Full text of "Proceedings of the United States National Museum"

SMITHSONIAN INSTITUTION
UNITED STATES NATIONAL MUSEUM

COPY.

PROCEEDINGS

OF THE

UNITED STATES NATIONAL MUSEUM

VOLUME 90

UNITED STATES

GOVERNMENT PRINTING OFFICE

WASHINGTON : 1942

The scientific publications of the National Museum include two
series, known, respectively, as Proceedings and Bulletin.

The Proceedings series, begun in 1878, is intended primarily as a
medium for the publication of original papers, based on the collec-
tions of the National Museum, that set forth newly acquired facts
in biology, anthropology, and geology, with descriptions of new
forms and revisions of limited groups. Copies of each paper, in
pamphlet form, are distributed as published to libraries and scientific
organizations and to specialists and others interested in the different
subjects. The dates at which these separate papers are published are
recorded in the table of contents of each of the volumes.

The present volume is the ninetieth of this series.

The series of Bulletins^ the first of which was issued in 1875, con-
tains separate publications comprising monographs of large zoologi-
cal groups and other general systematic treatises (occasionally in
several volumes), faunal works, reports of expeditions, catalogs of
type specimens, special collections, and other material of similar
nature. The majority of the volumes are octavo in size, but a
quarto size has been adopted in a few instances in which large plates
were regarded as indispensable. In the Bulletin series appear vol-
umes under the heading Contrihution.s from the United /States National
Her-harimrb^ in octavo form, published by the National Museum since
1902, which contain papers relating to the botanical collections of
the Museum.

AlJEXANDER WeTMORE,

Assistant Secretary^ Smithsoniun Institution.

COPv

CONTENTS

Pages

Babiy, p. p. a new genus of psammocharid wasp from China.
No. 3119. October 24, 1941 ^ 543-546

New geuus : SinotocypJius.

New species: SinotocypJius chinensis.

Berry, Willard. Pamlico fossil ecliinoids. No. 3113. July

5, 1941 1 443-445

Clark, Austin H. Notes on some North and Middle American

danaid butterflies. No. 3118. November 4, 1941 ' 531-542

New species : Danaus candidtis.

New subspecies: Danaus plexipptis tohagi, D. p. portoricensis.

Cl^vkke, J. F. Gates. Revision of the North American moths
of the family Oecophoridae, with descriptions of new genera
and species. No. 3107. November 6, 1941 ^ 33-286

New genera : Martyrhilda, Bibarrambla, ApacJiea, Hinunacia,
Mathildana, Carolana, Anoncia, Antequera.

New species : Agonopterix arcucUa, A. oregonensis, A. fuscitermi-
nella, A. cajonensis, A. thelmae, A. dimorphella, A. costimacida,
A. antennaricUa, Martyrhilda sordidella, M. sphaeralceae, M.
hildaella, Semioscopis traunae, 8. mcdunnoughi, Depressaria
atrostrigella, D. palousella, D. eleanorae, D. whitmani, D. yaki-
niae, D. angustati, Psilocorsis caryae.

New variety : Agonopterix rosaciliella echinopanicis.

New combinations : Biharramhla aUenella (Walsingham), Apachea
barberella (Buscls), Eimmacia huachucella (Busck), Inga hu-
mata (Meyrick), /. concolorella (Beutenmiiller), /. canarieJla
(Busck), /. trigama (Meyrick), I. cretacea (Zeller), /. ohscitro-
maculella (Chambers), /. ciHella (Busck), Mathiidana newma-
nclla (Clemens), Carolana ascriptella (Busck), Anoncia episcia
(Walsingham), A. conia (Walsingham), A. fasciata (Walsing-
ham), A. orites (Walsingham), A. diveni (Heinrich), A. splia-
celina (Keifer), A. marinensis (Keifer), Antequera acertella,
(Busck).

Fisher, Walter K. A new genus of sea stars {Plazaster) from
Japan, with a note on the genus Parasterina. No. 3114.

June 18, 1941 1 447-456

New genus : Plazaster.

Fowler, Henry W. New fishes of the family Callionymidae,
mostly Philippine, obtained by the United States Bureau of
Fisheries Steamer Albatross. No. 3100. April 8, 1941 ^ 1-31

^ Date of publication.

IV PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Pages

New subfamilies : Amorinae, Draculinae.

New genus : DicalUonymus.

New species: Callionymus vmbrithoraof, C. scabriceps, C. hole-

ogenys, C. marisinensis, C. hudsoni, C. longi, C. irunneus, C.

pimctilateralis, C. keeleyi, C. distethommatus, C. leucobranchi-

alis, Synchiropus paUidus, 8. grinnelli, Elcutherochir mccad-

deni.

Gahan, a. B. a revision of the chalcid-flies of the genus
M onodontomenis in the United States National Museum.
No. 3116. August 19, 1941 ^ 461-482

New species: Monodontomerus dianthidii, M. cuhensis, M. viridi-
scapus, M. mexicanus, M. indiscreUis, M. mandibularHs, M. bakeri,
M. emarginatus, M. subobsolctus.

GiLMORE, Charles W. A history of the division of vertebrate

paleontology in the United States National Museum. No.

3109. August 5, 1941^ 305-377

Greene, Charles T. Two new species of cecidomyiid flies from

Phlox. No. 3120. October 30, 1941 ^ 547-551

New species : Hyperdiplosis phlox, Lestodiplosis macuUpennis.

Henderson, E. P. (See under Preston, F. W.) 387-416

Humes, Arthur G. A new harpacticoid copepod from the gill

chambers of a marsh crab. No. 3110. August 5, 1941 ^ 379-386

New species : Cancrhicola plumipes.
Linton, Edwin. Cestode parasites of teleost fishes of the

Woods Hole region, Massachusetts. No. 3112. July 15,

1941 ^ ____ 4i7_442

Preston, F. W., Henderson, E. P., and Randolph, James R.

The Chicora (Butler County, Pa.) meteorite. No. 3111.

June 17, 1941^ 387-416

Randolph, James R. (See under Preston, F. W.) 387-416

Stejneger, Leonhard. Notes on Mexican turtles of the genus

Kinostemon. No. 3115. July 25, 1941 ^ 457-459

New subspecies: Kinosternon cruentatmn consors.
Webber, Rat T. Synopsis of the tachinid flies of the genus

Tachinomyia^ with descriptions of new species. No. 3108.

June 30, 1941 ^ 287-304

New species: Tachinotuyia cana, T. acosta, T. nigricans, T. dako-
tensis.

Wetmore, Alexander. Notes on the birds of North Carolina.

No. 3117. October 31, 1941 ^ 483-530

New subspecies: Dryobates borealia hylonomus.

^ Date of publication.

ILLUSTRATIONS

PLATES

Following
page

1. North American Oecophoridae, Cosmopterygidae, and Ethmiidae:

Lateral aspect of heads 286

2. North American Oecophoridae : Lateral aspect of heads 286

3. North American Oecophoridae, Cosmopterygidae, Blastobasidae, and

Heliodinidae : Lateral aspect of heads 286

4. North American Oecophoridae and Cosmopterygidae: Wings and

lateral aspect of heads 286

5. North American Oecophoridae : Wing venation 286

6. North American Oecophoridae and Blastobasidae: Wing venation- 286

7. North American Oecophoridae, Cosmopterygidae, Ethmiidae, and

Heliodinidae: Wing venation 286

8. North American Oecophoridae and Blastobasidae: Ventral aspect

of male genitalia 286

9. North American Oecophoridae and Cosmopterygidae : Ventral aspect

of male genitalia 286

10, 11. North American Oecophoridae : Ventral aspect of male genitalia 286

12. North American Oecophoridae, Ethmiidae, and Heliomidae : Ventral

aspect of male genitalia 286

13. North American Oecophoridae and Cosmopterygidae : Ventral aspect

of male genitalia 286

14, 16. North American Oecophoridae : Ventral aspect of female gentalia__ 286
15. North American Oecophoridae and Heliodinidae : Ventral aspect of

female genitalia 286

17. North American Oecoplioridae and Ethmiidae : Ventral aspect of fe-

male genitalia and abdominal spines 286

18. North American Oecophoridae and Cosmopterygidae : Ventral aspect

of female genitalia 286

19. North American Oecophoridae, Cosmopterygidae, and Blastobasidae :

Ventral aspect of female genitalia 286

20-38. North American Oecophoridae : Ventral aspect of male genitalia 286

39-48. North American Oecophoridae : Ventral aspect of female genitalia,. 286

49. Main exhibition hall for fossil vertebrates, U. S. National Museum :

Diagonal view from the southeast 3-'0

50. Main exhibition hall for fossil vertebrates, U. S. National Museum :

View from the west, second floor 320

51. General view of the vertebrate paleontology laboratory, U. S. Na-

tional Museum 320

52. View of storage room for reserve or study collections of fossil ver-

tebrates, U. S. National Museum 330

53. Storage cases and cupboards for vertebrate collections, showing utili-

zation of corridor, U. S. National Museum 330

54. The two individuals of the Chicora meteorite fall 390

55. Map of terrain covered by witnesses of the Chicora meteorite fall 390

VI PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Following
page
5G. Map showing observations, in tliree dimensions, of the Chicora

meteorite fall 390

57. The appearance of the smoke cloud as seen from the wvjst at an

infinite distance (Chicora meteorite) 394

58. Drawing illustrating the absorption of kinetic energy of the meteorite

by the air (Chicora meteorite) 394

59. Fragment of the banded olivine chondrule and the fibrous upper half

of the field contains a fragment of a pyroxene chondrule (Chicora
meteorite) 410

60. Cestodes from fishes: Caryophyllaeus terebrans (Linton) ; Llyula

sp. ; Spathehothriiim simplex Linton ; Bothrimonus intermedius
Cooper; Bothriocephalus scorpii (Miiller) .. 442

61. Cestodes from fishes: Bothriocephalus scorpii (Miiller) ; B. claviceps

(Goeze) ; B. sp. ; Clestohothrium crassiceps (Rudolphi) 442

62. Cestodes from fishes: Ahothrium rugosum (Batsch) ; Ancistro-

cephalus aluterae (Linton) ; Bothriocephalus sp. ; Proteocephalus

macrocephalus (Creplin) ; Corallobothrium flmbriatum Essex 442

63, 64. Encope michelini L. Agassiz 445

65. Encope michelini L. Agassiz and Mellita guinquiesperforata (Leske) 445

66. Plazaster borealis (Uchida) 456

67. Lahidiaster annulatus Sladeu; L. radiosus Liitken 456

68. Plazaster borealis (Uchida) : Abactinal and lateral surfaces of five

rays and about half of disk 456

69. Plazaster borealis (Uchida) : Actinal surface of three rays and part

of disk 456

70. Patiria granifera Gray ; NepaMhia crassa (Gray) 456

71. Danaus plexippus plexippus, male ; D. p. megalippe, male ; D. p.

tobagi, new species, male 536

72. Danaus candidus, new species, male; D. plexippus portoricensis,

new subspecies, male 536

73. Danaus jamaicensis, male; D. berenice berenice, male; D. b. strigosa,

male 536

74. Danaus eresimus, female; D. cleothera, male 536

TEXT FIGURES

Page

1. CalUonymus umbrithorax, new species: Type 3

2. CalUonymus scabriceps, new species: Type 5

3. CalUonymus boleogenys, new species: Tyi)e 6

4. CalUonymus marisinensis, new species: Type 8

5. CalUonymus hudsoni, new species: Type 9

6. CalUonymus longi, new species: Type 10

7. CalUonymus brunneus, new species: Type 12

8. CalUonymus punctilateralis, new species: Type 14

9. CalUonymus keeleyi, new species: Type 15

10. CalUonymus schaapii Bleeker 17

11. Callioyiymus distethommatus, new species: Type 17

12. CalUonymus leucobranchiaUs, new species: Type, male 20

13. CalUonymus leucobranchiaUs, new species: Paratype, female 20

14. Synchiropiis pallidus, new species: Type 23

ILLUSTRATIONS VII

Following
page

15. Synchiropus grinnelli, new species : Type 25

16. Eleutherochir mccaddeni, new species: Type 28

17. Tachinomyia characters: Lateral aspects of anal forceps of male

genitalia (several species) ; hind tibia of T. apicata; apex of abdomen
with drawn genitalia showing formation of fourth and fifth sternites
in lateroventral aspect of T. apicata and T. variata 298

18. Cancrincola plumipes, new species 383

19. Sketch of clouds and explosion aa noted from "north side" Pittsburgh

(Chicora meteorite) 392

20. Nepanthia crassa (Gray) : Abactinal plates near base of ray 453

21. Nepanthia crassa (Gray) : 6 lateral plates of third and fourth series

above superomarginals near base of ray, to show pedicellariae 453

22. Nepanthia variabilis Clark: A plate from lateral area near base of ray

to show pedicellaria 454

23. Sinotocyphus chinensis, new species 1545

24. Two new flies reared from Phlox: nyperdiplosis phlox; Lestodiplosis

maculipennis 549

PROCEEDINGS OF THE UNITED STATES NATIONAL MUSEUM

by the

SMITHSONIAN INSTITUTION

U. S. NATIONAL MUSEUM

Vol.90 Washington: 1941 No. 3106

NEW FISHES OF THE FAMILY CALLIONYMIDAE, MOSTLY
PHILIPPINE, OBTAINED BY THE UNITED STATES BU-
REAU OF FISHERIES STEAMER "ALBATROSS"

By Henry W. Fowler

In my studies of the Indo-Pacific fishes of the family Calhonymidae,
with special reference to those of the Phihppines, I found a number of
interesting new species in the collections of the United States National
Museum. These are described and figured herein, with the same
methods employed as in an earlier paper.' Each figure is drawn in
lateral view, with an insert showing a dorsal view of the head and
predorsal region and a ventral view showing the under surface of the
head, together with the chest and breast. The head is measured to
the gill opening.

ANALYSIS OF GENERA

a'. Two distinct dorsal fins.

6'. Callionyminae. No supraorbital tentacle; body smooth.
cK Lateral line single.

dK Gill opening superior, above opercle; ventral without free ray.

eK Ventral with last membrane connected to middle of base of pectoral.
/'. Gill opening exposed, superior on neck; opercular membrane not
free posteriorly.

.g'. Lateral line simple Callionymus

g^. Lateral line jointed Paracallionymus

/-. Gill opening above or hidden by opercle; opercular membrane

forms free flap posteriorly Synchiropus

C-. Ventral with last membrane not united with pectoral; gill opening
hidden by opercle Eleutherochir

1 Proc. U. S. Nat. Mus., vol. 85, pp. 31-135, ftps. l-(il. 1938.

Z PROCEEDINGS OF THE XATIOXAL MUSEUM vol. 90

(P. Gill opening behind opercle; opercular membrane without free lobe;
ventral with free ray, posterior part of fin with ventral membrane

united to middle of anterior pectoral base Dactylopus

c2. Two lateral lines Dicallionymus

62. Amorinae. Each eye with long tentacle above; gill opening behind pre-

opercular spine; body spinulose Amora

a2. Draculinae. Only one dorsal, corresponds to second dorsal of other gen-
era Draculo

Callionyminae, new subfamily

Type genus. — Callionymus Linnaeus.

No supraorbital tentacle. Body smooth. Two distinct dorsal fins.

Genus CALLIONYMUS Linnaeus

ANALYSIS OF SPECIES

a'. Maxillary entire, without serrae.

61. First dorsal spines not prolonged, or but little longer than first ray of second
dorsal,

ci. Chest and breast dark or blackish brown umbri thorax

c2. Chest and breast pale to whitish like rest of under surface of body.

d>. Preopercular spine slender, straight, with 7 to 9 small antrorse serrae
along its inner edge.
eJ. Parietal region rather coarsely rugose; snout but little longer than

eye scabriceps

e^. Parietal region with more obscure and radiating striae; snout much

longer than eye . boleogenys

d?. Preopercular spine more robust, with 4 recurved denticles; eye

greatly exceeds short muzzle raarisinensis

d?. Preopercular spine with 2 recurved denticles; eye subequal with

muzzle '. hudsoni

■6*. Spines of first dorsal more or less prolonged or filamentous.

/'. Only first dorsal spine free terminally or prolonged.

g^. Eye subequal with muzzle; inner edge of preopercular spine
with 4 or 5 antrorse spinules.

h}. Preopercular spine inserted close behind eye longi

h?. Preopercular spine inserted well behind eye, or nearer gill

opening than eye brunneus

g^. Eye 2 in muzzle; preopercular spine with 7 antrorse spinules

along its inner edge punctilateralis

p. First 2 dorsal spines elongated; preopercular spine with 4 recurved

denticles keeleyi

p. First 3 dorsal spines elongated filaments; preopercular spine short,
curved, with 2 curved denticles on inner edge, -distethommatus
a^. Maxillary with outer edge denticulate in males, entire in females; preopercular
spine long, with 4 recurved denticles on its inner edge; chest with a con-
trasted white band on each side and approximating forward.

leu c ob ran chi alls

NEW PHILIPPINE CALLIONYMIDAE FOWLER 6

CALLIONYMUS UMBRITHORAX, new species

Figure 1

Depth 7; head 3}^, long as wide. Snout 3 in head; eye 3, subequal
with snout; maxillary not quite reaching eye, entire; mouth cleft short,
reaches halfway to eye, lower jaw very slightly shorter and included in
upper jaw; lips narrow; teeth very minute, villiform, in moderate
bands in both jaws; interorbital very narrow bony frenum, with
longitudinal groove. Preopercular spine large, nearly long as eye, ends

H.W.F.del,

Figure I. — Callionymus umhrithorax, new species. Type.

incurved in point followed by four recmwed denticles along its inner
edge, and a small subbasal denticle in front pointing forward.

Lateral line complete, high along side of back, simple.

D. IV-10, I fh-st spine 1% in head, fh'st ray l)i; A. 9, i, first ray 3;
caudal 2)^ in rest of fish, cuneate; least depth of caudal peduncle 5 in
head; pectoral 1^, rays 17; ventral rays I, 5, fin 1}^ in head. Anal
papilla elongate, conic, length half of eye.

Color in alcohol, rather light brovm above, with many obscure or
faint darker rings and arcs on trunk and tail above. Six dark- to
blackish-brown spots close below along lateral line. Various blackish-
brown spots scattered about upper surface of head ; pair on each side
of snout, 2 each side before preopercular spine, and several on opercle
especially distinct. Under surface of body largely whitish. Breast and
chest blackish brown, dark area extending between bases of vcntrals.

4 PROCGEEDINGS OF THE NATIOIfAL MUSEUM vol. 90

First dorsal shaded with dark brown, and large black ocellus on second
membrane. Second dorsal largely transparent, with dark basal spot
in front. Anal dark brown marginally, pale basally. Caudal white,
with 10 transverse rows of dark-brown spots, these chiefly on mem-
branes. Pectoral transparent, with 2 dark spots subbasally above.
Ventral becomes dark brown terminally, also with some ill-defined
scattered brown spots on rays.

Philippines. A species defined chiefly by the dark-brown color of
the breast of the male. It is quite different from Callionymus maldi-
vensis Regan,^ the males of which are said to have an oblong blackish
blotch on the tkroat. Callionymus margaretae Regan,^ based on
material 47 mm. long without caudal, is described with an oblong
blackish or triangular patch on the throat in the male. It differs in
the male having the first dorsal spine produced in a filament 2% in the
fish excluding the caudal. Its preopercular spine is said to be straiglit,
with serrated inner edge and forwardly directed spinous extension at
base in front.

Type.— U.S.N. M. No. 99433. D. 5345. Cliff Island, S. 43° E.,
4.4 miles (lat. 10°50' N., long. 119°22'03" E.), Malampaya Sound,
Palawan Island. December 26, 1908. In 7 fathoms. Length 47
mm. Male.

Paratype.— U.S.N. M. No. 99434. Same data. Length 38 mm.
Female.

(umbra, dark place + thorax.)

CALLIONYMUS SCABBICEPS, new species

Figure 2

Depth 6; head 3%, length iKo in width. Snout 2% in head; eye 2%,
subequal with snout, well protruded on upper profile of head; maxillary
reaches three-fourths to eye, entire; mouth cleft reaches halfway to
eye, upper jaws greatly protractile and mandible shorter or included in
upper jaw; teeth very fine, minute, villiform, in narrow bands in jaws;
no interorbital space, eyes closely set in apposition. Preopercular
spine long, slender, greater than orbit, inner edge with eight antrorse
spinules, and small spine at front of base directed forward.

Lateral line elevated, high along side of back, complete. Along
and above course of lateral line four short vertical bars of papillae or
pores, last over caudal peduncle before caudal fin base. Parietal
region of head with striate rugae, rather coarse and irregular.

D. IV-9, I, first spine 1% in head, first ray 1%; A. 8, i, first ray 2^;
caudal 2% in rest of fish, cuneate ; least depth of caudal peduncle 4 in

> Trans. Linn. Soc. London, ser. 2, voL 12, ZooL, p. 247, pi. 30, flg. 3, 1908 (type locality: Maldives, S.
Nilandu. in 30 to 36 fathoms).

3 Journ. Bombay Nat. Hist. Soc, vol. 16, No. 2, p. 326, pi. 3, fig. 3, 1905 (type locality; Muscat, in 15 to 30
fathoms).

NEW PHILTPPIN'E CALLIONYMIDAE FOWLER 5

bead; pectoral rays 19, fin 1 in head; ventral 3 in fish without caudal,
ra^/s I, 5.

Color in alcohol, back brown, with six darker saddlelikc cross bands,
though ill-defined and more or less obscured by paler rings or blotches,
the various shades producing more or less mottled appearance above.
Dark-brown blotches also on sides of head and prepectoral region.
Eye gray-brown, with darker mottling and spots. Under surface of
body uniform whitish. First dorsal with dark-brown spots, obscurely
ocellated with paler and with some h'regular white spots on last two
membranes. Second dorsal transparent, with three and sometimes

H. W. F. del

Figure 2.-^Callionymus scahriceps, new species. Type.

fom' longitudinal rows of black ocelli, subbasal and median series most
contrasted. Anal with rays tipped white and submarginallj'- dark
brown, shading paler basally on fin. Caudal transparent, with eight
transverse dark-brown bands on lower two-thirds of fin, and in each
band brownish-black spot, only distinct as fin is expanded. Pectoral
largely colorless. Ventral variegated with brown, as pale to whitish
spots, producing a mottled appearance.

Philippines. Distinguished by its combination of characters, espe-
cially the prominent long slender attenuated preopercular spine, which
is \% times the eye and furnished with 8 antrorsc spinules along its
inner edge, besides a small spine directed forward at its base in front.
Top of head rugose. Snout subequal with eye.

PROCEEDINGS OF THE NATIONAL MUSEUM

TV^e.— U.S.N.M. No. 99406. Jolo, Jolo Islands. February 8,
1908. Length, 58 mm.

Paratype.—lJ.S.l<iM. No. 99407. Surigao, Mindanao. Fifty-foot
seine. May 8, 1908. Length, 38 mm.

{scaber, rough +«€0aX77, head.)

CALLIONYMUS BOLEOGENYS, new species

Figure 3

Depth 7K; head S%, width 1% in its length. Snout 2}i in head; eye
2^0, 1% in snout; maxillary reaches four-fifths to eye, entire; mouth

H. W. F. del.

Figure 3. — Callionymus holeogenys, new species. Type.

cleft reaches two-fifths to eye, long premaxillaries well protruded and
a little protractile, lower jaw shorter; teeth very minute, villiform, in
narrow bands in jaws, none on palate ; interorbital narrow bony frenum,
with median longitudinal shallow concave groove. Preopercular
spine long, straight, slender, ends in short spine and also short basal
spine directed forward; eight fine antrorse serrae along inner edge.

Lateral line complete, high along side of back, and continued on
caudal base. Parietals with very fine or weak radiating striae.

D. IV-8, I, first spine l)i in head, first ray \)i; A. 7, i, first ray 2)^;
caudal 3}^ in rest of fish, cuneate, with upper rays graduated shorter;
least depth of caudal peduncle 4; pectoral \)i, rays 15; ventral rays I,
5, fin equals head. Anal papilla small, short, depressed or subconic
fleshy point.

NEW PHILIPPINE OALLIONYMIDAE FQi^VLER 7

Color in alcohol, back and upper surfaces very pale bro\\Ti, washed
with rose, with still paler to whitish blotches, ringed areas and irregular
and slightly darker arcs, circles, etc. Eye gray-white, with pink tinge
above, and variegated with gray to dark gray. Under surface of body
uniform whitish, with six longitudinal gray bands, variable and irregu-
lar. Caudal with seven rows of large blackish blotches on lower half.
Fins otherwise all pale to transparent.

Pliilippines. Characters of distinction are the long, slender pre-
opercular spine which equals the eye, protractile mouth, very pale
coloration washed with rose and with pale to whitish spots. The low
first dorsal has gray longitudinal lines and narrow bands.

TVpe.— U.S.N.M. No. 99408. Pandanan Island. Seine, 130 feet.
March 23, 1909. Length 51 mm.

Paratope. — U.S. 'NM. No. 99409. Same data. Length 32 mm.

(j3oXts, diirt-\-ykvv<i, cheek ; with reference to the slender preopercular
spine.)

CALLIONYMUS MARISINENSIS. new species

Figure 4

Depth 8}i; head 4, length 1}^ in its width. Snout 4 in head; eye
2Ko, about twice snout length, enters upper profile of head; maxillary
reaches front eye edge, miarmed; mouth cleft reaches halfway to eye,
lower jaw slightly shorter or included in upper jaw; teeth minute,
villiform, firm, simple, in narrow bands in each jaw; interorbital
narrow firm bony keel and separating eyes. Preopercular spine long
as eye, ends in rather short and slightly curved in spme, followed by
thi'ee or four more slightly recurved spines along its inner edge, first
longest and fourth (anterior) smallest; at base in front short spine
directed forward. Gill opening superior, small, about half long as
pupil.

Lateral line high along side of back, simple, complete to caudal base.

D. IV-9, I, fu'st spine 1^ in head, first ray IK; A. 9, i, first ray 2%;
caudal 3% in rest of fish, rounded behind; least depth of caudal pe-
duncle 4 in head; pectoral 1, rays 18, fin rounded; ventral rays I, 5,
long as head. Anal papilla short, depressed, pointed.

Color in alcohol, brown above, with innumerable darker brown
lines forming a marbled appearance, due to arcs, curves, rings, and
blotches darker and paler. Iris silvery gray. Under surface of body
uniform whitish. Dorsal dark or gray black with four whitish
oblique bands. Soft dorsal pale or whitish, with faint brown spots on
each ray. Anal pale or whitish. Caudal pale, with six brown spots
on each membrane, forming dark transverse bands in retracted fin.
Paired fins pale, pectoral with brown cross lines and ventral with
faint brown spots on each ray.

PROCEEDINGS OF 'THE NATIONAL MUSEUM

China Sea. Greatly resembles Callionymus hudsoni, but with an
entirely different preoperciilar spine long as eye.

Type.—V.S.NM. No. 99410. China Sea, vicinity of Hong Kong,
lat. 21°44' N., long. 114°48' E. In 34 fathoms. August 9, 1908.
Length 48 mm. [D. 5303.]

H. W. F. del

Figure 4. — Callionymus marisinensis, new species. Type.

Porafype.— U.S.N.M. No. 99411. Tinakta Island (N.), S. 80° W.,
3.3 mUes (lat. 5°12'30" N., long. 119°55'50" E.). February 21, 1908.
Length 36 mm [D. 5157.]

(marisinensis, of China Sea.)

CALLIONYMUS HUDSONI, new species

Figure 5

Depth Q%; head '3%, width 1%. Snout 3/8 in head (m profile) ; eye 3,
slightly greater than snout; maxillary not reaching eye, length 4K in
head; mouth cleft 5% in head from snout tip, extends halfway to eye,
lower jaw well protruded in front; teeth very minute, in narrow villi-
form band, firm, more easily felt than seen without a lens; interorbital
narrow, with large eyes set close. Preopercle ends in spine directed
up and back, with small one at base in front and two large, subequal,
recurved spines along its hind edge. Gill opening small, high, faces
upward.

Lateral line high, complete, simple.

NEW PHILIPPINE CALLIONYMIDAE FOWLER \)

D. IV-9, I, first 2 spines end in short filaments, first spine IY2 in
total head length, first ray 1^^; A. 7, i, first ray 2%; caudal 1, rounded
or convex behind; least depth of caudal peduncle 4%; pectoral iKo,
rays 18; ventral rays I, 5, fin 2% in fish without caudal.

Color in alcohol, very pale gray brown above, with slightly darker
variable spots, blotches, and bars. Iris silvery to coppeiy gray.
Entire under surfaces of head and body white. Fins all largely whitish.
First dorsal with dark or blackish brown blotch on most membranes
marginally. Few scattered pale brown spots on ventral rays.

^^^^-'^~—

H. W. F. del.

Figure 5. — Callionymus hudsoni, new species. Type,

Known only from the Philippines. Characterized by its moderate
caudal, large ventrals, and coloration, the inner edge of the preoper-
cular spine with only two recurved hooks.

r?/2Jg.— U.S.N.M. No. 99412. Pandanan Island. Seine, 130 feet.
March 23, 1909. Length 35 mm.

Para^?/pes.— U.S. N.M. No. 99415. Cabugao Bay, Catanduanes
Island. From seine. June 9, 1909. Length 40 mm.

U.S.N.M. No. 99417. Cebu market, Cebu. March 20, 1909.
Length 50 mm.

U.S.N.M. No. 99414.
1908. Length 38 mm.

U.S.N.M. No. 99416.

Mantacao Island. From seine. April 8,
Port San Vicente, Luzon side (beach).

November 18, 1908. Seine 130 feet. Length 29 mm.

PROCEEDINGS OF THE NATIONAL MUSEUM

U.S.N.M. No. 99413. Reef opposite Cebii, Cebu Island. AprU 7,
1908. Length 44 mm.

(Named for Capt. Charles B. Hudson, whose valuable figures of
Japanese dragonets were published in 1903.)

CALUONYMUS LONGI, new species

Figure 6

Depth 9; head 4Ko, length iKo in its width. Snout 2%, in head;
eye 2%, subequal with snout; maxillary nearly reaches orbit, length
3)8 in head; mouth cleft reaches tlu-ee-sevenths to eye, lower jaw

Tnrrt.

H.W.F.del

Figure 6. — Callionymus longi, new species. Type.

shorter or included m upper; teeth villiform, fine, close set, simple,
conic, in bands in jaws, narrowing along sides of jaws; interorbital
bony frenum separating large close set eyes. Preopercular spine very
long, slender, but slightly ciu*ved and ends behmd in a long slender
spine, fom* low broad spinules along its inner edge, and small
forward directed basal spine in front.

Lateral line complete, high along side of back, with loop above
close before caudal fin, and extendmg out on caudal base. Occiput
finely striate, with striae not greatly developed.

D. IV-9, 1, first spine slender, ends in free filament 3% in fish without
caudal; A. 9, i, first ray 2% m head measured to gill opening; least
depth of caudal peduncle 4K; pectoral Iji, rays 20; caudal 2% in rest

NEW PHILIPPINE CALLIONYMID'AE FOWLER 11

of fish; ventral 3%, rays I, 5. Anal papilla conic, pointed, nearly
long as pupil.

Color in alcohol, back and upper sui'faces light brown, warmer
tint on trunk and tad. Back also variegated with pale and dark
rings and marbling. Under surfaces of body white, with pale buff
and brown tints. Side of head with rather large brown spots inter-
spersed with white dots and small spots, and on under sides of head
variable, inclined, short white lines and bars. Eyes brown, variegated
with dark blotches above. First dorsal whitish, with black pale-edged
ocelli, on last two membranes largest and with dark to black bands.
Second dorsal pale or transparent with several brown spots on each
ray. Anal transparent. Caudal transparent whitish, with six trans-
verse bars made up of dark-brown spots, chiefly on upper half of fin.
Pectoral white, with small brown spots mostly on upper part of fin.
Ventral pale or whitish, with scattered brown spots on hind half of
fin.

Philippines. Known by the long preopercular spine 1 + % times eye,
ends in a backward dhected barbed point behind; four low broad
spinides along its upper edge and a small basal forward directed spine
in front. The first dorsal spine is an extended filament, little shorter
than head. Entire under surfaces uniformly white.

Type.— [J. S.N. M. No. 99418. Pajumajan Island (W.), S. 2° W.,
2 miles (lat. 5° 22' 55" N., long. 120° 15' 45" E.), Sulu Archipelago,
Tawi Tawi Group. In 34 fathoms. February 18, 1908. Length 98
mm. Biu-eau of Fisheries (No. 22922). [D. 5152.]

(Named for Bayard Long, of the Department of Botany, Academy
of Natiu-al Sciences of Philadelphia, to whom I am indebted for
many American fishes.)

CALLIONYMUS BRUNNEUS, new species

Figure 7

Depth 9)^; head 3%, long as wide. Snout 2% in head; eye 3%, 1%
in snout ; maxiUary reaches halfway to eye, length 4% in head ; mouth
cleft reaches two-fifths to eye, lower jaw included; premaxillaries very
protractile; teeth fine, conic, pointed, villiform, in bands in jaws,
broader anteriorly and on sides narrowing back; interorbital narrow
bony frenura, separating large closely set eyes. Preopercular spine
large, slender, slightly longer than eye, nearly straight, ends in
slender long spine, with five recurved denticles along inner edge, and
small subbasal denticle in front directed forward.

Lateral line complete, runs little high along side of back and out on
caudal base. Parietal region smooth, only some very feeble radiating
obscure striae, little visible except under a lens.

PROCEEDING'S OF THE NATIONAL MUSEUM

D. IV-9, I, first spine 1)^ in head, first ray Iji; A. 9,
caudal Sji in rest of fish, cuneate; least depth of caudal peduncle 4:% in
head; pectoral 1%, rays 19; ventral rays I, 5, fin 3% in fish without
caudal. Anal papilla depressed, narrowly triangular, two-fifths of eye.

Color in alcohol, back and upper surfaces brown, variegated with
slightly darker rings, arcs, circles, and spots, all producing little con-
trast. Under surface of body uniform whitish, with buff and pale-
brown suffusions. Several well-inclined dark-brown bars along lower
side of head and cheek. Opercle with brown spots, these extending on
connecting membrane below to ventral fin. Obscure brown spots

^.:^^^m^

H.W.F.del

Figure 7. — Callionymus brunneus, new species. Type.

little inclined back and below along side of tail. Eyes gray, brown
above with darker spots and blotches. First dorsal with pale brown
membrane, large black blotch on two last membranes, with several
dark streaks on fu'st two membranes. Second dorsal transparent,
greatly variegated with dark brown spots, ocelli and blotches. Anal
transparent basally, with blackish brown shading terminally on each
membrane and lower edge of each interradial membrane white in
front. Caudal pale to whitish with six transverse dark bars, mostly
made up of spots. Pectoral whitish, upper portion with small numer-
ous dark-brown spots. Ventral whitish basally, dark brown termin-
ally, upper outer surfaces also thickly spotted with still deeper brown
Philippines. A species easily known by its general brown appear-
ance. Its long slender straight preopercular spme ends in a slender

NEW PHILIPPINE CALLIONYMIDAE FOWLER 13

spine directed straight back, with five small antrorse spines along its
upper edge, and a small spine directed forward from its base. First
dorsal spine a little longer than tlie others in the male and less than
the head. Last two dorsal membranes black and anal with a black
siibmarginal line.

Type.— V.S.N M. No. 99419. Balayan Bay, Taal. January 19,
1908. From fishermen's seine. Length 80 mm.

Paratypes. — U.S.N.M. No. 99420. Same data as for type. Length
67 to 78 mm. Seven specimens.

U.S.N.M. No. 99421. Subig Bay, Subig, southern Luzon. From
seine. January 7, 1908. Length 34 to 52 mm. Two specimens.

(brunneus, brown.)

CALLIONYMDS PUNCTILATERALIS, new epecicB

Figure 8

Depth 8; head 4%, length %o of width. Snout 2 in head; eye 4, 2 in
snout; maxillary reaches two-thirds to eye, ridges and edges entire;
mouth cleft reaches halfway to eye, lower jaw slightly shorter and
included in upper when mouth is closed; teeth in villiform bands in
jaws, bands broken anteriorly; teeth strong, fu-m, conic, close set, and
in quite narrow bands on sides of jaws; interorbital narrow, concave,
width about equals diameter of pupil. Preopercular spine slender,
1% times eye, ends in point behind; seven recurved small denticles
along inner edge; small basal spine directed forward. Gill opening
small, an orbital diameter behind eye.

Lateral line complete, but little waved or undulate, high along side
of back. Parietals each with radiating striae.

D. IV-9, 1, first spine largely detached, prolonged, 3% in fish without
caudal, first ray 1% in head; A. 9, i, first ray 2J^; caudal 2% in rest of
fish; least depth of caudal peduncle 3K in head; pectoral rays 20, fin
iKo in head; ventral rays I, 5, fin 3% in fish without caudal. Anal
papilla slender, tapering, depressed, three-fourths length of orbit.

Color in alcohol, burnt umber or warm brown above, variegated
with many close-set intricated darker marblings, arcs, circles, and
vermiculating lines. Lower surfaces uniform whitish. All along
sides of head and body very numerous thickly strewn, dark to black-
ish-brown spots. On lower side of head and thorax spots larger or as
blackish blotches, especially contrasted with whitish of under surfaces.
Iris gray, variegated with darker spots on orbit above. First dorsal
with first and second membranes largely white, marked with some
black white-ringed ocelli; other membranes blackish, with four or five
oblique whitish lines. Second dorsal with graj^ to black ocelli, variable
on fm membranes. Anal largelj'' shaded gray-brown below or toward
margins. Caudal marked with large white-ringed ocelli on greater

PROCEEDINGS OP THE NATIONAL MUSEUM

upper portions, and fin otherwise with variable more or less transverse
gray waved parallel lines or narrow bands. Pectoral whitish, with
numerous small blackish-brown spots on upper part. Ventral whitish,
rays dark brown terminally and axillary membrane finely spotted with
black.

Philippines. Known by its small dark spots scattered over the side
of the body in the male.

H.W.F.del.

Figure 8. — Calltonymus punctilateralis, new species. Type.

Type.— One example. U.S.N.M. No. 99422. Tigbauan, Panay
Island. May 14, 1929. Male. Length 162 mm.

Paraty pes. —Four examples. U.S.N.M. Nos. 99423 and 99424.
Below mouth of Mindanao River, Cotabato, Mindanao. Seine.
May 20, 1908. Two males and two females. Length 52 to 84 mm.

(punctum, spot -\- lateralis, of the side.)

CALLIONYMUS KEELEYI, new species

Figure 9

Depth 7%; head 3%, long as wide. Snout 2% in head; eye 3%, IK in
snout; maxillary concealed; mouth cleft reaches halfway to eye, low^er
jaw little shorter; teeth villiform, fine, conic, simple, in rather wide
short bands in jaws, form narrow band along each side of jaw; man-
dible shallow, rami not elevated inside mouth; interorbital narrow
bonj'- ridge separating large close set eyes. Preopercular spine % of

NEW PHILIPPINE CALLIONYMIDAE ^Ol^VX,ER

eye, ends in small slightly recurved denticle, four more mostly larger
ones along inner edge, and small basal denticle in front pointing
forward.

Lateral line complete, high along side of back, simple, and extends
out on caudal base. Parietal region smooth.

D. IV-9, I, first 2 spines prolonged into long filaments, first longer
or 1% in fish without caudal, first ray VA in head; A. 9, i, first ray 2%;
least depth of caudal peduncle 4; pectoral 1%, rays 19; ventral rays I,
5, fin Sji in fish without caudal; caudal 1%. Anal papilla conic,
simple, depressed, long as pupil.

Color in alcohol, back and upper surfaces brown, variegated with
darker and paler marblings. Eight diffuse or obscure median darker

Figure 9. — Callioyiymus keeleyi, new species. Type.

blotches on back. Sides of head and body with more defined and
contrasted dark spots on sides. Eyes gray, with silvery tints and
darker markings. All under surfaces of body immaculate whitish.
First dorsal dark gray to gray-black, especially on second and thu'd
membranes; some paler longitudinal lines on fu-st two membranes.
Second dorsal transparent, with scattered dark spots or specks on
rays. Anal pale, dark to blackish brown m.arginally. Caudal whitish,
gray terminally, and with a dozen irregular transverse rows of dark to
kackish spots. Pectoral whitish, with numerous small, irregular,
blackish-brov/n spots above. Ventral pale to whitish basally, dark
to blackish brown termuially, and inner half of fin with connecting
membrane marked with small dark to blackish dots.

16 PROCEEDINOS OP THE NATIONAL MUSEUM vol. 90

Philippines. Known by the orbit 1}^ in snout and first dorsal with
two spines elongate filaments, fin dark, on last membrane black with
white lines. Caudal with a dozen rows of dark spots and anal border
black. Pectoral with many fine dark spots and ventral blackish
terminally.

Type.— U.S.N. M. No. 99425. Cebii. April 17, 1929. Lt. H. C.
Kellers. Length 80 mm.

(Named for Frank J. Keeley, of the department of mineralog}^,
Academy of Natm-al Sciences of Philadelphia.)

CALLIONYMUS SCHAAPII Bleeker

Figure 10

Callionymus schaapii Bleeker, Nat. Tijdschr. Nederl. -Indie, vol. 3, p. 455, 1852
(type locality: Banka). — Gunther, Catalogue of fishes in the British Museum,
vol. 3, p. 148, 1861 (copied). — Bleeker, Versl. Meded. Akad. Wet. Amster-
dam, ser. 2, vol. 14, p. 94, 1879 (Singapore; Banka; Java).

Depth 7% to 8K; head 4:% to 4%, length lYi in its width to width 1}^ in
its length. Snout 3 to 3% in head; eye 3}^ to 3%, subequal with snont;
maxillary reaches three-fifths to eye, entire; mouth cleft with rictus
halfway in snout, lower jaw little shorter than upper, which protrudes
in front; teeth very small, fine, simple, conic, close set, villiform, in
narrow bands in each jaw; interorbital narrow, concave, with median
longitudinal groove, bony width wide as pupil. Preopercular spine
large, bent, ends in small terminal point, also similar small basal
point in front directed forward, and along inner edge four or five
rather large recurved denticles.

Lateral line high along back to caudal base, simple and continuous
with loop above over caudal peduncle posteriorly, Parietals with
fine, narrow striae.

D, IV-9, I, first spines end in long filaments, second spine 1% to Sji
in fish without caudal, first ray 1% to 1% in head; A, 9, i, first ray 2%
to 2%; caudal 2% to 3 in rest of fish, convex behind; least depth of
caudal peduncle 3K to 4 in head; pectoral iKo to 1 + Ko, rays 20;
ventral rays I, 5, fin slightly longer than head to twice length of
head. Anal papilla simple, conic, three-sevenths of eye.

Color in alcohol, dull brown above, with many obscure darker
spots, specks, and blotches; also many arcs, rings, and circles on
trunk and tail above; on tail form four obscure dark saddles, Dark-
bro^vn spot about half size of orbit, on opercular flap. Dorsals and
anals largely transparent. Caudal with five transverse darker bars.
Pectoral with five or six dark transverse bars, more or less forming-
imperfect pale rings over upper part of fin. Ventral pale to whitish,
each ray with four or five brown spots.

KEW PHILIPPINE CALLIONYMIDAE FOWLER

H.W.F.del

Figure lO.—CaUionymus schaapii Blecker.

H.W.F.del.

Figure U. — Callionymus distethommatus, new species. Type.

274411—41 2

18 PROCEEDINGS OF THE KATIONAL MUSEUM vol. 90

East Indies, Philippines. Resembles Callionymus hudsoni and
Callionymus marisinensis but differs in having the preopercular spine

1 + /^ times the eye, first three dorsal spines filamentous and pro-
longed, also the head striate behind eye. The species may be known
by the very conspicuous ocellated black blotch below and before the
pastoral origin.

Fifteen examples: Buenavista, Guimaras Island. Seine. January
14, 1909. Length 54 to 88 mm.

One example. Cebu. April 14, 1921. Lt. H. C. Kellers. Length
70 mm.

Four examples. Iloilo market, Panay. March 28, 1908. Length
57 to 85 mm.

CALLIONYMUS DISTETHOMMATUS, new species

Figure 11

Depth 7M; head 4, broad as long. Snout o% in head; eye 3%, in-
vades upper profile of head; maxillary reaches below front of eye,
length 3K in head; mouth cleft reaches two-thirds to eye, greatly
protractile down and forward; teeth fine, villiform, in narrow bands
in jaws; interorbital very narrow frenum, or bony ridge with median
fine longitudinal groove, separating two large close-set eyes. Pre-
opercle with moderate spine, ends in recurved denticle and followed
by two others along hind edge; small basal spine in front directed
forward.

Lateral line high along upper side of body, simple, complete.
Parietal with very fine rugae.

D. IV-8, 1, spines all prolonged in filaments so third 1% in fish with-
out caudal, first ray 1% in head; A. 7, i, first ray 2%; least depth of
caudal peduncle 4; pectoral 1, rays 18; ventral rays I, 5, fin 2% in
fish without caudal; caudal 2%, cuneate. Anal papiUa conic, slender,

2 in orbit.

Color in alcohol, pale or light grayish above variegated with gray-
brown and brown, forming five transverse dark bars or blotches on
back. Snout pale above. Many small black to blackish-brown ocelli
scattered on sides of head and top of head and on fins, also two below
head at front of branchiostegal region. Iris gray and pearly. First
dorsal with large black ocellus on third membrane. Second dorsal
with some faint dark spots on rays. Anal blackish brown marginally.
Caudal pale to whitish, with large pearly ocellus basally and many
dark ocelli on lower and terminal part of fin. Pectoral largely whitish,
only very faint trace of gray spots. Ventral whitish, grayish
terminally and each ray with a few dark spots.

Philippines. A very handsome and ornamental species, with very
definite color pattern.

NEW PHILIPPINE CALLIONYMIDAE FOAVLER 19

Type.— U.S.N. M. No. 99426. Cebii market. August 28, 1909,
Length 69 mm. Male. [1835.]

Paratypes.—V .S.N .M. No. 99429. Cebu market, Cebu. ISIarch
20, 1909. Length 40 mm. Female.

U.S.N.M. No. 99431. Cebu m.arkct. March 28, 1909. Length
41 mm. Male.

U.S.N.M. No. 99428. Cebu market. August 26, 1909. Length
52 to 58 mm. Males. Three examples.

U.S.N.M. No. 99427. Cebu market. August 28, 1909. Length
45 to — mm. Males. Two examples.

U.S.N.M. No. 99432. Guijulugan, Tanon Strait, east coast of
Negros. Shore seme. April 2, 1908. Males. Length 48 to 59 mm.
Two examples.

U.S.N.M. No. 99430. Port Matalvi, off western Luzon. Seine
150 feet. November 23, 1908. Females. Length 35 to 42 mm.
Two examples.

{8is, twice + (jT^Oos, breast + ^fifia, eye; with reference to the
double ocellated spot on each side of breast.)

CALLIONYMUS LEUCOBRANCHIALIS, new species

Figures 12 (male), 13 (female)

Depth 8K to 10%; head 3%o to 4:%, long as wide. Snout 2% to 3 m
head; eye 2% to 3, subequal with snout; maxillary reaches seven-
eighths to eye or little below front of eye; outer ridge in males with row
of denticles, entire in female; mouth cleft horizontal, rictus three-
sevenths in space to eye; lower jaw slightly included in male, jaws
subequal or lower slightly protrudes in female; teeth fine, minute,
villiform, crowded and in narrow bands in both jaws; interorbital
narrow bony frenum, with longitudinal groove, bony width about half
diameter of pupil. Preopercular spine long as eye, ends in short
curved point behmd, with three or four recurved denticles along its
inner edge, and short denticle basal in front directed forward.

Lateral line complete, distinct, extends high along side of back
out on caudal base, with or without loop over caudal peduncle above
and just before caudal base. Parietal region smooth, radiating striae
feeble and not very distinct.

D. IV-9, I, or 10, I, spines all prolonged into slender free filaments
terminally reaching well mto caudal base in males, first dorsal spine
VA in head in females, first dorsal ray 1% to 1%; A. 9, i, first ray 2Ko
to 2% in males; caudal nearly long as rest of fish, in females caudal
3 m rest of fish; least depth of caudal peduncle 4% to 5 in total head
length; pectoral greater than head in males or 3% in fish without caudal,
subequal with head in females, rays 17 or 18; ventral rays I, 5, fin
3% in fish without caudal or 1}^ in head. Anal papilla long, slender.

PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

H.W.F.deI.

Figure 12. — Callionymus leucobranchialis. Type. Male.

xe-r-.

H.W.F.deI.

Figure 13. — Callionymus leucobranchialis. Paratype. Female.

NEW PHILIPPINE CALLIONYMIDAE^ FOWLER 21

Color in alcohol, light brown, on back with many faint and in-
distinct brown rings, arcs, and mottlings. Under surfaces largely-
whitish. Groove of lower lip each side of mandible, dark brown.
Throat, chest, and breast rather light to pale brown, with strongly
contrasted white gills. Iris silvery gray, with brown variable lines or
arcs. First dorsal variegated with white marks on dark gray -brown
backgroimd in male, less contrasted in female. Second dorsal pale or
transparent whitish, with dull brownish spots on rays. Anal similar
to second dorsal, or with oblique dark streaks below. Caudal pale or
whitish basally, with five to seven transverse series of large rounded
dark-brown spots, in closed fin forming transverse bands; terminally
fin darker or brown with still darker mottling or obscure blotches.
Pectoral pale, with about nine transverse brown lines. Ventral
whitish basally and marked with many inconspicuous brown spots;
fin gray black terminally.

Philippines. Apparently approaches Callionymus neptunia Scale ^
l)ased on examples 160 to 190 mm. It differs from that species in the
preopercular spine robust, but with the terminal spine small and
straight anterior basal spine directed forward well developed, and only
four large strong denticles along the upper edge of the spine. More-
over the four dorsal spines are greatly elongated and reach the basal
part of the caudal fin. The coloration is also different.

Type.— V.S.N.M. No. 99393. San Fernando Point Light, N. 39°E.,
8.4 miles (lat. 16°30'36" N., long. 120°11'06" E.), west coast of
Luzon. In 45 fathoms. May 11, 1909. Length 99 mm.

Paratypes.—V .S.N M. No. 99399. Same data as for type. Length
43 to 97 mm. Six specimens.

U.S.N.M. No. 99400. Bacoor, Luzon. Seine. June 15, 1908.
Length 60 mm.

U.S.N.M. No. 99395. Same data as precedmg. Four specimens.
Length 38 to 59 mm.

Other specimens. — U.S.N.M. No. 99403. Bolalo Bay, Malampaya
Sound, Palawan Island. Dynamite. December 21, 1908. Length
42 to 73 mm. Two examples.

U.S.N.M. No. 99394, Cavite and San Koque markets. June 27,
1908. Length 53 mm. 12819.

U.S.N.M. No. 99396. Cebu market. April 5, 1908. Length 73
mm. 8564.

U.S.N.M. No. 99401. Manila Bay, Luzon. December 12, 1908.
Length 42 mm.

U.S.N.M. No. 99402. Manila Harbor, Luzon. Seine of 34 feet.
March 16, 1908. Length 68 mm.

U.S.N.M. No. 99397. Outside harbor wall, Manila Bay, Luzon,
Dredge. December 12, 1907. Length 40 to 43 mm. Three examples.

* Philippine Journ. Sci., vol. 4, No. 6, p. 539, 1909 (type locality: Balayan Bay, Luzon).

22 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

U.S.N.M. No. 99404. Philippines. Length 48 to 65 mm. Two
examples.

U.S.N.M. No. 99405. D. 5442. San Fernando Point Light, N.
39° E., 8.4 miles (lat. 16°30'36" N., long. 120°11'6'' E.), west coast
Luzon. May 11, 1909. Length 28 to 73 mm. Five examples.

U.S.N.M. No. 99398. Sandakan Bay, Borneo. Seine. March 2,
1908. Length 58 to 66 mm. Three examples.

{XevKos, white +/8pd7xtQ;, gills)

Besides the above several other mteresting species in the collection
are:

CALUONYMUS SAGITTA Pallas

Callionymus sagitta Pallas, Spicilegia zoologica, vol. 8, p. 29, pi. 4, figs. 4-5, 1770'
(type locality: Amboyna). — Jordan and Richardson, Bull. U. S. Bur.
Fisher., vol. 27, p. 283, 1908 (Manila). — Borodin, Bull. Vanderbilt Marine
Mus., vol. 1, art. 2, p. 61, 1930 (Manila Bay).— Herrb, Fishes Herre 1931
Philippine Expedition, p. 94, 1934 (Manila).

22169. Davao, Mindanao. May 16, 1908. Length 58 rmn.
One example. Cebu. April 24, 1909. Lt. H. C. Kellers. Length
66 mm.

callionymus BENITEGURI Jordan and Snyder

Callionymus benileguri Jordan and Snyder, Proc. U. S. Nat. Mus., vol. 23, p,
370, pi. 17, 1900 (type locality: Tokyo Bay). — Jordan and Fowler, ibid.,
vol. 25, p. 956, 1903 (Misaki, Otaru, Wakanoura, Kobe, Hakodate, Aomori,
Hiroshima, Nagasaki, Matsushima Bay).— Jordan and Hubbs, Mem.
Carnegie Mus., vol. 10, No. 2, p. 317, 1925 (Toba, Kobe, Tatoku Island,
Mikawa Bay). — Schmidt, Trans. Pacific Committee Acad. Sci. U. S. S. R.,
p. 143, 1931 (NagasaKi).

Calliurichthys beniteguri Franz, Abh. Bayer. Akad. Wiss., vol. 4, Suppl. Band 1,
p. 84, 1910 (Yokohama).

U.S.N.M. No. 49402. Bay of Tokyo, Japan. K. Otaki. Length
185 mm. Type.

One example. Miyazaki, Japan. Rev. Cyrus A. Clark. Oberlin
College. Length 105 mm.

CALUONYMUS VALENCIENNEI Schlegel

Callionymus valenciennei Schlegel, Siebold's Fauna Japonica, Poissons, pts. 7-9,..
p. 153, pi. 78, fig. 3, 1845 (type locality: Nagasaki, Japan). — Jordan and
Hubbs, Mem. Carnegie Mus., vol. 10, No. 2, p. 317, 1925 (Kobe, Toba).

Callionymus valenciennesi Bleeker, Versl. Meded. Akad. Wet. Amsterdam, ser. 2,
vol. 3, p. 238, 1869 (Japan).

Calliurichthys valenciennesi Franz, Abh. Bayer. Akad. Wiss., vol. 4, Suppl. Band
1, p. 84, 1910 (Sagami Bay; Aburatsubo).

Callionymus japonicus (not Houttuyn) Valenciennes, Histoire naturelle de
poissons, vol. 12, p. 299, 1837 (Japan).

Callionymus simplicicornis Valenciennes, ibid., p. 303, 1837 (type locality:
Guam) .

Callionyjuus reevesi (part) Richardson, Zoolog}^ of the voyage of H. M. S. Sul-
phur, Ichthyology, p. 60, pi. 36, 1844 (female; on drawing made in Canton,
not type).

NEW PHILIPPINE CALLIONYMIDAE FOAVLER

Callionymus flagris Jordan and Fowler, Proc. U. S. Nat. Mus., vol. 25, p. 952,
fig. 7, 1903 (type locality: Tsuruga; Tokyo; Aomori; Kobe; Nagasaki; Owari
Bay; Matsushima Bay),

Two examples. China. A. de W. Sowerby. Length 64 to 75 mm.
Genus SYNCHIROPUS GILL

ANALYSIS OF SPECIES

o'. Second dorsal elevated; pectoral long, reaches halfway to caudal pallidus

a^. Second dorsal lower; pectoral moderate, reaches Yi to caudal griimellj

SYNCHIROPUS PALLIDUS. new species

Figure 14

Depth 6K; head Sji, wide as long. Snout 4 in head (in profile) to
gill opening; eye 2%, greatly exceeds snout; maxillary reaches below

mm.

H. W. F. del.

Figure 14. — Synckiropus pallidus, new species. Type.

front of eye, length 3% in head to gill opening, entire; mouth cleft
reaches tlu-ee-fourths in snout, lower jaw very slightly shorter; jaws
greatly protractile down and forward; teeth small, very close set,
villiiorm, in bands in jaws which narrowed posteriorly or along sides
of jaws; bony interorbital narrow, width two-thirds of pupil, shallowly
concave; preopercle spine less than e3^e, curved, ends in slender spine
and rather large forward curved denticle on inner upper edge; no
basal spinule in front.

24 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Lateral line complete, simple, extends on caudal base, and its course
little high along side of body. Parietals striate, rather smooth.

D. IV-7, I, first spine terminally free, equals head, first ray 2)2 in
fish without caudal; A. 7, i, first ray 3% in head to gill opening; least
depth of caudal peduncle 4; pectoral l}(o, rays 19; ventral rays I, 5,
fin 3Ko in fish without caudal; caudal 2, cuneate. Anal papilla very
short, conic, simple.

Color in alcohol, very pale brown above, whitish below. Iris copper
color, dark gray on eyes above. Fins all pale to transparent. First
dorsal with eight transverse and slightly inclined pearl-white lines
and about six on caudal.

Known by its very pale general coloration and the second dorsal
with eight transverse, pearly white, wide-spaced, narrow bands, the
caudal with six.

TV^e.— U.S.N.M. No. 99437. No. 1355. Length 190 mm. to end
of broken caudal.

(pallidus, pale.)

SYNCmROPUS GRINNELLI, new species

Figure 15

Depth 6%; head 3%, width 1. Snout 4% in head; eye 2%, greatly
exceeds short obtuse snout, with orbits conspicuously impinging on
upper profile of head; maxillary reaches below front part of orbit;
mouth cleft slopes little down and forward, rictus extending very
nearly opposite front edge of orbit and lower jaw slightly shorter than
upper; teeth small, simple, conic, close set or crowded, uniformly
villifoi-m and in moderate bands in jaws, which little broader anteri-
orly; no teeth on palate; interorbital narrow bony frenum separating
veiy large orbits and traversed by a shallow longitudinal groove.
Preopercular spine large, strong, ends in slightly upturned spine and
another on inner edge recurved, and both invested in connecting mem-
brane. Gill opening rather large, nearly opposite first dorsal origin.

Lateral line complete, high at first along back, posteriorly falls
until axial along middle of side of tail and caudal peduncle on to
caudal base. Parietal areas of cranium osseous, with pits and short
anastomosing radiating bony branches. Preorbital broadly tri-
angular, with fine radiating striae.

D. IV-9, I, first 2 spines with ends flexible and filamentous, first
spine IK in head, first ray 1%; A. 7, i, first ray 3^; caudal cuneate or
hind edge obliquely and slightly convex down to lower branched
rays, which longest and fin 3 in rest of fish; least depth of caudal
peduncle 5 in head; pectoral 1%, rays 20; ventral rays I, 5, length 1
in head Anal papilla very short, depressed, blunt, cutaneous flap.

NEW PHrLTPPIN'E CALLIONYMID.'LE FOi\\T.ER

Color in alcohol, pale buff -brown , inclining to whitish below. Upper
part of back with very faint traces of light blotches or spots, variable
though rather large. Iris grayish. First dorsal whitish, with large
black ocellus terminally on third membrane, and other membranes
with white and brownish shades. Second dorsal pale, with some
pure-white blotches medially or in upper half, in some lights these
markings reflected as brownish. A pale or transparent like second
dorsal, with subterminal dark-brown band, beyond or on edge of nar-
row white line. Caudal transparent with white markings as three
imperfect transverse bands sloping little back, as upper part of fin,

Synchiropus grinnelli, new species. Type.

and two large white basal blotches. Pectoral transparent, with
obscure w^hitish basally and tw^o very imperfect whitish bars above
transversely. Ventral pale to whitish, with two dark blotches on
each ray, outer or subterminal darker, and fin edge narrowly white.

A handsome species with large orbits greatly exceeding length of
snout. Preopercular spine half length of orbit, ends in upturned
spine, besides another spine above slightly bent forward, but no
lower basal spine. The species is known by its peculiar physiognomy,
the greatly enlarged orbits, and bony broad triangular postorbital.
Dorsal and anal both with membranes notched marginally.

T?/i>g.— U.S.N.M. No. 99436. D. 5475. San Bernardino Light,
S. 27° W., 11 miles (lat. 12°55'26" N., long. 124°22'12" E.), east

26 PROCEEDINGS OF 'THE NATIONAL MUSEUM vol. 90

coast of Luzon, San Bernardino Strait to San Miguel Bay, In 195
fathoms. June 24, 1909. [1699.] Length 118 mm.

(Named for the late Dr. Joseph Grinnell, of the Museum of Verte-
brate Zoology, Berkeley, Calif.)

Besides the above, there are several very handsome specimens of
the following:

SYNCmROPUS OCELLATUS (Pallas)

Callionymus ocellatvs Pallas, Spicilegia zoologica, vol. 8, p. 25, pi. 4, figs. 1-3,
1770 (type locality: Amboina). — Weber, Sihoga Expedition, vol. 57, Fische,
p. 524, 1913 (Beo; Kabaena Island; Ambon; Nusa Laut; Tiiir).

Synchiropus ocellatus Jordan and Richardson, Bull. U. S. Bur. Fisher., vol. 27,
p. 282, 1907 (1908) (Calaj^an).— Herre, Fishes Herre 1931 Philippine Exped.,
p. 94, 1934 (Dumaguete).

Callionymus pundulatus LACi^pfeoE, Histoire naturelle des poissons, vol. 2, pp.
328, 340, 1800 (type locality: No locality).

Three examples. Gubat Bay, Luzon. Tide pools. June 23,
1909. Length 31 to 73 mm. [1693, 1696.]

SYNCmROPUS SPLENDIDUS (Herre)

Callionymus splendidus Herre, Philippine Journ. Sci., vol. 32, p. 416, 1927 (type

locality, Bungau). — Whitley, Rec. Australian Mus., vol. 16, No. 4, p. 222, pi.

17, figs. 1 a-b, 1927 (Hayman Island reef, Whitsunday Group, Great Barrier

Reef, Queensland).
Synchiropus splendidus Giltay, M^m. Mus. Roy. Hist. Nat. Beige, Hors ser.,

vol. 5, fasc. 3, p. 84, 1933 (reference).

One example. Biri Channel. June 1, 1909. Dynamite. Length
70 mm. [1545.]

Genus ELEUTHEROCHIR Bleeker

Eleutherochir Bleeker, Versl. Meded. Akad. Wet. Amsterdam, ser. 2, vol. 14,
p. 103, 1879. (Type, Callionymus opercularioides Bleeker, orthotypic.)

BrachycalUonymus Herre and Myers, in Herre, Proc. Biol. Soc. Washington,
vol. 49, p. 12, 1936. (Type, BrachycalUonymus mirus B.erre = Callionymus
opercularioides Bleeker, orthotypic.)

There is hardly any question that the small specimens named as
BrachycalUonymus mirus Herre are the young of Eleutherochir opercu-
laris.

Eleutherochir differs from the other genera especially in its even
head with straight profile. The gill opening resembles that of Synchi-
ropus, but is larger and hidden under the opercle, which is extended
back in a free-pointed lobe.

ELEUTHEROCmR OPERCULARIS (Valenciennes)

Callionymus opercularis Valenciennes, Histoire naturelle des poissons, vol. 12,
p. (228) 305, 1837 (type locality: "L'embouchure de la riviere d'Arian-
Coupan").— Day, Fishes of India, pt. 2, p. 323, 1876 (Pondicherry; Madras).
— Weber, Sihoga Expedition, vol. 57, Fische, p. 522, 1913 (Menado; Ambon).

NEW PHILIPPINE CALLIONYMIDAE FOWLER 27

Synchiropus opercularis Gill, Proc. Acad. Nat. Sci. Philadelphia, 1858, p. 130

(reference). — Herre, Fishes Herre 1931 Philippine Expedition, p. 95, 1934

(Lemery) .
Callionymus opercularioides Bleeker, Nat. Tijdschr. Nederl.-Indie, vol. 1, p.

(30) 32, 1850 (type locality: Padang, Sumatra).
Eleutherochir opercularioides Bleeker, Versl. Meded. Akad. Wet. Amsterdam,

ser. 2, vol. 14, p. 103, 1879 (Sumatra; Bali; Celebes; Batjan; Ceram; Amboi-

na; Guebe).
Brachycallionymus ?mrus Herre, Proc. Biol. Soc. Washington, vol. 49, p. 12, 1936

(type locality: Celebes; Philippines).

Two examples. Camp Overton, Mindanao. August 15, 1909.
Electric light. Length 24 mm.

Nine examples. Malabang. May 2 1,1908. Electric light. Length
20 to 23 mm.

One example. Nato. June 18, 1909. Electric light. Length 24
mm.

One example. Taal. February 20, 1909. Electric hght. Length
20 mm.

U.S.N.AL No. 98827. Romblon. March 25, 1908. Albatross
Collection. Length 16 mm. Paratypes of Brachycallionymus mirus.
Two examples.

U.S.N.M. No. 98828. Nasugbu, Batangas Province. January 15,
1908. Albatross Collection. Length 15 mm. Paratype of B. mirus.

22170. Davao, Mindanao. Seme 150 feet. May 16, 1908. Length
59 to 85 mm. Six examples.

20824. Lingayen Gulf, Luzon. May 11, 1909. Seine 500 feet.
Length 77 mm.

One example. Nasugbu, Luzon. Seine. January 16, 1908.
Length 52 mm.

13959. PaluanBay, Mindoro. Seine 130 feet. December 11, 1908.
Length 37 to 70 mm. Three examples.

24132. Port San Vicente, Luzon. Seine 130 feet. November 18,
1908. Length 57 mm.

Two examples. San Vicente Harbor, Luzon shore. November 13,
1908. Seine. Length 54 to 59 mm.

ELEUTHEROCHIR MCCADDENI, new species

Figure 16

Depth 7; head 3, width 1%. Snout 0% m head; eye 3, slightly
greater than snout; maxillary reaches opposite front of eye, entire;
mouth but little inclined from horizontal, mouth cleft with rictus
reaching halfway to eye, and lower jaw well protruded in front; teeth
fine, conic, simple, close set, villiform, very minute, and set in inconspic-
uous narrow band above; lower teeth uniserial, rather large in com-
parison with upper, little curved in, situated along outer edge of

PROCEEDINGS OF THE NATIONAL MUSEUM

mandible and always visible, even in closed mouth ; rami of mandible
well elevated inside mouth; no teeth on palate; tongue small, entire,
rounded and free in front; intcrorbital narrow, bony width two-fifths
of eye, flattened. Preopercular spine curved back and up terminally,
ends in strong spine, and two spines curved up and back on its upper
edge. Opercular flap extends well back on pectoral base.

Lateral line complete, little high at first, becomes axial along side of
tail.

D. IV-9, I, spines not free, short, flexible, first 2% in total head
length, first ray 2; A. 10, i, first ray 2%; caudal 3K in rest of fish^

H.W.F.del

Figure 16. — Eleutherochir mccaddeni, new species. Type.

rounded convexly behind; least depth of caudal peduncle 3^5 in headp
pectoral 1%, rays 20; ventral rays I, 5, fin 1}^ in total head length.

Color in alcohol, back and upper surfaces dull brown, everywhere
more or less finely specked, dotted and spotted with darker, though
markings nowhere- much contrasted. On sides of head and opercle
they are larger, coarser, and fewer. All over back above traces of
dark arcs and circles, nowhere prominent. Entire under surfaces
whitish, with dull-gray or brown tints. Eyes dark gray above, with
dark blotches and mottling, pale to whitish below. First dorsal gray
black, with some obscure pale blotches on first membrane. Second
dorsal pale, with two dark spots on each ray. Caudal pale or light
brown, with five transverse dark bars, last subtenninal and empha-

NEW PHILIPPINiE CALLIOXYMIDAE FO'WLER 29

sized with several blackish-brown spots. Anal transparent. Pectoral
pale, with four very slightly darker transverse bars above. Ventral
pale.

Philippines. Known chieily by the long preopercle spine and white
€olor of the thorax.

Characterized by its dentition and especially by the long opercular
flap encroacliing on the pectoral fin base. Ventral without upper
basal membrane connecting fin to lower outer part of pectoral. It
differs from Eleutherochir opercularis in the projecting lower jaw, the
crooked or arched terminal preopercular spine, narrower bony inter-
orbital, and pale ventrals.

Type.— V.S.N. M. No. 99435. Hinunangan Bay, Leyte. Seine.
July 30, 1909. Length 52 mm.

(Named for David McCadden, formerly taxidermist in the Academy
of Natural Sciences of Philadelphia, to whom I am indebted for many
American fishes.)

DICALLIONYMUS, new genus

Type. — Callionymus goramensis Bleeker,

Body elongate, slender, depressed anteriorly. Head moderate,
depressed, as seen from above tapering into rather narrowly triangular
muzzle. Snout nearly pyramidal. Eye rather large, well invading
upper profile of head, upper velum of eye invading pupil above.
Mouth protractile down and forward, jaws subequal or lower little
shorter. Maxillary reaches front of eye, entire. Teeth in villiform
bands in jaws. Interorbital very narrow bony frenum, between large
close-set eyes. Long preopercular spine Iji times eye in length; with
small basal denticle directed forward, terminal spine upturned and
with five others along its upper edge. Gill opening small slit, con-
cealed by opercular flap, which rounded and not reaching pectoral
base. Upper lateral Ime normal, along upper side of back; lower
lateral fine extends only along lower side of tail, and like upper ex-
tended on caudal base. First dorsal spine elongate, filamentous,
reaches back well into middle of second dorsal. Last dorsal ray nearly
long as head and reaches well into caudal base. Edge of soft dorsal
entire. Anal with a slight notch along edge of each membrane.
Caudal but little longer than head, cuneate, lower median rays longest.
Pectoral broad, little shorter than head. Ventral longer than
pectoral, broad with inner membrane well connected up on pectoral
base. Anal papilla simple point. Coloration pale, variegated.

Distinguished from Synchiropus, with wliich it was later placed by
Bleeker, this genus is unique among the Callionymidae in the presence
of two lateral lines. It resembles Synchiropus in its gill openings.

(5ts, two + Callionymus; with reference to the two lateral lines.)

30 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

DICALLIONYMUS GORAMENSIS (Bleeker)

CalUonymus goramensis Bleeker, Nat. Tijdschr. Nederl. -Indie, vol. 15, p. 214,

1858 (type locality: Goram Island).
Synchiropus goramensis Bleeker, Versl. Meded. Akad. Wet. Amsterdam, scr. 2,

vol. 14, p. 101, 1879 (Goram).

Eleven examples. Pandanan Island. Seine 130 feet. March 23,
1909. Length 33 to 71 mm.

Amorinae, new subfamily

Type genus. — Amora Gray.

Long tentacle above each eye. Gill opening behind preopercular
spine. Body spinulose. Two distinct dorsal fins.

Genus AMORA Gray

Amora Gray, Illustrations of Indian zoology, Hardwicke, vol. 2, pi. 90, fig. 1,.

1833-34. (Type Amora tentaculata Gray, monotypic.)
Anaora -Gray, ibid., in directions for arranging the plates. (Type, Amora

tentaculata Gray.)

Body moderately elongate, little broader than deep. Head large,
depressed moderately. Snout rather long. Eye large, elevated, but
little advanced in head length. Mouth terminally inferior. Above
each eye a filament, twice eye length. Gill opening behind pre-
opercular spine. First dorsal begins over gill opening, with four
spines, about high as second dorsal. Second dorsal with eight rays,
last branched. Anal similar with seven rays; also last branched.
Paired fins little shorter than head.

This genus, long overlooked, was originally established on the
colored figure published and named by Gray as Amora tentaculata.
Bleeker in 1879 appears to have been the last author that accepted it.
Though he diagnosed the genus he failed to call attention to any of the
specific characters, some of which are of sufficient importance to be
readily made out. He mentioned that the body is spinulose, and this-
is better indicated by Gray's figure la, giving a dorsal view. This
suggests that CalUonymus Jimbriatus Herre,^ described with short
conspicuous cutaneous filaments on the body, and otherwise largely
in agreement, is a synonym of Amora tentaculata Gr&j.

AMORA TENTACULATA Gray

Amora tentaculata Gray, Illustrations of Indian zoology, Hardwicke, vol. 2, pi.

90, figs. 1, a-b, 1833-34 (type locality: Amboina). — Bleeker, Versl. Meded.

Akad. Wet. Amsterdam, ser. 2, vol. 14, p. 107, 1879 (reference).
Synchiropus tentaculatus Herre, Philippine Journ. Sci., vol. 35, p. 33, pi. 3, 1928'

(type locality, Puerto Galera).

One example. Port Caltom. December 15, 1908. Caught in
seine. Length 46 mm.

» The fishes of the Herre 1931 Philippine Expeditiorf, p. 94, 1934 (type locality: Sitanki Reef).

NEW PHILIPPINE CALLIONYMID'AE^ FOWLER 31

Two examples. Port Galera, Mindoro. June 9, 1908. Caught in
a seine. Length 36 to 48 mm.

One example. Romblon. March 26, 1908. Caught in seine.
Length 62 mm. [817].

19319. Sandakan Bay, Borneo. March 2, 1908. Caught in seine.
Length 39 mm.

[2158.] Great Tobea Island. Tide pool. December 15, 1909.
Length 42 mm.

Draculinae, new subfamily

Type genus. — Draculo Snyder.

Only one dorsal, corresponds to second dorsal of other genera.

U. S. eOVERNMENT PRINTING OFFICE] l»4l

PROCEEDINGS OF THE UNITED STATES NATIONAL MUSEUM

issued v.m*v\^^ Q^^S ^y '^*

SMITHSONIAN INSTITUTION

U. S. NATIONAL MUSEUM

Vol.90 Washington: 1941 No. 3107

REVISION OF THE NORTH AMERICAN MOTHS OF THE
FAMILY OECOPHORIDAE, WITH DESCRIPTIONS OF
NEW GENERA AND SPECIES

By J. F. Gates Clarke

INTRODUCTION

This study of the North American Oecophoridae was begun several
years ago at the suggestion of August Busck, of the United States Bu-
reau of Entomology and Plant Quarantine. In the beginning I had
intended to do only a specific revision of the genera Agonopterix and
Depressaria. It soon became apparent, however, that it would be nec-
essary to study carefully all the species known from North America
together with many from other parts of the world. Hence this paper
has become a revision of the entire family.

In North America the family Oecophoridae comprises a large group
of small to medium-sized moths most of which are inconspicuously
colored. A few, however, are brilliantly marked. The moths are
chiefly nocturnal, but some may be seen flying about in the bright sun-
light. Because of their retiring habits they are seldom seen and are
best secured by rearing the larvae.

The majority of the Oecophoridae are leaf and flower feeders in the
larval stage. Those that feed in the inflorescence usually attack plants
of the family Umbelliferae, while the leaf feeders attack a large variety
of plants in many families. Some are forest insects.

Besides the leaf and flower feeders there are others that feed on
stored products, such as dried foods, bulbs, and tubers. Some are
scavengers and feed in the refuse occurring in the nests of mice and
birds. Some live in the cracks of bark, others beneath the bark where

286614—41 1 33

34 PROCEEDINGS OF THE NATIONAL MUSEUM vol. so

they feed eitlier on refuse or bits of dried vegetable matter. One
{Martyrhilda sphaeralceae, new species) is a leaf miner.

Among the flower feeders the most important is Depressaria herac-
liana (Linnaeus) (the parsnip webworm), which does considerable
damage to commercially grown parsnips by destroying the flower-
heads and immature seeds. A closely related species, D. cinereo-
costella Clemens, has been recorded from caraway but is most
frequently found on closely related indigenous plants such as Sium
lineare Michx.

Among the forest insects we find Agonopterix argillacea (Walsing-
ham), which has been reared in quantity from willow {Salix spp.).
The species of the genus Psilocorsis attack beech {Fagus grandifolia
Ehrh.), oak [Quercus spp.), and pecan {Carya pecan Aschers. &
Graebn.) and hickory {Carya ovata (Mill.) Koch). P. caryae^ new
species, is of considerable importance in the South, where it is de-
structive to pecan. In the immediate vicinity of Washington, D. C,
the larvae of P. fagineUa (Chambers) do a great deal of damage
to beech.

In addition we have the European Carcina quercana Fabricius,
which is an oak feeder and has been introduced on Vancouver
Island, British Columbia, and Machimia tentoriferella Clemens,
which feeds on Castanea^ Juglans^ Quercus^ and Prunus.

The scavenger bulb moth, Hofmannophila pseudospretella (Stain-
ton), is recorded from stored bulbs and tubers of tulip, dahlia, and
gladiolus and from dried fruits and other stored products (see hosts
under species) and is a common pest in houses. It is found through-
out Europe and America and in New Zealand.

Endrosis lactella (Schiffermiiller) does considerable damage on the
Pacific coast and in Europe, feeding on stored cereals, fruits, and
other products.

The pupae are formed in debris, in leaves, or in hollow stalks and
do not protrude at the time of emergence of the moth. Many species
hibernate as adults.

Several species are of considerable economic importance. Notable
among these are Endrosis lactella (Schiffermiiller), Hofmannophila
pseudospretella (Stainton), and Depressaria heracliana (Linnaeus).

The distribution of the North American Oecophoridae appears to
center about the Southwestern States, particularly Arizona, Colorado,
and California. The genus Agonopterix^ the largest in the family, is
particularly well represented in western North America.

Since Busck^ published his paper on the North American moths
of this family no attempt has been made to revise the group. He

* Busck, A., A generic revision of American moths of the family Oecophoridae, with
descriptions of new species. Proc. U. S. Nat. Mus., vol. 35, pp. 187-207, 1908.

REVISION OF THE OECOPHORIDAE — CLARKE 35

recognized 19 genera and 121 species of North American Oecophori-
dae. In the Barnes and McDunnough list^ the authors followed
Busck in recognizing 19 genera but enumerated only 98 species, the
remainder being transferred to other families. Meyrick ^ recognized
17 North American genera and 104 species.

EXCLUSION OF UNRELATED FORMS

Within the family I recognize 22 genera and 117 North American
species. Six genera, 19 species, and 1 race are described as new in
this paper.

The genus Etidrosis is retained in the family with doubt. Busck *
transferred TAclonella to the Cosmopterygidae in 1932. I have re-
moved Eumeyrickia and Gerdana from the family and have placed
them in the families Ethmiidae and Blastobasidae, respectively. For
part of the species formerly placed in Borkhausenia I have erected the
genus Anoncia and have referred it to the Cosmopterygidae. For
Semioscopis acertella Busck I have erected the genus Antequera and
have placed it in the family Cosmopterygidae.

CHARACTERS OF THE FAMILY

Head usually smooth, with loosely appressed scales; often with
raised side tufts. Antenna simple to strongly ciliated, usually with
pecten on basal segment but frequently with pecten absent or slightly
developed. Labial palpus well developed, usually long, upwardly
curved, variously scaled; terminal segment acutely pointed. Maxil-
lary palpus short, filiform, appressed. Prothoracic and mesothoracic
legs normally slender, moderately long; posterior tibia with long rough
hairs above.

Fore wing with 12 veins (or 11 by coincidence of veins 7 and 8) ;
lb furcate at base; Ic always preserved, at least at margin; 7 and 8
stalked or coincident, 7 to costa, apex or termen.

Hind wing with eight veins or rarely seven by coincidence of veins
5 and 4 {Endrosis) ; veins 6 and 7 remote, subparallel; veins 3 and 4
normally stalked or connate (rarely separate); 8 free; 5 nearer to
4 than to 6 (except Oecopliora and Mathildana).

Male genitalia symmetrical; harpe with or without clasper, base
of harpe broadly attached; sacculus well defined; cucullus rounded
or pointed except in Carcina^ where it is greatly reduced and weakly
sclerotized. Anellus a simple plate or with moderately or well de-
veloped lateral processes. Aedeagus with very small "blind sac" or

* Barnes, W., and McDunnough, J., Check list of the Lepidoptera of Boreal America, pp.
160-1C2, 1017.

* Meyrick, E., in Wytsman, Genera Insectorum, fasc. 180, 1922.

* Busck, A., Proc. Ent. Soc. Washington, vol. 34, p. 19, 1932.

36 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

none. The entrance of the penis is dorsal, near the proximal end.
Vinculum bandlike or moderately produced. Gnathos present.^ Socii
and uncus present or absent. First and eighth segments simple or
with hair pencils.

Female genitalia : Ostium opening in median part of genital plate,
the latter simple or modified {Fabiola, Inga) ; signum present or
absent; abdomen weakly or strongly sclerotized; sometimes spinose.

The foregoing characters will serve to distinguish this family from
all other Lepidoptera except a few Blastobasidae and Ethmiidae. The
Blastobasidae may be separated from the Oecophoridae by the widely
separate veins 11 and 10 of the fore wing, the proximity of veins 2
to 10, and the thickening of the membrane below the costa (the "stigma"
of Zeller). Certain of the oecophorid genera have some of the char-
acters of the Blastobasidae while lacking others. Endrosis possesses
a spinose abdomen, a character frequently encountered in the Blasto-
basidae,** but lacks the characters listed in the foregoing paragi-aph
as being tyjDical of that family. The larva of Endrosis further sug-
gests Blastobasidae. The Ethmiidae may be distinguished from the
Oecophoridae (except Oecophora and Mathildana) by the proximity
of vein 6 to 6 of the hind wing; and the characteristic male genitalia,
which exhibit a primitive segmented type of harpe.

I have removed the genera Oecophora and BorJchausenia from our
American lists, as the only species formerly included are referable
to other genera.

For haydenella and pseudospretello I have resurrected the names
Chamhersia Riley and Hofmminophila Spuler, respectively. For
ascriptella Busck I have erected the genus Garolana. I propose the
genus Mathildana for newmaneUa and have restricted Oecophora
to hractella and its congeners. I retain all these in the family
Oecophoridae.

Larva.'' — ^With primary setae only (except in Apachea., which has
a few secondary hairs in group VII on most of the abdominal segments
and two or three on the pro thoracic shield) ; three setae on prespiracular
shield of prothorax ; setae IV and V approximately and directly below
or (frequently) ventrocephalad of the spiracle on proleg-bearing seg-
ments; on eighth abdominal segment seta III usually dorsocephalad
of the spiracle, rarely directly above, never dorsocaudad or wdth a

*The genera of the family fall into two groups on characters of the gnathos. Agonop-
terix, Bibarramila, Apachea, Depressaria, Seminscopis, Martyrhilda, Machimia, Himmacia,
and Psilocorsis fall into a group with a spiny gnathos. The remaining genera are without
spines on the gnathos. This suggests a division that may be of subfamily significance.

• The spinose abdomen is found in at least seven other oecophorid genera. They are
Martyringa, BorJihausenia, nofmannophila, Carolana, Pleiirota, Inga, and Semioscopis.

'' The setal characters of group VII should be used with caution as occasional abnormal
specimens occur in nearly all species. A hair may be missing from the group on either the
first or eighth abdominal segment, but only on one side of the insect. The opposite side
is nearly always normal.

REVISION OF THE OECOPHORIDAE — CLARKE 37

pigmented sclerotized ring surrounding the base of the tubercle; on
ninth abdominal segment seta I ventrocephalad of II, rarely approxi-
mate to II {Endrosis^ Eofmannophila)^ usually about equidistant
from II and III, III much nearer to IV-V than to I, IV and V closely
approximate, VI rarely (Psilocorsis) on the same pinaculum with
iy_Y^ sometimes approximate to VII, VII unisetose. Legs and pro-
legs normal. Crochets in a complete circle, biordinal and, occa-
sionally, irregularly triordinal, never in a penellipse (or a ring broken
outwardly).

Head with adfrontals extending to incision of dorsal hind margin
of epicranium; frons not extending to incision of hind margin; a
longitudinal ridge always present; epicranial seta L^ usually closer
to A^ than A^ is to A^ Ocelli normally six (reduced to 4 or 2 in
Hofmannophila and Endrosis) , arranged approximately in a parallelo-
gram ; 3 and 4 mostly in a straight line with 2 and 5 ; 3 and 4 closely
approximate.

Pupa.— Smooth, or pubescent. Body usually depressed. Epicranial
suture present; frontoclypeal suture not distinct for its entire length
or reaching the meson ; maxillary palpi present, large, usually reach-
ing proximolateral angles of maxillae; caudal portions of antennae
lying adjacent on the meson, separating at their distal ends to expose
metathoracic legs ; maxillae from one-half to two-thirds the length of
wings; labial palpi rarely exposed {Endrosis) . Prothoracic femora
sometimes exposed, often not. Abdomen with segments 4 to 6 movable
and with deep incisions between these segments on dorsal and ventral
surfaces; genital and anal openings slitlike in both sexes; no hooked
setae on ventral surface of ninth segment; cremaster absent or shortly
developed.

CLASSIFICATION

In the classification of this family venation, palpi, general habitus,
and genitalia have been employed in this revision. The genitalia are
especially helpful and have been used to separate species as well as
genera. I have found the anellus of the male genitalia to be particu-
larly useful, and in most cases it is possible to separate species as well
as genera by this structure.

The palpi are normally useful in the separation of genera but in a
few cases cannot be relied upon. The eyes and vestiture of the head
may be useful taxonomically, and Meyrick laid gi-eat weight on the
length of the ciliation of the antennae, but the modifications of these
characters appear to be too gradual for safe generic differentiation and
have not been employed in this paper.

Venation is usvially reliable within a genus, but occasionally unusual
venation is encountered. A large niunber of genera have nearly iden-
tical venation, but such cases are easily separable on genitalia or ex-

38 PROCEEDINGS OF THE NATIONAL MUSEUM vol. so

ternal characters. In a few the venation is unstable. This is par-
ticularly true of veins 2 and 3 of the fore wing of Semioscoj)is and
veins 3 and 4 of the hind wings of Agonopterix and Depressaria. The
genitalia of these three genera distinguish them, however, from all
others. The species fall, with one or two exceptions, into natural
groups on habitus as well as on structure, so that the definition of
genera is comparatively simple.

In a few genera it is not entirely clear what we are dealing with.
For example, in Depressaria there are five distinct species groups,
which may actually represent separate genera. The leptotaeniae-mul-
tifdae group is especially interesting and represents a series of
"species" that may represent only simple Mendelian variants of oTie.
species. As will be noted, in the proper place, all these except yahiTmae.
are of practically the same color and have closely similar habits and
hosts, yet aU have distinct genitalia.

In the genus Psilocorsis it is particularly difficult to separate many
of the species {re-ftexella-faginella group) , and here we are confronted
again with the problem of what actually constitutes a species.

These problems can be settled only by careful breeding, and in the
meantime we shall have to content ourselves with the supposition that
genitalic differences represent specific entities or genera depending on
their nature. Nevertheless, I strongly suspect that Mendelian varia-
tion may be transmitted to the genitalia as well as other structures and
that eventually we shall be able in most instances to use genitalic
characters for group separation only.

ACKNOWLEDGMENTS

I wish to express my gratitude to August Busck, of the U. S. Bu-
reau of Entomology and Plant Quarantine, for suggesting this most
interesting study and for much advice and help received through
correspondence while I was at Washington State College and since I
came to the Bureau of Entomology and Plant Quarantine. To Dr.
R. L. Webster, Head of the Department of Zoology, Washington State
College, my sincere thanks are due for the use of his library and the
release from many duties which enabled me to do research and spend
much time in the field ; to Carl Heinrich for the larval diagnoses and
notes and for many helpful suggestions and much useful criticism;
to Dr. J. McDunnough, Chief of the Division of Systematic En-
tomology, Department of Agriculture, Ottawa, Canada; to Dr.
Annette F. Braun, Cincinnati, Ohio ; to H. H. Keif er, Department of
Agriculture, Sacramento, Calif.; and Mrs. Florence Blackmore, Vic-
toria, British Columbia, for much material lent for study; to Dr.
Lincoln Constance, Department of Botany, Washington State Col-

REVISION OF THE OECOPHORIDAE — CLARKE 39

lege,« for his keen interest, the identification of host plants, and the
coUection of very interesting material; to Dr. Harry F. Clements, of
the same department,^ who kindly arranged for greenliouse facilities
for rearing purposes; and to Miss Grace Val Sisler for much kmd
help in cataloging and recording. In addition Dr. Braun kindly sub-
mitted notes on specimens in her collection and material for study. I
wish also to express my thanks to Nathan Banks, of the Museum of
Comparative Zoology, Cambridge; to E. T. Cresson, Jr., and J. A. G.
Rehn, of the Academy of Natural Sciences of Philadelphia, for mak-
ing it possible for me to study types of Chambers and Clemens re-
spectively; and to the National Research Council for a grant-in-aid,
which enabled me to obtain needed equipment and supplies and to
take many field trips in Wasliington, Idaho, and Oregon.

The drawings for this paper were made by the author. Whenever
possible the genitalia were figured from the type, either male or
female, and where the type was not available, from paratypes or
authentically determined specimens. I have been able to examine
the genitalia, both male and female, of all the genotypes and of either
male or female, or both, of all the species of Agonopterix and_ De-
pressaria except A. endryopa (Meyrick), A. murmumm (Meyrick),
D. nymphidia Meyrick, D. corystopa Meyrick, Borkhausenia axiiculata
Meyrick, and Paratheta astigmatica Meyrick. I am unable to recog-
nize these species.

For most of the Canadian records no reference is made to the sex

of the specimens. These records were sent to me by Dr. McDunnough.

The following keys to the genera include all genera discussed by

Busck (1908), with the addition of two old and six new genera

described in this paper.

ARTIFICIAL KEY TO THE GENERA

1. Basal segment of antenna with pecten (figs. 3, 4, 6, 9, etc.) 2

Basal segment of antenna without pecten (figs. 7, 10, 14, 18, etc.) 13

2. Second segment of labial palpus long, straight, porrect (fig. 6)

12. Pleurota (p. 230)

Second segment of labial palpus otherwise (figs. 3, 4, 9) T" oox

3. Antenna longer than fore wing 13- Carcina (p. 233)

Antenna shorter than fore wing ','"," nA^\

4. Veins 7 and 8 of fore wing coincident (fig. 55) 16. Decantha (p. 241)

Veins 7 and 8 of fore wing not coincident ^

5. Veins 2 and 3 of fore wing stalked (figs. 29, 44, 45) o

Veins 2 and 3 of fore wing separate

6. Costa of fore wing arched; costa of hind wing arched, veins 4

and 5 well separated (figs. 29, 45)

Costa of fore wing straight or slightly concave; costa of hind
wing concave, veins 4 and 5 closely approximate at bases
(f^g 44) 2. Martyrhilda (p. 12o)

• Now at the Department of Botany, University of California, Berkeley. Calif.

• Now at the Department of Botany, University of Hawaii, Honolulu.

40 PROCEEDINGS OF THE NATIONAL MUSEUM vol. oo

7. Abdomen flattened; labial palpus with well-developed brush on

second segment; hind wing lobed at anal angle... 1. Agonopterix (p. 43)
Abdomen not flattened; labial palpus without well-developed
brush on second segment; hind wing without lobe at anal angle.

3. Bibarrambla (p. 146)

8. Costa of hind wing excavated (fig. 51) 22. Endrosis (p. 262)

Costa of hind wing not excavated g

9. Second segment of labial palpus with brush (figs. 11, 16) 10

Second segment of labial palpus without brush (figs. 3, 4, 12) _. ._ H

10. Brush of labial palpus broadly triangular (fig. 16) 6. Apachea (p. 197)

Brush of labial palpus not broadly triangular (fig. 11)

5. Depressaria (p. 163)

11. Fore wing lanceolate, apex pointed; veins 3 and 4 of hind wing

stalked (fig. 49) 15. Carolana (p. 239)

Fore wing not lanceolate, apex not pointed; veins 3 and 4 of

hind wing connate or closely approximate (figs. 43, 46) 12

12. Veins 2 and 3 of fore wing remote (fig. 46) 19. Chambersia (p. 252)

Veins 2 and 3 of fore wing approximate (fig. 43)

21. Hofmannophila (p. 258)

13. Fore wing: Veins 8 and 9 out of 7 (fig. 39) 11. Martyringa (p. 228)

Fore wing: Vein 9 separate 14

14. Labial palpus with longitudinal stripes 9. Psilocorsia (p. 204)

Labial palpus without longitudinal stripes 15

15. Fore wing with veins 7 and 8 coincident (fig. 41) 17. Fabiola (p. 244)

Fore wing with veins 7 and 8 otherwise 16

16. Hind wing with vein 5 nearer to 6 than to 4 (fig. 50) 14. Mathildana (p. 236)
Hind wing with vein 5 nearer to 4 than to 6 17

17. Discocellulars of hind wing strongly outwardly oblique between

veins 4 and 6 (fig. 31) 8. Himmacia (p. 202)

Discocellulars not outwardly oblique between veins 4 and 6 18

18. Fore wing with vein 2 distant from 3 (figs. 33, 47, 56) " 19

Fore wing with vein 2 approximate (sometimes connate or

stalked with 3 in Semioscopis) to 3 (figs. 35, 37) 21

19. Costa of hind wing excavated (fig. 47) 20. Epicallima (p. 254)

Costa of hind wing not excavated (figs. 33, 56) 20

20. Fore wing broad; termen straight (fig. 33) 7. Machimia (p. 198)

Fore wing narrow, pointed; termen confluent with inner margin

(fig. 56) 18. Schiflfermulleria (p. 246)

21. Fore wing with costa and inner margin parallel; vein 11 from

middle of cell (fig. 37) 10. inga (p. 217)

Fore wing with costa and inner margin strongly divergent; vein

11 from well before middle of cell (fig. 35) 4. Semioscopis (p. 149)

KEY TO THE GENERA BASED ON MALE GENITALIA

1. Gnathos strongly spined (figs. 59, 62, 64, etc.) 2

Gnathos not spined (figs. 60, 76, 78, etc.) '" 10

2. Transtilla and anellus fused (fig. 86) 6. Apachea (p. 197)

Transtilla and anellus not fused (figs. 62, 64, etc.) 3

3. Anellus with long, free lateral processes (fig. 70) 7. Machimia (p. 198)

Anellus without long, free lateral processes (figs. 59, 64, etc.) 4

4. Uncus well developed (figs. 59, 64) '_ " 5

Uncus absent, or if present, poorly developed (figs. 62, 65, 67, etc.) 6

5. Clasper present (fig. 64) 8. Himmacia (p. 202)

Clasper absent (fig. 59) 9. Psilocorsis (p. 204)

REVISION OF THE OECOPHORIDAE — CLARKE 41

6. Sacculus extended as a furcate process or as a broad truncated

expansion with a small toothlike clasper; or vesica armed
with a forked or strongly curved single cornutus. 4. Semioscopia (p. 149)
Sacculus and cornuti not as above 7

7. Clasper absent or present; if present then the aedeagus twisted;

or sacculus with process from base; or aedeagus with basal

process 5. Depressaria (p. 163)

Clasper always present but without the above combinations

(figs. 62, 65, 67) 8

8. Clasper divided (fig. 67) 2. Martyrhilda (p. 125)

Clasper simple (figs. 62, 65) 9

9. Uncus and socii fused forming hood (fig. 65) 3. Bibarrambla (p. 146)

Uncus (if present) and socii not fused (fig. 62) 1. Agonopterix (p. 43)

10. Anellus with lateral processes (figs. 60, 61, 62, etc.) 14

Anellus without lateral processes (figs. 69, 72, 78, 80) 11

11. Clasper present (figs. 78, 80) 12

Clasper absent (figs. 69, 72) 13

12. Clasper concave distally; aedeagus coiled, bandlike basally

(figs. 78, 78a) 21. Hofmannophila (p. 258)

Clasper never concave distally; aedeagus never coiled or band-
like basally (figs. 80, 80a) 10. Inga (p. 217)

13. Gnathos flat, broad, shovel-shaped; vinculum narrowly

rounded (fig. 69) 15. Carolana (p. 239)

Gnathos narrowed distally; vinculum strongly produced an-
teriorly (fig. 72) 11. Martyringa (p. 228)

14. Extension of sacculus with tuft of strong, curved setae (fig.

71) 13. Carcina (p. 233)

Sacculus without such tuft 15

15. Sacculus with pointed, curved or nearly straight, free extension

(figs. 60, 61, 66, 81) 16

Sacculus without such free extension (figs. 74, 76, 79, 82) 19

16. Vinculum spatulate (fig. 60) 22. Endrosis (p. 262)

Vinculum rounded (figs. 61, 66, 81) 17

17. Vesica unarmed (fig. 61a) 17. Fabiola (p. 244)

Vesica strongly armed (figs. 66a, 81a) 18

18. Lateral processes of anellus flattened and expanded distally

(fig. 66) 16. Decantha (p. 241)

Lateral processes of aneUus not appreciably flattened; pointed

distally (fig. 81) 20. Epicallima (p. 254)

19. Vesica strongly armed (fig. 74a) 14. Mathildana (p. 236)

Vesica unarmed (figs. 76a, 79a, 82a) 20

20. Lateral processes of anellus sharply pointed; aedeagus small,

weak (figs. 79, 79a, 82, 82a) 21

Lateral processes of anellus dilated distally; aedeagus stout

(figs. 76, 76a) 12. Pleurota (p. 230)

21. Lateral processes of anellus very broad basally; gnathos pointed

(fig. 79) 19. Chambersia (p. 252)

Lateral processes of anellus of about equal width throughout;

gnathos not pointed (fig. 82) 18. SchiflEermiilleria (p. 246)

KEY TO THE GENERA BASED ON FEMALE GENITALIA

1. Anterior apophyses branched (figs. 101, 116) 2

Anterior apophyses not branched (figs. 95, 97, etc.) 3

42 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

2. Ovipositor always extended after death; ductus bursae not con-

voluted (fig. 116) 22. Endrosis (p. 262)

Ovipositor not extended after death; ductus bursae convoluted

(fig. 101) 21. Hofmannophila (p. 258)

3. Bursa copulatrix single (figs. 97, 99, etc.) 4

Bursa copulatrix double (fig. 105) 9. Psilocorsis (p. 204)

4. Ductus bursae with saclike evagination from ventral surface

(fig. 118) 14. Mathildana (p. 236)

Ductus bursae without such evagination 5

5. Inception of ductus seminalis at, or anterior to, middle of ductus

bursae (figs. 87, 91, 95, 98, 115) 6

Inception of ductus seminalis posterior to middle of ductus

bursae (figs. 109, 110, etc.) 10

6. Signa several (fig. 87) 12. Pleurota (p. 230)

Signum, if present, single 7

7. Signum a weakly developed toothed plate (figs. 91, 115) 8

Signum absent or, if present, not as above 9

8. Ductus bursae membranous (fig. 115) 19. Chambersia (p. 252)

Ductus bursae at least partly sclerotized (fig. 91)--- 16. Decantha (p. 241)

9. Genital plate strongly convex, protruding (fig. 95) 17. Fabiola (p. 244)

Genital plate convex but not protruding (figs. 98, 213-215)

18. SchiflPermiilleria (p. 246)

10. Ductus bursae strongly sclerotized, broadened and somewhat

flattened posterior to inception of ductus seminalis (figs.

90, 92, 99) 11

Ductus bursae otherwise 13

11. Signum present; sclerotized portion of ductus bursae armed

on inner surface (fig. 99) 20. Epicallima (p. 254)

Signum absent; ductus bursae unarmed (figs. 90, 92) 12

12. Ostium protruding; ventroanterior edge strongly sclerotized

(fig. 90) 11. Martyringa (p. 228)

Ostium not protruding; ventroanterior edge membranous (fig.

92) 7. Machimia (p. 198)

13. Signum very large, broadly oval or elongate (fig. 100)

2. Martyrhilda (p. 125)
Signum, if present, otherwise 14

14. Signum absent (figs. 89, 229, 240, 272) 15

Signum present (figs. 96, 102, 109, etc.) 16

15. Area surrounding ostial opening always membranous (figs. 89,

206-210) 10. Inga (p. 217)

Area surrounding ostial opening always sclerotized (figs. 229,

240, etc.) 1. Agonopterix (part) (p. 43)

16. Genital plate dilated ventrolaterally; ostium transverse, slit-

like (fig. 88) 13. Carcina (p. 233)

Genital plate otherwise 17

17. Signum a toothed plate (figs. 102, 103, 109, etc.) 18

Signum a small plate with median keel (fig. 110) 8. Himmacia (p. 202)

18. Ductus bursae armed with small teeth for much of its length

(fig. 96) 15. Carolana (p. 239)

Ductus bursae otherwise 19

19. Signum a distinct cross; ostium very large (fig. 94) 6. Apachea (p. 197)

Signum and ostium otherwise 20

REVISION OF THE OECOPHORIDAE — CLARKE 43

20. Signum a moderately large plate with two or three large strong

teeth (fig. 109) 3. Bibarrambla (p. 146)

Signum otherwise 1. Agonopterix (part) (p. 43);

5. Depressaria (p. 163); 4. Semioscopis (p. 149)

1. Genus AGONOPTERIX HUbner

Plate 2, Figube 13 ; Plate 6, Figubb 45 ; Plate 8, FiGxmES 62, 62a ; Plate 17,

FiGUBB 103

Agonopterix Hubneb, Verzeichniss bekannter Schmetterlinge, p. 410, 1826. —
PiEBCE and Metcalfe, The genitalia of the British Tineina, pp. 35-38, 1935.
(Genotype: Pyralis ocelhina Fabricius, Systema entomologiae, p. 652, 1775.)

Agonopteryx Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198, 1908.— Walsingham,
Biol. Centr.-Amer., Lepidoptera-Heterocera, vol. 4, pp. 135-136, 1912.— Barnes
and Busck, Contr. Lepid. North America, vol. 4, pp. 231-233, 1920.— Sasscer,
Journ. Econ. Ent, vol. 13, p. 183, 1920.— Bbaun, Proc. Acad. Nat Sci. Phila-
delphia, vol. 73, pt. 1, p. 10, 1921.— Busck, Can. Ent., vol. 53, pp. 277-278,
1921.— FoEBES, Cornell Univ. Agr. Exp. Stat., Memoir 68, pp. 236-241, 1923.—
Clarke, Can. Ent., vol. 65, pp. 84-85, 1933.— McDunnough, Can. Ent., vol.
67, pp. 74-75, 1935.

Agonopterys Ely, Proc. Ent. Soc. Washington, vol. 12, p. 68, 1910.

Agnopteryx Babnes and McDunnough, Check list of the Lepidoptera of Boreal
America, p. 160, 1917.

Epeleustia Hubner, Verzeichniss bekannter Schmetterlinge, p. 410, 1826. —
Walsingham, Biol. Centr.-Amer., Lepidoptera-Heterocera, vol. 4, p. 136,
1912.— IMeybick, in Wytsman, Genera insectorum, fasc. 180, p. 169, 1922.
(Genotype: Tinea- hypericella Hiibner, Sammlung europaischer Schmetter-
linge, vol. 8, fig. 441, 1796.)

Pinaris Hubneb, Verzeichniss bekannter Schmetterlinge, p. 411, 1826. — Walsing-
ham, Biol. Centr.-Amer., Lepidoptera-Heterocera, vol. 4, p. 135, 1912. — Mey-
BiCK, in Wytsman, Genera insectorum, fasc. 180, p. 169, 1922. (Genotype:
Tinea gilvella Hiibner, Sammlung europaischer Schmetterlinge, vol. 8, fig. 96,
1796; synonym of Agonopterix areneUa Schiffermiiller.)

Tichonia HiJBNER, Verzeichniss bekannter Schmetterlinge, p. 412, 1826. — Walsing-
ham, Biol. Centr.-Amer., Lepidoptera-Heterocera, vol. 4, p. 136, 1912. — Mey-
EiCK, in Wytsman, Genera insectorum, fasc. 180, p. 169, 1922. (Genotype:
Phalaenae Tineae atomella Schiffermiiller, Systema tisches Verzeichniss der
Schmetterlinge der Wiener Gegend, p. 137, 1776. )

Haemylis Teeitschke, Die Schmetterlinge von Europa, vol. 9, p. 235, 1832. —
Walsingham, Biol. Centr.-Amer., Lepidoptera-Heterocera, vol. 4, p. 136,
1912. — Meyeick, in Wytsman, Genera insectorum, fasc. 180, p. 169, 1922.
(Genotype: Haemylis assitnilella Treitschke, Die Schmetterlinge von Europa,
vol. 9, p. 259, 1832.)

Head with appressed scales ; tongue developed ; antenna ciliated or
simple; basal segment elongate, with pecten; labial palpus long, re-
curved; second segment with well-developed furrowed brush; terminal
segment shorter than (or rarely as long as) second segment, sometimes
roughly scaled, acute. Thorax smootlily scaled to strongly crested.

Fore wing elongate (narrow to moderately broad) ; 12 veins; 2 and 3
stalked ; 7 and 8 stalked ; 7 to costa or apex.

44 PROCEEDINGS OF THE NATIONAL MUSEUM vol. oo

Hind wing as wide as or wider than fore wing, with costa nearly
straight, termen evenly rounded; 8 veins; 3 and 4 connate or short-
stalked; 5 curved, approximate to 4; 6 and 7 subparallel. Abdomen
flattened.

Male genitalia. — Harpe ample; clasper present, simple. Anellus
without well developed fingerlike processes. Vesica with or without
cornuti. Socii fleshy. Uncus reduced or lacking.

Female genitalia. — Bursa copulatrix with or without signum ; ductus
bursae membranous throughout (except fuVva).

Larva. — Ninth abdominal segment with setae I and II well separated
(I as near or nearer to III than to II) : seta VI not on a single pinacu-
lum with IV and V but not approximate to VII. Setal group VII
bisetose on first and seventh abdominal segments, unisetose on eighth
and ninth abdominal segments, normal (trisetose) on proleg-bearing
abdominal segments. Ocelli normal. Submentum without sclerotized
pit.

Pupa. — Pubescent. Prothoracic femora and labial palpi not
exposed. Cremaster absent.

Remarks. — Agonopterix is a valid genus readily separable from
Depressaria^ with which Meyrick and others continue to synonymize
it, and from Apach£a. The stalking of veins 2 and 3 of the fore wing
in Agonopterix is constant and is associated with the absence of the
long lateral processes of the anellus so frequently encountered in
Depressaria and other oecophorid genera. The pattern of the fore
wings of Agonopterix always differs from that of Deprcssaria by en-
tirely lacking conspicuous longitudinal streaks such as are found in
the latter genus.

The moths of this genus, with the exception of one small group,
form an extremely compact assemblage of species, often difficult to
separate. The male genitalia are strikingl}'^ similar, species being
best separated on characters of the anellus, clasper, and aedeagus.
The anellus seems to be the most stable of the three and may suffice
for separation of species where the other two fail. The length of
the harpes {psoraliella and others) and the shape of the cucullus are
apt to vary considerably. (In preparing mounts of the genital
organs great care must be exercised to eliminate distortion and
shrinkage.)

The remaining species of the genus all have fingerlike claspers,
small, numerous, few, or no cornuti on the vesica, and all the females
have a closely similar type of genitalia with or without a signum.
The wing form varies from moderately narrow to broad, and the
palpus has a well-developed brush on the second segment. There
are, however, in this last large group, sections that show divergence

REVISION OF THE OECOPHORIDAE — CLARKE 45

from the usual type. One section of the group, consisting of atro-
dorsella, scdbella^ pulvipennella, ptdeae, and eupatoriiella, forms a
very closely knit complex. Another section, consisting of gelidella,
hyperella, lythrella, nuhiferella, and arcuella, forms another com-
plex of species so closely related that separation on male genitalia
is difficult. The female genitalia, however, usually give good char-
acters, but where they fail larvae or host plants suffice to distinguish
the species.

In addition there are two species, fulva and amicella, which have
become modified and represent offshoots from the type stock.

The remaining species form a group of very closely related forms.

Busck^° lists 39 species as belonging to this genus, one {plum-
merella) being described as new. The latter falls as a synonym of
eupatoriiella. In 1920 Meyrick" described dryadoxena (synonym
of costosa) and sciadopa (and proposed the new name testifica for
the supposedly preoccupied hyperella Ely) . That same year Barnes
and Busck" described pteleae^ latipalpella, hlacella^ callosella, and
terinella, of which pteleae and latipalpella are valid species ; Macella
falls as a synonym of argUlacea^ terhiella falls to paUidella^ and
callosella to sabulella. In 1921 Busck^^ described 'blackmori (syno-
nym of costosa) and Braun" described nivalis^ the latter being valid.
In 1926 Braun ^^ added cogitata ( synonym of canella) , and in 1933 '^^ I
added serrae (synonym of pallideUa) . Keifer ^^ added clarkei in 1936.
In the present paper I have transferred eight species to the new genus
Martyrhilda and have described eight species and one race as new,
bringing the total niunber of described species for our fauna to 44.
There are additional species on hand at the present time, but these
are represented by poor or unreared material, and I deem it inadvisa-
ble to add more names for these. When long or reared series can be
obtained, then, and then only, can we safely add new names. The
larvae of comparatively few of the species of this genus are known,
but those known are found attacking a large variety of plants. Many
larvae roll the leaves of the host plants and feed within the tube thus
formed; some are leaf tiers and others feed in webs in the inflores-
cence or leaves. Pupation occurs in debris on the ground or,
occasionally, in the leaf roll made by the larva.

JO Busck, Proc. V. S. Nat. Mus., vol. 35, p. 198, 1908.

^ Meyricb, Exotic Microlepidoptera, vol. 2, p. 313, 1920.

^2 Barnes and Busck, Contr. Lepid. North America, vol. 4, p. 231-233, 1920.

" Busck, Can. Ent., vol. 53, p. 277, 1921.

1* Braun, Proc. Acad. Nat. Sci. Philadelphia, vol. 73, pt. 1, p. 10, 1921.

^ Braun, Can. Ent., vol. 58, p. 47, 1926.

"Clarke, Can. Ent., vol. 65, p. 84, 1933.

" Keifer, Bull. Southern California Acad. Sci., vol. 35, p. 10, 1936.

46 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

KEY TO THE SPECIES OF AGONOPTERIX BASED ON COLORATION

1 . Fore wing roughened with numerous raised scales 2

Fore wing smooth or with few raised scales 5

2. Fore wing light ochreous-brown with black markings 3

Fore wing ochreous-fuscous thickly mottled with black and

white scales ^ 4

3. A large blackish-fuscous spot at end of cell reaching costa

pteleae Barnes and Busck (p. 72)
No such large spot latipalpella Barnes and Busck (p. 118)

4. Terminal segment of labial palpus long, recurved

eupatoriiella (Chambers) (p. 74)
Terminal segment of labial palpus short, hardly recurved

scabella (Zeller) (p. 72)

5. Fore wing with crescentic discal dash before middle 6

Fore wing otherwise 10

6. Fore wing predominantly grayish ochreous

curvilineella (Beutenmiiller) (p. 62)
Fore wing otherwise 7

7. Discal spot at end of cell yellow or cream colored 8

Discal spot at end of cell white 9

8. Fore wing grajash lavender hyperella Ely (p. 56)

Fore wing brownish purple lythrella (Walsingham) (p. 59)

9. Fore wing predominantly reddish purple arcuella, new species (p. 58)

Fore wing purplish fuscous gelidella (Busck) (p. 55)

10. Fore wing some shade of ochreous or yellow 11

Fore wing otherwise 5

11. Base of fore wing blackish fuscous atrodorsella (Clemens) (p. 70)

Base of fore wing otherwise 12

12. Inner margin of fore wing immaculate, or with minute, incon-

spicuous dark irrorations 14

Inner margin of fore wing with conspicuous dark markings 13

13. Inner margin with fuscous subtornal blotch, posticella (Walsingham) (p. 119)
Inner margin strongly suflfused basally with blackish to reddish-
fuscous.. flavicomella (Engel) (p. 105)

14. Third segment of labial palpus with at least two dark annuh 18

Third segment of labial palpus with one or no dark annulus 15

15. Outer discal spot fuscous 16

Outer discal spot cream colored or yellow 17

16. Abdomen suffused with fuscous beneath; terminal segment of

labial palpus with brownish-red subapical annulus

nubiferella (Walsingham) (p. 60)
Adbomen with black lateral line on each side beneath; terminal
segment with subapical annulus and apex blackish fuscous

sabulella (Walsingham) (p. Ill)

17. Outer discal spot yellow; terminal segment of labial palpus fus-

cous with creamy-white median fascia; abdomen with black

lateral line on each side dimorphella, new species (p. 97)

Outer discal spot cream colored; terminal segment of labial
palpus with apex and subapical annulus black; abdomen with
row of black spots on each side beneath costosa (Haworth) (p. 113)

18. Outer discal spot absent or indistinct; subbasal annulus of third

segment of labial palpus brick red robiniella (Packard) (p. 92)

REVISION OF THE OECOPHORIDAE — CLARKE 47

Outer discal spot conspicuous; subbasal annulus of third segment

of labial palpus never brick red 19

19. Outer discal spot preceded by an orange-red streak

tlielmae, new species (p. 96)
Outer discal spot not preceded by any such streak 20

20. Outer discal spot whitish or whitish ochreous 21

Outer discal spot black or blackish fuscous 24

21. At basal third of fore wing two conspicuous black discal spots

surrounded by a pale ochreous shade lecontella (Clemens) (p. 94)

Discal spots at basal third of fore wing not surrounded by a pale

ochreous shade 22

22. Ground color of fore wing reddish ochreous; outer discal spot

white pulvipennella (Clemens) (p. 76)

Ground color otherwise 23

23. Ground color of fore wing whitish ochreous; outer discal spot

white with black ring arnicella (Walsingham) (p. 90)

Ground color of fore wing pale grayish ochreous; outer discal

spot sordid whitish argillacea (Walsingham) (p. 98)

24. Fore wing ochreous overlaid with reddish fuscous; third segment

of labial palpus with subbasal and supramedial annuli and tip

black fusciterminella, new species (p. 80)

Fore wing light ochreous irrorated with blackish fuscous and
shaded with fuscous; third segment of labial palpus with sub-
basal and supramedial annuli blackish fuscous, tip whitish
ochreous pallidella (Busck) (p. 88)

25. Fore wing predominantly red, tawny-red or reddish brown 26

Fore wing otherwise 33

26. Fore wing tawny-red fulva (Walsingham) (p. 53)

Fore wing otherwise 27

27. Costa of fore wing conspicuously grayish 28

Costa of fore wing otherwise 29

28. Brush of second segment of labial palpus trumpet-shaped

antennariella, new species (p. 108)
Brush of second segment of labial palpus not trumpet-shaped

oregonensis, new species (p. 65)

29. Fore wing crimson-red irrorated with black and cinereous

scales; costa lighter, tawny-red walsinghamella (Busck) (p. 78)

Fore wing otherwise 30

30. Fore wing reddish brown 31

Fore wing ochreous-white suffused with reddish fuscous and

sparsely irrorated with black scales 32

31. Third segment of labial palpus with subbasal and subapical

annuli and apex black; abdomen with a lateral row of black

spots on each side beneath psoraliella (Walsingham) (p. 121)

Third segment of labial palpus with subbasal and subapical an-
nuli black; apex yellowish white; abdomen with four longi-
tudinal rows of fuscous spots beneath clemensella (Chambers) (p. 68)

32. Fore wing strongly suffused with reddish fuscous; discal spots

usually not sharply contrasted rosaciliella (Busck) (p. 83)

Fore wing liberally sprinkled with fuscous to black scales, but
these and discal spots usually sharply contrasted

echinopanicis, new subspecies (p. 86)

33. Fore wing some shade of gray 34

Fore wing otherwise 38

48 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

34. Brush of second segment of labial palpus trumpet-shaped; fore

wing lavender-gray nebulosa (Zeller) (p. 110)

Brush of second segment of labial palpus not trumpet-shaped;

fore wing not lavender-gray 35

35. Fore wing without row of dark subterminal spots; ground color

gray, carmine tinted; third segment of labial palpus light
grayish ochreous with black apex and black spot near base

anteriorly sanguinella (Busck) (p. 123)

Fore wing and labial palpus otherwise 36

36. Third segment of labial palpus with two conspicuous dark annul! 37

Third segment of labial palpus without conspicuous bands;

suffused with ochreous-gray outwardly muricolorella (Busck) (p. 64)

37. Fore wing pale yellowish gray with rosy tint on apical third of

wing and on basal half of inner margin canadensis (Busck )(p. 104)

Fore wing ochreous-gray without rosy tint senicionella (Busck) (p. 107)

38. Fore wing grayish fuscous 39

Fore wing otherwise 40

39. Fore wing without dark subterminal spots; third segment of

labial palpus with subbasal and supramedial annuli and tip

black cajonensis, new species (p. 82)

Fore wing with dark subterminal spots; third segment with sub-
basal and supramedial annuli black; apex sordid whitish

novi-mundi (Walsingham) (p. 87)

40. Fore wing some shade of brown or fuscous 41

Fore wing luteous, infuscated; cilia fuscous, tipped with luteous

ciarkei Keifer (p. 66)

41. Alar expanse, 16-17 mm 42

Alar expanse, 20-25 mm 43

42. Fore wing violaceous-brown; apical third of terminal segment of labial

palpus black amyrisella (Busck) (p. 124)

Fore wing ochreous-brown; terminal segment of labial palpus with
subbasal and supramedial annuli black; apical third ochreous-white

amissella (Busck) (p. 117)

43. Fore wing light brown, without row of dark subterminal spots

pergandeella (Busck) (p. 116)
Fore wing brownish or ochreous-fuscous and with row of dark subter-
minal spots 44

44. Third segment of labial palpus with basal three-fourths and apex black

nigrinotella (Busck) (p. 101)
Third segment of labial palpus with subbasal and supramedial annuli
and apex blackish fuscous costimacula, new species (p. 102)

KEY TO THE SPECIES OF AGONOPTERIX BASED ON MALE
GENITALIA

1. Clasper short, flattened, longitudinal; tegumen and socii strongly sclero-

tized (fig. 152) fulva (Walsingham) (p. 53)

Clasper not flattened and socii never appreciably sclerotized (figs. 153,

157, etc.) 2

2. Socii small and widely separated; tegumen truncated (fig. 153) 3

Socii and tegumen otherwise (fig. 62) 7

3. Hairs of the transtillar lobes coarse and strong (fig. 156) 4

Hairs of the transtillar lobes fine and weak (fig. 153) 6

REVISION OF THE OECOPHORIDAE — CLARKE 49

4. Coarse, strong hairs of the transtillar lobes three or less (fig. 156) 5

Coarse, strong hairs of transtillar lobes at least six

lythrella (Walsingham) (p. 59)
6. Clasper of harpe stout, curved (fig. 156) hyperella Ely (p. 56)

Clasper of harpe stout, straight (fig. 154) arcuella, new species (p. 58)

6. Anellus broader than long (fig. 161) nubiferella (Walsingham) (p. 60)

Anellus longer than broad (fig. 153) gelidella (Busck) (p. 55)

7. Costa and sacculus of harpe parallel; cucullus broad and rounded (fig.

186) amicella (Walsingham) (p. 90)

Costa and sacculus of harpe not parallel and cucullus not broad and
rounded (figs. 157, 158, etc.) 8

8. Clasper attaining or reaching beyond costa of harpe (figs. 174,

177, 190) 9

Clasper not attaining costa of harpe (figs. 169, 170, etc.) 11

9. Clasper reaching beyond costa of harpe (figs. 177, 190) 10

Clasper not reaching beyond costa of harpe (fig. 174)

costimacula, new species (p. 102)

10. Harpe long, slender; posterior edge of anellus strongly concave

(fig. 190) posticella (Walsingham) (p. 119)

Harpe broad; posterior edge of anellus nearly straight, with

shallow median cleft (fig. 177) psoraliella (Walsingham) (p. 121)

11. Anellus broader than long (figs. 173, 183, 184, 185) 12

Anellus longer than broad (figs. 181, 182, etc.) 16

12. Aedeagus short, stout (figs. 183a, 185a) 13

Aedaegus other\vise 14

13. Posterior margin of anellus strongly convex, smooth (fig. 183)

pergandeella (Busck) (p. 116)
Posterior margin of anellus not strongly convex or smooth (fig.

185) nebulosa (Zeller) (p. 110)

14. Clasper very stout, nearly reaching costa of harpe (fig. 184).

costosa (Haworth) (p. 113)
Clasper slender, not reaching far beyond center of harpe (figs.

159, 173) 15

15. Clasper straight, with small basal protuberance; distal end ser-

rate (fig. 173) flavicomella (Engel) (p. 105)

Clasper hooked, without basal protuberance; distal end not

serrate (fig. 157) curvilineella (Beutenmuller) (p. 62)

16. Clasper abruptly broadened at distal end (fig. 181)

sabulella (Walsingham) (p. Ill)
Clasper otherwise 17

17. Lobes of anellus large, prominent (figs. 158, 159) 18

Lobes of anellus small, inconspicuous (figs. 169, 170, 176, 179) 30

18. Posterior margin of anellus distinctly concave (figs. 158, 159,

160, 163, 164, 165, 180, 188) 19

Posterior margin of anellus not distinctly concave (figs. 162, 166,

172, 178, 182) .". 26

19. Clasper reaching well beyond middle of harpe (figs. 159, 160,

163, 164, 180, 188)_...l 20

Clasper not reaching much beyond middle of harpe (figs. 158, 165) 25

20. Aedeagus slender, sharply pointed (figs. 159a, 163a, 164a, 180a) 21

Aedeagus stout (figs. 160a, 188a) 24

50 PROCEJIJDINGS OF THE NATIONAL ]MUSEUM vol. 90

21. Anellus narrowed posteriori}^; clasper very slender (fig. 159)

clemensella (Chambers) (p. 68)
Anellus not appreciably narrowed posteriorly; clasper strong

(figs. 163, 164) ' 22

22. Anellus with a narrow, biramous, strongly sclerotized median

area pulvipennella (Clemens) (p. 76)

Anellus without such sclerotized area (figs. 163, 180) 23

23. Lateral lobes of anellus nearly attaining posterior edge of cen-

tral plate (fig. 180) cajonensis, new species (p. 82)

Lateral lobes of anellus not nearly attaining posterior edge of

central plate (fig. 163) eupatoriiella (Chambers) (p. 74)

24. Anellus strongly constricted anteriorly; harpe abruptly narrowed

beyond clasper (fig. 160) atrodorsella (Clemens) (p. 70)

Anellus not strongly constricted anteriorly and harpe not
abruptly narrowed beyond clasper (fig. 188)

latipalpella Barnes and Busck (p. 118)

25. Distal half of aedeagus much more slender than proximal half

(fig. 165a) walsinghamella (Busck) (p. 78)

Aedeagus of about equal thickness throughout its length (fig.

158a) muricolorella (Busck) (p. 64)

26. Clasper nearly attaining costa of harpe (figs. 162, 178) 27

Clasper not reaching much beyond middle of harpe (figs. 166,

172, 182) 28

27. Posterior edge of anellus convex (fig. 178) clarkei Keifer (p. 66)

Posterior edge of anellus not convex (fig. 162)

pteleae Barnes and Busck (p. 72)

28. Aedeagus long, slender (fig. 166a) rosaciliella (Busck) (p. 83)

Aedeagus stout (figs. 172a, 182a) 29

29. Anellus with a narrow, longitudinal, weakly sclerotized median

area; cucullus rather pointed (fig. 182) pallidella (Busck) (p. 88)

Anellus without such median area; cucullus rounded (fig. 172)

senicionella (Busck) (p. 107)

30. Clasper twisted (fig. 168) novi-ravxndi (Walsingham) (p. 87)

Clasper not twisted (fig. 167, etc.) 31

31. Clasper reaching at least three-fourths of the way to costa of

harpe (figs. 170, 187) 32

Clasper reaching to or slightly beyond middle of harpe (fig. 167, etc.).. 34

32. Clasper straight (figs. 170, 187) 33

Clasper definitely curved distally (fig. 176)__oregonensis, new species (p. 65)

33. Clasper stout (fig. 187) amissella (Busck) (p. 117)

Clasper slender (fig. 170) nigrinotella (Busck) (p. 101)

34. Posterior edge of anellus concave (figs. 171, 179) 35

Posterior edge of anellus not concave (figs. 167, 169, 175, 189) 36

35. Clasper slender, reaching slightly beyond middle of harpe (fig.

171) argillacea (Walsingham) (p. 98)

Clasper stout, not reaching beyond middle of harpe (fig. 178)

dimorphella, new species (p. 97)

36. Anellus nearly round (figs. 169, 189) 37

AneUus roughly rectangular (fies. 167, 175) 39

37. Posterior margin of anellus with shallow cleft; cucullus sharply

pointed (fig. 189) sanguinella (Busck) (p. 123)

Posterior margin of anellus without any trace of a median cleft;

cucullus dull-pointed (fig. 169) 38

REVISION OF THE OECOPHORIDAE — CLARKE 51

38. Cornuti fine, spiculate (fig. 169a) robiniella (Packard) (p. 92)

Cornuti coarse, small thelmae, new species (p. 96)

39. Posterior margin of anellus distinctly convex (fig. 167)

fusciterminella, new species (p. 80)
Posterior margin of anellus not distinctly convex (fig. 175)

antennariella, new species (p. 108)

KEY TO THE SPECIES OF AGONOPTERIX BASED ON FEMALE
GENITALIA

1. Bursa copulatrix with signuin (figs. 241, 255, 270, etc.) 5

Bursa copulatrix without siguum (figs. 229, 240, 272) 2

2. Ductus bursae short, not much longer than bursa copulatrix

(fig. 272) amyrisella (Busck) (p. 124)

Ductus bursae much longer than bursa copulatrix 3

3. Anterior edge of genital plate convex, entire; ostium large,

round (fig. 240) hyperella Ely (p. 56)

Anterior edge of genital plate produced (figs. 229, 259A) 4

4. Produced margin of genital plate before ostium truncate; ostium

round (fig. 229) diraorphella, new species (p. 97)

Produced margin of genital plate before ostium strongly convex;

ostium broadly oval (fig. 259a) thelmae, new species (p. 96)

5. Signum minute, round; area posterior to ostium with two small

sclerotized patches (fig. 259) robiniella (Packard) (p. 92)

Signum a small to large toothed or scobinate plate (figs. 242, 245,

250, 255, etc.) 6

6. Ductus bursae with a large sclerotized patch adjacent to bursa

copulatrix (fig. 248) fulva (Walsingham) (p. 53)

Ductus bursae without sucli sclerotized area 7

7. Anterior margin of genital plate cleft (figs. 230a, 249-253) 8

Anterior margin of genital plate not cleft 13

8. Lobe of ovipositor clothed with spines (fig. 230) clarkei Keifer (p. 66)

Lobe of ovipositor not clothed with spines (figs. 249-253) 9

9. Cleft narrow (figs. 251, 253) 10

Cleft semicircular (figs. 249, 250, 252) 11

10. Inception of ductus seminalis at the ostium (fig. 251)

pteleae Barnes and Busck (p. 72)
Inception of ductus seminalis a short distance before the ostium

(fig. 253) pulvipeiinella (Clemens) (p. 76)

11. Anterior points of genital plate nearly touching (fig. 252).

atrodorsella (Clemens) (p. 70)
Anterior points of genital plate widely separated (figs. 249, 250) 12

12. Ostium occupying over half length of genital plate (fig. 249).

eupatoriiella (Chambers) (p. 74)
Ostium occupying less than half length of genital plate (fig. 250).

scabella (Zeller) (p. 72)

13. Genital plate with pronounced anteromedian ventral evagina-

tion (fig. 274) posticella (Walsingham) (p. 119)

Genital plate without such evagination 14

14. Anterior margin of genital plate produced (figs. 255, 261, 265,

266, 269, 270, 273) 15

Anterior margin of genital plate not produced 26

15. Production of margin as wide or nearly as wide as genital plate is

long (figs. 255, 265) 16

Production of margin of genital plate narrower 17

52 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

16. Ostium large, round (fig. 265) nigrinotella (Busck) (p. 101)

Ostium spindle-shaped (transverse) (fig. 255) argillacea (Walsingham) (p. 98)

17. Ostial opening definitely in anterior half of genital plate (fig. 261).

costimacula, new species (p. 102)
Ostial opening not definitely in anterior half of genital plate

(figs. 262, 263, 264, 266, 267, 268, 269, 270, 273) 18

18. Ostial opening about middle of genital plate (figs. 266, 269) 19

Ostial opening definitely in posterior half of genital plate (figs.

262, 264, 267, 268, 270, 273) 20

19. Anterior to ostium a strongly sclerotized crescent-shaped bar;

posterior to ostium a small sclerotized rectangular area

(fig. 269) amissella (Busck) (p. 117)

Anterior to ostium an elongate, slightly curved, sclerotized mar-
gin; no sclerotized area posterior to ostium (fig. 266)

nebulosa (Zeller) (p. 110)

20. Signum distinctly 4-pointed (figs. 262, 264, 270, 273) 22

Signum not 4-pointed (figs. 267, 268) 21

21. Genital plate broad, with a gently concave, narrow, sclerotized

anterior margin; ostial opening on extreme posterior edge

(fig. 268) latipalpella Barnes and Busck (p. 118)

Genital plate narrow with a broad, convex anterior margin; ostial
opening not reaching extreme posterior edge of genital plate
(fig. 267) flavicomella (Engel) (p. 105)

22. Ostial opening occupying more than half length of genital

plate (figs. 262, 263, 270) 24

Ostial opening occupying half or less than half length of genital

plate (figs. 264, 273) 23

23. Signum large, with small, scattered teeth; lateral points much

larger than anterior and posterior points (fig. 273)

costosa (Haworth) (p. 113)
Signum small, with strong teeth; lateral, anterior, and posterior

points about equal in length (fig. 264) antennariella, new species (p. 108)

24. Posterior and anterior points of signum of about equal length

(fig. 263) senicionella (Busck) (p. 107)

Posterior and anterior points of unequal length (figs. 262, 270) 25

25. Anterior point of signum longer than posterior point (fig. 262)

canadensis (Busck) (p. 104)
Anterior point of signum shorter than posterior point (fig.
270). sabulella (Walsingham) (p. Ill)

26. Signum situated in extreme anterior end of bursa copulatrix

(figs. 242, 243, 245, 247a, 271) 27

Signum not situated in extreme anterior end of bursa copula-
trix (figs. 241, 246, 254, 256, 257, 258) 31

27. Ostial opening at extreme anterior margin of genital plate (fig.

271) psoraliella (Walsingham) (p. 121)

Ostial opening not at anterior edge of genital plate (figs. 242,

243, 245, 247) 28

28. Ostial opening at extreme posterior edge of genital plate (fig.

242) clemensella (Chambers) (p. 68)

Ostial opening not at posterior edge of genital plate (figs. 243,

245, 247) 29

REVISION OF THE OECOPHORIDAE — CLARKE 53

29. At posterior end of ostium two small, sclerotized lateral areas

(fig. 243) muricolorella (Busck) (p. 64)

No such sclerotized areas at posterior end of ostium (figs. 245,

247) 30

30. Signum a distinctly 4-pointed plate (fig. 247a) gelidella (Busck) (p. 55)

Signum irregular, not 4-pointed (fig. 245)

curvilineella (Beutenmliller) (p. 62)

31. Ostial opening at extreme posterior edge of genital plate (fig.

254) walsinghamella (Busck) (p. 78)

Ostial opening otherwise (figs. 244, 246, 256, 257, 258) 32

32. Bursa copulatrix symmetrical (figs. 244a, 256, 257, 258) 34

Bursa copulatrix distinctly asymmetrical (figs. 241, 246) 33

33. Signum oval; ostium oval (fig. 246) arcuella, new species (p. 58)

Signum diamond-shaped, ostium elliptical (fig. 241)

oregonensis, new species (p. 65)

34. Signum large (figs. 244a, 258) 35

Signum small (figs. 256, 257) 36

35. Signum a 4-pointed plate (fig. 244a) cajonensis, new species (p. 82)

Signum roughly diamond-shaped (fig. 258)

fusciterminella, new species (p. 80)

36. Signum bilobed (fig. 257) rosaciliella (Busck) (p. 83)

Signum oval (fig. 256) pallidella (Busck) (p. 88)

AGONOPTERIX FULVA (Walsingham)

Plate 25. Figuees 152, 152a ; Plate 42, Figxjee 248

Depressaria fulva Walsingham, Trans. Amer, Ent. Soc, vol. 10, p. 175, 1882. —

Riley, in Smith, List of the Lepidoptera of Boreal America, No. 5251, 1891.—

Busck, Proc. U. S. Nat. Mus., vol. 24, p. 741, 1902 ; in Dyar, U. S. Nat. Mus.

Bull. 52, No. 5871, 1903.— Keakfott, in Smith, List of the Lepidoptera of Boreal

America, No. 6414, 1903.— Meybick, in Wytsman, Genera insectorum, fasc.

180, p. 176, 1922.— Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 824,

1939.
Agonoptcryx fulva (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198,

1908.— Forbes, Cornell Univ. Agr. Exp. Stat., Memoir 68, p. 240, 1923.
Agnopteryx fulva (Walsingham) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6453, 1917.
Agonopterix fulva (Walsingham) McDunnough, Check list of the Lepidoptera

of Canada and the United States of America (Part 2, Microlepidoptera), No.

8446, 1939.
Labial palpus with second segment tawny-red outwardly irrorated
with whitish and fuscous scales, whitish inwardly ; third segment sor-
did whitish with a poorly defined narrow, light tawny-red subapical
band. Antenna pubescent, fuscous, narrowly and faintly annulated
with ochreous. Head, thorax, and ground color of fore wing tawny-
red ; thorax suffused with fuscous and irrorated with white posteriorly ;
fore wing irrorated with fuscous and with a large fuscous shade at end
of cell; in the center of this shade is a white discal spot (absent in some
specimens) ; at extreme base of wing on inner angle a small transverse
whitish patch not reaching costa ; veins, beyond cell, marked with fus-
cous scales ; cilia grayish fuscous edged lightly with rosy red. Hind

54 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

wing and cilia grayish fuscous, the latter lightly edged with rosy red.
Legs whitish inwardly, overlaid outwardly with tawny-red and with a
rosy-red suffusion; tarsi fuscous narrowly annulated with whitish.
Abdomen grayish-fuscous above, whitish beneath, strongly overlaid
with fuscous.

Male genitalia. — Strikingly different fi'om those of any other de-
scribed North American species : Harpe only moderately clothed with
hairs at the cucullus and along the costal edge ; sacculus deeply folded
and very strongly sclerotized; clasper short, flattened, longitudinal,
constricted at middle, broadly rounded at its extremity. Anellus
broadly oval, moderately sclerotized, with a deep excavation on the
posterior edge. Vinculum broadly rounded. Aedeagus sharply bent
near the middle, slender, dilated at the distal end ; armed with many
short, stout, cornuti. Tegumen strongly sclerotized, more so along
the ventral edges. Socii strongly sclerotized flaps, very sparsely
clothed with hairs. Gnathos a small, round, spined knob.

Female genitalia. — Genital plate broad, strongly sclerotized except
for a narrow, median, longitudinal membranous portion posterior to
ostium. Ostium large with deeply concave anterior edge, and open-
ing near anterior edge of genital plate. Ductus bursae membranous
except for a large sclerotized patch adjacent to bursa copulatrix and
a short portion before ostium ; anterior to the latter the ductus bursae
is constricted at the inception of the ductus seminalis. Signum a
small, irregular sclerotized plate.

Alar expanse, 22-24 mm.

Type. — In the United States National Museum.

Type locality. — Not designated.

Food plant. — Unknown.

Distribution. — Rocky Mountains region of the United States and
Canada, and eastern Canada.

United States records

Arizona: White Mountains, Apaclie County, near McNary P. O., 2 {1-15-IX-25,
O. O. Poling).

Canadian records

Alberta: "Head of Pine Creek," Calgary, $ (VIII-5-05, F. H. Wolley-Dod).
British Columbia: Jesmond, S (2-VIII-1937, J. K. Jacob).
New Brunswick: Fredericktown, $ ("August 20").
Ontario: Stittsville, $, 2 (21-VIII-1939, E. G. Lester).

I have seen other specimens without locality labels.

Remarks. — An easily recognizable species not to be confused with
any other described from North America.

The type, which is before me, shows only a single white scale in the
single discal spot at the end of the cell as described by Walsingham,
but a "homotype" from Alberta, which is clearly this species, shows a

REVISION OF THE OECOPHORIDAE — CLARKE 55

well-defined spot at the end of the cell. A third specimen, from
Beutenniiiller's collection, has only a few blackish scales suggesting
this same spot.

The female from Arizona, which I have identified as this species,
undoubtedly belongs here.

AGONOPTERIX GELIDELLA (Busck)

Plate 25, Figures 153, 153a ; Plate 42, Figxikes 247, 247a

Depressaria gelidella Busck, Proc. Ent. Soc. Washington, vol. 9, p. 90, 1908. —

Meyrick, in Wytsman, Genera insectorum, fasc. 180, p. 177, 1922. — Gaede,

in Bryk, Lepidopterorum catalogus, pt. 92, p. 325, 1939.
Agoywpteryx gelideUa Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198, 1908.
Agnopteryx gelideUa (Busck) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6450, 1917.
Agonopterix gelidella (Busck) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera ) , No.

S451, 1939.

Labial palpus yellowish white ; second segment purplish black out-
wardly tinged with carmine; third segment strongly suffused with
purplish black anteriorly, tip white. Antenna purplish black on
basal segment, remainder fuscous. Head grayish fuscous. Thorax
purplish gray; tegula purplish black tipped with whitish. Ground
color of fore wing dark purplish fuscous, sparsely irrorated with
black; near base, from inner margin, a transverse yellowish-white
streak, not reaching costa; on the disk, at basal third, a crescent-
shaped black dash with a few carmine scales mixed and followed by
a white patch ; at the end of cell a white spot edged with black ; costa
faintly spotted with whitish; from costa a poorly defined, narrow,
outwardly curved whitish line to inner margin before tornus ; around
termen a series of blackish- fuscous spots forming an almost continu-
ous fine line before cilia; inner margin and apical portion of wing
carmine tinted ; cilia grayish fuscous. Hind wing yellowish fuscous ;
cilia light fuscous with considerable white mixed. Legs yellowish
white inwardly, purplish black outwardly. Abdomen grayish fuscous
above, yellowish white beneath, suffused and irrorated with fuscous
and with a longitudinal row of black spots on each side.

Male genitalia. — Harpe only weakly sclerotized and moderately
clothed with hairs; cucullus rounded; clasper very stout, somewhat
dilated distally; outer edge rough; reaching two-thirds distance to-
ward costa. Anellus narrow, longer than broad, roughly rectangular,
emarginate on the posterior edge; lateral lobes weak. Vinculum
broad, rounded. Aedeagus moderately stout, gently curved and ter-
minating in a sharp upturned point; just before middle a forked
sclerotized arm by which it is attached to the anellus. Transtilla a
weakly sclerotized band with large, hairy lateral lobes, the hairs fine

56 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

and weak. Gnathos an oval, spined knob. Socii very small, widely
separated, sparsely clothed with fine hairs. Tegumen truncated.

Female genitalia. — Genital plate broad with a row of strong hairs
on posterior margin. Ostium large, round, nearer to anterior than
to posterior edge ; anterior edge well sclerotized. Ductus bursae long,
rather stout, constricted just before ostium at inception of ductus
seminalis. Bursa copulatrix large with a well developed 4-pointed
signum in the anterior end.

Alar expanse, 19-20 mm.

Type. — In the United States National Museum.

Type locality. — Winnipeg, Manitoba, Canada (A. W. Hanham)

Distribution. — Alberta, eastward to Ontario.

Canadian records

Alberta: Nordegg, $ (l-VIII-1921, J. McDunnough).

Manitoba: Winnipeg, 2 $ $ (no date, A. W. Hanham).

Ontario: Ottawa, $, 5 (&-VII-1905, 25-VII-1906, C. H. Young) ; Trenton, $

(27-VI-1911, Evans).
Saskatchewan: Earl Grey, $ (24-VII-1925, J. D. Ritchie) ; Indian Head, 9

(3-VIII-1925, J. J. de Gryse).

Remarks. — This and the following species are similar in pattern
to the European conterminella (Zeller) .

The abdomen of the type is missing, but I do not hesitate to place
the series before me under this name.

Through the courtesy of Dr. McDunnough I have been able to
dissect one of the only two females I have seen and have figured the
genitalia.

This species has been recorded fi'om British Columbia, but all
specimens so determined are referable to oregonensis (new species) .

AGONOPTERIX HYPERELLA Ely

Plate 25, Figxjbes 156, 156a ; Plate 42, Figube 240

Agonopterys hyperella Ely, Proc. Ent. Soc. Washington, vol. 12, p. 68, 1910.

Agonopteryx hyperella (Ely) Foebes, Cornell Univ. Agr. Exp. Stat. Memoir 68,
p. 240, 1923.

Agonopterix hyperella (Ely) McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera ) , No.
8438, 1939.

Agnopteryx hyperella (Ely) Baenes and McDunnough, Check list of the Lepi-
doptera of Boreal America, No. 6457, 1917.

Depressaria testiflca Meykick, Exotic Microlepidoptera, vol. 2, p. 316, 1920; in
Wytsman, Genera insectorum, fasc. 180, p. 176, 1922. (New synonymy.)

(Not Agonopteryx hyperella McDunnough, Can. Ent., vol. 67, p. 74, 1935.)

Labial palpus creamy white, exteriorly heavily overlaid with
fuscous; third segment with a broad black subapical annulation.
Antenna dark purplish fuscous annulated with gray. Head creamy

REVISION OF THE OECOPHORIDAE — CLARKE 57

white in front, grayish above. Thorax lavender-gray overlaid with
cinereous to white scales. Fore wing dark grayish lavender shaded
with yellowish brown, especially toward inner margin; at base of
wing, from inner angle almost to costa, a transverse yellowish line
preceded by two black spots, one near costa, the other at inner angle ;
in center of cell a black crescent-shaped dash margined with yellowish
brown and followed by a grayish shade ; between the upper point of
this dash and the costa a small black spot similarly edged with yel-
lowish brown ; at the end of cell a conspicuous, though small, cream-
colored or yellowish spot edged with brown and preceded and fol-
lowed with a few scattered black scales; costa somewhat strigulated
with fuscous and grayish; around termen, at base of cilia, a thin
fuscous to black line; cilia light purplish fuscous mixed with gray.
Hind wing smoky fuscous; cilia somewhat more brownish with a
fuscous basal band. Legs creamy white to yellowish shaded with
fuscous exteriorly except at joints. Abdomen grayish fuscous above;
beneath, yellowish overlaid with fuscous and with indistinct black
lateral lines.

Male genitalia. — Harpe slender, moderately clothed with fine hairs;
clasper very stout, gently curved inwardly, its outer edge rugose;
cucullus bluntly pointed; sacculus short, broadly folded. Anellua
deeply incised on posterior margin, concave laterally; lateral lobes
scarcely developed. Aedeagus curved, pointed, with a short ventral
arm which articulates with the anellus; vesica armed with fine
cornuti. Vinculum rounded. Transtilla a narrow sclerotized band
with well-developed lateral lobes; each lateral lobe bears three long,
stout hairs. Gnathos a spined oval knob. Socii small, widely sepa-
rated. Tegumen truncated.

Female genitalia. — Genital plate broad, moderately sclerotized;
anterior edge convex, entire. Ostium large, round; at center of
genital plate. Ductus bursae membranous; inception of ductus semi-
nalis shortly before ostium. Bursa copulatrix without signum.

Alar expanse, 15-18 mm.

Type. — In the United States National Museum.

Type localities. — Great Falls, Md., and Great Falls, Va.

Food plant. — Hypericum proli-jicum L.

Distribution. — Middle Atlantic States.

United States records

Maryland: Great Falls, 2 $ $, 9 (V-26-09, Chas. R. Ely) ; Plummors Island, $

(V-30-09, Chas, R. Ely).
Virginia: Great Falls, $ (V-31-10, Chas. R. Ely).

Remarks. — Meyrick proposed testifica in place of hyperella think-
ing that the latter name was mtended for hypencella of Hiibner and
further states that hyperella is preoccupied by hypeHcella. Ely's

58 PROCEEDINGS OF THE NATIONAL MUSEUM vol. so

species is distinct from the European hypericella, and hyperella is not
invalidated by Hiibner's name. I, therefore, reinstate Ely's species
and sink Meyrick's name.

The stout, gently recurved clasper of hyperella distinguishes it from
both lythrella and arcueUa. The clasper of lythrella is slender and
straight, and that of arcueUa is stout and straight. The separation
of lytJvrella from gelidella is discussed under lythrella. The females,
however, give the best characters for separating hyperella from
arcueUa. The signum is present in arcueUa, absent in hyperella.

AGONOPTERIX ARCUELLA. new species

Plate 25, Figure 154 ; Plate 42, Figube 246
Agonopteryx hyperella McDunnough [not Ely], Can. Ent., vol. 67, p. 74, 1935.

A small reddish-purple species closely related to the foregoing two
and to the following species.

Head yellowish ochreous, mixed with golden-brown; labial palpus
whitish ochreous ; brush of the second segment narrow, heavily over-
laid with reddish purple and black outwardly and beneath; terminal
segment thickened, reddish purple and black exteriorly, with incom-
plete reddish-purple basal and subapical annuli ; antenna with basal
segment blackish fuscous, with narrow ochreous apical annulus;
remainder of antenna fuscous, ochreous annulated, and the whole
with a purplish sheen.

Thorax strongly crested, yellowish ochreous mixed with golden-
brown ; tegula and crest yellowish purple, the former strongly shaded
with fuscous. Ground color of fore wing reddish purple strongly
suffused with fuscous along costa and at base ; the fuscous basal shad-
ing terminated by a sharply contrasting, narrow, whitish ochreous
line and including a sharply contrasted black spot near dorsal edge;
another similar black spot at base of radius; costa strigulated with
whitish and carmine ; at basal third an outwardly curved black cres-
cent preceded by carmine and followed by whitish scales ; discal spot
at end of cell white ; from middle of costa to center of wing a strong
whitish shading from costa; at apical third an indistinct fascia out-
wardly curved to between veins 6 and 7, then turned back, parallel
to termen, nearly to dorsum; cilia purplish fuscous, carmine tipped,
with black basal line from veins 3 to 7. Hind wing brownish fus-
cous; cilia a shade lighter with dark subbasal and light basal line;
underside with strong black line at base of cilia. Legs blackish fus-
cous, with whitish outwardly on femora and reddish purple on tibiae
and tarsi; tarsi annulated yellowish ochreous. Abdomen fuscous
with much yellowish ochreous beneath.

Male genitalia. — Harpe moderately sclerotized and sparsely clothed
with hairs ; cucullus rounded ; clasper stout, straight, rugose on outer

REVISION OF THE OECOPHORIDAE — CLARKE 59

edge; anellus roughly oval, emarginate on posterior edge; lateral
lobes undeveloped; aedeagus stout, pointed, slightly curved, with
large spinulate patch. Vinculum rounded. Transtilla a sclerotized
band with large lateral lobes, the latter with few strong, coarse hairs.
Gnathos an oval spined knob. Socii small, widely separated, with
few hairs. Tegumen truncated.

Female genitalia. — Ostium large, oval at about center of genital
plate; genital plate membranous or sclerotized posterior to ostium;
ductus bursae long, membranous, slightly dilated at point of inception
of ductus seminalis ; bursa copulatrix large, asymmetrical, bulging on
left side ; oval, with oval, spined signum.

Alar expanse, 16-18 mm.

Type. — In the Canadian National Collection.

Paratypes. — U. S. N. M. No. 52078. Also in Canadian National
Collection and collection of Dr. A. F. Braun.

Type locality. — White Point Beach, Queens County, Nova Scotia.

Food plant. — Hypericum virginicuTn L.

Remarks. — Described from the $ type, IQ $ $ and 10 9 5 para-
types all from Wliite Point Beach, Queens County, Nova Scotia
(VIII-7 to 13-34, J. McDunnough) ; 2 5 $ , $ , Sparrow Lake, Ontario
(VII-1-26, A. F. Braun); $, Black Mountains, North Carolina
("VII-20").

McDunnough misidentified this species and reported it under the
name A. hyperella Ely. In addition to the above I now have two
specimens from Chamcook, New Brunswick, Canada (4— 5-VIII-1938,
T. N. Freeman; Rf. Hypericum sp.) submitted by Dr. J. McDunnough,
and three specimens from Orrington, Maine (24-VII-1922, A. C.
Ward; Rf. Hypericum sp.), submitted by J. V. Schaffner.

AGONOPTERIX LYTHRELLA (Walsingham)

Plate 25, Figuke 155

Depressor ia lythreUa Walsingham, Ins. Life, vol 1, p. 257, 1889. — Busck, Proc.
U. S. Nat. Mus., vol. 24, p. 744, 1902; in Dyar, U. S. Nat. Mus. Bull. 52,
No. 5879, 1903.— Ke,\rfott, in Smitli, List of the Lepidoptera of Boreal
America, No. 6422, 1903. — Meyrick, in W.ytsman, Genera insectorum, fasc.
180, p. 176, 1922.— Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 332,
1939.

Agonopteryx lythrella (Walsingham) Busck, Proc. U. S. Nat Mus., vol. 35, p.
199, 1908.— FoEBES, Cornell Univ. Agr. Exp. Stat., Memoir 68, p. 240, 1923.

Agonopterix lythrella (Walsingham) McDunnough, Check list of the Lepidop-
tera of Canada and the United States of America (Part 2, Microlepidoptera),
No. 8437, 1939.

Agnopteryx lythrella (Walsingham) Barnes and McDunnough, Check list of
the Lepidoptera of Boreal America, No. 6468, 1917.— Bbimley, The Insects
of North Carolina, p. 304, 1938.

60 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Labial palpus yellowish mottled exteriorly with purplish fuscous;
third segment with indistinct basal and subapical fuscous annuli.
Antenna fuscous broadly annulated with tawny. Head and thorax
cinereous, the former with a fuscous median shade, the latter irrorated
with purplish fuscous ; tegula purplish fuscous. Fore wing brownish
purple dusted with fuscous and with much cinereous scaling along
costa; at base, from inner angle almost to costa a narrow, cream-
colored, transverse line preceded at inner angle by a distinct black
spot; on disk, before middle, a short outwardly curved, black
crescent-shaped dash edged with reddish and followed by cinereous;
at end of cell a small cream-colored spot edged with fuscous; from
costa to termen at apical third a more or less distinct cinereous fascia ;
inner margin narrowly shaded with reddish ; around termen, at base
of cilia, a narrow blackish-fuscous line; cilia purplish gray. Hind
wing and cilia brownish gray, the latter edged with cinereous. Legs
cream-colored strongly overlaid and irrorated exteriorly with pur-
plish fuscous except at joints. Abdomen purplish gray above;
beneath, cream-colored strongly suffused and overlaid with purplish
fuscous.

Alar expanse, 13-18 mm.

Tyfe. — In the British Museum.

Type locality. — "Illinois."

Food plant. — Lythrum alatum Pursh.

Distribution. — Eastern United States.

United States records
Illinois.
North Carolina: Black Mountains (July 20, from larvae).

Remarks. — The male genitalia of this species are nearly identical
with those of hyperella, arcuella, nuhiferella, and gelidella. The
clasper of the harpe (fig. 155), however, is much more slender than
that of any of the other four. Each of the hairy lobes of the
transtilla bears about half a dozen strong hairs. In hyperella and
arcuellu there are only two or three hairs and in gelidella the hairs
are numerous but weak. I have not seen the female genitalia of this
species. All females examined are without abdomens.

AGONOPTERIX NUBIFERELLA (Walsingham)

Plate 26, Figuees 161, 161a

Depressaria nuhiferella Walsingham, Proc. Zool. See. Lonclou, ISSl, p. 316,
pi. 36, fig. 6.— BuscK, Proc. U. S. Nat. Mus., vol. 24, p. 745, 1C02 ; iyi Dyar,
U. S. Nat. Mus. Bull. 52, No. 5S81, 1903.— Kearfott, in Smith, List of the
Lepidoptera of Boreal America, No 6424, 1903. — Busck, Proc. U. S. Nat. Mus.,
vol. 27, p. 764, 1904. — MeyeicKs in Wytsman, Genera insectorum, fasc. 180,
p. 176, 1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 335, 1939.

Agonopteryx nuMferella (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35,
p. 199, 1908.

REVISION OF THE OECOPHORIDAE — CLARKE 61

Affonopterix nuUfereUa (Walsingham) McDunnough, Check list of the Lepi-
doptera of Canada and the United States of America (Part 2, Microlepidop-
tera), No. 8450, 1939.

Agnoptcryx nuUfercUa (Walsingham) Barnes and McDunnough, Check list of
the Lepidoptera of Boreal America, No. 6470, 1917.

Labial palpus light ochreous, second segment overlaid with brownish
red exteriorly ; third segment with subapical annulus of the same color.
Antenna fuscous with slight reddish luster. Head, thorax, and ground
color of fore wing ochreous ; tegula and anterior part of thorax over-
laid with brownish red. Extreme base, except costa, of fore wing
unmarked, remainder of wing suffused with brownish red and irrorated
with scattered fuscous scales ; from costa, across end of cell, almost to
inner margin, a dark brownish-red shade with a fuscous central discal
dot ; this shade is preceded by two more or less distinct brownish-red
discal dots, obliquely one above the other ; along costa a series of red-
dish-fuscous spots; around termen, at base of cilia, a line of the same
color ; cilia ochreous, shaded with brownish red. Hind wing grayish
fuscous ; cilia yellowish basally, grayish fuscous apically. Legs light
ochreous; anterior and median pair strongly overlaid with reddish
fuscous except at joints; posterior legs irrorated and suffused with
light fuscous except at joints. Abdomen ochreous overlaid with
fuscous beneath.

Male genitalia. — Harpe clothed with rather coarse hairs; cucullus
rounded; clasper short, reaching just beyond middle, but very stout,
bluntly pointed. Anellus a lightly sclerotized plate, broader than long,
indented at the posterior edge ; lateral lobes undeveloped. Vinculum
rounded. Aedeagus stout, dorsoventrally depressed, with a slightly
upturned point; near the base is a bifid plate by which the aedeagus
articulates with the anellus. Transtilla a narrow band with large,
hairy lateral lobes, the hairs fine and weak. Gnathos an oval spined
knob. Socii very small, widely separated, and sparsely clothed with
fine hairs. Tegmnen truncated.

Alar expanse, 18-20 mm.

Type. — In the British Museum.

Type locality. — Eogue River, Oreg.

Food plant. — Hypericum perforatum L. (The Hypericum referred
to by Walsingham is probably this species also.)

Distribution. — Western United States and Canada.

United States records

California: Mendocino and Shasta Counties (June and July 1871, Walsingham).
Idaho: Lapwai (V-28 to l-VI-35, J. F. G. Clarke [reared]).
Oregon: Rogue River.

Washington: Logan Hill, Chehalis (IX-8-29, T. M. Clarke) ; Pullman
(VH-24-9S and VIII-10-9S, C. V. Piper).

62 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

Canadian records
British Columbia: Shawnigan Lake, Vancouver Island (VII-14-23, E. H.
Blackmore) .

Remarks. — This and the four foregoing species all have very sim-
ilar genitalia, but nubiferella has distinctly narrower harpes than the
others and is widely different in pattern and coloration.

All the specimens I have seen are males. The California and Oregon
specimens (including cotypes) are all considerably lighter in colora-
tion than specimens from Washington, Idaho, and British Columbia.
The northern specimens may represent a species distinct from that rep-
resented by the California and Oregon specimens, presenting a par-
allel case with that of arcueJla and hyperella in which the females
give the only reliable character for separation. The type locality
(from which I have two specimens) is not far from the locality where
the Idaho material was collected, and it does not seem likely, there-
fore, that the two color forms represent different species. Until fe-
males from both northern and southern localities are obtained I
believe it advisable to leave both color forms under one name.

AGONOPTERIX CURVILINEELLA (Bentenmuller)

Plate 26, Figures 157, 157a ; Plate 42, Figueb 245

Depressaria curvilineella BEXJTENMtiLLEit, Ent. Amer., vol. 5, p. 10, 1889. — Busck,
in Dyar, U. S. Nat. Mus. Bull. 52, No. 5874, 1903.— Kearfott, in Smith, List
of the Lepidoptera of Boreal America, No. 6417, 1903. — Meyeick, in Wyts-
man. Genera insectorum, fasc. 180, p. 176, 1922. — Gaede, in Bryk, Lepidop-
terorum catalogus, pt. 92, p. 321, 1939.

Depressaria curvilinieUa Rilet, in Smith, List of the Lepidoptera of Boreal
America, No. 5260, 1891. — Dietz, in Smith, Catalogue of the insects of New
Jersey, p. 474, 1900.— Btjsck, Proc. U. S. Nat. Mus., vol. 24, p. 742, 1002.

Agonopteryx curvilinieUa Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198, 1908.

Agonopteryx curviUneella (Beutenmiiller) Forres, Cornell Univ. Agr. Exp. Stat.
Memoir 68, p. 240, 1923.

Agonopterix curvilineella (Beutenmiiller) McDunnough, Check list of the Lep-
idoptera of Canada and the United States of America (Part 2, Microlepidop-
tera). No. 8439, 1939.

Agnopteryx curvilinieUa Ke^^kfott, in Smith, Catalogue of the Insects of New-
Jersey, p. 561, 1910.

Agnopteryx curvilineella (Beutenmiiller) Barnes and McDunnough, Check list
of the Lepidoptera of Boreal America, No. 6445, 1917.

Labial palpus white ; second segment irrorated with blackish fuscous
exteriorly; third segment with basal and subapical bands and apex
blackish fuscous. Head, thorax, and ground color of fore wing light
grayish ochreous; face white; base of tegula and fore part of thorax
suffused with brown; fore wing, except extreme base and basal half
of costa, strongly suffused with brown and irrorated with scattered

REVISION OF THE OECOPHORIDAE — CLARKE 63

in pale basal patch two small blackish-fuscous
spots; along costa and around termen a series of blackish-fuscous spots,
those around termen small and indistinct ; at end of cell a white discal
spot edged with blackish fuscous and preceded by a long, curved,
blackish-fuscous discal dash ; cilia light grayish ochreous, suffused with
brown. Hind wing gi-ayish fuscous; cilia white with grayish-fuscous
basal band. Legs white suffused with fuscous ; tarsi of hind legs with a
fuscous spot at the base of each. Abdomen grayish ochreous above
suffused with fuscous ; beneath, whitish sparsely irrorated with fuscous
scales and with a row of blackish-fuscous spots on each side ; between
these rows of spots two rows of smaller spots of the same color.

Male genitalia. — Harpe broad, profusely clothed with fine hairs, par-
ticularly basally; cucullus pointed; sacculus moderately sclerotized,
broad; clasper slender, heavily sclerotized, hooked, reaching two-thirds
distance toward costa and without basal protuberance ; anellus moder-
ately sclerotized, oval, broader than long, with weak lateral hairy
lobes. Vinculum with a well-developed dorsoanterior process.
Aedeagus nearly straight, wedge-shaped, bluntly pointed. Transtilla
a narrow sclerotized band, with sparsely hairy, lateral lobes. Gnathos
a finely spined, elongate-oval knob. Socii weakly sclerotized, broad,
hairy flaps.

Female genitalia. — Genital plate moderately sclerotized. Ostium
near middle of genital plate. Ductus bursae long, slender, mem-
branous. Bursa copulatrix oval; signum a small, irregular, sclero-
tized plate, armed with short thornlike teeth.

Alar expanse, 15-20 mm.

Type. — In the United States National Museum.

Type locality. — "New York."

Distribution. — Eastern United States and Canada.

United States records
District of Columbia: Chain Bridge, 2 9 5 (October 20, 1920, A. Busck).
Maryland: Cabin John (11-26-11, F. Knab) ; Plummers Island, 29 5 5, 14 9 9

(March and April dates ; A. Busck and H. S. Barber).
New Jersey: Essex County Park, 9 ("June 10," W. D. Kearfott).
New York: 2 9 9 (Beutenmiiller) ; Ithaca, 2 9 9 (6-15-V-30, A. B. Klots) ;

Rochester, 2 9 9 (8-VII-33, A. B. Klots).
Pennsylvania: New Brighton, 10<J 5, 10 9 9 (May and August to November

dates, H. D. Merrick) ; Oak Station, Allegheny County, 3 $ S, 9 (July and

September dates; Fred Marloff).

Canadian records

Manitoba: Aweme (2S-IX-23, N. Criddle) ; Winnipeg, $ (A. W. Hanham.

no date) ; 9 ("1-5-98," no collector).
Ontario: Ottawa (28-VIl-2-VIII-190o, C. H. Young).

64 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

AGONOPTERIX MURICOLORELLA (Busck)

Plate 26, Figitbes 158, 15Sa ; Plate 42, Figuee 243

Depressaria muricolorella Busck, Proc. U. S. Nat. Mus., vol. 24, p. 741, 1902;
in Dyar, U. S. Nat. Mus. Bull. 52, No. 5873, 1903.— Kearfott, in Smith,
List of the Lepidoptera of Boreal America, No. 6416, 1903.— Meyrick, in
Wytsman, Genera insectorum, fasc. 180, p. 176, 1922,— Gaisde, in Bryk,
Lepidopterorum catalogus, pt. 92, p. 333, 1939.

Agonopteryx muricolorella Busck, Proc. U. S. Nat. Mus., vol. 35, p. 199, 1908.

Agonopterix muricolorella (Busck) McDunnough, Check list of the Lepi-
doptera of Canada and tlie United States of America (Part 2, Microlepidop-
tera). No. 8441, 1939.

Agnopteryx muricolorella (Busck) Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6455, 1917.

Labial palpus creamy white; outside of second and all of third
segment suffused with ochreous-gray ; second segment sparsely ir-
rorated with fuscous. Antenna dark ochreous-gray. Head, thorax,
base and basal third of costa of fore wing light grayish brown ; fore
wing dark mouse gray with sparse blackish-fuscous irrorations and
with a purple tint around edges and at apex; at end of cell a
white discal spot edged outwardly with a semicircle of blackish-
fuscous scales ; first discal spot indicated by a few scattered blackish-
fuscous scales at basal third; on costa and around termen a series
of indistinct blackish-fuscous spots; cilia mouse gray with some
mixture of pale whitish ochreous. Hind wing grayish fuscous;
cilia a shade lighter with light fuscous band and whitish tips.
Legs light grayish ochreous suffused with fuscous. Abdomen dark
purplish gray above; beneath grayish ochreous shaded with fuscous.

Male genitalia. — Harpe moderately clothed with fine hairs ; cucul-
lus pointed ; clasper rather stout, lightly sclerotized, constricted just
before middle and having a twisted appearance; apex heavily scler-
otized, rough. Anellus oval, longer than broad, with large hairy
lateral lobes; posterior edge concave. Vinculum rounded with a
well developed dorso-anterior process. Aedeagus stout, cur\xd, and
terminating in a sharp, slightly upturned point. Transtilla a scler-
otized band with well developed hairy lateral lobes. Gnathos oval,
clothed with spines. Socii broad, hairy lobes. Tegumen terminating
in a single short median projection.

Female genitalia. — Genital plate moderately broad; deeply cleft
on posterior edge. Ostium small; near anterior margin of genital
plate ; at posterior end of ostium two small, sclerotized lateral areas.
Ductus bursae long, gradually enlarged to form the small oval
bursa copulatrix. Signum an oblong, sclerotized plate with pro-
jections on the anterior and posterior edges.

Alar expanse, 17-18 nun.

Type. — In the United States National Museum.

REVISION OF THE OECOPHORIDAE — CLARKE 65

Type locality. — Golden, Colo.

Food 'plants. — "An umbelliferous plant" (Dyar) ; Lomat'mm grayi
Coult. and Rose (Clarke).

Distribution. — Western United States.

United States records

Colorado: Golden, $ (Dyar and Caudell, no date).

Washing'ton: Snake River, Whitman County, opposite Clarkston, 3 5 5,39 9
(V-3()-31, VI-4 to 18-33, and V-11-34, J. P. G. Clarke).

Remarks. — With Busck's unique type male I associate six reared
specimens from the Snake River, AVash. (Clarke Nos. 2944, 4416, 4417,
4418, 4419, and 5209) . The Washington specimens fit the description
in every detail except that some of them show two discal spots, the
single discal spot not being present. In specimen 5209 veins 3 and
4 of the hind wing are short stalked as described, but in the remaining
specimens they are connate. In spite of tlie variation in venation it
would be illogical to consider the Washington specimens anytl.dng
but one species. The male genitalia show some slight variation in
the length of the harpe and shape of the cucullus, but this is not
significant.

AGONOPTERIX OREGONENSIS. new species
PrAiTc 31, Figures 176, 17Ga ; Plate 42, Figuee 241

A small reddish species with contrastingly lighter costa.

Antenna with basal segment blackish fuscous; remainder shining
grayish fuscous, darker distally; narrowly annulated with blackish
fuscous; the whole with a faint reddish sheen. Labial palpus with
second segment light whitish ochreous, the brush pink beneath both
inwardly and outwardly ; outwardly, and at apex inwardly, irrorated
with fuscous; third segment blackish fuscous with a pink-tinged
whitish-ochreous median fascia. Head light brownish fuscous with
a pink suffusion; scales tipped with whitish ochreous; face shining
whitish ochreous. Thorax, tegula, and fore wing brownish fuscous,
the fore wing so heavily overlaid with brick-red scales that it appears
dark reddish. Thorax and tegula with a strong reddish suffusion
anteriorly and much whitish-ochreous scaling posteriorly. Base of
fore wing and costa to middle a contrasting light gray; beyond middle
of wing the light-gray scaling diminishes; extreme costal edge bright
pink spotted with black; along costa, inside the grayish area, and
along veins toward the apex, with abundant black scaling. At the
end of cell a conspicuous white spot encircled with black scales; at
basal third two small, obliquely placed black spots followed by white
scaling; cilia fuscous with pink suffusion. Hind wing dark smoky
fuscous; cilia fuscous; pink tinged apically. Legs whitish. ochreous

286614—41 3

66 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

much suffused and irrorated with fuscous ; tarsi fuscous above except
for narrow band of whitish-ochreous at distal ends of segments.
Abdomen grayish fuscous above, whitish ochreous below; posterior
edges of segments above whitish ochreous ; on underside a lateral row
of black spots ; anal tuft tinged with pink.

Male genitalia. — Harpe clothed with many long hairs; cucullus
pointed; costa strongly sclerotized; clasper stout, pointed, reaching
three-fourths of way to costa; definitely curved distally; sacculus
moderately sclerotized. Anellus an elongate, moderately sclerotized
plate, lateral edges concave; anterior edge convex, posterior edge
deeply cleft, lateral lobes small. Vinculum rounded. Aedeagus
slender, gently curved, pointed. Transtilla a narrow, strongly scle-
rotized band, lateral lobes large, hairy. Socii large, fleshy, hairy
lobes. Tegumen pointed.

Female genitalia. — Genital plate broad. Ostium large, broadly
elliptical. Lobes of ovipositor clothed with long coarse hairs. Duc-
tus bursae long, membranous; inception of ductus seminalis just
before ostium. Bursa copulatrix large with well developed diamond-
shaped signum.

Alar expanse, 15-18 mm.

T'y^e.—U.S.N.M. No. 62079.

Type locality. — Salem, Oreg. (March; E. Y. Lansing, Jr.)

Described from the $ type and 5 5 5 and 10 ? 9 paratypes as
follows: 5 , Pullman, Wash. (8-II-30, J. F. G. Clarke) ; 2 5 5 , Dun-
can, British Columbia (26-11-21, W. Downes) ; 5 , Duncan, British
Columbia (25-III-21, A. W. Hanham) ; 5 , Fitzgerald, British
Columbia (14-V-22, E. H. Blackmore) ; 5, Salem, Oreg. (March;
E. Y. Lansing) ; 2 , Logan Hill, Chehalis, Wash. (15-X-29, T. M.
Clarke) ; 9 , Departure Bay, British Columbia (^11-29) : 5 ,
Fitzgerald, British Columbia (14-V-22, E. H. Blackmore) ; 9 , Era-
ser Mills, British Columbia (22-XI-23, L. E. Marmont) ; 9 , Quam-
ichan Lake, British Columbia (l-IV-23, E. H. Blackmore) ; 2 9 9 ,
Victoria, British Columbia (21-III-21, E. H. Blackmore; 23-VI-23,
W. R. Carter) ; 2 9 9 , no locality but British Columbia, specimens
under Blackmore No. 160 (13-IV-23; 5-V-23).

Remarks. — This species has been placed under the name gelidella
in collections.

AGONOPTERIX CLARKE! Keifer

Plate 31, Figubes 178, 178a ; Plate 40, Figures 230, 230a

Agonopteryx clarkei Keifer, Bull. Southern California Acad. Sci., vol. 35, p. 10,

pi. 4; pi. 7, fig. 6, 1936.
Agonoptcrix clarkei (Keifer) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera), No.

8415, 1939.

REVISION OF THE OECOPHORIDAE — CLARKE 67

Dcpressaria clarkei (Keifer) Gaede, in Bryk, Lepidopterorum catalogus, pt. 92,
p. 315, 1939.

Labial palpus liiteous ; second segment irrorated with fuscous out-
wardly and in brush; third segment somewhat darker than second
with poorly defined blackish-fuscous basal and subterminal annuli.
Antenna fuscous with narrow, darker annulations, the latter obscured
below. Head, thorax, base and cost a of fore wing luteous, the head
infuscatecl above; thorax infuscated particularly anteriorly; costa
with fuscous spots and dashes throughout its length; beyond luteous
basal patch deeply infuscated, this color rapidly fading; at middle a
dark blotch from costa; first and second discal spots minute, black,
obliquely one above the other; third discal spot at end of cell whitish
followed by a black scale or two ; all discal spots practically obsolete ;
cilia fuscous tipped with luteous. Hind wing luteous, darker api-
cally; cilia with a fuscous basal band. Legs luteous overlaid and
suffused except at joints ; posterior legs somewhat lighter. Abdomen
luteous, darker above than below; on each side, beneath, a blackish-
fuscous line.

Male genitalia. — Harpe heavily clothed with long hairs; cucullus
bluntly pointed, sacculus moderately sclerotized, narrow; clasper
long, slender, straight, nearly reaching costa. Anellus a simple
sclerotized plate with deeply excavated lateral, and convex posterior
edges; longer than broad; hairy lateral lobes well developed. Vin-
culum evenly rounded. Aedeagus short, stout, strongly bent; vesica
armed with many small spinules. Transtilla a narrow sclerotized
band with well-developed hairy lateral lobes. Socii large, hairy,
fleshy flaps.

Female genitalia. — Genital plate broad, moderately sclerotized,
anterior margin cleft. Ostium small, situated near posterior edge
of plate. Ductus bursae membranous, moderately long ; inception of
ductus seminalis just before ostium. Bursa copulatrix large, with a
large, strongly sclerotized, toothed signum. Lobe of ovipositor with
a few long, strong hairs and numerous moderately long stout spines.

Alar expanse, 18-20 mm.

Type.—V.S.l^M. No. 52092.

Type local'itij. — Missouri Flat, Placerville District, Calif.

Food plant. — Artemina vulgaris L. var.

Remarks. — Keifer has discussed and figured this species in his
paper, but I have refigured it for completeness' sake.

I have before me the specimens from Aweme, Manitoba, which
Keifer discusses, but I do not believe these represent a different
species. Although they could possibly be considered as a race, they
differ from the typical specimens only in the slightly paler pink
coloration in the cilia of the fore wing and in the apparently slightly

68 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

shorter spines of the lobes of the ovipositor. Until we know more
about the specimens from Aweme I decline to give them a separate
designation.

The type of ovipositor found in this species is unique for the genus.

AGONOPTERIX CLEMENSELLA (Chambers)
PLATE! 26, FiGUKES 159, 150a ; Plate 42, Figure 242

Qelechia clemensella Chambers, Can. Ent., vol. 8, p. 173, 1876.

Dcprcssaria clemensella (Chambers) Bxjsck, Proc. U. S. Nat. Mns., vol. 27, p. 766,
1904.— Meyeick, in Wytsman, Genera insectorum, fasc. 180, p. 176, 1922. —
Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 315, 1939.

Dcpressaria applana (Fabrlcius) Walsingham, Trans. Amer. Ent. Soc, vol. 10,
p. 175, 1882. — RiLET, in Smith, List of the Lepidoptera of Boreal America,
No. 5251, 1891.— BuscK, in Dyar, U. S. Nat. Mus. Bull. 52, No. 5862, 1903.—
Kearfott, in Smith, List of the Lepidoptera of Boreal America, No. 6405,
1903.

Affonopteryx clemensella (Chambers) Busck, Proc. U. S. Nat. Mas., vol. 35, p.
198, 1908.

Agonopterix clemensella (Chambers) McDunnough, Check list of the Lepidop-
tera of Canada and the United States of America (Part 2, Microlepidoptera ) ,
No. 8430, 1939.

Agnoptcryx clemensella (Chambers) Babnes and McDuxnough, Check list of
the Lepidoptera of Boreal America, No. 6447, 1917.

Agonopteryx applana clemensella (Chambers) Forbes, Cornell Univ. Agr. Exp.
Stat. Memoir 68, p. 239, 1923.

Labial palpus yellowish white; second segment irrorated with fus-
cous exteriorly and with a strong pink suffusion in the brush ; third
segment with basal and broad subapical tumuli black. Antenna fus-
cous, somewhat lighter beneath. Head, thorax, and base of fore wing
very light yellowish gray with fuscous and reddish-brown irrorations
and a rosy suffusion. Ground color of fore wing reddish brown ; costa
and termen blotched with alternate fuscous and yellowish- white spots;
whole surface of wing marked with scattered yellowish-white scales;
discal spots four, white; one at end of cell preceded by one at middle
of wing; at basal third two, one obliquely above the other; the outer
surrounded by a dull fuscous cloud, the inner, oblique pair preceded
by a few black scales ; cilia reddish brown suffused with fuscous and
irrorated with yellowish white. Hind wing and cilia shining yellow-
ish fuscous, the latter with a fuscous basal band. Legs yellowish
white overlaid and irrorated with fuscous exteriorly; posterior pair
lighter in color, all with a rosy tint. Abdomen grayish fuscous above,
yellowish white beneath with four longitudinal rov.s of fuscous spots.

Male genitalia. — Harpe slender, heavily clothed with hairs; sacculus
moderately sclerotized ; clasper slender, nearly reaching costa of harpe,
slightly thicker at middle, strongly curved toward the cucullus. Anel-
lus broadly rectangular, slightly constricted posteriorly; posterior

REVISION OF THE OECOPHORIDAE CLARKE 69

margin concave and with well developed lateral lobes. Vinculum
broad with a pointed dorso-anterior process. Aedeagus long, curved,
terminating in a sharp slender point; basally there is a sclerotized
biramous arm by which tlie aedeagus articulates with the anellus.
Transtilla a narrow sclerotized band, with large, profusely hairy,
lateral lobes. Gnathos long, oval, slender, clothed with fine spines.
Socii broad fleshy lobes.

Female genitalia. — Genital plate broad, lightly sclerotized; ostium
small oval, at posterior edge of genital plate. Ductus bursae mem-
branous, inception of ductus seminalis rather remote from ostium;
bursa copulatrix oval. Signum a small, more or less rectangular
plate, slightly cupped and covered with thornlike teeth.

Alar expanse, 16-20 mm.

Type. — In the Museum of Comparative Zoology, Cambridge, Mass.

Type locality. — Easton, Pa.

Food plant. — Parsnip; wild parsnip?.

Dis-trihution. — Northeastern United States and eastern Canada.

United States records

Connecticut: $ (Beutenmiiller).

District of Columbia: Chain Bridge, 5 ("2-4-85") ; Washington, 2 ("21-4-85").

Illinois: Putnam County, S (M. O. Glenn).

Maryland: Plummers Ishmd, S (March 1908, H. S. Barber).

New Hampshire: Hampton, S (IV-2G-04, S. A. Shaw).

New York: Ilion, $ (IX-3-11, McBlhose) ; Ithaca, 2 9 9 (5-19-V-31, A. B.
Klots) ; Lockport, 9 (XI-30-35, L. L. Pechumau) ; Long Island: Orient, $
(VII-27-26; Ray Latham) ; Montvale, 9 (November 26, 1923, F. M. Schott) ;
Rochester, 9 (3-IV-33, A. B. Klots).

Pennsylvania: New Brighton, 12 $3,7 9 9 (March and July to November
dates, H. D. Merrick) ; Oak Station, Allegheny County, 2 9 9 (Oct. 15,
1908, and March 30, 1910, Fred Marloit) ; Pittsburgh, $ (IV-7-06, Henry
Engel).

Vermont: Clarendon, $ (no date or collector).

Wisconsin: Millwood County, $ ("X-14-08").

Canadian records
Ontario: Ottawa (VIII-21-05, C. H. Young) ; Toronto ("III-2 to 8-95").

Remarks. — Busck ^® pointed out that in all probability cle^rnemella
is distinct from the European applana. There can be no doubt about
the close relationship between the two species, but they can be separated
easily on genitalic characters as well as pattern.

The ground color of applana is much lighter than that of clemrnsella;
in the former there is a tendency to coalescence of the discal spots,
whereas in the latter the spots are smaller and distinctly separated.

The male genitalia of clemensella show a slender, gradually finely

« Eusck, Proc. U. S. Nat. Mus., vol. 27, p. 766, 1904.

70 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

pointed aedeagus and a very slender clasper as compared to a more
abruptly pointed aedeagus and a stouter clasper in applana. The
female genitalia show fewer differences, but those examined indi-
cate that the ductus bursae of applana is longer in proportion to the
other structures than that of clemensella. I have examined 16 Euro-
pean and 41 American specimens, all of which bear out the above
conclusion.

In view of this evidence, I believe it is certain that the European
applana does not occur in North America.

AGONOPTERIX ATRODORSELLA (Clemens)

Plate 26, Figuees 160, 160a ; Plate 43, Figxjbe 252

Depressaria atrodor sella Clemens, Proc. Ent. Soc. Philadelphia, vol. 2, p. 124,
1863. — Packard, Guide to the study of insects, p. 349, 1869. — Robinson, Ann.
Lye. Nat. Hist. New York, vol. 9, p. 156, pi. 1, fig. 7, 1870.— Clemens, in Stain-
tou, Tineina of North America, p. 91, 1872. — Chambers, Can. Eut., vol. 4, p.
91, 1872.— Zeller, Verh. zool.-bot. Ges. Wien, vol. 23, p. 233, 1873.— Chambebs^
U. S. Geol. Geogr. Surv. Terr. Bull. 4, p. 138, 1878.— W.\lsingham, Proc. Zool.
Soc. London, 1881, p. 312.— Coquillett, Papilio, vol. 3, p. 98, 1883.- Beuten-
MULLER, m Smith, Catalogue of the insects of New Jersey, p. 355, 1890.—^
Riley, in Smith, List of the Lepidoptera of Boreal America, No. 5255, 1891. —
DiETz, in Smith, Catalogue of the insects of New Jersey, p. 473, 1900. — BuscK^
Proc. U. S. Nat. Miis., vol. 24, p. 736, 1902 ; in Dyar, U. S. Nat. Mus. Bull. 52,
No. 5854, 1903. — Kearfott, in Smith, List of the Lepidoptera of Boreal Amer-
ica, No. 6397, 1903. — Meyrick, in Wytsman, Genera insectorum, fasc. 180, p.
173, 1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 310, 1939.

Agonoptenjx atrodor sella (Clemens) Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198,
1908.— Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 238, 1923.

Agonopterix atrodorsella (Clemens) McDunnough, Check list of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera),
No. 8409, 1939.

Agnopteryx atrodorsella (Clemens) Smith, Catalogue of the insects of New
Jersey, p. 561, 1910. — Barnes and McDunnough, Check list of the Lepidop-
tera of Boreal America, No. 6433, 1917.

Labial palpus and face yellowish white ; second segment of palpus
irrorated with fuscous exteriorly and in the brush, the latter also
tinted with pink; third segment with basal and subapical annuli
and tip blackish fuscous. Antenna with black, ochreous-tipped basal
segment; remainder fuscous annulated with ochreous-fuscous. Head
reddish ochreous with median fuscous area. Thorax and extreme
base of fore wing blackish fuscous; collar whitish. Fore wing yel-
lowish ochreous with sparse blackish irrorations; costa with four to
six or seven blackish-fuscous spots; at basal third a black discal dot
followed by a white one at the end of cell ; from center of cell a longi-
tudinal reddish blotch extending to apical third and enclosing the
white discal spot; on costa, from apical third to apex, a longitudinal
fuscous and ochreous cloud ; cilia reddish ochreous. Hind wing light

REVISION OF THE OECOPHORIDAE — CLARKE 71

reddish fuscous; cilia whitish with basal and two subapical bands
fuscous. Legs yellowish white strongly overlaid with blackish fus-
cous except at joints; the latter suffused with pink. Abdomen light
brownish ochreous, lighter on posterior margins of segments; beneath,
yellowish white with broad, blackish-fuscous longitudinal, lateral
stripes and two rows of similarly colored spots between.

Male genitalia. — Harpe broad basally but abruptly narrowed be-
yond clasper; moderately clothed with hairs; clasper long, slender,
slightly bent, not quite reaching costa. Anellus a sclerotized plate,
longer than broad, concave on posterior edge, constricted anteriorly,
with prominent hairy lateral lobes. Aedeagus stout, slightly curved,
pointed, with large spinulate patch in basal two-thirds of vesica.
Transtilla a narrow sclerotized band with well developed, hairy,
lateral lobes. Gnathos an elongate, oval, spined knob. Socii large
fleshy, hairy flaps.

Female genitalia. — Ostium large, near posterior edge of genital
plate. Genital plate very broad, produced anteriorly; anterior
edge deeply cleft, the cleft semicircular, the anterior points nearly
touching. Ductus bursae long, membranous, gradually tapering to
form the large oval bursa copulatrix; inception of ductus seminalis
just anterior to ostium. Signum a large diamond-shaped, spined
plate.

Alar expanse, 18-24 mm.

Type. — In the Academy of Natural Sciences of Philadelphia.

Type locality. — Pennsylvania,

Food plants. — Bidens frondosa L., Myrica asplenifolia L., Eupa-
torium sp.

Distribution. — Eastern United States and Canada.

United States records

Connecticut: $ ^ Beutenmiiller, coll.).

District of Columbia: W^ashington, S (X-25-02).

Illinois: Flosmoor, S, $ (no date or collector) ; Evanston, $ ("4-99").

Massachusetts: Cummington, $ (no date or collector) ; Dover, $ (8-7-33, no

collector) ; Marthas Vineyard, $ ("1-3," George D. Eustis) ; Winchendon, $

(IX-2S-02, no collector).
New Hampshire: Hampton, 3 $ $ (V-10-13, S. A. Shaw).
New Jersey: Cedar Grove, $ (IV-9-24, F. M. Schott).
New York: Ilion, $, ? (V-3-13, H. McElhose) ; Ithaca, ? (X-24-35, J. G.

Franclemont) ;2 3 $ (12-V-31, 5-V-31, A. B. Klots) ; Liberty, 9 (no date or

collector) ; New Windsor, 2 $ $ (13-VIII-'93; 23-VIII-'97, E. L. Morton).
Pennsylvania: New Brighton, 4$$ (IX-7-02, IX-27-02, V-25-06, IX-29-06,

H. D. Merrick).
Vermont: Clarendon, S (W. D. Kearfott).
"Wisconsin: Cranmoor, Wood County, ? (IV-22-0S, C. B. Hardenberg) ;

Madison, $ (XI-10-23, S. B. Fracker).

72 PROCEEDINGS OF TPIE NATIONAL MUSEUM vol. 90

Canadian records

Ontario: Bobcaygeon (July 24, 1931, J. McDunnough) ; Hymers, S (IX-16-23) ;
Merivale (May 1, 1936, W. J. Brown) ; Ottawa, $ (IV-21-1900) ; (3 speci-
mens. May 9, 1904, C. H. Young; March 29, 1906, May 1, 1906, J. Fletcher) ;
Toronto (no further data) ; Trenton (May 5-21, 1911, Evans).

Quebec: Aylmer (April 29, 1904, C. B. Hutchings) ; Chelsea (July 17, 1915,
J. McDunnough) ; Fairy Lake (April 24, 1932) ; Knowlton (September 11,
1929, J. McDunnough); Meach Lake (March 29, April 25, 1900; Sep-
tember 25, 1902; May 4, 1901, C. H. Young).

Rema7'ks. — In addition to a long collected series of this species I
have a specimen before me from Madison, Wis., reared from Myrica
asplenifolia by S. B. Fracker, and four from Dover, Mass. (Gipsy-
Moth Lab.), reared from '"'■Goieopsis'''' (synonym of Bidens). The
wide difference in food plant is very surprising in view of the fact
that nearly all species are either host specific or feed on very closely
related species of plants. The specimen from Bobcaygeon, Ontario,
was reared fi-om Eupatorium.

AGONOPTERIX SCABELLA (Zcller)

PI.ATE 43, FiGTJKB 250

Depressaria scabeUa Zexleb, Verii. zool.-bot. Ges. Wien, vol. 23, p. 236, 1873. —
Chambers, U. S. Geol. Geogr. Surv. Terr. Bull. 4, p. 138, 1878.— Rim:y, in
Smith, List of the Lepidoptera of Boreal America, No. 5280, 1891. — BuscK,
Proc. U. S. Nat. Mus., vol. 24, p. 749, 1902 ; in Dyar, U. S. Nat. Mus. Bull. 52,
No. 5892, 1903. — Kb;aefott, in Smith, List of the Lepidoptera of Boreal
America, No. 6435, 1903. — Meyeick, in Wytsman, Genera insectorum, fasc.
180, p. 177, 1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p.
348, 1939.

Depressaria scahrelJa Walsingham, Proc. Zool. Soc. London, 1881, p. 312.

Agonopteryx scahella (Zeller) Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198,
1908.— FoEBES, Cornell Univ. Agr. Exp. Stat., Memoir 68, p. 239, 1923.

Ago7ioptetix scabella (Zeller) McDunnough, Check List of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera ) ,
No. 8453, 1939.

Agnoptenjx scabella (Zeller) Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6439, 1917.

Alar expanse, 24 mm.

Type. — In the British Museum.

Type locality.— ''Ohio y

Reviiarhs. — I have one specimen, supposedly this species, before
me and have figured the genitalia (fig. 250, ? ) , which show it is very
close to atrodorsella.

AGONOPTERIX PTELEAE Barnes and Busck

Plate 27, Figukes 162, 162a ; Plate 43, Figure 251

Agonopteryx ptclcac Baenes and Busck, Contr. Lepid. North America, vol. 4,
p. 231, pi. 28, fig. 13 ; pi. 38, fig. 1, 1920.

REVISION OF THE OECOPHORIDAE CLARKE 76

Agonoptcrix pteleae (Barnes and Busck) McDunnoiigh, Check list of Lepid-

optera of Canada and the United States of America (Part 2, Microlepidop-

tera), No. 8426, 1939.
Depressaria pteleae (Barnes and Busck) Meyrick, in Wytsman, Genera insec-

torum, fasc. ISO, p. 175, 1922. — Gaede, in Bryk, Lepidopterorum catalogus,

pt. 92, p. 342, 1939.

Labial palpus light ochreous ; second segment with fuscous base and
iiTorations exteriorly; third seginent with fuscous basal and sub-
apical annuli. Antenna fuscous. Head reddish ochreous above ; face
light silvery ochreous, Tliorax and fore wing light ochreous-brown
mottled and suffused with black and brown, the entire surface rough-
ened with raised scales ; extreme base at inner angle whitish ochreous
broadly edged with black; from costa to end of cell a large blackish-
fuscous blotch; along costa and around termen a series of blackish-
fuscous spots ; first and second discal spots black, the latter obscured
by the large costal blotch. Hind wing light ochreous fuscous; cilia
light ochreous brown with basal band a shade lighter. Legs light
ochreous brown suffused and annulated with blackish fuscous. Ab-
domen light ochreous-brown, with two lateral longitudinal rows of
black spots on the underside.

Male genitalia. — Harpe moderately sclerotized, pointed, and almost
entirely clothed with fine hairs; at base, just inside costa, is a group
of long fine spines; clasper and a narrow "spur" of the sacculus adja-
cent to it, heavily sclerotized. The clasper tapers gently to a fine
point, is slightly curved, and reaches almost to costa. Anellus a
heavily sclerotized oval plate, longer than broad, with truncated pos-
terior edge ; lateral lobes large, hairy. Transtilla a narrow sclerotized
band with well-developed, hairy lateral lobes. Aedeagus rather stout,
curved, and pointed, with a lateral, lightly sclerotized flap from about
the middle to the apex. Gnathos a short oval knob armed with many
spines. Socii small, mainly indicated by hairs. Tegumen with a
single apical projection.

Female genitalia. — Ostium large, near posterior edge of genital plate.
Genital plate greatly produced anteriorly and narrowly cleft on an-
terior margin. Extreme posterior portion of ductus bursae abruptly
constricted before ostium, where ductus seminalis enters ; ductus bursae
gently tapering to form the large bursae copulatrix. Signum a roughly
diamond shaped, sclerotized, spined plate.

Alar expanse, 20-22 mm.

Type. — In the United States National Museum.

Type locality. — Decatur, 111.

Food plant — Ptelea trifoliata L. (hoptree).

Remarks. — This species is exceedingly close to pulvipennslla but dif-
fers from it by the rough-scaled fore wing, smaller anellus, and less
sharply pointed anterior margin of the genital plate.

74 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Besides the typer series in the National collection from Decatur, 111.
(June 1-15), there are two specimens from Rockford, 111, (collector
"O, H. S.") among the undetermined material, which, although some-
what larger and darker than the types, undoubtedly belong here. This
species has an unmistakable roughened appearance due to the presence
of many long, slender, almost hairlike, upturned scales on practically
the entire insect.

AGONOPTERIX EUPATORIIELLA (Chambers)

Plate 27, Figukes 163, 163a ; Plate 43, Figihse 249

Depressaria eupatoriiella Chambers, U. S. Geol. Geogr. Surv. Terr. Bull. 4, p. 82,

115, 1878. — Riley, in Smith, List of the Lepidoptera of Boreal America, No.

5261, 1891.
Agonopteryx plummerella Busck, Proc. U. S. Nat. Mns., vol. 35, p. 199, 1908. —

Forbes, Cornell Univ. Agr. Exii. Stat. Memoir 68, p. 239, 1923. (New

synonymy. )
Agonopteri.v plummerella (Busck) McDunnough, Check List of the Lepidoptera

of Canada and the United States of America (Part 2, Microlepidoptera), No.

8454, 1939.
Agnoptenjx phimwcrella (Busck) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6456, 1917.
Depressaria plummerella Meyeick, in "Wytsman, Genera insectorum, fasc. 180, p.

177, 1922. — Gaede, in Bryk, Lepidopterorum catalogus. pt. 92, p. 340, 1939.
Depressaria pnlvipennella Busck, not Clemens (in part), Proc. U. S. Nat. Mus.

vol. 24, p. 737, 1902.

Labial palpus light ochreous-gray : second segment lightly irrorated
with black scales ; third segment almost wholly black, with only nar-
row, poorly defined, median and apical bands of the light ground
color showing; long, recurved. Antenna dark fuscous narrowly an-
nulated with light ochreous. Head, thorax, and fore wing ochreous-
fuscous; face shining light ochreous-gray; thorax and fore wing
speckled with numerous black and white scales, the majority of the
former raised ; at the end of cell a white discal spot surrounded by a
few black scales and preceded by two black discal spots at basal third,
the latter obliquely one above the other ; along costa a series of poorly
defined blackish spots; cilia ochreous-fuscous. Hind wing shining
fuscous, lighter basally than apically; cilia light fuscous, whitish
distally with a narrow dark basal band. Legs whitish ochreous over-
laid and mottled with fuscous except at joints. Abdomen ochreous-
fuscous above, whitish ochreous beneath with four longitudinal rows
of blackish-fuscous spots.

Male genitalia. — Harpe rather short, broad, with the costal two-
thirds clothed with fine hairs ; cucullus rounded ; sacculus and clasper
moderately sclerotized; clasper slightly curved toward cucullus dis-
tally and reaching about two-thirds distance across harpe. Anellus
oval, longer than broad, well sclerotized, with posterior edge concave;
lateral lobes clothed with fine hairs. Vinculum rounded. Aedeagus

REVISION OF THE OECOPHORIDAE — CLARKE 75

slender, bent just before middle, apex pointed, upturned. Transtilla
a narrow, weakly sclerotized band with well-developed, hairy lateral
lobes. Gnathos an oval spined knob. Socii broad flaps, weakly sclero-
tized and with few hairs.

Female genitalia. — Ostial opening very large, occupying over half
t]ie length of the genital plate ; genital plate broad ; anterior margin
with semicircular cleft, the anterior points of which are widely
separated. Ductus bursae long, gradually broadening into the large
bursa copulatrix ; at posterior end of ductus bursae an elongate, lightly
sclerotized patch. Signum a large, strongly sclerotized, diamond-
shaped plate with the anterior and posterior points of the diamond
curled. The plate is armed with short stout teeth.

Alar expanse, 22-24 mm.

Types. — In the Museum of Comparative Zoology, Cambridge, Mass.
{eupatoriiella) ; in the United States National Museum (No. 11943)
{phmimerella) .

Type localities. — "Kentucky" {eup at or Hell a) ; Plunnners Island,
Md. (Busck) ; Cincinnati, Ohio (Dr. A. F. Braun) {phmwierelkf) .

Food plant. — Eupatorium.

Distrihutioii. — Eastern United States probably as far north as New
England.

United States records

Illinois: Putnam County, 9 (14-IV-1939, M. O. Glenn).

Maryland: Plummers Island, 10 $ S , Q 9 9 (March to August dates, 1905-1908,

A. Busck and H. S. Barber).
Ohio: Athens, 3 $$,2 $9 (ll-XI-39, 29^1-40, 10-11-40, R. C. Barnes);

Noble County, $, 9 (ia-IX-39, 31-XII-40, R. C. Barnes).
Pennsylvania: Pittsburgh, $, 9 (4-IV-06, lS-XI-06, Henry Engel).

Remarks. — This species is apparently very close to scahella Zeller,
but since I have no authentic material of the latter species, I carniot
be sure of the genitalia. I quote, in part, a letter from Mr. Stringer
of the British Museum, to Mr. Busck concerning plummerella and
scdbella.) which was written before I had an opportunity to examine
eupatoriiella :

"I have carefully compared plwnmerella with our type of scdbella
Z., and have no doubt that they are distinct. The labial palpi of
scdbella have the Deprcssaria "furrow" on second joint but much more
compact and the terminal joint is shorter, not so pointed or recurved
as in your species and nebvlosa Z. ; in color markings there is no
difference. In size and color of fore wings your species is nearer to
nehulosa; scaheUa is a smaller species with mucli narrower fore wing
and is brown-gray, not salt and pepper ; it is also devoid of any mark-
ings on the underside of either wing as in jjlummereUa and almost all
of the species of the genus."

76 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

I have examined the type of eupatoriieUa in the Museum of Com-
parative Zoology and have made a slide of the female genitalia.
This and plumineTella appear to be identical ; ewpatoriiella was syn-
onymized with pulvipeiineUa Clemens through misidentifications made
by Lord Walsingham. The specimens in the United States National
Museum, referred to by Busck,^^ are clearly p^dvipenneUa; pulvipen-
nella lacks the rough scaling which is found on the wings of eupatori-
ieUa and so cannot be confused with it. Chambers clearly states in
his description of eupatoriieUa^ "the palpi and wings dusted with
blackish atoms, each of which is a minute tuft * * *"

AGONOPTERIX PULVIPENNELLA (Clemens)

Plate 27, Figures 164, 164a ; Plate 43, Figure 253

Depressaria pulvipennella Clemens, Proc. Ent. Soc. Philadelphia, vol. 2, p. 421,
1864.— Robinson, Ann. Lye. Nat. Hist. New York, vol. 9, p. 157, 1S68.—
Chambers, Can. Ent., vol. 4, p. 91, 1872. — Clemens, m Stalnton, Tineina of
North America, p. 244, 1872. — Riley, in Smith, List of the Lepidoptera of
Boreal America, No. 5276, 1891.— Busck. Proc. U. S. Nat. Mus., vol. 24, p. 737,
1902 ; in Dyar, U. S. Nat. Mus. Bull. 52, No. 5858, 1903.— Kearfott, in Smith,
List of the Lepidoptera of Boreal America, No. 6401, 1903. — Meybick, in
Wytsman, Genera insectorum, fasc. ISO, p. 176, 1922. — Gaede, in Bryk,
Lepidopterorum catalogus, pt. 92, p. 342, 1939.

Depressaria fulvipcvnelJa Dietz, in Smith, Catalogue of the insects of New
Jersey, p. 474, 1900.

Depressaria soUdagivis Walsingham, Ins. Life, vol. 1, p. 255, 1899. — Riley, in
Smith, List of the Lepidoptera of Boreal America, No. 5281, 1891.

Agonopteryx pnlvipcnneUa (Clemens) Busck, Proc. U. S. Nat. Mus., vol. 35,
p. 198, 1908— Forbes, Cornell Univ. Agr. Exp. Stat. Memoir G8, p. 238, 1923.

Agonopterix pulvipennella (Clemens) McDunnough, Check list of the Lepidop-
tera of Canada and the United States of America (Part 2, Microlepidop-
tera). No. 8410, 1939.

Agnopteryx pulvipennella (Clemens) Smith, Catalogue of the insects of New
Jersey, p. 561, 1910. — Barnes and McDunnough, Check list of the Lepidoptera
of Boreal America, No. 6449, 1917.

Agnopteryx pulvipenella Brimley, The insects of North Carolina, p. 304, 1938.

Labial palpus whitish ochreous tinged with pink; second segment
irrorated with blackish fuscous exteriorly; third segment with basal
and subapical annuli and tip black. Antenna fuscous. Face shining
whitish ochreous. Head reddish ochreous suffused with fuscous pos-
teriorly. Thorax and fore wing light reddish ochreous mottled and
irrorated with blackish fuscous; extreme base of the light ground color
and followed by a rapidly fading fuscous shade that does not reach
costa; at basal third two black discal spots obliquely one above the
other, followed by a third, white discal spot at the end of cell; in
center of wing beyond the first and second discal spots a blackish-
fuscous shade surrounding the third discal spot ; along costa and around

"Busck, Proc. U. S. Nat. Mus., vol. 24, p. 737, 1902.

REVISION OF THE OECOPHORIDAE — CLARKE 77

termen a series of blackish-fuscous blotches and spots; cilia light
ochreous-fuscous. Hind wing light fuscous; cilia somewhat lighter
with a dark basal band. Legs whitish ochreous suffused and mottled
with blackish fuscous except at joints ; at joints the ochreous is strongly-
tinged with pink. Abdomen grayish ochreous above ; beneath whitish
ochreous with broad blackish-fuscous longitudinal lateral lines and
two rows of blackish-fuscous spots between the lateral lines.

Male genitalia. — Harpe broad, clothed with fine hairs; clasper
straight, slender, reaching well beyond middle of harpe; cucullus
bluntly pointed. Anellus a large oval plate, longer than broad, with
concave posterior margin, narrow biramous, strongly sclerotized me-
dian area and well developed hairy, lateral lobes. Aedeagus slender,
sharply curved, pointed; vesica with large spinulate patch. Vincu-
lum rounded. Transtilla a moderately broad, sclerotized band, with
well developed, hairy, lateral lobes. Gnathos an oval, spined knob.
Socii large, sparsely hairy, fleshy flaps.

Female genitalia. — Ostium small, near posterior margin of genital
plate. Genital plate very broad, produced anteriorly and deeply and
narrowly cleft o\\ median anterior edge. Ductus bursae long, mem-
branous, gradually tapering into the large bursa copulatrix ; inception
of ductus seminalis very near ostial opening. Signuni a large, dia-
mond-shaped, spined plate; posterior and anterior points curled.

Alar expanse, 18-24 mm.

Type. — In the Academy of Natural Sciences of Philadelphia.

Tyfe locality. — "Illinois."

Food plants. — Solidago and Urtica.

Distribution. — Eastern United States and Canada.

United States records

Connecticut: East River, 5 (September 3, 1908, Chas. R. Ely).

District of Columbia: Washington, 3 5 5, 9 (IV-21-85) ; S (IV-7-1900) ;

S (IV-21-1900).
Louisiana: Natchitoches, $ (August, G. Coverdale).
Maine: Orono, 2 S $ (October 8, '88, one, no date).
Maryland: Plummers Island, 17 $ S , 4 9 9 (March, April, and September dates,

collected by H. S. Barber, A. Busck, E. A. Schwarz).
Massachusetts: Chilmark, $ (IX-3-1935, George D. Eustis) ; Cohasset, 9 (May

11, '07, Owen Bryant) ; Framingham, 5 (April 21, 1905) ; Springfield, $

(nodate, G. Dimmock) ; Winchendon, S (IX-28-02).
Missouri: (One specimen, no date.)
ISTew Hampshire: Hampton, 6 (IV-30-04, S. A. Shaw).
Few Jersey: Denville, 5 ("X-14") ; Essex County, 9 (IV-20-09, W. D.

Kearfott).
Nev/ York: Ilion, $,2 99 (IX-3-11, H. McElhose) ; 9, same (IX-2-12) ;

Ithaca (many $ $ and 9 9 , various dates in coll. CornellUniv. ) .
Ohio: Athens, 2 9 9 (29^1-40, R. C. Barnes).

Pennsylvania: New Brighton, c5 , 9 (IX-27~02, X-^S-OT, II. D. Merrick).
Virginia: Vienna, 9 (February 14, 1915, R. A. Cushman).
Wisconsin: Cranmoor, V/ood County, $ (IV-22-08, C. B. Hardenberg).

78 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Canadian records

Manitoba: Cartwright, 9 (X-11-03, E. F. Heath).

Nova Scotia: Bridgetown, $, ? (5-9-IX-1912, G. E. Saunders).

Ontario: Ottawa ( 5, April 28, 1905; $, May 7, 1905, C. H. Young) ; Trenton,

$ (25-IV-1911, Evans).
Quebec: Meacli Lake (9, April 19, 1902; October 12, 1907, C. H. Young);

Montreal, S (V-3-04, "A. F. W.").

Remarks. — The variety of food plants indicates that this species
is a rather general feeder. I believe we are dealing with at least
two very closely related species, which can be separated only by a
careful comparison of larvae and pupae, the adult characters not
being sufficient for separation. Moths reared from Solidago are
darker, and the coloring is more suffused than in the majority of
specimens. This may be due entirely to the difference in food plant,
but there is not sufficient material at hand to determine this
definitely.

The species appears to be confined to midwestern and eastern North
America.

AGONOPTERIX WALSINGHAMELLA (Busck)

Plate 28, Figukes 165, 165a ; Plate 44, Figuee 254

Depressaria walsinghamella Busck, Proe. U. S. Nat. Mus., vol. 24, p. 739, 1902. —
Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 356, 1939.

Depressaria fernaldella Walsingham, Ins. Life, vol. 1, p. 256, 1889. — Busck, in
Dyar, U. S. Nat. Mus. Bull. 52, No. 5S64, 1903.— Kearfott, in Smith, List
of the Lepidoptera of Boreal America, No. 6407, 1903. — Meyeick, in Wyts-
man, Genera insectorum, fasc. ISO, p. 176, 1922. [Not Depressaria fernal-
della Chambers, U. S. Geol. Geogr. Surv. Terr. Bull. 4, p. 82, p. 138, 1878.]

Agnopteryx fernaldella Baenes and McDunnough, Check list of the Lepidoptera
of Boreal America, No. 6443, 1917.

Agonoptcrix fernaldella (Walsingham) McDunnough, Check list of the Lepidop-
tera of Canada and the United States of America (Part 2, Microlepidop-
tera), No. 8443, 1939.

Agonopteryx walsinghamella Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68,
p. 239, 1923.

Agonopteryx icalsinghamiella Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198,
1908.

Agonopterix walsingTiamiella McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera ) , No.
8443, 1939. [As synonym of fernaldella (Walsingham).]

Labial palpus pale cinereous strongly suffused with reddish ; second
segment irrorated with blackish scales exteriorly ; third segment with
blackish scales exteriorly; third segment with blackish-fuscous basal
and median annuli (the former poorly defined) and apex. Antenna
fuscous annulated with dull cinereous and tinged with reddish. Head
tawny-red with slight cinereous irrorations; face pale cinereous.
Thorax and base of fore wing, except costal part, cinereous ; fore part

REVISION OF THE OECOPHORIDAE — CLARKE 79

of thorax and tegula suffused and irrorated with reddish. Fore wing
deep crimson-red sparsely irrorated with black and cinereous scales;
costa, nearly to apex, broadly tawny-red with veins 9 to 12 strongly
indicated by cinereous and irrorated with black scales and with poorly
defined fuscous spots on extreme edge; discal spots white; first discal
spot at basal third edged with carmine; second at end of cell preceded
by a few carmine scales; cilia reddish fuscous edged with carmine.
Hind wing light fuscous, darker apically than basally; cilia light
fuscous with whitish irrorations and pale pink suffusions. Legs pale
cinereous overlaid and irrorated with fuscous, except at joints, and
suffused with reddish. Abdomen pale grayish fuscous above ; beneath,
pale cinereous with two black longitudinal lateral stripes.

Male genitalia. — Harpe moderately clothed with fine hairs, tapering
gently to a pointed cucullus; clasper short, stout, straight, scarcely
exceeding middle of harpe, slightly rugose on exterior edge. Sacculus
narrow, moderately sclerotized. Anellus longer than broad, concave
on posterior edge, narrowed basally, with well-developed, sparsely
hairy lateral lobes. Aedeagus stout, curved, much narrower in distal
than proximal half; vesica armed with numerous spinulate cornuti.
Vinculum rounded. Transtiila a narrow, lightly sclerotized band with
moderately well developed, hairy lateral lobes. Gnathos a spined oval
knob. Socii fleshy, hairy flaps. Tegumen terminating in a moderately
long, pointed process.

Female genitalia. — Genital plate broad, lightly sclerotized. Ostium
round, opening at posterior edge of genital plate; the area around
ostium slightly raised above the remainder of surface of plate.
Ductus bursae long, membranous; ductus seminalis opening well
before ostium. Bursa copulatrix large with small signum situated
about middle.

Alar expanse, 20-22 mm.

Type. — In the British Museum.

Type localities. — Orono, Maine; Wisconsin.

Food plants. — Myrica asplenifolia L. and Myrica carolinensls Mill.

Distiihutian. — Eastern United States and Canada.

United States records

Connecticut: East River, $ (August .j-13, Chas. R. Ely).

Maine: Bar Harbor, 8 S$, 2 (May to November dates, 1934-193S, A. E.

Brewer) ; Kennebunkport, 2 $ $ (September 24-30) ; Orono, 9.
Massachusetts: Springfield (a series of 4 5 5 and 3 2 9 reared from Myrica

asplenifolia by Dimmock).
New Hampshire: Hampton, 8 S S (March and April dates; S. A. Shaw, coll.).
New York: Albany.

Pennsylvania: Hazel ton, $ ("9-12-04").
Wisconsin: (Ace. Forbes.)

80 PROCEEDINGS OF THE NATIONAL ]VIUSEUM vol. 90

Canadian records

Nova Scotia: Digby, $ (IX-19-07, J. Russell) ; White Point Beach, Queens

County, 5 (VIII-20-34, reared, J. McDunnough).
Ontario: Biscotasing (May 16, 1931, Karl Schedl) ; Constance Bay (October 3,

1932, W. J. Brown).
Quebec: Kazubazua (August 13-23, 1933, G. S. Walley) ; Laniel (June 3, 1932,

H. S. Fleming).

Remarks. — The identity of this species and the invalidity of
Walsingham's name have been established by Busck.-"

AGONOPTERIX FUSCITERMINELLA, new species

Plate 28, Figuees 167, 167a ; Plate 44, Figuke 258

Depressaria yeatiana Walsingham (not Fabricius) Proc. Zool. Soc. London,

1881, p. 316.
Depressaria arenella Wai^singham (not Schiffermiiller), Trans. Amer. Ent. Soc,

vol. 10, p. 175, 1882. — Riley, in Smith, List of the Lepidoptera of Boreal

America. No. 5253, 1891.— Busck, Proc. U. S. Nat. Mus., vol. 24, p. 743, 1902 ;

in Dyar, U. S. Nat. Mus. Bull. 52, No. 5877, 1903.— Kearfott, in Smith,

Check list of the Lepidoptera of Boreal America, No. 6420, 1003. — Meyeick,

in Wytsman, Genera insectorum, fasc. 180, p. 174, 1922.
Agonopteryx arencUa Busck (not Schiffermiiller), Proc. U. S. Nat. Mus., vol.

35, p. 199, 1908. — Barnes and McDunnough, Check list of the Lepidoptera

of Boreal America, No. 6446, 1917.

Antenna fuscous, faintly and narrowly annulated with ochreous;
apex and underside of basal segment whitish ochreous sprinkled with
black or dark-brown scales outwardly; palpus with brush well de-
veloped; terminal segment with black subbasal and supramedial
annuli and black tip. Face and head whitish ochreous, the latter
intermixed with brownish scales. Thorax whitish ochreous ; dorsally
the ground color is liberally mixed with brown and blackish scales.
In the posterior half are two crests of raised scales. Fore wing with
ochreous ground color strongly overlaid with reddish brown and
liberally sprinkled with fuscous to black scales ; extreme base ochreous
followed, in dorsal half, by a strong red-brown patch, the latter
mixed with black in basal part; at the base of cell two small obliquely
placed black spots followed by a strong fuscous or black cloud; the
latter surrounded by a reddish-brown suflfusion; at the end of cell
another small black spot. Costa and termen with conspicuous black
spots. Cilia ochreous with brown scales intermixed and the whole
with a distinct rosy tinge. Hind wing whitish, strongly suffused
apically with fuscous; scales at base of cilia, along termen. and fre-
quently around posterior margin, black. Cilia light brownish
ochreous. Underside of hind wing with many black scales in apical
half. Legs ochreous ; fore leg with femur fuscous inwardly ; anterior
surface of tibia bright pink with overlying brownish scales, the

2« Busck, Proc. U. S. Nat. Mus., vol. 24, p. 739, 1902.

REVISION OF THE OECOPHORIDAE — CLARKE 81

latter having a purplish sheen: tarsi broadly edged with fuscous.
Mesothoracic legs like anterior pair but without fuscous femora and
with more fuscous on tibiae and tarsi. Hind legs almost wholly
whitish ochreous except for delicate pink suffusion on tibiae and
fuscous tibial spurs. Abdomen ochreous with broken black lateral
lines beneath and suffused with fuscous above.

Male genitalia. — Harpe elongate, narrow ; weakly sclerotized except
for clasper, sparsely clothed with fine hairs; clasper straight, blunt,
pointed, reaching about two-thirds distance across harpe. Anellus
longer than broad, subrectangular, moderately sclerotized; posterior
edge convex; lateral lobes little developed. Transtilla a narrow
sclerotized band with weak lateral lobes. Aedeagus moderately slen-
der, with sharp, upturned point. Gnathos an oval, spined knob.
Socii well developed hairy flaps.

Female genitalia. — Base of lobe of ovipositor with a row of long
stout hairs. Ostium moderately large, round. Genital plate broad,
moderately sclerotized. Ductus bursae long, membranous; inception
of ductus seminalis near opening of ostium. Bursa copulatrix large,
oval, symmetrical, merging gradually with the ductus; gignum a large,
roughly diamond-shaped, spined, sclerotized plate.

Alar expanse, 22-25 mm.

Type.—V.S.'^.M. No. 52077.

Type locality. — Duncan, Vancouver Island, British Columbia.

Food plant. — Cynoglossum grande Dougl. ex Lehm.

Remarks. — Described from the $ type and b $ $ and 13 $ 2
paratypes as follows : Type $ , Duncan, Vancouver Island, British
Columbia (July 1, 1910, Hanham) ; paratypes, Z $ $ and 7 $ 9 ,
Duncan, British Columbia (June and July dates, Hanham) ; $
and 2 5 2, Wellington, British Columbia (April and October dates,
G. W. Tavlor) ; $ and 4 5?, Phoenix Lake, Marin County, Calif.
(V-11 to VI-2-1927, H. H. Keifer).

Paratypes in the U. S. National Museum and H. H. Keifer collec-
tion, Sacramento, Calif.

In superficial appearance very much like the European arenella
but easily differentiated from it by the genitalia; in the male of
arenella the clasper is very broad and flattened, with a deep excavation
on the outer edge, while the clasper of fuscitemiinella is straight,
fingerlike ; the harpe of arenella is shorter and broader than that of
fusciterminella. In the female of arenella the anterior edge of the
genital plate is strongly convex but that of fusciterminella is much
less strongly so.

The California specimens are somewhat darker than those from
British Columbia, but this may be due to their having been reared.

286614—41 4

82 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

The northern specimens show more pink on the legs also, but they are
identical in other characters, including genitalia.

This species has been confused with pallidella Busck and has been
placed in collections under that name.

AGONOPTERIX CAJONENSIS, new species

Plate 31, Figures ISO, 180a ; Plate 42, Figures 244, 244a

This species is much like argilJacea but lacks the median dark shade
of the fore wing.

Labial palpus light ochreous-white, lightly mottled exteriorly on
second segment with fuscous ; third segment with black subbasal and
supramedial bands and tip; the basal band is poorly defined. An-
tenna fuscous with ochreous-white scaling on basal segment and basal
third. Head, thorax, and ground color of fore wing grayish fuscous
heavily overlaid with ochreous-white ; face light ochreous-white ; base
of fore wing and basal third of costa ochreous-white ; the light basal
portion is followed by a dark fuscous shade; whole surface of fore
wing sparsely irrorated with small fuscous spots; at the end of cell
a conspicuous ochreous-white spot narrowly edged with fuscous;
cilia grayish fuscous. Hind wing grayish fuscous; cilia concolorous
with light basal band. Legs grayish fuscous strongly overlaid with
ochreous-white. Abdomen grayish fuscous above with posterior mar-
gins of segments ochreous-white ; beneath ochreous-white with a well-
defined lateral stripe on each side and a pair of black spots on the
posterior margin of each segment between the lateral lines.

Male genitalia. — Harpe rather sharply narrowed before cucullus;
cucullus bluntly pointed; clasper long, curved, slightly dilated
distally ; sacculus moderately sclerotized. Anellus longer than broad,
slightly constricted at about middle; posterior edge concave; lateral
lobes large, extending nearly to posterior edge of central plate.
Aedeagus slender, curved, pointed, with ventral sclerotized arm by
which it is attached to the anellus; vesica armed with an elongate
patch of fine cornuti. Vinculum rounded, with well-developed dorso-
anterior process. Transtilla a moderately broad sclerotized band
with well-developed lateral lobes. Tegumen pointed.

Ferruile genitalia. — Genital plate very broad, strongly sclerotized.
Ostium small, crescentic, opening slightly posterior to middle. Ductus
bursae long, membranous; inception of ductus seminalis just before
ostium. Bursa copulatrix large, oval, symmetrical, with well-
developed 4-pointed signum.

Alar expanse, 20-23 mm.

Type.—V.S.^.M. No. 52948.

Type locality. — Cajon Valley, Calif.

Food plant. — Unknown.

REVISION OF THE OECOPHORIDAE — CLARKE 83

Remarks. — Described from the type 5 and 1 9 paratype (Cajon
Valley, Calif., 25-VI-1937, Grace H. and John L. Speiry).

Both specimens were received from ISIr. Sperry in material sub-
mitted for determination.

AGONOPTERIX ROSACILIELLA (Busck)

Plate 28, Figures 166, 166a ; Plate 44, Fiquke 257

Depressaria rosacilieUa Busck, Proc. U. S. Nat. Mus., vol. 27, p. 763, 1904.^ —
Anderson, Catalogue of British Columbia Lepidoptera, No. 1093, 1904. —
Gaede, in Bryk, Lepidoptororum catalogus, pt. 92, p. 346, 1939.

Depressaria ciliella Walsingham (not Stainton), Proc. Zool. Soe. London, 1881,
p. 316.— Busck (not Stainton), Proc. U. S. Nat. Mus., vol. 24, p. 739, 1902; in
Dyar, U. S. Nat. Mus. Bull. 52, No. 5863, 1903. [Not Depressaria ciliella
Stainton, Trans. Ent. Soc. London, 1849, p. 161, pi. 17, fig. 7; or Rebel, in
Staudinger and Rebel, Catalog der Lepidopteren des palaearctischen Faunen-
gebietes, vol. 2, No. 3234, 1901.]

Depressaria rosiciliella Meyekick, in Wytsman, Genera insectorum, fasc. ISO, p.
176, 1922.

Agonopteryx rosacilieUa Busck, Proc. U. S. Nat Mus., vol. 35, p. 198, 1908l —
Braun, Proc. Acad. Nat. Sci. Philadelphia, vol. 73, pt. 1, p. 10, 1921.

Agonopterix rGsaciliella (Busck) McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera), No,
8431, 1939.

Agonopteryx ciliella Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 239, 1923.

Agnopteryx rosacilieUa (Busck) Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6448, 1917.

Labial palpus ochreous-white suffused with pale reddish; second
segment mottled exteriorly with blackish fuscous ; third segment with
basal and subterminal bands and tip black. Head, thorax, and fore
wing ochreous-white heavily overlaid with reddish, suffused with red-
dish fuscous, and irrorated with sparse blackish scales ; face whitish ;
light basal portion of fore wing containing a fuscous spot and fol-
lowed by a deep reddish- fuscous shade, the latter diminishing in in-
tensity outwardly; costa and termen with a series of more or less
distinct fuscous spots, those on costa alternated with small whitish-
ochreous spots; discal spots four, white, edged with black; first and
second obliquely one above the other, third and fourth in line with
second, the fourth at end of cell ; cilia fuscous with strong reddish suf-
fusion. Hind wing and cilia light fuscous, the latter with a strong
reddish suffusion. Legs ochreous-white with reddish tinge and
strongly overlaid and irrorated with blackish fuscous except at joints.

Male genitalia. — Harpe long, lightly sclerotized, bluntly pointed,
very hairy, especially in costal half. Clasper straight, slender, grad-
ually tapering to a blunt point, reaching beyond middle of harpe.
Vinculum produced anteriorly to form a broad, rounded, short pro-
jection. Anellus a more or less rectangular plate, longer than broad,
slightly emarginate on posterior edge, with large, hairy, lateral lobes.

§4 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Aedeagus slender, not heavily sclerotized ; dilated basally and sharply
pointed distally; the point is slightly upturned; vesica armed with
numerous fine, long cornuti. Transtilla a weakly sclerotized band
with slightly hairy, lateral lobes. Gnathos conical, slender; covered
with fine spines. Socii fleshy lobes mainly indicated by hairs.

Female genitalia. — Genital plate broad, moderately sclerotized.
ostium situated at about center of plate. Ductus bursae very long
(permitting the bursa coupulatrix to reach extreme anterior portion
of abdomen), slender, gradually becoming larger to form the sym-
metrical bursa copulatrix; inception of ductus seminalis well before
ostium. Signum a small, lightly sclerotized, spined, bilobed plate.

Alar expanse, 20-26 mm.

Type. — ^In the United States National Museum.

Type locality.— ''CdiXiv^ Watson," Oreg. (III-IV-1872, Walsing-
ham).

Food plants. — OsmorJiiza (Dr. A. F. Braun, 1921) ; 0. occidentalis
Torr. (Clarke, 1935).

Note. — Osmorhiza occidentalis Torr. Although the three species
of Osmorhiza [occidentalis., divaricata., and hi^evipes) that are in-
digenous to the State of Washington were found at Godman Springs,
Blue Mountains, at an altitude of approximately 6,000 feet, larvae
infested only O. occidentalis. An extended search was made to
determine the host specificity of the species, and in all localities where
the food plants were found rosaciliella attacked only O. occidentalis.

Distribution. — Northwestern United States and southwestern
Canada.

United States records

Montana: Glacier National Park, $ (August 7, 1920, A. F. Braun).
Oregon: "Camp Watson," 9 (III-IV-1872, Walsingham).

Washington: Godman Springs, Blue Mountains, alt. 6,000 feet, 8 S S (VIII-3
to 8-35) ; Pullman, 2 (III-1-98, C. V. Piper).

Canadian records

British Columbia: Kaslo, $ (Dyar and Cockle) ; Trout Lake, $ (lS-VIII-1937,
H. Leech).

Saskatchewan: Scott, 9 (20-X-1923, Kenneth M. King).

Remarks. — I have before me a series of 12 specimens reared from
Osmorhiza occidentalis and 90 from Echinopanax horridum. The
former series is from the interior of Washington State, while the
latter is from the Puget Sound (coastal) region.

For some time I have thought that these two series represented two
distinct but closely related species. I am now of the opinion, how-
ever, that they are conspecific, but that the coastal specimens represent
a distinct race. The interior specimens are, with one exception, con-
siderably darker and more suffused than those from the Puget Sound

REVISION OF THE OECOPHORIDAE — CLARKE 85

area, but on genitalia characters they are identical, and for this
reason I do not feel justified in separating them specifically. The
larvae show a few differences in coloration, but larvae of this group
are likely to vary even though they may come from eggs of the same
female. No doubt the color variation existing between the two groups
of specimens is induced by differences in food plants. The habits of
the larvae are much the same, both forming rolls, but those on
Osmorhiza have an easier time of it because of the small size of the
leaves on which they work; those on Echinopanax have to cut the
leaf from the edge inwardly first and then form the roll.

I am including a description of the lai-^'ae from both localities so
that their similarities and differences may be appreciated better. In
the main the larvae are the same.

Dr. Braun has been kind enough to send me a specimen from Yellow-
stone National Park reared by her from Osmorhiza. It resembles the
Puget Sound series more closely than do the eastern Washington
specimens.

The larva is as follows : Length, 15-16 mm. Head dull yellowish
brown, spotted with a darker brown posterolaterally ; from the base of
the antenna, extending in a line to the posterior edge of the head, is a
broad blackish-brown band constricted in the middle (in some speci-
mens the band is considerably broader than in others) ; ocelli light
brown. The margins of the underside of the head are blackish brown.
Cervical shield pale green, broadly edged laterally and posteriorly with
black and bisected with a fine whitish-green median line. Thoracic
and abdominal segments bright green heavily suffused with reddish
purple in the dorsal half when the larva is mature. Posteriorly on
the segments the reddish-purple color entirely obscures the green. In
the dorsal half there are three brownish-fuscous longitudinal stripes,
the outer ones fully twice as broad as the median one. On abdominal
segments 2-7 inclusive, near the dorsoanterior margin, on each side
of the dorsal stripe, is an outwardly oblique line of 4 to 6 small green
spots. Between the longitudinal lines are several small green spots
generally following folds in the skin. Anal plate green, heavily suf-
fused with reddish purple. Tubercles black centrally, whitish green
outwardly. Spiracles ringed with black, with a whitish-green suffu-
sion outwardly. Thoracic legs concoiorous with segments.

Dr. A. F. Braun first discovered larvae of this species in Yellow-
stone National Park. She has sent me one bred specimen from this
lot of larvae. Although the food plant is widespread throughout the
inland empire, the larvae are very difficult to locate. Much time has
been spent in a search for these larvae in order to clear up the status
of this and related species. On July 13, 1935, a stand of Osmorhiza
was found at Godman Springs, Blue Mountains, Columbia County,

5b PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Wash., at an altitude of 6,000 feet. After a somewhat prolonged
search ten larvae and one pupa were collected, the latter being the
first found.

The larva is distinctly a leaf roller, not invading the umbels as is
done so commonly by members of Agonopierix and Depressaria. The
pupa was found in an umbel, but there was no sign of feeding having
taken place, and I do not think it is the usual habit for pupation to
occur there. It seems more likely that pupation generally occurs
about the base of the plant in debris. This larva is one of the most
active in the group, being easily disturbed and very difficult to capture.

Pupa : Normal for group : "Wing, antenna, and leg sheaths bright
green. Abdominal segments green heavily suffused with reddish pur-
ple dorsally. On the second day the color begins to deepen; the
sheaths become a duller and darker green and the abdominals a light
reddish brown. The color gradually becomes darker until the pupa
is almost black.

The pupal period is lS-14 days.

This species is very close to the European cilieUa and was identified
as such by Walsingham. The American rosaciUeJla has a much more
slanting termen and appears to be a more narrow-winged species than
cilieUa. The genitalia of the male of rosaciliella show a truncated
anellus, while in cilieUa the anellus is concave. The harpes of the
former species are less pointed and broader than those of the latter.

The abdomen of the type is lost, but I do not hesitate to place the
reared series under this name.

AGONOPTERIX ROSACILIELLA ECHINOPANICIS, new variety

Similar to typical rosoxiilieUa but may be distinguished from it by
the generally lighter color and less suffused and less smoky appear-
ance. The black scaling is sparser and more sharply contrasted.

The male and female genitalia are the same as those of typical
ros(wiliella.

Alar expanse, 20-2G mm.

Type.—^]. S. N. M. No. 52252.

Type locality. — Skyline Eidge, Mount Baker district, Whatcom
County, Wash. Altitude 2,500-3,000 feet.

Food plant. — Echinopanax horridwm (Smith) Dene, and Planch.

Remarks. — Described from the type $ and 78 $ and $ paratypes
as follows : Skyline Eidge and Bagley Creek, Whatcom County, Wash.,
altitude 2,500-3,000 feet, 35 5 5 and 40 $ 9 (September 5-12, 1933
and 1935, J. F. G. Clarke) ; Fraser Mills, British Columbia, 2 9?
(IV-10-21, IV-4-23, L. E. Marmont) ; Vancouver, British Columbia,
9 (no date).

REVISION OF THE OECOPHORIDAE — CLARKE 87

Paratypes in the United States National Museum, Canadian Na-
tional collection, and the collection of H. H, Keifer, Sacramento,
Calif.

The larva is as follows: Length, 14-16 mm.; head dark brov/n to
black. Thoracic shield greenish to light brown bisected by a whitish
median line and broadly bordered with black on the posterolateral
edges. Thoracic and abdominal segments green, the latter suffused
dorsally with reddish; from the mesothorax, in the dorsal half, two
broad longitudinal brown stripes; a third median longitudinal stripe
of the same color beginning at the first abdominal segment and con-
tinuing posteriorly the full length of the body. Thoracic legs green-
ish. Tubercles black. Spiracles edged with black. Anal plate light
brown.

Pupa : At first yellowish green with the abdominal segments suffused
with reddish dorsally. In 24 hours the wing sheaths become green
and the abdominal segments reddish brown. The pupa finally turns
deep reddish brown; wing sheaths nearly black, but they still retain
a greenish sheen.

The pupal period of 12 to 15 days.

AGONOPTERIX NOVI-MUNDI (Walsingham)

Pl.\te 28, Figures 168, 168a

Depressaria paHJeUa novi-mundi Walsingham, Ins. Life, vol. 1, p. 256, 1889'. —
Riley, in Smith, List of the Lepidoptera of Boreal America, No. 5273,
1891.— BuscK, Proc. U. S. Nat. Miis., vol. 24, p. 74, 1902.

Depressaria novimiindi (Walsingham) Busck, in Dyar, U. S. Nat. Mus. Bull.
52, No. 5866, 1903.— Keakfott, in Smith, List of the Lepidoptera of Boreal
America, No. 6409, 1903. — Meyrick, in Wytsman, Genera insectorum, fasc.
ISO, p. 175, 1922.— Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p.
335, 1939.

Agonoptcryx novimundi (Walsingham) Busck, Proc. U. S. Nat. IMus., vol. 35,
p. 198, 1908.

Agonopterix novimundi (Walsingham) McDunnough, Check list of the Lepi-
doptera of Canada and the United States of America (Part 2, Microlepidop-
tera), No. 8428, 1939.

Agnopteryx noinmundi (Walsingham) Barnes and IMcDunnough, Check list of
the Lepidoptera of Boreal America, No. 6454, 1917.

Labial palpus sordid whitish; irrorated with blackish fuscous
inwardly and outwardly on the second segment and in the brush;
third segment with black subbasal and supramedial annuli. An-
tenna blackish fuscous with grayish annulations. Head, thorax, and
ground color of fore wing grayish fu.scous, the latter heavily overlaid
with dull purplish fuscous ; head scales and dark scales of fore wing
tipped with cinereous; at base of fore wing an indistinct fuscous
patch followed by a poorly defined yellowish-brown shade; at basal
third an outwardly oblique black dash in disk edged with yellowish

88 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

brown ; at end of cell an indistinct white discal spot ; between veins,
in apical third, yellowish-brown shading; along costa and around
tei-men a series of poorly defined fuscous spots; cilia fuscous tipped
with white. Hind wing shining yellowish fuscous; cilia concolorous,
lighter apically. Legs cinereous strongly overlaid with shining yel-
lowish fuscous except at joints.

Male genitalia. — Harpe broad, sparsely clothed with hairs; clasper
slender, pointed, dilated about middle, reaching two-thirds distance
to costa. Anellus more or less rectangular, longer than broad, con-
cave on each side before posterior end and also on posterior margin;
lateral lobes weak, hairy. Vinculmn rounded. Aedeagus stout, bent,
pointed, the point slightly upturned; vesica with a weak spinulate
patch about middle. Transtilla a broad, sclerotized band with large
hairy lateral lobes. Gnathos an oval, spined knob. Socii large hairy,
fleshy flaps.

Alar expanse, 16 mm.

Type. — In the British Museum.

Type localities. — Mount Shasta, Siskiyou County, Calif., and
"North Oregon."

Remarks. — ^Various specimens in collections have been associated
with this species, but the only two I have seen that belong under this
name are paratypes of Lord Walsingham's that he deposited in the
National collection. I have seen no females.

AGONOPTERIX PALLIDELLA (Bnsck)

Plate 32, Figubes 182, 182a ; Plate 44, Figure 256

Depressaria pallidella Busck, Proc. U. S. Nat. Mus., vol. 27, p. 765, 1904. — Ander-
son, Catalogue of British Columbia Lepidoptera, No. 1095, 1904. — Meybick,

m Wytsman, Genera insectorum, fasc. 180, p. 174, 1922. — Gaede, in Bryk,

Lepidopterorum catalogus, pt. 92, p. 337, 1939.
Agnopteryx pallidella (Busck) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6461, 1917.
Agonopterix pallidella (Busck) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera), No.

8418, 1939.
Agonopteryx terinella Barnes and Busck, Contr. Lepid. North America, vol. 4,

p. 232, pi. 28, fig. 15, 1920.
Agonopterix terinella (Busck) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera), No.

8418, 1939. (As synonym of pallidella (Busck).)
Depressaria terinella Meybick, in Wytsman, Genera insectorum, fasc. 180,

p. 174, 1922.
Agonopteryx serrae Claeke, Can. Ent., vol. 65, p. 84, pi. 5, 1933.
Agonopterix serrae (Clarice) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera), No.

8418, 1939. (x\s synonym of pallidella (Busck).)
Depressaria serrae (Clarke) Gaede, in Bryk, Lepidopterorum catalogus, pt. 92,

p. 350, 1939

REVISION OF THE OECOPHORIDAE — CLARKE 89

Labial palpus whitish ochreous, second segment irrorated with
blackish fuscous exteriorly; third segment with subbasal and supra-
medial bands blackish fuscous. Antenna fuscous. Head, thorax,
and fore wing light ochreous. Face shining light silvery ochreous;
head with light infuscation anteriorly; thorax and fore wing irro-
rated with blackish fuscous and shaded with fuscous; extreme base
whitish ochreous, with a single blackish fuscous spot in the fold and
followed by a rapidly fading fuscous shade; along costa and around
termen a series of blackish fuscous spots; at basal third, in cell, two
black discal spots, obliquely one above the other, followed by a fuscous
cloud; the latter is in turn followed by a blackish fuscous spot at end
of cell; cilia light grayish fuscous irrorated with wiiitish ochreous.
Hind wing grayish fuscous, cilia concolorous, with a slightly darker
basal band. Legs whitish ochreous ; fore and mid legs heavily over-
laid with fuscous except at joints; hind leg overlaid with fuscous on
femur with slight infuscation on tibia and tarsus. Abdomen fuscous
above; beneath whitish ochreous with broad, fuscous lateral line on
each side.

Male genitalia. — Harpe almost entirely clothed with fine hairs;
cucullus rounded, with several long, coarse marginal hairs;
clasper straight, reaching just beyond middle of harpe; anellus
a moderately sclerotized plate (with the posterior edge convex),
with large sparsely hairy lobes laterally. Vinculum rounded, with
dorsoanterior process well developed. Aedeagus stout, slightly bent
about middle, and bearing a flat, sclerotized basal process by which
the aedeagus articulates with the anellus. Transtilla a narrow, lightly
sclerotized band. Gnathos a heavily spined cone. Socii flat, hairy
lobes. Terminal portion of tegumen pointed.

Female genitalia. — Genital plate narrow. Ostium semicircular.
Ductus bursae membranous, becoming gradually larger to form the
symmetrical bursa copulatrix. Signum an oblong-oval, toothed plate.

Alar expanse, 17-24 mm.

Types. — In the United States National Museum.

Type localities. — Kaslo, British Columbia {pallidella) ; Silverton,
Colo, (terinella) ; Pullman, Wash, {serrae).

Food plant. — Senecio serra Hook.

Disfrihution. — ^Western United States and southwestern Canada.

United States records

California: Warner Mountains, 3 miles east of Davis Creek, Modoc County, alt.

5,500 feet, 2 $ $ and $ (VII-8 to 15-22, A. W. Lindsey).
Colorado: Silverton, 3 $ $ and 5 ( August 24-30 ) .
Utah: Stockton, 2 5 5 and 5 (VII-8-07, VII-30-13, VII-21-07, Tom Spalding) ;

Warner Ranger Station, La Sal Mountains, 9,000 feet, (July 1933, A. B.

Klots).

90 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Washington: Pullman, SI $ $ and 37 $ 2 (VI-15 to VII-3-1932-35, J. F. G.
Clarke [reared]).

Canadian records

British Columbia: Kaslo, $ (Dyar and Coclde) ; Keremeos, 3 $ $ (VI-30 to
VII-11-23, C. B. Garrett).

Remarks. — I do not hesitate to accept the above synonymy after
carefully studying the types of all three species. It is not surpris-
ing that the three species should have been described because of the
apparent differences. When carefully compared these differences are
reduced to mere shades of coloration. The specimens of teri/tiella are
somewhat darker than the type of pallideUa., and this is no doubt
due to the fact that they are in better condition than the rubbed type
of the latter species. A long reared series of serrae shows all but one
or two decidedly darker than either of the two types above, but this
depth of color is purely environmental. There are no differences
in the genitalia of either the males or females.

Under this name a long series of both reared and captured speci-
mens from British Columbia and California has previously been
placed. Although the genitalia of these specimens are similar to
pallidella they are amply different in other respects. They are de-
scribed on an earlier page of this paper as A. fusciterminella^ new
species.

AGONOPTERIX ARNICELLA (Walsingham)
Plate 33, Figtjee 186

Depressaria arnicella Walsingham, Proc. Zool. Soc. London, 1881, p. 313, pi. 36,
fig. 3. — Riley, in Smith, List of the Lepidoptera of Boreal America, No. 5254,
1891.— BusCK, Proc. U. S. Nat. Mus., vol. 24, p. 738, 1902 ; in Dyar, U. S. Nat.
Mus. Bull. 52, No. 5859, 1903. — Kearfott, in Smith, List of the Lepidoptera of
Boreal America, No. 6402 1903. — Anderson, Catalogue of British Columbia
Lepidoptera, No. 1089, 1904.— Kearfott, Can. Ent., vol. 37, p. 296, 1905.—
Metbick, in Wytsman, Genera insectorum, fasc. 180, p. 174, 1922. — Gaedb,
in Bryk, Lepidopterorum catalogus, pt. 92, p. 307, 1939.

Agonoptcryx arnicella (Walsingham) BuscK, Proc. U. S. Nat. Mus., vol 35, p.
198, 1908.

Agonopterix arnicella (Walsingham) McDunnough, Check list of the Lepidop-
tera of Canada and the United States of America (Part 2, Microlepidoptera )
No. 8414, 1939.

Agnoptcryx arnicella (Walsingham) Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6452, 1917.

Labial palpus pale whitish ochreous ; second segment heavily over-
laid with fuscous exteriorly and with a pink spot, about the middle
of brush, inwardly ; third segment with apex and subterminal annulus
black and a poorly defined subbasal annulus fuscous. Antenna dark
grayish fuscous. Face shining whitish. Head yellowish ochreous;
tuft beneath eye brilliant carmine. Thorax and fore wing pale whitish

REVISION OF THE OECOPHORIDAE — CLARKE 91

ochreous ; thorax suffused and irrorated with fuscous ; pale basal shade
of fore wing with black suffusion on extreme base of costa, a black
spot in fold and followed by a fuscous shade that does not reach costa ;
on the inner margin, at angle, an elongate black blotch in the fuscous
shade; along costa and around termen a series of blackish-fuscous
spots; apical third of costa roseate; at basal third two black discal
spots, obliquely one above the other, followed by a fuscous shade ; at
end of cell a white discal spot edged with black; cilia grayish fuscous
suffused with reddish. Hind wing shining gray; cilia concolorous,
with strong reddish suffusion. Legs whitish ochreous strongly over-
laid with fuscous except at joints; tip of tibia of first leg and the tips
of tibia and first tarsal segment of second leg, carmine ; tibia of hind
leg with strong reddish suffusion.

Male genitalia. — Harpe broad, moderately sclerotized, sparsely
clothed with fine hairs ; sacculus slightly emarginate at base of clasper,
otherwise parallel with costa; harpe not narrowing toward the broad,
rounded cucullus; clasper very short and slender, scarcely reaching
past the middle of the harpe. Anellus an oval plate constricted basally
and produced posteriorly, with hairy, lateral lobes. Vinculum
rounded. Transtilla a broad but lightly sclerotized band with small
narrow, lateral lobes. Gnathos an oval, spined knob. Socii mod-
erately clothed with fine hairs. Aedeagus slender, curved, pointed;
dilated slightly basally and with a well developed sclerotized arm by
which the aedeagus articulates with the anellus; vesica armed with
numerous fine, weak cornuti.

Alar expanse, 22 mm.

Type. — In the British Museum.

"^we locality. — Mount Shasta, Calif.

Food plants. — Arnica angusti folia (Vahl) (?) ; Erigeron sp. (Dr.
Braun's record).

Remarks. — ^I have received a male and a female of this species from
Dr. Annette F. Braun, of Cincinnati, Ohio. This is the first record of
the occurrence of this species since its discovery at Momit Shasta,
Calif., by Lord Walsingham in 1871. Dr. Braun's material was reared
from larvae collected at Hoh Lake, Olympic Mountains, Wash.,
August 13, 1936. The moths emerged September 2, 1936. This record
extends the known range of the species 1,000 miles northward.

Notes made in the field by Dr. Braun are as follows : "Larvae in the
top of Erigeron shoot, webbing together the uppermost leaves and
flowerbuds ; very dark purplish in color, with paler tubercles."

The food plant, as recorded by Dr. Braun, casts some doubt on
the correctness of Lord Walsingham's determination of the host of
ai^icella. Both food plants are Compositae, but I question Lord
Walsingham's record.

92 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

The female submitted by Dr. Braim is the first I have seen. Lord
Walsingham did not specify the sexes of the five specimens in his
type series, and, although he probably had a female or two, the one
before me is the only authentic one in this country. There are two
male paratypes from Lord Walsingham's series in the National col-
lection and one male in the Museum of Comparative Zoology at Cam-
bridge, Mass. I have compared the genitalia of one of the National
Museum paratypes with the genitalia of Dr. Braun's male and find
them identical. The male genitalia of arniceUa are unique in the
genus so there can be no doubt about the identity of the Washington
specimens.

I am gTeatly indebted to Dr. Braun for this interesting record and
also for the male specimen she has deposited in the National collection.

AGONOPTERIX ROBINIELLA (Packard)

Plate 29, Figures 169, 169a ; Plate 44, Figure 259

Depressaria rotlniella Packard, Guide to the study of insects, p. 349, pi. 8, fig. 14,
1869.— Chambers, Can. Ent., vol. 4, p. 91, 107, 1872 ; Cincinnati Quart. Journ.
Sci., vol. 1, p. 208, 1874.— Walsingham, Proc. Zool. Soc. London, 1881, p. 312.—
Riley, in Smith, List of the Lepidoptera of Boreal America, No. 5278, 1891. —
BuscK, Proc. U. S. Nat. Mus., vol. 24, p. 745, 1902 ; in Dyar, U. S. Nat. Mus.
Bull. 52, No. 5882, 1903.— Kearfott, in Smith, List of the Lepidoptera of
Boreal America, No 6425, 1903.— Traver, Psyche, vol. 26, p. 78, 1919.— Meyrick,
in Wytsman, Genera insectorum, fasc. 180, p. 176, 1922. — Gaede, in Bryk,
Lepidopterorum catalogus, pt. 92, p. 346, 1939.

Agonopteryx robiniella (Packard) BuscK, Proc. U. S. Nat. Mus., vol. 35, p. 199,
1908.— Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 241, 1923.

Agmiopterix ro'bimella (Packard) McDunnough, Check list of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera),
No. 8448, 1939.

Agnopteryx roMnieUa (Packard) Smith, Catalogue of the insects of New Jersey,
p. 561, 1910. — Barnes and McDunnough, Check list of the Lepidoptera of
Boreal America, No. 6471, 1917. — Brimley, The insects of North Carolina,
p. 304, 1938. — Procter, Biological survey of the Mount Desert region. Part 6,
The insect fauna, p. 273, 1938.

Depressaria hilarella Zellee [not Coquillett, Papilio, vol. 3, p. 98, 1883], Verh.
zool.-bot. Ges. Wien, vol. 23, p. 234, 1873.— Walsingham, Proc. Zool. Soc.
London, 1881, p. 312. — Smith, Catalogue of the insects of New Jersey, p. 355,
1890. — Riley, in Smith, List of the Lepidoptera of Boreal America, No. 5266,
1891. — Dietz, in Smith, Catalogue of the insects of New Jersey, p. 474, 1900.

Agonopterix hilarella (Zeller) McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera), No.
8448, 1939. (As synonym of roUniella (Packard) .)

Labial palpus whitish ochreous basally, gradually becoming more
yellowish toward the apex ; second segment mottled with brick red and
fuscous exteriorly and in the brush ; third segment with subbasal and
subterminal annuli brick red and fuscous, respectively, the former
redder, the latter more fuscous. Antenna fuscous with considerable

REVISION OF THE OECOPHORIDAE CLARKE 93

red scaling basally, especially above. Head, thorax, and <i;i-ound color
of fore wing yellow, mottled and overlaid with brick red and irrorated
and shaded with fuscons and black; at basal third two black cliscal
spots, the one nearer the inner margin less distinct than that toward
eosta ; discal spot at end of cell absent or very indistinct ; replaced by
a snbtriangular fuscous shade; before termen a poorly defined dark
band not attaining costa; along costa and around termen a series of
indistinct blackish spots; cilia light fuscous, tinged with red. Hind
wing grayish fuscous with terminal edge narrowly blackish fuscous;
cilia light fuscous with narrow subbasal and subterminal bands. Legs
whitish ochreous suffused and mottled with brick red and fuscous.
Abdomen whitish ochreous strongly overlaid with fuscous above; be-
neath sparsely irrorated with fuscous and black and with a row of
black spots on each side.

Male genitalia. — Harpe elongate, clothed with fine hairs, cucullus
pointed; clasper short, stout, straight. Anellus oval, slightly longer
tlian broad, posterior edge truncated; lateral lobes w^eak, clothed with
short hairs. Aecleagus stout, slightly curved ; vesica with an elongate
patch of fine, spiculate cornuti. Vinculum rounded. Transtilla a
broad sclerotized band with large lateral lobes. Gnathos an elongate,
oval, spined knob. Socii large, hairy, fleshy flaps.

Female genitalia. — Genital plate broad; area posterior to ostium
with two small sclerotized patches. Ostium oval, near center of plate.
Ductus bursae long, membranous, dilated at point of inception of
ductus seminalis. Bursa copulatrix with minute signum.

Alar expanse, IJ— 20 mm.

r^/z^e.— Lost?

Type locality. — ^Massachusetts ?

Food plants. — Hohinia psexidoacacia L. ; Saniculaf

Distribution. — Northeastern United States and eastern Canada.

TJyiited States records

Illinois: Oconee, $ (July 1-7).

Maine: Freedom, S (J- C. Purlin) ; Salisbury Cove (July 27; ace. Procter).

Maryland: Plummers Island, 2 S S (August 1903, A. Busck ; 2-VIII-20, H. S.
Barber).

Massachusetts: Manchester, $ (Beutenmiiller coll.) ; Marthas Vineyard, 5 S S ,
2 ? $ (July and August, F. M. Jones) ; Vineyard Haven, $ ("VII-13,"
F. M. Jones) ; Woods Hole {$, 19-VII-1919; $, l.WIII-1014, W. T. M.
Forbes ) .

New Hampshire: Hampton, 3 9 9 (VII-2-05, VII-8--08, S. A. Shaw).

New York: Bedford, 2 5 5 (ir)-VII-34, A. B. Klots) ; Ithaca (5 $ S , July and
August dates, W. T. M. Forbes ; S , 8-VIII-33, E. C. Hodson ; S , ll-VII-1928,
S, 22-VII-1928, A. B. Klots; 5, 27-VII-1929, A. G. Richards) ; Mattituck,
Long Island, 5 (4-VI-1933, Roy Latham) ; New Windsor, $ (June 1891) ;
Orient, Long Island, 8 $ S, 8 5 5 (April to October dates, 1933-1934, Roy
Litham) ; Rochester, 5 (31-VII-33, A. B. Klots) ; Southold, Long Island,
2 (August, Roy Latham).

94 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

North Carolina: Black Mountains, 2 ("July 4").
Ohio: Cincinnati, $, 5 ( VI 1-11-04, A. F. Braun).

Pennsylvania: New Brighton, $, 2 (VIII-9-07, VII-20-02, H. D. Merrick);
Oak Station, Allegheny County, $ (26-VI-1911, Fred Marloff).

Canadian records

Nova Scotia: Bridgetown, 2 (VIII-8-1912, "G. E. S.") ; Truro, 2 (25-VII-1913,
R. Matheson).

Ontario: Point Pelee, $, 2 (26-VII and ll-VII-1927, F. P. Ide [reared from
Rohinia pseudoacacla]) ; Port Hope, 2 2 2 (one, 24-V 1 1-1897 ; the other
without date or collector) ; Toronto, 2 (no date or collector) ; Trenton, $
(12-VII-1908, Evans) ; Vineland Station, 22 2 (5-VII-1038, W. L. Putnam
[reared from Rohinia pseudoacacia]).

Remarks. — This and the following three species have been confused
repeatedly in collections because of the difficulty of separating them
on superficial characters. The moths of all four intergrade to some
extent, but dimorpheUa, and less so lecontella, can be distinguished
rather readily. Under the respective species I have discussed their
separation from robinieUa.

I have described what I consider to be typical rohinielJa. It must
be borne in mind that there is considerable variation between speci-
mens and that many lack the red coloration found in tlie typical
form. The characters I have used in the key, and those I have dis-
cussed in my remarks under the various species, should suffice to
place any doubtful specimens.

AGONOPTERIX LECONTELLA (Clemens)

Plate 44, Figure 260

Depressaria lecontella Clemens, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 174. —
Robinson, Ann. Lye. Nat. Hist. New York, vol. 9, p. 157, pi. 1, fig. 9, 1870. —
Clemens, in Stainton, Tineina of North America, p. 137, 1872. — Chambers,
Can. Ent, vol. 4, p. 146, 1872; U. S. Geol. Geogr. Surv. Terr. Bull. 4, p.
138, 1878.— Walsingham, Proc. Zool. Soc. London, 1881, p. 312.— Riley,
in Smith, List of the Lepidoptera of Boreal America, No. 5268, 1891. —
BuscK, Proc. U. S. Nat. Mus., vol. 24, p. 745, 1902; in Dyar, U. S. Nat.
Mus Bull. 52, No. 5883, 1903. — Kearfott, in Smith, List of the Lepidoptera
of Boreal America, No. 6426, 1903. — Meyrick, in Wytsman, Genera insec-
torum, fasc. 180, p. 176, 1922. — Gaede, in Bryk, Lepidopterorum catalogus,
pt. 92, p. 330, 1939.

Ago^iopteryx lecontella (Clemens) Busck, Proc. U. S. Nat. Mus., vol. 35, p.
199, 1908.

Agnopteryx lecontella (Clemens) Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6472, 1917.

Agonopterix lecontella (Clemens) McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera), No.
8447, 1939.

REVISION OF THE OECOPHORIDAE — CLARKE 95

Labial palpus pale ochreous; second segment irrorated exteriorly
with fuscous ; third segment with subbasal and median bands fuscous.
Antenna fuscous. Head, thorax, and ground color of fore wing light
ochreous; the thorax, except collar, and fore wing, except base and
basal third of costa, strongly overlaid and suffused with brown ; fore
wing sparsely irrorated with black scales, more so in distal half; a
spot at extreme base of costa, one subcostally and another on inner
angle near base, black; at basal third two conspicuous, black discal
spots, one above the other, surrounded by a pale ochreous shade, the
latter followed by a poorly defined fuscous cloud ; at the end of cell
a poorly defined whitish-ochreous spot; along costa, around termen to
inner margin a series of small fuscous spots; cilia pale ochreous-
fuscous edged outwardly with pale ochreous. Hind wing grayish
fuscous ; cilia somewhat lighter with ochreous-fuscous terminal band.
Legs pale ochreous slightly irrorated and suffused with fuscous. Ab-
domen light ochreous, suffused with fuscous.

Female genitalia, — Genital plate moderately narrow with a strongly
sclerotized, slightly produced anterior edge. Ostium large, occupying
most of length of genital plate. Ductus bursae very long, gradually
tapering into the large asymmetrical bursa copulatrix. Inception of
ductus seminalis on right side of ductus bursae just before ostium.
Bursa bulged out to the right, the evaginated part annulated with a
series of concentric ridges ; the signum is a moderately large, strongly
sclerotized, toothed plate with well developed anterior and posterior
points and is situated at the periphery of the evaginated portion of
the bursa copulatrix.

Alar expanse, 18-25 mm.

Type. — In the Academy of Natural Science of Philadelphia,

Type locality. — "Pennsylvania."

Food plant. — Baptisia tinctoj^ia R. Brown.

Dis-tnhution. — Known only from the type locality (presumably
Pennsylvania) and one other specimen from Arendtsville, Pa. (7-4-29,
S.W.Frost).

Remarks. — In the National collection there is a single female of
this species. Aside from the type it is the only other specimen I
have seen. Everything else I have seen in the collections under this
name is referable to other species and is dealt with accordingly.

This species can be distinguished from the others with which it
has been confused by the pale area surrounding the two conspicuous
black discal spots at basal third, by the pale whitish-ochreous spot
at the end of the cell, and by the strong signum of the bursa.

This species is not included in the key based on gentalia.

96 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

AGONOPTERIX THELMAE, new species

Plate 44, Figtjke 259 A

Depressaria leeontella Smith [not Clemens], Catalogue of the insects of New

Jersey, p. 355, 1890. — Dietz, in Smith, Catalogue of the insects of New

Jersey, p. 474, 1900.
Agonopteryx leeontella Fokbes [not (Clemens)], Cornell Univ. Agr. Exp. Stat.

Memoir 68, p. 241, 1923.
Agnoptery.v leeontella Smith [not (Clemens)], Catalogue of the insects of New

Jersey, p. 561, 1910.

Labial palpus whitish ochreous ; second segment irrorated exteriorly
with reddish fuscous; third segment with blackish-fuscous subbasal
and supramedial bands. Antenna with basal segment whitish ochre-
ous indistinctly annulated with fuscous basally, finally becoming fus-
cous in apical third. Head, thorax, and ground color of fore wing
whitish ochreous. Thorax and fore wing irrorated with fuscous and
suffused with dull reddish ; beneath inner edge of tegula, on each side,
a fuscous spot ; extreme base of costa and a narrow line beyond light
basal area, fuscous; at basal third two superposed blackish-fuscous
spots ; a similar one at end of cell preceded by an orange-red streak ;
between this streak and the costa a small fuscous shade; along costa
and around termen a series of indistinct fuscous spots, those around
termen more reddish; cilia whitish ochreous with a reddish tint.
Hind wing fuscous ; cilia whitish ochreous banded with fuscous. Legs
whitish ochreous suffused and irrorated with reddish fuscous except
at joints. Abdomen whitish ochreous heavily overlaid with fuscous
above and with a row of black spots on each side beneath.

Male genitalia. — Similar to rohiniella but with slightly narrower
transtilla and with coarse, though small, cornuti.

Female genitalia. — Genital plate narrow, convexly produced an-
teriorly. Ostium large, broad, oval, about middle of genital plate.
Ductus bursae membranous except for a strongly dilated, lightly sclero-
tized area near posterior third; inception of ductus seminalis near os-
tium. Bursa copulatrix moderately large, without signum.

Alar expanse, 17-25 mm.

Type.—\J. S. N. M. No. 53175.

Type locality.— Oak Station, Allegheny County, Pa. (21-VIII-1910,
FredMarloff).

Food plant. — Unknown.

Bemarks. — Described from the $ type, 16 $ and 14 5 paratypes as
follows: 9 , "Connecticut"; S $ S , Plummers Island, Md. (July, Au-
gust, 1903, A. Busck; 2-Vn-1920, H. S. Barber) ; 9 Marthas Vine-
yard, Mass. ( September 1, F.M.Jones) ; $ , Ithaca, N. Y. (2-IX-1924,
W. T. M. Forbes) ; 2 , Lenox ([New York?] ; Beutenmiiller) ; 2 9 5
"New York"; S , Linwood Hill, N. Y. (14-VII-1888, H. G. Dyar) ; $ ,

REVISION OF THE OECOPHORIDAE — CLARKE 97

Sea Cliff, Long Island, N. Y. ("August") ; 12 5 $,9 Oak Station,
Allegheny County, Pa. (July and August dates, 1908-1911, Fred Mar-
loff) ; ? , Lobo, Ontario (29-VIII-1924, H. F. Hudson) ; 2 $ ? , Port
Hope, Ontario (17-Vni-1896, 15-VIII-1897, no collector); 2 9 9,
Toronto, Ontario (no date or collector) ; 2 $ ? , Montreal, Quebec
(lO-IX-1904, no collector).

Paratypes in the United States National Museum, Canadian Na-
tional, and Cornell University collections.

This species is difficult to separate, ^Yith certainty, from rohiniella
on superficial characters and on male genitalia. All the specimens I
have seen, however, have a distinct orange-red bar or dash in the cell
before the outer discal spot; all specimens of 7'ohm.iella I have seen
lack it. The females can be easily separated by genitalia. The gen-
ital plate of thelmae is narrow, that of rohiniella broad.

In addition to the type series I have seen 2 5 9 from Massachusetts
(Chilmark, "VIII-6'' George D. Eustis ; Vineyard Haven, VIII-10-36,
F. M. Jones) and 1 2 from Maine (October 12, 1938, A. E. Brower).

AGONOPTERIX DEWORPHELLA, new species

Plate 31, Figures 179, 179a ; Plate 40, Figure 229

A medium-sized species superficially resembling lecontella and
Tohinella but darker and smaller than either.

Labial palpus with second segment creamy white irrorated with
fuscous outwardly; third segment fuscous with each scale tip and
a median, inner fascia creamy white. Antenna with basal segment
and proximal fourth creamy white beneath and fuscous above; distal
three-fourths fuscous with narrow, lighter annulations. Head,
thorax, and fore wing reddish ochreous, suffused with fuscous.
From costa a fuscous median shade extending almost to inner mar-
gin; well before termen a similar but narrower, less conspicu-
ous outwardly curved shade; before middle of cell two small black
discal spots, one above the other ; at end of cell a conspicuous yellow
discal spot; cilia fuscous, irrorated with creamy white, and wnth a
distinct pink tinge; underside of fore wing blackish fuscous. Hind
wing blackish fuscous; cilia a lighter shade of the same color. Fore
leg with femur fuscous inwardly, creamy white outwardly; tibia
creamy white overlaid or irrorated with fuscous outwardly; tarsi
fuscous annulated with creamy white. Mid legs and hind legs creamy
white with tibiae fuscous outwardly; tarsi fuscous with creamy white
annulations. Abdomen fuscous above, creamy white beneath with a
black lateral line on each side.

Male genitalia. — Harpe moderately clothed with hairs; cucullus
pointed; clasper stout, short, scarcely exceeding middle of harpe,

286614 — 41 5 . .rv-^H

98 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

bluntly pointed; sacculus moderatelj^ sclerotized. Annellus longer
than broad, constricted basally; lateral edges convex; posterior edge
concave; lateral lobes poorly developed. Aedeagus stout, slightly
curved, bluntly pointed; vesica armed with a large patch of small
cornuti. Vinculum rounded, with prominent dorsoanterior process.
Transtilla a narrow sclerotized band with well developed, hairy
lateral lobes. Socii moderately large hairy lobes. Tegimien pointed.

Female genitalia. — Genital plate moderately sclerotized, with pro-
duced, truncate anterior margin. Ostium small, round. Ductus
bursae membranous ; inception of ductus seminalis well before ostium.
Signum absent.

Alar expanse, 11-18 mm.

Type.—\].^.'^M. No. 52947.

Type locality. — Henry, Putnam County, 111.

Food plant. — Amorplia fruticosa L.

Remarks. — Described from the $ type, 78 $ and 65 5 paratypes
all from the same locality. These are all reared specimens, bearing
May and June dates, from larvae collected by Murray O. Glenn.

Paratypes in the collections of Murray O. Glenn, Magnolia, 111.;
H. H. Keifer, Sacramento, Calif. ; Dr. Annette F. Braun, Cincinnati,
Ohio. ; Los Angeles Museum ; and Canadain National Museum.

In this species the males are strikingly darker and smaller and the
markings less contrasting than in the females.

I am indebted to Mr. Glenn for the long series of reared specimens
which make up the type series. Some of the specimens were reared
by Mr. Glenn while others were reared in Washington from
larvae submitted by him. The specimens reared at Washington are
identical with those reared in Illinois, although the emergence dates
are earlier for the former than for the latter.

Of this species Mr. Glenn writes, "The first emergence was June 4
(1938), and the heaviest from June 10-13. Pupation in the field
occurs immediately (after the larva matures) at the base of the
plant. In many instances this is the only place available as the
shrub is often completely surrounded by water, except for a small
hummock, about a foot in diameter, formed by the plant roots."

This species may be separated readily from roMniella or lecon-
tella by the yellow discal spot of the fore wing.

AGONOPTERIX ARGILLACEA (Walsingham)

Plate 29, Figures 171, 171a ; Plate 44, Figuee 255

Depressaria argillacea Walsingham, Proc. Zool. Soc. London, 1881, p. 313, pi.
36, fig. 2. — Riley, in Smith, List of the Lepidoptera of Boreal Amei'ica, No.
5252, 1891.— P.uscK, Proc. U. S. Nat. Mus., vol. 24, p. 788, 1902; in Dyar,
U. S. Nat. Mus. Bull. 52, No. 5860, 1903 ; Proc. U. S. Nat. Mus., vol. 27, p.

REVISION OF THE OECOPHORIDAE — CLARKE 99

763, 1904.— Anderson, Catalogue of British Columbia Lepidoptera, No. 1090,
1904. — Kearfott, in Smith, List of the Lepidoptera of Boreal America, No.
6403, 1903; Can. Ent., vol. 37, p. 296, 1905. — Meybick, in Wytsman, Genera
insectorum, fasc. 180, p. 174, 1922.— Gaede, in Bryk, Lepidopterorum cata-
logus, pt. 92, p. 306, 1939.

Agonopteryx argillacea (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35,
p. 199, 1908.- Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 238, 1923.

Agonopterix argillacea (Walsingham) McDunnough, Check list of the Lepidop-
tera of Canada and the United States of America (Part 2, Microlepidoptera),
No. 8413, 1939.

Agnoptcryx argillacea (Walsingham) Barnes and McDunnough, Check list of
the Lepidoptera of Boreal America, No. 6459, 1917.

Agonopteryx Uacella Barnes and Busck, Contr. Lepid. N. Amer., vol. 4, p. 232,
pi. 38, tig. 2, 1920. (New synonymy.)

Agonopterix Nacella (Barnes and Busck) McDunnough, Check list of the Lepi-
doptera of Canada and the United States of America (Part 2, Microlepidop-
tera), No. 8422, 1939.

Depressaria bJacella (Barnes and Busck) Meyrick, in Wytsman, Genera insec-
torum, fasc. 180, p. 174, 1922.

Labial palpus, antenna, head, thorax, and fore wing pale grayish
ochreoiis; labial palpus suffused and lightly irrorated exteriorly on
second segment with fuscous; third segment with subbasal and sub-
apical annuli (the former poorly defined) and tip blackish fuscous;
antenna darker apically; face lighter than rest of head; pale basal
patch of fore wing diffused along costa and bounded below it by an
outwardly diffused but inwardly distinct blackish-fuscous shade; re-
mainder of fore wing sparsely irrorated with black scales ; along ccsta
and around termen a series of indistinct fuscous spots ; at basal third, in
cell, two black discal spots (sometimes containing some brown scales)
obliquely one above the other, followed by a blackish fuscous cloud
above the middle of the wing and reaching a sordid-whitish discal
spot at the end of cell ; the spot at end of cell is narrowly edged with
blackish fuscous and brown scales and usually preceded by a similar,
brown-edged white spot; cilia concolorous with ground color of fore
wing. Hind wing and cilia pale grayish ochreous, the latter with
light-fuscous basal band. Legs pale grayish ochreous suffused with
blackish fuscous except at joints. Abdomen light grayish fuscous
above; beneath, pale grayish ochreous with a lateral row of black
spots on each side.

Male genitalia. — Harpe ample, moderately sclerotized and clothed
with hair; cucullus rounded; clasper slender, straight, reaching be-
yond center of harpe. Anellus broadly rectangular, somewhat nar-
rower basall}^, longer than broad and with weakly developed lateral
lobes. Vinculum rounded. Aedeagus moderately sclerotized; stout,
curved with apex pointed and sometimes slightly dorsally upturned.
Gnathos an elongate oval spined knob. Socii fleshy, hairy lobes.

100 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

Female genitalia. — Ostium spindle-shaped (transverse). Genital
plate moderately broad, produced anteriorly, the production of the
anterior margin as wide as the genital plate is long. Ductus bursae
membranous, variable in length; inception of ductus seminalis just
anterior to ostium. Bursa copulatrix not abruptly defined but
formed by the gradual broadening of the ductus bursae. Signum a
sclerotized, toothed, diamond-shaped plate.

Alar expanse, 19-24 mm.

Types. — In the British Museum {argillacea) ; in the United States
National Museum {hlacella).

Type localities. — Newville, Tehama County, Calif, {argillacea) ;
Shasta Eetreat and Truckee, Calif, (blacella).

Food plant. — Salix spp.

Distribution. — Western United States and western Canada and
northeastern United States and eastern Canada.

United States records

California: Sacramento, 11 $ $,2 9 9 (V-10 to 24-33, H. H. Keifer [reared]) ;

Truckee, 8 S $, 8 9 9 (August and October dates, Ximena McGlashan).
Idaho: Johnson's Bar, Snake River, 9 (IV-l()-26, J. F. G. Clarke).
Oreg-on: Fort Klamath, Fort Watson (Walsingham).
Pennsylvania: New Brighton, 9 (VI-20-07, H. D. Merrick).
Washington: Dieringer, 9 (VIII-1-32, W. W. Baker [reared]) ; Battleground,

Clark County, $ (VIII-22-30, J. F. G. Clarke) ; Logan Hill, Chehalis, $,

9 (II-5-30, III-28-30, T. M. Clarke) ; Pullman (4 $ $, S 9 9, March to

November dates, T. M. and J. F. G. Clarke [reared]).

Canadian records

British Columbia: Duncan, 9 ("1-10-12", Hanham) ; Vancouver, $ (IV-6-03) ;

Victoria, $, 9 ("7-9-03" ; "3-7-03") ; Wellington, 2 $ $ (IV-30-02), 9 9

("2-5-02"; "20-4-07") 4 S S (April, G. W. Taylor).
New Brunswick: Frederickton, 9 (Aug. 27, 1929, R. P. Gorham).
Ontario: Biscotasing, $ (VIII-4-31, Karl Schedl) ; Bobcaygeon, $ (VIII-16-32,

J. McDunnough) ; Stittsville, $ (21-VIII-1939, E. G. Lester).

Remarks. — The study of 22 male and female genitalia slides, to-
gether with host records, convinces me that the above synonymy is
correct.

A paratype of Walsingham's argillacea is before me. This speci-
men, in good condition, although somewhat faded, exactly matches
specimens of hlacelln. All the male genitalia are identical except
for slight variations in the harpe, a phenomenon frequently encoun-
tered in this gi'oup. The female genitalia show wide variation in the
length of the ductus bursae, but a study of Busck's cotypes of hlacella
show this wide variation in length even in the type series. The length
of the ductus bursae is not a reliable character for the separation of
species of this group.

REVISION OF THE OECOPPIORIDAE — CLARKE 101

I have before me long bred series from various localities, all show-
ing some variation. Specimens from Sacramento, Calif., match Wal-
singham's description of argillacea^ while other bred specimens from
Washington State match the types of hlacella. The pattern of both is
identical. The depth of color and the definition of the discal spots
vary throughout the series. These characters cannot be used for dif-
ferentiation of species since, in any long bred series, all degrees of
depth of color or definition of spots may be found.

AGONOPTERIX NIGRINOTELLA (Busck)

Plate 29, Figukes 170, 170a ; Plate 45, Figure 265

Depressaria nigrinotella Bxjsck, Proc. Ent. Soc. "Washington, vol. 9, p. 88, 1908. —

Meykick, in Wytsman, Genera insectorum, fasc. 180, p. 176, 1922. — Gaede, in

Bryk, Lepidopterorum catalogus, pt. 92, p. 334, 1939.
Aganopteryx nigrinotella Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198, IGOS. —

Barnes and Busck, Contr. Lepid. N. Amer., vol. 4, p. 232, 1920. — Forbes,

Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 240, 1923.
Agonopterix nigrinotella (Busck) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2. Microlepidoptera), No.

8444, 1939.
Agnopteryx nigrinotella (Busck) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6444, 1917.

Labial palpus pale yellowish brown ; second segment sparsely irro-
rated with blackish fuscous exteriorly; third segment with basal
three-fourths and apex black. Antenna fuscous. Head light reddish
brown. Thorax, base, and basal half of costa of fore wing light yel-
lowish brown; anterior edge of thorax and tegula somewhat darker
brown ; posterior tip of thorax blackish fuscous. Fore wing brownish
fuscous faintly irrorated with blackish fuscous; beyond light basal
part of wing a blackish-fuscous shade, which fades rapidly into the
lighter ground color ; at basal third, in cell a black discal dot followed
at end of cell by a yellowish- white discal spot ; cilia brownish fuscous,
more whitish at tornus. Hind wing shining light yellowish fuscous;
cilia somewhat lighter with pale fuscous subbasal and subterminal
bands. Legs pale yellowish brown, irrorated and suffused with black-
ish fuscous except at joints. Abdomen yellowish.

Male genitalia. — Harpe broad, pointed, clothed with long hairs;
clasper moderately sclerotized, slender, reaching almost to costa.
Anellus a rectangular plate longer than broad, with the posterior
edge truncated ; lateral lobes weak with only a few hairs. Vinculum
rounded with a well developed dorsoanterior process. Aedeagus ro-
bust, curved, pointed. Transtilla a narrow sclerotized band with
elongated lateral lobes. Gnathos an elongated cone armed with fine
spines. Socii moderate, clothed with many long hairs.

102 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Female genitalia. — Ostium large, round, near anterior margin of
genital plate ; anterior edge of genital plate produced, the production
nearly as wide as the plate is long. Ductus bursae membranous,
long ; ductus seminalis entering well before ostium. Bursa copulatrix
relatively small; signum a weakly sclerotized, oval, toothed plate.

Alar expanse, 20-23 mm.

Type. — In the United States National Museum.

Type locality. — Cincinnati, Ohio.

Food plants. — Xanthoxylum amsricanum Mill, and Oarya (sp. ?).

Distribution. — Eastern United States and Canada and probably
as far west as Texas in the South, following the distribution of the
food plant.

United States records

Ohio: Cincinnati, 2 S S , 4 $ 5 (June and July dates, A. F. Braun).

Canadian records
Ontario: St. Davids, 2 5 2 (VII-16-34, W. L. Putnam).

Remarks. — Busck^^ recorded this species from Ptelea trifoliata.,
but the food plants of nigrinotella are Xanthoxylum annericanum
and Carya (sp. ?). The species referred to by Mr. Busck is an
undescribed species, the description of which follows.

AGONOPTERIX COSTIMACULA, new species

Plate 30, Figures 174, 174a; Plate 45, Figure 261

Agonopteryx nigrinotella Barnes and Busck (not Busck), Contr. Lepid. North
America, vol. 4, p. 232, 1920.

Superficially much like nigrinotella but lighter, less immaculate,
and averaging larger with correspondingly wider wings.

Head light ochreous-f uscous ; face whitish ochreous; labial palpus
light ochreous-fuscous, lighter interiorly on second segment and
irrorated with blackish fuscous exteriorly; terminal segment with
blackish- fuscous subbasal and supramedial annuli and apex ; antenna
ochreous-fuscous, darker toward apex, narrowly annulated with
blackish fuscous. Thorax light ochreous-fuscous mixed with fuscous
anteriorly; ground color of fore wing ochreous-fuscous, the base and
costa to about middle slightly lighter; light basal area followed by
a blackish- fuscous shading; at basal third two obliquely placed black
discal spots, the upper, inner one large, the outer, lower one small;
at end of cell a light whitish-ochreous discal spot narrowly edged
with fuscous; on costa, six or eight conspicuous fuscous spots and a
series of smaller ones at ends of veins around termen; the whole

Busck, Contr. Lepid. Nortli America, vol. 4, p. 232, 1920.

REVISION OF THE OECOPHORIDAE — CLARKE 103

wing is irrorated with black scales ; cilia ochreous-f uscous with many
lighter tipped scales. Hind wing light smoky fuscous, lighter
basally, underside heavily shaded with black toward apex ; cilia light
fuscous with narrow, dark, subbasal band. Legs, except tarsi, yel-
lowish ochreous shaded with blackish fuscous; tarsi blackish fuscous
annulated with yellowish ochreous. Abdomen fuscous and whitish
ochreous above and whitish ochreous below with broad, black, lateral
lines.

Male genitalia. — Harpe long, moderately wide; cucullus pointed;
clasper long, stout, straight, reaching costa ; sacculus narrowly folded.
Anellus oval, truncated on posterior edge; lateral lobes weak with
few hairs. Vinculum rounded. Aedeagus long, curved, bluntly
pointed; vesica with large spinulate patch in basal half. Transtilla
a narrow sclerotized band with large, hairy, lateral lobes. Gnathos
an oval spined knob. Socii large hairy flaps.

Female genitalia. — Genital plate long; anterior edge folded nar-
rowly. Ostium near anterior margin of plate. Ductus bursae long,
membranous; inception of ductus seminalis well before ostium.
Bursa copulatrix large with small signum.

Alar expanse, 21-25 mm.

Type.—\]. S. N. M. No. 52080.

Type locality. — Plummers Island, Md.

Food plant. — Ptelea trifoliata L.

Remarks. — Described from the $ type and 51 $ and ? paratypes
as follows : 13 $ $ and 5 $ 9 , Plummers Island, Md. (March and
April dates, H. S. Barber and August Busck) ; Decatur, 111., $ and
2 9 5 (June 8-15, no collector) ; Cincinnati, $ and 2 5 5 (X-8-04,
X-8-07, VI-25-08, A. F. Braun) ; Clermont County, Ohio, 2 9 5
(VI-14-14, A. F. Braun) ; Point Pelee, Ontario, 13 5 5,8 55
(VII-15 to VIII-11-27, F. P. Ide; VII-30-31, G. S. Walley; VII-
27-31, W. J. Brown) ; Pelee Island, Ontario, 2 5 5 (VII-30-31, G. S.
Walley).

Paratypes in U. S. National Museum, Canadian National collection,
and collection of Dr. A. F. Braun, Cincinnati, Ohio.

This species has been mixed in collections, being placed vnider
eupatoHiella {=^ plummerella) ., argillacea., and nigrinotella. Ob-
scurely marked specimens of any of these species might easily be
confused. The long clasper which reaches the costa of harpe im-
mediately distinguishes the present species from the others.

I am indebted to Dr. J. McDunnough for a long, reared series from
Point Pelee, Ontario, which has greatly supplemented the material
here.

104 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

AGONOPTERIX CANADENSIS (Busck)

Plate 45, Figure 262

Depressaria canadensis Busck, Proc. U. S. Nat. Mus., vol. 24, p. 744, 1902 ;

in Dyar, U. S. Nat. Mus. Bull. 52, No. 5878, 1903.— Keaefott, in Smith,

List of the Lepidoptera of Boreal America, No. 6421, 1903.— Meyrick, in

Wytsman, Genera insectorum, fasc. ISO, p. 174, 1922. — Gaede, in Bryk,

Lepidopterorum catalogus, pt. 92, p. 311, 1939.
Agonopteryx canadensis Busck, Proc. U. S. Nat. Mus., vol. 35. p, 199, 1908. —

Forbes, Cornell Univ. Agr. Exp. Stat., Memoir 68, p. 240, 1923.
Agonopterix canadensis (Busck) McDunnough, Check list of the Lepidoptera

of Canada and the United States of America (Part 2, Microlepidoptera),

No. 8424, 1939.
Agnopteryx canadensis (Busck) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6467, 1917.

Labial palpus pale ochreous- white ; second segment evenly
springled with blackish fuscous exteriorly ; third segment with broad
subbasal and subapical annuli and tip black. Antenna fuscous with
indistinct black annulations. Head, thorax, and fore wing pale yel-
lowish gray; face creamy white; thorax with a few light-reddish
scales mixed ; pale base of fore wing diffused along costa, containing
a small black spot in fold and bounded below with a rapidly fading
blackish-fuscous shade ; remainder of fore wing irrorated with black-
ish-fuscous and black ; along costa and around termen to near middle
of inner margin a pronounced series of well-defined black spots; on
costa, in apical third of wing, and on basal half of inner margin a
rosy tint; at basal third a pair of rather large black discal spots,
obliquely one above the other and followed by a blackish-fuscous
shade above the middle of the wing ; at the end of cell a black discal
spot ; cilia yellowish gray, slightly darker than ground color of fore
wing. Hind wing light fuscous; cilia sordid whitish with yellowish
tint. Legs ochreous-white mottled and suffused with blackish fus-
cous except at joints.

Female genitalia. — Genital plate narrow, weakly sclerotized, pro-
duced anteriorly. Ostium occupying nearly all central portion of
genital plate ; inception of ductus seminalis well before ostium. Duc-
tus bursae membranous, very long, slender, gently tapering to form
the small bursa copulatrix. Signum of bursa a small, narrow, 4-
pointed plate, with anterior point longer than posterior.

Alar expanse, 17 mm.

Type. — In the United States National Museum.

Type locality. — ^^Yinnipeg, Manitoba (A. W. Hanham).

Remarks. — The female type is the only specimen of this species
I have seen. It is very distinct and should not be confused with
any other described North American species.

REVISION OF THE OECOPHORIDAE — CLARKE 105

Specimens from British Columbia and Ontario have been sent to
me under this name, but all these are referable to other species
{klamathiana or sciadopa) to which there is, in some cases, a super-
ficial resemblance. The genitalia of canculensis show no close re-
semblance to those of the other species, and canadensis is further dis-
tinguishable from them by lacking discal spots that thei others possess.

AGONOPTERIX FLAVICOMELLA (Engel)

Plate 30, Figures 173, 173a ; Plate 45, Figuee 267

Depressaria flaricomella Engel, Ent. News, vol. 18, p. 276, 1907. — Metriok, in

Wytsman, Genera insectorum, fasc. 180, p. 174, 1922. — Gaede, in Bryk,

Lepidopterorum catalogus, pt. 92, p. 324, 1939.
Agonopteryx flavicomella (Engel) Busck, Proc. U. S. Nat. Mus., vol. 35, p. 199,

1908.— Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 240, 1923.
Agonopterix flavicomella (Engel) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera ) , No.

8420, 1939.
Agnoptcryx flavicomella (Engel) Smith, Catalogue of the insects of New Jersey,

p. 561, 1910. — Barnes and McDunnough, Check list of the Lepidoptera of

Boreal America, No. 6463, 1917. — Brimi.et, The insects of North Carolina,

p. 304, 1938.

Labial palpus and head pale whitish ochreous; second segment of
palpus strongly irrorated and suffused with blackish fuscous exteriorly
and in brush ; third segment immaculate ; sides of head slightly darker.
Antenna grayish fuscous narrowly annulated with fuscous. Thorax
and basal part of fore wing (except costa) light ochreous; thorax
suffused and lightly irrorated with reddish brown and with a black-
ish-fuscous spot on each side beneath inner edges of tegulae. Ground
color of fore wing light ochreous heavily overlaid with reddish brown
and fuscous and irrorated with black; beyond the light basal patch
a dark blackish-fuscous shade, which rapidly fades to reddish, then
to ochreous at apical part of wing; basal part of costa suffused with
fuscous; costa and termen marked with a series of black spots, those
around termen more conspicuous than those on costa; at basal third
a conspicuous black discal spot followed by a similar but larger one
at middle and another larger more diffused one at end of cell; be-
tween the first and second discal spots and the inner margin (on vein
Ic) a somewhat diffused black spot; cilia fuscous, darker basally.
Hind wing grayish fuscous with a series of blackish- fuscous dashes
around apex and outer margin ; cilia yellowish fuscous with an indis-
tinct dark subbasal band. Legs light whitish ochreous strongly over-
laid or suffused with shining sooty black except at joints and on pos-
terior tibiae. Abdomen pale whitish ochreous irrorated with fuscous
above; ventrally with a broad blackish-fuscous longitudinal line on

106 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

each side and two indistinct median longitudinal rows of similarly
colored spots.

Male genitalia. — Harpe rather broad basally, tapering gently to the
bluntly pointed cucullus, profusely clothed with coarse hairs; costa
narrowly sclerotized ; sacculus moderately sclerotized ; clasper slender,
gently excurved, bluntly pointed, with small basal protuberance,
reaching well beyond center of harpe ; distal end serrate ; clasper and
inner edge of sacculus heavily sclerotized. Anellus broader than
long, convex laterally and slightly concave on posterior edge ; lateral
lobes large, sparsely hairy. Aedeagus stout, gently curved, pointed;
vesica armed with a large elongate patch of strong but moderately
small cornuti. Vinculum rounded with small dorsoanterior process.
Transtilla a narrow lightly sclerotized band with large, hairy, lateral
lobes. Gnathos rather long, slender. Tegumen pointed. Socii
large, hairy, fleshy flaps.

Female genitalia, — Genital plate moderately narrow; weakly
sclerotized around ostium with a broad convex anterior margin.
Ostium round, slightly nearer to anterior than posterior edge ; margin
narrowly sclerotized except posteriorly. Ductus bursae long, mem-
branous with entire inner surface studded with numerous minute
stout teeth anterior to inception of ductus seminalis ; ductus seminalis
well before ostium. Bursa copulatrix rather small, inner surface also
with numerous small teeth, but fewer than in ductus bursae ; signum
a moderately large irregularly shaped toothed plate.

Alar expanse, 15-17 mm.

Type. — In the United States National Museum.

Type locality. — New Brighton, Pa.

Food plant. — Heraclemn sp.

Distribution. — Eastern United States and Canada west to British
Columbia.

United States records

Illinois: Chicago, $ (no date or collector) ; Monee, 2 $ S (VI-21-12, A. Kwiat).
New Jersey: Essex County Park, 5 S $ (VII-2-^6, W. D. Kearfott).
Pennsylvania: New Brighton, 22 5 5 , 9 (June and July dates, H. D. Merrick) ;
Oak Station, Allegheny Coimty, 16 S o , 4 5 $ (June dates, Fred Marloff).

Canadian records

British Columbia: Rolla (5-VIII-1927, P. N. Vroom).

Manitoba: Riding Mountain Park, 2 $ S,S 9 9 (25-VI-33, J. McDunnough).

Ontario: Trenton (VII-22-12, Evans).

Saskatchewan: Indian Head (VII-22-25, J. J. de Gryse).

Remarks. — The abdomen of the type is missing. The genitalia are
figured from a "cotype" male and a typical female.

REVISION OF THE OECOPHORIDAE — CLARKE 107

AGONOPTEUIX SENICIONELLA (Busck)
Plate 30, Figukes 172, 172a ; Plate 45, Figures 263

Depressaria senicionella Busck, Proc. U. S. Nat. Mus., vol. 24, p. 742, 1902; in

Dyar, U. S. Nat. Mus. Bull. 52, No. 5875, 1903.— Ke^vrfott, in Smith, List of

the Lepidoptera of Boreal America, No. 6418, 1903.— Engel, Ent. News,

vol. 18, p. 276, 1907.
Depressaria seniciella Busck, Proc. U. S. Nat. Mus., vol. 24, p. 743, 1902.
Depressaria senecionella Meyeick, in Wytsman, Genera insectorum, fasc. 180, p.

174, 1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 350, 1939.
Affonopfcnjx senecionella Busck, Proc. U. S. Nat. jNIus., vol. 35, p. 199, 1908.—

Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 240, 1923.
Af/onoptrrix soiicioiiella (Busck) McDunnougii, Check list of the Lepidoptera

of Canada and the United States of America (Part 2, Microlepidoptera),

No. 8423, 1939.
Agnoptenjx senicionella (Busck) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6464, 1917.

Labial palpus light ochreous ; second segment irrorated with black-
ish fuscous exteriorly ; third segment with subbasal and subterminal
annuii and apex black. Antenna fuscous with grayish-ochreous an-
nulations. Head and thorax light ochreous; face yellowish white;
thorax with brownish-ochreous suffusion and irrorations. Fore wing
dark ochreous-gray, strongly overlaid and suffused with brownish
ochreous; light basal patch suffused on costa with fuscous, contain-
ing a blackish-fuscous spot in fold and bordered outwardly by a
rapidly fading fuscous shade ; at basal third, in cell, two black discal
dots, obliquely one above the other ; the lower one of these two is often
lacking ; at the end of cell an inconspicous black discal dot preceded
by an indistinct fuscous shade; termen and costa indistinctly marked
with poorly defined fuscous spots ; cilia ochreous-gray with two nar-
row ochreous-white lines, one at middle and one around outer edge.
Hind wing grayish fuscous, darker apically; cilia light yellowish
fuscous with grayish-fuscous subbasal and subterminal bands. Legs
light ochreous overlaid with fuscous except at joints. Abdomen gray-
ish ochreous above; beneath light ochreous with a broad blackish-
fuscous line on each side ; between these two lines a pair of indistinct
longitudinal rows of blackish-fuscous spots.

Male genHaUa. — Harpe rather broad, well sclerotized along costa;
clothed with fine hairs; cucullus rounded; clasper nearly straight,
modei'ately sclerotized and reaching slightly beyond middle of harpe.
Anellus oval, longer than broad, lateral lobes well developed, clothed
with few fine hairs. Vinculum narrow, rounded, with well-developed
dorsoanterior process. Aedeagus stout, slightly curved, bluntly
pointed ; ventral arm by which it is attached to anellus short, broad,
and situated near base of aedeagus. Transtilla a narrow sclerotized

108 PROCEEDINGS OF THE NATIONAL MUSEUM tol. 90

band with large, hairy, lateral lobes. Gnathos an oval spined knob.
Socii very large flaps clothed with many fine hairs. Tegumen pointed.

Female genitalia. — Genital plate narrow, strongly sclerotized and
produced anteriorly. Ostium round. Ductus bursae membranous,
gradually becoming larger to form the large oval bursa copulatrix.
Signum a large four-pointed plate, the anterior and posterior points
of about equal length but smaller than the lateral points.

Alar expanse, 18-22 mm.

Type. — In the United States National Museum.

Type locality. — Cabin John, Md.

Food plant. — Senecio aureus L.

Distribution. — Eastern United States.

United States records

District of Columbia: Washington, 8 5 (J, 9 (May 28 to June 16, 1900, August

Busck).
Maryland: Cabin John, 2 S S (IV-28 and 30-1900, August Busck) ; Plummers

Island, 3 5 5 , 11 $ $ (May and June dates, A. Busck and Chas. R. Ely).
Virginia: Great Falls, 9 (May 28, 1900, no collector),

AGONOPTERIX ANTENNARIELLA, new species

Plate 30, Figuees 175, 175a ; Plate 45, Fig-dbe 264
A medium-sized red-brown species unlike any other described from
North America.

Head dark reddish fuscous above mixed with carmine and whitish-
ochreous-tipped scales ; face shining whitish ochreous ; second segment
of palp)us whitish ochreous tinged with carmine inwardly and irrorated
with fuscous exteriorly; brush trumpet-shaped; terminal segment
whitish ochreous with broad blackish-fuscous basal and subterminal
annuli. Antenna fuscous, aimulated with whitish ochreous.

Thorax, base of fore wing, and costa to well beyond middle cine-
reous; anterior portion of thorax and extreme base of costa suffused
with rich brown ; ground color of fore wing red-brown, scales lightly
tipped with carmine ; at basal third two small obliquely placed black
discal spots followed by a few cinereous scales ; at end of cell a third
white discal spot edged with black; apical third of wing heavily
shaded with cinereous; cilia reddish fuscous, tipped with carmine; a
row of fuscous spots around tennen at base of cilia. Hind wing light
fuscous above ; with much white scaling beneath in apical third and a
black line at base of cilia ; cilia light fuscous with distinct rosy hue,
and darker subbasal band. Legs fuscous overlaid with whitish
ochreous and strongly suffused with carmine ; hind tibiae whitish, suf-

REVISION OF THE OECOPHORIDAE — CLARKE 109

fused with carmine; tarsi annulated with whitish ochreous. Abdo-
men grayish fuscous above, whitish ochreous below; ventrolateral
lines poorly defined, black; anal tuft yellowish ochreous, suffused
with carmine.

Mode genitalia. — Harpe moderately sclerotized and covered with
hairs over entire inner surface; costa and sacculus parallel beyond
middle then converging to form a narrowly rounded cucullus. Clasper
heavily sclerotized, smooth, short, terminating in a blunt point. Anel-
lus longer than broad, truncated on posterior edge; lateral lobes weak.
Vinculum with a very short, small dorsoanterior process. Transtilla a
very narrow, moderately sclerotized band with small lateral lobes.
Aedeagus stout, rather short and evenly tapered, terminating in a
sharp point. Armature consisting of many fine spinules. Gnathos a
heavily spined cone.

Female genitalia. — Genital plate moderately sclerotized, broad,
slightly produced anteriorly. Ostium near posterior margin of plate.
Ductus bursae slender, gradually becoming larger till it merges with
the bulbous bursa copulatrix. Signum a 4-pointed, moderately sclero-
tized small plate with strong teeth; lateral, anterior, and posterior
points about equal in length.

Alar expanse, 17-24 mm.

Type.— U. S. N. M. No. 52081.

Type locality. — Kamiack Butte, Whitman County, Wash.

Food plant. — Antennaria luzuloldes T. and G.

Remarks. — Described from the $ type, 7 $ and 4 $ paratypes, as
follows: Kamiack Butte, 3 5 3,25$ (V-18 to 22-34) ; 4 5 5 , 1 9
same (VI-15 to 19-35) ; $ , Pullman, Wash. (1-28-31 [indoor
record] ) ; $ , Godman Springs, Blue Mountains, Wash., 6,000 feet
(VIII-1-35) ; all reared from larvae collected by the author.

Paratypes in collections of United States National Museum, Cana-
dian National Museum, and H. H. Keifer, Sacramento, Calif.

Dr. J. McDunnough, of Ottawa, has sent me three specimens of
this species from British Columbia. The larvae, from which these
moths were reared, were collected by A. N. Gartrell at Shingle
Creek Road, Keremeos. The adults emerged from 30-VI to
&-VII-35.

Mr. Gartrell has labeled his specimens as reared from wild red
currant. The food plant record is rather surprising since most of
the species of this group are host specific, and it will be noted that the
type series was reared from Antennaria. I have carefully compared
the genitalia of specimens of the two lots and find them identical.

110 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

AGONOPTERIX NEBULOSA (Zeller)

Plate 32, Figuees 185, 185a ; Plate 45, Figueb 266

Dcpressaria nehulosa Zeller, Verb, zool.-bot. Ges. Wien, vol. 23, p. 237, 1873.— i
Chambers, U. S. Geol. Geogr. Surv. Terr. Bull. 4, p. 138, 1878.— Walsing-
HAM, Proc. Zool Soc. London, 1881, p. 312.— Riley, in Smith, List of the
Lepidoptera of Boreal America, No. 5269, 1891.— Busck, Proc. U. S. Nat.
Mus., vol. 24, p. 741, 1902 ; in Dyar, U. S. Nat. Mus. Bull. 52, No. 5870,
1903. — Kearfott, in Smith, List of the Lepidoptera of Boreal America, No.
6413, 1903. — Meyeick, in Wytsman, Genera insectorum, fasc. 180, p. 177,
1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 334, 1939.

AgonopteryoR nebulosa (Zeller) Busck, Proc. U. S. Nat. Mus.. vol. 35, p. 198,
1908.— Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68 p. 239, 1923.

Agonopterix nclulosa (Zeller) McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera),
No. 8452, 1939.

Agnopteryx nehulosa (Zeller) Smith, Catalogue of the insects of New Jersey,
p. 516, 1910. — Barnes and McDunnough, Check list of the Lepidoptera of
Boreal America, No. 6438, 1917.

Labial palpus shining creamy white ; second segment sprinkled with
black scales outwardly and with brush suffused with white-tipped
fuscous scales; brush abruptly expanded, trumpet-shaped; third seg-
ment with broad subbasal and subapical annuli black; the black scales
of these annuli white tipped. Antenna fuscous annulated with gray.
Head, thorax, and fore wing lavender-gray, the scales white tipped;
face creamy white, tegula with a bluish iridescence; along costa and
around termen a series of poorly defined light fuscous spots ; at basal
third two black cliscal dots, the upper, inner one small, the outer,
lower one dashlike and both bordered outwardly with white scales;
at the end of cell a white black-edged discal spot preceded by a similar
one; sometimes these two are confluent, forming a white and black
longitudinal dash; cilia grayish fuscous with narrow median and
terminal white lines. Hind wing and cilia light yellowish fuscous, the
wing darker apically ; cilia with several, somewhat darker bands. Legs
ochreous-white, overlaid and irrorated with grayish fuscous above,
ochreous-white beneath; the usual longitudinal lines or rows of dark
spots lacking or only faintly indicated by a few blackish-fuscous scales.

Male genitalia. — Harpe moderately broad, entirely clothed with fine
hairs; cucullus narrowly rounded; clasper stout, curved, slightly
hooked, reaching two-thirds of distance to costa. Anellus broadly
oval with poorly developed lateral lobes. Vinculum rounded. Aedea-
gus short, stout, straight, with a blunt point; vesica armed with
numerous spinulate cornuti. Transtilla a very narrow sclerotized
band with moderately large, hairy, lateral lobes. Gnathos an oval,
spined knob. Socii hairy flaps.

REVISION OF THE OECOPHORIDAE — CLARKE 111

Female genitalia. — Genital plate narrow, emarginate behind ; ante-
rior margin in the form of a narrow, sclerotized, curved bar; ostium
occupying nearly whole ventral portion of genital plate and without
sclerotized area posterior to it. Ductus bursae membranous, abruptly
constricted before ostium. Bursa copulatrix oval ; signum an elongate
plate with posterior pointed process.

Alar expanse, 19-20 mm.

Type. — In the British Museum.

Type locality. — Cambridge, Mass.

Food plant. — Unknown.

Distrihution. — Northeastern United States.

United States records

Maine: Monmouth, S ( "November 30, '05" ) .

Massachusetts: Amherst, 2 S S (January 21, 1906).

New Hampshire: 2 $S without further data; Hampton, $, 2 (III-26-05,

III-22-05, S. A. Shaw).
New York: New Windsor, $ (June 24, 1903, E. L. Morton).

Remarks. — Differs from all other gray American species by the
accentuated trumpet-shaped palpus.

AGONOPTERIX SABULELLA (Walsingham)

Plate 32, Figotes 181, 181a ; Plate 46, Figure 270

Depressarla sabulella Walsingham, Proc. Zool. Soc. London, 1881, p. 313, pi. 36,
fig. 1. — Riley, in Smith, List of the Lepidoptera of Boreal America, No. 5279,
1891.— BusCK, Proc. U. S. Nat. Mus., vol. 24, p. 743, 1902 ; in Dyar, U. S. Nat.
Mus. Bull. 52, No. 5876, 1903. — Keaetott, in Smith, List of the Lepidoptera
of Boreal America, No. 6419, 1903 ; Can. Ent. vol. 37, p. 296, 1905.— Meyeick, in
Wytsman, Genera insectorum, fasc. 180, p. 174, 1922. — Gaede, in Bryk, Lepi-
dopterorum catalogus, pt. 92, p. 348, 1939.

Agonopteryx sabulella (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35, p.
199, 1908.

Agonopterix sabulella (Walsingham) McDunxough, Check list of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera ) , No.
8412, 1939.

Agnoptenjx sabulella (Walsingham) Barnes and McDunnougii, Check list of
the Lepidoptera of Boreal America, No. 6465, 1917.

Agonopteryx callosella Barnes and Busck, Contr. Lepid. North America, vol. 4,
p. 231, pi. 38, fig. 4, 1920. (New synonymy).

Agonopterix callosella (Barnes and Busck) McDunnough, Check list of the
Lepidoptera of Canada and the United States of America (Part 2, Micro-
lepidoptera) No. 8421, 1939.

Depressarla callosella (Barnes and Busck) Meyrick, in Wytsman, Genera insec-
torum, fasc. 180, p. 174, 1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt.
92, p. 311, 1939.

Labial palpus pale ochreous; second segment with slight fuscous
shading and blackish-fuscous irrorations exteriorly; third segment

112 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

with subapical aniiulus and apex blackish fuscous. Antenna blackish
fuscous. Head, thorax, and fore wing pale ochreous; face ochreous-
white; base of costa and spot in anal angle blackish fuscous; forewing
sparsely irrorated with fuscous and shaded with reddish scales; at
basal third two fuscous discal spots obliquely one above the other;
at end of cell a fuscous spot preceded above by a less conspicuous one
of the same color; between the outer and inner pairs of spots a poorly
defined fuscous cloud in costal half of wing; apical half of costa and
termen and the cilia with a reddish suffusion, cilia light ochreous.
Hind wing grayish ochreous, cilia paler, with alternating light fus-
cous and whitish bands. Legs ochreous-white strongly suffused and
overlaid with blackish fuscous except at joints; posterior tibia and
tarsus hardly more than irrorated with fuscous. Abdomen grayish
fuscous above, pale ochreous beneath ; on each side ventrally, a black
longitudinal line.

Male genitalia. — Harpe narrow, long, pointed, clothed with very
fine hairs; clasper stout, slightly curved toward cucuUus, and termi-
nating in a knob, Anellus a roughly oval plate, longer than broad,
truncated posteriorly, and with well developed, hairy, lateral lobes.
Vinculum rounded. Aedeagus stout with a long slender upturned
point and a large, spinulate patch in basal half. Transtilla a nar-
row, lightly sclerotized band with well-developed, hairy, lateral lobes.
Gnathos a spiny cone. Socii broad flaps clothed with fine hairs.

Female genitalia. — Ovipositor naked except for a few strong long
hairs at base and a few weak hairs distally. Genital plate moderately
broad, slightly produced anteriorly. Ostium is posterior half of
plate. Ductus bursae membranous, slender, widening abruptly to
form the large oval bursa copulatrix. Signum a diamond-shaped
plate, with the anterior point shorter than the posterior.

Alar expanse, 19-23 mm.

Types. — In the British Museum {sahulella) ; in the United States
National Museum {callosella).

Type localities. — Mendocino County, Calif, {sabulella) ; San Ber-
nardino, Calif, {callosella).

Food plants. — Eriophyllum stachaedi folium Lag. and Eriophyllum
lantanum typicum Constance,

Distribution. — ^Western United States probably as far north as
southern British Columbia in the interior.

United States records

California: Land's End, San Francisco, $ (VII-23^09, F. X. Williams) ; San
Francisco, $ (VI-12-26, H. H. Keifer) ; Monachee Meadows, Tulare
County, 8,000 feet, $ ("July 8-14") ; Croville, S (IV-14-28, H. H. Keifer) ;
San Bernardino, 9 ("June 1-7") ; San Diego, $, 5 ("May 24-30").

Idaho: Kamiah, $ (VI-27-35, L. Constance).

REVISION OF THE OECOPHORIDAE — CLARKE 113

R&marks. — The above synonymy is based on a comparison of Wal-
singham's figure and description with the types of callosella.
Although there is no authentic material of Walsingham's species
here, there appears to be no doubt about the synonymy. Mr. Busck
concurs with me in this opinion.

H. H. Keifer, of Sacramento, Calif., was kind enough to send me
three specimens, which I associate with this species, two of which
were reared by him from ErioyhyUmn stachaedi folium. These speci-
mens are identical with the types of callosella^ which are before me.

I have another specimen before me from Kamiah, Idaho, reared
from Eriophyllwm lantanum typicwm^ which was collected by my
friend Dr. Lincoln Constance, of the Department of Botany, Uni-
versity of California. This specimen is close to the types of callosella
but differs in details. I am placing it here, however, until more
material can be obtained and its specific limits ascertained.

AGONOPTERIX COSTOSA (Haworth)

Plate 32, Figtxres 184, 184a ; Plate 46, Figure 273

Depressaria costosa Hawobth, Lepidoptera Britarmica, vol. 3. p. 508, 1811. —
Wood, Index entomologicus, p. 172, pi. 38, fig. 1182, 1845. — Stainton, Trans.
Ent. Soc. London, vol. 5, p. 153, 1849 ; Insecta Britannica, p. 84, 1S54. — Zeu^er,
Linn. Ent., vol. 9, p. 198, 1854. — Stainton, A manual of British butterflies and
moths, vol. 2 p. 320, 1859. — Fbey, Die Tineen und Pterophoren der Schweiz, p.
82, 1856. — Heinemann, Die Schmetterlinge Deutschlauds und der Schweiz,
vol. 2, p. 143, 1870.— Meykick, Ent. Monthly Mag., vol. 13, p. 281, 1870.—
Fkey, Die Lepidopterea der Schweiz,. p. 351, 1880. — Rossleb, Jahrb. nassau.
Vereins Naturk., vol. 33, p. 282, 1881. — Snellen, Die Vlinders van Nederland,
vol. 2, p. 577, 1882. — Steudetl and Hofmann, Wiirttemberg. Vereins vaterl.
Naturk., vol. 38, p. 196, 1882. — Joubdheuiixe, Mem. Soc. Acad. I'Aube, vol. 47,
p. 176, 1883. — Sorhagen, Die Kleinschmetterlinge der Mark Brandenburg, p.
175, 1SS6.— Meyrick, A handbook of British Lepidoptera, p. 620, 1895.—
Reutti, Verb. Naturw. Vereins Karlsruhe, vol. 12, p. 228, 1898. — Stangk, Die
Tineinen der umgegend von Friedland in Mecklenberg, p. 20, 1899. — Stau-
dinger and Re:bel, Catalog der Lepidopteren des palaearctischen Faunenge-
bietes, vol. 2, No. 3177, 1901.— Car.\dja, Bull. Soc. Sei. Bucarest, vol. 10, p. 149,
1901.— Malloch, Ent. Monthly Mag., vol. 37, p. 186, 1901.— Disque, Deutsche
Ent. 'Zeitschr., Iris, vol. 14, p. 205, 1901. — Longstaff, Ent. Monthly Mag., vol
38, p. 28, 1902.— ScHtJTZE, Deutsche Ent. Zeitschr., Iris, vol. 15, p. 12, 1902.—
Crombbugghe, Mem. Soc. Ent. Belgique. vol. 14, p. 43, 1906. — Gibbs, Eutomolo
gist, vol. 39, p. 7, 1906 [as D. costana, nom. nud.]. — MUlleb-Rutz, Bull. Soc,
Ent. Suisse, vol. 11, p. 346, 1909.— Spulee, Schmetterlinge Europas, vol. 2, p,
333, pi. 89, fig. 13, 1910.— GiANEixi, Ann. Accad. Agr. Torino, vol. 53, p. 94,
1910. — SCHIULE, Krakow. Akad. Umiejetnosci Sprawozdania Komisyi fizyo
graficznej, vol. 45, p. 29, 1911.— Sk.u.a, Int. Ent. Zeitschr., vol. 5, 303, 1912
Verb, naturf. Vereins Bi-unn, vol. 51, p. 312, 1913. — Vorbbodt and Muller-
Rutz, Schmetterlinge der Schweiz, vol. 2, p. 456, 1914. — Buxton, Ent. Rec,
vol. 27, p. 183, 1915.— Galvagni, Wien. Ent. Vereins Jahresb., vol. 25, p. 32,
1915.— Martini, Deutsche Ent. Zeitschr., Iris. vol. 30, p. 153, 1917.— Resjel,
286614—41 6

114 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Sitzungsb. Akad. Wiss. Wien, vol. 126, p. 808, 1917.— Galvagni, Wien. Ent.
Vereins Jahresb., vol. 28, p. 63, 1918. — Strand, Archiv fiir Natnrg. vol. 85A,
pt. 4, p. 9, 1919. — Meyrick, Entomologist, vol. 55, p. 254, 1922; in Wytsman,
Genera inseetorum, fasc. 180, p. 173, 1922. — Zimmerman, Verb, zool.-bot.
Ges. Wien. vol. 71, p. (43), 1922. — Preissecker, Wien. Ent. Vereins Jahresb.,
vol. 30, p. 187, 1924.— Blair, Entomologist, vol. 58, p. 10, 1925.— Stephan,
Deutsche Ent. Zeitschr., Iris, vol. 39, p. 123, 1925.— Larsen, Ent. Meddel., vol.
17, p. 73, 1927.— Zeirny, Eos, vol. 3, p. 480, 1927.— Wickham, Entomologist,
vol. 60, p. 43, 1927. — Meyrick, A handbook of British Lepidoptera, p. 683,
1928. — Hayward, Entomologist, vol. 62, p. 50, 1929. — Lhomme, L'amateur de
papillons, vol. 4, p. 209, 1929. — Uffeln, Westfalischer provinzial-Verein fiir
Wissenschaft und Kunst, Miinster, Abhandl., vol. 1, p. 76, 1930. — Amsel,
Deutsche Ent. Zeitschr., Iris, vol. 44, p. 121, 1930. — Ve3ibrodt, Deutsche Ent.
Zeitschr., Iris, vol. 45, p. 126, 1931.— Chater, Bull. Ent. Res., vol. 22, p. 231,
1931. — Rebel and Zerny, Denkschr. Akad. Wiss. Wien, math.-nat. Kl., vol.
103, p. 150, 1931. — Hering, in Brohmer, Ehrmann, and Ulmer, Die Tierwelt
Mitteleuropas, vol. 1, p. 142, 1932. — Eckstein, Die Kleinschmetterlinge
Deutschlands, p. 118, pi. 5, fig. 213, 1933. — Sterneck, Prodromus der Schmet-
terlingsfauna Bohmens, vol. 2, p. 105, 1933.— Rapp. Beitrage zur fauna
Thiiriugens, vol. 2, p. 131, 1936; Beitrage Schmetterlingsfauna Harz, p. 28,
1936. — Gaede, in Bryk, Lepidopterorum catalogus pt. 92, p. 318, 1939.

Agonopterix costosa (Ha worth) Pierce and Metcalfe, The genitalia of the
tineid families of the Lepidoptera of the British Islands, p. 35, pi. 19, 1935.—
McDuNNouGH, Check list of the Lepidoptera of Canada and the United
States of America (Part 2, Microlepidoptera ) , No. 8410, 1939.

Agonopteryx blacktnori Busck, Can. Ent. vol. 53, p. 277, 1921.

Depressaria Nackmori (Busck) Meyrick, Entomologist, vol. 55, p. 254, 1922. (As
synonym of D. costosa (Haworth).)

Depressaria dryadoxena Meyrick, Exotic Microlepidoptera, vol. 2, p. 315, 1920;
Entomologist vol. 55, p. 254, 1922 ; in Wytsman, Genera inseetorum, fasc. 180,
p. 173, 1922. (As synonym of D. costosa (Haworth).)

Agonopterix dryadoxena (Meyrick) McDunnough, Check list of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera),
No. 8410, 1939. (As synonym of D. costosa (Haworth).)

Agonopterix hlackmori (Busck) McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera), No.
8410, 1939. (As synonym of D. costosa (Haworth).)

Tinea depunctella Hxjbner, Sammlung Europaisher Schmetterlinge, vol. 5, pi.
56, fig. 378, 1816.

Pinaris depunctella (Hiibner) Hubner, Verzeichniss bekannter Schmetterlinge,
p. 411, 1825.

Depressaria depunctella (Hiibner) Koli.ar, Linz. Oberosterreicher musealverein
Jahresb., vol. 2, p. 92, 1832.— Zeller, Isis von Oken, 1839, p. 195.— Duponchel,
Historie naturelle de lepidopteres . . . de France, vol. 8, p. 148, pi. 291, fig. 2,
1838. — Hekrich-Schaffer, Die Schmetterlinge von Europa, vol. 5, p. 123, 1853.

Haemylis depunctella (Hiibner) Teeitschke, in Ochsenheimer, Die Schmetter-
linge von Europa, vol. 9, p. 260, 1832 ; vol. 10, p. 185, 1835.

Haemylis costosa (Haworth) Stephens, Illustrations of British entomology,
Haustellata, vol. 4, p. 203, 1835.

Labial palpus, head and face creamy white; second segment irro-
rated with blackish fuscous exteriorly; third segment with subapical

REVISION OF THE OECOPHORIDAE — CLARKE 115

aniiulus and apex black; head frequently suffused above with fuscous;
spot above base of antenna brown; tuft below eye pink. Antenna
brown, darker basally than apically. Thorax and ground color of
fore wing whitish ochreous; posterior tuft on thorax yellowish ochre-
ous; light base of fore wing diffused along costa to apex, sparsely irro-
rated with black scales; beyond light base a dark ochreous shade,
which covers posterior two-thirds of wing but becomes lighter toward
termen. The whole surface of wing more or less irrorated with black
scales ; on costa, slightly beyond middle a sooty cloud ; at basal third,
two black spots obliquely one above the other sometimes edged by a
few dark reddish-ochreous scales ; between the outer of these two spots
and inner margin a small dark reddish-ochreous spot; at the end of
cell a cream-colored discal spot edged with dark reddish ochreous and
preceded by a dark reddish-ochreous spot. Sometimes the latter
spot and the reddish-ochreous scales of the outer discal spot are con-
fluent, forming a dash ; cilia whitish ochreous strongly suffused with
reddish fuscous, especially toward apex. Hind wing light yellowish
fuscous; cilia yellowish, banded with light fuscous. Legs whitish
ochreous overlaid with fuscous and tinged with carmine. Abdomen
light ochrex)Us above, whitish ochreous beneath; a row of black spots
on each side ventrolaterally.

Male genitalia. — Harpe long, slender, tapering into a long, pointed
cucullus ; moderately covered with hairs ; clasper very stout, somewhat
dilated distally, nearly reaching costa ; sacculus short, strongly sclero-
tized. Anellus broader basally than apically, broader than long; pos-
terior edge convex; hairy, lateral lobes well developed. Aedeagus
stout, slightly narrower in middle, nearly straight, pointed; vesica
armed with a large patch of spinulate cornuti. Vinculum rounded.
Transtilla a narrow sclerotized band with large hairy lateral lobes.
Gnathos a spined oval knob. Socii hairy, fleshy flaps.

Female genitalia. — Genital plate moderately broad, sclerotized ; an-
terior edge slightly produced. Ostium round; situated about middle
and occupying less than half the length of the genital plate. Ductus
bursae membranous; inception of ductus seminalis just before ostium.
Signum distinctly 4-pointed, with slender anterior and posterior points ;
teeth small and scattered.

Alar expanse, 18-22 mm.

Type.—\\\ the British Museum.

Type locality. — "Europe."

Food plants. — Genista, Qu-ercus (f), Lahiuvmm, and Cytisus scopa-
rius (L.) Link.

Distrihution.—'ExtvQmQ northwestern United States, southwestern
and eastern Canada and Europe.

116 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

United States records

Washington: Bellingham, 3 $ S, 9 (IX-1-29, VIII-4-31, IX-7-35, VIII-2a-33,
J. F. G. Clarke) ; Tacoma, 9 (VII-15-2S, T. M. Clarke).

Canadian records

British Columbia: Victoria, 19 5 (J, 9 9 5 (July to October dates, E. H. Black-
more, W. Downes, W. R. Carter collectors) ;2 $ S, 9 (9-15-VII-1923, K. F.
Auden).

Ontario: London, $, $ ( 19-26-IV-1933 [Good's Greenhouse]).

Note. — ^Well established in the Puget Sound region and no doubt
wider spread than the above records indicate. May not have escaped
at London, Ontario.

Remarks. — There appears to be no doubt of the synonymy as given
above. Ahhough I have not seen drijadoxena^ Meyrick's description
tallies with costosa and the locality for the former species agrees with
that of hlackmori. The food plant of dryadoxena {Quercus) is ques-
tionable, since the species of this genus rarely feed on such widely
separated species of plants and are more frequently host specific as
has already been pointed out.

The genitalia of the males and females are identical except for
slight variation in the male harpes, a condition frequently encountered.
Pierce ^^ figures the genitalia of costosa showing the ostium opening
near the anterior margin of the genital plate. In the specimens I have
seen (4), both European and North American, the ostium opens
slightly nearer to the posterior than to the anterior edge (see fig. 273).
Furthermore, the signum as figured by Pierce lacks the anterior and
posterior points, present in all specimens I have examined. The sig-
num, it must be admitted, may be somewhat variable, but in this
species it appears to be unusually constant.

AGONOPTERIX PERGANDEELLA (Busck)

Plate 32, Figures 183, 183a

Depressaria pergandeeUa Busck, Proc. Ent. Soc. "Washington, vol. 9. p. 89, 1908. —

Meybick, in Wytsman, Genera insectorum, fasc. 180, p. 174, 1922. — Gaede,

in Bryk, Lepidopterorum catalogus, pt. 92, p. 339, 1939.
Agonopteryx pergandeeUa Busck, Proc. U. S. Nat Mus., vol. 35, p. 199, 1908.
Agnopteryx pergandiella Baenes and McDunnough, Check list of the Lepi-

doptera of Boreal America, No. 6462, 1917.
Agonopterix pergandiella McDunnough, Check list of the Lepidoptera of Canada

and the United States of America (Part 2, Microlepidoptera), No. 8417, 1939.

Labial palpus ochreous- white ; second segment sparsely irrorated
with fuscous exteriorly, brush suffused with brown [third segment of
both palpi missing]. Antenna with basal segment fuscous above,
whitish beneath [remainder of both antennae missing]. Head

22 Pierce, F. N., The genitalia of the Tineina, pi. 19, 1935.

REVISION OF THE OECOPHORIDAE — CLARKE 117

ochreous with a median fuscous band; face shining creamy-white.
Thorax and fore wing light brown; thorax strongly suffused with
brownish fuscous ; surface of fore wing irrorated with poorly defined
blackish-fuscous spots and with a series of similarly colored spots
around termen ; at basal third, in cell, two black discal spots obliquely
one above the other ; another similar spot at the end of cell preceded
by a poorly defined blackish-fuscous cloud; extreme base of costa
fuscous; a well-defined black spot in inner angle; cilia light brown
suffused with fuscous. Hind wing shining light yellowish fuscous;
cilia more whitish. Legs whitish ochreous overlaid and mottled with
fuscous except at joints. Abdomen dark yellowish fuscous.

Male genitalia. — Harpe moderately clothed with long hairs [cucuUus
broken from each harpe] ; sacculus only moderately sclerotized ;
clasper nearly straight, slightly dilated distally, reaching fully two-
thirds of the distance toward the costa. Anellus broadly oval,
posterior edge strongly convex, smooth ; with very small lateral lobes.
Vinculum roimded. Aedeagus nearly straight, stout, and deeply
notched distally. Transtilla a broad, sclerotized band with large
lateral lobes. Gnathos a finely spined cone. Socii small, weak,
sparsely clothed with hairs. Tegumen produced into a moderately
large broad flap.

Alar expanse, 21 mm.

Type. — In the United States National Museum.

Type locality. — "Nebraska."

ReTiiarks. — The only specimen I have seen is the type male. This
is very distinct from any other North American species.

AGONOPTERIX AMISSELLA (Busck)

Plate 33, Figubes 187, lS7a ; Plate 46, Figure 269

Depressaria amissella Busck, Proc. Ent. Soc. Washington., vol. 9, p. 89, 1908. —

Meykick, in Wytsman, Genera insectorum, fasc. 180, p. 174, 1922.— Gaede,

m Bryk, Lepidopterorum catalogus, pt. 92, p. 300, 1939.
Agonoptcryx amissella Busck, Proc. U. S. Nat. Mus., vol. 35, p. 190, 1908.
Agonopterix amissella (Busck) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera), No.

8425, 1939.
Agnopteryx amissella (Busck) Babnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6460, 1917.

Labial palpus ochreous- white ; second segment with the brush suf-
fused with fuscous and sparsely irrorated with fuscous exteriorly;
third segment with subbasal and subterminal annuli black. Antenna
fuscous with light ochreous- fuscous annulations; basal segment
ochreous- white beneath. Head, thorax, and fore wing light ochreous-
brown; face shining ochreous- white ; thorax suffused with fuscous;
surface of fore wine: irrorated with black and blackish-fuscous scales

118 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

and with a series of blackish-fuscous spots along costa and around
termen; extreme base of costa fuscous; light basal patch bordered out-
wardly by a transverse fuscous dash, which contains some jet-black
scales and does not reach costa; at basal third, in cell, two conspicuous
jet-black spots of raised scales, obliquely one above the other and fol-
lowed outwardly by a poorly defined fuscous shade ; discal spot at end
of cell inconspicuous or absent, but when present indicated by a few
whitish scales; cilia light ochreous-brown the scales tipped with
ochreous-white. Hind wing light fuscous, darker apically than
basally; cilia light yellowish fuscous tipped with ochreous-white and
with a narrow, fuscous subbasal band. Legs ochreous-white strongly
suffused and overlaid with blackish fuscous except at joints.
Abdomen ochreous-fuscous.

Male genitalia. — Harpe rather narrow, pointed, only sparsely
clothed with hairs ; clasper very stout, straight, reaching three-fourths
of the distance to costa. Anellus roughly rectangular with trun-
cated posterior edge; lateral lobes small. Vinculum rounded.
Transtilla a narrow sclerotized band with large lateral lobes.
Aedeagus stout, curved, with a heavily sclerotized band basally;
vesica armed with a large patch of weak, spinulate cornuti. Gnathos
an elongate, oval knob clothed with fine spines. Socii weakly sclero-
tized, small, with few hairs. Terminal portion of tegumen pointed.

Female genitalia. — Genital plate narrow ; anterior edge in the form
of a strongly sclerotized crescentic bar. Ostium broad, oval, with a
small, strongly sclerotized area posterior to it. Ductus bursae mem-
branous. Bursa copulatrix small; signum of bursa a small, oval,
spined plate.

Alar expanse, 17 mm.

Type. — In the United States National Museum.

Type locality. — Kissimmee, Fla. (Wm. Beutenmiiller) .

Remai'hs. — The only specimens I have seen of this species are the
type male and a paratype female. The species is quite distinct from
any other North American species.

AGONOPTERIX LATIPALPELLA Barnes and Busck

Plate 33, Figuees 188, 188a ; Plate 46, Figxjbe 268

Agonopteryx latipalpella Baenes and Busck, Contr. Lepid. North America, vol.

4, p. 233, 1920.
Agonopterix latipalpella (Barnes and Busck) McDunnoxjgh, Check list of the

Lepidoptera of Canada and the United States of America (Part 2, Micro-

lepidoptera). No. 8432, 1939.
Depressaria latipalpella Meyrick, in Wytsman, Genera insectorum, fasc. 180,

p. 176, 1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 330,

1939.

The entire insect presents a roughened appearance due to the raised
scales that cover almost the whole surface. Labial palpus ochreous-

REVISION OF THE OECOPHORIDAE — CLARKE 119

white; second segment irrorated with black and fuscous exteriorly
and suffused with fuscous in the brush; third segment with broad,
poorly defined blackish-fuscous basal and median annulations.
Head, thorax, and fore wing light ochreous-brown, the scales suffused
with fuscous and tipped with sordid whitish; collar narrowly black;
base somewhat lighter than the rest of wing, the light color diffused
for a short distance along costa and bordered outwardly by a strong
black shade, the latter from the middle to inner margin of wing; at
basal third an inconspicuous white discal spot (sometimes two) pre-
ceded by a few black scales, at the end of a cell a small white discal
spot broadly edged with black; cilia light ochreous-brown with a
narrow median fuscous band. Hind wing ochreous-fuscous, darker
around margins; cilia ochreous with narrow fuscous subbasal and
subterminal bands. Legs ochreous-white suffused and mottled with
blackish fuscous and fuscous. Abdomen ochreous-fuscous edged lat-
erally with numerous light ochreous-brown hairlike scales; beneath,
two broad, black, longitudinal lines broken into spots in posterior
half; between these two lines two indistinct parallel rows of spots.

Male genitalia. — Harpe elongate, moderately narrow; sparsely
clothed with hairs; cucullus pointed; clasper long, slender, slightly
enlarged in middle; sacculus broad. Anellus rectangular; posterior
edge emarginate ; lateral lobes well developed, hairy. Aedeagus stout,
slightly bent, pointed ; vesica with a weak spinulate patch of cornuti
in basal half. Vinculum rounded. Transtilla a narrow sclerotized
band, with large hairy, lateral lobes. Gnathos a spined, oval knob.
Socii hairy flaps.

Female gemtalia. — Genital plate broad, with a gently concave, nar-
row, sclerotized anterior edge. Ostium small, round, situated on pos-
terior margin of genital plate. Ductus bursae sclerotized behind incep-
tion of ductus seminalis, short, membranous before, gradually widen-
ing into the very large bursa copulatrix ; signum large, 3-pointed, with
a few large teeth on posterior margin.

Alar expanse, 17-19 mm.

Type. — In the United States National Museum.

Type locality. — San Benito, Tex.

Remarks. — The species is represented only by the type series from
Brownsville and San Benito, Tex.

AGONOPTERIX POSTICELLA (Walsingham)

Plate 33, Figures 180, 190a ; Plate 46, Ficuee 274

Depressaria posticeUa Walsingham, Proc. Zool. Soc. London, 1881, p. 315, pi. 36,
fig. 5. — Riley, in Smith, List of the Lepidoptera of Boreal America, No.
5274, 1891.— BuscK, Proc. U. S. Nat. Mus., vol. 24, p. 744, 1902 ; in Dyar, U. S.
Nat. Mus. Bull. 52, No. 5880, 1903.— Kearfott, in Smith. List of the Lepidop-
tera of Boreal America, No. 6423, 1903. — MEYracK, in Wytsman, Genera in-

120 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

sectorum, fasc. 180, p. 173, 1922. — Gab3)e, in Bryk, Lepidopterorum catalogus,

pt. 92, p. 340, 1939.
Agonoptcryx posticella (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35, p.

190, 1908.
Agonopterix posticella (Walsingham) McDunnough, Check list of the Lepidop-

tera of Canada and the United States of America (Part 2, Microlepidoptera ) ,

No. 8411, 1939.
Agnopteryx posticella (Walsingham) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6469, 1917.

Labial palpus dull whitish ochreous ; second segment irrorated with
fuscous exteriorly ; third segment with apex fuscous and, in a few speci-
mens, a faint indication of a fuscous subterminal annulus. Antenna
fuscous. Head, thorax, and fore wing dull whitish ochreous; thorax
with slight inf uscation ; fore wing irrorated with black and fuscous and
suffused with reddish especially toward apex ; on costa at base and on
inner angle near base a blackish-fuscous spot; at basal third a black
discal spot followed by a similar one at end of cell, the latter frequently
obsolete ; on inner margin, before tornus a fuscous blotch not attaining
costal half; cilia fuscous with reddish tinge. Hind wing light grayish
fuscous; cilia lighter with rosy tinge. Legs pale whitish ochreous
heavily overlaid with fuscous except at joints and on posterior tibiae,
the latter with faint rosy tinge. Abdomen pale ochreous, suffused with
f uscuous above and with a row of more or less confluent spots on each
side beneath ; anal tuft usually with rosy tint.

Male genitalia. — Harpe slender, almost entirely clothed with long
hairs; clasper reaching beyond costa, slightly narrowed about the
middle and somewhat enlarged at the distal end, terminating in a
sharp point. Anellus a sclerotized plate, posterior edge concave, and
with weak lateral lobes. Vinculum rounded, broad, with well de-
veloped dorsoanterior process. Aedeagus slender, slightly curved,
gradually tapering to a sharp, dorsally upturned point; at the base
is a bifid sclerotized arm by which the aedeagus articulates with the
anellus. Transtilla a narrow band, with well developed, hairy, lat-
eral lobes. Gnathos a long, spined cone. Socii rather small, clothed
with fine hairs.

Female genitalia. — Genital plate broad, with pronounced anterior
median ventral evagination. Ostium large, oval, in posterior half
of genital plate. Ductus bursae membranous, long; inception of
ductus seminalis well before ostium. Bursa copulatrix large, oval;
signum a small, oval plate, with serrate edges and a pointed process
from the posterior margin.

Alar expanse, 17-21 mm.

Type. — In the British Museum.

Type locality. — Lake County, Calif.

REVISION OF THE OECOPHORIDAE — CLARKE 121

Food, plants. — Psoralea physodes Dougl., P. macrostachya DC,
and P. tenuiflora Pursh.
Distribution. — Western United States.

United States records

Arizona: Yavapai County, $ (no date).

California: Fresno, $ (no date; E. A. Schwarz) ; Sacramento, S $ $ (V-11-S3),

and Applegate, Placer County, $ (VII-14-32), all reared and sent to me

by H. H. Keifer.
Colorado: Boulder, 2 ("May 14, Cockerell") ; other specimens (Dyar and

Caudell, labeled "Colorado"); Chimney Gulch, Denver, $ (6-13).
Oreg-on: McMinuville, $ (VII-9-22, S. E. Keen) ; Salem, 9 ("March," E. Y.

Lansing Jr.).

Remarks. — This far-western species seems to be confined to the
area south of the Cohimbia River. A diligent search in the spring
of 1934 by W. W. Baker, of Puyallup, Wash., failed to reveal the
presence of this species in Washington State. Mr. Baker did, how-
ever, locate larvae of psoraliella (referred to in this paper), which
is found feeding with posticella in California. The food plant {P.
physodes) is rather widely distributed on the Pacific slope west of
the Cascades in both Washington and British Columbia, so the
species may eventually be found in both places.

AGONOPTERIX PSORALIELLA (Walsingham)

Plate 31, Figxjres 177, 177a ; Plate 46, Figure 271

Depressaria psoraliella Walsingham, Proc. Zool. Soc. London, 1881, p. 317,
pi. 36, fig. 7. — Riley, in Smith, List of Lepidoptera of Boreal America, No.
527.5, 1891.— BuscK, Proc. U. S. Nat. Mus., vol. 24, p. 740, 1902 ; in Dyar,
U. S. Nat. Mus, Bull. 52, No. 5865, 1903.— Kearfott, m Smith, List of the
Lepidoptera of Boreal America, No. 6408, 1903. — Meyrick, in Wytsman,
Genera insectorum, fasc. 180, p. 176, 1922. — Gaede, in Bryk, Lepidopterorum
catalogus, pt. 92, p. 342, 1939.

Agonoptcryx psoraliella (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35, p.
198, 1908.

Agonopterix psoraliella (Walsingham) McDunnough, Check list of the Lepidop-
tera of Canada and the United States of America (Part 2, Microlepidop-
tera). No. 8449, 1939.

Agnopteryx psoraliella (Walsingham) Barnes and McDunnough, Check list of
the Lepidoptera of Boreal America, No. 6442, 1917.

Labial palpus pale gray; second segment strongly suffused with
fuscous and dull red exteriorly; third segment slightly tinged with
red and with subbasal and subapical annuli and apex black. An-
tenna fuscous with reddish tint. Head gray, lightly suffused with
fuscous and tinted with red laterally; face shining white. Thorax,
base of wing, and basal fourth of costa gray irrorated with deep
red-brown and black (thorax in one specimen wholly red-brown) ;

122 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

posterior tuft of thorax light red, beneath the inner edge of each
tegula a small black spot ; fore wing deep red -brown irrorated with
blackish fuscous and gray, especially toward costa and along veins;
at basal third two small yellowish discal dots, one above the other,
followed by another at the end of cell ; frequently there is a fourth,
though smaller spot between the outer and inner discal spots; all
spots edged with deep red; along costa and around termen a series
of indistinct blackish-fuscous spots; cilia fuscous with a reddish
tinge. Hind wing dark grayish-fuscous ; cilia lighter, reddish tinged
with a well-defined fuscous basal band and several less distinct sub-
terminal ones. Legs gray overlaid with blackish fuscous, except at
joints, and tinted with red. Abdomen gray, suffused with fuscous
above ; beneath, lightly suffused with red, irrorated with fuscous and
with a row of black spots on each side.

Male genitalia. — Harpe broad and short (when compared with
those of other members of the genus) , clothed over most of the sur-
face with fine hairs; cucullus rounded (in one specimen from Cali-
fornia the cucullus is long and slender) ; clasper long and slender,
reaching to, or slightly beyond, the costa; occasionally the clasper
is slightly hooked at the distal end. Anellus an oblong-oval plate;
posterior edge nearly straight with shallow median cleft; lateral lobes
well developed. Vinculum rounded, with dorsoanterior process.
Aedeagus slender, tapering distally to a point just past the middle,
then becoming larger and finally terminating in a sharp point. At
the base of the aedeagus is a bifid arm by which it articulates with
the anellus. Transtilla a broad sclerotized band, with large, hairy,
lateral lobes. Gnathos a slender spined cone. Socii large hairy
flaps.

Female genitalia. — Genital plate broad. Ostium small, at ex-
treme anterior margin of plate. Ductus bursae long, membranous;
inception on ductus seminalis well before ostium. Bursa copu-
latrix scarcely larger than the ductus. Signum a lightly sclerotized,
oval plate.

Alar expanse, 20-24 mm.

Type. — ^In the British Museum.

Type locality. — Sonoma County, Calif.

Food plants. — Psoralea physodes Dougl. and Psoralea macro-
stachya DC.

Distrihution. — The Pacific slope of the United States.

United States records

California: Pope Creek, Napa County, ,^,2 5 5 (V-27 to V-30-32, H. H. Keifer
[reared] ; Mills College, Alameda County (1 specimen in Dr. Braun's col-
lection) ; Dry Creek, Sonoma County, $ (V-21-1S71, Walsingliam [reared]).

Washington: Puyallup, 4 S $, 5 9 $ (V-7 to 17-34, W. W. Baker [reared]).

REVISION OF THE OECOPHORIDAE — CLARKE 123

Remarks. — The species is easily recognizable from Walsingham's
description and figure. There is one specimen from Washington State
that has the head and thorax dark reddish brown instead of the usual
gray.

AGONOPTERIX SANGUINELLA (Busck)

Plate 33, Figures 189, 189a

Depressaria sanguineUa Btjsck, Proc. U. S. Nat. Mus., vol. 24, p. 738, 1902. —
in Dyar, U. S. Nat. Mus. Bull. 52, No. 5861, 1903.— Kearfott, in Smith, List
of the Lepidoptera of Boreal America, No. 6404, 1903. — IMeyrick, in Wyts-
man. Genera insectorum, fasc. 180 p. 176, 1922. — Gaedb. in Bryk, Lepidopter-
orum catalogue, pt. 92, p. 348, 1939.

Agonoptenjx sanguineUa Busck, Proc. U. S. Nat. Mus., vol. 35, p. 199, 1908.

Agonopterix sangiiinella (Busck) McDunnough, Check list of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera),
No. 8445, 1939.

Agnoptenjx sangui^icUa (Busck) Baenes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6458, 1917.

Labial palpus pale grayish ochreous; second segment sparsely ir-
rorated with black exteriorly; third segment with a small spot near
base anteriorly, and apex black. Antenna dark fuscous. Face, head,
and collar of thorax light straw color, the scales of the head with a
grayish suffusion. Thorax and fore wing gray, sparsely irrorated
with black and with a carmine tint, the carmine color more pronounced
along costa and apex of fore wing; extreme base of wing and costa
pale grayish ochreous, the former followed by and the latter narrowly
edged with black; at basal third two obliquely placed black discal
spots edged with a few carmine scales; at end of cell a white discal
spot conspicuously edged w4th carmine; from the discal spot at end
of cell an inwardly oblique, blackish dash not reaching costa; cilia
gray with reddish suffusion and sparse black irrorations; termen with-
out row of dark spots. Hind wing light ochreous fuscous ; cilia con-
colorous with subbasal and two jDarallel fuscous bands and with whit-
ish irrorations apically. Legs pale grayish ochreous irrorated and
suffused with blackish fuscous except at joints. Abdomen pale
grayish ochreous irrorated with black and with a row of black spots on
each side beneath.

Male genitalia. — Harpe slender, clothed with long, fine hairs ; cucul-
lus pointed; clasper short, straight, bluntly pointed; sacculus mod-
erately sclerotized. Anellus slightly longer than broad, v/ith small
lateral lobes; posterior margin with shallow cleft; basal portion con-
stricted. Aedeagus stout, slightly curved, bluntly pointed; vesica
armed with numerous strong spinulate cornuti. Vinculum rounded.
Transtilla a sclerotized band with well-developed lateral lobes.
Gnathos a spined knob. Socii fleshy, hairy flaps. Tegumen termi-
nating in a blunt point.

124 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Alar expanse, 21 mm.

Ty^e. — In the United States National Museum.

Type locality. — Pinal Mountains, Ariz. (R. Kunze).

Remarks. — I have seen no specimens certainly referable to this
species, although I have before me a series from White Mountains,
Ariz., which may belong here. The specimens from this locality aver-
age considerably larger than the type of sanguinella but may be well
within the range of variation for this species.

AGONOPTERIX AMYRISELLA (Busck)

Plate 46, Figure 272

Depressaria amyrisella Busck, Proc. U. S. Nat. Mus., vol. 23, p. 233, 1901. — Dyab,
Proc. Ent. Soc. Washington, vol. 4, p. 476, 1901. — Busck, Proc. U. S. Nat. Mus.,
vol. 24, p. 741, 1902 ; in Dyar, U. S. Nat. Mus. Bull. 52, No. 5872, 1903.— Keab-
FOTT, in Smithy List of the Lepidoptera of Boreal America, No. 6415, 1903. —
Meybick, in Wytsman, Genera insectorum, fasc. 180, p. 175, 1922. — Gaede, in
Bryk, Lepidopterorum catalogus, pt. 92, p. 300, 1939.

Agonopteryx amyrisella Busck, Proc. U. S. Nat. Mus., vol. 35, p. 198, 1906.

Agonopterix amyrisella (Busck) McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera ) , No.
8427, 1939.

Agnopteryx amyrisella (Busck) Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6446, 1917.

Labial palpus yellowish white ; second segment with blackish-fuscous
base and with black irrorations ; third segment with apical third black,
subbasal fourth light reddish fuscous ; the whole palpus with a reddish
tinge. Antenna dark metallic greenish brown. Face yellowish with a
few brown scales. Head with erect scales, yellowish at bases, purplish
black toward tips, tips white with reddish tinge. Thorax yellowish
brown with violaceous scales intermixed and with a transverse crest of
six tufts of scales. Fore wing dark violaceous-brown sparsely irro-
rated with black scales ; extreme base yellowish, the color not extend-
ing to costa, and containing a black spot near inner angle ; beyond this
light basal patch a purplish-black fascia not attaining costa ; at basal
third, in cell, a few scattered purplish-black scales; at end of cell a
small white discal spot broadly edged with black ; on costa five rather
large blackish spots, the fourth (from base of wing) the largest, ex-
tending into cell and confluent with the black-edged white discal spot ;
around termen six to eight small black spots; cilia yellowish brown
with a narrow fuscous subbasal line and tinged with reddish. Hind
wing yellowish brown, much darker apically; cilia light yellowish
fuscous with a broad fuscous subbasal band. Legs pale yellowish
brown irrorated and suffused with purplish black except at joints.
Abdomen pale yellowish brown infuscated above and with a row of
black spots on each side beneath.

REVISION OF THE OECOPHORIDAE CLARKE 125

Female genitalia. — Genital plate moderately sclerotized; anterior
edge lobed. Ostium small, round, opening about middle of plate.
Ductus bursae short, not much longer than bursa copulatrix; incep-
tion of ductus seminalis just before ostium. Bursa copulatrix large,
oval, without signum.

Alar expanse, 16-17 mm.

Type. — In the United States National Museum.

Type locality. — Palm Beach, Fla.

Food plant. — Ainyris -jioridoma Nutt.

Remarks. — This species is represented only by the type series.
These are females.

2. MARTYRHILDA, new genus

Plate 2, Figure 17 ; Plate 6, Figuee 44 ; Plate 10, Figures 67, 67a ; Plate 16,

Figure 100

Genotype. — Depressaria canella Busck, Proc. U. S. Nat. Mus., vol.
27, p. 764, 1904.

Similar to Agonopterix but palpus more slender, smooth, brush on
second segment poorly developed or absent. Fore wing narrow,
elongate, bluntly pointed; costa straight. Hind wing with costa
straight or slightly excavate. Abdominal wall strongly sclerotized.

Male genitalia. — Clasper divided, with transverse arm and longi-
tudinal arm, the former sometimes with lateral process from inside;
gnathos broadened, sometimes renifonu; vesica usually armed with
strong cornuti.

Female genitalia. — Signum always large, broadly oval to elongate,
never diamond-shaped. Ductus bursae membranous or partly sclero-
tized, sometimes with strong thornlike processes from inner surface.

Larva. — ^As in Agonopterix.

Pupa. — Pubescent. Prothoracic femora exposed. Labial palpi not
exposed. Cremaster absent.

Remarks. — ^This genus is closely related to Agonopterix., differing
from it by the poor development or absence of the brush of the second
segment of the labial palpus, the divided clasper of the harpe, the
unusually large broadly oval or elongate signum, and the exposed
prothoracic femora of the pupa.

With the genotype I associate ten other species, three of which I
have described as new.

The larvae of only three species of this genus are known. The larva
of sphaeralceae is a leaf miner ; those of umhraticostella and canella
are leaf tiers.

126 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

KEY TO THE SPECIES OF MARTYRHILDA BASED ON
COLORATION

1. Ground color of fore wing white or whitish 2

Ground color of fore wing otherwise 4

2. Costa with a large fuscous blotch at middle canella (Busck) (p. 128)

Costa without large dark markings 3

3. Basal segment of antenna sordid whitish; hind wing smoky

fuscous sordidella, new species (p. 132)

Basal segment of antenna fuscous; hind wing white, nivalis (Braun) (p. 139)

4. Base of fore wing dark reddish brown or blackish fuscous 5

Base of fore wing otherwise 7

5. Ground color of fore wing straw yellow gracilis (Walsingham) (p. 133)

Ground color of fore wing otherwise 6

6. Ground color of fore wing light reddish ochreous

umbraticostella (Walsingham) (p. 130)
Ground color of fore wing pale ochreous-gray.

thoracenigraeella (Chambers) (p. 135)

7. Costa, to about middle, lighter than general color of wing 8

Costa not contrastingly lighter than remainder of wing 10

8. Third segment of labial palpus immaculate., hildaella, new species (p. 140)
Third segment of labial palpus maculate 9

9. Fore wing suffused or marked with reddish or brownish; alar

expanse 20 mm. or more klamathiana (Walsingham) (p. 142)

Fore wing without reddish or brownish suffusions or markings;

alar expanse 19 mm. or less sciadopa (Meyrick) (p. 144)

10. Ground color of fore wing grayish fuscous strongly overlaid with

whitish ochreous sphaeralceae, new species (p. 138)

Ground color of fore wing brownish fuscous sparsely irrorated

with whitish ochreous thoracefasciella (Chambers) (p. 136)

KEY TO THE SPECIES OF MARTYRHILDA BASED ON MALE
GENITALIA

1. Gnathos reniform (figs. 100, 143) 2

Gnathos not reniform (fig. 149) 4

2. Transverse arm of clasper extending beyond middle of harpe (figs.

67, 143) 3

Transverse arm of clasper short, stout, not extending beyond

middle of harpe (fig. 145) sordidella, new species (p. 132)

3. Harpe slender; both arms of clasper of about equal length; trans-

verse arm reaching costa of harpe (fig. 143)

umbraticostella (Walsingham) (p. 130)
Harpe short, broad; transverse arm of clasper longer than longi-
tudinal arm but not attaining costal edge of harpe (fig. 67)

canella (Busck) (p. 128)

4. Lobes of transtilla fused (figs. 146, 147) 5

Lobes of transtilla not fused (figs. 148, 149, etc.) 6

5. Sacculus as broad as one-third width of harpe at base; transverse

and longitudinal arms of clasper of about equal length;
aedeagus about one-half length of harpe (fig. 146)

thoracefasciella (Chambers) (p. 136)

REVISION OF THE OECOPHORIDAE — CLARKE 127

Sacculus broader than one-third width of harpe at base; trans-
verse arm of clasper appreciably longer than longitudinal
arm; aedeagus longer than one-half length of harpe (fig. 147)

sphaeralceae, new species (p. 138)

6. Transverse arm of clasper short, hardly extending beyond

middle of harpe (fig. 148) hildaella, new species (p. 140)

Transverse arm of clasper long, at least extending beyond middle

of harpe (figs. 142, 144, etc.) 7

7. Aedeagus sharply bent at middle (fig. 142a)- .gracilis (Walsingham) (p. 133)
Aedeagus slightly curved (figs. 149, 150, etc.) 8

8. Transverse arm of clasper arising at middle, or slightly before

middle of harpe; aedeagus strongly compressed (figs. 149, 151) 9

Transverse arm of clasper arising well before middle of harpe;

aedeagus not appreciably compressed (fig. 150) 10

9. Transverse arm of clasper reaching to or slightly beyond costa

of harpe ; free from below middle of harpe

klamathiana (Walsingham) (p. 142)
Transverse arm of clas^jer not reaching costa of harpe; free

from middle of harpe or beyond (fig. 149) sciadopa (Meyrick) (p. 144)

10. Both arms of clasper very slender, sharply pointed; vesica armed
with an elongate patch of fine cornuti (figs. 150, 150a)

nivalis (Braun) (p. 139)
Arms of clasper otherwise; vesica armed with an elongate patch

of strong cornuti (fig. 144) thoracenigraeella (Chambers) (p. 135)

KEY TO THE SPECIES OF MARTYRHILDA BASED ON FEMALE
GENITALIA

1. Ductus bursae partly sclerotized (figs. 100,239) 2

Ductus bursae membranous (figs. 232, 233; etc.) 3

2. Sclerotized portions of ductus bursae consisting of one moder-

ately large and one small area, the former bearing several
thornlike teeth inwardly (fig. 239) .thoracenigraeella (Chambers) (p. 135)
Sclerotized portion of ductus bursae consisting of one large
area armed inwardly wdth 17 or 18 short, stout teeth (fig. 100)

canella (Busck) (p. 128)

3. Inner surface of ductus bursae armed with a group of 11 stout

teeth (fig. 238) sordidella, new species (p. 132)

Inner surface of ductus bursae unarmed (figs. 232, etc.) 4

4. Signum broadly oval (figs. 231, 233) 5

Signum elongate (figs. 232, 235, etc.) 6

5. Ostium with a strongly sclerotized area laterally (fig. 233)

gracilis (Walsingham) (p. 133)
Ostium without such sclerotized area (fig. 231) nivalis (Braun) (p. 139)

6. Signum with several long teeth projecting from posterior end

(fig. 232) umbraticostella (Walsingham) (p. 130)

Signum without such teeth (figs. 235, etc.) 7

7. Bursa copulatrix distinctly asymmetrical (figs. 234, 236) 8

Bursa copulatrix symmetrical (figs. 235, 237) 9

8. Genital plate with two narrow raised ridges posterior to ostium

(fig. 234) klamathiana (Walsingham) (p. 142)

Genital plate without such ridges (fig. 236) sciadopa (Meyrick) (p. 144)

9. Ostium with sclerotized, cuplike structures laterally (fig. 235)

thoracefasciella (Chambers) (p. 136)
10. Ostium without such structures (fig. 237).. sphaeralceae, new species (p 138)

128 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

MARTYRHILDA CANELLA (Busck)

Plate 2, Figure! 17; Plate 6, Figure 44; Plate 10, Figures 67, 67a;
Plate 16, Figure 100

Depressaria canella Busck, Proc. U. S. Nat. Mus., vol. 27, p. 764, 1904.—

Meyrick, in Wytsman, Genera insectorum, fasc. ISO, p. 176, 1922. — Gaede,

in Bryk, Lepidopterorum catalogus, pt. 92, p. 311, 1939.
Agmopteryx canella Busck, Proc. U. S. Nat. Mus., vol. 85, p. 198, 1908.—

Forbes, Cornell Univ. Agr. Exp. Stat., Memoir 68, p. 238, 1923.
Agonopterix canella (Busck) McDunnough, Check list of the Lepidoptera

of Canada and the United States of America (Part 2, Microlepitloptera),

No. 8433, 1939.
Agnoptcryx canella (Busck) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6437, 1917.
Agonopteryx cogitata Braun, Can. Ent., vol. 58, p. 47, 1926.
Agonopterix cogitata (Braun) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera ) , No.

8433, 1939. (As. synonym of canella (Busck) ).
Depressaria cogitata (Braun) Gaede, in Bryk, Lepidopterorum catalogus, pt.

92, p. 316, 1939.

Labial palpus white strongly irrorated with fuscous. Antenna,
head, and collar of thorax fuscous to black; face whitish. Thorax
and ground color of fore wing pure white, the latter marked with
black, brown, and fuscous ; basal third with a few inconspicuous black
and fuscous spots and fine strigulae; from costa, at middle, a large
fuscous blotch extending to near middle of cell edged above and below
with brown and preceded by a black crescentic dash; around termen
a series of black or fuscous spots ; cilia whitish strongly suffused with
fuscous. Hind wing light fuscous; cilia whitish. Legs white
strongly mottled and overlaid with fuscous or black. Abdomen
white with a black lateral stripe on each side beneath.

Male genitalia. — Harpe lightly sclerotized except for sacculus,
which is heavily sclerotized and deeply folded ; sacculus produced to
form a broadly bifid clasper, transverse arm long, extending beyond
middle of harpe, the other produced toward cucuUus. Anellus a
broad, sclerotized plate, dorsoventrally concave; two dorsal projec-
tions laterally produced. Aedeagus stout, blunt; vesica armed with
several stout cornuti, the terminals (1-3) at right angles to the long
axis of the aedeagus forming hook. Vinculum broad, rounded.
Lobes of transtilla not fused. Gnathos a reniform, heavily spined
knob.

Female genitalia. — Ostium round. Genital plate heavily sclerotized
at edges of ostium ; ductus bursae short, constricted just before bursa
copulatrix ; the posterior portion broad, heavily sclerotized ; near the
anterior edge of the sclerotized portion of the ductus a group of 18
short, stout spines (this probably varies) . Signum of bursa copulatrix
a large oval plate.

REVISION OF THE OECOPHORIDAE — CLARKE 129

Larva.— Length 11-12 mm. Head light brown with a heavy suffu-
sion of dark brown on the margins, epicranial sutures and, especially,
beneath. Body subcylindrical, considerably thicker at middle and
tapering toward each end. Thoracic and abdominal segments light
yellowish green, lighter ventrally. Prothoracic shield light yellowish
brown. Thoracic legs yellowish with joints edged with brown.
Tubercles large, black; spiracles edged with brown.

The larvae of canella are often abundant, and their work is char-
acteristic and easily recognized. Early in spring the tubes, formed
by the larvae from tied terminal leaves, are conspicuous as "beaked"
processes at right angles to the long axis of the stem. Frequently
several tubes are constructed by one larva so that it becomes necessary
to inspect several tubes before the larva is finally located. From the
terminal portion of the tube the woolly covering of the leaves is ejected,
this often forming a conspicuous mass. Pupation occurs in debris
at the base of the plant. In the laboratory larvae pupated on April
24 and 25, and the moths emerged May 8, 1934.

The species is exceedingly difficult to rear. In the spring of 1935
nearly 400 larvae were collected but only 19 moths were obtained.
This might suggest faulty rearing conditions but three systems were
used with the same results. This and the fact that in nature the
moths are scarce, although the larvae are abundant, suggest a natural
high mortality.

Alar expanse, 16-20 mm.

Type. — In the United States National Museum {canella) ; A. F.
Braun collection, Cincinnati, Ohio {cogitata).

Type localities. — Pullman, Wash, {canella) (Piper) ; Aweme,
Manitoba {cogitata).

Food plants. — Antennaria luzuloides T. & I. (Clarke) ; Gtmphalium
(J. McDunnough).

Distribution. — ^Western United States and Canada in the Rocky
Mountain and intermountain areas; northeastern United States and
eastern Canada.

United States records

California: Warner Mountains, 3 mi. E., Davis Creek, Modoc County, alt. 5,500
feet, 3 (8-10-VII-1922, A. W. Lindsey).

Connecticut: New Haven, $ (Dr. Britten),

Idaho: Viola, Moscow Mountains, 5 (12-VI-35, J. F. G. Clarke [reared]).

ITe-w Hampshire: Portsmouth, $ (VI-9-05, C. E. Montgomery).

New York: Big Indian Valley, Catskill Mountains, 9 (IX-2-10, R. F. Pearsall) ;
Iliou i$, VI-25-13, 2 5 9, VI-25-13, VlI-11-17, H. McElhose) ; also Wil-
mington and Ithaca (according to Forbes) ; Ithaca, 9 (30-VI-31, A. B. Klots) .

V/ashington: Kamiack Butte, Whitman County, S $ S and 9 9 9 (May 8, 1934;
Jlay 16-June 18, 1935, J. F. G. Clarke [reared] ) ; Pullman, S (2 September
'98).
286614—41 7

130 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Canadian records

Alberta: Waterton Lakes (12 July, J. McDuunough).

British Columbia: Chilcotin, $ (VIII-22-25, George V. Copley).

Manitoba: Aweme.

Quebec: Kazubeque, $ (VI-24-27, J. McDunnough [reared]).

Remarks. — There is no doubt about the synonymy of this species.
Dr. Braun distinguishes her cogitata from canella on the basis of the
absence of the bhick anterior border of the thorax and the white apical
portion of the hind wing of the latter species. Although Busck did
not mention the black anterior portion of the thorax in his description,
all specimens I have seen, including the type, have it present. The
use of a white apical portion of the hind wing to distinguish further
between the two is impractical, no two specimens of this species having
the same amount of black scaling at the apex of the hind wing and some
lacking it entirely. The difference in the male genitalia is confined
to the number of cornuti present on the vesica; but this character is
useless since the number differs in different specimens.

This species closely resembles the European alstroemeriana but is
easily distinguished from it by the black or very dark-brown collar.

MARTYRHILDA UMBRATICOSTELLA (Walsingham)

Plate 24, Figtjres 143, 143a ; Plate 41, Figure 232

Depressaria umbraticostella Walsingham, Proc. Zool. Soc. London, 1S81, p. 318,
pi. 36, fig. 8. — Riley, in Smith, List of the Lepidoptera of Boreal America,
No. 5283, 1891.— Busck, Proc. U. S. Nat. Mus., vol. 24, p. 736, 1902 ; in Dyar,
U. S. Nat. Mus. Bull. 52, No. 5855, 1903.— Keaefott, in Smith, List of the
Lepidoptera of Boreal America, No. 6398, 1903.— Busck, Proc. U. S. Nat.
Mus., vol. 27, p. 763, 1904. — Meyrick, in Wytsman, Genera insectorum, fasc.
180, p. 177, 1922. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 356,
1939.

Agonopteryx umhraticostella (Walsingham) Busck, Proc. U. S. Nat. Mus., vol.
35, p. 198, 1908.— Bbaun, Trans Amer. Ent. Soc, vol. 51, p. 197, 1925.

Agonopterix umhraticostella (Walsingham) McDunnough, Check list of the
Lepidoptera of Canada and the United States of America (Part 2, Micro-
lepidoptera). No. 8455, 1939.

Agnopteryx umbraticostella (Walsingham) Barnes and McDunnough, Check
list of the Lepidoptera of Boreal America, No. 6434, 1917.

Labial palpus light ochreous suffused with light fuscous; third
segment with median and basal bands, the latter poorly defined.
Antenna fuscous. Head ochreous irrorated with fuscous. Thorax
and base of fore wing blackish fuscous to black. Fore wing light
reddish ochreous with small diffused blackish-fuscous spots on costa
and around termen ; on middle of costa a conspicuous, outwardly dif-
fused blackish-fuscous shade preceded by two minute (sometimes
inconspicuous) discal dots of the same color; from costal patch,
around termen to near middle of inner margin, a light fuscous shade;

REVISION OF THE OECOPHORIDAE — CLARKE 131

cilia light fuscous. Hind wing shining grayish fuscous. Legs
whitish ochreous, the fore and mid legs strongly overlaid with fus-
cous; hind legs lightly so with their tarsi fuscous and annulated with
whitish ochreous. Abdomen light fuscous above, whitish ochreouS
beneath with a lateral stripe on each side and irrorations blackish
fuscous.

Mate genitalia. — Harpe lightly clothed with hairs; costa heavily
sclerotized; clasper divided, the arms long, of about equal length;
one arm, parelleling the sacculus, stout and sharply pointed ; the trans-
verse arm curved, reaching costa and having a lateral thornlike
process projecting toward the sacculus. Anellus a more or less
shield-shaped plate, deeply excavated on the posterior margin, the
excavation, in the main, being formed by two heavily sclerotized,
winglike processes. Vinculum broad, rounded. Aedeagus stout,
straight, bluntly pointed, with a large spinulate patch about the
middle. Transtilla a broad sclerotized band. Gnathos reniform.
Tegumen bifid. Socii rectangular flaps bearing a few hairs.

Female genitalia. — Genital plate narrow at the middle, the entire
width being taken up by the rectangular ostium, then broadening
laterally. Ductus bursae short, broad, and membranous. Bursa
copulatrix large, oval; signum a large, elongate oval plate, slightly
broader anteriorly than posteriorly, with long, spinelike projections
at the posterior end.

Alar expanse, 16-19 mm.

Type. — In the British Museum.

Type localities. — Mount Shasta, Calif., and "North Oregon."

Food plants. — Balsamorhiza sagittata (Pursh) Nutt. and Helian-
fhits pumilits Nutt.

Distribution. — Western United States and Canada.

United States records

Arizona: White Mountains, Apache Indian Reservation, alt. 7,000 ft., 5 $ ^,
3 5 9 (June 1925, O. C. Poling) ; 5, 9 (July 5-15, 1925) ; ? (August 1-15,
1925).

California: Mount Shasta, $ ; Placer County, $ (September),

Colorado: Williams Range, $ (August 8, A. J. Snyder).

New Mexico: Fort Wingate, $ (July 16-23).

Oregon: "Camp Watson" 2 $ S (III, IV, 1872, Walsingham).

Utah: Stockton, 2 $ $ (IX-6, IX-7-07, Tom Spalding) ; Eureka {$, VII-13-11,
$ , VIII-14-11, 9 , VIII-29-11, Tom Spalding) .

Washing'ton: Almota, Whitman County, $ (IV-5-30, J. F. G. Clarke) ; Pull-
man, 2 $ $ (III-1-98, C. V. Piper), 9 (11-17-30, D. H. Brannon),
$ (VII-26-32, T. M. Clarke [reared]), $ (IV-13-26, J. F. G. Clarke), 9
(II 1-26-30, J. F. G. Clarke) ; Snake River, Whitman County, opposite
Clarkston, $ (IV-28-32, J. F. G. Clarke) ; Wenatchee, 2 $ $ (VI-5-30,
VI-10-30, A. Spuler), 9 (rX-5-29, A. Spuler).

132 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Canadiayi records
British Columbia: Vernon (20-XI-1927, A. A. Dennys).

Remarks. — This species seems to be confined to far-western North
America, where it is rather widely distributed. It was originally
described from specimens collected by Lord Walsingham on Mount
Shasta, Calif., and in North Oregon. Busck ^^ also records specimens
from Arizona. He further states that Dyar bred a specimen from
Helianthus pumilus collected near Denver and Sedalia, Colo.
Braun -* bred three specimens from BalsamorMza sagiftata collected
near Logan, Utah.

MARTYRHILDA SORDIDELLA, new species

Plate 24, Figttbes 145, 145a ; Plate 41, Figure 238

Labial palpus, head, thorax, abdomen, and ground color of fore
wing sordid whitish; second segment of labial palpus with sparse
brown and fuscous irrorations exteriorly ; third segment immaculate ;
antenna with basal segment sordid whitish; remainder of segments
fuscous with dull golden-yellow scaling above. Thorax and fore wing
irrorated with dull golden-yellow and brown; from apical third of
cost a around termen to inner margin a series of indistinct brownish
spots, with a few dull golden-yellow scales mixed; at basal third a
brown discal spot; below this spot in fold (vein Ic) an elongate dull
golden-yellow patch ; at end of cell a conspicuous brown discal spot ;
cilia white, rather shining; underside of fore wing fuscous, except
around edges. Hind wing smoky fuscous; cilia white with a brown
subbasal line. Legs sordid whitish, slightly overlaid and mottled
with fuscous exteriorly; tarsi fuscous except at joints.

Male genitalia. — Harpe moderately broad, tapering gi^adually to
the bluntly pointed cucullus, and lightly clothed with coarse hairs;
sacculus broad and very heavily sclerotized ; clasper divided, the longi-
tudinal arm produced as a strong, long, bluntly pointed extension of
the sacculus, the transverse arm as a short, stubby, moderately sclero-
tized projection about middle of harpe. Anellus a broad rectangular
plate with concave posterior edge, moderately well developed hairy
lateral lobes and a broad median process from basal edge. Aedeagus
stout, moderately long, nearly straight, bluntly pointed ; vesica armed
with an elongate patch of strong cornuti. Vinculum broadly rounded.
Transtilla a narrow sclerotized band with large, hairy, fleshy, lateral
lobes. Gnathos reniform. Tegumen pointed. Socii large fleshy,
hairy lobes.

Feinale genitalia. — Genital plate moderately broad, lightly scler-
otized. Ostium large, round, with an elongate, strongly sclerotized

23 Busck, A., Proc. U. S. Nat. Mus., vol. 24, p. 736, 1902.

2< Braun, A. F., Trans. Amer. Ent. Soc, vol. 51, p. 197, 1925.

REVISION OF THE OECOPHORIDAE — CLARKE 133

patch on each side inwardly ; anterolateral edge narrowly but heavily
sclerotized. Ductus bursae membranous, short, broad, with 10 or 11
short, stout, strongly sclerotized, thornlike teeth inwardly ; inception
of ductus seminalis adjacent to ostium. Bursa copulatrix moderately
large, round ; signum a large, round, toothed plate.
Alar expanse, 18-21 mm.
Type. — In the Canadian National collection.

Paratyfes. — U. S. N. M. No. 53258 ; also in the Canadian National
collection.

Type locality. — Shingle Creek, Penticton, British Columbia.
Food plant. — Unknown.

RemarJcs. — Described from the $ type, 10 $ and 3 $ paratypes
from British Columbia as follows: Brent's Landing, Penticton,
Z $ $ (30-V-1935) ; Shingle Creek, Penticton, & $ $ (1&-V-1936) ;
Shingle Creek Road, Keremeos, $ (8-VI-1935) ; Summerland, 2 9 $
(25-V-1935) ; Vaseaux Lake, 9 (18-V-1936), all collected by A. N.
Gartrell; Penticton, $ (7-VI-1935, J. McDunnough). The entire
type series was submitted by Dr. McDunnough.

This is the third species of the genus from North America with a
white ground color to come to my attention.

The costa of the fore wing of the female is somewhat straighter
than that of the male and is more nearly parallel to the inner margin,
producing a wing that is narrower and of nearly equal width
throughout its length.

MARTYRHILDA GRACILIS (Walsingham)

Plate 24, Figotjes 142, 142ji ; Plate 41, Figube 233

Depressaria gracilis Walsingham, Ins. Life, vol. 1, p. 257, 1889. — Busck. Proc.

U. S. Nat. Mus., vol. 24, p. 737, 1902; in Dyar, U. S. Nat. Mus. Bull. 52,

No. 5857, 1903.— Keaefott, in Smith, List of the Lepidoptera of Boreal

America, No. 6400, 1903. — IVIeyrick, in Wytsman, Genera insectorum, fasc.

180, p. 177, 1922.— Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 325,

1939.
Agonopteryx gracilis (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35, p.

198, 1908.
Agonopterix gracilis (WalsinglVam) McDunnough, Check list of the Lepidoptera

of Canada and the United States of America (Part 2, Microlepidoptera),

No. 8457, 1939.
Agnopterjix gracilis (Walsingham) Barnes and McDunnough, Check list of the

Lepidoptera of Boreal America, No. 6436, 1917.

Labial palpus straw yellow ; second segment blotched with reddish
brown outwardly; third segment with a reddish-brown subapical
annulus. Antenna with basal segment dark brown, remainder brown.
Head straw yellow with a light reddish-brown patch above the base
of each antenna. Thorax and base of forcwing dark reddish brown.
Ground color of fore wing and cilia straw yellow; discal spots dark

134 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

brown, the first two, obliquely one above the other, before middle,
the third at end of cell ; from costa, just before apex, around termen
a series of six or seven dark brown spots; cilia strongly mixed with
dark brown. Hind wing pale grayish fuscous; cilia pale straw
yellow with a faint, narrow, brown subbasal line. Legs straw yellow ;
anterior femora and tarsi strongly overlaid with brown; tarsi of
middle and hind legs tinged with brown. Abdomen brownish above
with posterior edges of segments and ventral surface straw yellow.

Male genitalia. — Harpe moderately broad; costa heavily sclerotized
to a point just before cucullus ; clothing of hairs confined to the apical
half except for a group of coarse hairs on base just below the costa ;
clasper divided, one arm, paralleling the sacculus, short and acutely
pointed, the other, long, reaching to a point just below the costa, with
a short, blunt lateral projection. Anellus a broad, sclerotized plate
with an elongated troughlike articulation on which the aedeagus
rests. Vinculum rounded. Aedeagus stout, bent sharply at middle
and tapering gently to a blunt point. Transtilla a narrow band with
hairy lateral lobes. Gnathos an oval, spined knob. Tegumen rounded
at apex. Socii fleshy flaps with a few hairs.

Ferrbale genitalia. — Genital plate broad, rather heavily sclerotized
around the ostium. Ostium deeply concave. The edges inunedi-
ately around the ostium and the posterior portion of the ductus bursae
are spinulate. Just before the ostium the ductus bursae is constricted.
Ductus bursae short, membranous, unarmed on inner surface, widen-
ing gently into the large bursa copulatrix. Signum a large, broadly
oval, slightly sclerotized plate.

Alar expanse, 15-18 mm.

Type. — In the British Museum.

Type locality. — Texas.

Food plant. — Unknown.

Distribution. — ^Western United States.

United States records

California: Laguna Beach, 9 ; Loma Linda, San Bernardino County, ? (Oct.
&-15) ; San Diego, 2 $ $ (Nov. 21, 1921, Karl R. Coolidge), S $ S (May
24-30), 9 (June 16-23), 9 (April 16-23), 9 (IV-26-08, Geo. H. Field).

Colorado: Colorado Springs (Fountain Valley School), 3 $$ ( 20-31- VIII-32,
A. B. Klots) ; Denver, $ (Oslar) ; 2 9 9 (no data).

Iowa: Iowa City, 9 (G. G. Ainslie).

South Dakota: Elk Point, 9 (Aug. 1918, C. N. Ainslie).

Texas: 2 $ $ (X-06, H. Lacey).

Remarks. — This very distinct species cannot be confused with any
other described North American Martyrhilda, but it closely resembles
the European culcitella.

REVISION OF THE OECOPHORIDAE — CLARKE 135

MARTYRHILDA THORACENIGRAEELLA (Chambers)
Plate 24, Figuee 144 ; Plate 41, Figure 239

Oelechia thoracenigraeella Chambers, Cincinnati Quart. Jouru. Sci., vol. 2, p. 246,

1875 ; U. S. Geol. Geogr. Surv. Terr. Bull. 4, p. 147, 1878.
Depressaria thoracenigraeella (Chambers) Rilet, in Smith, List of the Lcpidop-

tera of Boreal America, No. 5495, 1891.— Busck, Proc. U. S. Nat. INIus., vol. 24,

p. 736, 1902 ; in Dyar, U. S. Nat. Mus. Bull. 52, No. 5856, 1903.— Kearfott, in

Smith, Check list of the Lepidoptera of Boreal America, No. 6399, 1903.—

Meyrick, in Wyt&man, Genera insectorum, fasc. 180, p. 176, 1922. — Gaede, in

Bryk, Lepidopterorum catalogus, pt. 92, p. 355, 1939.
Agonopteryx thoracenigraeela (Chambers) Busck, Proc. U. S. Nat. Mus., vol. 35,

p. 198, 1908.
Agonopterix thoracenigraeella (Chambers) McDunnough, Check list of the

Lepidoptera of Canada and the United States of America (Part 2, Micro-

lepidoptera ) , No. 8456, 1939.
Agnopteryx thoracenigraeella (Chambers) Barnes and McDunnough, Check

list of the Lepidoptera of Boreal America, No. 6435, 1917.

Labial palpus whitish ochreous; second segment irrorated with
fuscous; third segment with a broad, well-defined black subapical
anmilus and a narrower, poorly defined basal annulus of the same
color. Antenna fuscous. Head, thorax, and ground color of fore
wing pale ochreous-gray ; fore part of thorax and base of fore wing
blackish fuscous ; from costa to middle of cell a fuscous shade edged
above and below with brown and preceded by a conspicuous outwardly
oblique black dash; on costa and around termen a series of grayish-
fuscous spots; cilia light fuscous irrorated with wliitish ochreous.
Hind wing brownish fuscous ; cilia somewhat lighter with a white ter-
minal edge. Legs light grayish ochreous overlaid and mottled with
fuscous ; hind tarsi fuscous, annulated with whitish ochreous. Abdo-
men grayish ochreous above, beneath whitish ochreous with a black
lateral line on each side.

Male genitaUa. — Essentially like gracilis^ diflfering chiefly in the
characters of the aedeagus. Harpe moderately broad and clothed with
fine hairs, especially in apical half; clasper divided, one arm parallel-
ing sacculus, the other transverse, arising well before middle of harpe,
reaching nearly to costa, and without lateral process ; sacculus heavily
sclerotized; anellus a sclerotized plate with median trough on which
the aedeagus articulates ; lateral hairy lobes present. Aedeagus stout,
only slightly curved ; cornuti numerous, strong, in an elongate patch.
Vinculum rounded, broad. Transtilla a narrow sclerotized band with
lateral hairy lobes. Gnathos an oval spined knob. Socii fleshy hairy
lobes. Apex of tegumen with median cleft.

Female genitalia. — Ostium moderately large, located near anterior
edge of genital plate. Ductus seminalis entering ductus bursae just
anterior to ostium. Ductus bursae membranous except for two sclero-

136 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

tized patches, one large, tlie other small, each set with sharp thornlike
projections. Signiim of bursa very large, oval.

Alar expanse, 16-17 mm.

Type. — In the Museum of Comparative Zoology, Cambridge, Mass.

Type locality. — Behrens, Calif.

Food plant. — Unknown.

Distribution. — Known only from California.

United States records
California: Carmel, 4^5,9 ("June," A. H. Vachell).

Reviarhs. — The type of thoracenigraeella, in the Museum of Com-
parative Zoology, Cambridge, is in verj^ poor condition. The right
fore wing, in good condition, is still attached to the thorax, but the
abdomen, most of the legs, and all the other wings are gone.

Despite the poor condition of the type I have been able to recognize
five specimens as this species. The figures of the genitalia have been
made from two of these specimens.

MARTYRHILDA THORACEFASCIELLA (Chambers)

Plate 24, Figures 146, 146a ; Plate 41, Figure 235

Gelechia thoracefasciclla Cuaubers, Cincinnati Quart. Journ. Sci., vol. 2, p. 246,

1875; Can. Ent., vol. 10, p. .50, 1878; U. S. Geol. Geogr. Surv. Terr. Bull. 4,

p. 147, 1878.
Depressaria thoracefasciella (Chambers) Riley, Iji Smith, List of the Lepidop-

tera of Boreal America, No. 5494, 1891.— Busck, Proc. U. S. Nat. Mus., vol.

24, p. 740, 1902 ; in Dyar, U. S. Nat. Mus. Bull. 52, No. 5867, 1903.— Keakfott,

m Smith, List of the Lepidoptera of Boreal America, No. 6410, 1903. — Gaede,

in Bryk, Lepidopterorum catalogus, pt. 92, p. 354, 1939.
Depressaria thoracifasciella (Chambers) Meyeick, in Wytsman, Genera insec-

torum, fasc. 180, p. 176, 1922.
Agnoptcryx thoracefasciella (Chambers) Babnes and McDunnough, Check list

of the Lepidoptera of Boreal America, No. 6451, 1917.
Agonopterix tJioracefasciella (Chambers) McDunnough, Check list of the

Lepidoptera of Canada and the United States of America (Part 2, Micro-

lepidoptera), No. 8442, 1939.

Labial palpus with second segment ochreous-white, mottled with
fuscous exteriorly; third segment blackish fuscous with a poorly
defined ochreous-white median band and ochreous-white tip.
Antenna blackish fuscous. Head, thorax, base and basal fourth of
costa of fore wing whitish ochreous. Ground color of fore wing
brownish fuscous sparsely irrorated with whitish ochreous; extreme
base of costa, and shade beyond whitish-ochreous base, blackish
fuscous; first two discal spots small, black, obliquely placed one above
the other ; at the end of cell a conspicuous white or whitish-ochreous
discal spot surrounded by a blackish-fuscous suffusion; along costa
and around termen a series of poorly defined blackish-fuscous spots ;
cilia fuscous, mixed with ochreous-white. Hind wing brownish

REVISION OF THE OECOPHORIDAE — CLARKE 137

fuscous, lighter basally; cilia fuscous. Legs blackish fuscous irro-
rated with ochreous-white ; tarsi annulated with ochreous-white.
Abdomen fuscous above; beneath, ochreous-white overlaid and irro-
rated with fuscous.

Male genitalia. — Harpe moderately sclerotized; cucullus rounded
and clothed with fine hairs. Transverse and longitudinal arms of
clasper of about equal length, the former extending beyond the costa ;
sacculus as broad as one-third the width of harpe at base. Anellus
a roughly rectangular sclerotized plate emarginate on the posterior
edge; lateral lobes minute. Transtilla a narrow band with fused
hairy lobes. Vinculum rounded, broad. Aedeagus slender, about
one-half the length of harpe, slightly curved, unarmed ; apex slightly
recurved. Gnathos an oval spined knob ; supporting arms moderately
sclerotized. Socii small, clothed with few hairs; widely separated.
Tegumen truncated, slightly emarginate.

Female genitalia. — Ostium large, rounded; on each side of the
ostium the genital plate is produced into a strongly sclerotized pocket.
Inception of the ductus seminalis slightly anterior to the sclerotized
pockets. Ductus bursae membranous, unarmed on inner surface,
tapering gradually into the symmetrical bursa copulatrix. Signum
an elongate scobinate plate attenuated and broken posteriorly.

The abdominal wall of this species differs from that of sphaeralceae
in being much less strongly sclerotized. The harpes of the male are
broader and correspondingly shorter, and much less sclerotized. The
aedeagus is shorter than that of sphaeralceae and the apex is more
strongly recurved. The female genitalia show marked differences.
The signum of sphaeralceae is smaller and less strongly sclerotized
than that of thoracefasciella and the sclerotized pockets of the latter
species are totally wanting in the former.

Alar expanse, 15-17 mm.

Type. — In the Museum of Comparative Zoology, Cambridge, Mass.

Type locality. — Behrens, Calif.

Food plants. — Siclahea malvaeflora A. Gray (Keifer rearing) and
Malvaf (C. M. Dammers).

Distrihution. — Southwestern United States.

United States records

Arizona: Hualapi Mountains, Mojave County, 2 $ $, 9 (May 24-31, no col-
lector).

California: Diamond Spring, Eldorado County, 3 $ $, 3 5 9 (V-26 to V-3-35,
H. H. Keifer) ; Canyon Valley, San Bernardino County, $ (VII-1-32, no
collector) ; Riverside, 9 (June 1930, C. M. Dammers) ; Mill Valley, Marin
County, 4 $ $ (5-12-III-20, E. P. Van Duzee).

Reinarks. — I have examined the type of this species and have made
a slide of the male genitalia. These agree exactly with specimens
from our reared series.

138 PROCEEDINGS OF THE NATIONAL MUSEUM vol. »o

MARTYRHILDA SPHAERALCEAE, new species

Plate 24, Figures 147, 147a ; Plate 41, Fiqube 237

A small, gray, narrow-winged western species, closely related to the
foregoing, but easily distinguished from it by the genitalia.

Head, palpus, and antenna dark grayish fuscous; face whitish
ochreous ; second segment of labial palpus whitish ochreous inwardly
and above with a liberal sprinkling of whitish-ochreous scales out-
wardly; third segment whitish ochreous above, the color extending
around middle to form an incomplete annulus ; apex whitish ochreous ;
antenna with narrow fuscous annulations. Thorax and ground color
of fore wing grayish fuscous, the tips of the scales narrowly whitish
ochreous ; basal part of fore wing, usually with more whitish-ochreous
scaling, appearing lighter than ground color and followed by a dark
fuscous shading ; at basal third two black obliquely placed discal spots
followed by white scales, the spots frequently confluent, forming an
outwardly oblique discal dash ; at end of cell a white discal spot sur-
rounded by blackish-fuscous scales; along costa and around termen
a series of indistinct fuscous spots ; underside of costa whitish ochreous
to apex ; a row of black scales around termen ; cilia gray mixed with
fuscous and whitish-ochreous scales. Hind wing grayish fuscous with
light fuscous cilia; a fuscous band around margin at base of cilia;
a blackish-fuscous line around apex and termen beneath. Legs fus-
cous irrorated with whitish ochreous; tarsi annulated with wliitish
ochreous. Abdomen fuscous above with grayish at the posterior mar-
gin of each segment ; grayish laterally and beneath with a row of black
lateral spots.

Male genitalia. — Harpe ample, strongly sclerotized ; cucullus blunt-
pointed, clothed with fine hairs ; transverse arm of clasper appreciably
longer than longitudinal arm, reaching well beyond costa; sacculus
broader than one-third the width of harpe at base. Anellus roughly
rectangular, emarginate on the posterior edge; lateral lobes scarcely
developed. Transtilla a narrow band with well-developed, fused,
lateral hairy lobes. Aedeagus slender, slightly curved, longer than
one-half the length of harpe, pointed, unarmed. Vinculum rounded.
Gnathos an oval spined knob; supporting arms strongly sclerotized.
Socii fleshy hairy lobes, well separated. Tegumen with the apex
emarginate.

Female genitalia. — Ostium small, round at extreme anterior edge
of genital plate. Genital plate broad, weakly sclerotized. Ductus
bursae moderately stout, membranous, tapering gradually into bursa ;
inception of ductus seminalis just anterior to the ostium. Bursa
copulatrix symmetrical, oval; signum a small, weakly sclerotized,
elongate plate.

Alar expanse, 15-18 mm.

REVISION OF THE OECOPHORIDAE — CLARKE 139

Larva. — Length 9-11 mm. Head and cervical shield light brown,
the former broadly suffused laterally and posteriorly with dark
brown ; epicranial sutures dark brown ; ocelli light brown surrounded
by a blackish area. Underside of head yellowish brown. The shield
is bisected by a narrow light-green or yellowish longitudinal line;
posterior half of shield dark brown; beyond this, on dorsal half,
prothorax whitish. Thoracic and abdominal segments apple green
with a broad suffusion of whitish around the dark brown tubercles
and on the posterior edges of the segments. Setae long, yellowish
brown proximally, whitish distally. Anal plate pale green with
whitish posterolateral edges. Thoracic legs yellowish brown with
lighter annulations at the joints.

The larva is a leaf miner forming a blotch mine. In the early
stages frass is ejected from the mine through a small silken tube,
which is constructed at one edge of the mine, usually parallel to
the midrib of the leaf. In the later instars the larva folds a leaf
and completely mines out the spongy material between the upper and
lower epidermal layers. The larva remains in this large, roomy mine
until ready to pupate. Pupation occurs in sand or soil at the base
of the plant.

The larvae are abundant in the Grand Coulee, where nearly all
plants of Sphaeralcea mMnroana, which are numerous, are heavily
infested. The leaves are badly discolored owing to the thoroughness
with which they are mined.

ry/>e.— U.S.N.M. No. 52076.

Type locality. — Park Lake, Grant County, Grand Coulee, Wash.

Food plant. — Sphaeralcea munroana (Dougl.) Spach.

Remarks. — Described from the $ type, 5 $ and 4 9 paratypes
(V-15 to VI-12, 1935, J. F. G. Clarke). Paratypes in United States
National Museum, Canadian National, and H. H. Keifer collections.

MARTYRHILDA NIVALIS (Braun)

Plate 25, Figures 150, 150a ; Plate 40, Ftgltre 231

Agonopteryx nivalis Braun, Proc. Acad. Nat. Sci. Philadelphia, vol. 73, p. 10, 1921.

Agonopteris nivalis (Braun) McDunnough, Check list of the Lepidoptera of Can-
ada and the United States of America (Part 2, Microlepidopteru), No. 8419,
1939.

Depressaria nivalis (Braun) Gaede, in Bryk, Lepidopterorum catalogus, pt. 92,
p. 334, 1939.

Labial palpus white except for some light-brown scaling on the out-
side of second segment. Antenna fuscous. Head, thorax, and fore and
hind wings white; slightly beyond base of fore wing, in inner angle
and along inner margin, a faint light-brown shade ; two discal spots,
obliquely one above the other, at basal third, another, larger, white-
centered discal spot at end of cell, a series of spots along costa and

140 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

around termen and sparse irrorations over entire surface of fore wing
fuscous to blackish fuscous. Legs and abdomen white, the former
strongly suffused with fuscous.

Male genitalia. — Harpe ample, gently tapering toward cucullus, the
latter rounded. Sacculus strongly sclerotized; both arms of clasper
very slender, sharply pointed, the transverse arm arising well before
middle of harpe and reaching two-thirds distance toward costa. Anel-
lus a weakly sclerotized broad plate with well developed, slightly hairy,
lateral lobes. Transtilla a weakly sclerotized band with well developed
lateral lobes. Aedeagus slender, slightly curved, not appreciably com-
pressed, bluntly pointed with elongate, spinulate patch on vesica. Vin-
culum rounded. Gnathos an oval spined knob. Socii weakly sclero-
tized, sparsely hairy, fleshy lobes.

Female genitalia. — Ostium small, round, without strongly sclero-
tized area laterally. Ductus bursae membranous, slender, inception of
ductus seminalis on right side just before ostium. Bursa copula trix
large, pear-shaped, with strongly sclerotized oval signum.

Alar expanse, 21-23 imii.

Type. — In collection of Dr. A. F. Braun, Cincinnati, Ohio.

Type locality. — Two Medicine Lake, Glacier National Park, Mont.

Food plant. — Unknown.

Distribution. — ^Western United States and Canada in Mountains.

United States records

"Washington: Skyline Ridge, Mount Baker district, Wliatcom County, alt. 6,000
feet, 5,399 (26-VII-25 and 16-VIII-30, J. F, G. Clarke).

Wyoming: Green River Lake, Wind River Range, 12 5 5 , 9 (July 24 to August
7, 1935, A. B. Klots).

Canadian records

Alberta: Lake Louise, $ ("VIII-1918").

Remarks. — ^This is one of three known white species in this genus
from North America. The others, canella and sordidella (new spe-
cies) , cannot be confused with it.

Dr. Braun has compared a male with her type and has kindly
verified my determination of the species.

The venation is subject to considerable variation, one female having,
on the right side, an accessory cell in the fore wing between veins 9
and 11 that is included in the discal cell ; vein 7 of the hind wing is
forked.

MARTYRHILDA HILDAELLA, new species

Plate 24, Fioxtkes 148, 148a

Labial palpus sordid whitish ; second segment with sparse black and
fuscous irrorations exteriorly, mostly confined to basal half, and with
slight infuscation in the undivided slender brush; third segment im-

REVISION OF THE OECOPHORIDAE — CLARKE 141

maculate. Antenna grayish fuscous with narrow fuscous annulations.
Head, thorax, and base of fore wing sordid whitish, the latter diffused
along costa to slightly beyond basal third ; tegula with a pale brownish
suffusion toward apex ; fore wing strongly irrorated with contrasting
black spots, especially along veins; in the light basal patch, slightly
below costa, a conspicuous black spot; beyond basal patch a transverse
dark fuscous dash, which does not reach costa and which rapidly fades
to a pale brownish ochreous, becomes stronger in color from middle to
apical third, where it is followed by sordid whitish or cinereous and
is narrowly diffused along costa to apex ; at middle of cell a conspicuous
though small black discal spot ; at end of cell a white discal spot edged
with black ; above and below the latter spot considerable black scaling
fusing with the brownish-ochreous shade; from apical third of costa,
around termen to inner margin a series of blackish dashes edged in-
wardly and narrowly with pale yellowish brown; whole surface of
wing somewhat lustrous and appearing predominantly gi'ay; cilia
light fuscous, darker basally. Hind wing shining grayish fuscous;
cilia brownish fuscous with a fuscous basal band. Legs ochreous-white
strongly overlaid and suffused with fuscous except at joints. Abdomen
light grayish fuscous with some cinereous and white scales mixed and
with a median longitudinal fuscous shade beneath.

Male genitalia. — Harpe rather broad, clothed with coarse hairs in
outer half; basal half heavily sclerotized except for a small naem-
branous median area; costa and ventral margin parallel and evenly
curved ; cucullus rounded ; sacculus rather narrow, very strongly sclero-
tized; clasper divided, the longitudinal arm produced as a strong
pointed extension of the sacculus, the transverse arm a slender pointed
process extending but slightly beyond middle of harpe. Anellus a
broad, subrectangular plate with large hairy lateral lobes, concave
posterior edge and broad sagittate basal process. Aedeagus stout,
moderately long, gently curved; vesica armed with a large elongate
patch of rather strong cornuti. Vinculum narrov. ly rounded. Trans-
tilla a narrow sclerotized band with large hairy lateral lobes. Gnathos
oval. Tegumen truncate. Socii fleshy, hairy lobes.

Alar expanse, 18-21 mm.

Type. — In the Canadian National Collection.

Paratypes. — U. S. N. M. No. 53259 ; also in the Canadian National
Collection.

Type locality. — Cameron Bay, Great Bear Lake, Northwest Terri-
tories, Canada.

Food plant. — Unknown.

Remarks. — Described from the $ type and 5 $ paratypes all from
the type locality ( 8-12-VII-1937, T. N. Freeman). I have seen no
females.

142 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

MARTYRHILDA KLAMATHIANA (Walsingham)

Plate 25, Figure 151 ; Plate 41, Figube 234

Depressaria Idamathiana Walsingham, Proc. Zool. Soc. London, 1S81, p. 314,
pi. 36, fig. 4. — Riley, m Smith, List of the Lepidoptera of Boreal America,
No. 5267, 1891.— BuscK, Proc. U. S. Nat. Mus., vol. 24, p. 740, 1902 ; in Dyar,
U. S. Nat. Mus. BuU. 52, No. 5868, 1903.— Keaefott', in Smith, List of the
Lepidoptera of Boreal America, No. 6411, 1903. — BuscK, Proc. U. S. Nat. Mus.,
vol. 27, p. 762, 1904.— Dyae, Proc. U. S. Nat. Mus., vol. 27, p. 934, 1904.—
Andeeson, Catalogue of British Columbia Lepidoptera, No. 1091, 1904. — •
Meyeick, in Wytsman, Genera insectorum, fasc. 180, p. 176, 1922. — Gaede, in
Bryk, Lepidopterorum catalogus, pt. 92, p. 328, 1939.

Agonopteryx JdamatMana (Walsingham) Busck, Proc. U. S. Nat. Mus., vol. 35,
p. 198, 1908.

Agonoptcrix klamathiana (Walsingham) McDunnough, Check list of the Lepi-
doptera of Canada and the United States of America (Part 2, Microlepidop-
tera). No. 8435, 1939.

Agnopteryx klamathiana (Walsingham) Baenes and McDunnough, Check list
of the Lepidoptera of Boreal America, No. 6441, 1917.

Depressaria cinifiuaella Walsingham (not Zeller), Ins. Life, vol. 1, p. 256, 1889. —
Riley, m Smith, List of the Lepidoptera of Boreal America, No. 5259, 1891. —
BuscK (not Zeller), Proc. U. S. Nat. Mus., vol. 24, p. 740, 1902; in Dyar,
U. S. Nat. Mus. Bull. 52, No. 5869, 1903.— KKVEForr, in Smith, List of the
Lepidoptera of Boreal America, No. 6412, 1903. — Andeeson, Catalogue of
British Columbia Lepidoptera, No. 1092, 1904. — Meyrick, in Wytsman, Genera
insectorum, fasc. 180, p. 176, 1922 (part). — Gaede, in Bryk, Lepidopterorum
catalogus, pt. 92, p. 314, 1939.

Agonopteryx ciniflonella Busck (not Zeller), Proc. U. S. Nat. Mus., vol. 35, p. 198,
1908.

Agnopteryx ciniflonella Baenes and McDunnough (not Zeller), Check list of
the Lepidoptera of Boreal America, No. 6440, 1917.

Agonopterix ciniflonella McDunnough (not Zeller), Check list of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera),
No. 8436, 1939.

Labial palpus gray; second segment strongly mixed with reddish
fuscous; third segment with a broad, fuscous annulus before apex.
Antenna reddish gray. Thorax, base of fore wing, and costa to about
middle sordid whitish ; anterior part of thorax suffused with reddish
fuscous ; ground color of fore wing reddish gray strongly suffused with
reddish fuscous; at basal third two obliquely placed black discal dashes
of raised scales followed by white or grayish scales ; beyond this light
patch a fuscous shade ; at the end of cell a white discal spot margined
by blackish fuscous ; around termen a series of fuscous dots ; cilia dull
reddish with a narrow, fuscous median band ; under side of costa nar-
rowly edged with carmine. Hind wing grayish fuscous; cilia light
fuscous with a rosy tinge and a fuscous basal band. Legs grayish
overlaid with fuscous and strongly suffused with carmine. Abdomen
grayish fuscous above; beneath sordid whitish with a broken, black
lateral line on each side.

REVISION OF THE OECOPHORIDAE — CLARKE 143

Male genitalia. — Harpe with the apical and basal portion just below
costa clothed with coarse hairs ; marginal hairs fine ; sacculus heavily
sclerotized; clasper divided; transverse arm straight, slender, arising
at or before middle of harpe and reaching to or slightly beyond costa ;
at the base of the arm a sclerotized portion forming a short, sharp
thorn (in some specimens this is scarcely noticeable) ; the other arm
is short, sharply pointed and parallel to the sacculus; cucullus
rounded. Anellus a rectangular plate, deeply cleft on the posterior
edge; lateral lobes weak. Vinculum rounded. Aedeagus broad
dorsoventrally and compressed laterally. The dorsal edge is strongly
sclerotized ; both ends slightly enlarged ; vesica with a spinulate patch
at middle. Transtilla a narrow sclerotized band ; lateral lobes large,
not fused. Gnathos a spined oval knob. The gnathos varies slightly
in different specimens. Socii broad hairy flaps.

Female genitalia. — Genital plate broad, with two narrow, raised
ridges posterior to ostium. Ostimn small, roimd and situated at ex-
treme anterior edge of plate. Ductus bursae membranous, short, and
tapering into the bursa copulatrix. The latter is large, asymmetrical.
Signum an elongate-oval, sclerotized, scobinate plate, extending almost
the entire length of the bursa.

Alar expanse, 20-25 mm.

Type. — In the British Museum.

Type locality. — Fort Klamath, Oreg.

Food plant. — Apple?

Distribution. — Western United States and Canada as far east as
Ontario.

United States Records

Montana: Ravalli County, 3 $$,9 (1-29-33, from magpie nests).
Oregon: Fort Klamath, 2 $ $ (IX-20-71, Walsingham).

Washington: Bellingham, ? (VIII-22-26, J. F. G. Clarke) ; Logan Hill, Chehalis,
3 5 5,29? (11-16 to in-28-30, T. M. Clarke).

Canadian records

British Columbia: Duncan, $ (Apr. 1-7, Hanham) ; Fraser Mills, 5 (IX-5-20,
L. E. Marmont) ; Hazelton, $ (IX, 14-21, W. B. Anderson) ;' Kalso, $
(IV-24-15, J. W. Cockle) ; Marron Lake, $ (Apr. 1924, C. B. Green) ; Quami-
chan Lake, Vancouver Island, $ (IX-27-02) ; Vancouver, $ (III-18-96,
Livingston) ; Victoria, $ (IV-18-17, E. H. Blackmore), $ (VIII-29-23, K. F.
Auden), $ (III-10-23, W. R. Carter) ; Wellington, 3 5 5,6 5 $ (V-25-07,
April [3 specimens] ; 3 specimens, III-22-03, G. "W. Taylor).

Ontario: Hymers, 2 5 5 (no date or collector) ; Ottawa, 5 (II 1-18-05, C. H.
Young).

RenypLvks. — After studying 45 specimens and 29 wing and genitalia
slides, I am convinced that all the above material is referable to this
species. I admit that the series studied shows many minor superficial
differences existing between specimens. In genitalia, however, even

144 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

though slight variations occur, there are no major differences and none
which would separate one group of specimens from another.

In addition, some specimens are more gray than others, having little
of the red scaling that is characteristic of the species ; others possess a
white, rather than gray, head.

The European ciniflonella is a gray species with very narrow wings.
The costa is not strikingly lighter as in most specimens of Mamathiana.
One or two North American specimens I have seen approach cmi-floneTla
very closely. The wing form appears to vary considerably due to the
differences in length of veins ^9 of the fore wing. In some specimens
these veins are considerably longer than in others, and the fore wings
are correspondingly longer. It is possible that what I consider Ma-
mathiana actually includes more than one species, or one in a state of
flux, but this entire group {Jdmnathiana^ sciadojm, cinifloneUa, and a
fourth unnamed) represents a very difficult complex of species. Only
careful rearing will determine the status of these.

MARTYRHILDA SCIADOPA (Meyrick)

Plate 24, Figures 149, 149a ; Plate 41, Figuee 236

Depressaria sciadopa Meykick, Exotic Microlepidoptera, vol. 2, p. 315, 1920 ; in

Wytsman, Genera iiisectorum, fasc. ISO, p. 176, 1922. — Gaede, in Bryk,

Lepidopterorum catalogus, pt. 92, p. 348, 1939.
Affonopterix sciadopa (Meyrick) McDunnough, Check list of the Lepidoptera of

Canada and the United States of America (Part 2, Microlepidoptera), No.

8434, 1939.

This species is like the foregoing, kh?7iathiana, but lacks the red
or brown scaling of that species, is without carmine suffusion, and
averages smaller in size.

Labial palpus whitish gray; second segment irrorated and suffused
with grayish fuscous exteriorly; third segment with broad supra-
medial grayish-fuscous annulus. Antenna grajdsh fuscous above,
whitish gray beneath. Head, thorax, and ground color of fore wing
whitish gray. Head lightly irrorated with grayish fuscous. Thorax
strongly overlaid with grayish fuscous and irrorated with black, the
darker colors almost obscuring the lighter ground color. Fore wing
suffused with grayish fuscous and irrorated with black; extreme
base of wing and basal half of costa whitish gray strigulated with
grayish fuscous and with a small black spot near base slightly inside
the costal edge; beyond the light costal and basal areas a strong
blackish to grayish fuscous shade rapidly fading to the light ground
color slightly beyond basal third; a similar, but smaller, dark shade
at middle of wing; at basal third a pair of small black discal spots,
one above the other, followed by white scales; sometimes these two
spots are confluent, forming a short, outwardly oblique black dash ; at
the end of cell a small white spot edged with black ; at apical fourth a

REVISION OF THE OECOPHORIDAE — CLARKE 145

narrow, faint, outwardly curved grayish-fuscous shade; along costa
and around termen a series of small black spots ; cilia whitish gray ir-
rorated with grayish fuscous. Hind wing whitish gray basally, gray-
ish fuscous apically; cilia whitish gray with a grayish-fuscous
subbasal band. Legs whitish gray overlaid and suffused with grayish
fuscous except at joints. Abdomen whitish gray suffused with gray-
ish fuscous above and with a poorly defined row of black spots on
each side beneath.

Male genitalia. — Harpe rather heavily sclerotized except for the
cucullus and a small area about the middle; cucullus and the sclero-
tized costal area moderately clothed with fine hairs (the cucullus of
one male is considerably rounded while those of four other males are
somewhat pointed) ; clasper strongly sclerotized; the free transverse
arm originating about the middle of the harpe; longitudinal arm
short, sharply pointed ; sacculus heavily sclerotized. Anellus a narrow
concave plate forming a trough on which the aedeagus articulates;
lateral lobes weak. Vinculum rounded. Aedeagus stout, slightly
curved, very much compressed, rather broad dorsoventrally, constricted
at about the middle; dorsal edge heavily sclerotized; vesica armed
with a patch of very small cornuti. Transtilla a narrow sclerotized
band with well-developed lateral lobes. Gnathos an oval spined knob,
Socii fleshy lobes clothed with fine hairs. Apex of tegumen bluntly
pointed ; ventral edges with rather well developed outgrowths. Abdo-
men of male heavily sclerotized.

Female genitalia. — Genital plate broad with the small romid ostium
situated at the extreme anterior edge. Ductus bursae short, membra-
nous, stout, tapering into the large asymmetrical bursa copulatrix;
signum an elongate sclerotized plate with small thornlike projections
covering the surface.

Alar expanse, 15-19 mm.

Tyjye. — In the British Museum.

Type locality. — Field, British Columbia, Canada.

Distribution. — Canada and northeastern United States.

United States records

New Hampshire: Jefferson, 2 $ $ (30-III-36, 21-X-1936, A. E. Brower).
New York: Oswegatchie, 5, 2 $ $ (31-III-33, A. B. Klots).

Canadian records

Alberta: Edmonton, 2 S $, 2 ? 5 (IV-10-21, K. Bowman) ; 3 9 ? (IV-ij to
13-24, Owen Bryant) ; Red Deer, 7 5 5,1? (VI-1 to 16-23, K. Bowman).

Manitoba: Aweme, 1 5,5 5? (XI-9-05), $ (III-31-04), ? (IX-29-20), $, ?
(X-16-20), all collected by N. Griddle; Cartwright, $ (XI-23-05, E. F.
Heath) ; Winnipeg, $ (no date, A. W. Hanham).

Ontario: Ottawa, $ ( X-1 7-07, C. H. Young).

Quebec: Chelsea, 5 (IV-21-23, J. McDunnough).
2S6G14 — 41 8

146 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

RemarJcs. — This species, Mamathiana^ ciniflonella^ and a few mis-
cellaneous unnamed specimens form a very complex group in which the
species are exceedingly difficult to separate. On genitalia it is difficult
to separate one from the other, only one specimen, an unnamed unique
female from the Moscow Mountains, Idaho, showing good specific
characters in the genitalia. The harpes of sciadopa are clothed with
seemingly finer hairs than those found in Jdaviathiana or cinifionella^
but this character, even though probably sufficient to distinguish
sciadopa from the others, fails in separating the latter two. The point
of origin of the transverse arm of the clasper and the comparative
lengths seem to be the safest characters for separating the males of
the two species. The length of the wings of cinifioneUa is proportion-
ately greater than in klamathiana, but I am skeptical of the value of
this character. In some long reared series of other species both long-
and short-winged forms are found. In some specimens the lengthen-
ing of the wings seems to be retarded. The venation shows some vari-
ation in all species, thus eliminating venation as a means of specific
separation.

On pattern and size it is possible, with exceptions, to distinguish the
three; sciadopa may be separated from the other two by its much
smaller size, grayer appearance, and total absence of brown or red
scales. Under this name I place with some doubt the specimens listed
above. Meyrick's description of sciadopa could actually fit any one of
two or three species before me, but because of the type locality and the
fact that Meyrick has placed sciadopa in the '■'■cini-fionella group." I
prefer to use his name instead of proposing another. An examination
of the type will be necessary to determine definitely what the name
really represents.

3. BIBARRAMBLA, new genus
PL.VTE 4, FiGTTRES 29, 30 ; Plate 9, Figxtbes 65, 65a ; Plate 18, Figube 109

Genotype. — Semioscopis alleneUa Walsingham, Trans. Amer. Ent.
Soc, vol. 10, p. 174, 1882.

Similar to Agonopterix but the second segment of the labial palpus
without furrow ; socii and uncus fused to form hood.

Head with appressed scales; side tufts spreading; antenna simple
in both sexes ; basal segment short, with pecten. Labial palpus long,
recurved, second segment much longer than third, without furrow.
Thorax with two minute crests. Abdomen not flattened. Fore wing
with arched costa ; scale tufts present ; 12 veins ; 2 and 3 stalked from
angle; 2 and 3, 4 and 5 closely approximate; 7 and 8 stalked, both to
costa; stalk of 7 and 8 approximate to 9; 11 from well before middle;
termen straight, oblique.

Hind wing as broad as fore wing ; 8 veins ; 3 and 4 connate ; 6 and 7
subparallel.

REVISION OF THE OECOPHORIDAE — CLARKE 147

Male genitalia. — With clasper. Anellus well developed, without
lateral processes. Uncus and socii fused. Gnathos a spined knob.

Female genitalia. — Genital plate strongly sclerotized, signum
present.

Remarks. — This genus is closely allied to Agonopterix.

BIBARRAMBLA ALLENELLA (Walsingham), new combination

Plate 4, Figures 29, 30 ; Plate 9, Figures 65, 65a ; Plate IS, Figure 109

Semioscopis alleiieUa Walsingham, Trans. Amer. Eat. Soc, vol. 10, p. 174, 1882. —
BuscK, in Dyar, U. S. Nat. Mus. Bull. 52, No. 5894, 1903 ; Proc. U. S. Nat.
Mus., vol. 35, p. 201, 1908.— Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 6489, 1917. — Meyeick, in Wytsman,
Genera insectorum, fasc. 180, p. 186, 1922.— McDunnough, Check list of the
Lepidoptera of Canada and the United States of America (Part 2, Micro-
lepidoptera ) , No. 8464, 1939. — Gaede, in Bryk, Lepidopterorum catalogus,
pt. 92, p. 369, 1939.

Agonopteryx allenella (Walsingham) Forbes, Cornell Univ. Agr. Exp. Stat
Memoir 68, p. 241, 1923 ; in Leonard, Cornell Univ. Agr. Exp. Stat. Memoir 101,
p. 545, 1928. — Procter, Biological survey of the Moimt Desert region: The
insect fauna, p. 273, 1938.

Labial palpus sordid whitish; second segment shaded with fuscous
on basal half and irrorated distally; slightly before apex a naiTOW,
incomplete, brownish-fuscous annulus; third segment with a spot an-
teriorly at base and a broad, supra-medial annulus brownish fuscous.
Antenna sordid whitish, narrowly annulated with fuscous; basal seg-
ment fuscous above. Head, thorax, and fore wing grayish white, suf-
fused and irrorated with fuscous ; at basal third, in cell, two fuscous
discal spots (sometimes confluent) of raised scales followed by some
ochreous scaling and with a few scales of white mixed; at the end of
cell a black-edged white discal spot of raised scales followed by some
ochreous scaling; along costa, and around termen to inner margin, a
series of fuscous spots; costa narrowly edged with pink (this pink
tint is obscure in some specimens but is strongly continued through the
cilia in others) ; cilia sordid whitish with a broad, pale grayish-fuscous
subbasal band. Hind wing pale grayish fuscous, darker apically;
cilia sordid whitish with a broad, pale grayish fuscous subbasal band.
Legs sordid whitish suffused and annulated with fuscous except at
joints and on hind tibia, the latter with pale ochreous suffusion and a
faint pink tint. Abdomen pale ochreous suffused with fuscous above.

Male genitalia. — Harpe broad ba sally, tapering to the pointed cucul-
lus, hairy ; sacculus broad, strongly sclerotized, clasper stout, dilated
distally, recurv^ed. Anellus a large sclerotized plate, pointed an-
teriorly, slightly concave posteriorly and without lateral hairy lobes.
Aedeagus long, slender, narrowly S-shaped; basally, on ventral side
a small, flat protuberance; vesica armed with minute cornuti. Vin-
culum broad, rounded, with a well-developed dorsoanterior process.

148 PROCEEDINGS OF THE NATIONAL MUSEUM vol. oo

Transtilla a narrow, weakly sclerotized band with small hairy lateral
lobes. Gnathos a spined, oval knob. Socii small, hairy, fused with
uncus to form a narrow hood.

Female genitalia: — Genital plate narrow, produced anteriorly,
strongly sclerotized ; anterior edge produced ventrally to form a small
shelf. Ostium small, elongate ; on each side a shallow, narrow cavity.
Ductus bursae membranous, slender, gradually tapering into the large
bursa copulatrix; ductus seminalis opening just before ostium. Sig-
num a small, strongly sclerotized plate with two or three strong teeth.

Alar expanse, 19-22 mm.

Ty^pe. — In the British Museum.

Type locality. — Maine ( ? ) .

Food plants. — Alniis sp. ; oak; birch.

Distribution. — Northeastern United States and eastern Canada.

United States records

District of Columbia: Washington, S (May 1902, A. Busck).

Maine: Kingsfield, $ (lO-VII-1936, no collector) ; Rangeley, 5 (12-VII-1936,

v. H. dos Passos) ; Sebec Lake, $,299 (June and July; no collector) ;

Wales, $ (21-VI-1907; uocoUector).
Maryland: Plummers Island, 3 $ $ (May and August 1903, 1919, A. Busck).
New Hampshire: Center Harbor, $ (July 27, 1902, H. G. Dyar) ; Dublin, 2 $ $

(no date; A. Busck) ; Hampton, $ (4-VII-1906, S. A. Shaw).
New Jersey: Essex County Park, $ (ll-VI-1899, W. D. Kearfott).
New York: McLean, Rhinebeck, Lond Island, etc. {teste Forbes).
Pennsylvania: New Brighton, 19 S $, 8 5 5 (May and June dates, 1901-1907,

H. D. Merrick).
Virginia: Upton, 5 (25-IV-1913, F. Johansen [Hopkins No. 9861d]).

Canadian records

Nova Scotia: Petite Riviere (July 11-18, 1935, J. McDunnough) ; S. Milford
(June 29, 30, 1934, J. McDunnough) ; White Point Beach (July 1934 and
Feb. 8, 1936 [indoor record?] J. McDunnough).

Ontario: Biseotasing (June 14-20, 1931, K. Schedl) ; Ottawa (June 3-24, 1906,
C. H. Young; July 19, 1926, C. H. Curran).

Quebec: Alcove (July 8, 1936, F. A. Urquhart) ; Brome (June 4, 1936, G. S.
Walley) ; Gracefield (June 16, 1937, O. Peck) ; Kazubazua (June 8, 1927, F. P.
Ide) ; Knowlton (Feb. 13, 1930 [indoor record?], J. McDunnough) ; Meach
Lake (July 20, C. H. Young) ; Mount Lyall (July 15, 16, 1933, W. J. Brown).

Reniarhs. — Forbes correctly removed allenella from Semioscopis, but
I do not agree with him in placing it in Agonopterix. The genus is
close to Agonopterix^ but the absence of the furrow or brush on the
labial palpus, absence of lateral lobes of the anellus, fused socii and
uncus in the male, and the unique signum of the female indicate that
this species is generically distinct.

The genotype is the only species I have seen referable to this genus.

REVISION OF THE OECOPHORIDAE — CLARKE 149

4. Genus SEMIOSCOPIS Hubner

Plate 2, Figuke 15; Plate 5, Figuee 35; Plate 11, Figures 73, 73a;
Plate 16, Figure 97

Semioscopis Hubnee, Verzeichniss bekannter Schmetterlinge, p. 402, 1826. —
Walsingham, Trans. Aruer. Ent. Soc, vol. 10, p. 174, 1882. — Riley, m Smith,
List of Lepidoptera of Boreal America, p. 99, 100, 1891. — Dyar, Can. Ent., vol.
34, p. 319-320, 1902.— BuscK, in Dyar, U. S. Nat. Mus. Bull. 52, p. 523, 1903.—
Keabfott, in Smith, List of the Lepidoptera of Boreal America, p. 114, 1903.—
BuscK, Proc. U. S. Nat. Mus., vol. 35, p. 201, 1908. — K&vrfott, in Smith,
Catalogue of the insects of New Jersey, p. 562, 1910. — Busck, Journ. Ent
Zool., Claremont, vol. 5, p. 100, 1913. — Barnes and JIcDu^nough, Check list
of the Lepidoptera of Boreal America, p. 161, 1917. — ^Meyrick, in Wytsman,
Genera insectorum, fasc. 180, p. 186, 1922. — Foubes, Cornell Univ. Agr. Exp.
Stat., Memoir 68, p. 234-244, 1923. (Genotype: Phalaenae Tortrix steinkell-
ncriana Schiffermtiller, Systematisches Verzeichniss der Schmetterlinge der
Weiuer Gegend, p. 130, 1776). — Gaede, in Bryk, Lepidopterorum catalogus,
pt. 92, p. 369, 1939.

Epiffraphia Steiphens, Catalogue of British insects, p. 304, 1872. — Grote, North
Amer. Ent., vol. 1, p. 53, 1880.— Walsingham, Trana. Amer. Ent. Soc, vol.
10, p. 174, 1882. — Beutenmtjlleb, in Smith, Catalogue of the insects of New
Jersey, p 355, 1890. — Dietz, in Smith, Catalogue of the insects of New Jersey,
p. 473, 1900. — Meyrick, in Wytsman, Genera insectorum, fasc. 180, p. 185-186,
1922. (Genotype: Tinea avellanella Hiibuer, Sammlung europiiischer Schmet-
terlinge, No. 8, fig. 27, 1796.)

Head smooth, side tufts somewhat spreading. Tongue developed
but short. Antenna shortly ciliated in male, simple in female ; basal
segment without pecten. Labial palpus moderatel}'^ long, curved ; sec-
ond segment with appressed scales which protrude at apex ; terminal
segment much shorter than second, slender, acute.

Fore wing ample, elongate ; 12 veins ; 2 and 3 approximate, connate
or stalked, 7 and 8 stalked, 7 to costa or apex, 11 from well before
middle.

Hind wing as broad as fore wing, ovate; 8 veins; 3 and 4 closely
approximate or connate.

31 ale genitalia. — Harpe elongate; sacculus frequently produced.
Anellus with lateral processes that are sometimes reduced. Transtilla
membranous; lateral lobes usually slender, digitate but greatly re-
duced. Gnathos a spined, oval knob. Socii mainly indicated by hairs.

Female genitalia. — Ductus bursae membranous or partially sclero-
tized ; signum present.

Remarks. — Meyrick ^^ separates Epigraphia from Semloscopis on the
condition of veins 2 and 3 of the fore wing (stalked or separate), yet
he ignores this same character in the case of Agonopterix and Depres-
saria except to use it to divide Depressaria into two sections. In Semio-
scopis veins 2 and 3 are much more unstable than in Agonopterix and
Depressaria., indicating that Semioscopis is in a state of flux and not

Meyrick, in Wytsman, Genera insectorum, fasc. ISO, p. 185-186, 1922.

150 PROCEEDINGS OF THE NATIONAL MUSEUM vol. »o

yet clearly definable into two genera. I have examined several hun-
dred specimens and find that veins 2 and 3 of the fore wing vary from
remote to stalked in every species, frequently differing on the two sides
of a specimen. The genitalia support the view that in the case of the
species of Semioscopis we are actually dealing with one genus. In
Agonopterix and Depressaria their separation is supported by geni-
talic evidence.

Busck ^® recognized six species in this genus. Forbes '^ later removed
alleneUa to Agonopterioo. For this species I have erected a new genus.
Two species, mcdunnoughi and hraunae^ are described as new in this
paper bringing to a total of seven the number of species for the genus.

KEY TO THE SPECIES OF SEMIOSCOPIS BASED ON
COLORATION

1. Fore wing with a dark, outwardly curved bar at end of cell 2

Fore wing without such bar at end of ceU; ground color gray or

whitish, strongly irrorated with grayish fuscous and without

large, conspicuous dark spots or dashes inornata Walsingham (p. 155)

2. Discal mark continued toward base as a straight or curved bar or

series of short lines 4

Discal mark not continued toward base 3

3. Fore wing sordid whitish marked with small contrasting black

irrorations braunae, new species (p. 159)

Fore wing whitish gray marked with small black irrorations and

generally suffused with fuscous megamicrella Dyar (p. 157)

4. Discal mark continued toward base as a strongly contrasted

curved bar 5

Discal mark not continued toward base as a curved bar 6

5. Discal bar reaching base of fore wing packardella (Clemens) (p. 151)

Discal bar sharply terminated at basal third of fore wing.

merriccella Dyar (p. 153)

6. Second segment of labial palpus suffused with blackish on distal

half; third segment contrastingly marked black and white

aurorella Dj-ar (p. 160)
Second segment of labial palpus almost wholly blackish exteri-
orly; third segment weakly mottled black and white

mcdunnoughi, new species (p. 162)

KEY TO THE SPECIES OF SEMIOSCOPIS BASED ON MALE
GENITALIA

1. Ventral margin of sacculus with prominent extension (figs. 137, 138, etc.). 2
Ventral margin of sacculus without prominent extension (fig. 135) 6

2. Extension of sacculus slender, simple, pointed (fig. 137)

inornata Walsingham (p. 155)
Extension of sacculus otherwise 3

3. Extension of sacculus bifurcate (figs. 140, 141) 4

Extension of sacculus otherwise (figs. 138, 139) 5

M Busck, A., Proc. U. S. Nat. Mus., vol. 35, p. 201, 1908.

ST Forbes, W. T. M., CorneU Univ. Agr. Exp. Stat. Memoir 68, p. 241, 1923.

REVISION OF THE OECOPHORIDAE — CLARKE 151

4. Vesica with one or two slender cornuti (fig. 141) _ merriccella Dyar (p. 153)
Vesica with cluster of slender cornuti (fig. 140)

packardella (Clemens) (p. 151)

5. Outer dorsal edge of sacculus evenly curved (fig. 138)

megamicrella Dyar (p. 157)
Outer dorsal edge of sacculus sharply angulate (fig. 139)

braunae, new species (p. 159)

6. Vesica armed with a stout, moderately short curved cornutus (fig.

135a) aurorella Dyar (p. 160)

Vesica armed with a stout, long, straight cornutus (fig. 136)

mcdunnoughi, new species (p. 162)

KEY TO THE SPECIES OF SEMIOSCOPIS BASED ON FEMALE
GENITALIA

1. Ductus bursae with a conspicuous, large sclerotized area (figs.

223, 224, 226) 2

Ductus bursae membranous or with a minute subtriangular

sclerotized area adjacent to ostium (figs. 225, 227, 228) 4

2. Ductus bursae sclerotized adjacent to bursa copulatrix (fig. 223)

aurorella Dyar (p. 160)
Ductus bursae membranous adjacent to bursa copulatrix (figs. 224, 226) — 3

3. Ostial opening as narrow as or narrower than portion of genital

plate posterior to it; loop of ductus bursae sclerotized for less

than half its length (fig. 224) bravmae, new species (p. 159)

Ostial opening wider than portion of genital plate posterior to it;
loop of ductus bursae sclerotized for more than half of its length
(fig. 226) megamicrella Dyar (p. 157)

4. Ovipositor lobes armed with hooked macrosetae (fig. 225)

inomata Walsingham (p. 155)
Ovipositor lobes without hooked macrosetae 5

5. Bursa copulatrix elongate, definitely asymmetrical; signum

minute, in posterior part of bursa (fig. 228a) _ packardella (Clemens) (p. 151)
Bursa copulatrix oval, not definitely asymmetrical; signum small,

about middle of bursa copulatrix (fig. 227) merriccella Dyar (p. 153)

SEMIOSCOPIS PACKARDELLA (Clemens)

Plate 23, Figuke 140, 140a ; Plate 40, Figubes 228, ^8a

Enicostoma pacJi-ardclla Clemens, Proc. Ent. See. Philadelphia, vol. 2, p. 125,
1S63 ; in Staiuton, The Tineina of North America, p. 231, 1872.— Bxtsok, Proc.
Ent. Soc. Washington, vol. 5, p. 214, 1903.

Semioscopis packardella (Clemens) Dyab, Can. Ent., vol. 34, p. 319, 1902. —
Busck, m Dyar, U. S. Nat. Mus., Bull. 52, No. 5893, 1903; Proc. Ent. Soc.
Washington, vol. 5, p. 214, 1903. — Keakfott, in Smith, Check List of the Lepi-
doptera of Boreal America, No. 643G, 1903.— Busck, Proc. U. S. Nat. Mus.,
vol. 35, p. 201, 1908. — Baknes and aicDuNNOUGH, Check list of the Lepidop-
tera of Boreal America, No. 6484, 1917. — Fokees, Cornell Univ. Agr. Exp.
Stat. Memoir 68, p. 244, 1923; in Leonard, Cornell Univ. Agr. Exp. Stat.
Memoir 101, p. 546, 1928.

Epigraphia packardella (Clemens) Walsingham, Trans. Amer. Ent. Soc. Phila-
delphia, vol. 10, p. 174, 1SS2. — Meyrick, in Wytsman, Genera insectorum, fasc.
180, p. 186, 1922.— McDunnough, Check list of the Lepidoptera of Canada

152 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

and the United States of America (Part 2, Microlepidoptera), No. 8459, 1939
[cited as synonym of Epigraphia steinkellneriana (Scliiftermiiller)].
Epigraphia eruditeUa Gkote, North Amer. Ent., vol. 1, p. 53, pi. 5, fig. 12, 1880.—
Meybick, in Wytsman, Genera insectornm, fasc. 180, p. 186, 1922. — Mc-
DuNNOUGH, Check list of the Lepidoptera of Canada and the United States of
America (Part 2, Microlepidoptera), No. 8459, 1939 [cited as synonym of
Epigraphia stemkelhieriana (Schiffermiiller)].

Labial palpus sordid white ; second segment overlaid with blackish
fuscous exteriorly, except at base and apex, and lightly suffused with
pink ; third segment with a small black spot at base anteriorly and a
broad, black subapical annulus. Antenna light fuscous, narrowly an-
nulated with gray. Head with the scales light brown, tipped with
white and appearing gray. Thorax gray suffused with brown ante-
riorly and with a faint carmine tint. Fore wing light shining gray
marked with numerous faint, short transverse strigulae ; at the end of
cell an outwardly curved transverse black bar continued as a longitudi-
nal curved black line to extreme base of costa ; between the bar at the
end of the cell and the costa two longitudinal brown-edged black
dashes (sometimes fused) followed on costa by a small, indistinct light
brown shade; costa marked with poorly defined, light-brown spots
and dashes and narrowly edged with pink to slightly beyond middle ;
around termen to inner margin a series of small black spots; cilia
pale gray with a light-brown suffusion outwardly. Hind wing pale,
shining gray, darker apically; cilia paler, with narrow light brown
bands. Legs pale ochreous-white overlaid exteriorly, except joints
and hind tibiae, with blackish fuscous; pale areas with a faint pink
tint. Abdomen sordid ochreous with a slight fuscous suffusion
beneath.

Male genitalia. — Very similar to memccella^ differing primarily in
characters of the aedeagus.

Harpe long, rather narrow, slightly wider before cucullus ; costa and
cucullus sclerotized, area between membranous; cucullus narrow,
rounded ; sacculus produced to form a large forked process. Anellus
long, strongly curved posteriorly to form a semicylinder ; basolateral
lobes reduced to slight swellings, chiefly indicated by hairs. Aedeagus
long, sharply curved, pointed; vesica armed with an elongate patch
of small straight cornuti. Vinculum rounded. Lobes of transtilla
long, digitate, hairy, fused with anellus at base. Tegumen roimded.
Socii reduced to small hairy lobes.

Female genitalia. — Much as in merriccella but differing by the
longer bursa and minute signum. Genital plate narrow, broadened
at middle to form the cup-shaped ostium. Ductus bursae a long,
slender, convoluted tube with a small, subtriangular sclerotized area
near ostium; inception of ductus seminalis at the sclerotized part.

REVISION OF THE OECOPHORIDAE — CLARKE 153

Bursa copulatrix large, definitely asymmetrical, elongate; sigiium a
minute toothed plate in posterior end of bursa.

Alar expanse, 21-28 mm.

Type. — In the Acadamy of Natural Sciences of Philadelphia.

Type locality. — Not stated; probably Massachusetts.

Food plant. — Unknown.

Distribution. — Northeastern United States and eastern Canada.

United States records

Maine: Bar Harbor, $ (2-V-1936, A. E. Brower).

Michigan: No specific locality, $, 9 (male without date; 2, 14-V-1885,

Gillette).
New Jersey: Montclair, S, 9 (13-V-1900, l-V-1899, W. D. Kearfott).
New York: Ithaca, Big Indian Valley, Albany (ace. Forbes).
Ohio: Cincinnati, 2 $ S, 9 (18-IV-1911, A. F. Braun).
Pennsylvania: Franconia, Montgomery County, 2 (no date or collector) ; New

Brighton, 3$S, 35$ (April 1902-1908, H. D. Merrick); Oak Station,

Allegheny County, 76$, 52$ (April and May dates, 1902-1915, Fred

MarlofE) .

Canadian records

Manitoba: Cartwright, 2 (no date; E. F. Heath).

Quebec: Burbridge (May 25, 1937, F. A. Urquhart) ; Meach Lake (April 27, 1899,
C. H. Young).
Remarks. — This species will probably be found throughout the
mid western United States and Canada as far west as British
Columbia.

SEMIOSCOPIS MERRICCELLA Dyar

Plate 23, Figitee 141 ; Plate 40, Figure 227

Semioscopis merriccella Dyab, Can. Ent., vol. 34, p. 319, 1902. — Keaefott, in
Smith, Check list of the Lepidoptera of Boreal America, No. 6437, 1903.—
BuscK, Proc. U. S. Nat. Mus., vol. 35, p. 201, 1908.— Barnes and McDun-
NOUGH, Check list of the Lepidoptera of Boreal America, No. 64S5, 1917. —
Forces, Cornell Univ. Agr. Exp. Stat., Memoir 68, p. 224, 1923 ; in Leonard,
Cornell Univ. Agr. Exp. Stat., IMemoir 101, p. 546, 1928.

Epigraphia merrickella Meyiiick, in Wytsman, Genera insectorum, fasc. 180,
p. 186, 1922.— McDunnough, Check list of the Lepidoptera of Canada and
the United States of America (Part 2, Microlepidoptera), No. 8459, 1939
[amended spelling for S. mcrrUccUa Dyar and cited as synonym of
Epigraphia steinkellneriana (SchifEermiiller) ].

Labial palpus white ; second segment suffused with blackish fuscous
exteriorly ; third segment with a fuscous spot at base anteriorly and a
broad black subapical annulus. Antemia with basal segment blackish
fuscous ; remainder light fuscous, narrowly and indistinctly annulated
with gray. Head and thorax gray, the latter rather strongly suffused
with fuscous. Fore wing light, shining gray lightly shaded and
strigulated with brown ; costa, from base to middle of wing, lighter ;
at the end of cell an outwardly curved blackish-fuscous bar followed

154 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

by a rapidly fading and spreading brownish shade; between the discal
bar and costa a short, poorly defined blackish-fuscous bar followed by
a blackish-fuscous spot before apex; before the discal bar, in cell, a
conspicuous, broad, curved, longitudinal, blackish-fuscous bar, which
does not reach base of wing but is preceded by a similarly colored
narrow dash and one or two spots from base of wing at costa ; from
apex, around termen, a series of blackish-fuscous spots; cilia pale
gray lightly suffused with brown. Hind wing shining gray with a
fine terminal brown line ; cilia lighter, shining, with narrow subbasal
and two subterminal, faint brown bands. Legs shining creamy white
strongly overlaid with blackish fuscous except at joints and on hind
tibiae. Abdomen sordid ochreous, faintly suffused with fuscous
beneath.

Male genitalia. — Harpe long, narrow, slightly wider before cucul-
lus; clasper absent, sacculus produced as a forked process; cucullus
narrow, bluntly pointed; costa and sacculus sclerotized, with area
between them membranous. Anellus a broad plate produced pos-
teriorly as a semicylinder ; basolateral lobes indicated by a few hairs.
Aedeagus long, slender, curved, terminating in a slender, curved
point; vesica armed with two, long, slender cornuti, one about half
the length of the other. Vinculum rounded. Lobes of transtilla long,
digitate, hairy, and fused with anellus at base. Tegumen rounded.
Socii mainly indicated by a few hairs.

Female genitalia. — Genital plate narrow. Ostium cup-shaped.
Ductus bursae a long convoluted tube with a small subtriangular
sclerotized area near ostium ; inception of ductus seminalis just before
ostium. Bursa copulatrix large, oval; signum a small toothed plate
about middle of bursa copulatrix.

Alar expanse, 24-31 mm.

Type. — In the United States National Museum.

Type locality^ — New Brighton, Pa.

Food plant. — Unknown.

Distribution. — Northeastern United States and Canada to western
British Columbia.

United States records

Maine: Bar Harbor, $ (13-V-1937, A. E. Brower).
New Hampshire: Hampton, 5 (l-IV-1907, S. A. Shaw).
New York: Big Indian Valley, Albany (ace. Forbes).

Pennsylvania: New Brighton, 12 S S, 4 $ $ (March and April dates, 1902-
1904, H. D. Merrick).

Canadian records

British Cohimbia: Salmon Arm, $ (16-V-1922, W. R Buckell).
Manitoba: Aweme, $ (15-V-1905, N. Griddle) ; Cartwright, $ (no date, E. F.
Heath).

REVISION OF THE OECOPHORIDAE — CLARKE 155

Remarks. — This species is very much like packardella but may be
distinguished from it by the broken discal bar, its more striate
appearance, and usually larger size. The genitalia of the two species
are strikingly similar, but they can be distinguished by the characters
given in the keys.

The single male from British Columbia is worn but undoubtedly
belongs to this species. The distribution of the species indicates this
specimen might be mislabeled or only a stray although further col-
lecting may reveal that the species is well established in the far West.

SEMIOSCOPIS INORNATA Walsinghara

Plate 22, Figuees 137, 137a ; Plate 40, Figure 225

Semioscopis mornata Walsingham, Trans. Amer. Ent. Soc, vol. 10, p. 174,
1S82.— Dyae, Can. Ent., vol. 34, p. 320, 1902.— Busck, in Dyar, U. S. Nat.
Mus, Bull. 52, No. 5895, 1903. — Keaefott, in Smith, Check list of the Lepidop-
tera of Boreal America, No. 6440, 1903.— Busck, Proc. U. S. Nat. Mus.,
vol. 35, p. 201, 1908. — Baenes and McDunnough, Check list of the Lepidop-
tera of Boreal America, No. 6488, 1917.— Meyeick, in Wytsman, Genera
insectorum, fasc. ISO, p. 186, 1922.— Fokbes, Cornell Univ. Agr. Exp. Stat.
Memoir 68, p. 244, 1923; in Leonard, Cornell Univ. Agr. Exp. Stat.
Memoir 101, p. 546, 1928. — McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera), No.
8460, 1939.— Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 372, 1939.

Semioscopis inornatella Busck, Proc. U. S. Nat. Mus., vol. 35, p. 201, 1908.

Labial palpus white ; second segment black externally, especially in
distal half except extreme apex; third segment with a subbasal spot
and a subapical annulus black. Antenna blackish fuscous and
broadly amiulated with white on basal third, the color fading to a
light yellowish fuscous, and the annulations disappearing, apically.
Head and thorax sordid white suffused and clouded with grayish
fuscous. Fore wing sordid white, profusely irrorated and suffused
with grayish fuscous; outer discal mark reduced to two indistinct
grayish- fuscous spots edged with whitish scales; at basal third two
blackish-fuscous spots, one above the other; costa spotted with gray-
ish fuscous especially before apex; around termen a series of poorly
defined blackish-fuscous spots ; cilia sordid white with basal and nar-
row subterminal bands light grayish fuscous. Hind wing shining
grayish fuscous, cilia sordid white with subbasal and narrow subter-
minal bands light grayish fuscous. Fore and middle legs sordid white
strongly overlaid with blackish fuscous except at joints; posterior
leg pale ochreous- white ; femur and tarsus suffused with fuscous.
Abdomen dull ochreous above, whitish ochreous beneath with a poorly
defined fuscous line on each side.

Male genitalia. — Harpe broad basally, narrowed beyond the ventral
extension of the sacculus, sacculus moderately and narrowly sclero-

156 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

tized and produced as a long bluntly pointed process; cucullus nar-
row, rounded ; clasper absent. Anellus an elongate sclerotized plate,
convex laterally beyond middle and basolaterally produced to form
small, hairy lobes; posterior edge deeply and broadly V-shaped.
Aedeagus stout, long, twisted, terminating in a blunt point; vesica
armed with a single, stout, forked cornutus. Vinculum rounded.
Lobes of transtilla rather thick, clothed with thick, short hairs apic-
ally and fused to harpe. Tegumen rounded. Socii poorly developed,
moderately hairy lobes.

Female genltal'm. — Lobes of ovipostor with conspicuous, hooked
macrosetae. Genital plate narrow, especially so at ostium where the
sclerotized portion is no more than the narrow anterior edge of the
ostium. Ductus bursae a long convoluted membranous tube; in-
ception of ductus seminalis well before ostium. Bursa copulatrix
large, oval; signum a small toothed plate.

Alar expanse, 25-35 mm.

Type. — In the United States National Museum.

Type locality. — Orono, Maine?.

Food plant. — Unknown.

Distribution. — Northeastern United States, Canada, west to British
Columbia and Alaska. The species probably will be found in some
of the Northwestern States.

United States and Alaskan records

Alaska: Rampart, 6 $ $ (no date or collector).

Illinois: Putnam County, S (8-VI-1936, Murray O. Glenn).

Maine: Orono, 3 $ $ (April; no collector).

New Jersey: Essex County Park, $, 9 (29-III-1905, 15-V-1906, W. D.

Kearfott).
New York: Ithaca, Karner (ace. Forbes).
Pennsylvania: New Brighton, 5 ( 25-1 V-1908, H. D. Merrick).

Canadian records

Alberta: Calgary (April 24, 190S, F. H. Wolley-Dod) ; Edmonton, 5 S $
(25-IV-1924, R. Aitcheson ; IV-26-29-1924, Owen Bryant ; 19-IV-28-IV-1921,
no collector) ; Red Deer, $ (lS-IV-1903, no collector).

British Columbia: Victoria, S $ $ (29-III-1920, 7-IV-1922, W. Downes).

Manitoba: Aweme, 7 $ S , 2 ? $ (April 3 to May 2, 1904-1905, N. Criddle; also
April 14 to May 12, 1920-1924) ; Cartwright, S $ $ (3 $ 5 -11-13-IV-1913,
no collector; 5 $ S, 7-IV-1905, E. F. Heath) ; Miniota, 9 (18-IV-1903, no
collector).

Ontario: Bells Corners (25-IV-1937, G. S. Walley) ; Biscotasing (16-V-1931,
K. Schedl) ; Constance Bay (26-IV-1935, J. McDunnough) ; Ingersoll (1-V-
1934, G. S. Walley); Merivale (2-V-1934, W. J. Brown; l-V-1936, G. S.
Walley) ; Ottawa, (21-IV-1906, C. H. Young; 18-IV-1927, G. S. Walley);
Port Hope (H. Bowers) ; Toronto (no date or collector).

Quebec: Aylmer (3-V-1923, C. H. Curran) ; Wright (19-V-1932, J. McDun-
nough).

REVISION OF THE OECOPHORIDAE — CLARKE 157

Remarks. — The British Columbia specimens, although larger and
more contrastingly marked, are otherwise indistinguishable from the
average specimens.

Walsingham did not state the type locality when he described the
species, but it is presumably Orono, Maine ; several specimens before
me with serial numbers corresponding to that of the type bear that
locality label. The type label bears an "O" ( = Orono?) in Walsing-
ham's handwriting.

The hooked macrosetae of this species are found on the European
strigidana also. These and other similarities lead me to believe that
inomata is only a race of strigulana. For the time being, however,
I am retaining inomata as a distinct species.

SEMIOSCOPIS MEGAMICRELLA Dyar

Plate 23, Figukes 138, 138a ; Plate 40, Figube 226

Semioscopis megamicrella Dyak, Can. Eiit., vol. 34, p. 820, 1902.— Ke.\efott, in
Smith, Check list of the Lepidoptera of Boreal America, No. G439, 1903. —
BuBCK, Proc. U. S. Nat. Mus., vol. 35, p. 201, 1908.— Barnes and Mc-
DiTNNOUGH, Check list of the Lepidoptera of Boreal America, No. 6487, 1917. —
Meyrick, in Wytsman, Genera insectorum, fasc. 180, p. 186, 1022. — Forbes,
Cornell Univ. Agi-. Exp. Stat. Memoir 68, p. 244, 1923 ; m Leonard, Cornell
Univ. Agr. Esp. Stat. Memoir 101, p. 546, 1928. — McDunnough, Check list
of the Lepidoptera of Canada and the United States of America (Part 2,
Microlepidoptera), No. S4G1, 1939. — Gaede, in Bryk, Lepidopterorum cata-
logus, pt. 92, p. 372, 1939.

Labial palpus white; second segment blackish fuscous in apical
two-thirds except for a narrow longitudinal area inwardly ; third seg-
ment with a minute subbasal spot anteriorly and subapical annulus
blackish fuscous. Antenna with basal segment blackish fuscous
above, whitish beneath; remainder light fuscous, narrowly and in-
distinctly annulated with grayish fuscous. Head, thorax, and ground
color of fore wing whitish gray irrorated with black and brown scales
and lightly shaded with fuscous ; at the end of cell a blackish-fuscous,
outwardly curved, crescentic bar (sometimes inconspicuous or broken
into a series of dots) preceded by some white scaling; at basal third
a pair of superposed blackish-fuscous dots ; in some specimens a longi-
tudinal dash of the same color; costa narrowly edged with pink (this
absent in some specimens) and spotted with blackish scales; a sub-
marginal and terminal row of blackish-fuscous spots, the former usu-
ally poorly defined ; cilia light gray, with pale fuscous subbasal and
apical bands. Hind wing shining pale grayish fuscous, cilia some-
what lighter, with pale fuscous subbasal and apical bands. Legs
sordid whitish overlaid with blackish fuscous except at joints and
on hind tibiae. Abdomen light fuscous.

158 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Male genitalia. — Harpe short, broad, sacculus produced as a broad
truncate sclerotized process, cucullus bluntly pointed ; clasper a small
tooth about middle of harpe. Anellus a broad oval plate narrowly
produced posteriorly, basolateral lobes small, sparsely hairy.
Aedeagus long, slender, curved and terminating in a long, slender,
curved point ; vesica armed with a slender, curved cornutus. Vinculum
narrowly rounded. Lobes of transtilla short, broad, hairy, and closely
attached to the harpe. Tegmnen rounded. Socii moderately well
developed hairy lobes.

Female genitalia. — Genital plate moderately broad at ostium, nar-
rower laterally. Ostium broad with a strongly sclerotized anterior
edge. Ductus bursae convoluted, the loop of the ductus sclerotized
for more than half its length ; ostium preceded by a broadly dilated,
sclerotized part of ductus bursae; before this a narrow membranous
band, anterior to which is a longer sclerotized portion of the ductus
bursae ; inception of ductus seminalis at the dorsoposterior edge of the
latter sclerotized part. Bursa copulatrix large oval with a moderately
large toothed signum.

Alar expanse, 16-27 mm.

Type. — In the United States National Museum.

Type locality. — New Brighton, Pa.

Food plant. — Unknown.

Distribution. — Northeastern United States and Canada westward to
Idaho and Alberta.

United States records

Idaho: Wallace, $ ( 17-1 V-23, Otto Huelleman).

Massachusetts: Foi'est Hills, $ (25-III-1930, no collector) ; Newton, $
(25-IV-1909, William ReifE) ; Winchendon, $ (12-IV-1902, no collector).

New York: Ithaca, Long Island (ace. Forbes).

Pennsylvania: New Brighton, 72 S $ , 15 $ 9 (March 1902-1907, H. D. Mer-
rick) ; Oak Station, Allegheny County, $, $ (4-IV-1909, 23-III-1907, Fred
Marloff ) .

Canadian records

Alberta: Edmonton, $, $ (2-V-1924, Owen Bryant).

Quebec: Aylmer, $ (9-V-1932, W. J. Brown) ; Meach Lake (April 28, 1903,
O. H. Young).

Remarks. — In this species there is unusually great variation in the
size of the specimens. This variation is not associated with sex or
locality.

The Idaho and Alberta specimens are considerably darker than
the types, but I can find no other external differences, and the genitalia
are identical.

REVISION OF THE OECOPHORIDAE — CLARKE 159

SEMIOSCOPIS BRAUNAE, new species

Plate 23, Figukes 139, 139a ; Plate 40, Figure 224

Semioscopis megamicrcUa Forbes (not Dyar), in part, Cornell Univ. Agr. Exp.
Stat. Memoir 68, p. 244, 1923.

Labial palpus white, second segment suffused with blackish fuscous
except for a narrow line inwardly and at tip ; third segment with sub-
basal spot anteriorly and a subterminal annulus black. Antenna
fuscous, spotted with white above on basal fourth. Head, thorax,
and ground color of fore wing white ; the head sordid, thorax strongly
infused with grayish brown and with posterodorsal crests blackish
fuscous, and the fore wing irrorated with blackish fuscous and black ;
at the end of cell an outwardly curved crescentic discal bar and at
basal third two spots, one above the other, black ; along costa and around
termen a series of blackish-fuscous spots ; before apex, subcostally, one
or two conspicuous black spots; cilia pale grayish white, with light
brown subbasal and subterminal bands. Hind wing shining grayish
fuscous, with a narrow fuscous terminal line; cilia pale grayish white,
with light fuscous basal band and pale brownish terminal suffusion.
Legs light creamy white strongly suffused and overlaid with blackish
fuscous except at joints and on hind tibiae. Abdomen pale shining
ochreous- fuscous, suffused with fuscous beneath.

Male genitalia. — Harpe broad, short; cucullus long, narrowly
rounded; sacculus narrow basally and with a broad, strongly sclero-
tized expansion distally; outer dorsal margin angulate; clasper mi-
nute, pointed. Anellus a moderately small oval plate, somewhat nar-
rowed posteriorly ; posterior edge convex ; lateral lobes absent or indi-
cated by a few hairs. Aedeagus long, slender, narrowed at middle,
curved and terminating in a slender, curved point ; vesica armed with a
single slender, pointed cornutus. Vinculum narrowly rounded.
Transtilla membranous; lateral lobes mainly indicated by a few
hairs. Tegumen rounded. Socii small fleshy lobes.

Female genitalia, — Genital plate narrow, slightly broader posterior
to ostium. Ostium as narrow as or narrower than portion of genital
plate posterior to it, bordered anteriorly by a strongly sclerotized
and dilated portion of the ductus bursae. Ductus bursae membra-
nous except for less than half the length of a posterior loop, and the
dilated part before the ostium; inception of ductus seminalis at
anterior edge of dilated section. Bursa copulatrix large oval; sig-
num a moderately large, narrow, transverse, toothed plate about
middle of bursa.

Alar expanse, 22-26 mm.

Type.—[]. S. N. M. No. 53118.

160 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Type locality. — Cincinnati, Ohio.

Food 'plant. — Unknown,

Remarks. — Described from $ type and 5 2 paratypes all from the
type locality (March and April dates, 1904^1908, Dr. Annette F.
Braun). Paratypes in the Canadian National and Dr. A. F. Braun
collections.

This species is much like inegamicreXla but is lighter in color, lacks
the fuscous or brown suffusion, and is more contrastingly and more
sparsely marked than that species. In male genitalia it differs from
megamwreTl<i in the stouter expansion and the angulate outer dorsal
margin of the sacculus. In the female the ostium is smaller as is
the posterior dilated portion of the ductus bursae adjacent to it.
The posterior loop of the ductus bursae is less strongly sclerotized.

This is the species placed under megamicreUa by Forbes.

I take pleasure in naming this species after Dr. Annette F. Braun,
who collected the type series.

SEMIOSCOPIS AURORELLA Dyar

Plate 22, Figure 135, 135a; Plate 40, Figure 223

Semioscopis aurorella Dyar, Can. Ent., vol. 34, p. 319, 1902.— Keabfott, in
Smith, Check list of the Lepidoptera of Boreal America, No. 6438, 190®.—
BuscK, Proc. U. S. Nat. Mus., vol. 35, p. 201, 1908.— Barnes and
JilcDuNNouGH, Check list of the Lepidoptera of Boreal America, No. 6486,
1917. — Meyrick, in Wytsman, Genera insectorum, fasc. ISO, p. 186, 1922. —
Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 244, 1923 ; in Leonard,
Cornell Univ. Agr. Exp. Stat. Memoir 101, p. 546, 1928. — McDunnough,
Check list of the Lepidoptera of Canada and the United States of America
(Part 2, Microlepidoptera), No. 8463, 19.39. — Gaede, in Bryk, Lepidopter-
orum catalogus, pt. 92, p. 370, 1939.

Labial palpus white with a faint roseate tinge; second segment
overlaid with blackish fuscous exteriorly on distal half except at
apex; third segment with a black spot anteriorly above base and a
black subapical annidus. Antenna light fuscous, with poorly defined,
narrow, grayish annulations. Head and thorax gray, the former
more whitish and with some brown intermixed, the latter suffused
with fuscous and with fuscous posterior tufts. Fore wing shining
gi-ay suffused with brown, with a roseate tinge and variously marked
with blackish fuscous; at the end of cell an outwardly curved
blackish-fuscous bar preceded by a whitish or light-gray patch; in
cell a pair of superposed dashes, sometimes fused to form an elongate
V-shaped mark ; a short basal subcostal dash and a basal dash follow-
ing vein Ic to slightly beyond basal third, blackish fuscous; a sub-
terminal row of five more or less well defined dashes between the
veins and a series of spots from costa around termen, blackish
fuscous; cilia pale grayish fuscous, darker apically and with a

REVISION OF THE OECOPHORIDAE — CLARKE 161

roseate tinge. Hind wing shining grayish fuscous, darker toward
margin and with a fine fuscous terminal line; cilia pale grayish
fuscous, with subbasal and terminal bands fuscous. Legs yellowish
white heavily overlaid with blackish fuscous except at joints and on
hind tibiae. Abdomen light fuscous narrowly annulated with pale
yellowish fuscous at posterior ends of segments ; anal tuft yellowish
fuscous.

Male genitalia. — Harpe narrow, long ; costa gently and evenly con-
cave; cucullus bluntly pointed; sacculus narrowly but strongly
sclerotized; clasper long, slender, curved inwardly, attaining or
slightly exceeding costa. Anellus a subrectangular plate with convex
lateral edges; posterior edge concave, basolateral lobes indicated by
a few hairs. Aedeagus stout, curved, moderately long, bluntly
pointed; vesica armed with a single stout, curved cornutus. Vincu-
lum broadly rounded. Transtilla indicated only by a weak mem-
brane; lateral lobes absent. Tegumen rounded. Socii moderately
well developed hairy lobes.

Female genitalia. — Genital plate moderately broad, strongly sclero-
tized. Ostium large, oval, with a narrow, strongly sclerotized an-
terior edge. Ductus bursae membranous in posterior half, strongly
sclerotized, convoluted and broadened in anterior half; inception of
ductus seminalis well before ostium. Bursa copulatrix large, round;
signum a small toothed plate.

Alar expanse, 24—31 mm.

Type. — In the United States National Museum.

Type locality. — New Brighton, Pa.

Food plant. — Unknown.

Distribution. — Northeastern United States and eastern Canada.

United States records

New York: Ithaca, Big Indian Valley (ace. Forbes).

Ohio: Cincinnati, 2 $ (27-III-1903, A. F. Braun).

Pennsylvania: New Brighton, 20 5 (J , 16 $ $ (March and April dates, 1901-

1905, H. D. Merrick); Oak Station, Allegheny County, 9 5 5 , 10 9 5

(March and April dates, 190S-1914, Fred Marloff).

Canadian records

Alberta: Red Deer, 4 S S, 2 5 $ (16-24 April, 1922-1923, K. Bowman).
Manitoba: Aweme, $ (3-V-1924, N. Griddle) ; Cartwright, $ (no date, E. F.

Heath).
Ontario: Constance Bay, 2 $$ (26-IV-1935. W. J. Brown; 26-IV-193.'>,

J. McDunnough) ; Ottawa, 5 9 9 (26-IV-1909. l-V-1906, C. H. Young).

Rcnmrks. — The specimens from Canada, which I have before me,
are considerably darker and, for the most part, are less conspicu-
ously marked than the specimens from the United States, owing to

286614 — 41 9

162 PROCEEDINGS OF THE NATIONAL MUSEUM tol. 90

a strong brownish or fuscous suffusion which appears to be char-
acteristic of the northern specimens. The genitalia of the two forms
are identical.

SEMIOSCOPIS MCDUNNOUGHI. new species

Plate 22, Figuke 136

Semioscopis aurorella Blackmore (not Dyar), Kept. Prov. Mus. Nat. Hist.
British Columbia for 1921, p. 32, pi. 4, 1922.

Similar to aurorella but slightly broader winged, less distinctly
marked and with a longer, straight cornutus.

Labial palpus white with a carmine tinge; second segment
strongly overlaid with blackish fuscous exteriorly for its whole
length; third segment with poorly defined blackish-fuscous subbasal
spot and subapical annulus. Head whitish with some brown scales.
Thorax cinereous strongly infused with fuscous and with median
longitudinal streak and tufts blackish fuscous; tegula blackish fus-
cous, white-edged. Fore wing shining gray suflPused with brown and
with a roseate tinge; discal bar, reduced to a poorly defined cres-
centic dash or a spot, a dash in cell, a basal dash following vein Ic,
and a basal shade, blackish fuscous; along costa and around termen
a well-defined series of blackish-fuscous spots; cilia light fuscous
with a median grayish band and a whitish pink-tinged terminal
line. Hind wing grayish fuscous, darker toward margins and with
a narrow fuscous terminal line; cilia pale yellowish fuscous with
basal and subterminal fuscous bands. Legs yellowish white strongly
overlaid with blackish fuscous except at joints and on hind tibiae
where the lighter ground color is carmine tinted. Abdomen
ochreous-fuscous lightly suffused with fuscous beneath.

Male genitalia. — Harpe narrow, long; cucullus rounded; clasper
long, reaching almost or quite to costa, slightly curved, pointed;
sacculus narrow, strongly sclerotized. Anellus a broad, oval plate,
narrowed, produced, and curved posteriorly to form a semicylindrical
articulation for the aedeagus; lateral lobes absent or at most only
indicated by a few hairs. Aedeagus stout, moderately long; apex
flattened, rounded; vesica armed with a single long, stout, straight,
sharply pointed cornutus. Vinculum broadly rounded. Transtilla
membranous ; lateral lobes vestigial, mainly indicated by a few hairs.
Tegumen bluntly pointed. Socii small hairy lobes.

Alar expanse, 28-29 mm.

Type.—U. S. N. M. No. 53117.

Type locality. — Bellingham, Wash.

Food plant. — ^Unknown.

Remarks. — Described from the ^ type (Bellingham, Wash.,
14-in-1923, J. F. G. Clarke, ¥0. 573) and 1 5 paratype (Fraser

REVISION OF THE OECOPHORIDAE — CLARKE 163

Mills, British Columbia, 14^111-1921, L. E. Marmont), the latter
in the Canadian National collection.

This species is much like aurorella and is the one reported by
Blackmore from British Columbia by that name. It can be dis-
tinguished from aurorella by the poorly defined markings of the
third segment and the blackish-fuscous shading of the second seg-
ment of the labial palpus, the absence or only faint indication of the
subterminal row of spots on the fore wing, the presence of a rather
strong basal blackish-fuscous suffusion and the long straight
3ornutus.

There is another known specimen in the collection of the Uni-
versity of British Columbia, the one figured by Blackmore, from
Millardville, British Columbia.

I have seen no females certainly referable to this species. There
are before me, however, two females from southeastern Washington
that may belong here, although they are considerably smaller and
probably represent an undescribed species. Females must be ob-
tained from the coastal region before these two can be definitely
placed here or described as new.

5. Genus DEPRESSARIA Haworth

Plate 2, Figuee 11 ; Plate 6, Figxjre 48 ; Plate 10, Figures 68, 68a ; Plate 17,
Figure 102

Depressaria Haworth, Lepidoptera Britannica, p. 505, 1812. — Chambers (in
part). Can. Ent., vol. 4, p. 91, 146-148, 1872.— V^.vlsingham, (in part),
Proc. Zool. Soc. London, 1881, p. 311-319; Trans. Amer. Ent. Soc, vol. 10,
p. 175, 1882. — BEUTBrNMXJTXER (in part), In Smith, Catalogue of tlie insects
of New Jersey, p. 355, 1890. — Riley (in part), in Smith, List of the
Lepidoptera of Boreal America, p. 99, 1891. — Dietz (in part), in. Smith,
Catalogue of the insects of New Jersey, p. 474, 1900. — Busck (in part),
Proc. U. S. Nat. IMus., vol. 24, p. 731-749, 1902; in Dyar, U. S. Nat. Mus.
Bull. 52, p. 520-522, 1903.— Kearfott (in part), in Smith, List of the
Lepidoptera of Boreal America, p. 114, 1903. — Busck (in part), Proc. U. S.
Nat. Mus., vol. 27, p. 763-766, 1904.— Dyar (in part), Proc. U. S. Nat.
Mus., vol. 27, p. 934, 1904. — Anderson (in part). Catalogue of British
Columbia Lepidoptera, p. 54, 1904. — Busck (in part), Proc. Ent. Soc.
Washington, vol. 9, p. 88-91, 1908.— Kearfott, in Smith, Catalogue of the
insects of New Jersey, p. 561, 1910. — Barnes and McDunnough, Check list of
the Lepidoptera of Boreal Axoerica, p. 161, 1917. — Meyrick (in part), Ex-
otic Microlepidoptera, vol. 2, p. 223, 1918; p. 315, 1920; pp. 391-^92, 1921;
p. 513, 1922; m Wytsman (in part), Genera insectorum, fasc. 180, pp. 169-
177, 1922.— Forbes, Cornell Univ. Agr. Exp. Stat., Memoir OS, p. 241-243,
1923.— Pierce, The genitalia of the tineid families of the Lepidoptera of
the British Islands, p. 33, 1935. — McDunnough, Check list of the Lepidoptera
of Canada and the United States of America (Part 2, Microlepidoptera),
p. 78, 1939.— Gaede, in Bryk, Lepidopterornm catalogus, pt. 92, p. 269, 1939.
(Genotype: Plialaena Tortrix lieracliana Linnaeus, Systema naturae, ed. 10,
p. 532, No. 225, 1758.)

164 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Biganorosis Waui^ngken, Ent. Tidskr., vol. 2, p. 94, 1881. (Genotype: Phalaetm
Tortrix Iwracliana Linnaeus, Systema naturae, ed. 10, p. 532, No. 225, 1758.)

Head with appressed scales ; tongue developed ; antenna simple or
moderately ciliated; basal segment elongate, with pecten. Labial
palpus long, recurved ; second segment reaching base of antenna, with
rough, projecting scales and furrowed beneath; terminal segment
shorter than, or rarely as long as, second, acute. Thorax smooth or
crested. Abdomen flattened.

Fore wing with 12 veins; 2 and 3 separate, 7 and 8 stalked; 7 to
costa or apex, 11 from middle.

Hind wing as broad as or broader than the fore wing; 8 veins, 3
and 4 connate or short stalked; 6 and 7 subparallel.

3Iale genitalia. — Harpe with or without clasper and with or with-
out process from base of sacculus. Anellus a well sclerotized plate
without long lateral processes. Aedeagus stout with or without
cornuti. Gnathos an oval, spined knob. Socii well developed.

Female genitalia. — Genital plate strongly sclerotized ; ductus bursae
frequently sclerotized for a considerable portion of its length; signum
present (in all species seen) though frequently weak.

Larva. — Ninth abdominal segment with setae I and II well sep-
arated; seta VI not on the same pinaculum with IV and V, remote
from VII. Setal group VII bisetose or trisetose on first abdominal
segment, bisetose on seventh and eighth abdominal segments,-^ and
unisetose on ninth abdominal segment. Ocelli normal. Submentum
without pit.

Group A : Setal group VII on first abdominal segment &^setose.
licracliana
juliella
dracunouU

Group B : Setal group VII on first abdominal segment ^Wsetose.
multifidae
leptotaeniae
angustati

Pupa. — Pubescent. Prothoracic femora exposed. Labial palpi not
exposed. Cremaster absent.

Remarks. — The separation of Depressaria from Agonopterix is dis-
cussed under the latter genus.

In this genus there are five distinct species groups. The first, con-
sisting of atrostrigella and dracuncul't., and probably palousella (males
not known), is characterized by the process from the costa of the
harpe in the male and the broad, somewhat dilated sclerotized band
in the ductus bursae of the female. The second group consists of

** Rarely unisetose on eigbth segment and if so trisetose on first.

REVISION OF THE OECOPHORIDAE — CLARKE 165

jidiella^ eleanorae^ heracliana, and cinereocostella with the European
nervosa. These have a strong basal process from the sacculus and no
clasper in the male and an elongated, sclerotized section of the ductus
bursae posteriorly. We have two species in the third group, artemisi-
ella and alienella, in which the clasper is present but the basal process
of the sacculus is absent in the male and the ductus bursae is wholly-
membranous. The fourth group, consisting of the togata-angustati-
multi-fldae complex, forms a perplexing assemblage of species charac-
terized by their similarity in coloration, by the peculiar spined proc-
ess from the base of the sacculus in the male, and by the frequently
present sclerites anterior to the genital plate in the female. The fifth
and final group is composed of three broad-winged forms, maculatella^
'betulelJa, and groteUa. In the males the divided clasper, extending
beyond the ventral margin of the harpe, and the spiraled ductus
bursae of the female are characteristic.

Although rather a large number of species have been reared, the
larvae of only six were available for study. By a comparison of the
larval characters it will be seen that the first two groups (Group A
under larvae) are closely similar. The fourth group (Group B under
larva) suggests generic separation, and I am inclined to believe that
this group does represent a distinct genus. Likewise I believe that
the fifth group should be given a separate generic designation, which
would be consistent with other generic separations in this paper.
Nevertheless I prefer to wait for the collection of more larvae before
making the separations which are suggested or before concluding that
the species all belong to one genus and merely represent species groups.

KEY TO THE SPECIES OF DEPRESSARIA BASED ON
COLORATION

1. Thorax white alienella Busck (p. 180)

Thorax otherwise 2

2. Fore wing distinctly reddish 3

Fore wing otherwise 5

3. Costa contrastingly lighter than ground color of fore wing

cinereocostella Clemens (p. 171)
Costa not contrastingly lighter than ground color of fore wing 4

4. Alar expanse 21 mm. or more juliella ^9 Busck (p. 176)

Alar expanse 20 mm. or less eleanorae, new species (p. 178)

5. Ventral side of abdomen with two distinct rows of blackish spots

or lines 6

Ventral side of abdomen without such spots or lines 8

6. Fore wing with light discal spot at end of cell 7

Fore wing without such spot heracliana (Linnaeus) (p. 173)

29 1 have examined over 150 specimens ot juliella none of which measures as small as 20 mm. The larger
specimens of eleanorae approach the smaller specimens ot juliella in size but noce has been found exceeding
20 mm.

166 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

7. Fore wing with distinct elongate blackish dash between two

discal spots. grotella Robinson (p. 192)

Fore wing without such dash betulella Busck (p. 195)

8. Second segment of labial palpus white irrorated with black and

fuscous 9

Second segment of labial palpus otherwise 11

9. Third segment of palpus white with blackish-fuscous subbasal

annulus and apex maculatella Busck (p. 194)

Third segment otherwise 10

10. Third segment of palpus fuscous irrorated with whitish and

with whitish apex yakimae, new species (p. 185)

Third segment whitish with black basal and subapical annula-

tions palousella, new species (p. 171)

11. Fore wing blackish fuscous or brownish fuscous, at least always

dark 12

Fore wing grayish or light brownish 14

12. *" The two species of this couplet cannot be separated on char-

acters of coloration; genitalia and food plant must be used.
Costa of fore wing with a distinct pinkish cast on underside;

Upper Sonoran or Arid Transition Zones.

whitmani, new species (p. 182); leptotaeniae Clarke (p. 184)
Costa without such pink color 13

13. The two species of this couplet (angustati and multifidae) cannot

be separated by the use of superficial characters. Collected
specimens may be separated as follows:

(a) Hudsonian Zone, altitude about 6,000 feet, Cascade

Range angustati, new species (p. 189)

(b) Upper Sonoran Zone to Arid Transition timbered Zone

in "intermountain" area; altitude 1,000 to 5,700 feet

multifidae Clarke (p. 187)

14. Alar expanse 19 mm. or less.. artemisiae dracvinculi Clarke (p. 169)

Alar expanse over 20 mm 15

15. Fore wing with conspicuous blackish-fuscous streak in cell.

atrostrigella, new species (p. 168)
Fore wing without conspicuous blackish-fuscous streak in cell.

artemisiella McDunnough (p. 181)

KEY TO THE SPECIES OF DEPRESSARIA BASED ON MALE
GENITALIA

1. Harpe with process from base of sacculus (figs. 198-205) 8

Harpe without such process (figs. 191-197) 2

2. Costa with prominent process before cucullus (figs. 194, 197) 3

Costa without such process (figs. 191-193; 195, 196) 4

3. Vesica armed with two or three straight terminal cornuti and a

cluster of stout curved ones near middle (fig. 194a).

atrostrigella, new species (p. 168)
Vesica armed with a cluster of straight, strong cornuti about
middle (fig. 197a) artemisiae dracxinculi Clarke (p. 169)

•• The species of this group are very difflcult to separate and must be carefully examined for characters
under the microscope. A careful examination of characters coupled with data on habitat will serve to
distinguish them.

REVISION OF THE OECOPHORIDAE — CLARKE 167

4. Clasper slender, armed with small spines and never projecting

beyond ventral edge of harpe (figs. 195, 196) 6

Clasper broad, without spines and at least part of it extending

beyond ventral edge of harpe (figs. 191-193) 6

5. Lateral edge of anellus strongly produced (fig. 196)

artemisiella McDunnough (p. 181)
Lateral edge of anellus not strongly produced (fig. 195)

alienella Busck (p. 180)

6. Transtilla roughly diamond-shaped (figs. 192b, 193b) 7

Transtilla roughly rectangular (fig. 191b) maculatella Busck (p. 194)

7. Width of clasper half, or less than half width of harpe (fig. 193)

betulella Busck (p. 195)
Width of clasper more than half width of harpe (fig. 192).

grotella Robinson (p. 192)

8. Clasper present (figs. 198-202) 9

Clasper absent (figs. 203-205) 13

9. Aedeagus as long or longer than harpe (figs. 198a-200a) 10

Aedeagus much shorter than harpe (figs. 201a, 202a) 12

10. Anellus longer than wide; spines of basal process of sacculus

long, stout (fig. 199) multifidae Clarke (p. 187)

Anellus much broader than long; spines of basal process of sac-
culus fine, slender (figs. 198, 200) 11

11. Posterior edge of anellus deeply cleft; transtillar lobes large,

broad; harpe short, broad (fig. 200) whitmani, new species (p. 182)

Posterior edge of anellus not cleft; transtillar lobes large,

narrow, harpe narrow (fig. 198) angustati, new species (p. 189)

L2. Basal process of sacculus curved toward cucullus; spines few,
large; clasper sharply angulate; transtillar lobes weak (fig. 201)

yakimae, new species (p. 185)
Basal process of sacculus transverse; spines numerous, rather

small; transtillar lobes strong (fig. 202) leptotaeniae Clarke (p. 184)

L3. Basal process of sacculus pointed (figs. 68, 205) 14

Basal process of sacculus not pointed (figs. 203, 204) 15

14. Costa of harpe deeply excavated before cucullus (fig. 68)

heracliana (Linnaeus) (p. 173)
Costa of harpe not excavated before cucullus (fig. 205).

cinereocostella Clemens (p. 171)

15. Basal process of sacculus armed with spines for most of its length

(fig. 203) juliella Busck (p. 176)

Basal process of sacculus armed with spines only at distal end

(fig. 204) eleanorae, new species (p. 178)

KEY TO THE SPECIES OF DEPRESSARIA BASED ON FEMALE
GENITALIA

1. Ductus bursae membranous (figs. 102, 275-278, 282) 2

Ductus bursae at least partly sclerotized (figs. 279-281; 283-289) 7

2. Ductus bursae spiraled (fig. 275) 3

Ductus bursae not spiraled (figs. 102, 278, 282) 5

3. Ostium a longitudinal slit between raised edges (figs. 275, 277) 4

Ostium nearly round, within triangular raised portion of genital

plate (fig. 276) betulella Busck (p. 195)

168 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

4. Ostium considerably shorter than width of genital plate; genital

plate with a long row of hairs posteriorly (fig. 277) .grotella Robinson (p. 192)
Ostium as long as width of genital plate; posterior row of hairs

reduced to about half a dozen (fig. 275) maculatella Busck (p. 194)

5. Ductus bursae bulbous before ostium (fig. 102) .heracliana (Linnaeus) (p. 173)
Ductus bursae not bulbous at any point (figs. 278, 282) 6

6. Anterior edge of ostium convex (fig. 278)-artemisiGlla McDunnough (p. 181)
Anterior edge of ostium cleft (fig. 282) alienella Busck (p. 180)

7. Ductus bursae sclerotized for at least two-thirds of its length

(fig. 279) eleanorae, new species (p. 178)

Ductus bursae otherwise (figs. 280, 281; 283-289) 8

8. Sclerotized portion of ductus bursae limited to a narrow band

(figs. 283, 284) 9

Sclerotized portion of ductus bursae otherwise (figs. 196, 197, 285, 289). 10

9. Sclerotized portion of ductus bursae armed with small teeth

(fig. 284) palousella, new species (p. 171)

Sclerotized portion of ductus bursae unarmed (fig. 283)

artemisiae dracvmculi Clarke (p. 169)

10. Sclerotized portion of ductus bursae strongly curved (figs. 286-288) 11

Sclerotized portion of ductus bursae straight or only slightly

curved (figs. 280, 281, 285, 289) 13

11. Genital plate with a pair of shallow, cupped sclerites on anterior

margin (fig. 288) multifidae Clarke (p. 187)

Genital plate without such sclerites (figs. 286, 287) 12

12. Genital plate abruptly narrowed laterally; ostium a narrow slit

(fig. 287) angustati, new species (p. 189)

Genital plate of nearly equal width throughout, ostium trian-
gular (fig. 286) whitmani, new species (p. 182)

13. Signum in posterior part of bursa copulatrix (figs. 285, 289) 14

Signum in anterior part of bursa copulatrix (figs. 280, 281) 15

14. Sclerotized ventral sclerites of genital plate equal to length of

sclerotized portion of ductus bursae (fig. 285)

yakimae, new species (p. 185)
Sclerotized ventral sclerites of genital plate shorter than length
of sclerotized portion of ductus bursae (fig. 289)

leptotaeniae Clarke (p. 184)

15. Ductus bursae sclerotized for less than half its length; a small

protuberance from the side of the sclerotized part (fig. 281)

cinereocostella Clemens (p. 171)
Ductus bursae sclerotized for about half its length and without
small protuberance from the sclerotized part (fig. 280)

juliella Busck (p. 176)

DEPRESSARIA ATROSTRIGELLA, new species
PlxVte 35, FiGUKES 194, 194a

Labial palpus light oclireous- white ; second segment suffused with
fuscous in the brush; third segment immaculate. Antenna light fus-
cous somewhat mottled basally with gray; basal segment gray be-
neath. Head, thorax, and fore wing ochreous-white. The whole
so strongly suffused with deep gray that the ground color is nearly
obliterated. Vertex and the head behind the antennae infuscated.

REVISION OF THE OECOPHORIDAE — CLARKE 169

Thorax liglitly irrorated with fuscous. Inner angle of fore wing,
veins, a strong longitudinal dash in cell and a series of poorly defined
spots around termen, blackish fuscous. The veins are, for the most
part, weakly marked, but vein Ic is marked with blackish fuscous for
its entire length ; cilia deep gray, tipped with ochreous-white. Hind
wing pale smoky gray; cilia lighter. Legs ochreous-white, the fore
and mid legs suffused with grayish fuscous exteriorly. Abdomen
pale grayish fuscous, faintly annulated with dull ochreous-white,

Male genitalia. — Harpe moderately broad, clothed apically with
numerous stout, straight hairs; cucullus rounded; on costa, before
cucullus, a large flat process that terminates in a small hook ; sacculus
moderately broad and sclerotized, without basal process; clasper a
minute papillate process about middle of harpe, with three or four
strong hairs from surface. Anellus a large subrectangular plate con-
stricted posteriorly; posterior edge truncate. Aedeagus rather long,
stout, tapering gradually to a slender, curved point ; vesica armed with
two or three long straight cornuti and a cluster of short, stout, curved
ones about middle. Vinculum rounded with well developed dorsoan-
terior process. Transtilla a broad, sclerotized, curved band; lateral
lobes indicated chiefly by hairs and constituting the lateral extremities
of the transtilla proper. Socii long digitate hairy lobes. Tegumen
reduced, truncate.

Alar expanse, 21-24 mm.

Type. — In the Canadian National Collection.

Type locality. — Aweme, Manitoba.

Paratype.—U. S. N. M. No. 53299.

Food plant. — Unknown.

Remarks. — Described from the $ type and 1 S paratype, both
from the type locality (6-IX-1928, N. briddle).

The genitalia of this species are nearest to dracunculi but differ
from it chiefly by the cornuti and the transtilla. The figures of
the two will show these differences adequately. I have seen no
females of atro^trigella.

The paratype is not marked so contrastingly as the type male and
is the larger of the two specimens.

DEPRESSARIA ARTEMISIAE DRACUNCULI Clarke

Plate 35, FiGxniES 197, 197a ; Plate 48, Figure 2S3

Depressai-ia dracunculi Clarke, Can. Ent., vol. 65, p. 90, pi. 5, 1933. —
McDuNNOUGH, Check list of the Lepidoptera of Canada and the United States
of America (Part 2, Microlepidoptera), No. 8395, 1939. — Gaede, m Bryk,
Lepidopterorum catalogus, pt. 92, p. 282, 1939.

Labial palpus creamy white ; second segment suffused and irrorated
in the brush with fuscous; third segment with subbasal and supra-

170 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

medial annuli black, these sometimes poorly defined and almost
wholly replaced by the lighter ground color. Antenna fuscous with
grayish or white annulations. Head light fawn, the scales tipped
with sordid white. Thorax light fuscous, the scales tipped with
gray or white; anteriorly the fuscous is more pronounced. Fore
wing with costal third fuscous mixed with gray, remainder light to
dark fawn irrorated with fuscous and gray; toward apex the colors
merge and are indistinguishable; at base, in anal angle, a small
blackish-fuscous patch; in cell a series of three blackish-fuscous
spots, sometimes confluent, forming a single longitudinal dash; cilia
grayish fuscous with a broad, dark, subbasal band. Legs creamy
white, strongly overlaid with fuscous except at joints. Abdomen
silvery grayish above; creamy white beneath suffused with fuscous
laterally.

Male genitalia. — Harpe broad, with very few coarse hairs ; cucullus
rounded, densely clothed with coarse, stiff hairs; before cucullus,
from costa, a prominent dull-pointed projection; clasper minute;
emitting a few long hairs ; sacculus broadly folded and without basal
process. Anellus a moderately small sclerotized plate, constricted
at middle ; anterior and posterior edges strongly convex. Vinculum
broad, rounded. Aedeagus long, slightly curved, terminating in a
long, attenuated point ; vesica armed with a cluster of straight, strong
cornuti. Transtilla a very broad, sclerotized band with well-devel-
oped hairy lobes, the whole fused with the posterior portion of the
anellus. Socii small, narrow, hairy lobes.

Female genitalia. — Genital plate moderately broad. Ostium large,
round, situated at the extreme anterior edge of the genital plate. A
narrow area of the ductus bursae, just posterior to the bursa copulatrix,
moderately sclerotized, unarmed; the remainder of the ductus mem-
branous; inception of the ductus seminalis about midway between
the sclerotized part of the ductus bursae and the ostium. Bursa copu-
latrix large with a very small signum in the anterior half.

Alar expanse, 12.5-19 mm.

Tyfe.—\n the United States National Museum.

Type locality. — Snake River, Wliitman County, opposite Clarkston,
Wash.

Food plant. — Artemisia dracunculoides Pursh.

DistHl)Ution.—^OYi\mQsiQvr]. United States and southwestern
Canada.

United States records
Washington: Almota, Whitman County, 12 $ $ , 12 9 ? (5-23-IV-34) ; Snake
River, Whitman County, opposite Clarkston, 5 $ $, 1 9 9 (February to
May dates, 1931-32 ; the February and March dates are for specimens reared
in a greenhouse) ; Truax, Whitman County, 7 5 5 , 10 9 9 (4-12-V-35) . (All
these specimens were reared from larvae collected by the writer.)

REVISION OF THE OECOPHORIDAE — CLARKE 171

Canadian records

British Columbia: Vernon (2 $ 5, 6-VI-1926; 2 , 12-VII-1927, E. P. Venables).

DEPRESSARIA PALOUSELLA, new species

Plate 48, Figure 284

A distinct, medium-sized moth resembling yakimae but smoother in
appearance.

Labial palpus with second segment shining white with much black
scaling exteriorly ; the white scales show a faint pinkish iridescence ;
third segment white with black basal and subapical annulations. An-
tenna with basal segment blackish fuscous with a few whitish scales
mixed; remainder grayish with blackish-fuscous annulations. Head
light fuscous, the scales white tipped. Thorax, base of tegula, and
base of fore wing black. The posterior two-thirds of tegula, the costa
to middle of fore wing and the major portion of the thorax strongly
marked with cinereous. Ground color of fore wing a light brownish
fuscous, darker basally and faintly irrorated with cinereous; in the
cell a strongly marked, longitudinal black dash edged anteriorly with
cinereous; veins, especially 9, 10, and 11 strongly marked with black
and around termen a series of indistinct, blackish spots at ends of veins ;
cilia light brownish fuscous. Hind wing light grayish-fuscous but
darker at extreme apex and around termen ; cilia light brownish fus-
cous, darker basally. Legs blackish fuscous irrorated with white.
Abdomen grayish fuscous above, blackish fuscous with much white
scaling below.

Female genitalia. — Genital plate broad, moderately sclerotized, not
appreciably produced ventrally around ostium. Ostium oval, trans-
verse. Posterior fourth of ductus bursae broadly expanded with the
major portion of this section strongly sclerotized and armed with
small teeth ; the remainder of the ductus is membranous. Inception
of ductus seminalis on the posterior margin of the sclerotized part
of the ductus bursae. Bursa copulatrix moderately large with a small
but well-defined signum.

Alar expanse, 19-20 mm.

Type and paratypes. — U.S.N.M. No. 52074.

Type locality. — Pullman, Wash.

Remark.^. — Described from the $ type (26-VI-30) and 1 9 para-
type (20-IX-30), both collected by the author.

DEPRESSARIA CINEREOCOSTELLA Clemens

Plate 38, Figures 205, 205a ; Plate 48, Figure 281

Depressaria cinereocostella Cij:mens, Proc. Ent. Soc. Philadelphia, vol. 2, p.
422, 1864.— Robinson, Ann. Lye. Nat. Hi.st. New York, vol. 9, p. 155, pi. 1,
fig. 6, 1869. — Clemens, in Stainton, Tineina of North America, p. 245, 1872. —

172 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Chambers, Can. Eut., vol. 4, p. 91, 1872; U. S. Geol. Geogr. Surv. Terr.
Bull. 4, p. 138, 1878.— Walsingham, Ins. Life, vol. 1, p. 255, 1889.— Riley, in
Smith, List of the Lepidoptera of Boreal America, No. 5258, 1891.— Busck,
Proc. U. S. Nat. Mus., vol. 24, p. 749, 1902; in Dyar, U. S. Nat. Mus.
Bull. 52, No. 5891, 1903 ; Proc. Ent. Soc. Washington, vol. 5, p. 217, 1903.—
Kh:abfott, in Smith, List of the Lepidoptera of Boreal America, No. 6434,
1903.— Busck, Proc. U. S. Nat. Mus., vol. 35, p. 200, 1908.— Babnes and Mc-
DuNNouGH, Check list of the Lepidoptera of Boreal America, No. 6482,
1917. — Meyrick, in Wytsman, Genera insectorum, fasc. 180, p. 172, 1922. —
Forbes, Cornell Univ. Agr. Exp. Stat., Memoir 68, p. 242, 1923.— McDun-
NOUGH, Check list of the Lepidoptera of Canada and the United States of
America (Part 2, Microlepidoptera), No. 8408, 1939. — Gaede, in Bryk,
Lepidopterorum catalogus, pt. 92, p. 278, 1939.
Depressaria clatisella Waxker, List of the lepidopterous insects in the collec-
tions of the British Museum, vol. 29, p. 564, 1864. — Walsingham, Proc. Zool.
Soc. London, 1881, p. 312. — McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera), No.
8408, 1939 (cited as synonym of cinereocostella Clemens).

Labial palpus pale cinereous; second segment with submedial and
subterminal spots exteriorly blackish fuscous and with the brush
suffused with reddish; third segment with a dull blackish-fuscous
supramedial annulus. Antenna cinereous, narrowly annulated with
fuscous basally and suffused with reddish; beyond middle strongly
suffused with fuscous ; extreme tip cinereous. Face, head, thorax, and
ground color of fore wing cinereous ; collar fuscous ; head and thorax
irrorated with fuscous and suffused with red, the thorax and tegula
especiall}'- so anteriorly. Fore wing, except costa, heavily overlaid
with dull brownish red, so much so that the wing appears red with a
contrasting light costa, suffused with blackish fuscous in basal half
and irrorated with black and cinereous scales ; in fold and along veins
a series of longitudinal black dashes; extreme base of costa and anal
angle blackish fuscous; from middle of costa, around termen to inner
margin, a series of poorly defined blackish-fuscous spots; cilia fus-
cous suffused with red. Hind wing grayish fuscous, lighter basally ;
cilia pale fuscous, white tipped, with rosy tinge and with dark fuscous
subbasal and subterminal bands. Legs cinereous with dull blackish
irrorations and suffusion except at joints. Hind tibia with a dull
smoky streak exteriorly; outer pair of spurs dull black. Abdomen
cinereous with fuscous suffusion dorsally and a blackish-fuscous longi-
tudinal line on each side beneath.

Male genitalia. — Harpe broadest at middle, sparsely clothed with
fine hairs ; cucullus rounded ; clasper absent ; sacculus broadly folded,
with long, pointed basal process; the basal process of the sacculus is
clothed with short, stout spines over its entire length. Anellus rec-
tangular, narrowed posteriorly; lateral edges concave in anterior
part. Vinculum bluntly pointed. Aedeagus long, slender, and
slightly curved ; bluntly pointed ; vesica with three to seven stout

REVISION OF THE OECOPHORIDAE — CLARKE 173

cornuti. Transtilla a large rectangular plate, produced anteriorly
at middle, with well-developed hairy lobes, the latter fused. Socii
large fleshy, hairy flaps.

Female genitaUa. — Genital plate broad. Ostium a long transverse
slit. Posterior two-fifths of ductus bursae sclerotized, straight; a
small protuberance from the side of the sclerotized portion of the
ductus bursae; inception of ductus seminalis just before ostium.
Bursa copulatrix small with well developed signum in the anterior
end.

Alar expanse, 15-22 mm.

Type. — In the Academy of Natural Sciences of Philadelphia.

Type locality. — "Virginia."

Food plants. — Carum carvi L., Sium lineare Michx., and
Ligusticum scoticum L.

Distribution. — Northeastern United States and eastern Canada.

United States records

District of Columbia: 2 S S (19-VII-99, A. Busck).

Iowa: Ames $ (22-VIII-1918; A. W. Lindsey) ; Iowa City, $ (15-IV-1917).

Missouri: St. Louis, S (2(>-VIII-05, H. McElhose).

New Hampshire: Hampton, 9 5 5 , 10 $ 9 (February to August dates, 1906-

1908, S. A. Shaw [reared]).
Pennsylvania: Philadelphia (no date or collector).

Canadian records

Manitoba: Cartwright, S (no date; E. F. Heath).

Nova Scotia: White Point Beach, Queens County, 13 cJ 5 , H 5 ? (2-ia-VIII-

1934, J. McDunnough [reared]).
Ontario: Toronto, ? (2-III-95).

Remarks. — The number of cornuti on the vesica seems to be of no
significance in this species. Such differences in the number of cornuti
are usually specific but in the present case all intergrades may be
found in any one group of specimens. It is also rather strange that
cinereocostella should feed on so many species of plants, a habit which
is unusual for species of this genus. The specimens reared from the
different food plants exhibit no characters by which any one group
can be separated from another.

This species is of some economic importance. Although it has not
done great damage to crops of caraway, it is potentially dangerous
because of its unusual habit of feeding on a number of food plants.

DEPRESSARIA HERACLIANA (Linnaeus)

Plate 2, Fi;;xjee 11 ; Plate 6, Figuee 48 ; Plate 10, Figures 68, 68a ;
Plate 17, FiotJKB 102

Phalaena Tortrix heracUana Linnaeus, Systema naturae, ed. 10, p. 532, No.
225, 1758; ibid., ed. 12, p. 380, No. 326, 1767.— De Geee, Memoires pour
servir a I'histoire des insectes, vol. 2, p. 407, No. 5, 1771.

174 PROCEEDINGS OF THE NATIONAL MUSEUM vol. eo

Phalaena Tortrix heracleana Linnaeus, Fauna Suecia, p. 347, No. 1334, 1761.

Phalaena heraclia7ia (Linnaeus) Mulleb, Natursystem, p. 731, No. 326, 1774.

Phalaena Jwracleana Retzius, Genera et species insectorum, p. 45, 1783.

Pyralis heracleana Fabeicius, Systema entomologiae, p. 655, No. 56, 1775;
Entomologia systematica, vol. 3, pt. 2, p. 286, No. 178, 1794.

Depressaria heracliana (Linnaeus) Zellek, Isis von Oken, vol. 4, p. 305, 1839;
Linnaea Ent., vol. 9, p. 312, 1854.— Bethune, Can. Ent., vol. 2, p. 19, 1870.—
LiNTNER, Can. Ent., vol. 5, p. 82, 1873.— Zeixeb, Verb, zool.-bot. Ges. Wien,
vol. 23, p. 235, 1873.— Chambers, U. S. Geol. Geogr. Surv. Terr. Bull. 4, p.
138, 1878.— Walsingham, Proc. Zool. Soc. London, 1881, p. 312.— Riley, in
Smith, List of the Lepidoptera of Boreal America, No. 5265, 1891. — Dietz,
in Smith, Catalogue of the insects of New Jersey, p. 474, 1900. — Rebel, in
Staudinger and Rebel, Catalog der palaearctischen Lepidopteren, vol. 2, No.
3280, 1901.— BuscK, Proc. U. S. Nat. Mus., vol. 24, p. 748, 1902; in Dyar,
U. S. Nat. Mus. Bull. 52, No. 5889, 1903.— Kearfott, in Smith, List of the
Lepidoptera of Boreal America, No. 6432, 1903.— Pettit, Michigan Agr. Exp.
Stat. Bull. 233, p. 189, 1906. — Chittenden, Insects injurious to vegetation,
p. 187, 1907.— Jarvis, Ontario Ent. Soc. Rept., vol. 37, p. 48, 1907.— Busck,
Proc. U. S. Nat. Mus., vol. 35, p. 200, 1908.— Bethune, Ontario Agr. Coll.
Bull., vol. 171, p. 28, 1909. — Fulton, Wright, and Gregg, Pennsylvania Agr.
Exp. Stat. Bull. 110, p. 29, 1911. — Sanderson, Insect pests of farm, garden,
and orchard, p. 417, 1912. — Gooderham, Proc. Ent. Soc. Nova Scotia, vol. 1,
p. 94, 1915.— Brettain and Gooderham, Can. Ent., vol. 48, p. 37, 1916.—
Du Porte, Ontario Ent. Soc. Rept., vol. 46, p. 50, 1916.— Barnes and Mc-
DuNNOUGH, Check list of the Lepidoptera of Boreal America, No. 6478,
1917.— Bethune, Ontario Dept. Agr. Bull. 251, p. 24, 1917.— Gibson, Ontario
Ent. Soc. Rept., vol. 47, p. 16, 1917.— Gossard, Ohio Monthly Bull., vol. 4,
p. 379, 1919.— Mosher, Journ. Econ. Ent., vol. 12, p. 261, 1919.— Traver,
Psyche, vol. 26 p. 77, 1919.— Gibson, Can. Dept. Agr. Ent. Circ, vol. 14, p. 13,
1920. — Leonard, Journ. Econ. Ent., vol. 13, p. 491, 1920.— Hudson, Ontario
Ent. Soc. Rept., vol. 21, p. 35, 1921. — Meybick, in Wytsman, Genera in-
sectorum, fasc. 180, p. 171, 1922.— Forbes, Cornell Univ. Agr. Exp. Stat.
Memoir 68, p. 243, 1923.— Ellis, Journ. Agr. Res., vol. 30, p. 789-790, 1925.—
Caesar, Ontario Dept. Agr. Bull. 325, p. 27, 1927.— Brettain, Nova Scotia
Dept. Nat. Res. Bull. 12, p. 86-87, 1927.— Drake and Decker, Iowa Agr. Exp.
Stat. Circ. 103, p. 16, 1927.— Pettit, Michigan State Board Agr. Rept., vol. 66,
p. 343-344, 1927.— Caesar, Ontario Dept. Agr. Bull. 359, p. 29-30, 1931.—
DusTAN, Can. Dept. Agr. Ent. Bull., vol. 32, p. 51, 1932.— McDunnough,
Check list of the Lepidoptera of Canada and the United States of America
(Part 2, Microlepidoptera), No. 8404, 1939. — Gaede, in Bryk, Lepidopterorum
catalogus, pt. 92, p. 283, 1939.

Phalaena heraelei Retzius, Genera et species insectorum, p. 45, 1783.

Pyralis umhellana Fabricius, Entomologica systematica, vol. 3, p. 286, No.
177, 1794.

Haemilis pastinacella Duponchel, Histoire naturelle des Lepidopteres de la
France, vol. 11, p. 153, pi. 291, figs. 4 and 5, 1S38.— Beuand, Ann. Ent. Soc.
France (ser. 2), 1844, p. 189, pi. 6, fig. A, B, a, b.

Haemylis heraeleella Zetterstedt, lusecta Lapponica, p. 999, 1840.

Depressaria ontariella Bethune, Can. Ent., vol. 2, p. 3, 19, 1870.

Labial palpus wliitish oclireoiis; second segment suffused in the
brush and irrorated exteriorly with fuscous and brown ; third segment
with subbasal and supramedial annuli blackish fuscous, the former

REVISION OF THE OECOPHORIDAE — CLARKE 175

narrow and poorly defined. Antenna luteous annulated with fuscous ;
basal segment blackish fuscous above. Face pale, shining grayish
fuscous. Head, thorax, and fore wing luteous shaded and streaked
with brown and blackish fuscous; extreme base of costa, anal angle,
discal spot at the end of the cell, and a series of spots from apex,
around termen to imier margin, blackish fuscous ; veins rather strongly
indicated by blackish-fuscous scaling; apical area with whitish irrora-
tions and an indistinct, narrow, outwardly curved fascia of the same
color from apical third of costa; cilia grayish-fuscous. Hind wing
pale yellowish- fuscous, darker apically and with a narrow, fuscous
terminal line ; cilia paler with a fuscous subbasal band. Legs luteous
shaded and irrorated with fuscous except at joints. Abdomen luteous
above, lightly infuscated; beneath, whitish-ochreous with a row of
blackish-fuscous spots on each side.

Male genitalia: — Harpe with cucullus and distal half of ventral
margin clothed with stout hairs; costa excavated before cucullus;
clasper absent, sacculus moderately broad, strongly sclerotized, and
with a long slender process from base. Anellus an elongate slightly
sclerotized plate narrower at distal than at proximal end. Transtilla
a broad band with well-developed hairy lobes. Aedeagus stout, elon-
gate, pointed; vesica armed with eight or ten long slender cornuti.
Vinculum rounded. Gnathos a spined knob. Socii elongate hairy
lobes, widely separated. Tegumen truncated ; uncus absent.

Female genitalia. — Lobe of ovipositor somewhat sclerotized basally.
Genital plate broad, strongly sclerotized. Ostium round; anterior
edge strongly sclerotized. Ductus bursae membranous, dilated at pos-
terior three-fourths; inception of ductus seminalis just before ostium.
Bursa copulatrix moderately large with small diamond-shaped signum.

Alar expanse, 21-29 mm.

Types.— Lo^fi ''^

Type localities. — Europe {heracliana, heracUi, umhellana.^ pastina-
cella) ', Ontario, Canada (ontariella) .

Food plants. — Parsnip {Pastinacea sativa Ij.) ; Angelica sp. (boring
in stem).

Distribution. — Throughout United States and southern Canada.

United States records

Arizona: Walton, Yuma County, 9 (1-6-III-1925, O. C. Poling).

Illinois: Chicago, S S $ (June, 1900; 12-VII-'02, W. D. Kearfott ; one without

date, K. Wyatt collector) ; Decatur, 2 $ $ (July 1-7).
Indiana: Morgan County, $ (lO-VII-1931, B. E. Montgomery; "reared from

wild par.snip").

SI Dr. McDunnough states (in lift.) : "Regarding the type of ontariella Bethune, I have no
information. It is certainly not in Ottawa and I imagine that most of Bethune's types
have been destroyed by Dermestes. . . . What remains of his collection has been scattered
about, but Dr. Ide of the Royal Ontario Museum who went over the specimens stated that
there were no types amongst them."

176 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Massachusetts: Cliftondale, 2 $ $ (August 1920, J. D. Caffrey).
New Jersey: Caldwell, S $ $,5 9 9 (August, W. D. Kearfott).
New York: Rye, Q $ S , 10 9 9 (2-20-VIII-1939, J. F. G. Clarke) ; $ (no date

or locality: Wm. Beutenmiiller).
Oregon: Portland, $ (l-VIII-1914, L. Leland).
Pennsylvania: Bristol, 8 $ S , 8 9 ? (June to August dates, 1S88, T. Pergande;

reared from parsnip) ; New Brighton, 3 S $,2 9 9 (5-13-VIII-1907, H. D.

Merrick) ; Oak Station, Allegheny County, $ (30-VII-lO, Fred MarlofE) ;

Pittsburgh, $, 9 (10-IV-'06, 13-IV-'06, Henry Engel) ; West Chester,

2 5 (5, ? (July, 1919, F. M. Trimble; reared from parsnip).
Rhode Island: Bristol, $ (15-VII-1920, no collector).
Utah: Logan, 9 ( lO-VIII-1907, E. S. G. Titus; reared from wild parsnip);

Murray, $ (12-VII-1913, Timberlake) ; Vineyard, 9 (l-IX-1912, Tom

Spalding).
Washington: Bellingham, 12 S $, 9 (4-VIII-1930, W. W. Baker; reared from

parsnip) ; Oroville, $ (18-VII-1933, Judson Murray; reared from parsnip) ;

Pullman, $ (29-IV-24, J. F. G. Clarke.)

Canadian records

British Columbia: Fraser Mills, 9 (18-IX-1925, L. E. Marmont) ; Lillooet
(2-VIII-1931, A. W. A. Phair) ; Summerland (12-IV-1934; 22-24-VII-1935,
A. N. Gartrell).
Nova Scotia: Truro (17-VIII-1915, no collector).
Ontario: London (2S-29-VII-1931, G. S. Walley) ; Ottawa (19-VIII-1903;

ll-VIII-1908, J. Fletcher); Port Hope (S-IV-1895, no collector).
Quebec: Meach Luke (29-VII to 6-VIII-1903, C. H. Young).

Re?narks. — This is the notorious "parsnip webworm," which has
appeared so frequently in the literature in America and abroad.

The name has repeatedly been credited to De Geer (1T71) but
should be attributed to Linnaeus (1758). Sherborn (Index Ani-
malium, 1902) credits the name to Linnaeus, but others have failed to
acknowledge this authorship.

The species is undoubtedly more widespread in America than is
indicated by the distribution given; but I have included in the dis-
tribution only the localities from which I have seen specimens. At
New Castle, Del., however, I have seen abundant evidence of the work
of this species.

I have been unable to locate any of the types, although some may
be in existence.

DEPRESSARIA JULIELLA Busck

Plate 38, Figures 203, 203a; Plate 47, Figxjee 280

Depressaria juUclla BuscK, Proc. Ent. Soc. Washington, vol. 9, pi. 91, 1908;
Proc. U. S. Nat. Mus., vol. 35, p. 200, 1908.— Baenes and McDunnough,
Check list of the Lepidoptera of Boreal America, No. 6475, 1917. — Metrick,
in Wytsman, Genera insectorum, fasc. 180, p. 171, 1922.— Clarke, Can.
Ent., vol. 66, p. 178, 1934.— McDunnough, Check list of the Lepidoptera of
Canada and the United States of America (Part 2, Microlepidoptera),
No. 8399, 1939. — Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 286,
1939.

REVISION OF THE OECOPHORIDAE — CLARKE 177

Labial palpus ochreous; second segment with a dark reddish-fus-
cous spot near base externally and with the brush suffused with red
and irrorated with reddish fuscous; third segment suffused with red
anteriorly and with poorly defined reddish-fuscous subbasal and
supramedial annuli. Antenna light reddish fuscous, narrowly and
faintly annulated with fuscous; basal segment ochreous beneath.
Face shining whitish ochreous. Head, thorax, and fore wing light
ochreous suffused and mottled with red. Below antenna, in front,
a vertical reddish-fuscous patch ; from base of costa, following fold,
a reddish-fuscous streak to about basal fourth ; in anal angle a small
reddish-fuscous patch; both of these dark markings mixed with black
scales. Along costa and inner margin and in apical third numerous
white scales; all the veins are more or less indicated by reddish fus-
cous and black mixed ; discal spot at end of cell faintly indicated by
a few red scales; cilia fuscous, strongly suffused with red and with
a blackish-fuscous subbasal band. Hind wing fuscous, lighter
basally; cilia red with a fuscous subbasal band. Legs ochreous suf-
fused with red and mottled with reddish fuscous. Abdomen ochre-
ous suffused with fuscous dorsally and with a blackish-fuscous line
on each side ventrally.

Male genitalia. — Harpe gradually tapering to the rounded cucullus ;
clasper absent, sacculus broadly folded, with a large, elongate, trun-
cated process from its base ; this elongate process is more or less armed
with spines over most of its length; at the distal end the spines are
numerous. Anellus a sclerotized plate ; broad anteriorly, narrow pos-
teriorly; anterior edge with a shallow median cleft; posterior edge
convex. Vinculum produced anteriorly. Aedeagus long, nearly
straight, dilated at proximal end ; vesica armed with 7-10 stout cornuti.
Transtilla membranous, with small hairy lobes. Socii long, narrow,
hairy flaps.

Female genitalia. — Genital plate broad, moderately sclerotized.
Ostium transverse, elliptical, situated at anterior margin of the genital
plate. Ductus bursae sclerotized, slightly curved and dilated in pos-
terior half, without protuberance; membranous in anterior half; in-
ception of ductus seminalis just anterior to ostium. Bursa copulatrix
large with a small, but well developed signum near anterior end.

Alar expanse, 21-25 mm.

Type. — In United States National Museum.

Type locality. — Pecos, N. Mex.

Food plant. — Cicuta occidentalis Greene.

Distribution. — ^^Vestern United States.

178 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

United States records

Colorado: Alamosa, $ (Oslar).

New Mexico: Pecos, $ (September, T. D. A. Cockerell).

Utah: Prove, 5 S $, S $ 9 (August and September dates) ; Vineyard, IQ $ S ,

17 9 9 (September and October dates, Tom Spalding).
Washington: Pullman, 56 $ $ , 52 9 9 (reared, July, August dates, 1933-1935,

J. F. G. Clarke).

Remxirhs. — I have already given ^^ a summary of the habits and have
briefly described the larva of this species but it seems appropriate to
give a brief description of the pupa at this time.

Pupa. — First day : Wing, antennal and leg sheaths, also dorsal part
of head and thorax light lemon yellow. Abdominal segments light
whitish ochreous except the last caudal segment which is strongly
tinged with reddish brown. The last five segments are free and
movable.

Second day: Ptipa light reddish brown; last segment somewhat
darker. From the second day on the color becomes progressively
darker. The wing sheaths become very dark brown first. The ab-
dominal segments gradually become darker until at the time of emer-
gence the entire pupa is nearly black.

Busck's remark ^^ concerning the close relationship of julAella to the
European nervosa is borne out by a study of both the male and female
genitalia. The differences in genitalia (although admittedly slight)
and the difference in food plant will serve to distinguish the two. The
coloration of juliella^ as indicated by a reared and a collected series of
over 150 specimens, is usually brighter and lighter than that of nervosa.

DEPRESSARIA ELEANORAE, new species

Plate 38, Figukes 204, 204a ; Plate 47, Figure 279

Similar to D. juUella Busck but smaller.

Antenna reddish ochreous above, lighter beneath and toward tip;
narrowly annulated with reddish fuscous. Second segment of labial
palpus whitish ochreous strongly suffused with red and reddish fus-
cous exteriorly; third segment whitish ochreous with broad reddish-
fuscous basal and subapical fasciae. Head light reddish ochreous;
face light yellowish ochreous. Thorax, tegula, and fore wing ochreous
strongly overlaid with red, reddish fuscous, and white scales; at base
of wing a black spot on costa and a similar one on dorsum ; narrowly
but strongly tinged with bright red; in middle of cell an indistinct
longitudinal streak of reddish fuscous and white scales mixed; cilia
light reddish fuscous. Hind wing light fuscous, lighter basally with
a distinct reddish hue ; cilia light reddish fuscous with a fuscous sub-
s'" Clarke, J. F. G., Can. Ent. vol. 66, p. 178, 1934,
«3 Proc. Ent. Soc. Washington, vol. 9, p. 91, 1908.

REVISION OF THE OECOPHORIDAE — CLARKE 179

basal band. Legs: Femora whitish ochreous; fore and mid femora
strongly marked with reddish fuscous exteriorly; tibiae and tarsi of
fore and mid legs reddish fuscous with whitish ochreous inwardly.
Hind tibia whitish ochreous strongly tinged with pink ; tarsi strongly
marked with fuscous. Abdomen whitish ochreous; basal segments
suffused with light fuscous; underside with well-defined lateral row
of fuscous spots on each side.

Male genitalia. — Harpe sparsely clothed with fine hairs; clasper
absent; cucullus rounded; sacculus broadly folded and with a long,
distally dilated, curved basal process; the latter armed with spines
only at distal end. Anellus a broad, sclerotized plate abruptly nar-
rowed in posterior half ; posterior edge strongly convex ; anterior mar-
gin indented. Vinculum with a strongly produced, acutely pointed
anterior process. Aedeagus long, slender, nearly straight ; vesica with
12 or more stout cornuti. Transtilla membranous with weakly de-
veloped, haiiy lobes. Socii long, slender, hairy flaps.

Female genitalia. — Genital plate broad, moderately sclerotized;
ostium transverse, slitlike; anterior edge of ostimn slightly convex.
Posterior two-thirds of ductus bursae strongly sclerotized. Bursa
copulatrix large ; signum small and situated at the anterior end.

Alar expanse, 18-20 mm.

ry/?6'.— U.S.N.M. No. 52082.

Type locality. — Hymers, Ontario (August 16-23).

Remarks. — Described from the $ type, 1 $ and 6 $ paratypes as
follows: Ottawa, Ontario ?> $ $ (28-VIII-1906 ; 5-IX-1908, James
Fletcher; 19-III-1933, C. H. Young) ; 5 (20-V-1925, C. H. Curran) ;
Trenton, Ontario, $ (9-IX-lO, Evans). The other two paratype
$ $ are without locality labels ; one with "In office, 21-X-1903" and the
other "23-X-04 J. F." (James Fletcher?). Paratypes in U. S. Na-
tional and Canadian National collections. I am indebted to Dr. J.
McDunnough for seven of the eight specimens of the type series.

This species is the eastern analogue of the western juliella but is
immediately distinguished from juliella by its smaller size. The
genitalia of the two are similar but differ as follows : In the male of
juliella the vinculum is bluntly pointed, in eleanorae sharply pointed ;
the basal process from the sacculus of juliella is spiny over practically
its entire length while that of eleanorae has spines only on the distal
half; on the vesica of jidiella there is one comutus which is noticeably
much larger than the rest, but in eleanorae the differences in the
lengths of the cornuti are not so striking. The ductus bursae of the
female genitalia of eleanorae exhibits a sclerotized portion of much
greater length than that found in juliella.

I take pleasure in naming this species for Mrs. Eleanor A. Carlin,
staff artist for the Bureau of Entomology and Plant Quarantine.

180 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

DEPRESSARIA ALIENELLA Busck

Plate 35, Figubks 195, 195a ; Plate 48, Figuee 282

Depressaria alienella Busck, Proc. U. S. Nat. Mus., vol. 27, p. 765, 1904. — Ander-
son, Catalogue of British Columbia Lepidoptera, No. 1094, 1904. — Busck,
Proc. U. S. Nat. Mus., vol. 35, p. 200, 1908. — Barnes and McDunnough, Check
list of the Lepidoptera of Boreal America, No. 6473, 1917. — Meykick, in
Wytsman, Genera insectorum, fasc. 180, p. 171, 1922. — McDunnough, Check
list of the Lepidoptera of Canada and the United States of America (Part. 2,
Microlepidoptera ) , No. 8393, 1939. — Gaede, in Bryk, Lepidopterorum cata-
logus, pt. 92, p. 273, 1939.

Depressaria emcritella Walsingham (not Stainton), Proc. Zool. Soc. London,
1881, p. 381. — Riley, in Smith, List of the Lepidoptera of Boreal America, No.
5261, 1891.— Busck, Proc. U. S. Nat. Mus., vol. 24, p. 746, 1902 ; in Dyar, U. S.
Nat. Mus. Bull. 52, No. 5884, 1903.

Depressaria emeriteUa alienella (Busck) Forbes, Cornell Univ. Agr. Exp. Stat.,
Memoir 68, p. 243, 1923.

Labial palpus white ; second segment irrorated exteriorly, and suf-
fused in the brush, with reddish fuscous and with a distinct blackish-
fuscous spot near base exteriorly ; third segment with a broad blackish-
fuscous supramedial annulus. Head and thorax white ; thorax sparsely
irrorated with reddish fuscous anteriorly ; tegula dark reddish fuscous.
Fore wing light fuscous strongly overlaid with red or reddish fuscous,
irrorated with cinereous and fuscous and streaked with blackish
fuscous; at the end of cell a white discal spot preceded and followed
with fuscous; around termen a poorly defined row of fuscous spots.
Legs white, suffused and irrorated with blackish fuscous except at
joints. Abdomen pale fuscous above, ochreous-white beneath, irro-
rated with fuscous.

Male genitalia. — Harpe moderately clothed with long fine hairs,
clasper long, pointed, not projecting beyond ventral edge of harpe,
armed with numerous fine spines; sacculus narrowly folded, without
basal process ; cucullus bluntly pointed. Anellus broadly oval ; lateral
edge not strongly produced. Vinculum bluntly pointed. Transtilla
a broad sclerotized band with hairy lobes moderately well developed.
Aedeagus long, slender, gently curved. Tegmnen truncated; socii
small, hairy lobes.

Female genitalia. — Ostium slitlike transverse, with shallow median
cleft on anterior edge ; situated near the anterior edge of the genital
plate. Duetus bursae membranous, straight; inception of the ductus
seminalis well before ostium and anterior to the sclerotized portion
of the ductus bursae. Bursa copulatrix large with well-developed
toothed sig-num.

REVISION OF THE OECOPHORIDAE — CLARKE 181

Alar expanse, 18-21 mm.

Type. — In the United States National Museum.
Type locality — Kaslo, British Columbia, Canada.
Food plant. — Artemisia sp., yarrow.

Distribution. — Northeastern and Western United States and Canada
from the Atlantic to Pacific.

United States records

California: Shasta Retreat, Siskiyou County, 2 S $, S 9 9 (July 1-7).

Maine: Bar Harbor, 4 $ $ (24-VIII-35; 26-IX-36, A. E. Brewer).

Massachusetts: Worcester, $ (2-VIII-1902).

New York: Ilion, 5,299 (2-3-IX-ll, H. McElhose).

Oregon: Rogue River, Josephine County (VI-1872, Walsingham).

Canadian records

Alberta: Nordegg, S $ S (16-18-IX-1921, K. Bowman),

British Columbia: Kaslo.

Manitoba: $ (lO-VIII-1905, L. E. Marmont) ; Rounthwaite, 9 (lO-VIII-06,

L. E. Marmont).
New Brunswick: Fredericton, $ (22-VII-35, A. E. Brower).
Nova Scotia: Round Hill, $ (28-VII-1935, F. C. Gilliatt).
Ontario: Ottawa, 3 S $ (30-VII-07, ll-VIII-1905, C. H. Young; 17-VII-1933

[reared from Yarrow] C. H. Young) ; Trenton, S $ $ (9-IX-ll, 5-IX-04,

5-IX-12, Evans).
Quebec: Meach Lake, 9 (16-VII-1902, C. H. Young).

Eemarhs. — The specimens I have placed under this name show con-
siderable variation in coloration, but all have one character in com-
mon, white head and thorax. The variation in the color of the fore
wings is of no value in distinguishing this species, and, further, the
color forms cannot be separated on genitalia.

The only North American species with which alienella might be
confused is art&misiella., to which it is very closely related, but the two
may be easily distinguished on characters of the anellus, and artemi-
siella lacks the white head and thorax so characteristic of alienella.

The European eirueriteTla has the thorax white as in alienella., but the
two are easily distinguished by their genitalia.

DEPRESSARIA ARTEMISIELLA McDunnongh

Plate 35, Figukes 196, 196a ; Plate 47, Figuee 278

Depressaria arfanisiella McDunnough, Can. Ent., vol. 59, p. 271, 1927 ; Check list
of the Lepidoptera of Canada and the United States of America (Part 2, Mi-
crolepidoptera), No. 8402, 1939. — Gaede, in Bryk, Lepidopterorum catalogus,
pt. 92, p. 273, 1939.

182 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

Labial palpus ochreous ; base of second segment deep brown and the
brush suffused and profusely irrorated with brown; third segment
with i)oorly defined brown subbasal annulus. Head ochreous mixed
with brown; anterior margin of front deep brown. Thorax deep
purplish brown mixed with ochreous. Ground color of fore wing
light wood brown, shaded with deep brown on costal half at base and
streaked with white and black; subcostal vein black irrorated with
white on outer half; veins 5 to 9 indicated by black scaling; at apical
third a transverse, outwardly curved, crescentic, whitish fascia; at
basal third, in cell, an indistinct black discal spot followed by a white
streak, the latter confluent with the poorly defined white outer discal
spot at the end of cell ; vein 2 largely white with brown shading above
and below ; veins 3 and 4 liberally sprinkled with white, 5 sparsely so ;
from apical third of costa, around termen to middle of inner margin,
a series of poorly defined blackish spots ; cilia concolorous with slight
pink tinge. Hind wing and cilia pale smoky; wing darker apically
with blackish terminal line. Legs ochreous suffused and irrorated
with fuscous except at joints. Abdomen ochreous.

Male genitalia. — Similar to alienella except that the lateral edges of
the anellus of artemisiella are strongly produced (see key and fig. 196).
The clasper of artemisiella appears to be armed with fewer and coarser
spines than that of alienella., but I have not seen sufficient material to
determine whether this character is constant.

Female genitalia. — Similar to alienella except that the anterior
edge of ostial opening is convex in artemisiella but slightly cleft in
alienella.

Alar expanse, 22 mm.

Type. — In the Canadian National Museum.

Type locality. — Seton Lake, Lillooet, British Columbia, Canada.

Food plant. — Arteinisia.

Distrihution. — Known only from the type locality.

Remarks. — In addition to the differences in genitalia the thorax
in artemisiella is not white as it is in alienella and the fore wing lacks
the red coloration of the latter species.

DEPRESSARIA WHITMANI, new species

Plate 36, Figures 200, 200a ; Pl.\te 48, Figure 2S6

A medium-sized dark species similar to leptotaeniae.

Labial palpus with tuft of second segment blackish fuscous below
and outwardly; inwardly and above this segment is olive-buff; ter-
minal segment blackish fuscous; apex olive-buff. Antenna blackish
fuscous narrowly annulated with grayish fuscous; basal segment
shining black above, olive-buff below. Head light fuscous, scales

REVISION OF THE OECOPHORIDAE — CLARKE 183

gray tipped. Thorax and tegula blackish fuscous anteriorly, drab
posteriorly. Fore wing blackish fuscous irrorated with drab and
olive-buff scales; wing, at exti-eme base in angle, three short dashes
at basal third, the bases of veins 9, 10, and 11, and a series of small
spots at the ends of all veins around termen, black; at end of cell a
well-defined olive-buff spot preceded by a few black scales ; at apical
third a poorly defined, narrow, outwardly curved olive-buff fascia;
extreme edge of costa f aintlj^ pink ; cilia grayish fuscous. Hind wing
light fuscous, darker apically; cilia very light shining fuscous with
a darker subbasal line. Legs fuscous strongly overlaid with olive-
buff ; tarsi annulated with olive-buff. Abdomen fuscous, the posterior
edges of the segments dorsally grayish ; under surface heavily over-
laid with olive-buff.

Male genitalia. — Harpe broad, sparsely clothed with hairs; clasper
stout, abruptly excurved just below middle; at base of clasper a
series of four or five short, stout bristles; sacculus narrowly folded
but strongly sclerotized; from base of sacculus an elongate, gently
excurved process armed with fine, slender spines. Anellus a broad
sclerotized plate broader than long; posterior edge narrowly pro-
duced to middle and deeply cleft. Aedeagus longer than harpe,
slender, evenly curved ; at base a broad, flat dorsal winglike expan-
sion; opposite this a broad concave plate by which the aedeagus
articulates with the anellus. Vinculum rounded with prominent
ventroposterior ridge. Transtilla membranous with large, hairy,
broad lateral lobes. Gnathos a spined, oval knob. Socii minute,
mainly indicated by a few hairs.

Female genitalia. — Genital plate narrow, of nearly equal width
throughout; ventroanterior edge produced into a narrow flap.
Ostium small, somewhat triangular. Ductus bursae membranous ex-
cept for a small, strongly sclerotized, curved section from middle to
posterior three-fourths ; entire posterior half rigid and slightly curved ;
inception of ductus seminalis just before ostium. Bursa copulatrix
large; signum a large, sclerotized, toothed plate with prominent
anterior and posterior points.

Alar expanse, 18-22 mm.

7V/^c.— U.S.N.M. No. 52083.

Type locality. — Snake River, Whitman County, Wash., opposite
Clarkston.

Food plant. — Lotnatium macrocarpum (Hook, and Am.) Coult.
and Rose.

Remarks. — Described from the $ type, 2 $ and 1 $ paratypes
all from the same locality. These specimens were reared from larvae
which I collected. The moths issued 26-27-V-1935.

184 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

The species appears to be closest to leptotaenkie in coloration but
nearest angustati on characters of the genitalia. In coloration it is
inseparable from leptotaemae but may easily be separated from
angustati by the presence of the pink cast of the costa which angustati
lacks. D. lohitinani can be separated from leptotaeniae by its long
slender aedeagus ; from angustati by the more evenly curved aedeagus,
the more strongly excurved spine cluster from the base of the sacculus,
the series of four or five short bristles as the base of the clasper, and
the deeply incised posterior margin of the anellus. The females are
at once distinguishable by their genitalia. D. leptotaeniae lacks the
curved, sclerotized posterior portion of the ductus bursae; angu^stati
lacks the produced ventroanterior flap of the genital plate and the
pronounced anterior and posterior points of the signum ; the sclerotized
portion of the ductus bursae is longer in angustati than in whitmani.

DEPRESSARIA LEPTOTAENIAE Clarke

PL.VTE 37, Figures 202, 202a ; Plate 48, Figure 289

Depressaria leptotaeniae CLmArke, Can. Ent., vol. 65, p. 87, pi. 4, 1933. — McDun-
NOUGH, Check list of the Lepidoptera of Canada and the United States of
America (Part 2, Microlepidoptera ) , No. 8396, 1939.— Gaede, in Bryk, Lepidop-
terorum catalogus, pt. 92, p. 286, 1939.

Male genitalia. — Harpe rather short, broad, sparsely clothed with
hairs; clasper stout, long, curved toward the cucullus ; cucullus pointed ;
sacculus narrowly curved, with a prominent transverse, spinous basal
process, the spines numerous, rather small. Anellus a long sclerotized
plate ; posterior and anterior edges deeply concave ; lateral edges con-
vex. Vinculum rounded. Aedeagus stout, pointed, curved. Trans-
tilla a moderately broad sclerotized band with well-developed hairy
lobes. Socii small, hairy.

Female genitalia. — Genital plate narrow with narrow, strongly scle-
rotized ventral sclerites, the latter narrower than the sclerotized part
of ductus bursae. Ostium triangular. Posterior portion of ductus
bursae broad, slightly curved, sclerotized ; inception of ductus seminalis
just anterior to ostium. Bursa copulatrix large with a well-developed
signum near posterior end.

Alar expanse, 17-23 mm.

Type. — In the United States National Museum.

Type locality. — Pullman, Wash.

Food plants. — Leptotaenia rrmlti-flda Nutt. and L. salmom-flora
Coult. and Kose.

Distribution. — Northwestern United States and Canada in the inter-
mountain area.

REVISION OF THE OECOPHORIDAE — CLARKE 185

United States records

Idaho: Viola, Moscow Mouutains (=Thatuna Hills), altitude 3,000 feet, 11 S $ ,
7 $ 9 (10-1&-VI-35).

Utah: Eureka, S, 5 (13-VII-1911, 2S-VI-1911, Tom Spalding).

Washington: Almota, Whitman County, altitude, 2,000 feet, $ (ll-VI-35) ;
Cashmere (Hay Canyon), 2 $ S,7 5 $ (24-V to 7-VI-35, I. W. Bales) ; Dry
Falls, Grant County, 2 $$, 9 (14-V-35) ; Eatiat, $ (9-V-1034, A. N.
Gartrell) ; Grand Coulee City, 17 $ S , 10 9 ? ( 20-22-1 V-34; 12-21-V-35) ;
Kamiack Butte, Whitman County, altitude 3000 ft., 6G $ $ , G3 9 9 (5-13-
V-34; 27-V to 1&-VI-35) ; Park Lake, Grant County, 15 5 5 , 12 9 9
(9^21-V-35) ; Pullman, altitude 2,500 feet, 5 5 5,899 (30-V to 1&-VI-32).
(Unless otherwise stated all were reared from larvae collected by the writer.)
Canadian records

Alberta: Waterton Lakes, 5 (18-VII-1923, J. McDunnough [rf. yellow flowered
umbel] ) .

Remarhs. — In this large series are two specimens (approximately
1 o/o) with short broad wings which otherwise are quite normal and
apparently functional. These two are not crippled in any sense, and
were reared under the same conditions as were the rest, which suggests
that this short-winged form may be the result of a Mendelian factor
such as has been demonstrated in experiments with DrosophUa.

I mention these short- winged forms merely to show that if they were
collected in nature they would undoubtedly be described as a distinct
species (distinct from that to which they actually belong).

Since this species was described I have been able to rear a series of
over 200 moths from various localities in the states of Idaho and
Washington. The Utah specimens are somewhat smaller than those
from Idaho and Washington and show slight variations in the gen-
italia but unquestionably belong here.

DEPRESSARIA YAKIMAE, new species

Plate 37 Figxjbes 201, 201a ; Plate 48, FictrEB 285

A very distinct, medium-sized, grayish species close to leptotaeniae.

Labial palpus with second segment white strongly irrorated with
blackish fuscous; third segment blackish fuscous sparsely irrorated
with white; apex white. Antenna with basal segment blackish
fuscous with a whitish-ochreous patch beneath at apex; remainder
of antenna brown above, whitish ochreous beneath, narrowly an-
nulated with blackish fuscous. Head light brownish fuscous, the
tips of the scales whitish. Thorax and ground color of fore
wing brownish ochreous mixed with white scales. Bases of tegula
and fore wing blackish fuscous; costa of fore wing fuscous mixed
with white; an indistinct longitudinal dash, and streaks following

186 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

the veins, black; at apical third an indistinct outwardly curved,
broken, white fascia formed by short series of white scales following
the veins; around termen a series of seven indistinct black spots;
cilia light smoky. Hind wing fuscous, lighter basally; cilia light
brownish ochreous with a distinct fuscous subbasal line. Legs
blackish fuscous outwardly, mixed with whitish ochreous inwardly ;
annulations of tarsi whitish ochreous. Abdomen gi-ayish above,
whitish ochreous beneath.

Male genitalia. — Harpe broad, sparsely clothed with hairs ; cucullus
broadly rounded; clasper stout, sharply and outwardly angled;
distal end with small scobinations ; sacculus narrowly folded, with
spinous process from base curved toward cucullus; elements of
spinous basal process long, stout, rather few in number. Anellus
longer than broad; anterior margin deeply cleft; lateral edge con-
vex, undulating; posterior margin concave. Aedeagus stout, gently
curved, sharply pointed; base with large, winglike dorsal expansion
and spoon-shaped ventral process by which the aedeagus articulates
with the anellus. Vinculum rounded. Transtilla broad, narrowly
hood-shaped; lobes of transtilla mainly indicated by hairs. Socii
small, hairy.

Female genitalia.. — Ostium large, oval ; genital plate broad, strongly
sclerotized, somewhat evaginated ventrally around ostium ; anteriorly
the ventral sclerites are produced to form two conspicuous, shallow,
sclerotized pockets equal in length to the sclerotized part of the duc-
tus bursae ; ductus bursae strongly sclerotized in posterior third, the
remainder membranous; inception of ductus seminalis just before
ostium; bursa copulatrix large, with a conspicuous, strongly sclero-
tized, toothed signum in posterior part.

Alar expanse, 20-22 mm.

ry;?^.— U.S.N.M. No. 52073.

Type locality.— Ya]nmn, Yakima County, Wash. (18-V-31, Fred
P. Dean, collector).

Food plant. — ^Unknown.

Remarks. — ^Described from the 9 type, 3 $ and 3 9 paratypes as
follows: Walla Walla, Wash., ^ $ $ and 3 9 9 (Clarke, No. 4731-
4736; [8-14-VI-1931, D. H. Brannon]).

This very distinct species cannot be confused with any other de-
scribed from North America. A few species {leptotaeniae, mulfi-
fldae, and angustati) show a slight tendency toward the formation
of the shallow anterior pockets of the genital plate but none has
them so well developed as yahimae.

I suspect that the larva will be found on some species of Lomatiwm
or a closely allied plant.

REVISION OF THE OECOPHORIDAE — CLARKE 187

DEPRESSARIA MULTIFIDAE Clarke

Plate 3G, Figures 1D9, 199a; Plate 48, Figure 288

Dcpressaria multifidae Clarke, Can. Eiit., vol. 05. p. 85. pi. 4, 1933; vol. 66,
p. 179, 1934. — McDuNNouGH, Check list of the Lepidoptera of Canada and
the United States of America (Part 2, Microlepidopteva), No. 8394, 1939.—
Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 288, 1939.

Male genitalia. — Harpe broad, sparsely covered with fine hairs;
cucullus pointed; clasper slender, fingerlike, curved toward cucullus;
sacculus narrowly folded with a prominent basal process armed with
long stout spines. Anellus a sclerotized plate longer than wide, with
sides convex; posterior and anterior margins concave. Vinculum
rounded. Aedeagus longer than harpe, slender, pointed; sharply
curved at distal third, less strongly so at basal third ; base of aedeagus
with a small bulbous process dorsally and a long, broad, shovel-shaped
process ventrally. Transtilla a broad, lightly sclerotized band with
moderately developed hairy lobes. Socii very small, mainly indicated
by hairs.

Female genitalia. — Genital plate narrow with a pair of shallow,
cupped sclerites anteriorly. Ostium a longitudinal slit, wider at
anterior end. Ductus bursae with a sharply curved sclerotized portion
near its middle; inception of ductus seminalis just before ostium.
Bursa copulatrix large with a well-developed signum near posterior
end.

Alar expanse, 17-21 mm.

Type. — In the United States National Museum.

Type locality. — Snake River, Whitman County, opposite Clarkston,
Wash.

Food plant. — Lomatium grayi Coult. and Rose.

DistrihiUion. — Northwestern United States.

United States records

Idaho: Lapwai, Nez Perce County, 23 ^ 5 , 20 ? 5 ( 21-26-V-35 ) .

Washington: Penawavv'a, Whitman County, 6 $ $, 4 9 9 (3-12-V-35) ; Snake
River, Whitman County, opposite Clarkston, 6 5 (J , 10 9 2 (May dates
1932-34) ; Truax, Whitman County, 5,299 (4-9-V-35) ; Godman Springs,
Blue Mountains, Columbia County, altitude 5.700 feet, 2 SS, 4 9 9
(23-31-VII-35).

(All the above specimens were reared from larvae collected by the writer.)
Remarks. — Although the male and fem.ale genitalia were character-
ized in the original description, it seems advisable to make necessary
corrections at this time. They have been correctly redescribed above.
The name of this insect is unfortunate and resulted from a mis-
identification of the host. The plant, upon which the larva feeds,
grows to a very large size on the sandy or gravelly bars of the Snake

188 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 90

River. Because of the large size of these individual plants they have,
for many years, been considered conspecific with the true Leptotaenia
multifida of the high plateau of the Palouse region around Pullman,
Wash.

Suspecting some error in my rearing records of this and allied
species, or in the identification of the hosts, I supplied larvae feeding
on true Leptotaenia with food from the Snake River locality. They
refused this substitute food. Wlien this evidence was obtained further
investigations were carried out which proved, beyond doubt, that the
food plant is Lomatium grayi.

A long series of moths has been reared from larvae collected in
several localities. These moths show some variation in structure and
habits, but not enough, I think, to warrant separation. Superficially
the moths cannot be distinguished, nor can the larvae or pupae.

The differences in habits are directly the result of the variations
in the host plants themselves. As already pointed out, the plants
growing in the sandy or gravelly bars of the river are unusually large,
while those growing on the dry banks a few yards away are scarcely
recognizable as the same species. This plant grows also in the Blue
Mountains of Washington and Oregon up to altitudes of slightly over
6,000 feet. As might be expected it varies greatly in its different
habitats there, from a small, sessile plant to one almost reaching the
proportions of the individuals of the Snake River bars.

The moth, however, apparently does not go above 5,700 feet,
although it is common at that altitude at Godman Springs, in the
Blue Mountains.

The larvae collected at Godman Springs show some colorational
differences but I think these unimportant specifically; one often finds
such differences in larvae from one locality. The larvae from God-
man Springs measured 12-14 mm. in length, 2 to 3 mm. smaller than
the mature larvae from the Snake River. This difference, I believe,
is purely the result of the different environments.

The larval habits of the mountain insects are much the same as
those of the desert-inhabiting individuals. The former differ, how-
ever, in producing much more webbing of the parts of the plants
affected, with pupation frequently taking place in the webbed leaf
sheaths. I failed to find a pupa (living or dead) in the stalks of the
mountain plants although I did find a few larvae in the flower stalks.
All of the larvae found in the stalks were parasitized. Because of
the small number of larvae and pupae found I suspect that many
larvae pupate in debris at the base of the plant. A brief description
of the pupa is as follows : Wing, antennal and leg sheaths bright green
gradually becoming darker and duller, tinged with brown. Abdomi-

REVISION OF THE OECOPHORIDAE — CLARKE 189

nal segments yellow-brown shaded with green. Just preceding
emergence the wing, antennal and leg sheaths become very dark brown
while the abdominal segments remain a deep reddish brown. Last
five segments free. Pupation period 8-10 days.

DEPRESSARIA ANGUSTATI, new species

Plate 36, Figures 198, 198a ; Plate 48, Figure 287

A medium-sized dark species close to multi-fldae and tohitmam.

Second segment of labial palpus ochreous with much blackish
scaling intermixed, especially exteriorly; third segment blackish
fuscous, ochreous tipped. Antenna blackish fuscous, slightly lighter
beneath, the whole becoming lighter toward the distal end. Head
ochreous with much fuscous scaling above. Thorax and fore wing
blackish fuscous with scattered ochreous scales. Tip of tegula and
posterior portion of thorax ochreous. Discal spot at middle of cell
black with some ochreous scales mixed; frequently this spot is very
indistinct; at end of cell a usually conspicuous ochreous spot pre-
ceded and followed by black scales. From the costa, at the middle,
to the center of the cell an indistinct broad ochreous fascia ; at two-
thirds from costa, outwardly to vein 6 then inwardly to inner margin,
a similarly colored but narrower fascia ; costa sprinkled with ochreous
scales above and ochreous beneath mixed with fuscous scales; cilia
light brownish fuscous. Hind wing fuscous, lighter basally; cilia
light brownish fuscous with a narrow fuscous subbasal line. Legs
with femora ochreous, strongly (usually) suffused with fuscous;
tibiae and tarsi ochreous inwardly, fuscous outwardly ; tarsi ochreous
annulated. Abdomen grayish above with ochreous at posterior
edges of segments; beneath fuscous except midventrally where it is
ochreous; anal tuft ochreous.

Male genitalia. — Harpe narrow with the cucullus bluntly rounded ;
clasper strongly sclerotized, moderately slender and strongly curved
outwardly in distal half; sacculus moderately broad with a promi-
nent spinous basal process the elements of which are fine, slender and
compactly associated, Anellus a very broad, sclerotized plate, broader
than long; posterior edge not cleft; from each basolateral edge of
the plate a pointed projection articulating with the sacculus of the
harpe. Vinculum very broad and only slightly curved on its anterior
edge. Transtilla scarcely sclerotized but w^ith well-developed, nar-
row hairy lobes. Aedeagus about as long as harpe, slender, pointed
and sharply curved at basal and distal ends ; at the base are a small
bulbous process dorsally and a broad spoon-shaped sclerotized por-
tion ventrally by which the aedeagus articulates with the anellus.
Tegumen broad, rather short. Socii small.

190 PROCEEDINGS OF THE NATIONAL MUSEUM vol.90

Female genitalia. — Genital plate narrow, more so laterally. Os-
tium small, elongate. Ductus bursae membranous except an elon-
gate, curved, sclerotized area on one side about the middle ; inception
of ductus seminalis just before ostium. Bursa copulatrix large with
well developed signum.

Alar expanse, 17-21 mm.

Type.—\J. S. N. M. No. 52075.

Type locality'. — Skyline Ridge, Mount Baker District, Wliatcom
County, Wash., altitude 6,200 feet.

Food plant. — Lomatium angustatuni (Coult. and Rose) St. Jolin.

Remarks. — Described from the 5 type, 9 $ and 6 9 paratypes all
from the type locality. All were collected by the author. Paratypes
in the United States National, Canadian National, and H. H. Keifer
collections.

The larvae, from which the type series was reared, were collected
on August 17 and 18, 1933. From the larvae collected 25 pupae were
obtained. From these pupae 16 moths emerged. Pupation began
on August 21 and ended August 24; emergence of the moths began
on August 31 and ended September 2.

The host of this species is a small plant growing on open gravelly
slopes or on rock outcrops where considerable fracturing of the basic
rocks has taken place. The basal leaves of the plant are frequently
closely appressed to the ground and it is in these leaves that the larva
does most of its feeding. Frequently, however, the more erect leaves
are involved by the tubes and webs spun by the larva. The silken
tubes in which the larva lives are constructed among the broken
rocks and soil and frequently extend several inches down cracks in
the rock, thus providing an excellent hiding place for the larva when
not feeding.

This species is clearly related to multifidae but may readily be
distinguished from that species by its genitalia.

DEPRESSARIA TOGATA Walsingham

Plate 37, Figure 200B

Depressaria togata Walsingham, Ins. Life, vol. 1, p. 254, 18S9. — Riley, in
Smith, List of Lepidoptera of Boreal America, No. 5282, 1891. — Busck, Proc.
U. S. Nat. Mus., vol. 24, p. 746, 1902; m Dyar, U. S. Nat. Mus. Bull. 52,
No. 5885, 1903.— KEARroTT, in Smith, Check list of the Lepidoptera of Boreal
America, No. 6428, 1903.— Busck, Proc. U. S. Nat. Mus., vol. 35, p. 200,
1908. — Baenes and McDunnough, Check list of the Lepidoptera of Boreal
America, No. 6474, 1917. — Meybick, in Wytsman, Genera insectorum, fasc.
ISO, p. 172, 1922.— Forbes, Cornell Univ. Agr. Exp. Stat. Memoir 68, p. 242,
1923. — IMcDuNNOUGH, Check list of the Lepidoptera of Canada and the
United States of America (Part 2, Microlepidoptera ) , No. 8397, 1939.—
Gaede, in Bryk, Lepidopterorum catalogus, pt. 92, p. 294, 1939.

REVISION OF THE OECOPHORIDAE — CLARKE 191

"Antenna purplish-fuscous. Palpi cinereous, speckled with fuscous
externally on the second joint; apical joint entirely suffused with
fuscous, with the exception of the apex which is ochreous. Head,
dull grayish-ochreous ; face paler. Thorax, cinereous, speckled with
fuscous. Fore wings pale grayish-ochreous, thickly suffused and
streaked with purplish-fuscous, the markings ill-defined, consisting
of a dark fuscous patch at the base of the dorsal margin, a dash of
the same color immediately above the middle of the wing at one third
from base, followed by some pale grayish-ochreous scales; a pale
grayish-ochreous spot on the middle of the wing at about the end of
the cell is preceded and followed by fuscous scales, and beyond and
above it are several fuscous dashes radiating outwards to the costal
and to the upper half of the apical margin, where is a row of obscure
fuscous spots preceding the somewhat paler mottled cilia. Hind
wings, pale shining whitish-gray, with the cilia scarcely darker in
which a slight tinge of grayish-ochreous is traceable. Abdomen,
grayish-ochreous."

Male genitalia. — Essentially like angustati but with the posterior
margin of the anellus narrower and the lateral edge entire, convex.
The clasper is outwardly angulate instead of evenly curved.

Alar expanse, 20 mm.

Type. — In the British Museum.

Type locality. — "Montana."

Food plan t. — Unknown.

Disfrihution. — Known only from the type locality.

Remarks. — The above color description is taken from the original.
I have drawn up the brief description of the male genitalia from a
sketch (see fi