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Das Tierreich.
Eine Zusammenstellung und Kennzeichnung der
rezenten Tierformen.
Herausgegeben
von der
Deutschen Zoologischen Gesellschaft.
Greneralredakteur : Franz Eilhard Schulze.
3. Lieferung.
Acariria.
Kedakteur : H. L o li m a n n.
Oribatidae
bearbeitet von
A. D. Michael, P. L. s., F. z. s., p. B. M. s., etc.
in London.
Mit 15 Abbildungen im Texte.
Ausgegeben im Juli 1898.
Berlin.
Verlag von R. Friedlander und Sohn.
1898.
Das Tierreich.
Herausgegeben von der Deutschen Zoologischen Gesellschaft.
Generalredakteur : Franz Eilhard Schulze.
Acarina. 3. Lieferung. Rcdakteur: H. Lohmann.
Oribatidae
A. D. Michael, F. L. s., F. z. s., F. B. M. s., etc.
of London.
With 15 figures.
Ausgegeben im Juli 1898.
'
Berlin.
Verlag von R. Friedlander und Sohu.
1898.
Complete to 31 March 1897.
All rights reserved.
Abbreviations of the technical terms.
Abd. - - Abdomen (see pag. 2)
Ant. - - Anterior
Bid. - - Bidactyle
Cephth. Cephalothorax (see pag. 2)
C. lam. - - Cusp of the lamella (see
pag. 3)
Dors. - - Dorsum, Dorsal
Hetd. - - Heterodactyle (see pag. 4)
Homd. - - Homodactyle (see pag. 4)
Im. - - Imago
Intlam. hair Interlamellar hair (see
pag. 4)
Lam. Lamella, Lamellar (see
pag. 3)
Lat. Lateral
Lv. Larva
Mdb. - - Mandible (see pag. 3)
Mond. - - Monodactyle
Nph. Nymph
Post. - - Posterior
Psstg. - - Pseudostigma (see pag. 4)
Psstg. orcj. - - Pseudostigmatic organ
(see pag. 4)
Ptm. - - Pteromorpha (see pag. 4)
Rost. Eostrum, Rostral (see pag. 2)
Tectp. - - Tectopedium (see pag. 4)
Trid. -- Tridactyle
Trlam. - - Translamella (see pag. 3)
Ung. - - Unguis
Ventr. Ventral
L. - - Length
/ff
Abbreviations of the titles of literature.
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zu Bremen. Bremen. 8.
Amer. Natural. The American Naturalist. Salem, etc. 8.
Ann.Sti.nat. Annales des Sciences naturelles. Zoologie. Paris. 8.
Arch. Dorpat. Ges. Archiv fiir die Naturkunde Liv-, Ehst- und Kurlands. 2. Ser.
Dorpat. 8.
Arch. Mus. Paris -- Archives du Museum d'Histoire naturellc. Paris. 4.
Arch. Naturg. Archiv f'iir Naturgeschichte. Berlin. 8.
Atti 1st. Veneto Atti del Reale Istituto Veneto di Scienze. Lettere ed Arti. Venezia. 8.
Atti Soc. Veneto-Trent. Atti della Societa Veneto-Trentina di Scienze natural!, resi-
dente in. Padova. Padova 8.
Ber. Ges. Bamberg Bericht der naturforschenden Gesellschaft in Bamberg. Bamberg. 8.
Berlese, A. M . S. - - A. Berlese, Acari, Myriopoda et Scorpiones hucusque in Italia
reperta. fasc. 179. Padova, 188296. 8.
Berlese, A. M. S., Crypt. II -- A. Berlese, Acari. Myriopoda et Scorpiones hucusque
in Italia reperta. Ordo Cryptostigmata (Oribatidae). Portici, 1896. 8.
Berlese, A. M.S., Repert. A. Berlese, Acari, Myriopoda et Scorpiones hucusque in
Italia reperta. Repertorium Specierum novarum. Padova. 8. [in tegu-
mentis fasciculorum.]
Berlin, ent. Z. Berliner entomologische Zeitschrift. Berlin. 8.
Bibl. zool. Bibliotheca zoologica. Original- Abhandlungen aus dem Gesammtgebiete
der Zoologie. Cassel. Stuttgart. 4.
Biol. Centr.-Amer., Arach. acar. - F. D. Godman and O. Salvin, Biologia Centrali-
Americana. Arachnida acaridea by 0. Stoll. London, 1886 93. 4.
Buffon, Hist, nat., ed. Sonnini. Ins. G. L.L. Comte de Buff on, Histoire naturelle.
generale et particuliere. Nouvelle Edition. Red. par Ch. N. S. Sonnini. -
Crustaces et Insectes par P. A. Latreille. 14 v. Paris. 1792 1805. 8.
Bull. Dep. Agric. Ent. - - U. S. Department of Agriculture. Division of Entomology.
Bulletin. Washington. 8.
Bull. Soc. ent. Ital. - - Bullettino della Societa entomologica Italiana. Firenze. 8.
Bull. Soc. Veneto-Trent. Bullettino della Societa Veneto-Trentina di Scienze naturali.
Padova. 8.
Canad. Ent. The Canadian Entomologist. Toronto, London. 8.
Canestrini, Prosp. Acarof. -- G. Canestrini, Prospetto dell' Acarofauna Italiana.
7 v. Padova, 1885 97. 8. [& in: Atti Soc. Veneto-Trent.]
Carus & Gerstaecker, Zool. J. V. Carus und C. E. A. Gerstaecker, Handbuch der
Zoologie. 2 v. Leipzig, 186375. 8.
Donnadieu, Rech. Tetranyques A. L. Donnadieu, Recherches pour servir a 1'Histoire
des Tetranyques. Lyon, 1875. 8. [& in: Ann. Soc. Linn. Lyon, n. ser. v. 22. 1876.]
Enc. meth. Encyclopedic methodique, ou par Ordre de Matiere. Par une Societe de
Gens de Lettres. Histoire naturelle. 10 v. & 17 atl. Paris, 17821832. 4.
Expl. Algerie, An. artic. - Exploration scientifique de 1'Algerie pendant les Annees
1840 42. Sciences physiques. Zoologie. Histoire naturelle des Animaux
articules par H.Lucas. 3 v. & atl. Paris. 1849. 4.
Abbreviations VII
Fabricius, Ent. syst. J. Ch. Fabricius, Entornologia systematica. emendata et aucta.
secundum Classes. Ordines .... 4 v., suppl. & indices. Hat'niae, 1792 99. 8.
Fiirstenberg, Krdtzm. 31. H. F. Fiirstenberg. Die Kratzmilben der Menschen und
Thiere. Leipzig. 1861. 2.
Gay, Hist. Chile. - - Cl. Gray, Historia fisica y politica de Chile segun Documentos
adquiridos .... Publ. bajo los Auspicios del supremo Gobierno. Zoologia.
8 v. & atl. Paris, 184754. 8 & 2.
Hermann, Mem. apt. J.F.Hermann, Memoire apterologique. Ouvrage couronne
en 1790 par la Soeiete d'Histoire naturelle de Paris. Publ. par F. L. Hammer.
Strasbourg, 1804. 2.
Hoeven, Handb. Dierk. J. van der Hoeven, Handbook der Dierkunde. 2v. Rotterdam.
1828- -33. 8. (ed. 2, 1849.)
Institut L'Institut. Journal universel des Sciences et des Societes savantes en
France et a 1'Etranger. Sect. I, Sciences math., phys. et nat. Paris. 4.
Isis Isis. Encyklopadische Zeitschril't, von L. Oken. Leipzig. 4.
,7. Ac. Philad. J ournal of the Academy of natural Sciences of Philadelphia. Philadelphia. 4.
Jahresh. Ver. Wiirttemb. - - Jahreshefte des Vereins fiir vaterlandische Naturkunde in
Wiirttemberg. Stuttgart. 8.
J. Anat. et Physiol. --- Journal de 1' Anatomic et de la Physiologic normales et patho-
logiques de 1'Homme et des Animaux. Paris. 8.
./. ,Y. York ent. Soc. - - Journal of the New York entomological Society. New York. 8.
J. R. micr. Soc. Journal of the Royal microscopical Society. London. 8.
C. L. Koch, C.M. A. - C.L.Koch, Deutschlands Crustaceen. Myriapoden und Arachniden.
Hrsg. von G. A. W. Herrich-Schaffer. 40 fasc. Regensburg, 1835 44. 12.
C.L.Koch, Ubers. Arach. - - C. L. Koch, Ubersicht des Arachnidensy stems. 5 v.
Nurnberg, 183750. 8.
Latreille, Gen. Crust. Ins. P. A. Latreille. Genera Crustaceorum et Insectorum
secundum Ordinem naturalem in Familias disposita, Iconibus Exemplisque
plurimis explicata. 4 v. Parisiis etc., 1806 09. 8.
Latreille, Precis Caract. Ins. P. A. Latreille, Precis des Caracteres generiques des
Insectes disposes dans un Ordre naturel. Paris, 1796. 8.
Linne, Fauna Suec., ed. 2 . Linne, Fauna Suecica sistens Animalia Sueciae Regni.
ed. 2. Stockholmiae, 1761. 8.
Linne, Syst. Nat., ed. 10 C. Linne, Systema Naturae, ed. 10. v. 1. Holmiae, 1758. 8.
Mag. enc. Magasin encyclopedique ou Journal des Sciences, des Lettres et des Arts.
Paris. 8.
Math, naturw. Ber. Ungarn Mathematische und naturwissenschaftliche Berichte aus
Ungarn. Budapest, Berlin. 8.
Mem. Ac. Washington Memoirs of the national Academy of Sciences. Washington. 4.
Michael, Brit. Orib. A.D.Michael, British Oribatidae. 2 v. London (Ray Soc.),
1884, 88. 8.
Miss. Cap Horn Mission scientifique du Cap Horn 1882 83. 8 v. Paris. 1885 91. 4.
Mt. Mus. Hamburg. - - Mitteilungen aus dem naturhistorischen Museum in Hamburg.
Hamburg. 8.
Murray, Econ. Ent. Apt. A.Murray, Economic Entomology. Aptera. South Ken-
sington Museum Science Handbooks. London, 1877. 8.
N. Acta Soc. Upsal. Nova Acta Regiae Societatis Scientiarum Upsaliensis. Upsaliae. 4.
Nouv. Diet. Nouveau Dictionnaire d'Histoire naturelle applique aux Arts, a 1' Agri-
culture Par une Societe de Naturalistes et d'Agriculteurs. 24 v. Paris.
1802-04. 8.
Of a. Ak.Forh. Ofversigt af KongligaVetenskaps-AkademiensForhandlingar. Stockholm. 8.
Perty, Allg. Naturg. M. Perty, Allgemeine Naturgeschichte, als philosophische und
Humanitatswissenschaft. 4 v. Bern, 1837 45. 8.
P. zool. Soc. London Proceedings of the zoological Society of London. London. 8.
Rep. Ins. Missouri Annual Report on the noxious, beneficial and other Insects of
the State of Missouri. Jefferson City. 8.
Rep. Ins. N. York - - Report on the noxious, beneficial and other Insects of the State
of New York. Albany. 8. [& in : Tr. N. York Agric. Soc.]
VIII Abbreviations
Rev. biol. Nord France Revue biologique du Nord de la France. Lille. 8.
Rev. Sci.nat. Quest Revue des Sciences naturelles de 1'Ouest. Paris. 8.
Riv. Ace. Padova - Rivista periodica dei Lavori dclla Reale Accadeniia di Scionze,
Lettere ed Arti in Padova. Padova. 8.
Schrank, Fauna Boica F. v. P. Schrank, Fauna Boica. Durchgedachte Geschichte
der in Bayern einheimischen und zahmen Thiere. 3 v. Niirnberg, Ingolstadt,
1798, 180103. 8.
Schrank, Ins. Aiijttr. F. v. P. Schrank, Enumeratio Insectorum Austriae indigenorum.
Augustae Vindelicorum, 1781. 8.
Scopoli, Ent. Carniol. -- J. A. Scopoli, Entomologia Carniolica. exhibens Insecta
Carnioliae indigena .... Vindobonae, 1763. 8.
Svenska Ak. Handl. Kongliga SvenskaVetenskaps-AkademiensHandlingar. Stockholm. 4.
Tijdschr. Ent. - Tijdschrift voor Entomologie. La Haye. Leiden etc. 8.
Tr. Ac. St. Louis The Transactions of the Academy of Science of St. Louis. St. Louis. 8.
Tr. Amer. ent. Soc. Transactions of the American entomological Society. Philadelphia. 8.
Tr. Benvickshire Club - - Transactions of the Berwickshire Naturalists Field Club.
Walckenaer, Ins. Apt. Baron Walckenaer, Histoire naturelle des Insectes. Apteres.
4.&atl. Paris, 183747. 8.
Z. iviss. Zool. Zeitschrift fiir wissenschaftliche Zoologie. Leipzig. 8.
Systematic Index.
p
Fam. Oribatidae ....
ag.
1
21.
22.
23.
i
0. aquatica (Banks) ....
berlesei Michael ....
,. languida Nic
>ag.
18
18
1R
A. Subfam. Oribatinae . . .
1. Gen. Pelops C.L.Koch . . .
1. P. acromius (Herm.) . . .
2 auritus C L Koch
;>
6
6
7
24.
25.
26.
27.
ovalis C. L. Koch ....
quadricornuta Michael . .
quadridentata (Banks) . .
.. tecta Michael
19
19
19
^0
3 farinosus Nic
7
28.
a'ilis Nic
20
4 fuligineus C L Koch fP.
29.
,. alata (Herm )
00
plicatus (C. L. Koch)] .
5. ,. OCCllltllS C. L. Koch, Nic. .
6. tardus C. L. Koch ....
7. torulosus C. L. Koch . .
8. uraceus C.L.Koch . . .
9. glaber G. & R. Can. . . .
10. phaeonotus C L Koch . .
7
8
8
8
8
8
q
30.
31.
32.
33.
34.
35.
36.
avenifera Michael . . .
centro-americana Stoll .
CUSpidata Michael . . .
dorsalis (C.L.Koch) . .
,. halleri Michael ....
intermedia Michael . . .
,. lucasi Nic
20
21
21
21
21
21
99
2 Gen Oribata Latr
q
37.
mucrouata (I &R, Can
00
1. 0. antarctica Michael . . .
2. ,. clvpeata Nic
12
1?
38.
39.
pelopt&ides Berl
,, rugifrons Stoll ....
22
o,9,
3. depauperata Berl . . .
13
40.
,, dentata Berl
09
4 ,. edwardsi Nic
IS
41
fusigera Michael ....
?S
5. fuscipes C.L.Koch [O.ephip-
piata (C.L.Koch)] . .
6. gilvipes C.L.Koch . . .
7. ,. flobula Nic ...
13
13
1 1
42.
43.
44.
45
mirabilis (Banks) ....
.,. monodactyla Haller . .
,. parmeliae Michael . . .
rilevi Haller
23
23
23
1
8. gracilis Michael ....
11
affinis Banks
<>\
9. . lapidaria H Luc TO rubra
arborea Banks
M
(C.L.Koch)] ....
10. lucens L.Koch ....
11. microptera Can ....
14
15
IT
,. calcarata C.L.Koch . . .
depressa Banks ....
. flammula C L Koch . .
24
24
05
12. mollicoma C L Koch . .
I")
hll'SUta Banks
05
13. ,. orbicularis C. L. Koch . .
14. ,. pallidula (C.L.Koch) . .
15. ,. piriformis Nic ....
15
16
IB
,. honiodactyla Karp. . . .
,. latirostris (C.L.Koch) . .
,. magna Banks
25
25
05
16. punctum C.L.Koch, Berl. .
17. ,. rubens C L Koch . . .
16
17
minuta (Banks) ....
IBOesta Banks . . . .
25
05
18. setosaC.L.Koch[0. trima-
culata (C. L Koch)] . .
17
,. palustris Banks ....
,. pinnata Gerv
25
9,6
19. sphagni Michael ....
20. ,. americana Haller . . .
17
18
pratensis Banks ....
,. reticulata L. Koch . . .
26
26
Systematic Index
robusta Banks ....
Pag.
. 26
5. Gen Liacarus Michael
Pag.
4(1
signata (Banks) . . .
simplex Gerv. . . .
. 26
. 26
. 26
1. L. coracinus (C. L. Koch) . .
2. globifer (P.Kram.) . . .
3. globoSUS (Berl )
40
41
41
B. Subfam. Serrariinae . .
1. Gen. Serrarius Michael . .
1 S fusifer (C L Koch)
. 26
. 26
27
4. lativentris (Nic.) ....
5. longipilis (Monz.) . . .
6. marginatus (Nic.) . . .
7. nitens (Gerv )
41
41
41
11
2. microcephalus (Nic.) .
C. Subfam. Zetorchestinae
1. Gen. Zetorchestes Berl.
. 27
. 27
. 27
OG
8. ovatus (C.L.Koch) . . .
9. palmicinctus (Michael)
olivaceus (Grube) . . .
6. Gen. Notaspis Herm. . . .
1. N. exilis Nic . . ...
42
42
42
43
4">
D. Subfam. Notaspidinae .
1. Gen. Scuto vertex Michael .
. 28
. 28
2. plantivaga (Berl.) . . .
2ct.,, var. sicula (Berl.) .
3. similis Michael ....
4. tibialis Nic
45
45
45
15
. 8. bilmeatus Michael . .
2. caelatus Berl ....
. 29
. 29
5. aequalis Michael . . .
46
6. bipilis Herm
46
3. COlTUgatuS Michael . .
4. .. harioti Michael ....
. 29
. 30
6a.,, var. quadridentata
Haller
16
5. maculatus Michael . .
6. SCulptuS Michael . . .
.. SB.. (Berl )
. 30
. 30
. 31
7. buiTOWSi Michael . . .
8. conformis (Berl.) . . .
46
47
2. Gen. Cepheus C. L. Koch .
1. C. bifidatus Nic ....
. 31
. 32
9. glabra Michael ....
10. hepatica (C.L.Koch) . .
47
47
2. latus Nic
. 32
11. licnophora Michael . . .
47
3. OCellatuS Michael . . .
4. tegeocranus (Herm.)
5. variolosus Berl . . .
. 32
. 32
. 33
12. lucorum (C.L.Koch) . .
13. microptera (Berl.) . . .
14. oblonga (C. L. Koch) . .
47
48
48
punctlllatus Banks . .
3. Gen. Tegeocranus Nic, . .
1. T. cepheiformis Nic. . .
2. dentatus Michael ....
3. hericius Michael . . .
4. latus (C.L.Koch) . . .
5. leporosus (Haller) . .
6. velatus Michael . . .
4. Gen. Carabodes C. L. Koch .
1. C. apicalis Banks ....
. 33
. 33
. 34
. 34
. 34
. 35
. 35
. 35
. 36
. 37
15. pilosa (C.L.Koch) . . .
16. serrata Michael ....
17. clavipectinata Michael .
18. juncta Michael ....
19. lacustris Michael . . .
20. lanceolata Michael . . .
21. ., longilamellata Michael .
22. monilipes Michael . . .
23. pectinata Michael . . .
24. quadricarinata Michael .
48
48
49
49
49
50
50
50
51
51
2. brevis Banks ....
. 37
25. splendens (C. L. Koch) .
51
26 trigona Michael
51
3. clypeatus (Nic.) . . .
4. coriaceus C. L. Koch
5. elongatus (Michael) . .
6. femoralis (Nic.) . . .
7. granulatus Banks . . .
8. .. labyrinthicus (Michael)
. 37
. 37
. 37
. 38
. 38
. 38
distincta (Karp.) . . .
longiuseula (C. L. Koch) .
E. Subfam. Damaeinae . . .
1 Gen Ameriis Berl ....
52
52
52
"iH
9. marginatus (Michael)
. 39
1 4. troisi (Berl )
53
10. papillosus (H.Luc.) . .
cephalotes C.L.Koch .
., dorsalis Banks ....
. 39
. 39
. 39
2. Gen. Damaeus C. L. Koch . .
1. D. auritus C.L.Koch . . .
2 beiiesei Michael ....
53
55
niffer Banks .
39
3. ,.. clavines (Herm.)
55
Systematic Index
XI
Pag.
4. D. concolor C. L. Koch . .
56
4. N.
5. flagellifer Michael . . .
56
5.
6. geniculatus (L.), C.L.Koch
56
6.
7. globipes (Can. & Berl.)
57
7-
8. nigro-femoratus (L.Koch)
57
8.
9. nitens (C.L.Koch) . . .
57
9.
10. patelloides Michael . .
58
10.
11. phalangoides Michael . .
58
11.
12. ., pulverulentus (C. L. Koch)
58
12.
13. Sllfflexus Michael . . .
58
13.
14. tecticola Michael . . .
58
14.
15. ., tenuipes Michael . . .
59
15.
16. verticillipes Nic. . . .
59
16.
17. bicostatus C. L. Koch . .
59
17.
18. craterifer Haller . . .
60
18.
19. femoratus C.L.Koch . .
60
19. ,
20. ornatissimus Berl. . . .
60
n
21. SetoSUS Berl
60
bulbipes (Karp) . . . .
61
11
carinulatus (Berl.) . . .
61
n
n
formica G-erv
61
5. Gen.
, riparius Nic
61
1. L.
2.
F. Subfam. Nothrinae . . .
61
6. Gen.
1. Gen. Hermannia Xic. . . .
62
1. H.
1. H. arrecta Nic
62
2-
2. bistriata (Nic.) [H.palliata
3.
(C.L.Koch)] . . . .
63
3. carinata P. Kram. . . .
63
n
4. convexa (C. L. Koch) . .
63
G. Subfai
5. . granulata Nic . . . .
64
1/~*
. Gen.
6. nanus (Nic.)
64
1. H.
7. ovulum (Haupt) ....
64
2
8. ., reticulata Thor
64
*** it
3.
9. scabra (L.Koch) . . . .
65
** n
4.
2. Gen. Neoliodes Berl
65
* 11
5
1. N. concentricus (Say) . . .
65
" n
6
2. .. peltifer (C.L.Koch) . .
65
\j. .,
7
3. theleproctus (Herm.) . .
66
1 n
8
kovnhuberi (Karp.) . . .
66
0.
venOSUS (Grube) ....
66
11
3. Gen. Cymbaeremaeus Berl. .
66
n
2. Gen.
1. C. brevipes (Michael) . . .
67
1. P.
2. cymba (Nic )
67
9
3. .. fimbriatus (Michael) . .
67
* n
3
4. tibialis (Nic.)
68
** n
4
minutus (C.L.Koch) .. .
68
* n
5
4. Gen. Nothrus C.L.Koch . .
68
u.
6
1. N. anauniensis Can.&Fanx. .
70
V.
7
2. ., banks! Michael . . . .
70
1 * 5?
3. bicarinatus C. L. Koch
70
H
Pag.
biciliatus C. L. Koch . . 70
biverrucatus C. L. Koch . 71
echinatus C. L. Koch . . 71
exClSUS Banks .... 71
flagellatus Berl 71
glaber Michael .... 71
horridus (Herm.) ... 71
invemistus Michael ... 72
monodactylus Michael . 72
palustris C. L. Koch . . 72
mguloSUS Banks .... 73
segnis (Herm.) .... 73
spinifer C. L. Koch ... 73
sylvestris Nic 73
tardus Michael .... 74
targionii Berl 74
pileiformis Karp. ... 74
quadracanthus Haupt . 74
simplex Banks .... 75
sordidus C. L. Koch . . 75
Lohmannia Michael . . 75
murcioides (Berl.) . . . 75
paradoxa (Haller) . . . 75
Hypochthonius C.L.Koch 76
brevis Michael .... 76
lanatus Michael .... 76
pallidulus C. L. Koch . . 76
rufulus C.L.Koch ... 77
tectorum Berl. 77
1. H. anomalum (Berl.) . .
carinatum (C.L.Koch)
dasypus (Ant, Dug.)
globosum (C. L. Koch)
magnum (Nic.) . . .
pulcherrimum (Berl.) .
striculum (C. L. Koch)
variolosum (Berl.) . .
crinitum (C.L.Koch) .
quadriseriatum (Haupt)
Phthiracarus Perty .
1. P. arduus (C. L. Koch) . .
berlesei Michael . . .
canestrinii Michael . .
contractilis Perty . .
longulus (C. L. Koch) .
nudus (Berl.) ....
retaltecus (Stoll) . .
glabratus (Say) . . .
77
77
78
79
79
79
79
79
80
80
80
80
80
81
81
81
81
82
82
82
82
XII
Systematic Index
Pag.
Acai'US L.. em. Latr 82
coleoptratus L 82
piger Scop 82
,. rupestris F 82
Sp., G. Shaw 83
Belba floridana Banks 83
.. minuta Banks 83
Carabodes nitens Johnst 83
Celaeno aegrota C. L. Koch ... 83
.. coccinea C.L.Koch ... 83
.. obsoleta c. L. Koch .... 83
rhodomela C. L. Koch ... 83
Cepheus ovalis C.L.Koch .... 83
.. pertllSUS Haupt 83
CepllUS nitidllS Banks 83
ClavicepS Can. & Fanz 83
.. Mrtus Can. & Fanz 83
.. laeviusculus Can. & Fanz. . . 83
.. ruber Can. & Fanz 83
.. I'UgOSUS L. Koch 83
.. trimaculatus L. Koch ... 83
Damaeus bulbipedata Pack. ... 83
CUl'tuS Haupt 83
OnUStuS C. L. Koch .... 83
Eremaeus asperulus Berl 84
.. lineatus Thor 84
Hermannia trinebulosa Riley . . 84
Hoplophora straminea C.L.Koch . 84
testacea C. L. Koch .... 84
Murcia acaroides C. L. Koch ... 84
., ephippiata C.L.Koch ... 84
fumigata C.L.Koch ... 84
Notaspis cassideus He 84
.. castaneus He >84
conchaeformis Koll. ... 84
.. corynopus He 84
.. globulus Koll 84
.. humeralis Herm. . . . . 84
Nothrus bicolor C. L. Koch 84
Pag.
Nothrus bispiuosus C. L. Koch . . 84
canaliculatus C. L. Koch . . 84
corticalis Karp 84
,. doliaris C. L. Koch .... 84
,. ellipticuS Haupt 84
,, marinus Banks 84
,. minimus C. L. Koch ... 84
,. OvivorilS Pack 84
posticus C.L.Koch .... 84
Oppia hungarica Karp 84
Oribata aspidioti Ashm 85
CVClonotuS (ierv 85
demersa Duj 85
,. marina Duj 85
notactis Gerv 85
,. quadripilis Fitch . . . . . 85
Oribates augulatus C. L. Koch . . 85
,. atemmus C. L. Koch ... 85
facula C. L. Koch .... 85
,. globosilS C. L. Koch ... 85
picipes C.L.Koch .... 85
Oribatella obesa Banks 85
perfecta Banks 85
Oxynamba liliputana Karsch . . 85
Pelonia (irube 85
crinita Grube 85
Miosa Grube 85
tricuspidata Grube .... 85
Scutovertex concolor Banks ... 85
piloSUS Banks 85
Zetes cespitum C. L. Koch . . . 85
,. dorsatus C.L.Koch .... 85
,. flavipes C. L. Koch .... 86
gilvulus C. L. Koch .... 86
,, laevigatus C. L. Koch ... 86
,. latirostris C.L.Koch ... 86
mortacinus C. L. Koch , . 86
semirufus C. L. Koch 86
. Oribatidae
1834 Oribatei, Ant. Duges in: Ann. Sci. nat., ser. 2 v.l p. 21 1842 Carabodides,
C. L.Koch, tibers. Arach., i:3 p. 96 | 1849 Notaspidea. Hoeven, Handb. Dierk., ed. 2
p. 669 ; 1855 Oribatides, Nicolet in: Arch. Mus. Paris, T.I p. 384 etc. 1859 Carabodides,
Grube in: Arch. Dorpat. Ges., ser. 2 v.l p. 417 ; 1861 C., Fiirstenberg, Kratzm., p. 208 |
1863 Oribatidae, Gerstaecker in: Carus & Gerstaecker. Zool., r.2 p. 342 ' 1864 Oribatides,
Andersen in: Ofv.Ak.F6rh.. v. 20 p. 181 i 1875 ,,0ribatides", Donnadieu, Rech. Tetra-
nyques, p. 9 j 1876 ,,0.", Megnin in: J. Anat. et Physiol., v. 12 p. 293 1877 Oribatidae,
Murray, Econ. Ent. Apt., p. 93 | 1877 0., P.Kramer in: Arch. Naturg., .43I p. 215 j
1877 Oribatidi, Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v. 3 p. 481 ] 1877
Oribatini; Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 76 ! 1884 Oribatidae,
Michael, Brit. Orib. \ 1885 Oribatina, Berlese in: Bull. Soc. ent. Ital., ' v. 17 p. 215 , 1891
Cryptostigmata, Canestrini in: Atti 1st. Veneto, ser. 7 v. 2 p. 717 1892 Oribatidae,
Trouessart in: Rev. Sci. nat. Quest, v. 2 p. 38* 1894 Oribatoidea, Banks in: Tr. Amer.
ent. Soc., 0.21 p. 209 ] 1896 Oribatei, Oudemans in: Tijdschr. Ent., e.39 p. 53 | 1896
Cryptostigmata, Berlese, A. M. S., Crypt. II.
Acarina with bodies of various forms, those of the Imago
oftenest more or less discoidal, hemispherical, pyriform, ellipti-
cal, oblong or fusiform, but these shapes are frequently broken
by many irregularities. The bodies of the immature creatures
oftenest of the above forms, but frequently extremely irregular.
The cuticle of Imago usually fully chitinized and hard, occasion-
ally leathery. The Cephalothorax and Abdomen are usually,
but not invariably, divided by a transverse constriction between
the second and third legs. Imago usually tracheate; tracheae
simple and unbranched. Immature forms without tracheae. Stig-
mata, when existing, sunk in the acetabula of the coxae. No
eyes. A pair of Pseudo-stigmata, each bearing a pseudo-stigmatic
organ, on the dorsal surface of the Cephalothorax. Legs of five
free joints, terminated by tridactyle or monodactyle ungues
without sucker or caruncle. Maxillary palps of five free joints
(rarely four). Mandibles chelate working perpendicularly, rarely
non-chelate and serrated. Maxillary-lip (Hypostome) with dentate
organs (Maxillae) working horizontally. Genital'opening abdomi-
nal in both sexes. Not any external sexual dimorphism. Larvae
and Nymphs with soft or leathery cuticle and monodactyle
ungues.
The Oribatidae are never parasitic, they are vegetable feeders. They
are usually oviparous, sometimes ovo-viviparous, rarely viviparous. The
Larva (Lv.) emerges with three pairs of legs; it becomes a Nymph (Nph.)
and acquires the fourth pair at the first ecdysis. The Nph. undergoes three
ecdyses, at the third (the fourth if the larval ecdysis be included) it becomes
an Imago (Im.). The Lv. and Nph. are usually totally unlike the Im., but
Das Tierreich. 3. Lief. : A. D. Michael, Oribatidae. i
Oribatidae
in the genera Neoliodes, Nothrus and Hypochthonius there is often a certain
resemblance. Legs fitted for walking or crawling, except in the genus
Zetorchestes, where the fourth pair are fitted for jumping. The Nph. often,
and the Im. less frequently, carry on the dorsal surface of the Abdomen the
abdominal portions of the dorsal skins cast at all prior ecdyses, they are
usually earned concentrically ; when they are present the age of the creature
may be known by the number of cast skins. In some few species the
external layer of the cuticle of the Im. (the epiostracum) is easily detched
and is apt to be rubbed off, either wholly or more frequently partly, from
the dorsal surface; it generally bears all the markings, so that when it is
gone the cuticle of a rough species may appear smooth.
XeglF
Fig. i. Oribata dorsalis
Dorsal surface.
The anatomy of the Ex o skeleton, so far as it is necessary for the
identification of species and for understanding this work, may be very
shortly sumarized as follows. The Cephalothorax (Cephth.) and Ab-
domen (Abd.) are usually divided by a well-marked constriction or trans-
verse line between the second and
third legs; this demarcation however
is lost in the 2 genera Scutovertex
and Amerus, and almost lost in the
one species Tegeocranus vellatus. The
Cephth. of the Im. is usually firmly
anchylosed to the Abd., but that of
the immature creatures commonly
has a slight motion upon it. In the
one subfam. Phthiracarinae however
the Cephth. of the Im. is moveably
articulated to the Abd. and is capable
of being folded downward so that its
ventral surface then rests against the
ventral surface of the Abd.
The Cephalothorax (Cephth.) may be considered as divided into
two parts; an anterior part or Rostrum (Rost.) and a posterior part.
These are divided in many species by a trans-
verse line, which may probably be the horno-
logue of the cervical groove in Crustacea
(Huxley, Astacus); in many other species
however there is not any such line and it
would then not be possible to fix a posterior
limit to the Rost.
The Rostrum (Rost.) is usually a
species of hood covering and protecting the
Trophi, its sides are called the Genae and
each Gena ends anteriorly in a small free
point in some few species, which is useful
in identification. The Rost. bears 2 curved
hairs, called the Rostral hairs (Rost. hairs).
The hood of the Rost. in partly closed
below by the articulated maxillary-lip (or
Hypostome), it is formed by the coalesced
bases of the maxillae, but the distal portions of the maxillae form large
projecting organs springing from, but at right-angles to, the lip. These
Kasth.Xamh.
. Anal covers
Dorsplate
Fig. 2. Oribata orbicularis ( K i\).
Ventral surface.
Oribatidae
organs are curved and blade-like, and bear teeth on their anterior edges;
they move horizontally and are highly developed in Hoploderma. The
maxillary-lip bears the Palpi, which are very mobile, Duges has called them
fusiform" ; the basal joint is usually the shortest and the second the longest.
In the Phthiracarinae there are only four joints,
apparently from the coalescence of the second and third.
The Mandibles (Mdb.) are of two joints and
chelate, the moveable arm of the chela is articulated
below the prolonged and narrow anterior end of the
first joint, and moves perpendicularly.
The posterior part of the Cephth. bears
two outfoldings of the dorsal cuticle, which when
well developed are large longitudinal blades usually
standing nearly on edge ; but they may be horizontal
or nearly so ; or they may be mere thickened ridges,
or entirely absent; they are called the Lamellae
(Lam.): they vary greatly in size and form. The
two Lam. are frequently joined at or near their an- Fig. 3. oribata piriformis.
terior ends by a transverse blade or ridge also formed Mdb - (SO ' l) -
by an out-folding of the dorsal cuticle which is called the Translamella
(trlam.). When the Lam. continue anteriorly beyond the Trlam. their
lower edges are not joined to the dorsal cuticle, thus they form free points,
which may be very long; these are called the
cusps of the Lamellae (C. lam.). The cusp
often exists although there is not any Trlam. The
cusp, or the Lam. itself if there be not any cusp,
almost always bears at or near its anterior end
a hair, usually long, which is called the Lamellar
hair (Lam. hair). In species where there is not
any Lam. these Lam. hairs generally persist, arising
from that portion of the dorsal cuticle, which would
be folded outward to form the anterior end of
the Lam., if it existed. The portion of the dorsal
Fig. 4. oribata piriformis ( I8 ,,). surface of the Cephth. bounded by the Lam. laterally,
Cephth., dorsal aud lateral surface, the Trlam. anteriorly, and the Abd. posteriorly,
is called the Dorso-vertex.
The ,,Tectum" of Nicolet, and of many subsequent authors following
him, is an imaginary organ which has not any existance in reality ; although
appearances are very deceptive in this respect; a few sections will at once
convince any-one of the error. It was supposed to be a sort of chitinous
shelf springing from the hinder part of the Cephth. where it adjoins the
Abd. and over-hanging the whole or part of the Cephth., and either standing
quite free or anchylosed to its surface; and this shelf was supposed to
have upturned lateral edges called the wings of the Tectum". In com-
paring the writings of these writers with this book, or with the creatures, it
must be remembered that where the Tectum is said to have wings these
wings are the Lam., and the Tectum is the dorsal surface of the Cephth.
itself, i. e. the Dorso-vertex, and the Trlam., when present. - - Where the
Tectum is said to be without wings it represents the Lam., which in these
cases are usually large and horizontal.
At or near each outer posterior corner of the Dorso-vertex; and con-
sequently close to the Abd. a very short chitinous tube directed upward.
4 Oribatidae
or somewhat laterally, projects from the dorsal surface of the Cephth.;
these two tubes look like stigmata and were formerly supposed to be so;
they are now known as the Pseudostigmata (Psstg.), from each a hair-
like or flesby organ projects, known as the Pseudostigmatic organ
(Psstg. org.) (Fig. 1, 4). The form of this organ varies greatly in different
species, and is most useful in identification; the whole apparatus, which is
peculiar to the Oribatidae and some few other Acari, is probably an organ
of hearing.
From the Dorso-vertex, between the Psstg., and consequently between
the Lam., arise a pair of hairs, often very large; these are the Inter-
lamellar hairs (Intlam. hairs) (Fig. 4).
From the sides of the Cephth. certain blade-like longitudinal out-
foldings of the article, not unlike the Lam., are frequently found; there are
generally two or more on each side when they are present ; they are some-
times prolonged anteriorly and form cusps; their office is to protect the
two anterior pairs of legs, which can frequently be bent and pla.ced so as
to lie entirely in the deep cavities between these organs which are called
Tectopedia (Tectp.) (Fig. 2, 4).
The Abdomen (Abd.) is usually much larger than the Cephth. its
dorsal surface is generally wholly covered by a thick chitinous plate, which
usually bends over a little and embraces the edge of the ventral plate. In
the Oribatinae the anterior part of each lateral edge of this dorsal plate
projects considerably beyond the ventral plate, forming a wing-like organ,
more or less flexible, which is capable of slight movement from the action
of the muscles of the Abd., not of any special muscles of its own. These
wing-like processes are called the Pteromorphae (Ptm.) (Fig. 1, 2). -
The genital and anal openings are large and placed in the median line of
the ventral plate ; each is closed by a pah' of chitinous folding-doors, called in
this treatise the genital covers and the anal covers. - - The expulsory
vesicles are two paired sacks in the Abd. of the Nph. of some Oribatidae
and in certain other Acari (,,Excretionstaschen"), each of which discharge
an excretory fluid to the enterior through a pore in the cuticle.
The legs are always of five free joints, which are called the coxa,
femur, genual, tibia, and tarsus. In some cases the coxae and femora
( or one ^ them) nave a thin blade-like border on their
lower edges. The tarsus is ended by a monodactyle
(mond.) or tridactyle (trid.) unguis (ung.) without
sucker or caruncle. The bid. ung. may be either homo-
dactyle (homd.), with all three ung. similar, orhetero-
dactyle (hetd.), with the central ung. different from the
lateral ones. In one single known instance, i. e. the'
European specimens of Nothrus anauniensis, the lateral
ung. on one side is abortive leaving a bidactyle (bid.),
hetd. ung. The principal parts useful in the identification
of species (in addition to the general shape and appea-
rance) are the Psstg. org., the Lam., the Intlam. hairs,
the Trlam., the Ptm., and the ung.
Oribatidae have been found in all parts of the world.
Oribata mollicoma. , . , , ,
Lee 4 f 180 ") which have yet been searched for them; they vary extremely
little in different countries the distribution being remarkably wide.
Xo genera are known from Tropical America which are not known in Europe, and
the species from Franz- Joseph- Land are either identical with, or closely allied
Oribatidae, Oribatinae 5
to some of those found in Central Europe. Their principal habitats are in moss, under
bark of trees, on leaves of trees either growing or fallen, on lichens and fungi,
and under stones. They are chiefly terrestrial but some are known which lead aquatic
lives crawling on water-weeds, not swimming. One doubtful species has been de-
scribed by Dujardin as marine, but it is not sufficiently described and possibly was
not one of the Oribatidae at all.
This work only includes recent species; it would make but little difference as
regards the Oribatidae if it were otherwise, as there are not any recorded species
which have any claim to be considered fossil except four found in amber. The
places, or strata in which the amber was found are not mentioned, but supposing it
to have been true tertiary amber these creatures would of course belong to that
period. The species are: Oribates convexulus C. L. Koch & Berendt. Nothrus
sulcatus Karsch, Nothrus kuhli Karsch, Nothrus punctulum Karsch. The descriptions
are somewhat imperfect, indeed it would scarcely be possible that they should be
otherwise with such minute creatures imbedded in amber.
In the localities given for the various species most of those in the Netherlands
have been taken from Oudemans' n List of Dutch Acari" (in: Tijdschr. Ent.. 1896, v. 39
p. 53); and many of those in Sweden from: C. H. Andersen's n Bidrag till Kannedomen
om Nordiska Acariden" (in: Of. v. Ak. Forh., 1863 (1864), v.20 p. 181). It is doubtful
whether the identification of species in these papers is quite reliable.
I regret having felt compelled to place so many of W. Banks' American species
in the doubtful list, but that gentleman was unable to publish any illustrations, and
the descriptions rely greatly upon Nicolet's erroneous anatomy, so that I do not think
that it would be possible to indentify the species from many of them; I understand
however that the author proposes publishing illustrations of some at least of them
which will probably remove the difficulty.
7 subfamilies, 20 good and 3 uncertain genera. 199 good and 115 doubtful
species, 2 varieties.
Key to the subfamilies.
f Abd. with chitinous wing-like expansions . A. Subfam. Oribatinae ... p. 5
| Abd. without wing-like expansions (Apterogasterea) 2.
/ Cephth. anchylosed to Abd. Ventr. plate
anchylosed to dors, plate 3.
9. i Cephth. hinged to Abd. and folding down
on its ventr. surface. Ventr. plate not
anchylosed to dors, plate GK Subfam. Phthiracarinae . p. 77
i Mdb. rod-like, serrated, not chelate . . . B. Subfam. Serrariinae . . p. 26
\ Mdb. chelate 4.
i Legs arranged first 3 pairs near together,
pair 4 distant, jumping organs . . . . C. Subfam. Zetorchestinae . p. 27
4 f Legs arranged first 2 pairs near together,
last 2 pairs near together. Pair 4
I crawling organs 5.
J With Lam D. Subfam. Notaspidinae . p. 28
5 \ Without Lam. 6.
J Legs thin, much longer than body . . . . E. Subfam. Damaeinae . . . p. 52
\ Legs short and thick F. Subfam. N"othrinae . . . p. 61
A. Subfam. Oribatinae
1884 Pterogasterinae, Michael, Brit. Orib., v. 1 p. 202 i 1885 Subfam. Oribatidae,
.Berlese in: Bull. Soc. ent. Ital, v. 17 p. 127 | 1891 0., Canestrini in: Atti 1st. Veneto,
ser.7 v.2 p. 717 j 1892 Oribatinae, Trouessart in: Rev. Sci. nat. Quest, v.2 p. 38* [ 1896
.Fam. Pterogasterinae, Berlese, A. M. S., Crypt. II p. 61.
Abd. with chitinous wing-like expansions (Pteromorphae, Ptm.).
Oribatinae, Pelops
2 genera, 55 good and 19 doubtful species.
Key to the genera.
i Mdb. broad at base, suddenly becoming and continuing
-| slender and rod-like, chelae small; Intlam. hair
usually spatulate and horizontal ........ 1. Gen. Pelops .
f Mdb. thick and stout, not slender nor rod-like in any
\ part; Intlam. hair usually setiform and erect . . 2. Gen. Oribata
[>. 6
p. 9
1. Gen. Pelops C.L.Koch
1835 Pelops (Im.) -f- Celaeno (Nph.) (part), C. L. Koch, C. M. A., fasc. 2. 3
P. (Im.) -f- C. (Nph.) (part.), C. L. Koch, Ubers. Arach., v. 3 p. 102, 108.
Mdb. broad at base, suddenly becoming stiliform, termi-
nated by minute chelae. Intlam. hair usually large and spatu-
late and horizontal ; usually inserted below a rectangular (square
or oblong) projection of ant. margin of Abd. Ung. trid.
10 species.
Key to the species.
, With a rectangular projection from ant. margin of Abd. (Div. I.
p. 6) - 2.
| Without any rectangular projection from ant. margin of Abd.
(Div. II. p. 8) 9.
{Dors, of Abd. and Ptm. deeply pitted 3.
Dors, of Abd. and Ptm. not pitted 4.
/ Almost the whole median line of
Dors, of Abd. occupied by 2
j shallow depressions 5. P. occultus .
* No such depressions 7. P. torulosus
f Hairs on Abd. all setiform .... 4. P. fuligineus
\ Hairs on Abd. all or partly spatulate 5.
Spatulate hairs on Dors, of Abd. - 6.
Spatulate hairs on actual post, margin only 8.
No true Lam., 4 rows of spatulate hairs on Dors, of
Abd. 7.
Lam. well marked and meeting anteriorly, 2 rows
I only of spatulate hairs near lat. margin .... 8. P. uraceus .
J Psstg. org. clavate 1. P. acromius
| Psstg. org. falciform 3. P. farinosus
| 2 pairs of spatulate hairs on post, margin .... 6. P. tardus . .
1 pair of spatulate hairs on post, margin .... 2. P. auritus .
p. 8
p. 8
p. 7
p. 8
p. 6
p. 7
p. 8
p. 7
/ Intlam. hair large, broadly falciform; Dors, of Abd.
smooth, very finely punctured; 8 or 10 hairs on post.
' margin not spatulate ............ 9. P. glaber . . .
I Intlam. hair small and fine; Dors, of Abd. coarsely
I pitted; 2 pairs of spatulate hairs on post, margin 10. P. phaeonotus
p. 8
p. 9
Div. I.
Sp. 18. With a rectangular projection from ant. margin of Abd.
1. P. acromius (Herm.) 1804 Notaspis acromios (corr. acromius), Hermann,
Mem. apt., p. 91 t. 4 f . 1 1855 Pelops a., Nicolet in : Arch. Mus. Paris, v. 7 p. 425 t. 3
f. 1 I 1877 P.a., Canestrini&Fanzagoin: Atti 1st. Veneto, ser. 5 w.4 p. 78 (var.?) | 1884
P.O., Michael, Brit. Orib., v. 1 p. 208 t. 1 f. 1 12; t. 2 f. 13 | 1835 Celaeno spinosa,
C. L. Koch, C. M. A., fasc. 3 1. 17 (Nph.).
Im. Black, nearly circular, broad posteriorly. Abd. large, broad poste-
riorly, flattish, rough, with irregular vermiform ridges. The ridges look
Pelops 7
black but are secreted matter not present when the creature emerges. Cephth.
bulbous posteriorly. Rost. pointed. Rost. hair fine. No true Lam. or Trlam.
Psstg. hidden by ant. margin of Abd. Psstg. org. moderate length,
gradually clavate, slightly recurred. Int. lam. hair very large, light,
spatulate, horizontal, directed forward passing Rost. Tectp. 1 with free
points. Ptm. short, rough, like Dors, of Abd. Ant. margin of Abd. has
a square projection with undulating margin of 3 lobes. Dors, of Abd. has
elliptical depressed, slightly convex space smooth and light. 4 rows of
spatulate hairs on dors, of Abd. and a row of similar hairs round post,
margin. Genital covers larger than anal. - - Nph. Red-brown, or yellow-
brown. Cephth. yellow, hairs white. Pyriform, Rost. narrow.- Cephth. divided
into spaces by regular raised ridges. Psstg. org. almost globular on short
peduncles. Intlam. hair thick, slightly spatulate. Abd. has lateral portions
raised, central area slightly convex; covered by irregular wavy soft ridges
running backward and inward on lateral, and transversly on central area ; a
row of large, curved, spatulate hairs round margin. Lv. like Nph., but
has only 2 pairs of the marginal hairs. Im. L. ca. 650 |u.
On Oak and white thorn trees and among fir needles and elsewhere. Algeria,
England, France, Germany, Italy, Netherlands, Switzerland.
2. P. auritus C. L. Koch 1840 P. a., C. L. Koch. C. M. A., fasc. 30 1. 11.
Im. Blackish-brown. Intlam. hair awl-shaped. Psstg. org. fusiform.
A square projection from ant. margin of Abd. 2 thick clavate hairs
on post, margin directed straight backward. No others on Abd. Abd.
as broad as long. L. ,,mittelgross".
In moss on trees. Germany.
3. P. farinosus Nic. 1855 P.f., Nicolet in: Arch. Mus. Paris, t>. 7 p. 425 t. 3
f. 2 1884 P.f., Michael, Brit. Orib., v.l p. 212 t. 1 f. 13.
Very like P. acromius, but smaller and with Psstg. org. falciform in-
stead of clavate, secreted ridges often remain white instead of turning black.
- Im. L. ca. 550 |u.
England, France. Probably same distribution as P. acromius.
4. P. fuligineus C. L. Koch*) 1835 Celaeno plicata, C. L. Koch, C. M. A., fasc. 3
t. 18 (Nph. !) ; 1840 Pelops acromios (non Notaspis acromios Hermann 1804!), C.L.Koch,
C. M. A., fasc. 30 t. 9 | 1844 Oribataa., Gervais in: Walckenaer, Ins. Apt., v.3 p. 258
1840 Pelops fuligineus, C.L.Koch, C. M. A., fasc. 30 1. 10 ! ?1844 Pelops hirsutus, C.
L.Koch, C. M. A., fasc, 38 t. 15 | 1855 P. laevigatus, Nicolet in: Arch. Mus. Paris, v. 7
p. 426 t. 3 f. 3 ! 1884 P. L, Michael, Brit. Orib., v.l p. 213 t. 2 f. 1 7 j ?1882 P.
sculptus, Haupt in: Ber. Ges. Bamberg, v. 12 p. 29.
Im. Black, dull, smooth but not polished ; shortly after emerging Dors.
of Abd. becomes coated with white secretion which splits irregularly all over
and sometimes rubs oif. Rost. sharp. Intlam. hair as in P. acromius.
Psstg. org. like Intlam. hair in form but smaller. Abd. almost circular,
*) According to the synonyma enumerated above by the author and in
accordance with the ,,Regeln fur die wissenschaftl. Benennung der Thiere" ( 2a: ,,Namen,
welche nur auf einem unentwickelten Zustand .... basieren, sind zulassig, falls die
Kennzeichnung zur Wiedererkennung der betreffenden systematischen Einheit ausreicht.")
the name of this species is P. plicatus (C.L.Koch). The Editors.
I am not able to agree in this opinion. As the immature forms of the Ori-
batidae are usually wholly unlike the Imagines I do not think that specific names founded
upon them are permissible under the rules referred to, and they are expressly forbidden
by the rules of nomenclature of the zoological congress at Moscow. The Author.
8 Pelops
slightly narrowed anteriorly. Ptm. longish with straight ant. margin. Central
projection of ant. margin of Abd. broad, tri-lobed. No clear depressed
space on Dors, of Abd. 4 rows of fine hairs on Dors, of Abd., some of
those on post, margin thicker. - - Nph. Red or yellow, legs darker, hairs
whitish; somewhat like Nph. of P. acromius, but hairs at sides of Abd.
thick but not spatulate. 2 very long hairs on post, margin. - - Im.
L. ca. 550 |u.
In fir-woods, moss etc. England. France, Germany, ? Netherlands.
5. P. occultus C. L. Koch, Xic, 1835 P. o., C. L. Koch, C. M. A., fasc. 2 t. 15
1855 P.O., Mcolet in: Arch. Mus. Paris, v. 7 p. 427 t. 3 f. 5 | 1887 P.O., Berlese, A.
M. S.. fasc. 35 nr. 4.
Im. Dark brown, coarsely punctured, Psstg. org. with thin peduncle
and clavate-fusiform hairy head (Berlese draws, them hairy but describes
them as ,.nudis"). A square projection from ant. margin of Abd. Abd.
with almost whole median line occupied by 2 shallow depressions
of equal width each bordered by a ridge; post, depression oblong, more
than twice as long as ant, which is oval. Abd. hairless. L. 430 \JL.
France, Germany, Italy.
6. P. tardus C. L. Koch 1835 P. t., C. L. Koch, C. M. A., fasc. 2 t. 16 j 1884
P. auritus, Berlese, A. M. S., fasc. 15 nr. 8.
Im. Very black, rough. Square projection of ant. margin of Abd.
present. Intlam. hair spatulate. Psstg. org. with thin peduncle and fusi-
form hairy head. Abd. smooth but usually covered with secretion in irre-
gular undulations. No hairs on Dors, of Abd. 2 pairs of spatulate hairs on
post, margin, the inner pair curved inward, the outer outward. Abd. a very
short ellipse. L. ca. 500 ju (350 ju in Berlese, A. M. S., Crypt. II p. 68).
In moss. Germany, Italy.
7. P. torulosus C. L. Koch 1840 P. t ., C. L. Koch, C. M. A., fasc. 30 1. 13 | 1877
P. t., Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 79 | 1896 P. t., Berlese, A.
M.S.. Crypt. II p. 68 | 1855 P. variolosus, Nicole tin: Arch. Mus. Paris, v.7 p. 427 t.3 f. 4.
Im. Black. Psstg. org. clavate. Abd. deeply pitted; pits large,
close together, irregular in shape; those on Ptm. smaller than those on
Dors, of Abd. Square projection from ant. margin of Abd. narrow, white.
3 pairs of clavate hairs on post, margin caducent. L. 500 m.
France, Germany, Italy.
8. P. uraceus C. L. Koch 1840 P. ., C. L. Koch, C. M. A., fasc. 30 1. 12 ' 1884
P. u.. Berlese, A. M. S., fasc. 15 nr. 7.
Im. Yellow-brown, roughish, strongly punctured. Lam. blades on edge,
touching anteriorly. Intlam. hair large fusiform. Psstg. org. long, with
thin peduncle and clavate, subfusiform head. Square projection of ant.
margin of Abd. large. Abd. long-shaped narrow, much narrowed and truncated
anteriorly; rounded posteriorly. A clear smooth oval space near ant. margin
of Abd. 1 pair of clavate hairs on hind margin curving outward; 2 pairs
on Dors, of Abd. near lat. margin and 1 near post, margin, their ends pro-
jecting between the pair on the actual margin. L. ca. 500 JLI.
On bushes, moss etc. Germany, Italy, Switzerland.
Div. II.
Sp. 9, 10. Without any rectangular projection from ant. margin of Abd.
9. P. glaber G. &R. Can. 1882 P. g., G. & R. Canestrini in: Atti 1st. A'eneto.
ser. 5 .8 p. 915 | 1895 P. g., Berlese, A. M. S., fasc. 74 nr. 2.
Pelops, Oribata 9
Im. Reddish - brown, smooth, very finely punctured. Resembles
P. fuligineus, but no hairs on Dors, of Abd. 8 or 10 round hind margin;
a small elliptical space of thin, light-colored polished chitin in median line
not far from ant. margin of Abd. No square chitinous projection from
ant. margin of Abd. Psstg. org. moderately clavate. L. 700 850 ju.
Italy.
10. P. phaeonotus C.L.Koch 1844 P.p., C.L.Koch, C. M. A., fasc. 39 t. 23
j 1877 P.p., Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 0.4 p. 80 | 1884 P.p.,
Michael, Brit, Orib., v.l p. 216 t.2 f.8 12 ; 1887 P.p., Berlese, A. M.S., fasc. 35 nr.3.
Im. Red-brown or black. No secreted matter. Shape pyriform, very
narrow anteriorly. Lam. large ; C. lam. nearly reaching tip of Rost. Psstg.
org. clavate, rather long; Psstg. sunk in deep depressions. Intlam. hair
small and fine. Abd. bordered by thin edge but without square pro-
jection from ant. margin of Abd. Ptm. attached by anterior part of
their edges only. Dors, of Abd. with depressed, oval, smooth, light space
as in P. acromius ; rest of Dors, of Abd. with depressed areolations, no hairs
on it, 2 pairs of spatulate hairs on post, margin. L. ca. 450 |u.
In moss. England, Germany, Italy.
2. Gen. Oribata Latr.
1802 Oribata, Latreille in: Buffon, Hist, nat., ed. Sonnini, Ins. .3 p. 65 1803
O., Latreille in: Nouv. Diet, r. 16 p. 349 : 1811 0., Olivier in: Enc. meth., Art. Oribate
| 1855 0., Nicolet in : Arch. Mus. Paris, v. 7 p. 427 j 1859 0., Grube in : Arch. Dorpat.
Ges., ser. 2 v.l p. 417 1884 O., Michael. Brit. Orib., B.I p.218 [ 1891 0., Canestrini in:
Atti 1st. Veneto, ser. 7 v.2 p. 579 ' 1892 0., Trouessart in: Rev. Sci. nat. Quest, v.2 p. 48
| 1826 Oribata -4- Galumna, Heyden in: Isis. p. 612 ' 1834 Oribates, Duges in: Ann.
Sci. nat., ser. 2 v.l p. 21 ' 1877 0., Canestrini & Fanzago in: Atti 1st, Veneto, ser. 5 p. 4
p. 81 | 1885 0., Canestrini, Prosp. Acarof., v.l p. 6 ] 1885 0. (0. -+- Achipteria -J- Sphae-
rozetes), Berlese in : Bull. Soc. ent. Ital, v. 17 p. 127, 128 | 1896 0., Berlese, A. M. S.,
fasc. 74 nr. 7 j 1835 Oribates -f Zetes (part.) -f Murcia (Nph.) (part.), C. L. Koch, C. M. A.,
fasc. 2 etc. ; 1895 Oribata-}- Oribatella -\- Oribatodes, Banks in: Tr. Amer. ent. Soc., v. 22
p. 5. 8, 10.
Mdb. of the form usual in the family (Fig. 3). Intlam. hair never
spatulate.
45 good and 19 doubtful species.
Key to the good species.
Ung. trid. (Div. I. p. 12) 2.
Ung. mond. (Div. IL, Gen. Oribatodes Banks, p. 23) 40.
/ Having a Trlam. (Subdiv. la. p. 12) 3.
\ Without any Trlam. (Subdiv. Ib. p. 18) 21.
Trlam. a blade on edge 4.
Trlam. a mere line 10.
Lam. hair, Intlam. hair and Rost. hairs setiform or absent 5.
Lam. hair. Intlam. hair and Rost. hairs pectinated 8.
/ Ptm. normal, truncated anteriorly, running posteriorly to about middle of Dors.
of Abd. 6.
' Ptm. projecting forward forming a point
of moderate length, but only running a
short distance posteriorly. L. 800 ju 6. O. gilvipes p. 13
{Psstg. org. fusiform, horizontal, directed
forward. L. 400 |u 16. O. punctum p. 16
Psstg. org. clavate 7.
!
10 Oribata
Abd. round. L. 650 n 13. O. orbicularis p. 15
Abd. pyriform. L. 650 |U 15. O. piriformis p. 16
Dors, of Abd. bearing a series of long hairs
near margin 9.
Dors, of Abd. hairless. L. 450 |ui . . . 17. O. rubens p. 17
C. lam. truncated; Lam. hair set in middle
of ant. truncated edge. L. 630 |u . 5. O. fuscipes p. 13
C. lam. long and pointed ; Lam. hair springing
from a shoulder near base. L. 540 \i 18. O. setosa p. 17
Dors, of Abd. hairless 11.
Dors, of Abd. bearing hairs -18.
f Ptm. strap-like; aquatic. L. 320 \n , . 19. O. sphagni p. 17
10
I
Ptm. normal; terrestrial 12.
J Psstg. org. clavate 13.
| Psstg. org. setiform or rod- like 17.
{Lam. hair and Intlam. hair setiform 14.
Lam. hair and Intlam. hair pectinated.
L. 500 |U 3. O. depauperata . . . . p. 13
{Femur of leg 2 greatly enlarged at distal
end. L. 500 n 14. O. pallidula p. 16
Femur of leg 2 normal 15.
{Ptm. very small, confined to quite the ant.
part of Abd., Psstg. org. pointed
anteriorly. L. 1 mm 11. O. microptera p. 15
Ptm. normal; Psstg. org. rounded anteriorly 16.
{Host, slender; C. lam. long; legs long; no
light patch on Dors, of Abd. L. 480 n 8. O. gracilis p. 14
Rost. obtuse; C. lam. short; legs moderate;
light patches on Dors, of Abd. L.SOOju. 9. O. lapidaria p. 14
{Abd. globular; Psstg. org. rod -like.
L. 1.05mm 7. O. globula p. 14
Abd. long, truncated posteriorly. Psstg. org.
setiform. L. 1.13 mm 2. O. clypeata p. 12
io / Hairs on Dors, of Abd. very long 19.
\ Hairs on Dors, of Abd. short and fine 20.
Lam. nearly as long as Rost.; Femur 1, 2,
with blades; leg 4 hardly passes post.
end of Abd. L. 500 (a 12. O. mollicoma p. 15
Lam. much shorter than Rost.; Femur 1, 2,
without blades; whole Tarsus of leg 4
passing post, end of Abd. L. 750 |n 4. O. edwardsi p. 13
(Rost. hairs, Lam. hair and Intlam. hair
serrated. L. 700 |n 1. O. antarctica p. 12
1 Rost. hairs, Lam. hair and Intlam. hair
setiform. L. 600 |U 10. O. lucens p. 15
JLam. large, horizontal, attached by post,
edge only, the 2 near together, only a
~JL . narrow cleft between. (Sect. Ib 1. Gen.
Oribatella, Banks) 22.
v Lam. normal on absent (Sect. Ib 2) 29.
iPtm. truncated anteriorly, not projecting
forward beyond ant. margin of Abd. 23.
Ptm. projecting forward nearly to tip of
Rost. 28.
Oribata 11
(Lam. deeply excavated anteriorly so as to
end in 2 long free points of about equal
length 24.
' Lam. truncated anteriorly 27.
Dors, of Abd. with a row of hairs near
margin; terrestrial 25.
I Dors, of Abd. hairless ; aquatic. - L. 550 \JL 21. O. aquatica p. 18
Ptm. reticulate with a dentate ant. margin.
- L. 600 M. 26. O. quadridentata .... p. 19
Ptm. not reticulate nor with dentate margin 26.
/ Psstg. org. clavate; hairs round Dors, of
Abd. about 12. L. 580 JLI 25. O. quadricornuta . . . . p. 19
26 ' Psstg. org.. setiform. strongly curved toward
!
median line ; hairs round Dors, of Abd.
about 18. L. 450 |u 22. O. berlesei p. 18
( Outer lat. edge of Lam. strongly and
27 | irregularly dentate. L. 390 |u . . . 20. O. americana p. 18
1 Outer lat. edge of Lam. plain. L. 210 n 27. O. tecta p. 20
{Dors, of Abd. with transverse furrows.
L. 420 in 23. O. languida p. 18
Dors, of Ab. without transverse furrows.
L. 650 |LI 24. O. ovalis p. 19
I Ptm. projecting forward nearly to tip of Rost. 30.
29 < Ptm. truncated anteriorly not projecting
beyond ant. margin of Abd. 34.
j Projection of Ptm. pointed anteriorly. -
30 | L. 520 n 35. O. intermedia p. 21
I Projection of Ptm. rounded anteriorly 31.
j Abd. with 2 large chitinous points projecting
31 < from centre of hind margin L. 650 ju 37. O. mucronata p. 22
\ Abd. without such points 32.
| Psstg. org. short clavate. L. 600 |u . . 29. O. alata p. 20
32 < Psstg. org. with long slender recurved
peduncle and fusiform head 33.
{Dors, of Abd. with fine longitudinal winkles.
- L. 850 |u . . 39. O. rugifrons p. 22
Dors, of Abd. plain. L. 700 n .... 33. O. dorsalis p. 21
f Dors, of Abd. hairless 35.
\ Dors, of Abd. bearing conspicuous hairs 39.
{Abd. and Cephth. plainly divided 36.
No demarcation between Abd. and Cephth.
- L. 900 |n 31. O. centro-americana . . ]t. 21
/ Lam. hair strongly serrated and nearly
twice as long as Cephth. Intlam. hair
36 ' sharply recurved. L. 430 |u . . . . 28. O. agilis p. 20
I Lam. hair, simple, setiform; Intlam. hair
I not recurved 37.
{Lam. much shorter than Rost. 38.
Lam. nearly as long as Rost.; Psstg. org.
fusiform, horizontal, directed forward.
- L. 320 n 30. O. avenifera p. 20
1 Psstg. org. short, clavate. pointed anteriorly,
directed forward. L. 350 |u . . . . 32. O. cuspidata P- 21
Psstg. org. long, recurved, with slender
peduncle and fusiform head. L. 600 |u 36. O. lucasi p. 22
12 Oribata
2 longitudinal rows of hook-shaped hairs on
Dors, of Abd. and a bordering row;
Psstg. org. with short peduncle and
. almost globular head. L. 440 |n . . 34. O. halleri p. 21
4 longitudinal rows of ordinary hairs on Dors.
of Abd. and a bordering row, Psstg. org.
with long peduncle and recurved fusiform
head. L. 450 ILI 38. O. peloptoides p. 22
Dors, of Abd. bearing hairs 41.
Dors, of Abd. hairless 43.
Ptm. joined anteriorly by a chitinous
bridge; a crescentic indentation in
, 1 j centre of post, margin, C. lam. hori-
zontal, truncated. L. 500 |l .... 45. O. rileyi p. 24
Ptm. not joined; no indentation of post.
margin; C. lam. vertical, pointed 42.
Dors, of Abd. with an irregular network
of ridges ; hairs on Dors, of Abd. long,
stiff, serrated, on post, margin clavate.
42 < - L. 500 n 42. O. mirabilis . . . . . . p. 23
Dors, of Abd. smooth; hairs on the same
short and simple, on post, margin
setifonn. L. 420 ILI 44. O. parmeliae p. 23
Psstg. org. short sessile, fusiform, directed
43 { horizontally forward. L. 300 |u . . . 41. O. fusigera p. 23
Psstg. org. long, recurved 44.
Psstg. org. pectinated along distal part of
44 \ ant. (outer) edge. L. 390 (LI . . . . 43. O. monodactyla . . . . p. 23
Psstg. org. not pectinated. L. 600 |ui . . 40. O. dentata p. 23
Div. I. Sp. 1-39. Uug. trid.
Subdiv. la. Sp. 119. Having a Trlam.
1. O. antarctica Michael 1895 0. a., Michael in: Mt. Mus. Hamburg, v. 12
p. 71 textf.
Im. Very like 0. gracilis but not identical, smooth, not quite so slender
as 0. gracilis nor legs quite so long. Lam. blades, much wider anteriorly
than posteriorly. C. lam. very long, standing quite free. Rost. hairs,
Lam. hair and Intlam. hair serrated. Psstg. org. with very short
peduncle and small, almost globular head. Legs of medium length, femora
of legs 3 and 4 with blades. Ung. trid. Tectp. 1 with long free points.
Ant. margin of Abd. convex in centre concave at sides. A few very fine
hairs on Dors, of Abd. L. ca. 700 fi.
South Georgia (Antarctic Ocean).
2. O. clypeata Nic. 1855 0. c., Nicolet in : Arch. Mus. Paris, c. 7 p. 437 t. 5 f. 7.
Im. Black, highly polished, legs brown. Lam. blades on edge, slightly
undulated; C. lam. very long. Trlam. a mere line strongly concave
forward. Tectp. 1 forming long free point anteriorly, bearing a hair.
Psstg. org. setiform, long. Abd. hairless broadest in centre, narrowed ante-
riorly and posteriorly but much more suddenly posteriorly; hind margin
truncated and much narrower than ant. margin of Abd., which is sinuated,
convex forward in centre, concave posteriorly. Ung. trid. L. 1.13 mm.
Under stones. France.
Oribata 13
3. O. depauperata Berl. 1887 Oribates depauperatus, Berlesc, A. M.S.. fasc. 35
nr. 6 ?1895 O.pusillus, Berlese, A. M. S., fasc. 77 nr. 3.
Im. Smoky-brown to chestnut ; polished Cephth. short. Lam. shortish
blades on edge ; Lam. hair springs from lower corner of Lam. C. lam.
rather long, tooth-like; Tectp. I with long free point. Host, hairs
minute. Lam. hair, Intlam. hair and a nearly terminal hair on Tectp. I.
pectinated. Psstg. org. shortish, strongly clavate, setose. Abd. a very short
ellipse, sub-spherical, hairless. Genual and Tibia 1. 2. each with one
strong spine. Ung. trid. L. ca. 500 |u.
In moss. Italy.
4. O. edwardsi Xic. 1855 0. Edwardsii, Nicolet in : Arch. Mus. Paris, c. 7
p. 438 t, 5 f. 5 | 1884 0. E., Michael, Brit. Orib., v. 1 p. 229 t. 4 f. 10 17 | 1887
Oribates E., Berlese, A. M. S., fasc. 43 nr. 5.
Im. Dark brown, dull, Dors, of Abd. punctured. Cephth. large ; Rost.
obtuse; Lam. very short blades on edge, not half the length of Cephth.;
no C. lam. Lam. hair large, setiform. Psstg. org. moderate length, clavate,
sometimes recurved. Legs long and thin, no blades to front legs; Ung.
large, central one very thick. Dors, of Abd. not much arched. Ptm. not
running for back; a row of large, flexible hairs of various lengths round
Dors, of Abd. and 2 short rows near centre of same. L. ca, 750 |u.
England, France, Italy.
5. O. fuscipes C. L. Koch*) 1835 Zetes ephippiatus, C.L.Koch, C. M. A., fasc. 3
t 7 (Nph.!) | 1836 Oribates fuscipes. C.L.Koch, C. M. A., fasc. 6 t. 8 | 1884 Oribata f.,
Michael, Brit. Orib., v. 1 p. 241 t. 7 f. 1, 2 | ?1895 Oribatella bidentata, Banks in: Tr.
Amer. ent. Soc., v. 22 p. 8.
Im. Dark red-brown, highly polished. Cephth. much narrower than
Abd. Lam. large, their ant. portions almost horizontal, not on edge.
C. lam. long, truncated, horizontal. Lam. hair long, curved, serrated;
springing from centre of truncated edge of C. lam. Trlam. a short wide
blade nearly horizontal; a strong pectinated hair on each gena the 2
hairs nearly meet. Psstg. org. long, almost upright, thin, rough, slightly
thickened toward ends. Intlam. hair serrated. Femora of first 2 pairs of
legs with broad blades; in pair 2 produced forward forming a large
obtuse point. Abd. a very long ellipse. Ant. margin of Abd. rounded;
a row of fine sparse hairs near margin ; a few others. Ptm. fade into Dors,
of Abd. posteriorly without clear demarcation. Nph. Yellowish-red or
tawny ; internal organs show as patches of dark color. Cephth. short, much
narrower than Abd. Psstg. org. long, filiform, doubly curved. Intlam. and
Rost. hairs setiform. Abd. broad, shield-shaped. Dors, of Abd. arched, with
fine granulations, but no hairs except at post, margin. - - Im. L. ca. 630 |u.
In moss. England, Germany, ?New York State America.
6. O. gilvipes C. L. Koch 1840 Oribates g., C.L.Koch, C.M.A., fasc. 30 1. 14.
Im. Black, polished. Lam. broad, undulated, slightly convex inward,
approaching very near each other anteriorly. C. lam. very long, nearly
reaching tip of Rost. horizontal, near together. Trlam. short but
*) According to the synonyma enumerated above by the author
and in accordance with the ,,Regeln fur die wissensch. Benennung der Thiere"
the name of this species is: O. ephippiata (C. L. Koch). The Editors.
See my note p. 7. The Author.
14 Oribata
strong. Psstg. org. long, filiform. Rost. rather trifid at tip. Abd. obovale,
broad, arched, hairless. Ptm. only reaching a very short distance
posteriorly, about to insertion of leg 3, but prolonged anteriorly,
forming a pointed horizontal blade light in color. Ung. trid. L. ca. 800 n*).
In woods. Germany, Sweden.
7. O. globula Nic. 1855 0. g., Nicolet in: Arch. Mus. Paris. i\ 7 p. 439 t. 5 f. 1
| 1884 0. g., Michael, Brit. Orib., v. 1 p. 234 t. 5 f. 6 12 | 1887 Oribates globulus,
Berlese, A. M. S., fasc. 43 nr. 6.
Im. Very black, highly polished, globular. Cephth. short, broad; Lam.
narrow blades on edge; C. lam. rather long, slightly truncated; each bears
a spine; Trlam. a thickened line. Psstg. org. long, filiform. Dors, of
Abd. much arched, no hairs on it; ant. margin of Abd. strongly sinuated;
post, margin truncated. - Nph. very like that of 0. alata. Color varies,
redish pink, straw-color, or olive green; internal organs often shew as dark
patches. Cephth. small, very little of it except Rost. seen. Psstg. org.
longish, recurved, with slightly lanceolate ends. Abd. globular, Dors, of Abd.
polished semi-transparent, without hairs. - - Im. L. ca. 1.05 mm.
On lichens and mosses. Algeria, England, France. Germany, Italy.
8. O. gracilis Michael 1884 0. g., Michael, Brit. Orib., v. 1 p. 225 t. 3 f. 9, 10 |
1895 Oribates g., Berlese, A. M. S., fasc. 74 nr. 6.
Im. Bright brown, highly polished. Cephth. long, slender, very
pointed; Dorso-vertex triangular, narrow anteriorly; Lam. blades on edge,
C. lam. long blunt-pointed, standing upward and forward, and with strong
hair at point; Tectp. 1 produced to sharp point bearing pectinated hair.
Psstg. org. straight, long, moderately clavate. Legs long, slender,
hind Femora flattened, with blades. Ant. margin of Abd. strongly undu-
lated. Ptm. reaching far back. No hairs on Dors, of Abd. Nph. Cephth.
legs and a large plate on Dors, of Abd. dull brown; remainder of Dors, of
Abd. ochre-yellow or greenish. 2 pairs of rostral hairs; Psstg. org. longer
and less clavate than in Im. legs shortish, Tibia of front pair enlarged at
distal end. Central and post, parts of Dors, of Abd. covered by a coarsely-
punctured chitinous plate. The ant. part bears 3 transverse rows of small
chitinous sclerites of various forms; a stout hair on each antero-lateral
angle of Abd., none on Dors, of Abd. - - Im. L. ca. 480 ju.
In moss on trees. England, Italy. .Rare.
9. O. lapidaria H. Luc. **) 1841 Murcia rubra, C. L. Koch, C. M. A., fasc, 31
t. 30 (Nph.!) i 1849 Oribata lapidaria, H.Lucas in: Expl. Algerie, An. artic., p. 318
t. 22 f. 11 | 1855 O.L, Nicolet in: Arch. Mus. Paris, v. 1 p. 437 t. 5 f . 6 | 1884 0.1,
Michael, Brit. Orib., v.l p. 230 t. 5 f . 1 5 | 1883 Oribates humeralis (non Hermann
nee C. L. Koch), Berlese, A. M. S., fasc. 3 t. 4.
Im. Dark rich brown, a large patch on Abd. near ant. margin of
Abd. and some smaller spots lighter; whole creature usually polished.
Genae shew anteriorly as small cusps. Cephth. short wide; Dorso-vertex
wide anteriorly, C. lam. very short; Tectp. 1 produced to a long point.
Psstg. org. short, clavate, Intlam. hair very long. Apodemata not reaching
*) Description supplemented from the present author's specimens.
**) According to the synonyma enumerated above by the author
and in accordance with the ,,Regeln fur die wissensch. Benennuug der Thiere"
the name of this species is: O. rubra (C. L. Koch). The Editors.
See my note p. 7. The Author.
Oribata 15
more than half across sternal plate. Abd. broad much arched, ant. margin
of Abd. strongly sinuated. Post, margin very round; genital and anal covers
far apart, former much the smaller. - - Nph. Orange-red or dark pink,
diamond-shaped. Cephth. not above '/. of entire length, a constriction in
front of Psstg. Psstg. org. globular heads on short peduncles. Abd. flat
and diamond-shaped when young, more convex and rounded lateral. 2 dark
red spots on ant. margin of Abd. with whitish space between; spots joined
by a red line ; a band of about 6 black clavate hairs a little behind red
line ; hinder part of Dors, of Abd. mostly covered by a flask-shaped patch
darker and thicker than rest of Dors, of Abd. and strongly dotted, broader
part forward. Crimson expulsory vesicles at sides of patch. - - Im. L.
ca. 800 n.
In moss and on bark of trees. Algeria, England, France, Germany, Netherlands.
10. O. lucens L.Koch 1879 0. 1, L.Koch in: Svenska Ak. Handl., v. 16 nr. 5
p. 115 t. 4 f. 2 | ?1879 Oppia oblonga, L.Koch, ibid., p. 118 t. 4 f . 6 j ?1879 Claviceps
bimaculatus, L. Koch, ibid., p. 135 t. 7 9. (Nph.)
Im. Black, very like 0. lapidaria but Dors, of Abd. said to be strewn
with short, fine white hairs distant from each other about their own length;
and not quite so wide as 0. lapidaria. L. 600 |u ? 1 mm.
Siberia (Yeniseisk), Novaya Zemlya.
11. O. microptera Can. 1896 0. m., Canestrini in: Bull. Soc. Veneto-Trent.,
v. 6 nr. 2 p. 90 | 1897 0. m., Canestrini in: Atti Soc. Veneto-Trent., ser. 2 /?. 3 p. 149 t. 4.
Im. Dark-brown, legs and Ptm. yellowish. Very like 0. globula, but
Abd. not truncated posteriorly. Psstg. org. more clavate with pointed distal
end. Ptm. small, narrow, confined to ant. part of Abd. Ung. trid. L. ca. 1 mm.
Bolivia.
12. O. mollicoma C.L.Koch 1840 Oribates mollicomus, C.L.Koch, C. M. A.,
fasc. 30 t. 20 | 1884 Oribatam., Michael; Brit. Orib., v. 1 p. 227 t. 4 f. 1 9.
Im. Black, legs and a patch on Dors, of Abd. red-brown. Dull but
not areolated. Rost. blunt; Dorso-vertex little narrowed anteriorly, Lam.
narrow blades on edge, nearly as long as Rost. Trlam. a line, far
forward. C. lam. pointed, turned down, bearing hair. Psstg. org. long with
filiform peduncles and discoidal head. Intlam. hair very long. Legs short,
2 front Femora with large blades. Abd. a long oval; Ptm. running far back;
4 rows of very long flexible hairs on Dors, of Abd. Nph. olive-
brown, polished. Rost. rounded. Cephth. behind Rost. much widened, bearing
2 small ridges. Psstg. dorsal, Psstg. org. long with filiform peduncles and
fusiform head. Legs short; pairs 3 and 4 almost hidden by Abd., which
is short wide and shield-shaped. Im. L. ca. 500 ju.
In moss. England, Germany, Sweden.
13. O. orbicularis C.L.Koch 1835 Oribates o., C.L.Koch, C. M. A., fasc. 3
t. 6 | 1855 Oribata o., Nicolet in: Arch. Mus. Paris, v. 1 p. 435 t. 5 f . 2 | 1877 Oribates o.,
Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 82 | 1883 0. o., Berlese, A. M. S.,
fasc. 9 nr. 1 | 1884 Oribata o., Michael, Brit. Orib., v. 1 p. 236 t. 6 f. 1, 3 9.
Im. Dark red-brown, a vague lighter patch near ant. margin of Abd.
finely punctured. Rost. blunt, genae show as points rather longer than frons.
2 pairs of Rost. hairs. Cephth. long. Lam. large blades nearly on edge, wider
anteriorly than posteriorly. C. lam. short, broad, truncated; Lam. hair in
middle of truncated edge; corner of C. lam. outside hair rounded.
16 Oribata
Trlam. a narrow blade on edge, of even width almost throughout-
Psstg. large, open, projecting; Psstg. org. short, spatulate, slightly recurved;
Tectp. 1 with long projecting point, 2 rounded. Abd. generally nearly
round, but varies, arched, slightly truncated anteriorly. Ant. margin of Abd.
strongly sinuated. Dors, of Abd. hairless. -- Nph. very like that of 0. lapidaria,
except in color. Pinkish-brown or yellow-brown; much dotted and slightly
rough; pyriform or diamond-shaped. Cephth. short, constricted in middle.
Psstg. org. short, heads nearly globular. Intlam. hair, Rost. hairs and some
of the hairs on legs serrated; Dors, of Abd. with large, flask-shaped,
thickened, dotted patch with expulsory vesicles at sides as in 0. lapidaria
Nph. A large, curved, serrated hair on antero-lateral angle ; 2 rows of short
fine hairs on Dors, of Abd. and 2 pairs on post, margin. - - Im. L. ca. 650 |o.
On oak-trees, etc. England, France, Germany, Italy, Sweden.
14. O. pallidula C.L.Koch 1841 Zetes pallidulus, C.L.Koch, C.M.A., fuse. 31
t. 9 j ?1841 Z.fuscomaculatus, C.L.Koch, C. M. A., fasc. 31 1. 11 (Nph.) | 1844 Z.
latipes, C.L.Koch, C. M.A.. fasc. 38 1. 14 j 1886 Oribates I, Berlese, A.M. S.. fasc. 30
nr. 3 j ? 1855 Oribata femoralis, Xicolet in : Arch. Mus. Paris, v. 1 p. 433 t. 4 f. 4.
Im. Red-brown or chestnut-brown. Lam. narrow blades widest poste-
riorly. No C. lam., Trlam. a mere line if visible. Psstg. org. short, clavate.
recurved. Intlam. hair very long, reaching tip of Rost. Abd. narrow, small
posteriorly. Ant. margin of Abd. convex forward in centre, concave laterally.
Ptm. narrow, but long. Dors, of Abd. hairless. Femur of leg 2
greatly enlarged at distal end. Berlese's drawing of Abd. is much
narrower posteriorly than Nicolet's. - - Im. L. 500 620 |u.
In moss. France, Germany, Italy.
15. O. piriformis Nic. 1855 0. p., Nicolet in: Arch. Mus. Paris, v. 7 p. 436 t. 5
f.3 | 1884 O.p., Michael, Brit, Crib., v. 1 p. 238 t. 6 f. 2, 10.
It is often impossible to distinguish the Im. of this species from
0. orbicularis, but the Nph. is quite different. 0. piriformis is generally
narrower than 0. orbicularis and has outer corner of C. lam. pointed instead
of rounded. - - Nph. Rost. and legs dark brown, rather rough ; provertex
light yellow polished, behind this 2 rolls of leathery skin greenish-brown.
Cuticle of Abd. whitish but Dors, of Abd. almost entirely covered by a finely
punctured chitinous plate which is dark olive-green in fully grown Nph.
Rost. pointed. Cephth. behind Rost. in 3 divisions. Intlam. hair and Rost.
hairs not serrated. Abd. veiy square, with rounded corners, not much
arched; Dors, of Abd. hairless. - - Im. L. ca. 650 ILI.
In moss. England, France.
16. O. punctum C. L. Koch, Berl. 1838 Oribatesp., C.L.Koch. C.M.A.. fasc. 30
t. 22 [ 1877 0. p., Canestrini & Fanzago in : Atti 1st. Veneto. ser. 5 v. 4 p. 16 | 1885 0. p.,
Canestrini, Prosp. Acarof., v.l p. 19 j 1886 O.p., Berlese. A. M. S., fasc. 30 nr. 2.
Im. Dark smoky-brown, smooth. Lam. very short blades on edge,
much nearer together anteriorly than posteriorly. Trlam. a blade as wide as
Lam., which distinguishes this species from 0. avenifera. No Trlam. Psstg.
org. directed horizontally forward with short peduncle and fusiform
head. Abd. sub-hemispherical. Ptm. wide, not running far back. Femur
1 with blade. L. ca. 400 ju.
Koch's species would not be distinguishable, Canestrini's contains erroneous
anatomy; the species really is denned by Berlese's figure and description.
In moss. Germany?, Italy, Sweden.
Oribata 17
17. O. rubens C. L. Koch 1841 Oribates r., C.L.Koch. C.M.A., fasc. 31 t, 10
1887 0. longipes, Berlese, A. M. S., fasc. 35 nr. 5.
Im. Chestnut-brown; smooth. Cephth. short. Lam. and Trlam. blades
on edge ; no C. lam. Psstg. org. moderate length, clavate, curved forward.
Lam. and Intlam. hairs slightly pectinated. Abd. globular, hairless.
Ptm. not reaching far back, terminated suddenly posteriorly, slightly undulated
anteriorly. Legs 1 and 2 long, femur 1 passing tip of Host. L. ca. 450 |u.
In moss. Germany, Italy.
18. O. setosa C.L.Koch*) 1835 Murcia trimaculata, C.L.Koch, C. M. A.,
fasc. 3 t. 21 (Nph. !) j 1840 Oribates setosus, C. L. Koch, C. M. A., fasc. 30 t. 19 | 1855
Oribata setosa, Nicolet in: Arch. Mus. Paris, v. 7 p. 436 t. 5 f . 4 j 1884 0. s., Michael,
Brit. Orib., v. 1 p. 243 t, 7 f. 312 : 1889 Oribates setosiis, Berlese, A. M. S., fasc. 43
t.4 1872 Oribata notata, Thorell in: Ofv.Ak.F6rh., v.28 p. H83.
Im. Dark brown, a lighter patch near ant. margin of Abd. Ptm. and
legs lighter. Broadly pyriform, round posteriorly. Lam. large blades on
edge; C. lam. almost horizontal, with points nearly as long as Eost.
Trlam. a blade with deep central indentation, and forming with
Lam. sharp angles which bear strong serrated Lam. hair. Tectp. 1
with long point bearing curved, serrated spine. Psstg. org. longish, rough,
clavate, slightly recurved. Intlam. hair long, rigid, serrated. Abd. slightly
longer than broad. Ant. margin of Abd. forms 3 shallow lobes, Ptm. vague
posteriorly. Dors, of Abd. smooth, dull. 1 or 2 rows of rough thick hairs round
margin ; a variable species. - - Nph. Rost. and legs pink-brown, chitinous,
Abd. bluish-white ; alimentary canal shows dark, often yellow-green ; expulsory
vesicles almost black ; a constriction between Rost. and remainder of Cephth.
2 pairs of rough Rost. hairs. Psstg. org. short, clavate ; Intlam. hair rough
spines. Ant. margin of Abd. slightly concave, several rough spines on
Dors, of Abd. - - Im. L. ca. 540 |u.
In long grass etc. England, France, Franz-Joseph Archipelago, Germany. Italy,
Netherlands, Spitzbergen.
19. O. sph'agni Michael 1880 0. s., Michael in: J. R. micr. Soc., v. 3 p. 179 t.4
f. 6 9 1884 O.s., Michael, Brit. Orib., v.l p. 223 t. 3 f. 1 8.
Im. Yellow-brown, polished. Rost. blunt. Lam. narrow, blades oa
edge ; C. lam. short with curved hair at point. Trlam. a line only. Psstg.
small, hidden by Ptm. Psstg. org. minute, entirely hidden within Psstg.
No Intlam. hair. Legs short, pair 4 not extending beyond Abd. Abd. oval,
ant. margin sinuous. Dors, of Abd. polished and hairless. Ptm. strap-like,
only covering about half the femur of leg 2. - Nph. Greenish-white or
yellow-grey, legs pinkish or yellow. Rost. bluntly-conical with ridges enclosing
spaces, Psstg. near together, Psstg. org. long and filiform. Intlam. hair
upright spines. Abd. shield-shaped, Dors, flat, covered with irregular wavy
ridges bearing scattered dots. Whole edge of Abd., except ant. margin
armed with long spikes. Im. L. ca. 320 |u.
On sphagnum. England.
*) According to the synonyma enumerated above by the author and
in accordance with the ,,Regeln fur die wissensch. Benennung der Thiere"
the name of this species is: O. trimaculata (C.L.Koch). The Editors.
See my note p. 7. The Author.
Das Tierreich. 3. Lief.: A. D. Michael, Oribatidae. 2
18 Oribata
Subdiv. Ib. Sp. 20-39. Without any Trlam.
Sect. Ib 1. Lam. large, horizontal, attached by post, edge only :
the 2 near together, leaving only a narrow cleft between. (Gen. Oribatdla
Banks.)
20. O. americana Haller 1884 0. a., G. Haller in: Arch. Naturg., v. 50 I p. 222
1. 15 f. 4.
Im. Red-brown, polished. Lam. horizontal, very large, almost covering
Cephth. standing free except at hind edge, straight and nearly touching at
inner edges as in 0. tecta, but outer edge strongly and irregularly
toothed, ant. end truncated, concave, and without Lam. h.? Intlarn.
hair extremely long, setiform. Psstg. org. moderate, somewhat clavate,
directed forward and inward. Eost. hairs clavate, hairy. Abd. almost
globular, with 2 longitudinal rows of long, setiform, strongly-curved hairs.
Legs short and thick. L. 390 ju.
America.
21. O. aquatica (Banks) 1895 Oribatella a., Banks in: Tr. Amer. ent. Soc.,
fl.22 p. 9.
Im. Dark reddish, a pale spot on Dors, of Abd. near ant. margin;
body very broad, almost circular, depressed, smooth. Lam. short, each
ending anteriorly in 2 spine-like projections, the lateral sinuate,
the median curved and bearing Lam. hair. Intlam. hair long, curved for-
ward; Psstg. org. moderate length, clavate. Ant. margin of Abd. convex
forward in middle. Ptm. oblong. Dors, of Abd. hairless. L. 550 |a.
On aquatic plants. New York State.
22. O. berlesei Michael*) 1883 Oribates calcaratus (err., non C. L. Koch 1835!),
Berlese, A. M. S., fasc. 9 nr. 2 & Crypt. II, p. 65 | 1885 0. c. (err.), Canestrini, Prosp.
Acarof., v. 1 p."17 || non 1880 Oribata quadricornuta, Michael in : J. R. micr. Soc., v. 3
p. 181 t. 4 f. 15.
Im. Like 0. quadricornuta, but considerably smaller, black instead of
brown. Psstg. org. very long and strongly curved inward, instead of clavate.
Intlam. hair setiform instead of pectinated. Lam. touching anteriorly and
joined in centre by a cross-bar with a central branch running backward;
hairs on Dors, of Abd. much more numerous and placed at edge of Abd.
instead of far in. No. pectinated Eost. hair? and strong spines on Genual
and Tibia instead of pectinated hairs. L. ca. 450 ju.
In moss and dead leaves. Italy.
23. O. lariguida Nic. 1855 0. 1, Nicolet in: Arch. Mus. Paris, v.7 p. 434 t.4 f. 8.
Im. Black, legs and ant. part of Ptm. brown. Like 0. ovalis, but
Dors, of Abd. covered with large, irregular transverse furrows,
or depressions (about 6). Ant. outer angle of Lam. a sharp, drawn out
point; post, margin of Abd. slightly undulated. Is this rare species a
deformity ? L. 420 |u.
France.
*) Berlese identifies the species with O. calcaratus C. L. Koch apparently in
error; consequently this is nameless. I have therefore named the species in honor
of Berlese. A.D.Michael.
Oribata 19
24. O. ovalis C. L. Koch 1835 Oribates ovalis, C. L. Koch, C. M. A., fasc. 3 t. 5
j 1877 0. o., Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 82 | 1855 Oribata
nitens, Nicolet in : Arch. Mus. Paris, v. 7 p. 433 t. 4 f. 8 j 1879 0. n., L. Koch in: Svenska
Ak. Handl., v. 16 p. 115 t. 4 f . 4 | 1885 O.punctata, Nicolet in: Arch. Mus. Paris, v. 7
p. 434 t. 4 f. 7; t. 2 f. 3 ' 1884 0. p., Michael, Brit. Orib., v. 1 p. 253 t. 9 f. 114 1883
Oribates Nicoletii, Berlese, A. M. S., fasc. 3 nr. 3 | ?1895 Oribatella armata, Banks in:
Tr. Amer. ent. Soc., v. 22 p. 9.
Im. Chestnut-brown to black; variable, typically dull and punctured,
var. nitens Nic. polished; Lam. very large, almost hiding Cephth.,
broad, horizontal, reaching tip of Rost., their inner edges almost
touching, truncated anteriorly; but with a prolonged point at outer angle;
Lam. hair from inner angles, short, hooked downward. No Trlam. Psstg.
org. medium length, slightly thickened toward tip ; Abd. arched, hind margin
truncated. Ptm. large, clearly denned posteriorly, prolonged anteriorly,
forming points which almost reach tip of Rost. A row of fine hairs
round Dors, of Abd. and 2 sparse longitudinal rows on same ; a very variable
species. Nph. brown, leathery, deeply wrinkled; Psstg. org. long, rough.
- Im. L. ca. 650 |u (but very variable).
England, France, Germany, Italy, Netherlands, ? New York State America.
25. O. quadricornuta Michael 1880 0. q., Michael in: J. R. micr. Soc., v. 3
p. 181 t. 4 f. 15 ] 1884 0. q., Michael, Brit. Orib., v. 1 p. 247 t. 8 f. 110.
Im. Chestnut-brown, a vague lighter patch near ant. margin of Abd.
legs light. Broadly pyriform; smooth not polished. Rost. rounded, with
flattened margin, Rost. hairs large, curved, strongly pectinated on
outer edges. Lam. very large, their hind margins, and those near
cusps almost meet, arched, but otherwise horizontal. Ant. edge of Lam.
deeply excavated, leaving 2 horns as long as Rost; Lam. hair in
centre of excavation, large, serrated; no Trlam. Intlam. hair long,
serrated; Tectp. very large ant. edges of pair 1 strongly serrated.
Psstg. org. long, clavate, pointed anteriorly rough. Ant. margin of Abd. much
sinuated. Ptm. large. A row of long, flexible hairs on Dors, of Abd. and
some on post, margin. - - Nph. Pale flesh-colour, Rost. and legs darker,
polished, transparent. Psstg. org. like Im. Intlam. hair extremely long,
serrated; a similar but smaller pair on Dorso-vertex. Rost. hairs similar.
Numerous serrated spines on legs. Dors, of Abd. shield-shaped, arched, a
large fold near ant. margin of Abd., a truncated conical projection in centre
of post, margin; numerous serrated spines on Dors, of Abd. - Im.
L. ca. 580 |n.
On dead wood etc. Algeria, England.
26. O. quadridentata (Banks) 1895 Oribatella 4-d., Banks in: Tr. Amer. ent.
Soc., v.22 p. 8.
Im. A species apparently resembling 0. quadricornuta and 0. berlesei,
but Psstg. org. stated to be short (shape not stated). Dors, of Abd. with
about 24 long stiff hairs; Ptm. stated to be ,,large, finely reticulate, ant.
margin finely dentate, terminating below in a small spine". Species im-
perfectly described from one specimen, not figured; but if the description
of the Ptm. be correct this is a good species. The ant. margin of the
Tectp. 1 is dentate in 0. quadricornuta and 0. berlesei, but the author
(Banks) never describes Tectp. L. 600 ju.
New York State.
20 Oribata
27. O. tecta Michael 1884 0. t., Michael, Brit. Orib., v. 1 p. 251 t. 8 f. 11.
Im. Dark-brown, polished, rather long-shaped. Lam. horizontal, very
large, almost covering Cephth. standing free except at hind edge,
their inner edges almost or quite touch, truncated anteriorly,
truncated edge has a shallow notch. Lam. hair near inner angle, rough.
Psstg. org. long, filiform or slightly clavate; Intlam. hair long, rough; legs
short, pair 4 not reaching far behind Abd. Dors, of Abd. hairless.
L. ca. 210 n.
In moss. England.
Sect. Ib 2. Lam. normal or absent.
28. O. agilis Nie. 1855 0. a., Nicolet in : Arch. Mus. Paris, v. 7 p. 432 t. 4 f. 3.
Im. Clear red-brown, polished. Lam. blades narrow posteriorly, widest
anteriorly. Lam. hair long and strongly serrated. Intlam. hair recurved.
No Trlam. Psstg. org. short, recurved, clavate. Abd. hairless. Ptm. trun-
cated anteriorly. L. 430 |u.
In moss. France.
29. O. alata (Herm.) 1804 Notaspis alatus, J. Fr. Hermann, Mem. apt., p. 92
t. 4 f. 6 | 1826 Galumna alata, Heyden in: Isis, p. 613 | 1841 Zetes alatus, C.L.Koch,
C. M. A., fasc. 31 t. 6 \ 1844 Oribata alata, Gervais in: Walckenaer, Ins. Apt., v. 3 p. 258
| 1855 0. a., Nicolet in : Arch. Mus. Paris, v. 1 p. 431 t. 4 f . 1 ] 1877 Oribates alatus,
Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 81 | 1884 Oribata alata (part.),
Michael, Brit. Orib., v. 1 p. 257 1. 10 f. 1 5 | 1886 0. a., A.S.Packard in: Mem. Ac.
Washington, 0.4 p. 42 1896 Oribates alatus, Berlese, A. M. S., fasc. 78 nr. 9 , 1841
Zetes satellitius, C.L.Koch, C. M. A., fasc. 31 1. 13 (Nph.) j ?1895 Oribata emarginata,
Banks in : Tr. Amer. ent. Soc., v. 22 p. 7.
Im. Brown or black ; short pyriform ; very round posteriorly ; polished.
Cephth. broad, bluntly-pointed, arched; no Lam. nor Trlam.; Psstg. org.
short (see 0. dorsalis) recurved, usually lanceolate or clavate but vary. Abd.
almost circular, Ptm. very large, flexible, light-colored, translucent,
they project nearly to tip of Rost. and are rounded anteriorly;
they have a sinuated (emarginate) lateral margin not seen from above ; they
have radiating lines. - - Nph. Broad, transparent, polished, straw-color to
brown; internal organs shew, dark. Rost. blunt, 2 pairs of Rost. hairs.
Psstg. org. dark, recurved, thin, slightly thickened distally. Dors, of Abd.
arched, hairless. - - Im. L. ca. 600 ju.
In moss and lichen. Algeria, America (Kentucky), Belgium, England, France,
Germany, Italy, Netherlands, Switzerland, Sweden.
30. O. avenifera Michael 1884 0. a., Michael, Brit, Orib., v.l p. 264 1. 11
f. 611.
Im. Reddish-brown, not dark; almost globular when Ptm. are pressed
to body, more shield-shaped when expanded; polished (punctured unter high
magnification). Cephth. short, broad; Lam. long, narrow blades on edge;
C. lam. long, blunt, Lam. hair terminal. Psstg. org. oat-shaped, or fusi-
form, sessile, almost horizontal, pointing forward and slightly
inward. Tibia of leg 4 with a projection. Ant. margin of Abd. advancing
greatly in centre, Dors, of Abd. arched, hairless; Ptm. short but usually
extended. L. ca. 320 |u.
In moss. Algeria, England.
Oribata 21
31. O. centro-americana Stoll 1891 O.c.-a., 0. Stoll in: Biol. Centr.-Amer.,
Arach. acar., p. 24 t. 15 f. 1.
Im. Black, polished, without hairs or markings on dors, surface. Very
broad and rounded posteriorly. No division between Cepth. and Abd.
Lam. small narrow. No Trlam. Psstg. org. medium length, slightly clavate,
slightly recurved. No Intlam. hair. L. ca, 900 |u.
Antigua, Guatemala, Brit. Honduras.
32. O. cuspidata Michael 1884 0. c., Michael, Brit. Orib., v. 1 p. 260 1. 10
f. 611.
Im. Light yellow-brown; appears polished but is very finely punctured;
shape broad, rounded posteriorly. Rost. pointed, genae with strong points
.which shew at sides of frons. Lam. very short, doubly curved, concave
on ant. margin; Lam. hair in centre of concavity. No C. lam. nor Intlam.
hair. Tectp. 1 with projecting points. Psstg. org. long, gradually
clavate. Ant. margin of. Abd. sinuous. Dors, of Abd. hairless. - - Nph.
Cannot be distinguished from Nph. of some Notaspididae, variable white or
pinkish, very transparent, internal organs shew, giving color; polished with
transverse markings like seratches; long, narrow. Kost. blunt-pointed, Cephth.
slightly constricted in middle. 2 pairs of Kost. hairs. Psstg. org. long,
slightly clavate. Abd. long, truncated anteriorly; a strong spine on each
antero-lateral angle ; 2 rows of short hairs on Dors, of Abd. and a row round
post: margin. Im. L. ca. 350 )^.
In moss. England.
33. O. dorsalis (C. L. Koch) 1835 Zetes d., C. L. Koch, C. M. A., fasc. 2 1. 14 |
1841 Z. elimatus (laps., corr. elimatus), C.L.Koch, C.M.A., fasc. 31 t. 5 | 1842 Z.elimatus,
C. L. Koch, Ubers. Arach., v. 3 p. 100 I 1877 Oribates elimatus, Canestrini & Fanzago
in : Atti 1st. Veneto, ser. 5 v. 4 p. 81 | 1886 0. elimatus, Berlese, A. M. S., fasc. 30 nr. 1 |
1884 Oribata alata (part.), Michael, Brit, Orib., v. 1 p. 257 1. 10 f. 1 5.
Im. Very like 0. alata, of which it may be a variety, it has long
Psstg. org. instead of the short ones of 0. alata. L. ca. 700 JLI.
Brazil, England, Germany, Italy, Netherlands, Paraguay, Sweden.
34. O. halleri Michael*) 1884 0. simplex (nonGervais!849!), G. Haller in: Arch.
Naturg., v. 501 p. 221.
Im. Light yellow-red. Lam. widest in middle, diminishing anteriorly
and posteriorly. Psstg. org. with short peduncle and suddenly clavate head.
Abd. almost globular, 2 longitudinal rows and a border of hook-shaped
granular hairs on Dors. ofAbd. Ptm. truncated anteriorly, scarcely passing
ant. margin of Abd., much folded under body. Legs short, thin, hetd.
L. 440 |u.
America. _
35. O. intermedia Michael**) 1855 0. ovalis (err., non C. L. Koch 1835 !), Nicolet
in: Arch. Mus. Paris, v. 7 p. 438 t. 4 f. 5 || non 1883 Chibates Nicoletii, Berlese, A. M. S.
fasc. 3 nr. 3.
*) Haller's name of simplex was preoccupied in the genus Oribata by Gervais'
species (in: Gay, Hist. Chile), which Haller apparently overlooked; I have therefore
renamed the species in honor of Haller. A.D.Michael.
**) Nicolets name 0. ovalis being preoccupied in this genus; and Berlese's name
O. Nicoletii which was intended to cure the difficulty ; being, in my opinion, applied
to a different species, I have renamed this species. A.D.Michael.
22 Oribata
Im. Light red-brown, polished, like 0. ovalis but Lam. different, vizt.
large thick blades not transparent, set at angle of about 45, con-
cave inward slightly wider and much further apart posteriorly than ante-
riorly, truncated anteriorly. Lam. hair from outer (upper) angle.
Abd. said to be hairless. L. 520 ja.
France.
36. O. lucasi Nic. 1855 0. Lucasii, Nicolet in : Arch. Mus. Paris, v. 7 p. 432
t.4 f. 2 | 1884 O.L., Michael, Brit. Orib., v. 1 p. 262 t. 11 f. 1 5.
Im. Light reddish-brown. Legs and Ptm. lighter, yellowish; polished.
Cephth. long; Kost. sharp; Lam. narrow blades on edge; no C. lam. nor
Trlam. Psstg. org. with fusiform head on long slender peduncles,
recurved. Kost. hairs long, curved; very large blades to femora 2. Abd.
oval, rather long; Dors, of Abd. arched but flattened centrally, hairless.
Ant. margin of Abd. slightly sinuous ; Ptm. small, transparent. A few short
hairs round post, margin. Ventr. plate much embraced by dorsal at sides.
Genital covers almost heart-shaped, far forward ; anal covers large, square.
- Nph. Very like a Notaspis Nph. White, transparent; internal organs
shew. Cephth. long, constricted in middle. 2 pairs of Eost. hairs. Psstg.
org. like Im., but directed forward. Dors, of Abd. hairless, without markings.
A fine hair on each antero-lateral angle and a few short ones on post.
margin. - - Im. L. ca. 600 ju.
Algeria, England, France.
37. O. mucronata G. & R Can. 1882 0. m., G. & E. Canestrini in : Atti 1st.
Veneto, ser. 5 v. 8 p. 916 t. 9 f. 2 | 1892 Oribates mucronatus, Berlese. A. M. S.,
fasc. 62 nr. 8.
Im. Dark-brown, Ptm. yellowish. Like 0. dorsalis; but. post, margin
of Abd. forms 2 chitinous points near median line, and between them a
hyaline, median, pointed projection divided longitudinally. L. 650 800 ju.
C. L. Koch, C. M. A., fasc. 2 1. 14 (1835). states that what is evidently this species
is one sex of his Zetes dorsalis and that he found them in Copo, but as no such sexual
variation is known in the Oribatidae and no one else has ever seen any species in
Copo and both sexes of Oribata (Zetes) dorsalis are known, this statement is probably
an error.
Germany, Italy.
38. O. peloptoides Berl. 1888 Oribates p., Berlese in: Bui. Soc. ent. Ital., v. 20
p. 215 1. 13 f. 3.
Im. Chestnut-brown. Lam. long but very narrow. No Trlam. Psstg.
org. (as in 0. lucasi) with long peduncle arid fusiform reflexed head. Dors,
of Abd. rough, punctured or areolated. Ptm. short, truncated anteriorly and
posteriorly. 4 rows of stout hairs on Dors, of Abd. and a row round edge.
L. ca. 450 |^.
Under stones. Brazil, Paraguay.
39. O. rugifrons Stoll 1891 0. r., 0. Stoll in: Biol. Centr.-Amer., Arach. acar.,
p. 25 1. 15 f. 2.
Im. Black, smooth, not polished, like 0. dorsalis, but dors, surface
marked with extremely fine longitudinal wrinkles. L. ca. 850 |u.
Brit. Honduras, Guatemala.
Oribata 23
Div. II. Sp. 40 45. Ung. mond.
40. O. dentata Berl. 1883 Oribates dentatus, Berlese, A. M. S., fasc. 9 nr. 3.
Im. Chestnut-brown, polished. Lam. very narrow blades on edge.
Rost. hairs and Lam. hairs long. Intlam. hair and Psstg. org. very long,
would reach far beyond tip of Rost. the former thin, setiform, flexible, the
latter recurved with thiupeduncle andfusiformhead. Ant. margin of Abd.
slightly convex ; 2 long hairs on post, margin, none on dors, surface of Abd.
Femur 2 with large blade ending anteriorly in large curved tooth. Ung.
figured trid. on fig. of whole creature, mond. on fig. of separate legs, nothing
said in letterpress; is mond. (Berlese, A. M.S., Crypt. II p. 65). L. 600 |u.
Found in hot-house of Bot. garden in Padua. Native country?
41. O. fusigera Michael 1884 O./ 1 ., Michael, Brit.Orib., v.l p. 268 t.12 f.611.
Im. Black, front of Abd. and legs lighter. Rost. short. Lam. nearly
on edge, near together, almost hidden by Psstg. org. No Trlam. ; Psstg.
org. long fusiform, terminated distally by a fine hair, they point for-
ward and inward. Ant. margin of Abd. sinuous, Dors, polished, hairless. -
Nph. Pinkish, coarsely punctured, long pyriform. Rost. long, narrow; Psstg.
org. with fusiform head on slender peduncles. Hind legs much shorter than
Abd. Dors, of Abd. shield-shaped, pointed posteriorly, not much arched,
hairless. - - Im. L. ca. 300 |u.
In moss. England.
42. O. mirabilis (Banks) 1895 Oribatodes m., Banks in : Tr. Amer. ent. Soc.,
v. 22 p. 10.
Im. Dark red-brown; short, broad, convex. C. lam. strong. Intlam.
hair very long, directed forward. Psstg. org. long, clavate. Abd. globose,
granular, with an irregular network of ridges. About 16 long stiff
serrated hairs in 2 longitudinal rows on Dors, of Abd. 4 clavate on post,
margin. Ptm. granulate. L. 500 n.
Under bark. New York State.
43. O. monodactyla Haller 1884 0. m., G. Haller in: Arch. Naturg., u 501
p. 221 1. 15 f. 3.
Im. Yellow- brown, short pyriform, Cephth. pointed. Lam. long, wide
posteriorly deminishing to nothing anteriorly. No Lam. hair nor C. lam.
Psstg. org. very long, slightly and gradually clavate, directed backward
and outward, distal half pectinated along ant. edge. Abd. long, oval,
slightly arched. Without hairs on Dors, of Abd. L. ca. 300 ja.
Said by Berlese (A. M. S., Crypt. II p. 67) apparently in error to be his Oribates
dentatus.
America.
44. O. parmeliae Michael 1884 O.p., Michael, Brit.Orib., v. 1 p. 265 t. 12
f. 15.
Im. Dark brown, smooth but not polished, punctured. Cephth. short,
Rost. blunt; Lam. very long blades nearly on edge; C. lam. long,
bearing strong curved hair; Trlam. a thickened line; Psstg. org. short,
pyriform recurved. Genuals of 2 first pairs of legs enlarged distally;
Ant. margin of Abd. veiy curved. A row of long flexible hairs round post,
margin and 2 rows on Dors, of Abd. - - Nph. Orangeyellow. Cephth.
24 Oribata
short. Psstg. protuberant; Psstg. org. short, strongly clavate. Rost. hairs
long, 2 pairs of hairs between Rost. hair and Intlam. hair. Legs short, pair
4 not seen from above. Abd. with straight sides; post, margin rounded.
Dors, of Abd. arched, divided in middle by a strong transverse indented
line. Ant. margin of Abd. curved, with 2 or 3 folds near it; 2 long flexible
hairs on each autero-lateral angle and about 6 round post, margin; some
smaller hairs on Dors, of Abd. Im. L. ca. 420 |u.
On lichens (Parmelia). England.
45. O. rileyi Haller 1884 0. R., G. Haller in : Arch. Naturg., v. 50 I p. 223.
Im. Dark brown, very pyriform, compressed anteriorly rounded poste-
riorly, granular, epiostracum loose and easily detachable. Lam. small. C. lam.
large, broad, truncated, almost horizontal, Trlam. a blade on edge.
Lam. hail' and Intlam. hair setiform, small. Psstg. org. rather short, clavate.
Abd. pyriform with crescentic indentation in middle of hind margin;
a small rounded projection lower in level seen in centre of crescent. Ptm.
reaching very far back, the 2 (one each side) joined anteriorly by a
long chitinous bridge. 2 rows of short serrated hairs on Dors, of Abd.
L. ca. 500 ju.
Described from a single specimen.
America.
O. afflnis Banks 1895 0. a., Banks in: Tr. Amer. ent. Soc., v.22 p. 6.
im. Resembles 0. pratensis Banks, but Lam. longer, no Trlam. Ptm. much
smaller (half the size); Psstg. org. longer. Dors, of Abd. hairless. Ung. trid. L. 800 |u.
America (Washington).
O. arborea Banks 1895 0. a., Banks in: Tr. Amer. ent. Soc., v. 22 p. 7.
Im. Yellow-brown, often with a white spot, on Dors, of Abd. near ant. margin,
moderately convex, highest posteriorly. Psstg. org. very short, clavate; Abd. smooth,
Dors, hairless; Ptm. as long in proximo-distal as in antero-posterior measurement,
tapering to a rounded tip. Ventr. surface smooth; post. Femur broad. Ung. trid.
L. 600 IUL
On cedar and peach trees. New York State.
O. calcarata C. L. Koch 1835 Oribates c., C. L. Koch, C. M. A., fasc. 2 t, 13 1|
non 1883 0. calcaratus, Berlese, A. M. S., fasc. 9 nr. 2 & Crypt. II p. 65 (v. 0. berlesei) \
non 1885 0. c., Canestrini, Prosp. Acarof., v. 1 p. 17.
Im. Small, very black, polished. Cephth. with numerous projections. Lam.
broad, white, far apart. Psstg. org. long clavate, reaching tip of Rost. Intlam. hair
long, thick, white. A pair of thick spatulate hairs or spines at distal end of Genual 1. 2,
and 1 at end of Tibiae 3, 4. Abd. globular, truncated anteriorly, 4 minute hairs on
post, margin, no others on Abd.
Species probably good but imperfectly described and figured.
In woods. Germany, Sweden.
O. depressa Banks 1895 0. d., Banks in: Tr. Amer. ent. Soc., v.22 p. 6.
Im. Red-brown. Psstg. org. moderate length, capitate. Abd. smooth, depressed
plainly longer than broad. Ptm. large, long, projecting in front of Abd., not sinuated
at distal margin. Dors, of Abd. hairless. Ung. trid. L. 450 \n.
Xew York State.
Oribata 25
O. flammula C. L. Koch 1840 Oribatesf., C. L. Koch, C. M. A., fasc. 30 1. 16 |
P1859 0. quadricuspis, Grube in: Arch. Dorpat. Ges., ser. 2 v.l p. 476.
Im. Like 0. calcarata but half the size, 2 longitudinal rows of hairs on Dors,
of Abd., none on post, margin. A large, vague, light patch on Dors, of Abd. near
ant. margin.
In wet meadows. Germany.
O. hirsuta Banks 1895 0. h., Banks in : Tr. Amer. ent. Soc., v. 22 p. 7.
Im. Pale brownish-yellow. Psstg. org. short, capitate. Abd. not high, a little
longer than broad, smooth. Ant. margin of Abd. with 2 incisions. Dors, of Abd.
bearing about 30 short hairs, those post, longest. Ptm. moderate, rounded, distal
margin incurved, ventr. surface granular. Ung. trid. L. 400 |u.
Under dead leaves. New York State.
O. homodactyla Karp. 1893 0. h., Karpelles in: Math, naturw. Ber. Ungarn,
v. 11 p. 82.
Im. Ung. said to be homd., other-wise species insufficiently described. Abd.
smooth, without hairs. Intlam. hair ,,chain-shaped". Leg-hairs feathered. L. ?
Hungary.
O. latirostris (C. L. Koch) 1844 Zetes I, C. L. Koch, C. M. A., fasc. 38 1. 13.
Im. Small, black, polished. No Lam., Rost. hairs, nor Intlam. hair. Psstg. org.
moderate length, clavate. Abd. sub-elliptical, broad, hairless. Ptm. short but wide,
ending abruptly posteriorly, slightly prolonged anteriorly.
In wet meadows. Germany.
O. magna Banks 1895 0. m., Banks in : Tr. Amer. ent. Soc., v. 22 p. 6.
Im. Apparently very like 0. globula, but Psstg. org. said to be ,,moderate,
clavate", and said to be no Intlam. hair. L. 1 mm.
New York State.
O. minuta (Banks) 1896 Oribatella m., Banks in: Tr. Amer: ent. Soc.. $.23 p. 76.
Im. Pale reddish brown; legs paler; a pale spot near ant. margin of Abd.
Body globose. Lam. each with 2 spine-like processes, as in 0. quadricornuta. Lam.
hair between the 2 spines and as thick as. the inner one. Rost. hairs pectinate on
their outer sides. Psstg. org. moderate-length, clavate. Abd. finely punctured. Ptm.
large, somewhat pointed and incurved at tip; about 20 fine curved hairs on Dors,
of Abd. L. 300 |n.
One specimen under hemlock bark. New York State.
O. moesta Banks 1895 0. m., Banks in: Tr. Amer. ent. Soc., v. 22 p. 6.
Im. Red-brown, no Trlam. Intlam. hair long. Abd. depressed, polished; Ptm.
small, truncated anteriorly. Psstg. org. moderate length, capitate. Dors, of Abd. hairless.
Ung. trid. L. 550 |u.
New York State.
O. palustris Banks 1895 0. p., Banks in: J. N. York ent. Soc., v. 3 p. 128.
Im. Red-brown, legs yellowish, a whitish spot on Abd. near ant. margin. Lam.
short. No Intlam hair. Psstg. org. moderate length, clavate, not as long as depth
of Ptm. which are much deeper than long at base. Body short, high, broad, globose;
Ptm. prominent. Ventr. surface granular. Legs short. Tibia 1 barely longer than
Genual Post. Femur very broad, margined below. Very like O. sphagni, but probably
a distinct species. L. 420 |i.
In a Sphagnum swamp. New York State.
26 Oribata, Serrariinae, Serrarius
O. pinnata Gerv. 1849 0. p., Gervais in : Gay, Hist. Chile, v. 4 p. 48.
Im. Body oval. Description insufficient for identification.
Chili.
O. pratensis Banks 1895 O.jp., Banks in: Tr. Amer. ent. Soc., v. 22 p. 6.
Im. Yellowish or reddish. Lam. short. Trlam. a ridge. Intlam. hair long, Psstg.
org. short capitate. Dors, of Abd. arched hairless, 3 small smooth spots each side.
Ptm. small, long, triangular; a short clavate figure in front of anal opening. Ung.
trid. L. 700 ja. Possibly 0. lapidaria.
New York State.
O. reticulata L. Koch 1879 0. r., L. Koch in: Svenska Ak. Handl. v. 16 p. 115.
Im. Imperfectly described from a single specimen and not figured, probably
very like 0. orbicularis, but long hairs on Dors, of Abd. L. 1 mm.
Siberia (Yeniseisk).
O. robusta Banks 1895 0. r., Banks in: Tr. Amer. ent. Soc., v. 22 p. 7.
Im. Black, yellowish near ant. margin of Abd. apparently very like 0. dorsalis,
distal margin of Ptm. said not to be sinuated (emarginate). L. 750 |u.
Chicago, Illinois, Texas.
O. signata (Banks) 1895 Oribatella s., Banks in: Tr. Amer. ent. Soc., v.22 p. 9.
Im. Dark reddish brown; a pale patch on Dors, of Abd. near ant. margin.
Abd. convex, smooth. Lam. long, reaching tip of Rost. truncate anteriorly. C. lam.
long, said to be a stiff bristle at each corner (2 all 4 or 2 only). Intlam. hair long,
curved, directed forward; Psstg. org. long, clavate. Abd. longer than broad. Ptm.
moderately large, triangular, projecting somewhat anteriorly. Apparently some-
what allied to 0. tecta. L. 450 |u.
New York State.
O. simplex Gerv. 1849 0. s., Gervais in : Gay, Hist. Chile, v. 4 p. 49.
Body oval. Ptm. scarcely visible. Description insufficient for identification.
Chili.
O. sp., Banks 1895 Oribatella setosa (non Oribates setosus C. L. Koch 1840!),
Banks in : J. N. York ent. Soc., v. 3 p. 128.
Im. Red-brown; a pale spot on Dors, of Abd. near ant. margin. Body longer
than broad. Lam. short; C. lam. long, spine-like. Intlam. hair erect, nearly as long
as leg 1. Psstg. org. nearly 2 / s the length of Intlam. hair, slightly clavate. Abd. smooth,
evenly convex. Ptm. large, somewhat triangular, tip rounded. Post, margin oblique.
Probably has a Trlam. Ung. trid. L. 480 |u.
In Sphagnum-swamp. New York State.
B. Subfam. Serrariinae
Without Ptm. Cephth. anchylosed to Abd. Ventr. plate anchylosed
to dors, plate. Mdb. rod-like serrated at distal end, not eh el ate.
1 genus with 2 species.
1. Gen. Serrarius Michael
1883 Serrarius, Michael in: J. R. micr. Soc., ser. 2 v, 3 p. 25 j 1884 S., Michael,
Brit. Orib., v. 1 p. 270 j 1896 S., Berlese, A. M. S., Crypt, II p. 56 1885 Neozetes, Ber-
lese, A. M. S., fasc. 20 nr. 6.
Serrarius. Zetorchestinae, Zetorchestes
27
Characters (at present) those of the subfamily. Ung. of
known species trid.
2 species.
Key to the species.
Abd. elliptical 1. S. fusifer . . . p. 27
Abd. globular 2. S. microcephalus p. 27
1. S. fusifer (C. L. Koch) 1841 Oribates f., C. L. Koch. C. M. A.,
fasc. 31 t. 3 ; 1885 Neozetes f., Berlese, A. M. S., fasc. 20 nr. 6 1896 Ser-
rarius f., Berlese, A. M. S., Crypt. II p. 56 || non 1855 Leiosoma micro cephala,
Nicolet in : Arch. Mus. Paris, v. 1 p. 443 ! non 1883 Serrarius microcephalus,
Michael in: J. R. micr. Soc., ser. 2 v. 3 p. 25 j non 1841 Oribates globulus,
C. L. Koch, C. M. A., fasc. 31 t. 4.
Im. Yellow-brown, smooth; Lam. narrow blades on edge.
C. lam. minute, scarcely recognizable. Psstg. org. very long,
with thin peduncle and fusiform head. Intlam. hair very long.
Abd. elliptical, slightly truncated anteriorly, 2 short hairs on
post, margin. Dors, of Abd. hairless. L. ca. 470 |u.
In moss. Algeria, Germany, Italy.
2. S. microcephalus (Nic.) 1855 Leiosoma m., Mcolet hi: Arch. Mus. Paris,
v. 7 p. 443 t. 6 f. 4 ; 1883 Serrarius m., Michael in: J. E. micr. Soc., ser. 2 v.3 p. 25 |
1884 S. m., Michael, Brit. Orib., B.I p. 272 t. 14 f. 7 13 1885 Neozetes bicornis,
Berlese, A. M. S., fasc. 20 nr. 7 ; 1896 Serrarius b., Berlese, A.M. S., Crypt. II p. 56 || non
1841 Oribates fusifer, C.L.Koch, C. M. A., fasc. 31.
Im. Red -brown, legs lighter; polished; almost globular. Rost.
slightly trilobed, 2 longitudinal ridges on its dors, surface; Cephth. short;
Lam. blades nearly on edge, widest anteriorly; C. lam rather long, slightly
truncated; Lam. hair terminal. Trlam. a thickened bar. Psstg. org. very
long with slender peduncles and small, fusiform head terminating
in a curved point; Tectp. 1 blade-like, large, 2 almost obsolete. Dors, of
Abd. without hairs or markings. 3 pairs of short hairs round post, margin.
Genital covers broader than anal, placed far forward. L. ca. 450 u.
England. France, Germany.
Fig. 7.
S. micro-
cephalus.
Mdb. (*/,).
C. Subfam. Zetorchestinae
Without Ptm. Cephth. anchylosed to Abd. Ventr. plate anchylosed
to dors. Mdb. chelate. Legs arranged first 3 pairs near together, pair
4 distant, much larger than the others. Coxae and Femora greatly enlarged;
formed for jumping.
1 genus with 1 species.
1. Gen. Zetorchestes Berl.
1885 Carabodes (part.), Canestrini, Prosp. Acarof., v. 1 p. 44 j 1885 Leptorchistis
(corr. Leptorchestes, non Leptorchestes Thorell 1869!), Canestrini & Berlese in: Atti Soc.
Veneto-Trent., t>.9 p. 207 j 1888 Zetorchestes, Berlese, A. M. S.. fasc. 63 nr. 2.
Rost. hairs dorsal, mounted on chitinous apophyses. Lam. present.
Second joint of palpus much the longest. Genital and anal covers close
together at extreme rear of Abd. Ung. mond.
1 species.
28
Zetorchestes, Notaspidinae. Scutovertex
1. Z. micronychus (Berl., Can.) 1883 Carabodes m.,
Berlese, A. M. S., fasc. 4*) j 1885 C. m., Canestrini, Prosj).
Acarof., v. 1 p. 44 t. 2 bis ] 1885 Leptorchistis m., Cane-
strini & Berlese in: Atti Soc.Veneto-Trent., v.9 p. 207 t.7
1888 Zetorchestes m., Berlese, A. M. S., fasc. 49 nr. 7.
Im. Blackish brown, polished; Lam. blades
on edge broadest and sharply truncated anteriorly,
no C. lam. Host hairs pectinated, borne on long
chitinous apophyses, set further back than usual.
Psstg. org. with longish slender peduncles and fusi-
form hairy head. Intlam. hair at extreme base
of Cephth. Abd. sub -globose, without hairs or
markings. Tarsus 4 with 2 and Tibia 4 with 1 short
lanceolate spine in upper median line. Ung.
minute. - Nph. with centre of Dors, of Abd.
divided into 6 broad bands of which the 3 broadest
are reticulated, the intermediate ones simply dotted.
Lat. and post, parts of Abd. irregularly wrinkled.
p t j^ j N R t h h
& &
L. 450 fl.
Fig. 8.
Zetorchestes micronychus ( 75 /i]
Ventr. surface. The Acetabulum
of leg 4 and the muscles moving the 1m.
Coxa are shown on one side.
Algeria, Italy, Switzerland.
D. Subfam. Notaspidinae
1885 Subfam. Nothridae (part.), Berlese in: Bull. Soc. ent. Ital., .17 p. 126 j 1891
N., Canestrini in: Atti 1st. Veneto, ser. 7 v.2 p. 579 ' 1892 Notlirinae (part.), Trouessart
in: Rev. Sci. nat. Quest, v. 2 p. 38* 1896 Nothridae (part.) -f- Tegeocranidae -\- Leioso-
midae (part.) -\-Damaeidae (part.), Berlese, A. M. S., Crypt. II p. 36, 41.
Without Ptm. Cepth. anchylosed to Abd. ventr. plate anchylosed to
dors. Mdb. chelate; legs 1 2 near together, 3 and 4 near together, craw-
ling organs. With lamellae.
6 genera, 62 good and 8 doubtful species, 2 varieties.
Key to the genera.
J Body rough and sculptured (Tegeocranidae Berl.) 2.
\ Body smooth 5.
{Cephth. and Abd. joined by a central projection of
the former, or otherwise 1. Gen. Scutovertex
Cephth. and Abd. separated by an unbroken line 3.
rt J Ung. trid., Femora without thin peduncles . . 2. Gen. Cepheus . .
\ Ung. mond. ; Femora with thin peduncles 4.
J Lam. very large, light colored blades 3. Gen. Tegeocranus
\ Lam. dark, low, solid ridges 4. Gen. Carabodes .
. J Last 3 pairs of legs inserted under the body . . 5. Gen. Liacarus . .
p. 28
p. 31
\ Last 3 pairs of legs inserted at the edge of the body 6. Gen. Notaspis
p. 33
p. 36
p. 40
p. 43
1. Gen. Scutovertex Michael
1879 Scutovertex, Michael in: J. R. micr. Soc., ^.2 p. 241
1884 S., Michael,
Brit. Orib.. v. 1 p. 296 1885 8.. Canestrini, Prosp. Acarof., v. 1 p. 7 j ?1895 S., Banks
in : Tr. Amer. ent. Soc., v. 22 p. 11 1896 S. -f- Ameronothrus, Berlese, A. M. S., Crypt. II
p. 32, 46.
A short note, printed inside wrapper of fasc.
A. D. Michael.
Scutovertex 29
Body rough and sculptured, dors, surface of Abd. not wholly divided
from Cephth., but joined by a central projection or otherwise.
6 good and 1 doubtful species.
Key to the good species.
\ Terrestrial 2.
\ Aquatic 4.
f Lam. blade-like; C. lam. large, rounded 3.
\ Lam. a mere thickened bar, no C. lam. L. 540 |u 5. S. maculatus . p. 30
I Ung. trid. L. 600 n 6. S. sculptus . . p. 30
\ Ung. mond. L. 500 |u 2. S. caelatus . . p. 29
i Psstg. org. with short peduncles and small globular
4 -j head; Ung. trid. 5.
I No Psstg. org. visible; Ung. mond. L. 620 |n . . 1. S. bilineatus . . p. 29
{Markings on Dors, of Abd. ridges usually more or
less zig-zag; 6 hairs on post, margin of Abd. but
no indentations. L. 680 in 3. S. corrugatus . p. 29
Markings on Dors, of Abd. projecting points, coalescing
near ant. margin of Abd.; 2 indentations in post,
margin of Abd. a spine in each. L. 700 |u . . 4. S. harioti ... p. 30
1. S. bilineatus Michael 1888 S. b., Michael, Brit. Orib., v. 2 p. 571 t. 54
f. 8-16 1 1896 Ameronothrus b., Berlese, A. M. S., Crypt. II p. 32.
Im. Dark brown ; dull, rough, rather like S. maculatus. Host, blunt ;
Maxillae with a single median indentation and a small black denticle in
the depression; Lam. thick, ill-defined ridges, usually with a similar Trlam.
Surface of Cephth. covered with raised dots. No Psstg. nor Psstg. org.
Abd. elliptical, with 2 large longitudinal, broken, irregular ridges on Dors,
of Abd. sometimes absent, 2 or 3 very irregular transverse ridges anteriorly
and posteriorly and raised dots like those on Cephth., but a little larger.
Ung. mond. - - Nph. very like that of S. sculptus but no Psstg. ' org. -
Im. L. ca. 620 |u.
Aquatic, on fresh-water algae. England.
2. S. caelatus Berl. 1895 B.C., Berlese, A. M. S., fasc. 74 nr. 1.
Im. Lightish smoky brown; dull, rough. Lam. very slightly raised;
Trlam. mere line; C. lam. very long and large, widest at distal ends
which are rounded. Psstg. org with thin peduncle and strongly clavate
setose head. Kost. dotted, vertex with irregular vermiform striae; Abd.
flatish, reticulated, elliptical, very slightly truncated anteriorly. 3 very small
projections from post, margin, the 2 lat. bearing minute hairs, 2 pairs
similar on lat. margin. Legs rough, apophysis on tibia 1 very long. Ung.
mond. L. ca. 500 |n.
In moss. Northern Italy.
3. S. corrugatus Michael 1888 S. c., Michael, Brit, Orib., v. 2 p. 567 t. 54
f. 17 j 1896 Ameronothrus c., Berlese, A. M. S., Crypt. II p. 32.
Im. Dark brown; dull. Cephth. short. Rost. blunt, slightly trifid.
Lam. straight ridges. Psstg. org. small with short peduncle and head which
looks globular, but is cup-shaped if magnified sufficiently. Dors, surface of
Cephth. with several irregular ridges. Abd. rounded posteriorly. Dors, of
Abd. slightly arched with numerous irregular ridges, mostly longi-
tudinal with short tranverse branches, rarely straight; 4 longitudinal rows
30 Scutovertex
of fine small hairs on Dors, of Abd. springing from minute papillae. Dors,
plate sufflexed. Ung. trid. - - Nph. Very like that of S. sculptus, but
Psstg. org. shorter. - - Im. L. ca. 680 ju.
Aquatic, on fresh-water algae. England.
4. S. harioti Michael 1891 S. h., Michael in: Miss. Cap Horn, v. 8 p. L59.
Im. Dark brown; cuticle hard and rough, Cephth. Dors, of Abd..
femora and coxae of legs 3, 4 thickly strewn with projecting points which
are confluent on Cephth. and ant. part, of Abd., larger, more dispersed, and
irregular on remainder of Abd., they are part of the epiostracum and rub
off with it. Rost. rounded, Lam. scarcely visible. Cephth. without other
markings. Psstg. org. with short peduncle and small globular head.
Intlam. hair very small. Ung. large trid., very hetd. Abd. without mar-
kings other than the points above mentioned; central part of ant. margin
of Abd. projects over Cephth. and coalesces with it. On each side of post,
margin of Abd. is an indentation with a spine inserted; 2 longi-
tudinal rows of about 3 extremely small hairs on Dors, of Abd. - Nph.
Very like that of S. bilineatus, but Intlam. hair very long and thick, flagelli-
form. - - Im. L. ca. 700 JLI.
On marine algae. Terra del Fuego.
5. S. maculatus Michael 1882 S.m., Michael in: J. R. raicr. Soc., ser. 2 v. 2
p. 13 1. 1 f. 15 j 1884 S.m., Michael, Brit. Orib., . 1 p. 302 1. 18 f. 9 16 : 1896
Ameronothrus m., Berlese, A. M. S., Crypt. II p. 32.
Im. Dark brown, almost black. Elliptical, rough, covered with raised
dots. Lam. and Trlam. thickened bars. 2 ridges from the 2 Lam. join
forming a V the point whereof lies in a semicircular indentation of ant.
margin of Abd. Psstg. org. very short with broadly pyriform head.
First 3 joints legs covered with raised dots. Dors, of Abd. with arched
centre and broad, flat, peripheral band; all covered with raised dots of even
size arranged irregularly on arched part, transversely on ant. margin of Abd.,
radiatingly on remainder of flattened band; about 10 short knob-headed hairs
on hind margin. Ung. trid. - - Nph. brown, broad, flat ; covered with ridges
and wrinkles of the cuticle which are arranged in a definite pattern on Cephth.
and ant. margin of Abd., irregularly on remainder of Dors, of Abd. 8 spatulate
hairs on post, margin, the 4 central of these the longest. Im. L. ca. 540 u.
On lichen near sea coast. England.
6. S. sculptus Michael 1879 S. s., Michael in: J. R. micr. Soc., v. 2 p. 242 j
1884 S.s., Michael, Brit. Orib., v. 1 p. 299 1. 18 f. 1 8 || non 1887 Eremaeus ovalis,
Berlese, A. M. S., fasc. 35 nr. 7.
Im. Dark brown or black. Rost. blunt. Cephth. small, narrow, dotted.
Lam. reticulated blades on edge; C. lam. long, blunt, curved; Trlam.
a bar. Psstg. org. long, spatulate, rough; legs with femora enlarged, hind
ones with blades. Tactile hair of leg 1 set on apophyses. Abd. oval,
produced to a long thin point which is anchylosed to Cephth. Edge
of Dors, of Abd. rough, post, margin with central lobe and indented on each
side of it. A bag-shaped space of clear smooth chitin bordered by a ridge
adjoins the centre of ant. margin of Abd.; a triangular dotted space bordered
by a ridge lies on each side of the clear space. Central part of Dors, of
Abd. irregularly and deeply pitted, lat. parts with small raised dots. 4 small,
spatulate hairs on post, margin; a few similar on Dors, of Abd. Ung. trid.
Scutovertex, Cepheus
31
- Nph. Light brown; very like that of Oribata ovalis. 2 square spaces,
one in front of the other, in central line of Cephth., each divided by a median
ridge. Psstg. dorsal near together. Psstg. org. long, lanceolate; Dors, of Abd.
flat, deeply and irregularly wrinkled. - - Im. L. ca. 600 ju.
In moss. Algeria, England.
S. sp., (Berl.) 1887 Eremaeus ovalis (err., non C. L. Koch 1841 !), Berlese, A.
M.S., fasc. 35 nr. 7 j 1896 Scutovertex o., Berlese, A. M. S., Crypt. II p. 46 |j non 1879
S. sculptus, Michael in : J. R. micr. Soc., v. 2 p. 242.
Im. Blackish brown; dull, rough. Lam. straight, narrow blades on edge. Trlam.
a low ridge; C. lam. very short, hardly existing. Psstg. org. with long thin peduncle
and fusiform, spinous head. Abd. prolongued anteriorly, rounded posteriorly. Dors,
of Abd. with clear polished, trapeze-shaped space near ant. margin, central parts
irregularly rough; several diagonal ridges on hind part, the 2 inner forming a. V. ;
4 small clavate. spinous hairs on hind margin. L. 500 (a.
In moss. Italy.
Var. (1887 Eremaeus ovalis var. siculus, Berlese, A. M. S., fasc. 35 nr. 8.) A
central ridge from ant. margin of Abd. along Vertex. 2 short longitudinal ridges on
dors, surface of Abd. near ant. margin. Markings on dors, surface of Abd. wavy irregular
lines of minute tubercles. No clear polished
space. No ridges. Sicilia (Palermo).
2. Gen. Cepheus C.L.Koch
1835 Cepheus, C.L.Koch. C. M. A.,
fasc. 3 1. 11 ! 1842 C., C. L. Koch, Ubers.
Arach., v.3 p. 104 ' 1855 C., Nicolet in:
Arch. Mus. Paris, 9. 1 p. 444 | 1884 C.,
Michael, Brit. Orib., v. 1 p. 284.
Body rough and sculptured,
dotted or reticulated. Dors, surface
of Abd. separated from Cephth. by
an unbroken line. Lam. large, undu-
lating, blade-like, light-colored, sloping
or horizontal. Tectp. usually very
large. Femora without thin peduncles.
Ung. trid.
5 good and 1 doubtful species.
Key to the good species.
/ Psstg. org. almost globular, sunk in Psstg. Markings
on Dors, of Abd. double rows of alternate pits;
1 ' a ridge between each double row. - - L. 600 n 3. C. ocellatus
I Psstg. org. projecting from Psstg. ; markings on Dors.
I of. Abd. irregularly scattered 2.
I Ung. hetd; Lam. separated anteriorly by a wide
2 space. L. 800 |u i.e. bifidatus
I Ung. homd. ; Lam. nearly touching anteriorly 3.
I Psstg. org. fusiform ; hairs on Dors, of Abd. long. -
3 \ L. 1,25 mm 5. C. variolosn
I Psstg. org. clavate 4.
/ Psstg. org. long and straight; Intlam. hair in the
usual position. L. 900 |u 4. C. tegeocranus . p. 32
4 / Psstg. org. short and recurved; almost pyriform;
Intlam. hair springing from base of Lam. itself.
C. bifldatus ( S5 /,).
p. 32
p. 32
p. 33
. 1 mm 2. C. latus
p. 32
32 Cepheus
1. C. bifidatus Nic. 1855 C. b., Nicolet in: Arch. Mus. Paris, p. 7 p. 446 t. 81 |
1884 C. b., Michael, Brit. Orib., v.l p. 290 t. 16 f. 8; t. 52 f. 2- -3.
Im. Dark brown or black, legs lighter ; dull, broad. Lam. wide, undu-
lated, reticulated blades set at angle of about 45 ; C. lam. broad, truncated ;
Lam. hair from middle of truncated edge; Lam. widely separated ante-
riorly ; Psstg. at edge of Cephth. directed forward and outward ; Psstg. org.
long, curved, almost filiform. Ung. very hetd. Abd. broad, rough,
rounded posteriorly, truncated anteriorly; antero-lateral angles elongated,
rough. 4 rows of large hairs on Dors, of Abd. - Nph. Of the type of
that of Tegeocranus latus, light buff, almost white, legs and Kost. pinkish;
rough granular, elliptical. Psstg. org. serrated spines; Intlam. hair
similar; Dors, of Abd. flat; cast dorsal skins of Abd. earned flat, centre of
each skin slightly in advance of that below it ; larval skin bordered by about
17 thin, curved, sparsely-serrated spines, and has 3 pairs on Dors,
of Abd. Each nymphal skin bordered by about 20 chitinous conical
papillae, each bearing an extremely long, slender, sparsely serrated
spine; some as long as the body, variously curved, hinder spines slightly
flexible. - - Im. L. ca. 800 M.
England, France.
2. C. latus Nic. 1855 C. I, Nicolet in: Arch. Mus. Paris, v.l p. 446 t. 7 f. 9 |
1877 C. I, Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v.4 p. 87 ' 1884 C.I.,
Michael, Brit. Orib., v.l p. 295 1. 17 f. 12 | 1888 C.I, Berlese, A. M. S., fasc. 49 nr. 5.
Im. Walnut-brown, legs lighter; dull, coarsely reticulated or pitted.
Rost. blunt, Lam. light colored, very wide, almost horizontal, rounded and
meeting anteriorly. Psstg. org. short, pyriform, recurved. No Intlam. hair
in the ordinary position, but a large rod-like hair springs from
the inside of each Lam. Tectp. very large. Legs shorter than those of
C. tegeocranus. Abd. broad, rounded posteriorly, truncated anteriorly, Dors,
of Abd. arched with 4 rows of very small spines. L. ca. 1 mm.
England, France, Italy, Netherlands.
3. C. ocellatus Michael 1882 C. o., Michael in: J. K. micr. Soc., ser. 2 v. 2
p. 1 1. 1 f. 19 1884 C. o., Michael, Brit. Orib., v. 1 p. 287 1. 16 f. 17 | 1896 Scutovertex o.,
Berlese, A. M. S., Crypt. II p. 46.
Im. Dark brown or almost black; dull without polish. Rost. sharp, Mdb.
small, Cephth. more than 1 / & of total length, flat. Lam. large, light colored,
fleshy-looking blades far apart, usually nearly horizontal, widest posteriorly
truncated anteriorly, a thick curved spine from lower angle of truncation
nearly touches the point of Rost., a longer thinner spine from upper angle
runs parallel to first; Trlam. a low thick ridge. Psstg. large, wide, facing
straight upward; Psstg. org. almost globular, slightly pyriform, sunk in
Psstg. Legs rough; Abd. oval, truncated anteriorly antero-lateral angles
produced forward. A broad flattened margin round Dors, of Abd. (except
ant. margin) ; this margin is transversely striated and bears a row of short,
blunt spines. Central part of Dors, of Abd. arched and divided by ridges
into bands each containing a double alternating row of pits. Nph.
Very like that of Liacarus palmicinctus, but scales much smaller and the
nervures branched irregularly instead of reticulated. - - Im. L. ca. 600 p.
On lichens. England.
4. C. tegeocranus (Herm.) 1804 Notaspis t., Hermann, Mem. apt., p. 93 t. 4
f . 3 5 [ 1844 Oribatat., Gervaisin: Walckenaer. Ins. Apt.. v.B p. 258 | 1877 Cepheus t.,
Cepheus, Tegeocranus 33
Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 -v. 4 p. 87 1884 C. t., Michael, Brit.
Orib.. v.l p. 293 1. 17 f. 1 11 j 1887 C. t., Berlese, A. M. S.. fasc. 36 nr. 2 ; 1855
C. vulgaris, Nicolet in: Arch. Mus. Paris, v. 7 p. 445 t. 31 f. 8.
Im. Color varies from light yellow-brown to black, legs lighter than
body ; coarsely reticulated (looks pitted under low amplification), rough. Host,
blunt; Lam. very wide, curved, light-colored blades meeting anteriorly,
inner ant. angles pointed. Lam. hair long, rough. Psstg. org. long, straight,-
clavate. Intlam. hair large, rod-like. Tectp. very large ; Ung. almost homd.
Abd. rounded posteriorly truncated anteriorly; Dors, of Abd. arched, with
4 rows of short spines, a smaller row round post, margin. - - Nph. About
twice as long as greatest breadth, polished, colorless, transparent; internal
organs shew and are brightly colored; appears divided into 3 parts. Psstg.
org. short, clavate, curving backward and outward. Intlam. hair very long,
flexible; a similar pair in front of them. Abd. oval, pointed posteriorly
about 12 long, flexible hairs round margin, 2 pairs on Dors, of Abd. -
Im. L. ca. 900 M.
Algeria, Belgium, England. France. Germany, Italy, Netherlands. Sweden.
5. C. variolosus Bed. 1888 C.v., Berlese in: Bull. Soc. ent. Ital., t.20 p. 216
t, 13 f. 4.
Im. Black, dors, and ventr. surfaces deeply pitted; Psstg. org.
long with fusiform head. Lam. undulated, separated anteriorly by a
narrow slit. Intlam. hair very long. Abd. round posteriorly. Femora 3 and
4 with spine-like process at distal end. Dors, of Abd. with long whitish
irregular rod-like hairs. L. ca. 1,25 mm.
Under bark of trees. Brazil (Matto-grosso).
C. punctulatus Banks 1895 Cephus (sic) p., Banks in: Tr. Amer. ent. Soc.,
e. 22 p. 10.
Im. Dark reddish ,,finely punctulate", Cephth. narrow, Lam. nearly reaching
tip of Host, truncated anteriorly n With several short and 1 long hair each side".
Intlam. hair moderate. Psstg. org. short, clavate. Abd. elliptical, with 2 short stiff
hairs on each shoulder; 6 near post, end, one each post, side; ventr. surface granular.
L. 850 |u. ~ A very insufficiently described species, not figured; doubtful if really a
Cepheus.
In decaying fungus and on weeds. New York State.
3. Gen. Tegeocranus Nic.
1835 Cepheus (part.), C. L. Koch, C. M. A., fasc. 3 1. 11 ! 1842 C. (part.), C.L. Koch.
libers. Arach., v. 3 p. 104 1855 Tegeocranus (part.), Nicolet in : Arch. Mus. Paris,
v. 7 p. 464 ; 1884 T. (part.). Michael, Brit. Orib., v. 1 p. 306 1896 T. + Tectocepheus,
Berlese, A. M. S., Crypt. II p. 46, 47 [ 1885 Carabodes (part.), Canestrini, Prosp. Acarof.,
v. 1 p. 9 i 1892 C., Trouessart in: Rev. Sci. nat. Quest, v. 2 p. 48*.
Body rough and sculptured ; dors, surfaces of Cephth. and Abd. usually
separated by a distinct unbroken line. Lam. large, thin, undulated blades,
usually more or less light-colored. Dors, of Abd. usually, but non invariably,
with a border different from the centre. Femora of first 2 pairs of legs
with thin peduncles. Ung. mond.
6 species.
Key to the species.
{Cephth. and Abd. divided by a distinct line 2.
Cephth. and Abd. not demarcated from each other.
(Tectocepheus Berl.) L. 320 \JL 6. T. velatus . . . . p. 35
Das Tierreich. 3. Lief. : A. D. Michael, Oribatidae. 3
34 Tegeocranus
/ Psstg. org. shortish, clavate 3.
2 < Psstg. org. long, setiform, pectinated on ant. edge
L. 320 |u 5. T. leporosus ... p. 35
Marginal band of Dors, of Abd. different from centre - 4.
Xo different band on Dors, of Abd. Lam. straight
and rather small. L. 590 |U 3. T. hericius . . . . p. 34
Upper edge of Lam. curved, without points -,5.
Lam. with 2 strong points or angles projecting
from upper edge. L. 800 y. 2. T. dentatus ... p. 34
I Lam. separated anteriorly by a considerable space
5 j - L. 900 n . . . . 4. T. latus p. 35
I Lam. joined at their ant. bases. L. 620 (a ... 1. T. cepheiformis . p. 34
1. T. cepheiformis Nic. 1855 T. c., Nicolet in: Arch. Mus. Paris, v. 1 p. 465
t. 9 f . 1 1885 T. c-i Michael in: Abh. Ver. Bremen, v. 9 p. 207 ; 1888 T c., Michael.
Brit. Orib., r.2 p. 340 t, 25 f. 1 7 || non 1886 T.c., Berlese, A. M. S., fasc. 33 nr. 7.
Im. Very like T. latus, but Lam. at their ant. ends joined at the base
and above the join leaving only a thin line between them instead of a
considerable space as in T. latus. Dors, of Abd. irregularly reticulated by
raised rough ridges, reticulations large, spaces between slightly depressed,
granular. - - Nph. Resembles that of T. latus, but the projections from the
edge of Dors, of Abd. and of each skin of the Dors, of Abd. different in form,
and each bears a transparent leaf-like scale with spinulated midrib instead of
a serrated spine. - - Im. L. ca. 620 u.
In fir woods etc. England, France, Germany.
2. T. dentatus Michael 1888 T. d., Michael, Brit. Orib., v.2 p. 338 t. 26 f. 17,9.
Im. Very like T. latus but differs in having Lam. with a blunt
point at ant. end bearing Lam. hair; post, to this outline concave but
rises to a large sharp point about the middle. Psstg. org. long, almost
filiform, distal end truncated. Dors, of Abd. (except border) reticulated,
reticulations large, rough, irregular, spaces between sunk and finely dotted.
- Nph. Like that of T. latus but the brown projections round margin
of i .Dors. of Abd. are quadrifid, not trifid as in T. latus, the extra point being
at the insertion of the serrated spine, which spine is longer than in T. latus
- Im. L. ca. 800 |u.
On dead wood. England.
3. T. hericius Michael 1888 T. h., Michael, Brit. Orib.. v. 2 p. 346 t. 25 f. 8 13;
t. 26 f. 10.
Im. Dark yellow-brown, very rough, pyriform. Cephth. long narrow;
Dorso-vertex reticulated. Lam. nearly straight, reticulated blades on edge,
smaller than those of T. latus, near together anteriorly. Psstg. org. almost
upright, with peduncles of moderate length, and large, shortly pyriform, rough
head. Whole edge of Dors, of Abd., except ant. margin, broken by strong,
rough projections bearing thick curved hairs. Dors, of Abd. not much
arched, edge slightly turned upward; no border, ornamentation similar
over whole Dors, of Abd. vizt irregular reticulations formed of very strong
rough ridges with deep granular pits between. - Nph. Of the type of
that of T. latus. but the projections round Dors, of Abd. and dorsal skins
mostly split into 3 bifid lobes and the large spines spinulated, not merely
serrated. Im. L. ca. 590 M.
On oak-trees. England.
Tegeocranus 35
4. T. latus (C.L.Koch) 1835 Cepheus I., C.L.Koch, C. M. A., fasc. 3 1. 11 (pro-
bably) 1884 Tegeocranus I, Michael, Brit. Orib.. v. 1 p. 310, 340 t. 19 f. 19 j 1885
Carabodes 1., Canestrini, Prosp. Acarof., v. 1 p. 42 ! ? 1886 Tegeocranus cepheiformis,
Berlese, A. M. S., fasc. 33 t. 7.
Im. Dark, dull brown. Rough; Shape broad, rounded ; Cephth. pyramidal;
Lam. large, clearly reticulated blades on edge with sinuated upper
edge without points, and separated anteriorly by a space. Trlam. a short
bar. Psstg. org. medium length, clavate, slightly recurved. Intlam. hair
rod-like. Abd. broad, rounded, truncated anteriorly, clearly divided from
Cephth. Dors, of Abd. with central part arched, covered with short irre-
gular rough corrugations; periphery (except ant. margin) has 2 borders, which
unite anteriorly; inner one broad, granular, sloping; outer with rough edge
and transverse striations; ant. corners rough, projecting. No hairs on Dors,
of Abd., a few very fine hairs on post, margin. - - Lv. Brownish -white,
semi-opaque, granular, oval. Intlam. hair strongly serrated, slightly curved
spines of clear chitiu ; 4 pairs of similar spines on dors, surface and 6 pairs
on post, margin of Abd. - Nph. Color and texture as in Lv., but rather
darker and rougher; form elliptical, getting broader with each ecdysis. Dors,
of Abd. nearly flat, cast skins carried, larval in centre, nymphal forming con-
centric borders. Round margin of each nymphal skin are about 20 large,
flat, chitinous, brown, trifid or quadrifid projections. The central lobe of
each projection bears a long, doubly curved, strongly serrated spine of clear
chitin. - - Im. L. ca. 900 |u.
On dead wood. Algeria, England, France, Germany, Sweden.
5. T. leporosus (Haller) 1884 Eremaeus I., Haller in: Arch. Naturg., v. 501
p. 225.
Im. Covered with irregularly scattered raised dots ; Lam. very com-
plicated forming 2 curves joined at their points, both concave in-
wards, the .hinder shell-like, much wider but not longer than the anterior.
Trlam. a line. C. lam. large. Rost. hairs and Lam. hair setiform, equal-
sized; Psstg. org. long, setiform, pectinated on the ant. edge, Abd. with a
distinct border marked like remainder of dors. 8 longitudinal rows of short
setiform hairs on Dors, of Abd. L. 320 ja.
America.
6. T. velatus Michael 1880 T. v., Michael in: J. E. micr. Soc., v. 3 p. 189 t. 6
f. 6 9 ; 1884 T.v., Michael, Brit. Orib., v. 1 p. 313 t. 31 f. 9 15 | 1895 Tectocepheus
v., Berlese, A. M. S., fasc. 77 nr. 2.
Im. Dull brown, not dark; roughened by irregular raised dots which
are on an outer easily detached skin which rubs off in parts leaving smooth
chitin below. Cephth. long; Lam. large, detached, nearly horizontal,
dotted blades. C. lam. long, nearly reaching tip of Rost. Trlam. a line
only but distinct. Psstg. org. long, with slender peduncles and almost dis-
coidal head bearing scattered points. No Intlam. hair. Legs not half the
length of body, coxae and Femora broad and flat. Dors, of Abd. not much
arched; without border; antero-lateral corners flat and very projec-
ting, making ant. margin; the widest part of Abd. 6 very short hairs round
post, margin. - - Nph. Pale grey, slightly granular. Rost. blunt. Cephth.
with 4 longitudinal and some transverse ridges. Psstg. org. much like Im.,
but heads more pyriform. Abd. and Cephth. almost coalesce. There is a
3*
36
Tegeocranus, Carabodes
raised, longitudinal, irregular band and about 4 raised transverse curved
bands each composed of 3 or 4 parallel wrinkles. - - Im. L. ca. 320 |u.
In moss. England.
4. Gen. Carabodes C. L. Koch
1835 Carabodes, G. L. Koch, C. M. A.,
fasc. 3 t. 15 ; 1842 C., C. L. Koch, libers.
Arach., v. 3 p. 107 j 1885 C. (part.), Cane-
strini, Prosp. Acarof., v. 1 p. 9 i 1895 C.,
Banks in : Tr. Amer. ent. Soc., v. 22 p. 12 |
1896 C., Berlese, A. M. S., Crypt. II p. 45 1855
Tegeocranus (part.). Xicolet in : Arch. Mus.
Paris, v. 1 p. 464 | 1884 T. (part.), Michael,
Brit. Orib., a 1 p. 306.
Body rough and sculptured, dors,
surface of Cephth. and Abd. separated
by a distinct unbroken line or deep
furrow. Lam. dark, low, solid ridges
or thick horizontal plates. Femora of
first 2 pairs of legs with thin peduncles.
Ung. mond.
10 good and 3 doubtful species.
Key to the good species.
Fig. 10. C. coriaceus (**/i).
1 1
Dors, of Abd. with a marginal band differently marked
from the central part 2.
Dors, of Abd. without any differently marked marginal
band -- 6.
Cephth. with longitudinal ridges between the 2 Lam.
Dors, of Abd. hairless 3.
No longitudinal ridges between Lam. Dors, of Abd.
bearing hairs 4.
3 toothed ridges between Lam. Markings on central
part of Dors, of Abd deep pits divided from mar-
ginal band by a raised ridge. L. 590 |n . . .
Only 1 central, untoothed ridge on Cephth. Markings
on central part of Dors, of Abd. a few almost
straight bands with undulated sides. L. 620 jn
Abd. broad ; hairs bordering it short, fine, radial - 5.
Abd. very long, hairs bordering it thick and curved,
lying along its edge. L. 680 u 5. C. elongatus . p. 37
Markings on central part of Dors, of Abd. numerous,
long, rough, irregular curved bands. - - L. 620 ja 4
Markings on Dors, of Abd. raised dots. - - L. 500 |u 9
Markings on Dors, of Abd. raised dots 7.
Markings on Dors, of Abd. ridges 9.
Hairs on post, margin short and fine 8.
Hairs on post, margin capitate. L. 400 |U . . . . 7.
Cephth. arched between Lam. L. 450 |n .... 8.
Cephth. with an elevated triangle, with concave sides
between Lam. L. 400 u . . . 2.
3. C. clypeatus . p. 37
C. femoralis . . p. 38
C. coriaceus . p. 37
C. marginatus . p. 39
C. granulatus . p. 38
C. labyrinthicus . p. 38
Ridges longitudinal 15 or 16. L. ca. 1,25 mm . . 10.
Ridges forming an irregular reticulation. L. 450 \JL 1.
C. brevis . . . p. 37
C. papillosus . p. 39
C. apicalis . . . p. 37
Carabodes 37
1. C. apicalis Banks 1895 C. a., Banks in: Tr. Amer. ent. Soc., v. 22 p. 13.
Im. Apparently very like C. labyrinthicus but some small ridges on
Dorsb-vertex. Dors, of Abd. irregularly reticulate with ridges; only 2
longitudinal sub-median rows of 3 spatulate hairs, but 10 or 12 similar
round post, margin; ventr. surface smooth ; genital opening circular, close
to anal. L. 450 |u.
New York State.
2. C. brevis Banks 1896 C. b., Banks in: Tr. Amer. ent. Soc., v. 23 p. 77.
Im. Dark red-brown, legs yellowish. Cephth. with a long trian-
gular elevated central region with concave sides. Lam. plate-like.
Psstg. org. with short peduncles and globular head. Abd. a little longer
than broad, truncated anteriorly, rounded posteriorly. Dors, of Abd. strongly
pitted and bearing 4 longitudinal rows of stiff curved hairs. Apparently very
like C. labyrinthicus except for the central triangle on Cephth. L. 400 |n.
In dead fungus. New York State.
3. C. clypeatus (Nic.) 1855 Tegeocranus c., Nicolet in: Arch. Mus. Paris, v. 1
p. 467 t. 9 f. 3.
Im. Black, legs dark brown. Psstg. orange-yellow. Rough, dull.
Lam. straight, thick, smooth ridges; between them 3 parallel toothed
ridges; Psstg. very lateral; Psstg. org. moderate length, clavate; Abd.
much longer than broad, sinuated and projecting in centre over Cephth.,
rounded posteriorly. An elliptical depression with bordering ridge and reti-
culated centre occupies centre of Dors, of Abd.; 3 converging conical
elevations, point forward, spring from inner post, edge of ridge. In front
of ellipse a trapeze-shaped depression bordered by a ridge and with a raised
oval in centre. Outside oval and trapeze a concave border with a raised
ridge at outer edge. 2 hairs on post, margin, none on Dors, of Abd.
L. 590 M.
France. Rare.
4. C. coriaceus C. L. Koch 1835 C. c., C. L. Koch, C. M. A., fasc. 3 1. 15 1884
Tegeocranus c., Michael, Brit. Orib., v. 1 p. 316 t. 20 f. 1 8 ; t. 22 f. 11 | 1896 Cepheus c.,
Oudemans in: Tijdschr. Ent., v. 39 p. 54 || non 1886 Carabodes c., Berlese, A. M. S.,
fasc. 33 nr. 8.
Im. Black, dull, very rough; chitin thick, Cepth. long broad, as wide
as ant. margin of Abd. Dorso-vertex lighter with 2 small curved ridges
which join a large, rough, transverse ridge with a central depression, a deep
trench between this ridge and ant. margin of Abd. Lam. large, thick,
rough, horizontal, almost reaching point of Kost. No Tiiam. Psstg. org.
long, proximal portion nearly straight, terminal clavate, rough. Legs short,
Femur rough. Abd. nearly as broad as long, margin (except ant.) with
broad, flat border with irregular edge and radiating ridges. Remainder of
Dors, of Abd. divided into numerous rough curved bands marked
like Morocco-leather. 4 rows of spatulate white hairs on Dors, of
Abd. - - Nph. Like a Tyroglyphus, semi-transparent, white; Rost. and legs
pink, polished, no markings. Body very thick with 2 rows of long hairs.
Legs short. - - Im. L. ca. 620 M.
In fungi etc. Algeria, England, Germany, ? Netherlands.
5. C. elongatus (Michael) 1879 Tegeocranus e., Michael in: J. R. micr. Soc.,
v. 2 p. 250 1. 10 | 1884 T. e., Michael, Brit. Orib., v. 1 p. 324 t. 22 f. 410 1888 Cara-
38 Carabodes
bodes e., Berlese, A. M. S., fasc. 50 nr. 1 j ?1893 Hermannia oblonga, Karpelles in: Math,
naturw. Ber. Ungarn, v. 11 p. 93 t. 20 f . 3 [ ?1895 Carabodes o., .Banks in: Tr. Amer. ent.
Soc., v. 22 p. 13.
Im. Black, rough, dull; very long-shaped, line of Cephth. and Abd.
continous. Cephth. as broad as ant. margin of Abd. Cephth. and Dors, of
Abd. coarsely pitted. Lam. narrow, reticulated. No Tiiam. Psstg. org. long,
recurved, gradually widened, square ended with 3 or more projecting points
on the end continued as ridges on the side. Lam. and Intlam. hair long,
thick, recurved. Abd. widest posteriorly. Dors, of Abd. (except ant. margin)
with narrow, transversely striated border and 4 rows of thick curved hairs;
a row round margin of smaller similar hairs recurved and arranged
along the margin, not radiating. - - Nph. White or grey, having the
form of the Im. ; otherwise very similar to that of C. coriaceus. Im.
L. ca. 680 ja.
In decayed wood, moss etc., Nph. burrows. Algeria. England, Italy.
6. C. femoralis (Nic.) 1855 Tegeocranus f., Nicolet in: Arch. Mus. Paris, v.7
p. 466 t. 9 f. 2 | 1884 T.f.. Michael. Brit. Orib., 0.1 p. 318 t. 20 f . 9 ; 1885 Carabodes f.,
Canestrini, Prosp. Acarof., ' v. 1 p. 43 | 1886 C.f., Berlese, A. M. S., fasc. 33 nr. 9.
Im. Black, rough; chitin thick. Cephth. narrower than ant. margin
of Abd., flat trapeze-shaped. Dorso-vertex with central ridge, and a trans-
verse ridge arid trench almost similar to those of C. coriaceus. Lam. thick,
horizontal, raised in median line forming an angle, massive. No Lam.
hair, Intlam. hair nor Trlam. Psstg. org. long filiform. Legs short,
femora rough. Abd. longer than broad, almost parallel sided, round poste-
riorly. Ant. margin of Abd. raised forming a ridge. Abd. (except ant. margin)
with border like C. coriaceus. Dors, of Abd. within margin divided into
5 or 6 longitudinal irregular raised bands. No hairs on Dors, of
Abd. L. ca. 620 |u.
England, France, Italy.
7. C. granulatus Banks 1895 C. g., Banks in: J. N. York ent. Soc., v. 3 p. 128.
Im. Black, Cephth. granular, broad, truncate anteriorly. Lam. ..plate-
like". Intlam. hair and Psstg. org. short clavate. Abd. rougly granular,
longer than broad, parallel sided, truncated anteriorly, rounded posteriorly.
4 pairs of short capitale hairs on post, margin. 4 longitudinal rows
of hairs on Dors, of Abd. Genital opening l 1 ^ times as long as broad.
Very like C. labyrinthicus, except capitate hairs on post, margin. L. 400 in.
Imperfectly described and not figured.
In sphagnum-swamp. New York State.
8. C. labyrinthicus (Michael) 1879 Tegeocranus L, Michael in: J. R. micr.
Soc., t>.2 p. 249 1. 11 f. 2, 3 | 1884 T. I, Michael, Brit. Orib., v. 1 p. 319 t. 21 f. 1 4,
68 ; t. 22 f. 3.
Im. Dark red-brown, rough, dull. Rost. blunt, Cephth. broad, rough,
reticulated. Lam. thick, narrow, horizontal, almost reaching tip of Rost.
No Trlam. nor Intlam. hair. Psstg. org. recurved with slender peduncles and
pyriform head excavated on one side. Femora enlarged, rough. Abd. longer
than broad, antero-lateral corners projecting. Dors, of Abd. much arched,
no border, covered with large irregular raised dots; 4 rows of short curved
hairs on Dors, of Abd., a row of straight hairs round post, margin.
- Nph. Almost white, Rost. and legs brownish; semi-transparent, thick
and fat-looking; a few transverse wrinkles near Dors, of Abd. 4 pairs of
Carabodes 39
longish straight hairs on post, margin; a few shorter and curved on Dors,
of Abd. and sides. - - Im. L. ca. 450 |u.
In moss. England.
9. C. marginatus (Michael) 1884 Tegeocranus in., Michael, Brit. Orib., c. 1
p. 322 t, 21 f. 5; t. 22 f. 12 i 1886 Carabodes coriaceus, Berlese, A. M. S., fasc. 33 nr. 8.
A species intermediate between C. coriaceus and C. labyrinthicus. -
Im. Dark red-brown, rough, dull. Cephth. large, broad, rough. Lam. large,
thick horizontal, reticulated, nearly reaching tip of Rost. Trlam. a line but
well marked. Transverse ridge and trench as in C. coriaceus. Psstg.
org. curved forward, gradually thickened till near end, then deminishing.
Rough with long spines on outer side of curve. Margin of Dors, of
Abd. bordered by a flat band with radiating ridges. Remainder
of Dors, of Abd. covered with large irregularly placed, raised
dots. - - Nph. Very like that of C. coriaceus. - - Im. L. ca. 500 |u.
In moss. Algeria. England, Italy.
10. C. papillosus (H.Luc.) 1849 Oribatesp., H.Lucas in: Expl. Algerie, An.
artic., p. 319 t. 22 f. 12.
Im. Black, rough, coarsely granular, a dusty shade in parts. Species
somewhat resembles C. femoralis. Cephth. narrower than ant. margin of Abd.
flat, trapeze-shaped. Dorso-vertex with central ridge and a transverse
ridge and trench almost similar to those of C. coriaceus. Lam. thick,
massive, raised. No Lam. hair, Intlam. hair or Trlam. (apparently) Psstg.
org. not described. A strong pointed projection between legs 2 3.
Abd. almost orbicular, with numerous (15 or 16) deep irregular longi-
tudinal sulcations with ridges between; 4 sparse longitudinal rows
of thick, spatulate whitish hairs on Dors, of Abd. No different
border. L. 1,25 mm.
Algeria.
C. cephalotes C. L. Koch 1835 C. c., C. L. Koch, C. M. A., fasc. 3 1. 16 1 1840
Nothnis cynocephalus, C. L. Koch, C. M. A., fasc. 30 t. 8 | 1842 Carabodes c., C. L. Koch,
Ubers. Arach., c. 3 p. 107.
Im. Small, dark brown, rough. Psstg. org. long, clavate; Abd. about twice
as long as Cephth. half as long again as broad bordered with rather distant short
straight hairs arranged radially. Apparently a good species and belonging to this
genus from description of Femora, but imperfectly described and not found since Koch.
Germany.
C. dorsalis Banks 1896 C.d.,. Banks in: Tr. Amer. ent. Soc., v. 23 p. 77.
Im. Pale brownish yellow. Cephth. convex with outlines of a broad T-mark.
Psstg. org. clavate, of moderate length. Abd. 1V-2 longer than broad, sides nearly
parallel; rounded posteriorly, coarsely granular; 4 longitudinal rows of stiff curved
hairs on Dors, of Abd. L. 450 ,u.
New York State.
C. niger Banks 1895 C.nigra, Banks in: Tr. Amer. ent. Soc., v. 22 p. 12.
Im. Black, legs yellowish or reddish; apparently very like C. marginatus,
possibly identical, but with spatulate Intlam. hair. Psstg. org. not stated to be curved
or spiniferous ; genital and anal openings approximate. L. 500 |a.
In fungi etc. New York State.
40 Liacarus
5. Gen. Liacarus Michael*)
1842 Oribates (part.). C. L. Koch, libers. Arach., v. 3 p. 99 ] 1855 Leiosoma
(non Stephens, Coleopt.. 1831!), Nicolet in: Arch. Mus. Paris, r.7 p. 439 j 1884 L.,
Michael, Brit, Orib., v.l p. 273 1885 L., Canestrini, Prosp. Acarof., v.l p. 7 ! 1896 L.,
Berlese, A. M. S., Crypt. II p. 58 | 1892 Liosoma, Trouessart in : Rev. Sci. nat. Quest,
.2 p. 48*.
Body smooth. Last 3 pairs of legs inserted under the body. Ung. trid.
in all known species.
9 good and 1 doubtful species.
Key to the good species.
| Abd. oval, elongated 2.
\ Abd. as broad as long 6.
f A central mucro beetwen the ant. ends of the Lam. 3.
\ No such mucro 5.
f Mucro short, scarcely if at all passing points of C. lam. 4.
\ Mucro long, almost reaching tip of Rost. L. 1,25 mm 7. L. nitens . . . . p. 41
( Psstg. org. fusiform with a terminal hair. L. 850 |u 1. L. coracinus . . . p. 40
4 ' Psstg. org. with short peduncle and globular head
- L. 450 M 2. L. globifer . . . . p. 41
- Abd. with a suture each side forming a'lat border.
. J Psstg. org. long and setiform L. 880 |u . . 6. L. marginatus . . p. 41
\ Abd. without lat. border. Psstg. org. short and
clavate. L. 650 (a 8. L. ovatus . . . . p. 42
f Abd. globular (hemispherical) 7.
) Abd. shield-shaped. L. 1,05 mm 4. L. lativentris ... p. 41
f Abd. with 4 long hairs on post, margin L. 700 ju 5. L. longipilis ... p. 41
I Abd. hairless 8.
Lam. far apart anteriorly connected by a slender
Trlam. Psstg. org. clavate. L. 1 mm ... 9. L. palmicinctus . p. 42
Lam. joined anteriorly. Psstg. org. fusiform sessile.
- L. 1,02 mm 3. L. globosus . . . p. 41
1. L. coracinus (C.L.Koch) 1841 Oribates c., C.L.Koch, C. M. A., fasc. 31
t. 1 1885 Leiosoma c., Canestrini, Prosp. Acarof., v. 1 p. 21 j 1885 L. c., Berlese, A. M. S..
fasc.' 20 nr. 3 ?1841 Oribates fuscus, C. L. Koch, C. M. A., fasc. 31 t. 2 | ? 1854 Oribates
politus, C. L. Koch & Berendt, Die im Bernstein befindlichen organischen Reste, v. 1 II
p. 110 t. 17 f. 112 ' 1855 Leiosoma similis, Nicolet in: Arch. Mus. Paris, v. 7 p. 442 t. 6
f. 2 j 1884 L. simile, Michael, Brit. Orib., v.l p. 276 t. 13 f. 1 7.
Im. Dark brown, polished, Rost. with a small point on each side.
Cephth. much narrower than ant. margin of Abd. Dorso-vertex raised in
centre, raised part pointed anteriorly. Lam. blades slightly sinuate, their
ant. ends deeply indented, terminating in 2 curved points. Lam. hair spines
springing between the points and passing tip of Eost. Psstg. org. fusi-
form, sessile, terminated distally by a fine hair. Blades to femora
of all legs. Abd. a long ellipse truncated anteriorly, almost pointed poste-
riorly ; Dors, of Abd. polished, hairless, much arched ; anal covers large,
square, almost reaching post, margin. - - Nph. Almost colorless except tip
of Rost. and Tarsus, polished, oblong, Psstg. org. short fusiform. Dors, of
Abd. with a sparse marginal row of flexible hairs. - Im. L. ca. 850 |u,
a large variety ca, 1,1 mm, according to Berlese 1,5 mm.
Algeria, England, France.
*) The name Leiosoma Nicolet is preoccupied. The name I have used is derived
from Xetoq (smooth) and dxapi (acarus). A. D. Michael.
Liacarus 41
2. L. globifer (P. Kram.) 1897 Leiosoma g., P. Kramer in: Bibl. zool., .20 p. 80.
Im. Light coffee-brown ; polished. Cephth. long. Host, rounded. Lam.
short, almost meeting at their ant. ends; a short unpaired spine directed
forward between them as in L. nitens. Psstg. org. short, with thin
peduncles and globular head. C. lam. longish, hearing a Lam. hair,
which reaches beyond tip of Rost. Tectp. 1 large and scoop-shaped. L. 450 ju.
Greenland.
3. L. globosus (Berl.) r'1844 Oribates subterraneus, C. L. Koch, C. M. A.,
fasc. 38 t. 11 ! ?1844 0. globosus, C.L.Koch, C. M. A., fasc. 38 t. 12 j 1885 Leiosoma
globosum, Berlese, A. M. S., fasc. 20 nr. 4.
Im. Black or black-brown, highly polished. Lam. blades on edge,
widest anteriorly, not distinctly separated from Trlam., which is also a blade
on edge. C. lam. short, broad, much truncated. Psstg. org. fusiform with
a terminal distal hair but no peduncle. Tectp. 1 with long terminal hair
similar to and close to Lam. hair. Abd. globular; 4 minute hairs on post,
margin ; Dors, of Abd. hairless. L. 1,02 mm (Beiiese's description).
It is not possible to say for certain from Koch's description and plate whether his
2 species (which are one only) are really the same as Berlese's, the present writer in-
clines to think them different and belonging to the genus Notaspis, but Berlese thinks
them identical.
In moss. Germany, Italy.
4. L. lativentris (Nic.) 1855 Leiosoma I. , Nicolet in: Arch. Mus. Paris, v. 7
p. 444 t. 6 f. t>.
Im. Cephth. and legs dull red -brown, Abd. black, polished. Like
L. globosus, but outline of Abd. instead of being circular is
,,Heraldic shield-shape d", i. e. .truncated and slightly sinuated ante-
riorly, centre slightly advancing and convex; lateral parts slightly concave.
Abd. somewhat pointed posteriorly, as broad as long. L. 1,05 mm.
France.
5. L. longipilis (Monz.) 1894 Leiosoma I, R. Moniez in: Rev. biol. Nord
France, v. 6 p. 201.
Im. Pitch-black, polished. Psstg. org. clavate, finely serrated.
Intlam. hair ca. 225 |u. Abd. hemispherical, truncated anteriorly. About
5 long radiating hairs ofca. 160 170 |u round post, margin. L. ca.
700 n.
In ants' nests. New Zealand.
6. L. marginatus (Me.) 1855 Leiosoma marginata, Nicolet in: Arch. 3Ius.
Paris, v. 1 p. 442 t. 6 f. 3.
Im. Red -brown, polished. Lam. blades on edge, widest anteriorly.
Lam. hair long; no Trlam. Psstg. org. as long as Cephth., setiform,
slightly recurved. Abd. elongated, truncated and slightly sinuated anteriorly
almost pointed posteriorly, hairless, but with a longitudinal suture on
each side of dors, surface forming a lat. border. L. 880 |u.
Under stones. France. Rare.
7. L. nitens (Gerv.) 1844 Oribata n., Gervais in: Walckenaer, Ins. Apt., v. 3
p. 259 t. 35 f. 7 | 1855 Leiosoma n., Nicolet in: Arch. Mus. Paris, v. 7 p. 441 t. 6 f . 4 |
1885 L. n., Berlese, A. M. S.. fasc. 20 nr. 2.
42 Liacarus
Im. Black, polished. Lam. short, much widest anteriorly. C. lam.
very long and large, slightly truncated anteriorly. Lam. hair very long,
passing far beyond tip of Host. Trlam. with a great central mucro
which nearly reaches tip of Rost. Psstg. org. setiform. Abd. oval,
rounded anteriorly narrower posteriorly, hairless except 4 short hairs on post,
margin. L. ca. 1,25 mm.
In moss and Woods. France, Italy, Netherlands.
8. L. ovatus (C. L. Koch) 1840 Oribates o., C. L. Koch, C. M. A., fasc. 30 f. 24
| 1855 Leiosoma ovata, Nicolet in: Arch. Mus. Paris, v.7 p. 443 t. 6 f . 5 ; 1884 L. ovatum,
Michael, Brit, Orib.. v.l p.278 1. 13 f.8 14; 1896 L. o., Berlese. A.M.S., Crypt. H p. 92
| 1876 L. truncatum, Canestrini & Fanzago in: Atti Soc.Veneto Trent., t. 5 p. 105 , 1880
L. pyrigerum, Berlese. A. M. S., i'asc. 50 nr. 2.
Im. Brown, shades vary in different specimens ; smooth, not polished,
finely punctured ; oval, rather broader than L. coracinus and more pointed
posteriorly. Dorso-vertex without any median point, Lam. blades on
edge, truncated anteriorly. No C. lam. nor indentation of the truncated
edge. Lam. hair from bottom of edge. Rost. hairs and Lam. hair very
slightly imbricated. Psstg. org. very short and clavate. Dors, of Abd.
not much arched, 2 rows of fine hairs on it and a marginal row; anal
covers like L. coracinus. Nph. Very like those of L. coracinus and
Cepheus tegeocranus. - - Im. L. ca. 650 ju.
England, France, Germany, Italy, Netherlands.
9. L. palmicinetus (Michael) 1884 Leiosoma palmicinctum, Michael, Brit. Orib. r
v. 1 p. 280 t. 14 f. 16; 1. 15 f. 16 j 1896 L.p., Berlese. A. M. S., Crypt. II p. 58.
Im. Dark brown, smooth, not polished. Rost. blunt, Cephth. broad,
short. Lam. narrow, slightly undulated blades on edge; C. lam. small but
longish, Trlam. distinct, forming a ridge. Psstg. org. of medium length,
peduncles thin, head slightly lanceolate; sternum very finely serrated. Abd.
almost circular, slightly truncated posteriorly, without markings; ventr. plate
much smaller than dors, the latter reflexed posteriorly. Nph. Very remar-
kable. Rost. blunt; Cephth. very short and broad; legs short. Abd. elliptical,
somewhat raised towards centre ; round the edge are about 1 6 large apophyses,
from each of which springs a large transparent scale of a Japanese fan shape,
or the shape of the scale of Lycaena argus, each scale has radiating
nervures and finer cross nervures, the scales overlap all round. Nph. carries
the cast skins of Lv. and of its own earlier stages, each skin is bordered
by these scales, which entirely cover the whole creature including Rost. and
legs, except the larval skin ; each nymphal skin also carries 4 apophyses from
which spring very long nairs, these are usually, but not invariably, broken
off from the cast skins. - - Im. L. ca. 1 mm.
England.
L. olivaceus (Grube) 1859 Leiosoma olivacewm, Grube in: Arch. Dorpat. Ges. T
ser. 2 v. 1 p. 476.
Im. Olive-brown; very finely granular; most like L. marginatus, but has margin
finely striated; Psstg. org. clavate; 2 longitudinal rows of 5 conspicuous hairs on Dors,
of Abd. near margin, 6 very small clavate hairs on post, margin. L. ca. 400 |u.
Germany.
Notaspis
43
6. Gen. Notaspis Herm.
1804 Notaspis, Hermann, Mem. apt., p. 87
| 1855 N., Nicolet in: Arch. Mus. Paris, v. 7
p. 446 j 1888 N., Michael. Brit. Orib., v. 2 p. 349
1891 N., Canestrini in: Atti 1st. Veneto, ser. 7
v. 2 p. 718 | 1892 N., Trouessart in: Rev. Sci. nat.
Quest, v. 2 p. 48* ] ?1826 Belba, Heyden in: Isis,
p. 613 | 1835 & 42 Eremaeus -f- Oribates (part.)
-\- Zetes (part.) -f- Oppia (part.) -f- Mttma(Nph.)
(part.), C. L. Koch, C. M. A., fasc. 3 etc. & Ubers.
Arach., v. 3 p. 99, 101, 102, 104 i 1859 Oppia,
Grube in: Arch. Dorpat. Ges., ser. 2 v. 1 p. 417
1877 0., Canestrini & Fanzago in: Atti 1st.
Veneto, ser. 5 v. 4 p. 77 | 1885 0., Canestrini,
Prosp. Acarof.. v. 1 p. 7 [ 1896 0., Oudemans in:
Tijdschr. Ent., v. 39 p. 54 | 1896 0. + Dameo-
soma (part.) -}- Oribatula -f- Eremaeus, Berlese,
A. M. S., Crypt, n p. 53, 54, 57, 59.
Body smooth. Last 3 pairs of legs
inserted at edge of body.
26 good and 2 doubtful species, 2 varieties.
Key to the good species.
I Ant. corners of Dors, of Abd. projecting as small free blades. (Div. I. - - Genus
Oribatula Berl.) 2.
Ant. corners of Dors, of Abd. not projecting (Div. II.) 5.
f Ung. trid. 3.
\ Ung. mond. L. 550 |u 3. N". similis
Fig. 11. N. pilosa C" /,).
3
f Having hairs on Dors, of Abd. 4.
Dors, of Abd. hairless. L. 450 fi ....... 2. N". plantivaga
/ With C. lam. L. 370 n ........... 1. N. exilis . .
. . p. 45
. . p. 45
. . p. 45
. . p. 45
n
) Without C. lam. L. 480 |u 4. N. tibialis
f Ung. trid. (Subdiv. Ha.) 6.
\ Ung. mond. (Subdiv. lib.) 17.
{Lam. blades on edge 7.
Lam. mere thickened bars or ridges 11.
J Lam. with C. lam. 8.
\ No C. lam. 10.
/ C. lam. very long; Psstg. org. Intlam. hair; Host,
hairs, hairs on Dors, of Abd. and a spine on coxa
3 strongly serrated. L. 650 |a 6. N. bipilis . . . . p. 46
I C. lam. short; Psstg. org., Intlam. hair; Host, hairs
v and hairs on Dors, of Abd., not serrated. 9.
Lam. equal width throughout; much nearer together
anteriorly than posteriorly; hairs on Dors, of
Abd. thick. L. 750 |u 7. N". burrows! . . p. 46
Lam. much wider anteriorly than posteriorly very
little nearer together anteriorly than posteriorly,
hairs on Dors, of Abd. very fine. L. 340 |n 9. N. glabra . . . . p. 47
Trlam. as broad as and similar to Lam., Dors, of
Abd. with long hairs. L. 420 ILI 5. N". aequalis ... p. 46
I No Trlam. Dors, of Abd. hairless. L. 1 mm . . 13. N". microptera . . p. 48
{Dors, of Abd. bearing hairs 12.
Dors, of Abd. hairless L. 600 |u 10. M". hepatica ... p. 47
{
44 Notaspis
J Hairs on Dors, of Abd. long and serrated. L. 560 |u 16. N. serrata . . . p. 48
\ Hairs on Dors, of Abd. not serrated 13.
{Psstg. org. clavate or with globular, fusiform or
pyriform heads on thin peduncles 14.
Psstg. org. flabellate, broad, flat, almost sessile.
-L. 190 |n 11. N. licnophora . . p. 47
Psstg. org. hirsute; Intlam. hair reflexed L. 680 ILI 8. N". conformis . . . p. 47
Psstg. org. not hirsute; Intlam. hair directed
forward 15.
Intlam. hair longer than Psstg. org. Ung. almost
homd. ; a spine on each antero-lateral angle of
Abd. Lam. arising from Psstg. much wider
apart posteriorly than anteriorly 16.
Intlam. hair shorter than Psstg. org. Ung. very
hetd; no spines on antero-lateral angle of Abd.
Lam. arising some distance, in front of Psstg.
- L. 600 n 14. N. oblonga . . . . p. 48
Abd. almost round; length of hairs on Dors, of
Abd. about 2 / 3 of width of Abd. Psstg. org. with
; elongated clavate heads. L. 450 (LI . . . . 15. N. pilosa . . . . p. 48
' Abd. elongated; length of hairs on Dors, of Abd.
about 1 /s of width of Abd. Psstg. org. with almost
globular heads. L. 670 |u 12. N". lucorum ... p. 47
Terestrial 18.
Aquatic. L. 500 |u. 19. N". lacustris ... p. 49
Trlam. present 19.
16
I
8 ' No Trlam. 20.
1C. lam. present; Tibia 1 2 almost globular, Dors,
of Abd. arched in centre with a flattened margin.
- L. 340 |n 22. N. monilipes . . . p. 50
v No C. lam.; Tib. 12 not globular; no flattened
margin to Dors, of Abd. 4 longitudinal ridges
on Dors, of Abd. L. 200 |ii ....... 24. N. quadricarinata . p. 51
i Psstg. org. pectinated 21.
I Psstg. org. not pectinated 22.
{Psstg. org. clavate with a pectinated club on a
smooth peduncle. L. 330 |a 17. N. clavipectinata . p. 49
Psstg. org. long, not clavate, with a setiform dist.
end. L. 420 n 23. N. pectinata . . . p. 51
2 short bars on Cephth. between Lam. 23.
f 2
\ N
No bars between Lam. 25.
{Bars not joined by a transverse bar; Rost. not
truncated 24.
Bars joined anteriorly by a transverse bar Rost.
truncated. L. 240 |u .......... 26. N". trigona . . . . p. 51
/ Lam. very long, almost reaching tip of Rost. -
(>JL ] L. 330 |u .... ............ 21. N. longilamellata . p. 50
(Lam. short, not reaching middle of Cephth. -
L. 310 |u ................. 25. N". splendens . . p. 51
C. lam. very long. Psstg. org. clavate without
any terminal hair. L. 220 n ....... 18. N. juncta . . . . p. 49
25 { No C. lam. Psstg. org. with long thin peduncles
and fusiform head with a terminal hair. -
L. 830 n ................. 20. N. lanceolata . . p. 50
Xotaspis 45
Div. I. Sp. 1 4. With ant. corners of Dors, of Abd. projecting as a
small, free blade.
1. N. exilis Nic. 1855 N. e., Nicolet in : Arch. Mus. Paris, v. 7 p. 448 t. 3 f. 7 |
1888 N. e., Michael, Brit. Orib., v.2 p. 359 t. 28 f. 1 5 | 1877 Oppia e., Canestrini &
Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 89 ! 1885 0. e., Canestrini. Prosp. Acarof., t.l
[>. 26 ; 1895 Oribatula e., Berlese, A. M. S., fasc. 77 nr. 6 j 1879 Oribata crassipes,
L.Koch in: Svenska Ak. Handl., v. 16 p. 116 t. 4 f. 3.
Im. Light-yellow-brown, polished, very finely punctured. Lam. long
blades on edge, broader at distal than proximal ends; C. lam.
short; Trlam. a thickened line. Psstg. org. with short peduncle and pyri-
form head; Ung. trid., very hetd. Antero-lateral angles of Dors, of
Abd. project suddenly like rudimentary Ptm.; projection terminates
sharply at post. end. 2 rows of fine hairs on Dors, of Abd. and some round
margin. - - Nph. White grey or yellowish ; semi-transparent, internal organs
shew, giving color. Kost. and legs pink. Polished. Antero-lateral angles
do not project. Cephth. without markings or Lam. In other respects Nph.
like Im. - - Im. L. ca. 370 ju.
In moss etc. England. France, Germany, Italy, ? Netherlands, Siberia, Novaya
Zemlya.
2. N. plant! vaga (Berl.) 1884 Leiosoma nitens (err., non Oribata nitens
Gervais 1844!). Targioni-Tozzetti in: Ann. Minist. Agricol., p. 75 | 1892 Oppia tibialis
(err., non Notaspis tibialis Nicolet 1855!), Berlese, A. M.S., fasc. 64 nr. 1 | 1896
Oribatula plantivaga, Berlese, A. M. S., Crypt. II p. 59 || non 1895 0. tibialis, Berlese,
A. M. S., fasc. 77 nr. 5.
Im. Yellow-brown. Lam. narrow blades deminishing anteriorly. No
C. lam. nor Trlam. Intlam. hair long, setiform, would reach tip of Host.
Psstg. org. veiy short, sessile, pyriform, almost hidden by advancing corner
of Abd. obovate; the ant. portion of edge of Dors, of Abd. very slightly
projecting; a few very fine hairs round margin, none on Dors, of Abd.
Tactile hairs of legs 1, 2, borne on apophyses. Ung. trid. L. ca. 450 ju.
In moss. Italy.
2 a. N. plantivaga var. sicula (Berl.) 1892 Oppia tibialis var. s., Berlese,
A. M.S., fasc. 64 nr. 1.
Abd. hairless, longer than type. Palermo.
3. N. similis Michael 1888 N. s., Michael, Brit. Orib., v. 2 p. 363 t. 31 f. 912
1896 Oribatula s., Berlese, A. M. S., Crypt. II p. 9.
Im. Almost exactly like N. tibialis but a trifle larger and ung. mond.
L. ca. 550 n.
In moss and leaves. England.
4. N. tibialis Nic. 1885 N. t., Nicolet in: Arch. MusT Paris, v. 7 p. 449 t. 3
f. 8 | 1888 N. t., Michael. Brit. Orib., v.2 p.262 t. 28 f. 6 8 | ?1895 Oribatula t.,
Berlese, A. M. S., fasc. 77 nr. 5 || non 1892 Oppia t., Berlese, A. M. S., fasc. 64 nr. 1.
Im. Chestnut -brown, polished, veiy finely punctured. Lam. long
blades on edge, narrower at distal than proximal ends. No C. lam.
Trlam. a line, Psstg. org. with slender peduncle of moderate length
and recurved fusiform head roughened by points. Ant. Tibia with
rounded elevation at distal end from which tactile hair springs. Ung. trid.
strongly hetd. Antero-lateral corners of Dors, of Abd. project strongly, like
rudimentary Ptm., they project more than those of N. exilis, but not so
46 Notaspis
suddenly, and the projection is earned further back. Hairs on Dors, of Abd.
as in N. exilis. - Nph. scarcely distinguishable from that of N. exilis. -
Im. L. ca, 480 p.
In moss and dead leaves. England, France, Germany, ? Italy.
Div. II. Sp. 526. Ant. corners of Dors, of Abd. not projecting.
Subdiv. II a. Sp. 516. Ant. corners of Dors, of Abd. not projecting
and Ung. trid.
5. N. aequalis Michael 1890 N.ae., Michael in: P. zool. Soc. London, p. 419
t. '61 f. 5.
Im. Yellow-brown, polished. Psstg. org. with long fine peduncle and
gradually clavate head. Lam. blades on edge. Lam. and Trlam. for-
ming a continuous band of equal breadth without demarcation between
them. No C. lam. Ung. very hetd. Abd. elliptical without markings.
4 longitudinal rows of long curved hairs on Dors, of Abd. a few on post,
margin, a spine on shoulder. L. ca. 420 ju.
Algeria.
6. N. bipilis Herm. 1804 N. b., Hermann, Mem. apt., p. 95 [ 1855 N. b., Nicolet
in: Arch. Mus. Paris, v.7 p. 448 t. 3 f . 6 | 1883 N.b., Berlese in: Bull. Soc. ent. ItaL,
v. 15 p. 219 j 1888 N.b., Michael, Brit. Orib., v. 2 p. 356 t. 27 f. 19 | 1844 Oribata b.,
Gervais in: Walckenaer, Ins. Apt., v. 3 p. 295 | 1877 Oppia b., Canestrini & Fanzago
in: Atti 1st. Veneto, ser. 5 v.4 p. 88 | 1885 0. b., Berlese, A. M. S. fasc. 20 nr. 8 j ?1840
Oribates badius, C.L.Koch, C.M. A., fasc. 30 t. 23 | ?1842 Oppia badia, C.L.Koch, libers.
Arach., v. 3 p. 195 J 1844 0. cornuta, C. L. Koch, C. M. A., fasc. 38 t. 8 | 1879
O. sphaerica, L.Koch in: Svenska Ak. Handl., v. 16, p. 117 t. 4 f. 5 j 1841 Murcia
acuminata, C. L. Koch, C. M. A., fasc. 31 t. 24 (Nph.)
Im. Red-brown, polished. Rost. long, pointed. Lam. blades on edge,
C. lam. pointed, reaching nearly as far as tip of Rost. Lam. hair and
Intlam. hair serrated spines directed forward. Psstg. org. serrated
spines directed transversly outward; coxa 3 bears a strong
serrated spine directed outward parallel to Psstg. org. Nph.
Translucent, white or grey, Rost. and legs pink, interior organs show through
skin, giving color. A few ridges on Cephth. Psstg. dorsal, near together.
Psstg. org. long, filiform, recurved. Abd. ovate; 2 papillae on ant. margin
of Abd. between Psstg. 2 larger papillae near centre of post, margin, from
each springs a hair as long as the body. Dors, of Abd. polished but dotted
with minute punctures. - - Im. L. ca. 650 |u.
6 a. M". bipilis var. quadridentata Haller 1882 N. b. var. q., G. Haller in:
Jahresh. Ver. Wiirttemb., v.38 p. 305.
In moss. England, France. Germany, Italy. Netherlands, Switzerland. Sweden,
Siberia. Nowaya Zemlya.
7. N. burrows! Michael 1890 N. Burrowsii, Michael in: P. zool. Soc. London,
p. 418 t. 37 f. 1 4 j 1892 Oppia lucorum, Berlese, A. M. S., fasc. 64 nr. 2.
Im. Darkish yellow-brown, smooth; Psstg. at base of Cephth. Psstg.
org. with thin shortish peduncle and lanceolate or nearly globular head.
Lam. blades on edge not above */ 3 the length of Cephth., much
nearer together anteriorly than posteriorly. C. lam. small but
Notaspis 47
distinct. Trlam. rather more than a mere line. Legs with finely pectinated
hairs. Ung. hetd. Abd. nearly round. 4 longitudinal rows of thick hairs
on Dors, of Abd. and some on post, margin, a thick hair on shoulder and
another on edge of Abd. all imbricated. L. ca, 750 ju.
Algeria, Italy, Canada (Lake "Winnipeg).
8. N. conformis (Berl.) 1895 Oppia c., Berlese, A. M. S., fasc. 77 nr. 7.
Im. Chestnut-brown, highly polished. Lam. mere lines, scarcely
noticeable. No C. lam. nor Trlam. Intlam. hair directed backward.
Psstg. org. clavate, setose, curving slightly forward. No markings on
Cephth. Abd. long, slightly sinuous and, advancing considerably in centre
anteriorly, pointed posteriorly. No markings on Dors, of Abd. a row of
fine hairs round margin, another a little within. L. ca. 680 |u.
In moss. Italy.
9. N. glabra Michael 1890 N. g., Michael in : P. zool. Soc. London, p. 419.
Im. Light yellow-brown, polished. Psstg. org. with short peduncle
and rough pyriform head. Lam. blades on edge, gradually increasing
in width from posterior to anterior ends. C. lam. very short hardly
visible. Trlam. not so wide as Lam. Ung. hetd. Abd. broad, short;
4 longitudinal rows of minute white hairs on Dors, of Abd. L. ca. 340 |u.
Algeria.
10. N. hepatica (C. L. Koch) 1835 Eremaeus hepaticus, C. L. Koch, C. M. A.,
fasc. 3 t. 23 ! 1884 E. h., Berlese, A. M. S., fasc. 15 nr. 9.
Im. Like N. oblonga but smooth, not punctured, and Dors, of Abd.
hairless. L. ca. 600 |u.
In moss. Germany, Italy.
11. N. licnophora Michael 1882 N. licnophorus, Michael in: J. R,. micr. Soc.,
ser. 2 .2 p. 10 t. 2 f. 7 8 j 1888 N. I, Michael, Brit. Orib., v.2 p. 379 t. 31 f. 6 8.
Im. Light yellow-brown, smooth. Lam. strong bars, not straight,
joined by a transverse bar near ant. margin of Abd. Trlam. very strong
going right across Cephth. Psstg. org. with very short. Peduncles and
extremely large, flat, Japanese fan shaped head. Abd. pointed
anteriorly and posteriorly, 2 rows of short spatulate hairs on Dors, of Abd.
and a row round margin. - Nph. Very like Im., but white and without
the Lam. and other ridges on Cephth. - - Im. L. ca. 190 |u.
On dead wood. England.
12. N. lucorum (C. L. Koch) 1841 Zetes L, C. L. Koch, C. M. A., fasc. 31 1. 18
1879 Notaspis I, Michael in: J. R. micr. Soc., v. 2 p. 241 1888 N. L, Michael, Brit.
Orib., v.2 p. 370 t. 29 f. 11 16 ', 1895 Oppia Berlesii, Leonard! in: Atti Soc. Veneto-
Trent,, ser. 2 v.2 p. 9 | 1895 0. B., Berlese, A. M. S., fasc. 77 nr. 4 || non 1892 0. lucorum,
Berlese, A. M. S., fasc. 64 nr. 2.
Im. Polished, finely punctured, punctures ca. 8 |u apart ; Rost. slightly
trifid. Psstg. org. short with obtusely pyriform head. Lam. hair and Intlam.
hair slightly serrated. Lam. very short, slightly raised, nearly straight blades.
Ung. almost homd. Abd. elongated, oval; ant. margin slightly sinuated and
bordered by a raised line except in middle; a strong, straight spine on
edge of Dors, of Abd. between legs 2 and 3. 4 rows of longish fine hairs
on Dors, of Abd. and an extra pair near post, margin. A very variable
48 Notaspis
species. - - Nph. Semi-transparent, white, internal organs show trough inte-
gument giving bright color, red yellow and green. Rost. and legs rosy
brown. Polished, a few dots and vermiform markings. Form, Psstg. org.
Lam. hair and Intlam. hair and hairs on Dors, of Abd. similar to those of
Im. Abd. sharply divided from Cephth. Spine on edge replaced by a hair.
- Im. L. ca. 670 |u.
Generally distributed, abundant on lichen. England, Germany, Italy.
13. N. microptera (Berl.) 1885 Oppia m., Berlese, A. M. S., fasc. 20 nr. 9.
Im. Yellow-brown, polished. Cephth. long, without markings. Rost.
pointed. Lam. narrow blades on edge, widest anteriorly, very long. No
C. lam. nor Trlam. Psstg. close to ant. margin of Abd. Psstg. org.
with long thin peduncle and fusiform, slightly recurved head
with terminal hair. Abd. as broad as long, widest behind leg 4 ; Ant. margin
slightly convex; post, margin round. Dors, of Abd. hairless. L. 1 mm
or larger.
In moss. Italy.
14. N. oblonga (C. L. Koch) 1835 Eremaeus oblongus, C. L. Koch, C. M. A.,
fasc. 3 t. 24 | 1855 E. o., Nicolet in: Arch. Mus. Paris, v.7 p. 451 t. 10 f. 1 | 1882 E. o. r
Haller in: Jahresh. Ver. Wiirttemb., v. 38 p. 306 j 1884 E. o., Berlese, A. M. S., fasc. 15
nr. 10 ; 1888 Notaspis oblonga, Michael, Brit. Orib., v.2 p. 375 t. 30 f. 6 11 1841
Murcia obsoleta, C. L. Koch, C. M. A., fasc. 31 f. 23 (Nph.).
Im. Dull red or yellow brown, smooth but not polished. Cephth.
constricted where it joins Abd. Lam. short nearly parallel bars
commencing some distance in front of Psstg. Trlam. a bar not
quite joined to Lam. Psstg. org. longish with slender peduncle and fusiform
head, both rough. Intlam. hair much shorter than Psst. org. Ung. very
hetd. Abd. oval, very variable in form. Dors, of Abd. punctured, no other
markings; 4 rows of curved hairs on same, and some round post, margin.
- Nph. Brown dark for a Nph., rough, Cephth. very like that of Im.
Abd. shield-shaped, a lat. border transeversely striated, remainder granulated.
Cast skins of Dors, of Abd. earned excentrically, the earliest furthest back, a
row of short curved hairs on ant. margin of Abd. - - Im. L. ca. 600 |u.
Algeria, England, France, Germany, Italy. Abundant.
15. N. pilosa (C. L. Koch) 1841 Zetes pilosus, C. L. Koch, C. M. A., fasc. 31
t.12 | 1888 Notaspis pilosa, Michael. Brit. Orib., v.2 p. 370 t.29 f.11,16 | 1896 Oppia p.,
Berlese, A. M. S., Crypt. II p. 57 j 1842 Zetes pilosulus, C.L.Koch. Ubers. Arach., v.3
p. 101.
Im. Red-brown; polished. Lam. short, slightly raised, nearly straight
ridges. Lam. hair and Intlam. hair long, serrated. No Trlam. Psstg. org.
with longish filiform peduncle and rough clavate head. Abd. short pyri-
form. A spike stands outward horizontally between legs 3 and 4. A ring
of very long setiform hairs round Dors, of Abd. and 2 rows of similar
hairs nearer centre. L. ca. 450 \JL.
On lichen. England. Germany.
16. N. serrata Michael 1885 N. s., Michael in: J. E. micr. Soc., ser. 2 v. 5
p. 389 t. 7 f. 2 | 1888 N. s., Michael. Brit. Orib., v.2 p. 366 t. 29 f. 110 j 1896 Eremaeus
serratus, Berlese, A. M. S., Crypt. II p. 36.
Im. Dull dark brown, coarsely pitted, and cuticle covered
with short villous processes, which usually retain fragments of lichen etc.
Notaspis 49
Pyriform, post, margin very round. No Lam. nor Trlam. yet Lam. hairs
persist and are serrated. Psstg. dorsal near together. Psstg. org. medium
length, with short peduncle and almost globular head. Intlam. hair long,
flexible, pectinated. Some of the hairs on legs pectinated. Dors, of Abd.
narrow anteriorly, broad posteriorly and there deflexed on to ventral surface.
A circular row of veiy long, flexible, serrated hairs round Dors, of Abd.
and a few similar hairs nearer centre. - - Nph. Usually appears like a heap
of small pieces of lichen etc. with a forest of long serrated hairs arising
from it ; when clean, pale fawn, leathery, almost oblong. Rost. narrow ; hind
part of Cephth. broad. The 2 rostral hairs, cross; these and Intlam. hair
and some hairs on legs serrated. Abd. flattish, parallel-sided. Notogastral
skins carried flat and concentrically, but shrivelled, each skin has a ring of
hairs like those on Dors, of Abd. Im. L. ca. 560 jn.
On lichen. England.
Subdiv. lib. Sp. 17 26. Dors, of Abd. without projecting ant. corners;
Ung. mond.
17. N. clavipectinata Michael 1885 ^V. c., Michael in: J. R, micr. Soc., ser. 2
r. 5 p. 392 t. 7 f. 6 ; 1888 N. c., Michael. Brit. Orib., v.2 p. 387 t. 32 f. 7 11 | 1896
Dameosoma clavipectinatum, Berlese, A. M. S., Crypt. II p. 54.
Im. Lightish yellow-brown, polished. No Lam. or Trlam. Psstg.
org. with long thin peduncle and short flat pyriform head;
bearing radiating bristles. Abd. elliptical; 2 rows of fine hairs on Dors,
of Abd. and a row round margin. - - Nph. Much like Im., but nearly white ;
but Psstg. org. without bristles of Im., but rough. Lam. hair and Intlam.
hair bipectinated. - - Im. L. ca. 330 ju.
England.
18. N. juncta Michael 1885 N.j., Michael in: J. R. micr. Soc., ser. 2 v.5 p. 390
t. 7 f. 3 i 1888 N.j., Michael, Brit, Orib., v.2 p. 365 t. 28 f. 9 12 j 1896 Oribatulaj.,
Berlese, 'A. M. S., Crypt. II p. 59.
Im. Light yellow-brown : polished, without markings. Short pyriform.
Lam. thin blades on edge, widely separated posteriorly, but approaching
so close anteriorly as to touch in centre of Cephth. slightly undu-
lated. C. lam. very long. Lam. hair terminal. Psstg. org. with short
peduncle and longish fusiform head. Femora broad, thin, with large blades.
Tibiae much enlarged at distal end. Ung. mond. Abd. a short ellipse,
truncated anteriorly. Antero-lateral corners of Dors, of Abd. slightly pro-
jecting, not as much as those of N. exilis, N. tibialis etc. L. ca. 220 n.
In moss on trees. England.
19. N. lacustris Michael 1882 N. I, Michael in: J. R. micr. Soc., ser. 2 v.2
p. 12 t. 2 f. 6 ; 1888 N. I, Michael, Brit. Orib., v.2 p.399 t. 33 f. 1-9; t.SOf. 12 j 1896
Scutovertex (sic !) 1., Berlese. A. 31. S., Crypt, II p. 46.
Im. Dull red-brown, generally partly covered with whitish deposit,
smooth, not polished, punctured, punctures about 7 jn apart. Maxillae divided
into 2 bifid lobes. No true Lam. or Trlam. but 2 pairs of irregular ridges one
of which may represent Lam. No other markings on Cephth. Psstg. minute
round holes near base of Cephth. Psstg. org. short, clavate, but almost
always broken off, leaving the part of the peduncle within Psstg. only
Das Tierreich. 3. Lief. : A. D. Michael, Oribatidae. 4
50 Notaspis
Ung. very large, minute scale -like irregular projections on each side of
ung. Abd. an ellipse, slightly truncated posteriorly and bearing 2 pairs of
hairs there, otherwise Dors, of Abd. without hairs or markings except a pale
patch near ant. margin. Nph. Dirty white or pale drab, legs and Host,
pinkish, leathery. Usually no Psstg. Psstg. org., Lam. hair or Intlam. hair
visible, but very minute ones do exist at some times of year. Dors, of Abd.
covered with broad, irregular, transverse wrinkles, each bearing a line of
knobs. Round post, margin are 6 flexible black hairs as long as body and
4 spikes. - - Im. L. ca. 500 |u.
Oudeman's (1896, in: Tijdschr. Ent., v. 39 p. 175) indentifies this species with
Acarus confervae Schrank (1781, Ins. Austr.. nr. 1054), but does not give any reason
for doing so, except habitat. It would not apparently be possible to say from Schrank's
description, which of the many Acari inhabiting confervae he referred to.
Aquatic in fresh water. England.
20. N. lanceolata Michael 1885 N. L, Michael in: J. R. micr. Soc., ser. 2 v.5
p. 394 t. 7 f. 8 1888 N. L, Michael, Brit. Orib., v. 2 p. 398 t. 32 f. 12 | 1896 Dame-
osoma lanceolatum, Berlese, A. M. S., Crypt. II p. 54.
Im. Light yellow-brown, polished, almost fusiform. Lam. short
ridges starting from Psstg. and running inward and forward until
their ends meet only a short distance from ant. margin of Abd.
Psstg. org. long, with slender peduncle and fusiform head terminated by
a bristle. L. ca. 330 |u.
In moss. England.
21. N. longilamellata Michael 1885 N.I., Michael in: J. R, micr. Soc.. ser. 2
0.5 p. 391 t. 7 f.4 ' 1888 N.L, Michael, Brit. Orib., v.2 p. 392 t, 28 f. 13 IB |' 1896
Dameosoma longilamellatum, Berlese, A. M. S., Crypt. II p. 54.
Im. Light yellow-brown, polished; long pyriform. Cephth. very
long, ca. 5 / 12 of whole length. Lam. thick ridges, reaching more
than half length of Rost. post, part converging, ant. part parallel. No
C. lam. nor Trlam. Psstg. org with long filiform peduncle and short fusi-
form head. 2 small curved ridges between proximal ends of Lam. Abd.
truncated anteriorly, rounded posteriorly. 2 rows of longish hairs on Dors,
of Abd. and a row round margin. L. ca. 330 M.
Algeria. England.
22. N. monilipes Michael 1882 N. m., Michael in: J.R. micr. Soc., ser. 2 ;.2 p. 16
t, 2 f. 15 1 1888 N. m., Michael, Brit. Orib., v. 2 p. 381 t.31 f. 15 1 1896 Cymbaeremaeus
m., Berlese, A. M. S., Crypt. II p. 37.
Im. Brown, shade variable, rough, leathery, covers itself with dirt.
Cephth. and Abd. very sharply divided. Cephth. in 3 regions, vizt. a smooth
anterior, a reticulated middle part between Tectp. 1 and a posterior roll with
raised dots. Lam. and Tectp. 1, which are dorsal rough straight bars.
C. lam. long, rough, curving downward; Psstg. org. long, clavate,
rough. Legs shortish, femora rough. Tibiae of legs 1 and 2, especially 1,
disproportionately large, with almost globular heads on short
thin peduncles, an apophysis from upper part of globe supports tactile
hair. Abd. shield-shaped. Antero-lateral angles produced forming points.
3 elevated lumps of which the second is the largest, occupy the whole median
part of Dors, of Abd., lateral border, flat ; whole dors, surface covered with
Notaspis 51
raised dots. A series of short hairs round margin of border. - - Nph.
Light-brown, rough. Cephth. with ridges forming a division into areas.
Psstg. org. long filiform, rough, sinuous. Abd. shield-shaped, cast skins of
Dors, of Abd. earned excentrically, earliest furthest forward. 3 hemispherical
lumps arranged longitudinally on larval skin. 2 projections at hind end bear
long spatulate hairs. Im. L. ca. 340 |u.
On dead wood. England. Rare.
23. N. pectinata Michael 1885 N.p., Michael in: J. R. raicr. Soc., ser. 2 v. 5
p.392 t.7 f.5 I 1888 N.p., Michael, Brit.Orib., v. 2 p. 389 t.32 f.l 6 j ?1892 Belbap.,
Berlese, A. M. S.. fasc. 61 nr. 8 | ? 1896 Dameosoma pectinatum, Berlese, A. M. S.,
Crypt. II p. 54.
Tm. Light yellow-brown, smooth, not quite polished, almost fusiform.
Lam. short ridges; no Trlam. a second pair of ridges at base of Eost. Psstg.
org. very long, doubly curved, directed backward, with a few
widely separated pectinations on one side of distal portion. Intlam.
hair, hairs on Dors, of Abd., and some on leg all pectinated. - - Nph. Almost
colorless, smooth not polished. Psstg. org. and Intlam. hair long, upright,
serrated, curving sharply forward at distal ends. Kost. hair similar but
smaller. Abd. almost oblong, 4 large papillae along each side 4 smaller
on post, margin, and 2 rows of about 4 on dors, surface, each papilla bears
a hair similar in character to Intlam. hair, but of various lengths. Im.
L. ca. 420 |u.
In dead leaves. England, ? Italy.
24. N. quadricarinata Michael 1885 N. q., Michael in: J. R. micr. Soc., ser. 2
v.~> p. 393 t.7 | 1888 N. q., Michael, Brit.Orib., v.2 p. 385 t. 31 f. 13 15 | 1896
Dameosoma quadricarinatum, Berlese, A. M. S., Crypt. II p. 54.
Im. Light yellow-brown; polished, short pyriform. Cephth. large.
Lam. and Trlam. strong ridges. No C. Lam. Psstg. org. slightly recurved
with fusiform head on short peduncles. Abd. short, arched, truncated ante-
riorly. 4 short longitudinal ridges on ant. part of Dors, of Abd.
4 rows of short hairs on Dors, of Abd. L. ca. 200 |u.
In moss. England.
25 N. splendens (C. L. Koch) 1841 Oppia s., C. L. Koch, C. M. A., fasc. 32
p. 6 | 1879 Damaeus s., Michael in: J. R. micr. Soc., v. 2 p. 247 j 1888 Notaspis s.,
Michael, Brit.Orib., v.2 p. 393 t. 33 f. 10 15 1895 Dameosoma s., Berlese, A. M. S.,
fasc. 77 nr. 10.
Im. Light yellow - brown ; polished. Lam. zig-zag ridges usually
springing from edge of Abd. so that a short part is behind Psstg. distal
ends of Lam. widely separated. No Lam. hair aor Trlam. 2 short
ridges nearer median line than Lam. Psstg. org. with longish slender
peduncle and rough fusiform head. Abd. oval, without markings; 4 rows
of small hairs on dors, surface. Nph. Almost white; no ridges or,
markings on Cephth. Hairs on Abd. very long, slightly serrated ; otherwise
Nph. like Im. Im. L. ca. 310 |u.
A variable species. Probably Koch's Oppia splendens included many species.
Algeria, England, Germany, Italy.
26. N. trigona Michael 1888 N. t., Michael, Brit. Orib., v. 2 p. 396 t. 28 f. 16
1896 Dameosoma trigonum, Berlese, A. M. S., Crypt. II p. 54.
4*
52
Xotaspis, Damaeinae
Im. Very like N. splendens; light yellow-brown, polished. Kost.
truncated, very angular with a median and 2 lat. points. Lam. short
bars commencing a short distance in front of Psstg. 2 very short
bars nearer median line starting from ant. margin of Abd. and joined at
their anterior ends by a strong transverse bar with 2 points directed for-
ward. A small pine apple-shaped lump on Dorso-vertex in front of trans-
verse bar. No Trlam. Psstg. org. with slender peduncle and fusiform head.
Abd. almost elliptical, slightly pointed posteriorly. Dors, of Abd. without
markings, 2 longitudinal rows of very fine hairs on same, and a few round
margin. L. ca. 240 ju.
England.
N. distincta (Karp.) 1893 Oribata d., Karpelles in: Math, naturw. Ber. Ungarn,
v. 11 p. 83.
Im. A Notaspis, probably N. bipilis. but described as having 2 longitudinal
rows of 4 short pectinated hairs on Dors, 'of Abd. Size?
Hungary.
N. longiuscula (C. L. Koch) 1841 Zetes I, C. L. Koch, C. M. A., fasc. 31 1. 19.
Im. Very small, yellow-brown, narrow, highly polished. Psstg. org. with long
thin peduncle and suddenly clavate head. Abd. a very long ellipse, narrow. Leg 4
not reaching post, margin.
In wet meadows. Germany.
E. Subfam. Damaeinae
Fig. 12. Damaeus geniculatus (' 20 /i).
1885 Subfam. Nothridae (part.), Ber-
lese in: Bull. Soc. ent. Ital., v. 17 p. 126 |
1891 N., Canestrini in : Atti 1st. Veneto,
ser. 7 v. 2 p. 717 j 1892 Nothrinae (part,),
Trouessart in : Rev. Sci. nat. Ouest, v. 2
p. 38 ; 1896 Damoeidae (sic) (part.), Berlese,
A.M.'S., Crypt, II p. 33.
Without Ptm. Cephth. anchy-
losed to Abd. Ventr. plate anchy-
losed to dors. Mdb. chelate. Legs
12 near together, 3 4 near together.
Leg 4 crawling organ. Without La-
mellae. Legs thin, much longer than
body.
2 genera, 22 good and 4 doubtful
species.
Key to the genera.
Cephth. and Abd. coalescing in the median
line ... 1. Gen. Amerus . p. 53
Cephth. and Abd. clearly separated
2. Gen. Damaeus . p. 53
Amerus, Damaeus
53
1. Gen. Amerus fieri
1883 Amerus?, Berlese, A. M. S., fasc. 3 nr. 5 ; 1896 A., Berlese, A. M. S.,
i'asc. 79 nr. 7.
Cephth. and Abd. coalescing in the median line without any demar-
cation between them.
1 species.
1. A. troisi (Berl.) 1883 Belba Troisii, Berlese, A. M. S., fasc. 3 nr. 5 | 1896
Amerus T., Berlese, A. M. S., Crypt. II p. 40.
Im. Very dark brown; polished. 2 pairs of Rost. hairs long and
flexible; Psstg. near together; Psstg. org. very long, setiform. Abd. long,
rounded posteriorly, with 2 sparse' longitudinal rows of very long flexible
hairs on dors, surface. Chitinous projections at side of body between legs
1 and 2, 2 and 3, 3 and 4. L. 1,1 mm.
Algeria, Italy.
2. Gen. Damaeus C.L.Koch
V1796 Acarus (non Acarus, Typ. : A. coleoptratus L.. Latreille 1795!), Latreille,
Precis Caract. Ins., p. 184 (Typ. : A. geniculatus L.) j 1835 Damaeus, C. L. Koch, C. M. A.,
fasc. 3 1842 D., C. L. Koch, Ubers. Arach., v.3 p. 106 1855 D., Mcolet in: Arch.
Mus. Paris, v.7 p. 459 j 1888 D., Michael, Brit. Orib., v.2 p. 404 | 1891 D., Canestrini
in: Atti 1st. Veneto, ser. 7 v.2 p. 580 ! 1892 D., Trouessart in: Rev. Sci. nat. Quest,
v. 2 p. 48* | 1877 Belba (non Heyden 1826!), Canestrini & Fanzago in: Atti 1st. Veneto,
ser. 5 v. 4 p. 77 | 1895 B., Banks in: Tr. Amer. ent. Soc., v. 22 p. 12 | 1896 B., Oudemans
in: Tijdschr. Ent., v. 39 p. 55 j 1885 Belba -f- Damaeus, Canestrini, Prosp. Acarof., v. 1
p. 8, 9 1896 B. + D. + Dameosoma (part.), Berlese, A. M. S., Crypt. II p. 38, 39, 54.
Cephth. and Abd. clearly separated.
21 good and 4 doubtful species.
Key to the good species.
J Ung. mond. (Div. I. Gen. Belba Berl.) 2.
I Ung. trid. (Div. II. Gen. Damaeus Berl.) 7.
J Abd. oval in outline 3.
| Abd. round in outline 4.
/ Abd. rounded posteriorly, Dors, of Abd. covered
with fine short hairs distant from each other
3 ' about their own length. L. 1 mm . . .
I Abd. pointed posteriorly, hairs on Dors, of Abd.
long and few. L. 510 |n 9. D. rfltens
i Psstg. org. long and flagelliform (i. e. setiform
4 J and flexible) 5.
' Psstg. org. not flagelliform 9.
Joints of legs mostly moniliform; i. e. with
very short peduncle and almost globular
head; Im. usually carrying cast skins of Dors,
of Abd. 6.
Joints of legs not moniliform, Im. not carrying
east skins 7.
8. D. nigro-femoratus
p. 57
p. 57
54
Damaeus
10
11
12
13
14
15
16
A strong curved point projecting from Abd.
between legs 1 and 2 and a similar point
between legs 3 and 4; hairs on Dors, of Abd.
shortish flexible, not projecting beyond its
margin except a little at post, margin. -
L. 360 |n
No curved points between legs 2 and 3 or 3 and 4 ;
hairs on Dors, of Abd. stiff, projecting half
their length beyond its margin. L. 550 |u
Leg 4 4-times as long as whole body. L. 520 n
Leg 4 not above twice the length of body 8.
Inner hair on Femur 4 much longer than the
Femur; hairs on Genual 4 three times as
long as the Genual. L. 650 |i ......
Inner hair on Femur 4 about 1 /% of the length
of the Femur; hairs on Genual 4 about
J /s the length of the Genual. L. 450 |u .
Hairs on Dors, of Abd. thick, straight or slightly
curved 10.
Hairs on Dors, of Abd. sharply hooked
L. 850 |u
Dors, of Abd. arched (hemispherical) 11.
Dors, of Abd. conical, usually plastered with
mud. L. 660 \i
Ventr. plate small, dors, greatly reflexed
posteriorly; covering about */ 3 of ventr.
surface. L. 650 (a
Ventr. plate large; dors, scarcely reflexed. 12.
Hairs on legs spatulate, Im. usually covered
with white powder. L. 500 |u
Hairs on legs not spatulate ; Im. without white
powder 13.
Psstg. org. setiform serrated. L. 650 |u . .
Psstg. org. not serrated 14.
Psstg. org. rod-like; a projection from Cephth.
between legs 1 and 2 15.
Psstg. org. clavate; no projection between
legs 1 and 2 16.
Femora with thin proximal part, distal suddenly
enlarged; projection between legs 1 and 2
pointed anteriorly. L. 1,05 mm ....
Femora gradually clavate, projection between
legs 1 and 2 blunt anteriorly. L. 1,5 mm .
Psstg. org. short with thin peduncle and suddenly
enlarged elongated head (like a pileus);
Intlam. hair short, rough, rod -like.
L. 670 n
Psstg. org. long, with thin peduncle and gradually
and slightly clavate head, pointed anteriorly
almost fusiform; Intlam. hair long setilbrm.
L. 690 n
16. D. verticillipes .
7. D. globipes . .
11. D. phalangoides
5. D. flagellifer . .
2. D. berlesei
p. 59
p. 57
]>. 58
p. 56
p. 55
1. D. auritus p. 55
10. D. patelloid.es .
13. D. sufflexus . .
12. D. pulverulentus
14. D. teeticola . .
j). 58
j). 58
p. 58
p. 58
3. D. clavipes . . .
6. D. geniculatus .
15. D. tenuipes . .
4. D. concolor . .
p. 55
p. 56
p. 59
p. 56
Damaeus
/ Dors, of Abel, convex; cast skins carried 18.
17 < Dors, of Abd. concave or flat, cast skins not
carried 19.
Psstg. org. flagelliform; 4 central hairs on hind
margin much longer than the others; a
square, central, post, projection from each
cast skin bearing similar hairs, cast skins
18 I shield-shaped. L. 650 |u 21. D. setosus p. 60
Psstg. org. thick, rod-like on slightly clavate,
only 4 hairs on hind margin, no central
projection, cast skins marked with radiating
lines 18. D. craterifer . . . . p. 60
i Dors, of Abd. discoidal 20.
19 j Dors, of Abd. truncated anteriorly, pointed
posteriorly. L. 1 mm 19. D. femoratus . . . . p. 60
Psstg. org. flagelliform, legs with broad rough
borders; a large curved projection between
legs 1 and 2. L. 1 mm 20. D. ornatissimus . . p. 60
Psstg. org. with long thin peduncle and clavate
spinous head; no borders to legs; no
projection between legs 1 and 2. L. 500 \\ 17. D. bicostatus . ... p. 59
Div. I. Sp. 116. Ung. mond.
1. D. auritus C. L. Koch 1835 D. a., C. L. Koch, C. M. A., fasc. 2 t. 11 j 1888
D.a., Michael, Brit. Orib., v. 2 p. 435 t. 40 j 1844 Oribata aurita, Gervais in: Walcke-
naer, Ins. Apt., v. 3 p. 257 j ?1885 Belbaa., Canestrini, Prosp. Acarof.. v. 1 p. 39 || non
1855 Damaeus auritus, Nicolet in: Arch. Mus. Paris, v.7 p. 462 t. 8 f. 3.
Im. Very like D. geniculatus, but smaller and hairs on Dors, of Abd.
short, fine, very sharply curved, almost hooked. Portions of the cast skins
of Dors, of Abd. usually earned. - - Nph. Like that of D. geniculatus but
smaller. - - Im. L. ca. 850 ju.
England, Germany.
2. D. berlesei Michael*) 1887 Belba nodipes (err., non Damaeus nodipes
C.L.Koch 1840!), Berlese, A. M.S., fasc. 43 nr.2 | 1896 Dameosoman., Berlese, A. M.S.,
Crypt. II p. 54.
Im. Cephth. without ridges or markings. Psstg. much raised;
Psstg. org. very long, whip-like (as in D. globipes). Abd. hemispherical,
polished; bearing 2 longitudinal rows of short curved hairs. Legs very
long. Femur of leg 4 considerably passing post, margin of Abd. joints
of legs with long thin peduncles and suddenly clavate heads.
L. ca. 450 |u.
In moss. Italy.
3. D. clavipes (Herm.) 1804 Notaspis c., Hermann, Mem. apt., p. 88 t. 4 f. 7
1888 Damaeus c., Michael, Brit. Orib., v. 2 p. 423 t. 38 1835 Oppia glaucina, C. L.
Koch, C.M. A., fasc. 3 t,9 (Nph.) | 1840 Damaeus nodipes, C. L. Koch, C. M. A., fasc. 30
*) Berlese identifies this species, I think erroneously, with Damaeus nodipes.
C. L. Koch (which is, in my opinion, a synonym of D. clavipes); the species therefore
appears to be unnamed and I name it in honor of Professor Berlese. A. D. Michael.
56 Damaeus
t. 6 ; 1855 D. auritus, Nicolet in: Arch. Mus. Paris, v.7 p. 462 t. 8 f . 3 |j non 1887
Belba nodipes, Berlese. A. 31. S.. fasc. 43 nr. 2.
Im. Dark brown, lighter when lately emerged; smooth not polished,
often powdery, often carries dirt. Cephth. with 2 transverse furrows, between
them are 2 short longitudinal ridges and rectangular projections bearing leg 1.
Psstg. very projecting on mamilliform elevations. Psstg. org. filiform not so
long as those of D. geniculatus, 4 short ridges behind Psstg. between
legs 1 and 2 is a chitinous projection longer narrower and more
pointed anteriorly than in D. geniculatus. Legs very long and
thin, joints mostly with long thin peduncle and suddenly enlarged head.
Abd. almost globular. About 14 black curved hairs on Dors, of Abd. and
4 on post, margin. Nph. Light yellow or reddish, legs pink, smooth ;
4 rows of large papillae on Dors, of Abd.; in young specimens and Nph.;
often only the post, pair persist in fully-grown Nphs. ; each papilla bears a
long, black, slightly serrated hair. Carries dirt and debris. - - Im. L. ca.
1,05 mm, L. of leg 4 ca. 1,8 mm.
All early writers except Hermann, and also Canestrini & Fanzago (1877. in: Atti
1st. Veneto, ser. 5 v. 4) confuse this species with D. geniculatus.
Algeria. England. France, Germany, Italy, Sweden, Switzerland.
4. D. concolor C. L. Koch 1840 D. c., G. L. Koch, C. M. A., fasc. 30 t, 6 | 1887
Belba c., Berlese, A. M.S., fasc. 43 nr. 1 1896)ameosoma c.. Berlese, A.M. S., Crypt. II
p. 54 I ?1882 Belba denticulata, G. &R. Canestrini in: Atti 1st. Veneto. ser. 5 v. 8 p. 914
t. 8 f.4 | 1885 B. d., Canestrini, Prosp. Acarof.. v.l p. 40. t
Im. Chestnut-brown Cephth. and Abd. without markings. Psstg.
org. long, slightly clavate, pointed at distal ends. Abd. globose.
Dors, of Abd. and post, margin with long flexible hairs. L. ca. 690 ju.
Germany. Italy.
5. D. flagellifer Michael 1890 D. f., Michael in: P. zool. Soc. London, p. 422
t. 38 f. 2.
Im. Dark brown, smooth, not polished, very finely punctured. Cephth.
slightly spotted, but without true markings. Psstg. org. long, setiform,
flexible, curled or undulated near distal end. Legs with joints clavate,
Femora suddenly so, leg 4 ca. 850 |u. One hair on Femur and Genual
of this leg like Psstg. org. and greatly larger than any other hair
on creature. L. ca. 650 |u.
Algeria.
6. D. geniculatus (L.), C.L.Koch ?1758 Acarus g., Linne, Syst. Nat., ed. 10
p. 617 | ?1761 A. g., Linne, Fauna Suec., ed. 2 nr. 1977 j ?1796 A. g., Latreille, Precis
Caract. Ins., p. 184 | ?1803 A. g., Schrank. Fauna Boica, v. 3 p. 208 nr. 2666 j ?1806
Oribata geniculata, Latreille, Gen. Crust. Ins., v. 1 p. 149 | 1835 Damaeus geniculatus,
C.L.Koch, C. M. A., fasc. 3 t, 13 1855 D. g., Nicolet in: Arch. Mus Paris, v.7 p. 460
t. 8 f. 1 I 1888 D. g., Michael, Brit. Orib., v.2 p. 428 t.39 | 1844 Belba geniculata,
Gervais in : Walckenaer. Ins. Apt., v. 3 p. 256 1877 B. g., Canestrini & Fanzago in :
Atti 1st. Veneto, ser. 5 v.4 p. 101 1887 B. g., Berlese, A. M. S., fasc. 36 nr. 4 j ?1835
Damaeus torvus, C.L.Koch, C. M. A., fasc. 3 f. 14 (Nph.) | ?1844 D. onustus, C. L.
Koch. C. M. A., fasc. 38 t. 7.
Im. Dark brown or black, dull, slightly rough. Cephth. with 2 trans-
verse furrows and 2 triangular depressions bordered laterally by ridges;
Damaeus 57
a chitinous projection carries leg 1. Psstg. on mamilliform projection, trumpet-
mouthed, ridged inside. Psstg. org. long, filiform; a projection between
legs 1 and 2 blunt-pointed anteriorly, rounded posteriorly. Legs thick for
genus; coxae of 2 hind pairs with projecting blades on post, edge; Femora
thicken gradually. Abd. sligthly pyriform; about 18 large black hairs
on dorsal surface. - - Nph. Pale yellowish, leathery; generally concealed
by dirt, legs pink. Psstg. org. S-shaped. Abd. usually concealed by mud,
but oval with 2 points posteriorly, bearing black hairs, 2 rows of similar
hairs on dors, surface. - - Im. L. ca. 1,5 mm, L. of leg 4 ca. 1,75 mm.
All the earlier writers and Canestrini & Fanzago confuse this species with
D. clavipes.
Belgium. England, France, Germany, Italy, Netherlands, Sweden, Switzerland,
Port Clarence (Yega-Exp.).
7. D. globipes (Can. & Berl.) 1884 Belba g., Canestrini & Berlese in: Atti
Soc. Veneto-Trent., v. 9 fasc.l p. 177 t.5 f.3 5 | 1886 B. g., Berlese, A. M.S., fasc. 33
nr. 4.
Im. Chestnut-brown, smooth. Psstg. org. very long, much longer
than Cephth. flagelliform; projections which cany legs 1 and 2 only
moderately developed. Abd. globular, with a curved row of 5 long, thick,
curved hairs near each lat. margin, and 2 pairs on post, margin. Femora
with thin peduncle and suddenly clavate head, other joints globose
(moniliform), each joint with a whorl of very strong, almost straight hairs.
Often carries cast skins on Dors, of Abd. excentrically, earliest posteriorly.
L. 550 |u.
In moss etc. Algeria, Italy.
8. D. nigro-femoratus (L. Koch) 1879 Nothrus n., L. Koch in: Svenska Ak.
Handl., r. 16 p. 112 t. 3 f. 23.
Im. Body Coxae and Femora black; remainder of legs yellow; smooth
not polished, Cephth. and Abd. covered with short fine hairs distant
from each other about their own length. Psstg. org. short, very clavate.
Abd. oval, widest and rounded posteriorly. L. 1 mm.
Yugor-Straits, North Russia.
9. D. nitens (C. L. Koch) 1835 Oppia n., C.L.Koch, C.M.A., fasc. 3 t.10 | 1888
Damaeus n., Michael, Brit. Orib., v. 2 p. 409 t. 34 f. 1 8 |j non 1896 Dameosoma
concolor, Berlese, A. M. S., Crypt. II p. 38 (C. L. Koch, C. M. A., fasc. 30 t. 6, although
said to be so by Berlese).
Im. Yellow-brown to chestnut-brown polished, oval. Psstg. org. with
long slender peduncle and slightly lanceolate head. Legs not so
long nor the joints so clavate as usual in genus. Abd. oval, rounded
anteriorly almost pointed posteriorly, no markings. 2 rows and 2 inner
pairs of longish filiform white hairs on Dors, of Abd. and 1 or 2 shorter
pairs on post, margin, all slightly imbricated if magnified 500 diam. - - Nph.
White, legs and Kost. pinkish; translucent. Psstg. org. long, rough, gradually
thickened. Intlam. hair slightly serrated ; Abd. truncated anteriorly, rounded
posteriorly; hairs on Dors, of Abd. long, serrated. - - Im. L. ca. 510 ju.
In moss and dead wood. England, Germany. Sweden.
58 Damaeus
10. D. patelloides 3Iichael 1890 D. p., Michael in: P. zool. Soc. London,
p. 420 t. 38 f. 3.
Im. Dark brown, smooth, not polished. Cephth. without sculpture
but a vague ridge between the 2 Psstg., which are raised, cup-chaped, but
outer side of cup produced to a point. Psstg. org. long, rod-like, slightly
tapering. Legs with gradually clavate joints; leg 4 ca. 680 ju long. Abd.
almost conical, leans a little backward; 2 rows of short, thick, rough
hairs on Dors, of Abd. forming almost a ring. L. ca. 660 fi.
Algeria.
11. D. phalangoides Michael 1890 D. p., Michael in: P. zool. Soc. London,
p. 421 t. 38 f. 1.
Im. Yellow-brown, smooth, not polished. Rost. narrow, hind part of
Cephth. broad. Intlam. hair and Psstg. org. long, fine, setiform,
flexible. Legs extraordinarily long and thin; leg 4 1,5 mm. Ung.
small. Abd. a short ellipse without markings; dors, with 2 longitudinal
rows of hairs of which the anterior are black, short and curved, the
posterior flexible and lighter. L. ca. 520 |u.
Algeria.
12. D. pulverulentus (C. L. Koch) 1840 Nothrus p., C. L. Koch, C. M. A.,
fasc. 29 f . 3 j 1855 Damaeus papillipes, Nicoletin: Arch. Mus. Paris, v.7 p. 463 t. 8 f. 4.
Im. Bright yellow or red, above, violet at sides. Legs red. Rost.
and sides of Cephth. rough, central and hind part of Cephth. covered by
a trapeze-shaped plate with raised edges and a transverse bar between the
2 Psstg., which are wide and oval. Psstg. org. very long clavate. Abd.
slightly elliptical, highly polished; a circle of long setiform hairs on Dors.
No spines on ant. margin of Abd. Hairs on legs spatulate. Creature
generally covered with white powder. L. 500 ju.
Berlese (1896, A. M. S., Crypt, p. 38) says this species is the same as I), verti-
cillipes Nic.
In woods. France. Germany. Uncommon.
13. D. sufflexus Michael 1885 D. s., Michael in: J. R. micr. Soc.. ser. 2 t: 5
p. 394 t, 7 f. 9 [ 1888 D. s., Michael, Brit. Orib., v. 2 p. 415 t. 34 f. 9, 10 | 1895 Belba
sufflexa, Berlese, A. M. S., fasc. 74 nr. 4.
Im. Dark brown, dull; Cephth. and legs roughish. Cephth. with a
transverse division behind leg 1 ; a median carination and 2 bosses each
side. Psstg. projecting. Psstg. org. long rod-like. Abd. globose finely
punctured. Dors, plate of Abd. turned over posteriorly on to ventr.
surface. Ventr. plate of Abd. small. A row of spines round hinder
part of Dors, of Abd. L. ca. 650 ju.
England, Italy.
14. D. tecticola Michael 1888 D. t, Michael, Brit. Orib., v.2 p. 416 t. 35
| 1895 Bella t., Berlese, A. M. S., fasc. 77 nr. 9 | 1893 B.mirabilis, Karpelles in: Math,
naturw. Ber. Ungarn, v. 11 p. 85.
Im. Brown with a purplish shade, often dusted with white powder.
Cephth. dull, roughish, no sculpturing but elevated opposite insertions of
legs, depressed between ; a small projection to which leg 1 is articulated, a
larger projection with a strong curved point anteriorly between legs
1 and 2, a smaller curved point behind leg 2. Psstg. org. serrated rod-
Damaens 59
like. Legs slender with clavate joints. Many hairs on legs imbricated.
Abd. almost globular, 2 rows of curved, black, imbricated hairs on Dors.
ofAbd., and a pair on post, margin. -- Nph. Light flaxen, polished. Psstg.
org. long, curved, setiform, often have particles of matter attached. Abd.
oval with bifid point posteriorly from which spring 2 long hairs, a pair of
long, black hairs on ant. margin and 2 rows on Dors, of Abd. Cast shrunk
skins of Dors, of Abd. carried concentrically on hind part of Abd. Im.
L. ca. 650 )u.
In thatch and moss. England. Italy.
15. D. tenuipes Michael 1885 D. t., Michael in: J. R. micr. Soc., ser. 2 v. 5
p. 395 t. 7 f. 10 | 1888 D. t., Michael. Brit. Orib.. v.2 p. 420 t. 36 | 1896 Belba t.,
Berlese, A. M. S., Crypt. II p. 39.
Im. Brown with a purplish shade, often dusted with white powder.
Cephth. very like D. tecticola, but projection between legs 1 and 2 rounded
and without anterior point. Psstg. org. short with rod-like peduncle
and rough bluntly conical head. Legs very long and slender, pair 4
twice as long as body. Abd. almost globular. Hairs on Dors, of Abd.
black, almost straight, deminishing in length from before backward. - - Nph.
Like that of D. tecticola but legs longer. Psstg. org. like those of Im. and
cast skins of Dors, of Abd. earned not flat but in a pyramid, supported by
an internal column. - - Im. L. ca. 670 |u.
In thatch. England.
16. D. verticillipes Nic. ? 1840 Nothrus pollinosus, C. L. Koch, C. M. A.,
fuse. 29 1. 12 (Nph. aut Im.) | 1855 Damaeus verticillipes, Nicolet in: Arch. Mus. Paris,
v.7 p. 462 t. 8 f. 2 | ?1888 D.v., Michael, Brit. Orib., v. 2 p. 412 t. 37.
Im. Usually covered with white or grey powder; when cleaned dark
red-brown, smooth not polished. Cephth. with large, lat., trifid projection
bearing leg 2. Psstg. near together, trumpet-shaped. Psstg. org. long
setiform with undulated distal ends, often matted up or broken. Intlam.
hair setiform, curved ; a shorter more curved hair on outside of each Psstg.
Legs short for genus; joints with thin proximal ends and sudden, almost
globular, enlargements; each joint with a whorl of curved hairs. Abd.
almost globular without markings; projecting points between legs. 2 hairs
standing forward from ant. margin, 2 rows of hairs on Dors, of Abd. Cast
skins of Dors, of Abd. often carried, earliest posteriorly. - Nph. Very like
Im., but yellowish white, soft, very finely reticulated ; legs a little shorter. Abd.
a little longer than Irn. Hairs on Dors, of Abd. black. - - Im. L. ca. 360 |u.
Berlese (1896, A. M. S., Crypt. II p. 38) identifies this species (as well as D. papillipes
Nic.) with Nothrus pulverulentus C. L. Koch. It is doubtful, if the species of Nicolet
and Michael are identical, if not Nicolet's has not been refound.
England, France, ? Germany.
Div. II. Sp. 17-21. Ung. trid.*)
17. D. bicostatus C, L. Koch 1835 D. b., C. L. Koch, C. M. A., fasc. 2 1. 12
1886 D.b., Berlese, A. M. S., fasc. 33 nr. 5 | 1877 Belba bicostata, Canestrini & Fan-
zago in : Atti 1st. Veneto, ser. 5 v. 4 p. 103.
*) The genus Damaeus is divided by Berlese, only this group being retained
in the genus, but this is not permissible, because Koch's type of the genus is auritus
a mond. species. A.D.Michael.
60 Damaeus
Im. Chestnut-brown, dull roughish, areolated and raised in scattered
hemispherical .dots. 2 pairs of Rost. hairs, the outer pair thick. Psstg.
org. directed backward and outward, with long thin peduncle
and clavate spinous head. Abd. discoidal, dors, plate concave,
but with slight longitudinal convexity along median line. Post, margin
with a small central projection bearing 2 short hairs, which turn sharply
forward. Tactile hair of Tibia 1 borne on an apophysis. L. ca. 500 ja.
Berlese suggests, that this species may be the young form of D. femoratus
C. L. Koch, but it is not. it is adnlt.
In moss. Algeria, Germany, Italy.
18. D. craterifer Haller 1884 D. c., G. Haller in: Arch. Xaturg., v.oQi p. 226
1. 16 f. 14.
Im. Dark brown or black. Cephth. simple, pointed. Psstg. org.
described short and thick, drawn long and thick. Abd. globular, 2 pairs
of fine hairs on post, margin. Haller draws and describes the species as
carrying a cast abdominal and cephalothoracic skin, the former
marked with radiating lines. Coxae enlarged, rest of leg thin. Tactile hair
on Tibia 1 mounted on a chitinous projection. Size ,.mittelgross".
Southern France.
19. D. femoratus C. L. Koch 1840 D. f., C. L. Koch. C. M. A., fasc. 30 t. 7 |
1876 Belba Dugesii, Canestrini & Fanzago in: Atti Soc. Veneto-Trent., v. 5 p. 104 j
1883 Damaeus D., Berlese, A. M. S., fasc. 3 nr. 6 j 1885 D. D., Canestrini, Prosp.
Acarof., v. 1 p. 33 1877 Belba gibba, Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5
v. 4 p. 104 (Nph.).
Im. Dark red-brown, smooth; large bifid projections between legs 1
and 2 and between 3 and 4. Psstg. org. long, filiform. Rost. hairs
most hairs on legs and those on border of Abd. clavate. Abd. oval truncated
anteriorly slightly pointed posteriorly; Dors, of Abd. hollow, edge
raised, centre very slightly arched, no hairs on it, only round border.
L. ca. 1 mm.
Algeria, Germany, Italy.
20. D. ornatissimus Berl. 1888 D. o., Berlese in: Bull. Soc. ent. Ital., r.20
p. 217 1. 13 f. 1.
Im. Chestnut-brown. Psstg. org. very long, flagelliform. Intlam.
hair large, slightly plumose. Abd. discoidal, concave with a raised border.
A strong curved, pointed horn-like projection between legs 1 and 2. Legs
with broad rough borders. L. ca. 1 mm.
Under bark of trees. Brazil (Matto-grosso).
21. D. setosus Berl. 1883 D. s., Berlese in: Riv. Ace. Padova. r. 33 p. 51 (
1887 D. s., Berlese, A. M. S., fasc. 43 nr. 3.
Im. Yellow-brown, smooth ; Cephth. without special markings. Psstg.
org. very long, flagelliform. Abd. sub-hemispherical, without mar-
kings or hairs on dors, plate, except a row of shortish hairs curved inward
springing from minute tubercules on post, margin, and 4 very long flexible
hairs from similar tubercles just within centre of post, margin close together
transversely. Abd. however usually hidden by cast skins of Dors, of Abd., which
are flat, shield-shaped, concentric, with square central projection
Damaeus, Nothrinae
posteriorly having 4 tubercles on post, margin, which bear hairs (usually
broken off). Whole creature usually covered with white dust. L. ca. 650 jn.
In moss. Sicilia.
D. bulbipes (Karp.) 1893 Belba b., Karpelles in: Math, naturw. Ber. Ungarn.
v. 11 p. 91 t. 13 f. 2.
Im. Like D. sufflexus, probably it, but some of the hairs on legs said to be
pectinated.
Hungary.
D. carinulatus (Berl.) 1888 Eremaeus c., Berlese in : Bull. Soc. ent. Ital., v. 20
p. 217 t, 13 f. 2,
Im. Brown, very like. D. bicostatus, but Psstg. org. not so much enlarged
at distal end; a few transverse ridges on Cephth. Described from a single specimen.
L. 800 |u.
Under bark of trees. Brazil (Matto grosso).
D. formica Gerv. 1849 D.f., Gervais in: Gay, Hist. Chile, v.-i p. 47.
Im. Somewhat like D. clavipes. Chestnut-brown, joints of legs not so much
incrassated as in D. crassipes. L. ca. 1 mm.
Chile.
D. riparius Me. 1855 D.r., Mcoletin: Arch. Mus. Paris, . 7 p. 461 t. 8 f . 5 |
1887 Belba aurita, Berlese, A. M. S., fasc. 36 nr. 5.
Im. Mcolet and Berlese identify this species with D. auritus C. L. Koch, if so
Mcolet's name fails, but his drawing and Berlese's show 4 concave longitudinal furrows
on Dors, of Abd., ending in points on ant. margin, these are not present in Koch's
drawing nor on the English specimens, and the hairs on Dors, of Abd. are larger and
less curved than those of D. auritus. probably therefore this is a good species, but it
may be a variety of D. auritus. L. ca. 1 mm.
France, Italy.
F. Subfam. Nothrinae
1885 Subfam. Nothridae (part.). Berlese
in: Bull. Soc. ent. Ital.. . 17 p. 126 1891 N.,
Canestrini in: Atti 1st. Veneto, ser. 7 v. 2 p. 579
| 1892 Nothrinae (part.), Trouessart in: Rev.
Sci. nat. Quest, v.2 p. 48* | 1896 Nothridae (part.)
-f- Damaeidae (part.), Berlese, A. M. S., Crypt. II
p. 2435.
Without Ptm. Cephth. anchylosed to
Abd. Ventr. plate anchylosed to dors. Mdb.
chelate, legs 1 and 2 near together, 3 and 4
near together. Leg 4 a crawling organ.
Without Lam., legs short and thick.
6 Genera, 42 good and 7 doubtful species.
Key to the Genera.
{Abd. not segmented 2.
Abd. transversly divided as if segmented
6. Hypochthonius . p. 76
j Dors, of Abd. convex 3.
2 < Dors, of Abd. flat, concave, or undu-
lated o.
Fig. 13. Nothrus biverrucatus
62 Nothrinae, Hermannia
3
!
I
Dors, of Abd. of Im. fully chitinized, not carrying
cast nymphal skins. Ung. mond 4.
Dors, of Abd. of Im. not fully chitinized, but carrying
cast nymphal skins of Dors, of Abd. Ung. trid.
Genital and anal covers quite separate and inserted
in a large ventr plate
2.
1
Neoliodes . . . .
Hermannia .
4
1
f
Genital and anal covers only separated by a rim
common to both; not inserted in a ventr. plate,
but touching the deflexed dors, plate at sides
Tectp. present at leg 2
5.
H
Lohmannia . . .
Cym.baerem.aeus
5
'
No Tectn. for letr 2 .
4
Nothrus .
p. 65
p. 62
p. 75
]>. 66
p. 68
1. Gen. Hermannia Nic.
!
1842 Nothrus (part.), C.L.Koch. Ubers. Arach., v. 3 p. 110 j 1855 Hermannia,
Nicolet in : Arch. Mus. Paris, v. 7 p. 468 | 1885 H., Canestrini, Prosp. Acarof., v. 1 p. 9
| 1888 H., Michael. Brit. Orib., v. 2 p. 439 | 1896 H., + Angelia (part.), Berlese. A. M. S.,
Crypt. II p. 24, 26, 31.
Abd. unsegmented. Dors, plate of Abd. convex, full chitinized, not
carrying cast larval or nymphal skins of Dors, of Abd. Genital and anal covers
separate and inserted in a ventr. plate. Dors, of Abd. rough in all known
species. Legs short, pair 4 scarcely passing post, margin. Ung. mond.
9 species.
Key to the species.
With a tubular projection on each side of Abd. 2.
No tubular projection 3.
I A quadrangular plate with sligthly raised edge on Cephth.
between Psstg. L. 750 ju. 5. H. granulata . p. 64
No such plate. L. 720 n 1. H. arrecta . . p. 62
{Dors, of Abd. without any margin differently marked
from central portion 4.
Dors, of Abd. with a lat. margin different from central
portion and separated from it by a ridge 8.
t Dors, of Abd. thickly strewn with raised dots or pits 5.
4 -j Dors, of Abd. very coarsely and irregularly reticulated.
- L. 850 n 8. H. reticulata . p. 64
Abd. globose or oval 6.
Abd. long, sack shaped L. 500 |u 6. H. nanus . . p. 64
Pstsg. org. long, filiform 7.
Psstg. org. short, clavate. L. 870 |u 9. H. scabra . . p. 65
{Hairs on first 3 joints of legs spatulate; 4 rows and
a border of hairs on Dors, of Abd. L. 820 |u . 4. H. convexa . p. 63
Hairs on first 3 joints of legs setiform; 2 rows and
a border of hairs on Dors, of Abd. L. 250 n . 7. H. ovnlum . . p. 64
/ Margin of Dors, of Abd. with transverse ridges; the
2 ridges which divide margin from centre of dors,
a I having a spoon-shaped depression between them
I posteriorly. L. 850 p 2. H. bistriata . p. 63
No transverse ridges on margin; no spoon-shaped de-
pression. L. 750 |u 3. H. carinata . p. 63
1. H. arrecta Nic. 1855 H. a., Nicolet in: Arch. Mus. Paris, 0.7 p. 470 t. 9 f . 5
| 1888 H. a., Michael, Brit. Orib., v. 2 p. 445 t. 43 f. 815.
Im. Dull red -brown, often carries cast skins. Cephth. triangular,
granular, no other markings. Psstg. projecting, cup-shaped. Psstg. org.
Hermannia 63
long, slightly incrassated towards distal end. Abd. finely granular with a
chitinous tubular projection on each side. Nph. Very light rosy
purple, almost white, legs and tip of Host, reddish, leathery; Abd. with small
scattered papillae and a roll of cuticle along ant. margin, otherwise resembles.
Im. - - L. ca. 720 |a.
On dead wood and moss. Algeria, England, France.
2. H. bistriata (Nic.)*) 1840 Nothrus palliatus, C.L.Koch. C. M. A., fasc. 30
f. 4 (Nph.!) 1896 Angelia palliata, Berlese, A.M. S., Crypt. IE p. 26 , 1855 Nothrus bi-
striatus (err., non C.L.Koch 1840!). Nicolet in: Arch. Mus. Paris, v.7 p. 397 t. 7 f . 7 |
1885 N. b., Berlese, A.M. S., fasc. 17 t. 9 | 1880 Hermannia bistriata, Michael in: J. R.
micr. Soc.. v. 3 p. 42 | 1888 H. b., Michael, Brit. Orib., v.2 p. 462 t. 42 f. 8 14 |
1876 yothrus cirrosus, Canestrini & Fanzago in : Atti Soc. Veneto-Trent.. v. 5 fasc. 1
p. 103 (Nph!).
Im. Dark brown or black. Cephth. rough, deeply pitted; Psstg. org.
moderately long, rod-like or clavate. Intlam. hair short, thick, white; legs
short, thick. Abd. long somewhat rounded posteriorly, ant. margin a rough
ridge. Dors, of Abd. sparsely but irregularly punctured, bears 6. (sometimes 4)
longitudinal ridges the whole length of Dors, of Abd. 2 form lat. edge ; short, irre-
gular-, transverse ridges unite lat. ridge to next (where there are 6). A basin-
shaped depression in centre of post, margin between the 2 median
ridges. Curved hairs round post, margin. Nph. Legs and part of Cephth.
red-brown, chitinous; remainder and Abd. yellowish white, leathery, granular.
No Psstg. or Psstg. org. visible. Abd. sack-shaped, slightly wrinkled, a row.
of curved hairs round post, margin, and 2 longitudinal rows on Dors, of Abd.
Im. L. ca. 850 n.
Im. terrestrial on moss. Nph. amphibious on land moss or Sphagnum. France.
Germany, Italy.
3. H. carinata P. Kram. 1897 H. c., P.Kramer in: Bibl. zool., 0.20 p. 81.
Im. Very like H. bistriata, but without the spoon -shaped
depression at post, end of Dors, of Abd. and without the trans-
verse ridges on the marginal band of Dors, and the hairs on the same
rather shorter, and ridges on Cephth. more curved than in H. bistriata.
There are 2 small apodemata on ant. part of Cephth. not found in H. bistriata.
L. ca. 750 n.
Greenland.
4. H. convexa (C.L.Koch)**) 1840 Nothrus convexus, C.L.Koch, C. M. A.,
fasc. 29 t, 1 | 1840 N.piceus, C.L.Koch. C. M. A., fasc. 29 t. 2 i 1886 N.p., Berlese,
A. M. S., fasc. 33 t. 6 j 1888 Hermannia picea, Michael, Brit. Orib.. v.2 p. 448 t. 41
f. 19 i 1896 H.p., Berlese, A. M. S., Crypt. II p. 91 | 1840 Nothrus gibbus, C. L. Koch,
*) According to the synonyma enumerated above by the author and in
accordance with the ,.Regeln fur die wissensch. Benennung der Thiere" the
name of this species is: H. palliata (C.L.Koch). The Editors.
See my note p. 7. The Author.
**) The well-known name of this species generally employed is H. picea. Koch's
Nothrus piceus and his N. convexus, are the same species (synonyms). Koch's work
was published in fasciculi of loose sheets (one to a species) without any arrangement
creatures of different families and orders being frequently mixed in the same fasciculus ;
they were intended to be arranged subsequently; but a number, to render reference
convenient, was put on each sheet; N. picea and N. convexa were in the same fasciculus
but the latter had the distinctive number 1. the former. 2. Under these circumstances
I should not have considered it necessary to disturb the well-known name of n picea"
but the Editors hold that n convexa" has the priority. A.D.Michael.
64 Hermannia
C\ M. A., fasc. 29 t. 4 1855 Hermannia crassipes, Nicolet in : Arch. Mus. Paris, v. 7
p. 469 t. 9 f. 4 ; 1882 H. c., Haller in: Jahresh. Ver. Wurtemb., v.38 p. 306.
Im. Black, rough, coarsely dotted. Cephth. almost as broad at base
as Abd. Psstg. org. long, filiform. Intlam. hah- thick, spatulate, slightly
curved. Legs thick, rough, with whorls of spatulate hairs Genual nearly as
long as Tibia Abd. very convex, raised dots on Dors, of Abd. about 90 to mm 6
longitudinal rows of thick spatulate white hairs on Dors, of Abd. Nph.
Yellow-brown, dull. Cephth. and legs granular. Cephth. with median longi-
tudinal sulcation and mamilliform elevation on each side. Psstg. org. and
Intlam. hair as in Im. Abd. less convex than in Im. Dors, of Abd. covered
with wavy wrinkles mostly transverse. Hairs on Dors, of Abd. as in Im. -
m . L. ca. 820 n.
On moss and wood. England. France. Germany, Sweden.
5. H. granulata Nic. 1855 If. g., Nicolet in: Arch. Mus. Paris, c.l p. 469 t. 9
f. 6 1883 H. g., Berlese. A. M. S., fasc. 9 nr. 4.
Im. Like H. arrecta except, that there is a quadrangular chitiuous
plate on Cephth., and Dors, of Abd. coarsely granular. L. ca. 750 |u.
In moss and wood. France, Sicilia.
6. H. nanus (Nic.) 1855 Nothrus n., Nicolet in: Arch. Mus. Paris, c.l p. 458
t. 7 f. 5 | 1888 Hermannia n., Michael, Brit. Orib., v.2 p. 455 t. 43 f. 1 -7 1892 H.
nana, Berlese, A. M. S., fasc. 63 nr. 1.
Im. Dark reddish brown, rough, dull, coarsely pitted, long and
narrow. Cephth. conical, deeply excavated for insertion of legs. Psstg. on
chitinous mammillae. Psstg. org moderately long, slightly thickened toward
distal end. 2 longitudinal ridges on Cephth. Dors, of Abd. very arched
with 8 longitudinal rows of white rod-like hairs. - - Nph. Very like Im.,
but Abd. white. - - Im. L. ca. 500 |u.
In moss. England. France, Italy.
7. H. ovulum (Haupt) 1882 Nothrus o., Haupt in: Ber. Ges. Bamberg, t'.12 p. 21.
Im. Said to be like Nothrus gibbus (i. e. H. convexa), but smaller, and
hairs on first 3 joints of legs setiform, not clavate; and only 2 rows
of about 3 hairs each on Dors, of Abd. but a marginal row of 4 hairs is
present. L. ca. 250 |uu
In moss. Germany.
8. H. reticulata Thor. 1871 H. r., Thorell in : Ofv. Ak. Forh.. v. 28 p. 696 1879
H. r., L.Koch in: Svenska Ak. Handl., v. 16 p. 112 t. 3 f. 21 | 1888 H.r., Michael,
Brit. Orib., . 2 p. 458 t. 42 f. 17.
Im. Black or dark brown, rough, dull. Cephth. finely and regularly
reticulated. Psstg. org. with very short peduncle and short pyri-
form head. Intlam. hair short spatulate. Legs reticulated, Genual as long
as Tibia. Abd. elliptical with a projecting post, lobe, rather long, covered
with irregular ridges forming very irregular reticulation; ridges composed
of coalesced dots; surface between ridges finely granular. 4 longitudinal
rows of spatulate hairs on Dors, of Abd. and 3 pairs on post, margin.
Ventral surface finely reticulated. - - Nph. Like a Nothrus ; light yellow
brown, leathery, rough; legs and Rost. reddish, chitinous. Cephth. and legs
resemble Im. Abd. oblong, constricted a little before post, margin, post angles
Hermannia. Xeoliodes 65
formed by raised lobes. Dors, of Abel, flat with raised lumps and rolls,
hairs resemble Im. - - Im. L. ca. 850 ju.
In lichen and moss near the sea. England. Spitzbergen. Xovaya Zemlya.
9. H. scabra (L.Koch) 1879 Nothrus scaber, L.Koch in: Svenska Ak. Handl.,
r. 16 p. 113 t. 3 f. 24 1888 Hermannia nodosa, Michael, Brit. Orib., 0.2 p. 452 t. 41
f. 10- 11 ; 1896 H. n., Berlese. A. M. S., Crypt, II p. 31.
Im. Very like H. convexa, but Psstg. org. short, clavate, blunt-ended,
roughened with small points near distal end. Abd. rather longer than in
H. convexa. Dots on Dors, of Abd. sometimes coalesce and form ridges or
network. L. ca. 870 ju.
Amphibious on moss lichen and fresh-water algae. England, Xovaya Zembla,
Siberia (Yeniseisk).
2. Gen. Neoliodes Berl.
1826 Liodes (non Leiodes, corr. Liodes, Latreille, Coleopt., 1796!), Heyden in: Isis
p. 608 | 1885 L., Berlese, A. M. S., f'asc. 17 nr. 10 | 1895 L., Banks in: Tr. Amer. ent.
Soc.. 0.22 p. 5 ; 1842 Nothrus (part.), C.L.Koch. Ubers. Arach., v.3 p. 110 [ 1877
N. (part.). Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 v.4 p. 77 | 1885 N. (part.),
Canestrini, Prosp. Acarof., v. 1 p. 28 | 1888 N. (part.), Michael. Brit. Orib., v. 2 p. 479 !
1888 Neoliodes, Berlese in: Bull. Soc. ent. Ital.. 0.20 p. 47 ; 1896 N., Berlese, A. M. S.,
Crypt, n p. 32.
Abd. unsegmented. Dors, of Abd. convex, not fully chitinized, but
carrying cast larval and nymphal skins of Dors, of Abd. Ung. trid.
3 good and 2 doubtful species.
Key to the good species.
I Dors, of Abd. without bordering hairs 2.
Dors, of Abd. bordered by strong semicircular
I hairs, which spring from margin and also touch
it with their tips . . . . 2. N. peltifer . . . . p. 65
Cast skins of Dors, of Abd. shield-shaped, pitted.
with a post, point bearing 2 spatulate hairs. -
L. 1 1,6 mm 3. N. theleproctus . p. 66
I Cast skins of Dors, of Abd. oval without post, point;
I markings on them radiating striae. L. 1,5 mm 1. W. concentricus . p. 65
J
1. N. concentricus (Say) 1821 Oribata concentrica, Say in: J. Ac. Philad.,
r. 2 p. 68, coll. writ. v. 2 p. 16 j 1895 Liodes concentricus, Banks in : Tr. Amer. ent. Soc.,
?.22 p. 15 j 1840 Nothrus scaliger, C.L.Koch, C.M.A., fasc.29 1. 11 \ 1890 N. s., Michael
in: P. zool. Soc. London, .p. 418 | 1883 N. theleproctus, Berlese, A. M. S., fasc. 3 nr. 1 |
1885 Liodes t., Berlese, A. M. S.. notes fasc. 3 p. 6 [ 1896 Neoliodes t., Berlese. A. M.S.,
Crypt, II p. 32.
Im. Like N. theleproctus, but Abd. less sharply truncated anteriorly,
hardly at all drawn out posteriorly and hairs at post, end much smaller,
no raised margin to cast skins: they slope down from the centre and
except the central part between larval skin and ant. margin of Abd. are
marked with radiating striations instead of areolations. L. ca. 1,5 mm.
Algeria. Germany, Italy, Sweden.
2. N. peltifer (C. L. Koch) 1840 Nothrus p., C. L. Koch. C. M. A., fasc. 29 t. 9
| 1879 N.punctatus, L.Koch in: Svenska Ak. Handl.. 0.16 p. 114 t. 3 f. 25 (Nph.?) | 1882
N. bicristatus -\- ?N. spirofilus, Haupt in: Ber. Ges. Bamberg. 0.12 p. 23, 24.
Das Tierreich. 3. Lief. : A. D. Michael, Oribatidae. 5
66 Neoliodes, Cymbaeremaeus
Im. Blackish brown, resembles N. theleproctus, but Abd. more oval and
bordered by curved setiform hairs which spring from the margin and form
about a semi-circle so that their tips also touch the margin. Size ,.mehr
als mittelgross".
In moss and dead wood. Germany, Sweden.
3. N. theleproctus (Herm.) 1804 Notaspis t., Hermann, Mem. apt., p. 91 t. 7
f. 5 ' 1826 Liodest., Heyden in: Isis. p. 608 j 1840 Nothrus t., C.L.Koch. C. M. A.,
fasc.29 t, 10 | 1888 N.t., Michael, Brit.Orib., v.2 p. 522 t.45 f. 1 9 | 1840 N.farinosus,
C.L.Koch, C. M. A., fasc.29 f . 8 (with one cast skin only) 1 1876 N. dorsattis, Cane-
strini & Fanzago in: Atti Soc.Veneto-Trent., v.o p. 101 (Nph.) I ?1882 N. circumv allot us
-\-N.dimera, Haupt in: Ber. Ges. Bamberg, v. 12 p. 22, 23 | ?1883 N. malleolus, Kar-
pelles in: Berlin, ent. Z., v. 28 p. 38 j 1883 N. Doderleinii, Berlese, A. M. S., fasc. 3 nr. 2
1885 Liodes D., Berlese, A. M. S., notes fasc. 3 p. 6 , 1896 Neoliodes D., Berlese. A.
M.S., Crypt. II p. 32 || non 1883 Nothrus theleproctus, Berlese, A. M. S., fast-. 3 nr. 1.
Im. Dark 'chestnut-brown, rough, dull. Cephth. broad, a sulcation
between it and Abd. covered with vermiform ridges ; 2 broad lobes on post,
median portion. Psstg. large, projecting. Psstg. org. of moderate length,
slightly clavate, directed backward. Legs thick, flat, rough; most leg-hairs
spatulate. Abd. truncated and straight anteriorly, curved laterally, drawn
out to a rounded point bearing 2 spatulate hairs posteriorly entirely
hidden by cast skins of Dors, of Abd., which are carried flat and almost
concentrically, each skin bordered by a raised roll, cast skins coarsely
areolated all over. Nph. Resembles Im. except in color. Im.
L. ca. 1 1,6 mm.
In moss. England, Germany, Italy, Paraguay. Switzerland. ? Pennsylvania.
N. kornhuberi (Karp.) 1883 Nothrus k., Karpelles in: Arch. Naturg., r. 491
p. 455.
Im. Dark brown, pentagonal; Cephth. small. Abd. large, arched; cast skins
carried on Dors, of Abd. in a cornucopia-shape. Body and legs covered with short
hairs. Ung. trid. Size ? Imperfectly described from a broken specimen in the Berlin
Museum. Probably a good species, probably a Neoliodes.
Japan.
N. venosus (Grube) 1859 Nothrus v., Grube in: Arch. Dorpat. Ges., ser. 2 r. 1
p. 476.
Im. Said to be something like Nothrus farinosus C. L. Koch, which is probably
Neoliodes theleproctus, without the larval and nymphal skins, but present species has
8 small processes on post, margin. L. ca. 400 jn. Probably a Neoliodes, possibly
immature, but not improbably a distinct species.
Gennany.
3. Gen. Cymbaeremaeus Berl.
1855 Eremaeus (part.) (non Eremaeus C. L. Koch 1835!), Nicolet in: Arch. Mus.
Paris, v.l p. 450 ] 1877 E. (part.), Canestrini & Fanzago in: Atti 1st. Yeneto, ser. 5 r. 4
p. 77 1885 E. (part.), Canestrini, Prosp. Acarof., v. 1 p. 9 j 1886 E. (part.), Berlese,
A. M.S., fasc. 33 nr. 10 ' 1888 E., Michael, Brit.Orib., r.2 p. 467 1896 E. (part.),
Oudemans in: Tijdschr. Ent., v. 39 p. 56 | 1896 Cymbaeremaeus, Berlese, A. M. S.,
fasc. 78 nr. 1.
Abd. unsegmented ; Dors, of Abd. flattened, undulated or concave, rough
and sculptured. Tectp. present to leg 2. Ung. trid.
4 good and 1 doubtful species.
Cymbaeremaeus 67
Key to the good species.
( Psstg. org. short, with short peduncle and small almost globular head 2.
\ Psstg. org. long, straight, rod-like L. 700 |u . . . 4. C. tibialis ... p. 68
I Markings on Dors, of Abd. coarse reticulations 3.
2 < Markings on Dors, of Abd. ridges, more or less
transverse, radiating on margin. L. 380 |U . . 3. C. fimbriatus . p. 67
j Margin of Dors, of Abd. turned up. - - L. 400
3 700 n 2. C. cymba . . . p. 67
' Margin of Dors, of Abd. turned down. -- L. 290 |u 1. C. brevipes . . p. 67
1. C. brevipes (Michael) 1888 Eremaensb., Michael, Brit. Orib., v.2 p. 475
t. 44 f. 1219 | 1896 E. b., Berlese, A. M. S., Crypt. II p. 36 | 1891 E. minimus, Berlese,
A. M.S.. fasc. 58 nr. 9 j 1891 E. Fockeui, Moniez in: Rev. biol. Nord France, v. 3 nr. 6.
Im. Light to dark brown, dull, rough; short pyriform. Cephth. with
raised dots but no regular markings. Psstg. org. with short peduncle and
globular head. Legs short, 4 scarcely reaches post, margin. Ung. almost
homd. Abd. widest near post, margin ; convex in centre; a broad margin
flat or sloping downward. Dors, of Abd. rather regularly reticulated,
reticulations about 9 jn ; a few very fine curved caducent hairs on Dors, of Abd.
and post, margin. - - Nph. Yellow-brown, dull, leathery. A sulcation between
Rost. and remainder of Cephth., which has longitudinal folds. Psstg. org.
like Im. Dors, of Abd. with some large cuticular folds regularly arranged,
6 short clavate hairs on post, margin in groups of 3. Im. L. ca. 290 M.
On trees, particularly Oaks. England.
2. C. cymba (Nic.) 1855 Eremaeus c., Nicolet in: Arch. Mus. Paris, v. 7 p. 452
1. 10 f. 3 I 1877 E. c., Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5 u4 p. 90 | ?1886
E.c., Berlese, A.M.S., fasc. 33 nr. 10 j 1888 E.c., Michael, Brit, Orib., v.2 p. 470 t.44
f. 1 11 j 1896 Cymbaeremaeus c., Berlese, A. M. S., Crypt. II p. 37.
Im. Light yellow-brown, dull, rough ; almost pyriform ; slightly pointed
posteriorly. Kost. very blunt and round; Psstg. org. with very short
peduncle and globular head. Cephth. coarsely granulated and bears
longitudinal and transverse ridges. Legs short, rough, femora large, those
of legs 1 and 2 with slender peduncles. Ung. hetd. Abd. oval, truncated,
ant. margin sharply raised, centre slightly convex. Whole Dors, of Abd.
coarsely and irregularly reticulated by ridges, interspaces more finely
reticulated. Some very short thick caducent hairs on dors, plate of Abd.
and post, margin. Ventr. surface reticulated like dorsal. Nph. Buff.
Legs and rost. reddish; dull, leathery, resembles Im., but rost. less broad,
femora without fine peduncles. Dors, of Abd. and ventr. surface irregu-
larly wrinkled instead of reticulated, hairs minute with short peduncle and
globular head. - - Im. L. ca. 400 700 \JL.
On Oak trees etc. England, France, ? Italy, Netherlands.
3. C. fimbriatus (Michael) 1890 Eremaeus f., Michael in: P. zool. Soc. London,
p. 422 t. 37 f. 6 j ?1896 E.marginalis, Banks in: Tr. Amer. ent. Soc., v.23 p. 76.
Im. Red-brown, rough, dull. Cephth. arched, bears numerous irregular
rough dots and ridges. Psstg. org. very short, peduncle hardly shows, head
globular. Legs short, 4. not reaching post, margin. Abd. compressed dorso-
ventrally. Ant. margin of Abd. truncated, post, margin round. Central
part of Dors, of Abd. arched, elliptical, has slightly turned-up edge; out-
side this a deep trench and then a broad, slightly arched border. Whole
68 Cymbaeremaeus. Nothrus
Dors, of Abd. bears numerous rough, broken, irregular, more or less
transverse ridges, which are radiating on margin. Ung. hetd.
L. ca. 380 M.
Algeria, ? America.
4. C. tibialis (Nic.) 1855 Eremaeus t., Nicolet in: Arch. Mus. Paris, v.l
p. 452 t. 10 f. 2.
Im. Dark red-brown, finely granulated, legs yellow, smooth. Psstg.
org. long, straight, rigid, directed obliquely backward. Abd. oval, Dors.
of Abd. concave, hairless, a red, polished, almost circular boss in
median line close to ant. margin, a similar projection, but rough, on post.
margin. - - Nph. Greenish-yellow with red shades, sub-fusiform, almost flat.
Rost. sharply demarcated from remainder of Cephth. 2 or 3 transverse lines
like segmentations on Cephth., not always seen on fully grown Nph.
Psstg. org. like Im. Abd. widest anteriorly much narrowed posteriorly,
terminating in 2 papillae each bearing a hair longer than body. - Im.
L. 700 |ii.
France. Rare.
C. minutus (C. L. Koch) 1835 Cepheus m., G. L. Koch, C, M. A., fase. 3 1. 12.
Im. Dark brown; dull, rough. Psstg. org. long, gradually clavate. Abd.
elliptical, centre slightly convex, finely wrinkled, a border with edge raised; 4 short
hairs on post, margin, generally a light smooth spot on Dors, of Abd. near ant.
margin. L.?
Germany.
4. Gen. Nothrus C.L.Koch
1835 Nothrus (part.). C.L.Koch, C. M. A., fasc. 2 j 1842 N. (part.), C. L. Koch.
Ubers. Arach.. v. 3 p. 110 i 1855 N. (part.). Nicolet in: Arch. Mus. Paris. r. 7 p. 453 |
1859 N. (part,). Grube in: Arch. Dorpat. Ges., ser. 2 v.l p. 417 j 1877 N. (part.). Cane-
strini & Fanzago in : Atti 1st. Veneto, ser. 5 v. 4 p. 77 I 1885 N. (part.). Canestrini, Prosp.
Aearof., ft 1 p. 9 | 1888 N. (part.). Michael, Brit. Orib.. v.2 p. 479 ' 1895 N., Banks
in: Tr. Amer. ent. Soc., c. 22 p. 13 | 1896 N. + Angeliu (part.). Eerlese. A. M. S., Crypt, II
p. 24, 26.*)
Abd. unsegmented, dors, plate flat, concave or undulated ; without Tectp.
Legs short.
19 good and 4 doubtful species.
Key to the good species.
} Ung. mond. 2.
1 \ Ung. trid. 4.
Abd. not pointed posteriorly; Psstg. of the ordinary type 3.
Abd. shield - shaped (pointed posteriorly) ; no
projecting tubular Psstg. L. 290 |n . . . . 12 N. monodactylus . p. 72
Post, margin of Abd. rounded, hairs thereon
arising from apophyses. L. 830 |ii . . . . 19. N. targionii . . . p. 74
Post, margin truncated, hairs thereon sessile. -*
L. 660 M. . . . . ............. 17. N. sylvestris . . . p. 73
r
<
*) The genus Angelia is formed to receive part of Koch's genus Nothrus including
N. palustris, which appears to be Berlese's type, but as this is Koch's type it cannot
be moved into a new genus but must remain the type of Nothrus. if the genus were
divided the other group should be renamed, but the present author does not consider
division necessary. A. D. Michael.
f
\
11
I
Nothrus 69
) Abd. shield-shaped, pointed posteriorly 5.
| Abd. not pointed posteriorly 6.
f Cuticle finely reticulated L. 360 |u 18. N. tardus .... p. 74
1 \ Cuticle smooth. -- L. 600 |u 9. N". glaber . . . . p. 71
f Psstg. org. filiform 7.
\ Psstg. org. clavate 10.
Abd. truncated posteriorly 8.
Abd. rounded posteriorly 9.
Hairs on Dors, of Abd. setiform; the pair on
post, corners 4 times as long as the others
on post, margin. L. 1,1 mm 13. KT. palustris ... p. 72
Hairs on Dors, of Abd. spatulate or rod-like ; the
pair on post, corners not twice as long as the
others on post, margin. L. 800 |u .... 1. M". anauniensis . . p. 70
Dors, of Abd. convex; hairs thereon clavate . . 4. N. biciliatus . . . p. 70
Dors, of Abd. concave, hairs thereon setiform.
[ L. 700 n 2. N. banks! .... p. 70
/ Spines on lat. margin of Abd. as long as width
of Abd. and arising from apophyses which
10 ' project beyond the margin 11.
I Spines (or hairs) on lat. margin of Abd. short;
no projecting apophyses - 12.
Apophyses long; Abd. oblong, truncated and
slightly concave posteriorly, the 2 apophyses at
postero-lateral corner of Abd. with bulbous
proximal parts. L. 900 |U 16. N. spinifer .... p. 73
Apophyses short; sides of Abd. somewhat curved,
it is narrowed and deeply bifid posteriorly,
apophyses at postero-lateral corners without
bulbous bases , . . 6. N". echinatus ... p. 71
/ Im. carrying the cast skin of post, margin and
of 2 long conical projections at post, angles
12 . of Abd. of Nph., these enclose the same part
in Im. as in a bag. L. 0,8 1,2 mm . . . 15. N". segnis . . . . p. 73
I Im. not carrying any cast skin. - - 13.
I Post, margin deeply incised (emarginate) 14.
13 { Post, 'margin truncated or slightly concave or
convex 15.
Incision comencing near lat. margin, causing post,
margin to be bifid; Dors, of Abd. flat without
raised margin ; 6 clavate hairs on post, margin.
L. 700 |u 7. N". excisus . . " . . p. 71
Incision only central; Dors, of Abd. with strongly
raised margin ; 4 pectinated hairs with reflexed
distal ends on post, margin; central pair
transverse and nearly as long as Abd. -
L. 700 n 8. N. flagellatus . . p. 71
j Post, margin with a broad central projection
lower in level than the Dors, of Abd. 16.
I Post, margin without any central projection 17.
/ Central projection with small tubercles at corners
each bearing a large thick hair. -- L. 800 |u 10. N. horridus . . . p. 71
' Central projection with very large, almost
cylindrical tubercles at corners, each bearing
a very small sharply curved hair. L. 1 mm 5. N". biverrucatus . p. 71
70 Nothrus
/ Hairs on margin of Abd. and Host, hairs setiform
I and pectinated 18.
1 Hairs on margin of Abd. and Rost. hairs clavate.
L. 700 n 14. N". rugulostts . . . p. 73
.o j Post, margin convex. L. 660 |u 11. N". invenustus . . p. 72
\ Post, margin concave L. 750 (a 3. N". bicarinatus . . p. 70
1. N. anauniensis Can. &Fanz. 1876 N. a., Canestrini & Fanzago in: Atti Soc.
V T eneto-Trent., v.5 p. 102 ' 1877 N. a., Canestrini & Fanzago in: Atti 1st. Veneto, ser. 5
0.4 p. 92 t. 2 f. 2 ; 1885 N. a., Berlese, A. M. S.. fasc. 17 nr. 7 | ?1895 lY. truncatus,
Banks in : Tr. Amer. ent. Soc., v. 22 p. 14 1896 Angelia sylvestris, Berlese. A. 31. S. r
Crypt, n ]>. 26.
Im. Like N. sylvestris, but ung. trid. or bid. through the atrophy of one-
lateral Ung. of trid. Ung. Post, and lat. margins of Abd. rounder. L. ca.
800 )n.
This Species is stated in error by Canestrini (1885. Prosp. Acarof., v. 1 p. 30) to-
be identical with N. palustris.
Algeria, Italy and ?New York State.
2. N. banks! Michael*) 1895 N. furcatus (non C.L.Koch 1844!), Banks in:
Tr. Amer. ent. Soc., v. 22 p. 14.
Im. Reddish brown. Apparently very like N. targionii, but hairs round
Abd. and Rost. hairs not stated to be pectinated, and there is stated to be
a curved transverse ridge between the post, ends of the raised lateral margin,
and in median line of Dors, of Abd. a forked ridge reaching
from ant. margin to near transverse ridge. 4 simple hairs on each
branch of fork. Ung. trid. L. 700 |u.
Washington State.
3. N. bicarinatus C. L. Koch 1840 N. b., C. L. Koch, C. M. A., fasc. 29 t. Iff
| 1855 N. b., Xicolet in: Arch. Mus. Paris, v.7 p. 456 t. 7 f . 3 ] 1877 N. b., Canestrini &
Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 94 | 1885 N. b., Berlese, A. M. S., fasc. 17 nr. 4
1888 N. b., Michael. Brit. Orib., v.2 p. 514 t. 47 A f. 15 | 1840 N. ventricosus,
C. L. Koch, C. M. A., fasc. 29 t, 17 | 1840 N. restrains, C. L. Koch, C. 31. A., fasc. 29
1. 19 (with dirt on Lam. hair) | 1844 N. pigerrimus, C. L. Koch, C. M. A., fasc. 38 t. 3.
Im. Resembles N. horridus, but has not central flat projection below
post, margin of Dors, of Abd. L. ca. 750 ju.
On trees etc.. especially Oaks. England, France, Grermany, Italy.
4. N. biciliatus C. L. Koch 1844 N. b., C. L. Koch, C. 31. A., fasc. 38 t. 2.
Im. Cephth. black, Abd. red-brown. Psstg. org. very long, thick
at base, but thin and setiform for the greater part of lenght.
Abd. oval, truncated anteriorly, flat; central part slightly raised and bearing
at its edge a row of curved clavate hairs, then comes a deep sulcation
then abroad raised border bearing a row of similar but larger haira
on outer edge. Legs with clavate hairs. Ung.'? L. ,,mittelgross''. -
A good species imperfectly described.
In moss on wood. Germany.
*) The name of furcatus used by Banks is preoccupied in the genus Xothrus,.
C. L. Koch having used it for a creature which is really a Nymph of Nothrus segnis,
but although Koch's name has failed it cannot, under the rules which govern ,,Das-
Tierreich" be repeated in the same genus. I have therefore named the species in
honor of Mr. Banks. A. D. Michael.
Xothrus 71
5. N. biverrucatus C. L. Koch ?1840 N. angulatus, C.L.Koch, C. M. A.,
fasc. 29 t. 14 | 1840 N. liver rucatus, C. L. Koch, C. 31. A., fasc. 29 1. 15 1888 N. b.,
Michael. Brit. Orib., v. 2 p. 510 t. 47 A f. 6 10 | ?1855 N. horridus, Nicolet in: Arch.
Mus. Paris, v. 1 p. 456 t. 7 f. 4 | ?1885 N. h., Berlese, A. M. S., fasc. 17 nr. 1.
Im. Very like N. horridus but larger, heads of Psstg. org. less globular,
and flat projection below post, margin of Abd. with a large truncated tubercle
at each post, corner bearing a minute sharply curved hair in the centre of
distal surface. L. ca. 1 mm.
England. Germany. France?, Italy?, Sweden.
6. N. echinatus C. L. Koch 1835 N. e., C. L. Koch. C. M. A., fasc. 2 1. 17
1882 N. ansatus, Haupt in : Ber. Ges. Bamberg. v. 12 p. 24.
Very like N. spinifer, but Abd. longer and bifurcating at post, end
so as to form 2 conical tails, as in N. segnis, when that species is carrying
the post, part of the nymphal skin of Dors, of Abd.
In moss etc. Germany.
7. N. excisus Banks 1895 N. e., Banks in: Tr. Amer. ent. Soc., v. 22 p. 15.
Im. Apparently intermediate between N. segnis and N. bicarinatus.
Pale brown, Cephth. truncate anteriorly. Psstg. org. short, capitate; Abd.
flat without raised margin; truncated anteriorly, deeply emarginate
posteriorly; a rough clavate hair on each post, angle, a sub-clavate
hair just ant. to angle; a pah- of shorter clavate hairs in centre of post,
margin. A sub-median ridge each side from near ant. margin, which near
post, margin curves outward to lat. angle; where curve starts is a trans-
verse ridge; 3 or 4 short clavate hairs on each lat. margin. Ventr. surface
and legs rough; numerous curved stiff hairs on legs. Most hairs on body
finely serrated. Gng. trid. L 700 ju.
On bark of spruce firs. New York state.
8. N. flagellatus Berl. 1888 N. f., Berlese in: Bull. Soc. ent. Ital., f. 20 p. 216
t. 13 f. 5.
Im. Earthy-brown. Abd. almost quadrangular, elongated, concave;
margins raised, with 4 transverse ridges on them ; post, margin with a central
incision its whole depth, a tubercle projects at each side of the incision
forming a prolongation of it; each tubercle bears a strong pectinated
hair almost as long as Abd. but sharply recurved toward distal end;
this hair directed transversely; one other pair of similar but shorter hairs
on post, margin directed backward. L. ca. 700 |u.
Under bark of trees. Argentina (Buenos-Ayres).
9. N. glaber Michael 1888 N.g., Michael, Brit. Orib., v.2 p. 525 1896 Angelia
gldbra, Berlese, A. M. S., Crypt. II. p. 26.
Im. Like N. tardus, but larger, smooth without reticulations ; Rost. more
pointed and ant. margin of Abd. without projecting corners. - - Nph. Like
Im., but lighter. - - Im. L. ca. 600 \JL.
Amphibious, in moss and sphagnum. England.
10. N". horridus (Herm.) 1804 Notaspis h., Hermann, Mem. apt., p. 90 t. 6 f. 3
| 1842 Nothrush., C.L.Koch, fibers. Arach., v. 3 p. 113 j 1888 N. h., Michael,
Brit. Orib., v.2 p. 503 t.47 f. 14, 713 \ 1844 Oribata horrida, Gervais in : Walckenaer.
Ins. Apt., v. 3 p. 254 | 1840 Nothrus mutilus, C. L. Koch, C. M. A., fasc. 29 1. 18 j
1840 N.Ustriatus, C.L.Koch, C. M. A., fasc. 29 t. 21 j 1840 N. sinuatus, C.L.Koch.
72 Nothrus
C. M. A., fast-. 29 t. 22 (Nph.) \ 1840 2V. runcinatus, C. L. Koch. C. M. A., fasc. 29 t. 23
| P1871 N.borealis, Thorell in: Ofv. Ak.Forh., .28 p. 697 | ?1885 N. angulatus, Eerlese,
A. M. S., fasc. 17 nr. 5 || non 1855 N. horridus, Xicolet in: Arch. Mus. Paris, r. 7 p. 456
t. 7 f. 1 | non 1885 JV r . h., Berlese. A. M. S.. fasc. 17 nr. 1.
1m. Brown, dull, rough, covered with short villous processes, usually
so concealed by dirt that neither color nor texture can be seen. 2 apophyses
on Kost. bear thick villous hairs, Psstg. org. with very short peduncle and
globular head. Legs fringed with thick hairs on apophyses. Ung. trid.,
almost homd. Abd. oblong, Dors, of Abd. flat or hollow, post, margin concave
with 2 apophyses near the angles bearing hairs; a flat projection at a
lower level bearing 2 hairs occupies the concavity. A sub-elliptical
raised ridge on Dors, of Abd. with 2 transverse ridges within and
4 diagonal from angles. 6 small apophyses on outside of this ridge bear very
short hairs but do not project beyond lat. margin of Abd. Nph. Yellowish-
white, covered with dirt. Somewhat like Im., but Dors, of Abd. and margins
of Abd. bear about 24 papillae, each with a thick curved hair. Im. L. ca.
800 |u.
At the roots of moss etc. Algeria. England. Germany, Switzerland.
11. N. invenustus Michael 1888 N. i., Michael, Brit. Orib., r. 2 p. 500 t, 47 A
f. 12.
Im. Very like N. horridus but post, margin of Abd. convex and
without central projection. Apophyses on Cephth. smaller than in N. horridus,
does not carry dirt. L. ca. 660 ju.
On lichen. England.
12. N. monodactylus Michael 1888 N. m., Michael, Brit. Orib., v. 2 p. 529
t. 45 f. 10 13 1896 Angelia monodactyla, Berlese. A. M. S., Crypt. II p. 26.
Im. Very like N. tardus but whitey- brown, slightly chitinized, not
reticulated. Legs 3 and 4 usually concealed beneath Abd. Ung. mond.
with a small square projection from tarsus on each side of Ung. A few
hairs round margin, none on Dors, of Abd. L. ca. 290 |u.
On moss and sphagnum. England.
13. N. palustris C. L. Koch 1840 N. p., C. L. Koch. C. M. A., fasc. 29 1. 13 |
1855 X.p., Nicolet in: Arch. Mus. Paris, v. 7 p. 457 t. 7 f . 6 I 1886 N. p., Berlese, A. M. S..
fasc. 30 nr.4 1888 N.p., Michael, Brit. Orib., v.2 p. 494 t.46 f.8 13 | 1896 Angeliap.,
Berlese, A.M.S., Crypt, II p. 26 ! 1844 Nothrus pollens, C.L.Koch, C.M.A., fasc. 38 t.4
(Nph.) ?1895 N.bipilis, Banks in: Tr. Amer. ent. Soc.. f. 22 p. 14.
Im. Dark red-brown, rough, dull; central parts coarsely pitted, pits
about 85 to mm peripheral parts finely granular. 2 longitudinal rolls of
cuticle on Cephth. ending posteriorly in mamilliform elevations, which bear
the Psstg. Psstg. org. very long, filiform, directed outward, usually
doubly curved. Legs long for genus; leg 4 articulated to a tubular projection
of ventr. plate. Ung. very hetd. Abd. wider posteriorly than anteriorly, its
lat. margins broad, curved, and raised at an angle, terminating posteriorly
in large bosses, median part of Dors, of Abd. arched. Ant. and post, margin
of Abd. almost straight, each post, angle bears a very long hair
directed backward and concave inward, 2 pairs of shorter hairs on post, margin,
2 rows on Dors, of Abd. and some on lat. edges. - - Nph. like Im., but Abd.
white and post, angles form conical projections. - - Im. L. ca. 1,1 mm.
In moss. England, France. Germany, Italy, Netherlands. Sweden and ?New
York State.
Nothrus 73
14. N. rugulosus .Banks 1895 K r.. Banks in: Tr. Amer. ent. Soc., c. 22 p. 15.
Im. Cephth. dark, Abd. pale brown. Like N. excisus but Abd. trun-
cated, slightly concave or convex posteriorly not deeply emarginate;
4 large clavate hairs on each lat. margin, margins roughened, but Dors, of
Abd. smooth. Rost. hairs very clavate. Ung. trid. L. 700 ju.
Under bark of dead trees. New York State and Illinois.
15. N. segnis (Herm.) 1804 Notaspis s., Hermann. Mem. apt., p. 94 t. 4 f. 8 |
1844 Nothrus s., C. L. Koch, C. M. A., fasc. 30 t. 1 1885 N. s., Berlese, A. M. S., fasc. 17
nr.2 j 1888 N. s., Michael, Brit. Orib., v.2 p. 517 t.48 f. 1- -6 | 1844 Oribatas., Gervais
in: Walckenaer, Ins. Apt., v.3 p. 254 V1840 Nothrus biurus, C.L.Koch, C. M. A.,
fasc. 30 t. 2 (Nph.!) j 1840 N. furcatus, C.L.Koch, C. M. A., fasc. 30 t. 3 (Nph.).
Im. Yellow-brown, dull, covered with raised dots, but usually partly
concealed by dirt. Cephth. flat, 2 apophyses extend beyond Rost. carrying
at their ant. ends thick, villous, curved hairs which cross. Psstg. org. with
short thin peduncle and pyriform head. Intlam. hair long villous. Legs
not reaching post, margin; broad, flat, bordered with curved spines. Ung.
trid. Abd. very long oblong, post, corners project and turn outward; ad-
joining inner side of projection is an apophysis bearing a thick, curved,
villous hair; but corners apophysis and hair are hidden in a conical
bag which is part of the nymphal skin; the 2 bags are joined by a
strip of nymphal skin which hides the post, margin and bears a thick curved
hair. An irregularly oval ridge on Dors, of Abd. - - Nph. Like Im., but
yellowish white, Rost. and legs reddish. Abd. bifurcate posteriorly, forming
2 long conical tails with a hair at post, end of each. Psstg. org. rod-like.
- Im. L. ca. 0,8 1,2 mm.
At the roots of mosses. England, Germany, Italy, Norway, Sweden.
16. N. spinifer C. L. Koch 1835 N. s., C. L. Koch, C. M. A., fasc, 2 1. 18 j 1885
JV. s., Berlese, A. M. S.. fasc. 17 nr. 3 | 1885 N. s., Canestrini, Prosp. Acarof., v. 1 p. 32 |
1855 N. spiniger, Nicolet in: Arch. Mus. Paris, v. 1 p. 455 t. 7 f . 2 j 1888 N. s., Michael,
Brit. Orib.. c. 2 p. 497 t, 48 f. 7 13 j 1882 N.concavus, Haupt in: Ber. Ges. Bamberg,
r. 12 p. 25.
Im. Chitinous brown, rough, leathery, granular, covered with villous
processes, but usually so hidden by dirt that neither color nor texture are
seen. Rost. hairs short, Intlam. hair long, both curved, villous, springing from
short apophyses; 2 very long similar hairs on long apophyses
between Rost. hairs and Intlam. hair, these cross in front of Rost.
Psstg. org. with very short peduncle and pyriform head. Legs thick, flat,
rough, with numerous thick curved hairs. Ung. trid. Abd. oblong, Dors,
thereof hollow, lat. margins and ant. margin raised. Ant. margin of Abd.
bears 3 pairs of villous curved spines on apophyses. Each lat. margin
has 5 similar spiniferous processes larger and more curved; a very
large apophysis and straight spine at post, angle, and 2 pairs with curved
spines on post, margin; all curved spines cross forming a basket. -
Nph. Like Im., but lighter and apophyses not so long or chitinized. -
Im. L. ca. 900 JLI.
Algeria, England, France, Germany, Italy.
17. N. sylvestris NIC. 1855 N. s., Nicolet in: Arch. Mus. Paris, v. 7 p. 458 t, 7
f. 4 1888 .V. s., Michael, Brit. Orib., r. 2 p. 490 t. 46 f. 1 7 || non 1896 Angelia s.,
Berlese, A. M. S., Crypt. II p. 26.
74 Nothrus
Im. Light chestnut-brown, dull, reticulated, almost oblong; reticulations
on Cephth. and lat. part, of Abd. ca. 8 |u on ventral part of Abd. ca. 16 |u.
Psstg. on chitinous mammillae ; a longitudinal sulcation between them. Psstg.
org. long, curved, filiform. Intlam. hair similar but shorter and thicker.
Coxae 3 and 4 inserted in tubular projections of sternal plate. Ung. mond.
Abd. almost oblong. Lat. parts of Abd. sharply raised at an angle, parti-
cularly anteriorly. Ant. margin of Abd. raised; central part of Dors, of
Abd. slightly arched; 4 rows of thick, curved, slightly spatulate, slightly
imbricated, white hairs on Dors, of Abd., 2 pairs on post, margin sessile,
and a few others. - - Nph. Very like Im., but Abd. yellowish white and
with raised dots instead of reticulations, but post, angles project and there
are 2 projections near centre of post, margin lower in level and bearing
hairs. - - Im. L. ca. 660 jn.
Algeria, England, France.
18. N. tardus Michael 1888 N. t., Michael, Brit. Orib., v. 2 p. 526 t. 47 f. 14 |
1896 Angelia tarda, Berlese, A. M. S., Crypt. II p. 26.
Im. Yellow-brown, finely and evenly reticulated, reticulations about 3 |u.
No exterior Psstg. of the ordinary type but a long setiform hair on each
side which is the homologue either of the Psstg. org. or the Intlam. hair.
Legs very short, thick, pairs 3 and 4 set under Abd. Ung. kid. Abd. flat,
shield-shaped, ant. margin straight, but antero-lat. corners of Abd. form
rounded projections each bearing a large hair directed outward. Central
and lat. parts of Dors, of Abd. slightly raised and rounded leaving a
shallow depression between, 2 longitudinal rows of about 3 hairs on Dors,
of Abd. - Nph. Like Im. but cuticle softer, not reticulated and with
numerous broad wrinkles. - - Im. L. ca, 360 ju.
On lichen. England.
19. N. targionii Berl. 1885 N. T., Berlese, A. M.S., fasc.17 nr.8 | 1888 N. T.,
Michael, Brit. Orib., v. 2 p. 488 t. 47 A f. 11 ! 1896 Angelia T., Berlese. A. M. S.,
Crypt. II p. 26.
Im. Dark brown. Cephth. pitted. Abd. smooth, not polished ; Host, small,
leg 1 set in deep excavation. Psstg. org. long filiform. Legs fringed
with thick curved hairs; Ung. mond. Abd. long, truncated anteriorly
rounded posteriorly hollow in centre; lat. and ant. margins sharply
raised, post, raised gradually; 4 large and 2 small apophyses on post,
margin bearing large, thick, sharply curved hairs; 4 similar hairs without
apophyses on each side of Abd. and some small hairs on Dors, of Abd., all
the large curved hairs on the creature are villous. L. ca. 830 |u.
This species is an approach to a Hermannia. particularly H. bistriata.
England, Italy; rare.
N. pileiformis Karp. 1883 N. p., Karpelles in: Berlin, ent. Z., v. 28 p. 87.
Im. Dark-brown, oval, broad anteriorly, slender posteriorly. Pileus-shaped
when seen from side; carries cast skins of Dors, of Abd. each of which has a wedge-shaped
incision. No hairs on body. Ung. trid. L. ? Probably a good species, but imper-
fectly described from broken specimens in Berlin Museum.
Pennsylvania.
N. quadracanthus Haupt 1882 N. q., Haupt in: Ber. Ges. Bamberg, v. 12 p. 24.
Im. Apparently like N. spinifer, but post, margin of Abd. more like N. horridus. L.?
In moss. Germany.
Nothrus. Lohmannia 75
N. simplex Banks 1895 N. s., Banks in: J. N. York ent. Soc., v. 3 p. 128.
Im. Pale yellow-brown. Cephth. ,,fat", triangular, concave in middle, sides
rounded. Intlam. hair long, Psstg. org. moderate length, clavate. Abd. much depressed,
Dors, of Abd. smooth with shallow depressions, truncate anteriorly broader and rounded
posteriorly, 2 simple hairs on margin, one central, one near post, end; ventr. surface
granular. Young said to be obovate with corrugated epidermis. Im. L. 900 |H.
In sphagnum-swamp. New York State.
N. sordidus C. L. Koch 1840 N. s., C. L. Koch, C. M. A., fasc. 29 t. 20.
Im. Yellowish-brown. Host, hairs not borne on apophyses. Abd. oblong, post,
margin concave. 2 very long hairs at post, angles standing outward and backward
but curving forward; a row of longish curved hairs on lat. margin and 2 on post,
margin. L. ,,mittelgross.".
In woods. Germany.
5. Gen. Lohmannia Michael*)
1884 (18. December) Michaelia (non Trouessart, Sarcopt.. November 1884 !), G. Haller
in: Arch. Naturg., w.501 p. 234 | 1885 Angelia (part.), Berlese, A. M. S., Crypt, II p. 26.
Cephth. and Abd. only divided by a simple line. Abd. almost cylindrical,
rounded posteriorly. Dors, plate much sufflexed on to the ventr. surface. Genital
and anal covers close together, only separated by a rim. common to both,
the latter reaching the post, margin, and these 2 pairs of covers occupying
the whole central part of the ventr. surface from a little behind the insertion
of legs 4 backward and touching the reflexed dors, plate on each side. Mdb.
very large. Maxillary palpus not so long as moveable arm of chela of Mdb.
Ung. mond.
Canestrini (1891, in: Atti 1st. Veneto, ser. 7 v.2 p. 718) and Banks (1895, in: Tr.
Amer. ent. Soc., v. 22 p. 2) state, that this genus is composed of immature creatures,
but this is an error.
2 species.
Key to the species.
Hairs on Dors, of Abd. setiform. L. 750 JLI 1. L. murcioides . p. 75
Hairs on Dors, of Abd. leaf-like. --- L. over 1 mm .... 2. L. paradoxa . . p. 75
1. L. murcioides (Berl.) 1896 Angelia m., Berlese, A. M. S.. fasc. 78 nr. 7.
Im. Similar to L. paradoxa, but indentations of Cephth. not quite
equal nor quite half-moon shaped. Pectinations on Psstg. org. longer, and
hairs on Abd. setiform, not leaf-like. L. 750 ju.
In moss. Italy; rare.
2. L. paradoxa (Haller) 1884 Michaelia p., G. Haller in: Arch. Naturg., 0.501.
p. 229 1. 16 f. 58.
Im. Light brown, twice as long as broad. Cephth. each side with
2 equal half-moon-shaped indentations, concave outward, occupying its whole
lat. edge except tip of Rost. anterior of the 2 bordered by a ridge. Psstg.
org. long, setiform, sparsely pectinated on one side. All the hairs except
some of those on the legs leaf-like, mostly fusiform or laceolate. Legs
short, not reaching post, end of Abd. L. over 1 mm.
In decaying Vines. Germany.
*) As the name Michaelia was preoccupied I have renamed Haller's genus in
honor of Dr. H. Lohmann. Berlese's gen. Michaelia (Trombidiidae), in: A. M. S., fasc. 16
nr. 6 & fasc. 57 nr. 8, was published 29. December 1884. A. D. Michael.
76 Hypochthonius
6. Gen. Hypochthonius C. L. Koch
1835 Hypochthonius, 0. L. Koch, C. M. A., fasc. 3 ; 1842 H., C. L. Koch. Ubers.
Arach.. v. 3 p. 109 j 1888 Hypodhonins, Michael, Brit. Orib., v.2 p. 530 j 1896 If.,
Berlese, A. M. S. } Crypt. II p. 24, 25.
Dors, surface of Abd. divided by transverse sutures as if segmented.
Cuticle imperfectly chitinized.
This genus is said by Nicolet (1855, in: Arch. Mus. Paris, -v.7 p. 389), Canestrini
(1885. Prosp. Acarof., v. 1 p. 4) and Banks (1895, in: Tr. Amer. ent. Soc,, r. 22 p. 2) to
be composed of immature creatures, but they are really adult.
5 species.
Key to the species.
Abd. of 2 segments 2.
Abd. of more than 2 segments 3.
. f Segments almost equal. L. 550 750 |u . ... 4. H. rufulus ... p. 77
\ Ant. segment much larger than post. L. 750 |u . 5. H. tectorum . . p. 77
j Abd. of 3 segments, central the narrowest, others about equal 4.
3 < Abd. of 4 segments, the 2 central narrow and equal,
the ant. and post, wider and equal. L. 330 |u 2. H. lanatus . . p. 76
, Psstg. org. long, filiform, curved, strongly pectinated.
j L. 360 |it 3. H. pallidulus . p. 76
j Psstg. org. with short thin peduncle and fusiform hairy
head. L. 180 |U 1. H. brevis . . . p. 76
1. H. brevis Michael 1888 Hypocthonius b., Michael, Brit. Orib., v. 2 p. 539
t. 49 f. 14.
Im. Yellow, dull, rather rough. Cephth. broad Rost. hairs serrated.
Psstg. dorsal; Psstg. org. white, directed backward, with short peduncle
and fusiform hairy head. Intlam. hair white spines. Abd. slightly
arched, truncated anteriorly, rounded posteriorly, divided into 3 segments
by transverse cuts, central segment narrowest, other 2 equal. 2 longitudinal
rows of thick white hairs on Dors, of Abd. and some round periphery Ung.
mond. L. ca. 180 |u.
On sphagnum etc. England.
2. H. lanatus Michael 1885 Hypocthonius L, Michael in: J. R. micr. Soc.,
ser. 2 t?.5 p. 396 t. 7 f.ll j 1888 H.I, Michael. Brit. Orib., v.2 p. 541 t.49 f. 15 22.
Im. Light yellow-ochre, dull, reticulated, slightly pulverescent. 2 low
ridges represent Lam. Rost. hairs, Lam. hair, Intlam. hair and 2 other
pairs of hairs on Cephth. all like bunches of white wool. Psstg.
org. long, straight, directed backward, with rod-like peduncle and fusiform
woolly head. Mdb. with fixed chela having 2 branches, both toothed.
Abd. pyriform, Dors, thereof coarsely reticulated; reticulations about 8 ju. Dors,
of Abd. divided into 4 segments by transverse cuts. 2 central segments
equal, ant. and post, segments about equal. On each segment a row of
colorless serrated spines which usually lie flat but can be erected. Ung.
mond. L. ca. 330 ju.
In wood, thatched roofs etc. England.
3. H. pallidulus C. L. Koch 1835 H. p., C. L. Koch, C. M. A., fasc. 3 f. 20
1888 Hypocthonius p., Michael, Brit. Orib., v.2 p. 537 t.49 f. 1 5.
Hypoehthonius. Phthiracarinae, Hoploderma 77
Im. Light yellow-brown redder at edges of Abel.; slightly polished.
Rost. long, pointed, narrow. Palpi and Psstg. dorsal. Psstg. org. as in
H. rufulus. Abd. almost diamond-shaped truncated anteriorly divided into
3 segments by 2 transverse cuts, central segment much the smallest, other
2 about equal. 4 longitudinal rows of thick white hairs on Dors, of Abd.
and about 5 on each side. Ung. mond. L. ca. 360 u.
In moss. England, Germany, Netherlands.
4. H. rufulus C. L. Koch 1835 H. r., C. L. Koch, C. M. A., fasc. 3 1. 19 j 1888
Hypocthonius r., Michael, Brit. Orib., v. 2 p. 534 t. 49 f. 6 13 i 1855 Leiosoma ovata
Xph., Nicolet in : Arch. Mus. Paris, v. 7 p. 395 t. 2 f. 5.
Im. Yellow -red (tawny) smooth, slightly polished. Cephth. much
narrower than Abd. Rost. pointed, palpi conspicuous. Psstg. dorsal Psstg.
org. very long, almost filiform, but pointed at distal end, doubly curved,
with sparse regularly placed hairs along the ant. edge. Intlam. hair long,
filiform. Abd. shield-shaped, divided into 2 equal segments by a
transverse cut. Middle of Abd. a continuation of the cone of Cephth. with
transverse sulcations, lat. parts broad, thin, semi-transparent blades; 4 longi-
tudinal rows of curved filiform hairs on Dors, of Abd. and 2 at edge of
ant. division of blade. Ung. mond. - - Nph. Like Im., but paler. - - Im.
L. 550750 ju.
In moss. England. France. Germany, Italy, Switzerland.
5. H. tectorum Berl. 1896 Hypocthonius t., Berlese. A. M. S.. fasc. 78 nr. 8.
Im. Varied with yellow-brown, flesh-color and smoky-brown. Psstg.
org. and all hairs on body clavate, setose. Abd. truncated anteriorly,
pointed posteriorly, divided by a transverse line into 2 unequal segments,
anterior the larger. Ung. trid. L. 750 JLI.
In moss, chiefly on roofs. Italy.
Gr. Subfam. Phthiracarinae
1841 Phthiracarea, Perty, Allg. Naturg.. v. 3 p. 874 1885 Subfam. Hoplophoridae,
Berlese in: Bull. Soc. ent. Ital., 0.17 p. 127 1891 H., Canestrini in: Atti 1st. Veneto.
ser. 7 r. 2 p. 718 | 1892 Hoplophorinae, Tronessart in: Rev. Sci. nat, Guest, p. 2 p. 38* \
1896 Phthiracarea, Oudemans in: Tijdschr. Ent.. v. 39 p. 53 | 1896 Hoplophoridae,
Berlese. A. 31. S., Crypt, II p. 14.
Without Ptm. Cephth. inoveably hinged to Abd. and folding down
against its ventr. surface. Dors, surface of Cephth. is covered by a simple,
more or less convex, chitinous plate (As pis), this frequently bears a median
longitudinal ridge (Carina) (Fig. 14 & 15). Ventr. plate not anchylosed to dors.
2 genera. 15 good and 3 doubtful species.
Key to the genera.
Genital and anal covers separate, Ung. mond 1. Gen. Hoploderma . p. 77
Genital and anal covers coalescing or joined; Ung. trid. 2. Gen. Phthiracarus . p. 80
1. Gen. Hoploderma Michael*)
1835 Hoplophora (non Perty. Orthopt,, 1830!), C. L. Koch, C. M. A., fasc. 2 | 1842
H., C. L. Koch, Ubers. Arach., r. 3 p. 116 | 1855 H., Xicolet in: Arch. Mus. Paris,
*) The name Hoplophora is preoccupied. The new name is derived trom oirXov
and bep|Lta. A.D.Michael.
78
V. 1 p. 470 | 1868 H., Claparede in: Z. wiss. Zool, v. 18 p. 507 ! 1877 H., Canestrini
& Fanzago in: Atti 1st. Veneto, ser. 5 v. 4 p. 77 ! 1885 H., Canestrini, Prosp. Acarof.,
v.l p. 9 i 1888 H., Michael, Brit, Orib.. v.2 p. 543 | 1895 JET., Banks in: Tr. Amor. cnt.
Soc., v.22 p. 16 i 1896 H., Berlese, A. M. S., Crypt. II p. 19.
Fig. 14 & is. Hoploderma magnum, open and closed (*/i).
Genital and anal covers separate and squarish. Ung. mond.
8 good and 2 doubtful species.
Key to the good species.
{Aspis with a median carina 2.
Aspis without carina 5.
{Ant. margin of Abd. with a large projection overhanging Cephth. 3.
Ant. margin of Abd. without any projection 4.
( Projection square and shelf-like. L. 0,75 1,6mm 1. H. anomalum
\ Projection rounded, concave, hood-like. L. 800 |U 6. H. pulcherrimum
{Abd. with a median carina its whole length, no
hairs on it; Psstg. org. setiform. -- L. 800 (a 2. H. carinatum
Abd. without carina, numerous large hairs on it.
Psstg. org. falciform. L. 0,9 -1,4mm ... 5. H. magnum . .
{ Abd. smooth - 6.
\ Abd. pitted 7.
j Aspis with a yellow, elliptical depression on each
side. L. 800 \i 4. H. globosum . .
' Aspis without depression. L. 0,5 1,3 mm . . 3. H. dasypus . .
, Pittings fine, Psstg. org. long and filiform -
L. 820 n 7. H. striculum . .
\ Pittings coarse; Psstg. org. short truncated.
L. 900 n 8. H. variolosum .
p. 78
p. 79
p. 79
p. 79
p. 79
p. 79
p. 80
p. 80
1. H. anomalum (Berl.) 1883 Hoplophora anomala, Berlese, A. M. S., fasc. 6
nr. 5 ! 1888 H. a., Michael, Brit. Orib., v. 2 p. 558 t. 51 f. 56.
Im. Resembles H. magnum but Psstg. more conspicuous. Psstg. org.
setiform, somewhat recurved, and a large squarish chitinous shelf-like
projection from ant. margin of Abd. occupying nearly its whole width
and overhanging Cephth. when creature extended. L. ca. 0,75 1,6 mm.
In moss. England, Italy.
Hoploderma 79
2. H. carinatum (C.L.Koch) 1841 Hoplophora carinata, C.L.Koch, C. M. A.,
fuse. 32 t. 9 1887 H. c., Berlese, A. M. S., fasc. 36 nr. 1.
Im. Coarsely granular. Aspis coarsely punctured, with a carina.
Psstg. org. long, thin, slightly fusiform. Abd. with smooth central
carina its whole length, remainder of dors, and lat. surface divided into
concave diamond-shaped spaces by fine ridges ; a raised knob at each angle
of diamond. Genital and anal covers do not occupy the whole ventr. surface.
L. ca. 800 n.
.In moss. Algeria, Germany, Italy.
3. H. dasypus (Ant. Dug.) 1834 Oribates d., Ant. Duges in: Ann. Sci. nat.,
ser. 2 v. 2 p. 47 ? 1883 Hoplophora d., Berlese, A. M. S., fasc. 6 nr. 4 | ? 1885 H. d.,
Canestrini. Prosp. Acarof., v. 1 p. 46 | 1888 H. d., Michael, Brit. Orib. v.2 p. 560 t. 50
f. 8 14 1835 H. decumana, C. L.Koch, C. M. A., fasc. 2 t. 9 ; 1882 H. d., Haller in:
Jahresh.Ver.Wiirttemb., v.38 p. 307 1841 H. lucida, C.L.Koch, C. M. A., fasc. 32
t, 14 1841 H. lentula, C. L. Koch, C. M. A., fasc. 32 t. 16 ; 1855 H. nitens, Nicolet in:
Arch. Mus. Paris, v. 7 p. 472 1868 H. contractilis, Claparede in : Z. \viss. Zool., v. 18
p. 507 | 1877 H. c., Murray, Econ. Ent. Apt,, p. 222 ; ? 1882 H. maxima, Haupt in : Ber.
Ges. Bamberg, v. 12 p. 27 || non 1887 Tritia lentula, Berlese, A. M. S., fasc. 36 nr. 3 [ non
1841 Phthiracarus contractilis, Perty, Allg. Naturg., v. 3 p. 874.
Im. Pale drab to red-brown; smooth, often polished when living;
elliptical, rounded posteriorly. Aspis without carina. Psstg. org.
minute, hardly to be seen without dissection, with short peduncle and
curved lanceolate head. 6 rows of long setiform hairs on Abd. Nph.
Not distinguishable from that of H. magnum. - - Im. L. 0,5 1,3 mm.
In decayed wood. Algeria, England, France, Germany, ? Italy, Netherlands,
Sweden, Switzerland.
4. H. globosum (C. L. Koch) 1841 Hoplophora globosa, C. L. Koch, C. M. A.,
fasc. 32 1. 12 ; 1883 H. g., Berlese, A. M. S., fasc. 6 nr. 3 1895 H. sphaerula, Banks
in: Tr. Amer. ent. Soc., v. 22 p. 16.
Im. Chestnut-brown, smooth. Aspis with a yellow depressed ellipse
each side, but no true carina. Psstg. org. sessile, fusiform. Abd. a
very short ellipse, almost globular, 2 longitudinal rows of long very fine
hairs on Dors, of Abd. L. ca. 800 |u. - - ? if really distinct from H. dasypus.
In moss. Germany, Italy, Netherlands.
5. H. magnum (Nic.) 1855 Hoplophora magna, Nicolet in: Arch. Mus. Paris,
f. 7 p. 472 t, 10 f. 4 | 1888 H. m., Michael, Brit. Orib., v.2 p. 556 t. 50 f. 1 7 1892
H. m., Berlese, A. M. S., fasc. 67 nr. 9.
Im. Brown of. variable depth, dull, coarsely pitted, pits on Abd. about
60 to mm finer on Cephth. Along oval, broader seen from side. Aspis
with strong rough longitudinal carina. Psstg.-org. short, fine,
sickle-shaped. Abd. round posteriorly. Dors, plate much sufflexed on to
ventr. surface. 2 longitudinal rows of long rod-like hairs on Dors, of Abd.
and 2 rows of smaller hairs along each side. - - Nph. Fat, soft, yellowish
white, can scarcely stand, Cephth. without aspis. Abd. very rounded with
mamilliform projection posteriorly. - - Im. L. ca. 0,9 1,4 mm.
Nph. bores in wood. Im. generally in moss. England, France. Germany,
Netherlands.
6. H. pulcherrimum (Berl.) 1887 Hoplophora carinata var. pucherrima.
Berlese, A. M. S., fasc. 36 nr. 10.
80 Hoploderma. Plithiracarus
Im. Like H. carinatum but a large, hood-like, rounded projection from
ant. margin of Abd. overhanging Cephth. L. ca. 550 |u.
Italy.
7. H. striculum (C. L. Koch) 1835 Hoplophora stricula, C. L. Koch. C. 31. A.,
fasi-. 2 t. 10 1855 H. s., Xicolet in: Arch. 3Ius. Paris, r, 1 p. 472 t. 10 f. 5 ! 1888 H. s.,
31 ic ha el, Brit. Orib. : r,2 p. 563 t. 51 f. 7 || non 1896 H. s., Berlese, A. 31. S., fasc. 78
nr. 3.
Im. Pinkish faun-color, dull, semitransparent, chitin thin, strongly
pitted, pits about 100 to mm. Aspis without carina. Psstg. org.
long sickle-shaped. Intlam. hair short rod-like. Abd. elliptical, bearing
8 longitudinal rows of rod-like, blunt-ended, imbricated hairs. L. ca. 520 |u.
H. stricula Berlese is similar, but has a Carina on Aspis and hairs on Abd.
are not imbricated. - - Berlese (1887, A. 31. S.. fasc. 36 nr. 2) says in error, that
H. stricula C. L. Koch is a young form of H. carinata C. L. Koch.
England. France. Germany.
8. H. variolosum (Berl.) 1888 Hoplophora variolosa, Berlese in: Bull. Soc.
ent. Ital., v. 20 p. 218 t. 13 f. 6.
Im. Brown, rough; Aspis and Abd. without carina. Psstg. org.
short truncated. Abd. coarsely pitted all over, ant. margin not
produced forward, 2 longitudinal rows of about 6 long hairs on Dors, of Abd.
Ung. mond. L. ca. 900 |u.
Under bark of trees. Brazil.
H. crinitum (C. L. Koch) 1841 Hoplophora crinita, (.'. L. Koch. C. 31. A.,
fasc. 32 t. 8 | 1841 H. ferruginea, C. L. Koch, C. 31. A., fasc. 32 t. 10.
Possibly the same as H. globulum. but Im. seems rather longer in Abd. and
hairs on Dors, of Abd. longer and more flexible. Im. ,,mittelgross".
Germany. Netherlands 'i
H. quadriseriatum (Haupt) 1882 Hoplophora quadriseriata, Haupt in: Her.
Ges. Bamberg. v. 12 p. 26.
Probably the same as H. globulosum, but Abd. rough, like shagreen, instead
of smooth, and hairs thereon thicker. Irn. ^mittelgross-'-
In moss. Germany.
2. Gen. Phthiracarus Perty
1841 Phthiracarus, Perty. Allg. Xaturg.. v.3 p. 874 ! 1896 P., Oudemans in:
Tijdschr. Ent.. v. 39 p. 53 | 1883 Tritia, Berlese, A. M. S., fasc. 6 nr. 1 1885 T., Cane-
strini, Prosp. Acarof.. c. 1 p. 9 '. 1892 T., Trouessart in: Rev. Sci. nat. Quest, r 2 p. 48* j
1895 T., Banks in: Tr. Amer. ent. Soc.. v. 22 p. 16 ' Hoplophora (part J), Autor. plurim.
Genital and anal covers anchylosed, or connected together, both long
and narrow. Ung. trid.
7 good and 1 doubtful species.
Key to the good species.
J Abd. bearing conspicuous hairs 2.
\ Abd. hairless 5.
J Abd. rounded posteriorly 3.
\ Abd. pointed posteriorly 4.
Phthiracarus 81
, Dors, of Abd. (when creature upright) J / 3 longer than
broad. Psstg. org. setiform. L. 1,5 mm .... 2. P. berlesei . . p. 81
\ Dors, of Abd. twice as long as broad. Psstg. org. very
clavate. L. 700 |u 5. P. longulus . . p. 82
j Psstg. org. filiform. L. 800 |u 1. P. arduus . . . p. 81
4 { Psstg. org. with thin peduncle and flattened expanded
head. L. 480 |u 3. P. canestrinii . p. 81
_ [ Abd. rounded posteriorly. L. 1.1 mm . . . . . . 4. P. contractilis . p. 81
:> \ Abd. pointed posteriorly - 6.
I Psstg. org. with thin peduncle and fusiform head. -
H L. 800 |u 6. P. nudus ... p. 82
I Xo Psstg. org. visible externally. L. 1 mm ... 7. P. retaltecus . p. 82
1. P. arduus (C.L.Koch) ?1841 Hoplophora testudinea, C.L.Koch, C. M. A.,
fasc. 32 t. 11 j 1841 H. ardua, C. L. Koch, C.M. A., fasc. 32 t. 15 1888 H. a., Michael,
Brit. Orib., v. 2 p. 564 t. 51 f. 815.
Im. Light red-brown and yellow-brown, varies in different parts of
creature ; smooth, dull, Abd. sometimes polished, marked with fine undulating
lines, frequently covered with fine dust. Psstg. org. long; upright, fili-
form. Intlam. hair very long; 2 parallel ridges on each side of aspis
running forward from Psstg. Abd. truncated anteriorly pointed posteriorly.
2 long and 2 short rows of long setiform hairs on Abd., and 2 pairs on
Cephth. besides the Rost. hairs and Intlam. hair. Ung. hetd. L. ca. 800 |u.
England, Germany.
2. P. berlesei Michael*) 1883 Tritia decumana, (err., non Hoplophora decu-
mana C.L.Koch 1835!), Berlese, A. M. S., fasc. 6 nr. 2 1885 T. d., Canestrini, Prosp.
Acarof., v. 1 p. 45.
Im. Chestnut-brown, smooth. Psstg. org. setiform. Abd. rounded
posteriorly a few very fine longish hairs near edge. L. ca. 1,5 mm.
In moss. Italy.
3. P. canestrinii Michael**) ?1885 Tritia lentula, Canestrini. Prosp. Acarof.,
v. 1 p. 45 '. 1887 T, I., (err., non Hoplophora lentula C. L. Koch 1841 !), Berlese. A. M. S.,
fasc. 36 nr. 3.
Im. Yellowish to reddish brown, polished, nearly pointed posteriorly.
Psstg. org. with longish thin peduncle and flattened expanded
head; curved and simple posteriorly; straight and pectinated anteriorly.
2 rows of long setiform hairs on Cephth. and Abd., post, part of anal covers
hairy. Ung. very hetd. Central claw thickest. L. ca. 480 |u.
In moss. Italy.
4. P. contractilis Perty 1841 P.O., Perty, Allg. Naturg., v. 3 p. 874 ; 1868,
Perty (in: Claparede) in: Z. wiss. Zool., v. 18 t. 36 f. 15 19 || non 1868 Hoplophora c.,
Claparede in: Z. wiss. Zool., v. 18 p. 507.
*) Berlese identifies this species, I think erroneously, with Hoplophora decumana
C. L. Koch (which is in my opinion a synonym of Hoploderma dasypus); therefore under
the rules which govern ,,Das Tierreich" the name cannot stand although in a different
genus, I have therefore named the species in honor of Prof. Berlese. A. D. Michael.
**) Berlese identifies this species. I think erroneously, with Hoplophora lentula
C. L. Koch, therefore under the rules which govern ,,Das Tierreich" the name cannot
stand, although in a different genus. I have therefore named the species in honor of
Prof. Canestrini. A. D. Michael.
Das Tierreich. 3. Lief.: A.D.Michael, Oribatidae.
82 Phthiracarus. Genera dubia et species dubiae
Im. Chestnut-brown semitransparent ; A b d. 1 o n g , r o u n d e d p o s t e r i o r 1 y.
hairless. Genital and anal covers broad for the genus. (Description
chiefly taken from Perty's figure in Claparede's plate). L. 1,1 mm.
In decayed wood. Germany.
5. P. longulus (C. L. Koch) 1841 Hoplophora longula, C. L. Koch, (.'. M. A.,
fast-. 32 t. 17 | ?1874 H.arctata, Kiley in: Rep. Ins. Missouri, v. 6 p. 52 & in: Tr. Ac.
St. Louis, v. 3 p. 215 ?1895 Tritia glabrata, Banks in: Tr. Amer. ent. Soc., v. 22 p. 16.
Im. Yellow-brown, polished. Psstg. org. short, very clavate, head
almost globular. Abd. very long-shaped, very narrow, thick from ventr.
to dors, surfaces, slightly truncated anteriorly, rounded posteriorly, numerous
long fine hairs on Dors, of Abd., the longest pair on the shoulders. L. 700 ju
(Kiley).
Kiley says his species is the shape of a mussel probably a Phthiracarus neither
of first two authors describe Ung.
In woods. Germany; at roots of vines America, supposed (probably in error)
to be feeding on Phylloxera and correctly on cotton-wood logs.
6. P. nudus (Berl.) ? 1844 Hoplophora laevigata, C. L. Koch, C. M. A., fasc. 38
1. 16 1 1887 Tritia nuda, Berlese. A. M. S., fasc. 35 nr. 21.
Im. Earthy to sooty brown. Almond-shaped, polished, pointed
posteriorly. Dors, surface hairless. Psstg. org. with short thin
peduncle and fusiform head. L. ca. 800 ju.
In moss and dead wood. Italy.
7. P. retaltecus (Stoll) 1891 Hoplophora retalteca, Stoll in: Biol. Centr.-
Amer., Arach. acar. p. 27 t. 15 f. 4.
Im. Like P. nudus, but no Psstg. org. visible externally, and central
Ung. said to be thinner than the 2 outer.
Guatemala.
P. glabratus (Say) 1821 Or ibata glabrata, Say in: J. Ac. Philad.. r. 2 p. 68,
Coll. Writ., -v. 2 p. 16.
Im. Black, spheroidal, somewhat oval, polished. Psstg. org. apparently with
globular head on very short peduncle.
Uncertain genera and species of Oribatidae. *)
Acarus L., em. Latr. 1758 A. (part.), Linne, Syst. Nat., ed. 10 r. 1 p. 615 j
1795 A., Latreille in: Mag. enc.. v. 4 p. 19 (Typus: A. coleoptratus L.).
Acarus coleoptratus L. 1758 A. c., Linne. Syst. Xat.. ed. 10 r. 1 p. 616 1761
A. c., Linne, Fauna Suec., ed. 2 nr. 1973 | 1795 A. c., Latreille in : Mag. enc.. v. 4 p. 19
(Typus generis Acarus L. !).
Acarus piger Scop. 1763 A. p., Scopoli, Ent. Carniol., p. 392.
Probably a Hoploderma.
Acai'us rupestris F. 1794 A. r., J. Ch. Fabricius, Ent. syst., r. 4 p. 433.
*) The following genera and species have been rejected as being either immature
creatures, too imperfectly described for identification, not Oribatidae at all although
called so; or for some other reason. A. D. Michael.
Genera dubia et species clubiae 83
Acarus sp., G. Shaw 1807 A. coleoptratus (err., non Linne 1758!), G. Shaw,
Zool. Misc.. <. 17 t. 714.
Probably a Notaspis.
Belba floridana Banks 1896 B. f., Banks in : Tr. Amer. ent. Soc., v. 23 p. 76.
Probably a Notaspis.
Belba minuta Banks 1895 B. m., Banks in: Tr. Amer. ent. Soc., v.22 p. 1.
Probably a Notaspis.
Carabodes nitens Johnst. 1847 C. n., G.Johnston in: Tr. Berwickshire Club,
u. 2 p. 221.
Probably a Notaspis.
Celaeno aegrota C. L. Koch 1841 C. a., C. L. Koch, C. 31. A., fasc. 32 t. 5.
A Gamasid.
Celaeno coccinea C. L. Koch 1841 C. c., C. L. Koch. C. M. A., fasc. 32 t. 1.
A Nymph.
Celaeno obsoleta C. L. Koch 1841 C. o., C. L. Koch, C. 31. A., fasc. 32 t. 4.
A Nymph.
Celaeno rhodomela C. L. Koch 1841 C. r., C.L.Koch, C. 31. A., fasc. 32 t. 2.
A Nymph.
Cepheus ovalis C. L. Koch 1841 C. o., C. L. Koch, C. M. A., fasc. 32 t. 7.
Cepheus pertusus Haupt 1882 C.p., Haupt in: Ber. Ges. Bamberg, c. 12 p. 27.
Cephus (sic) nitidus Banks 1895 C. n., Banks in: Tr. Amer. ent. Soc., v.22 p. 10.
Probably a Liacarus.
Claviceps Can. & Fanz. 1877 C., Canestrini & Fanzago in: Atti 1st. Veneto,
ser. 5 r. 4 p. 196.
Immature Oribatidae included in Acarini (= Tyroglyphinae), before their life-
histories were known.
Claviceps hirtus Can. & Fanz. 1877 C. h., Canestrini & Fanzago in : Atti 1st.
Veneto, ser. 5 v.4 p. 203.
Claviceps laeviusculus Can. & Fanz. 1877 C. L. Canestrini & Fanzago in :
Atti 1st. Veneto, ser. 5 v. 4 p. 204.
Claviceps ruber Can. & Fanz. 1877 C. r., Canestrini & Fanzago in : Atti 1st.
Veneto, ser. 5 v. 4 p. 205.
Claviceps rugosus L. Koch 1879 C. r., L. Koch in: Svenska Ak. Handl.,
v. 16 p. 135.
A Nymph, possibly of Hermannia convexa.
Claviceps trimaculatus L. Koch 1879 C. t., L. Koch in : Svenska Ak. Handl.,
v. 16 p. 134.
A Nymph, probably of an Oribata.
Damaeus bulbipedata Pack. 1886 D. b., Packard in : 3Iem. Ac. Washington,
v.4 p. 42.
Damaeus curtus Haupt 1882 D. c., Haupt in: Ber. Ges. Bamberg, v. 12 p. 28.
Damaeus onustus C. L. Koch 1844 D. o., C. L. Koch. C. 31. A., fasc. 38 t. 7
Not a species, only coated with dirt.
84 Genera dubia et species dubiae
Eremaeus asperulus Berl. E. a., Berlese. A. M. S., Repert.
Eremaeus lineatus Thor. 1871 E. I, Thorell in : Ofv. Ak. Fb'rh.. v. 28 p. 690.
Hermannia trinebulosa Riley 1885 H. t.. Riley in: Bull.Dep. Agric. Ent., p. 216.
Hoplophora straminea C.L.Koch 1841 H. s., C.L.Koch, C. M. A., fasc. 32 1. 13.
Hoplophora testacea C.L.Koch 1841 H.t., C.L.Koch, C.M.A., fasc. 32 1. 11.
Murcia acaroides C. L. Koch 1835 M. a., C. L. Koch, C. M. A., fasc. 3 t. 22.
A Nymph.
Mureia ephippiata C. L. Koch 1841 M. e., C. L. Koch, C. M. A., fasc, 31 t, 22.
A Nymph.
Murcia fumigata C. L. Koch 1841 M. f., C.L.Koch, C. M. A., fasc. 31 t. 21.
A Nymph.
Notaspis cassideus Herm. 1804 N. c., Hermann, Mem. apt., p. 93.
A . Gamasid.
Notaspis castaneus Herm. 1804 N. c., Hermann, Mem. apt., p. 89.
Notaspis conchaeformis Koll. N. c., Kollar -
Notaspis corynopus Herm. 1804 N. c., Hermann. Mem. apt., p. 89.
Notaspis globulus Koll. N. g., Kollar
Notaspis humeralis Herm. 1804 N. h.. Hermann, Mem. apt., p. 92 1840
N. h., C. L. Koch, C. M. A., fasc. 30 1. 18.
Nothrus bicolor C. L. Koch 1844 N. b., C. L. Koch. C. M. A., fasc. 38 t. 5.
A Nymph.
Nothrus bispinosus C.L.Koch 1840 Kb., C.L.Koch, C.M.A.. fasc. 29 t.24.
A Nymph.
Nothrus canaliculatus C.L.Koch 1840 N. c., C.L.Koch, C. M. A., fasc. 29
t. 7 1842 Carabodes c., C. L. Koch, Ubers. Arach., r. 3 p. 107.
Probably an immature Neoliodes.
Nothrus corticalis Karp. 1893 N. c., Karpelles in: Math, naturw. Ber. I ngarn,
r. 11 p. 86.
Nothrus doliaris C. L. Koch 1840 N. d., C. L. Koch, C. M. A., fasc. 29 t. 5. 6.
A Nymph.
Nothrus ellipticus Haupt 1882 N. e., Haupt in: Ber. Ges. Bamberg, r. 12 p. 22.
Nothrus marinus Banks 1896 N.m., Banks in: Tr. Amer. ent. Soc., r.23 p. 77.
Possibly a Scutovertex.
Nothrus minimus C. L. Koch 1844 N. m., C. L. Koch, C. M. A., fasc. 38 1. 1
A Nymph.
Nothrus ovivorus Pack. 1870 N. 0., Packard in: Amer. Natural., v.3 p. 366.
Nothrus posticus C. L. Koch 1840 N.p., C. L. Koch. C. M. A., fasc. 30 t, 5.
A Nymph.
Oppia hungarica Karp. 1893 0. h., Karpelles in: Math, naturw. Ber. Ungarn,
nil p. 89.
Genera dubia et species dubiae 85
Oribata aspidioti Ashm. 1879 0. a., Ashmead in: Canad. Ent., v. 11 p. 93.
Not an Oribata, probably a Nymph.
Oribata cyclonotus Gerv. 1849 0. c., Gervais in: Gay. Hist. Chile, v. 4 p. 48.
A Nymph.
Oribata demersa Duj. 1842, Dujardin in: Institut, v. 10 p. 316 j 1844 0. d.
Duj.. Gervais in: Walckenaer. Ins. Apt., v. 3 p. 253.
Oribata marina Duj. 1842. Dujardin in: Institut, v. 10 p. 316 j 1844 0. m. Duj.,
Gervais in : Walckenaer, Ins. Apt., v. 3 p. 254.
Oribata notactis Gorv. 1849 0. n., Gervais in: Gay, Hist. Chile, t?.4 p. 48.
Possibly Neoliodes concentricus.
Oribata quadripilis Fitch 1859 0. q., Fitch in: Hep. Ins. X. York v.3 sect. 159.
Oribates angulatus C. L. Koch 1840 0. a., C. L. Koch, C. M. A., fasc. 30 t. 21.
Founded on a deformed or injured type.
Oribates aterrimus C. L. Koch 1844 0. a., C. L. Koch, C. M. A., fasc. 38 t. 10.
Oribates facula C.L.Koch 1840 O.f., C.L.Koch, C. M. A., fasc. 30 t. 17.
According to Berlese same as 0. lapidaria.
Oribates globosus C. L. Koch 1841 0. g., C. L. Koch, C. M. A., fasc. 31 t. 4.
A Notaspis or a Serrarius.
Oribates picipes C.L.Koch 1840 O.p., C.L.Koch, C. M. A., fasc. 30 1. 15.
Oribatella obesa Banks 1895 0. o., Banks in: Tr. Amer. ent. Soc., v. 22 p. 9.
Oribatella perfecta Banks 1896 O.p., Banks in: Tr. Amer. ent. Soc., p. 23 p. 77.
Oxynamba liliputana Karsch 1881 0. 1., Karsch in: Berlin, ent. Z., t\25 p. 37.
Oribatidae? Included in Gamasidae (Oxynamba nomen nov.).
Pelonia Grube 1859 P., Grube in: Arch. Dorpat. Ges., ser. 2 v. 1 p. 477.
Immature creatures.
Pelonia crinita Grube 1859 P.O., Grube in: Arch. Dorpat. Ges., ser. 2 t\l p. 478.
A Nymph.
Pelonia foliosa Grube 1859 P.f., Grube in: Arch. Dorpat. Ges., ser. 2 v. 1 p. 477.
A Nymph.
Pelonia tricuspidata Grube 1859 P. t., Grube in: Arch. Dorpat. Ges., ser. 2
v.l p- 478 j 1863 P. t., Andersen in: Ofv. Ak. Fork, v.20 p. 188.
A Nymph.
Seutovertex concolor Banks 1895 S. c., Banks in: Tr. Amer. ent. Soc., 0.22 p. 11.
Apparently not a Seutovertex, probably a Notaspis (said to be so by Banks
in litera).
Seutovertex pilosus Banks 1895 S.p., Banks in: Tr. Amer. ent. Soc., .22 p. 11.
Apparently not a Seutovertex, probably a Notaspis (said to be so by Banks
in litera).
Zetes cespitum C. L. Koch 1841 Z. c., C. L. Koch, C. M. A., fasc. 31 t. 8.
Zetes dorsatus C. L. Koch 1841 Z. d., C. L. Koch, C. M. A., fasc. 31 1. 15.
A Nymph.
86 Genera dubia et species dubiae
Zetes flavipes C. L. Koch 1841 Z. f., C. L. Koch, C. 31. A., fuse. 31 1. 16.
Zetes gilvulus C.L.Koch 1841 Z.y., C.L.Koch, C. M. A., fasc. 31 t. 17.
A Xotaspis.
Zetes laevigatus C. L. Koch 1835 Z. I, C. L. Koch, C. M. A., fasc. 3 t. 8.
Zetes latirostris C. L. Koch 1844 Z. I, C. L. Koch, C. M. A., fasc. 38 t. 13.
Zetes mortacinus C. L. Koch 1841 Z. m., C. L. Koch, C. M. A., fasc. 31 t. 14.
A Nymph.
Zetes semirufus C.L.Koch 1841 Z. s., C.L.Koch. C. M. A., fasc. 31 t. 7.
Founded on some accidental monstrosity.
Alphabetical Index.
acaroides (Murcia) 84
Acarus 82
Acarus 53
Achipteria . 9
acromios (Xotaspis) 6, 7
acromios (Oribata) 7
acromios (Pelops) 6, 7
acromius (Pelops) 6
acuminata (Murcia) 46
aegrota (Celaeno) 83
aequalis (Notaspis) 46
affinis (Oribata) 24
agilis (Oribata) 20
alata (Galumna) 20
alata (Oribata) 20
alata (Oribata) 21
alatus (Notaspis) 20
alatus (Oribates) 20
alatus (Zetes) 20
americana (Oribata) 18
Ameronothrus 28
Amerus 53
anauniensis (Nothrus) 70
Angelia 62, 68. 75
angulatus (Nothrus) 71, 72
angulatus (Oribates) 85
anomala (Hoplophora) 78
anomalum (Hoploderma) 78
ansatus (Nothrus) 71
antarctica (Oribata) 12
apicalis (Carabodes) 37
Apterogasterea 6
aquatica (Oribata) 18
aquatica (Oribatella) 18
arborea (Oribata) 24
arctata (Hoplophora) 82
ardua (Hoplophora) 81
arduus (Phthiracarus) 81
armata (Oribatella) 19
arrecta (Hermannia) 62
asperulus (Eremaeus) 84
aspidioti (Oribata) 85
aterrimns (Oribates) 85
aurita (Bella) 55, 61
aurita (Oribata) 55
Pag.
auritus (Damaeus) 55
auritus (Damaeus) 56
auritus (Pelops) 7
auritus (Pelops) 8
avenifera (Oribata) 20
badia (Oppia) 46
badius (Oribates) 46
banksi (Nothrus) 70
Belba 42,53
berlesei (Damaeus) 55
berlesei (Oribata) 18
berlesei (Phthiracarus) 81
berlesii (Oppia) 47
bicarinatus (Nothrus) 70
biciliatus (Nothrus) 70
bicolor (Nothrus) 84
bicornis (Neozetes) 27
bicornis (Serrarius) 27
bicostata (Belba) 59
bicostatus (Damaeus) 59
bicristatus (Nothrus) 65
bidentata (Oribatella) 13
bifidatus (Cepheus) 32
bilineatus (Ameronothrus) 29
bilineatus (Scutovertex) 29
bimaculatus (Claviceps) 15
bipilis (Notaspis) 46
bipilis (Nothrus) 72
bipilis (Oppia) 46
bipilis (Oribata) 46
bispinosus (Nothrus) 84
bistriata (Hermannia) 63
bistriatus (Nothrus) 71
bistriatus (Nothrus). 63
biurus (Nothrus) 73
biverrucatus (Nothrus) 71
borealis (Nothrus) 72
brevipes (Cymbaeremaeus) 67
brevipes (Eremaeus) 67
brevis (Carabodes) 37
brevis (Hypochthonius) 76
brevis (Hypocthonius) 76
bulbipedata (Damaeus) 83
bulbipes (Belba) 61
bulbipes (Damaeus) 61
Alphabetical Index
Pag.
kurrowsii (Notaspis) 46
burrows! (Notaspis) 46
caelatus (Scutovertex) 29
calcarata (Oribata) 24
calcarata (Oribates) 24
calcaratus (Oribates) 18, 24
canaliculatus (Carabodes) 84
canaliculatus (Nothrus) 84
canestrinii (Phthiracarus) 81
Carabodes 36
Carabodes 27, 33
Carabodides 1
carinata (Hermannia) 63
carinata (Hoplophora) 79
carinatum (Hoploderma) 79
carinulatus (Damaeus) 61
carinulatus (Eremaeus) 61
cassideus (Notaspis) 84
castaneus (Notaspis) 84
Cdae.no -6
centro-americana (Oribata) .... 21
cephalotes (Carabodes) 39
cepheiformis (Tegeocranus) .... 34
cepheiformis (Tegeocranus) .... 35
Cepheus 31
Cepheus 33
cespitum (Zetes) 85
circumvallatus (Nothrus) 66
cirrosus (Nothrus) 63
Claviceps 83
clavipectinata (Notaspis) 49
clavipectinatum (Dameosoma) ... 49
clavipes (Damaeus) 55
clavipes (Notaspis) 55
climatus (Oribates) 21
climatus (Zetes) 21
clypeata (Oribata) 12
clypeatus (Carabodes) 37
clypeatus (Tegeocranus) 37
coccinea (Celaeno) 83
coleoptratus (Acarus) 82
coleoptratus (Acarus) 83
concavus (Nothrus) 73
concentrica (Oribata) 65
concentricus (Liodes) 65
concentricus (Neoliodes) 65
conchaeformis (Notaspis) . . .' . . 84
concolor (Belba) 56
concolor (Damaeus) 56
concolor (Dameosoma) 56
concolor (Dameosoma) 57
concolor (Scutovertex) 85
confervae (Acarus) 50
conformis (Notaspis) 47
conformis (Oppia) 47
contractilis (Hoplophora) 81
contractilis (Hoplophora) 79
Pag.
contractilis (Phthiracarus) 81
contractilis (Phthiracarus) 79
convexa (Hermannia) 63
convexulus (Oribates) 5
convexus (Nothrus) 63
coracinus (Leiosoma) 40
coracinus (Liacarus) 40
coracinus (Oribates) 40
coriaceus (Carabodes) 37
coriaceus (Carabodes) 37, 39
coriaceus (Cepheus) 37
coriaceus (Tegeocranus) 37
cornuta (Oppia) 46
corrugatus (Ameronothrus) .... 29
corrugatus (Scutovertex) 29
* corticalis (Nothrus) 84
corynopus (Notaspis) 84
crassipes (Hermannia) 64
crassipes (Oribata) 45
craterif'er (Damaeus) 60
crinita (Hoplophora) 80
crinita (Pelonia) 85
crinitum (Hoploderma) 80
Cryptostigmata 1
curtus (Damaeus) 83
cuspidata (Oribata) 21
cyclonotus (Oribata) 85
cymba (Cymbaeremaeus) 67
Cymbaeremaeus 66
cymba (Eremaeus) 67
cynocephalus (Carabodes) 39
cynocephalus (Nothrus) ...... 39
Damaeidae 28. 61
Damaeinae .' 52
Damaeus 53
Dameosoma 42. 53
Damoeidae 52
dasypus (Hoploderma) 79
dasypus (Hoplophora) 79
dasypus (Oribates) 79
decumana (Hoplophora) 79, 81
decumana (Tritia) 81
demersa (Oribata) 85
dentata (Oribata) 23
dentatus (Oribates) 23
dentatus (Tegeocranus) 34
denticulata (Belba) 56
depauperata (Oribata) 13
depauperatus (Oribates) 13
depressa (Oribata) 24
dimera (Nothrus) 66
distincta (Notaspis) 52
distincta (Oribata) 52
doderleinii (Liodes) 66
doderleinii (Neoliodes) 66
doderleinii (Nothrus) 66
doliaris (Nothrus) 84
Alphabetical Index
89
Pag.
dorsalis (Carabodes) 39
dorsalis (Oribata) . . 21
dorsalis (Zetes) 21
dorsatus (Nothrus) 66
dorsatus (Zetes) 85
dugesii (Bella) 60
dugesii (Damaeus) 60
echinatus (Nothrus) 71
edivardsii (Oribata) 13
edivardsii (Oribates) 13
edwardsi (Oribata) 13
elimatus (Oribates) 21
elimatus (Zetes) 21
ellipticus (Nothrus) 84
elongatus (Carabodes) 37
elongatus .(Tegeocranus) 37
emarginata (Oribata) 20
ephippiata (Murcia) 84
ephippiata (Oribata) 13
ephippiatus (Zetes) 13
Eremaeus 42. 66
excisus (Nothrus) 71
exilis (Notaspis) 45
exilis (Oppia) 45
exilis (Oribatula) 45
facula (Oribates) 85
farinosus (Nothrus) 66
farinosus (Pelops) 7
femoralis (Carabodes) 38
femoralis (Oribata) 16
femoralis (Tegeocranus) 38
femoratus (Damaeus) 60
ferruginea (Hoplophora) 80
fimbriatus (Cymbaeremaeus) .... 67
fimbriatus (Eremaeus) 67
fiagellatus (Nothrus) 71
flagellifer (Damaeus) 56
flammula (Oribata) 25
flammula (Oribates) 25
flavipes (Zetes) 86
floridana (Belba) 83
fockeui (Eremaeus) 67
ibliosa (Pelonia) 85
formica (Damaeus) 61
fuligineus (Pelops) 7
fumigata (Murcia) 84
furcatus (Nothrus) 70, 73
fuscipes (Oribata) 13
fuscipes (Oribates) 13
fuscomaculatus (Zetes) 16
fuscus (Oribates) 40
fusifer (Neozetes) 27
fusifer (Oribates) 27
fusifer (Oribates) 27
fusifer (Serrarius) 27
fusigera (Oribata) 23
Galumna 9
Pag.
geniculata (Belba) 56
geniculata (Oribata) 56
geniculatus (Acarus) 53. 56
geniculatus (Damaeus) 56
gibba (Belba) 60
gibbus (Nothrus) 63
gilvipes (Oribata) . 13
gilvipes (Oribates) 13
gilvulus (Zetes) 86
glaber (Nothrus) 71
glaber (Pelops) 8
glabra (Angelia) 71
glabra (Notaspis) 47
glabrata (Oribata) 82
glabrata (Tritia) 82
glabratus (Phthiracarus) 82
glaucina (Oppia) 55
globifer (Leiosoma) 41
globifer (Liacarus) 41
globipes (Belba) 5.7
globipes (Damaeus) 57
globosa (Hoploj)hora) 79
globosum (Hoploderma) ...... 79
globosum (Leiosoma) 41
globosus (Liacarus) 41
globosus (Oribates) 85
globosus (Oribates) 41
globula (Oribata) 14
globulus (Notaspis) 84
globulus (Oribates) 14. 27
gracilis (Oribata) 14
gracilis (Oribates) 14
granulata (Hermannia) 64
granulatus (Carabodes) 38
halleri (Oribata) 21
harioti (Scutovertex) 30
hepatica (Notaspis) 47
hepaticus (Eremaeus) 47
hericius (Tegeocranus) 34
Hermannia 62
hirsuta (Oribata) 25
hirsutus (Pelops) 7
hirtus (Claviceps) 83
homodactyla (Oribata) 25
Hoploderma 77
Hoplophora 77, 80
Hoplophoridae 77
Hoplophorinae 77
horrida (Oribata) 71
horridus (Notaspis) 71
horridus (Nothrus) 71
horridus (Nothrus) 71
humeralis (Notaspis) 84
humeralis (Oribates) 14
hungarica (Oppia) 84
Hypochthonius 76
Hypocthonius ' ... 76
90
Alphabetical Index
intermedia (Oribata) 21
invenustus (Nothrus) 72
juncta (Notaspis) 49
juncta (Oribatula) 49
kornhuberi (Neoliodes) 66
kornhuberi (Nothrus) 66
kiihli (Nothrus) 5
labyrinthicus (Carabodes) 38
labyrinthicus (Tegeocramis) .... 38
lacustris (Notaspis) 49
lacustris ( Scutovertex) 49
laeingata (Hoplophora) 82
laevigatus (Pelops) 7
lafivigatus (Zetes) 86
laeviusculus (Claviceps) 83
lanatus (Hypochthonius) 76
lanatus (Hypocthonius) 76
lanceolata (Notaspis) 50
lanceolatum (Damaeosoma) .... 50
languida (Oribata) 18
lapidaria (Oribata) 14
latipes (Oribates) 16
latipes (Zetes) 16
latirostris (Oribata) 25
latirostris (Zetes) 86
latirostris (Zetes) 25
lativentris (Leiosoma) 41
lativentris (Liacarus) 41
latus (Carabodes) 35
latus (Cepheus) 32
latus (Cepheus) 35
latus (Tegeocranus) 35
Leiodes 65
Leiosoma 40
Leiosomidae 28
lentula (Hoplophora) 79, 81
lentula (Tritia) 79.81
leporosus (Eremaeus) 35
leporosus (Tegeocranus) 35
Leptorchestes . . . 27
Leptorchistis 27
Liacarus 40
licnophora (Notaspis) 47
licnophorus (Notaspis) 47
liliputana (Oxynamba) 85
lineatus (Eremaeus) 84
Liodes 65
Liosoma 40
Lohmannia 75
longilamellata (Notaspis) 50
longilamellatum (Dameosoma) ... 50
longipes (Oribates) 17
longipilis (Leiosoma) 41
longipilis (Liacarus) 41
longiuscula (Notaspis) 52
longiusculus (Zetes) 52
longula (Hoplophora) 82
Pag.
longulus (Phthiracarus) 82
lucasi (Oribata) 22
lucasii (Oribata) 22
lucens (Oribata) 15
lucida (Hoplophora) 79
lucovum (Notaspis) 47
lucorum (Oppia) 46
lucorum (Zetes) 47
maculatus (Ameronothrus) 30
maculatus (Scutovertex) 30
magna (Hoplophora) 79
magna (Oribata) 25
magnum (Hoploderma) 79
malleolus (Nothrus) 66
marginalis (Eremaeus) 67
marginata (Leiosoma) 41
marginatus (Carabodes) .:.... 39
marginatus (Liacarus) 41
marginatus (Tegeocranus) 39
marina (Oribata) 85
marinus (Nothrus) 84
maxima (Hoplophora) 79
Michaelia 7o
microcephala (Leiosoma) 27
microcephalus (Leiosoma) 27
microcephalus (Serrarius) 27
microcephalus (Serrarius) 27
micronychus (Carabodes) 28
micronychus (Leptorchistis) .... 28
micronychus (Zetorchestes) .... 28
microptera (Notaspis) 48
microptera (Oppia) 48
microptera (Oribata) 15
minimus (Eremaeus) 67
minimus (Nothrus) 84
minuta (Belba) 83
minuta (Oribata) 25
minuta (Oribatella) 25
minutus (Cepheus) 68
minutus (Cymbaeremaeus) 68
mirabilis (Belba) 58
mirabilis (Oribata) 23
mirabilis (Oribatodes) 23
moesta (Oribata) 25
mollicoma (Oribata) 15
mollicomus (Oribata) . 15
mollicomus (Oribates) 15
monilipes (Cymbaeremaeus) .... 50
monilipes (Notaspis) 50
monodactyla (Angelia) 72
monodactyla (Oribata) 23
monodactylus (Nothrus) 72
mortacinus (Zetes) 86
mucronata (Oribata) 22
mucronatus (Oribates) 22
Murcia 9. 42
murcioides (Angelia) 75
Alphabetical Index
91
Pag.
murcioides (Lohmannia) 75
imitilus (Nothrus) 71
nana (Hermannia) 64
nanus (Hermannia) 64
nanus (Nothrus) 64
Neoliodes 65
Neozetes 26
nicoletii (Oribates) 19, 21
niger (Carabodes) 39
nigra (Carabodes) 39
nigro-femoratus (Damaeus) .... 57
nigro-femoratus (Nothrus) 57
nitens (Carabodes) 83
nitens (Damaeus) 57
nitens (Hoplophora) 79
nitens (Leiosoma) 41
nitens (Leiosoma) 45
nitens (Liacarus) 41
nitens (Oppia) 57
nitens (Oribata) 41
nitens (Oribata) 19
nitens (Oribata) 45
nitidus (Cephas) 83
nodipes (Belba) 55, 56
nodipes (Damaeus) 55
nodipes (Dameosoma) 55
nodosa (Hermannia) 65
notactis (Oribata) 85
Xotaspidea 1
Notaspidinae 28
Notaspis 42
notata (Oribata) 17
Nothridae 28,52,61
Nothrinae 61
Nothrinae 28, 52
Nothrus 68
Nothrus 62. 65
nuda (Tritia) 82
nudus (Phthiracarus) 82
obesa (Oribatella) 85
oblonga (Carabodes) 38
oblonga (Hermannia) 38
oblonga (Notaspis) 48
oblonga (Oppia) 15
oblongus (Eremaeus) 48
obsoleta (Celaeno) 83
obsoleta (Murcia) 48
occultus (Pelops) 8
ocellatus (Cepheus) 32
ocellatus (Scutovertex) 32
olivaceum (Leiosoma) 42
olivaceus (Liacarus) 42
onustus (Damaeus) 83
onustus (Damaeus) 56
Oppia 42
orbicularis (Oribata) 15
orbicularis (Oribates) 15
Pag.
Oribata 9
Oribatei 1
Oribatella 9,18
Oribates 9. 40, 42
Oribatidae 1
Oribatidae 5
Oribatides . . . . 1
Oribatidi 1
Oribatina 1
Oribatinae 5
Oribatini 1
Oribatodes 9
Oribatoidea 1
Oribatula 42
ornatissimus (Damaeus) 60
ovalis (Cepheus) 83
ovalis (Eremaeus) 30,31
ovalis (Oribata) 19
ovalis (Oribata) 21
ovalis (Oribates) 19
ovalis (Scutovertex) 31
ovata (Leiosoma) 42, 77
ovatum (Leiosoma) 42
ovatus (Liacarus) 42
ovatus (Oribates) . 42
ovivorus (Nothrus) 84
ovulum (Hermannia) 64
ovulum (Nothrus) 64
pollens (Nothrus) 72
palliata (Angelia) 63
palliata (Hermannia) 63
palliatus (Nothrus) 63
pallidula (Oribata) 16
pallidulus (Hypochthonius) 76
pallidulus (Hypocthonius) 76
pallidulus (Zetes) 16
palmicinctum (Leiosoma) 42
palmicinctus (Liacarus) 42
palustris (Angelia) 72
palustris (Nothrus) 72
palustris (Oribata) 25
papillipes (Damaeus) 58
papillosus (Carabodes) 39
papillosus (Oribates) 39
paradoxa (Lohmannia) 75
paradoxa (Michaelitt) 75
parmeliae (Oribata) 23
patelloides (Damaeus) 58
pectinata (Belba) 51
pectinata (Notaspis) 51
pectinatum (Dameosoma) 51
Pelonia 85
Pelops 6
peloptoides (Oribata) 22
peloptoides (Oribates) 22
peltifer (Neoliodes) 65
peltifer (Nothrus) 65
92
Alphabetical Index
perfecta (Oribatella) . .
pertusus (Cepheus) . .
phaeonotus (Pelops) . .
phalangoides (Damaeus)
Phthiracarea
Phthiracarinae ....
Phthiracarus
picea (Hermannia)
piceus (Nothrus)
picipes (Oribates)
piger (Acarus)
pigerrimus (Nothrus)
pileifbrmis (Nothrus)
pilosa (Notaspis)
pilosa (Oppia)
pilosulus (Zetes)
pilosus (Scutovertex)
pilosus (Zetes)
pinnata (Oribata)
piriformis (Oribata)
plantivaga (Notaspis)
plantivaga (Oribatula)
plicata (Celaeno) . .
plicatus (Pelops)
politus (Oribates)
pottinosus (Nothrus)
posticus (Nothrus)
pratensis (Oribata)
Pterogasterinae
pulcherrimaf Hoplophora carinata var
pulcherrimum (Hoploderma) . . . .
pulverulentus (Damaeus)
pulverulentus (Nothrus)
punctata (Oribata)
punctatus (Nothrus)
punctulatus (Cepheus)
punctulatus (Cephus)
punctulum (Nothrus)
punctum (Oribata)
punctum (Oribates)
pusillus (Oribates)
pyrigerum (Leiosoma)
quadracanthus (Nothrus)
quadricarinata (Notaspis)
quadricarinatum (Dameosoma) . . .
quadricornuta (Oribata)
quadricornuta (Oribata)
quadricuspis (Oribates)
quadridentata (Notaspis bipilis var.) .
quadridentata (Oribata)
quadridentata (Oribatella)
quadripilis (Oribata)
quadriseriata (Hoplophora) . . . .
quadriseriatum (Hoploderma) . . .
retalteca (Hoplophora)
retaltecus (Phthiracarus)
reticulata (Hermannia)
Pag.
85
83
9
58
77
77
80
63
63
85
82
70
74
48
48
48
85
48
26
16
45
45
7
7
40
59
84
26
5
i 79
79
58
58
19
65
33
33
5
16
16
13
42
74
51
51
19
18
25
46
19
19
85
80
80
82
82
64
Pag.
reticulata (Oribata) ^li
rhodomela (Celaeno) 83
rileyi (Oribata) 24
riparius (Damaeus) 61
robusta (Oribata) 26
rostratus (Nothrus) 70
rubens (Oribata) 17
rubens (Oribates) 17
ruber (Claviceps) 83
rubra (Murcia) 14
rubra (Oribata) 14
rufulus (Hypochthonius) 77
rufulus (Hypocthonius) 77
rugifrons (Oribata) 22
rugosus (Claviceps) 83
rugulosus (Nothrus) 73
runcinatus (Nothrus) 72
rupestris (Acarus) 82
satellitius (Zetes) 20
scaber (Nothrus) 65
scabra (Hermannia) 65
scaliger (Nothrus) . 65
sculptus (Pelops) 7
sculptus (Scutovertex) 30
sculptus (Scutovertex) 31
Scutovertex I . 28
segnis (Notaspis) 73
segnis (Nothrus) 73
segnis (Oribata) 73
semirufus (Zetes) 86
Serrariinae 26
Serrarius 26
serrata (Notaspis) 48
serratus (Eremaeus) 48
setosa (Oribata) 17
setosa (Oribatella) 26
setosus (Damaeus) 60
setosus (Oribates) 17, 26
sicula (Notaspis plantivaga var.) . . 4.")
sicula (Oppia tibialis var.) .... 45
siculus (Eremaeus ovalis var.) ... 31
signata (Oribata) 26
signata (Oribatella) 26
simile (Leiosoma) 40
similis (Leiosoma) 40
similis (Notaspis) 45
similis (Oribatula) 15
simplex (Nothrus) 75
simplex (Oribata) 26
simplex (Oribata) 21
sinuatus (Nothrus) 71
sordidus (Nothrus) 75
sphaerica (Oppia) 46
Sphaerozetes 9
sphaerula (Hoplophora) 79
sphagni (Oribata) 17
spinifer (Nothrus) 73
Alphabetical Index
spiniger (Xothrus) 73
spinosa (Celaeno) 6
spirofilus (Nothrus) 6.")
splendens (Damaeus) ">1
splendens (Dameosoma) 51
splendens (Notaspis) 51
splendens (Oppia) 51
straminea (Hoplophora) 84
stricula (Hoplophora) 80
striculum (Hoploderma) 80
subterraneus (Oribates) 41
sufflexa (Belba) 58
sufflexus (Damaeus) 58
sulcatus (Nothrus) 5
sylvestris (Angelia) 73
sylvestris (Angelia) 70
sylvestris (Nothrus) 73
tarda (Angelia) 74
tardus (Nothrus) 74
tardus (Pelops) 8
targionii (Angelia) 74
targionii (Nothrus) 74
tecta (Oribata) 20
tecticola (Belba) 58
tecticola (Damaeus) 58
Tectocepheus 33
tectorum (Hypochthonius) 77
tectorum (Hypocthonius) 77
Tegeocranidae 28
Tegeocranus 33
Tegeocranus 36
tegeocranus (Cepheus) 32
tegeocranus (Notaspis) 32
tegeocranus (Oribata) 32
tenuipes (Belba) 59
tenuipes (Damaeus) 59
testacea (Hoplophora) 84
testudinea (Hoplophora) 81
theleproctus (Liodes) 66
theleproctus (Liodes) 65
theleproctus (Neoliodes) 66
theleproctus (Neoliodes) 65
Pag.
theleproctus (Notaspis) 66
theleproctus (Nothrus) 66
theleproctus (Nothrus) 65
tibialis (Cymbaeremaeus) 68
tibialis (Eremaeus) - . . 68
tibialis (Notaspis) 45
tibialis (Notaspis) 45
tibialis (Oppia) 45
tibialis (Oppia) 45
tibialis (Oribatula) 45
tibialis (Oribatula) 45
torulosus (Pelops) 8
torvus (Damaeus) 56
tricuspidata (Pelonia) 85
trigona (Notaspis) 51
trigonum (Dameosoma) 51
trimaculata (Murcia) 17
trimaculata (Oribata) 17
trimaculatus (Claviceps) 83
trinebulosa (Hermannia) 84
Tritia 80
troisi (Amerus) 53
troisii (Amerus) 53
troisii (Belba) 53
truncatum (Leiosoma) 42
truncatus (Nothrus) 70
uraceus (Pelops) 8
variolosa (Hoplophora) 80
variolosum (Hoploderma) 80
variolosus (Cepheus) 33
variolosus (Pelops) 8
velatus (Tectocepheus) 35
velatus (Tegeocranus) 35
venosus (Neoliodes) 66
venosus (Nothrus) 66
ventricosus (Nothrus) 70
verticillipes (Damaeus) 59
vulgaris (Cepheus) 33
Zetes 9,42
Zetorchestes 27
Zetorchestinae . 27
