OCCASIONAL PAPERS
OF THE
California Academy of Sciences
No. 88, 65 pages, 30 figures.
NEW SCENOPINIDAE (Diptera) FROM NORTH AMERICA
By
Lewis P. Kelsey
?A R Y
NOV 8 1971
HOLE,
SAN FRANCISCO
PUBLISHED BY THE ACADEMY
October 1, 1971
COMMITTEE ON PUBLICATION
Dr. George E. Lindsay, Chairman
Dr. Edward L. Kessel, Editor Dr. Leo G. Hertlein
OCCASIONAL PAPERS
OF THE
California Academy of Sciences
No. 88, 65 pages, 30 figures.
NEW SCENOPINIDAE (Diptera) FROM NORTH AMERICA
By
Lewis P. Kelsey
Department of Entomology and Applied Ecology
College of Agricultural Science
University of Delaware, Newark, Delaware 19 711
Since the completion of "A Revision of the Scenopinidae
(Diptera) of the World" (Kelsey, 1969)2, the author has con-
tinued to receive additional specimens for identification.
Among these were the 30 species new to science whose descrip-
tions are included in this paper. Represented are members of
four of the five North American genera. These additions bring
to 127 the described North American (including Mexican) spe-
cies. The fact that these comprise nearly half of the known
species of the family is undoubtedly a reflection of more in-
tensive collecting rather than of actual distribution of the
group.
The author is indebted to the following institutions and
persons for furnishing the material for this study:
University of California, Riverside, particularly Michael
Irwin and Dr. Evert I. Schlinger;
Canadian National Collection, Ottawa, J. R. Vockeroth;
United States National Museum, Washington, D. C. , W. W.
Wirth; and
Washington State University, Pullman, Dr. M. T. James.
The keys to the North American genera have been revised
to include all of the new species.
1. Published as Miscellaneous Publication no. 621 with
the approval of the Director of the Delaware Agricultural
Experiment Station. Publication no. 402 of the Department
of Entomology and Applied Ecology.
2. United States National Museum Bulletin, no. 277: 336
pp. , 210 figs.
2 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
In the illustrations of the new species, the scale marks on
the plates equal 1/2 mm. The shorter mark applies to the wing
and head, while the longer relates to the terminalia and en-
larged details of the head and antenna.
GENUS SCENOPINUS Latreille
Kelsey (1969) recognized 37 species in the genus Scenopinus
from North America including Mexico. Of these, seven were of
the Fenestralis Group and 30 of the Velutinus Group. The addi-
tion of the material described here increases the number of
North American species by three for the former group, and by
six for the latter, for a total of 46 Scenopinus species found
in North America.
FENESTRALIS GROUP
As all three of the new species would run to S_. undulaf rons
Kelsey in the keys to the Nearctic species (Kelsey, 1969 p. 16),
the following key has been prepared to include all species.
1. Antennae long and narrow 2
Antennae short, pear-shaped 4
2. (1) Wings hyaline, halter knob white to brown 3
Wings smoky gray, halter knob yellow-brown
S. schroderi (Krober)
3. (2) Frons shining, halter knob white, legs orange-
brown S. glabrif rons Meigen
Frons rugose, halter knob brown to white, legs
red-brown S. fenestralis (Linneaus)
4. (1) Vein R4 branching before the middle of cell R5 . . . 5
Vein R4 branching at the middle of cell R5
S. acquelonius Kelsey
5.(4) Vein R4 branching from near the middle of cell R5 . . 6
Vein R4 branching from the basal fourth of cell
Re S. brevi terminus Kelsey
6. (5) Halter knob red-brown 7
Halter knob of female black, of male dark
red-brown with lighter base
S. vockerothi Kelsey, new species
7. (6) Antennae with last segment red-brown 8
Antennae with last segment black or black-
brown 9
8. (7) Vein R4 branching well beyond the line of the
m crossvein extended S. undulafrons Kelsey
Vein R4 branching from a point on line of m
crossvein extended . . . S. estesi Kelsey, new species
9.(7) Antennae with last segment black; a deep
transverse groove on lower frons
S. transversus Kelsey, new species
Antennae with last segment black-brown;
lower frons smoothly rounded with a shallow
median groove S. nidorcaupulus Kelsey
No. 88] KELSEY: NEW SCENOPINIDAE
Scenopinus estesi Kelsey, new species.
(Figure 1. )
MALE. Head black; eyes red-brown above, black-brown be-
low, extending to the back side of head; frons narrow, a trans-
verse depression with a central and two lateral points that
make it look like a crown on the lower frons over the antennae;
silvery pile surrounding the oral opening extending in a nar-
row band over the antennal bases; mouthparts brown, two- thirds
the length of the oral opening; palpi red-brown, slender,
shorter than mouthparts; ocellar tubercle black, prominent;
ocelli red-brown; antennae dark red-brown, third segment oval,
truncated at the tip, and only twice as long as broad; see fig-
ures for details.
Thorax black-brown dorsally, supraalar calli red-brown,
pleural areas dark red-brown; wings brownish hyaline, veins
brown, R4 branching from before the middle of cell on a line
with the m crossvein extended; halter stem red-brown, knob red-
brown; fore legs with femora dark red-brown, tibiae red-brown,
tarsi orange, darkening distally; middle and hind legs with
femora and tibiae dark red-brown, tarsi orange, darkening dis-
tally, femoro-tibial knees orange.
Abdomen black-brown; white membranous bands on segments 3-
6 very narrow (best seen from posterior aspect) , terminalia
black-brown; much like S_. fenestralis externally but differs in
accessory glands and apodemes internally; see figures for de-
tails .
FEMALE. Unknown.
LENGTH. Male body 4.3 mm. wing 3.2 mm.
TYPE LOCALITY. Stony Creek, Sequoia National Park, Cali-
fornia; 15 June 1964 in wet log, emerged 1 August 1964 (P. M.
Estes) .
TYPE DEPOSITED. California Academy of Sciences no. 10117
(ex University of California, Riverside) .
Scenopinus transversus Kelsey, new species.
(Figure 2 . )
FEMALE. Head black; eyes black-brown; postocular ridge
moderately wide, elevated on the posterior edge; frons moder-
ately broad, nearly twice as wide as the ocellar tubercle,
rugose, with a median groove bordered by transverse striation.
and a transverse depression above the antennal sockets; mouth-
parts brown, well developed; palpi black, half as long as the
oral opening; ocellar tubercle black, not set off; ocelli
brown; antennae black, third segment pilose, pear shaped,
barely twice as long as wide; see figures for details.
Thorax black, rugose with short silvery hairs; humeral and
supraalar calli red-brown; mesoscutellum not distinctly set off
from the notum; pleural areas black-brown; wings smoky brown
hyaline, veins brown, R4 distal to the extension of the m cross
vein extending straight to the edge of the wing; halter stem
black-brown, knob red-brown; legs with femora and tibiae dark
red-brown with orange-brown knees, tarsi orange-brown.
s
CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
Abdomen black, rugose, subshining. See figures for con-
formation of 8th sternum, and 9th sternum and bursa.
MALE. Unknown.
LENGTH. Female body 4.8 mm., wing 3.2 mm.
TYPE LOCALITY. Sagehen Creek near Hobart Mills, Nevada
County, California. 15 July 1964 (M. E. Irwin); Malaise trap
in meadow.
TYPE DEPOSITED. California Academy of Sciences, no. 10119,
(ex University of California, Riverside) .
PARATYPE. 1 female same data except 18 July 1962, (R. M.
Bohart) in University of California, Riverside.
Scenopinus vockerothi Kelsey, new species.
(Figure 3. )
MALE. Head black; eyes red-brown above, black-brown below,
extending to back of head; frons triangular, extending to medi-
an ocellus, upper portion nearly as wide as median ocellus,
roughened, dull but not striated, a very faint midline on lower
frons; ocellar tubercle black; ocelli orange; back of head con-
cave; mouthparts red-brown, filling oral cavity; palpi red-
brown, slender, more than half the length of the oral cavity;
antennae black-brown, third segment oval, not twice as long as
broad; see figure.
Thorax, black, rugose, subshining; humeral and supraalar
calli tipped red-brown; pleural areas black, except below wing
red-brown, rugose; wing hyaline, veins brown, R4 branching be-
fore the middle of cell R5 ; halter stem brown, knob dark red-
brown with lighter base; legs dark red-brown except tarsi
orange-brown with tips and claws red-brown.
Abdomen black-brown with thin white membranous bands behind
segments 3-5, 9th tergum four-lobed; see figure for details.
FEMALE. Head black; eyes black, with moderately broad,
shining postocular rim; frons broad, wider than ocellar tuber-
cle and rugose except shining along the eye margins and median
band, with a slight depression at the middle and two bulges on
the lower frons; ocellar tubercle black; ocelli red-orange;
antennae black, third segment tapering and not twice as long as
broad; see figures for details.
Thorax, black, rugose; humeral and supraalar calli red-
brown; pleura black, rugose; wings brownish, veins brown, R4
undulating to edge of wing from a point only slightly distal to
the line of the m crossvein extended; halter stem brown, knob
black; legs black except femoro-tibial joints orange, tarsi
orange, darkening to red-brown distally.
Abdomen, broad, shining, and black, except central portion
covered with brown pollen.
LENGTH. Male body 3.8 mm., wing 2.8 mm., female body 4.6
mm. , wing 3 . 3 mm.
TYPE LOCALITY. Old Chelsea, Quebec, Canada. Summit of
King Mountain 1150', July, 1964 (J. R. Vockeroth) .
ALLOTYPE. Female, Stittsville, Ontario, Canada, 26 July
1965 (J. R. Vockeroth).
TYPE AND ALLOTYPE DEPOSITED. Canadian National Collection
no. 11428.
No. 88] KELSEY: NEW SCENOPINIDAE
VELUTINUS GROUP
The following revisions of keys to the Nearctic species
(males) (Kelsey 1969, p. 110) will permit the determination of
all known males. The key to the females is completely revised,
Nearctic Species (Males)
14.(12) Halter knob completely or partially brown
to black 14a
Halter knob completely or partially red-brown ... 15
14a (14) Halter knob all brown to black. . S. valgus (D.E.Hardy)
Halter knob brown with ventral tip white
. . . . S_. reciprocatus Kelsey, new species (in part)
25. (24) Halter knob all brown or brown with a lighter
ventral area 26
Halter knob all red-brown, or with cream
ventral surface, or cream with a flush of
red-brown at base 27
26. (25) Halter knob all brown, hind margin of 9th
tergum produced at the midpoint . . . S. cooki Kelsey
Halter knob brown to black-brown above, a
white lateral band and a ventral tip
separating it from a lighter brown ventral
surface, 9th tergum with posterior margin
straight S_. johnsoni Kelsey, new species
27. (25) Halter knob all red-brown 28
Halter knob red-brown above, cream below or
cream with flush of red-brown at base,
9th tergum truncate
. . . . S_. reciprocatus Kelsey, new species (in part)
28. (27) Third antennal segment short .... S. barberi Kelsey
Third antennal segment long and narrow
. S. butleri Kelsey
Nearctic Species (Females)
1. Wings hyaline 2
Wings brown or white 11
2. (1) Halter knob white 3
Halter knob not white 5
3. (2) Palpi dark 4
Palpi orange . . S. aurantipalpus Kelsey, new species
4. (3) Lateral depressions on 8th tergum and
sternum S. beameri (D.E. Hardy)
No lateral depressions on 8th tergum
S_. whittakeri James
5. (2) Halter knob brown or red-brown 6
Halter knob tan or yellow 9
6. (5) Cell R5 tapering to margin of wing 7
Cell R5 sharply constricted before margin
of wing S. gossypius Kelsey
7. (6) Legs with tibiae red-brown 8
Legs with tibiae orange-brown to
brown S. magdalenai Kelsey
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
8. (7) Eighth tergum flat, with lateral
depressions S. beameri f uscus (D.E.Hardy)
Eighth tergum rounded and without
depressions S. valgus (D.E.Hardy)
9. (5) Halter knob tan 10
Halter knob yellow, tip of cell R5 open
S_. mirabilis Adams
10. (9) Antennae red-brown, third segment elongate,
more than twice as long as broad . S_. howdeni Kelsey
Antennae brown, third segment oval and barely
twice as long as broad
S. dietricki Kelsey, new species
11. (1) Wings brown or smoky brown 12
Wings white or smoky white 21
12. (11) Halter knob white to tan 13
Halter knob dark 15
13. (12) Halter knob cream or tan 14
Halter knob white S_. vanduzeei Kelsey
14. (13) Halter knob cream, antennae long,
straight S. kuiterti (D.E.Hardy)
Halter knob tan, antennae short,
pear-shaped S. cochisei Kelsey
15. (12) Cell R5 narrowed at margin of wing 16
Cell R5 open at margin of wing 19
16. (15) Halter knob red-brown to black-brown,
marked or lighter below 17
Halter knob red-brown to black-brown,
polished S. nubilipes Say
17. (16) Halter knob brown or black-brown 18
Halter knob red-brown above, white
below S. hagai Kelsey
18. (17) Halter knob black-brown above, with a white
lateral band and ventral tip, remainder of lower
surface red-brown . . S. johnsoni Kelsey, new species
Halter knob brown above, ventral tip of knob
light . S. reciprocatus Kelsey, new species (in part)
19. (15) Halter knob entirely brown or red-brown 20
Halter knob red-brown and velvety above,
tan below S. foxi Kelsey, new species
20. (19) Halter knob light red-brown, tarsi light
red-brown S. bryanti Kelsey
Halter knob brown, tarsi straw
yellow S. flandersi Kelsey
21. (11) Halter knob partially or entirely white 22
Halter knob partially or entirely yellow to brown . 26
22. (21) Halter knob all white 23
Halter knob white with base tan ... S. weemsi Kelsey
23. (22) Body without flattened scales 24
Body with flattened scales
S. albif asciatus (D. E. Hardy)
24. (23) Third antennal segment not parallel-sided 25
Third antennal segment uniform in
width s. chico Kelsey
25.(24) Third antennal segment oval, widest at
the middle S. alalacteus Kelsey
Third antennal segment pear-shaped, widest
at base S. cretatus Kelsey
26. (21) Halter knob brown or red-brown 2 8
Halter knob lighter 27
No. 88] KELSEY: NEW SCENOPINIDAE
27. (26) Halter knob orange-yellow, thorax without
white hairs S. seftoni Kelsey
Halter knob dark tan to light orange-brown,
thorax covered with short white hair
S_. albipilosus Kelsey, new species
28. (26) Halter knob entirely red-brown 29
Halter knob brown or red-brown above,
lighter below 30
29.(28) Antennae black-brown, elongate oval; frons
swollen above antennal bases and with a broad
median depression on lower fourth . . S. rossi Kelsey
Antennae red-brown, short; frons swollen above
antennal bases, a median groove dividing
lower two-fifths S. barberi Kelsey
30. (28) Halter knob brown above, white below . S. pecki Kelsey
Halter knob red-brown above, ventral portion
cream diffused with brown
• • • • S. reciprocatus Kelsey, new species (in part)
Scenopinus albipilosus Kelsey, new species.
(Figure 4. )
FEMALE. Head black; eyes red-brown; postocular ridge very
narrow; frons rugose, swollen above antennae, broader than the
ocellar tubercle, with a subshining median band and with short
white hairs on the lateral portions; ocellar tubercle black,
not distinctly set off; ocelli clear; mouthparts well developed,
nearly filling the oral cavity; labellum brown; palpi large,
paddle-shaped, clothed with short white hairs laterally; oral
cavity bordered by white pubescence which extends over the an-
tennal bases, genae below eyes shining; antennae with first two
segments red-brown, granular; third segment lost on the speci-
men; see figures for details.
Thorax black, rugose, covered with sparse but noticeable
short white hairs; humeral and supraalar calli orange; pleural
areas black-brown with white hairs; wings milky white, opaque,
with brown veins; halter stem brown, knob dark tan to light
orange-brown; legs with coxae to tibiae red-brown and covered
with short white hairs; tarsi and knees light orange-brown.
Abdomen, dark red-brown with short white hairs that are
particularly noticeable at posterior end. See figures for
details of 8th sternum, and 9th sternum and bursal cavity.
MALE. Unknown.
LENGTH. Female body 3.7 mm., wing 2.6 mm.
TYPE LOCALITY. El Centro, Imperial County, California,
10 June 1954, (R.A. Flock).
TYPE DEPOSITED. California Academy of Sciences, no. 10114,
(ex University of California, Riverside) .
Scenopinus aurantipalpus Kelsey, new species.
(Figure 5 . )
FEMALE. Head black-brown; eyes red-brown with a narrow
8 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
postocular ridge; frons broad, granular with a flattened median
depression; ocellar tubercle black-brown, ocelli orange-brown;
oral margin behind and below the antennae polinose , mouthparts
red-brown, palpi with base red-brown, distal portion large,
fleshy, orange colored, easily visible; antennae with two basal
segments black-brown, third segment elongate tapering, orange-
brown basally, black-brown distally, and half as long as the
head is high; see figures for details.
Thorax black-brown, dorsum rugose with sparse hairs; humer-
al and supraalar calli red-brown; wings hyaline with tan-brown
veins; halter stem brown, knob white; legs red-brown, lighten-
ing distally.
Abdomen red-brown subshining. See figures for details of
8th sternum, 9th sternum, and bursal cavity.
MALE. Unknown.
LENGTH. Female body 2.7-3.2 mm., wing 2.0 mm.
TYPE LOCALITY. Deep canyon, Riverside, California, 9 Au-
gust 1963 (E. I. Schlinger) .
TYPE DEPOSITED. California Academy of Sciences, no. 10115
(ex University of California, Riverside).
Scenopinus dietricki Kelsey, new species.
(Figure 6 . )
FEMALE. Head black; eyes black-brown with a narrow post-
ocular ridge; frons rugose, with a central elevated ridge that
has a shallow depressed midline and with a few hairs; gray pu-
bescence extending from the border of the oral cavity to above
the antennae; genae subshining; ocellar tubercle black, not
distinctly set off; ocelli orange; back of head concave, under-
cutting the ocellar tubercle; mouthparts brown, well developed,
and extending to end of oral cavity; palpi brown, only a little
more than half as long as the mouthparts; antennae brown, first
two segments short, third oval and only a little more than
twice as long as broad; see figures for details.
Thorax black, rugose; humeral and supraalar calli tan;
mesoscutellum black in anterior half, red-brown in posterior
half, and with a transverse median depression; pleural areas
red-brown; wings hyaline with brown microtrichia and light
brown veins; halter stem brown, knob tan above and lighter be-
low; legs with femora and tibiae red-brown with yellow knees;
tarsi yellow, darkening at tips.
Abdomen, dark red-brown, 8th segment with sparse long
hairs. See figures for conformation of 8th and 9th sterna and
bursal cavity.
MALE. Unknown.
LENGTH. Female body 3.5 mm., wing 2.2 mm.
TYPE LOCALITY. Indio, Riverside County, California, 20
July 1962. (E. J. Dietrich) on tomato.
TYPE DEPOSITED. California Academy of Sciences, no. 10116
(ex University of California, Riverside) .
No. 88] KELSEY: NEW SCENOPINIDAE 9
Scenopinus foxi Kelsey, new species.
(Figure 7. )
FEMALE. Head dark red-brown; eyes brown with golden glints
and a well developed postocular ridge; eye margins separated
above from the ocellar tubercle by a space equal to width of
the tubercle; frons broad, rugose, with prominent swellings
above the antennal bases and deeply cleft by a groove that
ends in a shallow pit on the lower third of the frons; ocellar
tubercle not prominent, ocelli orange; mouthparts brown, well
developed, filling two thirds of oral opening; palpi subequal
to mouthparts; antennae brown, the first two segments short,
the third oval, with a pile of hairs and about twice as long
as wide; see figures for details.
Thorax red-brown; humeral callus with tip orange; supra-
alar callus orange; mesoscutellum lighter in color than scutum;
wings a faint smoky brown, the veins light brown; halter stem
brown, knob red-brown and velvety above, tan below; legs red-
brown with lighter colored tarsi, the hind tarsi straw yellow.
Abdomen red-brown, its 8th segment dark red-brown. See
figures for details of 8th and 9th sterna and bursal cavity.
MALE . Unknown .
LENGTH. Female body 2.7 mm., wing 2.1 mm.
TYPE LOCALITY. Hollister, Idaho, 13 June 1931 (D. E. Fox),
Plot 6 outside.
TYPE DEPOSITED. United States National Museum, no. 70816.
Scenopinus johnsoni Kelsey, new species.
(Figure 8. )
This species is represented by a series of 65 specimens,
nearly equally divided between males and females, taken from
the nests of Ectemnius spiniferus (Fox) (Hymenoptera, Crabron-
idae) in cotton stems, pointing up again the fact that sceno-
pinids can be taken in abundance at the proper place and time.
MALE. Head black; eyes red-brown above, black-brown be-
low, and extending around to the back side of the head; frons
narrow, triangular, extending half the distance from the an-
tennae to the median ocellus, rugose with a central depression
on the upper fourth; ocellar tubercle black-brown, subshining;
ocelli brown; back of head concave; genae small, shining;
mouthparts brown, well developed; palpi black-brown, slender,
half as long as oral opening; oral opening bordered by silvery
pile that extends dorsally above the bases of the antennae;
antennae black-brown, first two segments short, third segment
oval, twice as long as wide, pilose/ and with a blunt tip. See
figures for details.
Thorax black-brown, a patch of silvery pile above the
humeral callus, and a few sparse hairs over the remainder;
humeral callus with a small red-brown tip, supraalar callus
tan; pleurum dark red-brown; wings milky white, the veins tan
becoming fainter behind; halter stem brown, knob brown to
black-brown above, with a white lateral band and ventral tip;
remainder of lower surface a lighter brown, legs with coxae
through tibiae dark black-brown, tarsi yellow-brown, darkening
10 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
to brown dis tally and posteriorly.
Abdomen black-brown, broad, segments 2 to 4 with narrow
white membranous posterior borders; aedeagal parameres weak.
See figures for details of 9th tergum and ejaculatory pump.
FEMALE. Head black-brown; eyes red-brown, with a moder-
ately wide postocular ridge which is glabrous near the eye and
rugose distally; frons rugose, swollen, with a distinct median
groove that fades on the upper fourth of the frons; back of
head concave; mouthparts as in the male; ocellar tubercle
black-brown; ocelli clear to red-brown; antennae as in the
male. See figures for details.
Thorax as in the male, except humeral callus with orange
tip; wings brownish hyaline, the veins brown, darker and
stronger than in the male; halter stem brown, knob black-brown
above with a white lateral band and ventral rip, remainder of
the lower surface red-brown; legs as in the male.
Abdomen black-brown, shining, 8th segment black-brown,
sternum excavated on midline. See figures for conformation of
8th and 9th sterna and bursal cavity.
LENGTH. Male body 2.6-3 mm., wing 2 mm. Female body 3.4-
3.8 mm., wing 2.3 mm.
TYPE LOCALITY. 3 miles North of Madera, Madera County,
California, 30 September 1965, (C. Johnson) from cells of
Ectemnius spiniferus (Fox) in cotton stems.
ALLOTYPE. Female: same data as male.
TYPE DEPOSITED. California Academy of Sciences, no. 10118
(ex University of California, Riverside).
PARATYPES. 31 males and 32 females deposited as follows:
20 males, 21 females in University of California, Riverside;
2 males, 2 females in United States National Museum; 2 males, 2
females in Snow Museum, Lawrence, Kansas; 4 males, 4 females in
University of Delaware, 1 male, 1 female in University of Cali-
fornia, Davis, 1 male, 1 female in University of California,
Berkeley.
Scenopinus reciprocatus Kelsey, new species.
(Figure 9 . )
This species is distinguished by cell R5 being sharply con-
stricted near its apex. The shape of this cell is not as bot-
tle-like as in S_. gossypius , yet the constriction is more pro-
nounced than in S. f landersi . The truncate posterior margin of
the 9th tergum of the male, though similar to those of the
above mentioned species, differs from the former in the short
aedeagal parameres, larger ejaculatory mechanism and accessory
glands, and from the latter in the more pointed claspers on the
9th segment. Though readily recognized as being distinct from
S. gossypius and S. f landersi , this species has been difficult
to classify- owing to individual variability in such characters
as halter color and form of the accessory gland of the ejacula-
tory pump. When the latter structure is viewed from the side
as in the illustrations, some individuals have the proximal
accessory gland anterior to the distal and the positions are
reversed in others. This suggested the specific name.
It is my feeling, based on the geographic range of the
specimens at hand, that speciation is taking place and that
No. 88] KELSEY: NEW SCENOPINIDAE 11
some time in the future several closely related but distinct
species may evolve.
MALE. Head black; eyes dark red-brown above, black-brown
below, extending over the back of the head and nearly touching
for half the distance from the median ocellus to the antennal
bases; frons narrow, triangular, with a median depression,
striated below and smooth above; ocellar tubercle black-brown,
shining; ocelli red-brown; back of the head concave; oral cavi-
ty bordered by a narrow band of white pubescence which extends
dorsally as two bands to meet on the midline above the antennal
bases; mouthparts brown, well developed; palpi black-brown,
only half as long as mouthparts; antennae black-brown, the
first two segments short, third segment oval and only twice as
long as wide.
Thoracic dorsum black-brown, striated, with a silvery patch
above the humeral callus; the transverse suture deeply grooved;
humeral callus black-brown, sometimes with an orange tip,
supraalar callus orange-brown; pleural areas black-brown; wings
brownish hyaline in the northern part of the range to milky in
the southern part, the veins brown to tan depending on wing
color; halter stem brown, knob brown in northern range to red-
brown southward, the ventral tip white in the northern speci-
mens to the ventral half tan to cream southward, or the knob
cream with a flush of brown over the basal half in the most
south-westerly portion of the range; legs black-brown with
brown tarsi, changing to red-brown with tan tarsi in the south-
ern part of the range.
Abdomen black-brown to red-brown, rugose; segments 3 to 5
with broad white posterior margins, the anterior portion of the
segment sclerotized, shining; terminalia as figured, 9th tergum
truncate, aedeagal parameres short, 9th segment claspers
pointed; internal ejaculatory organs nearly half again as long
as 9th tergum. See figures for details.
FEMALE. Head black; eyes red-brown with a moderately wide,
rounded, postocular rim; frons broad, narrowest just above the
antennae, broadening dorsally, only slightly swollen on the
lower portion, a shallow median depression with radiating stri-
ae on lower third; ocellar tubercle not prominent or set off
from the frons, anterior ocellus sunken; ocelli red-brown;
mouthparts and palpi as in the male; silvery pile bordering
oral cavity extending to laterad of antennal bases; antennae
as in the male.
Thorax as in the male except that light areas on ventral
portion of halter knobs are diffused with red-brown.
Abdomen dark red-brown to red-brown; 8th sternum broadly
truncate with median posterior margin gently convex; 9th ster-
num with a membranous margin and without prominent hairs; roof
of bursa rectangular, membrane on ventrally projecting face.
See figures for details.
LENGTH. Male body 2.7-3.0 mm., wing 1.9-2.3 mm.; female
body 2.7-3.1 mm., wing 2.2-2.4 mm.
TYPE LOCALITY. Craters of the Moon National Monument,
Idaho; 13 July 1965 (D. S. Horning) on Melalotus officinalis.
HOLOTYPE. Male, United States National Museum, no. 70818.
ALLOTYPE. Female, same data.
PARATYPES. 1 male and 1 female, same data as type except
male 5 July 19 65; 1 male, Las Cruces, New Mexico, 2 7 June
12 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
1961, (J. C. Watts) on alfalfa, 1 male, Phoenix, Arizona, 27
May 1938, (Christenson) , 1 male, Lindsay, California, 15 July,
(H. E. L. ) , all in United States National Museum; 1 male,
Riffle, Colorado, 6 July 1901 (C. W. Johnson) in the Museum of
Comparative Zoology, Harvard University; 3 males, Bishop, Cal-
ifornia, 28 July 1940, (D. E. Hardy), 1 male, Arivaca, Arizona,
26 July 1941, (R. H. Beamer) ; 1 female, Las Cruces , New Mexico,
3 July 1940, (R. H. Beamer) , 1 female, Sabine, Arizona, 7 July
1950, (L. D. Beamer), all in Snow Museum; 1 female, Riverside,
California, 1 October 1950, (Timberlake) , 1 female, Sierra
Madre Insectary, 20 October 1923, (Armitage) in Timberlake Col-
lection; 1 female, Harrison Hot Springs, British Columbia, 16
August 1916, (C. G. Hewitt), in Canadian National Collection;
2 males, Patagonia, Arizona, 27 June 1961, (P. H. Johnson)
ultra-violet light trap, 1 male, Continental, Arizona, 22 March
1956 (F. Werner, G. Butler) in swept alfalfa, 1 male, 8 miles
North of Vail, Pima County, Arizona, 30 August 1962, (F. Wer-
ner, W. L. Nutting), ultra-violet light trap, 1 male, Litch-
field Park, Arizona, 7 August 1956, (C. Williams) in swept
alfalfa; 1 male, Cortaro , Arizona, 7 August 1957, (G. D. But-
ler) in swept mesquite , 1 female, Uma, Arizona, 18 June 1953,
(Haga) on cotton, all in the University of Arizona; 1 male,
Hat Creek P. O. , Shasta County, California, 2 July 1965, (E.E.
Lindquist) , 1 male and 1 female, in copula 2 miles Northeast
Rodeo, Hidalgo County, New Mexico, 24 July 1963, (A. Raske) ,
in the University of California, (Berkeley) .
GENUS BREVITRICHIA D. E. Hardy
The genus Brevitrichia containing 27 species is distrib-
uted west of the Rocky Mountains from Idaho and Washington to
the north to Guatemala on the south. With the addition of the
15 new species described herein, the number of known species
is raised to 42. The addition of so many new species has made
it necessary to prepare completely revised keys to the genus
Brevitrichia to be used in place of those found in Kelsey
(1969) pp. 65-68.
Key To The Species Of Brevitrichia (Males)
1. Wings hyaline 2
Wings smoky white or brown 9
2. (1) Halter knob tan 5
Halter knob cream or cream-yellow 3
3. (2) Halter knob cream, legs varicolored 4
Halter knob cream-yellow, legs dark red-
brown with hind tibiae yellow-brown
B. wilcoxi Kelsey, new species
4. (3) Legs with femora red-brown dusted with gray
pollen, tibiae orange-brown, tarsi orange-brown
darkening distally. . . B. arena Kelsey, new species
Legs with femora light brown with tan knees,
tibiae tan, tarsi tan, but darkening
distally B. arnaudi Kelsey
No. 88!
KELSEY
NEW SCENOPINIDAE
13
5.
6.
7.
8.
(2
(5
(e;
(7
9. (n:
10. (9
11. (10
12. (9
13. (12
14. (13i
15. (13;
16. (15;
17. (12
18. (17
19. (18;
20. (19
21. (20
Halter knob
Halter knob
Halter knob
upper and
Abdomen with black-brown or red-brown dorsal
markings 6
Abdomen white dorsally B. oculivirida Kelsey
Abdomen with plain bands of color 7
Abdomen with red-brown "T's" on first two
segments/ remainder cream dorsally
B. badiclitella Kelsey, new species
Bands dark-brown or red-brown with cream or
tan membranes 8
Bands black-brown with white membranes B. halli Kelsey
Bands on abdomen red-brown with tan margins;
antennae black-brown with tuft of white between
tips B. albanota Kelsey, new species
Bands on abdomen dark-brown with light posterior
margins; antennae reddish-brown with two terminal
spines, but no tuft of white between. B.ordwayi Kelsey
Wings smoky-brown 10
Wings smoky-white 12
Halter knob tan and cream 11
tan B. hodgeni Kelsey
tan above, cream below . B. grif f ini Kelsey
tan with a creamy band separating
lower halves
B. argentaurba Kelsey, new species
Halter knob white 13
Halter knob cream or tan dorsally 17
Abdominal segments red-brown or black with
white posterior membranes 14
Abdomen pristine white or white with basal
segment red-brown 15
Abdominal segments black with white posterior
margins B. insulana Cole
Abdominal segments red-brown with white posterior
margins B. inferacanna Kelsey, new species
Abdomen pristine white above 16
Abdomen with second segment with a red-brown base
and a thick central stem, remainder of segment and
abdomen white . . B. forf icicruxa Kelsey, new species
R4 branching from beyond r-m crossvein equal
to length of r-m B. addacifons Kelsey
R4 branching from beyond r-m crossvein twice
the length of r-m B. castanea Kelsey
Halter knob tan 18
Halter knob cream 2 3
Abdomen predominantly white or marked with
pigmented "T's" 19
Abdominal segments red-brown with white posterior
margins, halter knob cream below. B.yucantani Kelsey
Abdomen marked with "T's" ~" 20
Abdomen without "T's", base of second segment
red-brown, remainder white . . . . B. pruinosa Kelsey
Halter knob all tan 21
Halter knob tan above and below separated by a
lateral band of white . B. nevada Kelsey, new species
Abdomen marked with black-brown "T" on second
segment, remainder essentially white 22
14 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
Abdomen marked with red-brown "T's", with creamy
lateral areas, over most abdominal segments ....
B. griseola Coquillett
22. (21) Antennae with third segment light tan
B. piscifonta Kelsey, new species
Antennae with third segment orange-brown
basally to red-brown distally
B. miraloma Kelsey, new species
23. (17) Halter knob all cream . 7 . . 24
Halter knob cream dorsally, white or yellow
below or with tan band 29
24. (23) Abdomen white beyond large basal segment 25
Abdomen banded with some color beyond large
basal segment 26
25. (22) Basal segment of abdomen with anterior half
red-brown continuing as a narrow longitudinal
band to posterior margin in the form of a "T" ,
lateral areas and rest of abdomen white
B. boharti Kelsey
Basal segment with anterior portion red-brown
with a median wedge-shaped extension with a
narrow stem meeting a broad lateral base
giving a "wineglass" appearance, lateral
areas and remainder of abdomen white
B. daileyi Kelsey, new species
26. (24) Abdomen with a "T" on basal segment, narrow
red-brown basal bands on some other seg-
ments; 9th tergum rounded at tip 27
Abdominal segments all with red-brown bands
with cream posterior membranes; 9th tergum
pointed B. beameri Kelsey
27.(26) Abdomen with "T's" on basal segment only,
bands on 3rd or 4th and remainder white
excepting expanded 9th tergites 28
Abdominal segments with median red-brown "T's"
decreasing in size toward tip of abdomen,
9th tergum white with brown base
B. coquilletti Kelsey
28. (27) 9th tergite white with a brown basal spot
B. downeyi Kelsey
9th tergite yellow with a tan basal spot
B. davisi Kelsey
29. (23) Halter knob white or yellow below 30
Halter knob with tan marginal band
B. timberlakei Kelsey
30. (29) Halter knob yellow below 31
Halter knob white below B. minuta Kelsey
31. (30) Abdominal segments yellow with black
basal bands B. helenae James
Abdominal segments brown with creamy
membranous margins B. schlingeri Kelsey
1.
2.
(1)
3.
(2)
4.
(3)
No. 88 J KELSEY: NEW SCENOPINIDAE 15
Key To The Species of Brevitrichia (Females)
Wings hyaline 2
Wings smoky-brown or white 12
Halter knob tan, cream, or orange 3
Halter knob white, 7 spines on 9th
tergum B. f locki Kelsey
Halter knob all or in part cream 4
Halter knob all or in part tan or orange 10
Halter knob all cream 5
Halter knob cream above, tan below; 9th
tergum with 7 spines B. downeyi Kelsey
5. (4) Abdomen brown or brown-red, orange, or
orange-brown 6
Abdomen brown grading to tan posteriorly 9
6. (5) Basal abdominal segments brown or red-brown .... 7
Basal abdominal segments orange or orange-brown . . 8
7. (6) Basal abdominal segments brown with white
posterior bands, 9th tergum with 7
spines B. arnaudi Kelsey
Basal two abdominal segments red-brown,
granular on mid-portion, hind margins
and lateral areas light orange, fourth
segment red-brown basally, but distal
portion and remainder of abdomen dark
orange; 9th tergum with 7 orange spines
B. septispina Kelsey, new species
8. (6) Abdominal segments 2-4 orange-brown with
cream posterior margins, frosted; remainder
red-brown shining; 9th tergum with s spines ....
B. arena Kelsey, new species
Basal abdominal segment orange with a median
granular brown spot, other segments orange;
9th tergum with 7 spines and a tuft of hair . . .
B. dicksoni Kelsey
9. (5) 9th tergum with 8 spines and a dorsal pit
B. boharti Kelsey
9th tergum with 6 spines .... B. schlingeri Kelsey
10. (3) Halter knob all or in part tan 11
Halter knob orange; 9th tergum with 8
spines B. davisi Kelsey
11. (10) Halter knob tan above, cream below; 9th
tergum with 8 spines and a ventral pit
B. beameri Kelsey
Halter knob entirely tan; 9th tergum with
9 short stiff spines B. halli Kelsey
12. (1) Wings smoky brown 13
Wings smoky white 20
13. (12) Halter knob cream, cream-yellow or yellow 14
Halter knob entirely tan, or tan above and
lighter below 16
14. (13) Halter knob cream yellow or yellow; abdomen
without "T's" 15
Halter knob cream; abdomen with red-brown
"T's", 9th tergum with 6 spines and much
hair B. coquilletti Kelsey
16 CALIFORNIA ACADEMY OF SCIENCES . [Occ. Papers
15.(14) Halter knob cream-yellow; abdomen black-
brown basally, lightening to orange-
brown distally, all dusted with gray
pollen; 9th tergum with 6 spines; post-
ocular ridge divided on midline
B. palmacana Kelsey, new species
Halter knob yellow; lighter below; abdomen with
basal segments red-brown with yellow posterior
margins, remaining segments red-brown with
orange lateral and posterior margins; 9th ter-
gum with 6 black spines; hair on 8th and 9th
segments directed anteriorly; postocular ridge
continuous behind ocellar tubercle
B. irwini Kelsey, new species
16. (13) Halter knob cream or white below 17
Halter knob all tan; abdomen dark red-brown;
9th tergum with 1 dorsal spine and 7
pits B. hodgeni Kelsey
17. (16) Halter knob cream below 18
Halter knob white below; 9th tergum with
7 spines B. grif f ini Kelsey
18. (17) Abdomen red-brown with white or cream 19
Abdomen brown; 9th tergum with 8 spines
B. melanderi Kelsey
19.(18) Abdomen red-brown, with white lateral
patches, grading to all red-brown, 9th
tergum with 4 spines B. yucatani Kelsey
Abdomen red-brown with prominent paired
dorsolateral longitudinal cream bands on
segments 2-7; 9th tergum with 8 spines
B. nayariti Kelsey, new species
20. (12) Halter knob white 21
Halter knob yellow-brown, cream, or tan 22
21. (20) 9th tergum with 9 spines and a ventral
tuft of hairs B. insulana Cole
9th tergum with 7 spines and sparse
hair B. palida Kelsey, new species
22. (20) Halter knob cream or tan 23
Halter knob yellow-brown; 9th tergum without
spines but with 8 pits B. aspinosa Kelsey
23. (22) Halter knob tan 24
Halter knob wholly or partially cream 26
24. (23) Halter knob all tan 25
Halter knob tan divided by a white band;
abdomen red-brown dusted with gray pollen;
8th segment orange, 9th tergum with 8
orange spines B. nevada Kelsey, new species
25. (24) Abdomen red-brown; 9th tergum with 5 spines
and much anteriorly directed hair . B. ordwayi Kelsey
Abdomen red-brown; 9th tergum with 8 light
orange spines. The dorsal 4 shorter than
the ventral 4 . . . B. piscifonta Kelsey, new species
26. (23) Halter knob entirely cream 28
Halter knob cream with tan lateral bands 2 7
No. 88] KELSEY: NEW SCENOPINIDAE 17
27. (26) Abdomen brown with white lateral patches;
9th tergum with hair only, no pits
B. addaci fons Kelsey
Abdomen with red-brown "T's" becoming
yellow-brown apically, 9th tergum
with 6 spines B. timberlakei Kelsey
28. (26) Abdomen brown grading to tan apically;
9th tergum with a fringe of hairs only
B. minuta Kelsey
Abdomen cream with red-brown central "T's"
grading to orange apically; 9th tergum
with 7 spines and very short hair . . B. kerni Kelsey
Abdomen black with white posterior
segmental bands; 9th tergum with 8
spines B. griseola Coquillett
Abdomen red-brown with white posterior
segmental bands; 9th tergum with 6
spines B. scitulacea Kelsey
Brevitrichia albanota Kelsey, new species.
(Figure 10. )
MALE. Head creamy white; eyes red-brown with a metallic
green sheen, darker below; frons narrow, slightly swollen, and
with an inverted V-shaped black-brown cleft above the anten-
nae; ocellar tubercle black-brown, pollinose; ocelli red-brown;
mouthparts well developed, but filling only half of the oral
cavity; palpi black-brown, nearly as long as rostrum; antennae
black-brown, second segment short with tan distal membrane,
third segment pear-shaped, scaled, and with a tuft of white in
cleft tip. It is from the latter character that the species
derives its name; see figures for details.
Thorax dorsum black-brown, dusted with gray pollen and with
two olivaceous mediolateral bands; lateral portions of notum
black-brown to olivaceous gray depending on how light strikes;
humeral callus cream at tip; scutellum black-brown, dusted with
gray pollen; pleural areas black-brown, dusted with gray pol-
len; wings hyaline with dark brown veins; halter stem brown;
knob light tan above and cream below; legs black-brown and pol-
linose, with knees of femora tan.
Abdomen granular, red-brown, with thin, tan, membranous
margins; terminalia granular, red-brown basally and tan dis-
tally, with sparse hair; aedeagal parameres long; see figures
for details.
FEMALE. Unknown.
LENGTH. Male body 2.9 mm., wing 2.0 mm.
TYPE LOCALITY. Six miles SE. of Parker, Arizona, 13 April
1965, (M. E. Erwin) (Sand dune association).
TYPE DEPOSITED. California Academy of Sciences, no. 10103
(ex University of California, Riverside).
18 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
Brevitrichia arena, Kelsey, new species.
(Figure 11. )
MALE. Head light brown; eyes extending to back of the
head, red-brown with a green metallic sheen and darker below;
frons very narrow, triangular, and with a darker central tri-
angle above the antenna which causes frons to have a V-shaped
appearance; ocellar tubercle black-brown, dusted with tan pol-
len; ocelli red-brown; oral cavity deeply sunken; mouthparts
with rostrum black-brown, labellum brown, palpi short and brown;
antennae with first two segments black-brown, third segment red-
brown, pear-shaped and with two distal points; see figures for
details .
Thorax dorsum black-brown dusted with bands of gray, olive,
and black-brown pollen in patterns that vary with position from
which viewed; lower edges of dorsum cream yellow; humeral and
supraalar calli cream yellow; scutellum largely black-brown
with cream yellow lateral areas, all dusted with gray pollen;
propleuron cream yellow; mesanepisternum largely red-brown and
dusted with gray pollen; mesokatepisternum red-brown with dor-
sal area cream and dusted with gray pollen; wings clear with
tan to opaque veins; halter stem brown, knob cream yellow; legs
with femora red-brown and dusted with gray pollen, tibiae
orange-brown, tarsi orange-brown, darkening distally.
Abdomen red-brown with cream posterior membranes; segments
2 to 5 dusted with gray pollen; 9th tergum narrow and with
black-brown base, remainder frosted whitish; see figures for
details.
FEMALE. Head orange-brown; eyes red-brown with a metallic
green sheen; postocular ridges moderately broad, extending
uninterruptedly behind the ocellar tubercle; a dark brown patch
behind the dorsal angle of each eye; frons broad, frosted, with
scattered light hairs and cut off from the postocular ridges by
grooves that extend from the upper corner of each eye to a
lateral ocellus; ocellar tubercle black-brown with olive pollen;
ocelli red-brown; oral cavity sunken; mouthparts with rostrum
red-brown, labellum brown, palpi short, orange-brown; antennae
red-brown, third segment pear-shaped, ending in two points with
a median peg and covered with short hairs with golden sheen;
see figures for details.
Thorax dorsum orange-brown with anteromesal and postero-
lateral dark black-brown bands of pollen, but yellow above hu-
meral callus and on sides back to scutellum; humeral and supra-
alar calli cream; scutellum with a narrow median band of red-
brown bordered by orange and with yellow margins; propleuron
mesanepisternum yellow with a ventral red-brown spot; mesakate-
pisternum red-brown with a broad dorsal yellow band and dusted
with gray pollen; wings hyaline with brown veins; halter stem
brown, knob cream-yellow; legs frosted orange-brown.
Abdominal segments 2 to 4 red-brown with cream posterior
margins and frosted with gray pollen; segments 5 to 7 red-
brown, shining; 8th segment orange-brown; 9th tergum with six
black spines and dark hairs; see figures for details.
LENGTH. Male body 2.6 mm., wing 1.8 mm; female body 4 mm.,
wing 2.0 mm.
TYPE LOCALITY. One mile W. of Glamis , Imperial County,
California, 1 October 1965, (M. E. Irwin) (Sand dune
No. 88] KELSEY: NEW SCENOPINIDAE 19
association) .
TYPE. Male and allotype female on same pin deposited in
California Academy of Sciences, no. 10104 (ex University of
California, Riverside).
Brevitrichia argentaurba Kelsey, new species.
(Figure 12 . )
MALE. Head tan; eyes dark red-brown above, with a green
metallic sheen, lower facets black-brown with a violet sheen
and occupying most of the head in side view; frons narrow tri-
angular, slightly swollen; genae cream; back of head brown;
ocellar tubercle red-brown, dusted with olive pollen; ocelli
red; oral cavity large, sunken, mouthparts filling two-thirds
of the oral opening, rostrum red-brown, labellum creamy, palpi
brown, half as long as mouthparts; antenna dark red-brown,
first two segments short, third pear-shaped and ending in two
points without a central peg; see figures for details.
Thorax black dusted with bands of brown and tan pollen
which change pattern when viewed from different angles; lower
half of humeral callus cream; mesoscutellum black-brown, dusted
with brown pollen, but lower posterior edges tan; propleura
tan; mesopleurum generally black-brown and dusted with brown
pollen, very little light area; wing brownish hyaline with
brown veins; halter stem brown, knob tan with a creamy band
separating upper and lower halves; legs black-brown, dusted
with brown pollen, but knees of femora tan.
Abdominal segments brown and dusted with brown pollen,
except intersegmental membranes narrowly whitish, 9th tergum
with base brown, dusted with brown pollen, remainder creamy;
10th segment distinctly visible and covered with long hairs;
aedeagal bristle short; see figures for details.
FExMALE. Unknown.
LENGTH. Male body 3 mm. , wings 2 mm.
TYPE LOCALITY. One mile S. of Sliver City, Grant County,
New Mexico, 1 June 1965, (M. E. Irwin).
TYPE DEPOSITED. California Academy of Sciences, no. 10105,
(ex University of California, Riverside).
Brevitrichia badiclitella Kelsey, new species.
(Figure 13. )
This species is named after the red-brown saddle-like
markings on the dorsum of the thorax.
MALE. Head tan; eyes a metallic blue-green with large
upper and finer lower facets; frons narrow, triangular, swol-
len, with a median cleft and darker dorsally; ocellar tubercle
black-brown with a dusting of brown pollen; ocelli black;
mouthparts well developed, with rostrum red-brown, labellum
tan; palpi tan; antennae brown, first segment nearly twice as
long as second; third antennal segment long, pear-shaped, cleft
at tip and pubescent; see figures for details.
20 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
Thorax dorsum with a solid red-brown, saddle-like area sur-
rounded by areas of yellow above the humeral callus, lateral
margins, and on either side of the midline of the posterior
scutum; all covered with a fine dusting of gray pollen; humeral
and supraalar calli cream; scutellum with a large red-brown
median spot and bordered by yellow and cream laterally; pro-
pleuron cream; mesanepisternum cream with a red-brown spot;
mesokatepisternum red-brown with a cream dorsal margin; wings
hyaline with brown veins; halter stem brown, knob tan with a
brown dividing line; legs red-brown except femora with white
knees .
Abdomen with second segment with a red-brown "T" which has
a broad base and a stem that does not attain posterior margin,
rest of tergum cream, except lateral margins red-brown; third
segment with smaller red-brown "T", remainder of abdomen cream
dorsally; 9th segment tan with a dark basal spot and covered
with long hairs; aedeagal spines long; see figures for details.
FEMALE. Unknown.
LENGTH. Male body 2.7 mm., wings 1.8 mm.
TYPE LOCALITY. Ternecuia, Riverside County, California, 25
June 1963 (E. I. Schlinger) .
HOLOTYPE DEPOSITED. California Academy of Sciences, no.
10106 (ex University of California, Riverside) .
Brevitrichia daileyi Kelsey, new species.
(Figure 14.)
MALE. Head cream; eyes red-brown above and black-brown be-
low, a very narrow postocular ridge just laterad of the ocellar
tubercle; frons narrow, triangular, and with a small triangular
area in front of the ocellar tubercle; ocellar tubercle black,
granular; ocelli red-orange; mouthparts orange, two thirds as
long as the oral cavity; palpi orange, half as long as the oral
cavity; antennae with first segment short and red-brown, second
and third segments orange; third segment pear-shaped and ending
in two points; see figures for details.
Thorax dorsum yellow with a dark red-brown median band ex-
tending two thirds the distance to the scutellar sulcus and on
each side of this a red-brown lateral stripe extending from the
humeral callus to short of the scutellar sulcus, all dusted
with gray pollen; humeral and supraalar calli cream; propleuron
cream yellow, mesanepisternum cream yellow with a red-brown
basal spot, mesokatepisternum red-brown with a dorsal yellow
band and all dusted with gray pollen; wings milky white with
pale tan veins; halter stem brown, knob cream; fore legs with
coxae cream and remainder yellow-brown; middle legs with coxae
cream and remainder orange-brown with tarsi darkening at tip;
hind legs with coxae cream, femora red-brown, tibiae orange-
brown and tarsi orange-brown, darkening distally.
Abdomen with segments 1-8 white, second and third segments
marked with red-brown, wine-glass shaped marks on dorsal sur-
face; 9th tergite with base brown and the remainder cream; see
figures for details of terminalia.
FEMALE. Unknown.
No. 88] KELSEY: NEW SCENOPINIDAE 21
LENGTH. Male body 3.1 mm., wing 2.2 mm.
TYPE LOCALITY. Kennewick, Washington, (7 miles SW. ) , grass
beside irrigation ditch, 17 July 1957 (E. F. Dailey).
TYPE DEPOSITED. Washington State University, Pullman,
Washington, no. 328.
Brevitrichia forf icicruxa Kelsey, new species.
(Figure 15 . )
MALE. Head cream; eyes red-brown above, black-brown be-
low, and may have metallic glints when fresh; frons narrow,
triangular, depressed on lower midline; ocellar tubercle
black-brown, dusted with gray pollen; ocelli black-brown; oral
cavity deeply sunken, with mouthparts filling opening; ros-
trum brown, labellum tan, palpi tan and only as long as ros-
trum; antennae red-brown (third segment missing) .
Thorax dorsum black-brown dusted with bands of gray and
olive pollen and with an orange patch above and behind humeral
calli; humeral calli with ventral surface orange and a red-
brown dorsal band; supraalar calli orange; scutellum with a
large red-brown median spot and orange lateral angles; pro-
pleuron orange, mesanepisternum orange with a ventral band of
red-brown, mesokatepisternum red-brown with a narrow dorsal
orange band, all dusted with gray pollen; wings milky hyaline
with tan veins; halter stem brown, knob white with a tan-
brown dorsal rib; legs red-brown dusted with gray pollen,
except knees of femora which are orange-yellow.
Abdominal second segment with anterior margin and thick
central stem red-brown, lateral areas and posterior margins
white, and remaining segments except 9th white dorsally with
lateral margins red-brown; 9th tergum with the base red-brown,
white laterally, and all dusted with gray pollen; see figures
for details.
FEMALE. Unknown.
LENGTH. Male body 2.3 mm., wing 2 mm.
TYPE LOCALITY. Scissors Crossing, San Diego County, Cali-
fornia, 26 June 1958 (R. M. Bohart) .
TYPE DEPOSITED. California Academy of Sciences, no. 10107
(ex University of California, Riverside) .
Brevitrichia inferacanna Kelsey, new species.
(Figure 16 . )
MALE. Head cream; eyes red-brown above, black-brown below,
and with a narrow postocular rim; frons small, triangular, with
an inverted "V" groove in the swollen portion above the anten-
nae, and covered with granular pollen; ocellar tubercle black-
brown and covered with gray pollen; ocelli red-brown; antennae
with basal segments tan, third segment red-brown, pear-shaped,
and granular; see figures for details.
Thorax dorsum black-brown, covered with gray pollen, and
with a white lateral band extending to the scutellum from above
and including the humeral and supraalar calli; scutellum with
22 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
the lateral margins and base white, midportion with a red-brown
patch that tapers to a point on the posterior midline, all
covered with gray pollen; me sanepi sternum white with a red-
brown ventral spot, mesokatepisternum red-brown with a white
dorsal band, all dusted with gray pollen; wings milky white
with tan veins; halter stem brown, knob white; legs red-brown
and dusted with gray pollen, except knees of femora which are
white.
Abdomen red-brown with white posterior bands on all seg-
ments, except 9th tergum white with a proximal ventral red-
brown spot, all dusted with gray pollen; aedeagal prongs short,
straight, not exceeding the 9th tergum. The terminalia were
lost on dissection, so could not be illustrated but similar to
those of B. minuta Kelsey 1969, figure 132.
FEMALE. Unknown.
LENGTH. Male body 2.3 mm., wing 1.6 mm.
TYPE LOCALITY. Deep Canyon, Riverside County, California,
23 August 1963 (E. I Schlinger) .
TYPE DEPOSITED. California Academy of Sciences, no. 10108
(ex University of California, Riverside) .
Brevitrichia irwini Kelsey, new species.
(Figure 17. )
FEMALE. Head cream; eyes red-brown with a moderately broad
postocular ridge continuing uninterruptedly behind the ocellar
tubercle and a dark spot behind the eyes extending mesad toward
the median ocellus past the eye margin along the front; frons
broad, tan except for a narrow cream band next to each eye, and
with a shallow groove extending two thirds of the distance from
the median ocellus to the antennal bases; ocellar tubercle red-
brown, covered with tan pollen; antennae with first segment
red-brown, second segment orange and shorter than first, third
segment red-brown, pear-shaped, ending in two points; all seg-
ments dusted with pollen; see figures for details.
Thorax dorsum orange with a broad brown median band on
anterior two thirds bordered by narrow lateral bands on poste-
rior two thirds, all dusted with pollen; anterolateral portions
of dorsum with moderately long, light hairs; humeral and supra-
alar calli cream; mesoscutellum orange with only a small brown
spot at the mid-dorsal base; propleuron cream and orange,
mesanepisternum orange with dark red-brown ventral spot, meso-
katepisternum red-brown with a dorsal band of orange, all
dusted with pollen; wings brownish hyaline with brown veins;
halter stem tan, knob yellow, but lighter below; legs light
brown and dusted with pollen, except tips of tibiae red-brown
and darker.
Abdominal second and third segments mostly red-brown, each
having yellow posterior band and lateral margins, remaining
segments red-brown with orange lateral and posterior margins,
all subshining. Terminalia red-brown, 9th tergum with 6 black
spines and a tuft of hairs, additional hairs anteriorly direct-
ed, as also are the hairs of 8th segment; see figures for de-
tails.
No. 88] KELSEY: NEW SCENOPINIDAE 23
MALE. Unknown.
LENGTH. Female body 4.2 mm., wing 2.1 mm.
TYPE LOCALITY. One mile W. of Glamis , Imperial County,
California, 28 January 1965 (M. E. Irwin) . Sand dune associa-
tion rearing no. 65-1-28 B j . with larval and pupal skins.
TYPE DEPOSITED. California Academy of Sciences, no. 10109
(ex University of California, Riverside).
Brevitrichia miraloma Kelsey, new species.
(Figure 18.)
MALE. Head creamy white; eyes red-brown above, black-brown
below, extending to the back of the head and separating the
irons from the ocellar tubercle; frons small, triangular, ex-
tending two-fifths of the distance from antennae to median
ocellus; ocellar tubercle black-brown, dusted with gray pollen;
ocelli red-brown; back of head with central portion black-brown
and lateral portions creamy white; oral cavity creamy white;
mouthparts , filling oral cavity, light brown, but palpi tan;
antennae with first two segments short and brown; third anten-
nal segment orange-brown basally, red-brown distally, pear-
shaped, and ending in two sharp points with a median peg be-
tween; see figures for details.
Thorax dorsum generally black-brown and dusted with tan
pollen, area above humeral callus cream to orange, as are also
the lateral areas of the scutum, a black-brown spot on the
posterior midline of the scutum preceeded by a square of, and
bounded by lateral bands of, orange; humeral callus cream with
the dorsal margin black-brown, supraalar callus cream; meso-
scutellum cream to orange with a spot of black-brown occupying
the central fifth; propleuron creamy, me sanepi sternum with base
black-brown and dorsal rim cream, mesokatepisternum with base
black-brown and dorsal area cream-orange, all covered with
pollen; wings smoky white with tan veins; halter stem brown,
knob tan; legs with coxae cream, femora red-brown dusted with
tan pollen, and tibiae and tarsi brown.
Abdominal segments essentially white with black-brown
markings along midline except second and 9th segments, second
segment marked with a black-brown "T" with broad anterior,
central stem, and posterolateral extensions, remainder of seg-
ment white; 9th tergum white with a dark brown basal spot; all
segments dusted with tan pollen; see figures for details.
FEMALE . Unknown .
LENGTH. Male body 2.75 mm., wing 2.25 mm.
TYPE LOCALITY. One mile N. of Mira Loma, Riverside County,
California, 30 July 1967 (M. E. Irwin) , sand dune association.
HOLOTYPE DEPOSITED. California Academy of Sciences, no.
10642 (ex University of California, Riverside) .
Brevitrichia nayariti Kelsey, new species.
(Figure 19.)
FEMALE. Head light tan; eyes black-brown; postocular area
2 4 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
limited; brown area behind the eyes broad; groove from corner
of eye to ocellar triangle not attaining the back of the head,
so the two sides meet narrowly behind the ocellar tubercle;
frons broad, half again as wide as ocellar tubercle, cleft by
a median groove, and with a band of light brown extending from
ocellar tubercle to antennae; this band broadens anteriorly and
has a few sparse hairs along the margins; ocellar tubercle
black-brown and dusted with brown pollen; ocelli amber; rostrum
light brown, labellum tan, palpi tan and short, only as long as
rostrum; antennae with first two segments brown, but third seg-
ment orange-brown, pear-shaped, and forked at tip; see figures
for details.
Thorax dorsum dark brown, heavily crusted with brown and
tan pollen, but orange above humeral callus and on either side
of posterior portion of scutum; humeral and supraalar calli
cream; scutellum tan with a relatively narrow median band of
red-brown; propleuron cream, mesanepisternum red-brown with a
dorsal band of tan, mesokatepisternum with ventral portion red-
brown and dorsal area orange, all dusted with tan pollen; wing
tinged brown with brown veins; r-m more distant from R4 than
usual for the genus; halter stem tan, knob light tan above, and
cream below; legs light brown, dusted with pollen which makes
femora look lighter along the midline, first tarsal segment
lighter.
Abdomen red-brown with prominent paired dorsolateral longi-
tudinal cream bands on segments 2 to 7 and sternum cream along
mid portion; 8th segment red-brown, subshining, and sparsely
haired, 9th tergum with 8 spines; all dusted with pollen; see
figures for details.
MALE. Unknown
LENGTH. Female body 3.7 mm., wing 1.9 mm.
TYPE LOCALITY. Ten miles N. of Txtlon del Rio, Nayarit,
Mexico, 28 April 1961 (Howden & Martin).
TYPE DEPOSITED. Canadian National Collection, no. 11429.
Brevitrichia nevada Kelsey, new species.
(Figure 20.)
MALE. Head cream; eyes light red-brown above, darker be-
low, and extending to back of head; frons narrow, triangular;
ocellar tubercle black-brown, dusted with tan pollen; ocelli
red-orange; mouthparts well developed and filling the oral
cavity, brown, except palpi tan and half as long as mouthparts;
antennae with first segment brown and short, second segment
brown basally and orange-brown distally, third segment brown,
pear-shaped, and with two distinct points; all covered with gray
pollen.
Thorax with dorsum black-brown, dusted with brown and olive
pollen; notum with a crescent of orange-brown laterad of the
midline on the posterior portion and orange marginal areas;
humeral and supraalar areas cream; mesoscutellum with dorsal
portion black-brown, dusted with gray pollen, lateral margins
and ventral portion cream; propleuron orange, mesanepisternum
No. 88] KELSEY: NEW SCENOPINIDAE 25
largely black-brown with a thin orange dorsal band, mesokate-
pisternum black-brown below and orange above, all dusted with
gray pollen; wings milky hyaline with tan veins; halter stem
brown, knob tan with lateral margin white; legs red-brown, with
orange-brown femoro-tibial knees, and dusted with gray pollen.
Abdominal second segment with anterior and median portion
dark black-brown in the form of a "T" that does not attain the
posterior margin, remainder of segment and succeeding segments,
except 9th, white except lateral margins of tergites black-
brown, all dusted with gray pollen; 9th tergum with base and
dorsal portion black-brown, dusted with gray pollen, lower mar-
gins creamy white; see figures for details.
FEMALE. Head cream; eyes red-brown, with a narrow post-
ocular ridge set off from the frons by sulci which extend from
the upper corner of the eye curving to the center of the later-
al ocelli; the two sides of the head joining uninterruptedly
behind the ocellar tubercle, a brown spot lateral to the upper
corner of the eye extending toward back of the head, frons
nearly twice as wide as the ocellar tubercle and divided on the
midline by a shallow groove which is bordered by a brown area
that widens anteriorly from lateral of the median ocellus to
above, but not attaining the antennal socket, dusted with gray
pollen; ocellar tubercle black-brown, dusted with gray pollen;
ocelli red-orange; mouthparts and antennae as in the male; see
figures for details.
Thorax as in the male except wings smoky brown hyaline with
brown veins.
Abdomen mostly red-brown with creamy membranes at the pos-
terior margins of segments, all dusted with gray pollen; 8th
segment orange-brown; 9th tergum with 8 orange spines; see fig-
ures for details.
LENGTH. Male body 2.8 mm., wing 2.1 mm., Female body 3.7
mm. , wing 2. 2 mm.
TYPE LOCALITY. South end of Topaz Lake, Douglas County,
Nevada, 5,000 feet, 20 July 1967 (M. E. Irwin).
TYPES DEPOSITED. Holotype male and allotype female (on
same pin) in California Academy of Sciences, no. 10643 (ex
University of California, Riverside) .
Brevitrichia palida Kelsey, new species.
(Figure 21. )
FEMALE. Head cream; eyes pale (brown?, specimen may be
teneral) ; postocular ridge cut off from frons by a groove from
the upper corner of the eye to behind the lateral ocellus and
continuing uninterruptedly behind the ocellar tubercle, marked
with red-brown behind the eyes; frons broad, with a median
groove from the antennae to the median ocellus, bordered by a
broad band of tan the width of the ocellar tubercle; ocellar
tubercle black-brown and granular; mouthparts cream, well devel-
oped, and filling the oral cavity, but palpi white and only
half as long as mouthparts; antennae with first two segments
cream, third segment orange, pear-shaped, forked at tip, and
with a median peg between the forks; see figures for details.
2 6 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
Thorax dorsum with a median longitudinal black-brown band
on anterior three-fourths bordered by orange bands on either
side which become joined posteriorly and two lateral black-
brown bands extending from over the humeral calli to over the
supraalar calli, lateral margins cream, humeral callus white,
and supraalar callus cream; mesocutellum cream, with a narrow
black-brown median band which extends for a short distance
across the scutoscutellar sulcus, all dusted with gray pollen;
mesanepisternum cream with a red-brown patch, mesokatepisternum
red-brown with a cream dorsal border, all dusted with gray pol-
len; wings milky white with creamy to white veins; halter stem
cream, knob white; legs with femora orange-brown, tibiae cream,
and tarsi cream basally, darkening to brown distally.
Abdominal segments 2 to 4 orange-brown with mid-lateral
white areas, remainder of abdomen orange-brown, 8th segment
elongate, 9th tergum with 7 spines and sparse hair; see figures
for details.
MALE. Unknown.
LENGTH. Female body 3.3 mm., wing 2.3 mm.
TYPE LOCALITY. Twelve miles S. of Bakersfield, Kern Coun-
ty, California, 6 August 1958 (E. I. Schlinger) . Collected by
vacuum cleaner in alfalfa field.
TYPE DEPOSITED. California Academy of Sciences, no. 10110
(ex University of California, Riverside).
Brevitrichia palmacana Kelsey, new species.
(Figure 22 . )
FEMALE. Head cream; eyes red-brown; postocular ridge mod-
erately broad, separated from the frons by sulci from the up-
per corners of the eyes to the back of the ocellar tubercle,
the two sides divided by a median sulcus at the back of the
head; a tan spot behind eyes extending upward from the back of
the head, but not attaining the eye margin; frons broad, half
again as wide as the ocellar tubercle, a tan band the width of
the ocellar tubercle extending from the latter to the antennal
bases, divided on the midline by a shallow groove; back of head
black-brown, dusted with gray pollen; mouthparts extending two-
thirds the length of the oral cavity, light brown, but palpi
cream and only half as long as the mouthparts; antennae with
two basal segments red-brown, the third a lighter red-brown,
dusted with gray pollen, pear-shaped, and ending in two points
with a median peg between; see figures for details.
Thorax with pronotum yellow; mesonotum striped with three
longitudinal black-brown bands, median band broad narrowing on
the posterior fourth and continuing to posterior margin of the
scutellum, lateral bands separated from the median band and
scutellar margin by orange areas, all dusted with gray pollen;
humeral and supraalar calli cream; pleural areas largely light
orange with lower two-thirds of mesoan- and mesokatepisterna
red-brown and dusted with gray pollen; wings brownish hyaline
with brown veins; halter stem light red-brown and dusted with
gray pollen, knob cream-yellow; legs with femora red-brown and
dusted with gray pollen, tibiae light brown basally, but
No. 88] KELSEY: NEW SCENOPINIDAE 27
darkening distally, and tarsi red-brown, but lighter at base.
Abdomen black-brown basally, lightening to orange-brown
distally and dusted with gray pollen; 8th segment orange-brown;
9th tergum with 6 black spines; see figures for details.
MALE. Unknown.
LENGTH. Female body 4.5 mm., wing 2.5 mm.
TYPE LOCALITY. At mouth of Palm Canyon, 25 miles S. of
Quartzite, Arizona, 31 March 1968 (M. E. Irwin).
TYPE DEPOSITED. California Academy of Sciences, no. 10644
(ex University of California, Riverside) .
Brevitrichia piscifonta Kelsey, new species.
(Figure 23. )
MALE. Head white except black at back center; eyes red-
brown with fine upper and coarser lower facets; frons short,
triangular and unmarked; ocellar tubercle black-brown and
dusted with gray pollen; ocelli orange-brown; mouthparts short,
filling only half of the oral cavity, light tan, with palpi
light tan and shorter than rostrum; antennae light tan with
third segment pear-shaped and bifurcated at tip; see figures
for details.
Thorax with dorsal portion of scutum black-brown, dusted
with bands of gray and olive pollen, and with lateral margins,
area above humeral callus , and two arcs laterad of the midline
anterior to the scutellum, orange; humeral callus white, except
for a dash of red-brown along the dorsal margin; supraalar cal-
lus white; scutellum with a large central black-brown spot and
lateral and ventral portions creamy; propleuron creamy with
red-brown areas, mesanepisternum orange-yellow with a black-
brown ventral spot; mesokatepisternum black-brown with a creamy
dorsal margin; all pleural areas dusted with gray pollen; wings
smoky white with tan veins; halter stem brown and knob tan;
legs yellow-brown, with each segment a darker brown distally.
Abdominal second segment with a thick black-brown, central-
stemmed "T" , remainder white, segments 3 to 7 white with later-
al margins black-brown; 9th tergum with base red-brown, remain-
der white; aedeagal spines short; see figures for details.
FEMALE. Head creamy white; eyes red-brown with a moderate-
ly wide postocular area which continues uninterruptedly behind
the ocellar tubercle, a black mark laterad of the upper hind
corner of the eye; frons broad, with a central band of red-
brown, strongly cleft on the midline, and with a few scattered
red-brown hairs; ocellar tubercle black-brown, frosted with
gray pollen; ocelli red-brown; mouthparts and antennae as in
the male; see figures for details.
Thorax dorsum with three areas of black-brown separated by
bands of orange, mesal band extending two-thirds the distance
to the scutellum, flanked by two lateral areas extending from
behind the humeral callus to near the scutosculellar sulcus,
lateral margins orange; scutellum with a dark red-brown patch
covering middle third and continuing anteriorly across scuto-
scutellar sulcus as a thin band on the midline; lateral areas
of scutellum orange, and entire dorsum dusted with gray and
28 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
olive pollen; humeral and supraalar calli cream-white; pro-
pleuron cream-white, me sanepi sternum orange above and with a
red-brown spot below, mesokatepisternum red-brown with an
orange dorsal band, all dusted with gray pollen; wings milky
white hyaline with tan veins; halter stem brown, and knob tan;
legs brown.
Abdomen red-brown with tan margins; terminalia orange-
brown; 9th tergum with 8 light orange spines, the 4 dorsal ones
shorter than the ventral 4, and with some short hair. Access-
ory gland in the figures milky white; see figures for details.
LENGTH. Male body 2.7 mm., wing 2 mm. female body 2.8 mm.,
wing 1. 8 mm.
TYPE LOCALITY. Fish Springs, Imperial County, California,
13 September 1965 (R. C. Dickson) Yellow sticky trap.
TYPES DEPOSITED. Holotype male and allotype female (same
data as the male) , in California Academy of Sciences, no. 10111
(ex University of California, Riverside).
Brevitrichia septispina Kelsey, new species.
(Figure 24.)
FEMALE. Head yellow-orange; eyes black-brown with red
metallic glints; postocular ridge moderate, not separated by
ocellar tubercle or sulci; brown spots laterad of the dorsal
corner of the eyes small, not attaining posterior margin of
ridge; frons broad, with an orange-brown patch on center as
wide above and wider below than the ocellar tubercle; ocellar
tubercle black and granular; ocelli orange; mouthparts with
rostrum black-brown, labellum cream, and palpi dirty white, but
brownish at tips; antenna with first segment red-brown, second
orange and twice as long as first, third pear-shaped and red-
brown, but lighter at base; see figures for details.
Thorax dorsum with a black-brown band along the midline
on anterior three- fourths , separated from similar lateral bands
extending along the posterolateral three-fourths by a thin band
of orange which extends from an orange area above the humeral
callus to an orange area laterad of the midline on the posteri-
or fourth, lateral margins orange and all dusted with gray and
olive pollen; humeral and supraalar calli cream, mesocutellum
orange, with a median red-brown patch covering top third and
extending across scutoscutellar sulcus as a thin band on the
midline, all dusted with pollen; propleuron cream, mesanepis-
ternum orange with a red-brown basal patch and mesokatepis-
ternum red-brown with an orange dorsal band; wings hyaline with
brown veins; halter stem brown and knob cream; legs with fore
and middle coxae and trochanters orange, hind coxae and tro-
chanters brown, femora red-brown, tibiae orange with distal
end red-brown, all pollinose, and tarsi with proximal portion
of first segment orange, but distal portion and remainder red-
brown.
Abdominal second and third segments mostly red-brown, gran-
ular on midportion, but hind margins and lateral areas light
orange; fourth segment red-brown basally c*nd dark orange dis-
tally; remaining segments dark orange; except terminalia light
No. 88] KELSEY: NEW SCENOPINIDAE 29
red-brown, 9th tergum with 7 orange spines and black-brown
hairs; see figures for details.
MALE. Unknown.
LENGTH. Female body 4.5 mm., wing 2.4 mm.
TYPE LOCALITY. Eighteen miles W. of Blythe , Riverside
County, California, 2 April 1963 (M. E. Irwin).
TYPE DEPOSITED. California Academy of Sciences, no. 10112
(ex University of California, Riverside).
Brevitrichia wilcoxi Kelsey, new species.
(Figure 25 . )
MALE. Head brown with genae orange-brown; eyes red-brown
with green metallic glints; lower one-fourth of eye, when
viewed from side, darker, because of finer facets; frons small,
triangular, and depressed on midline; ocellar tubercle red-
brown; mouthparts with rostrum and labellum brown-orange, palpi
tan, short, and half the length of rostrum; antennae red-brown,
third segment swollen at base, forked at tip, and with median
peg subequal to lateral prongs; see figures for details.
Thorax mostly black, covered by dark and light stripes of
pollen; humeral and supraalar calli cream with an orange spot
behind humeral callus; scutellum black, dusted with gray pol-
len, and with lateral angles orange-brown; pleurum generally
black, dusted with light gray pollen, but with a few light
areas; wings hyaline with brown veins; halter stem red-brown
and knob cream-yellow; legs dark red-brown, dusted with gray
pollen, but hind tibiae yellow-brown.
Abdomen with all segments red-brown, dusted with gray pol-
len, and with posterior membranes narrow, whitish; 9th tergum
red-brown basally, distal portion creamy white with long white
hairs; aedeagal bristles very long; see figures for details.
FEMALE. Unknown.
LENGTH. Male body 3.8 mm., wing 2.2 mm.
TYPE LOCALITY. Eighteen miles S. of Gila Bend, Arizona,
12 May 1965, (J. Wilcox).
TYPE DEPOSITED. California Academy of Sciences, no. 10113
(ex University of California, Riverside) .
GENUS PSEUDATRICHIA OSTEN SACKEN
The genus Pseudatrichia, containing 33 described species,
has a range from south central British Columbia through Central
America to Brazil. To date no species have been recorded east
of the Rocky Mountain chain. Two of the species reported here
represent the most easterly and northeasterly examples of North
American distribution of the genus. The three new species des-
cribed below raise the number of known Pseudatrichia species to
36.
30 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
The following changes in the key to the species of Pseuda-
trichia in Kelsey (1969, Pg. 223) will permit their separation.
Key to the species of Pseudatrichia (Males)
10. (8) Antennae orange-brown 10a
Antennae black-brown P. unicolor Coquillett
10a (10) Seventh abdominal segment elongate,
red-brown; 9th segment black-
brown P. ruf itruncula Kelsey, new species
Seventh abdominal segment short, shining,
black; 9th segment black. . . P. gracilipennis Kelsey
Key to the species of Pseudatrichia (Females)
5. (4) Abdomen with 8th segment long and narrow 5a
Abdomen with 8th segment only slightly
longer than wide, marked with black
areas dorsally and ventrally (Mexico) . P. longurio Loew
5a. (5) Median emargination of sternum
pointed (Costa Rica) .... P. longiventris Kelsey
Median emargination of sternum gently
rounded (Mexico and Texas)
P. rufitruncula Kelsey, new species
16. (13) Eighth sternum gently rounded on
posterior margin 16a
Eighth sternum notched on posterior
midline P. evergreeni Kelsey
16a (16) Vein R. arising from middle of cell,
undulating to attain wing margin
beyond tip of cell R5 . . . . P. unicolor Coquillett
Vein R4 arising from distal third cell
and reaching wing margin before tip of
cell R5 ; cell R5 pointed
P. schlingeri Kelsey, new species
20. (19) Vein R4 ending in wing margin opposite
tip of cell R5 20a
Vein R4 ending in wing margin well
beyond tip of cell R5 P. barnesi Kelsey
20a (20) Vein M^+2 joining vein R5 at nearly a
right angle, making cell R5 blunt; 9th
sternum trilobed. . P. idahoensis Kelsey, new species
Vein M]_+2 joining R5 more gradually,
cell R5 more pointed; 9th sternum
excavated on the midline P. bakeri Kelsey
Pseudatrichia idahoensis Kelsey, new species.
(Figure 26 . )
FEMALE. Head black; eyes black-brown; postocular ridges
prominent, the two ridges separated by the ocellar tubercle;
No. 88] KELSEY: NEW SCENOPINIDAE 31
frons moderately broad, not wider than ocellar tubercle,
roundly swollen above the antennal bases and with a faint medi-
an groove on the upper half; ocellar tubercle black, set off;
ocelli red; back of head pollinose; mouthparts brown, filling
oral cavity, palpi brown, half as long as rostrum; silvery pile
bordering oral cavity; antennae with basal segments black-brown
and short; third segment black-brown at base, orange-brown elon-
gate and truncated at tip; see figures for details.
Thorax black, rugose, transversely striated, and with a
pilose area above humeral callus; humeral and supraalar calli
orange-brown; pleural areas black; wings hyaline with black-
brown veins; vein M1+2 joins R5 at a right angle; halter stem
brown and knob white with a brown base and dorsal rib; fore
legs orange, middle legs orange-brown, and hind legs red-brown.
Abdomen black, rugose and with thin, white, intersegmental
membranes; 8th sternum hairy; see figures for details.
MALE. Unknown.
LENGTH. Female body 7.5 mm., wing 4.7 mm.
TYPE LOCALITY. Chatcolet, Idaho, 3-4 July 1963 (M. & H.
James) .
TYPE DEPOSITED. Washington State College, Pullman, no.
329.
Pseudatrichia ruf itruncula Kelsey, new species.
(Figure 27. )
MALE. Head black, not distinctly longer than high; eyes
black-brown and with no apparent division into coarse upper
and finer lower facets; postocular ridge distinct, elevated
and rugose; frons narrow and triangular, separating the eyes
by the width of the median ocellus, lower frons rugose with
the margins next to eyes shining; ocellar tubercle black and
rugose, sharply cut off from postocular ridge; ocelli clear;
back of head concave below ocellar tubercle; mouthparts or-
ange-brown and well developed, but palpi lighter, slender, and
two-thirds the length of the rostrum; antennae with first seg-
ment brown and short; second orange-brown and longer than
first; third segment orange-brown, oval about twice as long
as broad, and with two lateral points and a distinct median
peg at apex; see figures for details.
Thorax dorsum black, rugose, and with a patch of white
pilosity anterior to each humeral callus; humeral callus red-
orange; pleuron dark red-brown and shining; wings hyaline with
orange-brown veins; halter stem red-brown, knob with base red-
brown and remainder white; legs orange-brown but hind femora
darker distally.
Abdomen black, rugose, with a purple metallic sheen, and
with segments 3 and 4 showing narrow white membranes, except
7th segment red-brown and 9th segment black-brown; see figures
for details.
FEMALE. Head black, longer than high; eyes black-brown
and elongate; postocular flange broad, subshining, and extend-
ing posteriorly farther than ocellar tubercle; frons narrow,
covered with several rows of recumbent, anteriorly directed
hairs, and produced in a rounded bulge above antennae; ocellar
32 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
tubercle black and shining; ocelli orange-brown; mouthparts as
in the male; antennae with first two segments short, black, and
granular, third red-brown, oval truncated, and with median peg
at tip; see figures for details.
Thorax black, depressed, granular, and with anterodorsal
portion pointed; humeral and supraalar calli red-brown; pleural
areas black and subshining; wings tinged brown and with micro-
trichia and veins brown; halter stem brown and knob red-brown,
elongate, and pointed; legs orange-brown, with hind femora and
tibae darker.
Abdomen with first seven segments black, rugose, and with
a metallic sheen; 8th dark red-brown, elongate, with dorsum
pointed and sternum hairy; see figures for details.
LENGTH. Male body 7.5 mm., wing 4.5 mm., female body 7.5
mm., wing 3.7 mm.
TYPE LOCALITY. Chisos Basin, Big Bend National Park,
Texas, 20 April 1959 (Howden & Baker).
HOLOTYPE MALE DEPOSITED. Canadian National Collection,
no. 11430.
ALLOTYPE FEMALE. Chis , N. Chiapas (3 km. S. Oaxaca, Rt.
190, 5-600'). Mexico, 12 July 1962 (H. E. Milliron) , deposited
in Canadian National Collection.
Pseudatrichia schlingeri Kelsey, new species.
(Figure 28.)
FEMALE. Head black, eyes red-brown; postocular flange
broad and rising to a ridge which drops sharply to the back of
the head, narrowly separated on back of head by the ocellar
tubercle; back of the head nearly flat; frons subshining,
slightly broader than ocellar tubercle, and with two diverging
bands of anteriorly directed hairs extending from before the
median ocellus to laterad of the antennal bases; ocellar tu-
bercle prominent, cut off by two deep grooves running from the
upper corners of eyes to behind the ocellar tubercle, but not
joining on the back of the head; oral cavity deep, bordered
laterad of the antennae by silvery pile; mouthparts brown,
filling two- thirds of oral opening, and palpi short; antennae
black-brown, the first two segments short, the third oval, with
bluntly bifurcated tip, and covered with very short hairs that
give a brownish color; see figures for details.
Thorax black, subshining, and transversely striated;
humeral callus with posterior tip orange; scutellum black; pre-
coxal area of prothorax with gray pollen, remainder of pleural
areas black-brown and subshining; wings hyaline with brown
veins; halter stem brown and knob with base brown, but remain-
der white; legs with coxae orange, femora orange, darkening to
orange-brown, tibiae orange-brown and tarsi brown.
Abdomen black, except 8th sternum red-brown at base, proxi-
mal segments rugose with a faint metallic sheen; 8th sternum
smoothly rounded at tip; see figures for details.
MALE. Unknown.
LENGTH. Female body 5.7 mm., wing 3.7 mm.
TYPE LOCALITY. Riverside, Riverside County, California,
No. 88] KELSEY: NEW SCENOPINIDAE 33
1 May 1966 (E. I. Schlinger) .
HOLOTYPE DEPOSITED. California Academy of Sciences, no.
10645 (ex University of California, Riverside) .
PARATYPE. 1 female, Riverside, Riverside County, Califor-
nia, 6 April 1967 (Saul Frommer) in University of California,
Riverside.
GENUS BELOSTA D. E. HARDY
New material received recently from the Department of En-
tomology of the University of California at Riverside contained
the two new species of Belosta described below. Added to the
five previously described (Kelsey 1969), this increases the
known species to seven. Although one of the new species, a
pair, was collected on the windows of a laboratory, no new
clues on the biology of the genus were obtained.
The following key to the genus has been completely revised
to include all described species.
Key to the species of Belosta
1. Males; eyes close together 2
Females; eyes widely separated 5
2. (1) Halter knob white above, or marked with
red-brown 3
Halter knob tan; thorax with short white
hairs B. albipilosa Hardy
3. (2) Halter knob white above 4
Halter knob with base cream and a red-
brown band down center; thorax with
medium long white hairs
B. albicomula Kelsey, new species
4. (3) Halter knob all white; thorax with
long white hairs B. pilosa (Coquillett)
Halter knob white above with lateral margin
and ventral surface tan; thorax with very
short scale-like hairs . . B. viticolapennis Kelsey
5. (1) 8th sternum long, sloping, without a
brush of hairs
8th sternum short, abrupt, with a ventral
brush of stiff spines 6
6. (5) Halter knob orange; thorax with mixed
short red and white hairs . . . . B. albipilosa Hardy
Halter knob with base and dorsal rib brown,
a white semicircle around the brown
knob, ventral surface white; thorax
with short white hairs; frons with a
creamy diamond on upper portion
B. scutulamacula Kelsey, new species
7. (5) Halter knob red-brown or marked with
red-brown 8
Halter knob orange; thorax with long
white hairs B. pilosa (Coquillett)
34 CALIFORNIA ACADEMY OF SCIENCES [Occ. Papers
8. (7) Halter knob red-brown above, lighter below .... 9
Halter knob white, and with a longitudinal
band of red-brown, thorax with patches of
short white hairs . B. albicomula Kelsey, new species
9. (8) Halter knob with ventral tip white. B. telfordi Kelsey
Halter knob cream below .... B. termitophaga Kelsey
Belosta albicomula Kelsey, new species.
(Figure 29. )
This new species closely resembles B. pilosa (Coquillett)
but differs in many small ways. The pile hairs of the thorax
are shorter, the antennae are longer and less swollen in the
center, the halters are colored differently, and the terminalia
of both sexes show distinct differences.
MALE. Head black, higher than long; eyes black-brown
above, black below, and extending to the back of the head dor-
sally; mouthparts atrophied, palpi club-shaped, extending one-
third the length of the oral cavity; lower postgenae covered
with medium-long white hairs; frons shining, triangular, and
extending dorsally as a thin band between the eyes to the medi-
an ocellus; ocellar tubercle elevated and distinct; ocelli red-
brown; antennae black-brown, with the first two segments short,
third segment lighter at tip, diamond shaped, and three times
as long as wide, with greatest width at midpoint; see figures
for details.
Thorax with the dorsum black, covered with medium-long
white hairs, much like B. pilosa except shorter; humeral and
supraalar calli orange-brown; scutellum black and hairy; wings
hyaline, with tan veins; halter stem red-brown, knob cream with
red-brown longitudinal band; legs with coxae to femora red-
brown and with yellow knees and hairy; tibiae yellow, darkening
to red-brown distally; tarsi red-brown.
Abdomen black, sparsely haired on basal segments, with
intersegmental membranes white posteriorly and yellow lateral-
ly; terminalia short; sternal comb thin; aedeagus with short
parameres; 9th segment claspers broadly rounded at tip; see
figures for details.
FEMALE. Head mostly orange-brown, darker below eye margin,
with the back of the head black-brown medially, higher than
long; eyes black-brown; mouthparts atrophied, palpi black-brown
and club-shaped; frons broad, smooth, shining, with a median
depression and black bands extending from the eye margins
laterad of the antennal bases to the ocellar tubercle, giving
the frons the appearance of being black with an orange median
area and two lateral orange patches above the eyes; ocellar
tubercle black; ocelli black-brown; antennae black-brown, the
first two segments black, and short (third segment lost); see
figures for details.
Thorax dorsum black, shining, and thinly covered with
sparse patches of short white pile; humeral callus cream with
a median brown spot; supraalar callus orange-brown; propleuron
cream and brown, mesopleuron black, with short white hairs;
wings hyaline, with brown veins; halter stem black-brown and
No. 88] KELSEY: NEW SCENOPINIDAE 35
knob white with a longitudinal band of red-brown; legs with
coxae to femora dark red-brown and with yellow knees; tibiae
yellow darkening to red-brown distally; tarsi red-brown.
Abdomen black with white intersegmental membranes; 8th ter-
gum marked with orange laterally; 8th sternum elongate and
toothed distolaterally ; see figures for details.
LENGTH. Male body 5.7 mm., wing 4.1 mm; female body 7.5
mm., wing 5.0 mm.
TYPE LOCALITY. Riverside, California, 1 June 1964 (E. I.
Schlinger) .
TYPE DEPOSITED. California Academy of Sciences, no. 10101
(ex University of California, Riverside) .
ALLOTYPE. Female; same date as type; deposited in Califor-
nia Academy of Sciences.
Belosta scutulamacula Kelsey, new species.
(Figure 30. )
This species is quite close to B. albipilosa Hardy, but may
be separated from it by the white diamond-shaped marking in a
dark field on the upper frons anterior to the median ocellus,
the absence of the dark spot below the eye of the female, the
thorax being covered with a pile of very short hairs giving a
dusty appearance, the halter color, the appearance of the 7th
sternum, the shape of the 8th and 9th tergites, and the shorter
spermatheca.
FEMALE. Head color basically creamy white, but back of
head dark red-brown, head higher than long; eyes black-brown; a
groove extending from the upper corner of the eye to the later-
al ocellus; a narrow red-brown band posterior to this groove;
frons shining, two grooves extend from laterad of the ocellar
tubercle to laterad of the antennal bases; central area of
frons dark brown with a creamy diamond on the upper half, the
dorsal point of which touches the median ocellus; the area be-
tween the grooves and eye margins marked with brown; ocellar
tubercle dark red-brown; ocelli red-orange; oral cavity white;
mouthparts atrophied, palpi red-brown and short; antennae
brown, the first two segments short, third oval and about
twice as long as wide; see figures for details.
Thorax with dorsum black, clothed with short white hairs
giving a dusty appearance (hairs shorter than those of B.
albipilosa and none reddish) ; humeral and supraalar calli creamy
white; pleural areas black with long white hairs; wings hyaline,
irridescent under reflected light, with yellow-brown veins;
halter stem brown and knob with base and dorsal rib brown, a
white semicircle around the brown base, and ventral surface
white; legs dark red-brown, except knees of femora creamy white.
Abdomen red-brown with a white posterior membrane on all
segments; 7th sternum with a fringe of spines; 8th sternum
short, abrupt, with a median brush of stiff spines, basal por-
tion with pits only, 9th tergum pointed; spermatheca elongate,
but only a little over half as long as that of B. albipilosa;
see figures for details.
MALE. Unknown.
36
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
LENGTH. Female body 5.5 mm., wing 3.6 mm.
TYPE LOCALITY. Los Prietos Campground, Santa Barbara Coun-
ty, California, 30 July 1965, (J. Bath, B. Ruge) .
TYPE DEPOSITED. California Academy of Sciences, no. 10102
(ex University of California, Riverside).
FIGURE 1. Scenopinus estesi Kelsey, new species, male
wing; b. and c. , lateral and frontal aspects of head;
a. ,
d. , enlarged detail of antenna; e., f . , and g. , lateral,
ventral (physically dorsal) and posterior aspects of
terminalia.
No . 8 8]
KELSEY
NEW SCENOPINIDAE
37
FIGURE 2. Scenopinus transversus Kelsey, new species,
female: a., wing; b. and c. , lateral and frontal aspects of
head; d. , enlarged detail of antenna and transverse groove on
lower f rons ; e. and f . , lateral and ventral aspects of 8th and
9th segments; g. , 9th sternum and roof of bursa.
38
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 3. Scenopinus vockerothi Kelsey, new species, male
and female; a. , wing; b. and c. , lateral aspects of male and
female heads; d. and e., enlarged details of male and female
antennae; f. and g. frontal aspects of male and female heads;
h. , i., and j., lateral, dorsal (physically ventral), and pos-
terior aspects of male terminalia; k. and 1. , lateral and
ventral aspects of female 8th and 9th segments; m. , female 9th
sternum and bursa.
No. 88]
KELSEY: NEW SCENOPINIDAE
39
FIGURE 4. Scenopinus albipilosus Kelsey, new species,
female; a., wing; b. and c. , lateral and frontal aspects of
head; d. and e. , lateral and ventral aspects of 8th and 9th
segments; f . , 9th sternum and roof of bursa.
40
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE
female ; a. ,
head; d. and
segments; f.
5. Scenopinus aurantipalpus Kelsey, new species
wing; b. and c. , lateral and frontal aspects of
e., lateral and ventral aspects of
, 9th sternum and roof of bursa; g.
8th and 9th
spermatheca,
No . 8 8]
KELSEY: NEW SCENOPINIDAE
41
FIGURE 6. Scenopinus dietricki Kelsey, new species, fe-
male: a., wing; b. and c. , lateral and frontal aspect of head;
d. , enlarged detail of antenna; e. and f . , lateral and ventral
aspect of 8th and 9th segments; g. , 9th sternum and roof of
bursa; h. , lateral view of bursa; i., spermatheca.
42
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 7. Scenopinus foxi Kelsey, new species, female:
a. , wing; b. , lateral aspect of head; c. and d. , enlarged
lateral and frontal aspects of head; e. and f . , lateral and
ventral aspects of 8th and 9th segments; g. , 9th sternum and
roof of bursa.
No. 88]
KELSEY: NEW SCENOPINIDAE
43
FIGURE 8. Scenopinus johnsoni Kelsey, new species, male
and female: a., female wing; b. and c. , lateral and frontal
aspects of female head; d. , enlarged detail of female antenna;
e. , male wing; f . and g. , lateral and frontal aspect of male
head; h. , enlarged detail of male antenna; i. , j. , and k. ,
lateral, ventral (physically dorsal)/ and posterior aspects of
male terminalia; 1. and m. , lateral and ventral aspects of fe-
male 8th and 9th segments; n. , 9th sternum and bursa of female;
o. , spermathecae.
44
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 9. Scenopinus reciprocatus Kelsey, new species,
male and female; a., wing; b., c. , d. , and e., lateral and
frontal aspects of male and female heads; f. and g. , enlarged
details of male and female antennae; h. , i., and j., lateral,
ventral (physically dorsal), and posterior aspects of male ter-
minalia; k. and 1. , lateral and ventral aspects of female 8th
and 9th segments; m. , female 9th sternum and roof of bursa; n.
lateral aspect of female bursa.
No. 88]
KELSEY: NEW SCENOPINIDAE
45
FIGURE 10. Brevitrichia albinota Kelsey, new species,
male: a., wing; b., lateral aspect of head; c. , enlarged
detail of antenna; d. , enlarged frontal aspect of head; e.
and f . , ventral (physically dorsal) and lateral aspects of
terminalia.
46
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
and
f
Brevitrichia arena Kelsey, new species, male
and b. , male and female wings; c. , d. , e. , and
lateral and frontal aspects of male and female heads; g,
FIGURE 11.
female: a.
enlarged dorsal aspect of female head; h. , enlarged detail of
antenna; i. and j., lateral and ventral (physically dorsal) as-
pects of male terminalia; k., lateral aspect of female 8th and
9th segments; 1., dorsal, right, and ventral, left, aspects of
female 8th and 9th segments; m. , female bursa; n. , spermatheca,
No . 8 8]
KELSEY: NEW SCENOPINIDAE
47
FIGURE 12. Brevitrichia argent aurb a Kelsey, new species,
male: a., wing; b. and c. , lateral and frontal aspect of head;
d. , enlarged detail of antenna; e. and f . , lateral and ventral
(physically dorsal) aspects of terminalia.
48
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 13. Brevitrichia badiclitella Kelsey, new species,
male: a., wing; b., lateral aspect of head; c. , enlarged detail
of antenna; d. , enlarged frontal aspect of head; e. and f . ,
lateral and ventral (physically dorsal) aspects of terminalia.
No. 88]
KELSEY: NEW SCENOPINIDAE
49
FIGURE 14. Brevitrichia daileyi Kelsey, new species, male:
a., wing; b. and c. , lateral and frontal aspects of head; d. ,
enlarged detail of antenna; e. and f . , lateral and ventral
(physically dorsal) aspects of terminalia.
50
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 15. Brevitrichia forf icicruxa Kelsey, new species,
male: a., wing; b. and c. , lateral and frontal aspects of
head; d. and e., lateral and ventral (physically dorsal) as-
pects of the terminalia.
No. 88]
KELSEY: NEW SCENOPINIDAE
51
FIGURE 16.
male: a. , wing; b. ,
Brevitrichia infecacanna Kelsey, new species,
and d. , en-
lateral aspect of head; c
larged lateral and frontal aspects of head.
52
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 17. Brevitrichia irwini Kelsey, new species, fe-
male: a., wing; b., lateral aspect of head; c. , enlarged de-
tail of antenna; d. , enlarged dorsal aspect of head; e., dor-
sal, right, and ventral, left, aspects of 8th and 9th seg-
ments; f . , lateral aspects of 8th and 9th segments; g. , sper-
mathecae .
No . 8 8]
KELSEY: NEW SCENOPINIDAE
53
FIGURE 18. Brevitrichia miraloma Kelsey, new species,
male: a. , wing; b. , lateral aspect of head; c. , enlarged de-
tail of antenna; d. , enlarged frontal aspect of head; e. and
f . , lateral and ventral (physically dorsal) aspects of termin-
alia.
54
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 19. Brevitrichia nayariti Kelsey, new species,
female: a., wing; b., lateral aspect of head; c. , enlarged
dorsal aspect of head; d. , ventral, right, and dorsal, left,
aspects of 8th and 9th segments; e., lateral aspect of 8th and
9th segments; f . , bursa; g. , spermatheca.
No. 8 8]
KELSEY
NEW SCENOPINIDAE
55
and
FIGURE
female
20.
a. ,
Brevitrichia nevada Kelsey, new species, male
wing; b. , c. , and d. , lateral and frontal as-
pects of male head and lateral aspect of female head; e., en-
larged detail of male antenna; f . and g. , enlarged lateral and
dorsal aspects of female head; h. and i., lateral and ventral
(physically dorsal) aspects of male terminalia; j . , lateral as-
pect of female 8th and 9th segments; k. , dorsal, right, and
ventral, left, aspects of female 8th and 9th segments; 1.,
spermatheca.
56
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 21. Brevitrichia palida Kelsey, new species, fe-
male: a., wing; b. and c. , lateral and dorsal aspect of head;
d. , enlarged detail of antenna; e., dorsal, right, and ventral,
left, aspect of 8th and 9th segments, bursa outlined; f . ,
lateral aspect of 8th and 9th segments.
No . 8 8]
KELSEY: NEW SCENOPINIDAE
57
FIGURE 22. Brevitrichia palmacanna Kelsey, new species,
female: a., wing; b., lateral aspect of head; c. , enlarged
detail of antenna; d. , enlarged dorsal aspect of head; e., dor-
sal, right, and ventral, left, aspects of 8th and 9th segments,
bursa outlined; f . , lateral aspect of 8th and 9th segments;
g. , spermatheca.
58
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 23. Brevitrichia piscifonta Kelsey, new species,
male and female: a. and b., male and female wings; c. and d.
lateral aspects of male and female heads; e., frontal aspect
of male head; f . , enlarged dorsal aspect of female head; g. ,
enlarged detail of antenna; h. and i., lateral and ventral
(physically dorsal) aspects of male terminalia; j., dorsal,
right, and ventral, left, aspects of female 8th and 9th seg-
ments; k. , lateral aspect of female 8th and 9th segments; 1.,
female bursa; m. , accessory gland.
No. 88]
KELSEY: NEW SCENOPINIDAE
59
FIGURE 24.
female: a., wing;
head; d.
Brevitrichia septispina Kelsey, new species,
lateral and dorsal aspects of
c.
b. and
nca^, yu. , enlarged detail of antenna; e., dorsal, right, and
ventral, left, aspects of 8th and 9th segments; f . , lateral as-
pect of 8th and 9th segments; g. , bursa; h. , spermathecae .
60
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 25. Brevitrichia wilcoxi Kelsey, new species, male
a., wing; b. and c. , lateral and frontal aspect of head; d. ,
enlarged detail of antenna; e. and f . , lateral and ventral
(physically dorsal) aspects of terminalia.
No. 88]
KELSEY: NEW SCENOPINIDAE
61
FIGURE 26. Pseudatrichia idahoensis Kelsey, new species,
female: a., wing; b. and c. , lateral and dorsal aspects of
head; d. and e . ,
segments .
b . and c. ,
ventral and lateral aspects of 8th and 9th
62
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 27. Pseudatrichia rufitruncula Kelsey, new species,
male and female: a. and b. , male and female wings; c. and d. ,
lateral and dorsal aspects of male head; e., enlarged detail
of male antenna; f . and g. , lateral and dorsal aspects of fe-
male head; h. , enlarged detail of female antenna; i., j., and
k. , lateral, ventral (physically dorsal), and posterior aspects
of male terminalia; 1. and m. , lateral and ventral aspects of
female 8th and 9th segments; n. , spermatheca.
No. 88]
KELSEY: NEW SCENOPINIDAE
63
FIGURE 28. Pseudatrichia schlingeri Kelsey, new species,
female: a. , wing; b., lateral aspect of head; c. , enlarged
dorsal aspect of head; d. , enlarged detail of antenna; e. and
f . , ventral and lateral aspects of 8th and 9th segments.
64
CALIFORNIA ACADEMY OF SCIENCES
[Occ. Papers
FIGURE 29. Belosta albicomula Kelsey, new species, male
and female: a., wing; b., c. , d. , and e., lateral and frontal
aspects of male and female heads; f . , g. , and h. , lateral,
ventral (physically dorsal) , and posterior aspects of male
terminalia; i., lateral aspect of female 8th and 9th segments;
j . , dorsal, right, and ventral, left, aspects of female 8th and
9th segments.
No . 88]
KELSEY: NEW SCENOPINIDAE
65
FIGURE 30. Belosta scutumacula Kelsey, new species, fe-
male: a., wing; b. and c. , lateral and frontal aspects of
head; d. , e., and f . , dorsal, ventral, and lateral aspects of
8th and 9th segments; g. , spermathecae.
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