PROCEEDINGS
OF THE
ACADEMY OF NATURAL SCIENCES
OF
PHILADELPHIA
1 8 8 O.
I'UBLicATioN committee-
Joseph Leidy, M.D., Geo. H, Horn, M.D.,
William S. Vaux, Thomas Meehan,
John H. Redfield.
Editor: EDWARD J. NOLAN, M.D.
PHILADELPHIA:
ACADEMY OF NATURAL SCIENCES,
S.W. Corner Nineteenth and Race Streets.
1881.
^:3<^^
Academy of Natural t^ciENCES ok Philadklphia,
March, 1881.
I hereby certify that printed copies of the Proceedings for 1880 have been
presented at the meetings of the Academy, as follows : —
Pa:
iges 9 to oG
March 23, 1880
57 to 88
March 30, 1880
" 89 to 120
April 6, 18811
" 121 to 152
June 1, 1800
" 153 to 200
July 27, 1880
" 201 to 210
August 10, 1880
" 217 to 232 .
August 17, 1880
" 233 to 248 .
. September 7, 1880
'' 249 to 280
September 28, 1880
" 281 to 328
October 12, 1880
" 329 to 352
November 9, 1880.
" 353 to 384
February 22, 1881.
" 385 to 410
March 1,1881.
EDWARD J. NOLAN,
Becording Secretary.
PHILADELPHIA
W. P. KiLDARE, Printer.
LIST OF CONTRIBUTORS.
With referenrv tn tin sfrci'id iirticlen n>ntrll»{fi(l hi/ enrli.
For Verhiil Cuininuniciitiuns sue (Jfucnil Imlex.
AUeu, Harrisou. Desci iption of a foetal walrus 38
On the Temporal and Masseter Muscles of Mammals 385
Barbeck, Wm. On the Development of Lemna minor. (Plate XVIII.) 230
Beriih, R. On the Xudibranchiate Gasteropod ^lollusca of the Xorth
Pacific Ocean, with special reference to those of Alaska. Part II.
(Plates I.-VIII.) 40
Chapman, H. C. On the Structure of the Orang Outang. (Plates
XI.-XVII. ) r 160
Genth, F. A., Jr. The So-called Emery Ore from Chelsea, Bethel
Township, Delaware County, Pa 311
Hartman, W. D. Description of a Partula supposed to he new, from
the Island of Moorea , '.3"3tt
Heilprin, Angelo. On the Stratigraphical Evidence afforded by the
Tertiary Fossils of the Peninsula of ]Maryland 20
On some new Lower Eocene Mollusca from Clarke Co., Alabama,
with some points as to the Stratigraphical Position of the Beds
containing them ( Plate 20 1 364
Kingsley, J. S. Carcinological Notes, Xo. 1 34
Carcinological Xotes, X^o. 2.— Revision of the Gelasimi. (Plates
IX. and X.) 135
Carcinological Xotes, No. 3. — Revision of the Genus Ocypoda 179
Carcinological Notes, Xo. 4. — Synopsis of the Grapsidse IST
Leidy, Jos. Rhizopods in the Mosses of the Summit of Roan Moun-
tain, Xorth Carolina , 333
Lewis, Henry Carvill. The Optical Characters of some Micas 244
On Siderophyllite, a new 3Iineral 254
The Surface Geology of Philadelphia and Vicinity 258
The Iron Ores and Lignite of the Montgomery Co. Valley 282
On a new Fucoidal Plant from the Trias 293
The Trenton Gravel and its relation to the Antiquity of Man 296
On Philadelphite ( Sp. Nov. i 313
Lockington, W. N. On a Pacific Species of Caulolatilus 13
Description of a new species of Hemitripterus from Alaska 233
Description of a new species of Catostomus (Catostomiis Cyplio)
from the Colorado River 237
]McCook, Rev. H. C. The Shining Slavemaker. — Notes on the Archi-
tecture and Habits of the American Slave-making Ant, Polyergus
lucidus (Plate 19) , 376
Rand, Theo. D. On Randite 274
Report on Plants introduced by means of the International Exhibition,
1876 '. 132
Vodges, Anth. W. Description of a new Crustacean from flic Upper
Silurian of Georgia, with remarks upon Calymene Clintoni. ... 176
PROCEEDINGS
OF THE
ACADEMY OF NATURAL SCIENCES
OP
PHILADELPHIA.
1880.
January 6, 1880.
The President, Dr. Ruschenbergeb, in the chair.
Forty-two persons present.
The following papers were presented for publication:
" On the Nndibranchiate Gasteropod Mnllusca of the North
Pacific Ocean, with special reference to those of Alaska," by Dr.
R. Bergh, of Copenhagen. Part II.
" The Terrestrial Mollusca inhabiting tlie Cooks or Harvej^
Islands," by Andrew Garrett.
The deaths of Dr. S. S. White and Joshua T. Jeanes were
announced.
January 13,
The President, Dr. Ruschenbkrger, in the chair.
Forty-two members present.
On disarticulating branches in Ampelopsis. — Mr. Tuomas
Meehan remarked that some species of Ampelopsis threw off
their dead wood by disarticulation, as he pointed out some years
ago to be the case with Thuja and Taxodium among coniferous
trees. This was especially the case with Ampelopsis quinquefolia,
and A. hipinnata. Tiiese, in common witli many other climbing
plants, produced some portions of their annual growth of so low
2f
10 PROCEEDINGS OF THE ACADEMY OF [1880.
a vital power, that they were soon killed when severe weather
occurred. In the grape vine, for instance, the extreme ends of the
strong branches and whole lengths of weaker ones died during
the winter. These remained on till cut a,wa,j by the pruner, or
until they fell by natural decay. In the AmpeJopsis named they
were thrown oil' by an articulation, so that by spring no dead
wood of the past season's growth would be found on the plants.
Every node included in the dead poi'tion, separated ; so that under
the plants the pieces ma}' be gathered like the separate vertebra;
in a skeleton.
The Ampelopsis, when running up a tree or wall, seldom sent
out lateral branches till it reached the summit. When these side
branches were produced, they appeared, after a few 3^ears,as thick
bushy masses, having the look of a hedge annuallj^ pruned. It
appears that in these cases the annual growth is disarticulated at
just one node above that one made last year— the branch thus
gaining but one node a year. A bushy branch of a dozen years
old, will thus have but a dozen nodes of living wood.
The observations were of some interest just now, from the dis-
covery of a species of Vitis in the South racific, which produced
tubers at the end of the branches, which at the end'of the season
were thrown off by a disarticulation, and in this way aided in
propagation and distribution. Though the disarticulation in the
neighboring genus Amp)elopsis^ as now noted, results only in
ridding the plant at once of useless wood, it showed a relation of
powers in allied species that must be of service to those engaged
in studies of derivation.
Geo. Yaux was elected a member of the Council to serve for
the unexpired term of C. Newlin Pierce. Aubrey H. Smith was
elected to serve for the unexpired term of Edw. D. Cope.
January 20.
The President, Dr. Ruschenberger, in the chair.
Forty persons present.
Notice of the Cruel Thread Worm, Filaria immitis, of the
Dog. — Prof. Leidy directed attention to a specimen, presented by
Mrs. Laura M. Towne, of Beaufort, S. C, consisting of the heart
and part of one lung of a dog, containing thread worms. The
right ventricle of the heart and the pulmonary artery contained a
bunch of the parasites, and several also were contained in the
lung. A similar specimen, with the ventricle literally stuffed full
of worms, is preserved in the museum of the University of Penn-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 11
sylvania. The parasite was described, thirty years ago, in the
Proceedings of this Acadeni}^, under the names of Filar^ia Canifi
cordis and Filaria immitis (see Proc., 18.50,118; 18.5(5,2,5.5),
and since has been repeatedly noticed by observers as infesting
the dog in Europe, India, China, Japan and this country.
The specimen presented is accompanied witli a letter from
Mrs. Towne, giving an account of the occurrence and symptoms
of the parasite as follows :
" I lost several dogs of different breed, age and birthplace, with
the same symptoms — a severe and peculiar cough being the prin-
cipal one. A gentleman living on a neighboring island (the Sea
Islands of South Carolina) lost over thirty hunting dogs in two
or three years with the same symptoms. I watched my two re-
maining dogs closelj'. They were a large Newfoundland (mixed);
and a small terrier. Both had the peculiar cough, which was
excited by any movement, especially after sleeping. It always
ended, after a few coughs, in a violent ettbrt to bring something
up from the throat. This did not seem nausea or sickness, and
as the dogs ncAcr threw up an3'thing, I thought it was due to
hairs in the throat. The two dogs had another symptom.
When they began to run violently, as at hogs, or a strange dog,
they fell down, became stitf and insensible, but in a short time
would get up and resume the chase.
" The little dog died with hemorrhage from the bladder or kid-
neys ; but no post-mortem examination was made.
" The large dog soon began to cough up bloody phlegm, with
considerable fresh blood at times. I found in the phlegm one
morning two Filariie alive, and at least six inches long. I sent
word of. this discovery to the owner of the hunting dogs, and
when his next one died he had it opened, and found the heart and
liver filled with Filaria?.
" My large dog grew so ill that I had him shot. His symptoms
were drowsiness, sleeping with the upper eyelids raised, and the
inner lining showing very red ; holding his head to one side, one
ear drooped ; dragging of one hind leg ; turning round and round
whenever he attempted to go anywhere; and, finally, spasms, ini
which he rolled over and over and drew his head backward. He
was fat and had a good appetite to the last.
" The sister of this dog was given to me. She had a slight cough,
but it increased rapidly. After about three months' attempt to
cure her, I had her shot before her sutfe rings became severe. Her
heart is the one you have. She was fat when she died, and seemed
in good health, except for short breath in running, the cough and
unusual sleepiness.
" In the post-mortem of the first dog, I foimd one Filaria lying
at full length in the windpipe, and in the large artery others
stretched at length and crowded close. Upon cutting into the
12 PROCEEDINGS OF THE ACADEMY OF [1880.
heart, the worms burst forth in bunches, slowly uncoiling them-
selves. They were white, stiff and wire-like, and not in the least
stained with blood. They lived in water about twenty-four hours.
The large blood-vessels of the lungs were filled densel}^, and even
from the small ones long Filarial were with some difficulty with-
drawn. No worms were found in the kidne3S."
January 21.
The President. Dr. Ruschenberger, in the chair.
Nineteen persons present.
A paper entitled '' Careinological Notes, No. 2. Revision of
.the Gelasimi," by J. S. Kingsley, was presented for pul)lication.
The death of Thomas M. Brewer, a correspondent, Avas an-
nounced.
Chas. W. Pickering, John S. Jenks, Wm. H. Jenks, A. K.
Thomas, Ferris W. Price, John Wagner, Chas. P. Tasker, Henry
;F. Fonnad and George W. Biddle were elected members,
Angelo Heilprin, of New York, Dr. C. A. White, of Washing-
:ton, Albert De Selle, of Paris, R. Hoernes, of Vienna, Georges
Rolland, of Paris, and Victor Raulin, of Bordeaux, were elected
< correspondents.
The following were ordered to be printed : —
1880. J NATURAL SCIENCES OF PHILADELPHIA. 13
ON THE PACIFIC SPECIES OF CAULOLATILUS.
BY W. N. LOCKINGTON,
111 the Proc. Acad. Nat. Sci., Phila., 1865, pp. 66-68, Dr. Gill
enumerates four species of his genus Gaulolatilus, one of them,
O. chrysops (Latilus c7i r j/sops, Val.) from the Atlantic, the others
from the Pacific Ocean.
These species are ; G. anomalus (=Dekaya anomalus^ Cooper),
G. princeps (=Latilus princeps, Jenyns), and G. affinis, Gill.
The first and second of these are stated to differ in the proportion
between the length of the posterior dorsal spines and the distance
between the dorsal fin and the lateral line ; as well as in the length
of the pectoral ; while the third species (characterized from a
specimen about three inches long) is described as follows : " Pro-
file quadrant, in front almost vertical; Greatest height less than
four times (.21) in the length (exclusive of the caudal), that of
caudal peduncle about nine times. Head more than f\ of the
length, while its height is to its length as 22^ : 31. Diameter of
eye equal to almost ^ the height of the head. Preorbital very
narrow. Teeth of preoperculum strong and distant ; those of the
middle directed obliquely upwards. Sixth dorsal spine equal to
^ of the length. Anus behind the middle of the length. Caudal
rather exceeding the height of the head. Pectorals equal to ^ of
the length. Yentrals shorter (.18) inserted beneath the base of
the pectoral, its spine at the vertical of the upper axil. D., vii,
25. A., ii, 22. P., 18. Color reddish brown on head and back,
lighter on the sides. A veiy distinct blackish spot above the
axilla of the pectoral. Locality, Cape St. Lucas." Dr. Gill
states his belief that the large eyes and the narrow preorbital are
characters of youth ; and, moreover, hints a doubt as to the spe-
cific identity of G. princeps^ and G. anomalus, but thinks it
scarcely probable on account of the few species known to be
common to Lower California and the Galapagos, the localities
from which the types of G. anomalus and G. princeps were re-
spectively procured.
As I have lately obtained tM'o individuals of a species of Gaulo-
latilus in the markets of San Francisco, I contribute a tolerably
full description, embodying the characters of the two (which
e\'idently belong to the same species) and notes upon the
peculiarities of each. The difference in some of the proportions
14 PROCEEDINGS OF THE ACAPEMY OF [1880.
between these two individuals has almost convinced me of the
identity of G. princepa and G. anomalus ; and I am inclined to
think it probable that the type of G. affinis is only a somewhat
abnormal specimen of the same species. The chief differences
between the smaller of my specimens and the t^'pe of G. affinis
are the more quadrantiform outline and greater length of the head
and the smaller number of dorsal spines and anal rays in the latter-
As, however, the form of the head differs so considerably in
individuals evidently belonging to the same species, too much
stress must not be laid on the former character ; and the A^ariation
in the number of dorsal spines (viii-ix) and dorsal and anal fin-
rays in specimens of undoubted G. anomalus on record, forbid us
to think the latter a positive character.
It is quite possible that an individual may have acquired the
form of head of the adult, while still of small dimensions. The
dorsal spine may be expected (judging from the two specimens
here described) to increase in their proportional length inversely
to the size of the fish.
If my conjecture be correct (and I only give it as a conjecture),
then there is only one Pacific species at present known, ranging at
least from the Galapagos to the Bay of Montere}-, near San Fran-
cisco ; representing in this ocean the G. cluysops of the Atlantic,
and var3'ing somewhat according to age and locality. To thor-
oughly settle the question, a thorough examination of several
specimens from the Galapagos, and a comparison of them with
others from Lower and Upper California, will be necessary.
Presuming, for the occasion, that the}- are identical, the
synonymy will be as follows :
Caulolatilus princeps (Jenyns), Gill.
Latilus pi-inceps, Jenyns, Zool. Beagle, 53, pi. 11.
Latilus prin,cej)s, Gvinther, Cat. Fish. British Museum, II, p. 253.
Dekaya anomala, Cooper, Proc. Acad. Nat. Sci. Phil., 1865, p. 68.
Caulolatilus anomalus, Gill, Proc. Acad. Nat. Sci., Cal., 1865, p. 68.
Caulolatilus affinis, Gill, loc. cit.
Caulolatilus anomalus, Streets, Bull. U. S. Nat. Mus., No. 7, p. 48,
1877.
D. viii-ix, 25-26. A. ii, 24-26. P. 19-20. Y. i. C. ac. 13-14.
C. 13.
General Description. — Profile more or less decurved,the curva-
ture increasing with age ; posterior portion of dorsal outline
nearly straight ; abdominal outline regularly curved. Greatest
1880.J NATURAL SCIENCES OF PHILADELPHIA. 15
depth slightly less than four to rather more than fives times in the
total length ; head, 4j-4f in the same. Greatest thickness about
2^ in the greatest depth. Eye, 4-5 times ; snout, 3-3^ times in
the length of the head. Interorbital width, measured round the
curve of the forehead, 2|-2f in the same. Caudal peduncle, 3-4
times in the greatest depth. Distance from the spinous dorsal to
the lateral line, xg^-l^ times in the height of the last dorsal spine.
Nostrils conspicuous, on the horizon of the centre of the pupil, an-
terior with a valve posteriorly ; posterior larger, simple, subeircular,
distant from the eye about one-third of the diameter of tlie latter.
Ej^es large, lateral, subeircular, their posterior margin nearer
the tip of the operculum than that of the snout.
Mouth slightly ascending forwards ; tip of the intermaxillary
level with the lower margin of the orbit ; posterior extremity of
maxillary nearly vertical with the anterior orbital margin. Max-
illar\^ narrow throughout, its posterior portion free, but the
greater part of its upper edge concealed behind the large preorbital
in the closed mouth. Jaws equal in front in the closed mouth.
Teeth in jaws in several rows in front, diminishing to a single row
farther back on the sides, rather small, slender, acute, recurved at
tip, but those in front of the mandible in the outer row inclined
forwards. Teeth in front largest, those on the sides diminishing,
but the hindermost tooth on each side of each jaw more or less
developed as a canine, though still shorter than tlie anterior teeth.
Xo teeth on vomer or palatines. Upper phaiyngeals set with
sharp, irregularly spaced, eardiform teeth; lower pharyngeals with
an outer and inner row of similar teeth, with some irregularly placed
teeth between the rows. Lower phaiyngeals entirely separate.
(rill-rakers of front of first branchial arch slender, rather stiff,
ab6ut ^ the diameter of the eye, all the others tubercular.
Hinder border of preoperculum vertical, very slightl}- curved,
lower angle rounded, set with teeth which slightly increase in
size at the angle, but do not extend along the lower border.
Operculum ending behind in a broad flat spine.
Dorsal commencing above the upper pectoral axil, very long,
the tips of its terminal rays reaching the caudal accessories ; the
length of its base about half the total length of the fish ; spinous
dorsal 3f-4 times in the total length of the fin, and lower than the
soft portion. First dorsal spine shortest, the others increasing
rapidly to the fifth, more slowly to the eighth or ninth ; the longest
6^-8 times in the total length of the fin.
16 PROCEEDINGS OF THE ACADEMY OF [1880.
Soft dorsal continuous with the spinous portion, and almost
equal in height throughout, the last ray excepted. Last ray
much shorter than the others. Height of soft dorsal, in front
5|— If times in the total length ; man}^ of the rays simple, some
slightly bifurcate at the tip, the two or three last ra^'S twice
branched.
Anal commencing under fifth dorsal ray, its length about | of
that of the dorsal, with which it is coterminous. Anal spines very
small, closely adpressed to the first ra3s ; rajs similar and about
equal in length to those of the dorsal, the last much shorter than
the others. Pectoral lanceolate, the seventh ray longest, the rays
decreasing rapidly on each side, the lowest scarcely one-fifth as
long as the seventh. Length to tip of longest ray ly-H ^^ ^^^^
of the head. Most of the rays twice branched, tip of the longest
reaching a little bej'ond the anus. Base of pectoral slightl}'
oblique.
Veutrals inserted under the hinder margin of the pectoral base,
their tips not reaching to the anus ; their length about f that of
the pectoral ; the last four raj'S twice bifurcate.
Caudal about one-sixth of the total length, with numerous acces-
sory rays, causing a widening of the caudal base ; principal rays
three times forked ; hinder border deeply and triangularly emar-
ginate, almost forked.
Lateral line indistinct, tubes simple; about 145 scales in its
length, parallel or nearly so with the dorsal outline.
About forty scales between the ventrals and the lateral line, and
thirteen above the latter. Scales of body almost rectangular, their
longitudinal exceeding the transverse diameter, the free margin
finely ctenoid. All the scales small, those of the abdomen rather
smaller than the others, especially front of the paired fins.
Scales extending upon the cheeks and opercular apparatus, but
the snout and forehead to above the centre of the eyes, the upper
border of the orbits ; preopercular margin, jaws and gill-membrane
scaleless.
No scales ujion dorsal or anal ; caudal covered with small scales
over the greater portion of its surface. Pectorals more or less
scaly exteriorly near the base, the scales extending farther between
the central than between the lateral rays.
Color leaden-gray, becoming darker above, but fading to a dirty
creamy-white below. Vertical fins slaty-gray. Dorsal surface of
head darker than the rest of the body.
1880.]
NATURAL SCIENCES OF PHILADELPHIA.
IT
30.1 23^
16
25
5.1
81
12
101
24§
7-1
'3
321 29^
1.25 16
14|
The two specimens on which the above description is principally
founded were procured in the market of San Francisco, and were
l)rought from the vicinity of Monterej- Bay. One is an adult, the
other an immature individual, and the two present considerable
variation in external form, and in the proportions of some of the
parts, as will be evident by the dimensions and further description
of each specimen here appended.
Dimensions of the Two Specimens. No. 1. No. 2.
INCHES. INCHES.
Total length, including caudal, . . 17.75 10.05
Length without caudal, . . . 14.65 8.30
Greatest depth of body, . . . 4.50 1.96
Greatest thickness of body, . . . 2.38 .92
Length of head, 3.74 2.08
Circumference behind base of pectorals, 10.88
Longitudinal diameter of eye, . . .80 .52
Length of snout, 1.25 .63
Interorbital width, round curve of forehead, 1.75 .76
From tip of snout to dorsal, along dorsal
outline, 4.75 2.46
Length of base of dorsal fin, . . . 8.96 4.95
" " " spinous dorsal, . . 2.40
From tip of lower jaw to anal, along ab-
domen, 8.<^0 4.39
Length of base of anal, . . . . 5.02 3.32
Length of pectoral base, ... .87 .45
Lengtbof pectoral to tip of longest (7'-') ray, 3.36 1.82
From tip of snout to insertion of ventrals, 4.46 2.42
Length of ventrals, . . . 2.30 1.20
Height of first dorsal spine, . . . .25 .25
" " second " " . . . .56 ,40
" " third " " . . . .82 .55
" " fourth " " . . . .93 .68
" " fifth " " . . , 1.03 .70
" " sixth " " . . . 1.05 .73
" " seventh" "... 1.09 .75
" " eighth " " . . . 1.13 .77
" " ninth " " . . . 1.10 .70
Distance from 1st dorsal to lateral line, . 1 .50 .65
Height of soft dorsal, in front, . . 1.30 .90
Depth of anal, 1.30 .90
Width of caudal peduncle, . . . 1,12 .65
Length of lower jaw, . , , , 1,62 .82
" " maxillary, along its cui-ve, . 1.50 ,75
Rays of dorsal, ix-26 ix-25
"anal, ii-25 ii-25
541 511
40 40
23 22 .
301 29
151 14
If 3
4 5
H
61
8^
7
7.2 81
9
n
10^
10^
71
^
10
9
9
73-
11
10
n
18 PROCEEDINGS OF THE ACADEMY OF [1880.
^ Further Description of No. 1. — Snout veiy declivous, dorsal
outline in advance of the dorsal rising rapidl}'^, owing to a great
accumulation of adipose tissue about the upper part of the bod}^ ;
posterior part of dorsal outline regularly descending almost in a
straight line ; abdominal outline i-egularly curved.
Greatest depth a little less than four times ; head, 4| times in
the total length; greatest thickness, 1§ in the greatest depth.
Eye, 4-|J ; snout, 3, interorbital width (round curve of forehead),
2^ times in the length of the head. Caudal peduncle, four times
in the greatest depth. Distance from the spinous dorsal to the
lateral line, measured along the curve of the side, one-third longer
than the longest spine.
Denticulations of preoperculum rather blunt ; opercvdar spine
blunt.
Teeth somewhat irregular, canines less distinct than in the
o
vouno".
Anal spines short and weak, but stiff, and distinctlj^ recog-
nizable as spines ; the first very short, the second about half as
long as the first ray.
Lateral line less conspicuous than in the 3'oung.
Upper part of the head and along the line of the back approach-
ing a chocolate tint.
Vertical fins darker nearer the margin. No black spot above
pectoral axil.
The whole fish is exceedingly oily, and the abundant exudation
of this oil renders it exceedhigly disagreeable to handle.
Further Description of No. 2. — Dorsal outline from tip of loAver
jaw to vertical from posterior margin of eye, much less convex
than in the adult ; rise from thence to the origin of the dorsal
very slight ; a gradual descent in an almost straight line from
thence to the caudal peduncle. Abdominal outline regularly
curved to caudal peduncle. Greatest depth, 5g^ ; length of head.
4^ times in the total length ; eye, 4 times ; snout, about 3^ times
in the length of the head. Interorbital width, measured round
its curve, about one-fifth more than the length of the snout, or
2f in the length of the head. Caudal peduncle, 3 times in the
greatest depth.
Distance from the spinous dorsal, at its posterior part, to the
lateral line, nearl}' 1^ in the length of the longest spine, and less
1880.] NATURAL SCIENCES OF PHILADELPHIA. 1'.)
than one-third of the semicireumference of the body. Longest
(9th) dorsal spine, 2f in the length of the head.
Forehead and occiput transversely much less arcuate than in
the adult, the large deposit of fat on these parts in the latter being
absent in the young.
Opening of moutli slightly less oblique tlian in the adult, the
maxillary extending a little farther back. Teeth much as in the
adult, but the hindmost tooth in each jaw, but especially in the
upper, assuming more distinctl}' the proportions of a canine,
though still smaller than the front teeth.
Denticulations of operculum proportionate!}' more conspicuous,
and more acute than in the adult, opercular spine ending in three
denticulations.
Ninth dorsal spine, 6j times in the length of the fin, about 2i
in the greatest depth. Rays of soft dorsal about 2^^ in the great-
est depth, the antepenultimate ra}^ slightly produced. Anal spines
closel}' attached to the first ray, ver}' small, flexible, and scarcely
recognizable as spines.
A black spot above the upper axil of tlie pectoral ; upper parts
without the warm tint of the adult. No large development of
adipose tissue.
Since the above paper was written, a third specimen of Caulola-
tilus from the same localit}' has come into the possession of the
California Academy of Sciences. This example is about equal in
length to the larger of the two described, bnt the development of
fat upon the occiput is much less marked, so that its proportions
are very nearly those of the type of C. anomalus.
Although I am perfectly aware that specimens from the Gala-
pagos would be required to settle the question of the identity of
G. princeps with C. anomalus and C. affinis, I believe that the
comparison of these three examples, evidently all of one species,
and sharing among them characters relied upon as specific, cer-
tainly throws great doubt upon the distinctness of the three de-
scribed species. Dr. Bean {in lit.) doubts the specific identity of
the two specimens described in this paper, and draws attention to
certain differences of proportion, but the only differences of mag-
nitude are those caused by the development of fat on the occiput.
20 PROCEEDINGS OF THE ACADEMY OF [1880.
ON THE STRATIGRAPHICAL EVIDENCE AFFOKDED BY THE TERTIAEY
FOSSILS OF THE PENINSULA OF MARYLAND.
BY ANGELO HEILPRIN.
The Tertiary deposits of Maryland have from time to time
attracted the attention of investigators more or less eminent in
their special lines of research, the results of whose observations,
owing to the then imperfect state of American geological and pale-
ontological science, only very gradually tended to unfold the true
relations existing between the S3'nchronous formations of the
east-Atlantic and west-Atlantic countries.
Maclure, on the map accompanying his " Observations of the
Geology of the United States " (1817), classed all the late super-
tlcial deposits of Maryland imder the general term " Alluvial,"
which term was likewise applied to almost the entire border
deposits of the Atlantic and Gulf slopes. In 1824 (J. A. N. S.,
vol. iv) Sa}^ described about forty species of fossil shells collected
by Mr. Finch from the' same state, but excepting some passing
reflections on the nature of the deposit whence they were obtained,
and on the great resemblance existing between some of the forms
and forms still living on the coast, no special geological inferences
were drawn from the collection. From a comparative examination
of the contained fossils. Van Rensselaer (" Lectures on Geology,"
1825, p. 261) subsequently referred the deposits in question to
the Upper Marine formation, which view was concurred in by
Morton in a paper read before the Academy of Natural Sciences
of Philadelphia in June, 1828. In a previous paper (*' Geological
Observations on the Secondary, Tertiar}-, and Alluvial Forma-
tions," J. A. N. S., January, 1828), published conjointly by
Yanuxem and Morton, no attempt was made to correlate the various
divisions of the American and European Tertiary formations.
Conrad, who, more than any other American geologist, con-
tributed to advance our knowledge of the geology and paleon-
tology of this latest period, was the first to recognize the existence
of at least three distinct post-Secondary^ formations in Maryland,
the oldest of which he identified by a series of a few fossils found
near Ft. Washington, on the Potomac, as belonging to the Eocene,
and the newest, as exposed on the southeast extremity of the
peninsula, to the Post-Pliocene (J. A. N. S., vol. vi, and Bulletin
1880.]
NATURAL SCIENCES OF PHILADELPHIA.
of the National Institution, 1841). The intermediate deposits
were classed as the Upper Marine, but subsequently under Lyell's
designation of Miocene. Conrad's original observations were in
general confirmed by his later researches, and the relations of at
least a great portion of the Miocene of Maryland, as well as of
almost tlie entire Atlantic slope, were clearly pointed out V)y
Lj-ell in 1845 (Proc. of the Geolog. Soc, vol. iv, p. 547).
It is mainly in relation to this last formation that we wish to
draw special attention, there being but little question concerning
the original determination of the Eocene and Post-Pliocene
(Pliocene ?) deposits. That the great bulk of the deposits known
as the Medial Tertiary of Marjland are not sj-nchronous with
the South Carolina deposits classed by Tuomey and Holmes as
Pliocene, an assumed fact insisted upon by Conrad, and for which
there appears to be no evidence, an examination of the following
table of mollusca will clearly demonstrate :
Lameilibranchiata of the Medial Tertiary Formations of Maryland.
Leda concentvica.
Anomia epTiippium,* Cardita protracta,
Amphidesraa caxinata,* " granulata,*
" subovata, Cardium laqueatum,
Area callipleura,
(= A. dipleura?),
" idonea,
" incile,*
' ' subrostrata,
" Marylandica,
" triquetra,
" centenaria,*
" improcera,*
" stilicidiura,
Artemis acetabulum, *
" concentrica,
(=: A. elegansf],
Astarte vicina,
" cuneiformis,
" obruta,
" perplana,
" exaltata,
" varians,
" distans,
" planulata,
" undulata,*
Cardita arata,*
" acutilaquea-
tum,
" craticuloides,
" leptopleura,
Corbula cuueata,*
" idonea,
" elevata,
" inequalis,*
Crassatella Marylandica,
" turgidula,
" melina,
" undulata,*
Cytherea Sayana,*
" albaria,*
Lima pajiyria,
Lepton {?) mactroides,
Lucina anodonia,'^
(=L. Americana),
' ' Foremani,
" subobliqua,
" subplana,
'* cribraria,*
" crenulata,*
" contraeta,*
" divaricata,*
Mactra incrassata,
" ponderosa.
' ' fragosa,
" subcuneata,
" delumbis,
(= C. idonea), Modiola Ducatellii,
Marylandica, Mya producta,
Mytilus incurva,
Ostrea Virginica,*
" percrassa,*
Panopaa Americana,
" rettexa,*
" porrecta,
(=P. Goldfussi?)
" subnasuta,
Isocardia fraterna,
" Markoei,
Leda liciata,
" acuta,*
" (Yoldia) laevis,
" (Nucula) proxima,
22 PROCEEDINGS OF THE ACADEMY OF [1880.
Pecten Madisonius, Penia maxillata, Tellina lenis,
" Humpbreysii, Petricola centenaria,* Venus tetrica,
" Jeflfersonius, Plicatula marginata,* " Mortoni,
" concentricus, Pholadomya abrupla,* " alveata,
" Clintonius, Pholas ovalis, " inoceriformis,
" septenarius,* {= P. costata?)* " stamineus,
Pectunculus parilis,'^' Sazicava rugosa, " tridacnoides,*
" lentifor- Solen ensis,* " violacea,*
mis,* Tellina sequistriata, " Rileyi.*
" subovatus," " biplicata, '^
Tbe species in italics are still living on tbe American coasts; tliose fol-
lowed by an * are described by Tuomey and Holmes as occurring in the
Pliocene formation of South Carolina.
[Note. — The preceding table has been compiled as accnrately
as possible from the various papers pertaining to the paleontology
of the State, but owing to their number, and to the numerous
l)n1)lications in which they have been spread, it has proved impos-
sible to collect them all, and no doubt some few species will be
found occuriing in the State which have escaped our notice.
These will probably be ver}'^ few in number, and will not materially
affect the general conclusion. The following twenty-two species,
mainly those described by Say from the collection of Mr. Finch
(J. A. N. S., vol. iv), have no stated locality : Area centenaria,
A. improcera, A. incile^ Astarte distans^ Grassatella undidatd^
Leda acufa^ L. conceyitrica^ L. proxima, L. Isevis^ Lucina con-
tracta, L. divaricala. L. subobliqua^ Panopeea rejlexa, Pecten
Jeff'ej'sonius, P. Clintonius, P. concentricus, P. septenarius, Pec-
tunculus subovatus, Plicatula marginata, Tellina sequistriata,
Venus deformis (tridacnoides), and V. Rileyi.']
It will thus be seen, that of about one hundred species of
bivalves, only thirty-six (36 per cent.) are common to about an
equal number (105) from the South Carolina deposits; and
further, that, whereas, of the preceding enumeration of Maryland
mollusca only about fifteen per cent, are recent forms, no less
than forty per cent, (or according to Tuomey and Holmes, nearly
tifty per cent.) of the South Carolina Pliocene (Conrad's Miocene)
bivalve mollusca are still living. There remains, therefore, no
question regarding the relative ages of the two formations.
An examination of the fossiliferous strata exposed in sections
at various points on the western shore of Chesapeake Bay, in
1880.]
NATURAL SCIENCES OF PHILADELPHIA.
23
Anne Arundel and Calvert Counties, on the Patuxent River, near
Benedict, and on tlie St. Mar3''s River, St. Mary's Count}', tend
to show, moreover, that the series of deposits intermediate between
the Eocene of Fort Washington and the Pliocene of the south-
east extremit}' of the peninsula belong to two different periods of
formation, an older and a newer; those belonging to the latter
period being characterized by a fauna, the proportion of living
forms in which is far in excess of that in the former. Sections
of the newer deposits are exhibited in Calvert County, near Cove
I*oint, on the Patuxent River, below Benedict, at about water
level, on the same river, further north, in the deposits above the
Perna beds, and more especiall}' on the St. Mary's River, St.
Mary's County. The older deposits are best shown in the oyster
beds, rising a few feet above tide- water, at Fair Haven, Anne
Arundel County (which point was considered by Conrad as the
northern termination of the peninsular Miocene formation), in
similar beds, also only a few feet above water level, at a point
about twenty miles further south ('' Colonel Blake's," of Conrad),
in the sections exhibited by the Calvert Cliffs, and in the Perna
beds on both banks of the Patuxent River. There is, further,
strong, although not conclusive evidence, for considering the beds
containing Perna maxillata and Oatrea percrassa as the lowest
of the series.
The following tables exhibit as nearh* as possible the distribu-
tion of Lamellibranchiata in the deposits of both periods, those
of the newer being for conA^enience of comparison divided into
the Patuxent and St. Mary's groups :
OLDER PERIOD.
1 Area dipleura,
(= A. calilplenra ?),
2 " Marylandica,
H " subrostrata,
4 " triquetra,
.*> Artemis acetabulum,
H Astarte varians,
1 " exaltata,
8 Cardium craticuloides,
9 " leptopleura,
1 0 Corbula idonea,
1 1 Corbula elevata,
12 Crassatella melina,
13 " turgidula,
14 Cytherca subnasuta,
15 Isocardia Markoei,
16 Leda liciata,
n Lima papyria,
18 Lucina Foremani,
19 " subplana,
20 " crenulata,
21 Mytilus incurva,
24
PROCEEDINGS OF THE ACADEMY OF
[1880.
22 Modiola Ducatelii, 29 Perna maxillata,
23 Ostrea percrassa, 30 Pholas ovalis,
24 Panopseaporrecta(Goldfiissi) (= P. costata?)
25 Pecten Humphreysii, 31 Tellina lenis,
26 " Madisonius, 32 Veiius alveata,
2*7 Pectimculus parilis, 33 " staminea,
28 " lentiformis, 34 " 3Iortoni?
NEWER PERIOD. — J^ PATUXENT GROUP.
1 Anomia Conradi, 13
(= A. ephippium ?),
2 Area idonea, St. M., 14
3 Artemis acetabulum, St. M., 15
4 Astarte undulata, St. M. 16
5 Cardita protracta, 11
6 Cardium laqueatum, St. M.,
1 Carbula idonea, St. M., E., 18
8 Crassatella Marylandica, E., 19
9 Gytherea Saijana, St. M., 20
10 " Marylandica,
11 » albaria, 21
12 Isocardia fraterna, St. M., 22
Lucina Americana, E.,
(= L. Floridana),
Mactra incrassata,
Mya producta,
Panopsea Americana,
" porrecta (Gold-
fussi), St. M.
Pecten Madisonius, St. M., E.
Petricola centenaria,
Pholas ovalis,
(= p. costata ?), St. M.
Tellina biplicata, E.,
Vemis Mortoni ? St. M.
II. ST. MARy's GROUP.
1 Amphidesma carinata,f
2 " subovata,f
3 Area idonea,
4 " arata,f
5 " stilicidium,f
6 Artemis acetabulum,
T Astarte undulata,
8 " planulata,t
(^= A. perplana '().
9 '' vicina.f
10 Cardita granulata,t
11 Cardium laqueatum,
12 Corbula inequalis,f
13* " cuneata,!
14 Corbula idonea,
15 Gytherea Say ana ^
16 "
{Artemis) concen-
trica,f
{^ A. elegans /).
Isocardia fraterna,
Lucina cribraria,f
IT
18
19* Mactra ponderosa,f
20
21
22
23
24
25
" subcuneata,f
" fragosa,f
" delumbis,f
Ostrea Virginica,f
Panopjea porrecta,
Pecten Madisonius,
* Corbula cuneata and Mactra ponderosa are also found in the newer
deposits of Calvert Covinty, iiear Cove Point.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 25
26 Pholacloin3'a abrupta,t 30 Venus aloeata,
27 Pliolas arcuata, 31 *' Mortoni^
(= P. costata), 32 " tetrica,t
28 Saxicava rugosa,-f 33 " mercenaria^f
29 Solen ensis ?f 34 " inoceriformis.f
Note. — The italicized names represent species supposed to be
identical with living forms ; those (in the Patuxent group) fol-
lowed by the letters St. M. and E., species common to St. Mary's
and to Easton (Choptank River); and those (in the St. Mary's
group) followed by a f , species peculiar to the locality.
A comparison of the foregoing lists will show at a glance, that
of the thirt3^-four bivalves belonging to the older formations, at
most only three (or 9 per cent.) are found to be living forms
{Pholas ovalis [= P. costata?^^ Venus alveata, and Venus Mortoni),
and that only six (18 per cent.) and seven (21 per cent.) are corn-
common respectively to the Patuxent and St. Mary's exposures ,
viz. :
To Patuxent. To St. Mary's.
Artemis acetabulum,* Artemis acetabulum,
Corbula idonea, Corbula idonea,
Pholas ovalis,* Pholas arcuata (= costata),
Panopaea porrecta, Panopsea porrecta,
Pecten Madisonius, Pecten Madisonius,
Venus Mortoni, Venus Mortoni,
" alveata.*
* There appears to be much confusion regarding the species of Artemin
found fossil in the Atlantic tertiary deposits, and their relation to the
forms now living on the Florida coast. In 1833 ("Fossil Shells of
the Tertiary Formations," p. 20) Conrad characterized the species A.
acetabulum, which appears to have been until then confounded with the
A. concentrica, Con., non Borni^. discus, Reeve, "Conchologia Iconica,"
vol. vi, sp. 9), inhabiting the southern coast. No mention is there made
of its being found also in a recent state, but subsequently, 1838 ("Fossils
of the Medial Tertiary Formations," p. 29), we find the following statement :
"This fine species is very common in the localities named, and also occurs
recent on the Florida coast." In the list of shells inhabiting the Floi-ida
coast, prepared by the same author in 1846 (A. J. Science, 2d series, ii,
p. 393), only two species of Artemis are catalogued, A. elegans and A. con-
centrica, and it therefore appears highly probable that the statement con-
sidering A. acetabulum also as a living form was founded on a misconcep-
tion, the more especially, as an examination of the recent shells in the
3
26 PROCEEDINGS OF THE ACADEMY OF [1880.
Deducting two or three species that are also found at Eastoii,
we still have left twenty-three (or 68 per cent, of the whole
number) that are not found in the later deposits.
Museum of the Academy fails to reveal anything answering to Conrad's
original description. This species appears moreover to be identical with
the Venus concentrica described by Tuomey and Holmes in their work on
the Pliocene fossils of South Carolina (1857, p. 82), and to which Conrad,
apparently without good reason, applied the specific name of intermedia
{Dosinia {Artemis] intermedia) in his check list of Miocene fossils (Proc.
A. N. S., 1863, p. 575). The A. acetabulum is found fossil in the tertiary
deposits of Maryland, Virginia, North Carolina and South Carolina, and
must be carefully distinguished from tlie A. concentrica of Born, to Which
it bears only a distant resemblance. Another fossil species is probably the
A. elegans, Con. (living on the Florida coast) ; one almost perfect speci-
men, which agrees in all essential respects with the recent forms, is in the
Academy Miocene collections, but, unfortunately, the locality whence it
was obtained is not given. In his account of the geology and organic
remains of the peninsula of Maryland (1830, J. A. N. S., vol. vi, p. 312),
Conrad mentions the Cytherea {Artemis) concentrica, Lam., as occurring in
the St. Mary's exposure, but as subsequently ("Fossils of the Medial
Tertiary," 1838, p. 30), it is distinctly stated that the same does not occur
in the Miocene formation, it is highly probable that the original observa-
tion was erroneous. Certainly nothing corresponding either to the species
in question or to A. discus is to be found in the Maryland Miocene collec-
tion of the Academy.
The common species inhabiting the southern coast is not the A. concen-
trica of Born, with which it has been frequently confounded, and to which
it bears only a very slight resemblance, but the A. discus of Reeve {loc. cit. ).
A third species, the A. (Dosinia) Floridana Con., is unquestionably very
closely allied to the last, from which it differs essentially only in the
gi'eater obliquity of the pallial sinus. In other respects it agrees with the
figures and minute description of Born's species as given by Agassiz in his
'■^ Iconographie des Coquilles Tertiaires^' {JSlouv. Mem. de la Societr
Hehetiqtce, 1845, vol. vii).
I am disposed to consider the various forms of Ve7ius alveata and T^
latilirata as mere varieties of one and the same species, a series of inter-
mediate stages seeming to link them together. The V. athleta constituted
by Conrad to embrace the V. athleta of Say, V. latilirata of Tuomey and
Ilolmes, and the V. paphia of Lamarck, appears likewise to be nothing
but a variety of the same form. The V. alveata is included by Stimpson
among the living mollusca of the Atlantic coast (Smithsonian Check
Lists, I860;, but this fact appears very doubtful in the opinion of Try on
("American Marine Conchology," 1873, p. 160). It must be confessed,
however, that there exists a very striking agreement between the fossil
shell and specimens of the F. paphia, Linn., from St. Thomas, the main
1880.] NATURAL SCIENCES OF PHILADELPHIA. 27
On the other hand, the fossils of the newer deposits as exhibited
in the sections on the west bank of the Fatuxent show a verj'
decided similarit>' to those of St. Mary's, for out of tlie twenty-
two species of bivalves occurring there, no less than eleven ( or
just 50 per cent.) are also common to the last mentioned locality.
There can, therefore, I believe, be no reasonable doubt that the
deposits exposed on the Patuxent River immediately above the
Pernn beds constitute a direct continuation of the highly fossil i-
ferous strata bordering both sides of the St. Mary's River. These
last number among their fossil fauna also about thirty-four species
of Lamellibranchs, the same number as is found in what we have
designated as the older group, but of these thirty-four, about
twenty -two (or, deducting Corhula cuneata and Madra ponderosa,
twenty), or 65 per cent, are peculiar to the locality. Moreover,
of the entire number, about nine (or 27 per cent.) are still living
on the Atlantic coast. The dissimilarity of the two faunae cannot
fail to strike the least observant investigator, and Conrad has
dwelt at some length upon this curious manifestation (A. J.
Science, vol. xxviii, p. 282, and Bull. National Institution, 1841,
p. 176). That paleontologist singularly enough (apparently not
having made any exact numerical estimates either of the living-
forms, or of the forms found in one locality and not in the other) .
explains the differences as due solely to variable local conditions.'
difference being a tendency on the pai't of the latter to lose the full solidity
of its ribs some distance before they reach the posterior slope. The V.
alveata exhibits a similar tendency, but not quite to the same extent.
I have been unable to discover any description of the PJiolas ovalis.
Con., nor is there any mention made of it either in the Mioqene check
list prepared by Conrad in 1862, (Proc. A. N. S.), or in that of Meek, of
1864 (Smithsonian Miscell. Collections). I have, therefore, only doubtfully
referred it to P. costata.
' Thus he states (A. J. S. loc. cit.) : "If our coast were now suddenly,
elevated, we should find spots where the shells would consist chiefly of an
immense number of Modiola demissa mixed with Littorina littoirea and
Melampas bidentatus ; these are found on the margm. of the lagoons at
high water mark, the Modiola imbedded in a tenacious soil. At a little
distance would be found Venus mercenaria, Mya, avenavia^ Solen ensis,
Solecurtus Caribeus ; among these would be Ostvea. . ViTginiana, Fusus
cinereus, and a few of Pecten concentricus. Such is the group existing on
the sandy shore of the Estuaries. Hai'd by, would be a vast deposit of
oyster shells with Echinus, and immense masses, of Sevpula. These live
on the bottom of the lagoons, which is composed of a mixture of sand and
28 PROCEEDINGS OP THE ACADEMY OF [1880.
This interpretation might very satisfactorily account for the phe-
nomenon as far as generic distribution alone is concerned, and,
indeed, it would even hold good in its bearings on a limited number
of species, but it would hardly apply to a case such as the present
one, where the specific dissimilarity is so vast in such a compara-
tively very limited geographical extent.
Now, if the supposition that the deposits in question were
deposited at two different periods be a correct one, and paleon-
tological evidence goes far to prove that they were, we should
naturally expect to find also some direct stratigraphical evidence
afforded by the superposition of the strata themselves.^ The
following section was obtained by Conrad at a point on the Chesa-
mud. Then would be found another group of shells which live only in
deep water, the Astarte lunulata, Nucula limatula, N. proxima, Cardita
borealis, Pholas costata, in company with great numbers of Mytili. This
deposit we should recognize as having been formed in harbors, like those
of Newport and Charleston. ..."
It will be observed, that in the above conception Conrad has confined
himself entirely to generic and not specific disti'ibution.
' It may as well be remarked, that, although in the foregoing examina-
tion of the molluscous fauna I have dwelt exclusively upon the Lamelli-
hranchiata, the Gasteropoda offer equal, if not greater support to the
general conclusion arrived at. On comparing the lists of geographical
distribution given by Conrad in the Bulletin of the National Institution
(pp. 181-7), it will be seen, that not a single recent form occurs among
the eighteen enumerated from the Calvert cliffs at "Hance's;" and
further, that only two species, Valuta mutabiUs and V. solitaria, are
common to the forty-two found at St. Mary's. Of these last eight (or 19 per
cent.) were considered by Conrad to be recent forms :
Buccinum trivittatum, Natica duplicata,
" lunatum, Dentalium dentalis,
" quadratum, Fusus cinereus,
Natica heros, Scalaria clathrus.
Nearly all the species found on the west bank of the Patuxent also
occur at St. Mary's, and the same can be said of those collected in Calvert
county near Cove point the southern extremity. Singularly enough, that
although three species of Turritella — T. indenta, T. exaltata, and T.
perlaqueata—yf ere cdWected from Calvert cliffs in the upper portion of the
county, none of them appear to have been found near Cove Point, where
" vast quantities " of a new species, T. plebeia, "the common species of
St. Mary's River" .(loc. cit. p. 182), appear suddenly to make their
appearance.
1880.]
NATURAL SCIENCES OF PHILADELPHIA.
29
peake, near " Beckett's," about twenty-eight miles south of Fair
Haven :
Feet in Thickness.
5
3
Sand, without shells.
Sand, with innumerable shells.
20
Mingled sand and clay, without fossils, or very rare.
Same as below, less numerous.
Sand and clay, with a group of shells like that at Hance's.
The sliells obtained at Hance's, about four miles further north,
were the following
Bivalves.
Astarte varians,
" exaltata,
Artemis acetabulum,
Area subrostrata,
" dipleura,
Cythcrea subnasuta,
Cardium leptopleura,
Crassatella melina,
Corbula idonea,
" elevata.
Bivalves.
Isocardia Markoei,
Lima papyria,
Lucina Foremani,
" subplanata,
" crenulata,
Pectuneulus lentiformis,
Venus latilirata,
" Mortoni ?
" staminea.
Univalves.
Bonellia lineata,
Cancellaria biplicifera,
" engonata,
Dentalium tlialloides,
Fissurella Marylandica,
Yoluta mutabilis,
Infundibulum perarmatum,
Mavginella perexigua,
Pleurotoma Marylandica,
Univalves.
Pleurotoma bellacrenata,
Scalaria pachypleura,
Solarium trilineatum,
Sigaretus fragilis,
Trochus peralveatus,
Turritella indenta,
" exaltata,
" perlaqueata,
Voluta solitaria.
30 PROCEEDINGS OF THE ACADEMY OF [1880.
It will be at once noticed that in addition to the lowest fossil-
iferous stratum, extending to about seven feet above water level,
a second highly fossiliferous one manifests itself at a height of
about twenty-seven feet, in which were recognized among other
shells Artemis acetabulum and Pecten Madisonius. The mineral
character of this upper deposit is described by Conrad as being a
" quartzose sand, very incoherent," which is exactly what we meet
with in the arenaceous deposits on the west bank of the Patuxent
River, near Benedict, and which we have identified as equivalents
of the St. Mary's deposits. They are described by Conrad as
being composed of an " arenaceous, fossiliferous stratum," the
sand of which is "quartzose and incoherent" (B. N. I., p. 185).
We have thus exposed in one section two highly fossiliferous
strata, the upper of which shows a very decided analogy to what
we have designated as the newer group, and the lower of which
assumes a distinct personality for reason of its position, and the
paleontologieal characters impressed upon it. Proceeding from
this point southeastward, and therefore in the general direction
of the dip of the beds, we should naturally- expect to meet a point
where our upper stratum, or its equivalent, would descend nearer
to the level of the Bay, and in fact we do find just such a point
near Cove Point, where " the group most characteristic of these
tertiary deposits, imbedded in sand," descends to a height only
about fifteen feet above water mark (B. N. I., p. 183). The
fossils found here are also nearly all found at St. Mary's, and they
are, moreover, " highly ferruginous, as much so as many of the
crag fossils of Great Britain, which they greatly resemble, also,
in other respects " (Conrad, loc. eit.). On the St. Mary's River,
the southeasternmost extension of the formation, the same deposit
sinks almost to water level, as might well be expected on follow-
ing the general direction of the dip. Here, the Pliocene deposits,
well characterized by their fossils, make their appearance.
On proceeding from our first point almost due northwards, and
therefore at a considerable angle to the line of strike, we meet
with just the reverse phenomena met with on our southern
journey. At Fair Haven, where Conrad obtained the following
section,
1880.] NATURAL SCIENCES OF PHILADELPHIA. 31
Feet in Thickness.
50
Whitish Clay.
3
5
Bones of Cetacea.
Clay, with siliceous casts of marine shells and fragments of bones.
Clay, with Ostrea percrassa, Pecten Humphrey sii.
the highl}^ fossiliferous stratum found at water mark, at Beckett's,
is probably represented by a bed of clay three feet in thickness,
commencing at a height of five feet, and which contains " great
numbers of black, water-worn, siliceous casts of small shells,
chiefly Turritella, the species not yet determined." Below this
an entirely new deposit now makes its appearance, a bed of clay
of five feet thickness, characterized by Ostrea percrassa and
Pecten Humphreysii. This last, therefore, probably represents
the most ancient post-Eocene deposit exhibited on the Chesapeake.
Ostrea percrassa and Pecten Humphreysii were also found by
Conrad at Huntingtown, Calvert County, where in a " depression
or small valley " a race-way had been excavated through the
fossiliferous ''marls." The lowest member of the section was
" quartzose sand, with casts of Perna maxillata.^^ On the east
bank of the Patuxent River, moreover, near the mouth of St.
Leonard's Creek, Conrad observed innumerable casts of Perna
maxillata imbedded in a stratum of fine siliceous sand, and rest-
ing on tlie fragmentary rock considered by him as the " founda-
tion of the peninsula " (B. N. I., p. 184).
We should naturally look for some deposit contemporaneous
with that occurring on the west bank of the Patuxent, at some
point northeast of that locality where a section may present itself.
This we find at Easton, on the Choptank, where the mol-
luscous fossil fauna corresponds very closely with that observed
on the former river. The deposits of the older period, on the
other hand, reappear in Cumberland County, New Jersey, in the
" Miocene marl " of Shiloh, containing the following assemblage
of fossils (Cook, " Geology of New Jersey," 1868, p. 297) :
32 PROCEEDINGS OF THE ACADEMY OF [1880.
Bivalves.
Ostrea Mauricensis, Astarte Thomasii,
" percrassa, Venus Ducatellii,
Plicatula densata, Periploma alta,
Carditamera aculeata, Corbula elevata,
" arata, Saxicava myaeformis.
Craseatella melina,
Four species of the above are also found in Maryland, three of
which, Ostrea percrassa, Crassatella melina, and Corbula elevata,
are found, I believe, exclusively in the deposits designated as
those of the older period. None are recent forms.
The small percentage of living forms occurring in the " older
deposits," as compared with that of the " newer," leaves little
doubt for the inference that the deposits in question were formed
at two different periods, the latest of which clearly belongs to the
Miocene. A comparative examination of some of the peculiar
fossil forms of the older deposits, together with the extremely low
percentage of living forms, seems to indicate an age moi-e nearh'
Oligocene than Miocene, although perhaps not a single Eocene
species is represented. This last fact need not surprise us, howcA^er?
as the relationship of the Oligocene to the Miocene appears to be
greater in almost all the localities of its representation than to the
Eocene. The Eocene, moreover, of Maryland is represented only
by a very limited number of fossils, and Conrad, himself, has called
attention to the fact, that there appears to exist a greater amount
of difference between the Eocene and Miocene formations than
obtains between the Secondary and Tertiary, or between the
Devonian and Carboniferous systems (B. N. I., p. 177). The fol-
lowing comparison may serve to throw some light upon the rela-
tive age of the deposits in question :
Perna maxillata, Lam.
This species agrees thoroughly with the figure and description
of the same given b}' Goldfuss in the " Pectrefacta Germanise "
(vol. ii, p. 106), and to which the locality Weinheim (Oligocene)
is assigned. The sub-Apennine species, formerly classed under
the same name, is considered by Deshaj^es to be distinct, and he
has applied to it the specific name of Soldanii (Lamarck, '"'■ Animaux
sans Vertebres,"" 2d ed., vol. vii, p. 79). A second species of Perna,
the P. Sandbergeri, Desh., also occurs in the Oligocene locality of
1880.J NATURAL SCIENCES OF PHILADELPHIA. 33
Weinheim (Sandherger,^^ Conchylien des Maimer Tertiarheckens,''''
p. 367).
Mytilus incurva, Conr.
This large species of Mytilus may perhaps be taken as the rep-
resentative of M. Haidingeri^ Homes Q'-Fossilen 3IoUusken des
Tertidrheckens von Wie7i,^^ Abhand. d. k. k. geolog. Reichaanntalt^
iv, p. 356), found both in the Oligocene (Eggenbnrg) and Miocene
divisions of the Vienna basin. Rolle (Sifzungsberichte d. k. Akad.
d. Wissenschaften, 1859, p. 64) and Sandberger consider the M.
Tlaidingeri as the eqnivalent of M. Faujasi, Brongn., occurring at
numerous Olia-ocene localities of the Vienna and Mentz basins.
Isooordia Markoei, Conr.
Tliis Isocardia is, it appears to me, erroneously referred by
HVrnes {loc. cit., p. 165) to the /. cor. L., from which it is very
readily distinguished by its relatively much greater height, and
greater development of the umbones. It is a singular fact, that
this species of Isocardia was followed in the later period by the
T. fraterna,'^ Say, which is barel}' distinguishable from fossil
examples of the I. cor from Astigiana and Sicil}'.
It is worthy of remark, that Rolle (loc. cit., p. 81), as early
as 1859, only four years after Bey rich first applied the term Oligo-
cene to some of the middle Tertiary deposits of northern Germany,
hinted at the possible existence of the same formation on the
banks of the Patuxent, his conclusions being drawn from an
examination, among other fossils, of specimens of Lucina anodonta,
Say, Area idonea^ Conr., and Cardium laqueatum^ Conr.
* On comparison with si)ecimens from the English Crag this species will
be found to differ very broadly from the /, {Cyprina^ rustica of Sowerby,
with which it has been confounded,
34 PROCEEDINGS OF THE ACADEMY OF [1880.
CARCINOLOGICAL NOTES No. I.
BY J. S. KINGSLEY.
It is the intention of the writer in this series of notes to give
descriptions of new species, rectifications of synonj'my, facts
relating to geographical distribution, and other matters of im-
portance concerning the Decapoda. Unless otherwise stated all
specimens are in the collection of the Academy of Natural Sciences ^
of Philadelphia.
Genus PSEUDOTHELPHTJSA Saussure.
{Potamia Latr. et Boscia Edw. preoc.)
Fseudothelphusa latifrons.
Potamia latifrons Randall, Journal of the Academy of Natural
Sciences of Philadelphia, viii. p. 120.
Carapax smooth, regions and sutures indistinct. Frontal crest
very prominent, uninterrupted. Front reflexed, making with
the surface of carapax an angle of about 45°, its margin undu-
lating and its surface and margin granulate. From the front
arise processes which all but join the inferior margin of the orbit.
Superior margin of orbit crenulated. Anterolateral teeth more
prominent than in any other of the genus and extending back to
the posterior third of the carapax. Below, the carapax is ever}-
where granulate and especiall}^ so on the sub-branchial regions
and near the mouth. Inferior margins of orbits denticulate.
Chelipeds nearly equal. Anterior surface of meros granulate, as
are the outer portions of carpus and upper portions of the hands.
The dactyli with rows of small tubercles above.
The species is a true Pseudothelphusa, the antennae being as in
that genus, but the reflexed front gives it a peculiar appeai'ance
and with the larger anterolateral teeth will readily separate it
rom all other known forms. The emargination of the external
margin of the orbit is no more marked than in P. chilensis (Edw.
and Lucas) Smith, the type of which, by the waA', is in the
Museum of the Academy.
Pseudothelpusa sinuatifrons (A. M.-Elw.) Smith.
The locality of this species was not known to Alphonse Milne-
Edwards. There are two males in the Academy's collection from
San Domingo (W. M. Gabb}.
1880. J
NATURAL SCIENCES OF PHILADELPHIA.
35
Genus DILOCAECINUS.
Dilocaremus pardalinus Gerstsecker, Archiv fiir Naturgeschichte xxii,
p. 148, 1856.
Gerstaecker gives doubtfully South America as the habitat of
this species. There are specimens with the label " ? Upper
Amazon, Dr. Wilson."
Dilocarciaus spinifrons, nov.
Carapax regularly arcuate, regions obsolete, sides arcuate, armed
with four spines besides the spiniform angle of the orbit ; the
margins of the spines finely serrate. Superior margin of the orbit
obscurely crenulate, inferior denticulate with a strong spine near
the interior angle. Front advanced, with about fourteen spines.
A spine at the anterolateral angles of the buccal area. Chelipeds
sub-equal, meros with two spines at about the middle of the pos-
terior margin and a single one on the anterior margin at about
the middle, and one on the distal portion of the upper margin ;
the spine on the interior surface of the corpus long, slender, acute.
Hand with an acute spine above at the articulation of the dactylus,
fingers with the denticulations fine but acute. Ambulatory feet
less dilated than is usual in this genus. The spined front readily
separates this from all other species.
Upper Amazon, Dr. T. B. Wilson.
Genus THELPHUSA (including Geothelphusa Stm.)
Of this genus forty -five species have been described. The
localities from which I have examined specimens are marked with
an exclamation point (!).
africana A. M.-Edw. West Africa,
anchietae Capello. West Africa,
andersoniana Wood-Mason. Burmah.
anfustifrons A. M.-Edw. Australia,
aubryi M.-Edw.
West Coast Africa (!) ; Natal (!).
aurantia Ilerklots.
pelii Herklots.
atkinsoniana Wood-Mason.
Northern India,
aasteniana Wood-Mason. India,
bayonioa Capello. West Africa.
bayonica var. a, Capello. West Africa.
berardi Savigny.
Egypt, Nile (!); Red Sea.
ehilensis (Heller) A. M.-Edw. Chili,
corrugata Heller,
crassa A. M.-Edw.
oristata A. M.-Edw.
dehaani White.
berardi DeHaan.
japonica Herklots.
denticulata M.-Edw.
depressa Krauss.
difformis M.-Edw.
edwardsii Wood-Mason
flaviatilis (Bosc.) Latr. Mediterranean
Region, Greece (!), Gaarda Sea (!),
(Museum Peabody Academy).
grapBoides White. Manilla.
? siibqnadrata Gerst.
gondoti M.-Edw. Madagascar,
Madras, Java.
Australia.
East Indies(!).
Japan.
China,
Port Natal.
Red Sea.
Burmah.
36
PROCEEDINGS OP THE ACADEMY OF
[1880.
gaerini M.-Edw. India.
hispida Wood-Mason. Burmah.
hydrodromus Gerst.
indica Latr.
cam'cularis Westwood.
? aitranlia Gerstaecker.
? rotunda Freycinet.
infiata M.-Edw.
jagori von Martens.
Isevis Wood-Mason.
larnaudi A. M -Edw.
leschenaulti Edw.
India (!), Mauritius, Tahita.
lugubris Wood-Mason. India,
margaritaria A. M.-Edw. West Africa.
nilotica M. Edw. Nile.
India.
Pt. Natal.
Philippines.
India.
Siam,
obesa A. M.-Edw. Zanzibar,
obtusipes (Stm.) A. M.-Edw.
Japan, Philippine*,
perlata Edw. South Africa, Pt. Natal (!),
philippena von Martens. Philippines.
picta von Martens. Philippines*
planata A. M.-Edw. Bombay.
= ? guerini M. Edw.
siamensis A. M.-Edw. Siam.
sinuatifrons M.-Edw. Unknown,
stoliczkana Wood-Mason. Penang.
subquadrata Gerst.
= ? yrapsoides.
transversa von Martens. Australia,
tumida Wood-Mason. Burmah.
To this list I would add three more :
Thelphusa emarginata nov.
Carapax glabrous, longitudinally strongly arched. Post-frontal
crest continuous, nearly straight, obscurely orenulate, epibranchial
tooth obsolete, a tootli between the extremity of the post-frontal
crest and the angle of the orbit. Protogastric region very short,
front about one-fourth the width of carapax, slightly sinuate.
External angle of orbit slightly emargiuate. Anterolateral
margin cristate ; crest, however, soon becoming obsolete. Chelipeds
sub-equal, meros with the margins tuberculate and with a strong
spine on the distal portion. Upper and outer surface of carpus
with indistinct squamje, inner portion two-spined, the proximal
spine exhibiting a tendency to become bifid. Hands with the
upper margin obsoletely tuberculate, fingers roughened, not
gaping. Ambulatory feet slender, compressed.
Is very near T. depressa Krauss, but ditfers from that species in
the narrower and straighter front, the tooth just behind the angle
of the orbit, and in the non-gaping fingers of the chelipeds.
Length 34 mm., breadth 56 mm.
West Africa, Du Chaillu ; Fori Natal, Dr. T. B. Wilson.
The name is proposed on account of the emargination of the
orbit.
Thelphusa enodis nov.
Carapax smooth ; post frontal crest wanting. Epibranchial
tooth very small. Front narrow, strongly curved downward, its
margin concave, Chelipeds unequal, hands with the inferior
1880.] NATURAL SCIENCES OF PHILADELPHIA. 3T
margin regularly arcuate. Is very closely allied to T. Isevis, but
differs in the flatter carapax, the concave front, and the regularly
arcuate lo^j^er margin of the hands. In all other respects Mr.
Wood-Mason's description and figures (Journal Asiatic Society of
Bengal, vol. xl, p. 201, PI. xiv, fig. 1-6) would well appl}' to it.
Ceylon.
Thelphusa rugosa nov,
Carapax depressed, cervical suture and post frontal crest well
marked, the crest interrupted. Front nearly straight ; proto-
gastric region nearl}^ smooth ; epibranchial tooth small, directed
inward, lateral portions of carapax with transverse ruga? as in
mau}^ Grapsi, the margin of the anterolateral portion obscurely
crenulate. Chelipeds subequal ; the outer surface of meros and
carpus with squamose rugae, the rugae on the hands indistinct.
Carpal joints of the first three pairs of ambulator}^ feet with the
sides cristate ; dactjdi pointed.
Ceylon.
Length 26 mm., breadth 32 mm.
This species is nearest T. denticulata, but will be readily
identified from that species by the more crenulated margin
between the orbit and the epibranchial tooth, and by the rugae on
the lateral portions of the carapax.
Acanthocyclus gayi Edwards and Lucas.
The type of this species is in the museum of the Academ3^
38
PROCEEDINGS OF THE ACADEMY OF
[1880.
DESCRIPTION OF A F(ETAL WALRUS.
BY HARRISON ALLEN, M.D.
The Academy is the iiossessor of a foetal walrus, which was
presented by Dr. I. I. Hays, and brought by him from the Arctic
region of eastern North America. I have thought that a figure
with measurements of this rare, if not unique, specimen would be
of value.
The specimen is straight, or nearly so, and it is by this simple
test distinguished from other embrj^os of Carnivora. There is
neither flexure of the head upon the trunk,
or the trunk upon itself. The limbs are
folded close to the trunk, this feature being
most pronounced in the inferior pair, which
are inclined upward upon the ventral surface
of the body, and carry between them the
rudimentar}' tail. The median margin of the
first toe of the anterior extremity bears a
small, rounded membranous lobe, or lappet.
The muzzle exhibits the future position of
the vibrissas by six rows of minute papillae.
The muzzle projects slightly beyond the line
of the mouth. The position of the future
nostrils is seen by two slightly convergent
slits.
The vent is a semicircular slit-like opening
upon the lateral and posterior surfaces of a
rounded nipple-shaped organ, which is prob-
ably the future penis or clitoris.
The eye is closed, rather prominent, and
presents a palpel^ral fissure, which is directed obliquely upward
and forward.
The auricle is represented by a membranous fold laid close to
the head. The slit-like opening defining its position lies 3^'"
behind the e^-e, and extends slightly downwards and forwards.
The auricle extends in advance of this slit to the distance of 1'",
1880. J NATURAL SOIKNCES OF PHILAPEI.PHIA. 39
where it ends in a minute elevation. A pro])c can be readily
inserted in the slit, and can be passed forward.
The color of the specimen is a dull wliite or waxy.
No trace of hair is anywhere visible.
Lengtli of specimen. 1" 9'".
Length of head, 9".
Width of body at widest part, I".
Length of anterior margin of anterior extremity, 4^"'.
Length of posterior margin of anterior extremity, 2'".
Length of anterior margin of posterior extremity, 4"'.
Length of posterior margin of posterior extremity, A'"
Distance between vent and navel, 1^'".
40 PROCEEDINGS OF THE ACADEMY OF [1880.
ON THE NUDIBRANCHIATE GASTEROPOD MOLLUSCA OF THE NORTH
PACIFIC OCEAN, WITH SPECIAL REFERENCE TO THOSE OF ALASKA.
BY DR. R. BERGH, COPENHAGEN.
PART II.
DIAULULA, Bgh.
Diauhila, Bgh., Malacolog. Unters. (Semper, Philipp.II, ii), Ileftxiii, 1878,
p. 567 ; Heft xiv, 1878, p. xxxv. Gattungen nordischer Dorideu, Arch.
f. Naturg., xxxv, 1, 1879, p. 343.
Forma corporis subciepressa. Dorsum minutissime villosum, holo-
sericeum, molle. Tentacula digitiformia. Apertura branchialis ro-
tundata, crenulata ; folia branchialia tripinnata. Podarium antice
bilabiatum, labio superiore medio fisso.
Aimatura labialis nulla. Lingua rhachide nuda, pleuris multiden-
tatis, dentibus hamatis. Prostata magua ; penis inermis.
In their general form the Diaidulx^ somewhat resemble the Disco-
dorides and the TJwrdisee,'^ although their habitus still is peculiar.
The back is villous, as in these genera and especially as in the Thor-
disse^ but finer and more velvet-like. The tentacles are finger-shaped,
smaller than in the Discodorides, larger than in the Thordisse. The
branchial-f-lit is rounded, crenulated ; the branchial leaves tripinnate.
The anterior margin of the foot bilobed, the upper lip broader, with a
median fissure. As in the Thordisse^ there is no armature of the
lip-disk. The radula nearly agrees with that of the Diacodorides;
the rhachis is naked ; on the pleurae there is a rather broad series of
plates of the usual hook-shape. The stomach is enclosed in the liver
(not free, as in the Discodorides and in the Thordisse). As in the
Discodorides, there is a large prostate and an unarmed penis.
Only the following species appears to be hitherto known, from the
northern Pacific.
1. D. Sandiegensis (Cooper).
^ Diaulus, medicus, of. Martialis, I, 48, p. 40.
* Cf. my Malacolog. Untersuch. (Semper, Philipj^ II, ii), Heft xii, 1877,
p. 518, iDiscodoris) ; p. 540 {Thordisa).
1880.] NATURAL SCIENCES OF PHILADELPHIA. 41
1. D. Sandiegensis, Cooper. Plate V, fig. 3-9.
Doris {Aclinocyclus?) Sandiegensis, Cooper, Proc. of the California
Acad, of Nat. Sciences, ii (1862), 1803, p. 204;' iii (18G3); 1868, p. 58.
Color corporis e brunneo lutescens, annulis nigris maculatus ; vel
brunneus.
Habitat. Oceanum Pacificum orient. (San Diego Bay; Santa Bar-
bara ; Sitka Harbor; Puget Sound).
According to Cooper, numerous specimens of this species were
found from November to May among grass on mud flats in San Diego
Bay, at or near low water mark ; according to Cooper, it is a very
" active " species ; Cooper later obtained two specimens at Santa Bar-
bara Island, on rocks at low water. During the expedition to Alaska
a specimen was taken by Dall in Sitka Hai'bor, on alg«, in August,
1865, at the depth of six fathoms (another in August, 1873, in Puget
Sound, by Dr. Kennerly, on algae, at low water).
Through the kindness of Dall, I have seen the original (rather
rough) drawings of this species by Cooper; a colored one represents
the back bright chocolate-brown, with six black rings, of which there
are two smaller ones between the rhinophoria ; the rhinophoria, the
gill and the foot seem bright-yellowish ; one figure shows five, another
six branchial leaves.
The length of the first specimen, sent to me preserved in spirits,
was about 22.0 mm., the height reaching 9.0 mm., and the breadth
13.0 mm.; the breadth of the foot reached 10.0 mm., the height of
the rhinophoria 2.0 mm., the branchial leaves 3.3 mm. The color
was uniformly brownish-gray ; nearly symmetrically on each side of
the true back was an annular black spot.
The form of the rather soft body elongate-oval, not much depressed.
The head quite concealed between the mantle and the foot; the
outer mouth had the form of a vertical slit ; at each side a short
finger-shaped tentacle. The margin of the rather large rhinophor-
holes rather prominent, crenulate ; the rhinophoria strong, the club
' " Pale brownish-yellow, with large, annular, brown spots, irregularly
scattered, varying from twelve to twenty, or entirely brown. Surface
slightly rough ; sometimes a little tuberculated. Dorsal tentacles conical,
retractile ; branchiae large, rising in five jiarts, which become tripinnately
divided, expanding so as to cover the posterior third of the body like an
umbrella. Mouth proboscidiform, with two short lateral tentacles. Length,
3^ inches; breadth, 2| inches; height, ^ inch. — Coopeu, 1. c.
4
42 PROCEEDINGS OP THE ACADEMY OP [1880.
with about thirty leaves (on each side). The back all over minutely
and densely villous (fig. 3). The margin of the rather wide (5.0 mm.),
roundish branchial aperture like the margin of the rhinophor-holes,
prominent, finely crenulate ; the branchial leaves (retracted) six in
number, very strong, tri- or quadripinnate. The anus strong, about 1 5
mm. high, cylindrical, closing the branchial ring posteriorly ; the renal
pore as usual. The edge of the mantle rather thick, projecting about
2.0 mm. from the body ; the sides low. The genital opening as
usual, with two distinct apertures at the bottom. The foot strong,
broad, somewhat narrower towards both ends ; in the anterior margin
a strong furrow, towards the median line deeper and forming two lips ;
the superior broader and divided in the median line.
The cerebro-visceral ganglia kidney-shaped, the visceral larger than
the cerebral ; the pedal of roundish contour, scarcely larger than the
visceral. The buccal ganglia of oval form, connected by a short
commissure ; the gastro-oesophageal roundish, short-stalked, in size
about one-fifth of the former, with one very large and one large cell.
The eyes short-stalked, with black pigment and yellowish lens. The
otocysts scarcely smaller than the eyes, overcrowded with otokonia of
the usual kind. The leaves of the rhinophoria strengthened with long,
perpendicular spicula, calcified at the surface. The tentacula with a
mass of shorter, but otherwise similar spicules, lying irregularly.
The villi of the back closely set with perpendicular spicula (lig. 3).
The anal papilla with long, perpendicular spicules ; the stalk of the
branchial leaves with many shorter spicula, irregularly situated ; in the
leaves themselves were no spicules. In the interstitial connective
tissue large spicules were seen rather sparsely.
The oral tube was about 1.5 mm. long, wide, with strong longitudi-
nal folds. The bulbus pharyngeus only about 4.0 mm. long, by a
htvight of 2.0 mm., and a breadth of 4.0 mm. ; the rasp-sheath very
prominent on the hinder part of the under side of the bulbus ; the
inner mouth with a yellowish, not thin, cuticula. The tongue with
nine rows of teeth, in the rasp-sheath also eleven rows of developed
and two of not quite developed teeth, the total number thus being
twenty-two. In the posterior rows of the tongue the number of plates
was twenty-eight or twenty-nine, on each side, and seemed in the suc-
ceeding rows not to surpass thirty. The color of the teeth horn-
yellowish ; the height of the outermost 0.06 to 0.08 mm., the height
rising to about 0.18 mm. The form of the teeth as usual; the wing
rather narrow ; the innermost (fig. 5aa, h) not very different from the
1880.] NATURAL SCIENCES OP PHILADELPHIA. 43
Others (fig. .5, 6), the body of the outermost three or four (fig. iaa, T),
as usual, of reduced size.
The glanduliE salivales (5.0 or G.O mm.) long, in the anterior part
about one-third larger than in the rest, measuring 1.0 mm. in diameter,
yellowish ; in the rest of the length much narrower, whitish.
The oesophagus is about 9.0 mm. long, rather wide. The stomach
is included in the liver, not spacious. The intestine appears on the
surface of the liver in the usual manner, passing forwards, forming a
short flexure, and running straight backwards to the anal tube, which
has in its interior many fine longitudinal folds ; the total length of the
intestine about 20.0 mm., with fine longitudinal folds through its whole
length. The cavity was empty. The liver yellowish, about 17.0 mm.
long, by a breadth of 8.0 mm., and a height of about 6.0 or 7.0 mm. ;
the anterior end truncate, the posterior end rounded ; on the right side
of the forepart a flattened impression for the anterior genital mass.
The vesica fellea, as usual, behind and at the left side of the pylorus,
elongate-pyriform, grayish, taken together with its duct about 2.5 mm.
in length.
The heart as usual. The two gland, sanguines as usual, whitish ;
the foremost more triangular, about 3.5 mm. long; the posterior
broader, about 2.0 mm. long.
The gland, hermaphrodisiaca with a rather thick yellow layer
clothing the greater part of the surface of the liver (except the pos-
terior end) ; in the lobules of the organ were rather large o'jgene cells
and masses of zoJsperms The anterior genital mass large, com-
pressed, about 10.0 mm. long, by a height of 6.3 mm., and a breadth
of 3.0 mm. The ampulla of the hermaphroditic duct strong, grayish,
when unrolled about 25*0 mm. long, somewhat coiled on the anterior
end of the left side of the mass and on its inferior flattened edge be-
hind the large prostate ; it reaches a diameter of 1.2 mm. The male
branch of the ampulla (fig. 8a) thin, white, passing into the narrow
inferior end of the prostate, thus forms the fore-end of the whole
genital mass. The prostate (fig. 86) is of dirty yellow color, flattened
and irregularly pyriform, the length about 6.3 mm., by a breadth of
as much as 3.0 mm. ; the spermatoduct (fig. 8c) issuing from the
upper part of the posterior side of the gland, in its first thicker part
nearly as long as the prostate ; in the rest of its length thinner, mak- -
ing several coils and passing (fig. 9a) into the male organ. The re-
tracted penis (fig. Sd) strong, about 2.5 mm. long, the praiputium with
fine longitudinal folds (fig. 9), from the aperture upwards and nearly
44 PROCEEDINGS OF THE ACADEMY OF [1880.
filled by the glans, which had nearly the form of a human penis, with
a well developed head with round aperture ; this head seemed covered
with very small, low and rounded, soft papillae. The spermatotheca
were whitish, spherical, of the diameter of about 2.3 mm., filled with
epithelium, fatty matter and altered semen ; the chief duct a little
longer than the spermatotheca, gradually passing into the simple
vagina, that was about half as much in length (and was filled with
sperma). The spermatocysta of violet-gray color, somewhat flattened,
of oval outline, of the length of about 2.3 mm., filled with sperma.
The posterior half, or a little less, of the large mucous and albuminous
gland, chalk-white ; the anterior, more than half, of grayish or (on
the left side) yellowish color ; the structure as usual.
A variety of the species (according to Dall, it also belongs to this
species) was, moreover, obtained by Dr. Kennerly, in August, 1873,
on ulgas, at low water, in Puget Sound, Washington Territory
(fig. 6-9).
The single individual was rather large ; the length 40.0 mm., by a
breadth of 28.0 mm., and a height of 13.0 mm. ; the breadth of the
foot 15.0 mm., of the margin of the mantle 11.0 mm.; the height of
the rhinophoria 5.0 mm., of the branchial leaves nearly 5.0 mm. The
color of the upper side obscure olive-gray, with rather large (diameter
about 4.0 mm.) black and blackish spots ; the under side yellowish.
The general form and the head, with the tentacles, as above described.
The openings of the rhinophor-holes as above, the club with about
twenty-five leaves. The branchial opening as above (diameter, 3.5
mm.) ; the retracted branchial leaves six in number ; the anal tube
nearly 3.0 mm. high. The back villous, as in the typical individual.
The foot as above.
The peritoneum colorless, without larger spicula ; but in the region
of the ventricle of the heart the pericardium is brownish.
The central nervous system as above ; the proximal olfactory
ganglia bulbiform, a little larger than the buccal ; the distal ones
smaller than the proximal, at the root of the club of the rhinophoria.
The buccal ganglia of oval form ; the commissure between them
being about one-third of the largest diameter of the ganglia. The eyes,
the octocysts, the leaves of the rhinophoria and the villi of the back as
above.
The oral tube large, of a length and diameter of 4 0 mm. The
bulbus pharyngeus 4.0 mm. long, by a height of 4 and a breadth of
3.5 mm. ; the sheath of the radula less prominent than in the former
1880.] NATURAL SCIENCES OF PHILADELPHIA. 45
specimen ; the cuticula of the lip disk as above. The tongue with
ten rows of plates, further back eleven developed and two younger
rows, the total number thus twenty-three. In the posterior rows of
the tongue there were as many as thirty-four dental plates on each
side of the rhachis ; they resembled those above described (fig. 6, 7).
The salivary glands yellowish, ribbon-shaped. The stomach as
above. The anteriorly proceeding part of the intestine 7.0 mm. long,
by a diameter of about 2.0 mm. ; the receding part about 20.0 mm.
long, by a diameter of 1.5 mm. In the stomach and the rectum were
pieces of a Keratospongia and different Diatomacea. The liver 23.0
mm. long, by a breadth and a height of 11.0 mm. ; the anterior end
truncate, with a median deep and narrow slit for the oesophagus and
for the intestine ; the right anterior half of the liver rather excavated,
especially beneath ; the substance of the liver yellow.
The foremost glandula sanguinea about 4.5 mm. long, by a breadth
of 2.5 ; the posterior 4.0 mm. long, by a breadth of 2.5 mm. ; both very
flattened (about 0.8 mm. thick), grayish-yellow. The kidney with its
whitish network, contrasting prettily with the yolk-yellow hermaphro-
ditic gland ; the urinary chamber not wide ; the tube on its floor thin.
The hermaphroditic gland clothing nearly the whole liver (with its
posterior end), as in the former specimen. The anterior genital mass
about 11.5 mm. long, by a height of 9.5 and a breadth of 5.0 mm., the
ducts also projecting 3.0 mm. The ampulla of the hermaphroditic duct
yellowish-white, about 35.0 mm. long, by sr diameter of 1.25 mm., run-
ning across the upper part of the left side of the genital mass, and
forming several windings on the anterior part of the upper margin.
The large prostate as above (fig. 8b), dirty yellow ; 7.5 mm. long,
by a diameter at the upper end of about 4.0 mm. ; the part (fig. Sc),
from which the spermatoduct proceeds, much brighter than the rest of
the organ. The thin spermatoduct forming (fig. 8) a little coil at the
upper end of the penis; when unrolled about 12 mm. long. This last
(fig. 8rf, 9) organ strong, about 4.0 mm. long, by a diameter of 1.5
mm. ; the prominent orifice in the vestibulum (fig. 8e) with strong
longitudinal folds; the glans conical, filling nearly half (fig. 9) of the
cavity of the organ, the surface (under a power of 350) smooth. The
spermatotheca whitish, spherical, with a diameter of 3.5 mm. ; the
spermatocysta short, sausage-shaped, about 4.0 mm. long, of reddish-
yellow color. The duct from the spermatotheca to the vagina rather
thick, 3.5 mm. long ; the vagina larger than the penis, 6.0 mm. long,
by a diameter of 2.5 ; the inside with fine longitudinal folds, and with
46 PROCEEDINGS OF THE ACADEMY OF [1880.
sperma in the cavity. The mucous gland large, 9.0 mm. long, by a
height of 7.5 and a thickness of 4.0 mm. ; whitish, yellowish chalk-
white and yolk-yellow ; the duct rather short, with the usual strong
fold. The vestibulum with longitudinal folds.
JORUNNA, Bergh.
Jorunna, Bgh., Malacolog. Unters. (Semper, Philipp. II, ii) Heft x, 1876,
p. 414, note. Gatt. nord. Doriden, Arch, fiir Naturges., xxxv, i, 1879,
p. 34«.
Corpus subdepressum ; dorsum minutissime granulatum, sub-asperum,
branchia e foliis tripinnatis formata ; tentacula digitiformia ; poda-
rium sat latum, margine anteriore sulcatum, labio superiore latiore et
medio fisso.
Armatura labialis nulla. Radula rhachide nuda, pleuris multiden-
tatis, dentibus hamatis. Penis stylo armatus ; glandula et hasta
amatoria.
This genus was established by the author on the D. Johnstoni (1876)
in reference to the results of the anatomical examination of Hancock
and Embleton ; he regarded it as nearly allied to the Kentrodorides,
just founded by him.* After the present examination of the D. John-
stoni by the author he is not entirely certain of a generic difference
between the Jorunnce^ and the Kentrodorides. The latter have been
examined only from rather insufficient material, and the hasta has not
been seen in any of the species, only a papilla in connection with a
peculiar gland ; still the Kentrodorides are of a quite different habitus,
very soft, and the upper lip of the anterior margin of the foot is more de-
veloped, while the innermost plate of the tongue is somewhat different
from the others. If not identical with the Kentrodorides, the Jorunnce
are certainly very nearly allied to them.
The Jorunnce are rather depressed ; the back finely granulated,
covered with equal minute papillulog ; the retractile gill formed of tri-
pinnate leaves ; the tentacles digitifoi-m ; the foot rather broad, deeply
grooved in the front margin, and the upper lip of this larger and cleft
in the middle line. The lip-disk not armed, covered with a simple
cuticula. The rhachis of the radula naked, the pleurae with many
hook-formed plates. In the vestibulum genitale are four apertures :
^ R. Bergh, Malacolog. Unters. (Semper, Philipp. II, ii) Heft x> 1876,
p. 413 427, Tab. XLIX-LI.
^ Jorunna, Bjornis filia. Laxdiila-Saga. Hafniae, 1826, p. 21.
1880.] NATURAL SCIENCES OF PHILADELPUIA. 47
one for the penis, which is armed with a stylus ; another for a hasta
amatoria, through which opens a peculiar gland (quite as in the genus
Asteronotus) ;i a third for the vagina, and the fourth for the duct of
the mucous gland.
Only one species of the genus seems hitherto known, belonging to
the northern part of the Atlantic Ocean. The spawn of the species
is known from Alder and Hancock, but nothing else is known of the
biology of the animal.
1. J. Johnstoni (A. et H.).
Doris Johnstoni, A. et H. Oceauum Atlantic, septentr.
Jorunna Johnstoni (A. et H.). Plate VIII, fig. 19 ; Plate IX, fig. l-ll.
Doris Johnstoni, Alder et Hanc. Monogr. Part I, 1845, fam. 1, PI. .5 ; Part
V, 1851, fam. 1, PI. 2. f. 8-11.
Doris Johnstoni, Hanc. et Embleton, Anat. of Doris. Philos. Trans. 1853,
II, p. 212, 215, 216, 220, 233, PI. XII, f. 2, 10 ; PI. XIV, f. 9, 10 ;
PI. XV, f. 1-2 ; pi. XVII, f. 2-3.
Doris Johnstoni, Forbes and Hanley, Hist, of Br. Moll., Ill, 1853, p. 564.
.' Doris tomentosa, Cuv., Fischer. Jouru. de Conchyl., 3me Ser., x, 1870,
p. 290 293 ; XV, 1875, p. 211, note.
.' Doris tomentosa, C. Verany, catalogo. 1846, p. 16-21. Ver., Hanc. et
Embleton, 1. c. 1852, p. 220.'^
? Doris tomentosa, C. Philippi, En. Moll. Sic. I., 183, p. 104 ; II, 1844, p.
79, Tab. XIX, f. 9.
Color flavescens, dorso interdum maculis fuscis seriatis ornatus ;
rhinophoria fusco-maculata ; branchia albescens.
Hab. Oceanum Atlanticum septentr.
This species, that was first described by Johnston under the name
of D. ohvelata (Miiller), was (1845) established by Alder and Han-
cock. Hancock gave a series of anatomical remarks upon this very
interesting form and of figures referable to it. Since then nothing
new seems to have been published about the species ; but a few months
ago I (1. 0.) gave a short notice of the generic characters of the
group.
Of this form I have only examined a single specimen, captured in
March, 1870, in the neighborhood of Hellebak, on the north coast of
Seeland (Denmark).
^ R. Bergh, Ueber das Geschlecbt Asteronotus, Ehrbg. Jahrb. der Deut-
scheu Malakozool. Ges., iv, 1877, p. 161-173, Taf. I-II.
•^ According to Hancock and Embleton (1. c, p. 220), the dart (hasta
amatoria) in Doris Johnstoni is straight, in D. tomentosa, Ver., curved.
48 PROCEEDINGS OP THE ACADEMY OF [1880
The specimen was of a uniform yellowish color ; the rhinophoria
finely dotted with brown (but not the branchial leaves). The length
of the rather contracted and somewhat contorted individual was about
18.0 mm. by a greatest breadth of 10.0 and a height of about Y.O mm. ;
the height of the (retracted) rhinophoria 2.5, of the tentacles nearly
1.5, of the (retracted) gill 2.5 mm. ; the greatest breadth of the mantle-
margin 3.5 mm., of the foot 5 0 mm.
The form is elongate-oval, the mantle-margin rather thick, not very
broad. The back covered all over with very minute granules, some-
times, especially on the middle of the back, crowded in irregular and
roundish small groups ; the under side of the mantle-margin smooth.
The (contracted) openings of the rhinophor-holes appear as a simple
transverse slit, the granules of the back reaching forward to the open-
ing, those in this neighborhood not larger than the rest. The club of
the rhinophoria stout, with about thirty^ broad leaves. The opening of
the gill-cavity small, transverse, triangular-crescentic, with the convexity
forwards (as contracted) ; the granules of the back reaching to the very
margin of the gill-slit, but not larger than the rest. The gill consisting
of eleven branchial leaves,^ five lateral pairs and an anterior unpaired
leaf; the anal tube low, truncate, nearly central; the renal pore at its
right side. The head rather small ; the tentacles digitiform, somewhat
flattened. The sides of the body nearly imperceptible ; the genital
opening contracted. ^ The foot rather strong, somewhat pointed at the
end ; the anterior margin with a deep furrow, the superior lip rather
strong and prominent, cleft in the median line.
The peritoneum with very fine dark points (brown -black) spread
everywhere ; entirely without true spicules.
The central nervous system showed the cerebro-visceral ganglia
somewhat elongate, thicker and broader in the posterior part, nearly
not excavated in the exterior margin ; the pedal ones of oval form,
larger than the visceral. The olfactory ganglia very short-stalked,
bulbiform, a little smaller than the buccal ; a small optic ganglion, the
optic nerve short. At the inferior side of the posterior part of the
right visceral (fig la) ganglion is a short-stalked (fig. lb) ganglion
genitale giving off several nerves, one of them has at its root another
ganglion (fig. Ic). The common commissure not longer than the
' Alder and Hancock mention merely ten to fifteen leaves.
^ Alder and Hancock mention fifteen leaves.
' The representation of the penis (?) (1. c. PI. 5, f. 3) by Alder and Haa-
cock cannot be correct.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 49
transverse diameter of the pedal ganglion, rather strong. The buccal
ganglia of roundish form, connected through a very short commissure ;
the gastro-tiesophageal ganglia short-stalked, reaching scarcely one-
quarter of the size of the former, with one very large and some
smaller cells.'
The eyes with black pigment and shining, horn-yellow lens. The
otocysts at the slight emargination at the outer margin of the cerebro-
visceral ganglia, crammed with otokonia of the usual kind. The broad
leaves of the rhinophoria stiffened in the usual way by long, much
calcified spicula, perpendicular on the free margin of the leaves. The
skin of the back crowded with spicula,^ mostly very large and much
calcified; in the rather low (height 0.5 mm.) granules (fig. 2) crowded
erect spicules. In the interstitial tissue of the intestines true spicula
are neither many nor large.
The mouth-tube about 2.0 mm. long, strong, rather wide, quite as
usual. The bulbus pharyngeus .3.0 mm. long, with a height of 2.8
and reaching a breadth of 2.5 mm. ; the rasp-sheath also projecting
1.0 mm. from the hindermost part of the under side of the bullus. The
form of the bulbus and its retractors as usual ; the lip-disk whitish,
clothed with a yellowish cuticula. The tongue of usual form ; on the
shining horny-yellow radula eleven rows of teeth, further backwards
twelve developed and four younger rows ; the total number of rows
thus twenty-seven. 3 The teeth of yellowish color ; the height of the
outermost 0.06, of the next 0.08 mm. ; the height reaches at most
about 0.22 mm. The two foremost rows were rather incomplete, in
the fourth row were twenty-four, and the number of teeth then in-
creases to twenty-seven.^ The rhachis (fig. 3a) rather broad. The
plates of the usual form,^ with the usual wing-like expansion of the
exterior part of the body and of the root of the hook (figs. 4, 5) ; the
first (fig. 3) with lower hook, which on the succeeding teeth slowly
1 This representation of the central nervous system in most points agrees
with that of Hancock and Embleton (1. c. p. 233, PI. XVII, tig. 2, 3).
2 Collingwood (Annals and Mag. of N. Hist., 3 Ser., Ill, IS^jQ, p. 462)
mentions the spicules of this species (from the estuary of the Mersey) as
"very elegant, consisting of a broad embossed ph\te with a double and
beautifully serrated edge, tei'minating abruiitly in a blunt apex."
" Alder and Hancock mention twenty-four rows, whereof eleven were on
the tongue.
■• Alder and Hancock mention twenty-five plates in the rows.
^ Cf. my Malacolog. Unters. (Semper, Philipp. II, ii), Heft XIV., 1878,
(Asteronotus), p. 036.
60 PROCEEDINGS OF THE ACADEMY OF [1880.
increases in height ; then the teeth keep the same height and decrease
again in the outer part of the rows (fig. 5) ; the four to six interior
teeth are more erect, with shorter body and thinner hook (figs. 5, G).
The salivary glands long, thin, whitish.^ The esophagus about
6 mm. long, rather wide, with strong longitudinal folds.- The stomach
small, included in the liver ; the biliary apertures as usual.
The intestine issues through the liver behind the region of junc-
tion of the first and second third of the liver ; the first anteriorly pro-
ceeding part lodged in a groove on the superior side of the liver, not
passing beyond the antei'ior margin of that organ, about 2.5 mm. in
length ; the rest of the intestine about 10.0 mm. in length ; the diameter
of the intestine 0.8-1.3 mm. ; the longitudinal folds rather strong.
The liver of yellowish color, more grayish on the surface; 9.0 mm.
in length, by a breadth of 5.5 and a height of 4.0 mm. ; the posterior
end rounded ; more than the anterior half of the under side, especially
its right part, is excavated (for the anterior genital mass) and behind
this is a deep transverse groove. The vesica fellea lying at the left
side of the offshoot of the intestine, rather small, in height about 1.25
mm., reaching nearly to the surface of the liver, nearly cylindrical.
The heart as usual. The sanguineous glands whitish, rather
flattened ; the anterior obliquely triangular with the point, as usual,
adhering to the under side of the junction of the two cerebral ganglia ;
in length 2.0 by a breadth of 1.5 mm. ; the posterior transversely
elongate-oval, with a breadth of 3.5 by a length of 1.5 mm. The
renal syrinx melon-shaped, its largest diameter about 0.75 mm. ; its
free duct nearly three times as long ; a strong continuation of it pass-
ing along the floor of the rather large renal chamber, to the region of
the pylorus.
The hermaphroditic gland spread in large groups of ramifications
over nearly the whole liver and by its brighter yellowish color some-
what contrasted with it ; in its lobules were masses of zoJsperHis and
rather small ojgene cells. The anterior genital mass^ in length 5.0 by
a breadth of 2.5 and a height of 4.0 mm. ; the right side rather con-
vex, meeting the more flattened left side at the sharp superior margin,
^ They are in this way also mentioned by H. and E. (1. c, p. 215, PI, XII,
fig. 2cc) .
2 The dilatation on the oesophagus mentioned and figured by H. and E.
(1. c, p. 215, PI. XII, fig. 2d) could not be seen in the specimen examined
by me.
« Cf. the PI. XIV, f. 9, of Hancock and Embleton.
1880.] NATURAL SCIENCES OF PUILADELPHIA. 51
the under side flattened. The ampulla of the hermaphroditic gland
resting on the superior posterior part of the genital mass, whitish,
making a large curve, about 5.0 mm. long, with a diameter of nearly
1.5 mm. The spermatoduct in its first part, as near as could be de-
termined, rather thick than thin, not very long, forming (fig. He, 7e)
a little coil on the upper end of the penis. The penis (fig. 7/) cylin-
drical, curved, about 2.5 mm. long, by a diameter of about 0.8 mm. ;
the inside with many longitudinal folds ; at the upper end of its cavity
a low truncated conical prominence (fig. 115), with a rather wide
aperture (fig. lib), through which opens a little bag (fig. 11), whose
inside was clothed with a thin yellowish cuticula, and contained a
hollow, nearly colorless tube, that could be extended by tension ; it
was probably pointed (the point seemed broken ofl') ; its length was
about 0.9 mm. ; the spermatoduct opened (fig. 11a) in the upper part
of this bag. Hancock has (1. c. PI. XIV, fig. dc, 10; PI. XV, fig.
1, 2) seen the penis and the "stiletto," but he too seems (1. c. p. 220)
not at all clear about these organs. At the side of the opening for
the penis in the vestibulum genitale was another aperture which led
into a bag, from whose bottom projected a hard, whitish, somewhat
compressed conical spur (fig. Id, 10), that under the influence of nitric
acid grew more pellucid, but developed very little gas ; through the
axis of the organ down to the fine aperture on the point, passes a
slender tube (fig. 10), the continuation of the fine coiled duct of the
gland of the organ. ^ This gland (glandula hastatoria, fig. 7e, Sd)
overlies the upper part of the vagina (fig. 1a,i) ; it is heart-shaped,
of a transverse diameter (breadth) of 2.0, and a thickness of 1.0 mm.;
the gland did not contain any larger cavity. The spermatotheca (fig.
8a) whitish, nearly spherical, having a largest diameter of 2.5 mm. ;
filled with fatty cells and detritus ; the two ducts (fig. 8e, e) as usual,
the vagina rather wide (fig. 7a, b), with longitudinal folds on the in-
side. The spermatocysta yellowish, spherical, 1.5 mm. in diameter
(fig. 8b), filled with zoV.sperms ; short-stalked. The mucous gland
not forming quite half of the anterior genital mass, consisting of a
smaller anterior biconvex part, and a large flattened wing-like poste-
rior part ; the space between them nearly filled by the spermatotheca
> These organs, the gland and the spur, have also been seen (1. c, PI.
XV, fig. 9) by Hancock, but lie does not mention them (in the text, and
explanation of the figures). In another of his figures (fig. 105) the spur
is designated (1. c., p. 248) as "male intromittent organ, ' and the (fig.
lOe, /) true penis as "penis-like organ furnished with a stiletto "
52 PROCEEDINGS OF THE ACADEMY OF [1880.
4
and the spermatocysta, the color of the gland yellowish-white, on the
left side of the anterior part a central yellow mass; the duct of
the mucous gland rather short.
All the former genera of Dorididce belonored to the larfje "roup of
Dorididee cryptohranchiatse ;^ the following are to be registered in the
group of Dorididce eleiitherohranchiatse (D. phanerohrancliiatse).
This section is also characterized by the non-retractility of the gill,
by a sucking-crop connected with the bulbus pharyngeus and by a
peculiar armature of the tongue, consisting usually of a single large
lateral plate and a single or several outer plates. This group seems
chiefly limited to nortliern climes, and contains at present the genera
Akiodoris, Acantlwdoris, Adalarid, LameUidoris, Goniodoris and
Doridunculus,'^ also Ancula, Drepania^ and Idalia.
AKIODORIS, Bcrgh
Akiodoris, Bgh. Gattungen nordisclier Dorideu, 1. c, 1879, -p. 354.
Forma ut in LameUidoridibiis. Nothseum supra granulosum.
Branchia non retractilis, e fuliis tripinnatis non multis et ad modum
ferri equini positis formata. Caput latum, veliforme ; tentaculis
brevibus, lobiformibus. Apei turte rhinophoriales integrse.
Discus labialis sine armatura. Ingluvies buccalis bulbo connata.
Radula rhachide quasi nuda ; pleuris dentibus lateralibus depj'essis non
multis; (12-13) quorum duo intimi fortiores, quasi subhamati. Penis
glande uncis simplicibus, furcatis vel palmatis armatus. Vagina in-
dumento valloso peculiari instructa.
The animals belonging to this group resemble externally especially
the LameUidorides. The back is finely granulated ; the head large,
veil-shaped, with short tentacles, which are lobate and pointed The
openings of the rhinophor-holes with plain margins, surrounded by
several larger papilla?. The non retractile branchia nearly horseshoe-
shaped, consisting of a mediocre number of leaves. The lip-disk
^ Cf. my "Gattungen nordisclier Doriden," 1. c. p. 341.
^ The genu?, Doridu7iculus of G. O. Sars fMoll. regionis ai'cticse Xorveg.,
1878, p. 309. Tab. 27, fig. 2a~d, Tab. XIV, fig. 5), whicli externally ap-
proaches Goniodoris and other Dorididce eleutherobranchiaton in the char-
acter of the radula, is hitherto only known from the northeas'ern part of
the Atlantic (Lofoten), and by a single species {D. ecliinulatus, S. ).
^ In the Ancula and Drepanice the penis is armed as in so many Dorididce
with a series of small hooks.
1880.] NATURAL SCIENCES OF PHILADELPHTA. 53
without armature. The tongue with transverse thickenings of the
rhacliis ; the lateral plates somewhat depressed ; the two first different
from the rest, larger and with a denticle at the root of the hook ; the rest
without any such, the external quite without a liook. A sucking-crop
on the upper side of the bulbus pharyngeus, but sessile, depressed con-
ical, and not consisting of two symmetrical halves. The large
stomach free on the surface of the liver. The glans of the long
penis with a strong and quite peculiar armature, consisting of strong
hooks, partly simple, partly bifurcate and partly digitate, with strong
digitations. The vagina with a peculiar armature of high palisades.
This interesting genus externally most resembles the LameUidorides,
both in reference to the nature of the back, to the form and size
of the gill and in the want of armature of the lip-disk ; the region of
the openings of the rhinophor-holes differ in the want of a glabella
and by the presence of a larger number of surrounding papilte. The
genital opening somewhat recalls the Acanthodorides, as do also the
(tripinnate) branchial leaves and the sucking-crop, but this is not
divided in two distinct halves as in this last genus. The armature
of the tongue is very different from that of the LamelUdorides,
Adalarise and Acanthodorides ; the large hook-formed lateral plates
of these genera are wanting, and in their places are two large de-
pressed lateral plates, with small hooks ; the external plates somewhat
recalling those of the Adalarise ; the rhachis rather broad, with
transverse thickenings of the cuticula, corresponding to the rows of
plates. In the very peculiar form of armature of the glans penis, and
by the peculiar clothing of the vagina, the Akiodorides differ from all
the above-cited genera.
Only a single species of the genus is hitherto known, the new one,
that wnll be described below.
1. Ak. lulescens, Bgli., n. sp. Oceanum Pacificum.
1. Ak. lutescens, Bgh., n. sp. PI. IV, fig. 3; pi. V, fig. 11-U: pi. VI, fig. 1-20;
pi. VII, fig. 1-8; pi. VIII, fig. 1-2.
Color lutescens.
Habitat. Oceanum Pacificum septentrion. (Nazan Bay).
Of this form I have had a large single specimen for examination,
obtained in August, 1873, by Dall, on stony bottom, at low water, in
Nazan Bay, Atka Island, Aleutians.
According to Dal), the color of the living animal was "yellowish-
white;" preserved in spirits, it was of a uniform dirty yellowish color.
54 PROCEEDINGS OF THE ACADEMY OF [1880.
The length was 32.0 mm., by a breadth of 19.0 mm., and a height of
13.0 mm, ; the breadth of the foot 12.5 mm., of the mantle-brim 3.0
mm. ; the height of the rhinophoria 3.0 mm., of the branchial leaves
2.5 mm. ; the length of the genital opening 2.25 mm.
The form was elongate-oval, somewhat larger than that of the Lam.
hilamellata. The papillae of the back relatively smaller and more
rounded than in that animal. The openings of the rhinophor-holes
an oblique oval slit ; the margins plain ; several (six to eight) larger
papilliB (of about 1.0 mm. in height) in the immediate vicinity of the
holes ; the club of the rhinophoria with about thirty leaves. The
branchia with about ten leaves. The anal papilla low, with a stellate
aperture ; the renal orifice as usual ; the interbranchial space crowded
with rather pointed and high papillae. The head and tentacles as in
allied forms. The genital papilla of oval form, with a large, longitu-
dinal, crescentic slit. The rather broad foot with the usual anterior
marginal furrow. The peritoneum colorless, without spicula.
The central nervous system more flattened than in allied forms ;
the cerebro- visceral ganglia reniform, a little broader in the anterior
part ; the pedal ganglia less flattened than the former, larger than the
visceral ones, of oval form, on the outside of the cerebro-visceral. The
proximal olfactory ganglia a little smaller than the buccal ones, bulbi-
form ; distal ganglia could not be found. The commissure not broad,
not short. The buccal ganglia of oval form, closely connected ; the
gastro-oesophageal roundish, rather long-stalked, in size about one-
sixth of the former, with one large cell and several (three or four)
smaller ones.
The nervi optici rather long; the eyes with yellowish lens and
black pigment. The otocysts in the usual place, filled with otokonia
of the usual kind. The leaves of the club of the rhinophoria very
richly furnished with thick (diameter, 0.04 mm.) and long spicula,
more or less calcareous, and very often giving off a thick twig of
greater or less length (PI, V, fig. 12) ; for the most part set perpen-
dicularly or obliquely on the free margin of the leaves. The axes of
the organs and the short stalk stuffed with strong and very much cal-
cified spicules. In the skin of the back a mass of spicula of the same
kind (PI. IV, fig. 13) as above, or still more hardened ; the papillie of
the back solidified in the usual way (PI. V, fig. 11). In the interstitial
tissue fewer and smaller spicules.
The oral tube rather short, wide. The bulbus pharyngeus of usual
form, about 5.5 mm, long by a height of 4.5 ram., (and at the upper
1880.J NATURAL SCIENCKS OF PHILADELPHIA. 55
part of the sucking-crop of 5.5 mm.), and a breadth of 4.75 mm. ;
the sheath of the radula projecting about 1.3 mm. backwards and
downwards. The lip-disk large, clothed with a thick yellow cuticula ;
the true mouth forming a narrow vertical slit. The cap-shaped suck-
ing-crop almost exactly as in Ac. pilosa, but more conical and with-
out external signs of duplication : on the inside clothed with a yellow-
ish cuticula, opening into the buccal cavity through a wide slit. The
tongue rather broad ; on the fine reddish-yellow colored radula seven-
teen rows of teeth, also on the point of the tongue were traces of six
entirely vanished rows ; the two first rows very incomplete, reduced to
some external plates. Further backwards were seen forty-two devel-
oped and three younger rows, or, all in all, the animal presented sixty-
two rows of teeth. The most external plate of each row is quite
colorless, the next two or three pale yellowish, the following all of
horny-yellow color ; the rhachis colorless. The length of the most
external plate about 0.0.35 mm., of the next about 0.05 mm., of the
following 0.07 mm. ; the length of the second large plate about 0.2
mm., of the first 0.022 mm. ; the breadth of the rhachis about 0.22
mm. The rhachis thickened between the rows and forming arched
elevations between them (PI. VI, fig. la, 3 ; PI. VIII, fig. la). The
first two plates rather large (PI. VI, fig. Ibb, cc, 4-6; PI. VIII, fig.
16, c) ; with a short strong hook and a stout denticle at each side of
it, the outer denticle broader ; the hook of the second plate somewhat
larger than that of the first ; sometimes a slight crenulation on the
outer margin of the first plate (fig. 5). All the following ten or eleven
plates (PI. VI, fig. 2e, /; PI. VIII, fig. 2a, b) of the same type, by
degrees decreasing in size, consisting of a quadrilateral basal part,
from which (PI. VI, fig. 7-13), in most of them, rises a strong, short,
broad hook ; the two or three outmost plates (PI, VI, fig. 2/; PI. VIII,
fig. 2) formed of the basal part alone ; the rest with the hook gradu-
ally more developed
The salivary glands yellowish-white, flattened, ribbon-shaped, of
about 10.5 mm. in length, reaching to the cardia, where they
are agglutinated one to another ; the breadth in the foremost part
about 0.75 mm , in the middle 1.5 mm., the posterior part again some-
what narrower ; the duct of the gland rather short.
The oesophagus rather wide, about 13.0 mm. long, the inside with
rather strong longitudinal folds ; it opens into the stomach, which lies
free in a cleft on the upper side of the liver. This organ (PI. VI, fig.
17a) is of oval form, of about (3.5 ram. largest diameter; the inside
56 PROCEEDINGS OF THE ACADEMY OF [1880.
with rather strong longitudinal folds; the pylorus (fig. 17) in the
neighborhood of the cardia. The intestine advancing from the stom-
ach to the fore-end (fig. 176) of the liver, in this part about 10.0 mm.
long ; forming a knee and retrograding to the anal nipple in a length
of 23.0 mm. The contents of the stomach were indeterminable animal
matter, mixed with some diatomacea;.
The liver 20.0 mm. long by a height of 10.0 mm. and a breadth of
about 12.0 mm. ; the posterior end rounded ; a little more than the an-
terior half of the under side obliquely flattened (by the anterior genital
mass) showing the cardiac end of the a?5ophagus and the root of the
hermaphroditic duct. On the anterior part of the upper surface is a
cleft for the stomach and for the biliary sac ; the color of the surface
and of the substance of the liver is grayish-yellow. The biliary sac
(fig. 17c) lying before the stomach, on the right side of the intestine,
large fas the stomach), somewhat flattened, grayish, of rounded out-
line and about 4.5 mm. largest diameter ; the contents, as in the
stomach.
The heart as usual. The sanguineous gland whitish, entirely cover-
ing the nervous system, about G.O mm. long, by a breadth of 4.5 and a
height of only 1.0 mm.
The hermaphroditic gland yolk-yellow, covering the upper side of
the liver with a thick layer ; in its lobes large o'jgene cells and masses
of zoJsperms. The anterior genital mass large, about 14.0 mm. long
by a breadth of 9.0 and a height of 11.0 mm., flattened and a little ex-
cavated on the left side, with an excavation on the fore side, the right
side very convex. The hermaphroditic duct whitish, rather thin
(diameter about 0.75-1.0 mm.), passing straight over the left side of
the genital mass to its anterior end, without formation of any (distinct)
ampulla. The first part of the spermatoduct whitish, forming several
long windings on the upper part of the forepart of the mass and pass-
ing into the yellowish (PI. VI, fig. 18a) continuation ; this, with its
numerous coils, forms a large flattened layer on the fore-end of the
right side of the mass ; it then rather suddenly passes into a much
thinner whitish continuation (fig. I8b) about 6 mm. long, that slopes
(fig. 18c) into the penis, which (retracted) was lying on the lowest
anterior part of the right side of the mass. The penis was cylindrical,
of the length of 11.0 mm. by a diameter of 1.5 mm. ; the truncated,
cylindrical, yellowish (under a magnifier nodulous) glans forming
(PL V, fig. 13, 14) a prominence of the length of nearly 1.0 mm. in
the vestibulum. This glans was partly covered on the outer side
1880.] NATURAL SCIENCKS OP PHILADELPHIA. 57
(fig. 13, 14), but especially on the margin of the wide, gaping orifice
and on its inside for a length of about 4.0 mm. (PI. VII, figs. 2-4),
with rather crowded and apparently irregularly set claws. The claws
were very strong and for the most part broad and high (fig. 3, 4),
even reaching a height of about 0.3 mm. (fig. 4). In the interior of
the glans, especially in its posterior part (fig. 5c), the claws were less
broad and simply uncinate or bifurcated, otherwise mostly broader and
with digitations of the margin. The body of the claws was plain or
curved ; the end simply pointed, bi- or trifurcate or with digitations,
sometimes very strangely formed. They consisted of a cuticula and
its matrix ; very often, especially on the outside of the glans, the
cuticula was torn off and the (fig. 20) rounded or pointed naked matrix
was left. The whitish spherical spermatotheca (PL VI, fig. 19a) was
about 3.5 mm. in diameter, laterally communicating tnrough a short
petiolus adhering to tne upper end of the vagina, with a sinuosity intO'
which opens the elongate, yellowish spermatocysta (fig. 19i), which had
a length of about 2.0 mm., and from which issues the long duct of the
mucous gland (fig. 19c). The grayish vagina very strong (fig. 18e),.
about 7.0 mm. long, elongate-conical; the lowest part wide, having a
diameter of about 3.25 mm. ; the walls thick, with a very peculiar
internal lining, consisting of cylindrical palisades (PI. VII, fig. 6-8)
of a height of about 0.4 by a greatest diameter of 0.07-0.08 mm. ;
between the larger were seen smaller and very small ones. The pali-
sades seemed to be densely clothed (fig. 8) with cilia, and showed a
nearly colorless axis (fig. 6, 8) up to their points ; the axes were often
denuded (fig. 6) after the sheath has been torn away. This lining
continued up to the superior end of the vagina, but not beyond it.
The mucous gland large, whitish, and yellowish-white ; the anterior
half yolk-yellow, denuded on the fore-end of the genital mass ; the duct
short.
A variety (PI. VI, fig. 14-20) of this species has also been found
by Dall, in July, 1873, at low water, in Kyska Harbor (Aleutians).
According to Dall the color of the living animal was " yellowish." T he
animal preserved in spirits was of a uniform light yellowish color. The
length about 18.0 mm. by a breadth reaching 8.0 mm and a height of
6.0 mm. ; the breadth of the foot at the fore-end 5.0 mm., the margin
of the mantle freely projecting 1.5 mm. ; the height of the rhinophoria
' 1.5 mm., of the branchial leaves 1.5 mm. Around the plain margins
of the rhinophor-holes seven to nine large conical tubercles ; the club
of the rhinophoria with about twenty leaves Around the branchial
5
58 PROCEEDINGS OF THE ACADEMY OF [1880.
ring, as well as in the centre of it around the vent, rather large conical
tubercles 1.5 mm. in height; the branchial leaves, fifteen in number,
as far as could be determined.
The oral tube strong, 4.5 mm. long, wide. The bulbus pharyngeus
about 5.5 mm. long, by a height of 3.0, and a breadth of 3.75 mm. ;
the rasp-sheath about 1.75 mm., freely projecting, bent upwards. The
cuticula of the lip-disk yellowish. The tongue with about thirty-five
rows of plates (fig. 14-16) ; further backwards, twenty-five developed
and four younger rows ; the total number of rows sixty-four On the
posterior part of the tongue fourteen plates, the number increasing
backwards to fifteen or sixteen. The five anterior rows very incom-
plete, only represented by 1, 7, 9, 10, 12 plates (on each side). The
plates as above. The breadth of the rhachis reaching to about 0.17 mm.
The glandulae salivales 6.0 mm. long. ^The stomach (fig. 17o) about
4.0 mm. long. The contents of the digestive cavity a mass of sponge.
The vesica fellea (fig. 17c) about 2.5 mm. high, with strong folds on
the inside. The anterior genital mass quite as above, also the sper-
matotheca and the spermatocysta (fig. 19j, the penis (fig. 18, 20), and
the vagina (fig. 18, 19).
LAMELLIDORIS, Alder et Hancock.
Lamellidoris, A. et H., Monogr. Brit. Nudibi". Moll., Part VII, 1855, p. xvii.
Lamellidoris, A. et H., R. Bei-gh, Malacolog. Untersuch. (Semper, Philipp.
II, ii). Heft xiv, 1878, p. 603-615.
Lamellidoris, A. et H., R. Bergh, Gatt. nord. Doriden, 1. c, 1879, p.
362-365.
Corpus vix depressum, nothoeo granulate. Branchia (non retracti-
lis) e foliis (multis) simplicita pinnatis, ut plurimura in formam ferri
equini dispositis, formata. Caput latum, semilunare, angulis tentacu-
laribus. Aperturte rhinophoriales, margine integro ; tuberculis anticis
2-3, calvitie postica.
Cuticula apertura? oralis infra asserculis duobus incrassata, et ante
annulus papillarum angustus. Lingua rhachide lamellis humilibus in-
structa ; pleuris dente interno hamiforrai permaguo et externo com-
presso lamelliformi unco minuto pra^dito armatis. Ingluvies buccalis
(suctoria) petiolo bulbo pharyngeo connata, tympaniformis.
Penis apice (glaude) curvatus, non armatus. Vagina brevis.
The genus Lamellidoris was established (1855) by Alder and Han-
cock, to receive two small groups of Dorididee., one with the D. hilam-
ellata as type, to which especially the name of the group is here
1880.] NATURAL SCIENCES OF PHILADELPHIA. 59
restricted ; and the other, characterized by a more depressed form and
the naked rhachis of the tongue, with the B. depressa, A. et H., as
type. Hancock has given some anatomical remarks on the typical
form {D. bilamellata, L.) ; but nothing else had been since made
known about these animals' until my just cited notice and those of G.
O. Sars.2
The Lamellidorides approach the Acanthodorides, but differ even
here, externally, by the coarsely granulated surface of the back and
by the larger number of the branchial leaves, which are set in the
form of a horseshoe ; the openings of the rhinophor-holes, the tenta-
cles as well as the genital opening are also of a different shape.
More notable still are the anatomical differences ; the Lamellidorides
want the armature of the lip-disk, which is found in the other group ;
the armature of the tongue is quite different (1, I — 1 — I, 1), and the
buccal crop is connected with the bulbus pharyngeus by a stalk. The
penis is quite different from that of the Acanthodorides, and without
true armature ; the vagina is short. After all the Lamellidorides are
much more allied to the Adalarice.
The form of the body, as in the Acanthodorides, not very depressed.
The back covered all over with semi-globular and short club-formed
papillae. The openings of the rhinophor-holes with plain margins and
1 According to H. & A, Adams (the Gen. of Recent Moll., II, 1858, p.
657), Lamellidoris is a synonym of ^^ OncMdoris, Blv.," which name is
employed by Adams for a group, whose type should be D. pusilla, A., ec
H. (that scarcely belongs to the true Lamellidorides). Cf. also Gray,
Guide I, 1857, p. 307.
The genus Onchidoris of Blainville (Man. de Malac, 1825, p. 489,
PI. XLVI, f. 8.), ought to be rejected entirely, as founded very likely only
on bad observation ; the genus figures with nearly impossible characters,
both in relation to the tentacles ("quatre teutacules comme dans les Dori»,
outre deux appendices labiaux") and to the anus ("median a la partie
inferieure et posterieure du rebord du manteau"). The type of the genus
Blainville found in the British Mus. (London), where it seemed to have
disappeared, at least it was not to be found in the collection of nudi-
branchiates which I looked over in May, 1873 (while, on the coutraiy, I
found the long-lost type of the genus Linguella, Blv., in his original glass,
and so have re-established the denomination Linguella for the much later
(1861) Sancara, Bgh. Cf. my Malacolog. Unters., Heft vi, 1874, p. 248).
Later, Mr. Abraham (1. c. p. 235) seems to have found the original speci-
men again.
■^ G. O. Sars, Moll. reg. arct. Norv., 1878, p. 306. Tab. XIII, figs. 5, 6 ;
Tab. XIV, fig. 2, 3.
60 PROCEEDINGS OP THE ACADEMY OF [1880.
commonly two larger papillae before and a bare space behind them.
The gill (not retractile) consisting chiefly of several (usually 20-30)
tripinnate leaves, set in the form of a horseshoe. The head large,
veil-formed (semilunar), with produced and pointed side-parts, which
are adherent to the foot nearly to the point. The genital openings
not being a slit, but on a large tubercle.
The cuticula of the oral aperture is thickened below, near the median
line, into a ledge ; and on the outside is a ring of hard papillae.
The buccal crop, connected through a petiolus with the foremost part
of the upper side of the bulbus pharyngeus, is drum-shaped ; on the in-
side clothed with a strong cuticula. The tongue has on the rhachis
short compressed lamellce, on each side of these is a very large up-
right plate with large compressed body and a hook which on the inside
is either plain or denticulated j at the outside of this plate is another,
compressed but much smaller and with a little rudimentary hook. The
salivary glands forming a short, coiled mass at each side of the root
of the oesophagus. The oesophagus without diverticle at its origin.
The spermatoduct (as in the Acanthodorides) very long ; the penis
short, its glans curved and clothed with a rather thick cuticula, but
otherwise not armed. The spermatocysta imbedded in the mucous
gland ; ^ the vagina short.
About the biological relations of the animals belonging to this
group very little is hitherto known. Where the species occur, they
seem to be rather abundant in individuals (cf. about the Lam.
bilamellata, Collingwood, in Ann. Mag. Nat. Hist., 3 S. Ill, 1859, p.
463). The spawn of several species (L. bilamellata, L. diaphana, L.
inconspicua, L. aspera, L. depressa, L. pusilla) has been described
by Alder and Hancock, and that of a single species (Z. muricata) by
Sars, Meyer and Moebius, etc. The first stages of the development
of this last form have been followed by Sars ^
The group seems limited to the northern part of the Atlantic and
of the Pacific. To the same belong with certainty some properly ex-
amined species, and, besides, several others mentioned in the litera-
ture can, with more or less probability, be referred to it.
' The spermatocysta has not been seen by Alder and Hancock. Cf. 1. c,
1852. PI. XIV, fig. 8 (p. 219).
2 Archiv. fur Naturges, 1840 p. 210, Tab. 7.
1880,] NATURAL SCIENCES OF PHILADELPHIA. Gl
A.
I. L. Mlamellata (L.). Oc. Atlant.
2- L. varians, Bgh., n. sp. Oc. Pacif.
3. L. Jiystricina, Bgh., n. sp. Oc. Pacif.
4. L. muricata (O. Fr, Miiller). Oc. Pacif.
5. L. diaphana (Aid. et Hanc). Oc. Atlant.
B. diaphana, A. et H., Moiiogr. Part ii, fam. 1, PI. 10 ; Part vii, PL
46 suppl. fig. 9.
6. L. aspera (A. et H.).^ Oc. Atlant.
D. aspera, A. et H., 1. e., Part v, fam. 1, PI. 2, fig. 15 ; Part vi, fam.
1, PI. 9, fig. 1-9; Part vii, PI. 46, suppl. text ; PI. 48, suppl. fig. 2.
B.
7. L. sparsa (A. et H.). Oc. Atlant.
D. sparsa, A. et H., 1. c, Part iv, fara. 1, PI. 14 ; Part vii, PI. 46,
supi>l. test.
8- £. depressa {A. et H.j. Oc. Atlant.
D. depressa, A. et H., 1. c, Part v, fam. 1, PI. 12, fig. 1-8 ; Part vii,
PI. 46, suppl. fig 12.
f Villiersia scutigera, d'Orb., Mag. de Zool., 1837, p. 15, PI. 109, fig.
1-4.
9. L. ineonspicua (A. et H. ). Oc. Atlant.
D. ineonspicua, A. et H., 1. c. Part v, fam. 1, PI. 12, fig. 9-16 ; Part
vii, PI. 46, suppl. fig- 13.
10. L. oUonga (A. et H.). Oc. Atlant.
D. ohlonga, A. et H., 1. c, Part v, fam. 1, PI. 16, fig. 4-5; Part vii,
PI. 46, suppl. fig. 10.
II. L. pusilla (A. et H.). Oc. Atlant.
D. pusilla, A. et H., 1. c. Part ii, fam. 1, PI. 13 ; Part vii, PI. 46,
suppl. text ; app. p. iii.
12. L. luteocincta (M. Sars).* Oc. Atlant.
13. L. (?) ulidiana (Thomps.). Oc. Atlant.
D. ulidiana, Th., Ann. Mag., Nat. Hist., xv, 18, p. 31.
D. ulidiana, Th., Aid. et Hanc, 1. c. Part vii, p. 42, app. p. ii.
14. L. (?) tenella (Agassiz). Oc. Atlant.
D. tenella, Ag., Gould, Rep. ou the Inv. of Massachusetts, ed. Binney,
1870, p. 329, PI. XX, fig. 289j 290, 293.
15. L. (f) pallida (Ag.). Oc. Atlant.
Z>. pallida, Ag., Gould, 1. c, p. 229, PL xx, fig. 284, 287, 288, 291.
' According to Morcb (Synopsis Moll. mar. Daniae, Vidensk. Meddel. fra
naturh. Foren. i Kbhvn., 1871, p. 179) this species ought to be identical
with the D. muricata of Meyer and Moebius ; but this is, of course, im-
possible.
■^ The organs of the bulbus pliaryngeus of this species have just been
figured by G. O. Sars (MolL reg. arct. Norv., 1878, Tab. xiv, fig. 3).
62 PROCEEDINGS OF THE ACADEMY OF [1880.
16. L. (?) diademata (Ag.). 6c. Atlant.
D. diademata, Ag., Gould, 1. c, p. 230, PI. xxi, fig. 298, 300, 301-304.
17. L. [?) grisea (Stimps.j. Oc. Atlaut. Gould, 1. c, p. 232, PI. xx, iig.
292, 295.
18. L. (f) derelicta (Fischer). Oc. Atlant.
B. derelicta, F., Jouiu. de conch yl., xv, 1867, p. 7.
19. L. (?) iuberculata (Hutton). Oc. Pacif. (Nova Zeland.).
Onchidoris tuberculatus, Hutton, cf. Abraham, 1. c, p. 226.
20. L. (?) eubalia (Fischer). Oc. Atlant.
Doris eubalia, F., Journ. de couchyl., xx, 1872, p. 10.
1. L. bilamellata (L.), var. jooci/ica. Plate V, fig. 10; Plate XT, fig, 3-9.
Color albido-flavescens, maculis fuscis plus minusve variegatus.
Dentes laterales margine Itevi.
Hab. Oc. Pacific, sepentr. (Mar. Beringi).
Six specimens of this variety of the Atlantic species were taken by
Dall, in Bering Sea (Hagmeister Id.), in August, 1874, at low water,
on a gravel beach. Three were sacrificed for the anatomical examin-
ation.
According to Dall, the color of the living animal was "yellowish-
white with brown macule."
The length of the specimens preserved in spirits was 11-13.0 mm.
by a height of 4.5-5.5 mm. and a breadth of 6-10.0 mm. ; the height
of the rhinophoria 1.75-2.2, of the branchial leaves 1-1.2 mm, ; the
breadth of the foot at the fore-end about .5-8.0 mm. ; the margin of
the mantle projecting freely about 1.5-2.0 mm. The color of the
individuals on the back was yellow-white, marmorated with light
reddish-brown, this marbling always occupying the spaces between the
tubercles, which are nearly white (or light yellowir-h) ; the branchial
leaves of the same reddish color ; the club of the rhinophoria yellowish-
white ; the under side of the body yellowish-white or whitish.
The form was elongate-oval. The head flattened, nearly semicircu-
lar, with the tentacular edges a little prominent. The vicinity of the
posterior margin of the rhinophor-holes plain, at the anterior two
large erect tubercles ; the club of the rhinophoria with about twenty
leaves, the stem rather short. The back covered all over with semi-
globular and short club-shaped rounded tubercles of different sizes,
mostly small, mixed with many larger ones 0.'I5 mm. in diameter ; the
larger tubercles mostly showing a spinous surface (PI. V, fig. 10)^ when
magnified.
o
1 Cf. my "Malacolog. Unters." (Semper, II, 11) Tab. LXVIII, fig. 15-16.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 63
The openings of the rhinophor-holes and of the branchial area (fig.
Sbb) surrounded by large and small tubercles which also were spread
over the central part of it (fig. 3). The branchial leaves ( fig. 3aa) were
about twenty-four or twenty-five in number, set in a transverse reni-
form ring ; the leaves in the front part much larger than the rest.
The anus as usual, scarcely projecting. The under side of the margin
of the mantle quite smooth. The genital openings always quite con-
tracted. The foot large, with a fine line along its anterior margin.
The cerebro-visceral ganglia short-reniform ; the pedal ones not
much smaller, of oval form, set nearly at a right angle to the inferior
face of the former ; the olfactory ganglia bulbiform or ovoid. The
buccal ganglia rather flattened, of roundish contour, a little larger than
the olfactory ones ; the commissure between them very short ; the
gastro-oisophageal ganglia not very short-stalked, roundish, in size
about one-quarter of the buccal ganglia, with three large cells. The
three commissures very distinct, the sub-cerebral and the pedal con-
nected throughout most of their length ; the visceral thin, not giving
off a genital nerve.
The eyes with black pigment, yellowish lens ; the nervus opticus
nearly as long as half the breadth of the cerebral ganglion. The
otocysts as large as the eyes, crowded with otokonia of the usual kind.
The leaves of the rhinophoria without spicules ; the axis of these organs,
on the other hand, were filled with such spicules, partly circularly and
concentrically arranged. The tubercles of the back stuffed with ordi-
nary spicules (fig. 10) in the usual way, the larger spicules mostly very
prominent on the surface
The oral tube as usual. The bulbus pharyngeus of the usual form,
about 2.0 mm. long ; the lip-disk with a rather thick yellowish cuticula,
and inwards with the same belt of (about ten to fifteen) rows of small
denticles as in the L. hystricina (cf. below) ; the sheath of the radula
somewhat bent upwards, freely projecting behind the bulbus for as
great a length as that of the bulbus itself. The tongue (in the three
individuals) with ten or eleven series of plates, in the sheath ten or
eleven developed and tliree younger rows ; the total number of rows being
thus twenty four or twenty-five. The plates light j^ellowish in their
thicker parts, otherwise nearly colorless. The length of the median
plates reaching about 0.12 mm., the height of the external ones
0.10 mm. The median (fig. 7a) and exterior plates (fig. Ih) quite as
usual ; the large ones of the usual forms (fig. Ih), sometimes, especially
64 PROCEEDINGS OF THE ACADEMY OF [1880.
the foremost, with rather obtuse point (fig. 9). The buccal crop (fig.
4, 5) as large as the bulbus, of quite the usual form, rather petiolate.'
The salivary glands forming (on each side) a large, thick, whitish
mass between the bulbus and the central nervous system (with the
glanduloe sanguineje).
The oesophagus rather wide. The stomach and the intestine as
usual. The liver as usual, much flattened on the right anterior half
The heart rather large. The gland, sanguineae large, whitish, cover-
ing the upper side of the central nervous system, the foremost part in one
individual very narrow. The renal syrinx about 1.0 mm. long, with
strong longitudinal folds, its clothing as usual.
The anterior genital mass 4-4.5 mm. long by a breadth of 1.25-1.5
and a height of 3-3.3 mm., yellow-white, plano-convex ; the anterior,
and partly the superior portion formed by the coils of the whitish sper-
matoduct ; in one individual one coil embraced the sheath of the radula.
The first part of the spermatoduct strong, when unrolled about 25.0 mm.
long ; the succeeding part of the length of 4-5.0 mm., thinner ; the
rest about 7.0 mm. in length, stronger, nearly as in the first part. In
the beginning of this last part the true spermatic duct was rolled up in
tight coils, the remaining part of its length was nearly straight. The
penis about 1.5 mm. long, with the usual glans in the interior. The
spermatotheca (fig. 6a) spherical, its chief duct nearly twice as long as
the bag, the vagina short (fig. 6e). The spermatocysta appeared
pyriform (fig. Qd)
In color this form seems to differ from the typical one, as that is
represented by Alder and Hancock (Monogr., Part vi, 1854, fam. 3,
li. 9); in the anatomical relations no specific differences could be
detected.
A specimen of another variety was obtained by Dall, on a gravel
beach, at low water, in June, 1874, at Port Etches (Prince William
Sound . According to Dall, the mantle was of "brown " color.
The specimen had a length of 13.0 mm,, by a breadth of 8.0 mm.,
and a height of 5.0 mm. ; the height of the leaves of the gill was
about 1.0 mm. The color of the back was brownish and yellowish;
that of the gill, as well as of the rhinophoria, yellowish. The number
of leaves of the gill was about thirty.
The bulbus pharyngeus about 1.75 mm. long, by a height of 1.5 mm. ;
the sheath of the radula nearly as long as the bulbus ; the buccal crop
^ In one specimen the form of this organ was entirely as figured iu my
Malacolog. Untersuch. (Semper, Reisen). Tab. LXV, fig. 2.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 65
a little larger than the bulbus. The radula brownish-yellow, with
nine rows of teeth, further back fifteen developed and two younger
rows, the total number being twenty-six. The teeth quite as above,
dark, horn-colored in their thicker parts ; the median ones reaching a
height of 0.16 mm. The salivary glands as above-mentioned.
The biliary sac uncommonly small. The black contents of the
rectum consisting of undeterminable animal matter, mixed with larger
and smaller pieces of small Crustacea. The liver much flattened on
the riglit anterior half.
The anterior genital mass large, about 7.0 mm. long, 5.0 mm. high,
and 3.0 mm. thick. The ampulla of the hermaphroditic duct whitish,
forming a long ansa, about 5.0 mm. long. The spermatoduct shorter
than in the other form, otherwise, with the penis, as in that form.
The spermatotheca yellowish, short, sac-shaped, of a largest diameter
of 3.0 mm. ; the spermatocysts about 0.3 mm. long, pyriform. The
mucous gland chalk-white and brownish-gray.
Of another variety, Dall, in August, 1872, obtained six specimens,
in Sanborn Harbor (Shumagin Ids.), on stony bottom, at low water.
According to Dall, the color of the back of the living animal is
" red-brown, with whitish papilla3." The color of the backs of
the specimens preserved in spirits was rather uniformly, dirty brown-
yellowish, commonly much lighter on the middle, the papilliB whitish ;
the gill and the rhinophoria of the color of the back ; the under side
of the whole body yellowish ; more whitish on the mantle. The
length of the animals varied from 18.0 to 25.0 mm., by a breadth of
11.0 to 16.0 mm., and a height of 8.0 to 12 0 mm. ; the breadth of
the foot '1.5 to 12.0 mm. ; the height of the rhinophoria reaching 3.0
mm., that of the gill 2.0 mm. The form as usual. The horseshoe
shape of the gill very pronounced, the number of leaves, twenty-eight
to thirty. The gill was surrounded by higher papillae, which, in the
largest specimen, reached the height of about 2.5 mm. ; the space
inclosed by the gill closely set with similar papillas, the largest (as
large as the above mentioned) in the periphery. The gill can be so
deeply drawn back in its groove, that these external and internal
papillai shut over and quite conceal it ; the papilla? of the centre
smaller ; a crest or some few papilhe in the median line go from the
anus backwards, between the incurved ends of the gill. The anus
small, very slightly prominent ; the renal pore on the right side. The
openings of the rhinophor-holes as usual, before them the two usual
papilUc, behind them a bare space. The papilla? of the back quite as
66 PROCEEDINGS OF THE ACADEMY OF [1880.
in the previously examined form, the largest (in the largest specimen)
reaching the height and the diameter of about 1.5 mm., those in the
neighborhood of the gill somewhat larger.
Two smaller individuals were dissected, the larger being harder than
these and not so suitable for that purpose. The peritoneum w^as
colorless.
The central nervous system just as in the former specimens, but the
buccal ganglia smaller than the olfactoiy, and the gastro-oesophageal
short-stalked.
The eyes as above. The otocysts, under the glass, very distinct as
chalk-white points on the hinder and outermost part of the cerebral
ganglia. The leaves of the rhinophoria without spicula. The skin
and the papillae of the back as above or still more crowded with very
hard spicula.
The oral tube large, (in both individuals) about 2.5 mm. long. The
bulbus pliaryngeus of the usual form, (in both individuals) about 3.0
long, by a breadth of 1.8 mm , and the height nearly the same; the
sheath of the radula projecting straight backwards 2.0 mm. The
buccal crop, lying to the left side of the bulbus, somewhat compressed,
of about 3.0 mm. largest diameter, the stalk nearly half as long as
the largest diameter of the crop. The tongue with ten rows of teeth,
further backwards also eleven or twelve developed and three younger
rows, the total number thus being twenty-four or twenty-five. They
were entirely as in the form first examined.
The salivary glands, the pyloric part of the intestine, with its biliary
sac, and the liver as usual. The sanguineous gland whitish, much
flattened, covering the whole upper side of the bulbus pharyngeus and
the central nervous system ; a flattened cavity in its interior. The
hermaphroditic gland, through its more reddish color, contrasting with
the grayish color of the liver.
The anterior genital mass 11.0 to 12.0 mm. long, by a height reach-
ing V.O to 8.0 mm., and a breadth of 4.0 to 4.5 mm. The ampulla of
the hermaphroditic duct lying transversely on the lowest and most
anterior part of the back of the mucous gland, rather straight or
forming nearly a circle, about 5.0 to T.O mm. long, whitish. The
spermatoduct making many coils on and before the anterior part of
the mucous gland; the first part about 35.0 to 45.0 mm. long, the
second nearly 25.0 mm. long ; the penis about 1.5 to 2.0 mm., pro-
jecting freely from the vestibulum, conical ; the glans seemed rather
short. The spermatotheca of about 3.0 mm. diameter, whitish. The
1880.] NATURAL SCIENCK8 OF PHILADELPHIA. 61
spermatocysta (fig. 66) quite imbedded in and concealed by the mucous
gland, only a part of its chief duct free on the surface of this last ;
the spermatocysta scarcely shorter than the spermatotheca, pear-sluiped,
incurved ; the duct to the mucous gland (fig. Qd) passing from the end
of the bag, the other strong, longer (tig. (^c^, opening in the duct of
the spermatotheca, where it begins to be wider (vagina) ; the vagina
(fig. Ge) rather wide, but short. The mucous gland whitish, yellowish
and dirty yellow. ^
2. L. varians, Bgh. PI. XI, fig. 1.3, 14; PI. XIII, fig. 1.
L. varians, B. R. Bergh, Malacol. Unters. 1. c, 1878, p. 613, 014.
Color cu?rulescens vel albescens vel flavescens.
Dentes laterales margine interno denticulati fere usque ad apicem,
Hab. Oc. Pacif. (Ins. Kyska).
Of this species six specimens were taken by Dall, in July, 1873, at
Kyska Island, on sandy ground, at a depth of 9-14 fathoms. Four
specimens were sacrificed to the anatomical examination.
According to Dall the color of the living animal is " bluish." The
animals preserved in spirits were of a uniform whitish color, so too
the rhinophoria and the branchia. Their length was 9-12.0 mm. by
a breadth of 5.3-7.0 and a height of 3-4.5 mm. ; the breadth of the
foremost part of the foot 3.6-5.0 mm. The height of the rhinophoria
reached about 2.2 mm., of the branchial leaves 1.0 mm.
T he form almost entirely as in the typical form and as in the L.
hystricina. 'i he head as in the last species ; also the openings of the
rhinophor-holes, with their (mostly three) larger tubercles, set with
equal spaces ; the club of the rhinophoria Vf-ith about twelve to fifteen
rather thick leaves. The tubercles of the back as in the L. hystri-
cina ; the number of larger ones much exceeding that of the smaller,
which are scattered between them. The branchial disk as in the L.
hystricina, also the branchial leaves, whose number did not surpass
twelve to twenty. The foot as usual.
The central nervous system (fig. 1) nearly as in the L. hystricina.
The cerebro-visceral ganglia of roundish or oval form, as also the
pedal ones which were not much smaller than the former. The com-
1 In my "Malacolog. Unters." (Semper, Philipp. II, ii, Ileft xiv, 1878,
p. 60G-G13 ; Tab. Ixiv, tig. 13, 14-1'> ; Tab. Ixv, fig. 1-.5, G-13) I have given
some anatomical remarks on the typical L. bilaviellata and ou the Green-
landic variety {D. liturata. Beck).
68 PROCEEDINGS OF THE ACADEMY OF [1880.
missura pedalia nearly as long as the diameter of the pedal ganglia ;
the subcerebral lying rather close up to the pedal ; the visceral quite
free, much thinner. A very short-stalked smaller ganglion (fig. Ic)
connected with the under side of the right visceral ganglion, gives off
a nerve that swells into a new ganglion, which sends out three nerves
(N. genitalis). The olfactory ganglia short-stalked, spindle-shaped.
The buccal (fig. Id and the gastro-oesophageal ganglia (fig. le), nearly
as in the L. hystricina ; the commissure between the first extremely
short, the gastro-oesophageal somewhat smaller.
The nervi optici one to one and a-half times as long as the diameter
of the cerebral ganglia ; the eyes with black pigment, yellowish lens.
The otocysts (fig. 1) lying rather backwards, a little smaller than the
eyes ; the otokonia of the usual form, in number about fifty. The
leaves of the rhinophoria without spicula. In the skin were almost
no spicula and no larger or calcified ones on the surface of the rigid
papillag of the back, which thus were rather smooth. In the intersti-
tial connective tissue small calcified cells, but no larger spicula.
The mouth-tube as in the L. hystricina. The bulbus pharyngeus
as in that species, but the sheath of the radula shorter and less
prominent, bent upwards, sideways or down and forwards. On the
intei'ior part of the nearly colorless labial disk, the usual belt of
(about twelve to fifteen) rows of small denticles. The tongue strong,
rather long, with curved superior and nearly straight inferior margin.
In the mature radula twelve to fourteen or sixteen rows of teeth,
further backwards fifteen or sixteen to eighteen rows of developed,
and three of partly developed teeth ; the total number of rows thus
thirty, thirty-one or thirty-five to thirty-seven. The median plates
(fig. 14) of nearly the usual form, in the under side rather excavated,
with thickened margins. The large lateral plates (fig. 13) formed
nearly as in the L. hystricina, but larger, reaching a height of 0.12
mm. ; the denticulation of the interior margin of the hook stronger,
with more (about twenty) denticles and reaching farther out towards
the end of the hook. The exterior plates nearly of the same form as
in the last species, reaching to the height of about 0 6 mm.
The sucking-crop quite as in the former species.
The salivary glands much smaller than in the former species, re-
duced to a large, scarcely lobed, whitish mass on each side of the
root of the ojsophagus.
The oesophagus somewhat spindle-shaped. The stomach included
in the liver. The intestine issuing from the liver behind its middle-
1880.] NATURAL SCIENCES Or PHILADELPHIA. 69
The liver of grayish-white color, of the length of about 9.5 mm. by
a breadth of 4 and a height of about 3.75 mm. ; the hinder end
rounded, the fore-end rather truncated, the anterior one-third on the
upper and right side flattened by the anterior genital mass.
The heart and the renal syrinx as usual ; the median renal cham-
ber continued to the fore-end of the liver. The sanguineous glands
connected on the upper side of the central nervous system to a flat-
tened whitish mass.
The glandula hermaphrodisiaca clothing the upper side of the liver,
and scarcely distinct from it in color ; in its lobules were large oijgene
cells. The anterior genital mass compressed, plano-convex; 4.0 mm.
long, by a height of about 3.3 and a breadth of 1.2 mm. The albumi-
nous gland on the left side of the mass and forwards, yellowish, very
finely gyrated on the surface ; the mucous gland whitish, pellucid.
The spermatoduct as well as the (3.0 mm. long) penis as in the L.
echinata. The spermatotheca rather small, spherical.
L. varians, var.
To this same species belonged certainly five specimens of a Lam-
ellidoris, which were taken by Dall in July, 18'73, at Unalashka
Island (Aleutians), at the depth of sixty fathoms on mud and stones.
Nevertheless, the color of these animals in the living state was, accord-
ing to Dall, " yellowish-white."
The size and the particular measures accorded with those of the
more typical individuals, referred to above.
The central nervous system as just mentioned, so even the eyes and
the otocysts. The bulbus pharyngeus of the usual form ; on the tongue
eleven rows of teeth, farther backwards twenty-six developed and four
not quite developed rows, the total number thus forty-one. The plates
quite as formerly described. The sucking-crop quite as in the typical
form, also the salivary glands. The whitish sanguineous gland entirely
covering the central nervous system. The penis as usual.
Two specimens of another variety of this form were gotten by Dall,
In July, 1873, at Kyska Island, on sandy bottom, and at a depth of
nine to fourteen fathoms. In a living state they were, according to
Dall, of yellowish color.
The length of the animals preserved in spirits was 8.5 to 9.0 mm.,
by a breadth of 6.0 ram., and a height of about 3.5 mm. The color
was uniformly whitish or yellowish-white. One individual was dis-
sected.
TO PROCEEDINGS OP THE ACADEMY OF [1880.
The central nervous system was as above mentioned, and also the
eyes (their nervi optici rather long), and the otocysts (the number of
the otokonia about one hundred). The bulbus pharyngeus as usual ;
on the tongue sixteen rows of teeth, farther backwards eighteen rows
of developed and four of younger teeth ; the total number of rows,
thirty-eight. The plates as above ; the length of the median plates
005 to 0 058 mm. ; the height of the anterior large lateral plates
about 0.14 mm., of the posterior about 0.17 mm. ; the number of den-
ticles on these plates mostly fifteen to twenty. The vesica fellea wa?
at the left side of the pylorus.
S. L. hystricina, Bergh.
L. hystricina, Bcrgh, Mai. Untersuch., 1. c, 1878, p. 614, Tab. Ixviii.
fig. 17-23.
Color cocrulescens.
Denies laterales margine interno denticulati sed non usque ad
apicem.
Habitat. Oceanum Pacificum (insula Kyska).
One specimen of this species was found by Dall, at Kyska Island
(Aleutians), on rocky bottom, at a depth of ten fathoms, in June.
1873. According to Dall, the color of the living animal is bluish.
The specimen preserved in spirits was 9.5 mm. in length, reached
a breadth of G 0 mm., and a height of the true body (without the
papilUe) of .3.5 mm. ; the breadth of the foremost part of the foot
was 5.3 mm., the height of the rhinophoria was about 2.1 mm., of the
branchia about 1.2 mm., of the dorsal papilla^ 1.2 mm. The color
was uniformly whitish.
The form was oval, the back not very convex. The head rather
large, formed like a velum, that is radiately folded, and has its side
parts connected with the ends of the anterior margin of the foot ; in
the middle of the hinder part of the under side of the velum is a trans-
verse slit, in which the small mouth-pore opens. The opening of the
rhinophor-holes was nearly round, with the margin rather thin, here
were three papilla? of the same kind as on the back ; the rhinophoria
stout, the club with about twenty leaves. The back covered all over
with mostly stout, club-shaped papilhv, apparently set without order,
and extending nearly out to the very margin of the mantle, which is
thin and has on the upper side smaller, cylindrical or club-shaped
papilhe. The papilht all firmly adherent to the skin, the spicules shin-
ing through all over on the back and in the papillae. The branchial
1880.] NATURAL SCIENCES OF PHILADELPHIA. Tl
disk rather large, at the margin set with about fourteen papillae, irregu-
larly alternating in size. The branchia composed of twelve small
leaves of the usual kind. The centre of the disk and the anus as
usual. The foot somewhat shorter and narrower than the back,
broader in front, with the anterior margin rather straight, rounded
posteriorly.
The cerebro-visceral ganglia showed the visceral part a little larger
than the cerebral, the pedal somewhat smaller than the visceral; the
four commissures as usual; the offshoot of the nerva genitalis could
not be determined. The buccal ganglia rounded, connected through a
short commissure ; the gastro-oesophageal having about one-quarter of
the size of the latter.
The eyes with very rich black pigment ; the nervus opticus not short.
The otocysts as large as the eyes, filled with otokonia of the usual
kind. In the thin leaves of the rhinophoria no spicula. In the skin
of the back and in the dorsal papillas an enormous amount of irregular
or rounded particles, often coalescing together in larger, irregular
lumps, which very often were crowded together in irregular heaps ; in
the papillse also were long, strong and very much calcified spicula,
often of uneven surface, whose points, as usual, often projected on the
surface of the papillie. In the interstitial connective tissue, including
the ends of the different ducts of the genital organs (vagina, mucous
gland duct), masses of large and long (as much as 0.9 mm,), calcified
spicula.
The mouth-tube was about 1.0 mm. long, rather wide, with strong,
longitudinal folds. The bulbus pharyngeus of usual, irregular form,
the bulbus proper of the length of about 1.75 mm. ; the sheath of the
radula, nearly as long as the bulbus, curved downwards. The labial
disk oval, at the inner margin of darker color, and there showing
(fig. 17) a narrow belt of small, yellowish denticles, of a height of
0.007 to 0.015 mm. ;' this belt seems continued a short space up in
the mouth that is otherwise, like the rest of the buccal cavity, clothed
with a rather thick, yellowish cuticula. Ihe tongue rather long and
narrow, in the groove on its back sixteen rows of teeth, in the sheath
eighteen developed and six undeveloped rows, the total number conse-
quently forty. The color of the true lateral teeth yellowish, the others
nearly colorless ; the height of the outer pseudo-plates about 0.075 mm.
1 he median pseudo-plates elongate, narrow (fig. 21) ; the true (lateral)
' In the outer mouth was found a little Caprella, of the length of 3.0 mm.
72 PROCEEDINGS OF THE ACADEMY OF [1880.
teeth strong, finely denticulated (with six to eight denticles) on the
inner side of the hook, and with a strong, rounded prominence at
the base of this (fig. 18a, 19, 20) ; the external pseudo-plates with
the usual curved points (fig. 186). Irregularities in the form of the
last were often observed (fig. 23).^
The crop entirely as in the typical species, the largest diameter
1.3 mm. /
In the stomach indeterminable animal matter and a little, undeter-
minable worm, of the length of 2.0 mm.
The hermaphroditic gland as usual ; the lobules filled with sperma.
The anterior genital mass rather large, measuring in length 4.5 mm.,
in height 2.5 mm., and in breadth 2.3 mm. ; the left side flat or a little
excavated, the right rather convex. The mucous gland, as well as
the albuminous gland, white and yellowish-white. The spermatoduct
not very long, but rather strong, continued in the very strong penis,
that (retracted ) forms the fore-end of the whole mass. The penis has
a length of about 3.5 mm., by a diameter of 1.3 mm. ; the inferior
end rather constricted ; the superior three-quarters of the organ com-
pact, perforated through the axis by the dense coils of the spermato-
duct proper ; the inferior one-third hollow, including the curved and
pointed glans.
2. L. muricata (MilUer). Plate IX. fig. 18; Plate XI, fig. lO-Ii.
Doris muricata, O. F. Muller. Zool. Dan. Fas. Ill, 1789, p. 7, Tab.
LXXXV, f. 2, 3, 4.
Doris muricata, Muller. Sars, (forma /5) Loveu, Ind. Moll. Scand.1846,
p. 5.
Dori<i muricata, Meyer und Moebius. Fauna der Kieler Buclit, I, 1865,
p. 73-75, Taf. Vc, fig. 1-8.
f Lamellidoris muricata, Muller. G. 0. Sars, Moll. reg. arct. Norv.,
1878, p. 307, Tab. Xlll, fig. 6.
Color flavidus vel luteo-albus.
Dentes laterales magni hamo denticu'ato sed non usque ad apicem.
Hah. Oc. Atlanticum septentr.
The original specimen on which Miiller founded his Doris muricata
does not exist, and by his incomplete description it is now completely
impossible with full certainty to determine what species was meant by
his description. In future the species described by Meyer and Moebius
' From the presence of only one Individual, the examination of the radula
was extremely difficult and limited, as also that of the genital organs.
1S80.J NATURAL SCIENCES OF PHILADELPHIA. 73
and by me ought to be called by that name. To the same is without
doubt to be referred the second variety (/9) of the D. muricata
(Miiller, Sars) of Lov^n (the first being the D. Lov'ni of Alder).
Of this form, and under that name, I have had two well conserved
specimens for examination, kindly sent me by Mr. Friele, of Bergen,
and caught in the neighborhood of that place.
The individuals (preserved in spirits) were of light yellowish color.'
The length 9-10 mm. by a breadth of 5-6.0 and a height of nearly
8.0 mm. : the breadth of the foot reaching 3.5 ram. ; the height of the
rhinophoria 1.5, of the branchial leaves 1.0 mm. The form of the
animal as usual ; the warts of the back not large, mostly truncate,
clavate. The openings for the rhinophoria as usual, with two tubercles
before them, or one on each side; the club with about fifteen to twenty
leaves.- The branchial leaves about twelve to fourteen, as far as could
be determined ;3 the space inclosed by the gill covered with the usual
tubercles ; the anus presenting the ordinary features. The head rather
large, the side parts adhering to the foot throughout their whole length.
The genital groove with three openings j a foremost round, a median
spalt-formed, and a posterior large and round.
Both individuals were dissected ; the peritoneum was colorless.
In the central nervous system the cerebro-visceral ganglia appeared
rather short, reniform ; the pedal ones of roundish form, somewhat
larger than either of the former ; the commissures rather short. The
olfactory ganglion short-stalked, nearly spherical, situated rather
posteriorly on the upper side of the cerebral ganglia, and nearly as
large as the buccal ones. The buccal ganglia of oval outline, con-
nected by a short commissure ; the gastro-oesophageal nearly spherical,
in size about one-quarter of the former,, short-stalked: a secondary
ganglion lying above the last on the oesophagus.
The eyes not short-stalked ; with rich black pigment and yellow
lens. The otocysts a little smaller than the eyes, filled with otokonia
of the common kind. In the leaves of the rhinophoria rather few but
large spicula of the same kind as in the skin, more or less perpendicu-
lar on the free margin ; the axes of the club like the stalk still more
richly endowed with smaller and larger spicules. Under the glass the
' According to Loven the color is yellowish; to Meyer and Moebius white
or yellowish-white the rhinophoria orange-colored.
- According to Meyer and Moebius the club of the rhinophoria has but
nine or ten leaves.
' Meyer and Moebius mention eight leaves as nearly constant.
6
74 PROCEEDINGS OF THE ACADEMY OF [1880.
skin between the warts, as well as the warts themselves, showed the
white spicules everywhere shining through ; the spicules often project-
ing from the surface of the warts The spicules for the greater part
very large, long, and reaching a diameter of at least 0.05 mm. ; they
were strongly calcified, mostly straight or slightly curved, the surface
nearly even. In the interstitial tissue were rather many spicules, but
(as in the rhinophoria) less calcified than in the skin.
The mouth-tube rather wide. The bulbus pharyngeus of nearly
usual form, about 1.6 mm. long ; the sheath of the radula, moreover,
projecting backwards about 0.4 mm., bent somewhat upwards or down-
wards ; the lip-disk with a rather thick yellowish cuticula ; the sucking-
crop large, larger than the true bulbus, to which it adheres by a very
short petiolus. The tongue with nine rows of teeth, further back
twenty to thirty-two developed and three younger rows ; the total
number of rows, thirty-two to forty-four.^ The yellow median plates
(fig. 10a) about 0.05 mm. long, of the usual form. The large lateral
plates yellow, of about 0.12 mm. height ; the form as usual ; the hook
with about fifteen to sixteen fine denticles, and a strong tooth at the
inside of the base (fig. lObb). The external plate colorless, about 0.04
mm. in height, with the usual rudiment of a hook (fig. 10c, llb).^
The salivary glands white, rather thick, making two or three short
coils at the sides of the oesophagus. The oesophagus as usual. The
intestine emerging from the liver at about the middle of its length ;
the biliary sac (fig. 18) is at the pyloric part of it, situated deeply,
scarcely showing itself on the surface of the liver and opening (fig.
18a) into the stomach close to the pylorus. The liver about 6.5 mm.
long by a breadth of 3.0 mm. and a height of 2.0 mm., deeply excav-
ated in the anterior third of its right side, and of light yellow color.
The sanguineous gland much flattened, whitish, heart-formed, of about
1.5 mm. largest diameter. The renal chamber rather wide, the tube
on its floor strong.
' Meyer and Moebius (1. c. p. 73) mention twenty-nine rows ; Alder and
Hancock thirty.
- The representations of the external plate by Meyer and Moebius (1. e.
fig. 2, 6) are not natural. Alder and Hanc. (1. c, Part VII, p. ii, PI. 46,
supplem. text) mention two external plates in their D. muricata (as in their
D. diaphana) ; either the D. muricata of A. and H. must be another species,
or they must have fallen into error from the particular view which is some-
times had in certain positions of the hind ends of the large lateral teeth
with the external ones. " .
1880. J NATURAL SCIENCES OF PHILADELPHIA. 75
The lobes of the hermaphroditic gland without developed sexual
elements. The anterior genital mass about 2.5-3.0 mm. in length by
a height of 2.0 mm. and a breadth of 1.0-1.5 mm. The ampulla of
the hermaphroditic duct of yellowish color, rather thick ( — 0.75 mm.
diameter), making a wide curve, about 2.5 mm. long. The spermato-
duct long ; its first part thinner, about 9.0 mm. long, then through a
stricture of the length of nearly 1 mm., passing into the thicker part,
which in its last half increases in thickness, and, all in all, has the
length of about 6.0 mm. by a diameter of 0.75 mm.; the last part
(fig. 12c) passes into the penis', in whose cavity (fig. 12bb) the glans
(fig. 12«) projects as a short club, the proper seminal duct passing
down to the gland in nearly continual cork-screw windings, and often
shining through the walls of the external coat. The spermatotheca
whitish, nearly spherical, of about 1.3 mm. diameter, filled with sem-
inal matter and detritus ; the spermatocysta elongate, nearly twice as
long as the former, yellowish, deeply imbedded in the mucous gland,
filled with ripe semen ; its duct somewhat longer than the cysta. The
vagina short.^ The mucous gland yellowish and yellow.
The species approaches to the L. hystricina and L. varians (of
the Pacific), but differs entirely in its colors; still the possibility can-
not be denied that further investigations may show both the Pacific
" species " to be merely varieties of the old Lamellidoris muricata
of the Atlantic.
ADALARIA, Bergb.
Adalaria, R Bergh. Malacolog. Unters. (Semper, Philipp. II, ii). Heft
XIV, 1878, p. xl.
Adalaria, R. Bergh. Gattungen nord. Doriden, 1. c. 1879, p. 360.
Forma corporis fere ut in Lamellidoridibus. Notha^um papillula-
tum vel subgranulosum. Branchia (non retractilis) e foliis vix mul-
tis, in formam ferri equini ut plurimum dispositis formata. Caput ut
:n Lamellidoridibus, latum, semilunare, teutaculis vix uUis vel brevis-
simis lobiformibus. Aperturse rhinophoriales integral, tuberculis
anticis 2-3, calvitie postica.
Discus labialis non armatus. Lingua rhachide lamellis depressis
instructa ; pleuris dente laterali interno hamiformi majore et serie
' The exserted penis is figured by Meyer and Moebius (1. c. taf. fig. 4)
and mentioned as cylindrico-conical.
- The upper end of the vagina seemed to present a particular diverticle.
T6 PROCEEDINGS OP THE ACADEMY OF [1880.
dentium externorum sat applanatorum pneditis. Ingluvies buccalis
bulbo pharyngeo petiolo connata.
Penis glande parva inermi. Vagina brevis.
«
The genus has been established by the author (1878) to receive
the D. proxima and its allies. The Adalarice externally approach
nearest to the Lamellidorides ; their branchial leaves are also dis-
posed mostly in horseshoe form, but fewer in number. The head and
the' tentacles are more as in the Aeanthodorides. The back is nearly
as in the Lamellidorides, but the granules are sometimes more pointed.
The opening for the rhinophoria as in the Lamellidorides, with plain
margin ; before them two to three tubercles, behind them the glabella.
The lip-disk only covered by a strong cuticula. The armature of the
tongue approaching to that of the Acanthodorides. The rhachis of the
tongue carries depressed small yellow plates ; at each side of these a
large hook-formed yellow plate, and further outwards a series of
smaller, nearly colorless plates, of which the inner ones are more
compressed, the rest depressed. The sucking-crop as in the Lamelli-
dorides, through a petiolus fixed to the bulbus. The salivary glands
as in the Lamellidorides. The oesophagus wider at its root. The
penis unarmed ; the vagina short.
The Adalarice are Lamellidorides with a tongue resembling that of
the Acanthodorides ; they form a sort of connecting link between these
two groups.
Of the typical species, the spawn is known (through Alder and
Hancock) and some few notices have been published about their
biology (through Meyer and Moebius) ; Sars mentions' the swimming
of Ad. Loveni.
The genus seems to belong to the northern oceans ; only five species
.seem hitherto known.
1. Ad. proxima (A. et H.). Oc. Atlanticus sept.
'Z. Ad. pacifica, Bgh., n. sp. Oc. Pacif.
;J. Ad. virescens, Bgh., n. sp. Oc. Pacif.
4. Ad. albopapi'losa (Dall). Oc. Pacif.
5. Ad. Loveni (A. et H.). Oc. Atlant. sept.
1. Adalaria proxima (Alder et Hancock;. PI. IX, fig. 12-15.
Doris proxima, A. et H. Monogr. Part VI, 1854. Fam, 1, PI . 9, fig.«».
10-1 G ; Part VII, 1855. PI. 46, suppl. f. 8.
Doris proxima, Meyer u. Moebius, Fauna der Kieler Bucht, I, 1865.
P. 69-71 ; taf. V b, fig. 1-8.
' Sars, Bidr. til Soedyrenes. Naturhist. 1829, p, 17.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 7"
Color flavus vel e rubro flavus.
Dentes laterales (magni) hamo edentulo ; externi numero 10.
Hab. Oc. Atlant. septentr.
Of this form I have had for examination three specimens of nearly
equal size, kindly sent me by Prof. Moebius in Kiel, and caught in
the neighborhood of that town.
The individuals wpre of a uniform whitish color, the liver shining
reddish-gray through the foot. Alder and Hancock have already re-
marked this shining through of the liver. The length was 7-0-8.0 mm.,
by a breadth of 5.0-5.5, and a height of about 3.5 mm- ; the height of
the rhinophoria about 1.25, of the branchial leaves 0.75 mm. The
form nearly as in the Ad. pacijica, also the tubercles (fig. 12) of the
back and the surroundings of the rhinophor-holes ; the branchial
leaves nine to ten in number. The number of branchial leaves ac-
cording to Alder and Hancock is eleven, according to Meyer and
Moebius eight or nine. The rhinophoria with about fifteen to twenty
leaves. The lateral parts of the head nearly connate with the foot.
and only slight traces of true pointed tentacles. The foot as in the
next species.
The three individuals were anatomically examined. The peritoneum
colorless.
The central nervous system as in the Ad. pacifica, but less de-
pressed. The eyes and otocysts as in that species ; the last with
about 200 otokonia of very varying diameter, reaching about 0.02
mm. The spicula of the skin as described by English and German
authors ; a rather large quantity spread in the skin of the head.
The bulbus pharyngeus (with the crop) of the length of about 1.5
mm., by a height of 1.5 and a breadth of 0.8 mm. ; the crop making
about half of the bulbus; the lip-disk with strong yellowish cuti-
cula ; the sheath of the radula a little prominent, bent more or less
upwards. The tongue narrow and pointed, with seven to nine rows
of teeth, further backwards thirty or thirty-one rows of developed and
three of younger teeth ; the total number thus amounts to forty or
forty-three.'
The teeth as in the Ad. pacifica. The large lateral yellowish, the
rest nearly colorless. The length of the median teeth about 0.025 to
0.03 mm. '1 he large lateral (fig, 1366, 14) showed the prominence
' Alder and Hancock notice forty-one, Meyer and Moebius tbirty-nine
rows of plates.
18 PROCEEDINGS OP THE ACADEMY OP [1880.
ut the inside of the root of the hook quite as in the Ad. pacifica.
The external teeth (fig. 15) only nine or ten in number,^ fewer than
in that species, always absent on more than half the tongue.
The salivary glands as in the next species, also the oesophagus, the
stomach and the intestine. The liver also of nearly the same form,
the inferior part of the posterior end continued as a little cone ; the
surface (especially of the back part) yellowish-white ; the substance
yellow. The vesica fellea in its usual place, small. The heart as
usual, also the sanguineous gland. The renal syrinx and the urinary
chamber as usual.
The anterior genital mass rather compressed, of angular-roundish
outline, of about 1.15 mm. largest diameter. The spermatoduct seemed
shorter than in the next species, especially the second part ; the penis
short. The spermatotheca pyriform ; the spermatocysta of more oval
form, having only about one-quarter of the size of the former, and fiUed
with sperma. The mucous gland whitish and yellowish.
2. Adalaria paciflca, Bergh, n. sp., PI. IX, fig. 17 j PI. X, fig. 1-3; PI. XI, fig. 1&.
Color lutescens.
Denies laterales (magni) hamo edentulo ; externi numero 15.
Habitat. Oceanum Pacificum (Unalashka).
Of this species Dall caught three specimens, in September, 1874, at
Unalashka, on a bottom of mud and shells.
According to Dall, the color of the living animal is " yellowish ; '*
the specimens preserved in spirits were of a uniform yellowish color.
The length of the two larger specimens about 12.0 to 14.0 mm., by a
breadth of 8.0 to 9.0 mm , and a height reaching 4.5 to 5.0 mm. -,
the breadth of the foot 6.0 mm., the height of the rhinophoria about
1.5 mm., of the branchial leaves 1.2 mm.
The form as in the Ad. proxivia, a little broader anteriorly. The
back covered all over with a mass of rather stout, subglobose and sub-
petiolate tubercles quite as in the typical species, mixed with much
fewer smaller ones. The larger ones, under magnification, showing
the perpendicular spicula shining through, while other spicula were
detected irregularly scattered in the intervals between the tubercles.
The rhinophor-holes nearly without projecting margin ; the adjoining
part of the back, behind, smooth ; immediately before the holes, on
* The number of external plates is, according to Alder and Hancock,
ten, to Meyer and Moebius, eight or nine.
1880.] NATURAL SCIENCES OF PHILADELPUI A. 19
the contrary, two or three larger tubercles ; the club of the rhinophoria
with about thirty leaves. The branchial area surrounded by larger
tubercles. The branchial leaves in number, eleven or twelve ; imme-
diately before the two hindermost was the slightly prominent anus, and
at its right Bide the renal pore ; in the space between the anus and
the branchial leaves, three or four larger and two or three smaller
tubercles. The head large ; the tentacles short, pointed. The foot
broad, rounded behind, a little broader in front ; the furrow on the
anterior margin very indistinct. The three individuals were all dis-
sected. The peritoneum was colorless.
The central nervous system rather flattened ; the cerebral ganglia
larger than the visceral, which were lying at their outer margin and
were a little larger than the pedal ones ; the proximal olfactory ganglia
bulbiform, less large than the buccal ones, which were of short, oval
form, connected through a very short commissure ; the gastro-oesopha-
geal ganglia short-stalked, rounded, nearly half as large as the former,
with a very large cell. The subcerebral and the pedal commissures
conne^^ted, the visceral free.
The eyes with coal-black pigment, yellow lens ; the nervus opticus
in one individual with black pigment. The otocysts, under a mag-
nifier, very distinct as chalk-white points at the hinder margin of the
cerebral ganglia, nearly as large as the eyes, filled with ordinary oto-
konia. In the leaves of the rhinophoria scanty, scattered spicules,
perpendicular on the free margin, not much more calcified than in the
skin ; in the stalk of the organ the spicules larger and less scanty.
The skin, especially its tubercles, with many long spicules and calcified
cells and groups of such cells ; the form of the spicules different from
that of the Doris proxima, as figured by Alder and Hancock (Monogr.,
Part vi, fam. 1, PL 9, fig. 15), and by Meyer and Moebius (1. c, figs.
8, 9), much less calcified, more straight and of more uniform shape.
In the interstitial connective tissue of the chief ducts of the anterior
genital mass were scattered large spicules.
The mouth-tube wide, about 1.3 mm. long. The bulbus pharyngeus
of rather compressed form, about 2.0 mm. long; the sheath of the
radula strongly projecting from the hinder end, nearly as long as
the bulbus, more or less curved upwards ; the lip-disk oval, with a
very strong yellowish cuticula. The tongue with ten or eleven rows
of plates, further back twenty-nine to thirty-four rows of developed
and three of younger plates ; the total number thus forty-two, forty-
three, forty -seven. The median plate (PI. IX, fig. ITa ; PI- X, fig. 1)
80 PROCEEDINGS OF THE ACADEMY OF [1880.
yellowi'gh, of a length of about 0.045 mm., with a median furrow
along the upper side and with thickened margins. The large laterals
horn-yellow in color, reaching the height of about 0.1 mm. (PI. IX,
fig. lib; PL X, fig. 2aa), hook-shaped, with a strong, roimded prom-
inence at the inside of the root of the hook (fig. 17). On each side
(PI. X, fig. 2b, c) of the two large plates (in two individuals) con-
stantly fifteen smaller, nearly colorless plates of a length of about
0.06 mm. These plates were all somewhat depressed ; the five inner
ones smaller, somewhat compressed ( fig. 2, 3a, 15 ) ; the others (fig. 2, 3)
broader, with the upper edge broad and irregularly toothed ; the outer-
most (fig. 2c) a little smaller than the adjoining plates. The bases in
each of the^e (fifteen) plates large, forming nearly half, or at least
making more than a third of the size of the whole plate.' The crop
of the bulbus of the usual form, as large or a little larger than the bulbus
itself; with a very short stalk with strong longitudinal musculature,
its aperture opening immediately behind the lip-disk.
The salivary glands large, white, very elongate, in their foremost
part broader, and with several coils filling the space left between the
crop, the bulbus and the cesophagus.
The oesophagus long. The stomach small, enclosed in the liver ;
the intestine rather short, forming its knee behind the fore-end of the
liver. The large posterior visceral mass about 9.0 mm. long by a
breadth of 4.3 and a height of 3.5 mm. ; the posterior end somewhat
pointed, though rounded ; the fore-end broader, perpendicular, somewhat
flattened on the right side ; the color of the surface (hermaphroditic
gland) ash-gray, the interior (the liver) brown or black brown, or
quite yellow.
The heart as usual. The sanguineous gland irregularly reniform,
situated somewhat more towards the left side, rather thick, whitish,
covering the central nervous system and a large part of the bulbus
pharyngeus from above. The renal syrinx as usual.
The hermaphroditic gland without developed sexual elements. The
anterior genital mass proper rather small, compressed, of about 2.5 mm.
largest diameter, but the loop of the spermatoduct (and the penis)
nearly as large as the rest of the mass. The spermatoduct long,
in its first part white, rather strong ; nearly as long as the second in
which it passes through a stricture ; this last part is thicker, cylindrical,
elongated, about 5.0 n;m. long, passing without exact limits into the
' In both individuals the three to five foremost rows were without the
smaller plates, and the following two or three very incomplete in this respect.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 81
short penis. The spermatotheca pyriform, about 1.3 mm. long; the
ppermatocysta not having one-fourth of the size of the last ; botli
empty. The mucous gland whitish and yellow-whitish.
This seems even externally to differ somewhat from the typical form,
of which it nevertheless may prove to be but a variety. Neither Alder
and Hancock, nor Meyer and Moebius saw more than eight to (nine)
ten external plates on the tongue of Ad. proxima, while this Pacific
form always presented fifteen.
3. Adalaria viresoeas, Bgh., n. sp. Plate X, ti^. \, 5.
Color virescens.
Dentes laterales (magni) hamo edentulo ; externi numero 15.
Hab. Oc. Pacific, septentr. Unalashka.
Of this species Dall found four specimens at Unalashku, on gravel,
in a depth of nine to fifteen fathoms, in September, 1874.
According to Dall the color of the living animal was " greenish,"
and the animals preserved in spirits showed remains of the same color
as a uniform grayish green. The length of these was 11.5-12.0 mm.,
by a breadth of 8.0 mm. and a height of 5.0 mm, ; the height of the
rhinophoria about 2.0, of the branchial leaves about 1.0 mm.
The form, as well as the rhinophor-openings, were quite as usual ;
the club of the rhinophoria with about thirty-five leaves. The gill not
large, with nine to twelve leaves ; the space within the gill as usual,
also the arms and the renal pore. The back covered with granula-
tions and short clubs. The head, with the tentaculaj and the genital
opening as usual.
Three individuals were dissected ; the peritoneum was colorless.
The central nervous system showed the cerebral ganglia larger than
the visceral, which were lying on the outside of and behind the former,
very distinct from them ; the pedal ones being intermediate in size
between the cerebral and the visceral ganglia. On the exterior part of
each cerebral ganglion a little short-stalked ganglion (gang, opticum?)
was easily visible under a hand magnifier. The (proximal) olfac-
tory ganglia bulbiform, short-stalked, a little larger than the buccal
ganglia, which were short-oval, connected through a very short com-
missure ; the gastro-ojsophageal being about one-fourth to one-fifth of
the size of the former. In the neighborhood of the penis a little oval
ganglion (g. penis) having a largest diameter of about 0.25 mm.
(fig. 5), containing only rather small cells.
82 PROCEEDINGS 0¥ THE ACADEMY OF [1880.
The eyes with black pigment ; the otocysts with not very many and
not much calcified otokonia. No distal olfactory ganglion, as far as
could be seen ; no spicula in the leaves of the rhinophoria. The skin
as in other species ; the spicula projecting on the surface of the granu-
lations of the back.
The bulbus pharyngeus about 1-1.5 mm. in length ; the sheath ot
the radula projecting 0.75-1.0 mm., bent upwards; the sucking-crop a
little larger than the bulbus itself, short-stalked ; the lip-disk as usual.
The tongue compressed, rather prominent, with six, eight, and nine
rows of teeth, also further back twenty-four, thirty-two and thirty-three
developed and three younger rows ; the total number of rows thus
being thirty-five, forty-one, forty-five. The median plates, the large
lateral and the (fifteen) external ones scarcely different from those of
the last species.
The salivary glands rather strong, with two or three short coils fill-
ing the space at the sides of the oesophagus (fig. 4), white. The
(esophagus (fig. 4a) wide in its upper part, the rest narrow. The an-
teriorly proceeding part of the intestine 2.0 mm. long, the other retro-
ceding part <^.0 mm. long ; no biliary sac could be found either at the
pylorus or higher up. The liver about 9 0 mm. long by a breadth of
4.2 and a height of 4.0 mm. ; of brownish-gray color ; the anterior end
truncate, inclined downwards and backwards ; the anterior one-third of
the right side flattened for the anterior genital mass ; the posterior end
somewhat pointed, rounded at the point.
The sanguineous gland whitish, covering the anterior end of the
bulbus pharyngeus and the foremost part of the central nervous system
or this last and the hinder part of the bulbus.
The anterior genital mass about 3.5 mm. long by a breadth of 0.75
and a height of 1.5 mm., a very large part of it formed by the thick
part of the spermatoduct. The ampulla of the hermaphroditic duct
about 2.0 mm. long, rathf r thin, whitish. The spermatoduct long ;
the first part thinner, about 8.0 mm. long ; the rest making a large
curve, about 5.5 mm. long, about three times as thick as the first, with
a diameter of 0.6 mm. ; the spermatoduct proper making many coils in
its interior course downwards to the penis, which shows a little un-
armed glans in the bottom of its orifice ; in one individual the penis
was exserted as a conical prominence of the height of 1.0 mm. The
spermatotheca pyriform, about 1,0 mm. long, of grayish color; the
spermatocysta a little less large, spherical ; the vagina rather short.
The mucous gland rather small.
1880. J NATURAL SCIENCES OF PHILADELPHIA. 83
Even this species might perhaps be merely a variety of the former ;
still it is of a quite different color and the back much more coarsely
j^ranulated,
4 Adalaria albopapillosa (Dall). PL IX. H^. Ifi; PI. X, fig. 9-11.
Alderia (? ?) albopapillosa, Dall, Amer. Journ. of Conch., vii, 2, 1873,
p. 137.
Color pallide flavescens, papillis dorsalibus niveis.
Dentes laterales (magni) hamo basi denticulato.
Habitat. Oceanum Pacificum septentrion. (Sitka).
Of this curious animal Dall caught three specimens [in company
with the Doris (Archidoris) Montereyensis and the jEolidia {Her-
missenda) opalesceiis'], in July, 1865, on algte, at the depth of six
fathoms, at Sitka (Alaska).
According to the drawings of Dall, the color of the living animal is
very pale yellow,^ the back all over covered with chalk-white papillae ;
the length was 3, the breadth 2 lines. The three original specimens
preserved in spirits were of a length of 5.5 to 7.0 mm., of a gi-eatest
breadth of 4.0 to 4.5 mm., and a height of 2.75 mm. 1 he color was
uniformly translucent grayish and yellowish whitish. The form of the
animal was oval, the mantle a little larger than and hiding the rest of
the body. 'I he back convex, covered all over with a multitude of
cylindrical or fusiform, relatively rather large papilla?, reaching to the
height of a full millimetre, and with some few small ones spread
between them. The rhinophor-openings at their usual place, having,
as usual (with retracted organs *, thin margins ; before them always
two larger papillae, behind them a little naked space.^ The club of
the (yellowish) rhinophoria with about twenty-five leaves. The gill
rather small; the branchial leaves (yellowish), as usual, set in horse-
shoe form, lower or at least not higher than the dorsal papillte, in
number, ten to twelve ; the anal papilla rather low, with one of the
ordinary papilla? before and one behind it ; the space between the
' "Of an opaque white, the remainder of the animal except the eyes,
being translucent yellowish." — Dall.
- Dall did not detect the retracted rhinophoria ("tentacles none") ; the
"black eyes sessile on the anterior surface of the body, near the mantle
margin," did not exist in the figure, but in one individual two black sand-
particles were lying there. The true eyes of the animal could not be de-
tected through the skin, and were lying more backwards.
84 PROCEEDINGS OF THE ACADEMY OF [1880.
branchial leaves and the anus otherwise naked. ^ The genital open-
ing as usual. The foot rather large, with a very fine furrow in the
anterior margin. The head as usual ; the tentacles relatively rather
large
The three individuals were dissected. The peritoneum was color-
less.
The central nervous system quite as in the former species, the vis-
ceral ganglions lying outside of the cerebral ; no distal olfactory
ganglion could be detected ; the buccal ganglia connected through a
commissure at least as long as the diameter of the ganglion ; the
gastro-ocsophageal ganglia and the eyes as in the former species.
The otocysts could not be detected. In the leaves of the rhinophoria
the spicula much more scanty. In the skin the same kind of not
much calcified spicula as in the former species ; the papillae of the
back very richly endowed with such, and commonly with a mass of
them projecting with their points (PI. IX, fig. 16) on the surface of the
papilke.
The bulbus pharyngeus as in the former species ; the length about
1.5 mm., two-fifths of which is the straight, backwards projecting
sheath of the radula ; the cuticula of the lip-disk as usual ; the
buccal crop somewhat compressed, with rather long pedicel. The
tongue with nine or ten rows of plates, farther backwards sixteen or
seventeen developed and three younger rows ; the total number of
them, twenty-nine or thirty. The median plates (^fig. 9a, 10a) nearly
as in the former species, or a little shorter. The large lateral plates
(fig. 9b, 106) rising to the height of 0.12 mm., yellow ; their form as
in the former species, but at the inside of the hook at its root were
three to six or seven to eight small denticles. The external lateral
plates (fig. lOcd, 11) farther backwards, in number constantly eight;
the outermost (fig. 11a) very small, the others as in the former species.
The salivary glands, as far as could be determined, were as in the
last species ; so also the oesophagus and crop ; also the stomach and
the intestine, which seemed to have the usual bag (pancreas, biliary
sac) at the pyloric part. The sanguineous gland flattened, grayish,
cordate. The liver of brownish-gray color.
In the hermaphroditic gland no ripe elements were found, and the
anterior genital mass was very small
* According to Dall, the " anus is terminal under the edge of the mantle."
This was erroneous. He did not see the gill, but regarded the dorsal papillae
as "branchial appendages."
1880.] NATURAL SCIENCES OF PHILADELPHIA. 85
The species is easy to distinguish from the former, by its color and
especially by the denticulated hook of the large lateral plates.
5 Adalaria Loveni (Aldir et Hancock). PI. X, fig. 6-S.
Doris muricata? O. F. Miiller, Sars, Bidr. til Soedyreues Naturb.,
1829, p. 15. Tab. 11, fig. 7, 8.
Doris Loveni, Alder et Hanc. Ann. Mag. Nat. Hist., 3 Ser., X, 18(>2,
p. 262. *
Lamellidoris Loveni, Friele et Arm. Hansen, 1. c. p. 3.
Lameliidoris Loveni, G. O. Sars. Moll. reg. arct. Norv., 1878, p. 364.
Tab. XIV, fig. 1.
? Lamellidoris muricata (Mull.) Abildgaard. Morch, Faunula Moll.
Ins. Fiiroens. Naturb. Foren. Yidsk. Meddel., 1867, p. 75.'
Doris muricata, Miiller, Sars ('>), Loven, lud. Moll., 1846, p. 5.
Doris muricata, M. Sars. Reise i Lofoten og Fiumarken, 1851, p. 75.
Color dorsi et rhinophoriarum e brunneo lutescens, paginal inferioris
et branchiae lutescens.
Dentes laterales (magni) hamo edentulo ; externi (linguae)
numero 12.
Hob. Oc. Atlant. septentr.
This species was first noticed by Sars, who hesitatingly regarded it
as perhaps the Doris imiricata of Mueller. It is, moreover, the prin-
cipal form of the Doris muricata (" Mueller, Sars ") of Lovt'n (his
second variety being the true L. muricata) ; has been established
(1862) as a species by Alder and Hancock, and has as such been
adopted by Friele and Hansen, as well as by G. O. Sars, who lately
gave figures of the teeth on the tongue. The species has been much
confounded with the *' D. muricata," which is a Lamellidoris ; it is
certainly distinct from the Ad. proximn^ and seems also to differ from
the other described species.
Of this form I have had fifteen individuals for examination, kindly
sent me by Mr. Friele, of Bergen, and dredged in the neighborhood of
that place.
' According to Morch (Rink. Gronland, I, 1857. Tillaig 4, p. 78), the D.
muricata, Sars, should bo the D. liturata, Beck ; this last is a mere variety
of the Lamellidoris bilamellata, and with this should, on the other hand,
accoidiug to Morch (Faunula Mollusc. Isl. Naturh, Foren. Vidensk. Med-
del., 1868, p. 203), the D. proximo of Meyer and Moebius be synonymous,
which belongs to the quite diflferent genus, Adalaria. An example more —
if such were needed — of the way in which the Nudibranchiata have been
.synonymized and systematized.
86 PROCEEDINGS OF THE ACADEMY OF [1880.
The color of the animals preserved in spirits was uniformly yel-
lowish. The length was 13-15.0 mm., by a breadth of 8.5-9.5 and a
height of 4-5.0 mm.; the breadth of the foot 6 mm.; the height of
the rhinophoria about 2.5 mm., of the branchial leaves 1.0-1.3 mm. ;
according to M. Sars the height of the rhinophoria is four to five times
that of the tubercles of the back, (1. c. p. 16, also in one of his figures
fig> 7). The form as usual ; the back .covered all over with large
rounded tubercles, which rose to the height of 1.5 mm., and were of
a similar breadth ; they were sessile or more or less subpedunculate,
sometimes set in indistinct longitudinal rows ; between the larger
tubercles everywhere were smaller ones of different sizes ; on the
margin of the back were tubercles of middle size or smaller ; the
spicula rather indistinct between and in the tubercles. The rhinophor-
openings as usual, two large tubercles before them ; the club of the
organs with about twenty-five leaves. The gill with eight to twelve
leaves ; according to M. Sars, the number of branchial leaves is ten —
to LovL'n, eight to ten. A large (high) tubercle between the hindermost
leaves, before it the low anal papilla, and to the right side the renal
pore ; some few smaller papillae were spread over the space between
the anus and the branchial leaves The head large, broad ; the short
tentaculje pointed. The genital opening as usual.
Six individuals were dissected. The peritoneum was colorless.
The central nervous system rather flattened, especially the visceral
ganglia, which lay on the outer side of and behind the cerebral ones',
which were a little larger ; the pedal ones larger than either of the
other ganglia, situated perpendicularly upon the former. The proxi-
mal olfactory ganglia bulbiform, a little smaller than the buccal ones ;
no distal could be found. The length of the commissures equal to the
largest diameter of the pedal ganglia ; the subcerebro-pedal about
three times as thick as the visceral. '! he buccal ganglia of oval form,
connected through a short commissure ; the gastro-oesophageal about
one-sixth of the former in size, with one very large cell.
The eyes with black pigment, yellow lens ; the nervus opticus about
as long as the largest diameter of the cerebral ganglion. ']"he otocyst*
of the same size as the eyes, situated externally at the junction of
the cerebral and the visceral ganglia ; with about fifty ordinary
otokonia, but among them four to six larger ones, of a diameter of
about 0.025 mm. The leaves of t^e rhinophoria nearly without
spicula ; in the axes, and especially in the stalks, on the contrary, an
enormous quantity of large spicula, in great part transversely situ-
1830.] NATURAL SCIENCES OF PHILADELPHIA. St
ated. In the skin a rather large quantity of spicula. The broad
centres of the warts of the back chalk-white in transverse section, on
account of the mass of strong spicula which ascend in bundles through
the axes of the warts, their peripheral parts being free from spicula.
The spicula, for the most part, staff-shaped or cruciate, reaching a
diameter of about 0.08 mm. ; small and large rounded ones were
also very common ; the spicula mostly very strongly calcified. In the
interstitial tissue calcified cells were seen scantily.
The mouth-tube was 1.5 mm. long ; the bulbus pharyngeus about
I. .5 mm. long, the sheath of the radula projecting about 0 75 mm.,
bent upwards ; the sucking-crop nearly as large as the proper bulbus,
short-stalked. The Hp-disk with the cuticula rather thick, especially
at the inferior median line, here sometimes prominent and reminding
one of the two blades in the AcantJwdorides. The tongue (in the six
individuals examined) with seven to nine rows of teeth ; further back-
wards twenty-nine, thirty-one, or thirty-four (in three individuals)
developed, and three younger rows ; the total number of rows was
thus forty-two to forty-six. The median plates (fig. 8a) and the large
lateral (fig. 6«a, Y, 86) ones quite as in the Ad. Pac'fica, also the ex-
ternal ones (fig. 6 J, 8c), but the number of those never surpassed ten
or twelve ;^ frequently all gone from the tongue, and only existing in
the two to four posterior rows ; the height of the large lateral plates
rising to about 0.09 mm.
The salivary glands, as usual, white. The oesophagus somewhat
wider in its first part ; the stomach as usual ; the liver of usual form,
its substance of yellow color ; on the first quarter of the right side an
impression for the anterior genital mass. The vesica fellea rather
smaller, on the right side of and a little behind the pyloric part of the
intestine, with its upper end appearing on the surface of the liver ;
the duct nearly as long as the bag, opening in the stomach.
The sanguineous gland of subquadratic form, the largest diameter
about 2.3 mm., very much flattened, whitish. The tube on the floor
of the renal chamber rather strong.
The hermaphroditic gland clothing the liver with a thin, whitish-
gray layer. The anterior genital mass small, nearly undeveloped,
much compressed, of about 1.75 mm. in length, the height a little less.
The ampulla of the hermaphroditic gland thin, otherwise as usual.
' According to Friele and Hansen (1. c. p. 3) the number of external
plates is twelve ; the figure of these authors (Tab, II, fig. 1) is rather bad.
G. O. Sars has eleven to twelve external plates in his figure.
88 PROCEEDINGS OF THE ACADEMY OF [1880.
The spermatoduct as usual, also the penis.* The spermatotheca and
the spermatocysta as usual. The mucous gland very small, whitish
and yellow.
ACANTHODOEIS, Graj.
Acanthodoris, Gray, Figs, of Moll. Animals, iv, 1850, p. 103, Guide Moll.
Brit. Mas. 1857, p. 207.
AeaniJiodoris, Alder and Hancock, Mon. Brit. Nud. Moll., vii, 1855, p. 43,
app. p. xvii. G. O. Sars, Moll. reg. arct. Norvegise, 1878, p. 308,
Tab. xiv, fig. 4.
Acanthodoris, R. Bergh, Gattung. Nord. Doriden, 1. c, 1879, p. 35G-360.
Forma corporis subdepressa. Nothaeum supra sat grosse villosum.
Branchia (non retractilis) e foliis tripinnatis non multis et in orbem
positis formata.
Caput latum, veliforme ; tentaculis brevibus, lobiformibus. Margo
apertur«um rhinophorialium lobatus.
Discus labialis armatura e hamulis minutis formata et infra cuticula
incrassata prominenti instructus. Lingua rhachide nuda ; pleuris an-
gustis dente laterali, hamiformi permagno et dentibus externie minutis-
(4-8).
Ingluvies buccalis bulbo pharyngeo connata.
Penis armatura e hamulis minutis formata instruetus. Vagina
lonsissima.
'o-
The genus Acanthodoris was established by Gray, to receive the
Doris pilosa with its non-retractile gill. Alder and Hancock adopted
the genus, made an anatomical examination of the typical form and
gave it natural characters, which were then adopted by Gray. In
several new malacological publications of a systematic nature the genus
has been omitted, and in the last twenty years no new information has
been published, until G. O. Sars lately gave some notes on the bulbus
pharyngeus.
The Acanthodorides approach the Lamellidorides, yet differ ex-
ternally in the scattered soft villosities of the back and in the smaller
number of the leaves of the gill, which are arranged in a circle.
Internally they differ still more, in the presence of a strong, oral
armature, in a different dentition (4 + 84-1+0 + 1+8-1-4), by a pecu-
' Sars (1. c. p. 16) mentions and figures (fig. 8) the penis as "a large,
white, conical " organ.
1880.] * NATURAL SCIENCES OF PHILADELPHIA. 89
liarly armed penis and by the imbedding in the pharyngeal bulbus of
the buccal crop,^
The Acanihodorides are not much depressed. The back is covered
with soft villi or papillte ; the openings for the rhinophoria have lobed
margins. The gill is not retractile, and consists of several (generally
seven to nine) tripinnate leaves, quite distinct from one another.^
The labial disk is provided with a densely set armature of small
hooks, passing backward on the cuticula of the mouth. This last also,
in the lowest part of the mouth, at each side of the median line is
thickened and projects like two thin, lancet-shaped blades over the
bare space left between the lower parts of the prehensile collar.^ The
form of the bulbus pharyngeus is as in the Lamellidorides, but the
buccal crop is imbedded in the upper wall of the bulbus, opening into
it through a slit, and is not connected with it by a short stalk.
The tongue is not broad, but nearly fills the buccal cavity, with a flat
furrow for the radula. This last has a naked rhachis, with a low and
narrow, longitudinal fold. The pleuraj contain a very large, com-
pressed, upright, lateral plate, with a large body and a rather short,
strong hook, denticulated or plain along the inner margin ; at the
outer side of the large plate are several (four to eight) small, external
plates (increasing in number backwards). The salivary glands long,
thicker in their foremost part. The oesophagus with a little, crop-like
diverticle at its root. Above the pyloric part of the intestine opens a
' The genus Calycidoris, of Abraham (Notes on some new genera of
Nudibranchiate Moll., Ann. Mag. Nat. Hist., 4th ser., xviii, 1876, p. 132 ;
and Revision of the Anthobranchiate Nudibr. Moll., P. Z. S., 1877, p. 224 j,
which is said to be allied to the Acantlwdorides and Lamellidorides, still
differs by its " subretractile " gill, with simple pinnate leaves, and does
not possess external plates on the radula. The genus is very probably
apocryphal ; iu the phauerobranchiate Dorididm it often happens that the
gill appears as if more or less retracted in a cavity. A single new species
is mentioned, of unknown habitat, the C. OuniJieri, Abr., 1. c., p. 133, PL
vi, fig. 1.
■■' Alder and Hancock mention and figure (1. c, PI. 15, fig. 2, 3) the
branchial leaves as "united at the base ;" so do Meyer and Moebius (1. c,
p. 65) ; this is not the case. The leaves are quite isolated, but there are
usually one or two foliola standing between them, which might simulate a
coherence of the leaves (cf. also PI. xv, fig. 6, A. and H.).
' These thickenings of the cuticle have been regarded, both by Alder
and Hancock, and more lately by Meyer and Moebius (1. c, p. 64, taf. v
A, fig. 8, K 9), as "jaws," but have hardly anything in common with those
organs properly so called.
7
90 PROCEEDINGS OF THE ACADEMY OP [1880.
little sac, which seems to be homologous with the biliary sac (pancreas,
autt.) of other Dorididse. Alder and Hancock, therefore, have de-
nominated that part of the digestive tract as " stomach," although it
in no essential respect differs from the rest of the intestine, and is
just like that part in the Chromodorides, and should be undoubtedly
regarded as the pyloric part of the intestine, when that sac opened
lower down, as in the Chromodorides,^ in the cavity, which is included
in the liver, and seems to be the true stomach. The spermatoduct and
the chief duct of the spermatotheca (vagina) are of very considerable
■ length ; the former consisting of two different parts, a superior softer,
and an inferior very muscular part, internally clothed with an arma-
ture, which is continuous through the penis. This last is rather short,
the superior part solid and projecting as an armed glans into the in-
ferior, hollow part (prseputium). The armature consists of rows of
hooks continued in the interior of the organ, and, as mentioned above,
farther upwards ; quite like that of the Polyceridse^^ Fhyllidiidse^
and Doriopsidae.*
About the biological relations of these forms very little is yet known
and that only with reference to the typical species, through Alder and
Hancock, as well as Meyer and Moebius. The spawn is figured by
Alder and Hancock (1. c, PI. 15, fig. 9), and by Meyer and Moebius
(1. c, fig. 13, 14) ; about the development nothing is yet known.
The few known species of this genus seem limited to the northern
parts of the Atlantic and of the Pacific.
1. Acanthodoris pilosa (O, F. Miiller). Oceanum Atlanticum et Pacificum,
Doris pilosa, Cuv.
Doris stellata (6m.), Cuv.*
> Cf. my Malacolog. Unters. (Semper, Philipp., II, ii), Heft xi, 1877, p.
464-494 ; Neue Nacktschnecken der Siidsee, 11, Journ. der Mus. Godeffroy,
Heft viii, 1875, p. 72-82 ; idem, iv, 1. c, Heft xiv, 1879, p. 1-21.
2 Cf. my Malacolog. Uuters. (Semper, Philipp., II, ii), Heft xi, 1877
(Trevelyana. Nembrotha).
^ Cf. my Bidr. til en Monogr. af Phyllidierne, Naturh., Tidskr. 3, R. V.,
1869 ; Malacolog. Unters. (Semper, Philipp., II, ii), Heft x, 1876, p. 377-
387.
* Cf., 1. c, Heffc X, 1876, p. 384-387 ; Journ. der Mus. Godeffroy, Heft
viii, 1875, p. 82 94.
^ According to Fischer (Note sur quelques especes du G. Doris, decrites
par Cuvier, Journ de Conchyl. 3 ser. x, 1870, p. 290), the Doris stellata,
Cuv., and the D Icevis, Cuv., are identical with his D. pilosa, and this with
the typical form of Miiller.
The D. stellata of Philippi seems a quite different form, a Platydoris
1880. J NATURAL SCIENCES OF PHILADELPHIA. 91
Doris Imvis, Cuv.
? Doris fv sea, O. F. Mull., Zool. Dan. (descr.).'
? Doris tomentosa, Loven, Index Moll. 1846, p. 4.
3. A. subquadrata (Aid. et Hanc). Oceanum Atlanticum.
Doris subquadrata, A. et H. Monogr., Part. V, 1851, fam. 1, Plate
10, f. 1-3 ; Part VII, 1855, p. 43, and III, PI. 46, Suppl. f. 14.
? {D. stdlata, Cuv. ?). Lebert, Beob. iiber die Mundung einiger
Gasteropoden, J. Muller, Arch., 1846, p. 441-446, Taf. XII, fig.
10-13.^
S. A. cmrulescens, Bgh., n. sp. Oceanum Pacificum.
4. A, ornata, Verrill. Notice of recent additions to the mar. fauna of the
eastern coast of North Amer. XXXVIII ; Amer. Journ. of Sc. and
Arts, XVI, 1878, p. 318. Oc. Atlant.
5. A. siellata (Gm.), Verr., 1. c, p. 313, D. bifida, Verr. Oc. Atlant.
6. A. citrina, Verr., 1. c, p. 313. Oc. Atlant.
7. A. f mollicella, Abraham, 1. c, 1877, p. 228, PI. XXX, fig. 1-4. Oc.
Pacincum.
8. A. ? globosa, Abr., 1. c, 1877, p. 228, PI. XXX, fig. 5-9. Oc. Pacif.
I. Acanthodoris pilosa (0. F. MUller). Plate X, fig. 12-15; Plate XI, fig. 1-2;
Pl;.te XII; Plate XIII, fig. 2-5.
Acanthodoris pilosa (O. F. Muller), Alder and Hancock. Monogr. Br.
Nudibr. Moll., Part V, 1851, fam. 1, Plate I, f. 1, 3-5, 12; Plate 2,
f. 2-6; Plate 15; Part VII, 1855, Plate 46; Supph Plate 48, f. 1.
Doris pilosa (O. F. Miiller), Meyer und Moebius, Fauna der Kieler Bucht,
I, 1865, p. 63-67 c. tab.; taf. V, A.
" Color pagin'cB superioris corporis albus vel luteus vel fuscus vel
griseus vel rubro-brunneus vel niger.
Dentes radute barao pro parte denticulato.
Hab. Oceanum Atlanticum septentr., Pacific, septentr.
< Platyd. FMlippii, Bgh.). Cf. my Malacolog. Untersuch. (Semper, Philipp.
II, ii.). Heft, xii, 1877, p. 507.
' It is in most cases a quite useless task to try to elucidate the species of
Dorides of the elder authors ; their examinations vrere all too superficial
and their descriptions don't contain the data necessary for their verifica-
tion. The best way would be to wholly cancel these names {D. fusca, M.;
D, loRtis, L., etc.) which have given later authors so much trouble. On the
Doris fusca of O. Fabricius, Morch has even formed a genus Proctaporia
(Rink. Gronland. I, 1857. Tillag. 4, p. 78), that must be cancelled, too.
* The short statements of Lebert about form and color of the animal
examined by him can scarcely entirely prohibit the identification of it with
the species described by Alder and Hancock. The figures of the (tongue)
teeth given by Lebert, rough as they are, suffice, on the other hand, to
secure the identification with the D. subquadrata, or at least with a nearly
related species.
92 PROCEEDINGS OP THE ACADEMY OF [1880.
Of this species I have had a lot of specimens for examination, all
preserved in spirits; partly (two) from the neighborhood of Bergen
(Norway), kindly sent by Mr. Friele, partly (one) from the Frith of
Kiel, sent by Prof. Moebius ; but particularly (seventeen) from the
coast of Denmark (Striib, lille Bait.)
The individuals varied much in color. The variability of the color
is noted by Alder and Hancock, They were whitish, or whitish sprinkled
with brownish, or dark (bluish) gray, or yellowish, or brownish, or
reddish-brown on the back, with whitish or yellowish sides and foot.
The length reaching 12.0 mm., by a breadth of 80 and a height of
5.0 mm. ; the foot then about 4.0 mm. broad, the branchial leaves
reaching to the height of about 1.0 mm.
The back covered all over with the soft, slender, conical and pointed,
erect (or curved) papillas of very different sizes, most of them small ;
between these are larger ones ;^ some of the largest divided into two
or three points, and some of them connate and forming small crests,
divided above into two or three points. The margins of the sheaths
of the rhinophoria rather prominent, divided into several (six to eight)
smaller and larger pointed lobes ; the club of the rhinophoria with about
twelve to twenty leaves.^ The branchia, in both Norwegian specimens,
with eight tripinnate leaves, otherwise with seven to nine (as men-
tioned by Meyer and Moebius). The anal papilla low, with several
papillulag and a star-shaped aperture ; on a low crest, issuing from its
posterior, is a strong papilla. The head and the tentacles (Plate X,
fig. lib) as figured by Alder and Hancock (1. c, Plate 15, fig. 1).
The anterior margin of the foot with a fine transverse furrow (Plate X,
fig. 14a). The genital opening is a longitudinal slit (Plate XI, fig. 2).
The peritoneum was mostly of reddish-brown color.
The central nervous system showed^ the cerebral ganglia rounded-
triangular, not much flattened, a little larger than the more rounded
visceral, which lie behind and on the outside of them and show a slight
notch in the outside ; on the inferior side of the visceral ganglia the
pedal ones are set nearly perpendicular on the latter, connected by the
^ Alder and Hancock, also Meyer and Moebius give eighteen to twenty
leaves. Cf. the figures 7-8 of Meyer and Moebius.
* Collingwood (Ann. Mag. N, H., 3 ser. vi, 1859, p. 463) remarks that it
" when not in motion; bears a great resemblance to a miniature hedgehog."
^ The representation of the system given by Hancock and Embleton (On
the anatomy of Doris, Philos. Transact. MDCCCLH, Plate 17, f. 8) is not
very like nature.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 93
three distinct commissures, which are nearly as long as the diameter
of the ganglia. From the outer part of the right visceral ganglion
issues a nerve nearly as long as the transverse diameter of the whole
central nervous system and swelling to a rather large ganglion (gangl.
penis) at the root of the penis ; this ganglion contains only rather
small cells and gives off three or four strong and several thinner nerves
(Plate X, fig. 15). The part of the brain which gives off the nervus
opticus, simulates a ganglion. The proximal ganglia olfactcia bulbi-
form, somewhat smaller than the buccal ganglia, but much larger than
the distal ganglia olfactoria ; the buccal ganglia flattened, rounded, con-
nected by a rather short commissure ; the ganglia gastro-oesophagalia
rounded, having about one-fifth of the size of the last, containing one
very large cell and a few smaller.
The eyes with black pigment and yellowish lens. The otocysts
lying at the hinder part of the cerebral ganolia, as large as the eyes ;
with numerous small otokonia, which in the specimens from Kiel,
were not much calcified. No trace of spicula in the leaves or other
parts of the riiinophoria. The spicula of the skin were, so to speak,
limited to the margins of the mantle and of the foot ; in the last they
were chiefly arranged perpendicularly or obliquely against the margin,
except that in the foremost and hinder part of the sole some few spic-
ula were seen scattered.
The amount of spicula in the skin seems to vary notably in the
Acanthodoris piJosa, as seems to be the case in general in different
forms of Dorididce, especially, as far as hitherto known, in the Poly-
ceratidce {Polycera, Ancula). (Cf. Meyer and Moebius, Fauna der
Kieler Bucht, I, 1865, pp. 52, 60.) Frey and Leuckart (Beitr. zur
Kenntn. wirbellose Thiere, 1847, p. 145 ; described a very regular
position of the spicula, but not, as it seems, in accordance with nature.
In the margin of the mantle the spicula were arranged as figured by
Alder and Hanc., 1. c. Part VII, PI. 48, supplem. fig. 1, only more con-
centrically at the transition from the margin to the side of the body; a
narrow belt of spicula crossed the back before the region of the gill.
Some spicula were also seen in the tentacles. The spicula reached a
notable length (at least 0.6 mm.^, in old individuals they were more
calcified than in younger ones. The skin was filled with unicellular
glands, especially in the dorsal papilla?.'
The mouth-tube was wide and strong, about 1.5 mm. long; the
bulbus pharyngeus in the largest individuals about 2.75 mm. long, by
^ Cf. the (not very good) fig. 6 by Meyer and Moebius.
94 PROCEEDINGS OF THE ACADEMY OF [1880.
a breadth of 2.0 and a height of about 3.0 mm. ; the sheath of the
radula projecting backward nearly 1.0 mm. ; the lip-disk sometimes
surrounded by a ring of black pigment. The armature of the lip-
disk entirely as shown (PI. XII, figs. 1-4, 10-11) by me in the form
from the Pacific, also the crop (PI. XIII, fig. 2) of the bulbus.'
The tongue in the eight specimens examined was provided with five,
seven, eight, nine rows of plates, farther backwards also sixteen to
twenty developed, and three younger rows; the total number amount-
ing thus to from twenty-seven to thirty.^ The large latei-al teeth^ yellow
in the body, especially in the anterior-inferior part, with commonly five
to eight denticles on the inside of the hook ; sometimes, especially in
the younger plates, the number of denticles rose from eleven to fifteen,
sometimes the three to four outermost denticles were much larger than
the rest, sometimes the denticulation was quite irregular ; the height
of this plate reached 0.4 mm. The outer plates (PI. XI, fig. 1) com-
monly four to six, seldom seven to eight ; in a series of four on the
hinder part of the tongue, the outermost measured about 0.05, the next
0.09, 0.11, 0.125 mm.; they were quite colorless, compressed, with
the upper side flattened, and rather erect.
The salivary glands as in the purple-colored form from the Pacific.
No constant dilatation of the middle of the oesophagus (as figured,
PI. I, f. 12^', by Alder and Hancock), but a strong, particular one at
the root as figured (1. c. PI. I, f. 12/j by Alder and Hancock and by
me (Gatt. nordischer Doriden, 1. c. Taf. XIX, fig. 14r). The
stomach as in the Pacific form ; the intestine sometimes dilated in its
first part, sometimes absolutely of the same caliber as the rest, and
neither externally nor internally different from it ; a little bag
(biliary sac) which has been noticed by Alder and Hancock (1. c. PI,
I, fig. 12^'), opening into the right side of this part of the intestine.
The posterior visceral mass (liver) flattened and excavated on the
anterior-inferior right half. The sanguineous gland whitish, convexo-
concave, short and irregularly kidney-formed, with the excavation
^ The first specimens of the Nortliern Atlantic left at my disposition
being too small and too few for a thorough examination, I am obliged to
refer to my examination given herewith of the form fiom the Pacific. Cf.
moreover my figures iu "Gatt. uord. Doriden," 1. c. PI. XIX, figs. 10, 11.
The crop is rather well figured by Alder and Hanc. (1. c. PI. I, f, 12c).
* According to Meyer and Moebius, the number of plates ("of the
radula") is thirty-one, to Alder and Hancock, twenty-seven.
* Cf. my Gattungeu nordischer Doriden, 1 c. Taf. XIX, fig. 12.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 95
forwards, transversely situated, with a largest diameter of 3.0 mm.
The renal chamber and the syrinx as in the form from the Pacific.
The hermaphroditic gland as in this last variety, its white color con-
trasting with the hue of the liver. The anterior genital mass of short
pyramidal form, with the point outwards, about 4.75 mm. long, the
breadth and the height a little less. The ampulla of the hermaphro-
ditic gland yellowish-white, forming a single ansa, about 4.0 mm. long,
by a diameter of 0.75 mm. lying on the upper part of the back of the
mucous gland. The spermatoduct yellowish, about 15.0 mm. long,
constricted a little above the middle of its length ; strong, sloping into
the penis, which is about 1.0 mm. long. The armature of the penis
entirely as in the form from the Pacific, continued backwards in the
interior of the spermatoduct for a length of 6.0 mm. ; the hooks
rising to the height of about 0.035 mm., nearly colorless.^ The sper-
matotheca (PI. XIII, fig. 5a) spherical, of a diameter of about 2 0
mm., greenish or whitish ; the spermatocysta (fig. 56) much smaller,
pyriform, yellowish ; both filled with sperma The chief duct (the
vagina, fig. 5dd) very long, with several (four) longitudinal folds, which
are folded again transversely ; the structure seemed to resemble en-
tirely the form from the Pacific ; in the cavity was more or less sperma.
The mucous gland yellow and yellowish-white ; the fold of the duct
with brownish-gray points, but no black pigment on the lower part of
the vagina or penis.
One specimen of this typical form, with "brown mantle," and in
all respects agreeing with the Atlantic, was dredged by Dall at Kyska,
in June, 1873, on rocky bottom at the depth of ten fathoms.
An individual of a (in living state) " yellowish-white " variety was
dredged by Dall in Popoff Strait (Shumagin Islands), on rocky bot-
tom at a depth of six fathoms.
The animal preserved in spirits was 10.0 mm. long, by a breadth of
6.0 and a height of 4.5 mm. ; the rhinophoria 1.5 mm. high, the eill
1.0 mm., the foot 3.0 mm. broad. The color yellowish-white. In the
club of the rhinophoria about thirty leaves ; nine branchial leaves ;
the anal papill.i with three small protuberances ; the renal pore very
distinct on the right side. The genital opening very wide ; the bul-
bus pharyngeus 2.0 mm. long ; the tongue with seven rows of plates,
the total number of these twenty-six (16 + 3) ; five external
^ The armature of the penis has been first seen by H. Friele and G.
Armauer Hansen (Bidr. til Kundsk. om de Norske Nudibranchiar. Christi-
ania, Vidsk. Selsk. Forh., 1875, extras, p. 4).
96 PROCEEDINGS OF THE ACADEMY OF [1880-
plates. The diverticle of the oesophagus nearly as large as the true
bulbus. The spermatoduct and the penis as usual, also the vagina ;
the spermatotheca of 1.6 mm. largest diameter. No trace of pig-
ment on the vagina or penis, and the peritoneum was colorless.
Another variety of the species, with " brown mantle and yellowish-
white papilljB," was dredged by Dall, in Yukon Harbor (Shumagins),
in August, 18t4, on sand and stones, at a depth of six to twenty
fathoms.
The individual preserved in spirits was 9.0 mm. long, by a breadth
of 6.5 mm., and a height of 4.5 mm. ; the breadth of the foot 4.0 mm.,
the height of the gill 1.5 mm. The back of the animal densely brown-
dotted, especially the circumference of the gill and the free area left
in the middle of the gill ; the dorsal papillae all whitish ; the stalk of
the rhinophoria and the inferior part of the club densely dotted with
brown, also, in a somewhat slighter degree, the outside of the branchial
leaves. The under side of the mantle and the upper side of the margin
of the foot and, in a slighter degree, the sides of the body and the sole
of the foot dotted with an enormous quantity of brownish-gray points.
The form as usual. The gill with nine leaves, of which the two pos-
terior were much smaller than the others.
The central nervous system as usual ; the otocysts very conspicuous
under the magnifier as chalk-white points. The mouth-tube 2.0 mm.
long. The bulbus pharyngeus 2.0 mm. long ; the sheath of the radula
projecting 2.0 mm., bent downwards. The armature of the lip-disk
(PI. XII, fig. 10, m very like that of the var. albescens (cf. PI. XIII,
fig. 4). The buccal crop as usual. The tongue with nine rows of
plates; the total number of rows, twenty-five (13 f 3). The large
lateral plates as usual ; the djnticulations rather long and somewhat
irregular. The number of the external plates (fig. 12) reaching to six.
The salivary glands, the oesophagus with its diverticle, the pyloric
part of the intestine with its bag (biliary sac), and the liver, as usual.
The sanguineous gland rather large, covering, besides the central
nervous system, the whole of the bulbus pharyngeus.
In the lobes of the hermaphroditic gland, masses of zojsperms. The
anterior genital mass of the usual form ; the ampulla of the herma-
phroditic duct somewhat larger. The spermatoduct as usual ; so, too,
the penis, with its armature ; the length of the glans about 0.5 mm.
The spermatotheca and the spermatocjsta as usual; also the chief
duct (vagina), the cavity of the last filled with sperma. The mucous
gland yellowish-white and in the centre (albuminous gland) brownish-
1 880. J NATURAL SCIENCES OF PHILADELPHIA. 91
yellow. Very scanty black pigment on the inferior part of the vagina
and of the penis ; the peritoneum of the back, on the contrary, very
(lark brown.
2. Acanthodoris pilosa (0. F. MUUer), var. albescens, PL X, fig. 14, 15 ; PI. XI, fig. 2 ;
PI. XII, fig. 13-16.
Color flavescente-albidus.
Hamus dentium (lingua;) edentulus vel parce denticulatus.
Habitat. Oceanum Pacificum septentrion. (Aleutian Islands).
Two rather large specimens of this variety have been dredged by
Dall, in June and July, 1813, at Kyska Harbor (Aleutians), on sand
or on rocky bottom, at a depth of nine to fourteen fathoms.
According to Dall, the color of the living animal was "yellowish-
white ; " that of the specimens preserved in spirits was so, too, but
very likely much more whitish. The length was 16.0 or 17.0 mm.,
by a breadth of 6.5 to 8.0 mm., and a height of 6.5 mm. ; the h^^ight
of the rhinophoria 2.5 to 3.0 mm., of the gill 3.0 to 4.0 mm. ; the
breadth of the foot 5 0 or 6.0 mm., the length of the genital opening
2.0 or 3.0 mm. The form as in the typical D. jnlosa; the rhinophoria
showed about twenty-five broad leaves in the club ; there were nine
branchial leaves ; the anal papilla very low ; the renal pore rather large.
The central nervous system as previously described. The distal
olfactory ganglion small ; a large (diameter, 0.4 mm.) ganglion penis
(fig. 15). The eyes with rich, coal-black pigment ; the otocysts visible
under a lens as chalk-white points, with about one hundred and fifty
otokonia.
The bulbus pharyngeus 8.5 mm. long, with the sheath of the radula
projecting 1.3 to 1.5 mm.; the height of the bulbus, with the crop,
4.0 to 4.5 mm., its breadth 2.5 to 3.0 mm.
The older elements of the lip-plate (PI. XII, figs. 13, 14) agreeing
in form with those of the typical species, but oftener showing a granu-
lated interior ; the said elements reaching a length of about 0.04 mm.
The diameter of the disk and mouth about 3.0 mm. The breadth of
either half of the disk 0.66 mm.
The tongue showed nine or ten rows of teeth ; the whole number of
rows, twenty-nine (16 or 174-3). The large lateral teeth were as in the
typical species, reaching 0.65 mm. in height (PI. XII, fig. 15, 16),
without or with only a very slight denticulation of the hook (fig. 15).
The number of the outer teeth, three to five.'
' Cf. my Gatt. nordischer Doriden, 1. c, Taf. xix, fig. 13.
98 PROCEEDINGS OP THE ACADEMY OF [1880.
The salivary glands deeply imbedded in the cavity for the oesophagus
at the fore-end of the liver. The oesophagus with its rather large
(1.5 mm. long) diverticle, the stomach, the intestine with its little
(1.0 mm. long bag, as above. The liver 7.0 to 9.0 mm. long, 5.0 to G.O
mm. broad, 5.0 to 6.25 mm. high, of yellowish-gray color. The san-
guineous gland of irregular, oval form, of a largest diameter of 4.0
mm., by a thickness of 1.0 mm., and of grayish color. The renal syrinx
about 0.75 mm. long.
The anterior genital mass Q.O or 7.0 mm. long, 4.0 to 6.0 mm. high,
and 3.0 or 4.0 mm. thick. The ampulla as usual; also the (about
40.0 mm. long) spermatoduct and the (nearly 2.0 mm. long) penis,
with its armature ; the hooks often set in pairs. The spermatotheca
(diameter, 4.0 mm.) and the spermatocysta (diameter, 1.5 mm.) as
above ; the chief duct, with the vagina (about 23.0 mm. long, by a
diameter of 0.4 to 1.0 mm.), as usual, and also its internal cellular
clothing (PI. X, fig. 13) ; the yellow nucleoli somewhat brighter; the
cavity nearly filled with sperma. The mucous gland as usual. No
black pigment on the inferior part of the vagina or on the penis.
S. Acanthodoris pilosa (0. F. MUller), mr. purpvrea, PI. XII, fig. 1-9.
Color e purpureo brunneus et flavescente-albidus.
Habitat. Oceanum Pacificum septentrion. Insula? Aleutiana? (Una-
lashka).
Only two specimens of this species were dredged by Dall, in Sep-
tember, 1874, on mud and stones, at a depth of about sixty fathoms.
The color of the living animal was, according to Dall, '■ purple-brown
and jellowish-white." The length of the animals preserved in spirits
was 24.0 or 25.0 mm., by a breadth of 9.0 or 10.0 mm., and a height
of 7.5 mm.; the foot 6.0 mm. broad; the height of the rhinophoria
about 3.0 mm., of the branchial leaves 2.3 mm. The color of the
back reddish-brown ; the stalk of the rhinophoria brownish, the club
yellowish ; the branchial leaves yellowish-white, the last brownish at
the rhachis ; the under side of the mantle margin, with the sides of
the body, the head and the foot, yellowish-white, dotted with brownish-
gray all over, the color much more scanty on the sides of the foot and
still more so on the head and on the sole of the foot.
The form was somewhat elongate. The back covered all over with
pointed, rather (0.75 mm ) high, digitiform, soft papilla3 and with inter
mixed smaller ones. The margin of the rhinophor-holes with several
pointed, projecting, digitiform processes ; the stout club of the rhino-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 99
phoria with about twenty leaves. The branchial leaves stronjr, (in both
individuals) eiglit in number, the two hindermost separateil by a narrow
crest, which rises into a larger papilla ; before this the anal papilla,
covered with some papilloe, at its right side is the r^nal pore ; on the
space before it were several smaller papilla?. The under side of the
free margin of the mantle (about 2.0 mm, broad) smooth. The head
large, the tentacles short. The genital opening a rather large, cres-
centic orifice. The foot rounded behind.
The peritoneum was richly dotted on the back with brownish-red.
The central nervous system nearly quite as in Ac. pilosa; the
proximal olfactory ganglia of oval form, true distal ones could not be
detected in the root of the rhinophoria, but only a fusiform swelling of
the nerve, with scattered nervous cells. The subcerebral and pedal
commissures connected, the visceral isolated. The buccal ganglia
larger than the olfactory, of oval form, connected by a commissure
nearly as long as each ganglion ; the gastro oesophageal ganglia de-
veloped on the side of the nerve, which is a little longer than the
ganglion, and in size about one-fifth of the former ; the contents one
very large cell, three or four smaller and several quite small ones.
On the upper part of the penis the large ganglion genitale, of about
the diameter of 0,3 mm., rounded, partly covered with black pigment
consisting of only rather small cells ; in the first parts of the nerves
given off from the ganglion, one or two rows of nervous cells of the
same kind as in the ganglion.
The eyes with black pigment, yellow lens ; the optic nerve rather
long. As chalk-white points the otocysts were situated on the hinder
part of the cerebral ganglia, where they touched the pedal ones ; they
were filled with solid, yellowish otokonia of about the usual form and
size, but, in both respects, rather irregular. In the leaves of the
rhinophoria no spicula. In the margin of the mantle and of the foot
almost no spicula at all, but everywhere in the skin, especially on the
back and the papilla, were an enormous quantity of large and small
glandular openings. In the interstitial connective tissue were hardly
any calcified cells at all.
The mouth-tube was about 2.3 ram. long, wide, with a glandular belt
on the outside, not closed below ; on the inside lined with a yellowish
cuticula. The bulbus pharyngeus strong, about 4.0 mm. long, and
the sheath of the radula projecting nearly 10 mm. from the posterior
part of the under side, directed straight backwards or downwards ; the
height (through the buccal crop) 4.0 mm., the breadth 2.5 mm. The
100 PROCEEDINGS OF THE ACADEMY OF [1880.
buccal crop making nearly half of the whole bulbus, and of the usual
form ; the walls very thick ; the compressed and rather small cavity
communicating through a long cleft with the anterior half of the
small buccal cavity. The lip-disk (fig. 1) of rounded contour, clothed
throughout its whole breadth (on each side to about 0.5 mm.) with the
light, horn-yellow colored armature ; the lowest part of this, as usual
in the Acanthodorides, injured or wanting ; the breadth of the belt
decreasing towards the upper end, where it is interrupted in the middle
line, also at the lower end. The armature (fig. 2&&, 36, 4} composed
of hooks, whose points are directed forwards (towards the opening of
the mouth), nearly like, but still differing a little from those in the
typical Ac. piloHO, reaching the height of about 0.04 mm., yellowish,
with rounded, bifid or irregularly cleft points. The lancet-shaped
(fig. la, 2a, 3a) blades at the inferior angle of the mouth as usual.
The tongue with nine or ten sei'ies of plates, farther backwards
thirteen to fifteen developed and three undeveloped series ; the total
number in this way, twenty-five to twenty-eight. The large lateral
plates relatively larger than in the Ac. pilosa, and (fig. 5, 6) less
thick in the anterior-inferior part of the body, with relatively larger
hook ; the denticulation of this last much weaker and much more
irregular ; in one specimen generally two to four denticles, sometimes
only a few vei'y insignificant ones or none at all (fig. 6) ; and this was
the case with the other specimen, in which only some few plates showed
two small denticles.* The outer lateral plates as in the typical form,
scarcely more than from four to six.
The salivary glands whitish, rather strong at their short first part,
in the rest of their length thin (fig. 7), accompanying the oesophagus
to the cardia ; the duct rather short (fig. 7a).
The oesophagus forming a little crop,^ with thin walls and longitu-
dinal folds on the inside; in the rest of its length rather thin. The
stomach rather small, with the usual biliary apertures. The intestine
(fig. 8a) somewhat inflated in its first part, with many rather strong
folds and one particularly thick ; a little over the point, where it
appears on the surface of the visceral mass, on the right side, a little,
scarcely pedunculated bag (fig. 8&^, of the length of 1.0 to 1.25 mm.,
with fine, longitudinal folds ; the rest of the intestine (fig. 8c) some-
what narrower; the total length of the intestine about 12.0 to 13.0
^ Although very like the plates of the Atlantic form, they still bore a
somewhat peculiar aspect.
■^ Cf, my Gattungen uordischer Doriden, 1. c, Taf. xix, fig. 14.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 101
mm., by a diameter of 1.0 to 1.5 mm. The contents of the stomach
and of the intestine indeterminable animal matter, mixed with an
enormous quantity of different and partly very handsome forms of
Diatomacese, with some polylhalamia and some small copepoda, and
fragments of the same.
The liver about 9-9.5 mm. long by a breadth (at the forepart) of
6.5-5.5 and a height of 6.25-6.0 mm. ; the posterior half somewhat
pointed, the anterior notably flattened and excavated on the right side ;
around the cardia the liver appeared naked (not covered by the her-
maphroditic gland) of (greenish) gray color, in sections it was yel-
lowish.
The ramifications of the aorta nearly as in the typical Dorididce,^
the root of the posterior aorta still longer and the Art. syringis renalis
stronger and more ramified. The sanguineous gland yellowish-white,
rather flattened, of irregular triangular form, lobulated, about 3.5 mm.
long.
The renal chamber large ; the yellowish-white renal syrinx about
0.75 mm. long, its tube somewhat more than twice as long, imme-
diately continuous with the tube on the floor of the renal chamber.
The hermaphroditic gland easily distinguishable from the liver
through its more whitish color; the secondary (ovigerous) lobes rather
small; in the lobes zodsperms and large oogene cells. The anterior
genital mass of plano-convex heart-shape with the point down and
backwards; the length about 5.0 mm. by a breadth of 4.0 and a height
of 5.0 mm. The ampulla of the very thin and white hermaphroditic
duct resting on the upper posterior part of the mucous gland, yellow,
short and thick (4.0 mm. long by a diameter of about 1.25 mm. form-
ing a simple ansa. The vas deferens yellowish, strong, resting upon
the upper side of the genital mass with its large coils and freely de-
scending before its anterior margin to the penis, constricted about the
(fig. 9c) middle of its total length (30.0-35.0 mm.). The penis forming
the end of the spermatoduct somewhat thicker, about 2.0 mm. long,
somewhat curved ; its lower part hollow, the rest solid and prominent
in the cavity of the former as a cylindrical glans of the length of about
0 6 mm. The glans with about ten series of yellowish hooks, which
from a rather large basis raised to the height of about 0.04 mm. ; the
continuation of the armature reaching through the interior of the glans
and of the spermatoduct nearly up to the stricture of the last, but the
» Cf. my Malacolog. Unters. (Semper, Philipp.) Tab. XLVIH, fig. 11.
102 PROCEEDINGS OF THE ACADEMY OF [1880.
number of sei'ies here smaller, about five to eight. The spermatotheca
whitish, forming an oval bag of 3.0 mm. largest diameter ; the sperma-
tocysta yellowish, of 1.3-1.5 mm. largest diameter, the ducts as in the
typical Ac. pilosa. The chief duct, too, very (about 25.0 mm.) long,
rolled up in many coils, partly spiral, the diameter varying between about
0.3 and 0.15 mm. ; the last fourth of the duct (vagina) with scattered
black pigment, somewhat narrower and with a rather strong retractor
muscle at its commencement ; the interior of this duct with some few
strong longitudinal folds, clothed with a cuticula, and under the same a
very fine layer of round and angulated cells with a large round or oval
nucleus of the diameter of about 0.4 mm. and a rather large yellow
nucleolus (PI. X, fig." 13). In the cavity of the vagina more or less
sperma.i The mucous gland yellowish and white ; the central mass
(albuminous gland) yellow ; the duct with scattered black pigment on
the outside (also on the outside of the lower part of the penis), with
the usual fold. The vestibulum genitale with black pigment on the
folds, the same pignaent was seen in the lowest part of the cavity of
the penis and of the vagina and on the folds of the duct of the mucous
gland.
A very similar animal, but " with brown mantle,'' was dredged by
Dall in Kyska Harbor (Aleutians) in July, 1873, on sand, at a depth
of nine to fourteen fathoms.
It was of large size ; the length 21.0 mm., by a breadth of 11.0 and
a height of 9.0 mm. ; the margin of the mantle 2.0 mm. broad, the foot
6.0 mm. broad ; the height of the rhinophoria and of the gill 3 mm. ;
the genital aperture 3.0 mm. broad. The color dirty brown on the
upper side ; the rhinophoria and the branchial leaves yellowish, dotted
with grayish, especially on the stalk of the rhinophoria ; the sole of
the foot yellowish, the under side of the animal whitish ; the under
side everywhere with an enormous quantity of gray and black dots.
The number of branchial leaves nine.
The peritoneum black-brown ; the central nervous system, eyes,
otocysts, as previously described. The bulbus pharyngeus of the
length of 4.5 mm. by a breadth of 3.0 and a height (with the crop)
of 4-T5 mm. ; the sheath of the radula projecting 1.25 mm. ; the crop
alone of the height of 2.3 mm. and 3.25 mm. broad. The lip-disk as
above, the thickenings in the lowest part of the mouth 1.2 mm. long,
of which nearly half freely projected. On the tongue nine rows of
' The length of the spermatoduct and the duct of the spermatotheca
(vagina) was much more considerable than in the typical form.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 103
plates, farther backwards eighteen developed and three younger rows,
the total number thirty; the plates denticulated as previously men-
tioned, the height of the large plates rising to 0,7 mm. ; the number
of external plates four to five. The oesophageal diverticle of a largest
diameter of about 3.0 mm. The pars pylorica of the intestine of about
4.5 mm. length, with higher folds than in the rest of the intestine, which
had a length of about 15.0 mm. ; the bag at the first part of the intes-
tine 1.5 mm. long. The liver 12.0 mm. long by a breadth of 8.0 and a
height of 6.0 mm. The sanguineous glands whitish, 5.0 mm. long by
a breadth of 6.0 mm. and 2.0 mm. thick, convexo-concave, the fore-end
flattened (by the buccal crop), the hinder end with two transverse fur-
rows (produced by two coils of the spermatoduct ; the anterior genital
mass 8.0 mm. long by a breadth of 3.5 and a height of 7.5 mm. The
ampulla of the hermaphroditic duct 5.0 mm. long, whitish. The coils
of the spermatoduct and of the vagina in this individual covering the
upper side of the mucous gland, and ascending to the back between
the pharyngeal bulbus and the liver ; a coil of the former embraced
the sheath of the radula. The first part of the spermatoduct 12.0 mm.
long, the last of the length of about 25.0 mm ; the penis about 3.5 mm.
long, the armature as usual. The spermatotheca nearly spherical, of
3.5 mm. diameter ; the spermatocysta yellowish, round, with a diameter
of 1.5 mm. ; the chief duct (vagina) 33.0 mm. long with a general
diameter of 1.2 mm. ; the structure of the wall as above ; the last, nar-
rower part (from the m. retractor downwards), 5.0 mm. long. 'J he
vestibulum, as well as the inferior part of the vagina and of the penis,
with very scanty black pigment.
4. Acanthodoris caerulescens, Bgh., n. sp. Plate XIII, fig. 6-7 ; Plato XIV, fig. 16.
Color paginas superioris corporis caerulescens.
Dentes radulae hamo per totam fere longitudinem deuticulato.
Hah. Mare Beringianum (Nunivak Island).
One specimen of this species was found by Dall at the north end of
Nunivak Island, Bering Sea, in July, 1874, on stony bottom, at the,
depth of eight fathoms.
According to Dall, the color of the living animal was bluish. The
animal preserved in alcohol had the length of 14.0 mm. by a height of
5.0 and a breadth of 8.0 mm. ; the length of the foot was 12.5 mm. by
a breadth of 6.5 mm. ; the height of the rhinophoria 2.0, of the bran-
chial leaves 1.5 ram. The color uniformly yellowish-white, with the
back of a slightly bluiah hue.
104 PROCEEDINGS OF THE ACADEMY OF [1880.
The form elongate-oval. The back covered all over with irregular
(the greatest height reaching about 1.5 mm.), conical, rather soft and
flexible papillte, in general larger than in the typical species. The
margin of the rhinophor-holes thin, somewhat prominent, with two
anterior strong tubercles and a posterior much smaller one ; the stalk
of the club rather low, the latter with about twenty-five to thirty leaves.
The branchia consisting of nine to ten leaves, the adjacent border set
with several strong tubercles ; the branchial leaves quite isolated at
their base, apparently simply pinnate. The anus prominent, before
the same a small tubercle, behind it a much larger one. The margin
of the mantle rather thin, on the upper side covered with a mass of
smaller and larger papillse and tubercles, the under side smooth. The
head broad, flat, with prominent rounded, flattened tentacula. The
foot broad, rounded behind.
The central nervous system as in the typical species ; the buccal
ganglia rounded, the commissure between them very short. The eyes
with black pigment and yellow lens. The otocysts a little smaller
than the eyes, with numerous otokonia of the usual form, and reaching
a length of 0.03 mm. The leaves of the rhinophoria without spicula ;
in the axes of the organs large, molecularly calcified cells and groups
of smaller calcified cells. In the papillae of the skin of the back were
no spicula at all, on their surface the usual large quantity of glandular
cells ; in the skin beneath the papillte cells and groups of cells as in the
case of the rhinophoria.
The mouth-tube rather wide, with strong cuticula. The bulbus
pharyngeus formed apparently as in the typical species ; the lip-plate
composed of many rows of rather low (the height rising to about
0.02 mm.), very (fig. 6) finely striated columns. The tongue with ten
rows of teeth ; further back, twenty-six developed and three undevel-
oped rows ; the total number thus thirty-nine. The lateral plates
large, yellow, of usual form, with a series of denticles along nearly
the whole of the inner margin of the hook (fig. 16a). The external
plates colorless, eight in number; somewhat depressed (fig. 7, 16),
obKquely rising from the cuticula of the tongue (fig. 7), of nearly equal
size excepting the outermost (fig. 16&), which is much smaller.
The salivary glands seemed of the usual form. The oesophagus and
the stomach as usual. The intestine issuing from the liver at the
middle of its length on the left side, rather short. The liver of the
length of about 9.0 mm. by a breadth and a height of about 4.2 mm. ;
1880.] NATURAL SCIENCES OF PHILADELPHIA. 105
the right anterior half excavated (on account of the anterior genita
mass) ; the color brownish-gray.
The heart and the sanguineous gland as usual, also the renal cham-
ber and the renal syrinx.
The hermaphroditic gland by its yellowish color contrasting with
the liver, clothing the under side, part of the left side, and its right
anterior half. The anterior genital mass rather compressed, about
6.0 mm. long by a breadth of 2.0 mm. The ampulla of the hermaphro-
ditic duct rather short, sausage-shaped, about 2.3 mm. long, curved
and whitish. The larger part of the penis was gone, but hooks were
seen in the remaining part as in the typical species. The sperma
totheca rather large, bag-shaped, about 3.5 mm. long ; the vagina
rather wide, about 10.0 ram. long. The mucous gland white, and the
albuminous gland yello wish- white. ^
This species seems very distinct from the typical one, by its color
and by the different form of denticulation of the large plates of the
tongue.
POLYCERATID^.
This large family, so rich in generic forms, was found represented
in the northern Pacific only by two generic types, Polycera and
Iriopha.
POLYCERA, Cuvier.
Polycera. Cuvier, (1812?), Regne-anitn., 1817, ii, p. 390.^ Regne-anim.,
ed. 2, ill, p. 52.
Themisto, Oken, Lehrb. der Zool., 1815, p. 278.
Cufaa, Leach, Moll. Britann. Synopsis, 1852, p. 21.
Polycera C, Aid. and Hanc, Observ. on the genus Polycera, Ann. Mag.
of Nat. Hist., vi, 1841, p. 337-342, PI. IX.
Limacia, O. Fr. Miiller, Zool. Dan., i, 1781, p. 65-68.^
Phanerobranchus, A. Fredol (Moquin-Tandon), Le monde de la mer, 1864,
PI. xii, figs. 1, 2.
* The anterior genital mass was so hardened and altered, tliat the nature
of its different components could not be determined with certainty.
2 According to a note of Hermaunsen, under the genus Themisto, Oken,
(Ind. Gen. Malacoz, primordia, ii, 1849, p. 572), the genus Polycera was
established by Cuvier, 1812, [but this is probably a typographical error,
since, under the genus Polycera itself, he indicates only the year 1817 —
Dall,] (cf., 1. c, p. 314).
' Limacia, Hartm., Neue Alpina, i, 1821, p. 208 [Arion, Fer.).
8
106 PROCEEDINGS OF THE ACADEMY OF [1880.
Limbus frontalis digitatus vel tuberculatus. Branchia S-Y-foliata.
Appendices dorsales (extrabranchiales) 1-3. Tentacula brevia, lobi-
formia.
Lamelhe mandibulares laterales forte?, sat applanataj. Radula
rhachide nuda ; pleuris dentibus lateralibus hamatis duobus (margine
Isevi), interne minore, externo majore, et dentibus externus 4-8.
Prostata magna ; pleuris ut in omnibus Polyceratis.
The genus Polycera was established by Cuvier (1812 ?), to receive
the Doris quadrilineata of Miiller and (in 1830) allied forms ; a few
years afterwards (1815), and not knowing the genus of Cuvier, Oken
formed his Themisto, nearly identical with the Polycera of Cuvier.*
The Gufeea of Leach (1852), is entirety congeneric with the genera
of Cuvier and Oken, as is also very likely the Phanerobranchus of
A. Fredol (Moquin-Tandon). The Limacia of 0. Fr. Muller (1781),
contains a whole series of different Nudibranchiata, among them the
D. quadrilineata, and, as first-named species, the D. verrucosa; the
name cannot therefore be employed here.
Although, tlirough Cuvier and Alder (1841), their external char-
acters were somewhat made known, still Polycera, like so many
other Nudibranchiata, remained very superficially known, until the
laro-e monograph of Alder and Hancock, ^ that first really unveiled
their external and internal structure, although Frey and Leuckarf''
had given some anatomical notices of these animals. Lately more
light has been spread over the northern species of the group, through
the investigations of Meyer and Moebius,^ and of G. O. Sars.*
The true Polycera shows a form of body common to the whole
family. The well-developed frontal margin is more or less curved in
* A careful search has failed to find any other ground for supposing that
Cuvier described the genus Polycera in 1812, or at any date before 1817, so
that the 1812 of Hermaunsen is almost certainly merely a misprint. The
name Themisto, of Oken, if congeneric, should therefore take precedence.
— Dall.
1 Alder and Hancock, Monogr. Brit. Nudibr. Moll., Part 2, 1846, fam. 1,
PI. 23 ; Part 4, 1848, fam. 1, PI. 24 ; Part 5, 1851, fam. 1, PI. 22 ; Part 6,
1854, fam. 1, PI. 17 (anat. !) ; Part 7, 1855, PI. 46 supplem. figs. 20, 21.
'^ Frey and Leuckart, Beitr. zur Kenntn. wirbellose Thiere, 1847, p. 66-
70, taf. i, fig. 1§, 13.
3 Meyer and Moebius, Fauna der Kieler Bucht, i, 1865, p. 49-57, m. 2
taf. uud taf. iv. A, B.
* G. 0. Sars, Moll. reg. arct. Norv., 1878, p. 312, 813, Tab. xiv, fig. 14-16.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 101
the middle, with its free margin tuberculated or digitate. The frontal
veil is continued in a more or less tuberculated ridge, that limits the true
back, and posteriorly ends in a single strong or in several smaller
dorsal (branchial) appendices on the outside of and behind the region
of the gill. The true back with longitudinal rows of more or less
developed connected tubercles, sometimes forming low longitudinal
ridges. The number of leaves in the club of the rhinophoria is not
large. The gill is composed of a moderate number (five to seven) of
leaves, which are either simply pinnate or composite (bi- or tripinnate).
The tentacles are small, flattened or auriculate. The jaws or man-
dibular plates in form somewhat recall those of the u^olidiidce, strong,
flattened, sometimes with a peculiar superior process. The rhachis of
the radula naked ; on the pleura? two large hook-formed lateral teeth,
of which the outer is much larger than the inner ; at the outside of
the laterals are four to eight, somewhat flattened uncina^. A large
prostate gives the genital apparatus a particular feature ; the arma-
ture of the penis is of the usual kind.
About the biological relations of Polycera very little is known, as
usual among the Nudibranchiata. The spawn of the most common
northern species is known, and a part of the developmental history
has been investigated by Ray Lankester.'
A small number of species have been described by different authors
in the course of years. Alder and Hancock (Monogr. part T, 1855,
p. 45, XVIII) established and rather well characterized two groups
of Polycera ; according to these authors Gray soon after (Guide I,
1857, p. 213) denominated these groups Polycera (typical) and Palio,
which perhaps might be conserved as subgenera.
L POLYCERA (stricte).
Marge limbi frontalis digitatus. Folia branchialia simpliciter pin-
nata ; appendices dorsales (branchiales) singula? majores.
Lamellae mandibulares processu superiori aliBformi.
1. F. quadrilineata (O. F. Miiller). M. Atlanticum ; Mediterraneum.
2. P. horrida, Hesse. Joum. de Conchyliol.,3 S., XIII, 4, 1873, p. 345. M.
Atlanticum.
' Ray Lankester, Contrib. to the Developm. hist, of Moll., Philos.
Trans., MDCCCLXXV, p. 29, PI. 10, f. 1-9.
Meyer and Moebius have, moreover, given a figure of the shell of the
embryo of their Pol. ocellata (1. c, fig. 10 j.
108 PROCEEDINGS OF THE ACADEMY OF [1880.
3. P. plebeia, Loven. Index Moll., 1846, p. 6.^ M. Atlanticum.
4. P. doriformis (Quatref.). Phanerobrancbe doriforme. Moquin-Tandon
(pseud. A. Fredol) Le monde de la mer., 1864, PI. XII, fig. 1. M.
Mediterraneum.
5. P. eanteriata (Quatref.) Phanerobrancbe a chevrons. Moquin-Tandon
(do) 1. c, pi. XII, f. 2. M. Mediterraneum.
II. PALIO, Gray.
Marge limbi frontalis tuberculatus. Folia branchialia bi- vel tri-
pinnata ; appendices dorsales (branchiales) niinores, complures.
Lamellae mandibulares simplices (sine processu superiori).
6. P. Lessonii (d'Orb.). Pol. ocellata, A. et H. M. Atlanticum.
7. P. pudica, Loven. Ind. Moll., 1846, p. 6. M. Atlanticum.
8. P. pallida. Bgh.. n. sp. M. Pacificum.
9. P. diibia, Sars. Bidr, til Soedyrenes. Naturh., 1829, p. 13. Tab. 3,
fig. 5, 6. Loven, Ind. Moll., 1846, p. 6. M. Atlanticum sept.
10. P. ? Cookii, Angas. Journ. de Conchyl., 3 S., IV, 1, 1864, p. 58 ; PI. V,
f . 6. M. Pacificum.
11. P. ? Capensis, Quoy et Gaim. Voy. de I'Uranie. Zool., 1824, p. 417 ;
PI. 66, f. 4. M. Capense.2
P. pallida, Bgh., n. sp. Plate XV, fig. 14 ; Plate XVI, fig. 1-9.
Color flavescens. Branchia sexfoliata.
Lamellie mandibulares fere ut in FoL Lessonii, sed magis elongatae.
Armatura lingualis fere ut in Pol. Lessonii; denies externi 5.
JIab. Oc. Pacificum septentr.
Of this form Dall dredged a single individual in June, 18t3, at
Kyska Harbor (Aleutians), at the depth of ten fathoms on rocky
bottom. According to Dall, the color of the living animal was " yel-
lowish-white."
The length of the animal preserved in spirits was 7.0 mm., with a
height of 4.0 and a breadth of 3.0 mm. ; the height of the branchial
leaves about 1.0 mm., also that of the rhinophoria ; the breadth of the
1 "Viridifusca, sulpliureo maculata, papillis frontis 10, brancliiali
utrinque una postica majore ; 11 mm. Bohus," Loven.
This, as well as the other new Polycera of Loven, has not since been
seen (Cf. G. O. Sars, Moll. reg. arct. Norv., 1878, p. 313).
2 Of the three (not too naturally represented) "Polycerse " of A. Fredol
(Moquin-Tandon), the one (1. c. PI. XII, fig. 6) seems to be the Pel.
Lessonii, the other two (fig. 3, 4) belong undoubtedly to the genus
Thecacera.
1880.] NATURAL SCIENCES OP PHILADELPHIA. 109
foot 2.0 mm. The color of the animal whitish, that of the rhinophoria
and the branchial leaves more yellow ; the margin of the foot white.
'The form as usual. The head rounded, with a prominence on the
upper lateral part ; the mouth a vertical slit. The margin of the
rhinophor-grooves plain. The stalk of the rhinophoria nearly as high
as the club, cylindrical ; the club rather flattened, with about fifteen
leaves; before the rhinophoria a low transverse frontal veil with
scarcely more than two prominences ; the veil continued backwards as
a rather indistinct prominent line on each side of the smooth rounded
back ; the pericardial region a little prominent ; behind the middle of
the length of the back, the gill with six tripinnate leaves in a slight
curve ; behind them the quite low anal nipple, and towards the right
side the renal pore ; behind the gill a little flattened space with a slight
crest on each side with three papilliB. The sides of the body rather
high. In the region of the anterior angles of the foot the genital
papilla with the everted penis (without its recurved point, 0.75 mm.
high), and below it a folded lamella, the duct of the mucous gland. The
foot rather narrow, of nearly the same breadth ; the rounded anterior
angles somewhat prominent ; a fine furrow in the anterior margin.
The intestines indistinctly appearing through the walls of the body.
The peritoneum colorless, nearly without spicula.
The central nervous system (fig. 1) very depressed; the cerebral
ganglia of rounded-triangular form, a little larger than the more
rounded visceral (fig. la); the pedal ones more pyriforra, a little
larger than the last ; the (proximal) olfactory ganglia bulbifonn, not
quite as large as the buccal ones, which were (fig. lb) of rounded form,
connected by a not very short commissure ; the gastro-oesophageal
ganglia of about one-eighth of the size of the former, rounded.^ The
three inferior ( subcerebral, visceral, and pedal) commissures (or at
least the visceral one) free.
The eyes (fig. 1) short-stalked, with black pigment and pale yellowish
lens. The otocysts (fig. 1 ) in their usual place, very short-stalked,
with about eighty otokonia of the ordinary kind. In the stalk of the
rhinophoria some scattered yellowish thick spicula, of the same kind
as in the skin of the back ; none, on the contrary, in the leaves of the
club. In the skin some scattered, yellowish, thick, straight or curved
spicula, mostly of about 0.15-0.3 mm. in length, and of the usual
form. In the interstitial tissue very few larger spicula.
^ In the other species of Polycera I have examined, I never saw gastro-
CBSophageal ganglia, nor any in Ewplocamui or iu Plocamopherus .
no PROCEEDINGS OF THE ACADEMY OF [1880
The oral tube whitish, of about 1.0 mm. length, wide. The bulbus
pharyngeus clear brownish-yellow, somewhat pyriform, with oblique
flattened posterior end, in length about 1.6, by a height of nearly 1.3,
and a breadth of 1.5 mm. ; the sheath of the radula a little prominent
downwards, and to the left from the hindermost part of the under side of
the bulbus. The lip-disk clothed with a brownish-yellow cuticula, that
is continued into the two mandibular plates behind the lip-disk at the
entrance of the oral cavity, the form of the mandible could not be
determined with certainty ; a yellowish cuticula clothes the rest of the
cavity. The tongue with ten rows of plates, further backwards six
developed and two younger rows ; the total number eighteen.^ The
rhachis (fig. 2) not very narrow. The plates yellow. The length of
the first plate about 0.11, of the second 0.20, of the inmost of the ex-
ternal plates 0.14, of the following 0.12, 0.10, 0 08 and 0.06 mm. (all
from the hinder part of the sheath). The first lateral plate (fig. '■laa^
5, 6) formed somewhat as in the P. Lessouii, the hook still smaller ;
the second of the same form, but larger (fig. 2&i, 3), the hooks much
larger, especially the anterior, which is broader and excavated (fig. T).
More outwards five external plates (fig. 2cc), all with a crest, which is
larger in the two innermost ; adjoining the outermost of these plates
several longitudinal folds of the lingual cuticula, which sometimes
simulate one to two plates more (fig. 2).
The salivary glands whitish, elongate. The oesophagus rather wide,
the stomach inclosed in the liver. The intestine appearing at the
middle of the length of the liver a little to the left, at the bottom of a
deep and large cavity in the upper side of the liver ; the pyloric part
' According to Alder and Hancock (Monog. Part VII, 1855, PI, 41 sup-
plement, fig. 20, 21), the number of rows was fifteen in the Polycera qnad-
rilineata, sixteen in tlie P. ocellata. thirteen in the P. Lessonii ; Alder and
Hancock saw (1. c.) four external plates in the Pol. quadrilineata, five in
P. ocellata, and six in P. Lessonii. Meyer and Moebius saw five to seven
external plates in their Polycera ocellata, whilst the number of rows (1. c.
PI. 50) is noted as thirteen to fifteen ; in the P. quadrilineata they found
four to five external plates and twelve to thii-teen rows. In four specimens
of Pol. quadrilineata I saw six to eight rows on the tongue, more back-
wards six to seven developed, and one not quite developed row ; the total
number of rows was fourteen to fifteen. In all specimens there were but
four external plates. In four specimens of Pol. Lessonii I saw nine to ten
rows on the tongue, more backwards eight to seven or five developed,
and a single not developed row; the total number of rows was sixteen to
eighteen. In all the specimens there were eight external plates.
1880.] ' NATURAL SCIENCES OF PHILADELPHIA. Ill
of the intestine rather wide, its curve reaching to the bulbus pharyn-
geus. The liver about 5.0 mm. long by a breadth of 3.5 and a height
of 3.25 mm. ; the form conical, the posterior end rounded, the anterior
much broader, flattened and adjoining another flattening on the infe-
rior part of the right side of the organ ; the color was yellowish.
The sanguineous gland of quadrangular form, of a diameter of about
1.5 mm., whitish.
The hermaphroditic gland with its yellowish-white lobes covering
nearly the whole surface of the liver : in the lobes large oJgene cells.
The anterior genital mass of the length of about 4.0 mm. by a height
of 3.0 and a breadth of 1.5 mm. The ampulla of the hermaphroditic
dact resting on the inferior margin of the genital mass, whitish,
straight, of the length of 3.0 mm by a diameter of about 0.5 mm.
At the anterior end of the ampulla a flattened body (prostate) that
freely projects before the anterior margin of the rest of the genital
mass ; it was of about the same length as the ampulla, but nearly twice
as broad ; the cavity of the organ rather large and the walls rather
thin. The prostate slopes gradually into the thin but strong spermato-
duct, which is about 6.0 mm. long and terminates in the penis, which
was short, conical (fig. 8«, 9), about 0.75 mm. long, and terminated
in a somewhat flexible, yellowish glans (fig. 8, 9, 14), of the length of
about 0.37 mm. by a diameter at the base of about 0.09, and at the
point of 0.037 mm.; through the largest part of its length it was
covered with (in all about twelve) series of small chitinized crests,
which did not surpass the height of about 0.0025 mm. (fig. 14 i; the
armature only continued through a short part of the interior of the
spermatoduct. The spermatotheca spherical ; the spermatocysta pyri-
form, filled with sperma. The cordate mucous gland whitish and
yellowish-white (fig. 8b).
This species approaches to the Pol. Lessonn, but seems even dif-
ferent in color from that and the other Atlantic forms, and also differs
in the slight development of the frontal veil and of the lateral crests of
the back, as well as in the number of the external plates of tongue,
and' in the nature of the armature of the penis.
* The armature of the penis of Polyc. quadrilineata (hitherto the only
species in which an armature has been described) as figured by Friele and
Hansen (1. c. Tab. II, fig. 3) is very different from that of the Pacific spe-
cies, and that difference has been confirmed by my examination of typical
specimens.
112 PROCEEDINGS OF THE ACADEMY OF [1880.
TEIOPHA, Bergh, n. gen.
Forma corporis fere ut in Triopis, sicut quoque margo fronta-
lis ; margo dorsalis appendicibus nonnullis nodosis vel breve
ramosis. Tentacula eompresso-poculiformes (auriformia) ; rhino-
phoria retractilia, clavo perfoliato. Branchia quinqnefoliata,
foliis tripinnatis.
Os lamellis dnabus fortioribus e baculis rainutis eompositis
armatum. Lingua rbachide dentibus spuriis (4); pleuris denti-
bus lateralibus 3-4 (corpore processu alaiformi et hamo ap-
planato instractis) et serie dentium externorum (10-11) armatis.
Prostata ?
This interesting form, that forms a link between Polycera
and Triopa on one side, and the Euplocami on the other, ap-
proaches more nearly to the latter than to the former.
In the exterior, the Trioph(E^ resemble the Trio'pce^ but still
differ in some points sufflcienth'. The appendices of the back
are more composite ; the tentacles seem different from those of
the Triop<e (which have them folded lengthwise and obtuse at
the end; see for comparison, PL XV, fig. 12); they are com-
pressed cup-shaped or auriculate. The gill contains five leaves.
Whilst the Triojoce want an armature of the true mouth, ^ the
Triojphce are provided with two strong plates (composed of
densely set staffs). Whilst the rhachis of the tongue in the
TriopcB is naked, the Trio2:)hce show four false plates, ("bosses" of
Dall, simple thickenings of the base membrane of the radula),
here ; instead of the two peculiarly formed lateral plates on the
pleural in the Triopa^ the Triophm have three or four lateral
plates (with a wing-like process of the bod}^ and a depressed
hook) ; with, on the outside of these, a series of (ten to eleven)
uncinal plates, nearly as in the Triopce. After all, the Triophce
are closely allied to the Colgce^* and essentiall}^ differ from these
^ Having at first and rather superficially examined the exterior, I first
regarded the animal as a Triopa, and called it so [s. part I, p. 138 (73),
and the Plates (XIV, XV)].
^ See for comparison PI. XIII, fig. 19.
^ See for comparison PI. XIV, fig. 31, 22.
* The diagnosis of the Colgce would be :
Forma corporis fere ut in Triopis. Vaginae rhinophoriales calyciformes
obliquse ; rhinophoria retractilia, clavo perfoliato. Tentacula auriformia.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 113
only in the armature of the tongue, which in the Oolgce exhibits
only a single series of (false) rhachidian plates and (on each
side) two lateral plates in form approaching those of Polycei-a.
The nature of the prostate is unknown ; the armature of the
penis not differing much from that ordinary in the large group of
the PohjceratidcB.
Although somewhat approaching to the Euplocami in the form
of the appendices of the back, in the armature of the true mouth
and of tlie pleura of the tongue, the Triophce still cntirel}^ differ
in the form of the tentacles, in the number of the branchial
leaves and very likely in the nature of the prostate.
The Trioph(E have hitherto been only found in the Pacific
Ocean.
1. Tr. modesta, Bgh. n. sp. Oc. Pacificum.
2. Tr. Carpenteri, Stearns. Proc. of the Cal. Acad, of Sci., April 7, 1873,
p. 2, fig. 2. Oc . Pacificum (California).
Tr. modesta, Bgh. n. sp. PI. XIV. fig. 17-20; PI. XV, fig. 1-10.
f Triopa Carpenteri, Stearns. L c. p. 2, fig. 2,
Color e flavido albescens. Appendices dorsales paucpe ; folia
branchialia 5.
Hah. Oc. Pacif. septentr.
Of this form Dall has obtained a single individual at Yukon
Harbor (Shumagins), in August, 1874, at a depth of six to
twenty fathoms, on a bottom of sand and stones. The color of
the living animal was, according to Dall, " yellowish-white."
The animal preserved in spirits was of whitish color ; the
dorsal appendices, the gill and the rhinophoria more yellowish.
The length of the animal 16.0 mm., b}^ a height of 7.0 and a
Dorsum papilligerum, prsesertim margo frontalis et dorsalis. Branchia
pauci (4-5 I foliata,
Mandibulse triangulares, fortes. Radula fere ut in Polyceratis, dentibus
lateralibus (3) et externis (7), sed prseterea dentibus medianis (spuriis)
instinicta.
Mei-ely one species of the genus is yet known, one of the first described
Nudibranchiata, the Doris lacera of Abildgaard (Zool. Dan., IV, 1806,
p. 23, Tab. CXXXVIII, fig. 3, 4), which has been found too on the coast
of America (Cf. Verrill, notice of recent addit. to the Mar. Fauna of
North Am., XXXVIII. Amer. Jour, of Sc. and Arts, XVI, 1878, p. 211).
114 PROCEEDINGS OF THE ACADEMY OP [1880.
breadth of 5.5 mm. ; the height of the branchial leaves 1.25, of
the rhinophoria 2.0 mm. ; the breadth of the foot 3.5 mm.
The form as usual. The head flattened in front, semilunar;
the tentacles compressed-cup-shaped, rather short (about 1.0 mm.
long), truncated at the end, longitudinall,y folded and open at
the outer side. The frontal margin not projecting much, with
many smaller and larger short digitations and crenulatious ; in
front in the median line were two small conical papillse before
the region of the rhinophoria. The margin of the rhinophor-
holes somcAvhat projecting, smooth : the (deeply retracted) rhino-
phoria with rather short stalk ; the club with thirty-five to forty
rather broad and thin leaves.
The back rounded over from side to side, without certain limits
l)etween it and the sides of the body. At the lateral parts
of the back (on each side) five appendices ; the first standing a
little behind the end of the frontal margin; the next about in
the middle of the space between the first and third ; this last a
little before the region of the gill ; farther backwards were also
two similar ones. The appendices were club-shaped, with simple
or composite nodosities spread upon their bodies, and especially
at their bases ; the third was the largest, reaching the height of
about 2.5 mm.; all the others a little smaller, and all of about
the same size. Much smaller, conical or club-formed simple
pfipilloB were scantily and irregularly scattered on the back. The
gill consisting of five strong, tripinnate, quite separate leaves, a
single anterior and two lateral pairs. The anal nipjDle nearly
in the centre of the posteriorly open branchial circle, a blunted,
nearly cylindrical prominence, about 0.5 mm. in height ; at its
base on the right side and a little forwards was the very distinct
renal pore. The sides of the body rather high and smooth ; the
genital bpening a short longitudinal slit lying rather forwards,
with two openings at its bottom. The foot not very narrow, of
nearly the same breadth throughout its whole length ; the anterior
border emarginated in the middle, with a fine line.
The intestines did not shine through the integuments. The
peritoneum was colorless, without spieula.
, The central nervous system (PI. XV, fig. 1) flattened ; the
cerebro-visceral ganglia (fig. la) reniforra, a little narrower at
the fore-end ; the pedal ones (fig. Ibb) rounded, scarcely larger
than the visceral ; the large commissure (fig. 1) as usual ; small
1880.J NATURAL SCIENCES OF PHILADELPHIA. 115
optic ganglia (fig. 1). The proximal olfactoiy ganglia (fig. Ic)
bulbiforra, the n. olfactorii not very long; the distal olfactory
ganglia inverse pyriform. The buccal ganglia (fig. Idd) ovoid,
connected nearly without commissure ; the gastro-oesophageal
ganglia small (fig. le), with one large cell.
The eyes (fig. 1) with coal-black pigment and yellow lens.^
The otocysts at the usual place on the under side of the cerebro-
visceral ganglionic mass, crowded with otokonia of the usual
kind (fig. 1). In the leaves of the rhinophoria no spicula ; in
the axes and in the stalk, on the contrary, spicula of the same
kind as in the skin or often larger. The skin with few and
small spicula and calcified rounded cells, here and there lying in
groups. The marginal dorsal appendices covered all over with
above-mentioned nodosities ; at their points perhaps a similar
(but empty) bag as in the typical species (Cf. PI. XIII, fig. 16, 17).
The anal tube large, 3.0 mm. long. The bulbus pharyngeus
strong, of the length of 4.0 by a height of 3.0 and a breadth of
3.3 mm. ; the radula-sheath projecting about 1.0 mm. from the
hinder part of the under side of the bulbus. The lip-disk rather
convex, with vertical oral slit (PI. XY, fig. 2), clothed with a
pale 3'ellow cuticula, that behind the oral slit on each side is con-
tinued in a triangular, brownish-yellow lip-plate of a greatest
breadth of 1.0 ram (fig. 3), narrow at the inferior end, broader
at the supei'ior, and composed of simple, somewhat curved, erect
stag's (fig. 4, 5) about 0.18 mm. in height (fig. 4). The tongue
broad ; in the amber-yellow radula, thirteen rows of plates,
further backwards in the sheath, six developed and two younger
rows ; the total number thus twenty-one. The three foremost
rows of the tongue very incomplete, reduced to the outermost
(four to five, six to seven, nine to eleven) uncinal plates. The
rhachis rather broad, bearing two quadrangular thickenings of
the cuticula (PI. XV, fig. 6a) of the length of about 0.18-0.2
mm.-, more thickened and jellowish in the anterior margin, otlier-
wise colorless. At the outer side of these median plates is a
somewhat shorter and narrower plate (fig. 6&6), of yellowish
color ; in the posterior rows (PI. XIV, fig. 20) much broader.
The three succeeding plates brownish-j-ellow, hook-shaped, all
nearly of the same form and of the same but outwardly slowly
' Alder and Hancock (1. c. part VI) also saw small optic gauglia in the
Triopa clavigera.
116 PROCEEDINGS OP THE ACADEMY OF [1880.
decreasing size (PI. XY, fig. Gcd) ; the fourth lateral plate, on
the tongue especially, with a small hook (fig. 7a) that is more
developed backwards, and in the four youngest rows is developed
quite (PI. XI Y, fig. IT) as in the three plates mentioned. On the
lateral parts of the pleuroe ten to eleven external (uncinal) plates,
the four to five interior (fig. 7, 8a6, 10 ; llbc) with a more devel-
oped crest, the rest (fig. lb) narrower.
The salivary glands (PI. XY, fig. 11a) nearly as long as the
duct (fig. 116) ; both together about 5. ,5 mm. long, descending
along the whole back side of the bulbus pharyngeus ; the gland
whitish, smooth.
The oesophagus rather long (6.5 mm.), and Avide especially
in the posterior part (diameter 2.0 mm.), entering into the inferior
part of the liver ; with rather strong and numerous folds ; the
contents (as in the intestine) spongiary masses and different Ba-
diolarise of a diameter of 0.09 mm. The intestine issuinaf from the
liver a little before the middle of the upper side of this organ ;
the anteriorly proceeding part reaching the anterior margin of
the liver and about 4.5 mm. long by a diameter of 1.5 mm. ; the
retrocessive part Y.O mm. long by a diameter of 0.75 mm. The
liver divided b}'^ a deep furrow from the right margin into two
halves of nearly equal size; 6.0 mm. long by a breadth of 3.75
and a height of 3.4 mm. ; the posterior extremity rounded ; the
anterior half of the inferior side obliquely flattened ; the color
yellowish-gray ; the cavity of the interior rather small.
The pericardium of oval form, large, having the length of
3.5 mm. The sanguineous gland whitish, of the length of 2.5 mm.
by a breadth (at the anterior end) of 2.5 mm. The renal syrinx
short-pyriform ; the tube of the organ strong.
The hermaphroditic gland not much developed, paler than
the liver, with large oogene cells. The anterior genital mass
small, about 1.5 mm. long by a height of 0.75 and a breadth of
about 0.5 mm. The ampulla of the hermaphroditic duct yellow-
ish, rather long, forming corkscrew-like windings. The sperma-
toduct not long, passing into the short penis. This, with its
armature of very minute hooks, the spermatotheca, the spermato-
cysta and the vagina, as far as could be determined, as in the
typical Triopa} The gland whitish.
' See for comparison, PI. XY, fig. 13.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 117
This species ma}' perhaps be the Triopa Garpenteri of Stearns ;
it has, like that, five branchial leaves, and does not differ much in
the number of the dorsal appendices (six) or the form of the
frontal margin ; but the dorsal nodosities of the last species are
orange-colored, and the rhinophoria, the dorsal appendices, and
the branchial leaves tipped with orange. Through the great
kindness of Mr. Dall I have seen a drawing of the animal of
Stearns, from specimens secured after those he had described,
but they do not give more details than the original description ;
and Stearns seems not to possess the original specimens, which
very likely are lost forever. On the other side, it must be remem-
bered that Sars (Beretn. om en i Sommeren, 1849, forctagen
zoolog. Reise i Lofoten og Finmarken, 1851, p. T4) found "the
young individuals of Triopa lacera (M.) entirely white, also on
the tentacles and gills, merely the liver shines brownish through
the skin."
EXPLANATION OF THE PLATES.
An asterisk denotes that the drawing is by camera lucida, the
fraction denotes the magnification.
The serial numbers of the plates (Part I, plates i-viii, Part II,
plates ix-xvi) are solel}- referred to throughout the text. As
Part II appears in another volume of the Proceedings of the
Academy, the plates of Part II have been for that reason renum-
bered with a second set of numbers, Plate ix being Plate i, Plate x
being Plate ii, etc., in the new volume. The serial numbers re-
ferred to in the text, follow the new numbers for Part II in
parentheses throughout this explanation.
Plate I (IX).
Jorunna Johnstoni (A. and H.),
1. a, stalk of the (6) gangl. genitale; c, gangl. genit. secunda-
rium,* ^^.
2. Granules of the back, stiffened by spicula,* ^f^.
3. Part of the middle of the radula, with the two innermost
lateral plates ; a, rhachis,* ^f ^.
i. The hook of a plate from the back,* ^s.,
118 PROCEEDINGS OP THE ACADEMY OF [1880-
5. Outer part of two series of plates with 8 plates,* ^f^; aa,
outermost.
6. Outer part of another series with 3 plates,* ^^.
7. a-6, vagina; c, gland, hastatoria; d, opening of the bag of
the spur; e, spermatoduct ; /, penis,* -\'-.
8,9. Spermatotheca ; c, its chief duct; d^ gland, hastatoria ; h.
spermatocysta ; e, duct to the mucous gland,* -\^-.
10. a, Duct of the gland, hastatoria; b, the bag of the spur; c,
opening of the bag,* ^^-.
11. a, spermatoduct; &, opening of the bag at the bottom of the
penis ; in the interior a dart (?),* ^4--
Adalaria p7'0xima (A. and H.).
12. Tubercles of the back.
13. A part of the rhachis from above; a, median plates; &6, large
lateral plates,* ^^.
14. Part of the radula, obliquely, from the side , the hooks of the
large lateral plates of both sides,* ^f^.
15. Two series of (9) external plates ; a, the innermost ; b, the
outermost,* ^f^.
Adalaria albopapillosa (Dall).
16. Part of the surface of a tubercle of the back,* ^^.
Adalaria pacijica, Bergh.
17. a, median plate ; 6, large lateral plates from the side,* ^^.
" Lamellidoris muricata (0. Fr. Miiller).
1 8. The vesica fellea ; a, its duct.
Plate II (X).
Adalaria pacifica^ Bergh.
1. Median pseudo-plate (or boss), from the upper side,* ^^^.
2. 2. Part of the radula, with series of (5-7) lateral plates ; a-a,
1-2 complete rows of (15) external plates, and 1-2 incom-
plete rows ; 66, innermost plates of the row ; ce, outer-
most,* -f-.
3. Outer part of a row with 9 erect plates ; a, innermost,* ^^^.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 119
Adalaria virescens, Bergh.
4. a, esophagus, with its dilatation ; b, salivary gland ; e, its
duct.
5. Ganglion penis,"^ ^j-.
Adalai'ia Loveni (A. and H.).
6. Median part of the radula from above, with (aa) large lateral
plates ; 66, innermost part of two rows of external plates,
with 1-5 plates,* ^^.
1. Large lateral plate, from the side,* ^^^.
8. Piece of the left part of the radula;* ^f^ a, two median
pseudo-plates or bosses ; 6, large lateral plates ; c, two in-
complete rows, with 6-7 plates.
Adalaria albopapillosa (Dall).
9. a, (2) median pseudo-plates ; 66, (2-3) large lateral plates of
both sides,* ^K
10. a, (3) median pseudo-plates; 66, (2-4) large lateral plates of
both sides ; c, innermost part of three (right) rows of ex-
ternal plates, with 3-4 plates ; (Z, (left) row of 7 external
plates,* ^K
11. Four outermost plates of a row ; a, outermost,* ^-f-.
Acanthodoris pilosa (0. Fr. Mtiller).
12. End of the everted penis ; a, opening,* ^f^.
13. Epithelium of the vagina,* -f^. ,
Acanthodoris pilosa^ var. albescens (Pacijica).
14. a, anterior margin of the foot ; 6, edge of the tentacle.
15. Ganglion genitale from the penis,* if^.
Plate III (XI).
Acanthodoris pilosa (Miiller).
1. Three external plates; a, outermost,* ^f^.
Acanthodoris pilosa, var. albescens.
2. The genital opening with its everted margin ; a, the two fore-
most apertures.
120 PROCEEDINGS OF THE ACADEMY OF [1880.
Lamellidoris hilamellata (L.) var. Pacifica.
3. Part of tlie branchial area with (aa) some branchial leaves ; 66,
some of the larger surrounding tubercles. In the centre
the anal nipple, the renal pore and interbranchial tubercles.
4. The Slicking crop, from the edge.
5. The half of the same, from the inside ; a, stalk.
6. a, spermafotheca ; 6, spermatocysta ; c, duct of the last ; d,
duct to the mucous gland ; e, vagina.
7. a, two median pseudo-plates ; 6, a latei'al plate ; cc, three
external plates,* -^-f^.
8. External plate from the side,* ^f^.
9. Two of the foremost lateral plates with blunted end,* -^-|^.
Lamellidoris muricata (Miiller).
10. a, Median pseudo-plate shining through the left of the lateral
plates, 66 ; c, three external plates,* ^f ^.
11. aa, Basal edge of three lateral plates ; 6, external plates,* ^4^.
12. a, Glans penis ; 66, praeputium ; e, spermatoduct,* J^f^.
Lamellidoris varians, Bergh.
13. Lateral plate from the side,* ^f^.
14. Median pseudo-plate, from above,* -^f^.
Adalaria Pacijica, Bergh.
15. Innermost part of two rows of external plates,* -f-; a, two
innermost; 6, the third failing (in the anterior row); c,
eighth.
Plate IV (XII).
Acanthodoris pilosa (0. F. Miiller), var. purpurea.
1. Labial disk, with (a) the lancet-formed blades projecting in
the lowest part of the mouth proper.
2. The lancet-formed blades (a) with the adjoining part (6) of
the armature of the mouth,* J-^^.
3. a. The right lancet-formed blade ; 6, the adjoining part of the
armature,* ^^^.
4. Elements of the armature,* ^^^.
5. Lateral plate, from the side,* -^f^.
1880. J NATURAL SCIENCES OF PHILADELPHIA. 121
<;. The hook of a pltite, from the side.* ^f^.
7. Salivary gland ; a, duct ; 6, posterior end.
8. a, pa7's pylorica intestini ; b, vesica fellea; c, intestinum
descendens.
9. Part of the vas deferens^ with its stricture,* J-?-^-.
Acanthodoris pilosa (M.) rar-. hrunnea albopapillosa.
10. ab, Lancet-formed blades from the under side,* i?^.
11. ff, Part of left; 6, of right lancet-formed blade; c, adjoining
part of the armature of the mouth,* ^f^.
12. aa, Upper part of three lateral plates ; bb^ two series of exter-
nal plates ; from the sheath of the radula,* ^f ^.
Acanthodoris pilosa (M.) var. albescens.
13. Elements of the armature of the mouth,* ^f^.
14. Isolated element,* ^f^.
1.^. Upper part of a lateral plate, from the outside,* ^\-.
1 6. Upper part of a lateral plate, from the inside,* ^f ^.
Plate V (XIII).
Lamellidoris varians, Bergh.
1. The central nervous system, obliquely, from the under side.
* 5j5 . d^ ganglia cerebro-visceralia; bb, ganglia pedalia; c,
gangl. penis and gangl. genitale; d, ganglia buccalia; ee^
ganglia gastro-cesophagalia. The eyes and the otocysts
visible.
Acanthodoris pilosa (M.), var. albescens.
2. The bulbus phar^-ngeus, from the side ; a, cuticula and the
lancet-formed blades; 66, mm. retractores bulbi ; c, the
sucking-crop ; c?, salivary gland, above this the right buccal
and gastro-oesophageal ganglion ; e, the sheath of the radula
/, the crop of the oesophagus ; g, continuation of the oeso-
phagus.
3. Lateral plates, from the outside,* ---^-fi.
4. Partof the armature of the spermatod net, with its hooks,* -^(^ .
9
122 PROCEEDINGS OF THE ACADEMY OF [1880.
Acanthodoris pilosa (M.).
5. a, spermatotheca ; b, spermatocysta ; c, duct to the mucous
gland ; dd, duct to the vagina,
Acanthodoris csei'ulescens, Bergh.
6. Part of the armature of the mouth,* ^f^.
7. External plates, from the side ;* -^^f^ a, innermost.
Chromodoris Dalli^ Bergh.
8. The upper part of a branchial l^af,* J-^-^-.
9. Part of the lip-plate, from above,* ^f^.
10. Elements of the lip-plate,* ^K
11. Part of the rhachis, with three (bosses or) false plates,* ^p.
12. a, false plate, obliquely, from the side,* -^f^.
13. The 13th plate, from the side,* -if^.
14. The 9th plate, from the side,* ^^.
Triopa clavigera (0. Fr. Miiller).
15. Tubercles of the back.
16. Vertical section of one of the appendices of the back ; a, bag
at the point.
n. Elements of this last bag.
18. Spicula of the skin.*
1 9. Lowest part of the mouth, with its cuticula ; a, the free
margin,* ^f^.
20. Hindermost part of the bulbus ; a, tongue; 6, sheath of the
radula.
/
Plate YI (XIV).
Chromodoris Dalli^ Bergh.
1. The buccal (a) and gastro-oesophageal (b) ganglia,* ^s.,
2. Part of the median portion of the radula ; a, false plates, on
each side the 2-3 innermost (lateral) plates,* ^^.
3. Outer part of two series of plates with 11 plates; a, outer-
most; b, eighteenth,* ^f^.
4. a, spermatotheca ; b, spermatocysta; c, duct to the vagina;
d, duct to the mucous gland,* ^^-.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 123
Ghromodoris Calif orniensis, Bergh.
5. Hinder part of the body, from the under side, with 6 knot*
on the mantle-margin ; a, foot,* ^f ^.
6. Upper median part of the true mouth,* ^|^.
7. Part of 4 series of hooks of the lip-plate, from above,* ^f^..
8-10. Elements of the same, in different positions,* ^^^.
11. Three innermost plates ; a, the first,* ^f^.
12. One of the largest plates,* ^^^.
13. Hook of 3 larger plates, obliquely, from the foreside,* ^f^.
14. Four outermost plates; a, outermost,* ^f^.
15. Two irregular outermost , plates ; a, outermost,* ^f^.
Acanthodoris cxj-ulescens, Berg:h.
16. Series of plates; a, two lateral plates; 6, the outermost of
the external plates,* ^^^.
Triopha modesta, Bergh.
17. Part of one of the hindermost series of plates (in the slieath),
with (a) 4 lateral plates and {b, c) 2 external plates,* ---f^.
18. o, second and &, third large lateral plates, from above and
from the back,* ^K
19. a, fourth ; h, fifth plate (as in fig. 18 from the tongue),* -^^.
20. Outer false plate of the rhachis (from the sheath),* 3,\^.
Tr-iopa clavigej^a (M.).
21. a, second lateral plate; 6, two external plates,* ^f^.
22. First lateral plate,* ^|^.
Plate YII (XV).
Triopha modest a, Bgh.
1. Central nervous system,* ^\4 . q^ ganglia cerebro-visceralia ;
bb, pedal ganglia ; e, ganglia olfactoria proximalia ; dd^
buccal ganglia ; e, gangl. gastro-wsophagal.
2. The labial disk with the true mouth.
8. Upper commissure of the lip-plates,* -Y-.
4. Elements of the lip-plate,* ^K
.'). Upper ends of two elements,* ^f ^.
124 PROCEEDINGS OF THE ACADEMY OF [1880-
6. Median part of a series of the teeth ; a, (false) median plates
of the rhachis; 66, external plate of the same; ce, first
lateral plate ; c?, third lateral plate,* ^K
1. Continuation of the former; a, fourth plate; 6, outermost
plate,* ^^.
8. Four (inner) uncinal plates ; a, the second ; b, the fifth,* ^^.
0. First lateral plate,* ^f^.
1 0. Seventh and eighth external plates,* -^-|^.
11. Salivary gland; a, gland ; b, duct,* -Y-
Triopa clamgera (M.).
12. Tentacle.
1 3. Part of the armature of the penis.* ^f ^.
Polycera pallida, Bergh.
14. The glans penis,* -^-|^.
Plate VIII (XVI).
Polycera pallida^ Bergh.
1. Central nervous system, from the upper side,* Y- ; aa, visceral
ganglia; 6, ganglia buccalia and gastro-oesojyhagalia.
'J. Part of the radula with two rows ; aa, interior ; 66, exterior
lateral plates ; cc, uncinal plates,* -^-f
3. Exterior lateral plate, from the side,* ^^.
4. Under side of the two lateral plates:* aa and 6, as in fig, 2.
* 35 0
1
* 35 0
5. First lateral plate, from the side,* ^4'
H. The same, from above,* 3.^^.
7. Hook of the second lateral plate,* ^^.
8. Genital papilla and everted penis with its glans ; 6, prominent
fold of the duct of the mucous gland.
0. Glans of the penis, with the end of (6) the spermatoduct,*
^f ^ ; a, point of the glans.
Archidoris Monfereyensis (Cooper).
10. Large lateral plate, from the side,* ^f-^.
JI. Outer part of two series of plates with 4 plates; aa, outer-
most,* -y-.
1.880.] NATURAL SCIENCES Of PHILADELPHIA. 125
Aphelodoris Antillensis, Bergh.
(Cf. Malakozoolog. Blatter, N, S., i, 1879, p. 107-113).
12. a, ganglia huccalia^ with 6, ganglia gastr-o-ossophagalia ; c.
secondary ganglion,
* 200
1 3. Median part of two series of plates ; aa, innermost ; 66, second
plates,* ^^.
14. A large lateral plate,* ^f-.
15. Outermost double plates of two series,* ^|^.
16. Outer part of two series with two plates ; aa, outermost,* ^^•-
11 7. The sixth plate from the outer margin of the radula,* ^f ^.
! 8. Outer part of three series with 3 plates ; a, outermost,* ^^.
Polycera Holbolli (Moll.).
1 9 The genital papillae, from the front.
20 The same, from the side.
21. First lateral plate, from above,* ^y^.
January. 1880.
ERRATA FOR PART I.
On account of the inability of the author to read the proofs,
and from certain obscurities in the manuscript, some errors crept
into the first part of this paper, and the arrangement of the para-
graphs was somewhat confused by the printer.
The delicacy and beauty of the plates in their original state,
liaving been destroyed by the printer, the present ones have been
steel-surfaced, to avoid, if possible, a similar misfortune.
The specific name Calif orniensis ( Ghromodoris) was substituted
in the printed text for Calensis^ which appeared on the plate and
ill the manuscript under the idea that the latter was intended
merely as an abbreviation.
The following list of errata has been received from the author ;
it is believed that the present concluding part of the paper is
much less in need of such corrections.
Page 128 ( 72), line 15 : for Triopa modesta, B., read Triopha viodesta, B.
" 129 ( 73), line 22 : for mandibulse read . Mandibulte.
'' 130 ( 74), line 2 : for genus read penis.
•' 132 ( 76), line 30 : a comma to be put before the parenthesis, and
the comma after the parenthesis to be cancelled.
126 PROCEEDINGS OF THE ACADEMY OF [1880.
Page 135 ( 79), line 11 : for dentibus medianis denticulati read dentibus
medianis denticulatis.
" 135 ( 79), line 18 : for caducous read not caducous.
^' 135 ( 79', line 19 : a semicolon is needed before "the foot."
'^ 136 ( 8^*), line 5 : the comma after "laterales " to be cancelled.
" 136 ( 80), line 17 : a comma is needed after " 1, 5 "; the comma after
" rhinophoria " to be cancelled.
"■ 138 ( 82), line 5 : for Plate I, fig. 9, read PI I. fig. 9-12.
" 140 ( 84), line 39 : for (fig. 11, one to fourj read fpl. I. f. 11 ; pi. II,
f. 1-4).
" 141 ( 8-i), line 1 : for The intestines are read The intestine is.
" 141 ( 85), line 3 : for anal papillae read anal papilla.
" 141 ( 85), line 34 : for 2 w. pi. read w. 2 pi.
" 141 ( 85), line 35 : for 2te Heft read 2tes Heft.
" 141 ( 85), line 41 : for ab read ob.
" 142 ( 86^ line 6 : /or denticalis rert(i denticulis.
" 144 ( 88), line 16 : for M. retractoris read M. retractor.
" 145 ( 89), line ^ : for Z R. J. read 3 R. I.
" 145 ( 89), line 27 : for Dentes medianse read D. mediani.
" 145 ( 89), line 27 : for altamen read attamen.
" 146 ( 90), line 22 : for mantle read muzzle.
" 147 ( 91), line 11 : for anal read oral.
* 150 ( 94), line 4 : for Animal read Color animalis.
" 150 ( 94), line 3 : before Dendron. Dalli, B., insert "2."
' ' 152 ( 96), line 27 : for side, the read side. The.
" 153 ( 97 S line 17 : for Dalzell read Dalyell.
" 153 ( 97). line 27 : for Tr. glaucse read Tr. glamae.
' ' 154 ( 98 \ line 15 : for cucculata read cucuUata.
" 154 ( 98), line 19 : for Duvancelia read Duvaucelia.
" 155 ( 90), line 8 : for of the papillae read of the papilla.
" 156 (100), line 11 : for is contracted read was contracted.
" 156 (100), line 16 : /(??• The larger mucous gland read The larger
opening of the mucous gland.
" 156 (100), line 19 : for before which read , below which.
" 156 (100), line 38 : for in the hinder part read between the binder
parts.
" 159 (103), line 20 : for The cardia were wide, etc., read the cavity
was, etc.
" 159 (103), line 26 : for but backward at the front and end read bent
backward at the frontal end.
" 160 (104\ line 1 : for Fig. 65 a rend 15 a.
" 161 (105), line 33 : for bulbus, and read bulbus, or.
" 161 (105 \ line 38 : for Beitr. read Bidr.
'* 162 (106), line 17 : for dentates read dentatis.
" 163 (107), line 33 : for leaves 80 read leaves 8.
" 163 (107), line 9 : for Fig. 6, 7, read Fig. 10, 11.
«• 165 (109), line 25 : for Fig. 1-7 read Fig." 8-14.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 12Y
Page 166 (110), line 19 : for Fig. 1 read Fig. 8.
167 (111), line 4 : for Fig. 2 read Fig. 9.
167 (ill), line 6 : for Fig. 3 read Fig. 10.
167 (111), line 15 : for Fig. 4 read Fig. 11.
167 (111), line 10 : for Fig. 1 a read Fig. 2 a.
167 (111), line 16 : for Fig. 5 read Fig, 12.
167 (111), line 19 : for Fig. 4, 5, read Fig. 11, 12.
167 (111), line 23 : for Fig. 6, 7, 8, read Fig. 13, 14, 3 b.
168 (112), line 5 : for Plate XII read PI. XIV.
168 (112), line 6 : for punctus read punctis.
170 (114), line 5 : for Fig. 13 read Fig. 15.
170 (114), line 24 : for latiuni read latum.
170 (114), line 26 : for minutissimus read minutissimis.
170 (114), line 33 : for the gills read the gill.
171 (115), line 34 : for Branchiae read Branchia.
172 (116), line 17 : for Samso read Samso.
173 (117), line 30 : substitute a semicolon for the period.
173 (117), line 31 : substitute a period for the semicolon.
175 (119), line 23 : for 1.3 read 13.
175 (119), line 23 : for 7 7.0 read 7-7.8.
175 (119), line 24 : for the light read the right.
176 (120), line 7 : for individual read individuals.
176 (120), line 21 : for leg read bag.
177 (121 (, line 1 : for branchiae read branchia.
177 (121), line 32 : for of the right hand are read of the right hand
one, is.
180 (124), line 10 : for spermatocysts read spermatocyst.
180 (124), line 33 : substitute a semicolon for the period.
183 (137), line 3 : for c read a.
183 (127), line 18 : for (F.) read (O. F. Miill.)
183 (127), line 21 : for inside read outside.
183 (127), line 23 : for the same read the same from the inside.
184 (128), line 13 : for d read a.
184 (128), line 16 : for b read a.
186 (130), line 12 : for of read f.
186 (130), line 26 : for 2. read 2, 2.
186 (130), line 33 : for e read c.
187 (131), line 37 : for to the twelfth read to b, the twelfth.
188 (132), line 12 : for cuticle read skin.
R. BERGH.
128 proceedings of the academy of [1880
February 3.
Mr. Meehan, Vice-President, in the chair.
Twenty -one persons present.
February 10.
The President, Dr. Ruschenberoer, in the chair.
Twenty-six persons present.
The death of Adolph E. Borie, a member, was announced.
Sartorius Muscle of the Gorilla. — Mr. Howard A. Kelly
described the sartorius muscle in the right leg of the Gorilla
troglodytes (young), from the Ogode river, West Africa, partially
dissected, and described b}- Dr. Chapman in the Proc. Acad. Nat.
Soi., Phila., 1879.
The muscle is 10 inches long, and \ inch broad. Tendinous for
about ^ inch at its origin, and its insertion. It arises from the
iliac bone at the beginning of the middle third of the distance
from betw^een the anterior superior spine of the ilium, and the
symphysis pubis. Its insertion is on to the inner face of the tibia
(which is 5i inches long), 3 inches below the knee joint.
Six inches from its origin the muscle is reinforced by a muscular
slip \ inch in breadth. This slip arises at the lower part of the
middle third of the femur, betw^een the origin of the quadriceps
extensor, and the insertion of the adductors, it joins the sartorius
muscle opposite the knee joint.
In consulting the literature on the myology of the Gorilla, wo
reference to any such slip has been found. Among all the
numerous anomalies recorded of this muscle, in the human sub-
ject, no corres|)onding variation has been found.
February IT.
The President, Dr. Rusghenberger, in the chair.
Thirty-three persons present.
A paper entitled " Description of a New Crustacean from the
Upper Silurian of Georgia, with remarks upon Calymene Cliv-
toni,''' by Anthony W. Vogdes, was presented for publication.
Germination in Acorns. — Mr. Thomas Meehan referred to
some intei'esting facts in the germination of Quercus virens, as
brought to his attention by W. St. J. Mazyck, of Georgetown,
1880.] NATURAL SCIENCES OF PHILADELPHIA. 129
South Carolina. It was generall}' known that in this species the
cotyledon did not divide into two lobes as usual in acorns, but
seemed to be of one solid mass, without any trace of a division.
In germination, however, two petioles were developed as in other
aconis, but instead of these being veiy short, indeed nearly
sessile, as in the ordinary white oak, thej' were produced appar-
enth' in the much advanced specimens sent by Mr. Mazj'ck to 1^
inches in length before the plumule and hypocotyledonary portions
of the embrj'o commenced their growth. In respect to the latter,
a small ovate, striate tuber, apparent!}' as one might judge from
the shrivelled specimens on hand, nearly one-fourth the size of the
acorn was formed, and from this tuber the radicle proceeded, and,
afterwards, the plumule on its upward growth.
Mr. Meehan said he had since examined sprouting acorns of
Quercus alba, Q. nibra, Q. arenaria, and Q. prinoidea, noticing a
very slight tendencv to a tuberous condition, onl}'- in the last
named. But in regard to the lengthening of the petioles, he was
surprised to find a variation in each species. In Quercus pri-
voides, the petioles were nearly- an inch in length.
He believed the discovery would be of great value to sj'stematic
botanists in the determination of species in this very difficult
genus, and should examine and report after an examination of
manv other species, but thought proper to call the attention of the
Academy to the matter in this early stage that due credit might
be recorded to Mr. Maz^'ck for his interesting discovery.
Mr. Edw, Potts, at the request of Mr. Meehan, had made
sections of both the acorn and the spindle-shaped radicle, with the
result of finding the cell structure of the latter an almost exact
counterpart of that in the nut: L e., sub-spherical cells of uniform
size, gorged with starch grains. So similar were they that it
would be nearly impossible for an observer to say which he was
examining but for the cortical tissue surrounding the root. It
seemed that the food supply of the young plant had been thns
withdrawn from a position exposed to hot sun and drying winds,
to one protected by the earth and in the direct line of growth.
No line of specialized cells could be discovered in the sections of
the nut, indicating the possibilitj' of a separation as in other
species into two cotyledons ; so that to all intents and purposes it
might be called monocotjledonous.
February 24.
The President, Dr. Ruschenberger, in the chair.
Twent3-nine persons present.
A paper entitled " Carcinological Notes, No. 3," by J. S. Kings-
ley, was presented for publication.
130 PROCEEDINGS OF THE ACADEMY OF [1880.
The death of John Rice, a member, was announced.
R. S. Hiiidekoper, M. D., David Townsend, John B. Wood,
Thos. Miles, Frances Emily White, M. D., and John S. Capp were
elected members.
The following were elected correspondents : — Robert Caspary,
of Konigsberg, Agostino Todaro, of Palermo, J. E. Bommer, of
Brussels, Teodoro Caruel, of Pisa, H. T. Geyler,of Frankfort-on-
the-Maine, Robert Schomburg. of Adelaide, and A. Inostranzeff.
of St. Petersburgh.
March 2.
The President, Dr. Ruschenberger, in the chair.
Twenty-eight persons present.
The death of Wm. Maxwell Wood, M. D., a correspondent, was
announced.
On a Filaria Reported to have come from a Man. — Prof. Leidy
exhibited a large thread-worm, which had been submitted to his
examination by Dr. J. J. Woodward, U. S. A. It was recently
presented to the Army Medical Museum, at Washington, by Dr.
C. L. Garnett, of Buffalo, Putnam Co., West Virginia. Accom-
panying the specimen, is the copy of a letter from Dr. Garnett to
Dr. Woodward, from which the following is an abstract : " During
the winter of 18t6, a man, a common laborer, aged about fiftj^,
presented himself to me for treatment having a gleety discharge
from the urethra, with a burning sensation during and after mic-
turition. Previously, he had been treated for gonorrhoea, and I
prescribed accordingly. The patient not improving, applied to
other practitioners. In April, 1878, he came to me with a round,
vivid-red worm, twenty-six inches in length, (the specimen you
now possess) which was alive and very active in its movements,
instantly coiling up like a watch-spring on being touched. Having
no work on helminthology for reference, the only description I
found which appeared to answer to the worm was that of Strongy-
lua gigas, in Niemeyer, vol. II, p. 41. The patient is an illiterate
man, with no motive for deception. He informed me that he dis-
covered the worm protruding from his penis and drew it out
without pain or difficulty. He was in much agitation and alarm
about the occurrence, fearing, as he said, that "there might be
more behind that one." For a few days previous to its passage,
his urine was of a milky hue and some time subsequently of a yellow
cast and slightly tinged with blood and mingled with mucus. The
man is truthful, and no doubt exists in my mind, or in the minds
of his neighbors as to the correctness of his statements. I regret
1880.] NATURAL SCIENCES OF PHILADELPHIA. 131
exceedingly that I did not appreciate the scientific interest of the
subject, and send you the specimen in a fresh state, but the busy
routine of a country practitioner's life leaves no time for the study
of other than subjects of practical value in one's every day ex-
perience."
The worm preserved in alcohol is much coiled, of a clay color
and opaque, or only feebly translucent, but more so at the head end.
If it is really a human parasite, it appears to differ from all those
heretofore described, and also seems different from other known
parasites. It certainly is neither Eusfrongylus gigo-^^, nor is it the
Guinea-worm, Filaria medinensis^ though nearly related to this.
Its characters are as follows : Body long, restiform, nearly uni-
formly cylindrical, smooth, shining, elastic,
tough, without evident annulation other than
transverse wrinkling, with the anterior ex-
tremity evenly tapering in the continuous
head, the end of which is rounded and smooth
or without appendages of any kind ; the pos-
terior extremity not tapering, with the caudal
end incurved, bluntly rounded, without ap-
Fig. 1. Fig. 2. pendages and imperforate or without evident
1. Cephalic extremity ; 2. anal or genital apcrturc. Mouth a terminal
Caudal extremity ; the diaso- .., , i- -ii , /.
nai marking indicates the porc without lips, papillte, or ai'maturc 01 any
ret?e^gume'l''^Teram:'. ^md. Tharyux Cylindrical, and opening into
ters. a straight cylindrical intestine, apparently
ending in a blind pouch. Generative organs unobserved. Length
of worm, 26 inches, greatest thickness, TS mm. Width of head
just behind the rounded extremity, 0'375 mm.; opposite the com-
mencement of the intestine, 0*625 mm.; at the middle, 1*5 mm.;
at the incurved caudal extremity, 1*5 mm. Length of oesopha-
gus, 1-125.
The worm, of exceedingly simple character, is clearl}'" neither a
Gordius nor a Mei-mis, and though apparently more nearly allied
to Filaria, a, more intimate knowledge of its structure may prove
it to be different. For the present it was proposed to distinguish
it with the name of Filaria restiformis.
On Rochelia patens. — Mr. J. II. Redfield remarked at the
meeting of the Botanical Section, that Rochelia patens was
founded by Nuttall, upon a plant collected by W^eth on Flat
Head River, in the Rocky Mountains, and was described in the
Journal of the Academy, 1st series. Vol. YII, p. 44, in 1834.
Dr. Gray in the Synoptical Flora of North America, II, p. 197,
remarks concerning the plant that it may be an Eritrichium, but
has not been identified, nor was it in the Academy's Herbarium.
Mr. R. stated that this specimen had been recently found among
the Academy's specimens of Echinospermum, and had been pro-
nounced by Dr. Gray to be Echinospernnnn florihundum^ Lehm.,
a species widely diffused in Western North America.
132 PROCEEDINGS OF THE ACADEMV OF [1880.
The following report iipon the plants introduced through the
medium of the Centennial Exhibition was read :
EEPORT ON PLANTS INTEODUCED BY MEANS OF THE INTEENATIONAL
EXHIBITION, isre.
The committee appointed on the 10th of October, IStG, at the re-
quest of the United States Centennial Commission, to examine and
report upon the subject of the introduction of insects* and plants
through the medium of foreign exhibits, respectfully reports that
it has delayed reporting on the plants till now in the belief that
some solitary plants might be ovei'looked, which producing seed
and increasing in following seasons, might be then discovered by
their greater numbers. But only those named in the list have been
found, and only in isolated specimens showing no disposition
whatever to spread and remain with us. So far, therefore, as the
object of the committee appointment is concerned, it may be said
in effect that no plant has been introduced, to our knowledge,
by the agency of the exhibition.
It is but justice to say that the plants have been collected by
our esteemed fellow member, Mr. Isaac Burk, whose familiarity
with the botany of Fairmount Park, rendered him particularly
fitted to detect any new introduction. Some of the few plants
named are from the western portion of our country, others from
Europe, and a few from Japan.
Lepidium sativum, L. Killingia pumila , M:ic.
Bunias Erucago, L. Fimbrutylis miliacea, Muhl.
Crepu tectorum, L. Cyperus diandrus, Torr.
Centaurea nigra, L. Triticum viUosum, Beand.
HypochcRris radicata, L. Triticum clavatum, Stedl.
Desmodium tomentosum, D. C. Leucoea Langsdorffiana, Steudl.
Cycloloma platyphylla, Moq.
Respectfully submitted.
John L. LeConte,
Geo. H. Horn,
Joseph Leidy,
J. Gibbons Hunt,
Thomas Meehan,
Committee.
*The report upon the insects was printed in the Proceedings of the
Academy of Natural Scienees of Philadelphia, for 1876, page 267.
1880.j natural sciences of philadelphia. 133
March 9.
The President, Dr. Rusohenberoer, in the chair.
Twenty-three persons present.
Mammary Glarids of Bats. — Dr. H. Allen exhibited specimens
of bats dissected to show the position and peculiarities of the
mammary glands. These bodies have been described as post-
axillary and two in number. For Desmodus this account is cor-
rect. For PhyUorhina^ Nycferis and the common red bnt of this
country {Atalapha {-^ Lasiurvs) novehoracensu) it is incorrect.
In the first two the glands answering to the axilla are low down
and have their nipples on a line with the middle of the clavicle. In the
common red bat the gland answering to the so-called post-axillary
is outside and below the axilla, but on a line with it. It occupies,
indeed, the lower third of the side of the chest and borders upon
the inferior line of the chest. In addition to this there is con-
stantly present a pectoral gland situated as in Quadrumana and
the human species. These glands resemble one another in general
appearance and size, being circular in form, without hair, of a
dull yellow color, possessing a well-developed nipple, and meas-
uring 3 lines in diameter.
It is interesting to observe that the specimens of non-lactating
bats show no external signs of mammse. The mammary regions are
covered with fur of the same character as seen elsewhere.
Neither in a female with embryos 2 lines in length is there any
external development. If such a specimen be dissected, the
locality of a rudiment of the gland can be detected by the posi-
tion of a small circle of thin, dark skin with a central white spot,
such structures representing the patch of modified skin and nipple
ready to receive the future developing active gland. No mam-
mary structure in this stage is anywhere visible, nor is there any
subcutaneous fat. Dissection of the body of the lactating female
on the other hand shows the mamma to be as large as the external
conformation, and the pectoral and lateral thoi*acic regions to be
occupied by a large but sharply limited mass of fat, which runs
up into the axilla and encroaches upon the dorsal surface of the
trunk. The rest of the under surface of the animal is without fat.
It is likely that there exists in the bat the same provision noted
in analogous structures of many lower animals, — namely, the
presence of secondary sexual characters (among which the milk
gland may be placed) which practically disappear in the periods
between sexual activity.
134 proceedings of the academy of [1880.
March 16.
Mr. Vaux, Vice-President, in the chair.
Thirty-five persons present.
A paper entitled " Carcinologieal Notes, No. IV," by J. S.
Kingsley.
The death of Dr. Win. M, King, U. S. N., a member, was
announced.
March 23.
The President, Dr. Ruschenberger, in the chair.
One hundred and fifteen persons pi-esent.
The following papers were presented for publication : —
" On the Gestation and Generative Apparatus of the Elephant,"
by H. C. Chapman, M. D.
" On a New Species of Hemitripterus from Alaska," by W. N.
TiOckington.
The death of Hector Tyndale, a member, was announced.
March 30.
Mr. Vaux, Vice-President, in the chair.
Thirt^'-eight persons present.
The death of Jacob Stauffer, a correspondent, was announced.
Paris Halderaan, Geo. B. Heckel and Emlen Physic, M. D., were
elected members.
The following were ordered to be printed : —
1880.] NATURAL SCIENCES OF PHILADELPHIA. 135
CAECINOLOGICAL NOTES, No. II.— EEVISION OF THE GELASIMI.
BY J. S. KINGSLEY.
I have endeavored in this paper to straighten out the species of
the " Fiddler Crabs," basing my work on the large collections of
the Academy of Natural Sciences of Philadelphia and of the
Peabody Academy of Sciences at Salem, Mass. My material has
been ample, embracing more than half the known forms, among
which are types of Smith, Guerin, Eydoux, Leconte and Sa}-,
with other specimens from Guerin's collection which were identified
by comparison with the types of Milne Edwards. I have reduced
considerably the number of specific forms, and in so doing I have
been actuated not by any desire to overturn the work of others,
but merely to arrive at the true limits of the species. A similar
reduction in other genera must be made, and will be made, by
any one who attempts to stud}' the forms of the whole world, and
does not limit himself to those of a small portion of its surface.
Among the important features of this paper is the extension of
the range of many forms, which has been accomplished either
by finding new localities among the specimens studied, or by a
union of two or more so-called species which bore different names
in different portions of the world.
I have endeavored to give descriptions and figures of all known
forms of Gelasimi, and when possible I have taken them from the
specimens themselves ; when I had no specimens, I have given a
description compiled from- some other carcinologist, and have
followed it by the initial of his name. The same remark will
apply to the figures. Localities from which I have examined
specimens are followed by an exclamation point (!), and the
museum in which the forms are preserved is indicated by an
abbreviation ; these abbreviations are : Phila. Acad., Academy of
Natural Sciences of Philadelphia, Pa. ; Peab. Acad., Peabody
Academy of Sciences, Salem, Mass. ; U, C, Union College,
Schenectad}', N. Y.
Genus GELASIMUS Latreille.
Cancer (pars.) Linne, Herbst, Fabricius, De Geer. Ocypoda (pars) Bosc,
Histoire Naturelle des Crustaces, ii, p. 240 (1828)'; Latreille,
' I have never seen a copy of the first edition of this work published in
the "An X " of the first French Republic (1802-3 of accepted chronology),
and my references are either quoted from the second edition by Desraarest,
or at second hand from Milne Edwards, or some other author.
136 PROCEEDINGS OF THE ACADEMY OF flggO
u
Histoire des Crustaces et de la Insects, vi, p. 27, " An. XI " (1803-4.)
Uca Leach, Trans. Linn. Soc, London, xi, p. 309 (1815). Gelasimus
Latreille, Nouvelle Dictionnaire d'Histoire Naturelle, xii, p. 517
(1817) ; Henri Milne-Edwards, Histoire Naturelle des Crustaces, ii,
p. 49 (1837) ; Annales des Sciences Naturelles, III serie, xviii, p.
144 (1853) ; Dana, Crustacea of the United States Exploring Expe-
dition, pp. 312 and 315 (1852) ; Hess, Archiv fiir Naturgeschichte,
xxxi, Pt. II, p. 145 (1865); Alphonse Milne-Edward.s, Nouvelle
Archives du Museum d'Histoire Naturelle, xi, p. 271 (1873) ;
Oonoplax (pars.) Lamarck, Histoire Animaux sans Vertebres, v. p.
353 (1818).
The genus Oelasimus belongs to Cyclometopa {Ocypodoidea of
Dana), family Macrophthalniidse (Dana), and sub-family Ocypo-
dinae of the same author. It is characterized by the rhomboidal
carapax, broader in front, the elongate ocular pedicels, the eyes
proper being placed at the extremity, and by the great inequality
of the chelipeds in the male.
In my studies I have found the characters derived from the
larger cheliped of the male to be the most constant, vrhile the
I'elative proportions of the carapax, the front and the margins of
the orbit, are of but slight importance and very variable.
But two species ever referred to this genus b}'^ authors, are
now^ referred to other genera ; Gelasimus cordiformis forming
the type of the genus Helcecius of Dana, and Gelasimus tel-
escopicus Owen, which belongs to the genus Macrophthalmus.
The genus may be divided into two groups, possibly of sub-
generic value, according as the front between the eyes is wide or
narrow, and the wide fronted section again according as the male
abdomen is seven or five jointed.
§ A. Front very narroiv between the eyes.
1. Gelasimus maracoani Latreille. PI. is, f. 1.
Ocypoda maracoani Latreille, Hist. Crust, et Ins., vi, p. 46 (1803).
Oelasimus maracoani luAtveiWe, Did. A^Wxst. Nat., xii, p. 519 (1817);
Desmarest. Coneid. (pars) p. 133 (1835); Edw., Hist. Nat. Crust.,
ii, p. 51 (1837) ; Ann. Sci. Nat., Ill, xviii, p. 144, pi. Ill, f. 1 (1852) ;
Dana, U. S. Ex. Ex, Crust., p. 318 (1853) ; White, List. Crust. Brit,
Mus., p, 85 (1847). Gonoplax maracoani Lamarck, Hist. An.
Sans Vert., v, p. 354 (1818). Oelasimus armatus Smith, Trans.
Conn. Acad., ii, p. 133, pi, ii, f, 5, pi. iii, f. 4 (1870) ; Report Pea-
body Acad. Sci., iii, p, 91 (1871).
Regions distinct, each branchial ornamented with a longitudinal
ridge, from which branch off smaller ones. Ischium of larger
ISSO.J KATURAL SCIENCES OF PHILADELPHIA. 137
cheliped with a prominent tooth below ; meros with a tooth on
posterior margin at the artieukition with the carpus, otherwise
smooth and rounded, its upper and lower margins with spiniform
teeth which are more prominent above ; carpus elongate, with in-
(iistinct tubercles. Hand very large, compressed, externally
tuberculate on the basal portion, above with several teeth like
those of meros, inferior margin proximaLy tuberculate, outer sur-
face of thumb with large shallow punctte, the lower portion being
marginate. Occludent margin with three rows of tubercles, the
middle one forming a prominence at the basal two-fifths, the other
rows undulating, extremity contorted, acute ; inner surface nearly
smooth, with a tubercular ridge running from the articulation of
dactylus to the middle of lower margin of the palm ; dactylus
lamellate, externally granulate, lower margin nearly straight,
upper margin arcuate, basally tuberculate, tip acute and nearly at
right angles with occludent margin, inner surface nearly smooth,
somewhat concave, with a longitudinal tuberculate ridge near the
occludent margin.
Bahia, Brazil! Dr. Wilson. Natal! [f] Dr. Wilson (labeled G.
natalensis). South America ! (Phila. Acad.) West Coast of Nicara-
gua ! McNiel (Smith's types in Peab. Acad.). Cayenne (Latr.
Edw.), Brazil (Latr. White), West Indies (White,', Eio Janeiro
(Dana).
The only differences between Smith's types and specimens
from other localities, are the more crowded spines on the
upper border of the meros and the more sparse tuberculation of
tiie basal portion of the hand, characters surely not of specifi^c
importance.
2. Gelasimus heterccheles Kingsley. PI. ix, f. 2.
Seba, Thesaurus, iii, pi. xviii, f. 8 (1758) ; Cancer vocans major
Herbst, Naturgesch. Krabben und Krebse, pi. i, f. 1 (after Seba)
(1790). Ocypoda heterocheles Bosc, Edit. I, "torn, ii, p. 197, 1802"
(teste Auct.) ; Edit. II, i, p. 250 (1828) ; Cancer iika Shaw, Natur-
alist's Miscellany. XIV, pi. 588 (after Seba).i Gelasimus maracouni
(pars) Desmarest, 1. c, p. 123 ( 1825). Gelasimus platydactylus Edw.,
Hist. Nat. des Crust., ii, p. 51 (1837) ; Ann. Sci. Nat., Ill, xviii, p.
' I have been unable to ascertain the date of this volujne. The first
volume of the series bears the date 1790, the twenty-fourth (and last)
1813, but no others are dated ; it would, however, seem probable that the
fourteenth volume appeared in 1803, while the ''.\n X," in which Bosc's
first edition appeared, embraced parts of 1802 and 1803.
10
138 PROCEEDINGS OF THE ACADEMY OF [1880.
144, pi. iii, f. 2 (1852) ; Savissure, Revue et Magazin de Zoologie, II,
V, p. 362 (1853) ; Smith, Trans. Coun. Acad-., ii, p. 122 (1870).
Gelasimus princeps Smith, 1. c, p. 120, pi. ii, f. 10, pi. iii, f. 3-3e
'1870) ; Report Peab. Acad., iii, p. 91 (1871) ; Lockington, Proc.
California Acad., vii, p. 145 (1877).
Carapax transversely iiearl}^ flat ; meros of larger cheliped
rounded posteriori}^, its lower margin crenulate, its upper pro-
duced into a broad, arcuate, laminiform, dentate crest ; carpus
elongate, externally tuberculate, inner margin crenulate, the inner
surface with one or two tubercles. Hand large, compressed, palmar
portion swollen, upper and lower margins tuberculate, external sur-
face of palm tuberculate, of thumb smooth, except a crenulated
ridge below. The inner surface smooth, with a tuberculate ridge
winning from the lower margin at the base of the thumb obliquely
upward and backward, and meeting a similar ridge from the base
of the dactjdus ; occludent margins of thumb with three rows of
tubercles, (the middle the most prominent) and somewhat angulated
beyond the middle. Dactylus with the upper margin and outer
basal portion tuberculate, the occludent margin rather prominent
in the middle.
Mexico! (Guerin-Meneville). Jamaica! (Dr. Wilson) Phila. Acad.
Cayenne, Edw. W. Coast Nicaragua ! (McNiel, Smith's types
Peab, Acad.). Lower California (Lockington). Mazatla.n (Saus-
sure),
Seba's figure represents the carapax as granulate and the front
rather broad (in these respects he has been followed by Herbst
and Shaw), otherwise his figure answers well. Bosc says that the
species is black ! Smith's types agree well with the Jamaica,
specimens which I have seen, except that the meral crest in the
Nicaraguan specimens is more distinctl}' den.tate.
3. Gelasimus bellator White. PI. ix, f. ?i.
Petiver, Opera, i, PI. 78, f. 5 (1767) ; Gelasimus bellator White, Cata-
logue British Musenm Crustacea, p. 36 (1847) ; (sine descr. ) Voyage
of H. M. S. Samarang, Crustacea, p. 49 (1848) ; Edw., Ann. Sci.
Nat., Ill, xviii, p. 146 (1852).
Carapax arcuate, front but little enlarged below tlie eyes.
Meros of larger cheliped posteriorly with an oblique rounded
ridge, its ijiipper and lower margins crenulate, the former even
denticulate ; carpus externally polished, above granulate, inner
margin denticulate, outside of palm and basal portion of dactylus
granulate., inside of palm granulate but without tubercular ridges
1880.] NATURAL SCIENCES OF PHILADELPHIA. 139
except a short curved one near the occludent margin ; thumb ex-
ternally margined below, its occludent margin forming a promi-
nence at the distal third ; dactylus with the margins nearly
parallel, the occludent one with scattered larger tubercles, tip
acute.
Australia ! (Dr. T. B. Wilson) Phila. Acad. Luzon (Petiver) Phil-
ippines (White).
4. Gelasimus styliferus, Edw. PI. ix, f. 4.
Gelasimus platydactylus Edw., 111. Edit. Regne Animal, Crustaces, pi.
xviii, f , 1 a (without date) . Gelasimus styliferus Edw., Am. Sci. Nat.
Ill, xviii, p. 145, pi. iii, f. 3, (1852) ; Smith, Trans. Conn. Acad., ii,
p. 118, (1870).
A species very near G. platydacii/lus, but liaving the marginal
crest of the arm less developed and the eye stalks terminated by
a small stylet as in the Ocypodaa (Edw.). Is possibly but a variety
of heterochelos.
Ouayaquil, Equador, (Edw.).
5. Gelasimus heterophthalmus Smith. PI. is, f. 5.
Oelasimus heterophthalmus Smith, Trans. Conn. Acad, ii, p. 116, pi.
11, f. 6, pi. iii, f. 1 (1870) ; Rep. Peab. Acad. Sci. iii, p. 91 (1871).
Meros of larger cheliped with posterior margin rounded, the
inferior crenulate, superior with a broad ci'cst, carpus with the
upper outer surface granulate, elsewhere smooth. Hand inflated,
basal portion of palm externall}' granulate, thumb punctate, with
an external elevated ridge. Inner surface of palm smooth, with
two rows of tubercles much as in G. heterocheles. Fingers com-
pressed, the thumb with a deep emargination at the base and a prom-
inent tubercle just beyond, occludent margin of finger nearly
straight.
Oulf of Fonseca, West Coast of Nicaragua .' McNiel (Smith's types
in Peab. Acad.).
This species is closely allied to G. heterocheles. When I exam-
ined the specimens, the prolongations of the ocular peduncles
described by Prof Smith were broken off.
6. Gelasimus heteropleurus Smith. PI. is, f. 6.
Gelasimus heteropleiirus Smith, Trans. Conn. Acad., ii, p. 118, pi ii, f.
7, pi. iii, f. 2 (1870) ; Rep. Peab. Acad., iii, p. 71 (1871).
Carapax but slightly convex, one side produced laterally. One
eye with a stylet about as long as the cornea, similar to those
found in certain Ocypodse. Meros of larger cheliped with the
HO PROCEEDINGS OF THE ACADEMY OP [1880.
margins denticulate, the upper one produced distally into a crest,
carpus granulate above. Hand externally granulate on the basal
portion, the upper and lower margins denticulate ; the inner sur-
face of the palm has an oblique line of tubercles running obliquely
upward and backward from the lower margin at the base of the
thumb to near the articulation with the carpus. Fingers short,
compressed, the thumb with the lower margin regularly arcuate ;
the upper margin of dactylus nearly straight as are the occludent
margins of each.
Gulf of Fonseca ! McNiel (Peab. Acad., Smith's types).
7. Gelasimus cultrimanus White. PI. ix, f. 7.
Gekmmus vocans Edw., Annales des Sci. Xat., Ill, xviii, p. 145, PI.
Ill, f. 4 (1852) ; Stimpson, Proc. Phila. Acad., 1858, p. 99 (1858) ;
Heller, Reise der Novara, Crustacea, p. 37 (1865) ; Hilgendorf, iu
van der Decken's Reise, p. 83 (1867) ; Alplionse Milne-Edwards,
Nouv. Arch, du Mus. d'Hist. Nat., ix, p. 273 (1873). Gelasimus
cultrimanus White, Catalogue Brit. Mus. Crust., p. 35, sine descr.
(1847) ; Voyage of the Samarang, Crust., p. 49 (1848). Gelasimus
nitidus Dana, U. S. Expl. Exped. Crust, p. 316, PI. X, f. 5 (1852).
Carapax smooth, arcuate. Meros of larger cheliped witli an
oblique ridge on the upi^er posterior surface which gradually dis-
appears before the articulation with the carpus ; the inner margin
somewhat cristate, distally with a prominent tooth and sometimes
traces of a second ; carpus externally granulate, a portion near
the articulation with the meros smooth, inner surface with a strong
spiniform tubercle. Palmar portion of hand swollen and exter-
nally granulate, gi'anules larger below. On the inner surface there
is an oblique tubercvxlar crest near the lower margin but not ex-
tending to it, and a second near the occludent margin. Thumb
with an impressed line on the outer surfoce, the lower margin
granulous, the occludent margin broadly excavate ; this excava-
tion is sometimes regularl}' curved, but generally shows traces of
a division into two sinuses ; the distal fourth bends abruptly
downward to meet the inferior margin. Finger granulate above
near the base, occludent margin nearly straight,
Philippines? Dr. T. B. Wilson (Phila. Acad.) ; Moreton Bay, Aus-
tralia! E, Wilson (Phila. Acad.); Coast of Malabar ! Guerin's
Collection (Phila. Acad.). This specimen (labelled " G. crassi-
manus Coll. Mus.") has the excavation of the thumb of the larger
cheliped plainly divided into two pai-ts. Java, Malabar (Edw.) ;
Nicobars (Heller) ; Zanzibar (Hilgendorf) ; New Caledonia (A.
M.-Edw.).
1880.] NATURAL SCIENCES OF PIIILABELPHIA. 141
There is a considerable confusion regarding this species. p]d-
wards considers this as the (Jancer vocans of Linne. Linne in
his tenth edition .p. (i2 >, 175Y) quotes Rumphius, PI. XIV, f. E. ;
and Catesby's Carolina, ii, PI. XXV. Rumphius' figure (of a
specimen from Amboina) represents a form with the fingers regu-
larly tapering, and resembling G. tetragonon more nearly than any
other species with which I am acquainted, but the figure is not
accurate enough to have an}^ systematic value. Catesby's figure
is the well-known Ocypoda arenai-ia of North America. Linne
(in the Amoenitates Academici, vi, p. 416) gives a description,
which does not at all apply to this species, and quotes in addition
Marcgrave, Piso, Rumphius, Catesby, and Seba, in the order
given, showing a still greater confusion. In his 12th edition, p-
1041, Gronovius and Petiver are added to the list, but no hints
showing what should be regarded as the Cancer vocanii. As there
exists such confusion, it is impossible to apply the name vocana,
with certainty, to an}' species, and for that reason I have thought.
it best to allow it to lapse into sj'nonymy and take the first recog-
nis ible description for this species.
8. Gelasimas marionis Desmarest. PI. ix, f. 8.
Oelasimus marionis Desm., Consid. sur le Crust., p. 124, PI. XIII, f.
1 (1825) ; Edw., Ann. Sci. Nat., Ill, xviii, p. 145 (1852).
Carapax smooth, and with each margin terminated by an acute
angle directed forward; an H -shaped impression on the carapax.
Ocular peduncles slightly enlarged at the extremity, and without
a terminal point. Inferior border of the orbit creuulate. Right
hand greatly larger than the left, greatly' compressed, basallv
granulate; finger straight, its sides smooth, its occludent margin
granulate ; thumb arcuate below, with its internal border broadly
excavate in the middle, and armed with fine teeth. Length, 8
lines ; breadth, one inch (Desmarest).
Manilla (Desm.^. Malabar {'Edw.''.
I have not seen any form corresponding to this description or
figure.
S. Gelasimus dubius Stimpson.
Gelasimux dubius Stra., Proc. Acad. Nat. Sci., Phlla., 1858, p. 99.
Carapax and front as in G. cultrimanus. Inferior margin of
orbit crenulate, externally angulate. Meros of larger cheliped
spinulose, hand stout, externally granulate or tuberculate ; in-
142 PROCEEDINGS OF THE ACADEMY OF [1880.
ternally with crests as in G. cuUrimanus, but less prominent.
Digits rather broad, externally sulcata ; inner margin nearly
straight, irregularlj'^ dentate, two or three teeth larger than the
others (Stm.).
Loo Ghoo (Stm.).
10. Gelasimus forcipatus White. PI. ix, f. 9.
Qelanmus forcipatus White, Catalogue Brit. Mus. Crust., p. 36, sine
descr. (1847) ; Voyage Samarang Crust., p. 50 (1848). Gelasimun
coarctatus Edwards, Ann. Sci. Nat., Ill, xviii, p. 146, PI. Ill, f. 6
(1853; ; Heller, Crustaceen Sud. Europas, p. 100 (1863) ; Alph.
Milne-Edwards, Nouv. Arch, du Mus. d'Hist. Nat., IX, p. 272, PI.
XII, f. 4 (1873).
Carapax convex, naiTOwed behind. Meros of larger cheliped
externally granulate, its margins denticulate. Carpus granulate,
inner margin produced but without a prominent tooth. Palm
externally granulate, its upper border slightly margined, its lower
tuberculate ; on the inside a few tubercles in a curved line near
the base of the dactjdus, and an oblique line from the lower
margin runs up to the articulation with the carpus, dactylus
granulate at the base, otherwise the hand and fingers are smooth.
Thumb regularly tapering, with an external impressed line, its
occludent margin regularly arcuate, with genei'ally a prominent
tubercle near the middle. Dactylus with a prominent distal
dentate lobe.
[?1 Odessa/ Guerin (Phil. Acad.). Philippines! Drs. Wilson and
Burroughs (Phil, Acad.). Australia! E. Wilson (Phil. Acad.).
Borneo (Adams and White). Odessa (Edw. ). New Caledonia,
(A. M.-Edw.).
I have united these two nominal species from an actual com-
parison of specimens. In the collection of Guerin-Meneville now
in the possession of the Philadelphia Academy, is a specimen
labelled " Gelasimus coarctatus Edw., Cat. Mus., Paris, Odessa,"
and which was probably one of the original specimens which was
the foundation of Edward's description. I am strongly inclined
to doubt of the authenticity of the locality " Odessa," as I have
been unable to find any other authority than that of Edwards.
Marcussen in his Fauna of the Black Sea (-Archiv. fur Natur-
geschichte xxxiii, pp. .358-363, 1867) does not mention it. His
subsequent paper and that of Uljanin, I have not seen. Heller
merely quotes from Milne-Edwards.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 143
11. Gelasimns arouatus De Haan. PI. ix, f. 10.
Ocypode ( Gelasimns) arcuata De Haan, Fauna Japonica, Crustacea, p.
53, PI. VII, f. 2 (1835). Gelasimns arcuatus M.-Edw., Ann. Sci.
Nat. Ill, xviii, p. 146 (1852) ; (?j Krauss, siid Afrikanische Crus-
taceeu, p. 39 (1843) ; A. M.-Edw., Nouv. Arch, du Mus., ix, p. 373,
(187?,'.
Carapax with sides carinate, carina acute, scarcel.y granulate;
inferior margin of orbit granulate. Meros of larger cheliped,
above concave, below flat ; internally with an acute granular ridge.
Carpus externally convex, above flat, hand twice the breadth of
the carapax , fingers compressed, smooth, externally longitudinally
sulcate (De Haan).
Japan (De Haan). JSew Caledonia (A. M.-Edw.). [?J Natal Bay
(Krauss).
12. Gelasimns tetragonon Ruppell. PI. ix, f. 11.
Seba Thesaurus, iii, PL XIX, f. 15. ? Cancer serratan Forskal, Descr.
Animalium, etc., p. 87 (1775). Cancer tetragonon Herbst 1. c, i, p.
257, PI. XX, f. 110 (1790). Gelasimns tetragonon Ruppell, Beschrei-
bung und Abbildung 24 Krabbeu des rothes Meeres, p. 25, PL Y,
f. 5 (1836) ; Edw., Hist. Crust., ii, p. 52 (1837) ; Ann. Sci. Nat. Ill,
xviii, p. 147, PI. Ill, f. 9 (1859) ; White, Cat. B. M. Crust, p. 3G
(1847) ; Guerin, Voyage Coquille, p. 10 (1839') ; Heller Reise der
Novara, p. 37 (1868) ; Hilgendorf in van der Decken, p. 84 (1867) ;
Kossman Reise nach rothen Meeren, p. 52 (1877). Gelasimns
duperreyi Gueviu, 1, c, PL I (1826) ; Dana, U. S. Ex. Ex. Crust., p.
317 (1852). Gelasimns desjardinii Guerin, MS. Gelasimns tetra-
gonon var spinicarpa Kossmann, 1. c, p. 52. Kossman gives a
reference to a paper by Poulson, but as the title is written in Russian
I have not been able to verify it.
Carapax strongly arcuate, front not expanded below the eyes,
Meros of the larger cheliped with the upper margin terminating
distally in a strong spine, carpus smooth, the inner margin acute,
its basal portion sometimes expanded into a strong tooth. Hand
compressed, externally finely granulate, a shallow pit with coarse
punctffi near the base of the thumb ; internally granulate but
without tubercular ridges ; thumb with two prominences on the
distal lialf ; the finger regularly tapering.
Mauritius ! Dr. WiLson, Guerin's Collection ; Tongatabou ! Wilkes
Expedition; Tahiti! A. Garrett ; Sandwich Is ! Dr. W. N. Jones
' The title page of the volume bears the date 1830, but the introduction
to the Crustacea and Aiachnida is dated "15 Novembre, 1838," so that it
is probable that the volume did not appear complete until 1839. The
plates bear date 1826.
144 PROCEEDINGS OF THE ACADEMY OF [1880.
(Phila. Acad); Tahiti and Sandwich Is..' A. Garrett (Peab.
Acad.); Red Sea and Nicoiar Is. (Heller); Zanzibar (HilgCn-
dorf, ; Bourbon (Edwards) ; New Caledonia {A. Milne-Edwards).
13. Gelasimus acutns Sfm.
Gelashnus acutus Stm., Proc. Phila. Acad., 1858, p. 99.
Carapax narrowed behind, anterolateral angles prominent, acute,
marginal line distinct. P'ront narrow, not constricted, inferior
margin of orbit crenulate, externally acute, internal suborbital
lobe convex ; a crest on the sub-hepatic region parallel to the in-
ferior margin of the orbit, the included surface smooth. Larger
hand coarsely granulate, a tubercular ridge on the inner surface.
Fingers not longer than the palm, externally sulcate, inner margin
dentate, median tooth larger, but no sub-terminal tooth (Stimpson).
Macao (Stimpson).
14. Gelasimus forceps Milne-Edwards. PI. ix, f. 12.
Oelasimus forceps Edw., Hist. Nat. des Cmst., ii, p. 52 (1837) ; An-
nales des Sciences Naturelles, III serie, tome xviii, p. 148, PI. Ill, f.
11 (1852) ; White Cat. Brit. Mus. Crust., p. 36 (1847).
Cavapax narrowed behind, lateral angles prominent, acute;
orbits below with two denticulate margins. Meros and carpus
smooth, the lower margin of the meros crenulate, upper cristate,
finely dentate ; hand smooth or indistinctly granulate, fingers
long, slender, finely denticulate, the thumb with a distal lobe
(Edwards).
Australia (Edwards, White).
I have not forms referable to the two foregoing species.
15. Gelasimus longidigitum (nov.). PI. ix, f. 13.
Closely allied to forceps in shape of carapax, orbits below with
a simple smooth margin. Meros and carpus smooth, the inner
margin of the carpus acute, crenulate. Basal portion of the hand
externally obscurely granulate ; internally with an oblique tuber-
cular ridge, and a few tubercles near the base of the fingers.
Fingers compressed, long, finely denticulate, and narrower near
the base than at the middle point.
Moreton Bay, Australia ! E. Wilson.
16. Gelasimus smitMi (nov.). PI. ix, f. 14.
Carapax gibbous, front narrow ; meros with a strong, oblique
ridge on the upper outer surface, the inner upper mnrgin produced
into a prominent vertical crest. Carpus externally nearlj' smooth.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 145
the inner margin slightly produced and denticulate. I'ulni exter-
nally granulate above, smooth below, its upper margin granulate
and indistinctly indicated by an impressed line on the outer sur-
f;ice, and its inner surface smooth, without tubercular ridges,
except one at the base of the fingers. Fingers long, slender,
slightly compressed and regularly tapering, the extremity of the
dactylus somewhat expanded and excavate.
Natal! E. Wilsou (Phila. Acad.).
Named in honor of my friend Prof. S. I. Smith, of Yale College,
who has monographed the Amei'ican species of this genus.
17. Gelasimus urvillei M.-Edw. PI. i.\, f. 15.
GeUisiinus urvillei M.-Edw., Ann. Sci. Nat., Ill, xviii i). 148, PI. Ill,
f. 10 (1852).
Resembles closely G. forceps, but has the medio-frontal sulcus
nearly linear, and the fingers shorter, the anterior border of the
meros of the larger cheliped obtuse and granulate (M.-Edw.).
Vanikoro (M.-Edw.).
18. Gelasimus dussamieri M.-Edw. PI. x, f. 16.
Gelasimus dussumieri M.-^dw., Ann. Sci. Nat., Ill, xviii, PI. IV, f. 12,
(1852)? Hilgendorf in van der Decken's Reise in Ost Afrika,
Crustaceen, p. 84, PI. IV, f. 1 (1867) ; Alph. M -Edw., Nouv. Arch,
da Mus. d'Hist. Nat. IX, p. 274 (1873).
Resembles closely G. urvillei, but the accessory sub-orbital lobe
is less marked, the median sulcus of the front entirely linear and
the anterior border of the meros of the larger cheliped denticu-
late. Chela ver}- large, G. rubripes is closely allied, but appears
to be distinguished by the form of the fingers of the larger hand,
the larger tubercles of the carpus, etc., (Ex. auct.).
Malabar and Samarang (Edw.) ; New Caledonia (A. M.-Edw.) ;
Zanzibar (Hilgendorf).
19. Gelasimus rubripes Jacq. and Lucas. PI. x, f. 17.
Geliisimus rubripes Jacquinot and Lucas, Voyage des Astrolabe et Zelee
Crustacea, p. 66, PI. VI, f. 2 (1853) ; Heller, Reise der Novara Crus-
taceen, p. 38 (1867).
Orbits granulate above and below, carpus of larger cheliped
with the external portion granulate, its margins finely denticulate.
Hand prominently granulate, internally smooth except fine granu-
lations at the origin of the thumb ; below strongl^^ dentate, finger
smooth except at the base where it is granulate; the inner margin
of the thumb with three large teeth, the intervals between which
146 PROCEEDINGS OF THE ACADEMY OF [1880-
are finely denticulate. Thumb smooth below its inner margin
with several rows of granulations and a prominent tooth near the
middle (J. et L.).
Unknown (J. and L. ) Nicobars (Heller).
20. Gelasimus signatus Hess. PI. x, f. 18.
Oelasimus signatus Hess, Archiv fur Naturgeschichte, xxxi, p. 146, PI.
VI, f. 6 (1865).
" Front between the eyes not so small as a G. variatns, cheliped
one and a half times the breadth of the body; arm, carpus and
hand bright red, fingers white. Arm below with two rows of
pearlj' tubercles, fingers with an elevation at the middle of the
inner border, distallj' arcuate and pointed " (Hess).
Sydney, Australia (Hess).
21. Gelasimus crassipes White. PI. x, f. 19.
Oelasimus crassipes White, Cat. B. M. Crust., p. 36, sine descr. ;
Adams and White, Voyage Samarang Crustacea, p. 49 (1848).
? 0. brevipes Edw., Ann. Sci. Nat., Ill, xviii, p. 146 (1852).
" Carapace very much arched, suddenly narrowed behind, front
with a lobe without narrow stalk. Four hind pairs of legs thicker
and stronger than in the other species " (Ad. and White).
Philippine Islands (White).
There have been described three other species * belonging to
the narrow-fronted section, one of which has been made the type
of the genus AcanthopJax by Milne Edwards. A fourth species
from Bahia, Brazil, is in the collection of the Philadelphia
Academy-. So far as 1 am aware these are all females and are
represented by only a single specimen each, and as I am strongly
inclined to consider them the females of well-known forms I omit
descriptions of them.
* Oelasimus insignis Smitli, Trans. Conn. Acad., ii, p. 126 (1870).
Acayithoplaxinsignis'Edw., Ann. Sci. Nat., Ill, xviii, p. 151, PI. IV,
f, 23 (1852) ; Archives des Museum, vii, p. 162, PI. II, f. 1 (1854).—
Chili (Edw.).
Oelasimus ornatus Smith, Trans. Conn. Acad., ii, p. 125, PL II, f. 9,
PI. Iir, f. 5 (1870) ; Report Peabody Acad. Science, iii, p. 91 (1871).—
West Coast Nicaragua! McNiel (Peab. Acad.).
Acanthoplax excellens Gerstiicker, Archiv fiir Naturgeschichte, xxii,
p. 138 < 1856). -No locality.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 147
§ B. Front broad between the orbits.
* Male abdomen seven-jointed.
22. Gelasimus vocator Martens. PI. x, f. 20.
Cancer vocator Herbst, Bd. iii, h. iv, p. 1, PI. LIX, f. 1 (1804).
Gelasimus vocans Edw., Hist. Nat. Crust., ii, p. 54 (1837) ; 111. Edit.
Regne Animal, Cru.stacea, PI. XVIII, f. 1 (no date) ; White, Cat. B.
M. Crust., p. 36 (sine synon.), 1847.
Gelmimus vocans (pars) Gould, luvertebrata of Mass, p. 325 (1841).
Oelasimus vocans var. a Dekay, N. Y. Fauna Crustacea, p. 14, PI. VI,
f. 10 (1844).
Gelasim2is palustrisBdyr., Ann. Sci. Nat., Ill, xviii, p. 148, PI. IV, f.
13 (1852) ; Stimpson, Annals N. Y. Lyceum Nat. Hist., p. 62 (1860);
Smith, Trans. Conn. Acad., ii, p. 127 (1870).
Gelasimus pugillator Leconte, Proc. Acad. Nat. Sci., Philadelphia,
1855, p. 403.
Gelasimus brevifrons Stimps., Ann. N. Y. Lyceum, vii, p. 229 (1860) ;
Smith, Trans. Conn. Acad., ii, p. 131 (1870) ; Lockington, Proc.
Cal. Acad., vii, p. 147 (1877).
Gelasimus sp. Saussure, Memoirs Societe Phys. et Hist. Nat. Geneve,
xiv, p. 440 (1858,1.
Gelasimus vocator Martens, Archiv fiir Naturgesch., xxxv, p 1 (1869 ;
xxxviii, p. 104 (1872) ; Kingsley, Proc. Phila. Acad., 1879, p. 400.
Gelasimus pugnax, mordax et rapax Smith, Trans. Conn. Acad., ii, pp.
131, 135, 134, Pis. II, f. 1, 2, 3, IV, 2, 3, 4 (1870).
Gelasimus affinis Streets, Proc. Phila. Acad., 1872, p. 131.
Gelasimus crenulatus Lockington, Proc. Cal. Acad., vii, p. 149 (1877).
Carapax smooth, meros of the larger eheliped with its margins
denticulate or tuberculate, cai-pus externally granulate, internally
with an oblique tubercular ridge. Hand tuberculate, its inner
surface with a ridge running up from lower margin to carpal
groove ; in front of this are scattered granules. Thumb straight,
extremity obliquely truncate, finger strongl}^ arcuate, longer than
the thumb.
East Coast of America, from Cape Cod! to Para, Brazil! West Indies!
and Aspinwall ! West Coast of Mexico ! Panama!
The localities from which I have examined specimens number
over thirty and embrace several hundred specimens. I find in
the Guerin Collection two specimens from Mauritius which closely
resemble Cuban forms.
This is, without much doubt, the species intended by Herbst ;
Edwards quotes the Cancer jMlustris of Sloane as this species,
but aside from the fact that his History of Jamaica was published
in 1125, And his name is therefore ante-Linnean (and is also poly-
148 PROCEEDINGS OP THE ACADEMY OF [1880
nomial), Sloane gives not the slightest description, but says th^it,
it agrees perfectly with the figure of Marcgrave which is the G..
maracoani of authors. I think that any one studying as I have
large series of specimens, will agree with me in uniting these
various forms under one specific name, as the characters which
separate them are variable and not of specific importance. Prob-
ably G. minax should also be included here, as suggested by
Professor Smith.
23. Gelasimus minax LeConte. PI. x, f. 21.
Gelamnus minax LeConte, Pix)C. Phila. Acad., vii, p. 403 (1855; ;
Smith, Trans. Conn. Acad., ii, p. 128, PL II, f. 4, PI. IV, f. 1
(1870) ; Rep. U. S. Fish Commission for 1871-72, p. 545 (1875);
Kingsley, Proc. Phila. Acad., 1879. p. 400.
Carapax strongly arcuate longitudinally, the branchial regions
granulate anteriorly. Meros of larger chelipeds, with the upper
and lower margins tubei'culate as is the upper portion of carpus ;
inner margin of carpus with prominent tubercles, its inner surface
with an oblique tubercular ridge. Palm cristate above, externally
with large depressed tubercles above, smaller below, inner surface
also tuberculate and with a ridge of tubercles running obliquely
up from the lower margin at the base of the thumb to the depres-
sion into which the carpus folds, and a second curved one near the
base of the fingers. Fingers long, slender, regularly tapering,
finger longer than the thumb and distally strongly' arcuate.
Beesley's Point, Dennis Creek, N.J..' S. Ashmead (Phila. Acad.,
LeConte's types) ; Bhiffton, S. G. ! Dr. Mellichamp (Peab. Acad.) ;
Northampton Co., Va. / H. E. Websier (Union College); New
Haven, Conn., and St. Avgustine, Fla. (Smith).
24. Gelasimus annulipes M.-Edw. PI. x, f. 22.
Oekisimus annulipes M.-Edw., Hist. Nat. Crust., II, p. 55, PI. 18, 1.
10-13 (18371 ; White, Cat. B. M. Crust., p 36 (1847' ; Edw.. Ann.
Sci. Nat. Ill, xviii, p. 149, PI. IV, f. 45 (1852) ; Dana, U. S. Ex.
Ex. Crust., 317 (1853) ; Heller, Reise der Novara, Crustacea, p. 88
(1867); Hilgendorf in Baron Decken's Reise, p. 85 (1867) ; Monats-
berichte Berliner Akademie, 1878, p. 803; Kossmann, Reise nach
rothen Meeren, p. 53 (187>) ; Spence Bate in J. K. Lord's Natural-
ist in Vancouver. Gelasimus macrodactylus Edwards and Lucas in
D'Orbigny's Voyage, 27, PI. XI, f. 3 (1843) ; Nicollet in Gay's Hist.
Chili Zool., iii, 165 (1840); Edw., Ann. Sci. Nat., Ill, xviii, 149
(1852\ Gelasimus lacteus Krauss, Sud. Af. Crust., p. 39 (teste Hil-
gendorf j. Gelasimus pidchellus UtiTaitsou, Proc. Phila. Acad., 1858,
p. 100. Gelasimus annulipes \a,v albiinana kossmann I.e.. Gelmi-
mus rectilatus Lockington, Proc. California Acad. Sci., p. 148
(1877),
I
1880.] NATURAL SCIENCES OF PHILADELPHIA. 149
Carapax transversely nearly flat ; inferior margin of orbit
crenulate. Meros of larger cheliped smooth, angles rounded,
carpus the same with a few obsolete granulations on the upper
surface. Hand smooth, sub-marginate below, an oblique row of
tubercles on the inner surface, running up and back from near the
lower margin half way to articulation with the carpus, and two
sitttilar cuived lines near the articulation of the dactylus. Thumb
regularly tapering, a prominent tubercle near the middle, extremity
sub-excavate. Dactylus distally strongly curved, extending
slightly beyond the thumb.
Australia! E. Wilson; Singapore! Dr. McCartee (Phila. Acad.);
Zanzibar! (C. Cooke) "N. W. Boundary Survey, A. Campbell,
Commr., Dr. C. B. Kennerly" ! (Peabody Acad.) ; Seas of India
and Asia (Edw.) ; Ceylon, Nicobars Madras (Heller); Mozam-
bique Inhambeni (Hilgendorf) ; Pondicherry (White) ; Valparaiso
(Edw. and Lucas) ; Vancouver (Bate) ; Lower California (Lock-
ington) ; Tahiti [^im.) \ Red Sea (Kossmann).
25. Gelasimus laoteas DeHaan. PI. x, f. 28.
Ocypode (Gelasimua) lacteus DeHaan, Fauna Japonica Crust., p. 54,
PI. XV, f. 5 (183.3). Oelasimus lacteus Edw., Ann. Sci. Nat., Ill,
xviii, 150, PI. IV, f. 16 (1852) ; Stm., Proc. Phila. Acad., 1858, 100;
Miers, Proc. Zool. Poc, 1879, p. 36.
Carapax longitudinally strongly arcuate, transversely nearly
f.at; antero-lateral angles prominent; meros of larger cheliped
externally granulate, a constriction of the upper margin near the
articulation with the carpus, lower crenulate or even denticulate.
(Carpus externally smooth, inner edge acute denticulate ; hand ex-
ternally finely granulate, above more plainlj^ so ; a crenulated
ridge near the inner lower margin and one or two near the fingers.
Fingers elevated, strongly compressed, the thumb suddenly nar-
rowed near the apex.
Japan! E. Wilson ; Pondicherry ! Dr. T. B. Wilson fPhila. Acad.) ;
Japan (DeHaan) ; China (Edw. Stm.).
33. aelasimas splendidas Stm.
Qelasiinus splendidus Stm., Proc. Phila. Acad., 1858, p. 99.
Inferior margin of orbit crenulate, externally rounded. Larger
hand nearly smooth, internally with an oblique tubercular crest.
Crest at the base of the fingers nearly obsolete. Fingers long,
slender, slightly denticulate. Thumb with the apex excavate
(Stm.).
Hong Kong (Stm.).
150 PROCEEDINGS OF THE ACADEMY OF [1880.
I have not seen this species ; it, however, appears to be very
near annuHpes.
27. Gelasimus minor Owen.
Gelasimus minor Owen, in Beechey's Voyage of the Blossom ; Ap-
pendix ; Crustacea, p. 76, PI. XXIV, f. 2 (1831).
Oahu, Sandwich Is. (Owen).
Tliis species is very near the annuHpes of Edwards, the only
difference being the larger teeth of the fingers of tlie cheliped.
28. Gelasimus triangularis A. M.-Edw.
Oelasimus triangularis A. M.-E<Jw., Nouv. Arch, du Mus., IX, p. 275,
(1873).
Is distinguished from ctdtrimanus, forcipafns, arcuatus, tetra-
gonon^ duiisiimieri, perj)lexus &i latreiUei by the carapax greatly
larger in front and smaller behind ; the lateral angles are spini-
form and directed strongly forward, the front between the eyes is
large and rounded. Larger cheliped externall}^ smooth, palmar
portion long and proximally inflated. Inner surface with a gran-
ular ridge, inner margin of fingers dentate, finger a little longer
than the thumb. This species is allied to G. winor by the form
of the hand, but is distinguished by the more triangular carapax
(A. M.-E.).
New Caledonia (A. M.-Edw,).
29. Gelasimus gaimardi Edw. PI. x, f. 23.
Gelasirmis gaimardi Edw., Ann. Sci. Nat. Ill, xviii. lijO, PI. IV, f. 17,
(1852) ; Heller Reise Novara, Crust., p. 38 (1867).
Very near annuHpes, but having the front more prolonged and
more rounded below and the external [internal ?] crest of the hand
obtuse and not denticulate, resembling that of G. Jatreillei (Edw).
Tongatabou (Edw.) ; Tahiti (Heller).
30. Gelasimus panamensis Stm, PI. x, f. 24.
Gelasimus pan amensis Stm., Ann. Lye, VII, p. 63 (I860); Smith,
Trans, Conn. Acad., II, 137, PI. IV, f. 5 (1870),
Carapax depressed. Anterior and inferior margins of the meros
of the larger cheliped crenulated, posterior rounded. Carpus very
short, smooth • hand smooth externally and internally, fingers
regularly tapering.
Gulf of Fonseca! McNiel (Peab. Acad.).
31. Gelasimus pugillator.
Ocypoda pugillator Bosc, Hist. Xat. Crust., Edit. I, i, p. 197, 1802-3,
(teste Auct.) Edit. II, i, p. 250(1828) ; Latr. Hist. Crust, et Ins. vi,
47 (1803-4\ Ocypoda pugillator (■pars.^, Sa.j, Jour. Phila. Acad. I,
1880J NATURAL SCIENCES OP PHILADELPHIA. 151
71 and 443 (1817-18). Oelasimus pugillator Latr., Nouv. Diet, d'-
Hist. Nat. Edit. II, p. 519 (1817) ; Desmarest Consid. 123 (1825),
Edw., Ann. Sci. Nat. torn. cit. p. PI. IV, f. 14 (1852) ; Stm. Ann.
N. Y. Lye. VII, p. 62 (1859) ; Smith Trans. Conn. Acad. II, p, 136,
PI. IV, f. 7 (1870) ; Rep. U. S. Fish Comm. 1871-72, p. 545 (1875).
Gelastmus uocans (pars.), Gould, Invertebrata of Massachusetts, p.
325 (1841) ; Dekay, N. Y. Fauna, Crust., 14, PI. VI, f. 9 (1844).
Carapax polished, swollen, nearly quadrate. Meros of the
larger cheliped with the outer surface rugose, upper and lower
margins crenulate. Carpus granulate externally, its inner margin
acute; hand inflated, the basal portion granulate and margined
above and below ; inner surface rounded, granulate, but without
any trace of a tuberculate ridge except one formed by a continua-
tion of the inner margin of the thumb. Thumb nearly straight,
a ridge on the outer surface, a large tubercle near the middle of
the inner margin, the extremity obliquely truncate. The finger is
longer than the thumb, i-egularly tapering and distally strongly
arcuate. There is a specimen in the collection of the Philadelphia
Academy from Surinam which appears to be intermediate, in the
characters of the hand, between this and G. vocator. The fingers
are shorter, the granules on the outside of the palm much more
prominent than, in typical pugillator, and there are traces, though
faintly indicated of a tubercular ridge on the inside of the
palm.
JVew Jersey ! T. Say, Wm. Wood ; Manatee River ! S. Ashmead ;
Mauritius ! Guerin's Collection ; Greenpoint, L. I. ! S. F. Baird ;
Boston Earborf J. H. Slack (Phila. Acad.) ; Nantucket and Key
West, Fla. ! A. S. Packard ; Bluffton, S. G. ! Dr. Mellichamp ;
Savannah, Ga. ! no collector's name given (Peab. Acad.) ; Beaufort,
N. a H. E. Webster (Union College) ; New Haven, Conn., Eg-
mont Key and St. Augustine, Fla. (Smith) ; South Carolina and
Cayenne (Edw.).
32. Gelasimus chlorophthalmus Edw. PI. x, f. 2B, 27.
Gelasimus chlorophthalmus Edw., Hist. Nat. Crust. II, 54 (1837) ; Ann.
Sci. Nat. torn. cit. 150, PI. IV, f. 19 (1852); McLeay in Smith's
Zool. S. Africa, p. 64 (1838) ; White, Cat. B. M. Crust., p. 36 (1847);
Guerin, Iconog. Crust., PI. IV, f. 3; Hilgendorf in Decken's Reise
Crust, p. 85 (1867); Monatsberichte Berlin Akad., 1878, p. 803;
Gelasimus marionis Edw., Hist. Nat. Crust., II, 53 (1837) ; Gelasi-
mus perplexus Edw., Ann. Sci. Nat. torn, cit., 150, PI. IV. ,f. 18
(1852) ; Heller, Novara Crust, p. 38, PI. V, f. 4 (1867) ; A. M.-Edw.
Nouv. Arch. Mus. IX, 274 (1873) ; teste Hilgendorf.
i52 PROCEEDINGS OF THE ACADEMY OF [1880.
Carapax arcuate. Larger hand small, joints all smooth, fingers
short, frequently shorter than the palm ; the ridges on the inside
of the pahn either smooth or obsoletely granulate ; fingers dentic-
ulate, regularly arcuate.
Island of Bourou ! Guerin's Collection (Phila. Academy); Mauritius
(Edw,, White); Zanzibar, Mozambique and Mascarenes (Hilgendorf );
Java (Edw.), Ceylon and Madras (Heller); Neic Caledonia (A. M.
Edw.).
I also understand that Maillard found this species at Reunion,
Injt I have not seen the work.
Hilgendorf from an actual comparison of specimens says that
the perpJexus and chlorophthahnus of Edwards are the same. The
(r. stenodactylus of Lockington (Proc. California Acad., vii, p.
148, 1877), from West Coast of Lower California, would appear
from the description and a rough figure of the hand sent me by
the author to be near this species ; it certainly is not stenodactylus
of Edwards and Lucas.
33. Gelasimus subcylindricus Stimpson. PI. x, f. 29.
Gelasii/ius subcylindricus Stimpson, Ann. N. Y. Lye, vii, p. 63 (1859);
Smith, Trans. Conn. Acad., ii, p. 137, PI. IV, f. 6 (1870),
Carapax obscurely granulate. Margins of mei;os of larger
raeliped granulous. Hand internally without tubercular ridge
t'xcept two or thi'ee parallel curved rows near the base of the
Singers, externally granulate. Fingers closely resembling those
of G. vocator, the common east coast form (Smith).
Matamoras on the Rio Grande (Smith, Stm.).
3*. Gelasimus latreillei E Iw. PI. x, f. 31.
Gelasimus latreillei Edw., Ann. Sci. Nat., Ill, xviii, p. 150, PI. IV, f.
20 (1852) ; A. M. Edw , Nouv. Arch. Mus., ix (1873).
Carapax smooth, lateral angles far behind the front. Greater
cheliped smooth ; meros with the upper and posterior margins
rounded, the upper ending in a prominent tubercle, the lower
crenulate and prominent. Inner upper margin of carpus minutely
crenulate, the others rounded. Hand cristate above, externally
microscopically granulate, internally with a smooth ridge near
the lower margin, no tubercles present ; fingers slender, slightly
compressed, regularly arcuate, with fine tuberculations on the
cccludent margins, the extremity of the thumb subexcavate.
Philippines f Dr. T. B. Wilson (Phila. Academy; Isle of Borabora
(Edw.); New Caledonia (A. M. Edw.).
/^Froc. Ji. ]\r. S. PhO^id'. 2880. 1/ J^trdiAr TL ^.
my
J^-Ji ^Torimjuv JoAnjfoTu^ fA.ei^). J2-J^ AJ-alciricu proccifruz^ fA.tt^S).
X-
Da/l^'s JAvpi tn, ^fn^hcv. /Froc. ^.JV. S. Fhilud. I880. IlJ Au^zSr. JPL jr.
' Oy JO.
Jt ^<r^A,
^Q -Jl. ^d. aZhopapillosa (l>.). 12 -l3. ^4c<tnt7iod . pzlostv (M).
^//.;. JUy./- u. .■U..AU /I'tvc. ^.J^S. Fh^la^. JS80. III. M^r TIM.
J.6.
3~9 J..u,„-m,J f»f,rM,-/&,hi (Z.). J.:i-J4. Z r>ari<ins . J}
/I'roo. ^.2V. 6\ J^hi/cz^. J880. IVJ jf,uM>,
^ 3.
/'/. J//.
}t JlfJ-^A ■
J. —J6'. ^.r<7/tf/i«J»ri\i- />f/,>j;r f.M. ) t»rr .
f Froc.ji.N.S. P?iilcui.J''if^'K Vj j-r^^,7,r J-/ xi//
4-
7>.,ns A\vpl. in AJa^k,. /P7-OC. Ji.J\r.S. PhUa^. 1880. VI J M^br . J^l.^TT.
JJ — 20 . Tri^^pa Tnod^^fM . JB . 21-^22. Tt- r7<tt>i</eray ( Jf.) .
J -JO. Triop,i rru}€i*:rta.. Ji M ^/3 Tr r747t>i<fera jU
14. Jhl. p<tnt<i<i . B.
jhaLr £:r-pl. in, ^/.«--6^ f Proc .4.MS. FhUod. 1880. VIII J Jf,^i7>r- J-/-, xn.
J —</ -Po/t/o pa/fuiti , li IO~Jl. ^rcltftl . Jfonffrei/fnj-zj- (C).
rj—J8. ApJir-Uf/ . ^inttV/ , H. Jp-2/. Fo7y<^ . JTo/hoffi (Jf.J.
PROC.A, N,S, 1880
PL, IX,
KINGSLEYONCELASIMI.
PROC. A, N. S, 1880.
KINGSLEYONGELASIML
1880.] NATURAL SCIENCES OF PHILADELPHIA, 153
35. Gelasimus tangieri Eydous. PI. x, f. 30.
Oelasimus taiigieri Eydoux, Magazin de Zoologic, 1835, clvii, PL XVII;
Edw., Ann. Sci. Nat., 1852, p. 151, PI. IV, f. 21; Heller, Crustaceeu
des siidlichen Europas, p. 101 (1863j.
Carapax transversely nearly flat, the sides of the branchial
regions strongly arcuate ; above everywhere granulate ; lower
margin of orbit bimarginate. Meros of larger cheliped externally
roughened, the upper margin produced into an arcuate crest
which is fringed with hairs, the lower margin with two rows of
tubercles. Carpus elongate, externally with prominent tubercles,
an oblique ridge on the inner surface with a slender obtuse spine
at about the middle. Palm margined above, and armed with
spiniform tubercles, externally with depressed tubercles, inferior
margin denticulate to the tip of the thumb ; internally a row of
tubercles near the base of the fingers, a second runs obliquely
upward from the lower mai'gin, meeting a third running backward
from the articulation of the finger, elsewhere internally smooth.
Fingers elevated, strongh' compressed. Dactylus Avith the upper
margin and outer basal surface tuberculate ; the rest of the outer
surface finely granulate. Occludent margins of both fingers with
three rows of tubercles, the margin of the finger regularly arcuate,
that of the thumb with a prominence near the middle, Meral
joints of the ambulatory feet denticulate above and below.
Tangier! Guerin's Collection (Eydoux's Types); West Africa! (Du-
chaillu); [?] Bahia ! E. Wilson (Phila. Acad.); Cadiz and Coasts of
Morocco (Edwards).
Edwards' figure is very poor.
36. Gelasimus perlatus Herklots, PI, s, f, 25.
Gelasimus perlatus Herklots, Additamenta ad Famiam, p, 16 (1851';
Edw., Ann. Sci. Nat., Ill, xviii, p. 151 (1852); Hilgendorf, Monats-
berichte Berlin. Akad., 1878, p. 806.
Carapax arcuate in both directions, above with patches of
o-ranules more prominent on the anterolateral portions. Larger
cheliped much smaller than is usual in the genus. Meros granu-
late, the posterior margin rounded, the anterior produced in an
arcuate crest; carpus and hand externally granulate. Hand
cristate above, internally without tubercular ridges, fingers com-
pressed.
Guinea! E. Wilson (Phila. Academy); Boutry, West Const of Africa
(Herklots); Loando, Chinchoxo, Liberia (Hilgendorf),.
11
154 PROCEEDINGS OF THE ACADEMY OF . [1880.
This species is closely allied to tangieri^ but differs in the pro-
portionately smaller cheliped without tubercular ridges on the
inner surface and in the more sparse tuberculation of the carapax.
* * Male Abdomen five-jointed,
37. Gelasimus stenodactylus Edw. et Lucas. PI. x, f. 33-35.
GJagimus gtenodactylus Edw. and Lucas, in D'Orbigny's Voyage
Crust, p. 26, PI. XI, f. 2 (1843) ; Nicollet in Gay's Hist, of Chili,
Zoologie iii, p. 105 (1849); Edw. Ann. Sci. Nat. Ill, xviii, 149
(1852). Gelasimus gibbosus Smith, Trans. Conn. Acad., II, p. 141,
PI. II, f. 11, PL IV, f. 8 (1870), Lockington, 1. c. Gelasimus lepto-
dactylus et poeyi Giieriu MS.
Carapax smooth, transversely flat, the regions sti'6ngly gibbous.
Meros and carpus of larger cheliped elongate, meros smooth, its
angles rounded, carpus externally unconspicuously granulate, its
inner margin acute denticulate. Hand externally smooth or
granulate, a tubercular ridge on the inside of the palm, running
obliquely from the lower margin to the groove in which the carpus
folds. Fingers much longer than the palm, internally denticulate.
Mexico ! Cuba! Brazil ! Guerin's Collection (Phila. Acad.) ; Gulf
of Fonseca! McNiel (Peab. Acad.) ; Gulf of California I W. N.
Lockington (Brown University) ; Valparaiso (Edw. and Luc).
Of the following species I can say but little. I have not seen
specimens which would answer to the descriptions and figui'es,
while the descriptions are so meagre that I cannot decide regard-
ing their affinities.
38. Gelasimus variegatus Heller, Verhandlung der Zool. Bot. Gesellschaft, Wien,
1862, p. 521.
" G. annulari affinis sed brachium chelipedum ad raarginem super-
iorem carinatum et dentatum, index dactylo paulo brevior acumin-
atus. Madras.''^
This is described as one of the specimens collected b}' the
Novara in her voyage around the world, but in Dr. Heller's final
memoir on the Crustacea of that expedition, this species is not
mentioned.
39. Gelasimus variatus Hess, Archiv. fiir Naturgeschiclite, XXXT, 146, PI. VI, f. 7
(1865). PI X, f. 32.
Cephalothorax smooth, greatly swollen. Front between the
eyes small. Greater cheliped of male somewhat longer than the
breadth of the carapax. There is a large triangular depression at
1880.] NATTRAL SCIENCES OF PHILADELPHIA. 155
the base of the index finger ; the index finger is somewhat bent,
tlie thumb is straight ; both are tuberculate on the inner margin.
Sydney, Australia .
40. Gelasimus porcellanus White, Cat. B. M. Crust., p. 36 (sine descr.), Adams and
White, Voyage of the Samarang, Crustacea, p. 50 (1848), Edw., Ann. Sci.
Nat., Ill, .wiii, p. 151 (1852).
" Eye pedicels veiy long, frontal portion of earapax not nar-
rowed at the base ; hind part of earapax much longer than the
sides. Fore-legs with the lower claws thickened at the end, the
inner margins of both claws with four larger tubercles amongst
the smaller crenules. Hab., Borneo.'''' (Adams and White.)
41. Gelasimus inversus Hoffmann, Recb. Faun., Madagascar, p. 29, Pl. IV, f. 23-2G.
Madagascar.
I have never seen this work, the quotation being taken from the
Zoological Record.
EXPLANATION OF PLATES.
1. G. maracoani. 19. G. brevipes (after 5Iilne-Edw.).
2. G. heterocheles. 20. G. vocator.
3. G. bellator. 21. G. minai (drawn from type).
4. G'. s^/?//er«.s (after Milne-Edw.) 22. G. annuHpes.
5. G. heterophthalmus. 2-3. G. gaimardi.
6. G. heteropleurus. 24. G. panamensis.
7. G. cultrimaau^. 25. G. perlatus.
8. G. marionis (after Desmarest). 26. G. chlorophtkalmus.
9. G. forcipatus. 27. G. perplexus (after Edwards).
10. G. arcuatus. • 28. G. lacteun.
11. G. tetragonon. 29. G. subcglindricus (after Smith).
12. G'./orcf/Js (after Milne-Edwards). 30. G. taagieri {irova. ty^^^).
13. G. longidigitum. 31. G. latreillei.
14. G. smithii. 32. G. variaius (.after Hess).
15. G. M?-i'i7/e2' (after Milne-Edw.). 33. G. s(ert.odaclyhts{h-on\'QvA.zi\).
16. G. dussiimieri (after Milne-Edw.). 34. G. stenodactylus (after Edwards
17. G. rubripes (after Hombron et et Lucas).
Jacquinot). 35. G. stenodactylus (from ISIekico).
18. G. signatus (after Hess).
156 proceedings of the academy of [1880.
April 6.
The Tresideiit, Dr. Ruschenberger, in the chair.
Forty -two persons present.
The death of Wm. Theodore Eoepper, a correspondent, was
announced.
April 13.
The President, Dr. Ruschenberger, in the chair.
Twenty-eight persons present.
A paper entitled " Description of a New Species of Catosto-
mus (C. cypho), from the Colorado River," by Wm. X. Locking-
ton, was presented for publication.
The death of M. Laporte, Count de Castelnau, a correspondent,
was aimounced.
Remarks on Pond Life. — Prof. Leidy remarked, that at the
invitation of Mr. Joseph W. Griscom, he had recently visited
some little ponds in the vicinity of AVoodbury, New Jersey,
which were remarkable for the profusion of minute invertebrate
life. The ponds oceup}' hollows in the woods, and consist mostly
of accumulated rain water, though several are likewise supplied
by springs. Several are completely di'ied up during the summer.
Mr. Griscom says they continue rich in animal life even during
the winter.
Of animals, entomostracans are. exceedingly numerous and
varied. Amono- some of the most beautiful and conspicuous were
noticed abundance of Brancliipus, of which two species from,
the same locality have been recently described by Mr. Ryder,
under the names of Chirocephalus holmanii and Streptucejihalus
sealii. There are also wonderful multitudes of many species of
copepods, ostracods and cladoceres, several of which are con-
spicuous for their large size and bright red color.
In one of the ponds a bright green Hydra was frequent, and in
another a pinkish one was abundant. These appear to be the
H. gracilis and H. carnea of Agassiz, but it is a question whether
they are not the same as the H. viridis and H. fusca of Europe.
Some of the Hydras were of a bright red color, and Mr. Griscom
intimated that this was due to the pinkish variety feeding on red
entomostracans. This was confirmed by some of the pink ones
which were brought home and kept in a jar with abundance of
1880.] NATURAL SCIENCES OF PHILADELPHIA, 15*1
red Cyclops, becomiug, after a few clays, as a result of feeding- on
the latter, of the same orange-red hue. Subsequentl^y, when food
became scarce, the red Hj'dras lost their bright color.
In one of the ponds, the stems of rushes and dead branches of
trees were invested with a bright grass-green stratum, consisting
of a bright green Vorticella, probably the V. fasciculafa of Midler.
The green color is dependent on chloroph^'l granules, as an
element of the structure, and not on food. The body of the
animal ranged from O'lOS mm. long by 0"Ofi mm. broad, to 0'12
mm. long by 0'09 mm. broad. A few measured were 0'1.5 mm. long
by 0*102 mm. broad at the peristome. In a large active bunch,
most of them measured 0-09 mm. long and broad. The pedicels
were from five to eioht times the length of the bodv.
In another pond, the water was rendered turbid from the pro-
fusion of Volrox glohator. In a bay of this pond filled with dead
leaves, a portion of water taken into a jar appeared opalescent
from the quantity of minute white flakes it contained. These, on
examination, proved to be Spirostomum amhiguum. In the same
pond, the Spatterdock, Nuphar ad vena, was just about unfolding
its leaves, and mau}^ of these were thickly invested with a clear
jelly, dotted with bright green spots. These proved to be Sfentor
yjolymorphus. On the underside of a few open leaves on the sur-
face of the water, were many spots of bright green and dull red-
dish. The former consisted of groups of the green Vorticella
before mentioned, the other consisted of attached groups of a
lilac- or amethystine-colored Stent07\ probably S. igneus. Similar
groups of this Stenfor were observed on a floating log, which had
been in the water since last 3'ear, as it exhibited attached many
statoblasts of a Plumafella. Ehrenberg describes S. igneus as
bright yellow or vermilion ; Stein as blood red, or often lilac-
colored, or vermilion to brownish red. Ehrenberg found it at-
tached to Hottonia. Stein says he never saw it fixed, but alwaj'S
swimmmg.
The Woodbury variety which might be named S. amethystinus,
was abundant and invariably found in conspicuous groups, visible
to the unaided eye, and when detached, thougli the animals swam
about actively, they were not onl}' disposed to become fixed, but
they actualh^ gathered together in groups. They all contained an
abundance of chloropbyl, apparently derived from food, but the
exterior structure was invariably of a distinct amethystine hue,
dependent on fine molecules. The color was more pronounced in
the longitudinal bands approaching the peristome. The nucleus
was spherical.
In the attached state, when the animal was fullj^ extended and
presented a trumpet shape, it was 0*6 mm. long by 0"18 mm. wide
at the peristome. This was a common size, but some measured
were O'Si mm. long. In the conical form, when swimming, indi-
viduals ranged from 0'2!r to 0-45 mm. long. In the most con-
158 PROCEEDINGS OF THE ACADEMY OF [1880.
•tracted condition of oval shape, they measured 0"18 mm. long by
0"15 mm. broad. The nucleus, 0*03 mm. in diameter.
Ehrenberg and Stein give for S. igneus one-sixth of a line
length, so that the variety indicated would appear to be much
longer.
April 20.
Mr. Thomas Meehan, Vice-President, in the chair.
Twenty-nine persons present.
April 27-
The President, Dr. Ruschenberger, in the chair.
Thirty-four persons present.
Lionel S. Beale, of London^ was elected a correspondent.
May 4.
Mr. Thomas Meehan, Tice-President, in the chair.
Twenty -eight persons present.
May 11.
The President, Dr. Ruschenberger, in the chair.
Twenty-two persons present.
The following papers were ordered to be printed in the
Journal of the Academ}'.
" The Terrestrial Mollusca inhabiting the Cooks or Harvey
Islands," by Andrew Garrett.
" The Placenta and Generative Apparatus of the Elephant,"
by Henry C. Chapman, M. D.
1880.] natural sciences of philadelphia. 159
May 18.
The President, Dr. Ruschenberger, in the chair.
Twenty-six persons present.
A paper entitled " On the Structure of the Oraug Outang," by
Henrj' C. Chapman, M. D., was presented for publication.
The death of Wm. Logan Fox, a member, was announced.
A fine portrait in oil, by Uhle, of Isaac Lea, LL. D., was pre-
sented to the Academy, and the following i-esolution was unani-
mously adopted :
Re.Holved, That the thanks of the Academy be presented to
Dr. Isaac Lea, for his gift of an admirable portrait of himself,
which has been long desired by the societ}', and especially by the
senior members, who are cognizant of his valuable contributions
to science, as well as towards the prosperit}^ of the Academy.
May 25.
The President, Dr. Ruschenberger, in the chair.
Twenty-two persons present.
The " Proceedings of the Mineralogical and Geological Section
of the Academy of Natural Sciences of Philadelphia, for the
years 18lt,18t8 and 1879," was presented for publication.
Henry S. Gratz, R. S. Peabody, Mrs. R. S. Peabody and Wil-
liam Barbeck, were elected members.
Adolf E. Nordenskiold of Stockholm, Carl Ochsenius of
Marburg, Oscar Hertwig and Richard Hertwig of Jena, were
elected correspondents.
The following were ordered to be printed : —
160 PROCEEDINGS OF THE ACADEMY OF . [1880.
ON THE STRUCTURE OF THE ORA.NG OUTANG.
BY HENRY C. CHAPMAN, M. D.
Tarious parts of the Orang, Siniia satyi'iis, L., have Ibeen dis-
sected, described, and figured bj Tiedemann,^ Owen,^ Saiidilbrt,^
Cuvier,^ Schroeder yan der Kolk and Yrolik,^ Rolleston,^ Selb^','^
Huxley,^ Bischoff,^ Barnard,^'' Langer," Gratiolet,!^ Spitzka,^^ and
others. It was hardly to be expected, the subject having been
investigated by such eminent observers, that I could hope to find
■anj'thing particularly new to science. It occurred to me, how-
ever, that it might not be altogether useless to bring to the notice
of the Academy a general resume of the results of jny dissection
of the Orang that died at the Philadelphia Zoological Garden in
Februarj" last, more especially as the memoirs referred to below
are scattered through the journals, and are often limited to descrip-
tions of certain parts of the animal only, such as the brain, mus-
cular svstem, etc.
My Orang was a j'oung male, supposed to be about three years
old. The following measurements were taken : From vertex to
rump, 16 inches ; upper extremity, 20|^ inches ; arm, "7 inches ; fore-
arm, 8 inches; hand, 5j inches; lower extremity, 17^ inches;
thigh, 5 inches ; leg, 6 inches ; foot, 6^ inches. What struck me at
once was the length of the upper extremitj-, it being 3 inches longer
1 Tiedemann, Zeit. Phys. Darmstadt, 1837.
2 Oweu, Proc. Zool. Soc, i, 1830, 1831.
■^ Saudifort, Ontleerlavindige Beschryving, Leideu, 1840.
^ Cuvier and Laurillard, Planches, 1849.
° Schroeder van der Kolk and Vrolik, Verhandelingen Kou. Nied. Inst. ,
1849; Verslagen Kon. Acad., 1862.
fi Rolleston, Nat. Hist. Rev., 1861.
^ Selby, Nat. Hist. Rev., 1861.
^ Huxley, Med. Times, 1864.
« Bischoff, Munich Abhand. 1870.
^^ Barnard, Proc. American Assoc, 1876.
'' Langer, Sitzungsberichte, "Wien, 1879.
^'^ Gratiolet, Plis Cerebraux des Primates, no date.
" Spitzka, Journal of Mental and Nervous Diseases, 1879.
Note. — I regret that when dissecting the Gorilla I was unacquainted
with Mr. Macalister's valuable paper in the Proceedings of Royal Irish
Academy for 1873^.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 161
tlian the lower one, the Orang agreeing nearly in this respect with the
Gorilla' which 1 dissected, the difference in the extremities in that
animal being Sj inches, whereas in the Chimpanzee- I found only
a difference of If inches. The foot in the Orang, however, was
i inch larger than the hand, whereas in the Gorilla the hand Avas •
^ inch larger than the foot ; in the Chimpanzee the difference in
this respect was f inch in favor of the foot. The foot in the Orang,
however, resembled superficially a hand much more than it does
in the Gorilla. Indeed the distinctness of hand and foot super-
ficiall}^ is more marked in the Gorilla than in the other anthro-
poids. I found the thoracic, abdominal and pelvic viscerae per-
fectly healthy. The animal seemed to have died from congestion
of the brain ; there was also some cerebritis. As the osteology
of the Orang has been thoroughl}^ described by Prof. Owen^ and
others it will not be worth while for me to dwell on that part of
its organization. I will pass therefore to the muscular system,
and more particularly to that of the extremities, as being the
most interesting as compared with man.
Muscular Systems — In Prof. Bischoff 's* paper on the Gorilla an
excellent figure is given of the muscles of the face of the Orang,
from a preparation by Rudinger. These muscles were described
by Prof. Owen,-^ but not figured. The same facial muscles are
found in man and the Orang with the exception that there is but
one zygomaticus possibly corresponding to the zygomaticus minor
of man, though on account of its size it may represent both the
z3'gomaticus major and minor. The facial muscles in the Orang
are not as well differentiated as in man, rather hanging together. I
noticed that the digastricus had only the posterior head. There
was nothing peculiar, however, about the sterno cleido mastoid,
omohyoid, or the scaleni. The omocervicalis or elevator claviculte
passed from the transverse process of the atlas to the acromial
end of the clavicle, as I found it in the Chimpanzee and in the
Gorilla. The pectoralis major arose in three portions : the first,
from sternum and first intercostal space ; the second, from sternal
part of thiixl, fourth, fifth, and sixth ribs, and the third from costal
' Proc. of Acad, of Nat. Sciences, Philadelphia, 1878.
- Proc. of Acad, of Nat. Sciences, Philadelphia, 1879.
=* Trans, of Zool. Society, 1835.
* Beitrage, Munich Abhand., 1879.
» Proc. of Zool. Society, i, 1830, p. 28.
162 PROCEEDINGS OF THE ACADEMY OF [1880.
portion of fourth, fifth, sixth and seventh ribs. This distinction in
origin is partly visible even in man. There was nothing notice-
able about the pectoralis minor or subclavius, supraspinati or
teres. The latissimus dorsi, as in all monkeys, gave off the slip
the latissimo condyloides, which, however, in the Orang scarcely
reached the condyle, and was pierced hy the ulnar nerve. The
biceps, triceps, and brachialis anticus were well developed, and
the external cutaneous nerve passed through the coraco-bracllialis
as in man. The anterior aspect of the forearm was quite human.
The pronator radii teres arose b}^ two heads, between which passed
the median nerve. The flexor carpi radialis and ulnaris and the
palmaris longus were well developed. The flexor sublimis did not
differ from that of man. The flexor profundus was rather sepa-
rated into two portions, one for the under and the other for the
remaining fingers. There was no trace of a flexor longus pol-
licis either as a distinct muscle or as a slip from the flexor pro-
fundus. The abductor, flexor brevis, adductor and opponens
pollicis, abductor flexor brevis, and opponens minimi digiti, and
the lumbricales were all present. As regards the back of the fore-
arm, the supinator longus arose higher than in man. The supi-
nator brevis, and extensor radialis longior and brevior, extensor
ossi metacarpi pollicis and exterior secundi internodii pollicis did
not differ from those in man. The absence of an extensor primi inter-
nodii pollicis was noticeable, as was also the fact of the extensor
indicis giving a slip to the middle finger and the extensor minimi
digiti one to the ring finger, making eight tendons supplying the
back of the fingers with the four from the extensor communis
digitorum. The interossei were the same as in man. Briefly', the
upper extremity of the Orang in its muscles differed essentially
from that of man in the absence of the flexus longus, and primi
internodii pollicis and in the presence of the additional tendons
to the ring and middle fingers. The Orang agreed with the
Gorilla in not having a flexor longus pollicis, but disagreed with
it in having the pronator radii teres arising by two heads, in the
presence of a palmaris longus, in the additional tendons for ring
and middle fingers, and in not having the extensor primi internodii
pollicis. As compared with the Chimpanzee, the Orang agreed
in reference to the pronator radii teres and palmaris longus, but in
the extensor ossi metacarpi pollicis being single, and in the
absence of the flexor longus pollicis as a slip from the pro-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 163
fundus, and in the presence of the additional extensor tendons it
differed.
As might be expected from the elongated form of the pelvis and
the absence of the round ligament of the hip-joint in the Orang, the
glutei muscles differ somewhat from those of man. The glutseus
magnus (PI. 12, e) in the Orang — not as large or as fleshy as its glu-
teus medius — is inserted together with the tensor vaginae femoris,
which is scantily developed, if at all, into the fascia lata of the thigh,
the glutieus medius being inserted into the great trochanter. Parallel
with the lower edge of the glutteus medius (PI. 12, c), is seen a small
muscle rising from the edge of the great sciatic notch, and inserted
into the great trochanter (PI. 12, b). This muscle seems to corres-
pond to part of the pyriformis in man, the sacral portion of the
muscle not being developed in the Orang. The glutseus minimus
is represented by a muscle arising from the external edge of the
ileum, and passing almost verticall}^ downwards until inserted into
the great trochanter, close to the pyriformis (PI. 12, a). At first
sight this muscle seems much displaced if it is the gluteus mini-
mus, but if one can imagine the ileum (PI. 12, d) in the Orang to
be widened outwardly to the same extent as seen in man, there
would be little or nothing anomalous about the muscle. From
the position of the glutwus minimus in the Orang, it would seem
that this muscle would supplement, to a certain extent, the want
of the ligamentum teres, which, it Avill be remembered, is absent
in this ape.
In the Chimpanzee there is so little that is peculiar about the
gluteus minimus that I had no difficulty in identifying it, and the
same can be said of the Gorilla. In the account of the Chimpan-
zee by TrailP however, the glutreus minimus is described as a
distinct new muscle, the scansorius ; the muscle I have described
as pyriformis, Traill regarded as the glutanis minimus, the pyri-
formis, according to Traill, being absent. Since then, this so-
called scansorius muscle has been referred to b}^ Bischoff", Owen,
Huxley and others, as a distinct muscle. With all deference to
such eminent anatomists, I cannot see any essential difference
betAveen the scansorius of Traill, and the glutaeus minimus in man.^
1 Wernerian Transactions, p. 18, 1821.
^ On looking up the literature upon the anatomy of the Orang, I find
t'liat in 1876 Prof: Baruard, o]). cit., considered the scansorius as being
homologous with the glutaius minimus, and mentioned in his paper that
164 PROCEEDINGS OF THE ACADEMY OF [1880.
The obturators, gemelli and qiiadratus femoris, were well de-
veloped. There was nothing peculiar about the muscles of the
thigh either on the anterior or posterior surface ; the rectus
arose, liowever, only from the inferior spine of the ileum. In
the leg anteriorly, I noticed the tibialis anticus divided into
two tendons ; otherwise, the muscles were as in man. The
peroneus longus and brevis were well developed, but there was
no peroneus tertius. The soleus, as usual in monkey's, had only
the plantar head, and there was no trace of a plantaris, although,,
according to Sandifort, it is present. The flexor longus digi-
torum supplied the perforating tendons for the second and fifth,
the flexor longus hallucis those for the third and fourth digits.
There was no slip from the longus hallucis for the big toe,
that muscle, therefore, except from its origin, scarcely deserves
that name. The flexor brevis digitorum supplied the perforated
tendons for the second and third toes. Those for the fourth
and fifth came off from the flexor longus digitorum. The
tendon for the fifth toe was not perforated. There was a connect-
ing slip between the third and fourth tendons. The external head
only of the flexor accessorius was pi'esent. In addition to the ab-
ductor, flexor brevis and adductor of the hallux, there was a well-
marked opponens hallucis. The lumbricales for the second and
fifth digits came from the flexor longus digitorum, those for the
third and fourth digits from the flexor longus hallucis. The ab-
ductor and flexor brevis minimi digiti were well developed, but
there was no transversus pedis. The interossei were like those
of the hand, Briefl}^, as compared with man, the leg and foot of the
Orang difl'er in the absence of the peroneus tertius, plantaris, flexor
longus hallucis and transversus pedis, in the fibular origin of the
soleus, and external origin of accessorius onl}^, in the distribution
of the perforating and perforated tendons for the toes, in the inter-
ossei, and in the presence of an opponens for the big toe. In this
latter respect the Orang differs not only from man, but from all the
other monkeys and anthropoids, the foot having a verj' hand-like
appearance, as compared with that of the Gorilla and Chimpanzee.
The foot of the Orang differs further in the absence of a special
Prof. Humphrey held essentially the same opinion. I was not aware, until
I had finished my dissection, of the views previously published by these
anatomists, and am glad to have been able, independently, to come to the
same conclusion.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 1G5
flexor for the big toe. This is supplemented to a certain extent by the
opponeus, and in a parti}' developed accessorius. The perforated
tendon for the fifth toe in the Gorilla came from the flexor longus
hallucis, whereas in the Chimpanzee and Orang it is supplied hy
the tendon of the longus digitorum. If Prof. Huxley's canon be
accepted that the distinction between a hand and a foot consists
in the latter possessing tarsal bones, the peroneus longus and
brevis, the short extensor and short flexor muscles, then the pos-
terior extremity of the Orang terminates in a foot. It appears
to me, however, that the difference between the hand and foot in
man, the Gorilla, Chimpanzee, and the lower monkej^s, is greater
than that observed between the corresponding members of the
Orang,
Alimentary Canal, etc. — It is usually stated that the uvula is
absent in the Orang, and, on looking into the mouth, at first sight
this appears to be the case, as it does not hang down as in man
between the pillars of the fances — nevertheless it exists. I found
it pointing directly backwards in a straight line from the posterior
palatine spine. It contained the azygos uvula? muscle. Prof.
Bischofl"^ mentions also finding the uvula in the Orang, The cir-
cumvallate papillae of the tongue are disposed in the form of a
/\ , as in man ; I found this to be the case in the female Chimpan-
zee,- of which I gave an account, and also in a male which I had
the opportunity recentl}' of dissecting. The salivar}^ glands with
their ducts were well developed, the submaxillary being ver}-
large both relatively and absolutel}' , as compared with man. The
stomach in the Orang (PI. ISjfig. 1) is not so human in its form as
that of either the Gorilla or the Chimpanzee, the cardiac portion, two-
thirds of the stomach, beinsf more elongated and constricted from the
pyloric part, which was tubular. The greater curvature measured
6 inches, the less 4. The small intestine was 8 feet 4 inches in
length, the large 4 feet. The constant presen(-e of valvula? con-
niventes in the small intestine of the Orang appears even at the
present day questionable b}' some anatomists. In speaking of
these folds occurring in the Gorilla, Bischoff^ refers to Owen not
finding them in the Orang, while they are said to exist by Saudi-
fort. Maj-er and Barkow. As to his own opinion on the subject,
he expresses himself as folloAvs : " Die beiden jetzt auf's Neue
1 Beitrage sur Gorilla, p. 37, ^ Op, cit., p, 57. ^ Op, cit., pp. 40, 41,
166 PROCEEDINGS OF THE ACADEMY OF [1880.
von mir untersiichten Diinndai'me cles Orangs aiis Dresden nnd
aus der hiesio-en Z00I02:. Sammlnno- sowie der eines Zweiten
Chimpanzee aus Dresden, zeigen keine Spur der genannten Fal-
ten. Ich halte nach alle diesem ihre Gegenwart beim Orang nnd
Chimpanzee fur zweifelhaft ; beim Gorilla, wenn gleich in
schwaeher Entwicklung, fiir gewiss ; individuelle Yerseheiden-
heiten sind doch in einem solchen Punekte nicht wahrscheinlich."
I foimd indications of valvuhie conniventes in the Orang, but of
the most rudimentary character as compared with man. In places
they run parallel with the long axis of the intestine (PI. 14, fig. 2),
then transversely as in man (PI. 14, fig. 3), then again as at first,
and afterwards again transversel3\ They are found in parts of the
jejunum and ileum. The valvulse conniventes I found very well
developed in the male Chimpanzee (PI. 14, fig. 4), but not at all in
the female. I noticed in the Orang the villi and solitary glands ;
the Peter's glands were very well developed. I counted fifteen,
some of which measured 4 inches in length. The coecum and ileo-
colic valve did not ditfer from the same parts in man. The ver-
miform appendix attained a length of 6| inches absolutely, and
was relatively much larger than that of man, reminding one of
the condition of this structure in the human embryo. As regards
the large intestine, the only noticeable peculiarities were the
large size of the solitary glands, and the fact that the mucous
membrane of the ascending colon was thrown into well-marked
longitudinal folds, with transverse connecting ones, exhibi ting-
quite a reticulated appearance (PI. 14, fig. 1). This is not the
case in the Chimpanzee. The peritoneum was disposed as in
man. The transverse colon was connected with the stomach,
as was also the case in the Chimpanzee, and Prof. Bischoff"'^
noticed that this obtains also in the Gorilla. As is well known, the
transverse colon in the monkeys can be raised entirely without
drawing up with it the stomach, with the exception sometimes
of the Macacques, in which I have noticed a slight peritoneal
connection between pyloric part of stomach and colon, indi-
cating a beginning of a gastrocolic omentum ? I did not notice
anything peculiar about the spleen or pancreas. The quadrate lobe
of liver was absent ; the spigelian lobe, however, was very well
developed ; the hepatic duct opened at a little distance from the
pancreatic. I found in the small intestine, five fine specimens of
1 Op. cit., p. 39.
1880. J NAT ORAL SCIENCES OF PHILADELPHIA. 16t
the Ascaris lumhricoides, and one in the large, and in the coecum
a Trichocephalus dispar. I believe this is the first time these
entozoa have been found in the same anthropoid. According to
Diesingi the Ti'ichocephahis is found in the Orang, and Cobbold-
states that Murie sent him aa Ascaris from the Chimpanzee.
Respiratory System. — In the Orang, as in the Gorilla and Chim-
panzee, particularly in the males, the ventricles of the larj^nx are
prolonged into the so-called laryngeal pouches. In 3' oung speci-
mens of the anthropoids, these pouches, though not so well devel-
oped as in the adults, can usually, however, be perfectly identified.
In dissecting m3^ Orang, after removing the skin in the cervical
region, I noticed what appeared to me to be the laryngeal pouches,
and by passing a tube into one of the ventricles of the larj-nx, the
pouch of that side could be readily inflated. On tracing, however,
the anterior wall of the pouch dowuiward, I noticed that it was
attached to the front of the sternum and clavicle, and on opening
the pouch and following its posterior wall, I found it attached to
the back of the sternum and first rib. Thus the interior of the
jDouch corresponded with the space between the two laj^ers of the
cervical fascia in man, usually filled with fat and absorbent glands,
but in the Orang it is empty and communicating with the interior
of the larynx. The pouch was not lined with mucous membrane,
resembling the remaining fascia, which was indeed continuous with
it. Supposing that mj" dissection really- represented the true rela-
tion of these parts, then, morphologically- speaking, the larj'ngeal
pouch in the anthropoids would be homologous with and replace
the two laj-ers of the cervical fascia in man, so familiar to the
surgeon. There was nothing especially noticeable about the vocal
cords, epiglottis or trachea. The lungs (PL 13, fig. 2), however,
were not divided into lobes as in the Gorilla and Chimpanzee.
Vascular System. — I did not notice about the heart anything
especially different from the human. In reference to the origin
of the vessels, however, the innominate gave off" the left carotid
and continuing an eighth of an inch then divided into the right
carotid and right subclavian, the left subclavian coming oflT sepa-
rately from the aorta (PI. 13, fig. 2). In the Gorilla and male
Chimpanzee I found the disposition of these vessels the same as
in man, which is the case in the Orang, according to Sandifort. In
the female Chimpanzee there were two innominates, a long and a
^ Helm., vol. ii, p. 534. 2 Entozoa, p. 291.
168 PROCEEDINGS OF TflE ACADEMY OF [1880.
short one, the latter dividing into left carotid and subclavian.
The arteries and veins of the extremities did not differ from those
of the Gorilla and Chimpanzee. I found in the Orang, as in them,
the " long saphenous artery " accompanying the nerve and vein
of same name. The mesenteric vessels exhibited loops along the
borders of intestine.
Genito-urinarij Apparatus. — The general appearance of these
structures resembled strikingly those of man (PL 15). The
kidne}'- measured 1^ inches in length, and exhibits only one
papilla. The ureters were 5 inches long. The bladder was 2
inches in length and 1 in diameter. The testicles measured | of
an inch in length, and were situated near the inguinal canal. The
cavity of the tunica vaginalis testes was shut off from the general
peritoneal cavity. The vas deferens was 4 inches in length, the
seminal vesicle 1 inch ; the seminal duct was yery short. The
caput gallinaginis was well developed, as was also the prostate.
The penis measured 2 inches in length, the glans was of cylindri-
cal shape. There was no bone in the penis. The Cowper's glands
were relatively large.
Nervous System — The brain of the Orang has been figured by
Tiedemann, Sandifort, Schroeder van der Kolk and Yrolik, Gra-
tiolet, Kolleston, etc. On account, however, of the few illustrations
extant, and of the importance of the subject, I avail myself of the
opportunity of presenting several views of my Orang's brain (Pi's
16 and 11), which was removed from the skull only a few hours after
death. The membranes were in a high state of congestion, and a
little of the surface of the left hemisphere had been disorganized
by disease, otherwise the brain w^as in good condition. It weighed
exactly 10 ounces. The brain of tlie Orang in its oeneral contour
resembled that of man more than those of either of the Chimpan-
zees which I examined. In these the brain was more elonaated.
The general character of the folds and fissures in the brain of the
Orang, Chimpanzee, and man are the same, there are certain
minor differences, however, in their disposition in all three. The
fissure of Silvius in the Orang runs up and down the posterior
branch pursuing only a slightly backward direction, the anterior
branch is small. The fissure of Rolando, or central fissure, quite
apparent, is, however, situated slightly more forward in the Orang
than in man. It differentiates the frontal from the parietal lobe.
The parieto- occipital fissure is well marked, bordered externally
1880. J NATURAL SCIENCES OF PHILADELPHIA. 1G9
bj the first occipital fold it descends internally on the mesial side
of the hemisphere, separating the parietal from the occipital lobes.
In the Oraiig, the parieto-occipital fissure does not reach the cal-
carine, being separated from it by the " deuxieme plis de passage
interne " of Gratiolet, or " untere innere Scheitelbogen-Windung"
of Bischofl!^'. I have noticed this separation as an anomaly more
than once in man.
According to Bischoff, this disposition obtains in the Gorilla,
and seems to be usual also in the Chimpanzee. In the female Chim-
panzee, however, on the left side I found the parieto-occipital
fissure passing into the calcarine, as in man. The frontal lobe is
easily distinguished from the parietal by the fissure of Rolando,
and from the temporal by the fissure of Sjdvius. In the Orang it
is higher, wider, and more arched than in the Chimpanzee. The
anterior central convolution in front of the central fissure runs
into the post-central convolution above and below, as in man. It
is difficult, however, to identify the three frontal convolutions
seen in man and the Chimpanzee, the frontal lobe of the Orang
dividing ratlier into two convolutions, the middle one being badly
defined. This is due somewhat to the length of the pre-central
fissure, which is as long as the fissure of Rolando, extending
farther upward than in man. There was nothing particularly
noticeable about the base of the frontal lobe ; on the mesial
surface it ran into the parietal. The part above the calloso-
marginal fissure in the Orang is not as distinctly divided into
convolutions as in man, though these are not constantly present
even in all human brains. The parietal lobe is separated from
the frontal by the central fissure, from the occipital and temporal
incompletely, by the parieto-occipital and Sylvian fissures. The
posterior-central convolution is well defined. The parietal fissure
in the Orang is more striking than that of man, resembling the
Gorilla's ; it is twice as long as the corresponding fissure in the
Chimpanzee, extending from the transverse occipital fissure, as is
sometimes the case in man, almost into the fissure of Rolando. It
is unbridged and without a break, and divides the parietal lobe
completely into upper and lower parietal lobules. The upper
parietal lobule is bounded externally by the parietal fissure ;
posteriorly it is separated from the occipital lobe, internally by
the parieto-occipital fissure ; externall}' it is continuous with the
occipital lobe, as the first occipital gyrus, anteriorlj' it is sepa-
12
110 PROCEEDINGS OF THE ACADEMY OF [1S80.
rated from the posterior central convolution more completely
than in man, by a fissure which runs parallel with the central
fissure. There is in the Orang, also, a fissure running parallel
with the parietal, which subdivides the u^pper parietal lobule into
inner and outer portions. The precuneus, or the space on the
mesial side of the parietal lobe between the parieto-occipital
fissures and the ascending branches of the calloso-marginal, is
well defined. The lower parietal lobule in the Orang divides
naturally into the supra-marginal and angular gyri. The supra-
marginal fold curves around the upper end of the posterior
branch of the fissure of Sylvius and runs into the superior tem-
l^oral g3'rus. The angular gyrus, which is very evident, arches
around the first temporal fissure, and becoming continuous with
the second occipital fold, passes then into the upper temporal
gyrus. The occipital lobe, separated from the parietal, internally,
by the parieto-occipital fissure, is continuous with upper parietal
lobule through the first occipital gyrus, and by the second
occipital gyrus with the angular. There are no sharp lines of
demarkation between the occipital and temporal lobes. In the
occipital lobe of my Orang the transverse occipital fissure was
present, and received the parietal fissure. The calcarine fissure
was well marked, but was separated in the Orang from the pai'ieto-
occipital fissure by the " deuxieme plis de passage interne " of
Gratiolet, the " untere innere Scheitelbogen-Windung " of Bischofl",
The cuneus of the Orang is therefore somewhat diff"erent from that
of man. In man I have seen these two fissures separated as an
anomal}-. The calcarine passed into the hippocampal fissure, so
tliat in the Orang, as in monkeys generally, the gyrus fornicatus
was separated from the hippocampal gyrus , whereas in man these
convolutions are continuous. This disposition has been noticed
in the Hylobates, in Ateles, and in one Chimpanzee, where
the calcarine did not reach the hippocampal. The first occi-
pital gyrus is very well developed, and, as the late Professor
Gratiolet observed, is one of the most strildng convolutions in
tlie brain of the Orang. It rises so to the surface that the
internal perpendicular fissure or external part of the parieto-
occipital fissure is almost entirely bridged over, the operculum so
characteristic of the monkey almost disappearing. It is con-
tinuous with the upper parietal lobule arching around the parieto-
occipital fissure. This convolution comes to the surface in the
1880. J NATURAL SCIENCES OF PHILADELPHIA. Ill
H3'lol)ates and Ateles almost to the same extent as in the Orang,
but it is more developed in the latter than in the Chimpanzee. It
is called also the "premier plis de passage externe," by Gratiolet,
the " obere innere Scheitelbogen-Windung,'' by Bischott", the •' lirst
annectant gyrus,"' X)y Huxley, and " first bridging convolution,"
by Turner. The second occipital convolution connects the occipital
lobe with the angular gyrus. In my Orang it was partl}^ concealed
bj- the first occipital. It was not as superficial as in man. The
third occipital gj'rus is continuous with that part of tlie temporal
lobe below the first temporal fissure. I noticed, also, in my
Orang the " quatrieme plis de passage "" of Gratiolet. On the
mesial side of the occipital lobe in n\y Orang, was well seen the
" deuxieme 'plis de passage interne" of Gratiolet, the " untere
innere Scheitelbogen-Windung " of BischofF, which separates the
ealcarine from the parieto-occipital fissure ; and in both the
Orang and Chimpanzee, more especially on the left side, I had no
difficult}' in recognizing the " premier plis de passage interne '' of
Gratiolet, its convexity turning inwards, while that of the first
occipital gyrus, or the " premier plis de passage externe," turns
outward. These two convolutions, the first occipital gyrus and
the " premier plis de passage interne," in my Orang were con-
tinuous. They are regarded as one by Bischott', forming his
" obere innere Scheitelbogen-Windung," but as two b}^ Gratiolet,
constituting his " premier plis de passage externe et interne."
The temporal lobe in the Orang is much less convoluted than in
man, or even in the Chimpanzee. Tlie first temporal fissure and
first temporal convolution are well marked, but the second and
third are badly defined. The fusiform and lingual lobes are sep-
arated b}" the inferior occipito-temporal fissures, the collateral
fissures of Huxley. The island of Reil was perfectly covered in
both the Chimpanzee and the Orang by the operculum, but was
not convoluted in my Orang. The surface in places was slightl}-
roughened. I noticed, however, three or four convolutions in
the Chimpanzee. On making a section of the left hemisphere of
the Orang I noticed that the corpus callosum was relativeh^
smaller than in man, but that the ventricle exhibited an anterior,
middle and posterior cornu, the corpus striatum, trenia semi-
circularis, thalamus opticus and fornix were well developed, the
hippocampus major with corpus fimbriatum were perfectly evident,
and the hippocampus minor larger relatively than in man. I did
1'72 PROCEEDINGS OF THE ACADEMY OF [1880-
not see a trace of the emmenentia collateralis ; this is often,
however, absent in man.
The cerebellum in my Orang was relatively larger than that of
man, but smaller than that of either the Chimpanzees I have dis-
sected, and was just covered and no more by the posterior lobes
of the cerebrum. This relation is still retained in my Orang,
though the brain has been lying in alcohol for three months since
it was taken out of the chloride of zinc in which it was placed
until the pia mater could be removed. During this period it has
been subject to the conditions, such as the want of the support of
the membranes, the eftect of pressure, etc., urged by Gratiolet,
Huxley, Rolleston, Marshall, etc., as sufficient to explain why
after death the cerebellum was uncovered by the cerebrum in the
Orang and Chim[)anzee, as held by Owen, Schroeder van der
Kolk and Vrolik, and BischotT. Every anatomist knows that
the brain after removal from the skull, especiall}' without the
membrane, if left to itself, very soon loses its shape. It is abso-
lutely necessary therefore to examine the brain in situ, and after
removal from skull to place it in some hardening fluid in which it
will float. Even with these precautions, through the change of
the surroundings, shrinkage, etc., the brain is alwa3's somewhat
altered. It happens, however, that I have had lying in alcohol
for some 3'ears a number of human and monkey brains. Among
the latter, examples of the genera Gebus^ Ateles^ Mavacus, Cyno-
cephalus, Cercopithecus, etc., taken out of the skull suflBciently
carefully, but preserved in the rudest manner Avithout any
regard to the above precautions. Now, while all of these brains
have somewhat lost their natural contour, the^' are not so changed
that in a single one, human or monkey, do I find the cerebellum
uncovered by the cerebrum, and in every instance the posterior
lobes overlap the cerebellum to a greater extent than I find is the
ease in my Orang. If the cerebrum and cerebellum in the Orang
and Chimpanzee invariably bear the same proportion to each
other as they do in man and the monkeys, why should not the
brain of an Orang or Chimpanzee, after lying in alcohol for some
years, exhibit the cerebellum covered l)y the cerebrum as in them?
Why sliould it be necessary to replace the brain of the Chimpan-
zee or the Orang in the skull, to make plaster casts, etc., if there
is no difference between their brains and those of man and the
1880.] NATURAL SCIENCES OF PHILADELPHIA. 113
monkeys, for there is no necessit}' of having recourse to such
measures to prove that the cerebellum is covered in the latter?
In the account I gave of the female Chimpanzee,' I stated that
I found the cerebellum uncovered. I had the opportunit}^ a short
time since, of verifjnng that statement in the male, noticing in
'^itu that the cerebellum was uncovered b}^ the posterior lobes.
This was found to be the case b^^ Mr. Arthur Browne, the Super-
intendent of the Phila. Zool. Garden, in a third Chimpanzee
which died there. With all deference to Prof Marshall's ^ photo-
graph of a plaster cast of the brain of a Chimpanzee, and how-
ever it may truthfully represent the relations of the cerebellum in
his specimen, I must say that it would be simply monstrous if
accepted as an illustration of either of mine, and with profound
respect for Prof. Huxley's^ opinion regarding the interior of the
skull being a guide for the determination of the proportion between
posterior lobe and cerebellum, I find it an3'thing but a safe one as
regards the anthropoid apes. For the space between posterior lobes
of brain and dura mater and bone, both posteriorly and laterally, I
find very variable in situ, due to the state of the blood vessels and
amount of fluid in arachnoid and subarachnoid cnvities. In speak-
ing of the Gorilla, Prof. Bischotf ^ observes, p. 100, " Das es bei
ersterem am wenigsten von oben Hinterlappen der grossen Hemi-
sphiire bedecktwird und bei der Betrachtung des Schiidel gewiss von
oben mit seinem hinterern Rande sichtbar wird." And in reference
to the Chimpanzee,^ p. 95, " Die Hinterhauptslappen des grossen
Gehirns bei diesem Affen wie bei dem Menschen das kleine Gehirn
iiberzogen und von oben fast ganz bedecken."' And Vrolik" states,
p. Y, of the Orang : " Ce lobe posterieur ne se prolonge pas autant
que chez I'homme ; il ne recourve pas si bien le cervelet du moins il
ne cache pas completement surtout vers les cotes." The fact of the
cerebellum being covered by the posterior lobes in my Orang and
that figured b}- Gratiolet, and but slightly uncovered in that of
Vrolik's, is no more strange than that Bischoff"^ should find it
covered in one Hylobates, and Prof. Huxle}' ** having stated it to
be uncovered in another.
I did not observe anything particularly noticeable about the
1 Proceed, of Acad., 1879. ^ Natural History Review, 1861.
^ Man's place in Nature, p. 97. * Das Gehirn des Gorillas, 1877.
^ Gehirn des Chimpanzee, 1871. " Amsterdam Verslagen, Deel 13, 1862.
' Beitrage zur Hylobates, 1870. ^ Vertebrate Anatomy, p. 411.
174: PROCEEDINGS OF THE ACADEMY OP [1880.
pons or medulla, except that in the latter the olivary bodies are
not as distinct as in man. As regards the peripheral nervous
system it was essentially the same as the human. As the brain of the
Orang which I have just endeavored to describe is the property
of the Academy, the animal having been bought and presented by
Mr. Wm. S. Yaux, and as it was desirable to preserve it in its
present condition, I could not make use of it to examine the
structure minutely. I would refer those interested in the his-
tology of the anthropoid brain, to Dr. Spitzka's paper.^
What can be inferred from the general oi-ganization of the
Orang as to its relation to the other primates ? The Orang like
man has twelve ribs, whereas the Gorilla and Chimpanzee have
thirteen ; on the other hand the carpal and tarsal bones are nine
in number in the Orang, while the Chimpanzee and Gorilla agree
with man in having eight. The upper extremity of the Orang
resembles that of the Gorilla in the absence of the flexor longus
pollicis. The Chimpanzee and man are alike in this respect, at
least the slip from the flexor longus digitorum in the former is
functionally a flexor longus. In the absence of a flexor longus
hallucis, and in the presence of an opponens hallucis, the Orang
difl'ers from man, the anthropoids and all the monkeys. The great
blood-vessels arise from the arch of aorta in the Gorilla and
man in the same way ; the same disposition is usually seen in the
Chimpanzee, rarely in the Orang. The lungs in the Orang are
not divided into lobes as in the Gorilla, Chimpanzee and man.
The stomach in the Gorilla and Chimpanzee is human in its
form ; in the Orang, however, it is quite different. The peri-
toneum in the Gorilla, Chimpanzee and Orang is like that of man ;
in the lower monkevs it is diff"erent. The brain of the Orang- in
its globular form, in the cerebellum being usually covered by the
cerebrum, and in the development of the first occipital gyrus,
resembles man more than that of the Gorilla and Chimpanzee.
On the other hand, the frontal and temporal lobes in the Orang
are not as much convoluted as in the Chimpanzee, and still less
than in man, and the island of Reil is not convoluted at all, at
least in my Orang.
It will be seen from the above illustrations, of which many
others might be given, that the Gorilla and man, in some respects,
agree with and differ from the Chimpanzee and Orang ; while
1 Op. cit.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 175
from other points of view the Orang approaches man more closelj'-
than either the Gorilla or Chimpanzee, and that as regards certain
muscles, man and the lower monkeys agree in having them, while
they are absent in the anthropoids. From these facts we maj--
reasonabl}^ infer that the ancestral form of man Avas intermediate
in character as compared with the living anthropoids or lower
monkeys, agreeing with them in some respects, and differing from
them in others. The Orang is closely allied to the Gribbons, the
Chimpanzee to the Macacques, and the gap between these and the
Senviopithecus is bridged over by the Mesojnthecus of Graudr^-.
Until,' however, the paleontologist will have procured more
material like that from Pikermi, and interpreted it as ably, it
will seem to me premature to oifer any detailed genealogical tree
of the Primates.
176 PROCEEDINGS OP THE ACADEMY OF [1880.
DESCRIPTION OF A NEW CRUSTACEAN FROM THE UPPER SILURIAN OF
GEORGIA, WITH REMARKS UPON CALYMENE CLINTONI.
BY ANTHONY W. VOGDES, U. S. A.
Calymene rostrata Vogdes.
This species differs in one aspect from the usual forms classed
under the genus Calymene^ in having a projecting process arising
directly from the cephalic shield in front of the glabella, and in
this respect resembles Homalonotus rhinotrojns of Angelin, a
species which has been referred by Salter, in his monograph of
British Trilobites, to H. Knightii. Salter says "the front margin
^'a- 1- is of most singular structure and may be
described as tricuspid. The narrow edge
is so deeply indented, and at the same time
folded, that the front portion overhangs
the rostral shield, forms one projecting
angle flanked by two smaller projections
Calymene rostrata Vogdes.
The glabella and fixed cheeks Opposite the axial furrows, exactly like the
showing the projecting pro- ,. , , , . , « r. ^.
cess. salient and re-entering angles of a forti-
fication." Our species has only the central triangular projection,
the margins of which are deflected, and the marginal border unites
and forms a triangular projection, directly in front and on the
median line.
The following characteristics are drawn from three specimens,
consisting of the glabella and fixed cheeks, and many p^-gidia
found associated with them at the same locality.
The glabella is convex and widens out posteriorly, being-
contracted in front ; the sides are marked with three lobes,
the basal one large, the middle lobe nearly spherical, the third
is somewhat obscurely defined. The fixed cheeks are separated
from the glabella by deep dorsal furrows, but opposite the eyes
the furrows are restricted hy a buttress thrown across it, nearly
touching the middle side lobes ; the cheeks are gibbous but
not elevated above the glabella, they are narrow along the
sides of the glabella and widen out laterally from the eyes.
The facial sutures cut the posterior angles of the head, but
anteriorl}^ from the eyes these lines run almost straight with
a slight tendenc}- outward, and pass over the margin. The
neck furrow is continued nearly to the posterior angles of the
1880.] NATURAL SCIENCES OF PHILADELPHIA. 17t
head. The frontal limb is triangnlar in outline, and prolonged
into a prominent projection, the bourrelet of the limb is defined
by a triangular ridge which forms the base of the projection.
The projection is formed by the thickening of the crust and by
the union of the outer marginal borders along the median line, it
is pointed and has its sides deflected. The space between the front
of the glabella and the base of the projection is somewhat de-
pressed.
The p3'gidium is obtusel}" triangular, with the front greatly
arched in uncrushed specimens, but this character seems to be
confined to the medium-sized specimens ; the larger forms are not
so much arched, and correspond in this respect to typical pygidia
of G. hlumenbachii. The axis occupies along the anterior border
about one-third of the width of the tail, and gradually tapers
posteriorly into an obtuse point ; it is marked with about eight or
nine articulations, the anterior one being slightly arched forwards,
but the others are extended almost straight rig. 2.
across it. The dorsal furrows are well defined.
The lateral lobes are marked with five pairs of
ribs, four of which are grooved and double half-
way up ; they are contracted along the dorsal
furrows, but widen out laterally. The ribs curve caiymene rostrata
" Vogdes. The pygi-
downwards and backwards, and are separated dium usually found
associated with the
from each other by well-defined grooves, the head.
last pair unite and form a ridge extending around the posterior
termination of the axis.
Geological Position. — Clinton Group, Taylor's Ridge, near
Catoosa Station ; and also at Dug Gap, Georgia.
Among the trilobite specimens wbicli I have collected in
Georgia, there are three movable cheeks and one pygidium
showing a strong resemblance to the same parts of Galymene
Glintoni as figured by Prof. Hall in Pal. N. Y., vol. ii, pi. 6(1 a,
fig. 5. These fragments were found associated with two glabella,
having characteristics not shown by the illustrations of the species
just referred to; therefore, for the purpose of comparison, I
carried the specimens to the American Museum, and through the
courtesy of Prof. Whitfield was enabled to study the trilobites
found in the Clinton Group of New York. The Georgia forms
are almost identical with those of New York, but show some
lYS
PROCEEDINGS OF THE ACADEMY OF
[1880.
Fis. 3.
variations from the typical C. Clint oni ; I shall, therefore,
describe these fragments.
Calymene Clintoni Vanuxem.
Glabella slightly convex, the base broad, so as to form a nearly
equilateral triangle. The sides are marked with three lobes, the
posterior one being twice as large as the middle lobe, but the
anterior one is ill-defined. The dorsal furrows are deep. The
occipital ring triangular in front, and narrowing out laterall}'.
The frontal limb is broad, and equal to half the length of the
glabella, and arched in front. It is worthy of remark that this,
character is not common to the minute glabella found in the same
beds. The fixed cheeks have a buttress thrown across them
extending along the sides of the glabella, but this
does not elevate them above it. The movable
cheeks are triangular in outline, and posteriorly
extended into spines, and correspond to tliose
figured by Prof. Hall, Pal. N. Y., vol. ii, pi. 66 a,
fig. 5, c. The}^ are convex laterally along their
inner half, and grooved near the outer margin, which
is defined by a raised border. The pj'gidium is
Vanuxem. The triangular in outline, and resembles the figure of
glabella and fixed , '^ ' =" .
cheeks showingthe this part given by Prof. Hall, Pal. N. Y., vol. ii,
pi. 66 a, fig. 5 a, d, except in size. The axis is
marked with about eight articulations. The lateral
lobes are not marked with ribs, as usual in Caly-
mene^ and in this respect the pygidium bears some
resemblance to that of C. arago and C. salteri^ two
European forms found in the Lower Silurian, and
described b^- Rousault in 1849.
Geological Position. — Clinton Group, Catoosa
Calymene Clintoni Station; also in the Hematitic bed at Dug Gap,
Vanuxem. The
movable cheek.
Calymene Clintoni
wide Irontal limb.
Fig. 4.
Geox'gia.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 1T9
CAECINOLOGICAL NOTES, No. 3.— REVISION OF THE GENUS OCYPODA.
BY J. S. KINGSLEY.
The following paper is based on the Collections of the Academy.
I have observed the rule adopted in previous papers of following
the locality from which I have seen specimens, hy an exclamation
point. In all other cases the name of the person who has reported
a species from any place follows that of the locality. The same
conservative spirit which pervades m^^ paper on the " Fiddler
Crabs " (Gelasimi) will be found in this. Many of the characters
given by authors prove to be of no specific value, but I liave not
ventured to unite forms unless I had specimens which corre-
sponded to each nominal species.
OCYPODA Fabiicius.
Cancer I. c, Fabr. Ocypoda Fabr., Suppl. Eut. Syst.. 347 (1798) ;
Edw., Hist. Nat. Crust., 11, p. 41 (1837j ; Dana, U. S. Ex. Ex.
Crust., p. 324 (1853).
Carapax transverse, rhomboidal or nearly square. Eyes stout,
the cornea occupjing the larger portion. Meros of external
maxillipeds shorter than ischium. Chelipeds unequal.
In many species there is a stridulating organ composed of a
row of tubercles on the inner surface of the palm, which, by being
drawn across a ridge on the ischium of the cheliped, produces a
noise.
§ 1. Ocular pedicels prolonged beyond the cornea as a spine
or style.
1. 0. ceratophthalma Fabr.
Cancer ceratoplithalmus Pallas, Splcelegia, p. 83, PI. V, f. 17(1772).
Ocypoda ceratophthalma Fabr., 1. c, 347 (1788) ; Latreille, Hist.
Crust, et Ins., vi, 47 (1803-4) ; Encyc. Meth., x, Pi. 274, f. 1 ; La-
marck, Hist. Aulmaux sans Vert., v, 252 (1818) ; Desmarest, Consid.
sur le Crustaces, 121, PI. XII, f. 1 (1825) ; Edw., Hist. Nat. Crust.,
11, p. 48 (1837); III. Edit. Regne Animal, PI. XVII, f. 1 ; Ann. Scl.
Nat., in, xvlii, p. 141 (1852j ; Krauss, S. African Crust., p. 41
(1843) ; Btimpsou, Proc. Phlla. Acad., 1858, p. 100 ; Hess, Archlv.
fiir Naturgeschicbte, XXXI, 143 (1865); Martens, Verhandl. Zool,
Bot. Gesellsch. Wlen, 1806, p. 381 ; Heller, Relse Novara, Crust., p.
42 (1867) ; Hllgendorf In v. Decken's Relse, Crust., p. 82, 1867 ;
A. M.-Edw., Nouv. Arch, du Mus., Ix, p. 270 (1873). Cancer
cursor Herbst., PI. I, f. 8-9 (1790). Ocypoda brevicornis Edw.,
180 PROCEEDINGS OF THE ACADEMY OF [1880.
Hist. Crust., ii, 48 (1837) ; Ann. Sci. Nat., Ill, xviii, 142 (1852) ;
Dana, U. S. Ex. Ex. Crust., p. 326, PI. XX, f. 3 (1853).
Ocypoda brevicornis var longicornuta Dana, 1. c, 337, PI. XX,
f. 4 (1853). Ocypoda agyptica Gerstaecker, Archiv. fur Natur-
gescliichte, xxii, 134 (1856) ; Heller, Sitzungsbericbte Wien Akad.,
xliii, p. 361 (1861) ; Hoffmann, Recli. Fauna Madagascar Crust., p.
14 (1874 teste Zool. Record) ; Miers, Ann. and Mag. Nat. Hist., V,
ii, 409 (1878).
Carapax nearly square, granulate, front strongly deflexed, orbits
sinuate, oblique, the lateral angles being far behind the front.
These angles are nearly right angles. Orbits with an indistinct
fissure below. Eyes terminated with a st3de which in most cases is
long and cylindrical, extending far beyond the orbits. In the
young, however, it is small and in some cases even wanting ; that
form, when small and conical, characterizes the nominal species
brevico7mis ; maxillipeds granulate. Meros of larger cheliped,
with the margins armed with spiniform tubercles, more prominent
on the anterior margins. Carpus granulate, with internally one
or two teeth. Hand externally acute, tuberculate, serrate below,
the inner surface with scattered tubercles. Stridulating ridge at
some distance from the base of the fingers, straight and composed
of rounded tubercles. Ambulatory feet with acute granules,
which exhibit a tendency to arrange themseh'es in rugfe.
]!fatal ! (E. Wilson) ; Mauritius ! (Guerin's Collection) ; Anjir,
Ternate, Amhoina, Adenare, Zanzibar, Benkula (Hilgendorf) ;
Sandwich, Is., Tahiti, Bonin, Loo Choo, Hong Kong Ouisma (Stm.);
Egypt, Mauritius, Bombay, Australia ( Edw. ) ; Ceylon and Nicobars
(Heller) ; Madagascar (Hoffmann) ; Tongatabu (Dana).
A specimen collected by the Wilkes Expedition ('' East Indies ")
has the carapax intermediate between this species and cursor.
The ocular st^yles are wanting. Milne-Edwards' figure in the
Regne Animal is diff'erent from any specimens that I have seen.
I agree with Kossmann in considering segyptica as but a variety
of ceratophthalma.
2. 0. platytarsis Edw.
Ocypoda platytarsis Edw., Ann. Sci, Nat., Ill, xviii, p. 141 (1853) ;
Heller, Reise Novara Crust., p. 43 (1867;.
Carapax wider than is usual in this genus and covered with
large granules. Superior margin of orbit sinuate, the external
angles rounded ; sides parallel about one-fourth of their length.
Orbits with an indistinct fissure below. Eyes spined as in ceratoph-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 181
thalma (teste Edw., the single specimen I have seen has the ej-es
broken). External maxillipeds granulate ; meros of larger cheli-
ped with the upper margin produced and dentate, the lower
spined, the posterior with transverse granular rugae, carpus
granulate ; hand with large granules, serrate below, stridulating
ridges carved and composed of crowded granules. Ambulators-
feet with rugre and subspiniform tubercles, dactyli broad.
Pondiclierry \ Guerin's Collection (labelled by Guerin "Ocypoda
platytarsis, Edw., Cat. Mus., Paris") and probably one of the
original specimens). Tahiti and Nicobars (Heller).
3. 0. urvillei Guerin.
Ocypoda urvillei Guerin, Voyage Cofiuille, Crust, p. 9, PI. I, f.
1 (1836), Edw. Hist. Crust., H, p. 49 (1837), Ann. Sci. Nat. HI,
xviii, p. 141 (1852), Owen in Beechey's Voyage Crust., p. 80 (1839;,
Dana, U. S. Ex. Exp. Crust., 328, PI. XX, f. 5 (1852).
Carapax wider than long, superior margin of orbit sinuate,
angles acute. Ej-es moderate, ocular spines short, extending only
to angles of orbit. Meros of larger cheliped rounded above, its
two other margins denticulate. Carpus with a strong internal
spine. Hand externally granulate, serrate above and below ; the
stridulating ridge nearly straight, a little remote from the fingers
and extending from the lower margin of the hand two-thirds of
the wa}' to the upper. (Guerin.)
Ta/u'^i (Guerin) ; Me Bouron (Edw.) ; Sanduoich Is. (Dana).
4. 0. macrocera Edw.
Ocypoda macrocera Edw. Hist. Nat. Crust., II, 49 (1837), Ann. Sci. Nat.
IV, xviii, p. 142 (1852), Heller, Novara Crust., p. 142 tl8G7).
Orbits wide, oblique, angle obtuse, ej^es with a spine as in 0.
ceratophthahna. Larger hand very short, broad and a little
spinose above ; its palmar portion broader than long. The fingers
of the smaller hand lamellate and very broad at their extremities.
Ambulatory feet roughened above. (Edw.)
E. Indies, Pondiclierry, {?] Brazil (Edw.) ; Tahiti, Nicobars (Heller).
5. 0. gaudichaudi Edwards et Lucas.
Ocypoda gaudichaudi Edw. et Lucas in D'Orbigny's Voyage, Crust.,
p. 26, PI. XI, f. 4 (1843', Edw. Ann. Sci. Nat. Ill, xviii, 142 (1832),
Nicollet in Gay's Chili, Zool. Ill, p. 163 (1849), Stimpson, Ann. N.Y.
Lye. Nat. Hist., VII, p. 61 (1859) ; Smith, Rep. Peab. Acad. Sci.,
Ill, p. 91 (1871) ; Streets, Proc. Phila. Acad., 1872, p. 240,
Carapax longitudinall}- strongly arcuate, distal portion of front
nearl}^ vertical. Superior border of orbit sinuate internally, its
182 PROCEEDINGS OF THE ACADEMY OF [1880.
external three-fourths nearly straight and directed slightl}^ back-
wards. Lateral angles nearly right angles, the sides behind them
being concave for about a fifth of the length of the carapax ;
oi'bits with a deep median fissure below. Eyes with a short conical
stjde reaching to, or slightl}^ beyond, the angle of the orbit.
External maxillipeds nearly smooth, or with a few inconspicuous
granules. Meros of larger cheliped with the upper and lower
margins spino-tuberculate, the posterior with transverse rugaj.
Carpus granulate ; hand subspinose above, finely serrate below,
internally granulate and with a well-marked transverse stridulat-
ing ridge, fingers lamellate, the extremities truncate.
Chili \ Guerin ; Panama ! Capt. Field and McNiel (Pliila. Acad.) ;
Gulf of Fonseca ! McNiel (Peab. Acad.) ; Callao(E6.vf. et Lucas) ;
Valparaiso (Dana).
6. 0. fabricii E Iw.
Oei/podafabricii 'Edw., Hist. Nat. Crust., 11, p. 47 (1837), Ann. Sci.
Nat. XVIII, p. 143 (1853), Hilgendorf in Decken's Reise Crust., 83,
PI. Ill, f. 1 (1867).
Carapax convex, finely granulate, front strongly deflexed,
orbits strongl}' sinuate ; lateral angles acute and some distance
posterior to the base of the rostrum ; sides parallel for about a
third of the lengtli of the carapax. Orbits witliout emargination
below ; e3'es with a short conical style, not reaching beyond the
orbital angle. Anterior margin of mei*os of larger cheliped cren-
ulate, distally spinose, posterior margin rounded, rugose. Carpus
granulate, as is the outside of the hand ; inner surface of the
hand polished, with minute scattered granules ; stridulating ridge
straight, composed of small, closelj'' set granules ; lower margin
of hand finely serrate ; fingers of moderate length. Joints of
ambulatory feet with transverse rugoe.
Australia ! E. Wilson; Natal ! E. Wilson ; Oceanica (Edw.) ; Zan-
zibar (Hilgendorf),
7. 0. cursor.
Cancer cursor Linn., Syst. Nat. Edit., xii, p. 1039 (1766). Ocypoda
ippeus Olivier, Voyage, p. 334, PI. XXX, f. 1 (1807); Savigny, Egypt,
PI. I, f. 1; Lamarck, An. sans Vert., v, p. 353 (1817); Desm., Con-
sid. Crust., p. 131 (1835); Edw., Hist. Crust., ii, p. 47 (1837); Mose-
ley, Notes by a Naturalist on the Challenger, pp. 48-49, woodcut,
1879. Ocypoda cursor DeHaan, Fauna Japonica, Crust., p. 89; Edw.,
Ann. Sci. Nat., Ill, xviii, p. 143 (1853); Stm., Proc. Phila. Acad.,
1858, p. 100; Heller, Crust. S. Europa, p. 99 (1863).
1880.J NATURAL SCIENCES OF PHILADELPHIA. 183
Carapax arcuate, front strong!}' depressed. Upper margin of
orbits but slightly sinuate and nearly transverse. Lateral angles
acute, the sides converging posteriorly, making the carapax widest
at the angles. Orbits below with a slight median fissure. Ej-es
terminated bj' a short conical style armed with pencil of hairs.
Meral joint of cheliped with the spines and rugse much less
prominent than in 0. ceratophthalma. Carpus internally' with a
denticulated tooth. Hands small, externally with depressed
granules, the lower margin finely serrate, the upper rounded and
without spiniform tubercles. Internally the hand is nearly smooth
except below where there are granules similar to those of the
outside. The stridulating ridge is ver}' near the fingers and is
crossed by numerous fine striae. The propodal joints of the
ambulator}' feet spined below, the dactyli of the second pair hair3\
Senegal! (Guerin"! ; Syria, Egypt (Edw.) ; Cape Verdes (Auct.);
Greece (Guerin).
This is probabl}' the species described by Hasselquist (Iter
Palestinum, p. 433, 1T57) under the name Cancer anomalus^ but
he gives many characters which will appl}' to no known decapod
and so to avoid confusion I refrain from applying his name to
this species. McLeay, in Smith's Zoology of South Africa, men-
tions '■^Ceratophthalma cursor DeHaan," an expression not to be
found in the pages of that Dutch Carcinologist.
8. 0. ryderi Nov.
Carapax coarsel}^ granulate, the upper margins of the orbits
sinuate, transverse, lateral angles acute, sides arcuate, the carapax
being widest at the anterior third, as in 0. arenaria, the lateral
margins anteriorly finely serrate.
Eyes not reaching the extremities of the orbits, terminated by
a minute spine. Chelipeds like those of 0. arenaria ; ambulatory
feet roughened by subspiniform granules.
Natal! (E. Wilson).
This species is closely allied to the Ocypoda arenaria of the
coasts of America, but is readily separated by the ocular spines
and the o-ranulations on the ambulatorv feet. It is dedicated to
m}^ friend, John A. Ryder, of the Academy of Natural Sciences of
PhiladeliDhia.
184 PROCEEDINGS OF THE ACADEMY OF [1880.
§ II. Eyes without an ocular spine.
9. 0, arenaria Sny.
Cancer arenarius Catesby, History of the Carolinas, ii, PI. 35 (1731
and 1771*). Cancer quadratus Fabr., Ent. Syst., ii, p. 439 (1793).
Ocypoda quadraia Fab., Suppl., p. 847 (1798); Bosc. Edit., 1, "i, p.
194;" (teste Auct.) Edit., II, i, p. 247 (1828); Latr., Hist. Crust, et '
Ins., vi, p. 49 (1803-4). Ocypoda arenaria Say, Jour. Acad. Nat.
Sci., Phila., i, p. 69 (1817); Edw., Hist. Nat. Crust., ii, p. 44, PI.
XIX, f. 13-14 (1837); DeKay, N. Y. Fauna, Crust., p. 13 (1841);
Gibbes, Proc. Am. Assoc, iii, p. 180 (1851); Gerstilcker, Arch, fiir
Naturg., xxii, p. 36 (1856 ; Guerin in La Sagra's Hist. Cuba, Crust.,
p. 7 (1857); Martens, Arch, fiir Naturges., xxxviii, p. 103 (1872);
Smith. U. S. Fish Coram. Report for 1871-72, p. 545 (1875); Kings-
ley, Proc. Phila. Acad., 1878, p. 322. Ocypoda albicans Latr., Encyc.
Meth., X, PI. 285, f. 1 (after Catesby vix 0. albicans Bosc). Ocy-
poda rhombea Edw., Hist. Nat. Crust., ii, p. 46 (1837); Ann. Sci,
Nat., Ill, xviii, p. 143 (1852); Gibbes, 1. c, p. 180 (1851); Dana,
U.S. Ex. Exp.Crust.,p.322,Pl.XIX,f.8(1852);Heller,ReiseNovara,
Crust., p. 42 (1867); Smith, Trans. Conn. Acad., ii, p. 135 (1870);
Streets, Proc. Phila. Acad., 1873, p. 240. Monolepis inermis Say,
1. c, p. ^57 (Megalops).
Carapax convex, gi'anulate above, front but little deflexed ;
lateral angles of carapax acute, extending as far forward as the
middle of the front. Lateral margin crenulate, arcuate, the
carapax being widest at the anterior third. Orbits below with
spiniform tubercles and occasionally a distinct emargination.
Eyes with the extremities rounded and not reaching to the
extremity of the orbit. Chelipeds with spines and tubercles ; the
meros with the upper and lower margins spined, the posterior
rounded and crossed by tubercular rugaj. Carpus with the
tubercles obsolete above but pronounced near the margins, the
inner margin armed with one or more strong spiniform teeth.
Hands tuberculate, the upper margin spined, the lower serrate.
A tubercular stridulating ridge on the inside near the fingers.
Fingers strongly inflexed. Ambulatory feet compressed, hairy,
their sides smooth, the upper portion margined and crossed by
transverse rugse.
o
* There were at least two editions of Catesby published, bearing dates
respectively as above and as the second is post-Linnean, the names employed
by him must hold. I think that another edition exists in the librai-y of
the Essex Institute at Salem, Mass., the date of which is between those
quoted above, but I have nothing at hand by which to decide the matter.
1880.] NATURAL SCIENCES OP PHILADELPHIA. 185
Young specimens (less than 10 mm. broad) have the lateral
angle further back than in the adult, while the spines of the
chelipeds are wanting or but faintly indicated.
The 0. albicans of Bosc, Lamarck and Desmarest has the
eyes terminated by a style, a feature I have never observed in
an}- specimen of 0. arenaria. The locality given is South Caro-
lina. The 0. rhombea of Fabricius is not this species, as his
expression " Carpus ut7'inque unidentatis, manibus sublaevis " will
not apply to this form, but agrees better with 0. forcUmana.
Fabricius gives no locality- for his specimens. The Brazilian
forms {rhombea Auct.) show no differences from northern speci-
mens. I haA-e seen specimens from over thirt}- localities embraced
in the limits of Great Egg Harbor, N. J. (Say\s types), to Rio
Janeiro, Brazil, and also specimens from the west coast of Mexico
(Dr. W. H. Jones).
10. 0. convexus Quoy and Gaimard.
Ocypoda convexus Quoy et Gaimard, Yoy. Uranie, Zool., iii, p. 535, PI.
LXX\^II, f. 2 (1828); Edw., Hist. Crust., ii, p. 49 (1837j.
Carapax granulate, sides arcuate, front deflexed, orbits sinuate,
lateral angles behind the base of the rostrum, acute. Meros
internall}' entire, distally tuberculate ; carpus tnberculate, its
inner surface with a bifid tubercle. Hands cordate, externally
granulate, serrate above and below.
This brief description is taken from the figure of MM. Quo}'
and Gaimard. I have never seen the species. It is said to have
come from Australia.
11. 0. cordimana Desm.
? Ocypoda rhombea Fabr., Suppl. Eut. Syst., p. 348 (1798). Ocypoda
cordimana Desm,, Consid. sur les Crustaces, p. 121 (1825); Edw.,
Hist. Nat. Crust., ii, p. 45 (1837); Ann. Sci. Nat. Ill, xviii, p. 143
(1852); Jacquinot et Lucas, Voy. Astrolabe et Zelee, p. 64; Heller,
Reise Novara Crust., p. 42 (1867); A. M. Edw., Nouv. Arch. Mus.,
ix, p. 271 (1872). Ocypoda rhombea ? Desmarest, 1. c. p. 122; Ran-
dall, Jour. Phila. Acad., viii, p. 123. Ocypoda pallidula Jacquinot
et Lucas, 1. c, PI. VI f. 4. Ocypoda laevis Dana, U. S. Expl. Exped.,
Crust., p. 325, PI. XX, f . 2 (1852). ? Ocypoda convexa Stm., Proc.
Phila. Acad., 1858, p. 100.
Carapax arcuate, evenh' granulate. Front strongly deflexed.
Orbits sinuate above ; lateral angles acute, but not extending as
far forward as the base of the front. Sides in the adult slightly
13
136 PROCEEDINGS OF THE ACADEMY OF [1880.
arcuate, but iu the young the}* are parallel or even concave, con-
verging behind. Eyes, without st3-liform process, and extending
nearly or quite to the orbital angle. Meros of chelipeds with its an-
terior margin crenulate in the young, in the adult with spinifonn
tubercles. Carpus externally granulate. Hand short, broad,
cordate, granulate internally and externally, its lower margin
serrate, the stridulating ridge nearly obsolete. Fingers short,
compressed, the thumb slightly hooked at the extremity. Meral
joints of the ambulatory feet with transverse rugae. Carpal and
propodal joints similarly roughened and covered with a short
pubescence.
New Zealand ! Mauritius ! (Guerin) ; Australia ! (E. "Wilson); Sand-
wich Is. ! (J. K. Townsend) ; Tahiti] (A. Garrett) ; Mozambique
and Zanzibar (Hilgendorf) ; Red Sea, Manilla, Nicobars (Heller) :
Hong Kong, Loo Choo (Stm. ) ; Japan (Edw.).
The following are not true members of the genus :
0. angulatus Latr. = Gonoplax angulatus.
0. aurantia Bosc. ex Herbst ^ Thelphusa aurantia.
0. carnifex Latr. ex Herbst = Cardiosoma carnifex.
0. hcterochelos Bosc. = Gelasimus heterochelos.
0. hispana Bosc. ex Herbst = Sesarma sp.
0. hyJrodromus Latr. ex Herbst == Thelphusa hydrodromus.
0. longimana Latr. = Gonoplax rhomboidalis.
0. viaracoani Latr. == Gelasimus maracoani.
0. macrocheles Bosc. == ? Macrophthalmus sp.
Q. pugillalor Bosc. = Gelasimus pug illat or.
O. quadrata Bosc. = Sesama sp.
0. rufopunctata Latr. ex Herbst = Trapezia rufopunctata.
0. senex Latr. ex Fabr. = Thelphusa sp.
0. teirago7ion Bosc. ex Herbst = Gelasimus tetragonon.
0. tridens Latr. ex Fabr. = ? Pachygrapsus sp,
0. vocans Latr. = Gelasimus sp.
I have not been able to identify
O. granulata Bosc. (Edit, ii) p. 247.
0. viacleayana Hess, Archiv. fiir Naturgesch., XXXI, p 143, PI. VI, f. S
(1865). Australia.
O. unispinosa Rafinesque, Precis de decouvertes Semiologiques, p. 21, No.
35 (1814).
1880.] NATURAL SCIENCES OF PHILADELPHIA. 18T
CAKCINOLOGICAL NOTES, No. IV.— SYNOPSIS OF THE GRAPSID.ffi:.
BY J. S. KINQSLEY.
The following paper is a continuation of my studies of the
Catometopa contained in the Museum of the Academj- of Natural
Sciences of Philadelphia. In it I have endeavored to embrace
every known species of the family with sufficient references to
their geographical distribution. To aid in the identifications of
species I have compiled anal3'tical tables for most of the genera,
but descriptions are given of onl}- those species of which I, have
examined specimens. I have reduced considerably the number
of nominal species, but believe that I am fully wan-anted in rele-
gating to synon3'my mau}^ so-called species founded on size, color,
geographical distribution, or variations of minor importance. All
localities from which I have examined specimens are marked with
an exclamation point (!). The classification employed is mainly
that of Dana, in the U. S. Exploring Expedition Crustacea (1852) ;
those of Milne-Edwards (Annales des Sciences Naturelles, III
Series, Zoologie, tome xx, pp. 163-200, 1853), and Kossmann
(Zoologische Ergebnisse, Reise in die Kiistengebiete des rothen
Meeres, 18YT), being comparatively'- worthless. Owing to the
limited amount of space at my disposal, the synonymy and bibli-
ography have been condensed as far as possible.
Family GRAPSID.31 Dana. {Ora-psoidiem M. Edw.).
Carapax subquadrate, depressed. Front generally broad. Eyes
short. Antennulge transversely plicate. Epistome short, some-
times linear. Meros of the external maxillipeds bearing the palpus
at the summit or at its external angle. Second joint of the abdo-
men of the male nearly as wide as the adjacent portion of the
sternum.
The Orapsidee are all inhabitants of the temperate or tropical
waters, and generally \i\Q near the shores. A few, however (e.g.
Nautilograpsus and Varuna)^ live on the high seas. The family-
may conveniently be di^nded into two^ sub-families, by characters
derived from the antennae. In the Grapsinse the antennae are
' The characters given by Dana for the Sesarraiuae I do not consider of
sufficient importance to warrant its retention as a sub family, and would
rather consider it as a group of the Grapsinse.
188 PROCEEDINGS OF THE ACADEMY OP [1880.
covered by the front ; in the Plagusinse they are received into
notches in the front and are visible from above.
Analytical Key to the Genera of the Grapsidse.
Antennse covered by the front. GRAPSIN^.
External maxillipeds without a piliferous ridge. Grapsini.
External maxillipeds widely gaping.
Antennse excluded from the orbit.
Meros of the external maxillipeds as long as the ischium. Goniopsis.
Meros of the external maxillipeds shorter than the ischium.
Metopograpsus.
Antennae entering the orbit.
Front less than half the width of the carapax.
No tooth behind the orbital angle. Epigrapsus.
One tooth behind the orbital angle.
Fingers of cheliped excavate. Grapsus.
Fingers acuminate.
Sides straight. Orthograpsus.
Sides arcuate in front. Geograpsus.
Two teeth behind the orbital angle. Leptograpsus.
More than two teeth behind the angle of the orbit.
Male abdomen five-jointed, anteunulse transverse, front
straight. Grapsodes.
Male abdomen seven-jointed, antennulse oblique, front exca-
vate . Cyrtograpsus.
Front more than half the width of the carapax.
Carapax transverse. Paehy grapsus.
Carapax longer than broad. Nautilograpsus.
External maxillipeds without a rhomboidal gape.
Palpus articulating with the inner angle of the meros of the external
maxillipeds.
One tooth behind the orbital angle. Br acliy grapsus.
Several teeth behind the orbital angle. EucMrograpsus.
Palpus articulating with middle of the anterior margin of the meros
of the external maxillipeds.
Exognath of external maxillipeds very broad. PtycJiognaihus.
Exognath much narrower than the ischium.
Sides of carapax entire. Acmceopleura.
Sides of carapax dentate.
External distal angle of the meros of the external maxilli-
peds expanded.
Joints of ambulatory feet rounded, dactyli styliform.
Pseudograpsus.
Ambulatory feet compressed, dactyli flattened, natato-
rial. Varuna.
Meros of external maxillipeds not expanded.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 189
Meros of external maxillipeds broader than long.
Meros as long as ischium. Glyptogra'psus.
Meros shorter than ischium. Utica.
Meros as long or longer than broad.
Front nearly half as wide as carapax. Heterograpsus.
Front not over one-third as wide as carapax. Eriocheir.
Palpus articulating with the outer angle of the meros of the ex-
ternal maxilliped.
One tooth behind the orbital angle. Perig7'(ipsus.
More than one tooth behind the orbital angle. Platygrapsvs.
External maxillipeds with an oblique piliferous ridge. Sesarmini.
Meros of external maxilliped elongate, its apex rounded.
Antennae excluded from the orbit. Metasesdrma.
Antennae not excluded from the orbit.
Carapax subquadrate, sides ai'cuate.
Joints of ambulatoiy feet entire. Sarmatium.
Joints of ambulatory feet dentate. Bhaconotus.
Carapax quadrate, sides straight. Sesarma.
Carapax elongate, narrowed behind. Aratus.
Mei'os of external maxilliped short, its distal border truncate or even
excavate, and bearing the palpus.
Antennae excluded from the orbit. Clistocoeloma.
Antennae entering the orbit.
Sides of carapax straight. Helice.
Sides arcuate.
Sides entire. Cyclograpsns.
Sides emarginate or toothed. Chasmagnathus.
Antennae lodged in notches in the front, and visible from above.
PLAGUSm^.
Meros of external maxillipeds large, as broad as ischium. Plagi/sia.
Meros small and much narrower than ischium. Leiolophus.
Sub-family Grapsinse {Grapsinm et Sesarmince Dana).
Antennulse more or less transverse, and covered by the front.
Tribe GRAPSINI (Sub-family Grapsinse Dana).
§ External maxillipeds -without an oblique piliferous ridge on
the ischial and meral joints.
Genus GONIOPSIS De Haan, 1835 (Goniograpsus (pars) Dana, 1851).
Carapax flat ; front vertical, over half as wide as carapax ;
sides straight, one-toothed. Suborbital lobe broad, reaching the
front and excluding the antenna from the orbit. External max-
illipeds slender ; meral and ischial joints of equal length.
190 PROCEEDINGS OF THE ACADEMY OP [1880,
1. G. cruentatus De Hann ex Latreille.
Cancer rvricola De Geer. Memoirs pour sei-vir a I'Hist. Insectes, vii,
417, PL XXV, 1778 (non Linne).
Orapsxis criientattts Latreille. Histcrire Naturelle des Crust, et Ins.,
vi, p. 70 (1803-4).
Goniopsis cruentatus I>e Haan. Fauna .Japonica Crust., p. 33 (1835).
Grapsus longipes Randall. Jour. Pliila. Acad., viii, p. 125 (1839).
Goniopsis ruricola White. List Brit. Mus. Crust., p. 40 (1847).
'Grajysus pelU Herklots. Additamenta ad Faunam Carcinologicam
AfricjB Occidentalis, 8, PI. I, f. 6-7 (1851).
'Goniograpsus cruentatus Dana. U. S. Expl. Exped. Crust., p. 342,
PL XXI, f. 7 (1852).
Front granulate, supra-frontal lobes four, margins crenulate,
oi'bits entire above, distally emarginate. Carapax with oblique
transverse ridges. Anterior margin of nieros of chelipeds ex-
panded, dentate, the upper and lower margins with spiniform
tubercles, as is also the upper margin of carpus. Hands with
spiniform tubercles above and below, the middle of the outer sur-
face smooth, the inner surface with scattered prominent granules.
Thumb and finger sub-excavate, the latter spinose above. Ambu-
latory feet compressed and armed with still' black bristles. Pos-
terior angle of meros of last pair rounded, in the other feet
dentate.
Floridti.! (H. E. Webster, in Union College Museum) ; Bahamas!
Cuba! (H.F.Baker); Surmam! (Dr. Hering, Randall's type of
G. longipes) ; Gaboon, W. Africa (Du Chaillu) ; West Coast of
Nicaragtia! (J. A. McNiel, in Museum of Peabody Academy);
Tropical Seas of America (Auct, ),
Genus METOPOGRAPSUS M.-Edw., 1858.
Front more than half the width of carapax, deflexed. Sides
straight. Internal suborbital lobe very broad, reaching the front
and excluding the antenna from the orbit. Meros of external
maxilliped short, much broader than long.
Key to Species.
Antero-lateral margin entire.
Frontal margin sinuate. messor.
Frontal margin straight. latifrons.
Antero-lateral margin toothed. oceanicus.
M. messor Edwards ex ForskaL
Cancer messor ForskaL Descr. An. in Itin. Observ., p. 88 (1775).
Grapsus gaimardii Andouin, Expl. PL Savigny (teste Edw.).
Grctpsus messor ^Avf ■ Hist. Nat;" Crust., ii, p. 88 (1837).
1880.] NATURAL SCIENCES OF PHILADELPHIA, 191
Grapsus thukuJiar Owen. In Beechey's Voyage, Zoology, p. 80, Tl.
XXIV, f. 3 (1839).
Graps>is paniUelus Randall. .Jour. Acad. Nat. Sci., Phila. viii, p.
127 (1839).
Metopograpsus messor, timkuliar, eydouxi et intermedius Edw. Ann.
Sci. Nat., Ill, XX, p. 165 (1853).
Carapax slightly narrowed behind, plications and rugse more
or less distinct. Frontal lobes rather prominent, frontal margin
sinuate, smooth or crenulate at the angles. Meros of chelipeds
with the posterior surfiice rugose, the anterior margin expanded
and distally truncate, a few spinose teeth near the base and
several on the truncate margin. Carpus externall}' rugose, inter-
nallj' with a prominent bifid or quadrifid tubercle. Hands with
oblique folds above and below, and a longitudinal ridge on the
lower outer surface. Fingers sub-excavate. Last joint of male
abdomen but slightly narrower than penult joint.
Sandwich Is. ! (Nuttall, Pease, Jones, Wilkes' Expedition); Tahiti!
(A.J. Garrett); Avstralia / (E. Wilson); Mauritius/ (Guerin);
Aden; ! Natal! (Dr. T. B. AVilson) ; Indian and Pacific Oceans
(Auct.)
M. latifrons Edwards ex White.
Oi'apsus latifrons White, in Jukes' Voyage of the Fly, ii, p. 337, PI. II,
f. 2 (1847j.
Metopograpsus latifrens et maculatus'Eidw., Ann. Sci. Nat. Ill, xx, pp.
166 and 165, PI. VII, f. 1 (1853).
Metopograpsus pictus A, M.-Edw., Ann. Soc. Ent. France, vii, p. 283
(1867) ; Nouv. Arch, du Mus., ix, p. 289, PI. XIII, f. 2 (1873).
Carapax narrowed behind, plications indistinct ; frontal lobes
granulate on the edge. Front broad, nearly straight, mai'gin
denticulate. Chelipeds similar to those of M. messor. Base of
last joint of male abdomen much narrower than the extremity of
penult joint.
Batavia, ! (Dr. "Wilson); Singapore (White); Java (Edw); New Cale-
donia (A. M.-Edw.).
M. oceanicus Jacq. et Lucas.
Metopograpsus (Grapsus) oceanicus Jacqiiinot et Lucas, Voyage Astro-
labe et Zeelee, Crust., p. 73, PI. VI, f. 9 (Text 1853, Plates 1842-53).
Metopograpsus quadridentatus Stimpson, Proc. Acad. Nat. Sci. Philadel-
phia, 1858, p. 102.
Pulo Han (J. et L.); near Hong Kong (Stm.) ; Nicobar Is. (Heller).
192 ' PROCEEDINGS OF THE ACADEMY OF [1880.
Geuus EPIGEAPSUS Ileller, 1S621 {Nedograpsus Heller, 1865).
Sides arcuate, entire orbits externally open ; external maxilli-
peds gaping, meros shorter than the ischium, rather broad abdo-
men of the male five-, of the female seven-jointed.
E. politus Helltr.
Epigrapsus politus Heller, Verb. Z. B, Ges. Wien, 1862, p. 522.
Nectograpsus politus Heller, Reise der Novara, Crustaceen, p. 57, PI. V,
f. a (1805).
Nicobars ; T«7i^<^■ (Heller).
Genus GEAPStJS Lamarck (restrict).
Sides arcuate, with one tooth behind the orbital angle; front
narrow, deflexed ; antennae entering the orbit. External maxilli-
peds slender, gaping ; meros oblong. Fingers of chelipeds exca-
vate.
G. maculatus Edwards ex Catesby.
Pagttrus maculatus Catesby, Nat. Hist, of the Carolinas, ii, Plate
XXXVI, f. 1 (1743 et 1771).^
^ Dr. Heller, in his preliminary account of the Crustacea collected by
the Novara Expedition (Verbandl. Zool. Bot. Gesellscbaft, Wien, 1862, pp.
519-528), characterizes several genera and species of which no mention is
made in the final report. These are MenmtJiius brevirostris which is prob-
ably Acantlionyx consobrinus A. Edw. , of the final report ; Xantho tetraodon
is Eudora tetraodon, Carpilodes granulatus appears to be C. tristis Dana.
Lupa Mrsuta was probably referred to Neptunus sanguinolentus. Thel-
phusa willlerstorfl appears to be /. leschenmidii, Parathelphusa dentipes to be
P. tridentata, Eelcecius areolatus to be H. cordiformis, Metaplax Jiirtipes is
apparently referred to a new genus, Orapsus depressus is probably Oeo-
grapsus crinipes, Orapsus dedivifrons is apparently rechristened Pachygrap-
sus iniermedius, Eeterograpsus barbigerus has its specific name altered
to barhimanus, Epigrapsus nov. gen. reappears as Nectograpsus nov.
gen. with no reason assigned for the change. Metasesarma granulatus is
redescribed as new under the name rugulosa. Plagusetes elatus is prob-
ably, as pointed out by Mr. Miers (Ann. and Mag. Nat. Hist., ix, p. 147,
1878), Acuntliocydus gayiof the final report. Oelas^m^ls vcmegatus appears
to have been finally referred to G. annulipes. Palinurus paulensis was
afterward apparently referred to P. lalandii and Pelias notatus is refen-ed
to Anchistia. All this shuffling of names is made without the slightest hint
to aid one in correlating the two papers, and is a proceeding which cannot
be too strongly condemned.
- There were at least two editions of Catesby bearing dates as above,
and as the second appeared thirteen years after the tenth edition of the
Syslema Natures oiJAmw, and five after the twelfth edition, the names em-
ployed by Catesby will hold.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 193
Cancer grapsus Liim., Syst. Nat., Edit., x, p. 630 (1758).
Qrapsus pictus Latieille, Hist. Crust, etiiis., vi, p. 69, PI. XL VII, f. 3
(1803-4.)
Goniopsis pictus De Haan, Fauna Japonica Crust., p. 33 (1835).
Grapsus strigosus Brulle, in Webb et Berthelot Hist Canaries, ii, PI.
II ; Crustacea, p. 15, 1836-44 (teste'Edw.)'
Grapsus maculatus, webbi, ornaius et pharaonis Edw., Ann. Sci. Nat.
pp. 167-8, PI. VI, f. 1 (1853).
Grapsus aUifrons Stimpson, Annals N. Y. Lyceum Nat. Hist., vii, p.
230 (1860).
Carapax depressed, transversel}^ plicate, folds anteriorly broken
lip into squamiform tubercles. Frontal crest four-lobed, median
lobes the larger, their margins subtuberculate. Frontal margins
crenulate, regularly arcuate. Lateral margin arcuate. Inferior
border of orbit with a deep fissure. Anterior border of ischium
and meros of cheliped spinose, the lower margin of the meros spino-
tuberculate, the posterior surface plicate. Carpus with distant
tubercles, its interior margin with a laminate spine. Hand above
tuberculate, externally with longitudinal ridges, below with
oblique folds. On the inner surface the tubercles and folds are less
prominent. Fingers short, tips excavate. Ambulatory feet com-
pressed, propodal and dactylic joint spinose.
Florida Keys! (Webster, Ashmead) ; West Indies/ (Lawrence,
Wood, Wilson, Goes, Lea) ; San Lorenzo I (Wilkes' Expedition) ;
Pernambuco! (Dr. Wilson); Tahiti! (A. Garrett); W. Coast
Mexico! (Dr. Jones); Central America I (McNiel) ; New Zealand!
(Dr. Wilson); Mauritius! (Guerin) ; Natal! (Dr. Wilson);
Georgia, California, Peru, St. Helena and Cape Verde Is. (Miers) ;
Paumotu and Hawaian Is. (Dana) ; Honduras! (no collector's
name).
The genus Graj)sus, as well as several others, is divided into
sections by Milne-Edwards, characterized either by having the
posterior distal angle of the meros of the last pair of ambulatory
feet regularly rounded, or dentate ; but in specimens of G. macu-
latus., I have occasionally found this angle on one side entire, and
the other dentate.
G. maculatus var. tenuicristatus Alartens ex Herbst.
Cancer' tenuicristatus Herbst. Krabbeu und Krebse, PI. Ill, f. 33-34,
1790 (teste Martens).
Grapsus rudis Edw., Hist. Nat. Crust., il p. 87 (1837).
Grapsus hirtus Randall, Jour. Phila. Acad., viii, p. 124 (1839).
^ Brulle gives not the slightest description which will distinguish his
specimens from either maculatus or strigosus.
194 PROCEEDINGS OF THE ACADEMY OF [1880.
Is distinguished from the typical forms of G. maculatus only
by the hairy carapax and nieral joints of the ambulatory limbs,
and the narrower carpal spine. All other characters Avhich have
been given prove inconstant. Dr. Martens, by an examination of
Herbst's type, has shown the identity of Herbst's teniiicriatafus,
and Edwards' rudis.
Hfiwaian Is.! (J. K. Townseiid, Randall's type) ; OoJiu! (Dr. W.
H. Jones) ; Ceylon (Heller) ; Bonin (Stm.).
G. Strigosus Latreille ex Herbst.
Cancer strigosus Herbst, PL XLVII, f. 7 (17991.
Orapsus strigosus Latr., Hist. Crust, et Ins., vi, p. 70 (1803-4).
Grapsus alboUneatus Lamarck, Hist. Animaux sans Vertebres, v, p.
249(1817).
Ooniopsis strigosus De Haan, Fauna Japonica Crust., p. 33 (1835).
Grcqysus granrtlosus, peronii et pelagicus Edw., Ann. Sci. Nat., Ill,
XX. p. 169 (1853). .
Grapsus longipes et suhquadratus Stimpson, Proc. Acad. Nat. Sci.
Philadelphia, 1858, pp. 102 et 103.
Carapax but little convex, posteriori}' with oblique transverse
lines, anteriorly with sqnamiform tubercles. Frontal lobes mod-
erate, frontal margin crenulate. Orbits with a deep fissure below.
Epistome short. Meros spined anteriorly, its other margins
rounded, the posterior surface with transverse rugae- Carpus
granulate and bearing internally a slender spine. Hands much
as in G. maculatus. Posterior distal angle of meros of last pair
of ambulatory feet denticulate.
Natal! Australia! (Dr. T. B. Wilson) ; Sandtdch Is.! (in Peabody
Acad.). Its distribution is embraced within the above limits and
Hong Kong (Stimpson).
G. gracilipes Milne-Edwards.
Grapsus gracilipes Edw., Ann. Sci. Nat., Ill, xx, p. 168 (1853).
China (Edwards).
Subgenus Orthograpsus, nov.
Carapax transverse, broadest behind. Sides straight, with one
tooth behind the orbital angle. Antennas entering the orbit.
Fingers of chelipeds acute.
0. hillii nov.
Carapax depressed, plications faint. Supra-frontal lobes mod-
erate ; front straight, narrow, deflexed. Sides of carapax straight,
post-orbital tooth small. Meros of external maxilliped a little
1880.] NATURAL SCIENCES OF PHILADELPHIA. 195
longer than broad. Chelipeds much as in Metopograpsus onessor-,
the hand granulate above, fingers acuminate. Posterior distal
angle of meral joints of ambulator}^ feet rounded, entire or finely
serrate, there being a variation in the sides of the same specimen.
West Indies ! (Dr. Wilson) ; Key West, Fla. ! (Dr. A. S. Packard, Jr.,
Peab. Acad).
0. longitarsis Kingslcy ex Dana.
Grapsus longitavsis Dana, Proc. Acad. Nat. Sci., Phila., 1851, p. 249.
U. S. Expl. Exped. Crust., p. 339, PI. XXI, f. 4 (1852).
Pmmiotu Archipelago (Dana).
Genus GEOGEAPSTJS Stimpson, 1868 (Biscoplax, Am. Ed., 1867).
Carapax depressed, sides curved in front, straight behind, one
tooth behind the angle of the orbit. Front narrow, strongly de-
flexed. Internal suborbital lobe large. Antennae entering the
orbit. Dactyli of chelipeds acuminate.
Synopsis of Species.
Meros of chelipeds with a laminiform expansion of the anterior margin.
Front nearly straight.
Folds of carapax transverse. Udidus.
Folds of carapax oblique. gvayi.
Front arcuate. crinipes,
Meros not expanded, carapax tuberculate anteriorly. longipes.
G. lividus Stimpson ex Milne-Edwards.
Orapsus lividus Edwards, Hist. Nat. Crust., ii, p. 85 (1837).
Grapsus brevipes Edw., Ann. Sci. Nat., Ill, xx, p. 170 (1853).
Geograpsus litidus Stimpson, Proc. Acad. Nat. Sci. Philadelphia,
1858, p. 101.
Geograpsus occidentalis Stimpson, Annals N. Y. Lye, vii, p. 230
(1860).
Carapax much broader than long, depressed. Plications nearly
transverse. Frontal lobes prominent, front deflexed, its margin
nearly straight. Sides of carapax slightly arcuate. Orbit with a
deep fissure below. Meros of chelipeds above and below with
transverse rugae, its anterior margin expanded, proxiraall}^ den-
ticulate, distall}' the teeth are larger. Carpus granulate and with
a short spine on the inner margin. Hand and dactylus tubercu-
late above, externally and below with short, oblique rugae ; fingers
acuminate. Distal angle of meral joints of the last pair of ambu-
latory feet rounded.
Isle Bartholomeio, W. I.! (A. Goes) ; Chili! (Guerin) ; West Indies
(Auct.) ; Cape St. Lucas (Stm.).
196 PROCEEDINGS OF THE ACADEMY OF [1880.
G, criuipes Stimpson ex Dana.
Grapsus crinipes Dana, Proc. Acad. Nat. Sci, Philadelphia, 1851, p.
249. U. S. Expl. Exped. Crust., p. 341, PI. XXI, f. 6 (1852).
Geograpsus crinipes Stimpson, Proc. Acad. Nat. Sci. Philadelphia,
1858, p. 101.
Grapsus depressus Heller, Verh. Z. B. Gess. Wien, 1862, p. 521.
Carapax depressed, the sides nearly parallel, folds of the cara-
pax oblique, frontal lobes but little prominent, front arcuate.
Ischial joint of cheliped spined in front ; meros with the anterior
margins expanded, finely serrate proximally, more coarsely so at
the apex ; carpus and hand roughened above, a few inconspicuous
lines on the lower outer surface of the palm. Distal angle of
meros of the last pair of ambulatory feet rounded.
Sandioich Is. ! (Dr. W. H. JonesJ ; Tahiti (Heller).
G. grayi A. Milne-Edwards ex H. Milne-Edwards.
Grapsus grayi Edw., Ann. Sci. Nat., Ill, xx, p. 170, 1853.
Geograpsus ruhidus Stimpson, Proc. Acad. Nat. Sci, Philadelphia,
1858, p. 103.
Geograpsus grayi A. M.-Edw., Nouv. Arch, du Mus., ix, p. 288 (1873).
Carapax somewhat inflated, its folds oblique ; frontal lobes
prominent, front nearl}^ straight. Orbit with a slight fissure
below ; chelipeds much as in G. ciHnipes. Distal angle of meros
of last pair of feet rounded, entire or dentate.
This is probabl}' the adult of the preceding species.
Tahiti! (A. Garrett); Australia, Mauritius, Zanzibar (Hilgen-
dorf) ; Madagascar, India, Bonin, New Caledonia (A. M.-Edw.).
G. longipes Kingsley ex A. Milne-Edwards.
Discoplax longipes A. M.-Edw., Ann. Soc. Ent. France, vii, p. 284
(1867). Nouv. Arch, du Mus., ix, p. 294, PI. XV (1873).
New Caledonia (A. M.-Edw.).
Genus LEPTOGRAPSTJS M. -Edwards (pars), 1853, Stimpson.
Carapax with the sides arcuate, two-toothed. Front less than
half the width of the carapax, not deflexed. Internal sub-orbital
^ lobe small, antennae entering the orbit. Meros of external max-
illipeds as broad as long, but shorter than the ischium.
Leptograpsus variegatus Milne Edwards ex Fabricius.
Cancer varkgatus Fabr., Ent. Syst., ii, p. 450 (1793).
Grapsus marginatus Latr., Hist. Crust, et Ins., vi, p. 71 (1803-4).
Grapsus personatus Lamarck, Hist. An. sans Vert., v, 249 (1817).
Grapsus jyictus Quoy et Gaimai'd, Voyage Uranie et Physiciene, p.
523, PI. LXXVI, f. 2 (1824).
1880.] NATURAL SCIENCES OF PHILADELPHIA. 191
Grapsvs strigillatus White, in Gray's Zoological Miscellany, p. 78
(1843).
Grapsus variegatus Edwards et Lucas, in d'Orbigny's Voyage, p. 27
(1849).
Grapsus planifrons Dana, Proc. Acad. Nat. Sci. Philadelphia, 1851,
p. 249. U. S. Expl. Exped. Crust., p. 638, PL XXII, f. 3 (1852).
Leptograpsus variegatus Edw., Ann. Sci. Nat., Ill, xx, p. 171 (1853).
Leptogiapsus hertheloti, xerreauxi, ansoni et gai/i Edw., 1. c, p. 172
(1853).
Carapax nearl}' flat, transversely plicate. Protogastric region
concave, with squamose tubercles, protogastric lobes but little
prominent. Front slightl}^ depressed, its margin crenulate and
nearly straight. Orbits with a narrow, deep, external fissure.
Meros of chelipeds with the anterior border expanded, dentate ;
the other angles rounded, the posterior surface rugose. Carpus
tuberculate and with a short spine on the internal surface. Hand
tuberculate above, externall}'' smooth. In the 3'oung there is an
elevated line along the outside of the palm. Ambulatory feet
with stiff setae.
Pernambuco / (Dr. Wilson); Chili/ (Wilkes' Expedition) ; Austra-
lia / (E. Wilson); JVeic So. Wales! (Capt. Putnam, Peabody
Academy) ; Isle Guam (Quoy and Gaimard) ; Canaries (Edw.);
Norfolk I. (Miers) ; Shanghai (Heller).
Genus GEAPSOPES Heller, 1865.
Carapax depressed, sides arcuate and dentate in front, behind
straight. Front less than half the width of the carapax, strongly
deflexed. Orbits externally open. Internal sub-orbital lobe
small, antennae entering the orbit. Meros of external maxilliped
longer than broad. Male abdomen five-jointed.
G. notatus Heller.
Grapsodes notatus Heller, Novara Crust., p. 58, PI. V, f. 2 (1865).
Nieobars (Heller).
Genus CYETOGRAPSUS Dana, 1851.1
■ Carapax broader than long, front narrow, excavate, sides arcu-
ate, with three teeth behind the orbital angle. External maxilli-
peds widely gaping, without a piliferous ridge. Epistome very
1 For some reason, Prof. Smith in his paper on Brazilian Crustacea
(Transactions of the Connecticut Academy of Arts and Sciences, Vol. ii,
pp. 1-42, 1869), and in his notes on Ocypodoidea (1. c, p. 154), refers
several times to this genus, and always as Cryptograpsus.
198 PROCEEDINGS OF THE ACADEMY OF [1880.
short. AnteniiiB entering the oi'bit. Male abdomen seven-
jointed, the second joint verj^ short.
*
C. angulatas Dana.
Cyrtograpsus angulatus Dana, Proc. Acad. Nat. Sci. Phila., 1851, p. •
250. U. S. Expl. Exped., p. 352, PI. 22, f. 6 (1852).
Cryptograpsus angulatus Smith, Trans. Conn. Acad., ii, p. 37, 1869.
Carapax uneven, granulate. Sides three-toothed, second tooth
small. Orbits with a slight fissure above. Feet all granulate.
Hands inflated; fingers acuminate.
Rio Negro, Patagonia! (U. S. Expl. Exped.).
C. cirripes Kingsley ex Smith.
Cryptograpsus cirripes Smith, Trans. Conn. Acad, ii, p. 11, PI. I, f. 3
(1869). .
Carapax depressed, areolate ; front narrow, slightly excavate.
Sides of carapax strongl}'^ arcuate, with four teeth behind the angle
of the orbit, the second and last teeth much smaller than the
others ; all of the borders of the carapax are crenulate. Chelipeds
stout, granular. Propodal joints of first, second, and fourth, and
dactylus and carpus of fourth pair of ambulatory feet haired.
Bio Janeiro! (Captain Harrington, Peabody Academy of Science,
Salem, Mass., types).
Genus PACHYGRAPSUS Randall (1839); Stimpson (1858).
Carapax somewhat narrowed behind, and with transverse striae-.
Front more than half the width of the carapax, sides entire, or
with one or two teeth ; inner sub-orbital lobe small, allowing the
antennae to enter the orbit. External maxillipeds widelj' gaping,
meros as broad as long. Type, P. crassipes.
Synopsis of Species.
Sides entire.
Front straight or nearly so.
Numerous transverse folds on carapax ; lower margin of hand spined.
corrugat^ls.
Carapax but little plicate, hands smooth below. mthiopic^is.
Front strongly sinuate.
Hands smooth. minutus.
Hands externally with longitudinal ridges. j)^'^'^'^^'^^-
Sides with one tooth behind the orbital angle.
Posterior distal angle of meros of fifth pair of feet rounded.
Front with a prominent tooth at angle. crassipes.
Front slightly sinuate without prominent teeth. maurtis.
Posterior distal angle of meros of fifth feet dentate.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 199
Fingers of cheliped smooth. transversus.
Fingers dentate or spined above. gracilis.
Bides two-toothed.
Transverse lines of carapax naked. marmoratus.
Transverse lines of earapax haired. pubeseens.
Unknown to me. latipes.
P. crassipes Randall.
Paehygrapms crassipes Randall, Jour. Acad. Nat. Sci. Philadelphia,
viii,p. 127 (1839).
Grapsus eydouxi ^dw., Ann. Sci. Nat., Ill, xx, p. 170 (1853).
Leptograpsus gonagrus Edw., L c, p. 173 (1853).
Carapax somewhat arcuate, sides with a single tooth behind the
orbital angle ; frontal lobes prominent ; front deflexed, its margin
nearly straight, the angles with a prominent tooth. Meros of
Ghelipeds with the anterior margin produced, distally truncate and
dentate. Hands inflated, margined above and with a longitudinal
ridge on the lower outer surface ; fingers excavate. Distal angle
of meros of posterior ambulatory feet roimded ; dactyli of the
ambulatory feet spinulose.
(?) Sandwich Is. ! (T. Nuttall, Randall's type); California from San
Francisco! to San Diego ! (Many coUectoi's) ; ? New Providence,
W. I. ! (H. C. Wood, Jr.); Tolcohama (Tozzetti).
P. maarus Lucas.
P achy grapsus maurus Lucas, Expl. Algiers, Crust., p. 20, PL II, f. 5
1849).
Ooniograpsus simplex Dana, Proc. Acad. Nat. Sci. Phila., 1851, p.
24'J ; U. S. Expl. Exped. Crust, p. 344, PI. XXXI, f. 8 (1852).
Pachygrapsus simplex Stimpson, Proc. Acad. Nat. Sci. Phila., 1858,
p. 102.
Algiers (Lucas) ; Madeira (Dana); Bio Janeiro (Dana, Heller).
P. transversus Gibbes.
Pachygrapsus transversus Gibbes, Proc. Am. Assoc. Adv. Science, iii,
p. 182 (1850).
Ooniograpsus innotatus Dana, Proc. Acad. Nat. Sci. Phila., 1851, i>.
249 ; U. S. Expl. Exped. Crust., p. 345, PI. XXI, f. 9 (1852).
Leptograpsus rugulosus Edw., Ann. Sci. Nat. Ill, xx, p. 172 (1853).
Pachygrapsus lavimanus Stimpson, Proc. Phila. Acad., 1858, p. 102.
Metopograpsus duhiusetminiatus, Saussure, Mem. Soc. Phys, etd'Hist.
Nat. Geneve, xiv, pp. 444-445, PI. II, f. 16, 17 (1858).
Grapsus declivifrons Heller, Verhandl. Z. B. Gesellschaft, Wien, 1862,
p. 521.
Pachygrapsus intcrmedius Heller, Novara Crust, p. 44 (1865).
Pachygrapsus socius Stm., Ann. N. Y. Lye. x, p. 114 (1871).
Pachygrapsus advena Catta Ann. Sci. Nat. VI, iii, No. 1. p. 7, PI. I
(1876).
200 • PROCEEDINGS OF THE ACADEMY OP [1880.
Carapax depressed, shining, witli transverse plicae, oblique on
the branchial region. Sides generally slightly arcuate, with one
tooth behind the orbital angle. Frontal lobes prominent, front
sinuate. Meros of chelipeds with transverse rugse, the inner
margin dentate ; carpus rugose, with an internal rounded tubercle.
Hand minutely granulate, a longitudinal ridge on the lower outer
surface, margins rounded; dactylus with the upper margin smooth.
Postero-distal angle of the meros of last pair of ambulatory feet
dentate.
Florida! {K.. S. Packard; Peab. Acad.; Brown Univ.; H. E. Web-
ster, Union College) ; West Indies! (S. G. Morton, H. C. Wood,
Jr.); Brazil! (Dr. T. B. Wilson); California! (J. L. Leconte);
New Zealand ! (E. Wilson); Tahiti {h.. Garrett); W. Coast Nicar-
agua! (J. A. McNiel, Peab. Acad.); Australia (Stm.); Mio Janeiro
(Heller); Madeira (Stm.).
P. gracilis Stimpson ex Saussure.
Metopograpsus gracilis Saussure, 1. c, p. 443, PI. II, f, 15 (1858)
Orapsus guadulpensis Desbonne et Schramm, Crustaces de la Guada-
loupe, p. 48 (1867).
Pachygrapsus gracilis Stimpson, Ann. N. Y. Lye, x, p. 113 (1871).
Orapsus {Leptograpsus) rugulosus Martens, Archiv fiir Naturges-
cliichte, xxxviii, p. 102 (1873).
Carapax much as in P. transversus, but with no folds on the
cardiac region; lateral margins nearly straight, one-toothed.
Frontal lobes nearly obsolete ; front nearly horizontal, regularly
arcuate and minutely crenulate. Chelipeds and ambulatory feet
nearly as in P. transversus, the hand and dactylus, however, being
spined or toothed above.
Florida! {A. S. Packard, Jr., Peab. Acad.); West Indies (Auct.).
P. corrugatus Kingsley ex Martens.
Orapsus {Leptograpsus) corrugatus Martens, 1. c, p. 107, PI. IV, f . 8
(1872;.
Cuia (Martens).
P. aetMopicus Hilgendorf.
Orapsus {Pachygrapsus) a'thiopicus Hilgendorf, in von der Decken's
Reisen in Ost-Afrika, Crust., p. 88, PI. IV, f. 2 (1869).
Ugurunga, Fast Africa (Hilgendorf).
P. plicatus Stimpson ex Milac-Edwards.
Orapsus plicatus Edwards, Hist. Nat. Crust., ii, p. 89 (1837).
Orapsus A;rff?<sw Edwards, Ann. Sci, Nat. HI, xx, p. 170 (1853).
Pachygrapsus plicatus Stimpson, Proc. Acad. Nat. Sci. Phila., p. 102
(1858).
Pachygrapsus striatus A. M.-Edw., Journal Museum Godeflfroy, iv. p.
82 (1873).
1889.] NATURAL SCIENCES OF PHILADELPHIA. 201
Carapax broader than long, everywhere crossed b}' plications
which are bordered by short hairs ; frontal lobes prominent, front
sinuate. Sides of carapax entire. Meros and carpus of chelipeds
externally' plicate, inner margin of meros expanded, proximally
denticulate distall^- with spiniform teeth. Carpus Avith a promi-
nent internal spine. Hand and dactjdus granulate above, exter-
nally the hand bears several longitudinal rugie. Fingers short,
gaping, extremities excavate.
Oahu! (Dr. W. H. Jones); Tafdti! (A. Garrett, Peab. Acad.); New
Caledonia; Samoan Is. (A. M.-Edw.j^ Natal (Krauss); Loo Choo
(Stimpson).
P. marmoratus Stimpson ex Fabriciup.
Cancer marmoratus Fabrieius, Ent. Syst., ii, p. 4.')0 (1793).
Grapsus varius Latreille, Hist. Crust, et Ins. vi, p. 69 (180;3-4i.
Grapsus marmoratus Desmorest, Considerations, p. 131 (1825).
Leptograp«us marmoratus Edw., Ann. Sci. Nat. Ill, xx, p. 171 (1853).
Pachygrapsus marmoratus Stimpson, Proc Acad. Nat. Sci. Pliila.,
1858, p. 102.
Carapax depressed, naked, transversely plicate; frontal lobes
prominent, front depressed, slightlv arcuate, or sometimes a
little sinuate. Sides with two teeth behind the angle of the
orbit. Meros of chelipeds expanded in front, expansion distally
truncate and dentate. Carpus and hand tuberculate above, the
former with a prominent internal tooth. Fingers slightly exca-
vate. Posterior distal angle of meros of last pair of feet rounded,
entire.
France!. (Guerin) ; Bospliorus ! (Smithsonian) ; Mediterranean ( Auct. ) ;
Madeira (Stm.).
P. pubescens Heller.
Pachygrapsus pubescens Heller, Novara Crust, p. 45, PI. IV, f. 4 (1865).
CMU (Heller).
P. minutus A. M. -Edwards.
Pachygrapsus minutus A. M.-Edw., Nouv. Arch du Mus., ix, p. 292,
PI. XIV, f. 2(1873).
New Caledonia (A. M.-Edw.).
P. simplex Kingsley ex Herklots.
Grapsus simplex Herklots, Additamenta, etc., p. 9, PI. I, f, 8 (1851).
Boutry, West Coast of Africa (Kevklot^).
Genus NAUTILOGRAPSITS Edwards {Planes BelP).
Carapax narrow, regularl}' arcuate, sides slightly convex, and
bearing a rudimentary tooth behind the orbital angle. Front
' The genus Planes is a MS. one of Leach. Bowdich, in his " Excursion
to Madeira and Porto Santo," p. 15, f. 2 (1825), figui-es and mentions a spe-
14
202 PROCEEDINGS OF THE ACADEMY OF [1880.
more than half the width of the carapax. External maxillipeds
broad, meros broader than long. Posterior feet compressed.
N. minutus Edwards ex Linne.
Cancer minutus Linne Syst. Nat. Edit, xii, p. 1048 (1766).
Grapsus minutus Latreille, Hist. Crust, et Ins., vi, p. 68 (1803-4).
Grapsus cinereus Say, Journ. Acad. Nat. Sci. Phila. i, p. 99 (1817).
Grapsus pelagiciis Say, 1. c, p. 443 (1818),
Planes clypeatns Bowdich, 1. c, p. 15, PI. f. 2 (1835).
Grapsns testudineum et pelagicus Roux, Crust. Med., PI. VI, f- 6-7
(1838-30).
Ocypoda ( Grapsus) pusillus De Haan, op. cit., p. 59, PI. XVI, f. 2
(1835).
Naiitilogra^isus minutus Edw., Hist. Nat. Crust, ii, p. 90 (1837).
Grapsus divis Costa, Fauna Napoli, Crustaces, PI. IV, f. 1 (1838-1851).
Planes mvtmtus Wliite, Cat. Brit. Mus. Crust, p. 43 (1847)
Kautilograpsiis Major et Smithii, McLeay in Smith Zool., South Africa,
Annulosa, pp. 66-67 (1849).
Planes Unneana Bell, British Stalk-eyed Ciiistacea, p. 135 (1851).
Planes eyaneus Dana, Proc. Phil. Acad., 1851, p. 250.
Wautilograpsus angustatus Stimpson, Proc. Acad. Nat. Sci. Pliila.,
1858, p. 103.
Carapax smooth, arcuate in both directions ; front nearly
straight, post-orbital tooth small, sometimes obsolete. Sides
arcuate. Meros of chelipeds with its inner distal border dentate ;
Carpus with a tubercle on the inner surface ; hand smooth, fingers
deflexed. Ambulatorj^ feet compressed, ciliate.
Gulf Stream! (Many Collectors); West Indies! (Dr. Griffith); Suri-
nam! (Dr. Hering); Falkland Is.! (Dr. Wilson); Peru! (Dr.
Ruschenberger) ; West Coast of Mexico ! Alaska ! (Dr. W. H. Jones) ;
China! (Capt. Putnam); N€^n Zealand! Natal! (Dr. Wilson); Rio
Ganibia! (J. Cassin); Mediterranean (Dr. Wilson); France!
(Guerin); " toutes des me7-8!^' (Guerin). Guerin's ideas of the dis-
tribution leave nothing more to be said.
Genus EUCHIROGRAPSUS M.-Edwards, 1853.
Carapax depressed, subquadrate, sides slightlj' arcuate, with
three teeth behind the orl;»ital angle ; orbits entire. Antennae long,
cies in these words : " A small crab, f. 3, a and b, which I conceive to be a
new species of Planes was found in gi-eat numbers amongst the anatiferse."
In a foot-note the species is described as follows : "It was of a delicate,
but bright, rose-color ; from the symmetrical form of its test (notched so
regularly as to increase the projection and distinctness of its chaperon), it
may be called P. clypeafi/s." This can hardly be considered as a sufficient
description to establish the genus, and hence I prefer to retain the com-
monly accepted name.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 203
entering the orbit; Meros of the external maxillipeds about half
the length of the ischium, its outer distal angle rounded, its inner
excavate and bearing the palpas.
E. liguricus Edwards.
EuclnrograptiitK Uguneus Edwards, Archives du Museum, vii, p. 153,
PI. X, f, 2 (1853).
Nice (Edwards).
Genus BRACHYGRAPSUS nov.
Carapax broader than long, arcuate, without transverse lineation,
sides nearlj^ straight, with one tooth behind the angle of the orbit.
Meros of the external maxillipeds shorter than broad, its external
distal angle prominent, the internal one bearing the palpus.^
B. Isevis nov.
Front straight, external angles of orbit not prominent, tooth of
lateral margin spiniform. Meros of cheliped triquetral, bearing an
obtuse tooth on the upper border. Carpus with an acute internal
spine. Hands inflated, smooth ; fingers acute. Ambulatory feet
elongate, slender, but slightly compressed, the dactjdi longer than
the propodal joints.
New Zealand ! (E. Wilson).
Genus PTYCHOGNATHUS Stimpson, 185S {Gxnihoffrajims A. M.-Edwards).
Carapax flat, lateral border emarginate. External maxillipeds
verj' broad, nearly meeting, the exoguatli fully as broad as the
ischium. The carpus bears the palpus at the middle of the ante-
rior margin, and has the external distal angle strongly produced.
Sijnoj^sis of Species.
Exognath of external maxilliped extending to or exceeding the external
distal angle of the ineros. riedelii.
Exognath extending only to the middle of the meros.
Oblique portion of branchial ridge bounded by a granulated ridge.
jnlipes.
Oblique portion without a prominent boundary. pvsUlus.
Insufficiently characterized. glaher.
P. glaber Stimpson.
Ptychognathns (jlahcr Stimpson. Proc. Acad. Nat. Sci. Philadelphia,
1858, p. 104.
Bonin I. (Stm. )
1 I am not certain as to the exact position of this genus, as it appears to
combine the characters of both the Cydometopa and Catometopa. In the
form of carapax and structure of the external maxillipeds it closely resem-
bles Trapezia. In the male genital appendages it is allied to the Grapsidce,
where for the present I prefer to allow it to remain.
204 PROCEEDINGS OF THE ACADEMY OF [1880.
Dr. Stimpson's short diagnosis presents no characters which are
not held in common by both P. riedelii and pilipes, excepting the
non-pilose hand, which in other species of the genus is only of
sexual importance.
P. riedelii Kingsley ex A. Milne-Edwards.
Gnathograiisus riedelii A. M.-Edw., Nouv. Arch, du Museum, iv, p.
182, PI. XXVII, f. 1-0 (1868).
Celebes (A. M.-Edw.).
P. pusillus Heller. ^ '
Pfi/chognathns pusillus Heller, Riese der Novara Crustaceen, p. 60, 1867.
Onathograpsus barbatiis A. M.-Edw., Nouv. Arch, du Museum, ix, p.
316, PL XVII, f. 4, 1872.
Carapax depressed, nearly smooth, with a few shallow impres-
sions anteriorly. Front slightly sinuate. Antero-lateral margin
with two indistinct teeth behind the orbital angle. Chelipeds finely
granulate, but without spines or tubercles. Hands of the male
with a lanose spot on the outside at the base of the fingers ; in the
female this is wanting. The exognath of the external maxillipeds
reaches only to the middle of the meros. Ambulatory feet slender,
compressed.
Mauritius! (Guerin'); Nicohars (Heller); New Caledonia (A. M.-
Edw. )
P. pilipes Kingsley ex A. Milne -Edwards.
Onathograpsus pilipes A. M.-Edw., Nouv. Arch, du Museum, iv, 184,
PI. XXVII, f. 6-10 (1868).
This species is scarcely more than a variety of P. pusillus, but
I prefer for the present to leave them separate.
Philippines and Celebes (A. M.-Edw.).
Genus ACM^OPLEUEA Stimpson, 1858.
Carapax depressed, the antero-lateral margins entire. External
maxillipeds nearly meeting the meros, bearing the palpus on the
middle of the anterior margin ; the exognath narrow.
A. parvula Stimpson.
Acmceopleura parvula Stimpson, Proc. Acad. Nat. Sci. Philadelphia,
1858, p. 105.
Japan (Stimpson).
Genus PSEUDOGRAPSUS M. Edwards (1837), reatr.
Carapax depressed, transverse ; sides arcuate, with two teeth
behind the orbital angle. Front less than half the width of the
1 These were labeled by Guerin " Sesarnia penicillata sp. ined."
1880,] NATURAL SCIENCES OF PHILADELPHIA. 205
carapax. Meros of external maxillipeds broader than long,
shorter than the ischium and with its external distal angle
strongly produced.
Synopsis of Species.
Hands inflated without elevated lines.
Fingers with many long hairs, carapax inflated. setosus.
Hairs on the hand between the bases of the fingers short,
carapax flat. albus.
Hands with an elevated line on the lower outer surface, fingers
without hairs. craisus.
P. setosus.
Cancer barbatus Rumph., PI. X, No. 2 (1705).
Cancer setosus Fabricius, Suppl, Ent, Syst., p. 339 (1798).
Grapsus penicilliger Latr., Reg. An, (I Edit.), iii, p. 16, PL XII, f. 1
(1817).
Eriocheir? penicilliger De Haan, Fauna Japonica, Crust,, p. 31 (1835),
Pseudograpsus penicilliger Edw., Hist. Crust., ii, p. 83 (1837).
Pseudogrupsus barbatus Edw., Ann, Sci. Nat., Ill, xx, p. 191 (1853).
Eastern Seas (Auct).
P. albus Stimpson.
Pseudograpsus albus Stimpson, Proc. Acad. Nat, Sci., Phila., 1858,
p. 104,
Japan (Stimpson) ; New Caledonia (A, M,-Edwards),
P. crassus A, Milne-Edwards.
Pseudograpsus crassus A, M.-Edw., Nouv. Arch, du Mus., iv, p. 176,
PI, XXVI, f. 6-10 (1868),
Celebes (A. M,-Edw),
Genus VARUNA Edwards, 1830 {Trichopus De Haan, 1835).
Carapax depressed, sides arcuate, two-toothed. Antennulae
oblique. Antennae entering the orbit ; external maxillipeds
slightly gaping. Meros much shorter than the ischium, its ex-
ternal distal angle expanded. Palpus articulating with the middle
of the anterior margin. Exognath half as wide as ischium.
Ambulatory feet compressed, natatorial,
V. litterata Milne-Edwards ex Fabricius.
Cancer litterata Fabr., Suppl. Ent. Syst., p. 342 (1798).
Trichopus litterata De Haan, Fauna Japonica, Crust., p. 32 (1835).
Vai-una litterata Edw,, Diet. Class. d'Hist. Nat., xvi, p. 511 (1830).
Hist, Nat, Crust., ii, p, 95 (1837),
Carapax smooth, cardiac region partly circumscribed. Front
straight, orbits fissured above, lateral teeth sepai-ated bj' slight
fissures. Posterior margin of meros of chelipeds acute, the
206 PROCEEDINGS OP THE ACADEMY OP [1880.
inferior granulate, the anterior with spiniform tubercles. Carpus
with a prominent internal spine and one or two smaller ones.
Hands inflated, rough, an elevated line on the lower outer margin.
Ambulatory feet strongly compressed, margins ciliate. In a
specimen from New Zealand the carpal spines are wanting.
Philippines / (E. & T. B. Wilson) ; Indian Ocean/ (Guerin) ; New
Zealand! (E.Wilson); China! (Capt. Putnam, Peab. Acad.) ;
Fenang! (J.P.Ward, Peab. Acad.) ; Japan (Miers); Mauritius
(A. M.-Edw.).
Genus TITICA "White, 1847.
Carapax depressed, sides more or less arcuate, two-toothed.
Antennulse oblique. Antennae entering the orbit. Meros of
external shorter than the ischium, its external angle not expanded.
Posterior feet compressed.
Synopsis of Sjiecies.
Inner margins of fingers strongly haired. hariimanus.
Hands naked.
Angles of front acute. gracilipes.
Angles of front rounded. glabra.
v. gracilipes White.
Utica gracilipes White, Proc. Zool. Soc, 1847, p. 86. Adams and
White, Voyage Samarang, Crust., p. 53, PI. XIII, f. 6 (1850).
Philippines ( White. )
U. glabra A. Milne-Edwards.
Utica glabra A. M.-Edw., Nouv. Arch, du Mus., ix, p. 296, PI. XIV,
f. 3 (1878).
New Caledonia (A. M.-Edw.).
U. barbimanus A. Milne-Edwards,
Utica barbimanus A. M.-Edw., 1. c, p. 297, PL XIV, f. 4 (1873).
New Caledonia (A. M.-Edw.)
Genus GLYPTOGEAPSUS Smith, 1870.
Carapax transverse, distinctly areolate, sides arcuate, three-
toothed. AntennfE entering the orbit. Extei-nal masillipeds
nearly meeting. Ischium and meros nearly equal in length, very
broad, the meros being broader than long, its external distal angle
not expanded. Ambulatory feet elongate, the dactyli quadran-
gular and spinose.
G. impressus Smith.
Glyptograpsus impressus Smith, Trans. Conn. Acad., ii, p. 154 (1870).
Acajutla, West Coast of Central America (Smith).
1880.] NATURAL SCIENCES OE PHILADELPHIA. 207
Genus HETEROGRAPSUS Lucas, 1849.
{Paeudograpsus, pars, Edw., Dana: Hemujrapsus Dana.
Carapax arcuate, front inclined, antero-lateral margins dentate-
External maxillipeds nearly closing. The meros as long or longer
than broad, and bearing the palpus on the middle of its anterior
border, the exognath narrow.
As the distinctions between the species are mainly comparative
and the descriptions of authors are very brief, no sj'nopsis can
be given. The species may however be divided into two sections,
according to the number of teeth on the antero-lateral margin.
A. Antero-lateral margin inth tico teeth behind the orbital angle.
H. Incasii Edwards.
Heterograpsus sexdentatus Lucas, Exploration Algiers, 1, p. 19, PI. II,
f. 4 (1849), (nee Edwards^.
Heterograpsus lucasii Edwards, Ann. Sci. Nat., Ill, xx, p. 193 (1853).
Carapax regularly arcuate, epigastric lobes but slightly indi-
cated. Front four-lobed. Antero-lateral margin with two promi-
nent, narrow, acute teeth. Chelipeds without spines or tubercles,
the hands of the male are smooth and rounded, in the female thej'
have a double crest above and two elevated lines on the outer
surface. 1 Ambulator}' feet slender, naked ; dactyli long and
slender.
Algiers! (Dr. T. B. Wilson) ; Candiu (Edwards).
H. sexdentatus Edwards.
Cyclograpsus sexdentatus Edwards, Hist. Nat. Crust., ii, p. 79 (1837).
Hemigrapsus sexdentatus Dana, U. S. Expl. Exped., Crustacea, p. 348,
PI. XXII, f. 2 (2850).
Heterograpsus sexdentat^ts Edwards, Ann. Sci. Nat., Ill, xx, p. 192,
PI. vii, f. 7 (1853).
Carapax arcuate, surface uneven, laterally granulate. Epi-
gastric lobes prominent. Front straight. Antero-lateral margin
with two teeth behind the orbital angle, the teeth broad, the
emarginations narrow. Chelipeds without spines or tubercles.
Hands small, fingers excavate. Ambulatory feet moderate, naked ;
dactyli short and stout.
Australia! (E. Wilson) ; Neic Zealand! (Dr. T. B. Wilson) ; Bay
of Islands (Dana).
1 Milne-Edwards (1. c, p. 192,) divides this section of the genus into
two groups, one with the hands roiuided and without longitudinal cre.sts,
the other with cre.sts a division which evidently cannot be maintained.
208 PROCEEDINGS OF THE ACADEMY OF [1880.
H. sanguineus Edwards ex De Haan.
Grapsus sanguineus De Haan, Fauna Japonica, Crustacea, p. 58, PL
XVI, f. 3 (1835).
Orapsus marmoratus White, Cat. Brit. Museum, Crust., p. 41, 1847
( sine descr. ) .
P-seudograpsus nudus Dana, Proc. Acad. Nat. Sci. Phila,, 1851, p. 349,
Expl. Exped., Crust., p. 335, PI. XX, f. 7 (1852).
Heterograpsus sanguineus, marmoratus et maculatus Edw., Ann. Sci.
Nat., Iir, XX, p. 193 (1853).
Heterograpsus nudus Stimpson, Proc. Acad. Nat. Sci. Phila., 1858,
p. 104.
Carapax posteriori}^ nearly flat, in front arcuate, with scattered
punctae and a carved line of larger depressions running inward
from the last tooth of the lateral margin. Front obsoletely two-
lobed ; antero-lateral margin with two teeth closely similar to
those of H. sexdentatus. Chelipeds smooth, with small red spots,
which persist in alcoholic or dried specimens. Hands with an
external crest, becoming obsolescent with age ; fingers excavate.
Ambulatory feet short, stout, naked, punctate ; the dactyli very
stout.
California ! (many localities and collectors) ; Vancouver Is. ! (Dr.
A. S. .Packard, Jr., in Peabody Acad. Science); San Lorenzo,
Gulf of California! (Wilkes' Expedition); Australia! (E.
Wilson) ; Japan (De Haan) ; Punipet and Auckland (Heller) ;
Hong Kong (Stimpson) ; Sitka (White) ; Polynesia (Edwards).
H. crassimanus Kin^sley ex Dana.
Hemigrapsus' crassimanus Dana, Proc. Phila. Acad., 1851, p. 250.
U. S. Ex. Exped., Crust., p. 349, PI. XXII, f. 4 (1852).
Hawaian Is. (Dana).
H. crenulatus Edwards ex GuTin.
Grapsus crenulatus Guerin, Voy. Coquille, ii, pt. i, p. 15 (1838).'
Cyclograpsus crenulatus Edw., Hist. Nat. Crust., ii, p. 80 (1837).
Hemigrapsus crenulatus Dana, U. S. Ex. Ex., Crust., p. 349, PI. XXII,
f. 3 (1852).
Heterograpsus crenulatus Edward.s, Ann. Sci. Nat. Ill, xx, p. 193
(1853).
Heterograpsus barbigerus Heller, Yerh. Z. B. Gesellschaft Wien,
1862, p. 522.
Heterograpsus barbimanus Heller, Novara Crustacea, p. 53, PI. IV,
f. 5 (1867).
Australia (Guerin) ; Mic Zealand (Edwards) ; Bay of Islands
(Dana) ; Punipet and Auckland (Heller).
1 The title bears the date 1830, the introduction to the Crustacea and
Arachnida, "15 Novembre 1838," and the plates 1826. Guerin in his de-
scription, refers to Milne-Edwards' classic work as then in manuscript.
1880.] NATURAL SCIENCES OP PHILADELPHIA. 209
H. elongatus A, M.-Edw.
IIetero(]raj)sus elongatus Alpb. Milne-Edwards, Nouv. Arch, du Mu-
seum, ix, p. 317, PI. XVII, f. 5 (1873).
Ifew Caledonia (A. M.-Edw.)-
H, oregonensis Stimpson ex Dana.
Pseudograpsus oregonensis Dana, Proc. Acad. Nat. Sci. Philadelphia,
1851, p. 248. Expl. Exped. Crust., p. 334, PI. XX, f. 6 (1852).
Heterograpsus oregonensis Stimpson, Proc. Acad. Nat. Sci. Philadel-
phia, 1858, p. 104.
Carapax depiessed, anteriorlj' iri-egularly roughened ; proto-
gastric lobes defined. Front four-lobed, the inner lobes the more
prominent. Antero-lateral margin with two prominent teeth.
Chelipeds without spines or tubercles. Hands with an elevated
line on the lower outer surface, the inner surface of the hand of
the male with a pilose spot. Ambulatory feet moderate, ciliate.
Pacific Coast of North America from Puget Sound! (Geo. Davidson);
to Santa Cruz! (Miss Hecox).
There are two si3ecimens belonging to this species in the Mu-
seum of the Academy, bearing the label " New Providence, W. I.,
Dr. H. C. Wood, Jr."
H. penicillatus Stimpson ex De Haan.
Briocheir penicillatus De Haan, op. cit., p. 60, PI. XI, f. 6 (1835).
Heterograpsus penicillatus Stimpson, Proc. Acad. Nat. Sci. Philadel-
phia, 1858, p. 104.
Japan (De Haan).
H, erythraeus Kingsley ex Kossmann.
Pseudograpsus erythra'us Kossmann, Reise in den Kiistengebiete des
rothen Meeres, p. 61, PI. 1, f. 5 (1877).
Bed Sea (Kossmann).
H. pallipes Milne-Edwards.
Pseudograpsus pallipes Edw., Hist. Crust., ii, p. 82 (1837).
Heterograpsus pallipes Edw., Ann. Sci. Nat., Ill, xx, p. 194 (1858).
Atistralia (Edw.).
B. Antero-lateral margin with three teeth behind the orbital angle.
H. octodentatus Edwards.
Cyclograpsus octodentatus Edwards, Hist. Nat. Crust., ii, p. 80 (1837).
Heterograpsus octodentatus Edwards, Ann. Sci. Nat., Ill, xx, p. 194
(1853).
Locality unknown.
H. affinis Kingsley ex Dana.
Hemigrapsus affinis Dana, Proc. Acad. Nat. Sci. Philadelphia, 1851,
p. 250. U. S. Exp. Exped., Crustacea, p. 350, PI. XXII, f. 5 (1852).
Patagonia (Dana).
210 PROCEEDINGS OF THE ACADEMY OF [1880.
H. spinosns Edw.
Heterograpsus spinosus Edw., Ann. Sci. Nat., Ill, xx. p. 194 (1853).
Vanikoro (Edw.) ; Australia (A. M.-Edw.
Genus ERIOCHEIR De Haan (1835).
Carapax quadrate, antero-lateral margin two-toothed. Front
much less than half the width of the carapax. Antennulffi
oblique. Antennae not excluded from the orbit. External
maxillipeds nearly closing. Meros as long as broad, the external
distal angle not expanded and the carpus articulating with the
middle of its anterior border.
Synopsis of Species.
Sides convex.
Mesial frontal lobes rounded. japonkus.
Frontal lobes acute. sinensis.
Sides straight. rectus.
E. japonicus De Haan.
Eriocheir japonicus De Haan, op. cit., p. 59, PI. XVII (1835).
Carapax nearly flat, surface uneven. Front four-lobed, mesial
lobes rounded, outer lobes acute ; protogastric lobes prominent,
granulate. Antero-lateral border two-toothed, with indications of
a third. Meros of chelipeds with the margins granulate, the
posterior terminating in an acute tooth. Carpus with a promi-
nent internal spine ; distal margin of the carpus and external
surface of the hand with thickly set long hair ; the inner surface
of the palm with a short horizontal line of granules. Fingers
sub-excavate. Ambulatory feet hairj^ above.
Japan! (no donor's name).
E. sinensis.
EriocJieirlxsl sinensis Edw., Ann. Sci. Nat., Ill, xx, p. 177 (1853).
Arch, du Mus., vii, p. 146, PI. IX, f. 1 (1854).
G?dna (Edw.).
E. rectus.
Eriocheir rectus Stirapson, Proc. Acad. Nat, Sci. Phila., 1858 p. 103.
Macao (Stimpson).
Genus PERIGRAPSUS Heller, 1862.
Carapax convex, sides arcuate, with one tooth behind the angle
of the orbit. Front narrower than half the width of the carapax.
Meros of the external maxilliped a little longer than broad and
beai'ing the palpus on the external angle. Dactyli of ambulatory
feet spined.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 211
P. excelsus Heller.
Perigrapsus excelsus Heller, Verb. Zool. Bot. Ges. Wien, 1862, p. 522.
Novara Crust., p. 50, PI. V, f. 1 (1865).
TaUti (Heller).
Genus PLATYGRAPSTJS Stimpson, 1858 {Platynotus De Haan, 18.35, preocc).
Carapax flat. Front horizontal. Sides nearly straight, with
two teeth behind the angle of the oi'bit. Meros of the external
maxilliped longer than the ischium and bearing the palpus on the
external angle.
P. depressus Stimpson ex De Haan.
Platynotus depressus De Haau, Fauna Japonica, Crust., p. 63, PI. VIII,
f. 2 (1835).
Platygrapsus depressus et convexiusculus Stimpson, Proc. Acad. Nat.
Sci. Phila., 1858, p. 104.
Carapax depressed, smooth; front horizontal , four-lobed , mesial
lobes the larger; sides with two teeth behind the angle of the
orbit, the posterior tooth indistinct. Chelipeds smooth and
unarmed ; meros with the anterior margin acute ; carpus without
spines or tubercles ; hand with an elevated line on the lower outer
surface ; fingers slender, gaping. Ambulatory feet elongate.
Japan! (no donor's name); Hong Kong (Heller); LooChoo (Stimpson).
Tribe Sesarmini (Sub-family /SesarmincB Dana).
Meros and ischium of the external maxillipecls crossed obliquelj-
by a piliferous ridge.
Genus METASESARMA Edw (1853).
Carapax quadrate, sides but slightly arcuate, entire ; front
broad, deflexed. Sub-orbital lobe large, meeting the front and
excluding the antennae from the orbit. Meros of external maxilli-
ped greatl}^ elongate, its apex rounded.
Si/nojjsis of Species.
Hands smooth, externally and above. roiisseauxi.
Hand roughened above. granulans.
Haud roughened above and externally. trapezium.
M. rousseauxi Edw.
Metasesarma rousseauxi Edw., Ann, Sci. Nat., Ill, xx, p. 88 (1853).
Arch, du Mus., vii, p. 158, PI. X, f. 1 (1854).
Zanzibar (Edw.).
212 PROCEEDINGS OF THE ACADEMY OF [1880.
M, granulans^ Heller.
Metasesarma granularis Heller, Verb. Z. B. Ges. "Wien, 1863, p. 523.
Metasesarma rugulosa Heller, Novara Crust., p. 65 (1865).
TaUU (Heller).
M. trapezium Stimpson ex Dana.
Sesarma trapeziu7n Dana, U. S. Expl. Exped. p. 354, PI. XXII, f. 8
(1852).
Metasesarma trapezium Stimpsou, Proc. Acad. Nat. Sci. Pbila., 1861,
p. 373.
Sandioich Is. (Dana).
Genus SARMATIXJM Dana, 1851 {Metagrapsua Edw., 1853).
Carapax convex, sides arcuate, entire or toothed. Front in-
clined, less than half the width of the carapax. External maxilli-
peds nearly as in Sesarma. Ambulatory feet with the margins
entire.
Synopsis of Species.
Sides of carapax with two teeth behind the orbital angle.
Hands externally smooth and rounded.
Carapax smooth, band transversely plicate above. crassum.
Carapax areolate, hand smooth above. curvatum.
Hands externally roughened.
Hands externally bearing a pectinate crest. pectinatum.
Hands without a prominent external crest.
Hands with an internal granulate ridge. punctatum.
Hands entire within. indicum.
Sides of carapax entire. integrum,
S. crassum Dana.
Sarmatium crassum Dana, Proc. Acad. Nat. Sci. Phila., 1851, p.
U. S. Expl. Exped., Crust., p. 358, PI. XXIII, f. 1 (1852).
Samoan Is. (Dana).
S. curvatum Kingsley ex Milne-Edwards.
Sesarma curvata Edw., Hist. Crust., ii, p. 75 (1837).
Metagrapsus curvatus M.-Edw., Ann. Sci. Nat. Ill, xx, p. 189 (1853).
Arch, du Mus., vii, p. IGO, PI. X, f. 3, 1854.
Senegal (Edw.).
S. pectinatum Kingsley ex Milne-Edwards.
Metagrapsus pectinatus Edw., Ann. Sci. Nat. Ill, xx, p. 189 (1853).
Martinique (Edw.).
S. punctatum Kingsley ex A. Milne-Edwards.
Metagrapsus punctatus A. M.-Edw., Nouv. Arch, du Mus., ix, p. 308,
PI. XVII, f. 2 (1873).
Neio Caledonia (A. M.-Edw.).
' I have here as in other places employed the earlier name ; what reason
Dr. Heller had for the change I cannot imagine.
1880.] NATURAL SCIENCES OF PHTLADELPHIA. 213
S. indicum Kingsley ex A. Milne-Edwards.
Mctagra'psus indicus A. M.-Edw., Nouv. Arch, dii Mus., iv, p. 174,
XXVI, f. 1-5 (1868).
Celebes (A. M.-Edw.).
S, integrum Kingsley ex A. Milne- Edwards.
Metagrapsiis integer A. M.-Edw., Nouv. Arch, du Mus., ix, p. 309,
PL XVII, f. 3 (1873).
New Caledonia (A. M.-Edw.).
Genus RHACONOTTJS Gtrstsecker, 1866.
Carapax sub-quadrate, sides areuate, toothed. Front narrow,
about one-third the width of the carapax. Mei'os of external
maxillipeds nearly- as broad as long and about half the length of
the ischium. Ambulatory feet compressed, the margins of the
joints serrate.
B. crenulatus Gerstrecker.
Rhaconotus crenulatus Gerstsecker, Arcliiv fiir Naturgeschicbte, xxi,
p. 142 (1856).
Locality unknown.
Genus SESARMA Say, 1818. (Pachysoma De Haan, 1835. Holometopus Edw., 1853.).
Carapax thick, quadrate, lateral margins straight, entire or
toothed. External maxillipeds with an oblique piliferous ridge
crossing the ischium and meros ; the meros elongate, its apex
rounded. Antenna? entering the orbit.
I have not attempted to revise the species of this genus on
account of a lack of sufficient material. I merely give a list of
the described' species, indicating in a few cases the synonymy,
but leaving the task of comparing a large number of poor descrip-
tions to some future carcinologist.
S. aflBnis Edw. (= ? quadrata).
Orapsus (Pachysoma) affinis De Haan, op. cit., p. 61, PI. XVIII, f.
5 (1835).
Sesarma affinis Edwards, Ann. Sci. Nat., Ill, xx, p. 183 (1853).
Japan (De Haan) ; China (Edw.) ; Natal (Krauss).
S. africana Edwards.
Sesarma africana Edw., Hist. Nat. Crust., ii, p. 73 (1837).
Senegal (Edw.).
S. americana Saussure.
Sesarma americana Saixssure, Mem. Soc. Phys. et Hist. Nat., xiv, p.
441 (1858).
St. Thomas, W, I. (Saussure).
S. angolensis Capello.
Sesarma angolensis Capello, Descr. tres sp. Nov. Crust, du Africa Oc-
cident, p. 4, f. 2 (1864).
Angola, West Africa (Capello).
214 PROCEEDINGS OF THE ACADE^n' OP [1880.
S. angusta Smith.
Sesarma angusta Smith, Trans. Conn. Acad., ii, p. 159 (1870).
Panama (Smith).
S. angustifrons A. Milne-Edwards.
Sesarma angustifrons A. M.-Edw., Noiiv. Arch, du Mus. Bulletin, v,
p. 26 (1869).
Sandwich Is. (A. M.-Edw.).
S, angustipes Dana.
Sesarma angustipes Dana, U. S. Expl. Exped., Crust., p. 853, PI. XXII,
f. 7 (1853).
Florida ! West Indies ! Brazil !
S. aspera Heller.
Sesarma aspera Heller, Novara, Crust., p. 63, PI. VI, f. 3 (1865).
Nicohars, Ceylon, Madras (Heller).
S. atrorubens Hess.
Sesarma atrorubens Hess, Archiv fiir Naturgeschichte, xxxi, p. 149,
PI. VI, f. 13 (1865).
Sydney, Australia (Hess).
S. aubryi A. Milne-Edwards.
Sesarma aubryi A. M.-Edw., Nouv. Arch., Bulletin, v, p. 35 (1869).
Nouv. Arch., ix, p. 307, PI. XVI, f. 3 (1873).
JS'ew Caledonia (A. M.-Edw.).
S. bidens Milne-Edwards ex De Haan. '
Grapsus [Pachysoma) bidens De Haan, op. cit., p. 60, PI. XVI, f. 4,
PL XI, f. 4 (1835).
Sesarma bidens Edw., Ann. Sci. Nat., Ill, xx, p. 185 (1853).
Japan (De Haan); Hong Kong, Nicobars (Heller); Friendly Is.
(Dana) ; Ceylon, Zanzibar (Hilgendorf).
S. boucourti A. Milne-Edwards.
Sesarma boucourti A. M.-Edw., Bulletin, 1. c, p. 38 (1869).
Siam (A. M.-Edw.).
S. chirogona Tozzetti.
Sesarma chirogona Targioni-Tozzetti, Zoologia del Viaggio della
Magenta, p. 186, PI. IX (1877).
Yokohama (Tozzetti).
S. cinereus Say ex Bosc.
Grapsus cinereus Bosc, Hist. Nat. Crust., i, p. 204 PI. V, f. 1, 1802-3
(teste Auct. ).
Sesarma cinerea Say, Jour. Acad. Nat. Sci., Phila., i, p. 443 (1818K
Virginia ! to Florida ! and the West Indies I
S. dentifrons A. Milne-Edwards.
Sesarma dentifrons A. M.-Edw., Bulletin, 1. c, p. 31 (1869).
Samoan Is. (A. M.-Edw.).
S. dehaani Milne-Edwards.
Grapsus (Pachysoma) guadratus De Haan, op. cit. p. 63, PI. VIII, f. 3
(1835).
Sesarma dehaani Edw., Ann. Sci. Nat., Ill, xx, p. 184 (1853).
Japan (De Haan).
1880.] NATURAL SCIENCES OF PHILADELPHIA. 215
S. dusumieri Milne-Edwards (= S. bidens).
Sesarma dus^wiieri "Ed-w., Ann. Sci. Nat., Ill, xx, p. 185 (1853).
Bombay (Edw.).
S. elegans Herklots.
Sesarma elegans Herklots, Addit. ad Fauuam Afric. Occident., p. 10,
PI. I, f. 10 (1851).
Boutry, West Africa (Herklots).
S. elongata A. Milne-Edward?.
Sesarma elongatum A. M.-Edw, Bulletin, 1. c, p. 30 (1869).
Madagascar (A. M.-Edw).
S. erythrodactyla Hess.
Sesarma erythrodactyla Hess, Arch, fiir Naturges., xxxi, p. 151, PI. VI,
f. 10 (1865).
Sydney, Australia (Hess).
S, eydouxi Milne-Edwards.
Sesarma eydouxi Edw., Ann. Sci. Nat., Ill, xx, p. 184 (1853).
Cochin China (Edw.); Madras (Heller).
S. fascicularis Hilgendorf ex Herbst,
Cancer fascicularis Herbst, Krabben nnd Krebse, PI. XLVH, f. 5
(1795).
Sesarma mederl'E.dyv,, Ann. Sci. Nat., Ill, xx, p. 185 (1853) teste Hil-
gendorf.
Batavia (Edw.).
S. frontale A. Milne-Edwards.
Sesarma frontale A. M.-Edw., Bulletin, 1. c, p. 27 (1869).
Madagascar (A. M.-Edw.).
S. germani A. Milne-Edwards.
Sesarma germani K. M.-Edw,, Bulletin, 1. c, p. 28 (1869).
Poulo Condore.
S. gracilipes A. Milne-Edwards.
Sesarma inipressa jiin. Homb. et Jacq., Voy. Ast. et Zelee, Crust., PI.
VI. f. 5.
Sesarma gracilipes Edw., Ann. Sci Nat., Ill, xx, p. 182 (1853).
Vaoa (Edw.); Nicobars (Heller).
S. guerini Milne-Edwards.
Sesarma guerini Edw., Ann. Sci. Nat., Ill, xx, p. 183 (1853).
Locality unknown.
S. gattatum A.Milue-Edwards.
Sesarma guttatum A. M.-Edw., Bulletin, 1. c, p. 26 (1869).
Zanzibar (A. M.-Edw.).
S. haematocheir Kingsley e.\ De Haan.
Orapsus {Pachysoma) hcematocJieir De Haan, op. cit. p. 67, PI. VII,
f. 4 (1837).
Holometopus hamatocheir^ Edw., Ann. Sci. Nat., Ill, xx, p. 188 (1853).
Japan (De Haan).
^ M. Milne-Edwards has elevated this species to distinct generic rank on
what seem to me wholly inadequate characters. A similar proceeding
with other Sesarmm would result in the creation of nearly a dozen genera.
216 PROCEEDINGS OF THE ACADEMY OP [1880.
S. impressa Milne-Edwards.
Sesarma impressa Edw., HLst. Nat, Crust, ii, p. 74 (1837).
Locality unknown.
S, indica Milne-Edwards.
Sesarma indica Edw., Hist. Nat. Crust., i i, p. 74 (1837).
Indian Seas (Edw.); Ceylon and Nicobars (Heller).
S. intermedia Milne-Edwards ex De Haan.
Grapszis (PacJiysoma) intermedia De Haan, op. cit., p. 61, PI. XVI, f.
5 (1835).
Sesarma intermedia Edw., Ann. Sci. Nat., Ill, xx, p. 186 (1853).
Sesarma Iceve A. M.-Edw., Bulletin, 1. c, p. 27 (1869).
Japan (De Haan); Sliangliai, Hong Kong (Heller);
Arrow Is. )A. M.-Edw.).
S. lafondi .Tacquinot et Lucas.
Sesarma lafondi Jacquinot et Lucas, Voyage Astrolabe et Zelee,
Crast., p. 70, PI. VI, f. 4 (1853).
Batavia (J. and L.).
S. leptosoma Hilgendorf.
Sesarma leptosoma Hilgendorf, in Decken's Reise, p. 91, PL VI, f. 1
(1869).
Zanzibar (Hilgendorf).
S. lividv.m A. Milne-Edwards.
Sesarma lividum A. M.-Edw., Bulletin, 1. c, p. 25 (1869), N. Arch., ix,
p. 303, PI. XVI, f. 2 (1873).
I^ew Caledonia {A. M.-Edw.).
S. longipes Krauss.
Sesarma longipes Krauss, Slid Afric. Crust., p. 444, PI. Ill, f. 2 (1843),
Umlass River, S. Africa (Krauss).
S. Mullerii A. Milne-Edwards.
Sesarma mulleri A. M.-Edw., Bulletin, 1. c, p. 29 (1869).
. Desterro, Brazil (A. M.-Edw.).
S. obesum Dana.
Sesarma obesum Dana, Proc. Pliiia. Acad., 1851, p. 250; U. S. Expl.
Exped., Crust., p. 35d, PI. XXII, f. 10.
Balabac Straits (Dana).
S. oblonga Martens.
Sesarma oblonga Martens, Monatsber. Akad. Wiss. Berlin, 1868, p. 611.
Philippines (Martens),
S. obtusifrons Dana.
Sesarma obtusifrons Dana, Proc. Phila. Acad, 1851, p. 250 ; U, S, Expl.
Exped., Crust., p. 355, PI. XXII, f. 9 (1852).
Sandwich, Is. (Dana).
S. occidentalis Smith.
Sesarma occideyitalis Smith, Trans. Conn, Acad,, ii, p. 158 (1870),
West Coast of Central America (Smith).
S. pentagona Hutton (=.? S. tetragona).
Sesarma pentagona Hutton, Trans, New Zealand lust,, 1875, p, 279,
Ne%ci Zealand (Hutton).
iS80.] NATURAL SCIENCES OF PHILADELPHIA. 217
S. quadrata Milne-Edwards ex Fabricius.
Ca7icer quadratus Fahr., Suppl. Ent. Syst., p. 341 (1798).
Ocypoda plicata Bosc, op. cit., i, p. 198, 1803-3 (teste A, 3I.-Edw.).
Orapsus {Pachysoma) pictus et affinis De Haaii, op. cit., pp. 61-66
(1835-37).
Sesarma quadrata Edw., Hist. Xat. Crust., ii, p. 75 (1837).
Sesarma picta Krauss, op. cit., p. 45 (1843 j.
Japan (De Haan) ; New Caledonia (A. M.-Edw.) ; Zanzibar (Hil-
gendorfj.
S. recta Randall.
Sesarma recta Randall, Jour. Acad. Nat. Sci., Pliila., viii, p, 123 (1839).
Surinam! (Randall).
S. reticulata, Say.
Sesarma reticulata Say, Jour. Acad. Nat. Sci., Pliila., i, pp. 73, 76 et
442, PI. IV, f. 6 (1818).
Sesarma cinerea De Kay, N. Y. Fauna, Crust., p. 15 (1842).
Virginia ! to Florida !
S. ricordi Milne- Edwards.
Sesarma ricordi Edw., Ann. Sci. Nat., Ill, xx, p. 183 (ISoS).
Hayti (Edw.).
S. roberti Milne-EJwards.
Sesarma reticulata McLeay in Smith Zool. S. Africa, p. 65 (18 ), vix
Say. .
Sesarma roberti 'Edw., Ann. Sci. Nat., Ill, xx, p. 183 (1853).
Gori! (Dr. Wilson) ; So. Africa (McLeay).
S. rotundata Hess.
Sesarma rotundata Hess, 1. c, p. 149, PI. VI, f. 9 (1865).
Sydney (Hess).
S. rotundifrons A. Milne-Edwards.
Sesarma rotundifrons A. M.-Ed., Bulletin,!, c. p. 30 (1869).
Samoan Is. (A. M.-Edw.).
S, rupicola Stimpson.
Sesarma rupicola Stimpson, Proc. Phila. Acad., 1858, p. 106.
Japan (Stimpson).
S. schiittei Iless.
Sesarma schuttei Hess, 1. c, p. 150, PI. VI, f. 11 (1865).
Sydney, Australia (Hess),
S. similis Hess (= S. atrorubens).
Sesarma similis Hess, 1. c, p. 150 (1865).
Sydney (Australia).
S. sinensis Milne-Edwards.
Sesarma sinensis Edw., Ann. Sci. Nat., Ill, xx, p. 186 (1853).
C7iina (Edw.). .
S. smithii Milne-Edwards.
Sesarma smithii Edw., Ann. Sci. Nat., Ill, xx, p. 187 (^1853); Arch, du
Mus., vii, p. 149, PI. IX, f. 2 (1854).
Natal (Edw.):; New Galedonia (A. M.-Edw.).
15
218 PROCEEDINGS OF THE ACADEMY OF [1880
S. sulcata Smith.
Sesarma sulcata Smith, Trans. Coun. Acad., ii, p. 156 riSTO).
Corinto, Nicaragua ! i .J. A. McNiel, Peab. Acad.
S. taeniolata Miers ej; White MS.
Sesarma tceniolata White MS., Miers, Proc. Zool. Soc., London (1877),
p. 137.
Philippines! (Dr. Wilson, with White's label).
S. tetragona Milne-E.lwards ex Fab icius.
Cancer tetragonon Fabriciiis, Suppl. Ent. Syst., p. 341 fl798).
Grapautt tetragonon Latr., Hist. Crust, et Ins., vi, p. 71 (1803-4).
Sesarma tetragona Edw., Hist. Nat. Crust., ii, p. 73 (1837).
Zanzibar (Hilgendorfj to New Caledonia (A. M.-Edw.).
S. trapezoida Milne-Edwards.
Sesarma trapezoida Edw., Hist. Nat. Crust., ii, p. 74 (1837).
Locality unJcnown.
S. ungulata Milne-EdwarJ?.
Sesarma ungulata Edw., Ann Sci. Nat. IK, xx, p. 184 (1853).
Celebes (Edw.).
S. vestita Stimpson,
Sesarma vestita Stimpson, Proc. Acad. Nat. Sci. Phila., 1858, p. 106.
Japan (Stimpson).
S. villosum A. Milne- Edwards.
Sesarma villosum A. M.-Edw., Bulletin, 1. c, p. 31 (1869).
Samoan Is. (A. M.-Edw.).
S. violacea Herklots.
Semrma violacea Herklots, op. cit., p. 10, PI. I, f. 9 (1851).
West Africa! (Du Chaillu).
Genus ARATTJS M.-Edw., 1853.
Carapax trapezoidal, elongate, narrow behind, sides straight^
entire ; front deflexed, verj- broad. External maxillipeds as in
Sesarma. Ambulatory- feet compressed, the dactyli very short-
A. pisoni Milne-Edwards.
Sesarma in»o»{ Edw.. Hist. Crust., ii, p. 76, PI. XVI, f . 4-5 (1837).
Ar a tus 2^isoni Fidw., Ann. Sci. Nat., Ill, xx, p. 187, 1853.
Carapax transversely arcuate, the branchial regions obliquely
plicate. Front vertical, its margin two-lobed. Meros of cheli-
peds triquetral, the margins denticulate^ the anterior one slightl\-
expanded distally. Carpus externally granulate. Hands every-
where granulate, the fingers ornamented with pencils of stiff
black hairs.
Florida! (H. E. Webster, Union College); West Indies! (many
collectors and localities); West Coast of Nicaragua! (J. A.
McNiel, Peab. Acad); Bio Janeiro (Heller i ; Praya, Brazil
(Martens).
1880.J NATURAL SCIENCES OF PHILADELPHIA. 219
Genus CLISTOCCELOMA A. M.-Edwards, 1873.
Cavapax sub-quadrate, sides dentate. Sub-ocular lobe large,
united to the front and excluding the antenna? from the orbit.
Meros of external maxillipeds short and rounded.
C. balansae A. Milne-Edward?.
Clistoca'lomn balanscv A. M.-Edw., Nouv. Arch, du Mus., ix, p. 311,
PI. XVII, f. 1 il873j.
New Caledonia (A. M.-Edw.).
Genus HELICE De Haan (1835).
Carapax quadrate, front deflexed, sides straight, with one, two
or three teeth behind the orbital angle. Antennaj entering the
orbit. Meros of external maxillipeds as long as or longer than
the ischium, its external distal angle prominent, its distal border
truncate.
Synopsis of Species.
Lateral margin with three teeth behind the angle of the orbit.
Ambulatory feet with a single distal spine on the meros.
A transverse ridge on the branchial regions. tridens.
No transveree crest on the branchial regions.
Hands smooth. spinicarpa.
Hands roughened. latreiUei.
Meral joints of ambulatory feet with several spines. dentipes.
Lateral margin two-toothed.
Hand strongly granulate. gdiuUchaudi.
Hand nearly smooth.
Meral joints of ambulatory feet with a spine on the
upper distal margin, the hands of the male with a
pilose spot at the base of the fingers. pilimana.
Meral joints without spines, hands of male without
pilose spots. crassa.
Lateral margin one-toothed. gibba.
Imperfectly characterized. leachii.
H. tridens De Hnan.
Helice tridens De Haan, op. cit., p. 57, PI. XI, f. 2, PI. XVI, f. 6 (1835).
Carapax longitudinall}- strongly convex, punctate, front curved
downward, its anterior border sinuate when vicAved from above.
Superior margin of the orbit sinuate, oblique ; lateral margin
with three teeth behind the orbital angle, the posterior tooth
rudimentary. Branchial regions with an oblique i-idge running
inward from this tooth. Orbits coarsel}' crenulate below. In-
ferior borders of the meral joints of the cheli]ieds with small
tubercles. Carpus spined on the inside. Hands externally
220 PROCEEDINGS OF THE ACADEMY OF [1880.
smooth, siib-cristate and granulate above, internally granulate ;
fingers excavate. Carpal and propodal joints of the first two
pairs of ambulatory feet pilose in front.
Japan!
H. spinicarpa Edward".
H. spinicarpa Edwards, Ami. Sci. Nat. Ill, xx, p. 190 (1853).
Locality unknown.
H. dentipes Heller.
Eelice dentipes Heller, Novara Crust., p. 62, PI. V, f. 5.
Ceylon (Heller).
H. latreillei Edwards.
Cyclogrupsus latreillei 'E^vnwAs, Hist. Nat. Crust., ii, p. 80 (1837).
Eelice Za/mto" Edwards, Ann. Sci. Nat. Ill, xx, p. 190 (1853).
Mauritius (Edwards).
H. gaudichaudi Edwards.
Eelice gaudichaudi Edwards, Ann. Sci. Nat. Ill, xx, p. 190, PI. VII,
f. 6 (1853).
Sumatra (Edwards).
H. pilimana A. Milne-Edwards.
Eelice pilimana Alph. Milne-Edwards, Nouv. Arch, du Mus., ix p.
313, PI. XVIII, f. 1 (1872).
New Caledonia (A. M.-Edw.).
H. orassa Dana.
Eelice crasm Dana, Proe. Phila. Acad. (1851), p. 252— U.S. Ex. Exp.,
Crust., p. 367, PI. XXIII, f. 8 (1853).
E. lucasii Edw., Ann. Sci. Nat. Ill, xx, p. 190 (1853).
Carapax closely resembling that of B. tridens, but with but two
teeth behind the orbital angle. Carpus of cheliped without an
internal spine, hands externally microscopically granulate, more
coarsely so internall}^, the upper margin acute. Carpal and pro-
podal joints of the first two i)airs of ambulator^' feet, pilose.
This is probably but a variety of H. tridens. Small females
show the elevated line on the hand characterizing H. lucasii.
New Zealand! (Dr. Wilson); Aukland (Heller); Australia (Dana).
H. leachii Hess.
Eelice leacMi Hess, Arcliiv fiir Naturgeschichte, xxxi, p. 153 (1865).
Sydney, Australia (Hess).
Genus CYCLOGRAPSUS Edw. (1837)(restrict). (Gnathochasmus McLeay.)
Carapax depressed, sides arcuate, entire front about half the
width of the carapax. Antenna not excluded from the orbit.
Meros of the external maxillipeds short, about as long as the
ischium; its external angle well marked, the palpus articulating
with the anterior margin.
1880. J NATURAL SCIENCES OF PHILADELPHIA. 221
C. punctatus Milne-Edwards.
Cyclograpsui punctatus Edw., Hist. Nat. Crust., ii, p. 78 (1837).
Gnathochasmus barbatus McLeay, in Smith, Zool. S. Africa, p. 65
(1838).
Sesarma barbata Krauss, Svid Af. Crust., p. 45, PI. Ill, f. 3 (1843).
Cyclograpsus audouinii, lavauxit, whitei, granulosus et reynaudi Edw.,
Ann. Sci. Nat. Ill, xx, p. 197 (1853).
Cyclograpsus Imvis Hess, Archiv fiir Naturj^escliicbte, xxxi, p. 152
(1865).
Oarapax smooth or slightl}' granulate ; sides arcuate in front,
straight behind. Front broad, nearly straight. Orbits externally
broadl}- emarginate, the emarginatiou continuing backward as a
groove for some distance. Hands externallj' smooth, internally
with a prominent longitudinal ridge. Male abdomen triangular,
regularly tai^ering from the third to the sixth joints, the seventh
much narrower than the sixth.
Neio Zealand! (Guerin); Australia! (E. Wilson and Wiikes' Ex-
pedition!; Cape of Good Hope, Madras, Java (Heller) ; New
Guinea (Edw.).
C. granulatus Dana.
Cyclograpsus granulatus Dana, Proc. Acad. Nat. Sci. Phila., 1851, p.
251 ; U. S. Ex. Exp. Crust., p. 361, PL XXIII, f. 4 ^1852).
Sandwich Is. (Dana).
C. cinereus Dana.
Cyclograpsus cinereus Dana, Proc. Acad. (1851), p. 251 ; U. S. Ex.
Exp. Crust., p. 360, PI. XXIII, f. 3 (1852*.
Cyclograpsus eydouxiY^dvf., Ann. Sci. Nat., Ill, xx, p. 198 (1853^
Valparaiso and Sandwich Is. (Dana).
C. longipes Stimpson.
Cyclograpsus longipes Stimpson, Proc. Acad. Nat. Sci. Phila. (1858),
p. 105.
Bo7iin Is. (Stimpson'.
C. integer Milne-Edward?.
Cyclograpsui integer Edw., Hist. Nat. Crust., ii, p. 79 (1837).
Florida! (A. S. Packard, Jr., Peab. Acad.); Brazil (Edw.).
Genus CHASMAGNATHUS DeHaan, 1835. [Paraijrapsns Edw.).
Carapax convex, sides arcuate, dentate, front curved down-
ward Antenna not excluded fi'om the orbit. Meros of external
maxillipeds longer than broad, widest distall}-, its anterior border
slightly excavate, the palpus medially articulated.
222 PROCEEDINGS OF THE ACADEMY OF [1880,
Sijnojjsis of Species.
Lateral margin with three tef th behind the orbital angle,
the posterior tooth inconspicuous. convexus.
Lateral margin with two teeth.
Front rounded. nrvillet.
Front nearly straight. gaimardii.
Front excavate.
Carapax and chelipeds granulate. granulatus.
Carapax and chelipeds smooth.
Epigastric lobes prominent. Icevis.
Epigastric lobes inconspicuous. suhquadratu^
Lateral margin with one post orbital tot>th. quctdridentatui,
C. convexus DeHaan.
CMamagnatlms eonvexus DeHaan, Fanna Japonica, p. 55, PL VII, f.
5 (1835;.
Jcifiin (DeHaan) ; Eastern Seas (Adams and White).
C. subquadratns Dunn.
CJiasmagnatTms siibquadratns Dana, Proc. Acad. Nat. Sci. Phila.^
1851, p. 251 ; U. S. F^x. Exp., Crust., p. 363, PL XXIIT, f. 5 (1852).
Ne^!) Zeulaiidf Avstraliaf (Dana).
C laevis Dana (=- ? C. subqnadratus.)
ChasmngnntMs laris Dana, Proc. Acad,, p. 252 ; Ex. Exp., p. 365, PI.
XXIII, f. 7 (18:;2).
Paragi-npsus verremixi 'Edw., Ann. Sci. Nat., HI xx, p. 195 (1853).
Paragrnpsus lavis Heller, Novara Crust, p. 55 (1865).
Carapax slightly convex, punctate ; regions not defined. Epi-
gastric lobes prominenf. Front deeply excavate in the middle,
when viewed from ahove. Antero-lateral teeth separated by
narrow fissures. Chelipeds everywliere smooth. Anterior surface
of carpus and propodus of first pair of ambnlatory feet tomentose.
Australia! (Guerin ; Ne^e Zealand (Miers).
C. urvillei Kingsl<y ex MiTne-Eduaids.
Paragrapsus iirvillei'Edyf., Ann. Sci. Xat., Ill, xx, p. 196 (185-3).
Va n ikoro I. ( E d w. ) .
C. granulatus Dara.
Clucsmognathns granulatus Dana, Proc. Acad., 1851, p. 251 ; U. S. Ex.
Exp., Crust., p. 364, PL XXIII, f. 6 (1852).
Helice granulata Heller, Novara Crust., p. 61 (1865).
Carapax convex, distinctly areolate, granulate ; the granules
on the branchial regions being larger ; epigastric lobes obsolete.
Front curved downward and, viewed from aboA'e, deeply excavate.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 223
Sides of carapax acute, the fissures between the teeth being very
slight ; all of the border of the carapax finely crenulate. Chelipeds
•externally granulate. Carpus produced internally ; the inner sur-
face of the hand with a patch of granules on the inner surface.
Carpal joints of the ambulator}- feet longitudiually sulcate.
Bio Janeiro I {Wilkes' Expedition); Bio Grande, Brazil! (Capt.
Harrington Peabody Academy).
C gaimardi Milne Edwards.
Cydograpsus gaimardi'Ed-w., Hist. Nat. Crust., ii,p. 79 (1837).
Paragrapsus gaimardi 'Edw., Ann. Sci, Nat. Ill, xx, p. 196 (1853 .
Australia (Edwards).
C, quadridentatus Kingsley <?•»• Milne-EiUvards.
Paragrapsiis quadridentatus Edw., Ann. Sci. Nat., IH, xx, p 19.")
(1853).
Australia (Edw.).
Sub family Plagusinse Dana.'
Carapax flattened, antennula? longitudinally plicate, lodged in
sinuses of the front, and visible from above.
Gtnus PLAGUSIA Latr., 18i)(j (restrict).
Meros of external maxilliped well developed, as broad as the
ischium.
P. speciosa Dana.
Cai-apax arcuate, covered everywhere with sqnamiform tubercles,
the inter paces being clothed with a short pubescence, these
tubercles Ijeing similar iu their arrangement to those of P. depressa
Sa}", but much more depressed than in that species. The margins
of the inter-antennular portion of the front is simple. Inferior
margin of the orbit acute, minutely denticulate. Sides of carapax
with two equal acute spiuiform teeth behind the angle of the orbit.
Feet closel}- resembling those of P. depressa., the ornamentatio»
being similar, but not so prominent. The hands, however, are
external!}- marked by six longitudinal impressed lines, the lowest
of them being on the inferior margin. The fingers are widely
gaping, the extremities deeply excavate. The dentiform process
1 This sub family having recently been revised by Mr. Miers (Annals and
Magazine of Natural Ilistoiy, V, ix, pp. 147-154, February, 1878), and as
I agree with his determinations and ideas of specific bmits, I omit the
synopsis of species from this paper, merely giving a few notes on the more
uncommon forms.
224 PROCEEDINGS OF THE ACADEMY OF [1880-
Oil the coxa of the third pair of aiiibulatoiy feet is niinutelj'
denticulate. The only additional character in the female is that
the squainag of the carapax are more depressed.
Mr. Miers (1. c, p. 151) remarks: "Only a carapax of this
species is known." The carapax referred to, Dana's type, was
destroyed in the Chicago fire. The Academy possesses two
specimens, male and female, sent b}^ Mr. Andrew Garrett, from
Tahiti.
Genus LEILOPHUS 1 Mi-rs, 1876. {Acanthnpus DeHaan.)
Meros of external maxillipeds very small, and much narrower
than the ischium.
L. pilimanus Miers ex A. M.-Edw.
Specimens of this rare species are in the museum of the Academy,
from the Sandwich Is. (J. K. Townsend) and Tahiti (A. Garrett).
So far as I am aware, the only other specimen, in an^^ collection,
is the type in Jardin des Plantes at Paris. The British Museum
has no specimens.
I am unable, either from the poorness of the descriptions, or
possible inaccurac}' of the figures, to assign the following species
to their proper generic positions.
Cyclograpsus ? tasmanicus Jacquiiiot et Lucas, Voyage Astrolabe et
Zelee, Crustaces, p. 76, PI. VI, f. 6 (1842-53).
Tasmania (J. et L.).
Cyclograpsus minutus J. et L., 1. c, p. To, PI. VI, f. 8 (1842-53).
Chili [S. et L.).
Grapsus inornaius Hess, Archiv fiir Naturgescliichte, xxxi, p. 148,
PI. VI, f. 11 (1865).
Sydney, Australia (Hess).
Grapsus Tiuzardi Desmarest, Consld. sur les Crust., p. 131 (1825).
Senegal (Desmarest).
Cancer tridens Fabricius, Suppl. Eiit. Syst., p. 340 (1798).
E. Indies (Fabricius).
Cancer Mspanus Herbst, PI. XXXVII, f. 1 (1796).
Goniograpsus pulcher Lockington, Proc. Cal. Acad., vii, p. 152 (1876).
Lower California (Lockington).
M. Henri Milne-Edwards (Archives du Museum, vii, p. 158,
1854) mentions a genus Holograpsus, possibly intending Holonie-
topus.
^ In the dismemberment of the genus Plugusia of Latreille, the name
Plagusia should have been retained for this section.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 225
June 1.
The President, Dr. Ruschenberger, in the chair.
Twenty-nine persons present.
A paper entitled " Description of a Partula supposed to be new,
from the Island of Moorea," bj' W. D. Hartman, M. D., was
presented for publication.
The Treasurer having announced the reception of a gift of
twenty thousand dollars from Jos. Jeanes, acting for the heirs of
the late Joshua T. Jeanes, who, in an unsigned codicil to his will,
had indicated his intention of bequeathing that amount to the
Academy, the following preamble and resolutions were unani-
mously adopted :
Whereas, The late Mr. Joshua T. Jeanes in a codicil to his
will bequeathed to tlie Academy twenty thousand doHars, an act
which may be regarded as significant of his appreciation and ap-
proval of the objects of the Society, but left this codicil without
his signature, and therefore legally inoperative ; and,
"Whereas, His executors have, placed in possession of the
Treasurer of the Academy the sum named, thus manifesting their
respect for the intention of their late brother in a most generous
and affectionate manner ; be it
Eesolved, That the Academy of Natural Sciences of Philadel-
phia highly appreciates and gratefully acknowledges the generosity
of the executors of the late Joshua T. Jeanes in bestowing on the
Academy- twenty thousand, dollars in compliance with his wish,
simply indicated.
Resolved, That the money thus bountifully given to the Society
be invested securely in the name of the Academy- of Natuial
Sciences of Philadelphia, to constitute a distinct and permanent
fund which shall be named the Joshua T. Jeanes Fund and the
income thereof shall be applied towards defraj'ing the ordinary
expenses of the Society.
Seiyenline Belts of Radnor ToirnsJup. Delaware Co. — At the
last meeting of the Mineralogical and Geological Section of the
Academy' of Natural Sciences, Theodore D. Rand read a paper
on the Seri^entine Belts of Radnor Township, Delaware Countj',
and the adjacent rocks. He adduced facts which he thought in-
compatible with Mr. Charles E. Hall's view, that the middle belt
consists of altei'ed Hudson River shales, and stated that the belt
was not continuous but was a succession of outcrops nearly east
220 PROCEEDINGS OP THE ACADEMY OF [1880.
and west from each oUier, the strike of which was, wherever ob-
servable, more N. E. and S. W. than the line joining them, thus
agreeing in stnicture with what Prof. Rogers states of the trap
dykes north of the serpentine in Chester County. He also called
attention to the existence of two trap dykes or two branches of
that extending through the Gulf Valley, and to curious markings
in quartz rock in the vicinity', suggestive of fossils in a formation
reaarded as azoic.
-o"
June 8.
The President, Dr. Ruschenberger, in the chair.
Twentj^-two persons jtresent.
A paper entitled " On the Development of Lemna minor," by
Wm. Barbeck, was presented for publication.
June 15.
The President, Dr. Ruschenberger, in the chair.
Eighteen persons present.
A paper entitled " A Bibliographical Catalogue of the Genus
Partula, with observations on the Species," b}' W. D. Hartman,
M. D., was presented for publication.
June 22.
The President, Dr. Ruschenberger, in the chair.
Eighteen persons present.
The deaths of AVni. G. E. Agnew and Morris L. Ilallowell,
members, were announced.
June 29.
The President, Dr. Ruschenberger, in the chair.
Eleven persons present.
The deaths of B. F, Lautenbach, M. D., and Wm. Kent Gilbert,
M. D., members, were announced.
On some Hoviologies in Bnnodont Dentition — Dr. Harrison
Allen, in speaking of the teeth of the Carnivora, Insectivora and
Chiroptera, dwelt upon the forms of the canines and premolars as
1880.] NATURAL SCIENCES OF PHILADELPHIA, 227
being valuable guides in interpreting the plan of the molars. He
traced the shapes of the last-named teeth from the sub-conical form
of the canine, with its associated cusplets or cingules characterizing
the canines, up to the complicated figures of the molars. Among
the seals, Leptonyx exhibits to the best advantage the figure result-
ing from the pronounced development of the antero-posterior cus-
plets, and is of still greater interest inasmuch as the molars retain
in all essential features the same parts. In genera where the form
of the molars is not so retained, the manner after which the depar-
ture takes place in the upper jaw is as follows :
1. The buccal cingulum becomes developed.
2. The buccal surface of the main' cusp is directed obliquelj^
backward and inward, and at the same time becomes concave.
3. In genera having the W-shaped pattern, the first V answers to
the concave, obliquel}^ placed buccal surface of the main cusp.
The second V is a vegetative repetition of the first, and is formed
from the posterior cusplet of the canine.
The W thus formed is a conspicuous feature in the molars of
most Insectivora and Chiroptera. It can be traced through its
several stages of development from the Carnivora. The genera of
the Procyonidjne exhibit the transition adYantageously. Tiie W
of the upper jaw, while forming a portion of the free under-surface
of the crown, is not functionally active as part of the grinder, but
is an extremeJij ohiiquehj placed portion of the i^hearing buccal
surface, and is not articular.
The V V seen from the palatal side of tooth form the summits
of two downward-projecting, prismoidal, shearing columns. Ex-
amined in relief from before backwards these columns are seen
to be of different relative lengths. In Vespertilio and Antrozous,
for example, where the appearance of the under free surfaces of
the crowns are almost identical, conspicuous differences in the
lengths of the columns are detected when the teeth are examined
with the columns in antero-posterior relief.
Tiie elevations placed to the palatal side of the base of the
columns are developments from the palatal fold of the cingulum
of the caniniforni tooth. If oue cingule be alone developed it lies
to the median side of the first V. Should a second be present, it
lies in an analogous position to the second V, and is much less
pronounced than the first.
The differences in the forms of the lower molars are traceable
to similar modifications of tlie simple cone and associated cusplets.
The second V is incomplete, the anterior limb not joining the
first to form a true W. There is no disposition to form a lingual
outgrowth. In its stead a tendency to backward projection from
the base of the second V exists. This projection is conveniently
called the " heel " of the tooth, and is always articular.
The forms of the canines and premolars are not as simple and
imiform as they- at first sight appear. They often present remark-
able differences in their details. This is especially true of these
228 PROCEEDINGS OF THE ACADEMY OF [1880.
teeth in the Chiroptera. The buccal, approximal and median surfaces
should be carefully studied in the different genera. Full descrip-
tions of these differences would be out of place in a communication
of this kind. One notable feature of man_y as seen in the canines
is especially well developed in the bats, viz., the junction of the
buccal and palatal surfaces resulting in forming a thin com-
pressed posterior edge. This may receive the name of the
" sabre " edge. It is repeated and exaggerated in the last pre-
molar and forms at least in Chiroptera (other than the Pteropidse)
the " sectorial " surface of the tooth. It constitutes a sharp
obliquelj'-placed ridge which is parallel with the last stroke of the
first V, and is doubtless serially homologous therewith.
The following were ordered to be printed :
1880.] NATURAL SCIENCES OF PHILADELPHIA. 229
DESCRIPTION OF A PARTULA SUPPOSED TO BE NEW, FROM THE ISLAND
OF MOOREA.
BY W. D. HART MAN, M. D.
Partula Mooreana, Hartman.
Shell sinistral, ovate, elongate, thin, translucent, pale yellowish
horn-color, apex darker ; whorls 5, flatly conA-ex, bod^^-whorl, with
or without from one to three narrow, pale, brown revolving bands ;
surface smooth, with fine, oblique striations, which are decussated
by crowded waved spiral striie ; a narrow white line beneath the
suture ; aperture hearh'- half the length of the shell, lip white,
moderately reflected, pillar tooth oval, prominent, situated nearest
the superior angle, umbilicus open, moderately compressed.
Length 18 mill., diameter 9 mill.
Hab. — Yaianai Valley, Island of Moorea (Andrew Garrett, Esq.).
In one hundred and fortj^-six species and varieties of Partula
represented in my collection, this shell possesses constant and
well-marked specific characters. Mr. Garrett informs me that
fifteen hundred specimens were all sinistral and dentate. The
surface of the shell resembles P. spadicea and varieties from
Moorea in possessing the thickly crowded waved spiral striffi.
This species is arboreal, and is not uncommon on bushes, in
Yaianai Valley, the metropolis of P. vexiUum Pse. = P. steiiostoma
Ph.
230 PROCEEDINGS OF THE ACADEMY OF [1880.
ON THE DEVELOPMENT OF LEMNA MINOR.
BY WM. BARBECK.
In the early part of last April, I found, in a little pond near
Gamden, N. J., among patches of Riccia fiuitans^ a number of
minute brownish bodies, which under the lens had very much the
appearance of germinating spores, showing at the top a greenish,
prothallium-like outgrowth. They were of an oval form, and less
than a millimetre in size.
I secured several of these little bodies, and, upon further exam-
ination under the microscope, I found that the}' contained a well,
developed embryo, which was enclosed by a comparatively large
cotyledon. Thus they were evidently the seeds of some mono-
cotyledonous plant.
I was not able to return to the pond until a week later. Within
this week the germination had been completed in a number of
specimens, and numerous little plants were developed, most of
them still in connection with the seed. These obovate, indis-
tinctly three-nerved individuals, with a single root hanging from
the under surface, were apparently Lemna minor. Thousands of
fresh seeds had meanwhile appeared at the surface of the water,
most of them germinating, and thus I could get the specimens in
all stages of their development. I have tried to show this gradual
development (up to the completion of the second frond) by a
series of illustrations, Plate XVIII.
Figures I and II represent longitudinal sections through a seed
in which the germination is about to commence. (Fig. I is from
the centre, Fig. II from a part nearer to the surface).
The seeds are seen surrounded by a comparatively strong
coat, the testa (^), which is considerably thickened towards the
top, where it covers the lid, or operculum (o), by means of which
the mycropyle is closed. In [c) we have the large cotyledon,
surrounded by a scanty layer of endospermium (sp.) ; in (u) and
(wj) the two lobes into which the cotyledon will afterwards split,
begin to be differentiated. The axis of the embryo (e) forms an
obtuse angle with the medial line of the cot3dedon. In {p) we
have the plumula, in {r) the radula of the embryo ; (/) indicates
a fissure inside of which the gemma of the second frond is being
formed.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 231
In Eig. Ill the testa has been removed from the cotyledon (c).
The two lobes are distinctlj' separate, iio) bearing the oi)erc'ulum
under which the upper part of the plumula is concealed. The
radula (r) is further developed ; in (g) we have the bud of the
second frond. The section in Fig. lY sliows the plumula (p)
fully developed into the first frond, which in (r) sends down its
radula. The angle formed by this frond and the axis of the cotj'-
ledon is about 120°. Corresponding to the first figures (c) and
(»'), are the lobes of the cotyledon. (^We have to bear in mind tliat
all the figures represent thin sections through the different parts.)
In rcalit}' the lobes of the cotyledon are two parallel obovate
sheets enclosing the basal part of the much larger, likewise obovate
frond. In this figure the gemma has been so far developed as to
show in ( f) the fissure in w-hich the bud of the third frond is
being clifierentiated. Its elongated inferior part (p') is the
secondarv plumula. In using a high power, the microscope will
show in the region indicated by (x) several rows of very wide cells.
Here the separation of the frond from the cotyledon will take place.
In Fig. V this separation is complete. In (p') we have the
yet more elongated plumula, in (r') the radula of the second frond,
and (/') shows again the fissure for the formation of the third
individual.
The section represented in Fig. YI goes through the radula (r),
showing a central vascular bundle (i) surrounded bj' a tissue of
very loose, almost hj-aline cells {I). In the further development
of the rootlet this outer tissue will follow the growth of the vascu
lar bundle to a certain extent ; then its basal part will be sepa-
rated from the frond. But, remaining in connection with the
more and more extending vascular bundle, this wide-celled tissue
will form at the top of the full-giown root the well-known hood or
calyptra, characterizing the roots in all Lemnacese.
The last two figures (YII and YIII) need no further explanation.
The^' show the formation and completion of the second frond (pO>
from which the third individual will be developed in the same
way as has been illustrated in the first figures. In i^p") we have
the plumula, in (r") the radula of the third fronds ; (/") in 'Fig.
YIII shows the fissure for the formation of the fourth individual.
In this way we see the propagation continued through the
summer, plant after plant being formed from a cleft of the pre-
ceding individual through a process of prolification.
232 PROCEEDINGS OF THE ACADEMY OF [1880
My investigations have been made only on the Lemna .minor ^
but there is no reason to doubt that in the development of the
"whole family of Lemnacese (analogous to our species) we have an
interesting instance of parthenogenesis, there being seeds (pro-
duced in autumn by a sexual process) from which, during the
course of summer, generation after generation is propagated with-
out any further fertilization.
1880.] NATURAL SCIENCES OF PHILADELrHIA. 233
DESCRIPTION OF A NEW SPECIES OF HEMITSIPTERUS FROM ALASKA.
BY AV. N. LOCKINGTON.
Hemitripterus cavifrons, nov. sp.
D. iv-xiv, 1-12. A. 14. P. 20. V. ^. C. 3-12-3. L. lat. 44.
Head very large and depressed ; abdomen protuberant, so that
the depth equals the width ; snout to tip of ascending process ot
pre-maxillar}' rising at an angle of about 45^ ; thence to occiput,
along the median line of the fish, deeply concave ; from occiput
to caudal peduncle regularly arched, the curve reaching its highest
point at about the tenth dorsal spine. Outline of anal base
corresponding to that part of the dorsal directly above it.
Depth, 3^ ; greatest width, 3^ ; length of head, 3^ ; length of
pectoral rather more than 4 times in the total length, caudal
included.
Axial length of snout, 3f ; longitudinal diameter of orbit, 6f ;
interocular width, 2|| times in length of head ; least depth of
caudal peduncle rather less than 5 times in greatest depth.
Anterior nostril on a level with the centre of tlie pupil, and
prolonged into a conspicuous tube ; posterior nostril somewhat
tubular.
Orbits elevated considerabl}' above the general surface of the
forehead, so tliat the concavity of the inter-ocular area is equal lo
about f of the transverse diameter of the eye ; e^^es lateral, some-
what elliptical.
Mouth very large, ver}' slightly oblique ; its width from tip to
tip of the opposite maxillaries, li in the length of the head, and
exceeding that of the upper jaw by more than one-third.
Pre-maxillaries not forming the whole of the margin of the
upper jaw, the maxillaries entering into it posteriorly.
Posterior extremity of maxillary considerably behind the orbit,
its upper margin not concealed by the pre-orbital in the closed
mouth.
Lower jaw slighth' projecting beyond the upper.
Several rows of sharp, recurved, cardiform teeth, forming a
broad band, in both jaws, also on the vomer, palatines and
phar^-ngeal bones. The teeth on the vomer and palatines slightly
longer than those on the jaws.
16
234 PROCEEDINGS OF THE ACADEMY OP [1880,
No gill-rakers ; pharyngeal hones large.
Supra-ocular and post-ocular ridges prominent, the former
curved inwards posteriorly, parallel to the post-ocular ; between
the two, at the posterior upper angle of the eye, are two small
spines or short ridges.
Occipital ridges with three tubercles, the anterior near the post-
oeular ridge, the two postei'ior near together and elongated
transversely ; a low ridge between the first and second.
Temporal ridge with three tubercles, the first immediately
exterior to the first of the occipital series, the second a longi-
tudinal ridge ; the third rounded, close to the second.
A long low crest across the operculum ; just above and anterior
to this a shorter ridge connecting it with the temporal series ; a
tubercle on the supra-scapula ; no spines upon the head, except
two upon the posterior border of the pre-operculum.
All the tubercles of the head and the spines of the pre-operculum,
covered b}' skin.
Maxillary with a fimbriated skinu}- flap near its posterior
extremit}' ; lower margin of mandible set along its whole length
with skinny flaps, of which three pairs are especially long and
fimbriated on both edges, while the posterior flap is very broad.
Lips well developed ; lower lip pendulous at sides, and to a
rather less extent in front, and bearing a fimbriated flap on each
side.
Two pairs of similar flaps on the snout, and two over each eye.
Gill-membranes continuous below the throat.
Branchiostegals, 6.
Origin of first dorsal slightly anterior to the lower pectoral
axil ; first two spines longest, about 2f in the length of the head,
fourth much shorter than the third, and a little shorter than the
fifth ; sixth, seventh and eighth much longer than fourth, the
remaining spines diminishing to the eighteenth, which is the
shortest.
A tag at the end of each spine, the membrane between the two
parts of the first dorsal notched considerably.
A spine at the commencement of the second dorsal, the base of
which is contained more than 2^ times in that of the first, the
rays increasing in length to the fifth, which is about ^ longer than
the longest spine of the first dorsal ; upper margin of second
dorsal, convex.
1880.J NATURAL SCIENCES OP PHILADELPHIA. 235
Anal longer than soft dorsal, arising opposite the last spine of
the first dorsal, and terminating somewhat posterior to the second,
Ninth to twelfth raj's slightly the longest.
Pectorals very broad and rounded, their base oblique, the tip of
the longest (sixth) ray reaching to about the sixteenth dorsal
raj- ; rays simple, the longest a little less than one-fourth the total
length of the fish.
Tentrals small, narrow, the longest (middle) ray rather more
than half the length of the longest pectoral ray, but not reaching
to the vent.
Caudal truncate on hinder margin, rather narrow, ra3's simple.
Yent midway between insertion of ventrals and origin of anal.
Lateral line with a series of skinny fimbriated flaps, similar to
those upon the head.
Body and head scaleless, but the former covered all over with
osseous papillre ; largest above the lateral line, smallest upon the
protuberant portion of the abdomen.
Color, in alcohol, blotches of dark purplish-brown on a lighter
ground ; the blotches on the fins conspicuous, and running into
transverse bars on the pectorals. Abdomen, light dirtj'-brown.
A single specimen of this interesting species was obtained by
Mr. W. J. Fisher, at St. Paul's, Kodiak. It is in the museum of
the California Academy of Sciences.
Hemitripterus cavifrons is the western representative of H.
acadianiis of the Atlantic, and differs from that species in the
following characteristics, among others : —
The great depression of the inter-ocular area, whence the specific
name ; the greater number of dorsal spines ; the shorter pectorals ;
the lesser depth of the posterior anal rays; the absence of hook-
like papillae along the lateral line, and the presence in their place
of fleshy slips ; and the smaller size of the bony papillae along
the dorsal region.
Dimensions.
Total length, 15.15
Greatest depth, 4.50
Least depth of caudal peduncle, 92
Length of head, 4.50
Width " 4.50
" of mouth, from tip to tip of maxillaries, . 3.75
Length of upper jaw along its curve, . . . 2.75
236
PROCEEDINGS OF THE ACADEMY OF
[1880.
Axial length of snout, .....
Longitudinal diameter of eye,
Interocular widtli, ......
Width of pectoral base, ....
Lengtli of longest (sixth) pectoral ray,
"Tip of snout to origin of dorsal, axial, .
" " " " " along top of head
'Length of base of spinous dorsal,
Height of first spine, .
" second spine,
" fourth spine,
" fifth spine, .
•" sixth spine, .
" eighth spine,
■^' eighteenth spine, .
•" spine of second dorsal,
" longest (fifth) ray of second dorsal,
Length of base of second dorsal,
Tip of lower jaw to ventrals, along abdomen,
" " " origin of anal, along abdomen
« '' " vent, ....
Length of ventrals, . .
base of anal,
longest anal lays (9-12)
1.25
.70
1.70
2:75
3.88
2.75
3.25
6.50
1.75
1.70
.90
.98
1.00
1.20
.'88
1.00
2.00
2.50
3.75
8.80
6.32
2.00
3.44
2.10
1880.J NATURAL SCIENCES OF PHILADELPHIA. 237
DESCRIPTION OF A NEW SPECIES OF CATOSTOMUS (CATOSTOMUS CYPHO)
FROM THE COLORADO RIVER.
BY WM. N. LOCKINGTON.
Catostomus cypho, sp. nov.
D. 3, 14. A. 2, T. C. M-16-1-Y. P. 18. T. 10. L. lat. 79.
Head conical ; snout long, much depressed ; dorsal outline
rising in a straight line to the occipital region, where commences
a prominent and considerably elevated liump, which attains its
greatest height at a distance from the occiput about equal to the
length of the snout, and thence descends to the origin of the
dorsal.
Along the base of the dorsal fin the dorsal outline descends
rapidly to about the end of the second third of the total length of
the fish ; caudal peduncle extremely elongated, and widening
considerably' toward the caudal base.
Abdominal outline almost straiglit to the origin of the anal,
thence diminishing to the caudal peduncle.
Greatest depth, at anterior pectoral axil, contained not quite
4^ times ; head a little more than 4 times in tlie total length ;
snout a little more tlian 2|, eye between 8 and 9 times in the
length of the head ; length of top of head not quite 2^ times in
the distance (in a straight line) from the tip of the snout to the
dorsal ; inter-ocular width equal to the length of the snout ;
pectoral about 1^ in length of head ; caudal peduncle about 3| in
the greatest depth.
Moutli rather wide, inferior. Lower lip small, in two distinct
ovoid lobes, covered with low, flat-topped papilla? ; the front of
the dentarj- bones covered by a well-developed, round-edged, horny
plate. Lower lip quite distinct from the upper ; the skin of the
cheeks forming an obliquely ascending crease, which does not,
however, cover the angle of the mouth. •
Anterior nostril horizontally sub-elliptical ; posterior large,
vertical, crescentic, entirely covered by its anterior flap.
Two distinct rows of pores on the top of the head ; connected
on the occiput with a series running behind and below the eye
almost to the tip of the snout.
Phar3Migeals arcuate, with numerous teeth, regularly' diminish-
ing posteriorly.
238 PROCEEDINGS OF THE ACADEMY OF [1880.
Opercular region well developed ; the distance from the posterior
margin of the eye to that of the operculum being, to the length of
the snout, about as eleven to nine. Posterior margin of operculum
and sub-operculum forming a continuous bold couA^ex curve.
Pectorals triangular-ianceolate, fourth and fifth rays longest ;
their tips extending to beyond the middle of the pubic bones,
ra3'S once or twice bifurcate, the first two excepted.
Yentrals reaching bej^ond the vent, the third rays longest, the
last about two-thirds as long ; all the ra^^s twice bifurcate except
the first.
Dorsal well developed, fourth and fifth rays longest, and con-
tained about 1^ times in the greatest depth ; first three rays
simple, the others twice bifurcate.
Anal considerably shorter than the dorsal, but equal in depth
to the height of the latter; the first two rays simple, the others
(except the last) twice or thrice bifurcate ; first ray about half as
long as the second.
Origin of the dorsal about one-sixth nearer to the tip of the
snout than to the centre of the base of the caudal (measuring
along the axis of the body), the base of its eighth ray above the
anterior axil of the ventrals.
The tips of the anal rays reach beyond the first caudal
accessories.
Caudal with numerous accessory rays, the longest about half as.
long as the outer simple principal ray ; the other principal rays
three times bifurcate ; posterior margin of fin triangularly emar-
ginate.
Scales C3'cloid, of variable size ; each scale with 8-16 conspicuous
radiating striie on its exposed portion ; ihe striae and their inter-
spaces crossed by numerous, much less distinct concentric striae.
Engaged portion of each scale with numerous diverging striae, less
distinct than those of the free portion. Scales along and near the
lateral line larger than those above and below, and increasing
considerably in size posteriorly, as do also those above and below,
so that the largest scales of the body are upon the peduncle of the
tail. The scales diminish much more rapidl}- in size downwards
than upwards, so that those of the abdominal region and behind
the pectoral base are by far the smallest. Scales somewhat
pentagonal, the length exceeding the height ; those upon the
caudal peduncle almost twice as long as high.
1880J NATURAL SCIENCES OF PinLADELPHIA. 239
Fins scaleless, as is also a small patch on the anterior part of
the dorsal hump.
Lateral line deflected near its origin, then running along the
median line of the bod^- to the origin of the caudal. Pores
simple.
Color of the preserved specimen silv£r3--gray above, light
straw-color or creamy on the abdominal region and under side of
the head ; fins light uniform slat^-gra}'. The color is produced
by numerous dark dots upon the scales and membrane between
them, but fewer upon the scales, the outlines of which are there-
fore quite distinct.
The hump is supported anteriorly by a very large trapezoidal
inter-neural, formed of a thick central pillar witli anterior and
posterior aloe, the latter twice as large as the former. The upper
margin of the bone is highest at the point of the central pillar,
from which it slopes anteriorly' and posteriorly. Tlie base of the
central pillar is broadly expanded transverseh', oftering a double
articulating surface on its under side. The next inter-neural is a
thin flat sub-rectangular plate, while the next three are expanded
above, attenuated below ; the fifth bent, and smaller than the
fourth, the loM'er portion of which is also bent forward. Inter-
neurals of dorsal fin with a central ray and an anterior and
posterior expansion dying out at their lower fourth ; symmetrical,
except that supporting the first two rays. Tliis is evidently
formed by two inter-neural bones, united by a thin bonj' plate,
which forms a broad expansion in front of the first, and a narrow
one behind the second.
Upon the first vertebra there is a broad articulating surface,
apparentl}"^ for the reception of the first inter-neural, as a thin
longitudinal perpendicular partition exactly fits into a notch
between the two articulating surfaces of that bone. Tlie trans-
verse processes of this vertebra are broadly expanded inferiorly,
and their lower edges suturally united to a pair of ver}^ large bony
plates of complex form, connecting tlie air-bladder with the back
of the skull.
From the anterior margin of each neurapophj-sis of the next
nine vertebne springs an upward-directed process, which, in the
first of these vertebnv, is almost as long as tlie neural spine, br,t
which diminishes in size on each successive vertebra.
The neural spines of the first two of these vertebriie are bifid.
240
PROCEEDINGS OF THE ACADEMY OF
[1880.
The single specimen from which the above description is taken
was brought from the Colorado River, at the junction of the Gila,
and was sent to the museum of the California Academj^ of
Sciences by John E. Currj^, Esq., Civil Engineer.
It is said that the species is not uncommon in the locality from
which this specimen was procured, and it is much to be regretted
that we have onl}- this example, especially since it is greatly
damaged by the extraction of the large inter-neural some two
years ago. The air-bladder is destroyed, so that it is impossible
to tell whether it agrees with the other species of Gatostomus, in
having that organ divided into two portions. The extremities of
the fins are also much broken, and the shape of the body distorted.
Dimensions
Total length,
Length to base of caudal.
Greatest depth, about .
Length of head, .
" top of head,
" snout, from e^'e,
Longitudinal diameter of eye,
Inter-ocular width,
Depth of head, at front of eye,
Snout, from front of uostrils,
Tip of snout to origin of dorsal, ii
Length of base of dorsal.
Height of longest dorsal ray,
Tip of snout to anterior portion of
Length of pectoral fin,
Tip of snout to anterior portion of
Length of ventrals,
'• anal base,
*■' longest anal ray, .
Tip of snout to origin of anal,
Width of caudal peduncle, .
Length of first inter-neural, .
Heiiiht of " '•
a straight line
pectoral
ventrals
base
INCHES.
Hi
n
n
2V
Wa
j>_
16
27
3 2
2|
1 15
2H
^ 3
1^
6|
3^
1 5
T«
29
32
1880.J NATURAL SCIENCES OF PHILADELPHIA. 241
P R O C E E D 1 N a S
OP THE
MiNERALOGICAL AND GEOLOGICAL SECTION OF THE ACADEMY
OF Natural Sciences of Philadelphia.
187 7-18 7 9.
May 28, 1877.
A New Polariscope — Mr. H. C. Lewis remarked that a cheap
and accurate polariscope for the measurement of the optic-axial
diveroence in minerals had Ions:: been a desideratum amona; min-
eralogists. He wished to direct attention to an instrument for this
purpose, lately made for him by Queen & Co., of this cit}', which
had proved very satisfoctory. The light was polarized b}^ reflec-
tion from a plate of black glass, converged upon the rotating stage
by two sets of adjustable lenses, and analyzed by a Nicol's prism.
A graduated circle of steel, having through its axis a sliding
forceps, is fastened at right angles to the stage. A pointer records
the amount of rotation of the forceps. The mineral to be examined
is either held in the forceps or is attached by a drop of oil to a
piece of thin glass which is held in the same way. Cross-hairs are
fixed below the eye piece, and the measurement of the divergence
of the optic axes is performed in the usual way. The instrument
was found to work admirably and could be recommended. The
adjustments were made quickly and the axial divergence could
be determined to within 30'. It is simple, absorbs but little light,
and gives good results even with very small fragments of minerals.
A Garnet with Inverted Crystallization. — Mr. Lewis ex-
hibited a garnet which he had found in Germantown, and stated
that it showed a very perfect example of inverted cr^'stallization.
Its form was a perfect trapezohedron except that one octant was
depressed, its apex lying within the crystal, one-halfway towards
the centre. The re-entrant angles corresponded in position with
the trihedral edges on the opposite octant of the crystal. The
garnet was an isolated one found in a matrix of gneiss. Atten-
tion was called to the fact that such inverted cr3'stallization was
apparently more common in the isometric than in other S3'stems
of crj'stallization and comment was made upon the cause of such
phenomena.
June 25, 1877.
Change of Serpentine into Quartz. — Mr. Theodore D. Eand
described and presented specimens showing the change of ser-
242 PROCEEDINGS OF THE ACADEMY OF [1880.
pentine into quartz, very strikingly shown near a quarry of
serpentine rock on tiie farm of John Stacker, about a tlaird of a
mile N. W. of Radnor Station, P. R. R., Delaware Co., Pa.
The outcrop of the serpentine is accompanied by a rock, locally
called " Ironstone," which however is a cellular quartz, generally
stained by oxide of iron. It occurs as loose masses in the soil,
generally of small size, but sometimes of over a hundred pounds
weight ; the cavities are frequently lined with drusy (piartz. This
rock is of common occurrence in connection with serpentine belts,
but that it has arisen from a decomposition of the serpentine,
has, he believed, not been observed elsewhere. On the south side of
Stacker's quarry a few feet below the original surface of the ground,
is a bed of soft serpentine much cracked ; a foot or two above,
these cracks are found lined with chalcedonic quartz, of paper-like
thinness ; above, the quartz thickens, the serpentine becomes more
and more decomposed, until near the surface the quartz onl3^ re-
mains, with the cavities empty, or filled with what appears to be
oxide of iron with alumina. It is an instance of pseudomorphism
on a large scale, the progress of which can be traced, step by
step, from almost unaltered serpentine to almost pure quartz.
Well-water. — In this connection the analysis of the water of a
well 50 feet deep in the serpentine, about 400 hundred feet from
the quarry, but under the same quartz outcrop, may not be
xniinteresting.
In a gallon of 70,000 grains, — mean of three analj^ses : —
Grains, per Gall. Parts in 1,000,000.
Silica, .....
2.753
89.3
Magnesia, ....
1.262
18.
Lime,
.262
3.7
Peroxide of Iron and Alumina, .
.577
8.2
Sulphuric Acid,
.687
9.9
Chlorine,
.124
1.7
.5.665 80.8
A New Locality for Siderite. — Mr. H. C. Lewis announced
Dunbar, Faj'ette Co., Penna., as a new locality for Siderite. It
there occurs in finely crj'stallized specimens in the interior of
nodules of amorphous Siderite. These nodules or concretions
are of various and often curious shapes. Doubly terminated
limpid quartz crystals and minute but ver}^ perfect crystals of
Pyrite are associated with those of Siderite, forming handsome
specimens.
Magnetite Markings in Muscovite. — Mr. Lewis made some
remarks on the markings in the Muscovite of Brandy wine
Hundred, Delaware. He proved that these markings were Mag
netite, by exhibiting their attractability by the magnet, and said
1880.] NATURAL SCIENCES OF PHILADELPHIA. 243
that in order to exhibit this properly, the section must be
exceedingly thin. He stated tliat an optical examination had proved
that the direction of the main lines of the markings corresponded
with or was at right angles to that of the crystallographic axes of
the Muscovite. He exhibited a plate of the mica shown distinctly
to be a twin by the two different groupings of Magnetite markings.
Examination in the polariscope confirmed this structure. Thus,
frequently, the crystalline structure of the mica and the direction
of its axes may he ascertained b}' an inspection of these markings
alone. It appeared, therefore, that the form and direction of the
markings was determined, not by independent crystallization of
the Magnetite forming them, but in part at least by the Muscovite
from which it had probably been derived. These markings are, in
some respects, pseudomorphs after Muscovite. He thought that
the statement in Dana's Mineralogy (p. 150), referring to these
markings, that "the brandling at angles of 60° indicates com-
position parallel to a dodecahedral face," was misleading, implying
that this form was produced by an inherent property of the Mag-
netite, and not, as he thought now appears, by the crystalline
structure of the Muscovite.
September 24, 1817.
A New Local if y for Asholite. — Mr. Lewis stated that he had
found Asbolite at Flourtown, Montgomery Co., a new locality for
this mineral. It is found in iron ore mines as an incrustation
upon Psilomelane. It is of a bluish-black color, is as soft as
graphite, and gives a shining streak when scratched by the nail.
The blowpipe indicates a considerable percentage of cobalt.
A New Locality for Fluorite. — Mr. W. W. Jefferis stated
that a few days since he was shown a massive specimen of Fluor-
6;par of a deep purple color, which was found in the limestone
near the village of Howellville, in Tredyffrin Township, Chester
County, Pa. This is the third locality of fluor in this county.
Epidote in Molybdenite. — Mr. Lewis mentioned that while
examining some Mol3'bdenite from Frankford, Phila., he had
found plates of a transparent hard mineral, of a light greenish-
yellow color, somewhat resembling Wulfenite, occurring in thin
laA-ers and minute scales between the folife of the Molybdenite,
and sometimes coating it as a thin film. It was not until after a
careful examination that it was proved to be Epidote in an
unusual form and situation.
October 22, 1877.
A New Locality for 3Iillerite. — Mr. Theo. D. Rand announced
the discovery of Millerite in Dolomite, from the Soapstone quarry
on the Schuylkill, in Philadelphia, near the Montgomery County
line. It occurred in capillary crystals in cavities of the Dolomite.
244 PROCEEDINGS OP THE ACADEMY OF [1880.
THE OPTICAL CHARACTERS OF SOME MICAS.
BY HENRY CARVILL LEWIS.
For the determination of the true characters of the micas — a
class of minerals rapidl}" gaining in importance — a knowledge of
their optical characters is almost as necessarj^ as is that of their
chemical composition. The optical is certainly the most ready
method of determination. The investigation here recorded is but
a partial one, and it is hoped that in the future it may be extended
so as to include most of the American micaceous minerals. The
measurements have been made for the most part upon minerals
which have never been optically examined, and are chiefly Ameri-
can. A few foreign species have been introduced for comparison.
The micas examined are largely those in the collection of the
Academy. Others were either in the writer's collection or have
been kindly given him by friends. The source from which each
specimen has been obtained is noted in the tables given below.
The polariscope used was made by Queen & Co., of this cit}',
and was described before this Section at its meeting last Maj-.
It reads to within 30'. The figures given below represent the
mean apparent optic-axial angular divergence for white light. As
the angle is somewhat different in different specimens and some-
times CA'en in different portions of the same plate, the figures must
be regarded as only approximate. In each case they represent
a mean of a number of separate measurements, and collectively'
are the result of over 1600 such measurements.
Phlogopite.
1. Sussex Co., N. Y. Hexagonal crystals, 3'ellow,
transparent. (Acad. Nat. Sci.) 6°.
2. Burgess, Oiit., Can. Clear brown. (A. N. S.) 6°45'.
3. N. Shore of Rideau Lake, Burgess, Can. Angle
varies in same piece. Clearbrown. (J.Willcox.) 6°-12°
4. Hammond, St. Lawrence Co., N. Y. Clear 3'ellow.
Hypei'bolas closer in the centre than thev are
near the edges of the crjj-stals. Crystals are
sometimes uniaxial in the centre and biaxial at
each end, while the plane of the optic axes at
1880.] NATURAL SCIENCES OP PHILADELPHIA. 245
one end is at right angles to that
y at the other end, viz. :
One crystal had angle at centre,
T^SO', angle at edge, 11°15'.
(A. K S.) 10^40'.
5. Jefferson Co., N.Y. Brownish-yellow. (A. N. S.) ll°21'-12°50^
6. Yrooman's Lake, Jefferson Co., N. Y. Wine-j'el-
low. (A.N. S.) 12°45'.
7. Oxboro', Jefferson Co., N. Y. Light yellow.
(A. X. S.) 13°12'.
8. Ottey Lake, Burgess, C. W. Brown hexagonal
crystals. (W. W. Jefferis). ' 13°20'.
A crystal from the same locality (J. Willcox)
gave for the outer part of crystal, 13°41';
centre of crystal, 11 °23'.
9. Calumet Is., Canada. Greenish-^^ellow, transpar-
ent. (A. N. S.) 13°20'-14°18'.
10. New Hampshire. Reddish-brown, similar to
Darby Biotite ; nearly uniaxial in thin plates. 13°10'-lt°.
11. Sparta, N. J. Dark brown; by reflected light
nearly black. 14°20'.
12. Yrooman's Lake, Jefferson Co., N. Y. Clear
pale 3'ellow. Some crA'stals show identical
phenomena with those from Hammond, St.
Lawrence Co.
13. St. Denis. "Plumose mica:" brown: thick,
nebulous hj-perbolas.
14. "Warwick, N. Y. Dark green; cleaving into
rhombs ; often mistaken for Biotite.
15. Pope's Mills, St. Lawrence Co., N. Y. Deep
reddish-bi'own. (W. W. Jefferis.)
16. Yesuvius. Black by reflected light, dark red-
dish-brown in thin plates. With icespar :
very opaque. (A. N. S.)
It. Clark's Hill, St. Lawrence Co., N. Y. Brown.
(W. W. Jefferis.)
18. Kennett Square, Del. Co., Pa. Brown ; in lime-
stone.
19. Edwards, N.Y. Pearly white. (W.W. Jefferis).
20. Rossie, St. Lawrence Co., N. Y. Yellowish-
brown. (A.N. S.) 15=52'.
14^
=24'.
W
=30'.
14<
=52'.
15°.
15
'°±.
15'
^10'.
15<
=20'.
15<
=30'.
246 PROCEEDINGS OF THE ACADEMY OF [1880.
21. S. Burgess, Can. Large brown crystal, purple
on edges. (A. N. S.) 16°38'.
22. Clark's Hill, near Rossie, N. Y. Brownish-
yellow. (A. N.S.) 16°45'.
23. Clark's Mills, N. Y. Light brown, transparent :
(probably identical with Nos. IT, 20, 22).
(A. N. s.) n°.
24. Canada. Asteriated Phlogopite. 19°.
25. S. Burgess. Clear yellow-brown. (A. N. S.) 19°.
26. Burgess, C. W. Yellowish-brown crystals, with
secondary cleavage along diagonal. (W. W.
Jefferis). 20°.
2*1. Rossie, N. Y. Black by reflected, reddish-
brown by transmitted light. (A. N. S.) 21°13^
28. Vesuvius. Black, crumbling, very opaque,
mixed with black hornblende. (A. N. S.) 21°20'±.
29. Burgess, C. W. Asteriated, not transparent,
silvery-brown. (A. N. S.) 21°35'±.
30. Rossie, N. Y. Black by reflected, dark brown
by transmitted light. Contains apatite. (A.
X. S.) 22°.
31. Chester Co., Pa. Feebly asteriated; locality
wrong ? ; probably from Rossie, N. Y. (A.
N. S.) 23°15'.
32. Alamutchie,X. J. Clear reddish-brown. (Frankl.
Inst.) 30°5'.
33. Van Arsdale's Quarry, Bucks Co., Pa. Red-
brown ; with graphite, etc. " 34°.
Biotite.
1. Easton, Pa. White, silver mica. 2°±:.
2. Antwerp, N. Y. Greenish-white. 0°.
3. Culsagee, N. C. White. 0°.
4. Vesuvius. White. 0°.
5. Darby, Del. Co., Pa. Deep red. 0°.
6. Delaware Co., Pa. Crystal in muscovite ; black
by reflected, brownish-red by transmitted light. 5°±.
7. Scotland. Brown. 0°.
8. Rossie, N. Y. Brown. 0
Probably several of these Biotites have an angle of l°-2°.
o
0°.
0°.
56'
=25'.
56
°50'.
59'
=20'.
60°.
60'
=40'.
61'
= 10'.
63°.
63°4'.
63'
=15'.
63^
'4r.
64
°23'.
1880. J NATURAL SCIENCES OF PHILADELPHIA. 247
Lepidomelane.
Arendal, Norway. Black; uniaxial.
Frankford, Phila. Black; uniaxial.
Muscomte.
1. Brunswick, Me. Briglit green scales. (A. N. S.)
2. Pennsbury, Pa. (A. N. S.)
3. Yesuvius. With adularia. (A. N. S.)
4. Button's Mills, Del. Co., Pa, (J. M. Cardeza.)
5. St. Lawrence Co., N. Y. Greenish-white, plu-
mose radiated crystals, showing Airy's spirals.
(A. N. S.)
6. Darby, Phila., Pa. Small scales in gneiss.
7. Siberia. (A. N. S.)
8. Germantown, Phila. Smokj- brown, clear crystals.
9. Plainfield, Conn, Margarodite. Contains 5 p. c.
of water.
10. Poorhouse, Del. Co., Pa.
11. Germantown, Pa.
12. Germantown, Pa. Containing enclosed crystals
of a black, uniaxial mica. 64°30'.
13. Frankford, Pa. In hornblende rock : in calcite,
Avith fluorite and epidote. (T. D. Rand). 64°50'.
14. Falls of Schuylkill, Phila. In hornblende rock. " 65°.
15. Cumberland, England. "Nacrite." (A. N. S.) 65°.
16. Goyaz, Brazil. (A. N. S.) 65°50'.
17. Brandywine Hundred, Del. Containing mag-
netite markings. 65°-67°30'.
After heating until it whitens, it has an
angle of 49°.
18. Litchfield, Me. (A. N. S.)
19. Portland, Conn.
20. Southern Colorado. Identical with mica of
Pennsbur}', Pa., and Brandywine Hundred,
Del., having magnetite markings,
21. Grafton, N. H.
22. Chandler's Hollow, Del. (J. M. Cardeza.)
23. Black Hills, Wyoming. (A. N. S.)
24. Zinnwald, Bohemia. (A. N. S.)
25. Buncombe Co., N. C. (A. N. S.)
65°
-68°34'.
66°.
66°7'.
66°12'.
66°40'.
66°48'.
66°51'.
67°30'.
6r
^30'.
67'
^45'.
61'
=45'.
68°.
69^
n9'.
69'
=38'.
69'
=45'.
70'
=14'.
W
=30'.
74'
°10'.
75°.
76'
= 15'.
248 PROCEEDINGS OF THE ACADEMY OP [1880.
26. Germantown, Pa. Large silvery plates.
27. Dixon's Quarry, Del. Pale green.
28. Connecticut. Green scales. (A. !N^. S.)
29. Georgetown, Col.
30. Upland, Del. Co., Pa. Pale green. (.J. M.
Cardeza.)
31. Germantown, Pa. Pale green.
32. Chester Co., Pa. (A. N. S.)
33. Westchester Co., N. Y.
34. Fabyans, White Mountains, N. H.
35. Glacier of the Aar, Switz. (A. N. S.)
36. Trumbull, Conn. Margarodite.
37. Paris, Me. Rose-color. (A. ]S^. S.)
Where not otherwise indicated, the above muscovites are of a
clear j^ellowish-brown tint.
Lepidolite.
1. Altenberg, Saxony. With Pycnite ; sometimes
distorted. (A. N. S.) 31°.
2. Zinnwald, Bohemia. Often very irregular. On
different parts of the same piece the angle
varies from 34°30' to 51°30'. (A. N. S.) 49°30'
3. Paris, Me. Much distorted ; several axes. (A.
N. S.) 60°±:.
4. Middletown, Conn. 66°.
Talc.
1. Lafayette, above Manayunk, Pa. Exfoliating :
fan-shaped cr3^stals : images much distorted. 12°40'.
2. Lafayette, Pa. Clear. 15°.
3. Lafayette, Pa. Foliated talc ; distorted images. 15°.
4. Harford Co., Md. White. 15°.
5. Shetland Is. Clear pale green, sometimes nearly
uniaxial. • 17°.
Pyrophyllite.
Westana, Sweden. 106°51'.
Serpentine.
Chrysotile from Chester Co., Pa., shows strong double refraction
when the fibres make an angle of 45° with the plane of polariza-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 249
tiou of the instrument. Bissectrix apparently parallel to the
fibres, Probabljr orthorhombic. Common serpentine and Wil-
liamsite show no double refraction.
Damourite.
1. Culsagee, N. C. In scales : analyzed by Koenig.
(F. A. Genth.) 66°lt'±.
2. Unionville, Pa. " Emerylite : " irregular h3-per-
bolas. (A. X.S.) 69°35'±:.
3. Unionville, Pa. On corundum. 72°.
4. Unionville, Pa. " Corundellite." (J. M. Car-
deza.) 12°.
5. Horsjoberg, Sweden. (T. D. Rand.) 72°25'.
6. Chester Co., Pa. " Margarite : " irregular, show-
ing sometimes four hyperbolas. (A. N. S.) T2°30'.
1. Haywood, N. C. "Altered from corundum." (F.
A. Genth ) 74°.
8. Unionville, Pa. Analyzed by Sharpless. (F. A.
Genth.) U°10'.
9. Unionville, Pa. Analyzed by Koenig. (F. A.
Genth.) 74°15'.
10. Newtown, Conn. With Cyanite. 74°24^
11. Newlin, Chester Co. "Margarite." (A. X. S.) 75°50'.
It is evident that the minerals labelled Emerylite, Corundellite,
Margarite, etc., are all Damourite.
Euphyllite.
.1. Chester Co., Pa. (A. N. S.) 37°-40°.
2. Unionville, Pa. "Original." (F. A. Genth.) 36°30'.
A thicker piece in which the h^-perbolas were very dim, had an
angle of 45°±.
This result is interesting, as the optical angle given by Silliman
is 71°.
Cookeite.
Paris, Me. In small scales. 42°40'.
17
250 PROCEEDINGS OF THE ACADEMY OF [1880.
Vermiculite.
1. E. ISTottingham , Chestei* Co., Pa. Hallite. In
green crystals : uniaxial. 0°.
2. Cecil Co., McL, Magnesia Quarry. Hallite. Con-
tains enclosed arrow-shaped crystals like Hal-
lite : uniaxial. 0°.
3. Chester Co., Pa., Brown's Quany. Uniaxial.
(T.D.Rand.) .0°.
4. Macon Co., N. C. Maeonite. In brown scales ;
uniaxial or with a divergence of 1°±. (F.
A. Genth.) 1°±.
0. MineralHill,Del. Co., Pa. Pale green. (A.N. S.) 19°.
6. Lenni, Del. Co., Pa. Brown and green ; some-
times a very small optic angle occurs. 19°-20°.
*7. Culsagee, N. C. Culsageeite. Yellowish-brown:
variable angle. Sometimes the angle varies
as different portions of the same piece are
moved into the field. One piece gave 9°,
and another was nearly uniaxial. The angle
given is the most constant one. 20° dz.
8. West Chester, Pa. Jefferisite. Variable angle :
a specimen gave at one part 16°30', and at
another 25°, the latter being the most distinct ;
a very tliin piece gave 11°30', and a thicker
piece 2t°20'. Apparentl}^ the optic-angle in-
creases with the thickness of the plate. Some
good specimens gave 22°, 25°, and 28°; mean
angle probabl}^ 26°±:.
9. Lafayette Soapstone Quarry, Montgomery Co.,
Pa. Brown scales in chlorite slate : constant
angle 32°-36°30' ; mean, 34° ±.
10. Germantown, Phila. Brown plates in hornblende
rock. Optic-angle constant w^ithin 31°20'-
39°30' ; the most constant angle is 37°±:.
It is very probable that, as suggested by Prof. Cooke, the varia-
tion in the optic-angle of the Yermiculites is caused by twinning
1880.] NATURAL SCIENCES OF PHILADELPHIA. 251
Ripidolite.
1. Patterson's Quarry, Xewliu Township, Chester
Co., Pa. Irregular green plates; with corun-
dum ; inclination of bissectrix to normal to
cleavage plane, 5^30': optic-axial divergence
variable on the same plate on account of twin-
ning, varying from 50° to 59°30'. (T. D.
Rand.) Generally as given. 59°30'.
2. West Chester, Pa. Green plates ; inclination of
bissectrix 10° : axial divergence, 78°3'0'.
3. Brinton's Quarrj', Chester Co., Pa. Fine clear
green plates ; inclination of bissectrix, 12°30'.
f) y> V. Axial divergence, 82°.
4. Dudleyville, Ala. Pale rose-color; on chromite.
Inclination of bissectrix, 1G°. o > v. (F.
A. Genth). 94° 15'.
In all of these, double refraction is feeble compared with that of
Muscovite. It is observed that the inclination of the bissectrix
to the normal to the cleavage plane increases with the divergence
of the optic axes.
Prochlorite.
Brewster, X. Y., Tillj- Foster Mine. Uniaxial. 0°.
Margarite.
1. Cullakanee, N. C. White, "altered from corun-
dum." Irregular figures. (F. A. Genth.) 110°±.
2. Chester, Mass. Rose-color, with corundum ;
irregular, in some places showing four hj^per-
bolas; one piece gave 89°30'. 1I2°45'.
3. Dudle3'ville, Ala. White, clear; inclination of
bissectrix, 1°±. (F. A. Genth). 122°15'.
4. Cullakanee, X. C. White, " altered from Zoisite."
Inclination of bissectrix to normal to cleavage
plane, 2°ih. (F. A. Genth.) 124°.
The large optic-axial divergence of Margarite readily distin-
guishes it from Damourite and other micas which resemble it. If
further observations agree in showing that the bissectrix is inclined
to the normal to the cleavage plane, it will show that Margarite
is Monoclinic and not Orthorhombic as has been supposed.
252 PROCEEDINGS OF THE ACADEMY OF [1880.
October 22, 1871.
A New Locality for Analcite — Dr. A. E. Foote called atten-
tion to the fact that Analcite had been found at Falls of Schuyl-
kill,— a new locality for that mineral.
November 26, 1877.
On the Measurement of Plave Angles. — Mr. Lewis described a
simple and quick way of measuring plane angles in minerals. It
was a method which he had found very useful in the measurement
of all edge angles, of cleavage and striatiou angles, the angles
of markings and dendrites in mica, and of other flat angles to
which a goniometer could not conveniently be applied.
A paper protractor was constructed, the radii of which, distant
each from each 1°, were continued from the circumference
to the centre. Horizontal lines, about twenty in number, are
drawn across these, parallel to the radius 0° and at right angles
to the radius 90°. These lines being parallel, the angles formed
by the intersection of any radius with each of them are equal. In
order to measure the angle of a crystal, it is laid on the protractor,
one of its edges is made parallel to a horizontal line, and then the
crystal is slid along that line until the other edge, forming with
the first the angle to be measured, becomes parallel to one of the
intersecting radii. The desired angle is now read oflT on the cir-
cumference of the protractor. Angles approaching 90° are read
on one of the upper horizontal lines, while those of less amplitude
are read correspondingly farther down. A magnifying lens is
conveniently used to determine the exact coincidence of the edges
of the ciystal with the lines of the protractor. Very large crys-
tals as well as crj'stals as small as a millimetre in diameter can be
measured in this way.
It was found that this method of measurement was very conve-
nient, and, if the protractor had been carefully made, was exact to
within 30' ; while it applied to those cases in which neither the
reflective nor the hand goniometer could be used.
December 17, 1877.
On an Exfoliating Talc. — Mr. Henry Carvill Lewis described
a variety of talc, occurring at the soapstone quarry above Mana-
yunk, which is in some respects new. It occurs in fan-like crystals
in Dolomite, and is much more similar to Pyrophyllite than to
common talc. It moreover differs from common talc by exfoliat-
ing when held in the flame of a candle or Bunsen bui'ner, and
was, therefore, at first mistaken for Pyrophyllite. In the closed
1880.] NATURAL SCIENCES OF PHILADELPHIA. 253
tube it exfoliates and gives off water. In optical characters
it is identical with common talc, having been found to have an
axial divergence of about 12°40', frequently distorted. . It is
marked with striations or cleavage planes crossing at angles of 60°
and 120°. In this respect it is like Jettei'isite or Culsageeite,
while in common talc such markings are rarely visible, and never
distinct. It has the chemical composition of talc, except that the
percentage of water is larger than usual, being 7-02 per centum.
None of this water is hygroscopic, as its weight remains constant
in a desiccator over sulphuric acid.
The water of two other talcs from the same localit}^ was deter-
mined. A massive talc whicli does not exfoliate in the Bunsen
burner flame or in the platinum crucible, but does so at the point
of the blowpipe flame, contains 4.23 per centum of water.
A foliated talc which is caused to exfoliate onlv very slightly
even in the blowpipe flame, contained 2.84 per centum of water,
and this was driven oS only at an extremel}'' high and long
continued heat.
In these three talcs, therefore, we have the interesting results :
1. That there is a direct ratio between the amount of combined
water and the amount of exfoliation.
2. That there is a direct ratio between the tenacity with which
the water is held and the temperature at which exfoliation occurs.
It is thought that perhaps these results may have a bearing in
an explanation of the propei'ties of the various Yermiculites.
January 28, 1878.
Tin in Nor^fh Carolina. — Mr. Lewis exhibited a small piece
of tin ore said to have been found in Surry Co., N. C, and which
had been handed to him for examination. It was a soft, light
earth}' mass of a brown color, crumbling when pres^^ed, which,
when held in a candle flame, became covered with small globules
of pure tin. The earthy base was a silicate of alumina, iron, and
lime, and was partially' soluble in acid. The tin was reduced b}^
ver}^ gentle heat, for less than that required to reduce Cassiterite.
It was suggested that the tin existed in it either native or as an
ochre or hydrous oxide. No sulphides were present. It was
questioned whether the specimen exhibited was a genuine native
product.
A New Locality for Gypxum. — Mr. Theo. D. Rand announced
his discovery of gypsum, as an eftlorescence upon gneiss, at a
quarry near Darby, Pa.
254
PROCEEDINGS OF THE ACADEMY OF
[1880.
ON SIDEEOPHYLLITE— A NEW MINERAL.
BY HENRY CARVILL LEWIS.
A.mong other interesting minerals which are found in the neigh-
borhood of Pike's Peak, Colorado, is a hard black mica, occurring
sometimes in large and fine crj'Stals, which the writer has been
unable to identify with any known species.
It is monoclinic, and has an eminent micaceous basal cleavage.
It has the following characters :
Hardness, 3.2. Specific gravit}^, 3.1. Lustre, bright micaceous.
Color, black b}- reflected light, and fine chrome-green by trans-
mitted light. Opaque except in very thin pieces. Streak, pale
green. Laminae very brittle. Biaxial ; optic-axial divergence 10° ±.
In its composition it appears to be an iron-alumina mica. The
analysis here given is a mean of two made b}^ the writer. In one
the mineral was fused with sodic carbonate before solution, and in
the other it was dissolved in hydrochloric acid. The anal3'ses
were performed in the usual way. Iron was estimated by solution
in sulphuric acid in a closed flask, and subsequent titration. The
percentage of alkalies was kindly determined by Mr. F. A. Genth,
Jr. The percentage of water is that given off on moderate igni-
tion. On strong ignition the mineral loses over 3 per centum of
its weight, some of the alkalies being driven off.
0 ratio.
SiO,
A 1,0s
Fe,0:,
PeO
MnO
MgO
CaO
Na.,0
Li,0
K.,0
H,0
36.68
20.41
1.55
25.50
2.10
1.14
.81
1.09
.37
9.20
1.01
2.44
1.19 I
.06 j
.n
.06
.06
.03
.03
.02
.20
.11
2.44
1.25
1.22
2.00
1.02
1.00
This gives R
99.86
R : Si = 1
Silica 1
basic.
1:2, and for the ratio of bases to
1. It is therefore a Unisilicate in which the water is
1880.] NATURAL SCIENCES OF PHILADELPHIA. 255
It has the formula
and the symbol
Kg, Tl , OJ3, U,.>.
Before the blowpipe it fuses with intumescence at about 2.5 to
a black glass. It sometimes gives a red lithia color to the flame.
It is soluble in hydrochloric and sulphuric acids, with separation
of silica. In its pyrognostic properties it is thus similar to Annite,
although Annite is less fusible. Its oxj^gen ratio is that of Biotite,
but the absence of magnesia, and its physical and optical proper-
ties, distinguish it from that mineral. It occurs in good crystals
back of Pike's Peak, Colorado. Amazon-stone and Astrophyllite
occur in the vicinity. The material upon which this investigation
has been made was obtained from Dr. A. E. Foote, of this city.
The name of Siderophyllite {(Tidrfpoq ifolkov) has been given in
allusion to the large percentage of iron which it contains.
256 PROCEEDINGS OF THE ACADEMY OP [1880.
February 28, 1818.
On Sterlingite and Damourite. — Mr. H. C. Lewis stated that
an optical examination of a number of American damourites had
shown that they all had a large optic-axial divergence. This angle
was generality 72°-H°. It is an angle somewhat larger than that
of niuscovite, and is remarkabl}^ constant in different specimens.
On the other hand, the original damourite of Delesse has, accord-
ing to Descloiseaux, an optic-axial divergence of only 10°-12°.
No such angle has been found in any of the American damourites.
As it has been shown that damourite (" h^'dro-mica ") is an im-
l)ortant element in our rocks, and is of wide distribution and
frequent occurrence, it is essential that its characters should be
well known.
The damourite of Sterling, Mass., conforming precisel}', both as
to composition and structure, with the t^- pe of American damourite,
and which Prof. Cooke has shown to have an optic-axial divergence
of 70° ±:, has been named by Inm, Sterlingite. This distinctive
name was given solely on account of its larger optic angle. But
it appears that this large angle is characteristic of all American
damourites, and probably of many European ones.^ It therefore
follows either that all of our damourites should be called Sterlingite,
or that the name should be dropped ; there would otherwise be
confusion. Notwithstanding the exceptional optical character of
the mineral examined by Descloiseaux, it is thought that identity
of chemical composition and of physical properties is sufficient
reason for retaining the original name of Damourite.
March 25, 1878.
Vanadium in Philadelphia Bocks. — Mr. Lewis said that he had
discovered the presence of Vanadium in hornblendic gneiss near
Wayne Station, Germantown. The presence of sphene in that
rock suggested the search for vanadium, recent researches having
shown that this element frequently accompanies titanium. The
following method was employed for its detection. The pul-
verized rock was slowlj' heated in a crucible with sodic carbonate
and sulphur. After partial fusion the mass was digested in warm
water and the filtrate acidified. The precipitate was washed, ignited,
and fused with sodic carbonate and sodic nitrate. It was now
digested in water, filtered, the filtrate concentrated, and solid
ammonic chloride added. A precipitate fell, which was found
by blowpipe and other tests to contain pure vanadium. An
exfoliating hydrous mica occurred at this locality, resulting
^ V. " The optical characters of some Micas :" by 11. C. Lewis, Proc.
Min. and Geol. Section, October 22, 1877.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 25Y
perhaps from the alteration of hornblende, and which was believed
to be a new species, in which there was .38 i^er centum of oxide of
vanadium-
A New Locatity for Epsomite. — Mr. Lewis reported having
found Epsomite in Sideling Hill Tunnel, E. Broad Top R. R.,
Huntington Co., Pa. It there occurs in small, colorless, acicular
ciTstals in an olive-colored shale in the lower part of the Vesper-
tine formation (No. X).
258 proceedings of the academy of [1880.
November 25, 1818.
the surface geology of philadelphia and vicinity.
by henry carvill lewis.
At intervals during the past year the writer has been devoting
some attention to the gravels and clays of our city, and although
the work as yet is only preliminary, and is still in progress, it is
thought that a sketch of what has been done ma}^ serve to show
what an interesting field is open for more thorough investigation.
A large number of localities have been examined and many sec-
tions have been made, but it is proposed at present merely to
summarize the facts observed.
The Upland Terrace. — 1. A traA'eller going from the city upon
the Germantown Railroad will notice in the cuttings for new
streets between Tenth and Broad Streets, and in the railroad cut
at New York Junction, numerous exposures of red or yellow
gravel, often overlaid by clay. The brickyards in the vicinity of
Nicetown expose large beds of brick-clay containing occasional
well-rounded boulders and pebbles. The land so far has been
comparatively level, and no rocks have been seen. Just before
reaching Wayne Station, rocks rise upon both sides of the road,
the clay and gravel disappear, and a rolling wooded country is
entered. A thin covering of light micaceous soil containing no
pebbles or boulders covers the gneissic rocks from here to Chest-
nut Hill. There is a great contrast between the two regions.
2. On the Pennsylvania Railroad it will be noticed that, soon
after leaving the depot, gravel covers the rocks along the Schuyl-
kill, and as the railroad turns back from the river, a plateau of
clay follows. The Centennial grounds lie upon this clay, and
boulders are frequent. Upon reaching Fift^^-seventh Street, op-
posite Belmont and George's Hill, the hill is entered by a cut, the
rocks come to the surface, and the drift is no more seen.
3. Again, on the North Pennsylvania Railroad gravels first
appear, then, on higher ground, clay, and soon after passing Green
Lane Station, the rocky uplands, free from drift.
4. So, too, on the West Chester Railroad, gravels and clays
cover the ground up to the base of the hill on which Swarthmore
College stands.
5. On the otlier hand, the New York division of the Pennsyl-
vania Railroad and the Philadelphia, Wilmington and Baltimore
1880.] NATURAL SCIENCES OF PHILADELPinA. 259
Railroad, which run parallel with the Delaware River, do not
rise out of the region of drift.
Now, connecting by a line the four points mentioned, it will
be found to represent a long straight hill 200 feet or more in
height, having a northeast and southwest trend, pai'allel to the
river, and lying at a mean distance from it of about four miles.
We have traced it from Bucks County, thi'ough Philadelphia and
Delaware Counties, into the State of Delaware, and find that it
uniformly defines the western boundarj^ of the drift. This hill is
easily recognized where uncrossed by creeks, being remarkably
straight and of uniform height. It forms the limit of tidewater,
and is recognized where it crosses streams by the occurrence of
rapids or falls. Being the first hill of importance west of the
Delaware, it often commands a fine view and is a favorite site for
residences. The geographical position of this ancient terrace
may be more exactly defined in the vicinity of Philadelphia, as
the hill which crosses Second Street Pike near Foxchase, and
crossing Tacony Creek farther south, runs nearly parallel with
it as far as Crescentville ; which crosses Green Lane and New
Second Street road near the place of Mr. J. L. Fisher ; crosses the
North Pennsylvania Railroad above Olney road, and the York
road below the Jewish Hospital ; which crosses Germantown
Avenue at the railroad bridge (being here called Negley's Hill),
and running along the railroad to beyond Wayne Station, passes
back of the Germantown Cricket Ground, past Old Oaks Cemetery
to Falls of Schuylkill. Thence, passing Chamouni, Belmont, and
Geoi-ge's Hill, it crosses the Pennsjdvania Railroad near Heston-
ville, and Haverford Road at Haddington ; passes back of the
Burd Orphan Asjdum into Delaware County, and runs north of
Kelleyville, Clifton and Morton to Swarthmore College, and
thence past Village Green into Delaware.
This hill, which is approximately parallel not only to the river,
but also to the shore of the Atlantic Ocean and to the line of
strike of the Cretaceous formations of New Jerse}^ forms, as we
have seen, the main dividing line between the ancient and the
modern formations.
We shall call it for convenience the Upland Terrace. The strike
of the gneiss forming it corresponds closely with the trend of the
terrace itself. A boulder-bearing clay rests upon its southeastern
slope at a uniform elevation of 150-170 feet above mean ocean-
level. While it is true that, as will appear hereafter, there are
260 PROCEEDINGS OF THE ACADEMY OF [1880.
patches of an ancient gravel on high points back of it, the Upland
Terrace nevertheless remains as the most important geological
feature in sontlieastern Pennsylvania.
Between the Upland Terrace and the Delaware, claj^s and gravels
cover the rocks in a continuous sheet except where eroded away
in the neigliborhood of streams. The amount of tlieir erosion is
in some respects a measure of the age of the surface formations.
It has been noticed that these foi'mations in the vicinit}' of Phila-
delphia have undergone very different amounts of erosion, the
amount of such erosion increasing as we recede from the Delaware ;
and this fact is regarded as offering evidence that the deposits are
of different ages ; those lying farthest from the river and highest
in elcA^ation being the most ancient, and those which are close to
the river, which have undergone but little erosion, being the most
modern of our surface formations. Examples of erosion of the
Philadelphia gravel may be well seen on the Philadelphia and
West Chester Railroad which crosses a number of creeks and runs
nearlj'' parallel to the terrace for several miles. As each creek is
approached the drift ' disappears and rocks come to the surface.
So on the Schuylkill, no gravel is seen on the river drive in the
East Park, but upon going back from the river and rising 100 feet
above it, as far as the East Park Reservoir, gravel appears abun-
dantly. Yet on the same riA^er, nearer the Delaware, a newer
gravel, made of different materials, not only forms its banks but
underlies it.
Recent Alluvium. — The most recent of all the surface deposits
is the stiff bluish clay which covers the low ground in the southern
part of the city. The Richmond meadows and the flats of Moya-
mensing, Greenwich and Tinicum are covered by this deposit. It
is bounded by a low terrace which may be called " The Floodplain
Terrace.'''' This terrace, up to which the river often comes in
times of flood, crosses South Broad Street diagonall}^ below Moya-
mensing Avenue, and crossing the Delaware extension of the
Pennsylvania Railroad near Penrose Ferry Road, winds around
Point Breeze Park back towards the Gas Works, and passing
below Suffolk Park crosses into Delaware Count3\ This terrace
is about ten feet above mean tide. It is the lowest and newest of
all the terraces and is formed of the next older formation, the
" River gravel." The mud or clay lying between this terrace and
' The term "drift" here includes all superficial formations of whatever age.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 261
the river is too stiff to 1)6 useful for brickmaking. Blackened
fragments of twigs, roots and leaves are frequent in it, and it is
said that trunks of the white cedar abound in it in some places.
There is here an indication that these beds are sinking and that,
as on the Atlantic coast, the water is encroaching. Frequently
a good peat covers the clay.
The River Gravel. — Forming the Floodplain Terrace and lying
back of it, is a light sand and gravel free from claj', which may
be designated the " River Gravel," since it formed the ancient river
bed. It is composed of a light micaceous sand made from the
wear of gneissic rocks, ovcrl3'ing a clean, loose gravel, whose peb-
bles are composed of the rocks which form the river bottom farther
north. The pebbles are generalh' flattened and are composed of
gneiss, Triassic red shale, Triassic argillite, etc. It is of a gray
color, white quartz pebbles being comparatively^ scarce. It under-
lies the river to a great depth and forms islands in it. Frequently
large boulders lie upon the river gravel. Bridesburg and the
Lazaretto are built upon it. The sand is used for building pur-
poses. It is bounded by the " i?u-e?- Gravel Terrace," a terrace
rising some twenty feet above mean tide, and which is capped by
the red gravel and brick-clay about to be described, while rocks
are frequently exposed at its base. The Chester Branch of the
Reading Railroad lies below this terrace, and the present line of
the Philadelphia, Wilmington and Baltimore Railroad is above it.
The Bed Gravel and Brick-Clays.— T\\q built-up portion of the
city stands upon an extensive deposit of brick-clay and gravel,
sections of which are exposed in ever}' cutting. The brick-clay
invariably overlies the gravel, and will therefore be first described.
By far the finest exposures of brick-clay are those on either side
of Long Lane, in the "Neck." The clay here is ver}" compact,
free from sand and gravel, and is often 15 feet or more in depth.
Loam lies above it, and is mixed with it for brick-making. Well-
rounded boulders of Potsdam, Medina, Trias, etc., are frequent.
The whole lies upon some 20 feet of stratified gravel. It is a
much finer and deeper clay than that of the northern part of the
city, as at Nicetown. It is interesting to note that while the clay
which is farthest from the Upland Terrace and lowest in elevation
is purest and deepest, on the other hand that near the terrace
and more than 100 feet above the river is both shallow and sand3\
It suggests that the former was deposited in deep water and the
latter near the shore. At the base of the terrace the clay is but
262
PROCEEDINGS OF THE ACADEMY OF
[1880.
two or three feet deep. The boulders of the Nicetown clay are
similar to those of the Neck clay, except in the fact that in the
latter there are numerous rounded and sharp fragments of triassic
red shale, while in the former boulders of that material are very
scarce. The boulders of both clays are invariably derived from a
northern source, ^o shells or organic remains have as 3'et been
noticed in this formation.
Beneath the clay, and often unconformable with it, is the Phila-
delphia red gravel. It is a claye^'^ gravel which packs well and is
much used on roads, and whose red color is caused by the ferru-
ginous clay in which the pebbles are imbedded. The pebbles are
composed of all kinds of rock and are not flattened as are those in
the river gravel. The predominant material is white quartz, but
pebbles of all other materials, as conglomerate, sandstone, fossil-
iferous hornstone, flint, red shale, etc., are numerous. Stratifica-
tion is observed in almost every section exposed. Good sections
of gravel are seen near the University of Pennsylvania. It has
here an elevation of about 50 feet, and comes to the surface of the
ground with but a very slight covering of clay. The gravel is
here over 15 feet deep, and as it is in some respects a typical
exposure, a section is herewith presented.
Ft. Fig. 1.
10
1
5
3
1
+
BRICK-CLAY.
RED GRAVEL.
BLACK GRAVEL.
YELLOW GRAVEL.
MICACEOUS SAND.
BOULDERS.
DECOMPOSED GNEISS.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 2G3
It will be noticed, in the first place, that the clay lies in the
form of crests and hollows upon the gravel. This is almost inva-
riablj'^ the case. Beautiful examples of wave motion may be seen
at Twenty-eighth Street and Columbia Avenue, at Tenth and
Tioga, at Fifteenth and Clearfield Streets, and in Fairmount Park.
In each of these we have apparently the action of a rushing flood
of water upon the gravel. Often the clay lies in a kind of pot-hole
in the gravel, and a concentric structure of clay and pebbles can
be seen. The following section, at Twent3'-eighth Street and
Columbia Avenue, shows six well-marked waves of gravel and
clay, the clay always filling the hollows between the crests of
gravel.
Fig. 2.
s N
The approximate dimensions of the weaves are given in the
diagram. Along the line of contact between clay and gravel there
are alternate streaks of fine and coarse gravel.
A ver}^ beautiful example of water action is exposed at Fifteenth
and Clearfield Streets, in a cut about 100 feet in length (Fig. 3).
The second point to be noticed in the section near the Univer-
sity is the stratification of the gravel, and its division into layers
of three diflTerent colors, — red, black and yellow. It is instructive
to note that this division is by no means a local one, but exists
along a line of about equal elevation (60 to 80 feet above ocean level),
in widely separated parts of the city. While the colors are of
course due simply to different states of oxidation of the iron, the
fact that the}^ mark continuous deposits through long distances,
indicates a uniformity in the condition of deposition which could
be due only to the presence of a large bodj^ of water.
In the third place, the section (Fig. 1) shows the important fact
that the gravel rests, not upon a hard floor of rock, as is usual with
the drift in more northern States, butupon a completely decomposed
gneiss. This is universall}' the case in every section examined in
the vicinity of Philadelphia. In no case does the gravel rest upon
264
PROCEEDINGS OF THE ACADEMY OF
[1880.
unaltered rock, except possibly in the
vicinity of streams where the water has
eroded away the soft rock. In such posi-
tions, however, the gravel also is usually
eroded, except in the case of the " River
gravel " proper. It will be observed that
a sharp stratified micaceous sand, made up
of the materials of the decomposed gneiss,
and often showing " flow and plunge "
structure, lies below the gravel. In the
section given, a well-rounded boulder of a
lower Silurian sandstone is seen partially
imbedded in the decomposed gneiss. This
fact offers two interesting deductions : —
(1.) That the gneiss was decomposed
before the deposition of the gravel.
(2.) That water, not ice, was the agent
of such deposition.
(1.) As additional evidence in support
of the first deduction, it has been observed
in several sections that portions of the de-
composed gneiss have been taken up and
interstratified in horizontal layers, either
with the gneissic sand, or with the gravel
itself That the steeply-dipping decom-
posed gneiss should be thus re-stratified, as
though by a flood, and that, on the other
hand, no such phenomena are ever observed
in undoubted glaciated regions, can only be
explained upon the assumption that the
gneiss was decomposed before the Glacial
epoch. That such decomposition took
place in a j^et earlier geological age, will be
indicated below under a description of the
" Bryn Mawr gravel."
(2.) Absence of a glacier in this region
is indicated by the wave-like junction of
gravel and clay, by the stratification of the
gravel, and by the presence of decomposed
gneiss. No polished surfaces of rock have
Fig. 3.
1+
1880.] NATURAL SCIENCES OF PHILADELPHIA. 265
ever been observed in this region, although the occasional slicken-
sides upon the gneiss in some quarries has been mistaken for
glacial strite. Frequently the lower yellow gravel is replaced by
a 3-ellow sand more or less fine, which is used for building purposes ;
and in this there are often good examples both of oblique lamination
and of " flow and plunge;" — structures attributable to flowing
water. Examples ma}'^ be seen on the North Penna. R. R. and in
the East Park. Tlie boulders of both clay and gravel, if not
brought down by water alone, have been dropped by floating ice.
The absence of life in either deposit indicates that the water was
too cold to support it.
The conclusion is therefore forced upon us tliat, during the
melting of the great Northern Glacier, whose southern terminus
crossed the river probably near Belvidere, the flooded Delaware,
then a great torrent five or ten miles wide and at least 150 feet
deeper than it is now, deposited at first gravels and afterwards,
when quieter, clays; while floating ice carried down already rounded
boulders and dropped them upon its bed.
The uniform elevation of the edge of the clay at the base of
the Upland Terrace can hardly be accounted for upon another
hypothesis.
The presence of an actual glacier over this region has, however,
been brought forward as the onl}^ explanation of our surface
deposits. Thus, in a recent paper, ^ the author, after inspection
of a gravel opening in West Philadelphia, concludes " that this
belt of drift deposit is no other than a glacial moraine formed by
the Schuylkill glacier receding from the site of the cit^." He
adds, " the surface of the gneiss where laid bare is comparatively
smooth, and shows evidence of having been polished, though so
soft as not to retain the marks of glaciation."' To us the very
locality described (Forty-fifth and Spruce) off'ers strong evidence
of the absence of all glacial action. The gravel, containing no
scratched pebbles, is horizontally stratified and shows flow and
l)lunge structure ; while the underlying decomposed gneiss, so far
from being polished, is seen in several places to have been taken
up by the swiftly flowing water and mingled with the gravel which
it bore along, so that several layers of decomposed gneiss, each
about half an inch in thickness^ and soon dying out, alternate with
the lower portion of the gravel.
' "On Glacial Depoi^itsat W. Pliila.," Pioc. Am. Fhil. Soc, Nov., 1875.
18
266 PROCEEDINGS OF THE ACADEMY OF [1880.
It has been supposed that the bending over of the outcrops of
steeply-dipping rocks, sometimes observed near Philadelphia, has
been caused by the pressure of a glacier. A very beautiful example
of such broken and l)ent-over strata is seen in a quarry at Edge
Hill. That such phenomena are to be explained, not by glacial
agencies, but by the force of gravitj^ onl}', — being the gradual
sliding-down-hill of the soil known as " creep," — is shown by the
facts, (1) that such bending over is always towards a lower eleva-
tion,— down hill; (2) that on the two slopes of the same hill the
strata have been seen to be bent over in opposite directions.
Thus at various points along the long ridge of altered Primal
slates known as Edge Hill, the slates on one slope are bent towards
the south, and on the other towards the north. A similar fact
has been noticed in the gneiss forming the Upland Terrace.
Moreover, such bending of the strata often occurs in regions quite
free from drift.
If, as we have conjectured, the Delaware Valley was filled with
a large body of water when the drift was deposited, it is reason-
able to suppose that the Schuylkill also was of far greater size, and
that some boulders would be brought down the valle}' of that
stream. Here again facts sustain the hypothesis. In the gravel
taken from the excavation for the East Park Reservoir, associated
with Triassic red shale and other boulders, we have found partiall}'^
worn fragments of chlorite slate containing octagonal crystals of
magnetite, evidently derived from the steatite quarry at Lafayette,
six miles above on the Schuylkill. At Twenty-eighth Street and
Columbia Avenue is a large boulder of trap, identical with that of
the trap-dyke which crosses the Schuylkill River at Conshohocken.
It thus appears that during the Glacial epoch the waters of the
Schuylkill em})tied into those of the Delaware at Falls of Schuyl-
kill, the city proper being entirely submerged.
Before closing our account of the Philadelphia red gravel — the
" University gravel," as it might be called for distinction — it will
be necessary to sa}' a word as to what occurs on the New Jersey
side of the river. If we are correct in ascribing this gravel and
brick-clay to a flooded river valley, similar deposits at the same
elevation must be found on both sides of the river. Although we
have been able to do but very little work upon this point in that
State, it has been obserA^ed : (1) That there is a sand at Camden
near the river, si.nilar to the sand of the " River gravel " of lower
1880.] NATURAL SCIENCES OF PHILADELPHIA. 267
Philadelphia ; (2) that at a higher elcA'ation there are deposits of
superficial yellow brick-cla}' quite distinct from the underlying
plastic clays ; (8) that boulders identical with those on this side
of the river occur in the brick-clay ; (4) that a stratified red gravel
containing Triassic shale, and similar to the University gravel
occurs ; and, (5) that there are indications of the existence of a
Terrace, several miles from the river, bounding the brick-clay and
its boulders, and composed of an older, and probably oceanic,
gravel and sand.
The Fossiliferoiis Gravel. — There seems to be evidence that
between the Upland Terrace and the River Gravel Terrace there
is an intermediate terrace, back of which is a gravel somewhat
different from the Philadeli)hia red gravel. It is characterized by
comparative absence of Triassic red shale, and by the presence of
numerous pebbles of flint, hornstone, or limestone, which are fre-
quently fossiliferous. These pebbles, as well as those of white
quartz, are not fresh-looking, but are eaten and weather-worn by
age. In both its position and its appearance it is an older forma-
tion than the red gravel. It is of a 3'ellowish color, becoming-
white when exposed to the weather, and is more sandy than the
red gravel. For these reasons it is less esteemed for road-making.
The Germantown Railroad cuts through this gravel at New York
Junction, We have found here pebbles containing Cyathophyl-
loid corals, Favosites, a Trilobite, etc. The Connecting Railroad
at Ridge Avenue Station cuts through the same gravel, and here
we have found Strophomena^ etc. Other fossils have been found
below the ela}' in the East Park and at the Centennial Grounds.
This gravel is found on the high level plateau which lies at the
base of the Upland Terrace, and is covered b}^ more recent brick-
cla3^ It lies farther from the river and at a higher elevation than
the red gravel, and there is a decided rise in the ground from the
latter to the former. This terrace has been observed in many
places near and in the city, but has not as yet been traced con-
tinuously, and its existence is doubtful. Nearly- all the brick-3'ards
in tlie city, exceijt those in the " Neck," lie upon this gravel and
back of this terrace, which lies at a mean distance of about a mile
inside of the Upland Terrace. It seems as though the flood,
diminishing in breadth, had eroded away the clay within this •' Red
Gravel Terrace." The red gravel comes to the surface, with
very little overlying cla}', at elevations below about 100 feet;
while at a higher elevation is the brick-yard plateau.
2C8 PROCEEDINGS OF THE ACADEMY OF [1880.
The Pennsj'lvania Hospital for the Insane stands upon the hill
forming this inner gravel Terrace. Its course is somewhat parallel
to the main Upland Terrace, and it crosses Walnut Street near
Fiftieth Street, and Broad Street near the Reading Coal Road cross-
ing. In Prof. Rogers' Geological Map of Pennsylvania, where a
rough attempt is made to represent the boundary of the drift, the
line in one place corresponds quite closely with what we have pre-
sumed to be the "Red Gravel Terrace ; " but it appears that in most
places in that map the boundary is meant to be merely a hypoth-
etical one. While the existence of this inner terrace is yet doubt-
ful, and while it is probable that red gravel will be found above it and
fossiliferous gravel below it, yet nothing has yet appeared to con-
trovert the assumption that the latter gravel is older than the
former. How much older, and whether of oceanic or of fresh-
water origin, is not yet determined. Here, again, a study of the
New Jersey gravels will be of assistance.
The Branch town Clay. — Having now described the surface
deposits lying between the Delaware River and the Upland Ter-
race, it remains to point out the existence of some isolated patches
of gravel and clay which have been noticed on some of the hills
back of and above this terrace.
In the village of Branchtown, on a plateau 250 feet above the
river, there is a local deposit of brick-cla}' \yh^g in an oblong belt
runnhig N. E. and S. W., perhaps a mile in length and an eighth
of a mile in breadth. That it is not a clay formed in place by
decomposition of the gneiss is shown hy the presence in it of
pebbles and rounded boulders of foreign rocks. The smaller
pebbles consist of quartz, and the larger of a friable quartz sand-
stone, prob?.bly Potsdam. Not a single fragment of Triassic red
shale, and not a single pebble of flint or fossiliferous rock was
found: and in this it is distinguished from any deposit heretofore
described. Nor were an^^ of the pebbles formed of the materials
of the bed of the Delaware River. Numerous sharp fragments, often
six inches square, of white or yelloAv siliceous sandstone and of
brown jaspery quartzite, both probably of lower Silurian age, were
f'jund. The peculiar conglomerate described below as "Mt. Holly
Conglomerate" does not occur. Decomposed gneiss lies below
the cla}^., which is two to three feet deep. The presence of sharp and
rounded boulders of a rock in place farther north suggests an
overland flood during glacial times ; but the complete absence of
1880.] NATURAL SCIENCES OF PHILADELPHIA. 269
all traces of Triassic red shale, a formation of large extent six miles
north of here, over which such a flood must have passed, is difficult
to explain upon that hypothesis. This belt of cla}', which may be
called for convenience the " Branchtown clay," extends S. W.
to Chelton Avenue and Chew Street, in Gerniantown, and to the
N. E. to Limekiln Pike and City Line Road, and is the site of
several brickyards. The clay plateau is bounded on the N. W. by
a hill 325 feet high. Doubtless this clay will be found in other
places, when more light will be thrown upon its origin and age.
The Bryn Maior Gravel. — Upon the summits of some of the
highest hills in the gneissic region back of Philadelphia, at a mean
distance of about nine miles from the river, and at elevations of from
325 to 450 feet above it, there are isolated patches of an ancient
gravel, different from any yet described, to which we have given the
provisional name of " The Brj'n Mawr Gravel." It can always
be recognized b}- the presence of sharp or partially rounded frag-
ments of a hard, heavj^ iron sandstone or conglomerate. Such
fragments are often covered by a brownish-black iron glaze. More
than ten years ago, the writer noticed in the soil of the upper part of
Gerraantown, pieces of this conglomerate, unlike any known rock,
and it is only of late that its origin has been suspected. It con-
sists of well-rounded pebbles of quartzite or siliceous sandstone
cemented by iron into a stone which is often very hard. This
conglomerate is found in occasional fragments upon ground over
300 feet high, but is not found in abundance until an elevation of
over 400 feet is reached. At these highest points it occurs in a
red gravel wliose pebbles are identical with those of the conglom-
erate.
One of such points is near Chestnut Hill, on the City Line
Road at its highest elevation, near Willow Grove Road. Here,
nearly nine miles from the river and 425 feet above it, is a patch
of this gravel and conglomerate. The larger pebbles and boulders,
like those of the Branchtown Cla}', consist of a friable quartzite
sandstone or a jaspery quartzite. Sharp fragments of quartzite
are numerous ; but there are no traces either of Triassic red shale,
of fossiliferous pebbles, or of rounded pebbles of the uiiderlying
gneiss. It rests upon a much decomposed gneiss. The conglom-
erate sometimes contains cavities filled with white sand. The
tract of gravel is of an oval form, whose major axis points N. E.
and S. W. It crosses the Township Line Road near the Bethesda
2T0 PROCEEDINGS OF THE ACADEMY OP [1880.
Home, near which place have been found a sharp boulder of conglom-
erate three feet in diameter, several fragments of ferruginous sand-
stone equally large, a partially rounded boulder of white quartz
nearly four feet long, and numerous fragments of quartzite and
Primal rocks. The gravel is here in part replaced by clay.
A similar tract of this gravel occurs at Bryn Mawr, extending
from that place to near Cooperstown. A good section is exposed
in the railroad cut below the station. From this locality, so easy
of access from the city, we have named the formation. It is
here about 430 feet high, and nine miles from the river. The
gravel is ten feet deep, and lies upon a steepl^'-dipping gneiss so
completely decomposed that it is as soft as cla3^ Underneath the
bridge, a soft Avhite kaolin-like material, conformable with the
gneiss, shows a decomposed steatite,^-being probably the con-
tinuation of that which crosses the Scimylkill at Laftiyette. Here,
as at Chestnut Hill, the gravel lies in an isolated patch upon a hill,
distant from any stream or other eroding agency. The gravel
holds sharp fragments of primal rocks and also the iron con-
glomerate. As at Germantown, the fields below, to the south,
contain occasional fragments of the conglomerate.
Another good exposure of the Bryn Mawr gravel is on a hill
crossed by the road leading from Haverford College to Coopers-
town. The conglomerate is here in large, sharp fragments, and
the gravel shows slight horizontal sti'atification. On the crest of
the hill, some 450 feet high, there is a weather-worn boulder,
four feet in diameter, of a soft, coarse, brown sandstone of Br^'n
Mawr age, apparently in place.
A fourth, precisely similar exposure of gravel Avith conglom-
erate, and at about the s:ime elevation, caps the hill back of Media,
near the lloseti'ee.
Without describing any further exposures, it already appears
that in these elevated patches of ancient gravel we have the last
remnants of a once continuous formation. The very great erosion
which has swept away all but these few traces is a sufficient proof
of its age. There are no points at all approaching the elevation
of these hills, between them and the Atlantic Ocean ; and it is at
once suggested that these patches are the remnants of an oceanic
deposit, possibly of Tertiary age. It is interesting to find that a
precisely similar formation caps some of the hills in New Jersey.
On top of the hill at Mount Holl}^, N. J., is an identical con-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 2tl
glomerate and gravel, similar in appearance, and composed of
the same materials as the formation in Pennsylvania. Tlie con-
glomerate has the peculiar ferruginous glaze already noticed.
It here overlies Cretaceous marls aud sands.
From its abundance at this place, and in order to show its con-
nection with Pennsylvania deposits, we shall call the conglomerate
of the Biyn Mawr gravel, " Ji?. Holly Conglomerate.'''' Prof. H.
D. Rogers' suggests that this rock at Mt. Holly may be of Mio-
cene age ; but Prof. Cook, not distinguishing it from the modern
iron crusts in the red Philadelphia graAel near the river, considers
it veiy recent. In the consideration of its age it is worth noting
that the sand of southern New Jersej^, apparently of late Plio-
cene age, frequently contains rounded pebbles of Mt. Holly con-
glomerate, thus showing that the latter is an older formation.
From the identit}^ of their contained boulders, it is probable
that the Branchtown cla}' and the Br3'n Mawr gravel are nearly
coeval. Being oceanic, it is presumed that they will be recognized
all along the gneissic hills of the southern Atlantic States.
We have given this detailed description of each of the surface
formations near Philadelphia in the hope that they may be recog-
nized elsewhere by other geologists. It has been found that a
careful examination of the materials comprising each gravel, taken
in connection with their elevation above tide, is the onl}- means of
- discriminating between them. Desultory observations in detached
localities are of little value. Should this work be extended in
Penns^dvania and New Jersey, and the distinctions between the
four gravels described be carried out, it is thought that, notwith-
standing the shifting character of the underlying strata in the
latter State, much may be done not only towards an exact deter-
mination of their age, but towards a settlement of some of the
vexed problems of surface geology in Eastern America.
Recapitulation. — The results obtained may be briefly summa-
rized as follows: —
Forming the N W. bound.iry of the Philadelphia gravel and
brick-cla}' is a hill of gneis?, rising 200 feet or more above the
river, which ma}^ be called the Upland Terrace. It has a N. E.
and S. W. trend, and in this vicinity is at an average distance of
five miles from the river.
1 Report ou the Geology of N. J., 1839.
272 PROCEEDINGS OF THE ACADEMY OP [1880.
Within the Upland Terrace, resting upon its slope, and extend-
ing to the river, is a series of stratified gravels and a boulder-
bearing brick-clay. Of these, the oldest is the " Fossiliferous
gravel ; " a gravel l^^ing near the terrace and under the brick-clay,
and containing pebbles which frequently are fossiliferous. Of more
recent age, and at a lower level, is the " Philadelphia red gravel,"
which is made up of the pebbles of the Fossiliferous gravel
mixed with fragments of Triassic red shale and other rocks
brought down the Delaware Valley. It is distinctly stratified,
rests upon decomposed gneiss, and contains rounded boulders
dropped by floating ice. Upon both of these gravels rests the
Philadelphia brick-clay, often l^ang unconformably upon them in
a series of pot-holes or wave-like forms, and apparently an aque-
ous deposit.
A yet more recent formation, the " River gravel and sand,"
lies within the others and close to the river, and is made up of
flattened pebbles composed of the rocks over which the river flows.
Upon this, in the river flats, lies a modern mud, the " Recent Allu-
vium."
Back of the Upland Terrace, isolated patches of two surface
deposits, more ancient than any 3ret described, lie upon the hills.
These are, the " Branchtown clay," at a height of 250 feet, con-
taining boulders of Potsdam rocks, but no traces of Triassic red
shale or of fossiliferous pebbles ; and the " Bryn Mawr gravel,"
which caps hills of a higher elevation, and which, containing
boulders and pebbles of identical material with those of the last,
is characterized by the presence of a hard iron conglomerate or
sandstone. This conglomerate, occurring also in New Jersey,
and named the " Mt. Holly Conglomerate," is conjectured to be of
Tertiary age.
In these seven formations is written the geologic history of the
Delaware Yalley.
Much remains to be done before any certain results can be
expected. It is hoped that the imperfect examination here
recorded may form the basis for a future and more thorough
study, which, extending to wider fields, shall make more exact
the knowledge of our surface geology.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 2t3
October 28, 1878.
'On a Belt of Steatite and Serpentine in Radnor, Del. Go. — Mr.
Theu. D. Rand read a paper on a belt of Steatite and Serpentine,
in Radnor Township, Delaware Co., Pa.
(Publisiied in Proc. Acad. Nat Sci.)
November 25, 1878.
Chromite near Radnor, Pa. — Mr. Theo. D. Rand annonneed
the occurrence of Chromite in considerable quantity in the
Southerly Serpentine Belt, near Radnor Station, Delaware Co., Pa.
274 PROCEEDINGS OF THE ACADEMY OF [1880.
February 24, 18*79.
ON RANDITE.
BY THEODORE D. RAND.
At the December meeting of the Mineralogical Section, Mr.
Goldsmith made a communication in regard to the uranium-yellow
coating found at the south end of the largest quarry at Frankford,
northeast of Adams Street, stating that he found in it, carbonic
acid, silicic acid, phosphoric acid, uranium, alumina and lime;
and that his conclusion was, that it was a mixture of autunnite
and calcite. The writer stated at the same meeting that he
had made an incomplete examination of the same mineral, which,
in great part, confirmed Dr. (roldsmith's observations, but that he
failed to find phosphoric acid, and promised the Section the result
of experiments then under way.
At the meeting of the Academy held December 31st, 1878, Dr.
Koenig communicated the results of a full quantitative analysis,
giving the composition, a hydrous carbonate of uranium and lime,
to which he gave the name Kandite.
The writer's results differ somewhat from those of Mr. Gold-
smith and Dr. Koenig. Owing to the very small amount of the
coating, and its close adhesion to the rock, proper separation was
impossible, and the first experiments were made by treating the
rock and coating, first with acetic acid, to remove calcite, then
with dilute hydrochloric acid. The coating was unaffected In* the
acetic acid, as proven by one specimen, in which, after solution of
a large amount of calcite, the Randite was left in tufts of acicular
crystals. The acetic solution contained chiefly lime, with a little
alumina, but no uranium.
The hydrochloric solution yielded a small amoimt of silica,
alumina, sulphuric acid, and phosphoric acid, with a large amount
of lime and uranium.
In the treatment with acetic acid, bubbles appeared to rise from
the coating — a multitude of tiny bubbles ; on the succeeding treat-
ment with hydrochloric acid, the bubbles were much larger, and
fewer in number, and appeared to rise from a cai"bonate in the
crevices of the rock.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 275
The proportion between the lime and uranium may be given as
follows :
Koenig. Rand, 1. Rand, 2.
Lime, 50 38 26
Uranium, 44 G2 14
10.708 gm. of coated rock, after treatment with acetic acid,
yielded to 8 p. c. liydrochloric acid, cold, in about five minutes
(the coating having disappeared), .122. On evaporating the
solution to dryness there was a residue less than .001 gm. The
solution Avas precipitated by ammonia, in the presence of chloride
of ammonium ; the solution with oxalate of ammonia gaA'^e car-
bonate of lime, .0365. The precipitate treated with acetic acid
dissolved wholly, except .001 of a white precipitate, which con-
tained phosphoric acid, and was probably phosphate of alumina.
The solution pi-eeipitated by phosphate of soda gave jjhos. uran.,
.0711 =U20;, .0569.
Per cent.
Uranic oxide, .0569 46.71
Lime, .0204 16.71
Phos. al? .001 .89
Undetermined, 35.69
100.
About 100 grams of the rock, free from the coating, were treated
with acetic acid in excess. A large amount of lime was dissolved,
and a trace of alumina. The residue, treated with hydrochloric
acid, yielded a little silica, some alumina, and considerable lime.
I infer from these tests that the mineral has not the composition
obtained b}^ Dr. Koenig, and that further investigation is needed,
if pui'e material can be obtained.
276 PROCEEPINGS OF THE ACADEMY OF [1880.
March 24, 1879.
Some Microscopic Enclosures in Mica. — Mr. Theo. D. Rand
described, and exhibited under the microscope, certain crystals,
etc., included in mica, chiefly from Swain's quarry, Chester Co. Pa.
Of these, the magnetite dendritic markings, and similar mark-
ings of red and brown colors, apparent!}' due to oxidation of the
magnetite, are most common and best known. Besides these the
following occur : —
Hexagonal crystals, black and opaque ; angles, 60° and 120°. In
the form of the crystal in this description, the form of the section
exhibited under the microscope is intended. A similar crj'stal,
brown in color, perhaps the same substance, translucent ; probably
biotite or lepidomelane.
Hexagonal or rhombic crj- stals of a bright red color, sometimes
with the angles modified; angles (;0° and 120°. There are some
specimens which indicate the change of the black into the red
rhombs. One of the red rhombs contained a black crystal, with
faces parallel to those of the red, and one, a very symmetrical and
simple crystal, from near Newtown Square, Delaware Co., Pa.,
was black for about one-fourth its length, the remainder red.
Rhombic crystals, polarizing light, giving very brilliant colors.
At first this was supposed to be due to films of the miea itself, but
the regularity and brilliancy of the rhombs, compared with the
mica, and their angles, seem to render this more than doubtful, the
angles being between 73^° and 78°. They are almost universally
accompanied by, and in contact with, the red or black rhombs,
and generally both.
Quartz cr3'stals, generally flattened, sometimes very minute,
sometimes large enough for the crystallization to be seen with the
naked eye ; generally masses of crystals, showing distinct crj- stal-
lization on the edges only, occasionally separate doubly terminated
prisms. Some of the specimens with polarized light are ver^^
beautiful.
A substance usually presenting the form of disks, jL inch and
less in diameter, showing, with polarized light, a radiation from
the centre, and a change of brilliant colors as the analyzer is
rotated. Apparently the same material occurs in acicular crystals,
often twinned at 60° and 120^, in a plumose form, and in a form
closely resembling a section of agate across the layers. Some of
these disks appear to be strictly a radiation of acicular crystals
from a centre, others to be made up of three or more oval masses ;
sometimes the latter are separate, or joined two, three, four, or
six together, showing apparent twinning at 60° and 120° ; these
oval masses, with polarized light, take each a single tint at a time.
This material was found also in mica from near Newtown Square,
1880.] NATURAL SCIENCES OF PHILADELPHIA. 2tt
Delaware Co., Pa., and from the Junction Railroad, above Girard
Avenue, Fairmount Park, associated with rhombs apparently of
lepidomelane or biotite, and also with quartz.
On the Bryn Maivr Gravel Mr. Henry Carvill Lewis re-
marked, that since the presentation of his paper on the " Surface
Geology of Philadelphia and vicinity," he had been able to extend
the investigation then begun, considerably beyond the limits of
Philadelphia. The " Upland Terrace " has now been traced con-
tinuously from near Trenton, through Bucks, Philadelphia, and
Delaware counties, to beyond Wilmington in Delaware. As far
as could be judged, the clay comes up to a uniform level along
this terrace. It has been gratifying to find that the main charac-
teristics of the different deposits, recorded in the paper referred to,
are constant throuohout the whole of this region.
The principal difficulty in the work has been want of topographi-
cal data. While within the limits of the city, the topographical
map of the Water Department had been of great service, but
beyond these limits elevations had to be estimated from occasional
railroad levels. Topography is an aid in all geological investiga-
tions, but in the study of surface geologj^ it is a necessity.
It is now desired to call attention to the great development of
the Bryn Mawr gravel in Delaware, and to the indications of its
assuming an important position in the geology of the Southern
States. In Bucks County, north of Philadelphia, the formation
has been recognized but scantily, but as we go south of the city
it increases largely in extent. Numerous hills in Delaware Co.
have been found to be capped by this formation, and in northern
Delaware it covers the gneissic hills in patches several miles long
and comes close to the river.
The Upland Terrace, after crossing the Delaware State line
about two and a-half miles back from the river, gradually ap-
proaches it, until near Bellevue Station, P. W. and B. R. R., its
base is but half a mile from the river. It forms the upper portion
of Wilmington, and then trends S. E. towards Baltimore, north of
the railroad and away from the river. In the neighborhood of
AVilmington the Bryn Mawr gravel lies directly upon and back of
the Upland Terrace, which is here about 300 feet high. It is
abundant to the southeast of Tallyville, Del., covering a large
tract of country, and it appears on the hills on both sides of the
Brandy wine in the neighborhood of Dupont's Powder Mills. It
is found on the Philadelphia and Wilmington Turnpike, two miles
northeast of Wilmington, and one mile from the river. In many
places it is five feet deep, and it seems less eroded than in Penn-
sylvania. It consists of sharp pieces of Mt. Holly conglomerate
and iron sandstone with well-rounded pebbles of quartzite and of
Potsdam sandstone, being identical with that of Chestnut Hill
and Br^n Mawr.
278 PROCEEDINGS OF THE ACADEMY OF [1880.
This forination, so abundant in Delaware, is thus proved to be
by no means a local one, and it is probable that it will be identified
with some of the formations grouped together under the name of
" Southern Drift."
The Bryu Mawr gravel has also recently been found in the
Montgomer}' County limestone valley, and there seems to be a
close connection between it and the surface or drift iron ores of
that valley. Some of these ores appear to be simply a verj' fer-
ruginous variety of the Mt. Holly conglomerate. They overlie
unconformably the steeply-dipping decomposed shales which hold
a more ancient and richer ore.
In Bucks Count}' there occurs a gravel different from any yet
described, M^hich at first occasioned some confusion. It has proved
to be the result of the decomposition of the lower Triassic con-
glomerate, the pebbles of which, loosened from their cementing
material, have been scattered tlirough the soil. These Triassic
pebbles are formed of gneiss, not Potsdam. Hills of red shale
border this gravel.
A preliminary map of the Surface Geology of Southeastern
Pennsylvania was exhibited, and it was suggested that its publica-
tion would be of service to man 3^ besides geologists.
April 28, 1879.
On some Enclosures in 3Iica. — Mr. Lewis exhibited some plates
of Muscovite which he had found on Shoemaker's Lane, German-
town, which contained microscopic ciystals of peculiar shape.
They consisted of a dark green mica, probably Lepidomelane,
in minute sharp crj^stals thickly- disposed throughout the
muscovite. These crystals were frequently arrow-shaped, and
generally much elongated. Large numbers of them were shaped
like a musket. They were very different from any of the enclo-
sures in the muscovite of Pennsbury, Del. Co., and were interesting-
objects under the microscope.
On Dendrites. — Mr. Henry Carvill Lewis made some obser-
vations upon dendrites and their mode of growth. He stated that
dendrites were not caused by filtration of metaliferous water, but
that they frequently grow upward by chemical or capillary action.
He described an exposure of white lower Triassic sandstone in a
quarry in the southern part of Noriistown, where dendrites of
oxide of manganese were seen upon the surface of the rock,
growing from below upwards. The dendrites were apparently in
process of growth, and were so soft that they could be scraped
with a knife from the rock. The material thus obtained gave a
bright metallic streak on the fingers, and was shown by the blow-
pipe to be hydrous oxide of manganese. It was otaserved that
while the rock above and below these dendrites was spotted with
minute rust-specks of manganese, the i)ortion upon which the
1880,] NATURAL SCIENCES OF rillLADELPHIA. 279
dendrites grew was pure white and free from such specks. It
seemed that the material of the dendrites is abstracted from the
rock and by some segregating force built up into tree-like forms.
An examination of their structures showed that the dendrites were
quite amorphous and that very frequently the upper extremities
of their branches were thicker than the stem portion, as though
some concretionary or capillary force acted most powerfully at
the growing points. No cr^'stalline structure was apparent, the
dendrites being bounded throughout by curved lines. It looked
as though they might have grown by a succession of concentric
metallic shells.
It was remarked that these dendrites were quite different from
those in muscovite and other crystals, which, frequently derived
from the substance of the crystal, have been so influenced by its
structure as to become often pseudomorphic. It was noted that
there are several distinct kinds of dendrites. They may be internal,
as in moss agate ; or external, as in the case now described. They
may also be either crystalline or amorphous. The crystalline
dendrites are subdivided into those which have been free to
crystallize of their own accord, and into those which have been
influenced b}' the crystalline structure of the mineral in which
they exist. Examples of each were cited.
On a Jurassic Sand. — Mr. Lewis directed attention to a fine
sand of considerable extent and depth, which he had found under-
lying the lower Cretaceous plastic clay. If this clay, as is supposed,
is the base of the Cretaceous formation, the sand below it may be
of Jurassic age. There is a fine exposure of this sand near Elkton,
Md. From its coherence it may be regarded as a fine-grained
sandstone. It is either white or pale yellow in color, and about
16 feet are hei'e exposed. Underneath the plastic cla^' south of
Trenton, N. J., the same sand is at least 30 feet deep. It is sug-
gested that, in the absence of fossils to fix its age, it may possibly
correspond stratigraphically with the " Hastings sand." The
overlying clay contains fossils at Baltimore, which Prof. Uhler
identifies as Wealden.
Upon the summit of the same hill, near Elkton, where the above-
described sand is exposed, " Bryn Mawr gravel" occurs in abun-
dance. It contains " Mt. Holly conglomerate," and has the same
features as in Delaware and Penn^lvania. Whether or not it has
any connection with the plastic claj^ is not known. This same
l)lastic clay, of probably Wealden age, occurs at Turkey- Hill, in
Bucks Countv, I'enna.
5
May 2G, 1879.
Potsdam Sandstone near King of Prussia. — Mr. Theodore D.
Rand called attention to primal (Potsdam) sandstone rocks in the
bed of a valley on the farm of Samuel Tyson, South Chester A'al-
ley Hill, near King of Prussia, Montgomery County, Pa.
280 PROCEEDINGS OF THE ACADEMY OF [1880.
A Kew Locality for Amethyst.— Mr. W. W. Jefferis announced
that Amethj'sts, well cr3'stallized, and of a rich purple color, had
been found this spring, for the first time, in the northern part of
Newlin Township, Chester Count}'. They were brought to the
surface by deep plowing, and were supposed to be derived from
a vein of this mineral.
September 22, 1879.
A New Corundum Locality. — Mr. W. W. Jefferis remarked
that a vein of blue Corundum, similar to that found in North
Carolina, was struck, on the south side of the Serpentine Kidge,
in Newlin Township, Chester County, a short time since. The
vein is well defined, being between walls of Culsageeite, in large
plates of a yellowish green color. Over 500 lbs. of massive blue
corundum has been taken out within ten feet of the surface.
The llinerals of Surry County, N. G. — Mr. H. C. Lewis commu-
nicated the following list of minerals which he had found near
Dobson, Surry Co., N. C, during a recent visit to that locality :—
Native sulphur, galena, pyrrhotite, p3n'ite,chalcopyrite, hematite,
menaccanite, magnetite, limonite, hausmannite, psilomelane, wad,
hornblende, actinolite, asbestos, garnet, talc, steatite, ripidolite,
chlorite.
The psilomelane occurred in a bed about 18 feet in thickness.
The magnetite was frequently polar. Native sulphur occurred
in cavities in quartzite as a coarse loose powder of rounded wax-
like grains, and was the result of the decomposition of pyrite.
It was also stated that rutile occurred in Alexander Co., N. C. —
a new locality.
Fossil (?) Casts in Sandstone. — Dr. J. M. Cardeza exhibited
specimens of quartz sandstone (Potsdam?) which he had found
lying loose upon the soil at Dutton's Mills, Pa., in which were
oblong rounded casts of sandstone, about an inch in length, and
similar to one another in shape. It was questioned whether they
mioht not be fossils.
*»
On a Peculiar Stratification in Gneiss. — Mr. Theodore D,
Rand stated that while much of the porphyritic gneiss of the belt
running southwest from the Falls of Schuylkill at the surface was
in rounded boulder-like masses, which had been mistaken for trap,
some of it presents at the surface a thin-bedded structure with,
apparently, very distinct stratification. Recently the cut of the
Pennsylvania Railroad through this belt, between Merion and Elm
Stations, about a mile from the boundary of the City of Philadel-
phia, has been widened, and on the south side ma}^ be seen an
interesting section. A mass of the gneiss, perhaps 15 feet across,
1880.] NATURAL SCIENCES OF PHILADELPHIA. 281
has ])een cut througli, and almost encircling it may he seen the
thin-bedded Aariet}', with its apparent stratification tangential to
the mass from which, by decomposition, it evidently was derived.
The true stratification of this bed of gneiss appears to be more
nearly horizontal and less contorted than that of any of the rocks
of the vicinit)- of Philadelpliia.
A Neio Locality for Lignite. — Mr. Henry Carvill Lewis
announced the discoveiy of lignite, or brown coal, in the lime-
stone valley of Montgomery Coanty, a mile and a-half from the
boundarj' of Philadelphia. He had found it, last June, at Marble
Hall, close to the marble quarry, within a few feet of diggings
for iron ore. In order to ascertain its extent and geological posi-
tion more definitely, he had caused a shaft to be sunk 40 feet deep
on the propertN' of Henry Hitner, Esq. After passing through 38
feet of decomposed hydromica slate, there was found a stratum 4
feet thick of a tough black fire-clay filled with fragments of lignite.
These fragments, sometimes a foot or more in length, lay in all
directions in the cla^^ They had the form of twigs and branches,
and, though completely turned into lignite, showed distinctly the
grain of the wood. The smaller pieces were generally flattened,
and offen as soft as charcoal, but the larger ones were quite hard
aud brittle and had the shining fracture of true coal. It burned
with a bright 3'ellow flame. Frequently l)alls of pyrite occurred
with the lignite.
The clay which contained it was underlaid by sand, and appeared
to dip south. It had an east and west strike, like that of the lime
stone and of the iron ores. In appearance it was similar to the
sub-Cretaceous plastic clays of New Jersey, which also contained
lignite I'esembling that of Marble Hall. White kaolin and white
and red potters' clay occur in the vicinity and are probably of
similar age. They are all older than the surface deposits and
gravel of the valley.
It was stated that while lignite is not uncommon in the Triassic
formation, its occurrence in a Silurian limestone valley is of great
interest. Whether referred to Tertiary or Jurassic age, it brings a
new geological epoch into this region and revolutionizes our ideas
of the age of man^' of the so-called *■' Primal " iron ores.
On Serpentine in Bucks County Mr. Lewis called attention
to the fact that while serpentine was abundant in Delaware Co., it
had not been recorded as occurring anywhere in Bucks Co. He
had recently noticed an exposure of it in that county, near the
village of Flushing, Bensalem Township. A narrow dyke of hard,
impure serpentine here crosses the road near the Neshaminy
Creek. He thought that the genesis of serpentine and its relation
to the gneissic rocks was still uncertain.
19
282 PROCEEDINGS OP THE ACADEMY OF [1880.
October 27, 1879.
THE IRON ORES AND LIGNITE OF THE MONTGOMERY CO. VALLEY,
BY HENRY CARVILL LEWIS.
The discovery ol lignite in the iron ore region north of Phila-
delphia introduces some new considerations in the study of its
geology, and has a direct bearing upon the age of its iron ores.
Lignite was found in this valley many years ago, but was supposed
to be Triassic, and therefore unimportant.' Before judging of the
connection that the occurrence of lignite in the Montgomery Co.
limestone valley will have with the geology of the Atlantic coast,
it will be important to enumerate other localities of a similar
nature where that mineral has been found.
In his Geology of Vermont, Prof. E. Hitchcock described an
occurrence of lignite in a similar position at Brandon, Yt., and
proposed a theory which excited much attention, but which has
been rejected by many geologists. It was shown that a steeply-
dipping stratum • of lignite lay within beds of plastic clay, kaolin
and iron ore, all dipping steepl}^ southeast. The iron ore deposit
was sometimes 100 feet deep, and all these beds rested against a
limestone which had the same steep dip. Mottled clays were
described as similar to those of Martha's Vineyard and the Isle of
Wight, and much of the formation was said to resemble a meta-
morphosed mica schist. The stratum of lignite was opened from
near the surface to a depth of 80 feet, and was used as coal. It
proved to be generally dicotyledonous, and to contain twigs and
fruits which belonged to a tropical climate, and which Professor
Xesquereux referred to a Tertiary epoch, probably Miocene. From
this discovery, Prof. Hitchcock proposed the theorj^ that all the
limonite iron ores of the Atlantic coast in similar geological posi-
tions were Tertiary and of oceanic origin. On the other hand,
it was argued that an isolated example was not sufficient to estab-
lish such a wide conclusion, and the lignite was regarded as locally
:formed by having been washed mto an existing cavern in the
limestone floor.
The next occurrence of lignite is a very similar one at Pond
Bank, near Chambersburg, Pa., described in an interesting
» F. Piof. Leidj, Proc. Acad. Nat Sci., Phila., 1861^ 77.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 283
paper by Prof. Lesle3\^ Here again it was found in a limestone
valley close to iron ore excavations. It was at a depth of 40 feet,
belo-sv strata of clay and sand. According to the superintendent
of the mine, it was in two strata, the lowest of which was 18
feet in thickness, and was separated from the upper bed, 4 feet thick,
b}^ a stratum of sand. Below it, at a depth of 65 feet, red and
white plastic clay occurred. The strata were nearly horizontal.
It was thought that the lignite was not necessarily connected with
the iron ores, but w^as a local deposit of late date, made in a shallow
pond, and that, as at Brandon, a sink-hole had been formed in tiie
underlying limestone. It was regarded as of the latest Tertiary
age.
Lignite has also recentlj^ been discovered by Prof Prime, in
Browm's iron mine, at Ironton, Lehigh Co., Pa.- He states that
it occurs in a Avhite plastic clay, but does not give the depth at
which it was fonud. He believes that it was transported by ice
and water in the Glacial epoch, and refers the iron oz'es of the
valley- to the same origin.
The writer believes that in the light of facts now developed,
this theory of the age of the lignite cannot be maintained. After
an inspection of the locality, he has found that the surface-drift
and boulders of that valley lie unconformably upon the forma-
tion containing the lignite. The lignite lies at a depth of 46
feet from the surface, in a tough plastic clay, which is entirely
free from boulders. About 30 feet of potters' clay and decom-
posed hydromica slate lie upon the lignitic stratum, and resting
upon the whole is 15 feet of drift. This surface drift, of yellow
brick-clay, boulders, gravel and drift iron ore, is thus of quite
different character from the strata below it, and is probably de-
posited b}^ glacial water's. The underlying formations have,
apparently, in some places, a dip like that of the adjacent lime-
stone, and are certainly more ancient than the surface drift.
The lignite recently found by the writer in the Montgomery Co.
valley, and described at the last meeting of the Section, occurs
under conditions very similar to those above indicated. In im-
mediate proximity both to a limestone outcrop and to iron ore
diggings, it was found at a depth of 35 feet, in a plastic cla}'-
which contains no gravel or boulders, and which is overlaid by
1 Proc. Amer. Phil. Soc, ix, 463.
^ Report DD., 2d Geolog. Survey of Pa., p. 76.
284
PROCEEDINGS OF THE ACADEMY OF
[1880.
clay and decomposed hydromica slate. A surface drift, containing
iron ore, gravel, and occasional boulders, lies unconformably upon
the whole formation. The section here presented was made in a
shaft which the writer was allowed to have sunk within a few feet
of Mr. Hitner's marble quarry, Marble Hall, Montgomery County.
FEET.
10
m
2^
LLLLU
" Top dirt," yellow, impure.
Soft white decomposed hydromica slate or im-
pure " kaolin," containing occasional broken
seams of sharp quartzite,'but no pebbles.
Coarse white sand and rounded pebbles ; appar-
ently a decomposed sandstone.
Tough mottled red claj'.
Blue plastic clay.
Lignite in a very tough, dark clay.
Coarse yellow sand, with fragments of stony
iron ore and with pebbles.
The lignite bed contains occasional streaks of fine gray sand,
and is underlaid by a coarser sand. So far as could be judged
from the very limited exposure, it dipped south, at an angle of
about 30° ; becoming thicker as it dipped. The lignite lies in
fragments in the clay, and consists of twigs and branches of land
plants, apparently all dicotyledonous. The lignite frequently
shows a brilliant black lustre when transversely fractured. The
small fragments are more like charcoal, and are often in the form
of flattened twigs. Some of these appear to be partially rounded
by attrition. No shells or marine fossils occur. Pyrite frequently
encrusts the lignite or forms nodules, and when exposed to the
air decomposes into ferrous sulphate.
At this same locality, lignite has been taken from three other
shafts in addition to the one just described. Two of these are
1880.] NATURAL SCIENCES OF PHlLADELrHIA. 285
close to the Marble Quariy, but the third is about 400 feet east of
these, behiud the barn of Mr. H. S. Hitner, who sa^-s that it Avas
found many feet below the surface. These facts indicate an east
and west strike to the lignitic strata, and an extent of 400 feet in
length and 40 feet in breadth. Shafts 100 feet north of these
struck iron ore, but no lignite. They exhibited the following
succession of strata: — (1) "Top dirt;" (2) Decomposed hydro-
mica slate; (3) White clay; (4) Yellow sand ; (5) Iron ore.
Recent explorations have shown that Marble Hall is not the
only locality where lignite is found, but that it occurs in a number
ol other places in the same valley. About a mile and three-
quarters west of Marble Hall it was formerly found in a field on
the Eidge Road, opposite a house once owned by W. Potts. It
was at a depth of about 40 feet. Red and white potters' clay,
white kaolin and iron ore are found close to the opening. The
lignite at this place is hard, and is said to burn well. Another
locality is on the farm of W. Wills, south of Plymouth Creek,
about one mile from Conshohocken. Considerable quantities of
lignite have been here exhumed, the pieces being often a foot in
length. This shaft was opened about thirt}'^ years ago, and was
probably the locality referred to by Dr. Leid}'. Lignite has also
been found in a number of iron ore pits south of here and east of
Conshohocken. It is said always to occur in fire-clay.
The above localities are all included in a strip of country about two
and a-half miles long and a quarter of a mile broad, Ij'ing in nearly an
eist and west direction. The lignite appears to form two distinct,
na. row lines of outcrop with a definite E. and W. or E. N. E. and
W. S. W. strike, — thus conforming with the limestone. While its
dip has not been actually established, the decomposed slates and
sandstones of apparently similar age have been observed to dip
40° S. 20° W.
From these facts it would appear that the lignite is not a mere
local wash or accidental deposit, but thatf it is part of a stratified
and distinct formation, having a trend like that of the limestone,
and of considerable extent ; and when the strata in the Mont-
gomery Count}" Valley are compared with those in other parts of
the countr}^, it will be seen that we have here to deal with a forma-
tion which, closely connected with the limonite iron ores of the
great limestone valleys, and having remarkably similar characters
throughout, may prove an important feature in American geology.
In entering upon a consideration of the age of the lignite, it will
286 PROCEEDINGS OF THE ACADEMY OP [1880.
be desirable brieflj^ to sketch the geolog}', and especially the
surfoce geolog}' of the valley in which it occurs.
The underlying rock is an altered lower Silurian limestone, the
"Auroral " of Rogers, which in the southern part of the valley is
crystalline marble and in the northern part is a sand}^ magnesian
limestone. It has an E. and W. strike and a steep south dip, and
is supposed to have an inverted synclinal structure. The lime-
stone rises to the surface in a series of parallel ridges, and between
these lie the iron ores and the lignitic strata. Bounding the valley
are hills of altered shale of probably Potsdam age. Beyond this
line of hills to the north, are the Triassic red shales and sand-
stones, while to the south is the Philadelphia gneissic district.
In many places the North Valley Hill has been eroded away and
Triassic strata lie directly upon the limestone.
The iron ores of this region probably belong to four different
geological ages, and may therefore be divided into four classes.
1. Gneissic Ore. This ore, never found in the valle3^, occurs in
the gneissic rocks of Chester County north of the Chester Vallej^,
and has been formed in place from the altered gneiss. It dips
with the gneiss, and is generally accompanied by scales of graphite.
Prof. Rogers^ supposed that this ore belonged to isolated patches
of Triassic red sandstone. The writer, however, has not been able
to confirm his sections, nor to show the presence of any more
recent formation than the aneiss.
2. Primal Ore. The hydromica slates which lie between the
Potsdam sandstone and the limestone liberate, when decomposed,
a rich limonite ore which is largely mined in jjortions of the
valley. Although in very irregular beds, a steep dip can be re-
cognized. It is perhaps derived from the decomposition of pyrite.
This is pi-obably the ore mined at Edge Hill.
3. Tertiary Ore. This ore, associated with which are the de-
posits of lignite, plastic clay, kaolin, fire-sand, etc., has been
hitherto confounded either with the Primal ore or Avith the Drift
ore of the valle^'. In that part of the valley under discussion
there are three distinct lines of outcrop of this ore, having nearly
an E. and W. trend. A ridge of limestone separates two of these
lines. The ore lies, sometimes at a great depth, below a re-stratified
decomposed h3dromica slate. This latter formation is almost
identical in appearance with the decomposed Primal slate in place
1 Geol. of Penna., I, 87.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 287
at the edges of the valle}', and has therefore been mistaken for it.
The discovery of lignite below it proves its re-stratification in a
later age. In manj- places shafts have been sunk over 100 feet
without coming to the limestone. The ore, originally derived
either from the limestone or from the primal slates, appears to lie
below the lignitic strata,
4. Drift Ore. Resting often unconformabl}^ upon these last, and
capping the elevations thi-oughout the valley, is a drift deposit of
gravel and boulders containing a workable iron ore. The compo-
sition of this drift is most interesting. Its boulders, almost with-
out exception, are composed of a loose-grained Potsdam sand-
stone,— a formation not now existing either on the North or South
Valley Hill at this place, and found only in a limited exposure at
the eastern end of the valley. The ScoUthiis linearis is frequently
found in these boulders. Moreover, notwithstanding the large
extent of Triassic red shale and sandstone immediatel}' to the
north of the valle}', and the occurrence of that rock resting often
directly upon the limestone, not a trace nor a fragment of Triassic
rocks have been found in this drift.
The evidence is here strong that this drift has not been caused
by any flood from the north in a modern age. Additional evidence
bearing upon the same point is found in the fact that the Triassic
region north of here is absolutely free from drift of any kind. A
careful study by the writer of much of that region has shown that
not a single drifted pebble is there found. The soil is formed
from the rocks below it, and such elaj^s as occur are bog clays of
local origin and recent age. That the pebbles of the valley drift
have not been derived by weathering from the neighboring lower
Triassic conglomerate, which holds often large pebbles, is shown
by the fact that such pebbles are here formed entirely of gneiss
or gneissic quartzite, and never of Potsdam, and therefore are
quite different from those in the valle3^
The drift ore and gravel does not lie in hollows, as though
locally washed, but is found in patches upon the elevated portions
of the valley, as though it were the remnant of a once continuous
deposit.
The facts above enumerated suggest a possible origin at an age
when cliffs of Potsdam sandstone, since eroded awaj-, stood as a
high barrier between the limestone valley and the Triassic rocks
north of it. Such a barrier would effectually prevent Triassic
fragments from mixing with the drift of the valle}', and would,
288 PROCEEDINGS 'of THE ACADEMY OF [1880.
during its degradation, offer the material for the pebbles and
boulders of that drift. In Triassic times some such barrier may
have formed the southern shore of the Triassic waters. It has
been interesting to discover that most of the pebbles belonging to
the sub-Cretaceous plastic clays of the Delaware are formed of
Potsdam sandstone, and that therefore during lower Cretaceous
times also, some such mountain of Potsdam must have offered
itself to eroding agencies. Again, it is found that Tertiary* gravels,
both in Pennsyh'ania and New Jersey, contain an abundance of
Potsdam pebbles. The hypothesis that the materials for the sub-
Cretaceous plastic clays and the Tertiary gravels were furnished by
hills now sunk beneath the Atlantic Ocean ^ is not sustained by
what is known of the configuration of the sea-bottom. The theory
now offered is supported by numerous facts concerning the power
of erosion, which geological considerations in other fields have
presented.
In a former paper on "The Surface Geology of Philadelphia and
vicinity," the writer showed that, in addition to the clays, four
separate gravels of different ages can be distinguished in that
region. These are (1) ''The River Gravel," the newest of all the
gravels; (2) "The Philadelphia Eed Gravel," of Champlain age;
(3 ) '' The Fossiliferous Gravel," recently proved by the writer to
be of upper Tertiary, perhaps Pliocene age, and now called the
" Glassboro Gravel ; " (4) " The Br^ai Mawr Gravel," the oldest
of the gravels, also oceanic, aud conjectured to be of upper
.^Miocene agfi. This last gravel, and this only, agrees in its char-
s.'Sicters with the valley drift now under consideration. In the
«;absence of .all Triassic fragments, in the presence of Potsdam
'boulders, and in the amount of erosion, these two gravels are iden-
:tieal, and it saems probable that the " Drift Iron Ore " of the one
is only a very ferruginous variety of the "Mt, Holly Conglom-
ero^te " of the olher. This being the case, we have here a formation
wh.icii, notwithstanding its boulders, suggestive of floating ice, ap-
pears ,to be older than an oceanic Pliocene gravel. There is per-
haps no good reason wh}'^ a glacier might not have existed in
upper Tertiary tim«s, boulders formed by which ma^^ still be
.fouit(i. However tikis may be, it appears that there are strong
grourjds for assiguiiag an upper Tertiary age to the drift ore and
:gra,vel «of the Montgoiiaer^^ County Valley.
Retui'Aing, finall}', to the lignite and associated strata, shown to
•^ .Report,ffla Clay Deposits of N. J., 1878, pp. 20.31.
1880.] NATURAL SCIEXCES OF PHILADELPHIA. 289
be older than the formation just described, and shown by its own
characters to bear no trace of glacial agencies, we may conjecture,
without au}^ reference to the phmts of the lignite, a middle or lower
Tertiary age. From the steep dip of the beds, — a fact difficult to ex-
plain,— and from the great resemblance of the plastic clays to those in
New Jersey, on the Delaware, the writer at first supposed them to
be of Wealden age. Some facts in connection with a gravel found
in Virginia and other Southern States, which, in both appearance
and position is ver}' similar to the Bryn Mawr gravel, were at first
thought to indicate a Jurassic age. But after a comparison with
the other lignite localities, and especiall}^ with that at Brandon,
where the fossils were shown to be of Tertiary age, this view can
hardly be sustained. The absence of shells or marine plants indi-
cates a period of inland waters, and the plants at Brandon belong
to a tropical climate.
It is now suggested that the period of the lignite mny corre-
spond most closely with that called b}' European geologists the
Oligocene. Since, in the present state of our knowledge, it is obvi-
ously' unsafe to make the age of these lignite deposits contempora-
neous with any exact geological epoch, and as there is a possibility
of their belonging to some period not recognized elsewhere, it will
probably be wiser for the present to group them together under
the name of The Brandon Period. As more facts develop and
wider comparisons can be made, more certain conclusions will be
possible ; aiid it must be understood that the theories here proposed
are brought forward onlj^ as those which now appear best to ex-
plain the facts observed.
Postscript. — Since the presentation of the above paper, the writer
has been in correspondence with Prof. N. A. Bibikov, of Augusta,
Georgia, who has recently discovered lignite in that vicinity. The
locality, called "Read's Brown Coal Mine,'' is in Richmond County,
two and a-half miles from Berzelia, and sixteen miles from Au-
gusta. It is described as h'ing back of the outcrops of gneiss and
limestone, and is apparently in a very similar geological position
to the Pennsylvania locality. Iron ore, plastic clay, kaolin, and
decomposed sandstone occur with the lignite. As in Pennsylvania,
the lignite was found in a plastic clay beneath 25 feet of a decom-
posed sandstone. Pour strata of lignite, separated by layers of
shale and clay, were found at a depth of from 30 to 45 feet from
the surface. A series of coarse and fine sands and clays under-
laid these deposits and were penetrated to a depth of 95 feet.
290
PROCEEDINGS OF THE ACADEMY OF
|_1880.
Three different shafts were sunk, the extremes being 600 feet apart,
in all of which lignite was found. The shaft in which the following
section was made is about 150 feet from an outcrop of hornstone
and quartzite, and 300 feet from a creek which lies 200 feet
below it.
24
1
3
2
1
3
6
1
8
1
2
8
11
+
rM
©;-=3):
Mottled clay.
Decomposed sandstone.
Crust of iron sandstone and spathic iron.
Decomposed sandstone
with thin layers of clay.
Nodules of pyrite.
Lignite.
Shale and clay.
Lignite.
Shale and clay.
Lignite.
Shale and clay.
Lignite.
Light-colored shale
with fossil plants.
Dark-colored (bituminous) shale.
White clay with streaks
of rose-color, etc.
Sandy clay.
Ferruginous coarse sand with nodules of clay iron-stone,
4 inches to 2^ feet in diameter with ochre, clay, etc.,
and quartz i^ebbles sometimes 3 inches in diameter.
Light-colored shale.
Yellowish sand.
Sandy clay.
White, very fine micaceous sand
with clay.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 291
The second stratum of lignite is the best, and contains fragments
of lignite sometimes three feet long. A number of fossil plants have
been found in this and other laj-ers. Some specimens were im-
bedded in a Isiyev of brown sandstone. The fossils appear to be
fragments of trees, grasses and other land plants, none of Avhich,
however, were sufficiently perfect to be determined. No shells
were found.
The whole section at Berzelia is rem.irkabl}- similar to those at
Brandon, Chambersburg, Ironton and Marble Hall, and with them
indicates the existence of a great inland fresh water Tertiar}' for-
mation in Eastern America, during the Brandon Period, once
fifty miles broad and nearly a thousand miles long.
292 PROCEEDINGS OF THE ACADEMY OP [1880.
All Enclosure in Quartz. — Mr. H. C. Lewis exhibited a crystal of
quartz from Herkimer County, N. Y., in wliich, hanging from a
bubble which moved in a cavity containing liquid, was a tuft of
minute acicular crystals of a pure white color. A microscopical
examination had failed to identify them with any known substance.
The crystals were similar to those of many organic salts. It was
conjectured that they had crystallized out from the liquid. Under
a power of 75 they looked like tufts of white wool, and it was
suggested that if future investigation failed to refer them to a
known mineral species, it might be convenient to give them the
name Erilite (from £V>:ov, wool).
In other cavities in the same crystal thei-e was an amorphous
yellow^ish-brown w^axy substance of unknown composition.
Menaccanite and Talc from Maryland — Mr. Wm. W. Jefferis
remarked that in Harford Count}^, Md., near the village of Dublin,
there is a vein of green foliated Talc in the serpentine, which has
been opened about 6 feet in length. It has furnished cleavage
foliated specimens over a foot in extent. The same vein contains
Menaccanite in tabular crystals, Avell crystallized. Yellow beryl
has also been found there, showing all three in the same specimen.
Sunstone in Lahradorite. — Mr. Jefferis stated that on examin-
ing a specimen of Labradorite in his possession, from the coast of
Labrador, he found that in addition to the usual play of colors
(blue and green), by turning it in another direction it showed
innumerable crystals of giithite, making it a beautiful sunstone,
which, he believed, was an unusual thing, and w4iich he had not
found mentioned in the books.
On a Probable Pseudomorphism of Gummite and Uranotile after
Uraninile. — Dr. A. E. Foote remarked that among a number of
specimens of gummite and uranotile, that he had recently' received
from Mitchell Co., N. C, he noticed some which were of remark-
ably regular form. The edges were slightl}' rounded, but they
were apparently simple prisms belonging to the triclinic system.
On breaking these open he found a solid core of uraninite, sur-
rounded by a layer of gummite, and this, in turn, surrounded b^^
a layer of uranotile. Although crystals of uraninite have never
been obsei-ved, he ventured to suggest that this is plainly a case
of pseudomorphism after uraninite. He hoped hereafter to obtain
crystals wdiose angles can be accurately measured.
He had observed nt least twenty specimens having evidentl}- the
same crystalline form, and all plainly pseudomorphs after some
pre-existing crystal. The majority of those that were broken open
showed the alteration of uraninite into gummite, and of gummite
into uranotile ; though in a few the uraninite had been changed,
and the crystal showed simply gummite and uranotile.
1880.J
NATURAL SCIENCES OF PHILADELPHIA.
293
November 24. 1879.
on a new fucoidal plant from the trias.
by henry carvill lewis,
The fossil described here as Palseophycas limaciformis^ sp. nov.,
is from a very beautiful and well-defined specimen, casts of which
stand out in relief upon a slab of Triassic sandstone. It was
found by the writer near Milford, New Jersey,' in upper Triassic
strata. Its general faeies is like that of some species of Palseo-
phycus and Astropthycus of the Carbonifei'ons age, and of Butho-
trephis of the Clinton group, and it belongs to the same order of
plants — that of the fucoid or marine algae. The general, rather
than generic name of Palaeophycus, which is quite as indefinite as
that of Fucoides, is well applicable to it.
The frond is cjdindrical and jointed. The ramuli, or short
l)ranches of the frond are flesh}-, tubular, elongated bodies of about
an inch in length. They are spindle-shaped, attenuate, and more
or less curved at both ends.
The}' are very frequentl}^ ag-
gregated in bunches of three
or more, radiating from a com-
mon point of growth, and are
generall}' detached from the
main frond. The form of
these bodies is characteristic
of the plant, being distinctl}'
snail-shaped; hence its spe-
cific name, " limacifomnis.-^
At the locality where it
was found there also occur
specimens of ripple-marked
sandstone, also of rain-prints
and mud-cracks. These show
the phj-sical conditions under
which the plant grew — that
of mudflats in shallow water
frequentl}' left exposed to
sun and rain. The rain-prints
have been compared with
Nat. Size.
PalcEophycus limaciformis Lewis, sp. nov.
294 PROCEEDINGS OP THE ACADEMY OF [1880.
modern rainprints made in mud, and their great similarity noticed.
One specimen of a fucoid found liere has apparently been so con-
fused and distorted by the beating of a heavy rain-storm that its
structure can scarcely be recognized.
Fucoids of somewhat similar apjjearance have been described
from more ancient geological horizons, but not, so far as can be
learned, from the Trias. The occurrence of a plant which has the
characters of marine algae in a formation supposed to have been
deposited by fresh water is interesting.
Postscript. — Since the above paper was presented, a photograph
of this fossil has been sent to Mr. Leo Lesquereux, the distin-
guished paleobotanist, who, in a letter to the writer, says : " Your
plate represents what I consider a new species of Palseo2:>hycus,
whose analogy is with P. (species undetermined) Hall, Paleont.
of N. Y., vol. i, PI. 70 (Hudson River group); also distantly
related to Fucoides auriformis and especially F. heterophyllus,
same author, 1. c, ii, PI. .3, figs. 3, 4. You may also find a kind
of analogy to what Hall considers and figures as roots or also as
stems of some marine plants, same vol., PI. 8, figs. 4, 5, and pi. 9,
fig. 4, and also pi. 10, figs. 5, 7 (all Clinton plants). The type is
evidently old, rather Devonian, even upper Silurian. European
authors have nothing like this from the Trias, Jurassic, Cretaceous
or Eocene." •
1880.] NATURAL SCIENCES OP PHILADELPHIA. 295
TJie Northern Belt of Serpentine in Radnor Township. — Mr.
Ranp made the followinaj communication.
Sometime ago, in a communication to the Academy, (Proc. Ac.
N. S., 1878, 402) I described a belt of serpentine in the valley of
the Gulf Creek, Radnor Township, Delaware Co., Pa. Recently a
trench for water pipe on the property of Judge Hare has enabled
me to procure specimens illustrating a section across the bed and
on both sides of it, which are presented herewith. The section is
on a line nearl}^ N. 75° W. The belt is probablj' not far from N.
70°-73° E., or about two-thirds that of the section, but this is not
certain. While deductions from these few specimens would not
be safe, 3"et the strong resemblance between the decomposed
gneiss of the easternmost exposure and the clearly magnesian
rocks 40 feet distant, points more to an alteration in place than to
a distinct bed.
Garnet mistaken for Corundum. — Dr. J. M. Cardeza called
attention to a garnet rock at Chelsea, Delaware Co., Pa., which is
quarried and used as corundum.
296 PROCEEDINGS OF THE ACADEMY OF [1880.
THE TRENTON GRAVEL AND ITS RELATION TO THE ANTIQUITY OF MAN.
BY HENRY GARVIIiL LEWIS.
In the course of an investigation of tlie Surface Geology of
Southeastern Pennsylvania, some facts have been developed in
connection with one of the gravels, which,, bearing directly upon
the Antiquity of Man in America, become of great interest..
Among the many scientific problems now attracting attention^
none perhaps holds a more prominent position than that of the
Antiquity of Man. It is a subject which, notwithstanding the
numerous facts gathered and the bulk of literature published, must
be regarded as still in an undecided condition.
As the Delaware is in many respects a tj'^pical river, and as
therefore deductions made here will hold good for the valle3^s of
many other rivers of the Atlantic coast, it is thought that a record
of the investigation will be of more than mere local interest. The
subject will be approached from a purely geological standpoint.
The main difficult}^ in inquiries of this kind has been the absence
of exact geological data. Hasty conclusions have been drawn
from an inspection of relics found in a gravel, which a more
accurate knowledge of the age of that gravel would not have
sustained.
The writer has shown in former papers ' that the gravels of the.
Delaware Valley belong to several distinct ages ; and if therefore
at any place the remains of man are shown to occur, it will be all
important to know to which of these gravels they should be
referred.
The surface formations of Southeastern Pennsylvania may be
divided into five clays and four gravels. The following is believed
to be the succession in which the^- occur, beginning at the
oldest: (1) Jurasso-Cretaceous plastic clay; (2) Tertiary clays,
(" Brandon Period ") ; (3) Bryn Mawr gravel, (upper Tertiary); (4)
Branchtown clay; (5) Glassboro gravel, (Pliocene); (6) Phila-
delphia red gravel, (Champlain) ; (7) Philadelpiiia brick-clay,
(Champlain) ; (8) Trenton gravel, ("Eskimo period"; ; (9) Recent
alluvium. Of clays, the oldest is the Jurasso-Cretaceous plastic
clay exposed at Turkey Hill, Bucks Co. A similar plastic clay,
1 "The Surface Geology of Phila. and vicinity." Proc. Min. and Geol.
Section, Acad. Nat. Sc. Phila., Nov. 1878.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 291
which, however, may be of later age, has been passed through by
artesian wells in the southern part of Philadelphia. The next
oldest clay appears to be the potters' clay of the Moutgomer}- Co.
limestone valley, which, containing sometimes lignite, and overlaid
by kaolin, decomposed hydromica slate, etc., belongs with its asso-
ciated limonite ores, to an inland Tertiary formation, the " Bran-
don Period,'' possibly of Oligocene age.^ A third clay, the
" Branchtown claj^" found at high elevations in a few places in
the gneissic region, containing occasional boulders, was made at a
period of general submergence and appears to be of a late Tertiary
age. The " Philadelphia brick-cla3' " of more recent formation,
of large extent, and with numerous boulders, is confined to the
river valley. This clay, deposited at the close of the Glacial period
by the waters resulting from the melting of the great Northern
Glacier, rests against the rocky " upland terrace " at a height of
about 150 feet above the present river. The fifth and newest
clay is the recent bog claj^ or mud in the flood-plain of the
river, still in process of formation.
The gravels are distinguished from one another both by their
composition and b}' their relative hj'psometrical positions. The
" Bryn Mawr gravel" — the oldest gravel of consequence in this
region — is readil^^ distinguished from others by the peculiar mate-
rials composing it, and is also known b}' being found at high
elevations (400 feet), in often isolated patches, capping the gneissic
hills. It is characterized by absence of fossiliferous or Triassic
pebbles and by the presence of an iron conglomerate, and is of
oceanic origin, and probably upper Tertiary age.^ A similar gravel
occurs on the heights of Georgetown, D. C. The next oldest
gravel, also oceanic, and which here occurs at lower elevations
than the last, the writer called in a former paper " The Fossilifer-
ous Gravel." It frequently contains pebbles formed of Niagara
limestone and other fossiliferous rocks, and has been found abun-
dantly' in New Jersey as well as in Pennsylvania. It is well
exposed in the railroad cut at Ridley Park, Del. Co. It is the
yellow gravel Avhich caps the watershed between the Atlantic
and the Delaware at a height of nearly 200 feet, and is now named
for distinction " The Glassboro gravel." Its pebbles are frequently
^ F. "The Iron Ores and Lignite of the Montgomery Co. Valley," by
the writer. Oct., 1879.
= 7. "On the Bryn Mawr Gravel," by the writer, Mar., 1879.
20
^98 PROCEEDINGS OF THE ACADEMY OF [1880.
weather-worn and eaten hy age, and have thus a much more ancient
appearance than the smooth, fresh-looking pebbles of later gravels.
It contains no boulders of consequence and is believed to be of
Pliocene age.
L^-ing at a lower level, within the Grlassboro gravel, and formed
of a mixture of its pebbles with others brought down the Dela-
ware valle}^, is a third gravel — the " Philadelphia red gravel."
This, like its overlying bi'ick-cla}', is confined to the river valley.
It is distinctly stratified ; it contains numerous fragments of
Triassic red shale and of gneiss, and smooth boulders of Silurian
rocks ; it shows flow and plunge structure and wave action on a
large scale ; and like the older gravels, it rests upon a decomposed
gneiss, which is sometimes interstratified with its lower laj^ers.
There are numerous exposures near the Universitj^ of Pennsjd-
vania. The writer has identified it on the Potomac and other
rivers, and it appears to belong to the age of the melting glacier —
the Champlain epoch.
The last and newest of all the gravels is one which, at Philadel-
phia, seemed to be of little importance. It lies close along
the river, and rising a few feet above it, extends but a short dis-
tance back from the river bank. It covers the flat ground of
Camden and the lower part of Philadelphia, and forms islands in
the river. It was called The River gravel and sand. It is this
alluvial gravel, the latest, except the recent mud-flats, of all the
surface formations, which is the subject of the present paper, and
which, from its great development farther up the river, is now
named The Trenton Gravel. It is in this gravel, and in this
gravel only, that traces of man have been found.
The Trenton Gravel at Philadelphia is composed principally of
a sharp micaceous sand, which, when below water-level, becomes
a "quicksand." Gravel lies below the sand. Unlike all the other
gravels, it contains but few pebbles of white quartz, and is of a
dark gray color. Its pebbles are made exclusively of the rocks
forming the upper valley of the river. Their shape is also very
characteristic. The pebbles of the older gravels are oval or egg-
shaped, but these are for the most part flat. This flat shape is
characteristic of all true river gravels. At several places along
the Delaware, gold has been obtained from this gravel. The
absence of clay in anj^ of its laj-ers indicates the action of swiftly-
running water. Data obtained from artesian wells have shown
1880."] NATURAL SCIENCES OF PHILADELPHIA. 299
that this formation has a depth on Delaware Avenue, Philadelphia,
of about 50 feet, and that it extends up to about Third and Market
Streets. On Smith's Island and on the bar in the river opposite
Cooper's Point, it is 100 feet deep, lying upon rock. It therefore
underlies the river, filling up its ancient channel. On Richmond
Street some verj- large boulders are seen lying upon the sand.
On tracing the Trenton gravel up the river, it is found to be
confined to its immediate vicinity, and that, from Philadelphia to
the Neshaminy Creek, its boundary is generally between the linO:
of the Pennsylvania Railroad and the Delaware. From this point
the bounding terrace trends directlv towards MorrisviUe and
away from the present river. Thus, at Bristol, the gravel and its
overlying sand extends two miles back from the river, and is.
bounded b^' a well-marked hill, upon which lie the older gravel
and brick-clay of Champlain age. These and the Tertiary gravels,
extend nearl}' seven miles inland. At Tullytown the Trenton
gravel extends two and a-half miles back, and at the canal shows
the following succession of strata : (1) sandy loam, 1 foot ; (2) fine
gray "moulding-sand," 2| feet; (3) shai-p "bar sand," 1 foot; (i)
clean gray river gravel of unknown depth. In other openings
near here the gravel is so full of boulders that these are dug in
large quantities and sent to Philadelphia for " cobble-stones."
Near Wheatsheaf Station, close to the railroad,. an opening which
has exposed a section of the Trenton gravel nearly half a mile in
lenoth. exhibits well the general features of the formation. The
pebbles, of characteristic shape and color, are made of gray Triassie
argillite, slate, red shale, sandstone, conglomerate, and various:,
other rocks found farther up the valley, while large and often
sharp boulders of red shale and other materials frequently occur.
The whole formation has a A-ery fresh appearance when compared
with older gravels. Xear Turkey Hill a large smooth boulder,
five feet in diameter, lies upon the sand.
At MorrisviUe the narrower portion of the valley begins, and
from here up, the river flows on arockA* bottom, and the gravel is
shallow and is confined to the immediate vicinity of the river.
The older gravels of oceanic origin continue across New Jersey
and do not appear above Yardleyville. The Philadelphia red
gravel is no more seen, but the brick claj' with its boulders occa-
sionally appears part way up the steep hills enclosing the valley,,
and is abundant in the side valleys formed by tributary streams.
300 PROCEEDINGS OF THE ACADEMY OF [1880.
Above Yardleyville, therefore, we have to deal with but two sur-
face formations, — the boulder-bearing brick clay, often much
eroded, and the Trenton gravel, confined to the bottom of the
valley and showing but little erosion. It will be well to bear in
mind the distinction between these two formations, — the one of
glacial, the other of post-glacial age. The writer has traced them
as far up as the Water Gap, past the great terminal moraine into
-laciated regions. It is interesting to note that while the modified
moraine material close to the river at Belvidere is in some points
similar to the Trenton gravel, and is the source of part of that for-
mation , the moraine on the Lehigh River at Stemton and at other
inland localities contains pebbles and boulders very similar to
those of the Philadelphia brick-clay.
Throughout the whole course of the Trenton gravel it is observed
that it lies within a channel previously excavated down to the
rock through the boulder-bearing brick claj- and its red gravel,
which, as shown in a former paper, belong to the Champlain epoch.
The Trenton gravel is therefore, later than the Glacial and Cham-
plain epochs ; and this is a fact which, when considered in connec-
tion with the human relics found in this gravel and the consequent
antiquit}^ of man, it will be most important to remember.
Having now sketched the character and position of the Trenton
gravel along the Delaware valley, we are prepared to examine the
formation as exposed at the locality whose name we have chosen
to distinguish it.
Trenton is in a position where naturally the largest amount of a
river gravel would be deposited, and where its best exposures
would be exhibited. It is at the point where a long, narrow valley
with precipitous banks and continuous downward slope, opens out
into a wide alluvial plain at a lower level. It is here that the
rocky floor of the river suddenly descends to ocean level and even
sinks below it, forming the limit of tidewater. Thus any drift
material which the flooded river swept down its channel would
here, upon meeting tidewater, be in great part deposited. Large
boulders which had been rolled down the inclined floor of the
upper valley would here stop in their course, and all be heaped up
with the coarser gravel by the more slowly flowing water except
sn<3h few as cakes of floating ice could carr^^ oceanward. On the
other hand the finer gravel and sand would be deposited farther
down the river.
1880.J NATURAL SCIENCES 01' PUlLAtlfiLi'fllA. SOl
This is precisely what occurs at Trenton. The material, Which
at Philadelphia is generally- fine, grows coarser as the river is
ascended, until at Trenton we find often immense boulders im-
bedded at all angles in the gravel. Moreover, the river has here
cut entirely through the gravel down to the rock, exposing at
one place a cliff of gravel 50 feet high. At Philadelphia, on the
other hand, as we have seen, the river still flows on the top of the
gravel. This fact may also be accounted for. Having heaped up a
mass of detritus in the old river channel as an obstruction
at the mouth of the gorge, the river, so soon as its volume dimin-
ished, would immediatel}' begin wearing away a new channel for
itself down to ocean level. This would be readil}^ accomplished
through the loose material, and would be stopped only when
rock was reached. On the other hand, that gravel wliich had
been deposited at places farther down the river where its bottom
was below ocean level, would remain un-eroded or nearly so. Wheii
the river had attained the level of the ocean there would be no
occasion to cut a deep channel, and it would therefore flow on top
of the gravel which it had deposited. It is necessary that this
point should be understood, as other geologists have brouglit for-
ward various theories to explain the high bank of gravel at Tren-
ton. The fact of the river having cut through the gravel at
Trenton, while at Philadelphia it flows upon it, is due to the con-
figuration of the rock floor of the river, which at Trenton rises
above ocean level, and at Philadelphia lies nearl}^ 100 feet below it.
In addition to the exposure upon the river bank, where the
whole depth of the formation is seen, the long railroad cuts
made by the Pennsjdvania R. R. at Trenton, afford excellent
sections of the gravel. It exhibits the distinctive characteristics
of a true river deposit, and is very different from the gravels which
are found at higher levels. It contains no clay;. its pebbles are
made of the rocks of the river bed and are flattened, and the strati-
fication of the whole deposit is well seen in the alternations of
sand and gravel. It extends several miles back from the present
river, covering the low ground along the Assunpink Creek, and
indicating the existence here of a former bay or arm of the Dela-
ware. This bay was shaped somewhat like a horseshoe, which had
one extremity in Trenton at the hill above the canal, and which
washed the base of the hill north of the Assunpink Creek, and,
extending about three miles back from Trenton, and sweeping
302 PROCEEDINGS OF THE ACADEMY OF [1880.
aronncT the " Bear Swamp," had its other extremity near the
house of Di\ C. C. Abbott, below Chambei'sburg. This village was
under water. Another ba}^ extended up the valley of Crosswicks
Creek. Boulders of Champlaiii age lie upon the Tertiary" gravels
which form the ancient banlv.
From the extent of the Trenton gravel in this vicinity, state-
ments have been published that it covered the whole southern part
of the state, and that at the time of its deposition the Delaware
Rivel' emptied into the ocean at Trenton. It is evident tliat the
distinction between the very different gravels of this region has
not been perceived. Careful examination will show the great dis-
similai'it}^ between the Trenton gravel and such gravels as occur
at Princeton Junction and interior New Jersej^, which are in
great part of Pliocene age, and will prove that it is confined to
the ancient river bed.
The presence of very large boulders on the river bank at Tren-
ton has led some geologists to suppose that the formation was a
glacial moraine. The occasional though ver}' rare examples of
scratched pebbles and polished boulders, which the flood had evi-
dently carried down from the moraine material north of Belvidere.
have been brought forward as supporting this theory. Yet the
absence of till and of angular masses of rock, and generally of
materials foreign to the Delaware Valley, when regarded in con-
nection with what we have shown to be the general characters of
the formation, can not be explained upon this theorj^ The char-
acter of the river banks along the valley render the presence of a
glacier at Trenton extremely improbable. These show no marks
of glacial action. We have, moreover, already shown that
the Trenton gravel is more recent than the deposits of Cham-
plain age, and that, lying in a channel cut within them, it is
the most recent of all the gravels. Clearly the Delaware Valley
and the channel of the river were excavated in a time previous to
the deposition of the Trenton gravel. The channel subsequently
having been filled up by this gravel, the diminished river still
later has cut a new channel either completel.y through it, as at
Trenton, or partially, as at Philadelphia. It is probable that
slight undulations of the level of the coast have aided in producing
these changes.
Before describing the human relics found in the Trenton gravel,
there are several facts bearing upon its origin and age which it
will be well to consider.
1880.] NATURAfi SCIENCES OF PHILADELPHIA. 30o
It has been noticed that from Trenton to Philadelphia the creeks
flowing into the river Delaware have a steep south bank, while the
ground north of the creek is flat. The writer finds that the flat
ground north of the creek is made of Trenton gravel, while the
southern bank is made of older formations which have been for-
merl}^ cut awaj^ by water action coming from the north. Thus,
the steep south bank of the Neshaminy is made of " Philadelphia
red gravel " of Champlain age, while a flat plain of Trenton gravel
lies south of the creek. The same configuration of the banks of
creeks on the New Jersey shore has been noticed by Prof. Cook.
By assuming that the river at the time of the deposition of this
gravel was of larger volume than now, this fact is of ready explana-
tion. The southern bank of the creek, often of Cretaceous or
Tertiarj" strata, in each case formed the shore of the ancient river,
and was worn away into a steep bank by the flood from the north.
Similar in cause and effect are the present banks of the Delaware,
which are steep on the outside of each curve of the river, and flat
and covered with recent alluvium on the inside.
Another fact showing river action is the frequent occurrence of
exposures of " flow and plunge structure " in this gravel. In these
the la3^ers are seen to dip up stream, as would be expected by
downward flowing water. It is interesting to find, on the other
hand, that the same structure in the Tertiary gravels, both of
Pennsylvania and New Jersey, shows layers dipping southeast, as
though deposited by incoming oceanic tides.
Another instance of the fluviatile character of the Trenton gravel
is found in the peculiar topography which it sometimes exhibits.
Frequently, instead of forming a flat plain, it forms higher ground
close to the present river channel than it does near its ancient
bank. Moreover, not only does the ground .thus slope downward
on retreating from the river, but the boulders become smaller and
less abundant. Both of these facts are in accordance wdth the
laws of river deposits. In a time of flood the rapidly flowing-
water in the main channel, bearing detritus, is checked by the more
quiet waters at the side of the river, and is forced to deposit its
gravel and boulders as a kind of bank.
In determining the comparative age of the Trenton gravel, a guide
may be found in the amount of its erosion. In this respect a
marked contrast exists between this and more ancient gravels.
Unlike the land covered bj' older surface formations, that covered
o04 PROCEEDINGS OF THE ACADEMY OF 1880.]
by the Trenton gravel is remarkably level and free from hillocks
or ravines. The change in topography may be well seen in the
neighborhood of Trenton, and can be noticed almost anywhere
along the valley. This fact alone would indicate a more recent
age than that of the clays and gravels of the Champlain epoch.
This difference is much more marked when comparison is made
Avith the oceanic gravels.
The actual time necessary for the Delaware to cut down to the
rock through 50 feet of this gravel at Trenton is by no means
great. Numerous facts have been adduced by geological writers
and by engineers to show how rapidly a stream of water can wear
through loose gravel material. When it is noted that the gravel
cliff at Trenton has been made, not b}" a straight downward cut,
but by a side wearing away as at a Ijank, and when it is remem-
liered that the erosive power of the Delaware was formerly very
much greater than it is now, it will be conceded that the presence
of the clitf at Trenton will not necessarily infer its high antiquity.
From what is known of the action of running water upon gravel,
it is thought that the time necessary to produce the erosion now
observed might be reckoned b}^ hundreds rather than by thou-
sands of 3'ears. While the gravel was of course formed in a
previous time, the rapid action of the flood which deposited it,
shown in many places by the character of the gravel, indicates
that the time necessary for its deposition need not have been long.
Having now shown that the Trenton gravel is a true river
deposit of modern age, it will be of interest to inquire how such a
flood as we have proved to exist could have originated. No flood
within the historical epoch has been known to at all approach in
magnitude that which deposited the Trenton gravel. No boulders
of the size found in and upon that gravel are ever carried down
the river by recent ice-cakes. In fact, at Trenton and below, the
boulders of this gravel are often much larger than any in the
Champlain gravel of that part of the valley.
We have seen that at the time of the Trenton gravel flood, the
lower part of Philadelphia, the whole of Bristol and Tullytown, and
almost all of Trenton were submerged. That the climate was then
cold is indicated not only by the suggestion that there were then prob-
ably very large masses of bouldei'-bearing ice floating in the river,
but also by the fact that, as the writer is informed by Dr. C. C.
Abbott, bones of Arctic animals (walrus, reindeer, mastodon).
1880.] NATURAL SCIENCES OP PHILADELPHIA. 305
often rounded by attrition, have been found in this gravel.
Although the Trenton gravel has none of the features of a moraine,
it is true that the cliff at the base of Riverview Cemetery, holding
immense boulders, has the appearance of having been deposited
b}' glacial waters. At other places, the boulders resting upon the
sand overlying the gravel suggests the grounding of large ice-
cakes derived from some mass of ice large enough to be called a
glacier.
It is difficult to imagine an origin for such a flood as we have
described other than the melting of a glacier. We have shown
that the flood was not an inroad from the sea, but that it
came down the valle}'. No rain-storms of modern experience
could have supplied such an amount of water. To call the time
of this flood a '■ Pluvial Epoch," will be of little assistance, since
no origin for such extraordinary rains is suggested, except under
a very different climate, or by evaporation from a melting glacier.
Yet such a glacier cannot be the great glacier of the Glacial epoch.
That was the glacier which in its melting deposited the brick-clay
and red gravel which Tve have shown to be much older than the
Trenton gravel. It must have been, if a glacier at all, another
and more recent one whose melting caused the flood which formed
this gravel. This last glacial flood flowed in a channel excavated
through the deposits of the first glacial period.
It appears, then, that there is evidence of a Second Glacial
Period — a period in which was deposited the last of the gravels,
and which has but lately passed away. From the limited extent
of its deposits it is inferred that the second glacier was much
smaller than the fti-st, and that its southern extremity was con-
fined to the valley. A second glacial period is recognized in
Europe under the name of the Reindeer Period.
It is thought that the hypothesis of a second and more local
glacier, long subsequent in age to the first great glacier, will
explain all the facts observed. The Trenton gravel cannot be
assigned to the first glacial period except by assuming that there
have been no river gravels deposited since that time ; — an assump-
tion which can hardly be maintained. Some European archje-
ologists have held that the Palseolithic Era, the era of the river
gravels, is antecedent to the Reindeer Period, the period of the
cave-men. No such distinction has been observed on the Delaware.
Should future researches show that a separate and second glacial
306 PROCEEDINGS OF THE ACADEMY OF [1880.
epocli cannot be proved in America, the facts here observed will
indicate a much more recent date for the disappearance of the
great glacier than has been assigned to it. The period of the
Trenton gravel flood, whether contemporaneous with a glacier or
not, is the period of the last geological deposits here known- the
recent mud-flats being alone excepted.
We have now glanced at the characters of the Trenton gravel,
and have indicated, so far as the facts at hand allow, its position,
origin, and relative age.
It is in this gravel that the writer's friend, Dr. Charles
C. Abbott, of Trenton, has made the interesting discovery of stone
implements of human workmanship, which, in their shape and
characters, are quite unlike those of the Red Indians of the Atlantic
coast.^ He has found them imbedded at various depths in the ap-
parently undisturbed gravel of the cliff at Riverview Cemetery
and in other places near Trenton. They are of palseolithic type,
and differ from Indian stone implements by being larger, ruder,
and made from a different material. They are composed of gray
argillite, a rock which is found in place farther up the river,
and which is a Triassic shale altered and hardened bj- the heat
from adjacent trap dykes. They occur in positions which render
it extremely probable that they belong to the same age as that of
the deposition of the gravel, or at least to an age when it was
overflowed b}' the flooded river. There are two points which
offer strong evidence in that direction.
The first is the fact that modern Indian implements, "neoliths,"
are never found associated with these " palaioliths " in the gravel.
Although abundant on the surface, it is stated that thej^ never
occur at a depth of more than a few inches in undisturbed soil,
while the palteoliths are found often ten or more feet from the sur-
face. This fact alone argues a different age for the two classes of
implements.
The second fact is that when found below the surface of the
ground, these palffioliths always occur in the Trenton gravel and
never in older gravels. The writer, in company with Dr. Abbott,
has gone over much of the ground where the implements occurred ;
and it was very interesting to find that it was onl^^ within the
limits of the Trenton gravel, previously traced out by the writer,
^ V. Tenth and Eleventh Annual Keports of the Peabody Museum of
American Archaeology.
1880.J NATURAL SCIENCES OF PHILADELPHIA. 30t
that Dr. Abbott had found implements below the surface. Beyond
the terrace of older gravels the pala^oliths sometimes occur with
implements of the modern tj'pe, but are not imbedded at any
depth. In Pennsylvania, moreover, the writer has found similar
pahieoliths in the region covered by the Trenton gravel and in that
region onh'. Here, then, is the strongest probability, even if the
implements were found upon the surface only, that they belonged
to and were of coeval deposition with the river gravel.
The implements of argillite found at the lowest depth in undis-
turbed gravel have been generally decided by archa-ologists to be
of human origin. It is, however, true that there are many sharp
fragments of this rock in the Trenton gravel which are of natural
origin, and that pebbles and partially rounded fragments of the
same rock are frequent. The writer has found several fragments
of argillite in the gravel exposed at the cut near Wheatsheaf
Station, Bucks Co., Pa., which, whether they were artificial or
natural, it was impossible to determine.
All the evidence that has been gathered points to the conclu-
sion that at the time of the Trenton gravel flood, man in a rude
state lived upon the banks of the ancient Delaware. He may have
been in the habit of spearing fish and seals with spears pointed by
his rough stone implements, and these having been dropped into
the flood may have sunk into the loose and shifting gravel. The
weathering upon the implements is so slight as to afford no evi-
dence of their high antiquity. Many of the paljeoliths found in
the river gravels of Europe, are of very similar tjqie. As a rule,
probably the implements of the Trenton gravel are somewhat more
rude. The writer is informed that even more primitive forms are
now in constant use among some of our Western Indian tribes.
It is interesting to find, as pointed out by archaeologists, that
until lately the Eskimos have used stone implements quite as rude
and similar in appearance to those found in the Trenton and other
river gravels, and it has been suggested that that race, now living
in a climate and under conditions perhaps similar to those once
existing on the Delaware, may have some kinship with the pre-
Indian people of this river. It ma}' be that an Eskimo race, living
here at the time of the flooded Delaware, were driven north by
the coming of the Red Indians. If future archaeological work
shows this surmise to be correct, the writer suggests that the
period of the Trenton gravel and of this palaeolithic people, — a period
308 PROCEEDINGS OF THE ACADEMY OF [1880.
perhaps following a second glacial age, — might appropriately be
called The Eskimo Period. This name, derived fi-om a higher order
of beings than that which gave the name Beindeer Period., is much
more suggestive and is probabl}' of full3^ as wide application as
the latter name, A term already in use, the Palaeolithic Era, is
also convenient.
It has been held that the occurrence of palaeoliths at Trenton
offered evidence of a very high antiquity of man in America, and,
the gravel being considered as a glacial moraine, that man's exist-
ence was carried back to interglacial and even pregiacial times. '
As we have seen, the geological investigations along the Delaware
Yallej", described in this paper, throw quite a new light upon this
subject. They show that the implement-bearing gravel is of post-
glacial age, and is a river deposit of comparativeh^ recent forma-
tion ; and that neither in the gravels of the Champlain epoch nor in
de])Osits of any previous age have any traces of man been
discovered. The evidence appears to indicate the origin of
man at a time which, geologically considered at least, is recent.
The actual age of the Trenton gravel, and the consequent date to
which the antiquity of man on the Delaware should be assigned, is
a question which geological data alone are insufficient to solve.
The only clue, and that a most unsatisfactory one, is afforded bv
calculations based upon the amount of erosion. This, like all geo-
logical considerations, is relative rather than absolute. The same
reasoning that showed that the modern river channel might have
lieen excavated in hundreds rather than thousands of years, will
indicate that no great length of time is necessary' to produce all
the surface features of the Trenton gravel. While the writer may
venture to express the opinion that there is no reason geologically
for carrying the age of this gravel and the antiquity of man on the
Delaware farther back than a very few thousand years at the most,
lie is fully aware that any close approximation can safely be
arx'ived at only by extended comparison with other river gravels and
by a much more complete series of observations than have yet
been possible. Ethnological considerations, which make paljco-
lithic man to antedate the oldest races of the mound-builders, will
Jiave a bearing upon this question. Meteorologists ma^^ show that
1 It will be remembered that Sir Charles Lyell, in his Principles of Geol-
ogy, 11th Ed., vol. 1, p. 286, conjectures the period of the gi'eat glacier to
have been about 200,000 years ago.
1880.J NATURAL SCIENCES OF PHILADELPHIA. 309
a cold climate and a period of a flood far larger than any of late ex-
perience may require a long lapse of time. These considerations
are not within the scope of this paper. It has been the aim of the
writer to define the antiquit}' of man in relation to geological rather
than to historical events. If, in showing that the Eskimo period is
the last of the geological ages, it does not necessarily follow that it
is by anj' means recent, it must be remembered, on the other hand,
that its high antiquity is not proven by the facts thus far
observed.
The conclusions to which the facts seem to point may briefly be
summarized as follows : —
1. That the Trenton gravel, the only gravel in which implements
occur, is a true river deposit of post-glacial age, and the most
recent of all the gravels of the Delaware valley.
2. That the palteoliths found in it really belong to and are a
part of the gravel, and that they indicate the existence of man in
a rude state at a time when the flooded river flowed on top of this
gravel.
3. That the data obtained do not necessaril}^ prove, geologically
considered, an extreme antiquit}- of man in Eastern America.
310 PROCEEDINGS OF THE ACADEMY OF [1880.
Note on Philadelphite — a new mineral. — Mr Lewis gave a pre-
limiiiaiy description of a new verniiculite from near Wayne
Station on the Germantown Railroad, wliicli he proposed to call
''■ Philadelphite." It occurs in plates of a brown color and talcose
lustre, existing as seams in an altered hornblende rock. When
heated, it exfoliates with great force to man}' times its original size
and becomes of a coppery bronze color. It was stated that while
exfoliating, it was able to lift over 50,000 times its own weight. It
had a hygroscopic power nearl}^ as great as that of chloride of
calcium. Its optical characters and its chemical composition were
given.
Analysis of Philadelphite. — Mr. Reuben Haines contributed
the following analyses of Philadelphite.
Specific gravity (determined in alcohol of 95 p. c.) 2.78-2.96.
I. XL
SiO^ 39.06 38.52
re.,03 20.59 20.01
AI2O3 14.75 14.82
Fe,0 2.04 2.04
CaO .99 1.08
MgO 11.49 11.32
MnO (traces) .... ....
Li20 (traces) .... ....
K,0 6.89 6.61
Na,.0 .90 .64
H,0 4.27 4.27
F (traces) .... ....
100.98 99.31
Per cent, of hygroscopic water in I, 3.12 p. c. ; in II, 3.43 p. c.
In these analyses the mineral was dried t.t 100° C, the hygro-
scopic water not being included in the determinations. Owing to
its very hygroscopic nature, it was found very diflicult to obtain
its weight at 100° C. accurately. It gains rapidly in weight while
being weighed upon the balance. Examples of its hygroscopic
power were given. The analyses were made l)y dissolving the
mineral in concentrated hydrochloric acid. Iron was estimated
volumetrically and the alkalies hj Smith's method effusion.
1880.J NATURAL SCIENCES OF PHILADELPHIA. 311
December 22, 1879.
THE SO-CALLED EMERY-ORE FEOM CHELSEA, BETHEL TOWNSHIP,
DELAWARE COUNTY, PENNSYLVANIA.
BY F. A. GENTH, JR.
At the November meeting of this Section, Dr. Cardeza called
the attention of the members to a garnet rock, mined as emery-
ore, at Chelsea, Bethel Township, Delaware Co., Pa., and subse-
»iuently left it with me for analysis.
The rock is composed almost exclusively of rounded rhombic-
ilodecahedral grains of red garnet, varjdng in size from a fraction
of a millimetre to over one centimetre ; also a little quartz, biotite,
muscovite, and magnetite. It is very friable, being easily crushed.
Its fracture is uneven, excepting in some of the larger grains,
which are so much intersected by mica, that, when struck by a
hammer, they break into angular fragments, apparently showing a
crystalline cleavage. Specific gravity = 4.028.
An analysis of the smaller and purer grains, obtained by wash-
ing and picking out, gave :
SiO, = 41.11
Fe.p3 = 2.11
Al.Os = 21.60
FeO ^ 25.86
MnO = 2.22
CaO = 1.89
MgO = 5.41
100.20
which proves it to be an ordinary iron-alumina garnet.
312 PROCEEDINGS OF THE ACADEMY OF [1880.
Some Neiv Mineral Localities. — Mr. Joseph Willcox announced
the following new mineral localities :
Burgess, Ontario, Canada, on the north shore of Rideau Lake :
Phlogopite, Green Pj-roxene, Apatite, Zircon. North Elnisley,
near Otty Lake, Canada : Phlogopite, in large and perfect crys-
tals. Bedford, Trontenac Co., Out., Can. : Apatite (unusually
fine). Black Pyroxene, Scapolite. Near Westport, Ontario, Can. :
Black Tourmaline. Russel, St. Lawrence Co., N. Y. : Steatite
pseudomorphous after Tremolite and Scapolite ; Black Tour-
maline, with modified terminations. Macon Co., N. C. : Crystals
of Biotite in Muscovite.
All the above were found in fine specimens, well crj^stallized.
Specimens were exhibited to the Section.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 313
ON PHILADELPHITE (Sp. Nov.).
BY HENRY CARYILL LEAYIS.
The mineral to which the above title has been applied was found
by the writer four years ago, in what was then a quarry of horn-
blendic gneiss, close to the boundary of the Twentj^-second Ward,
Philadelphia. The locality is on Germantown Avenue, at the
bridge crossing of the Germantown and Norristown Railroad,
near Waj-ne Station. The quarry is now walled u^d, and is used
as a coal and lime yard.
Geologically, the locality is just at the base of the terrace of
metamorphic rocivs which bounds the drift formations underlying
the greater part of the city. Quaternarj^ clays, boulders of the
Champlain period, and tertiary gravels appear within a hundred
feet of the quarry, and the waters of those different epochs have
successively eroded the hill rising above it. This hill, here called
Neglejf's or Logan's Hill, about 225 feet in height, is part of the
same hill or " Upland Terrace,'' which, trending nearl}' northeast
and southwest, has been traced continuously from here into Mary-
land, on the one side, and across New Jersey on the other, and
Avhich, though composed of quite different rocks in different places,
forms throughout, the boundary of the post-jurassic formations.'
The rock at this place is a hard black hornblendic gneiss, subject
to decomposition in its upper portions. It is well exposed in the
cut on "Waj-ne Street, where numerous minerals occur, and it is
the same which is quarried at Frankford and at MoKinney's
quarry, both noted mineral localities. In its altered state it
crumbles easil}-, and when heated exfoliates. In this condition,
after being crushed in a mill between heavj^ iron rollers, it is
sometimes used as a building sand.
The mineral here described as PhiladeJphife belongs to the ver-
miculite group of hj'drous silicates. It occurs both disseminated
in scales throughout the gangue-rock, and also in seams, an inch or
more in thickness and many feet long. Associated with it in the
same quarry are crystals of sphene, epidote and hornblende, and
specks of chalcopyrite. It has been found in small quantities also
at Waj'ue Street, at McKinney's quarry, and in Germantown.
^V. Proc. Miu. and Geolog. Section Acad. Nat. Sci., Phila., Nov., 1878.
21
314 PROCEEDINGS OF THE ACADEMY OF [1880.
Since most of the vermiciilites occur in serpentinous or chloritic
formations, it is to be noted that no such rocks occur here or in
the vicinit}'. The mineral is probablj' derived orio-iually from
hornblende.
Physical Characters, — Hardness, 1..3 ; Specific gravit}', 2.80
(taken in alcohol and referred to water). Lustre pearly. Color,
by reflected light, bronze ; by transmitted light, brownish red, and
in A'er}^ thin laminae, brownish yellow. Opaque, except in thin
pieces. Streak brownish yellow. Laminae unelastic, readily
flexible, tough, not brittle. Feel greasy.
CrystaUographic Characters. — Monoelinic. Cleavage ; basal, emi-
nent ; also, occasional!}', a cleavage parallel to the diagonals.
Striations crossing at about 90°, caiising the mineral to break
into nearly rectangvilar fragments, are sometimes observed, and
these are parallel to the plane of the optic axes and to the diagonals
of the rhomb. No triangular striations as in Jefferisite. Plates
often contorted and wrinkled. Twin crystals frequent, observable
by polarized light. Optically biaxial. Double refraction strong,
negative. Optic-axial angle, 31°20'-39°30' ; generally 3T°±:.
Crj-stals sometimes nearly 2 inches wide and \ inch high. The
hyperbolas are well defined in the polariscope, and the angle of
their divergence is more constant than in some of the other ver-
miculites. Twinning produces variations in the angle.^
Pyrognostic Characters. — In the closed tube it gives off water and
exfoliates with great force, in a direction perpendicular to its base,
to ten times its original volume. Upon exfoliation it becomes of a
bright copper color and takes a metallic lustre. It also becomes
brittle and more opaque. The exfoliated mineral has a far more
distinct and frequent secondary vertical cleavage than it has before
exfoliation, and the basal cleavage is also easier. It shows strong
double refraction in the polariscope, and has an optical divergence
of about the same amount as that of the unignited mineral (30° to
37°). The h^'perbolas are extremely ill-defined, and no exact
measurements could be taken. It is yellow b}- transmitted light.
It forms a fine object under the microscope by reflected light.
The fine copper color gained on exfoliation is characteristic, dis-
tinguishing it from the other vermiculites. The color is obtained
whether it is heated suddenly in the flame, or slowly in an air-bath
to exfoliation. Upon long-continued ignition in a platinum cru-
1 T'. Prof. Cooke's Paper on the Vermiculites, Proc. Amer. Acad.,
Boston, 1874, 35.
ISSO.] NATURAL SCIENCES OF PHILADELPHIA. 315
cible, heated without access of air, it becomes a steel-gray color, its
iron having been reduced. Before the blowpipe it gives the violet
flame of potash and fuses to a black magnetic globule, which does
not intumesce when further heated.
With the fluxes it reacts for silica and iron. It is readily dis-
solved by hot sulphuric acid, the pure white silica being left in the
original shape of the mica. It is dissolved in hydrochloric acid
upon long digestion .
Chemical Composition. — In the investigation of the chemical
composition of Philadelphite the writer has had the valuable
advice of his friend, Prof. F. A. Genth, of the University of
Pennsylvania. The method used in the estimation of vanadium
is entirely due to him. The writer is also indebted to his friend,
Mr, Reuben Haines, of Germantown, for two analyses, and for
some interesting experiments.
Of the four analyses given below, Numbers I and II are by Mr.
Haines; Nos. Ill and IV by the writer. Nos. I and II were
made upon the pulverized mineral, previously dried in an air-bath
at 100° C; the hygroscopic water, amounting to over 3 p. c, not
being included in the determinations. " In both the anal3^ses the
sample was dissolved in concentrated HCl, and the Si02 purified
by digestion with HCl. The Fe and Al were precipitated togetlier
by NH4HO and the Fe titrated by permanganate. The ferrous
oxide was found by dissolving the weighed mineral in sulphuric
acid in a closed flask from which the air was expelled by boiling
with sodic carbonate, and titrating as before. The magnesia was
weighed as pyrophosphate and the alkalies were separated by
Smith's method of fusion, and were determined by platinic chloride,
controlling the result by ignition of the platinic salt in hydrogen
and weighing as metallic platinum. Tlie combined H,0 is an
average of the results of experiments Nos. IV and VI (given
below) taken at a red heat on bottom of crucible."
Analyses Xos. Ill and IV were made upon the ignited mineral,
this being considered its most constant state. The atomic water
was determined separately, and the analysis of tlie anhydrous
mineral reduced when the percentage of water was added. The
ignited mineral being v\ith difficulty soluble in acid, it was decom-
posed by fusion with sodic carbonate for anal^'sis. After repeated
evaporation of the silica with HCl, it was found still to contain
titanic acid, which was extracted by evaporation with concen-
trated H,iS04 and precipitated by dilution and boiling. Addi-
316 PROCEEDINGS OF THE ACADEMY OF [1880.
tional titanic acid was separated upon boiling the filtrate from SiO,„
after reduction with H-S. In one analysis titanic acid was sepa-
rated from SiOoby volatilizing the latter with HF, dissolving the
residue in HsSOi, diluting and boiling. Ferrous oxide Avas deter-
mined in the air-dried mineral as in analyses I and II. Iron and
alumina were estimated by precipitation b}' boiling with sodic
acetate in a neutral solution, dissolving in HCl, reprecipitating
with NH4HO, igniting and weighing together. In the filtrate
MnO was precipitated by bromine and ignited.
The following method was employed for the detection of vana-
dium. 80 grammes of impure mineral were mixed with 90 grammes
of sodic carbonate and 100 grammes of sulphur, and the whole
heated slowly in a Hessian crucible covered by charcoal until
partially fused. It was then digested in warm water, filtered, and
to the filtrate dilute IICl was added, precipitating a copious
heavy flocculent brown mass of the sulphides of vanadium, copper,
cobalt and nickel. The precipitate was washed, ignited and evap-
orated with nitric acid, when it gave a red residue. This was fused
with a mixture of sodic carbonate and sodic nitrate, and extracted
with water in order to separate the oxides of copper, cobalt and
nickel. Solid ammonic chloride was now added to the aqueous
solution, when vanadate of ammonia was precipitated. Upon
ignition it was changed to vanadic oxide, and was found to be
pure, giving all the characteristic reactions.
For the estimation of vanadium the following method was em-
ployed. 4^ grammes of the pulverized ignited mica were fused
with a mixture of 3 parts NaCOj and 1 part NaNOs, the mass ex-
tracted with H.O, filtered, and the filtrate digested with HiS.
Traces of CuS and FeS were filtered off, and the silica eliminated
by evaporation to dryness and addition of dilute H2SO4. II2S was
again added, giving a blue solution. After driving off" the H,,S by
heat, the vanadic acid present was estimated volumetrically by the
addition of a measured portion of a standard solution of per-
manganate of potash.
Magnesia was determined as pyrophosphate, and the alkalies b}'
means of Smith's method. Phosphoric acid was precipitated as
phosplioraolybdate of ammonia, and weiglied as pyrophosphate of
magnesia.
On account of the remarkable h3'groscopic powers of Philadel-
phite, great difficult}^ was experienced in the estimation of the com-
bined water. Nearly one-half of the water in the air-dried mineral
1880.]
NATURAL SCIENCES OF PHILADELPHIA.
sn
is h^'groscopic, and may be driven off eitlier by long exposure over
sulphuric acid in a desiccator, or by drying in an air-bath at 100°
C. The percentage of water given in the analj'ses represents
approximately the amount of water in the mineral after such
desiccation.
Spec. grav. (taken in alcohol of 95 p. c.) 2.78-2.96.
SiO,
Al,,03
Fe.,03
Fe^O
MnO
MgO
CaO
Na,0
Li.b
K2O
F
H2O
H^-gr
Spec.
2.80.
SiO,,
TiO,,
Ai,d,
Fe^Og
V2O3
FeO
MnO
NiO|
CoO)
CuO
MgO
CaO
Xa,0
Li.,0
K.2O
PO,
CI
H,S04
H,0
I.
39.06
14.75
20.59
2.04
trace
11.49
.99
.90
trace
6.89
trace
4.27
II.
38.52
14.82
20.01
2.04
trace
11.32
1.08
.64
trace
6.61
trace
4.27
Mean.
38.79
14.78
20.30
2.04
11.40
1.03
.77
6.75
4.27
Quantivalent ratio.
2.587
.861
.761
.056
.570
.037
.025
.143
.474
2.587
1.622
5.45
3.42
.831
1.75
.474 1.—
100.98 99.31 100.13
oscopic water in I, 3 12 ; in II, 3.43.
grav. (taken in alcohol of 84 p. c on the air-dried mineral)
III.
35.94
1.30
15.23
19.48
.37
2.09
.46
trace
trace
11.41
1.38
1.42
trace
6.52
trace
trace
trace
4.34
lY.
35.52
.77
16.32
19.43
.36
2.28
.55
.06
.08
11.72
1.54
.38
ti'ace
7.11
.11
trace
trace
4.34
Mean.
35.73
1.03
15.77
19.46
.37
2.18
.50
.06
.08
11.56
1.46
.90
6.81
.11
4.34
2.43
5.05
Quantivalent ratio.
2.38
.05
.91 )
.73 V 1.65 3.43
.01 )
.06 ^
.01
5
3
.58
.05
.03
.14
.87
1.80
.48
.48 1 1
99.94 100.63 100.45
Hygroscopic water in III and IV, 3.24.
318 PROCEEDINGS OF THE ACADEMY OF [1880.
From both these pair of analyses we have the ratio
R: R : Si : H = 2 : 3 : 5 : 1 and R R : Si : H= 1 : 1 : i. The ratio
of bases to silica is 1:1, and for sesquioxides to protoxides.
vr II
S : R = 2 : 1.
Philadelphite dried at 100 ^C appears to be a unisilieate, the
water not being basic.
The formula ma}' perhaps be written
£.: \\ O. \\ r 2. (K ^^-^"l -L 3. Q f S2 ^^ "(I,, J- 2 „.,
•^- II ^-i II L 0 \^>i' -i^J ^ o P K^^- 1 ^~ )j^ ^ 5 '^4
The general symbol would be,
11 VI
R^jiVi, O20. Sio- 2 HjO.
The water will be regarded as water of cr3'stallization. Prof.
Cooke has shown the close chemical relation between the anhydrous
vermiculites and biotite. A like result is brought out by the fol-
lowing analysis of ignited Philadelphite. The analysis is a mean
of the two analyses of the anhydrous mineral which formed
analyses Nos. Ill and lY of the mineral dried at 100° C.
2
Quant, ratio
SiO,
TiO,
37.35
1.08
2.49|
.05)
2.54
2.82
ALO3 .
16.49
.96 ^
re,03
20.33
.76
1.73
. 1.92
V203
.38
.01)
FeO
2.28
.06 V
MnO
.52
.01 1
MgO
CaO
12.09
1.53
.60 (
.05^
.90
1.
Na,0
.94
.03 1
K.O
7.13
.15^
100.12
II VI
Here R : ?^ : Si ^ 1 : 2 : 3, the ratio of a typical biotite. Anal-
gous as the anhydrous mineral is to biotite in its formula, it has
been shown that physically and optically the two minerals are
quite dissimilar, and it is not proven that they have any necessary
connection. It is by no means a hydrous biotite in the sense that
margarodite is a hydrous muscovite, in which case the characters,
optical and phj'sical, are identical. Such hydrous biotites occur
in several places in the vicinity of Philadelphia, in a partially
1880.] NATURAL SCIENCES OF PHILADELPfflA. 319
altered micaceous gneiss, in which the muscovite has become
margarodite, and the orthoclase become white and crumbling.
Such mica exfoliates slightly when heated, is uniaxial, fusible
with difficult}^, and might be called Hijdr-ohiotite for convenience.
It frequentl}' occurs enclosed in cr^'stals of margarodite, or in
muscovite passing into margarodite.
Hygroscopic Properties. — In the determination of water in its
different states in PhiladeliDhite, the principal difiicult}^ was on
account of the strong hygroscopic properties possessed by the
mineral. After the water has been expelled by heat or desiccation,
it is rapidly absorbed again from the air, if exposed. Upon the
balance, the dried mineral gains so rapidly that it was found
necessary while weighing to enclose it in corked tubes. It appears
to absorb water with the avidit}^ of chloride of calcium. Even
when enclosed in watch-glasses clasped together and standing in
the closed balance-case with dry CaCL, it gains decidedly in weight.
The following experiments by Mr. Haines illustrate its h3^gro-
scopic properties :
G-rammes.
(1) Weight of undried mica, .9935
Heated at 100° C. for li hours, .9616
Weight after standing; in balance-case with CaCL
for 3 days, .9915
Reheated for 3 hours at 100° C, .9580
Left on balance 20 minutes. Gain in weight, .0070
Left on balance 2 hours. Total gain in weight, .0085
(2) Weight of undried mica, 1.1280
Heated at 100° C. for 3 hours, 1.0965
Left in balance-case with CaCl, for 1 hour, 1.1175
Left in balance-case with CaCI.^ for \\ hours, 1.1230
Left in balance-case with CaClj for 2^ hours, 1.1250
Left in balance-case with CaCl,^ for 2 days, 1.1260
(3) Undried mica heated at 100° C. for 6^ hours.
Loss, 2.49 p. c.
On standing in balance-case with CaCL for 2^
daj'S, regained nearly the whole of its original
weight (all but 2 milligrammes). Again
heated at 100° for 3 hours, loss of weight, 3.09 p. c.
These experiments, showing that nearly the total amount of
hygroscopic water is regained even in the presence of such an
320 PROCEEDINGS OF THE ACADEMY OF [1880.
active desiccator as chloride of calcium, indicate a remarkable
hygroscopic force in the dried substance ; a property not easy to
explain. It will be noticed that this force is exercised much more
powerfully immediately after desiccation than it is after a lapse of
time. Experiment No. (2) shows that two-thirds of the water is
absorbed during the first hour. It has been found that the amount
of water in the powdered mineral varies with the hygrometric state
of the atmosphere at the time of weighing. It is interesting to
note that several of the zeolites, a class of hydrous silicates whose
exfoliation by heat is very like that of the vermiculites, also have
strong hygroscopic powers, losing and regaining part of their
water with ease.^
Water of Crystallization — The water in Philadelphite probably
exists in three theoretical conditions, viz. : — H^-groscopic water,
water of crystallization and water of constitution. The first is
driven off by drying at 100° C. or by exposure to dry air over
HaSO^; the second by gentle ignition, and is accompanied by ex-
foliation ; the third by strong and prolonged ignition. The latter,
which probably does not much exceed 1 per cent., and whicli
the analyses have shown is not needed with the basic radicals to
complete the unisilicate formula, will be regarded with the water
of crystallization. The most satisfactory determinations of the
water of crystallization have been made by subtracting the hygro-
scopic water from the total water.
The following experiments haA'e been made upon the amount
and condition of the water.
(1). The dry mica, which had been out of the quarry for more
than a year, was cut into pieces about 5 mm. square, heated in a
platinum crucible to a bright red heat for 25 minutes, cooled in a
desiccator over HsSOi for half an hour, and then quickly weighed.
It lost 7.58 per cent., which will be regarded as the total amount
of water.
(2). The finely powdered mica holds more water. Different
experiments gave:— 7.81 (ignited 10 minutes), 7.89, 7.90, 8.11
(ignited 25 minutes), 7.50 (powdered just previous to ignition).
Strong ignition of the powdered mica probably volatilizes some of
the alkalies in addition to the water.
>• Damour (Ann. d. Mines, IV, x, 208) shows by an experiment similar to
those given above, that the water lost by lieulandite exposed over H^SO^ is
all regained in 1^ days.
1S80.J NATURAL SCIENCES OF PHILADELPHIA. 321
(3) The finely powdered mica was divided into two portions,
one of which Avas spread out on an open w^atch-glass, the other
placed in a crucible. Both were weighed, put in a desiccator over
sulphuric acid, and let stand unopened for two months. That in
the crucible lost 2.76 per cent, of water. That on the watch-glass
had lost 3. 87 per cent. On standing 3 or 4 minutes upon the
scale-pan it gained .53 per cent, of water from the air. Upon ex-
]iosure over sulphuric acid in the desiccator 24 hours longer and
then being quickly weighed, it was found to have lost 3.99 per
cent. It Avas now placed in an air-bath and kept at a temperature
of 100° C. for 4 hours. After cooling 15 minutes in the desiccator,
it was found to have gained in weiglit about ^ per cent., indicating
that the desiccation over sulphuric acid was more complete than
that in the air-bath at 100° C. That in the crucible lost on igni-
tion 5.97 per cent, of its weight.
(4). The powdered mica w.as placed in a watch-glass in a desic-
cator over sulpliuric acid.
Aftev27 days it had lost 2.28 per cent.
" 40 " " " 2.36 "
During weighing; it was enclosed in clasped w^atch-glasses. It was
now^ put in a crucible and ignited.
The dried mineral lost on 1st ignition, 5.18 per cent.
u u a 2d " 5.36 "
" " " 3d " 5.47 "
(5). The following direct determinations of water of crystal-
lization w^ere made from the mica, dried in a glass tube, corked
while weighing, and then ignited in a crucible.
Desiccation.
Time of
Desiccation.
If^nition
Loss of
water in
dried mineral
(a)
100° C. in air-bath.
24 hours.
15 min.
5.38 p. C.
(b)
u u
3 days.
20 min.
5.
(c)
over H,SO,
2 weeks.
3 times.
Mean,
5.00 "
5.32 "
Tliis determination is thought to be too high, including some
iiygroscopic water, since the mica in a tube cannot be perfectly
desiccated.
A mean of the three determinations of hygroscopic water ab-
sorbed over sulphuric acid gives 3.24 per cent., which deducted
322 PROCEEDINGS OF THE ACADEMY OF [1880.
from the total water, ^.5S per cent., giA^es for water of cr3'stalliza-
tion, 4.34 per cent. As will be seen below, a similar amount is de-
duced from Mr. Haines' experiments.
The following experiments by Mr. Haines have been kindly
placed at the disposal of the writer. They may be relied upon as
having been performed with great care.
I. The powdered mica is placed in a desiccator over concen-
trated sulphuric acid.
(a) Dried 15 days. Loss, 2.69 per cent.
(b) " 10 " " 2.89 "
II. The undried mica is heated in an air-bath at 100° C.
(a) Heated 3 hours. Loss, 3.14 per cent.
(b) " 3.33 "
(c) " 3.42 "
(d) " 51 " " 3.69 "
Weights. Per cent, of loss.
III. Weight before heating. 1.0880
Heated at 100°, 1| hours. 1.0613 2.46
" " 4i " 1.0598 2.59
Over H2SO4 and heated 5 hours
at 100°. 1.0558 2.96
Heated 2 hours at 100° and
cooled over H,SO,. 1.0613 2.46
"Weight. Total Loss from Inere- p.c. ofp.e. ol
loss. 100° C. ment total loss fr.
ofloss. loss. 100°O
lY. Weight before heating. .9035
HeatedatlOO° C. for Ihr. .8743 .0292 3.23
" * 100° " 2 " .8730 .0305 3 37
" 105° " 2^" .8715 .0320 .0015 .0015 3.54 0.15
" 119° " 1 " .8705 .0330 .0025 .0010 3.65 0.28
" full red heat 5 mill. .8350 .0685 .0380 .0355 7.58 4.34
" over blast lamp
1st time. .8270 .0765 .0460 .0080 8.46 5.27
" over blast lamp
2d time. .8280 .0755 .0450 8.35 5.15
1880.] NATURAL SCIENCES OF PHILADELPHIA. 323
Weight. Total L.OS? from Incre- p. c. of p. c. of
loss. 100^ C. ment total loss fr.
ofloss. loss. 100°r
V. Weight of iindried mica .S052
" at 100° C. .7827 .0225 2.79
" at 125° C. cooled
for 8 minutes. .7757 .0295 .0070 .0070 3.66 0.89
" at 150° cooled for
3 minutes. .7682 .0370 .0145 .0075 4.59 1.85
" at 170°-175° cool-
ed for 4 min. .7682.0370.0145 4.59 1.85
" 190° cooled for 3
minutes. .7647 .0405 .0180 .0035 5.03 2.30
VI. "Weight of undried mica. .9855
" at 100°C, heated
several hours. .9615 .0240 2.43
Below faint red heat. .9445 .0410 .0170 .0170 4.16 1.77
Heated to pale red at
bottom of crucible. .9320 .0535 .0295 .0125 5.32 3.07
Heated to bright red at
bottom of crucible. .9210 .0645 .0405 .0110 6.54 4.21
Heated to full red on
whole crucible. .9148.0707 .0467 .0062 7.17 4.85
VII. Total water.
(a) Loss of weight at red heat, 7.30 per cent.
(b) " " on ignition, 7.50 '•
(c) " " " 3 times, 7.86 "
From the above experiments of Mr. Haines in connection with
Nos. (1), (2) and (3) under "hygroscopic properties," we may
deduce the following percentages :
For total water, we have (IV), 7.58 p. c. ; (VI), 7.17 ; (VII, a,
b, c), 7.3a, 7.50, 7.86.
Mean total water, 7.48 per cent.
For h3'groscopic water, driven off at 100°, we have
Exp. (1) Exp. (1) Exp. (2) Exp. (3)
Analysis I. Analysis II. Heated 1| hrs. Reheated 3 brs. 3 hrs. 3 hrs.
3.12 3.43 3.21 3.57 2.79 3.09
Exp.IIaExp. lib Exp. lie Exp. II d Exp. III. Exp. IV. Exp. V. Exp. VI.
3 hrs. 5^ hrs, 5 hrs. 2 lirs. several hrs.
3.14 3.33 3.42 3.69. 2.96 3.37 2.79 2.43
324 PROCEEDINGS OF THE ACADEMY OF [1880.
A mean of these 14 determinations gives for hj'groscopic water
3.17 per cent.
Subtracting tliis fx'om the mean total water, 7.48 per cent., we
liaA^e for water of crystallization 4.31 per cent., an amount closely
agreeing with that deduced from the writer's experiments. The
desiccation over sulphuric acid in Exp. I is for too short a time to
completely extract the hj^groscopic water.
The exact state of the water cannot yet be regarded as certainly
established. There is no reason why a fixed temperature of 100°
C. should divide the hygroscopic water from the water of crystal-
lization. The above experiments show that the loss of water as
the temperature is raised above that point is a very gradual one.
It is difficult to see in what manner the water driven off at 190°
in experiment Y, differs from that driven off at 100°. It will be
seen hereafter that much of the water can be driven off without
exfoliation. Again, there is no sufficient reason why some of the
water absorbed by sulphuric acid in the desiccator may not be
water of crystallization. It has been long known that sulphate of
copper either at 100° C. or in a desiccator over sulphuric acid
loses much of its water of crj'stallization. M. Damour has shown
that chabazite loses nearl}^ half of its water in a desiccator. It
seems probable that Philadelphite, with other vermiculites, holds
its water in a similar manner. From the experiments here de-
tailed it would seem that we maj" define water of constitution to
be the more closely combined, and hygroscopic water the less
closely combined water of cr3-stallization ; and the distinction
between the three states of water then becomes a theoretical rather
than a practical one.
Temperature of Exfoliation. — The temperature at which exfo-
liation occurs is from 1,50° to 160° C. It has been found that the
exfoliation temperature is proportional directly to the original
volume of the substance, and inversely to the rapidity ^Df the ap-
plication of heat. The larger the piece experimented upon, the
higher the temperature necessary to make it exfoliate, and the
more rapidly the heat can be applied, the sooner will it exfoliate ;
as the following experiments will show.
(a) Yery small fragments heated on a watch-glass in an air-bath
began to exfoliate at 150° C.
{h) A large piece heated similarly did not exfoliate at 210° C.
(c) A piece was immersed in melted parafflne. At 100° C.
1880.] NATURAL SCIENCES OF PHILADELPHIA. .325
bubbles went off slowly, but there was no exfoliation. The tem-
perature being raised, it made tlie first movement at 160°, exfoli-
ated vigorously at 175°, and at 180° rose from its support to the
surface of the parafline.
(d) Another piece similarly immersed gave bubbles briskly at
1.30°, and began to exfoliate at 160°.
(e) Pieces thrown into melt'ed parafline whose temperature had
previously been raised to 1G0° C, immediatel}^ exfoliated and rose
to the surface.
(/) A large piece did not exfoliate even after the temperature
had been gradually raised to 225° C.
(g) Immersed in melted sulphur, it immediately exfoliated and
strongly effervesced.
(h) Immersed in concentrated sulphuric acid which had been
heated to 160° C, it immediately exfoliated and became pure
white, being completeh- and immediately^ decomposed. Immersed
similarh' at a temperature of 150° C. it exfoliated, but did not
become immediatel}^ white. At a lower temperature no exfoliation
occurred. A similar piece being similarly immersed and the tem-
perature raised, began to exfoliate at 130° C, and continued ex-
foliating as the temperature rose, though being meanwhile decom-
posed. This sudden change of form and color upon immersion in
hot sulphuric acid recalls a somewhat similar change in the efflor-
escence of protosulphate of iron when immersed in the same acid.
It is seen from these experiments that no absolute determination
of the exfoliation temperature is possible. By a verj- slow heat a
large proportion of the water (about 5 per cent.) can be driven off'
and the mica raised to a high heat without any exfoliation of
consequence. The following experiment illustrates this fact.
(/.) A piece of Philadelphite was cut into two equal portions.
One piece, heated suddenly on platinum foil to a red heat, exfoliated
to ten times its original volume. The other piece was slowly
heated in an air-bath. At 285° C. it had exfoliated but very
slightl\'. It was then taken out and heated on platinum foil to a
red heat, when it exfoliated very little more, becoming only one-
fourth the length of the first piece.
A similar experiment has been made upon heulandite and stilbite
from near Philadelphia. Both of these zeolites, as is well known,
exfoliate largely when held in the flame. It has been found that
if the}' are heated very slowlj- on platinum foil, they can be raised
326 PROCEEDINGS OF THE ACADEMY OF [1880.
to a white heat without exfoliation, and when afterwards held in
the flame, exfoliate but slightly. Apparently the water in Phila-
delphite is combined precisely as in the zeolites
It appears that it is as difficult to make a distinction between
water of crystallization ahd water of constitution as it is to make
one between the former and li3^groscopic water.
Amount of Exfoliation. — The amount of exfoliation is quite
constant at ten times the original volume.
Original thickness. Thickness after exfoliation.
Inches. Inches. Eatio,
.015 .13
.015 .17
.02 .2 1 : 10
.03 .25
mean 1 : 10
04 .47 I mean 1 : 10.4
.06 .68
/
These experiments were made by heating tlie mica on platinum
foil over the flame of a Bunsen burner. The heat must be sudden
in order to have a large exfoliation (i-. Exp't. k\ Exfoliation
takes place in one direction onlj', viz., at right angles to the
cleavage. No lateral expansion whatever occurs. When the
flame is applied to one side of the mica, that side exfoli^es the
most, and causes the exfoliating mineral to curve in the opposite
direction.
Force of Exfoliation. — It has been found that the force exer-
cised during the exfoliation of Philadelphite is enormous. In one
experiment a fragment of it while exfoliating lifted more than
50,000 times its own weight. The 'force of exfoliation is governed
by a law which is the inverse of that controlling the exfoliation
temperature. It may be stated thus : The force of exfoliation
increases directly with the rapidit}' of the expulsion of water, and
inversel}^ with the volume of the substance. The latter part ot
the law follows as a necessaiy consequence of the first part, since
the smaller the fragment, the more rapidl}- and completel}' can it
be heated. Various experiments were made, and though per-
lormed in an extremeh^ rough manner, will give an idea of this
force. To find what amount a given Aveight of the mica could lift
when exfoliating, iron pound-weights were placed upon the ring of
a retort stand and connected with the fragment of mica placed on
a support immediately below them. A pencil of chalk or gas
IsSO.]
NATURAL SCIENCES OF PHILADELPHIA.
32T
carbon resting loosel}' in a perpendicular position between the
mica and the centre of the weights connected them so that an^'
expansion of the mica would lift the weights from off the ring
on which thej^ rested. The blowpipe flame was now directed
from one side upon the mica.
G, retort stand.
b, ring.
(', support.
d, weight.
e, pencil of chalk.
/, fragment of mica.
In the following table of experiments, the first column repre-
sents the weight of the fragment of mica, and the second column,
the iron weight which was lifted by the exfoliating mica.
Philadelphite. Weight.
15 grains lifted 10 lbs. avoirdupois.
6
i,'.
10 lbs.
u
5i
ii
10 lbs.
u
n
U
10 lbs.
u
2
a
5 lbs.
u
u
u
2 lbs.
u
1
u
3 lbs.
n
2
3
a
2 lbs.
u
1
2
a
* 3 lbs.
u
1 .
2
u
4 lbs.
u
readily.
In the last experiment the four-pound weight was lifted up and
thrown ofl" the ring supporting it ; the weight lifted being 56, Ot 0
times the weight of the mica.
A remarkable motive power is here deA'eloped. That it is
OAving solely to the escape of the combined water is shown by the
tact that if the weights are so arranged that the mica can onlv
slightly expand, and, after heating, are removed, the mica will
expand no more, or very slightly more, upon further application
of heat, the water having been in great part expelled. If the mica
is confined under a weight so heavj' that it is impossible for it to
o28 PROCEEDINGS OF THE ACADEMY OF [1880.
exfoliate, and is suddenl}' heated by the flame, it occasionally
explodes with a loud report, throwing off fragments laterally into
the air.
It may be stated that the exfoliated mineral when powdered,
forms a handsome and permanent bronze powder not liable to
tarnish, and useful in the arts.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 329
A Potsdam Sandstone Outcrop on the S. Valley Hill of Chester
Valley. — Mr. H. C. Lewis remarked that an occurrence of Potsdam
sandstone on the farm of Mr. S. Tyson, near King of Prussia,
Montgomery Co., to which Mr. T. D. Rand had directed attention
last May, was of considerable interest. A recent examination of
the locality with Mr. Rand, had shown that the blocks of sand-
stone there found were not, as had been supposed, out of place,
but belonged to a narrow outcrop of the sandstone on the South
Valley Hill. It had a strike, and apparently a dip, nearly iden-
tical with that of the limestone in the valley below. In one place
the decomposed rock is quarried for white sand. Pits for iron
ore have been sunk in a very ferruginous variet}' of the same
rock. The exposure, which can be traced by blocks upon the
surface, suddenlj' comes to an end in a ravine, as though by a fault.
A tongue of sandstone blocks extends three hundred 3'ards or
more down this ravine, towards the valley, in a line at right angles
to the line of strike. On the farther side of the ravine, to the
east, no sandstone has been found, its place being filled by the
usual damourite slate of the South Valley Hill. The blocks of
sandstone therefore make an " L," the shorter arm of which
extends down the ravine. There is here an interesting example
of the work of erosion in carrying down these blocks to a lower
level. Whether or not the existence of a fault can be proved, the
occurrence of Potsdam sandstone at a new locality on the South
Valle}^ Hill is well worth}' of studv. This formation forms the
North Valle}^ Hill, but is almost totally absent on the South
Valley Hill. It is found here only in a few isolated patches. Its
place is supplied by a greenish damourite slate. If, as is sup-
posed, the North and South Valley Hills are opposite sides of a
synclinal trough which dips beneath the limestone of Chester
Vallej', it is curious that the rocks of each hill are so very dis-
similar. It is important that each one of the rare exposures of
sandstone on the South Valley Hill should be made known, and it
is thought that a determination of their relations to the adjoining
slates will greatly help to elucidate the geology of that region.
22
330
PROCEEDINGS OF THE ACADEMY OF
[1880.
July 6.
The President, Dr. Ruschenberger, in tlie chair.
Eleven persons present.
July 13.
The President, Dr. Ruschenberger, in the chair.
Six persons present.
July 20.
The President, Dr. Ruschenberger, in the chair.
Ten persons present.
July 27-
The President, Dr. Ruschenberger. in the chair.
Eleven persons present.
The death of Constantine Hering, M. D. was announced.
Fresh-water Sponges of Fairmount Park.^Mx. Potts reportetl
that he had found in a small stream within the limits of the late
Centennial grounds, Fairmount Park, Philadelphia, three distinct
species of Fresh-water Sponges, one of which appears to be un-
described and the others ditfer in important points from the pub-
lished descriptions. In anticipation of a more detailed paper
describing; these and some other forms which had come under his
notice, he said — that one of tliese known as the common green
sponge of this neighborhood, resembles the European Hjwngilla
Jacusfris in its general appearance and in the shape of its skeleton
and dermal spiculae ; but differs in that the seed-like bodies or
spherulfe are entirely smooth, showing no incrustation of curved
spined spiculae as described in the European species.
The second form was first seen as a thin rust-colored incrusta-
tion, afterwards discovered to consist of spherulfe forming a
continuous layer. Supposing this to be new he had named it
provisionally S. Morgiana : but later examinations of the living-
sponge had convinced him that it was identical with the N. fragilis
of Leidy.
The third was found creeping upon and around Aimcharis and
Willow roots, matting them together and thus forming loose, ir-
regular masses several inches in diameter ; color yellowish, light
or dark green, according to exposure to the light. Spherulae,
globular, light yellow or brown, rather numerous amongst the
1880,] NATURAL SCIENCES OF PHILADELPHIA. 331
roots and spicuke ; covered with long birotulate spicula^ radially
arranged ; foramen elongated into a tube flaring at its extremity
and dividing into 2 — 5 tapering, slender, curling or twisted tendrils,
believed to be as much as half an inch in length. The sareode
decomposes early in the season and most of the skeleton spiculte
are then washed awa^^ ; but these tendrils hold the mass of
spheruUie attached to the roots etc. above mentioned, awaiting the
spring germination. For this curious species he suggested the
name S. tentasperma or tendril seeded.
Dr. M. S. De Bey of Aix-la-Chapelle and Prof. Torquato
Taramelli of Pavia were elected correspondents.
August 3.
The President, Dr. Ruschenberger, in the chair
Sixteen persons present.
The death of James Ridings, a member, was announced.
Notes on Jarof^ite. — Prof. George A. Konig communicated his
<liscovery of Jarosife at the " Iron Arrow Mine,'' in Chaffee Co.,
Colorado.
The mineral occurs there in seams and cavities of silicious
thurgite and hematite, which iron ores crop out on the steep side
of a Porjihyry hill about (lOO feet above the Arkansas Rive]-, flow-
ing at a distance of two miles to the south.
The mineral appears in small, but Aery brilliant cr3''stals, isolated
and in groups ; also as aggregations of crystals which produce
crusts. It is remarkably crystalline, since no compact, or crypto-
crystalline masses were observed.
The crystals are rhombohedrons (resembling cubes), modified
by the basal plane. The speaker had not observed as yet a crys-
tal of sufficient size to be accurately measured. Hardness slightly
above selenite ; color, from light amber-yellow to deep brown.
Perfectly transparent. Lustre of crystal faces adamantine, resin-
ous on the fracture. Sp. gr. = 3.144.
The material used for analysis consisted of the aggregations
before mentioned, which showed an admixture of chalcedony and
of brilliant, black grains of thurgite ; these could not be separated
mechanically, being too small.
The mean of two analyses gave :
Fe,0:.
— 51.10
KoO
— 7.13
Na.,0
— 0.84
SO,
— 28.57
H,0
— 10.56
SiO,
— 2.40
100.80
332 PROCEEDINGS OF THE ACADEMY OF [1880.
Calculated from the analysis :
K.FeeS.O,, + 6H,0 .
FeiH.O, (Thurgite)
Excess of water
Chalcedony (SiO^)
= 89.58
= 8.6T
= 0.39
= 2.40
100.94
This result may be estimated as a confirmation of Richter's
analysis, which gave to Jarosite the formiila of " alunite,'' with
which it is isomorphous.
Thus 41unite = K^AlgS.O,, + 6A.,0
Jarosite = K^FceSiO,, + 6H,,0
The utmost care was given to the estimation of the alkalies, the
sulphuric acid and the water, as the question of constitution must
be dependent mainly upon them.
August 10,
The President^ Dr. Ruschenberger, in the chair.
Sixteen persons present.
August IT.
The President, Dr. Ruschenberger, in the chair.
Twelve persons present.
A paper entitled " Rhizopods in the Mosses of the Summit of
Roan Mountain N. C," by Jos. Leid}^, M. D. was presented for
publication.
August 31.
The President, Dr. Ruschenberger, in the chair.
Fourteen persons present.
The following was- ordered to be printed :
1880.] NATURAL SCIENCES OF THILADELPHIA. 333
EHIZOPODS IN THE MOSSES OF THE SUMMIT OF ROAN MOUNTAIN,
NOETH CAROLINA.
BY JOSEPH LEIDY, M, D.
In a trip to Roan Mountain, Mitchell Co., North Carolina, in
the early part of July, the writer was led to make some observa-
tions on the microscopic animal life, among the mosses on the
summit of the mountain. The top of Roan Mountain, at an alti-
tude of 6367 feet, forms an extensive grassy prairie, suitable for
pasture. It is adorned with broad patches of the beautiful Rhodo-
dendron catawbiense, and bordered Avith forests, chiefly of Firs —
Abies canadensis and A. Fraseri. The floor of the forests, made
up of broken granitic and gneissoid rocks and fallen timbers, is
thickly carpeted with a luxuriant growth of mosses, conspicuously
decorated at the time by the common Wood-Sorrel, Oxalis aceto-
sella. Chief among the mosses, each apparently attempting to
outvie the others in the exuberance of its growth, were the three
pretty Hjpnums — U. splendens^ H. crista-castrensis, and H. tri-
quetrum.
Clouds, dews, and frequent rains keep the mossy carpet more
or less moist or wet the greater part of the time, and it thus comes
to be a favorable habitation for man3^ of the humbler forms of
animal life. The shell-covered Rhizopods abound ; and the Wheel
Animalcule, Rotifer vulgaris, and the Water Bear, Macrobiotus
Hufelandii also find a suitable home in it. When the mosses
become more or less dr^', the animalcules they shelter become
torpid, and then again become active on the restoration of
moisture.
In water squeezed from the Il3^pnums, besides the animals just
indicated there were noticed a few young Anguillules, pollen
grains of Abies, starch grains, spores of lichens and fungi, ova,
vegetal hairs, etc. Few or no liAdng Diatomes or Desmids were
present.
The Rhizopods observed were as follows :
Nebela flabellulum. — Common. Nearly circular in outline,
usually slightly broader than long, and commonly with a short
neck or rim to the mouth ; colorless or with a feeble yellowish
tint ; composed of circular cancelli of variable size and propor-
OO-d
PROCEEDINGS OF THE ACADEMY OF
[1880.
tious and degree of distinctness. Individual specimens presented
the followinsi; measurements.^
a. Specimens of equal length and breadth, oi- of gr^eater breadth
than length.
1. Length 0.066, breadth 0.066, b'th of mouth or neck 0.018, 1'th of neck 0.006,
9
ii
0.066,
(i
0.069,
0.018,
0.006.
.3.
U
0.06(3,
U
0 072,
0.015,
0.006.
4,5.
t(
0.066,
i(
0.072,
0.018,
0 006.
6.
ii
0.072,
ii
0.078,
0.018,
0.003.
7.
4 i
0.078,
*(
0.078,
0.024,
0.006.
8,9.
4(
0.078,
(..
0.084,
0 018,
0.006.
10.
if
0.078,
(i
0.084,
0.024,
0-004.
11.
Ci
0.084,
fc(
0.084,
0.024,
obsolete.
12.
((
0.084,
((
0.09,
0.018,
0.006.
1-3.
-
0.084,
i(
0.09,
0.021,
0.006.
14-18.
a
0.084,
. (
0.09,
0.024,
0.006.
19.
t(
0.084,
t fc
0.096,
0.024,
0.006.
•20.
it
0.096,
£ i
0.096,
0.03,
0.006.
b.. Specimens slightly longer than broad.
21. Length 0.072, breadth 0.066, b'th of mouth or neck 0.018, 1'th of neck 0.006.
22,23." 0.084, " 0.078, " " " 0.018, " 0.006.
24. " 0.09, " 0.081, " " " 0.018, " 0.006.
25. '« 0.09, " 0.078, " " " 0.024, " 0.U06.
In many the sarcode was contracted into a ball, encysted, and
the mouth of the shell closed with an opercle. Sarcode colorless,
but sometimes mingled with yellow and brown food balls ; some
oleaginous in appearance. Mostly, a pore was readily distinguish-
able on each side of the body of the shell, about one-third the
distance of the length above the mouth ; and the wall of the shell
was observed to be sensibly thickened approaching the pore. The
shell was usually minutely and more or less distinctly cancellated,
the cancelli being circular, sometimes nearl^^ uniform, at others
greatly differing in size. Occasionally the cancellated condition
was so indistinct that the shell appeared fainth' granular and even
nearly structureless.
1 The measurements are given in divisions of the millepietre. As nearly
;ill the Ilbizopods indicated are compressed forms, the measurements are
given from their most convenient position as usually seen, that is to say, iu
the greater breadth.
1.
Length 0.06, 1
jreadt
h 0.036,
2.
0.066,
0.086,
3,
4. "
0.066,
0.039,
5.
0.066,
0.042,
6.
0.066,
0.042,
7.
0.072,
0 042,
8.
0.072,
0.048,
9.
0.096,
0 078,
1880.] NATURAL SCIENCES OF PHILADELPHIA. 335
Nebela coLLARis. — Occasional. Flask-like forms, with the
usual variations in the condition of the cancellated structure of the
shell ; sometimes finelj^ punctate, but mostly with distinct circular
oancelli, more or lesg uniform or greatly var3'ing in proportionate
size. In several specimens the cancelli of the shell appeared to be
like minute lenses or spheres, and to present an external con-
ACxity. Individual specimens measured were as follows :
" 0.015.
" 0.018.
" 0.015.
" " 0.018.
" " 0.018.
" 0.018.
" " 0.024.
Hyalosphenia tincta ? — One specimen onl3^ Sarcode encysted
as a ball 0.048 diameter, containing much brownish food and
bright yellow oil-like globules. Shell structureless, pale yellowish,
with a pair of pores piercing the body above the junction of the
neck. The specimen looked like a Nehela fiabellulum, but the yV
inch magnif3'iug power showed no structure to the shell.
Length 0.060, breadth 0.072, breadth of m lutU 0,024, length of neck 0.0045.
DiFFLUGiA GLOBULOSA. — Rare. Small forms with shells of fine
sand and dirt. From hemispherical to globular and with circular
mouth.
1. Breadth of shell 0.06, height 0.042, brei.lth of mouth, O.OIS.
2. '• " 0.06, " 0.018, " '• 0.024.
DiFFLUGiA CONSTRICTA. — Rare. Shell of yellowish dirt and.
sand.
Length 0.072, breadth 0.072.
DiFFLUGiA ARCULA. — Rare. Shell hemispherical, brownish, in-
eorporated with dirt and fine sand ; mouth trilobate.
Breadth 0.132, height 0.09, breadth of mouth 0.048.
Heleopera petricola. — Occasional. Purplish brown, with
variable proportions of incorporated sand.
1. Length 0.09, breadth 0.078, breadth of mouth 0.042.
2. " 0.096, " 0.078, " " 0.036.
336 PROCEEDINGS OF THE ACADEMY OF [1880.
AssuLiNA SEMiNULUM. — Somewliat frequent. Of varied size and
hue ; from nearly colorless to dark brown, and mostly lighter at
the neck. Mostly empty shells ; often living specimens.
].
Leng
th
0.0^.6,
b
re.idili 0.03 ;
b
readth of
moL
ith 0.009.
2.
0.042,
0.036 ;
0.012.
3.
0.048,
0.042 ;
0.012.
■i.
0.054,
0-048;
0.018.
•3.
0.072,
0.072 ;
0.024.
6.
0.078,
0.072 ;
0.024.
EuGLYPHA AREOLATA, Ehr. — Small compressed spineless forms,
occasional. A living specimen measured as follows :
Length 0.06, breadth 0.048 ; breadth of mouth 0.015.
EuGLYPHA STRiGOSA. — Occasional. An empty shell measured :
Length 0.06, breadth 0.042 ; breadth of mouth 0.012.
Trinema enchelys. — Common. Small forms of much variety.
1. Length 0.03 , breadth 0.024.
2, 3. " 0.036. " O.O.S.
4. " 0.036, " 0.036.
5. " 0.042, " 0.03.
In some positions, on dripping ledges and cliffs, from among
the greater profusion of H^'pnums, there grew cushion-like masses
of bright pea-green Sphagnum, sometimes rose tufted. Water
.squeezed from the Sphagnum, exhibited many Diatoms, a few
Desmids,' pollen of Abies, starch grains, spores, etc. It also
contained many Rhizopods, an occasional Infusorian, a few
Auguillules, but no Rotifers nor Water-bears, Of Rhizopods
the foUowino- were observed :
'&
Nebela collaris. — Abundant and of much variety in shape and
size and in the arrangement and distinctness of the cancellated
structure of the shell. Compressed pyriform, or flask-shaped,
with oval or ovoid body and with neck of variable proportionate
length and breadth. Cancellated structure very variable ; mostly
of circles, minute and nearly uniform, or greatly varying in size
among themselves or in proportion with the shell. Dead shells
most frequent ; in living ones observed, the sarcode mostly con-
tracted and inactive, often encysted ; least frequently in an active
1880. J NATURAL SCIENCES OF PHILADELPHIA. 337
locomotive condition. Small forms common ; giant forms few,
Individual specimens noted as follows : .
1. Length 0.078, breadth 0.06 ; breadth of mouth 0.018. Compressed oval ;
living.
2. Length 0.078, breadth 0.06 ; breadth of mouth 0.018. Oval with short
neck. Empty shell ; cancelli circular, variable, the largest 0.006, the smallest
0.003.
3. Length 0.078, breadth 0.06 ; breadth of mouth 0.012. Oval with short
neck; living, active ; nucleus, 0.12.
4. Len:;th 0.081, breadth 0.054; breadth of mouth 0.018. Empty shell,
with sharply defined circles, large and small, together with a few rods.
5 Length 0.084, breadth 0.042; breadth of mouth 0.018; with neck 0.018
long. Empty shell, with minute circles on the neck, but unusually large in
proportion to the shell on the body where they ranged from 0.006 to 0.012.
6. Length 0.084, breadth 0.048; breadth of mouth 0.018. Flask-like empty
shell, with minute circular cancelli 0.003 or less.
7. Length 0.084, breadth 0 0-54; breadth of mouth 0.012. Flask-like empty
shell minutely and uniformly cancellated.
8. Length 0.084, breadth 0.05; breadth of mouth 0.015. Oval, empty.
9. Length 0.09 , breadth 0.072 ; breadth of mouth 0.^2 1. Flask-like. '
10. Length 0.096, breadth 0.078 ; breadth of mouth 0.024. Pyriform ; neck
0.006 long; cancelli circular, variable in size, a few on the fundus to 0.012.
Sarcode an encysted ball, with yellowish oil-like food globules ; diameter of
ball 0.048.
Some giant forms especially noted were as follows :
11. Length 0.18; breadth 0.09; breadth of mouth 0 042. Living; shell
nearly replete with sarcode, colorless but containing a multitude of bright
yellowish and brown globules from 0.006 to 0.012.
12. Length 0.21; breadth 0.12; breadth of mouth 0.048. Empty shell of
faint yellowish tint ; basis of structure faintly and uniformly punctate with
only distinct minute circular cancelli approaching the fundus.
18. Same size as preceding. Shell mostly of minute circular cancelli, larger
near the fundus and there mingled with a few square ones. Shell closed by an
operculum. Sarcode contracted into an oval mass 0.144 by 0.072.
14. Length 0.192; breadth 0.102; breadth of mouth 0.048. Shell with
minutely cancellated structure. Sarcode in a ball 0.084 by 0.072.
Nebela flabellulum. — This form comparatively rare. Shell
nearly circular in outline, with a short neck, mostly composed of
minute circular cancelli more or less nearl}^ uniform or variable ;
rarely of elliptical cancelli. ^
338 PROCEEDINGS OF THE ACADEMY OF [1880.
1. Length 0.072; breadth 0.072; neck 0.003 long: breadth of mouth 0.009.
2. " 0.072; " 0.078; " 0.003 " ; " " 0.009.
3. " 0.678: " 0.078; " 0.0015 " ; " " 0.009.
4. " 0.084; '' 0.078; " 0.003 " ; " " 0.012.
Shell with minute elliptical caucelli.
DiFFLUGiA CONSTRICTA. — Rare. Only a few specimens seen.
Small forms, with shell of minute sand grains and yellowish dirt.
Pyriform viewed from the front or back.
1. Length 0.06 ; breadth 0.048
2. " 0.078; '• 0.048; breadth of mouth 0.024.
3. " 0.108; " 0.102; breadth opposite moulh 0.078.
DiFFLUGiA PYRiFORMis. — Rare. Shell of dirt and fine sand.
1. Length 0.084 ; breadth 0.048; breadth of neck and mouth 0.024.
DiFFLUGiA ARCULA. — Rare. Shell yellowish, incorporated with
more or less brownish dirt and sand. Form hemispherical ; mouth
trilobed.
1. Breadth 0.132; height 0.09.
Centropyxis aculeata. — Rare. Shell of pale brown chitinoid
membrane incorporated with more or less dirt and sand ; with
coarser grains of the latter along the course of the usually six
spines. Mouth oval, with a more or less sinuous border.
1. Length 0.096 ; breadth 0 084 ; breadth of mouth 0 024.
2. " 0.12 ; '• 0.096; mouth 0.03 by 0.024.
Heleopera petricola Occasional ; shell incorporated with
more or less dirt and sand, and of a purplish brown tint,
1. Length 0.09 ; breadth 0.078; breadth of mouth 0.042.
2. " 0.108; " 0.09 ; " " 0.036.
EuGLYPHA AREOLATA, Ehr. — Small compressed forms, without
spines or other appendages. Abundant. Apparently from six to
fifteen teeth to the mouth of the shell. Mostly emptj^ shells.
Often living specimens.
1,
2.
Le
agth 0.0i2,
bre
idth* 0.021;
bi^adih of
mou
h 0.012.
10 teeth.
.1
0.042,
0.024
0 009.
6 teeth.
4,
5.
0.U48,
0.024
0.012.
10 teeth.
6.
0.048,
0 03
0 012.
8 or 10 teeth?
7.
0.05 I,
0.024
0.012.
6 teeth.
8.
0 054,
0.03
0.012.
6 teeth.
9.
0.102,
0.72
0.036.
15 teeth?
1880.] NATURAL SCIENCES OF PHILADELPHIA. 339
EuGLYPHA STRiGOSA. — Compressecl, hirsute forms. Occasional,
Usually with about tea teeth to the mouth of tlie shell ; scales
distinct ; finel}' hirsute all over except near the mouth.
1. Length 0.102, b;-eadth 0.072; breadth of mouth 0.021 ; hairs 0.008 long.
2. " 0.108, " 0.06 ; " " 0.021; •' 0.012 "
EuGLYPHA BBACHIATA — One emptj^ shell observed, with but one
divergent spine, and five or six teeth to the mouth.
1. Length 0.102; breadth 0.04 ; breadth of mouth 0.012 ; length of spine 0.042.
EuGLYPHA CRiSTATA. — One empty shell, with acute fundu=;,but
without spines, and four teeth to the mouth.
1. Length 0.54 ; breadth 0.012; breadth of mouth 0.009.
EuGLYPHA ciLiATA. — Comprcsscd forms, with short spines or
hairs along the acute lateral borders. Rare. Nucleus 0.018.
1. Length 0.108; bre.idth 0.06 ; breadth of mouth 0.021 ; hairs or spines 0.012
long. Ten teeth lo mouih of the shell.
2. Length 0.102; breadth 0.072; breadth of mouth 0 024; spines 0.008 long.
Nucleus of sarcode disiinctly and uniformly granular (breaking up
into spores?), 0.018 diameter. Ten or twelve teeth to mouth of the
shell.
Placocista spinosa. — One specimen observed, living, but the
ample sarcode contracted and containing a transversely oval nucleus
with two nucleoli.
1. Length 0.084, breadth 0.054; breadth of mouth 0.021. Lateral spines
short, hair-like, single or in pairs, 0.009 long. Nucleus 0.021 by 0.018 :
nucleoli O.OOo.
AssuLiNA SEMINULUM. — Moderately frequent ; from nearl}'
colorless to dark brown, mostly lighter at or near the mouth.
Living and dead specimens observed.
1. Length 0 042, breadth 0.036, breadth of mouth 0.012.
2.
0.048,
0036,
" 0.012.
3.
0.072,
0.048,
" 0.018.
4.
0.072,
0.072,
" 0.024.
5.
0.078,
0.078,
" 0.024.
6,7.
0.084,
0072,
" 0.024.
Trinema enchelys. — FrequeiTt and of varied form and size,
though none of the largest variety observed. Usuall}' pyriform ;
often oval ; rarel}^ obovoid; of varied pi-oportionate length and
breadth, of narrowing opposite the mouth, and degree of obliquity-.
Most!}' dead shells, though frequent living individuals observed
Ranging from 0.024 to 0.0T2 in length. Specimens presented the
following measurements.
340
PROCEEDINGS OF THE ACADEMY OF
[1880.
1. Length 0.024, breadth 0.015; breadth of mouth 0.006. Pyriform.
2.
0.03,
3.
0.086,
4,5.
0.042,
6.
0.045,
less
spores from
7,9,
0.048,
10.
0.051,
11.
0.054,
12.
0.054,
13.
0.057,
14:
0.06,
15.
0.06,
16. 17.
0.072,
18.
0.072,
0.018;
0.024;
0.08,
0.03;
0.012.
0.012.
Obovoid.
Pyriform.
0.012. Contained 6 color-
0.08 ; breadth of mouth 0.012,
0.36;
0.03;
0.036;
0.086 ;
0.03;
0.042 ;
0.03;
0.08 ;
" 0.015. Nucleus 0.012,
" 0.015.
" " 0.015.
Lateral view of a specimen.
It is worth}' of special remark that among the Rhizopods of the
sphagnum of Roan Mt., there were observed no individuals of
Hyaloi<phenia impilio and H. elegans, which are so common in the
sphagnous swamps of the eastern plains.
1880.] natural sciences of philadelphia. 341
September 7.
Dr. R. S. Kenderdine, in the chair.
Fifteen persons present.
On large Sphene from Canada. — Dr. A. E. Foote recorded the
occurrence of crystals of sphene of unusual size in the county of
Renfrew, Canada, near tlie upper part of the navigable portion of
the Ottawa River. The largest crystal collected weighed 23^
pounds or raorcthan twenty times as much as the largest liereto-
fore found.
September 14.
The President, Dr. Ruschenberoer, in the chair.
Twenty-eight persons present.
The death of Prof. S. S. Haldeman, a member, was announced.
On the Timber Line of High Mountains. — Mr. Meehan re-
marked that on the tops of most high mountains we find a total
absence of ligneous plants. The highest alpine vegetation con-
sists for the most part of acaulescent perennials. Lower down
we may find some woody species, and often we come to dwarfed
forms of trees of species, which, still lower down, form forests of
considerable height, and which as timber trees make what is
known to mountain travelers as the ''Timber Line." Thus in the
mountains of Colorado, the forests commence at about 1000 feet
above the sea level, and continue up to about 11,000 feet, where
they suddenly cease, and form at that elevation what is there known
as the " Timber Line." On Gray's Peak he found Pinus aristata,
Pinus Jlexilis, Abies concoJor, and Abies Engelmannii, with
some willows forming the timber line. The Coniferous trees
were probably 30 or 40 feet high, and it was interesting to
note that this tall timber as suddenly ceased, as if a wood had
been cut half awaj' bj^ a woodman's axe. But at once commencing
where the tall timber ceased, the same species exist as dwarf
stunted shrubs seldom exceeding 3 or 4 feet in height, and often
but a foot, though trailing widel}' over the groinid. In tiiis
stunted condition the species would often extend some fifreen
hundred feet higher up, or half way from the recognized timber
line to the top of the mountain. Other observers have noted
that the average of 11,000 feet marks the entire timber line of
the Rocky mountain range.
So far as he knew this peculiar timber line has been referred
wholly to climatic conditions, of which temperature and moisture
342 PROCEEDINGS OF THE ACADEMY OF [1889.
have been regarded as the chief elements in producing the results.
That admirable botanist and energetic collector, Dr. C C. Parry,
in a paper on the Rocky mountain alpine region, published in the
"Proceedings of the American Association for the Advancement
of Science " for 1869, p. 249, remarks that the most satisfactory ex-
planation is that the so called timber line marks the extreme point
of minimuvi temperature below which no exposed phenogamous
plant can exist. All that survives above this point does so by
submitting to a winter burial of snow, beneath which protecting
cover it is enabled to maintain its torpid, existence.
The great objection which this purely meteorological view pre-
sented to Mr. Meehan's mind was that the dwarfed and gnarled
coniferae extending so many hundred feet up the mountain sides,
never produced seed, and we are reduced to the alternative of
believing either that the seeds have been carried up the mountain
sides in enormous quantities and to enormous distances from the
fruitive trees below by winds, or else that there were seed bearing
progenitors of these scrubby pines, beneath the tall protecting
branches of which they had their earliest stages of growth. He
was satisfied from subsequent observations in the mountains of
North Carolina, and in the White Mountains of New Hampshire,
that this last view is the correct one, — that large timber trees at
no very remote period extended much further up the mountain
sides than thej^ do now, and that they have since disappeared for
reasons presently to be stated, leaving only the 3'ounger trees to
struggle on as best they may.
Roan Mountain in North Carolina is about 6300 feet above the
level of the sea. Timber extends to its summit on some parts of
it, while in other parts it is destitute of timber for many hundreds
of feet down its sides. The species on the summit is Abies
Frazeri, and Abies nigra. Oak and other trees come occasionally
to near the top and at about 6000 feet he measured a black oak —
Quercus tinctoria, that was 5 feet in circumference at 3 feet from
the ground, and was perhaps 40 feet high. The places destittite
of trees were the steep declivities, — while those on which the
trees were growing were of a more level character. Further down
the mountain sides the steep inclines would be clothed with forest
growth, as well as those of a more gradual ascent. It is of the
summit only that the differences in inclination, presented different
forest aspects. But in the spaces clear of" Balsam "' as the Abies
Frazeri is popularlj^ known, an occasional one of good size would
be seen. In the close Balsam woods, both on the summit and
lower down the mountain sides, crops of young plants would be
found under the mature trees, but, what was very remarkable,
there had evidently been no young trees started till the parents
were near maturity. A large area with trees 30 or 40 feet
high would have an undergrowth of young ones a foot or so high,
while other areas of younger trees, would have innumerable small
seedlings growing among the damp moss beneath them, and it
1880.] NATURAL SCIENCES OF PHILADELPHIA. 343
was further interesting to note tliat in most cases the crops of
young plants in each area were about the same age in each case,
as if the seeds in the several locations had all started to grow
together in some one particular year, and probably at no other time.
On the naked ^^laces, where few or no trees were now found, the sur-
face would be closely covered bj^ a matted growth of a grass almost
peculiar to that I'egion, Danthonia compressa, but a close examina-
tion of the surface showed occasional tracts of deep vegetable
mould which had been formed by ages of deca^ying Hypnum or
Sphagnum moss, and the evident remains of roots, just as we now
find under the Balsam trees, and there is no doubt from these
facts that these steep upper declivities were once clothed with
trees and mosses, to which the grass previously named succeeded.
With these facts in mind he examined the arboreal features of
the White Mountains in New Hampshire. On Mount Washington,
which is a little over 6000 feet, the timber runs up to about 4000
feet ; while Mount Webster, a mountain forming the southern
peak of the same chain, and about 4000 feet high, has little timber
above 3000 feet. Clearly, climatic reasons will not account for
these peculiarities. On Mount Washington there is much of the
same character as distinguishes the foi'ests of the Rocky Moun-
tains. As already noted the timber line becomes marked at about
4000 feet. For at least another thousand feet we meet witli
sci'ubby bushes of Abies Balsamea, Abies yiigra^ and Abies alba,
with some Betula papyracea. Beyond this, and almost to the
summit, an occasional specimen of one or another of the conifers
may be seen. As noted in regard to the Colorado scrubbj^ growth,
none of these had ever produced seed ; nor was it at all probable,
from a careful survey of the locations, that many of the areas
could have been seeded by the winds, however strong, bringing
the seeds up these mountain heights. Moreover, there were many
cases where there were intermediate areas clear of all scrubby
spruce plants, and where seeds could be brought by winds in
these modern times much easier than to the heights above.
Besides this, it was evident that many of these dwarfed specimens
were of immense age. Some that he examined were certainly
fifty 3'ears old, though the stems at the ground were no thicker
than his wrist, and, trailing on the ground, occupied but
16 or 20 square feet of space. There seemed to be but little
doubt that at some time in the past Mount Washington had
forests of conifeme at much higher elevations than at present, if
not perhaps clean up to the summit ; that these scrubby plants
now there were seedlings that had sprung up under the elder
ones, and that in time the older ones were destroyed, leaving the
small ones beneath alone to their fate.
An examination of different parts of Mount Washington shows
not only that this is the true explanation of the absence of good
timber beyond what is known as the timber line, but that the
same law is in progress to day as in centuries past. Illustrations
344 PROCEEDINGS OF THE ACADEMY OF [1880.
of this are numerous. There is now a railroad running straight
up the mountain side from the base to the summit. Near the
timber line, a cut had to be made through an area covered by
mature Balsam Firs. This cut was about 8 or 10 feet deep.
Under the trees moss and dead roots and old fir leaves had made
an earthy strata of a foot, or in places, more in depth. The moss
was still green from the rains, melting snows, and fogs of this
elevated res-ion, and sustaining the various kinds of low veoeta-
tion common to these alpine heights. Young firs were springing
up in great abundance. But all the larger trees were dead, though
here and there might be seen a branch with a few lingering green
leaves. This mass of dead, standing timber occupied several acres.
The reason for their death was evident. The railroad cut showed
that the forest stood on a mass of large but loose gneiss rocks,
through which the waters from the two thousand feet of loose rock
above rushed as soon as the railroad cut was made, carrying with
it all the earthy matter on which the larger trees subsisted, but
leaving the tough turf}^ matter at the surface, on which smaller
trees of the same sort may live for many years, though the larger
ones cannot longer exist. With the death of the larger trees
there is, of course, an increase of light, and then the Sierochloe,
with other grasses and sedges, speedily take possession, holding
together the loose soil, and even permitting in many cases an
increase of the earthy layer, b}^ holding much of the disintegrated
rock which may be washed or blown on from above. Carefully
examining patches of scrubby spruces above the timber line, it is
not uncommon to find dark patches of vegetable mould evidently
the remains of large trees that have been growing where now only
the masses of small scrubby plants exist. In some places a sharp
stick may be pushed down among the scrubby firs and spruces,
and the earth found to be but a foot or so deep over the loose
rock below, from which the earth has been wholly washed away.
Again, there are some places often nearly an acre in extent where
the scrubby firs are still standing, dead, from the earth having
been washed away from below upwards, not leaving enough for
even the moderate demands of these little bushes.
In view of the facts detailed we may conclude that at the
elevation of these mountain chains, the lowland veafetation was
carried up at the same time. The summits, covered by luxuriant
forests would present a cooler surface to the moist clouds, and
there would be less condensation than on bare sun-warmed rocks,
and deep snows would be less frequent, and not suflicient to inter-
fere much with arboreal growth. But the rain would of necessity
carry down the earth and disintegrated rock to lower levels ; and
the melting snows, such as there were, would make this downward
progress of the soil continuous. In some mountains where the rock
was easily broken by frost, as in Colorado and the White Mountains,
it would be very difl!icult for the soil to hold its own against these
forces of gravitation ; but on more solid rock the mass of tree
1880.J NATURAL SCIENCES OP PHILADELPHIA. 345
roots protectino- the rock, and retaining the earthy matter would
longer hold its own. In the former case with the gradual washing
away of the earth the larger trees will have to find a lower
level ; the summit condensing more moisture, and having a cooler
atmosphere, would form heavier masses of longer enduring snow,
and thus keep down from tall growth the younger trees left as the
older and larger ones retired. They would have to be low bushes
by the absence of earth for vigorous groAvth, and remain trailing
bushes, through the superincumbent and long continued mass of
snow.
We thus see that though a long continued mass of snow has
much to do in marking a timber line, that line is precedent to the
snowy mass. The primar}^ cause is the gravitation of disinte-
grated rock— tlie movement of the hill top towards the sea. From
the moment the mountain reaches its highest point it commences
its downward march. The entire reduction of the highest to a
level with the plain is but a question of time. The frost and rain
and melting snow will do it all, and this reduction, bringing down
not only the earth, but cold-loving plants to warmer levels, must
continually change the aspects of vegetation, as well as perpetually
vary the timber line.
In low hills as well as in high mountains the forces of gravita-
tion are also at work. But the sides are seldom so steep as in the
loftier hills, — the rains do not gather with such force nor are
the melting snows of near the same duration. There are sudden
washes, but not the continuous roll of the earth to the bottom.
In time they maj^ exhibit the same phenomena of the disappearance
of species from their summits as their loftier brethren ; but the
centuries here will gather much more slowly to produce a similar
effect.
In conclusion he would say briefly that the " timber line " of
high mountain tops results from the washing down of the earth
from the higher elevations.
Mr. Redfield remarked that there could be no doubt that
influences other than climatic (such for instance as the washing
away of soil, mentioned by Mr. Meehan) do often modify and
change the timber-line upon mountains. But he was unable to
accept Mr. Meehan 's views as to the insignificant part played by
climatic causes, and still held them to be the prevailing factor in
the problem. Dr. Parry's explanation by the weight and depth of
winter snows might not always be the correct one, but snow and
ice must be very important agents, and Mr. R. thought that in
considering climate, we should have regard not merely to the
present period, but to past great secular periods. He then referred
to the glacial age, when not only the White Mountains, but all
New England was capped with a vast ice-sheet, which under secular
changes gradually retreated, leaving only the mountain tops
covered. The slow retreat of the glacial covering was followed
23
346 PROCEEDINGS OF THE ACADEMY OF [1880.
b}' the advance of fitting arboreal vegetation, until a point was
reached when the present climatic conditions were such as to
limit any higher advance of the trees.
September 21.
The President, Dr. Ruschenberger, in the chair.
Twenty-eight persons present.
Bone Caves of Pennsylvania. — Prof. Leidy remarked that in
the early part of August, in company with Dr. T. C. Porter of
Easton, he had visited Ilartman's Cave, in the vicinity of Strouds-
burg, Pa. They had been invited by Mr, T. Dunkin Paret, of
that place, who had recently undertaken the exploration of the
cave, and had obtained from it an interesting and important col-
lection of animal remains, which had been submitted to Prof.
Leidy 's examination.
The cave is situated about five miles from Delaware Water
Gap in a ridge which separates Cherry Valley from the valleys of
the Pocono and McMichael's Creeks. The ridge is an anticlinal
fold of the Ileiderberg or Upper Silurian limestone, and the cave,
occupies the axis of the fold and opens in the face of a clitf formed
by a cross section of the ridge. An accumulation of debris forms
a slope at the base of the cliflT, and above the deliris and just
below the arching roof of the cave, a low passage waj^ has long
been known into which adventurous bojs would creep.
Mr. Paret commenced the exploration by having a passage dug .
through the del)ris to the entrance of the cave, and then extended
the trench within the latter for upwards of a hundred feet, and to
a depth siifficient to walk erect. At one place within the cave the
digging was carried to^the rock floor. It would thus appear that
the cave is occupied by a bed of clay about 10 feet in depth. On
this is a thin layer of stalagmite and on this again about a foot of
black frial)le earth mingled with animal and vegetal remains.
No remains have been found imbedded in the. cla}^ nor on the
rocky floor in the pit dug through the latter. *
Prof. Leidy supposed that during the glacial period, a stream
of water, from melting snow and ice at a higher level, had made
a passage way through the fissured limestone of the anticlinal axis
and had left in it the abundant clay deposit. When the cave
ceased to be a water course the layer of stalagmite was formed
and subsequently the more frialjle eartli accumulated from mate-
rials, such as (lust and leaves, blown in and mingled with the
remains of animals, occupants of the cave, and of their food. The
recess of the cave above the clay floor api)ears to have been too
small to be inhabited by the larger carnivorous animals or mah,
and therefore no large entire bones of these have been found in
the ossiferous stratum.
1880.] NATURAL SCIENCES OF PHILADELPHIA. Z41
The remains thus far discovered are of such interest as to
encourage Mr. Paret to continue furtlier exploration Most of
these collected to the i)resent time were exhibited by Prof. Leidy,
and consist of the following :
Numerous fragments and splinters of limb bones of smaller and
large animals, man}- or most of which exhibit the marks of being
gnawed, whether by rodents or small carnivores is somewhat im-
certain. A few also show the marks of canine teeth, of medium
sized carnivores. Some of the splinters pertain to such large and
strong bones as to render it questionable whether they were pro-
duced by even our largest carnivores, and probably are the rem-
nants of human feasts, in which the bones were crushed to obtain
the marrow. Numerous bones and fragments of others of the
smaller and smallest animals. These include especially limb
bones, and lower Jaws, and less frequently skulls, fragments of
others and vertebrae. Many of these are also gnawed, while many
are not.
The fragments of larger bones may be supposed to have been
conveyed into the cave by small carnivores. A few pieces of bone
are somewhat charred ; and a small fragment of a lower jaw, con-
taining a molar tooth, of the Bison, also apparently exhibits
the marks of fire. This probably is a remnant from a human
feast, which may have been carried into the cave' by some small
gleaner.
All the bones and fragments together amount to about half a
bushel. Most of them pertain to animals of a kind still living,
though some of these no longer belong to the fauna of our state,
and a few of the remains are those of extinct animals. How far
the remains of different species are cotemporary is uncertain,
though it is most probable that they were introduced through a
long succession of years from the time following the glacial period.
The remains of extinct animals consist of an incisor tooth and
half a dozen molars of the great rodent Gasforoides ohioensis^
and portions of the upper and lower jaw, with teeth, of a young
Peccary, the Dieotijles nasutus, previously known only from a
single fragment of an upper jaw, discovered in Indiana, (Extinct
Mammalia of North America, 385, pi. xxviii, figs. 1, 2. Jour.
Acad. Nat. Sc, vii, 1869).
The remains of animals no longer living in Pennsylvania are as
follows :
Bones and teeth of the Caribou or Woodland Reindeer,
Rangifer caribou.
A fragment of the lower jaw containing the last molar tooth, of
the Bison, B. americanus.
Man}- lower jaw halves, and other bones and teeth of the Wood-
"rat, Neotoma fioridana. Most of these ai'c of comparatively large
size, and of the character of similar remains referred by Prof.
Baird to a supposed extinct species, with the name of Neotoma
348 PROCEEDINGS OF THE ACADEMY OF [1880.
magister, (U. S. P. R. R. Exp. & Surveys — Zoology, viii, 1857,
498).
Remains of other mammals are as follow : Lynx, Felis cana-
densis; Wolf, Canis lupus; Gray Fox, Vulpes virginianus ;
Skunk, Mephitis mephitica ; Weasel, Putorius ermineus ; Raccoon,
Procyon lotor ; Mole, Scalops aquations ; Dusky Bat, Vespei'tilio
fuscus; Little Brown Bat, V.subuJatus; Woodchuck, Arctomys
monax ; Porcupine, Erethizon dorsatus ; Beaver, Castor fiber ;
Muskrat, i^zfeer zibethicus ; Gray Squirrel, >Scf»rus carolinensis ;
Ground Squirrel, Tamias sfriatus ; Gray Rabbit, Lepus sylvati-
cus ; Meadow Mouse, Arvicola riparius ; White-footed Mouse,
Hesperomys leucopiis ; Deer, Gervus virginianus: Elk, Gervus
canadensis.
Arnong the remains, none have been identified as positively
pertaining to our domestic animals, unless, perhaps, a pair of
specimens are to be referred to this category. The specimens are
the complete isolated first and second large molars of a foetal or
new-born Horse 1
The collection further contains numerous bird bones, chiefly of
the Wild Turkey, Meleagris gallopavo ; some of turtles, the Box
Turtle, Gistudo clausa^ the Snapper, Ghelydra serpentina, etc. ;
and others of several species of snakes.
In the same stratum were also found a number of shells of
moliusks, chiefly Helix albolabris, H. alternata, and H. tridentata.
Also a valve of Unio complanatus.
Of A^egetal remains there were a few small fragments of charcoal,
and many seeds, consisting of those of the Dogwood, Gornus
iJorida, Pig-nut, Gary a porcina, and Walnut, Juglans nigra.
The human remains are of an interesting character. One is a
large stone celt of hard brown slate, obtained from the bone earth
some distance within the cave. There are five bone awls, several
of which exhibit marks of gnawing. Some of these were found
in the cave, and others in the outside debris. An implement con-
sists of the prong of an antler worked so as to be barbed on one
side, and was probably used as a needle for making nets.
A small implement of bone, resembles in its present condition
a crochet needle such as is iiow employed by ladies in making
worsted work. It is much gnawed away on one side, and looks
as if it may have been like an ordinary needle with a perforation,
and this now rendered incomplete from the gnawing.
Another implement is a fish-hook worked out of bone.
Such bone implements are among the rarest of human relics in
our portion of the countr}' .
Another remarkable relic is a cone shell bored through the axis
as a bead. The shell is a marine species, Gonus tornatus, found
on the western coast of Central America. Its presence among the
cave remains, would indicate an extended intercourse among the
inhabitants of early times.
1880.] NATURAL SCIENCES OF rHILADELPHIA. 349
The investigation of the interesting collection of remains secured
by Mr. Paret, had led Prof. L. to examine a small collection of
bones in the Museum of the Academy, which had been presented
upwards of thirty years ago, as a sample of many of the same
kind discovered in Durham Cave, Bucks Co., Pa. The cave
appears to have since been obliterated in the quarrying of lime-
stone. At the time of the presentation of the specimens of bones
the}' were recognized as pertaining to existing species of animals
and were therefore regarded as of little interest, though fortu-
nately they have been preserved.
Prof. Leidy remarked that these bones Avere of the same charac-
ter as those of Hartman's Cave, and he had distinguished among
them the followino- :
The Black Bear, Ursus americanus : Raccoon, Skunk, Gray
Fox, Deer, Moose, Alee americanu>< ; Woodland Reindeer, Bison,
Gray Squirrel, Rabbit, Beaver, Muskrat, Porcupine, Woodchuck,
Woodrat, Wild Turkey, Box Tortoise, Snapper, Snake, Sturgeon,
and Cattish.
The examination of these collections show that the exploration
of small caves may not only prove of ethnographic value, but
serve to give us information relative to the early fauna of the
countr}'. Thirty years ago Prof. Baird gave an account of the
exploration of some bone caves in this State (Proc. Am. Assoc,
II, 1849, 352). He refers to a vast accumulation of remains in
one of the caves, and remarks that the number of species of mam-
malia found is twice that of present existing species in Pennsyl-
vania. It is to be regretted that no further account has yet been
given of the species to which the remains belong.
Dimorphic Flowers in Hoiistonia. — Mr.Tnos. Meehan remarked
that flowers dimorphic in their sexual character were well known.
Generalh' there was little difference in the corolla between the
short styled or long styled flowers, but in Houstonia cceridea^ L.,
the long st^'led form was accompampanied by a thick tube, while
the tul)e in the short styled form was not more than half the
diameter of the other. In this species of Houstonia the anthers
were placed on a ledge which was at the base of the tube in the
long styled form. In the short styled form the anthers were
brought to the mouth of the corolla without any lengthening of
filaments, but by the bringing up of this ledge on which the
anthers are placed. The position of the anthers at the mouth or
at the base of the tube, was in fact decided by the modification of
the form of the corolla tube. This had been explained in the
first series of " Flowers and Ferns of the United States." In a
recent examination of Houstonia serpullifoHa^ Mx., on the top of
Roan Mountain in North Carolina, Mr. Meehan found preciselv
the same characters in that species. On the same mountain
Houstonia purpurea^ L., abounds, and also has a similar sexual
dimorphism, but_in]^this case the elevation of the anthers is due
350 PROCEEDINGS OF THE ACADEMY OF . [1880.
to the lengthening of the stamens and the form of the corolla-tube
is the same in both sexual forms. The number of plants repre-
senting each sexual form is about equal. In a handful of plants
gathered at random there were of Houstonia serpyllifolia thirteen
with long styles and short stamens, and eleven with short styles and
the anthers in the mouth of the tube. Of Houstonia j)urpurea
there were thirteen with long styles to fourteen with short ones.
An interesting fact in connection with these sexual differences is
that the dimorphism seemed to be wholly in the shortening of the
style or filaments, and did not seem to effect injuriously the
anthers or stigma. Both forms seemed to be equally fertile.
In answer to a question by Mr. Redfleld, Mr. Meehan said there
did not appear to be any intermediate forms. The stigma or the
anthers were either exactly at the throat of the flowers, or exactly
at the base. In regai'd to cross fertilization the long styled would
be in the best position for receiving pollen from foreign flowers,
but the short styled one would more readil}" receive its own. As
cross and self-fertilization had an equal advantage he would infer
that the dimorphism had little reference to fertilization as a final
cause.
Gleistogam.y in Oxo.lis Acetosella, L. — Mr. Meehan observed
that under the forests of Abies Frazeri^ on Roan Mountain, North
Carolina, early in August Oxalis Acetosella was abundantly in
flower.
In a large number of cases examined he could find no traces of
any disposition to produce seed vessels, but pushing out beneath
the soil or near the surfece were numbers of cleistogene flowers,
from which, in all probability, seeds would be produced in abun-
dance.
Most plants which produced cleistogene flowers, also produced
the usual open corolla bearing flowers at one time or another in
the season, and it had been suggested that this bright petalled
condition was for the purpose of attracting insects, and thus give
the species some chance to escape from the evils which cleis-
togamy, or close-breeding is supposed to involve. It is worthy
of note that no day insects were noted to visit the flower of the
Oxalis blooming in these dark fir forests, though nocturnal ones
might supply tlie deficiency. Mr. Meehan remarked, however,
that no rule could be deduced from single observations, or obser-
vations repeated in the same localities, as the behavior of
plants and insects varied with circumstances. The Oxalis might
not be cleistogene, might be visited by insects, and the open
flowers mioht be fertile elsewhere. He referred in illustration to
Amphicarpsea monoica^ Nutt., which near Philadelphia sometimes
produced no seeds from the petaliferous flowers, while at other
times these flowers were remarkably fertile. Again, all his exam-
inations in the locality named had resulted in finding that the
1880.] NATURAL SCIENCES OF PIIILADELPiriA. 351
petal-bearing flowers were as perfectly fertilized in the iinexpanded
corollas as were tlie cleistogene flowers. But he was prepared to
expect different results elsewhere.
September 28.
The President, Dr. Ruschenberger, in the chair.
Thirty-four persons present.
The following papers wei'e presented for pul)lication in the
Journal :
" The Parasites of the Termites," liy Jos. Leidy, M. D.
" Remarks on Bathygnathus horealis," l)y Jos. Leidy, M. D,
October 5.
The President, Dr. Ruschenberger, in the chair.
Thirtj'-two persons pi*esent.
The death of James C. Fisher, M. D., a member, was announced.
Sexual Variation in Castanea Americana, Michx. — Isaac C.
Martindale stated he had recently visited Pitman Grove, Glou-
cester Co., New Jersey, in order to examine some chestnut trees
growing there, and remarked that it was well known there are
but two species of chestnut trees in this country, the chinquapin,
Castanea pmnila^ found occasionally in New Jersey, and abund-
antly further south, and the common chestnut, Castanea Ameri-
cana. The chinquapin attains the height here of ten to tweh'e
feet, being a shrub rather than a tree. In Mar3^1and, Virginia and
North Carolina, it often reaches four times that height, and be-
comes nearly a foot in diameter. It is from those States that
most of the chinquapins found in our markets come. The burs
contain but a single nut, and it quite small, but as a cluster of
five or six together is not unfrequent, nearl^^ as many chinquapins
may be found on a branch, as chestnuts on the common chestnut
trees. The leaves of the dwarf chestnut, as it is sometimes called,
are woolly underneath, even in their mature condition, while on
the other they become green on both sides as they reach full size.
A peculiar feature in the flowering of the chestnut, namely, that
the trees bear two sets of flowers, was pointed out b^^ Thomas
Meehan, of the Oardener''s Monthly, a close observer of vegetable
growths, more than a year ago. His observations are recorded in
the Proceedings of the Academy of Natural Sciences of Phila-
delphia, for the year 1879. It may not be difficult for any one to
recall the manner of the infloresence, which consists of the
352 PROCEEDINGS OF THE ACADEMY OF [1880.
flowers growing on a stem, botanically called a spike, from four
to six inches in length, there being from thirty to sixty flowers
together. These come from buds in the axils of the first leaves
of the season, and are composed entirely of staminate (male)
flowers. They are very odorous when in full bloom, and often so
abundant as to give the trees a white appearance when seen at a
distance. As soon as these flowers fade, which is in a few days,
a disarticulation takes place close to the branch, and the spike
falls to the ground. About ten days later, a second flowering
takes place, these spikes coming from the later axillary buds of
the season, and instead of being all staminate as in the first in-
stance, at the base of the spike will be found one, sometimes two,
rarely more, pistillate (female) flowers. These are fertilized by
the staminate flowers that are in blossom at the same time ; the
staminate part 0f the spike falls away after flowering, but the pis-
tillate part remains attached to the branch, and developes into a
bur, containing from two to five or six nuts. What may be the
use of the first set of blossoms, has not yet dawned upon the
mind of man ; it would seem a great waste of energy to provide
for such an abundance without a purpose, but the prodigality of
nature is visible in numerous other instances as well.
The variety of forms of the nut was greater in the locality re-
ferred to than he had ever seen before. One tree was particularly
attractive, the shape of the bur being exactly pyriform instead of
globular ; its chestnuts, of course, corresponding somewhat in
shape, being long and slim.
Near the southern line of the tract was found one tree, and
afterwards in another part a second tree, which will require special
notice. The former was about twenty feet high and six inches in
diameter, while the other was at least seventy-five feet in height,
and more than two feet in diameter at the base, a very wide
spreading and thrifty looking tree. In these, the later blossoms
referred to, instead of Ijeing part staminate and part pistillate,
have been all pistillate, consequently were succeeded by burs all
along the spike, numbering in those counted from flft}- to sixty
together, and hanging from the branches like bunches of grapes.
Every branch of the tree that bore any at all, had them of this
character, so that there were doubtless hundreds if not thousands
of them. An important point is here manifested. These flowers
being all pistillate, and the staminate ones (the first blossoms re-
ferred, to) having fallen, there was nothing to fertilize them, con-
sequently they could not attain much size nor develop chestnuts
within the bur, except that rarely the first or second nearest the
base contained three or four ver^^ small nuts. These nuts, how-
ever, were without germs.
He had been unable to find any record of such an occurrence in
this country before, but Dr. Masters records it as having been
noted in France. The superintendent of the grove to whom be-
longs the credit of first detecting these trees, could not say
rroc. A IN. o. run. luuv
u.
T.Binclaii- &. S oti, liilh •F^.>■la ia
Simia Satyr^us. L.
PROC, A. N. S, PHILA,, 1880
PL, XII
CHAPMAN, ANATOMY OP ORANGOUTANG.
^ROC, A, N. S, PHILA., 1880,
PL, Xlil,
.ANATOMY OF ORANGOUTANG,
Proc A.N. S. Phil -1880.
Plate XIY
i^
W\
,..--^'
Sf§:f
^cjWi^'fliiytfWgtf&^j
w
r-
'^.:
<PUnik>>>
*;
T Sinclair* Sonlith.Philada
Chapman, Anatomy of Ourang Otan^.
PROC. A. N. S, PHILA,. 1880,
PL, XV.
CHAPMAN, ANATOMY OF ORANGOUTANG,
PROC, A, N, S, PHILA., 1880.
PL XVI,
CHAPMAN, ANATOMY OF ORANG OUTANG,
PROC. A, N, S. PHILA,, 1880,
PLXV!
CHAPMAN, ANATOMY OF OPANO OUTANG,
PROC. A, N. S, PHiLA
PL XVIII
BARBECKON LEMNA,
1880.] NATURAL SCIENCES OF PHILADELPHIA. 353
whether in past years they had borne burs in this manner or
not.
It will be remembered that occasionally in a field of corn the
tassel, which is the staminate (innle) flower, has a number of
grains of corn intermixed. These grains come from pistillate
(female) flowers, occurring among the staminate ones; thus it
may be observed that our chestnut tree is not the only instance
of deviation from the regular laws of development. It has been
aroucd that a want of nutrition will aecoiuit for this and similar
instances, but the healthy appearance and vigorous growth of
the trees in question is not such that a lack of nutrition can well
apply.
Mr. Thomas Meehan remarked that he believed instances of
the changes of flowers normally of one sex to the otlier, were oc-
casionally met with, though he could not refer to many without
further thought or investigation, but it occurred to him just then
that it Avas not unusual for some normally male spikes in Carex
to have female dowers among them. He had himself seen well
developed ovariums among the anients of Populus alba, and the
case of female flowers among the male catkins of willows, was
well known to teratologists. Reference had been made to his
papers on sex as influenced by nutrition. His view of sex, as
well known, was that in the earlier stages, between the cessation
of vegetative growth and reproductive growth, a vegetable cell
might be either male or female, and that the power of that cell to
assimilate nutrition, involved the question of sex. If a full sup-
ply was received, the female form resulted; if limited, the male
was produced. In most cases this assimilative power influenced
only the branches or cells in the immediate vicinity of the
flowers. There might be no difference in the cells of the whole
plant in a general way to avail themselves of a full supply of
nutrition. He did not know tiiat there was greater vegetative
strength in the plant of Maize, which bore some females among
the '' tassels '' or males, than there was in the normal plant.
There certainl}^ was no difl'erence in the vegetative strength
of plants of separate sexes in many classes of plants. But
there were instances which proved that the whole individual
plant was influenced by laws of nu.rition when the question
of sex was involved. The female Hemp, the female Spinage,
the female Croton, when the plants were wholly bi-sexual, were
cases he could readily call to mind where vegetative vigor favored
the whole plant.
The common Ambrosia artemisisefolia, which often grows so
thickly over cultivated fields as to a;)pear as a regular farm
crop, each plant fighting for nutrition with its neighbor, pro-
duces almost wholh' male blossoms ; the few females are found
at the base of the male spikes. But when we go to the maize or
the potato fields, where the plants are few and well fed, we may any
24
354 PROCEEDINGS OF THE ACADEMY OF [1880.
time find plants which have a great abundance of female flowers, —
indeed, sometimes plants which are wholly female.
In the case of these chestnuts he would not say it was a want
of nutrition which made these normally male flowers become fe-
male. That was not his view of the case. On the contrary, it
was that better nutritive advantages prevailed to influence the
female sex, and these long spikes of chestnut fruit proved the
fact rather than interposed an objection. It was a simple and
uncontroverted fact that these young chestnuts were being
nourished, were imbibing nutrition, wdiile if they had been nor-
mal male flowers, they Avould have been dead months ago. It
was evident to the senses that nutrition was in the end involved,
and w^e only had to consider at what point of early cell life its
influence was felt. The old idea would probably be that the
question of nutrition followed the " fiat " which made sex, while
his views deduced from the numerous facts he had published on
the question, were th;it nutrition, in its various phases, was itself
the law-maker. As to the greater power behind this, which
decreed that this should be the law, ;nid that the law should pro-
duce such even divisions in the proportion of the sexes, it was
another question. He only claimed that his discoveries had
brought us a step nearer to this greater cause.
Note. — I have since learned through au old resident in the vicinity, that
the large tree has borne such burs for many years, and that it is known
throughout the neighborhood as the "he " tree. — I. C. M.
OCTOBEK, 12.
Tlie President, Dr. RuscHENP.KiKiEU. in tlie cliair,
Thirly-five ])ersons i)rv'sent.
October 19.
Dr. I\. S. Kenderdine in the chair.
Twenty-eiglit persons present.
The Publiciition Committee reported in favor of pul>lishing the
following papers in the Journal of the Academy: —
" The Parasites of the Termites," by Jos. Leidy, M. D.
" Remarks on Bathygnathus orientalis," by Jos. Leidj-, M. D.
1880.] natural sciences of philadelphia. 355
October 26.
The President, Dr. Ruschenberger, in tlie chair.
Nineteen persons present.
The deaths of Dr. Chas. H. Budd and of Joshua Lippincott,
members, were announced.
Samuel R. Kuiglit, M. D., and Rev. Wm. F. C. Morsell were
elected members.
November 2.
The President, Dr. Ruschenberger, in the chair.
Twentj^-four persons present.
Rain Trees. Note on Yucca gloriosa.—'Mv. Thomas Meehan
referred .to a branch of Yucca gloriosa.^ exhibited a few evenings
ago, taken from a plant growing in his garden, and which had
flowered during September, the usual period for blooming near
Philadelphia. Walldng through his garden with Mr. Isaac C.
Martindale, the latter had called his attention to moisture which
covered the whole outer surface of the flowers, and collected in
drops at the drooping apices of each leaf of the perianth. The
plant was within, a few days' of going wholl}' out of bloom, but
during these few daj's the exhibition of moisture continued, and
the appearance of the leaves beneath showed that the dropping of
liquid had been going on for some time, and perhaps during the
whole flowering season. There was no perceptible sweetness in
the liquid, but the presence of ants indicated that it might possi-
bly have a slightly saccharine character, though not sensible to
the human tongue. It was diflicult to decide whether this liquid
was an exudation from the leaves of the perianth or Avas simply
an exercise of the power of condensing moisture in the atmo-
sphere which some plants possessed, notal)ly the Pithecdohium
Saman, Benth., famous as the " Rain-tree " of Peru, which watered
its own roots by the moisture condensed from the atmosphere,
thus en!d)ling the tree to live in almost rainless regions, if the
reports of travelers are to be fidly credited. He hoi)ed to make
further oliscrvatious on the Yucca another year.
November 9.
The President, Dr. Ruschenberger, in the chair.
Twenty-four persons i)resent.
The resijTuation of Mr. Geo. Vaux as a member of Council was
"read and accepted.
356 proceedings op the academy of [1880.
November 16.
The President, Dr. Ruschenberger, in the chair.
Fortj^-two persons present.
The death of Alexander Wilcocks, M. D., a member, was an-
nounced.
The following was unanimously adopted :
Resolved — That the thanks of the Academy of Natural Sciences
of Philadelphia be presented to Mrs. Isaac Hays for Waugh's
excellent portrait of the late President, Isaac Hays, M. D., whose
labors and influence during more than sixty 3'ears contributed
largely to promote the interests of the society.
Dioscism in Andromeda Cateshsei, Walter. — Mr. Thos. Meehan
remarked that in 1861 he had reported to the Academy the
dittjcism of Epigaea^ and he believed this had stood so far the
only case of unisexualit}^ reported in the whole of the large order
Kricacepe. He said he had now to add another in Andromeda
Gatesbsei of Walter, of which he exhibited specimens gathered last
year on the Catawba River, in North Carolina. In the course of
mau}'^ days' journeyings he had the opportunit}' of examining num-
erous plants in many different districts, and they were all either
wholly sterile or wholly fertile in separate plants, as in the speci-
mens exhibited. Occasionally, as often seen in dioecious plants,
a few capsules would be found on the sterile plants, but he could
not say whether the seeds in them were perfect.
Mr. Redfield inquired whether Mr. Meehan had examined the
flowers, and found intermediate stages of development in the
sexual organs ?
Mr. Meehan replied, that the plants were out of flower when
observed ; that it was the abundant fertility in some plants, and
absolute sterility in others that had attracted his attention. From
the remains of the few faded flowers he could And on the plants
the stamens appeared perfect on the staminate plant, with no trace
of pistil or ovarium, while in the fertile plants no trace of stamens
could be found about the remains, though it is probable from
analogy in Epigfiea^ these organs in the fresh flowers would be
found to exist in a rudimentary state.
On Fresh-water Sponges. — Mr. Potts, continuing the subject of
American forms of fresh-water sponges treated of some weeks ago,
said that the number of species noticed during the few months in
which they had claimed his attention gave some reason to believe
that the Order Spongida has many more representatives in our
fresh waters than has been generally supposed.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 357
On a former occasion he had described three species of Sp07ig-
illa from a small stream near Philadelphia, one of which, then
named S. tentaaperma, but which he now preferred to call S.
tenosperma, exhibited features so exceptional as almost to claim
for it generic distinction.
He had since found the S. frag His of Leidy plentifuU}' in the
Schuylkill river below the dam, ( Leidy 's original localit}'), and
above the dam a lacustrine form differing from that before alluded
to. A very slender green species creeping along stems of sphag-
num, etc., had been received from a swamp near Absecom, N. J.
As it appeared to be entirelj' withoiit spined spicule of either
class, he proposed for it the name S. aspinosa.
From the Adirondack lakes a beautiful species, believed to be
identical with S. stag)ialis, Dawson, had been received through
the kindness of Prof. H. Allen. Another lacustrine form which
yet is not ([uite S. lacustris, was brought from the lake near Cats-
kill Mountain House by Professors Cope and Hunt. Its status
has not been fully determined.
From the cellar of an okl ruin at Lehigh Gap, Pennsylvania, he
had obtained four species, all of which appeared to be new. These
were all thin, creeping or encrusting sponges, three of them of
the birotulate tj'pe, briefly described as follows :
S, argyrosperma — seed body or sphreruhv, large, silver-white,
densel}^ covered with radial spicule, the shafts of which are long,
stout, with numerous long spines, straight or curved ; the rotulae
at each end being replaced by 1-4 strong recurved hooks.
S. repens — found creeping over the stems and leaves of Pota-
mogeton ; sphjerula^ also closely covered with spicuLne, shorter and
more slender than those of the preceding species ; their shafts
nearly smooth, the ra3^s of the rotulfe, six, eight or more, uniformly
incurved like the ribs of an umbrella.
S. astrosperma — the sphairuhv haA'e the appearance of being much
smaller than in either of the former species, which is probably due
to the fact that the birotulate spicule surrounding the real cap^
sules are very short ; the length of the shaft being less than the
diameter of the rays. Thej' are rather sparsely scattered over
the surface of the nearl}' transparent sphere, suggesting the name
star-seeded.
The remaining form is considered a variety of 8. fragilis, and
called minuta ; sph.erula' much smaller than in the type species,
the dermal and superincumbent spicule terminated by sharp points,
while in the other they are universally truncate or rounded.
A more particular description with measurements, etc., is in-
tended.
Mr. Ezra T. Cresson was elected a member of Council, to fill
the vacancy caused by the resignation of Mr. Geo. Vaux.
358 proceedings of the academy of [1880.
November 23.
The President, Dr. Rusciienberger, in the chair.
Thirty members present.
November 30.
The President, Dr. Ruschenberger, in the chair.
Thirty-six persons present.
Note on the Seed-vessels of Wistaria — Mr. Thomas Meehan
remarked tliat most persons knew that special temperatures were
required to insure the germination of various seeds. The com-
mon chickweed germinated at a little above freezing point, while
one of TO'-' was required by most palms. Heat and moisture had
also a varying influence on the openijig of seed-vessels, some
requiring more or less than othei-s. He exhibited some seed-
vessels of Wistaria si7ie7isis and Wistaria fri(tescens, to illustrate
the point. A box, four inches deep, with some seed-vessels of
Chinese Wistaria was placed on a shelf in a cool room. A fire hap-
pened to be made in the room and kept up all night, and the next
morning the capsules had burst, and scattered the seeds and open
vessels about the room. So great was the force of the expansion
that some seeds were projected ten feet from the box. One large
seed-vessel had been lifted before opening by one or more beneath,
over the four-inch side of the box, and had fallen on the ground
at least two feet away from the box in a horizontal line. The few
that had been thrown on to the floor by the explosion of their
companions did not open, owing to the difference in the tempera-
ture of the floor from that of the shelf. Five seed-vessels of each of
the two species were then placed together on the shelf, where the
temperature of the atmosphere was about 45°. After four days
they were examined. The American species had all opened, but
without expelling the seeds, which were still attached to the car-
pel; but those of the Chinese Wistaria were still unopened. The
Chinese Wistaria required a much higher temperature to open the
capsules than the American, though it might be that hygrometri-
cal conditions would vary the exact degree required.
Mr. Martindale observed that the seed-vessels of the Chinese
Wistaria were much more indurated and rigid than the American
species, and required more force to open them. He had noted
that such hard seed-vessels always exerted a greater projectile
power when opening.
C. S. Turnbull, M. D., and J. M. Anders, M. D., were elected
members.
1880.j natural sciences op philadelphia. 359
December T.
The President, Dr. Ruschenberger, in tlie cliair.
Twenty-three persons present.
December 14.
The President, Dr. Ruschenberger, in the chair.
Thirt\-two persons present.
A paper entitled " On some Lower Eocene Mollusca from Clarke
Co., Alabama, with some points as to the stratigraphical position
of the beds containing them," by Angelo Heilprin. was presented
for pnblication.
The Phalanges of Bats. — Dr. Allen, in reviewing the manner
after whicli tlie phalanges in mammalia are enumerated, spoke of
the propriety of inelmling the terminal eart'laginous tip to the
fingers, present in many bits, in the series of phalanges.
Authors do not hesitate in namino- the terminal cartilaue to the
second finger in Rhinopoma a phalanx, nor should they, Dr. Allen
held, hesitate in so including the terminal segments in other genera.
It is interesting to observe that in 3Iolossus perotis the terminal
joint in tlie second finger is bony, and anchylosed to the first
phalanx. If this plan of numbering the phalanges in bats be
accepted, from one to three joints are present in all the fingers.
The position taken by recent writers that the Phyllostomidte are
distinguished from other families by the presence of the third
phalanx to the third finger cannot be sustained, since this phalanx
can be counted in other families, the terminal joint, however, re-
maining in them cartilaginous.
December 21.
The President, Dr. Ruschenberger, in the chair.
Ten persons present.
Note on a new Northern Gutting Ant. Atta sep!entrionalis. —
Dr. McCooK remarked that he had the pleasure of announcing
an interesting discovery of a species of cutting ant upon the
eastern central coast of the State of New Jersey. The dis-
covery was made by Rev. George K. Morris at a new watering
place called Island Heights, which is located upon a swelling
bluft' on the northern bank of Tom's River, near its mouth,
three miles from the Atlantic Ocean, in about Lat. 40° N.
360
PROCEEDINGS OF THE ACADEMY OF
[1880.
Mr. Monis, who has been much interested in
noting
the habits
of ants, observed this species carrying the needle-like leaves
of the pine into their nests, and thereupon followed tlieir
behavior until he found it to be quite like that of the cutting
ant of Texas, Atia ferveus. Dr. McCook having been informed
of the above discover^-, made a journey to Island Heights in
the early part of September, 1^80. Unfortunately a severe eastern
storm set in before the train reached Tom's River, and con-
tinued during his stay with such rigor as utterly to preclude
observation of the out-door behavior of the ants. However,
b}^ woiking in the storm, protected by rubber garments and a
temporary shelter, he was able to make a study of the internal
architecture of a nest.
The opening from the surface appeared to be a single narrow
tubular galler}', X, of about two inches in length, which pene-
trated the ground at an angle of near 45°, and entered a spherical
chamber, V — a sort of vestibule — about Ij inches in diameter.
Within this a few ants were found, nothing more.
This vestibule communicated hy a short gallery, Y, with a
second chamber or cell, 0,
having generally a spheri-
cal shape, but more irregu-
lar in outline than the ves-
tibule. It was about 3 in.
in diameter. Within this
were several small masses
of an ashen-gray, fibrous
pulp or papery material,
closely resembling that
found b}' him in the large
cells or caves of the Texas
cutting ant.^ This was
evidently the leaf-paper
formed by the manduca-
tion of the pine leaves. It
was exceedingly fragile,
even more so than the
leaf-paper of the Texas
Atta,and could not be kept
together in the original
mass for examination. It
appeared, however, to be
without the decided cellu-
lar arrangement first ob-
served b}^ him in the leaf-
paper of the Texas ant,
whose '• combs " — the analogue of those of other hymenopters, as
1 Proc. Acad. Nat. Sci. Phila., 1879, p. 37.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 361
the bee and wasp — were composed of irregular hexagonal cells of
various sizes.
None of these leaf-paper masses exceeded an inch in height ;
they lay upon the floor of the cave, C, or were upheld by the fila-
mentous rootlets which penetrated within the liollow, or, more
properly speaking, around which the cave had been formed. In
this respect also the habit of the northern ant resembles that of
the southern. Although, as said, the out-door beliavior of the ant
could not be observed Ijy Mr. McCook, the insects iiaving all been
driven in-doors by the storm, these were observed to some extent
by Mr. Morris, the discoverer, during the summer. Mrs. Mary
TVeat also visited the Heiglits and noted the same. From verbal
reports received from these persons the analogy between the two
insects, in respect of gathering and tiansporting leaves, appears
to be quite close. There are many i)oints, however, which remain
to be determined accurately, which it is to be hoped will be done
next summer.
The following account was communicated to the speaker by Mr.
Morris, and is incorporated, by his permission, in this note : —
" In answer to your questions I would say, when first observed,
there were two columns, one going each way, and moving ver3'
deliberately. If alarmed by any rudeness on my part, they sought
safety by remaining perfectly motionless for some time, making-
it difficult to find them, as they are nearly the color of the dry
leaves.
" Those in the column going homeward were carrying little pieces
of the pine needle or leaf, cut from seedling plants about 2 inches
high and upwards. In some instances the piece of leaf was not
as lono; as the ant itself, but in others it was longer than the
bearer. The appearance presented by the cohunn was very singu-
lar, for instead of carrying their burdens as other ants do, so far
as I have observed, they bore the load on the head, resting in a
saddle-like, V-shaped space between ridges on the head, running
from the base of the mandible on each side to tne top of the liead.
One end was held firmly by the mandibles. T'he eftect at a little
distance was to give them a ' shoulder arms ' appearance.
" Tracing the column back, I readily found tlieir foraging ground
a few feet from the formicary. There were the remains of several
seedling pines which had been strip])ed and cut doAvn nearly to
the ground. Some ants were at work on the only one left stand-
ing, and I enjoj'cd the pleasure of witnessing their operations.
There la}' on the ground a few pieces, which were picked up, as I
watched, and carried awa}'. On the plant there were two at work
cutting. Climbing out on a leaf to a position near tlie end, the
ant applied her mandibles, and moved around as she cut, until the
piece was severed, when she repeated the process, in most cases
allowing the severed pieces to fall. One cutter held on to her last
362 PROCEEDINGS OF THE ACADEMY OF [1880.
cutting, backed down with it, and started off home. In no in-
stance have I ever found one of these ants carrying a load of any
kind in an^' other way than as described above. Your account of
tlie Texas ant would answer as well for this.
" The little leaf-cutters at Island Heights have no mounds at the
entrance to their formicaries. I found many nests, but none
with a mound. The sand pellets in every case where I found ants
at work were carried off several inches distant on every side and
scattered, as if for secrecy. The ants' movements were exceed-
ingly deliberate, always. Some seemed overloaded and rested
often, but never laid down their load.
" A.11 the colonies were comparatively small. I was imable tO'
find any indication that the}^ were connected one with another.
•' There is not, so far as my observations go, any elaborate open-
ing and closing of gates, such as you describe, but in wet weather
I usually found a leaf over the entrance to their nest.
" The architecture of their caves is a miniature copy of that of
3'our Texas cutting ant.
" I found them cutting only the pine leaf and the leaf of a small
shrub called Cow Wheat (Melantpi/rinn omerU aiiuin). Of this
plant the^' took the petals also. They carried the dry curled leaf
as well as the green and freshly cut.
" They also carry and incorporate into the nest material the drop-
pings of certain larva> that feed on oak leaves."
Mr. Morris observed at the same place a second and larger form
of leaf-cutting ant, which he thinks to be a distinct species ; but.
as he was not able to point out any of the nests to Mr. McCook
and has not yet made thorough examination, this point remains,
to be solved.
The ant whose economy is described above closel}"^ resembles
the famous Aftafei^cens of our southern regions, having the same
leaf-brown color and the same characteristic spines. It is distin-
guished by a black longitudinal baud along the median dorsal
part of the abdomen, and by a similar band along the middle
part of the fjxce, marking the furrow formed by two ridges, the
prolongation (apparently) of the antennal ridges to the vertex of
the caput. A double row of spines extends along the entire tho-
rax and nodes, expanding at the prothorax into about four. Two-
castes of workers were found, appearing to be the workers major
and minor, in length respectively 4 and 3 millimetres, or about
one-sixth and one-eighth of an inch. The species appears to be
new, and Dr. McCook at least ventured conditionally to name it
the Northern Cutting Ant — Atta septentrionalis.
The discover}^ at so northern a point of this species, with habits
quite identical with those of tropical congeners, seemed to the
speaker to be a remarkably interesting fact in the distribution of our
ant fauna. He was at once impressed by the striking contrast be-
tween the vast myriads of workers, the extensive excavations,
and the formidable and vigorous activities of the Texas colonies^
1880.] NATURAL SCIENCES OF PHILADELPHIA. 363^
and the small numbers, slight excavations and apparently slug-
gish movements of their northern allies. And he could not for-
bear the thought that these New Jersey communities of Atta
septentj'ionalis seemed like the feeble remnant of a vigorous race
left or thrust by some untowai'd change upon unfavorable sites,
which must work toward their extinction.
December 28.
The President, Dr. Ruschenberger, in the chair.
Eighty persons present.
The following papers were presented for publication : —
•364 PROCEEDINGS OF THE ACADEMY OF [1880.
ON SOME NEW LOWER EOCENE MOLLUSCA FROM CLARKE CO. ALABAMA,
WITH SOME POINTS AS TO THE STRilTIGRAPHICAL POSITION
OF THE BEDS CONTAINING THEM.
BY ANGELO HEILPRIN.
The following species of fossil mollusca, for which I am indebted
to Dr. Eugene A. Smith. State Geologist of Alabama, were ob-
tained from sections exposed in that State on Knight's Branch
and Cave Branch, tributaries of Bashia Creek (Clarke Co.), and
from Wood's Bluff on the Tombigbee River, near the mouth of
Bashia Creek, and some twenty-eight miles north of St. Stephen's.
They occur in probably the oldest marine tertiary deposits of the
State, and occupy a horizon nearly parallel with that which is
characterized by the fossils of Upper Marlborough and Piscata-
w^ay River, Marjdand, and Pamunkey RiA'er, Virginia. The follow-
ing enumeration of fossils from the three localities first named,
will best illustrate the pala^ontological relations of the beds
containing them toward each other, and to the various Eocene
deposits of the Atlantic and Gulf slopes :
Fossils from Knight's Branch.
Astarte tellinoides, Conr. ( Va?'. A. sulcata, Lea.)
Cytherea, perovata, Conr.
Cytherea NnttaUiopsis, Heilpr. sp nov.
?• Cardita alticoda {Blandinyi)^ Conr.
Gorbula rugosa, Lam.
(C. oniscus, Conr; va?: C. gihbosa^ Lea.)
Ancillaria (Ancillopsis) subglobosa, Conr.
Natica settles, Conr.
TurbineMa {Ga^Hcella) Bandoni, Deshayes, sp.
{Valuta Bandoni, Desh.)
Lsevibuccinum lineafum, Heilpr. sp. nov.
Rostellaria (Calyptrophorus) trinodifera, Conr.
Solarium cupjola, Heilpr. sp. nov.
Fusus inter striaius, Heilpr. sp. nov.
Fusus sub-tennis, Heilpr. sp. nov.
Fusus (Strepsidura) subscalarinus, Heilpr. sp. nov.
Tornatella {TornatelWa) bella, Conr.
Ostrea.
Cylicosmilia. i
1880.] NATURAL SCIENCES OF PHILADELPHIA. SQb'
Fossils from Cave Branch.
Dentalium micro-dria , Heilpr. sp. nov.
Natica fetifes, Conr.
Natica Mississippien sis^ Conr.
Pyrula muUangulata, Heilpr. sp. nov.
Pyrula iricoxtata. Desh.
TurriteUa carinata, Lea.
Solarium cupola^ Heilpr. sp. nov.
? Pleurotoma acuminata^ Sowerby.
Pleurotoma moniliala, Heilpr. sp. nov.
Ga^ddaria (fragment). Closely allied to C carinata, Lam.
Voluta (Afhleta) Tuomeyi. Conr.
Fusus pagodiformis^ Heilpr.
Fusus interstriafui^^ Heilpr. sp. nov.
Fusus subfenuis. Heilpr. sp. nov.
Fusics iStrepstdura) suhscalariuus, Heilpr. sp. nov.
Leda protexfa, Conr.
Cardium {Protocardia). Young of G. McoUetif Conr.
Ostrea (same species as from Knight's Bi'anch).
Fossils from Wood's Blurt".
Dentalium micro-stria, Heilpr. sp. nov.
Natica limnla, Conr.
Pyrula midtangulata. Heilpr. sp. nov.
TurriteUa carinata, Lea.
Solarium cupola^ Heilpr. sp. nov.
Solarium deljjhinuloides, Heilpr. sp. nov.
Gancellaria evulsa, Brauder, sp.
( G. tortiplica ? Conr.)
Pleurotoma ( Gochlespira ) cristata^ Conr.
Pleurotoma^ n. sp.
AnciUaria (Ancillopsis') suhglobosa^ Conr.
Pseudoliva vetusta, Conr.
Pseudoliva scalina, Heilpr. sp. nov.
Voluta (Athleta) Tuomeyi^ Conr.
Fusus pagodiformis, Heilpr.
? Fusus (Levifusus) trabeatus^Coui:
Fusus inter st rial us ^ Heilpr. sp. nov.
Fusus. n. sp.
? Gardita alticosta {Blanditigi), Conr.
. Leda protexfa, Conr.
366 PROCEEDINGS OF THE ACADEMY OF [1880.
Pecten Foulsoni, Morton.
Ostrea (species difterent from that of Knight's Branch and
Cave Branch.)
From an examination of the above tables it will be seen that a
fair proi)ortion of the fossils from Knight's and Cave Branches
are held in common by both deposits, and therefore there can be
no reasonable doubt that they represent about equivalent horizons.
Of the hitherto undescribed forms Gytherea Nuttalliojysia and Lsevi-
huccinum lineatuvi appear to have been obtained only at the
former, and Pleurotoma moniliata at the latter locality, although
it is highly probable that further investigation will reveal their
mutual presence in both localities. The described American forms
are mainly those occurring at various heights on the Claiborne
exposure. A comparison of these forms with those obtained by
Tuomey ( First Biennial Report of the Geology of Alabama, p.
146) from the Basliia Creek sections near Choctaw Corner, shows
the two groups to be of a contemporaneous age, for from bed No.
2 of that section Prof Tuomej^ obtained (among others) species
of " Ostrea., Gytherea, Cardita^ Cardiuin, BosteUaria, Actseon.
Vohita, Infundibulum, and' Solarium,^'' wliich appear to have been
identical with the species obtained by Dr. Smith from the two
localities above mentioned.'
1 The siiecies enumerated by Tuomey are Ostrea coinpressirostra, Cardita
planicostd, Rostelhtrla vcluta, ArtiEon po)/nHii.s, Vol nUi Say ana? Cardium
Nicollctl, undinfundibulfuii trochiforinis. The specimens aj^pear to have been
submitted to Mr. Conrad, who considered the determinations of Tuomey as,
at least in part, imperfect, and substituted the following specific names {Am,
JoxLrri. Science, new series, xl, p. 266) : Ostrea CaroUnensis (species from
the Santee Canal, South Carolina), Volutilithes [Athleta] Tuomeyi (de-
scribed by Conrad [Proc. Acad. Nat. Sciences, vi, p. 449] in 185iJ from
Bashia C!reek), and Protoairdiu Virginiana ? The following remark in
pencil occurs in the volume of Toumey's Reports, contained in the library
of the Academy : "All doubtful except VenericAirdia planicosta. T. A.
Conrad." Tuomey's Rostellaria tetata and Acta'on pomilivs were in all
■pvdhahiWtY Ixostellaria trinodif era and Tornntella, bella, wliich would better
agree with the descriptions of obviously the same fossils as given in Hale's
report (C. S. Hale : The Geology of South Alabama, Am. Journ, Science,
new series, vi, p. 355).
1880.]
NATURAL SCIENCES OF PHILADELPHIA.
SECTION ON BASHIA CREEK.
367
1
2
3
4
5
6
Hard Limestone.
4 feet.
Marl, highly fossiliferous.
35 feet.
Blue Sand.
Variable.
Lignite and Clay.
6 feet.
Laminated Clay, Sand and Mud.
Thickness undetermined.
Lignite.
Thickness undetermined.
(Tuomey : First Biennial Report, p. 145.)
Note.— Beds 5 and 6 do not properly belong to the section, but "repre-
sent beds seen on another part of the stream below the preceding." (Loc.
cit. p. 146.)
The fossils from Wood's Bluff, some 15 miles W. of Choctaw
Corner, were obtained by Dr. Smith from a bed of indurated
jrreen sand rising about 10-15 feet above water line, which bed may
possibly represent the lowermost portion of bed No. 2 of the
Bashia- section. Some support is given to this view by the cir-
cumstance that at this point— Wood's Bluff — the basal lignite
(which in the above named section has a thickness of G feet) has
disappeared, and more especiall}' (at least, as showing it to possess
a distinctive character) by the general fades of the representative
molluscous fauna. Although there exists a close similarity be-
tween the general assemblage of its fossils and those of the two
*' Branches " of Bashia Creek, yet the numljer of peculiar forms is
considerably greater, and consequently the aggregate i)ossesses a
much more decided iudividualitj' than obtains with either of the de-
posits in question. Moreover, I am informed b}' Dr. Smith that
the fossil fauna of Knight's and Cave Branches corresponds most
closely with that of btid No. 4 ' of the Wood's Bluff section, an
aluminous deposit about :21-26 feet above water le^-el, and con-
taining species of Denfalium, TornateUa^ Solarium, Turritella,
and Boi^lellaria identical with forms from the two first named
localities. The disappearance of tlie basal lignites at Wood's
^ Section as yet unpublished, but communicated by letter to the author.
868 PROCEEDINGS OF THE ACADEMY OF [1880.
Bluff may be accounted for on the supposition that they have
dipped under, whicii would be in harmony with what we know
concerning the dip of the beds in this region. This is but locally
or at best, but partially indicated in Tuomey's reports, but judg-
ing from the contour lines of the cretaceous formation on the
general maps appended to the first and second Reports, and from
the north and south sections on the map of 1849, as well as from
the facts obtained in Mississippi, it must be in a direction west of
the southern line, or in other words, S. by W. Dr. Smith has
found the loss by dip in a southerly direction on the Tombigbee
River to be about 10 feet to the mile, which accords well with .
Hilgard's observations on the Upper Eocene and Oligocene forma-
tions of Mississippi.^
From pala?ontological evidence alone the three exposures in
question might readily be taken to represent rather an Upper than
a Lower Eocene horizon, for in addition to the species typical of
the American Middle Eocene, or Claiborne group proper ( Calca-
reous Claiborne of Hilgard), and to the new or undescribed forms,
we have the following which have not been hitherto recognized as
belonging to the formation, and which, on the contrary, were
originally described (at least the majority of them) from deposits
of newer date.
Caricella (Voluta) Bandoni, Deshaye?, sp. {Aniiiian.ic s. Vert'br., lidnKin de P<iiix,
II, PI. 1(12, tigs. 13 aud 14), frDiu the " caloaire grossier," Middle Eocene of most
geologists, Upper Eocene of Judd. Knighi's Branch.
Natica Mississippiensis, Cuurad (J. A. X. S. 2d series, i, p. 1 14), originally described
from the \'icksburg (<)ligocene) group, but also foind in the .Jackson ( Upper
Eocene) deposits. Cave Branch.
Pleurotoma acuminata, Sowc by (Mineral Conchology, Vol. ii, p. 105), from the
London Clay of Highgate (Lower Eocene of most geologists, Middle Eocene of
.ludd), and Barton clay (Upper Eocene)? (I have bad no specimens of this species
with which to institute direct comparisons, but from a careful examination of
Sowerby's and E<lvvards' figures and descriptions there appear to me to be no
justifiable grounds for se])arating the species figured on PI. 2(1, tig. 10, from its
European ally. Cave Branch.
Pyrula tricostata, Doshayes (('<><iiiiUm Fosxil «, ii, p. 584), from Retheuil and Uuise-
Laraothe, Middle Eccene (Suessonian of d'Orbigny). Cave Branch.
^ Hilgard fovind the dip of the Jackson and Vicksburg strata to be from
10 to 13 feet per mile S. by W., at "points where the great regularity of
succession for a considerable distance seemed to indicate a normal config-
uration." (A. J. Science, new series, xliii, p. 36.)
1880.] NATURAL SCIENCES OF PHILADELPmA. 369
Pecten Poulsoni, Morton (Synopsis Org. Rem. Cret. Group, p. 59), a companion of
Orbitoiden Mant-Ui, Mort. sp., and, according to Ililgard, an enseutialli/ Vicks-
burg (Oligoeene) fossil. Wood's Bhiff.
Cancellaria evulsa, Sowerby [rjuccinum evulaum, Brander] (Miner. Conchol., iv, p.
84), from tlie Barton clay (Upper Eocene) of England, and Grignon ("Calcaira
grossier") of France. i Wood's Bluff.
Pleurotoma (Cochlespira) cristata, Conr. (J. A. N. S., 2d ser. i, p. 115), originally
described from the Vicksburg group, but doubtful whether differing from the
Pleurotoma bella, Conr., from the Upper Eocene of Texas. Wood's Bluff.
In addition to the above, tliere is among the fossils from Wood's
Bluff an immature Gardium (Protocardia)^ which may possibly
represent the young of C. Nicolleti (Jackson group), with which
it agrees in outline and general ornamentation, or that of G. Vir-
giniana, Conr. (Pamunkey River), an undescribed species, but of
vrhich a labeled specimen is in the collections of the Academy.
The absence of asperulations on the posterior slope of the speci-
men in question, however, rendering it uncertain whether they
were ever present, or whether they are merely abraded or water-
worn, allows of no absolute specific determination.
Whatever ma}' be the palseontological facies of the deposits in
question, however, there can be no reasonable doubt as to their
true position, since Dr. Smith, as he informs me, has traced bed
No. 6 (or the uppermost bed immediately underlying the stratified
drift) of his Wood's Bluff section to the mouth of Witch Creek,
about 2 miles below on the Tombigbee River, where its relation to
the overlying " Buhrstone " is made manifest in an exposure just
beyond the mouth of the creek. White Bluff, about 250 to 275
feet in height, beautifully exhibits the white siliceous clay stones
and silicified shells so characteristic of the southern Buhrstone
formation. These occupy the uppermost portion of the l)luti', and
make up fully 100 feet of its vertical height; the intermediate
portion extending to the water's level, is mainly composed of
laminated lignitic clays, with occasional intercalated beds of pure
lignite. It becomes manifest from what has just been stated that
the fossiliferous beds of Wood's Bluff {et conseq. the equivalent
deposits on Knight's and Cave Branches and Bashia Creek) must
be between 150 and 200 feet below the base of the Buhrstone
' A very closely allied species, the Tritoniuvi{!) jiaucivaricatum of Gabb,
occurs in the Tejon group (Upper Cretaceous — Eocene?) of California,
associated with C'nrdita planicosta. and other characteristic forms of Ter-
tiary fossils.
25
310 PROCEEDINGS OF THE ACADEMY OF [1880.
(Siliceous Clairborne of Hilgard) tbnnation, or what has hitherto
been considered as the base of the Eocene formation in South
Carolina. Allowing a uniform southerly dip of 10 feet to the
mile, these same beds must be about 250 to 280 feet below the
"bed of green sand" mentioned by Tuomey (1st Biennial Report,
p. 148) as occurring at Baker's Bluff, a few miles above St.
Stephens, (stated to be " rich in organic remains, identical with
the fossils of Claiborne ") and which, immediately above St.
Stephens (Tuomey, Joe cit., p. 149), dips beneath the water-line.
This approximate determination of position agrees closely with
the observations made in the northeastern portion of the county,
for Dr. Smith found by actual barometric measurements that the
"chalk hills" (Buhrstone) near Lower Peach Tree on the Ala-
bama River, and at a locality about 1 to 8 miles south of Choctaw
Corner, were about 250 feet above Knight's and Cave Branches,
and the marl bed (No. 2) of Tuomey's Bashia section.
Whether these older Eocene deposits underly the blufl' at Clai-
borne has not yet been proved, but it is but fair to presume that
they do. Likewise, it remains to be shown what relation the basal
lignite on Bashia Creek bears to the "Northern Lignite" of
Hilgard.
CYTHEREA, Lam.
Cytherea Nuttalliopsis, n. sp. PI. 20, fig. 1.
Shell sub-elliptical, moderately ventricose, its surface covered
with fine concentric striae, which are apt to become roughly im-
bricate on the basal margin ; umbones not very prominent, rather
anterior ; lunule cordate, deeply impressed at about its middle, its
outline clearly pronounced by a sharply impressed line ; posterior
extremity regularly rounded, the anterior somewhat produced ;
margin entire ; pallial sinus somewhat angular, pointing toward
the centre of the shell.
Ijength, 1^ inch. Knight's Branch, Clarke Co., Ala.
This species most resembles among American species of Cy-
therea the C. NuttalU, Conr., from which it may be distinguished
by the greater production forward of the anterior extremity, and
by the median depression in the lunule. In this last character it
agrees with G. Poulsoni, Conr., from which, however, it very
materially differs in form, and in the much lesser development of
the umbones.
1880. J NATURAL SCIENCES OF PHILADELPHIA. 371
PSEUDOLIVA, Swainson.
Pseudoliva scalina, n. >■]>. PI. 20, lig 12.
Shell l)iiccinifonn, of about seven volutions ; the whorls rouglily
l)licated ; the folds on the bod}' whorl apjiearing as shoulder no-
dules; dentiferous sulcus well pronounced, followed by about five
impressed revolving lines, which slightlj' crenulate the margin of
the outer lip ; revolving lines on the Ijodj'-whorl above the sulcus
almost obsolete ; aperture slightly exceeding the spire in length ;
columella callous ; suture deeply channeled.
Length, H inch. Wood's Blutf, Clarke Co., Ala.
L.EVIBUCCINUM, Ouna .
(Amer. Jour. Couchol., i, p. 21. Genus uot characterized.)
Shell having the general form of Metula, H. & A. Adams, but
destitute of all traces of a jiosterior canal ; aperture between bucci-
niform and fusiform, about the length of the spire. This, genus
is distinct from Bucciy^anops of d'Orbigny, under which the Buc-
cinnm (Laevibuccinum) prorsum^ Conr., is erroneously classed in
the Pi'odrome de Paleontoloc/ie , ii, p. 369.
Laevibuccinum lineatum, n. s] . PI. 2i', fig. 5.
Shell fusiform, of about seven convex volutions, which are
throughout their whole extent covered by fine, but distinct, re-
volving lines ; aperture slightl}^ exceeding the spire in length,
sub-canaliculate anteriorly; columella gently arcuate; outer lip
striate within.
Length, 1 inch. Knight's Branch, Clarke Co., Ala.
This species mainly ditters from the L. prorsum, Conr., in
having the revolving lines equally distinct over the entire surface
of the whorls. The Murex {Fusus et Buccinum auct.) mitree-
f or mis of Brocchi, from the Oligocene and Miocene deposits of
France, Austria, and Ital}', is a closely related species.
FUSUS, Lamarck.
Fusus subtenuis, n. .'^p. PI. 20, fig. 4.
Shell fusiform, of about seven sub-angular volutions ; whorls
ornamented with somewhat obscure longitudinal folds, about twelve
on the body-whorl, which are cut by several prominent revolvinof
ridges commencing at the shoulder angulation ; shoulder of the
whorls more or less smooth, with an obscure median revolving line,
and a prominent sub-sutural one ; aperture about the length of the
372 PROCEEDINGS OF THE ACADEMY OF [1880.
spire, or slightlj- exceeding it, tlie canal gently curved, moderately
contracted, and somewhat expanding at the extremity ; outer lip
thin, and showing internally the external ornamentation ; base with
numerous revolving lines, which alternate in coarseness.
Length, 1^ inch. Kniglit's Branch ; Cave Branch, Clarke Co.,
Alabama.
Fusus interstriatus, n. sp. PI. 20, fig. 11.
Shell fusiform, slender, composed of about ten convex volutions,
the first three of which are smooth ; whorls ornamented with both
longitudinal plications and revolving lines, the last of which (about
eight in the upper whorls) alternate with finer intermediate stride ;
the longitudinal plications distinct on the earlier whorls, but
becoming much less so on the body-whorl, and the one preceding ;
aperture about the length of spire ; the canal somewhat tortuous ;
outer lip thin, dentate within.
Length, 2 inches. Knight's Branch ; Cave 'Branch, Clarke Co.,
Alabama.
Subgenus HEMIFUSUS?
Fusus (Hemifusus T) engonatus, n. sp. PI. 20, lig. 8.
Shell turreted, of about ten volutions, the first three whorls
smooth and convex, the remainder strongly carinated, and tra-
versed by numerous fine revolving lines, which on the median
portion of the body -whorl alternate with intermediate finer stride ;
body-whorl impressed immediately below the carination (shoulder
angulation) ; lines of growth sinuous, and approximating the
characteristic lines of the Pleurotomidae ; aperture considerably
exceeding the spire in length; columella slightly arcuate, and
presenting a rudimentary fold at a))out its central portion.
Length, 1^ inch. Wood's Bluff; Clarke Co., Ala.
This species resembles the Fusus bifascialus of Deshayes
{Animaux sans Vertebres, Bassin de Paris, II, pi. 84, figs. 15 and
16) from the Paris basin, but may be readily distinguished from
that species by its more slender form.
Subgenus STREPSIDURA, Swaiuson.
Fusus (Strepsidura) subscalarinus, n. sp. Pi. 20, fig. 7.
Shell somewhat bucciniform, whorls about eight, sub-angular,
the first three or four smooth, the remainder ornamented with
both longitudinal costae and revolving striae, the latter showing a
1880.] NATURAL SCIENCES OF PHILADELPHIA. 373
tendenc\^ to alternate in size ; the costte are arcuate, not in a
regular continuous series, those on the body-whorl extending
considerably below the middle of the whorl ; aperture about the
length of spire, the canal somewhat reflected ; columella covered
with a callous deposit, considerably twisted ; outer lip dentate
within.
Length. 1 inch. Knight's Branch; Cave Branch, Clarke Co.,
Alabama.
This species greatly resembles the Fusus scalarinus of Deshayes
(Goquilles Fossiles, II, p. 574, PI. LXXIII, figs. 21 and 2S), but
may be distinguished by the lesser prominence of its costse, and
b}^ the presence of well defined striae over the entire surface of
the whorls. In this last respect, as well as in the subangulated
form of the whorls, it also differs from the Fusus acalariformis,
Nyst (CoquiUes et Poh/piers Fossiks, p. 504, PI. XL, figs. 5a, 6),
from Lethen, Belgium.
TURBINELLA, Lamarck.
Subgenus CARICELLA, r^onrad,
Turbinella (Caricelia) Bandoni, Desha}'ps, sp. PI. 20, fig. 15.
The large species of Caricelia from Knight's Branch agrees sa
closely with the figures of Volata B a ndoni^ J) esh. (Animaux sans
Vertebres, Bassin de Paris, II, pi. 102, figs. 13 and 14), from the
Paris basin, that I do not feel justified in considering it a distinct
species. The American form ajipears to have been somewhat
more elevated, but this is probabl}^ no more than a varietal cir-
cumstance.
Length, 4 inclies. Knight's Branch, Clarke Co., Ala.
PLEUROTOMA.
Pleurotoma moniliata, n. sp. PI. 20, fig. 9.
Shell fusiform, elevated, of about eight volutions, the whorls
considerably contracted above the shoulder ; whorls ornamented
with a double series of nodes, the lower much the most strongly
developed, which giA^es to the upper portion of the spire a monili-
form appearance ; surface of entire shell traversed by fine revolving
lines, which become more distant, very prominent, and alternate
on the median portion of the body-whorl ; aperture about the
length of spire ; the relative position of the upper and lower nodes
corresponds to the sinuous lines of growth.
Length, 1 inch. Cave Branch, Clarke Co., Ala.
374 PROCEEDINGS OF THE ACADEMY OF [1880.
Pleurotoma acuminata T Powerby. PI. 20, fig. 10.
(Minei-al Conchology, II, p. 10"'.)
Shell fusiform, acuminate, of about nine volutions ; whorls flat-
tened, longitudinally plicated and traversed by fine revolving lines,
which become crowded on the concave upper portion of the whorls,
and alternate on the basal portion of the body-whorl ; suture bor-
dered inferiorly by an elevated line, which is somewhat crenulated
by the sinuous lines of growth ; aperture less than one-half the
length of shell.
Length, 1 inch. Cave Branch, Clarke Co., Ala.
This Pleurotoma corresponds ver}- closely with the descriptions
and figures of P. acuminata as given by Sowerb}^ in the " Mineral
Conchology," and by Edwards in his monograph of the English
Eocene mollusca ( Palfeontographical Society'- Reports, 1854, p.
230, pi. xxvii, figs. 3a, 6, c, d), and will probably prove, on direct
comparison, to be referable to that species.
PYRULA, Lamarck.
[Ficula, Swainson. )
Pyrula multangulata, n. sp. PI. 20, fig. 2.
«
Shell elongated, sub-claviform ; apex of spire obtuse, consist-
ing of three smooth volutions ; whorls about seven, covered with
revolving stria?, which are very fine on the upper portion and
shoulder of the body -whorl, but less so and attenuate on the basal
portion ; body-whorl occupying about three-fourths of the entire
shell, marked b}^ two prominent and one lesser carinse, and a
sino-le row of crenulations on the shoulder angulation ; the fourth
whorl (the first one bearing ornamentation) appears cancellated ;
columella curved.
Length. 1 inch. CaA^e Branch; Wood's Bluft', Clarke Co., Ala.
Pyrula tricostata, Deshaye?. PI. 20, fig. 6.
(Coquilles Fossiles, ii, p. 584, Atlas, PI. 79, figs. 10 and 11.)
Although I have no specimen of Pyrula tricostata for direct
comparison, I have, nevertheless, but very little hesitation in
referring the Alabama form above figured to the same species, as
it agrees in all essential respects with the figures and descriptions
of that form as given by Deshayes in the Coquilles Fossiles.
Three unnamed specimens of a Pyrula in the Academy collection
from Dax, France, which I believe to be the P. clava (Oligocene?)
of Basterot, somewhat resemble the Alabama species, but are
1880.] NATURAL SCIENCES OF PHILADELPHIA. 375
•comparatively much move robust, and have the costal nodes and
revolving stria? considerably' more developed.
SOLARIUM, Lamarck.
Solarium cupola, n. sp. PI. 20, fig. 14.
Shell convexlA-^ conical, moiuid-like, of about five volutions ;
whorls ornamented with alternating coarse and very fine concen-
tric lines, and appearing double from a medial impression (the
shell apparently- of twice the number of volutions that it actually
possesses) ; base similarly ornamented as the upper surface, con-
vex, and strongly margined by the prolongation inferiorly of the
outer wall ; umbilical margin finely crenulated, the umbilicus
broadly open, and exhibiting the concentrically striated internal
volutions of the apex ; aperture rhomboidal.
Length, | inch ; diameter, 1;^ inch. Cave Branch ; Knight's
Branch ; Wood's Bluff, Clarke Co., Ala.
Solarium delphinuloides, n. pp. PI. 20, fig. 13.
Shell convexly conical, of about seven volutions ; the whorls
ornamented with several beaded revolving lines, two or more of
which near the upper margin, and one near the basal margin being
the most prominently defined; base convex, sub-marginall}' chan-
neled, and ornamented with numerous finely beaded revolving
lines, which become most prominent in the umbilical region; um-
bilical volutions distinct to the apex, transverseh' striated, super-
medially carinated ; umbilical margin crenulated ; aperture sub-
circular.
Length, ^ inch ; diameter, f inch. Wood's Bluft\ Clarke Co., Ala.
DENTALIUM, L.
Dentalium micro-stria, n. sp. PI 20, fig. 3.
Shell slender, considerably curved and greatly attenuated,
faintly striated, the stritB most conspicuous on the attenuated
portion ; posterior aperture entire, there being no fissure ; anterior
aperture circular.
Length, 1| to 2 inches. Cave Branch ; Wood's Blufi", Clarke
Co., Ala.
fusus pagodiformis (Plriiroinma .' pagodi, Heilpr., Proceedings of the U. S. Nii-
tional Museuiu, 1880), si.ecific name /<a(/o(/« pre-occupiod by Adains and ReeTe.
316 PROCEEDINGS OF THE ACADEMY OF [1880.
THE SHINING SLAVEMAKER.— NOTES ON THE ARCHITECTURE AND
HABITS OF THE AMERICAN SLAVE-MAKING ANT,
POLYERGUS LUCIDUS.
By Rev. Henry C. McCook, D. D.
August 21st, 18*78, at the foot of the Allegheny Mountains, near
Altoona (Bellwood, Pa.), I discovered a nest of Polyergus lucidus^
Mayr, the American representative of the well-known European
P. rufescens. The latter is the Amazon or Legionary Ant of
Huber, and is associated with that author's discovery of compound
ant-hills, the term applied to those nests in which certain ants
have associated witli them, in a sort of slavery, ants of another
species. Huber made a full and interesting account of the pred-
atory excursions of P. rufesans^^ and other interesting behavior,
which Forel- has recently fully confirmed and completed. It is,
however, of interest, to discover the existence of tlie same habits
in a elosel}^ allied species in America, and this record is therefore
presented. Moreover, there are here some details of architecture
which maj^ prove of value in themselves.
The nest of Lucid us above referred to was situated in the
gravelly soil of a valley between the mountains and the Juniata
River. Tlie field was sown in clover, and had not been plowed for
several years. While passing through the field, I observed several
ants resembling at first sight the common mound-makers, i^ormtca
exsectoides, issuing from a hole. I stopped to note them more
carefully and saw a worker of Pohjergui^ lucidus come out and
return to the same nest. I at once began an exploration of the
nest, as my time was limited, and professional duties prevented
extended studies of tlie out-door habits of tire creatures. There
were four gates (fig. 1, PI. 19), separated a few inches from each
other. Two were simple tubular openings into tlie ground, about
three-fourths of an inch in diameter ; the others were two similar
openings removed several inches from the first named, and united
by a worn concave road, like a half tube. The four were arranged
upon the arc of a circle. The nature of the soil, which was filled
with coarse gravel and stones, prevented me from noting (as per-
1 " Natural History of Ants, " Johnson's translation. London, 1830.
^ "Les Fourmis de la Suisse."
1880.] NATURAL SCIENCES OF PHILADELPHIA. 3YT
haps it liad prevented the ants from making) any orderly arrange-
ment of galleries and rooms in stories. But chambers were
discovered, placed one above the other, united by tubular galleries^
and extending down at least twenty-two inches, the depth to which
the excavation was canned. The general character of these may be
shown b}' the following examples. Twelve inches from the sur-
face the trowel uncovered an opening into a cavity. By gently
removing the earth, a similar opening was made just opposite
(fig. 2, PL 19). When the little bridge between the two was cut
away there was exposed an ovoid room (fig. 3, PI. 19), in which
were a number of ants, chiefly males and females of Lucidus.
The room was an inch high at the middle, and an inch and a half
across from wall to wall ; a tubular gallery led from it into the
earth beyond. Another chamber, found at ten inches below the
surface, was a large irregular cavity, which appeared, on removing
a smooth stone, flush up against which it had been mined (fig. 4,
PI. 19). It was three inches long, one inch and a quarter high, at
the highest point, and extended inward at the deepest point nearly
two inches. The line of the roof against the stone was irregular,
falling to seven-eighths of an inch to five-eighths, rising to seven-
eighths, and at one end terminating in a gallery-like extension of
half an inch. A gallery opened downward near the stone and one
opened inward at the innermost point. This chamber was also
occupied chiefly by males and females. This suflficiently charac-
terizes the internal architecture.
Mingled with the Lucidus ants, in large numbers were workers
in three forms, major, minor, and dwarf, of the species Formica
Schauff'iissi.
August 23d, the excavated nest was visited, and these ants were
found to be busy in part upon the galleries, which they were clean-
ing out, dragging the pellets of sand to the opening with the
design apparently of closing them. None of the Lucidus ants
were engaged in this work. Another portion of the slaves was
engaged in an extensive migration.' A few of the slaves were
carr3'ing their fellows, but for the most part the deportation was con-
fined to the males and females of Lucidus. The manner in which
the latter were seized and carried oif was well observed and is as
follows : The slave approached the winged queen (for example)
^ I have referred to this migration in " The Agricultural Ant of Texas,'
p. 154.
STS PROCEEDINGS OF THE ACADEMY OF [1880.
and after the usual toucliino; and crossinof of antenna; the man-
dibles were tightly interlocked (fig. 5, PL 19 ) ; the head of the slave
was then raised, and simultaneousl}- the body of the queen drawn
back, stretched quite out in a straight line, and then doubled under,
the abdomen being thrown upward apparently resting against the
lower part of the face and the fore-part of the thorax (fig. 6, PI.
19). In this i)osition the large virgin queens were carried up the
perpendicular face of the cutting for eighteen or twenty inches, and
then for the distance of six feet over the ground and through the
grass. The time consumed in this journey was a few seconds over
one minute. I frequently observed this carrying of the workers of
Lucidus, in the artificial colonies which I afterw^arcls formed and
brought to Philadelphia. The process was substantially^ the
same, although often the master was simply dragged along the
surface. Moie than once a slight opposition was made to this
treatment. The slaves, or at least certain individuals of them (for
I am persuaded that ants have their personal peculiarities of dis-
position and moods like larger animals) seemed at times to have
a prejudice against the presence of the Lucidus ants above ground,
and would unceremoniously seize them and carry them below. I
have seen a master or more properly " mistress," thus served
several times, each time returning in a dogged sort of resistance
to the will of her servitor. These emmet mistresses too, appar-
ently know something of the bitterness of bondage to a capricious
domestic " help."
The wonderful muscular force of the grip which Lucidus takes
with her mandibles was thus illustrated : One worker had for
some reason fallen under the displeasure of another, who held her
firmly grasped by the middle thorax. Anxious to preserve my
colony from unnecessary loss, I lifted the fwo out on the point of
a quill toothpick, laid them in my hand, and thrust the fine point
of the quill betw^een the jaws of the aggressor, and so teased her
until she released her hold of her fellow. The rescued ant in-
stantly clasped the palm of my hand, threw her abdomen under,
and thus with back curved up like an angry cat, sawed and tugged
at the skin until an abrasion had been made. While watching
this operation the other ant was still clinging to the quill, and to
lier I next turned my attention. She was holding fast in her
mandibles the point of the toothpick, with her body stretched
straight out into space, her limbs stretched outward, except one
1880.] NATURAL SCIENCES OF PHILADELPHIA. 379
hind leg, which was a little bent upward. Thus, without an,y per-
ceptible siippoi't, except that which her jaws gave her upon the
quill i)oint, she hung outstretched for several minutes. How
long she would have kept this position I know not, for I dropped
her into the nest b}' clipping off with scissors the i)oint of the
quill, which, after hugging fiercely for a while, she finally abandoned
as an unresponsive and unworthy foe.
In the course of the above migration, one queen was seen to
resist carriage so vigorously that she was finally dropped, and,
refusing to give the slave a hold upon the mandibles, was seized
b^^ the wing and dragged off. The Lucidus ants seemed to have
no Aolition in nor direction of this movement. I released a num-
ber from their porters during various stages of the transit, who
alwaj's wandered about with a confused, aimless and irritated
manner until again seized and borne off by slaA'es.
The locality to which the formicary was being thus transported
was about six feet distant from the gates of the original nest. It
was either an old nest or a portion of the one just disturbed. The
quarters at least appeared to have been formerl}" prepared and
occupied. The gates of the nest were placed in one sloping side
and in the angle of a deep cross-furrow, and were quite well con-
cealed by tall grass and clover, tufts of sheep-shaw and various
small weeds (see fig. T). In the angle of the furrow was a cleft
in the earth nearly two inches long, one end of which was rounded
into a gate of the size and character of those first described, and
at the other end into a smaller similar vertical tube. This entrance
was so well concealed by grass that I did not see it for some time
(fig. 8, PL 19). Tsvo and a half inches diagonally above this was
a lateral cleft, three inches long, from a half to three-fourths of an
inch high, and penetrating into the earth laterally at various points
by galleries. The stalks of grass growing upon the side of the
slope above sent down their roots through the roof of this cleft vesti-
bule into the floor. On one side of the cleft, half an inch above it,
was an entrance, with a dome-shaped vestibule. On the other side,
three inches above, was a fourth gate, opening under a round
stone. While some slaves were engaged in deporting their For-
mica fellows and Polyergus associates into the new home, others
were busy bringing out sti'aws and sand as though preparing the
galleries and chambers Avithin. Occasionall}- a Lucidus worker
would show herself for a moment at the gate with outreaclied
380 PROCEEDINGS OF THE ACADEMY OF [1880,
antennae and open mandibles, as though on the watch for intruderSj
and then slowly return to the interior.
September lYth, twenty-scA^en days after the discovery of this
formicary, I was again at Bellwood, and revisited it. The new
nest seemed to be deserted ; the ground around the gates seemed
to have been recently disturbed by a visitor, and no ants were
visible. The old nest, however, was abundantly peopled, and
numbers were found two and a half feet below the surface, from
which I was enabled to gather a large colony of slaves and workers
ofLucidus. The winged forms were gone. Mr. Edgar Kay, who
had assisted me in the excavations at the first visit, and had kept
an eye upon the nest, reported that a few days after my departure
(in the latter part of August), he had seen one male and several
females taking flight. They perched upon grasses, etc., and thence
flew eastward, at a height of forty or fifty feet, to the end of the
field, some 300 feet distant. It is probable that after this marriage
flight of the sexes, the workers returned to the old home.
After these ants were colonized, I was able to observe several
facts, chiefly eonfirmator}- of those recorded b^' Huber, Forel and
others, of the European Polyergus. The masters never work: the
colony was changed several times in order to incite to new work
in mining galleries and rooms ; clusters of Lucidus were placed by
themselves ; alwa^^s they remained idle. The slaves wrought with
the greatest industry and energy as long as there was any need ;
the masters would crowd into the galleries, and move about in an
aimless way, but I never could trace any attempt either at direct-
ing or aiding in the work. So also I never saw one attempt to
eat. Sugar was fed freel}'^ and the slaves freely partook, until
the}" became gorged, and their abdomens grew transparent with
the pouched supph^ of liquid sweets. The masters strode over
the grains of sugar, and even when I had supposed that I had pre-
pared them with a good appetite b}^ previous fasting, they partook
of nothing. Yet tlie^- are in good condition, and evidently well
fed. They doubtless are fed b}' the workers who must disgorge
the food, as when feeding larvae, callows, males, females, and even
each other. I have, however, never yet seen the actual passing of
nutriment from one to another, although often observing Lucidus
and Schauffussi in the posture which is commonl^^ assumed
when this mode of conveying food is being practiced.
In galleries and rooms the Lucidi hang upon the sides or to the
1880.] NATURAL SCIENCES OF PHILADELPHIA. 381
ceiling, or are snugged in little clusters with the slaves. In
changing formicaries they were found thus rolled together in
balls, slaves and masters — or more properly, mistresses, for all
workers are undeveloped females — mingled in a promiscuous mass.
One such change was made October 14th, the weather at the
time being chillj-, and the furnaces not yet fired in the house.
But little eftbrt was made to dig new galleries until evening, when
I warmed up the glass globes b}- a gas-lamp. My experience here
has been quite the opposite of Ruber's, who took such pains to
keep his artificial nests of P. rufescens from the light. Just as
with my agricultural ants, so P. lucidus at once turned to the
genial warmth and light, gathering in a great ball within the com-
fortable glow. A few of the slaves mounted the glass in the
warmest place to be found. As often as 1 would revolve the globe,
the cluster of snu^ffing; ants would unravel and transfer itself in
new mass upon the side toward the flame. The slaves also cheer-
lully work on the side toward the light, and indeed seem to prefer
to do so.
Lucidus cleanses herself quite freely, but also I have very
frequently seen her soliciting the slaves to this service, who some-
times consent, and go over the bodj- from antenn* to abdomen,
licking and scraping it. The need of this service especially
appeared upon examining a dead Lucidus. Its body at various
parts was covered with minute white, ovoid objects, apparently
parasite eggs. I thought them parasites, but could discover no
trace of life, or appearance of being living creatures. One seemed
to be suspended to the ant's body by a thread-like attachment.
Man}' of the slave-makers are thus aftected. While taking out the
colony one slave was found upon wliieh were fastened two small
white insects, apparently mites, which I could not then examine,
and unfortunately lost among ni}- specimens. I have often ob-
served ants to be infested with mites in natural site, and particu-
larly in artificial nests. The greatest care is required to keep
them in healthy condition while in confinement. The admirable
structural provision for cleansing the person given to ants,' is
certainly needed in view of the liability to such dangers.
The listless, heavy manner that is characteristic of Lncidus in
common, is Avholly changed at any alarm, or the presence of an
enemy. Her true character and duty to the community then
' See " Agricultural Ant of Texas," p. 130.
382 PROCEEDINGS OF THE ACADEMY OF [1880.
appear. Various experiments established the fact that some of
. these slave-makers (apparently) always keep on guard, and that
certainl}' some are ready to spring at once to repel any attack.
For exami)le, one of the slave-making Formica .'^angumea, found
in the same neighborhood, was dropped into the Polyergus colony.
The hostile presence was instantly discerned and a Lucidus
worker sprung upon the Sanguinea and seized her near the throat.
Several slaves ran to the fray, and took part by seizing legs and
antennae of the intruder. Xot wishing such an unequal conflict, I
lifted the principal combatants out, having teased away the others,
and set them down to fight it out fairl}-. Lucidus had Sanguinea
grasped by the face at the eye with her mandibles when first
removed. . This was not satisfactory, for she began cautiously
and deftly to release her hold, preparing herself meanwhile, so
that with a quick snap she seized her foe by the neck, then turned
up the abdomen, and, as I suppose, ejected poison upon the face
and mouth of Sanguinea. I separated the two before either had
been mortally hurt. However, Lucidus had lost the flagellum
of one antenna. I put lier back into her nest. The battle-scarred
warrior had no sooner struck the soil which she had so gallantl}'
defended, than she was violentl}^ seized by a slave, and dragged
up and down by her sound antenna, the poor jointless scape mean-
while thrust out and waving piteously. The late exalted mien
and ferocious aspect^ were now gone, and the warrior cringed her
body and drooped her limbs like — it is no mere fancy word-painting
this — a sullen criminal in the hands of a policeman. The two
disappeared from my sight in the mouth of a gallery ; but half an
hour afterward I saw the same warrior, whom I recognized b}^ the
mutilated antenna, in the clutch of one of her scarlet fellow-
soldiers, who was mounted upon her back and holding her by the
neck.
I am happy to record that two days thereafter I saw the
same veteran, evidently again in " good odor," perambulating the
surface of the formicary. It is probable that in the battle her
body had been tainted by some odor peculiar to her adversary,
which had made her obnoxious. It may be, indeed, that the loss
of the upper part of the antenna may have impaired recognition,
and so caused this hostile treatment. At all events I could not
but wonder whether any thought went through the little ci-eature's
brain analogous to our meditations upon the ingratitude of Repub-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 383
lies, and the vanit}' of military glory ! This incident, and many
other observations, go to establish that in the function of the
warrior is the true economy of this ant. The manner in which
her European congener Rufescens makes her raids upon the nests
of Formica fuse a and F. cunicularia, marching in solid column,
and conducting war with activity, intelligence and success, may
be read in the fascinating pages of Huber and Forel. There is no
doubt that our American species has precisely the same habit.
Mr. Joseph Jeanes, a well-known member of this Academy, has
described to me the raids of an ant observed by him upon his
conntrj'-place at Fox Chase, which, from his description of the
insect, without a specimen, I should have little hesitation in identi-
fying as our P. lucidua.
The slaves, however, are not deficient in the combative faculty
They spring to repel a hostile attack as freely and fiercely as the
masters. They do this independent!}-, too, just as they conduct
their mining operations, and their abilit}'^ to wage successful war-
fare seems to be quite in keeping with their mai'tial spirit. Dr.
Darwin has conjectured,^ that the slave-making instinct may have
originated from the unintentional rearing of pupfe collected for
food, who proving themselves useful and congenial inmates of the
nest, suggested the collecting of pupjae to be reared. Thus origin-
ated a habit, which by natural selection was strengthened and
made permanent, and finally increased and modified, until an ant
was formed as abjectly dependent on its slaves as P. rufescens.
Whatever credit we may give to this ingenious hypothesis, it
must be said, that in the case of our F. ScJwjiJf'ussi, natural selec-
tion has not operated to degenerate the soldierly courage and
faculty, and remand the duty of defense to those associates in
whom the military faculty has been specialized. In other words,
if Lucidus has become specialized as a warrior, dropping an origi-
nal disposition and ability to labor, her slave has not become
specialized as a worker, nor dropped her combative faculty, but
seems to be possessed in all respects of the normal habits and
nature of ants of her species. At least I could trace in her no
effects of slavery, other than the strange association with and care
of her abductor. One, therefore, who accepts Dr. Darwin's sug-
gestion, must allow that natural selection has wrought toward
specialization in one section of the colony, but has been suspended
' Origin of Species, p. 26.
384 PROCEEDINGS OF THE ACADEMY OF [1880.
in its operations upon the other section. It is doubtful if the
anomalous conditions thus raised by Dr. Darwin's explanation,
be not more difficult to explain than the original conditions to
which the hypothesis was applied.
It is important to note the wide distribution of this insect across
the American Continent. During the summer of 1819, while
encamped in the Garden of the gods, studying the Honey and
Occidental Ants, a nest of Lucidus was discovered just inside my
tent door. Its gate was a simple opening into the ground, into
which both Lucidus and her slaves were frequently passing.
There was a similar opening under a small bush about three feet
distant. The slave, or worker, was here precisel}' the same,
Formica Schaiiffussi, which is found so often in the compound
nests of both F. sa^iguinea and Lucidus in the Eastern States. On
one occasion I captured a slave carrying a winged queen from one
opening to another.
A comparison of a Lucidus taken at Bellwood, at the foot of the
Allegheny Mountains, Pennsylvania, with the Colorado specimens,
shows no difference except that the Pennsylvania example is
slightly more robust and of a somewhat darker color. The pecu-
liar uniform gloss which gives the American ant its specific or
varietal name, as distinguished from the duller color of the
European species, P. rufeftcens, marks equally the Eastern and
Western representatives. The European ant is decidedly smaller
than her American congener. The Colorado F. Schaujfussi is of
a more uniform and darker brown- color than the Allegheny
Mountain specimen.
I have no specimens of Lucidus from points intermediate of the
localities above named, but no doubt the species is spread over the
whole of our Continent.' That it carries with it its characteristic
habits, even its favorite domestic servant and associate, and that
in these respects it exhi])its the habits of its closely allied congener
of Europe, affords another interesting point in the geographical
distribution of our insect fauna.
' P. Tufescens of Europe has not yet been found in tlie warm plains of the
South of that Continent. (Catalogue Emory-Forel, p. 450, Mitth. d.
Schweizerischen Entomol. Gesellschaft. ) It would be a valuable contri-
bution to our knowledge of distribution were we to know whether or not
P. lucidus is found in our Southern States. We might venture the ano-
logical prediction from the above habit of its European congener, that it is
not found in the Gulf States.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 385
ON THE TEMPORAL AND MASSETER MUSCLES OF MAMMALS.
BY HARRISON ALLEN, M. D.
Systematic writers have described the temporal and masseter
muscles in mammals as being distinct from one another. I hope
to show that they are, in the great majority of forms, parts of the
same muscle.
I have found in my dissections that the temporal muscle,^ as a
rule, has a deep and a superficial set of fibres. The deep set arises
from the floor of the temporal fossa, and makes up the greater
part of the muscle. Most of the fibres unite to form a tendon,
which is inserted upon the apex of the coronoid process of the
lower jaw. Many of the fibres which do not so unite are inserted
upon the median surface of the coronoid process ; others again are
continuous with the superficial fibres. The superficial set of fibres
arise from the temporal aponeurosis. It is continuous in the main
with the deep fibres of the masseter, and the fibres are inserted
upon the lateral surface of the coronoid process. A partially
distinct slip arising from the median aspect of the malar bone,
and the ridge on the squama over the external auditory meatus,
is an accession to the superficial fibres, but possesses a tendency
to unite with the fibres of the deep set in the anterior portion
of the fossa. These fibres maj^ receive the name of the supra-
zygomatic portion of the masseter. They are inserted at the base
of the coronoid process, forming a thin glistening tendon within
and a little posterior to the anterior border of the body of the
masseter. The supra-zygomatic slip is merged with the large
superficial mass in the dog.
I believe that I have detected as part of the general plan of the
masseter muscle, when well developed, that it is composed first
of a tendino-muscular layer, rising tendinouslj' from the anterior
part of the zygoma or the maxilla near the infra-orbital foramen,
and is inserted muscularly into the angle ; second, of a nearly
vertical laj'er, tendinous below near the angle, muscular near the
zygoma ; third, of a nearh' vertical layer, having a disposition to
become tendinous, both near the angle and at the malar bone ;
fourth, of a smaller laj-er occupying the fossa on the lateral
surface of the ramus, and which exhibits a glistening laj'er of
1 For convenience the temporal and masseter will be held as distinct in
the descriptions.
26
386 PROCEEDINGS OF THE ACADEMY OF [1880.
tendon at tlie origin from tlie malar bone. The fibres beneath this
are continuous, in most mammals, with the superficial layer of
fibres of the temporal muscle, including the supra-zygomatic slip,
which, in some animals, is distinct in great part from the fibres
arising from the temporal aponeurosis. The masseter presents a gene-
ral resemblance to the internal pterygoid muscle, which, wherever
exampled, has shown these imperfect attempts at planal cleavage.
This outline being borne in mind, it may be well to turn to the
descriptions emplo3'ed by writers on comparative anatomy.
The descriptions of the muscles in Meckel (Vergleich. Anat.,
iv, 495) are ver}- general. The temporal is said to be covered by
a conspicuous aponeurosis ; the muscle to be more or less fan-
shaped, and gradually narrowed from above downward. The mas-
seter is said to be divided ordinarily into an outer, longer, stouter
and straight layer and an inner, shorter, weaker layer, in which
the fibres are more or less obliquelj^ placed from above downward
and before backward.
Cuvier (Legons d'Anat. Comp., 2d Ed., iv, Ime Part, 64 infra)
describes the temporal in the apes, bats, insectivora, rodents, two-
toed ant-eater, hog, ruminants and the cony. None of these includes
the arrangement of fibres above given. It is true that in the ant-
eater the masseter and temporal muscles are united, but no detail
of the character of the union is presented. Mivart (Elements
of Anatomy, 310) repeats this statement. It is evident that the
union of the muscles is here thought to be exceptional. Cuvier
and Laurillard further describe the masseter in the bats, rodents,
artiodactyles, ant-eater and the cony as composed of two portions,
a zygomatic and a maxillary. The former is present in all ; the
latter is seen in the rodents, artiodactyles, the ant-eater and the cony.
Mivart (1. c, p. 309) describes the masseter in Lagostomus and
Dasyprocta as follows, as of " great development :" "" The masseter
is divided into three portions, and traverses the singularly en-
larged infra-orbital foramen spoken of in describing the skeleton."
According to the interpretation used in this paper, the masseter in
rodents has even fewer subdivisions than in some other mammals.
Of these, at least one only passes in such direction as to permit
the expression that it " traverses the infra-orbital foramen;" and
this part is not separable from all the fibres lying on a plane lower
than that of the zygoma.
The descriptions of Cuvier and Laurillard of animals I have not
dissected, may be liere epitomized :
1880.] NATURAL SCIENCES OF PHILADELPHIA. 387
111 the horse a baaall superficial slip of the temporal exists,
which does not conceal the main tendon.
In the lion (PL 143 and 144, fig. 2) the parts marked vertico-
soufien and jugo-soutien answer in position to the superficial and
snpra-zygomatic fasiculi. The slips are identified, however, with
the muscles of the external ear.
In the description of the myology of Erinaceus ecaudatiis the
authors use the following language : " Between the crotophj'toe
(temporal) and the masseter muscles in the position of the absent
malar bone a red muscle-fasicle is seen, Avhich we have been un-
able to identify. It is lost posteriorly upon the <-emporal aponeu-
rosis, and passes under the masseter to be i .serted upon the
anterior border of the ascending ramus of the mandible."
I do not hesitate to classify this slip with the supra-zygomatic
slip of the masseter (see fig. 2, PI. TT of C & L.). Nothing to
invalidate such identification can be presented excepting the fact
that the slip has an alleged origin from the temporal aponeurosis.
It is in every way likely that either the glistening main tendon or
the aponeurotic texture of the superficial portion of the temporal
is here described.
In the above descriptions no mention is made of the union of
temporal and masseter except in 'the ant-eater, and in none is the
method of description the same as employed in this paper.
In making the dissections it was found convenient, after prepar-
ing the superficies, to dissect the masseter as far as the retention of
the zygomatic arch in position would permit. Then this arch was
sawn through at either end, and turned down. This exhibited the
continuity of the fibres attached to it, and the temporal. The
latter muscle was then studied carefully. After this the head was
sawed through from right to left vertically (frontal cut) in order to
expose the arrangement of fibres on the median aspect of mandible
in rodents, or, in lieu of this, an antero-posterior section was made.
Attention will now be invited to the detailed arrangement of
the parts in ditferent mammalian types.
In Macacus nemestrinus we find the fibres of aponeurotic origin
(superficial fibres) arising as in man. In addition, a slip of fibres,
arising from the inner surface of the malar bone and the occipital
crest, passes forward and obliquely downward, to be continuous
with the deep masseteric fibres. This slip is the supra-zj-gomatic,
and is believed to be an upward prolongation of the deep
388
PROCEEDINGS OF THE ACADEMY OF
[1880.
Fig. 1.
masseteric fibres. It joins the superficial fibres on a line with the
coronoicl process. The decn fibres furnish a brilliant tendon,
which extends forwards quite to the orbito-tcmporal septum;
thus differing strikingly from the arrangement in man. The deep
fibres are further seen to be imperfectly^ difterentiated from the
internal pterj^goid muscle.
The masseter muscle is composed of the following: — 1st. A
layer arising aponeurotically from the anterior third of the zygo-
matic arch, and pissing obliquely downward and backward to the
angle. 2d.- A k , er resembling the foregoing ; it arises from the
zygomatic arch at its middle. The fibres are nearly vertical and
end tendinously at the angle. 3d. A layer arising tendino-
muscularly from the posterior third of the zygomatic arch. Its
fibres are inserted upon the upper half of the ascending ramus of
the lower jaw. These layers merge
anteriorly ; indeed, are indistin-
guishable at the anterior border.
In the dog the superficial fibres
are much better developed than in
either the macaque or man, and
cover in the deeper fibres, while thej'
do not form at an}' part a supra-
zygomatic slip ; the general ar-
rangement of both temporal and
masseter muscles as in other mam-
mals. The masseter exhibits six
cleavages, of which the last or
deepest occupies the fossa on the
outer surface of the ramus, and is
directly continuous with the super-
ficial fibres of the temporal (fig. 1).
In the coati, Nasua narica^ the
superficial fibres, as in the dog,
completely conceal the deep fibres.
The supra-zygomatic slip is beauti-
fully distinct. Branchesof a con-
spicuoas venule which can be seen
lying upon the superficial portion
disappear abruptl}' as they appi'oach the upper border of the
supra-zygomatic slip. Subsequently dissection detects the trunk
Transverse section through masseter and
temporal muscles (one inch behind the
eye), Canis familiar is.
X, Superficial temporal fibres.
XX, Deep temporal fibres.
XXX, Superficial masseteric fibre, sepa-
rated from A' by a tendinous sheet.
XXXX, Internal pherygoid muscle, cut
transversely.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 389
of this vein lying between the masseter and temporal masses,
behind the zvo-oma. The central tendon is thicker at the root of
the zygoma and the bone over the external auditory meatus than
any other localitj'' in the temporal fossa.
The masseter has fine lajers, closely resembling those in the
dog. The deepest layer, namely', that one whose fibres occupy
the raraal fossa, has a much thicker aponeurosis than the other
la^-ers, the anterior portion of the first alone excepted.
In the opossum, Bidelphys virginianus, the superficial fibi'es of
tlie temporal are everywhere thick. The aponeurosis is well de-
veloped. The supra-zygomatic slip is not distinct. The deep
portion of the muscle exhibits a white glistening tendon, which
does not, however, extend as far as the orbito-temporal septum.
The anterior portion of the muscle is made up as is usual by the
union of the deep and superficial portion. In addition to its
forming the slip passing down to the front of the base of the
coronoid, it sends a powerful bundle to the median side of the
coronoid, a thin movable laj'er of muscular tissue, which passes
in front of the coronoid, between the medio-coronoid and pre-
coronoid portions.
The masseter is highly tendinous superficially. The tendency
to cleavage is not pronounced, and the continuity of the deep
fibres with the superficial fibres of the temporal is very noticeable.
In the squirrel, Sciurus hudsonicus, the superficial portion of
the temporal is less distinct than in man, and the supra-zygomatic
slip, while demonstrable, is not large. The superficial tendinous
layer of the masseter arises from a sijur on the maxilla below tUe
infra-orbital foramen. It passes, as is usual, downward and back-
ward toward the angle. This layer does not, as in most mammals,
form the entire superficies. A second layer arises from entire
inferior border of the zygoma, which appears to be lost upon the
foregoing about midway between the zygoma and the angle. Upon
turning this last layer downward, the third and last layer is seen,
which is continuous in the ordinary manner with the temporal
fibres. The ai-rangement of fibres on the median surface of the
mandible was not examined.
In the North American porcupine, Erethizon dorsatus, the
masseter consists of a superficial set of fibres arising tendinously
from the malar bone, and passing downward and backward to the
angle of the mandible. It arises from the anterior three-fourths
390 PROCEEDINGS OF THE ACADEMY OF [1880.
of the lower border of the Bialar bone, the entire lower border of
the enormous infra-orbital foramen. Its insertion is not only
npon the angle but the median surface of the ramus as well. The
last-named insertion occurs as follows : The anterior edge of the
muscle becomes stout and broad as it approaches the rounded
border of the bone in front of the angle. It winds round this
border, receiving as it does so a large accession from the angle,
and a portion of the adjacent median surface from the lower jaw.
This portion of the masseter lies below the jaw on the soft parts
of the neck. In addition to the above, a long, stout, fusiform
belly is inserted b}' fleshy fibres at a point half way up the ramus
in front, and above the upper border of the insertion of the in-
ternal pterygoid muscle.
Beneath the superficial fibres just described, the masseter in
Erethizon exhibits the usual tendinous fibres arising from the
angle and passing upward and forward. The third set is of great
importance in this animal. It agrees with the general plan of
arrangement in other mammals examined, but is remarkable for
its extent. It arises from the lower and median border of the
zygoma by tendinous fibres, from the side of the maxilla, at the
nasal region and supra-orbital surface of the same bone- by fleshy
slips, also fleshy from the upper concave border of the zygoma,
where fibres form the supra-zygomatic slip ; tendinous and fleshy
from the anterior and lower half of the inner wall of the orbit.
The insertion of this set of fibres is upon the ramus, between the
angle and the sigmoid notch. The anterior part of the insertion
is rounded and tendinous. It lies beneath the corresponding
border of the surperficial portion, and receives the fibres passing-
through the infra-orbital foramen. The remaining portions unite
to be inserted as already indicated.
The temporal muscle possesses a superficial portion, which
everywhere covers in the main muscle. Its aponeurosis arises
from the vertex, the upper border of the posterior half of the
orbit and the posterior and upper half of the inner wall of the
same depression. It thus covers in the post-orbital process of
the frontal bone. It is inserted entirely upon the main tendon,
and receives no fasiculus from the masseter. The deep or main
portion extends its aponeurosis forward, to be inserted stoutl}-
upon the post orbital process of the frontal bone. A thin apo-
neurosis passes downward, thence to the mandible behind the
coronoid.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 391
It will be seen that the plan of the muscles is the same as in
other mammals, but is remarkable for the muscles' subdivisions
remaining distinct from one another. In rodents having the large
infra-orbital foramen, the masseter muscle is described as having
a separate portion passing there through. Mivart, in his Elements
of Anatomy, page 309, says, in this connection: "In certain ro-
dents, e. g., Lagostomus and the Agouti, the masseter divides into
three portions, and traverses (that is, one of these portions tra-
verses) the singularlj" enlarged infra-orbital foramen." This is a
correct expression of the view usually taught. According to the
plan of description followed in this paper the masseter of Ere-
thizon in nowise differs from the muscles of the same name in
other mammals, except in the extent of development of the la3'er
to which the pre-foraminal fibres belong. I have had no oppor-
tunity of examining Lagostomus, but it is probable that the
masseters are much alike in all. The porcupine is further of
interest in the extent of encroachment of the muscular fibres
upon the orbital space. Both masseter and temporal appropriate
large surfaces. It is noteworth}^ in addition to find that the
l)ost-orbital process is here purely muscular in significance. It is,
indeed, imbedded in muscle. Notwithstanding its size, the process
has no septal significance in this rodent.
In Goelogenys the temporal is thin in the temporal fossa but
thick and massive on posterior wall of the orbital space. The
superficial layer and supra-zygomatic slip are distinct. Raising
these two portions of the temporal from the temporal fossa no
muscular fibres are seen beneath. A distinct tendon becomes
visible, however, underlying the junction of the superficial and
supra-zygomatic portions In the orbital space the superficial
portion is exceedingly robust and extends medianly the entire
depth of the posterior wall. The temporal is inserted into the
lower jaw as follows : The superficial portion arising from the
temporal fossa, and the zygomatic portion are inserted through
the main tendon upon the apex of the coronoid process ; the orbital
portion upon the median side of the same tendon and the median
surface of the coronoid its entire length.
Comparing the plan of this muscle to the others described it
may be said that the de6p part of the muscle is absent, unless the
greater bulk of the orbital portion is assigned to the deep part.
It has been generally found that the deep and superficial portions
392 PROCEEDINGS OF THE ACADEMY OF [1880.
are continuous anteriorly. It is probable that while the deep part
is absent from the temporal fossa proper, it remains in position in
the orbital space at a point answering to the post-septal depression
in animals having a partition between the orbit and the temporal
fossa. But while the deep part is absent from the proper temporal
fossa, a stout glistening tendon is here in the usual position of the
central tendon, and, as in Erethizon^ is concealed from without-
It is interesting to note that the supra-zygomatic slip is temporal?
it being doubtful whether any of its fibres are continuous with the
masseter.
The masseter bears a general arrangement to the muscle in
Erethizon. The anterior edge is less muscular than in the latter
genus. The mandibulo-zygomatic portion, whose origin from the
mandibular angle occupies the lower one-third of the surface, con-
stitutes the massive fleshy belly. The fibres are for the most part
nearly horizontal. It is covered for the upper half of its surface by
the enormous malar bone. The slip from the median surface of the
mandible is arranged as in Erethizon. It lies in part in front, and
in part beneath the internal pterygoid. Its junction with the main
body of the masseter conceals the tendinous anterior edge thereof
and is continuous with those fibres arising from the angle and the
basal third of median surface. The deep mandibulo-maxillary
portion is as in Erethizon in all essential features. It is continu-
ous with the superficial parts. A thin layer of orbital fibres
OA^erlies the temporal muscle in the orbit. Another layer is
apparently continuous with the buccinator.
In Dasyprocta the general plan of arrangement seen in Coelo-
genys is followed. The minute points of distinction therefrom not
being noteworthy save the continuance of the main tendon within
the orbital space, where it overlies the deep anterior vertical
fibres.
In Cuvier and Laurillard (PI. 245) the slips of the temporal
are represented as parts of a bi-peniform muscle. I find the supra-
zygomatic fibres more horizontal in position, darker in color and
more convex than the remainder of the muscle.
In the bats the superficial portion of the temporal may be small
or well developed. In the first variety a good example is seen in
Fhyllostoma hastatum and other American leaf-nosed bats in which
forms the superficial portion is confined to the anterior fourth
of the temporal fossa. The supra-zygomatic slip is also very
1880.] NATURAL SCIENCES OF PHILADELPHIA. 393
conspicuous. In Artibeus the superficial fibres occupy the anterior
half of the fossa. In Desmodus the fibres are confined to the
pj^ 2 anterior portion of the fossa ;
they are weak and unimportant.
^ The supra-zygomatic slip over-
lies the tendon of the main
''^^ muscle above the zygoma. In
) Lonchoglossa the muscle is
^ poorh'^ developed throughout.
The superficial fibres are re-
^, , , , , , ■ -c- duced to mere rudiments. The
The temporal and masseter muscles in a Fox
Bat {Epomophorus). supra-zygomatic slip is present.
.Yjf, Supei-flciai fibres of same. The deep portion does not
XXX, Supra-zygomatic slip of masseter ■i.po/-.|i the vertex,
muscle. 1 . -r> • 1
xxxx, Masseter muscle. The muscles in Ptcropme bats
resemble those in the American
leaf-nosed forms. The superficial fibres are confined to the anterior
third or half of the temporal fossa as seen in Pteropus medms,
Epomophorus and Cyonycteris amplexicaudata. The supra-
zygomatic slip is relatively smaller than in the Phjdlostomidae.
In 3Iegaderma frons and Phyllorhina bidens the paints bear a
general resemblance to the above group. The supra-zygomatic
slip is absent in the latter species.
In 3I0I0SSUS the superficial fibres are enormousl}' developed, they
entirely cover the deep, and arise from a continuous osseous sur-
face at the vertex which, being broad anteriorly , narrows gradually
toward the occiput. The fibres arising from the vertex-crest, when
such is present, are those belonging to the superficial set. The
supra-zygomatic slip is present. A similar arrangement is seen
in Noctilio^ in which form the main mass of fibres possess an un-
usually deep set central tendon, and the superficial layer extends
backward along the line of the vertex to the occiput. Lasionyc-
teris, Atalapha, Vesperus and Vesperugo have an arrangement of
the temporal fibres similar to the above but vary in the degree of
development of the anterior slip.
To sum up the knowledge possessed of the temporal muscle in
the Chiroptera it may be said that the deep portion is most ex-
posed in Pteropus and its congeners, and the family Phjdlos-
tomididae, owing to the small development of the anterior fibres.
In Yespertilionidse and Molossi the deep portion of the temporal
394 PROCEEDINGS OF THE ACADEMY OF [1880.
is more concealed owing to the greater development of the anterior
fibres. Those in Noctilio appear to be directly continuous with
fibres arising from the occipital crest and inserted on the outer
surface of the main tendon. With respect to the masseter it may
be said to be simpler than the typical description given at the
beginning of this paper. It possesses but slight tendencj'^ to
planal cleavage.
In the bovine type of the ruminants as seen in the head of a
calf the first or superficial layer of the temporal is continuous
with the masseter as shown In the above forms, the deep layer is
much less conspicuously developed than in them. The temporal
fossa being shallow — and not high — the central tendon is pro-
duced backward and is relatively small and insignificant. It is
not traceable over a short distance beyond the top of the coro-
noid.
The masseter muscle possesses six layers. The first is broad
and attached to the superior maxilla by an oblique line extending
the entire distance from the inferior border of the orbit to the
gum line over the first molar. The second arises tendJnously from
the angle of the lower jaw and extends obliquely upward and for-
ward, halfway up the ascending ramus. The third layer is tendi-
nous at the anterior superficies of the malar bone, the fibres
arising thence including the anterior half of the surface of the
zygoma. The fourth la3'er arises from the inferior border of the
zygoma at its anterior two-thirds, and is inserted muscularly upon
the ramus about midway between the zygoma and the lower border
of the mandible. It is this layer which is continuous with the
temporal as in other quadrupeds.
The masseter exhibits a fifth slip which appears to be a differ-
ential from the second or third layers, it overlies the temporo-
maxillaiy articulation in the form of a well-defined bundle which
arises tendinously from the root of the z3'goma. It is inserted
on the ramus near the posterior border at about its middle. A
sixth layer exists in the form of a narrow, bright tendon and asso-
ciated fibres arising from the I'oot of zygoma beneath the fore-
going.
Nothing similar to the fifth and sixth layers were seen in the
other animals examined. The sixth layer of the dog being rather a
sub-division over the ramus in front of and remote from the joint.
It is every way likely, however, that the number of the layers in
1880.] NATURAL SCIENCES OF PHILADELPHIA. 395
masseter will be found to be variable. The parts in the masseter
of the Virginian deer ( Gariacus virginianus) presented essentially
the same features as in the calf. The superficial layer of the tem-
poral resembles that of the calf, but the main tendon is small and
is without muscular fibres, as it lies behind the orbito-temporal
septum. The suiDerficial portion is small. It lies behind the
coronoid, in the posterior superior portion of the temporal fossa.
It must be said that the human anatomist seems warranted in
treating the masseter and temporal muscles distinct. Quain, in-
deed, affirms that some of the posterior temporal fibres arising
from the temporal fascia blend with the deep fibres of the masse-
ter, but the union of the muscles in man is a rare anomalj^
Macalister (Muscular Anomalies in Human Anatomy. Trans,
of the Roj^al Irish Academy, xxv, 1872, 18.) has met with it
but once. I have seen it once onl3\^ No mention is anywhere
made of the presence of the supra-zygomatic slip. It is quite-
likel}' that it maj' be occasionally seen in the cellulo-adipose tissue
above the zygoma. Of the presence of any peculiarities in the
anthropoid apes in these muscles I am uninformed.
The arrangement of the superficial layerof the temporal muscle
in man is ver}^ similar to that seen in the quadruped. This layer
arises from the temporal aponeurosis, and while thin posteriorly
is thick anteriorly, behind the orbital septum. If this layer of
fibres be divided posteriorly and the anterior portion turned for-
ward, a thick radiated tendon is displayed beneath. This is the
tendon of the deep set of fibres which here as in quadrupeds
constitute the mass of the muscle. The fibres of the superficial
and deep sets are continuous behind the orbital septum. This
method of displaying the temporal muscle has been for many
years employed b}' Prof. Joseph Leidy in his demonstrations at
the University of Pennsylvania.
From the above examination I have come to the following con-
clusions : —
(1) While it is convenient to separate these muscles it must be
remembered that in many mammalia the tendency is for the mas-
seter and temporal muscles to unite-^the deep part of the former
being continuous with the superficial part of the latter.
1 In a dissection of the muscles in a mulatto child at term, I found the
deeper plane of masseteric fibres arising from the external surface of the
temporal tendon.
396 PROCEEDINGS OF THE ACADEMY OF [1880.
(2) Man, and some of the Rodents — the latter illustrated in
Erethizon — are exceptions to the tendency. In these forms the
temporal is distinct from the masseter. In the same order, as in
Coelogenys and Dasyprocta, tlie deep portion of the temporal is
either absent or represented in a single orbitally disposed mass of
vertical fibres.
(3) The muscles in question have been much neglected. They
should be carefully dissected in all myological studies of the
mammalia.
18S0.] NATURAL SCIENCES OF PHILADELPHIA. 397
The following reports were read and referred to the Publication
Committee : —
REPORT OF THE PRESIDENT
For the Year ending November 30, 1880.
Nothing has occurred during the year to disturb the Societ}^ in
its usual course. It is a source of satisfaction that its financial
condition is better now than it was at the close of last j-ear.
Although its current income is not yet quite equal to the sum it
needs or desires, it still remains free from debt.
The effort begun more than a j'ear ago to collect subscriptions
for the purpose of establishing a maintenance or working-fund,
has not been as successful as was expected. The aggregate of
subscriptions is now $2680, of which $1550 have been paid. It
may be hoped that at the close of 1881 the Treasurer will be able*
to report that the Maintenance Fund has been largely increased.
The eftbrt should not be abandoned in despair.
In this connection it may be mentioned that an addition of
$3000 to the permanent fund of the Academy' has been made by
reserving for investment, under a by-law (Chapter IX) enacted
Ma}^, 1876, all moneys received from members for commuting
their semi-annual contributions, as long as they may retain their
membership. This commutation fee is designated life-membership,
and the fund accruing from it is convenieutl}^ called the Life-
membership Fund. The income from it is applicable to the pay-
ment of the ordinary expenses of the society.
The Charlotte M. Eckfeldt Fund, formed of money received,
June, 1879, from the executors of the late Mrs. C. M. Eckfeldt,
who made the Academy one of her residuary legatees, amounts to
$2466.86. The income from it has been temporarily assigned to
the use of the Publication Committee,
The heirs of the late Mr. Joshua T. Jeanes, who died suddenly
January 3d, 1880, have generously given to the Academy twenty
thousand dollars, the sum which he had indicated his intention to
bequeath to the Society in an unsigned codicil to his last will.
The money has been invested in approved mortgages, and b}^
order of the Academy constitutes the Joshua T. Jeanes Fund, the
income from which has been duly made applicable, like that of
the Maintenance Fund, to the general purposes of the society.
398 PROCEEDINGS OF THE ACADEMY OF [1880.
The Thomas B.Wilson Fund, the Elizabeth Phyle Stott,the Isaac
Barton, and Publication Funds are unchanged. Owing to cir-
cumstances over which the Academy has no control, the income
of the year from the I. Y. Williamson Librar}' Fund has been
somewhat diminished. But it is confidently conjectured tliat in
a short time it will be the same that it has been in the past.
The financial condition of the Academy will be found detailed
in the Report of the Treasurer, to whom the society is much
indebted for the time, care and labor which he bestows in the dis-
charge of the duties of his office.
Five 3'oung men have been receiving the benefit of the Jessup
Fund ; two for two months each, one for five, one for six, and one
for eiglit months during the j^ear.
A ])rief account of the origin of this fund, and the manner of
its application, may interest those especially who have become
members of the Society within the past few years.
Mr. Augustus E. Jessup, who became a member of the Academy
November, 1818, and died in Wilmington, Del., December 17th,
1859, gave the institution and its purposes a high place in his
estimation. He had expressed his intention to bestow on the
Academ}^ if ever able, a sum of money to constitute a perpetual
fund for specified purposes. His children determined that this
intention of their father should be realized, although he left no
written instructions on the subject.
In a letter dated March 6th, 1860, and addressed to Dr. Isaac
Lea, then President of the Academy, the^^ stated that, in accord-
ance with what they believed to be the intention of their father,
they proposed " to pay to the Academy one hundred and twenty
dollars per annum to be applied to its Publication Fund ; and the
further sum of four hundred and eighty dollars per annum, to be
used for the support of one or more deserving poor 3" oung man or
men who may desire to devote the whole of his or their time and
energies to the study of the natural sciences ; and that they
looked forward to investing in trust, at some not distant time, the
principals of the sums named, for the purpose of creating a per-
petual fund for the above-named uses."
Substantially these are all the instructions given to the Academy
for its guidance in the administration of these two funds — one to
be applied to its publications, and the other to the support of
students— aggregating six hundred dollars a year. They paid
1880. J NATURAL SCIENCES OF PHILADELPHIA. 399
this Slim regularly from March, 1860, until February, 1872, when
they ti'ansferred to the Academy — " the principals of the sums
named " — ten consolidated mortgage bonds of the Philadelphia
and Reading Railroad Company. Thus Mr. Jessup's children
generously fulfilled their promise and realized their father's inten-
tion. They have also consented that women may enjoy the
benefits of the student fund.
The action and language of the Messrs. Jessup imply, without
any doubt whatever, that their intention was to give to the
Academy six hundred dollars per annum : one hundred and
twenty dollars applicable only to the Publication Fund, and four
hundred and eighty dollars to the support of students. And to
secure this sum to the Academy annually forever, they gave to it
in trust an investment, the par value of which is ten thousand
dollars, an amount equal, at the rate of six per cent per annum,
to "the principals of the sums named for the purpose of creating
a perpetual fund," designed to be the impersonal successor and
never-dying agent of the Messrs. Jessup for the payment of six
hundred dollars annually to the Academy.
As trustee, the Academy is bound in honor, if not in law, to
adopt such proper measures as may be necessary from time to
time, to preserve entire not only the principal sum, but also
to prevent, if possible, the income from ever becoming less than
six hundred dollars, the specified sum it has been authorized and
directed to expend annually for the purposes named. Reduction
of this income must be detrimental to those who may properly ask
assistance from it, to the extent of any diminution it maj^ suffer.
It is designed to benefit students of the future as well as those of
the current time. The interests of those of the coming centuries
in it are entitled to present consideration and protection, if
needed.
The instructions under which the Jessup Fund for students
was established, describe in general terms the requisite qualifica-
tions of those upon whom the Academ}^ may bestow its benefits.
An eligible candidate for aid from the Jessup Fund, is required
by those instructions to possess the following qualifications :
1, Evident " desire" to devote the whole of his time and ener-
gies to the study of the natural sciences.
2. He must be so poor as to be dependent on his own labor for
a livelihood, and therefore, unless he can be otherwise supported,
400 PROCEEDINGS OF THE ACADEMY OF [1880.
he cannot devote the whole of his time and energies to the study
of the natural sciences, to which he seeks to dedicate himself.
3. He must be "deserving" of support in this connection.
This condition means much. To deserve any support from the
Jessup Fund, he should possess a quick natural intelligence,
above the average; a good and sufficient education, including,
perhaps, a knowledge of the German and French languages ; in-
dustrious and orderly ways ; integrity in every sense beyond sus-
picion, and lastly, a. manifest intention to dedicate his lifetime
and energies to the study of the natural sciences.
4. He must be " young "—say under twenty-five years of age.
Under such conditions, and with faculties suitably equipped
and disposed, the candidate ma}^ pass through an apprenticeship
here provided, and become a practical naturalist.
The application of the fund is entirely at the discretion of the
Academy. It would not violate the letter of the trust by using
it to support approved students of the natural sciences without
•giving them instruction, or granting them the use of its library
or museum or its hall as their workshop. The trust does not re-
quire that the Academy shall be the preceptor of the beneficiaries
of the Jessup Fund in any degree. But inasmuch as one of
the functions which the society has prescribed for itself is to im-
part and diffuse knowledge, it seems peculiarly proper that it
should direct and facilitate the studies of these beneficiaries.
The four hundred and eight3^ dollars may be given annually to
support one, or be divided between two or more, as may seem to
the Academy expedient. The time during which any one may
receive assistance from the fund, is limited at the discretion of
the Academy.
After due consideration of the subject at the start, it was
determined that the approved candidate should be received at
first on probation, for one month, and if the trial were satisfac-
tory, he might be appointed a beneficiary^ for two years, and then
retire in favor of another, unless there should be special reasons
for his continuance.
Inasmuch as the members of the society pay dues for their
right to use the library and museum, it is considered proper to
require the beneficiaries of the Jessup Fund to give, daily, a part
of their time and labor to the Academy, under the direction of
the curators, as compensation for instruction, and the use of the
1880 ] NATURAL SCIENCES OF PHILADELPHIA. 401
Academy's propert^^ This time is employed in work incident to
taking care of, mounting, and arranging specimens in the museum,
such as cleaning them when necessary, labeling, etc., a kind of
work which is pertinent to the vocation of a naturalist, and
throuffh which the beneficiaries become familiarized with natural
objects, more perfectl}^ than thej^ can be in any other way. It
should not be forgotten that the Academy has alwaj-s been de-
pendent, almost exclusivel}', upon the unpaid labor of its members
for the care of its museum, and this circumstance, perhaps, ex-
plains why beneficiaries of the Jessup Fund are expected to do
an}' kind of work in the Academ}- that the curators and other
members are in the habit of doing. They are, in fact, regarded
as almost apprentices, who should be ever ready to avail them-
selves of the opportunities afforded to learn everj'thing pertinent
to the career of a naturalist.
Applications for the benefits of the Jessup Fund, are considered
and decided by the Council of the Academy.
Between March, 18G0, and November, 1880, thirty-four persons
have received aid from the fund, for a longer or shorter period
than two years. Of these, five have died, well known and much
respected naturalists. Five of those, now living, are professors
and eminent men. It is believed that all of this class of gentle-
men have acquitted themselves satisfactorily, and that all grate-
full}^ appreciate the benificence of the Jessup Fund, as well as
the advantages derivable from it ; and it is hoped that none will
ever regret an}- of the work he has done, or the time he has spent
in the Academy.
The annual reports of the curators and librarian show the ex-
tent of increase of the museum and- library ; and those from the
several sections of the Academy indicate that they are active and
prosperous.
During the year, more than 600 pages of the Proceedings have
been published, and the fourth part of the eighth quarto volume
of the Journal of the Academy is passing through the press.
The proceedings of the Entomological Section are printed on
the premises by some of its members, and issued separately.
This section has published more than 370 pages and 7 plates
during the year.
The second volume of a " Manual of Conchology, Structural and
Systematic, with Illustrations of the Species," by George W.
27
402 PROCEEDINGS OF THE ACADEMY OF [1880.
Tiyon, Jr., published by the nuthorand issued from the Academy,
has been pviblished duriui' the year. It includes 289 pages of
text, 70 plates with 975 figures.
Professor Leidy's admirable work on " Fresh-Wster Rhizopods
of North America," forming Yol. XII of the final reports of the
United States Geological and Geographical Survey of the Terri-
tories, under the direction of Dr. F. V. Hayden, is so closely
connected with the Academy, that its publication during the j-ear
may be mentioned here. Dr, Leidy, at the stated meetings of the
society, gave verbal accounts of very many fresh-water rhizopods
which are described in his work, and the Academy's librarj^ was
the sole source from which he was enabled to prepare the bibli-
ography of the subject.
Few persons devote their whole time and energies to natural
history for a living. Generally, the study is an occupation for
leisure hours, and may be regarded as a secondary pursuit among
us, which yields little or nothing towards a livelihood. Satisfactory
study of natural history requires so much to aid its votaries, in the
way of collections and books, that it is extremely rare to find any one
person rich enough to procure all that is needed. For this reason
manj^ of like tastes associate, each contributing his quota, for the
purpose of gathering what is necessary or desirable to be used in
common for self-instruction.
In one sense the Academy may be regarded as an association
of this kind.
A prominent object of the Society is to afford opportunity to
those who desire to undertake self-culture in any or all the de-
partments of the natural sciences. From its beginning in 1812,
continuously to the present time, members have freely contributed
specimens to its museum, and books to its library. Besides
materials of this kind they have given money liberally, established
permanent funds for several specific purposes, and employed what-
ever time the}' could fairly take from their daily avocations in
working with their own hands to render the constantly increasing
means of stud}^ as easil}^ available as possible. The value of per-
sonal labor gratuitously given to establish and promote the growth
of this institution cannot be over-estimated. A result of the joint
efforts of the members of the societ}^ since its foundation is the
opportunity of self-instruction here liberally afforded to those who
may choose to avail themselves of it.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 403
Although the museum is deficient in many of its departments,
it is remarkably sufficient in some, and as a whole is very exten-
sive, and in every sense very valuable. The unequal development
or growth of the several departments is ascribable to the depend-
ence of the collections for increase on donations exclusively, and
the want of money to purchase desiderata, and not to indifference
or ignorance of those to whom the immediate care of the museum
is confided.
At this time the library as a whole, though not complete, is
perhaps the best collection of works on natural history in this
country, and the Library Fund, given by Mr. I. V. Williamson,
provides liberall}' for its increase.
The opportunity for self-culture to be found now in the museum
and library, with all their deficiencies, is a result of the generosity,
goodwill, industry and benevolence of very many members and
friends of the Society. Some expert naturalists may disparage
this opportunity of self-culture, such as it is, and take pleasure in
pointing out its defects and deficiencies, but those just entering
the field, as well as those not yet proficient will find it fully suf-
ficient for their use and worthy of cordial approbation.
Objection has been made to the regulation which restricts the
use of specimens and books to the premises of the Academy, sug-
gesting that study would be very much facilitated by loaning speci-
mens and books to members, especially to those who are advanced
students and experts. The answer is that the loan of specimens
and books, which may be regarded as a luxury rather than as a
necessity to students, would somewhat facilitate the work of one
borrower, but while they w-ere in his possession the studies of
several persons having occasion to consult the same specimens
and books might be much retarded or hindered. Besides, loaning
books and specimens increases the chance of their loss and injury.
After ample experience in the practice of loaning, and due con-
sideration of the whole question, the Academj'^ adopted the exist-
ing regulation of loaning specimens only on a recommendation of
a majority of the curators, approved by a vote of the Academj- ;
and by prohibiting the circulation of books, has made the library
a librar}' of reference exclusively. It is confidently believed that
the common interests of all concerned are best served b}' strict
adherence to this practice. It promises " the greatest good to
the greatest number" of those who have occasion to examine
401 PROCEEDINGS OF THE ACADEMY OP [1880.
specimens or consult books, because, come when they may, from
far or near, they are reasonably sure to find in the building what-
ever bclono^s to the museum or library.
The regulation in question is appi'oved by an experience of more
than a quarter of a century. It is supposable that it would have
been changed long ago had the menibers of the society believed
it to be injurious to their common interests.
It may be freely granted, however, that there are persons — those
who are seldom spontaneously considerate of the convenience and
rights of others to the use of property held in common, to Avhom
all restrictive rules appear unwise — especially when they are an
obstacle to the satisfaction of some transient interest or desire.
The same individuals would probably discover a grievance in the
loaning system, should they find that those very specimens and
books which they desire to refer to at the moment, had just been
borrowed and taken out of the building, to be returned at the end
of a fortnight or possibly a month. Persons of such temperament,
unhappily for themselves as well as their associates, are prone to
find that " All goes wrong, and nothing as it ought," where others
of more happy constitution discover nothing unpleasant.
The opportunity of self-instruction in the Academy is good as
it now is. It is accessible to those who may desire to make use
of it under the rules of the society. They must come spontane-
ously. The Academy is not prepared to enlist regulars, hire
mercenaries, or solicit volunteers in order to bring into more
extensive use the opportunity of self-instruction which it has
built up.
When both the museum and library were easily contained in
one small room, the effort of the Academy to increase its means
and opportunities of self-instruction was approved. No one com-
plained of its deficiencies. All cheerfully endeavored to use profit-
ably what it had. The propriety of admitting to its membership
those who possessed no other qualifications tlian friendliness to
scientific pursuits and personal respectability was not questioned.
But since the possessions of the Academy have grown to be ex-
tensive and of great value in every sense, there are individuals
who lament that they are not greater, and seem pleased to dis-
parage its condition, its course and its organization. Ignoring
all that is recorded in the sixteen volumes of the first and second
series of the Journal of the Academy and in the thirty-two
1880.] NATURAL SCIENCES OF PHI LAD ELPUr A. 405
volumes of its Proceedings, tliey imagine that it sadl}' lacks the
afflatus of pure science and does nothing to pi'omote research.
Their tone implies that the capabilities of the institution, the
potentialities of its possessions might be made more useful to
truly qualified investigators by reforming the present S3-stem and
policy, which are too broadly in the interest of beginners and
amateurs in science. They seem to believe that the collections
should be placed under the control of expert specialists, with
power to loan specimens at their discretion ; that the books of
the librarj" should be allowed to circulate freely, and finally, that
the society should consist of proficients exclusivel}', or at least
include a privileged class of experts.
Whether the Academy should now permit its extensive museum
and librar}', which have cost so much time, labor and mone}' to
form, to be diverted from their present ways of usefulness to
students generally, and appropriated by skilled investigators, is a
question too important to be hastil}"^ decided.^
The b3'-law of May, 1876, which provides for the appointment
of professors, remains inoperative. No candidate has presented
himself during the year. No report ha's been . i-eceived from the
Professor of Histolog}^ and Microscopic Technolog}', who was
appointed April 16, 1877.
In conclusion, it may be said that the condition of the Academy
has never been better since its foundation than it is at the present
time. It is independent of debt, and its income has been so far
increased that it is hoped, under a careful administration of its
financial afiairs, it will soon be sufficient to meet the usual demands.
The whole is submitted,
W. S. W. RUSCHENBERGER.
^ A society composed exclusively of proficients may be desirable and
even essential to the progress of original investigation in Philadelphia.
Those who are of this opinion might possibly form such a society at once,
and in the course of time acquire all it may need ; and, without coveting
or attempting to appropriate its possessions, permit the Academy to exist
for the benefit of those proficients who approve of its organization as well
as of beginners and amateurs. Some of these might become qualified to
be admitted to membership of any society composed exclusively of gener-
ally recognized masters in science.
406 PROCEEDINGS OP THE ACADEMY OF [1880.
REPORT OF THE RECORDIXG SECRETARY.
The Recording Secretary respectfully reports that during the
year ending November 30th, 1880, twenty-six members and twenty
correspondents have been elected.
Resignations of membership have been received from D. E.
Dallam and J. D. Thomas.
Records of the death of twenty members and four correspond-
ents have been published in the Proceedings under the dates of
announcement.
Twenty-five papers have been accepted for publication as follows :
H. C. Lewis, 7 ; J. S. Kingsley, 3 ; Jos. Leid}', 3 ; H. C. Chapman,
2 ; Harrison Allen, 1 ; R. Bergh, 1 ; Andrew Garrett, 1 ; A. W.
Yogdes, 1 ; W. N. Lockington, 1 ; W. D. Havtman, 1 ; Wm. Bar-
beck, 1 ; Angelo Heilprin, 1 ; T. D. Rand, 1, and F. A. (Jenth,
Jr., 1.
Twenty-two of these papers have been published in the Pro-
ceedings and three in the Journal. In addition , nine papers published
in the Proceedings, together with reports of a nnmber of important
verbal communications, formed the Proceedings of the Mineralo-
gical and Geological Section of the Academy- for the years 18 YT
to 1879.
Two hundred and eighty-eight pages of the Proceedings for
1819 and three hundred and fiftj'-two pages of the volume for 1880
have been printed during the year. The concluding number of
Yolume YIII of the Journal will be issued early in January.
The list of those making verbal communications at the meet-
ings includes the names of Messrs. Leidy, Meehan, Allen, A. J.
Parker, Wilcox, Koenig, Cope, Kellv, Ryder, Evarts, Frazer,
Dercum, Horn, McCook, Barbeck, Kingsley, Chapman, Potts,
Canby, Foote, Coates, Tasker, Martindale, Pike, Ford, Halde-
man, Redfield, Porter and Hough.
At the meeting held January 13th, 1880, Messrs. Aubrey H.
Smith and Geo. Yaux were elected to fill vacancies in the Council
caused by the absence from the meetings thereof for six consecu-
tive months of Dr. C. Newlin Peirce and Prof. Edw. D. Cope,
and on November 16, 1880, Mr. Ezra T. Cresson was elected to fill
a vacancy caused by the resignation of Mr. Geo. Yaux.
All of which is respectfully submitted.
Edw. J. Nolan,
Recording Secretary.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 407
REPORT OF THE CORRESPONDING SECRETARY.
In accordance with the By-Laws of the Academ}-, the Corre-
sponding Secretar}' presents the following Report of the business
of his office during the year ending November 30th, 18S0.
Thei-e have been elected twent}' Correspondents, as follows :
Angelo Heilprin,* New York City; Dr. C. A. White * Wash-
ington, D. C. ; Albert de Selle,* Paris, France ; Victor Raulin,*
Bordeaux, France ; R. Hoarnes,* Yienna, Austria ; Georges Rol-
laud, Paris, France; A. Inostranzeft',* St. Petersburg, Russia;
Dr. Robert Schomburgh* Adelaide, Australia ; Dr. Herman T.
Geyler, Frankfort a. M., Germany ; Robert Casparis, Konigsberg,
Germany ; Agostino Todaro, Palermo, Italy ; J. E. Bommer,
Brussels, Belgium ; Prof. Teodoro Caruel,* Pisa, Italy ; Lionel
S. Beale,"*" London, England ; Prof. Richard Hertwig,* Jena,
Austria ; Prof. Oscar Hertvvig,* Jena, Austria ; Dr. Carl Ochse-
nius,* Marburg, Prussia ; Dr. M. H. De Bey, Aix-la-Chapelle,
Prussia ; Prof. Adolf E. Nordenskjold,* Stockholm, Sweden ;
Prof. Torquato Taramelli,* Pavia, Italy ; all of whom have been
promptly notified, and acceptances have been received from those
whose names are marked with an asterisk, *.
The donations to the Museum have been numerous and valuable,
as will be learnerl from the Curators' report, and prompt acknowl-
edgments have been sent to the various donors, numbering in all
228.
Letters transmitting publications have been received from Cor-
responding Societies or Institutions, at home and abroad, to the
number of fifty-one ; from individuals, four.
Letters or other acknowledgments of the reception of the pub-
lications of the Academy have been received to the number of
fifty-two.
In addition to the above, thirteen letters of a miscellaneous
nature have been received, and those requiring an answer have
been in all cases replied to.
Respectfulh^ submitted,
George H. Horn,
Corresponding Secretary.
408
PROCEEDINGS OF THE ACADEMY OF
[1880.
REPORT OF THE LIBRARIAN.
During the past year, from December 1st, 1879, to November
30th, 1880, the library of the Academy has been increased by
2744 additions, mainly exchanges received for the publications of
the Academy. The accessions have included 310 volumes, 2.345
pamphlets and parts of periodicals and 89 maps, photographs,
portraits, etc.
They were derived from the following sources : —
and
Societies 1018.
I. V. Williamson Fund 522.
Editors 487.
Authors 218
Dr. I. Minis Hayes 118.
Wm. S. Beebe 61.
Wilson Fund 50.
Department of Mines, Nova Scotia,
23.
Geological Survey of Sweden 23.
Geological Survey of Belgium 19.
Department of the Interior 17.
Isaac Lea 14.
.Jos. Leidy 14.
Geological Survey of Pennsylvania
13.
University of Chili 12.
Dr. F. V. Hayden 11.
Wm. S. Vaux 10.
Geological Survey of Canada 10.
I)e])artment of Agriculture 10.
Smithsonian Institution 8.
Engineer Department, U. S. Army
7.
Geological Survey of India 6.
,1. H. Redfield 6
Geological Survey of Wisconsin 5.
(Jolonial Secretary N. S. Wales 5.
Geological Survey of Minnesota 5.
Treasury Department 5.
Geol. Surv. N. Zealand 4.
Yale College 4.
Minist. of Pub. Instruction, Bel-
gium 4.
British Museum 3.
The books and pamphlets thus acquired were distributed to the
several departments of the librar}^ as follows : —
War Department 3.
U. S. Coast Survey 2.
Kansas State Board of Agriculture
2.
Geol. Surv. N. J. 2.
Jos. M. Gazzam 2.
R. Schomburgli 2.
John Brazier 2.
Mrs. Charles Pickering.
S. S. Ilaldeman.
Rathmell Wilson.
U. S. Commission of Fish
Fisheries 1.
Commissioners of Fisheries, Cali-
fornia.
Mines Commissioners of Maryland.
Geol. Surv. Kentucky.
Geol. Surv. Indiana.
Geological Survey of Japan.
University of Minnesota.
Directors of City Trusts.
Meteorological Office, Canada.
South African Museum.
Government of Victoria.
Commissioners of Public Charities,
Penna.
East Indian Government.
Bureau of Education.
Public Library, Milwaukee.
Astor Library.
Library Co. Phila.
Mercantile Library Associations of
San Francisco, St. Louis, New
York and Cincinnati, each 1 .
Journals 1969.
Geology 132.
General Natural History 169.
Anthropology 88.
Conchology 68.
Botany 61.
Anatomy and Physiology 41.
Entomology 36.
1880,] NATURAL SCIENCES OF PHILADELPHIA. 409
Ornithology 34. Mammalogy 9,
Bibliography 26. Ichthyology 4.
Physical ir^cieiice 21. Voyages and Travels 4.
]\Iineralogy 16. Herpetology 3.
Helminthology 16. Microscopy 1.
Agriculture 13. Miscellaneous (Historj^, Statistics,
Chemistry 11. Politics, etc.) 12.
Encyclopedias 10.
From the above statistics and the accompanying list of addi-
tions it will be seen that, apart from exchanges received from
societies and editors, the gro-wth of the library has been raainlj-
dependent upon the I. Y. Williamson Fmid.
It gives the Librarian pleasure to be able to report the comple-
tion of the card catalogue of all the special departments of the
library coming within the province of the Acadeni}'. It is to be
hoped that some of the remaining sections, at present arranged
on the gallery, may soon be disposed of hy sale or exchange, as
the}- embrace books of a character rarely- or never consulted in
the Academy ; although many of them would be of importance
and Talue elsewhere. The revision of the catalogue of journals
and periodicals is progressing slowly, as time is taken after the
completion of each geographical section to apply for all deficiencies
noted. The answers to such applications thus far made have beeir
so satisfiictor}- as to warrant the hope that important additions
will be received from this source during the coming year.
The collection of portraits of the Presidents and benefactors of
the Academy has been increased b}^ the addition of a fine oil
painting of Isaac Lea, LL. D., by Ulile, one of Dr. Isaac Hays,
b}- Waugh and a life-sized crayon portrait of Mr. Isaiah V. Wil-
liamson. For these gifts, interesting not onh' as works of art,
but also as memoi'ials of men to whom the society is indebted for
many and permanent benefits, the thanks of the Academy
are due to Dl". Lea, Mrs. Dr. Isaac Hays and Mr. Williamson.
The Acadeni}^ now possesses the portraits of eight out of its ten
presidents, those of Dr. Thos. B, Wilson and Dr. Robert Bridges
being j-et lacking. It is to be hoped that these ma}' be supplied,
and that the series, which will certainl}' be of great interest here-
after, may be kept complete.
Fine framed photographs of Dr. Jos. Leid}^ and the late Prof.
Henr}^ have been received from Mr. F, Gutekunst,and Dr. C. W.
De Lannoy has presented a death mask of Di*. James Aitken
Meigs.
410 PROCEEDINGS OF THE ACADEMY OF [1880.
For the amount expended from the various funds for books you
are respectfully' referred to the report of the Treasurer.
Edw. J. Nolan,
Librarian.
REPORT OF THE CURATORS.
The Curators respectfully report that the Museum of the
Academj' continues in its usual good state of preservation. The
following report of the Curator in charge gives brief notice of
what has been done, and the additions which have been made
during the year.
Sir : — I would respectfully report, that during the year all the
collections of the Mviseum have been carefully inspected and cared
for, and that they are in good condition. The vertebrate fossils
are in process of arrangement.
Dr. J. Allen Kite has been engaged in the arrangement of the
collection of Bird-skeletons, and Mr. Angelo Heilprin in the ar-
rangement of the Invertebrate fossils.
The specimens received during the year have been labeled and
placed in their proper positions.
The contributions in the various departments during the year,
excepting those reported on by some of the special sections, are
as follows : —
Mamwah. — Zoological Society of Philadelphia : Two 3Iacacus
ocreatus, Macacus maurus^ Ateles ater, Gercopithecus lalandi,
Fterojyua vulgaris, Heiyesfes griseus, Bassaris astida, Viverra
indica, two Tragulus Javanicus, Dasyprocta acouchi, Ca?logenys
paca, Sciiiriis variabilis, HypsiprymniLs rufescens. Jacob Binder :
A colloidal mass with nodules of osteo-dentine embedded, from the
tusk of an Elephant. Dr. H. C. Chapman : Placenta of Asiatic Ele-
phant, born in Philadelphia. Dr. Geo. H. Horn : Two Atalapha
{Lasiurus) noveboracensis, Phila. Jos. Jeanes : Two young Ele-
phant skulls, Elephas indie us and E. africanas. Albert Koebele :
Nycticejus crepuscularis^Y'lorxdvi. Dr. Jos. Leid}- : Hesperoinys
(sp.). Roan Mt., N. C. ; Buffalo jaw, from a forest in the Uintah
Mts., portion of the great part of a skeleton observed by him in
the locality in which it is now extinct. Miss Miller : Horns of
Chamois, Alps. W. S. Yaux : Young Orang-Outang, from Phila-
delphia Zoological Gardens.
Birds. — Philadelphia Zoological Society : Sycalisjlaveola, Bro-
togerys xanthoptera, Brazil ; Anser indicus. F. W. Allen : Dio-
1880.] NATURAL SCIENCES OF PHILADELPHIA. 411
medea exulans. Hill : Otus vulgaris. Mrs. Herbert Russell
Walsh : Two hundred and ninety -seven (one hundred and twent}--
one species) Bird skins, collected and prepared by the late Robert
Frazer.
Amphibians and Fishes. — Albert Koebele : Eleven species Am-
phibians,' Florida. Dr. Jos. Leidy : Two species Salamanders,
two do. Fishes, Roan Mt., N. C. Dr. H. Allport : Erimyzon
sucetta, Centre Co., Pa. S. W. Ayer : Opercular bones, etc.,
Megalojys fhrissoides. Mr. Holbrook : Argyrieosus (Vomer) seti-
pinnis^ Atlantic coast, Md. Dr. W. H. Jones: Nine species of
Fishes, Atlantic and Pacific Oceans. J. E. Mitchell: Amhlyopsis
spelaeus^ Mammoth Cave, Ky. National Mus., through Smiths.
Inst.: Fifty-two species of North American Fishes.
Articidafe!<. — J. J. Brown : Lepas pectinata, Balanusyetc, Flor-
ida. Dr. H. C. Chapman : Lepidnotus, and Nymphon^ Mt. Desert,
Me. C. Chambers : Grillotalpa longipennis., Philadelphia. John
Ford: Libinia canaliculata^ Atlantic Cit}', N. J. Geo. Heberton :
Libinia canaliculata, Limulus pjolyphemus. Cape May, N. J. Dr.
W. H. Jones : Thirty-four species Crustacea, Atlantic and Pacific
Oceans ; Nautilograpsus minutus^ taken from side of ship Aca-
pulco. J. S. Kingsley : Six species Crustacea, in exchange. Dr.
J. A. Kite, Wasps' nest, Morgan Co., Pa. All>ert Koebele : Two
species Crustaceans ; two species Myriopoda, from Florida. J. E.
xMitchell: Nest of Tarantula, California. Dr. T. H. Streets:
Forty-two specimens Lepidoptera, Mantis (sp.), Yokohama, Japan.
U. S. Fish Commission, through Smiths. Inst. : Thirty named
species of Crustacea, Coast of New England.
Vermes., Echinoderms, Goslenterafes, Bryozoans and Porifera.
— U. S. Fish Commission, through Smiths. Inst. : Thirteen species
of Annelida, Coast of New England. Dr. W. H. Jones : Four spe-
cies of Annelida, Pacific Ocean. Dr. Jos. Leidy : Lice from the
interior of pouch of White Pelican {Menopjon perale, Leidy),
Florida. Laura M. Towne : Filaria immilis, from heart of dog,
Beaufort, S. C. J. J. Brown: Gidaris trihuloides, Haiti. John
Ford : Euryale (sp.), Palermo, Ital3^ U. S. Fish Commission,
through Smiths. Inst. : Sixteen species Ecliinodermata, Coast of
New England. Dr. W. D. Hartman : Rotula (sp. ), Madagascar?
U. S. Fish Commission, through Smiths. Inst. : Eleven species of
Coslenterata, Coast of New England. Dr. W. H. Jones : Thirteen
species Coslenterata, Pacific Ocean. Dr. H. C. Chapman: Hyd-
roids, from Mt. Desert, Me. U. S. Fish Commission, through
Smiths. Inst. : Fifteen species Brj'ozoans and Porifera, Coast of
New England.
Anna T. Jeanes : Glass models of Physophora niagnijica^
Diphyes Sieboldi, Cyanea cajnllata, Oceania phosphorica.^ Sagai^-
tia bellis., Palythoa auricula, Phellia pjicta, Corynaclis clavigera,
Nemacula primula, Peachia hastata, Phymactis Jlorida, Evactis
artemisia, Tubulay'ia indivisa, Gorymorpha nutans, Laomedea
amphora, Bougainvillia fruiicosa.
412 PROCEEDINGS OF THE ACADEMY OF [1880.
Fossils. — Dr. Carter, through Dr. Jos. Leidy : Fragments of
bones of Uintatherium, Palsposyops, and fossil turtle-eggs, near
Ft. Bridger, W3^onilng. Walter Collins : Eight species of Fossils
from the cretaceous marl, Blackwoodtown, N. J. Dr. Corson,
through Dr. Jos. Leidy: Femur and fragments of jaw of Pal aeo-
syops major, and foot-bones of Uintatherium, near Fort Bridger,
Wyoming. Persifor Frazer : Two fossils. Wm. M. Gabb : Fish
teeth, Martinez, Contra Costa Co., Cal. G. N. P. Gale: Shark's
teeth and fragments of bone, Ashley River beds, S. C. Gustavus
Guttenberg : Fossil Fucoid ? near Erie, Pa. M. Hotchkiss and
John Gibb : Three fossils. Central coal shaft, Fairburg, 111. Dr.
Geo. M. Lawrence: Ammonites (sp.). Cretaceous of Hemstead
Co., Ark. R. L. Lamborn : Fish tooth (carboniferous), Irwin
Station, Westmoreland Co.. Pa. Dr. Jos. Leidy : Ivory of Mas-
todon, bored by mollusks, Rib of Manatee, and two fish vertebrae.
Phosphate beds, Ashley River, S. C. ; Tibia of Palseosyops., fossil
turtle-eggs, near Ft. Bridger, Wyoming. Thomas Meehan : Four
coal fossils, Schuylkill Co., Pa. J. W. Pike : Forty-eight specimens
of fossil ferns, etc., Mazon Creek, Grundy Co., 111. J. H. Redfield :
Pentremites <TO?YZo?in(subcarboniferous). E. S. Reinhold : Thirty-
six specimens coal fossils, Mahanoy City, Pa., A. L. Siler :
Fossil (sacrum). West branch of South fork of Rio Virgin River,
above Glendale, Utah. Wm. Spillman : Gcelorh.ynchus ornatus,
Clark Co., Miss. Charles Wachsmuth : Forty-five species of
Crinoids, from the Burlington Limestone, Burlington, Iowa.
Harry H. Wheeler : Crinoid, cut from a side-walk paved with upper
Silurian Limestone, Wabash, Ind. ; Four fossil Fishes, near Fort
Bridger, Wyoming.
Ethnological and Miscellaneous. — Stephen Bowers : Sixteen
Indian skulls, from a burial place, Santa Barbara, Cal. ; Skull of
Flathead Indian, Oregon ; Skull of Peruvian, Onca, Peru. Miss
E. S. Boyd : Hawaiian skull, Sandwich Islands. W. C. Desmont :
Dart, used in salmon fishing by the Digger Indians, Sacramento
River, Cal., 1858. Jacob Geismar : Stone axe and arrow-head,
Haddonfield, IS". J. Prof. S. S. Haldeman : Eight pieces modern
Pueblo pottery. New Mexico; Tunisian drum, Tunis, N. Africa;
Stone tomahawk, two hammers, chisel and pestle, Gloucester Co.,
N. J. ; Stone chisel, Indiana ; Pestle, Ohio ; Stone implement
found on the farm of Mr. Wittmer, Lancaster Co., Pa. ; twelve
pieces Pueblo potter}', New Mexico ; two stone hammers, modern
Sioux, and fragments of an earthen pot, from a cave, E. Tenn. ;
fifty-seven pieces of native pottery, two Sorcerer's chairs. Con-
juror's rattle, Necklace, three bowls made of the gourd-like fruit of
the calabash tree (Grescentia cujeta), by the Indians of British
Guiana; Native cloth, Island of Corisco. Prof W. de M. Hooper:
Stone implement, used ifor skinning, Misner's Farm, Pittsburg,
Carroll Co., Ind. Wm. L. Mactier: Snow-shoes, made by the
Otonagon Indians of Michigan. Miss Miller : Antiqiie linen shawl,
vase, lamp, mummied human hand,' ibis, etc., Eg3'pt. Wm. J.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 413
Potts : Fragments of Indian skeletons, pottery, etc., bank of the
Delaware River, below Kaighn's Point, Camden, N. J. Jos. Wil-
cox : Fragments of pottery, from a mound, St. John's River, Fla. ;
Stone axe, pestle and arrow-head, Mitchell Co., N. C. ? Two
pieces Ancient Peruvian potter}^
C. F. Parker.
Respectfully- submitted by
Joseph Leidy,
Chairman Curators.
REPORT OF THE BIOLOGICAL AND MICROSCOPICAL
SECTION.
During the past j-ear eighteen (18) meetings were held ; the
average attendance being thirtj^ (30) persons.
The Annual exhibition was held on the evening of October 15th,
at which time a large and interested compam- was present.
The following is a summary of the principal subjects presented
during the 3'ear : —
Dec. 1st, 18T9.— Filarite in the Bronchial Tubes of Cattle, by
Dr. James McCoart.
Dec. 15th, 1819. — Modern Microscopical Work, by Dr. J. Gib-
bons Hunt.
Dec. 15th, 18t9. — Description of Psorosperms found in Cysts
of Fishes, by John Ryder.
Jan. 5th, 1880.— Pleuro-Pneumonia of Cattle, by Dr. John
Gadsden.
Jan. 15th, 1880. — The Microscope as a means of Investigation,
by Dr. Carl Seller.
Feb. 2d, 1880.— The Preparation of Material for Microscopical
Examination, by Dr. Seller.
Feb. 2d, 1880. — Nuclei of the Eggs of the Common Limpet, by
Mr. John Ryder.
Feb. 2d, 1880. — Observations upon the Nervous System of the
Common Centipede, hj Mr. John Ryder.
Feb. 2d, 1880. — Observations upon a specimen of Actinoptae-
riuvi, by Mr. Edward Potts.
Feb. IGth, 1880. — The Mounting of Microscopic Objects, by Dr.
Seller.
Feb. 16th, 1880. — Description of Rhipidodendroyi and Halteria,
b^'- Mr. John Ryder.
il4 PROCEEDINGS OP THE ACADEMY OF [1880.
March 1st, 1880. — Injecting and Special Methods of mounting
Microscopical Objects, by Dr. Carl Seller.
March 1st, 1880. — Observations upon Sponges, by Mr. John
Eyder.
March 1st, 1880. — A Plan to show Opaque Objects with the Gas
Microscope, by Persifor Frazer.
March 15th, 1880. — Lithological Studies with the Microscope,
by Persifor Frazer.
April 5th, 1880.— Histological Studies, by Dr. Seller.
April 19th, 1880. — Lantern Exhibition, by Mr. Holman and
Mr. Ryder.
May .3d, 1880. — Communication upon Fresh-water Sponges, by
Mr. E. Potts.
May llth, 1880. — Communication upon the Eggs of the matica,
by Mr. E. Potts.
Sept. Gth, 1880.— Life Forms at Atlantic City, by Mr. E. Potts.
Sept. 20th, 1880 — Communication upon the Larvis of King
Crabs, by Mr. E. Potts.
Oct. 14th and 15th, 1880.— Annual Exhibition.
Nov, 1st, 1880. — Report of the Committee on Exhibits and Im-
provements in Microscopical Science at the Annual Exhibition,
by Dr. Hunt.
Nov. 15th, 1880. — Communication upon the Development of the
Pyrulla, by Mr. Charles Perot.
The following Members and Associates were elected during the
year.
Memhei's: — John C. Wilson, Otto Luthy, Howard Kelly.
Associates: — Dr. Joseph Simsohn, Dr. James A. McCoart, Dr.
Edward T. Bruen, Dr. John W. Gadsden, Dr. Monroe Bond, Dr.
J. H. Wills,
RoBT. J. Hess,
Recorder.
REPORT OF THE CONCHOLOGICAL SECTION.
The Recorder of the Conchological Section respectfully reports
that during 1880, Dr. R. Bergh, Dr. W. D. Hartman, and Mr.
Angelo Heilprin have presented papers upon the Mollusca, which
have been accepted and published in the Academy's Proceedings.
18 so.] NATURAL SCIENCES OF PHILADELPUIA. 415
The Section has again lost a valued member, Professor S. S.
Haldenian, who died September 10th, 1880. Professor Halderaan
manifested his interest by frequent contributions to our Museum,
as well as by papers published in the American Journal of Con-
cholog}'. He also presented to us a number of copies of text and
plates of his celebrated monograph upon the Fresh-water Univalve
Mollusca of the United States. These the Section reissued, the
work having been long out of print, and the sale resulted to its
pecuniary advantage.
Mr. George W. Try on, Jr., Conservator of the Section, reports
as follows :
About fifty distinct donations and purchases of recent shells
will be found recorded in the detailed list hereunto appended,
aggregating 1216 species, represented by 4574 specimens. These
have all been carefully determined, labeled, mounted, and placed
in the cases.
Mr. Charles F. Parker has, as usual, afforded valuable assistance
in preparing these specimens for exhibition.
Mr. John Ford continues to prepare for us sections of univalve
shells, showing their internal form and structure. He has pi*e-
sented over fifty of these during the year. We are indebted to
Miss Anna T. Jeanes for a number of beautiful glass models of
mollusks, and to Mr. Joseph Jeanes for a fine suite of California
shells, and mounted Unguals of Ghitonidse, etc.
Mr. John H. Redfield has presented his entire and very com-
plete collection of Mar-gineUidae.
The U. S. Fish Commission, and Dr W. H. Jones, U. S. N.,
have presented numerous specimens, both in alcohol and dry.
We have received from Dr. Isaac Lea, the tjq^e series of
Claiborne (Ala.) Eocene fossils, described and figured in his
'• Contributions to Geology," numbering 228 species.
Mr. John A. Ryder has prepared a drawing in outline of the
gigantic Architeuthis princeps, Yerrill ; although onl^^ j^g of the
natural size, this drawing is upon a canvas twelve feet in length.
It is exhibited upon the wall of the Conchological gallery'.
Our collection of fossil shells, the systematic arrangement of
which has been so long neglected, has at length, under the com-
petent supervision of Mr. Angelo Heilprin, received that attention
which its importance merits. Mr. Heilprin has critically studied
416 PROCEEDINCxS OF THE ACADEMY OF [1880.
and arranged the whole of the North American Eocene Collection,
and is now engaged upon the Miocene. 'These shells have all been
labeled and mounted by Mr. Parker. A suitable label has been
placed upon the drawers containing the " Swift Collection," and
these have been made accessible to the public. The Cephalopoda^
Muricidse, Purpuridse, Fusidag, and Buccinidae of the general
collection have been rearranged in accordance with the latest
information upon these groups ; and it is proposed to continue
this work of revision upon the other families of marine shells as
opportunity offers.
A rearrangement of the Land shells in accordance with the
natural groups of Dr. Louis Pfeiffer's " Nomenclator Heliceorum
Viventium," and of the ITnionidse, in accordance with the latest
edition of Dr. Lea's " Synopsis," will be commenced as soon as
possible.
The Museum of Recent Conchology now contains 38,624 trays
and 136,387 specimens.
There have been no changes made in the Bj'-Laws of the Section.
The officers for 1881 are :
Director — W. S. W. "Ruschenberoer.
Vice-Director — John Ford.
Recorder — S. Raymond Roberts.
Secretary — John H. Redfield.
Treasurer — Wm. L. Mactier.
Conservator — Geo. W. Tryon, Jr.
Lihrarian — Edw. J. Nolan.
Respectfully submitted,
S. Raymokd Roberts,
Recorder.
The following are the additions to the Conchological Cabinet
received during 1880 ■:
R. Arango. Two hundred and sevent3'-four species and A'arie-
ties of Cuban shells.
W. G. Binney. Helix Ifayrani, Algiers. Testacella haliotoidea,
England.
John Brazier. Bythinia hyalina, from New South Wales.
Eighty -four species Land, Fresh- water, and Marine shells from
Anstralia.
J. J. Brown. A collection of mollusks from Florida and Haiti.
W. W. Calkins. Unio Blandingianus and Tritonidea t^'ncta,
Florida.
1880.] NATURAL SCIENCES OF PHILADELPHIA. 417
Caleb Cooke. Melania scabra^ Zanzibar.
W. H. Doughert}-. • BuUmm^ Schiedeanus, Coaliuila, Mexico.
John Ford. Arcn pexata^ Say, Newport, R. I. Five species
of marine shells, Atlantic City, N. J. Mytihi>t hamolns, Say, from
Seekonk River, Providence, R. I. Helix tuberculoma Conrad,
Sinaitic Desert. Natica duplicata and N. heros, with nidus, ova
capsules of Nassa frivUtata, Atlantic City, X. J. Fine specimen
of Cassis tuberosa, Bahamas. Over fifty specimens, sections of
shells.
Andrew Garrett. Pariula decussafa and P. ganymedes. Domi-
nique Is., Marquesas. P. inflata, Taiwata, Marquesas. Partula
(sp.), Moorea, Society Isles. Trochus trochoidea, Society Isles,
Gai^dium (sp.), Paumotus Is.
E. Hall. A collection of land and fresh-water shells from
various localities. Twenty-six species of fresh-water shells.
Dr. W. D. Hartman. Embryonic Partulae. Cijclostoma in-
eomptas, near Bogota, S. A. Helix simila7-is, Fer., Japan. Three
species of Paiiula from Marquesas Islands. Partula Raiatensis
(type) from Raiatea. Partula ajyproximata, Raiatea.
Henry Hemphill. Over two hundred species and varieties of
California shells.
J. Gr. Hidalgo. Murex Tryoni (type), Lesser Antilles. Pici-
nula nodosa, Brazil.
Anna T. Jeanes. Glass models of twelve species of nudibran- •
chiate mollusks.
Joseph Jeanes. Mounted linguals of thirty-seven species of
mollusks. Ninety-nine species and varieties of land, fresh-water
and marine shells from California.
Dr. W. H. Jones. Twenty-five species of pelagic mollusks from
the Atlantic and Pacific Oceans. Scyllsea pelagica. Oxygyrus
Keraudrenii.
Henry C. Lea. Twenty -four species of Claiborne Eocene fossil
shells, types of his descriptions.
Dr. Isaac Lea. Type collection of Claiborne (Ala), Eocene
shells, consisting of 228 species ; arranged as described and figured
in his " Contributions to Geology." Voluta Junonia, Edgmont
Key, Fla.
Joseph Leidy. Goniobasis proxima, Say, Piedmont Springs,
North Carolina
E. T. Nelson, Eupleura Tampaensis, Conr., Tampa Bay, Fla.
T. R. Peak*. Astarte castanea^ Say, Sandy Hook, N. j.
John H. Redfield. Collection of Marginellidj^, including several
hundred specimens of about two hundred species and varieties.
Spirifer mucronatus, Hamilton group.
S. R. Roberts. Macoma balthica, Linn., Collins' Beach, Del.
Helix cinnamomea. Trima pellucidula, Sandwich Isles. Cylin-
drella gracilicollis, Macroceramus Klatteanus, Bid. (ex auct.),
Port-au-Prince, Hayti.
28
418 PROCEEDINGS OF THE ACADEMY OF [1880.
Dr. W S. W. Riischenberger, Dione lupinaria,, San Bias.
Prof. D. S. Sheldon. Physa gyrina, yonng, Davenport, Iowa.
Hon. F. E. Spinner. Seven lots of marine shells (fifty -six spe-
cies) from the month of St. John's River. Fla.
U. S. Fish Commission. Fifty-fonr species of marine MoUusca
from the New England coast,
H. A. Ward (purchased). Glass models of six species of
Cephalopods.
J. F. Whiteaves. Eight species of fluviatile and marine sliells,
from Queen Charlotte's Isl. and Gulf of St. Lawrence.
REPORT OF THE BOTANICAL SECTION.
The Vice-Director takes much pleasure in reporting to the
Academy the continued prosperity of the botanical department,
which, in fact, is qnite equal to all that can be expected of it,
until, by the good fortune of an endowment, funds can be supplied
regularly to extend its work. Meetings have been held every
month throughout the year except July and August, at each of
which valuable communications have been made by various mem-
bers. Some of the more important of these have been communi-
cated to the general meetings of the Academy, and have found a
place in its published proceedings.
The Section now consists of thirty-one members, one having
been added during the year.
The Conservator's report to the Section of the condition of the
Herbarium has been adopted by tlie Section as its report to tlie
Academy, and is as follows : —
The accessions to the Academy's Herbarium during the past
year have been large and valuable, and the mounting and distri-
bution of the plants received have made large demands upon the
time of the Conservator and of the members of the Section who
have kindly aided him.
Among the valuable contributions received were the collections
of the late Dr. Charles Pickering, made in the years 1844-5, during
a journey to Malta, Egypt, Arabia and India, and presented to the
Academy by his widow, Mrs. Sarah S. Pickering. The number
of species is estimated at about 1500, and, as none of them were
named, the labor of distributing them in their proper natural
1880.] NATURAL SCIENCES OF PHILADELPHIA. 419
order (which has been performed by Messrs. Meehan and Biirk),
has been slow, but is now completed.
From Di-. Gray, of Cambridge, important contributions have
been received. Dr. Garber has presented a collection of 100
species collected by himself recently in Porto Rico, while some
'i.'SO species of Mexican plants, collected b}' Drs; Parr}- and Palmer,
have been added.
In the department of the Lower Cryptogams the additions have
been of a nearly complete series of the Mosses and Hepaticae of
X. America, collected and named by the late lamented Austin, and
presented by the liberalit}^ of members of the Section, and of 200
species of N. American Fungi, collected, determined and pre-
pared by J. B. Ellis, and presented l)y Mr. Martindale.
The total number of species contributed during the year is
estimated at 3100, a very large proportion of them being new to
the Herbarium.
Some progress has been made in the mounting of the Noi'th
American Herbarium, the orders from Polemoniacea? to Scrophu-
lariacej^} inclusive having been completed. Mr. Scriliner has con-
tinued his work upon the determination and the mounting of the
Grasses, though interrupted by long and serious illness. Those
familiar with Herbarium work can appreciate the amount of labor
yet required to sift the material now upon our shelves, to deter-
mine the doubtful species and to mount the whole.* Volunteer
labor is hardh' adequate to take care of the new accessions, and,
until some endowment shall secure to the Academy the constant
work of a competent botanist, the completion of the task must
remain for the future.
Much inconvenience has been heretofore experienced from the
want of a proper place to receive and display such seed-vessels
and vegetable products as were too large to place in the herbarium
sheets. Such objects have necessarily been placed in the gallery
of the Museum, too distant from the botanical working-room to
be readily consulted. The liberalitj' of a member of the Acad-
emy, whose aid has often supplemented its needs and sustained
the hands of its workers, has removed this difficulty, and we have
now in the botanical room a most convenient and capacious work-
ing table, containing sixty-four large drawers for the reception of
seed-vessels, pine-cones, wood-sections, etc.
The Conservator must acknowledge, as heretofore, tlie efficient
420 PROCEEDINGS OF THE ACADEMY OF [1880.
aid received from Mr. Chas. F. Parker in the work of poisoning
and arranging the collections received during the year and for
other material assistance.
John H. Redpield,
Conservator.
December \Wi, 1880.
The officers elected for the forthcomino- year are :
Director. — Dr. W. S. AV. Ruschenberger.
Vice-Director. — Thomas Meehan.
Recorder. — F. L. Scribner.
Gor. Secretary. — Isaac C. Martindale.
Conservator, — John H. Redfleld.
Treasurer. — J. 0. Schimmel.
Respectfully submitted,
Thomas Meehan,
Vice-Director,
Donations to Herbarium and Museum. — Mrs. Sarah S. Pickering,
of Cambridge, Mass. : 1200 species plants, collected by the late
Dr. Chas. Pickering, in the years 1844 and 1845, in Malta, Egypt,
Arabia, Zanzibar and India, also lot of seed-vessels, etc. Chas.
F.Parker: Lechea Novae Gesarise Austin, Bergen Co., N. Y.,
(author's type) ; Fracjaria Gillmani, Clinton, Detroit, Mich,
(author's type) ; Guizotia oleifera., D. C, African species, from
Ballast, Camden, N, J. ; Lycopus sessilifolius, Gr., Batsto, N. J. ;
Lycopus Europseus, Ballast, Camden, N. J. Wm, M. Canb}- :
Phlox Stellaria^ Gr., Nashville, Tenn. ; 5 species plants from Cali-
fornia, new to the collection ; 111 species of plants from Europe,
Syria, S. Africa, etc., many of them new to the collection. Dr.
Asa Gray: 161 species plants from California. Arizona, Oregon,
Washington Terr., Turkistan, and Micronesian Islands, mostly
new to the collection. Prof C, J. Sargent : Aster Herveyi, Gr.,
Tiverton, R. I. ; Photographs of Conifene, from Oregon. Geo. E.
Davenport, Boston : Gheilanthes viscida, Davenp., California.
Isaac C. Martindale : Third and fourth centuries of Ellis' North
American Fungi; Gorethrogyne filaginifolia, Nutt, San Diego Co.,
Cal. ; specimens of Oas/a ??pa re.sca, L., var. ^»;eKcana, with ab-
normal fertile spikes, from Pitman's Grove, N. J. ; Bark of Pinus
Tnitis.! Mx. ; Brickellia VincenUana.^ Greene, new species. New
Mexico ; Gorrigiola litteralis^ L., Ballast, near Philadelphia. Dr.
C. C. Parry, l)aven])ort, la. : Tithonia tubseformiSj Cass., cidt. at
Davenport, from Mexican seed ; Mexican mats and rope made
from fibre of ^f/aue heteracantha ; Fibre of Agave Americana.
A. L. Siler, Utah : Pentstemon Sileri^ Gr., nov. sp., Beaver Dam
Mts., Utah. Mrs, M. J. Myers, Syracuse, N. Y. : Epipactis Hel-
1880.] NATURAL SCIENCES OF PHILADELPHIA, 421
feborine Rich., var, viridans Gr., near S^yracuse, N. Y. Dr. Thos.
Lancaster : Poli/porus, growing on Hemlock Spruce, McKean
Co., Pa. Wm. H. Dougherty : Skeleton of stem of Opuntia im-
hricata; Capsule of Hanhuria; Stems of Euj^horbia anfis{/j:)hiUtica,
all from Chihuahua, Mexico. Aubrey H. Smith : MijoHotw versi-
color, Pers., Cobb's Creek, Darby, Pa. Thomas Bland, N. York :
Wax, from the leaves of the Carnauba Palm ( Copernicia ceriferay,
Whip from the bark of , Jamaica ; Paper from the fibre of
Plantain and Banana, Jamaica; llyri.stica fra grans, Houtt., Nut-
meg with its Mace and outer husk, Jamaica ; Seeds of the follow-
ing plants from West Indies : Lucuma mammosa, Gris., Feuillea
cordifolia, Ser., Coix lachryma, L., Sajnndus sajwnaria, L.,
Adenanthera paronina, L., Abi^us precatorius, L., GuUandina
Bonduc, L., Anacardium occidentale, L., GuUandina BonduceUa,
li. ; Wood of the Down tree {Ochroma Lagopus). Thos. Meehan :
Pellaea at ropurpurea^ Link., collected in Southern Utah, by A.
L. Siler ; Sedum Meehani, Gra}^ collected in Southern Utah, by
A. D. Siler. Isaac Burk : Wood of Herifiera, Africa. Dr. A. P.
Garber: 95 species of plants, collected by him at Yaueo, Porto
Rico, in 1880, named by' Dr. Gray and Mr. Oliver of Kentucky.
John H. Redfield : 217 species plants collected by Dr. C. C. Parry
and Dr. E. Palmer, near San Luis Potosi, Mexico, named at Kew
and Cambridge ; 57 species Ferns, collected by A. Fendler, in the
Island of Trinidad, W. I., in 1879-80, supplementary to a collec-
tion presented in 1878, named by Prof. D. C. Eaton; 7 species
plants from Florida and Tropical America. J. H. Redfield, J. C.
Martindale. Thos. Meehan, Wm. M. Canby, Dr. Chas. E. Schaffer,
Dr. J. Bernard Brinton : Complete set of the Mosses and Ilepaticifi
of N. America, collected and named by the late Coe F. Austin,
consisting of 518 species of Mosses and 124 species of Hepaticae.
REPORT OF THE ENTOMOLOGICAL SECTION.
Tiie Entomological Section, throughout the past j-ear has held
its regular meetings, excepting during the months of July and
August. Most of the meetings have been quite interesting, owing
to the man}' original communications, both verbal and v/ritten,
that have been presented. These lectures, as they may be called,
were devoted to illustrating some of the variations of form, etc.,
of many entomological species and genera. At the same time
opportunity has been aff"orded to those so desiring, to describe
new species of the entomological fauna, and to present their ob-
servations to the public. That the claims of priority of description
thus made, might not be lost by the several authors, such com-
munications have been condensed and published in the Proceedings
of the Section. The more full and complete papers are, as hereto-
422 PROCEEDINGS OF THE ACADEMY OF [1880.
fore, published in the Transactions of the American Entomologi-
cal Society. There have been eight pa})ers presented and pulilished
by the latter society during the 3'ear, comprising 338 pages of
printed matter in octavo form, illustrated by seven plates. This,
in connection with 24 pages of the published Proceedings of the
Section make a total of 362 pages of entomological publications
issued since last annual meeting.
The entomological collections of the Academy have been care-
fully attended to through the year, by the Conservator, Mr. Geo.
B. Cresson, and have been preserved from all infection or loss.
By the death of Mr. James Ridings, in Jul}-, the section lost
one of its most valued members. Mr. Ridings was one of the
founders of the American Entomological Society, and thi'ough his
many valuable discoA^eries was well known among the entomolo-
gists of the United States.
At the annual meeting of the section, held December 13th, the
following gentlemen were re-elected as officers for the ensuing year:
Director. — John L. LeConte, M. D.
Vice-Director. — George H. Horn, M. D.
Treasurer. — E. T. Cresson.
Recorder. — J. H. Ridings.
Conservator. — Geo. B. Cresson.
Publication Committee. — George H. Horn, M. P.
Samuel Lewis, M. I>.
At the last annual meeting of the American Entomological
Society the sum of seventy-five dollars was contributed towards
the funds of the Academy-.
Respectfully submitted,
James H. Ridings,
Recorder.
REPORT OF MTNERALOGICAL AND GEOLOGICAL
SECTIOX.
The Director of the Mineralogical and Geological Section would
respectfully report :
Meetings of the Section have been held monthly, except during
Jul}" and August. The attendance has been good. A number of
interesting papers were read, and man}' valuable communications
and donations made. During the year, the first volume of its pro-
ceedings was published, containing scientific papers and commu-
nications to January 1st, 1880. The collection of local rocks and
\
1880 J NATURAL SCIENCES OF PHILADELPHIA. 423
minerals has outgrown the i)hice provided for it. It is almost
complete as to the rocks of Philadelphia, and of Delaware, Mont-
gomery and Bucks counties. Believing, as he does, that this
collection will grow into one of great importance and interest, he
is glad to state that the desire of the Section for a better location
for it has been granted b}* the Council of the Academy.
Resi'>POtfully submitted,
Theo. D. Rand,
Director.
Philadelphia, December 27th, 1880.
To the Director of the Mineralogical and Geological Section :
The mineral collection of the Academy has been improved
during the past year by the addition of the usual number of dona-
tions. These have been carefully labeled and placed in the cases
b}^ Mr. Charles F. Parker, to whose industry and care we are
chiefly indebted for the satisfactory arrangement and labeling of
all our specimens. I submit with this a list of the donations
during the past year. The collection is in a satisfactory con-
dition.
Joseph Willcox,
Conservator.
Addition!< to Mineralogical Cabinet during the year 1880: —
Jas. W. Beath : Twenty -five specimens of polished Agates, from
Oberstein, Germany, and Paraguay ; Crocidollte, S. Africa. C. S.
Boutcher : Proustite, Gunnison, Co., Colorado. Walter Collins :
Asphaltum, Cretaceous Marl, Blackwoodtown, N, J. Chas. Doble :
Millerite, Chalcopyrite and Niccoliferous P^a-rhotite, Gap Mine,
Lancaster Co., Pa. W. 11. Dougherty : Xative Gold, also a fine col-
lection of Xative Silver, Silver ores, Argentiferous Galena, Ruby
Silver, Cassiterite, etc., Mexico; Green Sand, San Antonio River,
Texas. John Ford : Stilbite, Frankford. Philada. : Actinolite,
Hornblende, Lafayette. Montgomery Co., Pa, John Garvin:
Native Gold in Quartz, Battle Branch, Ga. E. Goldsmith: Lignite,
containing Fichtelite, Brazil Prof. S. S. Haldeman : Stalactite,
and six specimens of Agates, Argentine Republic. E. P. Hancock:
Two specimens Jetfersonite, Sterling, Sussex Co., N.J. ; Thorite,
Brevig, Norwa^^ W. W. Jefferis: Quartz pseud, after Dog-tooth
Spar; Picrolite (Slickenside,) Newlin, Chester Co., Pa ; AVavellite,
E. Whiteland, Chester Co., Pa. Dr. G. A. Koenig : Jarosite,
Chattee Co., Colorado. Dr. Isaac Lea : Amazonstone, and a fine
specimen of Sunstone, near Media, Del. Co., Pa. Dr. Jos. Leidy :
Three specimens of Talcose Slate, Soapstone Quarry, shore of the
Delaware River, above Easton, Pa. ; Corundum, Laurens Co., S. C. ;
Biotite, Steatite Quarry on Bushkill Creek, near Easton, Pa. H.
C. Lewis: Philadelphite, Phila. ; Hyalite, Germantown, Pliila. ;
424 PROCEEDINGS OF THE ACADEMY OF [1880.
Halite, Saltville, Va. Wm. Lorenz : Chrysotile, Canada. Mr.
Loyer : Corundum, Chester Co., Pa. Miss Miller: Crystals of
Silver, Lake Superior; Hematite, Gypsum, Stalactite, Cliloras-
trolite, Halite, etc., from various localities. Dr. Weir Mitcliell :
Silicified Wood, Missouri River, above Bismarck. L. Palmer:
Albite,Vermiculite,Del. Co.,Pa. Theo. D. Rand : Kammererite and
Chromite, Radnor, Del. Co., Pa. ; Crystallized Quartz in Potsdam
Sandstone, Mont. Co., Pa. ; Herrengrundite, Herrengrund, Hun-
gary ; Orileyite, Burmali. J. L. Reed : Asbestus, Italy ; Clirys-
otile, Ontario, Canada. T. W. Ried : Chalcopyrite, Montgomery
Co., Pa. Dr. W. S. W. Ruschenberger : Copper Slag, Caldera,
Chile, 1856. Dr. J. Richard Taylor: Cerargyrite, Chloride of
Silver, with fractured Wavellite crystals. Galena with free Sulphur,
Millerite, and argentiferous Carbonate of Lead, Leadville, Colo-
rado ; Ore from the Ohio Mine, bearing Gold, Silver and Copper,
Breckenridge, Col. C. M. Wheatley : Fine specimen of Byssolite,
Chester Co., Pa. ; Azurite on Chalcopyrite, Upper Salford Mine,
Montgomery Co., Pa. ; Aurichalcite on Calcite, and white Apatite
Avith Byssolite ; Chalcopyrite, Pyrite and Melaconite, Jones Mine,
Berks Co., Pa. Dr. Jas. W. White : Corundum, Zircon, Storeville,
Anderson Co., S. C ; Corundum, Concord, N. C. ; Andesite with
Corundum, Hog-back Mt , N. C. ; a collection consisting of Zincite,
Rutile, Phlogopite, Graphic Granite, Chesterlite, Fibrolite, Garnet,
Damourite, Stalagmite, etc., from various localities. Joseph
Willcox: Autunite (Uranite), Mitchell Co., N. C ; Pyroxene,
Biotite ?, Apatite, Burgess, Ontario, Canada ; Corundum coated
with Margarite, Iredell Co., N. C. ; Danburite, Russel, St. Law-
rence 00.,"^". Y. ; Black Tourmaline, Westport. Ontario, Canada;
Scapolite, Pyroxene, and four specimens of Apatite, Bob Lake,
Ontario, Canada. A. E. Foote, in exchange for duplicate books :
nine specimens of Apatite, Renfrew, Ontario, Canada; Chrysotile,
four specimens of Titanite (Spliene), Titanite (Lederite), two
Yesuvianite, Beryl, Triphyllite, Celestite, Wollastonite, Gummite,
Uranotile, Tourmaline, Octahedral Crystals of Fluorite, with
Apatite and Calcite, from various localities. Purchased : Limon-
ite, Superior Mine, Michigan.
Additions to Bock CoUeetion.—i o\m Ford: Hornblende, Soap-
stone Quarry, Lafayette Pa. ; Tourmaline and Hornblende Schist,
Tunnel near Girard Ave. Bridge ; Decomposing Gneiss with Mica,
ditto with Quartz, ditto with Manganese?, near west end of Cal-
lowhill St. Bridge, Philadelphia. G. H. Ivens : Geode of Limonite,
Kent Co., Md. W. W. Jetferis : Gneiss. John Hartman : Two
specimens of Crystalline Slag, taken from hearth of Blast Furnace,
Charlotte, N. Y. Dr. Jos. Leidy : Talcose Slate, Soapstone
Quarry, Pot Rock, Delaware River, above Easton, Pa. ; Indurated
Clay (Bridger Eocene), near F'ort Bridger, Wyoming. H. C.
Lewis : Glacier-scratched boulder, Belvidere, N. J. Theo. D.
Rand : Twenty-seven specimens of Rocks, from the neighborhood
of Philadelphia, for Local Rock collection; three specimens of
1880.] NATURAL SCIENCES OF PHILADELPHIA. 425
slags, from Puddling Furnace, Coatesville, Pa. Joseph H. TuU :
Six specimens of Rul)y Silver, near Austin, Nevada.
SUMMARY OF THE REPORT OF WM. C. HENSZEY,
Treasurer, for the year ending Nov. 30, 1880.
Dr.
To Balance from last account $1032 99
" Initiation fees 230 00
" Contributions (semi-annual contributions) 2244 98
" Life Memberships ; 500 00
" Voluntary Contributions from Life Members 615 00
" Admissions to Museum 455 30
" Sale of Guide to Museum 62 00
" " Duplicate Books 7 75
" Donation from Mineralogical and G. Section towards
Proceedings 35 00
" Donations towards Plates for Proceedings 10 00
" Interest on Deposits 69 04
" Interest on Phil, and Erie Railroad Bonds 30 00
" Life Member Fund. Interest on Investment 120 GO
" Maintenance Fund. " " " 30 00
" Publication Committee. W. S. Vaux, Treasurer 507 04
" Publication Fund. Interest on Investments 280 00
" Barton Fund. " " " 240 00
" Wilson Fund. Towards Salary Librarian 300 00
" Freight returned 4 30
" Phila. and Erie Railroad Bond, Transferred to Mainte-
tenance Fund 1000 00
$7763 40
Cr.
Salaries, Janitors, etc $2960 00
Freight 60 59
Inspecting Boiler 10 20
Repairs 188 05
Insurance
Jars and Bottles
Coal
Gas
Mounting Bird
Stationery and Postage Stamps 136 55
Books
Alcohol :
Publication Committee. W. S. Vaux, Treasurer
Newspaper Reports
Water Rents
Trays
Binding
Printing and Paper • 1539 63
Plates and Printing 142 52
Miscellaneous 448 54
Life Memberships transferred to Life Membership Fund. 500 00
30
00
74
11
195
50
177
27
1
25
136
55
6
50
37
60
93
33
64 00
26
15
42
00
118
40
6852 19
Balance^ $911 21
^ During the year there was received from voluntary contributions and
donations $660, which, with a Bond for $1000, used for general purposes,
indicates that the current expenses exceed the regular income over $1600, the
balance at the close of the year being a little less than at the commencement.
426 PROCEEDINGS OF THE ACADEMY OF [1880.
LIFE MEMBERSHIP FUND. (For Maintenance.)
Balance per last Statement $500 00
Life Memberships Transferred to this account 500 00
Interest 120 00
Sill20 00
Transferred to General Account 120 00
To Balance for Investment $1000 00
BARTON FUND. (For Printing and Illustrating Publications.)
Balance per last Statement $240 00
Interest 240 00
$480 00
Transferred to General Account 240 00
Balance $240 00
JESSUP FUND. (For Support of Students.)
Balance last Statement $551 67
Interest on Investments 560 00
$1111 67
Disbursed 590 00
Balance $521 67
MAINTENANCE FUND.
Total amount received $1550 00
Interest 30 00
Less paid for Printing ,
Invested in Bonds Phila. and Erie Railroad.
Interest Transferred to General Account
$1580
00
$23 65
1000 00
30 00
1053
65
Balance $526 35
I. V. WILLIAMSON LIBRARY FUND.
Balance $330 26
Rents Collected 97 00
Ground-rents Collected 1096 00
$1523 32
For Books ;«^577 56
Expenses Sale of Prop'ty for arrearages of Ground-rent... 167 25
Costs, Insurances, etc
Repairs to Properties
Taxes
Water Rents
Collecting
Balance $209 55
37 47
232 37
204 02
41 40
53 70
1313 77
1880.J NATURAL SCIENCES OF PHILADELPHIA. 427
PUBLICATION FUND.
Balance last Statement $:->38 25
Income from Investments 350 00
$688 25
Transferred to General Account 280 00
To Balance $408 25
THOMAS B. WILSON LIBRARY FUND.
Balance last Statement $141 57
Interests on Investments 570 00
Received from W. S. Vaux for Duplicate Books 8 00
$719 57
Paid for Books $300 00
" Binding 11 55
To General Account towards salary of Librarian oOO 00
$611 55
Balance $108 02
MRS. STOTT FUND. (For Publication.)
Twelve Months' Interest... $112 00
Paid to (W. S. Vaux) Publication Committee 114 00
JOSHUA T. .JEANES FUND. (For Maintenance.)
Bequest by him paid by Heirs $20,000 00
Invested in three Mortgages $7000 00
3000 00
10,000 00
$20,000 00
ECKFELT FUND.
To be invested $2466 86
428
PROCEEDINGS OF ACADEMY OF NATURAL SCIENCES. [1880.
The election of officers for
result : —
President,
Vice-Presidents,
Recording Secretary,
Corresponding Secretary,
Treasurer,
Librarian,
Curators,
Councillors to serve three
years, ....
Finance Committee,
1881 Avas held, with the following
W. S. W. Ruschenberger, M. D.
Wm. S. Vaux,
Thomas Meehan.
Edward J. Nolan, M. P.
Geo. H. Horn, M. D.
Wm. C. Henszey.
Edward J. Nolan, M. D.
Joseph Leidy, M. D.,
Wm. S. Vaux,
Chas. F. Parker,
R. S. Kenderdine, M. D.
Wm. L. Mactier,
Aubrey H. Smith,
Henry C. Chapman, M. D.,
Geo. y. Shoemaker.
Edw. S. Whelen,
Clarence S. Bement,
Aubre}^ H. Smith,
S. Fisher Corlies,
Geo. Y. Shoemaker.
ELECTIONS DURING 1880.
MEMBERS.
January 27. — A. R. Thomas, M. D., Wm. H. Jenks, John S.
Jenks, Chas. W. Pickering, Henry F. Formad, M. D., Charles P.
Tasker, John Wagner, Ferris W. Price, Geo. W. Biddle.
February 2^. — R. S. Huidekoper, M. D., Frances Emily White,
M. D., David Townsend, Thos. Miles, John S. Capp.
3Iarch SO, — Paris Haldeman, Geo. B. Heckel, Emlen Physick,
M. D.
May ^5.— Henry S. Gratz, R. S. Peabody, Mrs. R. S. Peabody,
Wm. Barbeck.
October 26.—Rq\. Wm. F. C. Morsell, Samuel R. Knight, M. Di
November 30. — Charles S. Turnbull, M. D., James M. Anders,
M. D.
For list of Correspondents elected see Report of the Corre-
sponding Secretary.
ADDITIONS TO LIBRARY. 429
ADDITIONS TO LIBRARY.
December 1st, 1879— November 30th, 18S0.
Abbot, E, H. Physics and hydraulics of the Mississippi River. Dr. I.
Minis Hays.
Allen, J. A. U. S. Geol and Geogr. Surv. of Ter. Miscellaneous publica-
tions, No. 12. History of North American Pinnipeds, 1880. The
Department of the Interior.
Allen, T. F. Charace* of America. Pts. 1 and 2. I. V. Williamson Fund.
AUis, O. A. Deformity from fractures at the lower end of the humerus. The
Author.
Alumni Association, Philadelphia College of Pharmacy. 16th annual report.
The Society.
American Angler's Guide. 3d Ed., 1849. S S. Haldeman.
American Museum of Natural History. 11th annual report. The Director.
Annual record of science and industry for 1878. I. V. Williamson Fund.
Arango, R. Contribucion a la fauna malacologica Cubana. The Author.
Archajological Section of the St. Louis Academy of Sciences. Contributions
to the archeology of Missouri. Part I, Pottery. The Academy.
Astor Library. 31st annual report, 1880. The Trustees.
Baillon, M. H Dictionnaire de botanique. 12me Fasc.
Natural history of plants. Vol. 6. I. V. Williamson Fund.
Balfour, F. M. Comparative embryology. I. V. Williamson Fund.
Barber, E. A. Comparative vocabulary of Utah dialects. The Author.
Biircena, Mariano. Terremoto del 17 de Maj o de 1879
Viaje a la Caverna de Cacahuamilpa, 1874. The Author.
Datos para el esludio de las rocas mesozoicas de Mexico, y sus fosiles
caracteristicos, 187-5. Di*. .Jos. Leidy.
Barrande, J. Brachiopodes. Vol. 5, 1879. The Author.
Beale, L. S. How to work with the microscope. 5th Ed. I. V. Williamson
Fund.
Bentham, G. and J. D. Hooker. Genera plantarum. Vol. 3, Pt. I. I. \ .
Williamson Fund.
Berg, Dr. C. Observaciones acerca de la familia Hyponomeutidre.
Apuntes Lepidopterologiques.
Hemiptera argentina, 1879.
La Reina de las Flores, 1880. The Author.
Binney, W. G. North American species of Zonites. The Author.
Board of Directors of City Trusts. 10th annual report. The Board.
Bocage, J. V. Barboza de. Melanges ornithologiques, V.
Liste des Antilopes d' Angola.
Algumas observa^oes e additamentos ao artigo do Sr. A. C. Smith intitu-
lado "A sketch of the birds of Portugal."
Note sur une nouvelle espece africaine du genre " Coracias."
Aves das possesoes portuguezas d'Africa occidental, 14th and 16th List.
Subsidies para a fauna das pos. portuguezas d'Africa occidental.
Aves da Zambezia e do Transvaal. The Author.
Bohnensieg, G. C. W. and W. Burck. Repertorium annuum literatune botan-
icte periodicoe. T. 6, 1879.
Encyklopedie der Naturwissenschaften. le Abth., 8 Lief., 1880. L V.
Williamson Fund.
Bolton, H. Carrington. Table showing the behavior of certain minerals with
Citric Acid alone and with reagents. The Author.
Bommer, .J. E. Monographie de la classe des Fougeres, 1867. Dr. F. V.
Hayden.
430 ADDITIONS TO LIBRARY.
Borre, A. Preudhomme de. Espcces de la tribu des Feronides qui se rencon-
ti-ent en Belgique. Ire Partie.
Kspeces des tribus des Panageides, des Loricerides, des Licindes, des
Chlwniides et des Broscides (jui se rencontrent en Belgique.
De la meilleure disposition il donner aux caisses et cartons des collections
d'insects
Note sur le Breyeria Borinensis. The Author.
Brazier, J. Synonyms of, and remarks upon Port Jackson, New Caledonian
and other shells, with their distribution.
Brief account of the natives of western Australia, 1879. The Author.
Brehms Thierleben. 8 Bd., 1-8 Heft. I. V. Williamson Fund.
Breviere, L. Catalogue des mollusques observes dans le Department de la
Nievre. I. V. Williamson Fund.
Bronn, H. G. Morphologische Studien iiber die Gestaltung-Gesetze der Na-
turkorper iiberhaupt, und der organischen insbesondere. 1858. Dr.
.Jos. Leidy.
Thier-Reichs. 6er Bd., III. Abth., 2-12 Lief. Wilson Fund.
Bruhl, C. B. Zootomie aller Thierklassen. Atlas. Lief. 14 & 15. I. V.
Williamson Fund.
Brun, J. Diatomees des Alpes et du .Jura. I. V. Williamson Fund.
Brush, Geo. J. Mineral locality at Bi-anchville, Connecticut. 4th paper.
The Author.
Buchanan, J. Manual of indigenous grasses of New Zealand. Wellington,
1880. Geological Survey of New Zealand.
Bureau of Education, circulars of information, Nos. 2 and 3. Department of
the Interior.
Bureau of Statistics, Treasury Department, quarterly reports, Sept. 30, 1879-
June 20, 1880. The Department
Butler, A. G. Illustrations of typical specimens of Lepidoptera heterocera in
the collection of the British Museum. Pt. 14. The British Museum.
Calkins, W. W. American conchology. Catalogue of Uniones. 1880. The
Author.
Cameletti, .J. II binomio di Newton. The Author.
Catalogue of the Queensland Court, International Exhibition. Sydney, 1879.
J. Brazier.
Catalogue of the officers and students of Yale College, 1879-80. The College.
Catalogus der ethnologische Afdeeling van het Museum van bet Bataviaasch
Genootschap van Kunsten en Wetenschappen. 2e D,ruck. The Society.
Central Park Menagerie, report of Directors, 1879. The Directors.
Chambers, V. T. President of Cincinnati Society of Natural History, annual
address, 1879. The Author.
Chapman, H. C. Memoir of James Aitken Meigs, M. D.
Lecture introductory to the course of Jefferson Medical- College for the
session of 1880-81. The Author.
Chatin, J. Les organes des sens dans la serie animale. I. V. Williamson
Fund.
Chief of Engineers, report, 1878. Parts 1, 2 and 3. Engineers' Department,
U. S. A.
Chief Gold Commissioner for the Province of Nova Scotia, report, 1863. De-
partment of Mines, Nova Scotia.
Chief of Oi'dnance, annual report, 1879. War Department.
Chili. Mcmoria de Guerra i Marina, 1878.
Memoria de Relaciones Esteriores, 1879.
Memoria de Ministro de Justicia, etc. 1879,
Memoria de Ministro del Interior, 1879.
Projecto de Codigo rural, 1879.
Anuaria hidrografico de la Marina de Chili, Ano V.
Anuaria de la Oiicina central meteorologica, 1874.
Anuaria estadistico, 1876-77.
ADDITIONS TO LIBRARY. 431
Sessioaes estraordinarias de la Camara de Seiiadores, 1878, Nos. 1 & 2.
Sessiones estraordinarias de la Camara de Disputados, 1878, Nos. 1 , 2 & 4.
Estadistica Agricola 1877-78.
Estadistica bibliografica de la Literatura Chilena, T. 2. University of
Chili.
riiurch, .1. A. New methods of ore concentration and gold amalgamation.
The heat of the Comstock lode. The Author.
Clement, Ch. Constitution Geologique de Luxembourg. 1864. Dr. F. V.
Hayden.
Cobbold. T. S. Parasites. 1879. I. V. Williamson Fund.
Cohn, Ferd. Biologie der Pflanzen. Ill, 1. I. V. Williamson Fund.
Colbeau, .J. A. .1. Materiaux pour la Faune malacologique deBelique. I. The
Author.
Colonial Museum and Geological Survey of New Zealand, 14th Annual re-
port. 1879.
Kaport of Geological Explorations, 1878-79. The Author.
Commission de la Carte gx^ologique de Belgique, Hoboken et Contich, Anvers,
Lennick-St-Quentin, Malines, Lierre, Heyst-op-den-Berg, Putte and
Boom, with maps. The Commission.
Commiss' oner of agriculture, report, 1878. The Author.
Commissioner of Fisheries of the State of California Report 1878 and 1879.
The Commissioners.
Commissioners of Public Charities of the State of Pennsylvania. Tenth an-
nual report, .Jan. 1880. The Board.
(,'omptroller of the Currency, annual report, 1879. The Author.
(.'ope, E. D. On the foramina perforating the posterior part of the squamosal
bone of the mammalia.
On tlie genera of the Creodonta. The Author.
(Wesson, E. T. and Edw. Norton. Tenthredinidne and Uroceridie of North
America. The Author.
Cuvier, G. Revolutions of the surface of the globe. 1831. Dr. I. M. Hays.
Dana, .J. D. Manual of geology, 3d Ed., 1880. I. V. Williamson Fund.
Dawkins, W. B. Early Man in Britain and his place in the Tertiary period,
1880. I. V. Williamson Fund.
The classification of the tertiary period by means of the mammalia.
Further discoveries in the Cresswell Caves' 1879. The Author.
Dawson, Geo. M. Geological map of a portion of the southern interior of
British Columbia. Geological Survey of Canada.
DeCandolle, A. and C. Monographiae phanerogamarum prodromi. Vol. 2,
1870. Wilson Fund.
Decken, C. van der. Reisen in Ost-Afrika. 3er Bd.^ III. Abth. I. V. Wil-
liamson Fund.
De Clercq, F. S. A. Het Maleisch der Molukken, 1876. Batavian Academy
of Sciences.
Department of Agricultux-e. Special Report, Nos. 20-27.
Circular regarding needs of. The Department.
Department of Mines, Nova Scotia, reports, 1862, 1864, 1865, 1867-1879. De-
partment of Mines, Nova Scotia.
Same, 1864. Dr. Jos. Leidy.
Department of Statistics and Geology of the State of Indiana, 1st annual report,
1879. The Department.
Deshayes, G. P. Anlmaux sans vertcbres decouvertes dans le bassin de Paris.
T. 2me, texte, pp. 641, et seq. Atlas, planches 40-107. I. V. Williamson
Fund.
Dohrn, A. Catalogus Hemipterorum, 1869. Dr. Jos. Leidy.
Doremus, C. A. and R. A. Witthaus. Chemistry of the Cobb-Bishop poisoning.
The Authors.
Drapei', J. W. Experiments on solar light. Dr. I. Minis Hays.
432 ADDITIONS TO LIBRARY.
Dumeril, A. and Bocourt. Mission scientifique au Mexique et dans TAmerique
Centrale. Recherches Zoologiques. 3me Partie. Etudes sur les reptiles
et les batraciens. I. Y. Williamson Fund,
Eaton, D. C. Ferns of North America. Pts. 22-27 and Title and Index to
Vol. 2. J. H. Redfield.
Ellet, C, Jr. The mountain top track, 1856. Dr. I. M. Hays.
Elliott, D. G. Monograph of the Bucerotidae. Pts. 7 and 8.
Monograph of the Felidse. Pt. 5.
Encyclopedia Britaunica. 9th Ed. Vols. 10 and 11. I. A'^. Williamson Fund.
Encyklopfedia der Naturwissenschaften. 1 Abth., 6-13 Lief. I. V. William-
son Fund.
Engelman, Geo. Revision of the genus Pinus and description of Pinus El-
liottii, 1880.
Acorns and their germination. The Author.
Engler, A. Entwicklungsgeschichte der Pflanzenwelt. I. Th. I. V. William-
son Fund.
Evarts, H. C, M. D. A new species of Ophrydium. The Author.
Exposition Universelle de 1878. Catalogue du Ministere de 1' instruction pub-
lique des cultes et des beaux-arts. Tomes 1, 2, 3. Section Beige. Catal.
officiel. Minister of Public Works.
Falsan, A., and E. Chantre. Monographic geologique des anciens glaciers et
du terrain erratique de la partie moyenne du Bassin du Rhone. Atlas,
1875. Society of Agriculture of Lyons.
Farlow, W. G. Impurities of drinking-water. The Author.
Financial reform almanac, 1880. Cobden Club.
Fisher, P. Subdivisions des ammonites The Author.
Fitzgerald, R. D. Australian orchids. Pts. 1-5. Colonial Secretary, New
South Wales.
Fol, H. Etudes sur les Appendiculaires du Detroit de Messine, 1872. The
Author.
Foote, A. E. Catalogue of minerals. 5tli Ed. The Author.
Friele, H. Tungebevtebningen hos de Norske Rhipidoglossa. The Author.
Fritsch, A. Fauna der Gaskohle und der Kalksteine der Permformation Boh-
mens. Bd. 1, H. 1 und 2. I. V. Williamson Fund.
Frommann, C. Structur und Bewegungserscheinungen des Protoplasma der
Pilanzenzellen. I. V. Williamson Fund.
Galvani, Luigi, portrait of. Academy of Sciences of Bologna.
Geological and Natural History Survey of Minnesota, reports, 1874, 1875, 1876,
1877-78. N. H. Winchell.
Geological explorations and surveys west of the 100th Meridian. Topographi-
cal Atlas. Wheeler, 1875. Engineer Department, U. S. A.
Geological Survey of Canada, reports of progress, 1844, 1848-49, 1850-51,
1851-52, 1852-53, 1853-56, 1857, 1858, 1875-76. The Survey.
Geological Survey of India. Records, Vol. XII, Parts 2 and 3. Memoirs, 8vo,
Vol. 16, Pt. 1. Memoirs, 4to. Palseontologia Indica, Ser. II., Vol. I,
4; Ser. XIII, I, 1 ; Ser. XIV. I, 1. The Survey.
Geological Survey of Kentucky. A general account of the Commonwealth of
Kentucky. Report on the timbers of Boyce and Mercer Counties. By
W. M. Linney. Chemical report of the sojls, coal, ore, etc. By Robert
Peter. The Survey.
Geological Survey of New Jersey. Annual report, 1879. The Author.
Geological Survey of Sweden. Eleven geological maps. The survey.
Geology of Wisconsin. Survey of 1873-79, Vol. Ill, with folio Atlas. Annual
report, 1879. The Survey.
Geyler, H. T. Ueber fossile Pflanzen aus der Juraformation Japans. The
Author.
Gibson, G. A. Sequence and duration of the cardiac movements. The Author.
Gilbert, G. K. Geology of the Henry Mts., 1877. Department of the Interior.
ADDITIONS TO LIBRARY. 433
Gilpin, E. The mines and mineral lands of Nova Scotia, 1880. Department
of Mines, Nova Scotia.
Gould, J. The birds of Asia. Pts. 28, 29, 30 and 31.
The birds of New Guinea. Pts. 3-11 inc. Wilson Fund.
Graf, L. Anleitung zur Kenntniss des Pferdes nach seiner iluszeren Korper-
form, 1846. Dr. Jos. Leidy.
Grateloup, Dr. Discours sur les sciences et les arts, 1837. Dr. I. Minis
Hays.
Gray, Asa. Botanical contributions, V. The Author.
Gray, A. F. Littorina litorea, Linn, on the American coast. The Author.
Gruber, W. Beobachtungen aus der menschlichen und vergleichenden
Anatomic. 1 and 2 H. I. V. Williamson Fund
Ueber den anomalen Canalis basilaris medianus des Os occipitale beim
Mecsehen. The Author.
Haast. J. von. Geology of the Provinces of Canterbury and Westland, New
Zealand. The Author.
Haeckel, E. Das System der Medusen, ler Th., le Halfte. Text and Atlas.
I. V. Williamson Fund.
Haldeman, S. S. Unsymmetric arrow-heads and allied forms. The Author.
Hamilton, G. Biographical sketch of James Aitken Meigs, M.D. The Author.
Hampe, E. Enumeratio muscorum, 1879. I. V. Williamson Fund.
Hanstein, J. v. Das Protoplasma, 1880. I. V. Williamson Fund.
Hayden, F. V. Five maps of portions of Wyoming, Idaho and Utah. Depart-
ment of Interior.
The Great West, 1880. The Author.
Hays, Dr. Isaac, framed portrait of. Dr. I. Minis Hays.
Hector, James. Handbook of New Zealand, 1879. J. Brazier.
Heger, A. Useful tables for finding specific gravity. The Author.
Heinemann, F. C. Catalogues, 188u. The Author.
Heller, A. A Kir. M. Termeszettudomanyi Tarsulat Konyveinek Czimjegy-
zeke, 1877. Royal Hungarian Society of Sciences.
Helmholtz. Sensations of Tone, 1875. I. V. Williamson Fund.
Heule, J. Handbuch der Eingeweidelehre des Menschen.
Henry, Jos. A summary of researches in sound, 1879. Smithsonian Inst.
Herman, 0. Ungarns Spinnen-Fauna. 3 Bd. Royal Hungarian Society of
Sciences.
Hermann, L. Handbuch der Physiologic. 2er Bd. 2er Th., 5 Bd. 1 Th. I.
V. Williamson Fund.
Hertwig. 0. & R. Die Actinien, 1879. Dr. F. V. Hayden.
Hesse-Martegg, E. von. Nord-Amerika. 1-4 Bd. Dr. F. V, Hayden.
Heude, R. P. Conchyliologie fluviatile de la Province de Nanking et de la
Chine Centrale. 5me and (5me Fasc. I. V. Williamson Fund.
Hewitson, Wm. C. Illustrations of Diurnal Lepidoptera. Pt. 1. I. V. Wil-
liamson Fund.
Hewitson, W. C. and F. Moore, Description of New Indian Lepidopterous
Insects. Rhopalocera, Heterocera. Asiatic Society of Bengal.
Hicks, H. On the Pre-Cambrian rocks of west and central Ross-shire. The
Author.
Hidalgo, J. G. Moluscos marines de Espana, Portugal y las Baleares,
Ent. 15, 16,
MdIuscos del Viaje al Pacitico, 1862-65. Pt. 3a, Ent. 1. The Author.
Hidegh. K. Chemische analyse ungarischer Fahlerze. Royal Hungarian
Society of Sciences.
Hind, H. Y. Report of the Waverly Gold District, 1869.
Report on the Sherbrooke Gold District, 1870.
Report on the Mt. Uuiacke, Oldham and Renfrew Gold Mining Districts
1872.
Report on a topographical survey of part of the Cumberland Coal Field
1873, Department of Mines, Nova Scotia.
39
434 ADDITIONS TO LIBRARY.
His, W. Anatomie menschlicher embryonen. I. Text and Atlas. I. V. Wil-
liamson Fund.
Hooker, J. D. The flora of British India. Pt. VII. The E. Indian Govern-
ment.
Hull, Edw. Geological age of the rocks forming the Southern Highlands of
Ireland.
On the upper limit of the essentially marine beds of the Carboniferous
Group of the British Isles adjoining continental districts.
On the geological relations of the rocks of the South of Ireland to those
of North Devon and other British and continental districts.
On the origin of the " Scalp,"
On a deep boring for coal at Scarle, Lincolnshire.
On the i-elations of the Carboniferous, Devonian and Upper Silurian Rocks
of the South of Ireland to those of North Devon. The Author.
Humphreys, J. T. Discoveries of minerals in Western North Carolina. The
Author.
Hunfalvy, P. Literarische Berichte aus Ungai-n. 8 & 4 Bd. The Hungarian
Academy of Sciences.
Hutton, F. W. Manual of the New Zealand Mollusca. Geological Survev, N. Z,
Huxley, T. H. The Crayfish. 1880. I. V. Williamson Fund.
Index Catalogue of the Library of the Surgeon-General's Office. Vol. I.
War Department.
InostranzefF, A. Metamorphosirte Gesteine im Qouvernement Olonez.' 1879.
I. V. Williamson Fund.
Ein neues, ausserstes Glied in der Reihe der amorphen Kohlenstofi'e. 1880.
The Author.
Inspector of Mines of the anthracite coal regions of Pennsylvania, Reports,
1878. .loseph M. Gazzam.
International Exhibition, Sydney, 1879. New Zealand Court. Appendix to
official Catalogue, 1880. The Commissioners.
Jeffreys, .1. G. The deep-sea mollusca of the Bay of Biscay. The Author.
Joly, N. E. Une lacune dans la serie teratologique remplie par la decouverte
du genre Ileadelphie.
Sur le placenta de I'Ai.
Nouvelles recherches tendant a etablir que le pretendu Crustace decrit
par Latreille, sous de nom de Prosopistoma, est un veritable insecte de
la tribu des Ephemerines. ,
Contribution a I'histoire naturelle et I'anatomie des Ephemerines.
Etudes sur I'embryogcnie des Ephumeres. The Author.
Julien, A. A. Spodumene and its alterations. The Author.
Just, L. Botanis'cher Jahrsbericht. 5er Jahrg., 1877, 3e Abth., 6er Jang. 1 H.
I. V. Williamson Fund.
Kansas State Board of Agriculture, quarterly report, Sept. 30, 1879. March
31, 1880. The Author.
Keller, Ferd. Lake dwellings of Switzerland, 2 vols., 1878. I. V. Williamson
Fund.
Keyserling, E.. Die Spinnen Amerikas. Laterigradae, 1880. I. V. William-
son Fund.
Kiener, L. C. Species general et iconographique des Coquilles vivantes.
Livr. 150-165. I. V. Williamson Fund.
Kingsley, J. S. Synopsis of the N. A. species of the genus Alpheus.
Notes on the N. A. Caridea in the Museum of the Peabody Academy of
Sciences. 1878.
Decapod Crustacea of the Atlantic coast, whose range embraces Fort
Macon.
North American Crustacea belonging to the sub-order Caridea.
Development of Moina.
Geographical distribution of Crustacea. The Author.
ADDITIONS TO LIBRARY. 435
Kingston, G. T. Report of the Meteorological Office of the Dominion of
Canada, 1879. Thp Superintendent.
Kjerulf, Th. Die Geologie des sudlicheu and mittleren Norwegen. 1880.
The Author.
Klaproth, M. II. Analytical Essays. 2 vols , 8vo. 1801. Dr. I. M. Hays.
Klein, E. andE. Noble Stnith. Atlas of histology, Pts. 0-12. I. V. Williamson
Fund.
Klunzinger, C. B. Die Korallthiere des Rothen Meeres. 3er Th. 1. V. Wil-
liamson Fund.
Kneass, S. H. Coal mines of the Lykens Valley Coal Company. 1844. Dr. I.
M. Hays.
Kobelt, W. lUustrirtes Conchylienbuch. 8e & 9e Lief. I. V. Williamson Fund.
Koch, A. Description of the Missourium, 1841. Dr. I. Minis Hays.
Kcilliker, Alb. Grundriss der Entwicklungs-Geschicbte des Menschen und
der hiiheren Thiere. 1880. I. V. Williamson Fund.
Kokscharow's Materialen zur Mineralogie Russland. Bd. VIII, sigs. 5-9. I.
V. Williamson Fund.
Kossmann, R. Zoologische Ergebniss einer im Auft.rage der K. Acad, der
Wissen. zu Berlin ausgefiihrten Reise in die Kiistengebiet^ des rothen
Meeres. 2e Hiilfte, le Lief. I. V. Williamson Fund.
Kiichenmeister, H. Die Parasiten des Menschen. 2e Aufl. 1 & 2 Lief. I.
V. Williamson Fund.
Kuntz, 0. Speciesbeschreibung und Rubus, 1879. I. V. Williamson Fund.
Liedy, Jos. Fresh-water Rhizopods of North America, 1819. Department of
the Interior.
Lente, F. D. Higher education of medical men. The Author.
Leuckart, R. Die Parasiten des Menschen und die von ihnen herriihrenden
Krankheiten. ler Bd., 1 Lief. 2e Auflage, 1879. I. V. Williamson
Fund.
Lewis, H. C. The Trenton gravel and its relation to the antiquity of man.
The optical characters of some Micas.
On Siderof.hyllite.
On Philadelphite.
On a fucoidal plant from the Trias.
The surface geology of Philadelphia and vicinity.
The iron ores and lignite of the Montgomery County Valley. The Author.
Librarian of Congress, annual report, 1879. The Author.
Library Co. of Phila., Bulletin, n. s. No. 5. The Library Co.
Lieberkiihn, N. KeimbUltter der Saugethiere, 1879. I. V. Williamson Fund.
Light-House Board, annual report, 1879. Treasury Department.
Lindsay, W. L. Mind in the lower animals. 2 vols. I-.V. Williamson Fund.
Linnarsson, G. Om Faunan i Lagr^n med Paradoxides Olandicus. Geological
Survey of Sweden.
Lippincott, J. S. The critics of evolution. The Author.
List of vertebrated animals now or lately living in the Gardens of the Zoologi-
cal Society of London. 1st Supplement. The Society.
Lockington, W. N. Notes on Pacific coast Crustacea, 1878. The Author.
Loewe, L. Nervensystem der Siiugethiere und des Menschen, 1880. I. V.
Williamson Fund.
Lyman, Benj. Smith. Geological Survey of Japan. Report of progress for
1878 and 1879. Tookei, 1879. The Survey.
McLachlan, R. Monographic revision and synopsis of the Trichoptera of the
European fauna. Pt. 9. I. V. Williamson Fund.
MacLean, J. P. Mound-builders, 1879. I. V. Williamson Fund.
Malaise, C. Description des gites fossiliferes Devoniens et d'affleurement du
terrain Cretace. Belgian Geological Survey.
Mallery, Garrick. Sign language among the North American Indians, 1880.
Introduction to the study of sign language among the North American
Indians. Smithsonian Institution.
436 ADDITIONS TO LIBRARY.
Mallet, J. W. La Barcenite, 1878. Dr. Jos. Leidy.
Marten, E. vod. Conchologische Mittheilungen. 1 Bd., 1-4 H. I. V. Wil-
liamson Fund.
Martin, E. Histoire des monstres. I. V. Williamson Fund.
Martin, E. Die Tertiiirtchichten auf Java. Palteontol. Th. 2-4 Lief. I. V.
Williamson Fund.
Martindale, I. C. Notes on the Bartram Oak. The Author.
Martini & Chemnitz. Systematisches Conchy lien-Cabinet. Lief. 284-296.
Wilson Fund.
Marsh, 0. C. New Jurassic reptiles. The Author.
Maryland, report on new map of, 1836. Dr. 1. M. Hays.
Mason. John J. Microscopic studies on the central nervous system of reptiles
and batrachians. 1 and 2. The Author.
Matteucci, C. Le90ns sur les pbenomenes physiques des corps vivants. Ed.
Francaise. Dr. Jos. Leidy.
Meehan, Tho3. Native flowers and ferns of the United States. 2d Ser., Pts.
1-24. 1879. The Publishers.
Meigs, J. A., death-mask of. C. W. De Lannoy.
Mencke, M, Beitrllge zur Biologic der Spaltpilze. I. V. Williamson Fund.
Mendell, G. H. Blasting operations at Lime Point, Cal., in 1868 and 1869.
Engineer Department, U. S. A.
Mercantile Library Association of the City of New York, .59th annual report.
The Directors.
Mercantile Library Association of San Francisco, 27th annual report. The
Trustees.
Mexican antiquities, 61 photographs of. (Poinsett Collection.) Wm. S. Beebe.
Milne-Edwards, M. Melanges Carcinologiques. Text and plates. Dr. Jos.
Leidy.
Leyons sur la physiologic et I'anatomie comparee de Uhomme et des ani-
maux. T. 14me, Ire Pt. Wilson Fund.
Mines Commission Report, 1880, Annapolis. Tha Commissioner.
Minks, A. Das Microgonidium, 1879. I. V. Williamson Fund.
Mission Scientifique au Mexique. Recherches Zool. 7me, Partie T. 2. 1880.
I. V. Williamson Fund.
Moleschott, Jac. Untersuchungen zur Naturlehre des Menschen und der
Thiere. XIII Band, ;! & 4 H. I. V. Williamson Fund.
MoUer, V. v. Die Foraminiferen des russischen Kohlenkalks. The Author.
Mongredien, Aug. Free trade and English commerce. 4th ed.
The western farmer of America, The Cobden Club.
Morse, Edw. S, Ancient and modern moUuscan fauna of Omori, Japan. The
Author.
Mueller, F. de. Index perfectus ad Caroli Linntei Species Plantarum. 1880.
The Author.
Miiller, N. J. C. Handbuch der Botanik. ler Bd., lerTh. I. V. Williamson
Fund.
Museum Ludwig Salvator in Ober-Blasewitz bei Dresden. The Author.
Nagle, J. T. Summary of births, mai-riages, still-births, deaths, etc., in New
York City. 1878. The Author.
Native Tribes of South Australia. 1879. R. Schomburgh.
Netoliczka, E Untersuchungen iiber Farbenblindheii und Kurzsichtigkeit.
2 Nos. 1879. The Author.
New York, geological reports, 18^7, 2d Ed. 1840, 1841 & 1851. Dr. I. M.
Hays.
Oiseaux dans la nature. Livr. 1. I. V. Williamson Fund.
O'Neill, T. Warren. The refutation of Darwinism, 1880. The Author.
Orr's Circle ot the sciences, Nos. 1 & 2. 1854. Dr. I. M. Hays.
Packard, A. S., Jr, Zoology. 1879. I. V. Williamson Fund.
Palasontographica. 26er Bd., 3. Lief., 27er Bd., 1. Lief. & Supplement III,
3., 8. & 9. Lief., H. 4. Wilson Fund.
ADDITIONS TO LIBRARY. 437
Palsontologie Fran^aise, Ire Ser. An. Invert. Ter. Jurassique Livr. 42 and 45,
2e Ser. Vegetaux. Ter. Jur. Livr. 29. Wilson Fund.
Palasontological Society's Publications, Vol. 34. I. V. Williamson Fund.
Pasteur, L. Studies on fermeatation, 1879. I. V. Williamson Fund.
Paulucci, M. Excursione scientifica nella Calabria, 1877-78. Fauna Mala-
cologica. L V. Williamson Fund.
Peabody Academy of Science, Salem, Mass. Visitors' Catalogue of the Mus-
eum. 1879. The Society.
Pennsylvania, Geological reports, 1833 and 1836. Dr. I. M. Hays.
Pfeiffer, L. Nomenclator Heliceorum viventium. 5. & 6. Lief. I. V. Wil-
liamson Fund.
Physical Science, Agriculture, etc. 81 pamphlets on. Dr. I. M. Hays.
Pickering, Cbas. Vocabulary of the Soahili language. Mrs. Chas. Pickering.
Porter, J. Topographical description of Plainfield, 1834. Dr. L M. Hays.
Prazmowski, A. Entwickelungsgeschichte und Fermentwirkung einiger Bac-
terien-.Arten. I. V. Williamson Fund.
Public Ledger Almanac. 1880. The Publisher.
Public Library, Milwaukee, 2d annual report. The Trustees.
Putnam, J. D. Biological and other notes on the Coccidaj, 1880. Jhe Author.
Quaritch, B. Letter to General Starring, Jan. 14, 1880. The Author.
Rand, B. H. Two lectures on impure air and ventilation. The Author.
Ratzel, Fr. Die Vereinigten Staaten von Nord-Amerika. 2er Band. I. V.
Williamson Fund.
Report of Commissioners appointed under resolve of 1856, Chap. 58, concern-
ing the artificial propagation of fish. Boston, 1857. Dr. I. M. Hays.
Reyer, E. Vier Ausfluge in die Eruptiomassen bei Christiania.
Zinn in Birma, Siam & Malakka.
Zinn in Australien und Tasmanien. The Author.
Riviere, E. Grotte de Saint-Benoit. 1878.
Le pliocene de Castel d' Appio en Italic, 1879.
De quelques hyperostoses de poisons trouvees dans les grottes ijuarter-
naires de Menton en Italie.
Note sur des instruments en obsidieune trouves en Grece, 1879. The
Author.
Robert, P. Les oiseaux dans la nature. Livr. 2-10. I. V. Williamson Fund.
Roberts, E. P. Directions for sowing, transplmting and raising the Mulberry
Tree, 1839. Dr. I. M. Hays
Roemer, F. Lethnea geognostica. I. Th. I. V. Williamson Fund.
Rossmiissler's Icjnographie der EuropUischen Land- und Siisswasser-Mollus-
ken. VII, 1-3. Wilson Fund.
Rothi'ock, J. T. Catalogue of trees and shrubs in the horticultural gardens
adjacent to Horticultural Hall, Fairmount. 1880. The Author.
Roucher-Deratte, C. Le9ons physiologic© meteorologiques sur les constitutions
des saisons. 1804 Dr. I. M. Hays.
Russ, Dr. K. Die fremdlilndischen Stubenvogel. Ill, 7-9. I. V. Williamson
Fund.
Rutherford, J. Coal-fields of Nova Scotia. Department of Mines, Nova Scotia.
Ryder, J. A. Ichthydium ocellatum, 1880.
On the occurrence of Freia producta, Wi-ight, in the Chesapeake Bay, etc.
The Author.
Sadler, John. Report on temperature during the winter of 1878-79 at the
Royal Botanic Garden, Edinburgh. The Author.
Saint-Lager, Dr. Reforme de la nomenclature botanique. The Author.
St. Louis Mercantile Library Association, annual report, 1879. The Directors.
Saunder, W. Tea-culture as a probable American industry. 1879. Depart-
ment of Agriculture.
Saussure, H. de. La Grotte du See. The Author.
Scheftler, H. Die Naturgesetze. 3ter Th, 6te, 7te und 8te Lief. The Author.
438 ADDITIONS TO LIBRARY.
Schimper, W. P., and K. A. Zittel. Handbuch der Palseontologie. 1. Bd., 3te
und 4te Lief. I. V. Williamson Fund.
Schlegel, H. Museum d'histoire naturelle des Pftys-Bas. T. 8, Monog. 41.
Wilson Fund.
Schmidt, Emil. Kraniologische Untersuchungen.
Mittheilungeu aus dei'anthropologischen Literatur Amerikas. The Author.
Schmidt, 0. Die Spongien des Meerbusen von Mexico. 2tes (Schluss-) Heft.
1880. Museum of Comparative Anatomy.
Schomburgh, R. On the naturalized weeds and other plants of South Australia.
On the Urari.
Catalogue of the plants under cultivation in the Government Botanic Gar-
den, Adelaide, South Australia.
Report on the progress and condition of the Botanic Garden and govern-
ment plantations during the year 1879. The Author.
Scott, W. B., and H. F. Osborn. Early development of the common newt.
The Authors.
Scudder, S. H., Writings of, compiled by George Dimmock. 1879. The Editor.
The Devonian Insects of New Brunswick. The Author.
Second Geological Survey of Pennsylvania. C-', 2 vols., G-, G^ H*, P, 0^, P^,
Q2, Q', R, T, "V2. The Commission.
Same. Report P. .Jos. M. Gazzam.
Seemann, B. Flora Vitiensis. 4to. 1865-73. L V. Williamson Fund.
Semper, C. Reisen im Archipel der Philippinen. 3ter Th., eter Bd., 5 H.
Wilson Fund.
Sheafer, P. W. Anthracite coal-fields of Pennsylvania and their exhaustion.
Diagram of the progress of the anthracite coal trade of Pennsylvania.
The Author.
Short, J. T. North Americans of antiquity. 1880. L V. Williamson Fund.
Sigsbee, C. D. U. S. Coast Survey. Deep-sea sounding and dredging. 1880.
C. P. Patterson.
Smith, F. Description of new species of Hymenoptera in the British Museum.
1879. Trustees of the Museum.
Smith, J. A Mutations of the earth, 1846. Dr. I. M. Hays.
Smith, .J. L. Minerals and mineral waters of Chili. Dr. I. M. Hays.
Progress and condition of several departments of industrial chemistry.
Paris Univ. Exp., 1867. Dr. Jos. Leidy.
Smithson, .Tames, Scientific writings of. 1879.
Smithsonian Institution and National Museum, visitor's guide.
Smithsonian Institution Miscellaneous Collections, Vol. 16 and 17. Contribu-
tions to Knowledge, Vol. 22.
Report, 1878.
Journal of Board of Regents, etc. The Institution
Smyth, R. B. The aborigines ot Victoria. The Government of Victoria.
Snellen van Vollenhoven, S. C. Pinacographia. Pt. 9. I. V. Williamson Fund.
Soret, L. Rapport du President de la Societe de physique et d'histoire
naturelle de Geneve, 1879. The Author.
South African Museum, report, 1879. The Trustees.
Spencer. H. Ceremonial institutions. 1880. I. V. Williamson Fund.
Stearns, R. E. C. New species or variety of land snail from California. The
Author.
Steenstrup, J. Sepiella, Gray.
De Ommatostrephagtige Blieksprutters indbyrdes Forhold. The Author.
Steiner, L. H. Annual address before the American Academy of Medicine at
New York, Sept 16, 1879. The Author.
Stille, A., M. I). Humboldt's life and character, 1859. Dr. I. M. Hays.
Memoir of Isaac Hays. The Author.
Stone, 0. Extra-meridian determination of time. The Author.
Strebel, H. Fauna mexikanischer Land- und Susswasser-Conchylien. Th. 4.
I. V. Williamson Fund.
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Strecker, H. Butterflies and moths, 1879. The Author.
Strobel, P. Material! per una malacostatica di terra e d'acqua dolci.
Disp. 1-4. •
Struckmann, C. Die Wealden-Bildungen der Umge^end von Hannover, 1880.
The Author.
Sveriges Geologiska Undersiikning. Ser. A A, Nos. 68, 69, 71 and 72 ; A B 4
and 5 : € 8vo, 31, 32, 34 and 35 ; C 4to, 29 and 33. The Survey.
Taplin. Rev. G. Folklore, manners, customs and languages of the south
Australian aborigine-e, 1879. R. Sehomburgh. ,
Tapparone-Canefri, C. Museum rauluccianum. Etudes malacologiques.
The Author.
Taramelli, T. Catalogo ragionato delle Rocce del Fruili.
Sulla formazione serpentinosa dell' Apennino Pavese.
Monograiica stratigrafia e paleontologica del Lias nelle Provincie Venete.
The Author.
Tate, R. Zoologica et , Plaeontologica Miscellanea, chiefly relating to South
Australia.
The natural history of the country around the head of the great Austra-
lian Bight.
The Adelaide Philosophical Society. Anniversary address of the Presi-
dent. The Author.
Taylor, R. C. Coal regions in the environs of Blossburg, 1833. Dr. I. M.
Hays.
Taylor, W. B. A memoir of .Joseph Henry, 2d Ed. Smithsonian Inst.
Thomas, C. H., M. D. Researches on hearing through the medium of the
teeth and cranial bones. The Author.
Traill, G. W. The Alga> of the Firth of Forth. The Author.
Trautwine, .J. C. Internal improvement system of the South, 1889. Dr. I.
M. Hays.
Tryon, G. W. Jr. Manual of Conchology, Pts. 5-8. The Author.
TurnbuU, C. S. Audiphone and Dentaphone. The Author.
United States Coast and Geodetric Survey. Pacific Coast Pilot. Coasts and
islands of Alaska. 2d Series, 1879. C. P. Patterson.
United States Commission of Fish and Fisheries. Part V, Report of the Com-
mission for 1877. The Commission.
United States Entomological Commission. Bulletin Nos. 3, 4 and 5. Depart-
ment of the Interior.
United States Geographical Survey west of the 100th meridian. Reports,
Vols. 2, 3, 4 and 5. Engineer Department, U. S. A
United States Geological and Geographical Survey of the Territories. 11th
annual report. Bulletin, Vol. V, No. 3. Department of the Interior.
United States National Museum. Bulletin Nos. 13 and 17. Department of
the Interior.
University of Minnesota, report of the Regents of, 1872. The Regents.
Vacek, Mich. Vorarlberger Kreide, 1879. The Author.
Van Beneden and Gervais, MM. Osteographie des Cetaces vivants et fossiles.
Text and atlas. Lief. 17 and 18. I. V. Williamson Fund.
Van der Berg, L. W. C. Verslag van eene Verzamelung Maleische, Arabische,
.Javaansche en andere Handschriftendoorde Regeering van Nederlandsch
Indie. Batavian Academy of Sciences,
Vanuxem, L. Experiments on anthracite, plumbago, etc., 1825. Dr. I. M.
Hays. ,
V^lain, Ch. Etude microscopique des verres resultant de la fusion des
cendres de graminees.
Mission de 1' He de Saint Paul. Recherches geologiques. 4to Paris,
1879. The Author.
Victoria. Reports of the Mining Surveyors, 31st December, 1878, 31st March,
1879.
Mineral Statistics of, 1878.
440 ADDITIONS TO LIBRARY.
Report of Inspector of Vines, 1878. Dr. F. V. Hayden.
Vogdes, A. W. Monograph of the genera Zethus, Cybele, Encrinurus and
Cryptonymus. The Author. t
Wagner, Wm. Five new fossils of the older pliocene formation of Maryland
and North Carolina'. The Author.
Waterhouse, Owen. Illustrations of the typical specimens of Coleoptera in
the collection of the British Museum. Pt. I, Lycidae. The British
Museum.
Watson, R, Chemical Essays. 7th Ed. 5 vols., 1800. Dr. I. M. Hays.
Watson, R. B. Mollusca of H. M. S. "Challenger" Expedition. Pt. 5. The
Author.
Watson, S. Geological Survey of California. Botany, Vol. 2. I. V. William-
son Fund.
Weismann, Aug. Uber die letzten Ursachen der Transmutationem, 1876.
Dr. Jos. Leidy.
Wex, G. V. Improvement of the Danube at Vienna, 1880.
Second treatise on the decrease of water in springs, creeks and rivers,
1880. Engineer Department, U. S. A.
White, C. A. Paleontological Field work for the session of 1877.
Cretaceous fossils of the Western States and Territories.
Note on the occurrence of Productus giganteus in California.
New invertebrate fossils from the Mesozoic and Cenozoic rocks of Ar-
kansas, Wyoming, Colorado and Utah.
Description of a very large fossil gasteropod from the State of Puebla,
Mexico.
Contributions to paleontology, Nos. 2-3. The Author.
White, F. E. Valedictory Address, Woman's Medical College, 1880-81. The
Author.
Wbiteaves, J. F. On some marine invertebrata of Queen Charlotte Island.
The Author.
Whitfield, R. P. Fossil crustaceans from the Upper Devonian rocks of Ohio.
The Author.
Whitney, J. D. The Geological Survey of California. An address delivered
before the Legislature of California.
Lecture on geology delivered before the Legislature of California, Feb.
28, 1862. Dr. .Jos. Leidy.
Wilhelm, K. Siebriihrenapparates dicotyler Pflanzen. I V. Williamson Fund.
Wilkes, Chas. Western America, 1840. Dr. I. M. Hays.
Wilson, T. B., engraved portrait of. Rathmel Wilson.
Winchell, A. Preadamites, 1880. I. V. Williamson Fund.
Witter, F. M., Catalogue of Iowa mollusca in the collection of. The Author.
Wood, H. C. and H. F. Formad. Research on the effects of inoculating the
lower animals with diphtheritic exudation. Dr. Wood.
Woodward's Gardens, illustrated guide to. 1880. The Author.
WooUs, W. Plants indigenous in the neighborhood of Sydney. The Author.
Yale College, Catalogue, 1880.
Obituary records of graduates, 1880 and supplement.
Yale College in 1880. The Librarian.
Yarrow, H. C. Mortuary Customs among the North American Indians. The
Author.
Young Men's Mercantile Library Association of Cincinnati. 45th annual
report, 1880. The Association.
Zeiler, R. Explication de la Carte geologique de la France. Tome 4me, 2e
Parte. Dr. F. V. Hayden.
Zesch, F. and 0. Reinecke. Coleoptera of Butfalo. The Author.
Zoological record, Vol. 15, 1878. I. V. Williamson Fund.
Zoological Society of London, list of vertebrated animals, 7th Ed., 1879.
Catalogue of Library. The Society.
Zuckerkandl, E. Morphologie des Gesichtsschiidels. I. V. Williamson Fund.
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Adelaide. Philosophical Society. Transactions, 1878-79. The Society.
Amsterdam, K. Akademie van Wettenschappen. Verslagen en Mededeelingen.
Afd. Letterkunde. 2e Reeks, 8 Deel. ; Afd. Natuurknnde, 2e Reeks, 14
Deel. Jaahrboek, 1878. Trocessen-Verbaal, Afd. Nat. Mei, 1878-Apr.
1879. Verhandlingcii, Deel 19. Alf. Nat. Deel 12. The Society.
Angers. Societo Nationale d' Agriculture, Sciences et Arts. Memoires, T. 19
and 20. The Society.
Atco Science Advocate, Vol. I, No. 1. The Editor.
Augsburg. Naturhistorische Verein, 25er Berichte. The Society.
Baltimore. American Chemical .Journal. Vol. 1, No. 1. The Editor.
American Journal of :Mathematics, Vol. 2, Nos. 3 and 4 : Vol. o. No. 1.
The Editor.
.Johus Hopkins University. Studies from the Biological Laboratory, Nos.
1, 2 and 4. Report of the Sd year. The University.
Peabody Institute, 12th and 13th annual reports. The Trustees.
Batavia. Genootschap van Kunsten en Wetenschappen Tijdschrift, Deel 23,
Afl. 5 and 6 ; Deel 24, Afl. 1, 2. 3 and 6 : Deel 25, Afl. 1 and 2 ; Notulen,
Deel 14, Nos. 2, 3 and 4; Deel 15, Nos. 2, 3,4; Deel 16, Nos. 1-4.
Gedenkboek, 1778-1878. The Society.
Natuurkundig Vereen in Nederlandsch Indie. Tijdschrift, Zevende Serie,
Deel 8. The Society.
Belfast. Naturalists' Field Club, annual report, Vol. 1, Pts. "> and 6. The
Society.
Natural History and Philosophical Society. Proceedings, Sessions 1878-
79, 1879-80. The Society.
Berlin. Archiv fiir Naturgeschichte, 44er .Jahrg., 5 H. ; 45er .Jahrg., 4 and 5
H. ; 46er Jahrg., 1, 2 and 3 H. The Editor. ■ ,
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1 H. The Society.
Entomologische Verein. Zeitschrift, 23er Jahrg., 2 H.; 24 Jahrg., 1 H. The
Society.
Gesellschaft Naturforschender Freunde. Sitzungs-Berichte, 1879. The
Society.
K. Preussischen Akademie der Wissenschaften. Monatsbericht, 1879, Juli-
1880, Juli. The Society.
Der N iturforscher, 12 Jahrg., No. 27-13 Jahrg., No. 13. The Editor.
Naturae Novitates, 1879, No. 19-1880, No. 8. The Editor.
Verein zur Beforderung des Gartenbaues. Monatschrift, 22 Jahrg., Jan.-
Dec. The Society.
Zeitschrift fiir die gesammten Naturwissenschaften, 3d Folge, 3 Bd., No.
4. The Editor.
Bern. Naturforscheude Gesellschaft. Mittheilungen, Nos. 937-978. The
Society.
Besan9on. Academie des Sciences, Belles-Lettres et Arts, 187(3-78. The
Society.
Beziers. Societe dEtude des Sciences Na*urelles. Bulletin. Ann(-es 1, 2 and
3, Fasc. 1 and 2. The Society.
Bonn. Archiv fiir Mikroskopische Anatomie, 17er Bd., 3 H-l8er Bd., 4 H.
I. V. Williamson Fund.
Naturhistorische Verein. Verhandlungen, 35er Jahrg., 2e and 36 Jahrg.,
le Halfte. The Society.
Bordeaux. Socic^te Linneenne. Actes, T. 33, Livr. 3, 4 and G. The Society.
Societe des Sciences physiques et naturelles. Momoires, 2e Ser., T. 3, 3e
Cah., and T. 4, le Cah. The Society.
442 ADDITIONS TO LIBRARY.
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1 and 2. The Society.
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Braunschweig. Archiv fiir Anthropologie, 12. Bd., 2-4 Viertelj. I. V. Wil-
liamson Fund.
Verein fiir Naturwissenschaft.. Jahresbericht, 1879-80. The Society.
Bremen. Natnrwissenschaftliche Verein. Abhandhingen, 6. Bd., 2tes und
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Brooklyn. Entomological Society. Bulletin, Vol. 2, Nos. 4-12. The Society.
Brunn. Naturforschende Verein. Verhandlungen, 17. The Society.
Bruxelles. Societi'- Beige de Microscopic. Proces Verbaux, 1879, Nov. 27 — ,
1880, No. 12. Annales T. 4. The Society.
Soci^te Entomologique. Compte Rendu, Ser. 2, No. 60-72. Annales T. 22.
The Society,
Soeiete Malacologique. Annales, T. 9me, 2e Fasc. Proces Verbaux, T.
8, 4 Oct 1879-7 Fev. 1880. The Society.
Buda-Pest. M. Tudom. Akademia. Ertekezesek amath. Tudomanyok Korebol,
7 Kotet, 1 Szam^l Kotet, 19. Math, es Termeszet. Kozlemenyek, 14
and 15 Kotet. Evkonyvei, Tizenhatodik Kiitet, 11 Uai-ab. The Society.
Ungarische National-Museum. Termeszetrajze Fuzetek, 4 Kcitet, 3. The
Museum.
Buffalo. North American Entomologist, Vol. 1, Nos. 6-12. The Editor.
Caen. Academic Nationale des Sciences, Arts et Belles-Letters. Memoires,
1879 The Society.
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Society.
Calcutta. Asiatic Society of Bengal. Journal, Vol. 47, Pt. 1, No. 4; Pt. 2,
No. 4: Vol. 48, Pt. 1, Nos. 1-4; Pt. 2, No. 1; Vol. 49, Pt. 1, No. 3.
Proceedings, 1879. Nos.' 2-10. The Society.
Same. .Journal, Vol. 48, Nos. 128-130. Proceedings, 1879, Nos. 5-10.
Isaac Lea.
Stray Feathers, Vol. 8. Nos. 2-6. I. V. Williamson Fund.
Cambridge. Appalachian Mountain Club. Appalachia, Vol. 1 and Vol. 2, Nos.
1 and 2. The Society.
Bussey Institution. Bulletin, Nos. 2-4, and Vol. 2, Nos. 1, 2 and 3. The
Institution.
Harvard University. Library Bulletin, Nos. 14, 15 and 16. Bibliograph-
ical Contributions, Nos. 1 and 2. The Trustees.
Museum of Comparative Zoology. Memoirs, Vol. 6, No. 1 : Vol. 7, No. 1
and No. 2, Pt. 1. Reports, 1878-79; 1879-80. Bulletin, Vol. 5, No.
16 and title; Vol. 6, Nos. 1-11 ; Vol. 7, No. 1. The Director.
Nuttall Ornithological Club. Bulletin. 1880, .Jan.-Oct. The Society.
Peabody Museum of American Archaeology and Ethnology. 12th and loth
annual reports. 'The Director.
Psyche, Nos. 65-76. The Editor.
Cap Rouge. Le Naturaliste Canadien, Dec. 1879-Oct. 1880. The Editor.
Cassel. Malakozoologische Blatter, Neue Folge, 2. Bd., 1. Bg.-3. Bd., 2. Bg.
I. V. Williamson Fund.
Verein fiir Naturkunde. Bericht 26 und 27. The Society.
Cherbourg. Societe Nationale des Sciences Naturelles. Memoires, T. 21 and
Catalogue of Library. The Society.
Chicago. American Antiquarian, Vol. 2, No. 3-Vol, 3, No. 1. The Editor.
Central Fishcultural Society. Proceedings, 1st An. Meeting. The
Society.
Oriental and Biblical Journal, Vol. 1, No. 2. The Editor.
Christiana. Archiv for Mathematik og Naturvidenskab. 4 Bd., 3 H.-5 Bd., 3
H. The Editor.
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Cincinnati. Society of Natural History. .Journal, Vol. 2, No. 2-Vol. o. No. 3.
The Society.
Copenhagen. Videnskabelige Medclelelser, 1877-78 ; ]87r'-80, 1 & 2.
Naturhistorisk Tiddskiift, Schiodte, 12 Bd., 1 H.. .also 1st Ser.. Vols. 1-4;
2d Ser., Vo'. 1. Nos. 4 and G, and Vol. 2; 3d Ser., Vol. 4-11 ^ I. V.
Williamson Fund.
Societc Royale des Antiquaries du Nord. Memoires, n. s. 1878-79. Til-
laeg, 1877, 1878. I. V. Williamson Fund.
K. D. Videnskabernes Selskab. Oversigt. 1879, No. 3. 1880, No. 1. Skrif-
ter, .5te Raekke, Vol. 9 No. f. Vol. 11 No. C. The Soc'ety.
Cordoba. Academia Nacional de Ciencias Exactas. T. ?>, Ent. 1 & 2. Boletin
T. 2, Ent. 4 : T. 3, Ent. 1. The Society.
Crawfordsville. Botanical Gazette, Vol. 5, No.s. 2-11. The Editor.
Danzig. Naturforschende Gesellschaft. Schriften. Neue Folge, 4. Bd., 4. H.
The Society.
Darmstadt. Verein fiir Erdkunde. Notizblatt. ?>. Folge, 18. H. The Society.
Davenport. Academy of Natural Sciences. Proceedings, Vol. 2, It. 2 : Vol.
3, Pt. 1. The Society.
Dijon. Academie des Sciences, Arts et Belles-Lettrrs. Annee, 1878-79. The
Society.
Dorpat. Naturforscher Gesellschaft. Sitzungsberichte, 4er Bd., 2. H. Ar-
chiv fiir die Naturkunde Liv-Ehst- und Kurbinds, 1. Ser.. Bd. 8, No. 4.
The Society.
Dresden. K. Leop. -Carol. -Deutschen Akademie der Naturforscher. Nova
Acta, Vol's 39 and 40. Leopoldino, H. 12-15. The Society.
Naturwissenschaftliche Gesellschaft Isis. 1879 Jan. — Dec. The Society.
Verein fiir Erdkunde. Jahresbericht 16er. The Society.
Dublin. Royiil Dublin Society. Proceedings, n. s. I, 1-3 ; II. l-(j. Trans-
actions, n. s. Vol. 1, Nos. 1-12; Vol. 2, Nos. 1 and 2. The Society.
Royal Geological Society of Ireland. .Journal, Vol. 15, No. 2. The
Society.
Easton. American Institute of Mining Engineers. Transactions, Vol. 7.
Edinburgh. Botanical Society. Transactions and Proceedings, Vol. 13, Pt. 3.
The Society.
Geological Society. Transactions, Vol. 3, Pt. 2. The Society.
Royal Society. Proceedings, Vol. 10, No. 103. Transactions, Vol. 28, Pt.
2, and Vol. 29. Pt. 1. The Society.
Scottish Naturalist, Nos. 37-40. The Editor.
Emden. Naturforschende Gesellschaft. Jahresbericht 64er. Kleine Schriften,
18. The Society.
Erlangen. Physikaliscb-medizinische Societtit. Sitzungsberichte, 11 H.
The Society.
Frankfurt a. M. Aerztliche Verein. Jahresbericht 22er. The Society.
Deutsche Malakozoologische Gesellschaft. Nachrichtsblatt, 1880, No. 1.
The Society.
Senckenbergische Naturforschende Gesellschaft, Abhandlungen 11. Bd.,
4 H. Bericht, 1878-79. The Society.
Der Zoologische Garten, 20 Jahrg. 7-12. The Zoological Society of Frank-
ford.
Freiburg, i. B. Naturforschende Gesellschaft. Berichte, Bd. 7, H. 4. The
Society.
Gand. Archives de Biologic, Van Beneden and Van Bambeke, T. 1, Fasc.
1-3. I. V. Williamson Fund.
Geneva. Schweizerische pallloctologische Gesellschaft. Abhandlungen Vol.
6. I. V. Williamson Fund.
Societe de Physique et d'Histoire Naturelle. Memoires, Vol. 26, No. 2.
The Society .
Genoa. Societa di Letture e Conversazioni Scientifiche. Giornale, Anno 3, No.
10— Anno 4, No. 4. The Society.
444 ADDITIONS TO LIBRARY.
Giessen. Oberhessische Gesellschaft fiir Natur- und Heilkunde, ISerBericht.
The Society.
Glasgow. Philosophical Society. Proceedings Vol. 11, No. 2. The Society.
Gottingen. K. Gesellschaft der Wissenschaften. Nachrichten, 187U. The
Society.
Guadalajara. Sociedad de Ingenieros de .Jalisco. Boletin T. 1, No. 1. The
Society,
Hamburg. Naturwissenschaftlicher Verein. Abhandlungen 7 Bd. 1 Abth.
The Society.
Harlem. Hollandische Maatischappij der Wetenschappen. Natuurkundige
Verhandelingen, Deel 4, 1 Stuk. The Society.
Musee Teyler. Archives Vol. 5, Pt. 2. The Director.
Socieie Hollandaise des Sciences. Archives, T. 14, 3e Livr. — T. 15, 2e
Livr. The Society.
Helsingfors. Finska Vetenskaps Societet. Ofversigt, 21. Bidrag, 32. Obser-
vationes Meteorologiques, 1877, 1878. Acta, XI. The Society.
SilUskapet pro fauna et flora Feiuiica. Ny serie, 2, 3, 4, 6-11 Haftet.
Acta I. Meddelanden 1—5. The Society.
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30
INDEX TO GENERA.
451
INDEX TO GENERA,
1880.
Abies 333, 341, 350
Acanthocycles o7
Acanthodoris 88
Actteon 366
Acmoecopleura 188, 204
Adalaria 75
Akiodorus 52
Alee m9
Ambrosia 343
Ampelopsis 5
Amphicarpiea o59
Amphidesiiia 21, 20
Anarcbaris 334
Analcite 252
Anciliaria 364, 365
Andromeda 356
Anomia 21, 24
Antrozovxs 227
Aratus 189, 218
Arctomys 348
Area 21, 23. 24, 29
Artemi;^ 21, 23, 24, 25, 29
Artibeus 393
Arvicola 348
Asbolite 243
Assulina 336, 339
Astarte 364
Astrophycus 293
Astarte 21, 23, 24, 29, 32
Atalapha 133, 393
Atta 359
Betula 343
Biotite 246
Bonellia 29
Brachygrapsus 188, 203
Branchipus 156
Buceinum 28
Bunias 132
Buthotrephis 293
Calymene 176
Cancellaria 29, 365, 369
Caneer 179
Canis 348
Cardita... 21, 24, 364, 365 369
Carditamera 32
Cardium 21, 23, 24, 29, 365. 369
Carex 353
Cariacus... 895
Caricella 368, 373
Carya 348
Cassidaria 365
Castanea 35 J
Castor 848
Castoroides 317
Catostomus 237
Caulolatilus 13, 14
Centaurea 132
Centropyxis 338
Cervus 348
Chasmagnathus 189, 221
Chelydra 348
Chirocephalus 156
Chromite 273
Cistudo 348
Clistocoeloma - 189, 219
Coilogenys 391, 396
Tonus 348
Cookeite 249
Corbula..21, 23, 24, 25,27, 29, 32, 364
Cornus 348
Crassatella 21, 23, 24, 29, 32
Crepis 132
Culsageeite 253
Cyelograpsua 189, 220
Cyeloloma 132
Cylicosmilia 864
Oyonyeteris 393
Cyperus 132
Cyrtograpsus 188, 197
Cytberea,.21, 23, 24, 29, 364, 366, 370
452
INDEX TO GENERA.
Calcydoris 89
Damourite 249,
Danthonia
Dasyprocta 386, 392,
Dekaya
Dentalium 28, 29, 365, 367,
Desmodium
Desmodus 133,
Diaulula
Dicotyles
Didelphys
Difflugia 335,
DilocarcinuH
Dolomite
Doridunculus
Doris
256
343
396
13
375
132
393
40
347
389
338
35
243
52
85
Echinospermum 131
Echinus 27
Epigrapsus 188, 192
Epomophorus 393
Epsomite 257
Erethizon 348, 389, 392, 396
Erinaceus 387
Eriocheir 189, 210
Euchirograpsus 188, 202
Euglypha 336, 338
Euphyllite 249
Eustrongy lus 131
Felis 348
Fiber 348
Filaria lO, 130
Fimbristylus 132
Fissurella 29
Flourite 243
Formica 376
Fucoides 293
Fusus 28, 364, 365. 371, 875
Garnet 241
Gelasimus 135
Geograpsus 188, 195
Gly pt ograpsus 189, 206
Goniopsis 188, 189
Gordius 131
Gorilla 128
Qrapsodes 188, 197
Grapsus.. 188, 192
Heleopera 385, 338
Helice 189, 219
Helix 348
Hemifusus 372
Hemitripterus 233
H esperomy s 348
Heterograpsus ..189, 207
Hierochloe 344
H olograpsus 224
Homalonotus 176
Hottonia 157
Houstoiiia 349
Hy alosph enia 335, 340
Hydra 156
Hypnum 333, 343
Hy pochoeris 1 32
Infundibiilum 29, 366
Isocardia 21. 23, 24, 29, 32
.Jarosit e 33"!
JefFerisite 253
.Jorunna 46
Juglans 848
Killingia 132
Lajvibuccinum 364, 371
Lagostomiis 386, 891
Lamellidoris , 58
Lasiony cteris 398
Lasiurus 183
Latilus 13
Leda 21, 23
Leiolophus 189, 224
Leda 365
Lemna 230
Lepidium 132
Lepidolite.. ..:... 248
Lepidomelane 247
Leptograpsiis 188, 196
Lepton 21
Leptonyx 227
Lepus 348
Leiicwa 132
Lignite 281
Lima 21, 23, 29
Linguella 59
Lonchoglossa 393
Lucina. 21, 23, 24, 29
Macacus 887
Macrobiotus 333
Mactra 21, 24, 27
Margarite 251
Marginella 29
Megaderma 393
INDEX T(i GENERA.
453
Melampusr 27
Melampyi'um Stj^
Meleagris o48
Mephitis 348
Mermis 131
Mesopi thecus 175
Meta^esarma 18'.t, 211
Metopo^rapsus 188. 190
Millerite 243
JVrodiola 21, 24. 27
Molo?sus 359, 393
Molybdenite 243
Muscovite 242, 247
Mya 21, 24, 27
Mytiltis 21, 23, 32
Kasua 388
Natica 27. 304,365, 363
Nautilograpsus 187, 188. 201
Nebela 3^3, 33G
Neotoma 347
Noctilio 393. 394
Nucula 28
Nupliar 157
Nycteris 133
Ocypoda 179
Onchidoi-is 59
Orbiiograpsns 188. 194
Ostre4 21, 23, 24, 32, 364-366
Oxalis 350
Pachygrapsus 188. 198
Pectuucnlus 22, 24, 29
Palseophycus 293
Palio 108
Panopu-a 21. 24, 25
Partiila 229
Pecten 22, 24, 25, 30, 866, 369
Perigrapsus •. 189, 210
Periploma 32
Perna 22, 24, 27, 32
Petricola .' 22, 24
Phikdelphite 310, 313
Phlogophite 244
Pholadomya 22, 25
Pholas 22. 24, 25
Phyllorhina 133, 339
Phylostoma 392
Pinus 341
Pithecolobium 355
Placocist-i 339
Plagusia 189, 223
Platygrapsus 189, 210
Pleiirotoma..29, 365, 366. 368, 369, 373
Plicatula 22, 32
Plumatella 157
Polycera 105
Polyergus 376
Populus 353
Prochlorife 251
Procy on 348
Pseudograpsus 188, 204
Pseudoliva 365, 371
Pseudothelphusa 34
Pteropus 393
Ptychognatlius 188, 20'^
Putorius 348
Pyropbyllite 248 ,
Pyriila .- 365, 367, 374
Quercus ; 128, 342
Randite 274
Kangifer 347
Rhaconotus 189, 213
Rliinopoma 359
Rhipidolite 251
Rhododendron 333
Riccia 230
Rochelia 131
Rostellaria 364
Rotifer 333
Sancara
Sarmatium 189,
Saxicava 22, 25,
Scalaria 28,
Scalops
Sciurus 348,
Semnopithecus
Serpentine 241, 248,
Serpula
Sesarma 189,
Siderite
Siderophyllite
Si garet us
Simia
Solarium '29, 364, 365,
Solecurtus
Solen 22, 25,
I Spiroslomum
Spongilla 330,
Steatite
Stent or
Sterlingite
Strepsidura '. ..
Streptocephalus
Talc
Tarn i as.
.248,
212
32
29
348
389
175
273
27
213
242
254
29
160
375
27
27
157
357
273
157
256
372
156
252
348
454
INDEX TO GENERA,
Taxodium 9 Variina 187, 188, 205
Tellina 22, 24 Venericardia 366
Thelphusa 35 Venus .22,24,25, 29, 32
Thuja 9 Vermiculite 250
Tornatella 364, 366 Vespertilio 227, 348
Trinema 336, 339 Vesperugo 393
Triopha 108 '. Vesperus 373
Triticum 132 ': Vitis 10
Tritonium 369 i Voluta 29, 365
Trochus 29 Volvox 157
Turbinella 364, 873 Vorticella 157
Turritella 29, 365 ! Vulpes 348
Unio 348 Wistaria 358
Ursus 349
Utica 189, 206 Yucca 355
INDEX.
455
GENERAL INDEX.
AtUlitions to Library, 429. |
Agnew, Wm. G. E., Announcement of
death of, 226.
Allen, H., Description of a foital
Walrus. 38 ; Mammary Glands of
Bats, 133; On some homologies in
Bunodont Dentition, 226 ; The Pha-
langes of Bats, 359 ; On the Tem-
poral and Masseter Muscles of
Mammals, 385.
Barbeck, Wm., On the Development of
Lemna Minor, 226. 230.
Bergh, R., On the Nudibranchiate
Gasteropod Mollusca of the North
• Pacific, with special reference to
those of Alaska, 9, 40.
Biological and Microscopical Section,
Report of, 415. .
Botanical Section, Report of, 418.
Borie, A. E., Announcement of death
of, 128.
Brewer, T. M., .•Vnnouncement of
death of, 12.
Budd, Dr. Chas. H., Announcement of
death of, 355.
Cardeza, .J. M., Fossil (?) Casts in
Sandstone, 280; Garnet mistaken
for Corundum, 295.
Chapman, H. C, On the Gestation and
Generative Apparatus of the Ele-
phant, 184, 158: On the Structure
of the Orang Outang, 159, IGO.
Conchological Section, Report of, 414.
Cresson, Ezra T., Election to Council,
357.
Elections during 1880, 428.
Entomological Section, Report of, 421.
Fisher, Jas. C, Announcement of
death of, 351.
Foote, A. E., A new locality for Anal-
cite, 252 ; On a probable Pseudor-
morphism of Gummite and Urano-
tile after Uraninite, 292 ; On a large
Sphene from Canada, 341 .
Fox, Wm. Logan, Announcement of
death of, 159.
Garrett, Andrew, The Terrestrial Mol-
lusca inhabiting the Cooks or Har-
vey Islands, 9, 158.
Genth, F. A., Jr., The so-called Emery-
Ore from Chelsea, Bethel Township,
Delaware Co., Pa., 311.
Gilbert, Wm. Kent. Announcement of
death of. 226.
Haines, R., Analysis of Philadelphite,
310.
Haldeman, S. S., Announcement of
death of. 341.
Hallowell. Morris, L., Announcement
of death of, 226.
Hartman, W. D, Description of a
Partula supposed to be New, from
the Island of Moorea, .225, 229; A
Bibliogrophical Catalogue of the
Genus Partula, 226.
Hays, Isaac, Resolution of Thanks for
Portrait of, 356.
Heilprin, A., On the Stratigraphical
Evidence afibrded by the Tertiary
Fossils of the Peninsula of Mary-
land, 20; On some Lower Eocene
Mollusca from Clarke Co., Ala., with
some points as to the Stratigraphical
position of the Beds containing them,
359, 364.
Hering, Constantine, Announcement of
death of, 830.
Hess, R. .J., Report of Biological and
Microscopical Section, 414.
Horn, Geo. H., Report of Correspond-
ing Secretary, 407.
Index to Genera, 451.
.leanes, Joshua, T., Announcement of
death of, 9 : Resolution regarding
Bequest of, 225.
Jefferis, W. W., A new locality for
Fluorite, 243 ; A new locality for
Amethyst, 280; A new Corundum
45(5
INDEX.
locality, 280; Menaccanite and Talc
from Maryland, 292; Sunstone in
Labradorite, 2.-t2.
Kelly, H. A., Sartorius Muscle of the
Gorilla, 128.
King, Dr. Wm. M., Announcement of
death of, 134.
Kingsley, .J. S., Carcinological Notes,
No 1, o4 ; Carcinological Notes, No.
2, Revision of the Gelasimi, 12, 1.35 ;
Carcinological Notes, No. 3, 179;
Carcinological Notes, No 4, 134,
187.
Kcenig, Geo. A., Notes on .Jarosite,
231.
Lautenbach, B. F., Announcement of
death of, 226.
Laporte. Count de Castelnau, An-
nouncement of death of, 150.
Lea, Isiac, Resolution of thanks for
Portrait of, 159.
Leidy, Jos., Notice of the Cruel Thread
Worm, Filaria immitis, of the Dog,
1" : On a Filaria reported to have
come from a Man, 130 : Remarks
on Pond Life, 156: Rhizopods in
the Mosses of the Summit of Roan
Mountains, N. C, 332, 333 ; Bone
Caves of Pennsylvania, 346 : The
Parasites of the Termites, 351, 354 ;
Remarks on Bathygnathis borealis,
351, 354 : Report" of Curators, 410.
Lewis, H. C, A new Polariscope, 241 :
A Garnet with inverted crystalliza-
tion, 241 ; A new locality for Sider-
ite. 242; Magnetite Markings in
Muscovite, 242 ; A new locality for
Asbolite, 243 ; Epidote in Molyb-
denite, 243 ; The optical characters
of some Micas, 244; On the Meas-
urement of Plane Angles, 252; On
an Exfoliated Talc. 252; Tin in
North Carolina, 253 ; On Siderophyl-
lite — a new mineral, 254; On Ster-
lingite and Damourite, 2.56 ; Vana-
dium in Philadelphia rocks, 256; A
new locality for Kp<omite, 257 ; The
surface geology of Philadelphia and
viciuity, 258 ; On the Bryn Mawr
Gravel, 277 ; On some enclosures in
Mica, 278; On Dendrites, 278; On
a Jurassic Sand, 279; The Minei-als
of Surry County, N. C, 280; A new
locality for Lignite, 281 ; On Ser-
pentine in Bucks Co., 281 ; The Iron
Ores and Lignite of the Montgomery
Co. Valley, 282; An enclo-ure in
Quartz, 292 ; On a new Fucoidal
plant from the Trias, 29:;; The
Trenton Gravel and its relation to
the Antiquity of Man, 296; Note on
Philadelphite — a new mineral, :>10 ;
On Philadelphite, 313; A Potsdam
Sandstone Outcrop on the S. Valley
Hill of Chester Valley, 329.
Lippincott, .loshua, Announcement of
death of, 355.
Lockington, W. N., On the Pacific
species of Caulolatilus, 13 ; On a
new species of Hemitripterus from
Alaska, 134, 233 ; Description of a
new species of Catostomus (C. cypho)
from the Colorado River, 156, 237.
McCook, Rev. H. C, Note on a new
Northern Cutting Ant (Atta septen-
trionalls), 859 ; The Shining Slave-
maker — Notes on the Architecture
and Habits of the American Slave-
making Ant (Polyergus lucidus),
376.
Martindale, Isaac C, Sexual variation
in Ca«tanea Americana, 351.
Meehan, Thos., On disarticulating
branches in Ampelopsis, 9 ; Germi-
nation in Acorns, 128; On the
Timber line of high mountains, 341 ;
Dimorphic flowers in Houstonia,
349; Cleistogamy in Oxalis aceto-
sella, 350; Sexual variation in Cas-
tanea Americana, 351 ; Rain Trees
— Note on Yucca gloriosa, 355 ;
Dioecism in Andromeda catesbtei,
356; Note on the Seed-ves-els of
Wistaria, 358 : Report of Botanical
Section, 418.
Mineralogical and Geological Section,
Proceedings of. 159, 241 : Report
of, 422.
Nolan, Edw. J., Report of Recording
Secretary, 406 : Report of Librarian,
408.
Officers for 1881,
Parker. Chas. F., Report of Curator in
Charge. 410.
Potts, Edw., Fresh-water Sponges of
Fairmount Park, 330 ; On Fresh-
water Sponges. 356
Proceedings of the Mineralogical and
Geological Section of the Academy,
for the years 1877, 1878 and 1879,
150, 241.
Rand, Theo. D., Serpentine belts of
Radnor Township, Del. Co., 225;
Change of Serpentine into Quartz,
241 ; A new locality for Millerite,
243 ; A new locality for Gypsum,
252 ; On a belt of Steatite and Ser-
pentine in Radnor, Del. Co., 273;
INDEX.
457
Chromite near Radnoi-, Pa., 27o ;
On Randite, 274; Some microscopic
enclosures in gems, 276 ; Potsdam
Sandstone near King of Prussia,
279; On a peculiar stratification in
Gneiss, 280; The northern belt of
Serpentine in Radnor Township,
2(io ; Report of Mineralogical and
Geological Section, 422,
Redfield, .J. H., On Rochelia patens,
131 ; On the timber line of high
mountains, 345 : Report of Botanical
Section, 418.
Report of Biological and Microscopical
Section, 413.
Report of Botanical Section, 418.
Report of Conchological Section, 414. .
Report of Corresponding Secretary, 407
Report of Curators, 410.
Report of Entomological Section, 421.
Report of Librarian, 408.
Report of Mineralogical and Geological
Section, 422.
Report of President, 397.
Report of Recording Secretary, 406.
Report of Treasurer, 42-5.
Report on Plants introduced by means
of the International Exhibition,
1876, 132.
Rice, J., Announcement of death of,
130.
Riding*, James, announcement of
death of, 331.
Ridings, .1. H., Report of Entomologi-
cal Section, 421.
Robert, S. R., Report of Conchological
Section, 414.
Roepper, W. T., Announcement of
death of, 156.
Ruschenberger, W. S. W., Report of
ihe President, 397.
Stauifer, .Jac, Announcement of death
of, 134.
Tyndall, Hector, Announcement of
death of, 134.
Vauj, Geo., Election to Council, 10;
Resignation from Council, 355.
Vodges, A. W., Description of a new
Crustacean from the Upper Silurian
of Georgia, with remarks upon Caly-
mene Clintoni, 128, 176.
White, S. S., Announcement of death
of, 9.
Wilcocks, Dr. Alex., Announcement of
death of, 356.
Willcox, Jos., Some new mineral locali-
ties, 312: Repor* of Mineralogical
and Geological Section, 422.
Wood, \V. M., M. D., Announcement
of death of, 130.
Proc.AlT.S.PMlajeeO.
PI. XIX
- ^•4-^^,>_,^ -•*' .•"
d*. *
4,^
3
ArchSt.Phila.
jyi.Ross. Lith.,
Mc Cook on Slave-making Ant.
Proc. fl,N,S.,PH[LA„1880.
Pl. XX.
.'I lS'ein 4 srs , ?"%
Lcv>t3 pc Co., Phila.. Kng's.
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Meehan, Thos. Germination in Acorns, .....
Leidy, Jos., M. D. On a Filaria reported to have conic from a
Man,
10
13
20
34
38
40
128
128
130
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CONTENTS — CONTINUED FROM NEXT PAGE.
Garnet mistaken for Corundum,
Trenton Gravel and its Relation to the
Rand. Theo. D. The Northein Belt of Serpentine in Radnor
Township,
Cardeza, Dr. J. M.
Lewis, H. C. The
Antiquity of Man, ....
Lewis, H. C. Note on Philadelphite — a new Mineral,
Haines, Reuben. Analysis of Philadelphite,
Genth, F. a., Jr. The so called Emery-Ore from Chelsea,
Bethel Township, Del. Co., Pa.,
Rand, Theodore D. Change of Serpentine into Quartz,
Lewis, H. C. A new locality for Sideritc,
Magnetite Markings in Muscovite,
A new locality for Asbolif e,
W. A new locality for Fluorite,
Epidote in ^Molybdenite,
Rand, Theo., D. A new locality for Millerite,
Lewis, H. C. The Optical Characters of some Micas,
A new locality for Analcite,
On the Measurement of Plane Angles,
On an Exfoliated Talc,
Tin in North Carolina,
D. A new locality for Gypsum,
On Siderophyllite —a new Mineral,
On Sterlingite and Damourite,
Vanalirum in Philadelphia Rocks,
A new locality for Epsomite,
The Surface Geology of Philadelphia and vicinity
Chromite near Radnor, Pa.,
On Randite,
D. Some Microscopic Enclosures in Mica,
On the Bryn Mawr Gravel, ■
On some Enclosures in Mica, .
On Dendrites,
On a Jurassic Sand, .
D. Potsdam Sandstone near King of Prussia,
WiLLCOX, Jos. Some new Mineral localities,
Lewis, H. C. On Philadelphite (Sp. Nov.),
Lewis, H. C. A Potsdam Sandstone Outcrop on the S
Hill of Chester Valley, ....
Potts, Edw. Fresh-water Sponges of Fairmount Park,
KoENiG, Geo. A. Notes on .Jarosite.
Leidt, Jos., M. D. Rhizopods in the Mosses of the summit of Roan
Mountain, N. C, .
FooTE, A. E. On Large Sphene from Canada,
Meehan, Thos. (Jn the Timber Line of High Mountain's,
Redfield, J. H. On the Timber Line of High Mountains,
Leidy, Jos., M. D. Bone Caves of Pennsylvania,
Meehan, Thos. Dimorijhic Flowers in Houstonia,
Meehan, Thos. Cleistogamy in Oxalis Acetosella, L.,
Martindale, Isaac C. Sexual Variations in Castanea Americana, •
Lewis, H. C.
Lewis, H. C.
Jefferis, W.
Lewis, H. C.
Foote, a. E.
Lewis, H. C.
Lewis, H. C.
Lewis, H. C.
Rand, Theo.
Lewis, H. C.
Lewis, H. C.
Lewis, H. C.
Lewis, H. C.
Lewis, H. C.
Rand, Theo. D.
Rand, Theo. D.
Rand, Theo.
Lewis, H. C.
Lewis, H. C.
Lewis, H. C.
Lewis, H. C.
Rand, Theo.
Valley
295
295
296
310
310
311
241
242
242
243
248
243
243
244
252
252
252
253
253
254
256
256
257
258
273
274
276
277
278
278
279
279
312
313
329
830
331
333
341
341
345
346
849
350
353
— 'Sfjl
CONTENTS.
Orang
Outang.
KiNGSLRY, J. S. Carcinological Notes, No. II.— Revision of the
Gelasimi. (Continued) ......
Leidy, Jos., M. D. Remarks on Pond Life, .
Chapman, H. C, M. D. On the Structure of the
(Plates 11 to 17). ....
VoGDEs, Anthony W. Description of a New Crustacean from the
Upper Silurian of Georgia, with remarks upon Calymene Clintoni,
Kingsley, J. S. Carcinological Notes, No. III.— Revision of the
Genus Ocypoda, ......
Kingsley, J. S. Carcinological Notes, No. IV.— Synopsis of the
Grapsidse, ••......
Rand, Theodoke D. Serpentine Belts of Radnor' Township, Dela-
ware Co., ••......
Allen, Harrison, M. D. On some Homologies in Bunodont
Dentition, ••......
Hartman, W. D., M. D. Description of a Partula supposed to be
New, from the Island of 3Ioorea, .....
Barbeck, Wm. On the Development of Lemna Minor. (Plate 18),
Lockington, W. N. Description of a new species of Hemitripterus
from Alaska, ........
Lockington, W. N. Description of a new species of Catostoraus
(Catostomus cypho) from the Colorado River,
Proceedings op the Mineralogical and Geological Section op
THE Academy op Natural Sciences op Philadelphia :
Lewis, H. C. A new Polariscope, . . . . .
Lewis, H.C. A Garnet with Inverted Crystallization, .
Jefferis, W. W. a new locality for Amethyst, .
Jepperis, W. W. a new Corundum locality,
Lewis, H. C. The Minerals of Surry Co., N. C,
Cardeza, Dr. J. M. Fossil (?) Casts in Sandstone,
Rand, Theo. D. On a peculiar Stratification in Gneiss, .
Lewis, H. C. A new locality for Lignite,
Lewis, H. C. On Serpentine in Bucks Co.,
Lewis, H. C. The Iron Ores and Lignite of the Montgomery
Co. Valley,^ •-.....
Lewis, H. C . An Enclosure in Quartz, ....
Jefferis, Wm. W. Menaccanite and Talc from Maryland,
Jepperis, Wm. W. Sunstone in Labradorite,
FooTE, A. E. On a probable Pseudomorphism of Gummite and
Uranotile after Uraninite, ......
Lewis, H, C. On a new Fucoidal Plant from the Trias,
153
156
160
176
179
187
335
336
339
330
333
387
341
341
380
380
380
380
380
381
381
388
393
393
393
393
393
^
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CONTENTS.
Martindale, Isaac C. Sexual Variations in Castanea Ameri-
cara. (Continued.) 352
Meehan, Thos. Sexual Variations in Plants, . . . 353
Meehan, Thos. Rain Trees. Note on Yucca gloriosa. . 355
Meehan, Thos. Dia3cism in Andromeda Catesbaii, Walt. . 356
Potts, Edw. On fresh-water Sponges. . ; . . 356
Meehan, Thos. Note on the Seed-A-essels of Wistaria. . 358
Allen, Harrison, M.D. The Phalanges of Bats. . . 359
McCooK, Rev. H. C. Note on a new Northern Cutting
Ant, Atta septentrionalis 359
Heilprin, Angelo. On some New Lower Eocene Mollusca
from Clarke Co., Alabama, with some points as to the
Stratigraphical Position of the Beds containing them.
(Plate 20.) 364
McCooK, Rev. H. C. The Shining Slavemaker.— Notes on
the Architecture and Habits of the American Slave-
making Ant, Polygergus lucidus. * (Plate 19.) . . 376
Allen, Harrison, M. D. On the Temporal and Masseter
Muscles of Mammals . 385
Report of the President. . 397
Report of the Recording Secretary 406
Report of the Corresponding Secretarj^ 407
Report of the Librarian • . 408
Report of the Curators. 410
Report of the Biological and Microscopical Section. . . 413
Report of the Conchological Section. . . . .414
Report of the Botanical Section 418
Report of the Entomological Section 421
Report of the Mineralogical and Geological Section. . . 422
Summary of the Report of the Treasurer 425
Election of Officers for 1881 428
Index to Genera 451
3.
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v;. p. K".2'-c, T.. 734 t. rz" ;Ari:rr/, st., philaow.
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