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Printed by Authority of the 
STATE OF ILLINOIS 


HENRY HORNER, Governor 


DEPARTMENT OF REGISTRATION AND EDUCATION 


JOHN J. HALLIHNAN, Director 


The Peach Tree Borers 
of Illinois ® 


S. C. CHANDLER 


NATURAL HISTORY SURVEY DIVISION 


THEODORE H. FRISON, Chief 


Circular 31 Urbana February 1939 


STATE OF ILLINOIS 
HENRY HoRNER, Governor 
DEPARTMENT OF REGISTRATION AND EDUCATION 


Joun J. HALLinAN, Director 


BOARD OF NATURAL RESOURCES AND CONSERVATION 
Joun J. HALLInAN, Chairman 


WILLIAM TRELEASE, D.Sc., LL.D., Biology 
Henry C. Cow.eEs, Ph.D., D.Sc., Forestry 
L. R. Howson, B.S.C.E., C.E., Engineering 


WitiiaM A. Noyes, Ph.D., LL.D., 
Chem.D., D.Sc., Chemistry 
Epson S. BAsTIN, Ph.D., Geology 


ARTHUR CuTTS WILLARD, D.Eng., LL.D., 
President of the University of Illinots 


NATURAL HISTORY SURVEY DIVISION 


Urbana, Illinois 


SCIENTIFIC AND TECHNICAL STAFF 
THEODORE H. Frison, Ph.D., Chief 


SECTION OF ECONOMIC ENTOMOLOGY 
W. P. Fuint, B.S., Chief Entomologist 
Cc. C. Comrton, M.S., Associate Ento- 


mologist 

M. D. FarRAR, Ph.D., Research Ento- 
mologist 

J. H. Biccer, B.S., Associate Ento- 
mologist 

S. C. CHANDLER, B.S., Southern Field 
Entomologist 

L. H. SHropsuHirE, M.S., Northern Field 
Entomologist 

W. E. McCautey, M.S., Assistant 
Entomologist 

C. J. Weinman, M.A., Assistant Ento- 
mologist 


C. W. KEARNS, Ph.D., Research Fellow 
in Entomology 

Dwicut PowELL, M.S., Research Fellow 
in Entomology 

ArtTHUR E. RITCHER, 
Fellow in Entomology 

R. C. RENpDTORFF, B.S., Research Fellow 
in Entomology 


B.A., Research 


SECTION OF INSECT SURVEY 

H. H. Ross, Ph.D., Systematic Ento- 
mologist 

Cart O. Monr, Ph.D., Associate Ento- 
mologist, Artist 

B. D. Burks, Ph.D., 
mologist 

G. T. RIEGEL, B.S., Assistant Entomolo- 
gist 


Assistant Ento- 


SECTION OF AQUATIC BIOLOGY 
Davip H. THompeson, Ph.D., Zoologist 
GEORGE W. BENNETT, M.A., Limnologist 
D. F. HANSEN, Ph.D., Assistant 
Zoologist 


SECTION OF GAME RESEARCH AND 
MANAGEMENT 


R. E. YEATTER, Ph.D., Game Specialist 
C. T. Buack, M.S., Research Fellow 


SECTION OF WILDLIFE 
EXPERIMENTAL AREAS 


A. S. Hawkins, M.S., Game Technician 


F.C. BELLROSE; JR eos 
Game Technician 


Assistant 


SECTION OF APPLIED BOTANY AND 
PLANT PATHOLOGY 


L. R. TeEnon, Ph.D., Botanist 
J. C. Carter, Ph.D., Assistant Botanist 
G. H. BoewE, M.S., Field Botanist 


SECTION OF FORESTRY 
James E. Davis, M.P., Extension For- 
ester 
LEE E. YEAGER, Ph.D., Forester 


SECTION OF PUBLICATIONS 
James S. Ayars, B.S., Editor 


This paper is a contribution from the Section of Economic Entomology 


(66787—4M—10-38) 


<oeU, 
cums 2 
as 


CONTENTS 


ERE BEACH “BORER 5:6. cme sc elec i ee a ales I 
Beach borer Life History Studies.........7...0....52... 1 
MMO ule nTenremCeas 5.2 Giics aek bho sues oie sine Wlcs wie ae | 
[eores iOMuOn MeiMGhAnCeS tt sae ec sens ow Oe le ee ee 3 
Goumolor the Peach Borer... 2.222 65.0. iy. as eee be oe 4 
Method of Treating for the Peach Borer.. Se ae 
Injury from PDB Crystals and aenement ar Vows 
Sipe GSrrartnse eet seat Seca ena toe vee Neh, Dat a 5 
reno emoln ViOtiinGdGHiyas ie otk ee. te Oe oe 7 
Amounts of PDB Crystals for Trees of Different Ages. 8 
PiKieROle A PlCALION goat ti Aa Jae ele tees eek dc oe 9 
CompaiaisonwOlSeasOMss 4.0 ys. asens cise i ae 10 
BesteOatesworm spring Mreatments, 2.54. . 20.525. 10 
Bes WanesmOneh alludbreatmentcmes ss. 2. eee ey 
Prequency ok Ireatmente.......-. ere: 
Removal of Grass and W aedks Before Taenianent See ey 15 
SMOsiibtLessOnk Dau Cry Stale... oon 2. «60s pees. wees. 15 
SOG! Sw OSHARERSS ngs Aa en eR Rac) WE ia aye 16 
IPG OUINS Tb UUES Ry aniste hfe ary Woes oie ee te oie oo 16 
SONG OurmMaMLiGguids=<*5.9 4 esa ee 10 
Compansonvor-solids and Liquids. 2... 22.45.9255. .2* 21 
Leroy Cite Rete Ry Sen ate ACN. 2s rte on ge nn Be, aA 
Bitectiveness at, Low Temiperatures.............. 21 
INGCESSILy Ore MOUmainge 6)... oa oe es OO 
RGuipmicntaweauiredas a... becuase. Se acl: 24 
fikitmerlineatnire cere. yok Scns a em ae ee a ce 2D 
(CORTES) Ba NY 2 025 6 211 SS 9 gee ne ne es Ny nee ge 26 
Summary of the Solid and Liquid Methods....... aA | 
EE LESSER PEACH BORER... 2-6. soc eee: 28 
PRPC HERO UBLED EL Vase ho Reet eee ae. eA Merah alos ae 29 
DUseIET OVC ASUITES He: Soo Ae crs ttat cta Ais PS eae Bore OEE doe. 30 
Bs eriimen tale WhO arn < aie il Ir ila Sac yas ee ak rly Sas 30 
ManlerOh A nea ii Citra ve a tate se Stet sche ani cs delves hee 8 33 


THE PEACH BARK BEETLE AND THE SHOT-HOLE 


Peach borer, Conopia exitiosa (Say). The peach borer passes the winter as a worm 

or larva, A, which, during the following spring and summer, feeds on the inner bark 

of the tree. In summer it spins a cocoon, B (right), in which it changes to a pupa, 

of which the skin is shown, B (left), and from which it emerges 18 to 30 days later 

as a moth: C, male moth; D, female moth. The female moth lays eggs from which 
larvae hatch. All figures four times natural size. 


The PEACH TREE BORERS 


of Illinois id S. C. CHANDLER 


sidered the most important insect enemy of the peach tree 

in Illinois. Although easily controlled, it is seldom eradi- 
cated and in this state is found to some extent in all peach 
orchards three years of age or older. Even orchards one and 
two years old may become seriously infested. 


? NHE peach borer, Conopia exitiosa (Say), is generally con- 


Peach Borer Life History Studies 


Peach borers pass the winter as whitish larvae, one-quarter 
inch to slightly over an inch in length. These worms are usually 
found most abundantly at the base of peach trees just below the 
ground line, but they may be found as much as 10 inches above 
or below this line. The presence of borers 1s indicated by masses 
of gum, mixed with frass or sawdust, exuding from the tree 
Erumkss te. 1. 

During the spring and summer, the worms (frontispiece) 
feed on the inner bark of the trees. They become full grown at 
any time between the middle of June and the first of September. 
They then spin cocoons of silk, incorporating in them bits of 
frass. In these cocoons they change to the brown pupal stage 
(frontispiece), emerging from 18 to 30 days later as clear-winged 
moths (frontispiece). 

The moths are in appearance similar to large wasps. The 
average life of the moths is six days, during which time the 
female, fig. 2, lays from 400 to 1,000 eggs. The eggs are placed 
singly on or around the base of the trunks of peach and related 
trees. In from 7 to 10 days, small white worms with brown 
heads hatch from these eggs. The worms crawl over the outer 
bark and, on finding a crack, they rapidly gnaw their way into 
the inner bark. Here they continue to feed until cold weather. 
There is but one brood a year. 


Moth Emergence 


Our life history studies have been concerned largely with 
moth emergence because of its direct bearing upon proper 
times of treatment for borers. For a period of six years, from 


[1] 


2 ILLINnoIs NATURAL HistoRY SURVEY CIRCULAR 31 February 


1924 to 1929, inclusive, emergence records were kept in a num- 
ber of orchards in the vicinity of Carbondale. During the early 
part of this period, moth-tight cages were constructed around 
the base of infested trees to catch the moths as they emerged. 


> 


Fig. 1.—Shown here at 
the base of a tree is the 
exudate (indicated by 
knife) resulting from 
peach borer attack. 
Frass and gum at the 
base of a peach tree 
are indications that 
peach borers are at 
work or have been at 
work on the inner bark. 


Fig. 2.—Adult female 
of the peach borer, 
natural size, on white 
arrow that points to the 
exudate at the base of 
the tree and the pupal 
skin from which the 
moth has emerged. 
The average life of the 
moth is six days, dur- 
ing which the female 
lays from 400 to 1,000 
eggs. There is but one 
brood each year; moths 
emerge between late 
June and early Oc- 
tober. 


Later, the daily emergence record was made by noting the 
number of fresh pupal skins at each examination. 

An examination of these data shows very little variation 
from year to year, and it seems best to combine the data from 
the six years, presenting them in the form of a graph. This 
necessitates grouping the records. In fig. 3, graphically re- 
cording emergence of peach borer moth from 1924 through 1929, 
the records are shown in weekly periods, from the last week in 
June until the middle of October. This graph shows emergence 
starting late in June and ending October 10. In the period 
covered by the graph, the peak was reached in the last week of 


1939 CHANDLER: PEACH TREE BORERS 3 


August. From the week ending August 22 to that ending Sep- 
tember 19, 67 per cent of the emergence took place. 


Location of Entrances 


Another phase of our life history studies which has a 
practical aspect relates to the amount of frass and exudation 
visible without the removal of any soil from the base of the 
tree. Growers frequently follow the practice of treating only 
those trees that show gum and frass, commonly called borer 
signs. The purpose of this jg 
phase of our study was to 
discover whether infestation 
by peach borers could be 
detected by superficial ex- 
amination of the tree. 


N 
(eo) 


Fig. 3—Graph illustrating composite 
records of peach borer emergence in 
orchards near Carbondale, 1924-29. 
The records for the six years are com- 
bined and shown in weekly periods. 
Emergence is shown as starting late 
in June and ending October 10; the 
peak as being reached the last week 
in August. The greater part of 
emergence, 67 per cent, is shown 
taking place between August 22 and 
September 19. The recommenda- 
tions contained in this circular for 
control of peach borers are based 


fo) 
(eo) 


h 
(e) 


8 


iy) 
[e) 


NUMBER OF PEACH TREE BORER MOTHS EMERGING FROM IO TREES 
Hn 
(e} 


upon these records because of the Ke 
direct bearing of moth emergence 
upon proper times of treatment for fo) 
the borers. 254 Il 1825 1 8 15 2229 5 |2 19 26 3 10 


JUNE JULY AUG. SEPT. OCT. 
FOR WEEKS ENDING 


In the fall of 1934, 50 trees, 10 in each of 5 orchards, which 
at the surface of the soil showed no signs of borers, were examined 
to determine the degree of infestation. 

Of the 50 apparently uninfested trees, borers were found in 
29. The borers, 107 in number, were distributed as follows: 
First orchard, 48 borers divided among 10 trees; second orchard, 
33 borers divided among 9 trees; third orchard, 2 borers divided 
between 2 trees; fourth orchard, no borers in any of the 10 
trees; fifth orchard, 24 borers divided among §& trees. 

It is evident from this investigation that presence of borers 
frequently cannot be detected by surface examination. How- 
ever, we found that, in most trees harboring borers, removing 


4 ILLinoris NATURAL History SURVEY CIRCULAR 31 February 


an inch or less of soil would have revealed the presence of gum 
and frass. 


Control of the Peach Borer 


Since the discovery in 1919 of the value of paradichloro- 
benzene,* commonly called PDB, for the control of the peach 
borer, much experimental work has been done by entomologists 
of Illinois and other states and of the United States Bureau of 
Entomology and Plant Quarantine on the many important 
details of the operation known to the peach grower as gassing 
the borer. In Illinois, control tests began in 1920, and almost 
every year since some studies for the control of this insect have 


Fig. 4.—Application of PDB for control of the peach borer. Left, the ring of PDB 
around tree trunk on leveled ground. Right, the mound around the base of the 
treated tree. 


been made. From time to time some of the information gained 
has been given out in the form of articles, state horticultural 
society reports and unpublished talks at fruit growers’ meet- 
ings. It is the purpose of this publication to summarize the 
results of our peach borer studies and bring them up to date, 
and to include information on the other borers of the peach 
tree, the lesser peach borer, bark beetle and shot-hole borer. 


Method of Treating for the Peach Borer 


The commonly recommended method for gassing the peach 
borer is as follows: Scrape earth from around the trees suffi- 
ciently to allow spreading of the PDB crystals. Apply the 
crystals in the form of a ring far enough from the trunk to 
prevent their touching the bark (cover and fig. +). Use from 


*As the name paradichlorobenzene (the chemical is sometimes called by growers ‘‘Painless Death 
to Borers’’) is difficult to remember and pronounce, the abbreviation PBD has become common and 
will be used hereafter in this publication. 


1939 CHANDLER: PEACH TREE BORERS 5 


one-fourth to one-half ounce PDB crystals on one- and two- 
year trees; up to 2 ounces on large, old trees. Recommended 
amounts for trees of various ages are given on page 8. Cover 


Se 
Pr «40 


Fig. 5.—Left, peach tree with mound sufficiently large for PDB crystals. Right, 
peach tree with mound needlessly large. 


the PDB with four to six spadesful of earth and tamp the earth 
down with the back of the spade. 

In our years of observation of growers’ treatments and 
in our own tests we have found that successful control of borers 
with crystals is obtained with much less mounding than some 
growers think necessary. On trees 10 years old, five or six 
spadesful of earth are required to go around the base. Control 
is not helped by adding more earth, nor should any appreciable 
reduction be made in the amount used. See fig. 5. 


Injury from PDB Crystals and Treatment of Young Trees 


Much experimental work on peach borer control has been 
done because of the fact that PDB may, under certain cir- 
cumstances, injure peach trees. Typical injury is shown in 
fig. 6. It may consist of only a slight flecking of the outer bark 
layers, or of a coalescing of the flecks into dark spots. The spots 
may be small, or they may be so large that they entirely girdle 
the tree. They may affect only outer bark layers or they may 
extend into the cambium. 

The subject of injury to young peach trees by crystaline 
PDB has been of little concern to Illinois growers. In 1920, 
1921 and 1922, tests were conducted in a number of places in 
southern Illinois on one-, two- and three-year old trees to study 
this question, with the result that no injury was observed so 
long as the crystals were not allowed to touch the bark. Asa 


6 ILLINOIS NATURAL HisTORY SURVEY CIRCULAR 31 February 


result of these’ tests we suggested in Circular 8* the use of 
three-fourths ounce of the material on trees as young as two 
years old and one-half ounce on younger trees, but we withheld 


Fig. 6.—Injury from PDB (paradichlorobenzene) crystals, ranging from few flecks, 
A, at the left side of the part of tree that has been cut, to blackened areas on the 
right side, B. 


unqualified recommendations until further experimental work 
could be done. In Circular 267, published in 1935, we suggested 
the use of one-half ounce on trees one full year old. 

In the fall of 1937, tests conducted jointly by the Illinois 
Natural History Survey and the United States Bureau of 
Entomology and Plant Quarantine on trees of different ages 
showed that the younger the trees the greater was the proba- 
bility of injury. The 5- and 13-year old trees used in the test 
were not injured. On the other hand, under circumstances of 
rather high soil temperatures, 20 per cent of the two-year old 
trees treated with one-fourth ounce PDB showed spots which, 
though small, penetrated into and severely injured the cambium. 

With nursery stock, 20 per cent of two-year rootstocks and 
100 per cent of all June buds were severely injured by one-eighth 
ounce PDB. From the tests made it appears unsafe to use 
PDB crystals on nursery stock. 

We are still of the opinion that orchard trees one full year 
old or older can be treated with comparative safety. For 15 
years, one- and two-year old trees in Illinois have been treated 
with PDB crystals, and in the close contact which we have had 
tore Sees See roe St Chanel: The Peach Borer and Methods of Control. Illinois Natural 


+Chandler, S. C., and W. P. Flint. Insect Enemies of the Peach in Illinois. Illinois Natural 
History Survey Circular 26. 1935. 


1939 CHANDLER: PEACH TREE BORERS if 


with the growers of the state we have never heard of or observed 
any serious injury to or loss of trees from PDB correctly applied. 
Undoubtedly, some injury takes place at times, but less than 
might occur if the trees were left to the borers. 

It is our suggestion that growers continue treating young 
trees as in the past but that they consider first the need for 
treatment. Many orchards are two or three years old before 
showing any appreciable infestation, and growers should make 
examinations to ascertain that treatment is needed before it is 
apphed. 


Removal of Mounds 

A number of experiment stations advise the removal of 
mounds from young trees in from three to six weeks after treat- 
ment, because of the assumption that gas will become concen- 


: e hig ¥: 


Fig. 7.—Removing mounds. This operation should be completed by the first of July. 


trated in the pores of the soil as increasing amounts of it are 
evolved from the PDB crystals. We have felt that under IIli- 
nois conditions sufficient gas will escape to prevent a damaging 
concentration. Removal of mounds shortly after treatment is 
almost never practiced in Illinois, and over a long period of 
years we have detected no ill effects from leaving the mounds 
until the following spring. 

In the spring or early summer, however, mounds made the 
previous year should be removed, fig. 7. A large proportion of 
borer eggs are laid on the tree trunks at the ground line, and 
many young borers enter the bark at the same place. To destroy 


8 ILLINOIS NATURAL HiIsTORY SURVEY CIRCULAR 31 February 


ae wes BS /- 
Fig. 8.—Peach trees ranging in diameter from 12 inches to 17 inches are common in 


old orchards in Illinois and require from 114 to 2 ounces of PDB crystals per tree for 
peach borer control. 


eggs and borers with the usual PDB mounding treatment, 
successively higher mounds must be made each year, unless the 
ground around the trees is leveled before the moths begin laying 
eggs. Fig. 3 indicates that, in the peach growing region of 
southern Illinois, moths begin emerging the last of June. There- 
fore, growers have until the first of July to remove mounds made 
the previous year. This gives them the opportunity to include 
mound removal as a part of their spring disking and hoeing, 
after first growth of grass and weeds has started. 


Amounts of PDB Crystals for Trees of Different Ages 


Fear of possible injury to trees has largely determined 
amounts of PDB crystals recommended. In the first years of 
our experimental work, we treated one-year old trees with as 
much as three-fourths ounce PDB per tree on several occasions 
without loss of a tree or even observable injury. Two tests with 
114 ounces per tree on two-year old trees gave similar results. 
Further tests, however, have shown that much smaller amounts 
of PDB give borer control. 

Basing our recommendations on data we have from Illinois 
and other states, we suggest the use of one-fourth ounce PDB 
crystals on trees one full year old, one-fourth to one-half ounce 
on two-year old trees and three-fourths ounce on trees three to 
five years old. 

Amounts used on older trees should be proportional to the 
circumference of the trunks. For most bearing trees, | ounce 
per tree is sufficient, but in many orchards 1% ounces are 
needed. We have in Illinois at the present time many peach 
orchards 12 to 20 years old with trees having diameters of 12 to 


1939 CHANDLER: PEACH TREE BORERS 9 


17 inches, fig. 8. A test conducted on 13-year old trees revealed 
that as much as 2 ounces of PDB may be necessary in some 
cases. Results of the test are given in table 1. 


Table 1.—Results of test in which various amounts of PDB were used on 13-year 
old trees treated at Centralia, October 19, 1922, and examined November 10, 1922. 


Amount PDB TREES BORERS BORERS TOTAL PER CENT OF 
PER TREE TREATED ALIVE DEAD Borers Borers DEAD 

1 ounce 10 4 23 27 85.2 

116 ounces 10 5 47 52 90.4 

2 ounces 10 1 59 60 98.3 


Accuracy 1s important in the application of PDB crystals. 
Such measures as spoonful or handful are inaccurate and may 
result in damage to trees or ineffective treatment. Measuring 
cups that hold an ounce of PDB crystals are supplied by in- 
secticide dealers. These may be used when an ounce is the 
specified dosage, or they may form the basis for the selection 
of other measures holding the exact amount of crystals pre- 
scribed. 


Time of Application 

Fall is the usual time of treating for the peach borer, but 
we have many inquiries from growers who have discovered 
after it is too late for fall treatment that they have a serious 
infestation. They wish to know whether they should treat in 


Fig. 9.—Method of obtaining results of peach borer tests; soil removed and worms 
cut out with knife. This slow and laborious job was the only method of peach borer 
control before the introduction of PDB. 


10 ILLINOIS NATURAL HisTORY SURVEY CIRCULAR 31 February 


the spring or wait until the following fall. Data pertinent to 
these inquiries have been collected, fig. 9. 

Comparison of Seasons.—Table 2 gives a summary of all 
our fall and spring tests in which PDB crystals were applied in 
the recommended amounts and at times and under conditions 
of soil temperature which would be considered advisable for 
orchard treatment. Data on those treatments applied under 
unusual conditions have been omitted. 

This table shows that spring treatments have given from 
77.8 per cent to 95.2 per cent kill and have averaged 83.4 per 
cent. The fall treatments have ranged from 85 per cent to 100 
per cent kill and have averaged 94.2 per cent. The variation 
over a period of 8 springs and 14 falls 1s shown in table 2. 


Table 2.—Results of spring and fall treatments with PDB crystals for peach 
borer in Illinois. This table includes data on all tests in which recommended 
amounts of PDB were applied under normal orchard conditions. 


SPRING APPLICATIONS FaLL APPLICATIONS 

YEAR Trees Total Per Cent of Trees Total Per Cent of 

Treated Borers Borers Dead ‘Treated Borers Borers Dead 
1920 20 162 717.8 40 47 100.0 
1921 90 93 81.7 130 127 98.4 
1922 30 21 OS? 30 139 92.8 
1923 20 36 83.3 60 120 85.0 
1924 — — -—-— 39 78 94.9 
1925 -— ae —— 30 66 87.9 
1926 10 19 84.2 20 88 98.9 
1929 -— = _— 20 23 95.7 
1930 — —- —— 15 1 100.0 
1932 --- = -— 20 49 98.0 
1934 —- —- -_—- 20 S2 93.8 
1935 10 41 92.7 20 56 98.2 
1936 10 20 95.0 70 144 93.8 
1937 25 55 te) 28 90 95.6 
All years 215 447 83.4 542 1000 04.2 


Best Dates for Spring Treatments.—Spring treatments 
should be made as early as temperature conditions are favorable. 
Temperature records kept in connection with tests indicate 
that adequate control can be obtained when soil temperatures 
in the mounds at the level of the material, fig. 10, are from 55 to 
60 degrees for a few days. More rapid volatilization of PDB 
crystals occurs when the temperature is somewhat higher. To 
the grower, however, this information is not so important as an 
answer to his question of when to treat. Date-of-treatment tests 


1939 CHANDLER: PEACH TREE BORERS not 


Fig. 10.—Soil thermograph and thermometer used in determining temperatures suit- 
able for volatilization of PDB crystals. 


were conducted in southern Illinois for a three-year period. A 
summary of these tests is given in table 3. 

Although not extensive, these tests indicate that treat- 
ments with PDB crystals made in southern Illinois in April 


Table 3.—Results of tests made to determine best spring dates for applying 
PDB for peach borer control in southern Illinois. These results indicate that in the 
latitude of Carbondale PDB crystals are consistently effective as early as the first 
of May. 


TREES BORERS BORERS ToTAL PER CENT OF 
DATE OF TREATMENT TREATED ALIVE DEAD Borers’ BoreERS DEAD 
March 31, 1920 10 42 12 54 DD) 2 
April 15, 1920 10 30 15 45 3568 
May 1, 1920 10 18 66 84 78.6 
May 15, 1920 10 18 60 78 76.9 
Check, no treatment 10 61 6 67 8.9 
April 14, 1921 10 11 0) 11 0.0 
May 2, 1921 5 0 5 5 100.0 
May 17, 1921 15 1 15 16 93.7 
April 15, 1922 10 0 3 3 100.0 
May 1, 1922 10 0 8 8 100.0 
May 15, 1922 10 1 9 10 90.0 
Check, no treatment 10 14 1 15 6.7 


12 ILLINOIS NATURAL HistoRY SURVEY CIRCULAR 31 February 


give unsatisfactory results but that those made the first of May 
and later are relatively effective. Spring treatments, however, 
cannot be expected to give as good results as fall treatments. 

Best Dates for Fall Treatments.—Two factors determine 
optimum times for fall treatments: (1) date of latest entrance 
by the borers and (2) soil temperatures. These are related to 
two questions which growers ask: (1) How early should we 
treat? (2) How late should we treat? 

The moth emergence chart, fig. 3, shows that in southern 
Illinois moths are still emerging in large numbers as late as 
September 19, in moderate numbers as late as September 26 
and in small numbers as late as October 3. Since all eggs are 
not laid for some days after the moths emerge, it is evident that 
borer attack in southern Illinois continues into October. The 
presence of tiny borers always found late in October substanti- 
ates this statement. 

In an effort to learn whether borer attack in southern 
Illinois orchards continues into October, trees of a Carbondale 
orchard were treated at different dates in the late summer and 
fall of 1923. Table 4 gives the results of these tests. It brings 
out the fact that, even as late as September 15, the treatment 
failed to kill 9.1 per cent of the borers which entered above the 
mound, but that no live borers were found following the October 
1 treatment. 


Table 4.—Total number of borers found and live borers found above mounds 
in treated orchard near Carbondale, 1923. 


PER CENT OF 


Date OF TREES NUMBER OF BORERS FOUND Bosna 
TRE ATEN fikEanED Total Above Mound MounpD 
August 18 10 7 4 Sel 
September 1 10 iil 7 63.6 
September 15 20 11 1 9.1 
October 1 20 3S) 0 0.0 
October 13 20 ia 0 0.0 


We recommend that growers in southern Illinois do not 
treat earlier than September 25 and that, preferably, they wait 
until October 1. 

The question of how late to treat probably could be 
answered, as already indicated, by reference to soil tempera- 
tures, 55 degrees F. being considered as about the minimum for 
effective volatilization of the crystal PDB. There are at times 


1939 CHANDLER: PEACH TREE BORERS 13 


some difficulties in interpreting soil temperature records, and 
they are not generally available. So the grower is desirous 
of knowing what calendar dates he can depend upon for effec- 
tive treatment. 

Tests conducted near Carbondale over a period of eight 
years show that treatment is consistently effective in southern 
Illinois as late as October 15. A summary of the data on this 
point, giving number of borers concerned and annual fluctuation 
in control, is presented in table 5. 

Table 5.—Results of tests made to determine best fall dates for applying PDB 


for peach borer control in southern Illinois. These results indicate that in the 
latitude of Carbondale PDB crystals are consistently effective as late as October 15. 


TREES BORERS BORERS TOTAL PER CENT OF 
DaTE OF TREATMENT TREATED ALIVE DEAD Borers Borers DEAD 
October 14, 1920 10 0 15 lS) 100.0 
October 17, 1921 10 0 5 5 100.0 
October 12, 1922 10 0 5 5 100.0 
October 19, 1922 20 1 59 60 98.3 
October 13, 1923 20 14 63 77 81.8 
November 1, 1924 10 D, 27 29 93.1 
October 24, 1925 20 4 39 43 90.7 
October 13, 1932 20 1 48 49 98.0 
October 15, 1937 8 3 34 37 91.9 
All treatments 128 25 295 320 92.2 


Differences in weather between southern and central Ili- 
nois indicate that the most favorable dates for treatments in 
the latitude of Centralia are between September 20 and October 
5. In peach districts north of Centrala, treatments should be 
made between September 15 and October 1. 

Every year we find numbers of growers desirous of treating 
later than October 15. They inquire how much later they can 
successfully treat. To answer this question, treatments have 
been made over a period of years as late as the first week of 
November. A summary of these data is given in table 6. 

In the tests recorded in table 6, the 1920 treatments were 
decidedly ineffective. In 1923 no treatments were made later 
tian October 13, but as the results of the treatment made at 
this date were only fair it 1s safe to assume that later treatments 
would have given even lower rates of kill. The results in 1924 
and 1937 were satisfactory, though not so good as would have 
been obtained under higher soil temperatures. 

From the data we have, we conclude that, in about three 
out of five years, satisfactory control of peach borers can be 


14 ILLiInois NATURAL HIsTORY SURVEY CIRCULAR 31 February 


Table 6.—Results of tests with PDB crystals to determine effectiveness in 
southern Illinois of treatments for peach borer control made as late as the first week 
of November. 


TREES BORERS BORERS TOTAL PER CENT OF 
DATE OF TREATMENT TREATED ALIVE DEapD Borers’ Borers DEAD 
November 1, 1920 10 16 10 26 38.5 
November 3, 1921 10 0 6 6 100.0 
October 13, 1923 20 14 63 Ha) 81.8 
November 1, 1924 10 2 27 29 93.1 
November 1, 1937 7 2) 22 24 91.7 


obtained by PDB crystals as late as November 1 in the latitude 
of Carbondale. We have made no similar tests farther north. 


Frequency of Treatment 


A very practical question asked by growers relates to 
frequency of treatment. For two successive years, 30 orchards 
were examined. These fell into three groups: One group was 
treated one year before examination; one, two years before; and 
one, three years before. In each orchard, 100 trees were ex- 
amined. The test involved in all 6,000 trees located in 60 
orchards. Table 7 gives the results of these surveys. 


Table 7.—Borer-infested trees per 100 in orchards treated one, two and three 
years before examination. 


INFESTED TREES INFESTED TREES AVERAGE FOR 
Trees Last TREATED PER 100 IN 1933 Per 100 In 1934 THE Two YEARS 


One year before 6.2 14.3 10.3 
Two years before 36.1 48.9 4225 
Three years before 54.6 S6mo 55.6 


This table indicates that on the average an orchard not 
treated for two years will have four times as many borer-infested 
trees as an orchard untreated for one year. The infestation does 
not continue to increase at the same rate for the years following 
the second, but trees untreated for three years have an accumu- 
lation of unhealed injury not indicated in these data. 

We conclude that if a. grower wishes to economize by de- 
creasing the number of peach borer treatments, he could proba- 
bly omit treatments for one year without much permanent 
damage to his trees, since he would kill most of the borers the 
following fall before they do considerable feeding. However, 
we believe he would be very unwise to omit treatment for two 
successive years. 


1939 CHANDLER: PEACH TREE BORERS 15 


A practice which has considerable merit and is economical 
is known as ‘‘treat a year and patch a year.’ Growers who 
follow it treat their trees thoroughly every other year. In alter- 
nate years they patch; that is, they treat only those trees that 
show, above the ground line, indications of borer infestation. 


Removal of Grass and Weeds Before Treatment 


Early directions for borer treatment included careful 
cleaning of the ground around the base of the tree before apph- 
cation of the crystals. As a result of these directions, many 
growers have spent and some now spend altogether too much 
time on this cleaning operation. Soon after the introduction of 
PDB into Illinois for borer control, tests were begun to de- 
termine the need for removal of grass and weeds or for prepar- 
ation of the soil. They are summarized in table 8. 


Table 8.—Results of tests to determine need for removal of grass and weeds 
prior to treatment with PDB crystals. 


Grass REMOVED Grass Not REMOVED 


Number Total Per Cent Number Total Per Cent 


TIME OF of Trees Number of Borers of Trees Number of Borers 
TREATMENT Treated of Borers Dead Treated of Borers Dead 
Fall, 1922 6 3 100.0 6 2, 100.0 
Spring, 1923 10 15 80.0 10 21 5), d/ 
Fall, 1923 30 87 82.8 30 33 90.9 
Fall, 1924 10 27 96.3 10 22 95.5 
Fall, 1925 10 60 98.3 10 28 100.0 
Fall, 1932 10 24 100.0 10 25 96.0 
Fall, 1934 10 11 100.0 10 16 81.3 
All treatments SO 227 Oil 2 SO 147 92.5 


These tests, extended over seven seasons, show conclusively 
that the removal of grass and weeds does not increase the control 
of the borer. The only reason for the operation is to facilitate 
the spreading of the crystals. The trees used in the plots in 
which the grass was not removed had in most cases a consider- 
ably thicker growth of grass around the base than is found in 
the average orchard and were selected because of this fact. 
Fig. 11 shows the extent of the growth in a typical orchard used 
in the test. 


Substitutes for PDB Crystals 


Four characteristics are sought in any substitute for PDB 
crystals: First, low cost; second, safety on young trees; third, 


16 ILLINOIS NATURAL HisToRY SURVEY CIRCULAR 31 February 


Fig. 11.—Typical tree with excess growth of grass and weeds around base as used in 
grass removal tests. 


efficiency; fourth, effectiveness at low temperatures. We con- 
sider the first two characteristics the most important of the four. 
Probably the hope of finding a safer substance has been the 
greatest incentive in the search for a new material over the 
country at large. 


Although Illinois growers have little fear of serious injury 
on young trees, there is real need for a safe material for use on 
nursery stock. Certainly some injury from PDB crystals occurs 
under certain circumstances to Illinois orchard trees. If a ma- 
terial could be found combining all or even most of the desirable 
characteristics suggested, a real contribution to peach borer 
control would be made. 

Solid Substitutes.—Four solid materials were tested during 
the seasons of 1925 to 1930 inclusive, all of them proposed by 
chemical or insecticide companies as possible inexpensive sub- 
stitutes for PDB. None proved to be of value. 

Liquid Substitutes.—Oliver I. Snapp of the United States 
Bureau of Entomology and Plant Quarantine, in tests over a 
period of years in Georgia, has shown that applications of cotton- 
seed oil emulsion impregnated with PDB may be sprayed around 


1939 CHANDLER: PEACH TREE BORERS 17 


the base of peach trees with the probability of less injury from 
it than from crystal PDB. His results show at least as good con- 
trol with this combination as with the crystal PDB. From time 
to time other liquids, not containing PDB, have been suggested 
and tried. 

In Illinois we have tested mineral oils as carriers of PDB 
and have favored them because of their cheapness and avail- 
ability. We have tested two materials not containing PDB, 
dichloropentane and Scalecide, suggested by manufacturers of 
insecticides. We have also tested soybean oil as a carrier of 
PDB. In table 9 are summarized our tests with these materials 
for the falls of 1934 and 1935 and the intervening spring. 


Table 9.—Results of tests with liquids on peach trees 10 years old for control 
of peach borer in southern Illinois, fall of 1934 and spring and fall of 1935. 


NUMBER OF TREES TOTAL BorRERS PER CENT OF 
TREATMENT SEASONS TREATED  BORERS DeEap Borers DEAD 


Liquids Containing PDB 


PDB in Dendrol 3 30 83 80 96.4 
PDB in Scalecide 2 20 Sil 49 96.1 
PDB* in Scalecide 1 10 29 7 58.6 
PDB in soybean oil 2 20 34 31 91.2 

Liquids Without PDB 
Dichloropentane, 

1 ounce per tree Y 20 5S 37 69.8 
Dichloropentane, 

2 ounces per tree 3 30 87 83 95. 
Scalecide, 1 in 8 2 20 45 10 DBP) 7s 
Check, no treatment 3 35) 103 3 2.9 
Crystal PDB, standard 

(1 ounce per tree) 3 50 129 WAS 95.4 


: *Bach tree received a dilution containing one-fifth ounce PDB as compared with 1 ounce in each 
of the three other treatments containing PDB. 


Our procedure when testing the PDB crystals in oils was 
to dissolve 2 pounds of the crystals in 1 gallon of oil and dilute 
with water to make 4 gallons. We then applied 1 pint of the 
diluted mixture to a mature tree. This gave each tree 1 ounce of 
PDB, the same dose it would receive in the crystal treatment. 
Mounding was done as in the crystal method. 

In the case of Scalecide without PDB, the directions of the 
manufacturer were followed. The dirt was removed from the 
base of the tree as for worming, gum rubbed off with a sack and 
1 quart of a dilute mixture, 1 part of Scalecide to 8 parts water, 


18 ILLINOIS NATURAL HIsTORY SURVEY CIRCULAR 31 February 


sprayed at the,base of the tree. At the suggestion of the manu- 
facturer, a similar dilution of Scalecide containing one-fifth 
ounce PDB was tested in the same way. 


Fig. 12.—Liquids for peach borer control may be poured around the tree, but this 
method of application increases the problem of carrying liquid in the orchard. 


Other liquids were either sprayed with a bucket sprayer or 
poured, figs. 12, 13, 14. In the case of dichloropentane, 1 or 2 
ounce dosages were applied per mature tree. 

All of the tests recorded in table 9 were on trees 10 years of 
age or older. No injury was observed following any of the treat- 
ments, but Mr. Snapp, working with some of the materials in 
Georgia at the same time on trees of various ages, reported very 
severe injury with dichloropentane used at strengths which gave 
good control. 

The tests recorded in table 9 show that control as good as 
that given by the standard PDB crystal treatment was ob- 
tained by the application of. 1 pint Dendrol or Scalecide dilution 
per tree when the dissolved PDB was used in amounts equal to 
the standard. They show approximately the same control with 
2 ounces of dichloropentane, and poor results with Scalecide 
diluted 1 in 8. 

In the late summer and early fall of 1937, the Illinois 
Natural History Survey and the United States Bureau of En- 


1939 CHANDLER: PEACH TREE BORERS 19 


tomology and Plant Quarantine conducted cooperative experi- 
ments in southern I]linois in which vegetable and mineral oils 
were tested as carriers of PDB. Also tested were ethylene 
dichloride and dichlorethyl ether, two liquids not containing 
PDB. In table 10 are given results of these cooperative tests, 
made under the direction of Mr. Snapp and the writer. 

Tests recorded in table 10 show that control as good as 
that with the standard PDB crystals was obtained with all ma- 
terials tested except dichlorethyl ether, and that almost as good 
control was obtained by a reduction of about 25 per cent in the 
amount of PDB incorporated into the carriers. Cottonseed oil 
gave a little better control than did the mineral oils when used 
as a carrier of PDB. Injury by PDB was discussed on page 5. 

Of the materials tested in this experiment, ethylene dichlo- 
ride looks especially promising. 

In tests conducted on nursery stock, so few borers were 
found that control tests were not conclusive and they are not 


Fig. 13.—Soil often needs some cup- Fig. 14.—A few shallow cuts may be 
ping or working to prevent run-off of made with a spade in the soil around 
liquid applied for peach borer control. peach tree to prevent run-off of liquid. 


included in table 10. However, the finding of more borers in 
the check plots than in the treated indicates that control was 
obtained by the use of liquids containing very small quantities 
of PDB. In one of the treated plots, each nursery tree received 
a one-eighth ounce dosage of PDB in liquid; in the other plot, 
each tree received one-sixteenth ounce. 

Spraying and Pouring Liquids.—In the tests recorded in 
table 10, most of the plots treated with liquids were divided into 


20 ILLINOIS NATURAL HISTORY SURVEY CIRCULAR 31 February 


Table 10.—Results of tests with liquids for control of peach borer, made in late 
summer and early fall of 1937, the Illinois Natural History Survey and the United 
States Bureau of Entomology and Plant Quarantine cooperating. 


PER CENT 
PLoT TREES TotTaL BorRERS OF BORERS 
No. TREATMENT TREATED BoRERS DEAD DEAD 
Materials Containing PDB 
1 Crystal PDB, standard 
13-year trees, 1 oz. per tree 10 32 32 100.0 
5-year trees, 34 oz. per tree 10 21 20 95.2 
2-year trees, 14 oz. per tree 10 0 0 —S 
2 PDB in cottonseed oil emulsion* 
13-year trees, 1 oz. in 14'pt. per tree 10 33 33 100.0 
13-year trees, 34 oz. in 14 pt. per tree 10 26 24 92.3 
5-year trees, 34 oz. in 14 pt. per tree 10 15 15 100.0 
5-year trees, 14 oz. in 14 pt. per tree 10 4 4 100.0 
2-year trees, 14 oz. in 4 pt. per tree 10 0 0 ——- 
2-year trees, 14 oz. in \% pt. per tree 10 0 0 -_— 
3. PDB in mineral oil emulsion? 
13-year trees, 1 oz. in !4 pt. per tree 10 32 29 90.6 
13-year trees, 34 oz. in 14 pt. per tree 10 41 39 95.1 
5-year trees, 34 oz. in 14 pt. per tree 10 8 8 100.0 
5-year trees, 4 oz. in 14 pt. per tree 10 39 38 97.4 
2-year trees, 14 oz. in \ pt. per tree 10 2 2 100.0 
2-year trees, 1g oz. in 4 pt. per tree 10 2. 1 50.0 
4 PDB in equal parts cottonseed 
and mineral oilf emulsions 
13-year trees, 1 oz. in 44 pt. per tree 10 56 52 92.9 
13-year trees, 34 oz. in 14 pt. per tree 10 61 60 98.4 
5-year trees, 34 oz. in 14 pt. per tree 10 17 17 100.0 
5-year trees, 14 oz. in 1% pt. per tree 10 6 6 100.0 
2-year trees, 4 oz. in 14 pt. per tree 10 4 4 100.0 
2-year trees, 14 oz. in 4 pt. per tree 10 5 5 100.0 
7 PDB in Dendrol 
13-year trees, 1 oz. in 44 pt. per tree 10 37 36 97.3 
5-year trees, 34 oz. in 14 pt. per tree 10 6 100.0 
2-year trees, 14 oz. in 4 pt. per tree 10 3 3 100.0 
Materials Not Containing PDB 
5 Ethylene dichloride emulsion 
13-year trees, 14 pt. 25% emulsion 
per tree 10 76 76 100.0 
13-year trees, }5 pt. 20% emulsion 
per tree 10 30 29 96.7 
5-year trees, 15 pt. 20% emulsion 
per tree 10 13 13 100.0 


5-year trees, 15 pt. 15° emulsion 
per tree 10 0 0 == 


1939 CHANDLER: PEACH TREE BoRERS 21 


Table 10—Concluded. 


PER CENT 
ROR TREES ToTaAL BoreERS OF BOoRERS 
No. TREATMENT TREATED BorERS DEAD DEAD 
2-year trees, 14 pt. 20% emulsion 
per tree 10 5 5 100.0 
2-year trees, 14 pt. 159% emulsion 
per tree 10 3 3 100.0 
12 Dichlorethyl ether dissolved in water at 
rate of 30 cc. in 1 gal. 
13-year trees, 1 pt. per tree 5 16 3 18.8 
13-year trees, 34 pt. per tree 5 15 9 60.0 
5-year trees, 14 pt. per tree 5 0 0 
6 Check, no treatment 
13-year trees 5 43 2 4.7 
5-year trees 5 35 3 8.6 
2-year trees 10 5 0 0.0 
*All emulsions, 50 per cent stock. Viscosity, 491.4. 


two parts; in one part the liquids were sprayed at the base of 
the tree with a bucket sprayer and in the other the liquids were 
poured, fig. 12. A summary of all the results gives the following 
percentages of dead borers: 96.1 per cent in plots in which the 
liquid was sprayed; 97.7 per cent in plots in which the liquid 
was poured. Under certain circumstances it might be of im- 
portance to know that liquids give as good results when poured 
as when sprayed. 


Comparison of Solids and Liquids 


The use of PDB crystals for peach borer control, now a 
long established practice, is relatively cheap, requires very 
little skill or equipment and is effective. Although under 
certain conditions some injury occurs, growers are not likely to 
make a change unless they can see sufficient advantages. For 
this reason we have tried to explore all the possible advantages of 
the liquid method and to compare them with those of the crystal 
method. 

Injury.—Injury by PDB has already been discussed on 
page 5. No material has yet been found that we are certain will 
give no injury under any circumstances, but some effective 
liquids, such as ethylene dichloride and PDB in cottonseed oil 
emulsion, are apparently a little safer than PDB crystals. 

Effectiveness at Low Temperatures.—Because of their 


Dy ILLINOIS NATURAL HisToRY SURVEY CIRCULAR 31 February 


physical characteristics, liquids are likely to become effective 
more quickly than solids. For this reason they might be effective 
at lower temperatures or might utilize to better advantage very 
short periods of high temperature. Tests with ethylene dichlo- 
ride made in New York state by Mr. Snapp gave good results at 
low temperatures. 

A test of ethylene dichloride emulsion was begun Novem- 
ber 18, 1937, in southern Illinois at soil temperatures which 
remained almost stationary at 35 degrees F. during the first 
four days, varied from 30 to 40 degrees during the next four 
days and attained a maximum of 52 degrees on the two days 
before the examination, 11 days after treatment. On December 
3, a treatment was made with this material and with crystal 
PDB for comparison. Temperatures following this treatment 
ranged only from 35 to 40 degrees. Examinations were made on 
December 16. Table 11 gives the results of these tests. 

Table 11.—Results of tests for effectiveness of ethylene dichloride used against 


peach borers in southern Illinois at low temepratures ranging from 30 to 52 degrees 
F., autumn, 1937. 


MATERIAL, DATE OF TREATMENT TREES TOTAL Borers PER CENT OF 
AND TEMPERATURE RANGE TREATED BORERS DeEap Borers DEAD 


Ethylene dichloride, 
November 18, 30-52 degrees 10 41 34 82.9 


Ethylene dichloride, 


December 3, 35-40 degrees 5 20 18 90.0 
PDB crystals, 
December 3, 35-40 degrees 5 10 1 10.0 


Apparently ethylene dichloride has the advantage of being 
effective at lower temperatures than is crystal PDB. . 

Further tests under conditions of cold soil were conducted 
in March, 1938, in Jackson and Johnson counties by the Illinois 
Natural History Survey, represented by the author, and the 
United States Bureau of Entomology and Plant Quarantine, 
represented by Mr. Snapp. Three weeks intervened between 
treatment and worming. In this time, soil temperatures 
averaged each week from 50.2 to 55.3 degrees with minimums of 
43 and a maximum one day only of 64. Ethylene dichloride 
emulsion and dichlorethyl ether, each at different strengths, 
were compared with crystaline PDB on trees of three different 
ages. Table 12 summarizes the results. 


1939 CHANDLER: PEACH TREE BORERS 23 


Table 12.—Results of tests for control of peach borer by ethylene dichloride 
emulsion and dichlorethyl ether under conditions of low soil temperature at Car- 
bondale, March, 1938. 


AGE OF AMOUNT TREES ToTaAL PER CENT OF 
TREES MATERIAL USED PER TREE TREATED  BorERS BoreERS DEAD 
10 years Ethylene dichloride VY pt. of 25% 20 134 97.0 

10 years Ethylene dichloride V6 pt. of 20% 20 186 9275 

10 years Dichlorethyl ether i joWrS* 10 60 96.7 

10 years Dichlorethyl ether 34 pt.* 10 46 OS 

10 years PDB crystals il Le 10 35 34.3 

10 years Check, no treatment == 5 28 0.0 

4 years Ethylene dichloride V4 pt. of 20% 10 85 97.6 

4 years Ethylene dichloride 14 pt. of 15% 10 53 96.2 

4 years Dichlorethyl ether ¥%, pt.* 10 96 92.7 

4 years Dichlorethyl ether Vy pt.* 10 112 a 

4 years PDB crystals 34 OZ. 10 116 87.1 

4 years Check, no treatment 5 80 0.0 

2 years Ethylene dichloride \y pt. of 15% 20 52 92.3 

2 years Ethylene dichloride 4 pt. of 124% 20 51 94.1 

2 years Dichlorethyl ether Vou per 10 35 82.9 

2 years Dichlorethyl ether Yn Dio 10 WD, 81.8 

2 years PDB crystals VY oz. 10 54 16.7 

2 years Check, no treatment 5 14 0.0 


*Diluted, 30 cc. dichlorethyl ether to 1 gallon water. 


This experiment indicates that ethylene dichloride emul- 
sion applied at strengths shown in table 12 will give very 
effective control of the peach borer at temperatures which are 
not high enough for good control by PDB crystals. It also 
indicates that dichlorethyl ether may give fairly good control 
under these conditions. The experiments made in the late 
summer and early fall, when soil temperatures were higher, 
were not very favorable to dichlorethyl ether, table 10. 

Necessity for Mounding.—Because of their physical state, 
it is conceivable that liquids applied for peach borer control 
might not need mounding. This conception was tested during 
the spring and fall of 1936. PDB was dissolved in Dendrol at 
the usual rate of 2 pounds of crystals to 1 gallon of the emulsion 
and diluted with water so that when each tree was treated with 
1 pint of material it received 1 ounce of PDB. Four tests were 
conducted in two orchards over two seasons. Trees were 10 and 
11 years old. Table 13 gives the results. 

Table 13 shows only one test in four in which control in 
unmounded trees equaled or bettered that in the mounded, 
and the averages for the 210 trees treated and nearly 500 borers 


24 ItLinors NATURAL HIstoRY SURVEY CIRCULAR 31 February 


concerned showed a kill that was 21.6 per cent less in the un- 
mounded than in the mounded trees. Obviously, liquid treat- 
ment does not lessen the necessity for mounding. 


Table 13.—Results of tests to determine necessity of mounding after liquid 
treatments. 


TREES MOUNDED TREES Not MoOuNDED 
TEST Trees Total Per Cent of Trees Total Per Cent of 
NuMBER Treated Borers Borers Dead Treated Borers Borers Dead 
1 0 22 100.0 10 22 86.4 
2 25 Sy 90.4 25 130 68.5 
3 50 107 oy a) 50 96 70.8 
4 20 40 Aes) 20 25 88.0 
All tests 105 291 04 1 105 273 (Zee 


It has been stated that mounds for trees given the liquid 
treatment need not be made so large as those for trees given the 
crystal treatment, fig. 5, and need not be tamped down. We 
have made no tests to determine the validity of this statement, 
but in view of the above experiment it seems rather doubtful. 
In the 1937 cooperative experiment, results of which are shown 
in table 10, liquid-treated trees were given a minimum amount 
of mounding and no tamping, but no comparison was made 
with crystal-treated trees with respect to size of mounds. 

Equipment Required.—The liquid treatment requires the 
handling of 8 to 16 times as much weight as does the crystal 
treatment. 

The most economical method of applying the liquid is 
usually with the power sprayer, which the orchardist already 
possesses. The life of a spray machine and the cost of mainte- 
nance is, in general, inversely proportional to the amount of use 
given it. Estimates from growers of large acreages in Illinois 
indicate that the cost is approximately $2 in depreciation, re- 
pairs and upkeep, not including gasoline and oil, to operate a 
sprayer for one day. In an actual test, described below, one- 
half gallon of gasoline at 18 cents per gallon was required to 
operate a sprayer one hour. To the cost of the sprayer must be 
added the expense involved in having it pulled by a team or 
tractor. 

A regulator has been devised, fig. 15, which may be at- 
tached at the end of the spray hose for liquid treatments. This 
is necessary so that the right amount of liquid, no more and 


bho 
on 


1939 CHANDLER: PEACH TREE BORERS 


no less, may be applied to each tree. Selling at the present time 
for $20, this piece of equipment could be made by a mechanically 
trained grower for less if his time were not considered, and 


Fig. 15.—Regulating device in operation. With this device a measured half pint or 
pint of liquid can be sprayed around the base of trees for peach borer control. 


probably if any great demand should develop it could be pro- 
duced commercially more cheaply than at present. Every man 
applying the liquid must be equipped with the device, and the 
number of men working at this operation is limited by the 
supply of regulators. 

The fact has been brought out on page 21 that, in the 
application of the liquid, pouring is as effective as spraying. 
If liquid were poured around each tree, the use of the sprayer 
and regulating device would, of course, be eliminated. How- 
ever, the problem of carrying the necessary large amount of 
liquid from tree to tree remains. The amount of time required 
by this method and present labor prices must be considered in 
figuring its cost. 

Time Required.—In the spring and fall of 1937, two tests 


26 ILLINOIS NATURAL History SURVEY CIRCULAR 31 February 


were run under orchard conditions to determine the actual time 
required to treat by the liquid and the crystal methods. In the 
Hartline Orchards, Union County, the tests covered an area of 
714% acres. In the Heaton Orchards, Johnson County, they 
covered about 18 acres. 

In all orchards, the liquid treatment was given by one 
man with a power sprayer and a regulating device; the pressure 
in the sprayer was reduced to 100 pounds. No driver except the 
operator was used, a saving that is possible only with steady, 
dependable teams. Two men followed the sprayer, mounding. 
Time-out was taken for every interruption, so that the records 
include only the actual time of work. In the crystal-treated 
blocks of the Heaton Orchards, records were kept of a crew of 
nine men, three to apply the crystals and six to mound. 

The average time required to treat and mound a tree was 
1.4 minutes by the liquid method and 2.0 minutes by the crystal 
method, or a saving of 30 per cent in time by the liquid method. 

On the basis of a common wage in Illinois orchards, 15 
cents per hour or one-fourth cent per minute, the following cal- 
culations were made: 

Liquid treatment, 1 tree (1.4 man-minutes) cost 
$0.0035 (about one-third cent). 

Crystal treatment, 1 tree (2.0 man-minutes) cost 
$0.0050 (one-half cent). 


It therefore cost $0.0015, approximately one-seventh cent, 
more per tree for labor to treat by the crystal than by the liquid 
method. 

In these tests, 3 gallons of gasoline were required to keep 
the engine running for 6 hours. This was equivalent to 5 
gallons for a 10-hour day. Oil and gas together for a day cost 
at least $1. The average number of trees treated by a sprayer in 
one day was 427. The cost of the gas and oil, therefore, amounted 
to $0.0023 (about one-fourth cent) per tree, approximately 50 
per cent more than the amount saved by the greater speed of 
the liquid method. The item of $2 per 10-hour day for deprecia- 
tion and repair of spray rig added another $0.0047 (about one- 
half cent) per tree to the cost of the liquid treatment. 

The cost of such items as filling the spray rig, heating and 
preparing the material, hitching and unhitching the team, and 
repairing possible spray-rig breakage was not considered in 
the calculations. 

Cost of Materials.—Price of the material is another im- 


1939 CHANDLER: PEACH TREE BORERS 27 


portant item in a comparison of liquid and crystal treatments. 
At a retail price of 18 cents per pound in 100-pound lots, the 
price which now prevails, and at the amount of 1 ounce PDB 
per tree, the cost of the material for the crystal treatment is 
slightly more than 1 cent per tree, or about twice the cost of 
the labor required to apply it. Obviously, the cost of oils used 
as carriers must be considered. Prices of these materials vary 
from year to year, but, if used at the dilutions indicated, even 
the cheapest of the carriers will add from three-fourths to 1 
cent per tree, which almost doubles the cost of materials. 

The greatest hope for economy is from the materials which 
do not contain PDB. Ethylene dichloride at the present price 
of 614 cents per pound in 50-gallon drums costs approximately 
1 cent per tree. Dichloropentane can now be obtained for 5 cents 
per pound in single barrel lots. If this is applied at the rate of 
2 ounces per tree, the cost is about $0.007 (slightly less than 
three-fourths cent) per tree. However, insufficient work has 
been done with this material, and because of severe injury in 
Georgia we do not at the present time recommend it. 

Summary of the Solid and Liquid Methods.—-Following is a 
summary of the facts that have become apparent through com- 
parison of the solid and liquid methods of treating for peach 
borer control: 

1.—To date only one solid, crystal PDB, is known to be 
both safe enough and cheap enough to use. 

2.—Four liquid substitutes tested, cottonseed oil emulsion, 
mineral oil emulsion and miscible oils, all impregnated with 
PDB, and ethylene dichloride emulsion (without PDB) are as 
effective under normal treating temperatures as PDB crystals. 
These substitutes appear to be a little less likely to injure the 
tree than are the crystals. 

3.—When the soil is cold, ethylene dichloride emulsion is 
more effective than PDB crystals. 

4.—Mounding is necessary after liquid treatments, though 
possibly the mounds need not be so large as when crystals are 
applied. 

5.—Cupping may be necessary under certain soil condi- 
tions to prevent run-off when liquids are used. 

6.—Liquid substitutes found safe and efficient are all more 
expensive than PDB crystals with the exception of ethylene 
dichloride emulsion, which costs about the same. 

7.—The liquid method of treating trees is 30 per cent 


28 ILLINOIS NATURAL History SURVEY CIRCULAR 31 February 


faster than the crystal method if the liquid is sprayed on the 
trees with a gasoline driven machine. 

8.—The use of a spray machine for applying the liquid 
entails the purchase of a special regulating device which limits 
the number of operators. 

9.—Under Illinois conditions the financial gain from the 30 
per cent saving in time, possible with the liquid treatment, is 
offset by any one of several items of expense connected with the 
use of a sprayer and team. 

10.—If the liquid material is poured around the trees, no 
time is saved, and the carrying into the orchard of weight eight 
times that of crystal PDB necessitates the use of a team and 
probably is more expensive and less simple an operation than 
the crystal treatment. 


THE LESSER PEACH BORER 


As the lesser peach borer, Synanthedon pictipes (G. & R.), 
works in crotches and wounds of peach trees, fig. 16, rather than 
at the base, it does not girdle the trees. It is termed “‘lesser”’ 


{ 


Fig. 16.—Injury caused by lesser peach borer, Synanthedon pictipes (G. & R.). 


1939 CHANDLER: PEACH TREE BORERS 29 


Fig. 17.—Wounded areas and weakened crotches on peach trees are especially sub- 
ject to attack of lesser peach borer. 


because the damage it does is not so great as that done by the 
peach borer, Conopia exitiosa (Say). 


Extent of Injury 


In our observations we have found the lesser peach borer 
working usually in places on the tree previously weakened or 
injured. Weakened crotches, areas on the trunk skinned by a 
tractor, sun-scalded and winter-injured branches, and branches 
broken by heavy crops are favored places of attack, fig. 17. 
There is evidence to show that some of the splitting of crotches 
which occurs would be avoided if it were not for the additional 
weakening caused by the borers working at a critical point. 
Needless to say, any horticultural practice that will avoid 
injuries to the trees will help to control this insect. 

In order to learn something of the prevalence of the lesser 
peach borer in actual percentages, examinations were made in 
10 representative commercial orchards of bearing age in south- 
ern Illinois. Fifty trees, scattered over each orchard, were ex- 
amined and a record made of the infestation. A tree with one 
or more injured areas was classed as infested. 

Degree of infestation among the orchards ranged from 10 


30 ILLINoIs NATURAL History SURVEY CIRCULAR 31 February 


to 32 per cent. Of all trees examined in the 10 orchards, 22.6 
per cent were infested with the lesser peach borer. 


Control Measures 


Since the lesser peach borer usually works too high on the 
tree to be covered with a mound, most growers have relied on 
the old jack-knife and wire method formerly used on the peach 
borer, fig. 9. With the introduction of the liquid method of 
peach borer control, it was suggested by Mr. Snapp that some 
of the materials used would control the lesser peach borer. 
In our control experiments for this latter insect, we used ma- 
terials at strengths sufficient for peach borer control and also 
at even greater strengths. 


Experimental Work 


In our observations of liquid applications for the peach 
borer in actual orchard practice, it was evident that the operator 
has neither liquid enough in the half pint or pint applied per 
mature tree, or time enough before his automatic cut-off stops 
him, to treat anything but the base of the tree, and he is not 
likely in this one operation to do any treating for the lesser 
peach borer. 

For this reason we confined our tests to painting the affected 
areas, fig. 18. When the gum could easily be removed, this 
was done before the material was applied, but when, as often 


Table 14.—Results of experiments for control of lesser peach borer by insecti- 
cides painted on affected areas. The experiments cover spraying seasons in the 
years 1934-37. 


STOCK, DILUTION, PER CENT 
POUNDS OUNCES OF 
MATERIALS PDB PER PDB PER SEASONS TREES TOTAL  BORERS 
UsED GALLON PINT TESTED TREATED BORERS DEAD 
PDB in Dendrol 2 1 3 38 47 Sie 
PDB in Dendrol 2 » 5 70 83 86.7 
PDB in Dendrol 4 4 D 35 27 100.0 
PDB in soybean 
oil 2 mal 1 10 23 8.7 
PDB in Niagara 
Tar Oil 2 2 1 10 16 93.8 
PDB in Niagara 
Tar Oil 2 4 (Un- 1 10 10 100.0 
diluted) 
Dichloropentane — Undiluted 1 10 24 70.8 
5 45 47 .0 


Checks — — 


1939 CHANDLER: PEACH TREE BORERS 31 


was the case, the gum had hardened it was left undisturbed. 
Good penetration of the gas, however, resulted even in the 
latter situation, and excellent control was obtained. The ma- 
terials tested were those used for peach borer control with the 
addition of one other. They are indicated in table 14, which 
gives the results of the experiments. 

No injury could be observed in the areas receiving any of 
the materials. The infested areas are usually somewhat corky 
and protected by outer bark and by several layers of inner bark. 
If the paint is applied chiefly to the place showing infestation, 
there seems to be little danger of injury, even at the strengths 
used. In one of the tests, the lower half of tender shoots of the 
current season, growing up in the center of the tree, were 
painted; the upper parts were left for checks. With the dichloro- 


“7? : ee t 
29r-eoe 


»< *.% 
af, + 


Fig. 18.—Control of lesser peach borer by painting affected areas of the tree with 
PDB-oil solution. Even a boy with a bucket of material can treat a large number of 
trees in a few hours. 


32 ILLINOIS NATURAL HisTORY SURVEY CIRCULAR 31 February 


pentane and the PDB in Dendrol diluted 2 ounces to 1 pint, 
injury occurred on from 10 to 25 per cent of the shoots, al- 
though no injury whatever could be found on the affected 
areas of the older wood. 

Examination of table 14 shows that rather disappointing 
results were obtained with the strength of PDB in Dendrol 
which gave very satisfactory control for the peach borer when 
mounding was done. This strength was 2 pounds PDB to 1 
gallon Dendrol, diluted with water so that each pint of the mix- 
ture contained 1 ounce of PDB. However, when that strength 
was doubled to 2 ounces PDB per pint of mixture, 86.7 per cent 
kall was obtained as a five-year average. 

An attempt was made to increase this strength by dis- 
solving 4 pounds PDB in Dendrol and diluting so that each 
pint of mixture contained 4 ounces PDB. Although this mix- 
ture gave better control than the one containing 2 ounces PDB 
to the pint, it was not satisfactory because of the trouble in- 
volved in incorporating such a large amount of the crystal into 
the oil, the difficulty of keeping it from crystalizing out on 
cooling and the danger of injury. The same strength could have 
been obtained by using 2 pounds of PDB to 1 gallon of Dendrol 
and applying without dilution, but it was found that spreading 
qualities of the paint are much improved when it 1s diluted with 
water. 

It is probably best in using Dendrol to keep close to the 
proportion of 2 pounds PDB to 1 gallon of the oil and to the 
dilution of 2 ounces PDB to 1 pint of mixture. 

It may be-seen in table 14 that the tar oil used gave in 
the two tests somewhat better control than the miscible oil. 
Although no injury occurred in our tests, it should be borne in 
mind that there is a greater probability of injury from a tar 
oil than from a miscible oil. After five seasons of tests, we do 
not hesitate to suggest the use of PDB in a miscible oil at the 
rate of 2 pounds PDB to 1 gallon of oil diluted with water so 
that there are 2 ounces PDB to 1 pint of mixture. 

In making up this material for use in the orchard, if all the 
stock is to be diluted, 2 pounds of the crystal dissolved in 1 
gallon of the miscible oil should be diluted with water to make 2 
gallons of mixture. If only a portion of the stock is used, it 
should be diluted at the rate of 6 parts by volume of the oil- 
PDB mixture to 4 parts of water. This will give slightly more 
than 2 ounces PDB to 1 pint of the diluted mixture. 


1939 CHANDLER: PEACH TREE BORERS 33 


This method of treating lesser peach borers is relatively 
cheap. A man or even a boy with a bucket of material can 
cover a large area in a day, examining trees and painting only 
the spots showing injury. In a trial test over a limited area, 1 
pint treated all the infested areas on eight trees. With the 
material used, PDB in Dendrol, the cost amounted to one-half 
Genii per tree: 


Time of Treatment 


Lack of information on the life history of the lesser peach 
borer in Illinois makes it impossible to state definitely the best 
times to treat for it. Our observations indicate that emergence 
starts earlier in the season than is the case with the peach borer, 
and it seems probable that treatments should be made in the 
fall rather than in the spring. The occurrence of a second brood 
in Illinois is indicated by studies already made in Ohio and 
West Virginia. With our present knowledge of the insect, we 
suggest treatment about the same time in the fall as for the 
peach borer. 


THE PEACH BARK BEETLE AND THE SHOT-HOLE 
BORER 


The peach bark beetle, Phthorophloeus limnaris (Harr.), 
and the shot-hole borer, Scolytus rugulosus Ratz., attack the 
trunk and branches of peach trees in Illinois, working especially 
on weakened trees and branches, figs. 19, 20, 21, 22. The best 
method of control is to correct whatever has caused the weak- 
ness, such as lack of nitrogen, poor drainage, attacks of scale 
or peach borers, lack of care and cultivation, or winter injury. 
This is not always possible, however, and at times these insects 
attack apparently healthy trees. There is need for direct con- 
trol by insecticides. 

Our control experiments with these insects began in 1931 
with a test of PDB in cottonseed oil sent by Mr. Snapp from 
the federal laboratory at Fort Valley, Ga. A kill of 96.5 per 
cent resulted from the first test. 

The following year the same material, possibly deteriorated 
by this time, was used in another test, and, though as good 
control was secured, very severe injury and killing of branches 
resulted, which discouraged us from further trials until the fall 
of 1935 when tests were started with PDB and miscible oil 
mixtures. 


34 ILLINOIS NATURAL HISTORY SURVEY CIRCULAR 31 February 


Fig. 19.—Weakened peach branch showing injury by bark beetles, Scolytus rugulosus 
Ratz. Note tiny holes made by beetles, some of holes filled with exuding gum. 


Fig. 20.—Branch of peach infested with bark beetles. The bark has been pulled off 
to show the white grubs in position. 


1939 CHANDLER: PEACH TREE BORERS 35 


In tests made in 1935, 1936 and 1938 for control of the 
shot-hole borer, PDB was dissolved in Dendrol at the rate of 2 
pounds of PDB to 1 gallon of Dendrol. This solution was diluted 


Fig. 21.—An infested peach branch that Fig. 22.—Results of bird attack on peach 
shows burrows of the bark beetle. branch infested with bark beetle. 


with water. Part was diluted so that each pint of mixture con- 
tained 1 ounce of PDB and part so that each pint contained 2 
ounces of PDB. Table 15 shows the results obtained. 

These tests indicate that with the PDB-Dendrol mixture 
it is possible to attain good control of the beetles but not of the 
grubs of the shot-hole borer. The reason for this apparently lies 
in the fact that the beetles stay close to the holes they have made 
in the bark, whereas, the grubs, after hatching, tunnel too far 

Table 15.—Results of 1935, 1936 and 1938 tests for control of shot-hole borer* 


by PDB in Dendrol, a miscible oil. Both beetles and grubs of the shot-hole borer 
were present in the 1936 and 1938 tests. 


DILUTION, 
OUNCES 

YEAR PDB PER TOTAL PER CENT OF TOTAL PER CENT OF 
TESTED PINT BEETLES BEETLES DEAD GRUBS GruBs DEAD 
1935 1 ounce 9 DD. —- — 

2 ounces 8 1S - 

Check 18 0.0 — 
1936 2 ounces 26 100.0 10 0 

Check 15 6.7 20 0 
1938 2 ounces 89 98.9 362 if faa 

Check 66 0.0 337 0.0 


*Determined by Dr. H. H. Ross as Scolytus rugulosus Ratz. 


36 ILLinois NATURAL History SURVEY CIRCULAR 31 


from the openings for the gas to penetrate. It appears necessary 
to treat when the beetles have first attacked the trees and before 
they have laid eggs. 


_ RECENT PUBLICATIONS 


of the Illinois State Natural History Survey 


A.—ILLINOIS NATURAL HISTORY SURVEY BULLETIN. 


Volume 21, Article 1.—The Effect of Petroleum-oil Sprays on Insects and Plants. By 
M. D. Farrar. November 1936. 32 pp., frontis. + 21 figs., bibliog. Contents: Fore- 
word; Properties of oil emulsions; Effect of petroleum oils on plants; Insecticide 
tests with the emulsions; Oils with fungicides. 


Volume 21, Article 2.—Responses of the Large-mouth Black Bass to Colors. By Frank 
A. Brown, Jr. May 1937. 23 pp., frontis. + 10 figs., bibliog. Contents: Problem 
of color vision in fishes; Materials for the experiments; Training and responses of 
large-mouth black bass; Interpretation of the responses; Summary. 50 cents. 


Volume 21, Article 3.—Studies of Nearctic Aquatic Insects. By H. H. Ross and T. H. 
Frison. September 1937. 52 pp., frontis. + 86 figs., bibliog. Contents: I. Nearetie 
alder flies of the genus Sialis (Megaloptera, Sialidae) by H. H. Ross; and IL. 
Descriptions of Plecoptera, with special reference to the Illinois species, by T. H. 
Frison. 50 cents. 


Volume 21, Article 4.—Descriptions of Nearctic Caddis Flies (Trichoptera) with 
special reference to the Illinois species. By Herbert H. Ross. March 1938. 83 pp., 
frontis. + 123 figs., foreword, index. $1.00. 


B.—ILLINOIS NATURAL HISTORY SURVEY CIRCULAR. 


28.—Rout the Weeds! By L. R. Tehon. August 1937. 34 pp., color frontis. + 8 figs. 
Contents: The importance of weeds; Weeds as economic factors; Weeds as harbor- 
ers of plant diseases; Relation of weeds to public health; Control methods; Eight 
pernicious weeds of Illinois—common ragweed, giant ragweed, poison ivy, poison 
sumac, wild parsnip, white snakeroot, pokeweed, common burdock. 


29.— Windbreaks for Illinois Farmsteads. By J. E. Davis. April 1938. 18 pp., frontis. + 
12 figs. Contents: Planning the windbreak; Planting the windbreak; Care of the 
windbreak; What the windbreak trees are like. 


30.—Outwitting Termites in [linois. By W. E. McCauley and W. P. Flint. June 1938. 
20 pp., frontis. + 19 figs. Contents: Termites and their habits; Structural control 
of termites; Chemical control of termites; Unified action against termites. 


C.—ILLINOIS NATURAL HISTORY SURVEY MANUAL. 


1.—Fieldbook of Illinois Wild Flowers. By the staff. March 1936. 406 pp., color 
frontis. -+ 349 figs., index. Contents: Introduction; Key to families; Description 
of species (650). $1.50. 


Address orders and correspondence to the Chief 
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Payment must accompany requests for publications, in the form 


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