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UNIVERSITY OF LONDON
FRANCIS GALTON LABORATORY FOR NATIONAL EUGENICS
EUGENICS LABORATORY MEMOIRS. IV.
ON THE MEASURE OF THE RESEMBLANCE
OF FIRST COUSINS
BY
ETHEL M. ELDERTON,
GALTON RESEARCH SCHOLAR IN NATIONAL EUGENICS,
UNIVERSITY OF LONDON
ASSISTED BY
KARL PEARSON, F.R.S.
LONDON :
PUBLISHED BY DULAU AND CO., 37, SOHO SQUARE, W.
1907
Price Tliree Shillings and Sixpence
THE FRANCIS GALTON EUGENICS LABORATORY.
University of London, University College, Gower Street, W.C.
The Laboratory is under the supervision of Professor Karl Pearson, F.R.S.,
in consultation with Mr Francis Galton, F.R.S.
Francis Galton Fellow in National Eugenics : David Heron, M.A.
Francis Galton Scholar : Miss E. M. Elderton.
Computer : Miss Amy Barrington.
Advisory Committee. — The following have kindly consented to aid the Staff
of the Laboratory in special forms of enquiry : —
W. Palin Elderton. E. Nettleship.
J. Macpherson, M.D. Edgar Schuster, M.A.
F. Mott, M.D., F.R.S. Lieut. -Colonel R. J. Simpson, R.A.M.C.
J. F. Tocher.
National Eugenics is the study of agencies under social control that may
improve or impair the racial qualities of future generations,
either physically or mentally.
It is the intention of the Founder, Mr Francis Galton, that the Laboratory
shall act (i) as a storehouse for statistical material bearing on the mental and
physical conditions in man and the relation of these conditions to inheritance
and environment, (ii) as a centre for the publication or other form of distribution
of information concerning National Eugenics. Provision is made in association
with the Biometric Laboratory at University College for training in Statistical
Method and for assisting research workers in special Eugenics Problems.
Short courses' of instruction will be provided for those engaged in social,
anthropometric; or medical work and desirous of applying- modern methods of
analysis to the reduction of their observations.
UNIVERSITY OF LONDON
FRANCIS GALTON LABORATORY FOR NATIONAL EUGENICS
EUGENICS LABORATORY MEMOIRS. IV.
ON THE MEASURE OF THE RESEMBLANCE
OF FIRST COUSINS
BY
ETHEL M. ELDERTON,
GALTON RESEARCH SCHOLAR IN NATIONAL EUGENICS,
UNIVERSITY OF LONDON
ASSISTED BY
KARL PEARSON, F.R.S.
LONDON :
PUBLISHED BY DULAU AND CO., 37, SOHO SQUARE, W.
1907
^v*V^
70709
Some reconstruction of the Francis Galton Laboratory having taken place, it
seemed desirable to provide the workers associated with it with a direct channel
of publication of their own, in which their more extended memoirs should appear.
It is hoped that the present sei'ies may be issued at short intervals. Subscribers
should notify their intention of taking in the memoirs as they are published
to Messrs Dulau & Co. Requests to exchange with similar publications, with
archives and journals dealing with demographic and sociological problems, or
with census reports, should be directed to The Editor, Eugenics Laboratory,
University College, Gower Street, London, W.C.
On a Measure of the Resemblance of First Cousins in Man.
J
By Ethel M. Eldertox, Galton Research Scholar in National Eugenics
in the University of London ; assisted by Karl Pearson, F.R.S.
1. Introductory. While a very large amount of data has been collected,
reduced and published relating to the degree of resemblance in physical and psychical
characters of a considerable number of pairs of relatives in man — especially in the
direct line and between collaterals of the first degree — but little has yet been done
with regard to collaterals in higher degrees. As far as we are aware tbe only quanti-
tative measures yet determined are those for eye colour in man between uncle or
aunt and nephew or niece*. No measure of resemblance has yet been determined for
cousins. Yet it is precisely among collaterals of the second degree that the question
of consanguineous marriages becomes in practical life of great importance, inequality
of age being bere less marked, and thus the degree of resemblance between such
collaterals has not only scientific but eugenic value. According to local law and
religious custom cousin marriages are permitted or forbidden ; thus it would appear
that we are here concerned with divergent human experiences, unconsciously formu-
lated, as to the relative value of endogamy and exogamy. If we take a character
whicb is detrimental to tbe individual, it will, at least in primitive communities,
be in the bulk of cases a hindrance to mating. Hence, as a rule, we must classify
such a detrimental character as recessive in the Mendelian senset, otherwise selection
would have weeded it out. Now consider for a moment a population of dominants
with notation DD, and suppose one of these to mate with an individual of detrimental
attribute and constitution RR. The result will be the hybrid sibship marked by DR,
in which the recessive character R will be latent. If brother-sister mating is for-
bidden the next generation will be obtained (assuming the recessive individuals RR
to be extremely rare) by mating with the population of dominant character, and the
result will be equal numbers of DD and DR. Thus the generation of cousins would
consist of 50 p. c. of dominants and 50 p. c. apparent dominants with the detrimental
character recessive. ' It therefore follows that it would be as detrimental for some
cousins to marry as for all brothers and sisters of the first hybrid sibship. That is to
* Pearson and Lee : Phil. Trans. Vol. 195, A, p. 114 et seq.
t Thus albinism and the tuberculous diathesis are, if not true and complete recessives in the Mendelian
sense, still more nearly recessive than dominant characteristics. But there are other abnormalities, e.g.
certain digital deformations, which are nearer to, if perhaps not true, dominants.
1—2
4 ETHEL M. ELDERTON
say while a brother and sister marriage would lead to 25 p. c. of the offspring having
the harmful character patent and another 50 p. c. having it latent, the intermarriage
of the cousins of the DR class among themselves would lead to the same baneful
results as this brother-sister marriage, while the intermarriage of the DR class of
cousin with the DD would also lead to 50 p. c. with the latent detrimental character.
In other words endogamy as far as brothers and sisters are concerned would lead to :
25 p. c. hale. 50 p. c. latent evil. 25 p. c. patent evil,
while endogamy in the cousinship would give us:
56"25 p. c. hale. 37"5 p. c. latent evil. 6-25 p. c. patent evil.
The explanation therefore of the wide-spread social feeling against endogamy in
the first degree, even between apparently hale individuals, is on the surface of it
explicable on the Mendelian theory ; also we see that, whether we look upon cousin
marriage as producing on the average more than six per cent, of patent evil, or in
the other aspect, that some cousin marriages are as detrimental as brother-sister
marriages, reasons can be found for their all being forbidden by tribal custom or
religious ordinance. But this is after all only to look on one side of the picture,
because the RR characteristic might be a patent good quality suddenly introduced
from outside into a population ; in such a case cousin marriage is distinctly to be
commended, and brother-sister marriage would be more effectual still. In this way
the endogamy of many early communities receives its due sanction. As long as a
species is likely to vary advantageously, endogamy between collaterals of the first
degree will produce 75 p. c. with patent or latent good quality, and between collaterals
of the second degree 62 5 p. c* ; even endogamy of ascendants and descendants may
be advantageous. It is probable that whenever selection is extremely stringent the
relative advantages of endogamy become apparent and are emphasised by tribal
custom. But the Mendelian theory cannot be considered as demonstrated, and if it
were, we could hardly at present apply it to man. We have no means of separating
the DR's from the DD's, short of that experimental breeding which the Mendelians
tell us is the only reliable guide to the gametic constitution. We cannot, however,
afford to bring defective children into the world to test where the endogamous union
will be an advantage, where a failure. The somatic characters of the individual and
of his or her ancestry are at present our sole possible guide to his or her gametic
constitution. From this standpoint we may ask what is the quantitative value of
the cousin in the problem of inheritance ? In predicting the probable offspring of an
individual is the cousinship of more or less importance than the parents' brothers and
sisters ? Is a knowledge of the grandparents' characters of greater value than that of
the cousin ? It will be clear that the cousinship while generally less accessible than
the sibship, is often far more accessible than the grandparentage, or in the case of
orphans than even the parentage ; and for the special purpose of medical diagnosis
* This supposes that endogamy in the first degree is forbidden.
MEASURE OF RESEMBLANCE OF FIRST COUSINS 5
may be of great relevance. The existing state of doubt as to the quantitative value
of cousinship may be illustrated from such a vital problem as that of the hereditary
predisposition to mental disease where some medical authorities would exclude entirely
evidence drawn from the cousinship*, while they retain the inquiry as to the direct
line and as to collaterals in the first degree. As the cousinship often combines many
lines, it requires of course careful handling, but its size and relative accessibility may
be factors which give it equal importance with the grandparentage or the parental
sibships.
2. With the object of throwing light on the value of the record of cousinship, an
inquiry as to the physical and psychological resemblance of first cousins was set on foot
by Karl Pearson some five years ago and a grant obtained from the Government
Grant Committee to assist the investigation. The assistance derived from this source
is here gratefully acknowledged. The plan followed was twofold. Two independent
collections were started. The first part of the investigation was based upon very
general inquiries as to the physical and psychical characteristics of families. At
present about 300 families have supplied very full particulars of ancestors and
collaterals as far as the personal knowledge of the recorders extend. These Family
Records supply the material upon which the bulk of the present paper is based, and
provide sufficient pairs of cousins to give a fair idea of the general intensity of re-
semblance in cousins. The family schedules asked for the following information:
(1) Present Age or Age at Death of each individual.
(2) Ailments in Life.
(3) Cause of Death, if dead.
(4) General Health under the Categories : Very Robust, Robust, Normally
Healthy, Delicate, and Very Delicate.
(5) Ability under the categories :
A. — Mentally Defective. — Capable of holding in the mind only the simplest
facts, and incapable of perceiving or reasoning about the relationship
between facts.
B. — Slow Dull. — Capable of perceiving relationship between facts in some
few fields with long and continuous effort ; but not generally or without
much assistance.
C. — Slow. — Very slow in thought generally, but with time understanding is
reached.
D. — Slow Intelligent. — Slow generally, although possibly more rapid in
certain fields ; quite sure of knowledge when once acquired.
Ex. — Fairly Intelligent. — Ready to grasp, and capable of perceiving facts in
most fields ; capable of understanding without much effort.
* Bucknill and Tuke : Psychological Medicine, 2nd edn. p. 266.
6 ETHEL M. ELDERTON
E... — Distinctly Capable. — A mind quick in perception and in reasoning
rightly about the perceived.
F. — Very Able. — Quite exceptionally able intellectually, as evidenced either
by the person's career or by consensus of opinion of acquaintances.
During a part of the investigation E, and E2 were classed together as E, but
a large number of D, E and E, F entries (i.e. 'Betwixt' entries) occurring,
this category of E was divided as above into Ej and E.,.
(6) Temper under the categories : Sullen Temper, Quick Temper, Even Temper,
Weak Temper (not ' even,' but weak good nature).
(7) Temperament — under three divisions (a) Reserved, Expressive or Betwixt ;
(b) Sympathetic, Callous or Betwixt ; (c) Excitable, Calm or Betwixt.
(8) Success in Life under the categories : Marked Success : An individual who is
not only marked above his family, but above his fellow citizens for achievement in
life. One who has made a name which would find a place in the Dictionary of
National Biography. Prosperous Career: An individual who has advanced beyond his
family level but not necessarily marked among his fellow men. An active successful
life or career. Average Career: An individual who has not fallen below the family
standard of life, whether in profession, trade or craft. Difficult Career : An individual
who has found it difficult to maintain the previous family standard. One who has had
a struggling and unprosperous career. Failure : An individual who has more or less
failed in life ; a bankrupt, or ne'er-do-well ; this letter (F.) may be used to cover the
black sheep of a family.
Considerable care was taken in distributing the schedules* among those likely to
be interested in the investigation and having a sense of responsibility for the frank-
ness and fullness of the information provided. A considerable number of schedules
were returned to the recorders for corrections or additions which were at once
supplied. In less than two per cent, of cases was it needful to reject a schedule
as untrustworthy, or so incomplete as to be useless. Each Family Record contains on
the average particulars of about 40 individuals and, as it is hoped to raise the total
number of records from the present 300 to 1000, we shall then possess an account of
a fairly random sample of the general population of about 40,000 persons.
3. The various types of cousinship distinguished in the schedules are :
(^4) Cousins are sons of two brothers. (B) Cousins are sons of two sisters. (C)
Cousins are sons of a brother and a sister. (D) Cousins are daughters of two brothers.
(E) Cousins are daughters of two sisters. (F) Cousins are daughters of a brother
and sister. (G) Cousins are son and daughter of two brothers. (H) Cousins are son
and daughter of two sisters. (/) One cousin is daughter of a brother, the other is son
of a sister. (K) One cousin is daughter of a sister, the other is son of a brother.
* They provided for information with regard to four generations in the direct line, and three
generations of collaterals.
MEASURE OF RESEMBLANCE OF FIRST CO.USINS 7
A, B and C are types of male, D, E and F are types of female, G, H, I, A' of male
and female cousins. There are thus ten types of simple first cousins. It was con-
sidered desirable to keep these ten types distinct in order to ascertain how far
resemblance was modified by change of sex in descent from a common ancestor.
Special cases of abnormal cousinship in the first degree were not included. All
individuals dealt with were adults. The four characteristics : General Health,
Ability, Temper and Success in Life, providing 5, 7, 4 and 5 categories, admitted
at once of tables of contingency being formed of at least 4x4 groups, and these were
at once reduced by the method of mean square contingency. This was done for all
the ten types of cousinship. In the case of Temperament there were only the
alternatives and the ' Betwixt ' groups. We were thus compelled to use either con-
tingency on a 3 x 3-fold grouping or else assume the material to have a Gaussian
distribution and apply fourfold table divisions. In the latter case the results will vary
somewhat according to the alternative with which the ' Betwixt ' group is associated.
The Temperament results are, however, in our opinion the least reliable of the
series. We believe this to be due to the fact that the ability, success, health and
to some extent the temper of an individual are matters of common knowledge or
repute ; but that temperament as we have classified it is less generally realised.
There is little doubt that some of the recorders had not previously formed a general
estimate of temperament, and have taken that of a particular individual as a standard
to classify other members of the family by. We should not be inclined accordingly
to place much stress on the Temperament results as proving anything beyond the
basal principle that temperament is an inherited charactei\
In the second series of investigations, it was considered that it would not be
without interest to deal with an entirely novel physical character, i.e. novel from the
standpoint of inheritance, and accordingly the hand was selected as easily accessible
and, at any rate for some characters*, capable of fairly accurate measurement. Other
physical characters readily ascertainable were eye and hair colours and general health.
Accordingly a cousin schedule was issued with the directions for measurement noted
below. See Appendix A, p. 21. Much time and energy had already been spent
over an endeavour to reproduce in a cheap manner the eye and hair scales used as
standards in the Biometric Laboratory. Ultimately we had to content ourselves with
an admittedly imperfect chromolithograph of hair coloursf. For the eye scale we used
a hand-painted scale. Miss Mary Beeton kindly painted on a printed blank the
irides of 24 eyes, painting one eye at a time in about 100 copies from a standard
glass eye. To these scales was added a cheap but quite efficient hand-spanner
prepared by the Cambridge Scientific Instrument Company. The scales, spanners,
directions and schedules were circulated in the same manner as the similar material
* R. S. Proc. Vol. 65, pp. 126—151 : "Data for the Problem of Evolution in Man. A First Study
of the Human Hand." By M. A. Whiteley and Karl Pearson,
t Reproduced Biometrika, Vol. v. p. 474.
8 ETHEL M. ELDEKTON
for the Family Measurements of six years ago* : they were loaned to College students,
personal friends of members of the Biometric Laboratory and others. It was, however,
soon obvious that we had miscalculated the ease with which pairs of cousins could be
found and measured. The work went forward extremely slowly, most investigators
sent in only two or three pairs ; and when the question of repeating or verifying
a measurement arose, the delay or even the impossibility of supplementing the data
was much more common than in the case of the Family Records. In fact cousins are
not like brothers and sisters, or pai'ents and offspring, in daily touch with each other;
and at the end of three or four years, we are far from having reached a sufficient
supply of cousin pairs. Hardly indeed have 300 pairs been yet measured. Accord-
ingly this side of the enquiry is incomplete and will only be used as a control series.
We need scarcely say that we shall be very glad indeed to loan spanner and scales to
any reader of this memoir who will undertake to measure pairs of adult cousins.
4. We now turn to the analytical methods by which the material was reduced.
We have already pointed out that contingency was used throughout the whole of the
first series, but that in the case of Temperament the 3 x 3-fold tables were not finely
enough grouped to make contingency thus obtained really comparable with that found
from higher-fold tables. Accordingly the temperament tables were only worked out
by 3 x 3-fold contingency in the case of the three groups : male cousin pairs, female
cousin pairs, and male and female cousin pairs. The fourfold table methodt was used
on the same material in thirty cases, namely the three classes of temperament in the
ten classes of cousins.
The mean value of the degree of resemblance between cousins as found from
40 contingency tables was "271 + '009 with a standard deviation of "083 + "006. The
mean value of the degree of resemblance in temperament as found from 30 tables by
fourfold process was -258 + -014, but the variability in this case was 35 p. c. greater,
the standard deviation being •115 + 'OrO.
The temperament tables worked by contingency with three types of cousins gave
the result -238 ±"010, with the reduced variability indicated by -045 + -007. These
results are collected in Table I. They suffice to show that mean square contingency
methods give more uniform results than the fourfold tables i. But the mean found
from contingency for Health, Ability, Temper and Success does not sensibly differ
from that found for the three divisions of temperament by fourfold tables leading to
the coefficient of correlation. If we combine the results of both methods so as to
* Directions and form of schedule for this case are reproduced in Biometrika, Vol. n. pp. 359-60.
t The fourfold tables were worked for two groupings for the alternatives Excitable or Calm ; the
Betwixts being thrown first into one group and then into the other, the average value of the correlation
coefficient is that given in Table III. In the case of the alternatives Reserved and Exjjressive the Betwixts
were thrown into the Reserved, and in that of the alternatives Sympathetic and Callous into the latter
group. This was done after some consideration and enquiry as to the popular weight of terming an
individual ' reserved ' or ' callous.'
\ The corresponding nine fourfold tables were somewhat erratic and gave a mean of only -19.
MEASURE OF RESEMBLANCE OF FIRST COUSINS
obtain a general average degree of resemblance between the ten types of cousins for
seven characters, we find for the whole seventy tables :
Mean value '267 ± '008. Standard Deviation = -093 + -006. Accordingly we may
conclude that the average degree of resemblance of cousins lies between -25 and "30,
say at '27.
Table I. Mean Results by Different Methods.
Characters
Method
No. of Cases
Mean
Standard
Deviation
Three Phases of Temperament
Ditto
Health
Ability
Temper
Success
Whole Series
Fourfold Division
1 Mean Square Contingency
\ 3 x 3-fold Table
Mean Square Contingency
4 x 4-fold and higher-
fold Tables
' Contingency
Fourfold Division
: ■
1-
40
30
•258±-014
-238+ -010
•271 ±-009
| -267* ± -008
•115±-010
•045 ± -070
•083 + -006
•093 t -005
Diagram of Frequency of Coefficients of Resemblance in Cousins and Brethren.
325" 375 425 475 525 575 625 675
Cousinships Sibships
Mean from grouped values ; ungrouped value = -265.
2
LO
ETHEL M. ELDERTON
The fluctuation is no doubt considerable in our results. But we think that it lies
far more in the difficulty of estimating psychical characters, than in any real variation
in the degree of resemblance. The fluctuation is greatest precisely in those characters
where personal bias and sex bias make the judgment more difficult.
We are now in a position to compare the intensity of resemblance between cousins
with that between brethren. Diagram I, p. 9, shows graphically the distribution of
the degree of resemblance of the 70 cases of cousins in this first series and of
65 cases of brethren, physical and psychical. The cases were numerically distributed
as follows :
Table II.
o
i
o
3
I
I
1
IN
I
1
1
1
I
!
1
1
1
Totals
Cousinsbip
Sibship
2
i
9
10
18
11
10
2
6
•5
1
14-5
2
22
15
8
3
70
65
Mean Cousinship : '27
Mean Sibship : '51
■008. Standard Deviation : -093 ± -005.
•006. Standard Deviation : -068 + -004.
An examination of the graph shows that the cousinship group clusters at 25 and
the sibship group at '5. These may, we think, safely be taken as working values for
cousinship and sibship resemblance for either sex, and we may safely assert that
brethren are on the average twice as closely related as cousins. This halving of the
degree of resemblance corresponding to the fact that normal first cousins have two
common grandparents, whole sibs have four.
It must be noted : (a) that our data for cousins are not drawn from the same
records as those for brothers and sisters. While the Family Records here used for
cousins enable us also to deal with brothers, these have not yet been tabled and
reduced, except in the one instance of Intelligence. Here the adult brothers gave "54
as against the average of three cases of adult male cousins giving '34. Schuster
found -56 from adult Oxford graduates for ability. Pearson found '52 for brothers
at school and Schuster for schoolboys "56. The half of these fraternal values would
be -27, which agrees well with the general cousin average, but not so well with the "34
which is a definitely higher value.
(b) that our data for cousins and sibs are neither from the same records, nor
for the same range of characters. In the case of the sibships 21 values were for
definitely measurable characters, a much more reliable class of material ; while the
consulships of the first series do not present a single measurable character, and only
one definitely physical estimate, that of Health. The characters which are common to
MEASURE OF RESEMBLANCE OF FIRST COUSINS
I I
both series are : Health, Ability and Temper in adult cousins, and in sibships of school
children ; the following table gives the results for three classes :
Table II a.
Type
Intelligence
Health
Temper
Cousinship
Sibship
Cousinship
Sibship
Cousinship
Sibship
Male and Male
Female and Female
Male and Female
■34
•34
•34
•40
•47
•H
•31
•33
•30
■52
•51
•57
• -18
•19
•25
•51
•49
■51
Mean
•34
•46
■31
•53
•20
•50
Mean Cousinship "28. Half Mean Sibship -25.
Treated alone these cases would show a definitely larger degree of l-eseinblance for
the cousinship, than for half the sibship, but this is not borne out for the whole
material. The differences also of the ages of the subjects, adults and school children,
and the methods of recording, by relatives and by school teachers, must also be borne
in mind.
We consider on the basis of this first series that "25 is a good round working
number for the cousinship. This denotes on the assumptions of linear regression and the
equal variability of the cousins, that two cousins of an individual selected from unrelated
stocks (i.e. maternal and paternal cousins) will give the same probable value for the
character of an individual, as a brother of that individual with the same character as
the mean of the cousins*. On the other hand the accuracy of the estimate will not be
so great. In the first case it is o^ v 1 — ('5)2 and in the second case o^ v 1 — (-25)2 — (-25)2,
which measures the variability of the array; these are as 8'6G to 9*35. Thus the
prediction from the brother would be somewhat better than from two mutually
unrelated cousins. It is clear, however, that a knowledge of two such cousins may be
very useful indeed, especially if facts as to the sibship are not forthcoming.
If we turn to other collateral relationships, the avuncular worked out for the eight
possible cases in eye colourt, is, as far as we know, the only one yet published. The
mean value of the eight cases is *265. We should accordingly conclude from this that
for purposes of inheritance a knowledge of the cousin is equally important with a
knowledge of the parental sibships. For example, there is no justification in medical
histories of lunacy for including the facts as to the parents' brothers and sisters and
* Regression equation for 1 on 2 and 3, the latter being independent, is
h, = 'Ji^i 1H + 'J^ h3 = rnh2 + ruh3
<T., 0-3
t Phil. Trans. Vol. 195, A, p. 114.
if o-,
•50 x J {lh + A,), if
12 ETHEL M. ELDERTON
omitting the cousins from the record. On the same ground the marriage of niece or
nephew with uncle or aunt seems to be a marriage of exactly the same degree of
kinship as a marriage between first cousins.
The only grandparental data at present reduced for man* are those for eye-colour
and the eight cases give a mean value of "32 1. This is somewhat higher than the
value ("27) for cousins, and pigmentation data in horses have given an almost equal
value. Still other species show rather smaller intensity, and until further data are
reduced for the case of man, especially for psychical characters, we are not convinced
that the grandparental relationship is definitely more important than the cousinship.
At any rate, even with our present values (-27 as against "32) it will be seen that it is
not reasonable for the purposes of medical or actuarial diagnosis to neglect the cousins,
and make a considerable point as to the grandparental constitution. The grandparent,
the uncle or aunt and the cousin are practically on the same footing with regard to
relationship or intensity of kinship as measured by degree of likeness of character ;
and it seems probable that any scientific marriage enactments would equally allow or
equally forbid marriage between grandparent and grandchild, uncle and niece, aunt and
nephew and between first cousins. This conclusion is reached on the assumption that
the undesirability of marriage depends on the closeness of likeness in the gametic
constitution, and that on the average the i-esemblance of the somatic characters may
be taken as a measure of the average gametic resemblance between any two classes.
5. We now turn to the details of Table III.
We first ask whether there is any sensible difference between the intensity of
inheritance in males and females. We note that the probable error of any individual
result runs from about -02 for cousins of same sex to "03 for cousins of different sexes.
Our table shows us that the average for pairs of male cousins is the same as that for
pairs of female cousins, i.e. "26. If we could lay any stress on the difference "02, we
should assert that cousins of different sexes were more alike than cousins of the same
sex. But we certainly cannot, and thus, as far as our data go, we can only conclude
that difference of sex makes no difference in degree of likeness.
In the next place we may consider whether type of cousinship makes any differ-
ence in the intensity of resemblance. Our mean values for all the characters range
from '22 to -31 according to the type, and it might be thought that this offered
sufficient range to answer the question. As defining the types there are two considera-
tions to be noted, (i) a difference of sex in either generation, parental or cousinal, and
(ii) a change of sex in descent. Neglecting the first we have :
No change of sex in descent in : A (SO), E (-27) or K (-29) ;
One change of sex in descent in : C (-23), F (-28), G (-24) or H ('31) ;
Two changes of sex in descent in : B ('24), D ('22), or / ('29).
* The Family .Records of the present series provide unreduced material for seven characters, and this
will shortly be dealt with.
t Phil. Trans. Vol. 195, A, p. 115.
MEASURE OF RESEMBLANCE OF FIRST COUSINS
13
The means for the three groups are '287, -265 and "250 respectively. It may
possibly be therefore that change of sex slightly weakens the intensity of inheritance
in the stock. If we turn to the first consideration the change of sex in the same
generation, the connected parents are of the same sex and the cousins of the same sex
in A and A. but in K the connected parents are of different sexes and the cousins
o\' different sexes. A" is not. however, the least of the three. In C and /''there is a
Table III. General Results of First Scries. Characters of Cousins.
Type of Cousins
Health
Intelligence
Success
Temper
Temperament
Reserved or
Expressive
Sympathetic
or Callous
Excitable or
Calm
Method -►
Male. Type A
„ B
„ c
M.S.C.
M. S. C.
M.S.C.
M.S.C.
F. T.
M.S.C.
F. T.
M.S.C.
F. T.
M.S.C.
•34
■32
•26
•-tl
•30
•32
•24
•15
•19
•23
•16
•15
■38
•06
•23
1 -20
•31
•36
•30
h
■21
•36
•17
!«
•30
•24
■23
Mean
•31
•34
•19
•18
•22
(■20)
■32
(•24)
•25
(•34)
•26
Female. Type D
„ F
•51
•24
•23
•34
■38
•31
•16
•27
■35
•14
•24
•18
•21
■30
•26
},
•03
•42
•37
1 -20
■15
•04
•23
|,
•22
•27
•28
Mean
•33
•34
•26
■19
•26
(•19)
•27
(■20)
■14
(•22)
•26
Male & Female. Tvpe G
„ H
„ I
•23
•32
•29
•37
•36
•27
■38
■34
•19
•33
•27
•26
■20
■30
•24
•25
•23
•28
•42
1,4
I
•52
■44
•19
■24
■24
14
■16
■IS
-•28
•24
•31
•29
■29
Mean
•30
•34
•26
•25
•29
(•24)
•34
(•24)
•18
(•28)
•28
General Means
•31
•34
•24
•21
•26
(•21)
•32
(■2.".)
•19
(•28)
■265
M. S. C. = Mean Square Contingency. F. T. = Fourfold Table,
difference of sex in the connecting parents, but not in cousins ; in G and H, there is
no difference in the connecting parents but one in the cousins. We might therefore
expect no difference in the four values. But we find C and G contrasted in magnitude
with F and H. In cither words two males related by a male and female go with a male
and female related by two males ; and again two females related by a male and female go
with a male and female related by two females. We can throw no light on this point,
and it may only be a strange result of random sampling. In the third group there are
no sex differences in the types B and D for either parental or cousinal generations.
14 ETHEL M. ELDERTON
For / there are such changes for both generations ; and yet / is larger than B and D.
We must thus consider that a difference of sex in the same generation makes no
difference in the intensity of resemblance so far as our present data go. Accordingly,
if change of sex in descent does to some extent weaken inheritance*, it does not
appear connected with sex differences in the same generation. The differences noted
are, however, too slender and the whole system of values too fluctuating to build up
any hypothesis as to sex influence in heredity.
If we now turn to the separate characters, and compare irrespective of cousin type
the general means of each, we find an even wider range of results ('19 to '34). We
attribute this only in part to real differences in the intensity of resemblance ; we
consider it more due to (a) difficulties of estimating some of the characters dealt
with, especially as in the case of cousins they are usually not in daily contact with
each other; (b) differences of method employed, and the assumption that temperament
follows a normal distribution of frequency.
Accordingly we shall draw no conclusions as to divergences in resemblance,
believing our data may be relied upon to give a " general average resemblance " of
cousins, but cannot be pressed beyond such a result to discriminate between individual
of character.
6. We now turn to the results of the second series of quantitative measure-
ments. These measurements as we have already noted are far from complete. They
give for the four measurements 107 pairs of female cousins, 34 pairs of male cousins,
and 111 pairs of male and female cousins, the two first sets giving 214 and 68 pairs in
the symmetrical table.
The following table gives the statistical constants of the series of measurements.
It will be observed that we have two series for each sex, but not all the individuals in
each series are different.
This table shows at once considerable irregularities, which may be due to the
paucity of data, or to the defective handling of the spanners. While the finger joint
measurements give a sex-ratio for the absolute lengths = '91, very nearly the usual
11/12 of stature and of bone measurements in man, the widths of hand and wrist
(involving a good deal more care in determination and allowing of more personal
equation) give ratios of about H'5/12. For these also the man is both absolutely and
relatively more variable than the woman. For the joint measurements the woman is
equally variable absolutely with the man, and relatively more variable. It seems
improbable that this equal absolute variability is correct. It is not true for the
majority of bone measurements in man and woman. It is further to be noted that in
the Male and Female Cousins series, where there was a much larger return of measure-
* The influence of change of sex has been very elusive ; it would appear to have some bearing on the
inheritance of eye-colour in man (Biometrika, Vol. II. pp. 237-40), but we have failed to find it in coat-colour
in horses (Ibid. Vol. n. pp. 229-34). It is doubtfully significant in the cases of coat-colour of Greyhounds
(Ibid. Vol. in. pp. 257-8), and of Shorthorns (Ibid. Vol. iv. pp. 449-51).
MEASURE OF RESEMBLANCE OF FIRST COUSINS
15
Hunts made by male students than in the case of the Female and Female Cousins
series, the absolute variabilities of the women are in every case less than in the
latter series. In a certain number of cases it was actually found that the user of the
hand-spanner had read from the sliding edge and not from the index point, but the
difference amounting to about 20 mm. was obvious on the face of the measurements
and at once allowed for by measuring the particular hand-spanner which had been
used. It is believed that no residual error has crept in in this manner, but the
point will be again dealt with below.
Table IV. Statistical Constants of Measurements of Hand in Man and Woman,
Sez
Series
Width of Hand
Width of Wrist
Joint,
Index Finger
Joint,
Little Finger
M.
S. D.
C. of V.
M.
S. D.
C.ofV.
M.
S. D.
C.ofV.
M.
S. D.
C.ofV.
Male
Male with Male
Male with Female
83-2
82-1
6-20
5-77
745
7-03
58-2
57-9
3 90
4-17
6-70
7-20
62-5
62 0
2-87
3-23
4-59
5-21
51-0
50-9
2-72
301
5-33
591
Mean
82-6
5-98
7-24
58-0
4-04
7-00
62-2
3-05
4-90
51-0
2-87
5-63
Female
Female with Female
Female with Male
72-5
71-4
4-79
3-64
6-61
5-10
51-8
51 0
3-38
2-78
6-53
5-45
57'2
56-3
3-47
3-06
6-07
5-44
46-4
46-1
2-94
2-70
6-34
5-86
"
Mean
72-0
4-21
5-84
51-4
3-08
5-99
56-8
3-26
5-74
463
2-82
6 09
Sex Ratio, Male and Female
•87
•70
•81
•89
•76
•86
•91
1-07
1-17
•91
■98
1 -08
The measurements are in mm.
The correlations as found by the product moment method without grouping are
given in Table V. Now if we looked simply at the general mean 336 of all 12 results,
we might conclude that the intelligence and health characters of our first series had
given us the more reliable results, the temperament and temper being more difficult
of estimate, and thence conclude that the average resemblance of cousinship was I .",.
Table V. Correlation of Measured Characters in Pairs of Cousins.
Character
Male and Male
Male and Female
Female and Female
Means
Width of Hand
Width of Wrist
Joint, Index Finger
Joint, Little Finger
■3.3
•17
•19
•29
•21
•26
■34
•37
•40
•43
•49
•56
•314
■286
•340
•404
Means -245
•295
•470
•336
16 ETHEL M. ELDERTON
This view might be confirmed possibly by noting that the less easy measurements,
those on hand and wrist, gave lower results than the joint measurements. But
on further inspection of the table we notice that it is the female-female series which
diverges so much from our previous results. The eight cases in which a male was one
of the pair, and presumably worked the spanner, give a mean of -270, agreeing
excellently with the -267 of our much larger first series. It is the pairs of female
cousins, with their excessive variabilities, which give an intensity of resemblance
equal to that of a sibship, and raising the average from -270 to "336.
To test the matter further the following steps were taken. A formula has been
given by Pearson* which is based on normal distribution of frequency and gives the
correlation coefficient in terms of the sum, S(x — y), of the positive differences of
correlated variates which have the same mean and s.D. Now this is precisely the
case of the 106 cousin pairs if we treat them as a symmetrical distribution of 212
pairs. The formula is :
,.-1 y{s(x-y)Y
Applied to the data for the joint of the little finger in pairs of female cousins, we find
r = '5578, while found by the product moment method the answer is "5579, a very
close agreement. But it will be clear that if the measurer had a personal equation of
the nature of a constant error for each pair, it would drop out in the difference x — y
for that pair. Hence the formula above is convenient to use when such an error for
the individual pair is suspected to exist, and the variability cr can be found from
other considerations. If in this particular case we adopt : (a) the standard deviation
of the women in the series of male and female cousins, (b) the standard deviation
found for the women on the assumption that the coefficient of variation for the women
ought to be (what it usually is) practically the same as for the men, i.e. if we take the
two values 2701 and 2-474, we find that the above formula gives r=-48 and "38
respectively. Thus indicating some considerable reduction from the value -56. It is
therefore possible that an adding or subtracting of a constant difference in some of
the measurements is the source of the exaggerated values of the female cousins
resemblance. Such an error would not only have exaggerated the standard deviation,
but it would have resulted at once, if a wrong correction had been applied to the
measurements of those helpers who read at the edge, and not at the index point, of
the spanner. It is believed that no spanners were removed from the numbered boxes
until the measurements had been corrected ; but the doubt, however slight, to those
who had the control of the instruments, is sufficient to. make it needful to repeat as
soon as possible the whole series of measurements on female-female cousins.
We are able to use this second series as a control series also for the characters,
hair colour, eye colour and general health. The method used was that of contingency
* "On further methods of determining Correlation," Drapers' Research Memoirs, Biometric Series IV.
p. 4 et seq. (Dulau it Co., Soho Square).
MEASURE OF RESEMBLANf 'E OK FIRST COUSINS
17
but it must be remembered that the series were short, i.e. treated as symmetrical
tables we had only 68, and 218 entries, and for male and female cousins 113. The
results given in the following- table were reached.
Table VI. Non-quantitative Characters, Second Scries.
Character Male and Male
Male and Female
Female and Female
Means
Health
Eye Colour
Hair Colour
•38
■44
•34
■29
•48
•26
•18
•38
•2G
•282
•434
•286
Means
■3S6
•343
■273
■33 1
It will be seen at once that (i) the preponderating intensity of pairs of female
cousins no longer exists, (ii) the eye colour values are, however, very high, in one case
at least approaching the intensity of the resemblance of siblings, and (iii) the generally
higher value obtained in the case of the measurable characters of the same series is
maintained. We have already noted that eye and hair scales were used in these
observations, 24 eye and 24 hair tints being given. There were 6 categories in the
Health graduation, but the " Very Delicate " and the " Very Robust " categories were
only very slightly represented, so that for Health merely 3 x 3-fold contingency tables, —
" Robust," " Normally Healthy," " Delicate " seemed possible. For eye colour the 24
eve tints were first classed as " Pure Blue," " Blue with some orange," "Pure Grey,"
" Grey with some orange," " Hazel-Green," " Hazel-Brown," " Brown " ; the two greys
were then clubbed together, as also the two hazels to form a 5 x 5-fold table for
contingency. The 7 x 7-fold table seemed far too fine for the numbers, 68, involved
in the male and male cousin tables, and it was desirable to treat all three tables alike.
The 24 tints of the hair scale were first grouped into : " Very Dark," " Dark-Brown,"
" Brown," " Light-Brown," " Fair," " Red." But for the male data only a single
"fair" and a single "red" occurred and only three "browns." Accordingly the 2nd
and 3rd categories and also the 5th and 6th were grouped together and a 4 x 4-fold
table used for the contingency of hair-colour. As samples, the Eye and Hair colour
tables for pairs of female cousins are given in Appendix B as Tables LXXX and
LXXXI. Now while we frankly admit that this Second Series, whether of
measurable or pigmentation characters, has a much too inadequate frequency to be
conclusive, still its drift is undoubtedly to confirm the view, that the average
resemblance of cousins is higher than that given by the Family Record results. It
approaches nearer the value indicated by the more precise of the " Record " characters,
and the more accurate of the hand measurements. The numbers in the first series are
large as compared with those of the second, and the second series also involves several
18 ETHEL M. ELDERTON
points of doubt and difficulty ; for this reason we have not yet modified the general
average of the First Series by including these higher results. But it is conceivable
that we may have to raise the general measure of resemblance of cousins from "28 to
•33, when other large series already observed have been tabulated and reduced.
7. One further point may be finally touched upon, namely the inheritance of
disease. We cannot in the least hope here for accurate numerical estimates, but the
data of our first series may suffice to show that cousins are of value even from the
standpoint of medical diagnosis. The difficulties of accurate determination are as
follows :
((() While on the schedules the record of brothers and sisters, of children, of parents
and of grandparents is fairly complete, that of cousins must necessarily be defective. It
is quite possible- — nay not infrequent— to have more than 50 first cousins. And while
one or two recorders actually were patient enough to enter details of a cousinship
as large or even larger than this, the bulk of recorders contented themselves with
entering a much more limited number, 10 to 12, and thus we have the first limitation;
our cousins, as the recorders themselves state, are a selection. It is probable, also, that
the selection has been made more frequently of living than of dead cousins, and more
frequently of accessible than possibly inaccessible cousins ; thus the individuals
suffering from phthisis or insanity, or having died from these diseases, may without
direct intention to deceive have been more frequently omitted than in the case of
relatives all of whom were included.
(b) The cousins in our family record schedules are those of the subject. In order
to get full ancestral information a young adult has been very often taken as the subject
and the cousins belong accordingly to the third generation, and are themselves often
young adults. It follows accordingly that their medical history is in many cases
incomplete. They have not passed wholly through the danger zone in the case of
either tuberculosis or insanity.
In the case of tuberculosis, we have for instance among males only 206 tuberculous
out of 2990 individuals, and among females only 205 out of 3242, whereas 10 p.c.
would probably be affected if we had the full record.
In our records for example there are in the case of women 130 cases of individuals
classed as cousins with some form of brain disease or mental defect*. These
130 individuals have 6 insane and 124 sane cousins. If in the remainder of their
lives 4 persons out of those 124 sane cousins were to suffer from some form of brain
attack, then our table would be as follows :
* "Insanity" for the purpose of this investigation has been taken to include the neuroses: confirmed
alcoholism and marked hysteria. These were not included by Heron in using Pearson's Family Records
(Eugenics Laboratory Pali/iratiiut.i, n. p. 33). Its use here approaches "want of mental balance."
MEASURE OF RESEMBLANCE OF FIRST COUSINS
First Female Cousin
I
L9
Insane Sane
Totals
Insane
Sane
14 120
120 2996
134
3116
Totals
134
3116
3250
instead of the actual
First Female Cousin
Insane Sane
Totals
Insane
Sane
i
6 124
124 2996
130
3120
Totals
130 3120
3250
The fourfold table method gives the correlation of the first table about -33 and that
of the second "03. Now it is not suggested that four additional cases of insanity are
what we have to expect in the case of 124 persons chiefly young adults of insane
stock. What we wish to point out is that with a disease so relatively rare as this,
the transference when the record is completed of comparatively few individuals from
the sane to the insane category is sufficient to raise the intensity of resemblance to a
value quite equal to that which we have found for other characters in cousins.
The following are the results reached for insanity and tuberculosis :
Table VII. Inheritance of Pathological Condition in Cousins with
incomplete Record.
Male Cousins Female Cousins
Male and Female Cousins
Means
Insanity -18
Tuberculosis ! -07
•03
■12
■08
•19
•10
•13
Means -12
•08
•13
•11
In all six series — and they number in each case about 3000 pairs — we have a
positive relationship, and the value is definitely significant in all cases but possibly
that of insanity in female cousins. Yet this, owing to the fact that a considerable
amount of insanity in the case of women is connected with change of life, is precisely
what we might have anticipated considering the ages of our cousins. The fact also
that insanity has a later average incidence than tuberculosis may explain why the
average value for tuberculosis is higlier than for insanity. We should conclude that
so far as our data go they show that the tendency to both insanity and tuberculosis
runs in stocks, and that with the incompleteness of the record there is no reason to
20 ETHEL M. ELDERTON
suggest that disease tendencies are not inherited at the same rate as physical and
psychological characters in cousins.
8. General Conclusions. Our memoir has dealt with two series of cousin
records. The quantitative measure of the resemblance of cousins is of great importance
— not only on account of its bearing on eugenic marriages, but because cousins form
often the principal living record to assist medical diagnosis. Its determination,
however, presents considerable difficulties. It is not hard to collect data as to the
characters of cousins, when these characters can be judged without the actual presence
of the cousins. This was done in our first series. But when we come to the
quantitative measurement of cousins our experience has been unfavourable to the
rapid accumulation of extensive material. The passing from brethren to cousins — -
although the latter are a far wider group — has more than trebled the difficulty of
obtaining measurements. Further our choice of the hand as the organ to be dealt with
has possibly led to difficulty, as the treatment and use of the spanner needed more care
than a simple measuring tape. It was possible to explain and illustrate the use of the
spanner to all the male students to whom it was loaned, but in the case of women
helpers we had often to trust to written directions. This may be the source of the
high values found for the resemblance of women cousins, but we confess frankly that
we are not satisfied that it is so, and we must await the reduction of further material
before settling this point. If we turn to the 70 cases dealt with on the basis of our
first series, we find an average resemblance of about -27, which tallies with the average
found from the eight quantitative series involving male cousins in our second
investigation. If this value be confirmed we should say that cousins have as much
significance as the parental brothers and sisters. On the other hand an examination
of our table shows that what may be treated as the more easily judged and reliable
results, show a rather higher value than '27, approximating rather to the '33 of the
grand parental resemblance. The pigmentation results of the second series tend to
confirm this view.
We should conclude accordingly from the present results that for the purposes of
eugenics cousins must be classed as equally important with uncles and aunts, and that
they may eventually turn out to be as important as grandparents. For practical
purposes it would hardly seem possible in the matter of marriage restrictions based
solely on the gametic resemblance judged by somatic characters, to differentiate
between the three classes. This equality of resemblance which may appear at first
sight paradoxical will be confirmed for uncles and aunts in a forthcoming memoir.
Its physiological bearing appears to us of fundamental importance as indicating that
a determinantal theory of heredity, emphasising alternate inheritance, must take pre-
cedence of any theory of simple blending for the bulk of the characters here dealt with.
We do not consider that our data show any difference between the inheritance of
physical, psychical and pathological characters, which could not be accounted for by
(a) the difficulty of appreciating temperament, and (b) the incompleteness of the
cousin record.
APPENDIX A.
HEREDITARY RESEMBLANCE OF FIRST COUSINS*.
I. OBJECT OF MEASUREMENTS AND GENERAL INSTRUCTIONS.
I.— The present state of our knowledge of the laws of inheritance in man may
be summed up as follows : —
We know well for a variety of organs direct inheritance from parent to offspring,
and the collateral relationship between brothers and sisters. We have less complete,
but still valuable data for the direct line in the case of grandparent and great-grand-
parents, and for the collateral line in the case of uncles and aunts. To supplement
our knowledge, one of the most urgent problems is the determination of the degree
of resemblance between cousins. It is with a view of solving this problem of cousin
relationship that I appeal for cooperative observations and issue the present paper
and schedules.
II. — For the purposes of the present investigation we are to understand by the
word cousin ;
(i) Full blood First Cousins, that is children of two whole (not half) brothers,
of two whole (not half) sisters, or of a whole (not half) sister and brother. Such
cousins are to have one and only one grandparental pair in common, and we term
them normal cousins.
(ii) " Abnormal " first cousins are to be excluded.
It may happen that two brothers of one family have married to two sisters
of another family, or that a sister and brother of one family have married a brother
and sister of a second. The issue of such marriages are "doubly" first cousins having
all their grandparents in common. Again, a brother and sister in one family might
marry an aunt and a nephew in a second family, or again, might marry a woman
and a man who are cousins in a second family, or, two brothers may marry two half
sisters in a second family. Indeed cases of abnormal cousinship occur in which the
abnormal cousins have 1, 2, 3 or 4 common grandparents. All such cases are excluded
from the present investigation, which is concerned only with normal cousinship as
defined under (i).
(iii) Normal cousins for the purpose of this investigation must be between 18
and 45 years of age.
* Issued by Professor Pearson, 1902 and onwards.
22 ETHEL M. ELDERTON
We cannot for a longer period consider the eye and hair colour to remain even
approximately constant. With a shorter period we might fail to obtain sufficient
material for statistical purposes.
III. — There are ten kinds of normal first cousins. Let A and B stand for the two
cousins, thus : —
Tiro (A and B may be sons of two brothers.
Male - A and B may be sons of two sisters.
Cousins\A and B may be sons of a sister and brother.
Two (A and B may be daughters of two brothers.
Female-, A and B may be daughters of two sisters.
( husins\A and B may be daughters of a brother and sister.
Male I A and B may be son and daughter of two brothers.
and ) A and B may be son and daughter of two sisters.
Female \A may be the son of a sister and B the daughter of a brother.
Cousins [A may be the son of a brother and B the daughter of a sister.
In this classification in the last group of " male and female cousins," A is taken
as the male and B as the female cousin. But in the actual schedule provision is
made for the case where the observer has taken A for the female and B for the
male cousin.
The observer, after entering his or her own name, should fill in the names of
the cousins A and B and their sex by putting a cross under male or female. Next,
under type of cousinship, a cross should be put in the last column against the special
type of the two cousins observed. This is very important, because we have reason
to believe from the grandparental and avuncular relationships that the degree of
resemblance varies a good deal with the type.
IV. — Any individual cousin A may be dealt with in any number of cases, but
it is not desirable to compare one cousin A with more than four other cousins who
are brothers and sisters to each other, and of these, not more than two should be
of one sex. Subject to this limitation A may appear, or A's brothers and sisters,
in any number of cousinships. A fresh schedule should be used for each such cousin-
ship. It is not, however, necessary to fill in on these additional schedules all the
measurements and characters. The name and sex only of the repeated cousin, and
the type of cousinship, need to be inserted. A cross reference to the number of the
observer's series in which the cousin is fully recorded will then suffice. A blank
is left for this reference under the name and sex of cousin. For example, if P, Q,
R, S be children of four different brothers and sisters, we first fill up a schedule
of P and Q, then one of R referring to the schedule containing P ; and another
schedule referring to the schedule containing R and to the one containing Q, but
giving the type of Q and R cousinship. Then we measure S and refer to P, and
MEASURE OF RESEMBLANCE OF FIRST COUSINS 23
finally two more schedules give merely the names of S and Q, and <S and R with
their types of cousinship, and refer to the proper schedules for the observations on S,
Q and R. Or again, the observer may till in one schedule for himself or herself,
and then with simple reference to the number of that schedule and the type of
cousinship, till in separate papers for twenty or thirty of his or her cousins. Then
another series of schedule papers may be filled with simple references to the indi-
viduals among these twenty or thirty persons (without repeating their measurements)
who are cousins among themselves apart from their relationship to the observer. In
each case the type of cousinship must he marked on the new schedule paper.
V. — Directions for recording Observations.
(1) Hair Colour.
On the hair-colour scale in the box, pick out the number of the hair corresponding
most nearly to the colour under observation. If the hair considered falls between
two tints, so exactly that you cannot say that it is nearer to the one than the other,
give both tints, thus 5 — 6. If the hair has turned grey before 45, say so ; and
if the hair is of tint distinctly not on the scale, fasten a very small sample, sufficient
to show colour, on the data sheet with the border of a sheet of postage stamp, or
other strip of gummed paper.
(2) Eye Colour.
In judging eye colour, first fix the attention on the amount of orange-brown
pigment in the iris. If there be no orange-brown pigment, the eye is (l) Dark Blue,
(2) Blue, (3) Light Blue, (4) Light Grey. With hardly visible amounts of the orange-
brown pigment we have next (5) Blue-green, (G) Dark Grey, (7) Hazel. Lastly,
with clearly marked orange-brown pigment, we have (8) Light Brown, (9) Brown,
(10) Dark Brown, (11) Very Dark Brown, (12) "Black." Samples of these eye types
are given on the eye-colour scale. Look at the eye with the light upon it from a
distance of about 1 8 inches and compare it with the scale. If the eye falls between
two types on the scale, give the numbers of both types ; if it agrees fairly well with
any type, give the number of that single type only. Thus G — 7 would mean that the
eye in question fell between 6 and 7 of the scale, but 7 would signify that it was
closer to the 7 than to the 6 of the scale.
(3) Health.
Place a cross against the category under which the general health falls.
(4) Measurements of Hand.
These are to be made with the hand-spanner which will be found in the box.
All the readings are to be taken to the nearest mark on the scale, and the observer
need not give fractions of the units on the scale, if the length falls between two
marks. If in any case the observer finds it quite impossible to determine which is
the nearer mark, then give both units, e.g., 34 — 35.
24
ETHEL M. ELDERTON
Fig. (i).
Self-measurement of the left hand by means of the hand-spanner.
(i) Width of Wrist. See Figure (i).
Feel for and satisfy yourself as to the positions of the bony protuberances on
either side of the main joint of the wrist. They
are the sides of the ends of the two bones of
the forearm. The space between the outer sides
of these has to be measured with tbe spanner.
Hold the spanner in the right hand, resting its
fixed jaw against tbe breast, and manipulate
the movable jaw with the spare fingers of the
right hand. Lay the left wrist back upwards
between the jaws of the spanner, so that the
bony protuberances come against the jaws.
Close the jaws with gentle pressure and clamp
the movable jaw with the clamping nut under-
neath. Repeat this at least once, and if time
will allow twice, taking the reading each time
and entering it on the schedule. Do not be
surprised if your measurements are not exactly
the same. Only suspect something is wrong,
difference in your results. If this be so, test carefully again. Do not fill in column
marked "mean," but leave this to those who have to reduce the observations.
(ii) Width of Hand. (Left hand, as before.) See Figure (ii).
Feel for and satisfy yourself as to the positions of the outer sides of the knuckles,
the one side being formed at the joint at
the base of the little finger, the other at
that of the forefinger. The hand is to be
placed with the fingers close together, with
the palm upwards, and all the knuckles
t muLiug the spanner. Measure the width
between the outer sides with the spanner
held with the fixed jaw against the breast
and the scale horizontally upwards. Bring
the movable jaw without pressure against
the knuckle at the base of the forefinger.
Clamp and read the scale. N.B. — Take
care to make two or three trials.
you
find
two units
MEASUIiF OF ItFSFMRLAXOE OF FIRST COUSINS
25
(iii) Length of First Joint of Index and Little Fingei
See Figure (iii).
Close the fist (thumb outside) and
thumb uppermost and spanner hori-
zontal— the lengths from knuckle to
first joint of (a) the Index Finger ;
(b) the Little Finger. The outside of
first joint is put against the tixed jaw
of the spanner, and the movable jaw
is brought against the outside of the
knuckle with gentle pressure. Clamp
and read as before, making two or three
trials.
All the measurements should be
made with care. The above instruc-
tions are intended for self-measurement,
but it is easy for one observer to measure
both of a pair of cousins, or him or herself and then the cousin
Fig. (iii).
VI. — In case of any difficulty, please apply at once to Professor Karl Pearson,
University College, London, W.C. The Box and papers should not be kept longer
than a month, unless the observer finds it possible to undertake a large series of
cousins. About a thousand pairs of cousins of each type, 10,000 in all, will be
required. Hence every co-operator will appreciate the necessity for rapid circulation
of the boxes, of which only a limited number can be provided. •
The name of the observer and address should always be given, in case it is
necessary to ask questions as to any special measurement or observation. The
cousins, if it be preferred, may be simply denoted by the initials of their christian
and surnames, as these will suffice for the observer to identify them*.
* Spanners and schedules are still (November, 1907) being issued,
measurements will be gratefully accepted.
id help in further cousin
26
ETHEL M. ELDERTON
II. SCHEDULE.
Kindly make no attempt to fill this Schedule in until the General Instructions
have been carefully read through.
Hereditary Resemblance of First Cousins.
Ages between 18 and 45.
Observer
Name
Number in
Observer's Series
Number in
whole Series*
* Leave this space blank.
against Sex of Cousin.)
1 Male
Female
Male Female
Cousin B.
Type of Cousinship. (Place cross against type in right-hand column.
Male Cousins
(i) A and B are sons of two brothers
(ii) A and B are sons of two sisters
r A is son of brother, B is son of sister
1 A is son of sister, B is son of brother
Female Cousins
(iv) A and B are daughters of two brothers
(v) A and B are daughters of two sisters .. .
f A is daughter of brother, B is daughter of sister
^Tl' \ A is daughter of sister, B is daughter of brother
Male and
Female Cousins
f A is daughter, B is son of two brothers
^V11' \ A is son, B is daughter of two brothers
. J A is daughter, B is sou of two sisters
\VU1' [ A is son, B is daughter of two sisters
f A is daughter of a brother, B is son of a sister
^'X' t A is son of a sister, B is daughter of a brother
1 A is daughter of a sister, B is son of a brother
^X' 1 A is son of a brother, B is daughter of a sister
A's measurements are already given on Schedule No.
B's measurements are already given on Schedule No.
only to be used if A and B have already been scheduled for other pairs of cousinships,
MEASURE OF RESEMBLANCE OF FIRST COUSINS
27
(1) Hair-Colour.
Insert number of nearest tint on hair-colour scale
(2) Eye-Colour.
Insert number of nearest tint on eye-colour scale
A
B
A
B
(3) Health.
Place a cross against the category
which seems best to describe A's
general health and a second in
the last column for B's health.
(4) Measurements of Left Hand.
To be made with the hand-spanner as described in the General Instructions.
A
B
Very Robust
Robust
Normally Healthy
Rather Delicate
Delicate ...
Very Delicate ...
Measurement
A
B
1st
Trial
•2nd
Trial
3rd
Trial
Mean
1st
Trial
•2nd
Trial
3rd
Trial
Mean*
(i) Width of Wrist
(ii) Width of Hand
(iii) Length of First Joint, Index Finger
(iv) Length of First Joint, Little Finger
* This column is to be left blank.
To ensure accuracy it is desirable that two or three trials should be made of these measurements,
if they are not taken by an independent observer who has measured already a considerable number
of pairs.
Kindly return this Schedule when tilled in to Professor Karl Pearson, University College,
London, W.C.
APPENDIX B.
TABLES OF DATA.
Health. Male Cousins.
Table I. Type A.
First Male Cousin
Very
Robust
Robust
Normally
Healthy
Rather
Delicate
Delicate
Totals
Very Robust
"
21
5
2
3
31
a
3
Robust
21
64
77
5
22
189
1
Normally Healthy
5
77
206
17
79
384
1
Rather Delicate
2
5
17
-
2
26
Delicate
3
22
79
2
26
132
Totals
31
189
384
26
132
762
Table II. Type B.
First Male Cousin
Table III. Type C.
First Male Cousin
V. R.
R.
N. H.
R. D.
D.
Totals
o
U
1
-a
o
1
V. R.
R.
N. H.
R. D.
D.
Totals
V. R.
30
31
28
1
5
95
V. R.
18
52
45
2
18
135
o
R.
31
98
77
5
2.3
236
R.
52
156
136
13
28
385
1
N. H.
28
77
196
13
35
349
N. H.
45
136
314
15
70
580
o
02
R. D.
1
5
13
-
2
21
R. D.
2
13
15
4
2
36
D.
5
25
35
2
14
81
D.
18
28
70
2
28
146
Totals
95
236
349
21
81
782
Totals
135
385
580
36
146
1282
MEASURE OF RESEMBLANCE OF FIRST COUSINS
29
Health. Female ( Iousins
Table IV. Type />.
First Female Cousin
Table V. Type E.
First Female Cousin
V. B.
B.
N.H.
B. D.
D.
Totals
15
Second Female Cousin
V. B.
B.
N.H.
B. D.
D.
Totals
V. E.
24
7
10
-
4
V. B.
-
(i
46
15
-
6
27
■5
0
c
R.
7
44
99
5
is
203
B.
6
60
1
14-
127
Is
a
X. H.
10
99
2 7 1'
9
127
517
N.H.
15
CO
200
4
78
357
I
B. D.
-
5
9
8
3
25
B. D.
-
1
4
-
-
5
D.
4
48
127
3
74
256
D.
6
14
78
-
22
120
Totals
45
203
517
25
256
1046
Totals
27
127
357
5
120
636
Table VI. Type F.
First F
-male
Cousii
V. E.
E.
N.H.
B. D.
D.
Totals
V. E.
8
31
22
-
14
75
1
E.
31
66
123
1
66
287
1
N.H.
22
123
354
17
136
652
—
a
z
I
X
B. D.
-
1
17
-
2
20
D.
14
66
136
2
58
276
Totals
75
287
652
20
276
1310
30
ETHEL M. ELDERTON
Health. Male and Female Cousins.
Table VII. Type G.
Male Cousin
Table VIII. Type H.
Male Cousin
V. R.
R.
N. H.
R. D.
D.
Totals
3
O
a
V. R.
R.
N. H.
22
R. D.
D.
Totals
V. R.
4
20
8
12
22
5
27
82
2
29
145
37
V. R.
18
6
-
2
48
.s
R.
10
66
85
200
R.
19
52
45
2
21
130
o
o
"3
N. H.
15
86
231
436
N.H.
27
63
163
14
58
325
b
R. D.
-
7
10
21
190
884
R. D.
1
3
4
-
1
9
D.
10
57
92
2
D.
16
31
84
1
33
165
Totals
39
236
126
38
Totals
SI
155
318
17
106
677
Table IX. Type I.
Male Cousin
Table X. Type K.
Male Cousin
V. R.
R.
N.H.
R. D.
D.
Totals
a
o
O
c
V. R.
R.
N.H.
R. D.
D.
Totals
V. R.
8
8
13
-
6
35
V. R.
14
13
9
-
5
41
p
R.
18
55
67
0
26
168
R.
25
46
41
2
16
130
o
O
N.H.
17
82
217
13
38
367
N.H.
11
95
219
18
47
390
i
R. D.
-
4
9
4
1
18
R. D.
-
-
5
2
-
'
D.
7
33
55
2
18
115
D.
17
34
81
1
24
157
Totals
50
182
361
21
89
703
Totals
67
188
355
23
92
725
MEASURE OF RESEMBLANCE OF FIRST COUSINS
31
Intelligence. Male Cousins.
Table XL Type A.
First Male Cousin
F&E2
E
E, & D
C,B*A
Totals
r & E,
14
42
36
5
97
E
42
208
:,:,
14
319
E, AD
36
55
L08
20
219
C.B&A
5
14
20
18
57
Totals
97
319
219
57
692
Table XII. Type B.
First Male Cousin
F&ES
E
Ej &D
C,B&A
Totals
F &E„
26
47
31
3
107
E
17 246
57
23
373
E, & D
31
57
54
19
161
C.B&A
3
23
19
6
51
Totals
107
373
161
51
692
Table XIII. Type C.
First Male Cousin
F&E2
E
E, &D
C.B&A
Totals
-.
F & E,
26
87
40
8
161
3
E
87
448
147
26
708
E, &D
40
147
118
30
335
X
C.BcfcA
8
26
30
30
94
Totals
16]
708
335
94
1298
32
ETHEL M. ELDERTON
Intelligence. Female Cousins.
Table XIV. Type D.
First Female Cousin
F&E2
E
Ej&D
C.B&A
Totals
•1
F&E,
6
23
29
1
59
o
E
23
336
133
14
506
"c«
£
e,*d
29
133
198
37
397
T3
C.B&A
1
14
37
8
60
Totals
59
506
397
60
1022
Table XV. Type E.
First Female Cousin
F&E2
E
Ej & D
C.B&A
Totals
F & E.:
8
9
17
1
35
E
9
262
97
22
390
E, &D
17
97
56
10
180
C,B&A
1
22
10
18
51
Totals
35
390
180
51
656
Table XVI. Type F.
First Female Cousin
F & E.,
E
Ej&D
C.B4A
Totals
F&E,
8
37
15
4
64
E
37
606
155
21
819
El(fcD
15
155
160
24
354
C.B&A
4
21
24
8
57
Totals
64
819
354
57
1294
MEASURE OF RESEMBLANCE OF FIIiST COUSINS
33
Intelligence. Male and Female Cousins.
Table XVII. Type G.
Male Cousin
F&E.
E
E, & D
C,B<S \
Totals
F&E,
11
30
15
56
B
3
0
E
09
266
60
31
429
"5
=
fa
E, ct D
63
93
124
50
330
C.B&A
•2
12
21
13
48
Totals
1 15
401
220
97
s.;:;
Table XVIII. Type II.
Male Cousin
1 A Es
E
Bj .v- D
CB&A
Totals
P & )•:,
13
46
12
2
73
c
3
O
O
E
29
279
74
20
102
S
E, & D
25
107
50
5
187
C.B&A
4
6
10
7
27
Totals
71
438
146
34
689
Table XIX. Type I.
Male Cousin
F & Ea
E
E, & DC, BAA
Totals
F & E,
9
23
13 ' 5
50
c
3
s
E
55
285
67 17
424
1
fa
E1 &T>
22
60
77 9
168
C.B&A
4
14
12 13
43
Totals
90
382
169 14
685
Table XX. Type A".
Male Cousin
FAE.
E
E, & D
CB&A
Totals
F & E,
17
23
9
1
50
1
E
34
334
93
19
480
"3
fa
E, & D
12
67
70
13
162
c.b&a
6
11
8
4
29
Totals
69
435
180
37
721
34
ETHEL M. ELDERTON
Success. Male CousrNS.
Table XXI. Type A.
First Male Cousiu
Marked &
Prosperous
Average
Difficult
Failure
Totals
Marked &
Prosperous
39
102-25
27-75
15
184
Average
102-25
196-5
40-25
16
355
Difficult
27-75
40-25
28
6
102
Failure
15
16
6
10
47
Totals
184
355
102
47
688
Table XXII. Type B.
First Male Cousin
M. & P.
A.
D.
F.
Totals
■1
M. & P.
90
96
35
14-5
235-5
0
8
0>
A.
96
139
47-5
14-5
297
1
£
D.
35
47-5
22
7
111-5
F.
14-5
14-5
7
,
44
Totals
235-5
297
111-5
44
688
Table XXIII. Type C.
First Male Cousin
M. & P.
A.
D.
F.
Totals
a
M. &P.
79
147-75
47-25
26
300
o
O
A.
147-75
272
80-75
24
524-5
D.
47-25
Si 1-75
51-5
6
185-5
1
F.
26
24
6
10
66
Totals
300
524-5
185-5
66
1076
MEASURE OF RESEMBLANCE OF FIIIKT COUSINS
35
SrccKss. Female ( 'oi sins.
Table XXIV. Typi />.
First Female I lonsin
M.A P.
A.
D.
F.
roi ii
M.A P.
21
5975
6-25
-
87
A.
59-75
450-5
30-25
8
548-5
D.
6-25
30-25
-
-
36-5
F.
-
8
-
-
8
Totals
87
548-5
36-5
8
680
I \i;i.k \\\ Type I:
Firsi Female Cousin
Table XXVI. Type F.
First Female Cousin
M.AP.
A.
D.
F.
Totals
M. & V.
64
94
6-5
12
176-5
A.
94
604-5
33-5
4
736
D.
6-5
33-5
5-5
-
45-5
F.
12
4
-
16
Totals
176-5
736
45-5
16
974
36
ETHEL M. ELDERTON
Success. Male and Female Cousins.
Table XXVII. Type G.
Male Cousin
M. & P.
A.
D.
F.
Totals
M. & P.
20
32-25
23-75
17
93
a
A.
104-5
224-75
83-75
43
456
a
D.
6-5
21
8-5
4
40
F.
4
1
-
-
5
Totals
135
279
116
64
594
Table XXVIII. Type H.
Male Cousin
M. & P.
A.
D.
F.
Totals
M. d- P.
23-5
24-5
5
11
64
A.
78-75
1 18-75
44
10
281-5
D.
6-25
21-25
24
6
57-5
F.
5
6
1
2
14
Totals
113-5
200-5
74
29
417
Table XXIX. Type I.
Male Cousin
M.&P.
A.
D.
F.
Totals
M.&P.
20-5
35-5
31-5
9
96-5
A.
91-75
228-25
54-5
9
383-5
D.
8-75
9-25
5
-
23
F.
1
-
-
-
1
Totals
122
273
91
18
504
Table XXX. Type K.
Male Cousin
M. & P.
A.
D.
F.
Totals
M. A P.
38
24-75
12-75
9-5
85
1
A.
80-75
191-75
75
17
364-5
1
D.
5-75
7-5
5-75
1-5
20-5
F.
3
2
4
•2
11
Totals
127-5
•226
97-5
30
481
MEASURE OF KFSEMBLANCF OF FIRST COUSINS
37
Temper. Male ( Iousinj
Table XXXF Type A.
First Male Cousin
Even
Quick
Sullen
Weak
Totals
a
Even
239
68
28-5
20-5
356
o
O
Quick
68
65
1625
12-75
162
a
-
Sullen
28-5
16-25
c,.-.
375
55
oa
Weak
20-5
1H-7--.
3-75
4
41
Totals
356
162
55
41
614
Table XXXI 1. Type 11.
First Male Cousin
Even
Quick
Sullen
Weak
Totals
_
Even
202
84-5
17-5
17-5
321-5
0
O
Quick
84-5
■_".i
9-5
7-5
130-5
C
8
V.
Sullen
17-5
9-5
-
5
32
Weak
17-5
7-5
5
2
32
Totals
321-5
130-5
32
32
516
Table XXXIII. Type C.
First Male Cousin
Even
Quick
Sullen
Weak
Totals
Even
419-5
179
43-5
24
666
Quick
179
96-5
22
17
314-5
Sullen
13-5
22
11
8-5
85
Weak
24
17
8-5
5
54-5
Totals
666
314-5
85
54-5
1120
38
ETHEL M. ELDERTON
Temper. Female Cousins.
Table XXXIV. Type D.
First Female Cousin
Even
Quick
Sullen
Weak
Totals
•i
Even
361
118-25
61-25
18-5
559
o
Q
Quick
118-25
77-5
27-75
8-5
232
Sullen
61-25
27-75
10
5-5
104-5
as
Weak
18-5
S-5
5-5
-
32-5
Totals
559
232
104-5
32-5
928
Table XXXV. Type E.
First Female Cousin
Even
Quick
Sullen
Weak
Totals
Even
269-5
75-5
16-5
6
367-5
Quick
75-5
64-5
7-5
4
151-5
Sullen
16-5
7-5
3
27
Weak
6
4
-
10
Totals
367-5
151-5
27
10
556
Table XXXVI. Type F.
First Female Cousin
Even
Quick
Sullen
Weak
Totals
.£
Even
545-0
194-5
49-5
11-5
801
6
Quick
194-5
116-5
22
6-5
339-5
5
Sullen
49-5
22
4
6
81-5
o
1
Weak
11-5
6-5
6
-
24
Totals
801
339-5
81-5
24
1246
MEASURE OF RESEMBLANCE OF FIRST COUSINS
39
Temper. Male and Female Cousins.
Table XXXVII. Type G. Table XXXVIII. Type II.
Male (
ousin
Even
Quick
Sullen
Weak
Totals
Even
265-5
98-5
500
23
137
s
6
Quick
104 23
72-25
•29-0
19
•224-5
"3
=
Sullen
33
8-5
9-5
2
53
Weak
12-75
6-75
3
7
29-5
Totals
415-5
186
91-5
«
744
Male C
ousin
Even
Quick
•Sullen
Weak
Totals
Even
206-5
74
25-5
25-5
33 1 -5
o
O
Quick
89
49
26
7-5
171-5
ta
Sullen
13-5
6-5
6
1
27
Weak
■2
1
-
5
8
Totals
311
130-5
57-5
39
538
Table XXXIX. Type I.
Male Cousin
Even
Quick
Sullen
Weak
Totals
Even
230
78
21
11-5
340-5
o
O
Quick
108-25
64-75
21
9-5
203-5
X
Sullen
1 1-25
10-75
4
2
31
Weak
6
5
-
-
11
Totals
358-5
158-5
46
23
586
Table XI, Type K.
Male Cousin
Even
Quick
Sullen
Weak
Totals
Even
2735
65-25
20-75
12
371-5
a
o
O
Quick
100-25
61-25
11
14-5
187
S
Sullen
14-25
14:5
5-25
3-5
37-5
Weak
10
3
1
2
16
Totals
398
1 11
38
32
612
40
ETHEL M. ELDERTON
Temperament — Reserved or Expressive. Male Cousins.
Table XLI. Type A. Table XLII. Type B.
First Male Cousin
Reserved
Betwixt
Expressive
Totals
»
o
O
1
Reserved
100
47
46
193
Betwixt
47
58
31
136
1
Expressive
46
31
68
145
Totals
193
136
145
474
First Male Cousin
Reserved Betwixt
Expressive Totals
Reserved
32
44
62
138
_0>
1
Betwixt
44
64 27
135
Expressive
62
27 50
139
Totals
138
135 139
412
Table XLII1. Type C.
First Male Cousin
Reserved
Betwixt
Expressive
Totals
Reserved
202
105
107
414
Betwixt
105
108
68
281
Expressive
107
68
112
287
Totals
414
281
287
982
MEASURE OF RESEMBLANCE OF FIRST CoUSlXS
41
Temperament — Reserved or Expressive. Female Cousins.
Table XLIV. Type D.
First Female Cousin
Table XLV. Type E.
First Female Cousin
Reserved
Betwixt
Expressive
Totals
1
<D
-c
1
Reserved
Betwixt
Expressive
Totals
s
•S3
1
Reserved
94
83
100
277
Reserved
32
34
53
119
1
Betwixt
83
110
62
255
Betwixt
34
74
50
158
a
o
a
Expressive
100
62
122
284
Expressive
53
50
134
237
Totals
277
255
284
816
Totals
119
158
237
514
Table XLVI. Type F.
First Female Cousin
Reserved
Betwixt
Expressive
Totals
o
O
Reserved
86
111
117
314
1=5
Betwixt
111
120
110
341
o
a;
02
Expressive
117
110
240
467
Totals
314
341
467
1122
42
ETHEL M. ELDERTON
Temperament — Reserved or Expressive. Male and Female Cousins.
Table XL VII. Type G
Male Cousin
Reserved
Betwixt
Expressive
Totals
a
Reserved
87
50
53
190
3
Eetwixt
57
87
41
185
&
Expn ssive
106
47
96
249
Totals
250
184
190
624
Table XLVIII. Type H.
Male Cousin
Reserved
Betwixt
Expressive
Totals
.2
Ki-rni'd
26
26
27
79
o
a
_2
Betwixt
72
69
33
174
£
Expressive
105
31
89
225
Totals
203
126
149
478
Table XLIX. Type I.
Male Cousin
Table L. Type K.
Male Cousin
Reserved
Betwixt
Expressive
Totals
5
O
1
Reserved
Betwixt
Expressive
Totals
Reserved
77
35
35
147
Reserved
63
59
25
147
Betwixt
64
36
61
161
Betwixt
70
72
45
187
Expressive
100
29
105
234
Expressive
71
27
83
181
Totals
241
100
201
542
Totals
204
158
153
515
MKASrKK OF liFSFMIUAXOF OF FIliST COUSINS
l::
Temperament — Reserved or Expressive. Cousins. All Types.
Table LI. Male Cousins.
First Male Cousin
Reserved
Betwixt
Expressive
Totals
71-")
Reserved
334
196
2 1 5
Betwixt
196
■230
126
552
Expressive
215
126
230
571
Totals
745
552
571
L868
Table LII. Female Cousins.
First Female Cousin
Keserved
Betwixt
Expressive
Totals
Reserved
212
228
270
710
Betwixt
228
304
222
754
Expressive
270
222
496
988
Totals
710
754
988
2452
Table LII I. Male and Female Cousins.
Male Cousin
1 Reserved
Betwixt Expressive
Totals
a
Reserved
253
170
140
563
o
O
<E
Betwixt
263
264
180
707
id
Expressive
382
134
373
889
Totals
898
568
693
2159
41
ETHEL M. ELDERTON
Temperament — Sympathetic or Callous. Male Cousins.
Table LIV. Type A.
First Male Cousin
Table LV. Type B.
First Male Cousin
Symp.
Betwixt
Callous
Totals
o
O
Symp.
Betwixt
Callous
Totals
Syrnp.
91
28
■2
121
Symp.
67
33
9
109
Betwixt
35
36
5
76
"3
3
Betwixt
24
49
5
78
Callous
31
15
2
48
o
0Q
Callous
6
7
10
23
Totals
157
79
9
245
Totals
97
89
24
210
Table LVI. Type C.
First Male Cousin
Symp.
Betwixt
Callous
Totals
Symp.
170
67
18
255
Betwixt
74
84
19
177
Callous
39
20
2
61
Totals
283
171
39
493
MEASURE OF RESEMBLANCE < >F MUST COUSINS
15
Temperament — Sympathetic or Callous. Female Cousins.
Table LVII. Type D.
First Female Cousin
Symp.
Betwixt
Callous
Totals
Syrup.
133
59
11
203
Betwixt
102
61
8
171
Callous
30
12
2
44
Totals
265
132
21
418
Table LVIII. Typt K.
First Female Cousin
Symp.
Betwixt
Callous
Totals
1
Symp.
151
40
7
198
o
~
Betwixt
26
27
1
54
r*
5
s
Callous
15
7
7
29
xn
Totals
192
74
15
281
Table LIX. Type F.
First Female Cousin
Symp.
Betwixt
Callous
Totals
o
O
Symp. ! 321
82
14
417
Betwixt
53
46
6
105
a
o
1
Callous
25
11
4
40
Totals
399
139
24
562
46
ETHEL M. ELDERTON
Temperament — Sympathetic or Callous. Male and Female Cousins.
Table LX. Type G.
Male Cousin
Symp.
Betwixt
Callous
Totals
Symp.
257
113
-
412
Betwixt
85
87
25
197
Callous
28
9
2
39
Totals
370
209
69
648
Table LXI. Type H.
Male Cousin
Symp.
Betwixt
Callous
Totals
Symp.
205
91
15
311
"3
c
Betwixt
41
92
12
145
3
b
Ph
Callous
13
4
12
29
Totals
259
187
39
485
Table LXII. Type I.
Male Cousin
Table LXIII. Type K.
Male Cousin
Symp.
Betwixt
Callous
Totals
o
O
f2
Symp.
Betwixt
Callous
Totals
.3
Symp.
293
66
43
402
Symp.
225
100
33
358
o
O
Betwixt
50
53
16
119
Betwixt
70
67
9
146
Callous
11
8
3
22
Callous
14
5
1
20
Totals
354
127
62
543
Totals
309
172
43
524
MEASURE OF RESEMBLANCE OF FIRST COUSINS
Temperament — Sympathetic oe Callous. Cousins. All Types.
Table LXIV. Male Cousins.
First Male Cousin
17
Symp.
Betwixt
Callous
Tola],
Symp.
656
261
105
1022
Betwixt
261
338
71
670
Callous
105
71
28
204
Totals
1022 670
204
1896
Table LXV. Female Cousins.
First Female Cousin
Symp.
Betwixt
Callous
Totals
Symp.
1210
362
102
1674
Betwixt
362
268
45
675
Callous
102
45
26
173
Totals
1674
r,::,
173
2522
Table LXVI. Male and Female Cousins.
Male Cousin
Symp.
Betwixt
Callous
Totals
a
Symp.
980
370
133
1 183
o
O
Betwixt
24G
299
02
607
3
Callous
66
26
18
110
Totals
1292
695
213
2200
48
ETHEL M. ELDERTON
Temperament — Excitable or Calm. Male Cousins.
Table LXVII. Type A.
First Male Cousin
Table LXVIII. Type B.
First Male Cousin
Excit.
Betwixt
Calm
Totals
o
O
"3
1
Excit.
Betwixt
Calm
Totals
o
1
Excit.
48
21
60
129
Excit.
40
18
42
100
Betwixt
21
68
41
130
Betwixt
18
90
34
142
o
1
Calm
60
41
126
227
Calm
42
34
94
170
Totals
129
130
227
486
Totals
100
142
170
412
Table LXIX. Type C.
First Male Cousin
Excit.
Betwixt
Calm
Totals
o
O
1
Excit.
78
56
122
256
Betwixt
56
166
93
315
CO
Calm
122
93
224
439
Totals
256
315
439
1010
MEASURE OF RESEMBLANCE OF FIRST COUSINS
i:»
Temperament -Excitable ob Calm. Female Cousins.
Table LXX. Type D.
First Female Cousin
Excit.
Betwixt
Calm
Totals
=
6
Excit.
58
30
110
198
Betwixt
30
70
99
199
8
Calm
110
99
226
435
Totals
198
199
435
832
Table LXXI. Type E.
First Female Cousin
Excit.
Betwixt
Calm Totals
Excit.
70
37
73 180
Betwixt
37
36
47 120
Calm
73
47
82
202
Totals
180
120
202
502
Table LXXII. Type F.
First Female Cousin
Excit.
Betwixt
Calm
Totals
I
=
fa
—
a
I
Excit.
136
54
163
353
Betwixt
54
110
76
240
Calm
163
76
322
561
Totals
353
240
561
1154
50
ETHEL M. ELDERTON
Temperament — Excitable or Calm. Male and Female Cousins.
Table LXXIII. Type G.
Male Cousin
Excit.
Betwixt
Calm
Totals
Excit.
45
39
81
165
•-
b
Betwixt
12
70
54
136
-
H
£
Calm
47
82
1G3
292
Totals
104
191
298
593
Table LXXIV. Type If.
Male Cousin
Excit.
Betwixt
Calm
Totals
Excit.
48
26
76
150
"53
o
Betwixt
31
87
66
184
~3
£
Calm
42
31
70
143
Totals
121
144
212
477
Table LXXV. Type I.
Excit.
Betwixt
Calm
Totals
Excit.
66
27
115
208
Betwixt
23
42
49
114
Calm
51
41
135
227
Totals
140
110
299
549
Table LXXVI. Type K.
Male Cousin
Exeit.
Betwixt
Calm
Totals
Excit.
44
21
87
152
Betwixt
18
65
32
115
Calm
60
50
152
262
Totals
122
136
271
529
MKASIIJK OF IIESKMBLANCE OK FIRST COI'SINS
51
Temperament — Excitable oe Calm. Cousins. All Types.
Table LXXVII. Male Cousins.
First Male Cousin
Excit.
Betwixt
Calm
Totals
a
1
Exoit.
166
95
221
485
Betwixt
95
324
168
587
c
q
I
Calm
•J24
168
444
836
Totals
is:,
587
836
1908
Table LX XVIII. Female Cousins.
First Female Cousin
Excit.
Betwixt
Calm
Totals
Excit.
264
121
346
731
Betwixt
121
216
222
559
Calm
346
222
630
1198
Totals
731
559
1198
2488
Table LXXIX. Male and Female Cousins.
Male Cousin
Excit.
Betwixt
Calm
Totals
_g
Excit.
203
113
359
675
3
o
Betwixt
84
264
201
549
£
Calm
200
204
520
924
Totals
487
581
1080
2148
52
ETHEL M. ELDERTON
PlGMENT-ATK >X CHARACTERS.
Table LXXX. Eye Colour, Female Cousins.
First Female Cousin
Tints
Pure Blue
Blue Orange
Pure Grey
Grey Orange
Hazel Green
Hazel Brown
Brown
Totals
c
Pure Blue
9
1-5
8
2-5
4-5
2-5
4-5
32-5
o
Blue Orange
1-5
6
7
9
3-5
3-5
8-5
39
(J
Pure Grey
8
7
4
5
1
2
1-5
28-5
2
Grey Orange
2-5
9
5
6
1
6
2
31-5
Ph
Hazel Green
4-5
3-5
1
1
1-5
4-5
3
19
"3
Hazel Brown
•2-5
3-5
2
6
4-5
8-5
5-5
325
tc
Brown
4-5
8-5
1-5
2
3
5-5
10
35
Totals
32-5
39
28-5
31-5
19
32-5
35
218
Table LXXXI. Hair Colour, Female Cousins.
First Female Cousin
Tints
Very Dark
Park Brown
Brown
Light Brown
Fair
Red
Totals
-3
Very Dark
9
11
3-5
11
1-5
36
o
O
Dark Brown
11
13
12
10
9
1
56
•%
Brown
3-5
12
8
9-75
3-25
3
39-5
ti
Light Brown
11
10
9-75
16-5
9-25
1
57-5
a
O
Fair
1-5
9
3-25
9-25
1
—
24
Red
1
3
1
_
_
5
Totals
36
56
39-5
57-5
24
5
218
M.KASU11K OF RESKMBLANOF OF FIRST COUSINS
53
Pathological Characters.
Table LXXXII. Tuberculosis.
First Male Cousin
Tuberculous
Non-tuberoulous
Totals
Tuberculous
Non-tuberculous
18
188
L88
2596
206
2784
Totals
206
2784
2990
Table LXXXII 1. Insanity*.
First Mala Cousin
Insane
Sane
Totals
Insane
Sane
38
221
221
2510
259
2731
Totals
259
2731
2990
Table LXXXIV. Tuberculosis.
First Female Cousin
Tuberculous
Non-tuberculous
Totals
Tuberculous
Non-tuberculous
20
185
185
2S52
205
3037
Totals
205
3037
3242
Table LXXXV. Insanity*.
First Female Cousin
.5
5
O
Insane
Sane
Totals
Insane
Sane
6
124
124
2996
130
3120
0
Totals
130
3120
3250
Table LXXXVI. Tuberculosis.
Male Cousin
Tuberculous
Non-tuberculous
Totals
Tuberculous
Non-tuberculous
20
237
148
2698
168
2935
Totals
257
2846
3103
Table LXXXVII. Insanity-
Male Cousin
Insane
Sane
Totals
Insane
Sane
12
187
128
2791
140
2978
Totals 199 2919
3118
Includes marked neuroses, alcoholism and hysteria
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A Cooperative Study of Queens, Drones and Workers in
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II. Natural Selection in Iklix Arbustomm. By A. P. di Notices and Bmhograpby.
Cesnola.
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Elder-ton, Ethel Mary.
On tht
measure of the
resemblance of first
cousins