species, known from Cuba, from Guatemala
to Brazil, from tropical and south Africa and Madagascar, seems
not to have been recorded hitherto from Mexico. Specimens
252 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
collected at Cordoba by Dr. Asa Gray and referred by him to the
above species, although no published record of them has been
found by the writer, bear somewhat larger fruit than my num-
ber 124, but differ in no other apparent regard.
Caesalpinia yucatanensis Greenman, sp. nov.
Shrub or small tree: stem covered with a light gray bark,
dotted with numerous lenticels, glabrous; cortex defoliating
in thin scarious layers; ultimate branchlets puberulent; leaves
alternate, bipinnate, petiolate, unarmed; petioles 2 to 6 cm.
long; pinnae 2 to 3 pairs; leaflets 2 to 4 pairs, oblong-elliptic,
1.5 to 4 cm. long, 0.7 to 2.5 cm. broad, obtuse to rounded at
both ends or slightly retuse at the apex, entire, glabrous on both
surfaces or somewhat pubescent in the early stages and glabrate;
midrib slightly sunken from the upper surface and, as well as the
lateral nerves, somewhat prominent beneath; petiolules i to 1.5
mm. long: inflorescence usually in terminal panicles, 0.5 to 1.5
dm. in length, occasionally terminating the lateral branches
in simple racemes, finely pubescent; pedicels i to 2 cm. long,
jointed above the middle, pubescent: calyx about i cm. long,
5 -parted; segments oblong-rotund, imbricated, densely soft-
pubescent on the outer surface: petals oblong to oblong-obovate,
about 1.5 cm. long, 8 to 10 mm. broad, narrowed at the base into
a villous-pubescent claw, chocolate-brown or dark red in color
and margined with pale yellow, covered externally in the lower
half with sessile or short-stipitate glands; the uppermost petal
producing a short fold on the inside near the base: stamens
barely exserted; filaments pubescent with more or less matted
hairs: ovary and lower part of the style densely pubescent:
mature fruit sessile, oblong, slightly oblique, 6 to 12 cm. long,
2 to 2.5 cm. broad, short-pubescent and closely beset with stipi-
tate tack-shaped glands; seeds suborbicular, flat, about i cm.
in diameter, smooth. Caesalpinia exostemma Millsp. Field
Col. Mus. Bot. Ser. i. 21 (1895), not Moc. & Sesse ex DC.
MEXICO. State of Yucatan: vicinity of Izamal, collection of
1895, Dr. Geo. F. Gaunter, no. 371 (hb. Field Mus.), type; near
Izamal, 13 January, 1895, Dr. C. F. Millspaugh, no. 75 (hb. Field
Mus.); Izamal, 22 February, 1906, /. M. Greenman, no. 417
(hb. Field Mus.); San Anselmo, Dr. Geo. F. Gaumer, no. 1715
(hb. Field Mus.); near Merida, Dr. A. Schott, without number
(hb. Field Mus.) ; on old hennequin plantations near Merida,
February, 1903, C. & E. Seler, no. 3844 (hb. Field Mus.) ; Colonia
San Cosme, 20 February, 1906, /. M. Greenman, no. 349 (hb.
Field Mus.); Itzimna, 19 February, 1906, /. M. Greenman, no.
335 (hb. Field Mus.); near Progresso, 5 March, 1899, Dr. C. F.
Millspaugh, no. 1660 (hb. Field Mus.); without definite locality,
coll. of 1896, Sr. Porfirio Valdez, no. 7 in part (hb. Field Mus.).
State of Campeche: without locality, Dr. Henry Perrine (hb.
Gray, and hb. Torrey).
This species is related to C. exostemma Moc. & Sesse ex DC.
DEC. 1907 SPERMATOPHYTES FROM MEXICO GREENMAN. 253
with which it has been confused, but from which it differs in
having a pubescent inflorescence, more oblong and copiously
glandular petals, in having also the inner or upper petal less
conspicuously clawed and bearing a scale-like fold on the inner
or upper side near the base, and finally by the shorter barely
exserted stamens.
Phaseolus (Drepanocarpos) polyanthus Greenman, sp. nov.
Stem robust, angulate-striate, sparingly pubescent with
ascending, spreading or even reflexed hairs: leaves petiolate,
trif oliolate ; petioles 4 to 10 cm. long, slightly pubescent; stipules
triangular-ovate, 5 to 6 mm. long, acute ; leaflets rhombic-ovate,
or the lateral obliquely ovate, 4 to 10 cm. long, 3 to 9 cm. broad,
mucronate-acute, entire, subtruncate to obtuse at the base,
dark green and substrigose-hirsute above, slightly paler and
hirtellous-puberulent beneath, more or less glabrate, 3-nerved
and bearing tufts of white villous hairs in the axils of the veins
on the under side; petiolules stoutish, about 5 mm. long, densely
tawny-hirsute above; stipels subfalcate-linear, 3 to 4 mm. long,
glabrous or essentially so: inflorescence in elongated axillary
racemes, 2.5 dm. or less in length; rhachis pubescent; bracts
lance-attenuate, about 7 mm. long, pubescent; pedicels becom-
ing 12 mm. in length, glabrous or nearly so, and as well as the
bracts persistent; bracteoles subtending the calyx, linear-
lanceolate to lance-oblong, 6 to 7 mm. long, i to 1.5 mm. broad,
acute, 3-5-nerved, ciliolate: calyx about 5 mm. high, subbila-
biate, or 2-lobed; tube 3 mm. long; upper lobe emarginate;
the lower lobe 3-toothed with the midddle tooth ovate, acute,
2 mm. long, the lateral teeth shorter and obtuse: vexillum
somewhat oblong-obovate, 12 mm. long, nearly or quite as broad,
short-unguiculate with a broad claw; disk barely exceeding
i mm. in length, crenate-margined : ovules commonly 6: mature
fruit not seen. MEXICO. State of Vera Cruz: on railroad
banks near Jalapa, 10 September, 1906, C. R. Barnes, C. J. Cham-
berlain & W. J. G. Land, no. 20 (hb. Field Mus., and hb. Uni-
versity of Chicago). The species is rather striking on account
of the large membranous leaflets, many-flowered inflorescences,
persistent pedicels, and the narrow bracts and bracteoles. It
suggests P. multiflorus Willd. and P. pedicellatus Benth, but is
quite distinct from either of them.
ASTROCASIA Rob. & Millsp. in Engl. Bot. Jahrb. xxxvi. Beibl. 80:
19 (i9S)-
This genus was first described from staminate specimens only.
Fertile plants are now at hand, and additional generic char-
acters may be given as follows: Pistillate flowers solitary or
fascicled. Ovary 3-celled; cells 2-ovuled; stigma sessile,
3-lobed, fleshy. Disk cupular. Capsule septicidally dehiscent,
each carpel splitting vertically into equal halves ; exocarp readily
separating from the endocarp. Seeds ecarunculate.
254 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
A. PHYLLANTHOIDES Rob. & Millsp. 1. c. 2o. Phyllanthus nutans
Millsp. Field Col. Mus. Bot. Ser. i. 306 (1896), as to Gaunter,
nos. 475, 685, & in Engl. Bot. Jahrb. 1. c. 19, not Sw.
A dioecious shrub i to 2 m. high: fully developed leaves 4 to
13 cm. long, two-thirds as broad: pistillate flowers few; pedi-
cels rather slender, 2.5 to 4.5 cm. long, gradually enlarged
towards the base of the calyx; sepals broadly ovate, ovate-ob-
long or slightly obovate, 2 to 3 mm. long, two-thirds as broad,
reflexed; petals 5, erect or nearly so, oblong-lanceolate, 4 to 5
mm. long, 1.5 mm. broad, crenate-undulate ; disk 5-lobed:
mature capsule about 8 mm. long, nearly or quite as broad,
smooth and glabrous; seeds two in each cell, ovoid, 4 to 5 mm.
long, smooth, 'brownish. MEXICO. State of Yucatan: Vicin-
ity of Izamal, Dr. Geo. F. Gaunter, no. 475 (hb. Field Mus.);
Temax, Dr. Geo. F. Gaunter, no. 685 (hb. Field Mus.); Calotmul,
Dr. Geo. F. Gaunter, no. 1795 (hb. Field Mus.); Chichankanab,
Dr. Geo. F. Gaunter, nos. 1261, 1794 (hb. Field Mus.); Mayapan,
C. & E. Seler, no. 3874 (hb. Field Mus.); Itzimna, near Merida,
C. & E. Seler, no. 3943, type, (hb. Field Mus.); near Izamal,
21 February, 1906, /. M. Greenntan, no. 392 (hb. Field Mus.).
State of Campeche: Apazote, near Yohaltun, E. A. Goldman,
no. 491 (hb. U. S. Nat. Mus., and hb. Field Mus.). Flowering
and fruiting specimens of this species were collected by the
writer in February of 1906 near Izamal, Yucatan, where the
plant is quite abundant, and where it is one of the most attract-
ive shrubs in the "scrub" formation.
Acalypha Seleriana Greenman, sp. nov.
Shrub, i to 2.5 m. high, branched; stem and branches covered
with a reddish-brown or grayish bark and dotted with numerous
lenticels; the younger branchlets densely pubescent with short
horizontally spreading tawny hairs: leaves petiolate, ovate
to oblong-lanceolate, 2.5 to 5 cm. long, i to 3 cm. broad, acute
or obtuse, dentate or crenate-dentate, obtuse to rounded at
the base, 3 -nerved, thin and membranous, at first pubescent
on both surfaces especially on the veins beneath, later more or
less glabrate; petioles 3 to 18 mm. long, densely pubescent;
stipules lance-linear, 1.5 to 2 mm. long; caducous: inflorescence
chiefly axillary: spikes of fertile flowers inconspicuous, slender,
few-flowered: pistillate flowers small, sessile, solitary in the
axils of minute 3 -parted bracts about 0.5 mm. high: calyx i
mm. long, 5-parted into narrowly lanceolate acute divisions,
sparingly pubescent: ovary muricate-hispid ; style 3-parted,
or occasionally 2 -parted; divisions thickened and roughish
at the base, branching into about 9 laciniate-fimbriate divisions :
spikes of staminate flowers numerous, uniaxillary, slender, sessile
or short-pedunculate, i to 8 cm. long, 2 to 3 mm. thick, erect,
spreading, or occasionally more or less reflexed : mature capsules
and seeds not seen. A. mollis Millsp. in Field Col. Mus. Bot.
Ser. i. 302 (1896), & in Engl. Bot. Jahrb. xxxvi. Beibl. 80: 19
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 255
(1905), not HBK. MEXICO. State of Yucatan : in forests about
Xkombec, 5 April, 1903, C. & E. Seler, no. 4028 (hb. Field Mus.),
type; in forests near Xcolumkin, 5 April, 1903, C. & E. Seler,
no. 4040 (hb. Field Mus.) ; in forests about Izamal, March-
April, 1895, Dr. Geo.F. Gaunter, no. 477 (hb. Field Mus., and hb.
Gray), and in the same locality coll. of 1888, Dr. Geo. F. Gaunter,
specimens without number (hb. Field Mus.); vicinity of Izamal,
21 February, 1906,7. M. Greenman, no. 390 (hb. Field Mus.).
Acalypha mollis HBK. to which species some of the specimens
above cited have been hitherto referred is described as an herba-
ceous plant with distinctly pedunculate spikes, and with 2-3-
flowered reniform-ovate-n-i5-dentate bracts. On these charac-
ters alone A. Seleriana may be readily separated. The species
here proposed seems to be quite unique on account of the shrubby
habit, the numerous sender sessile or subsessile spikes of stami-
nate flowers, and the pistillate flowers solitary in the axils of
exceedingly minute 3 -parted bracts.
Dalechampia Schottii Greenman, sp. nov.
Stems twining, covered below with a grayish bark; branches
terete, striate, pubescent with spreading or reflexed hairs:
leaves petiolate, simple and undivided, subtrinervate from the
base, ovate or ovate lanceolate, 2-7.5 cm - l n g> I -5~4 cm - broad,
rounded to acuminate at the apex, mucronate-acute, subentire
or somewhat dentate in the lower half, obtuse to subcordate at
the base, usually bearing on the upper side at the junction of
petiole and blade two subulate appendages, pubescent on both
surfaces, glabrate above; petioles 0.5-2.5 cm. long, pubescent;
stipules narrowly lanceolate to almost subulate, 2-6 mm. long:
peduncles 1-3 cm. long, striate, pubescent; the petaloid involucral
bracts small, ovate, 6-12 mm. long, 3-8 mm. broad, acuminate
or merely acute at the apex, obtuse at the base, 3-nerved, spar-
ingly denticulate, externally pubescent, ciliate: calyx of the
staminate flowers 6-parted; divisions lanceolate, 2-2.5 mm.
long, acute, entire, glabrous: calyx of pistillate flowers 7-12-
parted; divisions linear-lanceolate, about 5 mm. long during
anthesis, pectinate and hirsute-pubescent, persistent and becom-
ing i cm. in length at maturity; ovary 3-celled, puberulent;
style cylindrical, stoutish; stigma subtrilobed, not dilated:
capsule depressed-globose, inconspicuously puberulent, reddish-
brown or blackish in the dried state; seeds subglobose, about
3.5 mm. long, rugulose. MEXICO. State of Yucatan: Merida,
3 August, 1865, Dr. A. Schott, nos. 534, 956 (hb. Field Mus.),
type; Chichankanab, Dr. Geo. F. Gaunter, nos. 1430 in part,
1463 (hb. Field Mus.); Merida, February, 1903, C. & E. Seler,
no. 3836 (hb. Field Mus.) distributed as "Dalechampia den-
ticulata Griseb.?;" Izamal, 22 February, 1906, /. M. Greenman,
no. 422 (hb. Field Mus.).
Var. trifoliolata Greenman, var. nov.
Leaves simple or trifoliolate ; divisions lanceolate, entire or
256 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
somewhat irregularly dentate: other characters as in the species.
MEXICO. State of Yucatan : Chichankanab, Dr. Geo. F. Gaumer,
nos. 1512, 1430 in part (hb. Field Mus.).
Some of the specimens above cited have been hitherto doubt-
fully referred to Dalechampia denticulata Wright of the West
Indies. From this species, however, D. Schottii differs in having
uniformly smaller leaves, shorter petioles, smaller floral bracts and
a nondilated stigma. Moreover, in D. denticulata the leaves
and floral bracts are distinctly cordate, while in D. Schottii
the leaves are from obtuse to subcordate at the base and the
floral bracts are narrowed below to an obtuse base. The variety
trifoliolata suggests D. triphylla, var. mexicana Mull. Arg., but
the latter has petioles very much longer in proportion to the
length of the leaf-blade.
Jatropha Gaumeri Greenman sp. nov.
Tree, 5 to 10 m. high, much-branched: trunk 2 to 5 dm. in
diameter; branches and branchlets thick and somewhat fleshy:
leaves alternate, petiolate, palmately y-nerved, broadly ovate,
5 to 1 8 cm. long, 4.5 to 15 cm. broad, abruptly caudate-acumi-
nate, acute, entire, or occasionally subdenticulate in the lower
portion, rarely sublobate, deeply cordate to subtruncate at the
base, membranous, glabrous above, tawny-pubescent along the
veins at the base of the blade beneath, otherwise glabrous;
petioles 2.5 to 13 cm. long, glabrous except near the blade:
inflorescence in terminal or axillary short-pedunculate com-
pound cymes, 2.5 cm. or less in length, glabrous or with a few
tawny hairs in the axils of the deltoid or triangular-ovate acute
glabrous bracts ; peduncles i cm. or less in length : flowers sessile,
monoecious, whitish or cream-colored: calyx gamosepalous,
2 to 3 mm. high, glabrous, 5-lobed, persistent; lobes erect in
anthesis, subrotund, slightly unequal, entire: corolla 6 to 7 mm.
long, tubular for about two-thirds its length, externally gla-
brous, densely ferruginous-pubescent towards the base within;
lobes 5, erect or slightly spreading, oblong-ovate, rounded at
the apex: glands usually 5, occasionally 3: stamens 8, included;
the outer series or cycle consisting of 5 distinct stamens about
equalling the more or less coalescent filaments of the 3 inner
stamens; anthers oblong, acutish: capsule oblong-globose,
subtriangular in cross-section, 15 to 18 mm. long, nearly or
quite as broad, glabrous, septicidally dehiscent; the carpels
later splitting along the median line: seeds carunculate, oblong,
about 13 mm. long, n mm. broad, slightly roughened. Ficus
Jaliscana Millsp. Field Col. Mus. Bot. Ser. i. 293 (1896), not
Watson. Jacaratia Mexicana Millsp. 1. c. 35 (1895), not DC.
MEXICO. State of Yucatan: near Izamal, collection of 1895,
Dr. Geo. F. Gaumer, no. 365 (hb. Field Mus.); San Anselmo,
Dr. Geo. F. Gaumer, no. 1705 (hb. Field Mus.); near Izamal,
15 January, 1895, Dr. Chas. F. Millspaugh, no. 96 (hb. Field
Mus.); vicinity of Izamal, 22 February, 1906, /. M. Greenman,
no. 478 (hb. Field Mus.).
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 257
In leaf-outline and in the pubescence of the leaf the species
here proposed suggests /. yucatanensis Briquet in Ann. Conserv.
& Jard. Bot. Geneve, iv. 230 (1900), but it differs in having
larger leaves which are abruptly caudate-acuminate and termi-
nated by a very slender acumen, shorter peduncles, essentially
glabrous inflorescence, deltoid bracts, sessile flowers, and 8
instead of 10 stamens.
The plant grows as a rather profusely branching tree with
thick and somewhat fleshy branches and twigs. The almost
leafless condition of the tree and its light gray appearance render
it a conspicuous feature of the "scrub" and woodlands about
the City of Izamal. It passes under the Mayan name of " Pom-
olche;" and its stems are said to be used by the native people in
making the so-called "Chul" or whistles.
Gouania Conzattii Greenman, sp. nov.
Stem terete or slightly angulate above, sparingly pubescent;
leaves alternate, petiolate, ovate or subrotund-ovate, rounded to
short-acuminate and submucronate-acute at the apex, crenate-
dentate, shallowly cordate at the base, dark green and hirsute-
pubescent above in the younger stages, more or less glabrate,
subtomentose beneath ; midrib and veins prominent on the under
side of the leaf; petioles i cm. or less in length: inflorescence
terminating the stem and upper branches in spicate racemes
together forming a more or less leafy panicle: flowers sessile,
or on very short pedicels: calyx-limb 5-lobed; lobes triangular-
ovate, acute, entire, externally as well as the entire inflorescence
tawny-pubescent; disk distinctly 5-lobed and the lobes about
one-half as long as the lobes of the calyx, 2-dentate and more
or less persistent: petals strongly cucullate, i mm. long: mature
capsules triangular, 6 to 7 mm. high, including the strongly
developed wings 7 to 9 mm. in diameter, glabrous or nearly so;
seeds oval, 3 to 4 mm. long, smooth and shining, convex on the
outer surface, 2-faced and more or less 2-scalloped on the inner
surface. MEXICO. State of Oaxaca: Cerro San Felipe, altitude
1,700 m., 15 September, 1906, C. Conzatti, no. 1567 (hb. Field
Mus.). Habitally and in leaf-outline G. Conzattii resembles G.
tomentosa Jacq., but differs in having a sparingly pubescent
stem, larger flowers, and also in bearing capsules which are
nearly twice as long in the vertical axis, and producing seeds
which are fully twice larger than G. tomentosa.
MACROSCEPIS OBOVATA HBK. Nov. Gen. & Sp. iii. 201, t. 233 (1818);
DC. Prodr. viii. 599 (1844); Hemsl. Biol. Cent. -Am. Bot. ii.
320 (1881).
Specimens agreeing in all details with the original description
and illustration of this species were collected at Izamal, Yucatan,
by Dr. Geo. F. Gaunter, no. 1198 (hb. Field Mus.), and again
at Chichankanab by the same collector, no. 2239 (hb. Field Mus.).
This species has not been recorded hitherto from Yucatan.
258 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
Ipomoea Conzattii Greenman, sp. nov.
Stem ligneous, covered with a gray bark and dotted with
numerous lenticels; ultimate branches pubescent; leaves not
seen: inflorescence in axillary sessile, or short-pedunculate
i -several-flowered (i-i6).more or less nodding sericeous-hirsute
cymes; bracts triangular-acuminate, acute, caducous; pedicels
i to 2 cm. long, upwardly thickened, striate, pubescent: calyx
about 7 mm. high; sepals ovate-rotund to broadly ovate, 5 to
7 mm. long, nearly or quite as broad, rounded or slightly emar-
ginate and submucronate at the apex, the outermost densely
sericeous-hirsute on the outer surface, the inner slightly pubes-
cent to glabrous externally, scarious-margined and often tinged
with purple: corolla tubular-campanulate, 3.5 to 4.5 cm. long,
externally glabrous; tube subcylindrical 2.5 to 3 cm. long, more
or less abruptly expanded into the 5-lobed limb, purple or some-
what magenta-colored in the dried state: stamens included or
barely exserted; filaments bearing a tuft of coarse hairs at their
insertion near the base of the corolla : style more or less persistent :
capsule subglobose, about i cm. in diameter, smooth and gla-
brous; seeds 2 in each cell, oblong-ovate, 9 mm. long, bearing
from its apex a reflexed coma somewhat exceeding the body of
the seed. MEXICO. State of Oaxaca: Almoloyas, altitude
800 m., 25 December, 1906, C. Conzatti, no. 1666 (hb. Field Mus.).
It is with some hesitation that the writer describes a leafless
plant as new to science, but the present one is so distinctive
in its ligneous stem, inflorescence and floral characters that it
seems best to present the above characterization. The species
is named in honor of the distinguished botanist, Professor
Cassiano Conzatti, Director of the Normal School in the City of
Oaxaca, Mexico.
IPOMOEA TENTACULIFERA Greenm. Proc. Am. Acad. xxxiii. 482 (1898).
Specimens collected on the Cerro San Felipe, Oaxaca, Mexico,
altitude 1,700 m., 12 August, 1906, C. Conzatti, no. 1618 (hb.
Field Mus.), match perfectly the original material secured by
Mr. C. G. Pringle in Tomellin Canon in 1897. Professor Conzatti's
specimens in addition to perfect flowers show well developed
fruit. The capsules are spherical-ovate, nearly or quite 1.5
cm. high, fully i cm. in diameter, smooth and glabrous with a
single well developed seed in each cell.
Stachytarpheta purpurea Greenman, sp. nov.
Suffruticose : stems terete or slightly 4-angled, hirsute-pubes-
cent: leaves opposite, rhombic-ovate, 2 to 5 cm. long, i to 2.4
cm. broad, rounded or acute at the apex, crenate-serrate, rather
abruptly contracted below the' middle to an entire base, hirsute-
hispid and more or less rugose above, slightly paler and more
densely hirsute beneath: inflorescence terminating the stem
and branches in slender elongated spikes, 2 to 2.5 dm. or less
in length ; rhachis sparingly pubescent, 2 mm. or less in diameter;
floral bracts rather remote, ovate, abruptly acuminate, 4 to 5 mm.
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 259
long, 2 to 2.5 mm. broad, glabrous or nearly so, ciliate and, as
well as the entire inflorescence, more or less purplish: calyx
tubular, about 7 mm. long, minutely 4-toothed, glabrous except
along the ribs, posteriorly parted for about one-third its length,
or occasionally parted both posteriorly and anteriorly: corolla
trumpet-shaped, i to 1.5 cm. long, 10 to 13 mm. in diameter
when fully expanded; tube curved, glabrous without, hairy
within; limb 5-lobed; lobes broader than long: stamens
included: staminodia pubescent: style exserted: fruit oblong,
4 mm. long, glabrous. MEXICO. State of Vera Cruz: hillsides
near Chavarillo, 7 September, 1906, C. R. Barnes, C. J. Cham-
berlain & W. J. G. Land, no. 48 (hb. Field Mus., and hb. Uni-
versity of Chicago).
Citharexylum Altamiranum Greenman, sp. nov.
Stem and branches covered with a rough gray bark; ultimate
branchlets hexangular, brownish, short-hirsute pubescent: leaves
opposite, petiolate, ovate, ovate-oblong to subrotund, i to 4
cm. long, 0.5 to 2.5 cm. broad, obtuse to rounded or occasionally
emarginate at the apex, entire or not infrequently with one or
two teeth towards the apex, ciliate, hirsute-pubescent on both
surfaces, slightly paler beneath, usually bearing one to three
disc-like glands on the blade; petioles 3 to 15 mm. long, pubes-
cent: inflorescence terminating the branchlets in pubescent
few-flowered spicate racemes, i to 3 cm. in length; bracts
subulate, exceeding the short pedicels: flower not seen: calyx
persistent and in the fruiting stage becoming somewhat charta-
ceous, turbinate, 4 to 4.5 mm. high, 5-angulate-keeled at the base,
distinctly 5 -dentate with short erect acute teeth, slightly pubes-
cent on both inner and outer surfaces, ciliate about the orifice:
mature fruit oblong-elliptic, 7 to 8 mm. long in the dried state;
pyrenae elliptic, 6 to 7 mm. long, concavo-convex, smooth.
MEXICO. State of Queretaro: Hacienda del Ciervo, between
San Juan del Rio and Cadereyta, 20 August, 1905, /. N. Rose,
Jos. H. Painter & J. S. Rose, nos. 9666, 10,268 (hb. U. S. Nat.
Mus., and hb. Field Mus.); del Ciervo al cerro de la mesa, 20
August, 1905, Dr. F. Altamirano, no. 1566 (hb. U. S. Nat. Mus.;
fragment in hb. Field Mus.).
The plant here described suggests in general appearance
C. Berlandieri Rob., but differs in having smaller leaves, fewer-
flowered inflorescence and in its calyx characters. The species
is named in honor of the distinguished Mexican naturalist, Sr.
Dr. Fernando Altamirano, Director of the National Medical
Institute in the City of Mexico.
CITHAREXYLUM CINALOANUM Rob. in Bot. Gaz. xvi. 342 (1891).
To this species the following specimens are referred. MEXICO.
State of Sinaloa: near Rosario, on the road to Acaponeta, 27
July, 1897, Dr. J. N. Rose, no. 1858 (hb. U. S. Nat. Mus., and
hb. Field Mus.) ; between Rosario and Concepcion, 23 July,
1897, Dr. J. N. Rose, no. 3269 (hb. U. S. Nat. Mus., and hb.
260 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
Field Mus.). The affinity of C. cinaloanum is in all probability
with the obscure C. scabrum Moc. & Sesse, and it may eventually
prove to be conspecific. The leaf-margins on an individual
plant often vary from entire to conspicuously dentate in the
apical portion; and the inflorescence, here as in several other
species of the genus, may be either simple or compound.
Citharexylum Rosei Greenman, sp. nov.
Stem and branches covered with a grayish or reddish brown
cortex, terete or subtetragonal ; ultimate branchlets 4-angled,
cinereous-hirsute: leaves opposite, lanceolate or oblanceolate,
1 to 3 cm. long, i cm. or less broad, obtuse to rounded at the
apex, entire, narrowed below to a subpetiolate base, hirtellous-
pubescent on the upper surface, cinereous-tomentulose beneath:
inflorescence terminating the branchlets in few-flowered short
racemes; pedicels 3 to 4.5 mm. long: flowers not seen: calyx
persistent and in the fruiting stage 2 to 2.5 mm. high, shallow,
somewhat saucer-shaped, truncate, pubescent: mature fruit
oblong, 7 to 8 mm. long in the dried state; pyrenae elliptic, 6 to
7 mm. long, concavo-convex, strongly corrugated on the outer
or convex surface. MEXICO. State of Queretaro: between
Higuerillas and San Pablo near the latter station, 24 August,
1905, /. TV. Rose, Jos. H. Painter & /. S. Rose, no. 9827 (hb.
U. S. Nat. Mus., and hb. Field Mus.). This species resembles
C. Altamiranum to which it is closely related, but from which
it differs amply in having smaller leaves of different outline, a
more dense tomentum, and in its shorter, shallower and trun-
cate calyx, and corrugated pyrenae.
Vitex Gaumeri Greenman, sp. nov.
Tree, 10 to 15 m. in height: branches covered with a grayish
bark; ultimate branchlets tawny-pubescent: leaves opposite,
petiolate, palmately compound; petioles 3 to 9.5 cm. long, vel-
vety pubescent; leaflets 5 to 7, petiolulate, ovate to elliptic-
oblong, rarely tending to become obovate, 1.5 to n cm. long,
0.5 to 5. cm. broad, usually short-acuminate and acute or occa-
sionally rounded at the apex, entire, obtuse to subcordate at
the base, dark green and pubescent above, glabrate in age, pale
and densely tomentulose beneath; midrib and lateral veins
prominent on the under surface, but the anastomosing of the
ultimate veins not conspicuously reticulated; petiolules 0.5 to
2.7 cm. in length, velvety pubescent: the pedunculate panicu-
late cymes axillary, clustered at the ends of the branches, 2 dm.
or less in length: calyx small, 2.5 mm. long, subbilabiate, rather
acutely 5-toothed, externally pubescent: corolla irregular,
bilabiate, about 9 mm. long; tube erect, 5 mm. high, narrowly
funnel-form; the two posterior corolla-lobes small, subrotund,
2 mm. long and broad, reflexed; the lower lip or three anterior
lobes of the corolla spreading, 5 to 6 mm. long, the median
lobe slightly pulverulent on the upper surface near its base:
stamens 4, didynamous, exserted; filaments narrow, slightly
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 261
pubescent: style about equalling the two longer stamens: fruit
depressed-globose, 1.5 cm. in diameter in the dried state.
Vitex pyramidata Millsp. Field Col. Mus. Bot. Ser. i. 317 (1896),
not Rob. MEXICO. State of Yucatan: vicinity of Izamal,
March-April, 1895, Dr. Geo. F. Gaunter, no. 607, flowering speci-
men, (hb. Field Mus., and hb. Gray), type; near Izamal, coll.
of 1888, Dr. Geo. F. Gaunter, flowering specimen, without num-
ber (hb. Field Mus., and hb. Kew) ; vicinity of Merida, June,
1865, Dr. A. Schott, no. 582, fruiting specimen, (hb. Field Mus.).
The species here described is nearly related to V. pyramidata
Rob. with which it has been confused, but from which it differs
in being a tree instead of a shrub, in having longer petiolules
which, as well as the petioles themselves, are velvety pubes-
cent instead of pulverulent, in having a paler lower leaf-surface,
smaller flowers, in the absence of villous hairs at the base of the
anterior lip of the corolla on its upper surface, and finally in
having a more pubescent and somewhat more sharply dentate
calyx. Dr. George F. Gaumer in whose honor the above species
is named states that the plant here described is a tree growing
in the forests about Izamal, where it attains a height of about
15 meters. The flowers, moreover, are said by Dr. Gaumer to
be bright purple. The tree passes under the native Mayan
name of "Yaxnic."
SCUTELLARIA AUREA Rob. & Greenm. Am. Journ. Sci. 1. 163 (1895).
Specimens well representing this species were collected on the
Cerro San Antonio, Oaxaca, Mexico, altitude 1,800 m., 28 Octo-
ber, 1906, C. Conzatti, no. 1583 (hb. Field Mus.).
Var. Conzattii Greenman, var. nov.
Stem erect or ascending: leaves petiolate, ovate to ovate-
lanceolate, 2 to 7 cm. long, i to 3 cm. broad, acuminate, acute,
entire; petioles 1.5 cm. or less in length: other characters as in
the species. MEXICO. State of Oaxaca: Cerro San Antonio,
altitude 1,800 m., 28 October, 1906, C. Conzatti, no. 1584 (hb.
Field Mus.) . The variety differs from typical forms of the species
in having smaller and perfectly entire leaves.
Bacopa procumbens,* n. comb. Erinus procumbens Mill. Gard.
Diet. ed. 8, no. 6 (1768) & ed. 9, no. 13 (1797). Lindernia dian-
thera Sw. Prodr. Veg. Ind. Occ. 92 (1788) ; Mill. Gard. Diet. ed. 9,
no. 2 (1797); Willd. Sp. PL iii. 326 (1800). Herpestis chamce-
dryoides HBK. Nov. Gen. & Sp. ii. 369 (1817); Benth. in DC.
Prodr. x. 393 (1846) ; Gray, Syn. Fl. ii. part I. 280 (1878) ; Hemsl.
Biol. Cent. -Am. Bot. ii. 451 (1882). Microcarp&a americana
Spreng. Syst. i. 42 (1825). Monniera procumbens O. Kuntze,
Rev. Gen. PL ii. 463 (1891). Bacopa chamczdryoides Wettst.
*The strict use of the earlier specific name requires the restoration of
Miller's procumbens for this well known plant which has long passed under the
name of H erpestis ckamaedryoi des HBK.; and in accordance with the Inter-
national Rules of Botanical Nomenclature, in adopting Bacopa of Aublet for
this group of plants, it becomes necessary to transfer the two species following.
262 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
in Engl. & Prantl. Nat. Pflanzenf. iv. Ab 3 b . 76 (1895). Mon-
niera dianthera Millsp. Field Col. Mus. Bot. Ser. ii. 98 (1900),
as to binomial and synonomy, not as to specimen cited. Mecar-
donia procumbens Small, Fl. Southeastern U. S. 1065 (1903).
Var. Schottii, var. nov.
Habit and foliar characters as in the species : pedicels slender,
varying in length from somewhat shorter to twice the length
of the leaves : flowers 6 to 7 mm. long in anthesis ; the outer upper
sepal sharply denticulate. MEXICO. State of Yucatan : Merida,
29 August, 1865, Dr. A. Schott, no. 616 (hb. Field Mus.); vicin-
ity of Izamal, collection of 1888, Dr. Geo. F. Gaunter, specimen
without number (hb. Field Mus.); Izamal, collection of 1895,
Dr. Geo. F. Gaunter, no. 474 (hb. Field Mus., and hb. Gray); in
moist places near Izamal, 22 February, 1906, /. M. Greenman,
no. 464 (hb. Field Mus.) ; San Anselmo, Dr. Geo. F. Gaumer,
no. 1792 (hb. Field Mus.); Chichen Itza, 28 January to 10
February, 1901, E. A. Goldman, no. 555 (hb. U. S. Nat. Mus., and
hb. Field Mus.). Differs from typical representatives of the
species in the somewhat smaller flowers, slightly narrower parts
of the calyx, and in the rather striking denticulate character of
the outer upper sepal.
B. auriculata, n. comb. Herpestis auriculata, Rob. Proc. Am. Acad.
xxvi. 172 (1891).
B. decumbens, n. comb. Herpestis decumbens, Fernald. Proc. Am.
Acad. xxxiii. 91 (1897).
JUSTICIA FURCATA Jacq., var. terminalis, n. comb. Adhatoda furcata,
a terminalis Nees in DC. Prodr. xi. 398 (1847).
In Mr. Pringle's Plantae Mexicanae two different numbers,
namely no. 6498 and 11665, have been distributed under the
above name, but no authority assigned to the combination.
Definite mention of the variety terminalis under the genus
Justicia with complete reference to synonomy seems not to have
been made hitherto. To this variety may also be referred speci-
mens collected at El Parian, District of Etla, Oaxaca, Mexico,
altitude 1,400 m., 2 September, 1906, C. Conzatti, no. 1556 (hb.
Field Mus.).
JACOBINEA VIRGATA Hemsley. Biol. Cent. -Am. Bot. ii. 522 (1882).
Drejera virgata Oerstd. in Kjoeb. Vidensk. Meddel. 1854, p. 154;
Walp. Ann. v. 660.
Specimens collected at De Almoloyas Sta. Catarina, Oaxaca,
Mexico, altitude 1,000 m., 26 December, 1906, C. Conzatti, no.
1662 (hb. Field Mus.), agree well with the original description
of the above species.
Morinda yucatanensis Greenman, sp nov.
A more or less climbing shrub : stem and older branches covered
with a grayish bark; the younger branches and branchlets
yellowish and densely pubescent with short horizontally spread-
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 263
ing hairs: leaves petiolate, lanceolate to elliptic-lanceolate,
occasionally ovate, 5 to 12 cm. long, i to 5 cm. broad, acuminate,
acute, entire, gradually narrowed below to a 2 - lo-mm. long
petiole, pubescent on both surfaces more densely so beneath;
interpetiolar stipules i to 4 mm. high, abruptly apiculate-acumi-
nate or occasionally bidentate: inflorescence in spherical sessile
or subsessile axillary pubescent heads: flowers numerous, united
to the inconspicuous truncate calyx-limb: corolla tubular or
tubular- funnel-form, about 7 mm. long, rather deeply 5-lobed,
pubescent on the outer surface, glabrous in the lower portion of
the tube within, pubescent above: stamens and style of two
lengths, reciprocally exserted or included : fruit somewhat succu-
lent, spherical, about 1.5 cm. in diameter; pyrenae obliquely
oblong-obovate, 6 mm. long, 3 to 4 mm. broad, smooth or nearly
so. M. Roioc Millsp. Field Col. Mus. Bot. Ser. i. 321 (1896) & in
Engl. Bot. Jahrb. xxxvi. Beibl. 80: 28 (1905), not L. M. Royoc
Millsp. Field Col. Mus. Bot. Ser. 1. c. 392 (1898), not L. MEXICO.
State of Yucatan: Nojcacab, 21 November, 1865, Dr. A. Sckott,
no. 709 (hb. Field Mus.) ; in brush lands about Izamal, collection
of 1895, Dr. Geo. F. Gaumer, no. 362 (hb. Field Mus., and hb.
Gray), type; near Merida, Sr. Porfirio Valdez, no. 67 (hb. Field
Mus.); Chichankanab, Dr. Geo. F. Gaumer, no. 1700 (hb. Field
Mus.); Chichen Itza\ 28 January to 10 February, 1901, E. A.
Goldman, no. 547 (hb. U. S. Nat. Mus., and hb. Field Mus.) ; Titas,
17 March, 1903, C. & E. Seler, no. 3971 (hb. Field Mus.) ; in forests
near Xkombec, 5 April, 1903, C. & E. Seler, no 4032 (hb. Field
Mus.); near Izamal, 22 February, 1906, /. M. Greenman, nos.
421, 471 (hb. Field Mus.).
This species has been referred hitherto to M. Royoc L., but
it may be readily separated by the persistent pubescence over
the entire plant, the sessile or subsessile inflorescence and fruit
slightly shorter corolla, somewhat more complete union of the
flowers, and by the slightly longer and more oblique pyrenae.
The plant is known in Yucatan under the Mayan name of "Joyoc"
(Hoyoc).
CRUSEA VIOLACEA A. Brongn., ace. to Neumann, in Rev. Hortic.
Ser. II. iv. 368 (1846); v. 61, pi. 30, fig. 4 (1846).
The following specimens agree in every essential detail with
the descriptions and illustration of the above little-known
species, and for the present at least they seem best placed here.
MEXICO-. State of Vera Cruz: Teocelo, 8 May, 1901, E. A.
Goldman, no. 685 (hb. Field Mus., and hb. U. S. Nat. Mus.);
meadows, Orizaba, May, 1905, C. A. Pur pus, no. 447 (hb. Field
Mus.), distributed as " Spermacoce rubra Ch. & Schl."; along
the railroad near Cordoba, 25 January, 1906, /. M. Greenman,
nos. 188, 209 (hb. Field Mus.); railroad banks near Jalapa,
C. R. Barnes, C. J. Chamberlain & W. J. G. Land, no. 16 (hb.
Field Mus., and hb. University of Chicago).
Professor William Trelease has kindly furnished me with a copy
264 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
of the original descriptions and a photograph of the illustration
of the above species from the Revue Horticole.
I am indebted, moreover, to Dr. Casimir DeCandolle who
has kindly made for me a critical comparison of my no. 209 with
Crusea calocephala DC. in the Prodromus herbarium. From
Dr. DeCandolle's letter I quote the following: "I have com-
pared the Crusea specimens you sent me with that of C. caloce-
phala in the Prodromus herbarium. The case of that species
is unfortunately not quite clear as you will see by the following:
C. calocephala DC. is represented in the herbarium by Mocino's
plate n. 497 (an original plate, by the way, and not a tracing),
together with Dunant's specimen from Peru. Now it happens
that these two documents do not seem to belong to the same
species, for the plate (of which I sent you yesterday a good
tracing, in a separate roller) shows penninerved leaves, whilst
in Dunant's specimen the leaves have the same nervation as in
your specimen. Now on the other hand Dunant's specimen
differs from yours ist by the shape of its involucral leaves
which are more attenuate at the base; 2d by its calyx being
densely pubescent from its base upwards, whilst in your specimens
it is much less pubescent and mostly on its upper part only.
Consequently I consider your specimens as specifically distinct
from both Mocino's plate and Dunant's specimen.
Moreover I must also draw your attention to the fact that in
Mocino's plate, the corolla is coloured in red (which please
inscribe on the tracing when it reaches you) whilst your plant
seems to have blue flowers. As for C. rubra Cham. & Schlecht.
it is also certainly distinct from your plant, as it has much
shorter and strigose hairs and longer petioles."
Both C. calocephala DC. and C. rubra Cham. & Schlecht. are
somewhat confused, and very little is known about C. violacea
Brongn. In fact the entire genus Crusea, as well as Spermacoce,
needs a careful revision, which, however, can only be done
satisfactorily after a critical comparison of the ample material
of our recent collections with types existing only in European
herbaria.
LOBELIA REGALIS Fernald, Proc. Am. Acad. xxxvi. 503 (1901).
Excellent specimens of this species were collected at Ejutla,
State of Oaxaca, Mexico, altitude 1,300 m., 12 December, 1906,
C. Conzatti, no. 1638 (hb. Field Mus.). This collection records
another station for one of the most beautiful species of the genus,
and one which is well worthy of introducing into cultivation.
OAXACANIA MALVAEFOLIA Rob. & Greenm. Am. Journ. Sci. 1. 151
(1895).
This interesting monotype has been found at De Almoloyas
a Sta. Catarina, Oaxaca, Mexico, at an altitude of 1,000 m.,
26 December, 1906, C. Conzatti, no. 1654 (hb. Field Mus.). The
only other recorded locality for this species is that cited under
the original description, namely Tomellin Canon, Oaxaca,
where it was secured by Mr. C. G. Pringle in 1894.
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 265
Brickellia Kellermanii Greenman, sp. nov.
Stem erect, nearly i in. in height, lignescent at the base,
terete, tomentulose: lower leaves opposite, the upper alternate,
short-petiolate, oblong-lanceolate, 3 to 6 cm. long, i to 2 cm.
broad, obtuse or acute, crenate-dentate to entire, subhirtellous
above, paler and whitish-tomentulose beneath, rather strongly
reticulate-nerved; petioles about 0.5 cm. long: inflorescence
a subcorymbose rather leafy panicle : heads many, sessile or nearly
so, 12 to 14 mm. high, usually i2-flowered: involucre narrowly
campanulate or subcylindric ; bracts of the involucre about 7-
seriate, strongly unequal, imbricated, sparingly pubescent,
ciliate, striate, pale with greenish nerves, or purplish; the outer
bracts ovate-oblong, mucronate, strongly ciliate, the inner
lance-linear, acute: flowers somewhat exceeding the involucre:
pappus silvery white, 6 mm. long, about as long as the white or
purplish-tinged corolla: achenes 3 mm. long, pubescent.
GUATEMALA. Department of Baha Vera Paz: Sierra de las
Minas, altitude 1,158 m., 3 March, 1907, W. A. Kellerman, no.
6127 (hb. Field Mus.).
In leaf-outline B. Kellermanii resembles B. cylindrica and B.
Pringlei Gray, but it is amply distinct from both these species in
the nature of the inflorescence, tomentum, and in the technical
characters of the head.
EGLETES VISCOSA Less. Syn. Comp. 252 (1832). Cotula viscosa L.
Sp. PI. 892 (1753); Willd. Sp. PI. iii. 2167 (1800).
This species was based on material collected by Houston in
the region of Vera Cruz, Mexico. Specimens obtained at La
Purga, 31 kilometers southwest of the City of Vera Cruz, 27
January, 1906, /. M. Greenman, no. 257 (hb. Field Mus.), agree
well with the early although brief descriptions of the above
species, and are confidently referred to it. The species is of
rather wide distribution, occurring in southern Mexico, the
West Indies and, according to Hemsley, in South America;
it is, moreover, well characterized by its viscid-hirsute character
and somewhat lyrate or pinnately incised foliage, suggesting
vigorous forms of Senecio viscosus L.
In 1890 and again in 1892 Mr. C. G. Pringle collected on river
banks at Las Palmas, Mexico, an Egletes which was referred to E.
viscosa Less. The latter of the two collections made by Mr.
Pringle was distributed by clerical error as "Egletes viscida
Less." Mr. Pringle's specimens differ in several particulars
from Lessing's species. Moreover, they do not correspond to
any known species of the genus, hence they may be characterized
as follows:
Egletes Pringlei Greenman, sp. nov.
Annual: stem erect or essentially so, simple or branched
from near the base, somewhat geniculate above, viscid-hirsute:
leaves ovate to more or less obovate, 1.5 to 7.5 cm. long, one-
half to two-thirds as broad, thin or membranous, rather coarsely
266 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
and unequally sinuate-dentate, contracted below into a narrowly
winged petiole and subamplexicaul by a slightly auricular base:
inflorescence terminating the stem and branches in a paniculate
cyme: heads 4 to 5 mm. high, radiate: ray-flowers many-
seriate; corollas minute, 1.5 to 2 mm. long, less than 0.5 mm.
broad: disk-flowers numerous; corollas about i mm. long,
4-5-toothed; tube on the outer surface glandular-pilose : achenes
slightly compressed, produced above into a somewhat irregular
saucer-shaped subcartilaginous pappus. MEXICO. State of
San Luis Potosi: river banks, Las Palmas, 4 June, 1890, C. G.
Pringle, no. 3531 (hb. Field Mus.) ; and in the same locality, 18
June, 1892, C. G. Pringle, no. 4101 (hb. Field Mus.).
E. Pringlei is readily distinguished from E. viscosa Less, by
the ovate or obovate undivided leaves, the more numerous
heads, shorter hairs on stem and branches, and finally by the
smaller narrower rays, the more conspicuously flaring pappus,
and the less conical receptacle.
Erigeron pacayensis Greenman, sp. nov.
Suffruticose : stems erect or ascending, branched, i to 1.5
dm. high, pubescent with upwardly appressed or but slightly
spreading hairs: leaves narrowly linear-oblanceolate, i to 2
cm. long, i to 3 mm. broad, acute or obtusish, entire or bearing
a single tooth on one or both margins, gradually narrowed below
to a subpetiolate base, sparingly pubescent on either surface to
nearly glabrous: heads few, small, 5 to 6 mm. high, less than
1 cm. in diameter including the rays, terminating the stem
and branches on long slender appressed-pubescent peduncles
6 cm. or less in length: involucre subcampanulate ; bracts of
the involucre about 3 -seriate, unequal, lance-linear, acute,
appressed-pubescent, more or less purplish; the outer bracts
shorter: ray-flowers 3-seriate, many; rays narrow, white or
roseate : flowers of the disk numerous : pappus bristles slender,
2 to 2.5 mm. long: achenes pubescent. GUATEMALA. Depart-
ment of Amatitlan: crater of the Volcano of Pacaya, altitude
2,500 m., 6 January, 1907, W. A. Kellerman, no. 6m (hb. Field
Mus., and hb. Gray).
The species here proposed has its affinity with Erigeron Ervend-
bergii Gray, and *E. irazuensis Greenman. From the former
it differs in having smaller heads, appressed instead of spreading
hairs on the involucre, more unequal involucral bracts, etc.
Although similar in habit to E. irazuensis, yet it differs in several
particulars, namely in having narrower and less pubescent
leaves, also in the appressed pubescence on the stem and pedun-
cles, as well as in other details.
I am indebted to Mr. H. H. Bartlett of the Gray Herbarium
for a critical comparison of Professor Kellerman's plant with the
type of E. irazuensis.
* By oversight published as Erigeron irazuense.
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 267
Baccharis Kellermanii Greenman, sp. nov.
Stem terete, sulcate, hirtellous-puberulent with crisp sordid
hairs: leaves elliptic-lanceolate to oblanceolate, i to 3.5 cm.
long, 2 to 8 mm. broad, acute or obtuse, sharply dentate with
unequal and somewhat spreading teeth, narrowed at the base
into a short petiole, at first slightly puberulent above but soon
becoming glabrous, persistently hirtellous-pubescent beneath,
subtrinervate and rather prominently reticulate-veined: in-
florescence a terminal subcorymbose cyme: heads of the pistil-
late plant numerous, 6 to 7 mm. high: involucre subcampanu-
late; bracts of the involucre unequal, 4-5-seriate, linear-ob-long,
obtuse or rounded at the tip, more or less purplish on the
back and ciliate towards the apex, the outermost shortest:
flowers 18 to 24; pappus about 4 mm. long, slightly tawny,
exceeding the slender corolla- tube : mature achenes 1.5 mm.
long, glabrous: staminate plant not seen. GUATEMALA. De-
partment of Solala: Volcano of Atitlan, 16 February, 1906,
W. A. Kellerman, no. 5356 (hb. Field Mus.). In leaf-outline
Baccharis Kellermanii suggests B. thesioides HBK., but the
two species are amply distinct in habit, inflorescence, involucral
details, and in the character of the surface of stem and leaves.
Gnaphalium brachyphyllum Greenman, sp. nov.
An herbaceous perennial, lanate-tomentose throughout: stems
several, erect or nearly so, from a ligneous base, i to 2 dm. high,
slender, leafy: leaves linear-oblong or slightly oblanceolate,
0.5 to 1.5 cm. long, i to 4 mm. broad, obtuse, entire, slightly
repand-margined, sessile and semiamplexicaul, densely lanate-
tomentose on both surfaces : inflorescence terminating the stems
in rather dense cymes: heads 4 to 5 mm. high: involucre sub-
campanulate, tomentose at the base otherwise essentially
glabrous; bracts of the involucre 3-4-seriate, pale stramineous,
the outer ovate and about 3 mm. long, the inner linear-lanceolate,
3.5 to 4 mm. long, greenish at the base especially along the
median line: pistillate flowers numerous: perfect flowers 6 to 10:
pappus caducous: achenes glabrous, i mm. long, reddish.
GUATEMALA. Department of Quezaltenango : Cerro Quemada,
8 February, 1906, W. A. Kellerman, no. 5301 (hb. Field Mus.).
The numerous slender stems and short crowded leaves render
this species readily recognizable among all the other known
species of the genus.
MELAMPODIUM KUNTHIANUM DC. Prodr. v. 519 (1836); Hemsl.
Biol. Cent.- Am. Bot. ii. 146 (1881); Rob. Proc. Am. Acad.
xxxvi. 460 (1901).
To this little known species the writer refers specimens collected
on hillsides near Chavarillo, State of Vera Cruz, Mexico, 7 Sep-
tember, 1906, C. R. Barnes, C. J . Chamberlain & W. J. G. Land,
no. 8 (hb. Field Mus., and hb. University of Chicago). In all
essential characters these specimens agree with descriptions of
the above species. The leaves in our material, however, are
268 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
either linear or 3 -parted with linear divisions instead of lan-
ceolate-linear and entire, as indicated in the original character-
ization.
Gymnolomia scaberrima Greenman, n. comb. G. platylepis Gray,
Proc. Am. Acad. xix. 5 (1883) ; Rob. & Greenm. Proc. Bos. Soc.
Nat. Hist. xxix. 102 (1899), excluding plants of Pittier. G. decur-
rens Klatt, Leopoldina, xxiii. 90 (1887). Tithonia scaberrima
Benth. in Kjoeb. Vidensk. Meddel. 1852, p. 91 & Gen. PI. ii.
368; Walp. Ann. v. 223 (1858). T. platylepis Schz. Bip., ace.
to Benth. & Hook. f. Gen. PI. ii. 368 (1876). Mirasolia scaber-
3- rima Benth. & Hook. f. 1. c. 375; Hemsl. Biol. Cent. -Am. Bot.
ii. 1 68 (1881). Perymeniopsis perfoliata Schz. Bip., ace. to
Klatt., 1. c. In addition to the specimens cited under G. platy-
lepis Gray by Robinson and Greenman, 1. c. (excluding Pittier's
nos. 3136, 3735), the following collection may be recorded,
GUATEMALA. Department of Socatepdquez : Volcano Agua, alti-
tude 2440 m., 18 February, 1905, Prof. W. A. Kellerman, no.
5361 (hb. Field Mus.).
Following the International Rules of Botanical Nomencla-
ture, adopted by the International Botanical Congress held at
Vienna in 1905, it becomes necessary to form the above binomial
for this well known south Mexican and Central American species.
Wedelia rugosa Greenman, sp. nov.
An herbaceous perennial: stems erect or nearly so, 5 dm. or
more high, simple or branched, subterete, strigose-hispid with
upwardly appressed hairs: leaves opposite, short-petiolate,
oblong to oblong-lanceolate, 3 to 10 cm. long, 1.5 to 5 cm. broad,
acute or obtuse, entire or slightly crenate-dentate, rounded to
subcordate at the base, tuberculate-hispid and rugose above,
hirsute-hispid and strongly reticulate-nerved beneath; petioles
2 to 5 mm. long: inflorescence terminating the stem and branches
in few-headed cymes: heads radiate, 5 to 8 mm. high, including
the rays 10 to 16 mm. in diameter; peduncles 0.5 to 4 cm. long,
strigose-hispid: involucre campanulate; bracts of the involucre
biseriate, oblong to elliptic-ovate, 3 to 4 mm. long, i to 1.5 mm.
broad, obtuse or acute, externally appressed-pubescent : ray-
flowers 12 to 14, fertile; rays yellow, about 6 mm. long, 2.5 mm.
broad, 2-3-dentate; achenes 3-angled: disk-flowers numerous;
achenes at first laterally compressed, densely atomiferous-
glandular over the upper one-third, glabrous below, at maturity
oblong, 2.5 mm. long and subquadrangular in cross-section:
pappus of both ray- and disk-flowers a fimbriate crown of 2 (-3)
small scales at the angles of the achene with minute more or
less coalescent intermediate scales : pales of the receptacle rather
conspicuous, more or less uncinate- tipped especially in the
mature state. Wedelia reticulata Greenm. in Trans. Acad.
Sci. St. Louis, vii. 434 (1897), not DC. CUBA. Province of
Santa Clara: Abresus, 29 June, 1895, Robert Combs, no. 269
(hb. Field Mus.), type. Province of Havana: Managua, 19
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 269
July, 1904, Chas. F. Baker & Percy Wilson, no. 292 (hb. Field
Mus.); Managua, 23 September, 1904, Carl F. Baker, no.
1554 (hb. Field Mus.); Madruga, 23 November, 1904, A. H.
Curtiss, no. 528 (hb. Field Mus.). '
Var. tenuis Greenman, var. nov.
Similar to the species, but with smaller and more slender stems :
leaves 2.5 to 6 cm. long, 0.5 to 2.2 cm. broad. CUBA. Province
of Pinar del Rio: Herradura, 30 September, 1904, Carl F.
Baker, no. 2154 (hb. Field Mus.); Herradura, 24 August, 1905,
H. A. Van Hermann, no. 705 (hb. Field Mus.).
The species here described resembles W. reticulata DC. ; but
the oblong or oblong-lanceolate outline of the leaves with their
distinctly pinnate venation and entire or slightly crenate margin,
the shorter involucral bracts, and the peculiar uncinate-tipped
pales amply distinguish it from the De Candolle species.
Perymenium Goldmanii Greenman, sp. nov.
Stem terete or slightly tetragonal, striate, strigillose: leaves
opposite, petiolate, ovate-lanceolate, 3 to 8 cm. long, i to 4 cm.
broad, acuminate, acute, dentate to subentire, rounded to sub-
cordate at the base, scabrous above and becoming slightly
rugose in age, hirsutish-pubescent beneath, green on both sides,
3-nerved from near the base; petioles 3 to 10 mm. long; inflores-
cence terminating the stem and the horizontally spreading or
ascending lateral branches in rather dense cymose clusters:
heads numerous, small, 6 to 7 mm. high, about 3 mm. in diameter,
radiate; primary and secondary peduncles short, usually less
than 2 cm. in length, appressed-pubescent ; involucre subcampan-
ulate; bracts of the involucre unequal, 2-3-seriate, the outer
shorter, ovate to ovate-oblong, 3 to 4.5 mm. long, acute or ob-
tuse, externally strigillose, ciliate: ray-flowers commonly 5;
rays lemon-yellow: disk-flowers about 20: pappus of numer-
ous unequal setae: mature achenes laterally compressed or
somewhat 3 -angled, about 2.5 mm. long, ciliate or narrowly
winged, transversely rugose under a lens and slightly pubes-
cent on the surface especially in the upper half. MEXICO.
State of Campeche: Apazote, near Yohaltun, 27 to 29 December,
1900, E. A. Goldman, nos. 487, 502 (hb. U. S. Nat. Mus., and
hb. Field Mus.).
The affinity of the species here described is with P. micro-
cephalum Schz. Bip., and P. gracile Hemsl., but it differs from
the former by the larger leaves and from the latter in foliar and
achenial characters. The narrowly winged achenes of P. Gold-
manii suggest P. gymnolomoides DC., but that species has smaller
leaves, shorter petioles and fewer heads in the inflorescence.
Notoptera Gaumeri, n. comb. Salmea Gaunter i Greenm. in Field
Col. Mus. Bot. Ser. iii. 124 (1904).
The homogamous heads, the short accumbent involucre,
the characteristic ear-like wing on the posterior pappus-awn,
270 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
and the habit of the plant render it a congener of Professor
Urban's recently published genus Notoptera, Symb. Antil. ii.
465 (1901). In addition to Dr. Gaumer's no. 977, cited in the
original publication of the above species, the following is typical.
MEXICO. State of Campeche: Apazote, near Yohaltun, 26
December, 1900, E. A. Goldman, no. 483 (hb. U. S. Nat. Mus.,
and hb. Field Mus.).
ENCELIA ADENOPHORA Greenm. Proc. Am. Acad. xxxix. 109 (1903).
In addition to the stations recorded for this species under
the original description is the following: Hacienda Guadalupe,
State of Oaxaca, Mexico, altitude 1600 m., 7 October, 1906, C.
Conzatti, no. 1529 (hb. Field Mus.).
OTOPAPPUS VERBESINOIDES Benth. in Hook. Ic. PI. xii. 47, t. 1153
(1876); Hemsl. Biol. Cent.-Am. Bot. ii. 192 (1881).
A careful examination of a considerable number of specimens
representing this species shows a marked variation in the form
and size of the outer spreading subfoliaceous involucral bracts.
These vary from spatulate to lanceolate or linear and from
3 to 12 mm. in length. There is also some diversity in the size
of the rays. The following specimens are here referred.
MEXICO. State of Vera Cruz: Canton de Cordoba, altitude
1200 m., 25 December, 1897, C. Conzatti & V. Gonzdlez, no. 622
(hb. Gray). State of Chiapas: near Yajalon, 21 November,
1895, E. W. Nelson, no. 3409 (hb. Gray); Palenque, collection
of 1895, A- V- Armour, no. 7 (hb. Field Mus.). State of Cam-
peche: Apazote, near Yohaltun, 26 December, 1900, E. A.
Goldman, no. 482 (hb. U. S. Nat. Mus.; fragment in hb. Field
Mus.). GUATEMALA. Department of Alta Vera Paz: altitude
1310 m., June, 1882, H. von Tuerckheim, without number (hb.
Gray) ; Pansamala, altitude 1158 m., January, 1887, H. von
Tuerckheim, no. mo (hb. Gray). Depart, of Santa Rosa: Rio
Chiquito, altitude 700 m. September, 1893, Heyde & Lux, no.
6174 (hb. Gray, and hb. Field Mus.), an extreme form with
narrow elongated outer involucral bracts. COSTA RICA: Tur-
rialba, altitude 570 m., November, 1893, Ad. Tonduz, no. 8337
(hb. Gray); San Jose, altitude 635 m., December, 1898,
Ad. Tonduz, no. 12,739 (hb- Gray). Thus the species, as known at
the present time, has a geographical distribution ranging from
the State of Vera Cruz, Mexico, to Costa Rica.
Goldmania Greenman, gen. nov. of Composites (Coreopsidece}.
Heads heterogamous, radiate. Involucre campanulate ; bracts
3-4-seriate, free. Receptacle conical, paleaceous; pales thin,
membranous. Ray-flowers uniseriate, fertile: achenes com-
pressed dorsally. Disk-flowers regular; corolla-tube short,
gradually ampliated above into a 5 -toothed limb; achenes more
or less dorsally compressed. Anthers slightly sagittate at the
base, terminated by a short appendage. Style-branches elon-
gated, acute. Pappus of 2 to 4 short thick awns, or subcoroni-
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 271
form. Herbaceous perennials with alternate undivided leaves
and few-headed cymose inflorescence.
G. sarmentosa Greenman, sp. nov.
Stem prostrate or ascending, rooting at the lower nodes, terete,
glabrous below, slightly pubescent above: leaves sessile, or the
lowermost short-petiolate, ovate, 3 to 6 cm. long, two-thirds as
broad, acute or mucronate-acute, somewhat oblique or unsym-
metrical at the base, at first pubescent with a few scattered hairs
on both surfaces, soon glabrate: inflorescence terminating the
stem and branches in few-headed cymes: heads pedunculate,
including the rays about 1.5 cm. in diameter; peduncles slender,
i to 7 cm. long, naked or sparingly bracteolate, subappressed-
pubescent: involucre 6 to 8 mm. high; bracts glabrous, yellow-
ish with reddish-brown nerves, the outer shorter, ovate and
acute, the inner oblong, rounded at the apex and subscarious-
margined: ray-flowers 5 to 8; rays oblong or oblong-obovate,
about 6 mm. long, 3 to 4 mm. broad, 2-3-dentate, white or pale
yellow: disk- flowers about 20: pappus persistent as short thick
awns, sometimes subcoronate: mature achenes oblong, 2.5 to
3 mm. long, much thickened, glabrous but with a slightly rough-
ened surface, reddish-brown. MEXICO. State of Campeche:
Canasayal, 12.5 km. above Champotan river, 12 December,
1900, E. A. Goldman, no. 448 (hb. U. S. Nat. Mus., and hb. Field
Mus.). The plant here described in the appearance of the
involucre suggests some of the Galinsogece, particularly Calea and
Geissolepis, and certain of the Helenioidece, especially Jaumea.
The paleaceous receptacle, the dorsally appressed achenes, and
the character of the pappus place it, however, with the Coreop-
sidecB.
The genus is named in honor of its collector Mr. Edward A.
Goldman of the United States National Museum.
Bidens TJrbanii Greenman, sp. nov.
Perennial: stem ligneous, twining; branches terete, minutely
striate, glabrous, or puberulent in decussating lines: leaves
opposite, petiolate, triangular-ovate in general outline, exclusive
of the petiole 3 to 7 cm. long, 2 to 6 cm. broad, bi-tri-pinnatisect
with narrowly lanceolate acute submucronate-dentate divisions,
glabrous to minutely and sparingly hispidulous on the upper
surface, paler and glabrous or essentially so beneath; petioles
slender, 0.5 to 3.5 cm. long: inflorescence in terminal more or
less leafy paniculate cymes; the individual axillary pedunculate
cymes exceeding the leaves: heads radiate, during anthesis 8 to
10 mm. high, including the rays 2 to 2.5 cm. in diameter: invo-
lucre biseriate; the outer bracts of the involucre herbaceous,
lance-linear to narrowly spatulate, acute or obtusish, reflexed,
glabrous or slightly puberulent; inner involucral bracts thin,
dark brown or chocolate-colored with yellowish 'subscarious
margins: ray-flowers usually 5, sterile; rays narrowly oblong,
about i cm. long, obtuse or retuse, pale yellow, about 7-nerved
272 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
with dark brown or blackish nerves: disk-flowers 20 to 25:
mature achenes of the fertile flowers linear, 12 mm. or less in
length, erect or somewhat recurved, 2-4-awned with relatively
short retrorsely barbed persistent pappus-awns, striate, 2-4-
angled in cross-section, ciliate along the angles otherwise glabrous.
PORTO Rico. On slopes of Mt. Montoso, near Maricao, 23
November, 1884. Sintenis, no. 387 (hb. Field Mus. Catalogue
No. 79397), type. MEXICO. State of Campeche: Apazote, near
Yohaltun, 20 December, 1900, E. A. Goldman, no. 468 (hb. U. S.
Nat. Mus.; fragment and photograph in hb. Field Mus.).
This species has its affinity with a small natural group of
plants to which B. rubifolia HBK, B. tereticaulis and B. coreopsidis
DC. belong, but the smooth or lineate-puberulent stem and
branches, the pinnatisect leaves with their numerous narrow
divisions, the nature of the pubescence, and certain technical
characters of the head seem to the writer to warrant its specific
rank. B. Urbanii is apparently most closely allied to B. Coreop-
sidis, var. (?) incisa DC. v. 599 (1836) (Coreopsis incisa Ker-
Gawl. Bot. Reg. t. 7), but here again a careful comparison with
the original description and illustration shows several important
differences.
CALEA PRINGLEI Rob., var. rubida Greenman, var. nov.
Leaves short-petiolate, ovate, 1.5 to 3 cm. long, two-thirds
as broad, densely tomentose beneath; petioles 2 to 4 mm.
long, tawny-pubescent: involucral bracts elliptic-oblong, red-
dish-margined. MEXICO. State of Vera Cruz: on hillsides
near Chavarillo, 7 September, 1906, C. R. Barnes, C.J. Chamberlain
& W. J. G. Land, no. 4 (hb. Field Mus., and hb. University of
Chicago). Differs from the type of the species chiefly in the
short but distinctly petioled leaves, and by the slightly narrower
and reddish-margined involucral bracts.
Florestina Liebmannii Schz. Bip., in herb.
Stem erect or somewhat ascending from an indurated base, 3
to 5 dm. high, simple or sparingly branched, terete below,
striate-angled above, substrigillose and more or less closely
beset with stipitate glands: lower leaves opposite, simple,
petiolate, narrowly lanceolate-oblong, 2 to 3.5 cm. long, 4 to
12 mm. broad, obtuse or acute, entire or occasionally few-dentate,
obtuse at the base, substrigose-hispid on both surfaces, 3-nerved;
petioles 0.5 to 1.5 cm. long, hispid-pubescent intermixed with
stipitate-glandular hairs; the upper stem-leaves alternate and
gradually reduced to linear bracts: inflorescence terminating
the stem in a glandular-pubescent subcorymbose cyme: heads
homogamous, 8 to 10 mm. high: involucre subturbinate ; bracts of
the involucre 8, uniseriate or essentially so, elliptic-oblong to
obovate-oblong, 4 to 5 mm. long, 2 to 3 mm. broad, rounded at
the apex, scarious-margined and more or less erose-ciliate, ap-
pressed-puberulent on the outer surface : flowers 12 to 14: pappus
of 8 obovate or broadly spatulate hyaline-scarious pointless
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN.- 273
scales, these about 1.5 mm. long and narrowed below into a
thickened opaque base: corolla 4 mm. long, deeply 5-toothed,
externally somewhat pubescent: achenes narrowly obpyramidal,
3 to 4 mm. long, 4-5-angled, striate, pubescent. MEXICO.
State of Vera Cruz: Boca del Rio, Liebmann, no. 71 (hb. Copen-
hagen, fragment and good drawing in hb. Gray, photograph
in hb. Field Mus.) ; along the shore, north of the City of Vera
Cruz, 24 January, 1906, /. M. Greenman, no. 114 (hb. Field
Mus., and hb. Gray).
The general habit and simple leaves of this species suggest
a relationship with Palafoxia or Polypteris, but the technical
characters of the involucre, pappus, corolla and style-branches
place it rather with Florestina. The affinity of the plant, as
we'll as its specific rank, was first recognized by SchultzBipontinus,
and subsequently by the late Professor F. W. Klatt, but the
name seems not to have been hitherto published.
T am indebted to Professor Eug. Warming for the identifica-
tion of my no. 114 with the Liebmann plant in the Botanical
Museum at Copenhagen.
TAGETES JALISCENSIS Greenm., var minor Greenman, var. nov.
Leaves 3 cm. or less in length, about two-thirds as broad:
heads 1.3 to 1.8 cm. high; involucre i to 1.4 cm. long: mature
achenes 7 to 7.5 mm. in length. MEXICO. State of Oaxaca:
Cercainas de Oaxaca, altitude 1550 m., 8 November, 1906, C.
Conzatti, no. 1516 (hb. Field Mus.). The chief distinguishing
characters separating var. minor from the species are the smaller
leaves, shorter involucre and flowers.
Dysodia (Gymnol8ena) oaxacana Greenman, sp. nov.
Shrub: stem and branches covered with a grayish bark; ulti-
mate branchlets terete, somewhat striate, greenish or reddish-
brown, minutely pubescent in decussating lines: leaves opposite,
sessile or essentially so, lanceolate, i to 4 cm. long, 4 to n mm.
broad, acute, finely serrulate, narrowed below to an entire or
occasional subsetiferous subpetiolate base, pellucid-punctate,
slightly pubescent on both surfaces under a lens especially on
the midrib of the upper surface, or glabrous : heads few, hetero-
gamous, 1.5 to 2 cm. high, terminating the stem and branches
on minutely bracteate 1.5-5 cm. -long glabrous peduncles: involu-
cre at first narrowly campanulate or subcylindric, often split on
one side, 12 to 14 mm. long, (7-) 8-dentate, naked or essentially
so at the base, glabrous, bearing several linear-elliptic glands,
later separating into more or less distinct lance-linear bracts;
teeth, or the terminal portion of the individual bracts, short,
triangular, acute, pubescent-tipped: ray-flowers commonly 8;
rays including the slender tubular portion 1.5 cm. long, 4 to
5 mm. broad, deep orange-red: disk-flowers about 30: pappus
of numerous lacerate-fimbriate scales, tawny: achenes 6 mm.
long, pubescent. MEXICO. State of Oaxaca: Almoloyas,
274 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
altitude 800 m., 25 December, 1906, C. Conzatti, no. 1653 (hb.
Field Mus. Catalogue no. 195851 ; fragment in hb. De Candolle).
I am indebted to Dr. Casimir De Candolle for a detailed com-
parison of Professor Conzatti's plant with the type of Dysodia
serratifolia DC. From this species, as pointed out by Dr. De
Candolle, D. oaxacana differs in having smaller leaves, few heads
instead of a multiflorous corymb, and pubescent-tipped invo-
lucral bracts.
To this natural group also belong D. integrifolia Gray, and
D. Seleri Rob. & Greenm. From the former the species described
above is readily separated by the pubescent branchlets and
puberulent leaves ; and from the latter by the presence of pellucid
glands on leaves and involucre.
CHRYSACTINIA MEXICANA Gray, PI. Fendl. 93 (1849). Pectis taxi-
folia Greene. Leaflets Bot. Obs. & Crit. i. 148 (1905).
Specimens collected at Kingston, New Mexico, by Mr. O. B.
Metcalfe, and distributed under no. 1440 (hb. Field Mus.), as
"Pectis taxifolia, Greene, n. sp." are identical with the above
species of Chrysactinia.
LIABUM CADUCIFOLIUM Rob. & Bart. Proc. Am. Acad. xliii. 59-
(1907).
To this very distinct species must be referred specimens
collected at Paso del Rio, State of Colima, Mexico, November,
1906, Dr. G. M. Emrick, no. 187 (hb. Field Mus.). Dr. Emrick's-
specimens show the uppermost leaves to be ovate, acuminate,
acute, entire or remotely mucronate-denticulate and abruptly
narrowed below the middle to an acute base. The mature
achenes, moreover, are 4 mm. in length, pubescent and strongly
striate.
Schistocarpha platyphylla Greenman, sp. nov.
Stem terete, striate, glabrous or slightly pubescent above:
leaves opposite, petiolate, ovate, 10 to 25 cm. long, 6.5 to 22 cm.
broad, acuminate, acute, sinuate-dentate, subcordate to cuneate
at the base and decurrent on the subconnate petioles, 3-nerved,
sparingly pilose on both surfaces, slightly paler beneath, thin
and membranous: inflorescence a terminal subcorymbose many-
headed pubescent panicle; bracts linear, subsetaceous : heads
7 to 8 mm. high: involucre narrowly campanulate; bracts of
the involucre 3-seriate, lance-oblong, 3 to 5 mm. long, i to 1.5 mm.
broad, obtuse or obtusish, pale stramineous, striate, glabrous:
pales hyaline, subcuneate, about 3 mm. long, irregularly 3-5-
toothed: pistillate flowers usually 13; corollas tubular; tube
2 mm. long, externally pubescent: perfect flowers 14 to 18;.
corollas about 5 mm. long, slightly exceeding the pappus; tube
puberulent, gradually ampliated above into a 5 -toothed limb:
mature achenes i mm. long, glabrous or sparingly puberulent
under a strong lens. GUATEMALA. Department of Quezal-
tenango: Santa Maria, 5 February, 1906, W. A. Kellerman, no.
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 275
5295 (hb. Field Mus.), type; south side of the volcano of Santa
Maria, altitude 1370 m. ( 19 January, 1907, W. A. Kellerman,
no. 6114 (hb. Field Mus.). The affinity of this species is with
Schistocarpha paniculata Klatt, from which, however, it differs
in having a more sparse tomentum on stem and foliage, larger
leaves, smaller heads with fewer flowers, and also in the shorter
broader pales.
Senecio (Erexnophili) durangensis Greenman, nom. nov. 5.
ctenophyllus Greenm. Proc. Am. Acad. xliii. 20 (1907), not Phil.
Senecio ( Sanguisorboidei) coahuilensis Greenm.
An herbaceous perennial, glabrous or essentially so through-
out: stem erect, 3 to 4 dm. high, branched, striate: leaves
pinnatifid, 5 to 10 cm. long, 2 to 3 cm. broad, thickish in texture
and slightly glaucous, glabrous on both surfaces or puberulent
beneath; segments obtusely crenate-dentate, the terminal
segment largest and subreniform, the lateral ones obovate; the
lowermost leaves petiolate, the upper stem-leaves sessile and
amplexicaul: inflorescence terminating the stem and branches
in a compound corymbose cyme: heads about 7 mm. high,
radiate: involucre campanulate, calyculate with 2 or 3 small
bracteoles, glabrous; bracts of the involucre 13 (-15), linear-
lanceolate, 4 to 5 mm. long: ray-flowers 8 to 10; rays oblong,
3 to 4 mm. long, 4-nerved: disk-flowers 35 to 40: achenes gla-
brous, striate. Monogr. Senecio, I Th. 23 (1901) & in Engl. Bot.
Jahrb. xxxii. 19 (1902), without complete description. MEXICO.
State of Coahuila: Lerios, February to October, 1880, Dr.
Edward Palmer, no. 755 (hb. Gray, and hb. Kew). This species
has been confused with 5. Sanguisorbce DC., but it is amply
distinct in habit, size and foliar characters.
Senecio ( Sanguisorboidei) Ervendbergii Greenm.
Glabrous throughout: stem herbaceous, leafy, striate: upper
stem-leaves thin, membranous, sublyrately pinnatifid, sessile,
more or less expanded and subamplexicaul at the base ; segments
7 to 9, sublaciniate, the ultimate divisions usually 3-toothed and
the teeth tipped by a short mucro; terminal segment obovate-
rotund, the lateral oblong or somewhat obovate-oblong; midrib
more or less winged throughout and often bearing small tooth-
like expansions between the main segments; inflorescence a
terminal compound corymbose cyme: heads small about 5 mm.
high, on slender pedicels: involucre campanulate, ecalyculate;
bracts of the involucre about 2 1 , slightly shorter than the flowers
of the disk: ray-flowers 12 to 14: disk-flowers 75 to 80: achenes
hispidulous. Monogr. Senecio, I Th. 23 (1901) & in Engl. Bot.
Jahrb. xxxii. 19 (1902) without complete characterization.
MEXICO. State of Vera Cruz: Wartemberg, near Tantoyuca,
province of Huasteca, October, 1858, C. L. Ervendberg, no. 90
(hb. Gray). Nearly related to 5. tampicanus DC.
276 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
Senecio ( Sanguisorboidei) leonensis Greenm.
An herbaceous perennial, more or less lanate-tomentose through-
out, somewhat glabrate in age: stem 2 to 3 dm. high, leafy at
the base, essentially naked above: leaves petiolate, pinnatifid,
including the petiole 8 to 12 cm. long, about 3 cm. broad, at
first lanate-tomentose on both surfaces, later glabrate: segments
or lobes rather coarsely, somewhat unequally and sharply
toothed ; the terminal segment subreniform, the lateral ones
(3 to 6 on either side) obovate-cuneate, becoming smaller towards
the petiole: heads few, about i cm. high, radiate: involucre
campanulate, subecalyculate and, as well as the bracteate pedun-
cles, tomentulose; bracts of the involucre usually 21, lanceolate,
acute, 6 mm. long: ray-flowers about 13; rays oblong, 6 to 7
mm. in length, 4-5-nerved: disk-flowers numerous, about 60:
achenes pubescent. Monogr. Senecio, I Th. 23 (1901) & in Engl.
Bot. Jahrb. xxxii. 19 (1902), without complete characterization.
MEXICO. State of Nuevo Leon : Sierra Madre, near Monterey,
i June, 1889, C. G. Pringle, no. 2894 (hb. Gray).
Senecio (Aurei) cyclophyllus Greenman, sp. nov.
An herbaceous perennial: stem simple, 3.5 to 5 dm. high,
sparingly tawny-tomentose at the base and in the leaf-axils
otherwise glabrous, striate, purplish: radical and lowermost
stem-leaves subrotund, 4 to 7 cm. long, equally broad, cordate,
crenate-dentate, green and glabrous above, more or less purplish
and sparingly hirsute to glabrous beneath; petioles 5 to 8 cm.
long: upper stem -leaves sessile, amplexicaul, lyrately pinnatifid
with a relatively large terminal segment and narrowly obovate-
cuneate unequally dentate lateral divisions : inflorescence termin-
ating the stem in a many-headed subcorymbose cyme; bracts
of the inflorescence linear-lanceolate to subulate, tawny-tomentu-
lose, purplish-tipped: heads 7 to 9 mm. high, radiate: involucre
campanulate, sparingly calyculate with minute bracteoles; bracts
of the involucre about 21, lance-linear, 5 to 6 mm. long, acu-
minate, acute, more or less purplish-tipped, glabrous: ray-
flowers commonly 13; rays orange-yellow: disk-flowers 50 to
60: mature achenes 2 mm. long, hispidulous. MEXICO. State
of Nuevo Leon: near Monterey, 1906, C. G. Pringle, no. 10,230
(hb. Gray), type; Cerro la Scilla, near Monterey, 20 March, 1902,
E. W. Nelson, no. 6,672 (hb. U. S. Nat. Mus. and hb. Gray). A
species somewhat intermediate in general aspect between 5. aureus
L. and 5. Cardamine Greene. From the former it differs in having
shorter petioles and a more rotund leaf -blade to the basal leaves ;
besides the terminal segment of the stem-leaves is relatively
broader and more reniform, and finally the achenes are hispidul-
ous. From 5. Cardamine, on the other hand, S. cyclophyllus
differs in its larger stature, size of foliage and leafy stem, and in
the many-headed inflorescence.
Senecio ( Aurei) Kosei Greenm.
An herbaceous perennial: root-stocks rather short, bearing
several fleshy-fibrous roots: stem erect, 4 to 4.5 dm. high, un-
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 277
branched, glabrous below, slightly pubescent above, terminated
by a single large radiate head, or bearing in addition i to 3 re-
duced heads: leaves petiolate, ovate, 2 to 3 cm. long, two-thirds
as broad, obtuse, glabrous on both surfaces, subentire to spar-
ingly and obtusely dentate; the lower leaves on long petioles,
2 to 4 times as long as the blade; the upper stem-leaves some-
what laciniate-dentate or sublyrate, not infrequently expanding
at the base and subamplexicaul : peduncles somewhat enlarged
above, pubescent: heads about 12 mm. high, including the rays
3 to 3.5 cm. broad: involucre campanulate, essentially naked;
bracts of the involucre about 20, lanceolate-linear, 8 to 10 mm.
long, acute, green, scarious-margined, glabrous except at the
penicillate tip: ray-flowers 10 to 12; rays conspicuous, light
yellow: 'disk-flowers numerous: achenes glabrous. Monogr.
Senecio, I Th. 24 (1901) & in Engl. Bot. Jahrb. xxxii. 20 (1902),
without complete characterization. MEXICO. Territory of
Tepic: in the Sierra Madre, near Santa Teresa, 10 August, 1897,
Dr. J. N. Rose, no. 2,157 ( h b- Gray, and hb. U. S. Nat. Mus.).
Senecio ( Amplectentes) heterodontus Greenman, sp. nov.
An herbaceous perennial: roots fleshy-fibrous from a short
perpendicular root-stock: stem erect, simple or branched, 0.5 to
i m. high, nearly naked above, striate, glabrous to crisp-pubes-
cent: leaves lanceolate, i to 3 dm. long, i to 4 cm. broad, slightly
hirtellous-puberulent to essentially glabrous on the upper sur-
face, pubescent beneath; the lowermost leaves, obtuse or acute,
narrowed below, rather long winged-petiolate, unequally dentate
or dentate-lobed ; the upper stem-leaves, remote, sessile, amplexi-
caul, acuminate, more or less regularly denticulate, becoming
bract-like toward the inflorescence: heads numerous, 8 to 10 mm.
high, radiate, disposed in a terminal corymbose cyme: in-
volucre campanulate, calyculate, glabrous, or slightly pubescent
at the base: bracts of the involucre 13 (-21), lanceolate, 4.5 to
5.5 mm. long, acutish, terminated by a blackish penicillate tip:
ray-flowers commonly 8 ; tube pubescent ; pappus somewhat
exceeding the tube: disk-flowers 48 to 50: achenes striate, gla-
brous. 5. potosinus Greenm. Monogr. Senecio, I Th. 25 (1901) &
in Engl. Bot. Jahrb. xxxii. 21 (1902), without complete character-
ization, not 5. potosianus Klatt. MEXICO. State of San Luis
Potosi: Valley of San Luis Potosi, in mountains near San
Miguelita, September, 1876, Dr. J. G. Schaffner, no. 280 (hb.
Gray), type; in the region of San Luis Potosi, collection of 1878,
Parry & Palmer, no. 536 (hb. Gray); Alvarez, 13 to 23 July,
1904, Dr. E. Palmer, no. 237 (hb. Gray, and hb. Field Mus.).
This species may be found in herbaria under the name of "Senecio
multidentatus . "
Senecio ( Amplectentes) mohinorensis Greenm.
Stem erect, striate, puberulent: leaves sessile, amplexicaul,
pinnately lobed, lanceolate in general outline, 5 to 15 cm. long,
i to 4 cm. broad, puberulent. on both surfaces, slightly paler
278 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
beneath; lobes oblong, mucronate, acute, entire or few-toothed;
the uppermost leaves reduced and laciniate-dentate to entire:
inflorescence a terminal comparatively few-headed panicle; the
branches usually bearing 3 heads: heads about 1.5 cm. high,
radiate: involucre campanulate, conspicuously calyculate with
linear-lanceolate green bracteoles; involucral bracts proper,
lanceolate to lance-oblong, 10 to 12 mm. long obtuse, scarious-
margined and, as well as the bracteoles, black-tipped and hirsute-
pubescent with spreading jointed hairs: ray-flowers 8 to 10; rays
yellow, elliptic-oblong, 10 to 12 mm. long, 4-nerved, the two
middle nerves often branching into 3 divisions: disk-flowers
about 50: achenes canescent-pubescent. Monogr. Senecio, I Th.
25 (1901) & in Engl. Bot. Jahrb. xxxii. 21 (1902), without com-
plete characterization. MEXICO. State of Chmuahua: Mt.
Mohinora, i September, 1898, E. W. Nelson, no. 4881 (hb. Gray,
and hb. U. S. Nat. Mus.).
Senecio ( Amplectentes) platypus Greenman, sp. nov.
An herbaceous perennial: stem simple or branched, striate,
purplish, lightly floccose-tomentulose: leaves petiolate, ovate
to ovate-lanceolate, 3 to 9 cm. long including the petiole, i to
4 cm. broad, mucronate-acute, entire to irregularly dentate,
cuneate to subcordate at the base, floccose-tomentulose above,
densely and permanently white tomentose beneath, thin and
membranous; petioles 4 cm. or less in length, usually expanded
and clasping the stem by an auriculate-stipuliform base: inflo-
rescence a terminal few-headed tomentulose subcorymbose cyme;
bracts linear-setaceous: heads 10 to 12 mm. high, radiate:
involucre narrowly campanulate, calyculate with minute setace-
ous bracteoles; bracts of the involucre usually 21, lance-linear,
6 to 8 mm. long, glabrous except at the base, black-tipped: ray-
flowers about 13; rays yellow: disk-flowers 40 to 45: achenes
densely sericeous-hirtellous. MEXICO. State of Nuevo Leon:
Sierra Madre near Monterey, 21 December, 1906, C. G. Pringle,
no. 10,352 (hb. Gray), type; on limestone ledges near Monterey,
12 March, 1906, C. G. Pringle, no. 13,882 (hb. Gray). The
conspicuous stipular-like development at the base of the petioles
renders this species easily recognizable, and quite distinct from
any other species known to the writer.
Senecio ( Mulgedifolii) Cpnzattii Greenm.
An herbaceous perennial: stem erect, simple or branched
above, about i m. high, striate, subangulate, at first arachnoid-
tomentulose, more or less glabrate: leaves oblong-lanceolate to
lance-attenuate, 0.5 to 3 dm. long, 0.5 to 5 cm. broad, acute,
glabrous or sparingly arachnoid above, permanently arachnoid-
tomentose beneath over a green to more or less purple leaf-sur-
face, dentate to merely callous-denticulate; the lowermost
leaves gradually narrowed below the middle into a winged
petiolar base, coarsely dentate to lyrately subpinnatifid ; the
upper leaves sessile and amplexicaul, becoming reduced towards
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 279
the corymbose-paniculate inflorescence to attenuate bracts:
heads 10 to 12 mm. high, discoid, 35-4o-flowered: involucre
narrowly campanulate, calyculate with rather conspicuous
bracteoles and, as well as the peduncles, pubescent with spread-
ing hirsutish hairs; bracts of the involucre usually 13, lanceolate,
^cutish, slightly penicillate-tipped and, as well as the corollas,
more or less purplish : achenes striate-ribbed, glabrous. Monogr.
Senecio, I Th. 25 (1901) & in Engl. Bot. Jahrb. xxxii. 21 (1902),
without complete characterization. MEXICO. State of Oaxaca:
Cerro de San Felipe, altitude 3,000 m., 14 November, 1897, C.
Conzatti & V . Gonzdlez, no. 559 (hb.Gray), type; Oaxaca, altitude
i,75om., July-August, 1900, C. Conzatti & V. Gonzalez, no. 1,003
(hb. Gray) ; Hacienda de Caciques, District of Cuicatlan, altitude
2,130 m., 14 August, 1895, Rev. Lucius C. Smith, no. 613 (hb.
Gray); mountains of Oaxaca, Cuming (hb. Gray).
Senecio ( Mulgedifolii) decorus Greenm.
Stem erect, striate-grooved, glabrous or nearly so, more or
less purplish; leaves runcinate--pinnatifid, 0.5 to 2.5 dm. long,
2 to 14 cm. broad, glabrous above, arachnoid-pubescent beneath;
terminal lobe largest, somewhat triangular-acuminate; lateral
lobes lanceolate to oblong-lanceolate, acuminate, acute, margin
dentate with horizontally spreading and slightly unequal teeth;
upper stem-leaves sessile and amplexicaul: inflorescence a term-
inal rather compact panicle: heads 12 to 15 mm. high, not in-
frequently somewhat nodding, discoid: involucre campanulate,
conspicuously calyculate with broadish bracteoles: bracts of
the involucre lance-linear and acute to somewhat oblong and
slightly expanded above the middle to an obtuse apex, glabrous,
the inner with scarious and slightly lacerated margins: flowers
35 to 45; corollas exceeding the involucre and, as well as the
bracts and bracteoles, reddish-purple: achenes striate-ribbed,
glabrous. Monogr. Senecio I Th. 25 (1901) & in Engl. Bot.
Jahrb. xxxii. 21 (1902). GUATEMALA. Department of Zacate-
pequez: Volcan de Agua, altitude 3,350 m., April, 1890, John
Donnell Smith, no. 2,361 (hb. Gray); Todos Santos, altitude
3,045 m., 26 December, 1895, E. W. Nelson, no. 3,637 (hb. Gray,
and hb. U. S. Nat. Mus.).
Senecio ( Mulgedifolii) jacalensis Greenm.
An herbaceous perennial: roots fleshy-fibrous: stem erect,
3 dm. or more high, from a thickish perennial base, rather leafy,
striate, below glabrous, above especially in the inflorescence
pubescent: lower leaves oblanceolate, 0.5 to 1.5 dm. long, 1.5
to 2.5 cm. broad, obtuse or acute, shallowly callous-dentate,
narrowed below into a winged petiole, glabrous on both surfaces;
upper stem-leaves sessile, amplexicaul, acuminate and more or
less imbricated on the stem: inflorescence subracemose or pani-
culately racemose: heads large, 13 to 15 mm. high, discoid, not
infrequently somewhat nodding: involucre campanulate, calycu-
late with rather large bracteoles, glabrous or slightly pubescent
280 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
at the base; bracts of the involucre subbiseriate, about 21,
lanceolate-linear, i cm. long, obtuse or obtusish: flowers numer-
ous, about 70: corollas, as well as the bracts of the involucre,
often reddish: achenes striate, glabrous.- Monogr. Senecio,
I Th. 25 (1901) & in Engl. Bot. Jahrb. xxxii. 21 (1902), without
complete characterization. Senecio helodes, Hemsley, Biol.
Cent. -Am. Bot. ii. 241 (1881), in part, not Benth. Cacalia race-
mosa, Schz. Bip., ace. to Hemsl., 1. c. MEXICO. State of Gua-
najuato: Jacal, Ehrenberg, no. 1,293 (hb- Gray, hb. Roy. Bot.
Mus. Berlin, and hb. Kew).
Senecio ( Mulgedifolii) rhyacophilus Greenm.
Stem herbaceous, glabrous: leaves runcinate-pinnate with
rather unequally and irregularly mucronate-dentate or sublobate
segments and with rounded sinuses between the segments, gla-
brous on both surfaces, paler beneath, thin or membranous in
texture, more or less expanded below into a lac.iniate-dentate
amplexicaul base, 0.5 to 2 dm. long, i to 6 cm. broad; the upper-
most leaves undivided, lance-attenuate from a rather broad
base, coarsely toothed: inflorescence a terminal somewhat
leafy pubescent panicle: heads 12 to 15 mm. high, discoid, about
24-flowered : involucre calyculate with linear-setaceous bracteoles,
glabrous; bracts of the involucre usually 13, linear, about i cm.
long, acute to obtusish and, as well as the corollas, more or less
purplish: achenes striate, glabrous. Monogr. Senecio, I Th. 25
(1901) & in Engl. Bot. Jahrb. xxxii. 21 (1902), without complete
characterization. GUATEMALA. Department of Zacatepiquez :
Volcan Fuego, altitude 2,735 m -> November, 1889. Heyde &
Lux, no. 4,502 (hb. Gray), exsiccatas of John Donnell Smith.
Department of Quiche: Chiul, altitude 2,400 m., April, 1892,
Heyde & Lux, no. 3,379 (hb. Gray), exsiccatas of John Donnell
Smith.
Senecio ( Fruticosi) hirsuticaulis Greenm.
Shrub: stem above and the branches, as well as the petioles,
densely hirsute-pubescent with spreading hairs: leaves petiolate,
ovate-oblong, 5 to 8 cm. long, 3 to 6 cm. broad, subcordate,
slightly hirtellous above especially on the midrib and lateral
nerves, later nearly glabrous, densely and permanently hirsute-
tomentose beneath, subangulately 5-7-lobed with broad shallow
sinuses, mucronate-denticulate ; lobes conspicuously mucronate-
acute; petioles stoutish, about 2 cm. long: inflorescence a ter-
minal compound hirsute- tomentose corymb; peduncles seta-
ceous-bracteate : heads about i cm. high, radiate: involucre
narrowly campanulate, calyculate with setaceous bracteoles;
bracts of the involucre usually 13, linear to lance-oblong, acutish,
the inner with scarious suberose margins : ray-flowers commonly
8; rays yellow, 4-nerved: disk-flowers 20 to 25, exceeding the
involucre: achenes glabrous. Monogr. Senecio, I Th. 26 (1901)
. & in Engl. Bot. Jahrb. xxxii. 22 (1902), without complete char-
acterization. MEXICO. En route from San Luis Potosi to
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 281
Tampico, December, 1878 to February, 1879, Dr. E. Palmer,
no. 1,114 (hb. Gray).
Senecio ( Fruticosi) santarosse Greenman, sp. nov.
Shrub: stem below covered with a grayish cortex, above
more or less arachnoid-tomentulose : leaves petiolate, oblong-
lanceolate, i to 2.5 dm. long, 3 to 7 cm. broad, acute or acumi-
nate, shallowly sinuate-dentate, narrowed below to an obtuse
base, rather prominently reticulate-veined, glabrous on both
surfaces or slightly pubescent beneath especially on the midrib
and lateral nerves, glabrate; petioles 3 to 7.5 cm. long, at first
tomentulose, later becoming glabrous: inflorescence a terminal
many-headed corymbose panicle: heads about i cm. high,
radiate: involucre arachnoid-tomentulose to essentially gla-
brous; bracts of the involucre oblong, 5 to 7 mm. long, obtuse
scarious-margined : ray-flowers usually 2; rays 4 to 5 mm. long,
yellow: disk-flowers 3 to 5; corollas rather deeply 5-toothed r
somewhat zygomorphic: pappus about as long as the corollas,
of the disk-flowers, copious, white: achenes glabrous. Monogr.
Senecio, I Th. 26 (1901) & in Engl. Bot. Jahrb. xxxii. 22 (1902).
Senecio Ghiesbreghtii, var. pauciftorus Coulter, Bot. Gaz. xvi.
101 (1891), not 5. pauciftorus Pursh. GUATEMALA. Depart-
ment of Santa Rosa: La Vega, altitude 1525 m., February, 1893,
Heyde & Lux no. 4,520 (hb. Gray), exsiccatae John Donnell
Smith, type. Department of Guatemala: Sapote, altitude, 1,310
m., March, 1890, John Donnell Smith, no. 2,359 (hb. Donnell
Smith, and hb. Gray). Department of Quezaltenango : Santa
Maria, 5 February, 1906, W. A. Kellerman, no. 5,277 (hb. Field
Mus.). Department of Solala: Volcano of Atitlan, 16 February,
1906, W. A. Kellerman, no. 5,353 (hb. Field Mus.).
Senecio ( Palmatinervii) adenolepis Greenman, sp. nov.
Perennial : stem covered with a gray cortex ; ultimate branch-
lets glandular-puberulent : leaves petiolate, palmately nerved,
ovate-orbicular, 5-lobed, slightly peltate, in specimens at hand
7 cm. long, 7 to 8 cm. broad, dark green and hirtellous-puberu-
lent above, paler and crisp-hirtellous beneath, mucronate-denticu-
late; lobes triangular-ovate and terminated by a mucro; peti-
oles 6.5 cm. in length, slightly puberulent, purplish: inflorescence
a terminal few-headed granulose-glandular cymose panicle;,
bracts lanceolate to linear-setaceous: heads 10 to 14 mm. high,
radiate: involucre narrowly campanulate; bracts of the
involucre usually 8, oblong-lanceolate, about i cm. long, acumi-
nate, acute or acutish, externally short-glandular-pubescent
with a few hirsute hairs intermixed: pappus copious, 6 to 7 mm.
long, white: ray-flowers 5, about i cm. long; tube of the corolla
4 to 5 mm. long, externally sparingly pubescent; rays equalling
or somewhat exceeding the tube, yellow: disk-flowers about 15;
corollas 9 mm. long with a short tube gradually amplicated
above into the 5-toothed limb; achenes glabrous. MEXICO.
State of Morelos: Sierra de Tepoxtlan, altitude 2,285 m -> J 3
282 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
February, 1907, C. G. Pringle, no. 13,909 (hb. Gray). Related
to 5. cordobensis Hemsl., but differs in having smaller leaves,
longer involucral bracts and more numerous flowers in the heads.
Senecio ( Palmatinervii) eriophyllus Greenman, sp. nov.
Shrub : stem in the dried state of a dark gray or blackish wood,
covered with a light gray cortex; ultimate branches white
floccose-tomentose in the early stages, glabrate: leaves petiolate,
ovate to ovate-oblong, 6 to 10 cm. long, 5 to 8 cm. broad, sinuate-
angulate-lobed, entire or sparingly mucronate-denticulate, cun-
eate to subcordate at the base, at first densely and softly tomen-
tose on both surfaces, later arachnoid-tomentose especially
above; the lobes terminated by a short subcartilaginous mucro;
petioles 2 to 3 cm. long, tomentose: inflorescence terminating
the stem in a many-headed floccose-tomentose panicle: heads
about 12 mm. high: involucre calyculate with minute subulate
bracteoles, glabrous except at the base; bracts of the involucre
8, lance-linear to lance-oblong, 7 to 8 mm. long, obtuse or obtusish,
bluntly keeled on the back, stramineous : flowers 12 to 15:
achenes glabrous. MEXICO. State of Oaxaca: hills near Tula,
altitude 1680 m., 20 May, 1906, C. G. Pringle, no. 13,864 (hb.
Gray). This species has its nearest affinity with 5. albonervius
Greenm.
Senecio ( Palmatinervii) Gilgii Greenm.
Suffrutescent : stems at least above densely pubescent with
spreading tawny jointed hairs: leaves long-petiolate, rotund,
i to 3 dm. broad, cordate, 7~9-nerved from the base, sinuately
lobed and the lobes again subtrilobate, mucronate-denticulate.
hirtellous-pubescent on both surfaces; petioles 8 to 14 cm. long,
densely pubescent: inflorescence a terminal panicle; heads
large, 1.5 to 1.7 cm. high, radiate: involucre campanulate,
calyculate, densely hirsute-pubescent; bracts of the involucre
usually 13, oblong to oblong-lanceolate, nearly or quite i cm.
long, 2.5 to 4 mm. broad, obtuse, more or less overlapping so
as to appear subbiseriate, the innermost bracts with scarious
margins: ray-flowers 8 to 10; ligules 6 to 7 mm. long, yellow,
4-5-nerved: disk-flowers about 30: achenes conspicuously
striate, glabrous. Monogr. Senecio, I Th. 26 (1901) & in Engl.
Bot. Jahrb. xxxii. 22 (1902), without complete characterization.
- MEXICO. State of Chiapas: near Pinabete, altitude 2,000 to
2,460 m., 8 February, 1896, E. W. Nelson, no. 3,773 (hb. Gray,
hb. U. S. Nat. Mus., and hb. Royal Bot. Mus. Berlin).
SENECIO HETEROGAMUS Hemsl., var, Kellermanii Greenman, var.
nov.
Leaves long-petiolate, suborbicular or somewhat reniform,
7 to 15 cm. long, 9 to 20 cm. broad; petioles 4 to 14 cm. long;
inflorescence including the involucre villous-hirsute with long
jointed red or reddish-brown hairs: other characters as in the
species. GUATEMALA. Department of Sacatepequez: Volcano
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 283
of Agua, 15 February, 1905, W. A. Kellerman, no. 4,706 (hb.
Field Mus.).
Senecio ( Palmatinervii) lanicaulis Greenm.
Shrub: stem, at least above, densely lanate-tomentose : leaves
petiolate, subrotund to reniform, cordate, palmately 7-9-nerved,
0.7 to 2.5 dm. broad, shallowly sinuate-lobed, margined with
unequal conspicuous more or less curved mucronulations, at
first tomentulose above, densely and permanently lanate-tornen-
tose beneath; petioles stout, 0.3 to 1.5 dm. long: inflorescence
a terminal many-headed lanate-pubescent compact compound
corymb; bracts setaceous: heads i cm. high, radiate: involucre
narrowly campanulate, calyculate, tomentulose at the base,
glabrous above; bracts of the involucre about 13, lanceolate
to lance-oblong, acutish, essentially glabrous, the inner scarious-
margined: ray-flowers 6 to 8; ligules 4-nerved: disk-flowers
12 to 20, exceeding the involucre: achenes striate, glabrous.
Monogr. Senecio, I Th. 26 (1901), & in Engl. Bot. Jahrb. xxxii. 22
(1902), without complete characterization. MEXICO. State of
Chiapas: near Pinabete, altitude 1,800 to 2,400 m., 8 February,
1896, E. W. Nelson, no. 3,771 (hb. Gr., and hb. U. S. Nat.
Mus.). GUATEMALA. Department of Quiche": Chiul, altitude
2,400 m., April, 1892, Heyde & Lux, no. 3,377 (hb. Gray), exsic-
catae of John Donnell Smith.
Senecio ( Palmatinervii) Langlassei Greenm.
Shrub 3-4 m. high: leaves petiolate, palmately nerved, ovate-
rotund in general outline, 1-2 dm. long, equally broad, 7-13-
lobed, densely granulose on the upper surface with hirsutish
hairs intermixed, especially on the nerves, white-tomentose
beneath as well as on the petioles; lobes mucronate-apiculate,
denticulate-margined: inflorescence a terminal round-topped
many-headed paniculate cyme: heads i cm. or less high, radiate:
involucre narrowly campanulate, sparingly calyculate; bracts of
the involucre narrowly oblong-lanceolate, acutish, scarious-
margined, dorsally granulose-glandular, thickened along the
median line: ray-flowers 6 to 8 with a slender 5 mm. -long puber-
ulent tube equalling the narrowly oblong 4-nerved yellow ray:
disk-flowers 15 to 20: achenes glabrous. Monogr. Senecio, I
Th. 26 (1901) & in Engl. Bot. Jahrb. xxxii. 22 (1902), without
complete characterization. MEXICO. "Etats de Michoacan et
de Guerrero," Sierra Madre, altitude 1,600 m., 21 April, 1899,
E. Langlasse no. 1,005 (hb. Gray, and hb. Roy. Bot. Mus.,
Berlin).
Senecio ( Palmatinervii) reg-lensis Greenm.
Stout herbaceous perennial, i to 2 m. high: stem glabrous,
smooth, brownish-lineolate at least above: leaves petiolate,
palmately nerved, ovate-rotund, 5 to 7 cm. or more long, nearly
or quite as broad, rather deeply 5-7-lobed, hirsutish-pubescent
on both surfaces especially beneath; lobes acute mucronate-
284 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
apiculate; margins cartilaginous-denticulate: inflorescence a
. , terminal round-topped many-headed paniculate cyme: heads
subcylindrical, 10 to 12 mm. high, radiate: involucre calyculate
with bracteoles less than half the length of the 8 oblong-lanceolate
obtusish bracts of the involucre: ray-flowers mostly 6: disk-
flowers 1 8 to 20, equalling or slightly exceeding the involucre;
pappus about as long as the corolla : achenes glabrous. Monogr.
Senecio, I Th. 26 (1901) & in Engl. Bot. Jahrb. xxxii. 22 (1902),
without complete characterization. MEXICO. State of Vera
Cruz: Regla, Ehrenberg, no. 454 (hb. Gray).
Senecio ( Multinervii) Cooperi Greenm.
Stout herb ? : stem at least above pubescent with brownish hir-
tellous hairs: leaves large, oblong-ovate, including the petiole
3 to 3.5 dm. long, i to 1.5 dm. broad, rather blunt or narrowed
at the apex, mucronate-acute, merely denticulate on the margins
to somewhat sinuate and unequally dentate, the teeth tipped
with a cartilaginous mucro, few and irregularly lobed at the
base with rounded open sinuses, decurrent on the petiole, spar-
ingly puberulent on both surfaces; midrib and the numerous
lateral nerves prominent beneath: inflorescence corymbose:
heads 1.5 cm. high, radiate: involucre barely calyculate with a
few small inconspicuous bracteoles; bracts of the involucre
8, linear, acutish, slightly penicillate-tipped, otherwise glabrous:
ray-flowers about 8; rays linear-oblong, conspicuous: disk-
flowers with a long slender tube and a rather deeply 5 -lobed
limb: achenes pubescent. Monogr. Senecio, I Th. 26 (1901) & in
Engl. Bot. Jahrb. xxxii. 22 (1902), without complete characteri-
zation. COSTA RICA. Province of Cartago: Cartago, altitude
1,310 m., December, 1887, Juan J. Cooper, no. 5,803 (hb. Gray),
exsiccatae John Donnell Smith, distributed as "Senecio multi-
venius Benth."; "la Division, vallde du Ge"ne"ral", altitude
2,160 m., Pittier, no. 3,405 (hb. Gray), in part.
Senecio ( Multinervii) megaphyllus Greenman, sp. nov.
Stem above arachnoid-tomentulose, striate: leaves large,
oblong-oblanceolate, including the petiole 3 to 5 dm. long, i
to 2 dm. broad, acute or acuminate-acute, more or less sinuate,
cartilaginous-dentate, narrowed below into a winged petiole
half-clasping the stem, arachnoid-tomentulose on both surfaces,
somewhat glabrate above except on the midrib; midrib and
lateral nerves prominent beneath: inflorescence a terminal
corymbose-panicle: heads many, on slender setaceous-bracteolate
peduncles: involucre cylindrical; bracts of the involucre 8,
linear, 10 mm. long, i mm. broad, acutish, glabrous, stramineous,
scarious-margined : ray-flowers 3 to 5, yellow: disk-flowers 5 to 8,
rather deeply 5-lobed: achenes glabrous. 5. multivenius, var.
oliganthus Greenm. Monogr. Senecio, I Th. 26 (1901) & in Engl.
Bot. Jahrb. xxxii. 22 (1902), without complete characterization,
not S. oliganthus DC. COSTA RICA. "Bords du Paraita
Grande au Copey," altitude 1,800 m., A. Tonduz, no. 11,700 (hb.
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 285
Gray, and hb. Inst. Physico.- Geogr. Cost. Rica); "bords du
rio Paraita au Copey," altitude 1,800 m.,A. Tonduz, no. 11,844
(hb. Gray, and hb. Inst. Physico.-Geogr. Costa Rica). The
long slender cylindrical few-flowered heads with longer and
narrower involucral bracts, as shown by additional material,
render this species of easy separation from 5. multivenius Benth.
with which it has been hitherto associated.
Senecio ( Terminales) chicharrensis Greenm.
Fruticose: stem leafy and white-tomentose above: leaves
long-petiolate, oblong-ovate, 1.2 to 2.7 dm. long, i to 1.8 dm.
broad, subcordate to abruptly contracted below to an unequal
base, glabrous above, floccose-tomentose beneath especially
along the prominent midrib and lateral nerves, sinuately lobed;
lobes mucronate, acute, remotely mucronate-denticulate ; petioles
8 to 10 cm. long, more or less floccose-tomentose: inflorescence
abruptly terminating the stem in small and many-headed floc-
culent close compound corymbs on rather long peduncles : heads
about 8 mm. high, radiate: involucre 4 mm. high, essentially
ecalyculate or with a few small inconspicuous bracteoles;
bracts of the involucre 8, oblong or linear-oblong, obtusish,
glabrous, the inner scarious-margined : ray-flowers 2 or 3 ; rays
short, oblong 2 to 2.5 mm. long: disk-flowers about 9: achenes
glabrous. Monogr. Senecio, I Th. 26 (1901) & in Engl. Bot.
Jahrb. xxxii. 22 (1902), without complete characterization.
MEXICO. State of Chiapas: near Chicarras, altitude 1830 m.,
12 to 15 February, 1896, E. W. Nelson, no. 3,796 (hb. Gray, and
hb. U. S. Nat. Mus.). A species similar to 5. arborescens Steetz,
but with different foliage and involucre. It is also related to
S. Gurkei Hieron., a species of New Granada, and to S. grandi-
folius Less., but these again have a very different pubescence
and involucre.
Senecio ( Terminales) copeyensis Greenm.
Arborescent: stem above tomentose: leave petiolate, large,
ovate-oblong in general outline, 2 to 3 dm. long, 1.5 to 2 dm.
broad, pinnately lobed with deep narrow sinuses, abruptly or
somewhat gradually contracted at the base into the petiole,
reticulately veined above and sparingly puberulent over the
upper surface except along the tomentulose midrib, at first
tomentulose beneath later somewhat glabrate; lobes narrowly
oblong to lance-oblong, 2.5 to 10 cm. long, i to 2.5 cm. broad,
terminated by a stoutish cartilaginous mucro, entire or remotely
and inconspicuously cartilaginous-denticulate, revolute-mar-
gined; petioles nearly or quite i dm. long: inflorescence abruptly
terminating the stem in two or more compound many-headed
tomentose corymbose panicles: heads about 8 mm. high, discoid:
involucre calyculate with few setaceous-linear bracteoles, slightly
tomentose to essentially glabrous ; bracts of the involucre usually
8, lanceolate to oblong-lanceolate, 4 to 5 mm. long, brownish
or even blackish in the dried state, the inner with broad scarious
286 FIELD COLUMBIAN MUSEUM BOTANY, VOL. II.
margins: flowers about 10 with a rather slender tube gradually
ampliated above into a deeply 5-lobed limb: achenes glabrous.
Monogr. Senecio, I Th. 26 (1901) & in Engl. Bot. Jahrb. xxxii.
22 (1902), without complete characterization. COSTA RICA.
"Forts du Copey," altitude 1,800 m., February, 1898, Ad.
Tonduz, no. 11,663 (hb- Gray, and hb. Inst. Physico-Geogr. Costa
Rica) .
Senecio ( Terminales) serraquitchensis Greenm.
Suffruticose: stem above tawny-tomentose : leaves petiolate,
oblong-lanceolate to somewhat oblong-obovate, 1.2 to 2.5 dm.
long, 3 to 9 cm. broad, mucronate-acute, slightly sinuate, remotely
cartilaginous-denticulate, narrowed below to the tomentulose
2.5 to 6.5 cm.-long petiole, at first tomentulose on both surfaces
especially on the midrib and lateral nerves beneath, but soon
glabrate and rather strongly reticulate-veined: inflorescence
abruptly terminating the stem in one or more long-pedunculate
compound many-headed corymbs: heads about i cm. high,
radiate: involucre barely calyculate with minute bracteoles;
bracts of the involucre 8, narrowly oblong, about 4 mm. long,
obtuse, turning blackish in drying: ray-flowers 5; ligules short,
4-nerved; tube shorter than the pappus: disk-flowers 5 or 6;
corollas rather deeply 5-lobed: achenes glabrous. Monogr.
Senecio, I Th. 26 (1901) & in Engl. Bot. Jahrb. xxxii. 22 (1902),
without complete characterization. S. Ghiesbreghtii, var. Uspan-
tanensis Coulter in Bot. Gaz. xx. 52 (1895), in part. GUATE-
MALA. Department of Alta Vera Paz: Serraquitche", altitude
760 m., April, 1889, John Donnell Smith, no. 1,598 (hb. Gray).
Senecio ( Terminales) uspantanensis Greenm. Monogr. Se-
necio, I Th. 26 (1901) & in Engl. Bot. Jahrb. xxxii. 22 (1902).
S. Ghiesbreghtii, var. uspantanensis Coulter, Bot. Gaz. xx. 52
(1895), in part, as to Heyde & Lux, no. 3,368 (hb. Gray) and
Botteri, nos. 609, 820 (hb. Gray).
Although 5. uspantanensis is similar in habit to 5. serraquit-
chensis, yet the former differs markedly in its glabrous stem and
foliage, longer and fewer (5 instead of 8) involucral bracts.
TRIXIS PRINGLEI Rob. & Greenm. Proc. Am. Acad. xl. 10 (1904).
Specimens collected by Professor C. Conzatti at De Almoloyas
Sta. Catarina, Oaxaca, Mexico, at an altitude of 1,000 m.,
26 December, 1906, no. 1,646 (hb. Field Mus.) agree well with
specimens secured by Mr. Pringle on which the species was
based, except the leaves in the Conzatti plant have a maximum
breadth of 2.5 centimeters.
Jungia Pringlei Greenman, sp. nov.
Stem terete, pubescent: leaves petiolate, orbicular-ovate,
cordate, 7-9-lobed, hirtellous-puberulent and rather strongly
reticulate-nerved above, crisp-hirsute-pubescent and atomi-
ferous-glandular beneath; lobes ovate-triangular, dentate, mu-
DEC. 1907. SPERMATOPHYTES FROM MEXICO GREENMAN. 287
cronate-acute ; petioles 1.5 to 10 cm. long, estipulate, pubescent:
inflorescence a terminal many-headed open panicle; bracts of
the inflorescence more or less foliaceous, ovate-oblong, lance-
elliptic to linear, dentate to entire: heads about i cm. high,
i8-22-flowered: involucre narrowly campanulate; bracts of the
involucre linear-lanceolate, 8 mm. long, acuminate, acute,
pubescent with jointed hirsute hairs intermixed with a glandular
puberulence: flowers bilabiate; the outer lip of the exterior
circle of flowers ligulate, 3 mm. long, spreading, pale yellow or
whitish: pappus exceeding the involucre, setulose, silvery white:
immature achenes narrowed above, pubescent. MEXICO. State
of Michoacan: Uruapan, collection of 1906, C. G. Pringle, no.
10,357 (hb. Gray; fragment and photograph in hb. Field Mus.)-
