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N. ) THE 


QUARTERLY JOURNAL 


GEOLOGICAL SOCIETY OF LONDON. 


EDITED BY 


THE ASSISTANT-SECRETARY OF THE GEOLOGICAL SOCIETY. 


Quod si cui mortalium cordi et cure sit non tantum inventis herere, atque iis uti, sed ad ulteriora 
penetrare ; atque non disputando adversarium, sed opere naturam vincere; denique non belle et probabiliter 
opinari, sed certo et ostensive scire; tales, tanquam veri scientiarum filii, nobis (si videbitur) se adjungant. 
—Novum Organum, Prefatio. 


VOLUME THE THIRTY-SEVENTH. 
CULL "7 Visas ’ 
1881. KS 


LONDON: LURE Ta 
LONGMANS, GREEN, READER, AND DYER. 


PARIS: FRIED, KLINCKSIECK, 11 RUE DE LILLE; F. SAVY, 24 RUE HAUTEFEUILLE, 
LEIPZIG; T.O. WEIGEL. 


SOLD ALSO AT THE APARTMENTS OF THE SOCIETY; 


MDCCCLXXXI. 


List 


OF THE 


OFFICERS 


OF THE 


GEOLOGICAL SOCIETY OF LONDON. 


Present, 
R. Etheridge, Esq., F.R.S. 


Wice-Prestents. 


John Evans, Esq., D.C.L., LL.D., F.R.S. Prof. J. Morris, M.A. 
J. W. Hulke, Esq., F.R.S. H. C. Sorby, Esq., LL.D., F.R.S. 


Secretaries. 
Prof. T. G. Bonney, M.A., F.R.S. | Prof. J. W. Judd, F.R.S. 


PForeiqn Secretary. 
Warington W. Smyth, Esq., M.A., F.R.S. 


Creasurer, 
J. Gwyn Jeffreys, Esq., LL.D., F.R.S. 


COUNGEIL, 
H. Bauerman, Esq. 


J Gwyn Jeffreys, Esq., LL.D., F.R.S. 
Prof. J. F. Blake, M.A. Prof. J. W. Judd, F.R.S. 
Prof. T. G. Bonney, M.A., F.R.S. Sir John Lubbock, Bart., M.P., F.R.S. 
W. Carruthers, Esq., F.R.S. Prof. N. S. Maskelyne, M.P., M.A., F.R.S. 
Prof. P. M. Duncan, M.B., F.R.S. 


Prof. J. Morris, M.A. 
R. Etheridge, Esq., F.R.S. J. A. Phillips, Esq., F.R.S. 
John Evans, Esq., D.C.L., LL.D., F.R.S. F. W. Rudler, Esq. 


Lieut.-Col. H. H. Godwin-Austen, F.R.S. | Prof. H. G. Seeley, F.R.S., F.L.S. 
J. Clarke Hawkshaw, Esq., M.A. 


Warington W. Smyth, Esq., M.A., F.R.S. 
Rev. Edwin Hill, M.A. H.C. Sorby, Esq., LL.D., F.R.S. 
W. H. Hudleston, Esq., M.A. H. Woodward, Esq., LL.D., F.R.S. 
J. W. Hulke, Esq., F.R.S. 


Assistant-Secretary, Librarian, anv Curator. 
W.S. Dailas, Esq., F.L.S. 


Clerk. 
Mr. W. W. Leighton. 


Librarp and fMuseum Assistant. 
Mr. W. Rupert Jones. 


TABLE OF CONTENTS. 


ANDREWS, Rey. W. R. Note on the Purbeck beds at Teffont .... 251 


BuakE, Rey. J. F. On the Correlation of the Upper Jurassic Rocks 
of England with those of the Continent.— Part I. The Paris 
erasOTPMn Obs NONOVIE hore cs sie oie eile sarees sino) o sie lel ei s'= sige e oie 497 


Bonney, Rev. Prof. T. G. On the Serpentine and associated Rocks 
of Anglesey ; with a Note on the so-called Ser pene of Porth- 
TLE De 6 sin agibroidit: cid OxeRDON IG I ena EDs nee 40 


On a Boulder of Hornblende Picrite near Pen-y-Carnisiog, 
PMNS SVM Ee Petra Pela cio ah nei Stai's, hs = oie ois! at slausiia's: oe) sileyeie ele 5 « 137 


Notes on the Microscopic Structure of some Anglesey Rocks 232 


Bropis, Rev. P. B. On certain Quartzite and Sandstone Fossili- 
ferous Pebbles in the Drift in Warwickshire, and their probable 
Identity, lithologically and zoologically, with the true Lower 
Silurian Pebbles with similar Fossils in the Trias at Budleigh 


ACHE ORAM CM OTIGMILO 21) a ti c/s ta «aid eet olor Sv cack me wo elaieleres ee ene 430 
Buckman, JAmzEs, Esq. On the Terminations of some Ammonites 
from the Inferior Oolite of Dorset and Somerset ............ 57 
Buckman, 8. 8., Esq. A descriptive Catalogue of some of the Spe- 
cies of Ammonites from the Inferior Oolite of Dorset ........ 588 
CatLaway, Dr.CHarites. The Archean Geology of Anglesey; with 
an Appendix on the Microscopic Structure of some Anglesey 
fvecuompy Eton |. (G. Bonney. (Plate VE)... 3... 210 
The Limestone of Durness and Assynt.................. 239 
CarPENTER, P. H., Esq. On two new Crinoids from the Upper 
Chalk of Southern Sweden, GElater Valle arses crates seas oe 128 
CaRRALL, JAMES W., Esq. Notes on the Locality of some Fossils 
found in the Carboniferous Rocks at T’ ang Shan, China ...... 83 
@orewerr, Dr. R. W. On Soil-cap Motion..............0.006: 348 


Davis, JAMES W., Esq. Notes on the Fish-remains of the Bone-bed 
at Aust, near Bristol, with the Description of some new Genera 
207, Syoweresy (Celera 9.0.4 De i ee an nen nen 414 


On Anodontacanthus, a new Genus of fossil Fishes from 
the Coal-measures, with Descriptions of three new Species. 


TElghe OCLL) Seis RRR See inns aioe a aeeientain oi ee 427 


iV TABLE OF CONTENTS. 


Page 
Dawson, Dr. G. M. Additional Observations on the Superficial 
Geology of British Columbia and adjacent regions............ 272 
Dawson, Dr. J. W. Notes on new Erian (Devonian) Plants. (Plates 
GHC a>. C U0) NA aim SAMmin norma oh aboDe do Gs dccooso on cor 259 


Duncan, Prof. P. Martin. On the Coralliferous Series of Sind, 
and its Connexion with the last Upheaval of the Himalayas.... 190 


Dunn, E. J., Esq. Notes on the Diamond-fields, South Africa, 1880 609 


Ecciss, James, Esq. On the Mode of Occurrence of some of the 
Volcanic Rocks of Montana, U.S. AS. ee acer ee 399 


ETHERIDGE, R., Esq. On a new Species of Zrigoma from the Pur- 
beck beds of the Vale of Wardour; with a Note on the Strata 


by the Rey. W. ER. ANDREWS 9 .0).)02 66 «elec ieee eee 246 
. On Plant-remains from the Base of the Denbighshire Grits. 
(Plate XOXW ies ot a ae oeacteio ravens one tiene te BPRS cc i50 6 490 


Hicks, Dr. Henry. On the Discovery of some Remains of Plants 
at the Base of the Denbighshire Grits, near Corwen, N. Wales; 
with an Appendix by R. EruEripeGs, Esq. (Plate XXV.).... 482 


Hotiinewortu, G. H., Esq. Description of a Peat-bed interstrati- 
fied with the Boulder-diift at Oldham .....--.. 72 eeeeee 713 


Hormss, T. V., Esq. The Permian, Triassic, and Liassic Rocks of 
the Carlisle Basin. (Plate X05)... 3. oc). 9. 286 


Jupp, Prof. J. W. On the Occurrence of the Remains of a Cetacean 
in the Lower Oligocene Strata ofthe Hampshire Basin; with an 
Appendix by Prof. BG. Simtiny Pues ts. ee eee 708 


Krrpine, H., Esq., and E. B. Tawnry, Esq. On the Beds at 
Headon Hill and Colwell Bay in the Isle of Wight. (Plate V.) 85 


KEEPING, WALTER, Esq. The Geology of Central Wales; with 
an Appendix on some new Species of Cladophora, by CHARLES 


LAPWORTH, Hsq. (Plate VIL) 9.02.0. ee 141 
KENDALL, J. D., Esq. Interglacial Deposits of West Cumberland 
and North Lancashire. (Plate TMI)... co. a oe eee 29 


LaPworTH, CHAR Es, Esq. On the Cladophora (Hopk.) or Dendroid 
Graptolites collected by Prof. Keeping in the Llandovery Rocks 
of Mid Wales. (Plate VIL)... ......522 08 oe 171 


MackinTosu, D., Esq. On the precise Mode of Accumulation and 
Derivation of the Moel-Tryfan Shelly Deposits; on the Discovery 
of similar High-level Deposits along the Eastern Slopes of the 
Welsh Mountains ; and on the Existence of Drift-zones, showing 
probable Variations in the Rate of Submergence.............. 301 


Moor®, Cuaruzs, Esq. On Abnormal Geological Deposits in the 
Bristol District. 0 5.0.5. 620+ aes oe on ene 67 


Owen, Prof. R. On the Order Theriodontia, with a Description of 
a new Genus and Species (4lurosaurus felinus, Ow.). (Plate IX.) 261 


Description of Parts of the Skeleton of an Anomodont 
Reptile (Platypodosaurus robustus, Ow.). Part II. The Pelvis. 
(Plate) ee oa eee oes 266 


TABLE OF CONTENTS. Vv 


Page 
PaRKINSON, C., Esq. p Upper cope) and Chloritic Marl, Isle of 
Ne Nee hea. ores wi sialcs PATS Ss Stowell aGocuarl Siw Beat 370 
Puitiies, Joun ARTHUR, Esq. Note on the Occurrence of Remains 
gigcecent Plantsiim row? Irom-0re. .32)) ines ce aes Soe wie ee if 
. On the Constitution and History of Grits and Sandstones. 
perma SPU Orme: Meoes uateape aldlai beets cal sky os altho sya) Gai io:4 vise ale «2 36s 6 
REavDeE, T. MELLARD, Esq. The Date of the last Change of Level in 
1 IDOE SIDTRG eo SiS es APO Come SEO ER cP ene BOR ran 456 


RuTLeEY, FRanK, Esq. The Microscopic Characters of the Vitreous 
Rocks of Montana, U.S. A.; with an Appendix by JAMES EccLEs, 
IPT opm OL ALCP NO NG clare teccrs esol, oid agetd Sa cls aie ealocenepeianc ie ptt w ofiere, Sele 591 


On the Microscopic Structure of Devitrified Rocks from 
Beddgelert and Snowdon ; with an Appendix on the ae 
Rocks of Skomer Island. ’ (Plate O05) ete Peet pn see En NO 403 


SrEeey, Prof. H.G. On Remains ofa small Lizard from the Neo- 
comian Rocks of Comén, near Trieste, preserved in the Geological 
Museum of the University of Vienna. (Plate IV.) .......... 52 


The Reptile Fauna of the Gosau Formation preserved in 
the Geological Museum of the University of Vienna; with a 
Note on the Geological Horizon of the Fossils at Neue Welt, 
west of Wiener Neustadt, by Prof. Epwarp SvusEss. (Plates 
ETT OO. Ae ee eee cheno PRS ee Pa aa art 620 


Note on the Caudal Vertebra of a Cetacean discovered by 
Prof. Judd in the Brockenhurst Beds, indicative of a new Type 
allied to Balenoptera (Balenoptera Juddi.)... 1... cece ee eae 709 


SHRUBSOLE, G. W., Esq. Further Notes on the Carboniferous 
“PS ES YSIIEGISS J 5G hte Gasser Me NEy CRANE AEE 178 


Sounas, Prof. W. J. On Astroconia Granti, a new Lyssakine Hex- 
actinellid from the Silurian Formation of Canada ............ 254 


On a new Species of Plesiosaurus (P. Conybeart) from the 
Lower Lias of Charmouth; with Observations on P. megace- 
phalus, Stutchbury, and P. brachycephalus, Owen ; accompanied 
by a Supplement on the Geographical Distribution of the Genus 
Plesiosaurus, by G. F. Wurpporne, Esq. (Plates XXIII. & 


URN Pen cia ee ey ire ai Sia nah a tiene Rane! iz teieie dase the 440 
Susss, Prof. Epwarp. Note on the Gosau Beds of the Neue Welt, 
UE SDROMAY TENCI NGUSCAGCG. «206s syn ceunredials's a Se ds be eernae we 702 


Tawney, EK. B., Esq., and H. Kreprne, Esq. On the Beds at 
Headon Hill and Colwell Bay in the Isle of Wight. (Plate V.) 85 


Ving, G. R., Esq. Further Notes on the Family Diastoporide, 
Busk, § pecies from the Lias and Oolite. (Plate XIX.)...... 381 


Silurian Uniserial Stomatapore and Ascodictya.......... 613 


Waters, A. W., Esq. On fossil Chilostomatous Bryozoa from South- 
west, Victoria, Australia. (Plates XIV—XVII.)............ 309 


vi TABLE OF CONTENTS. 


Page 
Wutpporne, G.F., Esq. Tabular Synopsis of the Geographical 
Distrbucion onthieyieleslosaursrar | easels eee is ciel aes tele ioe renner 480 


WILLETT, Epear W., Esq., and Henry WILLETT, sq. Notes on 
a Mammalian Jaw from the Purbeck Beds at Swanage, Dorset. 376 


PROCEEDINGS. 

Page 
anual’ Reports. cecteeaela t reece) ite oe 8 
duist of Horeien Miembersi). caer eerie eee 16 
List of Foreign Correspondents .........-. +... eee eee eernneeee 17 
List of Wollaston’ Medallists\2 72); oni... 0 0 -isie ere eeee 18 
List of Murchison Medallists. 066.522 «joes aie eee 19 
Taist of lyellt Medallists ie. se ieee ae eeeeeie 20 
Mist ot Biesby Medallists) 2 ee retteeee 21 
Mimancial Report. ce. cre crete peters ellie rie Rents sie “ag eae 22 
Award of the Medals &. 0.05. 6 oie 1. sie she oes ile aie ketene ee 28 
Anniversary Address... 2). ciel eee cine Cle elo eee 37 
Donations to the Library (with Bibliography) .................. 244 
Presentation of Portrait of William Smith? 2.05. oon 2 
Haun, Dr. Otto. On Microscopic Sections of Meteorites ........ 7 


Lones, F. D., Esq. On some Specimens of Diastopora and Stomato- 
pora from the Wenlock Limestone. (Abstract.) ............ 2.39 


Bequest of Drawings of Fossil Fishes by Sir Philip Egerton ... ni 241 


LIST OF THE FOSSILS FIGURED AND DESCRIBED 
IN THIS VOLUME. 


[In this list, those fossils the names of which are printed in Roman type have 
been previously described. ] 


Name of Species. Formation. Locality. Page. 


Incerteé sedis. 


Tubutella ambig“ua .......csceceeceecees |Jurassic &c..,.... [pe Britain’ esse Pvt 
PLANT. 

Aitheotesta devonica. PI. xii. f. 14. \ | (Scotland ...... 306 
Archeopteris, sp. Pl. xiii. f.19 ... | | New Brunswick 305 
Asteropteris noveboracensis. PI. xii. | | 

RNS e so inincio'rsiesivn's «see ce sciees New York ... 299 
Cardiopteris eriana. P\. xiii. f. 18.. New Brunswick 305 
Celluloxylon prim@vum ....0..ee0.00e- | | New York ... 302 
CIOPLCTISN SD! gio cejsicess aise cetee eecie ss | . | New Brunswick 305 
Cyciostigma affine. Pl. xii. f. 11, 12 p PSOE Bac sccu: 4 New York ... 301 
Dicranophylium australicum. Pl. | 

sai Te, Aa ee Australia ...... 306 
Equisetides Wrightiana. Pl. xii. f.10, 

[P| STG 15 7A Dee eee ee New York) .2: 301 
Lepidodendron primevum. PI. xii. 

RBSRPRRC Rae ons Sieacincencinieesedseonss ) \ New York ... 302 
ue ematop ee, I STITHEIN Fagabeose WaleSiteces.c.8 494 
a fier oeamnosa: 4 El xi. } Devonian......... New Brunswick 305 
Pachytheca. P|. xxv. f.7-9 ........ Silurian sceece-e: Wealesm.css.cce: 483 
SPONGIA. 


Astroconia Granti ......... Gonocsdcdden UTTER ear apee | Camnada......... | 254 


Vill FOSSILS FIGURED AND DESCRIBED. 


Name of Species. Formation. Locality. Page. 
C@LENTERATA. 
(Hydrozoa.) 
Acanthograptus ramosus. PI. vii. \ | 
CBO ga cies sanceisiveisislncis sisters gaa femaees ( 174 
Calyptograptus’? digitatus. PI. vii. | : 

dees deaticoccingediansecucnanaeancenee 173 

plumosus. PI. vii. f. 4.......... | 174 
Die tyonema genome TL aL ( Llandovery ...... Central Wales { 

Se Caneee weelseceiacuislemece aaeacieise es | 172 
delicatulum. Pl. vii. f.2 ...... | | 172 
DONUSLUNE aly neville gl gece eectye: el a alee 

Odontocaulis Keepingii. PI. vii. f. 7) L| 176 

ECHINODERMATA. 

Antedon impressa. Pl.vi.f.8,9... |Upper Chalk ...| Sweden ...... 135 

Mesocrinus Fischeri. PI. vi. f. 1, 2: |Planerkalk ...... SaMONYV sess 130 
suedicuss Pl vie fade vasneeeoss Upper Chalk Sweden meee 132 

PoLyzoa. 

Ascodictyon radians «............+-+0. USERS A Sosecoe: Britain 618 
Stellatum: cecjaddesc-cssrseneeene 

Caberea rudis. PI. xviii. f. 86 ...... \ (| 322 

Canda fossitis. Pl. xvi. f. 51, 52... 322 

Catenicella alata. Pl. xvi. f.47, 49, 

OOo Gos Hei aeons mee cece ee er 317 
ampla. Pl. xvi. f. 46, 50...... | 317 
cribriformis. Pl. xvi. f. 39... 317 
elegans, var. Buskii. Pl. xvi. | 

fA 2 AG ee one rae ee e 317 
flexuosa. Pl. xvi. f. 40, 41 . 317 

—— internodia. Pl. xvi. f. 78, 79.. 318 
marginata. Pl. xvi. f. 44, 45.. 317 
soliday selexvie isso Or re ; 318 

Cellaria fistulosa. Pl. xiv. f. 1, 2, MMICBANS Goscesoe: se Australia 

DOV cir eine Biaeataseiy Satara ea eee 319 
qlouusosay abaleexivenie lon lezmee 321 

—  malvinensis. Pl. xiv. f.3...... 321 
ovicellosa. PI. xiv. f. 4-6, Pl. | 

NVI O Qiiciae ecseeaae tee ee coe 321 

Cellepora fossa. Pl. xviii. f. 89 ... 343 
UT ARTIS atic benconecebuebassoe | 343 
SAS DE sSecece tat scrote ce eee eee 344 

Cribrillina dentipora. P\. XY. 1 33. 326 
suggerens. Pl. xvii. f. 75.2... 327 
terminata. PI. xvii. f. és ey (| 326 

Diastopora cricopora. P|. xix. f. 18- 

2 ince dale nase echt Mone tenet ea eee Ooliter2 see Britaines cee 387 

— oolitica. PI. xix. f. 11-14 386 
es Ee a Britain &e. ... | 384 
ventricosa. Pl. xix. f.15-17.. |Oolite ............ Britain’. 2.2.4.9: 385 


FOSSILS FIGURED AND DESCRIBED. 1x 


Name of Species. - Formation. Locality. Page. 


Potyzoa (continued). 


HEMESUCIIAS CLASS onc c.0 eevee. ca cseeee \ 


(| 186 

DUMENSIS 55.25 sas ale sccco deen ig EST 

—— membranacea..................005 Carbonif Britain , 181 

== COS ee ee r |Carboniferous -.. race heey 183 

=== GE 179 

MOLY HOLAGA! 6.0 5os50cedeodeossecace ) 185 
Lepralia cleidostoma, var. rotunda. \ 

Pls SOU, 1 Ee een ee are 336 
corrugata. Pl. xvii. f. 60...... 339 
monilifera, var. armata. Pl. xv. | 

BPM ooo Sclch oaiacis vig ciencicanbawe's 335 
spumulata. Pl. xvii. f. 87...... 335 

Membranipora argus. PI. xiv. f. 20, é 
BUN en oo Sahin tins chew eeaicerinn 24 
GAME MOANA seo sicfoicie's saree <'s/ejeie as 323 
cylindriformis. Pl. xvii. f. 74 323 

concamerata. PI. xiv. f. 22, : 

PME ie icticinc nts aceeiseddesuees 24 
gemmara: Pl. xvi. f..95 ...... | 325 
MCE AMER ees S oSaaiso idee gies 323 
lusoria. Pl. xiv. f. 14, Pl. xviii. 

if, eos soclgonoSO OER CHOEe Ronee ee eeenCe 324 

—  macrostoma. Pl. xiv. f. 18, 4095 

tuaonicar | Ploxiv. £9) ..:...... 329 
WVIGCTONONOPAUMUEA oo. ceccceccssaacess 326 
Microporella enigmatica. Pl. xv. 

i, 2D AD) cocc ene g eee ae eee 331 
clavata. Pl. xviii. f. 84 ...... | 
coscinopora, var. armata. PI. | 

RNS MS OMENS oe a cis vacea cases ceovelen PlVitgeemer wane .ne: S.W. Australia 331 
elevata. Pl. xvii. f. 63, 64, 

TAL, SAT TE) Ree eee a 

3 


ceo Pl xvi, f. 72 ooik oc. .0s 
symmetrica. Pl. xviii. f. 83... 
violacea. Pl. xv. f. 26, Pl. xvii. 


yarraensis. Pl. xv. f. 27, 28... 
Mucronella duplicata. P\. xvi. f. 54 
Clemanseme llevar. f. 91s oo 2... 
mucronata. Pl. xvii. f. 66 ... 
Porella denticulata. Pl. xvii. f. 70.. 
emendata. Pl. xvii. f. 69...... 
Porina clypeata. Pl. xvii. f. 67 ... 
columnata. PI. xviii. f. 88 ... | 
cGononatas “RI xv.t. 97  ..L... | 


Retepora marsupiata. Pl. xv. f. 34— 
36, Pl. xvii. f. 59, 61, 76, 77 


| 

| 

| 

| 

| 332 
| 

| 

| 

\ 


rimata. P\. xvi. f. 48, 53...... 343 
Schizoporella amphora ............4+ 341 
australis. Pl. xiv. f.15. ...... | 341 
conservata. Pl. xviii. f. 81 ... 340 

— excubans. Pl. xvi. f. 56, Pl. 
PMO! Eos) ow eisicecy iocave chess 341 
PMCSEGRLOR Seth S. es oh xtostanwee. ) 339 


Name of Species. | 


FOSSILS FIGURED AND DESCRIBED. 


Formation. Locality. 


PoLyzoa (continued). 


Schizoporella phymatopora. Pl. xv. \ 
[ERO IMBY-) Sham be a remeninsarsr re cen Hors | 
spiroporina 
submersa. 
ventricosa 
vigilans. 
, Sp. 
Selenaria alata 
marginata. 
Smittia anceps. 
centralis 


Peeters eeesetesecsese 


CeCe ere ereeesereese esses 


secrcceee 


Pees ercereoesesssesesscoesesserecr 


Ceo eoeee sere eres secees 


Pixie! 
Pl. xviii. f. 94 


Cece eereSoescrsenes seessseae 


BUGS) 


Cech eereceesc ese sseesceseegsorr oe 


Pl. xvii. f. 65 
Stomatopora dissimilis 
inflata 


eoeoeecee 


eertesroecesoccseeesoeecs cece 


‘Miocene 


S.W. Australia 


oe I 


Silurian 
Silurian 


eoceecocees| BILIUGAILILI w.cccvcces 


Mo.uuusca. 


(Lamellibranchiata.) 


Trigonia densinoda 


seoneseescseesessccae 


|Purbeck 


Dorsetshire ... 


( Cephalopoda.) 


Amaltheus subspinatus ......0....004. 
Ammonites Braikenridgii 
—— Brongniarti 
cadomensis 
concavus 
Gervillii 
—- Humphriesianus 
linguiferus 
— Mansel 


eceveeserecsessesozsece 


eoceec® 08 Cesseoe 


eoceesseessscessss ene see 


eeceocesetoseocc cee ess cs eee 


Sececc SO Oeeeseeses seeceosecce 


@eeeeersccescereecosesossssecsece 


Subraditatusim--eencoscceneceteere 
Cosmoceras Caumontii 
Garantianum 
ParkimsOMlsnn.ccuscee reece Cees 
SOURED So5coabaccasoscocaoce 
Haploceras oolithicum 
Harpoceras adicrum 
concayum 
CONUS Cee s Pena 
— cycloides 
dispalisuimienennsesccsease cece: 
— Edouardianum 
—— fissilobatum ...............00-00- 
evesquellyasr sj. Seki eaccine dese 
Moorei 
Murchisone ......... ane Ne aeane ) 


@eececsccsccses 


e@oeeaeccesssece 


eons ectcoceesecsrce 


Coeee eo ee OOP eeoevesseses 


| 

> ball 

Inferior Oolite...| Dorsetshire ... + 
| 

\ 


| Page. 


338 
340 
340 
338 
338 
339 
345 
344 
337 
337 


337 
337 
615 
615 


| 246 


FOSSILS FIGURED AND DESCRIBED. Xl 
Name of Species. Formation. Locality. Page. 


Motuvsca (continued). 


( Cephalopoda, continued). 


Harpoceras Sowerbyi...............-+. Nei (; 602 
EYLOCCTUS CONFUSUIM .....00005..00000: | | | 601 
——— Budesianum ...........:.-.ses00e 601 
REPRE SUIS aw ores caicie-is\sie se oer «a | || 601 
Oppelia subcostata ......... Re ercniades '! 607 
SPUD ACUI AAS etter o/s wie oh tive sae bos 606 
PRR U ET ea eres = ctclaic cia ciciaicis'o eer ee | 606 
Perisphimctes Martinsii ............... 602 
BV SIESUIN ose se cca caceee ese | 602 
Spheroceras Brongniarti ............ 097 
———— GEMOEPNUM « ...--.2.22-20+eseeee | 598 
Mansel 02022070, F imferior Ootte....- Dorsetshire ...4 | 597 
REMI CMS 80 ce ccioosciciini<seles once: 597 
PoOlyMOTPHUM .2.2...52.....500- | 598 
RMAC eaten sacs fe sic oi's'alvinlon 034s 196 
IAL Dee c cies ou oor eiccie ns ocwe en 599 
Stephanoceras Blagdeni............... | 595 
RRC ONTIG SU). 20... ccc. ccecees ese 1 | 895 
Deslongchampsii ............00. 595 
— Humphriesianum ............... | 594 
Hineaiferm, 82.3. ...6..2.c0ees ee | | 596 
OUVAME TUT caer sci. sxle Saisie 2 ices | 596 
OPO eee see sarees cwess a aeesles ) {| 596 
VERTEBRATA. 
(Pisces.) 
Anodontacanthus acutus. PI). xxii. \ | 
EUR ees sobdiicanciide ccesteeseo- | fe 428 
spies. De sale ‘Coal-measures...) Yorkshire...... 498 
Gurusise ble xxi to 1 2.25%: 428 
Cladodus curtus. Pl. xxii. f.9 ...... \ (| 424 
Ctenoptychius Ordii. Pl. xxii. f. 8. | | 422 
PE CUUDANUIS Hence sc ccs scents oct tes | 424 
Hybodus austiensis. Pl. xxii. f.1... | | | 416 
pumetatus. Pl xxi. f.2-...... )|Rheetic.......c.-.: Bristol) 2.07.00. Slit Wf 
Nemacanthus minor. Pl. xxii. f.5. | | 419 
Paleosaurus Stricklandi. Pl. xxii. | 420 
SPROMPPRRO Ere cco cede ces cess vaste | 
Sphenonchus obtusus. PI. xxii. f. 7 ) \| 420 
( Reptilia.) 
; : re 
ae Sos FL ae Cretaceous ...... | Folkestone ... 634 
Adriosaurus Suessii. Pl. iv........ sei -NEOCOMMNanNn-ee-- | Comén,Trieste 55 
viluxosaurus felinus.- Pl.ix. ...... |KarroO.........s0« South Africa.. 261 
AGLOSAUTUS OTOCUIS ..0..cc00ss.0n see \ (| 700 
Crateomus lepidophorus. P|. xxvii. | | 
foo Pl xxvii. t.9; Pl xxix. \|Cretaceous .....-| Austria......... 4 
foo. blo xxx. f. 2,5; Pl. xxxi. | | 
Lé-5 ,.d er i (| 660 


xii FOSSILS FIGURED AND DESCRIBED. 


Name of Species. 


Formation. 


VERTEBRATA (continued). 


(Reptilia, continued). 


Crateomus Pawlowitschii. Pl. xxvii. \ 
f. 20; Pl. xxviii. f. 2-4; PI. XXIX. 
1 Zh BB Pls 18 Gi, 455 lel 20:0.4l. 


sp. Pl. xxvii.f.9-16 .......-- 
Crocodilus proavus. Pl. xxvii. f. 24- 
26 sPle xxvinits OS) eel exis 


f. 7-13; Pl. xxx. f. 6-14 ......... 
Doratodon carcharidens.......-++++++- 
Emys Neumayri. Pi. xxvii. f. 273 | 

Pixxx 6s, Plbsexxtstelioncse (" 
Hoplosaurus ischyrus. Pl. xxx. |» 

Dich ascliasnesicsaishs ac menmotrancsctstes 
Megalosaurus pannoniensis. P). xxvii. 
£223 reese once soeeee css: 
Mochlodon Suessii. P1. xxvii. f. 1.. | 
Ohigosaunustadelust penser reece 
Ornithocheirus Bunzeli ..............- 
Ornithomerus gracilis. Pl. xxviii. | 

POs sseet aoslecteos Oameee ceaae y) 

Platypodosaurus robustus. PIl.x.... 


Plesiosaurus brachycephalus. PI. \ 
D:D IN IY linen Nopenana) aon coddo stone | 
Conybeart. PI. xxiil., Pl. xxiv. } 


Cote ert cce ere ees ee eee COS Fees eroeceee 


megacephalusaseeneeeteeeeeen eee ) 
Pleuropeltus Suessiz, Pl. xxviii. f. 8, \ 
Os PS xoxixe of 0165 3 ea ee etree a | 
Rhadinosaurus alecimus. Pl. xxxi. | 
EG NO 3 eee cadence tae eee r 
Struthiosaurus austriacus. Pl. xxvii. | 
56 em ee 


’ 
(Mammalia.) 

Balenoptera Juddi.........cccccececeee Oligocene 

Ariconodonemordaxt esses. see Purbeck 2-2 


Cretaceous 


Cretaceous 


eecoeeeos 


Locality. 


Austriaeee eres 


South Africa.. 


Dorset 


Hampshire ... 


Dorset 


eeecccee 


~~ SS - SF 


Page. 


EXPLANATION OF THE PLATES. 


PLATE PAGE 
I. ) Microscoric Sections or Grits, and GraArns or Sanp, to il- 
100 instrates Mire dean milli pss paper ea secteessss-eeee sek ee = 6 


i Maps, Plans, and Sections of InrerauactAnL Deposits oF 
Til West OCumMBERLAND AND Nortu LANCASHIRE, to illustrate 


Mie ey Wendallignpaper  vannciensuiaecesnesesrecsssesce «ses acts. 29 
IV. f ApriosauRuS Sugssti, to illustrate Prof. Seeley’s paper on 
TL Gust TR ecb ae SO A eee 52 
SECTION OF THE CLIFFS IN ToTLAND AND CoLWELL Bays, 
v Iste or Wicut, to illustrate the paper by Messrs. Keep- 
mene t Cl Meh we MCI eee esac seta iapisicjotie wsiaielaclan “ccianisteeideeiisiansise.«s 85 
VI SwepisH Cretaceous Crinoips, to illustrate Mr. P. H. 
Wan Penber spa Peli eee ssc een ess ee caaceoecsc- Sens steoasae 128 
yy, { Weusn, Crapornora, to illustrate Mr. C. Lapworth’s ap- 
; pendix-to Mr: Walter Keeping’s paper .::.-..0.:--+..------0-- Wal 
VIII “Map anv Secrions or ANGLESEY, to illustrate Dr. Calla- 
‘ { WW LVES [DL CLAN sctueetccctoeeateia Mama nisme'sisl oleltccia-iale/sisiisiniciaaiierse ssitslesaicioe 210 
1x, J Hvvrosavrvs rerinvs, to illustrate Prof. Owen's paper on 
Elberta pbtlor mc racamemtesss sodeeaticenceesceteac aot sans suena toes 2 
x, { Puarypoposaurus RoBustus, to illustrate Prof. Owen’s paper 
Gin, WANG TSO hes waaneBa ston ynan dakke Lacea saeO Bec DRARUECE nem eoneaee 266 
XI Map sHowING THE Rocks OF THE CARLISLE Basin, to illustrate 
Mir eV eOlmeaiskpaner., a taenne ates tae sete seo. wesaee. 286 
Sate | Drvonran Prants, to illustrate Dr. J. W. Dawson’s paper... 299 
XIV. \ 
ae | Victorian Fossint Bryozoa, to illustrate Mr. A. W. Waters’s 
XVII. | } 02) 02 SSS SoS COCO OBOOS OOS SO COO COCOOO Sc OOOO COCOUC KOO GeOSoUOUnCOnOSOomon: 309 


XVIIL ) 


X1V EXPLANATION OF PLATES. 


PLATE PAGE 

JuRAssic DriAstoporrps, to illustrate Mr. G. R. Vine’s 

XLEXG 

DAPOL" seis ages ouacaninnk Getinamas eaten neice ean an eaeena cece ee eeenee 381 

Xx Microscoric Sections or Virreous Rocks or Montana, to 
illustrate; Marsh Roubley(s) papers... accent hee eeeeeeee 391 

XXI Microscoric Structure or WetsH LAVAS or Low&r SILURIAN 
ae Agu, to illustrate: Mir. Ee Rutleys paper) 2. ...s-4-e4-eeeeeees 403 

XXII FHISH-REMAINS FROM THE Aust BoNE-BED AND THE COAL- 
MEASURES, to illustrate Mr. J. W. Davis's papers ...... 414,427 

XXIII. ) Puestosaurus ConyBEARI AND P. BRACHYCEPHALUS, to illustrate 
XXIV Prof. W. J. Sollas’s paper on those reptiles .................. 440 


XXV. Sinurtan Piant-remarns, to illustrate Dr. Hicks’s paper ... 482 


ComPARATIVE Sections or Upper JURASSIC Rocks In FRANCE 


XXVI. | AND EnGuanD, to illustrate Prof. Blake’s paper ............... 497 


OTE \ 


VIII. 
XXVIII. | REMAINS oF REPTILIA FROM THE GosAu ForMATION AT WIENER 


a f Nevsrant, to illustrate Prof. Seeley’s paper.................. 620 


OO) 


ERRATA ET CORRIGENDA. 


Page 197, line 12 from bottom. for these read there. 

Pages 305 and 307, line 10 from bottom for Peny read Perry. 

Page 306, line 15 for Bodun read Brown. 

Page 307, line 30 for magnacensis read gaspiensis, and delete footnote. 
Page 326, line 15 from bottom, for DENTIPOR read DENTIPORA. 

Page 406, explanation of Fig. 1, for Fisst/e read Felsite. 

Page 597, line 3 from bottom, for Mansrniui read MANSELII. 

Page 597, line 2 from bottom, for Mansellii read Manseliv. 


THE 


QUARTERLY JOURNAL 


OF 


THE GEOLOGICAL SOCIETY OF LONDON. 


Vou. XXXVII. 


‘1. Nore on the Occurrence of Remarns of Recent Pants in 
Brown Iron-orzr. By J. Anruur Purtiirs, Esq., F.GS. 
(Read November 3, 1880.) 


Tuts bed of fossiliferous iron-ore is situated at Rio Tinto, in the 
province of Huelva, Spain, and is in close proximity to the cele- 
brated copper-mines of that name. 

In this portion of Southern Spain deposits of cupreous iron 
pyrites, consisting of a series of lenticular masses of ore, having a 
general direction a little north of east and south of west, extend 
from Aznalcollar, near Seville, in the east, for a distance of more 
than seventy miles westward to within the Portuguese frontier. 

At Rio Tinto the deposits of this mineral are very extensive, and 
consist of a compact and intimate admixture of iron pyrites with a 
little copper pyrites, through which strings of the latter mineral 
- sometimes ramify. 

Although these mines appear to have been worked, and the copper 
smelted upon the spot, from time immemorial, it is evident from the 
vast heaps of furnace-slags, and from the extent of the various other 
remains in which coins and inscriptions of the reigns of the 
Emperors from Nerva to Honorius have been discovered, that their 
great development under the Romans took place during the first four 
centuries of the Christian era. After the fall of the Roman empire 
they seem to have been abandoned down to as late as the year 1727, 
from which date they were intermittently worked by the Spanish 

Q.J.G.8. No. 145. B 


2 J. A. PHILLIPS ON THE OCCURRENCE OF REMAINS 


Government and by various private speculators until 1873, when 
they were purchased by an English company. The extent of the 
mining and metallurgical operations anciently carried on in this 
district will be understood when it is stated that at Rio Tinto alone, 
in addition to hundreds of Roman shafts and miles of Roman 
galleries, the heaps of copper slags resulting from the smelting at 
that period cannot amount to much less than one and a half million 
of tons, and that there are large accumulations of similar ancient 
refuse at Tharsis, Buitron, and omen mines. 

As illustrating the care and skill of the metallurgists of that period 
it may be stated that each ton of their slags seldom contains above 
three pounds of copper. 

The prevailing rock throughout this region is clay-slate, which, 
from the evidence of various fossils found by Mr. A. Hill, Mr. 
G. W. Clement, and other officers on the staff at Rio Tinto, is 
apparently of Silurian age. These specimens were kindly examined 
by Mr. R. Etheridge, who did not hesitate to identify them as 
belonging to that period. 

These slates are, in places, broken through by large dykes of 
quartz-porphyry, which frequently form one of the walls of the 
various deposits of cupreous pyrites. 

The fossiliferous iron-ore which is the immediate subject of this 
note forms a cap one kilometre long, with an average width of one 
hundred and thirty metres, on the top of the Mésa de los Pinos, 
nine hundred metres south of the open cutting at Rio Tinto. Its 
surface is approximately level, but it varies m depth from one to 
seventeen metres in accordance with the conformation of the surface 
of the slate upon which it lies ; the rock beneath it is bleached and 
to some extent decomposed. 

The order and relative positions of the several formations will be 
best understood on referring to the accompanying plan and section 
(figs. 1, 2), for which I am indebted to Mr. Neil Kennedy, a gentile- 
man in charge of a portion of the work at the mines, through whose 
kindness I am also enabled to lay specimens of the fossiliferous iron- 
ore upon the table. 

On the extreme right of the section is a broad porphyritic dyke 
forming the north wall of the south lode, next to which is the lode 
itself, which at this point has only one third of its greatest width. 
iNest in succession, to the south, comes a band of slate, which 
is again penetrated on the left by a broad dyke of quartz-porphyry. 
tt will be observed that the upper part of the vein has, to. a con- 
siderable depth, been converted into a ferruginous capping (gossan), 
of which a large portion has been removed by denudation. The 
stratum of iron-ore forming the surface at the Mésa de los Pinos 
is shown with precipitous sides; and a small patch of a similar 
formation occurs, at within a metre of the same elevation, at the 
Cerro de las Vacas. Numerous fissures occur in the surface of the, 
larger deposit of iron-ore, and out of these pine-trees formerly 
grew in considerable numbers, their presence giving the name to 
the locality ; these were eventually destroyed by sulphurous fumes 


OF RECENT PLANTS IN BROWN IRON-ORE. 3 


Fig. 1.— Plan of part of Rio Tinto. 


SY 


i ope SA L/w 
ee ay ROWE 

Y Ws 
MWg Cory, iy ae ,, = —. Ce TTOS¢ loan — 


Phe (E Z LUN 
. ) INEZ 


a ed 


: 1 
Seale of Plan and Section 15,000 


Fig. 2.—Section in line AB on Plan. 


p.W N.E. 


South Mésa de Rio Cerro de South Cerro 
Dam. los Pinos. Agrio. las Vacas. Lode. Salomon. 


ee 


‘ 
{ 
1 
1 
1 
1 
i 
1 
t 


aA Mian Ah) Ten YN OPEN DO 
Th I] i) “ TMM HH] ISO 
SUIS Hi HUH i lh HH MEMS MA) SESSEAIS S/R AEN 


See) 


7:'| Capping of lode. 


Mineral. 


resulting from the metallurgical operations which were carried on 
upon the declivity of the opposite hill. 

In some instances the decomposed clay-slate has been partially 
removed from beneath the iron-ore, from which blocks have been 
detached, and have slidden a considerable distance down the surface 
of the slaty declivity. 

A specimen of this iron-ore, analy a in my laboratory, afforded 
the following results :— 


BQ 


4 J. A, PHILLIPS ON THE OCCURRENCE OF REMAINS 


(Ghivesromlebnic me iar ieee 1:40 

Wis | Ree RO SAIa Tren 11°85 
ISU CAF Baro anno ei oo bod 6:5 1:53 
MerniCloxide ies. ener wm iw 84°65 
AU TAaMAT TG 3, coe eueoraye cs puede eset ee trace. 
Phosphoric anhydrides - 4. -4e)e- 14 
PiU 0) COU Ree AA anc ACA as cach ene 23 
99°80 


It follows that this is a rich ore of iron of fairly good quality ; and 
as there is now a well-appointed railway in its immediate proximity, 
it is probable that under moderately favourable conditions of the 
iron trade it may be worked with advantage. 

During the process of quarrying this ironstone for exportation, 
Mr. Kennedy first observed the presence of fossil remains of what 
appeared to be leaves and seeds of many of the plants still growing 
in the neighbourhood, as well as of several well-preserved beetles. 

All doubt as regards the recent character of these fossils is, how- 
ever, removed by the following communication from Mr. W. Carruthers, 
who kindly undertook their examination, and who says, “‘ The speci- 
mens you have sent me for examination from Rio Tinto contain the 
following fragments of plants which I have been able to identify :— 
leaves and acorns of Quercus Ilex, Linn.; leaves and seed of a two- 
leaved species of Pinus, most probably Pinus Pinea, Linn.; the cone 
of Equsetum arvense, Linn.; and a small branch of a species of Erica. 
There is also a well-marked leaf of a Dicotyledonous plant which 
I have not yet been able to identify. The greater portion of two of 
the specimens consists of a thick growth of moss, but it is impossible 
to say what the species are. ‘The whole is permeated with minute 
branching roots, showing that the vegetation was formed as a peat- 
moss, the oak- and pine-leaves being carried or blown into it. The 
plants are evidently, all of them, the same species as are still found 
growing in Spain.” 

In addition to these fossils this deposit sometimes aorta minute 
concretionary patches of imperfectly crystallized quartz. 

Every one who examines the section extending from the Cerro 
Salomon across the valley of the Rio Agrio to the Mésa de los 
Pinos will probably agree that the origin of this deposit of iron-ore 
can scarcely be doubtful. At the time of its formation a marsh or 
shallow lake extended from beyond the last-named point to the foot 
of Salomon, and into this flowed solutions of iron-salts resulting 
from the decomposition of the upper portions of the immense masses 
of pyrites constituting the south lode. 

From these salts oxide of iron was deposited, as in the case of 
bog iron-ores generally ; and, finally, the valley of the Rio Agrio was 
eroded, as well as that south of the deposit, leaving the Mésa 
capped with iron-ore, while a small patch of the same mineral 
was left at the Cerro de las Vacas. 

That the deposit took place at a comparatively recent date is 
evident from the fossils it contains; and it is equally certain that the 


OF RECENT PLANTS IN BROWN IRON-ORE. 5 


erosion of the valley is older than the occupation of the district by 
the Romans. Not only are numerous remains of buildings and other 
works belonging to the Roman period found in the valleys, but the 
Roman grave-stones, of which scores are still scattered over some 
parts of the district, are invariably made of this iron-ore. 


Discussion, 


Mr. CarrurHers pointed out that the deposit, though of such 
thickness and importance, was essentially a bog iron-ore and, like 
ores of that character, contained remains of recent plants. 

The Prestpent remarked on the difference in appearance of mas- 
sive hematites from ordinary bog iron-ores and those brought by the 
author of the paper. 

Dr. Sorsy asked if it were possible that these could have been 
deposits produced by springs, such as occur on the hill-sides of 
Yorkshire, and are still forming. 

Mr. Parrison stated that the bog-ore deposits in Ireland occur 
on the sides and summits of hills, and that the Rio-Tinto deposit is 
a true bog iron-ore. 

Mr. Bavprman remarked on the resemblance between this ore 
and that of Arklow, which is also derived from pyrites ; and con- 
sidered that the freedom from phosphorus was to be attributed to 
their rapid and direct formation as compared with ordinary bog 
iron-ore, which is contaminated by accessory products of decompo- 
sition from organisms and rock-masses. 

The AvutHor, in reply to Dr. Sorby, stated that the top of the 
deposit was regular and the bottom irregular, leading to the belief 
that it was probably a deposit in water. Further he indicated that 
there is an outlier on the same level. The hard cap is sometimes 
undercut by denudation. He remarked that springs are scarce over 
the whole district, and that there are none containing more than 
traces of iron in solution. 


6 J. A. PHILLIPS ON THE CONSTITUTION AND 


2. On the Constitution and History of Grits and SanpstonEs. By 
J. Antour Puittres, Esq., F.G.8. (Read December 15, 1880.) 


[Puares I. & IT] 


Te careful and exhaustive researches relative to the constitution 
and mode of formation of arenaceous rocks which have recently been 
published by Professor Daubrée, Dr. Sorby, and others, leave open a 
comparatively restricted field for the pursuit of similar investiga- 
tions. Having, however, during the last two years paid considerable 
attention to the study of rocks of this class, I now venture to bring 
before the notice of the Geological Society some facts and deductions 
- therefrom which would appear to have escaped the attention of 
previous observers. 

For the convenience of developing certain ideas relative to this 
subject, I propose in the present paper, in the first place, to describe 
various grits and sandstones which have been microscopically and 
otherwise examined. In doing this the older rocks will be con- 
sidered first, and those of more recent age subsequently noticed in 
the order of their geological sequence. The chemical composition 
of some typical rocks will also be given. 

Secondly, the results of observations bearing on the effects pro- 
duced by the action of flowing water on particles of sand and gravel 
transported thereby will be described ; and finally, the more impor- 
tant observed facts will be summarized,.and their bearings discussed. 

The difference between grit and sandstone is one not always 
distinctly marked ; and numerous definitions of the two rocks, often 
somewhat contradictory, have been given by various geologists at 
different times. It has even been stated by an eminent authority 
that rocks which in the north of England would be called grits, 
receive the name of sandstones in the south *, 

In order, therefore, to avoid misunderstanding upon this point, 
I may state that in the following descriptions the term grit is 
applied only to coarse-grained arenaceous rocks of which the com- 
ponent fragments are for the most part angular, and which, although 
frequently crystalline in structure, seldom contain either perfect or 
nearly perfect crystals. The cementing material of such rocks is, 
as a rule, highly siliceous. 

Sandstones differ from grits in being finer in ‘structure than the 
latter, and in their component ¢ erains being usually less completely 
incorporated with the cementing median The quartz in many 
sandstones occurs principally in the form of perfect erystals, or in 
that of crystalline aggregations. 

In quartzites the spaces between the component grains are com- 
pletely filled by a siliceous cement, in which respect they closely 
resemble some varieties of fine-grained grit. 


* “Manual of Geology,’ by John Phillips, F.RB.S., p. 654. 


Quart. Journ.Geol. Soc Vol. XXXVII. P11. 


imp. 


Mimtrn Bros. 


8 


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del. 


Frank Rutl ey, 


vi CMON Ob GRIMS: 


AN ee 


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: 


~ HISTORY OF GRITS AND SANDSTONES. 


NI 


Composition and Structure. 


Cambrian.— The well-known Barmouth Grits of North Wales, 
which occupy a large extent of country lying between Barmouth 
and Harlech, are usually of a greenish-grey colour. They are ex- 
tremely hard, and often enclose angular fragments of quartz above 
a quarter of an inch in diameter. Sometimes these grits become 
fine-grained, and assume a purple tint ; they are intermingled with 
occasional bands of greyish-green and bluish slates, which, especially 
towards the lower part of the series, attain considerable develop- 
ment. Of these rocks Professor Ramsay remarks :—‘“ The beds seem 
to have been formed principally by the direct waste of rocks of a 
granitic character, or at least into the composition of which crystal- 
line quartz and felspar largely enter ”*. 

When a thin section of this rock is examined under the micro- 
scope, it is seen to consist, mainly, of an aggregation of fragmentary 
quartz and felspar united by a siliceous cement, which is throughout 
permeated by a moss-like greenish mineral, a portion of which is 
probably chlorite. ‘The larger pieces of quartz and feispar are often 
distinctly rounded, although they also sometimes present irregular 
and perfectly sharp outlines. Hig. 1, Pl. 1I., drawn by Mr. F. Rutley, 
represents, in black and white, a section of this rock as seen in 
polarized light, magnified 18 diameters, and containing much felspar. 

The quartz occasionally contains liquid-cavities enclosing moving 
bubbles, but these are by no means numerous; the greenish mineral 
of the cement sometimes penetrates into fissures 1n the siliceous 
grains. ‘Two distinct species of felspar are present in considerable 
quantities, the larger grains being, for the most part, somewhat 
rounded fragments, which, after having assumed the form of 
pebbles, have sometimes been broken across their smaller dia- 
meter, thus presenting one angular and one rounded termination. 
The orthoclase is not much altered; and a triclinic felspar, which, 
from the optical properties it exhibits, is probably oligoclase, shows 
brilliant lines of twinning when seen in polarized light. Some 
of the quartz encloses hair-like crystals of rutile, while calcite, 
magnetite, iron pyrites, and a few imperfect garnets are present in 
the cementing siliceous base. An analysis of this rock is given, 
page 21. 

The grits in the neighbourhood of Harlech are usually finer in 
grain than the foregoing, but otherwise differ from it only in con- 
taining a few imperfect crystals of epidote. 

Stlurvan.—Stiper Stone, from the neighbourhood of Shrewsbury, is 
a fine-grained and exceedingly hard sandstone, the grains of which 
are so closely cemented together by crystalline silica as to form a 
quartzite. Many of the fragments of quartz, of which this rock 
is mainly composed, are somewhat cloudy and are considerably 
rounded, while others are colourless and transparent, with but few 
fluid-cavities, which are, for the most part, full. The average dia- 
meter of the grains is about +, inch, and some of the most pebble- 


* Geology of North Wales, p. 17, ed. 1. 


8 J. A. PHILLIPS ON THE CONSTITUTION AND 


like among them form the nucleus of a crystalline growth of colour- 
less transparent quartz, which converts them into more or less perfect 
crystals of that mineral. Felspar is not abundant, and is often 
considerably altered ; the cementing material occasionally contains 
minute crystals of a mineral which is probably epidote. 

A grit of a grey colour, speckled with minute white points, from 
the neighbourhood of Aberystwith, is seen under the microscope to 
be composed of an almost equal amount of quartzose and felspathic 
grains united by a siliceous cement, everywhere permeated by a 
moss-like chloritic mineral, to which reference has already been 
made. In this rock the constituent fragments are sometimes as much 
as = inch in length, and their angles are usually to a certain extent 
rounded, although they occasionally exhibit very sharp and irregular 
outlines. The quartz, which exceptionally contains a few needles 
of schorl, is colourless and transparent, containing exceedingly 
minute fluid-cavities, of which the majority are full, although others 
enclose constantly moving bubbles. 

The felspar has been subjected to considerable alteration, and is 
not unfrequently obscured by flocculent microlths resulting from 
chemical re-combinations ; a portion of it, however, evidently belongs 
to a triclinic species. In addition to the above, there are inclusions 
which are undoubtedly fragments of a volcanic rock of basaltic 
character. 

The siliceous cement contains a few crystals of iron pyrites, as 
well as small flakes of brown and colourless mica, of which the 
edges are much rounded. 

It is evident that some portion, at least, of the quartz constituting 
this grit has been derived from the disintegration of quartz-felsite 
(quartz-porphyry), since its general characteristics are not only 
similar, but it, moreover, includes the blebby masses of an ater ee 
ground-mass SO characteristic of the quartz of such rocks. 

A fine-grained foliated rock belonging to the same series, from 
ae ae Botmetmnes locally called a grit, of which the grains 
vary from <4, to zgbp inch in diameter, has a composition gene- 
rally similar to that of the foregoing. In addition, however, it con- 
tains numerous water-worn flakes of mica, which occur chiefly in 
distinct bands, and which are arranged with their cleavage-surfaces 
parallel to the plane of foliation of the rock. The fragments of 
quartz, which are of a nearly uniform size, are all sharply angular, 
and elongated or flattened grains are rare. 

An examination of sections prepared from May-Hill Sandstone, 
containing numerous casts of Pentamerus oblongqus, aoe that this 
rock is mainly composed of angular grains about z00 inch in dia- 
meter, united by a turbid siliceous cement, suggesting the idea of 
its “haraime been deposited from waters holding clay in suspension. 
In addition to the smaller grains, of nearly equal dimensions, there 
are a few fragments of, at least, four times the size above stated. 
These, like the smaller ones, sometimes contain a few hair-like 
crystals of rutile; but fluid-cayities containing bubbles are exceed- 
ingly rare. 


HISTORY OF GRITS AND SANDSTONES. 9 


Sandstones, or rather quartzites, of Upper-Llandovery age are 
developed in the Lower Lickey hills in Worcestershire. A specimen 
from this locality was found to be chiefly composed of much-rounded 
grains of quartz, having an average diameter of = inch, cemented 
together by a growth of transparent crystalline quartz. "This mode 
of | formation is rendered evident by the circumstance of the rounded 
grains being frequently composed of cloudy or slightly turbid quartz, 
while the cementing silica is perfectly colourless and transparent. 
When examined in polarized light, the cementing quartz is seen to 
exhibit, fora certain distance around each grain, the same colour as 
the grain itself, and appears to have been deposited in crystalline 
continuity therewith. Fluid-cavities, with bubbles, are numerous in 
some of the grains, while others are entirely without them. Rutile 
and schorl are sometimes present in the form of minute crystals. 

A specimen of Denbighshire grit from Pont Cletwr Yspytty, 
when examined, was found to consist of an exceedingly fine-grained 
mosaic of cementing concrete containing minute granules of quartz, 
and enclosing larger fragments of the same miner fle with Hel sien and 
colourless or brown mica, each grain being from = to >j> inch in 
diameter. Some of the quartz fragments are so traversed by the 
moss-like greenish mineral, often forming a constituent of the cement 
of sandstones, as almost to suggest the idea of their being pseudo- 
morphs after a mineral which has disappeared. If seen in polarized 
light, however, they will be observed to be each made up of several 
distinct grains, in the fissures between which the substance referred 
to has obviously been deposited. The quartz of this sandstone occa- 
sionally encloses a few needles of schorl or hair-like crystals of 
rutile ; cavities containing bubbles are rare. 

A rock belonging to the Coniston-Grit series from Green-quarter 
Feli, Westmoreland, consists of angular grains of quartz and felspar, 
united by a siliceous cement traversed in all directions by numerous 
greenish BEerouths. The average size of its constituent particles 
does not exceed 5,45 inch, allow there are a few larger ones, 
measur.ng about 1, inch in came cen! The quartz contains on fluid- 
- cavities with bubbles ; but when these occur they are extremely 
minute. This rock contains a little iron pyrites, and the cement is 
sometimes stained by hydrated ferric oxide; a few flakes of colour- 
less and dark-brown mica are occasionally seen between the grains 
of quartz and felspar. 

Devonian.—The majority of the siliceous grits of Cornwall are 
usually regarded as being of Devonian age; but it is probable that 
some of them may be of older date. 

-Two distinct beds of such rock, of a greenish-grey colour, which 
are worked for road-metal, are quarried on the farms of Tregian and 
Dairy, in the parish of St. Ewe, near St. Austell, and were noticed 
in a previous paper under the name of slaty agglomerates*. 

In both these localities the grit contains angular fragments of a 
soft clay-slate of a greenish-blue colour, and is exceedingly hard 
and tough. The rock quarried at Tregian is composed of a mixture 

* Quart. Journ. Geol. Soc. 1878, vol. xxxiv. p. 476. 


10 J. A. PHILLIPS ON THE CONSTITUTION AND 


of angular pieces of quartz and felspar, of which some of the 
larger fragments have a diameter of =, inch; they are cemented by 
a siliceous concrete enclosing particles of granular quartz, through 
which minute greenish microliths are plentifully disseminated. 
Comparatively few fluid-cavities, either with or without bubbles, 
are present. In addition this rock contains a few crystals of schorl 
enclosed in the quartz, some water-worn flakes of silvery-white 
mica, a few crystals of pyrites, and perhaps a little altered mag- 
netite. The felspar chiefly belongs to a triclinic species, but ortho- 
clase is also present. The rock at Dairy differs from that at Tregian 
only inasmuch as it contains a few water-worn crystals of horn- 
blende and a little magnetite. 

In addition to the foregoing, through the kindness of Mr. J. H. 
Collins, I have been enabled to examine four other specimens of 
Cornish grit, namely, one from St. Allen, four miles north of Truro, 
two from Ladock, five miles further east, and one from Perranzabuloe, 
on the Bristol Channel. 

Hand specimens of all these rocks closely resemble one another, 
excepting that those from Ladock enclose numerous angular frag- 
ments of a greenish slate, which the others do not, and that one of 
them contains a number of rounded quartzose and other grains 7 inch 
in diameter. 

When examined under the microscope, the St. Allen grit differs 
little from those at St. Ewe; the quartz is angular and transparent, 
the largest fragments having a diameter of about = inch, and they 
sometimes, though rarely, enclose minute crystals of tourmaline. A 
little hornblende, with white mica and epidote, are also present. 
The felspar is, to a large extent, triclinic, but there is also some 
altered orthoclase ; the quartz contains but few fluid-cavities. 

In the rock from Ladock, which contains small rounded grains of 
quartz, felspar, and other material, these bodies are sparsely dissemi- 
nated throughout the mass of the normal grit; and a microscopical 
examination shows that some of them are fragments of volcanic 
rocks closely akin to the “‘ greenstones” and “ dunstones” of many 
parts of Cornwall, but which have often become so altered as to be 
recognizable only by their felspars and general structure. 

Fig. 2, Pl. I., represents, in black and white, a fragment of volcanic 
rock which occurs in this grit, as seen in polarized light, magnified 
18 diameters. 

The second specimen from this district is made up chiefly of angular 
fragments of quartz and felspar, united by the usual cementing con- 
crete. It contains a considerable amount of felspar, a large propor- 
tion of which is triclinic. The largest pieces of quartz are about 
;'z inch in diameter ; and fluid-cavities, although by no means abun- 
dant, are more plentiful than they are in the Cornish grits before 
described. Some of the quartz contains a notable quantity of dis- 
seminated epidote (?), and flakes of white mica are frequently jammed 
between the fragments of which the rock iscomposed. A few minute 
garnets are present, as well as some fragments of a volcanic rock. 

The microscopical structure of the gritty rock from Perranzabuloe so 
closely resembles that of those from the other localities that a special 


HISTORY OF GRITS AND SANDSTONES. J1 


description of it is unnecessary. The felspar, of which a large pro- 
portion is triclinic, is present in large quantities; and, as in the 
case of the other specimens examined, the quartz contains but few 
fluid-cavities. It will be observed that in this respect the quartz of 
these grits materially differs from that of the Cornish granites, in 
which fluid-cavities with bubbles are abundant. 

Carboniferous.—A. fine-grained yellowish-white sandstone from 
Shalk Beck, Cumberland, belonging to the Yoredale series, much 
spotted by stains of hydrated ferric oxide, and rendered somewhat 
mealy a) the presence of kaolin, contains but few fragments of more 
than ;4, ich in diameter. The quartz is in angular pieces, colour- 
less and transparent, and, to a large extent, free from fluid-cavities, 
which, when present, are, for the most part, full, and consequently 
without bubbles. In addition to quartz, with a little felspar and 
kaolin, the only recognizable mineral is white mica. For the 
analysis, see p. 21. 

A sandstone from Brigham, Cumberland, belonging to the Mill- 
stone-Grit series, was examined both microscopically and chemically, 
and is essentially composed of a of quartz with a little fel- 
spar, the grains being usually about +45 inch in diameter, united by 
a siliceous cement, which j is sometimes a little cloudy. Between the 
- constituent fraements of this rock there are sometimes minute 
erystals of a mineral which may perhaps be epidote; and the quartz, 
which is colourless and transparent, encloses a few needles of tour- 
maline, besides containing occasional fluid-cavities, but few of which 
contain bubbles. For the chemical composition of this sandstone, 
see page 21. 

AtSpinkwell quarry, near Bradford, a foliated siliceous sandstone, 
which can be raised in the form of very large slabs, is worked in the 
Lower Coal-measures, and is much used in the construction of chemical 
apparatus on a manufacturing scale. This sandstone is mainly com- 
posed of fragmentary quartz and felspar, of the latter of which a 
portion is triclinic, united by the usual siliceous cement. The 
quartz is colourless and transparent, and contains but few fluid- 
cavities, although it sometimes encloses needles of schorl. 

In addition to quartz and felspar, this rock contains kaolin, with a 
few minute garnets, and flakes of dark-brown and colourless mica, 
which are more abundant along certain lines of foliation than else- 
where. A few minute crystalline scales of micaceous oxide of iron 
were observed in the eas of this sandstone. The component 
fragments rarely exceed 51, inch in diameter. For analysis of this 
rock, see page 21. 

The fine-grained yellowish-grey Coal-measure sandstone of Stony- 
hough, Workington, Cumberland, is, to a large extent, oe of 
minute crystals. or crystalline ageregations oi quartz of about 51, 
inch in diameter, somewhat loosely united by a siliceous cement, 
often much stained by hydrated ferric oxide. 

A considerable number of sandstones belonging to the Carboni- 
ferous period are chiefly composed of quartz crystals, which have 
evidently crystallized in situ, since they exhibit the freshness of out- 
line peculiar to crystals which have not been subjected to the slightest 


1s J. A. PHILLIPS ON THE CONSTITUTION AND 


amount of abrasion subsequently to their formation, not a point being 
broken or an angle removed. Sometimes the original grains of 
quartz have, by the subsequent deposit of silica upon their surfaces, 
become converted into complete double-terminated crystals; but 
the forms are frequently less simple, and the faces bounding the ex- 
terior cannot all be referred tothe same crystal. That this crystal- 
lization is produced by a deposit of silica around the original grains 
of quartz, subsequently to their having become members of an accu- 
mulation of sand, was first noticed in British rocks by Professor 
Bonney*, and has been subsequently demonstrated by Dr. Sorby 7. 
It is almost equally certain, as will be subsequently shown, that a 
portion, at least, of the silica so deposited has been derived from the 
decomposition of felspar. 

Among highly crystalline Carboniferous sandstones may be men- 
tioned one belonging to the Mountain-Limestone series, which occurs 
at Yeathouse, in Cumberland, that of Parkhead, in the same county, 
and that of Augill, near Brough, Westmoreland; the two last of 
Yoredale age. Another sandstone in the Lower Coal-measures, 
worked at Barngill quarry, in the county of Cumberland, is also 
crystalline. 

Permian.—The St. Bees Sandstone, at Rheda, Cumberland, is a 
fine-grained reddish-brown rock, composed of a mixture of angular - 
fragments and minute crystals of quartz with a little felspar, the 
whole being united by a cement rendered, to some extent, opaque 
by ferric hydrate. The grains and crystals of quartz have usually a 
diameter of about 54, inch, and contain but few fluid-cavities. The 
colour of this sandstone, like that of the majority of similar rocks, is 
caused by a coating of hydrated oxide of iron over the surfaces of 
the grains and crystals of which it is composed, but which is readily 
removed by digestion in acids. Felspar is present in notable quan- 
tity, and is often considerably altered; no triclinic species was 
observed. A small amount of colourless mica is present in the form 
of water-worn flakes, together with a few imperfect crystals of 
schorl. 

Numerous other crystalline sandstones of Permian age might be 
cited; but those of Penrith, which have been described by Dr. Sorby, 
are probably the most interesting and remarkable examples. 

Triassic.—Among the Bunter sandstones of Lancashire and Che- 
shire are certain reddish-brown friable beds, possessing but little 
cohesion, and of which the constituent grains are all so completely 
rounded, that the disintegrated sand flows between the fingers as 
readily as shot. Deposits of such sandstones, which are distin- 
guished by the name of “ Millet-seed beds,” occur plentifully in 
the Lower Mottled series, and occasionally among the Upper Mottled 
sandstones, as well as in the Frodsham beds of the Keuper. 

A specimen of millet-seed sandstone from the Lower Bunter, 
obtained in the form of a core, at a depth of 1039 feet, from the 


* Quart. Journ. Geol. Soc. vol. xxxy. p. 666. 
+ Address delivered at Anniversary Meeting of the Geological Society of 
London, 20th February, 1880, p. 36. 


HISTORY OF GRITS AND SANDSTONES. 13 


Bootle bore-hole of the Liverpool water-works, was, with a large 
number of others from that district, kindly furnished to me by Mr. 
Charles HE. De Rance, of the Geological Survey, but was found to 
be too friable to admit of the preparation from it of thin sections. 
On examination by reflected ght, however, it was found to be so 
entirely made up of rounded grains, varying in diameter from +, 
to st, of an inch, that I did not hesitate to suggest to Mr. De 
Rance the probability of its origin being due to blown sands united 
by a ferruginous cement. 

These grains, of which the majority are quartz, are so rounded as 
to represent miniature pebbles, while a few, consisting of partially 
decomposed felspar, are often corroded into deep cavities on one or 
more of their sides. 

The granules of quartz, as well as those of felspar, have been 
covered by a thin coating of hydrated ferric oxide; while on the 
surfaces of the former a beautiful growth of crystals of trans- 
parent quartz has frequently taken place. 

These crystals do not often exceed +4, inch in length; but they are 
sometimes very perfect, with sharply defined angles, and frequently 
exhibit both plagihedral modifications and horizontal striation. A 
few crystals of pyrites and of calcite have also been formed on the 
surfaces of the rounded quartz-grains. 

Figs. 1, 2, 3, and 5, Pl. Il., drawn by Mr. F. Rutley from 
specimens which I selected for that purpose, and magnified one 
hundred diameters, represent crystals of quartz attached to rounded 
grains of the same mineral forming the basis of this sandstone. 
Fig. 4, magnified to the same extent as the others, is a grain of 
quartz which exhibits a depression at its point of contact with 
another similar body. Specimens presenting this appearance, which 
are not very numerous, may sometimes be the result of one grain of 
quartz having been forced or ground into the substance of another. 
A careful examination, however, of such depressions leads to the 
conclusion that in some cases when this pitting of a grain is ob- 
served a deposit of silica may have taken place upon all parts of its 
_ surface, excepting where it has been protected by contact with 
adjoining grains. 

Angular cores of a siliceous material which have been deposited 
in cavities at the point of junction of several grains are sometimes 
detached when the rock is carefully disintegrated by friction with 
a hard brush. When these adhere to one only of the adjacent 
grains, haying separated from the others, they obviously might give 
rise to depressions of the kind referred to. 

If, after treating this sandstone by hydrochloric acid, the residue 
be examined under the microscope, the presence of these siliceous 
bodies becomes at once apparent. They are sometimes slightly 
coloured by ferric oxide, and do not always exhibit colours when 
seen in polarized light; in other cases they afford evidence of im- 
perfect crystalline structure, and are occasionally colourless and 
transparent. 


Fig. 6, Pl. II., is a grain of felspar which has become so corroded 


14 J. A. PHILLIPS ON THE CONSTITUTION AND 


that two distinct cavities have been produced in its substance. A 
erystal of iron pyrites attached to a rounded grain of quartz is seen 
in fig. 7, which, as well as the preceding figure, is represented as 
magnified to the same extent as the other illustrations contained in 
the Plate. 

Prolonged digestion in hydrochloric acid removes the oxide of 
iron, leaving the surfaces of the rounded quartz granules clean 
and colourless. The minute crystals of quartz which have been 
formed upon them, however, adhere firmly to the rounded grains, 
after this treatment, and no stain of ferruginous matter can be 
observed between their point of attachment and the grain of sand 
on which they have been formed. It would therefore appear 
that, although the sand had been covered generally by ferric hy- 
drates previous to the growth of quartz crystals, these have never- 
theless originated at those points only where a chemically clean 
surface of the quartzose nucleus was exposed. An analysis of this 
sandstone is given, p. 21. 

Sandstones chiefly composed of rounded weather-worn grains occur 
in the Lower Mottled series, at a depth of 80 feet from the surface 
at Stock’s Well, belonging to the Widnes water-works, and at 
Scott’s bore-hole near St. Helens, as well as at a depth of 260 feet 
in the Winwick boring of the Warrington water-works. 

Beds of loose, rounded sand of the age of the Lower Mottled 
Sandstone are known to occur at Chapel Bridge, Prescot, and in a 
boring a little east of Newton Bridge, near Warrington. 

Sandstones of this character are met with north of Eccleston hill, 
and asimilar bed belonging to the Upper Mottled group comes to the 
surface in the yard of the Bridgewater Foundry at Runcorn. By 
no means, however, do all the sandstones of Lancashire and Cheshire 
which belong to this geological age exhibit characteristics suggestive 
of their formation from eolian sands. At Wirral, in Cheshire, as 
well as sometimes in the neighbourhood of Liverpool, the Pebble- 
beds of the Bunter are represented by a brownish-yellow sandstone 
containing numerous pebbles, which is much employed for building- 
purposes. 

In many of these beds the quartz is almost entirely in the form of 
minute crystals, or crystalline aggregations, often united by a ferru- 
ginous cement, which has manifestly been introduced subsequently 
to the covering of the original grains with crystalline quartz. 
An excelient example of a non-ferruginous crystallized sandstone 
belonging to the Upper Mottled group occurs at Town Green, near 
Ormskirk. This rock is mainly composed of crystals of transparent 
quartz, of which the edges and angles are beautifully perfect. It 
is of a far too friable nature to allow of the preparation of thin 
sections ; but it appears to have little or no cementing material, and 
to be, to a large extent, merely felted together by the intergrowth 
of its constituent crystals. It will be needless to remark that the 
grains of sandstones of Bunter age are not always oleae rounded or 
enveloped i in crystals of quartz. 

Among the Keuper division of the Triassic candauinen which are 


HISTORY OF GRITS AND SANDSTONES. 15 


mainly composed of well-rounded siliceous grains, may be cited a 
grey, friable, fine-grained rock from the base of this formation, 
which is exposed in the railway-cutting at the Runcorn station, 
and a dark-red sandstone of still finer texture, belonging to the 
Frodsham beds, at no great distance from the same locality. The 
Lower Keuper cupreous sandstones of Alderley Edge, Cheshire, are 
frequently made up of quartz crystals on which still more minute 
crystals of vanadinite may sometimes be distinguished by the aid 
of a lens. 

In many localities the quartz pebbles occurring in crystalline 
arenaceous rocks have their surfaces, and more particularly their 
upper surfaces, covered by minute crystals of that mineral. ‘This 
may be observed in the case of pebbles found in Lower Keuper 
sandstone in a quarry near Litherland. 

At Dymoke, Worcestershire, there is a Lower Keuper sandstone 
which is sufficiently coherent to admit of the preparation of thin 
sections. This is a fine-grained, quartzose, distinctly DALES 
rock, of which the component grains sr 
in diameter. These to a large extent consist of quartz, Jeena: 
enclosing hair-like crystals of rutile, and occasionally fluid-cavities, 
in some of which bubbles were observed. A certain amount of 
felspar, a portion of which is triclinic, is present in this rock. A 
few flakes of colourless mica, and a little of the- fibrous mineral 
which has been referred to as often occurring in the cement of 
certain sandstones, were also observed. The cementing material, 
which contains a little kaolin, encloses a few minute garnets, and 
is frequently stained by hydrated ferric oxide. 

The Waterstone beds belonging to this series enclose numerous 
angular fragments of dark-coloured slaty rock, some of which are as 
much as 3 inch in diameter. The quartz grains, many of which are 
; inch in diameter, are usually much ‘younded, and not unfre- 
quently enclose fluid-cavities. In addition to quartz and the 
cementing material felspar is present, as is also, in small quantities, 
another mineral of a light yellowish-green colour, which I have 
been unable to identify, but which occasionally forms part of the 
cement. 

A fine-grained sandstone of Upper Keuper age, which occurs at 
High House, Warwickshire, is to a large extent composed of quartz 
erystals, while a bed of loose sand, found 25 fect below the surface 
at Frodsham, above the Keuper Marl, is, on the contrary, entirely 
made up of much-rounded grains. 

The rounded quartz grains of Triassic sandstones, when examined 
in a suitable medium, after the removal of their external ferruginous 
coating, are found to be colourless and often transparent. Grains con- 
teens ‘fluid-cavities are comparatively rare, but they are apparently 
more numerous in the Keuper sandstones than in rocks of Bunter 
age. <A few crystals of schorl occasionally present themselves in the 
quartz of these rocks, which is not unfrequently rendered turbid 
by the enclosure of what is probably a little ferruginous clay. It 
is, however, probable that the grains containing “fluid-cavities in 


16 J. A. PHILLIPS ON THE CONSTITUTION AND 


which crystals of schorl are also found owe their origin to a different 
source from. that whence the supply of quartz which does not contain 
such cavities was derived. 

Jurassic.—A fine-grained Upper Lias sand, of a greenish colour, 
from Seizincote, Stow-on-the-Wold, Gloucestershire, effervesces 
when treated with hydrochloric acid, leaving grains of transparent 
quartz, which are generally angular. In some instances, however, 
their more acute angles appear to be slightly rounded, although 
the mean diameter of the fragments is only 5,5 inch. Besides 
quartz containing occasional fluid-cavities, usually without bubbles, 
there are present a few pieces of somewhat.doubtful felspar, together 
with numerous fragmentary crystals of schorl and garnet. 

The only coherent arenaceous rock of this age which I have had 
an opportunity of examining is that quarried at Egton, near Whitby, 
under the name of “‘ Moor Grit,” which is locally much employed 
for road-metal. It is white and fine-grained, being often so 
compact as to be entitled rather to the name of quartzite than to 
that of grit. Its geological horizon is above the Grey Limestone in 
the estuarine series of the Lower Oolite in North-east Yorkshire. 

Under the microscope this rock is seen to be almost entirely 
composed of CSIDENEDE, colourless quartz, of which the largest 
pieces are about -; inch in diameter, and of which the ineleal are 
usually more or. less removed. Around and between these grains 
a deposit of transparent crystalline quartz has taken place, thus 
forming a cementing medium. A few small garnets are present, 
but no “fluid- cavities ath bubbles were observed, although some of 
the quartz encloses minute crystals of a yellowish mineral which 
I have been unable vo identify ; these are exceedingly minute, 
often not exceeding ;>4y)5 mch in length. Many of the smaller 
grains in this rock exhibit, when examined in polarized light, that 
complex structure so frequently observed in the quartz of clay-slates 
and other somewhat similar rocks. 

A sand resulting from a disintegrated Portland Stone at Fonthill 
Giffard, Wiltshire, is largely composed of ovoid grains of calcite. 
After being attacked by hydrochloric acid, a rounded quartzose sand, 
amounting to about one quarter of the total bulk of the mixture, 
remains behind. This sand, of which the grains vary from =), to 545 
inch in diameter, contains but few fluid-cavities, and these, as a rule, 
are without bubbles. The fragments of quartz are associated with, 
and not enclosed by, the ovoid grains of calcic carbonate. 

Cretaceous.—The Tilgate Sandstone, Ashdown Sands, from Mewaee 
Park, Sussex, 1s composed of slightly rounded grains of colourless 
transparent quartz, united by a cement consisting partly of calcic 
carbonate and partly of flint. The quartz is almost entirely free 
from fluid-cavities, but encloses a few hair-like crystals of a mineral 
which is probably rutile. If felspar be present it has become too 
extensively altered to admit of identification. i 

Sections have been examined of the chert known as Sevenoaks 
Stone, as well as of several others of Lower Greensand age. They 
all contain numerous fossils, particularly sponge-spicules, and in 


HISTORY OF GRITS AND SANDSTONES. 17; 


some cases consist, to a large extent, of amorphous silica; in others 
the rock becomes crystalline and distinctly chalcedonic. The flinty 
varieties often contain crystals of calcite, which cluster around an 
included fossil as a nucleus. All the specimens examined contain 
glauconite, and occasionally grains of ordinary quartz, some of which 
are much rounded, while others are angular. 

The “‘ Carstone” of Hunstanton, near King’s Lynn, Norfolk, and 
of various other localities, is a friable ferruginous sandstone belonging 
to the upper portion of the Lower Greensand formation, and occurring 
in beds of which the relations have not, as yet, been accurately 
determined. A series of specimens from this locality was kindly 
furnished me by Mr. 8. B. J. Skertchly, of the Geological Survey, 
who is at present occupied in working out the geology of the district. 
With the exception of certain variations in colour, these beds so 
closely resemble one another in their general characteristics that a 
description of one of them will suffice for the present purpose. 

A specimen of Carstone from immediately below the Red Chalk 
exposed in the cliff at Hunstanton was found to be mainly composed 
of a mixture of somewhat rounded grains of quartz, with small 
pebble-like granules of dark-brown iron-ore. The individual grains 
of these minerals vary in diameter from +4, inch to the most minute 
sand, although small pebbles of larger size than the highest limit 
quoted are not of unfrequent occurrence. The quartz contains 
schorl and rutile, together with a few fluid-cavities, of which the 
majority are without bubbles. In addition to ordinary quartz 
grains, this rock, when carefully disintegrated, exhibits numerous 
examples of the angular bodies resulting from the breaking-up of a 
siliceous deposit formed between the grains of the original sand, 
which have been noticed (p. 13) in connexion with millet-seed 
sandstones of Bunter age. 

By the prolonged action of hydrochloric acid the quartz of this 
rock is rendered colourless, while the globules of ferric hydrate are 
dissolved, with the exception of a siliceous skeleton which preserves 
the exact form of the original grains. These bodies do not usually 
exhibit colours when mounted in balsam and examined by polarized 
light; but in a few cases the presence of a dark cross indicates a 
pisolitic structure in the siliceous residue*. In addition to the 
foregoing, this rock contains a few minute scales of mica and a very 
small quantity of felspar. The majority of the grains of quartz 
have their angles distinctly abraded; in some instances they 
have been completely removed, and a pebble-like form has been the 
result. 

A specimen of Carstone obtained from a bed directly beneath the 
Red Chalk, afforded on analysis the following results :— 


* Both Dr. Percy and Professor Judd have described siliceous skeletons which 
occur in the pisolitic grains of Northamptonshire iron-ore:—Metallurgy, Iron 
and Steel, pp. 225, 226; Memoirs of the Geological Survey, Geology of Rutland 
&e. p. 119. 


@.5.G.S. No. 145. C 


18 J. A. PHILLIPS ON THE CONSTITUTION AND 


Ao [fab PeTRONINE TLIO 54 ooo 55.0 v 3°85 
Water (eee CEG dire a ok 6:56 
SiGe hy he SeGec Peat tice eee engin A981 
Phosphoric anhydride yee eee 0-42 
PAWL UUAAATN Gute we loy nee Rete es eRe ee ene eC Salen 

*Reric Oxide y tea ena eee 29°17 
errous OxIde « Yds sie: 0-35 
APART) A eee) Seat a ee aaa i ocean a 2°43 
Macon esiae Ac eeciee cea ee Reet ee 0°95 
POEASSA EL. Sei ie Reale 0°48 
SOdariicls eae Re Ray erp acters 0-84 

100-03 


Another variety of this rock from the same locality, but darker 
in colour, was found to contain 37 per cent. of ferric oxide and 4o 
per cent. of silica; the amount of phosphoric anhydride was nearly 
the same as in the first specimen analyzed. 

An examination of the spherules of various pisolitic iron-ores 
shows that they exhibit all the characteristics of the globular 
ferruginous grains found in these sandstones; and it may therefore 
be inferred that they have had a similar origin. A pisolitic iron- 
ore of Middle Neocomian age, which occurs at Market Rasen in 
Lincolnshire, consists to a large extent of spherules very closely 
resembling the ferruginous grains in the sandstones at Hunstanton. 

Tertiary.—Hertfordshire Puddingstone, Lower Kocene, is a conglo- 
merate of flint pebbles united by a concrete consisting of fragments 
of transparent quartz and greyish flint held together by a flinty 
cement. In this concrete the quartz is considerably in excess of 
the flint, and sometimes contains fiuid-cavities. Its fragments are 
all angular, and vary in diameter from =, to 54, inch. 

A specimen of sand from Hordwell, Hampshire, equivalent in age 
to the Headon beds, contains no recognizable felspar. All the quartz 
down to a diameter of +, inch is completely rounded; and even the 
smallest particles have had their angles entirely removed. Fluid- 
cavities with bubbles are abundant in some of the quartz consti- 
tuting this sand. 

Sand from the Marine beds, near the top of the Hempstead series, 
Isle of Wight, was, after treatment with hydrochloric acid, found to 
be composed chiefly of grains of quartz, of which about three 
fourths had a diameter of less than 4, inch. These, down to the 
finest particles, are much rounded, although still roughly retaining 
the form of the original fragments. 

The fine-grained brilliantly coloured sands at Alum Bay, Isle of 
Wight, of Upper Eocene age, usually classified as Lower Bagshot, 
have not, as yet, been definitely identified with the beds of the London 
Basin. By digestion in hydrochloric acid the quartz becomes colour- 
less; and, although not completely rounded, the angles even of the 
smallest fragments have generally been modified by attrition. Fluid- 
cavities are not plentiful, and when present seldom enclose bubbles. 


HISTORY OF GRITS AND SANDSTONES. 19 


Needle-like crystals of schorl are sometimes enciosed in this 
quartz. 

The most considerable bed of sand at Bovey Heathfield, Devon- 
shire, no. 27 of Mr. Pengelly* (Miocene?), consists largely of 
quartzose fragments, nearly all of which are sharply angular, trans- 
parent, and colourless. ‘They contain fluid-cavities with bubbles ; 
but the latter appear to be less numerous than in the quartz of some 
Cornish granites. Schorl is present in considerable quantity, both 
as detached crystals and as portions of crystals, also as needles 
penetrating quartz. 

Post-Tertiary.—Sand washed from the Lower Boulder-clay at Holy- 
well, Flintshire, is largely composed of small quartz pebbles, rounded 
grains of various felspathic and other rocks, and numerous fragments 
of millet-seed sandstone. A few unworn quartz crystals resulting 
from the disintegration of crystalline sandstones, and some angular 
grains of quartz, were also observed. Even the smallest particles of 
this sand are often rounded. 

The larger grains of a sand of Middle Glacial age which occurs in 
this locality are either rounded grains of quartz or of some other 
rock, or small pebbles of millet-seed sandstone. Those of medium 
size are millet-seed quartz grains, mixed with a few unworn crystals 
and angular pieces. of the same mineral. 

A specimen of Middle Glacial sand from Bagilt in the same 
county differs in no respect from the foregoing, excepting that 
erystals of quartz derived from crystalline sandstones are rather 
more numerous, angular fragments are less rare, and broken millet- 
seed grains are of more frequent occurrence. 

The Middle Glacial drift at Colwyn Bay is mainly composed of 
small pebbles of various rocks, principally of quartz, with a few 
unworn crystals of the same mineral, resulting from the disintegra- 
tion of sandstones. In this drift the smallest fragments, although 
generally rounded, have not been converted into minute pebbles. 

At different times I have examined numerous specimens of recent 
water-borne sands. Among these, that on the sea-shore at Pentewan 
in Cornwall is, as described, p. 24, perfectly sharp and angular, as 
is the sand on the beach at Par, about six miles further east. Accord- 
ing to Dr. Sorby such is also the case with regard to the sands of 
the modern beach at Scarborough, and those of the river-terraces at 
Dunkeld. 

A large proportion of the quartz in the sands of the Thames 
valley is sharply angular, although mixed with rounded grains of 
the same size. The grains of the auriferous sands collected on the 
coast of Northern California are hkewise for the most part angular, 
although perfectly rounded ones are at the same time present. 

Among the blown or eolian sands which have been examined is 
one from the Great African Desert, and another from Arabia 
Petrea. The grains of these are, without exception, much worn; 
and there is no admixture of the angular fragments found in 


we 


* “The Lignites and Clays of Bovey Tracey,” Phil. Trans. 1862, vol. clii. 
p. 1019. 
c2 


20 J. A. PHILLIPS ON THE CONSTITUTION AND 


all subaqueous deposits; the majority are in the state of minute 
well-rounded pebbles. As in the case of water-borne sands, the 
effects of attrition are more conspicuous in the larger fragments 
than in the smaller ones; but even the most minute particles are in 
these sands much rounded. Exceptionally the quartz encloses 
fragments of a felspathic material; and fluid-cavities with bubbles 
are not entirely absent in African specimens. 

A bed of sandstone, said to occur in a salt-producing district 
sixty miles south-east of Tebessa, of which some years ago I brought 
a specimen from Tunis to London, is entirely made up of rounded 
orains. In that respect this rock resembles the millet-seed sand- 
stones of Lancashire and Cheshire; but it is unlike them, inasmuch 
as no crystals of quartz, or of any other mineral, have been deposited 
upon the surfaces of the rounded granules. 

Modern blown sands, of which we have numerous examples in 
this country, differ from desert sands and from those of certain 
sandstones only in being usually somewhat less completely rounded. 
Among the sands of this description which have been examined are 
specimens from the dunes at Rhyl, Flintshire, Colwyn Bay, Den- 
bighshire, Lytham, Lancashire, and from Perranzabuloe and Lelant, 
in Cornwall. Speaking generally, the sands from the northern loca- 
lities have been more completely rounded than those from Cornwall, 
and consist of a mixture of worn quartz and various slaty and 
other rocks, with a little felspar and a few fragments of shells. 
Fluid-cavities with bubbles are rare in the quartz of these sands. 
In addition to rounded grains of various slaty and other rocks, 
quartz, felspar, and fragments of shells are present; among these 
quartz largely predominates. The Cornish sands contain a few 
partially rounded prisms of tourmaline. 

Chemical Composition of Sandstones §c.—A microscopic examina- 
tion of a large number of sections of grits and sandstones haying 
led to the conclusion that many of the published analyses of such 
rocks must be of a very imperfect character, five different specimens 
were selected for analysis. 

In making these analyses I have received the valuable assistance of 
Mr. E. W. Voelcker, A.R.S.M.; and in each case fusion with alkaline 
carbonates was adopted. The estimation of alkalies was made after 
an attack by hydrofluoric acid, and was checked by a fusion with 
carbonate of calcium and chloride of ammonium. 


HISTORY OF GRITS AND SANDSTONES. 


PAA 


Analyses*. 
I. 1 III. TV. V. 
Winter { hygrometric ...... 125 150 050 “400 150 
Combined) J/.05.5-.+- ‘935 “700 | 1:290 "850 "300 
STLE ES) coo Ogee See 80-600 | 75°750 | 87-400 | 85:550 | 87-150 
ALOT eee ee eee 9200 | 8227 | 3997 | 7570| 3-948 
Carbonic anhydride ......... IE OZON ers cce IES7Oa | iecsaccc: 1-200 
Phosphoric anhydride ...... ‘076 0-15 | trace. 070 | trace. 
Sulphuric anhydride ...... trace. 171 060 | trace. ‘094 
IEEEIGIOXICO) -o 20... 20. -c25e0e- trace te LO ile rsa leases ses 1-352 
IERCOUSIORIGE ........5..002- 2370 1352 1366 1-915 
Herre persulphide (ReS,) .|....2..5. | ..-..---- 800 ‘73 °203 
Manganous oxide ............ 232 | trace. PALS) A paceoagee 
IDEID® 48-0 /0dsseeeee ee 1-330 532 1-932 588 | 2°681 
WEEN GEIC)) ieee en oe 1-285 "360 684 612 1-080 
I°QUSINEC) SASe nae ROneeee pe eeeeeee 1-647 1-059 Weal "915 1:273 
SG)” cocccseeCsn senate USB | es} "332 | 1-113 "840 
100-197 | 100-120 | 100-307 | 100:336 | 100-271 
Specific gravity ......... 2-689 | 2-464 2710 | 2-531 2-660 


J. Grit, Cambrian: Barmouth, North Wales. 

II. Sandstone, Carboniferoust: Yoredale Series, Shalk Beck, Cumberland. 
Iii. Sandstone, Carboniferous: Millstone-Grit Series, Brigham, Cumberland. 
IV. Sandstone, Carboniferous: Lower Coal-measures, Spinkwell quarry, 

Bradford. 
VY. Sandstone, Triassic: Bunter, Bootle Well, Liverpool. 


Examination of Water- borne Sands. 


With the view of to some extent studying the action of running 
water upon the mineral fragments which it transports, a microsco- 
pical examination was made of the sands of the St. Austell river, in 
Cornwall. This stream, which during the summer months is a mere 
rivulet of moderate size, sometimes in winter becomes a considerable 
torrent. Its eastern arm arises at a distance of two miles and 
three quarters from the town, and at a height of 470 feet above the 
foot of the weir at the “Old Bridge.” Its western arm, which meets, 
the other a little north-west of the town, takes its rise in a small 
valley only a mile from the point of origin of the more easterly 
branch. From the bridge the distance to the sea at Pentewan is 
four miles, while the total fall is only 114 feet. 

This stream formerly carried with it into the bay vast quantities 
of the granitic sand which is separated by washing from china-clay 
at the different clay-works in the district. As, however, catch-pits 
have of late years been employed for the purpose of retaining it at 


* Since the above were completed, my attention has been directed to some 
analyses published in U.S. Geol. Survey, XLth Parallel, vol. ii. pp. 35 & 246, 
which agree very closely with those here given. 

+ A white fine-grained sandstone, much spotted with brown. - 


22 J. A. PHILLIPS ON THE CONSTITUTION AND 


the several works, the quantities thus transported are now very 
small. 

On the eastern branch a certain amount of granitic sand escapes 
into the stream almost immediately at its source; this is repeated 
at short intervals for a distance of a mile and a half, and finally 
ceases a mile and a quarter above the Old Bridge, at a height of 
about 200 feet above the weir-foot. 

The first introduction of sand into the western branch takes place 
nearly a mile below its source, and is discontinued half a mile 
further down its course, but at a somewhat lower level than in the 
case of the eastern fork. 

It follows that, before arriving at the bridge, each grain of sand 
must have travelled over a distance of at least a mile and a quarter, 
with a fall of above 150 feet, while a portion of it has been trans- 
ported two miles and three quarters through a channel thickly 
strewn with granite boulders, and having a fall in that distance of 
470 feet. 

Before arriving at the sea, therefore, the whole of the sand must 
have travelled at least five miles and a quarter, with a minimum 
fall of about 270 feet, while a portion of it will have been transported 
a distance of six miles and three quarters over a total declivity of 
584 feet. 

The first samples of material were collected from the bed of the 
river a little below the bridge, and a mile and a half from the point 
at which the last granitic sands are discharged into the stream. 

For the purpose of facilitating a microscopical examination of 
these sands, they were divided into four different parcels by a series 
of sieves, the first sieve allowing to pass through it all fragments 
less tesa =|, inch in diameter, the second those haying a less diameter 
than 4 inch, and the third all particles having a smaller diameter 


fo} Wi 
than —~ inch. 


The lar rgest fragments, retained upon the coarsest sieve were about 
7 inch in inmeten eraduating to a diameter of =, inch. 

This sand consists of a mixture of quartz, felspar, schorl, and 
mica, in which the last-named mineral is present in smaller pro- 
portion than any of the other minerals. When examined by re- 
flected light, and magnified 20 diameters, the edges and points of 
the different fragments of quartz and schorl are found to be sharp 
and unrounded ; the only exception being in the case of certain 
grains of quartz, which Dr. Sorby suggests may have been corroded 
by the action of alkaline waters, but which may have perhaps never 
possessed other than rounded outlines. 

The angles and edges of the felspar and mica are, on the contrary, 
distinctly ‘Tounded ; and although this might have been anticipated 
as regards the mica, it is at first’ sight not so easily understood in 
the case of felspar, whose density and hardness differ but slightly 
from those of quartz. It must, however, be remembered that the 
grains of felspar had become externally kaolinized while still form- 
ing an integral portion of the decomposed granite, and that, on the 


HISTORY OF GRITS AND SANDSTONES. 23 


removal of this coating of china-clay by washing, a rounded central 
nucleus will remain. 

Sand which had passed through the ;5-inch apertures of the first 
sieve, but which was retained by the ,-inch openings of the second, 
has a similar composition to that of coarser grain. The quartz and 
schorl are both angular; the felspar is more rounded than the quartz; 
and the mica, which is more plentiful than in the coarser sand, is 
much worn on the edges. 

The sand, which after passing through the j,-inch sieve was 
retained on the ,1,-inch sieve, consists of a mixture of angular 
quartz, unworn crystals of tourmaline, grains of felspar (some of 
which are rounded) and flakes ‘of mica (which are much worn on 
the edges). 

An examination of the material which passed the sieve having 
apertures ;}, inch in diameter, shows that its grains are entirely 
unwaterworn. The proportion of mica is much larger than in the 
coarser sands, and there is less felspar; but most of the grains have 
had their angles removed. ‘The quartz is generally in the form of 
tabular flake-like fragments, while the schorl often oceurs as small 
acicular crystals. 

The quartz of these sands is frequently penetrated by needles of 
schorl; and when mounted in balsam it is seen to be full of fiuid- 
cavities containing bubbles—in this respect differing entirely from 
the quartz of the Cornish grits, as well as from that of the majority 
of sandstones. 

Specimens of the sandy deposit were taken from down the course 
of the river, at intervals of a mile apart, the last having been obtained 
at a point slightly above the sea-level at high water at Pentewan. 
In every case, however, they so exactly resembled those first taken 
from below the Old Bridge at St. Austell as not to require detailed 
description. The quartz and schorl are angular, the felspar is more 
or less rounded, and the larger flakes of mica are worn at the edges. 
With regard to the distribution of sand along the river-bed, it is 
needless to remark that the coarser fragments are found towards the 
centre of the stream, while the finer silt, with minute flakes of mica 
and quartz, accumulates in less rapidly moving currents near the 
banks. 

Having found that quartz grains below —4, inch diameter are not in 
the slightest degree rounded by a minimum transit of five miles 
and a quarter down the course of the stream, it was thought desirable 
to ascertain the effect of a prolonged action of the waves upon the 
sand lying on the sea-shore. 

It must be here remarked that since the first opening of china- 
elay works in this district, now about sixty years since, millions of 
tons of granitic sand have been carried into the sea by the streams 
into which it was discharged. The effect of this at Pentewan has 
been to silt up the harbour to a very serious extent ; while the whole 
of the sands upon the sea-beach bear evidence of having been 
derived from the same source. 

The point from which the specimens were taken for examination 


94. J. A. PHILLIPS ON THE CONSTITUTION AND 


is situated at a distance of half a mile west from the present mouth 
of the stream by which the sands were brought down. They were 
collected from the water’s edge at half-tide; and as the discharge of 
granitic sands into the river has been very small during the last ten 
or twelve years, and this point is considerably removed from its 
mouth, it is evident that a large proportion of the grains taken must 
have been subjected during many years to the wearing action of the - 
waves. 

An pao of this sand shows that quartz having a diameter 
between =} go and zy 5 inch is usually angular, although some of the 
larger pieces are “distinctly (but not considerably). rounded. The 
schorl, like the quartz, generally” presents sharp angles, although 
somewhat abraded grains ‘of this mineral are occasionally met with. 
Nearly all the felspar is rounded to a considerable extent, as is also 
the small Graal of mica which is present. 

Below =; inch in diameter the angularity of the fragments of 
quartz and schorl is perfect, with the exception of ‘occasional 
‘ corroded ” grains ; the felspar has, for the most part, rounded 
outlines; and mica is almost entirely absent. 

At page 32 of his Address to the Geological Society of London 
(1880) Dr. Sorby remarks :—“ Unfortunately I am not acquainted 
with sufficient facts to prove how long it would require to thoroughly 
wear down and round a grain 4, inch in diameter. It is evident it is 
avery different thing from the wearing of a pebble, and may require 
a longer period of wear than we might suspect, if we did not bear 
in mind that when buoyed up by water the friction of such small 
particles on the bottom must be always small.” Again, at page 34, 
he says :—‘‘ It appears to me sufficiently proved that a great amount 
of drifting and mechanical action must be segue to wear down 
angular fragments of quartz into rounded grains 7 )q inch i in diameter, 
ith i have taken as the standard for comparison.” 

Professor Daubrée states that the diameter ot erains capable of 
floating in slightly agitated water is about 4, millimetre, or, say, 
sti inch, and remarks that all smaller grains must of necessiby 
remain angular *. He subsequently says ‘That a current or wave 
capable of carrying off in suspension particles of that diameter, with- 
out in any way affecting their form, would cause larger fragments 
of the same mineral to so rub one against another as gradually to 
produce rounded sand. According to an experiment quoted by this 
STN, a sand of which the grains have a diameter of ;°, millimetre, 
say =, inch, to which a movement of one metre pe Tagan 1s Im- 
ee becomes rounded, with a loss equal to 754 pp of its weight 
per kilometre traversed. 

This experiment appears to indicate that a grain of quartz zy inch 
in diameter requires, before becoming completely rounded and 
assuming the form of a miniature pebble, an amount of abrasion 
equal to ‘that which would result from having travelled a distance 
of three thousand miles. In arriving at this conclusion the fact 
must not be lost sight of that, after the first rounding of the 

* Géologie Expérimentale, p. 256. 


HISTORY OF GRITS AND SANDSTONES. 25 


angles and edges, the operation will gradually become slower as the 
surfaces become more worn and the weight of the grain decreases. 

That angular fragments of quartz having a diameter of less than 
=i; inch remain unrounded by the continuous action of breakers after 
many years’ exposure, is evident from an examination of the sands 
at Pentewan. It has been shown by other evidence, as well as by 
the recent experiments of Professor Daubrée, that the rounding-down 
of such sands by the action of running water must be an exceed- 
ingly slow operation, and one requiring a somewhat active current 
with an amount of friction equivalent to transport over enormous 
distances. Grains of quartz of similar dimensions are, in blown 
sands, completely rounded. 


Summary and General Conclusions. 


The Cambrian grits of Barmouth contain quartz and felspar, both 
in the form of angular fragments and also as rounded pebbles. The 
materials presenting these different forms have probably been de- 
rived from two distinct sources; while the large size and complete 
sharpness of the angles of mauy of the irregular grains appear to 
indicate that they cannot have been transported from any consider- 
able distance, and that the felspar cannot have been derived from 
kaolinized granite. 

All the arenaceous rocks of Silurian age which have been ex- 
amined contain a small proportion of felspar, the grains of the 
various constituent minerals being in some cases angular, and in 
others rounded. 

Many of the rocks belonging to this period are composed of a 
mixture of grains of both forms. Among rocks mainly composed of 
rounded grains are the Stiper Stones of Shropshire and the Lower 
Lickey Quartzites of Westmoreland. Some of the grits from the 
neighbourhood of Aberystwith enclose fragments of a volcanic rock 
of doleritic character. 

The grits of Cornwall, which are of at least Devonian age, include 
flakes of soft slaty rocks, the edges of which are perfectly sharp, 
together with angular fragments of the well-known “ greenstones ”’ 
and ‘‘ dunstones” of that county. 

A large number of the Carboniferous, Permian, and Triassic 
sandstones are composed almost entirely of quartz crystals, which 
have undoubtedly been produced in sztu, as they not only penetrate 
and interpenetrate one another, but also exhibit the most perfect 
sharpness and freshness of outline. As confirmatory of this hypo- 
thesis it may be mentioned that in a quarry at Foggen Tor, on 
Dartmoor, the felspar has in places become decomposed into soft 
kaolin, in which the liberated silica is imbedded in the form of 
ageregations of well-formed and transparent quartz crystals *. 
Unworn double-pointed crystals of quartz, likewise resulting from 
the decomposition of felspar, have. recently been found near St. 
Austell, Cornwall, in soft china-clay; one of these, more than three 


* Quart. Journ. Geol. Soc. vol. xxxvi. p. 9. 


26 J. A. PHILLIPS ON THE CONSTITUTION AND 


inches in length, is now in the collection of the Museum of Practical 
Geology. Sandstones of this description are not unfrequently with- 
out any kind of cementing matrix, being merely felted together by 
a matted intergrowth of their constituent crystals. 

Professor Daubrée entertains the opinion that crystalline sand- 
stones frequently owe their origin to chemical agencies resulting 
from an outpouring of igneous rock; but, although this may some- 
times haye been the case, many of the most completely crystalline 
British sandstones are situated at distances of many miles from any 
known rock belonging to this class. The same author maintains 
that the presence of anhydrous ferric oxide in sandstones affords 
evidence of their having been subjected to a high temperature*. It 
must, however, be remembered that the carnallite of Stassfurt, which 
has evidently never been highly heated, contains crystals of specular 
iron-ore. 

Numerous fine-grained sandstones, particularly among those of 
Triassic Age, are composed of quartz grains so completely rounded 
as, under the microscope, to resemble well-worn pebbles. These 
‘* millet-seed”’ sandstones are often coloured either red or brown 
by variously hydrated oxides of iron; and in some cases minute, 
perfectly formed, and beautifully transparent crystals of quartz have 
been developed upon their surfaces. 

On attacking the sand of such sandstones with hydrochloric acid, 
the oxide of iron is easily removed, put the crystals of quartz still 
remain firmly attached to the surface of the grains upon which they 
have grown. It would also appear that crystals have been formed 
upon those parts only of the grains which, having been free from a 
coating of oxide of iron and from every other extraneous material, 
have admitted of direct chemical contact between the silica of the 
rounded quartz and that of the subsequently formed crystals. 

In addition to silica in the form of perfect crystals of quartz, that 
substance has often been deposited in such a way as to fill cavities 
existing between the original grains of sand. 

As this silica has frequently been thrown down upon a thin deposit 
of ferric hydrate, it is generally detached by prolonged digestion 
in hydrochloric acid, by which the intervening ferruginous coating of 
the grains is ultimately removed. When such a deposit of silica 
adheres to only one of the adjoining grains it may give rise to a de- 
pression upon its surface of the kind represented in fig. 4, Pl. II. 

On examining a considerable number of modern sands, none of 
them, excepting such as had long been subjected to the wearing 
effects of wind action, were found to resemble those of the millet- 
seed sandstones in having all their grains reduced to a pebble-like 
form. Among these the grains of blown desert-sands most com- 
pletely resemble those of millct-seed sandstones. 


* Géologie Expérimentale, pp. 226-230. 

+ With regard to the Torridon Sandstones of the Central Highlands it has 
been observed by Professor Bonney that wherever “dirt” has been deposited 
upon the siliceous grains their agglutination has been prevented (Quart. Journ. 
Geol. Soe. vol. xxxvi. p. 106). 


HISTORY OF GRITS AND SANDSTONES. 27 


The above facts would appear to render it probable that the 
rounded grains of these sandstones may be of eolian origin, and 
that, during certain periods of Triassic time, desert areas with blown 
sands extensively prevailed in this country. 

Mr. De Rance has observed that the millet-seed beds are usually 
free from pebbles, shale-beds, pseudomorphs after common salt, and 
from all traces of life*—conditions which are characteristic of de- 
posits produced by wind-currents. 

The granules of brown iron-ore which are so plentiful in the 
**Carstones”’ of Hunstanton are pisolitic grains, and not fragments 
of that mineral rounded by attrition. 

An instructive example of the occurrence at the same time of 
rounded and angular grains is met with in the Interglacial sands of 
Flintshire, where seme of the pebbles are fragments of a millet- 
seed sandstone, while many of the smaller particles are grains 
detached from the same rock. . 


HXPLANATION OF PLATES L. & II. 


Puate I. 
Magnified 18 diameters. 
Fig, 1. Group of felspar crystals in Cambrian Grit. Polarized light: p. 7. 


2. Grit from Ladock, Cornwall, enclosing a fragment of a volcanic rock. 
Polarized light: p. 10. 


Prats IT. 
Magnified 100 diameters. 
Figs. 1, 2,3, & 5. Crystals of quartz deposited upon rounded grains of the same 
mineral in Bunter Sandstone: p. 18. 
4. Depression in a grain of quartz from the same sandstone. 
6. Corroded grain of felspar from the same. 
7. Rounded grain of quartz with attached crystal of iron pyrites. 


Discussion. 


The Presrpent expressed his sense of the value of Mr. Phillips’s 
communication. 

Dr. Sorpy expressed his agreement with the paper, to which he 
had listened with great interest, especially as the author had ap- 
proached the subject from a point of view somewhat different from 
his own. He was especially glad to find that his opinions were 
confirmed by the author, especially as to the crystals of quartz in 
certain sandstones. The observations as to the time required to wear 
down a grain of sand were especially valuable. He had found the 
drift sands of the Yorkshire coast almost all angular; but then those 
examined by Mr. Phillips were from another locality, which might 
explain the difference in their observations. He should only regard 
sand as eolian when a very large proportion of grains were rounded. 


* “Further Notes of Triassic Borings near Warrington,” read before the 
Manchester Geol. Soc. June 29th, 1880. 


28 ON THE CONSTITUTION AND HISTORY OF GRITS AND SANDSTONES. 


So far as his observation had gone, the sands of the dunes on our 
coasts were not much more rounded than other sands. 

Dr. Hicks said that the condition of the Barmouth Grit led him to 
think that the materials had not drifted from far. He had also 
observed that there was a large proportion of rounded grains in 
the Stiper Stones; were we to suppose these to be blown sands? 
Rounded grains were still more common in the quartzites of the 
N.W. of Scotland ; we must believe therefore that there was, still 
remaining at the time they were deposited, an adjoining great land- 
area or the materials from one not far away. 

Mr. Dr Rance said that the Keuper beds under the Marls were 
now divided into the Water-stones, soft current-bedded sandstones 
called Frodsham beds (which denoted entirely different physical 
conditions and contained the millet-seed grains), and then the Lower 
Keuper building-stone (Labyrimthodon-beds). Then came a line of 
erosion. In the Bunter series were the Upper Mottled Sandstones 
(with the millet-seed grains), then the Pebble Beds (which had a 
different kind of current-bedding from that of the Frodsham beds), 
then another line of erosion and the Lower Mottled Sandstone with 
millet-seed grains again. The bedding of the sandhills of Lanca- 
shire much resembled that of the Frodsham and other millet-seed 
beds in their high angle and rapid change. 

Mr. Ruttey said he had examined eruptive rocks rather than 
sedimentary, but could not but express his gratification at the 
agreement between two such observers as Dr. Sorby and Mr. 
Phillips. He called attention to the presence of felspar in many of 
the sandstones described, and suggested that it was possible for 
such sandstones to be changed into felstone. There was often 
much difficulty in distinguishing between the finer-grained igneous 
and sedimentary rocks. He also called attention to the develop- 
ment of microcrystalline structure in felspar crystals. 

Mr. Branrorp said that some years ago he had examined the 
Indian desert, and found the grains of sand well rounded. They 
were mostly of quartz, with a few of felspar and occasionally of 
hornblende. The strongest wind there blows from the west; the 
sands had come from the coast and the river Indus; and the sand 
in the bed of the river was also rounded. The blown sand ap- 
peared unstratified. 

Mr. Puitiips said that the grains of millet-seed sandstone were 
much more rounded than was usual in sea-sand. He thought the 
rounding of felspar was often due to disintegration by decomposition. 
With regard to Mr. Blanford’s remarks, he could only say that he 
believed the grains of all the desert-sands which had been yet 
examined had been found to be much rounded. 


INTERGLACIAL DEPOSITS OF W. CUMBERLAND AND N. LANCASHIRE, 29 


3. LNTERGLACIAL Deposits of West CumBertAnp and Norra Lanca- 
sHirE. By J. D. Kenpatt, Esq., C.H., F.G.8. (Read Novem- 
ber 17, 1880.) 
[Puate IIT.] 


CoNTENTS. 


1. Introduction. 
2. Observed Facts. 
3. Deductions. 


1. IyrrRopuction. 


Tue glacial deposits of these districts are capable of a threefold 
division, as below :— 


1. Boulder-clay (Upper). 
2. Sand, Gravel, and Clay. 
3. Boulder-ciay (Lower). 


Seldom do we find the whole three members present in one 
section. Sometimes the Lower Boulder-clay alone is found; at 
others this is overlain by sand or gravel or clay, or by some or all 
of these rocks. In other cases we find the series complete. They 
occur almost continuously, in more or less completeness, from the 
sea-shore to an altitude of 500 feet; and from that level they appear 
in patches up to 1000 feet above the sea. 

The two Boulder-clays have the ordinary character, and are very 
much alike, except that the lower is tougher than the upper and 
contains larger boulders. ‘The character of the included stones is 
the same in both clays, as well as in the middle sands and gravel. 
Some most remarkable facts are presented by the distribution of 
these boulders*. 

Associated with these glacial beds, and occurring at various 
places, sometimes inland, sometimes along the sea-coast, between 
high- and low-water marks, there are a number of deposits of vege- 
table matter, which hitherto have been almost entirely neglected by 
geologists. 

When occurring on the sea-shore, these deposits usually pass 
by the name of ‘‘submerged forests.” I am, however, inclined 
to doubt the accuracy of this appellation. After a long and careful 
investigation into the nature of these deposits, I have come to the 
conclusion that they are not forests at all, nor the sites of forests, 
as will appear further on. 


* “On the Distribution of Boulders in West Cumberland, by iD) Kendall, 
C.E., F.G.S.,” vol. v. Trans. Cumberland Assoc. for the Advancement of Lit. 
and Science. 


30 J. D. KENDALL ON THE INTERGLACIAL DEPOSITS OF 


2. OBSERVED F Acts. 


Lindal deposit.—The first of these deposits to which I shall refer 
has already been dealt with by two other writers :— 

First, im vol. xvii. p. 274 of the Quarterly Journal, in a short paper 
by the late Mr. John Bolton, “On a Deposit with Insects, Leaves, 
&e.” ; secondly, in vol. xix. p. 19 of the same Journal, in a paper 
by the late Miss Hodgson. 

A comparison of these papers will show that the writers differed 
seriously about the facts with which they dealt. It is therefore not 
surprising that they arrived at very different conclusions. 

Owing to this discrepancy, I have been at some pains to ascertain 
the facts accurately ; and I give them below. 

The position of the deposit referred to by the above writers is shown 
on the map (Plate III. fig. 1). It occurs near Lindal in Furness. In 
figs. 2-4 a plan and two sections of the deposit are given. The data 
from which these sections were prepared are given below; they 
were obtained by a number of shafts and boreholes put down by 
the Ulverston Mining Company in search of hematite. The posi- 
tions of the boreholes are shown in the plan and sections. 


Sections of Boreholes. 


(Explanation of local terms used :—Pinel = Boulder-clay ; Black muck* = 
Vegetable deposit.) 


Borehole No. 41. 
Thickness Depth 


of each from 
stratum. surface. 
ft. 1. Tt, iA. 
Surface=soilk i. 25. cab onteenic an eee eee: 2 0 2 0 
Gikey spinel. seas <hq. came cteetiannste duets ease 66 0 68 O 
Iimestone. ce fhe eceeeseee cee eee 29 O 97 0 
Borehole No. 42. 
Stttace-soil: | Sea ey eee eee ane 1) 1 O 
Inte lay’ | L.. ca beset hee ae eee 3 0 4 0 
Gravel:andustones a peeeee ree eee eee eeeee 20 O 24 0 
Borehole No. 48. 
Surface-soil\ 722 aeece ae eee eee 1) 0 
GEBIVEL 256 oe nonce ee ee ee i @ 2 0 
Blue clay iii ee ee ei 8G o 6 
Pinel (blue) he eee eee 4 0 <1 6 
GAVE aco cacac cs ty gas Cee ee 9 O 16 6 
Gravellypimel 9). ees. eee ee ee eee 10 O 26 6 
black aniwek (woody) ss. © (5 
Blue Sarid 4.) a be 4 0 35 «6 
Grey. pine .2....85. 052 eal ee ee 5) 20) 40. 6 
MMMES TONE xy..::ideseteoteet cee Eee 2 0 A? 6 
Grrevajoimel i.) .0. Ges. anne eee 10 O 52 6 
Jointy limestone ....... waleie sa Gee 28 0 80 6 


MM c . . 

_* The use of the term ‘Black muck” is here clearly wrong, but quite a 
pardonable mistake for a borer to make. “ Black muck,” as that term is 
usually understood in Furness, does not contain any vegetable matter. 


2 


WEST CUMBERLAND AND NORTH LANCASHIRE. a) 


Borehole No. 44. 
Thickness Depth 


of each from 
stratum. surface. 
Hits abale ft. ine 
RECS Olan sae eee ena ae ee cose uae. 1 1 () 
ESN Gl aya 9 yee nce <a chtstecststiecsesseaa= 3 0 4 0 
STENT EL yu Ae 42 le ia ee oe 18 0O ya NY) 
PEPE TSLOMEH ete oe Ste eee oes cak dice ie DG 
Hed RUN CM ra. ceseds-. sores osaeset mata nce res 6 O 29 6 
STOW TTI ee ee ee a ene 9 0 38 6 
ICE: MUCH: (WOOGY) "2 y.5-.-csceeesc ees TO 5) 6 
LEE SON Lah seh ee ede eee ee pe Meri na oa 0 48 6 
LGLLON? GLa -taed sc tccRe seeeees tesa esa neEn ie a0 HIG 
Greys pinel) 25202... .5.)...2cses eee anaes 3.0 54 6 
Reddish pinel with limestone ............ oy 62 6 
REG! CLES conenee cee a eure mane ABE Eason eaenerrs 4 0 66 6 
{LEIDER ATS, 5 Bape Se ee re Pn ee 22 O 88 6 
SIGIUG GIR | RBRS noses eae saree nee aanee Bs AD) 91 6 
Borehole No. 45. 
Selb Gaecaill GE ee aan eee ei ae 113s ACG) i) 
PASSE psa Soe Boba Se ana ee 220 De 
aE NE a ots he le enc sc sleersaice ele ce Hae) 3f 0 
IBIQCTE MUCK (WOOGY ) ..200.020+020-2 0-00-2506 200 57) 
TE LIED SDUNGL TUES AS Aiea eal ee Be Rn Re foe Q 64 0 
CURRIN costes. st Ween some cae seoase CW (ie AD 
PREC lage es foe tees viewer ac secoe ncsecenis 10 O 81 0 
LD ASTESUOIIVO. 25 SSSR AUEe BAP RPE ER BURR Apne nthe ar Ae 2050 101 O 
Borehole No. 51. 
IEICE STUY IAS Lk a a bp aZe'O iy otk 
Meee piel os. . 525. 5. snacnnesciceccetssasseess 13 0 15 0 
ISeeye MINE esis oc vastsccsessccserscccsesce) | 40> O 58 0 
£2, DIG TLS Te EA ae eT 5 0 63°60 
Meme CLAY, |... 2. -e- wee Laseedesanecesazserees 7; 0 65 0 
Wellow SO8Sdn 2......i5<.csszszcccescsccctes 1 0 66. O 
[h.20, (UNDO) UG. Ae eee ee nen Asean erate 20 ua 
TCT OIHS sa cae Ae SBE aera ee eee 0 8 th ts 
Drarke pinelee sco< ccc) for seed cbt eee seed eons (0) TAS 
| TTT SST ROTI Ha 5, ae ee eee a 10226 88 2 
Berehole No. 52 
TL GGRCTOTL | 4 Bae eee ree tay See as YY) Zi 
LES, TELS) eee a ie ee ee 30 0 32 0 
S15) LNG) Se nee ee ae ee 36 0 68 0 
TE0) (UDnelh A On reer eee ener 9 0 TD 
2 SRE ETS E Gere ee ie Aa ee ee 10 O 87 O 
Borehole No. 58 
Brriietee SONNE och rat ee) eos oe te oo ck je car 
“Sse TOG) deg ee pace eee ee SiO 82 0 
22 UE TSG A Bane nee ae 20 86. GO 
eilack Muck (WOOdY).......2.c-s.0s00es0s-+ 24. 0 ITO. © 
“ELISE ROTI | RA a Oe Sat Oe ee ee 20 112 @ 
25G7, TID MIP Nee hee cote aera ek eee ee 3 O 11d O 
Red pinel and limestone .................. Dd 0 120; “0 
ison ore and’ StOneS ....2...c..6+0ssececeso- 2 0 176 
1 L POTTS) OTIS) gai oe eae le ae 0 10 122 10 


J. D. KENDALL ON THE INTERGLACIAL DEPOSITS OF 


Borehole No. 59. 
Thickness Depth 


of each from 
stratum. surface. 
ft. in. vita) 00 
Gin hacessOll wh esceesoceoe en Sas ee eee 2 0 20) 
(Civeyy jOMMVall, $4 Age ceoononoccdseosaso0q0esc0acuce 70 O 72 O 
Pack Much (WOOdYy))..0-.2-+ ease eee ene 10 eG 
Mplueusamd:) eee. va. acd. eats eee eee 2 0 75 O 
BLACIEGICULCK (WOOGTY, ics): 2-- cee eeecee ene il ©) 86 oO 
(Ble San yess vas she cciececeha ries mse eeeeeeee 4 0 90 O 
Regine), ese qucicec ce ss one eee eee 4 0 94 0 
Weimre stones. asactien seca ee cre nce eeee gy 103 0 
Borehole No. 60. 
Surface=soil’"... ote ccseeece scence ee eee eeee i) 0 
Red Spinel. 3c pki alo dare eee 30 0 3l 0 
Greyipimel ponent edocs ce coe earner ee 43 0 74 0 
THAD GOOG (CiOOENS)) anncoodescosescsesnsa%a: 12 © 36 0 
Bluewsand. “pe sves. cect scans cmesee cee enecene 2 0 838 0 
Wellowashiclaw ero suse eee ee ce eeeeeeee 2 0 90 O 
Broken ground (dragged by workings).. 22 0O ZO 
Limestone \ian.cecenes eee eee cee eee 4 0 116 @ 
Borehole No. 61. 
Surface-soil 7 ces cece eta 0 ib 
Grey pinell .2) a frase aes ancien eee 69 O 70 O 
Vellowapinelieereseenceceres Re, eee 20 ( Y 
Grey spinel 284 4e seen se ee eee 6 0 fier 
BIGche: MUch Ne die Oe eee eee eee Go) 3 S7aas 
iBlnevclaya(Sandiy)) eens eee ee eeeeee eee 3 0 90 3 
Limestone). ccc saanscce wer teden eae nee Oo 90 11 
Redspinell #224 4: gests cere eee ee netec ene 3 0 93 11 
Tumestonel. ears eee ee een eee i 94 11 
Borehole No. 62. 
Surface-soil’ 72. canoe eon nO ei) 
Grey pinel. 2.0). eaeareeerten ae meeenaee 88 0 89 0 
Black nick (woodsp\peeercceecsneeo ances If oO I0Oo oO 
Blue*clay -s.i)...ccda cree eee Caen 2A) 102 O 
‘Yellow sand’ \c)2) ahi ae eee eee 6 0 108 O 
IRedspinelli(withtoxe) my-eetee rer eee ene 4 0 112 O 
Noft stone Gwithiore)wee-eeeeeeeeeee eae 3 0 115 0 
Limestone with clay joints ..............- AL MD) 156 0 
Borehole No. 63. 
Surtace sot: os cucacc.carel ce heeeene eee 1 A) he @ 
Greyopimel co. 2. ...025. ah voce eRe eee eee ay O 88 0 
Black wniwek) (woody). ..2./s¢.cesanaa ete 10 Oo 98 oO 
BF uSielay i athe cncea dees Ahan nea ee 3 0 101 9 
ved apimell(wathyore) eee eee eee 4 0 105 O 
ivedipine li (wathistone) pase -oeeree eee 8 0 113 0 
iin CSLOME ransenecesscan eet ae eee eee On) ipa 


WEST CUMBERLAND AND NORTH LANCASHIRE. 33 


Borehole No. 64. 


Thickness Depth 

of each from 
stratum. surface. 
re aya foe ue 


Surface-soil ..... ME Can Gah seen reaes eae 1 O 10) 
BERG DUONG Bee dae siccncecsarciconiek asia stnies nessa 92°, 0 93 O 

RICH MUCH (WOOTY) .. icccccsecesclecedenes 20 0 13 0 
Bineclay  ....,<.- SOLOS ROREE CNeH OAC Beare Re 2 0 Milsy 
PROMEEC Ne iced wainacine Sia te oresielee eter 9 0 124 0 
LEDS OT AS) GRR rey a ep Re ees f O ll 124 Li 
MNOHEERIGHOMERE cas cc ccs si canaccaan ce wacscuncuces 0 8 WEY 7 / 
[TRONMEGINE: SaRs AP SOG RE a ee ean LS 126 10 
PROC ISLON Omen ss one ccc. tec saan tieaein 0 8 Ie 6 
PEICORE ee ech escscaesewcac sec ccseeccnusl 2 6 130 O 
Be ecg MM ESTOBG secs is nc wowace aaesecee' 20 0 HOO TO 


SMRRESCE SOMME eonece socicceaeanieswesa oaccnaeet LG 1 O 
RPE CHPNIN CY es sate ecieh Se tentisas sesaqas sins 99 0 100 O 
NEGIIUGH (WOOGY))...0.2.06..00000cess00e5 AO @ 1W6) ©) 
PEPER G) AY gaifoe nics chicane ce cislecicacieiviisiswesieidsis.si my AY) 122 0 
SLELLORP EEOC YE ee RO 125 O 
HRPM WOM Ome tes secoetecattc clas felosccetionwatloalsae 3 O 128 0 
Borehole No. 66 
SUREKCC-SOL Uecasccecacemerss Seacoast tak ag) 1 O 
BCR CMM EG Wa aise is Sine ueueid se seeleatiina be tewans 96 0 ot 0 
2r BULONTEP SENTING | So et Ts A) 98 O 
EDOELIS CIES MEE RE ee 1) 116 Oo 
PENITIEN SM Clere een eH Ch ecais eaten laccns wea es 2 0 118 0O 
BWrellFous POSsaM” Mo. c sects tenes eat venules 6 O 124 0 
TL BTIDOSTGTING Sec eie ae IS Eee ae L6G 125 6 


The pinel or Boulder-clay overlying the Black muck or vegetable 
matter in the above sections is the Upper Boulder-clay, having a 
greyish and ochrey matrix, and containing numerous boulders, some 
as much as 2’ 0" x1' 3” x1 3”; but the majority are below 3” in 
diameter. Coniston grits and flags, from the adjacent highlands, 
are the principal rocks represented. Besides these, however, there 
are St.-Bees Sandstone, Eskdale granite, Carboniferous Limestone, &c. 

The extent of the deposit has not yet been proved in cither a N.E. 
or §.W. direction, as shown on the section AB (fig. 3). So far as is 
at present known, it covers an area of about 34 acres. 

For a list of the plants &c. found in this deposit, the two papers 
above referred to should be consulted, as I had only an opportunity 
of seeing some of the vegetable deposit as 1t came out of the bore- 
holes. It was then too much broken up to enable me to identify 
any of the plants. 

Crossgates Deposits —Several deposits similar to that at Lindal 
have been met with at Crossgates, in working the hematite-mines. 
In some cases they were covered up by 9 or 10 fathoms of Boulder- 
clay; and they invariably rested on clay. JI have seen a large 
quantity of the woody matter which came from some of these de- 


GedaG.s. No. 145. D 


b+ J. D. KENDALL ON THE INTERGLACIAL DEPOSITS OF 


posits ; but no correct information has been preserved as to their ex- 
tent and thickness. Accumulations of similar material have been 
met with in the solid rock, as shown in the following section :— 


Section obtained by Boring at Crossgates. 
Thickness Depth 
of each from 
stratum. surface. 


fies a0 fie. nas 
Soil 


Danis Seer tec ate sie oars Uostasietdocioeeee aerate Het) 20 
Grayeland layer. cece. s-coen cee eee 24 0 26 0 
Decomposed limestone ............1--++-+-+ ge O) 43 0 
Yellow clay mixed with iron-ore......... 4 0 47 O 
TEUIE TUG a Me RE BORCSOE PEDO DOE ORCL EO C8.00% A (0) 5r To 
Tron-ore (dark-coloured) ...........,..0++ 2,10 53 0 
Black muck mixed with iron-ore......... Q, © 59 Oo 
PPOM=OVC, oe cerca sh oteael cle anen acenee eee 8 0 67 20 
Decomposed limestone m-.cse eeese eee os) 74 0 
LEUCOE THODUY WAVOSE see cnondccodscadnacecen: 2 86 0 
Decomposed limestone ........5....-....06 (ie 92 0 
Black mould and wood .......2...+.0.00+00¢ oy) 94 0 
Yellow clay mixed with iron-ore......... 16 0 110 O 
Black mould mixed with tron-ore......... LOE 120 <0 

ABU GCI I NOWLE SEN aot ee me ae ne Ly © nye) 


Black mould mixed with vron-ore and 
limestone 


Core cesses esesaseersessesseossossseons 


250 127 116 

Watney Deposit—On the western shore of the island of Walney, 
and about a mile south of the village of Biggar (fig. 1), there is a 
vegetable deposit, in many respects like that at Lindal; but not 
much of it is exposed. A plan and two sections of it are given in 

figs. 5-7. 

The deposit rests on Boulder-clay certainly ; but I have not been 
able to prove that it is overlain by the same formation. Still I 
think there is very little doubt about it when we look at the sec- 
tions, and w.cen we know that the shore is travelling rapidly land- 
wards. On the occasion of my visit I had not time to make any 
observations on the inner nature of the deposit with the view of 
determining the different kinds of plants enclosed in it; but I may 
state that, externally, it is very much like other deposits that I shall 
describe more in detail further on. 

Drigq Deposit.—On the shore opposite Drigg (Plate III. fig. 1). 
there is another woody deposit. A plan and several sections of it 
are given i figs. 8-11. In one part of it I sunk a shaft, as shown 
on the plan. A section of this shaft is given below. 

A. Vegetable matter (brown) and grey sand in alternating layers of various 
thicknesses, the vegetable matter predominating. I found in it the elytra 
of beetles, acorns, oak-leaves, hazel-wood, alder-wood and leaves, stems 
of common bracken, pieces of Sphagnum, seeds of various kinds and sizes, 
and rush-like stems and leaves, the stems standing on end and crumpled 
endwise, asif by downward pressure, the leaves lying on their side. The fol- 


lowing diatoms were found in this bed:—Cyclotella minutula, Pinnularia 


viridis, Pinnularia oblonga, Gomphonema acuminatum, Himantidium bidens, 
Himantidium pectinale. 


B. Yellow and brown sandy clay in irregular layers, 
C. Blue sandy clay. 


D. Red and blue sandy clay, the last 12 inches redder and more clayey. 


WEST CUMBERLAND AND NORTH LANCASHIRE. 30 


Some of the pieces of wood in A. were as much as 12 inches in 
diameter, but all lying on their side and partly flattened as if by 
pressure. 

Here, as at Walney, we find Boulder-clay under the deposit, but 
we have no direct proof of its having been overlain by that forma- 
tion. 

St.-Bees Deposit.—A few miles further north than Driggs, along 
the coast, opposite St. Bees (Plate III. fig. 1), there is another 
deposit, which I have examined more minutely than any of the 
others. A plan and two sections of it are given in figs. 12-14. 
In this deposit I have sunk several shafts and boreholes, sections 
of two of which (Nos. 1 & 8) I give below. 


Shaft No. 1. 


A. Vegetable matter (brown) and grey sand in layers of different thicknesses. 
The vegetable matter contaims seeds of various sizes, leaves and stems 
of rush-like plants, hazel-nuts, leaves and wood of the oak, alder, and 
hazel. The rush-like stems are numerous, and vary in length from 1 
to 3 mehes; they are standing erect, and crumpled endwise, like those 
found at Drigg. The following diatoms have been found in this layer :— 
Epithemia turgida, Epithemia granulata, Epithemia proboscidia, Pinnu- 
laria acuta. A vertebral column about the size of that of a rat was also 
found in this bed. 

B. Grey sandy clay containing a few rush-like leaves and the elytra of 

eetles. 

C. Hazel-wood, nuts, and leaves, also leaves of the oak, beech, and alder. 

D. Similar to A, but containing more leaves and hazel-nuts. 

(Boulder-clay.) 


Shaft No. &. 

A. Vegetable matter (brown), containing wood and leaves of the same kinds 
as those found in the same bed in Shaft No. 1. 

B. Fine oe clay with rush-like stems standing upright, and the elytra of 
beetles. 

C. Vegetable matter consisting almost entirely of leaves of the oak, alder, and 
willow, and hazel and alder wood. 

D. Brown sand and vegetable matter with some rush-like stems standing up- 
right and crumpled. 

On the surface of the deposit there are a considerable number of 
stems of oak and alder, the former turned black and the latter 
ee ; some of them are as much as 12 and 18 inches in diameter. 

T also found a large piece of the stem of a coniferous plant about 
18 inches in diameter. Under the microscope it was like the yew. 
These stems and branches all lie on their sides. In several parts of 
the deposit there are root-stocks of oak and alder with about 12 
inches of stem standing, as though they had grown where we see 
them: the rootlets, however, are imbedded only in vegetable matter ; 
they do not extend into the underlying earth. 

Near the upper edge of the shingle, on the beach, two boreholes 
were put down for the purpose of proving whether or not the vege- 
table deposit extended so far inland. One of these boreholes is 
shown in the section A B, fig. 13. The pees of it were as 
follows :— 

D2 


36 J. D. KENDALL ON THE INTERGLACIAL DEPOSITS OF 


Borehole No. 2. 
Thickness Depth 


of each from 
stratum. surface. 
fos yaa fits yum. 

Ths SIMMS Sorccsessogossscosdoopon0d0000000 1 6 LG 
2, (Senne eyarsl mph yell ccooansoancanséosnc0000 Ik 3.0 
By WGI QiOUG TUCIRIEP gboodbsedooonceHs50ce 3 6 11 6 
4, Boulder-clay (grey) ............se0ees ROO NEG 12 0 


The vegetable matter (No. 3) in this section was very soft and 
spongy, easily pierced by the boring-tool, and quite unlike that found 
in any of the pits or bores made on the shore, which was particularly 
difficult to bore through; in fact No. 3 is more like the soft peaty 
deposit which occupies the flat ground in St.-Bees valley. Two 
years ago a great length of drains was cut in this peaty matter ; 
and I had then a good opportunity of ascertaining the nature of it. 
It, however, was nothing like the vegetable matter which occurs on 
the shore, being much more spongy, and containing a far larger 
quantity of water, and altogether having more the appearance of peat, 
which in fact itis. The vegetable matter on the beach contains very 
little, or almost no water, notwithstanding that it is covered twice 
every day by the tide. Another difference between the two deposits 
is that the one on the shore is quite laminated, whilst that in the 
valley is totally devoid of lamination. About eighteen months ago 
Pow Beck, during a freshet, diverted its course just where it passes 
on to the shingle of the beach (see Section A B, fig. 13). In the 
new course a large patch of peaty matter, similar to that in the 
valley, was exposed. After seeing that, 1 had no doubt whatever 
that the vegetable matter found in borehole No. 2 was the same, and 
not at all like that found on the shore. 

Maryport Deposit.About halfway between Allonby and Mary- 
port, and about one third of the range of the tide from low-water 
mark (Pl. III. fig. 1), there is another vegetable deposit; but it 
is not very well exposed. 

I have not dug through this deposit, and therefore cannot say 
what is below it; and the soft silt, which covers the shore in 
the neighbourhood, prevents any information on that head being 
obtained. It has the same external appearance as all the other 
deposits, besides being laminated and very much more compact than 
peat. It contains a large number of seeds about the size of gun- 
shot, and a quantity of rush-like leaves, as well as pieces of hazel 
and alder wood. The wood is in every case flattened as if by 
pressure. | 

There is another and similar deposit about a quarter of a mile 
further on the shore towards Allonby; but only the upper surface of 
it can beseen. Near Beckfoot, I am told, there is another; but that 
I have not yet visited ; nor have I seen one which occurs at Cardunock, 
on the Solway. 


All the deposits described, whether on the sea-shore or inland, 


WIEST CUMBERLAND AND NORTH LANCASHIRE. oy 


have the same compact nature, and, so far as I have been able to 
ascertain, contain the same kind of plants. 


3. DEDUCTIONS. 


So little was known of the extent of the Lindal deposit at the time 
Mr. Bolton and Miss Hodgson wrote, that its real geological import- 
ance was missed. We now see that it is overlain by an immense 
mass of Boulder-clay, in some places nearly 100 feet thick; it is 
also underlain by Boulder-clay; so that I think we may fairly say it 
is interglacial. The impossibility of its being a recent introduction, 
carried down by means of swallow-holes, as suggested by Miss 
Hodgson, must be apparent to every one now that we know more 
of its extent and real nature. 

We may ascribe a like (that is, interglacial) age to the deposits 
at Crossgates. The deposits in solid rock were, I believe, thrown 
down in preexisting cavities in the Limestone at the same time as 
the deposits immediately below the Boulder-clay. 

The shore-deposits, so-called submerged forests, I think are also 
of the same age. We are not able to prove it so directly as in the 
case of the other deposits ; but I think that a careful consideration of 
all the facts must lead any one to the conclusion that they are 
interglacial. The fact upon which I most rely is their compact 
nature, which, to me, speaks most forcibly of their having been 
subjected to great pressure, such as would be the case if they had 
been overlain by the upper glacial beds. They are altogether unlike 
the spongy peat which occurs in St.-Bees valley, and quite as unlike 
any of the vegetable deposits which usually go by the name of peat, 
although many of the species of plants found in the shore-deposits are 
common to peaty accumulations. 

These shore-deposits all rest on the Lower Boulder-clay, a fact 
which is somewhat remarkable if they are of postglacial age. Why 
should we not find some of them on the Upper Boulder-clay ? 
Besides, how can it be for a moment doubted that the Walney 
deposit extends below the Boulder-clay, although this has not been 
directly proved. Referring to Plate III. figs. 6 & 7, we see that the 
deposit dips towards the land, and that it is only about 50 feet from 
high-water mark, where we have the Upper Boulder-clay. When to 
these facts we add that the shore is travelling rapidly inland, it seems 
to me we are bound to admit the Interglacial age of the deposit. 

Usually these deposits on the sea-shore pass by the name of 
“‘ submerged forests ;” but the conclusion to which I am led is, as 
already stated, that they are not forests at all, nor the sites of 
forests, but that the vegetable matter has been accumulated under 
water. All the facts but one point in this direction: the Diatoms, 
the rush-like stems and leaves (probably of a species of Sparganium), 
and the interbedded layers of sand, all speak of watery conditions ; 
the only fact which seems to me to indicate in any way that these 
deposits are the remains of ancient forests that grew on the spot, is 
the occurrence in them of a few root-stocks in their normal position. 


38 J. D. KENDALL ON THE INTERGLACIAL DEPOSITS OF 


But, as I have already pointed out, the rootlets from these stocks do 
not pass down into the underlying Boulder-clay, but are simply 
imbedded in the neighbouring woody matter. Some of these root- 
stocks belong to trees which must have been at least 18 imches in 
diameter; so that if such trees could have derived their necessary 
nourishment from a woody soil of this kind, it is perfectly certain 
they would not have been able to stand in it, because there would 
be nothing of any weight or tenacity for the roots to lay hold of. 
The presence of such root-stocks, it seems to me, may be better 
explained by supposing them to have been floated to the positions 
in which we see them. The position in which they now stand is 
that of flotation; that is to say, a root-stock would be floated and 
dropped in water with the same side up as when it was growing. 

The inner nature of the shore-deposits being precisely the same 
as that of the cavernous deposits found at Crossgates is also sug- 
gestive of drifting ; for clearly the latter are not on the site of an 
ancient ferest, but have been carried to their present resting-place 
by water. 

The facts presented by these deposits seem to me to have a most 
important bearing on the question of the formation of coal. We 
have here similar underclays to those which accompany cval-seams, 
and the same kind of intercalated clay-bands, both of which, in the 
case of coal, suggest watery conditions just as much as they do in 
the deposits under consideration. I think it is just as impossible 
that trees can have grown in the underclays of coal, which at the 
time would be soft and inccherent, as that they can have stood in 
the woody matter of the so-called ‘‘ submerged forests.” ‘The Stig- ~ 
maria rootlets of the coal may be accounted for in the same manner 
as the root-stocks which are found in vegetable deposits like that at 
St. Bees. The trees which have been found rising out of coal-seams 
and passing into the overlying strata, may be explained by supposing 
them to have been so loaded at the root when they were deposited 
that their position of flotation was erect. In this way the whole 
Coal-measures may have been deposited during a gradual subsidence 
without any of the periods of cessation demanded by those who hold 
that coal-seams are the remains of forests which grew in situ. 

On the assumption that the shore-deposits are “‘submerged forests ” 
of recent age, it has been often held that they indicate a subsidence 
of the coast; and writers, in consequence, have had to make the land 
rise and fall in a very remarkable and erratic manner, when dealing 
with the phenomena of “raised, beaches” on the one hand and 
‘‘ submerged forests” on the other. The view just enunciated sim- 
plifies matters very considerably, as, according to that, the vegetable 
deposits on the shore do not necessarily indicate either a rise or a 
fall of the land in recent times. 


} 
j 
| 
/ 
| 
j 
} 
| 
| 
} 
| 
| 
| 
| 


>= a 
SwncremavEN 
etek 
= 4 = 
= 
ae = 
= \ as => 
= = 
S % 
so 
= seice 
= . 
ms 


iromeere Bash £ Low Water 


Fig . 3. 
Section on Line AB Fig 2. 


Fig. 2 ae Fug. 4, 
, Sectton on Line © D 
Map of Lindal District ¢ 
wo" | 
2 c Fs 
ny *R , == 
[ew Te ; 
\ > = AX | — 9 
~ @LINDAL STATION | = - 
| th 
— | 
: to Se £ 
i Se ae me , 
= 2 ( = 
eee (EEN x Scale of Se 320 fe R cy 
Pinna are Mineslae pur ot cale of Selon Oe 7 uh. i 
: ° SUL SHAT. ‘ 4% FBedent of deposit. yt proved colored. nenlnal tint. } 
MEI SHAFT \ “ 7 [|| 
a Ss A Grey Pinel ( Boulder Gay) Zz 
x B Vegetable Deposit 
C Blue Sand (Cavey) 
D Red Pinel (Boulder Clav) 
E 
F 
. 


> Scale of Map 3 inches tw a Mile 


_ Journ. Geol. Soe. Vol. XATVIL Pi iit 


jae Ome 


Plan of Deposit om Shore oppose Prigg 


: Reference to Plan & Sections. 
™ Figs 8-11. 
A Vegetable Depostt 
, B Blue Any (Sandy, ) 
“4 C Red Cay in some places containing stunes(Boulde Tay) 
c De 3.2 5¢ | 
: \ Dip of Varrane of Woody atter- 


Carboniferous Limestone 

Gravel 

Borvholes 
evel liere 


Fig 1. 
Section on Line €E Fig-4. 
1 em A 


Fig. 10 
Section on Line © D. Fig.4 
Agt 


A 


Scale of Plan und Sections AB and CD 72 feet to an Inch. : 
Fig. 9. Secon on Line AB Fig. 4. 


Zz 
cS 
f os 
= 
= 
> 
=z 
oc 
> 
Zs: 
= 
Zz 
aa 7 
rats LINDAL 
esssorr 


Fig. 6. Section on Line AB Fug. 5. | 
High Water __ s Eyer 
eon 


C Boulder Clay 


A Fig. Ih. 
Scale of Plan & Section AB 100 feet to an Inch SM Plan of Deposit 
a y 2 ~ at S! Bees. 
velerence 4 
A Vegetable Deposu 1 ———— 
7 Glow 6 | fron ee Es SS SS \ 
Benet iiey . / Whitehaven; FURNESS RATEWAY 


Sa 
5 = 
Po, ! 


‘sts ‘ 
oy = eon is / 
red by Sand and 5°" er 0.47 

é i , : : 


SHINCLE 


Fig. 5. a H rid 
Plan of the Walney Depostt : { 


Fig 7 
Section on Line © D Fig. 5 
Chaves 8 
Seale of Section CD90 feet to an Inch 


= 


Norixontal Scale of 


Reference to Plan & Sections Fig. $5 1214. 


A Vegetable Deposit compact aa D® soft and spongy ke peat 

B Blue Sundv Clay 

C Roulder ve Sonatas Fig. 14. 

D Sand & Stones covering veg le osth = 

No Dep of lamina of Woody Matter eee rae Paes CEN Eee 
# iP 


Fig. 13. Section on Line AB Fig. R. = 
ee |) ; 


ot Scale of Section © DH) fet to an mck. 


Hugh Water 


Ww 
cl Y 


=== SS 


Plan & Section AB Iti hes 10 a Mile 


Vertical Scale cf Section AB IO feet to un uich. 


MAPS , PLANS AND SECTIONS OF INTERCLACIAL 


DEPOSITS OF WEST CUMBERLAND AND NORTH LANCASHIRE. - 


+ 
Frad® Dengerlieké Leth Luratim, 
=r 


# 


a 
Final 


WEST CUMBERLAND AND NORTH LANCASHIRE, 39 


EXPLANATION OF PLATE III. 


Plans and Sections illustrating the Interglacial Deposits of West Cumberland 
and North Lancashire. 


Rig. 1. Map of the district, showing the positions in which deposits of woody 
matter have been found, 
Figs. 2-4. Map and sections in the Lindal District. 
5-7. Plan and sections of the Walney Deposit. 
8-11. Plan and sections of Deposit on shore opposite Brigg. 
12-14. Plan and sections of Deposit at St. Bees. 


Discusston. 


Mr. ©. Rem said that in the Cromer Forest bed he had dug up 
many stumps of trees asserted to have been im situ, and had 
found them not to be so. He thought that great caution was 
needed in asserting trees to be wm situ. They would usually sink in 
a vertical position with their roots downwards. Sometimes portions 
of the soil in which the trees had grown were retained among the 
roots, and differed entirely from the matrix. 

Rey. H. H. Wrywoop pointed out. that trees often grew in a soil, 
but the roots did not pierce the underlying clay. Might not this 
be the case with the Cromer trees mentioned by Mr. Reid ? 

Rey. J. F. Braxz spoke of a boring at York with peaty matter in 
the midst of the Boulder-clays. 

Mr. Tippeman asked whether it was quite certain that there was 
Boulder-clay above and below these peaty deposits. 

The Sxcrerary replied that the author stated so distinctly in the 

aper. 
: The PRESIDENT spoke of the importance of comparing together 
the results of investigations in different areas. i 


40 PROF. T. G. BONNEY ON THE SERPENTINE 


4. On the Serpentine and assocrateD Rocks of ANGLESEY ; with a 
Nore on the so-called Srrprntine of PoRtHDINLLEYN (CAERNAR- 
VONSHIRE). By Prof. T. G. Bonnzy, M.A., F.R.S., Sec. G.s. 
(Read November 3, 1880.) 


1. ANGLESEY witH HonyHEAD IsLAND. 


TE serpentine from the vicinity of Rhoscolyn, though known within 
a limited area as an ornamental stone, does not appear to have 
received much attention from geologists. It is dismissed in Pro- 
fessor Ramsay’s memoir on North Wales in a few brief sentences, 
and has not, so far as I know, met with fuller notice in any other 
quarter. On the Geological-Survey map it is delineated as forming 
an elongated lozenge-shaped patch about a mile and three quarters 
in length from EK. to W., extending inland north of Rhoscolyn from 
the west shore of Holyhead Island, and occurring in several smaller 
patches on or near the opposite coast of Anglesey. In each of these 
localities I have studied the rock in the field, and have examined 
microscopically the specimens there collected. The results, I hope, 
may be of some use, though, after three separate visits, I have not 
been able to visit every one of the outcrops or examine minutely 
every part of the district. Owing to the intricacy of the coast in 
certain places, a map on a considerably larger scale than 1 inch to 
the mile would be required before an elaborate study could be 
undertaken. 

The questions which I have attempted to solve (as in all former 
studies of serpentine), were (1) its relations to the other associated 
rocks, and (2) its nature and origin. 

The ordinary rock in the vicinity of these masses of serpentine is 
a dull bluish or greenish schist, composed chiefly of minutely 
crystallized micaceous or chloritic minerals with some quartz, the 
foliation being parallel to the bedding. The latter is generally very 
distinct and sometimes exhibits very remarkable crumplings. The 
serpentine occurs in low rugged knolls, cropping out here and. there 
from the fields over which in the map the colour denoting its presence 
is extended—the general aspect of the rock masses reminding us of 
the serpentines of Cornwall, Scotland, and Italy. It differs markedly 
in all its characters from the ordinary schists of the district, and, 
where associated with the latter, occurs in such a manner as to 
suggest either intrusion or sporadic metamorphism of a very singular 
character. This is also suggested by the mapping; but it is even — 
more conspicuous in the field. 

My examination, however, showed me that under one name two 
distinet rocks had in some cases been confounded, viz. a true serpen- 
tine and a gabbro, especially in the Anglesey group. Jor instance, 
if we alight at the Valley Station on the Bangor-Holyhead railway, 
and follow the road to Four-mile bridge, we observe in the fields on 
our left, near Ty Newydd, a long craggy mass of serpentinous 


AND ASSOCIATED ROCKS OF ANGLESEY. 41 
aspect, the northern edge of the largest patch in Anglesey, 


according to the map. But on reaching this we find it a mass of 
gabbro, generally rather coarse—considerably altered, no doubt, but 


Fig. 1.—Junction of Schist and Serpentine. 


! ; 
\y { Ly, = t= 
~ La AP ZF- : ——— Ss 


AEF, SSS 


GLY 
Le il 


——s 


So : 
_——DS Sq 
TELL 
ee Se , 
—— SS Se 
a Se eee a 
A. Schist. B. Serpentine. C. Sand. 


unmistakable. There is, however, a small inconspicuous outcrop of 
true serpentine, separated from it by some grass, about seventy yards 
to the north of its eastern end. From the west end of this gabbro 
massif we made* for the shore. Here we found an irregular rocky 
coast-line fringed with skerries and islets, mostly dry at low tide. 
Some are schist, marvellously contorted, others gabbro, a few ser- 
pentine. Without a large-scale map, it would be impossible to 
make the details intelligible; but it may suffice to say that we 
worked carefully along the shore to the inlet by Tyddyn Gob, 
examining the different rocks. In one place the evidence, though not 
perhaps absolutely conclusive, seemed strongly in favour of the serpen- 
tine being intrusive in the schist; for if the junction were due to a 
fault, this would be a very strange one (fig. 1)7. Further on (beneath 
a wall) gabbro is seen intrusive in serpentine, which has assumed, 
as is not uncommon in such cases, a locally schistose aspect. <A 
large mass of gabbro forms an island near here. On the bank 
of an inlet near Tyddyn Gob is a rather schistose rock, which in 
the lower part resembles a foliated gabbro, in the upper a schist. 
Further to the south the schists are wonderfully contorted, and 
there is a small mass of serpentine near Ty Ucha (reached by 
crossing a causeway over an inlet); this is greatly crushed and 
slickensided. Appearances suggest an intrusive junction; but both 


* On my first visit to this place I was accompanied by Mr. F. T. 8S. Houghton, 
to whom | am much indebted for kind help on this and a subsequent occasion. 

t In the face of a crag facing S.E. and rising from the shore about the top of 
the letter ¢ in mile on the map. 


42 PROF. T. G. BONNEY ON THE SERPENTINE 


schist and serpentine are so much affected by subsequent changes 
that it is difficult to be sure. 

On the north-eastern edge of this massif of serpentine (Survey 
map), near Cruglas Farm, five shallow pits have been opened in a 
low ridge east of the road, two being close together by the road 
and outside the enclosure. In one the rock is a normal serpentine, 
as it is on the right-hand side of the other; but on the left we find a 
tougher variety (see below, p. 46). I failed, however, to find any 
line of demarcation between the two varieties. The next two pits 
are in normal serpentine, veined in places with calcite. The last is 
a kind of ophicalcite, hard and tough. On my first visit I found a 
loose block, near one of the western pits, of a dark green serpentine 
full of lustrous erystals, up to 0-3 inch diameter, of a mineral of 
nearly the same colour. I could not then discover any of this rock 
ar situ, and on a second visit, after a most careful search, aided by 
some of my students, was equally unsuccessful, though in one or 
two places the presence of a few minute crystals in the normal 
serpentine indicated a slight approach to it*. On the western side 
of the road a pit has been opened in a large mass of gabbro, some 
of which has a rather serpentinous aspect. 

About a quarter of a mile further down the road we passed over a 
reddish serpentine, rather brecciated and veined with calcite. We 
again touched a little serpentine (dull green) near Gwrthya. All 
the south-western end, adjacent to the shore, of the large mass near 
Melin Carnau, coloured as serpentine on the map, is gabbro. 
Thence we worked along the shore northward till opposite Ynyslas. 
There is no serpentine here, the gabbro being succeeded by schists, 
some of which are, indeed, massive and serpentinous in aspect; but 
the difference is evident, even in the field, on careful examination. I 
failed to find any serpentine at the shore end of the slope opposite 
Ynyslas; the rock there is the green serpentinous-looking schist ; nor 
did I see any in the isolated crags inland in the immediate neigh- 
bourhood. The railway crosses the mass which is represented on 
the map as running southward from Llyn Treflas; I walked along 
this, and found nothing in the cuttings but the green schist sueceeded 
by the ordinary schists of the district. 

A much larger mass of serpentine than any of the above is indi- 
cated on the map near Rhoscolyn, at the south end of Holyhead 
island. Specimens from this locality are familiar to me; but I have 
only examined the western portion of it in the field. As on the 
other side of the water, both schist and serpentine crop out here and 
there in rough knolls and ridges among the fields. The two most 
interesting exposures are on the western side of the western of two 
roads leading to Rhoscolyn. The first, near Ceryg Moelion, is a 
quarry approached by a driftway. Along this we pass a large mass 
of gabbro, which is succeeded by an outcrop of dark green serpentine ; 
and beyond this we come to the quarry. It has been opened in a mass 


* See below, p. 45, for a microscopic description. 


AND ASSOCIATED ROCKS OF ANGLESEY. 43 


of ophicalcite which is bordered by true serpentine on either side. 
The annexed diagram (fig. 2) roughly represents the relations of the 


Vig. 2.—Quarry at Ceryg Moelion. 


at 
Semmerene’ 


A. Serpentine, crushed and slickensided. 
B. Ophicalcite. C. Débris. 


two rocks. The boundary between them is generally quitesharp; and 
the serpentine is shattered, greatly slickensided, and in parts almost 
flaky, as if it had undergone great pressure. The ophicalcite varies 
much: in parts nearer the edge it is evidently a breccia of angular frag- 
ments of dark serpentine united by crystallized calcite ; in other parts 
the latter predominates and is mottled by a green serpentinous 
mineral without distinct evidence of fragments. The section is per- 
plexing; but, after careful examination aided by the microscope, I 
think that we have here a fissure which has been connected with 
some faulting and has subsequently been filled up. As the serpen- 
tine itself contains little or no lime, and as the schists of the island 
are not markedly calcareous, one is disposed to attribute the calcite 
to infiltration from overlying masses of Carboniferous Limestone 
which have been subsequently removed by denudation. 

Following the road for perhaps a quarter of a mile further south, 
we pass some more serpentine and several craggy outcrops of schist, 
and then in a field on the left find another interesting pit * which 
has been opened in order to work a band of steatitic rock. The 
annexed diagram (fig. 3) expresses the relations of these rocks. 

A is a green schist with a rather high N.N.W. dip. B is the 
band quarried. It is about 4 feet thick, but is rather irregular, 
seemingly conformable with A, and distinctly schistose in structure. 


* T believe it is called Plas Goch ; but it is most difficult to obtain informa- 
tion as to names in these wild parts of Wales; the spot must be very near the 
division between the two quarter sheets of the Ordnance map, 


44 PROL. T. G. BONNEY ON THE SERPENTINE 


Fig. 3.—Quarry at Plas Goch. 


A. Green schist, well marked dip. 
B. “ Steatite” schist, partially quarried out. 
C. Green schist, most massive and serpentinous-looking at x. 


It is very soft, like tale, and greasy to the touch, of a pale dull green 
or leaden colour. C is a less pure steatitic schist, which at D 
assumes a more normal character, and it dips to N.N.W. or N. xX is 
more like a serpentine, and has more the aspect of an intrusive rock ; 
it is of a dull-green colour, and has a slightly schisty structure, is 
harder than B, but can just be scratched with the nail. It contains 
numerous crystals of magnetite, commonly minute, but in places very 
distinct octahedra, which are sometimes nearly 3 inch in diameter. 
The matrix, however, on close examination, rather resembles a 
massive chloritic rock, like some of the lapis ollaris of the Italian 
Alps (e.g. that from near Chiesa in the Val Malenco) than a true 
serpentine. To the latter, under the microscope, it has little 
resemblance. It consists of a thickly felted mass of a scaly or 
fibrous clear mineral, slightly tinged with green, and feebly dichroic. 
The brightest tint shown with crossed Nicols is a dull greyish white ; 
and the mineral is either hexagonal or orthorhombic; minute gra- 
nules; some fairly clear, some brown to opaque, are scattered about 
the slide (the larger crystals have been apparently torn out in pre- 
paring it), part of which are probably chromite. After two visits 
to the pit, and the best study that I can give to the rock, I am dis- 
posed to think that it is more probably a massive chloritic schist 
than a true serpentine, and that the appearance of intrusion is 
illusory. The quarried rock (B) under the microscope exhibits a 
thickly felted mass of almost colourless folia, which, in transverse 
section, do not show conspicuous foliation, together with some few 
scattered granules of opacite. With crossing Nicols they afford fairly 
bright pink and green colours, and are almost certainly tale, so that 
the rock may be regarded as a tale-schist.* 

The following are the results of the microscopic examination of 
some of the above-mentioned rocks. The porphyritic rock found near 


* Tam since indebted to F. T. 8. Houghton, Esq., F.G.S., for the following 


AND ASSOCIATED ROCKS OF ANGLESEY. 45 


the quarry south of Cruglas consists of three minerals :—(a) a clear 
transparent mineral, with a satiny texture and occasional very faint 
indications of two cleavage-planes ; with crossed Nicols it presents 
a dark base crowded with irregular patches of minute fibrous micro- 
liths of a dull yellowish colour. (6) About an equal quantity of 
roundish or irregularly polygonal grains of a pale yellowish green 
mineral, which exhibits one marked set of cleavage-planes, and a 
second less perfect at right angles; these, especially the former, are 
further indicated by numerous grains and rods of opacite, probably 
magnetite. With crossed Nicols this second constituent is either 
black or shows a pale fibrous dull bluish mineral. The greatest 
absorption takes place when the cleavage-planes are parallel to one 
of the vibration-planes of the Nicols. (c) Scattered black grains and 
erystals of magnetite or possibly chromite. No structure resembling 
olivine occurs in the slide; so far as it 1s possible to conjecture the 
nature of the unaltered rock, it most resembles one that has consisted 
almost wholly either of two kinds of enstatite*, one richer than 
the other in iron, or of enstatite and hypersthene. I much regret 
that I could not succeed in finding the rock 7 situ. 

The Ty-Ucha rock, on the contrary, has the granular structure 
(defined by wavy irregular strings) which is often seen in an altered 
olivine rock. These, with crossed Nicols, exhibit a slightly fibrous 
structure and bluish-white colour, with an occasional golden-yellow 
tint, aS is common in serpentines, the interstices being dark or 
relieved by patches of dull bluish fibres. Opacite and larger grains, 
generally rounded, but sometimes clearly crystals of the isometric 
system, are scattered about the slide; probably these are magnetite. 
No appreciable quantity of any other mineral is present in the slide. 
This serpentine then probably results from the alteration of an olivine- 
rock, a kind of dunite. 


analyses:—I. is the dull-green chloritic rock, and II. the talcose schist. 
No. L., however, has not made it easier to identify the chief mineral 


it Ta 
a “fC 

eae Se Water’) Vane 286 
Pe age oh SLO ee 56°34 
50, ie eo NGOs. ai elieabe Deis 821 
TOM es, 2-87 TNS ORy Oe Sees aoecr ct eechoe Ue 
(Cie, Oe <RRbde pave en eea nee eee traces. ae % Rr Saas agg 2°00 
isa C) ea een Te) OV terete yareeroeee neat ce, traces 
ih Oo Je 1-73 Wa OE eRe Oe 0-52 
MrOM 1579 Mg Oversees. 25°43 
Gintepee 0-70 Na,O  vieeeeeteeee eines 0°79 

101-27 99-19 


* Prof. Maskelyne has described an enstatite rock (without olivine or other 
mineral, except perhaps diopside) from 8. Africa. Another slide cut from an 
outcrop in this neighbourhood presents some resemblance to the above, though 
the constituents are more minute and more highly altered, and there is less 
iron. 


A6 PROF. IT. G. BONNEY ON THE SERPENTINE 


The Ty-Newydd rock* is similar to the last, but contains a little 
of a mineral which, optically and in all other respects, resembles a 
rather altered enstatite. One of the black grains shows a parallel 
cleavage, as if it were a pseudomorph of enstatite in magnetite, a 
thing not very uncommon in the case of augite. A few strings of 
chrysotile traverse the slide. 

The serpentine from Ceryg Moelion consists partly of a fibrous- 
looking doubly refracting mineral akin to chrysotile, and partly of a 
clear, rather satiny mineral, dark or faintly granulated with bluish 
light, resembling one of thoso described in the block found at 
Cruglas (a), with thickly clotted opacite in irregular strings and 
some scattered grains. It may be an altered olivine rock. 

The rock from a small quarry about a quarter of a mile north of 
Plas Goch appears to be mainly composed of flaky mica-like plates 
of a doubly refracting serpentinous mineral, with the usual granules, 
rods, and clotted strings, probably of magnetite, and a very little of 
the mineral described above (5) in the block found at Cruglas. Of 
all those examined, it most resembles an altered olivine rock. 

A slide cut from the tougher serpentine in the quarry south of 
Cruglas shows a rather peculiar structure. The slide is traversed by 
irregular strings of clotted opacite, and is composed of two closely 
associated minerals—one the scaly serpentine described above, the 
other exceedingly minute scales of a doubly refracting mineral, 
dichroic, changing from pale yellowish to dull grey-brown, and with 
the two Nicols giving fairly brilliant colours. There is nothing 
inconsistent with “this serpentine being the result of the alteration of 
an olivine rock; but, so far as my experience goes, its appearance is 
exceptional. As stated above, I could see no evidence that this 
variety was intrusive in the normal serpentine. 

Microscopic examination confirms the view taken in the field, that 
several of the so-called serpentines are true gabbros. The specimen 
from the great mass near Ty Newydd shows the ordinary structure 
of a typical gabbro, though the felspathic constituent is wholly re- 
placed by secondary products (much being an opaque dust), being 
thus a variety of the saussuritic mineral so common in old gabbros. 
There is also plenty of a rather coarsely cleaved diallage, with pro- 
bably some ordinary augite, and a few grains of a more finely 


* JT am indebted for the following analysis of this ser penline to the kindness 
of B. T. 8. Houghton, Hsq., F.G.S. :— 
Wrater! 2120.0 ge nase decsaeeceae nee enanee 12°52 
No) is © RPA ACA as ioseubhencindn condoan wan udsoooo: 3862 
v0 O SR REPREP POM! isn cn nen Ut ane isco Qhudbccaences 415 
NCE 0 Riper eres eres Séscadeedoe bol Dicer 5:21 
1D @ MORENO Re SUNG Nobo cd oor hyedbedonansate 4°34 
VEO) Se con «cade ae One e nere Seee traces. 
OL 0 RRR AEs See orice ad uoasa)scuondosbuon co traces. 
A Fa 0 ERP RRM REN Ce Chadonscbancdnas an sacsoc 33°83 
Faull. <0 (21: Speen eR eREM iE Moca aciquhaaanuoraocdsac 0-70 


AND ASSOCIATED ROCKS OF ANGLESEY. AT 


cleaved fibrous mineral of an altered aspect. This may be only 
the result of pseudomorphic action on the diallage; but it may have 
been from the first a separate mineral. I incline to the latter view, 
as the normal constituent is sometimes altered, but seems then to 
produce a different mineral. 

A specimen from the islet near the south shore closely resembles 
the last. Here the pyroxenic mineral sometimes contains microliths, 
apparently of augite, sometimes exhibits lamellar twinning ; it is 
occasionally altered as in the last case. A third specimen, from a 
knoll on the shore between an outcrop of schist and another of ser- 
pentine, corresponds generally with the above. 

Specimens of the normal schist from near the chapel on the Holy- 
head island, from near Tycoch, and within about 5 feet of the ser- 
pentine at the presumed intrusive junction (see above, fig. 1, p. 41) 
have been examined. They are representatives of a class of schist 
of which I have seen and received numerous examples from Anglesey. 
Under the microscope they are seen to be composed mainly of two 
minerals, both of minute size:—one quartz; the other a greenish 
mineral in little scales*, fairly dichroic, changing from a greenish 
yellow to a strong green, occurring also in veins in little tufts: 
it resembles a chlorite more than a mica. There is an occasional 
granule of epidote and a good deal of a sort of grey earthy dust, 
sometimes in clots, probably in part decomposed magnetite or ilme- 
nite. There are also some yein-like bands of clear quartz, probably 
segregation-products. The third specimen has a rather coarser 
structure, and contains some scattered quartaz-grains which may 
indicate original constituents. These rocks show a banded structure, 
probably due to original bedding; but they are very highly altered. 
The schistose rock at Tyddyn Gob, as might be supposed from its 
aspect, is very different. It has a marked. foliated structure, being 
chiefly composed of wavy bands of an earthy mineral, almost opaque, 
and of a at colourless fibrous mineral, probably a variety of 
actinolite. Among the latter, in the lower and coarser rock, are 
the remains of a mineral with well marked parallel cleavage, from 
which the other may have been produced by secondary change. In 
appearance the mincral more resembles augite, and it is practically 
not dichroic; but in other optical properties it agrees better with 
hornblende; a little epidote is present. I cannot help suspecting 
the possibility of this rock being an altered gabbro with pressure 
foliation ; but if so, it is a most exceptional ease. 

From the above observations it will appear that the whole of this 
district needs remapping; and this task will not be an easy one. 
That the gabbro is an ordinary igneous rock cannot, I think, be 
doubted. “AS regards the serpentine, the evidence is a little less 
clear. The microscopic structure, owing probably to the great age 
of the rock, is rather obscure and anomalous ; its relations to the 
schist are not clearly displayed. Still the structure in several cases 
coincides with that of serpentines which we need not hesitate to 


* Not more than ‘001 inch diameter. 


A8 PROF. T. G. BONNEY ON THE SO-CALLED 


consider altered olivine rocks, such as those of Cornwall, Scotland, 
Italy, &c.; the general aspect of the rock in the field perfectly agrees 
with the same; and its relation to the schist would be most difficult 
to explain, except on the theory that it was intrusive in the latter. 
We must also remember that olivine rock partially altered has been 
detected in the Lleyn peninsula*. The evidence in favour of the 
serpentine being produced from the schist by some kind of selective 
or local metamorphism is of the weakest possible description. We 
therefore need not hesitate to add the Anglesey serpentines to the 
list of altered peridotites. 


2, THE SO-CALLED SERPENTINE OF PoRTHDINLLEYN. 


The northern and larger part of the promontory at Porthdinlleyn 
(west of Nevin) is coloured as serpentine in the Survey map, and 
thus noticed (by Mr. Selwyn) in the memoiry :—‘“* The rock at Porth- 
dinlleyn is a kind of coarse green and purple serpentine, with nests 
of red jasper ; veins of the serpentine are observed to dash in among 
the slaty series at Porthween.” So far as true serpentine is con- 
cerned, [ might have modelled this note on the well-known chapter 
concerning the snakes in Iceland; for there is none at Porthdinlleyn. 
The rocks, however, which I examined in my search for serpentine 
are so interesting that I am tempted to add a few words concerning 
them. 

Specimens of several of these rocks have been examined microsco- 
pically ; and one or two exhibit structures of much interest. On the 
present occasion, however, I shall not enter into details foreign to 
the immediate,subject, but only endeavour to identify the specimens. 
The first rock seen (by a landing-stage) as we approached Porth- 
dinlleyn along the shore from Bwlchdinlleyn is a massive green rock 
of serpentinous aspect. In the craglets and reefs near the hamlet 
itself we find a similar rock which also shows in parts a distinctly 
brecciated structure, pale-green fragments of various sizes (generally 
up to 2 or 3 inches diameter, but in one case about 18 inches) 
occurring in a darker and more ashy-looking matrix. In other 
parts the structure more resembles brecciation in situ ; sometimes 
the rock is almost spheroidal, and reminded me of certain instances 
which I have seen in old igneous rocks of a rather basic nature, where 
the outer shells of the spheroid had become decomposed, simulating 
a conglomerate with but little matrix. A reddish variety of the 
compact rock also occurs ; and these rocks continue for some distance 
to the north of Porthdinlleyn, the “ nests of red jasper ” (hematite 
with quartz, calcite, &c.) being in places very conspicuous. In the 
field it is very difficult to pronounce upon the nature of the above 
compact rock. It is hard, with a subconchoidal to rather argil- 
laceous fracture, traversed by well-marked divisional planes, but 


«x Geol. Mag. dee. ii. vol. vii. p. 208. 
t The Geology of North Wales, p. 170. 


SERPENTINE OF PORTHDINLLEYN. 49 


with no certain indications of bedding. The rock exhibits spots of 
epidote, varies in colour from a dull sap-green to a greenish-purple 
colour, is traversed by thin veins of quartz and calcite, and 
might be taken either for a compact chloritic rock resulting from the 
alteration of a uniform fine sediment, or for a very compact diabase. 
Under the microscope both varieties are seen to be rather decom- 
posed, the slides being rendered more or less opaque with fine 
powdery ferrite, which sometimes also occurs in granules. There 
are indications of numerous microliths, some doubtless being felspar, 
and probably a plagioclase ; these, in places, show a tufted, sometimes 
an almost spherulitic arrangement. In this are scattered some 
larger augite crystals, much altered but still recognizable. The 
green variety has some faint indications of a fragmental structure, 
and might be a tuff in which the outlines of the fragments were 
nearly obliterated. The structure of the redder variety is more 
distinctly that of a true igneous rock; and as I could see no signs of 
a division in the field, we may venture to identify these as compact 
diabases*, very probably old flows of a basaltic lavat. The breccia 
exhibits under the microscope a base of filmy pale-coloured viridite, 
in which are scattered numerous granules of ferrite, crystals of 
epidote, with other minerals of secondary formation, altered augite, 
and felspar, and scattered fragments, apparently of more or less 
glassy lapili. Of the nature of some of the last there can be no | 
doubt. The rock, then, is a ‘“diabase tuff,” of which probably a 
good deal was once a very fine dust. 

For some distance to the north of Porthdinlleyn the dubiously 
“‘ conglomeratic ” aspect of the rock as above described continues ; 
and then we find, on arriving within a short distance of the life- 
boat house, sections that, even in the field, are perfectly conclusive. 
The best occurs in a little cliff (an old quarry-face) looking seaward. 
On the sides of this (the section is about a dozen yards wide) we 
find a dull-green ashy-looking matrix, full of angular and subangular 
fragments of all sizes up to 3 or 4 inches diameter of a compact 
paler-coloured rock; and in the middle a mass of rounded blocks (a 
foot or two in diameter) which seems almost solid passes irregularly, 
but with fairly marked boundaries, into breccia. There can, then, be 
no doubt that we have here a true volcanic agglomerate, including 
either a thickly fallen mass of bombs or a small lava cowlée. This 
view is fully confirmed by microscopic examination,—the breccia 
exhibiting a ground-mass, as above described, full of scattered frag- 
ments obviously of volcanic origin, some & brown slaggy glass, like 
a, kind of tachylite, others compact basaltic scoria. The examination 
of a spheroidal mass from the agglomerate shows it to be highly de- 
composed compact diabase. 


* The name diabase is given, though little or no chlorite can be recognized, 
because I know of none other that exists. 

t+ I may mention that I have in my collection a rock microscopically identical 
with the above, which I collected between Glyn Garth and Beaumaris. ‘These 
old basalts are, i suspect, not uncommon among the Welsh Pre-Cambrians, and 
in the field are hard to recognize. 


Q. J.G.S. No. 145. E 


50 PROF. T, G. BONNEY ON THE SERPENTINE 


Proceeding a little further, we find, about fifty yards south of the 
boathouse, a compact green rock with indubitable spheroidal strue- 
ture. This rock now continues along the northern and western face 
of the peninsula, becoming in places very distinctly spheroidal. The 
face of a crag (a joint-plane) near the sea at the N.H. angle exhibits 
one of the most conspicuous instances of the structure that I have 
ever seen. The rock, where not spheroidal, is compact and sharply 
jointed, in one or two cases a little platy in structure. Under the 
microscope it is seen to be an altered basalt, the structure being 
recognizable, though the felspar is replaced by earthy decomposition- 
products, and the grains and crystals of augite are much altered. 
1 also noticed one or two small dykes, seemingly of late date. The 
eastern side of the peninsula had occupied so much of my time that IT 
was unable to examine into the relations of the above group of rocks 
with the schist of the district (which is not to be seen on that side), 
and had to be content with satisfying myself that, though there are 
igneous rocks, there is no true serpentine at Porthdinlleyn. 


Discussron. 


Mr. Baverman had examined the steatite-deposit of the locality, 
and had arrived at much the same conclusions as the author. 
There were true serpentine rocks, and doubtful or “‘ serpentinous ” 
ones. He considered an examination of the composition of these 
rocks desirable. 

Dr. Hicks congratulated the author on speaking of the mtrusive 
origin of the Anglesey serpentines with a certain degree of doubt. 
He was himself inclined to regard these rocks not as intrusive, but 
as altered representatives of certain ancient sediments. He repudi- 
ated the notion of “ selective metamorphism ” being generally ap- 
plicable to these rocks. The rocks at Porth Dunbar he regarded 
as mainly indurated and altered ashes. 

Prof. Huenrs pointed out that when the Survey map was made 
the definition of “serpentine” was much wider than that which 
was now adopted. He was not convinced of the intrusive cha- 
racter of the serpentines in the section given by the author, but 
thought it might be one of the numerous cases in which during 
the crumpling of the gnarled series the harder masses were pro- 
truded through the schists, producing small local faults. He believed 
that calcareous beds in the gnarled series, and magnesian minerals 
in the dykes, generally associated with the serpentinous rock and 
ophicalcite, were sufficient to account for both. 

Mr. Hupirston had examined the rocks of the district, and 
thought they differed so greatly from the Cornish serpentines as to 
make it somewhat doubtful whether they could have been formed 
by the alteration of olivine rocks of intrusive origin. He was in- 
clined to think that the Anglesey serpentines, like those of the 
Shetlands described by Dr. Heddle, might in part be altered acti- 
nolitic rocks, 


AND ASSOCIATED ROCKS OF ANGLESEY. sy | 


The Avrnor doubted whether the so-called steatite is really the 
normal mineral. Dr. Hicks had entirely misapprehended him ; for 
he had no doubt that the true serpentines of the district were 
altered olivine rocks, and not metamorphosed schists. He con- 
sidered the use of the term “ serpentine” by the Survey, if it were 
as stated by Prof. Hughes (which he doubted), was one which was 
quite unjustifiable ; nor could he agree with Prof. Hughes’s expla- 
nation of the section showing an apparent intrusion of the serpentine 
into the schist. He could not admit that either the Anglesey ser- 
pentines or any Scottish serpentines which he had himself examined 
could have been formed by the alteration of actinolitic rocks. 


5y4 PROF. SEELEY ON A SMALL LIZARD FROM THE 


5. On Remains of a Smart Lizarp from the Neocomian Rocxs of 
Comin, near Trreste, preserved in the GrotogicaL Musgun of the 
University of Vienna. By Prof. Srxrny, F.R.S., F.G.S8., of 
King’s College, London. (Read December 1, 1880.) 


[Puars IV.] 


Prorrssorn Epuarp Svzss, F.M.G.8., recently received from Comén, 
near Trieste, a specimen showing the hinder half of the skeleton of 
a lizard which he has desired me to describe. Unlike the two fine 
slabs from the island of Lesina, preserved in the k.-k. geologische 
Reichsanstalt, which are in a pale yellowish limestone matrix, 
this specimen is from a limestone slab nearly black; the animal 
shown upon it has lain exposed for some time in the quarry and 
suffered by the solvent action of the rain. Prof. Suess mentioned 
to me that the colour merely indicated one of the many alterations 
in the limestone, and that, since it was collected by a former pupil, 
no doubt could attach either to its stratigraphical or geographical 
position. Prof. Kornhuber does not appear to have been quite cer- 
tain as to the position of the Lesina rocks in the Cretaceous series ; 
but while [ was in Vienna, Professor Pisani mentioned to me that 
he had identified thirteen species of fish with Upper Neocomian 
species ; and as fish constitute the chief fossils of the deposit, this 
must be held conclusive evidence of the geological age of these 
lizards. 

This new fossil (PI. IV. fig. 1) at first sight presents a considerable 
resemblance to the Hydrosaurus lesinensis of Kornhuber, as was 
pointed out to me by Prof. Suess; but the differences are so 
remarkable and important that I find myself unable to inelude it in 
the same genus. The specimens in the Museum of the Imperial 
Geological Survey were shown to me by the Director Franz Ritter 
von Hauer. They are admirably preserved, and, as Koruhuber has 
stated, appear to indicate an animal with 9 cervical, 30 dorsal, and 
2 sacral vertebrae, and a tail of which only the 24 anterior vertebre 
are preserved. The type 1s distinguished by the remarkable stoutness 
of the dorsal ribs, by the very long and large neural spines and early 

caudal vertebrae; and it possessed | well-developed limbs, of which the 
hinder pair were much larger than the anterior pair. The specimen 
which I have now to describe has only the hindermost 12 dorsal 
vertebre preserved. There are presumably 2 sacral vertebre ; and 
then succeeds the tail, of which about 65 vertebra are preserved 
or indicated by impressions ; and it is probable that more remained 
in the slab which had been adjacent but was not collected. 

The length of the 12 dorsal vertebra is about 55 millims.; and 
the remainder of the vertebral column, as preserved, measures along 
the curves of the tail nearly 200 millims. The dorsal region is ex- 
posed so as to display the attachment of the ribs. The ilium is the 


Quart. Journ. Geol. Soc. Vol. XXXVIL.PL IV. 


ST 


tf) 
g 
re 


A.S.Foord del. et lith. dias Mintern Bros. mp. 


ADRIOSAURUS SUESSII. 


NEOCOMIAN ROCKS OF COMEN, NEAR TRIESTE. D3 


only pelvic element clearly exhibited; and that is directed backward in 
the usual lacertian manner (fig. 1,1). The hind limbs are preserved, 
though they were fast becoming obliterated by weathering. The caudal 
vertebrve lie upon one side, and consequently do not give a very dis- 
tinct idea of their forms. They, however, show no trace whatever 
of the neural spine, though the chevron bones are well developed, 
and are preserved down the greater part of the tail. The dorsal 
vertebrae apparently rest upon the neural surface and expose the 
visceral surface. This may be inferred from the form of the cen- 
trum, the curvature of the ribs, which are concave in length as 
exposed, and the fact that the ilinm underlies the femur; but the 
face of each centrum has been somewhat dissolved, so that 1b cannot 
in any ease be said to exhibit the unaltered appearance of the bones. 
Each centrum in the dorsal region is rather less than 3 a centim. 
in length, and is about 6 millims. widein front. The sides converge 
posteriorly to about 3 millims. ; so that, besides being a far smaller 
animal, the lower dorsal vertebree appear here to be relatively 
shorter, and the centrum, instead of having the concave lateral 
outlines of Hydrosawrus, has its side rather convex in length; 
and the anterior surfaces on each side of the anterior articulation 
look more forward and less outward than in that genus. There 
1s, moreover, no indication of the transversely concave outline of 
the intervertebral union—but in place of it a notch in the ante- 
rior border, as though there were a small ossicle at the junction of 
each two vertebre; but this apparent notch may be nothing but 
the neural canal exposed in this position by the thin base of the 
centrum being there dissolved. As the vertebree pass downward 
towards the sacrum, their aspect seems to be less massive, and the 
posterior end becomes a little more compressed from side to side. 
There are slight indications of two ridges running longitudinally 
on the base of the centrum from the outer corners of the anterior 
cup towards the posterior articular ball. All the dorsal vertebree 
were furnished with ribs; but they become shorter and smaller 
towards the sacrum. On the right side of the specimen they lie 
together, touching each other along their lengths, except in the case 
of the last three or four, which are only indicated by impressions. 
The ribs appear to be flat on the under side and moderately curved ; 
the rib of the third vertebra preserved is 2 centims. long and 2 
millims. wide at the proximal end. The ribs appear to be tubular 
and single-headed; they taper evenly to a blunt point, but scarcely 
give the impression of being relatively so stout as the ribs of Hydro- 
saurus. The sacral vertebra are no better defined than those of the 
dorsal region, being partly covered by femoral bones. ‘There is an 
angular bend in the tail, beyond which the vertebre le on their 
sides more perfectly displayed than in the case of the first few caudals. 
The neural arches of the caudal vertebre were low, without any indi- 
cation of neural spines, the neural arch being concave superiorly from 
front to back, and articulating with the arches of adjacent vertebre 
by zygapophyses, which were elevated high above the neural sur- 
face. The neural arch widens in front, and is smooth at the sides. 


ot PROF. SEELEY ON A SMALL LIZARD FROM THE 


There is no trace preserved of transverse processes, such as may be 
presumed to have existed. The chevron bones are relatively long and 
slender and directed backward parallel to each other. The vertebrae 
rapidly decrease in size, and in the latter half of the tail preserved 
are small. These hindermost vertebre (that is, after about the 
thirty-fifth) appear to develop a slight neura: spine, which is slender, 
directed backwards nearly horizontally, and terminates without any 
decrease in thickness, in a rounded end; but these spines can only 
be detected in some eight vertebra. The neural arch appears to be 
preserved to the end of the series, where the whole lateral measure- 
ment of the vertebra is less than 2 millims., and its length does not 
ereatly exceed 2 milims. The height from the base of the centrum 
to the middle of the neural arch at about the eighth caudal is 34 
millims., the neural arch forming less than half of this height. The 
union between the centrums is not well defined. The chevron bones 
in the first dozen vertebre appear to be about 53 millims. long. 
There thus appear in the tail to be differences from Hydrosaurus 
desinensis in the relatively small development of the neural spine, 
which never extends upward as a broad plate in this form, and, 
when it does exist, is a slender backwardly-directed process. The 
earlier chevron bones in the present fossil appear to have been rela- 
tively broader; and in the absence of any indication of transverse 
processes, it is impossible to affirm that those processes existed. 
The pelvis is imperfectly seen on both sides. The ilium extends on 
the right side parallel to the vertebree (fig. 1, 1), but its anterior part 
is covered by the head of the femur. The part exposed is fully 7 
millims. long; its outer edge is rounded; it expands anteriorly a 
little, but appears to be distinguished by its slender form and parallel 
sides. On the left side of the specimen, underlying the other femur 
and articulating with a bone in front, which may well be the arti- 
cular part of the ilium, is a curved bony element (nearly 8 millims. 
long, wider than the ilium), which from its position might well be 
the ischium. ‘The bend in it occursin its anterior third ; 1t is com- 
paratively slender, and only expands a little at its distal end. The 
slender form of the ilium and its relative length are points of differ- 
ence from the species with which this has been compared. Both hind 
limbs are fairly well preserved. The femur of the left side (fig. 1, m) 
is about 11 millims. long, a good deal constricted in the middle, and 
flattened and expanded at the distal end, which is concave from side 
to side; the anterior margin of the bone is concave in length, while 
the posterior margin is more straight. The characters of the arti- 
cular head are not well defined, owing to the way in which the bone 
is compressed; but the head appears to have been well rounded, and 
to have measured about half a centim. from front to back. There 
are no indications of the distal epiphyses represented by Kornhuber. 
The tibia (fig. 2, ¢) has its anterior margin straight, and its pos- 
terior margin concave. It is about 7 millims. long. The proximal 
end is greatly expanded; and the shaft is relatively more slender than 
in Kornhuber’s species. The fibula (fig. 2, f) is a rather more slender 
bone, without any indication of proximal expansion ; it widens at 


NEOCOMIAN ROCKS OF COMEN, NEAR TRIESTE. 55 


the distal end to 3 millims., which about corresponds to the proximal 
width of the tibia. The tarsal bones (fig. 2, ts) are not well preserved, 
but appear to consist of one large bone below the tibia, which is appa- 
rently polygonal, and two smaller bones placed below the fibula; but 
there has probably been at least a third bone, which was distal in 
position and is not preserved. The metatarsus (fig. 2, mt) is not very 
distinct ; and it is difficult to say whether the metatarsal bones were 
entirely distinct from each other. The specimen would appear to 
indicate that there were at least three metatarsals. The bone 
below the large tarsal element is short and broad, reminding one in its 
proportions of that of a Plesiosaur. It articulates by a large proximal 
facet with the large tarsal bone, and by a small facet at the proximal 
end with the distal tarsal element adjacent to it. Its width at the 
proximal end is over 2 millims.; and it appears to be about 4 millims. 
long. The metatarsals on the outer side are not preserved. There are 
five digits (fig. 2,1-v) formed of slender bones with a median groove 
on the dorsal surface of the first row. There are two bones in the first 
digit, which is short and terminates in a pointed bone, the distal end of 
which is a little curved and apparently carried a claw. There are 
three bones in the second digit, which is about twice the length; 
but the fracture in the slab passes through the middle of the ter- 
minal small phalange. In the third digit there may have been four 
phalanges; but the fracture passes through the third, which is so 
large that it is not likely to have carried a claw. There appear to 
have been four phalanges in the fourth digit; and in the fifth the 
number cannot be satisfactorily determined ; but there do not appear 
to haye been more than three. The three inner digits have the 
proximal surfaces of the proximal row of bones in the same line ; 
but the two outer digits look as though placed a little higher up, 
which is in accord with the ordinary Lizard plan. The totallength 
of the longest digit as preserved is about 9 millims.; and the total 
length from the proximal end of the tibia is a little over 21 millims. 
It will thus be seen that the tarsus and digits differ considerably 
from those indicated by the animal described as Hydrosawrus lesi- 
nensis. 

It is of course with this type that the present specimen must be 
chiefly compared; and the form and proportion of the dorsal ver- 
tebree, the mode of articulation of the ribs, and the characters of the 
caudal vertebrae, especially in the neural spine and transverse pro- 
cess, indicate a distinct type. The pelvis is also distinct, while the 
proportions of the segments of the limb differ in as remarkable a 
way, and necessitate placing this fossilin a distinct genus. Though, 
from the imperfect preservation of the specimen, its more important 
characters remain unknown, it may be conveniently distinguished 
as Adriosauwrus Suessit. Its affinities need the assistance of more 
perfect remains for their elaboration. 

I am indebted to Prof. Suess for the opportunity of making this 

ecord, and adding another species to the Secondary representatives 
f the lizard group. 


56 ON A SMALL LIZARD FROM THE NECOMIAN ROCKS. 


EXPLANATION OF PLATE IV. 


Fig. 1. Tail of Adriosawrus Suessit, natural size: 1, ilium; m, femur. 
2. Hind limb, from right side, enlarged: 7¢, tibia; f, fibula; ¢s, tarsus ; 
mt, metatarsus ; I, I, Il, Iv, V, digits. ; 


Discussion. 


Mr. Huzxe said that he had no doubt that the species described 
by Prof. Seeley belongs to a genus distinct from Hydrosaurus. He 
compared the former with the Geckos in respect of their having 
bony scales. 

The Avurnor stated that the distinction of the vertebrae had been 
rendered difficult through chemical action. He doubted as to the 
existence of scutes in the specimen. 


AMMONITES FROM THE INFERIOR OOLITH. oi 


6. On the Terminations of some Ammonites from the Inrertor 
Oorrre of Dorser and Somerset. By James Buckman, Hsq., 
E.G.S., F.L.S., &e. (Read June 23, 1880.) 


Durine the progress through the press of D’Orbigny’s ‘ Palconto- 
logie Francaise, ‘‘ Terrains Jurassiques,” I was busily engaged in col- 
lecting the fossils of the Inferior Oolite and Lias in the neighbour- 
hood of Cheltenham. In both of these rocks I had the good fortune 
to find a somewhat large series of Ammonites, the greater propor- 
tion of which I could readily identify from D’Orbigny’s figures*. 

But as these fossils were in bad condition, especially when com- 
pared with the French drawings, I was at first almost led to think 
that these latter had been somewhat unjustifiably made up or, as 
we should say in plain English, “ fudged.” 

Thus, in reckoning up the plates of admitted Inferior Oolite Am- 
monites in the ‘ Paléontologie Francaise,’ we find them to be thirty- 
eight, out of which no less than twenty species (a httle over two 
thirds of the list) are drawn with the mouth of their shells more 
or less perfect; and yet, strange to say, I do not recollect a single 
example of their Cotteswold prototypes having occurred to me in 
this perfect condition. 

Since those days of scepticism, however, D’Orbigny’s book has 
become my constant and trusted companion, as the specimens in 
our Dorsetshire list of Ammonites appear to be in much the same 
condition as those figured by that talented author. 

During the last seventeen years my lot has been cast in the 
pleasant county of Dorset; and, curiously enough, the farm that I 
haye occupied is situate on the Inferior Oolite ; the Halfway-house 
quarry is within a mile of my own residence; whilst a quarry in 
one of my own fields, within a hundred yards of my house, has 
proved to be one of the richest in England, if not in the world, in 
Oolitic fossils, and especially in the species of the Cephalopoda 
and Gasteropoda, in each of which great classes we may safely 
reckon as many as from sixty to seventy species. From this quarry 
and the surrounding district, there haye been determined as many 
as twenty-eight species of Brachiopoda; and I probably possess 
nearly, if not quite, a hundred forms of Lamellibranchiata, which, 
not to name other remains, cannot but be considered as a goodly 
list. 

If, again, we consider that the greater part of the individuals of 
these lists occur in a bed varying from 2 to 3 feet in thickness, 
the surprise becomes still greater ; and net only are the species so 
numerous, but it is one mass of specimens, as, instead of our merely 
finding an Ammonite here and there (as in the Cotteswolds), this par- 
ticular bed is a mass of them, so that several species will be huddled 
together in a small space; and, in so far as the Ammonites are 


* The part on the ‘‘ Céphalopodes” was finished in 1849. 


58 J. BUCKMAN ON AMMONITES FROM THE 


concerned, on the same slab will be representatives of several of the 
zonal divisions of different authors. This may be judged from the 
fact that on the surface of slabs from this bed other Ammonites 
besides the following have been noted. 


Ammonites jurensis, Zev. Ammonites Gervillii, Sow. 
—— Sowerbyi, Miller. —— Brongniarti, Sow. 
—— Humphriesianus, Sow. —— subradiatus, Sow. 
—— Braikenridgii, Sow, —— leviuseulus, Sow. 
—_— linguiferus, D’ Orb. —— discus, Sow. 

—— Sauzei, D Ord. —— Parkinsoni, Sow. 


Some species prevail to such an extent that hundreds of indivi- 
duals may be found in a square yard of the bed, one species prevail- 
ing at one quarry, and another at other quarries. 

So numerous are individuals in some quarries, and so varied do 
they seem, that it is hardly possible to escape the conclusion that the 
family must have hybridized to a considerable extent. Anyhow, 
under such circumstances, it is exceedingly difficult to accurately 
define species—much more so than in the Cotteswold district, where 
it would be difficult to find more than half a dozen forms in a 
day. 

But besides this, when we compare the Dorset and Somerset Am- 
monites with the Gloucestershire forms, we find that the former 
are usually sharp and well preserved, while the latter are consider- 
ably rougher and more fragmentary. Thus it is that shells with 
their terminations are rare in the latter county, while with us they 
are comparatively common. 

The specimens laid before the Society are a sufficient evidence of 
this fact; and they are fast increasing, now that this matter is being 
more carefully investigated, the truth being that collectors seem to 
have vied with each other in squaring up the mouths to make them 
neat and tidy. Many among them, indeed, would cut away ald of 
the outer chamber, preserving only the inner chambers, which, being 
more or less filled up with carbonate of lime, were said to be alive, 
whilst the outer one, filled up with indurated mud, was pronounced 
to be dead, and so was removed as useless. 

But not only do we find our Dorset specimens to be much like 
the French ones (as figured by D’Orbigny) in condition and state of 
preservation, but the species from the two sides of the Channel are 
almost identical: thus we have within jour of the whole of D’Or- 
bigny’s figured specimens from the Inferior Oolite of France. 

But, besides this, there are several species in our own Cephalopoda- 
bed, now admitted to be high up in the Inferior Oolite, which 
D’Orbigny has allocated to the Lias. This, we take it, has been 
mainly due to the fact that this bed has been confounded with the 
ene at the base of the Inferior Oolite in Gloucestershire. 

In this list are the following :— 


Ammonites insignis, Schiidl, Ammonites Germaini, D’ Orb. 
—— variabilis, D Ord. —— jurensis, Zict. 
—— Murchisone (?), Sow. —— cornucopia, Young § Bird. 


—— discoides, Ziet. —— torulosus, Schiidl. 


INFERIOR OOLITE OF DORSET AND SOMERSET. og 


Now as to the forms which up to this time we have found with 
terminations to their shells, we may state that as yet we have only 
met with fourteen; but it happens, as a rule, that the species so dis- 
tinguished number several individuals, though some three or four 
of them have only yielded single specimens. 

Our list, then, includes the following species :— 


Terminations. 

. Ammonites concavus, Sow. 
. -—— subradiatus, Sow. 
—-- Kdouardianus, D’ Orb. 
—— Braikenridgii, Sow. 
—— linguiferus, D’ Orb. 
—— Sauzei, D’ Orb, 

—— Martinsii, D’ Ord. 
-—— subcostatus, Buckman, MS. 
—— Gervillu, Sow. 

—— Brongniarti, Sow. 
—— Manselii, Buchman. 

. --— Humphriesianus, Sow. 
Moorei, Lycett. 

. —— boscensis, Legnés. 


Lanceolate ...... J 


Ovate-lanceolate 
or spathulate. 


semicircular. 


WWieivedes focal. : 


HA Oo NYS oO OO I TB Go BO 


l 
\ 
Delphinulate, | 
| 


Here, then, we have given fourteen species of Ammonites with 
their terminations more or less complete ; and for these we have at- 
tempted to make a provisional classification. 

So far as we have been enabled to examine these more complete 
Specimens of Ammonites, we arrive at the conclusion that, the 
terminations to the shells are capable of greatly aiding in the 
determining of species, as trimmed-up examples of A. linguiferus, 
with their broadly oval or spathulate mouths, are difficult to distin- 
guish from A. Humphriesianus ; but when we have the semicircular 
rugose mouth of the latter for comparison with the former the 
difficulty vanishes. Thus, again, the lanceolate termination of the 
discoid A. subradiatus is very different from the spathulate or ovato- 
lanceolate ending of A. subcostatus (Buckm.); and yet these two forms 
have been referred to one species by different authors. 

The dephinulate mouth in A. Gervilliz is a complicated ter- 
mination when compared with the plainer semicircular terminations 
of A. Brongmarti and A. Humphriesianus ; and yet without this 
knowledge we had always confounded the first two forms. 

From these remarks it seems but reasonable to conclude that the 
terminations of Ammonites may become important in the distinguish- 
ing of species, though, from what we have seen, their preservation is 
of somewhat rare occurrence; at the same time our own experience 
seems to point to the fact that many fine examples of more com- 
plete shells have been lost from the want of due observation upon 
the subject. We may further mention that, with the vast numbers 
of Cephalopada which are everywhere around us and which are so 
perfect in other respects, we have not yet seen a single example either 
of Rhyncholites or Trigonellites. Mr.S. P. Woodward, however, has 
figured an Ammonite with the operculum from Dundry*. The shell 

* See ‘ Geologist, vol. iti. p. 328. 


60 J. BUCKMAN ON AMMONITES FROM THE 

is referred to A. subradiatus ; but it is very different from that which 
is now recognized by this name,—A. Brightii, which it is said to 
resemble, being an Oxford-Clay fossil, while the Dundry Oolite is 
the exact equivalent of the Dorset Cephalopoda-bed. I hope there- 
fore some day to see this specimen in its place at the British Mu- 
seum. 

I now proceed to pen a few notes on these fourteen forms which 
we have observed to present more or less perfect terminations to 
their shells; and in doing so I would express the hope that ere 
long other forms will be met with to enable us more clearly to un- 
derstand the value of a more complete structure, while at the 
same time it may not be too much to express a hope that better 
specimens may yet be found than those we already possess or, 
through the kindness of friends, have had access to. 


Nores ON THE SPECIES. 


Fig. 1.—Termination of Ammonites concavus, Sow. 


Toe S 
WWM 
SA 


1. AmnmonrTEs concavus, Sowerby, pl. 94, fig. 2. (Fig. 1.) 


This is a very common shell in the Cephalopoda-bed in Dorset 
and Somersetshire, occurring less frequently in the same horizon in 
Gloucestershire. It is a very variable shell; and hence its syno- 
nyms are unusually numerous. | 

In this district we have usually named it A. subradiatus; and it is 
here only retained because A. concavus was figured by Sowerby from 
a specimen obtained from this neighourhood. 


INFERIOR OOLITE OF DORSEY AND SOMERSET. GL 


Fig. 2.—Termination of Ammonites subradiatus, Sow. 


SS 


Z) 
Ze) 


2. A. suprapiatus, Sow. pl. 421. fig. 2; D’Orb. pl. 118. fig. 3 (now 
ie 2); also D’Orb. pl. 129. fig. 3.. (Fig. 2.) 


The difference insisted upon seems to be that in A. concavus the 
curved radii are seldom branched, whilst in A. subradiatus, the 
radii are either branched or have short intermediate ones between 
the longer ones. There is, however, so much variation in this 
respect that we still hold these to be mere varieties. 

We have now several complete examples of the termination ; but 
the first specimen was obtained by Captain Kennedy from our own 
quarry so perfect that it might have sat for the portrait given by 
D’Orbigny, pl. 129. fig. 3. It is a very common fossil, and in 
some quarries occurs mixed with the former so thickly that hun- 
dreds of examples could be got ina day ; and yet a specimen with even 
the base of the termination 1s very rare. 


3. A. Epovarpianus, D’Orb. pl. 130. figs. 3-5. 


Is found sparingly with the examples just quoted. Our two 
specimens are the only ones we have meth wit in the Bradford- 
Abbas quarry ; and, curiously enough, they have both indications 
of the termination. 


4, A. Brarkenripert, Sow. Min. Conch. t. 184. (Fig/3.) 


This shell was figured by Sowerby in the second volume of Min. 
Conch., which bears date 1818 ; the specimen, a very imperfect one, 
was from Dundry, near Bristol. In his description he has the fol- 
lowing remarks* :— 

“Perfect terminations of the Ammonites are scarce. I have, 
however, met with several specimens indicating the form of the lip; 
but none of them exhibit much out of the usual way, excepting 
some French ones and those now before us: in one of the French 
specimens the aperture is much contracted by the lip; in another 
the lip forms a single arched lobe slightly bent inwards.” 


* Min. Conch. vol. ii. p. 187. 


62 J. BUCKMAN ON AMMONITES FROM THE 


Fig. 3.—Ternunation of Ammonites Braikenridgii, Sow. 


SY 


yA 


a. Krom the side. 6, From the front. 


Our Cephalopoda-bed is on the same horizon as the bed from 
which the Ammonite just quoted was obtained, Dundry being the 
extreme western extension of the Oolitic system which prevails in 
Dorset and Somerset, which latter has so much resemblance, as evi- 
denced by fossils, to the Inferior Oolite of France. 

This fossil is usually found most abundantly at Clatcombe, near 
Sherborne, where examples with the curious terminations are not 
uncommon; and it illustrates one of those cases in which a different 
species of the genus occurs at one quarry from that which prevails 
at another. It is occasionally found at Bradford Abbas, where the; 
prevailing Ammonites are A. concavus and A. Sowerbyt. 

The terminations in this species are among the most perfect, as 
well as the most frequently met with. The spathulate projection 
varies greatly both as to length and breadth. 


5, A. trincurrerus, D’Orb. pl. 136. figs. 1 & 2. 


Has a neat form and a less broad spathulate termination than the 
preceding, though closing up the aperture of the shell more by 
reason of its bending inwards. It is thinner in the whorls; and 
the ribs are in threes at the back of the shell, uniting into one, with 
a tubercle in front. Sometimes there is an intermediate rib between 
a pair, with three-ribbed vaultings. 

This shell is by no means so frequent as A. Braikenridgu, but 
occurs in the same places, intermixed with it. 


6. A. Savzer, D’Orb. pl. 139. figs. 1-3. 


The termination of this shell is much like the former; but its 
ribs are much larger, and the shell broader. It is also inclined to be 
unsymmetrical, and has but few whorls. 


INFERIOR OOLITE OF DORSET AND SOMERSET. 63 


This is not uncommon with the termination both near Sherborne 
and at Bradford Abbas. 

These three last species are admirably figured by D’Orbigny ; and 
yet we are happy to think that our Dorset examples are quite up 
to the perfection of the French ones. 


fake Martins, D’Orb. pl. 125. figs. 1, 2. 


Ts not uncommon; but the small specimen we have seni is the 
only case in which we have observed the terminal Jip. It seems to 
have been the first lip formed; and if fig. 3 of D’Orbigny’s plate 
represents the lip of an older specimen, it would tend to show that 
the lips vary very much with age. Our specimen is from Bradford, 
where the species is not uncommon. 


gs. A. suscostatus, Buckman, MS. 


This shell, from near Sherborne, belongs to the discoid section of 
Ammonites, and is therefore only placed here on account of its spa- 
thulate termination, which differs so much from that of the shell 
with which it has been confounded, namely A. subradiatus. 

Probably this is the A. subradiatus of D’Orbigny, T. J. pl. 118. 
figs. 1 & 2.; but if so it differs so much from his A. subradiatus 
previously referrred to that we feel justified in proposing a fresh 
name for it. Our specimen is the only example we have met with 
with the termination: still it is not an uncommon fossil near Sher- 
borne, though it occurs very sparingly at Bradford Abbas and Half- 
way House. 


Fig. 4.—Termination of Ammonites Gervillii, Sow. 


9. A. Gervitim, Sow. p. 184, tab. A. fig, 3; D’Orb. pl. 140.. 
figs. 1-7. (Fig. 4.) 
This is a shell with a very curious and interesting termination, 
which we have named delphinulate, because its side view 1s so much 
like that of the head of the classic Dolphin. Sowerby says, ‘‘ This 


64 J, BUCKMAN ON AMMONITES FROM THE 


species occurs in a marly limestone which is replete with grains of 
iron ochre. It is from Bayeux, in Normandy ” *. 


Fig. 5.—Ternunation of Ammonites Brongniarti, Sow. 


= yy 


10. A: Bronentartr, Sow, t. 84 A, ‘ie. 2: DiOrcbampiaailane 
figs. 1-5. (Hig. 5.) 

The lip of this shell is so decidedly distinct from the preceding that 
we cannot help separating them, although we at one time inclined 
to a different opinion. It is not at all so complicated, but is of a 
semilunar or, rather, semicircular shape, with a deep depression be- 
neath, but without the elaborate form of the lip of A. Gervilla. It 
is met with at Bradford Abbas and other places near Sherborne. 


11. A. Mawserit, Buckman, n. sp. 


Is perhaps related to the A.-Brocchi group ; but the fineness of its 
ribs and the absence of tubercles is a sufficient distinction. We 
have in one single example the usual deep depression before the 
terminal semicircular depression. We possess several examples of 
this shell from Bradford, Chaleombe, and other places ; but only the 
one sent is an example of the termination. 


12. A. Humenrimstanus, Sow. t. 500. fig. 1; D’Orb. pl. 133 & 134. 
(Fig. 6.) | | 

This is a very variable shell, sometimes having few (perhaps five) 
thick whorls, at others from eight to ten, exceedingly slender and 
delicate; yet the markings are the same, and the termination of both, 
starting from a deeply contracted furrow, is simply semilunar in 
shape, and is the same in both large and small examples. 

This is a common shell about Sherborne, but is rarely met with 
at Bradford or Halfway House. 


* Sowerby, Min. Conch. vol. u. p. 190, 


INFERIOR OOLITE OF DORSET AND SOMERSET. 65 


Fig. 6.—TZermination of Ammonites Humphriesianus, Sow. 


SS 


a. Krom the side. 6. From the front. 


13. A. Mooret, Lycett, ‘The Cotteswold Hills,’ pl. i. figs. 2, 7. 


The flexure of the mouth of the shell, with its elegantly curved 
line, like that of a modern Nautilus, is distinctive of this shell. It 
is common to the sandy part of our Oolite below the usual Cepha- 
lopoda-bed. The mouth is simply a termination of a flat slightly 
ribbed shell, without any previous depression. It is a common 
shell in the limestone bands below the Cephalopoda-bed ; but the 


specimen sent is the only one in which we have observed the com- 
pletion of the mouth of the shell. 


Fig. 7.—Termination of Ammonites Moorei, Lycett. 
e 
We. 


14. A. Boscrnsts(?), Reynés. 


We quote this, with some degree of hesitation, from Beneke, inas- 
Q.J.G.S. No. 145. F 


66 AMMONITES FROM THE INFERIOR OOLITE. 


much as we have only a single specimen. This, however, is from 
the Cephalopoda-bed of Bradford Abbas, and in some points it is so 
much like the A. Moorei, both in its form and the outline of the 
mouth, that even now we must confess to fecling great difficulty in 
the matter. 

Note.—Since the above was in type, we have found other species 
and examples with terminations. ‘These, with the new forms, will 
described in a paper now preparing for the Society : 


_ Discussion. 


Mr. Cuartesworta called attention to the early work in this 
direction of the late Mr. Chaning Pearse, who studied the Ammo- 
nites of the Oxford Clay of Christian Malford. That author was 
led to believe that the peculiar prolongations of the mouth were 
periodically absorbed and reproduced. 


ABNORMAL GEOLOGICAL DEPOSITS IN THE BRISTOL DISTRICT. 67 


7. On AxsnormMat GrotocicaL Deposits tn the Bristot District. 
By Crartes Moors, Esq., F.G.8. (Read November 17, 1880.) 


In making excavations at Redland, on the edge of Durdham Down, 
about 45 years ago, some conglomerates, associated with Carboni- 
ferous Limestone, were opened up, which contained the teeth and 
scattered and broken bones of reptilia, described by Messrs. Riley 
and Stuchbury in the ‘ Proceedings of the Geological Society’ for 
1836 under the names of Thecodontosaurus and Palcosaurus. As 
they were then the oldest known reptilia and of a high order, much 
interest has always attached tothem. <A few years since, in drainage- 
works at the same spot, this conglomerate was again crossed, and 
some other bones added to the series deposited in the Museum of the 
Bristol Philosophical Society. The collection has since been reviewed 
and described by Professor Huxley, F.R.S.*, and an account of 
the physical characters of the district given by Mr. Etheridge, 
LAI NES By g ; 

Much uncertainty has prevailed as to the geological age of the 
Durdham-Down conglomerate. At first it was supposed to be Per- 
mian; but we have as yet no conclusive evidence of true Permian 
beds in the West of England. Mr. Etheridge has placed it on the 
horizon of the Dolomitic Conglomerate of the Keuper ; whilst, owing 
- to the discovery by myself of the same genera of reptilia, under 
somewhat similar physical conditions, in the Rhetic deposits of 
Holwell, and since then of true Rhetic remains on Clifton Down, I 
had referred them to the latter age—a point to be reviewed below. 

In my paper on the abnormal conditions presented in the Frome 
district I especially described numerous Secondary unconformable 
deposits and vein-fissures resting upon or passing down through 
the Carboniferous Limestone, some of them haying organic remains 
whereby they could be referred to different geological periods. 
Since then I have discovered several other features of interest at 
Holwell; and as the phenomena of this district will serve as a 
key to unlock hitherto unnoticed physical conditions in the geo- 
logy of the Bristol area and some of the paleontology connected 
therewith, it will be desirable first shortly to notice the special fea- 
tures presented near Frome. 

The Carboniferous Limestone has here its last south-eastern ex- 
posure before being entirely covered up by Secondary deposits, and 
is to be seen in very narrow ravines at the Vallis, Him, Mells, What- 
ley, Nunney, and Holwell. The prettiest combe is that of the Vallis, 
from which, at its northern end, bifurcates that which passes 
to Elm and Mells. At the entrance to the Vallis at Hapsford the 
first sections show irregularly bedded Rhetic conglomerates resting 

* “On the Classification of the Dinosauria, with observations on the Dino- 
sauria of the Trias,” Quart. Journ. Geol. Soc. vol. xxvi. (1870) p. 32. 

tT “On the Geological Position and Geographical Distribution of the Rep- 


tilian or Dolomitic Conglomerate of the Bristol Area,” Quart. Journ. Geol. 
Soe. vol. xxvi. (1870) p. 174. 
F2 


68 CG. MOORE ON ABNORMAL GEOLOGICAL 


in depressions on the edges of the inclined Carboniferous Limestone. 
They are separated by thin blue clays with Avicula contorta and 
also Discona Babiana. I have no doubt they would yield important 
vertebrate remains, as I found a very perfect Dinosaurian vertebra ; 
but, unfortunately, these beds have not been worked for some years. 
Vertical mineralized infillings are present, passing down through the 
Carboniferous Limestone in this section. In the southern corner, but 
occupying a higher position than these conglomerates, there is a 
patch of close-grained cream-coloured limestone undistinguishable 
from the White Lias; but it has yielded me no trace of organic re- 
mains. The deposits of one section usually differ from those in 
another, or die out altogether, although they may be in close proxi- 
mity. For instance, in the next quarry the conglomerate is a single 
thick, dense bed, with pebbles united by calcareous matrix, in which 
are occasional nests of Lstheria minuta and insects, whilst above are 
a few inches representing the Lower Lias, capped by about 10 feet of 
Inferior Oolite. 

In addition to the unconformable Secondary deposits which lie on 
the Carboniferous Limestone, a special feature of the latter formation 
is the many vertical fissures passing down through it, with infillings 
containing Secondary remains, or those of a still later age. In the 
Vallis these veins are best seen in the face of a large quarry at Ege- 
ford, where, in addition to several smaller ones, the workings are 
bounded by veins at both ends, that on the north being of some 
thickness, and showing a vertical side about 50 feet in height, where 
it meets a patch of Inferior Oolite. The matrix of this vein is Liassic ; 
and numerous organisms may be traced init. All the veins are much 
mineralized, and contain sulphate of barytes in concretionary layers, 
with occasional traces of galena and blende. At Elm and at Nunney 
hematite iron-ore has been extracted ; at the latter place in close 
proximity to a deposit of Inferior Oolite. 

The hamlet of Holwell occupies a depression at the southern end 
of the Vallis gorge, and is on every side surrounded by sections 
which, if looked down upon from a little distance, without closer 
inspection, would appear to most geologists to be entirely composed 
of Carboniferous Limestone. The road from Frome to Shepton 
Mallet passes through this spot, and crosses the little stream from 
which the hamlet is named. On the eastern side are two quarries I 
have already described*, viz. :—that of the Marston road, in which, 
within a few yards, are present stratified Inferior Oolite, Liassie con- 
glomerate (with many organisms), and a thin band of Rheetic clay, all 
resting on Carboniferous Limestone, with a thick vein filled with calc- 
spar in their midst ; and also a large quarry below, with a face pa- 
rallel to the stream, in which are to be seen many vertical dykes, 
some with Rhetic and others with Liassic organisms ~. They present 
much mineralogical variety ; and the somewhat impure limestone 


* « Abnormal Conditions” &c., Quart. Journ. Geol. Soc. vol. xxiii. (1867) 
p. 483. 

+ A section of this has been given in ‘‘ Abnormal Conditions” &c., Quart. 
Journ. Geol. Soc., 1867, p. 484. 


DEPOSITS IN THE BRISTOL DISTRICT. 69 


within the veins shows, by its frequently being in very thin lamine, 
that it has been deposited slowly. It is desirable to bear in mind, 
whilst comparing the same facts in the Bristol district, that in the 
quarries above noticed the limestone of these Secondary infillings is 
often left standing out as butresses by the workmen, whilst they 
work back between them for the purer Carboniferous beds. 

Postpliocene, Liassic, and Rhetic Deposits. The Microlestes- 
Quarry.—On the Shepton road, immediately west of the hamlet, 
there is a large quarry on each side facing the road. The first, 
on the south, is the Microlestes-quarry; in the other, inclined 
Carboniferous Limestone has been worked between thick veins of 
Middle Lias, the matrix of which occasionally showed carbonate of 
lead and barytes. The rubbish-tippings from the limekiln have 
since obliterated these points; but I have preserved them in pho- 
tographs. It is probable that all the veins on this side of the 
hamlet differ from and are to the south of those in the large 
section east of the river. 

In my former examinations of the Mcrolestes-quarry I was told 
by the workmen that they had occasionally crossed a fissure down 
which, by the aid of a rope, a man might descend; but, as it was 
becoming dangerous, it was filled up. From this I have suspected 
the presence of a Postpliocene cavern at the spot; but although it 
has not been found, they have worked back upon a vein filled with 
bones, which I have lttle doubt may be connected with one. It is 
about a foot in thickness, and, like all the other fissures and their 
infillings, appears to have its own special individuality. Like an 
ordinary mineral vein, the walls have vertical mineralized layers, the 
innermost being large crystals of dog’s-tooth spar. Towards the top 
there are anguiar pieces of Carboniferous Limestone, which below 
sive place to material composed of about one third of brown marl, 
crystals of carbonate of lime, and dismembered jaws and bones of 
Arvicola and frogs, with, rarely, small vertebree of birds and fish. 
The jaws of Arvicola are very numerous, whilst their vertebree and 
other bones are scarce. They appear to belong to A. arvalis or A. 
saxatilis. Bones of the larger mammalia, though few, including ox 
and deer, were found. Amongst them was found a tooth not to be 
distinguished by itself from a human incisor; but the subsequent 
discovery of the remains of wolf leads to the inference that it may 
belong to that animal, the difference in the incisors being with diffi- 
culty distinguished. Several Rhetic teeth, showing from their worn 
condition that they are derived, and a few Carboniferous-Limestone 
corals and shells are mixed with the above, as ; well as pebbles of 
hematite and bog-iron ore. 

The deposit, with its Rhetic remains, was in a north-and-south 
fissure a little west of the above; but it is now all but exhausted. 
_ Without reference to other mineralized veins in which no organisms 
have been found, it will be seen that at Holwell alone, and in the 
line of a single quarry of but a few hundred feet length, Post- 
pliocene, Liassic, Rheetic, and Carboniferous-Limestone formations are 
represented. ‘These later deposits are the rule, not only where they 


C. MOORE ON ABNORMAL GEOLOGICAL 


70 


~>, 


‘sdagg Axren% 


LORS 


*‘q1g0dep 
uel ado 


“OnAdTIOM 


‘umog moypung fo abpy ay, yo worn 


ay 


—"T “OL 


“£urend 


DEPOSITS IN THE BRISTOL DISTRICT. Tt 


fringe the outcrop of the clder rocks, but high up on their table- 
lands, as in the case of the Charterhouse lead-mine, near 


Cheddar. 
Tue BristoLt AREA. 


Before proceeding with a description of the Bristol area and 
comparing it with the above, it is desirable I should advert to a new 
palzontological feature which hitherto has not attracted any atten- 
tion, in the presence of a multitude of minute Serpula-like calcareous 
tubes found in various deposits under examination. Years ago I 
noticed them in the freshwater brick-earth of Salisbury, which is of 
Postpliocene age, and put them aside as minute Serpule; but after- 
wards learning that all species of this family are marine, I thought 
they might possibly be analogous to caddis-cases, and belong to some 
insect. When they were afterwards found in enormous numbers, 
and under many diverse circumstances, I saw the desirability of 
learning more about them, and sent them to friends who were 
authorities in special departments of natural history for their 
opinions, at the same time describing the circumstances under which 
they were found. Without mentioning names, it proved a “ pursuit 
of knowledge under difficulties.” First, they were doubtfully re- 
ferred to the cases of some Dipterous insect ; and if not such, it would 
be worth considering if they were Serpule. An entomologist decided 
that they were not insect-tubes; a good microscopist was of opinion 
they were the calcareous cases left by rootlets of vegetation passing 
down from the surface; next, an eminent zoologist intimated, with 
some uncertainty, that they somewhat resembled the genus of Ser- 
pulide, Filograna; then a good botanical friend and microscopist 
pronounced them to be vegetable; upon which I sent them to a 
first-class botanical expert, who was of opinion they had nothing 
to do with vegetables, and that he had consulted a zoological friend, 
who also believed they were worm-tubes allied to Annelids, and 
that again they presented some resemblance to /ilograna. 

They are to be found very abundantly in some of the vein- 
fissures of the Carboniferous Limestone (to be referred to hereafter) 
in the Bristol district, and also, though not in such numbers, in 
some of the later stratified marls. My impression is that they 
are due to freshwater conditions, and that, though they may not 
be rootlets which have passed down from the surface (some of 
them being found under conditions apparently precluding this idea), 
yet they may, notwithstanding, be due to freshwater vegetation. 
They are sometimes free, but often in clusters united by the ma- 
trix of the deposit. It will be desirable to determine their syste- 
matic position, as they will have a bearing on the age and other 
conditions of the deposits in which they are found. I propose 
that they should be recognized under the designation of Tubutella 
ambigua. 

Durdham and Clifton Downs.—I now propose to show that the 
same physical conditions prevail at Durdham Down and at various 
outlying spots near Bristol, as in the Frome district. The area to 


72 C. MOORE ON ABNORMAL GEOLOGICAL 


_be first noticed will be that embraced in the southern escarpment of 
the Carboniferous Limestone, extending from Redland, its eastern 
end at Durdham Down, to the suspension-bridge at Clifton, over- 
looking the river Avon on the west. 

Near the Old Black Boy inn, just under the edge of Durdham 
Down, there are places known as The Quarry and The Quarry 
Steps (see fig. 1). comming on the latter, we look down into a large 
excavation, which musi have been formerly extensively worked for 
stone, but is now occupied by small houses and gardens, most of which 
are probably a century old. One of these houses is close under the 
flight of steps leading to The Quarry, and has been built against a 
natural vertical wall of mottled red or yellow unstratified limestone, 
differing altogether in colour and texture from the grey Carboniferous 
Limestone of the district, by which, on either side, itis bounded. It 
is about 8 feet in thickness, and may be traced on the east side of 
the excavation by its brighter colouring; and there is no doubt that 
it continues into the limestone of Durdham Down immediately above. 
As was the case with the workings on the Mendips, the old quarry- 
- men here extracted the purer limestone, leaving the impurer some- 
what conglomeratic infillings standing out, and terminated their 
quarry on the east by this large dyke. As it has not been opened 
up, little examination could be given to it for organic remains, which, 
from its close proximity to the Thecodont deposit, would have been 
desirable; but that some are present therein is sufficiently indicated 
by the fact that, after examining a few pounds weight of the softer 
material taken from the interstices or sides of the vein, I obtained 
numerous minute fragments of bone or teeth, one very small fish- 
tooth, an Hchinus-spine, a few joints of Carboniferous-limestone 
Encrinites, and some of the tubes previously referred to. 

The platform of the Quarry Steps rests upon the surface of the 
above dyke. Looking from it, along the Down escarpment to the 
west, the eye takes in Bellevue Terrace, on the edge of the Down ; 
and it was between these houses and the quarry, a distance probably 
of 200 yards, along the same face of limestone and on the same 
horizon, that the deposit containing the Thecodontosaurian remains 
was found. Unfortunately the precise spot is unknown; and, from 
its being built over, there is not much hope of its being again 
identified. 

My late friend, Mr. W. Sanders, F.R.S., of Clifton, gave me, some 
years ago, a sketch showing the conglomerate on the edge of the lime- 
stone, with what was then considered to be New Red Sandstone at its 
base (fig. 2); and it is significant that he does not so much represent 
it as a basin-shaped depression in the limestone as indicate a deposit 
following the slope of the escarpment, similar to the case of a vein 
the top of which was opened up, but from which the limestone still 
resting below against its side had not been removed. Mr. Sanders 
also marks the spot where the reptilian bones were supposed to be 
found. . 

At the time when these reptilia were discovered, the peculiar 
conditions of deposition I have indicated were unknown. In the 


DEPOSITS IN THE BRISTOL DISTRICT. 73 


Fig. 2.—Sketch Section of the Thecodontosaurus-bed of Durdham 
Down. (Drawn by W. Sanders, Esq., F.R.S.) 


Cis Pretest BORE SEAC 
< ZING ‘ Z 
NY iN ENG ZG 
RS yy 4, Y, es OX 
COw: aN Oe & G 
“nN NEE N UAW <4 SO OX 


Sf 


ON, 


a. Conglomerate of New Red Sandstone. 
b. Locality of Thecodontosaurus. c. New Red Sandstone. 
d. Carboniferous Limestone. 


section described below, which embraces a line of working from the 
Quarry Steps to a large quarry which I call the ‘‘ Avenue Quarry ” 
(from its being immediately north of the Avenue Road), a distance 
all together of about 680 yards, I shall show the presence, as at Hol- 
well and elsewhere, of a series of veins with infillings derived from 
different geological ages; and my interpretation of the reptilian 
deposit therefore is that it 1s one of such a series in Carboniferous 
Limestone, by which, on both sides, it is surrounded. 

Durdham Down Section.—From the Quarry Steps to the end of 
Bellevue Terrace, within which area the reptilia were found, is about 
300 yards. Until lately these were the last houses fringing the Down 
to the west; and there then stood up by itself, like a wall, a large 
vein-infilling (the limestone having been worked up to it on both 
sides) separating the garden of the last house from an adjoining 
quarry. Other buildings have now occupied the line of the quarry, 
covering up four smaller veins; but the one above referred to, from 
14 to 16 feet in thickness, is still utilized as a boundary between the 
gardens. It has the same general hthological character as that at 
Quarry Steps, but has occasionally small pockets of iron-ore, and is 
more mineralized. I purposed to examine these veins closely for 
organisms, but have been prevented. 

Passing along the roadway from this quarry, the next vein seen 
was one containing hematite iron-ore, at its thickest about 16 feet. 
It had been worked up to the roadway on the edge of the Down. 
At my last visit this also was being filled up, and a house built 
across it. 

Alluvial Veins with Inassic and Rhetic Remains.—For about 
200 yards from the iron-ore vein the limestone has been unworked. 
Following the road to a spot near the ventilating-shaft of the 
Avonmouth Railway, an archway facing the road leads into the 
Avenue Quarry, which presents some interesting features. For 
many years a lump of seemingly stratified yellow marl had been left 
in the bottom of the quarry, the limestone having been worked 


74 C. MOORE ON ABNORMAL GEOLOGICAL 


around it. Its presence often puzzled me; but there is little doubt 
that it is due to an alluvial infillmg into a small limestone cavern, 
and is connected with one of three thin veins of alluvial material 
which are present in this quarry. A cavern was many years ago 
discovered on the Down, containing, amongst other things, the 
remains of Hippopotamus, which are now in the Bristol] Museum. 
These three veins are filled with an ochreous or brown clay, which, 
when critically examined in its natural condition, appears to differ 
slightly in character in each, as though they might have received 
their infilling at somewhat different times ; still, to save labour in the 
examination of their contents, they were mixed together. Their 
organic remains are very varied, and not less so their mineral con- 
stituents. Their Postpliocene or still later age is indicated by the 
presence in them of frequent portions of the incisor teeth of Arvicola, 
and a single shell of Hew. Although all the remains are rare, those 
from the Lower Lias are most abundant. These consist of young 
forms of Ammonites of two species, Myacites and Astarte, Cylindrites 
and portions of three other univalves, a Pentacrinite-stem, and a 
single valve of an Entomostracon. ‘There are teeth and scales of 
fishes, and a fragment of bone which may be either Liassic or Rheetic. 
The interesting presence of Rhetic remains, however, is shown by 
teeth of Saurichthys and Lophodus and a small palate identical with 
species from Holwell. Some of the teeth are bleached and worn by 
water-erosion. The little calcareous tubes accompany them. 

The residue of these veins, after washing, shows a great abund- 
ance of black pisolitic granules. The following analysis of its 
mineral contents has been obligingly made by Mr. Gatehouse, City 
Analyst of Bath :— 


Soluble in acids: 


Watery Mos) ioc ts Sisieoe Gade a Bee a 7°80 
Calciumycarbonmate tame ar ae eee 20°50 
Magnesium carbonate...... ee eB os 0°73 
Oxide ‘Of Tromso.) eee os sil eens 6:50 
Zime carbonate (calamine) 2-5. 02) 24.45 0°54 
Manganese, dioxaideds saucy. a)-Nelcie cna 0-50 
ead=sulphide\(eallena)) ye. eee 0°90 
ho) Det Mra AUR eM NAME ie Bestia i ny'ss 6 8°16 
PeMADNACUNGl: PREM ML HARE OIRO 6 oa.3 2°60 
Pobassivamn'y 6,552.05 eng sieeucie ds oe ieee ne 0:07 

. 48°30 

Insoluble in acids: 

Calcium sulphate............ Pale ee 0-39 
SUHICALC Of, ZINC Cerrar. cele n anand eae 0-25 
Oxadeyor ron Gnsoluble) ie; ariel 6°55 
Alumina Pe ROMA NHRC MRO Wind suis ics 10°45 
Silica RS UIM Rn NENTS Mae fee ka hes 33°70 
WOPDCE MN ce cas walneal ele emete trace: 


DEPOSITS IN THE BRISTOL DISTRICT, 75 


Inassic Vein.—The working of the limestone in this quarry has ter- 
minated on the western side against a vein from 10 to 12 feet thick, 
which has received an infilling of yellow clay of an entirely different 
lithological character from those last mentioned. So far as the evi- 
“dence goes, it is apparently a vein of true Lower Lias, with occa- 
sional weathered blocks of Carboniferous Limestone. All the remains 
found are Liassic, and consist of :— 


Teeth and scales of fishes. Gryphea. 
Ammonites (young). Terebratula. 
Univalves (several species). Cidaris (teeth, plates, and spines). 


Astarte. Pentacrinites. 
Modiola. Cristellaria. 
Lima. Webbina. 
Cardium 


All the phenomena I have mentioned, from the Quarry Steps to 
the Avenue Quarry, are embraced in less than half a mile. Without 
including the Reptilian deposit, eleven veins are shown to be pre- 
‘sent; and but for a considerable part of the edge of the Down being 
covered, no doubt many others would have been seen. 

The thetic Bone-bed—There are no workings between the 
Avenue Quarry and the neighbourhood of the Clifton suspension- 
bridge. ‘The road to the latter is cut through Carboniferous Lime- 
stone containing many veins, which are usually filled with cale-spar 
and other mineralized matter. Looking towards the Observatory, 
there are two veins. One, filled with limestone, has been left boldly 
standing up by the quarrymen; but nothing can be said of its age. 
Close to the toll-house, however, on the Clifton side, there is a 
deposit of considerable interest, having a face of about forty feet, in 
which the Rhetic bone-bed and its associated remains are present. 
It is partly composed of irony and yellow sandy-looking marl, 
with many free crystals of carbonate of lime, as in the Holwell 
fossiliferous infilling ; and there are also patches of finely laminated 
rock, similar to the Rhetic “ White Lias.” Some of the associated 
blocks of stone appear to be fossiliferous ; but as the deposit forms the 
boundary-wall of the toll-house, they cannot well be broken down 
for examination. The bone-bed is two inches thick, with teeth of 
Saurichthys apicalis, Lophodus minimus, and many fish-scales; and 
the clay on either side contains fish-remains of the same age. 

In Professor Ramsay’s ‘ Physical Geology and Geography of 
Great Britain, he has given a very pretty sketch of the Gorge of the 
Avon, with the Suspension-bridge and its beautiful scenery, which, 
through the kindness of Prof. Ramsay and of Mr. Stanford, the pub- 
lisher, I am permitted to use (fig. 3). Curiously enough, it includes 
the Rheetic bone-bed, and, on the shoulder of limestone looking down 
the river on the east, the conglomerate mentioned below. The Car- 
boniferous Limestone dips rapidly to the south under the Bedminster 
coal-beds, and disappears under beds of New Red Sandstone. A 
lighter line in the sketch, between the eastern abutment of the bridge 
and the houses on the level of the roadway, with the Observatory 
seen beyond it in the distance, shows the exact position of the Rheetic 
deposit. 


76 C. MOORE ON ABNORMAL GEOLOGICAL 


Fig. 3.—Gorge of the Avon at Clifton. 


Before referring to other outlying sections near Bristol, it may 
be noticed that although the true New Red Marls are present 
around, the red and variegated marls which lie on the east of the 
basin between the Durdham-Down and Clifton escarpment, and 
extend towards Bristol, are alluvial. They have been opened up in 
brick-pits repeatedly, as well as the conglomerates below. From 
the former, after long search, I obtained several small bits of Inferior 
Oolite and fragments of shells of Ostrea. The conglomerates are 
bedded, and are usually extracted on the site of each building 
erected; and although hundreds of thousands of tons have been so 
used, I have never detected any contemporaneous organism in 
them, nor have I heard of such having been found. They get coarser 
and more irony as they approach the limestone below; and in 
several sections its rugged surface has received pockets of hematite 
iron-ore. The pebbles are comparatively small and angular, and 
give the idea that they may be due to a subaerial denudation of the 
Carboniferous tableland above. 

Conglomerate of the Avon Gorge.—Passing down the river, not far 
below the Suspension-bridge, the ‘ Dolomitic Conglomerate,” of 
which Mr. Etheridge has given a section, is reached. It rests on 
the edge of the Carboniferous Limestone at the side of the road 
leading up to the Down, and is, as a friend has remarked, ‘a 
gigantic heap of conglomerate” with pebbles of great size at the 
base, getting gradually smaller at the top. It is very irony, and 
appears to occupy but a small area. Some red marls gathered from 


DEPOSITS IN THE BRISTOL DISTRICT. 77 


the sides and interstices of the deposit yielded me part of a fish- 
palate, which is probably of Carboniferous-limestone age, and great 
numbers of Encrinital stems, a comparatively recent freshwater 
operculum, and numerous examples of Tubutella. Whatever may be 
the age of this deposit, groups of the latter are usually surrounded 
by an irony matrix which seems part thereof. 

Vein at Ashton.—On the west side of the Avon, about a mile 
beyond Ashton, considerable deposits of iron-ore occur ; but I have 
been unable to examine them. They lie at the foot of the lime- 
stone escarpment. On the tableland above them, at Longwood 
Farm there are disturbed Carboniferous Limestones, in which a 
vein occurs containing occasional lumps of galena and some ¢a- 
lamine. Tubutella is present in great abundance. The upper 
part of this vein is somewhat honeycombed; and the tubes are 
found adhering to thin flakes of calc-spar. No other organisms 
were found at this spot. 

Westbury-on-Trym— Carboniferous Limestone with Minerals and 
Oolitic Remains.—On passing from the Durdham-Down quarries 
(previously mentioned) along the eastern edge of the Carboniferous 
Limestone, at a distance of two miles, some large quarries are to be 
found. On the eastern side of one at Southmead, worked by Mr. 
Kennedy, there may be seen what appears to be an ordinary mineral 
vein, about a foot thick, passing down through the section. It con- 
tains good hematite iron-ore, ochre, galena, and calamine. Although 
it appeared nothing but a mass of mineral matter, I still hoped a 
sample might yield some evidence of its age. In this I succeeded 
beyond my expectation; for on washing it I at once found many 
angular pieces of Inferior Oolite, which, from their being stained 
with iron, were not before visible. On a still closer examination 
I obtained the oolitic organisms given in the list below, asso- 
ciated with TZwubutella; the specimens show very little sign of 
attrition. No Oolitic deposit from which they could be derived, 
however, is in this district nearer than Dundry, six miles to the 
south, or the Cotteswolds, many miles to the east, with, in the latter 
case, the area of the Bristol coal-field intervening. 

In the same quarry there is at another spot a pocket of light- 
green clay. Although very intractable, I was able to disintegrate 
it sufficiently to find that it contained Alge &c.; and although 
there remains a little doubt as to their age, it is hkely they are of 
comparatively recent date. 


Westbury vem, Oolitic Organisms. 


Fish-tooth. Thecidium. 

Nerina. Bryozoa, several species. 
Solarium. Echini, teeth, plates, and spines. 
Univalve, sp. Entomostraca. 

Turbo. Serpule. 

Dentalium. Foraminifera ? 

Astarte, Tubutella, 

Cardium. Pentacrinite joints. 

Ostrea. Encrinite stems, Carboniferous. 


Algeze (in clay vein), several genera 


78 C. MOORE ON ABNORMAL GEOLOGICAL 


The sections remaining to be noticed are along the eastern edge 
and to the north of the Bristol coal-field. On the eastern side the 
Carboniferous Limestone does not come to the surface between the 
Mendips and a very small outcrop at Grammar Rock, under Lans- 
downe, near Bath (which is not recorded on the Ordnance Map), and 
others at Wick Rocks and Codrington. From Chipping-Sodbury 
a narrow belt continues to Yate snd Wickwar, and entirely sur- 
rounds the north of the coal-basin. 

At Wick, where the quarries are extensively worked, mineral 
veins are to be seen having their usual vertical bands of barytes, 
galena, &c. passing down them. In their softer pockets Tubutella 
is abundant; in one which has a marly infilling it is present in 
great numbers. Occasionally small patches or pockets of grey marl 
lie near the surface; but I have not yet found any other organisms 
in them. 

The Yate Rock Sections—These are between Chipping Sodbury 
and Wickwar. On the ragged surfaces of the limestone there are 
here also pockets containing more mineralogical materials with 
Tubutella. A vein passes down from the surface of one of the 
quarries a foot in thickness, containing soft mineralized material in 
which are myriads of these little tubes; they are often attached to 
the broken-up pieces of barytes and other minerals. 

Netilebury Quarry, Yate, and Clevedon.—The Nettlebury quarry 
is a large one, nearest Wickwar, and is the last I shall refer to on 
this side the Bristol coal-basin. A section of it has been given by 
Mr. Etheridge *, in which are shown to the east horizontal step-like 
beds, overlying highly inclined Carboniferous Limestone, whilst 
on the west side equivalent beds dip at the same angle as the 
limestone towards the coal-basin. At the present time it would be 
difficult to recognize the locality from the above section, from the 
almost entire absence of the beds on the eastern side, though 
they are present from 12 to 14 feet thick on the west. Mr. 
Etheridge considers these beds to be of the age of the Dolomitic 
Conglomerates, and the representatives in lithological condition and 
age of the supposed Magnesian Limestones of Clevedon. The latter 
are thick, irregularly bedded, yellow limestones, used in the district 
for building-purposes. It is apparently a local deposit, resting in 
the quarry near the hotel on Old Red Sandstone, which crops out on 
the beach and abuts against Carboniferous Limestone to the east, the 
Old Red being fringed on a level with the Severn by a continuous 
belt of Dolomitic Conglomerate, continuing north for some miles. . 
Almost every parting or crack between the blocks of Clevedon 
stone shows the presence of galena, carbonate of copper, or other 
minerals. In only one instance have I found a block of stone with 
organic remains. This contained afew Encrinital stems, an im- 
perfect Rhynchonella, and also several imperfect Strophomene. Were 
it not that the specimens put on the lithological appearance of the 
enclosing matrix, I should be disposed to think them redeposited 


* Quart. Journ. Geol. Soe. vol. xxvi. (1870), p. 179. 


DEPOSITS IN THE BRISTOL DISTRICT, 79 


from the Carboniferous Limestone *.” Looking at the Yate deposit as 
it lies on the limestone in the section, it has much resemblance to the 
Clevedon beds, but on closer examination this ceases. The latter 
are almost pure Magnesian Limestone, whilst the upper 9 feet of 
the former are of sandy yellow marls, which on being washed float 
away and leave a sandy residue. Next follow eighteen inches of 
yellow limestone, in two irregular beds, which in structure and 
colour somewhat resemble the Clevedon stone. On treatment with 
acid these also leave a sandy residue. Between the above and the 
Carboniferous beds there is a thin deposit in pockets, almost com- 
posed of fine grey sandstone. There are no organic remains special 
to these beds beyond fucoids and Tubutella ambiqua, which is rare. 
For these reasons, and knowing how varied are the deposits on the 
outcrops of the older limestones, I am disposed to think that the 
yellow beds at Yate are of comparatively recent age. On the 
surface of the Carboniferous Limestone there are occasional thin 
patches of what appears to be a comminuted shelly breccia with a 
grey sandy matrix, which I do not think the equivalent of the 
bouldered Dolomitic Conglomerates found in other parts of the 
district. 

Thornbury Railway and Secondary Veins—The branch line which 
leads from the Yate station to Thornbury exhibits some interesting 
geology. At Tytherington fine sections of Carboniferous Limestone 
are seen, and thick deposits follow of what are, no doubt, true 
Dolomitic Conglomerates. On emerging from the tunnel towards 
Thornbury these have some marly divisions, in which, in a flat over 
the tunnel, some galena has been found. Not far beyond, the Lower 
Limestone shales pass into the Upper Devonian beds, the conglo- 
merates also resting upon them. Just before reaching this point 
there are several thin veins in the limestone containing sulphate of 
barytes and galena, in a matrix of gossany iron-ore. A sample 
from the soft ochreous part yielded me three Conodonts of Carboni- 
ferous-limestone age—and of Secondary remains, Pentacrinite joints 
and a single specimen of a Foraminiferous shell, Planularia pau- 
perata. On a second visit I discovered deposits towards the surface 
which, lithologically, I cannot distinguish from a ferruginous marl 
of the Middle Lias, which contains hollow casts of shells and crushed 
specimens of what appear to be Rhynchonella tetrahedra. Tubuiella 
is present in great numbers, surrounded, as at Yate Rocks, by a re- 
deposited ferruginous matrix. 

Age of the Deposits—From the foregoing examples of abnormal 
deposition, all of which have been accidentally revealed by quarrying- 


* Since writing the above, I have sent the specimens to my friend Mr. 
Davidson, who says:—‘‘ The specimen you send for my examination you say is 
from the Magnesian limestone. In colour it looks like a rock of that for- 
mation ; but I have never hitherto seen from our British Permian rocks a Sfro- 
phomena or Streptorhynchus shell similar to the one I observe on both sides of the 
specimen, and which looks like Strophomena crenistria. 'The Rhynchonella is 
not sufficiently complete for specific determination. If not Carboniferous, at 
least one of the species would be new to our Permian rocks or Magnesian Lime- 
stone. I almost fear your enclosed specimen is Carboniferous.” 


80 C. MOORE ON ABNORMAL GEOLOGICAL 


operations in the Carboniferous Limestone, it must be manifest that 
there are a multitude of other examples not yet opened up that 
would yield an interesting study to the geologist. One of the most 
difficult problems regarding some of them is to arrive at satisfactory 
conclusions as to their exact age; but there can be little doubt 
that the physical conditions to which the deposits are due were the 
same both in the Bristol and the Mendip areas. Supposing the 
fissures in any district had all been caused by the same shrinking or 
change of level, they would have been subject to the same refilling 
influences, and would contemporaneously have received a mixture 
of materials derived from the denudation of that time; but although 
the alluvial infillmgs in the Avenue Quarry have a mixture of 
organisms, it is a singular fact that in a series of parallel veins 
coming to the surface on the same horizon, not far removed from 
one another, and some of them but a few inches in thickness, each 
appears to have an individuality of its own, and to represent in 
geological time intervals clearly distinct from one another. As at 
Holwell, so at Durdham Down, the worked face of the escarpments 
reveals infillings of allnvium, Oolite, Lias, and Rheetic and Keuper 
beds, whilst mineralized or iron-ore veins show conditions specially 
their own. 

Reference has repeatedly been made to the Tubutella ambiqua, 
which I have found in almost every deposit that could be examined 
under favourable conditions, from Maidenhead to those of Gloucester- 
shire and Somersetshire. When they occur, as in the brick-earths 
of Salisbury, in association with freshwater shells, and also with 
Postpliocene mammalia, there seems little reason to doubt that 
they belong to freshwater deposits. If they can be traced in older 
formations, they may be a guide in determining the conditions under 
which those formations have been laid down. I have reason to 
suppose this may be the case between the Upper Devonian and the 
Carboniferous series. All veins, mineral or otherwise, come to the 
surface; and if the Tubutelle be found therein, they will probably 
indicate the presence of freshwater conditions. ‘The upper portions 
of those I have mentioned in this district represent the gossans of 
the lodes in more ancient rocks ; and if, as at Yate and Tytherington, 
the Tubutella is caught up or surrounded by the mineral matter of 
the vein, there has either been a remodification of that portion of 
the vein, or it must have been contemporaneous with the organisms 
enclosed. 7 

I have already shown that most of the mineral veins of the 
Mendips and South Wales are at least of Liassic age; and on this 
point I have much confidence in the belief that a careful exami- 
nation of the gossans and other mineral constituents of the veins in 
our more ancient rocks will repay the labour, by giving either more 
precise indications of their age or of the physical conditions under 
which they were deposited. 

Age of the Bristol Reptilia.—The varied points mentioned in this 
paper have drawn me away from the chief object which led to their 
consideration, viz. the age of the Bristol reptilia. As before 


DEPOSITS IN THE BRISTOL DISTRICT, ont 


remarked, the peculiar circumstances attending their discovery has 
always left this in some obscurity. They are now assigned by 
Mr. Etheridge to the Dolomitic Conglomerates at the base of 
the Keuper. ‘These conglomerates are in great part composed of 
rounded boulders of the Carboniferous Limestone, some of them of 
great size, and seemingly requiring glacial conditions for their 
removal and transportation. Not only do they fringe the outcrops 
of the limestone from which they are derived, but they form an 
almost continuous deposit of considerable thickness, extending for 
many square miles in the Somersetshire coal-basin, the inclined 
ragged limestone edges of which are, in places, rendered by their 
denudation quite horizontal. The removal and redeposition of the 
conglomerates indicate very troublous times, during which it seems 
impossible for any organic life to have existed; for nothing could 
have withstood the grinding-processes to which it would have been 
subjected ; and it is a significant fact that no organic remains have 
ever been referred to this period except the reptilians under notice. 

For these reasons, and from my having found the teeth of The- 
codontosawrus and Palcosaurus in the Rheetic deposit at Holwell, 
and also from my subsequent discovery of the Rhetic bone-bed and 
remains of that age almost alongside the Clifton reptilia, I had 
come to the conclusion that the latter belonged to this period—a 
view which further investigation respecting both Keuper and Rheetic 
reptilia requires me to modify. Seventeen Thecodont teeth, more or 
less perfect, are in my Holwell series. On comparison with those 
from Bristol, they are more robust, have a more wrinkled or 
striated surface, with the serrations on the edges smaller, less 
oblique. and more numerous. In my paper ‘‘On Abnormal Con- 
ditions” &c.*, I gave a section of variegated Keuper marls at 
Ruishton, near Taunton, one bed of which I described as a “ Gritty 
conglomerate, with occasional sandy bands and intermediate layers 
of marl, with fish, reptile, and batrachian remains, fourteen inches 
thick.” In this bed I have lately found some teeth of T'hecodonto- 
saurus, which appear in all respects identical, in form, structure, 
and the character of the serrations on their edges, with those from 
Bristol. It contains also Acrodus keuperinus, Hybodus, Diplodus, 
&e. Itisrather a coarse sandy bed than a conglomerate; and, owing 
to its being rather unconsolidated, its remains are very fragile. 
There seems little doubt that this bed is on the horizon of that in 
Warwickshire which has yielded identical vertebrata; and if so, the 
Bristol reptilia will have to be removed one stage later in time, from 
the Dolomitic Conglomerate to the middle of the Upper Keuper. 

Té is an interesting paleontological fact that, although most of 
the generic forms of the Keuper recur in the Rheetic beds, so far as 
I have ascertained, the species differ, and are special to the two 
formations. 


* Quart. Journ. Geol. Soc. vol. xxiii. (1867), p. 468. 


Orie G.c. No, 145. G 


82 ABNORMAL GEOLOGICAL DEPOSITS IN THE BRISTOL DISTRICT. 


Discusston. 


The PresipEent spoke of the great industry and skill of the author 
in collecting the evidence on which this paper was based. 

Mr. Tawney stated that Mr. Sanders held that the Dolomitic 
Conglomerate is of different ages in different parts of the district, 
and that the Vhecodontosawrus is high up in the Keuper series. He 
thought that the fact of these remains being imbedded in solid con- 
glomerate was scarcely reconcilable with the notion that they came 
from vein-infilling. 

Dr. Duncan, with reference to the origin of the so-called Tubu- 
tella, stated that similar tubes might be seen in course of formation 
by the escape of air-bubbles from the surface of oysters and other 
shells covered with Algze in turbid water containing carbonate of 
lime. 

Prof. Sretny spoke of the great interest attaching to the question 
of the age of the Thecodontosawrus. He considered the specimens 
exhibited to belong to at least two genera. He stated that the ilium 
of Thecodontosaurus is Crocodilian, with Dinosaurian affinities. 

Rev. H. H. Winwoop supported the views of the author concerning 
the position of the veins of Durdham Down. He remarked upon the 
difficulty of understanding the mingling of different faunas in the 
same vein. 

The Presipent supported the views of Mr. W. Sanders, as ex- 
pounded by Mr. Tawney, and bore testimony to the great value of 
Mr. Sanders’s map of the Bristol area. 

The AutHor agreed that the Magnesian Conglomerate is of diffe- 
rent ages. He thought the Thecodontosaurian remains were ob- 
tained from the top of one of the veins. He stated that, while the 
veins occasionally contained the remains of fossils belonging to more 
than one geological period, others contained organisms which 
appeared special to a single period only, implying a denudation and 
refilling in veins, at the present time difficult to account for. 


J. W. CARRALL ON CHINESE CARBONIFEROUS PLANTS. 83 


8. Norzs on the Locaritry of some Fossits found in the CARBONIFEROUS 
Rocks at T’ane Suan, Cuina. By Jamus W. Carratt, Esq., 
F.G.S. &c., of H. 1. M. Customs Service, China. (Read Novem- 
ber 3, 1850.) 


Tue fossils which form the subject of this paper were found at 
T’ang Shan, Chiao Chia T’un, in Lan Chow of the Province of 
Chih Li, and about 120 miles from the treaty port of Tientsin, 
in a N.N.E. direction from that port. 

A Chinese company has been formed, known as the “ Chinese 
Engineering and Mining Company,” to work the bituminous coal 
deposits there found on the European system. 

Mining operations were commenced in 1878 by ascertaining the 
locality of the coal-seams by boring with the diamond boring- 
machine. The bore-holes, three in number, averaged about 40U feet 
apart; the third and deepest of the three driven reached a depth of 
536 feet. 

The seams dip at an angle of 45° to the north, calculated by the 
angle of the strata found in the bores. 

The thickness of the coal-seams, not taken at the slant but 
parallel to the beds, is as follows :— 


ft. in 
iNowi-seam, N. of King’ seam: o..4.... 6 O odd. 
TSO? GOUT USS nO Ire el cee era Ibs 
Wosileseam, o: of King seam ...... 5... Oued 
No. 2 seam cy Mn a ete ec ie Ra Ia nd 
No. 3 sear ey abla Meeps ONS ae oe Oe Id 
No. 4 seam Boy) 1 aI ae ae ere ALD 


The seams are in curves or folds; one seam that comes to the 
surface at the back of the colliery, again appears above ground 
halfway from the colliery to Kai Ping (say three miles from the 
first outcrop), and again five miles further on, trending in a north- 
erly direction. 

Some very good magnetite, containing between 45 and 8U per cent. 
of metallic iron, has been found at Pai Mah Shan, about seventeen 
miles from T’ang Shan. It is intended to erect roiling-mills near 
the colliery, and place them also under foreign superintendence, 
trams being laid down (after permission has been obtained) between 
Pai Mah Shan and the hills for the transit of the ore. 

The most striking feature of the geological formation of the 
country round ‘l’ang Shan is, that above the Carboniferous system 
is first loose sand and then loam, the loam being uppermost, and 
extending but deepening all the way to Tientsin. Decomposed 
red sandstone was seen in the distance to the north. 

A gradual ascent commences four miles before coming to Lu 
T’ai by land from Han Ku, and continues all the way to T’ang 
Shan. The colliery is situated in this incline, and is about a mile 


84 J. W. CARRALL ON CHINESE CARBONIFEROUS PLANTS. 


from the nearest hill, T’ang Shan, which I do not think is higher 
than 300 feet or thereabouts. 

From the roundness of the hills and the way the chain is detached 
in places, there is not the least doubt in my mind that this part of 
the country was under water, and 1s now gradually rising, this also 
being proved by the fact that, according to Chinese history, the city 
of Tientsin was at one time situated on the seaside, and itis now some 
twenty-five miles inland. 


Note on the Specimens, by W. Carrurumrs, Esq., F.R.S., F.G.S. 


The specimens from China belong to a single species of Annularia ; 
and I have no doubt that it is A. longifolia, Brongn., which is so 
abundant in our coal-measures in Britain, and is found on the con- 
tinent and in North America. It occurs throughout the whole of 
the coal-bearing beds, from the lowest to the highest. 

Newberry and others have described fossil plants from China asso- 
ciated with beds of coal; but these belong to Trias or still later 
formations. The great interest of this communication is that it 
records the existence of the true coal-measures in China, and gives 
a range to a well-known Carboniferous plant, in harmony with what 
we already know of other western contemporaneous Paleozoic 
plants. 


DISCUSSION. 


THe PrEstipent remarked on the great interest attaching to the 
discovery of these Coal-measure fossils. 

Mr. Carrurners pointed out the interest attaching to the com- 
mencement of coal-mining in China. He remarked upon the world- 
wide distribution of certain Paleozoic species of plants. 

Pror. Jupp spoke of the importance of this discovery in a country 
the geology of which was so little known. 

Mr. Branrorp suggested that possibly future observation might 
show that these plants were associated with Mesozoic forms. 
The Palseozoic flora had not a worldwide distribution. Mesozoic 
types of plants had been observed in Australia in Paleeozoic beds. 

Pror. Seetey demurred to the conclusions of Mr. Carruthers as 
to the distribution of Paleozoic plants. 

Mr. Buaxs supported Mr. Carruthers’s views. 

Mr. Carrutuers, in reply to Mr. Blanford, said he believed that 
the Glossopteris-beds of Australia are really Mesozoic and not Palee- 
ozoic in age. . 

Mr. Buanrorp said the late Rev. W. B. Clarke had found Glos- 
sopteris-beds intercalated amongst Paleozoic marine fossiliferous 
beds in coal-pits in New South Wales. 

THe PRESIDENT supported the views of Mr. Carruthers and Mr. 
Blake, in opposition to those of Prof. Seeley. 

Pror. Jupp supported Prof. Seeley’s views as to the existence of 
life-provinces in Palzeozoic times. 


tor Ho 


fl 


Geol. Soc. Vol. XXXVII. PLV. 


WIGS 


_ 400 te. 


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Bagshot “Sands 


‘s tation. 


How-ledge 
Lime: stone 


RL Lyne 


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786 


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| 
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SECTION OF THE GILINEIRS IN TROMME/AIN(D) ANID) = OIL TE TEIL, ANS), = WISI =) WHIT 
' W/6? E<— > Wiss. 
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OT ft. 

if 0 t 1 a nr d B a y = . 2 
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Teclend Bey Firting Ree 

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Varden sands Wardein sands 

G 0 Zz n e l L I @ v 

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were ji 4 NIGE<—> S16°W 

i Warden. Battery (Eoerbeppenth 
: pe Bramble Post- Tertiary Gravel cap eer 

Petiory Linstone Colwell Limestone 
—— ee U. Head: Np eee U. Heador Chine = : ____ Me Headon 
ee f Ue Sands & hazelnut bed » Hea ae z tipper ee ae — 
} Grass slopes - 


EN 


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pero = TO ry : with Lymnea 
Limestone = Cventricosaor bed. gime re = 5 

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Scale for Vertical Heights 2500. Scale for Horizontal Distances 5280, or 12 inches w the Mile (one inch 0 46 yards.) 


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Fral® Dangerfield Zit: 22 Bedtord. S* Covent Garden 
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THE BEDS AT HEADON HILL AND COLWELL BAY, ISLE OF WIGHT. 895 


9. On the Beds at Heapon Hitt and Cotwett Bay in the Isle of 
Wight. By H. Kunrrne, Esq., and E. B. Tawney, Esq., M.A., 
F.G.S., of the Woodwardian Museum, Cambridge. (Read De- 


cember i, 1880.) 
[Puate V.] 


I. InrRopuctIon. 


In a recent communication laid before the Society* the opinion was 
expressed that a serious error had been made by almost all previous 
writers in regarding the marine beds at Colwell Bay and Headon 
Hill as on the same geological horizon ; we read :—‘* We shall now 
demonstrate that the Colwell-Bay marine beds are not, as has been 
hitherto supposed, the equivalent of those of Headon Hill and Hord- 
well Cliff, but that they occupy a distinct and much higher horizon.” 
Upon the correction of this supposed error a new classification and 
nomenclature for the Upper Eocene formation of Britain was pro- 
posed. . 

The author further, after a review of the paleontclogical evidence, 
arrived at the conclusion that, on the one hand, the fossils in the 
Headon Hill and Hordwell Cliffs were identical, while, in the second 
place, those of Colwell Bay, White Cliff, and Brockenhurst presented 
the closest agreement among themselves. Then, comparing the 
former two localities, taken together, with the latter three, taken 
together, he considered (1) that the fauna of the first group was 
largely estuarine, and that of the second group marine; (2) that 
less than half the forms found in the former occur in the latter; (8) 
that the fauna of the former approximated more to that of the Barton 
beds, having about one third in common with them, while not more 
than one fifth of those from the latter three localities occur at Barton ; 
(4) that the fauna of the former two agreed with that of a series of 
beds on the Continent which underlay and were older than beds 
containing the fauna of the last three. 

In the following communication the authors attempt to traverse 
these points in the paper above referred to, in succession. By reference 
to detail-sections they argue that the stratigraphical evidence is 
plainly demonstrative of the identity of the Colwell-Bay and Headon- 
Hill marine series, the beds being continuous through the cliffs and 
easily traceable. 

Referring to the paleeontological evidence, it is shown from collec- 
tions, made with their own hands this year, (1) that the fauna of the 
Colwell-Bay and Headon-Hill beds are identical; (2) that this fauna 
differs considerably from that of the Brockenhurst bed, which occupies 
a lower horizon; (3) that the Colwell-Bay bed has less than one 
third of its species common to Barton beds, while the Brockenhurst 
fauna has nearly one half in common with Barton beds. 

* «On the Oligocene Strata of the Hampshire Basin,” by Prof. J. W. Judd, 
E.R.S., Sec. G.S., Quart. Journ. Geol. Soe. vol. xxxvi. p. 1387. 

Q.J.G. 8. No. 146. H 


86 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


We conclude therefore that no reason has been shown for upsetting 
the classification of the strata adopted by the Geological Survey, and 
which, for nearly a quarter of a century, has been received among 
geologists. 

Certainly we wish to uphold in its integrity the work of the late 
E. Forbes *, and the classification of beds adopted by him when 
Palxontologist to the Geological Survey, and subsequently confirmed 
by Mx. H. W. Bristow, F.R.S., in the second Survey Memoir? on the 
Isle of Wight. 

One of the authors, from his long residence in the district and his 
constant occupation with these beds, has been long satisfied that 
E. Forbes’s account of the beds is true to nature, and his classification 
fully borne out by lithological identity of beds, as well as by distri- 
bution of the fossils. The present notes, then, are based on his part 
upon an aquaintance with the district, and the work of the Geological 
Survey there during its progress, supplemented by subsequent visits, 
and specially this summer by a joint examination by both, including 
measurement of beds and collection of fossils, which, however in- 
complete, was made bed by bed, and represents the prevailing fauna 
of each—a point on which we lay great stress. 

We do not wish to delay over the history of previous opinion, 
which has been sufficiently treated in Forbes’s and Prof. Judd’s 
memoirs ; but the latest criticism of Forbes’s work (op. cit. p. 141) 
may be alluded to. 


II. Tornanp anp Cotwett Bays. 


The Survey Horizontal Section east of Headon Hill_—The first 
point at issue between Forbes supported by “ nearly all observers, ” 
on the one hand, and Prof. Judd on the other, is whether certain 
marine beds known as the Middle Headon (including the “* Venus-bed” 
of local collectors) in Colwell Bay are rightly associated with similar 
marine beds in Headon Hill. The Survey identify them, and 
correlate the freshwater beds immediately above and below as Upper 
and Lower Headon respectively in both localities. This succession, 
however, is stigmatized (op. ct. p. 142) as a “ mistake ” of which the 
‘‘ primary cause ” is considered to be an “ assumed” existence ‘“ of 
a great anticlinal fold of which the summit is supposed to be seen 
in Totland Bay. The manner in which this supposed anticlinal is 
regarded as having affected the strata is illustrated in Prof. E. Forbes’s 
memoir, pl. vii. fig. 1, and also in Sheet 47 of the Horizontal Sections 
published by the Geological Survey. And yet further on we read 
(op. ert. p. 146), “ at Totland Bay there is undoubted evidence of the 
presence of a slight anticlinal fold having its summit near Widdick 
Chine, to the westward of which the beds for a short distance have 
a slight dip to the south;” so that after all the only mistake the 
Survey could have made would have been to exaggerate the dip. 

We are next told of the E. and W. flexure, which causes a slight 

* On the Tertiary Fluvio-marine Formation of the Isle of Wight, by Prof. 


E. Forbes, F.R.S., 1856 [Memoirs of the Geological Survey of Great Britain]. 
+ The Geology of the Isle of Wight, by H. W. Bristow [Sheet 10], 1862. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 87 


dip to the W. in Headon Hill; and it is implied that the Survey 
section is false, owing to the neglect of this consideration. We 
must point out, however, that this Section, Sheet 47, fig. 2,in which 
alone the anticlinal is shown, does not go through the summit of 
Headon Hill at all, and passes about a half mile inland at the latitude 
of Widdick Chine : init the Upper Bagshot beds are made to appear a 
little above the sea-level at that spot; and we have great confidence 
that the calculations on which this is grounded are correct*; they 
would be brought up by the anticlinal of which the existence has 
just been acknowledged. So far then, we may observe, no reason 
has been shown why the Survey Section should not be received as 
correct. 

Prof. Judd’s Section (op. cit. pl. vii..—We do not find any 
indications of the direction in which this section is drawn ; but, from 
the names of chines which occur in it, we presume that it is intended 


to start from Alum Bay in a N.E. direction through Headon Hill, 


and after that to follow the coast-line; in this case it is not precisely 
comparable with the Survey Section, whose direction is indicated on 
the map as passing inland and crossing from sea to Solent. It will be 
seen at once why it does not correspond to nature, and agree with 
the views of other observers. It will be noticed that the Marine 
bed of the Headon Group (3 of fig. 3, pl. vii.) is made to exist at the 
sea-level near Widdick Chine; and in the letterpress we read 
(op. cit. p. 147) “it is admitted on all hands that at the north-east 
angle of Headon Hill the marine band [ Middle Headon beds | makes 
its appearence just above the sea-level.” On the contrary, we cannot 
imagime any one putting the bed in this position. At the spot indi- 
cated the top of the Middle Headon is about 105 feet above the sea- 
level; so that the section, in our opinion, is erroneous: the dip 
thereby is exaggerated ; and 105 feet of beds are intercalated which 
do not exist. We shall prove this presently by a detail-section at 
this point of the hill; at the present moment we wish to point out 
that, with the correction of this error of 105 feet, the argument 
against the accuracy of the Survey section entirely breaks down. 

- Thus, we are told that the height of the Bembridge Limestone 
above the sea-level at this point is 250 feet (op. ct. p. 147); then, 
the marine band being put at the sea-level, it follows that 250 feet 
of strata must intervene between that and the Bembridge Limestone ; 
“but the Geological Survey [vertical] section shows less than one 
half of that thickness of beds, and in Colwell Bay the distance 
between the Bembridge Limestone and the marine band is 120 
meew yy SS ss But 250 feet of strata is precisely the thickness required 
by my interpretation.” Since we have to subtract 105 feet from 


* The greater thickness of Upper Bagshot beds above the sea-level in the same 
line of section in the old edition of Sheet 47, and in the Plates of the Memoirs, 
seems due to the outline of the ground at the S. end being raised too high above 
Ordnance datum; probably the accurate height of the Beacon Hill was not 
obtainable till 1870, when the revised edition of Sheet 47 was published ; other- 
wise the sections are practically identical. 

t It would be 133 feet, according to Mr. Bristow’s estimate (Forbes’s Mem. 
p. 142), to the top of the Bembridge Limestone in Colwell Bay. 

H2 


88 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


the estimate in the paper above referred to, we shall certainly have 
to abandon this section in favour of that of the Survey; for the 
thickness left, viz. 145 feet, more nearly corresponds with the 
thickness of the Bembridge, Upper Headon, and Osborne beds, which 
are stated by the Survey to exist at this spot, and whose thickness 
would be 144 feet *. 

It would appear, therefore, that there is no necessity for supposing 
“that, in a distance of little more than one mile, a mass of 
beds 120 feet thick has expanded to 250 feet, and, further, that the 
beds have been entirely changed in their mineral character.” 

We do not understand the warning (op. ct. p. 146) against 
“trusting to the general impression which is produced by viewing these 
beds from a distance,” nor the purport of the following statement :— 
“ The strata of How Ledge and Warden Point are seen in such a true- 
scale section to be clearly continued in precisely similar beds ap- 
pearing underneath the gravel of Headon Hill.” The section offered 
to us is on rather too small a scale to show detail ; but, in our opinion, 
the beds are inaccurately laid down in Warden Cliff, and no such 
bed as the Brockenhurst bed occurs at all in Headon Hill. 

Before we commenced drawing our section, we traced the beds 
along the cliffs, measured their thickness, and obtained their height 
above the sea-level at various points, but found it possible only to 
show general results in the horizontal section ; the details are embodied 
in the vertical sections. 

Vertical Section at North-east Corner of Headon Hill (fig. 1, 
p- 91).—We may now proceed to a more detailed account of the 
beds. We will begin near the N.E. corner, where the Bembridge 
Limestone is seen, indicated on the section (op. cit. pl. vii. fig. 2) 
with an asterisk, and lettered 250 feet altitude. (The quarry there 
is not now at work; but it is the place at which one of us has ob- 
tained most of the finest specimens of Palwothertum which have been 
found in the Isle of Wight.) We take the thickness from the 
Survey Memoir as 25 feet. 

Beneath this, in the section (fig. 3, pl. vil.), we notice a blank 
space with the legend “slopes covered by gravel and landslips.” 
We think this scarcely a correct description. Landslips exist in that 
the clays and marls tumble and form taluses ; but we saw no gravel 
covering the slopes between the Bembridge Limestone and Widdick 
Chine ; nor indeed does it entirely conceal the beds between the 
Bembridgeand Upper Headon Limestones all the way to Heatherwood 
Point in the other direction; at intervals tumbled gravel conceals 
a limited portion of them. 

The gravel west of Widdick Chine is not accurately depicted: the 
thickness is exaggerated, while its horizontal extent is overestimated 
here t. It does not appear in the cliff certainly beyond 80 yards 

* Taking the Bembridge Limestone at 25 feet, the Upper Headon and 
Osborne at 119 feet, as read off their vertical section by scale, after altering 
their lower boundary a few feet to correspond with our own. 

t Mr. Bristow (Forbes’s Mem. p. 105) gives the entire horizontal extension 
of the Lacustrine beds (Post-Tertiary) on both sides of the chine as 350 yards ; 
the section under review makes the gravel extend about 720 yards. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 89 


from the chine on the west side (private grounds interfere with 
examination nearer the chine). 

We should say, from our examination of the ground which inter- 
venes between the escarpment of the thick Upper Headon Limestone 
and the Bembridge, that there is no difficulty in seeing what beds exist 
there. It is true they are sufficiently interrupted by local taluses to 
cause trouble in making a continuous measurement ; but the tumbled 
portions are partial and affect only a few feet of beds at a time, so 
that by moving the observer’s position laterally it is possible to see 
all the bedsinturn. ‘This we claim to have done; we do not pretend 
that our measurement of the beds is any thing but rough, though 
controlled by repetition, because we had no levelling-instruments with 
us, and in shitting horizontally from one spot to another there might 
be frequently a slight error in picking up the next bed to be 
measured. We might describe the beds immediately below the Bem- 
bridge Limestone, in descending order, as follows:—yellowish and 
ochry marls, red and grey mottled marls, marly clays with nodular 
bands, greenish-grey clays, pale greyish clays, grey and ochry clays, 
stiff pale or whitish clays with calcareous lenticular bands, red and 
green mottled marls, ‘“ cherry marl” with calcareous bands. These 
are the Osborne beds of the Survey ; and they come in the precise 
position assigned to them in the Survey Memoirs. They are well 
characterized by the ‘cherry marl”—the mottled red and pale 
greenish marl which distinguishes the group from the Upper 
Headon ; and we consider the subdivision a very useful one in the 
classification of beds. Their thickness here, by our comparatively 
rough way of measurement, is 70 feet. The vertical section of the 
Survey Sheet 25 gives 71 feet, reading off by scale down to the bed 
which we have taken as our boundary; their measurement seems to 
have been taken near Heatherwood Point, where this series contains 
a thick limestone of 18 feet; as noticed by previous observers, this 
limestone thins out to the east, and is only represented by nodular 
calcareous bands at the east end of the hill ; its loss will probably be 
compensated by an increase in the clays. Our results are perhaps suffi- 
ciently near to those of the Survey to prove that the same series of 
beds has been examined in both cases. Weshould remark that from 
the lower red beds it is perfectly easy to draw a continuous vertical 
section to the beds below; and from here our measurements to the 
Lower Headon are uninterrupted in a vertical line. The Osborne 
beds yield few interesting fossils; Lomnea is abundant in the 
calcareous bands ; but, as noticed by Forbes (Mem. p. 81), the shell- 
substance is not preserved. These beds are identical lithologically 
with the mottled red and pale green series at Cliff End. 

The beds next below the red series are grey and ochry clays, very 
rich in Potamomya gregaria and Paludina lenta beautifully pre- 
served: we place these in the Upper Headon; they are about 15 
feet thick *. 

* Tn the Survey Section no. 5, at Headon Hill (sheet 25, Vert. Sect.), these 
clays are included in the Osborne series; but in the Vert. Section. no. 4, at Col- 
rh the boundary is so drawn that analogous claysare put into the Upper 
Headon. 


90 ON THE BEDS AT HEADON HILL AND COLWELL BAY. 


This brings us to the vertical escarpment of the thick limestones, 
so conspicuous a band along Headon Hill cliffs that it is indicated 
on the Ordnance maps, both on the 25-inch and the 6-inch. We 
pause awhile to draw attention to the fact that we have accounted 
for about 110 feet of beds from the top of the Bembridge to the top 
of the great limestone (Upper Headon) ; and the Brockenhurst series 
does not exist here. There is not a single marine fossil to be found 
am that interval; nor is there any bed with the faintest resemblance 
either lithologically or palceeontologically to the Colwell-Bay Venus-bed. 
This is in opposition to the view (I. ¢. p. 176 e¢ passim) that the 
Colwell-Bay series exists here “‘entirely concealed” by some supposed 
gravel talus*; yetit is upon the existence of such a second marine series, 
thus supposed to be added above the Venus-bed that the presence of 
a Brockenhurst series at the west end of the island is inferred. 

Next we turn to consider the thick limestone of the escarpment, 
the Limneea-limestone of the Upper Headon. It is in several beds, 
of which details are given by the Survey; we measured it by sus- 
pending a tape, and found it 27 feet (fig. 1, p. 91). 

The only difficulty in correlating the Headon-Hill beds with those 
of Colwell Bay is centred in this limestone: it might be a difficulty 
to those who would have expected a priori that the limestone would 
have maintained its thickness in direction of dip for a mile or two 
to the north; for we identify it with a limestone at Cliff End not 
above 1 foot 8 inches thick. It would be equally a difficulty aecor- 
ding to the correlation which identifies it with the How-lLedge 
limestone (op. cit. p. 144); im this case the 27 feet has thinned to 
3 feet at How Ledge, a distance of 14 mile in a straight line, while 
in Warden Cliff (only 926 yards distant from the Headon-Hill bed) 
it is about 5 feet; so that it must have thinned very rapidly at the 
first stage. The limestone in the Upper Headon at Cliff End, with 
which we identify it, is distant 1 mile 926 yards. In either case it 
thins out considerably to the north, as noticed by E. Forbes (Mem. 
p. 84). We shall be able to prove that it does not occupy the same 
position as the How-Ledge bed; for we have recognized that bed, 
which forms the summit of the Lower Headon, at a lower position 
in Headon Hill and in its natural position, viz. below the marine 
series (Middle Headon), as in Warden Cliff, where it was last seen. 
As a paleontological distinction between the Upper Headon limestone 
of Headon Hill and Cliff End + and the Lower Headon limestone 

* With respect to the “inextricable difficulties and confusion” (7. ¢. p. 144) 
in which the Survey is supposed to be involved by their not allowing the Col- 
well Marine bed to come where the Osborne beds are placed, and which is sup- 
posed to be shown by 48 feet of strata being classified in the letterpress (Forbes’s 
Mem. p. 81) as Osborne, while in the plate they are classed as Upper Headon, 
this is merely a, question of classification and the drawing of a boundary-line, 
matters entirely subjective and not affecting the total thickness ; their vertical 
section shows almost the same thickness of beds consistently, notwithstanding 
certain irregularities in the boundaries and classification. 

t In the legend to the Survey Vertical Section, sheet 25, no. 4, this limestone 
in the Upper Headon is said to form How Ledge; this is plainly an oversight 
or clerical error, as is also the statement in Forbes’s Memoir, p. 132, that the 
How-Ledge bed is faulted in Warden Cliff. The fourteen faults mentioned affect 
the Upper Headon limestone at Cliff End: their section is fairly correct; but 


there seems to have been some confusion between Warden Point and Cliff-Hnd 
Point in the letterpress. 


Fig. 1.— Vertical Section of Beds at the North-east corner of Headon Hill. 


63-8 0 


UppEerR HEADON. 
bs 
oOo 


® 8 
9 in.-2ft 
6 in 

6 in 

ft." 1m: 
1 10 


MIDDLE HEADON. 


~~ A 
ow 
f=) 


seen. 


LowER HEADON. 
CO 
for) 


(Scale, 8 feet to the inch.) 


BEMBRIDGE LIMESTONE ...... a0 feet. 


OSBORNE BEDS 


SO esac r esos ersorccras 


WPPER HIBADON ....0:0.0-0s00008 9 a (total). 


al 


& aye 36 


CO 


Part of thick Limnea-limestone. Limnea fusiformis, &c. 


Laminated greenish clay, with broken Paludina. 


Whity-brown to buff sands, with layers of lignitic matter. 


Greyer sands below. Potamomya, Melania muricata, Unio, Paludina lenta. 


> Lignite. 
* Greyish-green oye. C. ven- \ Vicarya concava, Marginella vittata, Neri- 
tricosum-bed With ...........026 tina concava, Melania muricata, &e. 


--- Limnea-limestone, soft and crumbling, with a thin lignite at top. 
-*-Verdigris-green clay, with rootlets. 


Limnea-limestone. 


Stiff green clays with conchoidal fracture in drying. 
Oyster-bed towards the base. 


Fusus labiatus, Mel. fasciata, M. muricata, 

Clay becoming greyer below. ) Nerita aperta, Cer. variabile, C. pseudo- 

IM OSSIUS |i cccestcseerstessseccceees cinctum, Ostrea velata, Mytilus affinis, Cor- 
bicula obovata, Lucina colvellensis. 


Alternating grey and ochry clays. 


Cyth. incrassata, Mactra fastigiata, 

“‘ Venus-bed,” richest portion, con- Mya angustata, Corbicula obovata, 
tains scattered flints, brown sandy | Wucula lissa, N. headonensis, Trig. 

- clay becoming green clay and sand deltoidea, Fusus labiatus, Cancell. 


Ibelow-m Ue ROSSILSSscsceseccs-0s.-sanenescess elongata, Melanopsis fusiformis, 
/ Voluta spinosa, Vic. concava, Natica 
Thin grey sandy clays, Studert. 


weathering brown. 


Cytherea incrassata &c. scattered throughout. 


Mya angustata, especially near base. 


Trig. deltoidea, Cer. pseudocinctum, 
Chocolate-brown or blackish sands. { Natica labellata, Melan. fusiformis, 


Trigonocelia-bed. 


Blackish-brown sands, Weritina-bed: WV. concava, M. fusiformis, C. obovata. 
Very stiff tenacious clay. 


Limnea-limestone, ‘‘ How-Ledge limestone:” ZL. longiscata, fusiformis, &c. 


Whity-brown or yellow sands and sand-rock, with layers of Paludina and 
Potamomya. 


[The base concealed by tumble and undercliff. | 


92 H, KEEPING AND E. B. TAWNEY ON THE BEDS AT 


of How Ledge, we may adduce that the former is very rich in 
Planorbis, while in the latter- these shells are comparatively rare, 
the fossils being chiefly Limnea. 

As the Upper Headon limestone is the strongest bed in the 
section and forms an escarpment through the greater part of Headon 
Hill, in it we may obtain a datum line. We take as a point of 
reference the spot where the top of the escarpment cuts the surface 
of the ground or outline of the cliff; this 1s seen on the 6-inch or 
25-inch map to be about halfway between the 100 feet and 200 feet 
contour-lines. From these points of known altitude, by the aid of 
the barometer, we obtain the height of our point of reference: ibis 
about 140 feet above Ordnance datum. 

The beds below the Limnea-lmestone are green clay, 1 foot, 
with broken Paludina, then pale butt or whitish sands, varying from 
63 to 8 feet, with occasional layers of lignitic matter, Potamomya, 
and Melana muricata; below, where it is sometimes grey, Unio 
Solandri and Paludina lenta may occur. We take this bed with 
the 3-inch lignite below as the base of the Upper Headon. The 
boundary chosen is, of course, arbitrary ; but the fact of the next 
bed being decidedly brackish inclines us to draw the division from 
the Middle Headon here ; the Survey vertical section, sheet 25, places 
it a few feet lower. 

Our estimate for the Upper’ Headon at Headon Hill amounts to 
a thickness of 50 feet; the thickness on the Survey vertical section 
is given as 37 feet ; but if we read off the distance between the beds 
which we have taken as boundaries, it becomes 48 feet. The com- 
bined thickness of Osborne and Upper Headon beds, according to 
the Survey section, is 119 feet, 2.¢. adopting the top of the C. ven- 
tricosum bed as the boundary ; our estimate, taken at the north-east 
end of the hill, is 120 feet. The agreement is sufficiently close to 
render it probable that a thickness above the average of the cal- 
careous portion is accompanied at the same spot by a diminution in 
the clays; so that the balance of average thickness is maintained at 
both ends of the hill. As we have said, we think it convenient to 
retain the name “ Osborne Series ” for the red and greenish mottled 
clays and marls and pale greenish-white limestones, since these 

‘physical characters distinguish them at once along this side of the 
Solent. We must decline to accept the statement (op. cit. p. 169) that 
‘‘under this name beds lying below the Brockenhurst series, as at 
Headon Hill, have been confounded with others on a totally diffe- 
rent horizon, above the Brockenhurst series.” 

We next come to the Middle Headon. HH. Forbes relates (Mem. 
p. 85) that the uppermost and lower portions at Headon Hill are 
brackish-water beds abounding in Cerithium ventricosum, C. pseudo- 
cinctum, C. concavum, Neritina concava, and Nemature, the condi- 
tions being less purely marine than at Colwell Bay. This is, no 
doubt, true of the series as a whole ; for below the C. ventricosum bed 
there are two freshwater Limnea-limestones. But it appears to us 
that too much has been made of this ; for instance, the lower Neri- 
tina-bed is identical in Headon Hill with the similar bed at Warden 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 93 


Cliff, and again in Colwell Bay, both physically and as to its fossils ; 
when we come to the portion of the series richest in Cytherea incras- 
sata known as the ‘“ Venus-bed” proper, we find identically the 
same fossils as in Colwell Bay; we cannot detect any difference, so 
far as our researches go. 

The uppermost bed is dark greenish clay, varying from 9 inches 
to 2 feet in thickness. When at its thickest the sands above are 
proportionally thinner. It is extraordinarily rich in fossils for the 
depth of one foot, chiefly Meritena concava, Cerithium (Viearya) 
concavum, C. ventricosum, Corbicula obovata, Marginella vittata, 
Melania muricata, Melanopsis fusiformis. We may callit the C. ven- 
tricosum bed; for this fossil seems almost confined to this horizon, 
while the Vicarya concava is found here all through the Middle 
Headon, though especially plentiful in this bed; its underside is 
occupied by an impure lignite band, with freshwater shells (Limnea 
and Planorbis), and rootlets, lying on a thin crumbly buff Limnza- 
limestone. Both together are 4 to 6 inches. Below is verdigris- 
green clay, 6 inches; next a buff freshwater limestone with Limnea, 
1 foot 8 inches to 2 feet, the shells frequently blackish in colour. 
These beds betoken certainly a recurrence of freshwater conditions 
after the brackish bed above. Next follow truly marine beds—first 
stiff green and grey clay, about 9 feet, with a conchoidal fracture 
when dry. ‘The abundance of oysters, O. velata (Wood), is the chief 
fossil feature ; these and the other fossils occur mostly towards the 
base. Such fossils are Musws (Pisania) labiatus, Nerita aperta, Me- 
lana fasciata, M. muricata, Cerithiwm variabile, C. pseudocinctum, 
Mytilus affinis, Corbicula obovata. 

The fossils cited are merely the most common, such as may be 
found in a few minutes’ search; but these being the most charac- 
teristic, seem to us precisely those required for the identification 
and correlation of beds. 

Comparing the beds noticed so far with those at Colwell Bay, 
we observe practically identity of fossils; this last, which we may 
call the oyster-bed, is identical with the oyster-bed of Colwell Bay ; 
in both localities OC. ventricoswm occurs above this bed, and there 
only, so far as we know. 

Next below follow alternations of grey and ochry silts, 3 feet, in 
which we observed no fossils. Below are 11 feet of beds, brown 
sandy clay above becoming green clayey sands below, and then grey 
sandy clays: this is the ‘ Venus-bed ” of collectors ; Cytherea imcras- 
sata occurs near the top and is scattered throughout the whole bed, 
but is most abundant for the space of one foot. The fossils obtained 
from the Venus-bed in a few minutes’ search were Cytherea incras- 
sata, Mya angustata, Mactra fastiqiata, Psammobia rudis, P. westu- 
arina (Hid. MS8.), Nucula headonensis, N. nudata, Trigonocelia del- 
tordea, Corbicula obovata, Vicarya concava, Ancillaria buccinordes, 
FPusus labiatus, Cancellaria elongata, Natica Studeri, N. labellata, 
Voluta spinosa, Melanopsis fusiformis, Nematura parvula, Limnea 
longiscata, Planorbis obtusus, crab-claw (Callianassa), coprolite. 


94 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


Towards the base Mya angustata and Corbicula obovata were parti- 
cularly abundant. 

Next below the Venus-bed is the Trigonocelia-bed, chocolate- 
brown sands, 34 feet, with sometimes a blackish tint ; we so call the 
bed from the principal fossil which occurs at precisely the same 
horizon in Warden Cliff and Colwell Bay; other fossils are Ceri- 
Ehium pseudocinctum, Melanopsis fusiformis, Natica labellata. Below 
are blackish-brown sands, 6 inches; at base is enough carbonaceous 
matter almost to amount to a lignite band; this may be called the 
Neritina-bed, the chief fossils are Weritina concava, Melanopsis fust- 
formis, Corbicula obovata, the latter in perfect condition, very large, 
and showing concentric colour-bands. The Nertina-bed occurs in 
the same position towards the base of the series in Warden Cliff 
and Colwell Bay. This can only be explained by admitting that 
the marine series in Totland Bay and Colwell Bay are identical ; 
the Ventricosum bed at the top, and the Neritina and Trigonocela 
beds at the base, identical in physical and fossil characters, are 
strong presumptive proof of this. 

Below is very stiff dark-grey clay, 1 foot to 1 foot 3 inches; 
fossils occur in patches, Neritina concava, Cerithium pseudocinctum, 
Melana muricata, Limnea, Corbicula obovata. This is the lowest 
bed of the Middle Headon here. Summing up, we obtain a thickness 
of from 31 feet 9 inches to 33 feet for the Middle Headon of Headon 
Hill at the N.E. end. Reading off the Survey vertical section by 
scale, we obtain 35 feet for it between the boundaries adopted by 
us * for the thickness towards the west end. 

The height of the base of the Middle Headon above the sea at 
this point, viz. about 120 yards in horizontal distance west of our 
reference-point, is by subtraction 72 feet. Direct barometric obser- 
vations gave about 70 feet. We have already used these figures 
when alluding to the position assigned to this series. 

The first bed of the Lower Headon is a Limnea-limestone of the 
usual buff colour: it is 2 feet thick at this point; but a little further 
west we obtained a measure of 4 feet. This is in our opinion the 
well-known bed which forms the top of the Lower Headon in Warden 
Clif, where it is quite a marked feature. It has there and in Col- 
well Bay precisely the same position in the series, supporting the 
Middle Headon—recognized by the Neritina-bed with all its charac- 
teristic features, the Trigonocelia-bed, and so on. From Warden 
Cliffit is traceable uninterruptedly to How Ledge, where it disappears 
beneath the sea-level ; we therefore speak of it as the ‘“‘ How-Ledge 
limestone.” It is correctly drawn on the Survey vertical sections, 
sheet 25, nos. 4 and 5, where in the legend is a clerical error, to 
which we have already alluded. This bed is so distinctly lacustrine, 


* Some irregularity in the boundaries of the Survey vertical section is to be 
noticed : viz. in the Headon-Hill Section, no. 5, the boundary is placed below 
the How-Ledge Limestone ; in the Colwell-Bay section, no. 4, the boundary is 
placed at one bed above the How-Ledge Limestone; this seems an error of the 
engraver, and of course does not affect the thickness of the beds. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 95 


being almost made up of Limnea shells and their débris, that. it 
seems most convenient to include it in the Lower freshwater group ; 
and it makes an especially good boundary. 

The beds which follow next below the Limnza-limestone at this 
spot are whity-brown or yellowish sands, and sand rock with 
Paludina and Potamomya in layers ; 8} feet are seen; this is all 
that is exposed of the Lower Headon here. At this spot all the 
eround below is tumble or undercliff; and for a more complete 
section of the Lower Headon in the present state of the cliffs we 
shall do best to take the one exposed in Warden Cliff and under 
Totland Bay Hotel, where the fresh road-cuttings to the New Pier 
have been of service to us. 

We consider that we have already sufficiently proved the identity 
of this lower limestone in Headon Hill with the How-Ledge bed of 
Warden Cliff; but if any objection be raised that its relations to 
certain specified bands in the (Middle Headon) Marine series being 
found identical in both localities is not conclusive—even though it 
has been shown that no other marine bed exists in Headon Hill— 
we have further means at our command. ‘There are, however, only 
two beds in Headon Hill with which the How-Ledge bed could be 
continuous: viz. either it is the same as the thick Upper Headon 
limestone (which we consider impossible, as the beds both above and 
below would then just be reversed, viz. freshwater above and marine 
below, instead of vice versa), or it must be identical with the one to 
which we assign it. It is possible, however, to settle the point by 
ocular demonstration*. Though the How-Ledge limestone is 
denuded from the top of the curve between Weston and Widdick 
Chines, some of the lower beds are traceable the whole way. Ac- 
cordingly we can join on the section in Headon Hill to that in 
Warden Cliff. We account thus for a continuous section of beds 
from the lowest seen beds of the Lower Headon, through the Middle 
and Upper Headon of Colwell Bay, to the Bembridge Limestone on 
the north, and again from the same base of the Lower Headon, 
through the succeeding Lower Headon beds in the cliffs between 
Weston and Widdick Chines, to the sand below the How-Ledge 
limestone at the N.E. corner of Headon Hill, and thence up to the 
Bembridge limestone on the south ; and we find that the section is 
identical in both cases. There is only one Marine (Middle Headon) | 
series, lying between two freshwater ‘series, the Lower and Upper 
Headon. 

Of course, all this has been done before by the Geological Survey, 
and our work is nothing but a confirmation of results already suffi- 
ciently established by E. Forbes and Mr. Bristow. 

On our horizontal section of the coast we have endeavoured ‘to 
represent the position of the beds in the cliffs and the extent to which 

* We are indebted to the Rev. O. Fisher, F.G.S., who specially visited Tot- 
land Bay this autumn, for the information that the Venus-bed is found in the 
Totland-Bay brickyard some little way above and behind the top of the cliff 
between the chines. He points out that since this is the only part where it is 


missing from the cliff, itis the link needed to prove that the bed is continuous 
all through. 


96 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


they are traceable ; but though the vertical scale is more than twice 
the horizontal, it is not possible to show the details; for these 
we must refer to the vertical sections. 

Lower Headon Beds of the cliffs between Weston and Wraddick 
Chines.—We left off with the Potamomya-sands, 83 feet below the 
How-Ledge limestone ; these are easily traceable through the grass- 
slopes, exposures of a few feet occurring at intervals all the way to 
Widdick Chine, about 230 yards distant. There good sections are 
seen on each side of the roadway; here the sands have in the upper 
part more clay mixed with them, as in Warden Cliff. Whiter sands 
are below; above are alternating whity-brown sands with bluish 
silts. Melania turritissima occurs in the latter, a shell which occurs, 
indeed, in the Bembridge Marls at Hamstead, but which, in this 
district, we only know at one horizon, viz. the Unio Solandri bed, 
and one above that, at Warden Cliff and Colwell Bay; we remark 
its analogous position at Widdick Chine. Below the sands, again, is 
pale greenish clay, 3 inches ; in descending order, soft buft Limnea- 
limestone, 1 foot; brownish sands with Potamomya and reptile 
dermal ossifications : these occupy the position of the Crocodile-beds 
in the Lower Headon at Hordwell; they continue along the cliff as 
we walk northwards. Below is a carbonaceous band or impure lig- 
nite, 6 inches, then a repetition of clays with carbonaceous layers, fol- 
lowed by another Limnza-limestone, 10 inches, Potamomya-clays, 
4 feet, another Limnea-limestone, 1 foot, greenish clays with Palu- 
dina lenta, Potamomya plana, Melanopsis brevis, Limnea &e. ; 
another Limneea-limestone, 8 inches, full of Gyrogonites, below that 
clay with carbonaceous layers passing to drab sands, about 2 feet ; 
then a lignite layer and impure Limnea-limestone, soft and crumbling 
at the outcrop. The limestone full of Gyrogonites is noteworthy, as 
it occurs only low down in the Lower Headon, and serves to mark 
our position in the series at this spot. Now this Chara-limestone is 
exposed again at the back of the Reading Rooms, where there are | 
the same five limestones seen as near Widdick Chine, the Chara-bed 
being the lowest but one; this is well seen behind the Reading 
ltooms, where there has been a cutting through the Lower Headon 
beds for a new pathway. It will be noticed that we have passed 
five thin Limnea-limestones in the lower part of the Lower Headon 
in the cliffs immediately north of Widdick Chine, and again behind 
the Reading Rooms; we see them again as they rise from beneath 
the sea-level beyond the new pier under Warden Cliff; they are 
seen also in the recent scarping under Totland Bay Hotel. The 
continuity, then, of the section from the five lower limestones under 
Warden Cliff through Weston Chine to Widdick Chine is undoubted ; 
and from there we continue through the sands above to the How- 
Ledge limestone in Headon Hill. The beds in the cliff here belong 
entirely to the Lower Headon*. 

* The top of the cliff at the back of the Reading Rooms has a capping 
of about 7 feet of Post-tertiary sand ; at the base of this is a layer of flints and 
derived marine fossils, Cytherea incrassata, Ostrea velata, &e., showing that 


the marine Middle Headon series existed here above this level. This Post- 
tertiary sand lies on the (Lower Headon) Warden sands. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 97 


Lower Headon of Warden Cliff.—The lowest beds seen are below 
the Totland Bay Hotel at Weston Chine. We therefore start from 
here in making a measure of the Lower Headon series, meeting higher 
beds as we go north towards Warden Point. ‘The details of this 
measurement are given in the vertical section, fig. 2(p. 98); we need 
here allude to only a few of the beds. The lowest bed seen in 
Totland Bay is a few yards south of the Coast-guard boathouse ; 
here are visible, in ascending order, whity-brown sands, 13 foot, a 
hard purple sandy ironstone, 6 inches, greenish clay, 4 feet. 

In our vertical section, fig.2, we have drawn an interruption in these 
clays, because we are not absolutely sure that they are identical with 
the similar clay in the road-cutting at the pier-head on the other 
side of the boathouse; there is little doubt of it, however. At the 
pier-head are seen 3 feet of clay in the road-cutting (after our 
section was drawn, on a later visit, a drain-opening showed 41 feet) ; 
and from this point the section 1s continuous without interruption. 
There is therefore only a possibility of error of a few feet. 

Five thin Limnea-limestones will be noticed. The two lowest 
contain Chara-seeds at the pier-head; on following them to their 
outcrop along the shore it is found that the fourth of them also 
has Chara-seeds at that point; the fifth, 1 foot 3 inches thick, is 
50 feet. below the top of the How-Ledge limestone, or top of the 
Lower Headon. 

From the top of the cliffs these limestones may be seen at low 
water, forming five submarine ledges parallel to the great ledge at 
Warden Point; they strike N. 36° W. 

As there is great uniformity in these freshwater beds, and their 
fossils mostly occur in the Upper Headon also, we pass over many beds 
to notice the sand-rock bed, which is a conspicuous feature in Warden 
Chiff. This sandstone, somewhat calcareous at places and friable at 
others, forms Warden Ledge, and runs out at the top of the cliff 


_close below the flagstaff of the coast-guard station. 


About 11 feet above that comes the Unio Solandri bed* with 
Melama turritissema, a bed of which we have already noted a portion 
in the cliff at Widdick Chine; the MW. turritissima occurs through 
a greater thickness of the 114 feet of clays than does the Unio. 
Next comes the How-Ledge limestone, 5 feet, in two beds containing 
Limnea fusiformis, with a carbonaceous or lignitic band at the base. 
The shells are more crowded at the base; and the lower surface of 
the blocks fallen from the cliff is a sight pleasing to the collector 
of fossils. The dip of this bed in Warden Cliff, calculated from its 
horizontal extension on the 25-inch map, between its position at How 
Ledge at sea-level and a point of known elevation near the flag- 
staff, is a slope of about 1 im 45. This How-Ledge bed crops out at 
the top of the cliff, a little north of the coast-guard flagstaff; it is 


* 'The Unio-bed with Melania turritissima occupies an analogous position at 
Hordwell Cliff, being at about the same distance below the Lymnza-limestone 
(which is a diminutive representative of the How-Ledge bed), where it has pre- 
cisely the same lithological characters with the same abundance of black seeds 
(Carpolithes) as at Colwell Bay and Warden Cliff; it crops out again with the 
same fossils on the shore immediately south of Milford. 


98 H. KEEPING AND E. B, TAWNEY ON THE BEDS AT 


Fig. 2.—Vertical Section of Lower-Headon (freshwater) Beds eaposed 
between Weston Chine and Warden Cliff. 


(Scale, 12 feet to the inch.) 


Limnea-limestone, in two beds, buff colour. 


How-Ledge bed; extends from How Ledge to Coast-guard 
Station.—Z. fusiformis. 
= Lignitic or carbonaceous band. 


Pale grey clays, rather mottled. Paludina at intervals. 


but more so above. WMelania turritissima and other fos- 


{batons greenish and greyer clays, rather sandy below, 
sils in layers at intervals, 


Unio-bed. 
Laminated grey sandy clay, with Unio Solandri, Mel. turritissima. 


Stiff clays, grey or brown in places, 


=| Sandy grey clay, more sandy above, with Paludina lenta, Mela- 
nopsis brevis, Potamomya plana, Unio Solandrt. 


>:| Pale whity-brown crumbling sand-rock, - . 
‘| with strings of clay. } Cyr ena depen 


| Ferruginous sand-rock, variable in thickness, with Limnea, &e. 


Concretionary calcareous sand- 


rock, | 7; 
forms Warden Ledge. } wer alp BMSEIOCIL LG: 


LowER HEADON. 
eee 
on _ bo bo lor} ler) oO nn = 
(S) (=) Ne) (>) (=) fer) (=) =) = 


White and whity-brown sands, Potamomya in layers at intervals. 


10 0 
20 Stiff grey or sandy clay, with Potamomya crowded together. 
=, Lignitic band. 

S150 Greenish clay. Potamomya gregaria. 

1 3 Buff-coloured limestone, with Limnea. 
Lignitic or carbonaceous band. 

33-4 0 = Greenish-grey clays. 

0 6 Soft crumbling buff limestone, with Limnea longiscata. 
Grey marly clay. 

2 0 

0 8 ==| Bulf limestones, with Limnea. 


50 SS Greenish clay. 


[bed. 
= Buff compact limestone, with Chara-seeds (Gyrogonites) Chara- 
2:27] Whity-brown sand-rock, with lignitic band below. 

“| ,..Butt limestone, with a few Chara-seeds. 
-*-Carbonaceous layers. 


Green clay (behind wall at Pier-head). 
...Green clay, a little south of Coast-guard boathouse. 


=| ---Nodular purple-red ironstone band. 
<| Pale yellowish or whity-brown sands. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 99 


denuded away in the centre of Totland Bay. It cannot, however, 
have been many feet above the present cliffs near Widdick Chine, 
while a little south of that it exists in the cliff. At the N.E. corner 
of Headon Hill we noted it at a height of 70 feet above the sea ; it 
is well seen also in the cliffs near Heatherwood Point, where it has 
a somewhat higher elevation. 

There is evidence of an anticlinal in Totland Bay, as indicated in 
K. Forbes’s sketchy diagram ; the summit of the anticlinal we infer 
to be near the old wooden pier. 

If we add up, we obtain a thickness for the Lower Headon exposed 
in Warden Chiff of 714 feet (supposing no interruption at the point 
specified above). It remains to settle the relation of the lowest bed 
here seen to the Upper Bagshot Sands. 

The junction of the Lower Headon with the Upper Bagshot is well 
seen north of Alum-Bay Chine; immediately above the Upper Bagshot 
sands come greenish-grey clays ; then, in ascending order, alternation 
of clays and sands ; next, pale greenish-grey sands ; then a stiff marly 
clay: total 16 feet. Above is a purplish red clay-ironstone band; 
and succeeding that is the first thin Limnea-limestone. 

If we consider this red iron-band to be at the same horizon as 
that noticed under Weston Chine, as is extremely probable (though 
it is some 4 feet nearer to the lowest Limnza-limestone), then we 
must add 12 or 16 feet to the 71 feet obtained for the Lower Headon 
in Warden Cliff, making a total of 83 or 87 feet before we reach the 
yellowish sand of the Upper Bagshot. 

Knowing now the full thickness of the Lower Headon, we are 
able to test the argument as to the position of the Upper Bagshot, or 
Headon sands, as they were once called by E. Forbes (Mem. p. 34-6), 
in Totland Bay. It is stated (op. at. p. 147) that the Survey 
actually represent these as occurring, in both the Vertical and Hori- 
zontal Sections, near the summit of the anticlinal in Totland Bay*. 

There seems a little inconsistency in the Survey Vertical Sections 
concerning both boundaries of the Lower Headon; if we may classify 
these green clays in the Lower Headon, and then read off by scale from 
the top of the How-Ledge bed, the Survey Section would give a thick- 
ness of 85 ft. for the Lower Headon in Totland Bay. 

It is urged “that the Headon-Hill sands do not occur in the 
position indicated by the Geological Survey ;” and the crucial test of 


* We pointed out above that the Survey Vertical Section [edition 1870] shows 
them only just above the sea-level at a point some way inland. The Vertical 
Section we are inclined to interpret in this way—that the beds denoted Upper 
Bagshot in Totland Bay are what we have classed in the Lower Headon ; for the 
legend states, below the “‘clay-ironstone ” are 6 inches sand in “ Totland Bay,” 
then “ green clay with lenticular patches of sand” 15 feet, so that the de- 
scription agrees fairly with our lowest bedsat Heatherwood Point. The Survey 
Section does not state that these were exposed in the centre of Totland Bay; but 
as Heatherwood Point is the western point of the bay, we may assume that 
they may have been seen anywhere short of that point along the base of Headon 
Hill, where it is quite certain that they exist, and must have been open at that 
time at more points than one; for the white glass-house sands were then being 
actively worked in Headon Hill, and the yellower sands above them may still 
be seen about a mile from the N.E. corner of Headon Hill. 


100 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


the excavations at the new Reading Rooms is supposed to prove 
“‘ that beds are [there] found which have their exact counterpart in 
the Headon-Hill section, not at the base, but at a much higher part 
of the series.” We can readily understand that the Upper Bagshot 
sands were not found in the excavation; we could even have 
predicted the fact beforehand ; from a rough calculation, we estimate 
that the sands are above a dozen feet below the foundation. In 
favour of this view, we may add that, when the old wooden pier was 
being made, one of us heard from those engaged in the work, that 
the piles were driven with difficulty because of their piercing solid 
sand! *, 

But putting aside the question as to their exact depth here below 
the Reading Rooms, we are able to refute the notion that any thing 
higher than Lower Headon beds exist here. As mentioned above, we 
recognized immediately behind the Reading Rooms the five Lymnea- 
limestones which come below the Potamomya-sand, both at Widdick 
Chine on the south, and Weston Chine on the north, between which 
places the sands may be traced almost continuously. The whole of the 
cliffs between Weston and Widdick Chines are occupied solely and 
throughout by Lower Headon beds (neglecting the cap of Post- 
tertiaries). All this part of the section (op. ct. pl. vu. f. 2) is in- 
accurate, in consequence of the Middle Headon being placed too low 
in Headon Hill. 

Middle Headon of Warden Cliff—Again, we cannot agree with 
that part of the section between Weston Chine and Warden Point. 
Here no marine bed is indicated ; for in fig. 3, the section drawn to 
true scale, the Colwell bed is made to die out before the Warden 
battery is reached, which is occupied by an exaggerated thickness of 
gravel 7. 

There is no fact more patent to any observer than that the Col- 
well-Bay marine bed extends all through Warden Point and Cliff, 
where it is supported by the How-Ledge limestone. We made a 
measure in detail of the Colwell marine bed (Middle Headon) at a 
point about midway between Warden Battery and Weston Chine ; it 
is here 344 feet thick ; we noticed there the Neritina-bed + with its 
characteristic features and fossils below the Venus-bed as at Col- 
well Bay and Headon Hill. It would be wearisome to give all the 
details ; but the Colwell bed here is easily recognizable as identical 
with the Middle Headon of Headon Hill, both physically and pale- 
ontologically. At Warden Battery, above the Middle Headon, comes 
some Upper Headon § besides the Post-tertiary cap. 

In the section drawn to true scale (pl. vu. fig. 3) the Colwell- 
Bay marine bed has its horizontal extension curtailed by almost one 


* The iron columns of the new pier are stated to pierce a bluish clay; we 
should interpret this to indicate the clays immediately above the Upper Bagshot 
Sands, and which are described as greenish in the Survey section. 

t We cannot reconcile this with fig. 2 of the same plate, where the marine 
bed is more nearly correctly drawn. 

{ Previously noted as bed 16 by Dr. Wright, Proc. Cotsw. Club, i. p. 95, and 
Ann. & Mag. Nat. Hist. s. 2, vol. vii. 

§ Described by Dr. Wright in 1850 as bed 5, Proc. Cotsw. Club, i. p. 90. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 101 


half; the result is that the dip is considerably exaggerated. Again, 
in Headon Hill an exaggeration of dip is produced by the error of 
105 ft. in plotting the marine beds. 'The effect is that the identical bed 
(Middle Headon) is split into two beds separated by 120 ft. of beds. 
From our point of view this could only be done by counting more 
than 100 ft. of beds twice over. In the legend attached to the diagram 
section fig. 2, “‘ new interpretation,” the beds g are, in our opinion, 
the Lower Headon; f is the Middle Headon ; e the Upper Headon ; 
d and ¢ are the Lower Headon; 6} is the Middle Headon again. 
According to the old view, which we certainly should prefer, this last 
105 ft. has no existence in fact*. 

Lower Headon of Colwell Bay.—The section in Colwell Bay is 
continuous with that of Warden Cliff; but in the bay, as we go 
north, a few lithological changes occur in the marine beds 7, as 
noticed below, and which cause the marine beds at one part of Col- 
well Bay to differ far more from the same beds at the centre of the 
bay than the latter do from the marine beds of Headon Hill. On 
rounding Warden Point, beyond the sea-wall, is a small rifle-target ; 
and from here the beds are fairly well exposed throughout the bay, 
though tumbled portions or a diminutive undercliff may conceal 
some of the beds in places, sufficiently to give considerable trouble 
in measuring the beds. ° 

The Unio-Solandri bed with Melania turritissima has been fre- 
quently worked by one of us below the How-Ledge limestone here 
but this summer we could only find tumbled portions of it. The said 
limestone rises from beneath the sea-level at How Ledge +, whence 
its appellation, and crosses Colwell Chine; here and at the target it 
has the same lignitic band and clays beneath it as on the south side 
of Warden Cliff; it thins down to 8 ft. north of the chine. We 
mention these upper beds of the Lower Headon to show that the 
Colwell-Bay marine bed, as at Warden Cliff, reposes on the same 
succession of beds as in Headon Hill. 

Middle Headon of Colwell Bay.—The lowest or Neritina-bed at 
the S.W. end of the bay, by the target, is now covered by tumbled 
matter, but is well seen a little further on about 50 yards short of 
Colwell Chine. Here, 7. ¢. between the target and the chine in the 

* The diagram in Forbes’s posthumous work is so schematic that it omits 
the higher part of Headon Hill, and, perhaps for clearness sake, the effect of 
the anticlinal is exaggerated. It is rather severe to treat it as if it were drawn 
to scale, and, because the Upper Bagshot Sands were brought up too much in 
the centre of the roll, to say that H. Forbes was “‘mistaken in his interpretation” 
(op. cit. p. 176) of the beds. Forbes’s diagram, in fact, with this qualification, 
represents the beds in their right position ; thus the Lower Headon, no. 6, occu- 
pies the summit of the cliffs in the centre of Totland Bay, while no. 7, Middle 
Headon, is denuded from above it—all which is perfectly correct. 

Tt Since Warden Battery has been built it is forbidden to search for fossils 
on the slopes at Warden Poimt. Many years ago one of the authors was in the 
habit of frequently exploiting the beds here for fossils, and many of them are 
incidentally described by Dr. Wright, Proc. Cotsw. Club, i. pp. 91, 92 (1850). 
The engineers, however, have not succeeded in grassing all the slopes, and fallen 
fossils may still be picked up at the base. 

_ ¢ Bed 18 of Dr. Wright, who describes it correctly in Warden Cliff, but 
appears to have mistaken its position in Headon sub (2b. p. 95). 


Q.J.G.S8. No. 146. I 


102 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


Marine series above the Neritina-bed,is some foxy-coloured sand- 
rock, while lenticular layers of white sand are seen thinning out in 
the grey clays, which also contain red-clay ironstone nodules, a dif- 
ferent lithological condition from what occurs further north in the bay, 
or to the south of Warden Point. Above comes the richest part of the 
Venus-bed with abundant Cytherea incrassata ; these fossils not only 
strew the tumbled clays, but with other fossils are commingled by the 
waves with recent shells on the strand. Atthis point of the bay Ostrea 
velata is very abundant above the part richest in Cytherea. 

In the centre of the bay between Colwell and Bramble Chines | 
this oyster is in extraordinary abundance; at one place it nearly 
crowds out most of the other fossils, and forms a massive oyster- 
bank about 21 ft. thick *, of which the lower 12 ft. is almost 
made up of these shells. The ordinary character of the Venus- 
bed is quite altered here, though its fossils occur mixed up with 
the oysters; we may notice as especially abundant Murex seaden- 
tatus, Pisania labiata, Natca labellata, Nerita aperta, Cerithium 
variabile, Ostrea velata, Nucula headonensis, Cytherea merassata, &c. 
On either side of this spot, near both How Ledge and Bramble Chine, 
the excess of oysters has disappeared, and they are chiefly abundant in 
a zone above the richest part of the Venus-bed, though they do occur 
sparingly throughout. Cytherea merassata occurs through several 
feet, but the richest part of the Venus-bed consists of about 9 inches 
of bluish-green sand ; in this the shells are found in the best state 
of preservation. Above the grey and greenish-grey sandy clays 
richest in marine forms are some pale bluish-green clayey sands, 
between Bramble and Linstone (or Lynchen) Chines ; in these, at 
the level of about 5 feet from the base, is a band very rich in Ceri- 
thium ventricosum, C. variabile, Melania muricata, Corbicula obovata, 
with, occasionally, Verita ; this bed is seen just beyond the spring at 
Linstone Chine, it is only a few feet below the base of the Upper 
Headon. We wish to draw attention to the first-named fossil, as it 
occurs also near the top of the marine beds at Headon Hill, 7. ¢. in 
quite an analogous position. 

Upper Headon of Colwell Bay.—The slate-coloured grey clay with 
Potamomya ~ (ammediately succeeding the bluish-green sands) we 
take as the base of the Upper Headon. A detail vertical section 
is given (fig. 3), in order to show the lithological differences 
existing between the series here and equivalent beds in Headon Hill. 
We do not, however, think them greater than the differences already 
noticed as occurring in the marine series in different parts of Col- 
well Bay, while the resemblances are sufficiently great to allow of 
their perfect correlation—not to mention their position between the 
Osborne beds above (so identical with the Osborne beds of Headon 
Hill) and the marine series below, which we have shown, on strati- 
graphical grounds, is most certainly identical with the Middle Headon 
of Headon Hill. The differences consist here in a greater develop- 
ment of sand and a reduction of limestone: the sands at the base 


* Bed 12 of Dr. Wright (zd. p. 92), who, however, much underrates its thickness. 
t Base of bed no. 5 of Dr. Wright (2d. p. 91). 


HEADON. 


UPPER 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 103 


Fig. 3.— Vertical Section A Upper Headon Beds in Colwell Bay. 
(Scale, 8 feet to 1 inch.) 


Paludina lenta, Pota- 


Light-grey clays, with layers of shells. ; momya gregaria, 8c 


Pale brownish-grey and light-brown clays in alternations. 


ft. in 
15 0 
5 : _ 3. § Corbicula obovata, Cy- 
0 6-9 Nodular band offerruginous sand-rock.) “7 Ty» Sete 2 x 
Whitish sand above, yellow and ORE NYRI, I MAE 
24 eee eee ears z lenta, Corbicula obovata, 
Torn y ; Melanopsis fusi ormis. 
Th al a " { Mi) | Buff-coloured TZimnea-limestone, with carbonaceous layer 
li th ali | below. 
16 > Pale greenish-grey clays. 
SS Zone of Cerithium trizonatum. 
———————=| Pale greenish clay. 
ew == Ferruginous sand layer. 
== Paler sandy clays. 
Cyrena Wrightii, Corbicula obo- 
io Stiff slate-coloured grey clay.4 vata, Melanopsis fusiformis, 
= Paludina lenta. 
5 6 Varying alternations of paler and darker grey clays. 
= : Pale buff-coloured sands, with lignitic or carbonaceous bands. 
15 0 2 
rede ssene ts 
re, ee 2 ee or | Greyer sands below. Potamomya, 
a roonn 
ee. aa: AO ee 
otal’... 463 feet 


Bo 


104 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


are considerably thicker here than at Headon Hill; their character 
is very variable even in different parts of Colwell Bay. The 
Limnza-limestone has here a much reduced thickness, as mentioned 
previously. 

Among the other beds we may specially notice the horizon of 
Cyrena Wrightw; it occurs in the nodular ferruginous sand-rock 
with Corbicula obovata, also in the sand immediately above the 
Limnza-limestone, and again in some of the clays below. Another 
fossil that we have only observed at one horizon is the Cerithium 
trizonatum (Morr.); it occurs in the pale greenish clays beneath the 
limestone, occupying only a narrow band in these clays. Equally 
characteristic is Serpula tenuis (Sow.), which occurs at the same 
horizon both here and at Headon Hill, viz. in the Upper Potamomya- 
clay just above the limestone. 

The Paludina-clays at the top are identical with those of Headon 
Hill at the top of the thick limestones. Measurement by tape here 
gave 15 feet ; total of the Upper Headon near Cliff End 46} feet. 

Osborne Beds of Cliff Hnd.—The red and greenish mottled marls 
of the Osborne series follow. These beds show for afew yards only, 
and then become hidden under the grass which the engineers have 
grown on the artificial slopes below the battery. One of us well 
remembers the numerous little faults (14 are enumerated by the 
Survey, as cited above) which repeated the Limnza-limestone. 
Beyond the battery the Osborne beds form the tumbled cliff; a 
measurement is no longer to be made with profit. Mr. Bristow 
gives 62 feet for the series here. 


III. PanxontToLogicaAL EVIDENCE. 


The question now arises, Does the distribution of fossils bear out 
the separation of the Colwell-Bay and Headon-Hill marine beds 
and their reference to different horizons ? and does it sanction the 
notion of the Brockenhurst bed being equivalent to the Colwell-Bay 
bed ? 

Two lists of fossils are laid before us, viz. one which mixes up the 
fossils from the Brockenhurst, Whitecliff Bay, and Colwell-Bay 
localities, and the other which gives those from the marine beds of 
Headon Hill and Hordwell Chiff; of the hundred forms (in round 
numbers) which occur in the latter list, it 1s said (op. cit. p. 150) 
‘less than one half occur at the other three places.” 

We may urge at the commencement that it comes rather near to 
begging the question to mix up Colwell-Bay with Brockenhurst-series 
localities. We conceive one of the main points in dispute to be 
whether the Colwell-Bay bed has any more affinities with the 
Brockenhurst fauna than has the Headon-Hill bed; and to this sub- 
ject we shall address ourselves after we have first compared the 
fauna of the Colwell-Bay and Headon-Hill marine beds. 

The first thing to be done is to separate the faunas of all the 
localities which are to be compared together; this we have done in 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 105 


the lists at the end of this communication. These have been drawn 
up from an inspection of the Edwards collection of Tertiary fossils 
in the British Museum ft, while we have added many occurrences 
from our researches this summer, and we believe our list to be fairly 
correct. 

We do not admit that the Edwards collection is sufficient for a 
full knowledge of the distribution of English Tertiary fossils: ¢.g. if 
a certain shell does not exist in the Edwards collection from Colwell 
Bay, it is no proof that it does not occur there, but simply that 
Mr. Edwards had better specimens of it from Headon Hill, which 
was considered, as, indeed, it is, an equivalent bed. The greater 
part of the Edwards collection was made by the hands of one of the 
authors; but we do not think it possible to establish from this col- 
lection that Colwell Bay contains so many marine forms which do 
not occur in Headon Hill, and therefore that the marine beds at these 
localities are not on the same horizon. 

The following reasons induce us to say this:—It was the habit of 
the local collectors to exploit the Colwell-Bay bed far more than the 
Headon-Hill locality, because it was more accessible and showed 
a larger extent exposed ; the fossils were better preserved, and in 
every respect it was easier to collect from. Again, some species 
which one of us supplied to Mr. Edwards from Headon Hill were 
not incorporated in this collection, probably owing to his having 
better ones from Colwell Bay; and doubtless they were exchanged 
with foreign correspondents or given away. These two con- 
siderations would serve to account for his Colwell-Bay collection 
being richer than his Headon-Hill one. Corroborative of this is 
the fact that in a few days’. search this summer we have found 
several species in the marine bed at Headon Hill which do not exist 
in the Edwards collection from that locality. 

Moreover we hold that the best test as to the contemporaneity 
of these beds is not to be obtained from the rarer forms, which may 
be evidenced only by a single example, but from a comparison of the 
commoner, which we should consider the more characteristic forms. 
Accordingly we add a list of the species obtained by us this summer 
from the Middle Headon of both localities: H indicates Headon 
Hill; C the equivalent bed at Colwell Bay. 


List of all the Shells obtained on the ground by the authors for the 
purposes of this paper (Aug. 1880). 


(Those with an asterisk pass up from Barton beds.) 


*Lamna contortidens (4g.) ... C,H. | *Borsonia sulcata(fouw.)......... C. 
Marginella vittata (Hdw.)...... H. | *Rimellarimosa (Sol.) ......:.. OVE 
*Voluta spinosa (Lam.) ......... C, H. Murex sexdentatus (Sow.)...... Opal 
or otoma headonensis (Hdw.).C, H. BPs Seen eee Cees on awetces lek 
denticula (Bast.), var. Pisania labiata (Sow.) ......... CO, H. 
odonbellat(Hd.) os. 5.00.42 C,H. | *Cominella flexuosa (Hd. MS.) O, H. 


t We are greatly indebted to the Keeper of the Geological Department, Dr. 
H. Woodward, F.R.S., for his courtesy in allowing us access to the collection, 
even during the laborious process of packing up and moving the national col- 
lection to the new buildin g at South Kensington. 


106 


H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


List of Shells (contenued). 


*Ancillaria buccinoides (Lam.) C, H. Planorbis euomphalus (Sow.) C, H. 
*Natica Studeri (Bronm)......... C, H. obtusus (Sow.) .........00. C,H. 
labellata (Lam.) ......... @), dal Paludina lenta (Sow.) ........- C,H. 
Cancellaria muricata (Wood) C, H. Ostrea velata (Wood) ......... C, H. 
elongata (Nys?)..........+ C,H. | *Mytilus affinis (Sow.) ......... C, H. 
Odostomia gracilis (Hd. MS.) C. *Trigonccelia deltoidea (Lam.), OC, H. 
Scalaria tessellata (Hd. MS.)... C. Leda propinqua (Wood) ...... C, H. 
*Cerithium yariabile (Desh.) ... C, H. Nucula headonensis (forbes) C, H. 
we pseudocinctum (d’ Ord.) CO, H. nudata (Wood)... <2... C, H. 
duplex GSow:) ae. sees H. Lucina (Strigilla) colvellensis 
ventricosum (Sow.) ...... OC, H. (Ed « MS.) 05 occ eeceeeeeee C,H. 
varians (Hd. MS.) ...... C,H. | Cytherea suborbicularis 
—— (Vicarya) concayum (Od MIS. Vins Sine eee C,H. 
(SOU) ca tn owilnvecturboaecmees Cons incrassata (Sow.) ........- C, H. 
Melania fasciata (Sow.)......... C, H. Tellina headonensis (Hd. MS.) OC, H. 
muricata (Sow.) ......... C,H. | [*]Psammobia compressa (Sow.), 
Hydrobia bulimoides (Hd. MS.) C, H. var. estuarina (Hd. MS.)... C, H. 
Melanopsis subfusiformis rudis: (Lane) eee eee C, H. 
(GUGIS anecion waoaatocadoasec SG, Ee Mactra fastigiata (Hd. MS.).,. C,H. 
fusiformis (Sow.) ......... C,H. | *Mya angustata (Sow.)=pro- 
Acton dactylinus (Ed. MS.) . C. ducta (2d MiS)) ieee eee C,H. 
Nematura parvula (Desh.) C,H. | *Corbula cuspidata (Sow.) ...... C,H. 
Nerita aperta (Sow.) .........5 55) (O4dab Corbicula obovata (Sow.) ...... CoE 
zonula) CHW 00d) <...2-2ke-- C,H. Cyrena cycladiformis (Desh.). C, H. 
Neritina concava (Sow.) ...... C,H. | *Balanus unguiformis (Sow.). C, H. 
Bulla estuarina (Hd. MS.) ... CO, H. Callianassa Baylu (Woodw.). C,H. 
Limneea longiscata (Sow.)...... C, H. 


Faunas of Middle Headon from Colwell Bay and Headon Hill 


compared. 


The above list contains only the commoner forms, such as may be 
found in a few days’ search. Out of a total of 58 species it will be 
observed that all but 7 were found by us in both localities *, and all 
but three are known to be common, or, again, a proportion of 94 per 
cent. of commoner Colwell-Bay forms occur.at Headon Hill. Surely 
from this we may presume a very close .affinity if not idence of 
these beds. It is stated (op. cit. p. 150) that less than one half of 
the Headon-Hill and Hordwell species occur at Colwell Bay—a 
result, it seems to us, only obtained by mixing up fossils from 
Brockenhurst and Whitecliff Bay in the same list with the Colwell 
forms. We shall show below that the fossils cited from these two 
latter localities belong to a lower zone. 

Next as to the statement that at Colwell Bay “ the strata are of 
purely marine origin” while “ the so-called Middle Marine beds of 
Headon Hill and Hordwell Cliff are of totally different character ” 


* Some of these do not exist from both localities in the Edwards collection, 
val have not found their way into the Headon-Hill.and Hordwell list in the 
paper referred to, though previously cited by Dr. Wr ight: from Hordwell 

ib. p. 124). 
~ Comparing the whole known fauna from the Middle laid of Colwell Bay 
and Headon Hill, we obtain the following result, viz..74-per cent. of the 
Colwell-Bay marine forms have been found at Headon Hill. ‘This is counting 
as separate species many names in the Edwards collection which are founded on 
imperfect and single specimens. As we have said, we think a surer guide in 
comparing faunas is to take only the characteristic and less rare species. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT, 107 


&e. (l.¢. p. 148). Of the brackish-water genera which are supposed 
to be found in Headon Hill only, we may remark that we found 
Cerithium, Cyrena, Hydrobia, Liamnea, Paludina, Planorbis, Melania, 
and Melanopsis fully as plentiful m the marine series of Colwell 
Bay as at Headon Hill; e.g. in a quarter of an hour we turned out 
half a dozen specimens of Limnea longiscata* from the richest 
nine inches of the Venus-bed, the best zone for Voluta, Cancellaria, 
Murex, Cytherea, &c. It has always been the opinion of one of us, 
who has worked these beds for so long, that these freshwater forms 
were either drifted down by flood-waters or were dead shells washed 
out of lacustrine or brackish deposits. They cannot have lived in the 
waters depositing the marine bed at Headon Hill any more than at 
Colwell Bay. 

Another argument brought forward in opposition to the views of 
the Geological Survey is, that certain species of Cerithiwm are 
confined to Headon Hill and do not occur in Colwell Bay; and by 
this means have been “ detected the serious errors which have crept 
into our classification and correlation of the strata we are now con- 
sidering” (op. cit. p. 149). Cerrthiwm ventricosum and C. concavum 
are said to be entirely confined to the Headon Hill and Hordwell 
localities. We cannot agree with the statement as to the distri- 
bution of C. ventricosuwm in the Headon-Hill beds and its “* prodigious 
abundance.” It is there, as far as we have observed, found only in 
one bed; moreover, it is equally abundant in a bed in a precisely 
similar position at Colwell Bay, viz. at the top of the Middle Headon. 
Its analogous position in these two localities we consider as fossil 
evidence confirmatory of the stratigraphical. 

Nor do our observations confirm the statement of “ prodigious 
abundance ” of C.(Vicarya) concavum at Hordwell Cliffin the Middle 
Headon. One of the authors who worked that bed when a special 
excavation was made for the purpose‘, considers that V. concava 
was extremely rare in the Hordwell bed; but, as is well known, it 
occurs abundantly in the Upper Bagshot sands further west at Long 
Mead End. 

As to the supposed absence of V. concava from Colwell Bay, we 
remark that we had not been many minutes at work on the richest 
portion of the Venus-bed before we found a specimen, subsequently 
followed by a dozen more. It can scarcely be maintained, therefore, 
that the Colwell-Bay bed does not belong to the C.-concavwm zone.» 
This species is here, however, not so common as at Headon Hill +. 

* Also noticed by Mr. Bristow, F.R.S. (Mem. 10*, p. 61), as well as by previous 
writers. 

t The Middle marine or Middle Headon bed at Rook Cliff, Hordwell, has not 
been exposed for the last twenty-eight years ; it is covered up by a great thick- 
ness of gravel, and its precise position is known but to few geologists. It was 
quite a thin bed, but rich in fossils, especially minute forms. Fossils in 
existing collections were all obtained about a quarter of a century ago. 

{ This species exists, however, in the Edwards collection, labelled as from 
Colwell Bay. The absence of a shell in the Edwards collection from Colwell 
Bay is no proof that it did not occur there; the local dealers might not have 


thought of picking up V’. concava at Colwell Bay. For this species they went 
to Headon Hill, where it was more abundantly found and in better preserva- 


108 H, KEEPING AND E. B. TAWNEY ON THE BEDS AT 


Summary.—There is but one marine bed here, namely that in 
the Middle Headon ; for the Colwell-Bay bed can be traced strati- 
graphically into the Headon-Hill Venus-bed, and the paleontological 
evidence 1s in harmony with the stratigraphical. The place of the 
Brockenhurst bed is at a lower horizon in the Middle Headon ; but 
it does not appear anywhere in the west end of the island. 


TV. Watreciirr Bay anp New Forest. 


Middle Headon of Whitechff Bay—We next have to raise a 
more serious objection to the way in which the Whitecliff Bay 
section has been interpreted. In mixing up all the beds in the 
marine series there together and calling them Brockenhurst series, 
it seems to us that the question has been obscured, if not begged. 
The statement is that the 100 feet of marine beds at Whiteclifi Bay 
are the equivalents of the 25-feet of marine beds at Colwell Bay and 
of the beds in the New Forest with the Brockenhurst fauna (op. cit. 

148). Hence the Colwell-Bay bed is placed in the Brockenhurst 
series, which is said to occupy a higher horizon than the Headon- 
Hill and Hordwell marine bed; and this view is indicated by dotted 
lines in the vertical sections on p. 170. Since the 100 feet of 
marine beds are classed together and called *‘ Brockenhurst Series,” 
we suppose that the Brockenhurst fauna is imagined to occur 
throughout them. Asa matter of fact, that fauna is confined to 
one zone, and that the very base of the series. 

Though we have worked over this part of the section bed by bed, 
we need not here give all the details, but will refer to the description 
of it on the Survey Vertical Section on Sheet 25. This series is 
there justly referred to the Middle Headon, since it lies between the 
freshwater Lower and Upper Headon, its total thickness read off 
by scale being 90 feet. At the top are clayey sands and yellow 
sands about 19 feet; then the “ Venus-bed” clays &c., 15 feet; 
next, below, are compact sands with nodules about 42 feet, said to 
contain Sanguinolaria Hollowaysii*; then come 14 feet of brown 
clays, the base ‘‘ greenish and brownish clay, very fossiliferous.” 
Now there is no doubt at all about the bed above and the Venus- 
bed here being any thing but the Colwell-Bay and Headon-Hill 
marine bed—its position and its fossils prove that; the characteristic 
Brockenhurst fossils are absent from it, and it is therefore certain 
that it differs entirely from the Beeeonheret beds. 

The Sanguinolaria-sand differs lithogically from the lower part 
of the marine series, both at Colwell Bay and at Headon Hill. Such 
few fossils as we observed in it are distinctive, not of the Venus- 


tion. Moreover these collectors sought to obtain as many forms as possible, 
but were not concerned in finding the same species in both localities; and if 
they found only a few stray examples at one locality of a species of which they 
had a great number from the other, they were liable not to put a separate label 
for the odd few, but to mix them up with the larger parcel. 

* A wrong determination; the shell is Psammobia compressa, var. estuarina, 
Kd. MS. It occurs in the natural position of life, 7. ¢. across the bedding. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 109 


bed, but of Brockenhurst beds; but though we have not sufficiently 
worked out this fauna, we may say that we are satisfied that they 
belong to a lower zone than any of the marine beds at Colwell Bay 
or Headon Hill, the Middle Headon being more fully developed at 
Whitecliff Bay than elsewhere. 

Brockenhurst Zone at Whatecliff Bay.—The succeeding 14 feet 
are the equivalents of the Brockenhurst beds; the lowest two feet 
we shall call the Brockenhurst zone ; the remainder of the thickness is 
not nearly so rich in species, and their grouping, as well as the litho- 
logical character, is more lke that of the Roydon beds. 

At the time the Survey section was made, the interesting bed at 
Brockenhurst had not been discovered nor its fauna described; 
hence such Brockenhurst fossils as were found in this zone here 
were not rightly determined (thus in the Survey section we must 
read Cardita deltoidea, Sow., for C. acuticosta), or specific names 
were withheld from them. Subsequent observers* have recognized 
the Brockenhurst fauna in this lowest bed. As we have obtained 
more fossils from it than previous observers, we have embodied our 
results in a separate column in the lists at the end of this essay ; 
that column contains nothing except what we have collected with 
our own hands this summer from the lowest two feet +, lying on an 
eroded surface of the freshwater Lower Headon. Comparison of this 

list with the fauna from Brockenhurst itself will convince most, we 

think, of the perfect equivalence of the zone in the island and in 
the forest, while its position at Whitecliff Bay shows that it is at 
the base of the Middle Headon. 

Brockenhurst Zone in the New Forest.—The greater part of the 
fossils from Brockenhurst were collected by the hands of one of 
the authors, and thence were dispersed into various public and 
private collections. They were obtained during the doubling of the 
line and widening of the cutting at Whitley Ridge, near Brocken- 
hurst t, about twenty-three years ago. During this work he had 

- the advantage of seeing more of the beds than any other geologist. 
He found the rich Brockenhurst zone (which varied from a few 
inches to nearly a foot) lying immediately upon the freshwater Lower 
Headon ; while about half a mile up the line, near the bridge by 
Lady-Cross Lodge, the Middle Headon Venus-bed was seen, followed 
by the freshwater Upper Headon beds above, the beds having a very 
gentle dip up the line oreasterly§. It is evident that the succession 


* Videlicet Von Konen, Quart. Journ. Geol. Soc. vol. xx. p. 98; Rev. O. Fisher, 
Quart. Journ. Geol. Soe. vol. xviii. p. 67, footnote; Mr. T. Codrington, Quart. 
Journ. Geol. Soc. vol. xxiv. p. 519; Dr. Duncan, Pal. Soc., ‘ Fossil Corals,’ i. 
p- 40 (1865). 

t In the Edwards collection the label “ Whitecliff Bay” includes many 
Venus-bed forms, indeed Lower and Upper Headon, or it may be any thing 
from the London Clay to the Bembridge Marls; there is therefore good reason 
for not allowing this collection to stand as evidence of what is found in the 
Brockenhurst zone at Whitecliff Bay. 

{ The railway-cutting at Brockenhurst (op. cit. p. 152) refers to the same 
spot as Whitley Ridge. 

§ We visited the New-Forest localities together this summer, and found the 
Whitley-Ridge cutting entirely grassed over (the rich zone was below the level 


110 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


here is, in ascending order, freshwater Lower Headon, Brockenhurst 
zone, Venus-bed, then freshwater Upper Headon, which agrees 
with the succession in Whitecliff Bay. Yet in the New-Forest 
section (op. cit. p. 170) we find the Brockenhurst bed placed above the 
marine band or Middle Headon of Totland Bay—in other words, 
the natural succession is inverted. 

Again, in Headon Hill as we have seen, an imaginary Brocken- 
hurst bed (of which the Colwell-Bay Middle Headon is stated to be 
the equivalent) is placed above the Upper Headon, in ground which 
is really occupied by the Osborne beds. If the Brockenhurst bed is 
at a higher horizon than the Middle Headon of Headon Hill, then 
where is the marine Middle Headon at Whitecliff Bay ? 

We can scarcely adopt a theory which makes the Colwell-Bay bed 
occupy a higher horizon than that of Headon Hill, because it is 
supposed to contain more Brockenhurst fossils, when the latter 
fauna is found below the zone with (Colwell-Bay or) marine Headon 
fossils both at Whitecliff Bay and near Brockenhurst. When once 
the position of the Brockenhurst fauna is recognized (and it has 
been correctly described by previous observers), the inconsistency of 
the theory is apparent, 

Affinities of the Brockenhurst Fauna.—Seeing that the Brockenhurst 
fauna, if different in age from the Marine Headon, is older, instead 
of being younger, it would be rather anomalous to find that “ while 
nearly one third of the Hordwell and Headon-Hill marine shells are 
Barton forms, not more than one fifth of those occurring at Brocken- 
hurst, Colwell Bay, and Whitecliff are found at Barton.” We have 
already mentioned one feature in the lists on which this statement 
is based by which the question is almost begged. We must next 
allude to what seem to us clerical errors, in order to justify the 
very different statistics which we have obtained by inspection of 
the Edwards collection, supplemented by our own researches. 

In the Headon-Hill list we observe nine species* that are said 
to pass down into Barton beds, while in the Brockenhurst list this 
range is denied to them; and besides these nine, the range into 
Barton, as proved by the Edwards collection, is omitted in the Brock- 
enhurst list in the case of twenty-two other species. Discrepancies 
of this sort must seriously detract from the value of any statistics 
based on such lists. 


of the rails, and will never be seen again here); the upper beds were yellowish 
clayey sands, poor in fossils. Sufficient characteristic Brockenhurst fossils may 
still be seen, however, on the old spoil-banks of the date of the making of the 
original single line, about forty-two years ago. By Lady-Cross Bridge the 
cutting is also grassed over ; but evidence can still be found of the Venus-bed 
in the side drains and of the Upper Headon in the slopes above it. 

* These species are—Borsonia sulcata, Nematura parvula, Mytilus strigil- 
latus, Cardium obliquum, Trigonocelia deltoidea, LIncina obesa, L. concava, 
Panopea subeffusa, and Scintilla angusta. On the other hand, an error on the 
opposite side, omitting the range into Barton beds in the case of Marginella 
simplex and Corbula cuspidata, goes only one quarter of the way towards 
redressing the balance. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. pa 


So far from the Brockenhurst zone having fewer forms common 
to the Barton than the Headon-Hill marine bed, we consider that 
it has rather more, as its position at the base of the Middle Headon 
at Whitecliff Bay would lead us to expect. Thus, if we take the 
whole Brockenhurst fauna (including the 15 corals which are mostly 
special to the zone), we obtain a total of 151 species, of which from 
74 to 81 pass up from Barton, or a proportion of about one half. 
Summing up the Headon-Hill forms in the same way—out of a 
total of 79 species, 23 pass up from Barton beds, or a proportion of 
29 per cent. But, instead of including the rarer forms, if we take 
only the more characteristic and abundant species of the Brocken- 
hurst zone, it would be perhaps a preferable course. 

The following list is a catalogue of the fossils in the Woodwardian 
Museum from the Brockenhurst zone, obtained by one of us many 
years ago at Whitley-Ridge railway-cutting, New Forest; and it 
may be taken to include the chief characteristic fossils of the zone. 
We have found all, except two, in the 2-feet bed at Whitecliff Bay 
this summer. 


xInfundibulum trochiforme = obli- 
quum (Sow.). 


*Hippocrenes (Rostellaria) ampla. 
*Rimella rimosa. 


Murex hantoniensis (Hd. MS.). 
xTyphis pungens, 
*Strepsidura armata. 
Cancellaria muricata. 
Pisania (Fusus) labiata. 
*Clavella (Fusus) longeeva. 
Leiostoma ovatum, 
xCassis ambigua. 
Ancillaria buccinoides. 
Pleurotoma transversaria. 


cymezea. 
headonensis. 
¥ denticula. 
% pyrgota. 
oo decora (Beyr.) = maga 
é). 
* spinosa. 
suturalis. 
geminata. 


xActezon simulatus. 
Marginella estuarina. 
xNatica hantoniensis. 
* Studeri. 
x— , var. grossiuscula (Hd. 
MS.). 
x—— labellata. 
Chenopus Margerini, var. speci- 
osus, 


Phorus eretifer (Ed. MS.). 
Ostrea ventilabrum = prona (S. 
Wood). 
xAnomia tenuistriata. 
Pecten bellicostatus, 
Modiola Nysti. 
xAvicula media. 
xLucina bartonensis (Ed. MS.). 
x Cardium porulosum. 
Protocardium hantoniense (Hd. 
MS.). 
Cardita deltoidea. 
xCytherea incrassata. 
suborbicularis (Hd. WS.). 
Solandri. 
Cyprina Nysti. 


* 


[x]Crassatella Sowerbyi, var. hanto- 


niensis (Hd. MS.). 
xCorbula ficus. 
x cuspidata. 
[x]Psammobia compressa, var. arcu- 
ata (Hd. MS.). 
Panopeea sulculosa (Hd. MS.). 
Madrepora anglica. 
Dendrophyllha. 
Lobopsammia cariosa. 
Balanophyllia granulata. 
Solenastrza cellulosa. 


Of this shorter list a proportion of about 50 per cent. pass up 
from Barton or Bracklesham beds; so that, taking the whole fauna 
or the more characteristic members of it only, in either case nearly 
one half pass up from Barton beds—a very different thing from one 


12 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


fifth, which was the proportion given in the paper referred to (op. 
cit. p. 150). If we compare with this the ratio of Barton forms in 
the list of commoner Headon-Hill given above (supra, p. 105), we find 
that 30 per cent. pass up from Barton beds (while in the complete list 
of the Headon-Hill fauna the porportion is also about 29 per cent.). 
Moreover, if we first eliminate the forms that occur also at Brocken- 
hurst, so as to obtain what we may consider as specially Headon 
species, the proportion becomes even lower. On all hands the 
paleeontological evidence seems perfectly in accord with the strati- 
eraphical. 

Relation of Colwell Marine to Brockenhurst Fauna.—In order to 
see whether the Colwell bed is more nearly related to the Brocken- 
hurst than is the Headon-Hill bed, we will first take the proportion 
of Barton forms in it for comparison with similar treatment of the 
Headon-Hill catalogue. Examination of the list gives us a pro- 
portion of 29 per cent. of Barton forms in the Colwell-Bay bed; we 
saw above that, in the Brockenhurst bed, the ratio was about 50 per 
cent. and in the Headon marine bed 29 percent. An inspection of 
the list of more characteristic Colwell and Headon marine fossils 
(supra, p. 105) showed that these faunas are practically identical 
Now we see that their proportion of Barton forms is nearly equal, 
and far lower than in the Brockenhurst bed. 

To complete the proof from fossils, if any such is needed, we may 
inquire whether there are more Brockenhurst forms peculiar to 
Colwell Bay than to Headon Hill. Examination of the lists 
shows that only the following Brockenhurst species occur at 
Colwell Bay and not at Headon Hill, viz. Scalaria tessellata 
and Tellina affinis, the latter passing up from Barton beds; 
while those occurring at Headon Hill and not at Colwell Bay are 
Marginella estuarina and Cardita paucicostata—two only in each 
case, which amounts to perfect equality. If, on the other hand, we 
count those common to the Colwell and Headon marine beds, ail not 
occurring at Brockenhurst, we find twenty-six species in this 
category. We are at a loss to understand how any one could 
imagine that the Brockenhurst fauna is identical with that of the 
Colwell-Bay bed and newer than that of Headon Hill. 

That the Colwell-Bay bed is stratigraphically identical with the 
Headon Middle Marine we hope that we have sufficiently proved ; 
and the fact is confirmed by fossil evidence. The same twofold 
proof has been brought forward to demonstrate that the Brocken- 
hurst bed, where present, lies at the base of the marine Headon 
beds and immediately above the Lower Headon. This bed is 
absent at Colwell Bay and Headon Hill, but is seen at Whitecliff 
Bay, Brockenhurst, and Lyndhurst. 

Since the Middle Headon includes every thing between the fresh- 
water Upper and Lower Headon, it must be allowed to include fhe 
Brockenhurst beds, though that special fauna was not known when 
names were given to these groups of strata. It would cause the 
ereatest inconvenience to abandon the term Middle Headon, as it 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. Lis 


would entail the abandonment of the names Upper and Lower also. 
There is therefore no room for the term ‘“ Brockenhurst Series” in 
the sense proposed in the paper referred to (op. ct. p. 168)—a 
classification which would be in conflict with the best authorities, 
and founded, as we have endeavoured to show, on a defective appre- 
ciation of the beds. We may urge that no new facts have been 
discovered concerning the succession of the strata to make any 
correctionary classification or nomenclature in the main groups of 
the Upper Eocene at all necessary or desirable, and we should prefer 
to retain the local groupings which have been so long familiar to 
geologists. We may denote as the “ Brockenhurst Beds ” the lower 
part of the Middle Headon with the rich Brockenhurst fauna. It 
is not always developed; the absence of admixture of fresh water 
was evidently the necessary condition of its abundance of marine 
mollusca and of the existence of its corals. 

It may perhaps be subdivided into the “‘ Brockenhurst zone” and 
the ‘‘Roydon zone”*. The correlation of this fauna was justly made 


* We found the Roydon brick-yard pit in a good state for examination this 
summer, and obtained from it twenty-eight species. They all came from the 
sandy clays with bands of iron-ore septaria; the lowest beds were below the 
level of the standing water. ‘The section is as follows :-— 


Gravel, Post-Tertiary. 


2-3 feet. Bluish to yellow-grey clay. 
( Murex -sexdentatus, Cardita ob- 
| longa var. transversa, Pisania 
Gr hies { ‘“Shell-bed ;” clay very full of 4 labiata, Trigonocelia deltoidea, 
5 shells. |  Ostrea velata, Cytherea incras- 
| sata, Cyrena obovata var..sub- 
\ 


regularis (Ed. MS.). 


Voluta geminata, Voluta spinosa, 
Strepsidura armata, Pleuro- 


( Grey clay. 
| toma transversaria, Pleuro- 


| Two nodule-bands of iron-ore 
4 feet. |  septaria separated ° Sune: ae 
7 feet 4 1 pare by grey toma  hantoniensis, Natica 
} sandy clay. ; 1 , . 
eee 9 epiglottina, Bulla Lamarckit, 
| Stiff bluish clay for the lower 2 . Sete 
leet Protocardium hantoniense, 
: Cytherea suborbicularis, Psam- 
mobia estuarina,  Corbula 
pisum, &e. 


7 feet. Greenish-grey clayey sands. 


— 


Reposing on Lower Headon fresh- 
water clays. 


The shelly bed, we consider, represents part of the Venus-bed or Headon- 
Hill marine zone, since it contains the characteristic oyster and Murex sexden- 
tatus, &e. 

The clays and clayey sands below, of which we examined 74 feet, while, 
according to the statement of the men employed, the remaining sandy beds 
below are another 7 feet, we propose provisionally to term the “ Roydon zone.” 
Tt is characterized paleontologically by the abundance of Voluta geminata, 
differing from the “ Brockenhurst zone” by the absence or great rarity of Voluta 
suturalis, Pleurotoma cymea, and Cytherea Solandrz, for the latter shells are 


114 H. KEEPING AND E, B. TAWNEY ON THE BEDS AT 


by Von Konen in 1864, and by Dr. Duncan for the corals in 1866, 
whose work is not in any way affected by any thing in the present 
essay. We have merely striven to prevent the beds in the Isle of 
Wight and the New Forest being thrown again into confusion, and 
the accurate work of EK. Forbes and the Geological Survey being 
rejected on such insufficient grounds as have ‘been recently put 
forward. 


abundant in that rich zone, a few inches thick, in the Whitley-Ridge railway- 
cuttin 

At the time of obtaining the Roydon fossils in the Edwards collection, one 
of the authors sank a pit to the base of these beds, and they were found lying 
immediately on the freshwater Lower Headon clays, the Brockenhurst zone 
being absent; the latter has apparently thinned out here, as the Roydon zone 
itself thins out a little further west. 

Of the White-Cliff Bay beds we are disposed to place i in the Roydon zone all 
those between the ‘‘ Venus-bed ” clays of the Geological Survey Vertical Section 
and the lowest two feet of sandy clays lying immediately on the eroded surface 
of the freshwater Lower Headon, which said bed we have described above as the 
“ Brockenhurst zone.” The Roydon zone will thus include the 42 feet of yellow 
and green sands with ironstone-nodules, of which the chief fossil is Psammobia 
estuarina (=Sanguinolaria Hollowayst of the Geological Survey Section), also 
the remaining 12 feet of beds described in the legend as “brown clay,” but 
which in their unweathered condition are slate-colour to greenish grey. 

We had unfortunately not sufficient opportunity to work out the fauna com- 
pletely; but such fossils as we found induce us to parallel these beds with those 
of the Roydon brick-pit. The lithological character of these lower beds, as seen 
below low-water mark at equinoctial tide, is singularly like the clayey sands of 
the Roydon brick-yard, while their chief fossils are the Psammobia, lying in the 
natural position of life, and Cardita deltotdea. At some future time we hope 
to work out the fauna more completely. 

At Cutwalk hill, Lyndhurst, both the Brockenhurst and Roydon zones occur. 
In the pits which one of the authors sank in obtaining fossils for Mr. Edwards 
and others from this locality, the Brockenhurst zone was found lying, as usual, 
immediately on the freshwater clays of the Lower Headon. 

Hence we may divide the Middle Headon into three zones, distinguishable 
easily by fossils (though, of course, many species are common to all three), viz. 
the Brockenhurst, the Roydon, and the Venus-bed or Headon-Hill zone. The 
percentage of Barton forms diminishes as we ascend in the series. 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. aly 


Fossils of the Middle Headon series, including the Brockenhurst 


beds *. 
Mippie Heapon. 
UprEr 
Brockenhurst  |Baagsuor. 
beds. 
Be : 
. os} ~ 
See es Loe, 
SCRA rk se iene lees 
® qd]jc sl | SS 5 Si ly. 
= S) a 5 2ijf}] 4 lines 
mS | = | SO = S 3) ceo 
Ei slaielel/e12| #3 
ee] fe tS Ge) ee tase | ea | 9s 
xLamna contortidens (Ag.) ......... ee mae eg « |K.T. K.T. 
Marginella simplex (Hdw.)......... shew lien ec Mi ctestet torte 50 * 
estuarina (LdW:) 2 ssccss- dae | pe * * 
—— Vittata (Hdw.) .........sceeseees con ||: 
Voluta geminata (Sow.) ............ noo ees x |K.T.) x 
— var.tereticosta(Hd.MS.)| ... |... | ... | * | * 
% decors @Bey7,.)—maga (Hd.).| ... ||... |... |... (KT) x | x 
suturalis (Mysz) = contabu- 
[EUG (JEG) ih ORO ie Aa ee eee A iy Oem eae Glico at Gee Rn 2 lia 
* SIMON CHAI)! Vesetaesacko ces x |K.T.) « | x |K.T.) «x | * 
Mitra gracilenta (Hd. MS.)......... Sisteeall meted esate | ie 
apmcenmiatan (HO neVLS. yess ectesclcoae || veces |v eines. vooes (Woelyl) eae |) 96 
— polygyra (Hd. MS.) ......... Woe hrceiseahlpravo sis Wacetece litsione [itevictant| thee 
[¥ |Conorbis dormitor(So/.), var. semi- 
TUTOR CLUGAVE Vcasescce tase odasecsases ser ea semlinatisedl mend lieteate ta 9a! (UE 
[x] procerus (Beyr.)=alatus, var. 
Ienatllissey CHA ac. ce ces sede ecumese x |K.T.) x | x 
Pleurotoma transversaria (Lam.) x |K.T.| x | x 
Gyameear(HG.)) cli ecbecc noses ees Soe eo Ren pean eae ome ea De * 
— var. nana (Hd.)......... Gon ates aay re 
* YGC OA CHG.) ccs vepecsceset ec Shou epee toes es x 
[ayes Dio kes (27000 Meee ee eee Bel ee Ae vse ate % 
Pa NVOOUU CHA.) 20. ccececsesoseee Paes le oe 
headonensis (Hdw.)............ x | « | « |K.T. i le as 
* denticula; (Base.) 0.20.2. <s06: aes il ace Watereteel [iar 
x— var. odontella (Hd.) .... x | « | * |K.TK.T.) x 
—— leviuscula (EHd.) ...........-... ee alia w ales. Kane el) Serio peer [Ae 
subdenticulata (Goldf.) = 
hantoniensis (Hd.) ..........0.06: Bek Mal ncsatele| aceite | oes RGAE SSO Rae 
* Borsonia sulcata (Mov.).........08. Ea ee cae eee 2a peer el 8 
IOMMEP EER eitoncesen avd eriavioicieoa ve Cai leg, cabilitaeet tia ceht | ead Des 
Chenopus Margerini(deKon.), var. 
speciosa (Schlot.) ...........0e0.e0. Peeters i sacsc fh amie a | acer oe i ee 
¥Rimella rimosa (Sol.) ............... San iB 36. =| 96S Ile ses KG aicseda aire 
xHippocrenes (Rostellaria) ampla 
DIGGS.) ), SACS CBAC BREA ee aan een ee Sse eee eh | eter | retoet VGA tea (3) lise 
Carried forward ...... Bs Dey yy ep Ue’ IP Alte) 1 


t Those marked with an asterisk pass up from Barton or Bracklesham 
beds. The asterisk enclosed in brackets signifies that the type species exists in 
Barton beds, but not the special variety. The initials K.T. denote that the 

_ citation is the result of our own researches instead of being founded on the 
Edwards collection. 


116 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


Fossils of Middle Headon series, Jc. (continued). 


Mippie Heapon. 


Upper 
Brockenhurst |Bacsuor. 
beds. 
= A x 
P| Sis | 8 | 3 |e eee 
sore as ss | “eS 3S an 
Se ls | Sy a | Seems 
ee popes |e te | 
Brought forward je. sace re AND | SO oa aes 1 
Murex hantoniensis (Hd. J7S.) 2.2)... |... | ..2 |) ce. (IK) See ee 
sexdentatus (Sow.) ............ x | * | x 
—— , var. cinctus (Hd.)...... Baca sea a liane 
= MM (YSO/5) sestaccocosdtocads ck lela pili slau thy cme 9 Ges ee 
*x—— obtusus (Desh.) ...........005- PP reel elton | age.|| 3 
S)Db. MenoSdccbooqoocanonseos Sag Gelb 
xTyphis pungens (Sol.) ............- wae if pete Te Bec] SRN ROS ea ie 
Cantharus subcostatus (d.) ...... sae jegaaiel | Meee TN Meetsiona ected RRS 
Fasciolaria crebrilinea (Hd. MS.)| ... | * 
Pisania (Fusus) labiata (Sow.) ...| ... | * | * | * |K.T) « | x 
-——, var. concinna ...........- Se ese 
acuticosta (Hd@.) ............--- ae x 
Phos scalaroides (Lam.) .....-...0+ x 
*Clavella (Fusus) longeeva (Sol.)...) ... | --- | .. | * |K.T) x | x 
Chrysodomus (Fusus) Sandber- 
BENIN CL C0/774) gel eeacey eee a cre ets wae [cage ae AN Dec G [Pee eae eee 
Leiostoma ovatum (Beyr.) ........- wes vesicle «/fteine a)! ceanall eee een Mi 
xStrepsidura (Buccinum) armata 
(CSIZTOD TENA Rkecr ne Bee EP A Len Sen wee |) bey | KCTS ia ae 
semicostata (Hd. MS.) ...... * 
xCominella (Buccinum)  deserta 
(ISOLDE tee eeneru oan ee von af vse ||) ateov'l|lleciesel| ACTS Pe ioe 
flexuosa (Hd. MS.) .......4...: oie (KT |e 21) 20 a ie 
ventricosa (Hd. MS_)......... MOR ore a ee 
xAncillaria buccinoides (Lam.). ...) * |K.T.) *« | * |K.T) x | x 
xCassis ambigua (Sol.) ............4.. wots [iedeisie vl toi fh pe | ERE Sa 
*Natica hantoniensis (Sow.) ....... ase cose: | Senin. |. serie yt CGEIE ieee 
% Obowatan (Sows) eaceeeenee eee uate md ata 
conulus (Bo. MES iho ease. wee [icesecla'eo es [yee eee ieee Moe 
Tea eaten wee Nes 96410. AE wal gael Ea a NG * 
* yar. grossiuscula 
(Ed. MS.) wie) Sia pee O eae Renee soe |. 9 | RAUS 20077|o2 Sn ee x 
% epiglottina (Lam.) .........--- Dell Geers \useoy| Kaede 
labellata (Lam)... ese S| eK ees aie 
; var, dubia) Cid SES 9) seal) eee [leet me lceate | iratrell tater * 
Cancellaria muricata (8. Wood) 
Sloymexoes GIL WIS) Soosonncosee KOT.) | 30 KEM rn ae 
elongata (Nyst) .....<......00. x [e56. | .3e L | ceo] Ree 
* EMU (ISOUS)s<css ns seel eee BME Aare, /a|ttere [tee of eS 
roydonensis (Hd. MS.) ...... SAS) Il ocean One esc 
xPyramidella (Turbonilla) obscura 
(LITE ULSI ONS ese ea nrnpmris ann eal ieee tessa oe 
Turbonilla plicatella (Hd. ae seettllieee cine 1 toe 
semilzevis (Hd. MS.) ......... cca etal eee Slane 
plicatilis (Hd. MS.)............ noo || ooo || #8 
Carried forward ...... 11 | 20 | 22 | 81 | 31 | 25 | 39] 8 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. ley 


Fossils of Middle Headon series, dc. (continued). 


MippiE Heapon. 


aa ae |. UPPER 
| Brockenhurst |Bacsuor. 
beds. 
a : 
a |S fe le 
Ss a) 
See ale | alae & 
slzlfi/sl/2l\2/2| es 
Sao lov) Seley Ea ee ae 
SiS Ole |e lala] A. 
Brought forward ............ 20 22) Sloe 25: 839 3 
*xTurbonilla obliquecostata(Hd.MS.)) ... | x 
Guia (2d. MS.). 0.0 .sccscess S|. 296 | 
sorella (did. MS.) ............ See 
Odostomia loxodonta (Hd. MS.)...) ...) * | * 
subumbilicata (Hd. MS.) ...) ... | * | x 
geminata (Ed. MS.) ......... gee) | een |t 3 
—— multispirata (Hd. MS.) ...... Sage |) 36 
gracilis (Ed. MS.) ............ ser eed 
x*—— hordeola (Zam.) ...........+08- | 
—— , var. angusta (Hd. MS.)| ... | ... |... | * 
Hulima gracillima (Hd. MS.)...... eel Cec ee a 
xCerithium variabile (Desh.) ...... Jeol Wt hl (ae Sal dies Salhi x % 
[*] submarginatum (Hd. MS.), 
var. recentior (d’ O7b.)..........+. se erie (eae? “ie x 
pseudocinctum (d’ Ord.) ...... Bee |e he Seem * | * 
—-— duplex (Sow.) .........sesseeees ei okey 
parvulum (Ed. MS.) ......... ee! 3 
—— pliciferum (Ed. MS.)......... Seen sete: «oe 
ventricosum (Sow.)..........45 x | * | x 
subeonoideum (Hd. MS.) ...| ... x 
—— contiguum (Desh.)? ......... seul 3 
—— multispiratum (Desh.) ...... {Pesto [bre | 
—— gyrostoma (Hd. MS.) ......+. sacl 1% 
concavum (Sow.)  ....sseeeee {SSG al, 9625]. 5 ewer 
yertenisn (HG IMS.) ...030000.% [eee edh ae 9. 
Melania fasciata (Sow.) .........66 eam sein! 236 |; * 
MUA (SOW.)\ 0. se0cererees [read ce eel ee ae 
x-—— brevicula (Hd. MS.) ......... % 
*xHydrobia anceps (Wood) ......... x x | % 
— Dubuissoni (Bowillet), var. 
petit (Hd. MOS.) ccacscsscceeees * 
bulimoides (Hd. MS.) ...... he | Gel A | 
Melanopsis subfusiformis (Morr.)| * | * | * % x 
masitormis (SOW.)  ..<.2..--.9 x |K.T.| x ae * 
Scalaria levis (Morr.) ..........0.008 saekip ae Wh Se 
tessellata (Hd. MS.) ......08. seal pete “esters e 3 
Nematura parvula (Desh.) ......... x (KAD) | oe. LN Gate 
—— pygmea (Forbes) ............ Wee, oceed eles: | 
lubricella (Brawn) ...........- | * 
Cecum Morrisii (Hd. MS.) ...... | aaceail ae 
Trochus pictus (Hd. MS.) .........) * | | | | 
*Trochita (Infundibulum) trochi- | | | | 
formis (Lam.) =obliqua (Sow.), ... | ... jy seo Kea x | 
Phorus cretifer (Hd. MS.) ......... K.T.| * 
Teinostoma minutissimum (Ed. 
MMe rasa casas cvvcnceoseesseeses ces leeooe| | eas 
Carried forward....... 25 | 46 | 47 | 384 | 33 | 26 | 44 | petal 


Q.J.G.8. No. 146. K 


118 H, KEEPING AND EB, B. TAWNEY ON THE BEDS AT 


Fossils of Middle Headon series, &c. ee 


ee Heapon. 


a ae | 
+ | 5 
2) a 
BS 
ro | 
5 | 6 
| 
Brought forward ............ | 20 | 46 
Teinostoma micans (Hd. MS.)......! ... as 
iINerita aperta(Sows ners cee: x |K.T 
[estuarina (Hd. MS.)|=N. 
Zouellai US: OOd) iene eeesseee seen * |K.T. 
Neritina concava (Sow.) ......066...! * |K.T 
[x]Rissoa carinata (Hd. MS.), var. 
denticulata (Hd. MS.)............ Hear ae 
ditropis (Hd. MS.)............ | * 
Tornatella Minato mis (Sandb.).| x 
alteran(WDeS/9) aa asensnese she 
| dactylina (Desh.) ............ sas 
| % simulatum (Sow.) ............ a5 
| Adeorbis apertus (Hd. MS.) ...... * 
zstuarinus (Hd. MS.) ...... | % | 
| xOrthostoma crenatum (Sow.)...... haa 
| --——retiarium (Hd. MS.) ......... ieee 
xRingicula parva (Hd. MS.)......... a a 
Bulla estuarina (Hd. MS.)......... x | x 
lopvaney ye sain (SLAIA)) Snooeensosse 
| —— curta (Hd. MS.) .....0......0.- 
simillima (Hd. MS.) |e y 
PeSCUATIMA ||| Pesce uneaceR cents a 
% attenuata (Sow.) ......scc0cce00| e+ 
navella (Hd. MS.) ..........+. ire 
Sowerbyi (Wyst) .......0.02.06- hase 
; —— tenuicula (Hd. MS.) ......... | * 
| # elliptica, (Sow:)it accrneesseeecl | x 
Cylichna globulus (Hd. MS.)...... S00 
OyeH NS (Cte, AHISE) Gcadaccoobocass Laas 
Dentaliumyyspie suede cece coos 
xAnomia tenuistriata (Desk.) ...... Ls soe tee 
Ostrea velata (8. Wood) ............ x |K.T 
*——- flabellula (Lam.) = venti-| 
| labruna (Goldie) merece cee Urea es 
|  Pecten belli aeeee (S. Wood) . ae 
xAvicula media (Sow.)............-+-- Nes 
xMytilus affinis (Sow.) ............... \K.T.| 
Modiola Nysti (Kieh/, MS.) ...... a 
demota(HaiS)) nee f 
sod ea biangula (Lam,) .)0...2.0.2---- He | 
appendiculata (Sow.) ......... one! sede 
ata (OGtL0) cctenvacncan leeseuellin ase 
* duplicata (Sow.)=sulcicos- 
Lata GING/SE) ilar cies cia rate awiane CHOIRS, 
| ¥Trigonoccelia deltoidea (Lam.) .... #* | * 
Nucula headonensis (forbes) ...... [ae 
Carried forward...... | 41 | 56 


Colwell Bay. 


* ss 


a 


Upper 
Brockenhurst |BaAqsHort. 
beds. 
2 4 bre) 
| 2) oe 
a [3 =) r= bend 
ey oS Sl | ce! 
ra) SS ures) Oo =| 
ZS || a S iS & 
a | | oe eae 
34 | 38 | 26 | 44 La 
| + 
se 2 Gr 
Ke a ee 
% 
coe CS *% 
. |K.T. 
K.T. 
Pre goad be 
Kea % 
pan OM Weiner |. oe 
cinco (RG FIDA eel lero 
3 ECA areca ee 
1 Hes * 
6 % ea 
39 | 45 | 31 | 57 14 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 


119 


Fossils of Middle Headon series, jc. (continued). 


Mippirz Heapon. 


UPPER 
Brockenhurst (Bacsnor. 
beds. | 
ie B 3 
set | Ee Pee lc aa eae Ve ng 
eee eee ae 
> a | to =| z ee 
elelelel2|sla| we 
Sele ele ie |e] ee 
pal eel tS) | ee |e es) | el ns 
Brought forward ............ AL SGP Gl toon aoninclenieon 14 
Nucula nudata ( Wood)...........066. weer [Fe Se le 
* BUMS (SOW)! 6.2 jcoeacte nsec tordle cea lavaren ol aoe EET 
* Lng! (17/12) aan eec ae Ws [baceat | notes Goods 
Leda propinqua (Wood) ............| «+ KET ioe Ke Ke: 
¥—— minima (Sow.)...ccecceccceceene att 5 OSE Be * 
Cardita simplex (Hd. MS.)......... Bad * 
paucicostata (Sandb.)=nodi- 
Celeb HAWS) ses cy vas ocoed sede ee ens Xe tele oi |i 9e 
deltoidea (Sow.) .........c0008. ae oy ESSAI HERG ASSN) Pee alee 
orbicularis (Goldf.) ......... Lil Selteta lias «Re 
[+] oblonga (Sow.), var. trans- 
SENAY MIS.) atcseoes ee stoacsc: Se GOIN Wicca [has alee oe % 


——— 


[*|Crassatella Sowerbyi, var. hanto- 
MINEMSIS A CHA s)) osc secvdeerccesss ens 
xLucina obesa (Hd. MS.) 
CONCAVACDELT.) sss <ceeees cine 
—— pulvinata (Hd. MS.) ......... 
x—— bartonensis (Hd. MS.) 
maitiatan(Hid. MS.) ..- csc eceae 
Strigilla colvellensis (Hd. M/S.) ... 
pulchella (4g.) 
Diplodonta suborbicularis (Ed. 
M, 


@eceeeossons 


@eocoee 


% 


@oereceshoscces 


Pooeereesrersose ee SeSoorees 2 2 oe 


e@ocvece 


AIS: Sep ccaGoa aga GUpee eR EE Eee Eee 
xCypricardia pectinifera (Sow.) ... 
Isocardia transversa (Nyst) 
xScintilla angusta (S. Wood) ...... 
Lepton nitidulum (S. Wood) ...... 
ae fomidum (Hd, MS.) ......-.. 
Cyprina Nysti (He6.) 
«Cytherea incrassata (Sow.)......... 


== — ftumida (Hd. MS.) ..........00 
— suborbicularis (Hd. MS.) ... 
subelliptica (Hd. MS.) 


* 


eoceece 


, var. serratina (Hd. MS.) MG 


eee 


M 
ww 


eee 


* 


turgescens (Hd. MS.)......... i 


—— hantoniensis (Hd. MS.) ...... ore 


* Solandri (Sow.), var. attenu- 

ERED AVES.) \\..0.000 cscs sonwesses 

[*] elegans (Lam.), var. 0 
¥—— Suessonensis (Desh.) 


ee CS | 


Kea 
KT: % 
ae ‘ 
a ea 
* 
~~ |K|laee 
x 
Pe KE Ke % 
Heese IKE Sle % 
oe Ad B eau Nh aes 
* x IK.T x * 
Kea. * 
San rd NB 
71 | 54} 58 | 34 | 79 17 


120 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


Fossils of Middle Headon series, Jc. (continued). 


Mippir Heapon. 


| UPPER 
| Brockenhurst | BaasHot. 
| beds. 
Ss) 8 | as 
= as tl © 
Elo |ts | 6 | eo mene r 
‘Shs 16) eS Saar eo S 
° OMaS Ss) = Ss £ a 
sl men pedapics er i) 
Brought forward ............ 51 | 68 | 71 | 54 | 58 | 34 | 79 il 
[x]Psammobia compressa (Sovw.), | 
var. estuarina (Hd, MS.), and HO] 36 | eat = * 
var. arcuata (Hd. MS.) ...... | | | 
rudis (Lam.)=solida (Sow.).| -..| * | * | 2 || * 
xTellina affinis (Ed. MS.) ......... Looe ldo) cae eg Kee Teme 
headonensis (Hd. MS.) ...... [tedees) ara seaaal | | | 
sphenoides (Hd. MS.) ...... buresiealey ae gl nse | 
Syndosmya colvellensis (Hd. MS.)| ... |... | * 
Mactra fastigiata (Hd. MS.) ...... x |K.T| x * 
SUSE (73K LUISE) cagaccmaaecoe meena | | * 
xMya angustata (Sow.)=M. pro- | | | | 
Guctai(HdaViS)\ine eee ence * | x | * |K.TIK.T, * 
xCorbula pisum (Sow.).............5- cre fle oe] 05.6 eM eR ; 
x Cuspidata (SO) eee sseeeseases % | 3% | a [KOT KO | se 
DRAM USER Ve Goeehcos secebeoa <8 acs | cos |e 
*Panopza subeffusa (Hd. MS.)...... secre & ae au ae | 
sulculosa (Hd. MS.) .........| »» Sees ofomewe |) «88a le eeanl een 
*Solen gracilis (S0.) vo .sccwececcwcels m7) |) =} tectell| ucceete Meret teen 
Corbicula obovata (Sow.) ......... | gee |. 36. | 3600) 2 een oa cane ae 
Cyrena cycladiformis (Desh.) ..7...|| “KOKO es ere ee x 
pisuna (Ves) cseewcec eee mca | | | 
«Clavagella coronatia (Desi.)rsc.ccecc| seo |) 0 || eo) || enn een Mneeean mE 
Gol ditussii(@P i217) ee eee wes. [ewe [oleate alte ter | rea teen Im 
Fistularia Heyseana (Phill) ...... coy ld ccewelmene Bee bcc ||) “5? 
SED SCENE S\Dporasqoooodobdeoagondpoacnos wae |. sentel Wont | seth een eee gan 
Pola Spi msec seca eee rere aera: voee | gee o| eet aeee | Aeeean eeeaa Mie 
Peredo,; Sp.. :.s.c0ieadecesubecsensessscn|aese: ic ve,1| soe Meee al iNeeens ene 
Serpula; (Spungen ee eearee [te23)|| 220 || see | 208 K.T.| 
Pollicipes reflexus (Sow.) ......... lees jee | 
*Balanus unguiformis (Sow.) ...... coe fSGMts) Sone CGI, | x 
Callianassa Baylii (Woodw.) ...... pape UG bohe ee IU C oT | | 
Solenastrzea cellulosa (Dune.). ...| ... |... | «-» |)... (KOE) 2.5 
IEG soon (YG) sococoneoner cae ode? W abe al coe shel Re ae Ieneo | 
Reussi CD 21c)) Were eee | see | ewe of dere leess Rect leet oe 
gemmans (Dune) ae.2..-- + Jo sicee Hew |/celees | (teri Gel eee 
—— Beyrichii (Dune.) ............ ccs Peace |leosc || obo | cco | cos | 
granulata (Dunce.) ..........+: | see i) sSe-oi) swine |e eee ee 


Balanophyllia granulata (Dune.)..|.... |... |... |-s-- | cee. 
Mendropliviliale see serra eee | 
Eobopsammia granulata (Dune:).-| ...%| 22.0) ee. secon eee nee 

Titharzea brockenhurstii (Dunc.)..| ... | ... |... |... [KAT] ... | # | 


Asxopora) Michelint (Dwne.)-.2cc0| ss-+ |. se | =e ecen eee eee 
Madrepora Solandri (Defr.) ...... wes | ‘aveaili Mee) eee cal aleeyeen eee Ie 
Roemer (Ya7es)) wos seen Kern Recs eoreilscs | cos | 8 
amelical(27e>)\iacsecceasascess wos: |: wales | Me se 0 eet ENGIN s IRterereial 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 121 


Notes relating to Divergences from Prof. Judd’s lists of Fossils, with 
Observations on the Edwards Collection of Middle-Headon Fossils. 


Marginella simplex is not in the Edwards collection as from 
Barton ; this and the following ten species, viz. Mytilus strigillatus, 
Borsonia sulcata, Nematura parvula, Trigonocelia deltoidea, Lucina 
obesa, L. concava, Cardita oblonga var., Cardium obliquum, Scintilla 
angusta, and Panopea subeffusa, are given in one list as occurring in 
Barton beds; and in the other list this range is denied to them. 

The range into Barton or Bracklesham beds (as shown by the 
Edwards collection) has been overlooked in the case of many species 
(viz. twenty-two) in the lists of Colwell-Bay and Brockenhurst 
fossils (op. cit. pp. 153-156). 

Voluta depauperata (Sow.) has been cited in error by Forbes from 
Colwell Bay ; it occurs only in Barton or Bracklesham beds, and is 
therefore omitted from our list. V. spinosa (Lam.), type, occurs in 
Barton and Bracklesham beds; the form from Brockenhurst and 
Middle-Headon localities might be recognized as a distinct variety. 

V. tereticosta (Kd. MS.) is plainly only a variety of V. geminata 
(Sow.) in which the coste are a little less spiny; all intermediate 
degrees occur. 

Clavella longeva, var. egrequt (Beyr.). Von Konen mentions 
this from Brockenhurst; but we find the absence of ridges as rare 
as in Barton examples, and therefore omit the varietal name. 

Hippocrenes ampla (Sow.) is not in Edw. coll. from Headon Hill 
or Hordwell ; and we consider it does not occur there. 

Murex seadentatus, var. cinctus (Kd. MS8.), is labelled in Edw. coll. 
as from Barton; but we suspect this to be inerror; it appears to be 
from Colwell Bay, as in Prof. Judd’s lists. 

Natica obovata (Sow.) occurs at Bracklesham, as shown by the 
Edw. coll. WV. grossiuscula (Kd. MS.) is probably only a variety of 
NV. Studer, as transitions exist between them. NV. dubia (Kd. M8.) 
we consider only a large variety of NV. lamellata (Lam.). NV. epi- 
glottina (Lam.) is in Edw. coll. labelled as from Hordwell; we 
found it at Roydon. 

Cancellaria elongata (Nyst) is in Edw. coll. from Headon Hill 
and Hordwell, and C. muricata from Hordwell. C. roydonensis 
(Kd. MS.) seems a doubtful species. : 

Cerithiwm pyrgotum (Ed. MS.) we consider a Lower-Headon form, 
and omit it therefore. C. varians (Kd. MS.) isin Edw. coll. from 
Headon Hill. C. cavatum (Kd. MS.) may be only a variety of C. con- 
cavum ; both it and C. speculatum (Kd. MS.) occur only at Long 
Mead End, and should be omitted from the list. C. ventricosum 
(Sow.) is in Edw. coll. from Colwell Bay; C. subventricosum (Kd. 
MS.) and C. deperditum? (Lam.) in Edw. coll. seem to be worn 
specimens of C.. ventricoswm; we agree with Prof. Judd in omitting 
them. C. marginatum (Kd. MS.), var. recentius (d’Orb.), occurs as 
in our list; its title to a distinct specific appellation seems doubtful. 
O. (Vicarya) concavum (Sow.) is in Edw. coll, labelled as from 


122 H. KEEPING AND E, B. TAWNEY ON THE BEDS AT 


Colwell Bay. C. trizonatum (Morr.) is in Edw. coll. from Hordwell 
&c.; it would therefore seem to be Lower Headon as well as 
Upper. C. subconordeum (Kd. MS.) seems doubtfully a distinct form. 
C. estuarinum (Kd. MS.) as from Hordwell, in Edw. coll., is founded 
on a minute fragment, and we do not insert it. C. headonense 
(Hd, MS.) is in Edw. coll. from Headon Hill; but we do not insert 
it, as it may be Lower Headon only. C. pulchrum (Kd. MS.) is in 
Edw. coll. as from marine Headon beds of. Hordwell. C. pseudo- 
cinctum (d’Orb.) is in Edw. coll. as from Barton. 

Lurbonilla plicatella (Kd. MS.) is in Edw.:coll: from Roydon only ; 
T. plicatilis (Kid. MS.) is there as from Colwell Bay.and Barton. 

Melania brevicula (Kd. MS.) is in Edw. coll. as from Hordwell 
and Barton; M. Woodi (Kd. MS.) from Hordwell seems a Lower- 
Headon shell, and is not inserted; the names M. conica, M. polygyra, 
M. minima (Sow. ),we could not ‘find represented in the Edw. coll., 
and omit them. 

fiissoa carinata (Kd. M8.) occurs at Barton, but. not the var. cden- 
ticulata, according to the labels in Edw. coll. 

Conorbis dormitor (Sol.) and C. procerus (Beyr.) occur at Barton, 
but not the special varieties to which MS. names are anes in the 
Kidw. coll. 

Hydrobia polita (Kdw.) from Headon Hill is Upper Hesnal only, 
and therefore omitted; H. anceps (Wood) is in Edw. coll. as from 
Hordwell, Colwell Bay, Long Mead End, and Barton; H. Dubuis- 
soni, var. rimata (Ed. MS.) is labelled as from Hordwell marine bed. 

Trochus pictus (Ed. MS.) is in Edw. coll. from Hordwell ; it is in 
the Woodwardian museum from near Setley Common, Lymington. 

Melanopsis ancillaroides (Desh.) was in Edw. coll. subsequently 
labelled MZ. subfusiformis (Morr.) ; the former name may be omitted. 
M, subulata (Sow.) occurs twice in Prof. Judd’s list, and with diffe- 
rent ranges. MZ. swbcarinata (Morr.) is in Edw. coll. as from Bem- 
bridge and Hordwell marine bed; at the latter locality it probably 
came from Lower Headon (freshwater) to judge from the aspect of 
the shell. 

The species of Adeorbis in Edw. coll. are founded on single indi- 
viduals ; A. estuarina (Kd. MS.) we could not find. 

Orthostoma crenatum (Sow.) is in Edw. coll. from Brockenhurst 
and Barton. 

Tornatella altera (Desh.) is determined from a single individual 
in bad preservation, and seems to us very doubtful. 7’. hinnefor- 
mis (Sandb.) [ste] of the Brockenhurst list 1s the same shell as 
Acicon limneiformis (Sandb.) of the Headon-Hill list; it is in 
Edw. coll. as from those two localities. A. stimulatus is inserted on 
the faith of our own researches. A. dactylinus (Desh.) is in Edw. 
coll. from Colwell Bay ; we have found it there also. 

Rtingicula ringens (Lam.) occurs only in Barton or lower beds. 

Trochita obliqua (Sow.) was described originally as “a small but 
perfect specimen from. Brakenhurst [sc], in Sussex ; the species is 
found much larger in the: cliff at Barton ;” the type was only 1 ineh 
long, and was recognized by Sowerby as occurring at Barton, “while 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. I 5S 


at Brockenhurst it would have been found in Bracklesham beds 
probably ; we consider it a young example of 7. infundibuliformis 
(Lam.). We do not know why Edwards should have referred 
Brockenhurst forms to a different species from the common Barton 
species: we have compared numerous examples both from Brocken- 
hurst and Whitecliff Bay, and consider them identical with Lamarck’s 
species ; we therefore rank 7’. obliqua as a synonym. 

Nerita estuarina (Ed. MS.) seems identical with 1. zonula (8. 
Woo 

Be pha neritopsidea (Kd. MS.) is from the Upper oe we 
therefore omit it. 

Ostrea flabellula (Lam.) is in Edw. coll. labelled as from “ Col- 
well Bay or Headon Hill” [sic], but in error; the specimen has 
evidently come from the Barton Clay. O. ventilabrum is not in 
Edw. coll. from Colwell Bay, and it does not occur there; we are 
not sure that the distinctive differences given by Mr. Searles Wood 
between this species and O. flabellula are constant ; we have com- 
pared examples of this oyster, so common in the Brockenhurst bed 
at Whitecliff Bay, with other examples from Barton beds; and 
some we consider perfectly identical with Barton and Bracklesham 
forms. 

Avicula media (Sow.) is not in Edw. coll. as from Hordwell or 
Headon Hill, but from Long Mead End, 7. e. probably from the 
Beacon Bunny (Barton) beds. 

Dreissena Brardii (Fauj.) is in Edw. coll. from Hamstead and 
Hordwell (Long Mead End), ranging thus through all the fresh- 
water series. It occurs in social groups. We found a single derived 
specimen just above the Lower-Headon boundary at Whitecliff Bay. 

Anomia tenuistriata (Desh.) is in Edw. coll. labelled as from 
Barton, Brockenhurst, and Hordwell; if the Brockenhurst example 
is to be identified with A. Alcestiana (Nyst), probably the Barton 
ones are so also. We follow Edwards in considering them all one 
species. 

Mytilus strigillatus (Wood) in Edw. coll. is only from Barton 
beds; we therefore omit it. M. affinis is abundant at Colwell Bay, 
and ranges up from Barton beds. 

Nucula semilis (Sow.) is in Edw. coll. only from Barton beds ; we 
found one imperfect specimen, however, at White Cliff Bay in the 
Brockenhurst bed. NV. lissa (S. Wood) is said by Mr. Wood to oecur 
at Brockenhurst; but it is not in Edw. coll. as from there; the 
Hordwell examples are possibly from the Upper Bagshot Sands. 

Arca appendiculata (Sow.) and A. levigata (Caill.) are in Edw. 
coll. as from Barton and Bracklesham beds. . 

Cardita deltoidea (Sow.) is not in Edw. coll. from Colwell Bay ; 
and we doubt the fact of its occurring there. 

Incina Menardi (Desh.), as so determined, is not in Edw. coll. 
L. gibbosula (Lam.) and LZ. pratensis (Kd. MS.), in Edw. coll. as 
from Long Mead End, are not from Headon beds: we therefore 
omit them. JZ. obesa (Kd. MS.) and Z. inflata (Kd. MS.) are in 
Edw. coll, as from Barton beds; the former is not distinguishable 


124 H, KEEPING AND E. B. TAWNEY ON THE BEDS AT 


from those labelled Z. concava; indeed Edwards admitted it as 
doubtfully distinct. 

Diplodonta obesa (Kd. MS.) exists in Edw. coll. from Barton and 
Bracklesham beds. D. dilatata (Sow.), determined as such, is not in 
Edw. coll. ; Dixon cites it from Bracklesham. 

Strigilla colvellensis (Kd. MS.) we found to be not unfrequent at 
Headon Hill. S. pulchella (Ag.) is in Kdw. coll. determined from 
a single imperfect valve: it is impossible to say whether it is a 
second species ; and we therefore omit it. 

Cardium Edwards (Desh.) is not in Edw. coll. from Brocken- 
hurst; it is'a Bracklesham shell. 

Cytherea suborbicularis (Kd. MS.) we found at Colwell Bay and 
also in the Brockenhurst zone at Whitecliff Bay. C. swessonensis 
(Desh.): under this name we find in Edw. coll. Barton and Lower- 
Kocene shells usually referred to C. tenwstriata (Sow.); we have it 
from the Brockenhurst zone in Whitecliff Bay. C. partemsulcata 
(Ed. MS.) is from Long Mead End only; we therefore omit it. 

Cyprina scutellaria (Desh.) is solely from Lower-Hocene locali- 
ties. OC. Nystt (Héb.) is the only species from Brockenhurst in 
Edw. coll.; we have it from Whitecliff Bay. 

Psammobia compressa (Sow.) is in Edw. coll. from Barton; the 
var. arcuata (MS.) is from Roydon and Brockenhurst; and var. 
estuarina (MS8.) is, according to the labels, from Hordwell, Colwell 
Bay, and Roydon. We have compared these examples, and cannot 
see any valid differences; we consider them all one species. 

Sanguinolaria Hollowaysuv (Sow.) is not in Kdw. coll. from Lynd- 
hurst, but is a Bracklesham shell. The Geological Survey cite it in 
error from Middle Headon beds of Whitecliff Bay. 

Tellina corbuloides (Kid. MS.) in Edw. coll., from Colwell Bay, is 
a crushed specimen, undeterminable, but probably not the Hamstead 
species; we omit it. 7’. ambigua (Sow.) is in Edw. coll. labelled as 
from Hordwell, but is probably not from Middle Headon beds, 
although Forbes cites it from Colwell Bay. 7. reflewa (Kdw.) is a 
Bracklesham shell, and does not occur above the Upper Bagshot of 
Long Mead End. 7. headonensis (Edw.)is in Edw. coll. from Hea- 
don Hill and Colwell Bay. TZ. sphenoides (Kdw.) is from Colwell 
Bay. 7’. affinis (Kd. MS.) is in Edw. coll. from Brockenhurst and 
Barton ; we found it also at Whitecliff Bay. 

Syndosmya colvellensis (Kd. MS.) seems founded on a single 
minute valve. 

Scintilla angusta (Kd. MS.) is from Colwell Bay, Hordwell, and 
Barton beds. 

Mactra fastigiata (Kd. MS.) which occurs abundantly at Headon 
Hill, is from Hordwell in Edw. coll. M. jilosa (Kd. MS.) from 
Colwell Bay and Long Mead End (Upper Bagshot) we are unable to 
consider a separate form, so rank it as a synonym. 

Solen gracilis (Sow.) is in Edw. coll. from Brockenhurst and Bar- 
ton beds. 

_ Mya angustata (Sow.) was originally found by Prof. Sedgwick at 
Colwell Bay; Edwards has an apparently identical shell which he 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT. 125 


at first identified with Sowerby’s species, but subseqnently altered 
to M. bartonensis (Ed. MS.) ; we cannot, however, see any distinc- 
tion between these and M. producta (Kd. MS.) from Colwell Bay ; 
we consider them all one species. Mr. Searles Wood doubts the 
Hamstead form M. minor, Forbes, being a distinct species either. 

Corbula pisum (Sow.) is not in Kdw. coll. from Headon Hill or 
Hordwell; and we doubt its occurrence there; it is, however, fairly 
abundant in the Brockenhurst zone of Whitecliff Bay. C. mtida 
(Sow.) is in Edw. coll. from Roydon and Long Mead End; it was 
originally described from Prof. Sedgwick’s specimens brought from 
Middle Headon beds. C. fortisulcata (Kd. MS.) we consider merely 
a variety of C. pisum; we omit it, since it comes from Long Mead 
End, probably from Barton beds. C. jicus was found at Brocken- 
hurst by one of the authors, and is now in the Woodwardian Mu- 
seum. 

Panopea corrugata (Sow.) is, according to the Edw. coll., only a 
Barton and Bracklesham shell ; but quite possibly P. subeffusa (Hd. 
MS.) is not really separable from this species; in either case one of 
the names must be omitted. 

Cyrena subreqularis (Kd. MS.) seems to pass into C. obovata (Sow.). 
C’, deperdita (Lam.) is in Edw. coll. from Headon Hill and Barton ; 
it is probably from the Lower Headon. C. arenaria (Forbes) in 
Edw. coll. is scarcely separable from the preceding; it is from 
Headon Hill and Hordwell, but apparently from Lower Headon. 
Mr. Searles Wood figures a different form as Forbes’s species. C. 
gibbosula (Morr.) is scarcely a distinct form. C. estuarina, C. alti- 
rupestris, C. obliquata, MS. names in Kdw. coll., are not inserted ; 
' they may probably be Lower-Headon shells. 

Balanus wnguiforms (Sow.) we found as frequent at Headon Hill 
as at Colwell Bay. 

Pollicipes reflecus (Sow.) is cited by Forbes from Colwell Bay. 

Callianassa Baylii (H. Woodw.) we have from all the zones of 
the Middle Headon. 


EXPLANATION OF PLATE V. 


The figure represents the coast-section from Headon Hill east of Heather- 
wood Point to Cliff End, Colwell Bay ; it passes inland at Warden Point 
to avoid the projecting promontory. An attempt is made to represent 
where beds may be seen iz situ, and where they are concealed by fallen 
material. Notwithstanding that the vertical scale is more than double 
the horizontal, it is impossible to show much detail; and for this refe- 
rence is to be made to the accompanying vertical sections (pp. 91, 98, and 
103). 

The bends of the coast-line are approximately indicated by the com- 
pass-bearings given, 


DIScuUssIon. 


The Prestpent remarked that the paper was one of great impor- 
tance. The question at issue was one sharply defined but difficult 
to come to a conclusion upon without visiting the sections. 


126 H. KEEPING AND E. B. TAWNEY ON THE BEDS AT 


Rey. O. Fisuur said he had visited the locality with Mr. Tawney’s 
paper and sections in manuscript, and that he agreed with the 
authors’ conclusions. He thought the error on the part of Prof. 
Judd might have proceeded from the fact that at the N.E. corner 
of Headon Hill the Middle-Headon fossils were found by the 
sea-shore; these, however, were not truly im situ, but had been 
brought down by aslip; this he thought possibly the key to the erro- 
neous interpretation. In confirmation of the view that the Colwell- 
Bay and Headon Venus-beds are one stratum, he had found it with 
its fossils in Totland brick-field, near the Hotel, exactly where it 
should occur to connect the disjoined portions. He had worked 
personally at the Brockenhurst locality at Whitley Ridge, and had 
identified the bed at the base of the Middle Headon in Whitecliff 
Bay. Consequently, if the Brockenhurst bed is to be called Oligo- 
cene, the Middle Headon can no longer be called Eocene. 

Prof. Jupp said that the paper rested largely on assumptions. His 
method of work in the field and in the museum had been made a 
matter of assumption. He had not hastily arrived at his conclusions, 
but for twenty years he had worked on these British beds, and for 
twelve years had studied their continental equivalents and collec- 
tions of fossils made from them. ‘The series of Kocene and Oligo- 
cene strata in Western Europe is perfectly clear; but when we 
come to Britain a difficulty has always existed. This confusion was 
removed by distinguishing the zone of C. concavum from the Broc- 
kenhurst series. ‘The authors’ sections were supposed to support 
those of the Survey; he thought on examination they would not do 
so. The mistake had really originated from using Cytherea encras- 
sata to fix the so-called “ Venus-bed” of fossil-collectors—the fact ~ 
being that that shell has a wide range, and there is more than one 
‘‘Venus-bed.” In asserting that the different ‘‘ Venus-beds” are 
upon the same horizon the authors begged the whole question. This 
autumn he again visited the island, and found that an excavation had 
been opened by the authors in a Venus-bed in Totland Bay, but 
in one quite different from that in Colwell Bay. Had the authors 
searched the Headon cliff they might have found other Venus-beds. 
The authors had confirmed his own conclusion that the Headon-Hill 
sands do not occur in Totland Bay; and this is fatal to their reading 
of the section. As regards the paleontological evidence, he thought 
that the method of comparison of most abundant fossils was often 
misleading, as might be shown in the case of the Cornbrash and 
Ragstone of the Lower Oolite. The authors say that the Brocken- 
hurst bed is not above but below the Venus-bed. Now the former is 
the equivalent of the Tongrian beds of Belgium; and foreign geolo- 
gists all regard the zone of C. concavwm as the top of the Bar- 
tonian—that is, of the Kocene. Hence the result of their interpre- 
tation of the section was to place beds with an Upper-Hocene fauna 
above those containing a Lower-Oligocene fauna. 

Mr. Starxrz Garpner said he had always thought that in the 
particular section under discussion there was only one Venus-bed : 
the section of Headon Hill till last year had been fairly clear; and 


HEADON HILL AND COLWELL BAY IN THE ISLE OF WIGHT.” 127 


he had never seen more himself. He thought ‘“‘ Upper Eocene” and 
“Oligocene” equivalent terms, and the question, which should remain 
in use? one of priority. 

Mr. Wuitaxsr said that cliff-sections in soft beds were apt to 
vary from time to time, so that observers who saw them under 
different conditions of exposure were likely to differ in inter- 
preting them. The examination of other parts of the island, 
and especially the mapping of limestones or other well-marked 
beds, might. partly help to settle the question in dispute. Perhaps 
the Geological Survey map had been constructed rather too much 
on theoretical grounds. 

The PresrpEnt said that on the one hand we had the minute 
measurements of Messrs. Keeping and Tawney, and, on the other, 
the wider views of Prof. Judd. At any rate these views were now 
on both sides well laid before the Society ; and the question, although 
a difficult one, as he had himself found in working over the ground 
25 years since with Dr. Wright, would be now carefully examined 
by many others. 

Mr. Tawney said that he thought Von Konen, in 1864, had rightly 
correlated the German and English beds. As for Mr. Whitaker’s 
remarks, he thought a person who was puzzled by a clitf-section 
would make but little of a drift-covered country where no sections 
were to be seen. He still denied what Prof. Judd had said about 
there being more than one marine series. The Certthiwm concavum 
zone of Prof. Hébert at Hordwell did not occupy the position attri- 
buted to this zone by Prof. Judd in Headon Hill. He maintained 
that there was but one Venus-bed. The 6-inch Ostrea vectensis bed 
in the Bembridge beds could not be confused with the Middle-Headon 
Venus-bed. 


(QS0a0" P. H, CARPENTER ON TWO NEW CRINOIDS FROM 


10. On two new Crrnorps from the Upper Cuarx of SovTHERN 
Swepen. By P. Herpert Carpenter, Esq., M.A., Assistant 
Master at Eton College. Communicated by Prof. P. Martin 
Duncan, M.B. Lond., F.R.S., F.G.S. (Read February 2, 1881.) 


[Puare VIL] 


Tux large work* of Prof. Geinitz on the fossils from the valley of 
the Elbe in Saxony contains a description of a small stalked Crinoid 
from. the “ Pliinerkalk’’ of Strehlen that has long been known to 
contain stem-joints of the type to which d’Orbigny gave the name 
Bourgueticrinust. Although no calyx was ever met with, the 
characteristic stem-joints were supposed to be sufficiently indi- 
cative of the presence of the common JB. ellipticus. Some ten years 
ago, however, a singularly perfect specimen was discovered, con- 
sisting of a complete stem with radicular cirrhi, and a calyx 
on the top of it (Pl. VI. fig. 1). But this calyx (fig. 2) proved 
to be totally different in its characters from those of the species 
of Bourgueticrinus described by @’Orbigny. In B. ellipticus the 
calyx is widest round the basal circlet, and tapers gradually 
downwards into the stem, while the outer surface of the radials 
has a considerable slant from above downwards and outwards. In 
@Orbigny’s other species, B. equalis, both radials and basals, 
especially the latter, are relatively narrower and higher, and the 
whole calyx, together with the top stem-joint, is almost uniformly 
cylindrical ; but in each case the top stem-joint is very large and 
relatively higher than that of any Apiocrinus, while in B. ellupticus 
it widens considerably from below upwards. Its height may be as 
much as or more than that of the basal and radial circlets together ; 
and the joints immediately below it gradually diminish in width 
until they resemble the ordinary stem-joints. 

In the Strehlen fossilt, however, the calyx is widest at its upper 
end, around the upper and outer edges of the radials, so that its 
diameter diminishes gradually from above downwards (Pl. VI. 
fig. 2a). The broad external faces of the radials slope downwards 
and inwards; and the basal circlet narrows still more, so that the 
diameter of its lower face is less than two thirds that of the upper 
surface of the radials (fig. 2b). But the top stem-joint, on which the 
calyx rests, only expands a little from its lower to its upper margin, 
and its increase in thickness over the one below it is far less marked 


* “Das Hlbthalgebirge in Sachsen,” Palzontographica, Band xx. Theil 2, 

Ds 1S Sh 
oS Histoire naturelle générale et particuliére des Crinoides vivans et fossiles 
(Paris, 1840), pp. 95, 96. 

¢ Thanks to the kindness of Prof. Geinitz, who has made a second exami- 
nation of this specimen, I am enabled to give a slightly more accurate figure 
of it than that published by him in the ‘ Elbthalgebirge’ (Taf. vi. fig. 9a). 
In the older figure only two joints are represented between the calyx and the 
enlargement on the stem. In that given here three joints are shown instead of 
two, this later interpretation of the markings on the upper part of the stem 
being considered by Prof. Geinitz to be the more accurate one. 


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THE UPPER CHALK OF SOUTHERN SWEDEN. 129 


than is the case in Bourgueticrinus. Both it and the joints imme- 
diately below it are decidedly smaller than those forming the lower 
part of the stem, which is just the reverse of what we find in Bour- 
gueticrinus. 

Prof. Geinitz was unable therefore to refer this specimen to 
Bourgueticrinus, despite the resemblance of its stem-joints to those 
of that type; but he supposed it to belong to the genus Antedon on 
account of the resemblance of its calyx to that of Ant. Sarsw, as 
represented by M. Sars in his well-known ‘ Mémoires pour servir a 
la connaissance des Crinoides vivants.’ In the specimens figured 
by Sars*, ‘‘the upper end of the stem is not thickened, and the 
calyx widens from its base to the upper end of the first radial,” just 
as in the fossil from Strehlen (Geinitz Tt). This resemblance is but 
an imperfect one, however; for Sars’s specimens were merely the 
stalked larvee of Ant. Sarsei, not more than an inch long. In the 
later stages of these larvet the uppermost stem-joint or future 
centro-dorsal piece of the mature and free Antedon not only becomes 
considerably enlarged, so as entirely to conceal the basals, but it 
also bears cirrhi. This, however, is not the case with the corre- 
sponding joint of the Strehlen fossil. 

Prof. Lundgren § has already pointed out that this specimen is 
too completely developed to be a larval Antedon like those figured 
by Sars; but neither he nor any other paleontologist has referred 
it to any definite position among the Crinoids. 

The ‘‘Mucronatenkreide” (= Upper Chalk) of Kopinge, near 
Ystad, in Southern Sweden, contains a quantity of stem-~joints 
(Pl. VI. figs. 3-6) which have been considered as belonging to 
Bourguetierinus ellipticus, though no calyx of this species has yet 
been found associated with them. Some years ago, however, a 
singular calyx was met with (Pl. VI. fig. 7), which was presented 
to the Geological Museum of the University of Lund by Herr Rector 
Bruzelius, of Ystad, into whose hands it had come. Its discovery 
was announced in the ‘Neues Jahrbuch ftir Mineralogie’ by Prof. 
Lundgren, who at once recognized its resemblance to the so-called 
Antedon Fischert of Geinitz (Pl. VI. fig. 2). It is, however, con- 
siderably larger and much less conical; but it has even less resem- 
blance than the Strehlen fossil has to the cylindrical Bowrgueticrinus 
equalis. During my recent visit to Lund for the purpose of 
examining the Comatulw of the Retzian collection, Prof. Lundgren 
showed me this fossil, and was kind enough to intrust it to me for 
description. For this and for many other acts of kindness I offer 
him my heartiest thanks. 

For these two fossils, so similar in their general characters though 
differing in points of detail, I believe that not only a new genus, 
but also a new family must be established. I propose to call the 
genus Mesocrinus; for while allied to Bourgueticrinus in the cha- 
racters of its stem-joints, it is quite as closely allied to the Penta- 
crinidg in the characters of its calyx. The only real resemblance 


* Op. cit. pls. v., Vi. T Op. cit. p. 18. t Op. cit. pl. vi. fig. 24. 
§ Neues Jahrbuch fiir Mineralogie, i876, pp. 180-182. 


130 P. H. CARPENTER ON TWO NEW CRINOIDS FROM 


to the Apiocrinide is in the nature of the faces of the stem-joints, 
which resemble those of Bourgueticrinus and Rhizocrinus. But 
similar joints occur in the larval Antedon and in Platycrinus * ; so 
that this resemblance does not go for much, especially when it is 
remembered that d’Orbigny speaks of the articular faces of the 
stem-joints in the Apiocrinide as being most frequently marked 
with radiating striee 7. 

On the other hand, Mesocrznus differs from the Apiocrinide as 
defined by d’Orbigny and by Zittelt, in the presence of verticils of 
cirrhi on the stem, and in the small size of its upper joints. 
Further, the central funnel of the calyx (Pl. VI. fig. 7a) is small, 
not wide and patent; while the articular faces of the radials are 
large, and not separated or barely in contact, but meet one another 
along the whole length of their sides, and bear distinct fossee for 
the attachment of muscles and ligaments. All these characters are 
more or less distinctive of the type of the Pentacrinide ; but Meso- 
crinus cannot be referred to that family, as it lacks the petaloid 
markings on the faces of the stem-joints. 


MzsocrInvs, 0. &. 


Calyx more or less conical, composed of five basals forming a 
complete ring, and five radials with high articular faces which bear 
distinct muscular and ligament-fosse, and are in contact for the 
whole length of their sides. Upper stem-joints the smallest. Lower 
ones with elliptical faces, the long axes of which are occupied by 
articular ridges. ‘The planes of these ridges on the two faces of 
each joint are more or less inclined to one another. The joints 
may bear single cirrhi, or two may combine to form a node for a 
verticil of two cirrhi. MRadicular cirrhi at the base of the stem. 


MusocRINvs suEDIcUS, n. sp. (Plate VI. fig. 7.) 


The lower part of the calyx is formed by five pentagonal basals, 
which are in complete contact laterally, so as entirely to cut off the 
radials from the top stem-joint. The lower surface of the basal 
pentagon is very concave, with a large central perforation ; but it 
is quite simple and devoid of any kind of ornamentation (Pl. VI. 
fig. 7b). The radials have very high outer surfaces (nearly twice as 
high as the basals), which slope very slightly upwards and outwards. 
Their articular faces are also rather high and trapezoidal, with large 
muscle-plates standing up around the centralfunnel. The thickened 
edges of the two muscle-plates on each radial are separated by a 
slight intermuscular notch. The transverse ridge expands into a 
large articular surface perforated by the opening of the central canal, 
to which it forms a thick rim on all sides. From this surface short 
ridges proceed upwards and outwards, separating the large muscle- 
fosse from the small ligament-fosse. The dorsal fossa for the 
elastic ligament is relatively small, being represented by little more 

* A Natural History of the Crinoidea (Bristol, 1821), pp. 34, 75. 
t Op. cit. p. 1. 
{ Handbuch der Palaontologie, Band i. p. 388. 


THE UPPER CHALK OF SOUTHERN SWEDEN. USE 


than the pit beneath the transverse articular ridge, around which 
is a simple smooth surface. 

Size. Height 5 millim., greatest diameter 5 mm., least diameter 
4 mm., greatest height of basals 1:5 mm., least height 1 mm., width 
3 mm. 

As already pointed out by Prot. Lundgren, the stem-joints which 
occur associated with this calyx are referable to three principal 
types. It is, of course, possible that they belong to anothe™ species 
altogether ; but, like Prof. Lundgren, I am inclined to refer both 
calyx and stem-joints to one and the same species. ‘The question 
cannot, of course, be decided until the Swedish collectors are fortunate 
enough to meet with a perfect specimen. 

Type 1. Thin circular disks, about 3 millim. in diameter, with 
faces perforated in the centre, but without markings of any kind. 
They probably belong to the upper part of the stem immediately 
beneath the calyx. Both in some forms of Bourgueticrinus* and in 
Rhizocrinus the upper stem-joints have simple faces without any 
of the characteristic sculpture which occurs lower down the stem. 
The thin penultimate joint of Mesocrinus Fischert (Pl. VI. fig. 1) 
would seem to have been of this nature; and one can readily 
imagine that in the larger WM. suedicus the number of such simple 
stem-joints was larger, as is actually the case in Bourgueticrinus. 

Type 2. The joints of this, the commonest type (PI. VI. fig. 3.), are 
higher, with oval articular faces, the long axes of which are occupied 
by transverse ridges. ‘The planes of the ridges at the two ends of 
each joint are inclined to one another at angles of from 60° to nearly 
90°; and the centre of each ridge expands considerably around the 
opening of the central canal into a well-marked articular surface. 
A median groove extends along each half of the ridge from the 

central opening towards the margin of the joint-face; and short 
shallow branches proceed from it on each side so as to cut out the 
upper portion of the ridge into a double row of small teeth. 

According to Prof. Lundgren these joints vary in size from 3 
to 8 millim. in diameter, usually 5 or 6 millim.; and their height 
is about equal to their diameter. 

Type 3 (Pl. VI. figs. 4-6). Wider but lower joints, the oval faces 
of which are much more pointed than in those oftype 2. They differ 
very much in the proportion of height to diameter. In the thicker 
ones (fig. 4) the expansion of the transverse ridge around the 
opening of the central canal is very distinct, and there is a crescentic 
pit on either side of it. But these features are much less marked 
in the thinner joints, the faces of which are flatter (fig. 5), while the 
transverse ridges scarcely expand at all around the central canal. 
Some of these joints bear portions of the cirrhus-sockets, as already 
pointed out by Prof. Lundgren. As in some species of Pentacrinus, 


* This is certainly the case in several specimens that I have examined, some 
of which, contained in the University collection at Berlin, were kindly shown 
to me by Prof. Beyrich. On the other hand, Quenstedt figures a top stem- 
joint of B. ellipticus with a distinct transverse ridge and articular facet on its 
under surface (Petrefactenkunde Deutschlands, Band iv. tab. 104, fig. 76). 


132 P, H. CARPENTER ON TWO NEW CRINOIDS FROM 


two joints contribute to the formation of a socket, and the sockets 
are at the ends of the long axes of the two apposed faces, so that 
there were two cirrhi at each node, and at least three nodes in im- 
mediate succession (fig. 6). The two grooves in the respective 
transverse ridges form by their apposition a canal which lodged the 
vessels proceeding to the two cirrhi from the central vascular axis of 
the stem, and opened at the bottom of each cirrhus-socket*. 

In the absence of complete specimens of Mesocrinus suedicus it is, 
of course, impossible to determine where these nodes or, rather, 
groups of nodes occured in the stem, or even whether a stem of this 
character was associated with the calyx under description. I suspect 
that the verticils of cirrhi were limited to the wide lower part of the 
stem ; but it is not possible to form any opinion as to whether they 
were simple cirrhi like those of Pentacrinus, or irregularly branched 
radicular structures like those on the lower stem-joints of Rhizocrinus 
lofotensis. 

Failing direct evidence to the contrary, it certainly seems to me 
most probable that the calyx and stem-joints from Kopinge all belong 
to one species, which would then differ from dMesocrinus Mischert in 
other characters than those of the calyx; for the lower joints of 
M. Fischert (P1\. VI. fig. 1) are all much longer than wide, as in 
Rhazocrinus, Bathycrinus, and the larval Antedon, and they rarely 
bore cirrhi(Pl. VI.fig. 1). In this latter respect Z. swedecus must have 
been related to MW. Fischert much in the same way as Lhizocrinus 
lofotensis with abundant radicular cirrhi on the lower part of its stem 
is related to &. Rawsoni f which had “ very few radicular cirrhi.” 

The wide lower stem-joints of MW. swedicus (Pl. VI. figs. 4-6) 
have much less resemblance to the corresponding joints of 1. 
Fischert than to those of Bourgueticrinus ellipticus, as represented 
by @Orbigny ; and they further resemble these last in having the 
transverse ridge continuous across the articular face. In B. con- 
strictus, however, the ridge is interrupted in the centre, and the 
more or less marked excavations in the lateral portions of the joint- 
face are connected with one another round the central canal, very 
much in the same manner as the two cornua of the grey matter 
unite around the central canal of the spinal cord +. ‘This feature is 
also distinctive of Rhizocrinus and of the Antedon-larva$; and in 
both of these types the transverse ridge is cut up into a double row 
of minute teeth. So far as I know, this character has never been 
described in any species of Bowrgueticrinus; and Geinitz neither de- 
scribes nor figures it in Mesocrinus Fischert, though (as mentioned 
above) it occurs in J. swedicus. 


* Quenstedt (op. ciz. p. 368) has described cirrhus-sockets on stem-joints from 
the White Chalk of Rugen, and has remarked that the grooves in the trans- 
verse ridges of these nodal joints are often very distinct; but there seem to 
have been more than two cirrhi at each node (tab. 104, fig. 63). 

t ‘Zoological results of the Hassler Expedition. I. Hchini, Crinoids, and 
Corals,’ Ilustr. Catalog. Mus. Comp. Zool. at Harvard College, no. viii. p. 27. 

{ This seems to be also the case in the so-called B. ellipticus from the Hocene 
of Traunstein (Quenstedt, op. cit. iv. tab. 104. fig. 82). 

§ Sars, ‘ Crinoides vivants,’ pp. 5, 6, pl. ii. fig. 27, and pl. vi. fig. 17. 


THE UPPER CHALK OF SOUTHERN SWEDEN. 133 


One of the stem-joints of WM. Fischeri which is figured by Geinitz 
is very singular. There isa thick articular rim around the opening 
of the central canal of the oval-oblong face, as in the lower stem- 
joints of MW. swedicus; but the ridge proceeding from this to either 
end of the oval is Y-shaped and not simple. Some of the stem-joints 
of B. ellipticus and also of Platycrinus levis which were figured by 
Miiler * present very similar characters; but I have not found 
them in any of the joints of J. suedicus. 

The genus Bourgueticrinus is rather in confusion just at present, 
all those stem-joints occurring in the Mesozoic rocks being referred 
to it that have elliptical articular faces with transverse ridges upon 
them, which are in different planes at the two ends of each joint; 
and we are unable to classify these joints properly in the absence of 
sufficiently perfect specimens of whole individuals. 

In any case, however, the so-called Antedon Fischeri and its 
Swedish ally have stem-joints closely approximating to the Bour- 
gueticrinus type (Pl. VI. figs. 3-6). Nevertheless the calyx is 
quite different (Pl. VI. figs. 2 & 7). Seen from the side it has a 
certain resemblance to some forms of Millericrinus Miinsterianus, 
d’Orb., and of WM. Nodotianus, dOrb. But this resemblance dis- 
appears altogether when the upper surface of the calyx is examined ; 
for the central funnel of Millericrinus is very large, and the articular 
faces of the radials which surround it are wide and low, barely 
meeting laterally; while Mesocrinus has quite a narrow central 
funnel (fig. 7, a) and relatively high articular faces, which diminish 
considerably in width towards their upper ends, but are in contact 
for the whole length of their sides. | 

On the whole the calyx most resembles that of a Pentacrinus, or 
rather of that section of the genus with a closed basal circlet, which 
is referred by de Loriol ¢ to Cawnocrinus. Had I either calyx alone 
before me, I should certainly refer it to Pentacrinus, among the recent 
species of which there is a considerable variation in the proportions 
of the different parts of the calyx t. In Carnocrinus, Millericrinus, 
Bourgueticrinus §, and Rhizocrinus, the composition of the calyx is 
the same. In all these genera there is a closed circlet of five basals, 
on which the five radials rest, just as they do in Mesocrinus. But 
there are considerable differences among these five types in the nature 
of the articular faces of the radials. In Afllericrinus these faces 
are very wide and low, and are nearly or quite separated laterally, 
while the fossee for the muscles and interarticular ligaments are 
usually but shghtly developed ||. In Bourgueticrinus the articular 

* Op. cat. pp. 34, 75. 

t ‘Monographie des Crinoides fossiles de Ja Suisse,’ Geneva, 1877-1879, 
p- 111. See also ‘On the Genus Solanocrinus, Goldfuss, and its Relation to 
Recent Comatulz,” Journ. Linn. Soc., Zool. vol. xv. 

{ Compare figs. 21 and 23 on pl. xi. of Journ. Linn. Soc. Zool. vol. xv., 
with fig. 3 on pl. vi. of the Popular Science Review, new ser. vol. iv. 

§ In some specimens of “ 6. ellipticus” in the national collection at South 
Kensington the basals appear externally as small triangular pieces which are 


not in contact laterally. 
|| According to de Loriol, Millericrinus differs from Apiocrinus in the presence 
of an articular facet on the first radials. In the latter genus “leur surface 


134 P. H. CARPENTER ON TWO NEW CRINOIDS FROM 


faces are much reduced and quite insignificant, while in Rhizocrinus 
they are wide, low, and separate, and more like those of Milleri- 
crinus, enclosing a wide central funnel. 

In both species of Mesocrinus, however, and especially in 7. suedicus, 
the articular faces are much higher relatively to their width, are in 
contact for the whole length of their sides (figs. 2 a, 7 a), and have 
distinct muscle-plates, which stand up around the opening of the 
small central funnel. These are largest and best developed in MW. 
suedicus, but are distinctly recognizable in Geinitz’s figure of I. 
Fischer. 

These features are eminently characteristic of the Pentacrinide 
and Comatulide, though I have seen no calyx of Pentacrinus which 
could be said to be precisely like that of Mesocrinus. But then no 
two Pentacrinus-calices that I have seen are precisely like one 
another; for they differ very much among themselves, not only in 
the relative development of the basals, but also in the shape and 
proportions of the outer surface of the radials and of their articular 
faces. At the same time it must be remembered that we are only 
just beginning to become acquainted with these modifications ; for 
neither species nor individuals of living. forms are at all abundant 
in collections, much less are they available ;for anatomical investi- 
gation. The calyx of Mesocrinus Fischer. (fig. 2) finds its nearest 
ally in that of P. Wyville-Thomsoni*. ‘It is in a side view that 
the resemblance is most evident, the chief point of difference being 
the greater height of the radials and of their articular faces in 
Mesocrinus. These features are more marked, however, in P. 
asteriay ; but in this species the basals are small, and do not meet 
laterally as they do in Mesocrinus. Except in this point the calyx 


supérieure ne servait pas de point d’attache” (oc. cit. p. 32). I regret that I 
cannot altogether agree with this opinion of the distinguished Swiss paleeon- 
tologist, who must have been unfortunate in some of the specimens he examined. 
The large concave surface in which the second radials of Apiocrinus rest is some- 
times an expansion of what in recent Crinoid#is the dorsal fossa lodging the elastic 
ligament. ‘This is well shown in de Loriol’s own figures of the ventral aspect 
ot the first radials of A. Meriani (pl. ii. figs. 4a, 5a), and also in d’Orbigny’s 
figure of A. Murchisonianus (pl. vi. fig. 7). The central end of this large fossa 
is bounded by the transverse articular ridge pierced by the opening of the 
central canal ; and rising up from this ridge so as to form a part of the rim of the 
central funnel are larger or smaller plates for the muscle- and ligament-fosse. 
In some specimens of Aptocrinus in the national collection these fossz are re- 
latively large and are separated by a well-marked ridge; but in most species 
they are greatly reduced in size, as is also frequently the case in Millericrinus. 
In the latter genus the dorsal fossa never reaches the enormous size that it does 
in Aptocrinus, and, though it is sometimes relatively large (as in some Comatule), 
it is occasionally comparatively small. 

I am not prepared to say, however, that no Apiocrinus had the first and second 
radials united otherwise than by muscles and ligaments. They must, in some 
cases, have been joined by a syzygy, e.g. Ap. Parkinsonti (VOrbigny, pl. v. fig. 6) ; 
but such cases are very anomalous; for in the other Articulate Crinoids 
syzygial union, though common enough between the second and third radials, 
never takes place between the first and second. Hyen in Marsupites there is a 
distinct articular facet on the first radials. 

* Journ. Linn. Soc., Zool. vol. xv. pl. xi. fig. 23. 

t Journ. Linn. Soc., Zool. vol. xy. pl. xi. fig. 21. 


THE UPPER CHALK OF SOUTHERN SWEDEN. aa 


of P. asteria is not unlike that of M. suedicus (Pl. VI. fig. '7), which 
has much more resemblance to it than to the calyx of any species 
of Millericrinus. 

Despite the resemblance of their calices to the Pentacrinus-type, 
the so-called Antedon Fischeri and its Swedish ally obviously belong 
to another genus than Pentacrinus. They lack the typical stem of 
this genus, with its verticils of cirrhi from the top downwards and 
the petaloid markings on the joint-faces. Neither does Mesocrinus 
possess the typical stem of Apiocrinus or Millericrinus ; but while 
chiefly resembling Bowrgueticrinus in the character of its stem, it 
differs from that genus and approaches /thizocrinus in the compara- 
tively small size of its upper stem-joints; and it especially resembles 
ft. Rawsoni in the relatively slight increase in the diameter of its 
calyx from below upwards. In neither case does the calyx pass 
gradually downwards into the thickened upper end of the stem, as 
it does in the typical Apiocrinide and in Bourqueticrinus, to which, 
in other respects, Wesocrinus has a considerable resemblance. 

The similarity of the articular faces of the stem-joints of the 
young Antedon Sarsii to those of Rhizocrinus, and the imperfect 
radicular processes that proceed from the lower part of its stem, are 
characters which connect the Comatulide with the Apiocrinide. 
The Pentacrinide, however, are sharply marked off from the latter 
family by the striking differences in the character of the stem. It 
is therefore of no small morphological interest to find a type in 
which the characters of a Pentacrinus-calyx are combined with those 
of a Bourgueticrinus-stem. It is quite possible that some of the 
stem-joints now referred to Bourgueticrinus may belong to other 
intermediate forms, the calices of which are still unknown to us, as 
that of Mesocrinus was but a few years ago. 


Prof. Lundgren has been kind enough also to intrust to me the 
description of a new Antedon, two specimens of which were found 
by himself and by Mr. J. Chr. Moberg in the Ignaberga Limestone 
at Balsberg, in the province of Scania, 8. W. Sweden. 


ANTEDON IMPREsSA, n. sp. (Plate VI. figs. 8, 9.) 


Centrodorsal a thin convex pentagonal disk with very indistinct 
traces of a small five-rayed impression at the dorsal pole. In the 
larger specimen the dorsal surface is somewhat flattened and tolerably 
free from cirrhi (fig. 8), but in the smaller and younger specimen 
there is but a very small cirrhus-free space (fig. 9,a). There are 
about 40-50 sockets arranged in three rather indistinct rows, with 
occasional traces of a fourth; but they are too much worn to show 
any structural details. 

The ventral surface is entirely obscured by matrix in the larger 
specimen, and is only partly visible in the smaller one. It is 
markedly concave; and its angles are somewhat produced upwards 
and outwards. In the middle line of each radial area is a median 
erooye, the central end of which is deepened and forms a radial pit. 


The outline of the central opening was probably decagonal; and 
L 2 


136 CRINOIDS FROM THE UPPER CHALK OF SOUTHERN SWEDEN. 


from its interradial sides there proceed five linear oblong basals, 
which do not quite reach the circumference. The outer end of each 
is marked by an oval oblong impression. 

Diameter, larger specimen 8 millim., smaller 5 mm. 

Height, larger specimen, 2 millim., smaller 1°5 mm. 

Remarks. I do not know of any Comatula, either recent or fossil, 
with a centrodorsal at all like that of this species, except Antedon 
Tourtie. Schliiter * figures five slight grooves radiating outwards 
from the radial pits of this last species ; but he makes no mention of 
them in the text. The ventral surface of its centrodorsal is slightly 
concave, with the angles raised; but this is far less distinctly the 
case than in A. impressa ; and the centrodorsal is half as high as it 
is wide, and bears four vertical rows of cirrhus-sockets, characters 
which distinguish it sharply from A. impressa. . 

The persistence of the basals in connexion with the centrodorsal 
rather than with the radials is also interesting. They are probably 
the original embryonic basals (or orthobasals), as in A. Lundgreni 
and a few other speciest. If they were merely basal rays connected 
with a rosette, some trace would have remained of such a connexion ; 
but I can find none. | 


EXPLANATION OF PLATE VI. 


Fig. 1. Mesocrinus Fischeri, Geinitz, sp., natural size. Copied from an 

amended drawing of the original, kindly lent by Prof. Geinitz. 

2. Calyx of Mesocrinus Fischeri, enlarged: a, side view; 0, dorsal 
surface. 

3-6. Stem-joints from Kopinge, 8. Sweden, probably belonging to 
Mesocrinus suedicus, all X 2: a, face views; 0, side views. 

7. Calyx of Mesocrinus suedicus, nov. gen. et sp., X 6: a, veutral 
surface; 0, dorsal surface; c, from the side. 

8 & 9. Centrodorsals of Antedon impressa, n. sp.: fig. 8. Dorsal sur- 
face of the larger specimen, X 4; fig. 9. Smaller specimen, x6: 
a, dorsal surface; 6, from the side; c, ventral surface. 


Discussion. 


The Prestpent expressed his sense of the value of the author’s 
communication, and hoped he would continue his work. 

Prof. Duncan expressed the same view. 

Prof. Srrrey said the variations of Bowrgueticrinus were very 
remarkable, both in form and in structure of the calyx. He had 
never seen one with the structure of that described, but had seen 
some approaching it. He eulogized the description given by the 
author, and expected from him important contributions to the 
knowledge of the Cretaceous Crinoids. 


* “Ueber einige astylide Crinoiden,” Zeitschr. d. deutsch. geol. Gesellsch. 
1878, p. 41, Taf. i. figs. 4-6. 

t Quart. Journ. Geol. Soc. 1880, vol. xxxvi. p. 550; Journ. Linn. Soe., Zool. 
vol. xv. p. 213. 


ON A BOULDER OF HORNBLENDE PICRITE. 137 


11. On a BovtvER of HorneienveE PrcritE near Prn-y-Carntsioe, 
Anetzsry. By Prof. T. G. Bonney, M.A., F.R.S., Sec. G. S. 
(Read January 5, 1881.) 


Last summer, through the kindness of Professor Rosenbusch, I was 
able to examine several specimens of picrite in: the Geological 
Museum at Heidelberg, and to study the rock in the field, near the 
village of Schriesheim, a few miles to the north of that town. In 
September I was walking with some students along the road which 
leads out of the village of Pen-y-Carnisiog northwards to Bwlyn 
(Anglesey), when I observed, in a field on the left, the fractured 
face of a boulder, in which a number of large crystals resembling 
augite, glittering in the sunlight, in a dull dark matrix, recalled at 
once the characteristic aspect of the Schriesheim picrite. The 
boulder had been broken, apparently rather recently, into three 
pieces, one much smaller than the other two ; and its volume must 
have been not much less than a cubic yard. In its weathered sur- 
face and toughness under the hammer it also resembled the Schries- 
heim rock. In both, the larger crystals (which are often about 
two thirds of an inch long) contain a number of dark serpentinous- 
looking enclosures, giving to the cleavage-faces an interrupted lustre 
somewhat resembling (except in the absence of a metallic gleam) 
that of bastite. The Pen-y-Carnisiog rock looks a little more de- 
composed ; but macroscopically the resemblance between my two 
specimens is so great that one could believe them to have been 
broken from different parts of the same mass. 


Part of a Slice from a Boulder of Hornblende Prerite near Pen-y- 
Carnisiog, Anglesey. (Magnified 30 diameters.) 
sa 


ATTAY 


4 ip \pbind “ 
, ym, 
Tin BY UALR \ NaN 
INS fF SON PAu) 
— +S D2 WP IS 1 
Mast SON ; Td 
< \ 


a. One of the grains of altered olivine. 
6, 6, b. Aggregated small crystals of hornblende, probably of secondary origin. 


138 PROF, T. G. BONNEY ON A BOULDER OF HORNBLENDE 


When the Pen-y-Carnisiog rock is examined microscopically, the 
difference between the two is rather more marked. In it the pre- 
dominant mineral is undoubtedly hornblende. This occurs under 
three forms :—(a) inumerable small acicular or blade-like crystals, 
in irregular tufted groups, forming a kind of ground-mass; these 
vary from a pale green tinge to almost colourless, and are generally 
very feebly, if at all, dichroic; the comparatively small extinction- 
angle shows them to be hornblende (actinolite) ; and there can be 
no reasonable doubt that they are of secondary origin; (6) small 
crystals, exhibiting often characteristic cleavages and even external 
forms (combinations of oP and oP), green-coloured and strongly 
dichroic; (c) large crystals (those mentioned above as supposed 
augite), including grains &c. of more than one other mineral, to be 
presently noticed. Augite occurs not unfrequently in almost 
colourless grains and crystals, some of which show a characteristic 
cleavage, and, in one or two cases, the characteristic outline (section 
of the combination of oP, oP‘, aP%s). The extinction-angle 
of the longitudinal sections of these crystals is large, ranging com- 
monly from 30° to 40°. They usually occur interspersed in a dull 
olive-green serpentinous or chloritic mineral. No olivine can now be 
recognized with certainty in the slides; but there are a number of 
irregular grains associated in them with the other minerals, and 
abundantly included (these being of smaller size) in the large horn- 
blende crystals, which there is every reason to believe are pseudo- 
morphs after the former mineral. Opacite and rounded crystal- 
line grains resembling magnetite abound in these, often clustered 
together or lining roughly parallel cracks, which remind us of the 
uregular cleavages of olivine; from these a brown staining often 
extends inwards for some little distance. ‘The pseudomorphic con- 
stituents vary considerably: sometimes, as described above, they 
are a brownish or yellowish green, of filmy granular structure, act- 
ing upon polarized light, but not greatly changing their colour; 
sometimes they exhibit a finely-speckled aspect, as though aggre- 
gates of extremely minute grains or folia of a mineral that acts 
strongly upon polarized light; sometimes they are aggregated small 
folia of a mineral resembling tale; and sometimes a clear isotropic, 
or nearly isotropic, mineral, such as is common in ordinary serpen- 
tines. Two or three of the inclusions in the larger hornblende 
crystals exhibit a radial aggregate structure with the usual black 
crosses. Though most of the above microlithic products are not the 
most usual replacements of olivine, I have seen them occasionally 
in my studies of peridotites, and have no doubt that this mineral 
was formerly present in the Pen-y-Carnisiog rock (probably a 
rather ferriferous variety). Magnetite is not uncommon as an in- 
clusion in the larger hornbiende crystals; the slide also contains a 
little altered brown mica, and a few small crystals of apatite. 

With regard to the large black crystals already mentioned, 
although, from their optical properties, one cannot but regard them 
as a brown hornblende, I doubt much whether this is not due to 
subsequent paramorphic change, and believe that they were for- 


PICRITE NEAR PEN-Y-CARNISIOG, ANGLESEY. 139 


merly a true augite. The extinction-angles are generally less than 
20°; but in one slide are two crystals which, though dichroic, give 
angles of about 30°. The larger crystals of the Schriesheim picrite 
agree with augite in their feeble dichroism and general appearance ; 
but the results of several measurements of the extinction-angle are 
less than 20° in the case of the largest crystal, while in a smaller 
one they are over 30°; and a very considerable quantity of horn- 
blende, similar to the varieties (a) and (6), is present in the body of 
the slide. A little olivine has escaped change; and the structure of 
the rest is rather more characteristic. Still, except that a mica is 
decidedly more abundant in the Schriesheim than in the Pen-y-Car- 
nisiog rock, the main difference appears to me to be, that the latter 
has undergone more alteration than the former, so that it, too, has 
once been a true picrite. 

Another rock is very abundant in boulders in this district of 
Anglesey, which, at first sight, has some resemblance to the picrite, 
though less porphyritic in structure. Closer examination, however, 
shows that felspar is always a constituent of this, though it is often 
not very conspicuous. I have examined a specimen microscopically, 
and find it to consist of a green hornblende, an altered felspar, a 
brown mica more or less changed, apatite, and a chloritic mineral. 
The hornblende and felspar are rather irregular in external form: the 
latter is almost wholly replaced by microlithic products; but one or 
two grains still retain sufficient traces of their original structure to 
show they have been plagioclastic. It is somewhat singular that a 
rather similar rock, but with more brown mica and better-preserved 
felspar, occurs at Schriesheim, within a short distance of the picrite, 
also intrusive in the granite. This is named a Labrador-diorite by 
the German petrologists. 

The only other instances known to me of the occurrence of picrite 
in the british Isles are two in Fifeshire, described by Prof. Geikie 
in his excellent monograph on the Carboniferous Volcanic Rocks of 
the Firth-of-Forth basin*, and one described by myself, to which, 
as I had at that time never examined a typical picrite, and had 
failed to obtain a very clear notion of the rock from such descrip- 
tions as I had seen, I did not venture to attach the name. In this, 
however, olivine (still very fairly preserved) is the dominant mi- 
neral; so that it comes nearer to a normal peridotite. The rock was 
collected many years ago by Professor Sedgwick, near Penarfynydd, 
in the Lleyn peninsula; and Mr. HK. B. Tawney, who lately visited 
the locality expressly to search for it in sitw, failed to find it, and 
believes that the specimen must have been taken from a boulder. 
Here also, as described by Mr. Tawney, are olivine-diabases and 
hornblendic diabases. 

I have ventured to draw especial attention to this Anglesey spe- 
cimen, in the hope that some geologist may succeed in discovering 
a like rock in situ. As the picrite just described is so uncommon 
and of so marked a character, we might assume with much confi- 


* Transact. Roy. Soc. Edinburgh, vol. xxix. p. 437. 
+ In a paper by Mr. E. B. Tawney, Geol. Mag. Dee. 2, vol. vii. p. 208. 


140 . ON A BOULDER OF HORNBLENDE PICRITE. 


dence (supposing no physical difficulties presented themselves) the 
Pen-y-Carnisiog boulder to have been derived from that parent rock. 
This would afford us most valuable evidence as to the direction in 
which the agent of transport (doubtless ice) had formerly moved. 
The south side of the Lleyn peninsula, even if the rocks corresponded 
(which they do not), seems excluded by physical considerations ; 
but I may remark that, though I have examined many boulders in 
Anglesey, I have failed as yet to identify any of them with rocks . 
from the Lake District or from Scotland, and think that we must 
look to the mountains of North Wales for the home of those which 
cannot be found in Anglesey itself. 


ON THE GEOLOGY OF CENTRAL WALES. 141 


12. The Gronoey of Cenrran Wares. By Watrer Kezerine, Esq., 
M.A., F.G.S., Lecturer on Geology in the University of Cam- 
bridge. With an ApPENDIX on some new Species of CLADOPHORA, 
by Coartes Lapwortu, Hsq., F.G.S. (Read June 23, 188U.) 


[Puate VIT.] 


ConTENTS, 


Part I. More distant sections. 
Introduction and Historical. (1) Llanbrynmaer area. 
Illustrative section from Aberystwyth (2) Dovey ~wallsy area. 
to the Devil’s Bridge and Plyn- (3) Corris area. 


laniaion. (4) The Cardigan area. 
(1) The Aberystwyth Grits. General results. 
(2) The Metalliferous-slate Group. Part IIT 
(3) The Plynlimmon Group. art 1h. 
Paleontological evidence. 
Part II. General summary. 
Brief notes upon other sections. 
(1) Aberystwyth through Pont APPENDIX. 
Erwyd to Builth. Appendix by Mr. C. Lapworth, F.G:S., 
(2) Llandeilo to Aberaeron. on new Species of Cladophora. 
(3) Rhyader to the Teifi Pools. 
General results. 
Part I. 


Tue following communication contains some of the results worked 
out in frequent field-excursions during three years’ stay at the 
University College of Wales, Aberystwyth. In many of these ex- 
cursions I was accompanied by my students of the College; and to 
them I am indebted for much help both in the museum and the 
field*. Our principal field of work was, naturally, within a radius 
of some fifteen to twenty miles around the town of Aberystwyth ; 
and most of this country we have carefully searched and traversed 
through and through. The more distant areas to the south of 
Cader Idris, at Llanbrynmaer, Llandovery, Llandeilo, Cardigan, &c., 
have also been visited with a view to the determination of the 
extent and variation of the Cardiganshire rock-groups, and the dis- 
covery of their stratigraphical relations to other and better-known 
geological horizons. 

Little has yet been done by geologists to elucidate the structure 
of this part of Wales: while the most careful labours of our greatest 
authorities have been devoted to the study of the northern counties 
and eastern borders of Wales, and also, of late years, to the south in 
Pembrokeshire, this barren and chaotic area of Mid Wales has been 
always neglected, and is even now very rarely touched with the geolo- 
gical hammer. For the bibliography of our subject we have therefore 
but little to say, the most important contributions being :—the work 

* In particular, I have received much assistance from my former pupils 


Mr. T. Roberts, now of St. John’s College, and Mr. E. Evans of Sidney College, 
Cambridge. 


142 WALTER KEEPING ON THE 


of our great leader Sedgwick, published in the Society’s Journal in the 
year 1847 (vol. ii. p. 150); a slight account of the Dol Fan conglome- 
rate, and a description of the Lampeter worm-trails in Murchison’s 
‘Silurian System, pp. 316, 317; some scattered remarks by Mr. 
Salter and Sir Roderick Murchison in ‘ Siluria, and by the former in 
the Cambridge ‘Catalogue.’ The Pont Erwyd district is referred to 
in Symonds’s ‘ Records of the Rocks,’ pp. 132,133; some Graptolites 
from Aberystwyth are recorded by Mr. J. Hopkinson, F.G.S., in the 
‘Journal’ of the Quekett Microscopical Club, vol. 1. p. 151; and 
the mineral veins of Cardiganshire are described by Mr. Warington 
Smyth, M.A., F.R.S., in the ‘Memoirs of the Geological Survey,’ 
vol. ii. part 2, p. 485. 

The present communication, although worked out in a close net- 
work of foot-routes in parts of the district, and cross traverses over 
the rest, is offered only as a first reconnoissance report, which must 
be followed by many labours betore the structure of this great and 
complicated district can be thoroughly mastered. Some general 
order amidst the whirl of contortions, however, is now made clear, 
such as the general succession of the Aberystwyth, Metalliferous, 
and Plynlimmon groups, and the great Plynliimmon synelinal. The 
enormous apparent thicknesses of the rocks are shown to be in part 
due to a series of inversions; and, further, the interpretation and 
correlation of the beds are greatly helped forward by discoveries of 
fossils, mostly Graptolites, in a number of localities, which afford 
excellent data for comparison with the more distant Scotch and 
Cumbrian Silurians. 

Even Sedgwick’s work (1846) was but of a very superficial kind ; 
for he tells us, ‘“‘ I profess not to know well this most contorted and 
perplexing country.” He makes four rock groups, namely the 
(1) Aberystwyth, (2) Plynliimmon, (3) Upper South-Wales Slate 
with the Rhyader Slate, and (4) Cambro-Silurian groups, which 
appear in ascending order from west to east; but he adds, “the 
sections are singularly contorted, the groups ill defined, and the 
actual order of superposition obscure.” His first group I still main- 
tain under the name of the Aberystwyth grits; but nearly all the 
remaining rocks in the line of section might, I believe, have been 
included in his second great series—the Plynlimmon ‘group. On 
the other hand, I have considered this great Plynlimmon group of 
Sedgwick under two distinct headings, namely the (inferior) slate 
series or Metalliferous-slate group, and the overlying Plynlimmon 
grits, so that we now have the following succession of deposits :— 

(3) The Plynlimmon grits, forming a line of high country 
in the centre of Wales, including Plynliimmon. 


(2) The Metalliferous-slate group, forming a broad zone of 
Cardiganshire contorted country on each side of (8). 
group. (1) The Aberystwyth grits, best developed between Aberyst- 
wyth and Aberayron. 
Nos. (1) and (2), being closely bound together by their fossils, are known 
together as the Cardiganshire group. 


For an illustrative section of our area the best I can offer is that 
from Aberystwyth, through the Devil’s Bridge, to the Plynlimmon 


143 


GEOLOGY OF CENTRAL WALES, 


Fig. 1.—Section from Aberystwyth to the Devil’s Bridge. 


(Horizontal scale, $ ch to a mile.) 


Ww. BE. 
Aberystwyth. Devil’s Bridge. The Arch, 


RAS Zz 
PP pos 
Vi 


S 
= 


PUG 


Aberystwyth grits. Metalliferous-slate group. 


The continuous dotted line indicates the probable course of the Cefn Hendre and the Devil’s-Bridge fossil-zone. 


. 


Fig. 2.—Continuation of the Section in fig. 1, from Owm Ystwyth, through Craig Lluest, to Rhyader and Gwastaden. 
(Horizontal scale, 1 inch to a mile.) 


W.<—e@ m—>T, W.N.W.<—= Onc 


Popty. Cwm Ystwyth. Craig Lluest (about 1200 feet), Pont ur Elan, Pen Rhiev Wen. Gwyn Llyn. Rhyader, Gwastaden, 


Za 
A 
(j---~---+--=- 


i 


Large-llag series. Grits and paper shales. Pale slate, thin Shaly slates. Hard pale Conglo- 
The Plynlimmon Synclinal. grits and shales. slates. merates. 


144 WALTER KEEPING ON THE 


Mountains (figs. 1 & 2). The lowest beds form the cliffs at the coast, 
while the highest go to form the central mountains. All these rocks 
are amazingly contorted ; and we find the intermediate or Metallifer- 
ous group especially thrown into frequent and violent twistings, 
with, in many places, actual inversion. A continuation of the section 
eastward, beyond Plynlimmon (fig. 2), shows a similar appearance of 
the rock groups in reversed order of outcrop and with contrary dips, 
thus showing that Plynlimmon, like its more ancient and greater 
fellow Snowdon, stands in a geological valley or synclinal. We 
proceed to the detail of this section. 

1. The Aberystwyth grits consist of beds of hard, compact, dark 
grey grit or greywacke, and dark shales, rabs, and imperfect slates 
in strikingly regular alternation, as may be seen in the ordinary 
photographs of the Aberystwyth cliffs. 

The grey grits or greywackes are of great sameness and regularity 
both in structure and composition around Aberystwyth, being hard, 
grained rocks, often felspathic, regularly and sharply jointed. A 
cross fracture often shows a remarkable contortion in the lines 
of lamine, this being, I believe, mostly of subsequent “‘ concre- 
tionary” origin; many of the beds themselves are also, in part, 
of the same concretionary growth. ‘The beds measure very con- 
stantly about 4 to 6 inches in thickness; and their under sur- 
faces exhibit an abundance of raised markings, which are irregular 
or tortuous, branching, net-lke or worm-like, these being also, in 
part, of concretionary origin. 

The argillaceous partings are usually of about the same thickness 
as the greywacke-beds, sometimes thicker (especially to the east and 
south), sometimes thinner (as In many places around Aberystwyth). 
Most of their varieties are the result of subsequent metamorphic 
changes acting differently upon the rock according to a slight 
diversity of original constitution, or depending upon slightly dif- 
ferent mechanical conditions. Thus have been produced the various 
forms of shivery shale, large platy shale, rubbly rab* of various 
forms and sizes, soft shaly slate, and even very well-marked regular 
slate in the Aberystwyth district. They are uniformly of dark 
colour, and never greatly indurated. Lenticular nodules with ‘‘ cone- 
in-cone”’ structure are of frequent occurrence both in this and the ~ 
following (or Metalliferous) series. 

Fossils—Fucoidal and worm-like markings are of frequent and 
wide-spread occurrence throughout this series, appearing for the 
most part in the form of raised markings upon the under surfaces of 
the grits. I have also found Graptolites in several localities. 


Quarry at Cwm, on the south side of the Clarach Valley. 
Monograptus Sedgewickii, Portt. Monograptus tenuis, Portl.? 


—— Clingani, Carr. Buthotrephis, small species. 
lobiferus, M/‘Coy. 


* A rab is a fine-grained rock, usually argillaceous and not indurated, which 
readily breaks up into a rubble of cuboidal or prism-like fragments. 


GEOLOGY OF CENTRAL WALES. 145 


From the Bryn-y-Carnau Quarry near the old Water-reservoir, 
Aberystwyth. 
Monograptus tenuis ? 
Dictyonema delicatulum, Lapw. 
(n. sp.). 


But the most important zone of fossils is found somewhat higher 
up in the series, exposed in a quarry formerly worked for building- 
stone and road-metal, in a field below Cefn Hendre, where we have 
collected 


Monograptus Sedgewickii, Porél. 
, var. distans, Port. —— tenuis, Portl.? 
—— Clingani, Carr. Oxthoceras, sp. 

lobiterus, M*‘Coy. Calymene. 
turriculatus, Barr. | 


Monograptus Sedgewickii, Port. 
crenularis, Lapw.? 
lobiferus, M/‘Coy. 


Monograptus Hisingeri, Carr. 


Many of these fossils were found in a set of thin, dark-grey, mica- 
ceous flags (the large-flag series), which are sometimes to be obtained 
of large size (4-6 feet square). 

Dips and Foldings.—Many rock-foldings, some of them very 
violent and accompanied by fractures, are seen in our line of 
section, good examples being exposed near the second milestone 
from Aberystwyth ; so that for a mile anda half along the Upper 
Devil’s Bridge-road we appear to have only the same set of beds, 
repeated again and again by a number of rock-foldings (see section, 
fig. 1). But beyond this the easterly dip becomes more constant, 
and we appear to be traversing the outcrop of a continually ascend- 
ing series. 

2. The Metalliferous-slate Group.—In the Cefn-Hendre quarry, 
only about a mile and a half east of Aberystwyth, we already find a 
larger proportion of shales to grits than in the coast-section around 
Aberystwyth ; and this change becomes still more marked in another 
quarry on the road-side further on down the hill towards Gogerddan. ° 
The same gradual though not perfectly regular disappearance of 
the grits to the east may be seen in our present section along the 
Devil’s-Bridge road, as, indeed, in any other of the east and west roads 
from Aberystwyth*. 

A change in the character of the argillaceous rocks appears in 
regular correlation with the loss of the grits. They become more 
and more indurated and cleaved, until, as the boundaries of the 
grit-series are reached, the normal cleavage of the district (striking 
N.N.E. and 8.8.W.) is found even in the partings between thick 
erit-beds. Thus gradually do we enter into the territory of our 
second series—the Metalliferous Slates. 

The hills of this district are barren and desolate, even more so 
than in the Aberystwyth-grit country ; but they are decidedly more 
rounded and regular in their contours. Here, in the uplands, is a 


* The limit of the grit series may be placed, as indicated by the yellow dots 
upon the Survey Map, at about three miles and a half east of Aberystwyth. In 
my pocket-book I find “‘at three miles from town, grits fewer and thinner than 
at Aberystwyth,” at four miles ‘‘a few grit-beds as much as 6 inehes thick,” and 
at five miles “still a few thin grits.” 


146 WALTER KEEPING ON THE 


vast dreary mountain-region of bare sheep-walks scantily marked 
out into districts by poor stone walls and wire fences, with much 
waste bog and peat land. The rocks are of uniform pale blue and 
grey colours, varying from small papery shales (rarer) to large 
irregular platy shale or regularly cleaved slates, also (in some areas) 
much indurated small slate rock cross-cut into fragments by frequent 
bedding- and joint-planes. Some zones of softer rab, like that of the 
Aberystwyth group, and a pale mudstone rab are sometimes found, 
especially near the junction with the grit series. 

As we reach the central mining district, some seven miles east of 
Aberystwyth, beds of hard, pale, indurated slate-rock with frequent 
bed-bandings occur; and in the immediate neighbourhood of the 
mineral veins such induration is nearly always well marked. 

Occasionally a bed of grit, 2-6 inches thick, occurs in this series ; 
but such occurrences are very rare, so that the building- and road- 
stones of the central ‘‘ Metalliferous” country have to be carried 
for miles, either from the Aberystwyth grit quarries in the west, or 
from the Plynlimmon group further east. 

Thin ferruginous gritty bands, about 4 inch thick, however, are 
more frequent, some of them being highly micaceous. Many such 
are seen between the fourth and fifth milestones and around the 
ninth milestone on the Devil’s-Bridge road. 

Rock-foldings.—Several excellent exposures of the contortions in 
this slaty series are seen along our line of section. There is a neat 
little synclinal at seven miles, an anticlinal towards the eighth mile- 
stone; and several folds may be detected about nine miles east of 
Aberystwyth; but overriding these foldings the prevailing dip is 
seen to be clearly and determinately to the east. 

Fossils are rare in this series. The curious branching structure, 
Nematolites Edwardsii, Keep., occurs at Ty Llwydd; and I have 
found the Fan Algal (Buthotrephis major, Keep.), the Nematolites, 
and worm-trails (Werettes) in several other places. But in other 
areas, especially at Cwm Symlog and near Machynlleth, out of our 
present line of section, a rich Graptolite-fauna has been discovered, 
and will be described later on. 

The Devils Bridge, great Inversion of the Rocks (fig. 1).—As we de- 
scend the hill to the Devil’s Bridge we pass over pale, hard, shaly 
slates* with thin gritty bands about 1 inch thick, marked with worm- 
tracks (Nereites Sedgwickit), impressions of the Fan Angal (Buthotrephis 
major, Keep.), and Nematolites Hdwardsi, Keep. Coarse roofing- 
slates have been worked about half a mile west of the hotel. These 
rocks belong to the Metalliferous-slate series; but coming to the 
waterfall we meet with a set of alternating thin grits and large 
pale shaly slates, together with some large flags of laminated grit, 
with fossils exactly resembling those already noticed from Cefn 


Hendre. 
Many of the grit bands are thin and little jointed, so that large 


* Although the splitting of these rocks is so irregular and shale-like, yet the 
planes of division are of subsequent cleavage origin ; therefore I call them shaly 
slates rather than slaty shales. 


GEOLOGY OF CENTRAL WALES. 147 


flags may be readily extracted, from which feature I have called this 
zone the ‘‘ Large-flag series” (fig. 2). The dip is to the west, running 
under the slaty series. Fossils may be found here in the open quarry 
above the Devil’s Punch-bowl, and along the sides of the waterfall 
itself; but all the Dendroid Graptolites (Cladophora) are from a 
band above the iron bridge in the waterfall-grounds ; while some 
other beds below this bridge are so crowded with Crinoid ossicles as 
almost to merit the name of Limestone. 
Fossils from the Devil’s Bridge :— 


Monograptus Sedgewickii, Port/.? Odontocaulis Keepingii, Lapw. 
—— spiralis, Geinitz. (n. sp.). 

turriculatus, Barr. Chonetes lzevigatus, Sby.? 
Dictyonema venustum, Lapw.(n.sp.). Orthis, sp. 
—— corrugatellum, Lapw. (n. sp.). Other fragmentary Brachiopods. 


Calyptograpsus (?) plumosus, Lapw. Phacops, n. sp. 


n. sp.). Fragments of Encrinites. 
Rhizograptus (?) digitatus, Lapw. Nereites Sedgwickii, Murch. 
(nu. sp.). Myrianites tenuis, I‘ Cov. 


—— ramosus, Lapw. (n. sp.). 


Now in this list of fossils the species of Graptolites are seen to 
correspond with those of Cefn Hendre, the occurrence of Mono- 
graptus turriculatus in both places being an especially important 
fact, this being a species of very limited range. 

And we have seen that they occur in identical rocks in the two 
places; I therefore cannot doubt that these are really one and the 
same set of beds, seen in the upper part of the Aberystwyth grits at 
Cefn Hendre and reappearing at the surface in an anticlinal at the 
Devil’s Bridge. | 

This conclusion, however, is in direct antagonism with the strati- 
graphical appearances ; for, notwithstanding the numerous folds, the 
predominance of the easterly dip is most determined and impresses 
itself strongly upon the mind. 

Being convinced of these appearances, it was determined to test 
the thickness of the beds by actual measurement; and our exact 
observations and calculations, made at more than 100 exposures 
along the Devil’s-Bridge road from Aberystwyth, strikingly con- 
firmed our earlier impressions, giving, indeed, a result of nearly four 
miles thickness of strata (3 miles 1612 yards). In such apparent 
conflict of evidence, and in the absence of large faults, the only 
reasonable explanation is that the original natural order of the 
rocks has been destroyed by the formation of a great inversion, or 
rather, as I believe, a series of inversion folds in the Metalliferous- 
slate series (see fig. 1). And indeed this interpretation, in con- 
formity with the fossil evidence, is independently almost demanded 
to explain away the enormous apparent thicknesses of similar rock- 
beds as measured from their present dips. 

We may then safely conclude that in the Devil’s-Bridge rocks we 
are again upon the upper part of the Aberystwyth-grit series. 

Continuing our section eastward over the hill through the Arch 
to Cwm Ystwyth (fig. 2), we still traverse a series of shaly slates of 
the “‘ Metalliferous” type, with here and there thin grits apparently 


148 WALTER KEEPING ON THE 


belonging to the Large-flag series. The presence of Fucoidal mark- 
ings, including the Net Algal (Reteofucus extensus, Keep.) and Nema- 
tolites at one mile and a half from the Devil’s Bridge, also serves to 
indicate our proximity to the Aberystwyth-grit series. The general 
apparent dip is clearly eastward, with some foldings. 

Descending beyond the Arch towards Kglwys Newydd the only 
matter of special interest is the occurrence of a thin seam of rotten- 
stone (a decomposed limestone) which may perhaps correspond with 
the crinoidal zone in the Devil’s Bridge. Such rocks are of very 
rare occurrence in Mid Wales, so that it is a popular saying that 
‘‘ there is no lime in Cardiganshire.” 

The rocks around Popty, near Eglwys Newydd, are again seen to 
be very much of the Devil’s-Bridge type; and I am inclined to think 
that a set of rock-folds with reversals have here brought that fossil 
zone near to the surface again. We now reach a region of manifest 
great rock-foldings; and as we ascend Cwm Ystwyth a number of 
exposures display an important westerly fold in the rocks. The 
beds appear still to belong to near the junction of the Aberystwyth 
grits with the Metalliferous-slate group. 

Next we pass through the rich metalliferous district of Cwm 
Ystwyth with its network of mineral veins in the Metalliferous- 
slate group, beyond which, at Blaen y Cwm, thicker grits (about 
2 feet) with east and south-east dips come in amongst the slates; 
and these latter become replaced over a considerable area by dark 
shaly slate, rab, pencil-rab, and other softer forms of the argillaceous 
rocks. 

Thick grit-beds, which are cleaved, next form a prominent feature 
at Craig Lluest (fig. 2); and these I regard as belonging to the 
lower part of our upper grit-series—the Plynlimmon group. This 
series is better developed in Plynlimmon itself, to which we must 
refer (infra, p. 156) for a more detailed description. The dip is 
eastward. 

Some three quarters of a mile beyond the top of the pass*the dip 
changes, a westerly slope setting in; and this remains well sustained, 
though with some east foldings, on to the hills west of Rhyader. 
Here, then, we see the eastern half of the great axial synelinal of 
this part of Wales—a great fold in which Plynlimmon lies; ‘but 
its apparent magnitude is greatly exaggerated by the phenomena 
of reversed dips. The lithological details of this part of the section 
in its frequent slight variations, but general monotony, till we 
reach Rhyader, would occupy much space and be of little value. 
Some pale shaly slates approaching the character of the Tarannon 
shales, and thin gritty bands, are seen on the dreary bog- and moor- 
land of the higher plateau and on the east of the pass, where also 
some of the grits exhibit irregular fucoidal markings and the Net 
Algal (Retiofucus extensus, Keep.) upon their under surfaces. 

One good anticlinal fold in “ Metalliferous slates” with some 
erits has its axis about one mile and a half east of Aber Gynwy; and 
a synclinal in the slaty series is indicated lower down on the road 
to Pont ur Elan. ‘The well-sustained general westerly dip of the 


GEOLOGY OF CENTRAL WALES. 149 


great series of thin-bedded slate, pale- or dark-coloured, seen in the 
ascent from this bridge to the top of the high part of Rhyader, 
appears to show these rocks as in regular serial continuity with 
the beds passed over on the great slope next to the west, the whole 
forming the vast westerly fold complementary to the great easterly 
dip of the Metalliferous-slate series between Aberystwyth and the 
Devil’s Bridge. And just as the latter is now proved to be not one 
continuous series, but reallya much smaller group affected by fre- 
quent inversion, so we are, I think, justified in adopting a like inter- 
pretation for these beds to the east of the Plynlimmon axis. The 
correspondence of the rocks on the two meridional sides of this axis 
is further illustrated as we reach the eastern edge of the great plateau 
above Gwyn Llyn (fig. 2), where we find a zone of pale banded and 
indurated slate rock identical with the Strata Florida slate rock used 
in the Aberystwyth stone pier. These pass under a series of grits 
and conglomerates in the beautiful valley of Cwm Elan. A little 
further south these latter beds reappear in the Gwastaden grits 
and conglomerates of the hills east of Rhyader. Thus our accumu- 
lated facts support the view that there are two distinct series of 
grits in Cardiganshire, separated by the Metalliferous-slate group. 
The Rhyader grits and conglomerates appear to be nearer to the 
Plynlimmon than the Aberystwyth group. 

The Aberystywth Grits, extent, varrations, Jc-—A glance at the 
yellow-dotted area upon the Geological Survey Map will show that 
the Aberystwyth grits form a crescent-shaped patch of country in 
the centre of the western Welsh border, extending from near Borth, 
some five miles north of Aberystwyth, to Traeth Bach, south of 
Llangrannog, a distance of thirty miles, and with a maximum 
breadth, at Mynydd Bach, of nine miles. 

A set of characteristic surface-features marks off its boundaries 
with tolerable distinctness, its barren or gorse-covered ridgy hills, 
elongated along the line of rock strike, and with minor sets of 
parallel crests and ledges (sometimes forming sets of small step- 
like structures or inclined ribs in the lines of more durable grit-beds), 
giving to the group an almost unmistakable appearance*.. 

The general characters of the rocks have already been described 
in our Deyil’s-Bridge line of section; and the variations from those 
types are neither numerous nor very striking. The greywacke and 
erits are very constant in grain, never becoming conglomeratic; but 
some coarser varieties are found in a few localities—for example, 
around the Hiddwen lakes, Mynydd Bach, and at Pen Craig, north of 
Llanilar. In these places the greywacke is almost granitoid in 
appearance, the quartz grains being large, flakes of mica common, 


* It was pointed out long ago by Sir Henry de la Beche that the same type 
of physical features appears in the area of the grits of Penrhyn ddu in the 
Lleyn peninsula, which are marked Lower Cambrian upon the Survey Map. 

To me this resemblance also appeared very striking. The Penrhyn-ddu beds 
are totally unlike any of the Longmynd group known to me; and I cannot 
believe they are of such an age; nor do I think with de la Beche that they are 
of the same age as the Aberystwyth beds, but regard them as an exceptionally 
developed type of the Tremadoc series. 


Q.J.G.8. No. 146. M 


150 WALTER KEEPING ON THE 


and crystals of felspar, sometimes perfect but more often fragmen- 
tary, very numerous and conspicuous. These constituents are held 
together in the dark argillaceous greywacke matrix. The rock often 
closely simulates a volcanic ash*. 

In thin sections, examined under the microscope, the Pen-Craig 
rock exhibits irregular angular and subangular fragments of quartz 
and felspar, the interstices filled with the dark, opaque (argillaceous) 
matrix. The quartz is somewhat cracked, and includes numerous 
cavities and minute spicular microliths; these latter are grouped 
into wavy stratoid zones, between which the principal cracking of 
the quartz runs. ‘The felspar is mostly in the form of.very angular 
crystal-fragments, usually much decomposed, and of powdery ap- 
pearance; but some of the better-preserved fragments show the 
characteristic ribbon banding of the Plagioclase group when ex- 
amined with polarized light. 

The ordinary finer-grained beds are usually of darker colour than 
these, and contain less felspar; they are, however, for the most part 
quite similarly constituted, the felspar crystals often being readily 
recognizable when decomposed into irony spots over the weathered 
portions of the rock. Paler thin bands, very compact in texture, 
occur in the southern part of the district: around Llangrannog. 

The presence of crystals of iron-pyrites occasionally gives a marked 
feature to the grits; and at Pen Craig, Llanilar, and in the Garthen 
valley, Eglwys Fach, some beds occur beautifully studded with these 
brassy cubes. 

The ordinary thickness (4—6 inches) of the beds is very constant 
throughout the area of the Aberstwyth grits; but more massive beds 
(1-13 foot) occur under Allt wen and at Aberaeron, in the Garthen 
valley, and at Llangwyryfon (24 feet). At Cefn Coch, Pen Pegwyns, 
and along the coast south of Aberystwyth still thicker beds are seen 
(3-4 feet); they are worked for building-stones. At the southern 
limit of the series, near Llangrannog, the grits become thin, irreguler, 
and inconstant, thus gradually dying away, to be replaced by the ar- 
gillaceous slate-rocks. | 

Two structural peculiarities are very characteristic of the Car- 
diganshire grits—namely, the remarkable contorted lamination seen 

on a fractured surface, and an abundance of fucoid markings, to- 
- gether with strange-looking ripplings, ridgings, volutings, and other 
raised forms of structure found upon the undersurfaces of the grit- 
beds?. The rock is jointed, sometimes into large blocks good for 
building- and flag-stones, but often much more closely into regular 
oblong or rhomboidal fragments. At Allt-wen cliff and elsewhere 
it is divided up into oblong, prismatic, or rudely lozenge-shaped 
fragments a foot long, forming a kind of coarse grit-rab. 


* This abundance of felspar crystals, so general in the Silurian rocks (Upper 
Silurian of North Wales, South Wales, and the Lake-district), points to the 
neighbouring presence of a vast mass of early, perhaps primeval, igneous 
rocks as the great source of sediment-supply in Silurian times. 

+ It is proposed to give a more particular account of the rock-structure of 
Cardiganshire in a separate paper. 


GEOLOGY OF CENTRAL WALES. 151 


Little need be added to what we have said on the alternating 
argillaceous beds. The rab-type is best seen upon tho Allt-wen 
cliffs, south of Aberystwyth, where also au elongated type, serving 
very well as rough pencils, forms the ‘“pencil-rab;” these beds 
commonly appear much more solid in the heart of the rock, the 
fragmentary structure only becoming developed after exposure to 
weathering. 

The cleavage phenomena here are often of peculiar interest from 
their frequently incomplete, half-developed character. The dark shaly- 
. looking rock of Constitution Hill, Aberystwyth, would not at first be 
suspected of a slaty structure ; but Specimens may be gathered with 
the cleavage seen distinctly cutting across the stripe- lines ; and many 
of the dark, soft, shaly rock-beds of the neighbouring cliffs, when 
seen in situ, will be found to have their shetsme) ies running 
across the bedding. 

A pale, homogeneous, hardened mud, or pale mudstone, splitting 
with curved, conchoidal surfaces into 2-3-inch blocks, is of additional 
importance, as probably marking a zone; I have found it some half 
a mile north-east of Nant Kos, at Pen Craig, Llanilar, on the road-side 
N.N.E. of Llanbardarn, and at five miles east of Aberaeron. This 
appears to indicate a distinct zone near the top of the Aberystwyth 
grits. The same kind of rock occurs at Llyn Fyrddyn Fawr, and at 
Rhos Rhydd, near Llantrisant, where itis again not far removed 
from the thin-flag series—another zone at the top of the Abery- 
stwyth grits. 

The Large-flag series.—This series, which belongs to the upper 
part of the Aberystwyth group, is best marked to the east, in the 
neighbourhood of the Devil’s Bridge and Llantrisant, where its 
presence is soon made manifest in the construction of piggeries and 
other low huts, whose sides and roofs are covered with the flags. 
These are grit-beds, about 1 inch thick, but little jointed, so that 
large slabs, frequently 4 and 6 feet square, are commonly extracted 
for use in the neighbourhood. ‘The rock is fine-grained, and usually 
exhibits a complete contortion in the lines of lamine; altogether 
it much resembles some of the grit-beds of the middle Lingula- 
flags of North Wales. The surfaces are gencrally undulated, and 
often show true ripple-marks, over which we find Graptolites and 
other fossils spread out: the Devil’s-Bridge Dendroid Graptolites 
occur in this series. 

The further extensions of these beds, north and south, are seen 
around Pont Erwyd and south of Llantrisant; and the same series 
seems to be brought up to day by folds, on the east at Hglwys - 
Newydd, and to the west near the head of Cwm Symlog. Bands of 
rottenstone (decomposed limestone) occur at the Devil’s Bridge, near 
Eglwys Newydd, and again in the hills some four miles east of Tal 
y bont. 

Around Aberystwyth these flags are not so well developed; but 
they may be recognized at Cefn Hendre, as already described, where 
they are put to the usual purpose of hut-making for the work- 
men’s shelter. 

M 2 


12 WALTER KEEPING ON THE 


Fossil localities, other than those already given, are :— 
Parson’s Bridge, and Temple Mine, south of Pont Erwyd. 


Monograptus lobiferus, MM‘ Coy. Myrianites M‘Leayi, Murch. 
Climacograpsus scalaris, His. Buthotrephis major, Keep. 
Nereites Sedgwickii, Murch. 


Cefn Coch, near Aberystwyth. 
Monograptus Sedgewickiu, Pordd. | Retiofucus extensus, Keep. 
Buthotrephis major, Keep. Paleochorda tardifureata, Keep. 
minor, Keep. | 

Constitution Hill, Aberystwyth; Wern Grug, Llanilar; Mynydd 
Bach; Plas Crag, Llanbardarn, and other places have also yielded a 
few fossils, mostly Alovw,—Buthotrephis major, B. minor, B. minimus, 
Keep., Paleochorda tardifurcata, Retiofucus extensus, and Nema- 
tolites Hdwardsu, Keep. 

Thickness.—We have seen that much of the apparent thickness of 
the Aberystwyth grits is deceptive, being explained away by great 
folds. Still a great series remains, well attested even by the simple 
heights of many hills which are built up entirely of this group. 
Mynydd Bach appears to be over 1000 feet high (by aneroid); and 
I should estimate the group at certainly not less than 1000 feet of 
maximum thickness. Our detailed measurements along the Devil’s- 
Bridge road show a thickness of 1639 yards, or nearly one mile. 

The Metalliferous-slate Growp—A very large part of Cardi- 
ganshire is constructed of this series. In the special area of our 
work it forms a broad semicircular zone, some eight to ten miles 
across, sweeping around the dome of the Aberystwyth grits to form 
all the slaty country from north of Cardigan up to Machynlleth. 
At the latter place it forks into two, the main curve being continued 
out to sea by the Dovey valley, while a broad but rapidly attenu- 
ating arm runs northward to Dinas Mowddwy, at which place its 
much diminished representative is found between the Bala series 
and the Tarannon shale. 

The above area, however, is not unmixed metalliferous slate, the 
upper part of the Aberystwyth group being brought up by folds, 
along north-and-south lines, running through Ystrad Meurig, and 
the Deyil’s Bridge; and possibly some outliers of the Plynlimmon 
group may be folded in. Still the general absence of arenaceous 
beds is marked over miles of country without a single grit-bed to 
serve for road- or building-stone. ‘Tin Llidiart village uses the hard 
pale slate of Goginan for building ; and at Llanafon the numerous 
boulders serve for road-metal. 

Further south, in the line between Cardigan and Llandovery, the 
slate series appears in still greater extent, which is due to its there 
representing also the Aberystwyth group, and perhaps also the 
Plynlimmon grits, these groups not being developed as such in that 
area. 

A marked lithological character distinguishes this group as a 
whole, the terms “‘ indurated shaly slate,” ‘irregular slate,” « flagey 
slate,” ‘ pale blue slate,” from my note-book, serving to indicate the 
usual character; I also find “hard pale slate-rock, often much 


GEOLOGY OF CENTRAL WALES. 158 


jointed,” and “pale slates with ribbon bandings.” ‘The prevailing 
colour of the rocks is a pale blue-grey ; and they generally break up 
into moderately small fragments, shale-fashion. 

Slate.—In many places where the slates are better cleaved, and 
less frequently jointed, they are worked in small quarries for inferior 
local purposes, as around Llancynfelin, in the hills south of Machyn- 
lleth, in the island mounds of the Gorsfochno flats, and near the 
Devil's Bridge. These rough slates are pale-coloured, hard, and 
coarse; but around Machynlleth thoroughly good slates are worked 
in anumber of large quarries. One of the largest of these is at Pont 
Faen, where large slate-flags are extracted ; but the most interesting 
to the geologist is that at Morben, two and a half miles south-west 
of the town of Machynlleth. These are thick-hedded slate rocks, 
with a few thin grits, dipping at high angles to the W.N.W. 

This, however, is an inverted dip, a8 is proved by the position of 
the prominences on the grit-bands, which are upon the upper sur- 
faces of the grits, the worm-trails being also manifestly in inverted 
position. ‘The cleavage-dip diverges only slightly from the bedding-. 
dip, so that the Graptolites usually run for some distance along the 
surface of the slate, and then gradually skim under it. The slates 
are very good, dark and soft. Nodules of iron-pyrites occur; and 
nearly all the Graptolites are converted into this bright mineral. 
This quarry has yielded the finest of our fossils, the Monograptus 
Sedgewicki being especially magnificent. 


Fossils from Morben Quarry, Machynlleth. 


Rastrites peregrinus, Barr. Monograptus Hisingeri, Carr. 
maximus, Carr. -—— tenuis, Portl. 

Monograptus Sedgewickii, Pord/. myolutus, Lapw. 

— , var. spinigerus, Wich. Diplograpsus Hughesu, Nich. 

—. , var. distans, Pordd. Climacograpsus normalis, Lapw. ? 

—— intermedius, Carr. Myrianites M‘Leayi, Murch. ? 
spiralis, Geinitz. Lapworthii, Keep. 

——- lobiferus, M‘Coy. Buthotrephis major, Keep. 


Below this in present position, but by inversion really above it, is 
a zone of pale blue and olive shaly slate, pointed out to me by 
Mr. J. EK. Marr, F.G.S., as being similar to certain zones in the 
Coniston mudstones. The great slate district of Corris, usually 
referred to the Llandeilo age, is in this series, as will be shown 
later on; also the excellent slates of Dol y Mynach and Cwm par 
Adwys, west of Rhyader. 

Other rock-varieties, such as dark paper-shales, rab, and mud- 
stone, approach near to the Aberystwyth types; and some of the 
occurrences of such rocks doubtless indicate the uprising of that group 
as inliers in the metalliferous-slate area. In the Devil’s-Bridge 
road such rocks appear in several places. 

Grit-beds as thick as 6 inches may occur; and thin zones, about 
4 inch to 2 inches thick, are frequent; these are generally red- 
stained at the surface, and micaceous. ‘They are trustworthy guides 
to the dip of the beds, which in their absence is often not easy to 


discover. 


154. WALTER KEEPING ON THE 


At Cwm Symlog, nine miles east of Aberystwyth, pale soft shales. 
and slates with thin grits, rich in fossils, are found. 


Fossils from Cwm Symlog. 
Rastrites peregrinus, Barr. Monograptus palmeus, Barr. 
Monograptus Sedgewickii, Port. Climacograpsus spiralis, His. 
crenularis, Lapw. 


FORAMINIFERA. 
cog ee ape _Dentalina. 
eregarius, Lapw. : 
intermedius, Carr. Textularia. 
—— Clingani, Carr. Rotalia. 
spiralis, Gezn. ANNELIDA. 
lobiferus, M*Coy. Nereites Sedgwickii, Murch. 
runcinatus, Lap. Myrianites M‘leayi, Murch. 
—— Hisingeri, Carr. , Nemertites Olivantii, Murch. 
tenuis, Portl.? 
involutus, Lapw. ALG. 
Diplograpsus tamariscus, Nich. ? Buthotrephis major, Keep. 
Hughesii, Nich. minor, Keep. 
sinuatus, Nich. —— minimus, Keep. 


At Ystrad Meurig, forming the high ground twelve miles south- 
east of Aberystwyth, and in near association with a fossil zone, is a 
series of hard pale grits with thin indurated slate partings, firmly 
bound together, to form hard massive rocks (a); we refer to this as 
the “Strata Florida rock.” 

Another well-marked rock demanding attention is (6) a pale indu- 
rated slate rock, usually cross-banded by thin arenaceous stripe- 
zones. It is well seen at Machynlleth, to the east of Kglwys Fach, 
and east of Tal y bont. Again, the same type of rock occurs (c) in 
the hills east of the Rheidol gorge; I find blocks of it around Llan- 
trisant; and it is also well developed at Goginan, to the east of 
Llanddewi Brefi, and in the hills west of Rhyader. These rocks 
are essentially similar to those of Ystrad Meurig, only less arena- 
ceous; and I include them all under the name of “Strata Florida 
rock.” 

Now, classifying these various localities of the Strata Florida rock, 
we find they fall into a set of definite lines having similar relations 
to neighbouring rock-beds, and indicating, as I believe, definite 
zones. First, the more arenaceous rock of Ystrad Meurig is in the 
same north-and-south line with Cwm Symlog and with Llantrisant ; 
and in the first two of these places it is in near relation with a rich 
fossil zone. Also it is about in the same line with the Large-flag 
series, as seen in the Devil’s-Bridge road, nine miles east of Aberyst- 
wyth, also found at Llantrisant aud Cwm Symlog. In these places 
we have, then, evidence of the occurrence of this hard pale slate rock 
along a definite line of country in near association with the Grap- 
tolite fossil-zone and the Large-flag series. Next we find that the 
group b also falls into a line east of Aberystwyth; and here, again, 
it is associated with the great fossil zone at Machynlleth, while near 
Tal y bont it is connected with a band of rottenstone like that of 
the Large-flag series at the Devil’s Bridge. 

Altogether, then, the result of these lithological correlational is to 
support our theory of four different lines of north-and-south foldings 


GEOLOGY OF CENTRAL WALES. BUS, 


between Aberystwyth and Plynlimmon—namely, (1) the principal 
axis of Aberystwyth, (2) the principal axis of Plynlimmon, and (3) 
the two minor axes running (a) from Ystrad Meurig through Llan- 
trisant and Cwm Symlog, ‘and (5) along the gorge of the Rheidol 
about Pont Erwyd and the Devil’s Bridge. The further northward 
extension of (a) is probably indicated by the grits of Yr Carreg and 
Moel y Llyn. These minor folds have not been recognized further 
south ; and, indeed, we might expect them to disappear as the lateral 
spread of the beds becomes restricted. 

At Lampeter the well-known olive shales and grits with Werettes 
&c. are exposed in quarries. The grits are in thin beds (2-4 inches), 
hard and laminated, with partings of shale either light (olive) or 
dark in colour; no cleavage is seen. The fossils are :— 


Nereites Sedgwickii, Murch. Nemertites Olivantii, Murch. 
-— cambrensis, Murch. Palzochorda tardifurcata, Keep. 
Myrianites M‘Leayi, Murch. Retiofucus extensus, Keep. 


also other worm-like and algoid markings. -Altogether this is a 
pretty typical metalliferous slate, though more arenaceous than is 
common, 


Fig. 3.—Railway-cutiing between Traws Coed and Caradoc 
Waterfall, 1877. 


Contortions.—The woodcut (fig. 3), representing a railway-cutting 
east of Traws Coed, illustrates some of the frequent and sharp contor- 
tions of this series. All the perceptible foldings, however, numerous 
and striking as they are, are far from adequately showing the real 
complexity of the infinite twistings and inversions of this series, the 
existence of which is proved principally by considerations from the 
fossil evidences. Without fossil evidences it is, as a rule, impossible 
to detect inversions; but still, in those places where grits with 
fucoidal markings &c. occur, we have another key to the true rela- 
tions of the rocks—namely, in observing the positions of the curious 
prominences upon the surface of the grit, these being normally 
always upon the under surfaces; but where the beds are inverted 
(as at Morben, Machynlleth, &c.) they are found upon the upper sur- 
faces, the true worm-trails (Nereites &c. ) being in this case Sapresed 
upon the under surfaces, in inverted position. 

These inversions explain away in great degree the enormous 
apparent thickness (miles of strata) of this series; but a very great 
vertical series still remains well attested in actual steep and preci- 


156 WALTER KEEPING ON THE 


pitous mountain-sides, from 500-1000 feet high, and I cannot esti- 
mate the group at less than 2000 feet. 

Fossils are almost entirely wanting in this series, except, I be- - 
lieve, near the base, where I would place the rich beds of the 
Morben Quarry, Machynlleth, and Cwm Symlog. The fauna is 
intimately related to that of the Aberystwyth group, as indeed was 
to be expectec ; for the fossils of this latter group occur in its higher 
part, not far removed, in serial order, from the Metalliferous-slate 
fossil-zone. 

The worm-like and fucoidal markings, Nereites, the Fan Algal, 
and the Nematolites are widely distributed throughout the series, 
worm-trails being very characteristic. 

Other fossil localities are :— 

Llantrisant, with the large Fan Algal (Buthotrephis major) and 
Nematolites tubularis, Keep.; the Llyfnant valley, with Climaco- 
grapsus scalaris, His. ; Dyffryn Castell, with Climacograpsus scalaris, 
worm-tubes, and the Nematolites, also some of the shells recorded 
by Sedgwick ; and Steddfa Gurig (west of), with Monograptus Sedge- 
wickri, “Portl., M. tenuis, Port. ?, Climacograpsus scalaris, His., and 
Orthoceras. ‘Above Taliesin, frasments of dendroid Graptolites and 
Nemutolites tubularis occur ; sm the latter is also found at Llantri- 
sant, Ty Llwydd, on the Rheidol, and elsewhere. 

Richer localities are :— 


West of Lisburne Mine, Ystwyth Valley. 
Climacograpsus scalaris, His. 


Nereites Sedgwickii, Murch. 
Buthotrephis, 


Rastrites ? 
Monograptus Sedgewickii, Pordl. 
spiralis, Gein. 


-—— lobiferus, MM‘ Coy, 


Garthen Valley, Melin Newydd. 


Monograptus Sedgewickii, Por?/, Orthoceras, sp. 
Climacograpsus scalaris, Hs. 


The numerous metalliferous veins (lead and silver) in the rich 
mining-districts of Cardiganshire are almost confined to this set of 
beds; hence the name here given it. 

3. The Plynlimmon Group.—As we ascend the mountain-slopes 
along the valley of the Rheidol, above Pont Erwyd, we meet with 
the rocks of the upper grit series, which form the upper part of the 
Plynlimmon mountain. Around Garn Fach some of the grit-beds 
are seen, of rather coarse type, sometimes assuming a very grani- 
toid aspect, there being a profusion of felspar crystals and large 
blebby quartz grains. The felspar crystals may be of large size; and 
some of them are perfectly preserved and glassy. Doubtless the 
rock is the direct result of the degradation of an ancient acidic 
igneous rock. 

Conglomerate beds, consisting of pebbles of white quartzite and 
vein quartz in a grit matrix, are also met with; but they are few. 
The group here is still, in the main, a slate series, some of the slate 
being pale and papery, of the “ pale-slate type.’ Iron pyrites is 


GEOLOGY OF CENTRAL WALES. 157 


common in both grits and slates. The main dip is decidedly 
easterly. 

Mounting the higher ground above Nant y Moch the grits become 
more conspicuous, forming a terraced structure by their outcrops 
around the hill; and near the summit a rugged steplike structure is 
produced by the regularly bedded fine-grained grit series. The 
finest exposure of these Plynlimmon grits is in the magnificent pre- 
cipices fronting the Rheidol lake (Llyn Llygad Rheidol), where 
many of the beds are very thick (7-10 feet) and of moderately 
coarse grain. A particular feature in them is their regular columnar 
jointing; this is quite as regular as in many columnar felsites, 
numerous columns, 7 feet long, being seen running right across the 
beds. Conglomerates are uncommon; but some three or four beds 
of beautiful “pudding stone,” with white quartz pebbles in a fel- 
spathic sandstone matrix, are seen upon the western slope of the 
mountain. 

Coarser beds of this group are probably indicated by a large 
boulder upon the hill-top above Gogerddan House, near Aberystwyth, 
which has pebbles as big as cannon-balls. 

These grits form a crescent zone from north of Plynlimmon 
through the high ground of Mid Wales, running east of Tregaron 
and east of Lampeter; but they die out beyond this to the south. 
They constitute the genuine backbone of South Wales, lying in a 
synclinal. The highest beds form the upper ground of Plynlimmon 
itself. In the Teifi area the beds are remarkable for being coarsely 
eleayed into large tiles about 4 inch to 1 inch thick, a structure 
which graduates into jointing on the one hand and true slaty 
cleavage on the other. Similar cleaved grits occur around Llyn 
Fyrddyn and on the hills east of Lampeter, all of which I would 
place on the same horizon. There is, however, some doubt in my 
mind as to the exact position of this horizon in the geological 
series. 

Foldings.—As a group, the Plynlimmon grits are characterized, 
not by violent contortions or sharp bendings with fracture, but by 
broad and gentle foldings. These are beautifully seen around the 
Teifi lakelets and Llynodd Ieuan; also at Llyn Fyrddyn Fach and 
Llyn Fyrddyn Fawr, and Llyn Bugeilyn, where the ridges and minor | 
hog-backs of the surface correspond with the structural anticlinal 
and periclinal foldings of the rocks with great regularity and 
clearness. 

Amongst the rock-varieties we may notice blue shaly slates, blue 
slates in 6-inch beds, cleaved into oblong flags about 3 inch thick 
(Llynodd Ieuan), and arenaceous mudstone, broken up into coarse 
rab, the fragments measuring 1-3 feetx3 inchesx1 inch, also 
smaller rab. 

The lithological differences between this series and the Aberyst- 
wyth group are the greater thickness of the grit beds, the presence 
of quartz conglomerates, and the rarity of curious rock-surfaces and 
fucoidal markings. The slates also are, as a whole, of paler type. 

Fossils have not been discovered in the grit series of the Plyn- 


158 WALTER KEEPING ON THE 


limmon mountain; but large boulders of coarse conglomerate upon 
Gogerddan Hill, Aberystwyth, which surely came originally from 
this mountain, contain casts of various fossils, of which I can men- 
tion crinoid ossicles, Nebulipora?, Petraia, a large cup coral, and 
Meristella. 

The thickness of this group must also be very great; for Plyn- 
limmon rises some 1500 feet above Garn Fach, the level where the 
Plynlimmon grits first appear. A thousand feet is probably an 
under-estimate. 


Part Il. Brief Notes upon other Sections. 


(1) From Aberystwyth through Pont Erwyd to near Builth (fig. 4),— 
From Aberstwyth to Pont Erwyd the section is similar to that from 
Aberystwyth to the Devil’s Bridge; but the western dips are more 
important towards Pont Erwyd, so that the inversions required are 
here smaller or less numerous. Beyond Pont Erwyd is a great slate 
and shale series running for many miles, nearly to Rhyader, and 
forming by the apparent dips an important synclinal under the highest 
ground south of Plynlimmon. Fossils occur at Pont Erwyd and 
Dyffryn Castell, also near Rhyader. The hard pale-slate series comes 
in at Goginan and east of Pont Erwyd, and there is a similar rock 
west of Rhyader; but the Plynlimmon grits do not actually appear 
as such, unless the grits of Llynodd Ieuan belong here; they are, 
however, well developed in the mass of Plynlimmon further north. 
Some of the paler slate along our line of section may represent this 
series. Grit beds similar to those of the Aberystwyth group are 
found in two places near the ‘Glansevern Arms,’ and again further 
on east of Langwrig. ‘They are to be regarded as special local de- 
velopments inthe Metalliferous Slates. Other grits and conglomerates 
seen at Gwastaden are probably the representatives of the Plyn- 
lmmon grits. 

(2) Section from Llandeilo to Aberaeron.—Three principal grit- 
areas are met with in the line of this section—namely, at Talieris 
(west of Landeilo), east of Lampeter, and at Aberaeron. The Talieris 
conglomerate beds are surmounted by dark shales and shaly slates 
of the Metalliferous types ; and beds of the Rhyader pale-slate type 
next underlie the grits of the country east of Lampeter. By an 
exceptional appearance the Metalliferous slates of the west seem to 
underlie the Aberaeron grits. No pale slates are seen beneath the ~ 
Talieris conglomerate beds. The principal axial fold in the section 
seems to be a synclinal in the Teifi valley by Lampeter—the 
mountains to the east and west of this being (that is, if the rock- 
dips have any truth in them) great anticlinals in the Metalliferous- 
slate and Rhyader Pale-slate series. The beds of grit and con- 
glomerate at Talieris are not of the nature of basement beds of a 
stratigraphical group, but indicate nothing more than such slight 
physical variations as a shallowing of the sea-bed or a change in the 
direction of the currents, resulting in the formation of sand and 
pebble banks ; for the pebbles are not fragments of the underlying 
rocks, and there is no trace of any physical break or even change of 
lithological character above and below them. 


159 


GEOLOGY OF CENTRAL WALES, 


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160 WALTER KEEPING ON THE 


(3) Rhyader to the Teift Pools 
(fig. 5)—In this section three 
main folds are indicated, namely a 
central anticlinal fold at Disgwylfa 
in the Rhyader pale slates, with a 
synclinal on its east and west. 
The former is a comparatively 
simple basin of pale slates beneath 
the grits and conglomerates of 
Cwm Elan and Gwastaden; the 
latter, corresponding with the 
Plynlimmon synclinal, is a much 
more complicated structure, pro- 
bably much affected by reversals. 

General Result of the Hast and 
West Traverses (see fig. 6).—The 
sections just indicated show that 
the great area of Mid Wales is 
made up of a thick series of im- 
perfect slates, pale slates, shales 
and grits, having a general re-. 
semblance and intimate connexion 
throughout, as of one continuous 
group, but divisible into the sub- 
groups indicated in the beginning 
of this paper. 

These rocks form one great pri- 
mary synclinal extending from the 
Aberystwyth axis to the east of 
Rhyader, and from the west of 
Llandeilo to Aberaeron. For in 
the west of Cardiganshire we find 
a great and continuous course of 
easterly dips running inwards from 
the coast; and in the eastern half 
of the area, in Radnorshire and 
Caermarthenshire, is a correspond- 
ing grand set of westerly dips. 
Subordinate but still very great 
anticlinal folds along north and 
south axes bring up on the east 
tee Metalliferous slates between 
Plynlimmon and Rhyader, and on 
the west the upper part of the 
Aberystwyth grits, in the line of 
Pont Erwyd and the Devil’s 
Bridge. Another, smaller fold 
with the same result as the last, 
runs from Ystrad Meurig north- 
wards towards Machynlleth, thus 


Rhyader. 


3. Rhyader pale slates. 
f. Plynlimmon grits 


Plynlimmon. 


rous-slate group. 


Fig. 6.—Diagram of the General Structure of Central Wales. 
yth grits. 


1. Aberystw 
Metallife 


2. 


Aberystwyth. 
{ 


GEOLOGY OF CENTRAL WALES, 161 


greatly swelling out the surface-area of the Metalliferous-slate 
series. 

~ Minor foldings, also with north-and-south axes are infinitely 
abundant over the whole district, with frequent inversions which 
obscure the original order of the rock-beds. 

Included in these foldings are four principal crescent zones of 
grit, greywacke, and conglomerate, namely :—1Ist, the western or 
Aberystwyth grits; 2nd, the eastern grits—an imperfect line of 
grits seen at Talieris, west of Llandeilo, and perhaps continued in 
other grits Kast of Gwastaden. These I regard as the diminutive 
representatives of the Aberystwyth grits. 3rd. The Plynlimmon 
grits and conglomerates, with much associated pale slate; and 
Ath, the grits and conglomerates of Cwm Elan and Gwastaden, which 
are also associated with, and, indeed, included in, a great pale-slate 
group, the Rhyader pale slates. The last two are upon the same 
stratigraphical parallel, occupying synclinals. These grits I do not 
regard as necessarily holding exactly the same horizon over wide 
areas, they having been probably shingle banks over the old 
sea-bottom. The pale slates seen in so many sections appear 
to belong to two principal horizons—(a) the lower or Strata Florida 
pale slates in the lower part of the Metalliferous-slate group, and 
(6) the Rhyader pale slates, associated with the Cwm Elen, Gwastaden, 
and Plynlimmon grits. 


More distant Sections (see Table of Vertical Sections, p. 164). 


In order to work out the relations of these rocks to the other 
lower paleeozoic groups, I have visited various districts where the 
Cardiganshire series might be studied in association with rocks of 
some definite and decided horizon—namely, amongst other places, 
Llanbrynmaer, Dinas Mowddwy, Cardigan, N. Wales, &c. 

In the Llanbrynmaer area we find (1) an upper series of grey 
grit and greywacke, often very felspathic and micaceous, cleaved, 
and with fragments of fossils (the Denbighshire grits). These pass 
down by gradual transition into (2) a series of pale blue and green, 
grey and purple shales, with some darker shale, rab, and green grits, 
which make up the true Tarannon shales; some obscure fragments 
of Graptolites have been found here. Again, there is no break, but 
a simple passage between this series and (3) the lower series of 
grits, greywacke, and dark shales which belong to the Cardiganshire 
type, and contain the fossils of our Metalliferous group. To the 
west of the Tarannon plateau we find the Metalliferous slates fully 
developed in the Pennant valley, with the characteristic fossil species, 
and the usual associated lead-mines of that group. 

The whole character of this Metalliferous slate is different from 
that of the Tarannon shale; and our evidence points uniformly and 
decisively to the Metalliferous slate being a more ancient series, the 
two groups being separated in this area by a zone of grits. The 
Plynlimmon grits and associated pale slates, as seen in Plynlimmon 
and around Rhyader, appear to be wanting in this area, there being 
no well-developed grit series between the Llandovery group and the 


162 WALTER KEEPING ON THE 


Denbighshire grits. We shall find reason to believe that they are. 
represented, feebly it is true, by the thin grit-beds above described 
in the Tarannon shale. 

The Dovey Valley : Section across the valley near Llany Mowddwy 
(fig. 7)—This section gives, in a narrow area (measuring less 
than 2 miles across), the complete series from the Lower Bala slates 
tothe Denbighshire grits. The Denbighshire grits pass down into 
the Tarannon shales, which are pale-slate rocks resembling the 
Rhyader pale slates. There is, again, no break between these and 
the series of greywackes and dark shales and slates beneath. ‘This 
latter group (which is in part the Lower Llandovery of the Survey) 
agrees both in its lithological characters and in its fossils, so far as 
known, with the Aberystwyth grits. 

There is no distinct basement bed of conglomerate or grit to this 
series, such as might indicate the existence of an important strati- 
graphical break ; but, on the contrary, there is every appearance of 
a passage from the underlying great slate group up into the true 
Silurian series. This great slate group corresponds in its general 
lithoisgical characters and its “ cone-in-cone” nodules with our 
Metalliferous slates; but I have not detected any pale slate in them. 
In the absence of fossils, it is not possible to determine their exact 
age; but they should probably be regarded as belonging for the most 
part to the Upper Bala slate group. I should, however, expect 
some of its upper beds to be the representatives of our Cardiganshire 
Group, the lower part (Upper Bala) corresponding with much of the 
slate group between Cardigan and Llangrannog, about to be de- 
scribed. 

It is further noteworthy that im this line of section, as in the 
Cardiganshire Group, the newer rocks are much better cleaved, and 
appear altogether more highly metamorphosed than the more ancient 
rocks to the west. 

Corris Avca.—The great slate district of Corris, usually referred 
to the Llandeilo age, belongs to our Metalliferous-slate group, as is 
proved by the following fossils from Corris and Taren y Gesail :— 


Monograptus Sedgewickii, Portl.? Climacograpsus scalaris, His, 
—- tenuis, Portl.? Orthoceras, sp. (same as the Cefn- 
Hendre shell). 


The Cardigan District.—The pale felspathic grits and black slates 
of Newport Bay and Cardigan, hitherto placed in the Lower Llan- 
dovery series “‘b 4,” are not of this age, but belong to the Middle Bala 
or Caradoc Group ; as to this the fossil evidence is conclusive. Above 
these come rolling beds of pale slates and shales, then darker shaly 
slates, with a zone of pale felspathic grits; these also I refer to 
the Bala period. The overlying shaly slates and rab are passage- 
beds of Caradoc-Llandovery age, presenting the gradual incoming 
of the Aberystwyth grits. 

Reviewing the several rock-groups and their distribution, we find:— 

(A) The Plynlimmon grits, seen upon Plynlimmon and around 
Rhyader, also probably in the hills west of Tregaron and Lampeter ; 


163 


GEOLOGY OF CENTRAL WALES. 


Dark slates (Lower Bala). 
Bala ash-bed. 

Bala limestone. - 

Bala phosphate-bed. 


Calcareous slates and flagey 


shales. 


ig. 7.—Section through the Dovey Valley at Llan y Mowddwy. 


(Horizontal scale, about 23 inches to the mile.) 


Lian y Mowddwy. Cwm Maenydd. Waterfall. 


6. Imperfect slates, dark shaly and 9. Hard pale slates, banded slates, well 
rubbly slates and shales. cleaved (‘Tarannon group). 

7. Dark-coloured irregular slates, with 10. Dark rab and coarse gritty rab, and 
“‘eone-in-cone” nodules. grit bands (Passage-beds). 

8. Grits, dark shaly slates, and rab. 11. Denbighshire grits, with dark rab 


Fossils (Lower Llandovery). and slates. 


164 | WALTER KEEPING ON THE 


but they die out to the north and south of these localities. No 
fossils have yet been detected in this series in Cardiganshire, except, 
perhaps, in the boulder-block at Gogerddan, near Aberystwyth. 
Stratigraphically they lie above the Metalliferous slates; and these 
latter are proved, in the Llanbrynmaer, Dovey, and other areas, to 
be inferior to the Tarannon shales. Therefore, with reference to the 
Noedd Grug section, they must either represent the uppermost part 
of the Llandovery rocks or a special gritty development in the Taran- 
non shales. I adopt the latter view for the following reasons :— 
because of (1) their dissimilarity to the grits (Llandovery grits) 
which underlie the Tarannon shales at Llanbrynmaer, Llan y 
Mowddwy, and elsewhere ; (2) their association with pale slate rocks 
similar to the Tarannon shales of the Corwen area; and (3) the oc- 
currence of such a group of rocks, the Gala grits*, in this position 
in the south of Scotland. Moreover, in the Tarannon district the 
first appearance of such a development of grits in the Tarannon 
shale is already seen in the numerous thin grits, with contorted 
structure, of Tarannon Hill, Llanbrynmaer. The Rhyader pale 
slates thus appear also to belong to the true pale slate or Tarannon- 
shale series. 

(B) The Metalliferous slates, which are enormously developed, 
spreading over a very wide area of Mid Wales. ‘They maintain one 
general lithological character of hard, pale, shaly slates, also containing 
a zone of pale slates. The group is generally plumbiferous. 

(C) The Aberystwyth grits may be taken as an arenaceous de- 
velopment of the Metalliferous slates, in its lower part. Like the 
Plynlimmon grits they die out to the north and south, their southerly 
attenuation being well exhibited around Llangrannog. They dip 
persistently under the Metalliferous slates ; and the truth of this dip 
is proved by the position of the contorted raised structures upon the 
under surfaces of the gritst. 

In the Dovey valley the Aberystwyth Grits are represented by 
eritty beds of a similar character; but here they appear to belong 
to a slightly different horizon, namely to the upper part of the 
Metalliferous slates; and the same is true of the grits exposed in 
the deep Talerddig cutting, Llanbrynmaer. The grits and conglome- 
rates of Talieris, west of Llandeilo, very probably belong to the 
horizon of the Aberystwyth beds. In the Noedd-Grug section these 
beds, together with the Metalliferous slate, are represented by the 
Llandovery group of the Geological Survey, probably the Upper and 
part of the Lower Llandovery ; but from the fossil evidence we can- 
not recognize the lowest part of the Llandovery Group (Lower Birk- 
hill) in Central or West Cardiganshire. Part of the lower set of 
slates beneath the grits, described in the Cardigan and Dovey areas, 


* Mr. Lapworth, F.G.S., writes me that “all the paleontological affinities 
of the Gala beds are with the Tarannon shales, with which, and not with the 
Upper Birkhill beds, they must eventually be connected.” 

+ Ihave confirmed this test by examining the beds of Lingula-flags, Trema- 
doc and Bala rocks of South Wales, where these curious worm-like, fucoidal, 
and irregular prominences are found uniformly upon the wader surfaces of the 


grits. 


les and othe 


6. Abery, 
and Plyn 


Pale |. 
Shales, | ~~ 


Mudstone 
Group. 


Bala 
Group. || 


a 


ae 


a we 


wit. 


ore 


Fr 


2 South Scotland 
(Lspworth). 


3. South-west 
Cardiganshire. 


Comparative Vertical Sections of Silurian Rocks in Central Wales and other Localities. 


5. West of 
Llandeilo, 


6. Aborystwyth 


4, Rhyader, 


Plynlimmon | 
Grits. | 
Grits. 
alee peers 


Pale Slates. 


Metalliferous 
Slates. 


MUM 
WOME 
CMM 


TT) 


WMUMLLA 


Aberystwyth [= 
Gnits. 


ee 
MM PTL 


LYM 
| HMMM 


WW 


and Plynlimmon. 


7. Lianbrynmaer, 


Denbighshire 
Grits. 


Tarannon 


Shales. (AF 


LZ 
V7 


8, Llan y Mowddwy, 


| 
ie 


Bala - 
Limestone. 


[Vo fuce puge 164, 


9, Clwydd Valley 
(Prof, Hughes). 


Corwen === 
Grits. | 


GEOLOGY OF CENTRAL WALES. 165 


probably belong here and complete the series ; for we have seen there 
is no break of continuity down into the Bala formation. 


Part III. Paleontological Evidence. 


Although the order of succession of the forms of life have in all 
eases been first determined from the results of stratigraphical re- 
search, yet, that order once established, the organic remains, as 
considered in groups, become the supreme test of the age and rela- 
tive order of rock-masses—superseding, and in some cases actually 
overthrowing, the apparent evidence of succession in the rock-beds. 
Such we find to be the case over a large area of Cardiganshire, 
where, as in the section from Aberystwyth to the Devil’s Bridge, the 
stratigraphical series, seeming to be continuously ascending through 
some three or four miles of thickness to higher and higher beds, 
is proved by a handful of Graptolites to be radically misleading ; 
for, the fossils being identical at almost the extreme ends of the 
section, the true reading is shown to be a series of inversions, by 
which the original order of the beds is obscured. Thus, over a large 
part of Mid Wales, it is to the organic remains alone that we can 
look with confidence for unravelling the apparently inextricable, and 
often delusive, tanglement of the strata, and establishing their true 
succession. 

The most important element in our paleontological evidence is 
the group of Graptolites, these fossils having now become, through 
the most successful work of Mr. C. Lapworth, F.G.S., of the highest 
value in Cambrian and Silurian geology. All my work in this 
branch of study is based upon the published results of that geologist : 
and I am also particularly indebted to him for examining many of 
my Graptolites, as indicated in the Table (p. 170), and for valu- 
able notes upon them conveyed to me in letters, from which I have 
quoted below. 

A study of the Table brings out a number of important facts. 
At the first glance it is clear that all our fossil lists exhibit 
one and the same general geological fauna. Our richest localities 
are Cwm Symlog (eight or nine miles east of Aberystwyth) and 
Morben (near Machynlleth), also Cefn Hendre, Cwm, and Bryn y 
Carnauin the neighbourhood of Aberystwyth). A single one of these 
localities (Cwm Symlog) yields every species, except four, of the 
true Graptolites known in Cardiganshire ; and this, together with 
the intimately allied fauna of Morben Quarry, includes all our spe- 
cies except one (Monograptus turriculatus). 

The other forms of life show no less clearly the unity of our 
fossiliferous Cardiganshire rocks. The Orthoceras is wide-spread 
in the Metalliferous Slates, and occurs also amidst the Greywacke- 
Flags of Cefn Hendre; and the various forms of Alge and worm- 
trails are also characteristic of the whole range of the Cardiganshire 
group. 

For the typical area of reference for Graptolitic deposits we must 
unquestionably look to the rich beds in the south of Scotland, of 
Llandeilo to Wenlock age, which have been so thoroughly worked 


Q. J.G.S. No. 146. N 


166 WALTER KEEPING ON THE 


out by Mr. Lapworth. And here the reading is decisive: our Cardi- 
ganshire series belongs not to the Glenkiln (Llandeilo) or the Hartfell 
(Bala) periods, when the Didymograpti and Diplograpti flourished, but 
to that part of the newer or Silurian era when the Monograptide 
were predominant, and in their greatest development, namely the 
Llandovery period (Birkhill Shales). 

The more detailed affinities of our fauna will appear by a study 
of the last five columns of the ‘‘ Table of Distribution.” Every one 
of our species of true Graptolitide (excluding the Cladophora) occurs 
in the Upper Birkhill beds of Scotland. Eight species are common 
to the Lower Birkhill Shales, and nine (as also two species of Cla- 
dophora) to the overlying Gala and Tarannon groups. 

Of the fossils of the two principal Graptolitic localities, Cwm 
Symlog and Morben, Mr. Lapworth writes that they ‘‘lie within the 
same general Graptolitic zone.” .. . “‘ Of the nine species recognized 
in the slates of Morben, five (namely Rastrites maximus, Carr., 
Monograpius spinigerus, Nich., MW. distans, Portl., M. Hisingere, 
Carr., and Diplograpsus Hughesiz, Nich.) make their first appearance 
in the Upper Birkhill Shales of 8. Scotland. astrites maximus is 
confined to the highest zone of that formation, in the Moffat area, 
and, together with its common associate, Dipl. Hughesii, seems to 
have become extinct before the deposition of the overlying beds of 
the Gala group. 

“¢ The second fossil locality (Cwm Symlog) is even more strikingly 
marked by its Upper Birkhill Graptolites. Sixteen forms have been 
identified from this spot. Of these, seven species (viz. Monograptus 
crenularis, Lapw., MV. pir Ge Carr., M. Clingani, Carr., MW. 
runcinatus, Lapw., M@. Hisingeri, Carr., Dipl. Hughes, Nich., and 
D. sinuatus, Nich.) are forms which are known for the first time in 
the Upper Birkhill of S. Scotland, and its equivalents in Girvan, 
Treland, and the north of England. J. intermedius, M. runcinatus, 
and M. crenularis are peculiar to the Upper Birkhill, as also is M1. 
Clingani, which is confined to a small seam in the very centre of the 
Birkhill beds.” 

Of the nine species of true Graptolites common to our Mid-Wales 
rocks and the Gala and Tarannon, not one is a special Tarannon 
form. Monograptus turriculatus, however, is most frequent in those 
beds; and the two species of Cladophora Dictyonema venustum, 
Lapw., and Rhizograptus ramosus, Lapw., are only known, else- 
where, in the Gala group (Scotland). 

From the fossil evidence, therefore, there can be no hesitation in 
referring our Mid-Wales rocks to the same age as the Upper Birk- 
hill of Scotland; and beyond the occurrence of the three species 
just mentioned in the rocks of the Devil’s Bridge, there is nothing 
that conveys the slightest hint that any of our strata are newer than 
Upper Birkhill. 

Lake District.—Fifteen of the Mid-Wales Graptolites are known 
from the Coniston or Graptolitie Mudstones of the Lake District; so 
that, in Mr. Lapworth’s ,words, ‘‘ the general facies of this Mid- 
Wales fauna is distinctly that of the Coniston Mudstones.” These 


GEOLOGY OF CENTRAL WALES. 167 


species are nearly equally distributed in the two zones (tenwis- and 
argenteus-zones) of that area. Also, amongst the other fossils, Mr. 
Marr recognizes the fragment (tail) of a Phacops, found by him at 
the Devil’s Bridge, as a new species also occurring in the “ Mud- 
stones” of the Lake District. 

Lithologically we find some of the Graptolite-bearing beds not 
unlike part of the Coniston Mudstone series; and some paler 
shales and slates, associated with the Graptolite beds at Morben, 
Machynlleth, and occurring again at Cwm Symlog, have their repre- 
sentatives also in the same series. Still, regarding them as a whole, 
our Cardiganshire rocks decidedly do not resemble their representa- 
tives in the English lake country. 

We have seen that our verious fossil lists indicate that all the 
fossiliferous beds belong to one general horizon. Certain minor 
differences, however, are to be observed, of which the most important 
are the presence of Monograptus turriculatus at Cefn Hendre and the 
Devil’s Bridge, and the species of Cladograpt: of the latter place, 
which are common to the Gala group of Scotland. 

From the Graptolitic evidence Mr. Lapworth would arrange the 
beds in the following order :— 


D. The M. turriculatus beds of the Devil’s Bridge. 

C. Next below (or perhaps identical with (D)) the Cefn Hendre 
erits. 

B. Then comes the rich Graptolitic zone of Morben and Cwm 
Symlog. 

A. Lowest of all lie jointly the Diplograpsus beds of Taren y 
Gesail, Corris, and Steddfa Gurig. 


It will, however, be seen that this order does not correspond with 
our well-established succession of the greater groups as given in 
the first pages of this :paper; for it would make our Metalliferous 
slates newer than the Aberystwyth grits. Now, with all my general 
confidence in fossil evidence, | consider that the paleontological data 
are here insufficient to maintain this order against the very strong 
stratigraphical evidence to the contrary., The principal fact is the 
occurrence of a single species (Monograptus turriculatus) in certain 
localities ; and this does occur, only less commonly, in the Birkhill © 
shales as well as the Gala group. Moreover the similar (more 
arenaceous) physical conditions common to our Devil’s-Bridge bed 
and the Gala group may have to do with their having some fossils 
im common. 

Mr. Lapworth’s researches have established the fact that, in the 
Sulurian and Cambrian periods, zones of Graptolites characterize defi- 
nite stages, just as do the Ammonites in the Jurassic rocks. But these 
latter have their exceptions, and the minor zones are only trust- 
worthy within local limits; much more so should we expect this 
to be the case with the more lowly organized group of Graptolitide. 
The occurrence of Monogr. colonus with Birkhill species in N. Wales 
is, indeed, an example of such. On the other hand the stratigra- 
phical succession of the Metalliferous slates over the Aberystwyth 

nN 2 


168 WALTER KEEPING ON THE 


erits is clearly marked over a large area, extending from near Borth, 
through Aberystwyth, and southwards to Llangrannog*, a gradual 
transition between the two groups being everywhereseen. And that 
the rocks here occupy their original relative positions is proved by 
the position of the raised rock-markings, which are uniformly found 
upon the wnder surfaces, as is normal in the lower paleozoic rocks, 
whereas when the beds are inverted these structures are found 
upon the upper surface. 

The only plan of exact harmonization would be, it seems to me, 
to suppose that the Aberystwyth grits had died out to the east 
before we reach the fossiliferous beds of Morben and Cwm Symlog, — 
and older beds had been somehow brought up in the confusion of 
folds and synclinals. This may be considered possible; but it 
again increases our difficulties in the eastern area (Devil’s Bridge 
&c.), where the usual order of the strata persists. I therefore 
adhere to my original classification as given in the earlier pages of 
this work; for, indeed, the paleontological difficulty is very insig- 
nificant. 

Comparing the faunas of the Aberystwyth grits and Metalli- 
ferous slates, we find that these do not afford any important 
data for their separation, but rather demonstrate their intimate 
connexion. ‘Twelve of our species of Graptolites are known only 
from the Metalliferous slates—namely Rastrites maximus, R. pere- 
grinus, Monograptus spinigerus, cyphus, gregarius, intermedius, 
runcinatus, and mvolutus, and all the Diplograpt:: but this signi- 
fies probably little more than the less favourable conditions for 
Graptolitic life in the Aberystwyth grits; for the two rock-groups, 
are bound together by the common possession of Monograptus dis- 
tans, Clingam, Sedgewickir, crenularis, spiralis, lobiferus, Hisingeri, 
and tenuis, and Climacograpsus scalaris. The Orthoceras, Nereites, 
Fan-Alge (Buthotrephis), and Nematolites(?) tubularis are also 
common to the two groups, and serve further to demonstrate the 
paleontological unity of the Aberystwyth and Metalliferous series. 

But there is another group (the Plynlimmon group) whose exact 
age has not yet been fully considered. ‘These beds contain no fossil 
evidence in themselves; but still, from their relations to the under- 
lying metalliferous slates and to the more eastern sections, there is 
little room for hesitation as to their stratigraphical position. 

The underlying rocks are shown to belong to the highest part of the 
Lower Llandovery group, or Upper Birkhill series, some of the species 
being actually of Gala types. Paralleling, then, our rocks with 
those of the south of Scotland, this overlying group of grits must 
either be close upon or actually in the Gala and Tarannon group. 
We have already seen how the stratigraphical and lithological con- 
siderations support the view that they really are an arenaceous 
development of the lower part of the Tarannon shale, thus being 
the true representatives of the Gala group of Scotland. The asso- 
ciated pale slates of Plynlimmon and around Rhyader (Rhyader pale 


* There is a difficulty, as before mentioned, at Aberaeron, which I have not 
been able thoroughly to investigate. : 


GEOLOGY OF CENTRAL WALKS. 169 


slates) thus fall into line with the great pale-slate group of Britain 
—the Tarannon group. 

N.E. Wales.—The Silurian rocks of Corwen, Llangollen, and the 
Vale of Clwydd have been described by Prof. T. M‘K. Hughes * ; 
and fossils have been collected from the Tarannon shales of the 
Conway area by Mr. C. Lapworth, F.G.S. Those species which are 
common to our Cardigan rocks are indicated in the Table; but, 
besides these, other species, characteristic of newer groups, are 
found—namely fetrolites, and Monograptus priodon, colonus, and 
galaensis. From these fossils Mr. Lapworth concludes that the 
Conway beds “ correspond with the earlier portion of the Gala beds” 
of Scotland. Thus the results of paleontology correspond well 
with the stratigraphical evidences, which show, in my opinion, that 
the great mass of our Cardiganshire series is unrepresented in the 
N.E. of Wales (as in the English border counties), or rather that 
it is represented by the break which Professor Hughes has worked 
out in that district at the base of his Corwen Grit. The black 
bands in the Clwydd valley, from which Prof. Hughes obtained 
Graptolites, do appear to lie in the parallel of the higher part of 
our Metalliferous group. 

My friend Mr. J. HK. Marr has just published the details of the 
sections at Cerrig y Druidion, N. of Balat. He there finds beneath 
the Tarannon shales the equivalent of the Graptolitic mudstones, 
containing fossils of the same species as our Cardiganshire series, 
but also including Monograptus colonus, which is characteristic 
of much higher beds. ‘The section appears to show a greater deve- 
lopment of Graptolitic “‘ mudstones ” than in the Clwydd valley, and 
so far to represent more fully our Cardiganshire group; but here, 
again, as at Corwen, the series appears to be incomplete, because 
of the existence of the Silurian unconformity. 


General Summary. 


Central and West-central Wales is made up almost entirely of a 
great series of imperfect slates and Greywackes belonging to our 
Cardiganshire group, together with the overlying pale slates and 
grits of Rhyader and Plynlimmon. ‘The Cardiganshire group is sub- 
divided into the (1) Aberystwyth Grits, and (2) Metalliferous Slates ; 
and part of the underlying slates may, perhaps, hereafter be proved 
to belong to the same group. Some minor subdivisions are also 
distinguishable. The arenaceous rocks are not constant over large 
areas, but die out both to north and south. 

The rock-beds are astonishingly folded into violent contortions, 
with frequent inversions, especially in the Metalliferous series, so 
as often to produce the misleading appearance of a regular and 
continuous ascending series exceeding five miles in thickness, All 
the important axes of elevation in the country havea common N. and 
S. direction, two of the main folds being the Aberystwyth anticlinal 


* Quart. Journ. Geol. Soc. vol. xxxiii. p. 207, vol. xxxv. p. 694. 
t Ann. Nat. Hist. 1880, vol. ix. p. 49. 
$ Quart. Journ. Geol. Soe. vol. xxxvi. p. 277. 


170 ON THE GEOLOGY OF CENTRAL WALES. 


and the Plynlimmon synclinal. Secondary axes of upheaval bring 
up the lower beds of the series at the Devil’s Bridge and near Cwm 
Symlog. Innumerable minor foldings preserve the same north-and- 
south strike. 

The included fossil remains, especially the Graptolites, prove the — 
Aberystwyth Grits and Metalliferous Slates to belong to the same 
general geological horizon—namely, on the parallel of the upper 
Birkhill series of 8. Scotland and the Coniston “‘ Mudstones” of the 
English Lake district. The Plynlimmon Grits are probably an 
arenaceous development of the Tarannon Shales; and the Cwm-Hlan 
conglomerates and Rhyader Pale Slates belong to the same series. 

Following up our Cardiganshire series to the lines of junction 
with older and newer groups in parts of Montgomeryshire, South 
Cardiganshire, Caermarthenshire, and Merioneth, we find no evidence 
of a break in any part of the rock-groups; but there 1s concordant 
evidence of lithological passage from the Bala to the Llandovery 
groups, and from these upwards into the Tarannon Shales and the 
Denbighshire Grit series. On the contrary, in Denbighshire and 
N.E. Merioneth, Professor Hughes has shown that, just as in the 
English-Welsh border districts, there is an important stratigraphical 
break at about this Llandovery period. Now the beds below that 
break in N. Wales are the Bala group, i. e. inferior to our Cardigan- 
shire series; those above it are basement-grits and conglomerates 
(the Corwen Grit), covered by the pale slates, with some black 
Graptolitic bands. These latter contain some species In common 
with the Cardiganshire group, but also others characteristic of a 
higher horizon than our fossiliferous beds. 

Thus all the facts are harmonious in pointing to a continuity of 
the Silurian and Cambrian deposits in West and Central Wales, 
while an important break exists in the east and north-east. Our 
Cardiganshire group is only partially developed (its upper part) in 
the latter district, being, in fact, there represented by a great strati- 
graphical break—Sedgwick’s original May-Hill unconformity. 

I conclude that, while in the latest Cambrian times(Sedgw.) and the 
dawn of the Silurian era the elevatory forces, acting in a north-and- 
south direction, lifted up the sea-bed to form a land-surface over the 
west of England and the Welsh borders, these forces influenced the 
greater part of Wales only in a less degree—producing, it may be, 
the shallower water in which the Aberystwyth grits were laid down, 
but not interfering with the continuous deposition of sediment and 
the unbroken sequence of the geological record from the Cambrian 
to the Silurian eras. 


te ceee 


. 
: 
N 


Lance showing Distribution of Fossils in Central Wales. [fo fitce page 170. 


Specimens marked thus (+) have been identified by Mr. O. Lapworth, F.G.S, 


ee 
: Scotland. Goniel 
| 4 $ 7 E 3 5 : o a 2 ack , es = = loniston 
a =) =) . = BS |e sc bb gb gE 2 = F Mudstones. | = 
SS SE eS Me Sle | ES d S/ = |da/ 2/3 |ae| 2 | 2 ae 
= (=) y s = i el =| BD 5 iS) is} & Er Gi |f dg a ] fs 
= = f is} e |Ce § 5 a: S 2 [2] A 5 3 6 
Sst | el is] Se ee ep eta 2 Bs SRS elle eee a a. WE Ale 
SS eee ee hee 2 (ee |e ca |e E/E) e] 2 lee} | & [te] 6 | 2] es | Bele 
3 o S$ = 5 2A gy] 
SSPE ES ads (Euston ae | ae (ee 8 |e ee ee ee se ie 
= 
we eres pi saesee: [eeeeee fiimesene [i CHES {| SE eeseee [ii seuwen || cuncne | cvauen f susece | savese | (senses | srene * * * 
mf | feces iceewee | Pf YP ceecce | ceceee | | wevene | ceeeee fi ceeeee | ccceee | ceceen | ceesee | teneee | cesses * 
S| eee | aaa | A eseeadl|eseeeod lessees 
Sef I cee leanne |i eecea eeeno lll Seeeee [ic osewn) || wecurn | vecees ||| eeene= | uunmac'G ccvses | eevee | secone | cevene * 
1} Be * 
; J rere) 
* MH cecece 
Ho] wacees * 
MO oceeece * 
* 
* 2 
= * . 
* | 
« | 
* | HO osneene 
* . 
en l(encceceul| Gastipe 
Hf ewaee | cervee 
* * 
* 
el 
| ee ee eee 
. an eter Ts * 
2 * * 
* * * 
* * . 
— << i * 
ee ‘ conece: | herrecoe |) ceooce | acetce: |) atacos 1} cocece ? 
Nemeriies Obvantii, Murch, 
> * 
Tertoerrs. 
Calymene. sp. 
coerey ¥ 
| | 
Bs ass |. serene Wiesarecd|it-cssmillsrenee | teraction | [tensor || tsaseas oh lWhecerxo * 
oP ecco * 
esacee toe A peek | recap Woseree 2 WN eeeceadd| nccee \Weeeepe | ecacce || cone |) Goch * 


| Gala Group. 


Z 
* Confined to the highest 1 nari 
Pageant Dae pata qu she Br Birkhill of South Scotland eae ° A few fraginents only in the highest seams of U, per Birkhill (Lapw.), 
2 Known only in ee Birkhill bed Shale and its equivalents (Lapw,), ° M, runcinatus is peculiar to Upper Birkhill Shales Lapw,). 
* Restricted to a single zone in the Te Moffat and Girvan (Lapw.), 7 Makes its first appearance in middle of U: per Birkhill (Lapw.) 

oper Birkhill Shales (Lapu.). * Diployrapsus Hughesié is confined to the Upper Birkhill Shales (Lapw.). 


Quart. Jour. Geol. aha Volo ea ae 


Mintern Bros. imp. 


INES Hoomdeliti 


WELSH CLADOPHORA. 


ON SOME NEW SPECIES OF CLADOPHORA. 171 


APPENDIX. 


On the Cravopnora (Hopk.) or Drenproip GRaprorirss collected by 
Professor Kuxpine in the Llandovery Rocks of Mid Wales. By 
Cuas. Lapwortu, Esq., F.G.8. &e. 

[Puate VII.| 


Tse forms of Cladophora collected by Professor Keeping from 
the Llandovery rocks of Cardiganshire are, regarded collectively, of 

a type almost new to British paleontology. “Although intimately 
allied to the well-known dendroid species of the Quebec and Arenig 
formation, they are very distinct in their minor features. They are 
essentially of a Silurian (Upper) facies, and they remind us strongly 
of a group recently made known to us through the researches of Mr. 
Spencer, of Toronto, which characterizes the Clinton and Niagara 
Groups of New York and Upper Canada *. 

None of our examples shows the complete polypary, or affords dis- 
tinct proof of the presence of all the more minute classificatory 
features ; but there is satisfactory evidence of the presence of at 
least four distinct genera and seven species. The genera repre- 
sented are Dictyonema, Hall, Calyptograptus, Spencer, -Acantho- 
graptus, Spencer, and Odontocaulis, Lapw. Dictyonema is a well- 
known British genus; the remainder are new to British paleon- 
tology. Calyptograptus and <Acanthograptus have been already 
briefly noticed from American strata by Mr. Spencer, but have not 
hitherto been figured. Odontocaulis is a new genus, of a peculiar 
type. 

: Of the seven distinct species recognizable, four (viz. Dictyonema 
corrugatellum, Calyptograptus plumosus, C’. digttatus, and Odontocarulis 
‘Keepingw) are as yet peculiar to the Mid-Wales area. The remaining 
three have already been recognized by myself in the Silurian rocks 
of the south of Scotland.  Dictyonema venustum certainly occurs in 
the Upper Gala group of Selkirkshire (Llandovery-Tarannon), and 
doubtfully in strata of a little older date in the Girvan area. 
Dictyonema delicatulum has been met with in the Llandovery (Lower) 
of Shalloch Forge, near Girvan. Acanthograptus ramosus occurs 
also im the Girvan area, at the base of the representative of the 
Tarannon Shales. 

We do not yet know enough of the Cladophora of North America 
to enable us to identify any of these Mid-Wales forms with corre- 
sponding species already described or figured from the Silurian strata 
of that continent. 


Genus DictyoNnEMA. 


Dictyonema, Hall, Paleontology, New York, vol. ii. 
1. DicryoNEMA VENUsIUM, sp. nov. Plate VII. figs. la—le. 


Polypary cyathiform in the growing state, at least 10 inches in 
- transverse diameter. Branches uniform, about one eightieth of an 


* “Graptolites of the Niagara Formation,” by J. W. Specs, F.G.S8., Canadian 
Naturalist, 1878-9, pp. 457-463. 


172 C. LAPWORTH ON SOME NEW 


inch in transverse diameter, radiating from the centre with few and 
remote bifurcations; approximately parallel distally; having 25 
branches to the inch, connected by frequent capillary dissepi- 
ments 12 or 13 to the inch. Calycles biserial, closely adpressed to 
the polypary, 50 to the inch; apertural margin acute, rarely spinose. 

The largest specimens obtained are distal fragments of the 
polypary, about two inches in length ; and from the characteristics 
presented by these fragments the foregoing diagnosis has been 
drawn up. 

This form is undoubtedly new to British paleontology. It may, 
however, be proved in the future to be identical with some of the 
imperfectly known American forms from the Clinton and Niagara 
eroups. 
~ Locality, Llandovery of Devil’s Bridge, Aberystwyth. Dictyonema 
venustum is a rare fossil in the Llandovery rocks, of the south of 
Scotland, at Abbotsford, Winterhope, and Penwhapple Glen. 


2, DicrYoNEMA DELICATULUM, sp. nov. Plate VII. figs. 2a, 26. 


Polypary cyathiform ? probably three or four inches in transverse 
diameter. Branches 40 to the inch, about 54, of an inch in 
width, distally parallel, connected by indistinct capillary dissepi- 
ments 16 or 17 to the inch; bifurcations? Calycles biserial, 80 to 
the inch ; aperture horizontal, devoid of ornament. 

The best example of this species collected is shown in the figure, 
Plate VII. fig. 2a. . The characters it presents show that it 
belonged to a distinct species from that already described, from 
which it differs mainly in the narrower and more closely-set 
branches, and in the more numerous calycles and dissepiments. 

Horizon and Locality. Llandovery of Bryn y Carnau, Aberyst- 
wyth. 

a single specimen has also been collected from the Pentamerus- 
beds of Shalloch Forge, Girvan, South Scotland. 


3. DicTYoNEMA CORRUGATELLUM, sp. nov. Plate VII. figs. 3a, 36. 


Polypary cyathiform, about one aud a half inch in height. The 
branches are nearly parallel, about ;}5 of an inch in diameter, and 
zis of an inch apart, connected by fine transverse dissepiments at 
regular distances of about =, of an “inch, undulating in direction, the 
edees being thrown alternately from side to side by the projection 
of the outer margin of the calycles. Calycles 120 to the inch, 
alternate, with gibbous outer margin, and deep rounded aperture 
lying interior to the edge of the branch. 

The most striking feature of this elegant little species is formed 
by the wrinkled or undulating outer edges of the branches, which 
wave from side to side owing to the projection of the ventral 
margin of the calycles. The slender branches divide more 
frequently than is the case with the generality of species of 
Dictyonema ; and the transverse dissepiments are stouter and appa- 
rently polypiferous. 

The calycles have their apertural margin strongly introverted, 


SPECIES OF CLADOPHORA. LS 


in the manner of those of Dicellograptus, Hopk.; and the line 
of the aperture lies wholly within the ventral margin of the 
branch. 

Horizon and Locality. Llandovery of Devil’s Bridge, Aberyst- 
wyth, Cardiganshire. Collection of Dr. Humpidge, Aberystwyth. 


Genus CALYPTOGRAPTUS. 


Calyptograptus, Spencer, Canadian Naturalist; 1878-79, p. 459. 


“Gen. char. Polypary cyathiform, with numerous bifurcating 
branches, which are dichotomous at the termination, but are not 
connected by lateral processes. The branches are marked with 
strie resembling rhomboidal pits; the axis has a black corneous 
exterior; and the radicle is composed of a thickened mass of the 
same texture as the branches. In appearance and texture this genus 
resembles Dictyonema; but the branches are all independent, not 
being connected by transverse dissepiments as in that genus, and are 
only united in one mass at the root” (Spencer, loc. cit.). 

One of the Cladophora collected by Professor Keeping appears to 
possess many of the foregoing characters, which are given by Mr. 
Spencer as the peculiarities of his genus Calyptograptus. ‘There are, 
however, some marked distinctions between our species and his 
typical examples; nevertheless it is most undoubtedly a closely 
allied form, and may therefore most conveniently be provisionally 
placed in this genus until we know more of the proper generic 
distinctions of these obscure fossils. 


4, CaLYPTOGRAPTUS? PLUMOSUS, sp.nov. Plate VII. fig. 4. 


Polypary cyathiform?, in the growing state, about one inch and 
a half in height, composed of numerous bifurcating polypiferous 
branches united into a short stem, longitudinally striated, not 
connected by transverse dissepiments. Calycles closely arranged, 
biserial. Basal disk elliptical in form, about one fourth of an inch 
in diameter. 

The stem, which is very short and stout, rises from the centre of 
the basal disk, and divides for the first time within about one eighth 
of an inch of its origin; the second, third, and fourth subdivisions 
are about one tenth of an inch apart; the later bifurcations are a 
little more remote. The primal branches are about one thirtieth 
of an inch in diameter ; the final branchlets are almost capillary. 

The substance of the polypary appears to have been somewhat 
membranous in character, and is wrinkled or striated longitudinally 
in the fossil. Two series of calycles are just discernible in the 
proximal portions of some of the branches, their apertural margins 
being éxhibited as short slits crossing the branches transversely, 
almost at right angles to their general direction. 

This form agrees with Dictyonema and Callograptus in the regular 
dichotomous nature of the method of subdivision of its principal 
branches. It differs, however, most markedly from both these 
genera in the apparent absence of the transverse dissepiments 


174 Cc. LAPWORTH ON SOME NEW 


uniting the branches. In the mode of branching and in the form of 
the calycles it approaches the genus Odontocaulis, to be presently 
described, from which, however, it is easily separated by its short 
non-polypiferous stem. 

Horizon and Locality. Llandovery of Devil’s Bridge, Aberyst- 
wyth. 


5, CALYPTOGRAPTUS? DieiTaTUS, sp. nov. Plate VII. figs. 6a—6b. 


Polypary short, composed of numerous compound branches about 
roy of an inch in width, dividing irregularly, and terminated distally 
by a group of palmatifid branchlets about one tenth of an inch in 
length. Calycles of the type of those of Callograptus, about 50 to 
the inch. 

The magnified drawing gives a fair idea of the external features 
of the only specimens seen, which are mere fragments. The 
proximal extremity of the stem is unknown; but the irregular mode 
of subdivision of the branches is different from what generally 
occurs in Dictyonema and its allies. There is a doubtful appearance 
of reticulation among the secondary branches. 

The most remarkable feature of this form is the strange digitate 
character of the final spine-like branchlets. Mr. Spencer has noticed 
the same feature in the genus Rhizograptus (op. cit. supra, p. 461), 
to which itis possible the present species properly belongs. 

Horizon and Loculity. Llandovery of Devil’s Bridge, Aberyst- 
wyth. 


Genus ACANTHOGRAPTUS, Spencer. 


Gen. char. “ Polypary shrublike, consisting of thick branches, 
principally rising from near the base, with little divergence and 
some bifurcations. One side of the branches is furnished with 
prominent spines or denticles, which appear to mark the cell- 
apertures. Test corneous and indistinctly striated.” 

‘«« This generic form resembles Dendrograptus, but is stronger and 
more bushy than species of that genus, and has conspicuous spines 
indicating a different cell-structure ” (Spencer, op. cat. p. 463). 

The foregoing is Spencer’s diagnosis of his new genus Acantho- 
graptus. I have provisionally assigned to it one of Professor 
Keeping’s species, which answers fairly enough to Spencer’s defini- 
tion. It is impossible to ascertain from Spencer’s diagnosis whether 
the denticles that ornament the calycles are horizontal or inclined. 
if the former be the case, I suspect that Acanthograptus will be found 
to lie somewhere near Thamnograptus of Hall, in which the calycles 
are provided with long projecting spines, and that the present form 
must be regarded as the type of a new genus. 

Acanthograptus occurs both in the Bala and Llandovery strata 
of Girvan, South Ayrshire, where, however, it is an uneommon 
fossil. None of the species collected by myself from the Scottish 
locality appears to be precisely identical with the Mid- Wales form. 


6, ACANTHOGRAPTUS RAMOsUS, sp. nov. Plate VII. fig. 5. 
Polypary short, shrublike, with thick rigid branches, repeatedly 


SPECIES OF CLADOPHORA. WG 


and irregularly branching and rebranching. Base unknown. Calycles 
monoprionidian ?, often with long acute denticles, which have an 
ascending direction. 

We have only one fair example of this genus from the Mid-Wales 
beds; and the foregoing characters are all that can be made out. 
The height of the visible portion of the polypary is less than half an 
inch. The main branches are apparently about one fortieth of an inch 
in diameter below, and are strongly striated throughout the whole of 
their extent. The right-hand branch, which is the better preserved, 
throws off three secondary branches in the upper half of its length. 
These secondary branches are of various lengths, being so extended 
that their outer terminations are on, or about, the same level. Each 
secondary branch splits at its summit into two secondary branchlets 
or polypiferous spines, about one twentieth of an inch in length, 
and terminating outwards in a short, stiff, mucronate point. The 
remaining primary branches present essentially the same general 
features. 

One margin of the branches exhibits small denticulations at short 
and regular intervals (about 50 to the inch), marking the mouths 
of the calycles, the margins of which are usually concave. The 
outer angle of the aperture is sometimes very slightly projecting and 
more or less rounded off, as in the genus Dictyonema. Generally, 
however, the outer margin of the aperture is prolonged into a stout 
denticle, which stretches outward and upward at an angle of about 
45 degrees to the main axis of the branch, and is terminated in 
a blunt point; the inner margin of the calycle, shown by a little 
groove upon the surface of the branch, running for some distance 
almost parallel with the outer margin. 

The colour of the test is black, and the texture corneous. 

Horizon and Locality. Llandovery of the Devil’s Bridge, Aberyst- 
wyth. 

ODONTOCAULIS, gen. Nov. 

Gen. char. Polypary cyathiform, composed of numerous indepen- 
dent and frequently bifurcating polypiferous branches, originating 
from the distal extremity of a short stem, which is likewise poly- 
piferous, and is terminated proximally in an irregular corneous 
expansion. Hydrothece of the type of those of Dictyonema, 
biserial, subalternate. 

The chief peculiarity of this genus is afforded by the character of 
the stem, which is identical in every respect with the main branches, 
and, like them, is denticulate or polypiferous throughout the whole of 
its extent. It commences proximally in a flattened expansion, with 
irregular or frayed-out edges, possibly the remains of a disk or bulb 
of attachment. 

The mode of branching is rigidly dichotomous, the first two 
branches being formed by the subdivision of the main stem itself. 
Each arm branches and rebranches again and again in the same 
manner, at irequent and close intervals,-composing an elegant cyathi- 
form or fanlike polypary, very symmetrical in form. The branches 
retain their original width to their final division, which gives rise to 
two minute branches less than one tenth of an inch in length. 


176 C. LAPWORTH ON SOME NEW 


The hydrothece are more prominent than those upon Dictyonema. 
The distal extremity of each appears to have been free and slightly 
introverted, as in the majority of the bilateral family of the 
Dicranograptide. 

Odontocaulis is separated from Dictyonema by the absence of the 
transverse dissepiments, and by the polypiferous character of the 
stem. From Callograptus, which it much resembles, the same 
features effectually distinguish it. In Dendrograptus the stem is 
stout and devoid of polypes, while the branches are irregularly 
disposed ; in the present genus the stem is no thicker than the 
branches, is polypiferous, and the branches are regularly and 
symmetrically subdivided. It has probably its nearest ally in 
Rhizograptus (Spencer, ‘ Canadian Naturalist,’ 1879, p. 460); but in 
that genus the stem appears to be barren, and the branches are 
possibly united at intervals. 


7. OpontocavLis KEEPINGII, sp. nov. Plate VII. figs. 7a, 70. 


Polypary composed of numerous elegant flexuous branches, 
frequently divided and subdivided in a regularly dichotomous 
manner, originating from the distal extremity of a polypiferous 
stem about one fourth of an inch in length, and forming a 
cyathiform frond one inch and a half in height. Hydrothece 50 
to the inch, free, distally patulous ; aperture prolonged, introverted ; 
denticle obtuse, rarely spinose. 

In the only specimen coliected the polypary commences proximally 
with a flattened expansion or disk with irregular or frayed-out 
edges. The stem is curved ventrally, and bears a single series of 
hydrothecze, which are most distinctly shown. There are also indi- 
cations of a second series. These hydrothece are a little more widely 
separated than those upon the branches, and appear to have been 
stronger and more projecting. At its summit the stem divides into . 
two, originating the primary branches. Within one twentieth of 
an inch of their origin each of these again divides in a correspond- 
ing manner. Within the next tenth of an inch the branchlets 
are again subdivided, and so on till as many as six of these 
dichotomous divisions have been made. The terminal branchlets 
are very short, less than one tenth of an inch in length; but they 
are polypiferous to their visible extremities. 

The hydrothecz upon the branches average about 55 or 60 to the 
inch. The majority are seen as scaliform impressions; those 
visible in profile are not unlike those in Dicellograptus, Hopkinson. 

The branches are of subequal diameter throughout, about one 
fortieth of an inch. They are quite free and independent, neither 
inosculating, asin some forms of Dictyonema, nor being connected 
by transverse dissepiments, as in that genus and its ally Callo- 
graptus. 

This beautiful little species is dedicated to its discoverer, Professor 
Walter Keeping. 

Horizon and Locality. Llandovery of Devil’s Bridge, Aberyst- 
wyth, Cardiganshire. 


SPECIES OF CLADOPHORA. 177 


- EXPLANATION OF PLATE VII. 


Fig. 1. Dictyonema venustum, sp.nov.: 1 a, natural size; 10, magnified 10 dia- 
meters. Locality: Devil’s Bridge, Aberystwyth (coll. W. Keeping). 
1 ¢, nat. size. - Loc.: Williamshope, Gala group (coll. C. Lapworth). 
2. Dictyonema delicatulum, sp. nov.: 2a, nat. size; 26, magnified 10 dia- 
meters. lLoc.: Bryn y Carnau (coll. W. Keeping). 
3. Dictyonema corrugatellum, sp.nov.: 3a, nat. size; 3b, magnified 10 dia- 
meters. lLoc.: Devil’s Bridge (coll. Dr. Humpidge, Aberystwyth). 
4. Calyptograptus plumosus, sp. noy.: nat.size. Loc.: Devil’s Bridge 
(coll. W. Keeping). 
5. Acanthograptus ramosus, sp. nov.: magnified 3 diameters. Loc.: Devil’s 
Bridge (coll. W. Keeping). 
6. Calyptograptus digitatus, sp.noy.: 6 a, nat. size; 66, magnified 5 dia- 
meters. Loc.: Devil’s Bridge (coll. W. Keeping). 
7. Odontocaulis Keepingit, gen. et sp. nov.: 7 a, nat. size; 7 4, basal por- 
tion magnified 5 diameters. Loc.: Devil’s Bridge (coll. W. Keeping). 


Discussion. 


The Presipenr said that the paper was an important contribution 
to the geology of Mid Wales. The fossils were interesting and 
remarkable, especially the Graptolites. 

Prof. HueHEs pointed out that the North-Wales Silurian was 
quite different from that of South Wales, and that the area described 
by the author was far from the typical sections of either north or 
south and differed from both. He considered the paper a valuable 
contribution to our knowledge. 

Dr. Hicks was glad to hear that in this area also, as in Pembroke- 
shire, there was no break between the Upper and Lower Silurians. 
It was also satisfactory to see that fossils had turned up on 
examination in these rocks, formerly reported to be barren or 
nearly so. 


178 G. W. SHRUBSOLE—-FURTHER NOTES ON 


13. Further Notes on the CARBONIFEROUS IENESTELLIDE. By GuORGE 
Wau. Surussore, Hsq. F.G.S. (Read January 19, 1881.) 


In a former communication to the Society L endeavoured to show 
that, as the result of a careful comparison of the existing individual 
forms, as figured by previous describers, with some tolerably 
perfect specimens in my own possession, it would be necessary to 
redescribe the species of Wenestella and considerably reduce their 
number. I accordingly append a redescription of the more promi- 
nent species ; but before introducing that, it will be necessary to say 
something about the foundation of the genus Menestella, since it can 
readily be imagined that the same causes which have thrown con- 
fusion into the species, have tended also to obscure the genus— 
namely the fragmentary and imperfect state of the specimens ex- 
amined, and the want of attention to the stages of individual growth 
and other details. Accordingly I find the existing descriptions of the 
genus Menestella inaccurate in many essential details. The genus 
has been defined by several paleontologists, but each time with 
reference to the fossils of a particular period; and no one description is 
sufficiently comprehensive to embrace all the members of the family 
Fenestellidee as they occur in the various Palseozoic formations. A 
fresh description of the genus therefore appears to be desirable. 

Without entering upon a full history of the genus Fenestella, I 
may mention that the first to describe it as such was Lonsdale, in 
1839*, who adopted the name from the MS. of Miller of Bristol, 
who had been engaged on a work on the Corals of the Mountain 
Limestone, which he did not live to complete. He gave the name 
Fenestella to the lace corals. enestella Millert of the Silurian is 
so named after him. Lonsdale twice defined the genus—first in the 
‘Silurian System,’ and again in the ‘ Geology of Russia’r. In the 
former he restricted the cells to one row on each side of the keel : 
and in the latter he placed no such limit to the order or arrangement 
of the cells ; hence it included Polypora, Retepora &c., in fact all the 
fenestrate species. His otherwise elaborate definition was also 
faulty to the serious extent that he described the appearance of a 
common incrusting coral as the mature growth of the Menestella. 
This [ explained in a recent communication to the Society. 

Phillips next in order of time (1841) gave a very good description 
of the genus, so far as relates to the Devonian groupt; but much vf 
its details are not applicable to the Silurian group. 

Prof. M‘Coy, in 18448, restricted the genus to forms with two rows 
of pores on the branches, so as to include only the true Lenestelle. 
The pore-cells he limits to the external face, whereas in the majority 
of the species they are on the inside of the polyzoary. He further 


x Murchison’s ‘Silurian System,’ p. 677. 
t Geology of Russia, vol. i. Appendix A, p. 629. 
t Paleeozoic Fossils, pp. 22, 25. 


§ M‘Coy’s Syn. Carb. Foss. Treland, p. 290. 


THE CARBONIFEROUS FENESTELLIDZ. 179 


states that the perfect condition of Fenestelia will be found in his 
genus Henutrypa. ‘This latter I have shown* to be a mistake due 
to an incrusting organism on Jenestella membranacea, Phil. 

In his second definition, in 1854 ++, which was intended to include 
both Carboniferous and Silurian species, M‘Coy is more successful ; 
no mention is made of the particular face on which the cells are 
placed. He does not adequately describe the shape of the polyzoary, 
and adopts one of Lonsdale’s errors respecting a layer of vertical 
capillary tubes on the reverse face. 

The last definition we have was by Prof. King t in 1848, after 
arranging the Permian Polyzoa. He ignores M‘Coy’s work in 
1844, and says that the genus, as constructed by Lonsdale in 1845, 
requires subdividing. As no trace of the incrusting coral was found 
in the Permian species, all reference to Lonsdale’s supposed mature 
condition of the Fenestella is omitted. Prof. King’s short and other- 
wise exact account is inexact in stating that the cells are distributed 
in two or more linear series.. The true Fenestclle have never 
more than two rows of cells on the interstice ; the supposed row on 
the keel does not really consist of cell-pores, but of the bases of hollow 
Spiny processes, which oftentimes, in the case of the Permian /ene- 
stelle, have been unduly distorted by a subsequent deposition of 
carbonate of lime. 

From the foregoing it is evident that none of the origina] deseri- 
bers of the genus Fenestella has grasped the main features of it as 
a whole. What is wanted is a definition which shall embrace the 
whole of the Paleozoic Menestelle. 

The following definition is therefore proposed, after a careful study 
of the several species ranging from Silurian to Carboniferous times. 


Genus FrnEsTELLA, Lonsdale. 


Polyzoary a calcareous reticulate expansion, either flat, conical, 
or cup-shaped, formed of slender bifurcating branches (interstices), 
poriferous on one face, connected by non-poriferous bars (dissepi- 
ments) forming an open network. Ced/s immersed in the interstices, 
and arranged in two longitudinal rows divided by a central keel, on 
which are often prominences. Cell-mouth small, circular, and 
prominent when preserved. 


FENESTELLA PLEBEIA, M‘Coy, Syn. Carb. Foss. Ireland, p. 29, 
fig. 3. 


Fenestella antiqua, Lonsd., M‘Coy, Syn. Carb. Foss. Ireland, 
p. 200. | 
carinata, M‘Coy, Syn. Carb. Foss. Ireland, pl. 28. fig. 12. 
devonica, Semenow et v. Moller, Bull. de l’Acad. de St. 
Pétersbourg, t. vii. p. 233, pl. 3. fig. 16. 
formosa, M‘Coy, Syn. Carb. Foss. Ireland, pl. 29. fig. 2. 
flustriformis, Phill. Geol. Yorkshire, pl. i. figs. 11, 12. 
fossula, Lonsd., Darwin’s Obs. on Volcan. Isl. p. 166. 


* Quart. Journ. Geol. Soe. vol. xxxv. p. 282. 
t Brit. Pal. Foss. p. 49. ¢ Permian Fossils. p. 34. 


180 G. W. SHRUBSOLE—-FURTHER NOTES ON 


Fenestella fossula, Dana, Geol. U.S. Explor. Uxped. p. 710, pl. 1. 
fig. 3. 
**___ nlebeia, POrbieny, Prod. de Paléont. strat, t. i. p. 152. 
—— irregularis, Phill. Geol. Yorkshire, pl. 1. figs. 21, 22. 
—— retiformis, Schloth., King’s Perm. Foss. pl. 2. figs. 8-19. 
—— trituberculata, Prout, Trans. Acad. St. Louis, vol. 1. p. 228. 
— undulata, Phill. Geol. Yorkshire, pl. 1. figs. 16-18. 
—— virgosa, Kichwald, Lethea Rossica, tom. 1. p. 358, pl. 23. 
9 


Sp. char.—Polyzoary a flat expansion, slightly convoluted, circular 
or oval in outline, depressed in the centre, by which it is attached. 
Interstices regular on the obverse face when well preserved, somewhat 
rounded otherwise, sides angular, keeled. Drsseprments thin, expand- 
ing at junction with interstice, more markedly on obverse than re- 
verse face. enestrules regular, oblong, equal to width of interstice 
on obverse face, twice the width on reverse face ; four in the space of 
two lines longitudinally, and six fenestrules in the space of two lines 
transversely. Acel rounded and strong, with two or three nodes in 
the length of a fenestrule. Pores round, prominent, their diameter 
apart, two or three in the length of a fenestrule, and often one more 
prominent than the rest in the angle formed by the junction of the 
dissepiment with the interstice. 

Obs. This species, whether from Scotch, Irish, or English localities, 
is everywhere the predominant form of Menestella. It attained to 
the largest expansion of polyzoary of any of the species. It de- 
serves in every way to be regarded as the typical species of the 
Carboniferous varieties. I have traced its growth through all its 
stages, from a speck with two interstices on a stalk which clasped, it 
might be, the spine of a Productus or a fragment of Serpula, to the 
adult form with its strong and numerous rootlets. The early leaf- 
like growth on a footstalk soon underwent a change; the footstalk 
became one of many rootlets, and the polyzoary coralliform at the 
base, and ultimately a more or less circular expansion, the edges of 
which terminated in slightly convoluted lobes. Its attachment to 
the rock was secured by a cluster of rootlets from about the base, 
and, indeed, from any part of the polyzoary which offered convenience 
of attachment. 

The actual size to which this species attains it is difficult to 
estimate, owing to the cleavage of the shale in which it occurs. I 
have seen indications which lead me to believe that an adult poly- 
zoon might attain a diameter of nearly two feet. Specimens of 
this species will be found to differ from each other somewhat in 
appearance. ‘This I have found is more owing to the nature of the 
matrix in which it has been imbedded than to any other cause. 
The more calcium carbonate present in it, the fuller and more life-like 
the organism appears, while it is flat and shrunken in the ordinary 
black aluminous shale of the Carboniferous beds. 

In diagnosing the species the first thing to be noted is the size of its 
interstice, in which it is intermediate between Fenestella nodulosa, 


THE CARBONIFEROUS FENESTELLID&. 181 


Phill, and Lmestella polyporata, Phill. There is also a peculiar 
smoothness and regularity, in the growth alike of interstice and 
dissepiment, which is attained by none of the other species and is 
a distinguishing mark. The presence of three or four pore-cells in 
a given space, the dissepiments, and the oblong fenestrules complete 
the identity. Prot. M‘Coy’s fine drawing of this species: is taken from 
a partly worn-down specimen in which the sides of the interstice have 
become sharp and angular, and the keel scarcely visible; whereas the 
interstice should have been full and rounded, and capped with a ridge 
or keel, bearing the remains of spiny projections, while on either side 
the cells stood out prominent and free. , 

There is no doubt about the identity of this species with Gorgoma 
antigua, Goldt., although his figure is drawn from a much weathered 
and mutilated fragment. Tt also agrees with Fenestella retiforinis, 
Schloth., the leading Permian species. Jam aware that a contrary 
opinion has been expressed. Virtually the species are the same. 
Phillips was, in this country, the first to give a descriptive account 
of it. His Menestella wreqularis, F. undulata, and F. flustriforimis 
are all different portions of the polyzoary, under somewhat varying 
conditions of growth and preservation. I cannot agree with Prof. 
M‘Coy in assigning Fenestella flustriformis to Ptylopora» flustri- 
jormis. As figured by Phillips, it is only the cast in limestone 
of the reverse face; and, as such, its relationship to Ptylopora 
is purely conjectural. Prof. M‘Coy was the first to recognize the 
importance of this species, and to do justice to it in ‘the way 
of description; and hence, although not its author, that it should 
retain the name which he gave to it is eencrally conceded. In 
this view I am borne out by Prof. de Koninck, who states ‘+ that 
Lonsdale was the first to describe this species as Fenestella fossula, 
but that his description was incomplete, and insufficient to recognize 
it with certainty, whilst M‘Coy’s description was full and accurate’ oe 
Phillips’s notice of it some years prior to that of Lonsdale was even 
more incomplete. For the foreign synonyms of Menestella plebeia 
I am also indebted to Prof. de Koninck’s work on the Carboniferous 
Fossils of New South Wales. 


FENESTELLA MEMBRANACEA, Phil. Geol. Yorks. pl. i. figs. 1-6. 


Fenestella flabellata, Phill. Geol. Yorks. pl. i. figs. 7-10. 
henuspherica, M‘Coy, Syn. Carb. Foss. Irel. pl. xxix. fig. 4. 
—— Shumardiu, Prout, Trans. Acad. St. Louis, p. 232. 
tenurfila, Phill. Geol. Yorks. pl. 1. figs. 93, 24, 25. 

Sp. char.—Base cylindrical, tapering to a “fine point, vw with strong 
non-poriferous rootlets attached. Upper part widely expanded. 
From nine to twelve inches in length, and five to eight ches in 
width. Jnéerstices rounded, straight, in parallel lines, keeled. Dis- 
sepiments fine, shghtly expanded at junction with interstice. Henes- 
trules oblong, slightly wider than insterstice, from two to three times 
as long as wide. At one inch from base, five dissepiments in the 
space of two lines measured vertically, and eight dissepiments in. 

* Hoss. Pal. Nouy. Galles du Sud, 1877. 

Q.J.G.8. No. 146. 0 


182 G. W. SHRUBSOLE—-FURTHER NOTES ON 


two lines transversely. Fenestrules gradually increasing in size 
with the growth. Pores small, round, more than their own diameter 
apart, placed on the outer face of the polyzoarium. At the base 
three pores within the length of a fenestrule ; in the upper portion 
four in the same space. 

Obs. There are many points of considerable interest attaching to 
this species. In shape and appearance it is an enlarged type of 
the Silurian species generally, and particularly of my Fenestella 
lineata. It is the only representative of the old type which has 
survived to Carboniferous times. In addition to its shape, it has 
most of the features of its prototype, viz. interstices seldom bifurca- 
ting, proceeding in parallel lines, and pores on outer face of polyzoary. 
Its early growth, from a minute point, was a tapering, often curved 
root-base, from which grew a hollow and gradually widening cone, 
which ultimately expanded in slightly folded and lobed outlines 
around the aperture. ‘To secure in position so fragile a structure, 
numerous solid rootlets grew from various points of the base, and 
attached themselves to surrounding objects. Being somewhat cylin- 
drical, 1t did not need to make the usual amount of lateral growth 
by bifurcation, as in the ordinary open type of Carboniferous 
enestelle. Owing to this comparative absence of bifurcation in 
the insterstice, its lines are singularly regular, forming a series of 
parallel rows. Further, its lateral growth was obtained by a gradual 
enlargement of all parts of the structure, proceeding from the base 
upwards—so much so that while at the base four fenestrules may be 
counted in one line, at the distance of two inches there are only two 
in the same space, thus doubling the circumference of the polyzoary 
without the aid of division of the interstice. This enlargement in 
the growth is more or less persistent throughout the polyzoarium, 
but not to the same extent, and furnishes at once a key to the 
synonyms. Accompanying the increased growth, an additional 
pore-cell may be noticed between the dessepiments. ‘Thus the 
difference between the base and the upper portion, both as to size 
and shape, is of a very marked character, and quite accounts for the 
several species into which this one has been divided. Phillips was 
the earliest worker at it; he named the extreme base VFenestella 
membranacea, the upper and enlarged growth Fenesiella flabellata, 
and the more delicately formed and intermediate portion Menesiella 
tenurfila. 3 : 

I have alluded to the fact that in the more cylindrical portion of 
FF’, membranacea bifurcation of the insterstice was arrested in part. 
At times a variety of circumstances may have hindered the longi- 
tudinal extension of the polyzoary ; then we find a rapid bifurcation 
which gives a globose outline to the expansion, and the form is then 
the Ff. hemispheriea of M‘Coy, while all its other details as to the 
pores, interstices, &c. clearly point to its identity with /. membra- 
nacea, Phill. 

The only species likely to be confounded with the foregoing is the 
F, nodulosa of Phillips. The square form of the fenestrule in the 
latter will at once indicate its character. 


THE CARBONIFEROUS FENESTELLID2:. 183 


Locality. This species is comparatively rare in the Mountain- 
Limestone beds of England and Scotland, and very abundant in 
certain localities in Ireland. 


FENESTELLA NopuLosA, Phill. Geol. Yorks. pl. i. figs. 31, 32, 33. 


Fenestella bicellulata, R. Kth., Jun., Mem. Geol. Surv. Scotl. Sheet 
Bo, p. 101. : 

—— fruiex, M‘Coy, Syn. Carb. Foss. Ireland, pl. xviii. fig. 10. 

— Popeana, Prout, Trans. Acad. St. Louis, p. 229. 
subretiformis, Prout, Trans. Acad. St. Louis, p. 233. 


Sp. char.—Polyzoarium, early growth foliaceous, having a distinct 
stem or footstalk, becoming an oval or circular expansion. Jnter- 
stices regular, rounded, carinated and bifurcating ; remains of spiny 
processes along the keel. Dussepiments thin, rounded, regular, a 
little arched in the early stage, not so thick as interstices. enes- 
trules square, regular in the early growth, four fenestrules in the 
space of one line measured longitudinally, and four fenestrules in 
the same space measured transversely. In the later and upper 
erowth there are three fenestrules only each way in the same 
measurements. Cells small, round, their diameter apart, one com- 
monly at the end of each dissepiment, and one between, or three 
to each fenestrule: this is in the early growth; the later growth 
has four in the same space. When the cells occur in the angle 
formed by the junction of dissepiment with insterstice, the former 
is expanded atthepoint. ‘This feature is not constant. Cell-mouth, 
when preserved, nearly on a level with the keel. 

Obs. This is a very marked and easily recognized species ; its 
square fenestrules and the round-wire-like nature of the interstices 
and dissepiments on the reverse face at once distinguish it from all 
others. In mature specimens the reverse has a peculiar and 
characteristic nodular aspect. Im size it is somewhat minute, 
being intermediate between Venestella membranacea, Phil., and 
Fenestella plebeca, M‘Coy. Both Phillips and Prof. M‘Coy were 
unfortunate in the fragments which they selected for description ; 
it is not surprising, therefore, that when a good representative 
specimen was found it should be described as a new species. 
Hence by far the best description of Fenesiella nodulosa, Phill., is 
that given for Fenesiclla bicellulata, R. Ether., jun. 

In 1874 Dr. Young and Mr. John Young announced the disco- 
very of a new Carboniferous polyzoon, Actinostoma fenestraium, in 
which we have all the characters hitherto observed by Phillips, Prof. 
M‘Coy, and Mr. R. Etheridge, Jun., in Menestella nodulosa, with the 
addition of the cell-aperture terminating in a nipple-shaped pro- 
jection, the orifice of which was furnished with eight radiate denti- 
cles. To regard Actinostoma as the full development of /. nodulosa 
seemed the right course: as such I alluded to it in my former 
paper on the Carboniferous Fenestellide. Since then Mr. G. R. Vine, 
@ most accurate observer of the paleeozoic polyzoa, informs me that 
he has noticed the denticulate cell-aperture in Fenesiella plebaa, 
M‘Coy; while recently Mr. John Young, F.G.S., in a paper read 

02 


184 G. W. SHRUBSOLE—-FURTHER NOTES ON 


before the Glasgow Natural-History Society, mentions the occurrence 
of the same in Menestella tenuifila, Phill. Thus we have the record 
of this peculiar cell-aperture in two if not three species of /enestella. 
Notwithstanding this evidence in favour of the denticulate aperture 
in Fenestella, I now hesitate to give this feature wholly or in part 
to it, since | made the discovery that some of the species of 
Glauconome possessed a fenestration not very dissimilar to that of 
Fenestella, and might easily, in a fragmentary condition, be mistaken 
for it, indeed have been so. ~ There is no doubt about the fact that 
some of the species of Glauconome have the denticulate cell-mouth 
in question—Gilauconome stellipora, Young, for instance; on the 
other hand, it is not equally well established that the cell-mouth of 
Fenestella had the same characters. There is the possibility that 
some of the fragments of reputed Jenestella upon which the 
denticulate aperture was seen, may prove to have belonged to 
Glauconome. This discovery of the fenestrate polyzoary in Glau- 
conome considerably complicates the question of the nature and 
relationship of the palzeozoic Polyzoa; and it will require careful ob- 
servation on the part of palxontologists to work out the distinctive 
characters of the several genera, and assign to the various species 
the right fenestration. The result will have an important bearing 
upon Glauconome, more than on Fenestella, since we know so little 
of the life-form of the former, whereas the latter is better under- 
stood. It may be that both Glauconome and Fenestella and kindred 
Polyzoa, possessed the denticulate aperture. It is so far certain, 
as regards Glauconome; it may ultimately prove to be true of 
Fenestella. But for the present I consider that Glauconome has 
absorbed the existing evidence of the peculiar cell-aperture in favour 
of its claims. The other problems remain to be worked out. 

I will now allude to a connexion which has become apparent 
during this inquiry between #’nestella nodulosa, Phill., and Palwo- 
coryne, a hydrozoan originally described by Prof. Martin Duncan 
and Mr. Jenkins from the Lower Limestone shales of Ayrshire. 
With regard to my facilities for observing Palwocoryne, 1 may 
remark that it was described * from specimens washed from the 
shale, a process necessarily destructive of many of its more 
delicate and distinctive features. All the specimens in my posses- 
sion, on the contrary, are 7m setw on the shale or limestone in which 
they were found. Prof. Duncan, speaking of it, says that ‘“ usually 
they are attached by a dactylose pseudo-cellular base to the margins 
of the polyzoaria of Henestella’’>. My observations would lead me 
to limit the attachment of Paleocoryne to one species of Menestella, 
viz. Fenestella nodulosa, and to the pore-face generally, rather than 
the margin of the polyzoarium. ‘The frequency with which I 
noticed this association of Palwocoryne with Fenestella nodulosa, 
led me to go carefully over my collection, and ascertain definitely 
the particular species of enestella with which it was most fre- 
quently allied. The result was, that, out of ninety-seven specimens 


* Hixplan. Sh. 23, Geol. Surv. Scotland, p. 96. 
Tt Quart. Journ. Geol. Soc. vol. xxix. p. 413. 


THE CARBONIFEROUS FENESTELLIDZA. 185 


of Paleocoryne, eighty-five are either attached to the polypite- 
face of Fenestella nodulosa or associated with it, while twelve 
only are free and unattached. In no one instance is there a sus- 
picion that the species is any other than the one mentioned. Simi- 
lar evidence to this I get in another way. I have more than one 
hundred specimens of /. nodulosa from the shales of Halkin 
Mountain, but out of them not one example showing the poriferous 
face ; all without exception present to view the reverse side. This 
I explain by supposing that, as it is on the poriferous face of Penes- 
tella that Palcocoryne finds its seat, that face has in consequence 
held the more firmly of the two to the shale. ‘This is really so; for 
the base of it may be detected by a slight bulging of the polyzoary 
of the enestella, and its presence demonstrated by removing a 
portion. There is no similar difficulty in obtaining the obverse 
face of the other species from the same beds; nor do the other 
species present the appearance on the reverse face which I have 
noticed in /’. nodulosa. ‘The concurrence of these two species may 
be accidental; but I scarcely think so; the extent of its occurrence 
is not in favour of that idea; so that I think we may conclude 
that we have good evidence that Palwocoryne in the upper beds of 
the Mountain Limestone is exclusively confined to the polyzoary of 
Fenestella nodulosa, Phill. I may mention that I have found 
Palewocoryne associated with . nodulosa in the middle beds of 
the Mountain-Limestone series of North Wales, and also directly 
seated upon the poriferous face of the same from the Calciferous 
Sandstones of Scotland. I have previously pointed out that several 
inferior organisms are parasitic upon or incrust the polyzoaria of /e- 
nestella, from Silurian times upwards—such as Hemitrypa, Aulopora, 
Alveolites, and Diastopora ; and now Palcocoryne has to be added to 
the list. In the case of the previous incrustations there is no doubt 
that the parallel branches of the Menestella afforded suitable base- 
lines for the attachment of the incrusting coral. Whether there 
was any thing more than this in the preference shown by Palwo- 
coryne, is one of the problems to be worked out; and it will, IL 
have no doubt, receive due attention at the hands of Prof. Martin 
Duncan, to whom I have handed over my specimens of Palcocoryne 
for further elucidation. 


FENESTELLA PoLYPORATA, Phill. Geol. Yorks. pl. 1. figs. 19, 20. 


Fenestella multiporata, M‘Coy, Syn. Carb. Foss. Ivel. pl. xxvii. 
fig. 9. 
intermedia, Prout, Trans. Acad. St. Louis, p. 231. 
variabilis, Prout, Trans. Acad. St. Louis. p. 231. 


Sp. char.—Polyzoarium foliaceous at first, arising from a stem, 
ultimately becoming a flat circular network. Jnterstices large, 
broad, rounded, keeled. Dissepiments thin, one third the thick- 
ness of the interstice, somewhat irregularly placed, not expanding 
at junction with interstice. enestrules large and elongated, three 
times as long as wide. Four fenestrules in the space of two lines 


186 G. W. SHRUBSOLE—FURTHER NOTES ON 


measured transversely, and two fenestrules in the space of two lines 
longitudinally. cel rounded and well developed, marked along its 
course with numerous spiny processes. ores small and round, 
their own diameter apart, prominent when preserved, from five to 
nine in the length of a fenestrule. 

Obs. This is the largest species of Carboniferous Lenestella, as 
regards the size of its interstice and fenestrules, although not so as 
to the ultimate growth of its polyzoary, which is considerably less 
than that of F. plebea, M‘Coy. This peculiarity at once dis- 
tinguishes it from any other species. It is subject to considerable 
variation, and will be met with both larger and smaller than the 
one described. Prof. M‘Coy, in his arrangement of the Fenestelle, 
assigned the smaller type to Phillips’s /. polyporata, and the type 
with the larger development and greater number of pores he de- 
scribed as I”. multiporata; but as both conform so well to the type in 
other respects, there is no reason for this division. This species 
often, on the reverse, attains to the size and character of some of the 
Polypore, from which it may readily be known by having only a 
double row of pore-cells. 

Lenestella polyporata never occurs very freely in any locality, but 
seems generally distributed throughout the Carboniferous strata. 


FryEsrecta crassa, M‘Coy, Syn. Carb. Foss. Irel. pl. xxix. fig. 1. 


Fenestella laxa, Phill. Geol. Yorks. pl. 1. figs. 26-30. 

My previous notice of this species was limited to the extent of 
showing the identity in character between the species respectively 
described by Phillips and Prof. M‘Coy as VFenestella laxa and 
Fenestella crassa. Since then a wider acquaintance with the Car- 
boniferous Polyzoa, and more especially with the Irish species in my 
possession, has caused me to hesitate about including it among the 
Fenestelle, since I have good reason for believing that the frag- 
ment which has been described as such will be ultimately found 
to belong to another fenestrate genus of Polyzoa. The original 
drawing of this species by Phillips clearly included two species of 
Polyzoa—one a Polypora with three rows, and the other apparently 
a Henestella with two rows of pores—the latter from Ireland, be it 
remembered, where in certain localities Jchthyorhachis Newenhamu, 
M‘Coy, and Glauconome grandis, M‘Coy, are not uncommon. 

Specimens in my possession lead me to say that I have little 
doubt of being able to show that the fenestrated form described as 
Frenestella crassa, M‘Coy, is likely to prove to be the network or 
polyzoary of one or other of the above species, or some kindred 
form. 

A glance at the drawings of Fenestella crassa given by Prof. M‘Coy 
would seem to confirm this view. The coarseness of the interstice, 
and irregularity of the dissepiment and growth generally, are not 
characteristic of the Fenestellide. Its true affinities have yet to be 
ascertained. or the present it is enough to say that its claims to 
be considered a Fenestella are very doubtful. 


THE CARBONIFEROUS FENESTELLID ZA. 187 


FrnESTELLA HALKINENSIS, Sp. Nov. 

Sp. char.—Polyzoarium a flat, oval, or circular expansion, folia- 
ceous in its early growth, having a stem and expanded root-base. 
Interstices flat, broad, very slightly rounded, keeled, the sides often 
showing a flap or fringe. Dissepiments irregularly placed, very 
thin in early stage, gradually thickening, rarely more than one 
third of the thickness of the interstice. enestrules elongated, 
becoming oval with increased growth, four in the space of two lines 
measured longitudinally, and six in the same space transversely. 
Pores small, round, often twice their diameter apart, three in the 
length of a fenestrule, four and sometimes five in the later and 
larger growth. cel a narrow round wavy line, having three 
nodes in the length of a fenestrule. 

This species at first sight would seem to have a strong resem- 
blance to Fenestella plebeia, which, however, is soon dispelled by 
careful scrutiny ; for it will be found to possess scarcely a feature 
incommon. Its broad flat interstices at once distinguish it from 
the round or more often angular stem of J”. plebera ; while the greater 
distance between the cell-pores, and the thread-lke nature of the 
keel, complete the distinction. Again, the growth of /. plebeva is 
noticeable for its regularity and smoothness, while 7. halkiensis is 
by comparison coarse and irregular. The root-base of F. plebeia is 
secured in position by numerous rootlets; while /. halkinensis has 
a broad, expansive and adherent base, which is further strengthened 
by the interstices which, in favourable positions, directly adhere to 
the rock by a similar flat calcareous base. Its expansion was not only 
much smaller in size than that of F. plebeia, but the last details would 
seem to indicate a species of dissimilar habits, by attaching itself 
to rock-surfaces which the rootlets of /. plebeia would fail to enter. 
The affinities of this new species are, on the other hand, clearly with 
Fenestella polyporata, Phill., rather than with F. plebeia. It has 
very much the appearance of being a diminutive variety of enes- 
tella polyporata, especially on the obverse face. 

Locality. Fairly abundant in the Cement-stone Quarries near Pen 
yr Wylfa, Halkin Mountain, North Wales. 

It will be noticed that I have described only five species of 
Carboniferous Fenestelle. I by no means wish it to be understood 
that I do not believe in the existence of species other than those I 
have described. Of these F. plebeia, M‘Coy, F. nodulosa, Phill., 
F, polyporata, Phill., and F. membranacea, Phill., are really the 
principal and prevailing forms, and will be found in more or less 
abundance in association with other Polyzoa, from the Calciferous 
Sandstones of Scotland to the topmost beds of the Mountain- 
Limestone series of North Wales and elsewhere. The only partial 
exception to the rule is Fenestella membranacea, Phill., which is 
either absent or not so abundant at certain points and localities. 
Having proved the existence and association of these species at 
various horizons in the Carboniferous series, I speak confidently as 
to the fact that the leading species of the Carboniferous Fenestelle 
are few in number, while the reputed species are made up from the 


188 G. W. SHRUBSOLE—FURTHER NOTES ON 


mutilated and altered fragments of the polyzoarium in all stages of 
its growth, not only of Fenestella, but of Ptylopora, Glauconome, 
and Ichihyorhachis, all of which presented a reticulated character, 
which under certain conditions might have been (and, indeed, has 
been) mistaken for that of Fenestella. Hitherto it has been the practice 
to assign any free fenestrate fragment of a polyzoan to Wenestella. 
With the discovery to which I have alluded, that Fenesteila was not 
singular in possessing a fenestrate polyzoary, the error of that 
method of determining polyzoan fragments becomes apparent. 

The few species to which I have reduced the British Carboniferous 
Fenestellide is, I find, in perfect. agreement with the results arrived 
at on the American continent. Prout has published * a list of eight 
species of /enestella from the Carboniferous rocks of North America ; 
although no plates are given, the details of the species are so minute 
as to leave nothing to be desired in the way of description. Of 
these eight species there is only one (Lenesiella Norwoodiana) that 
has any claim to be considered a new species ; they correspond exactly 
with the types of our various English species. Prout did his work 
without the aid of Prof. M‘Coy’s book on the Carboniferous Fossils 
of Ireland, in which the Fenestellide are for the first time 
adequately described. The result of Prout’s independent research, 
apparently with ample material, is that only five good species are 
made out. These American synonyms I have placed for the first 
time under their respective English species. 

There are possibly a local species or two of Fenestella which I 
have not described. Ltnestella halkinensis is one of these local 
forms. I have not met with it elsewhere than at Halkin Mountain. 
These local species require to be very carefully worked out before 
they can be definitely pronounced to belong to the Fenestellide. I 
would suggest, with a view to the prevention of a needless mul- 
tiplication of species of Fenestella in the future, that, before descri- 
bing a new species, the nature of the attachment of the fenestration 
should be definitely ascertained, whether to a root-stalk with root- 
lets, as in Fenestella, or to a midrib or stem, as in Péylopora and 
Glauconome. Nor do I consider this standard too high. I have 
adopted it with the five species which I have described. Owing to 
the complicated surroundings which I have shown to be connected 
with Henestella and the forms allied to it, I am strongly of opinion 
that some such course of procedure is necessary. 

The following table of the measurements of the leading features in 
the several Fenestelle will be found of considerable service in es- 
tablishing their identity. 


* Trans. Acad. St. Louis, vol. i. p. 228-286. 


THE CARBONIFEROUS FENESTELLIDA, 189 


N ie 


NT | Number 
ne mee of fenes- | of fenes- 
aaa gees aren Shape of | trulesin. | trules in 
co Lissep;. | fenestrule.| two lines | two lines | 
Seer : trans- | longitu- 
; versely. | dinally. 
| 
| Fenestella plebeia, M‘Ooy..., 3-4 Oval. 6 ce 
membranacea, Phill...., 3-4 Opreng, 8 | : 
nodulosa, Phill.......... | 3-4 | Square. | 6-8 noes 
polyporata, Pile 6-9") | | Elongate. 4 | 2 
halkinensis, Shrubsole| 3-4 | Elongate. | 6 ay 4 | 
Discussion. 


The Presrpent stated that this group, which ranged from the 
Llandeilo to the Permian, is often represented by very imperfect 
specimens, and that there is great difficulty in discriminating the 
forms. Nowhere are they so well preserved as in the Carboniferous 
rocks of Flintshire and Scotland. 

Dr. Mucrre spoke of the value of the method adopted by the 
author in studying the varieties of growth and development in each 
species. 

Prof. Snetey regarded the principle on which the author had 
worked as a very sound one, and he thought that such revisions of 
certain life-groups were calculated to be of the greatest service to 
geology. 

The PrestpentT remarked upon the difliculty that arises from the 
fact that many type specimens are inaccessible for reference. He 
deprecated the creation of new specific names before a rigid com- 
parison had been made with the old ones. 


190 PROF. P. M. DUNCAN ON THE 


14. On the Corarttirerovs Suriss of Srnp, and ets Connexion with 
the last Upuuavat of the Himatayas. By Prof. P. Marty 
Duncan, M.B. Lond., F.R.S., F.L.8., &. (Read February 2, 
1881.) 

ConrTENTS. 
I. Introduction: the History of the Geology of Sind; Questions involved. 
II. The Stratigraphical Position of the Series and of the Ossiferous Manchhar 
deposits. 

III. General Results regarding the Alliances and Peculiarities of the Corals of 
the Series. The Prenummulitic, Nummulitic, Oligocene, and Miocene 
Coral-faunas of Sind. 

IV. The Equivalence of the Mancbhar and Sivalik deposits. 

V. General Considerations regarding the Age of the last Himalayan Uplift. 


I. Introduction §e. 


A memoir by Grant, illustrated by Sowerby, which appeared in the 
‘Transactions’ of the Geological Society (series 2, vol: v. 1837), first 
brought the countries west and east of the Indus under the notice 
of European geologists. Fossils from Sind, Baluchistan, and Cutch 
were therein described ; and the types were presented to the Society. 
But it is to Mr. Vicary that science owes the first attempt at a com- 
plete description of the geology of Sind*. Written in 1847, the 
fossils which should have illustrated his paper, and some others 
collected by Lieut. Blagrove in Cutch, were handed over to 
MM. d’Archiac and Jules Haime for examination and publica- 
tion. Their fine work, the ‘ Description des Animaux Fossiles du 
groupe Nummulitique de Inde, was published in 1852. They only 
recognized one geological horizon, the Nummulitic, although Grant 


had expressed an opinion that there was more than one fossili- — 


ferous series. Messrs. Cook and Carter added to the knowledge of 
the Baluchistan area in 1860; and the last-named naturalist felt 
it necessary to give a Miocene age to some fossils which Sowerby 
had figured for Grant. On the other hand, MM. d’Archiac and 
Jules Haime severely criticised M. d’Orbigny for stating that some 
of their Sindian species were of Falunian age, and decided against 
Dr. Carter’s grouping of some of the marine Tertiary beds as 
Miocene x. 

In 1863 Mr. Henry M. Jenkins, F.G.S., at that time Assistant- 
Secretary of this Society, and myself were endeavouring to learn 


something about Tertiary deposits situated as remotely as possible 


from European types. 

A collection of Mollusca and Corals from Java had been sent to 
the Society by M. de Groot; and we proceeded to examine them; 
and in order to determine the affinities of some, which seemed to be of 
younger age than the Nummulitic, it became necessary to study the 
work of MM.d’Archiac and Haime on India and to examine their types. 

We found that there was a species in Java which my friend called 

* Quart. Journ. Geol. Soc. 1847, vol. ui. p. 334. 

t Cook, Trans. Med. Phys. Soc. Bombay, vol. vi. pp. 1-45; Carter, Journ. 
Lombay Royal Asiatic Society, vol. vi. p. 184. 

t See Carter, Geol. Papers on Western India, pp. 628-776, and the general 
résumé in their work already noticed. 


CORALLIFEROUS SERIES OF SIND. 191 


Vicarye callosa, and that it was closely allied (the distinctions being 
only of ornament) to Vicarya Vernewlit, D’Arch., from Sind. 

This Vicarya of Java was associated with species which are closely 
allied to those of the Miocene of Western and Eastern Kurope. 

A Coral (Heliastrea Herklotzi, Dunc.) from Java resembled the 
Miocene Heliastreeans more than those of the Kocene*. 

Following up the subject carefully, 1 examined all the collections 
submitted to MM. d’Archiac and Jules Haime; and to my surprise I 
found that they had neglected many specimens which had not only 
a Miocene but even a Pliocene facies. The identity of some species 
with members of the West-Indian Miocene Coral-fauna was tolerably 
evident; and whilst admitting the similarity of the mineral con- 
dition of all the fossils, I divided the genera into those which else- 
where are found in Hocene, Miocene, and Pliocene formations. 

During the following yeart I examined and described twenty- 
six species of Corals from Sind, from unknown geological horizons 
near Karachi. Some had been previously described by D’Archiac 
and Haime; but the bulk had not. ‘he evidence of the existence of 
three Tertiary deposits instead of one seemed overwhelming. 

Subsequently the Geological Surveyors of India, Mr. W. T. 
Blaniord, F.R.S., and Mr. Fedden, made an elaborate survey of 
Upper and Lower Sind, and Messrs. Wynne and Fedden of Cutch.: 

In the last-mentioned district fossils were found which proved 
the equivalency of the deposits with those of Sind, and that there 
was a higher Tertiary horizon than the Nummulitic. 

The Corals collected in Sind by the Survey, under the superinten- 
dence of Messrs. Blanford and Fedden, and carefully marked with 
figures denoting their geological horizons, were sent to me by Mr. 
Medlicott, F.R.S., the present Superintendent of the Survey, so that 
they might be described in a volume of the ‘ Paleeontologia Indica.’ 

But simple description was not all the requirement of the Survey 
and myself. Certain questions of great importance had arisen in 
the geolcgy of India; and it was probable that the determination of 
the age of the coralliferous strata would assist in deciding the ages 
of the Himalayan upheaval, of the Sivdlik deposits, and of certain 
olive shales underlying a trap at the base of the Nummulitic series— 
questions which had been attempted and had been answered by 
Messrs. Medlicott, Blanford, and Lydekker, and which required 
some confirmatory evidence. 

There was another reason why I should be honoured by the Geo- 
logical Survey of India asking me to undertake the description of 
their splendid collection of fossil Corals from Sind. In my Presi- 
dential Address for the year 1878 I criticised the opinions of my 
friends Messrs. Medlicott and Blanford regarding the age of the 
Himalayas, and inferred that their decision regarding the Post- 
pliocene date of the so-called Nerbudda fauna, was influenced by 
the discovery of a human implement in the containing deposit. 

They took the very philosophical course of affording me the means 
of converting myself to their opinions. 


* Quart. Journ. Geol. Soe. vol. xx. p. 45. 
t Ann. & Mag. Nat. Hist., April 1864 (Fossil Corals from Sind). 


192 PROF, P. M. DUNCAN ON THE 


J 
The description of the Corals has been published in a volume of 
the ‘ Paleontologia Indica ;’ and although the evidence regarding a 
Pliocene marine fauna failed, there is no doubt about the former ex- 
istence of Lower and Middle Tertiary coralliferous deposits in Sind. 


Il. The Stratigraphical Position of the Series and of the Ossiferous 
Manchhar Deposits. 

The detailed description of the geology of Western Sind is to be 
found in the Memoirs of the Geological Survey of India, vol. xvii. 
part 1 (1879), by W. T. Blanford, F.R.S. &e., Deputy Superintendent 
of the Geological Survey of India; andit is therefore only necessary - 
to explain those parts of it which relate to the general succession of the 
stratigraphical series and the position of the strata yielding Corals. 

The mountain-ranges west of the Indus run nearly north and 
south, in irregular parallel series. The longest range, the Khirthar, 
is slightly curved, the concavity being to the east; and it extends 
from slightly south of 26° N. lat. to close to 28° N. lat. The Laki 
range, more to the east, is nearly coincident with the sixty-eighth 
parallel of east longitude, and extends from the Indus north of 
26° N. lat. to nearly a degree to the south. Other minor but 
nearly north-and-south ranges occur; and the whole were com- 
prised by MM. d’Archiac and Haime under the title of the Hala 
range (a name unknown to the natives). 

The following is the list of geological formations in Western 
Sind, and which are to be recognized in one or other of the moun- 
tain-ranges * :— 


| amantell | 
Group. Subdivisions. pe Age. Remarks. 


feet. 
ALYUN EA RG oe |eae eeleneeeene ? Post-Tertiary. 
i (Uipperyic sce OOO shacsonae Plioceneteeee Unfossiliferous. 
MESSCHEAB 00. { mae eno 3000-5000 ...| Old Pliocene or} Vertebrate remains. 
Upper Miocene. 
GAS PRR SEER WDC ER RE at 1000-1500 ...| Miocene ............ Coralliferous; no 
Nummulites. 
| Wiper meeeeacs 4000-6000 | scl brad epee eee Unfossiliferous. 
NARI wecrccree Oligocene. ( Coralliferous, with 
| Tower:.......: 100-1500 | VaR ile nnn game Nummulites garan- 
Sensis. 
{ Wipperyeeees.: 500-3000...| Nummulitic ...... Nummulitie lime- 
KHIRTHAR .. stone. 
I OWers acces GOOO'? a..36 sacle erence Unfossiliferous. 
IRANIKOT cuca digester cee D000 eae Lower Nummu-| Fossiliferous. 
litic. Corals and Nummu- 
lites. 
RAP ys so AO | gr Mea Rr Cana AD-90 ewesea oe Decean Trap. 
(| Cardita Beau-\ 350-450 ...... Transition beds ...| Fossiliferous. 
|| sandstones ..| 700 © Hori t det 
andstonesi.|) (OO meesccnecs si. retaceous. orizon not deter- 
ee racrous 1 Limestones | 320. mined. 
|| with Hip- 
(| purites. 


* From the Memoir by Blanford, p. 32, slightly modified. 


CORALLIFEROUS SERIES OF SIND. 193 


Explanatory Sections.—In the Laki range, south-west of Amri 
on the Indus, are dark-coloured hills which contrast with the cliffs 
of grey and white Nummulitic limestone behind them. A section 
close to the hill called Barrah is given by W. T. Blanford, F.R.S. ; 
and it shows that the range consists of three parallel ridges (see fig. 1, 
p- 194). 

The outer, to the east, is composed of Tertiary rocks, while the 
intermediate one consists of Cretaceous beds faulted to the eastward 
against the Lower Eocene strata, and dipping under them io the 
westward. 

This section shows the normal sequence of the groups of strata 
from the Cretaceous to the Khirthar inclusive. Above the Nummu- 
litic limestone of this last group the Nari and Gaj series are wanting, 
and the Manchhar succeeds*. 

On the Gaj river, in the Khirthar range, W. T. Blanford remarks 
that a thickness of at least 25,000 feet of strata is exposed (see 
fig. 2,p.194). The succession from west to east 1s :—unfossiliferous 
strata, probably of Cretaceous age, followed by Khirthar strata (the 
Ranikot series, the lowest Eocene, being absent); then the lower 
and upper Nari series come in, and are followed by the Gaj and 
the Manchhar deposits. 

The lowest coralliferous deposits occur in the soft olive shales and 
sandstones with volcanic ash, belonging to the Cardita Beaumonti 
Series below the trap. The accompanying remains are those of 
Amphiccelian Crocodilia and Echinodermata; and the deposit was 
neither a reef-structure nor a deep-water one. 

The lower part of the Ranikot series, resting immediately on the 
trap, consists of soft sandstones, shales, clays with gypsum and lignite, 
and pyritous shale. A few fragments of bones and some dicoty- 
ledonous leaves occur. ‘These freshwater strata are succeeded by 
highly fossiliferous marine limestones, often brown in colour, inter- 
stratified with sandstones, shales, clays, and ferruginous bands. 

Nummulites appear for the first time, and there is a grand develop- 
ment of Corals, Echinodermata, Gasteropoda, and Cephalopoda. It 
was not a very deep-water formation. Hrosion of the surface of the 
Ranikot strata occurred before the deposition of the next series. 

The Khirthar series includes in its highest portion a massive, pale 
or dark grey, hard, compact Nummulitic limestone whose extreme 
thickness is 3000 feet. It thins out to the south-west, and disap- 
pears within a distance of twenty-five miles of its greatest develop- 
ment. Other Nummulitic limestone-beds are found, which may be 
lower in the series ; and they and the main group are represented 
elsewhere by shaly limestones and sandstones with calcareous bands. 
In some districts flint occurs in a limestone with Alveolinw. The 
lower members of the series are often wanting, and are well repre- 
sented by shales, marls, and sandstones and where these are present ; 
unconformity with the underlying .Ranikot beds is not seen; but 
where they are absent the Nummulitic limestone (as in the Laki 
range) rests unconformably. The compact limestone is of course 

x Memoirs Geol, Survey of India, vol. xvii. pt. 1, p. 151. 


PROF. P. M. DUNCAN ON THE 


194. 


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CORALLIFEROUS SERIES OF SIND. 195 


highly fossiliferous ; but it has not yielded very good specimens of 
Corals. The indications of a fringing-reef-building fauna, or a bank 
of coral, are distinct; but the species when compared with their 
modern analogues do not indicate a great reef-development. 
Amongst other fossils, Lamellibranchiata prevail, and the Echini are 
numerous; but the Nummulites and Orbitolites, Alveoline, and 
Patelline are the most important organic remains. Amongst the 
Nummulites there are WV. Ramondi, N. biaritzensis, N. Beaumonin, 
NV. granulosus, and N. Leymerier. 

The Nari series rests conformably on the Khirthars; and there is 
occasionally an apparent passage from the Khirthar limestone into a 
yellow or brown rock of the Nari group. There is, however, a 
biological break; for the Nummulites of the upper group are 
characteristic, and differ from those of the lower. The Khirthar 
forms are not found, and Nummulites garansensis appears with Orbi- 
toides papyracea, in the Nari series. The rock is a limestone with 
intercalations of sandstones and shales. Asa rule, shales, fine sand- 
stones, and occasional bands of limestone form the base of the Nari 
group, and pass upwards into coarse, massive, thick-bedded sand- 
stones, attaining a thickness of from 4000 to 5000 feet on the eastern 
flank of the Khirthar range. 

A local break occurs to the south, and the Upper Nari beds rest 
unconformably on the denuded edges of the Lower Nari brown 
limestones; and still further south, fifty miles east of Karachi, there 
is a well-marked distinction between the upper members of the 
group with Orbitoides papyracea and the lower with Nummulites 
garansensis; and in one locality the upper member overlaps the 
lower, and rests on Khirthar limestone. 

To the east of the Laki range the Nari beds are wanting, and the 
Mancbhar series rests unconformably on the Khirthar, with some 
pebble-beds of the Gaj series intervening. But to the west of the 
range the Nari and the Gaj series are found in their normal sequence ; 
and towards the coast the exact distinction which can be drawn 
elsewhere, stratigraphically and petrologically, between the Ter- 
tiary series is not possible: this is mainly due to the disappear- 
ance of the limestone element of the Khirthar and Nari series, and 
to the prevalence of sandstones and shales. The fossils, however, 
distinguish the groups: but the horizons of the zones of Nummu- 
lites and Orbitoides vary, being higher or lower in their proper series 
according to locality. 

The upper sandstones of the Nari group have not yielded marine 
fossils, and in Upper Sind they contain the remains of plants. 

The Nummulites become fewer in their species at the base of the 
Nari group, and cease to be found in the lower beds of the suc- 
ceeding series, the Gaj. 

The Gaj group, with a base of highly fossiliferous limestones and 
calcareous beds, more or less shaly and stratified, overlies the softer 
shales and sandstones of the Nari series. The development of the 
calcareous series is great; but it is subordinate to an arenaceous 
element. ‘The sandstones are intercalated with clays with gypsum; 


196 PROF, P. M. DUNCAN ON THE 


and shales and bands of limestone, highly coralliferous, are very 
constant. 

The Gj series rests conformably on the Nari series; although 
there is a mineralogical break, the passage is so oradual that cu 
careous bands of the Gaj series are found interstratified with the 
uppermost Nari sandstone. The Gaj series overlaps the Nari to the 
south and rests on the Khirthars, and is wanting in Lower Sind to 
the eastward of the Laki range 

North-east of Karachi the series is highly developed, and massive 
limestones occur ; they reach as far as the coast. 

The uppermost beds of the Guaj series are variegated clays and 
grey sandstones, which form a passage into the everlymg Manchhar 
strata, and contain species of Ostrea, Corbula, Arca, Scalaria, 
Buccmum, and Turritella. A crab of the genus Typilobus and the 
Vicarya Vernewliw already noticed have also been found there. 
The oyster is like Ostrea multicostata. 'The Echinoderms are of the 
genera Maretia, Meoma, Breynia, Clypeaster, &e. The Corals found, 
principally associated with the lmestone-beds, are very different in 
their aspect from those of the underlying Nari group. They belong 
for the most part to reef-building genera, and some are represented 
in modern reefs by allied species, which grow in the surf and in the 
most exposed parts. ‘The rest were dwellers ; in quiet water. Great 
masses of the coral limestone consist of Stephanocena maxima in 

casts, and the presence of the genera Madrepora, Hahastrea, and 
Porities is very significant of s shallow-water conditions. 

Resting on the Gaj strata is the Manchhar group; and where the 
Gaj beds are wanting, and even where the Nari series is deficient, 
this upper group rests on the Khirthars. 

Of ereat thickness (10,000 feet on the flanks of the Khirthar range) 
the Manchhar series is divisible into an upper and a lower group. In 
the lower group much grey sandstone, soft and fine-grained, and 
composed of quartz with some felspar and hornblende, is found ; and 
red sandstones and conglomeratic beds exist towards the base, as 

well as red, brown, and grey clays. The conglomerates do not 
contain pebbles of older Tertiary rocks; but cream-coloured clay, 
soft sandstone, quartzite, and micaceous shale are found in them. 
The conglomerates near the base are ossiferous; and the vertebrate 
remains are teeth or bones more or less rolled. There is no satis- 
factory distinction to be made on the Gaj river between the estua- 
rine beds at the top of the Gilj series a the lowest beds of the 
Manchhar; and osseous remains have been found in the upper Gaj 
series. 

The upper part of the Manchhars has a conglomerate in it with 
stones derived from the older tertiaries ; below it are red, brown, or 
buff sandstones, with some clays; and there ure no fossils. 

Although usually conformable to the Gaj series, in places the 
Manchhars rest on the older rocks ; ; and there are proofs of the Gaj 
strata having been greatly denuded before the deposition of the 
Manchhars. 

In the Laki range the Manchhars have an ossiferous conglo- 


CORALLIFEROUS SER1ES OF SIND. 197 


merate near the base. In the Vero plain, running southward along 
the eastern side of the Laki range, south of Ranikot, large frag- 
ments of silicified wood are common, and some trunks of trees are 
upwards of 30 feet in length and 10 in girth. In the south, near 
Karachi, the Manchhars pass into Gaj strata, and marine fossils are 
associated with the lowest beds. 

The general conformity of this great tertiary series is evident— 
although local unconformities occur, and there is evidence ‘in 
fayour of there having been some disturbance of the older rocks 
before the deposition of the Lower Manchhar group. The sub- 
sidence which took place during the deposition of these thick shallow- 
water deposits was vast; and there were occasional slight upheavals. 
The final epoch of the mountain-formation occurred after the depo- 
sition of the Manchhars. The strike of the chains is in the main 
north and south; and the thrust came from the west to the east, and 
from east to west. 

The date of the grand mountain-formation is subsequent to the 
deposition of the sedimentary strata forming the Upper Manchhars. 


III. General Results of the Study of the Corals of the Series. 


The Corals of the strata with Cardita Beaumonti below the trap. 
are shallow-water forms; and their development indicates conditions 
untavourable to vigorous coral-growth. There was no reef. The 
species are new ; and there are no characteristic Cretaceous or Hocene 
forms present. The Smelotroch: and the Lhabdophyllic are Secondary 
genera; but the first-named had species inthe European Tertiaries, and: 
may be represented in the shallow seas of the present day. The Cary- 
ophyllice of the deposit have four cycles of septa only ; and this gives. 
them somewhat an ancient look ; and the Litharwa would pass either 
as a Cretaceous or as an Eocene form. 

The fauna as a whole is deficient in characteristic forms, and may 
be considered transitional. 

It has been stated by the Geological Surveyors that the trap re- 
sembles the Deccan and Malwa trap; and the position of that vast 
outburst is anterior to the Nummulitic age, and subsequent to some 
Cretaceous strata in Western Central India. Certainly there are no- 
Nummulites in the sands and shales which contain the fessils; and 
these are the remains of a Cretaceous crustacean. 

The Corals of the Ranikot group are numerous in genera and 
species ; and the fauna is remarkable for the number of simple forms: 
it contains, and for the predominance of the family Fungide. 

The compound Corals of the family Astraide, which so largely 
enter and have entered into the composition of reefs, are but feebly 
represented ; and the few species which have been described did not 
ageregate to form massive limestones, but were seated on small flat 
circular epithecate bases. Fine as were many of the growths, yet 
it was a stunted Coral-fauna; and the shape of the majority of the 
forms would rather indicate that they lived in still water, and not 
in the rush of the waves. Certainly the Corals were beyond the 


Q.J.G.8. No. 146. P 


198 PROF. P. M. DUNCAN ON THE 


rolling or scouring action of the sea; for any such movement would 
have displaced most of them. 

As a generic assemblage, the Ranikot Corals are Kocene in facies; 
and of the twenty-seven genera three are new, and oneis known in the 
Miocene of the West Indies. Out of fifty species seven are identical 
with European Eocene types, which are not found, however, as a 
whole, on one geological horizon in Europe. Some of them are 
found in the Lower Eocene; and the others are to be noticed in the 
deposits of San Giovanni Ilarione and of the Oberburg, in Styria, and 
in the higher horizon of Castel Gomberto. These forms are located 
in one series in Sind, and at the bottom of the Nummulitic group. 
Five species are closely allied to Kuropean forms out of the same great 
vertical series; and about seven others have a distinctly European 
facies. These 19 species give the Eocene charaeter to the fauna. 

The most distinctive genera are Stylocwinia, Plocophyllia, Stepha- 
nophyllia, Stephanocenia, Prronastrea, Reussastreea, Cyclolites, and 
Litharea. It is remarkable that five well-marked species of the 
genus Turbinoseris (nobis) should have lived on the Indian area; 
they are allied to those of the Eocene of St. Bartholomew in the 
West Indies.* ‘The genus is Lower Cretaceous in England ; and, under 
another name, it has lately been introduced into the N attheim 
oolitic Coral- fauna. 

The presence of a species of St yline and of Thamnastrea recalls 
the Jurassic and Cretaceous ages: ‘the first is found also in the next 
or Khirthar series of deposits; and'thé last probably lived longer in 
the southern seas. Only one genus, Placocyathus, is a Miocene type ; 
and its distribution is West-Indian. 

There is one very remarkable species, Stylocenia maxima, nobis, 
in the Ranikot group; and it is so called from the great size of the 
intercalicular projections, which are little monticules in the British 
Stylocenia emarciata, Lamk. sp. ae emeniun of the Cages base 
of the coral is 3} inches; an A 
concentric epitheca is on the base; and the upper surface i is @ mass 
of calices, and tall columns between them, some reaching 75 inch in 
height. The minute columns in the British | ‘species are proved by 
the examination of these gigantic growths to be stunted or aborted 
calices. Some of the processes of Btylocoenia maxima have a perfect 
calice on the top, and the costs come down the side; others have ill- 
developed calices, and a mere representation of a columella and septa. 
It is acommon form, and is characteristic of ‘the Ranikot series; 
its nearest ally is Stylocania macrostyla, Reuss, which is associated 
with Nummultes planulatus and Cerithium ¢ Uh iganteum in the district 
of San Giovanni ilarione. 

Young specimens, of course, greatly resemble the European Stylo- 
coenia emarciata of Bracklesham age. 

‘Two other species of Stylocoenia are very common: one, S. Vecaryt, 
was described by D’Archiac and Haime ; and the other is new. It has 
a large circular epithecate base ; and the calices are large. 

Astr ocoenta, that very widely distributed genus, whose range, ver- 

* P. M. Duncan, Quart. Journ. Geol. Soc. vol. xxix. p. 554, 


CORALLIFEROUS SERIES OF SIND. 199 


tical and horizontal, is enormous, has several species in the Ranikot 
eroup: and Astrocenia ramosa, Sowerby, a Cretaceous type from the 
Turonian of Gosau, found also in the Hocene of St. Bartholomew 
in the West Indies, is present in Sind, and has the usual tuberose 
branchlets so characteristic of the form. 

Large solitary Corals of the genus Montliwvaltia abound, and also 
a characteristic series of simple forms which I have had the oppor- 
tunity of naming after Mr. Fedden, to whom the geology of Sind 
owes so much. The genus Meddenia had its young forms growing 
on small shells, which, with the growth of the base of the Coral, 
were enclosed, and the Coral became free. This is not an unusual 
habit in some two or three species of Corals of another family, 
now living; and an elongate base is produced. One of the 
results of this change of condition was that the coste did not 
radiate upwards from the peduncle, as is seen in the larger forms, 
but from the whole length of the enclosing base. Another pecu- 
harity is that the epitheca is like a broken mosaic. 

There are three well-marked species of this genus; and one has 
two good varieties in the Ranikot series. It is near Montlivaltia as 
a genus; and, admitting the great variability of all simple or 
solitary Corals, it is very possible that the unusual method of 
early growth may have produced correlative changes in the general 
shape and septal arrangement. The genus was restricted to this 
Lower Eocene horizon. Amongst the great series of Fungide found 
in the Ranikot group there are two European forms of Cyclolites ; 
and the other seven species of the genus are well differentiated. 

Finally, the genus Stephanophylliia, amongst the Kupsammines, 
ought to be represented, to give the full early Tertiary facies to 
the Cyclolitic assemblage. Stephanophylla indica, from Jhirk, in 
the Ranikot group, maintains the character of the genus for beauty, 
and is an exquisite gem. Its affinities are rather with the Cre- 
taceous species; and it differs but slightly from S. Bowerbanki of 
the Lower Chalk in its main characters, and from S. discoides of 
the London Clay more decidedly. Like most of the Ranikot corals, 
the individuals of the new species began life by settling down on a ~ 
Nummulite; this formed the permanent base of the coral; and the 
usual radiating costal arrangement of Stephanophylha either does 
not exist or cannot be seen for the Nummulite. 

Khirthar Serives.—Sixteen species of Corals were found in this 
series; but ten of them were taken from beds so high up that it 
is not satisfactorily proved whether they are at the top of the 
Khirthars, or form the base of the Nari group, next in vertical 
succession. By separating these ten species, an indubitable 
Lower-Khirthar fauna is decided to have existed: its biological 
conditions were unfavourable to vigorous coral-growth ; and there 
were no littoral or reef-building forms. From the analogy of 
recent forms, which are identical generically with those of the 
Khirthar series, or whose shape resembles that of the ancient species, 
a sea-floor at the depth of from 20 to 200 fathoms or more would 
be indicated. 

RP 2 


200 PROF. P. M. DUNCAN ON THE 


The species belong to the genera TZ'rochocyathus, Leptocyathus, 
Stylophora, Montlivaliia, Calamophyllia, and Astrocema. The 
Trochocyathus is one of the discoid group, and the Leptocyathus is 
of course a low flat Coral. The Stylophora is a minute Coral; and 
so is the Astrocenia. But the Montlvaliia is a finely grown form ; 
and so is the Calamophyllia. Of these, Stylophora contorta, 
Leymerie, has also been found in the Kocene of the West indies and 
at La Palareain Europe; and Astrocenia nunusma, Defrance, sp., is 
a Nummulitic form at Gap, in the district of Nice. This little 
assemblage is thus clearly of Hocene age. No distinction can be 
made between this and the preceding deposit, as regards age, from the 
Corals. ; 

Now the ten species, the horizon of which may be of Upper 
Khirthar or Lower Nari age, belong to genera whose morphology in- 
dicates the presence of totally different physical conditions from those 
which environed the stunted, simple, and the few large Corals of the 
lower horizon. They were shallow-water and reef-building forms ; 
and the genera are Stylina, Latimceandra, Hydnophora, Favia, 
Isastrea, Pterastrea, Plesiastrea, and Porites. Thereis nota simple 
Coral amongst them ; and the facies is singularly mixed, old and new, 
Mesozoic and Cainozoic genera existing together. 

The species Hydnophora malirrensis, nobis, is allied to H. venusta, 
Catullo, of the European Nummulitic. Porites Pellegrinit, D’Achiardi, 
is found in Kurope at San Giovanni Llarione; and Porites indicus, nobis, 
is allied to a species from Crosara, Oberburg. The distinctness of 
this fauna from that of the Ranikot group is evident enough ; but 
it is still Nummulitic in facies and character. 

The Narv Series—From the remarks already made upon the 
localities of some of the presumed Khirthar Corals, it is as well to 
admit that some upheaval took place at the close of that period, and 
produced fringing-reef-building or bank conditions. But they did 
not prevail throughout the accumulation of the whole of the Nari 
Series; and it is somewhat remarkable that no species passes up 
from the reef-building horizon into that of the Nari proper. 

The genera of the Nari group above the base, indicate rather 
vigorous coral-growth, and both deep- and shallow-water conditions. 
The genera Trochocyathus, Trochosmilia, Montlivaltia, Cycloseres, 
and COyclolites are the simple forms; and the vigorous compound 
forms are included in the genera Dasyphyliia, Rhabdophylhia, Lep- 
toria, Meandrina, and Prionastrea. | . 

The Trochocyatht of the Nari series are remarkable for having 
commenced their growth in a discoid shape. Some retained that 
shape; but one in particular, Trochocyathus nariensis, nobis, grew 
upwards from its disk-shaped base in a perfect cylinder, reaching 2 
inches in height. 

Trochocyathus cyclolitoides, Kd. & H., is a widely distributed 
Huropean Eocene coral, and is found in the Nari series of Sind. 

Stylophora pulcherrima, D’Achiardi, from the Hocene of Friuli, 
Trochosmilia varicosa, Reuss, from Crosara, Stylocenia taurinensis, 
Kd. & H., from European Eocene and Miocene strata, and Cycloseris 


CORALLIFEROUS SERIES OF SIND. 201 


Perezi, Ed. & H., from the Nummulitic of Nice, are also amongst the 
Nari corals. Hence, out of the twenty species of fossil Corals in the 
Nari series, one is found at a higher horizon, and one fourth of the 
number are found in the Upper Nummulitic and Oligocene deposits 
of Hurope. 

Taking the Ranikot, Khirthar, and Nari Coral-faunas as a 
whole, there are 76 species of Corals in them and severai varieties ; 
and of these 16 are identical with European forms which are found 
in strata yielding Nummulites to the top of the Oligocene. There 
are also eight species in the Sind series, closely allied to those of the 
European fauna; and whilst some species are common to the West- 
Indian Eocene, an important genus is also common to both localities. 
The absence of species passing up from one series to another is 
very remarkable. 

The Gdj Series——This series of strata contains a large number 
of Corals in bands ; but although most of the forms are massive and 
compound, and suited for reef-building, the majority are peduncu- 
late instead of covering a large surface and incrusting. Some are 
very massive ; and the presence of species of Madrepora, an essentially 
reef-building genus, with Porites, Agaricia, Echinopora, Prionastrea 
Plesiastrea, Brachyphyllia, Leptoria, and Dasyphyllia would 
indicate, at the present time, very active shallow-water coral- 
growth. The majority of the Gaj genera still flourish; and most 
of those which are extinct had the structures requisite for reef- 
buildmg. The Astreide as a family preponderate, and simple 
Corals are rare, in the collection. But although the facies of the 
Gaj coral-fauna is very recent, and there are very few extinct 
genera, still the modern Coral-fauna of the Eastern and Red Seas 
is not represented by a single species. Two species are identical 
with Miocene West-Indian forms; and one is found in the Nari series. 

Tt is interesting to find the West-Indian Miocene and recent genus 
Antillia represented. Nevertheless the evidence afforded by the 
Corals is in favour of a mid-tertiary age being given to the 41 
species from Gaj. 

In the communication to the ‘ Annals and Magazine of Natural 
History’ (1864, vol. xiii.) I described several species of fossil Corals 
from unknown geological horizons in Sind; and it is not difficult, 
with the lists of the genera and species which have been collected 
by the Geological Survey of India, to decide whence the majority 
eame. Thus Oculina halensis, nobis, Antillia dentata, nobis, An- 
tillia ponderosa, Ed. & H., Cladocora haimei, nobis, Hydnophora 
rudis, H. plana, H. hemispherica, nobis, point, with Cyathoseris val- 
mondorisiaca, Kd. & H., and Agaricia agaricites, Ed. & H., to a late 
Miocene horizon. They were picked up in the neighbourhood of 
Karachi, and doubtless came from the highest beds of the Gaj 
series. 

Finally, many specimens of Isidinee have been found in the Gaj 
series; and some of them closely resemble modern forms. No Num- 
mulites occur in the Gaj series ; and the Zoantharian eviderce indi- 
cates a Miocene age, and not an early one. 


202 PROF. P. M. DUNCAN ON THE 


The Sindian formations, including the transitional beds with Car- 
dita Beawmonti beneath the trap, and the Gaj series at the top, 
contain 136 species of Corals and many varieties. Of these, 9 
belong to the transitional series, and 56 to strata in which Nummu- 
lites exist in profusion. The Nari series, with few species of Num- 
mulites and with WV. garansensis, a Kuropean type characteristic of 
the Upper Nummulitic or Oligocene, contains 20 species. The 
Miocene strata of Ga} contain 41 species. I omit from these 
numbers all species from doubtful localities. Thus, as only one 
species transgresses, there are five Coral-faunas ; and if the separa- 
tion I have proposed at the base of the Nari series is right, there are 
six in the same area, included in a prodigious depth of sedimentary 
strata. The study of the Corals does not indicate that there was 
ever a sea there with a depth of many hundred fathoms; on the 
contrary, shallow-water conditions (within 20 fathoms) usually pre- 
vailed during the slow oscillation of the area, in which subsidence on 
the whole predominated. 


IV. The Lquivalence of the Manchhar Strata of Sind and the 
Swalik Group of the Himalayas. 

The Geological Survey of Sind by Blanford and Fedden has proved 
that in some places the Manchhar strata succeed transgressively to 
the Gaj series, whilst in others there has been erosion of the marine 
deposit before the deposition of the freshwater one, or else the 
lower group is absent. Bones of Rhinoceros siwalensis have been 
found in the uppermost Gij strata. There is no doubt that the 
greater part of the coralliferous strata of the Gaj series accumulated 
in shallow water, and yet beyond the reach of the wash-down of a 
coast-line. But the lower Manchhars were deposited in shallow 
water within the scope of terrestrial denudation. There must have 
been considerable general changes in the physical conditions of the 
area: and they persisted ; for coral life has never since prevailed 
there. The first change probably was one of slight general up- 
heaval; and subsequently a slow and progressive subsidence 
occurred, during which the vast vertical development (8000 to 10,000 
fect at least) of the lower and upper Manchhars accumulated. 

Conglomerates, coloured sandstones (grey, green, and red in tint), 
and clays, are the prevailing deposits on the inner flanks of the 
vast. mountain-system which surrounds Peninsular India, from Sind 
to Burma inclusive. Made up of stone brought down by tne rivers 
of the extra- peninsular mountain-system, before it became of very 
ereat height, and when its breadth was probably much greater than 
it is now, these deposits are to be traced to the north of Sind in the . 
Suleiman Mountains, in the Salt range, covering much of the 
surface of the Northern Punjab, and then, forming part of the sub- 
Himalayan range, as far east as the Brahmapootra. Similar 
deposits are found, in diminished thickness, in the Assam range ; and 
they became important in the Burmese territories. The Sindian suc- 
cession of the strata is the normal one; and it extended to a certain 
distance northwards ; moreover it is exemplified in the Burmese 


CORALLIFEROUS SERIES OF SIND. 203 


area. ut in the great central area of the Himalayas proper there 
was no marine Miocene. No marine beds, the equivalents of the 
Gaj series of Sind and Burma, underlie the Sivalik deposits in the 
Sub-Himalayas. 

There was open sea during the Nummulitic age where the great 
semicircle of extra-peninsular mountains now exists ; and marine con- 
ditions persisted on the west and east during the Miocene period ; 
but a post-Nummulitic upheaval developed a land-surface and_hill- 
tracts from Kashmir to Assam. 

This upheaval commenced before, but culminated after, the close 
of the Eocene age; and a considerable thickness of purple sand- 
stones, red clays, and grey and purple sandstones containing plants 
accumulated in the swamps on the plains of that age, near the 
mountains on the edge of the Nummulitic sea-floor. 

These plant-bearing strata and the underlying Nummulite- 
bearing strata have a minimum thickness of 2000 feet, and they 
form the Sirmuir series of Indian geologists. 

This series underlies unconformably the vast freshwater sedi- 
mentary formation comprising the Nahun and Sivalk strata, which 
attain a thickness of about 15,000 feet, and, except where buried 
beneath recent deposits in one locality, extend, with a varying 
development, along the south of the great mountain mass, and are 
found on one of the great tablelands to the north of the Central 
Himalayan axis. The lower, or Néhun unfossiliferous, series con- 
sists mainly of grey lignitiferous sandstones. On it the upper, or 
fossiliferous Sivaliks, accumulated as sandstones and clays; and on 
the top of all are conglomerates. A great fauna is represented in 
all parts of the Sivalik deposits above the Nabun beds. 

From the lie of these sedimentary strata, it may be very reason- 
ably inferred that the Nahun and Sivalik deposits are the equivalents 
of the Lower and Upper Manchhars of Sind; and the inference may 
be extended to the Upper Tertiaries of Burma. There is an outlier 
of this series and of its lower member in the Gulf of Cambay or 
Perim Island. 

In attempting to establish exact parallelism between the Sindian 
and the Himalayan deposits called Manchhar and Sivalik, it must 
be noticed that the vertical development of the last-named rocks is 
the greatest, and that whilst the lowest beds of the Sindian series are 
fossiliferous, those of the Nahun beds of the Sivdliks are not*. On 
the other hand, osseous remains are found throughout the Sivaliks 
proper (above the Ndhun beds), but not in the Upper Manchhars 
in a recognizable form. 

The Manchhar and Sivdlik series have been upheaved, uptilted, 
and in the last instance greatly contorted. Both series were the 
youngest implicated in the great orographical development; and 
although they are on different lines of strike, they were affected 
during the same geological period. 

The denudation of their exposed edges has been great. Both are 


* Possibly the ossiferous deposit at Kushalghar, near Attock, is of Nahun 
age. See further on. 


204 PROF, P. M. DUNCAN ON THE 


covered in some places with high-level gravels; and the Sivaliks of 
the great tableland of Hundes underlie deposits referable to the 
glacial period, whose effects were not felt so far south as Sind. 

In considering the paleontology of these deposit it may be stated 
that the osseous remains are found on several horizons. 

First, the Rhinoceros-remains which were found included in the 
Marine Miocene series of Gaj in Sind, lead to the inference of the 
existence of neighbouring contemporaneous land-surfaces and shallow 
seas—that is to say, of Miocene land washed by a Miocene sea. The 
Rhinoceros, according to Mr. W. T. Blanford, F.R.S., is Rhinoceres 
sivalensis. 

Secondly, the fragmentary bones and teeth found in the conglome- 
rate near the base of the Lower Manchhar formation in Sind accu- 
mulated later on than the Gaj series. 

The following is the list of the vertebrate remains which were 
collected by Mr. Fedden, and named by him and Mr. Lydekker in the 
publications of the Geological Survey of India :— 


Amphicyon paleindicus. Anthracotherium silistrense. 
Dinotherium pentapotamiz. Hyopotomus palzeindicus. 
indicum, Hyotherium sindiense. 
, Sp. nov. Dorcatherium majus. 
Mastodon perimensis. minus. 
latidens. 
Falconeri. EDENTATA. 
Rhinoceros paleindicus. Manis sindiensis. 
, sp. near R. deccanensis. 
Acerotherium perimense. Rerriita. 
Sus hysudricus, Crocodilus, sp. 
Hemimeryx, sp. Chelonia, sp. 
Sivameryx, 2 sp. Ophidia, sp. 


Chalicotherium sivalense. 


The conglomerate containing the bones was composed of the 
wash-down of the Lower Manchhars themselves. 

Thirdly, the fossil bones which have been discovered in consider- 
able quantities in a conglomerate on Perim Island (although their 
place in the geological series cannot be determined, from the absence 
of a succession of rocks) appear to have belonged to a fauna allied 
to that of the Manchhars and to that about to be noticed. 

The species which have been determined by Falconer and Lydekker 
are :— 


Dinotherium indicum. Brahmatherium perimense. 

Mastodon latidens. Camelopardalis sivalensis. 
perimensis. Capra perimensis. 
sivalensis. Antilope, sp. 

Rhinoceros, sp. Sus hysudricus. 


Acerotherium perimense. 


The remains are included in sandstone blocks. 

The same, or a slightly higher, horizon is recognized far away to 
the north, not, however, by its geological position, but by the fossil 
remains, in the neighbourhood of Kushdlghar, forty miles south of 
Attock, in the Punjab. The deposit contained, according to Falconer 
and Lydekker, the following genera and species :—Dznotherwum 


CORALLIFEROUS SERIES OF SIND. 205 


pentapotamic, Mastodon, Listriodon pentapotanie, Rhinoceros, 
Merycopotamus, Dorcatherium, Sanitherium Schlagintweitii, and 
Amphicyon palemdicus. 

It is remarkable that all these horizons should contain Dzno- 
therium. 

Fourthly, the Sivalik clays, sands, and conglomerates above the 
Ndahun series present an assemblage of genera and species second 
to none in importance, and much grander generally than the existing 
fauna, which is very slightly represented. A number of genera 
which had lived in the Miocene elsewhere were associated with 
genera not known in mid-Tertiary deposits, and which have lasted 
either into Post-Pliocene or into Recent times. The Dinotheria are 
absent. Omitting the common Miocene genera and stating the 
others, an African element is noticed, as in the instance of Pikermi 
in Greece. The characteristic genera are Paleopithecus, Macacus, 
Semnopithecus, Stegodon, Elephas, Lowodon, Hippopotamus, Camelo- 
pardalis, Camelus, Bos, Bubalus, Bison, Cervus, Equus, Canis, Ursus, 
Mellivora, Meles, Lutra, Enhydriodon, Tapirus, Hystrix, Mus, Rhizo- 
mys, Crocodilus, Gavialis, Varanus, Ballia, and Emys. 

Moreover many of the freshwater Mollusca are identical with 
recent forms. 

It appears that these wide horizons are separable into an older 
serles, with Dinotherium, Hyotherium, Hemimeryx, Sivameryx, Hyopo- 
tamus, Anthracotherium, Acerotherium, and Manis, but without 
Elephas and the later bovines; and into a younger series, in which 
genera of an African type, such as Hippopotamus and Camelo- 
pardalis, are found, and without the older types. 

The first series is in relation with the Lower Manchhars, and the 
second with the Sivalik deposits above the Nahun beds. But the 
interesting fauna from Kushalghar appears to be probably of Nahun 
age; and if that should be proved eventually, the succession of two 
fairly distinct faunas, linked together by some species, will be 
evident. 

Finally, the Post-Pliocene deposits of the Jumna area, and also 
of the Nerbudda and other peninsular rivers, contain some species of 
the Sivalik horizon. 


V. General Considerations regarding the Age of the last Himalayan 
Upheaval. 


It might be supposed, considering the ready manner in which 
European mammalian faunas are placed in the scale of geological 
succession, that the contemporaneity of some of them with the 
Sivalik-Manchhar assemblage of vertebrates would be easily deter- 
mined, and a geological age given for the latter without doubt. 

But the critical examination of the positions in which many of 
the European mammaliferous deposits have been found, indicates 
that the manner in which the Tertiary deposits have been classified 
by means of the fossil Mammalia is open to exception. 

Two well-known examples of European mammaliferous deposits 
which resemble, in their succession, the Sivalik-Manchhar series are 


206 PROF. P. M. DUNCAN ON THE 


those of Eppelsheim and Heppenheim and of Pikermi. At Eppels- 
heim sands in small patches are at the top; and they contain a few 
specimens of fossil Rodentia, Insectivora, and Carnivora. A con- 
glomerate is beneath, and contains the Dinotherian remains, and 
also those of Rhinoceros and Mastodon; and all are remanic¢. At 
the base of the series there is a clay with freshwater shells; and 
it rests conformably on a marine limestone; and this is of late 
Miocene age. : 

At Pikermi the deposit, the bones from which have been so ably 
described by Gaudry and commented on by W. T. Blanford, rests 
on a freshwater stratum of Phocene age, and a Miocene lacustrine 
series underlies the whole unconformably. 

Now at Sind as at Eppelsheim the underlying marine beds are 
Upper Miocene in age, and freshwater conditions prevailed subse- 
quently, during which the osseous remains were depoisted. 

What is the age given to the Dinotherium-conglomerate at Eppeis- 
heim? Carl Vogt, influenced apparently by the presence of 
Dinotherum, and regardless of any stratigraphical arguments, and 
not considering the important changes which had occurred in the 
area, in the relative level of the land and sea-floor, decided that the 
overlying freshwater beds are of mid-Tertiary age. But Credner, 
dealing with the subject more philosophically, and not being so 
much impressed with the presence of a genus which elsewhere is 
represented in Pliocene strata as by the evidence of the considerable 
mutations which had occurred in the physical geography of the 
district, and which had brought a marine deposit above the original 
sea-level, places the bone-bearing conglomerates in the Pliocene age. 
Thus one geologist associates the land and marine elements together, 
and the other separates them. | 

The comparison which can be instituted between the Manchhar 
and Sivalik deposits and those at Pikermi is very close; and, accord- 
ing to the ordinary rules of stratigraphy, if the osseous remains at 
the last-mentioned locality overlie Pliocene deposits, the animals 
which left their bones could not have been of Miocene age. Yet 
Prof. W. Boyd Dawkins, in his contributions to this Society, asserts 
that the Pikermi fauna flourished in the Miocene. Evidently there 
is a great diversity of opinion regarding the age which should be 
given to land-surface remains overlying marine strata, freshwater 
and estuarine deposits intervening. And if this subject is studied 
it will be found that great discrepancies of opinion have existed in 
regard to similarly placed deposits in many of the great formations. 

About the relative age of some superincumbent terrestrial remains 
there is no discrepancy of opinion. ‘The Coal-measures are associated 
with the underlying grits and limestones; and the land remains of 
the Inferior Oolite are similarly connected in classification with the 
marine deposits beneath them. When the lateral extension of the 
strata can be traced, and distant marine equivalents of the overlying 
series can be proved to contain fossils representative of or identical 
with those of the underlying series, the land-surface is classified with 
the formation in which the marine strata are placed. Or when there is 


CORALLIFEROUS SERIES OF SIND. 207 


a succession, on the same area, of marine beds over the terrestrial 
and freshwater series, if the fossils of the upper strata resemble 
those of the deeply seated marine ones, the whole belongs to one 
great aspect of nature. 

It will be found that there can be no exception made to placing 
the Woolwich and Reading series out of the formation which includes 
the white chalk; but the propriety on any grounds of linking the 
Purbecks on to the marine Portland is open to exception. 

Except im instances similar to those of the Carboniferous and 
Inferior Oolite, and where there is also decided unconformity, the 
question can only be answered after a careful consideration of the 
amount and extent of area implicated in the changes in the physical 
geography which may fairly be assumed to have occurred since the 
underlying marine deposit was completed. The upheaval of limited 
marine deposits recognizable as belonging to a particular formation, 
and the accumulation upon them of freshwater deposits and conglo- 
merates, would hardly necessitate the belief in such a change in the 
aspect of nature as would warrant their classification under two 
great geological ages. But when the same phenomenon is witnessed 
over widely separated areas, and a conglomerate is followed by some 
thousands of feet of fluviatile and other freshwater strata (the 
wreckage of high land close at hand), it becomes certain that the 
physical change has been wide enough to admit of an alteration in 
the geological nomenclature. 

With regard to Sind, the Lower Manchhars usually rest conform- 
ably on the Gaj marine Miocene, and marine, estuarine, and fresh- 
water intercalations exist at their base and before the conglomerate 
is fully developed. In one locality the Manchhars rest on a greatly 
denuded surface of Gaj strata. 

Upheaval (slow, irregular, and on a very grand scale) occurred 
subsequently to the deposition of the Gaj series; and ‘a marine tract 
became estuarine, fluviatile, and a region of wearing of high land. 
Subsequently the enormous subsidence took place, doubtless almost 
synchronously with the deposit of the thousands of feet of the 
Mafnchhars ; and yet the sea never broke in: it was far away. 

The area of change was vast; and it appears to be unreasonable 
to associate all these deposits under one geological formation. 

The disassociation of the Manchhar and Gaj series is a necessity ; 
and the nature of the fauna, so singularly allied to that of Pikermi, 
necessitates its relegation to the early Pliocene time. 

In following up this subject it must be remembered that the 
Sivalik strata, the horizon of which is above the Lower Manchhars, 
have a vast vertical as well as horizontal development, and that 
osseous remains have been found in them throughout their height. 
The fauna as a whole has a later facies than that of the Lower 
Manchhars, and resembles, even in its African element, that of 
Pikermi. : 

On the ground of inferred equivalency with the Upper Manchhars, 
and of faunal alliance with the assemblage at Pikermi, it must be 
credited that the Sivalik strata are of Pliocene age. 


208 PROF. P. M. DUNCAN ON THE 


There is another argument which has not hitherto been employed 
and which favours this theory. 

The Sivdlik strata rest on, or are in contact with (along a line of © 
fault), certain plant-bearing strata of the Sirmur series which are 
associated with the underlying Nummulite-bearing Eocene rocks of 
Subathu. 

The nature of the flora, so far as it has been examined, is not 
very foreign to that of India at the present time; and by the same 
kind of reasoning which asserts the separation of other terrestrial 
and underlying marine strata, these beds, called Kasaoli and Dugr’hai, 
may well be the remains of the Miocene land. 

Whence were the vast thicknesses of the sands, clays, and con- 
glomerates of the Manchhar-Sivaliks derived? ‘They represent a 
ruined mountain-chain in bulk; and they are found not only on the 
flanks of, but also within, the orographical systems of which they 
form parts. 

It appears from the study of the history of the Himalayas by 
Strachey, Stoliczka, Medlicott, and those Indian Geological Sur- 
veyors who have laboured so industriously of late years, that a low 
mountain-chain existed on the area after the Nummulitic age—the 
results of a post-Nummulitic earth-movement. It was probably 
a broad chain, and not a sufficient barrier to prevent the roaming of 
the animals on and over it in the subsequent geological age. This 
chain appears to have had an axis of old rocks; and the whole 
suffered denudation during the age of the Sivalik Mammalia. In 
Hundes Strachey found the great tableland (now at an altitude of 
from 14,000 to 16,000 feet) to consist of sediments filling up a basin 
in old rocks to the depth of 3000 feet—sediments which included 
osseous remains of animals that could not have traversed high and 
difficult mountain-ground. . 

Part of the Sivalik sediments were formed out of low mountain- 
ground by fluviatile denudation; but to account for the vast deve- 
lopment of the rest on the flanks of the present mountain-system, it 
is necessary to admit that the upheaval, and much of the north-and- 
south crush of the Himalayas, occurred part passu with the slow 
accumulation. Elevation and diminution of the breadth of the area 
progressed ; and doubtless much of the great crush which folded and 
often reversed the flanking strata was final. 

The Sivalik strata rest on the flanks of the chain and on old rocks 
within the range; and they were the youngest deposits affected by 
the mountain-making. Hence the Himalayas, as a grand system, 
culminated during and subsequently to the collection of these strata, 
which have been pronounced to be Pliocene in age. 

The Sivalik deposits in the tableland of Hundes are overlain by 
relics of the great glacialization of the Himalayas. Hence, before 
the vast glaciers of the glacial period accumulated, valleys had been 
Worn out and denudation had proceeded. So it is necessary to 
recognize that the culmination of the movements which developed 
the height of the Himalayas occurred in preglacial times and 
during the Pliocene age. 


CORALLIFEROUS SERIES OF SIND. 209: 


To complete the serial changes, it is to be remembered that a 
Postglacial fauna was found in the old alluvium of the Jumna, 
a wash-down of the Himalayas. 

With regard to the Upper Manchhars, they were implicated in the 
great orographical movement, which was contemporaneous through- 
out the extrapeninsnlar area of India; and they are of Pliocene age. 

The details regarding the succession of strata and of many of 
their organic remains are to be found in the publications of the 
Geological Survey of India, and in the communications to this 
Society by Strachey, Grant, Falconer, and others. The ‘Manual of 
Indian Geology, by Medlicott and Blanford, an abstract of the 
labours of the Survey, contains them in a condensed form. I am 
under great obligations to all those writers, and also to Messrs. Med- 
licott and Blanford for much unpublished information ; and in thus 
heartily acknowledging my obligations, I am glad to have the 
opportunity of expressing my assent to their conclusions regarding 
the age of the Himalayas. 


Discussion. 


The Presrprent remarked on the persistence of Mesozoic types in 
the Tertiary strata of the Himalayas. 

Mr. Branrorp expressed the obligations of himself and other 
Indian geologists to Dr. Duncan for his researches on the Corals. 
which they had collected. The base of the series of Sind consists 
of a limestone containing Hippurites; and above these Cretaceous 
beds are strata partly unfossiliferous and the representatives of the 
Deccan traps, the whole being overlain by the Lower Eocene and 
Nummulitic. This succession is shown both in the Laki and the 
Khirthar ranges. The Nari or Oligocene group is 5000 or 6000 
feet thick ; its upper subdivision, which is much thicker than the 
lower, is of freshwater origin, and contains imperfect plant-remains. 
This is overlain by the Gaj and Manchhar. The unconformities which 
occur in the series are purely local. He was gratified to find 
that Dr. Duncan had arrived at the same conclusion as the Geolo- 
gical Survey of India as to the age of the Sivdlik beds. He 
replied to the opinions expressed by Prof. Boyd-Dawkins and 
Mr. Bose on this subject. 

Lieut.-Col. Gopwiy-Auvsten remarked upon the greater contortion 
of the Nummulitic strata in the Western Himalayas as compared 
with those of Assam. 

The Autor stated, in reply to the President’s remarks, that 
while the genera of Corals are remarkably persistent, the species are 
not. He bore testimony to the great value of the volume published 
by the Geological Survey of India. He doubted the value of the 
terrestrial Mammalia as fixing the age of the strata overlying the 
Kocenes. 


210 C. CALLAWAY ON THE ARCH XAN 


15. The Arncoman Guotocy of Anetusny. By C. Carraway, Hsq., 
M.A:, D.Sc., F.G.S. With an Appenprx on the Microscopic 
SrRuctuRE of some ANGLESEY Rocks, by Prof. T. G Bonney, 
M.A., F.RB.S., Sec.G.8. (Read January 5, 1881.) 


[Prats VIIT.] 
Introduction (p. 211). 
A. Description of Areas and Sections. 
i. Menai Anticline (p. 211). 
a, Gmeissic series. 
(1) South of Pentraeth. 
(2) Mynydd Llwydiarth. 
(3) Gaerwen to Menai Bridge. 
b, Slaty series. 
If. Llangefni “Syncline” (p. 215). 
a. Slaty series. 
(1) Llangristiolus slates and grits. 
(2) Llangefni conglomerates and shales. 
(3) Cerrig-Ceinwen slaty and calcareous group. 
6. Gneissic series. 
(1) Coast section from Porth Nobla to Aberffraw Sands. 
(2) Railway-section from Ty Croes to Bodorgan. 
(3) Distribution of the subdivisions in localities to the north-east. 
IIY. Central Zone (p. 218). 
a. Slaty series. 
6, Gneissic series. 
(1) Bodafon Mountain. 
(2) Section between Llangwyllog and Llanerchymedda. 
(3) Area south-east of Paris Mountain. 
(4) Structure of the Zone. 
ITV. Northern Area (p. 221). 
(1) Voleanic group of Paris Mountain. 
(2) Chloritic schists of Mynydd Mechell. 
(3) Llanfechell grits. 
(4) Rhosbeirio shales. 
(5) Sharply contorted group south-east of Amlwch. 
(6) Slates and limestones of Amlwch and Cemmaes. 
Y. North-western Area (p. 224). 
Section from Porth y defaid to Pen bryn ’r Eglwys. 
VI. Western Area (p. 225). 
(1) The Mainland. . 
(2) Holyhead district. 
(3) Rhoscolyn district. 
B. Summary of Results. 
1. Distribution of the Rocks (p. 227). 
a, Gmeissic series. 
(1) Geographical. 
(2) Stratigraphical. 
b. Slaty series. 
Ii. Evidence of Age (p. 229). 
a. Relations to Palzozoic groups. 
6. Relations to each other. 
c. Relations to other areas. 
(1) Caernarvonshire. (3) Shropshire. 
(2) St. David's, (4) Charnwood. 
Conclusions (p. 232). 
Appendix (p. 232). 


eee ee ee 


Quart. Journ. Geol, Soc. Vol. XEXVIL. Pl. Vill 


Fig. 6. Section ir lower Quarry, Nebo. 


Fig. 5. Section in upper Quarry, Nebo, 


jes = 
yy en ob s 
U Yf S S 


Uf) 


Yi 
Yi 


Fig.3. Section across lynydd Llyydiarth . 


we a a OT Oy 


Mi 
, 
' 
. 
‘ ‘ 
‘ 
as v 
, he 4 
i 


Quart. Journ. Geol. Soe. Vol. AMV Pl. Vi 


Fig. 6. Section in lowe Quarry, Nebo. 


Fy. 1. 


MAP of ANCLESEY 


Fig.2.; Generalized Section trom Holyhead Mountain to Menar Strats. 
ESE. 5.5.W. EE. Malllraeth 
Tlangefne Marsh 


Werrrre EMAL A= 


Ss 


with a, overlying 


Fred* Donyerlield Lith Lorudon 
18 


GEOLOGY OF ANGLESEY. PU: 


INTRODUCTION. 


[I wave to submit evidence in support of the conclusion that there 
are represented in Anglesey two Precambrian or Archean groups, 
each distinguished by well-marked lithological characters. They 
may be named respectively the Slaty and Gneissic formations. The 
‘“‘ Pebidian ” schists, as recognized by Dr. Hicks, I include in the 
eneiss group ; but, in other parts of the island, I have worked out 
a great assemblage of fragmental felspathic rocks, which I believe 
to be of true Pebidian age. I shall also attempt to prove that the 
quartz rocks of Anglesey are included in the gneissic series. ‘The 
microscopic notes, kindly furnished by Prof. Bonney, are based upon 
the specimens only, and have all the value of independent evidence. 
The growing importance of these ancient rocks would seem to re- 
quire that the provisional term ‘“ Precambrian” should give place 
to a word which can be permanently used; and I have ventured to 
adopt the term “‘ Archean,” so widely employed by Continental and 
American geologists. To the words ‘“‘ Azoic” and “ Hozoic” there 
are obvious objections. 


A. Description oF AREAS AND SECTIONS. 


I. Menat ANTICLINE. 


For convenience I accept this name for the rocks lying between 
Menai Straits and the great fault which throws down the newer 
Paleozoic rocks of Malldracth Marsh, though the term is not strictly 
applicable. It is true that the strata on the south-east side of the 
area dip to the south-east, and those on the north-west side to the 
north-west; but between these two extremities the undulations are 
frequent, and some of the dips are probably overthrows. 

a. Gineissec Series. 

_ wo Varieties of Gnerss—The structure of the area is rendered 
much clearer by the recognition of two prevailing gneissic types— 
a dark micaceous or hornblendic variety, associated with chloritic 
schists*, and a greyish or light-red rock clearly foliated into an or- 
dinary ternary gneiss. in the Craig-yr-allor anticline ft the two 
varieties are also recognizable, the grey gneiss passing up through 
the dark type into the granitoidite. In the Menai district the dark 
schist holds the same relation to the grey variety, and the passage 
between the two may be seen at many points. 

1. Gneiss south of Pentracth.—Next to the fine section at Gaer- 
wen #, the rocks south and east of Pentraeth require notice. The 
band coloured “ greenstone,” running from Tai hirion north-north- 
east to Plas-gwyn lodge, appears to be a highly-altered hornblendic 
gneiss, with a north-west dip. In the field to the south-east of the 
lodge is a small quarry of typical grey gneiss; and in the plantation. 

* Throughout the paper the terms “schist ” and “schistose ” are strictly con- 
fined to foliated rock. 

t Geol, Mag., March 1880, p. 119. + Geol. Mag., March 1880, p. 121. 


Dal C. CALLAWAY ON THE ARCH HAN 


a few yards to the north-west we find the dark-green schist in its 
usual position above, both dipping to the north-west. 

Following the strike to the north-east, the gneiss is seen to con- 
tain angular pieces of quartz, some of which are flattened in the 
foliation-planes, so as to appear like ordinary folia. 

2. Mynydd Llwydiarth.—This elevated ridge, trending to the 
north-east for nearly a mile and a half, is composed of highly con- 
torted gneiss. The general dip is north-west; but in some places 
the beds undulate to the south-east ; and the thickness cannot be 
great. The rock is more quartzose than usual; and the quartz frag- 
ments become much more numerous,—the two facts suggesting the 
proximity of quartz land. The gneiss, rolling in frequent contor- 
tions, is finely exposed in low vertical cliffs ; and the angular pieces 
of white quartz studding and projecting from the surface produce a 
very striking effect. Both types of gneiss appear to be present;. 
but the felspar being less abundant, the rock is brought into closer 
resemblance to the micaceous and chloritic schists of Holyhead. On 
the north-western slope the gneiss is extensively brecciated, and 
the presence of a fault is further inferred from an abrupt transition 
to younger formations. At the foot of the slope, near the fault, the 
eneiss becomes somewhat granitoid, suggesting the commencement 
of the passage into granitoidite, which is seen so clearly in the Craig- 
yr-allor anticline. At the extreme north-east of the ridge, on Red 
Wharf Bay, the green schist is exposed for some distance across the 
strike to the east ; but south of Wern we come onto the grey gneiss, 
all the dips being north-west. Further to the east, at Bryn y Castell, 
the grey type is again seen, but with the dip reversed; and at Hafodty, 
still further to the south-east, the dark schist comesin again. This 
is a clear anticline, the grey gneiss exposed at the apex throwing off 
green schist in opposite directions. The structure of this district is 
shown in Pl. VIII. fig. 3. 

3. Gaerwen to Menart Bridge.—Crossing the anticline from Gaer- 
wen, we find at the railway-junction that the dark schist has rolled 
over to the south-east; but half a mile to the south-east, near 
Llanddaniel, the series, represented by contorted quartzose chloritic 
schist, dips north-west. Exposures then are rare till we come to 
the Anglesey column, where dips are in opposite directions, but 
usually to the north-west. This rock is of the ordinary dark green 
type, occurring everywhere above the grey gneiss in both the Menai 
and Craig-yr-allor areas. It has been described by Prof. Bonney *. 
On the Straits, near the west end of Menai Bridge, green schists 
occur of the ordinary varieties. 

From the above facts it is clear that the rocks of the anticline 
belong to the gneiss series. Well-foliated gneiss occurs at numerous 
localities; and the green schist 1s so intimately associated with it, 
as an upper band, as to remove all shadow of doubt that the two 
types form one unbroken group. In this area I have not found a 
trace of any thing but true crystalline schists. 

The thickness of the group is not great. The foldings are nume- 

* Quart. Journ. Geol. Soe. vol. xxxv. p. 308. 


GEOLOGY OF ANGLESEY. 213 


rous, the lower gneiss being brought up to the surface at many 
points. Probably the entire series is exposed west of Gaerwen, 
where the total thickness can hardly reach 1000 feet. 

An isolated mass of schist occurs south-west of Pentraeth. It is 
about a mile long by half a mile broad, and is bounded by faults on 
all sides. Carboniferous Limestone surrounds it on three sides. On 
the east it is separated from the main mass by a faulted strip of the 
formation now to be described. 


b. Slaty Serves, 


This wedge is composed of rocks of a newer Archean group. 
At Tan y graig, south-west of Pentraeth, is a greenish, coarse, 
slaty variety. In the fields to the north-west, a jaspery rock 
projects in small knolls through the turf; but its relation to 
the bedded type is obscure. Crossing a hollow to the east, we 
come to a ridge, the western slope of which is occupied by 
massive breccias. The fragments and the matrix are similar, and are 
composed of a grey and purplish dolomite (Note 40, p. 235). Further 
to the north-east, above Wugan bach, is a light-purple slaty rock, 
which in places is literally smashed. Between this point and Plas 
ewyn, along the ridge, the prevailing type is a purplish ashy rock, 
sometimes brecciated, and in places altered almost to a hornstone. 
The dip is high to the north-west. The breccias of this ridge are 
obviously connected with faulting. North of Plas gwyn, on the 
same strike, is a puzzling rock, which, under the microscope (Note 
41, p. 236) appears to be a limestone rendered impure by volcanic 
mud. Slaty beds are again seen to the north-east of Pentraeth, 
dipping south-east. 

For the greater part of its length this mass is separated from the 
Menai anticline by a faulted strip of Paleozoic shale; but north- 
east of Rhiwlas it comes up against the gneiss, the fault between 
the two formations running along a hollow. The heavy breccias 
which fringe both groups point to excessive fracture and crushing. 


Il. Luaneernt *‘ SyNcLINE.” 


Crossing the Carboniferous strata of Malldraeth Marsh, we come 
to an altered and contorted group of rocks which bear a superficial 
resemblance, especially in their colour, to the Menai schists, and 
have hitherto been regarded as a part of the same group. But on 
a careful examination of the lithology of this series, fundamental 
differences are observable. Even my first day’s work in the district 
convinced me of the existence of Pebidian rocks of the St.-David’s 
type; and fuller working-out of the area brought to light analogies 
with other Pebidian localities. 

Slaty rocks occupy only the south-eastern side of the apparent 
syncline lying between the Carboniferous on the south-east and 
the granitoid band on the north-west, the north-western side being 
composed of gneissic strata. The two groups are brought together 
by a fault or faults; but as they dip towards each other, an appa- 
rent basin is formed (see section near Aberffraw, Pl. VIII. fig. 2). 


Q.J.G.8. No. 146. f°) 


aa C. CALLAWAY ON THE ARCHAZAN 


a. Slaty Serves. 


1. Llangristiolus Grits and Slates—The lowest beds I have ob- 
served are exposed near the Llangristiolus turnpike. The rock is 
clearly bedded; but its lithology is obscure, and the jointing is so 
close that clean fractures are hard to get. It appears to be a quartzo- 
felspathic grit, altered into a sort of hilleflinta. Some bands are 
coarser, grains of quartz and ielspar being visible to the naked eye. 
A similar rock is common in the Pre-Cambrian rocks of the Chureh- 
Stretton district (Shropshire). 

A little further to the north, near Cerrig ddwyffordd, a quarry- 
section is very interesting. At the base is a purplish grey felspathic 
grit (Note 42, p. 235). This is overlain by a pale-green slaty rock of 
Charnwood type, associated with, and apparently passing into, a 
sort of porcellanite (Note 43, p. 234), singularly like a rock at Caer 
bwdy, St. David’s. ‘The Charnwood facies of these varieties is also 
noted by Prof. Bonney. The section is capped by a repetition of 
gritty bands. The dip is northerly, at from 40° to 50°. Similar 
rocks are exposed in neighbouring fields. 

2. Llangefni Conglomerates and Shales.—TYo the west and north- 
west of the Llangristiolus sections are several exposures of a greenish 
conglomerate. ee is seen on the Holyhead road, near Waen She and 
north of Cerrig ddwyffordd, at the arrow mark, but is ver y fully ex- 
posed in the railway- -cuttings north-west of Llangefni, the enclosed 
fragments, weathering white, being clearly visible even from the rail- 
way carriage. The matrix is green shale. The pebbles are very 
varied, the following being the principal types :—quartzite, common ; | 
pinkish grit (Note 38, p. 235) of quartz and felspar; greenish fel- 
spathic grit; and green and grey hornstone. Of these, quartzite is 
the only ingredient which is certainly foreign to the slaty series ; the 
others are such as are found in some part of the group. The con- 
glomerate would then seem to have originated in contemporaneous 
denudation, such as is common in yolcanic rocks, together with the 
wearing-down of a preexisting land composed of quartzite. 

Interstratified with the conglomerate are beds of purple and 
green felspathic shale, with some bands of hornstone.. The resem- 
blance of these rocks to the St.-David’s types is unmistakable. 

The road-sections in this district display a prevailing north-west. 
dip; but the more complete exposures in the railway-cuttings 
reveal frequent undulations, so that the thickness is not great. 

A similar group, probably on the same horizon, is well displayed 
in the cuttings east of Bodorgan station, 6 miles south-west of 
Llangefni. The prevailing rocks are purple, green, and grey ashy 
shales, with some felspathic breccias. Conglomerates occur con- 
taining pebbles of felsite, in which microscopic examination (Note 39, 
p. 236) reveals a very interesting resemblance to modern lavas. 

Associated with the shales at Llangefni station is a thick bed of 
quartzose grit, such as might have been preduced by the denudation 
of an ancient quartzite. 

Still following the strike to the south-west, we find the slaty 
series well exposed at numerous points on the coast from Bodowen 


GEOLOGY OF ANGLESEY. 215 


round to the Aberfiraw sands. On the shore near Bodowen, Llan- 
gefni conelomerate is underlain by grit. On the east side of Porth 
twyn mawr, it overlies slaty rock, and contains large unrounded 
pieces of it. The same conglomerate, repeated by folding, occurs at 
intervals as far as the Aberffraw sands. 

Quartzite occurs in lenticular bands in pale-green slate west of 
Porth Cadwaladr. This association is also very common in Northern 
Anglesey. : 

Near, or at, the base of the slaty series of this district are several 
masses of quartz rock of obscure origin. The most prominent of 
these is the craggy hill* called Craig fawr, above Llangefni. It 
1s a massive white rock, in which the quartzite structure is still 
apparent. It is surr ounded by Carboniferous strata, and it probably 
formed an island in the Carboniferous sea. The same rock may be 
traced at intervals along the strike to the south-west. One conspicuous 
crag of it is seen at Bethel, north-east of Bodorgan station. I can 
only suggest that we may have in these bosses the remains of an 
ancient land, from which the Pentraeth gneiss derived its angular 
fragments, and the Llangefni conglomerate its rounded pebbles and 
its quartz grits. 

3. Cerrig-Ceinwen Slaty Group.—These rocks lie to the north- 
west of the last group, with the same (north-west) dip ; and as they 
present important lithological differences, they can hardly be repe- 
titions. 

North east of Cerrig Ceinwen, a little south of the Holyhead 
road, are green and purplish slaty beds. Some bands are calcareous ; 
and the formation on the whole resembles the rocks forming the 
ridge at Wugan bach. Large nodules of jasper are abundant. Their 
origin is not absolutely clear; but 1 am disposed to regard them as 
included pebbles. The beds have a high dip to the north-west. 
On about the same strike to the south-west are pale green ashy bands. 
Close to Cerrig-Ceinwen church, to the north, are prominent ridges, 
composed of a greenish rock, so tough that it was difficult to obtain a 
specimen. It has a superficial resemblance to a greenstone, but is 
undoubtedly an indurated sedimentary rock, presumably an ash. It 
suggested the “‘ greenstone” of Clegyr Foia, St. David’s. 

A fine exposure in a quarry west-south-west of Cerrig Ceinwen, 
on perhaps a little lower horizon than the jasper conglomerate, 
deserves attention. The rock is typical of the slaty series of 
Anglesey. In the field it appears asa pale green slate or indurated 
shale, sometimes faintly banded. Comparing it with some of the 
Charnwood slates (Note 44, p. 234), the only difference I could detect 
was that the Anglesey type displayed a slight lustre indicative of 
incipient metamorphism. It sometimes passes into a rock in which 
the alteration has been carried further, sometimes into a sort of 
hornstone. ‘The dip is still to the north-west. 

At Bod enlli we are apparently on the horizon of the jasper 
conglomerate. The conglomerate itself is not exposed; but we have 
limestone, purple slates, and green breccias and shales, not unlike 
those associated with the j jasper. 


* Coloured ‘‘ greenstone” on the map. 


216 C. CALLAWAY ON THE ARCH AN 


Over a mile to the west-south-west, at Cerrig engan fawr, there 
is a considerable exposure of a very tough, green, ashy-looking rock, 
similar to the variety north of the church. 

All the above rocks dip to north-west ; but a little further to the 
west, at Ty’n-y-buarth, the beds dip to south-east. The strata at 
this locality are highly contorted on a small scale, and are very 
similar to the gnarled rocks east and south-east of Amlwch, the 
alteration, as in that district, having proceeded further than at 
Cerrig Cemwen. In all other respects these altered rocks resemble 
the Cerrig-Ceinwen slates. ' 

b. Gnerssice Serves. 

The band of gneissic rocks is hardly a mile in width at Bodwrog, 
where it is separated from the Pebidian by a faulted strip of Pale- 
ozoic shale ; but it gradually expands to a breadth of over two miles 
in the coast section on the south-west. The fault separating it 
from the slaty series runs from near Bodwina, north of the 
Holyhead road, in a south-south-west direction, losing itself under 
the sands of Aberffraw. On the west of the sands are nothing but 
schists ; on the east are nothing but rocks of the slaty series. It 
will be necessary to describe two sections across this zone. 

1. Coast Section from Porth Nobla to Aberffraw Sands.—On the 
coast, west of Porth Nobla, is a considerable exposure of greenish 
felspathic rock similar to the halleflinta seen on the strike to the 
north-east, near Ty Croes. It appears to form an anticline. On 
Porth Nobla it clearly dips south-east, and passes gradually up into 
grey gneiss. To the west it presents but slight indications of 
Stratification ; but as we proceed eastward planes of separation grad- 
ually appear, which, at first obscure, become more and more distinct, 
till the rock is undistinguishable from a true gneiss. I have long 
since noticed an unbroken passage between hilleflinta and gneiss 
in the Wrekin; and Dr. Hicks has recorded the presence of an 
incipient foliation in the hilleflinta (‘‘ Arvonian’’) of St. David’s. 
The grey gneiss soon passes up into quartzose gneiss, quartz-schist, 
and quartzite. These siliceous types, which are of much theoretical 
interest, occur in great force, and form the main part of the 
headlands of Mynydd baen and Pen y cnwe, which project between 
Porth Nobla and Porth Trecastell. On the west side of Porth 
Trecastell is a band of calcareous quartz-schist, or quartzose foliated 
limestone. The silicate appears to be tale or chlorite. ‘There is 
a break in the section at the centre of the bay; but as the cal- 
careous schist reappears on the east side, there can be but a slight 
break, if any, in the succession. Overlying the last-named rock is 
a considerable exposure of decomposed grey gneiss interstratified with 
bands of quartz rock, the whole being much contorted and broken. 
A greenstone dyke appears here. After a few undulations, the de- 
composed rock disappears, and ordinary white and grey gneiss, some- 
times rather quartzose, dips steadily to the south-east for some 
distance. Then comes in the well-known green schist of the 
Menai type, which is continued to the Aberfiraw sands. It is well 


GEOLOGY OF ANGLESEY. ATE 


seen on Porth gwyfen with the normal south-east dip. On the 
shore, near the church, an undulation brings up grey gneiss. 
Then the south-east dip is resumed; but at Trwyn du, just before 
reaching the Aberffraw estuary, the beds turn up to the south-east, 
and a syncline is formed. The sucgession described is shown in 
fig. 4. 

52. Section on the Railway from Ty Croes to Bodorgan.—tThe suc- 
cession is substantially the same as the last; but it furnishes one 
or two additional facts. At Felin bont is the hilleflinta first 
noticed by Dr. Hicks, who rightly conjectured, as the last section 
proves, that it passed to the south-east beneath the schist at Ty 
Croes*, though he was in error in placing it above the granitoidite, 
which les at the very summit of the gneiss series. West of Ty 
Croes quartzose schist is exposed in the road. 

Coming to the railway-cuttings, we find to the east of the station a 
good section of grey quartzose gneiss (Note 34, p. 233) dipping south- 
east. This rock is one of the most prominent types seen west of Porth 
Trecastell. Grey and white gneiss are seen for more than a mile 
with the same dip. In the cutting from near Bodgedwydd to near 
Graig bach, the grey gneiss grows gradually darker and darker, and 
passes without a break into the dark green Menai type, which is 
continued to the end of the exposure. The average dip is 40° to 
south-south-east. Leaving the line, we find in a road-section north- 
east of Tre’ Iddon that the dip of the dark schist has changed to the 
north-west. Hast of this point we come almost at once to the slaty 
beds and conglomerates of the newer series. 

The railway-section is thus seen to agree precisely with the suc- 
cession on the coast. In some points it is less complete; but it dis- 
plays very clearly the passage of the grey into the dark schist, so 
well seen in the Menai anticline. 

3. Distribution of the gneissic Subdivisions to the North-east. Halle- 
jinia.—This band gradually narrows towards the north-east, being 
cut out by the fault that brings the lower part of the gneiss series 
against the granitoidite ; so that at Gwalchmai it is hardly visible, 
and at the section; north of the Holyhead road the grey gneiss is 
in immediate contact with the granitoidite. Some of the gneissic 
rocks of Gwalchmai display a transition towards the halleflinta 
types. 

Quartz-schist. This zone is not well exposed near ‘'y Croes. To 
the north-east it is seen at Melin Ddrydwy andGlan’rafon ; but it may 
be best studied at Gwalchmai near the church, on the road to the south- 
west of the church, and on the Holyhead road. In the last locality 
the granitoidite is brought against it by the fault. The dip is to 
the south-east, except on the Holyhead road, where it is reversed. 
There is probably a broken anticline at this point. The prevailing 


* I was at first disposed (Geol. Mag. March 1880, p. 128) to associate this 
halleflinta with the granitoidite, since similar rock, together with quartz- 
felsites, is included in that group; but the coast-section has cleared up several 
difficulties. 

t Geol. Mag. March 1880, p. 124. 


21 


oa) 


C. CALLAWAY ON THE ARCHAAN 


type is a true quartz-schist, sometimes passing into a quartzose 
eneiss. As it was important to critically compare this rock with 
the quartz rocks of Holyhead and Bodaton Mountains, I have had 
four specimens cut; and Prof. Bonney appends full descriptions 
(Notes 25-28, p. 233). 

Grey Gneiss. From the section east of Ty Croes station, this 
band may be traced to the north-east, forming the low ridge from 
Tal y llyn to Gwalchmai. Passing to the east of Gwalchmai church, 
it 1s well exposed about Bodkin tae. and on both sides ef the 
marsh (Cors Bodwrog) and pool. At Bodwrog church it is in contact 
with granitoidite, the quartz-schist having in its turn been cut out. 
In the ridge to the east of the church the gneiss is interstratified 
with a highly crystalline limestone, or calcite-schist (Note 51, p.236). 
From the microscopic description 14 will be seen that this rock re- 
sembles the quartzose limestone of Porth Trecastell ; and it is pro- 
bably on the same horizon. The normal south-east dip is preserved 
as far as the old Holyhead road, north of the marsh ; but near here 
a change takes place, and black Paleozoic shales come up to the 
eranitoidite. 

Dark Schist. This zone is not well seen north of the Holyhead 
railway. The fault which limits it on the south-east gradually cuts 
it out, so that east of Gwaichmai the slaty series approaches the 
grey gneiss. The gneiss is exposed at Pen bryncele, dipping south- 
east ; and half a mile to the south-east, near the +2 milestone on 
the Holyhead road, we find green slaty rocks with a north-east dip. 


IIL. Centrrat Zone. 


This band extends across the island from sea to sea. It is the 
most complicated part of this broken and contorted area; but by 
steady attention to the lithology, assisted by some stratigraphieal 
indications, clear results may be secured. 


a. S laty Series. 


In following up the gneissic rocks towards Bodafon Mountain in 
order to ascertain the relation between the gneissic and quartz 
groups, I was surprised to find an area of slaty rocks between the 
two. At Plas Llanfihangel dark gneiss and granitoidite dip to the 
south-west; and a little further to the north, at Ma’n-addwyn, 
felspathic shales dip as if they would pass below the gneiss. The 
rock is of a typical St.-David’s type, and unquestionably pelongs to 
the younger series. High up on the south-east slope of the moun- 
tain are similar rocks dipping south-east. The dips in both cases 
being away from the quartz rock, it was natural to infer that the 
shales rested immediately upon it. Further examination proved 
that such was not the case. 

At the south-west end of the quartz ridge north-west of Tyn— 

‘Wlidiart the flanking rock is a sort of hornstone, banded pink and 
green, of a common Pebidian type. Approaching the hill, the horn- 
stone is seen to dip away from it to the south-west; but surmounting 
the low ridge made by the. hornstone, the beds are observed to roll 


GEOLOGY OF ANGLESEY. 219 


over and plunge directly towards the quartz at the very junction. 
A little further to the north, above the farm, the evidence of dis- 
cordance is, if possible, still clearer. In the quarry is a good section 
of a greenish slaty rock, with the beds striking directly at the quartz 
forming the ridge at a distance of about 20 yards. 

It is thus evident that the slaty series is faulted against the quartz 
group of Bodafon Mountain. 


b. Gneassic Series. 


1. Bodafon Mountain.—This mass cannot be brought into direct 
stratigraphical relation with the gneiss group, for the reasons just 
explained; but the lithological resemblance of this rock to the 
quartz-schist of Gwalchmai and Mynydd baen is so close as to 
render the correlation highly probable. In hand specimens some of 
the varieties are undistinguishable. Prof. Bonney, who was not 
aware of my views when working on the specimens, groups a typical 
variety (Note 29, p. 233) from the summit of the mountain with 
the Gwalchmai schists. The rock of the western ridge, which is a 
little more schistose, is also closely represented in the south-western 
localities. The inlier, surrounded by newer Paleozoic rocks, which 
lies to the south-east is precisely similar to the eastern part of the 
hill. 

2. Section between Llangwyllog and Llanerchymedd.—The sue- 
cession between these points is similar to that of the coast-section 
between Ty Croes and Aberfiraw; but the granitoidite is wanting 
in the latter, and the dip is reversed, so that the two sections repre- 
sent the two sides of a broken anticline. 

The granitoidite occupies a large area south of Llanerchymedd. 
Round Coedana is a grey variety, rather small-grained. To the south, 
at Rhydgoch, it is much coarser, consisting of fragments of pink 
felspar in a matrix of smaller bits of quartz, felspar, and chlorite. 
The older geologists would have called it a porphyritic granite. 

The dark: schist is exposed about half a mile further to the south, 
at Glanrhyd, being the band of “ greenstone” of the Survey. Prof. 
Ramsay originally suspected the metamorphic origin of this rock ; 
and, after microscopic examination,: Prof. Bonney is disposed to 
consider it a hornblendic gneiss, containing felspar, epidote, horn- 
blende, and some quartz, the felspar being decomposed, and the 
whole a good deal “‘ messed.” Any obscurity arising from such an 
altered rock is removed if we follow the strike to the north-east. 
To the south-west of Graig Ilwyd there is a good exposure of 
hornblendic gneiss interstratified with granitoidite. The gneiss 
sometimes passes into hornblende-schist. The hornblende is well 
erystallized ; and the alteration is very slight. 

The grey gneiss is well seen about two furlongs south of Glanrhyd, 
in a quarry north of the stream. The dip is high, to the north-west ; 
and the rock is of the ordinary type. 

No rock is seen on the road for two or three furlongs to the 
south ; but north of Llangwyllog church there is a slight exposure 
of hallefinia too small to display dip. There is room for the 


220 C. CALLAWAY ON THE ARCH EZAN 


quariz-schist between this rock and the grey gneiss; but I cannot 
affirm its presence. 

3. Arca south-east of Paris Mountain.—Vhe gneissic rocks of this 
district are on the north-easterly prolongation of the strike of the 
central zone; and there is no doubt that they belong to the same 
series: the dip, the succession, and the prevailing rock-types are 
the same. 

On the coast between Dulas Bay and the fault north of Porth 
lygan the succession is not satisfactory, owing to great disturbance. 
South of Porth lygan the shore is mainly occupied by sandstones 
and conglomerates, presumably of newer Paleozoic age, deposited 
on an eroded surface of gneiss, and sometimes wrapping round little 
promontories of the older rock. ‘The prevailing dip is easterly. On 
Porth lygan the gneissic series is well exposed. Grey gneiss, dark- 
green schist, granitoidite, and quartzite (all the types which occur 
on the strike inland) lie in confused, contorted and shattered masses. 
At the north of the bay the rocks are less disturbed, and dip to the 
north-west. Black Ordovician (Lower Silurian) shales are faulted 
down to the north, resting, in a clear coast-section, against the 
granitoidite. 

Between Llanwenllwyfo and Nebo the succession is more dis- 
tinct. Thin-bedded gneiss is seen at several points north-west of 
Llanwenllwyfo and north of Plas uchaf; the overlying green schist 
is well exposed around Rhos manarch; and the capping grani- 
toidite, with associated schist-bands, forms the ridge between Nebo 
and ‘'y-Newydd, the whole dipping to the north-west. In two 
quarrics south-east of Nebo there is well exposed a strong band of 
quartzite and quartzose conglomerate, similar to some of the masses 
on the coast. I have elsewhere* given reasons for believing that 
this is the equivalent of the quartzose conglomerate of Twt Hill; 
and I stated that in these quarries the rock is unconformably oyver- 
lain by black shales of (at least) Bala age. If the shales are 
Tremadoc or Arenig, as Prof. Hughes has since maintained, the fact 
is still more strongly confirmatory of the Archsan age of the 
quartzite. As the point is important, I submit a section (figs. 5 & 
6) from each of these quarries. 

Since this quartz-rock is Precambrian, is conformably underlain 
and overlain by gneissic strata, and occurs on the coast in the closest 
association with gneiss and granitoidite, there can be no hesitation 
in referring it to the older Archean series. It must not. however, 
be confounded with the quartz-schist of Bodafon Mountain, which is 
lower in the succession. 

Granitoidite and dark schist are found along the ridge to the 
west, and at the base of Paris Mountain. I have not examined the 
latter locality ; but Mr. Allport kindly allows me to use some notes 
made by him in May 1868. The numbers refer to specimens now 
in the British Museum. 

“443, Fine-grained granite? South-east flank of Paris Mountain. 

“444, Granite? 100 yards from last. 

* Geol. Mag. March 1880, p. 118. 


GEOLOGY OF ANGLESEY. DN 


© 338. Hornblende-schist, passing into a more compact rock (339), 
not schistose, or very slightly so; and this, again, into hornblende 
and felspar rock, very similar in appearance to that at Malvern (340). 

‘«‘This is an interesting case, as the above varieties were taken 
from different parts of the same large block. As in Malvern rocks, 
epidote occurs near the joints. No.340 might be called hornblendic 
eneiss; the felspar 1s well crystallized in many parts of the mass. 
In this place there is also hornblende and mica rock; and in some 
parts hornblende and felspar occur in separate layers. The granite 
has a gneissose structure; and to my mind the evidence is clear that 
the whole form a series of metamorphic rocks.” 

I am glad to quote confirmatory evidence by so competent a 
ithologist, and trust that it will be unnecessary to multiply retuta- 
tions of the old “‘ granite-vein”’ hypothesis. 

A, Structure of the Zone.—-For structural purposes it is necessary 
to include in this band the gneissic area between Ty Croes and 
Aberfiraw. The whole forms a complex and shattered anticline. A 
central fault margins the granitoidite on the east from Caernarvon 
Bay to a mile north-east of Bodwrog; then 16 curves a little to the 
east, and, passing near Llangwyllog, leaves room for the underlying 
gneisses. ‘The strata dip in opposite directions on each side of the 
fault. This dislocation thus appears to pass along the summit of 
the anticline; but it does so obliquely, permitting the south-east 
side of the arch to open out towards the south-west, and the north- 
west side to expand towards the north-east ; so that we have an 
almost complete section between Llangwyllog and Llanerchymedd 
as we have between Porth Nobla and Aberfiraw, but with the oppo- 
site dip. 

The large area occupied by the granitoidite may seem to require 
explanation. As this rock rarely exhibits bedding, its relations 
are frequently obscure; but, fortunately, in the district between 
Gwalchmai and Llechyn farwy the dark Holyhead schist, which 
immediately underlies it, is brought up by repeated undulations. 
Near Gwalchmai, for example, the granitoid rock dips north-west ; 
but to the west 1t is thrown off in opposite directions in the Craig- 
yr-allor anticline. The granitoidite is, then, a band of no great 
thickness, repeated by contortions. ‘There is also evidence of repe- 
tition by faults. 

The generalized section (fig. 1) represents the relations of the 
principal rock-groups in Anglesey. 


LV. NorrHern AREA. 


This district is bounded on the south by the curved fault which, 
starting from near Carmel’s Point on the west, passes round by way 
of Llanfilewin and Paris Mountain to the east coast at Porth-y- 
corwg. The rocks are very much contorted and broken; but there 
appears to be, on the whole, an ascending series from the fault to the 
north coast. The dip is generally to the north; but on the west it 
is to the north-east. 

1. Volcanic group of Paris Mountain.—The chief types of this 


922 C. CALLAWAY ON THE ARCH AN 


ridge are a felspathic breccia and a compact felspathic rock like 
hornstone; but both varieties are quite distinguishable from any 
other rocks in Anglesey. At the east end of the hill, by the Pen- 
sarn chapel, the breccia is well exposed. The fragments, some of 
which are a yard in diameter, are a very compact felstone; and the 
matrix is a sort of greenish ash. The rock, as a whole, reminded 
me of the compact breccias at Clegyr Bridge, St. David’s. Prof. 
Bonney has since examined a shde; and he is of opinion that it 
exhibits both perlitic and fragmental structure and is in all proba- 
bility a rhyolitic ash. The dip of the series is to the north, agreeing 
with that of the adjoining slaty group. About 100 yards to the east 
of the chapel, and therefore on the same strike, is a knoll composed 
of a pale-green and purple rock, very compact, almost like jasper. 
Under the microscope (Note 53, p. 236) it is seen to be a trachyte, 
probably from a lava-flow, and is, in Prof. Bonney’s judgment, as ~ 
modern in its appearance as the rock composing the enclosed pebbles 
near Bodorgan. The purple variety is not unlike some of the fiinty 
felstone common in the Precambrian of Shropshire. From litho- 
logical characters the Archean age of this group appears not 
improbable, though it must be conceded that it does not resemble 
very closely the volcanic group south of Bangor. Field work throws 
little ight upon the problem. On the north side the ridge is bounded 
by afault; and at the base of the southern escarpment black shales 
seem to pass conformably under the volcanic group. The two rocks 
appear within three or four feet of each other, dipping in the same 
direction into the steep face of the hill. There are, however, no 
signs of a passage between the black shales and the compact felspa- 
thic rock. The appearance of conformity might be produced by 
faulting, accompanied or followed by lateral pressure; and there are 
abundant proofs of both in the district ; but, under the circumstances, 
I hesitate to include this group in the slaty series. 

2. Chloritic Schists of Mynydd Mechell.—This group is the “ foli- 
ated grit” of Ramsay. In the field it has the appearance of a 
quartzose grit, with chlorite covering lamination surfaces ; but Prof. 
Bonney, after microscopic examination, regards it as a true schist 
(Note 45, p. 234) of the Holyhead type, though he admits that the 
chloritic constituent is ‘rather minute.” These rocks are much 
contorted, and are frequently penetrated by dykes of felstone and 
dolerite running with the strike. At a higher horizon, at Cas 
Clock and west of Rhos-y-pill, the rock is an undoubted schist with 
silvery lustre (Note 46, p. 234). Some of the strata north-east of 
Paris Mountain are not unlike this schist, as may be seen at Cerrig, 
on the Amlwch road. 

It is important to observe that Prof. Bonney agrees with me in - 
recognizing (Notes under C, p. 284) a difference between these 
schists and those of the gneissic districts ; but in such a broken 
country it is difficult to prove that these rocks should go with the 
slaty series. My reasons for so associating them are the following :— 

(1) The rocks are not uniformly foliated. To the west of Cas 
Clock, for example, chloritic schist is overlain in the same quarry 


GEOLOGY OF ANGLESEY. 995 


by hard grits. In the same neighbourhood are bands of hornstone, 
similar to ordinary Pebidian types, associated with schist. 

(2) These rocks appear to pass up through less altered varieties 
into slightly metamorphosed grits and shales, as seen south of 
Llanfechell and in localities to the east. 

(3) The alteration is not carried so far as is usual in the older 
series. 

(4) In the undoubted siaty districts, as west of Cerrig Ceimwen, 
the rock frequently undergoes partial metamorphism, and bands 
occur which might almost pass for true foliated schists. 

(5) This area is scored with intrusive dykes. I do not suppose 
that these are the cause of metamorphism; but they indicate the 
proximity of a source of heat, which may have been concerned in 
the change. 

3. Llanfechell Grits—The altered rocks of Mynydd Mechell appear 
to pass to the north into a series of green grits, well exposed round 
Llanfechell. This rock has undergone much alteration, to which 
probably the colour is due. Microscopic examination (Note 47, p. 234) 
brings to light the important result that it is “ almost certainly 
derived from the older gneissic and schist rocks of this region of 
North Wales.” These grits are also seen on the strike to the south- 
east near Bodewryd. 

4, Rhosbeirio Shales.—W est of Rhosbeirio church, and in a quarry 
near the farm, are very interesting exposures of felspathic rocks of 
the true St.-David’s type. These shales are soft, well laminated, 
and fine-grained; but here and there are thin seams of grit like the 
variety just described. While the body of the rock is but slightly 
altered, the gritty bands have a very schistose look. The prevailing 
colours are pale green and purple; but there are also some beds of a 
soft yellow shale, like a common variety in North-western Anglesey. 
The dip is to the north at 30°. Under this group, at Nant-y-cyntin, 
slaty beds of a more altered character, intermediate between these 
shales and the Cas-Clock schist, are seen. 

5. Sharply contorted Group south-cast of Amlwch.—At Crogan 
goch and on towards Llaneilian are exposed the remarkable rocks 
described and figured by Prof. Ramsay. The foliated structure (Note 
49, p. 234) of these slaty-lcoking beds is not very evident to the 
naked eye. Similar rocks compose the tongue-shaped promontory 
of Point A‘lianus; but sometimes contortion is wanting. At the 
base of the promontory the beds are gritty and partially altered 
(Note 48, p. 235). 

6. Amlwch Slates and Cemmaes Limestones.—South of Amlwch, 
near Crogan goch, the last group, with the intervention of an ashy 
band, is succeeded by uncontorted pale-green chloritic slaty beds 
(Note 50, p. 235), which are continued to the coast and all along the 
west side of Bull Bay to Ogo’ goch. The dip is steadily to the 
north at a moderate angle; and the thickness must be considerable. 
Towards the west, on Porth wen, bands of limestone and quartz- 
conglomerate come in; and further west, on Cemmaes Bay, limestone 
and quartz rock predominate over the slaty beds. I am not certain 


D4 C. CALLAWAY ON THE ARCH.BAN 


that the Amlwch and Cemmaes groups are on the same horizon ; 
but the evidence appears to point in that direction. The limestone 
(Note 52, p. 236) is very similar to a rock at Llanfaethlu; and the 
associated slaty beds are alike in both localities; but it would be 
rash to correlate the groups. 

The northern area (omitting Paris Mountain) is thus seen to 
consist of chloritic schists, felspatho-quartzose grits, felspathic shales, 
chloritic slates, quartz-conglomerates, and grey limestones, with some 
limestone bands on more than one horizon. In such a shattered 
district a true succession.can hardly be indicated. North of Paris 
Mountain some of the horizons are probably faulted out of sight, 
and it is difficult to correlate the rocks of this district with those 
further west. Most of the groups appear to pass into each other, 
either vertically or laterally ; and I cannot avoid the conclusion that 
they all belong to the same epoch. 


V. NortH-wESTERN AREA. 


In Western Anglesey are two well-marked groups of strata, which 
I regard as the equivalents respectively of the Gneissic and Slaty 
series. They are brought together by a fault. 

Porth-y-defaid Fault.—This dislocation is situated on the west 
coast, about midway between the northern and southern extremities. 
It 1s well seen on the shore, striking inland to the east. Greenstone 
is erupted along the line of junction, and alters the strata for a 
short distance on each side. South of the fault are thoroughly crys- 
talline quartzo-micaceous and chloritic schists dipping north-west ; 
while close at hand on the north side green ashy shales dip north- 
east. This fault is the boundary between the two series ; for true 
schists with a south-west strike occur everywhere to the south, 
while a comparatively unaltered group, with a prevailing northerly 
dip, stretches right up to the north-west corner of the island. 

Section from the Fault to Pen brywv Eglwys.—Proceeding north- 
ward, we find at Trefadog ashy shales dipping north-east. In the 
quarries north of Llanfaethlu church is a good section of grey lime- 
stone and pale-green ashy slate with easterly dip. The former is com- 
pact, and apparently as unaltered as any Paleozoic limestone. A 
careful search revealed no trace of fossils. The slaty rock is of a 
common Anglesey type; and the whole group is like the Cemmaes 
series. About Llanrhyddlad exposures are numerous. At Porth 
Swtan, on Church Bay, is a considerable thickness of yellowish fel- 
spathic slate and breccia. The dip is north-east, with slight contor- 
tion on a small scale. Higher beds are seen in quarries at Rhyd 
Ngharad; the rocks are brecciated in part and very felspathic. 
Greenish colours predominate. One variety, a pale-green slate, 
contains cubic pyrites. On about the same horizon, at Ogo Lowry, 
are green brecciated rocks more highly indurated. 

The district between Ogo Lowry and Penbryn’r Eglwys is one of 
the most faulted parts of the island ; the section consists of alterna- 
tions of Archean and Paleozoic rocks, repeated by faults, and dipping 
to the north. The wedges of dark shale let in amongst the older 
group are very clearly seen in the sea-cliffs. The younger shale, of 


GEOLOGY OF ANGLESEY. 225 


course, frequently appears to pass under the felspathic series. Such 
facts suggest great caution in trusting to dips in areas where the 
faulting is not clearly seen. The lithology here, as generally in 
Anglesey, is the true guide. The Precambrian chiefly consists of 
felspathic breccia. The fragments are usually of similar composition 
to the matrix; but some are of quartzite. The felspathic fragments 
weather out sharply where the cliffs are washed by the spray. On 
the whole this rock strongly suggests the breccia near Nun’s Well, 
St. David’s. Good exposures are seen at Yynys y fydllyn, Porth yr 
hwch, and west of Pant yr Kglwys. The thickness, owing to such 
frequent faulted repetitions, probably is not great. 

The section at Pen bryn’r Eglwys is of much interest. In this. 
great headland, which juts out at the north-west corner of Anglesey, 
are rocks which are described in the section of the Survey Map as 
‘‘ oneissic rocks pierced by granite veins ;” and these are represented 
as passing on the south into “ metamorphic foliated rocks.” The 
latter are the felspathic shales and breccias already noticed. The 
description of the former also requires modification. 

The breccias pass up into the southern slope of the headland, 
where they are succeeded by a band of quartzite. This is over- 
lain by a considerable thickness of greenish felspathic beds of ordinary 
Pebidian type and not much altered. ‘The highest ridge of the 
promontory consists of this rock. Following these strata across the 
strike to the north, they are seen gradually to change, the rock 
putting on a glazed aspect, and mica appearing in small quantities 
on the lamination-planes. This altered material soon passes into 
thoroughly foliated gneiss and granitoidite. 

It was rather startling to find true metamorphic rocks associated 
with comparatively unaltered felspathic beds of the newer series. 
The granitoidite is very similar to that of the older gneiss; and my first 
impulse was to refer it to that group. In sucha shattered district 
as Anglesey, especially in such an area as its north-west corner, a 
faulting-up of the older series seemed not unlikely. But, after careful 
examination, I found it impossible to accept this supposition. The 
‘transition between the unaltered rock and the schists is gradual and 
complete. <A close comparison furthermore revealed hthological dis- 
tinctions between the gneiss of the two series, the most important 
of which is a difference in the colour and lustre of the mica. 

The cause of metamorphism was not apparent. The “granite 
veins ” of the Survey are probably the granitoid bands in the gneiss. 
Some quartz veins running across the cliffs may have suggested the 
intrusion of granite. As this metamorphosed mass occurs in the 
neighbourhood of the Mynydd-Mechell schists, the alteration in both 
cases may be due to the same general cause. 


VI. Western AREA. 


All the rocks south of the Porth-y-defaid fault and west of the 
Paleozoic area belong to the older series. 

1. The Mainland.—This district is chiefly occupied by green 
chloritic schists (Note 37, p. 234) similar to the rocks described from 
the Menai anticline, Aberffraw, Craig yr allor, and east of Paris Moun- 

& 


226 C. CALLAWAY ON THE ARCHAAN 


tain. ‘The dip is usually to the south-east; but the beds frequently 
roll over to the north-west, as may be seen south of Porth twyn- 
mawr, and at Caer ceiliog on the Holyhead road. There is no doubt 
that these schists represent the dark green band which in the 
central zone underlies the granitoidite. 

2. Holyhead District.—Chloritic schists are well seen at many 
points in and round the town. <A typical specimen from Porth felin 
(Note 36, p. 234) is of the same character as the schist at Porth y 
defaid. The strikeisto thesouth-west; and undulations arenumerous. 
These rocks are separated from the quartz group by faults: one, on 
the south-west, runs from the south coast, at Porth y corwel, to the 
south end of Holyhead Mountain ; and another, on the north-west, 
hes along the south-eastern base of the mountain and cuts the coast 
at Porth yr Ogof. 

Porth-yr-Ogof fault.—Prof. Ramsay * states that the “ quartz 
rock” dips under the ‘“ fohated schists ;’ and he gives a section in 
support of his opinion. ‘This view would have greatly simplified 
my work ; but on visiting the spot I was disappointed to find that the 
two groups were brought together by a fault. There was a notch 
in the cliff; on one side was quartz rock, and on the other green 
schists (Note 35, p. 234), with no signs of a passage. The break 
was filled in with fallen rubbish. It was very fortunate that this 
dislocation was discovered, as it compelled me to seek the true 
succession elsewhere. 

Holyhead Mountain.—The microscopic examination (Note 31, 
p. 233) of a typical specimen of the quartz rock from the great quarry 
shows that it is a true quartz-schist of the same type as the Bodafon, 
Gwalchmai, and Mynydd-baen quartz groups. The structure of the 
mountain is interesting. Ascending the south-east slope, separation- 
planes are seen to dip to the north-west at about 80°; but on the 
north-west side the dip changes to south-east at the same angle. The 
planes are not close and even, as in slaty cleavage, but they cause 
the rock to split in large thick flat flakes. This difference is doubtless 
due to the coarseness of the material; and the planes must represent 
true cleavage. ‘The cleavage-strike agrees with the strikes of the 
contorted bedding and of the ridge. 

The Holyhead district is thus brought into clear comparison with 
other parts of Anglesey. The chlorite-schists undoubtedly repre- 
sent the dark schist which underlies the granitoidite in the central 
zone; and the quartz-schists may with the highest probability be 
placed on the same horizon as the foliated quartz rocks of Bodafon 
and Gwalchmai. The absence of the grey gneiss is easily explained. 
On both the west and south the junction between the quartz and 
chloritic groups is a fault; but the two formations dip in the same 
direction, and the downthrow is apparently on the side of the chlo- 
rite-schist. As the latter group is repeated in numerous shallow 
curves, there is no reason why the grey gneiss should appear at the 
surface, as it does amidst the deeper undulations of the Menai anti- 

line. 
* Geology of North Wales, p. 185. 


4, 


GEOLOGY OF ANGLESEY. QF. 


The chloritic rocks of this area are rather more quartzose than in 
most Anglesey localities; but they resemble some of the varieties in 
Mynydd Liwydiarth, and there are indications in both districts 
that a source of quartz-derivation was not far distant. 

3. Rhoscolyn District.—The south-eastern half of Holyhead Isiand 
is similar in structure to the Holyhead area; but the rocks are 
affected by an upheaving force to the south, so that the prevailing 
dip is to the north. The green schists, which occupy the chief part, 
need no further description. ‘The southern fault, which separates 
the quartz and chloritic groups south of Holyhead, passes under the 
sea, and holds the same relation near Khoscolyn. It is first seen 
near Bwa du, and again appears on the coast tothe south-east at Borth 
wen, passing in its course near Rhoscolyn church, where the quartz- 
schist (No. 32, p. 233) is exposed, dipping northerly. A few yards to 
the north the chloritic group dips in the same direction, at about the 
same angle. At Bwa du the junction is well seen, both series dip- 
ping northerly, the green schist at a lower angle than the other. 
The quartz group forms an elevation, Mynydd Rhoscolyn, as is usual 
in Anglesey. 

At Borth Saint, south of Bwa du, the contortion is intense. There 
is evidence, from the folding of quartz veins, that the rock has been 
squeezed to one fourth of its original bulk. At one spot there is 
a very curious specimen of ripple-mark, distorted so as to be almost 
unrecognizable. Quartzite dips to the north, and is underlain by 
soft schist. The under surface of the former overhangs, and is 
covered with rounded projections, which are evidently the squeezed 
easts of ripple-marks. The schist was the mud which received the 
impressions of the ripples; and the quartzite represents the sand 
which covered in and preserved the marks. I have elsewhere* 
figured and described the above phenomenon. It is important to 
record ripple-marks in so ancient a formation. 

South of the last locality is a considerable thickness of chloritic 
quartz-schist (Note 33, p. 233). 


B. Summary or REsvutts. 


I. Distrrpurion or tHE Rocks. 
[See Map (Pl. VIII.) and Fig. 1.] 
a. Gneessic Serres. 

1. Geographical Distributcon—This series occupies the greater 
part of the Menai anticline. It reappears west of the Llangefni 
slaty area, and forms the whole of the central zone, except the 
faulted mass south-west of Bodafon Mountain. It then reemerges 
west of the Paleozoic, and covers Western Anglesey south of the 
Porth-y-defaid fault. About three fifths of the area coloured 
“‘ altered Cambrian and Silurian” consists of eneissic and schistose 
rocks. 

* ‘Science for All,’ part xxxi. pp. 205 and 205. 


228 C. CALLAWAY ON THE ARCHAAN 


The hdlleflinta forms a band extending from Porth Nobla to 
near Gwalchmai. It is probably represented north of Llangwyllog. 

The quartz-schist extends on the strike from Mynydd baen to 
Gwalchmai. It may occur in its place north of Llangwyllog; but 
it emerges in the craggy mass of Bodafon Mountain and in the 
small inlier to the east. In Western Anglesey it forms the eleva- 
tions of Holyhead Mountain and Mynydd Rhoscolyn. 

The grey gneiss is brought up through the dark schist on the 
western side of the Menai anticline at Gaerwen and east of Pentraeth. 
In central Anglesey it runs from near Porth Trecastell to north- 
east of Bodwrog, reappearing soon after on the opposite side of the 
anticline north of Llangwyllog, and further to the west in the Craig- 
yr-allor anticline. ‘To the north-east it is seen in the area south- 
east of Paris Mountain. 

The dark schist oceupies the greater part of the Menai anticline. 
It emerges west of the Llangefni “‘ syncline,” but is gradually faulted 
out towards Gwalchmai. On the opposite side of the central anti- 
cline it forms a band as far to the north-east as Plas Llanfihangel, 
and is brought up through the granitoidite in the oval dome of Craig 
yr allor. Italso occurs south-east of Paris Mountain. In Western 
Anglesey it constitutes the schistose area south of the Porth-y-defaid 
fault and north of the quartzose masses. 

The granitoidite is the “ granite” band of the Survey. The details 
are inaccurately laid down in their map at many points; but little 
theoretical importance attaches to these errors. There are slight 
traces of this rock on the west side of the Menai anticline. 

2. Stratigraphical Distribution.—One of the most difficult pro- 
blems in Anglesey was to make out the succession of the subdivisions 
of this group. In no single section, owing to the faulting, is the 
series entire; but the subdivisions are recognizable by their litho- 
logical characters; and by comparing localities the fragments 
are pieced together into a complete succession. The facts will be 
best expressed in the following table, the groups being taken in de- 
scending order :— 


| | | 
| j . | 
| 1. Holy-| 2. Craig 13. Coed-}4. Dulas} 5. Ty 6. Menai 
| : cr ee : Croes, |O- Menai. 

head. | yr allor.| ana. Bay. roes. 

| | 
| Granitoidite ...!... % * | x iy Traces. | 
| Dark schist ... * * x * * x ; 
Greysomeiss i: mie * x * % * | 
Quartz-schist ... x a ? as 
Halleflinta ... wee ie ? vee * : 


The star indicates the existence of the subdivision in the locality 
at the head of the column. By comparing 5 with 2, 3, and 4, it 
will be seen that two groups are common to the four areas; and 
they serve to link together the lowest and highest subdivisions. 


GEOLOGY OF ANGLESEY. 229 


The series is probably complete in 3; but in running the section L 
could get no exposure between an obscure opening of hiilleflinta and 
the grey gneiss, and I had not time to search beyond the highroad. 


b. Slaty Series. 


This group occupies four areas, viz. the small mass south- 
west of Pentraeth, the east side of the Llangefni “syncline,” northern 
_ Anglesey, and the western division north of the Porth-y-defaid 
fault. The chief facts are thus summarized :— 

(1) South west of Pentraeth (Wugan-bach ridge). 

Slaty and dolomitic rocks. 

(2) Llangefni district. 

Slates (some hypometamorphic*), grits, green conglomerates, 
purple and green shales, hornstone, calcareous beds. 

(3) Northern district. 

Chloritic schists, altered grits, green and purple shales, slates 
(some hypometamorphic), quartz-conglomerates, limestones, 
hornstone. 

(4) North-western district. 

Yellow felspathic shales and breccias predominating, quartz- 
conglomerates, pale-green slates, and limestones. 

J am not prepared to submit ascheme of the succession. The rocks 
of area | are not unlike some of those near Cerrig Ceimwen in area 
2. Comparing 2 and 3, the rocks are similar as a whole; but the 
green conglomerate, so conspicuous in 2, is not seen in 3, while the 
quartz-conglomerates and grey limestone of 3 do not appear in 2. 
Portions of 3 are also altered to schist, while in 2 the transition 
into foliated rock is rarely complete. The felspathic breccias of 4 
do not certainly occur in any of the other areas; but its quartz- 
conglomerates, slates, and limestones are similar to those of 3. The 
interesting Rhosbeirio shales in 3 have less complete representatives 

n 2, but are fully exhibited in 4. 

The rocks of these four areas, though geographically isolated, are 
closely connected by their mineral and petrological characters, and 
I have no hesitation in placing them all in the same group. Should 
the chloritic schists of the north be separated from the newer series, 
it will not materially affect my general conclusions. The same 
remark applies to the small patch of gneissic rock at Bryn ’r Eglwys. 


II. Evipencr or AGE. 


a. Relations to Paleozoic Groups. 


Gneissic Series.—It is not necessary to recapitulate the evidence 
for Precambrian age. The sections in the quarries near Nebo would 
alone be sufficient to remove’ all doubt. 

Slaty Series—These rocks do not clearly underlie any part of the 
Cambrian ; they are, indeed, overlain by a purple conglomerate on 
the north coast, west of Tor Ilwyn. The pebbles of this rock are a 

* The term “submetamorphic,” used by some authors, is objectionable on 
grammatical grounds. 


Q.J.G.S. No. 146. age 


930 C. CALLAWAY ON THE ARCH EAN 


purple quartzo-felspathic grit; but I have no certain proof of its 
age. 

"The Precambrian age of the series may be inferred from included 
fragments. The rocks west of the granitoid zone are shown by the 
fossils collected by Prof. Hughes to be Tremadoc and Arenig. In 
the conglomerates west of Llanfaelog, which appear to be of at 
least equal antiquity, a large proportion of the pebbles are certainly 
derived from the Slaty series, pale green and purple hornstone and 
slate being the most common. It is also worthy of notice that 
the Harlech conglomerates of Caernarvonshire contain fragments 
of green and purple slate *, which must therefore be Precambrian 
and of no distant source of derivation. 

The relations of these series to the Cambrian rocks in contact 
indicate the same result. At numerous points the two groups are 
brought together by faults; but while the Slaty series is generally 
more or less altered, the Cambrian shales are quite unchanged, and 
Lhave never observed the slightest indications of a passage betweer 
the two, the line of demarcation being always clear and sharp. The 
Slaty series must therefore have undergone partial metamorphism 
before the Cambrian period, otherwise it is difficult to see how the 
Cambrian rocks could have remained unaltered. 

The black shales in some localities dip as if they would pass under 
the altered series; but at many points the latter also appear to pass 
beneath the former. In such a shattered district this evidentey is 
worth nothing in either case, unless an “unbroken sequence can be 
proved. 

I am not prepared absolutely to deny that true Harlech rocks 
occur in Anglesey. There are beds, especially near Llangristiolus, 
which lithologically do not appear to be. very distinctively Pre- 
cambrian ; but, as stratigraphically they are closely associated with 
the Slaty series, it is safer to place them in that group. 


b. Relations to each other. 


In all parts of Anglesey the junction of the two series, whenever 
they are in contact, isafault. In the east and centre they generally 
strike in the same direction; but in the north-east (near Paris 
Mountain), at the base of Bodafon Mountain, and. inthe western 
district, as seen north and south of the Porth-y. -defaid fault, the 
strikes are discordant, sometimes approaching a right angle. The 
much more intense metamorphism of the gneissic series would seem 
to point to a greater antiquity. Some additional light is thrown 
upon the subject by the microscope. Prof. Bonney, in note 47 (p. 234), 
already noticed, is decidedly of opinion that the Llanfechell grit, a 
characteristic variety of the newer rocks, is derived from the older 
series. 

c. Relations to other Areas. 


(1) Caernarvonshire. 


Comparing the Slaty series with the Bangor group, the re~ 
* Bonney, Quart. Journ. Geol. Soc. vol. xxxv. p. 511. 


GEOLOGY OF ANGLESEY. 31. 


semblance, it must be confessed, is not very close. Both formations 
are largely of volcanic origin; but the felspathic materials of the 
Anglesey rocks are derived from a more distant source, while the 
frequency of quartzite pebbles indicates the proximity of land not 
covered by volcanic products. 

The Gneissic series would appear to be closely related to the 
granitoid rock of Twt Hill. As the normal position of the dark 
schists is below the granitoidite, there seems to be no reason to 
doubt that the green schists on the Anglesey side of the Menai 
Straits pass conformably beneath the Twt-Hill beds. The post- 
Carboniferous fault on the west side of the T’wt-Hill ridge might 
not materially affect the true relations of the older subdivisions. 


(2) St. David's. 


_ Slaty Series.—The lithological resemblances between this group 
and the St. David’s Pebidian are numerous and striking. The 
varieties whose Pebidian facies is most marked are the pale- 
green, purple, and yellow felspathic shales of Rhosbeirio, the 
felspathic breccias north of Llanrhyddlad and east of Bodorgan 
station, the tough green rock at Cerrig Ceinwen and Cerrig-engan 
fawr, porcellanitic bands near Llangristiolus, and hornstones of 
many localities. The slaty beds are less distinctively of St. David’s 
type. 

Fe ncicsic Series—The Anglesey granitoidite has been compared 
by Messrs. Bonney and Hicks with the St. David’s Dimetian. If 
this correlation be conceded, the entire Anglesey series, from the 
hilleflinta upwards, must go with the Dimetian. But it must be 
admitted that the two.groups present important differences. At St. 
David’s the Dimetian is represented by a great thickness of grani- 
toidite and quartzite ; whereas in Anglesey the similar band is much 
thinner, and is underlain by a considerable thickness of gneiss and 
other schists. 


(5) Shropshire. 


The felspathic slaty beds of Lileshall Hill are hardly distin- 
guishable from some of the Anglesey rocks. The Lilleshall group 
is probably higher than the Wrekin volcanic rocks, which have 
many points of resemblance to the fragmenta! group south of Bangor. 
The purplish ash of the Wrekin, for example, is very similar to the 
srit at Bryn Ilwyd. Nothing certain, of course, can be inferred 
from such facts; but it is at least worthy of consideration, and the 
suggestion may be a guide in further inquiry, whether the Anglesey 
Pebidian may not represent a higher horizon than the Bangor group. 
Some of the gritty and slaty rocks of the Precambrian ridges east 
of Church Stretton also display affinities with the Anglesey series. 

It is worthy of note that the Charlton-Hill conglomerate, an 
undoubted part of the Wrekin group, contains, amongst enclosures 
from the Malvern series, pebbles of rocks which more closely 
resemble some of the gneissic and quartzose types of Anglesey. 

k2 


232, PROF, T. G. BONNEY ON THE MICROSCOPIC 


(4) Charnwood. 


The slaty rocks of Anglesey are nearer in their lithological 
characters to the Charnwood slates than to any other formation with 
which I am acquainted. The resemblance was very marked even 
in the field ; but in the case of a homogeneous rock like slate, the 
microscope is of special value, and Prof. Bonney’s examinations 
(Notes 42, 43, 44, pp. 234, 235) are strongly confirmatory of my 
opinion. 

CoNCLUSIONS. 

1. In Anglesey there are two Archean groups, the Slaty and the 
Gneissic. 

2, The Slaty series is composed of slates, shales, hornstones, grits, 
conglomerates, limestones, and chloritic schists, in which no definite 
order has been positively ascertained. The Gneissic series is divided 
into five groups, in which the following descending order is in- 
variable, viz. granitoidite, chloritic and hornblendic schists, grey 
gneiss, quartz-schist, and hilleflinta. 

3. The Slaty series is occasionally foliated, but is usually in a 
partially altered state: the Gneissic group is thoroughly meta- 
morphosed. 

4, The Slaty series has closer lithological affinities with the St.- 
David’s voleanic group, the Charnwood rocks, and the Lilleshall 
series than with the Bangor group. 

5. The Slaty series is undoubtedly Pebidian: the Gneissic series 
may with some probability be referred to the Dimetian. 


APPENDIX. 


Notes on the Microscopic Structure of some ANGLESEY Rocks. 
By Prof. T. G. Bonnny, M.A., F.R.S., Sec. G.S. 


In the following notes I shall endeavour to avoid repetition by 
grouping together those specimens which in some important cha- 
racteristics have a general resemblance one to another. I may 
add that, as the purpose of my examination was petrological rather 
than mineralogical, I have not felt bound to spend. much time in 
endeavouring to ascertain the exact species of some microlithic 
minerals which are accidentally present in one or two of the slides. 


A. Quartz-Schist Group. 

Nos. 25-29, 31-34. 

This group consists of a series of schistose rocks, in general highly 
metamorphosed, in which quartz is the most important constituent, 
associated with micaceous or chloritic minerals, ferrite or opacite, 
and more or less felspar. 

The fragmental origin of most of these specimens is indubitable ; 
but with regard to numbers 25 and 27 it is less easy to be quite 
sure. Their structure comes nearer to that of some microcrystal- 
line felsites ; but still I believe that I am right in ranking these also 
among the altered clastic rocks. 

25. (Gwalchmai, p. 218.) Contains numerous grains of quartz of 


STRUCTURE OF SOME ANGLESEY ROCKS. 233 


rather irregular outline in a finely granular and rather earthy-looking 
matrix—most probably the result, in the main, of the decomposition 
of rather imperfectly crystallized felspar, which consists now of 
an admixture of earthy dust, doubly refracting clear microliths, 
and minute scales of mica or chlorite. The quartz grains show 
an indistinct banded order, and contain many minute cavities. The 
rock appears to have been brecciated more than once zn situ. The 
older cracks are mainly filled up by quartz, the newer by calcite 
with a little chlorite. In one part of the slide the faulting of an 
older vein and subsequent infiltration of a newer one is very 
clearly shown. Macroscopically the rock has aresemblance to the 
Treffgarn halleflinta, but under the microscope it is more gneissic. 

26. (Gwalchmai, p.218.) Is certainly of fragmental origin, show- 
ing stratification foliation. The quartz grains have fewer inclusions; 
there is felspar much as before, and a fair quantity of mica; most of 
this is of a very pale golden-yellow colour, is feebly dichroic, and 
gives brilliant tints between the Nicols; the rest is dull green, 
probably an alteration produced after biotite. Some granular matter 
as above. 

27. (N.W. of Gwalchmai, p. 218.) More resembles 25, but is more 
compact. I think that it, too, is of fragmental origin, and is pro- 
bably the result of the metamorphosis of a fine quartzose silt. 

28. (N.W. of Gwalchmai, p. 218.) Is certainly fragmental, con- 
sisting of quartz with a fair amount of chlorite, some white mica, 
and a few grains (probably fragmental) of a closely twinned plagio- 
clastic felspar. 

29. (Summit of Bodafon Mountain, p. 219.) Is a quartz-schist 
with a fair amount of ferrite and minute chlorite (?). 

31. (Holyhead Mountain, p. 226.) Also a quartz-schist with a fair 
amount of a pale-green mi¢a, as above, the ground-mass consisting of 
this mineral with agglutinated very minute grains of quartz. In this 
are scattered quartz grains of larger size up to about 003 diameter, 
clearly of fragmental origin, some having a secondary deposit of 
quartz on their edges. Minute cavities are common in these grains, 
and one or two contain some dark hair-like microliths, a grain or two 
of epidote, and possibly one of tourmaline. 

32. (Rhoscolyn church, p. 227.) Also a quartz-schist resembling 
the last, but more uniform in structure; the same adventitious 
minerals as in the last. 

33. (S. of Borth Saint, p. 227.) Another quartz-schist with rather 
more mica and chlorite, a few grains of felspar, and the same adven- 
titious minerals. Numerous grains of a granular earthy mineral 
(? an epidote). 

34. (Cutting EK. of Ty Croes, p. 217.) Quartz and a while mica 
are the most conspicuous minerals, with an occasional grain of 
felspar; but a sort of granular paste in which these are imbedded, 
consisting of microliths of white mica and other minerals, may in 
some cases have replaced a felspar constituent. The rock is much 
altered, but, I suspect, still retains in its large grains traces of its 
original fragmental structure. 


234 PROF, T. G. BONNEY ON THE MICROSCOPIC 


B. Chlorite-Schists. 


A group of highly altered, distinctly foliated rocks, consisting 
mainly of rather minutely crystalline chlorite and quartz. To this 
belong Nos. 35-37, 45, 46, 49. 

35. (Porth yr Ogof, Holyhead, p. 226.) Contains very numerous 
rods and granules of a black mineral, perhaps magnetite, with a little 
sphene or staurolite; it exhibits beautiful contortions. 

36. (Porth felin, Holyhead, p. 226.) Foliation less marked ; a con- 
siderable amount of ferrite is present. Original structure perhaps 
less uniform than in the last. 

37. (Porth y defaid, p.225.) The same remark applies to this speci- 
men also, but itsfoliation is moremarked. A vein of quartz and calcite. 


rather minute. 
46. (Cas Clock, p. 222.) Highly altered and markedly foliated. 
49, (Crogan goch, p. 223.) The same; chloritic or possibly mica- 
ceous constituent extremely minute; beautifully foliated and contorted. 
In this group, as a rule, it is more difficult to identify with 
probability any original constituent of the rock, owing probably to 
differences in chemical composition and size of the materials. Some 
of the quartz grains in 37, 45, 46 may, however, be original. 


C. Slaty and other not highly altered Rocks. 


In this group there has, indeed, been a certain amount of altera- 
tion, but it is only what may be termed micromineralogical. The 
original fragmental character of the rock is still distinct ; the newer 
products are either extremely minute or such as result rather from 
decomposition than from recomposition. The alteration, in short, is 
of a kind which we commonly meet with in the earlier Cambrian or 
latest Precambrian rocks, not in the gneisses, in the conspicuously 
foliated schists, and in the most highly metamorphosed quartzites. Its 
members, then, are, as a rule, either much more modern than those 
previously described, or selective metamorphism must have operated 
upon the latter to a rather unusual extent. 

43. (S.E. of Cerrig ddwyffordd, pp. 214, 232.) One of the flinty 
argillites or indurated imperfectly cleaved slates. Under the micro- 
scope the slide consists of a brownish-grey earthy-looking material 
interspersed with minute specks of quartz, felspar, and micaceous or 
chloritic minerals, whether original constituents or of secondary origin 
it is in many cases difficult to say. The “stripe,” however, is well 
indicated, and the specimen in no important respect differs from 
many examples that I have examined from Charnwood and other 
localities where the rocks are approximately of Cambrian age. 

44, (W.S.W. of Cerrig Ceinwen, pp. 215, 232.) A similar rock 
of slightly coarser material and a little more distinctly cleaved. 
Among the constituents are many scales of a strongly dichroic green 
mineral with a marked wavy cleavage—chlorite or some member 
of the provisional viridite group, probably the result of alteration 
of a magnesia-iron mica fragmentally present. 

47. (Llanfechell, p. 223.) A coarser fragmental rock with a rather 


STRUCTURE OF SOME ANGLESEY ROCKS. 239 


«streaky ” structure, not unlike some of those which occur in the 
Borrowdale series, containing numerous microliths of the viridite 
group, some being certainly chlorite. The aspect of the rock suggests 
that it has undergone considerable pressure. Many of the imbedded 
fragments are from about 0:03 to 0-06 in longer diameter. Among 
them quartz, felspar, altered biotite (?), and a chloritic quartz-schist 
may be recognized, detrital materials almost certainly derived from 
the older gneissic and schist rocks of this region of North Wales. 
Other fragments of a less certain character are present, with grains 
of decomposed ilmenite or magnetite and of epidote, which perhaps 
has replaced some other mineral. 

A8. (Base of Point Athanus, p. 223.) A rock of a generally similar 
character ; but, as the fragments are smaller, their nature (except in 
the case of the quartz) is less easily ascertained ; the whole rock also 
seems slightly more altered than in the last case. 

50. (Coast N.W. of Amlwch, p. 223.) The materials are more 
homogeneous than in the other cases; small greyish clustered 
eranules frequent, and a vast number of microliths of a green 
mineral (chlorite?). These are very likely of secondary origin, but 
I consider the rock a true slate, and not a schist. 

AD, (S.E. of Cerrig ddwyffordd, p. 214.) A fine grit with much 
of the fibrous green (? hornblendic) mineral which I have often ob- 
served in the Charnwood rocks developed among the finer materials; 
the larger fragments (generally rather angular) are chiefly quartz, 
felspar (orthoclase and plagioclase), and a minutely microcrystalline 
or cryptocrystalline rock, much resembling bits of an acid lava. 
From certain minute peculiarities, I have strong suspicions that all 
three of these constituents have been derived from rocks of volcanic 
origin. 

38. (Fragment in Llangefni conglomerate, p. 214.) A grit, consist- 
ing of subangular and rolled grains, mostly quartz, in a fine quartzose 
or quartzo-felspathic matrix. It is difficult in the case of rocks of this 
character to draw an inference as to their age and amount of meta- 
morphism; but I believe that I am right in grouping it Tae the 
probably more modern series. 


D. Granitoid Gneiss. 


Of this rock there is one specimen from Pen Bryn yr KEglwys not 
at all in a favourable condition for microscopic examination. It 
resembles the group of coarse granitoid gneisses similar to those 
which I have examined from the neighbourhood of Llanfaelog 
rather than a true igneous rock. It consists of quartz with many 
minute enclosures, two felspars much decomposed, and various 
alteration-products probably replacing an iron-magnesia mica. One 
or two grains of another mineral are present, obviously much 
altered, which I cannot identify; possibly it may have been garnet. 


KK. Crystalline Limestones. 


40, (H.N.E. of Tan y graig, Pentraeth, p. 213.) A finally granular 
clear rock consisting almost wholly of calcite or dolomite. Judg- 


236 PROF, T. G, BONNEY ON THE MICROSCOPIC 


ing from the structure and optical characters, I should infer the 
presence of a considerable quantity of the latter mineral, and so 
name the rock a dolomite. 

51. (E. of Bodwrog, p. 218.) Is more coarsely crystalline with an 
admixture of quartz grains and some silicate, which I am unable to 
identify with certainty ; the carbonate, however, appears tc be mainly 
calcite. 

Al, (At “a” of “ Pentraeth,” p. 213.) To the unaided eye appears 
less crystalline and less pure, resembling some limestones which have 
an admixture of volcanic mud. Under the microscope the calcite 
appears in rather irregular grains of fair size and rather brecciated 
aspect, imbedded in a streaky-looking paste composed of a pale green 
serpentinous mineral, of black and dark brown dust, with a little 
quartz, and of a clear silicate in minute granules, probably some 
kind of zeolite. 

52. (Tau isa, N.H. of Cemmaes, p. 224.) Is moderately clear, 
chiefly composed of very minute granules of crystalline calcite and 
possibly dolomite, with occasional brown stains, and veined irre- 
eularly with more coarsely crystalline calcite. In no one specimen 
can I detect any distinct trace of an organism; but the first two 
rocks differ much from the last two, which might readily pass for 
members of the Carboniferous Limestone series. 


F, Igneous Rocks. 


39. (Fragment in green shale, EK. of Bodorgan station, p. 214.) The 
ground-mass is thickly crowded with small and not very sharply 
defined felspar crystals, which, in their elongated form and the 
twinning of larger specimens, generally resemble members of the 
plagioclase group, interspersed with numerous minute grains of a 
pale green colour, of which the larger are distinctly dichroic and are 
probably hornblende. Small grains and occasional rod-like micro- 
liths of an iron peroxide are also present. Scattered about in the 
ground-mass are roundish patches of tessellated aspect, chiefly 
composed of crystalline quartz; one, in which the crystals are larger 
and by their enclosures indicate lines of growth, has calcite at the 
centre; this, however, as in the case of a neighbouring vein, is 
probably a subsequent infiltration. The majority more resemble 
imperfectly formed spherulites; a portion of the slide shows a 
distinct fluidal structure, and the ground-mass has a general resem- 
blance to several modern lavas, ¢. g. to specimens in my collection 
from Astroni and the Solfatara (Phlegrzean Fields). 

53. (Carreg-winllan, Pensarn, 8.H.of Amlwch, p. 222.) Exhibits a 
rather clear base, in which numerous small roundish patches are 
defined by exceedingly minute opacite and ferrite, the latter some- 
times forming the inner edge of the boundary ; it is also more or less 
powdered about through the base. With crossing Nicols this last is 
seen to be crowded with rather acicular felspar microliths, and the 
majority of the patches are well-defined spherulites with the usual 
radial structure. The rock is undoubtedly a trachyte, like the 
other; and, I think it highly probable that each is from a lava-flow. 


STRUCTURE OF SOME ANGLESEY ROCKS. Dis ff 


Considering the locality and the consequent age of these specimens, 
their structure is most interesting. So far as my experience goes, 
there is nothing in either which would have awakened my sus- 
picions as to their age, had they been labelled as from some locality 
where Tertiary or even more recent trachytes are found. 


EXPLANATION OF PLATE VIII. 


. Map of Anglesey, showing the distribution of the Paleozoic, Pebidian, 
and Gneissic series. 
. Generalized section from Holyhead Mountain to Menai Straits. 
. Section across Mynydd Liwydiarth. 
. Section from Porth Nobla to Aberffraw. 
- Section in Upper Quarry, Nebo. 
. Section in Lower Quarry, Nebo. 
In all the sections the signification of the letters is as follows :~- 
d. Carboniferous. 
c. Cambrian. 
BG Pebidian. 
,- Granitoidite. 
;- Dark schist. 
a, . Grey gneiss. 
Limestone. 
Quartz schist. 
. Halleflinta. 
. Faults. 


Fig. 


o> Or 09 bO am 


2,8 


“We Ny v2 


| 
_ a, Gneissic series ja 
| 
| 
a, 


Q.8 


Discussion. 


Dr. Hicks agreed with Dr. Callaway that there are two Pre- 
cambrian series in Anglesey. He differed from the author, how- 
ever, in regarding the so-called granitoidite as constituting the 
lowest and not the highest member of the so-called Gneiss 
series. He stated that some of the breccias associated with the 
halleflintas contain pebbles of the granitoid rocks, and are therefore 
of younger age than the latter. He admitted, however, that some 
of the points must be regarded as in an unsettled state, owing to 
the faulted condition of the district. 

Prof. Ramsay argued against the principle of identifying rocks as 
of different ages by their mineral characters as studied by the micro- 
scope. He maintained that the altered rocks of Anglesey are the 
metamorphosed representatives of the Cambrian, because the un- 
altered Cambrian are found striking directly towards the altered 
strata, and both are overlain by the Arenig. 

Mr. W. W. Smyrtu also argued against the recognition of a number 
of different formations on mineral evidence alone without any aid 
from organic remains. He thought the so-called “ gneissose rocks ” 
differ widely from typical gneisses, and that the granitic series 
belonged to the class of ill-defined granite rocks known in Corn- 
wall as “bastard granite,” the plain Saxon of which was perhaps 
preferable to the cacophony of “ granitoidite.” He thought that the 
evidence brought forward was insufficient to upset the detailed map 
of the Survey. 

Prof. Bonnzy was inclined to agree with Dr. Callaway’s interpre- 


238 ON THE GEOLOGY OF ANGLESEY. 


tation rather than that of Dr. Hicks, as the latter undervalued the 
amount of metamorphism the Anglesey rocks by the Menai Straits 
had undergone. He differed from Prof. Ramsay as to the value 
to be attached to the microscopic study of rocks; he did not regard 
the Bangor rocks as partially metamorphosed, and he found no 
trace of the progressive metamorphism insisted on by Prof. Ramsay. 
He defended the use of the term “ granitoidite,” which, though 
open to objections, was less misleading than the base-born term 
proposed by Mr. Smyth. He thought that the microscopic study of 
the rocks supported Dr. Callaway’s conclusions. 

Dr. Cattaway agreed with Prof. Ramsay as to the great value of 
the Survey work. He contended that the principle of the identi- 
fication of rocks by their mineral characters is a safe one when 
applied in the same district. The sections described proved that the 
granitoidite was the summit of the gneissic series. Many of the 
gneissose rocks were true gneiss of very typical character. The 
difficulty through faulting was counteracted by comparing areas. 


ON THE LIMESTONE OF DURNESS AND ASSYNT, 239 


16. The Limzstone of Durnuss and Assynr. By C. Cattaway, Esq., 
M.A., D.Sc, F.G.S. (Read January 5, 1881.) 


Iv is well known that the late Sir R. I. Murchison based his deter- 
mination of the Ordovician (Lower Silurian) age of the greater part 
of the Scottish highlands upon the discovery by Mr. C. Peach of 
Ordovician fossils in the Durness limestone. The views of Mur- 
chison were shared by many distinguished geologists, and have been 
generally accepted as one of the most assured and important conclu- 
sions of modern geological science. Wishing to ascertain for myself 
the truth of this opinion, I devoted a portion of last summer to the 
study of the limestone at Durness and Inchnadamff. I selected these 
localities because they alone are alleged to have yielded fossils from 
the limestone, and because Murchison regarded them as of primary 
importance in the construction of his argument. My researches led 
me to the conclusion, not only that the sections were broken, and 
therefore untrustworthy, but that the relations of the several rock- 
groups were inconsistent with the supposition that the limestone 
passed below any part of the newer metamorphic series. On my 
return from Sutherland, I found that many of my observations 
agreed very closely with those of Prof. Nicol; and I am glad to 
bear testimony to the accuracy of an author whose work has hardly 
received due recognition. It is certain that he made out many im- 
portant points which escaped the observation of his distinguished 
rival; and I must, in candour, acknowledge that he has anticipated 
many of the results which I have now to submit to the Society. 
I can only claim to have ascertained some additional facts, which 
seem to me to strengthen the case against the received opinion. 


DuRNEss. 


a. The Limestone in Relation to the Flagqqy Gnevss.—In his later 
papers, Murchison admitted that the limestone was separated from 
the flaggy beds* by a “ great fault,” by which the “ upper quartzite ”’ 

was thrown down out of sight; but he, notwithstanding, held that 
the limestone was ‘“ overlain by the upper series of quartzose and 
gneissic rocks,” apparently on the ground that both dipped in the 
same direction, to the east. Admitting his facts, his reasoning is 
obviously fallacious, and might be employed to prove the most op- 
posite conclusions. But I cannot accept his facts. Neither the 
limestone nor the flagey group dips to the east. The true relations 
of the two formations are shown in the annexed plan (fig. 1), which 
is, in outline, a reduced copy of the new ordnance map. The most 
important dips are shown by arrows. 


* These flags are a true schist, though they are far less coarsely crystalline 
than the Lewisian. 


240 C. CALLAWAY ON THE LIMESTONE 


Fig. 1.—Sketch Map of the Durness Area. 
(Scale, about ? inch to 1 mile.) 


Mh. 
vy NW “ h ‘ B 
WwW WNW ees CET TTN 


== lauartaite. ie 
Koiitis Flagey Gneiss. Limestone. EEE] Blown Sand. f. Faults. 


The fault f#, runs from sea to sea, east and west, exactly at the 
base of Far-out Head, which is entirely composed of thin-bedded 
gneiss and quartz-schist, dipping steadily to the north-east, except 
at some points a mile north of the fault, where the dip is conse- 
quently unimportant. 

West of the ruined church, where the limestone contains numerous 
fossils, the dip is north-east, so that 1t might seem as if it passed 
beneath the schist. But here the nearest exposure of the latter is 
over half a mile to the north-east, the intervening area being occu- 
pied by blown sand, so that no junction can be seen. Further ex- 
amination proves that this dip of the limestone is exceptional. 

Following the strike of the schist to the south-east, we trace it 
right up to the fault. Both rocks are here clearly exposed, the flags 
on the shore and in the adjoining field, and the limestone in a low 
cliff, which is an inland extension to the west of the precipice of 
Creag Chearbach. The fault runs along the base of the cliff. The 
flags dip uniformly to the north-east, the nearest section being only 
twenty yards from the limestone, which, at this point, dips east- 


| 


22 S8050- Bay Gneiss. 


OF DURNESS AND ASSYNT, Q41 


south-east. Several dips taken in this locality were the same, others 
veered round to the east, but none to the north of east. 

Following the cliff along to the west, and noting the prevalenco 
of south-easterly dips, we come, at about two furlongs east of Baile- 
nacille, to a small arch of limestone, with dips ranging from south- 
east to north-east. Such local variations are obviously unimpor- 
tant. In the area south of the fault the dips are almost uniformly 
to the south-east, as may be seen at many points between the inn 
and Sangomor. 

Coming back to the west of the old church at Bailenacille, we find 
the north-east dip soon changes to east, and then to east-south-east, 
and on the Kyle of Durness the limestone forms an escarpment over 
a mile long, striking south-south-west and dipping clearly to the 
east-south-east. At many other points in the area round Durness 
the same dip is seen. 

It is then certain that the mass of limestone lying south of the 
flags, and in immediate contact with them, dips south of east, 
northerly dips being quite exceptional, and then occurring only at a 
distance from the schist. As the latter dips steadily to the north- 
east, it is difficult to see how it can be conformable to the limestone. 

But a more comprehensive view of the district presents the re- 
ceived view in a still more incredible light. East of Sango Bay is 
the Smoo mass of limestone. Though separated from the Durness 
area by a faulted strip of gneiss, it preserves the same south-south- 
west strike. Hast of Sangomor it steadily dips east-south-east, at 
Smoo it is about horizontal, but west of Sangobeag it turns up to 
the south-east and a syncline is formed. On the promontory of 
Leirinmohr some of the dips are nearer south than east; but in no 
case have I observed in the Smoo mass the north-easterly dip of the 
altered series. : 

As Murchison himself admits that higher up the valley the 
limestone is faulted against the old gneiss on both sides, it is need- 
less to follow it further. 

b. Geiss of Sango Bay.—Between the two limestone areas is a 
band of schist. On the shore it is well exposed, not ‘“ thrust about 
in dire confusion,” but forming a symmetrical half-dome facing to 
the north-east. On the east side, within a few yards of the lime- 
stone, and dipping towards it obliquely (that is, to east-north-east, 
at 40°), is hornblendic and chloritic gneiss. ‘Towards the west the 
dip curves gradually round to north-east and north, and the gneiss 
is then underlain by dark mica-schist, which soon occupies the 
shore, the dip turning round to north-north-west, and finally to 
north-west, as if it would pass beneath the limestone which crops 
up in the shore close at hand, and is seen in the cliffs with its usual 
low south-easterly dip. There is no doubt that this strip is faulted 
(f, and f,) against the limestone on both sides. 

his gneiss is of the same kind as that which underlies the flagey 
schist on Far-out Head and east of Loch Hrriboll, and it must not 
be confounded with the Lewisian, which in this district is very 
massive, coarsely crystalline, and almost vertical inits dip. Accord- 


242 C. CALLAWAY ON THE LIMESTONE 


ing to received views, this newer gneiss must overlie the limestone ; 


and, if so, the force which contorted the former must also have 
affected the latter. But though the limestone comes up to the 
gneiss on both sides, its gentle south-east dip is not changed. It 
would appear to be more reasonable to infer that the limestone was 
deposited on the contorted gneiss, and that the latter was subse- 
quently thrust up through the former between two parallel faults. 

c. The Quartzite——The bay between the limestone promontory 
of Leirinmohr on the west and the headland of Lewisian gneiss on 
the east is occupied by the quartzite, which is also seen sloping 
down from the fianks of Ben Keannabin to the bay and forms a 
small outlier on the headland just named. It dips uniformly to the 
north-east. In the western angle of the bay it is faulted ( f,) against 
the limestone. Both limestone and quartzite are crushed into thick 
breccias at the junction, and on the bank immediately above the 
limestone is seen dipping to the north-west, the strikes of the two 
formations being, as in the former case, nearly at right angles. 

To sum up these results, it is clear that the metamorphic rocks, 
quartzite and schist, have been affected by a force tilting them up 
to the north-east, while the limestone forms a syncline whose axis 
strikes to the south-south-west. How the limestone can hold a 
conformable relation to the altered groups is a problem which the 
advocates of the received view may fairly be called upon to solve. 

Though the metamorphic rocks occur in three distinct patches, it 
is probable that they form a true succession. To the east of Loch 
Kirriboll, the quartzite is overlain by gneiss of the Sango-Bay type, 
which is surmounted by the flaggy group on Loch Hope. In Durness 
the Sango-Bay gneiss must overlie the quartzite, since the latter 
rests on the Lewisian gneiss; and if the flags of Far-out Head were 
~ prolonged on the strike to the south-east, they would overlie the 
newer gneiss. 

ASSYNT. 

The section on which Murchison placed chief reliance is the suc- 
cession on the south-west slope of Cnoc an drein, above the church. 
As I entirely differ in my reading, I submit a section of the ground 
(fig. 2), in which I have carefully excluded hypothesis and have 
simply inserted the facts observed. 


Fig. 2.—Section above Inchnadamff Church. 
S.W. ) NE. 


L. Limestone. DL. Dark Limestone. F. Felsite. 
WL. White Limestone. Q. Quartzite. 7. Fault. 


— a ee 


OF DURNESS AND ASSYNT. 943 


Just above the road we come to a band of dark limestone, which 
is succeeded by a white zone. Beyond a little hollow, quartzose 
flags are followed by similar dark and white limestones, which are 
probably a repetition; then in succession we have felsite, lime- 
stone, and felsite. Hitherto the dip of the bedded rocks has been 
to the north-east. Above the felsite is a considerable thickness of 
quartzite dipping east, so that if it overlies the limestone it must be 
unconformable. Higher up the quartz-rock dips for some distance 
to the south-west. Approaching the summit of the hill, felsite 
appears, and just beyond is massive quartzite dipping steadily to the 
south-east for a considerable distance. 

It is obviously unsafe to base a succession upon such a broken 
section as this. Passing over the intrusions, it is certain that more’ 
than one fault * occurs, and that the strike of the limestone is at 
right angles to the main mass of the quartzite. But a still more 
decisive refutation of Murchison’s views remains. 

Following the strike to the south-east it is seen gradually to 
recede from the quartzite ridge, while opposite dips by degrees come 
in on the northern side of the limestone band, which thus expands 
in the plateau of Stronchrubie into a broad syncline, the northern 
side of which dips away from the quartzite, that is, to the south- 
west. Towards the ridge the limestone dips grow steeper and aft 
last approach the vertical. Climbing the face of the mountain 
above, the quartzite is seen to dip to the north of east, so that the 
dips of the two formations are in opposite directions. As the 
northern side of the syncline gradually disappears towards Cnoc an 
drein, it is obvious that it is cut out by a fault, so that the southern 
side of the basin is brought against the quartzite and appears to dip 
below it. ‘These facts are illustrated in fig. 3. 


Fig. 3.—Plan of Limestone and Quartzite at Inchnadamff. 


wy AN 
Ca ae 


* Almost certain at F, and F,. 


244 C. CALLAWAY ON THE LIMESTONE 


There is another difficulty in the reception of the old view. 
According to Murchison, the limestone is conformably overlain by 
the great mass of quartz-rock which rises into the lofty peaks of 
Ben More. If so, then the limestone basin of Stronchrubie must 
have been covered by an equal thickness of quartzite. It is for the 
followers of Murchison to show how denudation could have cleared 
off the entire mass of such an intractable rock as quartzite from the 
limestone, and yet have made so little impression upon the mountain- 
ridges which overhang. 

The quartzite which slopes down from the south-easterm face of 
Queenaig passes up through a continuous quartzite ridge into Ben 
More. It is not pretended that the limestone occurs in this line of 
section; but it is very singular that it should have thinned out 
just where its presence would have been of decisive value. 

One important and suggestive fact remains. At both Durness and 
. Assynt the limestone forms a symmetrical basin, the axis of which 
does not agree with the normal strike of the rocks which are sup- 
posed to overlie, but which coincides with the axis of the existing 
valley in which vt les. In Assynt the axes strike to the south-east, 
at Durness to the south-south-west. 

I do not at present press these conclusions beyond the districts 
observed, but submit them as an instalment towards the solution of 
a great question. 


Discussion. 


The PrestpEnt said he had twice visited the area, and felt diffi- 
culties in tracing the succession of the rocks, especially as to the 
asserted recurrence of the quartzites. The fossils proved the lime- 
stone to be of about Arenig age; certainly the limestones seemed 
to lie in a synclinal basin on the quartzites. 

Prof. Jupp said that, after several visits to the district, he felt 
great difficulty in offering an opinion as to the succession of beds ; 
he thought the simple sections commonly drawn to illustrate the 
geology of the country did not hold good. He considered that 
Prof. Nicol had made good his position as to there being only one 
quartzite and one limestone; but as to the relation of these to the 
gneiss, there was great difficulty in arriving at a satisfactory con- 
clusion. 

Dr. Hicxs agreed that the limestone series sometimes lay in 
synclinal folds of the quartzite; but he thought there might be two 
quartzites, as in a case he had recently noticed in another region 
along the same line further south. He was thoroughly satisfied that 
the series was newer than, and did not underlie, the eastern gneiss. 

Mr. Huptrston observed that Dr. Callaway’s mapping of the 
Durness area coincided with Sir R. Murchison’s section as to the 
limestone being troughed, and also as to the reversed position of 
the two gneisses. Further south the quartzites and limestones ap- 
peared to be intercalated, as at Loch Erriboll, where Orthoceras had 


OF DURNESS AND ASSYNT. 245 


been obtained from the quartzite, tending to show that the two 
rocks hang together. The Assynt series, though less distinct than 
that at Loch Erriboll, was not inconsistent with such an idea. The 
unequal tension of the two rocks might in part account for local 
discordance of strike. 

Dr. Cattaway could not agree with Mr. Hudleston as to his 
interpretation of the Durness section; such discordant strikes were 
incompatible with the idea of a conformable succession ; nor did 
he agree with his view of the Loch-Erriboll rock. He accepted the 


age of the limestone, 


Q.J.G.8. No. 146. S 


246 R. ETHERIDGE ON A TRIGONIA FROM THE 


17. On a New Srrcizs of Triconta from the Purseck Bens of the 
Vatz of Warpour. By R. Erazripes, Ksq., F.R.S., Pres. G.S. 
With a Note on the Strata, by the Ruv. W. R. AnpREws, 
M.A. (Read December 15, 1880). 


Earty in this year the Rev. W. R. Andrews, of Teffont Rectory, 
near Salisbury, placed in my hands what at first sight appeared to 
be a new species of Z’regonia from the Purbeck rocks in the Vale of 
Wardour. This new species was found by Mr. Andrews in the rail- 
way-cutting one mile west of Dinton Station in the Vale of Wardour ; 
and its geological or stratigraphical horizon is the “ Cinderbed ” so 
well known in the several sections in the Isle of Purbeck and else- 
where. This bed is here composed of hard grey marl and pale 
brown soft limestones, variable in their proportions. Ostrea distorta 
and casts of TZrzgonca abound in this Cinderbed, which here measures 
about 2 ft. 6 in. in thickness. The section accompanying this 
paper (fig. 2, p. 252) was constructed by Mr. Andrews, and most 
correctly represents the succession of the Lower and Middle Pur- 
becks at the spot in the vale whence the new Trigonza came. It 
was from this marine “‘Cinderbed ” at Durlston Bay that Hemicdaris 
purbeckensis, Forbes, was obtained, associated with Ostrea distorta, 
a Trigonia like 7’. gibbosa, Cardium Gubbsir, Perna sp., and remains 
of fish. No other locality or section of the Dorsetshire Purbecks 
has yielded the above. ‘There cannot be any doubt that this 7ri- 
goma is not T. gibbosa. The Rev. O. Fisher obtained T'rigonia gib- 
bosa(?) from the middle Cinderbed of Fovant, Wilts; and the Rey. Mr. 
Andrews has obtained four other specimens of the same species from 
his immediate area; so that the genus is well represented in the 
Middle Purbeck of the Vale of Wardour. The Lower Purbeck beds 
of the vale lie below the datum of the railway and the level at 
which many of the quarries are worked, the “‘ Juxeat ” beds being 
about the lowest seen. The beds exposed in the railway-section near 
Teffont include the base of the Wealden, all the Middle Purbecks of 
the Vale of Wardour, and the top of the Lower Purbecks. The middle 
group here is about the same thickness as the beds in the Isle of — 
Purbeck ; those at Ridgway Hill and Mewps Bay are about 50 ft. ; 
and at Worborrow Bay the middle series measures 90 ft., and at 
Durlston Bay 150 ft. There is no Upper Purbeck in the area 
under observation ; the hard marl with eroded surface is the highest 
member of the Middle Purbeck, the succeeding 17 ft. being probably 
Hastings Sand at the base of the Wealden. 


Description. 


Section Glabree. 
TRIGONIA DENSINODA, Ether. (Fig. 1.) 


Shell greatly elongated or lengthened posteally, the anteal or 


PURBECK BEDS OF THE VALE OF WARDOUR. 247 


anterior side rounded and somewhat steep; the base (inferior or 
ventral border) flattened; the postero-dorsal (umbonal or superior) 
-border concave and much elongated. 


Fig. 1.—TZriyonia densinoda, Ether. 


Umbones pointed, small, but prominent or elevated and slightly 
recurved, antero-mesial, or placed within the anterior third of the 
valves. 

Area moderately wide, flattened and filled with reticulated costelle, 
which are minutely and delicately nodulated or tuberculated ; it is 
bounded by two carine, the outer densely or closely nodulated, in- 
creasing in width from the umbo to near the extremity of the pos- 
teal side, then merging into the system of tuberculated transverse 
costee ; or the extreme posteal border possesses many irregular lines 
of growth due to the elongation of the tubercles. ‘The inner 
carina, or that bordering the escutcheon, consists of a series of minute 
tubercles which are immediately surmounted by the tranverse wavy 
ridges of the escutcheon. The median sulcus, or mesial furrow, 
is well defined. The escutcheon is remarkably large, having trans- 
verse Wavy rugee resembling those occurring in the Neocomian Qua- 
drate. The shell (valve) possesses regularly and concentrically 
arranged tuberculated or knotted rows of coste; about 40 range 
from the umbo to the posteal attenuated border. Those tubercles 
upon the posterior third of the valve, below the area become elongated 
and less regularly arranged than those of the umbonal region, where 
they are closely concentrically arranged. 

Affinities and Differences. —In outward ornamentation 7’. densinoda 

s 2 


248 EK. ETHERIDGE ON A TRIGONIA FROM THE 


mihi, much resembles 7’. tenwitexta from the Portland Oolite of 
Portland, Devizes, Crookwood and Tisbury; but the shell is more 
depressed and lengthened posteriorly than 7’. tenwtexta, and does 
not possess the antecarinal sulcus or space of that species, which 
occurs in all the known Jurassic Glabrae, and which is so essentially 
characteristic of the Portland group. 

The ornamentation is that of the Upper-Jurassic Glabree ; but the 
escutcheon, which is remarkably large and possesses transverse 
ruge, agrees with that of the Neocomian Quadratz, to which it 
allies the species. 

All the Neocomian Quadratze have the escutcheon ornamented, 
and therefore distinct from that part of the tuberculated Jurassic 
species in which itis plain or unornamented. Again, the Cretaceous 
Scabree have also ornamented escutcheons; but the latter are alto- 
gether different and far removed from the Jurassic Trigome. 

I therefore regard this new species of Trzgonia as atransition form, 
tending to connect the Portlandian Glabre, through its ornamenta- 
tion, with the Neocomian Quadrate, through the ornamented and 
peculiar or characteristic escutcheon. Its stratigraphical position 
at the highest portion of the Jurassic system indicates a zoological 
change in progress as regards the genus Zrigonia, constituting an 
important feature in the history of the genus, no other example of 
which, so far as I am aware, is known in rocks representing the 
Purbeck formation, either British or continental. 

This new species falls under the group or section Glabre, founded 
by Agassiz upon very insufficient “data, and described by him as 
beivg without ornamentation, having no tubercles or costa. The 
shells of this group are usually ‘“inflated-ovate or ovately-oblong ; 
and the area is only slightly separated” from the rest of the valve. 

Mesially, or anterior to the position of the marginal carina, there is 
a smooth space which commences at the apex (umbo) or near it, and 
gradually widens downwards to the posteal border. This smooth 
space is more depressed than any other portion of the valves, and is 
distinctly impressed by lines of growth. 

In this group the “ anteal ” portion of the valves possesses costee 
more or less prominent, usually small, closely arranged, and either 
plain or tuberculated. The number of species belonging to this sec- 
tion is seven, six of which are Jurassic, the remaining form being 
Cretaceous (2. excentrica, Park.). 

Our species is closely allied to those forms known in the Portland 
rocks under the name 7’. gibbosa, Sow., 7’. Damonana, de Lor., 7’. 
Manselli, Lycett, and 7’. tenwitexta, Lycett, and. especially to the 
last named, from which, however, it differs in form and characters. 


Nore on the Strata. By the Rev. W. R. Anprews, M.A. 


Mr. Andrews has drawn up the following description of the 
Purbeck beds of the Vale of Wardour ; and any of the sections would 


suffice to read aright the succession and position of the fossil under 
consideration. 


PURBECK BEDS OF THE VALE OF WARDOUR. QAO 


Mr. Andrews says, “the Purbeck strata occupy an area of from 
five to six miles square at Teffont Ewias, Chicksgrove, Lady Down, 
&c. in the centre of the Vale of Wardour, and rest conformably 
upon the Portland group, sometimes, but not always, separated by a 
thin band of dark clay. These estuarine deposits have been consi- 
dered to belong to the Jurassic system, in consequence of their pre- 
senting remains of animal life nearer in affinity with the Jurassic 
period than with the Wealden above, and also from the fact that the 
marine Portland beds pass into the freshwater Purbecks without un- 
conformity, and generally with an ancient land surface at or near the 
junction ; the same area too (for the Portland beds are always capped 
by the Purbecks) which formed the bottom of the Portland sea, became 
when raised, first the support of the ancient forest, and then, when 
again slightly depressed, the delta of some great river, and this 
without any unconformity or denudation, which would imply a great 
lapse of time. 

“<The Purbeck beds in the Vale of Wardour are comparatively much 
thinner than those exposed on the Dorsetshire coast, and here pre- 
sent only about 60 or 70 feet, belonging entirely to the lower and 
middle divisions. 

“‘ Whether the Upper Purbecks were ever deposited here, it is im- 
possible to say. These Purbeck beds thin out rapidly on the coast, 
passing from K. to W., or from Durlston Bay to Portland Bill, and 
also from 8. to N., passing from the same exposure at Swanage 
through the Vale of Wardour to the thin capping at Swindon. 
This thinning-out may have been due to the north and west sides 
of the estuary being raised and above water when the Upper Purbecks 
were being deposited in the south and east—a supposition which 
is borne out by the Purbeck at Swindon, shown by Mr. Blake to 
be ‘in point of time as old as some parts of the Portland.’ Or, 
on the other hand, the upper beds may have been deposited and 
denuded—a supposition not improbable when we remember that 
the Cretaceous system rests quite unconformably on these freshwater 
beds, e.g. the Gault in the Vale of Wardour overlapping the beds 
below. 

*« New quarries, opened since Fitton wrote his memorable paper, 
have supplied some very interesting fossils, amongst which is the 
new Trigonia (T. densinoda). 

“ At the junction of the Purbeck beds with the Portland strata 
there generally occurs a thin bed of dark clay; and some few feet 
above may be seen an ancient earth or land surface, which is, in 
places, as much as 2 feet thick. This bed has in it large pieces of 
coniferous wood and a Cycad, as in the Isles of Portland and Pur- 
beck. The “cap” as it called, is here visible, and has yielded 
some interesting Mollusca, besides several species of fishes, and 
a very much larger form of Archeoniscus than A. Brodie. The 
beds immediately above the Middle Purbecks are not well exposed, 
they consist (as on the coast) of soft shales with Cyprids and marls, 
affording no good stone worth the quarryman’s labour. These, 
I presume, would be equivalent to the soft cockle-beds of the 
Durlston section. 


250 REV. W. R. ANDREWS ON THE 


“‘ At the top of the Lower Purbecks and in the Middle Purbecks 
much useful stone occurs, and has been extensively quarried both 
for building-purposes and for burning into superior lime; from these 
exposures some interesting fossils have been obtained. 

‘Thick beds of a hard grey marl at the top of the Purbecks, very 
similar in appearance to the insect-beds of Durlston, but containing 
Cyprides, here afforded some few insect-remains, several species (6) 
of fishes, and Archwoniscus (the fishes are the following— 


Microdon radiatus, Ophiopsis breviceps) ; 
Plewropholis, 


also Turtle and Crocodile remains. The insect-remains are not so 
plentiful as at Durlston, a comparative scarcity arising from the 
terrestrial condition of these beds in the Vale of Wardour, as evi- 
denced by the presence of Cyprides. 

‘“‘ Passing up through the ‘‘ cherty freshwater ” beds, which here 
contain, as in Dorsetshire, Paludine and Cyclades beautifully imbed- 
ded in flint, we arrive at one of the most interesting beds of the whole 
series, a marine bed called, in the island, the “Cinder.” Although 
it is much reduced in thickness from 12 ft., crowded with Ostrea 
distorta, still, in the Vale of Wardour, it maintains its marine cha- 
racter, and, as elsewhere, from its hard enduring nature, has outlasted 
many softer beds. Beside the Ostrea distorta which is scattered 
through it, two species of Zragonia occur, 7’. gibbosa and a new 
species which Mr. Etheridge has named 7’. denstnoda, which has not 
occurred in any other formation and, according to Mr. Etheridge, is 
of much paleontological interest, arising from the fact that it has 
characters connecting it with the Jurassic Zrigonia-group Glabre, 
on the one hand, and Cretaceous forms of the group Quadrate, on 
the other. Occurring, as it does, in beds of a transitional character 
between the Jurassic and Cretaceous, it is more interesting still. 

‘‘ Higher up in the Middle Purbecks occurs an extraordinary abun- 
dance of the fossil Isopod Archconiscus Brodiei, sometimes so closely 
lying together that 250 specimens haye been obtained on a slab not 
larger than one foot square. The 8 or 10 feet of red and yellow 
stratified sandy clays which are here found on the top of the Pur- 
becks, are possibly the Wealden. They rest upon an eroded surface 
of limestone, but otherwise present no unconformity. It is remarkable 
that these Wealden beds so often cap the Purbeck series, a fact 
that seems to indicate that the same area which served for the delta of 
the Purbecks performed the same office for the Hastings series, with, 
however, this difference—that the source from which the materials 
came must have been different, although the ancient river may 
have served for both formations. 

“« Endogenites erosa, not hitherto clearly proved to belong to any 
other horizon than the Wealden, has been found in these red and 
yellow sandy clays in situ ; a stratified bed above contains a small 
Modiola and Cyrena; but whether Wealden or Purbeck, I am not 
able to say. Hndogenites erosa occurs in a similar section half a 
mile to the east.” 


PURBECK BEDS OF THE VALE OF WARDOUR. 251 


The Rev. Mr. Andrews has made a careful section of the railway- 
cutting at Teffont, which shows the Hastings beds and the Middle 
and Lower Purbecks. The Rey. O. Fisher marked the divisions on 
the section. Mr. Andrews is of opinion that no higher Purbeck 
beds occur in his neighbourhood than those shown on the section, 
and that the Hastings series were deposited upon the eroded Purbeck 
strata; the Purbeck strata continue 50 ft. or more below the base 
of the railway-cutting, as there are many quarries in the neigh- 
bourhood with many feet of Purbecks resting upon the Portland 
beds. The section in the railway-cutting ends 7 ft. below the 
“cinder bed;” but the quarry near the Rectory has been carried 
19 ft. below the ‘‘ cinder,” into hard grey marls, which are burnt 
for lime, and contain many fish, insects, Archwonscus, Cyclades, 
Cyprides, &c. 


The following 1s a detailed section of the Portland beds in the 
railway-cutting, drawn up by Mr. Andrews, which, although similar 
to the woodcut-section on p. 252, nevertheless enters more into the 
particulars of the beds, both as to thickness and succession, and will 
doubtless be found useful if the district is visited and examined : — 


No. ft. in. Surface. 

Rif 1. 2 Q Brown earth, with scattered flints, passing into 
S| 2. 2 Q  Redand yellow sand, with a few scattered flints. 
3 | 3. 1 6 Grey sand with red streaks, passing into 
=e | 4. 1 O Grey sandy clay, red and yellow layers. 

a { 5. QO 2  Purplish red clay, laminated. 
5 6. 0 6 Green clay, light-red lines. 
= | Te 4 Layer of iron in lamine. 

\ 8. 1 Green clay, red irony lines of variable thickness, resting 
unconformably on the bed below, and running down 
amongst the lumps. 

( 9. 1 90 Worn flint-shaped lumps of hard marl, vertical fracture. 

10. 1 6 Sandy, with perished shells in layers. 

| 11. O 6 Clay, red and grey, in lamine. 

12. 0 2 Very red sandy rock, sometimes blue and grey, with large 
bivalves, Ostree. 

| 13. 1 0 Limestone layers in sandy clay. 

14. 0 9 Thin laminated brown sandy limestone. 
@ 15. 1 0 Hard sandy stone, often red outside, ripple-marks on 
® the top. 
= 16. 1 0 Composed of perished shells, thin layers of limestone 
as and a line of ‘“‘ beef” about the middle. 

. | 17. 1 4 Redand yellow sand in layers. Ostree. 
eS Ss. Clay, thin-laminated, soft and yellow; impressions of 
2 Archeoniscus. 

2 19. 0 9 MHard blue and brown marl, Archeoniscus, Cyprides, 
Ay 4 Cyclas, Ostree, insects’ wings. 
x 20. 0 3 Laminated thin white limestone and sand. 
oS 21. 0 3 Hard red sand rock, occasional streaks of blue; oysters 
= and impressions of large bivalves. 
22. 0 2 Soft sandy limestone. 
Carried a 
forward }16 13 Le de 


Fig. 2.—Section of Purbeck-Beds in a Railway-cutting 11 mile west 
of Dinton ae Vale leh Wardour, (Scale, 1 inch ‘to 1 foot.) 


MIDDLE 
PURBECKS. 


LOWER 


PURBECKS. 


— 


i 


Me pal toon 


VILLE 
re 


Surface. 


Flints, “ trail.” 


Yellow and red sandy clays. Endogenites erosa. 


Hard marl, eroded surface. 


Sandy, perished shell in layers. 


Laminated clay. 
Red sandy rock. Ostrea. 


Limestone, laminated. 


Red and yellow sand. Ostrea. 


Archeoniscus, Cyprides, Cyclas, Ostrea. 


Hard blue marl. Archeoniscus, Insects. 


Hard red sandy rock. Ostrea. 


White fissile limestone. -Archeoniscus. 


Brown limestone rock. Ostree and Fish. 
« Beef.” 
Chert. 


Cinder-bed.  Ostrea distorta and Trigone 
(Lf. gibbosa and T. densinoda). 


Hard limestone. 


Sandy rock. 
Old earth-surface. 


“TLias.” Cyprides, Archeoniscus, Fish. 


Clay. 
Hard crystalline rock. 


ON THE PURBECK BEDS OF THE VALE OF WARDOUR. 953 


a 


Lower Purbecks. 


ING, Vy Teng 

Brought ‘ 

orward i 16 1a 
= 23. 1 4 Soft white limestone, laminated, ‘‘ White bed.” <Archeo- 
S NUSCUS. 
S 24. 1 0 Brown rock, shelly, large bivalves, oysters, fish-remains. 
iS | 25. Oolitic stone, brown. 
Ss | 20. Soft limestone, vertical fracture. 
2 s Pipe Oa os Beet. 
24 28. 0 6 Sand and limestone in layers. 
2 | & 29. 0 2 Chert. 
& | = 30. 0 2 Sandy clay, dark brown, shelly. 
© | w ol. 2 5 Hard grey marl, and brown soft rock, very varying in 
nS, 3 their proportions, sometimes the hard marl taking up 
= in all the space, and vice versd. ‘‘Cinder” containing 

5 scattered Ostrea distorta, and T'rigonie. 

6 32. 3 0 Hard crystalline limestone, in 1, 2, or 3 layers, with thin 
sand or clay between, blue outside, and containing 
lumps of chert, Cyclades in clay parting with vegetable 

| remains and Paludine. Cyclades and Ostree in the 
| chert. 
\ 03. 1 0 Sandy rock, yellow. 
( 
| 34. 0 2 Dark clay. 
4 39. 1 6 Hard grey marl. Insect-beds of Purbeck. 

36. 0 2 Clay. 
| of. Hard crystalline rock, shelly. 
c 

23 65 
Discussion. 


Mr. Hutxs remarked that the fauna of the “ Cinder-bed” being 
very limited, even where it was best known, namely at Swanage, 
any addition to it was of great importance. The discovery of the 
species described by Mr. Etheridge was especially important, the 
fossil being so strongly characterized that no doubt could be enter- 
tained as to its distinctness, while, at the same time, it was par- 
ticularly interesting as binding together two other forms, an older 
and a younger one, and thus, to a certain extent, bridging over the 
gap between the Jurassic and Cretaceous Trigonie. 

Dr. Duncan thought that the Wealden-Purbeck forms a series 
intermediate between the Cretaceous and the Jurassic. 

Prof. Srrtny agreed with the author that the form is a new 
species. 

The Aurnor stated that he had shown the specimen to Dr. Lycett, . 
who agreed with him as to its peculiarities of character. 


254 W. J. SOLLAS ON A NEW LYSSAKINE HEXACTINELLID 


18. On Astroconta GRANTI, a new LyssakINE HExACTINELLID from 
the SILURIAN Formation of Canapa. By W. J. Sottas, Esq., M.A., 
F.R.S.E., F.G.8., Professor of Geology in University College, 
Bristol. (Read February 23, 1881.) 


By the great kindness of Lieut.-Col. Charles Coote Grant, I have re- 
ceived a valuable collection of fossil specimens from the Silurian strata 
of Hamilton, Ontario. J hope, as opportunity serves, to offer de- 
scriptions of the more interesting of these fossils, and in the following 
short paper make a beginning by describing an incompletely silici- 
fied specimen of chert, which by its remarkable similarity to the chalk- 
flints of Trimmingham arrested my attention when studying those 
bodies. The specimen in question is a small piece (1 inch cube) of 
greyish siliceous dolomite with a gritty granular texture; on one 
face it bears a carbonized Hydrozoon, on another a silicified Polyzoon, 
and on a third shows the opening of a long winding cavity which 
extends into the interior. From the sides of this cavity some long 
cylindrical rods, very suggestive of sponge-spicules, were seen 
conspicuously projecting: ‘under the microscope by reflected light 
they were seen to be covered with an irregular crystalline deposit, 
which made the resemblance to spicules less striking than when exa- 
mined by the unassisted eye. The specimen was now broken into two 
pieces, and the part containing the rods and cavity placed in dilute 
hydrochloric acid. Solution with faint effervescence took place; and 
after standing for twenty-four hours the supernatant liquor was 
poured off for subsquent examination, while the sediment and 
small piece of siliceous dolomite remaining were well washed and 
further examined. From the sides of the cavity in the dolomitic 
chert, cylindrical rods were still seen projecting, and under the 
microscope were found to have lost their irregular outline and 
at the same time to have somewhat diminished in thickness ; 
evidently a crystalline coating had been dissolved away from them ; 
and in the slender siliceous rods which remained one recognized at 
once true sponge-spicules. 

The sediment was next examined. It contained a number a large 
siliceous spicules, some of which are figured on page 255, also some 
small colourless transparent hexagonal prisms capped with six-sided 
pyramids at each end, evidently quartz crystals, and, finally, some 
minute rhombohedra, which were subsequently found to be mag- 
nesite. 

The Sponge-Spicules.—By reflected light the spicules appear snow- 
white, with a vitreous lustre; by transmitted light they are glassy and 
translucent ; when immersed in water, glycerine-jelly, or Canada- 
balsam, they become quite transparent, except in places where they 
are traversed by a minute black spongy network which appears 
white and shining with reflected light, and evidently contains air: 
when this is displaced by the liquid medium, the spicule becomes 


FROM THE SILURIAN FORMATION OF CANADA, 255 


Figs. 1-11.—Spicules of Astroconia Granti, Sollas. (All magnified 
40 diameters, except figs. 38, 5, and 7, which are magnified 104 
diameters. ) 


256 W. J. SOLLAS ON A NEW LYSSAKINE HEXACTINELLID 


transparent throughout; it is frequently colourless, but sometimes 
presents a faint yellowish brown tint either in places or throughout. 
The surface of the spicule has a rough irregular appearance when 
examined dry, which disappears, however, on immersion in balsam, 
being evidently due to a crystalline incrustation (magnesite). When 
this is removed the sides of the spicules appear as neat continuous 
lines; the axial canal is also quite sharply defined, and is sometimes 
not more enlarged than in recent deciduous spicules, though 
occasionally it is so wide as to reduce the spicule toa mere shell. In 
some instances it has been infilled with silica, so as to appear as a 
cylindrical rod projecting from what remains of the spicule in which 
it was formed (fig. 8, p. 255). 

The spicules are not excavated by rhombohedral pits like those 
seen in Hyalostelia Smithi, Young, from the Carboniferous Limestone 
of Scotland ; but sometimes they are pitted all over with hemispherical 
cavities such as are seen in deciduous and fossil spicules, and which 
can be produced artificially in recent spicules by solvent agents. 

With polarized light all the spicules give brilliant colours, and 
much more vividly than those from the chalk of Trimmingham ; the 
colours frequently occur in spherical patches, as though crystallization 
had been set up about various centres within the spicular substance ; 
one might term it an internal botryoidal structure, though the 
spicules are never botryoidal externally ; somtimes the polarization- 
effects show clearly that the spicule is composed of minute prisms 
radiating outwards from the sides of the axial canal. 

In concluding this brief account of the mineral state of the spicules, 
I may remark that, though highly crystalline, they are almost if not 
quite as well preserved as those of Hyalostelia from the Carboniferous 
formation ; and this is certainly surprising when we consider their 
ereater age and much smaller size. The conditions under which the two 
sets of spicules were preserved were evidently very different, however ; 
for Hyalostelia does not appear to occur in association with deposits of 
chert as is the case with Astroconia. Hyalostelia is possibly preserved 
to us chiefly by reason of the magnitude of its spicules, Astrocoma 
chiefly by secondary silicification. 

We now pass on to a description of the forms of the several spicules, 
which are figured on page 255, all being magnified 40 diameters 
except figs. 3, 5, and 7, which are magnified 104 times. 

Fig. 1 is the commonest form, occurring in fragments of various 
lengths, that represented being the longest seen. It is almost exactly 
cylindrical; and the axial canal maintains a straight course and even 
width throughout. Its natural termination has not been seen; but 
probably it was pointed at both ends, and formed the acerate 
spicule of the sponge. 

Fig. 2 looks at first sight like the trifid spicule of a Tetractinellid ; 
but careful examination shows the presence of four rays, of which 
that marked a is one; though as here represented it appears 
merely as a bifurcation of the one in front of it. By examining the 
other side of the glass slide on which.the spicule is mounted its real 

nature is readily made out; moreover I was able to turn an almost 


FROM THE SILURIAN FORMATION OF CANADA, 957 


exactly similar spicule over on its head, so that its shaft projected 
vertically ; the four capitular rays could then all be seen in one view. 
The axial canal of the shaft is prolonged upwards past the common 
origin in the head, and, no doubt, once extended into a short vertical 
spicule ; indeed the companion spicule showsas much. In addition 
to this axial spine, containing a continuation of the canal, the head 
is ornamented by conical tubercles given off in the same direction 
from the upper surface of the capitular rays (as shown in the figure). 

I take this spicule to have projected radially from the surface of 
the sponge-body, the shaft being directed inwards and the capitular 
rays expanding parallel to the surface of the body, as in the similar 
spicules of Holtenia and Rossella; it has not the appearance of an 
anchoring spicule. 

Figs.3 and 3a. These two drawings represent the same specimen, 
the upper one taken before immersion in a liquid medium and by re- 
flected light, the lower one by transmitted light when mounted in 
Canada balsam. They are magnified 104 diameters: so that, if 
represented on the same scale as the other figures, they would not be 
quite so large as the spicule represented in fig. 6. he noticeable 
feature is the presence of spines on the rays, directed in all cases 
towards the ends. ‘The rays were evidently at least five in number ; 
probably there were six. | 

Fig. 4. The four rays of this spicule are directed downwards 

below the plane of the paper ; near their origin they each give rise 
to a large bifid spine inclined upwards away from the plane of the 
paper. It is probable that this specimen once formed the head of a 
spicule like fig. 2. 
' Fig. 5. This spicule (X 104) is chiefly remarkable for its spinose 
rays: on the longest but still broken ray three spines are given off 
from one side, and a fourth, represented in plan by a circle, from 
the adjacent side at right angles. 

Fig. 6. The proximal half of the rays is thicker than the distal 
end, which is long, slender, and directed downwards, the proximal 
half lying in the plane of the paper. The thicker part of the rays is 
spined—the spines (so far as one can judge from the three which 
remain on one of the rays) being arranged spirally, and two of 
them arising near the origin of the slender extremity. 

Fig. 7. This represents a fragment of a cylindrical spicule magni- 
fied 104 diameters, with the axial canal filled up with silica, which 
now projects at both ends like an acerate spicule run through the 
middle. 

. Fig. 8 is also a fragment of a cylindrical spicule; it shows the 
hemispherical pits which have been eaten out of its surface. 

Fig. 9. This is one of the most frequently occurring forms ; it is a 
large simple sexradiate—the rays having a cylindrical form, only 
slightly tapering towards their broken extremities. The axial 
canals are regular and clearly defined. ‘This is probably one of the 
staple body-spicules of the sponge. 

Fig. 10 is a smaller spicule of the same kind. 

Fig. 11. This is less like a Hexactinellid spicule than the others, 


258 W. J. SOLLAS ON A NEW LYSSAKINE HEXACTINELLID 


possibly owing to its being incomplete and yet simulating complete- 
ness through a sharpening of its ends by solution. The axial canal 
can be clearly traced through one half of it only, the other half being 
rendered opaque by the close firm network of cavities which exca- 
vate it. 

Leaving out of consideration the last, which is a doubtful form, 
and a few other fragments not figured and quite undeterminable, one 
may safely say that all the spicules extracted from the small 
fragment of dolomitic chert are distinctly Hexactinellid; and it 
would appear that we probably have here, not a heterogeneous mix- 
ture of spicules derived from several kinds of sponge, as is usually the 
case in chalk-flints, but the much rarer case of a natural assem- 
blage of forms derived from a single individual. For this group of 
forms we propose a distinct name, Astrocona Granti, the specific 
designation being given in honour of the indefatigable observer to 
whom. we are indebted for our knowledge of the sponge. 

The Quartz Crystals—These do not differ in character from those 
described from the Carboniferous Limestone by Mr. Wardle and in 
my papers on Catagma (Ann. & Mag. Nat. Hist. ser. 5, vol. u. p. 361, 
fig. 3), and on Flints (vol. vi. p. 445). I have also called attention to 
the important part which such such crystals play in rock-formation 
by pointing out that certain beds of sandstone in the Eden valley 
appear to partly consist of them. If we knew the bulk of these beds, 
we might calculate the amount of limestone which must have been 
denuded to produce them, on the assumption that this limestone con- 
tained 1 oz. of crystals in every 10 lb. of rock, as is the case with 
that at Buxton at the present day. I believe that in these quartz 
crystals, derived undoubtedly from siliceous organisms, we have also 
the key to the origin of the “krystallinische Quarzpsammite” of 
Naumann, which occur in several formations on the continent (Zir- 
kel, Lehrbuch der Petrologie, vol. 11. p. 575). They are nothing 
but the insoluble residues of limestone masses which have been 
dissolved away by the action of subaerial waters*. If this be so, 
we have in these crystalline sandstones the last stage of one of those 
beautiful cycles or, rather, spirals which appear to characterize the 
operations of nature : the silica derived from the disintegration of igne- 
ous rocks is carried in solution into the ocean and there built up by 
living organisms into forms of endless diversity and exquisite beauty ; 
this organic silica again yields to solvent influences and afterwards 
crystallizes out amidst calcareous sediments in perfectly formed — 
crystals of mineral quartz; the calcareous rocks elevated above the 
sea-level are exposed to water and the weather; solution proceeds 
apace; and after the carbonate of lime has become dissolved away, the 


* This statement is too absolute. The beautiful observations of Mr. J. A. 
Phillips, F.G.S., in complete accordance with those of Prof. Bonney and 
Dr. Sorby, show that crystalline quartz-sandstones are sometimes formed in a 
quite different manner—indeed, that the formation of quartz crystals has been 
subsequent to that of the sandstone as a whole. In deciding upon the origin of 
such sandstones, we shall have probably to be guided by the circumstances in 
each ease, before we can say certainly that the constituent quartz crystals are 
original or superinduced formations. 


pe 


FROM THE SILURIAN FORMATION OF CANADA. 259 


minute quartz crystals, uninjured and unworn, are washed into the 
sea to constitute afresh the mineral sediments of the earth’s crust. 

Rthombohedra of Magnesite.—These were left on the solution of 
the limestones in dilute hydrochloric acid, the majority of them 
occurring in a free and separate state, but a few imbedded in small 
fragments of chert, just like those figured by Professor Renard in his 
memoir on the ‘ Phthanites du Calcaire Carbonifere de Belgique’ 
(Bull. de PAcad. Roy. de Belgique, 2™° sér. t. xlvi. p. 471, 1878). To 
determine their nature, as well as that of the rest of the limestone, 
J made a preliminary qualitative analysis, first of the solution pro- 
duced by the action of hydrochloric acid, and next of the undis- 
solved residue. The former contained lime and magnesia, carbo- 
nate of lime and phosphate of magnesia being precipitated in appa- 
rently equal bulk ; the residue was boiled with strong hydrochloric 
acid, and the resulting solution analyzed; it contained a little iron, 
not a trace of lime, but a considerable quantity of magnesia; hence 
one might conclude that that part of the rock which is not chert, is 
dolomite, while its included rhombohedra, which were set free on 
solution, are magnesite; and a quantitative analysis is scarcely 
necessary. _ 

The Geological Horizon.—The rock specimen was obtained from 
the chert beds of the Niagara Limestone, which is homotaxial with 
our Wenlock Limestoue. It affords the oldest known instance of the 
associated occurrence of siliceous sponge-remains and chert. On the 
one side of a fragment of rock we find free spicules still siliceous, 
together with quartz-crystals imbedded in dolomite—on the other a 
mass of chert, in which spicules are rarely seen, and then usually as 
hollow casts, but which contains remains once calcareous (such as 
the Polyzoon before mentioned) now converted into silex. 


Discussion. 


The PresrpEnt said this was the oldest sponge next to Proto- 
spongia. The author’s remarks on it were most valuable, and those 
on the collateral subject very interesting. It was to be hoped that 
Mr. Maw would discover in the English Wenlock Limestone some 
similar microscopic forms. 

Prof. Duncan said it was interesting to see the modern Hexacti- 
nellids thus foreshadowed. Very lately he had seen one of the 
spicular forms described by Prof. Sollas in a form he had just 
described. He quite agreed with the author in assigning this form 
to the Lyssakine Hexactinellids. There could be no question as to 
the solution of the spicules in sea-water, as he had lately seen 
evidence of it in specimens from deep-sea dredgings. The results of 
Mr. Maw’s washing promised to be very interesting. He had 
examined many, but had not yet found either sponge-spicules or 
Foraminifera. 

Prof. Rupert Jones remarked that there are different kinds of 
“‘chert,” and expressed his opinion that Mr. Sollas had well 


260 ON A NEW LYSSAKINE HEXACTINELLID FROM CANADA. 


explained the origin and formation of the spicular strata which 
he had described on this and other occasions. He thought that 
Dr. Wallich’s hypothesis of the conversion of extensive layers of 
sponge-protoplasm into black flint elucidated many, but not all, of 
the phenomena connected with the origin of such siliceous strata as 
flint and chert. He stated that sponge-spicules, and numerous 
other Microzoa from the Upper Silurian shales of Shropshire, had 
been noticed lately by Mr. Smith of Kilwinning. 

Dr. Hicks said that it was remarkable that chert was not asso- 
ciated with Protospongia, as, on either Dr. Wallich’s or Prof. Sollas’s 
view, might have been expected. 

Prof. Jupp said that, as the solution of siliceous organisms had 
been recently doubted, Prof. Sollas’s observations were of additional 
interest. He himself fully believed that this solution did take 
place; now and then he had found, in examining the residues left 
by dissolving chalk in acid, the thickest portions of siliceous spicules 
still remaining not quite destroyed in chalk. 

Prof. Sottas replied that he believed a spicule had been described 
by Mr. Carter similar to that mentioned by Prof. Duncan. The one 
described now by him, however, was much more robust. He had 
never been able to find spicules in the Wenlock. He could not 
comprehend what Dr. Wallich’s views really were. That none of 
the fossil siliceous spicules which the author had described were 
originally calcareous was quite certain. As for Protospongia, it 
did not occur in hmestone, and bore a very small proportion to the 
mass of the bed; and this might account for the absence of the 
chert. 


Quart. Journ.Geol.Soc Vol XXXVILPI IX. 


WMintern Bros amp 


IGS WELAINTOS 


+ 
J 


AVLUROSAU 


PROF. OWEN ON THE ORDER THERIODONTIA. 261 


19. On the Order TumrtopontiA, with a Description of a new Genus 
and Species (ALUROSAURUS* FELINUS, Ow.). By Prof. Owen, 
C.B., F.R.S., F.G.S. (Read March 9, 1881). 


[Prats IX.] 


Or Permian and Triassic Reptilia the most interesting, those that 
help to fill the hiatus separating the mammalian Marsupials from 
the cold-blooded Vertebrates, seem to me to be the extinct species 
constituting, or referable to, the Order THERIoponmTrIA. 

To the characters of this Order given in my ‘ Catalogue of the 
Fossil Reptilia of South Africa’, viz. “‘ Dentition of the Carnivorous 
type, incisors defined by position and divided from molars by a large 
laniariform canine on each side of both upper and lower jaws,” may 
now be added “dentition ‘monophyodont’” +. At least I have not 
had evidence of an immature specimen showing a milk-series of 
teeth to be succeeded by a permanent series; but if such should be 
found in any of the extinct Reptiles of the present Order, such Order 
will be “ Diphyodont,” like the Mammalian Carnivora; for there is 
no evidence of any third set of teeth to follow those which may have 
been preceded (though I doubt it) by a first or deciduous set. Of 
the adult dentition, whether it be “ first ” or “ second,” the molars, 
as a rule, are inferior in size to the incisors, as both are markedly 
less than the canines. Add to these characters, “‘ humerus perifo- 
rated by an entepicondylar foramen”$. 

The Reptiles so distinguished or characterized are already refer- 
able te several genera ; and although I fully recognize the artificial 
character of a more or less forward extension of the ossified ‘‘ septum 
narium,” there was a convenience in disparting the Theriodont 
genera known in 1876 into “ Mononarialia” and “ Binarialia”||. 

At that date the ‘‘ Mononarials” included Cynodraco 4], Cyno- 
champsa**, Cynosuchus ++, Galesaurus tt, Nythosaurus §§, Scalopo- 
saurus || ||, Procolophon 44 ; the “ Binarials” included Lycosaurus *** 
and Tigrisuchus;ty.  Gorgonopsttt manifested a third narial 
modification. 

To this series have since been added species referable to some of 
the foregoing genera, and, also to a genus T%tanosuchus §$$. The 
latter was founded on fragmentary fossils not yielding a narial cha- 

* Gr. aidoupos, cat; catpos, lizard. t Ato, 1876, p. 15. 

t Anatomy of Vertebrates, 8vo, 1866, vol. ii. p. 268. 


§ Ib. p. 19, pl. xix. figs. 2&3 4,h; Quart. Journ. Geol Soc. August 1876, 
p: 361, cut, fig. 2, h. 
|| Catalogue, u¢ supra, pp. 15, 17. €| Ibid. p. 19. 
** Quart. Journ. Geol. Soc. vol. xvi. 1860, p. 61, pl. iii. 
tt Catalogue, ut supra, p. 21. 
tt Quart. Journ. Geol. Soc. vol. xvi. 1860, p. 58, pl. ii. 
§§ Catalogue, ut supra, p. 24. || || Ibid. p. 24. {4 Ibid. p. 25. 
F~ Ubid. p: 15. ttt Ibid. p. 17. {tt Ibid. p. 27. 
§§§ Quart. Journ. Geol. Soc. vol. xxxy. 1879, p. 189, pl. xi. 
Q.93.G.8. No. 146. a 


262 PROF. OWEN ON THE ORDER THERIODONTIA, 


racter; and with Tvtanosuchus, for the same reason, may be cited 
Kutorga’s Brithopus and Orthopus, from Russian Permian, V. Meyer’s 
Urosawrus, Fischer’s Rhopalodon, Hichwald’s Deuterosaurus, and 
Twelvetrees’s Cliorhizodon, from the same zone and locality ; to the 
Theriodontia belongs also Bathygnathus from the Trias of ‘“ Prince 
Edward’s Island,’ North America*. 

To add to this series of Theriodont genera, and, seemingly, to the 
“‘ Mononarial” section, I now submit to the Society evidence of 
another genus, dlurosaurus, exemplifying in a clear manner the 
typical Theriodont characters, under modifications generically distinct 
from those of the specimens above cited. 

It is a skull, including both upper and lower jaws, with the orbits, 
obtained by Mr. Thos. Bain from the Trias of Gough, in the Karoo 
district of South Africa, and in the usual petrified condition of the 
fossils of that formation and locality. The postorbital part of the 
skull is broken away, and the border of the nostril has been slightly 
fractured; but the rest of the specimen, with the dentition, is 
instructively preserved. 

The nostril (Pl. IX. figs. 1 & 2, 1) is terminal and vertical, and 
shows no part of a septum; in shape it is a full transverse ellipse ; 
what remains of the outlet yields in breadth 13 millim., in depth 
9 millim. Lach orbit (ab. ib. 0) gives a full obliquely vertical 
ellipse, 25 millim. by 20 millim. The facial part of the skull 
extends two and a half times the fore-and-aft diameter of the orbit 
in advance of that cavity. The hreadth of the upper jaw a little 
behind the nostril is 26 millim., and gradually increases to 35 
millim. near the orbits. The upper surface of the antorbital part 
of the skull is moderately convex ; the sides are less convex, but not 
flat; the vertical extent of the upper jaw at the middle of the molar 
series is 34 millim., and decreases to 20 millim. above the incisors. 

The skull has been subject to slight distortion ; but as this has 
been effected without fracture, it may be concluded to have been 
due to forces operating on the matrix after petrifaction, and when 
the fossil was so encased as to be equally supported on every side 
during the movements of the bed, such partial pressure haying 
chiefly affected the left orbit (Gb. fig. 3,0) and a small part of the 
same side of the skull in advance of it. From the degree in which 
the sutures are obliterated, I conclude it to have come from a full- 
grown and probably old individual, the state of the dentition sup- 
porting that inference. 

The premaxillo-maxillary alveolar border, as it recedes from 
below the nostril, follows a slightly sinuous course, concave above 
the incisors, convex above the canine and the molars; thence 
straight to beneath the orbit. 

The mandible is preserved, with the mouth close-shut; and the 
mandibular teeth are hidden by the overlapping ones of the upper 
jaw, requiring the sections made in two places for exposure. (ib. 
fig. 3,c'). The symphysis mandibule (ab. fig. 3,s) is 27 millim. in 
depth and 20 millim. in breadth where it is crossed by the upper 

* Quart. Journ. Geol. Soc. vol. xxxil. p. 352, 1876. 


AND ON ZLUROSAURUS FELINUS. 263 


canines. From its fore border, formed by the incisive alveoli, it slopes 
downward and backward, curving more directly backward where it 
terminates below. All trace of a median symphysial suture is obli- 
terated. The breadth of the mandible where the rami diverge from 
the symphysis is 20 millim. The preserved length of the mandible 
is 3 inches 3 lines; and this was probably that, or nearly that, of 
the skull. 

The suture between the maxillary (ib. fig. 1, 21) and malar (26) is 
distinct, and shows the slender pointed hind end of the maxillary 
terminating below the middle of the orbit, the lower or malar 
boundary of which is here 8 millim. in depth. 

Before the vertical parallel of the hind border of the orbit is 
reached, the malar abruptly descends (ib. fig. 1, 26) at almost a right 
angle to near the lower border of a deflected part of the mandible. 
This descending part of the malar is broken off on the left side 
Gb. fig. 3); and the mandibular depression which received it is there 
exposed. 

The mandibular ramus continued backward from the symphysis 
is subcompressed, 12 millim. in depth beneath the middle of the 
molar series, thence rapidly gaining depth, especially by a descent 
and slight inflection of the lower border; the corresponding rise of 
the coronoid plate is concealed by the malar. At the part where 
the vertical extension of the mandibular ramus begins there is a low 
tuberous outswelling of the external surface, behind which that 
surface gradually sinks and describes a moderate concavity to the 
angle of the jaw. | 

A suture indicative of angular and surangular is not visible, 
but that dividing such elements of the mandible from the dentary 
one is manifest, as is also, along the lower narrow border 
of the ramus (ib. fig. 3) the suture between the splenial and 
dentary, showing the splenial to contribute to the hind part of the 
large and strong symphysis (s) supporting the lower incisors and 
canines. 

The upper incisors (Pl. [X.figs. 1 & 3, 71-5) are ten in number, 
five in each premaxillary, arranged in a semicircle or part of a 
broad ellipse (fig. 3,2). The bases of the crowns are in contact, 
save between the outermost and next tooth in advance. 

There is but little difference in size; the first and last in each 
premaxillary are the narrowest, the second and third the broadest ; 
the exserted crown is best preserved in the fourth and fifth of the 
right side (fig. 1), and in the fifth of the left side, the basal breadth 
being 24 millim., the length about 10 millim.; they all appear 
to have had the same simple laniariform character. When entire 
they passed in front of and covered or concealed the crowns of the 
lower incisors, the base of one of which is exposed behind the frac- 
tured crown of its homotype above. 

After an interval of 8 millim. extent the crown of the upper 
canine (ib. c) extends downward, and with a feeble curve backward, 
along a depression of like size and shape on the outer surface of the 
mandible, which gains breadth for the socket of the lower canine 

Tt 2 


264 PROF. OWEN ON THE ORDER THERIODONTIA, 


(fig. 2, c’) at the fore part of the depression for the reception of 
the upper one. 

The breadth of the base of the exserted crown of the upper canine 
is 5 millim.; it rather suddenly narrows to the pointed end; the 
length of the preserved exserted crown is 12 millim., a small part 
of the apex being wanting. The crown is slightly compressed, with 
a trenchant border on the hinder part which inclines somewhat 
inward; there is an indication of a fine crenation of this border. 

On the left side (Pl. IX. fig. 2) I had the outer alveolar wall of 
the canine removed and exposed its root (c), extending upward and 
slightly backward for twice the length of the exserted crown. The 
root, as it rose, slightly expanded beyond the breadth of the crown, 
and as gradually and slightly narrowed to the open end of the pulp- 
cavity. There is not a trace of a successional canine; and the con- 
dition of the pulp-cavity and petrified pulp, discoloured by the iron 
of its blood, indicates renewal of the working part of the laniary by 
continuous growth. In the course of this expository operation the 
exserted crown of the mandibular canine (ib. c’) was exposed, 12 
millim. in length, nearly that of the corresponding part of the maxil- 
lary canine, in front of which it extends, but along the inner or median 
level, so as to have been wholly concealed (as shown on the right 
side, fig. 1) by the alveolar plate continued from the upper canine - 
to the upper and outer incisor. There are few instances of carni- 
vorous air-breathers in which the mandibular teeth are so completely 
covered and hidden by the upper jaw, when the mouth is closed, as 
in this and other Theriodont Reptiles. 

After an interval about equal to that between the upper canine 
and outer incisor, the molar series (ib. figs. 1 & 2, m) commences, 
behind the canine. Of this series the crowns of five are exposed on 
each side of the upper jaw; they are all of the simple, slender, 
laniary type; the four anterior ones are divided by intervals of 
about the basal breadth of those teeth, which is 14 millim. in the 
first and second molars, and diminishes to 1 millim. in the fifth; 
the crown, inclining a little backward in the three hindmost, gra- 
dually narrows in each molar to a sharp point; the alveolar extent 
of the five molars is 7 millim. 

The toothless extent of the maxillary to beneath the fore border 
of the orbit is 13 milim. The upper dentigerous tract in a straight 
line from the foremost incisor to the hindmost molar is 45 millim. 

On a cursory comparison of the Theriodont genera and species 
now made known we discern a considerable range of variety in both 
size and shape. 

The extremes of size are exemplified by T%tanosuchus ferox * and 
Scaloposaurus constrictus tT ; those of shape by the flattened head of 
Galesaurus t and the compressed head of #lurosaurus (Pl. IX.). 
The skull of Procolophon § is broad in proportion to its length ; that 


* Quart. Journ. Geol. Soc. vol. xxxv. (1879), p. 189, pl. xi. 
Tt Catalogue, wt supra, p. 24, pl. xvi. figs. 10-15. 

t Quart. Journ. Geol. Soc. vol. xvi. p. 58, pl. ii. 

§ Catalogue, wt supra, p. 25, pl. xx. figs. 4-7. 


AND ON XLUROSAURUS FELINUS. 265 


of Gorgonops* is narrow; but the ordinal characters are manifest 
in all; only in the dwarfer and weaker species the relative size of the 
canines decreases, as is the case with similar carnivorous Mammals. 

Galesaurus planiceps and Aflurosaurus felinus form an equal-sized 
pair of these ancient Triassic precursors of our existing cats, or rather 
cat-like Marsupials. Galesaurus had 4—4 upper incisors like Dasy- 
urus; Ailurosaurus had 5— > upper incisors like Didelphys; but the 
molars of the foregoing and other Theriodonts indicate the lower or 
earlier type that bore them in their simple acuminate form as in the 
antecedent teeth; they had not advanced to the more complex modi- 
fied character shown in the molars of the most carnivorous of either 


12—12 
marsupial or placental Fere. Galeswurus had as many as ;—, of 


such molars ; in “lurosawrus they did not exceed, or at most by 


one, the = manifest in the fossil here described. 


Galesaurus, in the subject of the paper in the 16th volume of 
our ‘ Quarterly Journal,’ still has the advantage over all the subse- 
quently discovered Theriodonts in the entireness of the skull, espe- 
cially in the occipital region; and we may infer, analogically, a 
repetition of the reptilian characters of the cranium, indicative of 
low cerebral development, in its coordinates. 

If we next compare lurosaurus with the skull of Lycosawrus 
curvimola ~, which, at the date of its extrication, was the next in 
completeness to that of Galesaurus, we find the nearer affinity to 
Ailurosaurus in the small number of molars, in the general propor- 
tions of the skull, and in the extent and slope of the symphysis 
mandibule ; but the incisor-formula is Dasyurine, and the facial 
part of the septum narium is prominently manifested. But, of all 
the previously described genera of Theriodontia I deem Lycosaurus 
to have had the nearest kinship to Zlurosaurus. 

I am indebted to Sir Bartle Frere, K.C.B., for kindly taking 
charge of, and placing in my hands, the unique subject of the 
present paper £. 


EXPLANATION OF PLATE IX. 
Ltlurosaurus felinus, 


(All the figures are of the natural size.) 


Fig. 1. Right side view of the skull. 
2. Left side view of the skull. 
3. Under view of the skull. 


* Catalogue,ut supra, p. 27, pl. xxi. if Dbids pf pe 8: 
{ For the discussion on this paper, see p. 270. 


266 PROF. OWEN ON THE SKELETON OF 


20. Description of Parts of the SkeLEron of an Anomopont REPTILE 
(PLATYPODOSAURUS ROBUSTUS, Owen). Part. II. The Penvis. By 
Prof. Owsn, C.B., F.G.8., &e. (Read March 9, 1881.) 


[Puate X.] 


Tue pelvis of Platypodosaurus, which has been relieved from the 
matrix since the communication of the former paper*, includes. five 
sacral vertebre (Pl. X. fig. 1, 1-5), the right ‘* os innominatum,” 
62-64, and a large proportion of the iliac constituent of that of the 
left side. Of the first, or foremost, sacral vertebra, s', the part of 
the centrum in advance of the transverse processes, d!, is broken 
away, but so as to show the apex of the anterior conical articular 
cavity. The lumbar vertebra, which was articulated therewith, but 
had become dislocated therefrom, is preserved in the contiguous 
matrix ; and the shape and depth of the articular cavity of its centrum 
are exposed (Pl. X. fig. 5). 

The transverse process of the first sacral, d1, including both di- and 
pleurapophysial elements, is short and massive: measuring one inch 
and a half in antero-posterior diameter at its middle and narrowest 
part, it rapidly expands to its articulation (and what appears to be its 
confluence) with the ilium, 62. Behind the transverse process the cen- 
trum is moderately constricted, but expands to form the joint with 
the second sacral, of which joint the transverse diameter is 2 inehes. 
The breadth of the entire vertebra is 44 inches. 

The centrum of this second sacral vertebra (s 2) shows a greater de- 
eree of constriction and a narrower inferior or ventral surface. The 
transverse process (ib. fig. 1, d 2) extends from the anterior two thirds 
of the upper part of the side of the centrum, is narrower than that 
of the first sacral, but is longer, curving outward, and backward, ex- 
panded at both ends, but most so where it abuts against the ilium. 
The breadth of the second sacral is 5inches. The line of confluence 
with and abutment against the ilium is better marked than in the 
preceding vertebra. 

The centrums of the third and fourth sacrals are less expanded 
at their mutual junction than are those of the first and second 
vertebrae. The transverse processes are rather shorter, as in that 
of the third sacral, and this is narrower than those of the second 

and fourth vertebre ; that of the fifth sacral is the narrowest. 

Each transverse process of the sacrum expands at its outer end 
so as to touch the contiguous one where it joins or coalesces with 
the ilium. The foramina or vacuities so circumscribed decrease in 
size from the foremost to the third pair, and change the oblong for 
the subcircular figure. The hinder articular end of the fifth sacral 
centrum, s5, is broken away, but in a minor degree than the fore 


* Quart. Journ. Geol. Soc. for August 1880, vol. xxxvi. p. 414, plates xvi. & 


XV. 


Quart Journ Geol. Soc Vol XXXVIL PL % 


Mintern Bros imp. 


pLATYPODs \SAURUS ROBUSTU 
A g 


AS Foord delethth 


AN ANOMODONT REPTILE. 267 


part of the first vertebra; and a larger proportion of the arti- 
cular concavity 1s there preserved. The entire breadth of this ver- 
tebra is 44 inches. 

Turning to the dorsal aspect of the sacrum (PI. X. fig. 3), the 
hinder end of a lofty but narrow crest of bone, ns, due to confluence 
apparently of the neural spines of the fourth and fifth sacrals, has 
been worked out. The entire breadth of the fifth sacral vertebra is 
5 inches. The condition of the spines of the antecedent sacrums 
could not be determined. In the portion of a Dicynodont sacrum 
described in the ‘ Transactions of the Geological Society,’ 2nd series, 
vol. vii. pl. xxxiii. figs. 4 & 5, the short and thick spines of the 
first and second sacrals are distinct, not confluent. 

The section of the circular area of the neural canal of the fifth 
sacral of Platypodosaurus gives a diameter of 6 lines; that of the 
first sacral has a diameter of 9 lines; that of the last lumbar ver- 
tebra one of 8 lines. ‘Thus we have an indication of an expansion 
in the sacral region for the lodgment of the part of the myelon 
transmitting the nerves to the pelvic extremities, which accords with | 
the development of the limbs indicated by the femur of Platypodo- 
saurus *, Comparing the sacrum above described with that of the 
larger and more entire pelvis, the subject of a paper in the ‘ Geolo- 
gical Transactions’ above quoted, I incline to regard “ five” as the 
total number of the sacral or pelvic vertebree in Platypodosaurus *. 

The degree of coalescence of these vertebre is such as to sustain 

the anthropotomical ormammalian consideration of the coalesced mass 
as one bone or “sacrum ;” but the ventral outlets are relatively 
larger and the wings consequently less ossified. The general shape, 
moreover, is quadrate rather than triangular, with deeper lateral 
concavities between the subcarinate bodies and theiliac bones. The 
present Reptilian sacrum consequently comes nearer in shape to that 
of the Megatherioid mammals: but it includes fewer vertebral 
constituents. 
_ The length of such “sacrum” in Platypodosaurus is 74 inches ; 
its breadth, greatest at the third vertebra, is 54 inches; it is con- 
sequently more mammalian in character than is the sacrum of Dicy- 
nodon =. In this larger example of the South-African reptilian pelvis 
the sacral centrums are more constricted between the articular ends, 
which are concomitantly more expanded. The space between these 
sacral bodies and the ‘ossa innominata” is relatively greater; the 
transverse processes are consequently longer, and retain more of the 
ordinary reptilian rib-like character. 

The ilac bone in Platypodosaurus rapidly expands from its attach- 
ment to or confluence with the first sacral transverse process ; it ex- 
tends and curves forward, outward and dorsad, with the convexity 
mesiad, the concavity lateral. The extreme breadth is 43 inches; 
and the anterior margin of the ilium runs almost parallel with the 


* Loe. cit. p. 422, pl. xvii. figs. 6 & 7. 

t Compare also with page 40, plate xxxvi. (pelvis of Dicynodon), in the 
‘ Catalogue of Fossil Reptilia of South Africa, 4to, 1876. 

{ Loe. cit. 


268 PROF. OWEN ON THE SKELETON OF 


portion of the long, slender, last lumbar rib (PI. X. fig. 5, pl) here 
preserved. The outer border of the above expanse of the ilium is 
produced into a narrow margin; but this subsides at the level of the 
articulation with the second sacral vertebra, where the ventral border 
of the ilium becomes thick, smooth, and convex. It thins off to the 
dorsal border, which is here sharp; and the breadth of the ilium is 
reduced to 24 inches. As the bone recedes it assumes a columnar 
character, with an oval transverse section, gradually expanding, 
where the right innominatum has been fractured, opposite its junc- 
tion with the hinder sacrals, to a dorso-ventral diameter of 24 inches, 
and a transverse one of 13 inch (ab. fig. 4). 

The expansion of the bone after quitting the sacrum rapidly aug- 
ments to the acetabulum (ib. fig. 2a), to which it contributes the 
anterior and dorsal walls, the posterior and postero-ventral wall 
(completing the circumference of the cup) being formed by the ischial, 
63, and pubic, 64, constituents of the ‘os innominatum.” The 
breadth of the iliac part of the acetabulum is 4 inches ; the diameter 
of the outlet of the cavity is5 inches. There is a well-marked oblong 
fossa, 14 inch in breadth, at the bottom of the cavity. The depth of 
the cavity here is 13 inch ; and the wall is entire, and nearly an inch 
in thickness. 

The ischium, Y, contributes a rather larger share to the cavity 
(GeIe Gailss 1 & 2, 63) than does the pubis..; Beyond the acetabulum 
the pein contracts to a breadth of 2 inches 4 lines, where it may 
contribute to the foramen (ib. fig. 20). This is subcireular and an 
inch in diameter. Beyond the foramen the ischium loses. thickness 
and gains breadth; but the confluence with the correspondingly 
lamellate pubis is such as to leave no trace of their relative shares 
in forming either the above foramen or the ventral wall of the pelvic 
outlet. This wall, 6 inches in breadth, is strongly concave outwardly, 
convex towards the pelvic cavity; both surfaces are smooth; and 
the plate of bone so formed thins off to a symphysial border from 
from three to four lines in thickness, and probably four inches in 
extent. 

The pubis (ib. figs. 1& 2, 64) gradually thins and auynik as it passes 
from the acetabulum to the foramen, o. ‘The border forming the 
subacetabular *‘ brim of the pelvis ” is from 7 to 8 lines in thickness ; 
its border has been chipped; but, though not entire, it is free from 
any indication of pectineal process, or of a prominence for the 
support of a marsupial bone. The utmost care in exposing this 
part of the pelvis failed to bring to light any such bones, or portions 
of them. | 

An accidental fracture, after exposition of the above-described part 
of the pelvis, about 2 inches in advance of the acetabulum, gives the 
transverse section (Pl. X. fig. 4) of that most contracted part of the 
iliac constituent. 

The dorso-ventral diameter of the anterior outlet of the pelvis is 
8 inches; the extreme transverse diameter is 54 inches. 

The place of the ischiadic notch, 2, which in most Bruta becomes 
a foramen, is marked only by a feeble concavity of the postaceta- 


an ae 
a 


AN ANOMODONT REPTILE. 269 


bular border of the ischium : the “‘ tuberosity ” is indicated at ¢, fig. 2. 
The extensive ischio-pubic symphysis (fig. 2, y,y) may have been 
obliterated by continuous ossification, as in the pelvis of Dicynodon 
above compared*, where it projects, ridge-like, between the broad, 
outwardly concave, ischio-pubic plates. The parts of the dorsal 
surface of the plates of Platypodosaurus exposed in the block of 
tmatrix show the hinder half of a continuous crest of bone (1b. fig.3,7s), 
due to confluence of the neural spines of the third, fourth, and fifth 
sacrals. It may have been continued further forward. 

The last lumbar vertebra (ib. fig. 5) shows the hinder concave 
articular surface of the centrum, ¢, which is 2 inches 3 lines in dia- 
meter. The margin of the concavity is thick and convex, as in the 
dorsal vertebra (loc. ct. p. 414, pl. xvi. fig. 3) of the preceding paper 
on Platypodosaurus. The neural canal, m, is 7 lines in diameter at 
its hinder outlet, m. The neurapophyses, x, n, rise to 10 lines before 
sending off the diapophyses, which are broad and flat above, and 
give attachment to a slightly curved pleurapophysis, pl, 45 inches 
in length and 9 lines in breadth; at its free end it almost comes 
into contact with the iliac labrum. 

Of all the examples of pelvic structures in extinct Reptilia which 
have come under my observation, the type of pelvis exemplified in 
Platypodosaurns departs furthest from any of the modifications of 
that part of the skeleton known in existing Reptilia, and at the same 
time makes the nearest approach to the Mammalian pelvis. This is 
seen, not only in the number of sacral vertebre, but in their breadth, 
due to the outward extension of their expanded transverse processes. 
The Mammalian character is more marked by the breadth of the iliac 
bones, and by the extent of confluence of the similarly expanded 


ischia and pubes, together with their further confluence at the 


ischio-pubic symphysis (which, though the bones are here fractured 
in the present example, may be inferred to have existed from the 
general resemblance of the pelvis of Dicynodon to that of Platypodo- 
saurus). 

The small solution of continuity between the ischium and ilium 
shown by the foramen (0, fig. 2, Pl. X.) closely accords with the 
foramen in a similar part of the pubis of many modern lizards, which 
gives passage to the “superficial femoral artery.” It may, however, 
if the ischium be actually continued into its periphery in Platypo- 
dosaurus, be regarded as a trace of a “‘ foramen obturatorium.” 


EXPLANATION OF PLATE X. 
Platypodosaurus robustus, 
Fig. 1. Front view of pelvis: + nat. size. 
2. Side view of acetabulum, ischium, and pubis: i nat. size, 
3. Back view of sacrum and iliac bones: +} nat. size. 
4, Section of ilium anterior to the acetabulum: nat. size. 
5. Back view of last lumbar vertebra: nat. size. 


* Loe. cit. 


270 PROF. OWEN ON THE SKELETON OF 


Discussion. 


Mr. Hurxe asked if the lateral view of the pelvis of Platypodo- 
saurus really bore out the view of the author as to its remarkable 
Mammalian affinities, and especially whether the acetabulum is in 
front of or behind the vertical of the sacro-iliac Junction. He further 
remarked that most reptiles have not one but many series of teeth ; 
and such would seem to have been necessary for carnivorous animals, 
in which the teeth are continually lable to damage. He stated that 
the canine described in this specimen seemed different from what had 
been described as characteristic of the order Theriodontia. 

Prof. SreLey inquired whether the ilium of Platypodosaurus re- 
sembled that of a Seal. So far as he could judge from the single 
diagram, the whole pelvis closely resembled that of Dicynodon ; and 
if so, its affinities with other orders of fossil reptiles were more 1m- 
portant than its presumed analogies with the Mammalia. This type 
of pelvis was approximated to by certain Dinosaurs, and, but for 
their prolonged iliac bones, was even more closely paralleled by the 
pelvis of some Solenhofen Pterodactyles. Even should the marsu- 
pial bones suspected by Prof. Owen have existed, the character 
need not be mammalian, since it is well developed in the Pterodac- 
tyle group. ’ 

With regard to the Theriodontia, he was unable to admit the 
importance of the characters and presumed characters on which 
the order was founded. The teeth of existing Lizards would, by 
the variety of types which they present, as well sanction ordinal 
subdivisions as the Anomodontia. The Dinosaurs also, with their 
carnivorous and herbivorous types of teeth, presented greater variety 
among themselves than that which was held to separate the fossil 
described from its Anomodont allies. He could not see how the 
circumstance of the so-called canine tooth being better nourished 
and growing larger could in these animals be an ordinal character. 
He thought it premature to infer the absence of successional teeth 
as an ordinal character—because the animals described were mature, 
and succession of teeth may well have occurred in early life. Not 
only were there no such characters dividing these animals from 
Anomodonts as separate Crocodiles from Lizards or Turtles from | 
Crocodiles, but there was absolutely no important difference of plan 
in the structure of the skulls of Anomodonts and the so-called 
Theriodonts. Prof. Owen had long ago classed these animals as 
Cynodontia, forming a family of the Anomodonts; that classification 
he thought excellent, but he could not accept the family as an order 
or adopt its new name. 

Mr. Twetverreers had just returned from the district of widely 
spread Permian deposits of Russia. So far as he knew, these rocks 
had only exhibited two types of Reptilian structure, the Labyrintho- 
dontia and the Theriodontia. He asked for information as to the re- 
lations between the latter and several genera described by Prof. Cope. - 

Prof. Owen said that the sacrum of the Dinosaurs approached 
that of birds rather than that of mammals. He was not aware of 


AN ANOMODONT REPTILE. Dal 


any Pterodactyle pelvis which resembled that of Platypodosaurus. 
He admitted that the dentition of the Lizards varied, but asserted 
that it never approached to that of the Theriodontia, in which the 
teeth were not ankylosed to the jaw. He was not able to compare 
these forms with the genera of Prof. Cope, in the absence of suffici- 
ently large and detailed figures to illustrate the descriptions given 
by that naturalist. 


272 G M. DAWSON ON THE SUPERFICIAL GEOLOGY OF 


21. AppITIONAL OnsERVATIONS on the SUPERFICIAL Guotoey of BririsH 
Cotompra and apsacENtT Reeions. By Groraz M. Dawson, 
D.Se., F.G.S., Assoc. R.S.M., Assistant Director of the Geological’ 
Survey of Canada. (Read March 9, 1881). 


ConrTENTs. 


Observations on the Southern part of the Interior of British Columbia. 
Observations north of the 54th parallel in British Columbia. 

Peace and Athabasca Basins. 

Additional Notes on the Coast. 

Glaciation of the Queen-Charlotte Islands. 

General Remarks and Conclusions. 


In two papers previously communicated to the Geological Society, 
the results of observations on the glaciation of the northern portion 
of the American continent from Lake Superior to the Pacific have 
been given*. The geological work of which these observations 
formed a part was carried on first in connexion with the North- 
American Boundary-Commission Expedition, and subsquently on the 
Geological Survey of Canada. In continuing the exploration of 
British Columbia on the Survey last named, during the seasons of 
1877, 1878, and 1879, many additional facts of interest have been 
gathered, which it is proposed here briefly to summarize and discuss 
with special reference to the second of the two papers above men- 
tioned, in which a description of the salient physical features of the 
province of British Columbia has been given, and a map published ; 
to these, which it is unnecessary here to repeat, reference should be 
made in considering the points now brought forward. 


Observations on the Southern part of the Interior of British Columbia. 


In the more detailed examination of that part of the southern 
portion of the province extending from the Fraser eastward to the 
Gold ranges, and including the whole breadth of the region formerly 
called the interior plateau, traces of a general north-to-south 
glaciation have been found in a number of additional localities at 
high levels ; and it would appear that the ice, whether that of a great 
glacier or water-borne, pressed forward to, or even beyond, the line 
of the 49th parallel, notwithstanding the generally mountainous 
character of that part of the country. With the facts previously 
recorded, these now extend the known area of north-to-south 
glaciation to a portion of the plateau over 400 miles in length. 

The most striking instance of this general glaciation, and that 
which carries it up to a height greater than elsewhere observed, 
is met with in the case of Iron Mountain at the junction of the 
Nicola and Coldwater rivers. This mountain is one of the more 
prominent points of that portion of the plateau, which, toward the 
eastern or inland borders of the coast-range, becomes rough and 
broken. It rises in a broad dome-like form to a height of 3500 


* Quart. Journ. Geol. Soc. vol. xxxi. p. 603, and vol. xxxiv. p. 89. 


— 


BRITISH COLUMBIA AND ADJACENT REGIONS. 273 


feet above the neighbouring river-valleys, or 5280 feet above the 
level of the sea. Its summit has been heavily glaciated, the projecting 
rocky masses being worn into ridges parallel to the direction of ice- 
movement, the indicated direction of which is nearly parallel to a 
bearing N. 29° W. to 8.29° E. If not due to the general glaciation, 
these markings can have been caused only by ice from the coast- 
ranges; and though ice has flowed from these as from the. other 
mountain masses of the province during the later portion of the 
elacial epoch, I believe the situation of Iron Mountain to be such as 
to preclude altogether this mode of explanation. The mountains of 
the coast-ranges are neither high enough nor so near as to supply a 
body of ice capable of overriding it. 

On the plateau south of Kamloops glaciated surfaces have been 
found in several places at an elevation of about 3200 feet above the 
sea. The locality is far removed from any mountain-ranges capable 
of giving rise to extensive glaciers, being situated in the very centre 
of the interior plateau. The rocks are broadly ice-shaped and not 
unfrequently polished, more rarely distinctly striated. The direction 
of movement varies from 8. 6° E. to 8. 27° E. On another part of 
the plateau, north of the course of the upper part of the Nicola 
River between Stump and Douglas Lakes, at an elevation of about 
3622 feet, are glacial traces similar to the last, consisting of polishing 
and striation without fluting, having a general direction of 8S. 9° E. 
Still another instance of this general glaciation is found on the 
eranite rocks near Chain Lake, between Lake Okanagan and the 
Similkameen River, in latitude 49° 40’ N. Here, asin the cases before 
mentioned, the circumstances seem entirely to preclude any expla- 
nation by local glaciers, as the portion of the plateau on which it 
occurs is fully up to the general level, and surpassed only by a few 
insignificant hills at a considerable distance. The rock-surfaces 
are beautifully polished, and show striation varying in direction 
between 8. 20° E. and S. 28° E., but no deep grooving. The ele- 
vation is 4075 feet. 

The Okanagan valley has been alluded to in the paper already 
referred to as the most important southern gateway of the interior 
plateau. The bottom of this valley, where it crosses the 49th 
parallel, is about 860 feet above the sea-level. It 1s wide, and must 
at one time have been much deeper, as its rocky floor 1s not now seen. 
It occupies the axis of a general depression of some magnitude, and 
appears to have carried the drainage of a great part of the interior 
of British Columbia at a former period. ‘This valley has probably 
been subject to heavy ice-action during the time of general glaciation ; 
but to what extent the features now found may bé due to this, and 
in how far to a subsequent period when, as a narrow arm of the 
sea or of a great lake, it carried southward ice produced by glaciers 
nearer the mountains, it is now difficult to ascertain. Glacial 
striation was observed descending obliquely from the sides toward 
the centre of the valley, and also in several places in the valley 
itself, but in both cases without distinct grooving. The rocks of 
the sides of the valley are often distinctly moutonnées; and, as seen 


274 G. M. DAWSON ON THE SUPERFICIAL GEOLOGY OF 


from a distance, those on the lower part of the slopes show 
flattened outlines, while those higher up are more abruptly rounded 
and have not been so thoroughly ground down. 

The general statements made in a former communication, in 
reference to the covering of Boulder-clay or unmodified drift spread 
over the entire area of the interior plateau, are borne out in the region 
now more particularly in question. From the rearrangement of 
this material the great systems of terraces subsequently mentioned 
have been formed. 

Details need not be given of the evidence in striation and rock- 
polishing of the existence of glaciers radiating from the various 
mountain-systems, though it may be mentioned that some of these 
seem to have had a very great extension down the lower valleys. 

In this southern portion. of the interior plateau, terraces are 
exhibited on a scale scarcely equalled elsewhere. ‘They border the 
river-valleys, and at greater elevations are found expanding beyond 
these and attached to the higher parts of the plateau and mountains.. 
None has yet been found here, however, equal in height to that 
previously described on [1-ga-chuz Mountain in the north at 5270 
feet above the sea. Many of the terraces and “ benches” of the 
valleys may be the result of the gradual cutting-down of the river- 
course in the hollow previously filled with glacial débris ; but for others, 
including more particularly those of the higher levels, no explanation 
short of the complete flooding of the plateau-region will suffice. 
Knowing therefore that the water must have stood successively at 
every lower level, it is of comparatively little importance that in the 
case of some of the lower terraces it becomes impossible to determine 
whether they belong to this period of the retreating waters or to a sub- 
sequent river-erosion. 

In this region the terraces frequently surpass 3000 feet in eleva- 
tion above the sea-level. The more prominent of those seen on the 
southward slope of Iron Mountain may be taken as an example 
of the arrangement of these old water-marks. These terraces are as 
follows, the approximate heights being given in feet—2386, 3063, 
3392, 3611, 3715. It is frequently observed, however, that the 
occurrence of a terrace at any particular level is merely a matter 
of local circumstance, probably dependent on the supply of material 
and other such causes; and in different places not very remote the 
the scale of terraces often differs. This is illustrated on Okanagan 
Mountain, situated east of the lake of the same name. On the 
south side of this elevation the principal terraces were baro- 
metrically determined as follows—1862, 2042, 2141, 2645, 2800, 
2839 feet; on the northern slope six principal terraces were again 
observed, as follows—1451, 1579, 1962, 2452, 2553, 2879 feet. 

The wide trough-like valleys which traverse the plateau are, over 
a considerable portion of its extent in the southern part of the province, 
partly filled with a deposit of white silt or loess-lke material com- 
parable with that described under the same name in the Nechacco 
basin to the north*. It is, however, unconnected with the latter, 

3 * Quart. Journ. Geol. Soc. vol. xxxiv. p. 105. 


BRITISH COLUMBIA AND ADJACENT REGIONS. 275 


and at a considerably lower elevation, reaching a maximum height 
of about 1700 feet above the sea. In the vicinity of Kamloops 
Lake and in the South Thompson and Okanagan valleys, it is well 
shown, generally forming the first terraces above the rivers. In 
origin it is probably, like that of the Nechacco region, a deposit from 
the turbid waters flowing from glaciers at a time when these had a 
considerable extension from the various mountain-ranges. At this 
time, either from general depression of the land, or the damming 
of the valleys by ice or moraines, a system of winding water-ways, 
lakes or fiords, must have occupied the main valleys. The heads of 
these valleys in the Gold ranges still hold long and deep lakes, on 
the banks of which, where they have been examined (more parti- 
cularly in the Shuswap region), drift deposits are comparatively un- 
important, and the white silts arenot found. The fine silty material 
must have been deposited in somewhat tranquil waters; but it 
appears difficult to explain its absence from the valleys on the 
flanks of the Gold ranges. It may be suggested that the currents 
in the upper parts of the valleys were so strong as to prevent 
the deposition of the silt; but, apart from the difficulty found in 
supposing such great bodies of water as the valleys must have held 
at this time to be in rapid motion, there is no such sudden widening 
in the valleys at the points at which the silt commenees as might 
account for the slackening of the current. 

It is perhaps on the whole most probable that the basins now 
occupied by the Shuswap lakes and others in a like position were 
filled with glacier-ice, from which the water flowed down the long 
valleys, while the abrasion of the rocky beds of the glaciers supplied 
in large quantity the material of the silt deposits. From the height 
at which the silts occur, their greater coarseness in the lower part 
of the Okanagan valley, and the evidence of current-action in that 
valley near Osoyoos Lake, it is probable that this depression has 
served as the main outflow of the white-silt lake or sound. At the 
last 16 would appear that the glaciers retreated with considerable 
rapidity, becoming extinct or dwindling to nearly their present 
size, and leaving the upper portions of the valleys which penetrate 
the Gold ranges almost free from débris and ready to form the 
basins of the lakes which now generally occupy them. 

The explanation here adopted to account for the existence of these 
lakes will, I believe, be found applicable to many in other parts of 
British Columbia, and is again referred to on a subsequent page. 
It is the same advanced by A. Helland for Norwegian lakes *. 
Whether any of the lakes in the region now in question lie in rock 
basins of glacial formation has not been determined, as the valleys 
below their outlets are generally filled to an unknown depth with 
detrital materials. 


Observations north of the 54th parallel in British Columbia. 


An exploratory survey of the remote region lying between the 
54th and 56th parallels in British Columbia and of part of the 


* Quart. Journ. Geol. Soc. xxxiil. p. 165. 


276 G. M. DAWSON ON THE SUPERFICIAL GEOLOGY OF 


Peace and Athabasca river-basins to the east of the Rocky Moun- 
tains, enables the characters of glacial evidence to be defined further 
north, and has aided in the decision of some theoretical points 
referred to in the sequel. Most of the facts observed to the west of 
the Rocky Mountains resemble so closely those previously described 
for the regions south and east of this that they do not require 
lengthened notice. The southward or south-eastward passage of 
glacier-ice in the valley of Babine Lake is indicated by glacial 
grooving, while the valley of the Skeena has formed a main channel 
of discharge of glacier-ice toward the coast. In the mountains 
between the valley of this river and Babine Lake a somewhat irre- 
gular, but still, I believe, distinct terrace-flat was observed on the 
watershed at an elevation of 4800 feet. Its surface is strewn with 
water-rounded stones differing from those of the mountains of the 
vicinity. The region north-east of Stuart Lake, extending to 
M‘Leod’s Lake and the Parsnip River at the base of the foot hills of 
the Rocky Mountains, is deeply drift-covered, the surface consisting 
either of Boulder-clay charged with erratics of varied origin, or 
terrace-flats formed by its rearrangement. This region lies to the 
north of and somewhat higher than the Nechaco basin, which is 
characterized by the white silts of a former paper*. ‘The highest 
part of its surface crossed by the trail has an elevation of 2900 
feet. 

In the valley of the Misinchinca, flowing westward from the 
summit of the Pine pass of the Rocky Mountains, glaciation was 
observed in a few places parallel to the direction of the main de- 
pression. In the Pine-River valley, draining eastward and joining 
the Peace, no glaciated surfaces were seen—a circumstance which 
may arise from the comparatively soft character of the rocks. 


Peace and Athabasca Basins. 


In the comparatively level country drained by the Peace and 
Athabasea rivers, to the north-east of the mountains, underlain 
by unaltered rocks of Mesozoic and Tertiary age, the chief evidences 
of the glacial period are found in the distribution of erratics, and 
the existence of extensive “drift” deposits. In travelling eastward 
from the mountains by the Pine-River valley, a remarkable absence 
of such deposits is noted in that part of the valley which traverses 
the eastern foot hills ; but at the Middle Forks the plateau, with an 
elevation of 1000 feet above the river, or 3000 feet above the sea, 
and at a distance of thirty miles from the indurated rocks of the 
mountains, is strewn with rounded pebbles of quartzite &e. from 
these rocks, though material of local origin preponderates. Highteen 
miles further east, at the Lower Forks, the superficial deposits are 
much more important, covering the surface of the plateau to a 
considerable depth, and consisting of gravelly beds passing upwards 
into finer silty materials; the elevation of the plateau is here 2350 
feet. In continuing eastward after passing over a summit of 3300 
feet on the line followed, Laurentian boulders which must have come 

* Quart. Journ. Geol. Soc. vol. xxxiv. p. 105. 


BRITISH COLUMBIA AND ADJACENT REGIONS. 277 


from the axis of these rocks to the east or north-east were first ob- 
served, and appear in abundance, at a height of from 2300 to 2500 
feet, near the D’Echafaud River, in latitude 55° 45’, longitude 120°. 

Kast of this point the wide Peace-River plateau extends, and the 
general character of the country in regard to its superficial deposits 
is so uniform that it is unnecessary to particularize localities in de- 
scribing it. Its surface is so thickly covered that exposures of the 
underlying rocks are, as a rule, found only in the larger river-valleys. 
The lower layers of the drift appear to represent the Boulder-clay of 
the great plains to the south and east and the northern part of British 
Columbia to the west; they are sandy clays with boulders and ~ 
stones in abundance, and their upper surface is somewhat irregular, 
rising in some places in ridges or broad gentle elevations, which 
stand out above the newer silty deposits in which a great part of 
the surface is enveloped. ‘The silt is generally pale grey or fawn- 
colour, and while in places passing almost into clay, becomes occa- 
sionally a fine sand. This sandy covering of the surface is found 
especially at the southern rim of the Peace basin, near the Atha- 
basca, where the plateau attains an elevation of about 3300 feet 
(long. 117°). The ridges at this elevation are still thickly strewn 
with Laurentian boulders. 

In regard to the material of the drift, the stones and boulders 
scattered over this great district are, in part, those of the Rocky Moun- 
tains to the west, in part derived from the Laurentian axis to the 
north and east. ‘The fragments from the first-mentioned source are 
generally of quartzite; the limestone and other softer rocks accom- 
panying these in abundance in the vicinity of the mountains, de- 
creasing rapidly as we recede from them. The Laurentian mate- 
rial is chiefly gneiss and granite of the usual well-marked types. 
Between the Athabasca and Saskatchewan (long. 113° 30’) the 
plateau attains a maximum elevation of about 2300 feet, and 
Laurentian boulders are everywhere exceedingly numerous. 


Additional Notes on the Coast. 


In the fiords penetrating the coast of the mainland of British 
Columbia, and channels intervening between the numerous islands 
lying off it, from the southern extremity of Alaska to the north end 
of Vancouver Island, marks of the passage of glacier-ice are to be 
found wherever the rocks are unweathered (see Map, p. 278). These 
marks are generally in strict conformity with the directions of the 
passages, which it is evident must have been filled with ice moving 
in the main seaward from the coast-ranges, in which many smaller 
glaciers are still found. Whether at any time the supply of ice has 
been so great as to form a confluent mass flowing toward the sea, at 
right angles to the general direction of the coast mountains, and with- 
out regard tothe smaller features of the surface, has not been definitely 
ascertained ; but it is highly probable that this has happened. The 
outer islands of the Shore archipelago have scarcely been examined ; 
but the little group called the Gnarled Islands (lat. 54° 39’), on the 
south side of the strait, thirteen miles wide, which lies between 


Q.J.G.S. No. 146. U 


278 G. M. DAWSON ON THE SUPERFICIAL GEOLOGY OF 


Dundas Island and Cape Fox at the southern extremity of Alaska, 
shows heavy grooving from N. 50° E. to 8. 50° W., proving that 
this strait must have been filled with ice. 


Sketch Map of part of British Columbia, showing the supposed exten- 
sion and general direction of flow of the glacier-ice when near its 
maximum limit. 


32° 
USAMA 
Fd ME Vii 
ra € y 


TuwHa Naga! 


SQNV1St a.t/Lo 


~S ud 
KA oho vtlll lll 
SAIS SYD VAI 
AIT IAAT IT SSE 


The arrows indicate the direction of flow of the ice. 
The dotted line shows the seaward margin of the confluent glacier. 


The scarcity of examples of well-marked terraces on the coast, 
and the comparatively small elevations at which they are found, has 
been remarked previously. At Fort Simpson, however, in lat. 54°34’, 
the surface bears a considerable thickness of detrital matter, and from 
a distance this appears to form an ill-defined terrace at a height of 
somewhat over 100 feet. A few miles further southward, at Metla- 
katla, there is a well-marked terrace, flat, with an elevation, baro- 
metrically determined, of 95 feet above high-water mark. 

In the previous paper, already several times referred to, evidence 
was brought forward in favour of a belief that during a part of the 
glacial period a vast glacier filled the entire Strait of Georgia, which 
separates the south-eastern part of Vancouver Island from the main- 
land, and that the ice swept across the south-eastern extremity of the 


BRITISH COLUMBIA AND ADJACENT REGIONS. 279 


island, and may even have passed some distance southward to Puget 
Sound, and westward by the Strait of Fuca. It still remained, how- 
ever, to determine whether the ice supply of this glacier was wholly 
derived from the neighbouring mountainous country, or whether (as 
might be supposed in accordance with some theories of glaciation) a 
great ice-sheet entered at Queen-Charlotte Sound, and passed con- 
tinuously southward between it and the mainland. Itis now found 
that the last-mentioned idea must be abandoned. In several places 
about the northern end of Vancouver Island, but more particularly 
on the little islands of the Masterman group near Hardy Bay, and 
on those in Beaver Harbour, are marks of very heavy glaciation 
from south-east to north-west, in bearings varying from N. 49° W. 
to N. 62° W. This not only passes over the islands, but has grooved, 
polished, and undercut vertical, or nearly vertical, faces on their south- 
eastern parts, while the north-western slopes are comparatively rough. 
These traces precisely resemble those found in the track of the 
Strait-of-Georgia glacier near Victoria*, and show that here, as 
there, the ice rode over the low extremity of Vancouver Island. 
The seaward margin of the continental shore is here also low, and 
the width of the glacier of Queen-Charlotte Sound can scarcely have 
been less than twenty or twenty-five miles, and may have been much 
greater. 

Some additional evidence of the movement of the upper parts of 
the Strait-of-Georgia glacier has been found at Nanaimo, on the 
inner coast of Vancouver Island, sixty miles north-west of Victoria. 
Hard sandstone rocks which have been bared on the colliery railway 
show heavy glacial grooving running parallel to the general trend 
of the coast and Strait of Georgia in such a way as to prove that 
the entire strait must here also have been filled with ice. No 
local glaciation, which would radiate from the mountains of the 
district, can account for the facts. In clays resting on these gla- 
ciated rocks, shells like those formerly observed at Victoria were 
found, a small collection comprising Sawicava rugosa, Mya truncata, 
and Leda fossa. The height of the locality is about 70 feet above 
the sea. 

Between Vancouver Island and the mainland, on both sides of 
the central region from which the ice spread in two directions to 
form the Queen-Charlotte-Sound and Strait-of-Georgia glaciers, well- 
stratified deposits of clays and sands occur, in some places forming 
cliffs 200 feet in height. In the course of the Queen-Charlotte- 
Sound glacier, Cormorant Island may be cited as an example of 
these deposits ; and in that of the Strait of Georgia, Harwood, Mary, 
Hernando, and Savary Islands. These deposits resemble those of 
Victoria, New Westminster, and the islands in the southern part of 
the Strait of Georgia previously described, but imply for the period 
of their formation a decreased length in the glacier, from its point of 
maximum extension, of at least 100 miles. Harwood, Mary, Her- 
nando, and Savary Islands lie about the entrance of Bute and 


* Quart. Journ. Geol. Soc. vol. xxxiv. pp. 94, 96, 99. 
v2 


280 G. M. DAWSON ON THE SUPERFICIAL GEOLOGY OF 


neighbouring inlets in such a position as to suggest that they may 
in part represent a moraine marking a stage in retreat of the ice. 
They form the projecting points of a comparatively shoal bank off 
these inlets, which, in their upper parts, are very deep. Boulders 
here occur in great abundance on the beaches, and are probably 
derived from a Boulder-clay or morainic material underlying the 
well-bedded deposits. 


Glaciation of the Queen- Charlotte Islands. 


These islands were the subject of geological examination in 1878. 
They form a compact archipelago widely separated from the southern 
extremity of Alaska to the north, and the western coast of British 
Columbia to the east, and may be regarded as a partly submerged 
mountain system, the axis of which lies in a N.N.W.-S.S.E. bearing. 
In its central part summits surpassing 4000 feet, and still bearing 
patches of perennial snow, are frequent, but it falls at both ends. 
On the north-east side of the mcuntain axis, at its north end, is a 
wide triangular attachment of flat land forming the greater part of 
Graham Island. 

In these islands we find everywhere evidence of the descent of 
glacier-ice from the mountains toward the sea, but (with one im- 
portant exception subsequently noticed) none of the passage across 
the group of any more ponderous ice-mass. The channels and fiords 
penetrating the southern portion of the islands show in general di- 
stinct and heavy glaciation which has evidently been local in cha- 
racter, the scoring and grooving being parallel to the main directions 
of the valleys, and changing with their course. In Houston- 
Stewart Channel, separating Prevost and Moresby Islands, the ice 
has evidently flowed from the axial mountains both eastward and 
toward the open Pacific to the west. Many of the boulders of the 
beaches are distinctly glaciated, and, as they lie in some places rudely 
packed together, seem to have been little disturbed since they were 
deposited by the ice. Sands, clays, and other detrital deposits re- 
ferable to the period of glaciation are here almost entirely wanting, 
and the water round the coast is deep.’ 

Further north, near Laskeek, where the width of the islands 
becomes greater, there is evidence, i in the comparatively slight de- 
gree in which the rocks at the outer ends of the inlets are slaciated: 
that the glaciers did not long stretch much further out than the pre- 
sent coast-line. At Cumshewa Inlet (lat. 53°), and further north at 
Skidegate Inlet, the character of the coast changes, becoming low; 
but both these inlets still head in the high axial mountains of 
the group. ‘Traces of the glaciers of these inlets are found nearly 
to their mouths ; but while the upper parts are still deep and fiord- 
like, they are partly blocked at their seaward extremities by trans- 
verse bars, and shallow water extends far off shore. 

Further north a series of fiord-like valleys are still found pene- 
trating the eastern side of the mountainous axis of Graham Island, 
and the shoal-water found off Cumshewa and Skidegate is repre- 


BRITISH COLUMBIA AND ADJACENT REGIONS 281 


sented by the wide stretch of flat land before alluded to. Several 
of the fiords here open together into a large sheet of water forming 
the upper part of Masset Inlet, which communicates with the sea to 
the north by the long narrow passage known as the Masset Sound. 
The fiords are heavily glaciated, bordered in most places by steep 
rocky shores, deep and free from drift deposits, and contrast in these 
respects markedly with the low-shoal eastern shores of the Masset 
expansion into which they open. 

The composition of the low land to the east and north-east is 
best shown in the cliffs forming its eastward-facing margin. A few 
miles north of Skidegate a low cliff or bank shows deposits which 
are evidently of glacial age, cut off above by a gently undulating 
surface of denudation, above which is 10 or 15 feet of material 
which shows no sign of blending with that below. The upper de- 
posit consists of sand and well-rounded gravel in regular and often 
nearly horizontal layers. It has here become in many places quite 
hard, being apparently cemented by ferruginous matter. Its lower 
layers hold small boulders, a few of which are from 18 inches to 2 feet 
in diameter. The lower deposit in one place is a typical Boulder- 
clay, with many half-rounded or subangular stones and occasional 
boulders of some size. The matrix is bluish grey, hard, and some- 
what arenaceous, the whole being irregularly mingled, and having 
no distinct bedding. Ata short distance this Boulder-clay begins 
to show bedding, and to become interleaved with hard clayey gravels 
composed of well-rounded pebbles. The stratification of these is 
undulating and rather irregular, and there is some local unconformity 
by erosion between the different layers. A few paces still further 
on these become interbedded with, and are eventually replaced by, 
hard, bluish-grey, arenaceous clays, which hold some pebbly layers 
and an abundance of broken specimens of mollusks, among which 
Leda fossa is the most common. A small Cardium-like shell and 
fragments of a Balanus were also observed. 

Further north on this coast the clays, with the overlying sandy 
deposit in greater or less thickness, form long ranges of cliffs ; and 
though locally irregular, their general character continues the same. 
The clays are, in some places, very hard, and were observed to hold 
fragments of trees quite brown in colour, but not mineralized. 
These deposits, as a whole, very closely resemble those previously 
described as occurring at Victoria, on the south-eastern extremity 
of Vancouver Island. 

Lying like Masset Inlet near the junction of the hilly and low 
countries is Naden Harbour, and between this and Masset Inlet are 
two large freshwater lakes, which doubtless occupy an analogous 
position, but have so far not been visited by any but Indians. 
Southward there is reason to believe that there are one or more 
basins in a similar relation between Masset Inlet and Skidegate. 

Boulders are very numerous on the coast of some parts of the 
northern portion of Graham Island; and these and the beach- 
gravel are in many cases formed of rocks which must have been 
transported from the mainland to the north or east, and quite unlike 


282 G. M. DAWSON ON THE SUPERFICIAL GEOLOGY OF 


those of the Queen-Charlotte Islands. Similar erratics appear to 
characterize in greater or less abundance the whole of the low 
country above described, but are not found about the heads of the 
south-western extremities of Masset Inlet. 

It has previously been shown that at the time when the Strait-of- 
Georgia glacier began to diminish the sea must have stood consider- 
ably higher in relation to the land than at present, and the glaciated 
rock surfaces about Victoria and Nanaimo no sooner appeared from 
beneath the glaciers than they were covered by deposits holding 
marine shells. Such must have been the state of affairs also in the 
Queen-Charlotte Islands ; and to this time are doubtless to be referred 
the clay and sand deposits of the low north-eastern part of Graham 
Island above described. The material of these must have been sup- 
plied from the glaciers of the islands themselves, but added to also (as 
the nature of the boulders proves) by the debris borne on floating ice 
from the larger glaciers of the mainland, the sea levelling and 
spreading abroad the material, and preventing the formation of any 
well-marked terminal moraines by the island glaciers. The rocky 
beds of the fiords and Masset-Inlet expansions must have been shaped 
to some extent by the ice; but the absence ef drift material from their. 
areas, and especially of the erratics derived from the mainland, are, 
with their situation, good reasons for supposing that they mark the 
regions last covered by glacier-ice, and from which it eventually re- 
treated with some rapidity, leaving the hollows formerly occupied 
by it to become first inlets, and then, with increasing elevation, in 
some instances lakes. 

The exceptional case which seems to show the impingement on the 
Queen-Charlotte Islands of ice not produced on them was found on 
the north coast on the little islands lying outside the entrance to 
Masset Inlet; but it is probable that similar traces might be found 
by search in additional localities in this vicinity. Wider exposures 
of basalt a few feet above high-water mark here show very heavy 
though somewhat worn glaciation in a direction 8. 10° E., or N. 
10° W., but probably the former. The depth and parallelism of the 
grooving would appear to show that it is glacier work. The moun- 
tainous axis of the islands in this their northern part does not ex- 
ceed in height about 1300 feet, and where nearest is about 15 
miles from the locality, while the direction of the marking is not 
that which would be followed by ice descending from the mountains 
under any circumstances, being more nearly parallel to than radiant 
from them. It is, however, just that which ice-masses floating up 
or down the strait separating the islands from the mainland must 
have taken, or glacier-ice pushing southward from the long fiords 
of the Prince of Wales group in Southern Alaska, sixty miles 
distant. It may, I believe, be attributed with greatest probability 
to the last-named agent; and in view of the great extension which 
the glaciers of other parts of the coast must at one time have had, 
that required for the Prince of Wales group and adjacent channels 
does not appear excessive. 


BRITISH COLUMBIA AND ADJACENT REGIONS. 283 


General Remarks and Conclusions. 


It is somewhat difficult to connect the various observed facts of 
the glaciation of British Columbia in a general theory of glaciation, 
owing to the complexity of its physical features and their marked 
character. Several conjectural schemes were advanced in a former 
communication ; but, abandoning the seemingly untenable theory 
of a great polar ice-cap, two probable hypotheses appear to remain. 
A general north-to-south movement of ice is indicated by striation 
in a number of places in the central-plateau zone, extending now 
for a length of over 400 miles. This region, from elevations ex- 
ceeding 5000 feet downward, is also covered thickly with drift-de- 
posits requiring, by their character and mode of arrangement, the 
action of water. ‘To account for these facts it was thought that 
either the flow of strong arctic currents bearing heavy ice during a 
period of great submergence might be supposed, or that the whole 
region may have been buried under a massive confluent glacier, the 
drift-deposits being laid down as it retreated in the water of the 
sea during a period of subsidence, or in that of a great lake held 
in by glacier-dams in the valleys of the several mountain-ranges. 

It was presumed that the gaps of the Peace and Pine rivers in the 
Rocky-Mountain range might have sufficed for the entrance from the 
north-east of such currents and masses of ice as would be required 
by the first theory; but the examination of the region, with this 
supposition in view, has convinced me that, notwithstanding the 
general decrease in elevation and width of the Rocky Mountains, 
the valleys of the rivers are too narrow and indirect, and the sur- 
rounding mountains too high, to allow the inflow of sufficient cur- 
rents with the degree of subsidence which would be required by 
most of the localities of glaciation and by the superficial deposits. 
Neither is there any evidence of the passage of drift-material in this 
region across the mountains either from east to west or in the oppo- 
site direction. 

It therefore appears to remain as the most probable hypothesis 
that a great glacier mass resembling the inland ice of Greenland 
has filled the region which may be called the Interior Plateau, be- 
tween the Coast Mountains and the Gold and Rocky Mountain 
ranges, moving (though perhaps very slowly) southward and south- 
eastward from the region of great precipitation and high mountains 
of the northern part of the province*, and discharging by the 
Okanagan depression and through the transverse valleys of the 
coast range. It still appears to me most probable, however, that 
this stage of the glacial period was closed by a general submergence, 
during which the deposit referred to as Boulder-clay was laid down 
in the interior plateau, and that as the land again rose it assumed 
its present terraced character. Conditions may bé suggested to 
account for the temporary existence of a great lake in the interior 


* Explorations in the northern part of the province in 1879 have shown 
that the mountains here are even higher and more extensive than had been 
supposed, several ranges exceeding 8000 feet in great portions of their extent. 


284 G. M. DAWSON ON THE SUPERFICIAL GEOLOGY OF 


plateau of British Columbia; but this will not explain the great; 
height to which water-action has extended on the east side of the 
Rocky Mountains*, which was probably synchronous. The last 
stage of the glacial period in the northern part of British Columbia 
appears to have given rise to the silts of the Lower Nechacco basin, 
while on the opposite side of the Rocky Mountains similar deposits 
were laid down over the Peace-River country, the elevation of the 
two districts being nearly alike. 

The general question of the origin of the drift-deposits of the 
Great Plains having been fully discussed elsewhere?, it will be 
unnecessary here to enter into it at length. The most remarkable 
feature of the glacial deposits of the plains is the Missouri Coteau, 
which it was supposed ran northward from the region near the 49th 
parallel, where it was more particularly studied, nearly following 
the margin of the third prairie steppe. This supposition has since 
been in great measure confirmed ; and on the journey from Edmonton 
to Winnipeg, in the autumn of 1879, I was able to examine cur- 
sorily the character of this feature where it touches the north 
Saskatchewan near the ‘“‘ Klbow,” and to observe the great accumu- 
lation of heavy boulders of eastern and northern origin in that 
vicinity. Further north, the facts now advanced show that with 
the general lowering of the surface of the country the well-defined 
zone of drift-deposits known as the Coteau is more or less completely 
lost, the material being scattered broadcast over the upper parts of 
the basins of the Peace and Athabasca rivers, and approaching in 
considerable mass the highlands near the base of the Rocky Moun- 
tains. 

Over the whole western portion of the plains, from the 49th to 
the 56th parallels, there is a mingling of the eastern and northern 
Laurentian débris with that from the Rocky Mountains to the 
west, the latter consisting largely of certain hard quartzite rocks, 
and the overlap seeming “to imply the existence of a sea in which 
ice derived from both sources floated freely. 


Discussion. 


The Presipent spoke of the care with which Dr. Dawson con- 
ducted his researches, and the value of his observations. 

Mr. Baverman stated that he was not acquainted with the district 
described by Dr. Dawson; but he thought, from what he had seen 
in Oregon and the Columbia valley, that many of the conclusions of 
Dr. Dawson could be established. He, however, doubted whether 
the ice had been quite so widely spread as Dr. Dawson supposed. 
He described some of the great terraces on the Barrier River ; there 
were sixteen, one over the other, on a stupendous scale. He had 
traced them on the Columbia River to 2300 feet above sea-level ; 


* Quart. Journ. Geol. Soc. vol. xxxi. p. 618. 
+ Quart. Journ. Geol. Soc. vol. xxxi. p.603. ‘Geology and Resources of the 
49th Parallel,’ p. 6. 


———— 


BRITISH COLUMBIA AND ADJACENT REGIONS. 285 


and they could be found still higher but for the degrading action of 
the climate. The rapid melting of the snow, followed by freezing, 
and slipping of the ice then formed, produced well-defined ice- 
scratches in a very short time. 

Prof. Boyp Dawxins said that he had studied the glacial phe- 
nomena in America, though he had not been so far north; and, 
so far as he could form an opinion, that northern area appeared to 
have been a great area of dispersal of ice. In the Western and. 
Pacific States, however, there was no evidence of a great ice-sheet, 
only a rather larger extension of local glaciers. On the eastern side 
the southern boundary of the confused glacial deposits, or the 
drift, passed from the latitude of New Brunswick in a N.W. direc- 
tion towards the area of the Mississippi, forming a low range of well- 
marked hills. To the south of this are the ‘‘ Champlain terraces” 
and traces of local glaciers on the higher hills. So that in North 
America there are two great systems of glaciation—one in the N.W., 
such as Dr. Dawson had described ; and another in the N.E. region, 
apparently pointing towards Greenland and Labrador. 


286 T. V. HOLMES ON THE PERMIAN, TRIASSIC, AND 


22. The Prrmtan, Triassic, and Liasstc Rocks of the Cartistx Basin. 
By T. V. Hotes, Esq., F.G.S8. (Read February 23, 1881.) 


[Puate XI. ] 


IT am enabled, by permission of Prof. A. C. Ramsay, Director-General 

of the Geological Survey, to lay before you the general results of 
the Survey examination of the, Permian, Triassic, and Liassic rocks 
in the country bordering the Solway Firth; but I do not propose 
to discuss here the glacial drift and other superficial beds by which 
the surface of the ground is almost entirely covered, and which are 
the main hindrance to an understanding of the rocks which form 
my subject this evening. 

Papers on the Permian and Triassic rocks of the North-west of 
England have been read before this Society by Prof. Sedgwick, by 
Sir Roderick Murchison and Prof. Harkness, and by Mr. E. W. 
Binney. But as few districts promise less, except as regards drift 
and peat-mosses, than that immediately around the Solway, it has 
hitherto been dealt with, as awhole, in a brief and cursory fashion. 
In addition, a knowledge of certain borings, the results of which are 
by no means generally known, is absolutely necessary to a correct 
view of the structure of the Carlisle basin (see Map, Pl. X1.). 

The lowest bed with which we are here concerned is the great 
brick-red Upper Permian sandstone, so well shown at St.-Bees 
Head, and named, therefore, the St.-Bees Sandstone. Between 
St.-Bees Head and Maryport the coast consists of the underlying 
Coal-measures, the St.-Bees Sandstone having been swept away by 
marine denudation. But at Maryport it again becomes visible, its 
most northerly appearance being below Swarthy Hill, on the fore- 
shore, about midway between Maryport and Allonby. North of 
Swarthy Hill none but superficial beds are exposed on the southern 
shore of the Solway. On the Scottish side St.-Bees Sandstone appears 
at Tordoff Point; but, with this exception, a walk along the coast 
between the Sark and the Nith shows no more than one between 
Allonby and Rockcliff, on the southern side. At a distance of one, 
two, or three miles from the coast-line, however, Scotland has de- 
cidedly the advantage, the corresponding part of Cumberland being 
entirely destitute of sections. 

Turning eastward from Maryport, the St.-Bees Sandstone is found 
to occupy a belt of country ranging towards Dalston on the Caldew — 
and Wetheral on the Eden. Between Maryport and the Caldew 
its southern boundary consists of Carboniferous rocks of various 
ages; and between the Caldew and the Eden of Lower Permian 
beds. The Carboniferous-Permian boundary is almost invariably a 
faulted one; in Shalk Beck, however, the unconformity between 
the two formations is very distinctly shown about a mile above 
Kast Curthwaite. The boundary between the St.-Bees Sandstone 
and the Lower Permian beds, between the Caldew and the Eden, 


Quart. Journ. Geol. Soc. Vol. XXXVIE Fi. X7. 


S 


BILE BES PREM 


SAGs RS 


ee. * 


Reference. 


| Sr i a 
| S =. Las = WE ee Shiites 
~ 
S hau i F 
| 3 SS Stance Marls = Feral S¢ Beas Sandst. 
S Ksridinton a 
& EST sandstone. l= wee 
S \ 
SP Ge borsterqns Rocks 
| re) Lil of varus Ages. 
—— Fazltis. 


The mark ———— 1 om the downthrow Side. 


0.4.8 Outlier of Kirklton Sandst. 


; Fea Daj afall La landon, 
SLE BASIN. 


i 
i Re ateaNA 
s 
4 
4 
+ 
any A 
Bynes 
5 a 
+ 
Con 
Fd 


\ 


Guart, Journ. Geol. Soc. Vol, XXXVIE Pl. XI 


0. Mi 
Gases 


Southerness 


MARYPOR 


tle 


Se 
oy 
7 
Sy 
/ 
Sloth 
ADUEV | 


4 


ye 
L 


ia 


i 


Ii 


G 


L hand? | A EN 
Hf). wale L 
b ifiT® Bangin gel : i 
[ee —— eS} 


Stanwie Marls 
Aarklinten 


| Sarudstone . 


—— Faults. 


BASIN. 


Reference. 


The mark —~— 1s on the downdrow Side. 
O.K.S=Outher of Kirklixton Sandst. 
| 


3 eA Preis Shales 
{EEE se Boas Sando 
Lasers Shales 

Perk Sindst. 


Jy] Garbonuterous Rocks| 
of runs Ages 


CARS. PE) 


Fred® Daniyerfotl bith, London. 


SHOWING THE ROCKS OF THE CARLISLE 


(Disregarding Superlfiaal Beds.) 
Seale __{__{ i _# Hfiles 


LIASSIC ROCKS OF THE CARLISLE BASIN. 287 


consists of two great faults, which abut against each other at New- 
biggin, on the Petterill. Both faults have a downthrow to the north. 
The more westerly ranges from a point alittle north of Rose Castle, 
on the Caldew, to Newbiggin ; the other from Newbiggin to Bracken- 
bank, on the Eden, about two miles above the railway bridge at 
Wetheral. This Brackenbank and Newbiggin fault is known to 
exist for many miles eastward of the Eden; it has also (as I believe) 
a considerable extension westward, in the direction of Cummers- | 
dale. 

The Lower Permian beds which are cut off by these faults are 
the Penrith Sandstone, a bright red rock remarkable for the scale 
on which it is false-bedded, and a red shaly series overlying it, and 
directly underlying the St.-Bees Sandstone. The Penrith Sandstone 
is well shown in the Petterill, a little below Wreay Bridge. The 
junction of the Penrith Sandstone with the overlying shaly beds (which 
in this locality contain gypsum) may be seen in the little streamlet 
which runs into the Petterill from the east, after crossing the Car- 
lisle and Penrith road about midway between Springfield and Carle- 
ton Hill. A little lower down the Petterill, but just above the road 
from Newbiggin Bridge eastward, there is a quarry in St.-Bees 
Sandstone. 

North of the faulted line the St.-Bees Sandstone is the only Per- 
mian rock seen*. About Brampton it occupies a broad belt of 
country between the Carboniferous formation on the east and the 
Triassic rocks on the west. In the Eden it is well shown between 
Brackenbank and Wetheral Bridge, and thence as far down as the 
junction of the Eden and Irthing. In the Hether Burn, the Line, 
and Cardwinley Burn sections are very abundant. In the Line the 
full thickness of the St.-Bees Sandstone does not appear, owing to 
Carboniferous rocks being brought in by a fault close to Brackenhill 
Tower, above which spot no Permian beds are seen; but in the 
Hether and Carwinley Burns there is an almost continuous series of 
sections, from Carboniferous rocks below the St.-Bees Sandstone to 
the Triassic beds above it. In both streams (as in Shalk Beck) a 
thin breccia is seen at the base of the Permian formation, and there 
is no fault at the junction with Carboniferous rocks. In the Esk, 
north of the railway bridge between Scots Dyke and Riddings junc- 
tion stations, the St.-Bees Sandstone may be seen, on the right bank 
of the stream, on both sides of the Border-line; it also appears 
in Moat Quarry and the cliff on the south side of the railway 
between Moat Quarry and the ancient fortress known as Liddel 
Strength. West of the Esk it is visible in the neighbourhood of 
Scots Dyke, both in the Glinger Burn and Sark ; alsoin many parts 
of the Kirtlewater, south of Kirtlebridge. It is well shown in the 
railway-cutting on the north side of the road from Annan to Kirtle- 
bridge, in the Annan water about Annan, and at Tordoff Point 
on the Solway. 

The dip of the St.-Bees Sandstone, which varies, as a general rule, 
from north to north-west between Maryport and the Caldew, slowly 


* Except a very thin breccia at its base. 


288 T. V. HOLMES ON THE PERMIAN, TRIASSIC, AND 


changes between the Caldew and Hether Burn, till, in the latter 
stream, it is about south-west. It retains this (general) south- 
westerly dip as far as the Esk, but west of it the dip becomes more 
or less east of south. At Annan it is nearly due south. 

No signs of the Lower Permian formations are visible east of the 


Annan Water. It is somewhat singular, however, that the Permian . 


rock between the Nith and Lochar Moss, in the neighbourhood of 
Dumfries, is precisely like the Penrith Sandstone in appearance. It 
is remarkable for the hardness and thickness of the beds of breccia 
it contains. The two great Permian sandstones of Penrith and St. 
Bees are so distinct in character, that there is usually no difficulty 
in distinguishing them even in hand specimens. 

Having traced the St.-Bees Sandstone around the circumference 
of the Carlisle basin, except where prevented from doing so by the 
waters of the Solway, the overlying beds now demand attention. 
First of these come the Gypseous Shales of Abbey Town and Bow- 
ness. So thickly and persistently covered by drift is the country 
immediately around those two places that were the only informa- 
tion obtainable that to be derived from natural sections the exis- 
tence of the Gypseous Shales would remain entirely unknown ; nor, 
indeed, would there be any reason to suppose the St.-Bees Sandstone 
to be overlain in that quarter by any thing but glacial drift, peat, 
and alluvium. 

Fortunately, however, two borings come to our aid, one of which 
discloses the fact that in the neighbourhood of Abbey Town more 
than 700 feet of Gypseous Shales rest upon St.-Bees Sandstone, and 
are themselves covered by nearly 200 feet of drift. The other 
boring was made near the west end of Bowness, at high-water mark, 
and proves the presence there of 367 feet of Gypseous Shales below 
41 feet of drift. It is not absolutely certain that the Bowness 
boring penetrated to the St.-Bees Sandstone, though it ended in red 
stone; but it is very highly probable that such was the case, St.- 
Bees stone being visible a short distance to the north, on the Scottish 
shore. At Bowness, therefore, the outcrop of the Gypseous Shales 
is beneath the Solway ; east of Bowness it probably ranges, under 
water, in the direction of Rockcliff Marsh. Towards Silloth, on the 
other hand, it is most likely outside the present coast-line, against 
which it may abut in the neighbourhood of Allonby. 

The southern outcrop of the Gypseous Shales can only be approxi- 
mately traced across an entirely drift-covered country by the help 
of the most northerly exposures of St.-Bees Sandstone. Its general 
direction is from Allonby to West Newton, and thence to Wigton 
and Dalston. 


East of Bowness and Abbey Town, the only evidence bearing on the 


existence of the Gypseous Shales is the record of an old boring near 
Great Orton, to which I shall have again to refer in treating of the 
Lias. It was made in search of coal in the year 1781. Below rock 
evidently Liassic was “ red stone or clay sometimes mixed with vems 
of white.” This description fits the Gypseous Shales very well, the 
gypsum in them being in the form of thin lamine, and beds of any 


= 


LIASSIC ROCKS OF THE CARLISLE BASIN. 289 


thickness being absent. On the other hand, in no other formation 
at all likely to be found underlying the Lias at Orton would “ veins” 
of a white substance be found. 

But if the Gypseous Shales are neither seen nor recorded as 
haying been bored through east of Great Orton, it may be asked 
why I have treated them as older than the formation which directly 
overlies the St.-Bees Sandstone north and east of Carlisle. On this 
point the evidence certainly leaves something to be desired, though 
there seems to me to be no doubt on which side the balance inclines. 
As will shortly be seen, the bed in question (which I have called 
the Kirklinton Sandstone) rests unconformably on that of St. Bees; 
while there is neither evidence nor presumption of any kind in favour 
of the existence of unconformity between the St.-Bees Sandstone 
and the Gypseous Shales. Again, the presence of gypsum is, so far 
as it goes, a presumption of Permian affinities, gypsum being found, 
as | have already mentioned, in the shales underlying the St. Bees 
stone, while it is not seen in any of the higher beds of this district. 

Leaving the almost sectionless district west of the Caldew and 
Eden till the time comes to treat of the Lias, it will now be most 
convenient to discuss the sections seen in the first-named stream 
about Dalston and Cummersdale. 

St.-Bees Sandstone is visible on the Caldew at Brackenhow and 
Buckabank, and here and there nearly as far down the stream as 
the outfall of the Pow Beck at Dalston. Opposite Dalston Hall 
the trap dyke, well known at Barrock Fell and Armathwaite, is 
slightly shown on the right bank, at the spot at which river and 
railway begin to run side by side. ‘This is the most westerly expo- 
sure of this dyke in Cumberland; but as I am informed by Mr. J. 
G. Goodchild that there is no ground for supposing (judging from 
his knowledge of it in the Eden-valley district) that it is also a fault 
of any magnitude, if a fault at all, I need say no more about it 
here. 

A few yards below the dyke, and on the same bank of the river, 
is a low cliff of greyish sandstone, and it seems to me that in this 
grey rock we probably have the uppermost beds of the St. Bees. The 
borizrg near Abbey Town shows that directly below the Gypseous 
Shales, and above the typical St.-Bees Sandstone, are about 40 feet 
of sandstone and sandy shale, mainly grey in colour. No sandstone 
like this is seen higher up the stream, and the rock next seen below 
at Cummersdale is evidently not St.-Bees Sandstone at all. In 
addition, Mr. J. G. Goodchild considers the beds at and below 
Buckabank to be higher beds in the St. Bees than any he has seen 
in the Eden-valley district, where the uppermost are cut off by the 
Penine fault. . 

The next section, that adjoining the rifle-butts at Cummersdale, 
disappoints any expectations that may have been formed of at last 
seeing the Gypseous Shales. Strange to say, they are nowhere visible 
at the surface. The rock at the rifle-butts is a very soft, red, shaly- 
looking sandstone, without any sign whatever of the presence of 
gypsum. Similar rock may be seen here and there in the river- 


290 T. V. HOLMES ON THE PERMIAN, TRIASSIC, AND 


bank, as far down as Cummersdale Print-Works. It is, in short, 
the Kirklinton Sandstone; but the dip at the rifle-butts is slightly 
west of north, and, being towards the Caldew, the surface of the 
low river-cliff is wet and slimy, and the characteristic bright colour 
and false bedding are both obscured. The soft Kirklinton stone, 
however, invariably weathers, as at Cummersdale, about, and for a 
few feet above, the surface of the river, along whose course it is 
exposed. A comparison of the sections at Cummersdale with those 
about Kirkandrews-on-Esk and elsewhere will show this very 
clearly. Fortunately the identity of the rock at Cummersdale is 
placed beyond dispute by the presence of a small but dry section 
about 100 yards below the rifle-butts. The spot is marked by the 
stump of an old tree, at the base of the bank bounding the alluvial 
flat on the right side of the stream. Here the bright and almost 
scarlet: colour and false bedding are both manifest. 

The question arises, in what way the absence of the Gypseous 
Shales at Cummersdale may be best accounted for. There can be 
no doubt that they suffered very much from denudation before the 
deposition of the Kirklinton Sandstone, as their total absence east of 
Carlisle bears witness. As regards the amount of their extension 
eastward, there are but two items of evidence. ‘The first is the old 
boring at Great Orton, already alluded to, in which 132 feet of what 
were probably Gypseous Shales were pierced and (apparently) their 
base not reached. ‘The second is a very recent boring at Justice 
Town or Iinehow, about a mile above the junction of Line and Hsk. 
This showed about 170 feet of Kirklinton stone resting directly on 
that of St. Bees. ‘'hese borings tend to show that the Gypseous 
Shales cease to exist more speedily north than south of Carlisle. On 
the whole, I am inclined to think that the Gypseous Shales would be 
visible at Cummersdale, but for an extension of the Brackenbank 
and Newbiggin fault. ‘This, if prolonged westward, might very well 
cross the Caldew between Dalston Hall and the rifle-butts, cutting 
off the Gypseous Shales and bringing in the Kirklinton Sandstone 
on its northern side; just as further eastward we find it bringing 
down the St.-Bees Sandstone on its northern side against the Lower 
Gypseous Shales on the south. 

The Kirklinton Sandstone is best seen in the parish from which 
it derives its name, and at Rockcliff and Netherby. I have already 
described its appearance at Cummersdale. Descending the Caldew 
it is again visible at Holmhead Bridge. Below Carlisle the upper 
beds are well shown at Skew Bank, north of Grinsdale, and lower 
ones at Rockcliff. Ascending the Eden from Carlisle it appears at 
Rickerby, in the river-bed at Low Crosby, and in the left bank a 
few yards N.E. of Holmgate. Thejunction with the St. Bees stone 
must be between the last-named spot and the junction of the Kden 
and Irthing. 

Between the Eden and Caldew the only evidence is the following. 
In the Petterill there is a small section, showing rock like that at 
Cummersdale, a few yards above the house called Petterill Bank ; 
and the record of a boring at Garlands Lunatic Asylum tells us that 


LIASSIC ROCKS OF THE CARLISLE BASIN. 291 


below 28 feet of drift 277 feet of red stone with white bands were 
pierced, which seemed to the borer to resemble “ Lazonby stone.” 
Now “ Lazonby stone” is, in other words, Penrith (the great Lower 
Permian) Sandstone ; but it is in the highest degree improbable that 
Penrith Sandstone would be found near the surface at Garlands, 
the natural presumption being that both St.-Bees and Kirklinton 
Sandstones overlie it there. A very simple explanation, however, 
suggests itself. The Penrith and Kirklinton Sandstones resemble 
each other very much in colour, and equally differ in that respect 
from the St. Bees. Both the Penrith and St.-Bees Sandstones are 
largely quarried—the former about Lazonby and Penrith, the latter 
near Curthwaite and Aspatria; but the soft Kirklinton stone is 
scarcely ever quarried, and is well shown only in localities that are 
but little generally known or visited. Hence the testimony of the 
borer, though decidedly against the supposition that the stone in the 
bore-hole was St. Bees, is not really against the view that it was 
Kirklinton Sandstone, as I have no doubt-was the case. Its outcrop 
hereabouts will keep a little east of Warwick and west of Scotby 
and Carleton, abutting against the Newbiggin and Cummersdale 
fault in the neighbourhood of Brisco Hall. 

The Kirklinton Sandstone is nowhere visible in the sectionless 
country between the Eden and Hether Burn; but in the latter 
stream it may be seen from Hether-Bank Bridge to its junction with 
the Line. At and above Hether-Bank Bridge are quarries in St.- 
Bees Sandstone, the dip being about south-west. At Cliff Bridge, 
Kirklinton, the Kirklinton Sandstone is extremely well displayed, 
and it may be seen, on ascending the Line, as far up as Shield Green, 
between Kirklinton Hall and the Muckle Linn. Between Shield 
Green and Brackenhill Tower is St.-Bees Sandstone, and above 
Brackenhill Tower Carboniferous beds (mainly sandstones and shales) 
are brought in by a fault which ranges nearly north and south, and 
may be seen crossing the river close to the Tower. 

Between Shield Green and Kirklinton Hall a bed appears in the 
Kirklinton Sandstone much resembling that of St. Bees; andin this 
harder band are two quarries, one in the northern corner of Hirst 
Wood, the other on the right bank of the river at Stag Ford. From 
the dip, these two quarries are in all probability in the same bed, 
and the St.-Bees-like stone need not be more than about 30 feet 
thick. 

There are no signs of faults, and the St.-Bees-like rock is evi- 
dently interbedded with Kirklinton Sandstone of ordinary appear- 
ance. This circumstance seems worth noting here, as, combined with 
the want of any evidence of unconformity between the two formations 
at Shield Green, it tends towards a totally different view from that 
pointed at by all the rest of the evidence bearing on the relations of 
these two beds. 

Below Cliff Bridge the Kirklinton Sandstone may be seen as far 
down as Metal Bridge, a little below the junction of the Esk and Line, 
being well shown at Westlinton. (I have already mentioned the 
boring at Justice Town, which showed 170 feet of Kirklinton Sand- 


292 T. V. HOLMES ON THE PERMIAN, TRIASSIC, AND 


stone above St. Bees.) Ascending the Esk it appears beneath Long- 
town Bridge and at various spots in the neighbourhood of Kirk- 
andrews Tower ; also about the outfall of Carwinley Burn, and up 
that stream as high as Carwinley Mill. At the mill the junction 
with the St. Bees is well shown. Itseems to me that there is some 
slight evidence of unconformity between them at this point, West 
of the Esk, Kirklinton Sandstone is seen in the lower part of the 
course of the Glinger Burn, and St. Bees nearer Scots Dyke. AIL 
that can be said of its western outcrop is that it must keep a little 
westward of the various exposures in the Glinger Burn and at Metal 
Bridge and Rockcliff, and eastward of the Gypseous Shales of Bow- 
ness. It probably runs below the Lias in the neighbourhood of 
ark Bampton. 

It will be remembered that the Kirklinton Sandstone seen at 
Cummersdale was invariably red, lke that of Kirklinton or 
Netherby ; but at Holmhead, in higher beds, a borehole recorded by 
Mr. EK. W. Binney* pierced through 108 feet of white sandstone, 
overlying 117 feet (not through) of red. At Carlisle Gaol the bore- 
hole was through 250 feet of red sandstone ; while at Messrs. Dixon 
and Co.’s, West Tower Street, I was informed by Mr. John Hamil- 
ton that 123 feet of white sandstone were penetrated. Other ex- 
amples might be given. Lastly, in a boring at Stainton, in still 
higher beds, 360 feet of white or grey stone were proved above an 
unknown thickness of red. Thus the Kirklinton Sandstone appears 
to consist of a lower red, a middle red and white, and an upper 
white series of beds. 

The Stanwix Marls probably nowhere attain any considerable 
thickness. Their most characteristic colours are red and greenish 
grey. They vary very much in hardness, stony bands being much 
more common in some places than others. They lie between Cliff 
Bridge on the north and Carlisle on the south, Houghton on the 
east and Beaumont on the west. They are well shown on the Eden 
in the lower part of the cliff at Etterby Scaur, from the North 
British railway-bridge to Grinsdale, and about Beaumont, the sec- 
tions in the two last-named localities being in the left bank of the 
river. On the Line their junction with the underlying Kirklinton 
Sandstone may be very plainly seen, both at Westlinton and near 
Cliff Bridge. At Westlinton it appears in the bank bounding the 
alluvial flat, a few yards east of the bridge. Nearer Cliff Bridge 16 
is plainly shown in the little plantation bordering the alluvium, 
about midway between Cliff Bridge and High Alstonby. The marls 
appear here and there in the little streams that unite and fall into 
the Line between Low Alstonby and Westlinton ; they are visible 
nearly as far eastward as Stonystonerigg. 

Before taking leave of the neighbourhood of Kirklinton, which is, 
as we have seen, perhaps supreme in geological interest among the 
localities mentioned in this paper, I may remark that the scenery of 
the Line above Cliff Bridge is very picturesque for many miles and 
very different from that below it. Both scenery and geology, how- 

* Mem. Lit. & Phil. Soc. Manchester, ser. 3, vol. ii. pp. 843-388. 


LIASSIC ROCKS OF THE CARLISLE BASIN. 293 


ever, have hitherto been entirely ignored by writers on the geology 
of North Cumberland, as well as by the authors of guide-books; 
for, though Cliff Bridge itself is only about two and a half miles 
from Longtown or Lyneside stations, trains stopping at those places 
are few and far between, and every mile higher up the Line increases 
the distance from the railway by that amount; while eastward lies 
the lone bare district of Bewcastle, destitute alike of railways and 
inns. 

The extent of the area covered by the marls, south of the Eden, 
is doubtful, but the evidence available tends to show that it is very 
small. Though they form the lower part of the cliff at Etterby 
Scaur, the three new railway-bridges across the Caldew, the lowest 
of which is close to the junction of the North-British and Caledonian 
Railways, are all founded on the underlying sandstone, the marls 
not having been met with at all; and they were absent in the wells 
at Messrs. Carr’s, Caldewgate, and Messrs. Dixon’s, West Tower 
Street : also in that at the Gaol. On the other hand, I saw them 
in an excavation at the foot of Gaol Brow, on the north side of the 
Gaol, and have observed traces of them at the bottom of deep drains 
in Bank Street, Lowther Street, and opposite Cavendish Place in 
the Warwick Road; but their thickness hereabouts, where they 
exist, must be very trifling. 

West of the Caldew the evidence is much scantier. In addition 
to the well at Messrs. Carr’s, already mentioned, Mr. KE. W. Binney 
records (in the paper before quoted) that at the pumping-engine for 
the canal by Edenside, immediately above the red and variegated 
marls, there was a section in the pump-well which distinctly showed 
the marls at the top gradually passing down into the red sandstone 
below. Again, at Stainton, on the north side of the Eden, between 
Carlisle and Grinsdale, a boring showed them to be only 23 feet 
thick. In the Eden, south of Stainton, the beds are lying nearly 
flat, except near the North-British railway-bridge, where north- 
easterly and north-westerly dips occur. There is no evidence as to 
their thickness near Beaumont. 

The above facts, when combined with the great change in direc- 


tion of the outcrop of the marls between Rickerby and Caldewgate, 


from about N.N.K. and 8.8.W. at the former, to nearly east and 
west at the latter place, seem to point to their very slight extension 
below the Lias. Most of the Lias probably rests, therefore, on the 
Gypseous Shales west of Great Orton, and on the Kirklinton Sand- 
stone east of that village, the Stanwix Marls underlying the Lias 
only in the neighbourhood of Bellevue. 

Fresh borings for coal in the Lias district not being probable, the 
existence of Rheetic beds is likely to remain an open question. No 
evidence of them has yet been discovered, all fossils hitherto found 
having been determined by Mr. Etheridge (our President) to be 
Lower Lias ; but so drift- covered is the country, and so few and small 
are the sections, that negative evidence must go for very little in 
settling the question. The Lias country is purely agricultural, and 
wells sunk for the supply of farm and other houses are usually only 


Q. J.G. 8S. No. 146. x 


294 T. V. HOLMES ON THE PERMIAN, TRIASSIC, AND 


from 15 to 20 feet deep, or perhaps half the average thickness of the 
Glacial*drift. Indeed a well-sinker, living near the southern border 
of the Lias*, informed me, mistaking the motive of my inquiry, 
that I need not be afraid of meeting with any rock below the drift 
in the locality around his home, as he had sunk at various places he 
mentioned to depths of 20, 30, or 40 feet without penetrating any 
thing but sand, gravel, or clay ; for instances in which water, 
attained on reaching a clayey stratum of drift, has been lost on 
touching porous sandstone below, have made well-sinkers very care- 
ful not to go a single foot lower than is absolutely necessary in the 
case of ordinary dwellings. 

The Lias area has, during the last 250 years, been much explored 
in search of evidence of coal. It will not seem strange that such 
has been the case, when it is remembered that the dark shale form- 
ing so large a proportion of the rock visible would naturally seem 
identical with the dark shale of the Coal-measures, and a striking 
contrast to the red Permian and Triassic beds around. The non- 
Carboniferous, and probably Liassic, nature of the formation was 
first discovered by Mr. R. B. Brockbanky, who, on finding Ammo- 
nites and other fossils in Thornby Brook, sent them to Mr. E. W. 
Binney, who pronounced them to be Liassic. Mr. Binney after- 
wards visited the district and recorded the result in a paper read 
before this Societyt. JI have since discovered but one section not 
described therein, in a brook between Great Orton and Flat. It is 
composed of dark shale with limestone bands. Ammonites John- 
stont was found there, as also in Thornby Brook and at Quarry Gill, 
near Aikton. By Mr. R. B. Brockbank’s assistance I was enabled 
to obtain evidence of the thickness attained by the lias between 
Great Orton and Flat, a spot nearly at its centre. A document, from 
which J have already quoted, which has been preserved by the Stordy 
family of Great Orton, gives the following details, according to a 
copy of it kindly made for me by Mrs. Hannah Pearson, of Station 
Hill, Wigton. In the year 1781 a boring was made by John Brisco, 
of Crofton, in John Stordy’s Gill close. A blue stone was found 
18 feet from the surface, and ‘different stone, mostly bluish,” till 
they arrived at a depth of 228 feet. Then they pierced the “red 
stone or clay, sometimes mixed with veins of white” (which I sup- 
pose to be the Gypseous Shales), till they came to a depth of 
360 feet. 

The Lias area forms a plateau, with a slightly greater general 
elevation than is attained immediately outside it. This plateau-like 
character is better marked in the country between Aikton and Great — 
Orton than eastward of the latter place. No person standing about 
Wiggonby, near the southern border of the Lias, can fail to notice the 
difference between the flat-topped plateau northward and the rolling 
drift-ridges to the south. But though this change of feature makes 
it possible to map the Lias boundaries with some approach to a 


_* Andrew Miller, Nealhouse. 
+ Of Moor Park, Crosby, near Maryport. 
+ Quart. Journ. Geol. Soc. vol. xv. p. 549. 


LIASSIC ROCKS OF THE CARLISLE BASIN. 295 


fairly good general line, the persistently drift-covered surface 
prevents any ‘thing like precision. 

In conclusion, ‘T will briefly pass in review the leading points 
bearing on the relations of the beds described to each other, which 
are illustrated in the diagram sections (figs. 1-3, p. 296). 

The St.-Bees Sandstone has been shown to surround the overlying 
beds, its general dip varying from north to a few degrees north of 

west, between Maryport and the Eden. In the Hether Burn, Line, 

and Carwinley Burn its general dip is more or less south of west ; 
and west of the Esk it varies between south-east and south. There 

can be no doubt that the St.-Bees Sandstone is the lowest bed of a 

true basin, the western limits of which are now below the Solway. 

Then, first of the overlying beds come the Gypseous Shales, which, 
strange to say, are nowhere exposed to view, but are known on the 
evidence of the boreholes at Abbey Town and Bowness, and, perhaps 
I may add, that at Great Orton. Their invisibility is mainly caused 
by the special thickness and persistence of the drift over the ground 
they occupy, but also, at Cummersdale, to the probable interposi- 

tion of a great fault. There is no evidence of any kind suggesting 
unconformity between the Gypseous Shales and the St.-Bees Sand- 
stone; and, on the other hand, as Gypseous Shales underlie the 
St.-Bees Sandstone near Carlisle, gypsum is evidently hereabouts a 
Permian characteristic. I have accordingly classed the Gypseous 
Shales as Permian. 

The Kirklinton Sandstone, however, appears to rest unconform- 
ably on the Gypseous Shales to the west, and on St.-Bees Sandstone 
to the east and north-east of Carlisle. But not only do the Gypseous 
Shales disappear towards the north-east, but the greater part of the 
St.-Bees Sandstone is also missing. Six or seven miles south-east 
of Carlisle the thickness of the St.-Bees Sandstone is estimated by 
Mr. J. G. Goodchild at from 1500 to 2000 feet; but in the Hether 
Burn there can hardly be more than 800 feet of Permian rock, from 
the breccia at its base to the outcrop of the Kirklinton stone below 
Hether-Bank Bridge ; and in Carwinley Burn, between the breccia 
at the base of the Permian formation and the outcrop of Kirklinton 
stone at Carwinley Mill, there cannot be more than 250 feet of rock. 
In both these burns there is an almost continuous series of sections, 
and there are no signs of faults. Both about Brampton on the one 
hand and west of the Esk on the other the thickness of the St.- Bees 
Sandstone must be very much greater. Two outliers of Kirklinton 
Sandstone exist—one at Canobie, opposite the church, resting on 
Carboniferous rock; the other on the Cambeck near Walton, resting 
on St.-Bees Sandstone. ‘Thus the band of St. Bees-like rock seen in 
_ the Kirklinton Sandstone on the Line goes for nothing when the 
whole of the evidence is considered, important as it would be did it 
stand alone. In poupcamence of this decided unconformity to the 
beds below, I have classed the Kirklinton Sandstone as Bunter. 

-  Itis evident that the Stanwix Marls, in their turn, repose uncon- 
_ formably on the Kirklinton sandstone ; for while at Cliff Bridge and 
_ Westlinton they rest upon the lower red beds, at Stainton and 

Kee 


T. V. HOLMES ON THE PERMIAN, TRIASSIC, AND 


296 


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LIASSIC ROCKS OF THE CARLISLE BASIN, 297 


Grinsdale they le upon the upper white ones. In consequence of 
this decided unconformity the Stanwix Marls are here classed as 
Keuper. 

And there seems to me to be little doubt that the Lias, in its turn, 
rests unconformably on formations all of which are unconformable 
to each other—the Gypseous Shales, the Kirklinton Sandstone, and 
the Stanwix Marls. But as in this case it is barely possible, though 
not, I think, probable, that the Stanwix Marls may overlap the two 
lower formations and underlie the Lias throughout its area, the 
possibility seems worth mentioning. 

In this paper such facts only have been brought forward as 
seemed necessary to establish the true relations of the various for- 
mations to each other, fuller details being reserved for a forthcoming 
memoir. 


Nore on the Lias. April 25, 1881. 


Since writing my paper, my attention was called by my friend 
Mr. H. B. Woodward to a passage in a paper by Rev. J. E. Cross 
*‘On the Geology of North-west Lincolnshire ” (Quart. Journ. Geol. 
Soc. vol. xxxi. p. 116). Mr. Cross remarks that in the neighbourhood 
of Appleby he has not been able to detect any Rheetic fossils, though 
he has searched diligently for them. He says, “ the first strata 
next above the Keuper are those which contain Ammonites angulatus 
and A. Johnston.” He also remarks that A. angulatus has the 
ereater range, and extends throughout a zone of 150 feet or 
more. 

In Cumberland A. Johnstont must have a range fully equal to that 
of A. angulatus in North-west Lincolnshire. 

In a letter received from Mr. Cross, dated February 22nd, 1881, 
he states that he has not since detected any Rheetic fossils in the 
above-mentioned locality, but that, as they may be seen both near 
Gainsborough on the south and across the Humber on the north, it 
is very possible that they nevertheless exist there. He also remarks 
that as “‘A. angulatus is one of the most persistent of Ammonites in 
its own place, and A. Johnstoni, whenever it appears, ranges from 
the middle of the A.-angulatus-beds to strata below those in which 
A. angulatus is ever seen, I think if you find A. Johnstoni and do 
not find A. angulatus, you must be below the A.-angulatus-beds 
altogether.” 

Prof. J. W. Judd, in his paper on ‘“‘ The Secondary Rocks of Scot- 
land” (Quart. Journ. Geol. Soc. vol. xxxiv. p. 697); remarks that the 
zone of Avicula contorta does not appear to be distinctly developed 
in the West Highlands, while the Infralias (Planorbis- and Angulatus- 
zones) attains a thickness of from 150 to 200 feet. 


EXPLANATION OF PLATE XI. 
Sketch Map showing the rocks of the Carlisle basin: scale 4 miles to | inch. 


298 ON THE PERMIAN ETC. ROCKS OF THE CARLISLE BASIN. 


Discussron. 


Mr. Baverman, having experienced the extreme difficulty of 
understanding the rocks of this district from the obscurity of the 
evidence, bore testimony to the great care and skill with which 
the author had worked out such an unpromising subject. 

Prof. Jupp remarked upon the great interest attachiug to the 
outlying patch of Lias near Carlisle. The author had now, for the 
first time, made us acquainted with the thickness of these Lias 
strata, and proved that it exceeded 200 feet. So far as the evidence 
went, they belong entirely to the Infralias, and the same member 
of the Lias was also abnormally thick on the west coast of Scotland. 
The numerous unconformities pointed out by the author as existing 
among the red rocks below seemed to show that but little value 
should be attached to such local unconformities in classifying the 
strata. 

Mr. Wartaker said that his colleague Mr. H. B. Woodward pro- 
posed to group the whole of these red rocks under the comprehensive 
term Poikilitic, and thought that the Rheetic beds are most likely 
present in the area, although not seen. He spoke of the unsafe 
character of the colour-test in classifying rocks. 

Rev. J. F. Buaxs asked as to the evidence of the existence of as 
much as 200 feet of Lias beds belonging entirely to the lowest part 
of the series, and as to the proofs of the existence of an uncon- 
formity between the Stanwix Marls and the Lias. 

Prof. T. Rurrrr Jones protested, with reference to the use of the 
word ‘ Poikilitic,” used by preceding speakers indicative of anti- 
Permian views, that there had been good grounds for John Phillips 
and others to determine and establish the Permian Series, and that 
there were no good grounds for its being again amalgamated with 
the overlying series. 

The PRESIDENT said that no evidence of the occurrence of Rheetic 
fossils in the area had ever been found, but that possibly the 
Stanwix Marls might have to be classed with the Rhetie Tea- 
green Marls of the Bristol area. The Lias of Adderley was let down 
by means of great faults below the ievel of the Keuper marls. 

The Avrnor said that he advanced his classification as a purely 
provisional one. He did not think it probable, though it is barely 
possible, that the Stanwix Marls underlie the Lias. In classing 
certain of these beds as Permian, Bunter, or Keuper, the author 
chiefly desired to mark their relative positions and affinities, the 
latter being determined by the existence or non-existence of un- 
conformities between them. 


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DEVONIAN PLANTS. 


Quart. Journ. Geol .5oc Vol OOWiiee eae 


A.S.Foord del. Mintern Bros. mp. 


DEVONIAN PLANTS. 


ON NEW ERIAN (DEVONIAN) PLANTS. 299 


23. Nores on New Ertan (Devontan) Prants. By J. W. Dawsoy, 
LL.D., F.R.S., F.G.S., Principal of M°Gill College, Montreal. 
(Read June 23, 1880.) 

[Prares XII. & XIIL] 


THE principal purpose of the present paper is to add a few new facts 
to our knowledge of the Precarboniferous flora of Eastern America. 
Since the publication, in the Journal of this Society, of my paper, 
“Further Observations on the Devonian Plants of Maine, Gaspé, 
and New York” (1863), and that on “Fossil Ferns of the Devo- 
nian ” (1871), a large addition was made to the knowledge both of 
the species of plants and of the general character and conditions of 
Devonian and Upper Silurian vegetation in my “Report on the 
Devonian and Upper Silurian Plants of Canada”*. In a subsequent 
report on the “ Plants of the Lower Carboniferous and Millstone 
Grit”7, I endeavoured, by the aid of the American formations, to 
remove the perplexities that had been caused by the disputes re- 
specting the age of the Kiltorcan beds in Ireland and the so-called 
“‘Ursa-stage ” of Bear Island, difficulties which, however, still appear 
to constitute subjects of discussion. 

In 1878 I contributed a paper to the Geological Society of Edin- 
burgh, entitled ‘‘ Notes on Scottish Devonian Plants,” comparing 
these plants with those of America, and at the same time establish- 
ing the genus Péilophytont for those remarkable pinnately-leaved 
Lycopodiaceous plants of which Lycopodites Vanuxemu of the De- 
vonian of New York and LZ. plumula of the Lower Carboniferous of 
Nova Scotia are types, and of which Pt. Thomsoni is a Scottish re- 
presentative described in the paper above referred to. 

Since the publication of these reports and papers several interest- 
ing new forms have come into my hands from the Devonian or Erian 
of North America, to which I shall add two new species, one from 
Scotland and one from Australia, kindly communicated to me by 
friends in those countries. 


Plants from the Erian (Devonian) of New York. 
1. AsTHROPTERIS NOVEBORACENSIS, gen. & sp.n. (Pl. XII. figs. 1-9.) 


The genus Asteropteris is established for stems of Ferns having the 
axial portion composed of vertical radiating plates of scalariform 
tissue imbedded in parenchyma, and having the outer cylinder com- 
posed of elongated cells traversed by leaf-bundles of the type of 
those of Zygopteris. 


* Geol. Survey of Canada, 1871, pp. 100, 20 plates. 

+ Ibid. 1873, pp. 47, 10 plates. 

{ Ptilophyton may perhaps he considered too near to Psilophyton, but the 
sound of the names is quite different. In a recent Report of the Regents of 
New York, Hall has stated his belief that Pt. Vanuxemii is not a plant but a 
zoophyte. He does not fully state the reasons for this conclusion ; but should 
this view be established with regard to this species, then Géppert’s Lycopodites 
penmneformis or my L. plumula may serve as the type of the genus. 


300 DR. J. W. DAWSON ON NEW 


The only species known to me is represented by a stem 2°5 cen- 
timetres in diameter, slightly wrinkled and pitted externally, per- 
haps by traces of aerial roots which have perished. The transverse 
section shows in the centre four vertical plates of scalariform or 
imperfectly reticulated tissue, placed at right angles to each other, 
and united in the middle of the stem (figs. 1-4). At a short distance 
from the centre, each of these plates divides into two or three, so as to 
form an axis of from ten to twelve radiating plates, with remains of 
cellular tissue filling the angular interspaces (fig. 3,6). The greatest 
diameter of this axis is about 1°5 centimetre. Exterior to the axis 
the stem consists of elongated cells (fig. 7), with somewhat thick 
walls, and more dense towards the circumference. The walls of these 
cells present a curious reticulated appearance, apparently caused by 
the cracking of the ligneous lining in consequence of contraction in 
the process of carbonization. Imbedded in this outer cylinder are 
about twelve vascular bundles (figs. 2, 3,d@), each with a dumb-bell- 
shaped bundle of scalariform vessels enclosed in a sheath of thick- 
walled fibres. Hach bundle is opposite to one of the rays of the 
central axis. The specimen shows about two inches of the length of 
the stem, and is somewhat bent, apparently by pressure, at one end. 

This stem is evidently that of a small tree-fern of a type, so far 
as known to me, not heretofore described *, and constituting a very 
complex and symmetrical form of the group of Paleozoic Ferns 
allied to the genus Zygopteris of Schimper. The central axis 
alone has a curious resemblance to the peculiar stem described by 
Unger (‘ Devonian Flora of Thuringia’) under the name of Cladoaylon 
nurabile; and it is just possible that this latter stem may be the axis 
of some allied plant. The large aerial roots of some modern tree- 
ferns of the genus Angiopteris have, however, an analogous radiating 
structure. 

The specimen is from the collection of Berlin H. Wright, Esq., 
of Penn Yan, New York, and was found in the Portage group 
(Upper Erian) of Milo, New York, where it was associated with 
large petioles of ferns and trunks of Lepidodendra, probably L. 
chemungense and L. primavum. 

In previous communications to the Society I have described three 
species of tree-ferns from the Upper and Lower Devonian of New 
York and Ohio; and this species is from an intermediate horizon. 
All four occur in marine beds, and were, no doubt, drift-trunks from 
the fern-clad islands of the Devonian sea. The occurrence of these 
stems in marine beds has recently been illustrated by the observa- 
tion of Prof. A. Agassiz, that considerable quantities of vegetable 
matter can be dredged from great depths in the sea on the leeward 
side of the Caribbean Islands. The occurrence of these trunks 
further connects itself with the great abundance of large petioles 
(Rhachiopteris) in the same beds, while the rarity of well-preserved 
fronds is explained by the coarseness of the beds and also by the 
probably long maceration of the plant-remains in the sea-water. 


* Prof. Williamson, to whom IJ have sent a tracing of the structure, agrees 
with me that it is new. 


ERIAN (DEVONIAN) PLANTS. 301 


2. Eauisutipes WrigHTIANA,sp.n. (Pl. XII. fig. 10& Pl. XIII. fig. 20.) 


This is a specimen in the same collection with the above. It is 
a cast in sandstone, 6 centimetres in diameter, with nodes from 4 to 5 
centimetres apart. The surface has a slight carbonaceous coating 
and is finely tuberculated, the tubercles being very regularly 
arranged, and representing the bases of very short hairs or bristles, 
which are seen entering the surrounding matrix. Impressions 
above the joints appear to indicate sheaths, each of about twelve 
broad leaves, which are abruptly narrowed and acuminate at the top, 
and show an indication of a median nerve or rib (fig. 10). The leaves 
of the sheaths are 1 centimetre broad and 1°7 centimetre long. It 
would be possible, however, to interpret these supposed sheaths as 
due to mere plications or foldings of the epidermis; and in this case 
the plant may have borne verticils of leaves, of which these sup- 
posed sheaths may be merely the remains. ‘The first explanation, 
however, appears more probable; and, if it is correct, the plant is a 
- true Hgwisetides, and the present specimen is the first occurrence of 
this genus in beds older than the Carboniferous. It is to be observed, 
however, that ‘Unger has described from the Cypridina-slates of 
Thuringia plants of the genera Kalymma and Asterophyllites (A. 
coronata) with sheaths at the nodes; and my A. scutigera, from St. 
John’s, has verticils of scales at the joints, which may represent 
sheaths. The present species has a remarkable resemblance in its 
markings and the form of its sheaths to a greatly magnified stem of 
the modern Hquisetum fluviatile, except that the leaves of the sheaths 
are shorter. 

The species is named in honour of its discoverer. Its essential 
characters will be as follows :— 3 

Stem stout, cylindrical or broadly ribbed, surface marked with 
short hairs or tubercles regularly arranged. Sheaths at the joints, 
of about twelve leaves, of the general form of those of Hquisetuimn 
fiiatile. 

The specimen is from the Portage group (Upper Erian) of Italy, 
New York. 


3. CYCLOSTIGMA AFFINE, sp.n. (Pl. XII. figs. 11 & 12.) 


Stem marked with alternate circular leaf-bases or areoles, slightly 
prominent below, evanescent above, and each with a circular dot or 
vascular mark. Scars scarcely two millimetres in diameter, and 
separated by finely corrugated bark, about twice their diameter 
apart. These markings occur on a stem about an inch in diameter. 
The Knorria, or decorticated form of this plant, presents irregular 
waving ridges, produced by the longitudinal confluence of the oblique 
vascular bundles. 

This plant is the nearest approach to the well-known C. kiltorkense 
of Ireland hitherto found in America. It differs chiefly in the more 
closely placed areoles. It was collected by Mr. Wright, and is from 
the Chemung (Upper Erian) of Italy, New York. The study of this 
plant has led me to the belief that Stigmaria ewiqgua of my Report 
of 1871 may, when better known, prove to be a new species, allied 
to the present, and a member of the genus Cyclostigma. 


302 DR. J. W. DAWSON ON NEW 


4, LEPIDODENDRON PRIMzVUM, Rogers. (PI. XII. fig. 13.) 


Mr. Wright’s collection contains fragments of a Leprdodendron 
from Milo, New York, which seems to belong to the species above 
named, but presents the curious peculiarity of having the leaf-bases 
depressed instead of being prominent. This may result either from 
some peculiarity of pressure or from the leaf-bases being deciduous 
and leaving depressed scars when removed. In either case these 
specimens illustrate this peculiarity as seen in the Lower Silurian 
Glyptodendron of Claypole, which may possibly have had decorti- 
cated leaf-bases. Specimens of this kind, of course, retain no distinct 
vascular marks, and the impression on the matrix resembles those 
decorticated Lepidodendra of the Coal-formation which used to be 
named Lyginodendron, but which, in Nova Scotia at least, usually 
belong to the species Lepidodendron rimosum, 


5. CELLULOXYLON PRIMMVUM, gen. & sp. n. 


A silicified trunk, showing in cross section large and somewhat 
unequal hexagonal cells, with an appearance of Knes of growth 
caused by concentric bands of smaller cells. No medullary rays. 
The longitudinal section shows either cells superimposed in vertical 
rows, or a sort of banded prosenchymatous tissue; but the structure 
is much masked by the crystallization of the quartz. 

This specimen is from the collection of Prof. J. M. Clarke, of Am- 
herst, Massachusetts, and was obtained from the Hamilton (Middle 
Erian) of Canandargua, New York. It was undoubtedly a woody 
stem and not an Alga; but its structure is even less specialized than 
that of Prototaxites, from which it differs in the want of medullary 
rays, and in its less distinctly elongated wood-cells without spiral 
markings. It has some resemblance to Aphyllum paradoaum of 
Unger, but is more uniform in its structure. It adds another to 
those mysterious woody stems of doubtful affinities which, in the 
Devonian or Erian of both sides of the Atlantic, represent the 
Taxinez and Conifers of later formations. 

Additional specimens received from Prof. Clarke show that the 
appearance of rings of growth is caused by large cells disposed in 
concentric narrow bands between the wider bands of fine fibrous 
tissue. In the longitudinal section these bands of large cells appear 
to be parenchymatous and not vascular. There are no medullary 
rays, but rounded patches of cellular tissue appear here and there 
in the fibrous layers. The structure is thus very peculiar, and ap- 
pears to have been the result of a kind of exogenous growth, in which 
coarse parenchymatous layers were deposited between the periodical 
rings of the stem, reminding one of the bark-like layers interposed 
between the growth-rings in G'netwm and in some tropical climbers. 
The stem of the present plant was, however, in all probability, of 
much more simple character, though woody and capable of resisting 
pressure. It is to be observed also that the specimens neither show 
the structure of the pith or bark, and that the finer structures of 
the tissues preserved must have been partially obliterated by the 


ERIAN (DEVONIAN) PLANTS. 303 


eranular crystallization of the quartz with which the specimen is 
mineralized. 


Miscellaneous Specimens from New York. 


Numerous large petioles of Ferns occur in collections sent to me 
by Mr. Wright and Dr. Parker of Ithaca, New York. Being desti- 
tute of the fronds, it seems unnecessary to describe them more 
particularly ; but they indicate the possibility that the Erian of New 
York may yet afford a rich Fern-flora comparable with that of St. 
John, New Brunswick. In collections made by Mr. Wright are 
also specimens of those singular plants, supposed to be Alge, 
which Hall has named Dictyophyton. A very fine specimen of one 
of the species was figured in my paper of 1863, under the name 
Uphantena chemungensis, originally bestowed on one of the species 
by Vanuxem, but which is rejected by Hall in favour of the generic 
name above given. The specimens sent by Mr. Wright do not give 
any additional information as to the mode of growth of these curious 
forms; but he has found in the Hamilton formation, not previously 
known to contain these plants, a species probably distinct from 
those described by Hall, and which may be named D. hamiltonense ; 
though if these plants were really Alge, the supposed species 
may be nothing more than varietal forms or stages of growth. 

The specimens referred to are unequally turbinate or unequally 
conical in form, rapidly expanding from the base, and marked with 
sharp longitudinal ridges, crossed with much finer and more 
frequent revolving lines. The largest specimen is almost 13 inch in 
diameter, narrowing to less than 1 inch in a length of less than 
2 inches. 

The remarkable spiral plant belonging to the genus Spirophyton 
of Hall, the “* Cauda-galli fucoid” of the earlier Reports on the State 
of New York, is found in the same beds with Dictyophyton. It is 
also found, as mentioned in my paper of 1863, in Gaspé, where it 
ranges from the Upper Silurian into the Lower Devonian. Plants 
of the same genus have been found by the late Prof. Hartt on the 
Rio Tapajos, in Brazil, in beds referred to the Carboniferous period, 
though some other plants found in the same beds might in North 
America be supposed to be Upper Devonian in age. In MS. descrip- 
tions of these plants sent to Prof. Hartt, and which may have been 
published in his Reports, I named this species S. brasiliense. It is 
of interest as showing the very wide distribution of this form in the 
paleeozoic seas. 


Plants from the Erian (Devonian) of St. John, New Brunswick. 


I have recently obtained, from the widow of the late lamented 
Prof. Hartt, the remainder of his Devonian plant-collections, con- 
sisting principally of duplicates of the more common species found 
at St. John, but with a few fragments indicating forms not pre- 
yiously known to me. 

Since the publication of my papers and reports on the fossils of 
the St.-John beds, they have been repeatedly referred to by Kuropean 


304 DR. J. W. DAWSON ON NEW 


palzobotanists as Lower Carboniferous, apparently on no better 
grounds than their superior richness in plants to the Devonian of 
Europe. On this account it may be desirable here to summarize 
the evidence now available as to their actual age. This may be 
stated thus:—(1) The Dadoxylon Sandstone and Cordaite shale 
of Southern New Brunswick are folded up and partially altered with 
the Silurian and Cambrian rocks of the district, and are overlain 
unconformably by the Lower Carboniferous conglomerates (Subcar- 
boniferous of some American geologists). These conglomerates are, 
further eastward, associated with beds holding the characteristic 
fossil plants of my Horton series, equivalent to the Tuedian or 
Calciferous series of Scotland. There is also evidence that the 
Devonian plant-beds are anterior to the great intrusive Devonian 
granite of this region, whose débris are found in the Lower Carbo- 
niferous conglomerates, but not in the underlying rocks. ' Additional 
facts illustrative of these points will be found in the Reports of 
Messrs. Bailey and Matthew in the publications of the Geological 
Survey of Canada for 1871 and 1875. 

(2) The flora of these beds is markedly different from that of the 
Lower Carboniferous, of the Millstone Grit, and of the true Coal- 
formation in New Brunswick and Nova Scotia, all of which have 
been studied and described. 

(3) The prevalent forms in the St.-John beds are those character- 
istic of the Devonian in Gaspé, New York, and Maine, such as Arche- 
opteris, Cyclopteris obtusa, Psilophyton, Calamites radiatus, Dadoxylon 
ouangondianwm, though several genera are common to these beds and 
the Carboniferous. The fact that the flora of these beds is richer 
than that of the Huropean Devonian, and contains types which 
appear later in Kurope, is in harmony with known facts as to the 
earlier appearance of plants in America in other stages of geological 
history. JI may add that some of the genera noticed in 1863 from 
St. John, and not then known in the Devonian of Europe, have sub- 
sequently been found there. ven as late as 1879 some of them 
were discovered by Peach in the Old Red of Scotland. 

(4) The new facts which have been disclosed, more especially those 
which indicate the great richness of the Devonian flora of New York 
in Ferns, now induce me to believe that these St.-John beds, though 
rivalling the Coal-formation in their abundance of fossil plants, are 
really of the age of the Hamilton group of New York, which in 
Europe would be regarded as Middle Devonian. 

(5) I would further add that the richness of this flora in species, 
as well as the discovery of rare and exceptional forms, such as insects, 
is in part due to the excellent exposure of the beds in the vicinity of 
St. John, and in part to the extensive and thorough nature of the 
explorations carried on with the aid of blasting by Messrs. Hartt 
and Matthew, under the auspices of the Natural-History Society of 
New Brunswick. It is probable that few fossiliferous beds in the 
world have been so thoroughly explored. In connexion with this 
it is to be observed that the mass of the specimens obtained repre- 
sents only a few species, while the greater number are represented 


ERIAN (DEVONIAN) PLANTS. 305 


by only a few fragments, which would undoubtedly have escaped 
the observation of ordinary collectors. 

In the collections now in my hands the following forms occur, 
which may be considered new, though most of them are too imper- 
fect for complete description. 


ODONTOPTERIS sQuaMOSA, sp. n. (Pl. XIII. fig. 17.) 


Petiole slender, bearing short pinnules placed at right angles to it, 
and each consisting of two rounded decurrent pinnule and a terminal 
pinnule of triangular form. Toward the end only the terminal 
pinnule appears. Veins obscure, diverging from a midrib, broad at 
base. Frond apparently of a thick or coriaceous texture. 

This would seem to have been a creeping or parasitic Fern. In its 
general habit it bears some resemblance to Cyclopteris dissecta of 
Unger, from the Devonian of Thuringia, but appears to have more 
affinity with the genus Odontopteris than with Cyclopteris. 


CARDIOPTERIS ERIANA, sp.n. (Pl. XIII. fig. 18.) 


Pinnules nearly round or slightly oblong, nearly equally cordate 
at base, somewhat crowded on a slender petiole. Length from 8 to 
14 millim. Veins regularly spreading from the centre of the base, 
curving toward the margin, and forking twice or thrice. 

This is the first appearance of this Lower Carboniferous genus in 
the Devonian. The species closely resembles Cyclopteris polymorpha 
of Goppert, though every way smaller and more delicate. 


Arc moprEris erspaae uel exunil fet al9).,) 


Petiole apparently woody, bearing broadly obovate decurrent 
pinnules, with strong, flabellate, straightish nerves. Pinnules over- 
lapping each other. 

This plant bears a general resemblance to Archwopteris of the type 
of A. (Cyclopteris) Maccoyana of Goppert ; but the woody petiole 
or branchlet, and the coarse texture of the pinnules, raise the sus- 
picion that the specimen may not be a Fern, but may have belonged 
to a coniferous tree of the type of Voltzia or Salisburya. 


CYCLOPTERIS, sp. 


Fragments of a very large cyclopterid leaf, with flabellate veins, 
and which, when entire, must have been three inches in diameter. 
It is too imperfect for description, but indicates a frond of the 
Same general character with Cyclopteris Brownw from Peny, in 
Maine. 

Other specimens indicate a small species of Archwopteris, more 
delicate than A. Jacksoni; and there are some fragments which 
seem to show, though not indisputably, that the submerged leaves of 
Asterophyllites latifolia were long and linear, approaching in form 
to those previously described as A. lenta. A fragment of Hymeno- 
phyllites, about the size and form of H. Gersdorffix, shows minute 
rounded spore-cases comparable with those of the modern genus 
Todea, which the Fern itself also closely resembles. 


306 DR. J. W. DAWSON ON NEW 


The species above described add to the number of small and 
delicate Ferns by which the St.-John beds are so especially charac- 
terized. 

Specimens from Scotland and Australia. 


/XTHEOTESTA DEVONICA, sp.n. (PI. XII. figs. 14, 144.) 


Fruit 4 millim. in diameter, oval in cross section. Testa less 
than 1 millim. in thickness, and consisting of radiating fibres. 
Nucleus represented by white mineral matter with coaly specks. 
The specimen shows only a cross section ; but there seems no reason 
to doubt that it is the seed of the above genus of C. Brongniart*, 
hitherto found only in the Coal-formation of France. It may be 
referred to Taxinee, and may have been the seed of trees of the 
genus Dadowxylon. 

The specimen is in grey sandstone, associated with fragments of 
carbonized plants. It was collected by the Rev. Thomas Bodun, of 
Edinburgh, in the Old Red Sandstone of Perthshire, where it is asso- 
ciated with Lycopodites Millert and Psilophyta. 


DICRANOPHYLLUM AUSTRALICUM, sp. n. (Pl. XIII. figs. 15 & 16.) 


Stem slender, 3 millim. in diameter, not tapering in a length of 
3 inches. It is marked with minute, narrow, elongated leaf-bases, 
spirally arranged. Leaves linear, 3 millim. long, bifurcating at an 
obtuse angle at their extremities. 

The specimen is in white sandstone and is well preserved. It 
was collected by Mr. R. L. Jack, F.G.S., of the Geological Survey of 
Queensland, in sandstones associated with limestone, on Fanning 
River, Burdekin, Queensland. The horizon is said to be under the 
Mt.-Wyatt and Star beds, and consequently lower than that of the 
plants collected by Mr. Daintree, and described by Mr. Carruthers 
in the Journal of this Society. 

The genus Dicranophyllum was established by Grand’ Eury* for 
certain plants of the French Coal-fields, which, though larger and 
better-developed than the present species, must have been some- 
what similar. Grand’Hury regards these plants as probably 
Coniferous. 

The plants described in this paper are fragmentary and imperfect, 
but they add six or seven types to the Hrian flora, and encourage © 
the hope that all the Carboniferous genera may yet be recognized in 
the older formation, together with others peculiar to itself, thus 
tending to vindicate the opinion expressed in a former paper that 
the plant-life of the Devonian was more varied or less monotonous 
than that of the Coal-formation. 


Supplementary Note. 


As some delay has occurred in the publication of the above paper, 
I may be permitted to add the following :— 
(1) In my paper on Devonian Tree-ferns in the ‘ Quarterly 
* Annales des Sciences Naturelles, vol. xx. t Flore Carbonifére. 


\ 


— 


ERIAN (DEVONIAN) PLANTS. 307 


Journal’ of this Society for 1871, I referred, under the names of 
Psaronius textilis and Caulopteris Lockwoodi, to certain remarkable 
trunks of Ferns from the Chemung formation (Upper Devonian) of 
Gilboa, New York, placed in my hands by Prof. Hall and Mr. Lock- 
wood, and which were stated to be from a locality where numerous 
erect trees exist. Prot. Hall has since extracted several of the 
largest of these trees, and they are now in the State Geological Col- 
lection at Albany, where I lately had the pleasure of examining them. 
They entirely confirm my conclusions as to their nature, derived 
from the fragments submitted to me, being evidently trunks of large 
tree-ferns surrounded by masses of aerial roots, in some cases 2 feet 
in diameter at the base, and apparently passing downward into a 
shaly bed or underclay filled with rootlets. Prof. Hall hopes shortly 
to publish illustrations of these remarkable trunks, representing the 
oldest fossil forest yet known. 

(2) In the course of last summer, the researches of Messrs. Ellis, 
Foord, and Weston, of the Geological Survey of Canada, have dis- 
closed, near the head of the Bay de Chaleurs, some interesting 
exposures of Devonian beds rising from beneath the Lower Carboni- 
ferous (Bonaventure formation of Logan). In some of these beds, 
probably Middle Erian, there are abundant remains of Psilophyton, 
similar to those of Gaspé Bay; but others, which are evidently 
upper members of the Hrian system, contain fossil fishes referred by 
Mr. Whiteaves to the genera Pterichthys, Tristichopterus, Phanero- 
pleuron, and Cheirolepis. In the same beds with these fishes occur 
fronds of three species of Ferns, of which I have myself collected 
specimens in a visit to the locality in July last, though the best 
examples have been found by Mr. Foord. One of the species is an 
Archeopteris, allied to A. hibernica and A. Jackson, but differing 
in the details of the fructification, which 1s well preserved (A. magna- 
censis*, MS.). Another is amagnificent fern, referable in the mean- 
time to the provisional genus Cyclopteris, and identical with that 
figured by Lesquereux in the Report of the Geological Survey of 
Pennsylvania (new series) as Archcopteris obtusa. Lesquereux’s 
specimen is from the Chemung or Catskill of Montrose, Pennsylvania. 
A third species is that described by me, in Quart. Journ. Geol. Soc. 
vols. xviii. & xix., as Cyclopteris Brounw. In the specimens from 
Bay de Chaleurs the large flabellate fronds of this fern are seen to 
be attached in dense groups to a rhizome or slender stem, showing 
that this plant was either, as I supposed in regard to the specimens 
which I described from Peny, in Maine, a low-growing ground-fern 
or an epiphyte. 

(3) In the “ discussion ” of my paper I observe a statement to the 
effect that Asteropteris noveboracensis may be a lycopodiaceous plant. 
In reply, I think it sufficient to refer to the description and figure, 
but may add that I have had occasion in previous papers to refer to 
the remarkable abundance and variety of ferns in the islands of the 
Devonian sea.. In accordance with this, the beds near Milo, New 
York, in which Asteropteris occurs, abound in stipes of large ferns, 

* Cape Magnach is the locality. 


308 ON NEW ERAN (DEVONIAN) PLANTS, 


while the only lycopodiaceous plant which they have afforded is 
Lepidodendron primevum. 


DESCRIPTION OF THE PLATES. 
Puate XII. 


Figs. 1, la. Asteropteris noveboracensis, cross section, natural size and enlarged. 
2. A. noveboracensis, portion enlarged, showing one vascular bundle. 
3. , portion enlarged 12 times, showing rays and vascular bundle. 
4, —— , stem restored-in cross section. 
5,6. —— , scalariform vessels, X 100. 
io == , prosenchyma of outer cylinder, x 40. 
8, 9. ——- ——, parenchyma of inner cylinder, x 40. 


In the above figures, a represents radii of axis, 6 cellular tissue 
of axis, ¢ outer prosenchyma, d leaf-bundles. 
10. Equisetides Wrightiana, leaf of sheath. 
ll. Cyelostigma affine. 
, leaf-base, enlarged. 
13. Lepidodendron primevum. 
14. Aitheotesta devonica, natural size and enlarged, 


Prats XIII. 


15. Dicranophyllum australicum. 
, enlarged. 

. Odontopteris squamosa. 

18. Cardiopteris eriana. 

19. Archeopteris, sp. n. 

20. Equisetides Wrightiana. 


Discussion. 


Mr. Carrutuers spoke very highly of the industry of Dr. Dawson 
in collecting fossils, but he could not agree with him in his con- 
clusions as to their systematic relations. He thought the form 
described as a fern should be referred to the Lepidodendroids. 


ON FOSSIL CHILOSTOMATOUS BRYOZOA FROM AUSTRALIA. 309 


4 


24, On Fossit Cuttosromatous Bryoz0a from SoutH-west Victoria, 
AvstRatia. By Arraur W. Warers, lsq., F.G.8S. (Read 
April 25, 1881.) 


[Puates XIV.-XVIII.] 


Parr of the material forming the subject of the present communica- 
tion I received in exchange from Miss E. C. Jelly, in a small test- 
tube, already washed out of the clay; and on two subsequent 
occasions she has kindly lent me a number of slides from her col- 
lection. The “lump of clay ” out of which they were washed was 
sent over to England marked ‘‘ Yarra-Yarra, Victoria,’ by Mr. John 
Allen some years ago; but the exact locality Miss Jelly has been 
unable to obtain for me. However, I find that Mr. H. Watts, in a 
paper “ On fossil Polyzoa” (in the Trans. Roy. Soc. Vict. vi. 1865, 
p. 82-84), mentions a deposit from which Mr. Allen sent fossils, and 
says, “‘ The deposit is described as being about thirty miles east of 
Warrnamboul, extending along the sea-coast for a distance of from 
six to seven miles, and is from thirty to forty feet in thickness.” A 
letter I wrote to Mr. Watts, on the possibility of its finding him, has 
elicited no response; and I therefore presume that he must be 
dead or have removed, and fear that the exact locality will not now 
be discovered. ; 

From the memoirs of the Geological surveys I conclude that it 
will be found to be what the Australian geologists call Miocene, though 
as yet this has not been shown to be of the age of the European 
Miocene formation. Mr. Etheridge, Jun., writes that Allen pro- 
spected in the ‘“‘ quartz cement which was considered by McCoy of 
Pliocene age,” and says “ the country traversed by Allen and party 
also consisted, especially near the coast, of beds which were referred 
by McCoy to the Miocene period.” Mr. Etheridge, Jun., thinks pro- 
bably the material marked Yarra-Yarra came from these beds, and 
there is reason for supposing that the material was not quite 
correctly labelled. The fossils which have been found in Muddy 
Creek, Hamilton, Victoria, seem to support the Miocene age of the 
beds; and similar Miocene clays, according to the Survey, occur in 
several places, as near the mouth of the Aire river and also near 
Geelong. 

Besides the Bryozoa, I found a large series of Foraminifera very 
well preserved; and I spent much time in picking out a large num- 
ber, probably representing 50-100 species, which I forwarded to 
Professor Karrer of Vienna, from whom I hear that he hopes to 
complete the examination of the series this spring ; and I anticipate 
that they will throw some light on the age of the formation, as 
the Foraminifera, both fossil and recent, have had much more 
attention than the Bryozoa. There were also a number of long slen- 
der Jsis-joints, which I submitted to one of our authorities without 


Q.J.G.8. No. 147. 4 


SWO) A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


any result. The few molluscan shells are probably fry, and do not 
give much assistance, but support the ‘“‘ Miocene” age of the beds. 
There are also a few Entomostraca. 

The fragments of Bryozoa are small; but their state of preserva- 
tion is often very perfect ; and in this lies their chief value. Their 
examination may well be used as an introduction to the study 
of the Mount-Gambier series of Australia, in which, so far as I am 
able to judge from the London Geological Society’s collection, kindly 
lent to me, and trom a collection belonging to Mr. Etheridge, 
jun., alsoim my hands, the state of preservation often does not 
admit of the details being seen; so that the determination and de- 
scription of these is sometimes very unsatisfactory if other material 
is not at hand to be used as a key to the structure. As examples I 
may mention Mrcroporella yarraensis, M. violacea, var., Porina cly- 
peata, and Retepora rimata, which occur in the Mount-Gambier beds ; 
but the details are wanting, and were it not for the “ Yarra-Yarra” 
specimens I should have been unable to classify them. 

The living British Bryozoa have recently been reclassified by 
Mr. Hincks, who has introduced in some sections an almost new 
classification. Since the appearance of his work no paleontogical 
papers of any importance have appeared; and it therefore becomes 
necessary to consider how far this classification is applicable in the 
determination of fossils; and it is here that we shall perhaps find the 
weak point in the modern classification: but, on the other hand, 
fossils more than recent forms show the utter unnaturalness of the 
older divisions. Although generic determination will often be diffi- 
cult, that is by no means confined to the present system ; for we often 
find fragments showing many important characters without being 
able to distinguish if they have grown in the Hschara or Lepralia 
form. Every change of classification should, of course, aim at 
making the system more natural; but at the same time special 
attention should be given to those characters which can be distin- 
guished in fossils, seeing that the number of known fossil forms is 
so many times more than all the known recent ones, and ultimately 
the relationship of the living ones must be worked out largely by 
means of the paleontological 1 record. 

In the classification used by Busk in his ‘Crag Polyzoa,’ and, with 
some modifications, by Reuss, the form of the ‘colonial erowth was 
made the first consideration ; so that colonies of cells of ‘a certain 
form incrusting stones or seaweed were called Lepralia, while quite 
similar cells, growing back to back, forming an erect coral-like 
stem, would be called Hschara or, if there was only one layer, 
Hemeschara. With more careful examination and comparison of 
recent and fossil forms this was found to be an absolutely untenable 
position, as the same forms of cells so frequently occur that any one 
well acquainted with recent and fossil Bryozoa could in a short time 
draw up a list of at least 40 or 50 cases where absolutely identical 
. cells are known in the Lepralia and Eschara forms. Smitt*, recog- 


* Krit. Fort. ofver Skandinaviens washichhote se af F. A. Smitt. Ofv. 
Vet. Ak. Forhandl. 1864-1868. 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. Olt 


nizing this, based his classification mostly upon the form of the oral 
aperture; and Mr. Hincks has followed Smitt’s example, introducing 
some modifications. But sometimes there isa peristome obscuring the 
oral aperture; so that it is difficult even in recent species to see the 
shape of the real aperture, unless there is sufficient material at hand 
to make the necessary preparations. As an example, in Tubucellaria 
cercoides, Ell. & Sol., the operenlum which closes the primary aper- 
ture is situated at some distance from the end of the peristome, 
which is produced into a long tube. In fossils, however, it is 
sometimes impossible to make out the exact form of the opercular 
aperture, even when other details are extremely well preserved ; for 
an example of which I only need refer to Porina clypeata (Pl. XVI. 
fig. 67), in which the details are so well preserved that I think there 
is no doubt it can be easily recognized when found elsewhere. 

Probably no naturalist at all thoroughly acquainted with the 
Bryozoa will again attempt to sustain such genera as the old Lepralia 
and Eschara ; but it may be well to examine carefully the growth of 
the Bryozoa before we entirely reject the form of the colony as of 
elassificatory value ; for in many cases it may be shown in this way 
from which part of a zocecium the following zocecium grows. The 
mode of growth of Lepralia and Eschara indicated no structural 
difference ; for the young zocecia in both grew out from the same part 
of the parent cells, and Hschara was only formed of Lepralia-cells 
back to back, often very slightly attached. For an example of the 
new zocecia arising in a different manner we may cite Bicellaria 
and Bugula, and fig. 33, provisionally placed with Cribrillina as C. 
dentipora. The form of the aperture must be the first consideration ; 
but especially among fossils we must carefully notice how they grow ; 
and it is to be hoped that this latter question may soon receive a 
thorough and conscientious investigation, as it is a point requiring 
still further elucidation. At one time I hoped to be able to devote 
some time to this question, which must be a laborious one; but I fear 
the state of my health will not allow me to carry it out. 

If the principles of the old classification had been adopted, it would 
have been necessary to make several new genera; but workers on 
recent Bryozoa have already overthrown the classification based upon 
the zoarial form, and therefore paleontologists should follow Smitt, 
Hincks, and others, and describe their species in such a manner, 
and use such a nomenclature, that fossils can be compared with 
living species. 

The Australian Bryozoa, both recent and fossil, have been as yet 
very imperfectly worked out. The most important works concerning 
living forms are Macgillivray’s* papers, and, for the Catenicellide, 
that of Prof. Wyville Thomsony. Prof Hutton has drawn up a list 


* MacGillivray, P. H., ‘‘ Notes on the Cheilostomatous Polyzoa of Victoria,” 
Trans. Roy. Soc. Vict. pt. ii. vol. iv. 1860; id. “ Descr. of some new Genera and 
Species of Australian Polyzoa,” loc. cet. vol. ix. 1869; id. in ‘ Prodromus of 
Zoology of Victoria,’ edited by F. M‘Coy, decades iii. & iv. 1879. 

t+ Wyville Thomson, ‘‘On new Genera and Species of Polyzoa,” Zool. Bot. 
Assoc. Dublin, vol. i. 1859. 


y¥2 


312 A. W. WATERS ON FOSSIL CHILOSTOMATUUS 


of the New-Zealand fauna ; but, as he does not seem to have devoted 
himself much to the Bryozoa, we cannot make comparison with the 
New-Zealand species until some specialist has worked at them *. 
There are several species described by Busk and Hincks in their 
papers and works; and lately the study has been taken up by Haswell, 
Goldstein, Maplestone, and J. B. Wilson, with interesting results. 
The fossils have attracted comparatively very little attention. In 
1859 the Council of this Society admitted a Note + by Mr. Busk on 
Mount-Gambier fossils ; but this only consisted of a list of 37 species, 
32 of which were new and were not described, but only christened ; — 
so that it still remains perfectly useless for comparison. These names 
have sometimes been quoted ; and workers have wasted their time in 
libraries searching for the descriptions; and in the various biblio- 
graphical and specific lists published by Mr. Etheridge, Jun., these 
names are all referred to in full, and thus some pages are filled up 
with empty names. As Stoliczka points out, such anticipatory pub- 
lication brings confusion without equivalent advantage; and cer- 
tainly in this case there has been much irritation and ink wasted, 
which might have been avoided if the paper had been entirely ig- 
nored. In Mr. J. E.T. Woods’s “Geol. Observations in Australia ” 
entirely unsatisfactory figures without description appeared ; and 
since then Mr. Woods has published a few short papers, to which I 
refer in the descriptive text, of which I will only mention his paper 
“On some Tertiary Australian Polyzoa” and “Australian Selenariade,” 
in which fossils from Mount Gambier and Muddy Creek, Hamilton, 
Victoria, are described, both being considered of the same age. 
Therefore we may say that, with the exception of the papers by Mr. 
Woods and one shortly to be mentioned by Mr. Wilson, no work has 
been done on the Australian fossil Bryozoa; but in the description of 
the Novara Expedition, vol.1. pt. 2, an important paper is published 
by Stoliczka + on the Bryozoa from the marine beds of the Waitemata 
Schichten of Orakei Bay, New Zealand; and as eight species are 
common to both formations, the two deposits are of somewhat the 
same age. ; 
At the time I commenced the study of this material no fossil 
Catenicellidz were known; but since mine were lithographed, Mr. 
J. Bracebridge Wilson § has described and designated by numbers 
twelve species, with none of which am I able to identify any of my 
species. But the characters on which Catenicelle must be grouped 
are as yet scarcely understood; and much change must be made in 
the classification, as undoubtedly some of the names now in common 


* BF. W. Hutton, Catal. of Marine Mollusca of New Zealand, Wellington, 
1873; id. ‘‘Corrections and Additions to the List of Polyzoa in the Catal. of 
Marine Mollusca of N. Zeal.,” Trans. New-Zealand Inst. voi. ix. 1876. 

+ “Note on the Fossil Polyzoa collected by the Rev. J. EH. Woods near Mount 
Gambier, South Australia,” by George Busk, F.R.S., F.G.S., &c., Quart. Journ. 
Geol. Soc. vol. xvi. 1860, p. 260. 

t Fossile Bryozoen aus dem tert. Griinsandstein der Orakei Bay bei Auck- 
land, von Dr. Ferd. Stoliczka, 1864. 

§ ‘* Fossil Catenicellze from the Miocene beds at Bird Rock, near Geelong,” 
Journ. Micr. Soc. Victoria, vol. i. nos. 2, 3, p. 60. 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. ols 


use are synonyms. As with other Bryozoa, so in Cuatenicella, the 
form of the aperture will have to be considered of primary impor- 
tance. This Mr. Wilson does not seem to have appreciated ; for 
only in one case does he mention the shape of the aperture, and, 
while in some the shape may not be distinguishable, we can hardly 
suppose that all the twelve are so badly preserved as to have this 
principal character destroyed. 1 have recently found two species 
of Catenicella in the Geological Society’s collection from Mount 
Gambier. Itis therefore notimpossible that when the figures which 
' Mr. Wilson promises are published, one or two may be found to 
occur near Geelong and in the present locality. 

One most extremely interesting form—=indeed, the most interest- 
ing specimen in the collection—is Catenicella internodia (fig. 78), 
consisting of long internodes with a double row of cells, whereas all 
the Catenicellidze now living have short beaded internodes, consisting 
of one, two, or even three cells; and we may find that forms with 
one or two cells in a node have developed from multicellular nodes, 
and should then ask, have not all jointed forms adapted themselves 
from unjointed ones? This and the Mount-Gambier collections 
furnish unjointed Crisiw. In the living fauna of Australia the 
number of jointed forms is very remarkable; but already in the 
European Chalk the number was very considerable. 

The Microporellidz are well represented, and also show that the 
genus Microporelia must be extended; for we are able to trace rela- 
tionship from WM. violacea, with a round pore, to the var. fissa, with an 
elongate pore (fig. 73); then we have M. yarraensis, with two or three 
denticulated pores in the depression ; and in this way pass on by . 
coscmopora and var. armata, to M. symmetrica (fig. 83). This group, 
with an area with several large pores, was well represented in the 
Hocene and Miocene of Europe by M. coscinopora &c.; and perhaps 
as the recent forms are further studied we shall find several living 
allies; for Microporella (Kschara) distomu, Busk, from Madeira and, 
in my collection, from Capri, from 150 metres, must evidently be 
looked upon as related ; and I find in my specimens from Capri that 
the pores are stelliform, which seems to be a frequent if not general 
character in the group, and is of great interest, as showing a corre- 
lation of characters, and supports the opinion of those who believe 
that we are now on the track towards a more natural classification. 
From the above remarks it will be judged that Porellina, Sm., which 
is separated from the other Microporellide in consequence of having 
the pore in a lunate form, is not considered a necessary genus. 

If the comparison is extended a little further we may find that 
such a form as M. symmetrica (fig. 83) is related to the Cribril- 
linidee through such forms as C. terminata (fig. 68). 

In studying both recent and fossil forms I have often been im- 
pressed with the frequency with which open pores are replaced by 
avicularia, and think that it is a matter worthy of most careful 
examination ; and, considering that there is reason to believe that 
avicularia may be only differentiated pores, I enter into the question 
when speaking of Cribrillina suggerens. 


314 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


The number of species of Chilostomata mentioned in this paper 
is seventy-two; and, so far as my preliminary examination goes, it 
seems that there are about thirty determinable species of Cyclosto- 
mata; so that from this one small lump of clay there cannot be less 
than two hundred determinable species when we add together 
the Bryozoa, Foraminifera, Hntomostraca, and other remains. It 
now becomes necessary to make comparison of these Chilostomata 
with those found in other strata; and we find that a large propor- 
tion are identical with those from the Orakei-Bay beds in New 
Zealand. Stoliczka’s descriptions and figures do not in all cases 
permit of comparison; but out of a total of twenty-nine comparable 
species, there are from Yarra-Yarra seven Chilostomata and seven 
Cyclostomata. With the Mount-Gambier beds the number is large ; 
but this is the result of direct comparison, whereas I have had no 
opportunity of seeing any fossils from Orakei Bay; and the number 
of species from Mount Gambier, known to me, and of which I hope 
shortly to publish a list, is greater than the Orakei-Bay list. The 
number of identical species from the two places is now twenty-three 
Chilostomata; but as I progress with the determination of the 
Mount-Gambier collection the number will no doubt be increased. 
We miss from this locality the Mount-Gambier species Cellaria 
(Melicerita) angustiloba, Busk, and Spiropora verticellata (a species 
common in the European Chalk). The C. angustiloba may be re- 
presented by C. globulosa. 

Twenty-two of the Chilostomata are known recent; and many 
more are very nearly allied to living forms. 

Three species are already known in the Miocene ? Muddy-Creek 
(Victoria) beds, and four Chilostomata and, at any rate, three 
Cyclostomata from the Hocene of Italy; but several others show 
relationship to the Miocene and Kocene forms, and we are frequently 
reminded of European Cretaceous ones, though in only one case 
could we feel justified in identifying the species with any from the 
Chalk. 

It will, however, be well to defer further comparison until the 
list of the known Mount-Gambier species, upon which I am now 
engaged, is completed. 

T have already * referred in various places to my reasons for ae 
the term Bryozoa, which is now universally employed by German, 
French, and Italian authors ; and Ido not therefore need to enter again 
into the question. And although I am sorry that those few English 
authors who have been before the public for a long time have not 
been induced to change the name which they have used for several 
years, yet I have indications which cause me to feel quite confident 
that the next generation of English workers will use the class name 
which has been employed in the rich literature of Germany and 
France ; and I am glad to see that Mr. Tenison Woods 7 has begun 


* “On the Terms Bryozoa and Polyzoa,” Ann. & Mag. Nat. Hist. Jan. and 
Feb. 1880; ‘“ Reply on the Term Bryozoa,” ibid. Aug. 1880. 

t “Corals and Bryozoa of the Neozoic period in New Zealand,” Palzontology 
of New Zeal. pt. iv., Colon. Mus. & Geol. Surv. Depart. 1880. 


| 8. Miicrepora patula ...........-...........- 326)... 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 315 


to use this name, and to hear from Australia that it is there likely 
to immediately replace the term Polyzoa. It is therefore with 
much pleasure that I use the name in this contribution to the 
paleontological literature of that continent. 

Though considering that modifications of the definitions of some 
genera is required through specimens now found, I leave that until 
I have completed my examination of the Mount-Gambier collection 
now in my hands. 

The figures are all drawn with the camera lucida, and, as far as 
possible, magnified twenty-five times; but this is in each case stated 
by the side of the figure. I regret that Plates XV. and XVI. are so 
unsatisfactory through want of success in transferring the drawing 
to the stone; and, in fact, Plate XVII. came out so badly that I 
was obliged to have it very considerably touched up upon the stone, 
and much detail is thereby lost or altered. The Plates are also un- 
satisfactorily arranged, in consequence of my receiving the material 
at various times. 


List of Specves. 


= 
a| 2 
o © A Allies and Localities. 
Sie 5 | 
3 2 )S18 
| Jae 1S 
1. Catenicella cribriformis ............... 317 
2. PERO oo aloe cidennsiinscocs i, 
Pe MATSINALA... 2... cesses eeseeess a 
4. FUDGE:  penebets= Sele see eee eee »> |---| % |...| C. ventricosa, B., living. 
5. PMR PAMEPHILONUSS on. os0cécccscness sss » |*|*|...| Australia. 
6. elegans, var. Buskii, Th. ...... me selec eAcustralia: 
ie SC_L13,  Sosueucolee 318 
8. BRERORILOO =< ja. 0 0022 ote cece aces - 
Me@elaria tistulosa, L.................0000- 319) x | * |...) Miocene and Eocene of Europe. 
10. MEAVEVONISIS. B.- 5c e.-ccoleec tos 321} «| *|...| Falkland Islands. 
11. qricelloces Si ee ee »» |---| * |] C. tenuirostris, var. a, Busk. 
12. 2.0 CLS) ceecgeeeee Pa oe eae ...| Eschara aspasia, d’Orb., Cretaceous. 
9) ORT gS B22)|...| 
mt. Caberea rudis?, Bush .................- » |*|...|...| Bass’s Straits. 
} 15. Membranipora lineata, Z. .....,...... 323) x |...|...] Widely distributed. 
| 16. catenularia, James. .........00008. », | ¥|---|:-.| Pliocene. 
Lid iG 5, |---| ¥|...| WM. Flemingzi, living. 
. —— macrostoma, RSs...............008 ,, |...|%*|* | Bartonian, Italy, and Miocene. 
ILD. Ce B74 doe sal bee Cretaceous. 
fe —— CONCAMECY ALA ..-.... 2.20202 -seneeces a 
see LS 2 a ta Nellia simplex, B., living. 
B —— maorica, Séol. ..........c.cec.cee0s 325| x |...| «| Upper Eocene, New Zealand. 
SEL LTUC. 6 eee eee 


316 


A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


List of Species (continued). 


S 
Pale 
a |= 
|CR Allies and Localities. 
. | alo 
o |S] 35 | 
a Le ios 
AY Hie |O 
25. Cribrillina terminata .................. 5 VA dllaaeload bec Lepralia scutulata, B., living. 
26. denbiporayactcet seo ee i 
27. Sugm@erens |: /.6 4. sscaseee eee 327 Cribrillina tubulifera, Hincks. 
28. Mucronella mucronata, Sm. sae gnats 328] x | x |...) Florida, living. 
29. duplicatal nots neeeee se ee ,, |*|*|...| Lyall’s Bay, New Zealand. 
30. —— elegans, Macg. ...............000655 329|%? 
ole Micropotella violacea, var. fissa ...... » |*|*|...| Indian Ocean. 
32. ferbeal Beare eee ome eet ke 330)...| * 
Do.) ——— Cleyata Hy GOGS ys eae een eee Pree tee 
34. ———— Varraenmsiseensss essence eee dol}...| * 
30. ———— COSCINO POA) eaeee-eseteeee encase a : 
36. —— eNIGMAatICA ...........eceecceveeees ei 
Ol. ———" SY MNMEGLICA) |. 252.56 son scsseoncccees 332 
38. ClavabapSCOlsoun co eneeenen omen uid tenes * 
oo. Porinds chy peata erect eee eee 1. | % 
40. CORE JHU: soepcsbdacsosadesaes 333| «|x| *| Low. Hoc., New Zeal., Bart., Italy. 
4], columnata RRM ited bane AL ce oh Sie esallccalicae Eschara heterostoma, Rss., Hoc., Italy.) 
42. Lepralia corrugata) 222-22. -0--0-6--0--2 335 | 
43. monilifera, var. armata ......... a0 bee ae ee Pliocene of Europe. 
44. ——'spatullatan ce isseecmscnaee .ea eae »» |---| * |---| L. muléespinata, B., Madeira. 
45, cleidostoma, Sm., var. rotunda |336|*?}...|...| L. elecdostoma, Sm., Florida. 
46.- Porellajemend@atay.......-.ess0-scen enone P 
47. denticulatas Stor. ss eee ss x | 
AS. (Smibtiareentralagieey acre erst ood 
49. -——— centralis, var. levigata ......... 5 
50. Mater: EV OOS ik vce aes Sellen tise 
1. aNCepS; ACh All sees caeceeeee near sre 
52. Schizoporella vigilans .................. 538 
538. phymatophora, Rss. ............ sp lescleeeiee .| Hocene of Italy. 
54. ventricosa,, Flasw:\ i.e..ssscseeee Paleo | 
55. === denestratays. 2700255..caescnnesee dence 309 
56. SB) Dslaae Shosenadsognaeec saonnogsocbone: 5 | 
57. SPi sessment meena cen en scen Leys lle ..| S. btaperta. 
D8. —— submersa ............ccseeceseneeees 340 | 
59. a= CONSOR Vata: (ci cccscocsconsockecenee RS * 
60; ———Ispiroporinapeeseeee eee cee a; 
61, ——‘excubans oe ae ee eee io41 
62... amphora ane ee 3 
63. AUS CALS aL OOO Seana eae nee ow glazelsee ...| Muddy Creek (Miocene ?). 
64. Retepora marsupiata, Sm. ............ 342) x | « |x?| Miocene (America). 
65. PUNUAGS Pa clsct tow aneenac soa ae 343]...| x 
66. Cellepora yarraensis................---.- A 
67. fossa, (ast a Ree eee iH x 
68. fe UR abies so aauee tacts esi at Seen’ SY 0 a oe C. pumicosa (B., non Linn.). 
69. Lunulites guineensis, Bush ............ ies 
70. cancellata., Business cr ereenee eae 
71. Selenaria marginata, T. Woods ...... ae 
72 ailatia. Lio WOOdS\ie eins eee Sisal lege eos Muddy Creek (Miocene ?). 


Lod 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 33) 


1. CATENICELLA CRIBRIFORMIS, sp. nov. Plate XVI. fig. 39. 


Zocecia wide, globose, with cribriform area below the aperture, 
with five pores on each side; large lateral avicularia (wings broken) : 
linear vittee nearly to the base; aperture rounded above (broken 
below, probably straight) ; reverse smooth. 

This is larger than C. hastata, and has more pores; and the lines 
in the cribriform area are smaller. 


2. CATENICELLA FLEXUOSA, sp.noy. Plate XVI. figs. 40, 41. 


Cells elongate, irregularly oval, large depressions above and at 
the side of the aperture; cribriform area small, with nine pores 
(fenestre) ; aperture rounded on the distal edge, straight below. A 
wavy thick tube on the front, which may be abnormal; reverse 
smooth, raised over the upper and lower part of the first zocecium 
and over the second zocecium. 

This has some of the characteristics of C’. alata, Thoms.; and, with 
only the one imperfect specimen, it is doubtful if it should be de- 
scribed as new. 


3. CATENICELLA MARGINATA, sp. nov. Plate XVI. figs. 44, 45. 


Zoecia nearly cylindrical, a coffin-shaped margin round the 
zocecium, and marginal bands enclosing depressions near the aper- 
ture of the corneous joint ; aperture rounded above, doubtful below, 
seven distinct pores surrounded bya line; back smooth. 


4, CATRNICELLA AMPLA, sp. nov. Plate XVI. figs. 46, 50. 


Cells large, subovoid, nine large pores (fenestre) on the border oi 
a large area (scutum); small longitudinal ridge above the aperture, 
with a depression on each side of it. Oral aperture rounded on the 
distal edge, proximal edge somewhat arched. Dorsal surface with 
a large grooved depression over the centre of each zocecium ; and by 
the side are two long adjoining chambers, which are probably covered 
by a membrane when living. 

The lateral chambers are very characteristic, and somewhat re- 
semble those in C. ventricosa, Busk; but in ventricosa they are 
lateral but turned partly towards the front, while in the present 
case the direction is dorsal. There is a faint median line, and 
lines branching off to each fenestra, which are not shown in the 
figure in consequence of the lithograph being badly put on the stone. 

Loc. Mount Gambier (Lond. Geol. Soc. coll.). 


5. CaTENICELLA aLata, W. Thoms. Plate XVI. figs. 47, 49, 53. 


Catenicella alata, W. Thoms., “On new.Genera and Species of 
Polyzoa from Coll. of Prof. Harvey,” p. 80, pl. vi. fig. 4 (Zool. Bot. 
Assoc. Dublin, 1859, vol. i.). 

Loc. Living. Fossil, Mount Gambier (Lond. Geol. Soc. coll.). 


6. CATENICELLA ELEGANS, Busk, var. Busxir, W. Thoms. Pl. XVI. 
figs. 42, 48. 


Catenicella Busku, Wyville Thomson, “On new Genera and 


318 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


Species of Polyzoa from the Coll. of Prof. Harvey,” Zool. Bot. Assoc. 
Dublin, vol. i. 1859, p. 83, pl. vii. fig. 2. 

Cells almost cylindrical, contracted towards the base. There have 
been avicularia at both sides above the aperture. Aperture not 
perfectly preserved, rounded on the distal edge, with a constriction 
on each side near the base ; proximal border (smaller than figured) 
slightly rounded. Vitte distinct, with double row of pores in a 
depressed area. 

The vittee of the Oreaee are not correctly figured and deseri- 
bed in the British-Museum Catalogue; for, instead of being raised 
areas with raised warts, they are sunken areas enclosed by raised 
edges, and along these sunken areas there are one or two rows of 
pores, according to the species. The vitte, however, when covered 
with a membrane, may appear like raised areas ; but anexamination of 
the British-Museum specimens shows that the structure was not quite 
correctly appreciated, and it would be a great advantage if any one 
who undertakes the revision of the Catenicellide would give figures 
with all the membranes removed. The size of the recent and fossil 
species is identical. 

In order to compare Catenrcella, Scrupocellaria, and other more 
or less corneous forms, I made a series of calcined preparations of 
most of the recent specimens in my possession. 

This is allied to C. perforata, B 

Loc. Living; Bass’s Straits (7.), Australia (my coll.). 


7. CATENICELLA SOLIDA, sp. nov. Plate XVI. figs. 37, 38. 


Single cell oblong to cuneate, double cells ovate to globular; a 
straight band or ridge down the front of each cell, and a similar 
raised band separating the two zocecia ; a small curved ridge forming 
a circle on each side of the aperture, and below the aperture a linear 
band on each side. On each side of the central bands large pores 
(sometimes double pores). Oral aperture round above, with a small 
sinus on the proximal edge; a denticle on each side of the top of the 
sinus. Dorsal surface with several large depressions, bounded by 
bands similar to those on the front ; the bands, both front and dorsal, 
are grooved. Avicularia very minute, angular or rounded (about 
half the length of the oral aperture), on the side of the cell near the 
distal end. 

This is common in the Yarra-Yarra formation, where, however, 
the cells are usually found double, and the number of single cells 
seen is very limited. The bands vary somewhat in shape; but fig. 38 
_ shows the typical form. This has many points of relationship with 
C. ponderosa, Wilson (Micr. Soc. Vict. vol. 1. p. 63, pl. v. figs. 1- 
3), a living species; also with C. carinata, Busk; and C. species xii. 
of Wilson (loc. cit. p. 63), a fossil from Spring Creek, but I am 
unable to identify it with any described species. 


8. CATENICELLA INTERNODIA, sp. nov. Plate XVI. figs. 78, 79. 


Zoarium in distinct internodes of several zocecia arranged in a 
bicellate series. Zocecia suboblong, with a wide ridge down the 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 319 


middle of the zocecium ; the band spreads out at the top immedi- 
ately below the aperture in an oval form, causing an oval depres- 
sion surrounded by the band. ‘The upper part of the zocecium, 
comprising the aperture and this oval, is slightly raised; in the 
longitudinal depression four or five pores. On the inner side of the 
zocecium, above the aperture; a small flattened boss; on the outer side 
a small triangular avicularium ; a large depression or pore in the 
middle of the central band. Oral aperture semicircular, with the 
proximal edge straight; on the back an irregular oval depression 
behind each zocecium, with from three to five pores in each of these 
depressions. 

This is allied to C. solida, Waters (figs. 37, 38); but the shape of 
the aperture is different, besides considerable difference in the 
structure of the back. At first I thought the large holes in the 
middle of the centre band were accidental; but this is really not 
the case, as in the cells, when there is no hole there, it is replaced 
by a dark round spot, probably of thinner shell. 


9. CeLtaria FistuLtosa, L. Plate XIV. figs. 1, 2, 10, 11. 


Cellaria fistulosa has occasioned systematists the greatest difficulty ; 
and it now possesses a mass of synonyms (for which see Hincks 
and Reuss), of which many can never be compared. When 
J examined these and the allied species from this Victorian mate- 
rial, I at first felt that it would be impossible to separate them, 
and thought with Reuss that the range of variation was so great 
that such forms as C. fistulosa, sinuosa, crassa, marginata, &c. must 
be included under one specific name. Since my preliminary exami- 
nation My. Hincks’s most valuable work on the British marine 
Polyzoa has appeared, in which he distinguishes three species, 
fistulosa, sinuosa, Johnson. I have therefore, with this book before 
me, again very carefully examined my recent specimens, first cal- 
cining considerable portions; and the conclusions I have come to 
are :—that the shape of the cell is so variable that it is perfectly 
useless as a character (this has already been mostly recognized by 
recent writers, but was the character on which the species were 
some time ago principally founded); then I next found that the 
bordering rim, which is a character of C. Johnsoni, Busk, is some- 
times found on one part of a colony of C. fistulosa, and absent in 
other parts; next I found the shape of the ovicellular opening 
equally unsatisfactory (for in most undoubted specimens of jistulosa 
from Naples it occurs in some cells as a minute orbicular opening, 
then it is elongate oval, and in other apparently older ovicells a broad 
semicircular line is formed, which changes to a transversely oval 
opening, resembling that figured by Mr. Hincks as a character of 
C’. sinuosa). In the same specimen, before any ovicells are formed, 
the aperture is very near the top of the zowcium; but afterwards 
its position is near the centre. Having found the position of the 
aperture, the shape of the ovarian opening, the shape of the 
zocecium and of the bordering rim unsatisfactory characters, there 
only remained the avicularia; and in all the specimens I have ex- 


320 A. W. WATERS ON FOSSIL CHILOSTOMA TOUS 


amined I have found one form constant: the jistulosa from the 
Mediterranean has its rounded avicularium above the zocecium ; the 
stnuosa has a diagonal avicularium pointed downwards, with the 
lower part raised; the Johnsoni, from Rapallo (Italy) and New 
Guinea, has a zocecial avicularium with a projecting hood above, 
as figured by Hincks. I find my observations on the recent species 
entirely confirmed by the examination of a large number of fossil 
forms. 

Some thick specimens, such as fig. 1, have the characteristic avicu- 
laria of the European jistulosa ; these are somewhat rare ; but slender 
specimens, such as fig. 1Z, are common, and probably these are the 
same species; but as I have been unable to find any avicularia in 
these, both from Yarra-Yarra and Mount Gambier, the point can- 
not be decided with certainty. In the specimens figs. 10, 11, how- 
ever, I find a small wide avicularium, as in larger fistulosa, with 
which these must therefore be united. The shape of these last 
resembles that of C. Johnsont, which is the C. marginata, Reuss 
(Lert. Wien, p. 59, pl. vu. fig. 29, not 28); and I have slender 
C. Johnsoni from off Raton (New Guinea) resembling figs. 10, 11 
in all except the avicularium, which is the large rounded one with 
which we are already acquainted in C. Johnsonz. 

In fig. 11 we have an interesting specimen, as showing how very 
slender they may sometimes be; this has only one longitudinal 
row of cells on each of the four faces. In specimens from Mount 
Gambier, like my fig. 12, I have occasionally found in the oral 
aperture two teeth above and two teeth below, and in the ovarian 
openings sometimes one set of teeth, sometimes two; but these 
cannot be looked upon as constant characters, and lead us to think 
that C. crassa must be considered a doubtful species; or perhaps 
two species are represented, seeing that avicularia of two kinds 
are figured. I also sometimes find in the recent C. Johnsoni, from 
Rapallo, two such teeth in the upper part of the aperture, as well as 
the two below. 

Fig. 1 shows that the joints of this specimen were attached by nu- 
merous horny tubes. In recent C. tenurrostris and C. malvinensis the 
joints are usually thus attached ; but this is not the case inC. fistulosa; 
but in specimens of C. Johnson, from Rapallo, there are sometimes, 
though not usually, several such connecting tubes. This leads us 
to the consideration of these joints; for when a stem divides and 
two new branches are formed, the calcareous wall is continuous, 
and in some the branches are already large (perhaps throwing out 
fresh branches) before this calcareous wall is broken through. Of 
specimens J examined, I found this calcareous structure remained 
continuous longest in C. Johknsonz, but scarcely at all in C. gracilis, 
while C. fistulosa occupied a mean position; hut much might de- 
pend upon the sea in which each specimen grew, as jointed structure 
in this and other genera must be looked upon as an adaptation to 
moving water. In fossil specimens from the Pliocene &c. perma- 
nent ankylosis, as already pointed out by Busk and Hincks, is 
frequent. I‘am not inclined to think that, as a rule, articulation 


tl er 


; 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 321 


can be made a basis of generic classification; and perhaps at some time 
the geologival record may show us that jointed forms are all derived 
from unjointed ones. 

Loe. Living, widely distributed. Fossil: Mount Gambier, com- 
mon ; and also from the Miocene and Pliocene of Europe; but as we 
cannot be quite sure from the descriptions of Reuss and others that 
Salicornaria farciminoides does not include other species, it is best 
to refrain from giving localities. 


10. Crttaria Matvinensis, Busk. Plate XIV. fig. 3. 


Salicornaria malvinensis, Busk, Mar. Poly. p. 18, pl. lxiui. figs. 1, 2. 

The zoarium is about the same size as Cellaria fistulosa, fig. 1; 
sometimes the cells are hexagonal, at others more acute above and 
below ; an acute avicularium replaces a zocecium. ‘The front of the 
zoceclum is very much depressed, being surrounded by a raised bor- 
der; distal edge of the oral aperture rounded, contracted above, so 
as to suggest a subtriangular shape; proximal edge curved inwards, 
with a small tooth at each side. 

Loc. Living: Falkland Island, South Patagonia (Darwin). Fossil : 
Mount Gambier, common (Lond. Gol. Soc. coll. & Eth. coll.). 


11. Crtraria ovicettosa, Stol. Plate XIV. figs. 4, 5, 6; Plate 
XVII. fig. 62. 


Salicornaria ovicellosa, Stol. Foss. Bry. Orak. p. 151, pl. xx. 
nes. 9, 10. 

From Yarra-Yarra I have several small fragments; and in some 
there is a rim round the cell, as figured by Stoliczka, and in others 
it is plain, as in fig.4. The most interesting specimen is one in the 
possession of Miss Jelly (fig. 62), which shows the great range of 
variation and the great difficulty in determining species on the 
characters generally used—as, for instance, the mouth varies in posi- 
tion in the fertile and unfertile cells, and the zoarium, in this case, 
is quite thin (as in fig. 4) in the upper and lower parts; but in the 
middle the diameter is more than twice as great, and in this 
thicker part, where the cells are fertile, there are very large acute 
raised avicularia above the zoccium, occupying nearly as much 
space as the zoecium. Fig. 5 shows the position of the distal 
rosette plates, which are not quite in the centre of the zoarium. 
Fig. 6 shows the two lateral plates on each wall. 

This is evidently very closely related to Cellaria tenuwirostris 
(especially var. a) of Busk, from Bass’s Strait, Tasmania, Florida, 
&c.; butit differs in having the upper part of the avicularium much 
projecting, and the rim is regular instead of being raised at the two 
sides. 

Loc. Orakei Bay, New Zealand (St.); Mount Gambier (Lond. 
Geol. Soc. coll.). 


12, CELLARIA GLoBULOSA, sp. noy. Plate XIV. figs. 16, 17. 


Zoarium consisting of joints, from the top of which arise two 
sunilar joints connected with the first by corneous tubes; internodes 


OLE A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


short, laterally subcircular, compressed. Zocecia quincuncially ar- 
ranged, hexagonal ; oral aperture arched above, lower lip straight, 
with two distinct denticles, aperture placed one third the length of 
the area from the top; zocecial area slightly depressed. 

This differs trom Cellaria (Kschara) aspasia, d’Orb. (Pal. Fr. 
p. 132, pl. delxvu. figs. 14, 16), in having the cells distinctly hexa- 
gonal instead of spatuliform, and from Melicerita angustiloba, Busk, 
in having the cells arranged quincuncially instead of in transverse 
rows; consequently the distal borders are straight and in con- 
tact in C. globulosa, while the lateral borders are straight in 
M. angustiloba, B. 

From specimens I have seen from Mount Gambier, Melicerita 
angustiloba certainly seems to be a jointed species, and should be 
united to Cellaria, The genera Melicerita and Hscharinella are 
separated by d’Orbigny in consequence of the first having the cells 
transverse, while they are quincuncial in the second. Latereschara 
was also divided from Hschara on the same grounds. 

Mr. Vine has in his possession a node from this locality, which is 
very irregular and elongate. 


13. Canpa FossILis, sp. nov. Plate XVI. figs. 51, 52. 


Cells biserial ; aperture elongate, rounded above, contracted below; 
upper margin recedent, with a spine on each side. Surface granu- 
lated. An avicularium (or spine) on the inner side of each zocecium, 
placed about one third of the length of the zocecium from the top. 
On the side a large pore for tubular fibre, above which is a vibra- 
culum ; on the dorsal surface cells arranged diagonally. 

This much resembles Canda arachnoides, Lamx., but differs in 
having a smaller aperture, and in the pieces found there is no 
median avicularium ; but possibly this may exist, as the diagnosis is 
based on a few small fragments. In C. arachnoides the cells on the 
dorsal surface are parallel with the median line.’ 

Loc. Mount Gambier (Lond. Geol. Soc. coll.). 


14. Caprrea rupIs, Busk?. Plate XVIII. fig. 86. 


Caberea rudis, Busk, Cat. Mar. Polyz. p. 38, pl. xlvi. 

Zocecia elongate, oblong; aperture elliptical, occupying about one 
half of the front of the zocecium; area sloping inwards, very minutely 
granular ; a spine on each side at the top of the aperture, and a small 
opening (spinous or radicular) beyond the rim of the area; a small 
avicularium on one side below the area. So-called operculum large, 
entire, elliptical. Ovicell much raised, opening arched, rather flat- 
tened in front, with a line round the flat region. A small avicu- 
larium on one side near the top of the ovicell. ; 

The fragment only consists of one row of cells; and therefore it is 
impossible to be quite sure of the determination ; but if not rudis, it 
approaches very closely to it, and the vibracula apparently cor- 
respond. 

The preservation of the operculum in so fragile a specimen was 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. O20 


surprising ; but I have also a specimen of Caberea from Mount 
Gambier with all the operculum remaining. 
Loc. C. rudis is found in Bass’s Strait (B.). 


15. Mempranrpora LInEATA, L. 


I have a small fragment of Membranipora consisting of two elon- 
gated cells with large area, with an acute avicularium above the 
cell, with beak pointing downwards, about sixteen large spines 
round the cell, and two smaller ones just above the border on the 
distal extremity. This seems to vary from the living lineata only 
in having more spines; but from so small a fragment it is impos- 
sible to speak with certainty. 


16. MEMBRANIPORA CATENULARIA, Jameson, 


For syn. see Hincks’s Brit. Polyz. p. 134. 

The specimens are small, consisting of only a few cells, but cor- 
respond exactly with recent specimens | have from the Semaphore, 
Adelaide, Australia. 

Living: Northern seas, Brit., Medit., Canada, Australia, Fossil: 
Pliocene of England and Italy. . 


17. MemMBRANIPORA CYLINDRIFORMIS, sp. nov. Plate XVII. fig. 74. 


Zoarium cylindrical. Zocecia not distinctly separated; area con- 
tracted above, expanded below, filled in for about a third of its 
length with a calcareous granular lamina. Aperture subcircular, 
flattened below, margin much raised ; oral spines, two large and two 
small ones; small acute triangular avicularium above the area, and 
larger more elevated ones below placed transversely. 

From the material marked “‘ Yarra Yarra” I have only found a 
small piece ; but from Mount Gambier I have larger cylindrical pieces. 
The form of the zocecia is very similar to that of M. Flemingi. 

Loc. Mount Gambier (Lond. Geol. Soc. coll. and Eth., jun., coll.). 


18. Mempranipora macrostoma, Rss. (in Vincularve forma). Plate 
MVS figs. 13, 19. 


Cellaria macrostoma, Rss. Foss. Polyp. d. Wiener Tert. p. 64, 
pl. vill. figs. 5, 6. 
" Biflusira macrostoma, Rss. Die foss. Anth. u. Bry. der Sch. von 
Crosaro, p. 274 (62), pl. xxxiil. figs. 12, 13; Ak. Wien, vol. xxix. 

? Biflustra papillata, Stol. Foss. Bry. Orak. p. 154, pl. xx. 
figs. 14, 14a. 

Flustrellaria macrostoma, Manzoni, I Brioz. foss. del Mioc. d’Aus- 
tria ed Ungh. p. 67 (19), pl. xiii. fig. 46 ; Ak. Wien. vol. xxxvii. 

Zoarium subcylindrical, or much compressed, a number of vari- 
cells. Zocecium distinct, with a well-marked border. Aperture 
elongate oval, occupying nearly the whole of the front of the cell; 
sides of the area folded inwards, so that the aperture is in a deep 
depression. Small raised avicularia above the zocecium in a hori- 
zontal or diagonal position. Distal rosette plates two, near the front 
of the distal wall. 


324 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


Aperture 0:46 millim. long. 

Loc. Oligocene (Bartonian), Val di Lonte (Ztss.), Brendola, Colle 
Berici, Ferrara di Monte Baldo (A.-W.-W. coll.) ; Miocene, Nussdorf 
(Manzon2), Orakei Bay (Stol.), Mount Gambier (Lond. Geol. Soc. coll.). 


19. Mrempranipora AReus, d’Orb. (in Vinculariew forma). Plate XIV. 
figs. 20, 21. 


Vincularia argus, @Orb. Pal. Franc. p. 253, pl. delxxxix. 
figs. 1-4. 

Zoarium erect, slightly compressed. Zocecia quincuncially ar- 
ranged in longitudinal lines, distinct, irregularly hexagonal, con- 
tracted below, not raised above the general surface. Aperture oval, 
occupying about one third of the zoecium. Two distal rosette 
plates near the centre of the distal wall; lateral rosette plates 
several close together (fig. 20, a). 

This differs from M. macrostoma in having a very much smaller 
aperture, which is not in so deep a depression. The aperture is 
0-31 millim. long. 

Loc. Cretaceous, Meudon (d’Orb.). 


20. MEMBRANIPORA CONCAMERATA, Sp. nov. (in Vencularie forma). 
Plate XIV. figs. 22, 23. 


Zoarium slender, erect, hexagonal in section, dividing dichoto- 
mously, about 0°5 millim. in diameter. Zocecia in longitudinal 
lines quincuncially arranged, distinct, with raised border, concave, 
elongate, contracted slightly below; border much raised above the 
aperture, forming a raised arch. Aperture oval, occupying from one 
third to one quarter of the length of the zocecium. Zocecial avicu- 
larium at the junction of each branch. One distal rosette plate 
near the centre of the zoarium (fig. 23); two lateral rosette plates 
(fig. 22, a). 

This is the most common fossil in the material, and much resem- 
bles Vincularia gracilis, d’Orb., from the White Chalk of France, 
but differs in being smaller and having a smaller aperture, and 
having six cells in a series instead of eight ; but we now know that 
this last is not an important difference ; perhaps the two should be 
considered varieties of one species. 

Aperture 0°14 millim. long. 


21. MEMBRANIPORA LUSORIA, Sp. NOv. (in Vincularie forma). Plate 
XIV. fig. 14; Plate XVIII. fig. 82. 


Zoarium ep lind eal) jointed, dichotomously dividing. Zocecia 
suboval, rounded above, contracted below, distinct, surrounded with 
an elevated margin raised above the aperture. Aperture large, about 
one third the length of the zocecium, oblong, rounded at the corners, 
contracted towards the middle. Very large elevated zocecial avicu- 
laria directed downwards, with acute mandible. 

Aperture 0-2 millim. long, 0-11 wide. 

The form of the cell suggests a battledore, or, better, snow-shoes ; 
and therefore, finding some difficulty in giving a distinctive name not 


Quart. Journ. Geol. Soe. Vol. MX PE 


Mintern Bros , imp, 
VICTORIAN FOS Sil BRVvOZzea. 


A.W. Waters lith. 


Quart. Journ. Geol. Soc. Vol. XXXVILPL XV. 


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VICTORIAN FO Soil, BRYOZOA: 


BRYOZOA FROM §.W. VICTORIA, AUSTRALIA. 325 


employed in this or neighbouring genera, I have allowed myself to 
stretch a point in choosing a name. ‘This would belong to Wellia of 
Busk; but that is a genus which, as defined, cannot possibly be 
maintained, and the present species with zocecial avicularia shows 
how unsatisfactory a character the absence or presence of avicularia 
is. Nellia, Busk, is almost synonymous with Quadricellaria, d’Orb. 

This is very closely allied to Nella simplex, Busk, from Prince of 
Wales’s Channel, Torres Strait ; the aperture in that case, however, 1s 
0-5 millim. long and a reguiar ellipse, while in M. lusoria it is only 
0-25 millim. long. Fig. 14 was drawn from a worn specimen with- 
out avicularia, before I had seen the one figured as 82; and then 
I could not make out all the structure. 


22. MemBranipora MAoRIca, Stol. (in Vincularie forma). Plate XIV. 
fig. 9. 

Vincularia maorica, Stol. Foss. Bry. Orak. p. 153, pl. xx. fig. 8. 

Vineularia? maorica, Hutton, on some Austral. Polyz. p. 23, 
Rep. R. Soc. Tasmania, 1877. 

Vincularia maorica, T. Woods, “ Corals & Bryozoa of the Neozoic 
period in New Zealand,” Paleont. of New Zeal., Col. Mus. & G.S. 
Dept. 1880, p. 27. 

Zoarium erect, with six cells in a series, or, thickening out, in 
places there are eight. ocecia with a raised border, rounded, with 
a tendency to be hexagonal, calcareous, front of the zocecia de- 
pressed, two tubercles above each zocecium. Aperture (usually 
rather larger than figured) slightly trifoliate, distal edge rounded, 
and proximal edge straight. Avicularia large, occupying the place 
of a zocecium. ‘Two round rosette plates near the middle of the 
distal wall. (In the same position as shown in fig. 21.) 

It is evident, from the description and the figure of the section, 
that Stoliczka’s specimen was somewhat worn; and in such a case 
the cells would assume an entirely hexagonal form. The presence 
of the tubercles is indicated in his figure of the section. The avicu- 
laria are the same as those so common in many of the fossil Mem- 
branipore-—for example, M. angulosa, Rss. (see my figure in Bry. of 
Bay of Naples, pl. xii. fig. 3, Ann. & Mag. Nat. Hist. s. 5, vol. ii1.); 
and it is interesting to find this with a distinctly Vincularian mode 
of growth. Smitt, when he broke up the old genera Lepralia and 
Eischara, indicated the colonial form by adding “ Lepralia-, Hschara-, 
or Hemeschara-forma”; and we may further extend the principle 
by speaking of Vincularca-forma: thus the form of cell and the 
generic relationship will be shown, together with the mode of 
growth. 

Loc. Living, Tasmania (Hutton). Fossil: Orakei Bay ; Hutchin- 
son’s Quarry, Oamaru (New Zeal:); Upper Eocene of New-Zealand 


geologists. 


23. MEMBRANIPORA GEMINATA, sp. nov. Plate XVI. fig. 55. 
I have only found a small fragment of this; but it seems to be 
a branching form, with two cells diagonally to one side, followed by 
Q.J.G.8. No. 147. Z 


526 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


two cells turned towards the other side. Area an irregular oval, with 
margin thickened, much raised, surrounded by numerous spines. 


24, MicRopora PATULA, sp. Nov. 


Zocecia distinct, divided by a scarcely raised thin border, sub- 
quadrangular, finely granulated with small pores, more numerous 
round the edges; distal portion of the zocecium much depressed, 
raised above the distal edge of the aperture and forming a hood over 
it. Oral aperture rounded above, straight below, with a much raised 
lip or rim. 

Aperture 0°25 millim. wide, 0°14 long. 

This has some resemblance to Vencularia binotata and cucullata of 
Reuss and Manzoni; but the fragment is so small that we do not 
know if it grewin the Vincularia-form. The mouth is much larger ; 
and also the zocecial border is less distinct. 

Loc. Fossil: Mount Gambier (Lond. Geol. Soc. coll.). 


25, CRIBRILLINA TERMINATA, sp. nov. Plate XVII. fig. 68. 


Zoarium in the Hemeschara-form. Zocecia but very slightly raised, 
the plain borders of neighbouring zocecia nearly confluent, so that 
the divisions of the zoccia are scarcely visible; cribriform area 
somewhat depressed, the large pores concentrically arranged. Oral 
aperture rounded on the distal edge, straight on the proximal, con- 
tracted below. Small rounded avicularium above the aperture, 
a small one immediately below, a little to one side, and a small one 
on each side, just below the line of the aperture. 

Oral aperture 0-13 millim. wide at base, 0°17 at widest part, 
0-18 long. ; 

This seems to have much in common with Lepraha scutulata, 
Busk (Q. J. M.S. vol. iii. 1855, pl. ii. figs. 1, 2); and it is difficult 
to know where it should be placed, as the pores do not seem to show 
any radial structure, and the aperture is contracted below, having 
the same shape as the oral aperture of Diporula verrucosa. It also 
has many points in common with C. punctata, and is one of those 
types which appear to be connecting links between widely separated 
forms. 


26. CRIBRILLINA DENTIPOR, sp. nov. (in Bactridii forma). Plate XY. 
fig. 33. 


Zoarium erect, with zocecia only on the front; zocecia growing 
from the side near the distal end of the zocecium below, the axis of 
each zocecium being thus diagonal to each of its neighbours. Zocecia 
with a large raised cribriform area, the pores being placed concen- 
trically, each pore with a denticle pointing away from the centre of 
the area. Oral aperture semicircular, with a straight proximal 
edge, 0°14 millim. wide. A small avicularium directed outwards 
on each side of the aperture. Dorsal surface with a central groove, 
and smaller depressions on each side. 

The fragments are small, and not sufficient to found a new genus 
upon, which may possibly be necessary when large pieces are found ; 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. By247( 


for the zocecia originating at the side is an important characteristic. 
The form of the cell, however, very closely resembles that of Cri- 
brillina terminata (fig. 68); and by the present designation we, at 
any rate provisionally, show the zocecial relationship and the mode 
of the zoarial growth. After finding that Schzzoporella phymatopora 
(fig. 32) showed, when broken, a structure like Bactridium, I re- 
examined the present fragments, which, however, seem to belong to 
a form with cells on one side only, growing erect with only one 
series of cells. The British-Museum specimen of Lepralia monoceros, 
Busk, shows that the pores in that species also are provided with a 
denticle ; but in that case the denticles are irregular. 


27. CRIBRILLINA SUGGERENS, sp. nov. Plate XVII. fig. 75. 


Zoarium in Aschara-form, consisting of two layers of cells grow- 
ing back to back. Zocecia oval, distinct, quincuncially arranged, with 
a double row of from twelve to fourteen erect tubular projections 
on the front, and a smaller row on the side, and some smaller ones 
in the central area ; large pores between the tubes. Oral aperture 
small, semicircular, with straight proximal edge; in one case two 
curved irregular spines form a sort of arch over the aperture. A 
short triangular avicularium above the aperture on one side. Two 
oblong lateral rosette plates, one oblong distal plate. 

Proximal edge of aperture 0°06 millim. wide. 

This is a most curious and instructive form, in which we are at 
the outset met by a difficulty as to its generic position; for, looking 
at the aperture, we find it might belong to Cribrillina or Mucronella. 
With the latter, however, in other respects there is little in common ; 
but with Crébrillina we find the radiating character of the pores, 
and, although no known species has such a bristling surface, yet in 
C. Gattye, C. cribrosa, Hell.*, C. figularis, &c. there is a row of 
slightly raised pores round the edge of the cribriform area. 

These very curious prominent tubes (about 0°07 millim. high) 
naturally lead to the consideration of the signification of the pores 
and of avicularia; for may not avicularia have originated in such 
tubes having covers? and we may again ask, is the function of 
these tubes different from that of the pores which we almost univer- 
sally find on the surface of the Bryozoa? The physiological signi- 
fication of these pores is, perhaps, not sufficiently appreciated ; and 
therefore a moment’s digression is necessary, as we may thus have 
the record even in fossils of physiological structure. When a recent 
cell is decalcified, the membrane remaining shows small disks where 
the calcareous pore has been, and a spot in the centre of this disk 
further shows that to each one a thread of the endosarc has extended ; 
so now we see that oxygenation, or a similar vitalizmg change, takes 
place by means of the communication with the exterior through these 
pores. The front of a cell, however, frequently becomes covered 
with mud or organic growth; and in such cases raised tubes might 
be of great advantage in the economy of the colony: and if these 

* See my fig. 4. pl. ix. in Bry. of Naples (Ann. & Mag. Nat. Hist. ser. 5, 
vol, iii. 1879), to which I have now to make addition. 6 

Z 


328 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


are provided with a lid or ‘“‘beak,” they may still longer render 
assistance to the colony ; for the polypide of a cell may be dead, 
and the cell itself covered with mud, but yet the zoarium may be 
living, and under favourable circumstances a new polypide may 
grow in this very zocecium. 

Modifications may take place, and the avicularia become highly 
differentiated, until we find them occupying the place of a zocecium 
and of equal individual value with the zocwcia. Tracing it thus 
from below instead of from above, as is usually attempted, we get 
a much more reasonable explanation of the origin and function of 
the avicularia; for certainly when we find very minute avicularia 
on the root, andon the back of the zoarium, besides large numbers on 
the front of the cell, the prehensile theory seems utterly to fail; and 
although the present idea requires further development, yet the 
examination of living forms seems to furnish much to support it. 

Smitt points out that in Nellia oculata, B., the papilliform pro- 
cesses are avicularia, sometimes with and sometimes without covers 
(mandibles) ; and we are constantly meeting with similar instances. 

Mr. Hincks has kindly furnished me with an advance plate and 
description of Cribrillina tubulifera, Hincks (see Ann. Nat. Hist., . 
July 1881, p. 8, pl. i. fig. 7), to which the present is closely related. 
The only important difference seems to be the relative size of the 
oral aperture, which is very small in the fossil. The specimen now 
described is very small; and it is to be hoped that a larger specimen 
may be found, upon which the affinities can be more exactly studied. 


28. MucronELLA MucRonaTA, Sm. Plate XVII. fig. 66. 


Kscharipora mucronata, Smitt, Floridan Bryozoa, p. 24, pl. v. 
figs. 113-115. 

? Eschara Lnversidge, T. Woods, ‘‘On some Tertiary Australian 
Polyzoa,” Tr. Roy. Soc. New 8. Wales, 1876, p. 3, figs. 11, 12, 13. 

Zocecia obscure, slightly elevated; aperture rounded above, straight 
below, with a mucro or broad, flat, plate-like expansion in front of 
the proximal edge; on the front of the zocecia usually five large 
pores partially closed by a large, simple or bifurcate denticle, some- 
times only three or four large pores, when there is usually a small 
rudimentary pore. Angular avicularia at the sides of the zocecia ; 
the two are usually unequal in size; a minute avicularium in the 
aperture. Probably incrusting or Hemeschara-form, from Yarra- 
Yarra. Aperture 0°23 mm. wide. 

In the Mount-Gambier specimens in the Lond. Geol. Soe. collec- 
tion there are usually only three pores, but sometimes four or five ; 
and although the avicularia are not so distinct, and were not figured 
or described by Mr. Woods, yet they are to be seen at the sides of 
the zocecia. 

Loc. Living: W. off Tortugas, Florida. Fossil: Mount Gambier 
(Woods and Lond. Geol. Soc. coll.). 


(29. MucRoneLna DUPLICATA, sp. nov. Plate XVI. fig. 54. 
The fragment is small; but some characters are shown which are 


BRYOZOA FROM 8.W. VICTORIA, AUSTRALIA. 329 


important, and by which it may be recognized again should it be 
found elsewhere. 

The aperture is orbicular, with a mucro turning inwards over the 
aperture. On each side, below the aperture, there is a large avicu- 
larium, also one on each side two thirds of the distance from the 
next aperture. Above the aperture there is a raised area with a flat 
surface, apparently an ovicell. Above the ovicell and down the sides 
of the zocecium there is a row of large pores. 

The peculiar flatness of the ovicell is not shown in the drawing. 

A specimen sent over to the British Museum by Mr. Hutton from 
New Zealand, marked and mentioned in his catalogue as Lepralia 
variolosa, seems to be this species. In the recent species the ovicells 
have a small umbo, and there is not a second pair of avicularia 
guarding the ovicells ; but the position of the upper avicularia is 
the same, and the shape is similar. JV. duplicata also occurs from 
Mount Gambier (Lond. Geol. Soc. coll.) with the zoarium erect, cy- 
lindrical, with one cell turned in each direction. 


30. MucroneLia ELEGANS, Macg., var.? Plate XVIII. fig. 91. 


? Eschara elegans, Macgillivray, “ Austr. Poly.” Trans. Roy. Soc. 
Vict. pt. ii. vol. ix. 1869, p. 138. 

Zocecia quadrate, surface finely granular with pores, except just 
below the aperture, a round avicularian pore immediately below the 
aperture. Oral aperture rounded on the distal edge, the proximal 
edge arched, forming a lip, a minute denticle on each side of the 
mouth immediately above the angles. Aperture 0°17 mm. wide ; 
0-1 long from the top of the lip to the distal end of the aperture. 

Only two cells are preserved ; and therefore it is impossible to be 
sure of even the generic position of the specimen ; but it is evidently 
related to the species from Queenscliff and Portland Bay, described 
by MacGillivray as Eschara elegans, a name that should not have been 
given, as there is already H. elegans of Milne-Edwards. In #. elegans, 
MacG., no suboral avicularia are mentioned; but according to Mac 
Gillivray avicularia are sometimes situated at the side of the mouth. 


31. MrcroporELLa vioLAceA, Johnst., var. rissa. Plate XV. fig. 26; 
Plate XVII. fig. 73. 


Microporella fissa, Hincks, ‘Contr. Gen. Hist. of Mar. Poly.,’ 
Ann. & Mag. Nat. Hist. ser. 5, vol. vi. no. xxxv., Nov. 1880, p. 381, 
pl. xvi. fig. 4. 

Zoarium in Eschara-form, flat, compressed. Zocecia pyriform 
or oval, much raised, surrounded by a row of pores, or in old and 
worn cells covered with pores; a raised protuberance on each side of 
the zocecium a little lower than the oral aperture; an avicularium 
immediately below the aperture, sometimes turned to the right, some- 
times to the left ; immediately below the avicularium there is a deep 
depression, and the opening of this depression seen from the interior 
is found to be an elongated denticulate pore (see fig. 73). Oral 
aperture rounded on the distal edge, straight on the proximal, 0°07 
mm. wide. 

It will be seen that this resembles the form of M. violacea called 


330 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


plagiopora, from the Crag, but differs in having an elongate pore; 
Escharipora trifora, Roem., from Sollingen, belongs to one of these 
forms. The old basal cells, where the oral aperture is obliterated, 
show a central depression for the pore and avicularium; and the 
whole cell stands out in a mamillated manner. 

I have one specimen in which all the zoccia are very much 
swollen on one side without any protuberance on the other; and 
in the same specimen, at the lower part of the avicularium, there are 
three openings, by which the mandible was probably attached. 

Loc. Specimen in the Geol. Soc. coll. from Mount Gambier: but 
from this alone the detail could not have been worked out. Living: 
Indian Ocean (Z.). | 


32. MIcROPORELLA FERREA, sp. nov. Plate XVII. fig. 72. 


Zoarium erect, in Hschara-form, compressed, few cells. Zocecia 
indistinct, slightly raised round the oral aperture, depressed towards 
the centre, with a median pore. Oral aperture rounded on the distal 
edge, contracted towards the proximal edge, which is straight, thus 
having the shape of a circle with the lower third cut off. Small, 
erect, oral avicularia in the aperture, in one row on the right, in the 
next on the left; small avicularia and large pores round the raised 
portion surrounding the aperture; large zocecial avicularia on the 
side of the zoarium, with the opening. looking downwards. Oral — 
aperture 0-14 mm. at proximal edge, 0: 1S at widest part, 0-14 mm. 
long. 

The large avicularium was (by an ucbeialte not figured ; but the 
beak is about as large as an ordinary zocecium, and it is just the 
same in form as the avicularia of Cellaria ovicellosa, fig. 62. 

This would be Diporula of Hincks ; but as I am doubtful about 
the necessity for the genus, I at present at any rate do not separate 
it from Microporella. A reference to my paper ‘‘ On the Use of the 
Opercula in the Determination of the Cheilostomatous Bryozoa,” Proce. 
Manch. Lit. and Phil. Soc. vol. xvi. no. 2, pl. i. fig. 28, will show 
that, in the form of the operculum, Microporella (Diporula) verrucosa, 
Peach, is very similar to the present species. 

Loc. Fossil: Mount Gambier (Lond. Geel. Soc. coll. 1 


33. MicroporELLA ELEVATA, T. Woods. Plate XVIL.: figs. 63, 64; 
Plate XVIII. fig. 90. 


Eschara elevata, T. Woods, “ On some Tert. Austr. Bry.,” Tr. Roy. 
Soc. N. 8. Wales, 1876, De Bo tates, IND). 

Zoarium in Eschara- form, foliaceous, compressed. Zocecia, in 
young cells (fig. 90), widely oblong to square, with a large short 
triangular avicularium pointing inwards on one side of the zoce- 
cium on a level with the aperture, or frequently one such ayicula- 
rium on each side; in the old cells the zocecia are indistinct, with a 
much raised peristome giving a crateriform appearance ; the general 
surface and the peristome divided by raised lines into angular areas, 
with one or two pores in each such area ; below the aperture a large 
pore, which, however, is not always readily distinguished on the front 
of the zocecium ; but when examined from the interior (fig. 64) the 


ee 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 331 


round pore is always seen. Aperture of peristome round; oral aper- 
ture, as seen from inside, apparently with a straight proximal edge. 
The right-hand side of fig. 63 is the lower part of the zoarium, 
being placed sideways to show the structure better. Itis difficult to 
decide if this should be placed with Microporella or Porina. 
Loc. Fossil: Mount Gambier (Lond. Geol. Soc. coll.). 


34, MicRoPORELLA YARRAENSIS, sp. nov. Plate XV. figs. 27, 28. 


Zoarium in Hschara-form, subcylindrical, compressed ; branches 
dividing dichotomously. Zocecium distinctly bordered, irregularly 
oval; peristome slightly raised. Oral aperture round on the distal 
edge, straight on the proximal; two large pores immediately below 
the aperture; the upper part of the zocecium, supporting the aper- 
ture, together with these pores, is raised; a large protuberance on 
one side of the zocecium, and a smaller one on the other. In the 
central portion of the cell a large depressed area with two or three 
openings ; below this area a small avicularium. The zocecium is sur- 
rounded with an indistinct row of pores. Three distal rosette-plates 
(see fig. 28). Oral aperture 9-6 mm. wide. 

When the front of the ceil is slightly worn and the aperture 
broken down, the aperture and pores together frequently give a 
subtriangular form to the resulting opening. 

The details could not have been deciphered from the Mount- 
Gambier specimen, though when compared with the better-preserved 
specimens marked Yarra-Yarra it is seen to correspond in every 
particular. 

Loc. Mount Gambier (Lond. Geol. Soc. coll.). 


35. MicROPORELLA coscINopora, Rss., var. ARMATA, nov. Plate XV. 
fig. 25. 

This has a large distinct depressed cribriform area, and an avicu- 
larium below this area; the protuberances on the side of the zocecia 
are wanting; and the surface of the zocecium is flat; the upper part 
of the oral area is raised. Aperture 0-1 mm. wide. 

In one specimen in Miss Jelly’s collection there are similar small 
avicularia scattered irregularly over the surface. 


36. MicRoPORELLA ZNIGMATICA, sp. nov. Plate XV. figs. 29, 30. 


Zoarium in Eschara-form, composed of two layers back to back, 
forming a very thin lamella, which apparently has considerable 
extension (as all the pieces are flat and show no signs of branch- 
ing). Surface of zocecium very flat, opening of peristome circular ; 
opening of oral aperture, as seen from the interior, arched above, 
straight below; depression below the aperture, closed by a cribri- 
form plate; large and small acute avicularia scattered abundantly 
over the surface. Aperture 0-1 mm. wide. 

The surface is very flat, and the openings of the oral aperture and 
avicularia being often of nearly the same size and irregularly placed, 
it is in some places impossible to decipher any structure. But in the 
best-developed and best-preserved parts the peristome is seen to be 
slightly raised; and at first it seemed that the form of the aperture 


333 24 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


indicated Lepraka; but the interior shows that the lower edge of 
the operculum was straight, and that it must be placed with Micro- 
porella. 


37. MicRopoRELLA SYMMETRICA, sp. nov. Plate XVIII. fig. 83. 


Zoarium in Hschara-form, foliaceous. Zoccia hexagonally oval, 
distinct, with a thin border round the cell, scarcely raised, with few 
pores near the edge, raised in the region of the aperture, especially 
above the aperture. Two large openings, directed laterally imme- 
diately before the aperture ; these are usually raised and form part 
of the peristome. An area on the lower part of the cell with stellate 
pores arranged in two series of 4 or 5, each regularly opposite to one 
another ; sometimes one or two pores between the lateral rows. This 
area usually forms a deep depression; but in some cells this is not 
the case. Oral aperture semicircular, with straight proximal edge, 
0:09 mm. wide, 0:07 mm. long. 

In the species we see a marked likeness to MW. yarraensis (fig. 27); 
and through these we are able to trace the relationship of this area 
from the round pore of M.violacea to the elongate pore of M. violacea, 
var. jissa (fig. 26), to the two or three pores of M/. yarraensis, and 
the cribriform pores of MW. coscinopora (fig. 25) and M. enigmatica 
(figs. 29, 30), and, lastly, to the depressed area of the present species ; 
and we seem justified in extending the comparison and comparing 
the non-depressed area of the present with that of Cribrillina 
terminata (fig. 68), which, in its turn, may be traced to typical 
Cribrillina. 


38. MicroporeLia ciavata, Stol. Plate XVIII. fig. 84. 


Flusirella clavata, Stol. Foss. Bry. Orak. p. 139, pl. xx. figs. 3, 5. 

Zoarium in Hschara-form, forming thin foliaceous dichotomous 
branches or small round branches. Zocecia suboval, contracted and 
truncate below; a characteristic semicircular depression immediately 
below the aperture, with three large raised pores, the two upper 
ones directed to the right and left. A laterally oval depression or 
opening about the middle of the zoccium, with the median pore 
~ below this. Oral aperture semicircular, with a straight proximal 
edge, 0:07 mm. wide, 0:05 mm. long. 

This is much the same as Eschara tetrastoma, Rss. Sitz. Ak. W. 
Wien. 1864, p. 9, pl. i. fig. 9. 

Loc. Fossil: Orakei Bay. 


39. Porina CLYPEATA, sp. nov. Plate XVII. fig. 67. 


Zoarium erect, cylindrical ; four zocecia in a complete series, al- 
ternating regularly with each other, and facing in four directions. 
Zoecia distinct, tubular; peristome much projecting; median pore 
placed about one half the length of the cell, opening at the top of a 
cone, the base of which is surrounded by an oval or circular rim, 
giving it ashield-like appearance. This rim is continued in a straight 
line above and below. A small obtuse avicularium on each side of 
the shield-like boss. 


a a oe 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA, S53) 


This is a very interesting specimen, as the shield-like elevation 
gives ita peculiar appearance different from any thing with which we 
are acquainted ; and at first I did not know in what way the struc- 
ture must be interpreted, and examined to see if it could be an ovi- 
cell, but think it must be considered homologous with the tubular 
pore of P. tubulosa. The specimens are all small. 

Loc. Fossil: Mount Gambier (Zth., jun., coll.). 


40. Portna coronata, Rss. Plate XV. fig. 57. 


Cellaria coronata, Rss. Foss. Polyp. des Wien. Tert. p. 62, pl. viil. 
fig. 3. 

Eschara conferta, Rss. loc. cit. p. 71, pl. vii. fig. 32. 

Acropora coronata, Rss. Foss. Anth. & Bry. d. S. von Crosaro ; 
Denk. math.-nat. Cl. k. Akad. der Wissensch. Wien, vol. xxix. 1868, 
p. 65, pl. xxxiv. figs. 33-55. 

Spiroporina vertebralis, Stoliczka, Foss. Bry. Orak. p. 106, pl. 
xvii. figs. 6, 7. 

Spiroporina veriebralis, T. Wocds, ‘‘ Corals & Bryozoa of Neozoic 
Period in New Zealand,” Pal. of New Zeal. pt. iv. Colon. Mus. & 
Geol. Survey Dept. 1880, p. 23. 

Porina Dieffenbachiana, Stoliczka, loc. cit. p. 135, pl. xix. fig. 20. 

Porina Dieffenbachiana, 'T. Woods, “C. & Bry. of Neoz.” loc. cit. 
p. 25. 

Eschara Buskii, T. Woods, ‘On some Tert. Austr. Polyz.,” Tr. 
R. Soc. N.S. Wales, 1876, p. 149, figs. xvi. & xvii. 

Myriozoum australiense, Haswell, “On some Polyzoa from the 
Queenland Coast,” Proc. Linn. Soc. N. 8S. Wales, vol. v. pt. 1, 1880, 
p. 43, pl. iii. figs. 9-11. 

This species has caused me much trouble, as its appearance in 
various conditions is very different ; but direct comparison of typical 
specimens with Acropora coronata from Val di Lonte showed that 
the size, form of the cell, peristome, median pore, and small pore 
were sometimes just the same. 

The zoarium is sometimes round and very slender, asin fig. 57; in 
others round and stout, as in Stoliczka’s figure of S. vertebralis ; but 
having a complete intermediate series, there is no doubt that they 
must be united. In other specimens, especially those from Mount 
Gambier, the branches are flattened as in P. Dueffenbachiana, 
Stol., fig. 20. The surface sometimes has small elongate pores, 
in other cases larger and round ones. In theslender specimens the 
pores are finer and more elongate, while in the stoutest they corre- 
spond with the pores of the Italian P. coronata. Again, the 
peristome in young cells is very thin and projects considerably ; 
while in older specimens the large pores in the walls of the peri- 
stome are more distinctly seen; and in very well preserved speci- 
mens the walls of these project, so that each peristomial pore may 
be said to have itself a peristome. The median pore is usually about 
one quarter of the length of the zocecium from the oral aperture, but 
sometimes much nearer ; and in old cells, where the peristomial wall 
is much thickened, it approaches very close to the peristome, and 


oot A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


almost becomes part of the peristomial crown. Ihave only one or 
two specimens in which there is a slightly projecting tube for this 
median pore ; and in all other cases it is depressed. In some spe- 
cimens, besides the small surface-pores, there are larger ones, which 
have in most cases been avicularia; and in a few specimens there is 
a larger spatulate avicularium above the oral aperture. 

In a recent specimen in the vertebralis condition, from Darnley 
Island, I found that a section made from one end showed the central 
spongy structure with which we are acquainted in the old genus 
Myriozoum, while a section made from the other showed the cells 
ranged round the circumference but not meeting in the centre: 
and a section is thus figured by Mr. Haswell (doe. cit. fig. 10). 

Stoliczka, in his figure, distinctly showed this Myriozoum character ; 
and it is strange to find that he nevertheless placed the species 
among the Cyclostomata. The slender forms so nearly correspond 
with Porina borealis, Busk (especially as figured by Smitt in his 
‘Floridan Bryozoa,’ pl. vi. fig. 144), that, if not the same, they are 
closely related to it; and the species is also related to Porima 
filograna, Goldf., figured in d’Orb. Pal. Frang. pl. 626. figs. 5-10, a 
common species in the Senonian, and also figured by Hagenow as 
schara Defrancei. Several of Hagenow’s species with other names 
may represent worn specimens, as pointed out by d’Orbigny. 

Although the different conditions pass so gradually from one to 
another that it is impossible to separate them, it may perhaps be 
useful to speak of them as condition (a) slender, as fig. 57 ; (6) stout, 
as vertebrals, figured by Stol.; (c) more flattened, and showing the 
quincuncial arrangement more clearly, as in Dieffenbachiana. 

Loc. Fossil: Bartonian, Val di Lonte (Ztss.), Montecchio Mag- 
giore (ftss.), Vienna (fss.); Ferrara di Monte Baldo, Crosaro, Bren- 
dola (all in my coll.); Hutchinson’s Quarry and Oamaru, New Zea- 
land, in Lower Eocene of New-Zealand geologists (as ¢, Woods); 
Hutchinson’s Quarry, Upper Eocene ditto ; Shakespeare Cliff, Upper 
Miocene ditto (as 6); Orakei Bay, New Zealand (6 & ¢, Siol.); 
Mount Gambier (Woods, 6 & c); and common in the Lond. Geol. 
Soc. coll., and also Eth., junr., coll. from Mount Gambier (as 6). 

Living: as 6, Holborn Island, Queensland, 20 fathoms (Hasw.), 
and Darnley Island, Torres Strait, in material sent me by Mr. Brazier, 
from soundings in 10-30 fathoms. 


4]. Pormya? cotumnata, sp. nov. Plate XVIII. fig. 88. 


Zoarium cylindrical, erect, with six cells in a complete series, con- 


sisting of two rows of three each. Zocecia irregularly oblong, with 
nearly parallel sides, distinct, separated by a wide rounded ridge, 
surface rounded, with much-raised acute granulations ; the peristome 
forms a closed arch over the lower part of the oral aperture ; the 
opening at the base of this arch is wide and laterally oval; a large 
tubular avicularium on each side of the peristome, also raised and 
_ continuous with the peristome. Oral aperture rounded below, pro- 
bably round to saddle-shaped, somewhat contracted above. ' 
The specimen is in a bad state of preservation; and diagnosis 


a 
a_i 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 335 


has to be made from the characters in various cells ; consequently 
the figure is, to a large extent, a restoration, which is not the case 
with the other figures. I have shown the effect when the aperture 
is broken down in different degrees: when completely worn away, 
three nearly equal openings are seen, the central one being some- 
what the largest ; in other cases an elongate opening is formed, 
bounded on the top by the distal end of the peristome, and below by 
the base of the subperistomial opening. 

This is closely related to Eschara heterostoma, Reuss (Bry. von 
Crosaro, p. =°7,, pl. xxvi. fig. 5), which we believe is synonymous 
with his #. duplicata (loc. cit. p. (61 sep.) 273, pl. xxxiii. figs. 8, 
10), from the Italian Eocene. 

The shape of the oral aperture would seem to be Lepralian, perhaps 
much the same as in Lepralia Pallasiana ; and itis somewhat a matter 
of doubt where this form should be generically placed. There are 
some species as yet undescribed from the southern hemisphere, with 
an arch more or less enclosing the aperture; and until these have 
been examined, it must be placed with Porina. 

Since the above was written, Mr. 8. O. Ridley has published, in 
the Proc. Zool. Soc. 1881, an “ Account of the Zoological Collection 
made during the Survey of H.M.S. ‘Alert’ (pt. Polyzoa),” and de- 
scribes and figures an incrusting species with cells much. resembling 
those of P. columnata. For this he forms the genus Gigantopora, 
and points out the similarity to Hippothoa fenestrata, Sm., which 
seems to sometimes occur in an erect form; and these three spe- 
cies are evidently closely related ; but we provisionally retain the pre- 
sent one under Porina until more complete specimens furnish further 
characters upon which the genus can be founded. 


42. LEPRALIA CORRUGATA, sp.nov. Plate XVII. fig. 60. 


Zocecia elongate, distinct, raised ; surface in irregular ridges and 
furrows; in the furrows a row of one or two pores; lower part of 
zocecia depressed, upper raised, with a large angular avicularium 
forming a kind of peristome over the lower part of the aperture ; 
one or two large pores at the lower part of the zocecium, just above 
the aperture of the zocecium next below. Aperture rounded. Zoa- 
rium either incrusting or in Hschara-form. 

Oral aperture 0°14 mm. wide. 


43, LEPRALIA MONILIFERA, M.Ed., var. arMATA. Plate XV. fig. 24. 


Zocecia elongate, pyriform, raised in the centre, with a row of 
pores round the border ; aperture oval, proximal edge slightly less 
rounded than the distal; a small angular avicularium on one or 
both sides of the aperture; in some cells larger angular avicularia 
below the aperture. 

Aperture 0-11 mm. wide; 0:14 mm. long. 

Only a small fragment, worn at the back. 


44, LEPRALIA SPATULATA, sp.nov. Plate XVIII. fig. 87. 
Zoarium erect, forming a solid stem, with zocecia all round. 


336 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


Zoecia hexagonal, distinct, with a thin border slightly rounded, 
raised towards the aperture; surface finely granular; the region 
above the aperture is much raised, with six spines, three on each 
side. Aperture nearly circular, slightly elongate, situated in a wide 
depression, caused by the elevation round the sides and distal part 
of the aperture. Aperture 0°15 mm. wide, 0°17 long. 

This much resembles Lepraha multispmata, Busk, from Madeira 
(Q. J. Micr. Sc. new ser. vol. i. 1861, p. 78, pl. xxxii. fig. 5); but 
the lip below the aperture is entirely wanting. 

In a specimen from the Mount-Gambier collection of Mr. Ethe- 
ridge, jun., there are many large spatulate zocecial avicularia scat- 
tered over the stem. 

Loc. Fossil: Mount Gambier. 


45, LEPRALIA CLEIDosToMA, Sm., var. roTuNDA. Plate XVIIL. fig. 92. 


Zoarium apparently flat and incrusting. Zocecia small, distinct, 
hexagonal ; surface coarsely granular. Oral aperture orbicular, con- 
tracted at the sides by two denticles; raised rounded avicularium 
(perhaps with vibracular characters) below the line of the aperture, 
on the side of the zocecium ; four lateral rosette-plates, one distal near 
the base of the zocecia. Oral aperture 0°1 mm. wide, 0°13 long. 

It is difficult to know where this should generically be placed. 
From the rounded distal and proximal edges of the aperture, it would 
have gone with Smitt’s earlier definition of Escharella. In the size 
and form of the cells it much resembles Schizoporella excubans, fig. 56; 
but this, instead of being an erect quadrilateral form, is incrusting, 
and has a rounded avicularium ; but we may perhaps here trace the 
passage from a definite sinus to a rounded proximal edge. It only 
differs from Lepralia cleidostoma, Smitt (Flor. Bry. p. 62, pl. xi. 
figs. 217-219), in having the rounded avicularium instead of an acute 
one, and should, perhaps, be united to it. 

I have an erect branching form, with ovicells, from the Mediter- 
ranean, Closely allied to this, which I hope shortly to describe and 
figure. 

Mr. Hincks has kindly informed me (in htt. ee that he has this va- 
riety from Bass’s Straits. 


46. PoRELLA EMENDATA, sp. nov. Plate XVII. fig. 69. 

Zoarium in Eschara-form. 

Surface nearly flat; cell-wall thickened round the oral aperture. 
Zocecia elongate, expanded above, contracted below; row of pores 
in a furrow on each side of the zoccium. Aperture round on the 
distal edge, straight on the proximal, with a very large denticle 
set deeply in the aperture. Small triangular avicularium below 
the mouth, usually placed somewhat diagonally. 

Aperture on proximal edge 0:07 millim. wide, 0-06 millim. long. 


47. PoRELLA DENTICULATA, Stol. Plate XVII. fig. 70. 


_Plustrella denticulata, Stoliczka, Foss. Bry. Orak. p. 138, pl. xx. 
fig. 2. 
Oral aperture arched above, contracted below, enclosing an ayicu- 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 307 


larium surmounted by a denticle ; faint lines on the surface indicate 
the zocecial divisions. Ovicell subimmersed, without any pores. 
Loc. Not very common in Orakei Bay. 


48. SMITTIA CENTRALIS, sp. nov. 


Zoarium slender, erect, quadrangular, with the zocecia on all four 
faces. Zocecia separated by a ridge, elongate, oblong, flat, with a 
double row of pores on both sides of the zocecia near the edge, 
with the peristome very much raised and projecting, with a large 
pore (probably avicularian) immediately below the or ifice ; SER DET 
transversely oval. 


49. SMITTIA CENTRALIS, var. LHVIGATA. Plate XIV. figs. 7, 8. 


Differs from the above in haying the surface of the zocecia slightly 
rounded ; and instead of the two rows of small pores, there are 
a few elongate pores near the edge of the zoccia; the peristome is 
less well preserved ; and from these small fragments it would be 
impossible to be sure of the generic position of the species. The 
distal rosette-plates (fig. 8) are near the centre of the zoarium, from 
which circumstance the name is chosen. 


50. Smirrra Tarrr, T. Woods. Plate XVII. fig. 65 


Eschara Tatei, T. Woods, “ On some Tert. Austr. Fossils,” Tr. R. 
Soc. N.S. W. 1876, p. 3, fig. xv. 

Zoccia elongate, with parallel sides, contracted below, surrounded 
by a narrow raised margin ; two perforated furrows down the front. 
Aperture raised, rounded, with a second rounded opening imme- 
diately below (this is probably avicularian). Seen from the inte- 
rior, there is a large denticle expanded at the top (fig. 65 a). 

Aperture 0°13 millim. wide. 

In consequence of the imperfect state of fossilization of the Mount- 
Gambier fossils, the aperture presents the appearances figured by 
Mr. Woods. A specimen in the collection of Mr. Etheridge, jun., 
shows cells as figured by Mr. Woods, and resembling those from the 
present collection. 

Loc. Fossil: Mount Gambier (Woods and collection of Mr. Ethe- 
ridge, jun.). 


51. Surrrra anceps, MacG. Plate XVIII. fig. 94. 


Lepralia anceps, MacGillivray, Prodr. of Zool. of Vict. decade iv. 
e7sp. 20, pl. xxxv. fig. 6. 

Zoarium cylindrical; in one specimen four or five cells in a series, 
in another a hollow cylinder 2-5 millim. in diameter. Zocecia sub- 
rhomboidal, sometimes hollowed in front, elevated towards the ori- 
fice, bounded by a prominent irregular sinuous line; surface very 
finely granulated, with large pores, more numerous near the edge. 
Oral aperture rounded on the distal end; the proximal edge is 
formed by the arc of a smaller circle. 

When the aperture is seen from inside, we find a broad denticle 
(fig. 94a); and when we look upon this denticle from above, it is 


Pate) A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


seen to bear avery minute avicularium. Aperture 0-1 millim. wide, 
0°14 millim. long. | 

‘If I had been unable to see the interior of the zocecium, I should 
have considered this to be Schizoporella sinwosa, B., with which it 
corresponds in most particulars. I have examined the interior of 
S. stnuosa from Shetland, in which there is no similar denticle or 
avicularium. Some of the zocecial areas have no aperture; and m 
' these there are few or no pores. 


52. ScHIZOPORELLA VIGILANS, sp. nov. Plate XIV. fig. 13. 


Zoarium erect, quadrilateral, with zocecia on the four faces. Zooscia 
irregularly oblong; surface flat, with elongated pores (reminding us 
of those of Hornera); upper part of the zocecium slightly raised. 
Oral aperture rounded on the distal edge, nearly straight on the 
proximal, with a well-marked sinus. Acute avicularia, placed hori- 
zontally in the middle of the front wall of the zocecium. 

The proximal edge of the oral aperture is slightly straighter than 
figured. Aperture 0:08 millim. wide, 0:07 millim. long. 


53. SCHIZOPORELLA PHYMATOPORA, Rss. Plate XV. figs. 31, 32. 


Eschara phymatopora, Reuss. ‘ Foss. Anth. u. Bry. von Crosaro,” 
Denschr. Ak. Naturwissensch. Wien, vol. xxix. 1869, p. 272 (60), 
jl soroxibbly 11k 1p | 

Zoarium cylindrical. Zocecia irregularly oval, not much raised ; 
border distinct. Aperture circular, with sinus below. Avicularia 
at about one third or the middle of the length of the zocecium, placed 
on one side; surface of zocecium covered with pores and fine gra- 
nulations. 

When I drew fig. 32, I supposed that this was Bactridiuwm, and 
allied to Bactridium Hagenowi, Rss.; and then I called it tr- 
serrata, aS in all the fragments I had there were three rows of 
cells. Since then fragments in my own and in Miss Jelly’s 
collection have shown that it is cylindrical. The species splits 
up with considerable regularity ; and then the other sides of the 
zocecia are seen with their lateral rosette-plates appearing like a row 
of pores. A central furrow is also seen with two rosette-plates for 
the entrance of the endosarc. The avicularia are usually much 
lower than figured by Reuss in his specimen from the Val di Lonte, 
but in some Australian specimens are higher than in my figure 31. 
Aperture 0:07 millim. wide. 

Loc. Val di Lonte (#ss.); Ferrara di Monte Baldo (A. W. W.): 
Bartonian. 


54, ScuizopoRELLA VENTRICOsA ?, Haswell (in Onchopore forma). 


Onchopora ventricosa, Haswell, “On some Polyzoa from the 
Queensland coast,” Proc. Linn. Soc. N. S. Wales, vol. v. pt. i. 1880, 
p. 36, pl. 1. fig. 3. ; 

There is only a small fragment, which has the zoarium cylindrical, 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 309 


and which seems to branch without joints. It seems to correspond 
with the above ; but there is reason for doubting whether too many 
species have not been made, as Onchopora ventricosa, Hasw., and 0. 
granulosa, H., are very closely allied. 

Loc. Holborn Island, Queensland, 20 fathoms. 


55. ScHIZOPORELLA FENESTRATA, sp. nov. 


Zoarium in Kschara-form. Zoccia indistinct, surface smooth. 
Oral aperture very large, in a very deep depression, rounded on the 
distal edge, with a sinus on the proximal; a small rounded avicularium 
within the oral depression, and a small rounded one below the aper- 
ture with avicularium opening with numerous denticles. Very large, 
erect, angular avicularium between the zoccia. Oral aperture 0°17 
millim. wide. 

The large deep depressions round ie large aperture, together with 
the smooth surface and small avicularia, gives the zoarium the ap- 
pearance of the network of Retepora; and the specimen was sent to 
me marked Retepora. I therefore call it fenestrata, although, of 
course, it is clear that itis here the magnified aperture which repre- 
sents in appearance the unmagnified fenestra of Retepora. 


56. ScHIZOPORELLA, sp. 


Zocecia perfectly parallel, oblong, distinctly divided laterally, ter- 
minal divisions scarcely distinguishable. Surface smooth. Oral 
aperture large, 0°3 millim. long, 0°22 millim. wide, rounded above, 
apparently triangular below ; frequently a notch on one side, as 
if regularly broken down; perhaps there is an avicularium inside 
the aperture below the notch. ‘The back of the cells is also smooth, 
showing the lateral parallel divisions distinctly, with an oval open- 
ing 0:4 millim. long, 2. ¢. about half the length of each zocecium. 

There are one or two species in which there is a similar oval 
opening on the dorsal surface; but at the»present moment I do not 
find any note or reference to such, except in Hemeschara gemi- 
nupora, Rss. 


57, SCHIZOPORELLA, SP. 


This somewhat resembles S. biaperta; but the state of preserva- 
tion is not sufficiently satisfactory for definite determination. The 
aperture is much smaller than in the last species; and there are 
numerous round avicularia scattered over the cell. The walls of 
the ovicells, which have been considerably raised, are broken down. 
This may be Hscharipora Lawderiana, Stol. (Bry. Orak. Bay, p. 136, 
pl. xx. fig. 1); but the figure and description are so unsatisfac- 
tory that it is impossible to know where Stoliczka’s specimen 
should be placed. The fossil specimen is in the Aschara-form. 

A specimen in the British Museum, sent by Mr. Hutton, marked 
Lepralia reticulata, seems to be this species. The oral aperture of 
the recent specimen is rounded on the distal end, with a smaller arc 
forming a sinus on the proximal. | 


340 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


58. ScHIzoPoRELLA SUBMERSA, sp. nov. Plate XVIII. fig. 85. 


Zoarium in Hschara-form, consisting of narrow foliations. Zo- 
cecia suboval, expanded above, contracted below, distinct, sur- 
rounded by a raised border, partially concave, few pores on the sur- 
face. Acute avicularium near the centre of the zocecium, sometimes 
directed diagonally upwards, sometimes downwards. Oral aper- 
ture very long, rounded on the distal end, with a small sinus on 
the proximal. ‘The aperture is much depressed, giving the ap- 
pearance of two wide denticles at the bottom of the oral pit. Aper- 
ture 0-07 millim. wide, 0°12 millim. long (to end of the sinus). 

The upper part of the zoceclum being depressed gives a concave 
appearance to the zocecia. 


59. ScHIZOPORELLA CONSERVATA, sp. nov. Plate XVIII. fig. 81. 


Zocecia suboval to oblong, distinct, not much raised, sometimes 
with a median raised ridge extending from the sinus; a few large 
pores near the edge; surface smooth; three spines above the oral 
aperture, and one on each side at the base of the aperture; surface 
slightly raised below the sinus. Oral aperture semicircular, rounded 
on the distal edge, straight on the proximal, with a small sinus 
which widens out below. An avicularium on one or both sides a 
short distance below the oral aperture; these vary from short 
triangular to acute lanceolate, pointing laterally outwards at right 
angles to the axis of the cell. Ovicells very large, elevated, with 
the central part plain, and nearly flat, the exterior walls beautifully 
ornamented with radiating lines with three pores between each. 
Aperture 0°16 millim. wide, 0°12' millim. long (to the straight 
proximal edge). | 

When the avicularia are small, they are raised ; but when large, 
they are immersed. The dorsal surface is hexagonally divided, with 
an elongate oval space (about half the length of each zocecium) of 
thinner shell. This perhaps corresponds with the “ flattened disk ” 
mentioned by Mr. Busk in Microporella Malusw, Aud. (Mar. Poly. 
p. 83); but it is by no means a constant character in Malusi, as 
the centre of the dorsal surface is usually depressed, with irre- 
gular elevations near the edge of the zoccium. The structure of 
the dorsal surface of Schizoporella conservata must probably be com- 
pared with that of Schizoporella, sp., No. 56. I presume this spe- 
cies grew in the Lepralia-form. 

Loc. Fossil: Mount Gambier (Lond. Geol. Soc. coll. and Eth., 
Jun., coll.). 


60. ScHIZOPORELLA SPIROPORINA, Sp. NOv. 


Zoarium cylindrical ; zocecia in annular series of six cells; series 
0-7 millim. apart. Zocecia indistinct, except towards the peristome ; 
_ peristome much raised in front, but very little behind. Oral aper- 
ture at the base of the peristome, nearly circular, rounded at the 
distal end, with a widesinus at the proximal. Large pores, perhaps 
avicularian, near the peristome. 

This may be Spiroporina immersa of T. Woods (Corals & Bry. Col. 


BRYOZOA FROM §.W. VICTORIA, AUSTRALIA. 541 


Mus. & Geol. Survey New Zealand, pt. iv. p. 23); but I must con- 
fess to being unable to understand Mr. Woods’s description of the 
mouth. The specimen from ‘ Yarra-Yarra” is very small and in 
imperfect preservation. 


61. ScHIZOPORELLA EXCUBANS, sp. nov. Plate XVI. fig. 56; Plate 
XVIII. fig. 80. 


Zearium erect, filiform, with a longitudinal row of zocecia on each 
of the four faces. Zocecia distinct, subhexagonal, upper part much 
raised, surface granular. Oral aperture depressed, rounded on the 
distal] edge, contracted on each side near the proximal edge, forming 
a large rounded proximal sinus. Triangular avicularium on raised 
prominence on a level with the oral aperture, directed downwards. 
Avicularia sometimes absent, usually on one side only. 

Aperture 0-08 millim. wide, 0-075 millim. long. 

Probably the genus Schizoporella will have to be broken up, and 
those forms in which the proximal edge of the oral aperture forms a 
large arc will be separated; for while the appearance is that of a 
large sinus, the structure must be different. It is also very diffi- 
eult to see where the division should be made between Lepralia and 
Schizoporella ; for the wide rounded aperture of Lepralia with two 
lateral denticles sometimes approaches very closely in shape that of 
a Schizoporella with a large and wide sinus. Fig. 56 was drawn 
from a small fragment in which it was impossible to see the form 
of the zoarium, and therefore, in this respect, is not quite correct. 
I have since received specimens showing the erect filiform growth. 


62. ScHIzoPORELLA AMPHORA, Sp. Nov. 


Zoarium slender, erect, with a single row of zocecia on each of the 
four faces, the opposite pairs alternating with the othertwo. Zocecia 
irregularly ovate, expanded _and raised in the middle, contracted above 
and below ; distal end of the zocecium much raised, forming a small 
peristome which is contracted on both sides near the lower part. 

The specimen is small; and more perfect ones may add many 
particulars. It may have been sometimes articulated, as seems to 
have been the case with Cellaria Schreibersi, Rss. 


63. ScHIZOPORELLA AUSTRALIS, T. Woods. Plate XIV. fig. 15. 


Tetraplaria australis, T. Woods, “On some Austr. Tert. Foss. 
Corals & Polyzoa,” Tr. Roy. Soc. of New 8. Wales, 1878, p. 5, fig. 4. 

Zoarium cylindrical, slender. Zocecia facing four ways, the op- 
posite pairs alternating with the other two, elongately pyriform ; 
surface very finely granulated, with extremely fine pores between 
the granules (but these can only be seen when the preservation is 
very good). Oral aperture rounded on the distal edge, slightly curved 
or straight on the proximal, with a small distinct sinus; aperture 
0-08 millim. wide, 0°07 millim. long. 

Since I drew the figure I have received a much better specimen, 
from which J have been able to make out the character of the 
aperture. The cells are nearly half as long again as in Cellaria 


Q.J.G.8. No. 147. 2A 


342 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


Schreibersi, Rss., with which Mr, Woods compares it; and, further, 
the two prominent avicularia are wanting; but from my Val-di- 
Lonte specimens of C. Schretbersi I find that both belong to the Schi- 
zoporellidse, and are allied in many particulars. 

Loc. Muddy creek, Western Victoria ( W.). 


64. Rerepora marsupraTa, Smitt. Plate XV. figs. 34, 35, 36; 

Plate XVII. figs. 59, 61, 76, 77. 

Retepora marsuprata, Sm. Floridan Bry. p. 67, pl. xiii. figs. 245- 
254; Svenska Vetensk. Handlingar, vol. xi. 1872. 

Phidolophora labiata, Gabb & Horn, ‘“* Polyzoa of Second. and Tert. 
Form. of N. Amer.” p. 138, pl. xix. fig. 21°(Journ. Ac. Nat. Sei. 
Philad. vol. v. pt. i1.). 

Zoarium reticulate. Zocecia suboval to hexagonal, separated by 
a distinct raised border; peristome elevated in young cells; in old 
ones the aperture is immersed, with sometimes a spine on each side 
of the aperture; in front of the peristome a ridge, at the base of 
which is sometimes a small pore (sometimes avicularian); this gives 
a distinctly sinuated appearance to the peristome: in some speci- 
mens large, erect, angular avicularia in the middle of the zocecium ; 
in others large erect more spathulate avicularia; in others very long 
and very narrow immersed ayicularia in a similar position, with the 
mandible in all cases pointing directly or diagonally downwards ; 
sometimes there are small rounded avicularia pointing downwards, 
with a central pore. Surface smooth, or with few granulations ; often 
two large pores near the proximal extremity of the zoccia. One 
ovicell is subimmersed, with one cleft, as described by Smitt ; ano- 
ther is more raised and more globose, with two depressed lines close 
together instead of the one central cleft; and there is also a small 
boss on the centre of the ovicell. The dorsal surface, which is 
divided irregularly by thin raised lines, has minute rounded avicu- 
laria scattered about; there are also long, triangular, immersed 
avicularia and large erect ones. In the basal portion the aperture 
is depressed, when we find the structure figured in figs. 59, 61, 
and 76, which are drawn from different specimens ; but interme- 
diate ones leave little doubt as to their identity. 

In these we see the median pore representing avicularia, and 
showing the relationship to Portna. Median pores are known in 
Retcpora tuberculata, Rss. Some of the basal branches seem to show 
that reticulation was much less frequent than in most Retepore ; and 
this and &. rimata would be Psilescharew of Busk, a genus which 
cannot be retained. ’ : 

It is impossible to be quite sure if this is the same as the fossil 
described by Gabb and Horn. It is evidently allied to R. Beanzana, 
King, and probably is the species described under that name by 
Stoliczka. The suboral pore represents the suboral avicularium in &. 
Beaniana; and small avicularia are found on the front of Beanana. 

Loc. Living: Floridan seas, 16-262 fathoms (Sm.); Teneriffe 
(Busk). Fossil: Miocene, 8. Barbara, Amer. (G. & H.); Mount 
Gambier (Lond. Geol. Soc. coll.). 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 343 


65. Rerepora rmmata, sp. nov. Plate XVI. figs. 48, 53. 


Zoarium probably reticulate; branches slightly compressed. 
Zocecia cylindrical, convex ; surface smooth ; peristome much raised, 
with a cleft in front the whole length of the peristome, expand- 
ing at the base, the edges on the two sides of the cleft turned 
inwards, peristome often surrounded in front with a raised rim. 
Small subimmersed avicularia below the peristome, subtriangular 
or suboval, pointing upwards, with a very minute opening near the 
distal end. Dorsal surface divided by white raised lines, with small, 
subcircular, acute avicularia and small bosses irregularly placed. 

This is common from Mount Gambier; but the state of fossi- 
lization would not have permitted all the characters to be made 
out if the “ Yarra- Yarra” specimens had not served as a key. 

Loc. Fossil: Mt. Gambier (Lond. Geol. Soc. and Hth., jun., coll.). 

In one specimen the avicularia in the front are replaced by asimple 
pore. This would unite the genus with Porinide; and we here see 
another example of the avicularia taking the position of pores, and 
are led to regard avicularia as modified pores. 


66. CELLEPORA YARRAENSIS, Sp. NOV. 


Zoarium cylindrical, dividing dichotomously, in one case throwing 
out a branch forming an angle of about 80°; diameter of stem 1-2 
millim. Zocecia very irregularly arranged, placed sometimes very . 
crowded, sometimes far apart, indistinct ; peristome much raised on 
the lower or avicularium side, usually not at all on the other, giving 
it, when seen in profile, a triangular appearance; at the proximal 
part of the peristome an avicularium ; when this is broken down the 
two large openings have the appearance so general in Cellepore ; 
between the zocecia few large much raised perforated protuberances, 
probably sometimes avicularian. Surface smooth, porcellaneous. 
Oral aperture semicircular, rounded on the distal edge, straight on the 
proximal, though when raised cells are seen the aperture appears 
rounded. Aperture 0-08 millim. wide, 0:1 millim. long. 

I have an undescribed thin branching species of Cellepora from 
Capri (near Naples) which also has the lower edge of the aperture 
straight and in many respects resembles the present. 


67. Ce~tepora Fossa, Hasw. Plate XVIII. fig. 89. 


Spheropora fossa, Haswell, “ On some Polyzoa from the Queens- 
land Coast,” Proc. Linn. Soc. New 8. Wales, vol. v. pt. i. 1880, p. 
42, pl. ii. fig. 5, 6. 

Zoarium subglobular, flattened above, specimens 2—5 millim. in 
diameter. On the upperside the zocecia are directed from the pole of 
the zoarium ; on the underside they are directed towards it. Zocecia 
subcylindrical or ovate or conical, very finely granular, erect (on 
the upperside) ; a few pores round the edge of the zoecium on the 
' upperside, but seldom on the back. Oral aperture rounded on the 


- distal wall, straight on the proximal. In the centre, immediately 


below the proximal margin, a large, wide, erect, obtusely conical or 
globular avicularium. Few large, spatulate, zocecial avicularia 
Za 2 


o44 - A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


scattered over the colony. Aperture 0:12 millim. wide, 0:1 millim. 
long. 

I cannot attach any importance to the pit round which the zocecia 
are formed and upon which Mr. Haswell’s new genus is founded. 
The form of the oral aperture is rare in the Celleporide ; but we see 
the same thing in C. sardonica, Waters (“ Bry. Bay of Naples,” Ann. 
Nat. Hist. 1879, p. 196), also in C. yarraensis, W., in C. intermedra, 
M‘G., and in C. compressa, Busk, with which perhaps the above 
should be united; and perhaps these together should form a sub- 
genus. This may be one of the Cellepore described by Mr. Tenison 
Woods in “ Tertiary Fossils in S. Australia,” p. 5 (Trans. Roy. Soe. 
of Victoria, vol. vi.); but identification with these is impossible, as 
the zoarial instead of the zocecial characters are described. In de- 
scribing fossil Bryozoa so little attention has been paid to the most 
important character, 2.¢. the form of the oral aperture, that com- 
parison is often not possible. 

Loe. Living: Holborn Island, Queensland, 20 fathoms. Fossil : 
Mount Gambier (Lond. Geol. Soc. and Hth., jun., coll.). 


68. CELLEPORA, sp. 


There are a few cells of a Cellepora which resemble those of 
C. pumicosa, Busk (non Linn.); but the fragment is too small to 
determine with certainty. 


69. LUNULITES GUINEENSIS, Busk. . 


Cupularia guineensis, Busk, Cat. Mar. Polyz. p. 98, pl. exiv. 

The specimen in Miss Jelly’s collection is very small; but as the 
few zocecia correspond in shape with those figured by Mr. Busk, 
we may safely conclude that the same species is ; represented. 

Loc. New Guinea (B). . 


70. LUNULITES CANCELLATA, Busk. 


Lunulites cancellata, Busk, Cat. Mar. Pol. p. 101, pl. exiii. figs. 4-7. 

Loc. Philippine Islands (Busk); off Raton, New Guinea, 7 
fathoms, and from Darnley Islands, Torres Straits, 10-30 fathoms 
(sent me by Mr. Brazier). 


71. Sevenarza MaRGINATA, T. Woods. Plate XVII. fig. 71. 


Selenaria marginata, T. Woods, “On some recent and fossil 
Species of Australian Selenariade (Polyzoa),” Trans. Phil. Soc. 
Adelaide, 1880, p. 9, pl. i1. fig. 9, a—d. 

The zocecial cells are very small, and in shape resemble those 
drawn by Mr. Woods. In some of the zocecia there is a plate, some 
little distance down the aperture, with a central perforation, so that 
the zocecium is almost closed. Sometimes the same thing is seen in 
Cellaria fistulosa both recent and fossil. I hardly understand 
Mr. Woods’s description or plate when he refers to the pore; but I 
think this is undoubtedly the species he describes. Aperture 0-06 
millim. wide. 

Loe. Living: Cape Three Points, 71 fathoms. 


BRYOZOA FROM S.W. VICTORIA, AUSTRALIA. 345 


72. Sermnapgia ALATA, T. Woods. 


Selenaria alata, Tenison Woods, “On some recent and fossil 
Species of Australian Selenariade (Polyzoa),” Trans. Phil. Soc. 
Adelaide, 1880, p. 11, pl. u1. fig. 12, a, 0, ©. 

Loc. Muddy Creek, Miocene Beds near Hamilton, Victoria (W.). 

The specimen is smaller than those described by Mr. Woods, 
only measuring 23 millim.; but otherwise the differences are very 
small and do not seem sufficient to make it a variety. The specimen 
may be described as follows :—Zoarium small, orbicular, moderately 
convex. Zoccia rhomboidal or semicircular, margins distinct ; area 
depressed, aperture large, narrowed below by triangular ale at each 
side. Avicularian areas few, not so large as the zoccia. Under 
surface with radiating grooves. 

All Mr. Woods’s figures for the above memoir are lesa upside 
down, perhaps by an error of the lithographer; and Mr. Woods 
seems to have described his species from the figure when he says of 
the aperture “‘ narrowed above ” instead of “ below.” 

Besides the specimens determined, there are several imperfect 
fragments of other species, among which there is a badly preserved 
Lunulites somewhat resembling L. aperta, T. Woods; and there are 
a few cells of a species (probably with Lepralia growth) with ven- 
tricose punctured cells and a long projecting spout below the aper- 
ture resembling that of L. mamillata (Crag Polyz. pl. vi. fig. 5), 
and a few zocecia of an erect form with elongate subtubular coarsely 
punctured zocecia with a projecting wide peristome ; each zoceclum 


- springs laterally from the one below, in the same manner as in C/r7- 


brillina dentipora, fig. 33. 


EXPLANATION OF PLATES XIV.-XVIII. 
Pruate XIV. 


Fig. 1. Cellaria fistulosa, L., 2°, showing openings for the horny connecting 
tubes. 
2 , 22, showing ee and ovicell. 
3. —— malvinensis, nice 25, showing avicularia. 
4 ovicellosa, Stol., 
5. Section of ditto. 
6, Central walls of ditto, showing rosette-plates. 
7. Smittia centralis, var. levigata, nov. 
8 
9 
10 
12 


. Section of ditto, showing eee plates. 
. Membranipora maorica, Biol Bs 
, Ll. Cellaria fistulosa, Le 
. Cellaria sp., 2° 
13. Schizoporella vigilans, sp. noy., 25 
14, Membranipora lusoria, sp. nov., 2 
15, Schizoporella australis, Ate Woods, 
16. Cellaria globulosa, sp. nov. 22. 
17. End view of do. 
18, 19. Membranipora macrostoma, Rss., 32. 
20, 21. argus, AOrb., 3,2: a, lateral rosette-plates. 
22, 23. concamerata, sp. nov. 332. 


346 A. W. WATERS ON FOSSIL CHILOSTOMATOUS 


Prare XV. 


26. violacea, var. fissa, 25. 

27, 28. Microporella yarraensis, sp. nov., 25. 

29. Microporella enigmatica, sp. nov., *. 

30. , seen from the interior, showing cribriform plate. 
31. Schizoporella phymatopora, Ras., 25. 

32. ——-, inner surface of broken piece. 

33. Cribrillina dentipora, in Bactridium-form, 2,3. 

34, 35, 36. Retepora marsupiata, Smitt, 2°. 


Pruate XVI. 


37, 38. Catenicella solida, sp. nov., 2°. 
39. Catenicella cribriformis, sp. nov., 25. 


40, 41. flexuosa, sp. nov., 2°. 
42, 43. elegans, var. Buskti, 22. 
44, 45. marginata, sp. nov., 32. 


46. Catenicella ampla, sp. nov., 2°. 

47. alata, Thom., 2,5. 

48. Retepora rimata, sp. nov., 7°. 

49. Catenicella alata, Thom., 2°., dorsal surface. 
50. ampla, sp. nov., dorsal surface, 25. 

51, 52. Canda fossilis, sp. nov., 2°. 

53. Retepora rimata, sp. nov., dorsal surface, 2,°. 
54. Mucronella duplicata, sp. nov., #2. 

55. Membranipora geminata, sp. nov., **. 

56. Schizoporella excubans, sp. nov., 2°. 

57. Porina coronata, Rss., 2,5. 

58. Catenicella alata, Thom., 25. 


Prats XVII. 


59. Retepora marsupiata, Smitt, 1,7, basal portion. 

60. Lepralia corrugata, sp. nov., 32. 

61. Retepora marsupiata, Smitt, 3,2. 

62. Cellaria ovicellosa, Stol., 2. 

63. Microporella elevata, T. Woods, 32. 

64. , seen from the interior, +2. 

65. Smittia Tatei, T. Woods, 25. 

66. Mucronella mucronata, Smitt, 32. 

67. Porina clypeata, sp. nov., 25. 

68. Cribrillina terminata, sp. nov., 72. 

69. Porella emendata, sp. nov., 3,2. 

70. denticulata, Stol., 32. 

71. Selenaria marginata, T. Woods, 25. 

72. Microporella ferrea, sp. nov., 32. 

7d. violacea, var. fissa, seen from the interior, showing the elongate 
median pore, 45: a, pore, $5. 

74. Membranipora cylindriformis, nev., 22. 

75. Cribrillina suggerens, sp. nov., 22. 

76. Retepora marsupiata, Smitt, 32. Basal portion. 

Gide marsupiata, Smitt, 3,2. 


Priate XVIII. 


78, 79, Catenicella internodia, sp. nov., 22. 
80. Schizoporella excubans, sp. noy., about 4°: a, aperture of ditto, 85. 
81. conservata, sp. noy., 2°. 


Fig. 82. 
2 83. 
84, 
85. 
86. 
87. 
88. 
89. 
90. 
91. 
92. 
93. 
94, 


BRYOZOA FROM 8.W. VICTORIA, AUSTRALIA. 347 


Membranipora lusoria, sp. nov., 25. 

Microporella see 2ea, sp. nov., 2 . 

clavata, Stol., 

Sehizoporella submersd, sp. nov., 35. 

Caberea rudis?, Busk, 25. 

Lepralia spatulata, sp. nov. fy a 

Porina columnata, sp. SOs ars 

Cellepora fossa, Hasw., 42, 

Microporella elevata, Woods, asa : a, transverse section of growing end. 

Mucronella elegans, MacG., 22. 

Lepralia cleidostoma, Sm., var. PETE: nov., 3. 

Retepora marsupiata, Smitt, 2 

Smittia anceps, MacG., 7: a, ae aperture, 5°; 5, aperture seen from 
the interior. 


Discussion. 


The Presivenr stated that the abstract read only gave the main 
points of a series of elaborate paleontological descriptions. In reply 
to Mr. Charlesworth, he said that Mr. Waters had settled the ques- 
tion between Ehrenberg’s name of “‘ Bryozoa” and Thompson’s name 
of ** Polyzoa” in favour of the former. 


345 R. W. COPPINGER ON SOILCAP-MOTION. 


25. On Somcar-Motion. By R. W. Coppinerr, Esq., M.D. 
(Read March 23, 1881.) 


(Communicated by the President.) 


1 wis to call attention briefly to a phenomenon which, so far as I 
am aware, exists to an unparalleled degree about the shores of 
Western Patagonia, and whose presence there is in a great measure 
due to the exceptionally wet nature of the climate. I allude toa 
slippage of the soilcap, which is, I believe, continually taking place 
over the basement rock wherever the latter presents a moderately 
inclined surface. Some of the effects of this soilcap-motion are apt 
to be confounded with those due to glacial action; tor the soilcap 
takes with it in its progress not only its clothing of trees, ferns, 
and mosses, but also a ‘‘ moraine profonde” of rocks, stones, stems 
of dead trees, peat and mud, whereby the hills of this region are 
being denuded, and the valleys, lakes, and channels, gradually 
filled up. 

On arriving at the Patagonian archipelago my attention was 
directed to this subject on noticing that the lower branches of 
trees fringing the sea-shore were in many places withering from 
immersion in the salt water, and that in some cases entire trees 


had perished prematurely from their roots becoming entirely sub- -— 


merged. On looking more closely I observed that the sodden 
snags of dead timber, mingled with stones, were often to be seen 
at the bottom of the inshore waters, and that the beds of fresh- 
water lakes were plentifully strewn with similar fragments of wood, 
the remains of forests prematurely destroyed. As the soileap, by 
its sliding motion, reaches the water, the soluble portions are 
removed ; and just as stones and boulders are often seen deposited 
in grotesque situations by a melting iceberg or a receding glacier, 
so are the phenomena of ‘perched rocks” to be here observed, 
although, in the class of cases to which I refer, due to a totally 
different cause. These facts are all the more interesting from their 
occurring in a region where the effects of old glacial action are to 
be seen in a marked degree. Planings, scorings, striz, and “ roches 
moutonnées ” may almost invariably be found wherever the rock is 
sufficiently capable of resisting the disintegrating influence of the 
weather to retain these impressions. Thus they are nowhere to 
be seen on the coarse-grained friable syenite, which is the com- 
mon rock-formation of the district; but where this rock is 
intersected by dykes of the more durable greenstone, the above- 
mentioned signs of former glacial action may be seen well deve- 
loped. There are therefore in this region ample opportunities of 
comparing and differentiating phenomena which have resulted from 
‘‘elacial action” and those which are due to “soilcap-motion ”—a 
force now in active operation. 

I may here observe that we did not see any glaciers worthy 


R. W. COPPINGER ON SOILCAP-MOTION. 349 


of the name either on the western islands or abutting on the main- 
land shores of Patagonian channels, although they undoubtedly 
exist further eastward, and discharge icebergs at the head of some 
of the deep fiords. In the main straits of Magellan there are fine 
examples of complete and incomplete glaciers, where one may ob- 
serve in all its grandeur the wonderful denuding power which these 
ponderous masses of ice exercise as they move silently along their 
rocky beds. 

Sir Wyvyille Thomson (vide ‘ Voyage of the Challenger, the 
“ Atlantic,” vol. ii. p. 245) attributes the celebrated “stone rivers ” 
of the Falkland Islands to the transporting action of the soilcap, 
which, among other causes, derives its motion from expansion and 
contraction of the spongy mass, due to varying conditions of moisture 
and comparative dryness ; and this hypothesis is to a certain extent 
supported by the occurrences which I am now endeavouring to de- 
scribe. Here, in Western Patagonia, are evergreen forests, and a 
dense undergrowth of brushwood and mosses clothes the hillsides 
to a height of about 1000 feet ; and this mass of vegetation, with 
its subjacent soil, resting as it frequently does upon a hillside 
already planed by ice-action, naturally tends, under the influence 
of gravitation, combined with that of expansion and contraction, to 
slide gradually downwards until it meets the sea or a lake or valley. 
In the first two cases its free edge is then removed by the action of 
the water, in a manner somewhat analogous to the wasting of the 
submerged snout of a Greenland glacier in the summer time; and 
in the last case the valley becomes converted into a deep morass. 

It appears to me that the conditions which are said to have 
resulted in the production of the ‘stone rivers” of the Falklands 
here exist in equal if not greater force. There is the thick spongy 
vegetable mass covering the hillsides and acted on by varying con- 
ditions of extreme moisture and comparative dryness; there are 
the loose blocks of disintegrating syenite to be transported; and 
there are the mountain-torrents, lakes, and sea-channels to remove 
the soil. Of actual motion of the soileap we have at least strong 
presumptive evidence ; but nowhere in the valleys have I found 
any thing resembling a “ stone river.” 

It might perhaps be thought that a slow and gradual depression 
of the land would account for some of the above phenomena ; but 
I have seen no reliable sign whatever of subsidence, and have, on the 
contrary, the evidence of numerous raised beaches and the work of 
stone-boring mollusca at heights above the present sea-level to prove 
that elevation of the land has taken place. 

The subject is one full of interest ; and feeling confident that it 
will repay further investigation, I take this opportunity of bringing 
the foregoing observations to your notice. 


Discusston. 


The Prestpenr said the theory brought forward in this paper 
would very well account for some cases of the infilling of valleys, 


350 R. W. COPPINGER ON SOILCAP-MOTION. 


lakes, and sea-margins, and was especially important as showing how 
some phenomena ordinarily attributed to ice could be produced. 

Mr. Ussuer asked what the nature of the subsoil was. 

Mr. Forpuam asked for information as to the origin of the soil- 
cap, and its formation on the high ground from which it was stated 
to be slipping. 

Mr. Hawxsuaw remarked that engineers, to their cost, were well 
acquainted with the unstable condition of the soilcap. Small dis- 
turbances often set in motion large masses not only of clays but of 
rocks. Good examples of rock-movement might be seen on the 
Ripponden branch of the Lancashire aud Yorkshire Railway. The 
surface once disturbed, continued to move for long periods. In some 
railway-cuttings the slopes had moved for twenty years. 

Mr. Sprartine called attention to the slipping of clay in the 
Brockley cutting, near new Cross. 

Mr. Huptezsron said this paper supplemented Sir Wyville Thomson’s 
observations on the “ stone rivers ” of the Falkland Isles, where he 
explained the accumulation of quartzite blocks by this kind of slip- 
ping. Such observations tended to show how some of the old 
brec ciasmight have been formed by other agencies than that of ice. 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 301 


26. On the precIsE Mopr of AccuMULATION and Derivation of the 
Moet-Tryran Suey Deposits ; on the Discovery of Smmiiar 
Hieu-Levet Deposits along the HKasrrrn Storrs of the WELsH 
Mountains; and on the Existence of Drirr-zonus, showing 
probable Variations wm the Rate of Susmercencr. By D. 
Macxintosu, Esq., F.G.S. (Read April 27, 1881.) 


ConTENTS. 


J. Introductory Remarks on the variable Character of the Lower Boulder- 
drift of the Lake District, North Wales, &c. 
II. Moel-Tryfan Deposits, &c. 
III. Deposits on Halkin Mountain, Flintshire. 
IV. Discovery of High-level Deposits of Gravel and Sand between Minera 
and Liangollen Vale, Denbighshire. 
VY. High-level Gravel and Sand near Llangollen. 
VI. Remarks on the High-level Gravel and Sand of Macclesfield Forest. 
VII. Arrangement of the Drift-deposits of North Wales into Vertical Zones, 
showing probable Variations in the Rate of Submergence. 
VIII. Concluding Remarks as to whether the Submergence was caused by 
Subsidence of the Land or Rising of the Sea. 


I. Inrropuctory REMARKS ON THE VARIABLE UHARACTER OF THE 
Lower Bovutprer Drirt or tHE Laxe Disrricr, Norra Wats, &e. 


Many years’ observations along the east coast of the Irish Sea, from the 
Solway Frith southwards to the estuary of the Dee, have led me to con- 
elude (as stated in former papers published in this Journal) that while 
the Upper Boulder-clay is a remarkably persistent and homogeneous 
formation, and while the line of separation between it and the 
cleanly washed, obliquely laminated, and boulderless sand and gravel 
(where the two formations are present) is always distinctly marked, 
the Lower Boulder formation varies both vertically and horizontally 
from compact stony clay to loam, gravel, and sand. This is more 
especially the case in the neighbourhood of the mountains and at 
comparatively high levels. Around the mountains of the Lake 
District (where it is called pinnel) it is often interstratified with or 
replaced by well-laminated and often contorted sand and gravel, as 
at Ulverston Railway station, between Ulverston and Arrad Foot, 
&ec. Inthe neighbourhood of Bangor, though a stiff Boulder-clay 
may here and there be seen lying under stratified sand and gravel, 
the two kinds of drift may quite as often be found in horizontal 
succession ; and they both agree in containing boulders (see VII. 
1 and 2). 


II. Mort-Tryran Deposrts, &e. 


1. Bruef History of Discovery.—In 1871, and again in 1880 (last 
year), I had opportunities of tracing the drift-deposits of Caernar- 
vonshire from the neighbourhood of Bangor to the top of Moel 
Tryfan, and of observing a number of facts connected with the Moel- 


352 D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


Tryfan deposits which would appear to have escaped the notice of 
previous observers. But before proceeding to the main subject, it 
may be desirable to give a brief account of the discoveries made since 
the first published account of the deposits. 

Trimmer, in the ‘ Proceedings’ of the Geological Society for 1831, 
has a short notice of the Moel-Tryfan drifts, in which he found Bucei- 
num, Venus, Natica, and Turbo (if correctly identified) beneath 20 ft. 
of sand and gravel. In vol. i. of the ‘ Journ. of the Geological Soc. 
of Dublin’ (1838), he wonders if the granite as well as the flints he 
found on Moel Tryfan came from Ireland. [He heard of quarrymen 
finding sea-shells on Moel Faban, near Bethesda, where Darwin 
afterwards could find no trace of drift likely to contain shells.| In 
his work on Geology, published in 1841, Trimmer gives a general 
section of the drift from Menai Strait over Moel Tryfan to Mynydd 
Mawr. He could find granite erratics only at eight points between 
Menai Strait and Snowdon. 

Buckland, in 1841, found rounded chalk-flints and white granite 
in the Moel-Tryfan deposits. His paper was read before the Geo- 
logical Society, and an abstract of it appeared in the ‘ Athenzeum,’ in 
1842. 

Darwin, in 1842, in the ‘ London, Edinburgh, and Dublin Phi- 
losophical Magazine and Journal of Science,’ gives a very sug- 
gestive description of the Moel-Tryfan deposits. He found boulders, 
chiefly from the neighbouring mountains, but likewise rounded flints 
and white granite. Under the drift he saw that the surface of the 
slate, to a depth of several feet, had been shattered and contorted in a 
very peculiar manner. He did not find shells; but near the summit 
of the hill, on the east side, he saw a thickness of at least 20 ft. of 
irregularly stratified gravel with boulders and layers of sand and 
fine clay. He attributed the shattering and contorting of the slates 
to icebergs grating over the surface, and lifted up and down by the 
tides. The shattered and rounded slate rocks were similar to what 
he had seen in Tierra del, Fuego. He believed that the Chalk tints 
had been brought by floating coast-ice. 

Darbishire gave an account of the Moel-Tryfan deposits in the 
‘ Proc. of the Manchester Lit. and Phil. Soc.’ for 1851-52, and in 
the ‘Geol. Magazine,’ vol. ii. He believed that a layer of yellowish- 
brown sandy clay 1 ft. 9 in. thick had preserved the shells in the 
underlying sand and gravel. He collected a great number of shells, 
a list of which may be found in Mr. Shone’s paper in the ‘ Quart. 
Journ. Geol. Soc.’ for May 1878, and in Dr. Gwyn Jeffreys’s paper 
in the same Journal for Aug. 1880. 

Lyell and Symonds, in 1863, visited Moel Tryfan, and found a 
mass of incoherent.stratified sand and gravel, 35 ft. thick, with 
fragments of shells, and a few whole specimens. In the lower beds 
they saw several large boulders of far-transported rocks glacially 
polished and scratched on more sides than one. 

Ramsay and Etheridge, in 1876, examined the Moel-Tryfan de- 
posits, and found that the boulders were chiefly local, and that the 
sand and gravel were obliquely laminated, similar to what may be 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTs, 353 


seen on a sea-beach. Ramsay gives a comprehensive account of the 
deposits in the last edition of his ‘ Physical Geography and Geology 
of Great Britain.’ He specifies the indirectly local erratics sw Bich 
came under his notice, and regards the mountain from what may be 
called the Snowdon stand-point. Though he had previously observed 
the drift-sections (in 1852), it would appear that the curved slaty 
lamine which I have been led to regard as the most interesting of 
all the Moel-Tryfan phenomena, were not during his visits, or indeed 
during the visits of any other geologist excepting Darwin, sufficiently 
striking to arrest attention, In this paper the mountain will be 
principally regarded from the N.W. and N. stand- -points. 

2. Deposition during Submergence.—lt seems to be generally 
admitted that deposition is in excess during submergence, and denu- 
dation during emergence ; and when we consider the thickness of and 
area covered by the Moel-Tryfan deposits, in connexion with the 
fact that they lie on a sloping surface, and reach up to within a short 
distance of the rocky crest of the hill, the idea of accumulation during 
the submergence or sinking of the land appears the most probable, 
while it is the most consistent with certain facts to be mentioned i in 
the sequel. 

3. Identification of the Local and Erratic Stones—The most pre- 
valent pebble, rock-fragment, or boulder in these deposits is a light- 
coloured felstone, which may have come from the Cambrian conglo- 
merate of the upper part of the hill, or from adjacent hills. The 
fragments of talcose or chloritic schists must have come from rocks 
like those which are now found in the tunnel (near the top of the 
hill), or from similar rocks in the hills around Moel Tryfan. The 
basaltic-looking diorite or greenstone may have come from bands in 
the hill or adjacent hills. Many small quartz pebbles in the drift 
may have come from the Cambrian conglomerate already mentioned*. 
The numerous fragments of slate may be very nearly i situ, or may 
likewise have come from neighbouring hills. There are many 
Eskdale granite pebbles, and a smaller number of granite pebbles 
from the south of Scotland (chiefly Criffel). Chalk flints are rather 
numerous (during my last visit they predominated in a particular 
part of the excavation), and reach a height of about 1350 feet. 
Eskdale granite on Moel Tryfan is found up to at least 1350 feet 
above the sea, or 64 feet above the highest range of the rock zn situ. 
Chalk-flints i situ, in Ireland, as Professor Hull has informed me, 
do not rise higher than about 1000 feet above the sea; so that on 
Moel Tryfan their height is 350 feet higher (see VIII.). On Moel 
Tryfan I found a pebble of red granite of unknown derivation, but 
exactly of the same kind as one I saw on the beach of West Cum- 
berland, and one on the Blackpool beach. 

4. Arrangement of the Gravel and Sand.—Most writers on the 
Moel-Tryfan deposits have noticed the extent to which the sand and 
fine gravel are obliquely laminated, and the resemblance they bear 
to what may be seen on a sea-beach. The sand is often as fine as 


* JT have to thank Dr. Hicks for assisting me in tracing the derivation of the 
local and indirectly local stones. 


304 D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


blown sand, and, when I last saw it, was in course of being blown 
by the wind, from the brink of the quarry-excavation, over the grassy 
flat above. On the right-hand side of the tunnel, where there is no 
clay above the sand, the latter presents a peculiarly involved 
appearance. 

On the left-hand side of the tunnel (1880) the sand inter- 
laminated with gravel had evidently been contorted to a great extent. 
The shell-fragments principally occurred in layers of fine gravel, in 
one place chiefly made up of angular flint chips. There the average 
thickness of the sand and gravel was about 10 feet, the clay above 
being about the same thickness. When Darbishire saw these drifts 
about thirty years ago the clay was only about 1 ft. 9 in. thick. In 
different quarry-excavations at different levels up the hill-side, sections 
more or less differing in their character have been revealed. In 
1880 the brink of the drift-cliff was only about 30 feet below the base 
of the abruptly rising rocky crest of the hill; and it is more than 
probable that the crest once rose as an isolated mass of rock above 
the surface of the glacial sea. The shelly sand and gravel (so far 
as aneroids can be trusted) extend up to quite 1350 feet above 
the present sea-level; and they have been found as low down as 
1170 feet. 

5. Position of Boulders.—One may go for miles along a railway- 
cutting without seeing a single boulder in situ ; but where a large 
clay- or gravel-pit has been excavated by the side of the railway, 
many boulders may be seen on the floor of the pit, the explanation 
being that boulders exposed in a railway-bank very soon tumble 
down and are covered up, blasted, or removed. It is therefore wrong 
to conclude that because there may be many boulders on the floor 
of a quarry in drift-covered rock, they were (as boulders) originally 
situated at the base or towards the base of the drift-deposits. Itis 
probable that many of the boulders of the Moel-Tryfan deposits 
which now lie on the quarry floor, were once dispersed at various 
levels in the overlying drift ; and in 1871 I saw a boulder at least 
3 feet long high up in the sand. This accords with the fact that at 
lower levels, including Anglesey, very large boulders may sometimes 
be found imbedded in sand or fine gravel containing shell-frag- 
ments, as well as in clay. 

6. Bent and shattered Hdges of Slaty Lamine. ata 1871 I noticed 
that the edges of the vertical slates were bent in the direction of 
Mynydd Mawr, or from about N.W. to S.E. Darwin, as already 
stated, observed somewhat similar phenomena in 1842, which he 
attributed to the impact of floating-ice. In 1880 I happened to see 
what may be called a magnificent display not only of the bending 
but likewise of the shattering of the edges of the slaty lamin, and 
of the extreme contortion of the laminz of sand by which the slates 
were covered (fig. 1). From the section (which from the quarry upper 
floor to the top of the clay is at least 20 feet thick) it is perfectly clear 
that the sand and overlying clay must have been deposited before 
the derangement of the clay, sand, and slaty laminee took place. 
The only ‘explanation which appears sufficient to account for this 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. B13)3) 


derangement is the violent stranding of an iceberg or mass of floating 
ice from the N.W. on the previously accumulated clay, which it 
pressed forwards and downwards until it reached the sand, which it 
ploughed up and inverted, and then bent and shattered the edges of 
the slaty laminz, so as to be able to roll up parcels of clay and 
slate-chips in the sand. The line marking the commencement of 
the slaty curvature is nearly horizontal ; but the upper termination 
inclines slightly in the direction of the movement of the floating ice 
(at first probably raised a little above the level of flotation by the 
impact, and afterwards shghtly lowered so as to regain its normal 
level). This section bears no real resemblance to some which are 
found in districts where traces of ice-action are absent, and which 
are believed by some geologists to have been caused by ordinary 
atmospheric action. On Moel Tryfan the finely laminated sand shows 
no trace of having ever been disturbed by the percolation of rain- 
water or by frost, while the preservation of the numerous shell- 
fragments in continuous though contorted layers of sand and fine 
gravel has evidently (as Darbishire long ago pointed out) been 
owing to the clay preventing the downward passage of rain-water. 
As Moel Tryfan is in the midst of a glaciated district, and as ice 
-eapable of bending the slates and contorting the sand must have 
brought the erratic stones, including chalk-flints from Ireland, why 
have recourse to any other supposition to account for the phe- 
nomena ? 


Fig. 1.—Laminated Sand and Bent Slates on Moel Tryfan. 
8.E. NW. 


9 


1S] 


Sat 
Ts 


lates ; B B, Sand, with parcels of slate-chips and Boulder-elay ; 


renin 


AA, 


op) 


7. Direction of Floating Ice in Ireland.—According to the Rev. 
M. H. Close the principal direction of floating ice in Ireland during 
the glacial submergence was from about N.W. to S.E. ; so that the 
local floating ice would only have to persevere in this direction, 
after leaving Ireland, in order to reach Moel Tryfan. 

8. Were all the Moel-Tryfan Shells brought by Floating Ice ?— 
This idea would appear to be untenable for the following reasons :— 
(1) The shells must at first have been somewhere én situ, and why 
not on Moel Tryfan? (2) The Mollusca may have lived where the 
erratic stones were imported as well as where they were exported. 


396 D. MACKINTOSH ON HIGH-LEVEL MABINE DRIFTS. 


(3) Shells have not been found in Eskdale on the old sea-coasts from 
which the granite erratics on Moel Tryfan were transported when 
the land was deeply submerged (see 11). 

9. Cause of the Absence of similar Deposits on the Lower Slopes of 
the Hill.—It has often been remarked that beach-like deposits with 
shells have not been found on Moel Tryfan excepting towards the 
summit of the hill. It is perhaps too early to speak positively on 
this point, as there is a bare possibility of patches of such deposits 
being yet discovered. But the cutting for the railway above Bryn- 
gwyn, which extends from near the base to near the summit of the 
hill, shows no trace of laminated sand and rounded fine gravel, but, 
on the contrary, is everywhere a Boulder-clay, or clayey loam, with 
stones, chiefly angular or subangular. The drift on the north side of 
the hill, so far as revealed by cart-roads, is somewhat similar to that 
above Bryn-gwyn, excepting towards the summit, where there are 
patches of gravel and sand interstratified with clay and loam. The 
cause of the difference in the deposits near the summit and lower 
down cannot (at least principally) be a difference in the sources of 
supply of the lower and higher drifts, because the materials for the 
elaboration of fine sand and gravel exist on the hill-slopes at low as 
well as at high levels. The most probable explanation seems to be 
that the submergence of the lower slopes of Moel Tryfan went on too 
rapidly to allow sufficient time for the accumulation of well-rounded 
beach shingle and sand (see sequel). 

10. Moel-Tryfan Shells and Erratics not pushed up hill by Land- 
1cé.—It has been asserted (though this is not the general opinion) 
that all the shells, along with the erratic stones, were pushed out 
of the bed of the Irish Sea as far south as Moel Tryfan, and then 
up the hill-slopes nearly to the summit by land-ice. But an inti- 
mate acquaintance with the character of the Moel-Tryfan deposits 
precludes this idea; for, if it were a true explanation, the shells and 
erratic stones would have diminished in number the higher up they 
were pushed. but, on the contrary, the shells and erratics in the 
drifts near the sea are fewer in number than on Moel Tryfan. 
This theory would likewise require to invest the land-ice with the 
power of rounding the pebbles derived from the upper part of the 
hill, and laminating the sand and fine gravel; for it ought to be 
remembered that though the sand and gravel are, in places, much 
contorted on Moel Tryfan, the contortion was evidently, in many 
instances, produced after their accumulation. 

11. Shells found by the Author.—As my main object was not to 
look for shells, only the following nine species were named for me 
by Dr. Gwyn Jeffreys out of a number of fragments I collected :— 


Cardium echinatum. Tellina balthica. 

edule. Mactra solida, var. elliptica. 
Cyprina islandica. Saxicava rugosa. 
Astarte suleata. Purpura lapillus. 

borealis. 


It is quite consistent with the remark made in section 8, that 
some of the Moel-Tryfan shells may have been brought along with 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 315i 


erratic stones from a distance by floating ice, provided we admit that 
Moel Tryfan may have returned the compliment; and a number of 
shell fragments which I found along with small flint chips certainly 
looked as if both had been brought and deposited at the same time, 
though their final juxtaposition in the lamina to which (in that part 
of the section) they were limited must evidently have been the 
result of rearrangement on the spot. 

12. High-level Gravel and Sand im other parts of Caernarvon- 
shire.—Near the summit-level of the pass of Llanberis, some years 
ago, | saw a striking section of obliquely laminated sand and fine 
gravel, probably about 1000 feet above the sea-level. The extent to 
which the stones were rounded, the arrangement of the lamin, and 
the position of the deposit (being away from any channel which 
could have conducted a freshwater stream), all pointed to its being 
of marine origin, though I did not see any shells. Above Bethesda, 
near a farm-house marked Gwaun-y-gwiail on the Ordnance map 
(probably about 1000 feet above the sea), a gravel-pit shows lami- 
nated and contorted gravel and sand, in which (for reasons which 
need not be stated) I had not time to look for shells; but Trimmer 
told Ramsay that he had found shells in that neighbourhood up to 
from 1000 to 1200 feet above the sea. It does not seem to be 
well known that Ramsay found shells in Boulder-clay on Fridd 
Bryn-mawr (west of Llanberis) at a height of about 1000 feet above 
the sea. 

13. High-level Gravel and Sand wm Ireland.—It may be neces- 
sary to complete this account of high-level marine drifts in the 
western part of Wales by adding a brief statement of what Close 
found on the Three-rock Mountain near Dublin (see Geol. Mag. vol. 1. 
Decade ii. 1874). He collected shells from gravel-pits at 850 feet, 
1000, 1100, and a little higher than 1200 feet above the sea. 
The Three-Rock Mountain consists of granite; but the stones com- 
posing the gravel were very nearly all limestone, which must have 
been brought by floating ice from the N.W. ‘They were nearly all 
subangular; but the deposits partly consisted of clean stratified 
gravel and sand [showing the action of the sea on the spot?]. He 
believed that the shells were brought along with the stones by 
floating ice, because both were scratched. He found more or less 
clay above the shelly deposits, and was led to believe that the 
mountain must have been submerged up to 1760 feet. The Three- 
rook Mountain is related to Moel Tryfan by the altitude of its shelly 
drifts, and by the direction of the floating ice which brought its 
erratic stones. 


Ill. Depostrs on Hatxin Mountain, FLINTSHIRE. 


Several years ago I read an account of these deposits before the 
Chester Natural-Science Society. Since then they have been mapped 
by the Geological Surveyors. The mountain, including its westerly 
continuation as far as the Vale of Clwyd, is a plateau surrounded on 
all sides by lower ground, so as to render it certain that the mounds 


Q. J.G.8. No. 147. 23 


358 D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


of gravel and sand on its summit could never have been accumulated 
by the action of fresh water flowing onto it from a higher level. The 
mounds range from about 750 to about 900 feet above the sea. The 
rounded stones they contain are chiefly local limestone and grit of 
Carboniferous age; but Eskdale granite and Lake-District felstone 
may be found in some places, and chalk fiints may occasionally be 
seen. The most striking mounds in which excavations have exposed 
sections are one situated near Brynford (not far from Holywell) 
and another towards the 8.E. end of the mountain. The latter is 
named Moel-y-Crio on the Ordnance map, and rises from the 
summit of a ridge about 950 feet above the sea, with ground falling 
all around it, except on one side, where for some distance it is 
nearly horizontal, and then falls to a lower level. It would appear 
to have been accumulated not so much by wave-action as by the 
piling agency of sea-currents while meeting or parting. 


Fig. 2.—Perched Gravel-mound on Halkin Mountain. 


The Halkin-Mountain deposits of gravel and sand occur at a 
lower level than the other high-level marine drifts described in this 
paper; and they are not very distinctly separated (excepting, in 
most places, along the east side) from the low-level gravel and sand, 
which extends upwards from the sea-coast, and inland along the pass 
between Mold and Bodfari, which is traversed by the Chester and 
Denbigh Railway. Exceedingly few shell-fragments have been 
found in the gravel and sand of Halkin Mountain ; but its compara- 
tively flat summit, in places surrounded by ridges, must have been 
very favourable to the accumulation of rock -fragments derived from 
the inner slopes of the ridges; while it is easy to understand that 
these fragments, instead of being washed down the mountain-sides 
into deep water, would remain for a considerable time at the mercy 
of sea-waves, so as to become more or less rounded and smoothed by 
attrition. It ought likewise to be remembered that the mountain, 
while an island, must have been exposed all round its coasts to 
winds, which, by increasing the power of breakers, may have expe- 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 359 


dited the process of rounding stones; so that in the case of the 
summit of this mountain (which is situated at a lower level than 
that of the gravel and sand on Moel Tryfan, Frondeg, Macclesfield 
Forest, &c.) it is not necessary to suppose that the land subsided 
more slowly than at the levels immediately below and above, so as 
to afford more time for elaborating rounded gravel and sand. As 
already hinted, the scarcity of shell-fragments accords with the 
idea that the rate of submergence was not sufficiently slow to allow 
much time for the growth and accumulation of Mollusca. It ought 
likewise to be stated that on many parts of the mountain the stones 
in the grayvel-beds are not much rounded. 


LY. Discovery oF HigH-LEveL Depostts oF GRAVEL AND SAND 
BETWEEN MINERA AND LLANGOLLEN VALE, DENBIGHSHIRE. 


For a long time, while travelling by railway between Wrexham 
and Ruabon, I fancied, from the suriace-configuration of the eastern 
or outer slope of the range of mountains between Minera and Llan- 
gollen vale, that marine gravel might there be found at about the 
same height as on Moel Tryfan; but, being then bent on finding 
sea-shells at higher levels than the Moel-Tryfan deposits, I did not 
explore the district until near the close of last year (1880). From 
the railway, for some distance westward, the ground is flat or gently 
undulating, with an average elevation of between 300 and 500 feet. 
In many places sand may be found under a deposit which is hori- 
zontally continuons with the upper Boulder-clay of Cheshire; and 
here and there the sand, graduating into gravel, rises in the form 
of mounds or knolls. From the commencement of the gradual rise 
of the ground in a westerly direction up to about 1000 feet the pre- 
vailing drift is a clay or loam, with angular stones and large Carbo- 
niferous-grit boulders from the mountain-range above mentioned. 
At about 1000 feet the ground rather suddenly rises, and the stones 
(as may be seen in ploughed fields &c.) become more or less rounded. 
A little west of Braich, in the district called Frondeg (see 1-inch 
Ordnance map), the ground abruptly swells into a series of ridges 
and hillocks, which consist of well-rounded gravel and sand. 

1. Surface-configuration and Character of the Deposits—In the 
Frondeg district the gravel and sand knolls, with intervening or ed- 
jacent deposits of a similar character, extend for about a mile and 
a half from north to south, and about one third of a mile in breadth 
from east to west; but as similar accumulations are repeated at inter- 
vals as far south as Mountain Lodge, the whole length of the deposits 
may be about three miles—that is, supposing they extend no further 
south. On walking from Braich to the summit of the mountain-range 
in a westerly direction, after passing the gravel mounds, the surface 
becomes very flat and covered with peat. ‘The breadth of the flat 
(which in some places extends, with a very gentle ascent, as far as 
the summit of the ridge, in other places not quite so far) is about 
half a mile. So far as can be seen in brook-channeils, the peat 


overlies a deposit of clay with angular stones and large angular 
2B 2 


360 D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


boulders. The summit of the ridge varies from about 1450 feet to 
about 1500 feet. The well-rounded gravel-and-sand commences 
about 1100 feet (west of Braich), and extends in a westerly direction 
up to about 1350 feet. The hillocks (west of Braich), in one of 
which I found numerous shell-fragments in a grayel-pit, range in 
height from about 1100 feet to about 1250 feet. Above Mountain 
Lodge (8. of Frondeg) there is a gravel-pit in a knoll about 1200 
feet above the sea, and one further west (in which I found shell- 
fragments) about 1230 feet. But the plateau, near the south edge of 
which these pits are situated, consists of rounded gravel-and-sand, 
which, in a westerly direction, rises to at least 1350 feet above the 
level of the sea*. In the gravel-pits and in the neighbourhood 
where sections are exposed in brook-channels, the stones, especially 
in the Frondeg district, are very much rounded, more so, in fact, 
than on the shores of the Irish Sea, where many of the stones (being 
derived from drift-deposits) have undergone a double process of 
rounding. This is the case especially with the Hskdale-granite 
pebbles, which, in the Frondeg gravel-pit, are almost as numerous 
as the local pebbles of Carboniferous grit or sandstone. In addition 
to these, there are chalk-flints, pebbles of Lake-district felstone 
and felspathic breccia, and local pebbles or fragments of Carboni- 

ferous limestone, with much decomposing coal and many coal-measure- 
fossils, which can be best accounted for by supposing them to 
have been locally worked up from strata now concealed under the 
drift-deposits. In the Mountain-Lodge gravel-pits there are many 
Carboniferous-limestone fossils, and much granite. 

2. Shell-fragments. --In the Frondeg gravel-pit shell-fragments 
are very numerous, but I could find no whole specimens. This can 
readily be explained by supposing that the excessive attrition to 
which the extra-rounded pebbles were subjected must have extended 
to the shells which were thrown up on the sea-beach. Among the 
fragments I collected from the Frondeg gravel-pit, and partly from 
one of the Mountain-Lodge gravel-pits, Dr. Gwyn Jeffreys has 
identified the following species :— 


Cardium echinatum, Linné. Tellina balthica, L. 
C. edule, Z. Mactvra solida, L. 
Cyprina islandica, L, Mya truncata, L. 
Astarte borealis, Chemnitz. Fusus antiquus, Z. 


3. Position of Boulders—So far as can be seen in the grayel- 
pits and brook-sections, the large angular boulders are chiefly to be 
found in the clay, which extends under peat, from the west border 
of the gravel-hillock zone up to near or (in some places) quite to the 
summit of the mountain-range. They are likewise found, some- 
times thickly strewn, on the surface of the clay, and to a less 
extent on the surface of large mounds, which partly, at least, con- 
sist of rounded gravel. They may also be seen scattered over the 
lower country as far east at least as the railway. In the Frondeg 


* T have to thank the Director of the Ordnance Survey, Southampton, for 
some of the above heights. 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS, 361 


district, where they consist of Carboniferous grit or sandstone 
(and partly of quartzose conglomerate), many of them must have 
been floated in an easterly direction from near the summit of the 
mountain-range after the submergence had reached that altitude ; 
for how otherwise can their position on the surface of the drift- 
deposits be explained ? 

4. Junetion of the Arenig-felstone and Eskdale-granite Dispersions. 
—West of Frondeg, on the summit-level of the mountain-range, about 
1500 feet above the sea, and extending a short distance eastwards, 
the stones, with the exception of those of local derivation, consist of 
Arenig felstone ; and large boulders of the same felstone are found 
on or near to the summit of the range further south, as well as 
strewn over the whole district between there and the Great Arenig 
mountain. I was surprised to find so great a number of Arenig 
pebbles on the ridge west of Frondeg, and equally surprised to dis- 
cover that they abruptly terminated in an easterly direction not far 
from the water-parting, where their place (with very little dove- 
tailing) was taken by Eskdale granite, reaching up to about 
1400 feet. I arrived at the conclusion that there the northern-drift 
current which floated the Eskdale erratics was sufficiently powerful 
to turn aside the current by which the Arenig erratics were floated 
from the west. I cannot believe that the latter could have been 
brought by land-ice, because 1b appears improbable, if not impos- 
sible, that land-ice proceeding from the Arenig mountain could 
have attained a surface-level of 1500 feet (above the present sea) 
by the time it reached the ridge west of Frondeg; and it may like- 
wise be remarked that as Arenig erratics have been found, not far 
from this ridge, on Cyrn-y-brain up to a height of 1830 feet, the 
supposed land-ice which brought them must have reached as high as 
that level, while further S8.W., on Moel Gamelin, it must have 
reached to about 1900 feet*. This is about the limit reached by 
the glacial submergence in the N.W. part of Wales, where (as well 
as in the Frondeg district) the upward termination of the zone of 
rounded gravel-and-sand may be explained by an increased rapidity 
in the rate of submergence having deprived the sea of the time 
required to round stones by a process of combined rolling and 
attrition +. 

5. Frondeg Erratic Stones and Shells not transported by Land- 


* See my paper on the Boulders of North Wales in Quart. Journ. Geol. Soe. 
vol. xxx. (1874), p. 711. 

t Mr. 8. V. Wood, F.G.S, has written to me to the effect that he regards 
the discoveries I made in the Frondeg district as corroborating the conclusion 
at which he arrived after reading my paper on boulders (Quart. Journ. Geol. 
Soc. vol. xxxv. (1879), p. 425), namely that land-ice extended from the Arenig 
mountain to the top of the ridge west of Frondeg, on which it left the Arenig 
erratics ; while floating ice from the Lake Districs brought the Eskdale erratics 
to the edge of the land-ice which prevented them from reaching further west, at 
the time when the submergence culminated at about 1400 feet. Mr. Wood 
likewise believes that the land-ice of the central part of North Wales spread 
out towards the N.K. and north, so as to prevent the erratics brought by floating 
ice from the north from getting into the interior of the country. 


362 D, MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


wce.—The idea of the granite pebbles of the Frondeg dis- 
trict having been brought by land-ice is opposed by the fact 
that few HEskdale-granite pebbles (so far as I have noticed) are 
to be found between Frondeg and the estuary of the Dee (the 
direction in which the ice would most probably move) excepting 
near the Dee estuary. There is likewise an equal absence of shell- 
fragments in the drift-deposits where the latter are present; but 
it is a remarkable fact that a great part of the area intervening 
between Frondeg and the Dee estuary is free from drift. Sup- 
posing the erratics and shells to have been pushed forward under 
the ice, they ought to be represented over the whole, or at least the 
greater part, of the intervening area, while no one would suppose 
that the numerous extra-rounded pebbles of Frondeg could have 
come in the form of a supraglacial moraine. 

6. Probable Decrease in the Rate of Submergence.—It has already 
been remarked that from about 500 feet to about 1000 feet on the 
east side and flanks of the mountain-range under consideration the 
stones in the drift are generally angular or subangular, while 
above 1000 feet they are generally rounded, and in many places 
extra-rounded. Itis likewise true that of the standard Frondeg 
erratics, namely Eskdale-granite, not one in fifty can be found in the 
comparatively low-level drifts between the mountain-range and the 
railway, while in the Frondeg district, above 1000 feet, Eskdale- 
granite pebbles are very numerous. We have no reason for sup- 
posing that the sea in this latitude was sufficiently warm to melt 
the floating ice, so as to cause the precipitation of many erratic 
stones. On the contrary, the great number of granite pebbles 
which have found their way as far south as Shrewsbury (where they 
are rather numerous in the lower Boulder-gravel, though not in the 
upper clay), and the much greater number (within a limited 
breadth of area) which reached as far south as Burton (S.W. of 
Broseley)* would seem to indicate that where many of these 
pebbles are found crowded in a small compass (as in the Frondeg 
district) they were left by the stranding and consequent breaking- 
up of floating ice. At this period the district was probably in the 
condition of a littoral zone, which may have lasted for a time 
sufficient to enable the waves to round the stones and to allow 
the Mollusca to multiply in the littoral and sublittoral zones, and 
thereby furnish many shells destined to be reduced to fragments by 
the rolling and grinding of stones on a much-exposed sea-coast. 
But a protracted sojourn of sea-waves in what is now the Frondeg 
district is hkewise indicated by the time required for the accumula- 
lation of the immense number of erratic stones. That these 
stones were rounded approximately in setw is evident from the fact 
that at the high levels in the Eskdale district, from which the 
stones must have been launched, the stones are all more or less 
angular. 

* See Quart. Journ. Geol. Soe. vol. xxxvy. (1879). p.425. The Frondeg erratics 
are not mentioned in this paper, as I had not discovered them when the paper 
was written. 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 363 


V.—HieH-LEVEL GRAVEL AND SAND NEAR LLANGOLLEN. 


Around Llangollen and elsewhere in North Wales a kind of 
loamy clay, or clayey wash, often forms the surface of the ground, 
and conceals what lies underneath. It may have been partly 
deposited as the land sank into deeper water, and partly during the 
rising of the land. The existence of an unknown extent of gravel 
and sand under this clay at a high level might never have been dis-: 
covered but for the excavation of a pit a short distance east 
of Rhos Pengwern farm, in a field nearly 1200 feet above the 
sea-level. Under about 3 feet of clay, with perfectly angular 
stones, there is an unknown thickness of fine and coarse gravel and 
sand arranged in lamine more or less arch-shaped. The stones are 
considerably rounded and smoothed, though the sand does not 
appear to have been very cleanly washed. ‘There can, however, be 
no doubt about its being a sea-coast or shallow-sea deposit (though 
I could not find any shell-fragments), as its situation precludes the 
idea of its having been accumulated by any freshwater stream. 
More extensive excavations are required to show both its horizontal 
and vertical extent. At lower levels down to about 500 feet or 
400 feet above the sea the drift contains scarcely any rounded 
stones, while it is particularly worthy of remark that in a pit at 
the east end of Grouse-box Hill (not far from Pengwern pit) about 
1300 feet above the sea, the gravel is perfectly angular, and con- 
tinues so up to the summit of the hill, 1715 feet above the sea. 


VI.—Remarks oN THE HigH-LEVEL GRAVEL AND SAND OF 
MaccLEsFIELD Forsst. 


The identity in level of the Frondeg and Macclesfield-Forest 
deposits renders it appropriate that some notice should be taken of 
the latter. They were discovered by Professor Prestwich in 1862, 
near the Setter Dog Inn, at an altitude between 1100 and 
1200 feet above the sea. He found more or less clay both below 
and above the shelly gravel and sand; but I should be inclined to 
regard the clay above as on a horizon distinct from that of the 
upper Boulder-clay of the plain of Cheshire, and more or less allied 
to the patches of clay or loam which overlie the high-level gravel 
and sand of North Wales. I found the shelly gravel near the 
Setter Dog Inn graduating eastward and upward, in the direction of 
Shining Tor, into angular gravel at a height of more than 1350 feet 
above the sea. At a greater height, in an easterly direction, all the 
gravel is angular; but S.E. of the Setter Dog, in Chapel Lane, 
rounded erratics may be found up to about 1400 feet. From these 
facts it would appear that the two gravel districts, the one on the 
west (Frondeg), and the other on the east (Macclesfield Forest), cor- 
respond almost exactly in level—Near Clulow Cross, some distance 
south of Macclesfield Forest, Mr. Sainter (of Macclesfield) has dis- 
covered gravel and sand with sea-shells about 1130 feet above the 
sea (see his interesting work entitled ‘Rambles round Macclesfield’), 


364 D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


So far as I can recollect, the western slopes of the Penine hills 
between the above-mentioned levels and the plain of Cheshire are 
comparatively, if not entirely, free from rounded gravel; but I 
hope to have an early opportunity of speaking with certainty on 
this subject. 


VII.—ARRANGEMENT OF THE Drirr-pEposits orf Norra Watzs 
INTO VERTICAL ZONES, SHOWING PROBABLE VARIATIONS IN THE 
Rate oF SUBMERGENCE. 


The existence of the above rounded gravel- and sand-deposits, 
with shells, at about the same level in different parts of North 
Wales, and likewise in England and Ireland, could scarcely have 
been the result of accident. At a somewhat lower level than these 
deposits the drifts are angular, with a few exceptions, which may 
have arisen from local conditions having been unusually favourable 
to the rounding of stones by rolling and attrition. Above the level 
of the shelly deposits rounded gravel and sand with shells would 
appear to be everywhere absent. If the time required for rounding 
stones be viewed in connexion with that necessary for the migration 
and multiplication of Mollusca (as already remarked), it will not 
appear too fanciful to suppose that the rate of submergence was 
slower (probably much slower) in what may be called the rounded 
gravel and shelly zone than in the zones above and below, especially 
as it 18 a priort improbable that the submergence (up to at least 
1350 feet above the present sea-level) progressed at a uniform rate. 
Many believe that the submergence terminated at the upper lmit 
of this zone ; but, as long ago advocated by Professor Ramsay, and 
recently repeated in his ‘ Physical Geography and Geology of Great 
Britain,’ a clay similar to that in which he found shells west of 
Llanberis may be seen rising to a height of from 1500 feet to 
1800 feet about the Turbary, and east of the river Ogwen. I noticed 
that the drift between the Turbary and the final ascent to 
Marchllyn Mawr (about 1700 feet) consisted either of clay or small 
chips and fragments arranged in a manner which could not possibly 
be explained by the action of fresh water; and in a continuation of 
this drift at a lower level the late Mr. Griffith Ellis, of Llanberis, 
told me he had discovered sea-shells. In the basins of the Llafar 
and Caseg (S.E. of Bethesda) the general surface-configuration of 
the ground up to about 1900 feet can be more easily explained by 
the former action of the sea than by freshwater agency; and this 
accords with the height reached by the Arenig boulder-dispersion 
in the nighbourhood of Llangollen (1897 feet on Moel Gamelin) 
and on the neighbouring hill-sides. In the present state of dis- 
covery it may therefore be said that the glacial submergence 
certainly reached an altitude of about 1350 feet (the extreme 
height at which sea-shells have been found), and very probably 
culminated at about 1900 feet. Below the middle (or Moel- 
Tryfan, Frondeg, and Macclesfield-Forest) zone of rounded and 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 365 


shelly gravel and sand, as already stated, there would appear to be a 
zone consisting of loam, with angular or subangular stones and no 
shells; but this zone is not uniformly continuous, as on Halkin 
Mountain (Sec. III.), and possibly in other places; it is varied by 
accumulations of more or less rounded gravel and sand. Below the 
third zone (order descending), and extending down to the existing 
sea-level, there is a zone of horizontally alternating clay, and 
angular and rounded gravel and sand, with boulders and shells*. 

Causes of the horizontal Discontinuity of the Middle Zone of 
rounded Gravel and Sand.—The question may naturally be asked, 
How is it that the rounded and shelly drift of Moel Tryfan, 
Frondeg, and Macclesfield Forest are horizontally separated by 
areas in which the drift is angular? We may, I think, be enabled 
to arrive at some explanation of this fact by supposing that rounded 
drift on this horizon may exist in many places under a covering of 
Boulder-clay, which may have been wholly or partly left during 
the rising of the land; but there are other considerations which 
might account for the local absence of rounded drift, one of them 
being that the sea, by merely lingering for a time near the same 
level, might not be able to accumulate rounded drift without being 
assisted by other favourable circumstances or conditions (positive 
or negative) such as the following :—(1) a sea-coast exposed to 
wave-producing winds; (2) the previous or contemporaneous 
accumulation of heaps of angular débris by melting or stranding 
icebergs or coast-ice, frost, rain, gravitation, &c.; (3) the absence 
of steep slopes which would so facilitate the washing-down of pre- 
existing angular débris below the reach of wave-action as to pre- 
vent the accumulation of rounded stones} ; (4) the absence on sea- 
beaches of clay, which would be unfavourable to the rolling of 
stones ; (5) the prevalence of stones susceptible of being easily 
rounded; (6) the existence of rapid currents capable of rolling 
stones below the limit of wave-action ; (7) the preservation of 
relative levels of the differently characterized zones during the 
emergence of the land; (8) the non-alteration of the character of 
the zones by deposits left during emergence; (9) the non-removal 
of preexisting rounded gravel and sand from valleys and slopes by 
glaciers during or after emergence. The last consideration is very 
important ; for, according to Professor Ramsay (to the great accu- 
racy of whose observations in North Wales I can bear humble 
testimony), the marine drift of the northern valleys was ploughed 
out by the second glaciation of the country. 


* Tt might unnecessarily complicate the main subject of this paper were I to 
_ refer particularly to a zone of fine rounded gravel and sand, with no boulders, 

at a low level in the plain of Cheshire and part of Lancashire (which I believe 
was deposited during the last stage in the rising of the land), or to the upper 
Boulder—clay at comparatively low levels, which I believe was the result of 
a re-submergence of the land. I lately found proofs of a land-surface between 
the upper Boulder-clay and underlying boulderless sand in excavations under 
Crewe railway-station, an account of which will soon be published. 

+ On the very steep slope of Mynydd Mawr, which is nearly opposite to the 
marine drift on Moel Tryfan, the stones are all angular. 


366 D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


VIII.—Concivpine REMARKS AS TO WHETHER THE SUBMERGENCE 
WAS CAUSED BY THE SUBSIDENCE OF THE LAND OR RISING OF 
THE SEA. 


In favour of the extension to the district under notice of Dr. 
Croll’s theory of a rise of the sea-level by the melting of circum- 
polar ice, and consequent displacement of the earth’s centre of 
gravity, and subsequent fall of the sea to its present level, it might 
be argued that this theory would most readily explain the coincidence 
in level between the marine drifts of Macclesfield Forest, Frondeg, 
Moel Tryfan, and the Three-Rock Mountain, especially when we 
take into account the great distance between the first- and last- 
named localities, and the comparatively very small vertical distance 
between the highest known shelly drifts and the present sea-level 
(about 700 to 1). It would likewise account for variations in the 
rate of submergence, by attributing the variations to changes in the 
rate of circumpolar liquidation, resulting from changes in tem- 
perature. Against the application of this theory to the district 
under consideration, we must take into account the differences in 
level between different parts of the land as it stood when the above 
marine drifts were deposited, and as it now stands. These dif- 
ferences would seem to indicate horizontal inequalities in the rate at 
which the land emerged, as might be expected on considering the 
extent of the submerged area. Thus the level of the greatest 
height reached by the chalk in Ireland (as Professor Hull tells me) 
nowhere exceeds 900 feet or 1000 feet. But at the time when 
chalk flints were transported to Moel Tryfan the chalk in Ireland 
must have reached to about 1350 feet above the present sea-level. 
The greatest height to which Eskdale granite extends is 1286 feet ; 
but at the time when when this granite was transported to Frondeg, 
it must, in situ, have reached to atleast 1400 feet. Ihave ventured 
to make these few brief remarks in conclusion with the view of 
stimulating to further inquiry on the subject, and partly because IL 
have reason to believe that so eminent an authority as Dr. Darwin 
is now inclined to the opinion that many of the apparent changes 
of level in the land were caused by changes of level in the sea. 


SUPPLEMENTARY NOTES. 


During the present month (April, 1881), having gone over the 
ground described in this paper a fourth time, I am able to furnish 
the following additional details:—On going south along the west 
side of the axial north-and-south ridge of Minera mountain, I saw 
numerous large and small angular stones (Carboniferous grit, sand- 
stone, and quartzose conglomerate); but I could find few or no 
rounded stones, excepting among the felstone erratics, which must 
have come from the west. After crossing the ridge and going in a 
S.E. direction down the eastern slope of the mountain, I arrived at a 
point nearly half a mile north of Krwau. Here it may be necessary 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 367 


to remark that though rounded stones in drift may often be seen in 
preok-channels and on roadsides, where the ground is flat, I have 
not examined any hillock or small mound which did not show signs 
of its being composed of rounded gravel and sand. Besides the tour 
hillocks with gravel-pits (in one of which I found shell-fragments) 
near Mountain Lodge, during my last visit I counted ten which 
evidently consisted of drifts similar to deposits on our present sea- 
coasts. North of a brook which joins another brook near Krwau, 
I saw three rather low mounds with surface-excavations sufficient 
to show that they consisted of rounded gravel and stratified sand. 
The furthest north of the mounds was quite 1300 feet above the 
sea-level. While continuing my journey in a northerly direction, 
I saw many well-rounded stones in drift exposed in a brook-channel 
about 1350 feet above the sea. On descending towards a house 
called Braich (1031 feet above the sea, as I have been informed by 


Fig. 3.—Section of Drift-deposits between Braich and the Summit 
of the Ridge, Frondeg, Denbighshire. (The levels, with the 
exception of the lowest and highest, are approximate.) 


W. E 
A ; 3 
¢ % ai 3 2 2 
o o A ao a a 
Ss set a 4 Ss] 9 & 
rs > O) ises ae - en 
wt Ye) » ° a v me) Be 
Se & Ge a Gro, ais) a8 = 
Do tant a ao +O a 
i ae Sun Teva ae iG 
S oO = oe) 045 3 
5c ete Romer” ole 3 
Ex Peat covering clay 3 By IB Gis) Bios a 
5 with angular stones. oO a S) Oo Q 
f | 4 i ' 
i t 1 1 I 
1 ! , , 1 1 
fo ; i} } ! 1 i 
D af 1 1 ' 1 { 
Tp i I \ 1 l I i 
ee ~. “ry ~ 1 1 ! 1 : ' 
=e SSS df A 4 eS 1 I 1 1 { 
—a... nos Lee 
a ~ Pein, 1 
ee ON | FER OUS c al ' f 
—_____ —_ >_—. Rl T AND S Fas 
aoe 


the Director of the Ordnance Survey), I encountered a knoll (fig. 3) 
about 1250 feet, showing the marks of a now disused gravel-pit, 
containing many rounded stones and sand. The shelly gravel-pit 
(described in the paper) lies between the above knoll and Braich. 
North of the latter there are three gravel hillocks between 
1100 feet and 1150 feet. West of the house called Cae-mynydd 
there is a rather large hillock showing the remains of old gravel- 
pits. West and N.W. of it there are several large mounds (the 
highest nearly 1400 feet above the sea), the character of which is 
doubtful; but in one of them several small excavations show a 
mixture of rounded and angular gravel. A similar kind of drift 
may be traced nearly to the north end of the mountain. I had pre- 
viously seen Kskdale granite only in the shape of small pebbles ; 
but on this occasion I stumbled on two large blocks, one N.W. of 
Cae-mynydd, and the other associated with numerous millstone- 
grit blocks on the summit of the axial ridge (about a mile west of 


368 D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS. 


Braich) at a height of about 1450 feet above the sea (fig. 3), which 
proved a submergence of the mountain to at least that extent. 

During a fourth visit to the Mountain-Lodge district (above 
Ruabon) I found that higher up than the extensive area of rounded 
gravel there was an interval of angular detritus, above which (and 
near to the summit of the mountain-range) at a height of about 
1550 feet, there was a mass of gravel as much rounded as that in 
which I found shells at a lower level. Gravel more or less rounded 
reappeared on the west side of the mountain at nearly the same 
height. 

During a third visit to Moel Tryfan (June 1881) I found that 
several new sections had been exposed, in which a close inspection 
showed that the edges of the slaty laminse were not continuously 
curved, though the broken-off chips were arranged along more or 
less curved lines which inclined from the N.W. 

I ought to mention that, in the observations which have resulted 
in this paper, | was again assisted by the Committee of the Govern- 
ment Fund for Scientific Research. 


Discussion. 


The President, in inviting discussion, spoke of the great value of 
Mr. Mackintosh’s paper. 

Mr. DuRance spoke of the laborious manner in which Mr. 
Mackintosh had carried out his work, and thought that he had dene 
much to refute the theory of a universal ice-cap overwhelming every 
thing. He said that he had examined some of the Molluscan fragments 
in Ireland, some of which were, indeed, scratched; but others, he 
thought, could only have been deposited during a submergence. He 
thought that the positive evidence in favour of submergence must 
overpower mere theory. As you ascend the slope of the Halkin 
mountain from the river Dee, you pass first over Glacial Drift con- 
taining erratics from the Lake district and south of Scotland ; mm the 
lower part a Boulder-clay resting on sands with Mollusca; but higher 
up the hill there is a different drift, local in character, and contain- 
ing only North-Wales rocks. 

Mr. UssHrr inquired if the Moel-Tryfan deposit abutted against 
higher ground, as then the subaerial débris shed upon the beach 
might have been mixed with it, and a local reversal of lamin pro- 
duced by the impact of an iceberg. 

Prof. Bonny said that he thought the facts. brought forward by 
Mr. Mackintosh would give a death-blow to the idea that the Mol- 
lusca had been brought up hill by glaciers. He doubted, however, 
whether the creep of the mingled sand and clays down hill would 
not account for the bending of the slate-edges better than the 
srounding of an iceberg; for would not that bend them up hill? and 
how would it ground on the lee side of the hill? If terraces were 
formed by the rising of the sea due to a polar ice-cap, then they 
should not be uniform in level, but rise along circles of longitude 
towards the pole. 


D. MACKINTOSH ON HIGH-LEVEL MARINE DRIFTS, 369 


Dr. Hicks thought that no theory could be founded on the special 
direction of the slaty curvatures, as he had seen them in all direc- 
tions. He asked if a glacier crossing Ireland along a north-west 
direction might not dam the sea up, so as to partially overwhelm 
Wales. He did not understand how the erratics could arrive at 
exactly the same points both from Ireland and the Solway. 

Mr. Macxintosu, in reply, said that he thought that a grounding 
iceberg might, if raised first up hill, then slip slightly down hill, 
and so bend the edges of the slaty lamine. To Dr. Hicks he replied 
that the currents in the sea were far from being necessarily parallel. 


370 C. PARKINSON ON THE ISLE-OF-WIGHT 


27. Upprr Greensanp and Canoritic Mart, Istz or Wicut. 
By C. Parkinson, Esq., F.G.S. (Read March 23, 1881.) 


AurHouer the Isle-of-Wight Greensands have been well explored by 
eminent geologists, such as Fitton, Mantell, Ibbetson, and Saxby, 
there yet remains ample work for paleontologists in the classifica- 
tion of the fauna characteristic of each horizon. Hitherto the 
fossils from different zones have not been clearly separated, the 
collections from the Malm rock being mixed with those of the 
upper cherts and rags, and the Chloritic Marl with both. In form- 
ing a collection illustrative of the above series, it has been the writer’s 
aim to divide every zone and to define the extent of each. 

At the base of the section a band of chert nodules is indicated, 
measuring 2 feet, which is possibly the true junction of Gault and 
Greensand, although in the Memoir on the Geological Survey of the 
Isle of Wight 50 feet of micaceous sands below this pomt have been 
included in the Greensand ; this has also been done more recently 
by Dr. C. Barrois, of Lille. It was from this section, exposed on 
the St.-Lawrence beach, that Captain Saxby procured a species of 
Crayfish ; and a second specimen has been found within the last 
eighteen months in the same locality. The importance of this 
horizon has apparently been overlooked ; for quite recently a further 
valuable addition has been made to the fauna by the discovery of a 
Chelonian previously unknown to paleontologists. From Captain 
Saxby’s day to the present time the locality has been neglected. 
About a year ago Mr. Mark Norman, a well-known local geologist, 
noticed a huge boulder of this blue chert lymg exposed to the wash 
of the tide; it had certain indications on the exterior which led 
him to examine the whole surface with care; there were, in fact, 
the perforations of bone clearly shown. With great care the 
hard matrix was chiselled out, revealing portions of the carapace 
and rib-bones of a Turtle of the family Paludinosa. The specimen 
is now in the Natural-History Museum, South Kensington ; and 
through the courtesy of Prof. Owen I am enabled to give the 
following notes, abridged from a description kindly made by him. 
It is referred to the 


Order CHELONIA. 
Fam. PALupINosa. 
Genus PLastremMys, Owen. 


PLASTREMYS LATA, Owen. 


A new genus and species of freshwater Tortoise from the Isle- 
of-Wight Greensand, remarkable for its breadth in proportion to 
length. In this character it approaches nearest to the Tertiary 
Emys levis; but the transverse dimension is not eked out, as in that 
species, by the accessory plastral plates, and the plastral portions of 
the marginal plates are respectively less broad in Plastremys, and 


UPPER GREENSAND AND CHLORITIC MARL. ort 


allow a large relative proportion of the plastron to the under- 
surface of the portable dwelling. 

The constituents of the plastron are in normal number, as in the 
typical Emydians; there are no accessory pieces interspersed 
between the hyo- and hypo-sternals, as in the Wealden genus 
Pleurosternon. 

The precise measurements, together with figures and minute 
description, will be published shortly. In the meantime the fact of 
the discovery of a freshwater Tortoise in the Upper Greensand is of 
interest, throwing additional light on the conditions of life in that 
epoch, and involving either the proximity of a continent or the 
remains of island faunas distributed in the marine deposits. 

The Crayfish obtained by Saxby, figured in Prof. Bell’s Mono- 
graph of the Crustacea, and the Chelonian found more recently by 
Mr. Norman, evidently prove that more thorough investigations 
among these chert rocks might result in further important additions 
to the Greensand fauna. 

The next 20 feet of compact red sand are for the most part 
unfossiliferous ; or rather the organic remains have been nearly 
obliterated. It is harder in texture than the succeeding 4 feet. 
Without any other lithological change than hardness, we suddenly 
come to a zone in which well-preserved casts of Mollusca abound, 
differing, as will be noted, from the Upper Cherts and Rags in a 
remarkable manner. At the base of this (that is, 20 feet from the 
lowest chert nodules) an Ammonite occurs, which, so far as I can 
discover, has not been described. Two specimens found by Capt. 
Ibbetson are now in the Jermyn-Street Museum, unnamed; a third 
is now in the Natural-History Museum, South Kensington, in 
my Isle-of-Wight collection, recently purchased by the Museum 
authorities. It has been placed provisionally by Mr. Etheridge as 
a species between A. auwritus, and A. rostratus ; though I think it 
will be found to differ from both sufficiently to rank as a new 
species. It is invariably of the same size, from 23 to 3 inches in 
diameter, and remarkably thin for its size, not having the pro- 
minent tubereles of A. auritus or the sharply defined chambers of 
A. rostratus. At the back the markings are alternate, more resem- 
bling the Gault species A. interruptus. Dr. Wright, of Cheltenham, 
was unable to identify my specimen with any species known to him. 

Immediately above, though never intermixed, the following 
fossils are found :— 


Ammunites auritus, Sow. 
rostratus, Sow. 
Trigonia aliformis, Park. 
T. ornata, D’ Orb. 
vicaryana, Lyc. 
Panopea plicata, Sow. 
mandibula, D’ Ord. 


Plicatula pectinoides, Sow. 
Ostrea canaliculata, D’ Orb. 


Pecten orbicularis, Sow. 


Cardium Gentianum, Sow. 


Area carinata, Sow. 
Cucullea glabra, Park. 
fibrosa, Sow. 

Natica sp. 

Cinulia sp. 

Vermicularia econeava, Sow. 
Rhynchonella latissima, Dav. 
Terebratula biplicata, Broch, 
Modiola sp. 

Serpula antiquata, Sow. 


aie C. PARKINSON ON THE ISLE-OF-WIGHT 


And suddenly as this horizon appears, so it disappears, the succeed- 
ing 32 feet being identical with the sands below in lithological cha- 
racter, with the exception of being less hard and gradually getting 
lighter in colour as we follow it up the section. Organic remains are 
few and far between, Holaster levis (on the authority of Dr. Wright, 
F.R.S.) being the only distinctive fossil. Up to this point the 
fauna may fairly be said to belong to the same epoch ; and collections 
from the lower division should be kept separate from those from the 
upper cherts and rags, which we now come to, and which indicate 
different conditions of life. The red sands give place to alternate 
bands of hard chert and coarse greensands (vide section). In the 
6 feet of inferior building-stone A. rostratus attains its greatest 
development, and casts of large Nautili are common. This is divided 
from the building-freestone (which hardens on exposure to the at- 
mosphere) by one foot of blue chert, having another chert band on 
the top little else but a mass of sponge-spicules imbedded in siliceous 
matrix. In the next bed of Greensand Pecten orbicularis is plentiful, 
though by no means confined to that band. Pecten cnterstriatus is 
not uncommon on the exposed surface of the succeeding Chert band, 
and species of Zima in other bands which could not be identified 
with certainty. The fossils may not all be confined to the various 
beds. Pecten asper, Lamk., never occurs in the Isle of Wight in 
these Cherts or Greensands; it is confined to a bed of phosphatic 
nodules in the Chloritic Marl, but is probably derived from older 
rocks, being in most cases much crushed and broken. 

The following fossils are from the Upper Chert and Rag beds :— 


Siphonia pyriformis, Park. Exogyra conica, Sow. 
Nautilus elegans, Sow. Ostrea frons, Park. 

sp. Lima aspera, Sow. 
Lima, large sp., strong ribs. Ostrea vesiculosa, Sow. 
Pecten, 14 ribs, (Galliennei ?). Cucullea glabra, Park. 
Cardium sp. Pecten orbicularis, Sow. 
Holaster sp. interstriatus, Lezon. 
Ammonites rostratus, Sow. — }-costatus, Sow. 
Clathraria Lyellii, Manz. sp. 


The Cycadeous plant Clathraria Lyell was first noted by 
Mantell, who obtained his specimen from a quarryman in Bon- 
church; it was not obtained from the Chalk Marl, but from the 
Greensand Rags. A second specimen of this rare fossil was found 
at Steephill, near Ventnor, last winter, 10 feet below the Chloritic 
Marl, in the same bed as the former one. The remains consist of the 
top of the stem surrounded by petioles or leaf-stalks, the leaves them- 
selves being either shed or decayed. It has usually been considered 
a plant of Wealden age. If, however, island floras and faunas can be 
shown to have existed during the formation of the Greensand strata, 
oth Chelonia and remains of plant-life are at once accounted for. 

The Chloritic or Glauconitic Marl, lying immediately over the 
Greensand, may be traced along the escarpment of the Undercliff, 
from Blackgang to Luccombe, cropping out finally near Culver Chiff. 
It varies in thickness from 6 to 7 feet, and may be divided into two 


UPPER GREENSAND AND CHLORITIC MARL. 373 


beds, the lower 23 feet, with few fossils, divided by a layer of phos- 
phatic nodules (with broken Pecten asper) from the upper 33 feet, 
which are rich in fossil remains. The best sections are at Ventnor- 
station quarries and below Old Park, St. Lawrence. 


A. Fossils derived, in phosphate. 


Ammonites varians, Sow. Solarium ornatum, Sow. 
Pleurotomaria sp. Pecten asper, Lamk. 
Hamites sp. Venus sp. 

A. curyatus, Sow. | Cinulia sp. 


Rynchonella Grasiana, D’ Orb. Gibbula levistriata, Seel. 


B. Fossils slightly phosphatic. 


Ammonites Coupei, Brongn. Emarginula sp. 

varians, Sow. Terebratulina striata, Wahl. 
—— Mantelli, Sow. biplicata, Sow. 
navicularis, Mant. semiglobosa, Sow. 


eurvatus, Mant. Plicatula pectinoides, Sow. 
sp. Rbynchonella Grasiana, D’ Orb. 
Hamites attenuatus, Sow. Pecten orbicularis, Sow. 
Scaphites zqualis, Sow. (very Beaveri, Sow. 
rare). interstriatus, Leym. 
Nautilus elegans, Sow. Terebratula pectita, Sow. 
expansus, Sow. Discoidea subuculus, Leske. 
Turrilites Morrisii, Sharpe. Micrabacia coronula, Goldf. 
tuberculatus, Bose. Cidaris vesiculosa, Goldf. 
—— Wiestii, Sharpe, Vermicularia concava, Sow. 


This Chloritic or Glauconitic Marl has caused considerable con- 
troversy as to its origin. I think the presence of phosphatic nodules 
between the two beds, together with crushed Pecten asper in great 
numbers, has not been sufficiently noticed. This would suggest a 
violent and sudden destruction in the phosphate bed, while in the 
upper 32 feet nearly all the Mollusca remain fairly periect, and 
Pecten asper is not found. 

Mr. A.J. Jukes-Browne suggested the name of “ zone of Scaphites 
equalis”’ for this marl ; but as it is extremely rare to find a specimen 
of this Scaphites lower than the Chalk Marl, I would propose ‘zone 
of Turrilites Morrisii,” which is peculiar to the Chloritic Marl. 

The best sections and measurements of the Isle-of-Wight Upper 
Greensands were made as far back as 1849 by the late Capt. Ibbetsor. 
These, together with measurements lately made by Dr. C. Barrois, of 
Lille, I prefix to my own section. 


Capt. Isprrtson. Coes a 

1. Zone of chert and rag with P. orbicularis, P. 5-costatus ... 15 O 
Pepurlomerate OF chert and Tag  ........-.0:..sseseecnneeceosonee 40 
PEE AMM HECSLONC' (5.05... )-ccoctccecacaseccedseecosscacsccencaccees Bye 
pempbbecensoneameulayers Of TAG... 2.2... ieee ee ce eeet vows eteewees [kG 
2 EE ibd, ela 0 eae ee is ek ee a ee 16:8 
6. Mammillary rag, sandy boulders with phosphate of lime.. 1 6 
“2, (LOL 922d) so AG Reese coe 6 8 
PEP MEAS THANE 2 52 css cs- sec co-escneccccnscecceesseesontevees 3 0 
2. DLE 200 i a a eee 40 0 
103 6 


2 3.G.S. No.147. 2Q¢ 


374. C. PARKINSON ON THE ISLE-OF-WIGHT 


Dr. Barrols. ft. 


: in. 
= i) Chioritie marl yin case eee 8 0 
<3 D Greensand andichert) .-.ns-.ce.s--cceesesee oe 26 0 
- |C Greensand, phosphatic nodules ............ 6 6 

A 

g 
8 es Yellow sands with bands.................065. 13 0 
= A Micaceous, glauconite sands ............... 117 0 

N 
170 6 


(The order of this section is reversed for uniformity.) 


Section of Upper Greensand and Chioritic Marl, St. Lawrence 
and Ventnor, Isle of Wight. 


{ Chloritic marl 33 feet (fossils list B, slightly phos- 
| phatic). 
G feet. nee cence Hard phosphatic nodules, with crushed Pecten asper. 
| 23 feet compact, darker grains: few fossils, mostly 
\ derived. 
( Blue chert bands ; casts of Cardium sp. 
Coarse greensand ; ; casts of Cucullea glabra, and 
| Holaster sp. Chert. 
94 feet Greensand: Ostrea, Cucullea, Arca. 
“| } —_—___| Chert: Lima sp. 
| Chert : Pecten interstriatus. 
Greensands: P. orbicularis. 
—-—] Chert with sponge-spicula imbedded 
(————-| 5 feet freestone, used for building-purposes: casts of 
| Nautilus pseudoelegans, N. elegans. 
Chert, with phosphatic nodules. 
14 feet... 1 6 feet inferior building-stone: casts of Nauwzilz. 
| Amm. rostratus here attains its greatest development. 
; ————| Blue chert. 
| 32 feet yellowish-red sandstones, varying in hardness. 
Organic remains few. 
Holaster levis (on the authority of Dr. Wright). 
| .| Zone of Amm. inflatus, Sow. A. awritus, A. rostratus, 
5G feet d Panopea, Cucullea, Arca, Trigonia. About 4 feet. 
“| ) —____| Compact red sands. 
—-—| Species of Ammonites undetermined, between A, rostra- 
tus and A. awritus. 
Compact red rock, harder than the above, 20 feet, 
unfossiliferous. 
D2 feeb. le eewset Nace Blue rag, fossiliferous. Astaciform Crustacea, Che- 
—__- —— lonia; passing into Gault sands. 
102 feet... 


It will be found that the three measurements agree very nearly. 
Dr. Barrois includes the 50 feet of sands below the lowest two-foot 
bed of chert nodules, as has been done in the Geological Survey 
Memoir. In Dr. Barrois’s hasty visit to the Isle of Wight, I think 
he inaccurately divided the zone of P. asper and A. inflatus, both of 
which are confined to extremely narrow limits. With regard to 
Capt. Ibbetson’s measurements, I think the zone of fossiliferous 
malm is placed too high up; otherwise the section is very accurate. 

In conclusion, I would call attention to the necessity for greater 
care in separating the fossils of each zone. Thus Pecten asper, P. 


UPPER GREENSAND AND CHLORITIC MARL. SMe) 


eretosus, P. Beaveri, Lima sp., P. Gallienner(?) may eventually 
be found to characterize different zones, while the entire fauna of 
the malm rock is different from that of the chert and rag beds. 

I would propose, also, that the 50 feet of sands included in the 
Upper Greensand by Mr. Bristow, and more recently by Dr. Barrois, 
should be referred to the Gault, taking the well-marked band of 
chert boulders with Chelonia &c. as the true base of the Upper 
Greensand. 

I also suggest, with some diffidence, that remains of island floras 
and faunas exist in the marine deposits of those Greensands, as 
evidenced by a freshwater Tortoise and Cycadeous plants, the former 
authoritatively stated by Prof. Owen to be a freshwater reptile, 
while the leaf-stalks and stems of tropical plants speak for them- 
selves. 

Dr. Barrois’s division of the Upper Greensand into two zones 
is misleading, as the fossils are confined to such narrow limits. I 
would suggest, in place of this, several zones in the upper cherts 
identified by characteristic Pectinide, which are the best-preserved 
of the Mollusca, the fauna of the Malm rock being clearly separated 
from that of the Rag. 

Mr. J ukes-Browne’s zone of Scaphites equals is imaccurate. 
20 feet higher up in the Chalk Marl S. equalis is abundant; in the 
Chloritic it is hardly ever found. Turrilites Morrisii is, on the 
other hand, characteristic of this horizon. 

The band of phosphatic nodules with crushed Pecten asper de- 
serves attention. ‘This species of Pecten is certainly not properly 
a Chloritic-Marl fossil; yet 1t only occurs in the peculiarly crushed 
state in this formation; it is never found in the Greensand proper 
in the Isle of Wight. 


Discussion. 


Mr. H. G. Forpuam stated that between Ballard Hole and Pun- 
field Cove the phosphatic nodules are scattered through the Chloritic 
Marl, and not confined to its base, and that no broken shells of 
Pecten asper occur at that point. He further remarked that the 
author gave the thickness of the Chloritic Marl as 6 feet, 7. ¢. greater 
than previous authors. 

The Presipent stated that the new Ammonite mentioned by the 
author seemed to be intermediate between A. rostratus and A. 
awritus, but nearer to the former. Pecten asper had not previously 
been noticed above the Upper Greensand. He referred to the great 
yalue of Dr. Barrois’s labours in connexion with the English Chalk. 

Rev. J. F. Braxe thought that the beds with Ammonites inflatus 
belonged to the Upper Gault, and not to the Greensand. 

Mr. J. Srarxte GarpneR thought that, as species of Pecten and 
Lima have often a long range, their value for characterizing horizons 
in the Cretaceous was doubtful. 

Mr. De Rance stated that Pecten asper in Dorsetshire never 
occurs above the Greensand. He agreed with Mr. Blak» in re- 


garding the zone of Ammonites wnflatus as Upper Gault. 
Zc 2 


376 E. W. WILLETT ON A MAMMALIAN JAW FROM 


28. Norss on a Mammarian Jaw from the PurBeck Bups at Swanace, 
Dorset. By Evear W. Wituerr, B.A. With an Introduction 
by Henry Witxert, Esq., F.G.8. (Read May 25, 1881.) 


(Communicated by the President.) 


Inrropuctory Remarks. By Henry Wittert, Esq., F.GS. 


The rarity of the fossil remains of mammals in Mesozoic strata 
is a paleontological fact so well known to the Members of thig 
Society, that it would be superfluous in me to do more than 
allude to it. 

All needful information will be found in the exhaustive. mono- 
graph (by one of the greatest living authorities on comparative 
anatomy, Prof. Owen) published in 1871 by the Paleontographical 
Society. From it I gather that in 1828 Mr. Broderip first dis- 
covered Didelphys in the Stonesfield-Slate quarry. 

In 1858 the teeth of Mrcrolestes were discovered by Mr. Moore 
in a breccia of Rheetic bone-bed and limestone filling a fissure in 
the Mountain Limestone at Frome, in Somersetshire. 

In 1864 my friend Prof. Dawkins discovered a worn molar of a 
Marsupial mammal in the Rheetic beds at Watchet, in Somerset- 
shire, called by him Hypsiprimnopsis rheticus (Microlestes rhe- 


ticus). But it was to the personal energy and perseverance of _ 


Samuel Beckles, Esq., F.R.S., that science was indebted for the 
discovery of the great variety of these interesting fossils, the de- 
scription of which occupies the largest portion of Prof. Owen’s 
work (loc. cit.). My own imagination was excited by a popular 
account of Mr. Beckles’s labours, written about 1858 by the late 
lamented Canon Kingsley; and the desire for further discovery 
caused me to pay several visits to the ‘‘ Dirt-beds” at Swanage, in 
the hope of making further additions to the catalogue of Mam- 
malian remains. fuddven I must have spent slbnee ther many 
hours in search on several oecasions, aided at times by local quarry- 
men, I could never succeed; but in 1878, having learnt that. 
Mr. Beckles had been obliged, unwillingly and prematurely, to 
abandon his researches at a posits which seemed rich in promise, I 
obtained leave from the Earl of Eldon, on certain reasonable con- 
ditions, to renew the inquiry at the point left by Mr. Beckles. The 
year 1879. was too wet and stormy to allow me to carry out my 
intention in so perilous a position. 

There are two so-called “ Dirt-beds ” in the Purbecks of Durdle- 
ston Bay, Swanage. The lower one can be readily examined from 
the shore, as it rises at varying angles from beneath the sea-level 
until it is lost in the débris and turf above. 

The upper dirt-bed, which also crops out at a similar angle, is 
less distinctly defined, until we reach a point about two thirds of 
the way up the cliff, just below the commodious refreshment-room 


THE PURBECK BEDS AT SWANAGE, DORSET. Ta 


erected on the summit of the cliff; and it was at this point (at 
which Mr. Beckles, hoping to resume his labours, had purposely cast 
down a large protecting accumulation of debris) that our attack 
was finally made ; in the summer of last year, 1880. 

I must here render my tribute of thanks to my able coadjutor, 
Mr. Henry Keeping, of Cambridge, for the skill with which he 
directed the work, at no. little peril to those engaged in it. 

We commenced operations by scarping down the overhanging 
strata for a depth of 40 feet, laying bare an area about 13 feet by 
10. This upper “ dirt-bed” is of varying thickness, from 2 to 
10 inches. It seems to have been a silt, filing up hollows and irre- 
eularities in the surface of the stratum immediately below it. 

Aided by those earnest geologists Prof. Dawkins of Manchester, 
Mr. Charles Potter of Liverpool, Mr. Griffith of Cambridge, and 
several members of my own family, the area was most carefully 
broken up and examined. The research occupied ten days; and 
although we found several teeth and jaws (hereafter to be described), 
it was a fortuitous blow of the hammer of a local quarry-man that 
laid bare the interesting specimen about to be described. 

The-time of the meeting has already been trespassed upon too 
long by these preliminary remarks. I hope on some future occa- 
sion to supplement them by a further description of the fossils dis- 
covered, a more ample account of the probable conditions under 
which the deposits were formed, and the reasons which explain 
. the rarity of discovery rather than the paucity in number of the 
Mammalian remains. 

With the jaw of Triconodon, and in the same bed, were found 

rocodilian remains (Zheriosuchus pusillus, Ni annosuchus g gracilidens, 
and Nuthetes destructor), with other Mammalian and Reptilan 
fragments not easily determinable. There appears to be a single 
tooth of Theriosuchus amongst the fragments, but whether of 7’. 
pusilus is doubtful. 


Ir will be remembered that rather more than twenty years ago 
extensive explorations were undertaken by Mr. Beckles, F.R.S., at 
Swanage, in search of Mammalian remains, and that he succeeded 
in unearthing some dozen new genera, including altogether sixteen 
new species of Mesozoic mammals. These fossils consisted prin- 
cipally of mandibles more or less broken, the only other bones 
found being portions of the upper jaw. 

In the early summer of this year the permission of. Lord Eldon 
was obtained to renew the search in Durdleston Bay, Swanage ; and 
although this search was not followed by such brilliant results as 
in Mr. Beckles’s case, it cannot be said to have been to no purpose, 
since a very good Mammalian jaw was obtained. 

The specimen consists of the larger part of a right mandibular 
ramus, of which the condyle, the upper border of the coronoid 
process, and the symphysial end anterior to the second premolar 
are wanting. Six teeth altogether are preserved in svtu, and, with 


378 E. W. WILLETT ON A MAMMALIAN JAW FROM 


the exception of the foremost, one are quite perfect. The genus to 
which the specimen belongs is Zriconodon, Owen; but it differs 


Fig. Lower jaw of Triconodon mordax, Owen, from Swanage. 
Nat. size. 


from those described by Prof. Owen in his Monograph on the 
Mesozoic Mammalia* in the fact that it has four teeth having the 
form of true molars, all those previously found (eleven or twelve in 
number) having each only three true molars, ‘a reduction rare in 
the Marsupial order,” to which these mammals are usually ascribed. 
It may be noticed here that the nearly allied genus T'riacanthodon 
has four true molars, but that the teeth now under consideration 
differ from those of this last-named genus in the following im- 
portant details :—The fourth premolar of Triacanthodon approaches 
the triconodont or true molar type; and the apex of this tooth 
reaches only to half the height of the main cone of the preceding 
premolar, whereas the main cone of the fourth premolar of the 
new jaw is by far the largest of the three cones, and is rather 
longer than the corresponding cone of the preceding tooth ; again, 
no hinder talon is found in any of the four molars of Trzacanthodon, 
while it is well marked here on each tooth. 

Of the six teeth preserved, four are, judging by form, true molars, 
and two are premolars, each of which is implanted by two roots; 
the two sockets for the next premolar are also plainly visible. 

The crowns of both the premolars consist of a principal sub- 
compressed cone, with a small and low anterior basal cusp, and a 
large and higher posterior one followed by a rudimentary talon ; 
of the two the anterior tooth is rather the smaller. 

All the true molars agree with the description .of the type 
specimen of Prof. Owen in being ‘“ subcompressed, antero-pos- 
teriorly extended, and divided into three nearly equal cones in the 
same longitudinal line, the middle cone being very little larger than 
the front or hind cone; further, there is no cingulum on the outer 
side of the crown, but at the posterior margin of the posterior cone a 
rudimentary talon is feebly marked off by a short vertical indent 
from the rest of the surface of that cone.” The first of the true 


* «Monograph of the Fossil Mammalia of the Mesozoic Formations,’ by 
Prof. Owen, F.R.S. D.C Ll. Printed for the Palzontographical Society. 
London 1871 


THE PURBECK BEDS AT SWANAGE, DORSET. 379 


molars is rather smaller than the others; and, as in the type speci- 
men alluded to above (plate iii. figs. 7, 7 4, loc. cit.), the base of 
the coronoid process hides part of the last tooth. Prof. Owen con- 
siders that this character may indicate that the jaw described by 
him belonged to an individual not quite fully grown. Assuming 
such to be the case, the fourth molar in the present specimen would 
point to a mature animal, though, for a similar reason, to one not 
yet adult. In point of size, the last premolar and the three anterior 
molars agree as nearly as possible with the corresponding teeth in 
the type specimen of 7’riconodon mordax. 

The question, therefore, to be settled is, Does the new specimen 
belong to this last-mentioned genus and species or not? 

Prot. Flower, who has kindly examined it, and compared it with 
those found by Mr. Beckles, which are now in the British Museum, 
thinks, on the whole, that it may probably be referred to this species, 
and ingeniously suggests two hypotheses to account for the extra 
tooth of true molar form. ‘They are as follows :— 

As is well known, the deciduous teeth of the Marsupials, to 
which group the Mesozoic mammals have hitherto been assigned, 
consist of a single tooth on either side, the tooth which replaces 
this deciduous one being the last premolar. Assuming, then, that 
this tooth is still in setu, and is represented by the third tooth, 
counting from the symphysis, the dental formula will bep.m. 4 
m. 3, which agrees with the type specimen of T'riconodon mordax 
in the British Museum ; and the first hypothesis is that the jaw 
belonged to a younger individual than any previously found, with 
the single milk-tooth in position. 

The second hypothesis is to assume that all the teeth preserved 
belonged to the permanent set, when the dental formula will be 
p-m.3,m.4. In this case the four molars agree in number with 
those found in the adults of all recent Marsupials, and indicate a 
more fully matured specimen than any hitherto discovered. 

A third hypothesis is to assume that it belongs to an altogether 
distinct species; but this, considering the close resemblance to 
Triconodon mordax, appears hardly necessary. 

The lower border of the ramus is, in its present condition, nearly 
straight; but it has been much crushed, and when recent was, with- 
out doubt, slightly curved. 

A small outlet of the dental canal opens under the foremost root 
of the third premolar (the first which is preserved); and there are 
traces of one, if not of two, other outlets anterior to this. 

The total length of the jaw is 13 inch. 


Discussion. 


Mr. H. Witterr said that he hoped the specimen would be accepted 
by the authorities of the School of Mines, and placed in the collec- 
tion there. 

Prof. Duncan spoke of the importance of the communication and 
the thoroughness which it showed. The jaw was a highly developed 


380 ON A MAMMALIAN JAW FROM SWANAGE, DORSET. 


one; and had not the marsupial nature been suggested, it might 
have been considered to belong to the Insectivora ; probably, how- 
ever, it was an insectivorous marsupial. But was it so certain that 
the jaw belonged to a marsupial of the present type? It was 
strange also that these remains of marsupials had, at present, only 
been found in Europe. He was under the impression that the 
Australian marsupial fauna was not very ancient. 

Mr. Cuartesworte said the history of the discovery of mammalian 
life in Britain was interesting. It had a important bearing on 
evolution. The explanation of the occurrence of lower jaws only was 
well known; but it was difficult to explain the disappearance of the 
phragmocones of Belemnites: also in Kast Anglia only teeth, antlers, 
and astragali of Cervz were found ; other bones were wanting. 

Mr. E. Witter said that mammalian remains had been recently 
found in great numbers in America, on about the same horizon, and 
that from them the discoverer, Prof. Marsh, was led to believe that 
all Mesozoic mammals belonged to a more generalized type than do 
the marsupials which at present exist. 


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Quart, Journ. Geol. Soc Vel XXXVI PLXIX. 


A.S Foord lth JURASSIC DIATOPORIDAS 


G. R. VINE ON THE FAMILY DIASTOPORID&. 381 


29. Further Notes on the Family Diastovoripe, Busk. Species from 
the Lias and Oottrr. By Grorer Rozert Vine, Esq. (Com- 
municated by Prof. P. Marrmy Duncan, M.B. Lond., F.R.S., 
F.G.S.) (Read January 19, 1881.) 


[Puate XTX.] 


Since my first paper, ‘‘A Review of the Family Diastoporide for 
the purpose of Classification” *, was written, a very important book 
has been published, namely ‘A History of British Marine Polyzoa,’ 
by the Rev. Thomas Hincks?. In this work the classification 
adopted by Prof. Busk in his Monograph of the Fossil Polyzoa of 
the Crag, 1859, and also in his british-Museum Catalogue, 
part ii. Cyclostomata, 1875, is set aside for one that J have little 
hesitation in saying will prove to be far more valuable to the 
working student. Instead of separating Diastopora, and making 
it the type of a family, Mr. Hincks places this genus with others in 
his second group of the Cyclostomata, which he calls Incrustata, 
DOrb. The family name which he adopts is Fam. IL. Tubuliporide, 
which includes the genera 


Stomatopora, Bronn. | Entalophora, Lamouroux. 
Tubualipora, Lamarck. Diastopora, Lamouroux (pt.). 
Idmonea, Lamourouc. 


Seeing that I have already committed myself to Prof. Busk’s 
arrangement, it is impossible for me now to retreat; and, with this 
explanation, | must be excused for still adhering to the famuy name 
I used at first ?. 

In his remarks on the Cyclostomata, Mr. Hincks says :—‘ Sim- 
plicity isin the highest degree characteristic of the group; the cells 
are universally tubular; the polypide is without complexity of 
structure, and has a small number of tentacles; all appendicular 
organs are wanting” §. In another place he says:—‘ In classi- 
fying the Cyclostomata we have to base our divisions mainly on 
habit or mode of growth, on the plan according to which the 
zocecia are aggregated together into colonies; the simplicity and 
general similarity of the cell throughout the tribe leave no other 
course open to us. We have to deal with very uniform struc- 
tural elements very variously combined; and the modes of com- 
bination chiefly supply us with the bases of our system. Under 


* Quart. Journ. Geol. Soc. vol. xxxvi. p. 856 (August 1880). 

Tt Van Voorst: 1880. 

t Of the two evils I choose the least. To call my paper “ Further Notes on 
the Tubuliporide” would be to commit myself to remarks on the other genera, 
which at present I have no intention of doing; and the retention of the family 
name Diastoporidz, Busk, may have its special advantages. 

§ Brit. Marine Polyzoa, Introduction, p. exxy. 


382 G. R. VINE ON THE FAMILY DIASTOPORIDZ. 


these circumstances we may not expcct very strongly marked 
boundary lines” *. 

The family Tubuliporide, Hincks, is thus defined :—“‘ Zoarium 
entirely adherent, or more or less free and erect, multiform, often 
linear, or flabellate, or lobate, sometimes cylindrical ; Zowcia tubu- 
lar, disposed in contiguous series, or in single lines; Owciwm an 
inflation of the surface of the zoarium at certain points, or a 
modified cell” +. 

These further Notes have reference to the Diastopore found in 
the English Lias and Oolite, and are the results of a very careful 
study of material kindly and liberally placed at my disposal by two 
good local workers, Mr. KE. A. Walford, of Banbury, and Mr. F. D. 
Longe, F.G.8., of Cheltenham. Whilst I was engaged in the study 

was continually perplexed with the ever-varying modes of growth 
of what may be considered to be true Diastopore. ‘To fix many of 
these forms under the descriptions already given by authors is 
indeed impossible; or to call them, arbitrarily, ‘ species” or 
‘“‘ varieties ” would give a false idea of their significance. The 
wisest course to adopt is to call them ‘‘types;” and in giving them 
specific names I have kept this suggestion constantly before me. 
Between the Liassic and Oolitic forms very little variation is per- 
ceptible if we select groups of the same or similar habit. If, as in 
the Inferior Oolite, for instance, we take the three or four different 
types, and try to correlate them under one specific name, the varietal 
or typical divergency is at once apparent. In this paper I have 
directed the attention of the paleontologist more particularly to 
these typical forms; and it remains now for local workers to mark 
the differences well, and then, by a rigid and philosophical scrutiny, 
to try and ascertain whether any of these varietal types creep gra- 
dually from one into the other. The doing of this as it ought to be 
done depends on whether material for the scrutiny is accessible to 
the student ; and besides the material, patience is needed to follow 
out a set design to its conclusion. To take away the stigma so con- 
tinually repeated to our disadvantage, “ the imperfection of the 
geological record,” work done in the direction indicated must be 
attempted, even if it be imperfect; but, after going over only a 
small portion of the labours of two local workers, I believe it to be 
possible to carry out the design to perfection, or nearly so, if keen 
eyes and willing hands are engaged in the task. 

I am not aware that previously to the labours of Prof. Quenstedt 
the attention of the paleontologist had ever been directed to Liassic 
Polyzoa. In his great work, ‘Der Jura,’ many Liassic fossils are 
described and figured, and amongst others we have figured and 
described the earliest known Mesozoic Diastopora. Quenstedt 
names it D.lasica; and he says of it, “ One usually recogn 
the primary cell; at first a rapid increase occurs; but the poly- 
zoarlium soon divides itself into two groups, draws itself back, and 


* Brit. Mar. Polyzoa, vol. i. p. 425. 
t Ubid. vol. 1. p: 424 


G. R. VINE ON THE FAMILY DIASTOPORID®. 383 


completes a perfect circle” *. He further says, D. liasica “ occurs 
so abundantly, and precisely on shells of the Jurensis zone, that you 
can often identify your horizon from it.” ‘The specimen he figures 
is found on Ammonites jurensis. 

Jules Haime‘, in correlating Berenicea striata, says that this 
species was discovered by M. Terquem in the Lias of Valiére. He 
says :—‘ D. lrasica, Quenstedt, Handb. der Petrefact. p. 637, pl. lvi. 
fig. 10, “is closely related to this species: it begins in the same 
manner with a plate in the shape of a fan; but it branches out more 
on the outside.”’ 

Dumortier, in his ‘ Paleontological Studies of the Jurassic De- 
posits of the Basin of the Rhone,’ recognizes another species, as he 
calls it, in the fourth part of the Superior Lias. This he names 
D. crussolensis ; and the only distinguishing feature between this 
and D. liasica is *‘ that it grows thicker than the last, and the form 
of the colony is more circular” $. 

In Mr. E. A. Walford’s pamphlet on some Upper and Middle Lias 
beds in the neighbourhood of Banbury §, the author relies upon 
Quenstedt’s description for the identification of his specimens found 
in the zone of Amm. spinatus, and also in the transition-bed, on 
corals and shells. 

In the best work on Oolitic Polyzoa that I have yet met with, 
‘Description of the Fossil Bryozoa of the Jurassic Formation’ ||, 
Jules Haime divides his typical Diastoporide ito two groups, the 
Berenicece and the Diastopore. The incrusting forms treated of as 
Diastopore in this paper belong to the first group of Haime. His 
species are:—B. diluviana, Lam., a common form of the Great 
Oolite; B. Archiaci, Haime; B. microstoma, Mich.; and B. lucensis, 
Haime. Both of these last are found in the Bradford Clay and the 
Great Oolite, very beautiful species with very characteristic cells. 
The Diastopore of Haime belong to the Foliaceous group. Another 
paper, by Prof. D. Brauns{[, contains some very valuable in- 
formation on both the foliaceous (especially lea) and incrusting 
Diastoporide. 

Mr. Walford has placed in my hands, for description or study, the 
whole of his local fossil Polyzoa; and so inadequately have the 
species been described and figured, that, for scientific purposes, the 
labours of these eminent paleontologists are almost useless. In 
giving the Liassic species a new name, I have had regard more 
particularly to its typical character than to any thing else. In this 
Liassic type I recognize a family likeness to later fossil and more 
recent Stomatopore ; and it will be advantageous to science to draw 
attention to the fact. 


* “Der Jura,’ pp. 279-292, fig. 1, tab. 40. 

t+ Bryoz. fossiles de la form. Jurass., in Mém. Soc. Géol. de France, 1854. 

{ Dumortier, /. c. p. 226, pl. 48. figs. 11, 12. 

§ Proceedings of the Warwickshire Naturalists’ and Archeological Club, 1878. 
| Loe. cit. 

{| Zeitschr. d. deutschen geolog. Gesellsch. 1879. 


384 G. R. VINE ON THE FAMILY DIASTOPORIDA. 


1. Drastopora stoMatToporiIpEs, mihi. Plate XIX. figs. 1-10. 


? D. liasica, Quenstedt, Handb. der Petref. p. 637, pl. lvi. fig. 10. 

? D. crussolensis, Dumortier, Paleont. Studies, p. 226, pl. xlviii. 
figs. 10 & 12. 

Zoariwum subcircular or orbicular, sometimes indefinite in its mode 
of growth, forming small isolated patches on shells or corals, varying 
in breadth from one and a half to three lines. Zowcia arranged 
linearly, or nearly so, long and slender tubes, many of them 
wrinkled or surface-roughened, and adherent by their whole length ; 
orifice, when perfect, oval, rarely circular. Primary zocecia either 
very excentric in the larger colonies, or proximal in the smaller 
ones, Which soon become excentric as the colonial growth increases. 
Oeca rare, when present pyriform, involving at lees two of the 
cells. Zocecial tubes very faintly punctate. 

Hab. On Amm. cornucopie, Up. Lias, Bloxham; on Cardinia 
hybrida, Sow., Appleton ; and on fornia Victoria, Mid. Lias, 
zone of Amm. Henleyr, Cherrington, Oxfordshire ; Sup. Lias, Crussol, 
Dumortier ; on Amm. jurensis, in zone of ditto, Southern Germany, 
Quenstedt. ‘ Deeper than this,” says Quenstedt, “ I have never 
found it.”’ 

A careful study of the figures given will convey to the paleeon- 
tologist a very fair idea of the character of this very early Mesozoic 
type of Diastopora. Unlike any of the palzeozoic types, it seems to 
be persistent, so far as the character of the cell is to be relied upon, 
high up into the Oolitic series, and, but for the peculiarity of its 
maar, might be recognized in the Stomatepora deastoporides, Norman, — 
and the Tubulipora lobulata, Hassall*. Indeed, of the first of these 
species Mr. Hincks says “ that it is the largest of British Stoma- 
topore, and has very much the look of Diastopora;” and of the 
other species, ‘“‘I can see no sufficient ground for placing Tubuli- 
pora and Diastopora in separate families; the two genera are nearly 
related, and have many common characters.” He said this without 
being aware of the existence of the forms now figured and described, 
which are in every sense confirmatory of the justness of his family 
arrangement. 


In the Oolitic series, beginning with the lowest beds—the Pea- 
we recognize altogether different types, 
not widely separated, but even on the same blocks. ‘These types 
belong to the foliaceous as well as to the crustaceous forms; and 
where to draw the line between the two it is difficult to say. In 
some few cases the boundary lines are broken down; and one at least 
of the typical Diastopore pass from the crustaceous into the foli- 
aceous form by a series of quiet gradations. Mr. Longe has given 
more attention to these forms than I have; and his remarks on these 
peculiar species may be referred to for exact informationy. After 


* Figures and description in Hincks’s Brit. Polyzoa, vol. i. pp. 434 & 444, 
pls. 61 & 63. 
t Geological Magazine, January 1881. 


G. R. VINE ON THE FAMILY DIASTOPORID®. 385 


a careful study of the crustaceous forms from several horizons, 
ranging from the Pea-Grit to the Coral Rag, I can detect at least 
three types that have not been specially noticed, so far as I am 
aware, by previous authors. ‘hey may, and I have not the least 
doubt that they have. been, casually identified; but that is all. 
They deserve, however, more than a passing notice, because some of 
these types, when they pass into the Bradford Clay and the Forest 
Marble, delicately preserved on branches of Terebellaria and frag- 
ments of broken shell, assume an altogether different character. 
Some of them are beautifully papyraceous ; others appear to have a 
kind of basal lamina extending slightly beyond their circumference, 
a character I have never observed in the Inferior-Oolite species. 

The papyraceous species of the Lower Oolite are also deserving 
of closer study than I can possibly give to them. Such work belongs 
rather to local students than to me. They, too, may study the 
types as they pass from one stratum to another; and in so doing, I 
would recommend them to mark the beginnings of the divergences, 
and the boundary lines of each of the four types given; and by 
doing so they will aid the paleontologist in classifying the evolu- 
tionary stages of a most important genus. 


2. DrastopoRaA VENTRICOSA, mihi. Plate XIX. figs. 15-17. 


Zoarium adnate, discoid in the earlier stages of growth, of most 
irregular outline in its later stages. Zowcia produced and partially 
free in the centre, gradually depressed towards the margin; tubes 
slightly bent and swelling towards the orifice, which causes a con- 
striction of the circular or subcircular mouth; cells well separated, 
the proximal ends heing immersed in the zoarium. Occia very 
largely developed, sometimes round the margin, at other times indis- 
eriminately all over the colony, involving two or three cells or only 
a considerable swelling of a single tube. In the best-preserved spe- 
cimens the cells and also portions of the ocecia are finely punctate. 

Hab. Ona weathered and partially smoothed pebble (No. 5), 
Inferior Oolite, Pea-Grit, Cheltenham: Mr. Longe’s Cabinet. On 
drift wood, Chipping Norton, lowest beds, Great Oolite: Mr. 
Walford’s cabinet. Good specimens also in Museum of Practical 
Geology, Jermyn Street. 

This species, or type, is a very peculiar one, well deserving especial 
study. I have it from several localitics, ranging from the Pea-Grit 
to the Great Oolite. The specimen in Mr. Walford’s cabinet 
contains innumerable colonies piled up very irregularly round a 
piece of water-logged coniferous wood of Oolitic age.. The wood 
was originally large; but the broken fragment submitted to me for 
examination was about three inches long, and from half to three 
quarters of an inch in diameter. The incrustation of the wood, 
made up wholly of colonies of Diastopora, varies in thickness from 
a quarter to half an inch, the margins of the newer colonies 
eradually becoming compressed into the general mass through 
successive stages of growth. The ventricose swellings are not so 
typical in this specimen as in the more beautiful specimens from 


386 G. R. VINE ON THE FAMILY DIASTOPORIDA. 


the Pea-Grit series (Mr. Longe’s cabinet). One colony is five lines 
by four; and the ocecia are well developed, for there are no fewer 
than from 25 to 27 swellings in various stages of development ; and 
it is from this specimen that the drawings (Plate XIX.) are taken. 
Measured across the cell-mouths, there are about six (or varying 
from five to seven) cells to a line. Three ocecia occupy about the 
same space. ‘The earliest stages of colonial growth are disks with 
free cells, flabelliform at later stages, ultimately presenting the 
appearance of one continuous mass of immersed cells. On the 
broken edge of a fragment of Mr. Walford’s specimen I can count 
from twenty to thirty layers, representing successive colonial 
growths. 

It may be that some at least of the specimens of this type are the 
Berenicea diluviana of authors, and the Diastopora verrucosa of 
Milne-Edwards. Berenicea Archiaci, Haime, is closely related to 
this species ; but the cells, and also the occia as figured by Haime*, 
are not so characteristic of our own Oolitic series as D. veniricosa. 


3. DrIAsToPoRA ooLiTicaA, mihi. Plate XIX. figs. 11-14. 


Zoarium circular or nearly so, completely adnate, and varying in 
size from one to three lines in diameter either way. Zocwcia short 
and bulging near the distal, gradually contracting towards the 
proximal end ; orifice variously shaped, the lower margin, in some 
cases, slightly mucronate (?); primary cell excentric. Central 
zocecia partially raised, becoming gradually depressed towards the 
margin. Oacia? Tubes faintly punctured, and no “ adventitious 
tubules.” 

Range from the Pea-Grit to the Great Oolite. 

Cabinets: (from several localities) my own; Miss Gatty’s, Kidling- 
ton, Oxon; Mr. Longe’s, very rich, from Pea-Grit; and also from 
Mr. Walford’s ; Museum of Practical Geology (several specimens on 
shells). 

Pa On stones and shells, forming small disk-like patches, more 
frequently isolated than clustering. 

This delicate little species is the most abundant of the Oolitic 
Diastopore. From its peculiar habit specimens are generally referred 
by collectors to the D. obelia of Busk; and in my early identifica- 
tions I was inclined to place it as a variety (var. oolztica) of that 
species. After drawing and carefully working out the type I soon 
found this to be impossible. The general habit is different ; the cells 
are more closely packed, and their shapes are altogether different ; 
and, above all, there is in none of the specimens I have examined 
any indication of ‘‘ adventitious tubules.” Judging from Manzoni’ 
figure t, which he refers to Berenicea striata, J. Haimet, a doubt na- 
turally suggested itself when correcting the proof of my first paper$ 
as to whether some specimens of this type may be referred to 
Haime’s species. There seems to be no identity either with that or 


* Bryozoa Jurassic Form. pl. ix. fig. 11. 

+ Fig. 79, Bryozoa of the Pliocene of Castrocaro. 

¢ Reuss, Die Bryoz. des braunen Jura von Balin, &e. 
§ Quart. Journ. Geol. Soc. vol. xxxvi. p. 357, note. 


G. R. VINE ON THE FAMILY DIASTOPORIDA. 387 


Busk’s D. obelia. D. oolitica is a very characteristic type, especially 
of the Pea-Grit Series, in the Cheltenham district at least. 


Another type, not the least important of the whole group, is a 
most peculiar one. In general habit and mode of growth it seems 
to be a true Diastopora ; but it must be looked upon more as a pas- 
sage form than as a constant one in all its characters. The type of the 
cells, when slightly worn, approaches nearer to that of the Paleozoic 
Ceramopore than to any Mesozoic Diastopore; but in the more perfect 
cell the largely developed peristome is unlike any thing in the more 
ancient or the more recent Diastoporide. If this be a true passage- 
form, rather than a constant one, the tendency is toward the Pus- 
tulopora-subverticellata type*. In this type the cells are connate ; 
but in the one under consideration only some few of the cells are 
connate; the great majority are “ separated pores,” as in ordinary 
Diastopore. My diagnosis is made from both the younger and older 
growths of several colonies ; and the description may be relied upon, 
however the species may be placed in the future. In the Pea-Grit 
Series the cells have a very bold outline, with interspaces (in some 
places) between cell and cell. In one specimen from Chipping 
Norton the cells have a Lepralia-like growth, the colonies being 
semicircular and piled one upon another; and some of the cells are 
so immersed that only the peristome can be seen. In naming this 
“type” or “species,” I desire to preserve a generic name, though 
the genus itself is now merged in that of Entalophora and Spi- 
ropora. 


4, Drastopora cRIcoporaA, mihi. Plate XIX. figs. 18-25. 


Zoarium adnate, forming small and large irregular patches, some- 
times self-attached, at other times incrusting other species of 
Polyzoa. In the early stages of growth the colony has a bicircular 
or oval outline; in its later stages the growth is most irregular. 
Zoecia short and stunted tubes, very coarsely punctate; orifice 
ring-like, with a largely developed peristome. Primary zoccium 
inconspicuous, being deeply immersed, giving off to the right and 
left secondary zocecia, which in their turn give off others. .The 
after colonial growth is thus early directed to two opposite points ; 
ultimately the proximal cells unite below the primary cell, so that 
it in time becomes centric or excentric. Owcia? 

Range from Pea-Grit to Great Oolite. 

Cabinets: several very fine specimens of this type are in the cases 
of the Museum of Practical Geology, Jermyn Street; Mr. Longe’s, 
Mr. Walford’s, and Mr. Windus’s. 

Hab. Chiefly on water-worn stones and broken shells. 

I have had very great difficulty in describing this type, more 
particularly on account of its peculiar preservation. One colony on 
Mr. Longe’s specimens (block 6) is very well preserved in its earliest 
“stages ; and from this specimen most of the figures are drawn. The 
natural size is shown at fig. 18, and is about three lines at its widest 


* See Busk’s ‘ Crag Polyzoa,’ pl. xviii. fig. 1, right-hand specimen. 


388 G. R. VINE ON THE FAMILY DIASTOPORID A. 


part. In fig. 19'(an enlargement of a portion of the same colony) the — 
tendency of the cells to the right and to the left is shown; the other 
figures are enlargements of special cells to show the ring-like cha- 
racter of the mouth and peristome. ‘There is another colony on the 
back of the same slab; but this is very much worn, and the cell- 
mouths are somewhat angular. The largest colony is on block 9*; 
and it is a very interesting study. ‘This block is a piece of coarse 
Oolitic Limestone, much worn by attrition; and, besides many 
Serpule, it contains on its surface several colonies of Polyzoa. 
Fig. 25 is a magnified portion of a colony of D. cricopora adherent 
to a colony of Hlea(?), or one of its nearest allies, the one almost 
wholly enveloping the other. In the fragment figured the different 
characters of the two fossils areshown. The under one of the Elea (?) 
type has all the cells in one plane, the walls of each cell so closely 
connected as to leave no interspaces. The normal orifice of the 
cell seems to be of a subcircular character ; when slightly worn it is 
large and circular, unlike any of the cell-mouths figured by Busk 
in his ‘Crag Polyzoa’ as Mesenteripora, which seems to have been 
used as a synonym ~ of Hlea foliacea (D. foliacea, Lamx.). Manzoni 
also figures t a “ Diastopora” ? having a habit somewhat similar to 
this, which he calls D. expansa, Manz. ; but the aperture is normally 
circular with a well-developed peristome. This flat adherent type 
of the Inferior Oolite becomes (so Mr. Longe informs me) foliaceous 
in its after stages; but whether it ever becomes really leaf-like with 
cells on both sides I cannot, as yet, satisfy myself. I have no desire, 
however, to put this type under the genus Diastopora. ‘The other 
incrusting form is a marginal portion of D. cricopora, having many 
of the characters of the group, but with cells altogether at variance 
with the general build of the true Drastopora-cell. 


The genus Diastopora has been, and is, in many instances, very 
much abused ; and before proceeding with these studies it may be as 
well to define and limit the genus. Lamouroux used two terms which 
have come into general use—the one Berenicea, and the other Dias- 
topora. Under these two names many divergent forms have been 
placed, so that to some extent Berenicea and Diastopora are synony- 
mous terms, the term Berenicea being used for one section of the 
Diastoporid and Diastopora for another section. In subdividing the 
foliaceous Cyclostomata, Milne-Edwards formed two great groups, 
‘« distinguished,” says Busk, ‘‘ by the character that in the one the 

ubes are almost wholly immersed, and in the other partially free.” 
To the former group Milne-Edwards applied the appellation ‘ Dras- 
TOPORES,” and to the latter that of “ Tuputrporss.” ‘ This division is 
natural ; but it seems convenient that it should be carried still further ; 


* These numbers refer to the specimens as numbered in Mr. Longe’s cabinet. 
It would be a good thing to have the types preserved in some public museum, 
so that they could be accessible to future students. Whenever I could, I have 
referred to specimens so preserved. 

+t D’Orb., Pal. Frane. terr. Crét. (p. 808). 

t Briozoi di Castrocaro, tav. vi. f. 83. 


G. R. VINE ON THE FAMILY DIASTOPORID®. 389 


and in the doing of this... no better classification can be adopted 
than that suggested by the same author, viz. into 1. Diastopores 
simples; 2. D. enveloppantes ; 3. D. bisérialaires. I propose, there- 
fore, to term the simple Diastopore of Milne-Edwards Diastopora, the 
enveloping or laminated forms Berenicea, and the biserial Mesenterz- 
pora’”*, This arrangement Mr. Busk follows in his ‘ Crag Polyzoa.’ 
But Mesenteripora is placed as a genus of the Tubuliporide, whereas 
in the Museum Catalogue, part 111. Cyclostomata, it is placed as a genus 
of the Diastoporide. This I have no objection to, because it is only 
those who have gone over the same ground who can possibly know 
what difficulties there are to encounter in classifying the species 
under discussion. The differences, however, between Mr. Busk and 
Mr. Hincks* are very striking on this point. In the ‘ Catalogue of 
Cyclostomata,’ Mesenteripora (a foliaceous form of the Diastoporide) 
is included in the Famity ; whereas in ‘ British Marine Polyzoa’ the 
foliaceous forms are included in the genus Diastopora. ‘This I entirely 
object to, on account of the confusion it is sure to create when we 
are dealing with Mesozoic forms. With this exception Diastopora, 
in the sense in which I have used it, may be defined as follows :— 

“ Zoarium adnate and crustaceous, usually discoid or flabellate, 
less commonly irregular in form. Zowcia tubular, with an elliptical 
or subcircular orifice, crowded, longitudinally arranged, in great 
part immersed.” 

The foliaceous species will have to be separately dealt with; and 
I think it very unwise to use the term Berenicea, as defined by 
Lamouroux, for other than Paleozoic species. IRPf Berenicea is used 
for Paleozoic and Oolitic species indifferently, simply because the 
*‘corallum incrusts foreign bodies ” and is “ composed of a thin cal- 
careous base” t, confusion will be sure to follow, for the simple 
reason that the characters of the Paleozoic and Oolitic species 
are altogether different. The Ceramopore and Berenicee of the 
Paleozoic rocks are not typical Diastopore. 

There now remain for investigation the Chalk- and Greensand- 
forms; and these I would gladly revise if Members would help me 
by the loan of material for this purpose. Some of the species 
catalogued are undoubtedly Drastopore; others are not Diasto- 
pore in the restricted sense used by me in this paper. 


My thanks are due to kind friends who have assisted me with 
material for the writing of this paper—to Mr. Walford for supply- 
ing me with extracts and tracings from Dumortier and Prof. Braun, 
and to Mr. J. D. Longe, F.G.S., for the loan of specimens from the 
‘Inferior Oolite. I also tender my thanks to Mr. R. Etheridge, 
F.R.8., and to Mr. E. T .Newton, F.G.S., for allowing me to examine 
the specimens in the Museum of Practical Geology. 


* Crag Polyzoa, p. 109. 
+ British Marine Polyzoa, p. 457. 
t M‘Coy, Brit. Pal. Foss. p. 44. 


Q.J.G.8. No. 147. 2D 


Figs. 


Figs. 


Figs. 


Figs. 


G. R. VINE ON THE FAMILY DIASTOPORIDA, 


EXPLANATION OF PLATE XIx. 


1-10. Diastopora stomatoporides, Vine. 


1, 2. Natural size of two different colonies. 

3,4, '7. Enlarged respectively about 10, and 15 diameters, to show the 
disposition and character of the cells. 

5, 6. X 25 diam., showing the elongated character of cells, together with 
the rugose markings on some. 

8. x 50 diam. 

9. xX 30 diam. 


10. Aperture of cell, X 50 times, to show the true Elea-foliacea (Dia- 


stopora, Lamx.) type of peristome; from colony. Fig. 4. As aboye. 
(Compare Berenicea striata, Haime, pl. vii. fig. 8, a,b, ‘ Foss. Bryozoa 
of the Jurassic Formation.’) . 


11-14. Diastopora oolitica, Vine. 


11. Natural size of colony. 

12. x about 8diam. There is much variation in this type; but it has 
generally a circular habit, as depicted. 

13. X about 33 diam. 

14. x about 50 diam. 


15-17. Diastopora ventricosa, Vine. 


15. Natural size of colony. This also varies as to size of colony. 

16. x 25 times, showing the disposition of the ocecial ‘“‘ goneecia,” Hincks, 
in the different parts of the colony. 

17. X about 50 diam. 


18-25. Diastopora cricopora, Vine. 


18. Natural size of colony from which the typeisdrawn. The specimens 
in the School of Mines vary considerably. 

19. Portion of colony, enlarged in the direction of line I, fig. 18, to show — 
the disposition of the partially immersed cells. 

20-22. x 25 times, giving a fair illustration of the ornamentation of the 
cells. 

23, 24. x 75 times. 

25. Two separate colonies: the uppermost, D. ericopora (marginal edge), 
incrusting Elea foliacea, Lamx. | 


Discussion. 


The PresipENT bore witness to the great value of the author’s 
study of this group of microscopic organisms. 

Prof. Srrtey stated that his study of the Polyzoa had led him to 
conclude that many of the supposed generic differences were mere 
accidents of age and growth. He thought that Mr. Vine’s paper 


was 


a valuable addition to science. He doubted the wisdom of 


inventing wholly new names for previously described species, as had 
been done in one instance by the author. 


@uart: 


Frank Rutley del. A.S Foord hth 


Journ.Geol. Soc Vol. XXXVI, PL XX 


Vie Os BOCKS OF MONTANA UES. 


N 


F. RUITLEY ON THE VITREOUS ROCKS OF MONTANA, U.S.A. 391 


‘30. The Microscopic Coaracters of the VirrEous Rocks of Montana, 
U.S.A. By Frank Rurtzy, Esqg., F.G.8. Woth an APPENDIX 
by James Eccrus, Esq., F.G.S. (Read April 6, 1881.) 


[Prats XX. ] 


THE specimens which have supplied the material for this paper were 
collected by Mr. James Eccles during a tour in Montana. They 
present so many interesting points of structure that a description of 
their microscopic characters:‘may prove acceptable to those who are 
engaged in the study of British vitreous and devitrified rocks. 
Among the latter we have already found that certain structural 
peculiarities may still be clearly recognized, notwithstanding the 
changes which have led to their more or less complete devitrification. 
Through these changes, once vitreous rocks assume the character of 
felstones; and as it is highly probable that many of our ancient 
‘“‘felstones ” and ‘* hornstones” were once vitreous, it becomes im- 
portant that we should note every structural peculiarity in glassy 
rocks of recent or late geological age which have undergone little or 
no change subsequent to their solidification. By doing this we are 
training our eyes to recognize similar structures in the devitrified 
obsidians, perlites, and pitchstones which ran over what are pro- 
bably the earliest land-surfaces of which we have any trace. 
Zirkel, in his ‘ Microscopic Petrography of the 40th Parallel N.,’ 
has described a large number of structures met with in the districts 
surveyed by Messrs. King, Hague, Emmons, and other members of 
the U.S. Government Survey ; and it is to him we owe much of our 
knowledge concerning these minute structures. In this country the 
researches of Prof. Bonney and Mr. Allport have also served to in- 
erease the interest which rocks of this class will always possess. 
The more carefully and patiently we study the unaltered examples, 
the better shall we be able to deal with the questions which concern 
their older and no longer vitreous representatives ; while researches 
upon artificial slags or glasses formed under known conditions, and 
modified by various known causes, either during or subsequent to 
solidification, will lend additional help in explaining minute struc- 
tural peculiarities which still remain to be worked out. 

The following is a description of the microscopic characters of 
ten of the most interesting specimens (which may be regarded as 
fairly typical of the whole series) collected by Mr. Eccles in this 
district. In each case the letter M precedes the description of the 
microscopic characters. 

No. 1. Yellowstone district. Black obsidian. 

A black and perfectly vitreous rock, speckled with rather sparsely 
_ disseminated small greyish-white flecks, which are generally imper- 
fectly developed felspar crystals. 

M. In this section it is transparent and colourless, and is a re- 
markably homogeneous glass. The porphyritic crystals (which are 

2p 2 


392 F. RUTLEY ON THE MICROSCOPIC CHARACTERS 


not very numerous) are, in some if not in all cases, orthoclase. 
Some are perfectly developed; others present the most ragged and 


irregular boundaries, as in the accompanying figure (fig. 1). In - 


Fig. 1.—Felspar Crystal in Obsidian (black), from the Yellowstone 
i District. (Magnified 55 times, crossed Nicols.) 


“aa - ate 
: ASN ow, &, 


this instance the spicular projections are sufficiently long and 
delicate to completely invalidate the supposition that the crystal 
is a broken fragment, and to render it absolutely certain that it 
has been developed during the cooling and solidification of the rock 
in which it occurs, or else that it is a partially dissolved crystal. 
This is an important piece of evidence, because less irregularly 
bounded imperfect crystals might often be erroneously regarded as 
included fragments taken up by and enveloped within the lava. 
Moreover it is interesting to see how, in such a remarkably homo- 
geneous matrix, this crystal, if aborted, fails to assume the globular 
form, rounded at the angles, which is so common in the porphyritic 
crystals developed in most vitreous rocks. Under an amplification 
of 300 or 400 diameters numerous clear granules and opaque tri- 
chites are visible, as shown in fig. 1, Pl. XX.; and there are many. 
small objects which may be elongated gas-pores ; but, as they mostly 
pass diagonally through the section from the upper to the lower 
surface, it 1s not easy in all cases to ascertain their true character, 
owing to the impossibility of bringing them into focus throughout 
their entire length. Those which coincide with the planes of section 
certainly appear to be elongated, and sometimes tortuous, gas-pores. 
The trichites are sometimes straight, sometimes curved, and frequently 
form stellate groups. 7 

No. 2. Yellowstone district. Obsidian. ots3 

Reddish-brown or Indian-red coloured rock, with some dark brown 
or black streaks and mottling, opaque, except in thin sections. 
Bright vitreous or slightly resinous lustre, and imperfect conchoidal 
fracture. 

M. It is seen bytransmitted light (fig. 2, Pl. XX.)to consist of delicate 
bands or strings of a clear yellow or orange-colour, with here and there 
_ afew bluish-black or grey strings. The material lying between these 
coloured strings, which are very closely packed, is a colourless or 


OF THE VITREOUS ROCKS OF MONTANA, U.S.A. 393 
almost colourless glass. The yellow fibres depolarize in all azimuths 
except those coimciding with the directions of the principal sections 
of the crossed Nicols. ‘There is therefore a marked optical difference 
between the yellow and the colourless bands, the latter being per- 
fectly isotropic*. The generally parallel disposition of the bands 
imparts an appearance to the magnified section which very closely 
resembles the structure seen in a longitudinal section of wood; and 
this resemblance is further heightened by the deflection of the bands 
around small porphyritic crystals and fragments of quartz, which 
may be likened to little knots in wood. Although the greater por- 
tion of the preparation appears dark between crossed Nicols when 
the general direction of the bands is parallel with one or other of 
the Nicol-sections, still the deflected bands around the porphyritic 
erystals transmit a strong brownish-yellow light, because they do 
not, in this position, coincide with either of the Nicol-sections, so 
that around each brightly depolarizing porphyritic crystal there are 
also depolarizing fringes resembling smoothly-combed tresses of hair. 
Some of the porphyritic quartz-crystals have irregular creeks of glass 
passing into them, in some cases the yellow fibres being mixed with 
the included colourless glass, thus clearly showing that these quartz- 
erystals were actually developed within the rock prior to or during 
its solidification. The preparation also shows numerous dark opaque 
granules and a few dark sections, apparently of minute octohedra, 
which are opaque, except at their margins, where they feebly trans- 
‘mit light; and this imperfect opacity seems to imply that they are 
not magnetite. The fine banding in this rock is a most perfect ex- 
ample of fluxion-structure. 

Representatives of this class of rock, similar in structure, but 
more or less devitrified, are to be found among the lavas of early 
Paleozoic age in North Wales. 

No. 3. Yellowstone district. Black spherulitic obsidian. 

A black glassy rock, containing numerous pinkish or pale-grey 
spherules irregularly distributed and varying from the size of a pea 
downwards to very small dimensions. 

M. The section is seen to be traversed by bluish-grey and colourless 
bands, the former consisting of streams of microliths (fig. 3, Pl. XX.). 
In this banded structure there is no depolarization, showing that the 
banding in this case is quite different from that in the red obsidian just 
deseribed. The spherules either occur isolated or in little colonies. 
Porphyritic felspar crystals occur here and there, and they show a 
few glass enclosures. 

No. 4. Yellowstone district. Spherulitic-banded obsidian. 

A rock consisting of dull dark grey and vitreous black bands, 
flecked with white porphyritic crystals of sanidine. 

M. The dull bands are seen to consist of radiately crystalline 
spherules ; while the deep-black bands visible in the hand-specimens 
appear in thin section as clear and colourless glass. In the drawing 

* Since this paper was read, I have satisfied myself that the double refraction 


in these coloured bands is probably due to strain, a thick bundle of spun glass 
exhibiting similar phenomena. 


394 F, RUTLEY ON THE MICROSCOPIC CHARACTERS 


(fig. 4, Pl. XX.) two portions of one and the same sanidine erystal, 
which between crossed Nicols undergo extinction synchronously in 
the same azimuth, are separated by spherulitic matter; and this 
seems to indicate that the spherules and crystals were probably 
developed at the same time. az 

No. 5. Yellowstone district. Black porphyritic obsidian. 

A black glossy rock, with white porphyritic crystals, some of them 
+ inch long. A smoothly-ground surface shows fine, continuous, 
greyish, parallel bands and porphyritic erystals in a deep-black 
ground-mass. The fractured surfaces of the specimen are very irre- 
gular or small conchoidal. 

M. The bands are seen to consist of closely aggregated spherules ; 
and the surrounding glass is filled with streams of microliths, which 
impart a finely banded appearance to it. The porphyritic erystals 
lie with their longest axes in the same direction as the bands. One 
erystal (part of which is shown in fig. 5, Pl. XX.) is completely sur- 
rounded by a border of small spherules; and in another instance a 
little crystal lies immediately within a rudely concentric series of 
perlitic cracks. The section is fissured in all directions, the cracks 
traversing the glassy matrix, the spherulitic bands, and the por- 
phyritic crystals; they must therefore have been produced subse- 
quently to the formation of those bodies. A few of the porphyritic 
crystals in this rock bear a close resemblance to olivine. One 
transverse section of a rhombic prism gives an angle of 94°, which 
corresponds with the angle in olivine. The face of the section shows 
the granulated surface so frequently seen in olivine-sections ; the 
angles of the crystal are slightly rounded; and the fringed cracks 
which traverse the crystal closely resemble those seen in olivine. 
The extinctions, moreover, in this crystal coincide with the directions 
of the crystallographic axes, assuming the section to be transverse 
toarhombic prism. If this crystal be olivine (as I believe it to be), 
we have a curious instance, possibly the only one yet recorded, of 
olivine occurring in an obsidian. Zirkel, however, describes the 
occurrence of olivine in a trachyte from the top of Whitehead Peak 
in the Elkhead mountains, which he says “ presents, beside sanidine, 
very many cracked quartzes as large as a pea, hornblende, and augite, 
and, what is remarkable, not very numerous but doubtless charac- 
teristic half-serpentinized olivines, the sections of which, measuring 
as high as 0°75 mm., are visible even to the naked eye in the slides. 
The peculiar quartz occurring here is therefore accompanied by a 
mineral which has never before been observed in a sanidine rock” *. 

No. 6. Yellowstone district. Spherulite rock. . 

A pale bluish-grey rock, slightly cellular and with yellowish-brown 
stains, with numerous little porphyritic felspar crystals (chiefly 
sanidine) and, exceptionally, some roundish grains of quartz. On 
a smoothly-cut surface the rock is seen to consist almost wholly of 
closely aggregated spherules, with a little dark glassy interstitial 
matter. It is, in fact, an obsidian almost completely devitrified by 


* “Microscopical Petrography,” p. 159 (U.S. Exploration of the Fortieth 
Parallel: Washington, 1876). 


OF THE VITREOUS ROCKS OF MONTANA, U.S.A. 395 


the development of spherules and small porphyritic crystals, and 
may be regarded as a spherulitic condition of sanidine trachyte 
or rhyolite. 

M. It is seen to consist of a multitude of spherules which possess 
a radiating, crystalline, or fibrous structure, and consequently show 
dark crosses between crossed Nicols (fig. 6, Pl. XX.). The spherules 
vary in diameter from about ;4, inch to extremely minute specks, 
when the section appears (in these parts) to pass into a microcrystal- 
line condition, apparently identical with felsite. Indeed, from the 
microscopic study of this and many other more or less closely allied 
rocks, it appears to me desirable to abolish any hard lines of classifica- 
tion which have hitherto been drawn between the trachytes, rhyolites, 
and felstones. The spherules frequently form continuous and some- ~ 
what tortuous strings, which, between crossed Nicols, resemble the 
chenille put round the bottom of glass shades to exclude dust. The 
porphyritic felspars frequently fail to show any definite crystal- 
lographic boundaries, and are occasionally penetrated by creeks of 
the surrounding substance, which in one or two instances is included 
and shut off from the matrix. The porphyritic quartz-grains are 
often irregular in outline, and do not appear to have any good fluid 
enclosures. 

No. 6a. Yellowstone Cafion. Quartz rhyolite. 

A pale bluish-grey compact rock, showing delicate wavy fluxion- 
bands, which, on a smooth surface, are seen to thin off, seldom being 
continuous for more than an inch. Numerous small porphyritic 
grains of quartz and a very few crystals of sanidine are also visible. 

M. The ground-mass of the rock is seen to be apparently micro- 
_ granular, while the bands also show a microcrystalline or crystalline- 
granular structure, but very much coarser. It is Just the same tex-- 
tural difference which is still visible in microscopic sections of our 
archean and paleozoic rhyolites. The ground-mass varies in texture 
in different parts of the section; and, where finest-grained, it is 
studded with little roundish blotches, which seem to depolarize more 
strongly, and which appear lighter than the ground-mass when the 
Nicols are crossed, and darker when they are set parallel, or when 
the section is seen by ordinary transmitted light. When examined 
with an amplification of about 250 diameters, these spots seem to be 
the result of devitrification; while by employing a quartz plate it 
is seen that the ground-mass is partially isotropic, and the doubly 
refracting spots appear to become more mixed with isotropic matter 
towards their margins. The spots, in fact, resemble segregations of 
doubly refracting granules, which impart a sort of spotty mottling 
to the ground-mass. The porphyritic quartz-crystals are nearly all 
rounded in outline, like those which oceur in quartz-porphyries ; but, 
unlike the latter, they seem to contain no fluid lacune. This rock 
affords interesting material for the study of devitrification. 

No. 66. Lower Geyser basin; is a pale grey or drab rock, with 
darker parallel bands. It is a banded spherulite rock, the bands 
consisting of small and the remainder of larger spherules. It closely 
resembles No. 6, except in the better definition of the bands. The 
rock contains some porphyritic felspar crystals. 


396 ¥F, RUTLEY ON THE MICROSCOPIC CHARACTERS 


No. 7. Quartz rhyolite. Gardiner’s River. 

Compact pale bluish-grey rock, containing numerous granules of 
quartz and a few crystals of sanidine. A rudely banded structure is 
visible on a smoothly cut surface of the specimen. 

iM. Shows curiously twisted and gnarled bands which end abruptly 
(fig. 7, Pl. XX.). This may, however, bein some instances due to 
their being cut off by the upper and lower surfaces of the preparation. 
The structure closely resembles the mottling on gun-barrels. in some 
of the thicker bands traces of a fibrous crystalline structure, transverse 
to the bands, is visible in polarized light ; and this, taken in conjunc- 
tion with the frequently looped or annular disposition of the bands, 
_may be accepted as evidence that, at all events to some extent, the 
general structure of the rock is due to an attempt to develop sphe- 
rules, but especially such as have an elongated axis, as in the axiolites 
described by Zirkel. 

No. 8. Yellowstone district, Lower Geyser basin. Vitreous tuff 
(“‘ obsidian sandstone”). 

A finely granular rock of a dark or blackish colour, with light 
specks. The granules have a vitreous lustre. 

M. It is seen to be a tuff composed of small fragments of vitreous 
rocks which in most instances show well-marked perlitic. structure 
(fig.8, Pl. XX.), and detached crystals and fragments of crystals which 
in most cases appear to be sanidine and occasionally plagioclastic 
felspar. The transverse sections of some of the former are lozenge- 
shaped, the boundaries being faces of the oblique rhombic prism and 
giving an angle of about 118°. These fragments of rocks and crystals 
are bound together by a cementing material which, in a not very thin 
section, appears brown or even absolutely opaque by substage illu- 
mination, and reddish brown by reflected ight. It is most likely 
limonite. As already mentioned, the section is a rather thick one; 
and between crossed Nicols the majority of the spheroidal spaces 
enclosed by the perlitic cracks in the rock-fragments exhibit more 
or less well-marked depolarization and a dark interference cross. 
The cross is sometimes rather irregular or distorted. In some 
instances depolarization takes place only along the bounding cracks ; 
in others it forms a well-defined zone, the central portion, where the 
arms of the cross would intersect, remaining dark and forming an 
approximately round spot, as in fig. 2. These phenomena appear 


Fig. 2.—Areas of Depolarization from Strain within Perlitic Bodies 
én Obsidian Tuff (“* Obsidian Sandstone”), Yellowstone Dastrict. 


most distinctly to be the result of strain; for Lommel* remarks, 


* “The Nature of Light,’ p. 330 (International Scientific Series, London, 1875). 


OF THE VITREOUS ROCKS OF MONTANA, U.S.A. 397 


“the double refraction of compressed and suddenly cooled glass is 
nevertheless essentially different from that of crystals;” and he 
adds that, in order to project the system of rings of a strained glass 
disk upon a screen, using Dubosq’s polarizing arrangement, it must 
be placed at a point where “ the rays by which it is struck are nearly 
parallel, and traverse the plate in the same direction and with the 
same length of path. ‘The difference of path which gives rise to the 
system of rings can therefore only be due to the fact that the double 
refraction, whilst the course of the rays remains unaltered, increases 
towards the periphery of the plate. In a crystal, on the contrary, 
the double refraction is at all points the same for the same direction 
Othe rays”... 

Before proceeding another step I wish to place side by side with 
this statement an extract from the late Hermann Vogelsang’s ‘ Kry- 
stalliten’*. Speaking of some crystallites in a piece of thick glass 
from the glass-works at Stolberg, he states that for a certain distance 
around these isotropic crystallites the glass exhibits double refrac- 
tion, a neutral cross also traversing the anisotropic area. In one 
case he noticed a slight disturbance of this cross in the neighbour- 
hood of an ellipsoidal body, which seemed to him to indicate that 
the strain in this instance was not constant in all directions; and 
he states, ‘‘ It is also to be remarked that the glass at the boundary 
of the polarization-picture is sometimes traversed by a fissure, a 
spheroidally-running cleft. The polarizing action is not thereby 
disturbed. Whether these cracks were produced during the cooling 
of the glass, or whether they have subsequently been developed in 
the splitting-off and grinding of the preparation, | am unable to say.” 
If the careful observer, who penned the lines I have just quoted, 
had seen the preparation which is now placed before you, I think 
that his doubts with regard to the origin of those spheroidal fissures 
would have vanished. We have here, I believe, additional confirma- 
tion of the views advocated by Professor Bonney, Mr. Allport, Mr. 
Cole, and myself with regard to the origin of perlitic structure. 
But we have something more. If Vogelsang’s crystallites be em- 
bryonic erystals (as I think we may certainly assume that they are), 
we have a close relationship between crystallogenesis and perlitic 
fission ; and, indeed, in the section to which our attention is now 
confined, there are plentiful examples of doubly refracting crystals 
which are immediately surrounded by perlitic cracks, but which do 
not, save very exceptionally, transgress those boundaries. 

In a paper upon a somewhat kindred subjecty I have already 
quoted a statement of Bischoff, to the effect that trachytic rocks, in 
passing from a vitreous to a crystalline state, undergo a shrinkage 
of nearly 10 per cent. of their original bulk. In those few instances 
in which a perlitic crack passes through a crystal, there is commonly 
another crystal developed by its side, which, with its surrounding 


* “Die Krystalliten,’ Bonn, 1875, p. 68. Admirable figures of these crystal- 
lites are given in plates ix. and x. of the vbove work. 

+ “On some Structures in Obsician,” erlite, and Leucite,” Monthly Micro- 
scopical Journal, vol. xv. p- 183. 


398 F, RUTLEY ON THE MICROSCOPIC CHARACTERS 


crack, may have been formed subsequently to the crystal which is 
traversed by that crack. What are we to infer from these crack- 
begirt crystals? Surely we have here something lke the contraction 
experimentally determined by Bischof in rocks of precisely similar 
constitution. The crystal is a constituent of trachyte; it may be 
regarded even as a small piece of trachyte developed within obsidian. 
The strain upon the surrounding glass, engendered by the formation 
of this crystal, is, I have no doubt, the cause of the rupture in the 
surrounding glass; and the gape of the fissure is the measure of the 
contraction for any one plane. ‘The polarization-figures seen within 
the perlitic areas surely, then, bespeak the incipient strain which 
heralds the production of the crystal; and I believe that the 
rounded boundaries and the curved cracks, so frequently seen within 
the angles of olivine and other crystals developed in vitreous rocks, 
are but another expression of the laws which crystallization enforces 
in a surrounding amorphous mass. In studying these questions we 
seem almost to stand upon the threshold of: crystallography; and I 
may well close this paper with the hope that the subject may be 
taken up by more able hands; for it requires further investigation, 
since there are periites in which no apparent relation exists between 
the perlitic structure and the porphyritic crystals—as in some of the 
perlites of Chemmitz, where even a score of small perlitoids may be 
seen to abut against the margin of a single crystal of sanidine or 
of magnesian mica. In these rocks we have, indeed, always assumed 
that the development of perlitic structure was subsequent to the 
formation of the porphyritic crystals, and not approximately syn- 
chronous with it, as 1 think has been the case in the materials of 
the obsidian tuff just described. In an inquiry of this kind thick 
sections, as well as thin ones, should be prepared and examined, ~ 
and conclusion after conclusion discarded if needful, until the truth 
is reached. Indeed, I am not unwilling to believe that some of the 
phenomena which we have just discussed may be due to different 
operations of one force, or of different forces; and, just as the same 
mineral may sometimes be formed by a wet or by a dry process, so 
it is possible that similar structures may not always be due to the 
same cause. 


AppEenvix.— Microscopic Characters of Volcanic Rocks of Montana. 


No.9 (the rock from Mount Washburn) is an andesite which seems to 
be intermediate in character between the augite- and the hornblende- 
andesites, and approximates in mineral constitution to a basalt. 

The constituents are chiefly triclinic felspar, augite, hornblende, 
magnetite, and vitreous matter. 

A little sanidine also appears to be present. The augite and 
hornblende crystals generally show a dark border when examined 
under the microscope with substage-illumination. By reflected light 
these borders appear of a bright rust-red, and are doubtless due to 
marginal decomposition of the hornblende and augite. These stains 
and granules of peroxide of iron impart a reddish brown-colour to 
the rock. The vitreous matter constitutes a considerable proportion 


OF THE VITREOUS ROCKS OF MONTANA, U.S.A. 399 


of the ground-mass. A little bright green matter is also present, 
which appears to occupy irregular vesicles, and which is either green 
earth or some closely allied mineral of secondary origin. The angles 
of extinction of the felspars seem to indicate that the triclinic ones 
are labradorite. 

The rock from Tower Falls closely resembles that just described, 
except that it contains apparently no augite and more hornblende. 
The erystals of the latter mineral are also surrounded by opaque 
borders, which are often broad, and consist of an internal zone of 
magnetite and an external one of hematite, which appears sharply 
defined at its contact with the magnetite. The rock is hornblende 
andesite. It is of paler colour than that from Mount Washburn, 
the hand-specimen being grey, with minute reddish brown, black, 
and white specks *. 


APPENDIX. 


On the Move of OcctRRENcE of some of the Votcantc Rocks of 
Montana, U.S.A. By James Kccrzs, Esq., F.G.S. 


Tue yoleanic district of the Yellowstone National Park, from which 
I obtained the various rock-specimens referred to in the foregoing 
paper, has been already described in detail by Dr. Hayden, Dr. A. 
C. Peale, and Prof. Bradley, in the reports of the U.S. Geological 
Survey of the Territories for the years 1871 and 1872. 

In the autumn of 1878 I had the good fortune to accompany Dr. 
Hayden and other members of his survey in some parts of the 
district referred to; and although I have no intention of giving a 
detailed description of the voleanic phenomena which were observed, 
a short notice of the localities whence the specimens were obtained, 
and of the general mode of occurrence of the rocks, may be of some 
interest as a supplement to Mr. Rutley’s description of their micro- 
scopic characters. 

The Yellowstone National Park, within which are the head-waters: 
of two of the great forks of the Missouri (the Yellowstone river and 
the Madison) and of the Snake river (one of the branches of the 
Columbia, comprises an area of about 3600 square miles, nearly the 
whole of which is covered up by voleanic rocks of great thickness. 
There are very few exposures of underlying sedimentary or other 
formations; and these are almost entirely limited to the extreme 
northern edge of the area. Some fifteen miles south of the southern 
boundary the extension of these volcanic rocks is seen to rest upon 
the northern spurs of the Teton range of mountains, at which point 
the underlying formation has been ascertained by Prof. Bradley to 
be of Carboniferous age. On approaching the Park from the south 
along the upper valley of the Snake, these volcanic rocks appear to 
form an irregular plateau densely covered with forest. From the 
point at which we first struck this plateau, as far as the Upper 
Geyser basin on the Firehole river (the chief branch of the Madison), 


* For the discussion upon this paper see p. 412. 


400 J. ECCLES ON THE VOLCANIC ROCKS OF MONTANA, U.S.A. 


Fig. 3.—Map of part of the Yellowstone Park. 
(Scale about 20 miles to 1 inch.) 


q ss A ee 
: =—_ ow =e = @ @ @ — - | 


TOWER FAULS Ko 4 


MOUNT 
Mi wasnauRn | 


Ane 


ba 


= 
1295 Pap nosnone 


awe eee ee ieee ie 


is a linear distance of about twenty-five miles, but three days of hard 
travelling. The rocks consist entirely of varieties of obsidian and 
trachyte, the latter almost invariably being found in great mass 
under the former. 

The obsidian, though frequent, is somewhat irregularly distri- 
buted. It is both black and reddish brown, is occasionally columnar, 
and is nearly always porphyritic. The most common form is a coarse, 
rapidly weathering variety, containing many crystals of sanidine. 
The disintegration of this rock produces some curious fine conglo- 
meratic deposits in the old watercourses and river-beds (No. 8). 

The trachyte is constant and of very great thickness, and is evi- 
dently closely allied to the obsidian. A cliff-section in the Upper 
Geyser basin suggests a transition from trachyte to obsidian. (Nos. 
4 and 6 were obtained from this section.) 


J. ECCLES ON THE VOLCANIC ROCKS OF MONTANA, U.S.A. 401 


* East of the Firehole river and across the divide between the 
Madison and Yellowstone rivers the section presents the same gene- 
ral characters as that between the Snake and the Madison. High 
up on the divide the trachytes are light in colour, and there is some 
pumice occurring on the surface. On the east of the divide the 
obsidian thins away for some distance, but is found again above 
the Great Canon of the Yellowstone, where it is capped by fully 
300 feet of fine sandstones and shales, probably of Quaternary age, 
which have doubtless been deposited by the Yellowstone lake in a 
former period of extension. The rocks in this basin, and especially 
in the section exposed in the Great Cafion, have been minutely de- 
scribed by Drs. Hayden and Peale; and I have nothing to add to 
their description. It is sufficient to say that the order and general 
character of the rocks is almost precisely the same as in the upper 
part of the Madison basin. . 

Up to this point the voleanic phenomena had been almost uni- 
form. The trachyte-flows, although dipping shghtly here and there 
(as much as 20° in the Upper Geyser basin), were approximately 
level; and the country through which I had passed seemed to be 
quite destitute of any features which I could recognize as vents from 
which such an enormous mass could have been poured out. Seven 
miles north of the head of the Cafion a change is observable. Mount 
Washburn here attains an elevation of from 2500 to 3000 feet above 
the trachytes of the Canon, and is a true volcanic cone. The summit is 
abroken-down crater; and the lava-flows, which are basaltic (No. 9), 
dip away in all directions, and conform generally to the slopes of the 
mountain. I found no contact between the basalts and the trachytes 
on the southern side of the mountain; but it is quite evident that 
the former overlie, and are more recent than, the great mass of tra- 
chytic rocks just described. 

Descending on the north side of the mountain, the Yellowstone 
river was reached again near Tower Falls, a distance of ten miles 
from the peak; and here the trachytes are again seen in the river- 
section, with columnar basalts resting on them. I «was not sure 
whether these basalts had flowed from Mount Washburn or from 
some vent on the east side of the Yellowstone; but, compared with 
the trachytes, they were comparatively insignificant both in extent 
and, except on the mountain itself, in thickness. ‘These basalts had 
been poured out before the formation of the present Cation; for the 
river here cuts its way through basalt and trachyte nearly 400 feet 
deep. 

The locality of Gardiner’s river, whence the specimen No. 7 was 
obtained, as well as the intervening spaee between this river and 
Tower Falls, was passed over most rapidly by me, owing to bad 
weather and Indian troubles. The trachyte in this locality resem- 
bles strongly that of the Great Canon in mineral character. Although 
there is a considerable distance between Tower Falls and Gardiner’s 
river where the trachyte does not exist, owing, probably, to denu- 
dation, I am inclined to regard this Gardiner’s-river trachyte as 
belonging to the great mass of trachytes further south. 


402 ON THE VOLCANIC ROCKS OF MONTANA, U.S.A. 


EXPLANATION OF PLATE XX. 
Vitreous rocks of Montana. 


Fig. 1. Black obsidian containing trichites. Yellowstone district. x 250: 

2. Red obsidian showing in thin section reddish-yellow bands of glass, 
and grains of quartz, ina nearly colourless glass. Yellowstone district. 
x 205. 

3. Black spherulitic obsidian with bands of microliths and radiately crys- 
talline spherules. Yellowstone district. Xx 77. Nicols at 85°. 

4. Spherulitic obsidian, consisting of dull dark-greyish and vitreous black 
bands, flecked with white glassy porphyritic crystals of sanidine. The 
dull bands are composed of radiately crystalline spherules, while the 
vitreous bands consist of glass, black in the hand-specimen and colour- 
less in thin section. Yellowstone district. x 25, 

5. Black porphyritic obsidian with spherulitic bands. The drawing shows 
part of a section of one of the porphyritic crystals of sanidine, which 
is surrounded by an isolated girdle of small spherules. Yellowstone 
district. x 18. 

6. Yellowish grey spherulite rock with small porphyritic crystals of sani- 
dine. Yellowstone. x 32. Crossed Nicols. 

7. Rhyolite with peculiar Damascene structure and grains of quartz. 
Between crossed Nicols it is seen to have a microcrystalline structure 
throughout. Gardiner’s River. x 55. 

8. “Obsidian sandstone,” a finely granular blackish tuff, composed of 
angular grains of vitreous rocks of different appearance, the majority 
being perlitic, others showing merely microlithie streaks, but all pro- 
bably derived from approximately the same source and representing 
the disintegration and cementing in place of vitreous rocks. The 
drawing shows part of one of the perlitic fragments, in which, between 
crossed Nicols, an interference-cross is seen in the centres of some of 
the perlitic spheroids, similar to the crosses seen under similar circum- 
stances in artificially stramed or compressed glass. In the lower part 
of the field a crystal of triclinic felspar is shown. Lower Geyser basin, 
Yellowstone district. x18. Nicols at 85°. 


Note.— Except when otherwise stated, the drawings have been made by ordi- 
nary transmitted lght. 


Quart.Journ.Geol. Soc. Vol XXXVIT PL XXL 


Frank Rutley del. A.S Foord lth 


Mirtern Brosimp. 


WELSH LAVAS OF LOWER SILURIAN AGE. 


ON DEVITRIFIED ROCKS FROM BEDDGELERT AND SNOWDON. 403 


31. On the Microscopic Structure of Drvirrirrep Rocks from 
BEeppDGELERT and Snownon ; with an Appendix on the ERUPTIVE 
Rocxs of Skomer Istanp. By Franx Rourrzy, Esq., F.G.S. 

’ (Read April 6, 1881.) 


[Puate XXI.| 


THz specimen from the south side of the Capel Curig road, about 4 mile 
from Beddgelert, which was collected by Mr. George G. Butler*, and 
which he kindly placed at my disposal, is a rather dark greenish- 
grey rock, spotted with pale greenish-grey spherules, some of which 
are over + inch in diameter, and which, besides occurring isolated 
and in approximately spherical forms, have also coalesced, forming 
bands from 4 inch to nearly 1 inch in breadth. On a smoothly cut 
surface they afford, by their pale tint, a strong contrast to the dark 
matrix. The isolated spherules and spherulitic bands are shown in 
fig. 3 (Pl. XXI.) as they appear when magnified about 6 diameters. 
A thin section, when examined between crossed Nicols, is seen to 
be studded all over with small doubly refracting specks. By ordi- 
nary transmitted light, and under a power of 25 diameters, a marked 
difference is visible in the microscopic characters of the spherules 
and the matrix, the former being almost colourless, save for a few 
pale greenish flecks, which become more closely aggregated at the 
margins of the spherules, forming a somewhat darker border, while 
the matrix appears to consist of a closely matted or granulated deep- 
green substance resembling chlorite, through which are interspersed 
a great number of clear spots consisting of nearly colourless matter, 
similar to that which composes the spherules. Between the sphe- 
rules and the matrix, forming a sharp. boundary, is a clear, narrow, 
colourless border (fig. 1, Pl. XXI.), while the matrix itself is tra- 
versed by more or less sharply defined lines, which also appear clear 
and colourless. | 

These lines seem, in places, to be nearly straight, and to divide 


* Extract from letter from G. G. Butler, Esq. :— 


“With regard to the fragment of spherulitic felstone, I have only to say that 
I knocked it off the corner a piece of ‘rock, perhaps 6 or 8 feet square and 
2 or 3 high, projecting from,a sloping field on the south side of the road from 
- Beddgelert to Capel Curig—about a quarter of a mile from the former place, and 
perhaps 200 yards from the road, the field sloping down towards the road. 
The rock appeared to be zz situ. I found similar specimens on other protruding 
- rocks near, but none so perfect as this. I fear I cannot give any more informa- 
tion about it. 

* A rock which abuts on the north side of the road, nearly opposite, but not . 
so far from Beddgelert (in fact, just at the edge of the village), presents a curious 
appearance from a number of globular bodies contained in it averaging 2 or 3 
inches in diameter and standing half out from its surface. When knocked in a 
careful way, they come out bodily, leaving an empty hemispherical socket. 

“T remain, very truly yours, 
“G. G. Buruzr.” 


404 F, RUTLEY ON THE MICROSCOPIC STRUCTUKE OF 


the rock into small irregular cuboidal masses. They also describe 
irregular circles or ellipses, the sections of spheres and ellipsoids, 
which appear nearly to fill the spaces enclosed by the rectilinear 
divisions. 

These straight and curved streaks have a very perceptible breadth 
(as shown in fig. 5, Pl. XXI., magnified about 12 diameters). Be- 
tween crossed Nicols they break up into an infinity of small doubly- 
refracting granules; and we may infer that they represent small 
fissures which have subsequently been filled by infiltration. 

The curved lines seem to indicate a coarse kind of perlitic or 
spheroidal structure; and their relation to the straight lines at once 
calls to mind the similar phenomena, on a larger scale, in the basalts 
of Le Puy and Rowley Regis, described by Pnokesscr Bonney *. 

In ane part of the matrix there i is a roundish patch, somewhat less 
than 54, inch in diameter, in which evident traces of perlitic struc- 
ture are discernible. A portion of this patch, magnified 25 diameters, 
is shown in fig. 4, Pl. XXI. 

The spherules and spherulitic bands are destitute of any definite 
internal arrangement. There is no trace either of radiating or of 
concentric structure. Under an amplification of 575 diameters the 
spherules are seen to consist of a confused aggregate of extremely 
minute, colourless, rounded granules and pale green scales: the 
latter appear to be chlorite. The little colourless granules closely 
resemble in appearance and in dimensions the granules of spes- 
sartine which occur in the Belgian honestones, and which have been 
determined and described by Renardt. Owing to their extremely 
small dimensions, they fail to occupy the entire thickness of the 
section ; hence they are always overlain or underlain by doubly- 


refracting matter, which precludes the possibility of ascertaining 
whether they are isotropic. On examining a section of the coticule 


of Dressante, near Hebronval, in Belgium, given me by Prof. Renard, 
I find just the same difficulty, except upon the extreme margin of 
the section, where a few of the granules have parted from the pre- 
paration, and can be examined independently. In such cases the 
light, of course, undergoes extinction during a complete revolution 
between crossed Nicols. | 

Leturning to the Beddgelert section, a few of the minute colourless 
eranules may also be met with in an isolated condition; and their 


isotropic character can then be readily recognized. Under these 


circumstances we may, perhaps, be justified in regarding them as 
garnets, and possibly the manganese garnet spessartiney. There is, 
indeed, a honestone, well known to the natives, and occurring in a 
quarry at Pen-y-Gwryd at the head of the Llanberis Pass, which, 
like its Belgian representative, contains numerous minute garnets 


* “On Columnar, Fissile, and Ppheroidal Structure,” Quart. Journ. Geol. 
Soe. vol. xxx1i. p. 140. 

t Mémoire sur la structure et la composition minéralogique du Coticule : 
Brussels, 1877. 

i Blowpipe examination of the finely powdered rock shows distinctly the 
presence of manganese. 


DEVITRIFIED ROCKS FROM BEDDGELERT AND SNOWDON. 405 


identical in appearance with the little grains in the Beddgelert 
rock. 

From the general character of this rock, from its spherules and 
spherulitic bands, and from the vestiges of perlitic structure which 
it presents, [ have no hesitation in regarding it as a devitrified obsi- 
dian or pitchstone. This might, however, be inferred by any practised 
geologist without recourse to the microscope ; and it is only right to 
add that Mr. Butler was well aware of its nature when he gave me 
the specimen. 

The rock next to be described is associated with Bala beds, and 
occurs at Clogwyn d’ur Arddu, a high ridge about 1 mile N.W. of 
the summit of Snowdon. The specimen from which the section has 
been cut was collected long ago by Professor Ramsay*, and is now 
in the rock-collection in Jermyn Street. 

The specimen is of a greenish grey colour, and shows an interest- 
ing weathered surface with projecting bands lying closely together, 
and separated by rather deep and narrow furrows. Speaking of 
these bands, Prof. Ramsay states that they “‘ probably originated in 
the same cause that produced the lamination in the lava of Ascension’’}. 
Under the microscope (fig. 2, Pl. X XI.) the section exhibits an irre- 
gular wavy-banded structure, such as might have been inferred from 
the banded character of the weathered surface. In polarized light this 
banding is marked by a strong difference in texture or grain; for the 
rock throughout is microcrystalline and is now felstone. 

A comparison of this rock with unaltered banded obsidians of com- 
paratively late geological age, such as those of Ascension, the Liparis, 
the Yellowstone, and other volcanic districts, leaves but little doubt 
that the structure has resulted from fluxion; and I think we may 
also assume that the rock was once vitreous. 

On the right-hand side of the road, between Pont-y-Gromlech 
and Gorphwysfa, as we ascend the Llanberis Pass, an outcrop of 
dark grey felstone-like rock occurs, which breaks or splinters under 
the hammer into irregular slabs or platy fragments. The fissile 
structure appears at first sight to be due to the presence of dark 
sreenish-black films with an oily lustre, resembling patches of 
talecose slate. It is, however, possible that other and more minute 
structure may also have some share in imparting this schistose cha- 
racter to the rock. In its present condition it may be termed a 
felsite schist ; and it is probable that many geologists would, from 
the general appearance of hand-specimens, regard it as an indurated 
voleanic ash. For a long time the microscopic character of this 
rock has been a source of perplexity to me. Under the microscope, 
by ordinary transmitted light, the section is seen to be made up of 
small fragments, strings, and shreds of every shape, separated by 
finely granular and less translucent matter, which is strongly im- 
pregnated with avery pale greenish chloritic substance forming very 


. * By permission of Prof. Ramsay this description is now laid before the 
ociety. 

T Descriptive Catalogue of Rock-specimens in the Museum of Practical Geo- 
logy, 3rd edit. (1862), p. 42, spec. 874, wall-case 41. 


ee. G.S. No..147. 2E 


406 ¥F. RUTLEY ON THE MICROSCOPIC STRUCTURE OF 


fine strings or films. Fig. 1 conveys a very fair idea of the general 
appearance of the section as seen by ordinary transmitted light, and 
magnified 55 diameters. Between crossed Nicols the shreds and 


Fig, 1.—issile Schist from near Pont-y-Gromlech. 
(Magnified 55 diam.) 


ZW 
RL J 
oe Yj 


\\ 


films are shown to be crystalline granular aggregates, which are in 
great part quartzose. Under a power giving about 250 diameters, 
the matter lying between the shreds appears to consist chiefly of 
very minute doubly-refracting granules; and I think that the whole 
rock simply presents different textural conditions of felstone, im- 
pregnated with some mineral of a chloritic or serpentinous character, 
possibly antigorite, since a porphyritic and schistose roek occurring 
just above Llyn Teyrn contains better-developed films of a similar 
character which closely resemble antigorite. The principal point of 
interest, however, is the meaning of the confused assemblage of 
many-shaped shreds which impart a distinctive character to thin 
sections of this rock. I think that a microscopic examination of 
the rhyolite from Gardiner’s River, in Montana, U.S., described and 
figured in the paper on the rocks of that country which has already 
been laid before the Society, will suffice to show that it is approxi- 
mately the same as the rock now under discussion. 

I do not believe that the structural peculiarities by means of 
which the osteologist correlates the bones of one animal with those 
of another, are more trustworthy than the structural peculiarity 
visible in these two rocks. A thin section of a deep-red obsidian 
from l'olesva in Hungary, given me by Professor Judd, is placed on 
the table for comparison with these rocks., When magnified (fig. 2), the 
pale glass which constitutes the ground-mass of the section is seen 
to be filled with minute strings of a reddish-brown glass, twisted in 
the most irregular manner. It will also be seen that the planes of 


DEVITRIFIED ROCKS FROM BEDDGELERT AND SNOWDON. 407 


Fig. 2.—Deep-red Obsidian from Tolesva, near Tokay, Hungary. 
(Magnified 250 diam.) 


section cut off parts of these convoluted strings, leaving irregular- 
shaped shreds similar to those in the rocks from Gardiner’s River 
and Pont-y-Gromlech, only very much smaller. The shreds in the 
latter rock appear, therefore, to be parts of convoluted strings or 
bands. 

We are thus comparing the Pont-y-Gromlech rock with a distinctly 
vitreous rock (obsidian of Tolcsva), on the one hand, and with a rock 
which does not present a glassy aspect (rhyolite of Gardiner’s River), 
on the other. A question of great interest now arises. Was the 
rhyolite of Gardiner’s River once vitreous like the obsidian of Tolesva? 
If so, then the Pont-y-Gromlech rock may be a devitrified obsidian. 
On the other hand we may be justified in assuming that the micro- 
crystalline condition of the rhyolite from Gardiner’s River, and, in- 
deed, that of any rhyolite, may be an wnmediate result of cooling, yet 
identical with the condition which often supervenes when solidified 
glassy rocks undergo devitrification. 

We may therefore, | think, be allowed to consider the Pont-y- 
Gromlech rock either arhyolite or a devitrified obsidian ; for, in the 
first case, it may be regarded as an obsidian devitrified at its birth ; 
in the second, as an obsidian devitrified in its oldage. The peculiar 
structure of this rock cannot, however, be reconciled with any pro- 
cess of crystallization, but must rather be regarded as the result of 
fluxion in what, at the time, must have been a nearly or quite 
amorphous magma. 

These considerations lead me to the conclusion that it is unwise 
either to employ too many names for rocks of the same character, 
or to give distinctly different names to rocks of different character 
which may once have been identical. Still there are difficulties in 
framing a new nomenclature; for, if we decline to speak of what 
we believe to be a devitrified obsidian as a rhyolite or a felstone, we 

2E 2 


408 F, RUTLEY ON THE MICROSCOPIC STRUCTURE OF 


commit ourselves to opinions concerning former lithological condi- 
tions, of which we sometimes have only very imperfect evidence. 
Moreover, if we decline to distinguish a rhyolite from an obsidian, 
we may with equal justice refuse to recognize a trachyte. It appears 
desirable to follow the classification of some of the continental petro- 
logists in grouping all the highly-silicated vitreous rocks together 
under the term “hyaline rhyolite;”’ and, as our knowledge of this 
group increases, we may possibly find that its members differ more 
in structure than in any other respect, and that even many of the 
structures are common. 

I have already stated that the Pont-y-Gromlech rock is now, to 
all intents and purposes, a felsite-schist * or felstone. Wherein lies 
the difference between a felstone and a rhyolite, so far as mineral 
constitution is concerned? The constituents are essentially felspar 
and quartz (similar to that in plutonic rocks), the felspar in felstones 
being chiefly orthoclase, and that in rhyolites the variety sanidine, 
which occurs only in volcanic rocks. In the majority of cases it is 
difficult to ascertain with certainty the precise nature of the minute 
crystalline granules of felspar in the ground-mass either of a fel- 
stone or of a rhyolite. The rhyolites commonly contain more or less 
vitreous matter; but in the older examples this, if it existed, has 
since undergone devitrification ; and the product is a microcrystalline 
ageregate which cannot be distinguished from the rest of the rock, 
except perhaps in some instances by difference in texture. The 
difference therefore between a rhyolite and a felstone is mineralogi- 
cally a very small one. The quartz-rhyolites or liparites are closely 
related to quartz-porphyry ; so that here, again, we have a mineralo- 
gical affinity in rocks which are, on the one hand, volcanic, and, on 
the other, plutonic; and in many cases, :especially among the older 
rocks, it is by structure alone that they can be distinguished. The 
quartz-porphyries may be regarded as spurs or dykes emanating 
from granitic masses ; these dykes, like the margins of the granitic 
masses, are commonly poor in mica; or that mineral may be totally 
absent. The dykes, again, like the margins of the granitic masses, 
‘re usually fine-grained. We have, then, a ae well-defined 
series, consisting of :— 


dues : Hyaline rhyolites (obsidian, pitchstone, perlite 
Eafe Sea sloet | &e.), quartz-rhyolite, and trachyte. 
Granitic series ... Quartz-porphyry, and granite. » 


Trachyte bears much the same structural relation to quartz-rhyolite 
that granite does to quartz-porphyry. Bathymetrical conditions pre- 
clude the granitic series from having any vitreous representatives. 
That there is a passage from the granitic to the rhyolitic series, as 
suggested by many petrologists, seems more than probable. So far 


* Tt is here worthy of remark that Daubrée has succeeded in superinducing 
schistose as well as fibrous and spherulitic structures in glass tubing, by heat- 
ing it in presence of water up to a temperature of about 400° C., and under a 
pressure which he estimates at more than 1000 aOR (Etudes Synthé- 
tiaues de Géologie Expérimentale, p. 156 eé7 seq.). . 


DEVITRIFIED ROCKS FROM BEDDGELERT AND SNOWDON. 409 


as I can judge from microscopic examination of the eruptive rocks 
of the Llanberis Pass, the work which remains to be done 1s to fill 
in in detail, on a larger scale, the lines now laid down upon the 
1-inch map. 

I have already laid before this Society a description of a perlitic 
rock which occurs at the top of the Glyder Fawr; and I then ex- 
pressed my belief that many more devitrified rocks would yet be 
found among the felstones of paleeozoic age. 

The present paper is probably a very small contribution to this 
list; and it is to be hoped that, as the list increases, we shall learn 
more precisely what a felstone is, and realize more fully what many 
of the felstones once were. The vitreous lavas were probably closely 
allied to trachytes. Other eruptive rocks, also occurring in the 
Snowdon area, such as those of Llyn-cwm-y-ffynon and Pont-y- 
Gromlech, are of a decidedly basic character. Thus we see that, as 
it 1s now, so it was in the vastly remote period which we call Silu- 
rian. The eruptive rocks of that age were both basic and acid; and 
their constituent minerals and structural features were similar to, 
if not identical with, those which exist in, but do not specially cha- 
racterize the rocks erupted at the present day. 

In view of these facts I think we may disclaim any power to 
determine the age of a rock by its mineral constitution or structure, 
and may protest, as Mr. Allport has done*, against the employment 
of different names for similar or once similar rocks, which differ only 
in point of age. 


APPENDIX. 


On the Eruptive Rocks of Skomer Island f. 


Since the preceding paper was written I have examined some 
specimens from Skomer Island, off the coast of Pembrokeshire. For 
many years they have remained undescribed in the collection of 
rocks in the Museum of Practical Geology; and my suspicion of their 
true nature was first aroused by the close resemblance which one or 
two of them bore to other devitrified lavas which I had previously 
examined. Microscopic examination of these specimens shows con- 
clusively that they are lavas of a once vitreous character. The sedi- 
mentary rocks with which they are associated are regarded as 
belonging to the Llandeilo or to the Bala series. 

Only a short account of the microscopic characters of these lavas 
is here given, as they will be examined and described in greater 
detail in the forthcoming edition of the official catalogue of the rock- 
collection in Jermyn Street. 

The specimens about to be described consist chiefly of banded and 
spherulitic rocks, now felstones, but once obsidians, the change being 


_* “On the Microscopic Structure and Composition of British Carboniferous 
Dolerites,” Quart. Journ. Geol. Soe. vol. xxx. p. 565. 

+ The following particulars are now laid before the Society by permission 
of Prof. Ramsay, by whom also the specimens were collected. 


410 F. RUTLEY ON THE MICROSCOPIC STRUCTURE OF 


due to the usual process of devitrification; while in some instances 
it 18 probable that the rocks have developed a spherulitic structure 
throughout, during solidification, in which cases they must be re- 
garded as having been spherulite rocks from the first. Although 
none of these rocks retain a vitreous appearance, the minute struc- 
tures which are developed in them are perfectly preserved, and are 
as clearly demonstrable as they would be in the most recent 
lavas. 

The close resemblance in minute structure between these Skomer- 
Island lavas and those of the Yellowstone district in the United 
States is very striking, although their respective periods of eruption 
are so far removed from one another, namely—by all the time which 
elapsed between the deposition of the upper beds of the Lower Silu- 
rian series and certainly the lowest, if not the highest, beds of the 
Tertiary epoch. There is also a close resemblance between the 
Skomer-Island lavas and those of the Snowdon district. The period 
of eruption in both areas is nearly the same. 

In vol. 1. of the ‘Memoirs of the Geological Survey of England 
and Wales’ an account of the rocks of Marloes Bay and Musclewick 
Bay is given by Sir Henry De la Beche; but, although Skomer Island 
is situated close by, he makes no mention of it. At Wooltack Park, 
on the north of Marloes Bay, there are fossiliferous shales with some 
sandstones, beneath which it is stated that trap occurs resting on 
conglomerate and associated with shales. The whole of the igneous 
rocks of Skomer Island and the adjacent promontory have been 
mapped as greenstone. 

It is, however, evident that Sir Henry De la Beche generalized to 
some extent in the mapping of these rocks, as the following extract 
from the ‘Transactions of the Geological Society ’* will prove :— 

‘¢ All the north of Skomer Island consists of massive trap, having 
the character principally of fine-grained compact greenstone, and 
sometimes approaching to cornean. ‘The smail peninsula, however, 
to the east of the landing-place must be excepted, where some strati- 
fied rocks of ambiguous appearance occur. The southern part of the 
island consists of stratified greenstone dipping at about 48° to the 
south-east. Between this and the greenstone belonging to the north 
of the island a quartzose cornean, mostly striped, occurs. In some 
parts of the island hornblende is the prevailing ingredient of the 
rock.” 

In the same paper (at p..2) Sir Henry states that he considers 
these traps to be ‘forcibly intruded amongst the other rocks at a 
period subsequent to their consolidation;” and he adds that, in 
applying the term “stratified” to trap, he only means to imply 
‘“‘ that there is a parallelism of texture in the trap, which it has in 
common with a contiguous rock belonging to some other formation, 
and that this texture is also parallel to the common surface of sepa- 
ration between the trap and that other rock.” 


* Second series, vol. ii. p. 8. 


DEVITRIFIED ROCKS FROM BEDDGELERT AND SNOWDON. 411 


Whether the banded cbsidians, now felstones, are the “ stratified 
rocks of ambiguous appearance” is a point which ean only be de- 
cided by these who know the ground; or perhaps they may be the 
“striped quartzose cornean”’ alluded to as occurring between the 
northern and seuthern masses of greenstone which constitute the 
island. At all events, Sir Henry’s description clearly shows that 
he had not overlooked these rocks, although he does not appear to 
have regarded them as lavas, or to have considered them of sufficient 
importaice to indicate their position on the Survey map. 


Rocks from Skomer Island. 


1. Devitrified, banded obsidian. 

A compact greenish-grey felstone, petrosilex or hornstone. 

Under the microscope the banded structure is very clearly shown ; 
and a minute spherulitic structure also occurs in places. ‘The gene- 
ral condition of the rock is microcrystalline. A greenish substance 
is present in it, which appears to be chlorite. This rock is of much 
the same character as the devitrified obsidian or rhyolite from Clo- 
gwyn dur Arddu at the base of Snowdon. 

2. Devitrified, banded and spherulitic obsidian. 

A light greenish-grey to dark blackish-green rock. 

The specimen shows a weathered surface, upon which numerous 
fine bands stand out in relief. The bands are much contorted. 

‘Under the microscope the banding is well shown, and the sphe- 
rules are well defined and very numerous. A perlitic structure is 
also clearly seen ; and greenish matter in many cases pervades certain 
portions of the section. This green matter has in some cases a finely 
granular structure, or else contains fine dust, and exercises a weak 
depolarization when rotated between crossed Nicols. Spherules of 
much larger size than those which constitute the bands are likewise 
developed. They interrupt and appear to obliterate the fine sphe- 
rulitic bands, which seem abruptly cut off by them. The crystal- 
lization in these larger spherules is much more confused than that in 
their smaller representatives ; and consequently they show no dark 
cross between crossed Nicols. 

The structures in this rock are as perfect as any to be met with 
in recent lavas. 

3. Basalt or andesite. 

An iron-grey rock with brown stains, compact in texture and. 
showing some minute glistening felspar-crystals. 

Under the microscope the rock is seen to be a finely crystalline 
admixture of triclinic felspar prisms, granules of augite, and crystals 
and grains of magnetite. Some isotropic matter seems also to occur 
in the matrix. 

4, Quartz-oligoclase trachyte (?). 

A greyish rock, with small white porphyritic crystals. 

Under the microscope it is seen to consist of crystals of quartz 


412 F. RUTLEY ON THE MICROSCOPIC STRUCTURE OF 


and triclinic felspars (oligoclase) and orthoclastic felspar, with some 
magnetite in a fine microcrystalline matrix *. 

The rock No. 4 is possibly the crystalline representative of the 
devitrified obsidians. The basalt (No. 3) is evidently an example of 
the greenstone which occurs so extensively at this spot. Some of 
the obsidians contain large spherical bodies sometimes an inch or 
more in diameter, which are best shown upon weathered surfaces. 
As these spherules are traversed by the fine bands which pass 
through the rocks in which they occur, it seems reasonable to assume 
that they have been developed subsequently to the solidification of 
the rock. 


DESCRIPTION OF PLATE XXI. 


Fig. 1. Devitrified spherulitic rock from Beddgelert, showing general character 

of spherules and matrix. X18. 

2. Devitrified obsidian or rhyolite from Clogwyn d’ur Arddu, Snowdon, 
showing banded fluxion-structure. x18. 

3. Devitrified spherulitic rock, No. 1. Beddgelert, showing spherules and 
spherulitic bands. x6. 

4, Ditto, showing portion of a perlitic patch. X25. 

5. Ditto, showing parts of spherules at top and right side of field, and 
infiltered shrinkage-cracks in matrix. X12. 


DIscussIon. 


The Cuarrman spoke of the value of the papers ‘and the interest 
of the agreement between rocks separated so widely in time or space. 
Some of those described by the author might be paralleled by 
instances from Arran and from the Auvergne. 

Dr. Sorsy said the most interesting part of the paper was the 
close relation of the structures described on the one hand to those of 
granite, and, on the other, to those of slags. The relation in struc- 
ture between slags and the older rocks was of great interest. 

Prof. Bonney expressed his sense of the value of the paper. In 
his opinion the only difference that could be maintained between 
rhyolite and felsite (he thought felstone should be used only as a 
group term) was structure; he would understand by rhyolite a 
trachytic rock in which a glassy base remained—by felsite those 
in which the matrix was crypto- or microcrystallme. He knew 
some of the rocks described by the author, and some remarkable 
spherulitic rocks, one showing a structure just like that of Pont-y- 
Gromlech on the east side of the Glyder. 

Rev. J. F. Braxe asked about the formation of the crystals with 
an inward growth, and whether large crystals did not indicate slow 
cooling. 

Mr. BaverMman considered that the irregular strains in the obsi- 
dian fragment described by the author might be due to contraction 


* The section cut from this specimen was hastily examined on the day upon 
which this paper was read ; and I am not sure than it is not a clastic rock. 


DEVITRIFIED ROCKS FROM BEDDGELERT AND SNOWDON. 413 


consequent on the passage of a glassy into a crystalline substance of 
sensibly the same composition ; and instanced the inverse case of the 
fusion of felspar, where the increase of volume, in passing to a glass, 
fissures the crystal, in the direction of its principal cleavage, into 
laminge, which are kept together by the glassy cement. 

The AvrHor said that the Arran rocks, so far as he knew, were 
intrusive; he quite agreed it would be well to use the term felstone 
in a wide sense. Mr. Blake’s question had been answered by Mr. 
Bauerman, whose remarks on the fused felspar were of much im- 
portance. 


414 J. W. DAVIS ON THE FISH-REMAINS OF 


32. Nores on the Fisu-remarns of the Bonz-Bep at Aust, near 
Briston ; with the Descriprion of some Nuw GunERA and 
Species. By Jamus W. Davis, Hsq., F.G.S. &c., Hon. Secretary 
of the Yorkshire Geological Society. (Read May 11, 1881.) 


[Puate XXII] 


I am indebted for the material on which the following paper is. 
based to Mr. W. T. Ord, of Bristol, to the Council of the Geolo- 
gical Society at London, who have kindly placed their collection at 
my disposal, and to Mr. Sollas, Professor of Geology at the 
University College, Bristol. 

The specimens are in good preservation, the smaller ones, con- 
sisting principally of teeth, being unbroken; the larger bones, as 
for example, ribs or other bones of Saurians, the larger spines of 
Fishes, &c., are generally found in a more or less fragmentary and 
broken condition. 

From the occurrence of the teeth which are characteristic of the 
older Carboniferous rocks, such as Psammodus porosus, Helodus, 
and Psephodus magnus of the Mountain Limestone, and Cteno- 
ptychius, which has hitherto been found in the Carboniferous 
series, and more especially in the Coal-measures, it appears pro- 
bable that some of the fossil remains found in the Rheetic beds at 
Aust have been derived from the disintegration of the older rocks. 
Hither this must have been the case, or the genera of fishes named 
had a considerably. longer period of existence than has hitherto 
been supposed. It may be objected that the remains are in a very 
perfect state of preservation (as, indeed, they are in most cases) and 
do not appear to have been exposed to much attrition by being 
washed on the shore or bed of the sea or a lake. Itis probable, 
however, that the area over which the bone-bed was deposited was 
composed, in the neighbourhood of Aust at any rate, of the blue 
clays which at present underlie it. During the formation of the 
bone-bed the nodular masses of blue-grey stone which are now 
found composing a great proportion of its mass were pieces of 
clay, rolled round by the action of the waves or tides, so soft that 
they received easily an impression of the bones or teeth which lay 
scattered along the shore with them. From the immense number 
of fossil remains of Saurians and Fish which occur in the bed, it will 
be inferred that it required a long period of time for their accumu- 
lation, and that throughout all that time there was a peculiar absence 
of sedimentary deposits, the nodular masses being derived from the 
adjoining Keuper beds, which also formed the floor on which the 
bone-bed was deposited. 

In a paper read to this Society in 1841* Mr. Strickland showed 
the bone-bed to extend over a surface of 120 miles; and since that 


* Proc. Geol. Soe. vol. iii. part ii. p. 585. — 


THE BONE-BED AT AUST, NEAR BRISTOL. 415 


time Mr. Charles Moore, in an elaborate series of papers*, has 
proved with extreme minuteness and care that there extended over 
the Somersetshire and South Wales Coal-fields, and over the 
Mountain Limestone, beds containing Rhetic fossils, the crevices, 
veins, and pot-holesin the Mountain Limestone being filled up with 
organic reliquiz of vertebrates as well as invertebrates of Rheetic age. 
These, according to Mr. Moore, have been washed in during the Rheetic 
and subsequent Liassic periods. Such being the case, the circumstances 
attending their deposition warrant the supposition that some of the 
fossils included in the Rheetic deposits were derived from the 
disintegration of the rocks on which they now rest. The bone-bed is 
variable in thickness, but rarely exceeds a few inches ; at Aust it is 
from 3 in. or 4 in. to 9 in. thick; in many other places it is less, 
ranging over considerable areas with a thickness not exceeding one 
inch. There is also much diversity in the prevalence of organic 
remains. In a few localities the fish-remains occur in abundance, 
as at Aust, Axmouth, Coomb Hill, &c., whilst in other places they 
are entirely absent or are found very sparingly. 

The fishes found in the bone-bed are comprised in the orders 
Plagiostomi and Ganoidei, the latter including, according to 
Prof. Miall t, the Ceratodus-remains. Besides Ceratodus, the Ganoids 
include the genera Saurichthys, Gyrolepis, Lepidotus, and Ambly- 
pterus. 

Amongst the Plagiostomous genera may be enumerated Hybodus, 
Acrodus, Sargodon (?), Nemacanthus, Sphenonchus, Lophodus, Squalo- 
raid. 

There are also large numbers of bones, teeth, and other remains 
of Saurians, including Ichthyosaurus, Nothosaurus, Scelidosaurus, 
and others found and identified by Mr. Moore ¢. 

A consideration of the characteristics of this mixed group of 
organic remains may afford some reasonable basis for deductions as 
to the circumstances under which they were accumulated. The 
Saurians would undoubtedly exist near and partly on land. The 
Ceratodonts, judging from a comparison of the ferms of their teeth 
with the still existing Ceratodus of Australia, were vegetable 
feeders, and would require a shallow-water area from which to 
obtain their food. With respect to the remaining Ganoids, it is 
probable that they could freely exist in deeper water, and, from 
the character of their teeth, were probably predaceous in their 
habits. The Plagiostomous Sharks, with their sharp teeth and 
strong fin-spines, often attaining a large size, as evidenced 
by the great length of the latter, would be equally adapted 
for either deep or shallow waters. Taking all the items 
together, it would appear that the Rhetic beds were deposited in 
a shallow sea not far from the coast; that the Saurians passed a 
large proportion of their existence in the water, the remainder on 


* Quart. Journ. Geol. Soc. vol. xvi. p. 483 (1860), vol. xxiii. p. 449 (1867), 
vol. xxvii. p. 67 (1881). 

t Palzontographical Soc., vol. xxxii. (1878). 

t Loce. cité. 


416 J. W. DAVIS ON THE FISH-REMAINS OF 


the land; of the fishes, the Sharks and some of the Ganoids 
frequented the shallow waters, probably in search of food. That 
they did so inlarge numbers, and spent considerable time there, is 
amply proved by the large number of coprolites. 

Microscopical sections of the coprolites exhibit abundant evidence 
that the food of the Fishes or Saurians to which they owe their 
origin consisted of smaller fishes—fragments of bone, teeth, and 
other similar objects belonging to smaller or more slightly armed 
species of fish being found in larger proportions than any thing else 
with structure. : 


Hyszopvus avstirnsis, Davis. (Plate XXII. fig. 1.) 


In the third volume of the ‘ Poissons Fossiles’ M. Agassiz described 
aspine of Hybodus, of which there were several specimens in the 
cabinets of Lord Enniskillen and Sir P. Egerton, and in the museums 
at Oxford and Bristol; and an example is figured from the latter, which 
is about 6 inches in length. ‘The spine is described as being round 
(that is, the sides are not so much compressed); and the ribs are 
more prominent, with deeper grooves between than in any other 
species described. The ribs do not anastomose, but run parallel to 
the anterior portion of the spine, and disappear along the posterior 
margin. There isa large cavity along the posterior surface; and the 
base of the spine is large proportionally to the remainder. The 
examples seen by M. Agassiz did not exhibit any traces of den- 
ticles sufficiently well preserved to enable him to describe them. 
Dr. Buckland and Sir Henry de la Beche knew ‘the spine, and had 
previously considered it a variety of their [chthyodorulites dorsetiensis 
(= Hybodus reticulatus, Agassiz) found at Lyme Regis. M. Agassiz 
further remarks that H. minor is not a small spine when compared 
with others he had described; but, at the same time, the spines 
found at Aust Cliff are not of large size, and are very different from 
the great spines found in the Lias at Lyme Regis. The teeth 
accompanying the spines are also different from those found at Lyme 
Regis; and altogether the differences are so great as to necessitate 
the institution of a new genus. 

Since M. Agassiz wrote his great work, many fossil spines of 
Hybodus have been found in the Rheetic beds of Aust; and the 
collection at the Bristol Museum contains specimens which very | 
nearly approach the sizes attained by the fine examples of Hybodus 
reticulatus and formosus of the Lias of Dorset. One of the largest 
examples would measure, when perfect, 13 inches in length, and 
fully aninch and a quarter in greatest diameter. The base pre- 
sents the usual fibrous structure ; it has been deeply and strongly 
implanted in the flesh. There is a large orifice or groove opening to 
the back of the spine at its base, and afterwards continued as an 
internal cavity towards the apex. The width of this orifice, from 
back to front, is about double that between the sides of the spine. 
The line dividing the base from the exposed portion extends with 
a convex curve from the anterior to the posterior portion of the 
spine, the convexity being towards the base. The anterior and 


THE BONE-BED AT AUST, NEAR BRISTOL. 417 


lateral surfaces are ornamented with strongly marked ridges of 
enamel, which anastomose frequently, and present a somewhat 
wavy outline on account of the enamel forming the ridges not 
being of uniform thickness, in some places presenting a beaded 
appearance. The ribs lie roughly parallel with the anterior surface 
of the spine, and run out along the posterior edge, but not in a 
very well defined manner. The junction of the ribs with the basal 
portion is not so well defined as in WH. reticulatus: in the latter 
the ends of the ribs rise above the surface of the base; whilst in 
the Aust specimen the ridges blend with the fibrous structure of 
the base, and the hollows between the ridges are below the level 
of the basal surface. The anterior and lateral portions of the spine 
have a circular or, rather, dome-shaped form in section. The 
posterior forms a wide base, not flat, but a little produced outwards 
towards the centre. The whole of the posterior portion higher 
than the open part of the cavity retains the fibrous structure of 
the base; and along each side of the median portion extends a row 
of blunt, laterally compressed, enamelled denticles; they are about 
-2 inch across the base, and rise *l inch from the surface of the 
spine. ach denticle is separated from the next by a distance a 
little greater than its own diameter. 

These spines differ from those of H. reticulatus in several particu- 
lars: their form is stronger and more robust ; the lateral surfaces are 
rounder, and the base and cavity wider. In comparison with its 
width, the spine is shorter, not so gradually pointed, and less curved. 
The posterior denticles are not pointed and recurved towards the base, 
as they are in 4. reticulatus; nor do its characters agree with 
those ascribed by M. Agassiz to H. minor, as already stated ; and it 
appears necessary that a specific name to indicate this spine should 
be instituted: I suggest that it be Hybodus austiensis. 


Hyszopus punoratus, Davis. (Plate XXII. fig. 2.) 


An imperfect spine from the bone-bed possesses characters which 
remove it from any other species hitherto described. It consists 
of the upper portion of a small spine 1 inch in length; the 
antero-posterior diameter is ‘2 of an inch at its broadest part; 
the transverse diameter is a little less; the spine is slightly 
curved, more so on the anterior than the posterior surface, and 
tapers gradually to a point. The lateral surfaces are covered with 
longitudinal ridges, separated by grooves of about equal diameter, 
numbering five on each side. Along the bottom of each groove 
there are a number of minute pittings extending in a line 
parallel with the groove. ‘Towards the points the ridges become 
less prominent and gradually disappear, the apex being quite 
smooth and without strie. There is a row of denticles along 
each latero-posterior surface ; they are prominent, obtusely pointed, 
laterally compressed. The internal prolongation of the pulp-cavity 
is comparatively small (less than one third the diameter of the 
spine), situated nearer the posterior than the anterior surface, and 
conforming in outline with the oval form of the spine. 


418 J. W. DAVIS ON THE FISH-REMAINS OF 


Besides Hybodus mimor, the only spine of this genus described 
by Prof. Agassiz from the Aust bed was H. leviusculus*. The 
original which served for the description of the latter was a small 
fragment about half an inch in length, in the museum at Bristol. 
It is described as having smooth sides, slightly compressed, with an 
internal cavity rounder than the external form of the spine. There 
are denticles along the posterior border ; and these are long, pointed, 
and recurved towards the base. 

The spine I have from Aust, the only other small form of 
Hybodus which I have seen, differs in every respect, except size, 
from H. leviusculus of Agassiz. Its sides are deeply furrowed; the 
posterior denticles are short and blunt; and the internal cavity is 
much longer than broad in section. 

The number of well-defined species of teeth of Hybodus found in 
the Rhetic beds of Aust would lead to the inference that there 
should be asimilar variety in the fin-defences of the fishes. Under the 
most favourable circumstances it is an extremely difficult matter to 
correlate the dermal defences, either spines or scutes, of the Sela- 
chians with the teeth of the same genus; but in this instance the 
difficulties are greatly increased by the rolled and mixed state in 
which the specimens are found; and it appears improbable that 
remains will be discovered whose relationship will be rendered 
certain by the position or circumstances under which they are dis- 
covered. ‘The pittings along the grooves suggest the name punctatus 
as appropriately designating this spine. 


Remarks on the Genus NeMacantuus, Ag. 


This genus was formed to embrace two species of fossil spines of 
Selachians found in the bone-bed at Aust. The spines are about 5 
or 6 inches in length and °7 inch in breadth in the larger species, viz. 
N. monilifer, and little more than half that size in the second one, 
N. fiufer. The genus is characterized by the spine having its sides 
much compressed, and finely striated, with a small posterior cavity 
reaching half the length of the spine ; where the cavity terminates on 
the posterior surface there commences. on the sides a number of 
rounded tubercles: they originate near the anterior surface, extend 
obliquely across the spine, and run in parallel lines thence to its 
apex, a row extending along the junction of the lateral with the 
posterior face having some resemblance to a row of small blunt 
tubercles along each side. Along the anterior portion of the spine 
there extends a round keel, which is marked off from the body of 
the spine by a lateral canal along each side. In the larger species 
the lateral keel is of about the same diameter as the tubercles arranged 
along the sides. The smaller species, NV. filifer, differs from the 
larger, V. monilifer, in the tenuity of its anterior keel, the smallness 
of the tubercles on its sides, and also of those extending along the 
posterior edges. 


* Poissons Fossiles, tome iii. p. 46, tab. 10. figs. 24, 25, 26. 


THE BONE-BED AT AUST, NEAR BRISTOL. 419 


In the species described by M. Agassiz the spines are as nearly as 
possible twice as much in their antero-posterior as in in their trans- 
verse diameter. In a specimen from the Bristol Museum (Plate 
XXII. fig. 3) the spine from back to front has a diameter of °35 inch ; 
and its width across the posterior surface is *3 inch, or very 
nearly equal to the antero-posterior diameter; the external pos- 
terior groove is shallow; and no denticles or tubercles are present. 
There is a very large median keel along the front of the spine. It 
is almost round, but rather wider than deep. It is composed of 
shining black enamel, and constitutes one fourth of the entire dia- 
meter of the spine. ‘The spine is imperfect, the apical portion 
missing ; so that itis impossible to say whether the sides were tuber- 
culated. It appears to have been less curved than the common 
forms, the portion preserved being straight. 

A second specimen, more closely resembling WN. filifer, is a flat-sided 
spine of the ordinary kind (Plate XXII. fig. 4), finely striated along 
each lateral face. ‘The anterior keel is small and threadhke; along 
the posterior surface are blunt, widely separated denticles. This 
spine is ‘25 inch in diameter, and, when perfect, would probably be 
about 2 inches long. The sides of the spine are covered with 
longitudinal striz as in the type specimens. 

Both the varieties noticed above are from the collection of Mr.Ord, 
of Bristol, and were collected from the bone-bed at Aust Cliff. 


Nemacantuvs minor, Davis. (Plate XXII. fig. 5.) 


Spine imperfect. Length 1:1 inch, diameter 1 inch, when 
perfect probably nearly or about 2 inches in length. In section 
it is circular. A canal or internal cavity of similar form ascends 
the centre of the spine towards the point (the latter broken off in 
this specimen). There is no evidence that the cavity was open 
along the posterior surface; but it appears to have been terminal. 
The spine is'slightly curved in form. Its surface is slightly and 
irregularly grooved, and is further ornamented by a number of mi- 
nute papille. In the latter respect the spine resembles the genus 
Nemacanthus ; but it differs in other essential respects from either of 
the two species described by Prof. Agassiz*. It does not exhibit 
any trace of having a ridge of any kind along the anterior 
surface ; instead of that, itis round and indiscriminately spotted 
with papilla or tubercles. The section of the spine is round, and not, 
as in the species of Agassiz, oval or triangular. It appears proba- 
ble that the spine may belong to the genus Memacanthus; but it is 
quite separated specifically. I propose the name JN. minor in allusion 
to its small size compared with those previously described. 

The specimen figured is in the museum of the Geological Society 
and is labelled “from the Fucoid bed, Wainlode Cliff,” and was 
presented to the Society by H. E. Strickland, Esq. 


* Poissons Fossiles, vol. iii. p. 26, tab. 7. figs. 9 & 10-15. 


420 J. W. DAVIS ON THE FISH-REMAINS OF 


Patmosaurus? Srrickianpi, Davis. (Plate XXII. fig. 6.) 


In the museum of the Geological Society, London, there is a tooth 
which I believe to be unique. It is from the Rhetic bone-bed of 
Combe-Hill, near Cheltenham, and was presented to the Society 
many years ago by Mr. H. E. Strickland. 

The base of the tooth is wanting ; the portion remaining is slightly 
more than ‘6 of an inch in length. In section (fig. 66) the 
front portion is seen to be more compressed than the back. The 
lateral extremities of the tooth are slightly produced, and end in a 
serrated margin ascending to the crown or apex of the tooth, which 
issmooth. The width nearest the part of the base preserved is -35 
inch. The surface of the tooth, except along the lateral margin, is 
covered with fine longitudinal striations, which finally disappear 
before reaching the point. 

This tooth has the appearance of having been washed and water- 
worn. The broken portion is smooth and polished ; and it is probable 
that it may have been derived from an older rock, and redeposited 
amongst the remains of the Fishes and Saurians of the Rheetic age. 

At a meeting of this Society held December 15th, 1841*, a paper 
was read by Mr. H. E. Strickland on the Bristol bone-bed, in which, 
along with other fossil remains, he mentions a “ portion of a tooth 
with two finely serrated edges, and considered as probably belonging 
to a Saurian allied to the genus Palwosaurus ;” there can be little 
doubt this is the same specimen. I have taken the hberty of asso- 
ciating the tooth with his name. 


SpuEeNoncuus (Hyzopvs) ostusus, Davis. (Plate XXII. fig. 7.) 


The genus Sphenonchus was originated by M. Agassiz for the de- 
signation of certain objects which are regarded as teeth. It appears 
probable, as I shall attempt to show, that these opjects were not 
teeth, but dermal defences. The specimen I am about to describe is 
‘45 inch in length and ‘35 broad at the base: it is perfect, with 
the exception of a small fragment which is broken from the right 
portion of the base. The object, which appears homogeneous in 
structure, contracts rapidly from the base for a distance of about half 
its length. At this point its diameter is little more than a third 
that of the basal portion; and it remains the same to within a little 
of the point, which is slightly wider and thinner than the stem. 
The general form is that characteristic of the genus. It is arched 
forward, and does not possess any secondary denticles. The upper 
portion is nearly cylindrical, with the point flattened out like a 
chisel; near the base the form is three-sided; right and left it is 
produced so as to form wing-like processes; whilst down the centre 
of the anterior curved portion a third process is developed, which is 
continued and increases in size to the lowest portion of the base. 
The posterior surface, in its basal portion, is slightly hollowed in- 
wards (fig. 76), compensating a little for the ridge in front. The 
whole of the surface is smooth and covered with shining enamel. 


* Proc. Geol. Soc. vol. iii. pt. 11. p. 585. 


THE BONE-BED AT AUST, NEAR BRISTOL. 421 


Agassiz, in the third volume of the ‘ Poissons Fossiles,’ describes 
three species of this genus :—S. hamatus, from the Lias of Lyme 
Regis, in the collection of the Karl of Enniskillen; S. elongatus, 
found by Dr. Mantell in Tilgate Forest; and S. Martini, Rob., from 
the Portland Oolites of Linksfield. These species vary greatly in form 
and general appearance; but in each the generic characters are well 
developed. ‘The cylindrical cone forming the upper tooth-like 
portion is bent over anteriorly, the basal portion spread out in 
aliform processes; and a third median ridge or process is developed 
from the anterior surface of the base. The specimen I have de- 
scribed from Aust shares these peculiarities ; but in detail it appears 
sufficiently distinct to form a new species. It is also from a ho- 
rizon earlier and lower than any of those mentioned above; and 
though it may probably be found necessary to modity the specific 
relationship of these objects when more is known of them, at 
present evidence is wanting to indicate that they are not distinct 
species. 

Sphenonchus hamatus, Agass., is remarkable for its extremely 
arched form and the pointed termination of the tooth-like part: it is 
an inch in length; and its base is widely expanded. S. Martini, Rob., 
is somewhat similar to S. hamatus, but is shorter and less curved. 
The third species, S. elongatus, is much larger than either of the 
others; the cylindrical portion is expanded at first, contracting nearer 
the extremity, and ending in a second expansion, “en sorte que sa 
forme ressemble un peu 4 celle d’une bouteille qu’on aurait recour- 
bée.” Sphenonchus obtusus differs from S. hamatus in its smaller size, 
less expanded base, and in its curvature being at a considerably 
smaller angle: its apical termination, wide, flattened and obtusely 
rounded, is in marked contrast to the finely pointed end of S. hama- 
tus. S. obtusus has altogether a finer and less stumpy form than 
that of S. Martini, whilst from S. elongatus it is easily distinguished 
by its smaller size and the nearly uniform diameter of the upper 
portion. 

I suggest the specific name obtwsus to designate this species, in 
reference to its wide and expanded apex. The specimens are from 
the cabinet of Mr. Ord. 

Prof. Agassiz described the genus Sphenonchus as a member of the 
family of Hybodontes, associated with Hybodus, Cladodus, and Diplodus. 
Since the classical work of Agassiz was completed, Diplodus has been 
proved to be the tooth of Plewracanthus, and must consequently be 
remoyed from the family of Hybodontes, and considered in con- 
nexion with the spine as nearly associated with the recent Rays, 
according to Agassiz; or, as I have attempted to show in a former 
communication, it may have some affinities with the group of the 
Siluroids. Sphenonchus and Diplodus are referred to in the ‘ Pois- 
sons Fossiles’ as offering considerable difference in microscopical 
structure from Hybodus and Cladodus, especially the former, which, 
whilst having a dense coating of dentine, has a large internal pulp- 
cavity, which is very different from the tooth-structure of either 


Q.J.G.8. No. 147. 2F 


499 , J. W. DAVIS ON THE FISH-REMAINS OF 


of the others. In the Magazine of Natural History* Mr. Charles- 
worth has described the fossil remains of a species of Hybodus, col- 
lected by Miss Anning at Lyme Regis. In connexion with the 
spine and teeth there is a bone which is undoubtedly an example of 
Sphenonchus, and is regarded by Prof. Agassiz in that light; im the 
same volume (p. 605) there is a letter from Miss Anning, saying . 
that the hooked tooth (Sphenonchus) is by no means new, but that it 
has been frequently found at Lyme Regis in connexion with the 
teeth and spines of Hybodus or the teeth of Acrodus. 

Mr. Charlesworth suggested that the hooked tosth is a dermal 
appendage or defence, probably situated immediately behind the 
head. Specimens since discovered prove that this suggestion was 
correct, and that two or three of these bodies were located on 
the occipital region of the head of Hybodus. The microscopical 
structure of the teeth and dermal defences of the Klasmobranch 
fishes is very similar, one of the principal differences being the 
large size of the pulp-cavity of the dermal processes as compared 
with that of the teeth ; and in this respect Prof. Agassiz has shown 
that there is a great difference between the teeth of Hybodus or 
Cladodus and Sphenonchus. The base of Sphenonchus is excessively 
expanded, especially in S. hamatus from the Lias; and its fibrous 
structure without enamel indicates that it was imbedded in the 
flesh ; it appears in this respect to resemble the dermal defences of 
Raia clavata from the Tertiary deposits. 

In addition to the specimens already named, there are a number 
of the bones of the head, including jaws with teeth, of Saurachthys ?; 
these I have handed to Mr. Sollas, who has already in preparation a 
paper on the same genus, derived from the examination of similar 
Specimens in his possession. 

Large numbers of fragmentary bones and small teeth are found 
scattered throughout the mass of the bed, but without sufficiently 
well established characters to enable an account to be given of them. 
A large operculum, nearly 24 inches in diameter, probably belong- 
ing to Ceratodus, and several bones with articular extremities, 
which may belong to the same genus, are included in the collection 
of Mr. Ord. 

Besides the palatal teeth of Psammodus and Cochliodus, in all 
probability derived from the Mountain Limestone which underlies 
the Rhetic beds in some parts of the area, the following species of 
Ctenoptychius have probably been derived from the Coal-measures, 
and the Petalodus from within the Coal-measures or the Limestone. 
Specimens of Helodus have also been met with; they appear to be- 
long or are very nearly related to H. semplew. 


Crenoptycuivs Orpi1, Davis. (Plate XXII. fig. 8.) 
Tooth. Length -55 inch. Depth -3inch. A portion of the base 


is wanting. 
The superior surface is folded so as to form a pectinated edge 


* Vol. ili. p. 242, 1839. 


THE BONE-BED AT AUST, NEAR BRISTOL. 493 


extending along the crown, which occupies the greatest diameter of 
the tooth. It is slightly circular towards each end, the central 
part being almost straight. It is very thin, and appears to have 
constituted a sharp cutting-edge. The foldings are produced at their 
extremities into small and separated denticles, about 24 in number, 
whose diameter is greater from back to front than laterally. Some 
of these are broken at the tips; and the section thus exposed, when 
magnified, shows that a hollow tube ascended in the centre of each. 
The sulci descending to the body of the tooth from the denticula- 
tions are much more marked towards the lateral extremities than in 
the median region. Towards the base the tooth becomes gradually 
thicker ; at the same time it also converges laterally to two thirds the 
diameter of the crown; from this part the tooth is broken off; but, 
from the impression on the matrix, it appears to have terminated in 
a broadly expanded rounded base. The tooth is attached to the 
matrix; and consequently the posterior surface is not exposed. The 
whole of the upper part of the anterior surface, above the root or 
base, 1s covered with a smooth polished surface of ganoine. From the 
base of the plications or foldings forming the crown of the tooth the 
surface extends towards the base in the form of a semicircular 
hollow. 

This genus of Selachians was instituted by Agassiz (Poiss. Foss. 
tom. 1. p. 99) for the accommodation of teeth obtained from the 
Coal-measures of Staffordshire and Lancashire. Since that time 
specimens have been found in the Limestone of Armagh, and also 
in the Coal-measures:and Limestones of Virginia, Illinois, and other 
localities in America. Hitherto specimens of the genus have been 
restricted to the Carboniferous group of rocks, 

The specimen now described may either have been derived from 
the disintegration of coal-measure strata, and washed into the 
Rhetic beds during their deposition; or it may have belonged to 
a fish which lived during the period when those deposits were 
accumulating. It is probable that the former is the correct sup- 
position. 

The species from Aust differs materially from the type species of 
Agassiz, C. apicalis. The latter is possessed of only seven or eight 
protuberances from the crown of the tooth, the centre one being 
considerably larger and forming an apex to those on either side. 
C. semicircularis, N. & W., irom the Coal-measure limestone of Ohio, 
bears a remarkably close resemblance to C. apicalis of Agassiz, 
and seems to be so little removed as scarcely to necessitate a sepa- 
rate specific name. Ctenoptychius Ordw bears some resemblance 
to C. denticulaius, Agass. (loc. ct. p. 101), in possessing a large 
number of serrations closely ranged along the crown of the tooth, 
whose lower portions form a series of plications extending to the 
body of the tooth and there disappearing; but in C. denticulatus the 
curvatures extend quite straight across the crown, whilst in the 
Aust specimen they form a semicircle. The base of the tooth in 
C. denticulatus is also much wider than in C. Ordii. 

Ctenoptychius serratus, Ord (Sedgw. and M‘Coy, Brit. Pal. Fos- 

2F 2 


424 J. W. DAVIS ON THE FISH-REMAINS OF 


sils &c. p. 626, pl. 3. i. figs. 21, 22, 23), from the limestone of 
Armagh, bears a greater resemblance to this specimen than, perhaps, 
any others; but it may be easily distinguished by the greater 
breadth of the crown of the tooth compared with its depth, the com- 
paratively wide and short, somewhat cone-shaped character of the 
denticles, and the tips of such denticles appearing minutely crenu- 
lated under the lens. 

The specimens serving for the above descriptions were collected at 
Aust by Mr. Ord, and are from his cabinet. In recognition of his 
energetic and painstaking labours, I have ventured to employ his 
name to distinguish this species of Ctenoptychius. 


CrENOPTYCHIUS PECTINATUS, Ag. 

A second species of Ctenoptychius has also been found at Aust, 
and forms a part of the collection of Mr. Ord, of Bristol. It is -2 
inch across the crown, and has a depth from the upper edge of 
the crown to the base -15 inch. The crown is composed of 
about 12 denticles, the tips of which extend almost in a straight 
line; at the apex the denticles terminate in short pointed cones. 
Towards the body of the tooth these speedily coalesce and form 
plications extending more than half the distance towards the base; 
they form a concave surface anteriorly, the base becoming thicker 
and convex. The two outermost denticles, stronger than the re- 
mainder, are continued in a semicircle, and converge towards the 
base. The whole of the exposed surface is covered with a black, 
shining enamel. 

In the third volume of the ‘ Poissons Fossiles’*, M. Agassiz 
describes and figures under the name of Ctenoptychius pectinatus, 
some small teeth obtained from the Burdie-House Limestone. The 
specimens figured present very considerable variations in form, but are 
each characterized by the denticles forming the crown of the tooth 
being less distinctly separated towards the apex, and the points, 
instead of extending vertically, as in C. apzcalis, diverge or radiate 
from the centre, somewhat in theform of a fan. The beautiful little 
specimen from Aust agrees sufficiently well with this description to 
warrant its inclusion in the same genus and species. Its more 
rounded basal termination is the most striking point of divergence, 
C. pectinatus haying a more contracted stem-like or prolonged basal 
portion. 


Crapopus curtus, Davis. (Plate XXII. fig. 9.) 

This tooth offers some peculiarities which give it a distinctive 
character. Itisimbedded in the matrix ; and on the lower part it is 
slightly imperfect. It consists of a wide and thick base, from the 
centre of which springs a cone-shaped cusp. On either side the 
central cone there are indications that at least one secondary cusp 
has existed. The bony structure is very dense, and now of a deep 
brown colour. The length of the base is -4 inch; and the 
height of the central cone is ‘3 inch. The latter is rather less 


* Page 100, tab. 19. figs. 2, 3, 4. 


ean 


THE BONE-BED AT AUST, NEAR BRISTOL. 425 


than °2 inch in diameter, and it ends in a rounded apex; rising 
from the body of the tooth, it first curves a little outward and 
backward, and then again bends forward, the end projecting, as 
shown in fig. 10a, Pl. XXII. It appears to be thin in proportion to 
its width, and has quite a tongue-shaped appearance. The surface 
of the part of the tooth forming the cone is deeply indented with 
numerous pituoles. The tip is smooth and is covered with a thick 
coating of ganoine. The basal line of the lower portion of the tooth 
is curved inwards from each lateral extremity ; and from the cone 
the tooth swells with a well-rounded forward curve to the base. 
Near each lateral extremity of the base a part has been broken away 
with the matrix. The portion left exhibits the base of a secondary 
denticle or cone, in the centre of which is the cavity which as- 
cended towards its point. The secondary denticles were placed 
forward in comparison with the principal centre one, and were 
rounder in section. 

This specimen more resembles Agassiz’s species C. marginatus 
than any other; it may be easily distinguished, however, by the 
absence of the deeply striated surface which characterizes the 
latter and by its shorter base. I suggest the nomen triviale C. 
curtus. 


EXPLANATION OF PLATE XXII. figs. 1-9. 


Fig. 1. Hybodus austiensts, Davis. 
2. Hybodus punctatus, Davis. 
2a. Transverse section. 
3. Nemacanthus monilifer, var. a. 
3a. Transverse section. 
4, Nemacanthus monilifer, var. 3. 
4a. Transverse section. 
5. Nemacanthus minor, Davis. 
5a. Portion of spine, magnified. 
5b. Transverse section. 
6. Paleosaurus Stricklandi, Davis. 
6a. Tooth, magnified. 
6. Transverse section, nat. size. 
7. Sphenonchus obtusus, Davis. 
7a. Ditto, magnified. 
76. Antero-lateral form. 
7c. Posterior view. 
7d. Postero-lateral form. 
8. Ctenoptychius Ordit, Davis 
8a. Longitudinal section. 
9. Cladodus curtus, Davis. 
9a. Longitudinal section. 


Discusston. 


Prof. SreLry remarked on the curious survival in the Aust deposit 
of Paleozoic types of fishes mingled with forms peculiar to the 
Mesozoic as offering a parallel to the mixture of Paleozoic with 
Secondary Mollusca in the Upper Trias of the Austrian Alps. He 


426 ON FISH-REMAINS FROM BONE-BED AT AUST, NEAR BRISTOL. 


stated that Sphenonchus always occurs on the head of Hybodont 
fishes in the Secondary rocks. 

Mr. Tawnzy thought the Paleozoic forms in the Aust bed are 
fossils derived from the Carboniferous strata, and not, as Prof. 
Seeley contended, surviving types. 

Mr. Lonek stated that there is a great difference between the 
contents of the bone-bed at Garden Cliff at Westbury and Aust Cliff 
respectively. He thought the fossils in the bone-bed do not belong 
to the period at which the materials were accumulated. 

Mr. Ussuer thought that the working-out of the Rhetic beds 
might afford evidence of the unconformities due to changed condi- 
tions of deposit. He alluded to an appearance of unconformity 
between Rheetic and Trias at Newark, where a thin band of derived 
fragments occurred at the base of the former. 

Rev. H. Wiywoop argued against the theory of the bone-bed 
being a remanié bed from the fact of the delicate fossils in it not 
pone waterworn. 

The Prestpent thought that most of the fossils at Aust Cliff and 
Westbury are not remanié, but are of the true Rhetic age; but 
some Carboniferous forms are undoubtedly derived. The specimens 
of Ceratodus were not at all worn. The form called Sphenonchus 
is certainly the head-spine of Hi yoda as proved by specimens 
from Lyme Regis. 

The ArHoR. stated that the details that he had given represented 
the examination of only three collections; and that there yet remained 
much work to be done in this field. 


Quart Journ. Geol. Soc Vol. XXXVI PLA 


Mintern Bros. ump 


AS. Foord del etlith 


=MAINS FROM THE AUST BONE-BED 
FISH BEM NE CDAL-MEASURES 


ae) 
wh 
Sie 
, 
* 
5 : s 
ae 
; 
5 
. 
. 
* 


J. W. DAVIS ON ANODONTACANTHUS, 427 


33. On ANODONTACANTHUS, @ NEW Gunus of Fosstt Fisnus from the 
Coat-MEASURES ; with Descriptions of three New Sprcres. By 
James W. Davis, F.G.S. &. (Read May 11, 1881.) 


[Pirate XXII, ] 


THe spines which form the subjects of the following descriptions 
are from the West-Riding Coalfield in Yorkshire, and the Ironstone 
occurring in the Lower Limestone series near Edinburgh. A strati- 
graphical description of the Cannel Coal at Tingley, from which the 
first two species have been obtained, may be consulted in the 
Quarterly Journal of the Geological Society issued for February 
1880, page 56. 

The three species of Ichthyodorulites forming the subject of this 
paper are the only specimens of which I have cognizance possessing 
their peculiar characteristics. In some particulars they resemble 
the genus Pleuracanthus; they are composed of a strong, close- 
grained, fibrous substance. ‘Their general form is also similar to 
that of the Pleuracanths: the internal cavity is terminal at the basal 
extremity ; and the walls become thinner near the base in comparison 
with the remainder of the spine; the method of implantation in the 
body of the fish must also have been the same. The great distin- 
guishing feature between the two rests an the absence in this genus 
of any form of denticulation. In the Pleuracanths there are two 
rows of denticles either along each lateral surface or on some portion 
between the lateral and posterior lines. Whether this may be a 
sufficient distinction to necessitate a second genus, or further ‘dis- 
coyeries will prove that it must be included in the genus Plewra- 
canthus, remains to be seen; but for the present it may be better 
to consider it as a distinct genus, for the following reason : hitherto 
remains of the fish Plewracanthus, where found in even tolerable 
perfection, as in the Stone-coal of Bohemia, have always had associ- 
ated together the three-pronged teeth of the so-called Diplodus, and 
a spine, the latter more or less straight, but always possessing, in one 
situation or another, two rows of denticles. From Tingley, where the 
spines under discussion have been obtained, I have several slabs of 
Cannel Coal covered with a mass of cartilage full of the mosaic-lke, 
minute, rhomboidal, bony centres characteristic of Plewracanthus, 
with numbers of teeth, and occasionally a spine in situ; but in all 
cases the spine has been denticulated. So far all the evidence goes 
to show that the spines of Plewracanthus were armed or ornamented 
by these denticular appendages; there is a possibility, however, that 
a specimen may be found with a spine without denticles in association 
with undoubted Plewracanthus-remains ; and should that happen, it 
will be necessary to modify the character of the genus so as embrace 
thisone. Until such evidence is forthcoming, it may be regarded as a 
distinct genus, for which the name Anodontacanthus may not be in- 
appropriate, signifying a toothless spine. 


428 J. W. DAVIS ON ANODONTACANTHUS, A NEW GENUS 


1. Anopontacantuvs acutus, Davis. (Plate XXII. fig. 10.) 


Spine. Length 2°5 inches, breadth :2inch. The spine is straight, 
and gradually tapers to a point; there are no denticles; and 
the surface of the spine is uniformly covered with little pittings, 
giving an appearance of reticulation. The walls are about one 
fourth the diameter of the spine in thickness; there is an in- 
ternal cavity with a wide open termination at the basal extremity, 
the walls as they approach the base becoming much thinner. The 
cavity extends along the interior of the spine almost to its apex. 
The spine has every appearance of having originally been circular in 
form. It has, however, become somewhat crushed along the basal or 
weaker portion of the spine since its deposition. The specific desig- 
nation acutus expresses its pointed character. 

Locality. Cannel Coal, Tingley, Yorkshire. 


2, ANODONTACANTHUS oBTUSUS, Davis. (Plate XXII. fig. 11.) 


A spine from the same locality and horizon as the one already 
described is of larger dimensions and different form ; the basal end 
is broken off; the remaining portion is 2°5 inches in length, and the 
diameter nearly 0°3 inch. Judging from the proportions of the 
spine preserved, it is probable that, including the missing portion, 
its length would be 3:50 inches when perfect. The surface is uni- 
forinly covered with minute longitudinal striations, which towards 
the apex become broken up into small pore-like indentations, the 
latter being finer than in the smaller species. The portion of the 
spline preserved maintains a uniform thickness to within an inch 
of the apex, where it becomes slightly contracted, and ends in a 
broad, flattened, obtusely rounded extremity, 0°2 inch across. The 
body of the spine is oval in section, with an internal canal occu- 
pying one third of its diameter. The spine differs from the one de- 
scribed before in its greater size, oval form in section, extremely 
broad apical termination, striated surface, and im the greater 
thickness and strength of its walls. All together these differences 
appear sufficiently great to render necessary a separate specific de- 
signation ; and I propose to give it the name of A. obtusus, in refer- 
ence to its obtusely-pointed extremity. 

Locality. Cannel Coal, Tingley, Yorkshire. 


3, ANODONTACANTHUS FasTIGIATUS, Davis. (Plate XXIT. fig. 12.) 


Spine imperfect, the base absent; part preserved 2-4 inches in 
length, extending from the point, apparently half the length of the 
spine, the lateral diameter greatest, being 0-4 inch. It is oval in 
section, least diameter between the anterior and posterior faces being 
0-2 inch; there is a central cavity, circular in section, which 
extends towards the point. From the largest diameter of the spine, 
it becomes gradually smaller, and ends in a somewhat worn-looking 
point. The whole of the surface is covered with longitudinal strie, 
most distinct near the basal extremity, anastomosing and becoming 
gradually less numerous towards the point, where the striz have 
disappeared and the spine is smooth, 


OF FOSSIL FISHES FROM THE COAL-MEASURES. 429 


This specimen is evidently closely related to the two species already 
described from the Cannel Coal, though it varies somewhat in form. 
It is larger and stronger in appearance ; as compared with the others 
it is very broad towards the base, and contracts in diameter more 
rapidly to the point. Itis oval in section, in this particular resembling 
A. obtusus, whilst it differs from A. acutus, which is round ; all the 
three agree in being straight, ending in a more or less pointed apex, 
and in the absence of denticles. The spines are of the same dense 
structure in each ; and the form of the internal cavity appears to be 
similar. In reference to its tapering form, I suggest the specific 
name fastigiatus. 

Locality. Blackband Ironstone at Loanhead, in the Middle Lime- 
stone series near Edinburgh. The specimen has been sent to me by 
Mr. W. Tait Kinnear of that city. 


EXPLANATION OF PLATE XXII. figs. 10-12. 


Figs. 10, 10a. Anodontacanthus acutus, Davis. 
Fig. 10d. Transverse section. 
11. Anodontacanthus obtusus, Davis. 
11 a. Transverse section. 
12. Anodontacanthus fastigiatus, Davis. 
12a. Transverse section. 


430 P. B. BRODIE ON CERTAIN QUARTZITE AND SANDSTONE 


34. On certain QuartzitE and SanDstonE FossItiFERous PEBBLEs in 
the Drirr in WARWICKSHIRE, and their probable Ippntity, litho- 
logically and zoologically, with the true Lownr Strur1an PEBBLES 
with similar Fosstts im the Trias at BupierenH SALrerton, 
DevonsuirE. By the Rev. P. B. Broprn, M.A., F.G.S. (Read 
May 11, 1881.) 


Specrat interest and some little difficulty attaches to the history 
and origin of certain quartzite and other pebbles in the Drift in a 
limited portion of the Midland Counties, which it is desirable, if 
possible, to determine. With this object I beg leave to lay a few 
more additional particulars before the Society, which will form a 
supplement to my previous paper in the Quarterly Journal, vol. xxiii. 
p- 210, 1867. It seems probable that the Keuper sandstones and 
marls in the more central parts of England were at one time much 
thicker, before denudation had reduced them, and that the pebble- 
beds existed in them as at Budleigh Salterton, and were afterwards 
broken up and the pebbles (like the flints of the Chalk) scattered 
about in all directions by powerful currents of water, helping to 
form a large proportion of the Drift in this and other parts of the 
Midlands. In a short notice in the Geological Magazine (July 
1878), Mr. 8. G. Percival supposes that the quartzose and many 
other pebbles in the Drift of the Midland Counties were originally 
derived from the Bunter conglomerate. This may be probable in 
some slight degree; and Mr. Jennings, in the Geological Magazine 
(May 1878, No. 167), states that he has found Orthis redux (bud- 
leighensis) in a supposed Bunter pebble near Nottingham, where a 
careful search should be made for other fossils which may occur 
there in situ, both in the Conglomerate and the Drift—viz. the 
igantic Lingule so characteristic of the lower Silurians in Nor- 
mandy, some of which I have already detected in the Drift at 
Rowington. If it can be shown that these pebbles in the Drift were 
originally derived from the Bunter, the latter must have been 
broken up before they were deposited in this newer portion of the 
Trias; then comes the question, Whence were these pebbles derived 
in the first instance, before they were washed, first into the Bunter 
(helping to form the Conglomerate), and afterwards into the later 
New Red Sandstone, as they certainly were, in Devonshire and 
other places? | 
The fossils found in these pebbles here and elsewhere show 
that many belonged to Paleozoic rocks, as evidenced by Orthis 
budleighensis, Lingula Leseuwru, Trachyderma serrata, and some 
others given in the list at the end of this paper. Some of these, 
especially the Lingulw, are species which as yet have not been 
noticed anywhere wm situ in this country, but are peculiar to Nor- 
mandy and Brittany, and were determined by the late Mr. Salter. 
Mr. Pengelly (Geol. Mag. No. 167, 1878) thinks that the occurrence 


FOSSILIFEROUS PEBBLES IN THE DRIFT IN WARWICKSHIRE. 401 


of Orthis budleighensis in the altered quartz rock (Lower Silurian) 
of Gorran Haven, Cornwall, shows that some of the Budleigh pebbles 
were derived from there, which may be the case so far as regards 
the Devonian area, but would hardly account for their presence so 
much further to the north-east, in some parts of the Midland Coun- 
ties for example. 

Therefore I suggested a more northerly or north-easterly exten- 
sion of Old Silurian rocks, in which view my late lamented friend 
Professor Phillips concurred. This would bring them much nearer 
to that portion of England now forming the Midland District; and 
the destruction of those ancient palzozoic deposits may have largely 
helped to supply the Bunter with pebbles, which were in later times, 
by the denudation of the latter, washed into the New Red Sandstone 
then forming. This, again, in its turn, was greatly attenuated, and 
the pebbles, much reduced in bulk, finally distributed, with many 
others derived from rocks of diverse ages and from all parts (north, 
south, east and west, notably from the north), as Drift. 

With reference to the Bunter Conglomerate on the northern edge 
of Cannock Chase, Professor Bonney, in a paper in the Geological 
Magazine for September 1880, concludes, from a careful comparison 
of the quartzite at Loch Nearn in Scotland, that many of the Staf- 
fordshire Bunter pebbles were derived from the north-west of 
Scotland. Other and different quartzites, he says, resemble more 
closely those of Budleigh, the Lickey, and Hartshill. He found 
Orthis budleighensis in a pebble at Rugeley which was identi- 
fied by Mr. Etheridge, and was, he states, lithologically and 
zoologically identical with the Cornish and Budleigh specimens. 
He also noticed a Rhynchonella, and probably Orthis calligramma. 
In Mr. Percival’s collection of pebbles from the Drift at Mose- 
ley, Birmingham, in the Jermyn-Street Museum, the following 
fossils are recorded—Orthoceras?, Cleidophorus amygdalis, Orthis 
budleighensis, Stricklandima lyrata, Spirifera disjuncta, Glypto- 
erinus, Petraia bina*. Professor Bonney is of opinion that the Lickey 
_ largely contributed to the Bunter pebbles about Birmingham and 
Bromsgrove. I quite agree with him in thinking that none of the 
Midland-Counties pebbles came from South Devon, and with Prof. 
Hull that very many have a northern origin; but there many others 
in certain places, and notably in the area referred to in this paper, 


* Some time since, I looked over a miscellaneous collection of rocks presented 
by Messrs Allport and Percival to the Midland Institute, obtained by them from 
the Drift in the neighbourhood of Birmingham, chiefly from Moseley and other 
places adjacent. There are many igneous and metamorphic rocks, basalt, gra- 
nite, syenite, and hard crystalline pebbles, including agates, all of which occur 
at Rowington. The fossiliferous rocks are chiefly Carboniferous, including chert 
with encrinite stems, probably from Derbyshire, several shells and corals and coal- 
plants. There are few (if any) Llandovery species. There are some Orthides, 
some of which occur in a dark-grey boulder very like the Snowdon rocks. 
There are only a few quartz pebbles similar to those described in this paper ; 
but I observed among them Orthis budleighensis and Trachyderma serrata of 
large size and well preserved, but certainly, taking the whole collection, not so 
numerous as in the Drift in this district. 


432 P. B. BRODIE ON CERTAIN QUARTZITE AND SANDSTONE 


which must have had a different origin. Very likely the altered 
Llandovery Sandstone (quartz rock) of the Lickey contributed many 
of the pebbles in this Drift, though it would be almost impos- 
sible to determine the source of all the quartz pebbles in the latter; 
I have, however, not yet detected a single well-defined Llandovery 
species in any of them in this district; and a very considerable num- 
ber, if not a majority of the fossils in the pebbles I have obtained are 
decidedly of Lower Silurian origin ; so that in a direction south of 
Birmingham and towards Warwick and Stratford-on-Avon the 
tendency evidently seems to be that pebbles of lower palseozoic age 
predominate; and the lithological and zoological resemblance which 
they bear to the pebbles of this date at Budleigh is sufficiently close 
to lead to the conclusion that they are in all respects identical and 
were derived from some Old Silurian rocks at a greater or less dis- 
tance, and so far had a common origin. Originally, of course, they 
must have come from their parent paleeozoic rock wherever it was, 
by the wear and tear of which, by the action of waves and currents 
and other processes of denudation, in the course of ages they became 
pebbles, and may have been in the first instance, perhaps, washed 
into the Bunter and have helped to form a portion, greater or less as 
the case might be, of its conglomerates in certain places. When this 
was in its turn fractured and denuded, some of these pebbles may 
have been afterwards redeposited in the New Red Sandstone, not 
everywhere, but locally, as at Budleigh in Devonshire, and parts of 
Warwickshire*. When the latter was also largely denuded, these 
Old Silurian bouldered remnants were finally scattered about and 
mingled with the Drift. If not derived from the Bunter or the New 
Red Sandstone they must have come directly from the relics of some 
ancient Silurian formation which was still in existence, and was 
finally but partially broken up at a more or less distant period, 
thus helping to add largely to the widely accumulating, often sifted 
and re-formed Drift. 

In a redistributed and miscellaneous superficial deposit of pebbles 
and other débris, like that of the Midland Counties in the districts 
referred to, it would, I think, be incorrect (at least we have not 
sufficient evidence as yet) to infer that most of the quartzite and 
other pebbles are derived from the Bunter. A large majority of 
these are more or less consolidated quartzite; and I have one pebble 
of quartz with an obscure impression of a shell, which reminds me 
of the Cornish Lower Silurian rock at Caerhayes. The most abun- 
dant are grey and brown, more or less striated, consolidated sand- 
stone boulders, varying in size and sometimes of a deep red colour; 
and these agree lithologically with the Budleigh pebbles containing 


* Although, with the exception of the Lower Keuper sandstone at Budleigh, 
there are no pebbles elsewhere found in situ in the New Red Sandstone, there 
is no reason why they should not have occurred in certain places in other di- 
rections in the same formation, either in the Upper or Lower Keuper. In the 
Midland Counties the upper division has here and there undergone very ex- 
tensive denudation ; or possibly pebble-beds may still exist in the New Red which 
have not yet been discovered. 


FOSSILIFEROUS PEBBLES IN THE DRIFT IN WARWICKSHIRE. 433 


certain well-known Silurian species and some of the rock specimens 
I have from Normandy. In fact placing the majority of the War- 
wickshire Drift pebbles in my collection side by side with those from 
Devonshire and France, it would be impossible to separate them, 
the character of the rock being identical, and the fossils, where pre- 
sent, the same. This is a point of much interest, and while fossili- 
ferous pebbles are on the whole rare (though, of course, very many 
must have been overlooked), there are many which have the same 
mineralogical character, and really form a considerable portion of 
the gravels, and were, no doubt, derived from the same source and 
originally belonged to a formation of the same age. With the 
exception of a few Carboniferous fossils and others of later date a 
very large percentage belong, as it appears, to the older Silurian 
rocks, and wherever they may have occurred a situ must have been 
originally derived from them. The absence, apparently, of Llan- 
dovery fossils is remarkable, and seems to show that a very small 
proportion of the Drift in this district had come from the Lickey 
within sight and not so very far off. In a list of fossils from 
the Bunter conglomerates near Cannock Chase, Mr. W. Molyneux*, 
F.G.S., on the authority of the late Mr. Salter, assigns all the species to 
Mountain- Limestone and Upper Silurian(May Hill sandstone, Llando- 
very) groups, and not one Lower Silurian form occurs; but Professor 
Bonney records Orthis redux (budleighensis) + from the same district 
on the authority of Mr. Etheridge. Mr. Davidson¢ is of opinion that 
the majority of the Brachiopoda in the pebbles at Budleigh are 
Devonian, which predominate; but the Lower Silurian are sufficiently 
numerous and well preserved to have enabled Mr. Salter || to deter- 
mine and identify a considerable number with certain species, giant 
Lingule and others, peculiar to the district of May and Gehard, 
(Armorican sandstone) in Normandy. 

The small collection of Lower-Silurian fossils which I possess, the 
result of several years’ work, from a limited area of the Midland 
District, must have been derived from a much nearer source than 
Devon or Cornwall; and Professor Bonney, in the same paper, justly 
observes that, as from physical considerations it is almost impossible 
that Cornish pebbles could have made their way into the country 
round Cannock Chase, the only possible inference is a nearer one by 
a further extension to the north-east of the old Silurian strata with 
their fossiliferous sandstones and quartzites. The idea that they 
can have been derived directly from Normandy is, of course, out of 
the question ; and Mr. Davidson also contends for an extension, as I 
do, of Silurian rocks in the Channel and nearer to Devonshire as a 


* Proceedings of the Dudley and Midland Geological and Scientific Society, 
1877, p. 139.. 

t Ate A. H. Atkins, B.Sc., one of the masters of King Edward’s School, Bir- 
mingham, has lately found Orthis budleighensis, in situ, in one of the Bunter 
pebble-beds, at Kinver Edge, near Stourbridge, which is another instance of 
the presence of this species in the lower division of the Trias. I recognized 
the species at once, and Mr. Davidson has since confirmed it. 

¢{ Quart. Journ. Geol. Soe. vol. vi., February, 1870, No. 101. 

|| Quart. Journ. Geol. Soe. vol. i1., August, 1864, No. 79. 


434 P. B. BRODIE ON CERTAIN QUARTZITE AND SANDSTONE 


probable source of the Budleigh pebbles which contain fossils of 
that date. The extension of this area of ancient rocks, or, indeed, of 
any geological age, is never improbable when we consider the 
enormous amount of denudation which all great formations have at 
one time or other undergone. I have made some additions to 
the Lower-Silurian fossils I have detected in the Warwickshire Drift 
since the publication of my former paper on this subject in the 
Journal of the Geological Society (vol. xxiii. p. 210, 1867); Orthis 
redux and Lingula Lesewrt were determined by Mr. Woodward some 
years ago ; Lingula, n. sp., by Mr. Davidson, and the remainder by 
Mr. Etheridge, who has been kind enough to examine them; and I 
here give the entire number, with the additional species, on his 
authority : 


Orthis budleighensis. Lyrodesma celata? 
Valpyana. Ctenodonta Bertrandi ? 
Lingula Leseurii, n. sp., Davidson. Arca Noranjoana ? 
Spirifer antiquissimus ? Palearca secunda. 
Davidis? Trachyderma serrata, 
Rhynchonella sp. Calymene Tristani. 
Modiolopsis lirata. Homalonotus, portions. 
sp. Fucoids, one or two branching forms. 


Orthis budleighensis is very abundant and characteristic. One 
of the Lingule determined by Mr. Davidson is, he says, a new 
species and totally unlike any of those from Budleigh. The 
Annelid Trachyderma serrata is frequent and generally in good 
preservation. The total number of Lower-Silurian genera and species 
figured and described by Salter from the pebbles at Budleigh Sal- 
terton is twenty-four; and there are several diligent collectors of - 
these fossils in Devonshire. From the Midland Drift, in a limited 
space and with less facilities for obtaining them, I have procured 
sixteen, leaving only a difference of eight in favour of Devon—which 
is somewhat remarkable, considering the much better opportunities 
for collecting them in the West. At present, I believe, this is the 
largest number of Lower-Silurian fossils of the age, probably, of the 
Bala and Lower Llandeilo formations, hitherto observed in the Mid- 
land Drift. This fact too, I think, strengthens my argument in 
favour of more extensive ramifications of old paleozoic rocks in a 
north or north-easterly direction; and these, when broken up, would 
furnish ample materials coming from different areas to supply the 
fossiliferous pebbles referred to. No doubt this list would be 
largely increased if the road-heaps, collected chiefly from the fields, 
could be more diligently searched; for many fossiliferous pebbles 
must be overlooked. 

Mr. Walter Keeping, in an interesting paper * on the Upware and 
Potton Greensand pebble-beds, observes that “some of the quart- 
zites are like those of the New Red Sandstone pebble-beds, and were 
probably thus derived.” He accounts for the presence of other older 
paleeozoic pebbles by the supposition that they were derived from a 
great paleozoic ridge extending northwards towards Cambridge. 


* Geological Magazine, No. 195 (New Series, No. 19), Sept. 1880, p. 414. 


FOSSILIFEROUS PEBBLES IN THE DRIFT IN WARWICKSHIRE. 485 


From this source at an earlier period the Upper and Lower Triassic 
pebbles were, as I have inferred, probably derived; and hence it 
formed the supply of a large proportion of the Midland Drift in the 
districts referred to in this paper. Mr. Etheridge suggested the 
possibility of the Wenlock Limestone, lately noticed in a boring at 
Ware, dipping to the south. This surmise, Mr. Keeping says, has 
since proved to be correct; and therefore a further extension of 
ancient Silurian and Cambrian rocks northwards is still more 
probable. 

This denudation of these old rocks may have come to a close 
during the Cretaceous or even some later period, when they finally 
disappeared beneath the sea, having been going on for ages, perhaps 
commencing at a date somewhat anterior to the deposition of the 
Bunter. There are, perhaps, few geological problems more difficult, 
to solve than the history (range, distribution, and origin) of these 
latest Pleistocene deposits, such as clays, till, gravel, and sands under 
the name of Drift, which are so widely spread and often composed of 
such heterogeneous materials. Where made up for the most part of 
formations present in situ in the immediate neighbourhood or not 
very far off, the solution is comparatively easy; but where they 
are mainly derived from a great distance, perhaps here and there 
from foreign sources, as some of our Midland Drift may be, it is very 
puzzling and extremely difficult to determine whence they originally 
came and by what means they were transported. Though many 
able papers have been written upon this subject, much yet remains 
to be done both here and elsewhere before definite and satisfactory 
results can be obtained. 


[ Note, July 21.—It has since been determined that some of the 
Brachiopoda belong to the Caradoc. | 


Discusston. 


The Presipenr remarked that the subject was a difficult one, 
but the species, as stated by Mr. Brodie, were no doubt correct. 
At Budleigh Salterton it was easy to tell whence many of the 
pebbles had been derived ; but in the Midland counties it was most 
difficult. 

Mr. Ussuer said that the occurrence of the same fossils did not 
prove that the pebbles were from the same area, and that the Drift 
of the paper was not quite clear from its title. 

Prof. Bonney stated that the author had not expressed this 
opinion in his paper; he thought it a most valuable addition to our 
knowledge, and was glad that such a contribution had been evoked 
by his own slight paper. He thought it almost certain that there 
were two sources for the quartzites, and that all the fossiliferous 
specimens could not have come from the Lickey, as some had been 
found at Nottingham. Probably ancient rocks had extended to the 
north-east of England, beyond those discovered in borings at 
Northampton, and to the north-east of those exposed at Charnwood. 


436 T. M, READE ON THE DATE OF THE LAST 


30. The Date of the tast Coance of Leven wm Lancasuire. By 
T. Mettarp Reapz, Esq., C.H., F.G.S., F.R.I.B.A. (Read 


April 6, 1881.) 


In estimating geological time the difficulty always lies in getting a 
reliable unit to measure with. Having surveyed and mapped out, 
on a scale of 6 inches to the mile, the whole series of Postglacial 
deposits between Liverpool and the mouth of the river Douglas, I 
have often asked myself, Can the age of any of these deposits be 
translated from mere sequence into years? Observation has led me 
to believe that an approximation may be made in the case of the 
blown sand; and this, as I will presently show, bears upon the title 


of my paper. 
A reference to the map (1 inch to the mile) and section, figs. 1 & 2, 


Fig. 1.—Sketch Map of the Coast at Blundellsands, near Liverpool. 
(Scale 1 inch to the mile.) : 


SUBMARINE 
FOREST ~ 


il 


ALLTEL ir Nw 


il 


a oF 
eee) 
- 


I 


Peat and forest- 
bed. 


MEASURED 
ACCUMULATION 


Y) OF BLOWN_SAN ' ts 


Boulder-clay. 


CHANGE OF LEVEL IN LANCASHIRE. 437 


and to those accompanying my paper on the Post-Glacial Geology 
of Lancashire and Cheshire, in the ‘ Proceedings of the Liverpool 
Geological Society,’ 1871-2, will explain the order and sequence of 
the deposits ; and it will be seen that the last movement of the land in 
Lancashire was downwards*. Submarine forests at the Alt mouth, 
the Rimrose brook, the Liverpool and Garston docks show that sub- 
sidence has taken place ; and (which is perhaps quite as good evidence) 
sections of stream gullies, cut in the Boulder-clay and filled with 
recent silt, at levels far below high water, are frequently met with 
in dock-excavations. 

Upon the superior peat- and forest-bed, which is an extension 
inland of the submarine forests, rests, as on a platform, some 22 
square miles of blown sand, in some cases rising 75 feet above 
Ordnance datum, and estimated by me to be at least 12 feet in 
average depth. 

This deposit is shown in my map, as well as in that prepared by 
the Geological Survey, by yellow dots; it varies in width at dit- 
ferent localities, its maximum being at Formby, where it reaches 
3 miles inland. It is quite evident that the whole of this Aolian 
deposit is an accumulation subsequent to the last subsidence of the 
land ; therefore, if we can calculate the time it must have taken for 
the deposit to form, we shall be in a position to determine the least 
time that can have elapsed since the subsidence. 

The whole of the blown sand has been derived from the shore 
between high- and low-water marks, but principally from between 
high water of springs and neaps when the shore is dry. On a windy 
day it is very curious to see the streaks of sand rushing over the 
shore, even when it is damp, shining like rays of a lighter colour 
pencilled over a dark ground. The shore is very flat, being in 
places more than a mile wide between high- and low-water marks 
of spring tides; so that the conditions for the generation of subaerial 
sand can hardly ever have been more favourable than they are now. 

In May 1866 I set out a plot of land at Blundellsands, in Burbo 
Bank Road North, for building-purposes ; it had a frontage of 350 
lineal yards to the sea, the western boundary being the then high- 
water mark of spring tides. In 1874, for the purpose of enclosing 
the said plot, I had to remeasure it to define the boundary, when I 
found that high-water mark was considerably beyond the western 
boundary, and that the sand had gained upon the sea. An open 
wire fence was then put up on the original high-water mark, when 
measurement showed that there were 15 yards of land in front of it 
at one end, and 5 yards at the other. The high-water mark of 
springs had, in fact, receded to that extent. I estimated the deposit 
of sand that had taken place in the eight intervening years at an 
average of 10 yards wide along the whole frontage and 2 yards 
deep. Allowing 1 yard more in depth for sand that may have been 


* At Meols, in Cheshire, on the opposite side of the Mersey estuary, was a 
Roman station ; and the land is now only a few feet above high-water ; therefore 
the land cannot have rzsen since the Roman occupation. 


Q.J.G.8. No. 147. 26 


438 T, M. READE ON THE DATE OF THE LAST 


blown over the top, which I am 
convinced is a large estimate, as 
very little sand blows on to Burbo 
Bank Road North, the eastern 
boundary of the plot, we shall thus 
have 350 x10x3=10500 cubic 
yards of sand deposited in eight 
years on a shore-frontage of 350 
lineal yards—or 3°75 cubic yards 
per lineal yard of frontage per 
annum. 

Taking this as my unit of mea- 
sure (and it is an exceptionally 
large one), I find, for the 16 miles 
of coast forming the western 
boundary of the deposit, it will 
give 105,600 cubic yards per an- 
num as moved by the wind; and 
dividing the 272,588,800 cubic 
yards contained in the 22 square 
miles, 12 feet thick, it will give 
2580 years as the age of the whole 
deposit of blown sand if accumu- 
lated at the assumed rate. 

Since these calculations were 
made, I have lately tested their 
accuracy in another manner. 
There is another plot in Serpen- 
tine Road, having a sea-frontage 
of 243 lineal yards, and contain- 
ing about 6500 square yards. 
Serpentine Road was made from 
my sections and under my super- 
intendence 1n 1866; so that I had 
every opportunity of ascertaining 
the respective levels. Taking 
every thing into consideration, I 
estimate that there has accumu- 
lated over its whole surface an 
average depth of less than 2 yards 
of sand (very little blows across 
the road); so I think we may 
fairly take an average depth of 2 
yards as representing the quantity 
of sand blown off the shore from 
a frontage of 243 lineal yards. 
This, extended over fourteen years, 
gives 3°81 cubic yards of sand per 
lineal yard of frontage per annum. 

It is quite evident that the sand 
cannot accumulate faster than it 
sweeps off the shore; but at other 
points on the coast the sea is 


(‘depq 00g) ‘paq-3se10y7 outavuqns v se a1oys oy} UO sdveddeoa pue souNp-puKY oY} LopuN senuUIZUOO SIq} YJNOM ITY 04} 4V x 


%* 


"kelQ Jopfnog, 


Highwatermark. 


Measured aceu- 
mulation of 
blown sand. 


Dunes. 


- Dunes. 


The Warren 
covered mostly 
with dwarf wil- 
lows. 


Railway. 


TL Sla Me gy amy ay7 buopn woyvg—s “Siz 


Links of the W. | 
Lancashire Golf 
Club. 


Marsh Lane. 
Dunes. 


CHANGE OF LEVEL IN LANCASHIRE, 439 


gaining on the land ; therefore what is deposited in front of the sand- 
dunes is again swept away, and ali that goes to the accumulation is 
that which is blown over the tops of the hills or sand-cliffs. 

This will also be the case where the coast-line is stationary. — 
Again, the high sand-dunes form a barrier that the sand cannot so 
readily surmount, the practical effect being that, the bases being 
swept by the tide, and the angle of repose remaining constant, less 
sand travels inland the higher they grow. At the mouth of the 
Alt there is a very extensive bank at low water swept by the north- 
west wind ; but the sand does not accumulate, as it is blown into the 
river Alt and washed out seawards again. The land on which the 
Altear rifle-range is situated has, in fact, as a protection to the river, 
been gained from the sea by the erection of artificial obstructions, 
against which the sand accumulates. This accumulation has now 
practically ceased; and the river and the sea combined are eating 
into the land by the Crosby lighthouse, and making sad havoc with 
the submarine peat- and forest-bed. : 

On referring to the map it will be seen that the blown sand is 
narrowest at this point, though the conditions are very favourable 
for its development, had the river Alt not intervened. 

If, then, it be conceded that the last change of level in South- 
west Lancashire was a downward one, I think the facts and cal- 
culations I have had the honour to lay before you pretty clearly 
prove that it did not take place within the last 2500 years. 


Discussion. 


The CHarrman (Mr. Hulke) remarked upon the economic as well 
as scientific interest of the communication from the proved increase 
of land in the area. 

Mr, De Rance, who had surveyed the district described by Mr. 
Reade in his paper, could corroborate many of the author’s conclu- 
sions, especially by the finding of Roman coins on the surface of 
the ar land and by a Roman bath only 5 feet above high-water 
mark. 

Prof. Jupp stated that Mr. Reade’s conclusions were entirely in 
accord with the most recent researches concerning the supposed 
changes in the level of the shores of the Firth of Forth since Roman 
times. 

The AvurHor stated that at Hoylake, in Cheshire, numerous 
Roman remains belonging to a Roman encampment a few feet 
above high-water mark prove that no appreciable downward move- 
ment has taken place since Roman times. He thought the actual 
period required for the formation of the blown sand was probably 
nearer 5000 than 2500 years. 


440 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


36. On a new Sprcizus of Piestosaurus (P. Conybearr) from the 
Lowsr Lias of Cuarmoutn; with Observations on P. MEGA- 
CEPHALUS, Stutchbury, and P. BRacHYcEPHALUS, Owen. By W. 
J. Sorzas, M.A., F.R.S.E., F.G.8., &c., Professor of Geology in 
University College, Bristol. Accompanied by a SuepLemEnt on 
the GrocRAPHICAL Distrisution of the Genus PLEstosauRrus, by 
G. F. Wuipzorne, Esq., M.A., F.G.S. (Read May 11, 1881.) 


[Puates XXIII. & XXIV.] 


Tae nearly complete and very fine specimen of Plesiosaurus 
(Pl. XXIII. fig. 1) which forms the subject of the present paper is 
the latest addition to the already large collection of fossil reptiles 
preserved in the Bristol Museum. 

It was found over a year ago, by Samuel Clarke of Charmouth, on 
the north-west corner of Blackven Water, half a mile west of the 
river Char, where it lay in a ‘“‘table-ledge” of the Lower Lias, some 
seven feet above the “ boulder-bed.” From its position and the 
species ** of Ammonite still associated with it, we may conclude 
that its geological horizon is that of the zone of Ammonites obtusus. 

From the pectoral to the pelvic girdle it is imbedded in a layer of 
hard impure limestone, thick enough to hold the parts securely 
together, but at the same time thin enough to let the skeleton be 
seen on both sides, dorsal as well as ventral. The head and neck 
were preserved chiefly in shale; so that, to ensure the safety of the 
neck, it has been found necessary to imbed it in plaster; but the 
head, being filled in and about with limestone, has been left free, 
and can be turned about, handled, and examined on all sides. 

The ventral surface of the fossil (Pl. XXIII. fig. 1) 1s exposed on 
the upper surface of the imbedding limestone; the coracoids lie side 
by side nearly in the position they would have occupied in the dead 
animal lying on its back, except that they are slightly displaced to- 
wards the left side; the left pubis and ischium are also nearly in 
position ; but the corresponding bones of the right side have been 
pushed to the left, so as to underlie them; the femora, which are all 
that is left of the hind limbs, remain on their proper sides, extended 
outwards and backwards. The fore limbs have been considerably dis- 
placed ; for though that of the left remains on its own side, the palmar 
surface of its hand looking upwards, that of the right has been com- 
pletely crossed over onto the left, so that its palmar surface would be 
directly superposed on that of its fellow were it not for a slight dis- 
location at the distal end of the humerus, which has carried the rest 
of the right limb backwards, and so left the left hand exposed. The 
scapule have turned on their axes, but have not shifted sides; and 


* This, according to the determination of Mr. Whidborne, is Ammonites 
planicosta, 


\ 
r 2 
{ 
af 
. 
; 
t 
S 
+ 
ify 
é a ‘ 

+ 7 . 


Quart Journ. Geol. Soc. Vol XXXVILPLXXI. 


C Beryean del ot Lith 


Minter Broa imp 
PLESIOSAURUS CONYBEART. 


Mmtern Bros amp . 


CONYBEARL AND BRACHYCEPHALUS 


So 


SUEY 


Quart. Journ. Geol. Soe Vol. XXXVIT.Pl XXIV. 


PLESIOSAURUS 


(g .Berjeau. del et hth. 


FROM THE LOWER LIAS OF CHARMOUTH. 441 


the “furculum,” or combined clavicles, is tilted up from right 
to left. 

So far as the appendicular skeleton is concerned, it has approxi- 
mately the position it would have in an animal lying flat on its back 
with its limbs extended outwards, except that it has been skewed 
over as a whole a little to the left, and the right fore limb completely 
crossed over. 

The axial skeleton is similarly disposed as far as the ribs are con- 
cerned ; but the head and vertebral column have been turned round 
90°, and lie on one side, the left. The neck and head are also 
curved backwards, in the manner so usual with Plesiosaurus, and 
which has been commented on by Professor Huxley as suggestive of 
death by opisthotonic contraction. 

This disposition of the parts of the skeleton may be readily ex- 
plained by supposing that the animal to which it belonged fell, after 
death, sideways through some depth of sea-water to an oozy bottom. 
The body, being broadest laterally, has settled on its back, the hind limbs 
sprawling outwards. The left shoulder touched the bottom first ; and 
the right fore limb heeling over, fell across the one to the left. The 
neck and, in this species, the head being broadest dorso-ventrally, 
settled on one side, the left, and so communicated a twisting strain 
to the rest of the vertebral column, which, being but slightly attached 
to the ribs and appendicular skeleton, readily yielded to it, and 
turned on its side also; thus the whole of the vertebral column 
came to lie on its broadest face (that is to say, laterally). The twist 
to the left given by the crossing of the right fore limb, and the sub- 
sequent pressure of overlying strata, led to the various other minor 
dislocations and displacements. 

That the specimen is undoubtedly the type of a new species is 
shown by the following summary of its chief characters :— 

1. The length of the skull from the anterior extremity of the 
lower jaw to the posterior margin of its articulation with the qua- 
drate bone is 19-75 inches, measured along the right side. 

2. The number of vertebra is 66, of which 59 are cervico-dorsal, 
2 sacral, and 5 caudal. Of the cervico-dorsal vertebrae 38 appear 
to be cervical and 21 dorsal. 

3. The length of the cervical region is 83 inches (6 feet 11 inches); 
so that the length of the head is to that of the neck as 24-1: 100. 

4, The length of the cervico-dorsal series is 136 inches (11 feet 
_ 4 inches); and the length of the head to this is as 14°6: 100. 

5. The length of the centrum of the anterior cervical vertebre is 
equal to the height and greater than the breadth of the articular 
face. Thus in vertebra xv the measurements are :—Length 2 inches, 
breadth 1-5 inch, height 2 inches. 

6. In the posterior cervical vertebre the breadth of the articular 
face is greater than the length or height, but the latter two dimen- 
sions remain equal, Thus in vertebra xxxv we find—length 2:7 
inches, breadth 3°5 inches, height 2-7 inches. | 

7. The neural spines increase in size up to the 40th to 44th ver- 
tebra, in which they measure 4-75 inches in length. 


442 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


8. The neural spines are inclined backwards as far as the 55th 
vertebra; past this up to the 57th they are inclined somewhat for- 
wards; but after this they again incline backwards to the end of the 
vertebral series. 

9. The humerus and femur are nearly equal in length, the femur 
being slightly the shorter. 

For the new species which these characters indicate, I propose 
the name of Plesrosaurus Conybeart, as a tribute to that classic au- 
thority who first made the existence and nature of Plesiosawrus 
known to ns. ‘This was sixty years ago; and it is singular that up 
to this date no one seems to have thought of calling some species of 
Plesiosaurus after the author of the genus. 

Plesiosaurus Conybeari agrees closely with P. Htheridgit in the 
relative length of head and neck; but if the length of the head be 
compared in each with that of the whole cervico-dorsal series, a 
marked difference is apparent; thus in P. Htheridgu the ratio is 
12°5 : 100, in P. Conybeart 14:6: 100. P. Conybeari further differs 
from P. Etheridgu in absolute size, being nearly twice as long; it 
also possesses a larger number of cervico-dorsal vertebra, P. Hthe- 
ridgw having only 53, or 6 less than P. Conybeari. In this latter 
character the new species more nearly agrees with P. homalospon- 
dylus, each having 388 cervicals, while the latter has 22 and the 
former 21 dorsal vertebree; but P. Conybeart has a larger head ~ 
than P. homalospondylus. Both haye the same length of neck; but 
P. homalospondylus has a longer dorsal series of vertebra (81 inches 
in length). The length of its head relatively to that of the neck and 
cervico-dorsal series is also much: less than in P. Conybeari, being 
to the neck as 10°6: 100, and to the cervico-dorsal series as 
04: 100. 


Detailed Description. 
The Skull (Pl. XXIV. fig. 1).—This, which has been very thoroughly 


freed from matrix, and is in an excellent state of preservation, pre- 
sents us with that very exceptional character amongst Plesiosaurs, a 
good profile. This is due to its having been compressed from side to 
side, and not, as is more usual, depressed from above downwards. 
Perhaps this indicates a difference in the original shape of the head. 
The right side of the skull has slipped a little upwards above the left; 
and some other displacements have occurred, but nothing like so great 
as one would have expected if the present greatly compressed head 
had originally been as broad from side to side as most Plesiosaur 
heads evidently were. 

Posterior Aspect.—In the middle of the back of the skull is a con- 
fused mass of bone comprising the axis and atlas vertebra, under 
which the foramen magnum lies concealed. Inferior to this are the 
posterior ends (articulare) of the rami of the lower jaws, bent to- 
wards the middle line; that on the right side is nearly perfect, 
clearly not needing more than a quarter of an inch to complete it. 
The articulare thins off rapidly behind its articulation with the qua- 


FROM THE LOWER LIAS OF CHARMOUTH. 443 


drate, not extending more than 1°5 inch beyond the posterior edge 
of the condyle. 

The quadrate appears to be directly continuous with the squa- 
mosal on each side ; and the latter bones are prolonged upwards into 
a process which, viewed from behind, has a somewhat sabre-like out- 
line, with the convexity outwards. Both these suprasquamosal 
processes are broken at the distal end, so that their junction over the 
parietal (a characteristic Chameleon feature seen in most Plesio- 
saurs) is not here observable. 

Superior Aspect.—Most posterior is the previously mentioned axis 
and atlas vertebral mass; then succeed in front the approximated 
ends of the broken suprasquamosal processes. The parietal comes 
next, a pent-roof-like bone, 5°75 inches long, compressed for the 
anterior three quarters of its length into a strong median crest, 
along which the persistent sagittal suture extends, expanding ante- 
riorly into the foramen parietale 0-3 inch wide, in front of which the 
parietal ends. Behind, the parietal is expanded into a form lke 
the bowl of a spoon, the bowl being supposed turned with the con- 
vexity upwards. 

The next bones in front are the frontals, longitudinally ridged in 
the middle, smooth at the sides, and separated by a median suture, 
which is slightly more open in the middle of its length than at the 
ends*. A splintery suture joins the frontals to the nasals, which 
are short longitudinally striated bones, not by any means clearly 
defined in front from the posterior prolongations of the preemaxille. 
The preemaxille, 9 or 10 inches long, are separated for their whole 
length by a simple suture; posteriorly they are smooth, or only 
faintly wrinkled, but in front much roughened, probably for attach- 
ment of integument. 

From the parietals a process is given off on each side of the fora- 
men parvetale, and continues backwards as far as the middle of the 
lateral margin of the bone; in uncrushed skulls this process is a 
plate of bone standing out nearly at right angles to the body of the 
parietal; its parallelism in this case must be due to compression. 

Lateral Aspect.—On the left side the upper jaw is 14°3 inches 
long; and the anterior 4°3 inches is furnished by the premaxilla ; 
this bone joins the maxilla along a line which runs obliquely 
upwards and backwards, to end just above the anterior nares. 
The maxilla is an irregularly triangular bone. Its base (10” long) 
furnishes the margin of the upper jaw; its anterior side bounds the 
premaxilla; its posterior side, just behind the apex, furnishes the 
lower anterior boundary to the external nostril, and further down 
the lower anterior margin of the orbit (being excluded from the upper 
part of it by the lacrymal and prefrontal bones, here badly defined) ; 
still further downwards and backwards it meets the jugal, by 
which it is excluded from the posterior half of the lower bonndary 
of the orbit; and along and beneath this bone it extends to its 

* This appears to indicate a fossa corresponding to that on the frontal suture 


of the Lizard’s skull, interpreted by Professor Parker as “the scarcely-closed 
anterior fontanelle’ of Clarias.”—Phil, Trans. clxx., 1879, p. 598. 


444 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


termination, which takes place a considerable distance (over an inch) 
behind the orbit. This is a marked character in several of the La- 
certilia, particularly the Agamide. The maxilla bears teeth at 
least up to within 1-8 inch of its termination. 

The jugal is bounded below by the maxilla; in front it forms the 
posterior lower corner of the orbit; above it joins the postorbital 
(postfrontal), which bounds the upper posterior corner of the orbit ; 
and behind it unites with the squamosal by a splintery suture, 
which is 1-4 inch long, and runs almost at right angles to the length 
of each bone. The jugal is convex outwards in front, and depressed 
behind; in the depression a vertical row of three oval pits (nutritive 
foramina) separated by intervening smooth ridges is situated. At 
the ends of the pits, which are elongated antero-posteriorly, strize 
appear on the surface of the bone, and are continued forwards, 
diverging at the same time, over the otherwise smooth anterior con- 
vexity. The external form and surface-markings thus described 
give to the jugal such a characteristic appearance that it is easily 
identified by them alone. 

The squamosal is a large and important bone, of which the 
general form and relations are not quite so clearly defined as could 
be wished. Its characteristic anterior or zygomatic process, how- 
ever, is well displayed; it is a thin bar of bone 1 inch broad, about 
23 inches long, and + inch thick, finely striated longitudinally, the 
strize sweeping somewhat obliquely forwards from above down- 
wards. 

The postorbital continues backwards from the jugal over the 
upper edge of the zygoma for a distance of 0°8 inch. These three 
bones, Jugal, postorbital, and squamosal, clearly meet in a T-shaped 
suture; and thus Professor Huxley is undoubtedly correct in assert- 
ing that, “‘ contrary to what is usually stated, the postfrontal appears 

. to articulate with a bone, the homologue of the squamosal 
of the Crocodile” (Quart. Journ. Geol. Soc. vol. xiv. p. 293). 

A slight extension downwards of the lower margin of the anterior 
end of the squamosal bar, rendering its inferior outline curved con- 
cavely, while that above is rectilinear, brings it within half an inch 
of the posterior end of the maxilla. From that part of the jugal 
which is exposed between the squamosal and the maxilla no bony 
bar is produced towards the quadrate; nor is there the slightest 
evidence of one having ever existed, so far as this skull is concerned. 
I feel persuaded that an inferior bony temporal arcade has never 
been present, and that, if a quadrato-jugal occurs at all, it must be 
as what we have called the anterior process of the squamosal, while 
for such a view I see no evidence. 

The quadrate is clearly enough identified at its articulation with 
the lower jaw; but on tracing it towards the squamosal it is impos- 
sible to say where it ends or the squamosal begins. The relations of 
the two bones are quite obscure; so that one cannot even tell in this 
specimen whether what we have already termed the suprasquamosal 
processes are the property of the bones to which we have assigned 
them, or whether they came off from the quadrate. They appear in 


FROM THE LOWER LIAS OF CHARMOUTH. 445 


other skulls, however, as continuations of the squamosals, though 
apparently divided from them by suture. Owen calls them “ supra- 
mastoids ;” and they may correspond to Parker’s ‘‘ second supra- 
temporals,” though, as they lie exterior to the squamosals and not 
beneath them, our term ‘“‘ suprasquamosal” is perhaps best retained. 

The articular end of the quadrate lies much below the general 
level of the upper jaw (over an inch). This is a character not found 
in Agamide or most Lacertilia; but it occurs in Iguana and the 
Chameleons. 

The Lower Jaw.—This, which has the usual reptilian composition, 
is 20°35 inches long from end to end; posterior to the symphysis its 
surface is smoothly striated by longitudinal thread-like ridges, the 
external expression of its fibrous structure ; in front, past the pos- 
terior end of the symphysis the surface is much roughened, and 
along the alveolar margin finely wrinkled*. The length of the 
symphysis is 3°35 inches; the height of the jaw from the top of the 
coronary to the lower margin of the ramus is 3 inches. 

In the following Table are given the more important measure- 
ments of the skull :— 


inches. 
Anterior extremity of preemaxilla to posterior margin of parietal......... 18:0 
Bs yp anterior mar gin OO,  Gocenooucoos 10:0 
- posterior margin of orbit ............ 13-0 
Ks a Bi left nostril ...... 8:2 
Le s s right nostril 9:2 
a Ys foramen’ parietale .-2%.25..2.0+.----0¢ 13-0 
Worn nereht of orbit ie SEGRE ase tO ota Sota cars dale ameetacucaeecagesaes 3:2 
8 TaTELINS: “nerd SOaecaeheis eae Re Se eer On 21 
Highest point of parietal to lower margin of lower jaw i ea eee ce 
Anterior end of dentary to posterior margin of articulation with quadrate 19°75 
3 x posterior end! of articulare) <..2:..<0.02..5:..c<00s 21:0 


Section across the Skull—The skull has been broken across in 
several places, so as to afford a view of its internal structure; but 
very little is clearly displayed, except in the most posterior fracture, 
which traverses the supratemporal fosse (fig. 1, p. 446). 

A large and originally bilaterally symmetrical mass of bone is 
seen in the middle of the section above the lower jaw; the greater 
part of it consists probably of the basisphenoid ; it is channelled in 
the middle line above by a deep narrow groove, through which a 
line can be drawn to the middle point of the inferior concave margin 
opposite, dividing it into nearly symmetrical halves; the right moiety, 
having apparently suffered but slightly from compression, is better 
fitted for study than the left, which is much crushed together. The 
right half is divided by a deep lateral excavation into an upper and a 
lower portion : the latter, descending outwards and downwards, soon 
bifurcates; and the upper of the two processes so produced, after 
diminishing to a narrow neck, widens suddenly into a triangular 


* The wrinkled appearance here referred to is very similar to that exhibited 
by parts of the integument described on page 466. 


446 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


area, the transverse section of a bone which is continued backwards 
on the side of the skull to terminate on the inner side of the lower 
end of the quadrate. In all probability this bone is the pterygoid, or 


Fig. 1.—Transverse Section through the Skull in the Region of the 
Supratemporal Fosse, (Scale 5.) 


at all events a part of it; and the narrow neck from which it extends 
may be partially furnished by the basipterygoid process ; but there is 
no evidence of the existence of a joint between it and the pterygoid. 
The lower process of the bifurcation may also be a part of the ptery- 
goid; if not, I do not know what nature to suggest for it. 

On the left side the pterygoid is crushed up against the basisphe- 
noid, almost obliterating the space which separates them on the 
right; and the descending process below it is thrust and broken 
against the lower jaw. 


FROM THE LOWER LIAS OF CHARMOUTH. 447 


On each side of the section above the basisphenoid the thin bar 
of bone which proceeds from the squamosal is seen, and at the 
summit the parietals with their median crest and persistent suture. 
The other bones appear to be undeterminable ; some of them are 
probably parts of the periotic mass. 

Dentition.—A fine series of teeth is well displayed on the left side 
of the head. They are slender, conical, slightly recurved, and finely 
striated from the apex for a considerable distance downwards, 2. ¢. 
over the crown. They vary greatly in size, the largest being those 
in the neighbourhood of the maxillo-premaxillary suture; an inch 
or so behind this they begin to diminish in size, and beneath the 
orbit have less than one half the average length of those in front, 
while behind it they dwindle to mere pointed tubercles. 

The largest tooth present is one in the right upper jaw, behind 
the maxillo-premaxillary suture; it is 2°45 inches long, the distal 
1°5 inch, or crown, finely striated, its diameter at the point where 
the striz begin being 0°6 inch. A smaller but more perfect tooth, 
the largest on the left side, measures 1-95 inches long; 1°35 inch is 
striated ; and the diameter at the commencement of striation is 
05 inch. 

Those teeth which still remain exposed to view are distributed as 
- follows :—on the left side in the premaxilla 5, in the maxilla 15, 
in the lower jaw 13; on the right side in the premaxilla 3, in the 
maxilla 11, in the lower jaw 8. The number of teeth in the left 
premaxilla and maxilla make the nearest approximation to the 
numbers originally present. 

The Vertebral Column.—tThere is a continuous series of 66 ver- 
tebre, of which 38 are cervical, 21 dorsal, 2 sacral, and 5 caudal. 
The caudal series is evidently incomplete, a considerable number of 
vertebre being missing from the distal end. 

Cervical Vertebre.—The first and second, as already mentioned, 
form a confused mass adherent to the back of the skull, but from 
the third onwards all are clearly defined and can be easily examined. 
m1. The centrum of the third is 1:12 inch long* (a.p.), 0°75 broad 
(1.1.), and 1:4 high (d.v.), the breadth and height being measured 
along the articular face. It is much compressed in the eaiddle, the 
edges of the articular ends projecting greatly, as though the more 
yielding cancellous interior had given way under heavy pressure, 
such as that of overlying strata; this feature is markedly present as 
far as the thirteenth cervical vertebra. ‘The neuro-central suture is 
a nearly straight or slightly tricurvate line, with the central con- 
yexity downwards. A tricurvate ridge, with the central convexity 
upwards, runs along the whole length of the centrum between the 
articular edges or rims, at a level 0-4 inch below the neuro-central 
suture; it defines the upper edge of the nearly oval costal pit, which 
is 0-9 inch long, 0:45 inch broad (d.v.), and obscurely divided into 
two by a faint median longitudinal ridge or closed furrow. The rib 
has been displaced downwards; and its ovate-lanceolate posterior 


* (a.p.) antero-posterior, (l.1.) from side to side, (d.v.) dorso-veutral. 


448 PROF, W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


prolongation extends with a downward direction as far back as the 
posterior edge of the centrum to which it belongs. 

Near the articular ends the centrum is roughened by a few small 
irregularly scattered tubercles, which become larger and more nume- 
rous in succeeding vertebra down to the twenty-fifth. The neural 
spine has been broken away ; so that the total height of the ver- 
tebra cannot be determined. The line between the zygapophyses is 
1°9 inch from the base, and 0:9 inch from the neuro-central suture. 

vu. The seventh vertebra has a total height of 2°95 inches; the 
centrum is 1:°375 inch long and 1°5 high. ‘The well-marked neuro- 
central suture is 1-1 inch from the base. The tubercles near the 
articular ends have become larger and more numerous. The rib is 
distinctly hatchet-shaped, and consists of a blade-like upper part and 
a lower handle-like horizontal process ; a deep incision separates the 
handle from the blade in front, and the front end of the handle does 
not reach the anterior edge of the centrum by about an inch. ‘The 
posterior margin of the blade slopes gradually down, and curves 
gradually into the handle, the posterior prolongation of which ex- 
tends a short distance beyond the posterior edge of the centrum, 
from which the rib proceeds. 

The anterior zygapophysis is, as in the other anterior cervical 
vertebree, turned inwards and upwards; below the line of the zyga- 
pophyses the neural arch is ridged in a direction crossing obliquely 
from the anterior edge of the anterior zygapophysial facet down- 
wards to the posterior edge of the neuro-central suture, the ridges 
being most marked near their origin and termination. The distance 
from the anterior to the posterior zygapophysis is 2°3 inches. From 
the line of the zygapophyses to the base of the centrum is 2°0 inches, 
to the top of the spine 1°35 inch. 

The neural spine has somewhat the outline of a Phrygian cap seen 
in profile; it has a gentle convex slope backwards in front, and a 
short sigmoid curve behind; it rises from the middle of the length 
of its centrum, and hangs over the anterior quarter of the centrum 
next behind. It is smooth below, but roughened towards the distal 
end. 

The vertebre increase in size and change in the relative size of 
their parts as they pass backwards: down to the fifteenth (xv), pro- 
bably to the twenty-second (xxir), the articular face of the centrum 
is an ellipse, with the major axis vertical (d.v.); at the twenty-third 
(xxur) the diameters are about equal, and continue so to about the 
twenty-fifth (xxv), beyond which the horizontal diameter (1.1.) be- 
comes the larger, and continues to increase over the vertical down 
to and beyond the end of the cervical series. The neuro-central 
suture becomes more sharply inflected in the middle, so that in the 
fifteenth (xv) the middle curve of the tricurvate line becomes trans- 
formed into a right angle. The zygapophysial facets acquire by 
degrees an entirely horizontal position ; they seem to have done so 
in the fifteenth vertebra. 

The neural spines increase more rapidly in size than the centra, 
and considerably change their form; at the eighteenth (xvm) the 


FROM THE LOWER LIAS OF CHARMOUTH. 449 


spine is broader at the distal end than in the preceding vertebra. 
The anterior margin is short and straight, sloping backwards; 
the upper margin curved, rising obliquely backwards; the posterior 
margin is sigmoid, convex backwards above, and concave below, 
nearly vertical. In the thirty-second (xxx1r) the outline is much 
sunpler: both anterior and posterior margins are straight, not quite 
parallel with each other, since they are further apart below than 
above; the upper margin is an elliptical curve, through which the 
anterior and posterior margins pass into each other. The base of 
the anterior part of the spine between the anterior zygapophyses is 
much compressed ; it broadens out immediately behind the origin of 
the anterior zygapophyses, and also over the posterior zygapophyses. 
The costal pits increase in size; but neither they nor the ribs show 
much sign of other change down to the twenty-sixth (xxvI) ver- 
tebra, when, however, preparations for change become evident. 

In the twenty-sixth, the rib being displaced allows the costal pits 
to be seen ; they are now quite separate oval depressions, 0°95 inch 
long and 0-9 inch distant from the anterior edge of the centrum, with 
which they are connected by a ridge which continues the anterior 
margin of each forwards. 

In the twenty-seventh (xxv) all trace of rugosity has disap- 
peared from the surface of the centrum, and it is now quite smooth; 
this continues to be the case throughout the rest of the vertebral 
column. The neuro-central suture is marked by a swelling ridge, 
particularly prominent in the central part of its course. 

In the twenty-ninth (xxrx) the thickening of the lower end of 
the neural arch becomes more marked, and the margin of the costal 
pits is somewhat elevated. 

In the thirtieth (xxx) faint signs appear of a ridge proceeding 
from the swollen end of the neural arch to the upper margin of the 
costal pit; in the thirty-first (xxx1) this and the ridge connecting 
the anterior margin of the costal pit with the anterior edge of the 
centrum have both become more marked. In the thirty-second and 
thirty-third (xxx and xxxi11) the costal pits begin to rise higher 
on the centrum, and become more posterior ; the upper edge of the 
pit swells into a marked ridge, and is connected by the previously 
mentioned ridge, which (now become very prominent) ascends from 
it to the swollen end of the neural arch. 

In the thirty-fourth (xxx1v) the lower costal pit has almost dis- 
appeared, and the upper and anterior margins of the remaining pit 
are swollen into a strong crescentic ridge, which is joined in the 
middle by the ridge descending from the neuro-central suture. 

In the thirty-fifth (xxxv) the thickened lower end of the neural 
arch and the ridge arising from the anterior margin of the costal 
pit form together a single vertically descending median ridge, which 
extends more than halfway down the side of the centrum. That 
part of the ridge contributed by the neural arch is more swollen 
than the other, and curves forwards as it descends, joining at an 
obtuse angle the part contributed by the centrum, which is sigmoid 
in outline ; the general form of the united ridge is much like that 


450 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


of a bracket (}), the middle point of the bracket standing for the 
point of junction of the two constituent ridges. 

In the thirty-sixth (xxxv1) the lower end of the neural arch is 
still more swollen, and has retreated further up the centrum; it 
more abruptly joins the costal ridge, which has become almost 
straight, and more prominent; it extends down the middle of the 
centrum to within 0°65 inch of an oval nutritive foramen, which lies 
on one side of the concealed middle line of the base of the centrum. 

(xxxvi1) An abrupt change in the character of the costal ridge 
takes place in the thirty-seventh vertebra; it has become greatly 
enlarged, to form a simple transverse process, which, curving down- 
wards and backwards from the neural arch, ends in an oval facet, 
looking obliquely backwards and downwards; the lower edge of the 
facet rests upon the centrum, the pedicel of the transverse process 
having as yet only an upper and not an inferior margin. The rib 
is no longer hatchet-shaped, but of the ordinary half-hoop form ; it 
is nearly cylindrical down to 1:3 inch from the transverse process, 
and then expands laterally so as to become somewhat triangular in 
section ; the line along which this change takes place is marked by 
a strong ridge, oblique to the axis of the rib. A transverse fracture 
across the front of this vertebra and the overhanging posterior zyga- 
pophyses of the preceding vertebra shows that the zygapophysial 
facets are not horizontal (as one might have conjectured), but much 
inclined, the posterior looking outwards and downwards, and the 
anterior inwards and upwards. 

(xxxviit) The base of the transverse process of this vertebra ex- 
tends a little more than halfway down the centrum; the facet is 
borne on a distinct pedicel, and looks a little less backwards than 
that of the preceding vertebra. This I take to be the last cervical 
vertebra, the transverse process of the next vertebra appearing to 
arise wholly from the neural arch. I use the word ‘“ appearing” 
definitely, since, in the absence of any well-defined neuro-central 
suture, it is difficult to say certainly what the exact constitution of 
the transverse process is. It is clear, however, that the process in 
this vertebra extends a little below the dorsal half of the centrum ; 
and this is presumptive evidence* that it is cervical; while in the 
next vertebra it does not, but is wholly confined to the dorsal half, 
and thus should be the first dorsal. Moreover, owing to a difference 
in the colour of the substance of the centrum and that of the neural 
arch, the latter being black and the former brown, in this region of 
the vertebral column, it is possible to detect in the transverse pro- 
cess of the thirty-eighth vertebra bone contributed by the centrum ; 
in the transverse process of the thirty-ninth no certain indication 
of bone so contributed is to be found. The possession of hatchet- 
shaped ribs was at one time included by Professor Huxley in the 
definition of a cervical vertebra ; if this should be regarded as an essen- 
tial character, then the vertebre thirty-seven and thirty-eight would 

* «There is reason to believe that the neurapophyses do not extend upon the 
bodies of the cervical vertebree beyond their dorsal half.”— Hux.ey, on Plesio- 
saurus Ktheridgit, Quart. Journ. Geol. Soe. vol. xiv. p. 282 (footnote). 


FROM THE LOWER LIAS OF CHARMOUTH. A51 


be excluded from the cervical series, and there would be only thirty- 
six cervical vertebree. If it were desirable to make a natural group- 
ing of the vertebre of this skeleton without reference to those of 
other species, one would not hesitate to draw the line between cer- 
vical and dorsal at the end of the thirty-sixth vertebra; for all 
down to the thirty-sixth are without transverse processes, but pos- 
sess hatchet-shaped ribs, while past the thirty-sixth the hatchet- 
shaped rib disappears and unmistakable transverse processes corre- 
spondingly arise. As, however, it is convenient to adhere as closely 
as possible to existing conventions, in order to facilitate the com- 
parison of species, I have been governed in my determination of the 
last cervical by the fact that a part of the transverse process borne 
by it does clearly seem to be contributed by the centrum as far back 
as the thirty-eighth vertebra; and thus I have included as cervical two 
vertebree which would certainly seem more in place in the dorsal 
series. Professor Seeley’s plan of calling those vertebre in which 
the transverse process is passing from the centrum onto the neural 
arch “pectoral” has much to recommend it, and might fairly be 
applied to the vertebree thirty-seven and thirty-eight. Indeed there 
is just a shade of doubt in my mind whether vertebra thirty-nine 
should not also be called pectoral; for its transverse process appears 
to havé a little brown bone like that of the centrum at its base, 
and the rib it bears is bifurcate near the head and short, while the 
succeeding vertebre bear ribs with a simple proximal termination 
only. 

The determination of the position of the last cervical vertebra is not 
only important as giving us the number of vertebre in the neck, but 
also because it furnishes us with a necessary datum for the measure- 
ment of the length of the neck, and hence for ascertaining that im- 
portant character, the ratio of the length of the head to that of the neck, 
or, as we may briefly term it, the cervico-cephalic index. Fortunately, 
in the case of the species under consideration the neck is so long that 
one or two vertebree more or less can make very little difference to 
the value of this index, the thirty-seventh and thirty-eighth vertebre 
measuring together not more than five inches. The thirty-eight 
cervical vertebree measure 83°25 inches, or 6 feet 11 inches; and 
the cervico-cephalic index is 24:1. A Table is here appended, 
giving the dimensions of the cervical vertebrx, so far as they are 
ascertainable: it will be seen that the constancy in length which 
Prof. Owen* regards as characteristic of the cervical vertebra of - 
the Enaliosauria, and only exceptionally absent in Phiosawrus, has 
no real existence, and also that no single vertrebra can well be taken 
as typical of the remainder. Hence the importance of such a Table 
as this as a help in the specific identification of separate vertebre 
will be apparent. It may be worth while to call attention to the 
abrupt manner in which some of the changes in dimensions occur, as 
for instance in the length of the centrum in passing from the twenty- 
second to the twenty-third vertebra; and again, to the abnormal 


* Report Brit. Assoc. 1841. Report on British Fossil Reptiles, p. 63. 


452 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


variations which occasionally appear, as, for instance, in the twenty- 
eighth vertebra, which is longer not only than its predecessor, but 
also than its successor; the same is also the case in the thirty-sixth 
vertebra: the difference is too large to be explained as an error of 
measurement, and can scarcely be the result of mechanical compres- 
sion; it seems rather to be concomitant with that wide departure 
from the ordinary Reptilian type which the Plesiosaurian neck 


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Table of Measurements of Cervical Vertebre. 
Centrum. Zygapo- | Zygapo- | 2Y82P0- | Neuro- eae 
a pie | Total | Physis | physis payeis cone suture Noe 
WO een || iitole toiendl suture | 
vertebral], ength.| Breadth.| Height.) “5 base. | of spine.| ZY84P0-| _ to costal | Vertebra. 
(a.p.) (1.L.) (d.v.) physis. | base. pit. 
Ih 
2. 
3 12 | 0°75 1-4 1:9 12 0-4 3 
4, 25 Sass 1:4 1E9 21 Teal 0-4 
5 295 1:4 2:0 1:35 1-1 0:4 9) 
6. 375 14 29) 2:0 ; 2°3 11 0:5 
7, 375 15 2°95 2:0 : 2°3 1-1 0:5 
8. 3795 15 2°95 2:0 1:45 2°3 11 0-5 
9. "379 15 32 2:0 2:3 11 0:5 
10 (i 15 2:25 2-45 | 1:05 | 0°5 10 
11 7 ey 2:3 ravasd) | 11) 0-5 
12 7 1-4 1-7 2°45 0 12 0-5 
13 8 145 | 1°75 2:45 14 | 05 
14 9 14 19 2: 1:9 1-4 0-5 
15 0 15 20 4-0 2:1 275 | 1:5 0-6 15 
16 ‘il aoe 2-1 4-6 2:2 2:35 | 1:55 | 07 
17 2 2-1 4:8 2°45 3°05 | 1:55 i 
18 2 2°2 oul 2°45 3°05 | 1:5 a 
19 2 2:2 5:2 1 
2 2: 5:4 2°8 1 20 
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FROM THE LOWER LIAS OF CHARMOUTH. 453 


Dorsal Serves. (Plate XXIII. fig. 2.) 


The dorsal vertebree are well exposed ina fine and complete series 
on the underside of the specimen. ‘They have the same general 
character as the last two cervicals—the centrum being smooth, not 
tuberculated, its height (d.v.) and length (a.p.) being about equal, 
and both shorter than the breadth (1.1.). The maximum dimensions 
of the centrum appear to be attained in the fortieth vertebra, in which 
the length and height are each 2°8 inches, and the breadth 4:8 inches ; 
its total height, however, is only 7:6 inches, being less than that of the 
forty-third vertebra, which is 8:2 inches high ; behind the fortieth the 
centra decrease in size, and a little more rapidly in breadth than in 
the other dimensions. ‘The transverse processes rise upon the verte- 
bre from the thirty-ninth (xxxrx) to the forty-second and forty-third 
(xLII and xt1r) beyond which they spring from the neural arch along 
the zygapophysial line ; they maintain this position down to the fifty- 
sixth (Lv), past which they begin to descend and also change in 
character. At first, as in the fortieth vertebra (x1) or second dorsal, 
the pedicel of the transverse process projects outwards at right angles 
to the vertical plane given by the flat side of the neural spine; 
passing backwards this angle is much diminished, so that in the 
forty-fourth (xxv) vertebra it is only 65°; behind this it begins to 
Increase again, and at length becomes 90° at the fifty-fifth (Lv) 
vertebra, or seventeenth dorsal. 

The length of the transverse process increases slowly down to the 
forty-seventh (xivir), in which it is 4-3 inches long; behind this it 
slowly shortens and becomes 1:5 inch at the fifty-eighth vertebra. 

The dorso-ventral diameter of the base of the transverse process 
is at first, as in the fortieth vertebra, 2-1 inches; but it rapidly 
diminishes, so that at the forty-third it has become 1:2 inch; 
past the forty-third it remains pretty constant as far as the fifty- 
sixth vertebra. At the fifty-sixth important changes commence ; the 
transverse process loses its straight boldly projecting form and droops, 
as it were, into a curve, assuming the character of the last cervical 
transverse process. In the fifty-eighth vertebra the facet is inclined 
downwards and backwards ; but its form cannot be fully made out, 
as its lower half is concealed by the head of its rib. 

The neural spines are parallel-sided, and truncated above by a 
straight or very gently curved distal margin. They attain their 
greatest length and breadth in the fortieth to forty-third vertebre, 
past which they diminish in size slowly. In the early part of the 
dorsal region the spines are inclined backwards at a slight angle ; 
thus in the fortieth to forty-second vertebre the axis of the spine 
makes with the zygapophysial line an angle of 64°, 2.¢. it slopes 
backwards 36° from the vertical ; posteriorly the backward inclina- 
tion diminishes and the spine becomes at length vertical; this is the 
case at the fifty-fifth vertebra ; still more posteriorly the inclination 
becomes reversed and the spine slopes forwards; thus in the fifty- 
seventh vertebra it makes an angle of 93° with the zygapophysial 


Q.J.G.8. No. 147. 25 


454 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS. 


line, 2. ¢. itis inclined forwards at an angle of 3° from the vertical ; at 
the sixtieth (Lx) vertebra (1st sacral) the spine has resumed its 
backward inclination. - 


The fabs. 


The early dorsal ribs for about two inches from the proximal end 
are almost straight ; they then somewhat rapidly bend into a curve, 
which is steeper near its origin and straighter towards the end. At 
the fifty-first vertebra the curvature of the rib has become less, in 
the fifty-third much less; and at the fifty-fourth the rib is straight. 
The longest ribs are those of the forty-seventh to the fiftieth ver- 
tebre ; behind the fifty-second they rapidly shorten, those of the 
fifty-sixth being only 5:4 inches, and of the fifty-ninth 2-7 inches 
long. 

For a short distance from the head the ribs are roughened with 
irregular longitudinal ridges, which are most marked in the anterior, 
and absent in the last few posterior dorsal ribs. All possess simple 
proximal ends, except the first dorsal, which gives off a short process 
just below the head. 


Sacral Vertebre. 


The two vertebre regarded as sacral are the sixtieth and sixty- 
first (Lx and 1x1). In them the transverse process has become very 
short, little more than a raised facet, the surface of which, however, 
is larger than that of the preceding transverse process of the last 
dorsal. It obviously consists of two nearly equal parts—an upper 
contributed by the neural arch, the articular face of which is a plane 
surface, meeting along a horizontal line at an obtuse angle the 
similar plane surface of the inferior moiety contributed by the cen- 
trum. ‘The ribs are short (1:9 inch long) and slightly expanded 
at the distal ends. The neural spines are inclined backwards, making 
in the sixtieth an angle of 80°, and in the sixty-first of 78°. 


Caudal Vertebre. 


If the determination of the sacral vertebree be correct, then there 
are five caudal vertebrae, the dimensions of which are given in the 
appended Table (page 455) of measurements for all vertebree past the 
last cervical. The spines are broken away from them all except the 
first, in which it is suddenly inclined backwards at a much greater 
angle than that of the last sacral. The zygapophyses are nearly 
vertical. . 

The transverse processes are now represented merely by pits with 
raised margins, only the upper part of which is furnished by the 
swollen end of the neural arch. The ribs remain short; but that of 
the first is longer than that of the last sacral. 


Table of Measurements of Postcervical Veriebre. 


. __|Inclina- a 3B 
Length | Length |Inclina- Peo ouapes 
Nor Centrum. ay | | pve et © of tion of pa Honeth & i S gq = a 
oem Total | @Y ee P- | to end | a ip of  |2nterior| spine | spine Bes Length fae 6 8 vs a8 a ae 
height. of ae margin} along to of rib. epee De : 

tebra. base. spine, | 2782P- | spine. of middie line of | Proves verse BF 4,8 verse 
Length. | Breadth.| Height. spine. | line. | zygap. ee Process: He cao See BCs a 
es pe eS 5 are ie oe eee eee 
in. in. in. in. in. in, in. in. in, in. . 5 in. in. in, in, in 


39. 2°6 3D 2'8 ae 4:2 
40. 2°8 4-1 2:8 


et et 


SvSr Or Or Ot 


Pt et tt tt tb 
ell \2 Meal NO We NO SUC Os 


8 

OLS ~ SiS 
Med DOMOODOMED DOO 

OU ON 


SCHWNHHARAAMWOWNNH OO 


° 


Over 


Or 


Or or 


on 
= 
eo 
fon) 
ES ° 
oo 
Or 
Ye) 
S 
We) 
S 


a 
° 


— 
a 
On 


(e2) 
j=) 
-J 
oe 
— 
= 
ler) 
H FOU SIS GO CO CO OE HO DY CO WONT Oo OVO 


. 
° 


Ourcr 


WMOH Odo WH OHOH 


M09 00 Gir: 


oO 
to 
bo 
> 
iw) 
bo 
Or 
bo 
tt et pt 


456 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


Pectoral Arch. 


The bones of the pectoral arch are all present, though somewhat 
displaced from their original position. They consist of the furculum, 
coracoids, and scapule. 


The Furculum. 


This (fig. 2) is a large bilaterally symmetrical plate of bone, convex 
ventrally from side to side, with two long thin tapering lateral wings, 
one on each side, directed backwards and slightly dorsally; a tri- 
curvate anterior margin, the central curve being a large semicircular 
excavation, which passes into a curve convex forwards on each side; 
and a gently curved posterior margin convex backwards, with a 
narrow deep incision running forwards along the median line, or 
axis of symmetry. The lateral curve of the anterior margin passes 
insensibly into the front margin of the lateral wing; the posterior 
margin meets the posterior margin of the wing in a rounded 
angle. 


Fig. 2.—Diagram of the Furculum. (Scale 3.) 


\ 


he 


If one draws a line from the central point of the curve, which we 
have just called a “ rounded angle,” parallel approximately to the 
anterior margin of the bone, we shall divide the body into two parts, 
the anterior of which is much thicker than the posterior, being at 
least 1 inch across ; it is of somewhat dense or close texture super- 
ficially, but loose and open in the middle: the posterior part is very 
thin, a mere lamella of bone. The anterior part may be distin- 
guished as the ‘‘ body” proper of the bone; the posterior, which is 
divided into two by its median longitudinal fissure, is a pair of 
“lapels;” and thus with the “wings” we have five distinct re- 
gions present, but of true sutures I cannot find a trace; the whole 
appears to be a single piece of bone, though having regard to the 
great difficulty there often is in discovering sutures which do really 
exist in fossilized bones, I should not wish to be thought too posi- 
tive on this point. 

The bilateral symmetry of the bone and its median longitudinal 


FROM THE LOWER LIAS OF CHARMOUTH. 457. 


notches, point to its connate nature; its position in front of the 
coracoids, between the prescapular processes of the scapule, points 
to its clavicular origin: I regard it therefore as representing a pair 
of fused clavicles, which repeating the behaviour of the coracoids, 
have expanded into extensive plates over the ventral surface. An 
interclavicular element appears to be absent; there is no room for it, 
except in the position conjecturally assigned to it by Protessor 
Seeley, who has suggested that it forms the anterior middle part of 
the bone. This, however, is a position which it occupies in no other 
known reptile, as it is always more or less posterior instead of an- 
terior to the clavicles. Since writing the first part of this paragraph 
I have been able to devote a few minutes to an examination (which L 
wish could have been less hasty) of a loose specimen of Plesiosaurian 
furculum, preserved in the British Museum, the same bone, I fancy, 
that is figured as a sternum in Hawkins’s monograph. It certainly 
shows traces of sutures, and is marked on the surface by strive, which 
appear to indicate a median and lateral elements. It has a suggestive 
resemblance to the clavicles and interclavicle of a Chelonian like, say, 
Trionyx. But it differs considerably in form and appearance from 
the furculum of our species ; so that it is doubtful how far it can be 
used as a guide. Very possibly the furcula of different Plesiosaurs 
may differ in composition, as they do in Birds, an interclayicle being 
sometimes present and sometimes absent. 

There is another difficulty attending the interpretation of the fur- 
culum ; and that lies in its position beneath the prescapular ends of 
the scapule, which overlap its posterior lapels. In all recent rep- 
tiles the clavicles are superficial to the scapule, while here just the 
reverse is the case. ‘This is proved by more than one well-preserved 
specimen in the British Museum, showing the scapular processes 
abutting on the body of the furculum, and also by Lord Enniskillen’s 
specimen of P. macrocephalus, which affords us a dorsal view of the 
left clavicle overlapping the dorsal surface of the scapula (fig. 3). 


Fig. 3.—Diagram showing the lefi side of the Pectoral Arch of 
P. macrocephalus, seen from behind. (The wing of the furculum 
conceals the termination of the scapula.) 


458 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


This anomalous position I altogether fail to explain : if the lapels 
could be shown to be precoracoids, all would be clear; but this view 
is not without difficulties. 

The dimensions of the bone are given below :-—— 


inches. 
Antero-posterior diameter in the median line 
(GS, 5. BAD)» 66-006 3°65 
Maximum antero- -posterior diameter (ib. ¢ dl) 6-4 
Antero-posterior diameter from a tow...... 5°25 
Breadth) (Gwaceitheslimeti)) eae er eee 8:6 
Length of wing (g h).. BU SEES eek nO ee 
Width of anterior excavation (¢ 9g) prance 2°55 
Depth of anterior excavation (av) ........ 1:05 
Depth of posterior incision .............. 2a 


The Coracords. 


The coracoids (fig. 4) have the usual Plesiosaurian form, presenting 
together a close and almost ludicrous resemblance to the front of a 


short jacket. 


Fig. 4.—Diagram of the Right Coracoid. (Scale 3.) 


The inner margins of the bones meet for half their length in a 
straight median harmonia, diverging gently outwards from each end 
of it. The outer margin has a simple concave sweep backwards 
behind the articulation for the humerus ; the anterior margin pro- 
jects in an elliptical curve in front, the outer edge of the curve sink- 
ing into a curve backwards, which joins the almost straight margin 
of the scapular articulation. The bone is thickest where it furnishes 
the articulations for the scapula and humerus. Thence it continues 
with only slightly diminished thickness along a ridge or keel, which 
extends transversely to its inner margin. This ridge, which rises 
from the ventral surface of the bone, and as shown by a transverse 


FROM THE LOWER LIAS OF CHARMOUTH. 459 


fracture, is scarcely, if at all, marked on the dorsal surface, is de- 
fined in front by a curved line, which commences near the anterior 
end of the scapular articulation, curves backwards to the middle of 
its course, and then forwards till it ends against the inner margin of 
the bone; posteriorly it is defined by a line very slightly curved, 
convex backwards, which commences from the hinder end of the 
humeral articulation and passes very slightly backwards, also to end 
against the inner margin. From this ridge the bone thins rapidly 
away in front and behind, more rapidly in front, till it ends in a 
thin edge. 

The posterior region of the right coracoid is traversed by several 
lines of fracture, on one side of which the surface of the bone remains 
higher than on the other, and thus forms little cliffs. A broken sur- 
face, dividing the bone across, reveals its internal structure, and shows 
between its dense outer layers a more cancellous open tissue in the 
middle ; not only so, but in some places the middle of the bone is 
occupied by a layer of calcite; and this is thicker on that side of a 
fracture where the surface is higher, and thinner on that side where 
it is lower. This calcitic layer is probably due to the replacement 
of cartilage ; and it is thinner where the bone is thinner on the side 
of a crack, because the cartilage had there been squeezed together by 
pressure of overlying strata. 

The measurements of the right coracoid are given below :— 


inches. 
Maximum length (fig. 4, mxv)............ 17:375 
Henethvot harmonia Gb: @p) ............. 10°5 
EBenea chee ens?) Nea c0-) she iced Seles Siete esses 86 
Breadth along median ridge (1b. hr). sox!) 


Distance from anterior end of coracoid to 
posterior edge of glenoid cavity (ib.mt).. 10:0 
Length of chord of anterior concave margin 


of coracoid (ie FOG)! ane hoe 2°75 
Height of are of anterior concave margin of 
EOLACOrdG (Tb fi) is ah uals ed. ifn se. A 0°375 


Length of articulation for thescapula(ib.ds) 3°3 
Length of articulation for the humerus(ib.s¢) 3°2 


Thickness at articulation for the humerus.. 1°6 
Length of chord of posterior and outer lateral 

ATEVO (Os G2) Ae eo eee 8:3 
Height of are of posterior and outer lateral 

GUIET® (GD. 70) elotame ieee atch eon ee 2°3 
Length of chord of posterior convex margin 

(10s BAD) 36 ous bthio aan a Geass ee ean eo 
Height of arc of posterior convex er 

(ib: 0). 33 


Thickness of median ridge where broken | in 
middle of its course (left coracoid)...... 0°91 

Thickness of anterior region of coracoid.... 0°25 

Thickness of posterior region of coracoid .. 0°6 


A60 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


The Scapula. 


This (fig. 5) consists of a ventral plate, from the outer lateral margin 
of which a lateral plate arises and ascends towards the dorsal surface, 
its plane being inclined to that of the ventral plate at an angle of 
about 90°. The ventral plate is longer than broad, thicker behind 
than in front, truncated by a slightly convex edge anteriorly, 
bounded by a widely open V-shaped margin’ behind, the inner stroke 
(cr) of the “ V ” representing its articulation with the coracoid, the 
outer stroke (v p)its share in the glenoid cavity tor the humerus. The 


Fig. 5.—Diagram of Right Scapula. (Drawn reversed, scale 3.) 


inner lateral margin is concave, the outer straight ; and from along its 
whole length the lateral plate arises. The latter is like a long sca- 
lene triangle in shape, the apex iying in front, the longest side being 
that common to it and the ventral plate, and the next longest its 
outer edge ; the base of the triangle is not a straight line, but a deep 
concave curve arising from the anterior edge of the glenoid cavity, 
passing forwards and upwards and then backwards to the end of the 
outer edge. The lateral plate, when complete, is not bounded by a 
simple outer or upper margin, as here described, but is prolonged 
upwards along its posterior third into a thin narrow ascending pro- 
cess: this process is broken off both scapule in our specimen; and 
the posterior third of the outer edge of the lateral plate is conse- 
quently a surface of fracture. 


Dimensions of the Scapula. 


inches 
Length of outer edge of ventral plate (fig. 5,ap) ........ 81 
Breadth at distal end (ib. ab) ............. Perr... 2:4 
Length of chord of concave inner margin (ib. 6c) ........ 5:05 
Height of arc of concave inner margin (ib. d) .......... 1:2 
Length of articular surface for coracoid (ib.cr).......... 22 
Length of articular surface for humerus (ib. 7p) ........ E93 
Length of chord of posterior concavity of ascending plate 
Gbl mm) Rien: 2-1 


Height of arc of ‘posterior concavity of ‘ascending plate 


FROM THE LOWER LIAS OF CHARMOUTH. 461 


Restoration of the Pectoral Girdle. (Plate XXIII. fig. 3.) 


The displaced bones retain their original outline so perfectly, that 
if it were possible to move them relatively to each other into their 
original positions the pectoral girdle would be well restored; as, 
however, the stony matrix in which they are imbedded renders this 
impossible, [ have made careful outline drawings of the several 
bones, and then cut these out and fitted them together, being guided 
in doing so by a direct study of the bones themselves. The result 
is given in the diagram (Plate XXIII. fig. 3). 

The furculum lies in front in the middle line; and the prescapular 
processes of the scapule abut each on its own side upon the outer 
posterior border of the furcular body proper, covering its wings, 
which appear to lie in the angle between the ascending scapula and 
its prescapular process. The lapels face the anterior projecting 
convexities of the co1acoids without touching them. There is thus 
produced a single continuous foramen, bounded laterally by the 
concave inner borders of the scapule, anteriorly by the furcular 
lapels, and posteriorly by the anterior margin of the coracoids; it is 
wide from side to side, narrow from before backwards, and roughly 
resembles in form two brackets joined face to face, thus :—@— 

In the fact that the foramen is not subdivided into two by the 
overlapping of the furcular lapels by the coracoids, this species differs 
from some other species of Plesiosaurs; a corresponding difference 
is displayed in the pelvic arch, the foramen between the pubes and 
ischia, likewise double-bracket-shaped, being continuous from side 
to side, and not divided into two, as happens in most other Plesio- 
saurs. 


The Humerus. 


The right humerus is carried over to the left side, and lies with 
its flat posterior surface uppermost (Pl. XXIII. fig. 1), For the proxi- 
mal third of its length it is a thick cylindrical bone, with an ellip- 
tical transverse section, the major axis being twice the length of the 
minor axis (4 inches and 2 inches respectively); it then widens out 
into a broad plate-like distal portion for the remaining two thirds of 
its length. The ulnar margin is almost straight, only slightly convex; 
the radial more curved and concave. It is covered superficially 
with irregular longitudinal ridges, more abundant on the radial than 
the ulnar side, which is almost smooth, and most marked near the 
ends of the bone; at the anterior end they become broken up into 
irregular tubercles. 

The left humerus is similar to the right; but its surface is 
smoother, and its proximal half less elliptical or more circular in 
section; it begins to widen a little past the middle of its length, 
widens and flattens then rapidly, becoming very thin (7 inch) 
towards its distal edge. The more rapid expansion of the left than 
of the right humerus is almost certainly due to compression, though 
without the latter bone for comparison we should have nothing to 


462 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


indicate this; hence a possibility to be borne in mind in making 
specific determinations of isolated bones. 


The Radius. 


The right radius is a straight, almost parallel-sided bone, elliptical 
in transverse section, truncated by slightly convex ends; it has a 
simpler outline than is usual in Plesiosaurs. The left radius, which 
has suffered more from compression, has more of the usual Plesio- 
saurian outlines: its outer edge is gently concave, its ulnar margin 
tricurvate, the middle curve being concave inwards; and its once 
cartilaginous articular surfaces have been squeezed beyond the edge 
of the flat surface of the bone. It is much ridged longitudinally, 
the ridges diverging a little from the middle towards the ends. 

Representing the length of the humerus by 100, that of the radius 
will be 37; and this number may be called the humero-radial index. 
Its value for other species is found in the table on page 477. For 
P. rostratus it will be seen to be almost the same as for P. Cony- 
bearer. 


Dimensions of Humeri, Radi, Ulne, and Femora. 


| Diameter of | Diameter at Diameter of Length 
| head. middle of length. distal end. 
[Saye ca from capi- 
Ra. (Ol ANG IP Wen, WAL wy TP. Ra. Ul. A. P. | end to end.' tular ridge 
_—— to end. 
Humerus :— chord.| are. 
Right...... 35 | 2:9 39 2°0 6:7 S21 1-22) ass | 15 A. 
14-25 P. 
Mhefti i teoe (?)36 | 3:2 39 2°0 nea BD) | oa 14:8 14°8 P. 
Right...... 35 1 3:6) 84) 0 2:0) 67 0s we alee 1425 | 12-75 M. 
13°5 A. 
ett s 35 | 38 37 16 ace aap allege 14:37 13°37 
Right...... eas aus SO os 35 375 | 0-7 5°45 ne 
ett: eee. 39 | 08 315 | 055 | 3:5 3°5 | 0:55 5:0 4°75 
Right...... ARESy Wo 39 ae 4°85 | 65 | O7 55 
MHGLt vate) 2.8 | OF 37 re 4-7 6:0 56 


Ra. Ul. means from radial to ulnar margin; A. P. means from anterior to 
posterior margin ; Chord means measured along chord of curve; Arc means 
measured along curve itself. In the last column A., M., P. mean measured to 
anterior, median, and posterior edge of distal margin respectively. 


The Ulna. 


The right ulna, like the right radius, retains more of its original 
form than the corresponding bone on the left ; for although its distal 
two thirds are much flattened, its proximal third still presents its 
original thickness. 

This uncompressed portion has a smoother surface than the other, 


FROM THE LOWER LIAS OF CHARMOUTH. 463 


which is distinctly ridged; the entirely compressed left ulna is 
ridged all over ; and this leads one to suggest that the ridges on fossil 
bones may in some cases have been subsequently produced by me- 
chanical pressure. Let the soft cancellous interior of a bone be 
crushed together, and the denser outer layers, in adapting them- 
selves to a more circumscribed area under pressure, may possibly 
become finely wrinkled, and thus give rise to a spurious appearance 
of ridges. 


The Carpus. 


The carpus, 7°25 inches in breadth, consists of two rows cf three 
bones each, which diminish in size from the ulnare to the radiale. 
They are polygonal bones, with the dimensions given in the table 
below. In the left manus the distal row of carpal bones alone 
bears the fingers, the radiale carrying one, the intermedium and 
ulnare two each. In the right manus the ulnare of the proximal 
row appears to bear one finger, the ulnare of the distal row two, 
the distal intermedium one (but it contributes a small facet for the 
adjoining digit of the ulnare), and the distal radiale, as in the left 
manus, one. 


Length. Breadth. 


; 
Radial. Inter- Ulnare. || Radial. | Inter- Ulnare. 


medium. | medium. 


Proximal series... 16 J3 2°65 215 | 25 2°75 


Distal series ...... 1:8 14 2:9 2 Pale ice | 2a 


The Manus. 


The hand, where broadest, is 8-5 inches across. It consists of five 
digits ; the first, with five phalanges, is incomplete in both hands; the 
rest are complete in the left manus—the second, third, and fourth 
having nine, and the fifth eight phalanges. The third and fourth 
fingers are the longest. The phalanges have the usual form, a 
compressed hour-glass outline,—except the most distal, which is 
triangular, and apparently equivalent to the proximal half of one 
of the other phalanges, the distal half being suppressed: it has 
much the appearance of an ungual phalanx, and may very well 
have borne a nail. 

The measurements of the phalanges are averages of those of the 
right and left manus, and in the case of the breadth, of the distal 
and proximal ends of each phalanx. 


464 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


Phalanges. 
Length. Breadth. 
Tih ta aoa ave {. 1 s/s er ane ave Vv. 
Tee tap 19 25 30 30 31 1:25 115 1:35 1:35 75 
BIS dered 18 23:5. 25 26 2:45 weet 1:85) cA 2 oie 
DID ics: SO MEDI AIP Seeks. Diva 12) 4b 1-4 ae eles 
LVR asthe oe 18 1°75 1:95 1:95 2:0 i ce, 1:25) L2o eo 
See id 1°75 2:05 1.7 1:65 1:66 PYb oo fi5: 131 O95 aes 
Wilson cnt 18S hei) leek hn4) a 9 10 O08 O07 
Video a esate Pa ESSs be) ee oils) 0:72; 072 107 tas 
VAUD Se cies 10 O8 1:0 0-45 055 045 0-45 0:35 
ch Coates 08 O7 O68. 044 03 03 


Total length a ? 17 18 18:25 16:8 


fingers ...... 
The Pelvis. 


A restoration of the pelvic girdle, obtained in the same way as 
that of the pectoral arch, is given on Pl. XXIII. fig. 4. 


The Pubes (fig. 6). 

These are more or less quadrangular plates, which meet each other 
in a straight median symphysis along nearly their whole length. 
The anterior margin is an undulating curve, with a general direction 
at right angles to the symphysial margin. The posterior margin, at 
first a little convex, becomes concave for half its length outwards 
from the symphysis, then almost straight and parallel to the ante- 


Fig. 6.—Diagram of the Right Pubis and Ischium. (Scale 3.) 


_Ylor margin as it articulates with the ischium; finally it curves 
forwards and outwards, to contribute its share to the acetabulum, 
and joins the outer margin, which has a slope obliquely outwards as 
it passes backwards from the anterior margin. 


FROM THE LOWER LIAS OF CHARMOUTH. 465 


The Ischium (fig. 6). 


This “ shoulder-of-mutton ” shaped bone meets its fellow for about 
half its length in the median symphysis. The m-shaped anterior 
margins of the two ischia meet to form together a bracket-like out- 
line (——~) ; the similarly shaped posterior margins of the pubes 
do the same; and the brackets facing each other give rise to a 
foramen, which is continuous from the pubic ischial smyphysis on 
one side to that on the other. In this character the pelvis resembles 
that of Murcnosaurus, Seeley, Q. J. G. S. vol. xxx. p. 206. 


The Ilium (fig. 7). 


The ilium has the usual Plesiosaurian form, a central shaft with 
expanded ends, compressed in different planes, which are inclined at 
about a right angle with each other. 


Fig. 7.—Diagram of Right Ilium. (Scale 3.) 
d 


P 


= 


Measurements of Pelvic Bones. 


Pubis. anne 
Womepieaisy lap ltysis (112. 6,09) ) Sicsasss os coc sce vonn sc scseccweesencc rues sas 5°5 
Breadth of bone from symphysis, and along a line at right angles to 
it, to anterior edge of acetabulum (ib. fl) ..........2...s.ceceeeeer eee ee 
Maximum antero-posterior diameter, drawn from d! .................. 81 
Length of diagonal from inner anterior angle to anterior edge of 
eice allo whtinie (AONE ae. oc Ania clas ene Ret acetates 10°3 
¥ from inner anterior angle to posterior edge of 
its share of acetabulum (@ 0) .................+..- 10:3 
3 “5 from outer antero-posterior to inner posterior 
EPELO (Lp) Meee fess 5, § Lavage Rai eases 9-0 
iy articular surface for acetabulum (f 0) .................0.s006 2°9 
a te Wn a {OIE TEG ATTN (DG) mesa seeaeossockosaceeseeenees 2°3 
a Chord Om posterior concavilby|(GIN)2 5,-2.0... eee ses eaetcass 39. 
Height of are a pk) RS ESR ee RRR a eae eS 2 1:25 
Ischium. 
Mienethohsympbysis (a p!, fig. G):..102:.-0.0.82.s0250002-cageceseep esses 5'5 to 6:0 
Breadth along a line at right angles to symphysis to posterior edge 
ni sapetnloml en (GD) ese owe nec eee aren AnE eR PPE Dose uaeaepcoddccore cou 
Length from outer edge of its articulation with the pubis to its pos- 
PeRAOIEAIS Loy (OM) kita Noses cerca eeeea toon seen adicc Soe ebatiies sane neces 10.1 
Length from anterior inner angle to posterior angle (N D)............ 7725 
Minima diameter across ‘neck 7 (72 &) , ~.....02-02sssecenssurmseeeacenee 2°4 
Length of symphysial margin for pubis (0 €) ................0 cee eseee eens 2°4 
ts Tee copula aneine (Oye ee te. aoc acincoveaemaceee acest ae 25 
. chord or anterior concavity (GIN) 5....2--s.a.cessoneseeeee sees 3°85 
ein onarc of anterior concavity (@) ..2....2-22-:0ssessscateecaeeeesson= 18 


466 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


Ilium. 
Right ilium. Left ilium. 
inches. inches. 
Mrenabla (07; O17) sos. we atecles idosin cite nyecednee Serene eae 6-9 72 
breadth of proximal end)(pp) \o 22... 224 scee= sees see 2°4 (2) 2°85 
3 distal (@'d)B3s eo ese e eee 2°3 1°9 
Minimum diameter of shaft (77 772)................0000 ae 1:0 


Note.—The proximal end of the right ilium is slightly broken, and its distal 
end compressed, probably by subsequent pressure. 


The Femora. 


These bones, which are smaller than the humeri, have the arti- 
cular head well defined by a sharp surrounding ridge; the articular 
surface itself is deeply pitted and tuberous, indicating the previous 
existence of a thick covering of cartilage which has since dis- 
appeared. 

The cylindrical proximal end is slightly constricted below the 
capitular ridge before it begins to expand. ‘The dorsal and ven- 
tral margins are almost straight lines, or only slightly concave 
curves, which diverge gently from the thick proximal to the broad 
and flattened distal end. ‘he distal end is truncated by a simple 
gently convex curve. 

The surface of the bone is roughened beneath the head, and 
strongly ridged at the distal end with-longitudinal lines, which 
diverge in conformity with the curves of the lateral margins of 
the bone. 


The Integuments. 


Remains of what appears to be some dermal structure have been 
stated by previous writers to occur in connexion with Ichthyosaurus ; 
and Charles Moore in particular has well described a thin layer 
having a wrinkled surface, which invests a large number of the 
Ichthyosaurs in the collection of the Bath Museum; but no one, so 
far as I know, has made mention of any similar investment in the 
case of Plesiosaurus. Great interest therefore attaches to the 
presence of a thin brownish film, with characteristic surface-mark- 
ings, which coats a considerable portion of the specimen under con- 
sideration. 

It is best displayed on the surface between the fortieth and 
sixtieth vertebrae, covering the bodies, transverse processes, and 
neural spines of the vertebra, the ribs, and the surface of stony 
matrix intervening between them. It also extends in a band over 
an inch broad, along the distal ends of the neural spines, at a little 
distance from them as now exposed. 

This band, which seems to restore the dorsal outline, ends in the 
pelvic region, where it covers the head of the right femur, and 
imbeds a small oblong bone, the smooth shining surface of which 
is raised into three parallel longitudinal ridges ; other fragments of 


FROM THE LOWER LIAS OF CHARMOUTH. 467 


similar bone are indicated near the same spot. Their presence can 
scarcely be accidental; and they may possibly be dermal plates. 
That they are not found elsewhere would simply point to the 
restricted distribution of dermal scutes in the species, they having 
originally been present in the pelvic region and nowhere else. 
The thin film, however, has nothing of the nature of scales and 
scutes, so far as we can see; it was a continuous membrane, not a 
collection of separate individual structures. It can easily be 
detached from the underlying surface, owing, it would seem, to the 
presence of a thin whitish layer, apparently calcite, which is more 
strongly adherent to the film than to the surface beneath. 

The surface of this film is variously marked ; but all the different 
markings may be described as essentially of the nature of 
wrinkles. In the film of the dorsal band they have the appear- 
ance of fine regular rounded ridges, giving the surface a re- 
semblance in some degree to “ corded silk;” elsewhere, as over the 
bases of some of the neural spines, the ridges lose to a great 
extent their straightness and regularity, take a tortuous course, 
though generally with one prevailing direction, and are more appa- 
rently mere wrinkles; but over the greater part of its extent an 
additional feature presents itself in the form of long, fine, parallel 
grooves, bordered by fine ridge-like margins, and looking as though 
they had been scored by a fine needle: they vary in distance from 
each other; but the best-marked are about 4, inch apart. They main- 
tain one general direction from before backwards on the bodies of 
the vertebree, the exposed outer sides of the ribs, and on the stone 
between them, from the forty-second to the fiftieth vertebra. 
Between the grooves minute wrinkles are abundant, mostly undu- 
lating, sometimes straight, not always confined to the space between 
two grooves, but sometimes crossing them without changing their 
course; they are inclined at all angles to the grooves, but are 
chiefly transverse to them. 

What the precise nature of this film may be is by no means 
clear. From its distribution one might infer that it originally 
formed a part of the integumentary investment. It closely resem- 
bles in character the structure which has been regarded by Mr. 
Moore as forming a part of the integument of Ichthyosawrus, and 
which this acute observer has compared to the wrinkled surface of 
the skin of the Porpoise*. The resemblance between this surface, 
as seen in our museum-specimens, and that of the investing film 
in Plesiosaurus is, indeed, great; and if such a skin were capable of 
fossilization, one might fairly allow that Plesiosawrus had been 
invested with it. It is very certain that the film in our fossil 
specimen was of a yielding flexible nature, or it could not have so 
neatly covered the exterior of the ribs and adapted itself to the 
ends of the transverse processes and the angles between the neural 
spines and the vertebral bodies as it has done. 


* Som. Archzol. & Nat. Hist. Soc. Proc. 186-566, p. 179. 


468 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


Classification. 


The pectoral arch in its essential characters is truly Plesiosaurian, 
though it differs from most Plesiosaurs in the fact, if it be a fact, 
that the coracoids do not extend in front so as to overlap the 
“lapels ” of the furculum. 

To make plain its relations to other members of the genus Plesio- 
saurus, I have constructed the following Table, in the last four 
columns of which the distribution of the vertebrae amongst the 
various regions of the spine is given for each species. 

In comparing the length of the head with that of the neck (or 
rather of the cervical series of vertebra) I have uniformly made use 
of the proportion :— 


length of heaqg_ I 
length of neck 100° 


The value found for I may be conveniently called the cervico- 
cephalic index. This index is given, for each species in which it has 
been determined, in the first column of the Table. 

In the second column I have similarly compared the length of 
the head and the dorsal series of vertebree (excluding sacral vertebree). 
The indices of this column are dorso-cephalic. 

The total length has not been made use of in comparison, since it 
is seldom possible certainly to obtain it, and variations in the length 
of the caudal region are of secondary value; but in the third column 
values are given for the length of the head compared with the cer- 
vico-dorsal series (exclusive of sacral vertebre). ‘To the cervico- 
dorsal cephalic index great value may be attached, since it can 
frequently be obtained and is independent of any error in the method 
or the practice of determining the position of the last cervical ver- 
tebra. It is true it may be affected by a mistaken determination of 
the first sacral vertebra; the sacral vertebrae, however, are more 
easily determined than the last cervical; and an error with regard 
to them will not cause a deficiency or excess of more than one or 
two vertebre to the cervico-dorsal series, while I am convinced 
much larger errors have been made with reference to the number 
of vertebree in the cervical series ; and, finally, in comparatively so 
great a length as that of the cervico-dorsal series one or two vertebree 
more or less will have but a trifling effect on the value of the index 
obtained fromthem. Another index of some interest is the cervico- 
dorsal, obtained by referring the length of the neck to that of the 
trunk taken as 100, 

The values now given for the various indices will certainly in 
many cases need revision, since the measurements on which they 
are founded are often extremely unsatisfactory. Sometimes they are 
unreliable or erroneous, sometimes vague (as when the length of the 
‘“‘trunk” is given without stating whether the sacral vertebrae are 
included or not, or when the length of one part is given in fractions 


FROM THE LOWER LIAS OF CHARMOUTH. 469 


of another*; and sometimes, finally, they are not comparable, as 
when in one place the length of the skull is given (as it too frequently 
is) as the length of the lower jaw, while in another it is taken as 
the length between the end of the snout and the basioccipital. 

On reference to the Table it will be seen that P. Conybeare has the 
same number of cervical vertebree as P: homalospondylus, but one 
less dorsal: its head, however, is much larger than that of the 
latter species; and hence there is a great difference in their re- 
spective cephalic indexes. In proportion of head to neck it agrees 
exactly with P. Hiheridgu, but differs widely in the number of cer- 
vical vertebre. It has also two dorsal vertebre fewer ; and its dorsal 
and cervico-dorsal cephalic indexes are distinctly different. Its 
cervico-dorsal index approaches nearest to that of P. Hawkins, nm 
which species this index attains its maximum value. 


; Cephalic indexes. Number of vertebree. Gadel odl ae ; 
2 | - umero- 
_ Plesiosaurus. a x oe dorsal | racial 
i O- Tso- in ° ina@ex 
ceuhalte. éeotalie. ome Cerv. | Dors. | Sac. Cau. ox 
ters 876 58°3 33°9 at Mee ae a 722 
BEGAGTIS....0c.c000. 63:9 Eyles 28:4 24 24 2 34 is SY a 
ropinguus ...... 60:0 Hore 28°85 |} 25 23 2 34. 92°6| 45°8 
Dramptoni......... 55-5 416 24:0 27 30 2 32 750| 28°5 
megacephalus 53°3 | 493 | 26°5 30 26 2 34 92:6; 33:5 
macrocephalus ...| 51:2 65°6 28°8 29 20 2 1280} = oO 
brachycephalus...| 35:3 | 403 | 188 31 24 2 21 | 1140; 333 
Ongirostris ...... 338 29-7 158 33 25 32 (2 We SS 
Hawkinsii ......... 30°0 46:9 18:3 31 23 2 TaG2 fo4s7 
Etheridgii ......... 24°3 26:0 125 30 23 2 34 106:0 
ponybeari ......... 24:1 37°4 14:6 38 21 2 5+) 153:0| 37:0 
lolichodeirus...... eT 23°6 10-1 4] 21 2 380+] 133°3|) 42:9 
macropterus ...... 12°8 17:0 UD 39 24 1? 28 1320} 50:0 
homalospondylus} 106 | 11:1 o4 38 22 2 105°0| 46-0 


There is a fine specimen of Plestosaurus in the British Museum, 
with the MS. name P. laticeps, Owen, which bears a close resem- 
blance to P. Conybeari. Its pectoral and pelvic girdles are well ex- 
posed on their ventral surface, and are strikingly similar in general 
form and arrangement to those parts in P. Conybeari; the dimensions 
of their component bones also show a general agreement, as will be 
seen from the following table. In each the humerus is longer than 
the femur; but these bones are each an inch shorter in P. laticeps 
than in P. Conybeari. 


* Thus, in his Monograph on Liassic Reptiles, Professor Owen says of 
P. rostratus that the skull is # the length of the neck. Now the skullis 1 foot 11 
inches long; and thus the length of the neck should be 2 feet 6 inches 8 lines. 
But he also says that the length of the neck is rather less than 4 the length of the 
spinal column; as the latter is 9 feet 9 inches long, the neck should be “‘rather less” 
than 3 feet 3 inches long. Whereabouts between these two quantities is the 
exact length? And would it not seem to have been easier to direcily state it? 
Unfortunately the instance here given does not stand alone. 


Q.J.G.8. No. 147. 21 


470 PROF. W. J. SOLLAS ON A NEW SPECIES OF PLESIOSAURUS 


Table of Measurements of corresponding Parts in P. laticeps, Owen, 
M.S., and P. Conybeari. 
P. laticeps. P, Conybeari. 


inches, inches 
Puneuluu’ leneth (aspscias) cesses steers eee sees 60 6-4 
@oracoid) tength (Gnaximiura) i ee5 ees eee eee 17-0 17-375 
is breadth: siajohiadscstenmecen. a costee merce eeee 80 8:0 
Pubis, length of symphysial margin (a.p. dia.) ......... 6°5 55 
< » parallel to outer margin (a.p. dia.) ...... 8:0 81 
is, » Oblique diameter from inner anterior to 
postero-exterion angles way see cee ase ee eee nee eee eee 10:0 90° 
Ischium, length |(@max.voblidias)im---sse- eee casera 9-0 10-1 
Humierus, length. £iirccsas. bec ecdoce cee cena nce ce eee 13°5 14:5 
Hémur; length: sa2.e. sae. tarenteate es cota eenece ee emcee 12:5 13°5 
Neck, from first cervical vertebra to anterior edge of 
furculumes..Uiten accaee sere. sane ee eee oem 56-0 780 
Trunk, from anterior edge of furculum to posterior 
edgesor pubis’ ast-ccusnnetecese eee cece rere 48:0 53°90 
Number of cervical vertebre from first to anterior edge 
Of furculumy Sincere casos et aaesas rae eee eres 27 38 
ength of posterior cervical vertebra ne-eea-eeeeeeeeeee 2°25 25 
Lengthrot dorselivertebra i. s-ss eee ee eee eee 2'375 2°65 


The anterior end of the head is brcken off and missing in P. lati- 
ceps ; so that the cephalic indexes cannot be determined. 

The regions of its vertebral column cannot be clearly defined, 
since the pectoral and pelvic girdles conceal to some extent the ver- 
tebree beneath them. There is no great difference in the length of 
the dorsal region of the two species ; as shown in the Table, in P. 
Conybeari it is 5 inches longer than in P. latieeps, a difference which 
may be accounted for by supposing the furcula of the former to be 
displaced a little forwards. 

The widest departure is seen in the neck, its length being much 
less and the number of its vertebree much fewer in P. laticeps; but 
there is an artificial look about the neck of this specimen which leads 
me to conjecture that some of the cervical vertebrae may be missing, 
so that, if those which remain were arranged as they were first 
found, several considerable lacunz would appear between them. 

The lengths of the centra of the cervical and dorsal vertebree 
make a close approximation in the two species, P. daticeps in this, as 
in several other characters being a little the smaller. The anterior 
cervical vertebra of P. laticeps are rugose or tubercular in the same 
fashion as those of P. Conybeari. 

Finally, both species come from the same “ gisement,” the Lower 
Lias of Charmouth. 

If my conjecture with regard to the identity of the two species 
should prove correct (and it will require a closer examination of 
Owen’s species than I have been able to give it to decide this), then 
of course the name “ latzceps ” will have to be suppressed. It occurs 
in print in the ‘Geological Magazine,’ vol. iv. p. 144, but without 
accompanying diagnosis or specific description. 

One other character of P. Conybeari alone remains for comparison ; 
and that is the relative dimensions of its vertebral centra. If we 
find the proportion of the breadth and height of a centrum to its 


FROM THE LOWER LIAS OF CHARMOUTH. 471 


length taken as 100, we shall obtain its latitudinal and altitudinal 
indices. These are given in the following Table for the cervical 
yertebree of a number of species. To make their comparison of 
yalue, corresponding vertebre should be selected for each species ; 
and for this tables like that on p. 452 would have to be constructed 
and discussed. As it is, I have had to make the best use I could of 
the material ready to hand in published papers, and to trust to the 
chance of different describers having given measurements of an 
average cervical vertebra. Most of the indices in the table have 
been derived from the thirteenth to the fifteenth vertebra, and 
probably are sufficiently comparable. 


Plesiosaurus Hea ees Number of vertebra. 
Wonywedey - 20.2. .62-.-5-- 70 100 XV. 
homalospondylus ......... 92 72 XIII & XIv. 

SUP USS apace ene 107 100 

ecelospondylus ............ 110 106 XV, 

(TIGRIS Caicos anceaeee enone 112 84 XV. 

BRP ACEOUNGUS soc occ edecccces 112 100 Middle. 

: Average of middle and 
infraplanus .............4. 113 101 ve a Pee ee 
dolichodeirus............... 113 94 

PRCWADIISTO cose cc access ese 116 116 

Sather 120 98 

SLOUL PAUSE ..2...:- +++ 123 115 

a WGINGID oc ch-os-c0000--5- 125 112 

TeLICING. ache seee eee 127 102 

Glenutheraxonm ............25. 127 100 

CORR 134 119 

SUDEFIZONUS | ...5....5.-004- 140 109 

GMOMPCTISNS .c20-- cts cciceccnse 140 116 

BACIVONAUS. 2.50.20 5000000 0: 42 110 

SULOCUIN Scenes 150 120 

eleutheraxon ............... 169 100 

Li 170 155 XV. 


It will be seen that the centrum of the fifteenth vertebra of P. 
Conybeari has a lower latitudinal index than any other known spe- 
cies, while its height remains about the average. 

P. homalospondyius, which makes the nearest approach to it, is 
equally remarkable for its exceptionally low altitudinal index. 

The compression which the cervical vertebrae of P. Conybear have 
undergone may to some extent account for the narrowness of their 
centra; but it cannot be altogether explained in this way, since it 
persists in a marked manner down to the twenty-eighth vertebra. 
Moreover our measurements were taken from the articular ends of 
the centra; and these show no obvious signs of compression. 

The Table shows a general tendency in the long-necked species 
towards a low latitudinal index, and in short-necked species towards 
a high one; but the rule is subject to exceptions, and we have not 
enough instances to reason from. The latitudinal has clearly a 


greater range of variation than the altitudinal index. 
212 


AT2 PROF. W. J. SOLLAS ON PLESIOIAURUS 


Notes on Plesiosaurus megacephalus, Stutchbury, and 
P. brachycephalus, Owen. 


The type specimens of these two species are preserved in the 
Bristol Museum ; so that I have had a good opportunity of making 
a close acquaintance with them, and have succeeded in elucidating 
some points in their anatomy which were hitherto obscure. 


PLESIOSAURUS MEGACEPHALUS. 


1. The Roof of the Mouth.—The skull of this specimen lies on its 
dorsal surface, separated from its matrix from the snout to a trans- 
verse fracture which traverses it across the orbits. The matrix 
has been carefully chiselled away from between the rami of the 
lower jaw, so as to clearly expose the roof of the mouth and the 
base of the skull for its entire length. The anterior part of the. 
base (fig. 8), which lies in front of the fracture before mentioned, is 
by a most lucky chance much better-preserved than that behind, and 
thus affords us an opportunity which has long been desired of ascer- 
taining more exactly the true nature of this part of the skull. 


Fig. 8.— Ventral View of the anterior Part of the Skull of 
| P. megacephalus. (Scale 3.) 


2, internal naris; f, palatal foramen. 


It presents, about 2 inches behind the end of the mandibular 
symphysis, two oval foramina (fig. 8, 7) longer than broad (1°45 inch 
in length) separated from each other by a bone which extends back- 


MEGACEPHALUS AND P. BRACHYCEPHALUS, 473 


wards in the middle line and unites by a splintery suture with the 
palatines behind. Itshows traces of a straight sutural union along its 
median antero-posterior diameter, and consists, without doubt, of the 
connate vomers. On their outer margin the foramina are bounded 
(fig. 9) for the anterior three quarters of their extent by the maxille, 
for the remaining quarter and along their posterior margin by the 
palatines, and along the inner border (as before mentioned) by the 
concave outer margin of the conjoined yomers. The vomero-palatine 


Fig. 9.—Diagram showing the left Internal Naris of P. megacephalus, 
bounded by the Maxilla, Vomers, and Palatine. (Scale 3.) 


Vo a P I} 


suture has more or less the shape of a W, as exposed on the floor 
of the skull, the apex of the W being on the median line, and 
the end of its lateral strokes cutting the inner posterior angles of the 
foramina. 

The palatines extend backwards for some distance as flat, hori- 
zontal plates, suturally united in the middle line, and completely 
roofing over the front of the mouth. Their sutural union is inter- 
rupted for a part of its course by an elongated vacant space (fig. 8, f) 
which clearly corresponds to the palatal foramen of many Lizards 
(e. g. Iguana, in which it is well seen). 

An oblique linear fissure starting from the middle of the outer 
stroke of the W of the vomero-palatine suture runs on each side out- 
wards and backwards, to disappear against the matrix bounding the 
roof of the mouth. These fissures might easily be mistaken for 
sutures, in which case they would be regarded as indicating the line of 
junction of the palatines; they are, however, simply fractures 
which have broken the palatines along a line where they become 
flanged upwards and outwards to join the maxilla. The fractures 
are, indeed, continued through the skull; so that the middle part of 
it, included between them, can be readily separated from the outer 
part on each side, and the form of the palatines and their union _ 
with the maxille clearly exposed (fig. 10). 

Another fracture traverses the skull nearly vertically, but oblique 
to the axis, passing through one of the external nares, and one of the 
oval foramina, or nares, as we may venture to call them, before 
mentioned. This fracture shows (fig. 11) a large central chamber, 
now filled with the Lias limestone; it is bounded above by the 


474 PROF. W. J. SOLLAS ON PLESIOSAURUS 


Fig. 10.—Oblique Fracture through the Skull of P. megacephalus, 
showing the line of union of the maxilla and palatine. (Scale 3.) 


In, Internai nasal passage. 


premaxille, below by the vomers and palatines, and on each side by 
the maxille, and also in some parts of its course by the palatines. 


Fig. 11.—Transverse Fracture through the Skull of P. megacephalus, 
crossing the nasal chambers. (Scale 3.) 


tK[ppyo 
yyy 
SLLLLINCL 


EN, external naris; JN, internal naris. 


The vomers (fig. 11, Vo) form a trough-shaped bone of considerable 
thickness, flat below but concave above, with a low ridge on each side 
of the middle line ; its sides extend upwards, together with a process 
from the palatines, as a curved wall for half the height of the central 
chamber, which is thus divided through its lower half into a middle 
and two lateral portions. ‘The lateral portions have the appearance 
of tubes sloping downwards and forwards. On each side of the 
premaxille the external nares open freely into the central chamber ; 
while the foramina on each side of the vomers communicate with its 
lateral passages. These passages are bounded externally by the 
maxille, and internally, as well as inferiorly, by the palatines. 

The oval foramina appear to represent the internal nares, since 
they are similarly situated with respect to the surrounding bones as 
the posterior nares of many Lacertilia ; and it is with this order that 
Plesiosaurus stands in the closest connexion. 


MEGACEPHALUS AND P. BRACHYCEPHALUS. 475 


It must be borne in mind, however, though I doubt whether it is 
generally known, that the posterior nares by no means occupy 
a constant position in the Lacertilia; for though they are most 
usually bounded in the manner above stated, they sometimes shift 
their position backwards and open at the back of the palatines. In 
such cases, however, the palatines are produced towards the middle 
line, each aleng its cuter edge, into an underlying plate, which roofs 
ever the mouth and forms a floor to the nasal passages. A section 
across the Plesiosaurian skull might be expected, therefore, to give 
some signs of an inflection of the palatine bones, converting them 
into incomplete tubes, if such a backward extension of the nasal 
passage obtained init. No such signs, however, are to be detected 
in the specimens under consideration. A diagrammatic sketch of a 
fracture passing transversely through is given below (fig. 12); it 
shows plainly the outward and upward bend of the palatines, but 
not a trace of an infolding. 


Fig. 12.—Transverse Section across the Skull of P. megacephalus, 
showing the palatal plates flanged upwards and outwards towards 
the Maxille, but not inflected to form a nasal passage. 


e) 


' O a 
a eee 


It is a fact too curious to be passed over, however, that the 
internal are situated in advance of the external nares of this 
Plesiosaur, the anterior margin of the latter being a trifle under 
2°8 inches behind that of the former. 

A transverse section through the upper and lower jaws is given 
in fig. 13: the upward flange of the palatine is seen meeting the 
maxilla; and the lower jaw has the usual reptilian composition. 

2. Redetermination of the Number and Distribution of the Ver- 
tebre and of the Length of the Regions of the Spinal Column.—Of 
cervical vertebre, twenty-nine are visible up to the anterior edge 
of the furculum; in all probability one more lies beneath this 
bone; and the total number may therefore be taken as thirty. 
In consequence of the concealment of a large part of the spine 
beneath the pectoral and pelvic girdles, the number of dorsal 


476 PROF. W. J. SOLLAS ON PLESIOSAURUS 


Fig. 13.—Transverse Section through the Jaws of P. megacephalus. 
(Scale 2.) 


Ans 


T, tooth. 


vertebre can only be indirectly arrived at. The total length of 
that part of the vertebral column which lies between the anterior 
edge of the furculum and the posterior edge of the ischium is 
70 inches; from this 2°3 inches must be deducted on account 
of the concealed last cervical vertebra, and 6-9 inches for three post- 
dorsal vertebree supposed to be concealed beneath the ischium, two 
of them being sacral and one the first caudal. This leaves 60-8 
inches (70—2°3—6:9=60°8), which is the length of the dorsal 
region. Divided by 2°3, the average length of a dorsal vertebra, this 
gives 26°43, or, neglecting the fraction, 26, which is the number 
of vertebre in the dorsal series. These numbers are embodied 
in the annexed Table (page 477), in which the measurements of a 
number of specimens of different species are compared together. 


PLESIOSAURUS BRACHYCEPHALUS, Owen. (Plate XXIV. fig. 2.) 


This species has not been figured, and has been only partly described. 
I do not intend here to do more than offer a few observations upon it, 
and to correct the previously made measurements. 

1. The Skull.—Although incomplete and broken, the skull is but 
slightly distorted, and presents several points of interest. 

The snout is broken away from the rest of the skull, and shows 
the under surface perfectly. It is 5 inches long, and does not 
exhibit the internal nares ; so that they must have been situated 
further back. Its broken surface extends at right angles to its long 


477 


MEGACEPHALUS AND P. BRACHYCEPHALUS. 


. Total Sa- -, | Fore | Hind | Hu- | Fe- | Ra- 1 Ma- 
Species. length. Head. | Neck. |Trunk.| am. | Tl. | imp. | limb. |merus.| mur. | dius. | 1b! | Ulna- Fibula ois. | Pes: 
*megacephalus ...............| 195 | 30 56:3 | 60°83 | 46 | 54:5 | 33+ | ...... | 137 | 135 | 46 | 45 | 43 4'5 
“© (Ovpsyaay ONAN Sec oodonsaneORoHOK 268 | 40 72 965 — J) eamecnd 66 GOD A misc 21 22 6 6° 6'5 6 
tmacrocephalus .............| 59+] 105 | 20°5 | 16 15 | 11+ | 12.75} 18:5 4 Aaa) |) 12745) RE 1-16 |" Whe 8 9 
tbrachycephalus ......... ...| 129 | 145 | 41 36 25 | 23 Sac ae | acoreeae poet OxGOnltenee sae IPAS) sonar. to) 
IGM OIROSiMis enema cmenr iO meorrO le LOl 980) so dou (POO) 44 bay cal OC limanecnialfs 40 vehea iene 
INGLE aehol-Ibh eoeeeeedeoneE ssocn, Wee (eon | 20 16 ee 20 Ome |aaliss Wate eo aerfoy oxo) fA) AD) te aS a accom | ee) 
PAWPMEHOCWeneewaceee esc) 08) OO 128 26:5 | 2°5 | 26 pee ln wataenlic GROE at oh ALO 
PIC OnV MCAT G cae. eveecvencs| Lik | 20 83 53'5 | 4:25) 10+ | 445 |...... | 148 | 14:25) 6°45) ..... DD) Geen oO 
LOMICHOCOUOTISN cmevretstetteven|) wiles 85 | 48 36 Taoran: (od sepia See alee a 7 3 3 s-tgtetaehe| Meo eecenel pele 14 
IMACKOPLELUS ...-crecree ses] LOa 9 70 53 escent RAO 42°5 | 46:5 | 12 11 6 5 Sheer leedeeee” 25 
homalospondylus ........./198 | 9 |85 |81 |....../32 | 44 | 45 { ae her Pee lo) ag | avn 24 
HOSHRENTUS | jogpoceooqoonouneene |f ets jp ae 1 Gls) 39 Fou, Oo 24 Pay 18) O7D | oo. | so Bide ses |e Le 
FOO PUM CUUIS reece veins 180 30 | 50 54 ncosee | Or4 44 Teucey owe meteor leOlOs | teas ale WOO 


ZELIENOIOUS savassceseeesccce 22S 4) | 52 72 Boies: |e stam hac ere ae 


The measurements are given in inches. 
In those marked * the length of the head is taken from the anterior edge of the premaxilla to the posterior edge of the quadrate ; marked 
thus, t, it is taken as the length of the lower jaw ; thus, }, from anterior edge of the pramaxilla to end of parietal. 


478 PROF. W. J. SOLLAS ON PLESIOSAURUS 


axis, and since it has been polished reveals very clearly the arrange- 
ment of the bones in this region of the skull (fig. 14). 


Fig. 14.—Transverse Section through the Skull of P. brachycephaius. 
(Seale 2.) 


The upper and backwardly prolonged processes of the preemaxille 
are seen on each side of the middle line above, the maxille bearing 
teeth on each side, the vomers in the middle line below conjoined to 
form a single trough-shaped bone ; a plate of bone forming the roof 
of the mouth on each side of the vomers is an extension inwards of 
the maxille. 

The left orbit is complete and undistorted ; it is bounded by the | 
usual bones, the sutures between them being exceptionally plainly 
shown. An additional bone, which looks as though it had been 
segmented off from the jugal, appears, however, between the jugal 
and postorbital ; its surface is marked in the same way as the Jugal ; 
and it bears a nutritive foramen, which completes the ascending 
series of these openings carried by the jugal. The suture between 
the two bones is distinct, however; and the striations on their 
surface are so directed as to indicate their separate nature. If a 
posterior supraorbital bone had worked its way in between the Jugal 
and the postorbital, it would have the position here described; but 
simulation of the appearance of the jugal bone wouid remain unex- 
plained. Hence it seems best to call it a suprajugal. 

The jugal and the suprajugal bones are not the simple bony plates 
which they appear to be in a lateral view of the skull, and as the 
jugal really is in recent Lacertilia. Both extend inwards behind the 
orbit as a bony plate, which meets and joins externally with a 
similar expansion of the parietal. In this way the orbit is com- 
pletely walled in behind. The maxillasimilarly extends inwards in 
front, no doubt accompanied by the lacrymal; and, below, a con- 
tinuous floor is afforded by the expanded palatine. The orbit is 
consequently very thoroughly walled round. 

The right orbit is incomplete ; and a horizontal fracture enables us 
to remove the jugal, the only outer boundary bone of it remaining, 
from the floor of the skull below. The jugal thus removed is a 


MEGACEPHALUS AND P. BRACHYCEPHALUS. 479 


triradiate bone—when looked at from below, something like the 
letter 7. ‘The crossbar of the T corresponds to the outer plate of 
the jugal, the stem to the process which extends from it inwards. 
The inner plate or process of the jugal does not join the outer bone 
abruptly, but curves outwards on each side into it. Thus a trian- 
gular space of considerable size is left at the junction of the two 
parts, or, in other words, at the origin of the inner plate. This 
space is occupied by coarsely cancellous bone, and lies immediately 
under the nutritive foramina, which occur on the outside of the 
proper jugal bone. The meaning of these foramina is thus made 
clear. 

The floor exposed by the removal of the jugal is very difficult 
to interpret. Immediately behind the palatine is a narrow bone, 
transverse to the axis of the skull, and apparently joining the 
posterior edge of the palatine. Behind this, again, is a flat 
parallel-sided bar or lath of bone, projecting from the middle of the 
skull outwards at right angles to the axis; it 1s united by a splintery 
suture with the inner margin of a large and important bone, which 
extends backwards, prolonging the line of the maxilla towards the 
quadrate. It consists of a vertical wall-sided outer plate, roughened 
on the external surface (which is a flat plane), and an interior 
horizontal plate, the inner angle formed by the divergence of the two 


‘plates being neatly rounded into a concave curve. The inner margin 


of the horizontal plate is a deeply concave curve. 

The vertical plate of a bone having a similar position is shown in 
the right side of the ventral surface of a skull numbered 14550 in 
the British Museum. I do not know what to make of this bone; but 
it appears to be that which Prof. Huxley has spoken of as quadrato- 
jugal in his paper on P. Hitheridgu. 

The posterior part of the skull of P. brachycephalus covers over 
the axis and atlas vertebre. It consists of the parietal and the ends 
of the two bones which have been called suprasquamosal. They 
join in a splintery suture over the middle of the parietal, and appear 
likewise to underlap it below ; so that this bone appears to proceed 
from between the upper and lower tables of the suprasquamosal 
bones. ‘This is a very singular feature; but as I have been able to 
examine this posterior fragment of the skull on all sides, and partly 
to take it to pieces, I entertain little doubt as to its existence. 

2. Redeternunation of Measwrements.—Professor Owen says that 
the vertebre, at least as far as the 28th, are cervical; but a careful 
examination leads me to include the 29th as an indubitable cervical 
vertebra. The succeeding vertebre have lost a slice from their 
exposed sides; but there is good reason to conclude that the 30th 
and 31st also belong to the neck. ‘The length of the cervical series 
is, then, as follows :— 


I and IT concealed+ III to XXIX undoubted cervicals+ XXX and XXXI doubtful. 
1S5inch + 36 inches + 3°5 inches 


=A] inches or 3 feet 5 inches. 


480 PROF. W. J. SOLLAS ON PLESIOSAURUS 


The remaining measurements are given in the appended Table 
(p. 477). 

meee Horizon.—Associated with this specimen, which came 
from the Lias of Bitton, are some Ammonites and Rhynchonelle, 
which Mr. Whidborne regards as A. Conybeart and R. variabilis ; 
hence he refers it to the A.-Bucklandi zone. 

In concluding this paper, I have to offer my best thanks to 
Mr. Etheridge and Dr. Henry Woodward for the kind assistance 
they have given me in many ways; to Mr. Whidborne I am espe- 
cially indebted for much useful help, and particularly for his care 
and assiduity in drawing up the appended list of species, with their 
geological positions, many of which have been determined by his 
own personal examination of the original types. 


EXPLANATION OF PLATES XXIII. & XXIV. 


Puate XXIII. 


Plesiosaurus Conybeart. 


Fig. 1. Ventral aspect of the skeleton. One twelfth nat. size. 

2. Dorsal aspect, showing the vertebral column from the thirty-seventh 
(xxxvi1) to the sixty-third (ux) vertebre. About one eleventh 
nat. size. 

3. Diagrammatic restoration of the pectoral girdle. One eighth nat. size. 

4, Diagrammatic restoration of the pelvic girdle, the ilia not being repre- 
sented. One eighth nat. size. 


Pruate XXIV. 


Fig. 1. Skull of P. Conybeari, left side. About one fourth nat. size. 
2. Skeleton of P. brachycephalus, Owen. About one twelfth nat. size. 


Discussion. 


Prof. Srerzy said that without an inspection of the evidence 
he was not in a position to criticise this elaborate paper; but it 
gave evidence of painstaking research of no ordinary kind, and he 
congratulated the author upon what he had put forward. He 
thought, however, that a part of the information was not entirely 
new. Still the species, he fully believed, was a new one, as several 
of the characteristics are not found in any other described Plesio- 
saur. He had an impression that the palatal foramina described 
by Prof. Sollas were shown in the species described by Mr. Stutch- 
bury fourteen years ago; and he believed they had been excavated 
after that description had been drawn up. As to the Lacertilian 
affinities of Plesiosaurus, he was unable himself, so far as he had 
seen, to recognize any of importance. In some respects it had 
affinities with Ichthyosaurs, Dinosaurs, and Crocodiles; so he thought 
the Lacertilian affinities could not be pressed. Though he dif- 
fered in some details, he thought the paper, as a whole, was done 
extremely well. 


ee 


X ney 
% 
4 
44) 
i 
4 
} 
i 
\ 
t 
as 


APPENDIX. 


*hexngonalis 
*infraplanus 


sp. - 
top. 


*gamma 
}t§erandis 
[:Bvansi (See 
*tmacromeras 
pachydoirus 


teimplox (PA 


} tttrochanteriv 


| “ep. 


*{tintorruptus 


[Mansellii (Hulke) 
‘megadeirus (Secley) 
‘plicatus (Phil.), “allied to” 


solidus (PAil.) 


Prtosaunus (Owen)..... 
taequalis (Phil.) .. 
§brachydeiras (Owen 

Owen, MSS.) . 

Owen) 

Phil: 
*nitidus (PAil).. 
‘See 


*tportlandicus (Owen) 


‘Geontinuns (Otsen) .. 


Lower Dias. | 


Mippre Lis, 


*ellipsospondylus (Oicen’ 
Fatt 2 ) 


Phil.) 


Pontuanp Bens. 


“carinatus (Phi) 1871 | Portland Rock... 
*svinspitensis (Seeley) -s{ 1871 | Portland Limestone... 
Onetackovs. 
“Bornardi (Oiven) .. 1850 | Neocomian to Uppor Ohalk\ 
*constrictus ( Owen) 1850. | Chalk 
§Gardneri (Seeley) 1877 | Gaul 
*Platispinus (Owen), 1854 | Lower Greensand 
*pachyomus (Owen) 1839 | Upper Greensand 
*planus (Owen). 1864 | Grecnsand. 


neocomionsis (Campiche) .. 


ley) .. 


il 


storrodeirus (Seeley, MSS. 


1s (Owen). 


| Porrrrrononox (Owen) .... 


(Owen) 


Kimmoridgo Ola; 
Kimmeridge Cla 
Kimmeridge Clay 
Kimmeridge Cla} 
Kimmeridge Olay 
Kimmeridge Olay. 


Kimmeridge Clay ....-1++ 


Upper Greensand. 
Lower Neocominn 
Bottom of L. Greensand 
Chalk 


Kimmeridge Clay. 
Kimmeridge Olay 


Portland Limestone 
Oxford and Kimmeridgo| 
Olay and Coral Rog 
Kimmeridgo Qlay.. 
‘Oxford and Kim. Olay 


Shanklin Sand ... 


L. Groonsand and Obalk ... 
Upper Greensand, Lower| 
‘and Middle Obalk 


jptarsuntus (Oe) 1899 | Planorbis? .| Street, Bitton, Charlton , 
fbrachycephalos (Owen) 1939 | Turneri | Bitton; Bott... 
; 1865. | Planorbis? -} Street... F 

fede (Ss) 1881 | Obtusus ... 1] Obarmouth Restricted go 

{dolichodeirus (Conybeare)... 1824 | Planorbis .. 4 Wein Ply Bristol, Bitton Type of genus... 
li 1865 | Planorbis ? ,| Stroot... i 

talien ( ) 1868 | Ostrea-beds 

Tavki Oven 1838 Ostren-beds a q Lyme, Weston . Restricted genus 

t Meeephalus (Conyea 1836. | Planorbis ‘| Street, Lymo, Weston . 

reatromos (Quen) 1830 | Marl” Near Lymo 

paesphalus (tule 1846 | Ostrea-beds, Streot, Wilmeote, 

Pee tea (Oiven) 1865 | Bucklandi Oharmouth 

Srost ea 2 a sa | CHSC) 
Ae 1 lanorbis, Bucklandi ymo, Weston Erot 

frugosus (Otren) Obtusus : §Grauby (Leicestershire), } a ee 

saubtrigoous (Otcen) 1839 | Bucklandi? . Weston (near Bath).. 


Oorwloy 
Shot 
Kimmeridge Bay 
Ely. 


Shotoyer 


Shotover, Cumnor, Baldon 


-] Quainton (Bucks). 


.| Winspit (I. of Purbeck) .. 


Reach, Arundel; Moscow. 
Steyning (Kent) 
ROLE E 

‘| Tquaudon Quarry, 
) Rea do) 
:| Cambridge 

:| Cambridge. 
|| Ste. Croix; Moscow 
‘|. of Wight 
-| Burnham 


.| Near Swindon, Shotover . 
|| Market Raisin, Shotovor . 
‘| Weymouth, Shotover . 

‘| Shotover, Kimmeridgo, 


yn: 
vin 


.| Shotoyer 
.| Groat Grant 
I. of Portland 
Shotoyer 


‘| Ohristian Maliord, Kimmeridgo, 


RK 


.| Bedfordshire -. 


Maik 


Cambridge, Frome, &e.; Moscow) 


lay pils ... 
Stanford, 


over, 
Murienosaurus 
Colymbosauras 
Elipsospondylic group 


.| Colymbosaurus (Seeley 2)... 


Manisaurus ., 


Oambri 
Moscow 


sbury, near St. Neotts 
ndon, Shotover 


“Genus doubtful” 


Or “now genus" (Ow. Kim, 
‘oxcombe O. R.). 


dstono, Sussex; Moscow ... 


.| Owen, Brit, Ass. Rep. 1839, p. 82.. 


_| Hulko, Q. 3, GS; vol. xxvi. p. GLI 
‘| Phillips, Valley o 


| Seeloy, Q. J. G. 8. vol. xxxii 


Owen, Brit. Ass. Rep. 1859, p.75. 
Owen, Brit. Ass. Rap. 1839, p. 69, 


Seeley, Ann. & Mag. N. H. eer. 3, vol. xvi. p. 352 
Sollay} described Meco peer 


Oren, Brit. Ass. Rep. 1859, p. 7 


Capricornis . Saltburn Tato & Blake, Yorkshire Lins, p, 252, 
Tamesoni |) Huntelit ate & Blake, Yorksbire Lins, p, 252. 
Urren Dias, 
*brachycophalus (Owen)... con 3 : , 2 York (Owe 
iGrampin (Car § Baily) 863 | Upper part of Bifrons......] Kettloness (near Whithy) Carto &Thily, Dubi, Roy. Soa. Journ, peu 
jlospandylus (sce) 1865) }| Communi =| puwntby, Owen, Pal. Soc., Linssie Rep. p. 12 Whitby 
faubin (Lite f Blake) 1876 | Communi | 5 .| Dato & Blake, Yorkshire Lias, p. 24 
thomalospondslos (Otcen) 1865 | Bifrons Whitby . | Owen, Pal. Soe., Linssio Rop. p. 1 ..]21, 80, 81, 33. ...| British, York, “Whitby . 
Varma (ate & Blake) Whitby . “| mato & Blako, Yorkshire Lite». 250, apes a 
et Lofthiouse Eretmosaurus 7? (T: § B.).| Sceloy, Ann, & Mag. N. H, ser. 3, vol. xv. p, 49 ......, 31. _} Woodvwardian 
ae Whith: F . "| Tato & Blake, Yorkshire Lias, p. 24 }i9 :| Whitby 
i s Eretmosaurus a| ri 
4 | Lofthouse ‘ Rep, Yorksh, Phil, Soc. 1851 | York, 
Lower Oouite. il 0 19 
*qsorrations (PAilli Stonesficld Slate; Stonesfield Phillips, Valloy of Thame: iy 
Tee Cee. Great Oolite . Minchinhampion Secley, Cat. of Rept. & Woodwardian. 
Minpix: Oourre. 
teurymorus (Phil. 1871 | Oxford Clay . Bedford ... % a Woodwardi 
Sectsii gabe, ) 1874 | Lower Oxfurd Olay Huntingdonshire Muranosaurus (Slcy) Coll Eacds 
osontensis (Phd) 1871 | Oxford & Ampthill Clay ...! Oxford, Long Marstoi Murmnosaurus 
philarchus (Seley, 1869 | Oxford Clay Peterborough eee 
*plicatus (PAIL) 1871 | Oxford Clay . ; * Bllipsospondylic group” 
Knnrenainor Cray. 
taffinia (Otven) « Kimmeridge Olay. Heddington, Shotovor Owon, B. A. Rop. 1839, p. 86 || Enniskillen, i 
*brachistospondylus (Auk) Kimmeridgo Olay . Kimmeridge Bay ... Hulke, Q. J. @, 8. vol. xvi, p. 611 
*brachyspoudylus (Otoen) .. Kimmeridgo Clay. Hedidington, Shotover, near Orvon, B, A. Rop, 1839, p. 78 || Oxford, Bristol, Woodwardian 
Weymouth, Foxmouth 
fdmdicomus (Oven), Kimmeridge Olay, Shotovor. Owen, B.A. Rep. 1899, p. 82. 


Phillips, Valley of Thames, p. 37: 
Phillips, Valley of Thames, p. 375. 
Phillips, Valley of Thames, p. 375. 


ley, Cat of Rept. & Birds, p. 97 
‘Thames, pp. 873, 374. 


Phillips, Valley of Thames, p. 370 


Phillips, Valley of Thames, p. 374..... 


Seoley, Ann. & Mag. Nat. Hist. cer. 4, vol. viii. p. 181 


Phillips, Valley of Tham 
Owen, Odontozraphy, p. 
Phillips, Valley of ‘Thames, p. 359, 
Owen, B. A. Rep. 1839, p. 83. 
Phillips, Valloy of Thames, piste 
Phillips, Valley of Thames, p. 360 
Secloy, Ont. Rept. & Birds, p. 116 
Owen, Pal. Soc,, Kim. Clay Rept. pt. 3, p.8 
Phillips, Valley of Thames, p. £66", 


Scoloy, Cut, RopseBirds, p_ 98 
Owen, B.A, Rep. 1839, p85. 


Seeley, Ann, & Mag. Nat. Hist. ser. 9, vol. xviii. p. 112) 


Owen, Odontogeaphy, vol. ii. p. 19. 
Owen, Odontogrephy, yol. ii. p. 19 


10, 


Dixon, Foss. Sussox, p. 396. .} 10,11, 12,15...) British. 

Dixon, Foss. Sussex, p. 398. -} 11, 12. 

Seeley, Q. J. G. S. vol. xxxiii. p. 541 liek oll. Gardner... 
Owen, Ont. Foss, in Ooll. Surgeons, p. 6 ll. .| British. 

Owon, B. A. Rep. 1839, p. 74... 41, 1 .| Woodwardian. 

Owen, Pal. Soo., Cret Rop. Sup. 15, | Woodwardian, British, 
Pictot & Campiche, Foss. Orét. de Ste. Oroix, ser. 2,| 11, 15. .| British, Woodwardian 


EVAR riored 


10, 11, 12, 15 


wardinn, 


..| Bristol (8 specimens) . 


Woodwardian. 
Bristol, British 


..| British (8 specimens) .... 


Woodwardian. 
Jermyn Street. 
British 
Enniskillen. 

Coll. “ Miss Phillpots” .. 
Bristol, Warwick, 
British .., 


§British, 
(York?) 


*Enniskillen, Bristol, 


*Bristol. 


Oxford, 


.| British & Coll. Mansell. 
.| Woodwardian. 
Oxford ..., 


British, *Bristol, Enniskillen, Wood- 
.| With 2A 


,| Syn. triatarsostinua (Haykins). 
-| Quoted by Stutchbury as" macrourus.” 


|] Tn beds with Anunonites heterophyllus, 


.| Near Pl. dadicomus. 
| With 


..| Seo Oxford Oloy. It is doubtful whethor Phillips 


To face page 480. 
TABULAR SYNOPSIS OF THE GEOGRAPHICAL DISTRIBUTION OF THE PLESIOSAURS. BY G. F. WHIDBORNE, Esa, F.G.S. 
. Tanre I.—English Species. 
*=Vertebra. t=Bones of tho limbs, thorax, or pelvis. }=Skull. §=Skeleton, [—=Imperfoct skeleton. §=Toeth. 
Date. Horizo Locality. Defined genus. Description. References. Museums, 4 
Presiosavnvs (Conybeare) « 1821. 
Ruetic. 

etttas (Oman 1839 Owon, Brit. Asso. Rep. 1839, p. 80 . “Bristol. 

*Hawkinsii (O1e" 5 ar Sco " Lowor Lins."” 
rugosus (Otoen) « Eretmosaurus *Bristol ‘Seo " Lower Lins." 
trigonus (Cuvier) ; 


Figured in Geol. Trans. ser. 2, yol. v, p. 44 “Not 

rom Whitby or Wurtemburg," Owen. 

nchonella variabilis (Schloth.) and Ammo- 

nites cbliquecostatus (Zict.) = A, semicostatus (1. & B,). 
Soo Wright, Q. J.G-S. vol. xvi. p. 406, 

Found with Ammonites planicosfafus, Sow. Min, Con. 
148, ono of the most frequent Ammonites of the 
Marston stone, as well as the Charmouth beds of 
this zone, 

\Bigured in Bridgwator Treatise.” 


Tho most perfectly preserved specimen known.” 


“Whitby,” Owen & Phillips, not Tato & Blake, 
The Granby skeleton found with 4, stellaris, 


| 
thy,” Owen, not Tato & Blake, Soo Lowor Lins. 
. below Dogger, with numerous Leda ctu.” | 
porhaps belong’ to P, longirosris” (‘I. & B,). 
“This is a synonym for P. macropferus, if tho latter is 

an Kremosaurus.” 


“Whit 
“60 


=P. grandipennis (MSS,). 
Founded on a specimen labelled " macrocephalus.” 
Sco Lower Lias, 


A small species, 


Am. Duncani, Lamberti, &e. Itis tho sano spe- 
cimen ag the“ Plesiosaurus grandis?" of Phillips, 


n 
Valley of Thames, p. S18. 


“Syn. Pl. recentior? (Qonyb. & You Meyer), and Pl, 
gigantews? (Gonyb. 1824),” Owen, 


makes (wo ord single species here, 


Small yortobran, probably but not cortainly from Port: 

Jand Rook, Tt Ouyier's nawo stands, this should bes 
como Pl, Phillipri. 
“Tt approaches H, megadaters! Sooloy. 


Coll. Leo. 
Coll, Swith. 


Oxford. 

..| Oxford, Woodwardian... 
Oxford. 
Oxford, Bri 
Woodwardian 
Oxford. 


Woodwardian. 
British. 
Oxford. 


Woodwardian. 
Enniskillen, Oxford, British. 


..| British. 
11, 12, 15 ¢..) Woodwardian, 


«| ‘Tho largest long-necked Plesiosaur known. 


Syn. "Pl, mamillatus (Owen). 


“Found with Am. vertebratis, whioh, howovor, occurs in| 
Uppor Oxford Olay at Cowloy,” Phil, 


Fouud with Arm, Duncani, Lamberti, &o. 


Tanre I.—Foreign Species. 


Pestosavnus. 


jonensis (Zen 
Nordmanni ( 


Bp. -. 


indicus (Lydekker) . 


i 
B 


pentagonus (Cue). 
profundus (Zonk 
suoyicus (Quon.) 


Norr.—Hawkins, in (6), 


Stercosurw 
complete li 


Omitting the most uncertain species, Table I gives 23 Rhuctio and Lisssie, 4 Oolitic, and 9 Oretaccous—making a total of 63 English species hitherto named or described. ‘There is, however, 


1836, 


2) from 


S24. 
182: 


1838, 
1839, 
1540. 
1s41, 
1846, Stutchbury, Q. J. G. S,, vol 
1848. 
1851. 
1850, Owen, in Dixo 


Exasuosxonvs. 
; : Owen, Brit. As. Rep. 1801, p. 122 -... 26 Ta : : Seoloy, Q. J. G.8. vol. xxii. p. 5Al. 
1525. | Oolite: Gur. Os. Foss. vol: p. 496, 9. “Provisionally veers (Cone) Niobsars, +} Cop, Bull. 0.8, Sury.vol iit no. 9, p.600, 
| Bay. of Concepcion Blake, Geol Mag 1803.” named,” Ouv, 
’ Waipara Bods (Australia) | Owen, Geol. Mag. vol, vii. p.49. Ma ; 
isin Neen » | Bich: Bull, de Moscow, 1846 Trasali (lector) Orelacoous,.....s.| New Zealand. rma, NZ, Inet, vol, vis p40) sven (99, 
as Owen, Geol, Mog. sol. vii. p.0, [p-30] 
eae Bydckker, Rep. Geol, Surv. Ind. vol. x] Puroasvnce- a ‘ 
if E Zauk. Beitr, rw. p. 65 Dedsbyspondslus (Oven), Wo |lerieecary .| DBichwald, Lethean Rossies, p, 1260 | Probably anerror’? 
1859? | Gris Neooom. Orimea D'Eichwald, Lethwa Rossi = % |) 15 
isp Honfleur, Cayier, Os. Foss. vol Worrinshit (Fisch). val 1616. | Neocomfan +o] Kiach. Bull, de Mfoscou,vol.i. pp. 11, 105) 10, 
Zenk. Beitr. Urw. 64, t. 6 
a i Quonstedt, Der Jura, pp. R 4 P 
1828 | Hample Zone | Hillisusen: Quenstedt) Dor Jura, t. 58. f. 3. NgSTE 1867 | Neocomian Fisch, Bull. do Mosoou, vol. i. p. 902.) 16, 
: 3 regoing Lis 16. 1861. Owen, Brit. Assn. Rep. p. 129, 9. 1876. Phillips, Geology of Yorkshire, od. 2. 
Books and PADIS ee a | 17. 1561-69. Owen, Bal. Soc, Kin, Clay Rept BI. 1875. Tate and Blake, Geology of Yorkshiro. . 
Conyb. Geol. Trans. ser. 2, vol. i. p- 103. On Pi. dolichodeirus. 18. 1861-70. Owen, Pal. Soe., Linssio Rept, R BE SF. Soy) Q UG: 8 vo, pH, on Muh Gardner p 710,08 
Her, Os. Rosa. vol 19, 1866. Carte and Baily, Dublin Roy, Soc. Lip, p.160. On Pl Oramp- rl, Beatie . 
Perisena Berdgowater ‘Treatise. On Pl. macrocephalus &o. toni. vs CE SeNE ROMS eae 83, 1878. Wright, Lias Ammonites. 


Owen, Geol. Trans. eer. 2, yol. v. 
Owen, Brit. Association Rep. p- 
Hawkins, Great Sea-Dragons. 

Owen, Brit, Assn. Rep. p. 60. 


Bronn, Index Pal. 
Owen, Brit \- Rep. 
Foss. of Sussex. 


1858. Quenstedt, der Jura. Phe am 
1859. D'Eichwald, Lethaa Rossics, vol. ii, sect 2 


has two fino figures; but his names being generic and o 
© Rhatic, Placodus (coll. Moore) and Tanys*ropheus (coll. 


16. On Pl. macrocephalus. 


i. p. 411. On Ph megacephatus, 


a 
Viant). "Owes’s ‘Walzontology’ and Von Meyer's Palasontographi 


1864. Geol, Mng. p. 47. 


Sceley, An. Sf. N. H. 
. xvi, p. 352. On Pl, 
Geol. Mag, 


7. Moore, Q- 


Mansellix. 


on the PI 


rently merged by Oren in his Report (1890 and 1841), it 


rp. 144. 
FG.S, vol. xii p. 448, Abnormal Secondary Deposits. 
). Hulke, Q J. G.S, vol. xxvi. p. O11. On Pl. brachistospondylus aud 


Seeley, Woodwardian Ostalogue of Rept. and Birds. 

). Owen, Geol Mag. vol. vii. p.49. On 
Phillips, Valley of Thames. 

Seeley, An. MLN. H. 

or QI.G.S vol. xxx. p. 197, on Murenosaurus 

lesicsaurian Pectoral Arch. 


49 & 992. On Pl, macropterus. 


sor. 8, vol. xv, 
Phi. cliduchs. 


eleutherazon & 


farians from New Zealand, 


ser. 4, vol. viii, On Pl. winspitencit. e 
Toe ee uras Leedsit; p40, 


has not 


ica” give many other foreign species and genera from 


been poreible to refer to them in the tables. Prof. Seeley, 


Notices of Plosiosauri alsa occur in the following works. 
1821, Do la Becho and Conybeare on Plesiosaurus, Cool. Trans. sor. 1, vol. vs p. O11. 


1837. Pusch, Polens Paliontologie (?). 
1546-47. ns alas do Moscou. 
1850, Owen, Odontography. . 
Gay, Historia iia y politics de Chile, 
1859. Pictet et Campiche, 
1858. Lucas Barrett, An. & Mo 
180. Owen, Palaontolozy. 
1874. Hector, New-Zealand Inst. vol. vi. 


N. IL 1808, 


1876. Lydekker, Rep. Geol. Surv. Tndia, vol. x. 


in Q J.G.S, vol. xxxili. p. 442, notes the fol 
the European ‘Trias. 


‘here are ssid to be Done from that of America. Table II docs not aim 


Fossiles da torrain Orétacé des environs de Sto. Croix. 


p00. 


Greensand, 


, : bridge 
lowing. genera :—{1) from the Cambridge Gromer 


at present no reason to suppose that some of the names 8r® not synonyms, 


abana taney 


a ahd Khe Nth ce 
ie. aor Pert> aN 4 
KU Lis a hake Ries 7 Oe PE 
ae Fe 

is: F 

5 ie 


in) 


4 


hy 
i 
Fa 
; 
a 


MEGACEPHALUS AND P. BRACHYCEPHALUS, 481 


Prof. Sorzas expressed his sense of the generous way in which 
Prof. Seeley had spoken of his paper. It was impossible to write a 
paper without making use of material already published; but he 
thought that there was but little put forward as new which was 
not really so in his paper. He thought the Lacertilian affinities 
were extremely well marked in the skull of Plesiosaurus. His de- 
termination of the structure of the anterior part of the roof of the 
mouth in P. megacephalus confirmed him in this opinion. 


482 H. HICKS ON THE REMAINS OF PLANTS FROM THE 


37. On the Discovery of some Remains of Prants at the Base of the 
DENBIGHSHIRE Grits, near Corwen, Norra Wares. By Henry 
Hicks, Esq.,M.D.,F.G.8. Wirthan Appendix by R. Krueriper, 
Esq., F.R.S., Pres. Geol. Soc. (Read May 25, 1881.) 


[Puats XXV.]_ 


In August 1875, when searching for fossils in the Pen-y-Glog 
slate-quarry, which is situated about two miles to the east of Corwen, 
I noticed some carbonaceous-looking fragments and markings on the 
shales in association with the massive grit beds towards the top of 
the quarry, which I thought at the time might possibly be plant- 
remains. Last summer I had another favourable opportunity of 
examining these beds; and I was fortunate enough to discover 
undoubted plant-remains scattered very abundantly over their 
surfaces. I submitted these for further examination to Mr. 
Carruthers, of the British Museum, and had the satisfaction to 
find that he entirely confirmed my views as to their nature. He 
said they were undoubtedly “ angular fragments of plants,” but that 
the specimens were not in a sufficiently satisfactory condition to 
determine the actual plants to which they belonged. I decided, 
therefore, not to bring the matter before the Society until I had 
another opportunity of visiting the quarry and of endeavouring to 
procure more perfect specimens. This I was able to do lately; 
and the additional materials now found have proved to exhibit 
structures sufficiently well marked to enable a very clear identifica- 
tion of several distinct plants to be made out. The specimens have 
been generally examined by Mr. Carruthers, and some specially 
by Mr. Etheridge and Mr. Newton, of the Jermyn-Street Museum. 
Amongst them have been found numerous small spherical bodies 
identical in general appearance, and in internal structure, with the 
Pachytheca described by Sir J. D. Hooker, from the bone-bed at 
the top of the Ludlow series. These are supposed to be the remains 
of spore-cases of land plants belonging to the order Lycopodiacee ; 
other specimens are supposed by Mr. Carruthers to be the micro- 
spores, and others to be fragments of the stems of the same plants. 
Some of the specimens would indicate the presence also of plants 
belonging to the genus Psilophyton. These, though tolerably abun- 
dant, occur chiefly as carbonaceous markings, and show little evi- 
dence of structure. The combined results are sufficient to make 
it clear that we have here a terrestrial flora of a tolerably high 
order. The majority of the fragments, however, belong to a curious 
plant not hitherto found in Great Britain. It was first discovered 
by Sir W. Logan in the Devonian rocks, in the peninsula of Gaspé, 
Lower Canada, and described by Dr. Dawson, of Montreal, in the 
Quart. Journ. Geol. Soc. for 1859, under the name Prototaxites. 
Dr. Dawson described it as a land plant of large size, belonging 


Mmtern Bros, mp. 


Sees a 


ss 


Fem acetone BE 


C 
Oo. 


8. 


REMAIN, 


SO rere 
etary oe 


9° 


I 


SILURIAN PLAN 


X 200 
ile 


os 


sooner it 


Quart. Journ. Geol. Soc Vol XXXVI TP: 


ith. 


4A.S Foord 


ai 


bie a, y 


DENBIGHSHIRE GRITS, NEAR CORWEN, NORTH WALES. 483 


to the Conifer, but yet differing from any Conifer known to him 
“in the cylindrical form and loose aggregation of the wood cells, as 
seen in the cross section, in which particular it more nearly resem- 
bles the young succulent twigs of some modern Conifers than their 

mature wood.” He maintained, however, that it was an “ exogenous 
tree, with bark, rings of erowth, medullary rays, and well- developed 
though peculiar woody tissue ”*, 

Mr. Carruthers subsequently examined the same plant and re-de- 
scribed it in an elaborate paper in the Monthly Microscopical Journal 
for October 1872, and gave numerous reasons for excluding it not only 
from the Conifer, but from land plants altogether, and for placing 
it in preference among the Algze. In doing so, however, he said it 
was an “anomalous Alga, and, indeed, that with the materials 
known, it was not possible to correlate it with certainty with any 
known group of Alge.” The identity of our plant with the above 
mentioned, which was re-named by Mr. Carruthers Nematophycus, 
is placed beyond doubt by the following note kindly given me 
by Mr. Carruthers :— 


‘«‘ The slides prepared by Mr. Newton show clearly that his deter- 
mination of the fragments of charcoal and petrified remains of plants 
in the Silurian rocks which you have found, belong to the same type 
of plants as that discovered by Logan at Gaspé, in beds which he 
considered to be of Devonian age. This was described by Principal 
Dawson in the Quart. Journ. Geol. Soc. (vol. xy.) under the name Pro- 
totaxites Logani. I made a careful examination of specimens which 
I owed to the kindness of Dr. Dawson, and published the results of 
this examination in the Monthly Microscopical Journal, giving the 
reasons for placing it among cellular plants and naming it Nemato- 
phycus Logani. The specimens show very distinctly the larger 
tubes of Nematophycus, running generally in a subparallel direction, 
but passing in and out amongst each other. The walls are not in juxta- 
position, leaving free space all around them, which was occupied, as 
is shown in the better-preserved specimens from Gaspé, with a dense 
tissue of more delicate tubes of smaller dimensions. That your speci- 
mens belong tothe plant called Nematophycus | havenodoubt. The 
conditions under which they are found are very different from those 
described by Dr. Dawson. His specimens were large trunks, some- 
times perfectly silicified and preserving their most minute structures. 
Your specimens, consisting of small fragments, consequently supply 
no help to the further knowledge of this remarkable plant, unless 
the occurrence on the same slab, in tolerable abundance, of small 
round bodies having the same form and structure as those found in 
the Ludlow bone-bed, which were figured and described by Sir J. 
D. Hooker under the name Pachytheca, indicate some possible re- 
lationship. The specimens found by you are perhaps smaller than 
those from Ludlow. ‘They present no indication of attachment, and 
no evidence of their relation to Nematophycus, except their being 
found together, which is not always a good basis for structural re- 


* * American Naturalist,’ vol. v. p. 249. 


484 H. HICKS ON THE REMAINS OF PLANTS FROM THE 


lations in fossil plants. The minute bodies, aggregated together, 
which you have also shown me are, I believe, spores; and as they 
are united in threes, they agree with the forms of the microspores 
of Lycopodiacez, both recent and fossil, and testify to the existence 
of a dry-land flora. Perhaps some of the anthracite fragments may 
belong to the stems of the plants of which these are the reproductive 
organs. The ribbon-like carbonaceous impressions, with a slender 
axis, must have been also dry-land plants ; they remind me of the 
plants discovered and described by Principal Dawson, C.M.G., as 
Psilophyton. 

“W. CARRUTHERS.” | 


The specimens found, hitherto, of Nematophycus are all in a frag- 
mentary condition, the largest pieces being generally under 2 inches 
in length, and a little over half an inch in thickness. The natural 
outline, however, is frequently preserved ; and if the majority of the 
fragments are any guide to the natural size of the mature plant, it 
is evident it must have been small as compared with the Devo- 
nian one of Sir W. Logan, which attained to over a foot in diameter. 
That the plant must have been plentiful at this early period is clear 
from the very great abundance of the fragments in some of the beds ; 
sometimes so closely compressed together are they, that they form 
an actual carbonaceous seam from one to two inches in thickness. 

The microscopical characters of this plant, which are peculiarly 
interesting, will be fully referred to in the Appendix by Mr. 
Etheridge *. 

The discovery of Pachytheca and other spore-like bodies in con- 
siderable abundance in association with Nematophycus is curious, but, 
as remarked by Mr. Carruthers, is no direct evidence of their relation- 
ship. We know, moreover, from geological evidence, that the shore- 
line at the time could not have been very far distant, and therefore 
that it is quite possible there may be here a mixture of marine and 
dry-land plants. ‘The broken condition of the specimens also tends 
to show that none of them lived in the actual positions in which they 
are now found, but that they were brought here by some accidental 
cause, possibly along with a great amount of sediment, and as the 
result of river-floods, or of depression followed by rapid marine 
denudation. | 

The almost abrupt appearance at this horizon of massive beds 
of grits upon fine muddy deposits of considerable thickness, such 
as the slates immediately below, evidently tends to show that a 
physical change was then taking place in some neighbouring area ; 
but besides this there is nothing to indicate a physical break at this 
point. 


* Dr. Dawson, in his reply to Mr. Carruthers, ‘Monthly Microscopical 
Journal,’ 1873, still insists on his former diagnoses; and in a letter addressed 
to me, dated June 16th, 1881, in reference to the published abstract of this paper, 
says :—“ I have perfect confidence in my genera Prototaxites, Nematoxylon, and 
Celluloxylon, as representing primitive types of land plants; and I maintain my 
judgment as to these genera, and I believe it will be vindicated by future dis- 
coveries. 


DENBIGHSHIRE GRITS, NEAR CORWEN, NORTH WALES. 485 


In addition to the above-mentioned tolerably well-preserved plant- 
remains from the shales associated with the grits, there are un- 
doubted evidences of a still earlier and probably equally important 
flora in beds of slate at the base of the quarry. Instead, however, 
of the remains being preserved chiefly in the condition of mineral 
charcoal, as in the upper beds, they occur here mainly in the state 
of a very pure anthracite. At the same horizon some large nodules 
are seen ; and in the centre of these anthracite is also occasionally 
found, evidently forming the nucleus. From some of the specimens 
examined, I conclude there can be no doubt that these plants must 
have been of considerable size; and the amount of carbon left on 
some of the surfaces, apparently from a single fragment only, would 
tend to show, as suggested by Mr. Carruthers, that it must have been 
derived from vascular plants. There can be little doubt, therefore, 
that there is in the slates and nodules, even in the so-called Taran- 
non slates, very clear evidence of a terrestrial flora of considerable 
importance*. The anthracite, as now found, is usually broken into 
innumerable small fragments; but it is perfectly clear that this must 
be due to changes to which it has been subjected since it was de- 
posited—changes which also produced induration, cleavage, and frac- 
tures in the argillaceous sediments. ‘The fissures in the anthra- 
cite, and in the charcoal in the other beds, are generally filled by a 
fibrous mineral, which occurs here in some places in considerable 
abundance. Mr. T. Davis has kindly examined this, and says that 
it is a “ fibrous form of a hydrated magnesian silicate.” In other 
cases the fissures are filled with calcite. 

The difference in the conditions of fossilization in which the 
remains are now found at the two chief horizons may doubtless be to 
a great extent explained by taking into consideration the manner in 
which they were imbedded in the deposits. The thick grit-beds were 
evidently thrown down rapidly, and covered over the fragments before 
decomposition had taken place in them to any great extent. The 
fine muddy deposits which compose the slates were evidently thrown 
down much more slowly, and in a tolerably quiet sea ; therefore vege- 
table material resting on the bottom would have time to decompose 
almost completely before it would be sufficiently covered over by the 
deposits. ‘To a certain extent the same cause has allowed remains 
of vascular plants only to be preserved, as cellular ones would be too 
readily and too completely destroyed to show indications of their 
presence in such deposits. This is, I believe, the reason why re- 
mains of Algee are not more frequently found in these older rocks, 
and why impressions only are seen in most cases. 

The Pen-y-Glog quarry, where the specimens were obtained, has 


* Since this paper was read I have received from Mr. Phillips, the manager 
of Pen-y-Glog quarry, to whom I am indebted for much assistance, specimens 
which show clearly that plant-remains occur in the pale shales below the Den- 
bigh-grit series in the same state of preservation as in the beds at the top of 
the quarry. These shales also contain numerous calcareous nodules. 


Oey. G.0. No. 147. 2% 


486 H. HICKS ON THE REMAINS OF PLANTS FROM THE 


been already referred to in several papers read before this Society ; 
and its geological position is therefore well known. In the Survey 
maps and sections its horizon is given as immediately at the base 
of the Denbigh-grit and Flag series. It is shown to be separated 
from the Bala or Caradoc beds by only a very thin band of shales, which 
also contain plant-remains. In the published sections across this 
neighbourhood approximate thicknesses of between 4000 and 5000 feet 
of beds belonging to the Denbighshire-grit and Wenlock series are 
shown to occur in direct superposition to the beds in which the plants 
are found. None of the beds belonging to the Ludlow series are 
mentioned as occurring in these sections, the highest given being 
Wenlock : hence, if the former were deposited in this area, they have 
all been since removed by denudation. 

The geological horizon of the plant-yielding beds in relation to 
typical sections in other areas is rather difficult to define, since 
hitherto the Denbighshire-grit series has not been satisfactorily 
correlated with any other group. In the Survey sections the true 
Wenlock beds are made to come in at an horizon somewhere about 
2000 feet above these beds, and to occupy the remainder of the sec- 
tions. ‘The only beds below the plant-beds which have been actually 
correlated by their fossil contents with those in other areas are the 
Bala and Lower Llandovery beds; hence, at present, the evidence 
goes to prove that they must be older than the Wenlock, and newer 
than the Lower Llandovery. In Mr. Ruddy’s paper* the Lower 
Llandovery beds are said to be represented in this area by the Corwen- 
grit series of Prof. Hughes; and the latter, in summing up the 
evidence as to the succession in this section, says 1t goes to prove :— 
‘that the Corwen grits are distinct from the Pen-y-Glog grits ; that 
there is more evidence of a discordancy at their base than at the 
base of the pale slates or the Pen-y-Glog grits; that there are 
generally some beds of conglomerate, sandstone, or limestone with 
sandstone on the horizon of the Corwen grits; that the general 
facies of the few fossils obtained from these beds in the district 
examined is that of May-Hill rocks”. 

He states, further, that these Corwen grits are succeeded by beds 
which ‘ pass up into the ‘ pale slates’ of the Survey, which in turn 
pass up into the striped flaggy beds of Pen-y-Glog, on the top of which 
come grits, to be referred to the true Denbigh Flag and Grit series.” 
As the latter are the higher beds mentioned where plants were 
obtained, we have ample evidence of their position in regard to the 
succession exhibited here, both from beds below and from those 
which rest upon them. In other areas the position occupied here 
by the Denbighshire flag and grit series, the pale slates, and the 
Corwen grits seems to be chiefly filled up by the Llandovery or May- 
Hill group and the Tarannon slates: the latter, I think, with the late 
Mr. Salter, however, should always be included in the May-Hill 


* Quart. Journ. Geol. Soc. vol. xxxv. p. 200. 
t Ibid. vol. xxxili. p. 207. 


437 


DENBIGHSHIRE GRITS, NEAR CORWEN, NORTH WALES, 


Diagram Section from Nant Liechos, near Corwen, to Moel Morfydd. (Horizontal scale 1 inch to a mile. Vertical scalo 
about 3000 feet to 1 inch.) 


S.W. 3 N.E. 


Poviuion of slates 
of Pen-y-glog : 
slate-quarry. Dee river. Moel Morfydd. 


RSE eA CER RB 


1. Bala beds. 5. Wenlock series. 
2. Corwen Grits and Shales (Lower Llandovery). x Chief positions of plant-remains. 
3. Pale slates. i Base of Upper Silurian of Survey. 


4. Denbigh Grit and Flag series. 


N 


be 
N 


488 H. HICKS ON THE REMAINS OF PLANTS FROM THE 


group. The accompanying section (p. 487) will explain more fully 
the succession in this area and the actual position of the chief beds 
containing the plant-remains. 

The evidence of the geological horizon, derived from the fossils, is 
at present imperfect; for the animal remains found in the slates, j in 
association with the plant-remains, are chiefly Graptolites, though 
fragments of Hncrinites, species of Orthoceras, and some Brachiopods 
are occasionally found. The following species of Graptolites were 
recognized by Mr. Hopkinson, in a collection made by him and my- 
self at different horizons im this quarry in 1875, viz. Cyrto- 
graptus Murchisom, Monograptus priodon, M. Sedgwicku, M. ake 
M. vomerinus, M, Hath, and Retiolites Geitzianus*. 

These forms, he considered, were ‘characteristic of beds 6 the 
summit of the Coniston Mudstones, or base of the Coniston Flags ”’ 
in the Lake district. The abundance of Graptolites found in these 
beds would tend to show that the deposits, for the most part, were 
thrown down in a tolerably quiet sea. 

It seems, therefore, so far as the evidence can be read at present, 
that this immediate area was not greatly affected by the physical 
changes which occurred in the neighbouring areas at the close of 
the Bala epoch—that if uplifted above sea-level, it must have been 
previous to the deposition of the Corwen Grits, as shown by Prof. 
Hughes. The physical break, therefore, if it exists here at all, must 
be placed at that point, and not, as formerly supposed, at the base of 
the so-called Tarannon Shales. There is, however, no visible uncon- 
formity between the Lower and Upper Silurians anywhere in the 
sections in this neighbourhood; and it is quite possible that the area 
may have remained under water during the whole of the Mid- 
Silurian epoch+. The parts raised above sea-level were chiefly to 
the suuth-east, south, and north-west of this area. JI am inclined to 
think that there was not a very extensive land area, but numerous 
islands, some of them of volcanic origin. They reached undoubtedly 
as far as Shropshire to the 8.E., and to Caernarvonshire (and probably 
beyond it) to the N.W. There is no satisfactory evidence to show 
that they extended much further to the 8.W. than the neighbour- 
hood of Builth, as the deposits apparently accumulated uninter- 
ruptedly during this time in part at least of Caermarthenshire, in 
Pembrokeshire, and in Cardiganshire. This is the only way in which 
we can account for the presence in those areas of some thousands of 
feet of beds between the topmost Bala and the Wenlock series, and 


* Mr. C. Lapworth has also kindly examined a small collection made by me 
recently in the same quarry, and mentions the following forms as recognizable. 
They were chiefly collected from the middle bands of slate and above the beds 
with nodules and anthracite:—Letiolites Geinitzianus, Cyrtograptus Murchisont, 
Monograptus vomerinus, M. personatus, M. priodon (and vars. riccantonensis, 
Lapw., and Flemingit, Salt.). He states, as to the correlation of these beds with 
those in other areas, that, so far as the evidence derived from the above Grap- 
tolites can be made out, they would occupy a position equivalent tothe ‘‘lower 
zones of the Wenlock shale of Shropshire and the west of England.” 

+ See paper by author, Brit. Assoc. Report, 1875. 


* 


DENBIGHSHIRE GRITS, NEAR CORWEN, NORTH WALES. 489 


their almost entire absence in the Longmynd and other districts. 
For the same reason also it must be expected that a great diversity 
will be shown in the sediments belonging to this period in ditterent 
areas, and particularly so in proportion as they approached to or 
were distant from any of the raised parts. The fauna would also 
in consequence vary considerably. Jor these reasons it becomes 
difficult to correlate with any satisfaction all the beds which are 
found between the Hirnant limestone of the Bala epoch and the base 
of the Wenlock, which are known in different places chiefly under 
the name of Llandovery or May-Hill, or Tarannon, and in the area 
under consideration as the Denbigh Grit and Flag series. The con- 
clusions, therefore, arrived at in regard to the plant-remains and the 
geological horizon in which they are found are :—that theage of the 
beds must be somewhere between the base of the Wenlock and the 
Lower Llandovery, probably not far from the horizon of the May- 
- Hill beds (Mid Silurian) ; that the plants did not live on the surfaces 
on which they are now found; that their position here is an acci- 
dental one; that they were not brought from a great distance, as 
they occur at several horizons; that the shore-line from which they 
were derived was towards the south or west; and that the land 
areas were chiefly formed towards the close of the Bala epoch. 

If we compare these plant-remains with those discovered in lower 
Paleozoic rocks in other areas in this country, we do not find so im- 
portant an assemblage anywhere so low in position, certainly not at 
a lower horizon than the Upper Ludlow rocks, and probably not 
below the Devonian. It is probable also that an equal number of 
important plants have not been found together at so low a geological 
horizon in any other part of the world. Those found in the Silu- 
rian rocks elsewhere are :—the branch of a fern, described by Count 
Saporta under the name of Hopteris Morieri, discovered by Professor 
Moriére in the Middle Silurian at Angers, France; the Glyptodendron 
of Prof. Claypole, from the Clinton group of Ohio, America; and the 
species of Psilophytum, Annularia, and Sphenophyllum described by 
Prof. Lesquereux, also from the Silurian rocks of Ohio. It is a 
curious fact that in each of these areas, in Britain, France, and 
America, the land plants are in a greatly broken condition, and 
occur in association with a marine fauna. 

Their geological position in each country seems to bear out the 
view that physical changes were taking place almost contempora- 
neously in Britain, in parts of the continent of Europe, and in 
America at this time. These changes, which took place towards 
and at the close of the Lower-Silurian (Ordovician) epoch, caused 
land to be formed in each of these areas of greater extent than_ 
could have existed since the earliest Cambrian times; therefore 
it is probable that unless we find land plants in the lowest Cambrian 
deposits, we are not likely to meet with them in the intermediate 
groups, which appear to have been deposited upon each other unin- 
terruptedly. That there were periods of shallow water, when de- 
posits were thrown down nearly at an equal rate with the depression, 


490 H. HICKS ON THE REMAINS OF PLANTS FROM THE 


is certain ; and marine plants probably lived in abundance. The cel- 
lular structure of the marine plants, however, rendered them so 
readily liable to decomposition, that it is not much to be wondered 
at that their remains are seldom found. ‘The various markings 
which have been attributed to land and marine plants in the earlier 
rocks may in some cases have been produced by them; but others, 
as shown by Salter, must have been tracks produced by worms. 
Of the most important of those about which doubt still remains may 
be mentioned Hophyton of Torell, from the Lower Cambrian rocks 
of Scandinavia, but which I have also found at St. David’s. Cruziana, 
from the Lingula-flags of North Wales, supposed by Salter to be a 
worm-track, I believe, from evidence I have been collecting for some 
time, will prove to be an Alga. Buthotrephis, found in the Lingula- 
flags and in the Arenig rocks in Wales, and by Prof. Nicholson in 
the Skiddaw Slates of Cumberland*, but first discovered and de- 
scribed by Prof. Hall in America, appears also to be allted to the 
Algee. Ot Hophyton? explanatum, which I found in the Tremadoc 
rocks of St. David’s, I fear the evidence is scarcely sufficient to ally 
it with land plants. Its strong tubular structure renders it unlike 
any known land plant; and the only other fossil found yet to which 
it can be compared is the Pyritonema of Prof. M‘Coy, placed by him 
amongst the Zoophytes, though its true nature is still a matter of 
much doubt. 


Appendix. By R. Erurriver, Esq., F.R.S.. Pres. Geol. Soc. 


Karty in the present month Dr. Hicks brought for my inspection 
several slabs of micaceous sandstone, having upon their surfaces 
numerous fragments of carbonaceous matter, which possessed no 
definite shape or apparent structure. Their general appearance, in 
some cases, was that of decomposed coniferous wood, in others, 
resembling bundles of finely striated black or dark-brown carbon, 
brittle or tough, the black portions being by far the most brittle ; 
the lighter and darker remains, however, undoubtedly belong to the 
same plant, but differently mineralized. Hitherto, in Britain, no 
true plant-remains are known to occur below the Upper Ludlow, 
and the only recognized species in that formation is Pachytheca, or 
Pachysporangium. Between the Upper Ludlow and the base of the 
Denbighshire Grits no traces whatever of plant-remains have yet 
occurred in British Silurian strata. 

Dr. Dawson, as far back as 1859, in his paper upon the “ Fossil 
Plants from the Devonian Rocks of Canada’’}, described, amongst 


* The fossils formerly placed by Prof. Nicholson in the genus Buthotrephis, 
from the Skiddaw Slates, have been since redescribed by him and Dr. Dawson 
under the generic name of Protannularia. 

tT Quart. Journ. Geol. Soc. vol. xv. p. 484. 


DENBIGHSHIRE GRITS, NEAR CORWEN, NORTH WALES, 491 


other things, a new genus which he called Prototaxites, and the 
species Loganz; he considered this to be the oldest known fossil 
tree in America, and, as such, gave it the name of Prototaxites, 
believing that it belonged to the Taxineee. Subsequently (1863), in 
vol. xix. of our Journal, he described other fossil plants from the 
Gaspé Devonians ; and apparently the same or a similar fossil is again 
described under the name of Nematoxylon crassum. 

Specimens of this plant (Prototaaites) were subsequently examined 
by Wm. Carruthers, Esq., F.R.S., who pronounced it to be a colossal 
Alga, or seaweed, and named it Nematophycus. Mr. Carruthers 
published an important paper upon this plant, wherein he gives his 
reasons for widely differing from Dr. Dawson *. 

Immediately on the receipt of Dr. Hicks’s specimens, and knowing 
how large a problem depended upon a right interpretation of the 
structure and affinities of the plant-remains, through their strati- 
graphical position or age, I at once obtained Mr. E.'T. Newton’s aid in 
preparing microscopical sections. The result has been in the highest 
degree satisfactory ; and he at once determined these remains from 
the Denbighshire Grits of Pen-y-Glog to be the Nematophycus of 
Mr. Carruthers. 

Mr. Newton and myself have microscopically examined several 
sections of both the lighter and darker portions of the carbonaceous 
matter placed at our disposal by Dr. Hicks; and we feel convinced 
that they both present the same structure, the difference seen being 
almost wholly due to the fact that the intensely opaque nature of 
the carbonaceous matter in the darker specimens obscures the 
structure. Mr. Newton’s observations were therefore made upon 
the light-coloured specimens, which we have carefully examined 
together. When examined with transmitted light and with a low 
power, the fragments show that the fibrous appearance is due to 
a number of dark rod-like tubes or cells running in a longitudinal 
direction, but in a more or less vermiform or wavy manner, some- 
times closely packed together (fig. 2), sometimes more widely 
separated: these tubes or cells have one general direction, and 
consequently appear nearly parallel; but closer examination shows 
that other cells irregularly curve in and out of plane, and are not 
parallel. 

In many parts a dark and apparently granular substance seems 
to fill the tubes, and may be resolved into round globules varying in 
size (fig. 3). 

In certain favourable spots some of the tubes appear to be crossed 
by very fine transverse lines (fig. 4) which strongly resemble spiral 
fibres ; these lines, however, may be due to the nature of the mineral 
which fills the cells. On others a network of fine lines may be 
seen spreading over the tubes; these might be thought accidental 
but for their frequent occurrence. In transverse section (fig. 1) the 


* Trans. of the Royal Microscopical Soc., Monthly Microscopical Journal, 
1872. 


499 H. HICKS ON THE REMAINS OF PLANTS FROM THE 


cells are more or less circular, and are separated by greater or less 
interspaces in different parts of the section. 

Compared with the Nematophycus Logani as figured by Mr. Car- 
ruthers, the minute structure of these carbonaceous fragments will 
be found to agree precisely, with the one exception that the finer 
tubes or cells filling the interspaces between the larger ones, which 
Mr. Carruthers figures and describes in Nematophycus, are not so 
distinct in Dr. Hicks’s specimens. The double spiral fibres figured 
by Dr. Dawson when he first deseribed his Prototavites (= Nema- 
tophycus, Carr.), are rather to be referred to the interlacing cells 
(network in Pen-y-Glog specimens) than to the fine spiral fibres 
seen in the latter (fig. 4). 

The rounded seed-like bodies found with the carbonaceous frag- 
ments are hollow and thick-walled. In microscopical sections the 
wall is found to be composed of radiating fibres (cells ?) arranged 
nearly parallel to each other and slightly wavy (fig. 8). These cells 
are irregularly filled with spore-like bodies. The seed-like bodies 
resemble the Pachytheca from the Upper Ludlow beds both in 
their outward form and in the radiated structure of their walls. 

The question naturally arises, Are the carbonaceous fragments 
so well known to occur in the Ludlow bed (Downton sandstone) of 
the same nature and structure as the Denbigshire-grit specimens ? 
Mr. Newton has not been able to obtain transparent sections of the 
Ludlow woody specimens; but, from what little we have been able 
to make out, these Ludlow fragments likewise show tubular struc- 
ture, but not quite of the same character as those from Pen-y-Glog. 

On first seeing. Dr. Hicks’s specimens I pronounced the remains 
to be those of some marine Alga; but I was not then acquainted 
either with Mr. Carruthers’s paper on Nematophycus or Dr. Dawson’s 
description of Prototavites, neither being then known as British 
plants; and no opportunity had occurred to me for examination. 
The sections made by Mr. Newton reveal in the most perfect and 
satisfactory manner the innumerable vermicular cellular filaments 
which constitute nearly the entire structure of what must have been 
the stem, and also the dense spongy nature produced by the smaller 
tubes, which seem to ramify irregularly. The larger tubes are not 
strictly parallel to each other, although they run in the same general 
direction ; they appear to be continuous, and their ends bluntly 
rounded. 

The description of the structure of Nematophycus, and the discussion 
of its affinities with certain orders in the Chlorospermee, are so ably 
and completely done by Mr. Carruthers, that little is left for further 
description, especially as there is no doubt whatever that the plant- 
remains in the Denbighshire Grit from Pen-y-Glog are unques- 
tionably Nematophycus of Carruthers, and the Prototaaites and 
Nematoxylon of Dr. Dawson, referred by Dr. Dawson to the Conifere 
through the Taxines or Yews. 

The fragments, small as they are, show unmistakably that we 
have at this low horizon the Gaspé plant (Prototaaites) which occurs 
in the Devonian rocks of America. ‘This is an important fact, as 


DENBIGHSHIRE GRITS, NEAR CORWEN, NORTH WALES. 493 


the so-called Fucoidal remains are mere impressions, and many, if 
not all, are trails or burrows of Annelida. 

The interest at first attached to these special remains was centred 
in the hope that they were portions of terrestrial vegetation grow- 
ing on a contiguous land surface during the deposition of the Den- 
bighshire Grits. Now, however, we have clear evidence that these 
remains, to which we more especially refer, formed portions of a 
colossal seaweed whose habit resembled that of the North-Pacific 
species of the genus Nereocystis and the arborescent JLessonia, 
and probably, as Mr. Carruthers suggests, the Macrocystis pyrifera, 
which attains the length of 700 feet. The arborescent Lessonie, 
belonging to the natural order Laminariaces, form large submarine 
forests, the stems of which, when dry, resemble exogenous wood, 
owing to a false exogenous growth. ‘This pseudo-exogenous struc- 
ture is well known in many of the Alge. Mr. Carruthers suggests 
that in the arborescent Lessonie we have a near approach to the 
Devonian Nematophycus or Prototawites. Many of the stems in 
Lessonia measure a foot in diameter and 30 feet in length. The 
Laminarie of our own shores exhibit a pseudo-exogenous growth ; 
but, as is well known, the difference in growth between the Antarctic 
Lessonia and the genus Laminaria consists in the “ exogenous in- 
crease in Laminaria being from below upwards, according to the 
growth of the roots, while in the genus Lessonia the growth is 
from above downwards, in proportion to the increase of the leaves”*. 
Macrocystis belongs to the same natural order (Laminariacez), but 
possesses a different habit. I may mention also D’Urvillea Harveyi, 
Hook., another of the Laminariaces, which possesses a stem-struc- 
ture most closely resembling Nematophycus in the vermiform nature 
of the cells or tubes, and their irregular semiparallel arrangement. 
Dr. Hooker describes and figures this plant in his ‘ Antarctic Voyage,’ 
vol. 11. t. 165-6 (fig. 2); a figure of the longitudinal structure of the 
stem is given on the plate. The Corwen Nematophycus is converted 
into amorphous silex. This condition interferes with the microscopic 
structure being readily made out, giving apparently false lines across 
the tubes or cells; and whether the spiral fibres are inside or out- 
side the large tubes, or independent structures outside, I am unable 
tosay ; neither can Mr. Newton or myself clearly make out that the 
so-called spiral fibres or concentric lines upon the tubes can be re- 
solved into minute dots. His figure (fig. 4) is most carefully drawn ; 
and one of the tubes containing the globular bodies is also delicately 
and similarly lined. Fresh observations may enable us to determine 
whether the spiral lines can be resolved into dots or not. The lon- 
gitudinal and transverse sections prepared by Mr. Newton show 
every feature described by Mr. Carruthers in his paper. The elon- 
gated cylindrical cells, of two sizes, appear completely and irregu- 
larly interwoven, as Mr. Carruthers well expresses it, into a kind of 


* Berkeleys Introduction to COryptogamic Botany, p. 57; Carruthers, 
Monthly Microscopical Journal, 1872, pp. 170, 171. 


494 H. HICKS ON THE REMAINS OF PLANTS FROM THE 


felted mass ; and in the figure of the longitudinal structure of tho 
stem in D’Urvillea Harveyr given by Dr. Hooker (loc. cit.) an irre- 
gular vermiform interlacing and interweaving of the cells or tubes 
is shown, remarkably agreeing with the structure of Mematophycus. 
It would be interesting to ascertain if the large round spore-like 
bodies (which are here termed Pachytheca or Pachysporangium, fig. 7) 
are the sporangia of Vematophycus: the cells which radiate from the 
central cavity are certainly filled with round spore-like bodies ar- 
ranged in single file, or disposed in such a manner as to be shed from 
the periphery or circumference of the body called Pachytheca. This 
is manifest in the structure of one of these cut through and shown 
under the microscope (fig. 9). The large tubes or cells, constituting the 
stem structure, which appear here and there to contain round spore- 
like bodies similar to these, do not really do so; they are, I believe, ac- 
cidentally scattered here and there, and appear to be within rather 
than upon the tubes in the microscopic sections. 

On two of Dr. Hicks’s rock specimens there are singular honey- 
comb-celled bodies having all the appearance of being coriaceous or 
membranous capsular bodies, or sporangia; the outer portion was 
definitely composed of either four- or six-sided cells. This sac-like 
body may have played some part in the history of Mematophycus. 
Under any circumstances they are worthy of notice, and are un- 
doubtedly organic. 

The plant called Prototaxites by Dr. Dawson, from the Gaspé 
beds, of Devonian age, was one of the oldest plants then known; the 
previously known oldest plants are mere fragments of woody matter : 
these occur with Pachytheca in the Upper Ludlow, of the Ludlow 
area; and no older British rocks have, until now, yielded any. 
Looking at the probable distribution of a marine flora the plants 
composing which were of such magnitude and probably widely 
spread over the Devonian sea-bed, we naturally turn to the origin or 
source of the Gaspé Devonian species. The presence of Nema- 
tophycus in the Denbighshire Grits is proved by these researches of 
Dr. Hicks; but no remains of a similar kind have been found 
through the higher beds of the Woolhope, Wenlock, and Ludlow 
rocks—except at the close of the Ludlow, in the Downton shales, 
if the seed-like bodies termed Pachysporangium or Pachytheca are 
to be regarded as portions of Nematophycus. If they have no relation 
to each other, then we have to account for the wide distribution in 
time of both—Nematophycus from the Denbighshire Grits to the 
Devonian, and Pachytheca from the Denbighshire Grits to the Upper 
Ludiow. We are warranted, I think, in believing that there is far 
less chance of these being one species than modified descendants 
from an older stock, the entire area never having been dry land at 
one time since the deposition of the Denbighshire Grits ; I therefore 
give this the specific name NMematophycus Hicks. I look forward 
to finding the remains of this group of Algee in rocks of far higher 
antiquity ; for it is evident that marine plants of such colossal size 
as the Gaspé species is known to have been could not then have 


DENBIGHSHIRE GRITS, NEAR CORWEN, NORTH WALES. 495 


made their first appearance. Again, the habitat of Nematophycus, 
if it at all resembled the present Lessonie in growth at the bottom 
of the ocean, may have survived many changes between land and 
sea during the deposition of the Upper Silurian rocks, while yet the 
sea-bottom or bed of the Silurian Sea was never exposed or became 
dry land. The finding and determining the nature of these remains 
opens up a great problem, so far as the age, continuity, and distribu- 
tion of Cryptogamic life in palzozoic time is concerned. 


EXPLANATION OF PLATE XXV. 
1. Nematophycus Hicksii: cross section, x 200. 
2. Ditto: longitudinal section, x 200. 
3. Ditto: tubes filled apparently with granular substance, x 329. 
4, Ditto: tubes covered by fine transverse lines, X 320. 
5. Ditto: portion of stem, natural size. 
6. Ditto: fragments, ditto. 
7. Pachytheca: X 6. 
8. Ditto: portion of wail, x 200. 
9. Ditto: fibre in wail, x 200. 
0. Microspores, probably of a Lycopodiaceous plant, x 15. 


Discussion. 


Mr. Carruruers spoke of the importance of Dr. Hicks’s discovery, 
and said that after the President’s note it was needless for him to 
add much. He had come in the main to the same conclusion. The 
view of the significance of the Pachythece was interesting : but the 
data which had come before himself hardly warranted his regarding 
them as Algee. He explained the reasons why he was unable to 
accept the President’s view, and agreed with that of Sir Joseph 
Hooker—namely, regarding them as Lycopodiaceous. He thought 
also that the signs of a vascular axis in some of the specimens 
proved a land-flora; also there were some spores, not sporangia, 
which bore out this view. 

Mr. Horpxryson said that the Graptolites were partly Middle 
Silurian and partly Upper Silurian forms, some being Llandovery 
species, here dying out, and others Wenlock species, first appearing 
here. The change in type seemed to imply an alteration in physical 
conditions. 

Mr. Dr Rance referred to some Ludlow pebbles at the base of the 
Carboniferous, and said that anthracite occurred in beds of the Lake- 
district of about the same age. 

Prof. Duncan spoke of the good fortune which had attended Dr. 
Hicks’s investigations in unpromising ground. The cellular plant 
discovered by him was a very remarkable one; it reminded him of 
Codium. The undulated underpart of the cellulur structure was 
very remarkable, indicating a general rugosity ora spiral fibre ; that 
fibre occurred in modern water-plants. In one part there was 
something like a dissepiment. As in the beds above the Gannister, 
there might be a common link here of land- and water-plants. He 


496 H. HICKS ON PLANT-REMAINS FROM DENBIGHSHIRE GRITS. 


commented on Pachytheca, and asked whether it could have any 
relation to the floats in the Sargassum. These old plants seemed 
capable of existing under conditions very similar to the present. 

Mr. Tawney said he thought a Lepidodendron had been found in 
the Silurians of America. 

Dr. Hicks said that Saporta had described a species from about 
the same horizon, supposed to be a mineralized fern. He thought 
that here was an admixture of land- and marine plants, and believed 
much more would be found. At. present Nematophycus was the 
best. preserved. 


Tt. Journ Geol. Soc. Vol. XXXVI. PL. XXVT. 


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ON THE UPPER JURASSIC OF THE PARIS BASIN. 497 


38. On the Corretation of the Upper Jurassic Rocks of ENGLanp 
with those of the Contrnent.—Part I. Tar Paris Basry. By 
the Rev. J. F. Buaxz, M.A., F.G.8. (Read April 27, 1881.) 


[Puarr XXVI.] 


In former papers on the Portland Rocks*, the Kimmeridge Clayt, 
and the Corallian Rocks +, as developed in our own country, it has 
been pointed out that, while the normal deposits of the period 
commencing with the Oxford Clay and continuing to the close 
of the Jurassic era were essentially argillaceous, the uni- 
formity has been broken by certain episodes which have resulted 
in the formation of distinct kinds of rocks, but that, in spite of 
these episodes, there is a continuousness both in the physical and 
biological features, uniting the whole into one great group, to which 
the term Upper Jurassic is appropriated. The present study ought 
therefore, logically, to include a correlation of the Oxford-Clay series ; 
but though the upper portions of that series come to be incidentally 
examined, the far wider range and greater constancy of the lower 
portion would render its examination a more arduous and _ less 
interesting task; and it is found convenient to have for base a 
thick mass of clay, which may almost everywhere be recognized, 
however much the upper part may be encroached upon lithologically 
by the various preludes to the Corallian series. In point of fact, 
the rocks hitherto called Corallian in England comprise much that 
is placed in the Oxfordian by the French geologists ; and our corre- 
lation is therefore only stopped when the rocks universally called 
Oxfordian are reached. 

The Upper Jurassic rocks of France lie in two distinct areas. 
The more northern is that which is drained by the Seine and the 
Loire and smaller rivers having a similar direction; the more 
southern is a continuation of the Swiss Jura, or lies to the south of 
the central plateau of Auvergne. The former constitutes the basin 
of Paris, round which city the Jurassic rocks form an irregular 
curve; and their development in this range forms the subject of the 
present study, the Upper Jurassic rocks of other districts being 
left for a future occasion. 

Much good work has been done by the French geologists in the 
description of the various portions into which this basin may be 
divided, and in the correlation of the rocks with one another, by 
the aid of which it is possible for a foreigner to pass from spot to 
spot, appreciating, confirming, or even correcting the stratigraphical 
succession and its interpretation. The advantage that such a 
student has is that, instead of being confined to one area, and 
being obliged to obtain his ideas from it, he can make the several 

* Quart. Journ. Geol. Soc. vol. xxxvi. p. 189. 


Tt Op. cit. vol. xxxi. p. 196. 
t Op. cit. vol. xxxiii. p. 260. 


Q.J.G.S. No. 148. Dh 56 


.498 J. F. BLAKE ON THE UPPER 


areas mutually throw light on each other, and present a complete 
idea of the whole and not merely of isolated fragments. Oppel, in 
his work ‘ Der Juraformation, made such a comparative study in 
1860; and Hébert, in his pamphlet ‘ Les Mers Anciennes,’ did the 
same for the basin of Paris in 1857. Since those dates much new 
material has accumulated; and it is possible also to bring English 
rocks into the comparison. Hence several fresh questions have arisen, 
and divergences of interpretation have been developed, on which 
it is impossible to come to any conclusion without a uniform and 
comprehensive study in the localities themselves, which will usually 
show that nature is much more simple than her various interpre- 
ters between them would make her. | 

The following numbered list gives some account of what has been 
written on the Upper Jurassic rocks of the Paris Basin :— 


Inst of Works and Papers consulted on the Upper Jurassic 
Rocks of the Paris Basin. 


(1) 1825-8. Caumont. Explication de la carte géologique de Normandie. — 

(2) 1842. Savvace & Buvienier. Statistique minéralogique et géologique du 
département des Ardennes. 

(3) 1844. Ravin. Mémoire sur la constitution géologique du Sancerrois. 
Bull. Soc. Géol. Fr. ser. 2, vol. it. p. 84. 

(4) 1844. Rover. Note sur les terrains jurassiques supérieurs et moyens de la 
Haute Marne. Bull. Soc. Géol. Fr. ser. 2, vol. ii. p. 705. 

(5) 1844. Royer. Comparaison des terrains jurassiques de l’Yonne avec ceux 
de la Haute Marne. Bull Soc. Géol. Fr. ser. 2, vol. 11. p. 714. 

(6) 1844. Corrzav, Ann. Stat. de ’Yonne, p. 236. 

(7) 1846. Luymertz. Statistique du département de l’Aube. 

(8) 1847. Graves. Essai sur la topographie géognostique du département de 
VOise. 

(9) 1847. Corrrau, Bull. Soc. Sci. Hist. et Nat. de Yonne, t. i. pp. 23 and 367. 

(10) 1850. Bounanerr & Bertera. Texte explicatif de la carte géologique 
du Cher. 

(11) 1851. Royer. Sur quelques failles dans la Haute Marne. Bull. Soc. Géol. 
Fr. ser. 2, vol. viil. p. 564. 

(12) 1851. Royer. Apergu sur les terrains Corallien et Oxfordien de la Haute 
Marne. Bull. Soc. Géol. Fr. ser. 2, vol. viii. p. 600 

(13) 1852. Buvienter. Statistique Géologique, Minéralogique, Minérallurgique 
et Paléontologique du département cle la Meuse. 

(14) 1852. D’Orsieny. Cours élémentaire de Paléontologie et de Géologie stra- 
tigraphiques. Vol. 2, 

(15) 1853. Ravin. Sur l Oxford Clay du département de l’Yonne. Bull. Soe. 
Géol. Fr. ser. 2, vol. x. p. 485. 

(16) 1855. Correav. Notice sur lage des couches inférieures et moyennes de 
Vétage Corallien du département de ?Yonne. Bull. Soc. Géol. Fr. ser. 
2, vol. xii. p. 693. 

- (17) 1855. Manis. Statistique géologique de la Charente Inférieure. 

(18) 1856. Corrzav. Compte Rendu de la session tenue par la Société Géolo- 
gique de France le 7 Septembre a Joinville (Haute Marne).—Compa- 
raison des terrains observés par la Société avex ceux du département 
de lYonne. Bull. Soc. Sc. Hist. et Nat. Yonne. 

(19) 1856. Réunion Extraordinaire de la Société Géol. de France a Joinville. 
Bull. Soe. Géol. Fr. ser. 2, vol. xiii. p. 787 (Royer). 

(20) 1856. Buvienter. Note sur les calcaires 4 Astartes et l’étage jurassique 
supérieur et moyen de la Meuse et de la Haute Marne. Bull. Soe. 
Géol. Fr. ser. 2, vol. xiii. p. 843. 


JURASSIC OF THE PARIS BASIN. 499 


(21) 1857. Huserr. Les Mers anciennes et leur rivages dans le bassin de Paris. 
Part I. Terrain Jurassique. 

(22) 1857. Buvienter. Observations sur le terrain jurassique de la partie 
orientale du bassin de Paris. Bull. Soc. Géol. Fr. ser. 2, vol. xiv. p. 595. 

(23) 1858. Coquanp. Description géologique de l’étage Purbeckien dans les 
deux Charentes. Bull. Soc. Géol. Fr. ser. 2, vol. xv. p. 577. 

(24) 1858. Coquanp. Statistique de la Charente. 

(25) on LeymericE & Ravuin. Statistique géologique du département de 
*Yonne. 

(26) 1858. Réunion extraordinaire de la Société Géologique de France a Nevers 
(Ebray). Bull. Soc. Géol. Fr. ser. 2, vol. xv. p. 680. 

(27) 1858. Opprt. Die Juraformation. 

(28) 1860. Héserr. Du terrain jurassique supérieur sur les cotes de la Manche. 
Bull. Soc. Géol. Fr. ser. 2, vol. xvii. p. 300. 

(29) 1861. Gousrrr. Note sur le gisement de Glos. [Ina “ Note sur les Tirigonies 
clavellées de VOxford Clay et du Coral Rag,” by M. Hébert.|] Journ. de 
ae Apr. 1861. See also Bull. Soc. Géol. Fr. ser. 2, vol. xviii. 
p. 520. 

(30) 1863. Dotirus. La Faune Kimméridienne du Cap de la Héve. 

(31) 1863. Hiserr. Observations Géologiques sur quelques points du départe- 
ment del’Yonne. Bull. Soc. Sc. Hist. et Nat. Yonne, 1863. 

(32) 1864. Esray. Etudes Paléontologiques sur le département de la Niévre. 

(38) 1865. Rieavx. Notice stratigraphique sur le Bas-Boulonnais. Bull. Soc. 
Acad. Boulogne. 

(34) 1865. Corrzav. Deux jours d’excursion dans le terrain jurassique des 
environs de Tonnerre (Yonne). Bull. Soc. Sc. Hist. et Nat. Yonne, 1865. 

(35) 1865. Waacen. Versuch einer allgemeinen Classification der Schichten des 
oberen Jura. 

(36) 1866. Punuar. Notes sur les assises supérieures du terrain jurassique de 
Boulogne-sur-Mer, &e. Bull. Soe. Géol. Fr. ser. 2, vol. xxiii. p. 193. 

(37) 1866. Hipert. Note sur le terrain jurassique du Boulonnais. Bull. Soc. 
Géol. Fr. ser. 2, vol. xxiii. p. 216. 

(38) 1866. Samann. Observations sur les communications précédentes, 1. ¢. 
p. 220. 

(39) 1866. Dz Lorton & Petuat. Monographie paléontologique et géologique 
de l’étage Portlandien des environs de Boulogne-sur-Mer. Mém. Soc. 
Phys. Genéve, tom. xix. 

(40) 1867. Tompecx. Note sur Vétage Portlandien de la Haute Marne et son 
parallélisme avec celui du Boulonnais. Bull. Soc. Géol. Fr. ser. 2, 
vol. xxiv, p. 187. 

(41) 1868. Pzuvar. Observations sur quelques assises du terrain jurassique 
supérieur du Bas-Boulonnais. Bull. Soc. Géol. Fr. ser. 2, vol. xxv. 
p- 196. 

(42) 1868. De Lorton & Corrzav. Monographie paléontologique et géologique 
de l’étage Portlandien du département de ’Yonne. Bull. Soc. Se. Hist. 
et Nat. Yonne, ser. 2, tom. i. 

(43) 1868. Corrzav. Nouvelles observations sur le terrain jurassique des en- 
virons de Tonnerre. Bull. Soc. Se. Hist. et Nat. Yonne. ser. 2, tom. ii. 

(44) 1868. Tompecx. Note sur le terrain Portlandien de la Haute Marne. 
Bull. Soc. Géol. Fr. ser. 2, vol. xxv. p. 456. 

(45) 1868. Tomsecx. Note sur les terrains Corallien et Kimmeridien de la 
Haute Marne. Bull. Soc. Géol. Fr. ser. 2, vol. xxv. p. 458. 

(46) 1870. Petar. Sur l’étage Portlandien du Paysde Bray. Bull. Soc. Géol. 
Fr, ser. 2, vol. xxvil. p. 683. 

(47) 1870. Prewuar. Nouvelles observations sur quelques assises du terrain 
jurassique supérieur du Boulonnais. Bull. Soc. Géol. Fr. ser. 2, 
vol. xxvii. p. 683. 

(48) 1870. Tompeck. Sur les étages jurassiques supérieurs de la Haute Marne. 
Bull. Soc. Géol. Fr. ser. 2, vol. xxvii. p. 687. 

(49) 1870. Lennier. Etudes Géologiques et Paléontologiques de la Haute 
Normandie. 

24 2 


500 J. F. BLAKE ON THE UPPER 


(50) 1872. Dr Lorton, Rover, & Tompzck. Monographie des étages jurassiques 
supérieurs de la Haute Marne. Mém. Soc. Linn. Norm. vol. xvi. 

(51) 1872. Savvacr. Note sur la position des couches a polypiers et a Terebra- 
tula insignis dans le Boulonnais. Bull. Soc. Géol. Fr. ser. 2. vol. xxix. 
p. 215. 

(52) 1872. Pruiat. Observations en réponse a la communication de M. Sau- 
vage sur la position stratigraphique des calcaires du Mont des Boucards. 
Ibid. p. 223. 

(53) 1873. Ricaux. Notes pour servir ala géologie du Boulonnais. I. Notes sur 
quelques sondages. Bull. Soc. Acad. Boulogne. 

(54) 1873. Hupert. Note additionelle 4 la communication relative a létage 
Tithonique. Bull. Soc. Géol. Fr. ser. 3, vol. i. p. 67. 

(55) 1873. Tomerck. Note sur l’Oxfordien et le Corallien de la Haute Marne. 
Bull. Soc. Géol. Fr. ser. 3, vol. i. p. 335. 

(56) 1874. Dz Lorton & Pettat. Monographie Paléontologique et Géologique 
des étages supérieurs de la formation jurassique des environs de Boulogne- 
sur-Mer. Parti. Mém. Soc. Phys. Genéve, tom. xxiii. p. 253. 

(57) 1874. TomBecx. Note sur l’Oxfordien et le Corallien de la Haute Marne. 
Bull. Soe. Géol. Fr. ser. 3, vol. ii. p. 14. 

(58) 1874. Tomsrcn. Note sur une excursion géologique faite au travers des 
terrains Coralliens et Oxfordiens de la Haute Marne. Bull. Soe. Géol. 
Fr. ser. 3, vol. ii. p. 251. 

(59) 1874. Bayan. Sur la succession des assises et des faunes dans les terrains 
jurassiques supérieurs. Bull. Soc. Géol. Fr. ser. 3, vol. ii. p. 316. 
Followed by observations by Tombeck and Pellat. Ibid. p. 348. 

(60) 1875. Dovvittx & Jourpy. Note sur la partie moyenne du terrain juras- 
sique dans le Berry. Bull. Soc. Géol. Fr. ser. 3, vol. iii. p. 93. 

(61) 1876. Dz Lorton & Pettat. Monographie, &e. des ét. sup. jur. de Boulogne. 
Part ii. Mém. Soc. Phys. Genéve, tom. xxiv. p. l. 

(62) 1876. Putiar. Emersion du sud et de lest du bassin Parisien a la fin dela 
période jurassique, et extension de la limite imférieure de l’étage Port- 
landien du Boulonnais. Bull. Soe. Géol. Fr. ser. 3, vol. iv. p. 364. 

(63) 1876. TripoLer. Sur les terrains jurassiques supérieurs de la Haute 
Marne comparés a ceux du Jura suisse et francais. Bull. Soc. Géol. Fr. 
ser, 3, vol. iv. p. 259. 

(64) 1876. Tompecx. Note sur le Corallien et lArgovien de la Haute Marne. 
Bull. Soe. Géol. Fr. ser. 3, vol. iv. p. 162. 

(65) 1877. Tompzcx. Coralliende la Haute Marne. Bull. Soc. Géol. Fr. ser. 3, 
vol. v. p. 24. 

(66) 1878. Toucan Sur la position vraie de la zone 4 Ammonites tenuilobatus 
dans la Haute Marne et ailleurs. Bull. Soc. Géol. Fr. ser. 3, vol. vi. p. 6. 

(67) 1878. Srruckmann. Der obere Jura der Umgegend von Hannover. 

(68) 1878. Prnuat, Résumé d’une description du terrain jurassique supérieur 
du Bas Boulonnais, &e. 

(69) 1879. Dz Lapparentr. Le Pays de Bray. 

(70) 1879. Tomprck. Réponse aux observations de M. Buvignier. Bull. Soc. 
Géol. Fr. ser. 3, vol. vi. p. 310. 


On studying these writings it soon becomes obvious not only that 
questions of theory are in dispute, but that facts are variously 
stated also, and that the latter must first be settled before the 
former can have a solid foundation. The study therefore becomes 
divided into two parts :—first, observations made for the verification 
or modification of the accounts of the true sequence in the various 
areas ; and, secondly, discussion of the opinions expressed on the 
relations and groupings of the rocks, with special reference to the 
equivalents in our own country. 

Within the area here spoken of as the Paris Basin there are five 
ranges, of very unequal size and importance, separated from each 


JURASSIC OF THE PARIS BASIN. 501 — 


other by intervals in which older or newer rocks occupy the surface 
of the ground. ‘These are 3 


I. From the Ardennes to the Cher. 
JI. The two Charentes. 
III. Normandy, with Orne and Sarthe. 
IV. The Pays de Bray. 
V. The Boulonnais. 


The first-named district, though not the nearest to England, nor 
presenting characters most similar to those of our own country, yet, 
from its large size, the continuousness of its deposits and the labour 
that has been bestowed upon them, presents itself as the most typi- 
cal for France, and the one therefore to be studied first, so that we 
may better understand the last three, which in some sense are in- 
termediate between the French and English types. 


I. From tHe ARDENNES TO THE CHER. 


Although this range has been studied continuously from one end 
to the other, it is necessary in description to subdivide it into those 
smaller areas which have been made the subjects of special mono- 
graphs by French geologists. 

1. The Ardennes Depariment.—The rocks of this area have received 
illustration at the hands of MM, Sauvage and Buvignier (2)*, who 
give the following classification 


1. Uprer Group. 
a. Marls with Exegyra virgula, 160 ft. 
6. Caleaire 4 Astartes, 21 ft. 
2. Corau Raa, 250 ft. 
3. OXFORDIAN. 
a. Ferruginous oolite, 28 ft. 
b. Marls and limestones, 300 ft. 
(1) Shelly siliceous limestone, (2) marls, (3) ironstone. 
e. Lower marls with ironstone, 30 ft. 


Commencing this series from the base, 3 ¢ may be passed over as 
Callovian, while 3 6 (2) and (3), forming the great mass of the 300 ft. 
and occupying the valleys, will represent the undoubted Oxford Clay. 
The series 36 (1) then commences the group of rocks now under 
study. If for “siliceous limestone” we read its equivalent, ‘“ calca- 
reous grit,” we have the term which will exactly suit the English 
geologist, who at once recognizes in it a representative of his ‘ Lower 
Calcareous Grit.’ At the base, as seen in quarries on the road from 
the station of Launois to Neuvizy, is hard blue cale-grit ; and above 
this, nodular beds of grit in a subargillaceous matrix. Some of 
these are very light, like the Upper Calcareous Grit of Yorkshire, and 
are said by Sauvage and Buvignier to be soluble in potash, indicating 
a possible organic origin, like the Renulina-grits of Scarborough ; 
the fossils, however, are in some cases ‘ beekized.’ These nodular 
_ beds, which begin to present some of that peculiar feature indicated 


* See list of works above. 


502 J. F. BLAKE ON THE UPPER 


Fig. 1.—Map of Country near Neuvizy. (Scale 1 : 80,000.) 


x L aunots L 


by the term ‘terrain a chailles’**, contain a characteristic fauna, viz. 
Ammonites cordatus, Modiola bipartita, Perna quadrata, Pecten fi- 
brosus, Avicula expansa, Exogyra spiralis, Ostrea dilatata, O. rastellaris 
Rhynchonella Thurmanni, and several others less certainly identified, 
but all indicating a horizon similar to that of the Nothe grits and 
clays or of the Lower Calcareous Grit of Yorkshire. At the top this 
series becomes much more calcareous, and finally contains abundant 
oolitic grains, while a change of fauna takes place. This has been 
very imperfectly examined; but the three chief species are Ammo- 


* This term “terrain a chailles” is a misleading one. It is often used as if 
indicating a fixed horizon ; but it appears from French writers that there is one 
of these ‘terrains’ in the Callovian, another in the Oxfordian (the present 
one), and a third in the Corallian ; so that it is really a mere petrological phrase 
similar to “nodular beds”. 


JURASSIC OF THE PARIS BASIN. 503 


nites oculatus of the variety called A. Bachianus, Oppel, Trigonia 
spinitfera, d’ Orb. (Prodr.), and Rhynchonella Thurmanni. The Tri- 
gonea is specially noticeable: it has been found by Mr. Hudleston at 
Snainton in Yorkshire, in a quarry belonging to the passage beds 
above the Lower Calcareous Grit, and named by Dr. Lycett 7’. snain- 
tonensis ; but, though never figured by the French geologists, it seems 
to be abundant in the Ardennes department, and occurs on the same 
horizon in the Vosges. The Ammonite serves to show how essen- 
tially Oxfordian these beds still are. Their total thickness is about 
50 feet. 

Next in the series is the Ferruginous Oolite, which, from its easily 
recognized mineral character and its rich fauna, has attracted con- 
siderable attention, and has been constantly used by authors as a 
term of comparison for beds in other districts. It is of no great 
thickness, 28 feet being given asitsmaximum. At Vieil-St.-Remy it 
is an earthy limestone with scattered oolitic grains of limonite and 
crowded with fossils. 

At Neuvizy almost every thing but the oolite grains has been dis- 
solved, and these remain as a loose deposit containing ‘ beekized’ 
fossils with vacuous interiors. The latter character, however, is 
exceptional, as the bed may be traced in the former character as far 
south as Commercy, if not further, a distance of 100 miles. It is, 
in fact, remarkable for its constancy, especially as compared with the 
beds above it; and if a division is to be made in the series in this 
district on stratigraphical grounds, it is certainly over this that 
the line must be drawn. From this rock more than 200 species 
have been recorded in this department, and 190 in the department 
of the Meuse. In such a fauna one naturally finds representatives 
of species which are abundant on several horizons; and one may 
therefore easily be misled to place it on any horizon that one hap- 
pens to be most familiar with. Two Ammonites are common, Viz. 
A. cordatus and A. Martelli, the latter being the A. plicatilis of 
dOrbigny. A form which may belong to the true A. plicatilis occurs 
in the same beds in the Meuse department ; and d’Orbigny and Buvig- 
nier both quote A. perarmatus from this horizon as well as from the 
beds below. Amongst the Gasteropoda quoted from here, Cerithium 
muricatum, Littorina muricata, Pseudomelania striata, Chemnitzra 
heddingtonensis and Bulla elongata are constantly met with in the 
lower Corallian limestones in England, the Pseudomelania being 
more characteristic of higher beds. Amongst the Lamellibranchiata, 
Gryphea gigantea, Pecten fibrosus, Lima gibbosa, Gervillia aviculo- 
ades, Avicula ca:pansa, A. ovalis, A. polyodonta (the same as our 
A. pteropernordes), Perna quadrata, Mytilus pectinatus, Cucullea 
oblonga, Opis similis, Astarte extensa, Tancredia curtansata, Iso- 
donta Deshayesia, Myacites decurtatus, and Pholadomya ovals are 
most noticeable as confirming the evidence of the Gasteropoda ; but 
Peeten vimineus points to arather higher horizon; while the presence 
of Rhynchonella Thurmanni, and of what is probably R. lacunosa, with 
abundance of Waldheimia buceulenta and Terebratula fileyensis, is 
still in harmony; and finally the echinoderms Willericrinus echinatus, 


504 J. F, BLAKE ON THE UPPER 


Echinobrissus scutatus, Holectypus depressus, and Collyrites bicordatus 
add their testimony to the great similarity which the fauna of this 
ironstone bears to the lower portions especially of the calcareous 
series of our Corallian rocks. 

This Ferruginous Oolite is referred by Hébert (21) to the middle 
Oxfordian on the ground that above it at Vieil-St.-Remy occurs a 
great thickness, 160 feet, of brown marls with a thoroughly Oxfordian 
fauna, such as Ammonites arduennensis, Trigonia perlata, Gervillia 
aviculoides, Ostrea dilatata, &e. The occurrence of any such marls is 
denied by Buvigner (22) who states that the Oolite seen below must 
be out of place. Certainly no such thickness of marl was observed 
where the beds were examined. About 40 feet of marls are seen at 
the base of the valley near Vieil-St.-Remy and in the railway-cut- 
ting near Neuvizy ; but these, from their position, are more probably 
below the Ferruginous Oolite, which is seen too near to the Corallian 
limestones on the road between Vieil-St.-Remy and Novion to allow 
of any intervening clay of more than a few feet thickness. It is 
probable therefore that the Grit and Ferruginous Oolite are hidden 
on the road to Wagnon in the 160 feet, the base of which contains the 
above fossils, while over it come the marly limestones recorded to 
contain Cidaris florigemma and Pecten articulatus. In many other 
places further south also some representative of the Coral Rag is 
found immediately overlying the Ferruginous Oolite ; so that its posi- 
tion is, in reality, perfectly fixed. 

The constancy of this bed in spite of its thinness is in striking 
contrast with the variability of the succeeding series on the horizon 
of the T'rzgonia-beds of Pickering or of the earliest limestones of the 
Corallian series, which are said by Buvignier to have nothing constant 
but their inconstancy. 

In the neighbourhood of Vieil-St.-Remy and Neuvizy the lowest 
beds referred to the Coral Rag are not well seen, though Hébert (21) 
describes about 4 feet to 5 feet of coral material with Cidaris forigemma 
on the road to Wagnon. Nowhere here, however, is there any great 
thickness of such growth ; and what there is is succeeded immediately 
by the white limestone which is seen so well at the quarries of No- 
vion. This limestone has been much acted on by chemical agents, 
many of the fossils being silicified and others represented only by 
casts. Among the latter the corals are of the greatest importance ; 
the majority are not Thamnastreean but Calamophyllian and Clado- 
phyllian. MM. Sauvage and Buvignier simply place the whole of 
these limestones, amounting, according to their estimate, to 250 feet, as 
Coral Rag, without further subdivision, and supply a rather defective 
list of fossils. At Novion a magnificent section, 100 feet in thickness, 
is seen, the Calamophyllic in places almost forming a reef at the 
base; then follow great false-bedded masses of fine white limestone, 
and towards the top huge crystalline nodules of Thamnastree. 
Lithologically therefore this might well pass as a Coral Rag; and it 
represents in this area all that would naturally be called Corallian. 
When, however, we examine it more closely, we note in the first 
place that this is scarcely a Thamnastrean reef, such as is usual in our 


JURASSIC OF THE PARIS BASIN. 505 


true English Coral Rag, the specimens at the top not being in situ— 
and also that there is an apparent absence or great rarity of Cidaris 
florigemma, the spines which occur being either smooth or belonging 
to C. Smith. At the base the most abundant fossil is a large 
Natica like N. millepunctata, Buv., but perhaps WV. grandis; while 
the topmost beds are crowded with silicified Nerinew, NV. sequana 
bring the most abundant. Other noteworthy fossils are rarer ex- 
amples of Diceras arietinum?, Cardium corallinum, Cerithium lime- 
forme, Pteroceras oceani (undistinguishable from those at Boulogne), 
Lima equilatera, Buv., and L. ornata, the Modiola wmbricata, Rom. 
(non Sow.), and Zerebratula. MM. Sauvage and Buvignier mention 
also among others Arca pectinata, Pecten articulatus, and Chemmitzra 
heddingtonensis. These characters taken together appear to indicate 
a higher horizon than the ‘ floregemma’ Rag, namely one which in 
spite of its abundant corals should be paralleled with Supracoralline 
beds elsewhere, or the upper part of theCorallian series when that has 
been divided as it has been in the Meuse. The upward succession 
is not well seen in this neighbourhood, the great limestones passing 
to large-grained oolites with undistinguishable fossils, and then to 
more compact limestones referred to the Astartian. 

The interpretation of the rocks seen in this district may now he 
checked by an examination of the country about 20 miles distant to 
the south. Starting from the town of Beaumont, and going westward, 
after passing over the Lower Oolites, the Kelloway hock and the 
Oxford Clay, which is here well developed in an argillaceous form, 
one comes, at the base of the escarpment of Stonne, to the same sandy 
nodular beds with Rhynchonella Thurmanni and Perna quadrata as at 
Neuyizy,. duly followed on the top of the hill by the Ferruginous Oolite ; 
but the bed with Trigomia spinifera was not observed. Above this, 
on the road to Ochsee, a village three miles south, the succession 
is perfectly observable on the rough cart-road mounting a barren 
hill; at the base are the Lower Calcareous Grit and the Ferruginous 
Oolite ; near the sammit are the great cavitary limestones of Novion, 
where the Calamophyllie have been, and above them flaggy Oolites 
&e. full of Nerinee ; but in the intervening space there is no sign of 
clay, all is calcareous. This portion, however, contains abundant 
Thamnastrean corals ; and with them Cidaris florigemma was easily 
found associated with Hemicidaris crenularis, Cerithium muricatum, 
and Littorina muricata ; but the Nerinee and Natice of the limestone 
above are absent. It thus appears that we are justified in regarding 
the Novion limestone as Supracoralline; while al] that can represent 
the series so well developed in England between the Lower and the 
Upper Caleareous Grit is the Ferruginous Oolite and the Rag Lime- 
stone. The upper portions of the series have not been examined 
in the Ardennes department, as the description given by the before- 
named authors indicated that they were similar to those to be seen 
in the northern part of the Meuse. 

2. The Meuse Depariment.—On entering this we come under the 
guidance of Buvignier alone (13), whose classical work is well known. 
He gives the following classification. 


506 J. F. BLAKE ON THE UPPER 


1. Upper JuRASSIC. 


I. Barrois limestone, 600 feet. 


a. Grey-green limestone. 
6. Carious limestones. 
¢, Lithographic limestones, 


II. Virgulian clays, 150 feet. 


III. Astartian limestones, 400 feet. 


a. Upper compact limestones. 
6. Lower marls. 


2. Mripp1E JurRAssIc, 
I. Coral Rag, 400 feet. 


a. Lithographic limestones. 
6. Various forms. 


II. Oxford clay, 500 feet. 


a. Ferruginous oolite. 
6. Siliceous limestones=Terrain 4 chailles. 
c. Woévre clay. 


Near the northern extremity of the department, in the neighbour- 
hood of Dun, the lower portions of the series may be easily taken 
up again; and they show a continuance of the conditions last noted 
in the Ardennes. On the slopes of the Cote St. Germain a fairly com- 
plete section may be seen: the lower half is occupied by the Oxford 
Clay, with the topmost part of the same nodular gritty character as 
before ; then may be seen the Ferruginous Oolite, of some thickness ; 
immediately over which come great crystalline irregular masses 
which, in spite of their present state, are easily recognized as remains 
of a Thamnastrean reef, with which Cidaris florigemma occurs ; 
this portion is also considerably oolitic, differing in this respect from 
the limestone that follows it, which, with its vacuous spaces once 
filled with corals and Nerineew, perfectiy represents the Novion 
stone. Here, then, we find confirmation of previous sections. Hébert 
(21) states that the Coral Rag alone is seen at the base of this hill- 
side, and that the Ferruginous Oolite must be some distance below, 
and separated by a great mass of clay. The sequence, however, is 
here perfectly clear; and on the opposite side of the valley, in the hill 
between Murvaux and Fontaines, the same succession is equally well 
seen. Only at this spot the true Coral Rag has greatly developed, as 
may be seen also by its examination in other places in the neighbour- 
hood. ‘Thus the great quarries to the south of Dun show perhaps as 
much as 60 feet, consisting of a lumachelle of broken shells at the base, 
a Thamnastrean reef with Ostrea solitarca in the middle, and oolite 
beds with Cidaris florigemma at the top, all of which must be about 
on this horizon, and which give an idea of the variability of the 
series ; while in the valley leading to Fontaines the still coral-bearing 
upper limestones are largely developed, but without the character- 
istic urchin of the lower reefs and their equivalents. 

Passing next to Verdun, the student who has traced the lower 


507 


JURASSIC OF THE PARIS BASIN. 


= ———— 


Prazncsezre 


Fig. 2.—Map of the Country round Verdun. 


i 
Se 


(Scale 1: 160,000.) 


ee 
Tater, Verdun Coté 
Ae oO ee bi 


208 J. F. BLAKE ON THE UPPER 


rocks so far has no difficulty in assigning to their true place the 
magnificent limestones which here astonish him and give him an 
idea of the grandeur of the development of the Corallian rocks in this 
district. The beds which in this locality overlie the Ferruginous 
Oolite are altogether different from any thing seen before, consisting 
at Haudainville of an unstratified mass of limestone 40 feet in thick- 
ness, made almost entirely of fragments of crinoids united by a cal- 
careous cement and yet distinct, forming a most admirable freestone : 
the only fossils seen in it are Cidaris florigemma and small oysters. 
Capping this mass in the quarry is 10 or 12 feet of Thamnastrean 
Rag with occasional Cladophyllic, but with abundance of Cidaris 
florigemma, Terebratula maltonensis, and Pecten vimineus, also Lima 
leviuscula and Venerwpis corallensis. 'The position, then, in this 
locality of the crinoid limestone is fixed; for it must be associated 
with the Rag above it, as forming part of Buvignier’s Coral Rag 0. 
The succeeding rocks, representing the Novion limestones, also intro- 
duce new features ; for here is first seen the lithographic stone which 
becomes so abundant further west. It is mixed with Oolites of 
various-sized grains; but the corals of all kinds are gone. ‘The ap- 
pearance of this group in the great quarries of St. Martin has been 
admirably described by Buvignier—its cherty bands, its vegetable 
remains towards the base, and crustaceans above. ‘The fossils are 
only abundantin parts: the commonest is Nerinea elongata, associated. 
with others, as V. Jollyana and Patella elegans; Natica globosa and 
Lucina mosensis were also collected. The upper part of this group 
is instructively seen on the north of Verdun, in the Cote St. Michel. 
The eastern portion is worked in pure lithographic limestone without 
a fossil; but the western shows great quarries of massive earthy lime- 
stones in which the characteristic fossil is Terebratula repelimana, 
d’Orb., as at Novion, but which shows its relation to the overlying 
rocks by its numerous Astarte suwpracorallina, &c. The long lists of 
fossils given by Buvignier as coming from his two divisions of the 
Coral Rag, of which the present is his a, contain many species in 
common, especially Diceras arietinum ; nevertheless there can be little 
doubt of their corresponding with beds showing more distinctness 
further north. We find therefore the same subdivision of rocks, here 
referred to the Corallian, as in the Ardennes,namely a variable mass of 
coral-bearing or crinoidal limestone below, with Cidaris florigemma, 
and a more compact and oolitic stone above, with many Nerinee, 
and an occasional Diceras. Hébert (21) thinks to divide the lower 
part into two portions, the base a shell-bed with Cidaris florigemma 
and the corals above; but the extreme variability of these rocks, and 
the occurrence of the characteristic urchin throughout all the modi- 
fications, render such a division valueless. 

In this neighbourhood the first examination of the higher: beds 
has been made along the road leading from Verdun westwards to 
Clermont-en-Argonne. Leaving the town by the fort on the north- 
west, one’s position is accurately determined by observing the great 
coral-reefs on which the foundations are built, associated with the 
gasteropod-bearing intercoralline brash, followed in the railway-cut- 


JURASSIC OF THE PARIS BASIN. 509 


ting by the massive rocks of the Nerinzan series as seen on the 
opposite side of the river. Succeeding these at once are more marly 
rocks than any seen before ; in fact they are alternations of marls and 
flagey earthy limestones, with here and there a bed of oolitic rock, 
a lumachelle of Exvogyra bruntutana, or alithographic limestone. The 
thin beds having their surfaces crowded with Astarte supracorallina, 
indicate with certainty the horizon. Rocks of this description occupy 
the country for a distance of five miles, till the lowest Virgulian 
lumachelle is reached, and seem tobe rather uninteresting and barren. 
Nevertheless Buvignier describes the lithographic stones as massive, 
and overlain by an irregular pisolite having the fossils as casts, in- 
cluding Diceras and Cardium Buvigniert, a species which appears to 
be not different from C. corallinum. This would seem to indicate a 
repetition of the Diceras-beds at a higher horizon than the first, namely 
immediately beneath the Virgulian; and the list of fossils givenasfrom 
the upper portion of the Astartian limestones contains several species 
usually of a lower horizon, such as Lima equilatera, Pecten articula- 
tus, Chemnitzia heddingtonensis, Natica globosa, &c. These pisolites 
are said to occur in the very district now under notice ; but though 
the various sections were duly searched, no such rock was observed 
in the required position. At Baleycourt quarries were seen in which 
the stone was very rubbly ; but it had the aspect entirely of a rede- 
posited material with the fossils in it remanié, and not at all like the 
true Diceras-beds of the Haute-Marne. With this exception, no 
change in the character of the rocks is observable till the lumachelles 
with Exogyra virgula are reached. If therefore any portion repre- 
senting the Pterocerian beds is to be found, it must be sought either 
in the beds containing #. virgula or in the lithographic limestones 
below. The latter are credited with an enormous thickness (400 feet), 
and obviously include much more than is usually placed in the 
Astartian beds. In point of fact, there seems little to characterize 
the beds here; and their subdivisions must be left somewhat 
doubtful. 

In the Virgulian marls which follow wehave fortunately an admir- 
able landmark, both because the abundance of the little oysters (which 
in themselves make half the bed) renders these marls immediately 
recognizable, and because the variety of their other fossils gives 
good materials for correlation. At the cutting of Nixéville the dark 
~ marls with Z. virgula occupy the lower 12 feet, and white chalky 
limestones the upper 20 feet ; neither therefore is seen in its full 
thickness. For a considerable distance to the west, alternations of 
these two forms recur, the marls being the last seen beneath the 
overlying lithographic limestone. The whole series does not appear 
to occupy so great a thickness as the lower Astartian beds. The white 
limestones are extremely fossiliferous, the most abundant shells being 
Pleuromya V olizu, Pholadomya acuticosta, Cucullea texta, and Terebra- 
tula bisuffarcinata. The other fossils observed were Péerocera Pont, 
Chenopus musca, Chemnitzia gigantea, Thracia lata, Mactromya ru- 
gosa,Ceromya eaxcentrica, Cardium Bannesianum?, Trigoma Merian 
T. Juddu, Pinna granulata, and Nucula Menkei and Astarte swpra- 


510 J. F. BLAKE ON THE UPPER 


corallina. In spite of the last-named, which is thus seen to have 
rather a wide range, the fauna is essentially Lower Kimmeridgian, 
and may be well matched in the body of that clay in England. For 
purposes of comparison in the basin of Paris itself, the Terebratula, 
Pholadomya, and Ceromya should be especially noted. Fragments 
of coronate Ammonites which might belong to A. gigas, recorded by 
Buvignier, were seen; and it may be noted that A. longispinus is 
mentioned as occurring; but there is nothing except the first-named 
to unite this to the Upper Kimmeridge of England. Hence, therefore, 
there is no probability of the overlying rocks being of Portland age, 
as they are called by Buvignier. 

In this locality the rocks which succeed the Virgulian beds at 
Dombasle are barren, lithographic, but somewhat argillaceous lime- 
stones, in remarkably uniform beds of from 8 to 12 inches thickness, 
with thin bands of intervening clay, thus presenting a well-marked 
lithological character by which they may be distinguished from the 
lower lithographic limestones. They become more rubbly towards 
the top, but, except for lumachelles of small oysters of unknown 
species, contain scarcely a single fossil. They occupy a considerable 
area, and must be nearly 100 feet thick; but even Buvignier only 
records four fossils :—Ammonites gigas, Pleuromya Voltzw, Cardium 
Dufrenoycum, and Patella suprajurensis. Nothing higher is to be 
seen in this part of the country, as the lithographic limestones ex- 
tend to the alluvial plain separating the Jurassic area from the 
cretaceous escarpment. 

We may next examine the country 25 miles further south from 
St. Mihiel and Commercy to Bar-le-Duc. The first-named is celebrated 
for the magnificent fauna of its Rag-deposits, and has been described 
separately by Buvignier. The escarpment begins about five miles 
to the west of the town, as at Apremont; but inliers close at hand 
enable us to recognize the Calcareous Grit, here still more nodular, 
with Rhynchonella Thurmanmi and Ostrea dilatata, and the overlying 
Ferruginous Oolite, retaining the same characters that it has further 
north. But the limestones which succeed these have a much more 
extensive development. At the grand section exposed on the de- 
scent to Apremont, the actual base is not seen ; but the rubbly un- 
stratified mass appears to begin at once. It is here full of unrecog- 
nizable Thamnastreean corals (which appear to have been rolled), and 
abundance of Hemicidaris crenularis, Cidaris florigemma, Aprocrinus 
Rossyanus, Terebratula maltonensis, and Pecten vimineus. There is 
a thickness of about 120 feet, all made of similar material except 
near the top, where about 16 feet is made of massive crinoidal lime- 
stone as near Verdun. ‘This latter type, therefore, as there, is sub- 
ordinate to the Coral Rag; and we hence learn the age of the magni- 
ficent freestones of this character worked about seven miles to the 
south at Lerouville, where the quarries have a 100-foot face, all of 
the same material, with occasional Cidaris florigemma, Terebratula 
insignis, and other fossils. We note also that, whereas near Verdun 
the coral-beds formed the capping to the crinoidal limestones, here 
the latter lie at the top or nearly so, and no definite position in the 


511 


JURASSIC OF THE PARIS BASIN. 


(3) = 


Chau vor cour€ psy aX 


Fig. 3.—Map of Country around St. Mihiel. (Scale 1 ; 80,000.) 


= ————_ 


wes lore, 


2 ee 


pepe 
-- 
(lor: Coble, 


LY ' ‘+, F, BLAKE ON THE UPPER 


Coral Rag can be assigned to either of these forms. Above the Coral 
Rag, on the road to St. Mihiel, are the more compact, almost litho- 
graphic limestones with Nerinee as before, occupying a wide expanse 
of country. So far, therefore, the Verdun form is fairly continued ; 
but when we examine the exposures seen in other valleys an inter- 
esting modification is observed. The deeper sections on the right 
bank of the Meuse show the base of the Coral Rag to be occupied 
by far more oolitic and regularly-bedded stone than any seen at 
Apremont; but on the left bank, quite close to St. Mihiel, a recent cut- 
ting gives a first real indication of something corresponding to our true 
Coralline Oolite. At the base of this cutting is 8 feet of Ferruginous 
Oolite crammed with fossilk—Ammonites convolutus 2, Pholadomya 
decemcostata and P. deltoidea, Mytilus pectinatus, Perna quadrata, 
Pecten articulatus, Ostrea dilatata, Rhynchonella lacunosa, Terebratula 
bucculenta, Collyrites becordatus, Dysaster ovals, and others. Next 
comes more than 6 feet of a rather sandy limestone, not exactly 
oolitic, but thick-bedded, in which no Cidaris florigemma or 
Pecten vimineus could be found, but such fossils as Ammoniies pli- 
catilis, Pseudomelania striata, Pholadomya deltoidea, Holectypus de- 
pressus, and Stomechinus sp. abound. ‘This mass is equally distin- 
euishable from the Ferruginous Oolite and from the Coral Rag, and 
occupies the place, hitherto unrepresented, of those deposits which 
in England form a basis for the latter. The thickness of this mass 
is not well seen here, and is comparatively unimportant ; but at 
Creué, 8 miles N.E. from here, Buvignier has described some ‘‘ lower 
white limestones ” occupying the same position, 250 feet in thickness ; 
and the same occur also to the south of Apremont at Liouville, and 
are therefore more or less alternative with the Rag, the great thick- 
ness of which may be due to its filling up the hollows between the 
lenticular masses of the ‘“ coralline oolite” below. These limestones 
contain a fauna very distinct from that of the Coral Rag: urchins are 
almost absent ; Myacide abound ; and the whole assemblage is much 
more like that in the Ferruginous Oolite, A. plicatilis being the most 
abundant cephalopod. It differs, however, from the fauna of the 
latter rock in the absence of the characteristic urchins, crinoids, 
brachiopoda, and belemnites. In Buvignier’s list 15 of its fossils 
are common to the Coral Rag, and 34 to the Ferruginous Oolite ; 
but the latter is only 18 per cent. of the number found in both, 
so that by any method of percentages these limestones should be 
reckoned distinct. 

Above the section already referred to, on the left bank of the 
Meuse, are about 80 feet of Coral Rag, most massive below, with old 
Thamnastree above, and containing all the usual fossils ; and on the 
ascent of the hill comes on at once a great thickness of beautiful 
pure white oolite containing scarcely a fossil, though a Corbis and. 
Diceras were noted ; all this, therefore, only continues what has been 
seen before. What lies above has not been seen; but detailed ac- 
counts of it asexposed in the cuttings between Lérouville and Loxé- 
ville are given by Hébert (21). At Vadonville is seen at the base 
a great mass of white limestones, finely oolitic, but in places almost 


JURASSIC OF THE PARIS BASIN. les 


pisolitic, stated to be 300 feet in thickness *; and above them occur 
150 feet ofoolitic beds with various-sized grains, some pisolites,nodules, 
and many specimens of Diceras. The whole of this mass is considered 
by Prof. Hébert to be above the white limestones of Novion and 
Verdun, which would place them in the Astartian of Buvignier, but 
that he considers them to be absolutely wanting to the north of St. 
Mihiel. From their position in relation to the great crinoidal 
limestones of Lérouville, however, not 1800 yards away, their lower 
portion can be nothing else than the Novion limestone; and we 
jearn that at this spot the peculiar pisolitic character of the Diceras- 
beds begins to be observable in this portion of the series towards the 
top. Itis remarkable that no mention is made of this in Buvignier’s 
work, though the spot is coloured as Corallian. The next cutting, 
27 miles te the west, is also described by Prof. Hébert. Here the 
base is occupied by the same pisolitic Diceras-beds, their continuance 
being accounted for by some folds and reversed dips observed. This 
bed is declared by Buvignier (22) to be distinct from that at Vadon- 
ville, and to belong to the pisolites he has described from the Astar- 
tian. His description, however, of the cuttingsis by no means clear 
(13), while Hébert’s is precise and definite. Above these pisolitic beds 
come a few feet of oolites, and then a mass of non-oolitic marly 
limestone with conglomerates and Ostrea deltoidea. The same series 
of cuttings reveal Virgulian beds further to the east at Loxéville, 
with many fossils common to the Astartian beds, and, finally, the 
lithographic “ Portland ” + limestones (see 21). This is well-deve- 
loped in the neighbourhood of Bar-le-Duc, forming the main mass of 
the hills. The carious limestones above are well-marked, with their 
hollow tubes filled with a brown earth. This is supposed by Tom- 
beck to be the original material, and the limestone to have formed 
afterwards ; but this can scarcely be the case, as there are in this dis- 
trict many fossils in the limestone with their interiors formed of the 
matrix but the shell gone. The assemblage of these appears to in- 
dicate very shallow-water conditions, as though here one were at the 
natural boundary of the deposit; they are mostly small and simple. 
Those found are Alaria dionysea, Mactromya rugosa, Plectomya 
rugosa, Lucina aspernata, Corbicella Morcana, Lithodomus vietus, 
and Cardium pesolinum?. We have very little assistance here to- 
wards correlation with our English rocks, and none towards placing 
these on a level with the Portland Limestone. 

The highest beds seen here are a variable series of rather dirty- 
coloured stones with sands &c.; but one remarkable rock requires 
special notice. It is the ‘‘ vacuolar oolite,” in which, so to speak, 
there are no oolitic granules, though once there were; each granule 
has been dissolved, the intervening matrix alone remaining to form 
a spongy rock. This becomes in places almost a lumachelle of Corbula 


- * The thicknesses given by Prof. Hébert are always very large when estimated, 
and require to be somewhat reduced to bring them into comparison with the 
estimates of other geologists. 

+ The names used by the describers are adopted until the amended nomencla- 
ture is proposed at the end of this paper. 


mes. G.S. No. 148. 2u 


514 J. F. BLAKE ON THE UPPER 


mosensis ; and the next abundant shell is Astarte rugosa. Buvignier 

records also Trigona gibbosa, but with doubt. These beds may very 

well represent the true Portland Limestone, which is also character- 
ized by Astarte rugosa; but this question will be decided later on. 

One more traverse completes 

Fig. 4.—Map of Country be- ourstudy of the Meuse department. 

tween the Meuse and Ornaim. This is about 25 miles to the 

(Scale 1 : 80,000.) south, but runs N.W. to 8.E., 

7 the general trend of the strata 
changing near the extremity of 
the department to be N.E. and 
S.W. instead of N. and 8. The 
whole series spreads about 84 
miles on either side of Gondre- 
court, actually commencing within 
the Vosges department. 

The lowest beds are exposed 
at Greux, on the Meuse; and we 
still see the nodular calcareous 
grits at the top of the Oxford Clay, 
here more approaching what could 
be called a “ terrain-a-chailles ” 
than anywhere else, and still cha- 
racterized by Lthynchonella Thur- 
manni. No sign, however, of the 
== 7. 6 |  Ferruginous Oolite was observed ; 
=e OU | a vac Buvignier does not place it 

eae See) here on his map. It has therefore 

: probably died out; for it is not 
seen again to the west. Neither is 
any thing like the Creué Limestone 
or “Coralline Oolite” seen; but 
the great mass of Coral Rag im- 
mediately succeeds. This consists 
of a rubbly rock, in places oolitic, 
without any corals, yet of that pe- 
culiar appearance which indicates 
the beds associated with them; 


= tase and all the usual fossils abound, 

ae SS (Liecers such as Cidaris florigemma, Hemi- 

= Zeomest) cidaris crenularis, Mytilus angula- 

= a tus (jurensis), &c. It is succeeded 

SS iN He nay f (Cigrky by a white chalky limestone with 
| aa \e deine Ok Lemes numerous hollow branches as of 
| Xd = Cora ie: d decayed corals, such as are seen 
== ; at Novion. Some beds are flaggy ; 

PALL. \. but all very barren of fossils. 

Greual | f Above comes 12 feet of strong beds 


’ 
Kay 
‘ A 
Domremy 3. \ Likleey g 


of oolitic and non-oolitic limestones, 
which are worked for building- 


JURASSIC OF THE PARIS BASIN. 515 


stone when thickened out in the neighbourhood; and, finally, coarse- 
grained thin-splitting oolites occupy the summit of the hill. There 
is thus no proof of our having here reached the top of the limestones ; 
and yet these are at least 300 feet above the Oxford Grit, all of 
which must be placed in the Coral Rag and Supracoralline beds. 
The continuation of the route, however, reads us a very instructive 
lesson. On consulting Buvignier’s detailed map it will be seen that 
behind this hill there is a long tongue of the lower rocks crossing 
the road in the valley. Little is to be seen on the descent of the 
hill; but on the next rise at Vouthon Bas, the nodular Oxford Grit 
is seen in the village, and above this successively a shelly limestone 
full of Cidaris florigemma, massive oolitic rocks, thin-bedded oolitic 
rocks, and then the lithographic lmestone—in a word the same 
succession as before, but in this instance crowded into a space which 
certainly cannot exceed 50 feet. So great a change in thickness in a 
distance of less than 3 miles gives an idea of the very local and 
fragmentary character of the Corallian rocks. 

The greater part of the distance to Gondrecourt, about 5 miles, 
is occupied by compact, almost lithographic limestones, with no 
fossils visible. ‘These appear to be placed by Buvignier, from the 
colouring of his map, as Astartian. They nevertheless occupy the 
position of the great masses to the north of Verdun, and perhaps 
also east of St. Mihiel, which are coloured Corallian; but in the 
absence of fossils it is difficult to say to which they should be joined. 
In this traverse, as in the others, but little sign is seen of the piso- 
litic Diceras-beds ; only some coarse-grained oolites are seen. Hence 
the peculiar character so well seen further west is here only occa- 
sional, or occupies too small a space to be easily discovered. 

Near the entrance to Gondrecourt these lithographic limestones 
are succeeded by and intermixed with rubbly beds with many 
fossils. A thickness of about 40 feet is seen, towards the top of 
which Terebratula subsella is abundant. Similar stone is continued 
for nearly a mile on the other side of Gondrecourt, and gives a very 
good idea of the Astartian beds in this district. The fossils here 
noted were Terebratula sp. (perhaps the young of 7’. subsella), 
Pterocera Thirrie, Thraca lata, Pholadomya Prote, Isocardia obovata, 
Ceromya excentrica, Pleuromya rugosa, Mytilus opisoides?, and 
Nautilus inflatus. Exogyra virgula is not seen in these beds, which 
are almost as argillaceous as the overlying series, especially if we 
include in them all that lies below the first undoubtedly Virgu- 
lian bed; but no pisolitic beds are seen along the road. The base 
of the Virgulian occurs at Houdelaincourt, about 3 miles north of 
Gondrecourt, where a mass of chalky limestones forms a bank 40 feet 
in height, the upper part of which yields all the fauna of Nixéville, e.g. 
Ammomtes longispinus, Pholadomya multicostata, P. Prote:?, Astarte 
supracorallina, Pinna granulata, and Pecten suprajurensis. Above this 
come the marls full of Hvogyra virgula, then more chalky lime- 
stones with Pleuromya Voltzi, and finally more marls and marly 
limestones such as are seen in a cutting further on. This cutting, 
made for the Marne-and-Rhine canal at Demanges, has been de- 

2M 2 


516 J. F, BLAKE ON THE UPPER 


scribed and drawn by Buvignier. It shows well how argillaceous the 
Kimmeridgian is here, being composed of quite as much clay as stone, 
which is an exception to its usual character in France. About 40 ft. 
is exposed; but this does not include any lumachelles of Haogyra 
virgula, though that oyster occurs with others. The nodular bands 
are very fossiliferous, and include more stony beds at the base, with 
Pholadomya multcostata, Gervilha kimmeridiensis, and Cucullea 
texta, and in more marly beds . terocera mosensis, Trigonia Voltzi, 
T. concentrica, Astarte scalaris?, Pleuromya Voltzu, P. donacina ?, 
Pecten suprajurensis, and Terebratula subsella. 

This description of the beds which intervene between the Corallian 
and ‘‘ Portland” limestones, though doubtless indicating that some 
division might be adopted, does not tend to magnify the importance, 
in this district at least, of the three groups—Astartian, Pterocerian, 
and Virgulian, all of which would be included in England in the body 
of the Lower Kimmeridge. There is a community of fauna which 
shows them all to be too intimately united for any wide separation. 
Above the fossiliferous Virgulian marls of the Demanges cutting 
comes immediately the unfossiliferous lithographic limestone of the 
so-called “‘ Portland” series, occupying all the summits of the hills; 
but no special observations were here made. 

3. The Haute-Marne Depariment.—The development of the Upper 
Jurassic rocks in this department has received more study and raised 
more discussions than that in any other place. It is consequently con- 
sidered by many the typical region in the Paris basin, and is used as 
a term of comparison in every attempt at correlation. Lying opposite 
the strait which connects the basin of Paris with the Jura, its lower 
part appears to partake of the character of the Swiss rocks ; and its 
upper part is more complete than anywhere else in the same range. 
Although it is therefore of the highest importance that its rocks 
should be correctly described and interpreted, the difficulties of its 
natural situation appear to have been increased by its illustrators, 
and one would suppose from their writings that it was of most ano- 
malous structure. The earliest description was given by Royer (4). 
Being the simplest, this was in some respects the best; and it will 
be well to quote it for comparison with the most recent and com- 
plicated. Royer divides the series as follows :— 


A. Portland. Oo.iTE or BaRrols. 


a. Carious limestones. 
6. Nodular compact limestones. 
c. Lithographic limestones. 
. Kimmeridgian. 
. Astartian. 
a. Nodular and oolitic beds. 
b. Compact limestone. 
. Coralline oolite. 
. Compact Corallian limestone. 
a. Thick limestone. 
b. Marly limestones passing to Oxfordian ; 6’. Rubbly coral limestone. 
. Upper Oxfordian marls, almost without fossils. 
. Middle Oxfordian. 
. Lower Oxfordian. 


Eo Qh 


FY C2 te 


JURASSIC OF THE PARIS BASIN. 7 


Various changes have since been made in this classification, partly 
by way of expansion, but partly by way of alteration; and an 
almost interminable series of Notes by M. Tombeck have led to the 
development of a classification which is exceedingly complicated, and 
which in several important points it seems impossible to accept. 
As published by MM. De Loriol, Royer, and Tombeck (50), and 
modified by the last named (5d), it is as follows :— 


PoRTLANDIAN. 


1. Zoue of Cyrena rugosa. 
a. Upper grey-green limestones. 
6. Vacuolar oolite. 
c. Lower grey-green limestones. 
. Zone of Cyprina Brongniarti. 
a. Tubulous limestones. 
6. Spotted limestones. 
¢. Carious limestones. 
3. Zone of Ammonites gigas. 
a. Bure oolite. 
6. Limestone with Amm. irius. 
¢. Marls with Hemicidaris purbeckensis. 
d. Lithographic limestone with Amm. rotundus. 


bo 


KIMMERIDGIAN, 


1. Zone of Amm. caletanus, = Virgulian. 
a, Alternations with Amm. erinus. 
6. Marls with Amm. eumelus. 
2. Zone of Amm. orthocera, = Pterocerian. 
a. Perforated limestones. 
b. Marly limestones with Dysaster granulosus. 
¢. Limestone with Jsocardia striata. 
d. Marls with Rhabdocidaris Orbignyana. 
e. Marls with Ceromya excentrica. 
Jf. Perforated limestone with Pteroceras. 


CoRALLIAN OB SEQUANIAN. 


1. Second zone of Zerebratula humeralis. 
Astartian limestone. - 
2. Second zone of Cardium corallinum. 
La-Mothe oolite. 
3. First zone of Zerebratula humeralis. 
a. Lithographic limestones. 
6. Saucourt oolite. 
ce. Limestones with Nautilus giganteus. 
d. Upper rubbly limestone with Cidaris florigemma. 
é. Limestone with Amm. achilles. 
f. Limestone with Amm. bimammatus. 
4. First zone of Cardium corallinum, and zone of Hemicidaris 
crenularis. 
a. Upper maris without fossils. 
b. Diceras-oolites, and lower rubbly limestenes with Cidaris 


florigemma. 


¢. Lower marls without fossils. 


OXFORDIAN. 


Zone of Amm. transversarius. 
a. Beds with Amm. Henrici and Amm. oculatus. 
6. Beds with Amm. Babeanus. 
¢. Beds with Amm. Martellé. 


518 J. F. BLAKE ON THE UPPER 


This is an exceedingly complicated classification ; but it 1s neces- 
sary to give it in full, because points of discussion arise depending 
on its details. According to the descriptions given by Tombeck, all 
kinds of extraordinary developments of the Corallian rocks take 
place in this district, and many differences are observable from any 


Fig. 5.—Map of Part of the 
Valley of the Rognon. 
(Scale 1 : 80,000.) 


thing found elsewhere; but though 
it may seem presumptuous to found 
any opinion upon a rapid survey, yet 
that survey, directed to the special 
examination of crucial spots, fails to 
reveal any thing very abnormal, 
though there are considerable differ- 
ences when the rocks are compared 


- with those already studied. In point 


of fact, an examination of the locality 
fails to comfirm M. Tombeck’s con- 
clusions in several important points. 
The chief point necessary to esta- 
blish is the true succession of the 
rocks which lie between the Virgu- 
lan and the Oxfordian beds. By 
the aid of a personal examination of 
the spots, aided by the definite facts 
recorded, when rightly interpreted, 
one may come to a pretty sound con- 
clusion. 
« This portion of the series may be 


- first examined in the valley of the 


Rognon. Passing down this valley 
from Andelot towards Donjeux, the 
Corallian rocks, in the form of rubbly 
limestones with Cidaris florigemma, 
are first seen on the left bank of the 
stream at Roche-sur-Rognon. Their 
base is not here well exposed; but 
is stated (19) to consist, in the neigh- 
bourhood, of disaggregated oolites 
with Glypticus hieroglyphicus, over- 
jain by. white coral-limestone. On 
the opposite. side of the stream the 
rubbly - limestones are themselves 
absent, their place being taken at 
Cultra by compact Oxfordian lime- 
stones, similar to those which occupy 
so much of the ground below them 
(by general dip) on the left bank. 
This is doubtless due to a fault; 
though Tombeck (65) actually thinks 
the rubbly limestones 100 feet thick 
have changed into this compact well- 


JURASSIC OF THE PARIS BASIN. 519 


stratified form at a distance of 500 yards. A second cross fault 
cuts off Cultra from the cliff of Bettaincourt, where the great 
Diceras-beds occupy the ground down to the stream. ‘The relation, 
therefore, of the two parts of the Corallian rocks is not here so clear 
as could be wished; but some portion of the Diceras-beds are seen 
to be lying on the top of the rubbly limestones further back at 
Roche-sur-Rognon; and at Reynel, about 3 miles to the east, Tom- 
beck (55) describes an admirable section, having at the base Ox- 
fordian marls with O. dilatata, then grey marls with Cidaris flori- 
gemma and Hemicidaris crenularis, next a mass of coral limestone 
with corals in situ, gradually becoming oolitic and then pisolitic as at 
Bettaincourt, the whole being 300 feet in thickness. Notwithstand- 
ing this section, however, he thinks (65) that the one form changes 
into the other. 

The Diceras-beds at Bettaincourt, first seen here in a journey 
westwards in their full development, are marvellous deposits. They 
consist of rolled fragments of all sizes, coated with calcareous cement, 
and which in one sense may therefore be called oolitic grains; but they 
have been subjected to a very different variety of the process which 
produced the concentric coats of the latter. In oolites the process 
is tranquil, in these tumultuous and rapid, the whole presenting the 
appearance of a substratified heap of rubbish from a neighbouring 
centre of life, borne along by the ancient currents which swept 
through the straits of Dijon. Such beds might be supposed to re- 
place, with almost any rapidity, the coral-growths from which they 
are derived, or to die out entirely. There are, however, great 
varieties in the Diceras-beds themselves—pisolites, large-grained 
oolites, suboolitic, and thin-splitting limestone alternating; and 
various groups of fossils are found in the various parts; but yet 
the whole is indivisible. With many Dicerata occur Cidaris flori- 
gemma, Cardium corallimum, and many rolled sponges. The other 
fossils noted were Nerinea sequana and other species, Littorina 
muricata, Isocardia striata, Corbis gigantea, Lima leviuscula, L. 
pectimformis, Trichites sp., Ostrea pulligera, and Terebratula mal- 
tonensis. What the real thickness of these beds may be it is im- 
possible to say; their stratification is so irregular, and their change 
of character in a lateral direction so marked, that all estimates are 
worthless. Perhaps 300 feet may be seen in the Bettaincourt hill-side if 
there is no real dip ; but even more may be introduced in the five miles 
over which the deposit spreads. It is, of course, easy (50) to pick 
out one of the oolitic portions, and call it the “ oolithe de Saucourt,” 
and another above this, and call it the “ oolithe de La Mothe;” but 
the actual oolites which occur at those places are not thereby proved 
to be superimposed, unless they are proved identical with these 
beds, which has not been, and cannot be done. The whole forms 
one great mass (here at its maximum of development), representing 
on the one hand the Novion and Vadonville limestones, and on the 
other the whole mass of the Diceras-beds further to the west. The 
upper portion is certainly more compact than the lower. On reach- 
ing Saucourt the peculiar character of the Diceras-beds is lost; and 


520 J. F. BLAKE ON THE UPPER 


on ascending the hill we find the oolites above worked in a 12-foot 
face, and overlain by a shelly limestone crowded with Rhynchonella 
pinguis; this is followed by more compact limestone, some portion 
of which may possibly be slightly oolitic; but there is nothing re- 
markable to distinguish the whole series, nor is its relation to the 
Diceras-beds below very clear. 

Succeeding these compact limestones comes no great thickness of 
irregular limestones, such as have been seen to characterize the 
Astartian beds near Gondrecourt ; and on the top of the hill follow 
the marly beds with Hxogyra virgula im lumachelles, and Phola- 
domya acuticosta. If any Pterocerian is to be made out here, it 
must be by a very close examination of fossils, which are not by any 
means abundant. 

At the forges of Donjeux, about a mile further on, is a quarry in 
which the oolite of La Mothe is supposed to be seen. Here the top 
beds consist of the irregular Astartian limestones; and the base of 
the quarry is worked for massive rocks, which are on the whole 
compact, varying on the one hand to lithographic stones, or on the 
other developing a fair proportion of oolitic grains, but totally un- 
like the Diceras-beds, because like any other semioolitie rock. Here, 
then, is no repetition; and on the whole, with the exception of 
the remarkable Diceras-beds, the succession in this valley is very 
similar to, and certainly consonant with, what we saw in the last 
section in the Meuse department. The chief noticeable point is that 
the peculiar pisolitic character is exhibited rather at the base than 
towards the top of the mass. 


Fig. 6.—Map of Part of the Marne Valley. (Scale 1: 80,000.) 


NY ty : cs 

y Buxr xere8//1/ of fH e' 4 a 
i j (Conf. z) Us, Bl eens) > 
1a bal Be e 


ee 
psiheem fet pb. L, 
ISG (erga) eds) 


Favin a 


2 Joe ers 
o fre TAN ay a fptee care) €or. ool} 


Bam HS Opforaisn) _| 
Now let us pass to the valley of the Marne. A complete section 
of the lower portion may be obtained by following the right bank 
of the river from Vouécourt to Buxicres, and then mounting the 


JURASSIC OF THE PARIS BASIN. 521 


hill-road. The base of the hill called “ Les Lavicres ” is formed by 
well-exposed sandy marls and flaggy beds, which are very poorly 
fossiliferous, and appear to represent the ‘‘ marls without fossils,” 
the only fossils obtained being Amm. marantianus and Area rhom- 
boidalis, the latter a fossil of wide range. The amount of this 
material must be considerable; in fact, in the neighbouring cutting 
of Buxieres about 50 feet is seen. Above these marls, on the road 
running north, are quarries of rock somewhat slipped out of place, 
but consisting of solid blocks of true oolites with Amm. plicatilis— 
a very good representative of a “coralline oolite.” Soon a river- 
cliff is reached, by the side of which the road runs, and we have 
a continuous section with scarcely a break. First there is a shell- 
limestone, in parts oolitic, containing Chemnitzia heddingtonensis, 
Neritopsis Guerrei, Cerithium mornatum, Nerinea sp., Trigona Etal- 
lon ?, Lama leviuscula, Pecten articulatus, Ostrea solitaria, Rhyncho- 
nella corallina?, R. pinguis, and Terebratula rotundata?. There are 
also numerous corals; in fact, in places towards the upper part the 
rock becomes a complete reef; but the corals are not commonly 
Thamnastrean, and the urchin is C. Smthit and not C. florigemma. 
Nevertheless these coral-growths must represent the Rag ; for, traced 
upwards, the rocks become more oolitic, then pisolitic, and finally, 
without our being able to draw any line, we are fairly launched into 
the characteristic Diceras-beds with all their great masses of rubble, 
their Diceras and Cardium corallimum. The whole of these beds 
constitute a magnificent assemblage of limestones which can scarcely 
be less than 300 feet in thickness. There is no change here in a 
horizontal direction, as stated by Tombeck. The beds dip slightly to 
the north, which brings higher beds to the same level; but the 
Diceras-beds are not met with till we begin to mount; it is there- 
fore, on the contrary, a vertical succession. The beds which overlie 
the Diceras-beds here gradually lose their pisolitic character and 
become marly and rubbly, but are often compacted into solid blocks. 
The fossils observed in these beds were Pholadomya Protec, Ceromya 
striata, Mytilus perplicatus, Trichites sp., Ostrea gregaria, and Tere- 
bratula Leymerw. These beds represent, apparently, the subdivisions 
C to F of Tombeck’s first zone of Ter. humeralis, and have a thickness 
of about 60 feet. They are here followed by a somewhat more oolitic 
block capped with a bed of Rhynchonella pinguis, which may well 
be the continuation of the block quarried at Saucourt. Above this 
come compact, nearly lithographic limestones, which occupy all 
the summits of the hills. The whole succession is here therefore 
perfectly clear, and it is singularly ike what has been recorded as 
seen at Reynel. 

There is nothing in all this to indicate any thing abnormal, fur- 
ther than the characteristic inconstancy of the Corallian recks. The 
Oxfordian strata, however, appear to have put on a different facies. 
There is here no Calcareous Grit with characteristic fossils, nor any 
Ferruginous Oolite ; and the species of Ammonites are not those which 
are common to the east. Further down in the Oxford clay the zones 
appear to be continuous, and hence the new form of marls with 


o22 J. F. BLAKE ON THE UPPER 


Ammonites marantianus &c. must have taken the place of the upper- 
most portion. 

On account, however, of the many questions which have been 
raised about the Haute Marne, taken by the French geologists as 
their typical area of Corallian rocks, it is needful not only to make 
out the succession from the rocks themselves, but also to discuss the 
evidence brought forward in support of a different and more complex 
reading. The peculiar changes supposed to be here effected in the 
relations of the recognized Corallian rocks and those hitherto con- 
sidered Oxfordian, and the generalizations founded on this, have been 
developed by degrees in numerous papers by M. Tombeck (45, 48, 50, 
55, 57, 58, 64, 65, 66). 

The ‘“ Marls without fossils,” stated by M. Royer to underlie the 
Corallian, were first made to be their lateral equivalents (55), and 
afterwards (64) were divided into two parts ; and the whole Corallian 
mass, when that occurred, was inserted between the two, which unite 
into one in its absence. Hence the fossils of these barren marls, espe- 
cially Amm. marantianus, were considered Corallian; and wherever 
that species is found we are to look for the Coral-beds and Pisolites 
beneath it. Hence, too, the beds with Amm. tenwlobatus, which are 
said to come above those with Amm. marantianus, are to be placed in 
the Astartian instead of the Oxfordian. Now, whatever evidence 
there may be elsewhere to show the true position of these zones, the 
only evidence obtainable from the Haute Marne shows that the first 
zone lies below the Corallian. 

The idea that the Corallian rocks change into marls depends on a 
comparison of the series in the valley of the Marne with that in the 
neighbouring valley of the Aube, and will be discussed later. The 
newer idea that they are to be intercalated is supposed to be proved 
in the valley of the Marne itself. 

The river-cliff before mentioned, which skirts the right bank of 
the Marne from Les Laviéres to Buxiéres, forms a semicircle of rocks 
haying a fairly constant dip to the north. The radius of this semi- 
circle is 1500 yards; and near the centre is a low hill, through which 
the railway passes in a cutting at a distance of only 1000 yards 
from the cliff. In this cutting it is said that the Diceras-beds, 300 
feet thick at 1000 yards distance, have died out to a thin wedge, 
and may be seen lying in the midst of the marls(65) and the under- 
lying rubbly limestones at the base. Neither of these statements 
can be accepted. ‘The marls at the cutting are the continuation of 
those of Les Laviéres, with exactly the same character: there are hard 
bands in them; but these are not Corallian ; and the top of the cut- 
ting on either side is covered by the shelly limestones of the cliff 
and their fragments, with corals and Pecten articulatus &e. Here, 
therefore, the Oxfordian marls are seen to underlie the Corallian 
limestones. The Corallian limestones lie very slightly unconform- 
ably on the marls; so there may be something wanting here. 

Another locality supposed to prove the same is at Vouécourt, 
two miles to the south (62). Above this village is a ravine leading 
east (Ravin du Heu), and a road sloping up the hillside to Viéville 


JURASSIC OF THE PARIS BASIN. ae 


on the south (Cote du Noeulon). In the first the marly limestone 
“4 A. Holbeim et A. marantianus” is said to be seen above the 
Diceras-beds ; and in the second the Diceras-beds are said to be re- 
duced tv almost zero in the midst of the clays. If “A. marantianus 
is found” (in beds which lie?) “‘ above 50 metres of Diceras-beds at 
Baxicres,” and (in beds which le?) “above 30 metres of the same 
at Heu,” still one swallow does not make a summer, and the rare 
occurrence of such a fossil would no more make the zone of Amm. 
marantranus Corallian than the occurrence of species so near to Amm. 
cordatus that they cannot be distinguished in Supracoralline or even 
Kimmeridgian beds makes the zone of Amm. cordatus a part of the 
Kimmeridge Clay. The actual section seen at Vouécourt is quite 
consonant with all that has been seen before, except that in the 
Ravin du Heu the lower part of the section is much distorted, and 
the Corallian beds on the north appear to be let down, as by a fault, 
to too low a level in relation to the Oxfordian on the south. The 
section commences at the top with compact limestones, as at Buxieres; 
then, below, come rubbly lhmestones with abundant Terebratula 
Leymern, T. tetragona, &e., also Rhynchonella pinguis, Plewromya 
Volizu, Anatina magnifica, Mytilus perplicatus, Pinna granulata, Pec- 
ten inequicostatus, and Nerita Royert. Next follow marls and marly 
flags containing the same Brachiopoda, Mytilus, and Pecten, with 
Arca rhomboidalis and Pholadomya cor? ‘These are the beds which 
contain Amm. marantianus, according to Tombeck ; if they do, it has 
not its usual associates, and is therefore out of its place. Below 
comes a great mass of limestone capped by a rosy large-grained 
oolite ; but the base is not seen, the whole being so disordered. In 
these were seen Lima leviuscula, Pecten articulatus, and Rhyncho- 
nella corallina. No doubt the Diceras-beds do not here make such 
a show; but the succession above shows that we are at their top, 
and there may be much more than appears. It is below all this 
that the whole road along the Noeulon is cut; and it shows only 
the marls and marly limestones of the Les-Lavicres and Buxiéres 
cuttings. 

All the other sections in and near the Marne valley quoted by 
Royer or Tombeck, and those seen by the Geological Society of 
France on their visit in 1856 (19), are perfectly consonant with the 
present reading, and may be easily interpreted. 

To complete the account of these lower beds, as developed in the 
department of the Haute Marne, we must make a digression towards 
the west to Maranville and La Mothe. At the first locality on the left 
bank of the Aube we are supposed to recognize the Corallian in the 
midst of the marls; there is, however, nothing Corallian about them. 
The rocks seen in the hills here are a series of marly limestones far 
more argillaceous than any seen before, having a thickness of 120 
feet. They are most lithographic above, and most marly below, the 
intervening portion being most fossiliferous. The fossils seen are Ox- 
fordian, viz. Alaria bispinosa, Astarte striatocostata, Littorina Meriani, 
Ostrea multiformis, Exogyra spiralis. There is nothing, therefore, 
here to identify any portion as having a position above the Diceras- 


a24 J. F. BLAKE ON THE UPPER 


beds ; but the whole corresponds to the marls below. Though nothing 
is seen above these Oxfordian rocks on the left bank of the Aube, 
the whole country on the right bank to near La Mothe is occupied 
by limestone rocks very similar in character to those at Buxiéres, 
though only occasionally observable. At La Mothe itself the most 
characteristic rock is exactly of the Doulaincourt and Buxieres type, 
and totally unlike any thing called La-Mothe Oolite elsewhere ; 
beneath it is a thick mass of large-grained oolite (perhaps 30 feet), 
worked for building-stone at Curmont, below which here, in an exca- 
vation, are seen the shelly limestones as at Buxieres. The succession 
is exactly the same, and the fossils similar. 

In Royer’s first description (4) these Diceras-beds were placed on 
the same level; and it was after his visit to the valley of the Yonne, 
where two Oolites are developed, that he endeavoured to find two 
also in the Haute Marne. The Oolite at La Mothe lies on a mass of 
compact limestone, which he traced to Soncourt, and there found 
rubbly beds at the base. These rubbly beds he apparently took 
for the thinned representative of the whole of the Corallian. 
Numerous sections seen and described show that (as at Buxieres) 
the base of the Corallian is rubbly and Coralliferous ; but it has 
never been proved that the Oolite of La Mothe overlies any such 
beds as the rubbly and oolitic limestones which cap the Diceras- 
beds at Buxieres; on the contrary, such beds, with their usual 
Brachiopoda, are seen above the Oolite in the neighbourhood of La 
Mothe ; and these are followed by compact, almost hthographic lime- 
stones; but whether immediately or not cannot be certainly made 
out. Above these come the irregular limestones which usually in- 
dicate the Astartian zone; and on the hill of Colombey-les-deux- 
Eglises are the marls full of Ewogyra virgula, the whole being very 
parallel to what is seen above Saucourt. The difference in elevation 
between Curmont and Colombey is about 150 feet. 

On the undisputed Astartian and Virgulian of the Haute Marne 
no special observations have been made; but it may be noted that 
the Geological Society of France found in the rubbly limestones of Don- 
jeux, associated with several Merinwe, Natica hemispherica, N. turbr- 
niformis, Pholadomya Prote, Plectomya rugosa, Ceromya eaxcentrica, 
C.obovata, Hxogyra virgula, Ostrea solitaria, Rhynchonella pingus(?), 
Terebratula subsella, T. Leymeri, and Holectypus corallinus. From 
Joinville northwards we have the most complete development within 
the basin of the “ Portland” rocks. At that town great masses of 
lithographic limestones are quarried, all with the peculiar clay bands 
dividing them into thin blocks, and containing virgulian lumachelles 
here and there. They have a great thickness, and are very unfos- 
siliferous; there are some alternations of marls and earthy lime- 
stones. and a block of oolite at what may be conveniently called the 
top. The Ammonites that have been found in them—A. gigas, A. 
suprajurensis, and A. rotundus, together with Trigoma Pellati, T. 
Cottaldr, Cardium Foucardi, Myacites jurassi, Pecten suprajurensis, 
with others recorded by Tombeck, are sufficient to correlate these 
limestones with what has been known elsewhere as Lower Portlan- 


JURASSIC OF THE PARIS BASIN. 525 


dian, though the facies is too different to permit us to compare it 
directly with the portion of the English Kimmeridge Clay corre- 
sponding to it. The next portion of the series, called by Tombeck 
the zone of Cyprina Brongniarti, consists of three subdivisions, each 
of which has sufficiently marked lithological characters to be recog- 
nized without fossils; and they are admirably shown to the north of 
Joinville. The thickness of this portion is estimated at 160 feet, and 
it is far more fossiliferous than the lower group, as we have seen it 
to be near Bar-le-Duc. Still the line between the two may be con- 
sidered arbitrary, and very probably does not correspond to the 
subdivisions in the Department of the Yonne. Pinna suprajurensis, 
after which the series is named in that locality, occurs here in 
abundance in some beds above the Bure Oolite, which are not cari- 
ous. The whole is well seen above Rachécourt, as Tombeck says. 
The chief fossils observed in this series are Cyprina Brongnarti, 
Mytilus waunensis, Anatina incequilatera, Corbula mosensis, Cardium 
Dufrenoycum, Cyprina implicata, Pecten nudus. These beds are 
here better developed than in the Meuse, and yield more data for 
comparison. ‘They are succeeded by a small thickness of unfossili- 
ferous, dirty-coloured stones, which form the possible representative 
of the “‘ Middle Portland” of Boulogne. 

Above all these rocks, and seen in several places to continue 
without discordance into them, are the remarkable beds placed as 
the “ Zone of Cyrena rugosa.” They partly consist of grey flags, 
sparkling with minute crystals and crowded with Corbula? inflewa, 
and partly of vacuolar oolite, worked in thick beds free from fossils, 
having here and there bands of shells, the principal of which are 
Astarte rugosa, Corbula inflewa, and <Avicula rhomboidalis. The 
beds have the appearance of having been formed in shallow water. 
These rocks, of which the 16 feet worked at Chevillon may be a 
maximum, appear to be separated from all below them, both in 
character and the greater number of the fossils; and under these 
circumstances, perhaps, the single shell (Astarte rugosa) which they 
have in common may suffice for a bond of union with the English 
Portland rocks. The development here is extremely restricted, these 
rocks not having a range of more than 30 miles. 

4. Department of the Aube.—We here come under the guidance ot 
Leymerie (7), whose work, though now old, is admirable; and very 
little attention appears to have been paid to the department since 
his time. His classification of the rocks is as follows :— 


Upper JURASSIC. 
1. Portland limestone, 330 feet. 
2. Marls and limestones, with Hxogyra virgula, 250 feet. 


MIDDLE JURASSIC. 


1. Astartian limestone, 320 feet. 
2. Nodular white limestones= Coral Rag, 40 feet. 
3. Lower Coral limestone. 

a. Compact limestone, 80-100 feet. 

b. “ Levique” limestone, 50 feet. 

e. Oolitic shell-limestone, 80-100 feet. 


Fig. 7.—Map of the Valley of the Aube, near Bar. 


526 J. F. BLAKE ON THE UPPER 


Our first traverse is down the valley 
of the Aube, on either side of Bar-sur- 
Aube. At Clairvaux, not very far north 
of Maranville, near the river, there 
BY fae” are great cement-works, where cement- 
We Sai const 


tema stones, more or less sandy, and inter- 
\\ eles spersed with marly beds, are worked over 
os a thickness of 40 feet or more. These 
contain abundant fossils ; the chief noted 
were Cerathium muricatum, Pholadomya 
canaliculata, P. cngulata, Trigona per- 
lata, Anatina magnifica, Goniomya sul- 
cata, Pinna lanceolata, Gervillia avicu- 
loides, and Exogyra spiralis. There really 
cannot be very much doubt whereabouts 
we are in the series here; it must be 
the top of the Oxfordian, somewhere on 
the level of the Lower Calcareous Grit. 
These, in fact, appear to be the beds that 
are missing a little further to the east 
(at Buxieres), and probably on a higher 
horizon than those at Maranville. In 
any case the fauna is perfectly distinct 
from that of the somewhat marly lme- 
stones which le above the Diceras-beds 
at Vouécourt. In Leymerie’s lists no 
such assemblage is noticed ; and it would 
seem that these beds were not open to 
his observation. 

The immediately overlying rocks are 
not well observable here; but there ap- 
pears no sign of rubbly Coral Rag or of 
Diceras-beds, in place of which the 
marly beds become more sandy, then 
more solid, and finally change into a 
massive limestone. ‘This is not to he 
wondered at, when we observe the great 
reduction the Diceras-beds have suffered 
between Buxicres and La Mothe. We 
are here introduced, in fact, to the fea- 
ture which characterizes much of the 
western extension of the Upper Jurassic 
rocks: they tend more and more to be- 
come a uniform mass in which recog- 
nizable rocks of peculiar constitution 
are mere occasional occurrences. The 
solidity of the limestones in this dis- 
trict is well shown in a quarry-face 
of 60 feet, the base of which is reached 
by a continual ascent through a stony 


JURASSIC OF THE PARIS BASIN. DOT 


ravine; and here, high above the cement-stones, perhaps 200 feet, 
we find a fossiliferous zone, similar to that at Saucourt, belonging 
to the compact limestones above the Diceras-beds. This contains 
Arca lineolata, Mytilus longevus?, Trigonia Etallom, Avicula obliqua, 
Pecten suprajurensis?, and Rhynchonella pinguis. Towards the 
top of the quarry the limestones become chalky. ‘These great masses 
of course occupy the country for several miles; and thus we find this 
same fossiliferous zone three miles to the north at Bayel. Here, too, 
the Rhynchonella-bed lies at the base, and the upper part is chalky. 
Cladophyllia occurs here and there; and in addition to the same 
Trigonia and Avicula, there are Pecten vimineus?, Cyprina callosa, 
Roém.?, Lerebratula tetragona, and Cidaris Smuthu. Traced still 
further upwards, the rocks become lithographic and unfossiliferous. 
The whole of this mass was considered Astartian by Leymerie; and 
hence he gives 320 feet as its thickness; but it plainly represents 
every thing which lies between the Oxfordian and Pterocerian. 
The next recognizable beds are irregular somewhat marly lime- 
stones, with Pteroceras Ponti and Mytilus acinaces. In such a mass 
it is difficult to recognize Astartian beds ; but it seems probable that 
some part at least of the lithographic limestones belong to them ; 
and hence the overlying beds, which are irregular in character, and 
contain one of the characteristic fossils, are the best representatives 
of any thing that could be called Pterocerian that has been hitherto 
seen. On the north of Bar-sur-Aube the marls with Hvoqyra 
virgula are well exposed, but do not yield many fossils, the chief 
noticed being Pleuromya tellina. This portion of the series pos- 
sesses but little interest, but is followed by the * Portland” litho- 
graphic limestones similar in character to those at Joinville, and 
containing a plicatiloid Ammonite like A. beplex. Above this 
is seen, near Bossancourt, another important Oolite in this series. 
Tt is the one which, in the Haute-Marne, is called the Bure 
Oolite® here developed to a bed of considerable importance, more 
than 6 feet in thickness, and worked in underground mines for 
building-stone. It is singularly free from fossils ; but below it are 
some dirty-coloured oolites, with Amm. suprajurensis and small 
oysters; whilst above are thin-bedded rather chalky limestones, 
with oysters also, and the following fossils—Cardium Dufre- 
noycum, Cardium Verioti, and Trigonia truncata. These pass into 
varying limestones, amongst which are the beds containing great 
branching earth-filled hollows, hence known as the carious lime- 
stones. They are well developed—partly chalky, partly rubbly— 
all the way to Trannes, where a large quarry shows the last of the 
jurassic rocks in this district. In these beds, which would probably, 
if searched, reveal a good fauna, occurred Lima suprajurensis, L. bo- 
loniensis, Pecten suprajurensis, Lucina fragosa, Terebratula subsella 
(var. minor), and teeth of Lepidotus. We here therefore do not see 
beds so far up in the series as those of the Marne valley, no repre- 
sentatives of the spotted limestones or of the vacuolar colite being 
preserved ; but the lower portions seem to become more and more 
fossiliferous as we pass westward. 


528 J. F. BLAKE ON THE UPPER 


Reel) 

CON ec (acs) 

i RA ia THAME 
ANS al Mt 


uh 
eed) 


SS 


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ealtes LeVannage 


Ruf \Ss 


(Oxf Gril € , ae 
Lani nee a ren Pee a a 


The next traverse, though not along the valley of a river, is an 
exceedingly instructive one, as we get an almost complete section 
from the Corallian to the uppermost beds. ‘This is across the hill 
that intervenes between Gyé-sur-Seine and the Riceys in the south- 
western portion of the department. Commencing at the former 
place, we find a cliff cut at the road-side, and all the rocks laid bare. 
At the base is (1) shelly limestone, similar to that at the top of the 
Clairvaux section, with indications of bastard oolite, like the Di- 
ceras-beds of Doulaincourt, and containing crinoid stems and Rhyn- 
chonella pingus. There is therefore still an absence here of the great 
Diceras-masses ; and thetypeis conformable to that of the valley of 
the Aube. Next (2) are from 30 to 40 feet of chalky limestones, split- 
ting into rather thin layers, with Pholadomya congulata, P. cor ?, and 
Amsccardia Legayt. Above this comes (3) a 2-foot band of a curious 
blue earthy limestone, with globular masses of calc tuff. This seems 
to be a bed of separation between two groups of rocks; it contains 
few fossils, except Pholadomye (cf. P. ampla). If any definite 


JURASSIC OF THE PARIS BASIN. 529 


line therefore can be drawn, it is above this that the Astartian zone 
should commence. Next comes (4) a mass of about 30 feet of solid 
light-coloured sublithographic stone, with two bands of remarkable 
character, made up of rolled fragments recemented, with Natice, 
Pecten striatus, Ceromya wabrensis, and abundance of Terebratula 
Leymerti. It is just such a band as this that Prof. Hébert (31) 
elsewhere thought a good line of separation, showing the distinctness 
of Astartian and Corallian beds; but, as it is obvious that they 
cannot both be such, they both lose their importance, and prove that 
there were frequent changes in the deposits about this period. This 
same feature is indicated by subsequent deposits in this section, in 
which are many rubble-beds alternating with lithographic lime- 
stones and occasional oolites. Such beds have athickness of about 
60 feet, and contain Pholadomya Protei, P. rostralis, Thracia tenera, 
Pecten Tombecki, Terebratula Leymeru, Rhynchonella ? matro- 
nensis, Cidaris Smithi, &ce. This is followed by the Virgulian 
marls, full of Hwogyra virgula and Pholadomya rostralis, for 
16 feet, exposed in the roadside, but extending far up the hill, and 
finally surmounted by lithcgraphic limestone, with Terebratula 
subsella. On continuing the route to the Riceys the descent of the 
hill shows a second section corresponding closely with the first, until 
the chalky limestones (2) are reached; but beneath these, instead 
of the shelly limestones with only traces of Diceras, is a large deve- 
lopment of true rubbly Diceras oolite, of which 15 feet in sheer 
height are seen in one spot, although there may be much more of it. 
These beds form the cappings of the hills, and assist in producing 
the magnificent escarpments which overlook both sides of the river. 
Wherever these Diceras-béds occur they never require to be looked 
for; and where they do not form a marked feature it is pretty certain 
they are absent. The principal fossils noted were Nerinea wmbili- 
cata?, Diceras arvetina, Nerita Royerr?, Trigonia Ktallon, and Rhyn- 
chonella matronensis. . Beneath these, on the west side of Ricey Bas, 
is cc a mass of 6 feet of a compact oolite, with few fossils in a 
crystalline state. Underlying this, again, is a considerable quantity 
of white chalky limestone, tending to be flaggy on weathering, and 
occasionally false-bedded. Of this there may be 60 or 70 feet, the 
principal fossils being Pholadomya striatula, Avicula Gesneri, and 
Rhynchonella pingms. In downward succession we reach, at Le 
Vannage, a quantity of shelly limestone, with abundance of corals, 
in which Pecten vimineus, Rhynchonella corallina, &e. abound, but 
in which no Cidaris florigemma could be found. On crossing the 
boundary into the department of the Cote d’Or, at a very slightly 
lower level, there are numerous quarries in a calcareous grit, whose 
horizon is accurately fixed by the occurrence of Trigonia spincfera. 
In the last-named department there is a very considerable develop- 
ment of the lower coral-beds, with Pecten vimineus and Rhynchonella 
corallina, followed in regular succession by the Oxford grit and marls. 

This traverse has been given in greater detail because it appears 
to be a very important one, not hitherto noticed or described, and 
serves for a term of comparison between the series of the Aube 


Q.J.G.8. No. 148. QN 


530 J. F. BLAKE ON THE UPPER 


and Haute-Marne and that of the Yonne. The thicknesses given 
by Leymerie must be maxima; but the succession he gives is well 
made out. It is obvious that he reckons the whole down to the 
Diceras-beds to belong to his Astartian: the first-named is his “* no- 
dular white limestone,” though none of his lists of fossils contain 
Diceras; and the underlying beds observed in the valley of the. 
Laignes are his “lower coral limestone ” in its three subdivisions of 
“ compact hmestone,” “ levique hmestone,” and * oolitic shell lime- 
stone.” Now it has been seen that the Diceras-beds occupy the 
position of the shelly limestone of Clairvaux and Bayel, and hence 
we ought to find the true Coral Rag below in the Aube valley. 
Unfortunately nowhere has Cidares florigemma been noted; but the 
false-bedded character of the ‘‘levique” limestones, and the record 
by Leymerie of a bed in them, at Gloire Dieu, entirely composed of 
Crinoids (a feature so characteristic of the Coral Rag of the Meuse), 
leads to the conviction that it is in these limestones we must look 
for the representative of the Rag, and that the coral shell-beds of 
the Cote d’Or must be regarded as “‘ Coralline Oolite,” so far as any 
separation of these can be effected. On the other hand the great 
development of beds below the Diceras-zone and the coralliferous 
character of the base leads on to what we shall seein the department 
of the Yonne, and show, to a certain extent, a repetition of the section 
in the valley of the Marne. 

5, Department of the Yonne.—The general description of the Upper 
Jurassic rocks in this department is by Leymerie and Raulin, who 
divide them as follows (25) :— 


Upper JURASSIC. 
1. Portland limestones, 135 feet. 
2. Kimmeridgian marls and limestones, 330 feet. 


Mipp.e JurASsIc. 
. Astartian limestone, 83 feet. 
. White coral limestone, 300 feet. 
. Upper Oxfordian limestone. 
. Middle Oxfordian limestones or marls, 260 feet. 
. Lower Oxfordian ironstone, 33 feet. 


Sue CO bD 


Equally important, however, in this case are the subdivisions of 
M. Cotteau (34, 42), though they are not very definitely formulated. 
They are as follows :— 

PoRTLANDIAN. 
1. Zone of Pinna suprajurensis. 


2. Zone of Ammonites gigas. 


KCIMMERIDGIAN. 


SEQUANIAN. 
1. White limestones. 
2. Lithographic limestones. 


CoRALLIAN. 
Diceras-beds. 
OXFoRDIAN. 
1. Zone of Ammonites plicatilis and Cidaris fiorigemma. 


JURASSIC OF THE PARIS BASIN. 531 


The lower portion of the Oxfordian is not brought into this scheme, 
the last subdivision being in another scheme made the base of the 
Corallian. 

Two river-valleys cross this district, and the traverses made along 
these give a complete idea of the development of the rocks under 
consideration, which here attain their maximum in the lower part, 
where they are truly marvellous. 


Fig. 9.—Map of the Valley of the Armangon, south of Tonnerre. 
(Scale 1 : 80,000.) 


We commence with the valley ofthe Armancon. At Ancy-le-Franc 
an admirable section commencing in true Oxfordian rocks is seen 


above the church. Towards the base are marly beds, with only an 
2n 2 


Hae J. F. BLAKE ON THE UPPER 


occasional calcareous band. ‘The fossils at once indicate the horizon, 
those noted being Amm. tricristatus, A. Martelli, Pholadomya an- 
gustata, and Arca rhomboidalis?. Traced upwards these marls 
become more calcareous and are almost limestones. They are there 
crowded with Pholadomya paucicosta, and have also P. flabellata, Cer- 
comya antica, Pinna lanceolata, a Trigoma like T. perlata, and Pecten 
demissus. ‘They become finally less fossiliferous, and are capped by 
shelly limestones. This series may be traced with the same 
character westwards to a great roadside quarry, where a face 
of 40 feet of sandy limestone is worked, and so on to Pacy, 
where the weli-xnown quarries have yielded many fossils, the 
chief of which are Amm. ef. plicatils, Belemnites ? Royert, Pholado- 
mya paucicosta, Myoconcha Rathervana, Trigoma ef. wrreqularis, 
Pecten demissus, Exogyra spiralis, and, according to Cotteau, Ostrea 
dilatata. There can be no doubt that these rocks represent true 
Oxfordian strata; but their relation to those in the valley of Jully 
has not been made out, though it is highly improbable that the 
latter are older, as they contain, according to Leymerie and Raulin, 
the characteristic fossils Ammonites perarmatus, Rhynchonella va- 
rians, and Pecten fibrosus, which have not been found here. The 
numerous changes thus observed in the character of the rocks which 
underlie the Corallian must be due either to an unconformity or 
to a very variable development of the Oxfordian strata; but we can 
hardly accept the arrangement of the strata by the above-named 
authors, who place the beds with Rhynchonella varians as the Lower 
Oxfordian, while those at Ancy and Pacy are considered to be one 
facies of the Middle Oxfordian, of which the other facies is the Coral 
Rag, to be noted presently at Merry-sur-Yonne andelsewhere. ‘The 
next succeeding series in this district is a mass of lithographic lime- 
stones, tending to split into thin plates, and for the most part un- 
fossiliferous. Here and there, however, in its mass are fossiliferous 
zones, apparently the representatives of strongly characterized beds 
further to the east. The lowest of these has not actually been met 
with ; but it is very instructively described by Cotteau (43), as seen 
in the neighbourhood of Gland, where the following upward suc- 
cession is determined :—(q) ferruginous Oxford clay; (6) sponge-bed ; 
(c) limestones of Pacy; (d) rubbly limestone; (¢) coral-bed, with 
Pecten subarticulatus and Cidaris florigemma ; (f) lithographic lime- 
stones ; (g) coralline limestone of Tonnerre. It is impossible not to 
see in the beds ¢ and d the continuation of those of Vannage and 
the Cote d’Or, the succession being perfectly consonant to what 
we might expect. There are, however, other bands in the litho- 
graphic limestones nearer the summit, one of which is seen on the 
roadside west of Commissey, associated with a rubble-bed ; another 
oceurs close to the village of Angy (below the white limestone), 
where it was seen by Hébert (31), and considered to be a junction- 
bed between the Corallian and Oxfordian, though lying upon one 
great mass of lithographic limestone and covered by another. The 
commonest fossil of this series is Rhynchonella coralliima. ‘This por- 
tion, though, being more lithographic, it is less clearly characterized, 


JURASSIC OF THE PARIS BASIN. 533 


will occupy the place of the compact and “levique” limestone of 
Riceys, if we find above it the representative of the Diceras-beds. 

Above the lithographic limestones comes an enormous mass of 
white chalky limestone, seen on the right bank of the Armangon be- 
tween Commissey and Tonnerre, and on the left bank at the well- 
known quarries of Angy and Tonnerre. The limestone of Angy is 
celebrated for its fossils, which occur in profusion and in admirable 
preservation. A face of 50 feet is worked, the rock being more sandy 
towards the base, and having masses of Septastrea, large Monthvaltic, 
Trichites Saussurer, Diceras sp., Rhynchonella matronensis, Terebratula 
maléonensis, and Cidaris, apparently not C. florigemma. As this is the 
<< White coral limestone” of Leymerie and Raulin, we find it credited 
with 300 feet of thickness. This it might well be judged to have from 
the hills in which it occurs; but it 1s not again well seen till the 
great quarries near Tonnerre are reached; and here it has a 60-feet 
face, which contains a different and higher portion, remarkable for 
the great masses of banded flints which lie more or less in lines. 
These are Nos. 7 to 15 of Cotteau’s section (34) and contain many 
unusual Echinoderms. We here also find the overlying rocks, 
which are, first, a solid block of 10 feet with rolled nodules and a 
few specimens of Diceras and Lhynchonella pinguis, and next some 
nodular and coralliferous beds. The corals are rolled and not wm situ 
(No. 56 of Cotteau). Next comes a coarse oolite, and then another 
rubbly bed with massive corals, Rhynchonella pinguis, Terebratula 
Leymeriz, and many other fossils, but no sign of Cidaris florigemma 
(No. 24 of Cotteau). On the opposite side of the river no good 
exposures of the white limestones occur at present; but Cotteau 
describes an important quarry, called ‘‘ Voceuses,” which contains, 
amongst other fossils, Cardium corallinum, Glypticus hieroglyphicus, 
and Cidaris florigemma. The upper part, however, is seen about 2 
kilometres south of Tonnerre, containing but few fossils, but in 
parts composed of white rubble of exactly the usual character. It is 
plain, therefore, that all this mass must be considered a develop- 
ment of the Diceras-beds of Riceys, in which the peculiar character 
is less regularly marked, and the fossils are therefore somewhat 
different. On this side of the river other quarries show rubbly beds 
above, then sublithographic stone, and then another rubble-bed 
full of Terebratula Leymern, which Hébert (31) regards as the base 
of the Astartian, though we have seen near Gyé that there may be 
several such overlying each other. Above comes a coarse brown 
oclite, which may, or may not, be the same as that seen on the west 
of the river. In any case the sequence on this side matches very 
well with the Gyé section, in which also rubbly beds with 7. Ley- 
merit were overlain by oolite. The upward succession is of no 
particular interest here; but the lithographic stones which overlie 
the oolites gradually become more chalky, and finally yield Evoqyra 
virgula and Pholadomya acuticosta, beyond which the beds have 
not been traced, as they are better known near Auxerre. 

The next traverse is up the valley of the Yonne, where it is best 
to commence with the uppermost beds. It has been seen that 


534 J. F. BLAKE ON THE UPPER 


Fig. 10.—Map of the Yonne Valley south of Aumerre. 
(Scale 1; 240,000.) 


= Harty Lith L) 


Vallen 


Y 


iy 4 

ty ( 

j R Yonne p | 
f 


aulty 2 Vilts } 
=~ 72 Chatvaw, oe 


WZ 
oo y 
P 
| a BA J © 2 
: 


=S5550 


Cotteau divides the “ Portlandian” rocks into two groups, the 
upper of which is the zone of Pinna suprajurensis. This is very 
well seen in the neighbourhood of Auxerre, and, like all these upper 
rocks, is more fossiliferous as we go west. After the splendid 
monograph by Loriol and Cotteau, it is superfluous to quote the 
fossils, the great majority of which come from the upper portion. 
We need only compare the series with those seen before. In the 


JURASSIC OF THE PARIS BASIN. 53D 


valley of the Yonne there is no oolite to compare with the Bure 
Oolite ; but the fossiliferous beds here correspond in character and 
fossils to the limestones of Trannes, and therefore to the “ various 
limestones.” It is also possible that, from the unevenness of the 
Neocomian denudation, portions of the higher zone, or “ spotted lime- 
stones,” may be represented, as at Villefargeau; but the series is 
certainly incomplete in its upper part; and the question of any passage 
here into the Neocomian, which De Loriol propounds (42), cannot 
possibly be seriously raised. The lower portion of the ‘“‘ Portlandian ” 
rocks is well seen on either side of the Yonne valley, where quarries 
eccur with a 40-feet face of a soft chalky limestone with intervening 
beds of marl, in which there is occasionally a lumachelle of Hvogyra 
vurgula. Some of the marls contain abundance of Thracia depressa ; 
and the other fossils noted were Amm. gigas, A. suprajwrensis, A. 
Gravesianus, Plectomya rugosa, Cyprina Brongnartc (in another 
quarry), and Pinna granulata. These beds correspond in position to 
the lithographic limestones of Joinville. The beds in this locality 
have a gentle rise to the south, and the sequence can be well taken 
up again near Vincelles and Bailly. The tops of the hills,here are 
occupied by the “ Portland” limestone, beneath which come earthy 
fossiliferous limestones with large examples of Terebratula subsella 
and Pholadomya multicosta, which graduate downwards into marls 
full of Hxoqyra virgula. The Astartian beds are not well seen in 
any section; but they consist partly of solid and partly of rubbly lime- 
stones, with bands of Terebratula Leymerii, containing Plectomya 
rugosa. Towards the base the beds become oolitic, then nodular, 
then bedded; then comes a 16-feet mass of brownish oolite* with Rhyn- 
chonella pinguis, then 2 feet of solid limestone. Below this comes 
an 8-feet block of solid character, here and there containing the 
peculiar pisolite of the Diceras-beds, which forms the cap to a mass 
of white limestones, varying from oolitic to chalky, and containing 
Ceromya excentrica, Homomya compressa?, Nerineea pseudospeciosa ?, 
Corbis gigantea, Trigonia Htallon, T. variegata, Pecten Tombeckt, and, 
perhaps, Cidaris florigemma. The series thus described is ob- 
viously identical with that seen at Tonnerre, with the exception 
that there are no beds of rolled corals over the Diceras-beds. The 
white limestones extend some way to the south, where they are 
succeeded by lithographic limestones. The basement has not been 
seen, but is stated by Cotteau (42) to contain Rhynchonella coral- 
lina, Terebratula humeralis, and Pholadomya paucicosta. At Mailly- 
la-ville are seen some 20 or 30 feet of very flaggy beds, succeeded 
by 12 feet of oolite, and then a development of the most extraor- 
dinary kind, totally different from any thing seen before and of great - 
interest. Along the banks of the Yonne are fine hoary cliffs, 
composed at first ef a rock resembling a consolidated cale tuff, 
without any stratification and weathering into peaks; the fossils 
are not so numerous as might be expected, though Pecten artrculatus 


* Above this bed, Hébert (21) draws the line between the Coral Rag and the 
Kimmeridge Clay. 


536 J. F. BLAKE ON THE UPPER 


and Rhynchonella corallina were noted. On the same level these 
change into Diceras-beds of characteristic material, with many ex- 
amples of that genus ; and finally, at Merry-sur-Yonne, an immense 
cliff, 200 feet in height, and carved into fantastic shapes, frowns over 
the river, and is geologically an unstratified heterogeneous mass of 
Coral Rag, Diceras-beds, and rubble all together. Huge masses of 
Thamnastrea, delicate branches of Caryophyllia, and fan-like growths 
of Thecosmilia here and there ornament the surface ; and in the inter- 
spaces innumerable specimens of Diceras, Cardium corallinum, stout 
spines of Cidaris florigemma, and Trichites a foot in length are crowded 
together. It is the finest example of a Coral Rag visible either in 
the basin of Paris or anywhere in England, and it is, undoubtedly, 
impossible to confound it with the much higher beds described as 
occurring at Bailly. Rocks of this description form the boundary 
of the picturesque valley through which the railway runs past 
Chatel Censoir to Coulanges. At the latter place there is an almost 
equally fine cliff, composed of a white limestone as beautiful as that 
of Angy, though apparently on a lower level, being really only 
a modification of the Coral Rag of Merry. Here, too, the fossils are 
in profusion and perfect preservation ; the finest examples of various 
species of Diceras occur, associated with Werinea Cottaldina?, N. 
subnodosa, Cardium corallinum, Hinnites, Terebratula wsignis, 
Cidaris florigemma, corals growing in reefs, and sponges. ‘This 
great corallian development proves conclusively that Dvceras, 
Nerinea, and Cidaris florigemma are not always characteristic in 
any sense of distinct horizons, and that the lowest portion of any 
possible rocks which may be referred to the Coralian may be 
characterized by the latter species. If the series has been rightly 
traced in our journey westwards, these great coralliferous masses 
must be the grander development of the coral-growths of the base 
of the series in the Aube, only represented by thin bands at the 
base of the lithographic limestones in the valley of the Armancon. 
Much discussion has arisen on the correct position of these lower 
coral-beds. At first they were confounded with the upper Diceras- 
beds and supposed to be repeated by some fault (3, 6). Certain 
marly limestones developed at Vermanton, with a proportion of 
Oxfordian fossils, were, at that time, considered to belong to Ox- 
fordian strata (9,14). These Vermanton lhmestones were after- 
wards proved by Raulin to overlie the Coral Rag of Chatel Censoir 
and Merry (15); and as they were still considered by him as Upper 
Oxfordian, it followed that the Coral Rag below should be called 
Middle Oxfordian, and as such it appears in the monograph by 
Leymerie and Raulin quoted above (25). Cotteau, however, showed 
by an exhaustive examination of the fossils (16), first, that the Coral 
Rag was essentially “Corallian” in the usual sense, and, secondly, that 
the Vermanton marly limestones, though from their similar lithology 
they appear more closely allied to the Oxfordian than to the coral- 
beds below, yet, on the whole, are really Corallian. It is noteworthy 
that though these marly limestones are quoted in the earlier deserip- 
tions as marls ‘a Anum. marantianus” (15), the only Ammonite given 


JURASSIC OF THE PARIS BASIN. 537 


in Cotteau’s exhaustive list is Amm. achilles; of the other fossils, 
Trigonia clavellata, Ceromya excentrica, Unicardium globosum, 
Phasianella striata, and Cidaris florigemma are the most noteworthy. 
These beds are believed by MM. Royer and Tombeck to correspond 
to the marly beds overlying the Diceras-beds of the Haute-Marne ; 
but by their position they must be part of the lithographic 
limestones which underlie those beds, though overlying the basal 
Coral Rag. 

The true position of these rocks is further certified by continuing 
the examination downwards. On the north side of the Yonne, at 
Coulanges, are seen, below the beds already described, 12 feet or 
more of uniform compact limestones with Gryphea dilatata, and 
then some sandy limestones, like those of Pacy, but more false- 
bedded and irregular, with abundance of Yerebratula bisuffar- 
cnata, also Trigonia spinifera, Nerinea allica, and many corals 
here and there. These, which are doubtless the same as the 
siliceous limestone of Druies, correspond very well with what we 
might expect, though under a form slightly different. It is at some 
distance below these that the fossiliferous zone of Amm. transver- 
sarius occurs on the road to Clamecy. 

6. Department of Niévre—The only description of the Upper 
Jurassic rocks of this department is by Ebray (32), supplemented 
by some notes inthe paper by MM. Douvillé and Jourdy (60). The 
series is thus described by Ebray :— 


KIMMERIDGIAN. 


Astartian Limestone. 


CoRALLIAN. 


Upper Lithographic Limestone. 

Oolite with small Diceras. 

Chalky Limestone. 

Lithographic Limestone. 

Oolite of La Charité with Diceras arietinum. 

Marly Limestone.and Lower Lithographic Limestone. 


ARGOVIAN. 


Sponge-marls with Amm. canaliculatus. 


The only traverse made in this district 1s along the valley of the 
Loire, from La Charité to the north of Pouilly. At the former 
place large masses of very marly limestone, having a markedly Ox- 
fordian aspect, are worked, in which fossils are very rare, but from 
which Amm.bimammatus and Amm. canaliculatus have been recorded. 
These continue some distance; but halfway to Meves are large 
quarries showing a 40-feet face of white limestone, with much of the 
rolled pisolitic material characteristic of Diceras-beds; and that genus 
here abounds, associated with Nerinwa. Had one never seen the 
limestone of Coulanges, one would, doubtless, take this for the 
ordinary Diceras-bed, and its low position would be a difficulty ; but 


538 J. F. BLAKE ON THE UPPER 


Fie. 11.—Map of the Loire 
between Sancerre and La 
Charité. (Scale 1: 80,000.) 


i iy) \sy CA ha 
Fiat SPACE. i | 


ery 
OrES 
oat 


UOve 
‘Raxs 


i 


TEAL SE LEED Se 


by iuy 


A ois 


PEE py 
SSS eae 


Vs 


SS 


= $< iower Dreerss 
= -Seds) 


Pe 
Re 
i 
oe 


there is no difficulty now in re- 
cognizing it as the tailing off of the 
great Coral-Rag-like mass which 
characterizes the valley of the 
Yonne, more especially as it has 
not been recognized on the left 
bank of the Loire. The whole 
space between these quarries and 
those of Pouilly appears to be oc- 
cupied by barren lithographic lime- 
stones, the thickness of which it is 
impossible to estimate. The quar- 
ries of Pouilly are worked in a 
finely oolitic stone, somewhat 
false-bedded and containing few 
fossils ; this must correspond to the 
white limestone of Tonnerre and 
Vineelles ; and we should expect to 
find representatives of the Diceras- 
beds above: of these no evidence 
was found at Pouilly; but Rau- 
lin (3) and Douvillé and Jourdy 
(60), speaking of the same beds 
seen on the opposite side of the 
river at Sancerre, describe a re- 
gular Diceras-bed as attaining its 
maximum there, though dying 
away to the west. On the hill 
north of Pouilly are seen, first, 
lithographic limestones, then some 
rubbly limestones with TYerebra- 
tula Leymerw, and at the summit 
the marls with Hwogyra virgula, 
and no more is apparent in this 
direction, though higher beds 
might well be developed. These 
present no special point of in- 
terest, beyond showing the con- 
stancy of this portion of the series 
in contradistinction to the varia- 
bility of the beds below. 

7. Department of the Cher.— 
Works on this department have 


- been published by Fabre in 1838, 


and Boulanger and Bertera in 
1850 (10); but the most recent 
description is that by Douvyillé 
and Jourdy, above quoted (60). 
These latter authors classify the 
series as follows :— 


JURASSIC OF THE PARIS BASIN. 539 


KIMMERIDGIAN. 

Limestone of Barrois. 
Virgulian marls. 
Astartian limestone. 

A. Marls and nodular limestones. 

B. Nerinzan Oolite. 

©. Kucoidal marls and limestones. 

CoRALLIAN. 


Upper lithographic limestones with Amm. achilles. 
D. Limestone with Pinna. 
HK. Compact limestones. 
Chalky limestone of Bourges. 
Lithographic lmestones. 
Sponge limestones, with Amm. marantianus, A. bimammatus, and A. plicatilis. 


The first traverse in this department 1s in theneighbourhood of Bour- 
ges, where the succession has been already well studied and described. 
From 3 to 6 kilometres north of the city the Virgulian marls appear, 
overlain to the north by lithographic limestones, belonging doubt- 
less to the so-called Vortland, and at the hill immediately outside of 
Bourges the different Astartian beds are seen. The highest are 
earthy limestones, partially lithographic and often extremely marly. 
The characteristic fossils are large examples of Ceromya excentrica and 
Pholadomya rostralis, Terebratula subsella and Plectomya rugosa; 
WNatica Royeri also occurs. The next seen are a few feet of rubbly beds, 
becoming very oolitic below, and more like the corresponding rock in 
the Boulogne area than any yet seen. It is the Nerinzan Oolite of 
the authors quoted, and contains Nerinea Desvoidyi?, Trichites 
Saussurei, Terebratula subsella, abundance of 7’. Leymert, Rhyn- 
chonella pinguis?, Pholadomya rostralis, and Trigoma Baylei, the 
last-named forming an interesting connexion with the rocks at Havre. 
The lower fucoidal marls have not been examined ; but from the list 
of fossils quoted, among which may be noted Mytilus perplicatus, 
Plectomya rugosa, Terebratula subsella, and Rhynchonella pinguis, it 
is obvious they form part of this series. On the south of Bourges a 
different group of rocks occur, the upper part being lithographic, 
unnecessarily divided into two parts. The fossils are few, and 
belong to Pinna obliquata (a broad species), Lucina cf. imbricata, 
and Terebratula ? tetragona. These are of considerable thickness, and 
are underlain by beautiful white limestone, which betrays its origin 
when closely examined. It is then seen to be an exceedingly fine 
rubble, haying here and there larger grains, and becoming almost 
like the Diceras-beds, to which it corresponds, and into which it 
is said to change at Sancerre. Terebratula cncta and Rhynchonella 
coralina are extremely abundant. Lima leviuscula, L. rigida, 
and Corbis gigantea were also observed. In the old quarries of the 
Chateau, what are probably lower beds are seen through 20 feet and 
more. Here are abundance of massive corals, with Crdaris flori- 
gemma and many shells, but fewer Brachiopoda. ‘This, then, is the 
true Coral Rag as usually seen, and corresponds to the white lime- 
stones of ''onnerre and Augy. Below this the beds have not been 
examined, as they are stated to consist of a uniform mass of litho- 


540 J. F, BLAKE ON THE UPPER 


Fig. 12.—Map of the Country round Bourges. (Scale 1: 160,000.) 


graphic limestone over a breadth of 20 kilometres, and to show 
no signs of the lower limestone of Coulanges and La Charite. 
Boulanger and Bertera, indeed, mention a coral limestone below the 
lithographic, to be seen near Venesme, which may represent it; but 
they correlate 1t with beds with Amm. hecticus, in which case it must 
be Oxfordian. 

In the last traverse of this range the swallowing up ofall portions 
of this series into uniform lithographic limestones, below, at least, the 
Astartian beds, is carried to its maximum ; for though, near Chapelle, 
the white limestone of the old Chateau quarries is still seen to be 
developed, it is much diminished in thickness, and the whole of the 
weary way from that locality to Chateauneuf-sur-Cher, along the 
borders of the river, is occupied by a dreary monotony of thick- 


JURASSIC OF THE PARIS BASIN. 541 


bedded, unfossiliferous, lithographic limestones. One peculiar fea- 
ture, however, is to be noted. About two miles south of St. Florent, 
and again at Lapan, at a lower level in the series, the soil appears 
of a peculiar rich red colour, and at the latter locality the cause has 
been ascertained. Instead of the ordinary limestones there are some 
beds, at least 3 feet or 4 feet thick, of a crystalline irregular mass, 
which are suggestive of the strangely altered rock produced from 
corals and coral-brash; and in this mass are cavities filled with 
large, round, concentrically coated, limonite concretions. It is by 
the decay of these that the fields are covered with the red soil; and 
they may be regarded as the last vestiges of coral-growths, the lower 
of which might well correspond to the Coral Rag of Merry. As we 
approach Chateauneuf the limestone becomes white and more marly, 
but still almost, if not quite, unfossiliferous ; and it is only to the 
south of the town that the canaliculate Ammonites set in, associated 
with abundant sponges and Amm. bimammatus and Amm. Martelli, 
on which little need be said, since they are so obviously Oxfordian. 
It may be noted, however, since the authors quoted place these beds 
as the base of the Corallian, that the first Ammonite to appear 
going down is Amm. canaliculatus; and this is far above the 
sponge-bed. 

The valley of the Indre has not been examined ; but Douvillé and 
Jourdy assure us that here even the white limestones are wanting, 
and the whole mass between the Astartian and Oxfordian consists 
of nothing but barren lithographic limestones, spreading over an 
expanse of 30 kilometres from Levroux to Chateauroux. 


Il. Tae rwo CHARENTES. 


This district scarcely forms part of the Paris basin, being rather 
the northerly extension of the Pyrenean, and will doubtless serve 
as a term of comparison between the two. Nevertheless, being so 
close to the termination of the great range, it 1s more conveniently 
studied on the present occasion. The main question, however, 
of which the solution has been sought, is the age of the so-called 
** Portland” and even ‘‘ Purbeck” beds—whether there are, in fact, 
in this area any rocks corresponding to those that are known by 
those names in England. 

1. Department of the Charente.—The Jurassic rocks of the depart- 
ment have been described by Coquand (24), who divides the upper 
portion as follows :— 


Upper J URASSIC. 
Purbeckian. 
Portlandian. 
Carious limestone. 
Limestones with Nucula inflexa. 
Oolitic limestone. 
Limestone with Cardium dissimile. 
Sands. 
Kimmeridgian. 
Virgulian marls. 
Pterocerian limestones. 
Astartian limestones. 


542 J. F. BLAKE ON THE UPPER 


MiIppLeE JURASSIC. 


Corallian. 
Oolite with Neringe. 
Coral limestone. 
Solid limestone. 

Oxfordian. 

Callovian. 


The whole series, as seen between Rochefoucault and Angouléme, 
is extremely calcareous, and no physical line can be drawn between 
the Oxfordian and Corallian; every portion is more compact than 
usual, and presents no such marked features as are often seen. The 
Oxfordian near Rochefoucault certainly contains coralliferous beds, 
and these give place to a sandy oolite. From Coquand’s descrip- 
tion it appears that the uppermost portion of the Oxfordian contains 
in some places such fossils as Pecten demissus, P. subfibrosus, and Rhyn- 
chonella Thurmanni, and in others Amm. oculatus, A. crenatus, and 
A. Henrici, and is followed by a brecciiform limestone, indicating that 
the Corallian here also lies on different portions and is really uncon- 
formable. The massive limestones of the last-named group occupy 
a wide expanse of country, but finally give way to rather rubbly 
limestones, doubtless the ‘‘Coral Limestone” of Coquand, which, 
however, scarcely forms a “ Rag”; and these, again, are replaced 
by compact suboolitic limestones, containing Diceras at Touvres, and 
in which Coquand records several Nerinee and Cardium corallinun, 
Terebratula insignis, and Rhynchonella corallina. It is obvious, there- 
fore, that the succession, though somewhat obscure from the similarity 
of the rocks to each other, is the same that we have seen in the great 
range before the lower Coral Rag appeared. The Astartian beds are 
not here of any great importance, consisting of marly limestones, with 
Pholadomya rostralis and Ceromya excentrica. The fossils quoted 
by Coquand, except the Astarte supracorallina (“minima”), are not 
specially characteristic; and it is to be noted that he considers 
some beds at La Rochelle, which d’Orbigny placed in the Corallian, 
to be the calcareous representatives of these more marly rocks. 
Traced upwards, however, they become still more marly, and 
contain abundance of Evogyra virgula. Indeed this part of the 
series is rather exceptional; and after seeing so much limestone 
one’s eye is refreshed by the sight of veritable clay, extending 
through a considerable thickness. These irregular beds may be 
taken up again in the railway-cuttings north of Angouléme; but 
the lower portion, distinguished as Pterocerian, which is to be seen 
some six miles to the north, at Vars, has not been examined. It 
appears from Coquand’s description to be more than usually distinct 
in this district by its fossils, which include Nautilus giganteus, 
Pteroceras oceant, Chemnitzia Danae, Pholadomya Prote., Mytilus 
medus, and Ostrea solitaria. The Virgulian marls near Angouléme 
are fairly fossiliferous, the species noted as associated with Hxogyra 
virgula being Natica ornata?, Plectomya rugosa, Arca ef. rhombor- 
dalis, Trigonia concentrica, T. monilifera?, Gervilliia kimmeridiensis, 


JURASSIC OF THE PARIS BASIN. 543 


Thracia depressa (towards the top), Mytilus virgulinus, Ceromya 
concentrica, and Pecten suprajurensis; and to these Coquand adds 
Ammonites longispinus, Pholadomya acuticosta, and Pleuromya 
tellina. The rocks therefore which here underlie the “ Portland” 
are the same that have been hitherto recognized as Virgulian, and 
represent the lower, or at most the middle, portion of our English 
Kimmeridge Clay. We are therefore prepared to find that here also 
the “ Portland” beds are no higher than those usually so called. Now, 
tracing upwards the beds towards Angouléme, we find aset of rather 
rubbly fossiliferous limestones, containing Pholadomya decemcos- 
tata, Trigonia concentrica, T’. ef. Bronnit, Arca laura, and Cerithium 
septemplicatum. These rocks become more sandy by degrees, and 
ai last form a peculiarly crisp calcareous sandstone. ‘The fossils in 
this sandstone, as noted, are Pseudomelania gigantea, Corbis Rather- 
wana, and Avicula Credneriana, all characteristic species of the 
* Lower Portland.” No more is seen in this section, as the Hip- 
purite limestone lies uncontormably upon the last-named stratum. © 
The higher beds of the district were therefore examined in the 
neighbourhood of Jarnac. The section at Souillac could not be 
verified as showing four subdivisions overlain by gypseous beds; but 
much oolitic stone is worked to the west of Jarnac, as far as Chassors 
and Nercillac, to a thickness of at least 12 feet; in this Cyprina 
Brongniarts is the characteristic fossil. From the fact of this species 
not being quoted, but in its place Cardium dissimile (only a single 
doubtful example of which could be found), and from the similarity 
of the casts of these two shells, is it possible to conclude that for 
the latter we ought to read the former in Coquand’s lists? Still 
something very like the Cardium dissimile was seen at Nercillac, and 
also, as will be seen, in the Lle d’Oleron. The other fossils noted 
at Chassors &¢. are those often accompanying Cyprina Brengniarta, 
or belonging to a higher zone, viz. Cyprina implicata, Astarie 
rugosa, Pecten nudus (= yarnacensis), Cardium Dufrenoycum, 
Astarte regularis, Corbula mosensis, and Mactra insularum. Over- 
lying these near Jarnac are more flagey beds, with many small 
oysters at the base. Comparing this with the more complete 
development in the Haute-Marne, the beds here shown belong 
obviously to the zone of Cyprina Brongniarti; and as several of 
the fossils indicate at least the upper part of it, though the 
gypseous beds above give no clue to their age, and we are 
therefore left without proof in this district of the equivalents 
of the vacuolar oolite representing the true Portland, it 1s pro- 
bable that such beds, though under a somewhat freshwater con- 
dition, are actually reached, being represented by the so-called 
« Purbecks.” 

2. Department of the Lower Charente.—The rocks in this de- 
partment, originally studied by d’Orbigny (14), have since been 
more fully described by Manes (17), who divides them as 
follows :— 


544 J. F. BLAKE ON THE UPPER 


Portlandian, 280 ft. 


Gypseous beds. 

Lumachelle limestones. 

Tabular compact limestones. 
Alternations of compact limestones. 


Kimmeridgian, 260 ft. 
White chalky limestone. 
Marls with Lxogyra virgula. 
Corallian, 330 ft. 


Coral limestones. 
Limestone with Nerineee. 


Oxfordian, 550 ft. 
Shaly marls. 
Clays and marly limestones. 


Whether the rocks in the Lower Charente are really different 
from those in the Upper, or whether their exposure in the sea-cliffs 
. gives a better opportunity for judging differences, certain it is that 
in the neighbourhood of La Rochelle one is immediately aware that 
one is studying the development in a new basin, and things are 
changed. If, in fact, we examine the lists of d’Orbigny as given 
in his ‘ Prodrome,’ we find that out of 222 Corallian species from this 
locality, no fewer than 90, or more than 40 per cent., are peculiar, 
while only 80 occur in any of the localities yet studied. 

The Oxfordian strata are of particular interest as occurring at 
Marans, whence the name of Ammonites marantianus is derived, 
which indicates that we should expect the overlying ‘“‘Limestones with 
Nerinee” to belong to the true Corallian series. The lower portions of 
these occur on the north side of La Rochelle; but the upper portion 
is the more fossiliferous, and occupies the first two headlands to the 
south, viz. the Point des Minimes and the Point de Roux. Here they 
are yery marly, with beds of limestone crowded with cavities formed 


Fig. 13.—Map of the Coast round the Pertuis d’ Antioch. 
(Scale 1: 240,000.) 


SEA 


DR aitipecul 
SSS AyD: 


2 SSN Se a7 adie 
s f de Chatelailbo We S g 


JURASSIC OF THE PARIS BASIN. 545 


by fossils which have decayed, leaving their exteriors perfectly 
moulded. The most abundant fossils are some large curved Montli- 
valtie (M. subnigra, VOrb.?), various peculiar Nerinee, small Cerithia, 
Mytilus pectinatus, Trigonia aculeata, VOrb., T. Tombecki? (LT. Me- 
riant of dOrbigny, but not that species), and <Astarte bicostata. 
Thus the common fossils are the peculiar ones, and one must fall 
back upon more detailed lists of rarer fossils. Amongst those 
quoted by d’Orbigny, the most important for correlation are Belem- 
nites Royert (probably from the lower portion), Ammonites achilles, 
Natica grandis, N. hemispherica, Turbo princeps, Cardiwm corallinum, 
Mytilus acinaces, Lima leviuscula, Avicula polyodon, Pecten subartr- 
culatus, Terebratula insignis, and Hemicidaris crenularis. None of 
the others having a contrary tendency, it is plain that in the great 
mass composing this series we have represented the shell-limestone 
usually included in the Corallian, up to and including the Coral Rag 
itself; though Crdaris florigemma does not appear to flourish here, 
and the corals are not Astrzean. Passing south towards higher beds, 
we find some more marly limestones with Pholadomya Proter and 
— Pinna obliquata not uncommon. If the latter shell may be trusted 
to keep to a single horizon, it should represent the lithographic 
limestones overlying the chalky limestones of Bourges. However, 
at the next point, Point du Ché, a fine development of a Coral Rag 
is seen. This is credited with more than 200 feet by Manes; but 
as the next series named is the Virgulian marls, it is obvious that 
the Astartian beds are included in this. ‘The cliffs have a height 
of 30 feet to 40 feet; and all is apparently Coral Rag, though on a 
higher horizon than usual. Fossils like Arca tetragona, Lima verdu- 
nensis, Terebratula, are abundant, with spines of Cidaris; to which 
may be added from d’Orbigny’s list Nautilus qiganteus, Nerinea 
Mandelslohi, Pinnigena Saussure, Diceras arietinum, and Rhyncho- 
nella pinguis (“ mconstans’’). 

This portion of the series may therefore fairly be placed on the 
horizon of the Diceras- or Supracoralline beds, like the far-off coral- 
liferous deposits at Novion; and it corresponds with the uppermost 
portion of the Corallian of the Charente, as described by Coquand. 
At the cliff near Angoulins, very little to the south of the Point du 
Ché, the last of the rubbly limestones, with Lima leviuscula and 
Ostrea cf. gregaria, is seen occupying the base, overlain by about 
5 feet of marly beds with Pholadomya Prote: and Ceromya ecacen- 
trica of large size, and then a hard limestone band with many fossils, 
amongst which are corals, Pecten striatus, and Cidaris Smithir?, 
but aiso a great abundance of Terebratula Leymerw. The presence 
of this last species would indicate Astartian beds, which are other- 
Wise unrecognized; and these are doubtless the beds to which Co- 
quand alludes when he says d’Orbigny has placed them with the 
Corallian on account of their being calcareous. As noted, however, 
by Hébert (54) only nine of the species found here are common to 
the Corallian beds at the Point des Minimes. In the same paper we 
learn that the marls of Chatelaillon, which show the upper succes- 


Q.J.G.S. No. 148. 20 


546 J. F, BLAKE ON THE UPPER 


sion, contain Amm. cymodoce, Natica hemispherica, Pteroceras ocean, 
P. Ponti, Mytilus jwrensis, Ceromya excentrica, Thracia suprajurensis, 
and Pinnigena Sausswret, and are therefore very fair represen- 
tatives of Pterocerian beds, especially when compared with those at 
Havre. Neither these beds nor the overlying Virgulian marls could 
be specially examined. 

The so-called Portland beds, however, of the Ile d’Oléron demand 
particular attention. As in the Charente department, the upper 
part is gypseous ; but M. Manes does not on that account call them 
Purbeck, but classes all with the Portland. The beds are well dis- 
played along the cliffs and shore of the island on either side of the 
harbour of St. Denis. The lowest beds seen are (1) massive blue 
cale grits, which stand out as reefs at low tide on the north of the 
harbour, and are broken up and stacked on the roads. These contain 
Ammonites Gravesianus and Trigonia ef. concentrica. They are suc- 
ceeded by many other beds, forming scars, which are better seen on 
the south of the harbour, though the succession may not be perfect. 
The lowest beds seen here are (2) marly limestones and grits, of 
unknown thickness, more than 4 feet; next is (3) blue sandy marl, 
3 feet; (4) hard blue sandy beds, with circular fucoid markings, 
with Mytilus Morrisu, Trigonia concentrica?, Cardiwm morinicum, 
and Haogyra nana, 2 feet; (5) earthy marls, 1 foot; (6) solid cal- 
careous block, with Cyprina Brongniart (broad var.), Lucina port- 
landica ?, Pecten suprajurensis, P. jarnacensis, Cardiwm dissimile, and 
Astarte rugosa?, 1 foot; (7) alternations of limestones with luma- 
chelles of Hwogyra nana and earthy beds with Cardiwn Morriseum, 
6 feet; (8) thin-bedded limestones, with shale and clay, 10 feet. 
In the cliffs on the north side, separated from the low-water rocks 
by masses of blown sand, are beds which must succeed the above, 
but perhaps not immediately. These are (9) alternations of thin- 
bedded limestones and black sandy marls, with gypsum, possibly the 
continuation of (8), 12 feet; (10) black and white sandy elays, no 
fossils, 20 feet; (11) thin-bedded and laminated clays, 34 feet; 
(12) black laminated clay and sandy limestones, 8 feet ; (13) purple 
lithographic suboolitic limestones in thin beds, the uppermost when 
weathered becoming a vacuolar oolite, and containing masses of 
shell-fragments, all undistinguishable, unless one is a flattened 
_ Cardium dissimite?, 2 feet. Above this are still some white chalky 
fragments; but no more is seen. Here, again, therefore the same 
argument holds as for the neighbourhood of Jarnac. The fossili- 
ferous beds cannot be lower than the zone of Cyprina Brongmart : 
and the higher beds must in all probability represent true Portland 
rocks. The occurrence of Cardium dissimile is especially note- 
worthy. It is found near the top of the series; and in no other 
locality does it occur lower than the true Portland. If the rocks 
which contain it be of such an age, we must place the overlying 
eypseous beds in the same category, unless we choose to call 
them Purbeck. This we should not be justified in doing un- . 
less we had the whole of the Portland limestone here, which is 
improbable. 


JURASSIC OF THE PARIS BASIN. 547 


II. Normanby witH ORNE AND SARTHE. 


1. Department of the Orne and Sarthe.—It has not been possible to 
examine the feeble development of the Upper Jurassic rocks in the 
narrow strip which leads northwards from Le Mans to Normandy. 
We may note, however, that Hébert (21) shows that the Oxfordian 
beds with Amm. perarmatus and A. cordatus, which he calls “‘ Middle 
Oxfordian,” consist of sand and calcareous grits; while the “‘ Upper 
Oxfordian,” with Zrigonia perlata (?‘ clavellata”), Pholadomya de- 
cemcostata, Perna mytilordes, and Gervillia aviculoides, is marly. This 
is a succession very like that to be seen near Trouville. Near Bel- 
léme are said to be seen, over this, calcareous grits full of Trigonia 
Bronnw (surely not the Supracoralline species!); next some oolitic 
limestones, with large Astarte mysa(?=A. ovalis). Next come piso- 
lites with Nerinee (vrobably equivalent to the limestones of Trou- 
ville and Osmington); and finally, at some distance above, come 
Diceras-beds, containing Cardium corallinum and Corbis gigantea, 
which have not been matched further north. These Diceras-beds 
elsewhere in the neighbourhood are said to be covered by an oolitic 
coral-limestone, very irregular on its surface. This is succeeded by 
marls and marly iimestones, with Ostreu deltoidea and several other 
of the lower fossils of Havre, such as Trigonia Meriani (“‘muricata”’), 
Cyprina cornuta, and Rhynchonella inconstans (?); and after a very 
slight interval come the Virgulian marls. These facts are very in- 
structive, as showing, amidst a general development similar to that 
of Normandy, some of the peculiar features, e. g. the Diceras-beds, 
of the great southern range. 

2. Normandy.—In studying this district it will be well to go, in 
the first instance, to the coast, because there the section is complete 
and has been described by several authors. The earliest description 
was that by Caumont (1), who gives some account also of the inte- 
rior, and recognizes the following subdivisions :—(1) Kimmeridge 
Clay, (2) Glos sand and Blangy limestone, (3) Coral Rag, (4) Oxford 
Clay. A later and more complete description of the coast only is by 
Hébert (28), who gives very detailed sections, and divides the rocks 
studied into (1) Coral Rag, (2) Upper Oxfordian, and (3) Middle 
Oxfordian. A further description of the higher rocks was promised 
at the time, but has never yet seen the light. The continuation 
of the series on the opposite side of the Seine, at the Cap de la 
Héye, has been described by Dollfus (30), who recognizes three 
portions of the Kimmeridgian, as (1) Ammonite-clays, (2) Pteroceras- 
marls, (8) Trigonia-clays and limestones ; and by Lennier (49), who 
gives a detailed section of the beds at the Cap de la Héve, and 
thence to Octéville. 

In the examination of this section the English geologist is at 
once struck with the extraordinary resemblance of the series to those 
at Weymouth and Osmington. So close indeed is it, that almost bed 
for bed can be recognized ; and the whole becomes therefore an admi- 
rable term of comparison between the French and English rocks, 

202 


J. F. BLAKE ON THE UPPER 


548 


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JURASSIC OF THE PARIS BASIN. 549 


if only the development in each country could be correlated respec- 
tively with these. 

The cliffs of Auberville are mainly composed of fossiliferous Oxford 
Clay (known as Argile de Dives); and Caumont’s section, in which he 
indicates the presence of Coral Rag, Upper Calcareous Grit, and Kim- 
meridgeClay beneath the Cretaceous rocks, is quite deceptive. Hébert, 
however, gives details of 33 beds seen in the central portion of the cliff, 
all referred by him to the Middle Oxfordian. His lower 19 divisions, 
comprising 195 feet of clays, with various bands of nodules, would 
correspond to the Weymouth Oxford Clay. To within 60 feet of the 
top this is pure clay, and bands towards the base are very fossiliferous, 
the horizon being indicated by such fossils as Amm. Lamberti, A. 
arduennensis, Turbo Meriani, Modiola imbricata, Nucula ornata, and 
Tthynchonella varians. These are succeeded by numerous bands of 
ferruginous oolite, scattered in the midst of the clays through a 
thickness of about 20 feet, after which the clay still continues for 
another 40 feet. The whole of these have Ostrea dilatata in great 
abundance. One might be tempted to regard the ferruginous bands 
as corresponding to the Nothe grits; but the latter are better 
represented by the mass which succeeds the Oxford Clay. This 
commences with a bed of oolitic grit of about 5 feet, then more 
clay for 5 feet, and then 12 feet of strong ferruginous oolite and 
grit, which is marly in parts, and has a lumachelle of small oysters 
at the top. ‘These beds are very constant: they may be traced 
all along the cliff of Auberville as far as Villers, and may be seen 
again at low tide on the shore east of Trouville. At this last 
locality the lower beds contain Ammonites plicatilis? and Pecten 
jfibrosus. ‘The intervening clays make no show; and the upper part 
are dark-brown oolitic grits, exactly similar to those at Osmington, 
erowded with large Gervillia aviculoides, Trigonia perlata, and 
Pecten fibrosus ; towards the top is a bed of Serpula tricarinata, 
and, finally, a regular Trigonia-bed like the lower one at Osmington. 
These are the beds Nos. 20-29 of Hébert’s Auberville section, and 
Nos. 2 and 3 of his Trouville section, which is rather differently 
described. The principal fossils noticed, besides those mentioned, 
were Astarte ovalis, Cerithium muricatum?, Unicardium globosun, 
Pleuromya tellina, and Hxogyra spwralis. The next portion of the 
series corresponds to the Nothe clays. It consists at Auberville of 
blue clay at the base with hard bands and fossils, and at the top of 
purple unfossiliferous clay, making a total of 6—8 feet. At Trouville 
the base is darker, and contains lumachelles of Hwogyra nana. There 
is a hard blue cale-grit with fucoidal marks at the top, which, from its 
position, may be taken as the representative of the Bencliff calc-grit. 
These are Nos. 30-32 of Hebert’s Auberville section, and apparently 
No. 3of his Trouville section, though the grit is not noticed. Above 
these come about 6 feet of alternations of clays and ferruginous oolitic 
bands at Trouville, scarcely recognizable at Auberville, in which 
Exogyra nana and Pecten fibrosus are still abundant, with Trigonia 
corallina, Pecten lens, Lucina Moreana, and Pseudodiadema sp. 
These must be reckoned with the succeeding beds, which are perfect 


550 ; J. F. BLAKE ON THE UPPER 


representatives of the Osmington volites. Like these same oolites 
as seen at Weymouth and Osmington, they are in one place more 
compact than in another. They are large-grained and more or less 
rubbly at Trouville, especially towards the base, and in alternating 
bands up to the top, where they are capped by a pisolite full of Verinee, 
forming a very characteristic horizon. At Auberville they are like- 
wise very oolitic towards the base, and this is the only part of them 
which is anywhere seen in the cliff; but as they have a dip towards the 
S.E., the upper portion 1s exposed in stream- and road-sections near 
Villers, beneath the Cretaceous strata, andis seen to consist of a fine 
massive oolite, perhaps as much as 20 feet in thickness, with scarcely 
any fossils except Pecten qualicosta. The more pisolitic beds, however, 
are crowded with fossils everywhere, the most abundant species being 
Echinobrissus scutatus, Opis Phillips, and Cerithium mwricatwn. 
Other fossils are Chemnitzia heddingtonensis, Pleuromya Voltzia, 
Nerinea elongata (or ? fascrata), Phasianella Buvigmert, and Lima 
subantiquata. Above this, at Hennecqville, are seen blackish oolitic 
beds, of no great thickness, alternately hard and soft, with Trigonia 
clavellata and Gervillia aviculordes, which may he paralleled with 
the Trigonia-beds of Weymouth. ‘The series above is variable, as 
usual with the Coral Rag. At Hennecqville there are 18 feet of dark 
unfossiliferous clay, rather sandy at the top, and rapidly changing in 
character towards Trouville, where it is almost concealed. Hébert 
mentions that at Villers, in some locality which is now imaccessible 
to observation, marls and limestones with Ctdaris florigemma succeed 
the oolites. Above this clay at Hennecqville the beds become oolitic 
again through 12 feet, and very similar to those below*, most fer- 
ruginous at the base, and cavitary towards the top by the decay of 
fossils, the most abundant of which here are Opis corallina and 
Nerinea fasciata?. There are also Cerithium muricatum, Littorina 
murieata,and [sodonta Deshayesia. The capping of this mass for about 
1 foot at Hennecqville is a ferruginous suboolitic limestone, full of 
fossils, especially Cedaris florigemma and some corals—also Nerina 
imbricata, Opis corallina, Cypricardia isocardina, Pleuromya tellina, 
Pecten vinmneus, P. qualicosta, Lima proboscidea, Exogyra nana, &e. 
This is the only representative of Coral Rag at this spot, where the 
clay is thick; but as we pass towards Trouville all that hes above 
the Osmington Oolite rapidly changes its character and becomes more 

calcareous. ‘The last exposure fomande the town shows massive lime- 
stones with rubbly Ovdaris-florigemma beds towards the top, through 
an aggregate thickness quite as great as at Hennecqville (31 feet) ; 
and in a large quarry in the town are seen beds which must be the 
representatives of this portion, though very dissimilar. Here at the 
top is a rubbly coral-reef mass, utterly irregular atits base, lying in 


* This clay and oolite are so similar to the clay and oolite below, that it is 
possible one may be deceived by slips, and they may be actual repetitions. At 
no place can the beds above the Lower Oolite be certainly determined ; but at 
Trouville they actually lie at a considerable depth below the beds saith Cidaris 
florigemma, which latter are underlain by a more compact limestone. If there 
be a slip, it is a very uniform one and ‘parallel to the face of the cliff. 


JURASSIC OF THE PARIS BASIN. 551 


ereat depressions of the subjacent limestones, and at places 20 feet 
thick. ‘The corals are Thamnastrea and Calamophyllia ; and Crdaris 
florigemma and C. Smithw also abound, with Littorima muricata, 
Pseudomelania Coquandi?, Natica corallina, Pholadomya Tombecki, 
Corbicella sp., Ostrea gregaria, and Acrosalena sp. Underlying 
this are 20 feet of shelly oolites, in which the fossils are badly pre- 
served, Trigona corallina(?) and Opis corallina being the chief noted. 
Below these again are 4 feet of rubble (with Hinnites velatus), and 
then 3 feet of oolite. This quarry does not appear to have been de- 
scribed before ; but it is of considerable importance as showing the 
rapid development of coral-growth, since, not a mile away, only the 
feeblest representative is seen in the cliff. In the other direction, 
towards Auberville, however, it becomes perhaps of still greater im- 
portance ; for at Benerville admirable coral-masses are developed, and 
slip down onto the sea-shore. These are of considerable thickness, 
and contain a crowd of all the usual Rag fossils, as at Trouville; they 
are underlain by oolitic stone and unfossiliferous clay. The whole 
mass, however, is out of place; and no stratigraphical data are obtain- 
able. Hébert does not notice this, though he describes the clay 
onto which it has slipped as crowded with Oxfordian fossils. At 
Villers only fragments are now visible; and, according to Heébert’s 
section, the reef is dying out; for he indicates only about 16 feet 
of limestones and marls, including the florigemma-rag. From the 
position which this rag occupies with relation to the beds below, and, 
as will be seen, to the beds above also, it follows that it occupies the 
place of the “ Sandsfoot Clay,” and possibly of the lowest bed also of 
the “‘ Sandsfoot Grit,’ many of the fossils of which are common to it. 
It is, however, so intimately connected with the subjacent oolites, as 
to make it difficult to range the two in different subdivisions of the 
geological series. 

Above the representative of the Coral Rag at Hennecqville, where 
alone any higher beds are seen on the coast, come calcareous grits 
and sands, admirably representing (as will be seen) the Sandsfoot- 
Castle beds. For the lower 15 feet they are marly sandstones with 
Pseudomelania striata and a narrow Nerinca; then comes a hard 
calcareous grit, called by Caumont the Blangy lmestone, from its 
being worked at a village of that name; and above this a series of 
beds varying very much in thickness, containing, towards the base, 
white hummocky calcareous grits with intervening yellow sand and 
beds of Tregonia muricata ; then fucoidal grits and sands, with great 
masses of black flints, dying out to nothing (one was noticed 3 feet 
thick by 16 feet long); and at the top honeycombed masses of soft 
and hard sandy beds, the cap being especially hard, and serving as 
a ledge on which the overlying clay appears to rest conformably, 
though it has doubtless slipped down and covered the intervening 
beds seen further east at Villerville, and blocks of which here strew 
the strand. The whole thickness of these is nearly 40 feet; and 
they contain also Ammonites varicostatus, Chemnitzia delia, and 
Pinna granulata. 

So far the section, instructive both by its resemblances and by 


Ou 


52 J. F. BLAKE ON THE UPPER 


its differences when compared with that at Weymouth, has been 
described before; but the succeeding beds which connect it with 
the section at Havre, and complete the similarity of the whole 
to the English series, appear to have escaped description. So com- 
pletely, indeed, have they done so that Hébert (see 41) states that 
the Trigonia-grits of Havre are separated from the base of the 
Kimmeridge Clay by 160 feet of marls which overlie the Corallian 
beds at Hennecqville, these 160 feet being obviously the overlying 
clays which have slipped forward and covered the grits which are 
seen i sitw further on. Though of considerable thickness, they 
have been subjected to so much disturbance that they make but 
little show even when exposed, as on the shore on either side 
of the village of Villerville. First, overlying the last-named grit of 
the Corallian series, which forms an admirable base line, are about 
8 feet of marls; and next a series of remarkable hard beds filled 
with small shells. These extend through about 4 feet, and are 
flaggy; some of the bands are crowded with a small Cerithium ; 
others have abundance of Astarte supracorallina, Cardium deli- 
batum?, and Haoyyra bruntrutana. Next come perhaps 10 feet 
(more or less) of grey marls; over which lies, apparently an situ, a 
gritty bed full of black pebbles; then more marls, 8 feet to 10 feet ; 
and finally, in this locality, 3 feet 6 inches of oolitie ironstone 
and associated marls with Pecten midas. Allthese are seen together 
to the west of Villerville. On the east side, presumably the same 
oolitic ironstone bands are seen in slipped masses on the strand, 
followed in an upward direction by flaggy sandy marls full of 
Pleuromya Volizi; then some more marls, 8 feet; next a bed of 
small oysters (Hvogyra nana or bruntrutana), 1 foot; next 3 feet of 
soft marl; and then the remarkable sandy grit with Tragome, so 
well known at Havre and Weymouth, about 1 foot in thickness ; 
above which come about 4 feet of clays with two bands almost made 
of O. deltoidea; and finally the ordinary completely argillaceous 
Kimmeridge Clay, which so often slips down and masks the beds just 
described. If any one compares this section with that at Sandsfoot*, 
he will see an almost exact identity, in which the ironstone, the 
lumachelle, and the Trigonia-grits hold their relative positions and 
almost their distances—the only difference being that the O.-deltoidea 
beds appear here to lie, as they are said to do at Havre, above the 
Trigonia-erits instead of below. ‘The fossils also are equally charac- 
teristic, those noted being Ammonites cymodoce, Belemnites mtidus, 
Inttorinma pulcherrima, Natica eudora, Pholadomya “ Protet,” Opis 
angulosa, Astarte Michaudiana (called ovata before), A. trigonarun, 
Cyprina Constantin, Cyprina cyreniforms?, Trigoma Meriani, 
I’. muricata, T. papillata, Mytilus peetinatus, Pecten midas, P. swpra- 
jurensis, P. minerva, Gervillia kimmeridiensis, Serpula gordialis, S. 
qunquangularis, S. tetragona, and Monilivaltia Lesueurn. The pre- 
sence of this latter fossil 1s of interest as showing that coral-growth 
had not died out here; for it must be to the lower portion of this 
series that the ‘“ Upper Coral Rag” seen in Ringstead Bay must be 
* Q. J. G.S. vol. xxx. p. 241, and vol. xxxili. p. 270. 


JURASSIC OF THE PARIS BASIN. aes 


referred, which is, at the same time, proved to be on a much higher 
horizon than the ordinary Coral Rag of Normandy. 

The section thus concluded at Villerville can be perfectly taken 
up on the other side of the estuary at Havre; for the “ Trigonia- 
erit” at the former is simply the continuation of the “ Calcaire a 
Trigonies” at the latter, of which the fauna has been so admirably 
illustrated by Dollfus. ‘These, including the associated marls and 
other hard bands, up to and including the grey clay over the fossili- 
ferous grit, are therefore the Kimmeridge passage-beds, as was 
ascertained by Waagen (35). Itis also obvious-that by their stra- 
tigraphical position and by many of their fossils they represent the 
Astartian beds of the rest of France *. Of the overlying beds at 
Havre little need be said, since their position cannot be otherwise 
than clear. ‘The beds included between No. 4 and No. 14 of Len- 
nier's section, the latter being a hard band with many gasteropods, 
represent Dollfus’s “* Marne a Pteroceres,”’ and the basal portion of 
the Lower Kimmeridge of Weymouth. In Normandy there is 
certainly much more reason for establishing a “ Pterocerian stage,” 
since Pteroceras Ponti is a very common fossil both at Villerville 
and at Havre, and has associated with it a sufficiently distinct 
fauna. It is therefore perhaps only due to the imperfect searching 
of the beds at Weymouth that a similar fauna has not been dis- 
covered there. Of the fossils in Dollfus’s list which are found only in 
the Pterocerian beds, the following occur in the Lower Kimmeridge 
of Weymouth—dAmm. decipiens, Pleuromya tellina, P. donacina, 
Pholadomya acuticosta, and Ostrea solitaria. Only one of these is 
at all characteristic ; and that is Phol. acuticosta, which, as we have 
seen, is a constant species in the Virgulian marls, whether these are 
divisible into two parts or not. In fact the fauna consists almost 
exclusively of Myacidze, Gasteropods, and Echinoderms, of which 
the latter two groups are so remarkable tbat they could scarcely 
escape attention if they occurred at all freely at Weymouth. Hence 
though the beds must correspond stratigraphically, we have no en- 
couragement in this country to recognize a ** Pterocerian” subdivision. 
Two other fossils may be noted. Avicula edilignensis occurs in these 
beds, which may hence be a guide to correlation: and about halfway 
between the Trigonia-grits and the Gasteropod-bed are bands full of 
Terebratula Leymeru, with associated nodular beds. It would 
therefore be perfectly defensible to raise the upper limit of the 
Astartian to this level, except that 4. virgula has already begun to 
be abundant. The “ Ammonite clays” of Dollfus present no points 
of interest, as they are but poorly seen and slightly fossiliferous, and 
they do not attain to the Upper Kimmeridge. 

The only other locality examined in Normandy is the neighbour- 
hood of Lisieux and Glos, which presents us with a greater develop- 
ment of a portion of the series seen at Trouville. This locality is 
included in the general description of Normandy by Caumont, who 

* This is not the conclusion I arrived at in my paper on the Kimmeridge 


clay ; and the correction shows the advantage of a personal over a literary 
acquaintance with the French series. 


554 J. F, BLAKE ON THE UPPER 


gives a section of the hill of Glos; and it is specially described by 
Goubert (29), who, with Zittel, illustrated the fossils found in the 
sands. Near Lisieux, on the road to Glos, is a quarry very similar 
to that at Trouville; for at the base is 6 feet of rubbly limestone, 
then 5 feet of a large-grained oolite, and on the top 20 feet of rubbly 
coral rag, almost a Thamnastrean reef in places, and containing 
Cidaris florigemma abundantly. This, therefore, represents the 
true Coral Rag. At a distance of about 3 kilom. from this quarry, 
on the north side of the bridge leading to Glos, is a very instructive 
exposure. At the base we have a rubbly coral rag of different 
character, and obviously either overlying the former, or a changed 
development of its upper part. ‘The beds are all vacuous by 
the decay of fossils, only internal and external casts being found. 
The corals are Calamophyllian, and not Thamnastrean: and there 
is an abundance of Nernea Goodhallu, with Littorina muricata, 
Cerithium muricatum, Trochotoma discoidea, Chemntza dera?, 
Natica corallina, Nerita sp., Lucina Moreana, and L. balmensis? 
The whole, by the absence or rarity of Cidaris florigemma, by the 
abundance of Nerinewe, and by the character of the corals, reminds 
one of the Novion hmestone, though the lithology is different. Over- 
lying this are oolites and oolitic sands in good beds, 34 feet; next a 
breccia of limestone-fragments with a hard 15-inch band of blue 
limestone in the middle, with Plewromya tellina, a total of 3 feet 
9 inches; and then the true sands of Glos. These are marly at the 
base, but become more sandy by degrees, with hard glauconitic 
bands or nodules containing fossils; but finally the sand is abso- 
lutely unfossiliferous, as it has usually been reported to be at this 
spot. The thickness here seen is 24 feet; but it certainly extends 
much further up the hill. The fossils noted are Ammonites serratus, 
Pterocera polypoda, Trigoma Brenna, Gervillia kimmeridiensis, 
and Pecten midas—a group rather characteristic of higher beds 
than these are usually considered, and which leads us to look on 
these sands as partly equivalent to the passage-beds of the coast, 
especially as Caumont records a section at Pont l’Evéque where the 
Trigonia-grit les immediately on the sands of Glos. The quarry 
just described does not appear to be the place where the usual fossils 
have been gathered, as they are said to be all small except the 
Trigonia. The common locality in fact is seen on the opposite side 
of the river, where a greater thickness of sand is seen containing 
bands white with fossils, the principal of which are Trigonia Bronnw 
and Lucina circumcisa. In any case, the exact position at which 
the sands commence in reference to the coast-section is clear, the 
lower beds in the quarry representing the first beds above the Coral 
Rag, and the blue lmestone the hard band (called ‘Calcaire de 
Blangy’’) in the Hennecqville Chiff. 


IV. Tue Pays bE Bray. 


The older works on this district, such as that of Graves (8), have 
been rendered out of date by the splendid monograph of M.de Lap- 


JURASSIC OF THE PARIS BASIN. 555 


parent (69), published as one of the memoirs of the Geological 
Survey of France. Only a small portion of the Upper Jurassic 
rocks is exposed in this district, brought up to day by a N.W. 
and S.E. elevation, from the summit of which the Cretaceous rocks 
have been worn away. It thus affords, as it were, a continuation 
of the coast-section of Normandy, commencing at the base where the 
latter ceases, namely in the Virgulian marls. M. de Lapparent 
classifies the rocks thus exposed as follows :-— 


Upper PorTnanp. 
Ferruginous sandstone, speckled clays, greensand. 


Mippie Portuannd. 
Blue marls. 


LOWER PorTLAND. 


1, Upper conglomerate. 

2. Glauconitic calcareous grit. 
3. Marly limestones. 

4. Calcareous grit with Anomie. 
5. Beds with Ostrea catalaunica. 


KIMMERIDGIAN. 


1. Upper clays and lumachelles. 

2. Compact lithographic limestone. 

3. Lower clays and lumachelles. 

4, Calcareous grit (Pterocerian or Astartian ?). 


The accuracy of this description has been verified so far as may be 
in a brief visit, and the correctness of the correlation determined— 
with one important exception, which may be the subject of discus- 
sion. According to verbal communications from M. de Lapparent, 
borings executed between the Pays-de-Bray and Boulogne have 
shown Cretaceous rocks lying directly on Paleeozoic ; so that we are 
not led to expect the Jurassic rocks to be continuous between the 
two districts. Nevertheless, as we have seen, the lower portions of 
the deposits under study are remarkably similar to those of England ; 
and as the true Portland rock of England reaches to Boulogne with 
little change, we may expect the intermediate beds to be also similar 
to those of England and differing from them in the direction of 
Bolonian characters. Judged in this way, the so-called “ Lower 
Portland” (used in the Bolonian sense) of the Pays-de-Bray should 
begin to show its episodal character somewhat lower in the series. 

The lowest bed visible, the sandy calcareous grit, is a very 
doubtful rock, seen only at one place. One cannot be sure that it is 
im situ; and its badly preserved Trigonie and Astartw suggest that it 
may be a mass belonging to No. 3 of the “ Kimmeridgian.” In any 
ease it has no particular resemblance to the Trigonia-grit of Havre, 
and cannot lead to the inference that we are here low down in the 
Kimmeridgian, as we should be if it belonged to the “ Pterocerian 
or Astartian.” The ‘‘ Lower Clays and lumachelles” with Kwogyra 
virgula present nothing worthy of note, except that towards the 
south they have a thickness of 200 feet, and on the opposite side of 
the same valley between Louvicamp and Mesnil, which contains the 


556 J. F. BLAKE ON THE UPPER 


Fig. 15.—Map of Part of the Pays de Bray. (Scale 1: 240,000.) 


¢ See . Zam Sn 
5 Sree ) SS ; 
fim Cay). LE EX 


ya ae 5 


grit above discussed, they are not much thinner. The upper portion of 
this, in which the lumachelles and the intervening material become 
very sandy, requires more discussion. M.de Lapparent says :—“ At 
the top of this system the lumachelles grow together into solid banks 
intercalated with very sandy material, and forming rather irregular 
masses than continuous beds.” ‘These sands and doggers are well 
i! seen to the north of Louvicamp. The sand is pure; and the doggers 
i are huge; and at the top is a lumachelle of Trigonie, which appear 
| to be 7’. Muniert. We have therefore the exact representative of the 
if sandy beds at Gris Nez and Mt. Lambert in the Boulogne area. The 
li same is exposed in the railway-cutting south of Haussez, where the 
beds seen are 10 feet or 12 feet of grits and sands, thickening to the 
south, and with abundance of Hwvogyra virgula. ‘The presence of this 
last fossil may at first appear a stumbling-block; and it undoubtedly 
proves the close connexion of these rocks with the Virgulian; yet, 
when we compare the Boulogne district, it proves nothing, since the 
| species is equally abundant in the sandy beds which in that area are 
iy referred to the “Lower Portland;” on the other hand the petrological 
ir similarity is of great weight. Overlying these comes the ‘Compact 
hy Lithographic Limestone,” a rock which can be scarcely be matched 
either in England or at Boulogne. It has a thickness of certainly 
more than 12 feet, and contains lumachelles of Trigonie too deeply 
imbedded for the species to be recorded. At Louvicamp it has also 
lumachelles of Kxvogyra virgula. The two fossils mentioned by M. de 
Lapparent are Ammonites cf. gigas, too characteristic a fossil to be mis- 
taken, and Gervilla kimmeridiensis, which might well be G. linearss, 
so similar are the species of this genus. Next comes another mass of 
clays which are said to be likewise 200 feet thick in the south, but are 
Ad certainly not so much in the central portion, where examined. There 
abs are few fossils in them beyond Lwogyra virgula, which occurs in luma- 
: chelles. It is seen underlying the cement-stone beds of the cutting 
at the station of Saumont-la-poterie. These cement-stones are 
regarded by M. de Lapparent as forming the base of the ‘‘ Lower 
Portlandian ;” and he states that Hwogyra virgula is entirely and 
suddenly replaced in them by Kw. catalaunica. ‘This is certainly 
A not the case: after the cement-stones begin there are no more 
lumachelles; but Ha. virgula occurs in tolerable abundance in the 
marls above, at least 6 feet of them. ‘This fossil, therefore, is no 
more available here for drawing a line at the base of the ‘‘ Lower 


~ 


JURASSIC OF THE PARIS BASIN. 557 


Portland” than itis at Boulogne. The fossils of these cement-stones 
are abundant; they are Ammonites swprajurensis, Natica athleta?, 
Pleuromya stnuosa, Thracia incerta, Cyprina elongata, C. implicata ?, 
Trigonia (clavellate sp.), Mytilus autissiodorensis, and Anomia lavi- 
gata. The lithological character of these limestones, and the abun- 
dance of Mytilus autissiodorensis and of T'hracia incerta, lead one in 
the first instance to recognize in these beds the cement-stones at the 
base of the Portland sands; but in face of the abundance of Ex. vir- 
gula and the presence of Amm. suprajurensis, it is safer to look upon 
these as the more marly portions of the “ Lower Portlandian,” such 
as are associated with the conglomerates at Portel and are abundant 
with the same fossils in Kimmeridge Bay. These cement-stones 
are cut off from less fossiliferous marly beds above by an inter- 
mediate bed of “ Calcareous Grits with Anomie.” These two, with 
the ‘‘ Glauconitic Calcareous Grit” and ‘‘ Upper Conglomerate” form- 
ing the cap, will then represent the uppermost portion of the Bolonian 
“¢ Lower Portland ” with Pteroceras oceant. That fossil is not recorded 
from the Pays de Bray; but the occurrence of Trigonia boloniensis, 
Pecten nudus, and a Hemeidaris called H. Hofmanni, but perhaps 
really H. purbeckensis, is highly characteristic. Of the upper beds 
little is now visible. The blue marls, described by de Lapparent as 
dark at the base, with Cardiwm morinicum and large Ammonites 
(? biplex), and calcareous towards the top, with Amm. biplea, Pleu- 
rotomaria Rozeti, Ostrea expansa, Perna Bouchardi, and Cardium 
Pellati, are obviously representatives of the ‘“‘ Middle Portland” beds 
of Boulogne, and are more similar to them than to the corresponding 
English series. Finally the ferruginous sands seen at Gournay are 
certainly distinct in appearance from any thing seen elsewhere: and 
they contain few if any fossils. At their base, however, they contain 
huge calcareous-grit doggers, which resemble very much those of 
Swindon and Shotover, the corresponding beds at Boulogne being 
more regular and calcareous. These contain clavellate Trigonie, 
Astarte, Corbula, and Thracia incerta? It is above these beds that 
ferruginous nodules are found, which contain Trigonia gibbosa and 
other true Portlandian Trigome, e. g. T. oncurva, the specimens of 
which from the Normanville cutting, near Gaillefontaine, preserved 
in the Ecole des Mines at Paris, are undoubtedly the true forms, 
though they can no longer be found in a recognizable state. While, 
therefore, the lower part of the sands and grits may well be the 
equivalents of our “ Portland Sands,” the upper part must at least 
attain the horizon of the Flinty series of Portland. 


V. Tue Bovunonnals. 


The coast of this district is almost English ground: so well is it 
known, and so long has it been studied by many of our native geolo- 
gists. They have, however, published little upon it; and we owe the — 
description of the area first to M. Rigaux (33) and Prof. Hébert 
(37), and later to MM. de Loriol and Pellat (86, 39, 41, 56, 61), 
the last-named author having given a final réswmé of his views in 
1878 (68). M. Pellat’s classification is now as follows :— 


508 


J. F. BLAKE ON THE UPPER 


Urrrer Portnuann. 
. Cypris-beds, 25 metres. 
. Siliceous limestone with Cardiwm dissimile, 24 m 
. Sands and grits with Natica elegans &e., 4 m. 
. Sands and cale-grits with Cardiwm Pellati, 3m. 


MippLE Portnanp. 
O,. Clays and glauconitic limestones with Ostrea expansa, 13 m. 
O,. Clays with Cardiwm morinicum, 15 m. 
Lowrr Portianp. 
Zone of Cyprina Brongniarte. 


N,. Sandstones with Preroceras oceani 10 
N,. Sands with Nautica Marcousana a 
N,. Conglomerate with Trigonia Pellati of Chatillon. 


Zone of Ammonites gigas. 


N,. Sands and grits with Amm. portlandicus, '7 m. 
M,. Shales and upper cale-grits with Hemicidaris purbeckensis, 17 m. 


Upper KIMMERIDGIAN. 
Zone of Anvmonites erinus. 


M,. Shale and lower limestones with Amm. erinus, 12 m. 
L. Sands and grits with Pygurus, 43 m. 


Mipp.ie KIMMERIDGIAN. 


Zone of Ammonites caletanus. 


Clays and upper lhmestones with Amm. caletanus, 18 m. 
. Sands and grits of Conincthun, 2 m. 


aia 


Lower KIMMERIDGIAN. 
Zone of Ammonites orthoceras. 


I. Clays and lower limestone with Amm. orthoceras, 22 m. 
H. 138 beds and little beds of Bréqueréque, 15 m. 


SEQUANIAN. 
G. Sands and grits of Wirvigne, 5 m. 
F,. White marls and oolites of Bellebrune, 4 m. 
. Clays with Ostrea deltoidea, 2 m. 
. Pisolite with large Nertne@e of Hesdin-l Abbé, 6m. 
E. Red limestone, sands and grits, with Triégonia Bronnii, 5 m. 
D. Clays with O. deltoidea of Brucdale and Mont des Boucards, 10 m. 


CorRALLIAN. 


A'. Coral Rag with Cidaris florigemma of Brucdale. 

Limestones of Mont des Bou- 
cards = Pyritous claysand lime- 
stones of the South of the Bou- 
lonnais, 50 m. 


Limestones with J/socardia. 
. Limestones with Jerebratule. 
. Coral Rag with Cid. florigemma. 


HO 


Upper OXxForDIAN, 


Houllefort limestone with Psewdomelania heddingtonensis, 1 m. 
Clays with Ammonites Martelli and sponges, 14 m. 


Mippue OxrorDiANn. 


Clays and limestones with Ostrea dilatata, 5 m. 
Black clays of Wast, 6 m. 


Lower Oxrorp1An (Callovian). 
Fissile limestones with Ammonites Lamberti, 2 m. 


Clays with Ammonites Duncani, 6 m. 
Ferruginous clays with Ammonites calloviensis, 5 m. 


ee 


JURASSIC OF THE PARIS BASIN. 559 


This classification is the result of several modifications of pre- 
vious ones, especially in the matter of names. 

The Upper Jurassic strata in the Boulogne area lie in an anti- 
clinal whose crest has a slope from E.N.E. to W.S.W. Hence the 


Fig. 16.—Map of Part of the Boulonnais. (Scale 1: 240,000.) 


Zz 


argunse 


OW. Bein ehurs 


lowest strata are seen in the E.N.E. corner, and the succession may 
be traced by passing thence to the sea at Boulogne, whence a diver- 
gence to the right or left will show the uppermost portions. The 
development is also greater towards the 8.W., as in the contrary 
direction we approach the shore-line. From M. Pellat’s beds E., 
or the base of the Nerineean limestones, downwards, the prevail- 
ing character is argillaceous, and the comparatively feeble lime- 
stones are merely intercalated masses, themselves usually more or 
less marly. This feature renders the correlation somewhat difficult, 
and has given rise to the complicated classifications of MM. Pellat 
and Rigaux. 

The lower groups of clays, estimated by M. Pellat at 26 metres 
in the north, but at 47 metres in the south-west, have not been spe- 
cially examined. The uppermost portion, with dAmm. cordatus and 
Millericrinus, doubtless corresponds to the Nothe grits and clays, and 


560 J. F. BLAKE ON THE UPPER 


to the beds with Trigonia spinifera at Neuvizy, and with Trigonia 
perlata at Trouville. The first mass of importance is the limestone 
of Houllefort, which is perfectly isolated in the midst of clays, some of 
which separate it above from the so-called limestones of the Mont - 
des Boucards*. It is extremely fossiliferous ; but the discovery of 
the fossils is dependent on their weathering out, and would be very 
unlikely in a boring. ‘Twenty-six species are enumerated by de 
Loriol from this limestone, and additional ones by M. Pellat. Those 
actually noted were Trochus houllefortensis, Purpura sp., Cerithium 
Struckmanni, Chemnitzia sp. (wrongly called Pseudomelania heddingto- 
nensis), Opis Phillips, Arca scabrella (near to A. quadrisulea), Nucula 
cottaldina, Lama rudis,and Cidaris florigemma. There is so much that 
is peculiar about this fauna that it gives little help in correlation; and 
the additional known species in de Loriol’s list, ¢. g. Amm. plicatilis, 
Alaria tridactyla, Pholadomya Prote and P. concinna, Trigonia moni- 
lifera, Lima rignda, Pecten vimineus, and Ostrea dilatata are not quite 
conclusive. Nevertheless, on the whole, the facies is Corallian, and 
indicates an horizon which may indeed be compared with that of 
Neuvizy or the Osmington Oolite, but is more like the base of the 
Corallian in the Haute-Marne and the Yonne departments. 

About the succeeding portion of the series there have been 
several changes of opinion among the French geologists, owing 
to there being different developments in different parts of the _ 
area. The difficulties arising from this are intensified in the 
Boulonnais by the deposits being exceedingly local and by several 
minor unconformities occurring in the series, owing to the proximity 
of the ancient shores. Four localities exhibit different arrange- 
ments, of which the sequence may be specially noted. First, in 
the extreme north near Bazinghen there is nothing but a few feet 
of clay between the Callovian grit with Terebratula humeralis (true) 
and Rhynchonella varians and the Nerinewan or so-called Astartian 
Oolite; thisisa minimum. Secondly, at the Mont des Boucards, there 
is a maximum; but the whole is essentially argillaceous though dig- 
nified by the name of limestone. Only at one spot on the hill-side 
are seen a large number of loose stones scattered.on so restricted a 
surface as to even suggest the question whether they may not be the 
débris of ancient buildings. These stones, surrounded on all sides by 
the clay, are probably a remanié collection, or are due to some special 
cause which renders their actual position, at a slightly lower level than 
the fossiliferous beds in situ, of little consequence; but their fauna 
is remarkable for its richness and abundance. The list published 
and illustrated by de Loriol is very defective, including only twenty- 
six out of eighty species which M. Pellat has, up to the present, dis- 
covered; and half an hour’s search obtained the following 30, 
several of which are not included in de Loriol’s list. Nautilus 
giganteus, Aporrhars elegans, A. musca, Pleurotomarva hesione, Ani~ 


* Tn his later descriptions (e. g. 56) M. Pellat places the “ Calcaire du Mont 
des Boucards” immediately above the Houllefort limestone; but in his earlier 
ones (e.g. 41) he recognizes that they are separated by a considerable mass of 
clay. 


JURASSIC OF THE PARIS BASIN. 561 


socardia elegans, Trigona monlifera, Arca Sauvage, Cucullea 
(called quadrisulca, de Lor.), Arca rhomboidalis, de Lor., Gastro- 
chena boucardensis, Nucula equilatera, Astarte Sauvage, A. num- 
mus, A. bruta, Lithodomus inclusus (in Isastrea), Modiola cequi- 
plicata, Mytilus pectinatus, Inma proboscidea, Pecten vimineus, P. 
intertextus, Avicula oxyptera, Plicatula horrida, Exogyra nana, 
Osirea rastellaris, Terebratula tetragona?, Rhynchonella corallina, 
Thecosmilia sp., Serpula Royert, Cidaris florigemma, C. Smithii, Pen- 
tacrinus cingulatus ?. These fossils being collected from the surface- 
stones, it is quite possible that some may be derived from overlying 
beds, which would account for their only being recorded by de Loriol 
as from such. But when we consider also the more important fossils 
recorded by him in addition to those above, such as Belemnites nitidus, 
Anatina striata, Astarte Michaudiana, Opis Phillipsi, Arca texta, 
Hinnites fallax, and Hemicidaris intermedia, we see that there is a 
very feeble minority, almost an absence, of Oxfordian species, and that 
the whole indicates a higher horizon than the usual florigemma-rag. 
If it be a remanié deposit, it might well be formed by the breaking- 
up of beds overlying the Coral Rag of Brucdale. The marly beds zn 
situ which, with the above exception, continue upwards at the Mont 
des Boucards the clays overlying the Opzs-limestone, are well seen 
in cuttings and quarries on the hill-side. Only here and there they 
have hard bands; and the term “limestones” is quite deceptive. 
Only one soft chalky limestone caps the whole, and forms the top of C 
of M. Pellat. At the base these beds are crowded with Terebratula 
insignis and Rhynchonella corallina (quite a repetition of the chalky 
limestones of Bourges), and here and there are lumachelles of Ostrea 
nana, and occasional deltoid oysters ; Mytilus pectinatus and Ostrea 
solitaria(rastellaris) are also abundant. The other fossils observed are 
Ammonites boucardensis [?|, Nucula Menke, Cardium Dufrenoycum, 
and Astarte nummus. A total of fifty-one species is recorded from the 
basal portion by de Loriol, an examination of which shows the same 
tendency as the above, namely to associate these with true Supra- 
coralline beds, or at most with the highest part of the Coral Rag, and 
to make it difficult to conceive how any one could have taken them 
for Oxfordian. The upper portion, C of M. Pellat, is, above all things, 
characterized by the presence of Ceromya excentrica, Isocardia striata, 
and Mytilus perplicatus, fossils which are everywhere more Astar- 
tian than Corallian ; the others noted are Arca rhomboidalis, de Lor., 
Cardiumn intextumn, C. orthogonale, Pleuromya tellina, and Exogyra 
bruntutana, which have little effect on the correlation. These marls 
are followed by more dark clays till the ferruginous grits are reached. 
In the third locality, that of Brucdale and the neighbourhood, are 
seen at the base 6 or 8 feet of Coral Rag zn situ, with the Tham- 
nastree in their place of growth ; but the whole thickness is not seen, 
nor any thing below the Rag. The fauna is an ordinary Rag one, incla- 
ding Cidaris florigemma and various Gasteropods and Lime. There is 
certainly some difference between it and that of the coral stones of the 
Mont des Boucards. ‘This is doubtless largely due to want of search 
among the latter; yet what difference there is indicates a higher 


Q.J.G.8. No. 148. QP 


562 J. F. BLAKE ON THE UPPER 


horizon for them than for the Rag of Brucdale; for the fauna of the 
Mont des Boucards contains seven species usually found on higher 
horizons, and that of Brucdale one only, the species of downward ten- 
dency being equally divided. ‘The surface of the Rag of Brucdale is 
eroded; and it was doubtless during the interval thus indicated that 
the peculiar beds of the Mont des Boucards were deposited, while 
the overlying marls spread over both areas. The fourth locality 
is the valley of the Liane near Hesdin-l Abbé and at Outreau, where 
two borings were made, by which it is supposed to be proved that the 
“‘ limestones” of the Mont des Boucards underlie the Rag of Brucdale. 
Beneath the latter,it seems, there were encountered some “ limestones” 
which were supposed to be identical with (51, 53) those of the Mont 
des Boucards ; but when the proofs are examined, these are found to 
consist solely in the fact that some shells were brought up which 
were thought to be the Ceromya of the Mont des Boucards—a fact 
not sufficient, even if the determination were right, to fix the horizon. 
No other fossils appear to have been obtained, which is rather extra- 
ordinary, considering the great abundance of Terebratula insignis at 
the Mont des Boucards. The supposed proof is therefore utterly in- 
adequate, and we may with more probability interpret the boring dif- 
ferently. At the top (see 51, 53) we have, below the alluvium, 22°65 
metres of undisputed beds, down to No. 18; No. 18 is 6-40 metres 
of the Nerinzan oolite of Hesdin-l Abbé; No. 19 and part of No. 20 
(say 3 metres) are the grits with Trigonia Bronnii. The remainder of 
No. 20 are the Ostrea-deltoidea marls, 12:90 metres; Nos. 21-23, 
which MM. Sauvage and Rigaux refer to a coralline limestone, though 
there are no corals recorded, are attenuated representatives of the 
Mont-des-Boucards limestones, not well developed and therefore 
without the Terebratula insignis, 4°75 metres. No. 24 is the Coral 
Rag of Brucdale with Cidaris florigemma, 6:60 metres. Nos. 25-29, 
referred by MM. Sauvage and Rigaux to the Mont-des-Boucards lime- 
stones on account of their containing a Ceromya at Hesdin-l’Abbé, are 
the marls above the Houllefort limestone, containing Cidaris flori- 
gemma and many fragments of fossils; and the poorly characterized 
beds below are the ordinary Oxford Clay. 

The next bed presents no point for discussion ; itis a ferruginous 
erit of little thickness whose characteristic fossils are Trigonia 
Bronnit and Astarte communis (Morine). Its place, its fossils, and 
its mineral character assimilate it at once to the sands of Glos, the 
corresponding beds at Trouville, and the Sandsfoot grits. It is suc- 
ceeded by the most constant limestone of this part of the Jurassic 
series. Varyingin thickness and somewhat in character, this latter 
may nevertheless be easily traced from the extreme north near 
Basinghen to the valley of the Liane. Like the others, it begins in 
the north by being thin and somewhat sandy, with large-grained 
oolites only here and there, and few fossils beyond Terebratula insignis. 
It has gained considerable thickness near the Mont des Boucards ; 
at Paincthun the oolite predominates, and at Hesdin-l’Abbé forms 
thick masses which are extremely fossiliferous, the most noticeable 
species being Nerinewa Desvoidyi, N. cecilia, Pholadomya hortulana, 


JURASSIC OF THE PARIS BASIN. 563 


P. Protei, Ceromya excentrica, Pinna pesolina, Pecten midas, and 
Rhynchonella corallina. From this oolite are also recorded Cidaris 
florigemma and Hemicidaris intermedia. Ail the more important 
fossils except the Pholadomye come up from below and finish in 
this bed. It is, however, coloured on the geological map with those 
above it as belonging to the “ Astartian.” It is denoted by F’ of M. 
Pellat. The succeeding beds F? and F°, are comparatively unfossi- 
liferous ; but their fossils are such as would unite them rather to the 
overlying than to the underlying beds, Ostrea deltoidea and Trigonia 
papillata being the chief recorded. The Grés de Wirvigne, assimi- 
lated by M. Pellat to the Trigonia-grits of Havre, is somewhat dif- 
ferent from them in mineral character, being more siliceous and full 
of oysters; it has a thickness of 6 feet, and is overlain by marly beds 
with hard bands. The fossils observed—Ammonites Berryeri?, 
Purpura sp., Leda venusta, Astarte supracorallina?, Corbula Des- 
hayesea, Trigoma Meriani?, Cyprina cyreniformis, Pleuromya tellina, 
Pecten solidus and P. strictus, as well as others recorded, such as 
Ammonites erinus, Lucina substriata, L. rugosa, Pinna granulata, 
Exogyra virgula, Terebratula subsella, Pygurus Blumenbachii, Rhab- 
docidaris Orbignyana—all have a tendency to confirm M. Pellat’s 
conclusion ; and scarcely one is more connected with beds below than 
with those above. It is possible that these grits and associated 
beds represent more than the Trigonia-grits of Havre and Wey- 
mouth, since there is nothing above them that can be called Ptero- 
cerian. : 

The overlying beds, in fact, are seldom seen. On the hill-slopes 
south of Mont Lambert but little can be made of them; and the 
same is the case in the north, near Bazinghen. At Boulogne they 
are hidden beneath the town, and merely a part is exposed at Bréque- 
recque, .nd contains no fauna which can be specially called Pteroce- 
rian. It is perhaps scarcely worth while to separate these beds from 
the overlying ones, called H and I; but we may usefully charac- 
terize this lower mass by the presence of Ammonites orthoceras, which 
is associated with Thracia depressa, Lucina minuscula, Pholadomya 
acuticosta, Arca texta, Gervilia kimmeridiensis?, and Anomia sp.; 
it then represents the zone of that Ammonite, and will be assi- 
milated to Pterocerian beds elsewhere. The succeeding series 
commences with a very characteristic group of rocks, part of which 
forms a great bank with white veins on the shore north of Boulogne ; 
and 18 feet of it is quarried along the sides of the great valley be- 
tween Mont Lambert and the sea. It is everywhere fossiliferous, 
the most abundant fossils being Ammonites longispinus, Trigonia 
Rigauxiana, Pholadomya acuticosta,Gervilliakimmeridiensis, Exogyra 
virgula, Terebratula subsella, and Serpula tetragona. This bed forms 
the base of a series of great thickness, in the centre of which comes 
the conspicuous mass of soft sand with large cheese-doggers (L of 
M. Pellat, 33 and 34 of M. Hébert), which is only seen at La Créche. 
This makes no show on Mont Lambert, and is probably quite a local 
deposit. For purposes of comparison it is well to pass over these 
minor distinctions, and to place all the mass K LM of M. Pellat in 

: 2P 2 


564 J. F, BLAKE ON THE UPPER 


one division, or zone, of Amm. longispinus. The upper portion is 
very shaly, and contains, on both sides of Boulogne, Lingula ovalis, 
Trigoma variegata, and Thracia depressa. The upper limit of this 
is assumed by M. Pellat to be the remarkable lumachelle of Hxogyra 
virgula, which stands out as a broad band in the cliff of Chatillon, 
but which unfortunately is absent from La Créche. The base of 
the ‘‘ Portland” was originally taken by M. Pellat (39), as by all 
others, below the great conglomerates ; he subsequently included all 
M as “ Portlandian ” (62), but finally (68) took only the upper halt. 
Certainly the beds become sandy by degrees, and, for some distance 
below the conglomerates, begin to indicate coming changes, as may 
be seen both at La Créche and Gris Nez. This change, being 
accompanied by the introduction of a new fauna, including Ammo- 
nites gigas, Cardium morinicum, Trigonia Mumeri and T. barrensis, 
Perna Bouchardi, and Hemicidaris purbeckensis, may well justify 
the lowering of the line of separation, as we so often have to do, 
the paleontological change being accomplished more quickly than 
the lithological. 

The next portion of the series is perhaps the most interesting of 
all; and as it is chiefly exposed along the coast, we here reap the 
benefit of M. Hébert’s admirable and accurate researches, which 
may be referred to for the closest details (37). He describes the 
three chief localities where these rocks may be seen, viz. :—to the 
south of Boulogne, from Chatillon to Kquihen; to the north, from 
La Créche to Wimereux, and at Gris Nez. The shales last noticed 
are the No. 26 of his sections, and from their thickness form an 
admirable base. The lowest bed of the ‘“ Lower Portlandian” at 
La Créche is a 3 to 4-foot band of excessively hard crystalline cale- 
grit, blue in the interior and without fossils ; it is followed by grey 
fucoidal doggers in sands or clays (not well seen) for 10 feet or 
more; above come 4 feet of hard yellow sandstone rock, full of a 
quartzose conglomerate, and a perfect lumachelle of Hxogyra virgula, 
covered by a layer crowded with Trigoma Pellats and T. Mumeri, 
and then passing into 4 feet more of ferruginous conglomeratic grit 
with the same fossils. These are Nos. 23-25 of M. Hébert. Seeking 
these on the Portel side, we find the crystalline grit attached here 
and there to the base of the great conglomerate, but very irregularly; 
towards the north it sinks by degrees till, opposite the Fort du Mont 
Couple, it is on the sea-shore. Here it encounters a considerable 
fault, running nearly parallel to the cliff, and just cutting behind the 
nearest cliff-quarries of Chatillon*. It is thus thrown to the top of 
the cliff, and forms the base or perhaps the greater part of the 
materials worked there, being in both places crystalline and ligni- 
tiferous, and overlying the shales more or less unconformably. 
Above it comes a variable mass of sands and clays, with huge grit 


* T at first thought the appearances were due to a local erosion and an over- 
lap of the Chatillon beds; but MM. Sauvage and Rigaux suggested a fault, 
the direction of which was made clear by the observations of Prof. Prestwich 
during an excursion of the Geological Society of France in September 1880. 


= 


JURASSIC OF THE PARIS BASIN. 563 


doggers, often becoming continuous, which develop to a considerable 
thickness at Chatillon, are worked there and at Mont Lambert, and 
constitute the base of Gris Nez. These are capped by the great 
ripple-marked conglomerate, distinguished everywhere by the abun- 
dance of Hwogyra virgula, Trigonia Pellati, and 7. Mumeri, which 
becomes, however, a mass of false-bedded sand with hard doggers 
at Gris Nez. These are so associated with the sands below, that 
the natural subdivision would seem to be made above them, where 
Hz. virgula ceases to be a common fossil; and these two parts, 
N,, N, of M. Pellat, 23-25 of M. Hébert, may form the zone of 
Ammonites gigas, which certainly occurs throughout. 

The next zone is the chief fossiliferous one. It consists, at the 
base, of soft marly sands with hard bands, N, of M. Pellat, charac- 
terized by Perna Bouchardi ; next, of some hard grits in more than 
one bed (N*), with Cyprina Brong gnarte, Plerocera oceant?, Natica 
Marcousana, and Hemicidaris purbeckensis; and, finally, of earthy 
beds and sands more or less ferruginous and consolidated, with 
rolled pebbles and local unconformities. The whole of this does 
not occupy more than 20 feet at La Créche. At Portel the middle 
portion is nodular, and the whole is thicker. At the quarry of 
Mt. Lambert, nearest Boulogne, the whole appears as massive sand- 
stones; but at Gris Nez the most instructive section occurs. On the 
east of that point the marly beds are crowded with Perna Bouchardt 
and many other fossils; over these come the hard grits with Cyprina 
Brongmartr, forming a well-marked feature; and above is a local 
band of green earth, followed by a conglomeratic sandstone and 
sand, which in the cliff on the south appear to thicken out to nearly 
30 feet. The whole in the first locality is overlain by the shales of 
the “Middle Portland,” not previously noticed here. The nodular 
character of these upper sandstones is noted by M. Hébert in his 
No. 17' of La Créche. With regard to the fossils, Trigonia Pellate 
extends to the top at Gris Nez, and so does Ammonites gigas at La 
Créche, according to M. Hébert; the others noted are Nerita transversa, 
Orthostoma Buvigniert, Corbula autissiodorensis, Corbicella Bayan, 
Mytilus Morrisu, Pecten suprajurensis, Corbula ferrugmea, from 
the Perna-beds; and from the grits and sands, Twrritella Semanna, 
Acteonina Davidson, Paleomya autissiodorensis?, and Ostrea rugosa 
in addition. 

The succeeding series, constituting the “ Middle Portlandian ” of 
M. Pellat, is very well marked. It commences at the base with 
some 30 feet of black, almost paper, shales, very similar to those of 
Hen-cliff and of the Upper Kimmeridge of Lincolnshire. These are 
followed by several cement-stone bands, two being especially con- 
spicuous; and then more shales, with a lumachelle of Ostrea dubi- 
ensis at the top, making a total of about 50 feet. The fossils are, as 
usual, flattened in the shales, and in places are innumerable. The 
most important species noticed were <Alaria cingulata, Astarte 
scalaria, Cardium morimicum, Lucina minuscula, Mytilus autissrodo- 
rensis, Anomia suprajurensis, Discina latissima, and Avicula octavia 


566 J. F. BLAKE ON THE UPPER 


in the lumachelle. These beds form the O, of M. Pellat, and 
the middle of Nos. 15-17 of M. Hébert. They are seen at the top 
of the cliff east of Gris Nez with the usual Astarte. Above these 
the beds become softer and more glauconitic by degrees, the lowest 
being a very fossiliferous band with Lima boloniensis, over which is 
about 48 feet of clays, the upper half of which is more sandy and 
full of hard bands, and forms a better cliff, the limit upwards being 
drawn where the sandy clays cease to be glauconitic and to have a 
blue tint. Lithologically these beds have a very close resemblance 
to the upper portion of the Kimmeridge Clay of Chapman’s Pool, 
which likewise becomes more glauconitic, and passes gradually into 
Portland sands, and thence to the Flinty series. The fossils here 
noted were, in addition to the Lima mentioned, Ammonites pseu- 
dogigas, Belemnites Sourchii, Plewrotomaria Rozett, Myacites jurassi, 
Astarte Semanni, Trigonia concentrica?, Mytilus unguiculatus, Pecten 
lamellosus?, P. Morini, Perna Bouchardi, Avicula octavia, Plicatula 
Boisdimi, Ostrea expansa, O. bruntrutana, Acrosalena Kengi, Cr- 
daris bolomensis, and Serpula triserrata. 

The highest beds of the series (P,, P,, P, of M. Pellat) are only 
seen in the cliffs on either side of Wimereux to the north, and of 
Portel to the south. The base consists of about 9 feet of sand and 
sandstone with fucoidal markings, like the flinty beds of St. Albans; 
and these contain abundance of large Cardiwm Pellati, and some 
Trigonia gibbosa and other shells closely imbedded. Above is a 
shell-bed crowded with Serpula gordialis, just as happens at Port- 
land and St. Albans. Natica ceres and many small Cerithia are 
here abundant. The succeeding rocks are more calcareous repeti- 
tions of the same kind, all the hard parts being fair limestones, and 
the softer quite sandy, making a total of about 15 feet. The lime- 
stones are exceedingly fossiliferous, having yielded M. Pellat nearly 
70 species; but the whole remains of the same character as the 
Flinty Series of St. Albans, though in the latter case the sands have 
been consolidated into flints. The other most noticeable fossils 
appear to be Ammonites bononiensis, Natica elegans, de Lor., Ceri- 
thium Manseli (near the top), C. pseudo-excavatum?, C. septempli- 
catum, Pleurotomaria Rozeti, Trigonia incurva, T. Carre, Astarte 
rugosa, Corbicella Pellati (?=Sowerbya Duker [cast]), Cardiwm 
dissimile, Pecten lamellosus, and Echinobrissus Brodier. The over- 
lying strata are of very peculiar character, bemg mostly composed 
of calcareous rubble and tuff, in which there are scarcely any recog- 
nizable fossils; and such as are recorded (Astarte socialis, Cypris, 
Cyrena Tombecki, &c.) are not known in English Portland rocks. 
The irregular manner in which these lie, aah ferruginous beds 
intercalated in places, according to M. Pellat, forbid us to look upon 
them as certainly equivalent to the Portland building-stones, which, 
as a lithological group, or as a paleontological horizon containing 
Ammonites giganteus (true), Bucconum naticoides, B. angulatwm, and 
Cerithium portlandicum, are not here to be found. There is, in fact, 
no proof that any thing but Wealden beds overlie the Flinty Series at 
Boulogne. 


JURASSIC OF THE PARIS BASIN. 567 


Discussion or Resutrts. 


From the above observations it is proposed to classify the rocks 
under consideration as follows :— 


1. PortTLANDIAN. 


Upper = Purbeck. 
Lower= Portland Limestone. 


2. Bowontan. 


Upper =“ Middle Portland.” 
Lower =“ Lower” Portland. 


3. KIMMERIDGIAN. 
Virgulian. 
Pterocerian. 
Astartian. 


4. CoRALLIAN. 
Supracoralline. 


Coral Rag. 
Coralline Oolite. 


. OXFORDIAN. 


(2) 


_ f Oxford Oolite. 
Upper = | Oxford Grit. 
Lower = Oxford Clay. 


The first point to be considered is the upper limit of the 
Oxfordian group. When the English classification is compared 
with the French, serious discrepancies are found to exist; for 
our Lower Calcareous Grit, and part at least of our Coralline 
Oolite, are in France universally placed in the Oxfordian. In the 
Ardennes department it has been seen that grits which underlie the 
Neuvizy ironstone correspond both in character and contents with 
the Nothe Grits and the Calcareous Grits of Yorkshire up to the 
Passage-beds ; yet these are placed as Middle Oxfordian by Sauvage 
and Buvignier(2). The richly fossiliferous ironstone forms the type 
of the Upper Oxfordian for all French geologists; and when we trace 
its range and that of the rocks which overlie it, this seems justified 
stratigraphically. For above it lies a series of very changeable lime- 
stones, contrasting in this respect with the uniformity of the iron- 
stone ; and these same limestones, after the ironstone has died out, 
are carried on in a recognizable form over other Oxfordian beds. 
Thus the independence of the two groups is most marked where 
the dividing line is drawn. The Paleontological separation is also 
well marked ; for out of the 154 species recordedby Buvignier from 
the ironstone in the Meuse, only 45 pass upwards to any member 
of the Coral Rag. Excluding the Creué limestone, which forms a 
kind of passage-rock, the difference is equally marked wherever the 
fossils have been sufficiently studied, as in the Yonne, where Cotteau 
shows (16) that, out of 323 species in the lowest Corallian beds, only 
17 occur in the Oxfordian, if we draw the line between the two 
groups as proposed; and even where the lithology is the same, 


568 J. F. BLAKE ON THE UPPER 


only 17 out of 58 are common. If, then, by stratigraphy, by 
paleontology, and by common consent, the Neuyizy ironstone 
and its equivalents are to be placed as the summit of the 
Oxfordian, the English classification must be modified to bring it 
into harmony with more widely-established facts. It has been seen, 
in former studies of our English Corallian rocks, how readily they 
lend themselves to such a change by the ‘“‘markedly Oxfordian 
character of the fauna” of their lower part. It is proposed, there- 
fore, to abolish the term ‘‘ Lower Calcareous Grit,” as associating 
the rock intended too closely to the “ Upper Calcareous Grit,” and, 
as it forms an integral part of the Oxfordian series, to call it the 
‘‘ Oxford Grit,” and, for some part at least of the overlying lime- 
stones, to revert to the old name of the “‘ Oxford Oolite.” These 
terms, however, are too descriptive to be generally applicable, and, 
when they are not so, may be replaced by the more general term 
‘¢* Upper Oxfordian.”’ 

We may now trace the range of this mass over the area 
studied. ‘The Ferruginous Oolite of Neuvizy has been placed as 
the Lower Oxfordian by Oppel (27), partly in reliance on Prof. 
Hébert’s erroneous section (21), which places a mass of clay 
above it, partly by placing the Crdaris-florigemma beds as Upper 
Oxfordian, but chiefly by including the whole of the underlying clay 
in the same subdivision, and thus quoting true Oxford-Clay fossils, 
which are not to be found in Buvignier’s lists, such as Amm. biarmatus 
and Amm. Henrici. Oppel does not, however, quote Amm. Lamberti, 
which is most characteristic of Oxford Clay. There is not sufficient 
evidence, therefore, to overcome the weight of that obtained from 
the list of fossils given above, which places this ironstone as the 
uppermost part of the Oxfordian, as it is stratigraphically by its 
immediately underlying the Cidaris-florigemma beds. Along the 
whole of the Eastern range, the Oxford Grit below remains constant, 
becoming more nodular to the south; and the ironstone reaches 
nearly as far. The only ‘deposit hereabouts which can give rise to 
any dispute is the limestone: of: Creué and Liouville, the highly 
Oxfordian character of the fossils in which has led Hébert (21) to 
place it in the Oxfordian ; while Buvignier (22) makes it Corallian. 
Paleontologically, itis undoubtedly more united to the Oolite below ; 
but by its extremely local character, and its alternating with the 
Rag above, it belongs more to the latter, and must therefore be left 
as on a kind of neutral ground. 

On the change of strike and on crossing the band of Lower Jurassic 
rocks which extends to the south, we find a change in the development. 
Grits and ironstones can no longer be found, but the rocks continue 
to be marly as below; while a new set of Ammonites characterizes 
the upper part, though the general facies remains the same, the true 
Oxford Clay, with Lon. Babeanus and Amm. Lamberti and others 
belonging to the group Harpoceras remaining constant. Below, it 
would appear that the beds which ‘intervene between these and the 
Corallian, with their Ammonites continuing from below, as Amm. 
marantianus &c., occupy the place of the Oxford Oolite and Oxford Grit, 


JURASSIC OF THE PARIS BASIN. 569 


and are therefore the Upper Oxfordian. Tombeck (50, 55,57, 64, 65, 
66) endeavours to show that the zone of the last-named Ammonite 
is above the Coral Rag ; but, his stratigraphy not being accepted, no 
proof or probability exists of its occupying so anomalous a position. 
Three are certainly great differences of opinion on the true position 
of the zone of Amm. tenwilobatus, which in the Jura is supposed to re- 
present this—some, as De Loriol, Mésch, Bayan, &c., considering it 
Astartian, and others, as Hébert, Dieulafait, &c., making it Ox- 
fordian. It would be hazardous to venture an opinion on this point 
before studying the localities; but the fossils have an Oxfordian 
facies; and if the zone really corresponds to that of dmm. marantianus 
in the Haute-Marne, it is certainly not Astartian. These are the so- 
called Marls without fossils, both Upper and Lower, which have never 
been seen separated by any true Diceras-beds. They form the base 
at Les Laviéres, Vouécourt, and Buxiéres ; and the lower part of the 
same mass constitutes the marly limestones of Maranville, and the 
upper the cement-stones of Clairvaux, in the Aube. Further west, 
in the valley of Laignes, the gritty character comes in again, with 
similar fossils to those in the Meuse. In the Yonne we find the 
true Oxfordian marls with their Harpocerata at Ancy-le-Franc, and 
above these the Upper Oxfordian, in the new form of massive cal- 
careous sandstone, at Pacy, in the valley of the Armancon. The iron- 
stone of Ktivey has unfortunately not been examined ; but by its 
fossils it might well represent the Oolite of Neuvizy, except for 
Ammonites lunula. In the neighbouring valley of the Yonne the 
Upper Oxfordian is recognized in the siliceous limestones underlying 
the great white limestones of Coulanges, referred to the Corallian. 
Great differences of opinion have been expressed about the correlation 
of this portion. In their general work Leymerie and Raulin (25) 
placed the Ktivey ironstone as Lower Oxfordian, as also the beds at 
Ancy-le-Franc, which they nevertheless considered lower still. The 
Pacy freestones they called Middle Oxfordian, and paralleled with 
the Coral Rag of Chatel Censoir, leaving the lithographic limestones 
only to represent the Upper Oxfordian. © This, however, has been 
proved to be wrong by Cotteau (16, 34), who showed the essen- 
tially Corallian character of the limestones of Coulanges and of 
Vermanton, and discovered the Coral Rag at the base of the litho- 
graphic limestones. The fossils and the stratigraphy are all in favour 
of the latter author’s view. M. Hébert (21, 31) recognizes in 
the Etivey ironstone an equivalent of the Neuvizy, and says it 
plunges beneath the limestones of Ancy-le-Franc. The latter thus 
becomes his Upper Oxfordian ; and in the same horizon with it he 
places the freestones of Pacy, which are the equivalent of the upper 
part only, and the limestones of Vermanton, which have been proved 
Corallian. He considers it also the equivalent of the Creué limestone, 
which is above the Ferruginous Oolite. Some of these correlations 
may be rejected : but there is unanimity on the Pacy freestones repre- 
senting the Upper Oxfordian, in spite of the recorded presence of 
A. Babeanus. In the valley of the Loire the Upper Oxfordian is lost 
in the unfossiliferous marls; and in that of the Cher it must lie in 


570 J. F. BLAKE ON THE UPPER 


the lithographic limestone mass, since the fossiliferous sponge-bed 
is characterized by Ammonites canaliculatus and other species be- 
longing to a lower horizon, though the recorded presence (60) of 
A. marantianus shows that we are there not far off the top. In the 
Lower Charente, the locality Marans has beds below any thing seen 
at La Rochelle. In the Sarthe the description of M. Hébert (21) 
would lead us to place his ‘‘ Middle Oxfordian” as Oxford Grit, which 
here returns to its usual character, and his ‘“‘ Upper Oxfordian,” 
which is here marly, as Oxford Oolite. On the coast of Calvados 
we find in the Trouville Oolites, seen in true succession in the cliff 
of Auberville, a good paleontological representative of the Neuvizy 
ironstone; and below comes the Oxford Grit, with an equally cha- 
racteristic fauna. The lower of these two beds is considered by 
Hébert (28) to be Middle Oxfordian, and therefore to correspond to 
the Neuvizy ironstone, and the rest to be Upper Oxfordian ; but the 
Neuvizy ironstone is certainly more allied paleontologically to the 
upper beds, and the grits associate themselves with these more 
completely than with the beds below. 

The series here, as has been seen, corresponds exactly with that at 
Weymouth—the Nothe Grits,the NotheClays, the Bencliff Grits, and the 
Osmington Oolites all following regularly. The last named, therefore, 
are to be specially paralleled with the Neuvizy ironstone, with which 
they have 29 per cent. of their fossils in common, and must be called 
Oxford Oolite,and the grits below, Oxford Grit. Whether the Trigonia- 
beds of Weymouth are to be also included is a more difficult matter, 
to be discussed hereafter ; but we must certainly place on the same 
horizon the pisolites of North Dorset and the oolite of Highworth, 
which has 57 per cent. of its fossils common to Neuvizy, and perhaps 
also the road-stones of Faringdon. Unfortunately at Oxford the ~ 
“Oxford” Oolite is not well represented, except near Marcham. At 
Upware the limestones of the north pit belong here. The great 
development in Yorkshire gives us for the Oxford Grit the whole 
mass up to and including the Passage-beds, and for the Oxford 
Oolite as far up as the Trigonia-beds of Pickering, the latter having 
55 per cent. in common with Neuvizy. In fact, the greater part of 
the limestones which underlie the Coral Rag proper belong to this 
portion. 

Our circle is finally completed in the Boulonnais, where the 
limestone of Houllefort, in the midst of clays, offers us the only 
representative of this part of the series, though to what exact part 
it may be considered to correspond may be doubtful. The excep- 
tional presence in it, however, of Cidaris florigemma indicates a high 
position. : 

We have next to discuss the Corallian series. De Loriol (50, 
56) and Tombeck would have us abolish the term altogether ; 
and Oppel (27) practically does so by including his “ zone of Cidaris 
florigemma” in the Oxfordian. The argument against its use is that 
coral-growth is only an accident which may happen at any time, 
and that there is much of the so-called Corallian which has no corals 
in it. These statements are doubtless true, and yet, to one who has 


JURASSIC OF THE PARIS BASIN. 571 


traced the series from end to end of the Paris basin, they seem to 
have no weight. The Corallian series, as now to be limited, is one 
of the best-marked groups it is possible to conceive; but, as Buyig- 
nier (22) has so well expressed it, its constant feature is inconstancy. 
Below this group we may have some continuous beds; above it the 
beds are markedly continuous ; but within it discontinuity is the rule. 
It is quite to miss the mark, therefore, to say, as Tombeck says 
(50 &c.), that its normal form is the compact limestone, and coral- 
growths and Diceras-beds abnormal. Noone kind of deposit is more 
normal than another, except in this way, that the variety which is 
most frequently repeated, and which occurs at several horizons and 
under the most varied forms, is a coral-growth, and neither above 
nor below the limits of this series can such coral-growths be found 
in the country examined. Hence, unless one were to invent a new 
term, such as Protean, to indicate variability, no better name could 
be chosen than Corallian to indicate this most name-needing group. 

In the study of the English Corallian we have stated* that the 
Coral Rag, with Cidaris florigemma, is always at the top of the 
limestones, while the French geologists have asserted it to be as 
constantly at the bottom. This anomaly is now explained. By 
marking off so much of the so-called Coralline Oolite to place it in 
the Oxfordian, we leave the Coral Rag nearly, and in most places 
quite, at the base. Sometimes, as near Oxford, it lies on the 
Oxford Grit ; sometimes, as at Upware, on the Oxford Oolite; and 
only here and there can we suppose, and that doubtfully, that any 
oolite belonging to this series rather than to the Oxfordian, and 
hence to be called Coralline Oolite, intervenes. It may beso at 
Malton and Seamer, in Yorkshire, especially where shell-beds are 
developed; and it may be so at Weymouth, where we may possibly 
call the Trigonia-beds by this name. As for Cidaris florigemma, it 
is characteristic of the Corallian as a whole, and is perhaps not con- 
fined to it. Tombeck describes (55) a bed abounding with it at 
St. Ansiau, in the Haute-Marne, supposed to lie in the Oxford Clay, 
though a fault is here possible ; and the Houllefort limestone, which 
cannot be called Corallian, contains it ; on the other hand, it is found 
in Astartian beds at Weymouth. But, after experience of its habit, 
it is impossible to say more than that it is commoner towards the 
base than towards the top of the Corallian series, and is usually asso- 
ciated with massive corals. As to there being a bed full of it con- 
stantly marking the junction with the Oxfordian, as Hébert says 
(21), this is a feature that seldom occurs. 

The upper line separating the Corallian from the Kimme- 
ridgian is rather hard to draw. Tombeck says (50) that, if 
we take the so-called Astartian as the base of the latter series, 
it cannot be drawn paleontologically; and de Loriol (50, 56), 
on the strength of this, wishes to call the Astartian and Coral- 
lian by one name, Sequanian. On the other hand, Hebert (19, 
31) says that one can put one’s finger on the line of junction, and 


* Blake & Hudleston, Quart. Journ. Geol. Soc. vol. xxx. p. 313 &e. 


572 J. F. BLAKE ON THE UPPER 


that the two series are perfectly separate by their fossils. The 
latter supposition we may dismiss, as it is exceedingly improbable ; 
and it has been shown(p.529)that in the Aube several such apparent 
junction-lines occur, one above the other. On the other hand, Tom- 
beck’s statements appear to be vitiated by his including the Oolite of 
La Mothe in the Astartian, instead of associating it with the Diceras- 
beds ; and if this rectification were made, we should find one palzon- 
tological distinction indicated by himself—namely, that while the 
Corallian contains the zone of Cardium corallinum, the Astartian con- 
tains that of Verebratula humeralis. Admitting, however, that the 
actual line may be rather arbitrary, and that some Corallian forms 
occur above it, and even Oxfordian ones, 1f Ammonites marantianus 
be truly recorded, yet if it be drawn not far above the Diceras-beds 
and ther equivalents a change is soon perceived after the line is 
‘passed, as will be shown in the sequel We shall then have three 
subdivisions in the Corallian series, which may or may not be deve- 
loped in any particular locality,and the distinctness of which may often 
be lost, but which are nevertheless sufficiently constant to make it 
important to recognize them. ‘The upper division, or “ Supracoral- 
line,” almost wants a better name than it has received. Its most 
remarkable form, the Diceras-beds, would almost induce us to apply 
that name, but that the form is too local. The term *‘ Supracoralline” 
1s applicable in the great majority of cuses ; but the beds must not be 
supposed to be always above the actual coral-growths. The middle 
division, or ‘Coral Rag,” is so called to denote the more common | 
position of the Astraan corals, generally associated with Crdaris 
florigemma, though this portion may be without such corals, which 
may occur above it instead. And the lower division, or ‘ Coralline 
Oolite,” represents those limestones which lie in certain places above 
the best representative of the Upper Oxfordian in the district, and 
yet below the beds identified as Coral Rag: it is a division which in 
most districts may be ignored. 

We must now trace these beds in their range. In the Ardennes 
we find, immediately above the Ferruginous Oolite, a comparatively 
thin bed of crystalline coralliferous limestone, with Cidaris flori- 
gemma, which may be recognized at once as the Coral Rag. Itis sur- 
mounted by the magnificent limestone of Novion, in which Crdaris 
florigemma is rare, if present, but which contains Cardiwm corallinum 
and some Dicerata. These, though full of corals, are the Supracoral- 
line beds, becoming oolitic above, and affording no well-marked upper 
limit. On entering the Meuse we immediately find the Coral Rag to 
have increased, and to have developed various forms, such as oolites 
and crinoidal limestones. There is, however, no regularity in these 
minor beds, since near Verdun the crinoidal limestones are beneath 
coral-growth, whereas near St. Mihiel they cap it. Near Verdun 
some shell-beds are found which might be called Coralline Oolite. 
The Supracoralline beds are much thicker. In fact, if we are to keep 
the Astartian beds at all comparable to each other, and to include 
all rocks of the Diceras-bed type in the Corallian, we must absorb 
into the latter a considerable portion of Buvignier’s Astartian. At 


JURASSIC OF THE PARIS BASIN. 573 


St. Mihiel we have a thin bed below the coral-growth, which is best 
ealled Coralline Oolite ; and it is probably the end of the great lime- 
stones of Creué, which also will be placed as such. Buvignier has 
himself (22) recognized that the Coral Rag of St. Mihiel is no other 
than that of Verdun, and it is only one facies of the beds of this age. 
The Supracoralline division retains its former character, but begins to 
have true Diceras-beds on the top, especially in the south (21). In 
the extreme south the Coral Rag develops into avery thick mass, which 
is very rubbly and fossiliferous towards the base, with Cidaris flori- 
gemma &e., surmounted by white chalky limestone, and finally by 
compact limestone beneath the Astartian beds; so that here the 
distinctions of the two parts of the Corallian are less marked. In 
the Haute-Marne, at Reynel, in the east, the rubbly Coral Rag occu- 
pies the base, and more compact limestone lies above it (55), with a 
development of the Diceras-beds above. Inthe valley of the Rognon 
there is a band of disaggregated Oolite at the base (19), then a fine 
development of the Rag, and a magnificent display of the Supracoral- 
line beds in the form of pisolites with Diceras, with compact and 
oolitic beds associated, up to the more marly Astartian beds. Inthe 
valley of the Marne the Supracoralline Diceras-beds have become the 
summit ; and beneath come the massive coral-growths with Cidaris 
florigemma, occupying a great thickness, and having below them 
massive limestones, which might be called Coralline Oolite ; but as, 
according to Royer(12), they have rubbly rag below them at Soncourt, 
they must be placed as a development of the Coral Rag. Passing 
west towards the Aube, the Diceras-beds become more restricted, and 
confined to the upper part, as at La Mothe, while the coral-growth 
or rubbly beds beneath lose their character, or remain as shelly lime- 
stones only. On reaching the valley of the Aube itself, all is united 
in an undistinguishable mass of limestone, with only a shelly repre- 
sentative of the Supracoralline beds; and Astartian and Corallian 
form one massive, lying on Oxfordian marls at Clairvaux, and 
covered by Pterocerian marls near Bar-sur-Aube. ‘Travelling west 
to the valley of the Laignes, new developments arise. The thin shelly 
band is replaced once more by Diceras-beds of considerable thickness, 
and the beds below put on a characteristic form at the base; for 
below a mass of compact and false-bedded limestone comes a rubbly 
mass of coral-growth, full of the shells which usually accompany it, 
though Cidaris florigemma was not noticed. This again might be 
called Coralline Oolite, but that it is believed to develop into Coral 
Rag later on. In the valley of the Armancon this lower shelly mass 
has again dwindled down to a narrow band; so that it was left to 
Cotteau, or rather to M. Lettéron (438), to discover it, filled with corals 
and the usual shells and echinoderms, underlying the lithographic — 
limestones. This discovery seems to give the coup de grace to the 
idea of the latter being Upper Oxfordian (25). They are, in fact, 
the continuation of the compact limestones of the Laignes valley ; and 
in various beds contain a Corallian fauna, the uppermost of which, 
met with by M."Hébert near Angy, seems to have led him (31)to place 
the limit of the Oxfordian too high, 7. ¢. above instead of below the 


574 J. F. BLAKE ON THE UPPER 


lithographic limestones, which are very different from the limestones 
of Pacy. In fact, the limestones of Angy and Tonnerre are shown 
to be the Supracoralline beds, and do themselves contain Diceras-beds, 
especially at the top, though the whole is enormously developed, so as 
easily to mislead one. On passing westwards, however, to the valley 
of the Yonne, still more is in store ; for while in the neighbourhood 
of Bailly and Vincelles the Supracoralline beds occur in all their 
characters, beneath them are still found the lithographic limestones, 
more than usually marly at Vermanton; and below these again the 
coral-beds of the Laignes valley, so much reduced in the Armancon, 
burst out into a magnificent mass, wherein corals of all kinds, 
Diceras, Nerinewa, and all the Rag fauna are confusedly mixed in ex- 
treme abundance, as may be seen so well at Merry-sur-Yonne, 
Chatel-Ceusoir, and, in a more uniform character, in the white lime- 
stones of Coulanges. It is this development that for ever negatives 
the idea that Diceras-beds are necessarily above the Coral Rag, though 
these Dicerata differ from those of the Diceras-beds in apparently 
having lived near the spot, and in not having been rolled. Beyond this 
valley the Corallian once more changes, and by degrees returns to the 
dead uniformity of a lithographic limestone. In the valley of the 
Loire the lower Diceras-bed is seen in diminished thickness near La 
Charité, succeeded in an upward direction by compact limestones, 
which show no more character than that of becoming oolitic near 
Pouilly. On the left bank of the river, however, it is stated (32, 60) 
that the Lower or Coral-Rag Diceras-beds are not to be found, but that 
the Supracoralline ones are once more developed at Sancerre. These 
last take up again their Armancon form near Bourges, and show, not 
quite at the summit, the white limestones of Angy and Tonnerre. 
Hence to the Cher is the path of degradation ; for in the latter valley 
even less can be distinguished than in that of the Aube, but all is 
lithographic. In the Charente department the Corallian rocks remain 
still massive and but slightly characterized, though coral-beds are 
developed in what may be considered the equivalent of the Coral Rag. 
The Supracoralline beds are still characterized by the abundance of 
Nerinee and by the presence of Diceras and Cardium corallinum. 
In the Lower Charente, on the shores of the Pertuis d’Antioch, the 
Corallian rocks put on a fresh form, the lower part being a white fos- 
siliferous limestone, which, by its fossils, may represent the Rag ; 
while the true coral-bearing beds are on a horizon usually “ supra- 
coralline,” and are parallel with the similar development in the 
Ardennes. It would appear that d’Orbigny included still higher beds 
in his ‘ Etage Corallien’ here ; but a comparison of the series in the 
two Charentes and the general fauna indicate the necessity of draw- 
ing the line separating the Corallian from the Astartian almost 
immediately above the coral-reef of the Point du Che. In the 
departments of the Orne and Sarthe, according to Hébert’s descrip- 
tion, there would appear to be no development of corals on the usual 
horizon ; but the Supracoralline beds contain Diceras and are over- 
lain by coral-growth, which is thus on a higher horizon than almost 
anywhere else. 


JURASSIC OF THE PARIS BASIN. 575 


On the coast of Calvados the sequence is tolerably plain; for 
the beds characterized by Cidaris florigemma succeed immedi- 
ately to the Trouville Oolite, and thus occupy the base. They 
change laterally into coral-growth, which represents the Coral 
Rag; so that there is little, if any thing, that can be called * Coral- 
line Oolite,” unless it be the mass that underlies the Rag in the 
Trouville quarry. The Supracoralline beds are here quite different 
in form from those in the southern range, being arenaceous, instead 
of calcareous or coralliferous ; nevertheless they contain calcareous 
matter, and are admirably marked off stratigraphically from the 
beds above by the difference of lithological character. The arena- 
ceous character of these beds appears to prevent their assimilation 
palezontologically with their calcareous representatives, the two sets 
having little in common; nevertheless the fossils afford no nega- 
tive to their stratigraphical correlation. In the well-known neigh- 
bourhood of Glos, in like manner, can be traced the Coral Rag in 
all its commonest characters, with perhaps a representative of a 
Coralline Oolite below. Above, we appear to have a twofold de- 
velopment of Supracoralline beds :—one, at the base, being calcareous 
and similar to the Novion limestone and other southern supra- 
coralline beds, wanting only the Diceras, but with abundance of 
ramose corals; the other, above, being an enormous development of 
loose sand, corresponding to similar beds on the coast, and containing 
a fauna which indicates a high position. These sands were placed 
by Zittel and Gouert (under the guidance of Hébert) as Upper 
Corallian, which corresponds to the above-assigned place. Their 
relation to the coast-beds, and the correspondence of the latter to the 
Sandsfoot-Castle beds of Weymouth, leave no doubt that the latter 
must, with them, be placed as Supracoralline. We come thus to the 
discussion of the Corallian of Dorset. There seems little doubt that, 
the Sandsfoot-Castle beds being Supracoralline, and the Osmington 
Oolite Upper Oxfordian, we must find the beds corresponding to the 
Coral Rag between these limits. What actually does intervene is 
the Sandsfoot Clay and the Trigonia-beds. The abundant fossils of 
the latter still leave some doubt as to its proper position. On the 
one hand, such fossils as Gervillia aviculoides, Ammonites cordatus, 
Pecten fibrosus, and Echinobrissus scutatus are extremely Oxfordian ; 
while the presence of Ostrea deltoidea, Cidaris florigemma, and 
Thamnastrea arachnoides attaches the rock to the Corallian. The 
knot can only be cut by calling it a passage-bed, or by placing it 
with other similar puzzles as ‘‘ Coralline Oolite.” It cannot, how- 
ever, properly represent the true Coral Rag ; and thus we are driven 
to look upon the Sandsfoot Clay as being its equivalent here, a cor- 
relation not without value when other northern localities are con- 
sidered. In North Dorset the lower limit of the Corallian, as now 
defined, is very well marked; for the uppermost Oxfordian is the 
false-bedded series, and the overlying rubbly beds with Cidaris flori- 
gemma cut off the upper edges. Nevertheless the paleontology is not 
altogether satisfactory ; and we are obliged to allow a wider range 
than usual for more than one characteristic fossil, ¢. g. Echinobrissus 


576 J. F. BLAKE ON THE UPPER 


scutatus and Hemicidaris intermedia. At Westbury the Corallian 
would appear to be almost absent ; for we must now refer the rubbly 
beds with Hehinobrissus scutatus to the Upper Oxfordian, and the iron- 
ore itself is nearly, if not quite, Astartian. The ferruginous cha- 
racter of rocks appears in some way connected with such gaps. In 
the neighbourhood of Westbrook, Wiltshire, a coral-bed has been 
described* from which Cidars florigemma is absent, and which, 
on that account, is considered older than the usual Rag. This may 
be so; but, in view of the similar Rag in the Aube department 
passing into a true florigemma-lKag in the Yonne, the difference can- 
not be of much importance, either here or elsewhere. It certainly 
need not be considered an Oxfordian reef. Atthe Calne quarries the 
lower clays, like those of Hillmarton, abound in Crdaris florigemma, 
and the bed with Hemicidaris intermedia occurs towards the base ; 
hence an examination of these quarries alone might lead a foreign 
geologist to call the freestones Supracoralline ; but the error of such 
a correlation is easily proved by the overlying coral-growths observed 
in the heights on the north and south immediately beneath the Kim- 
meridge Clay. There is little, in fact, in this district to represent any 
Supracoralline beds. The same is true all the way to Oxford, 
the ferruginous earth being the only deposit referable to them. 
The only Corallian rock is the Coral Rag itself, unless we call the 
great quarries at Wheatley Supracoralline, for which their great 
resemblance to the limestones of the Ardennes and the Meuse 
might be an argument. The arrangement of the Yorkshire beds is 
not very easy, because it is rather difficult to decide which of the 
beds underlying the Rag should be placed with the Oxford oolite, 
and which should be associated more closely with the overlying 
mass. Referring to our comparative sections (Q.J.G.8. vol. xxxiii. 
pl. 12), the abundance of Hchinobrissus scutatus in the Lower Oolites 
of Grimston mark them as Oxfordian; while the ‘“ mamillated-ur- 
chin series,” by its Cidaris florigemma, is as plainly Corallian. 
The Chemnitzia-lmestone of Malton is just one of those doubtful 
deposits to which it is best to apply the name Coralline Oolite. At 
Pickering the rubbly nature of the base of the Upper Limestones, 
g, and the introduction there of corais, added to the change of 
fauna, indicate that the base-line of the Corallian should be drawn 
almost directly above the Trigonia-beds. It will then be seen that 
the shell-beds at the top of the Lower Limestones are more con- 
nected with the uppermost Oxfordian than with the Corallian. As 
to the Supracoralline beds, they are simply equivalent to the ‘‘ Upper 
Calcareous grit,” which latter is parallel to the Sandsfoot grit, and 
not altogether above it as we previously supposed (Jl. ¢. p. 390), 
owing to the non-recognition of the Sandsfoot clay as the equiva- 
lent of the Coral Rag. 

In the Boulonnais the clays with marly limestones of the Mont 
des Boucards were first called Oxfordian by Rigaux (83), appa- 
rently because they seem continuous with the marls below; but on 
the discovery of the coral-bearing beds at the base or lower part 


* Quart. Journ. Geol. Soe. vol. xxxiii, p. 288. 


ld 


JURASSIC OF THE PARIS BASIN. aT 


(41), Pellat was compelled to place them at least as Corallian, as he 
identified the Coral Rag with that of Brucdale, and, at the same time, 
noted that the overlying marls are on the horizon of the compact 
limestone of Vouécourt and of Tonnerre, both of which are Supra- 
coralline. At a later date (56), owing to the supposed proof by 
Sauvage (51) and Rigaux (53), that the equivalents of the marls of 
the Mont des Boucards underlay, in the soundings, the Brucdale Coral 
Rag, he met them halfway, and placed the two as equivalent. With 
the breakdown of that proof is lost the necessity of disturbing the 
former more correct reading, by which the coral-bearing beds of 
Brucdale and the Mont des Boucards are the true Coral Rag, and the 
marls of the latter place are Supracoralline, though not developed 
at Brucdale. The occurrence, however, of Trigonia Bronnw in a 
ferruginous bed above is too close a resemblance to the development 
at Glos and Sandsfoot to allow us to place them in the Astartian ; 
and we must raise the limit of the Supracoralline to, at least, the 
top of this bed. 

The next division is the Kimmeridgian. Its limitation above 
will be discussed later on; as to its lower limit, a point much dis- 
puted is, whether the beds called Astartian are to be included with 
it, or with the lower beds. It is to this portion of the series, it 
appears, that the term Sequanian was first applied by Marcou*; and 
it was considered part of the Kimmeridgian, while Thirriay had 
previously called it Astartian. Neither of the original localities for 
these names has, as yet, been examined; but the latter is adopted 
as most generally in use for a distinct portion of the series. De 
Loriol (61) is probably the chief advocate for uniting the Astartian 
with the Corallian under the term Sequanian, led thereto, perhaps, 
by the supposed sequence in the Haute-Marne, and by his objection 
to the term Corallian. Itis, however, noteworthy that almost every 
stratigraphist separates the Astartian from the Corallian, to associate 
it with the Virgulian or Pterocerian in one great group. It issoin 
the description of the Ardennes(2), of the Meuse (13), and in Royer’s 
original description of the Haute-Marne (4). It is not so in Ley- 
merie’s description of the Aube (7), probably because he held all the 
limestones at Clairvaux to be Astartian, nor in the same author’s 
description of the Yonne (25). But it is practically so in Cotteau’s 
description of the same (42), as the white limestone is made the 
uppermost bed of his “ Sequanian,” which thus really is Supra- 
coralline, and the true Astartian beds are left for the Kimmeridgian. 
It is soin the Nievre (32), in the Cher (60), and in the Charente 
(24); while in the Lower Charente, the coral-limestone being the 
highest Corallian bed, it may be taken that the Astartian is not 
specially recognized. On the coast of Normandy no one thinks of 
associating the Trigonia-beds with the Coral Rag, or Supracoralline, 
though they succeed the latter immediately ; and a study of the rocks 
in the Boulogne area leads Pellat to the same conclusion (41). 


* Recherches géologiques sur le Jura salinois, p. 116: 1846. 
+ “ Notice sur le terrain Jurassique,’” Mém. Soe. 8. N. Strasbourg, 1830, p. 26. 


Q.J.G.8. No. 148. 2a 


578 J. F. BLAKE ON THE UPPER 


There may be some difficulty in drawing a line; but throughout the 
whole of the eastern and southern range one finds a marly charac- 
ter of deposit setting in again over the true Supracoralline beds, 
often showing much rubble and indications of local unconformity, 
as we should expect on the introduction of a new series. ‘The 
same is more remarkably true in the North and in England; only 
in the Boulonnais is there somewhat more difficulty in drawing 
the line. The fossils are usually sufficiently distinct, Terebratula 
Leymervi being a characteristic form. The conclusion, therefore, so 
tenaciously held by Hébert (21), seems to be the true one, that these 
beds are a part of the Kimmeridgian. The thickness of the beds 
assigned to the Astartian by the describers of the different areas is 
very varying. ‘Thus, in the Ardennes they are said to be 21 feet, 
but in the Meuse 400 feet; on the Haute-Marne, again, they are 
supposed to be thin. It is plain, therefore, that a large portion of 
the 400 feet (if truly determined) must really belong to the Supra- 
coralline, as supposed for other reasons. ‘Their assigned thickness 
in the Aube is due to a mistake, as before noted; in the Yonne 
again they are called 33 feet ; in the Cher, down to the base of the 
Nerinxan Oolite is 35 feet, though 80 feet more are included by 
Douvillé. -The whole is therefore of comparatively small thick- 
ness, and forms only a subordinate portion. The Kimmeridgian is 
thus divisible into three—<Astartian, Pterocerian, and Virgulian. 
The first of these is practically defined above. The Pterocerian is 
adopted solely in deference to its probable justification im the area 
where it was first introduced, namely the Jura, and to its distinct- 
ness paleontologically when the fossils of any locality have been 
carefully studied. Neither in the basin of Paris, nor in any other 
part yet studied, is it sufficiently distinct to be of much importance 
in the field. The Virgulian is the most easily recognized Kimme- 
ridgian deposit, because the characteristic oyster occurs constantly in 
lumachelles ; but itis not confined to this part of the series, either in 
an upward or downward direction. 

In the Meuse department, and probably in the Ardennes, the 
Kimmeridgian is highly calcareous, the Virgulian portion alone 
being marly. In the former there still remains some doubt as to 
the true limits of the beds, and even as to their stratigraphy. While 
the lower portion of the Astartian requires to be added to the Coral- 
lian, the upper group appears to represent the Pterocerian as well, 
from the abundance, as recorded, of Pteroceras ocean, &ce. The 
lower portion abounds in the characteristic Ostrea deltowdea. As 
Tombeck points out (40), the Virgulian must be made to include the 
marly beds at the base of Buvignier’s “ Calcaire du Barrois,” since 
these contain Pholadomya acuticosta as well as abundance of 
Exogyra virgula. In the south of the Meuse department, if we 
place the compact limestones in the Corallian, we find the Astartian 
well characterized as rubbly limestones; and a comparison of this 
locality with the Haute-Marne in the valley of the Rognon, would 
lead us to commence the Astartian above the lithographic limestones 
which overlie the Saucourt oolite. Nevertheless it is to be noted that 


land 


JURASSIC OF THE PARIS BASIN. O79 


both Buvignier in the Meuse, and Royer originally in the Haute- 
Marne, included the lithographic limestones in the Astartian, 
and 'fombeck and Pellat do the same. It is therefore possible 
that the line of junction ought to be carried further down. In the 
south of the Meuse the Virgulian beds are well developed and very 
fossiliferous; but the lower portion has not yet been satisfactorily 
marked off as Pterocerian, although from the recorded occurrence of 
Pteroceras oceant at Mauvage, and the distinct grouping of the fossils 
on the several horizons, such a subdivision might doubtless be made. 
Tombeck has divided his Kimmeridgian into two zones, those of Amm. 
orthocera and of Amm. caletanus, corresponding to the Pterocerian 
and Virgulian respectively ; and these subdivisions are accepted by 
Pellat. In the Aube, contrary to the description of Leymerie, the 
Astartian commences above the chalky limestone, and is characte- 
rized throughout by rubbly and oolitic beds, with abundance of Tere- 
bratula Leymerw. Near Bar-sur-Aube something of a Pterocerian 
group may be made out; but the fossils have not been sufficiently 
studied to draw a satisfactory line, which must be a paleontclogical 
one. In the valley of the Armancon we have the same difficulty 
as in the Yonne to decide how far above the Diceras-beds the Astar- 
tian must commence, the rubbly beds next above containing rolled 
corals as well as Terebratula Leymerui; but perhaps the line is best 
drawn above the solid oolite block, as is done by Hébert (21) in 
the neighbouring valley of the Yonne. No subdivisions correspond- 
ing to the Pterocerian and Virgulian have been made out in these 
western districts. Pholadomya acuticosta is the most characteristic 
fossil. As to the neighbourhood of Bourges, the chief feature is the 
strong development of a Nerinean oolite. The beds below this still 
containing a species of Diceras, seem certainly more referable to the 
Supracoralline, in spite of their lying so far above the white lime- 
stones certainly recognized as the latter ; and it might be a question 
whether the Nerinzan oolite ought not also to be placed below 
the line of separation; but the presence of Pholadomya Protez, 
Terebratula Leymern, and Trigonia Baylit may be allowed to decide 
it in favour of the consensus of opinion. In the Charente, the three 
portions of the Kimmeridgian are recognizable, the Astartian com- 
mencing where the beds become more marly and contain Ceromya 
excentrica abundantly. This latter, however, is not a fossil which 
can exclude beds from being Supracoralline; and we may still re- 
gard the line, therefore, as rather arbitrary. The upper two zones 
are well distinguished paleontologically. ‘The same may be said of 
the Lower Charente: the Astartian may be commenced with the 
marly beds of the Point d’Angoulins, with abundance of Terebra- 
tula Leymerii, also Pholadomya Prote: and Ceromya eacentrica, the 
latter certainly becoming locally characteristic ; and the Pterocerian 
and Virgulian are well distinguished, according to Hébert (54), at the 
Point de Chatellailon. In Normandy, as at Weymouth, the Astar- 
tian beds are better characterized than elsewhere. The change 
from the calcareous grits of the Supracoralline is very marked ; and 
the beds included in the zone, viz. theVillerville beds and the Trigo- 
202 


580 J. F. BLAKE ON THE UPPER 


nia-grits of Havre, with any Ostrea-deltoidea beds above them, have 
well-marked features of their own. 

There cannot be any doubt of the correspondence of these beds to 
those called Astartian throughout the whole of the southern range ; 
and from this it follows that the Kimmeridge passage-beds of Wey- 
mouth are the exact representatives of the same portion. It seems 
also most satisfactory to place the Abbotsbury and Westbury iron- 
stones in the same horizon, as they are apparently higher than 
Supracoralline. The lowest beds of the Kimmeridge Clay seen in 
the Wootton-Bassett cutting, with Rhynchonella inconstans, and the 
great Ostrea-deltoidea beds of Lincolnshire, belong here also. 

On the south bank of the Seine, it is prebable that at Honfleur 
Pterocerian beds may be developed, though not well seen; but the 
ereat mass of the clay at the Cap du Héve belongs to this division— 
that is, the ‘* Marnes & Ptérocéres ” of Dollfus, and the beds nos. 7 
to 14 of Lennier. In spite of the abundance of Hxogyra virgula, 
and even of Terebratula Leymerw in certain beds, this portion is 
well marked paleontologically by its numerous Pterocerata. The 
Ammonite-marls above, so far as seen, belong to the Virgulian. 

Doubtless the same subdivision might, with care, be made at Wey- 
mouth ; but as yet, in spite of the many collectors who formerly 
searched the shore, no Pterocerian fauna has been brought to 
light. The Kimmeridge Clay of England has been shown to 
be divisible into two groups*, formerly called Upper and Lower 
Kimmeridge. The term Kimmeridgian must now be confined to 
the latter; and though the subdivision into Pterocerian and Vir- 
gulian is not very clear, it may, perhaps, fairly be taken that the 
absence of Hwogyra virgula (as in Lincolnshire), indicates the former, 
while its abundance (as at Ely, Swindon, and in parts of the south- 
ern coast section) indicates the latter. The idea expressed in the 
paper quoted, that the Lower Kimmeridge Clay represented the 
Astartian, was founded on the abundance of an Astarte which at 
least is very like A. supracorallina. It may, however, be distinct ; 
and in any case its presence cannot be allowed to interfere with 
stratigraphical conclusions supported by the general paleeontological 
facies. 

In the Boulogne area we have the same difficulty in drawing the 
lower limit as at Bourges. While, on the one hand, there need be 
no hesitation in including the Grés de Wirvigne in the Astartian, 
or the Mont-des-Boucards marls in the Supracoralline, the three 
intervening deposits are doubtfully attached either to one or the 
other. It is certain that, in the Meuse, Buvignier would include 
such beds as the Ostrea-deltoidea clays in the Astartian; yet the 
ferruginous bed above with Trigoma Bronnw seems to unite both, 
through the Normandy sections, to the Supracoralline, as has been 
seen above, p. 563. The important MNerinwa-oolite, again, is con- 
nected with the lower series by the abundance of that genus, nowhere 
characteristic of Astartian, but constantly found in Supracoralline 


* Quart. Journ. Geol. Soc. vol. xxxi. p. 197. 


JURASSIC OF THE PARIS BASIN. 581 


beds; and its complete fauna, as given by de Loriol, shows that 
17 species are common to lower beds and only 10 to higher. Much 
therefore might be said for making this Supracoralline ; but as there 
is not much certainty about it, and it has hitherto been placed in the 
Astartian, it may be left there, with the similar rock at Bourges, for 
the present. Perhaps it would be preferable to draw the line im- 
mediately above this, and include the F, F, of Pellat with the Gres 
de Wirvigne. The fossils of the latter approximate closely to the 
Astartian at Havre. The remainder of the Kimmeridgian is well 
characterized here ; the lower portion, being apparently barren of 
Pieroceras, would be better named after its Ammonite; but the 
upper part is markedly Virgulian, and corresponds to the mass of 
the ‘‘ Lower” Kimmeridge Clay (so-called) of Dorset. In the Pays- 
de-Bray only this portion is seen. 

For the series of deposits which overlie the true Kimmeridgian or 
Virgulian, and underlie the true Portland beds, the name of Botonran 
is proposed. It has already been proved* that in the Boulogne 
area these rocks correspond to what had been hitherto considered 
an integral portion of the Kimmeridge Clay; and Waagen (35), by 
separating them as a zone above the Virgulian, came to practically 
the same conclusion. For the lower portion of them, therefore, which 
especially differs in lithological character from Kimmeridge Clay, and 
is of the nature of an episode in its midst, the name of “ Bolognian 
episode ” was formerly proposed. A further study of the same series 
in the basin of Paris shows that elsewhere they are not specially epi- 
sodal in character, but nevertheless require separation from the Virgu- 
lian. The name Portlandian has usually been applied to them; but 
since it is certain that they do not correspond to our Portland rocks, 
but to beds below them, this name is to the last degree mislead- 
ingy; and the only way out of the confusion is the use of a distinet 
name. Seemann is said by Pellat (39) to have proposed the name 
Pontidian; but as this is rejected by the latter, who alone mentions 
it, it cannot be said to have priority ; and it is not a good geogra- 
phical name. It seems therefore best to modify the name already 
applied, and extend it to all the continental beds which, not being so, 
have been called Portlandian. Some name connected with the term 
“ Calcaire du Barrois” would have been better, if some true Port- 
land stone had not been included in that term. 

These Bolonian beds admit of a twofold subdivision in almost every 
locality, though the apparently natural limits in the several places may 
not quite coincide. In the Boulogne area the two parts here distin- 
guished have been called ‘“‘Lowerand Middle Portland.” According 
to Pellat, the fauna of the Lower Bolonian commences in the clays — 
below the great conglomerates; hence the change of fauna was not 
brought about by the changes of physical character, but had already 
commenced. The paleontological line will therefore not coincide with 
the lithological one; but the upper parts of the clay series are inde- 

* Quart. Journ. Geol. Soc. vol. xxxvi. p. 189. 


T Some Swiss geologists have been led by this misnomer to declare that there 
are no Portland beds (in ¢heir sense) in the Isle of Portland! 


582 J. F. BLAKE ON THE UPPER 


pendently marked off by a change of fauna, and the “ Lower Port- 
land” comes in in the midst as a true episode. This episode is less 
marked in the Pays de Bray, and scarcely recognizable in the coast 
of Kimmeridge, where we find the northern argillaceous type of 
the Bolonian, equally marked off from the Virgulian by its fossils. 
During the same epoch the southern or calcareous type was 
being developed. The distinctions made in this, of lithographic 
limestones, carious limestones, and tubular limestones, are too local 
to be of great importance ; but the two zones of Ammonites gigas and 
elther of Pinna suprajurensis or Cyprina Brongniarti are of wider 
interest. These represent the bulk of the Bolonian in the southern 
range. It would appear that at Boulogne the argillaceous and cal- 
careous types overlapped for a while, and the paleontological divisions 
scarcely coincide with the lithological. Hence if we divide the Bolo- 
nian into the two parts which have the widest significance, the Lower 
Bolonian, or zone of Amm. gigas, will commence with M, and end above 
N, of M. Pellat (that is below the Perna-beds) ; and the Upper Bolo- 
nian (or zone of Cyprina Brongmarti in the south) will include N, 
and N,—that is, the remainder of the episode, together with argil- 
laceous beds called ‘“‘Middle Portland.” M. Pellat includes N, also in 
the upper zone; but asit contains Amm. gigas the reason of this is not 
evident. The northern type may also be divided paleontologically, 
independently of the forms which may be supposed introduced from 
the south. ‘The lower portion appears to be characterized by Amm. 
suprajwrensis (formerly quoted as “A. Thurmanni?”), and the upper 
by several species, amongst which it is difficult to choose the most 
characteristic. Belemnites Sowichii, Astarte Semanni, and Discina 
latissima are the chief species almost confined to this portion. These 
divisions would coincide very nearly with those of the southern range ; 
and if with the zone of Cyprina Brongnarti were included the un- 
fossiliferous grey-green limestones of the Meuse and Haute-Marne, 
the periods might be considered synchronous. ‘This is very nearly 
the correlation made by Pellat (59, 68); that is, he recognized in the 
two fossiliferous zones his ‘‘ Lower Portland,” and in the barren zone 
the possible equivalents of his ‘‘ Middle Portland.” His greatest line 
of division, however, is at a different place, namely above N,. This 
arrangement may be most suitable for Boulogne; but it makes the 
Upper Bolonian almost absent from the southern ranges, whereas it 
probably continued there in its varied forms long after the intro- 
duction of clay and clay-loving forms into the Boulonnais. 

Tracing these beds through the areas of their occurrence, 
we find only the lower Bolonian or lithographic limestones in 
the Ardennes and Northern Meuse; and from these the lower 
portion of the rocks assigned to the same series by Buvignier 
must be detached, as containing a more Virgulian fauna, especially . 
Pholadomya acuticostata. ‘The presence of Hwogyra virgula itself 
even in lumachelles cannot be made of any great importance, since 
at Boulogne, throughout the southern range, and even in England 
that oyster certainly survived the introduction of a very distinct 
fauna of far more interest than itself. There isno sign of thinning 


JURASSIC OF THE PARIS BASIN. . 583 


of the Bolonian beds in the Meuse, as Hébert supposed (21); for, as 
Buvignier (22) points out, the smaller quantity there visible 1s com- 
posed of the lowest beds only, and not of diminished representatives 
of all. On entering the Haute-Marne much more is seen, in fact 
the full development, which, however, commenced near Bar-le-Due 
in the extreme limit of the Meuse. The Lower Bolonian will here be 
the ‘¢ Lithographic limestones” of Buvignier and the zone of Ammo- 
mites gigas of Royer and Tombeck, of which, after them, the Bure 
oolite may be taken as the upper limit*. The Upper Bolonian in- 
cludes the more fossiliferous beds, distinguished as carious, spotted, 
and tubulous limestones, and also the lower portion of the series. 
called the zone of Cyrena rugosa, namely the porous limestones, 
which are mostly unfossiliferous, but in places appear to contain 
Natica Marcousana, avery characteristic Bolonian fossil. Passing 
westwards, the tubulous limestones and higher beds are rapidly lost, 
and the series is reduced in the valley of the Aube to the Lower 
Bolonian and the carious limestones. These two are of longer con- 
tinuance, and are found as fully developed and still more fossiliferous 
in the valley of the Yonne, while they have representatives as far as 
Bourges. In the Charente the sandy limestones near Angouléme give 
us fair representatives of the Lower Bolonian, though with a somewhat 
uncommon fauna, somewhat allying it to the Upper. This latter must 
be recognized in the wide-spread limestones with Cyprina Brongni- 
arte and other usual fossils of this horizon. Possibly representatives 
of the Lower Bolonian exist in the Ile d’Oléron, in the limestones at 
the base with A. Giravesianus ; but the great mass seen to the south 
of the harbour of S. Denis is undeniably Upper Bolonian, with the 
same paleontological characters as in the Charente. Neither in the 
Orne and Sarthe departments nor in Normandy are beds so high in 
the series reached. In the Pays de Bray the lowest Upper Jurassic 
rocks are the Virgulian marls; and above them the Bolonian beds 
are well developed. These are considered by M. de Lapparent (69) to 
commence with his “‘beds with Ostrea catalaunica ;” while his ‘‘upper 
clays and lumachelles ” and ‘‘ compact lithographic lmestones ” are 
referred tothe Virgulian. Ithas, however, been shown that the top 
- of the “ lower clays and lumachelles ” is exceedingly similar in cha- 
racter to the base of the Lower Bolonian at Boulogne, that in the 
north these contain the characteristic Tragonia Munceri, and in the 
centre the lithographic limestones contain Amm. gigas, while in the 
south, according to MM. Semann and Graves (39, 8), at Hodenc, a 
locality coloured Portlandian by M. de Lapparent, the upper marls 
(whieh are very thin in the north) contain Amm. gigas and Amm. 
Gravesianus, and are followed immediately by beds with Cyprina 
Brongmart. Hence it is more consonant with other localities to 
commence the Lower Bolonian towards the top of the “lower clays 
and lumachelles,” and end it at the top of the ‘‘ beds with Ostrea 
catalawmca.” The Upper Bolonian, as before noted, commences 
with the “calcareous grit with Anomias” and continues to the top 


* Perhaps the beds at Cirly recorded by Tombeck to contain Amm. gigas 
and A. swprajwrensis, ought to be also included in the Lower Bolonian. 


584 J. F. BLAKE ON THE UPPER 


of the blue marls called “ Middle Portland.” The latter undoubtedly 
correspond to the beds called by the same name at Boulogne; and 
the grits below represent the upper part of the so-called “ Lower 
Portland,” as has been shown by Pellat (46). 

In our own country the Bolonian strata are pretty nearly synony- 
mous with those shown to be separable under the title Upper 
Kimmeridge. Only in the coast-section of Dorsetshire has any dis- 
tinction into Upper and Lower Bolonian been possible as yet; but 
when once it is recognized that the whole is argillaceous, the two 
parts may be some day recognized by their fossils. Throughout the 
basin of Paris, however, and in the Dorset section, the lower beds are 
less fossiliferous, and therefore in England they will be less likely 
to attract attention in inland sections. In separating the Bolonian 
from the Kimmeridge below and the Portland above, there arises in 
England, as at Boulogne, the question as to their limits. In the 
latter place, the ‘‘ Lower Portland ” is taken by Pellat (68) to com- 
mence in the midst of marls, on account of the change of fauna. 
And this change is a remarkable one; for it consists in the intro- 
duction of species which do not specially characterize the episodal 
deposits, but which continue upwards through the whole of the 
Bolonian. So on the coast of Dorset the lower limit of the Bolonian 
must also be drawn in the midst of clays, where the most marked 
introduction of new species commences. This takes place at no very 
well defined line; so that the limit must remain open; in any case 
it will be below bed No. 29 of:the:Kimmeridge Bay section, and may 
be as low as No. 40. At Boulogne,'the ‘‘Middle Portland” has been 
taken to end upwards where the sands cease to be marly and are 
often consolidated into calcareous grits, the lowest beds of the next 
series containing a different fauna, the most remarkable species being 
Cardium Pellati. If we draw the same line in England, it will 
lie immediately beneath the Flinty series of the coast, the Tisbury 
freestone, the Swindon Trigonia-beds, and the rubbly limestones of 
Buckinghamshire. In other words, the ‘Portland Sand” must be 
thrown into the Upper Bolonian, care being taken that the glau- 
conitic and rubbly beds which form the base of the true Portland 
are not included. That the sands ought to be separated from the 
Portland Stone was perceived by Fitton ; and their distinctness from 
the general mass of the Kimmeridge Clay was equally clear to him. 
The fact of their containing a fauna much allied to that of the 
higher parts of what was then called the Kimmeridge Clay, enables 
us rightly to associate the two under a separate and common title, 
the Bolonian. This period both in England and the North of 
France, served as an introduction to the Portland, with the lowest 
beds of which its uppermost strata have several fossils in common. 

The localities in England which show Bolonian strata may now 
be enumerated. In general, of course, all that has been described 
as Portland Sand will now represent part of the Upper Bolonian, 
and, in addition, much that has been termed Kimmeridge Clay. The 
brick-yards at Upway, the fossiliferous beds at the base of the Tis- 
bury section, the sandy “‘ Kimmeridge Clay” of Devizes, so rich in 


JURASSIC OF THE PARIS BASIN. 585 


fossils, the sandy portions below the actual sand on the northern 
slopes of Swindon Hill, the lower part of Shotover Hill with the 
finely laminated clays, the uppermost part of the pit at Hly, and the 
whole of the mass called Upper Kimmeridge in Lincolnshire’, all 
belong to the Bolonian. In most cases no subdivision into Upper 
and Lower can be traced; but the record by Prof. Judd (Q.J.G.S. 
vol. xxiv. p. 237) of Amm. gigas and Amm. Gravestanus in the ‘‘ Port- 
landian ” portion of the Speeton Clay would appear to indicate the 
distinctness of the Lower Bolonian in that locality. It appears 
from this description that the Upper Bolonian in the northern area 
may be divided into 3 minor parts as in the south, viz. the zone of 
Cyprina Brongniarti, the zone of Discina latissima, and the Portland 
Sands, which last might be called the zone of Belemnites Souichi. 

The last series to be considered is the Portlandian. The line of 
junction of this with the Bolonian has already been discussed. The 
beds above this line which may be certainly placed among Jurassic 
deposits, are the Flinty Series, the Building-stones, and the Purbecks 
of the Isle of Purbeck. Towards the close of the Jurassic epoch 
freshwater conditions appear to have set in in various parts of the 
area under study. ‘There is, however, no reason to suppose that 
their introduction in every place was synchronous, but every reason 
to think the contrary; yet by the custom of calling all these beds 
“‘ Purbeck ” their synchronism has been practically affirmed. To 
obviate this it is proposed to include the typical Purbecks, as seen in 
the Isle of Purbeck itself, in a “ Portlandian ” group, of which it will 
form the upper member, while the Flinty series and Building-stones 
form the lower. ‘There can be no objection to this, if it renders cor- 
relation easier. 

It remains, then, only to indicate the localities where these rocks 
occur, and their position in the group. In the southern range 
no possible representative is met with till we reach Bar-le-Duc, 
shortly before entering the Haute-Marne. Here commences a small 
mass of rocks formerly considered Suprajurassic, of which the most 
noteworthy is the vacuolar oolite. From stratigraphical position, this 
might equally well represent the upper part of the Upper Bolonian ; 
but the occurrence in it of Astarte rugosa, a characteristic fossil of 
true Portland rocks, leads to its being placed as Lower Portlandian, 
a correlation to which Pellat (68) agrees (using the term in the 
sense now defined). These beds have been proved (40) to be over- 
lain unconformably by the Neocomian ; so that nothing higher is to 
be found ; but it is a remarkable circumstance that the greatest 
development upwards is just at that locality where all detrital beds 
are at their maximum, opposite the Straits of Dijon. These beds 
soon disappear ; and nowhere else in the same range can similar beds 
be found. In the Charente, however, if we correlate the limestones 
with Cyprina Brongniarti (or its representative) with the Upper 
Bolonian, the succeeding beds with Corbula inflewa must be the base 
of the Lower Portland. Indeed the Portlandian and Bolonian may be 


* In the map in Quart. Journ. Geol. Soc. vol. xxxi. p. 202, the names ‘“‘ Upper 
Kimmeridge” and “‘ Lower Kimmeridge” have been interchanged by error. 


586 J. F. BLAKE ON THE UPPER 


here less distinct than elsewhere ; for if Coquand is right in recording 
OCardium dissimile from the latter, and the shell called Astarte 
rugosa has been rightly determined from Chassors, there is a very 
Portlandian aspect even about the Lower Limestone, though it would 
be impossible, for stratigraphical reasons and from the general cha- 
racter of its fauna, to separate it from the Upper Bolonian. Thus 
there is nothing to absolutely prevent a geologist from considering, 
with Coquand (24), that the gypseous beds at the top belong to the 
Upper Portland, ¢.'e. Purbeck ; but it is more in keeping with proba- 
bility to regard them as taking the place, without any intervening 
gap, of the Lower Portland or true Portland limestone, as Manés 
does (17) when he groups the equivalent beds in the Ile d’Oléron 
with those below. In the latter locality, at St. Denis, it is undoubted 
that beds with Cardium dissimile follow very closely on others with 
Cyprina Brongniarts (or its representative) ; and these we are justified 
in considering Portlandian. They are followed conformably by the 
gypseous beds, which are associated with others containing fossils 
like the beds below; and these too, therefore, are placed as Lower 
Portlandian. In.the Pays de Bray it has been satisfactorily shown 
that above the Upper Bolonian clays come ferruginous grits contain- 
ing true Trigona gibbosa; and these are therefore rightly placed 
by de Lapparent (69) and Seemann on the horizon of the Lower Port- 
landian of the present paper. In the Boulonnais the beds hitherto 
called ‘‘Upper Portland” correspond without doubt to the true 
Portland limestone, but not to the whole of it. Throughout they 
are more or less arenaceous, and correspond lithologically to the 
Flinty series, while their characteristic Ammonite is A. bononzensis, 
and not A. giganteus, which latter is characteristic of the Building- 
stones. Here, therefore, the Portland series is incomplete; and the 
beds which lie at the top and contain Astarte socralis and Cyprids 
are, for this reason, scarcely likely to represent Purbeck beds. They 
are in fact so rubbly and irregular that they cannot be considered 
conformable, and may be of any age, either Purbeck or Wealden ; 
but there is not the slightest proof that they belong to the former. 
Finally, in our own country it has been shown (Q.J.G.8. vol. 
xxxvi.) that the Portland limestone is only complete in Dorsetshire 
and the Vale of Wardour; and in these two districts the freshwater 
(or partially freshwater) strata will belong to the Upper Portlandian, 
and be its only representatives. Justifications for a closer associa- 
tion of these rocks with the Portland marine limestones than is 
allowed by calling them Purbeck, may be found in the occurrence 
in them of Hemicidaris purbeckensis in Dorsetshire, a fossil found 
also in the Upper Bolonian, and of Trigonia densinoda, belonging 
to the Glabrae, recently described by Mr. Etheridge (Q. J.G.S8. vol. 
Xxxvil. p. 247) in the Vale-of-Wardour “ Purbecks.” It is true 
these beds exhibit local unconformities; but that is natural when 
freshwater strata succeed marine; and it is to be noted that “ Pur- 
becks” never lie on any thing but “ Portlands.” The so-called 
Purbecks of Swindon and those of Buckinghamshire, lying on lower 


JURASSIC OF THE PARIS BASIN. 587 


portions of the Portland limestones, will belong to the Lower 
Portlandian. 


In bringing this Part I. to a close, I must not omit to return my 
thanks to those who have assisted me ; especially to M. de Lapparent, 
who enabled me to study the literature of the subject in Paris, and 
who showed me the true Portlandian fossils from the Pays de Bray ; 
to Prof. Hébert, who gave me many valuable indications of the best 
localities to visit, without which my difficulties would have been 
greatly increased ; to M. Rigaux, who has on several occasions guided 
me in the Boulonnais; to MM. Cotteau and de Loriol, who have given 
me copies of some of their writings for reference; and, finally, to the 
Government-Grant Committee of the Royal Society, to whose recom- 
mendation I owe the means of carrying on these researches on the 
Continent. 


EXPLANATION OF PLATE XXVI. 


Comparative diagram sections of Upper Jurassic Rocks, Paris Basin 
and England. 


DiscussrIon. 


The Prestprnr stated that the correlations of the author went 
far to complete our knowledge of the Upper Jurassic rocks in the 
Anglo-Parisian Basin. 

Mr. Huptzston was able to confirm the author’s views concerning 
the correlation of the Upper Jurassic strata in parts of this country 
with those of the Ardennes as regards the lowest beds in question. 
The coincidence in the faunas of the beds in these widely separated 
areas was very remarkabie. He thought Mr. Blake’s observations 
tended to support the views of M. Hébert rather than those of 
M. de Loriol with respect to the importance of the Coral Rag as a 
formation. He did not agree with the separation of the so-called 
“« Pterocerian;” nor could he agree with the author in absorbing the 
greater part of the Kimmeridge Clay into his “Bolonian.” The 
author’s Bolonian was called by the French authors Lower Portlan- 
dian, and was claimed by them as the normal or characteristic de- 
posit; while the Portlandian in the south of England, on a higher 
horizon, was the less constant development, and therefore more 
truly the episode. 

Tne AvrHor, in reply to Mr. Hudleston, stated that the Astartian 
of this country resembled that of Normandy rather than that of 
the Boulonnais; but he agreed with him as to the Pterscerian. He 
thought that the calling of the Bolonian by the name of “ Port- 
landian” had been a continual source of error. 


588 S. S. BUCKMAN ON AMMONITES FROM 


39. A Duscriptive CataLoeur of some of the Spucizs of AMMONITES 
from the Inrertor Ootire of Dorset. By 8. 8. Buckman, Esq. 
(Communicated by James Buckman, Hsq., F.G.8.) (Read 
June 22, 1881.) 


BEFORE commencing a descriptive catalogue of these Ammonites, it 
may be as well to give some idea of the various beds from which 
they come. This has been done before ; but, various mistakes having 
crept in and new researches having given us fresh information, I 
deem it worth while to reintroduce the subject. 

The beds under consideration begin with the ‘‘ Sands” or ‘ Pas- 
sage-beds,” also called Midford Sands. They are from about 100 to 
150 feet, perhaps more, in thickness, with interpolated layers of 
comminuted shells about a foot or so thick, occurring at intervals of 
from 4 to 10 feet. These layers contain a large conglomeration of 
broken shells, so that good specimens are scarce. I have obtained 
from them a Rhynchonella which is probably R. cynocephala(Richard.), 
besides Harpoceras Moorei (Lycett), and several species of Trigonia 
(clavellated and costate), Loma, Astarte, &c.; and I am led to sup- 
pose that these sands are probably equivalent to the lower part 
of the Cynocephala stage of Lycett, in the Cotteswold Hills. 

Above these sands comes the stone of the Inferior Oolite. This 
stone changes at certain levels, both in composition, colour, and hard- 
ness. The Ammonites do not occur all in one bed, but this Inferior 
Oolite stone can be very well divided into four zones, which are ex- 
tremely well marked, but vary very greatly in thickness at different 
localities; and it is probably this variation in thickness, and some- 
times almost complete absence of a zone, that has led to very much 
confusion. 

At the base of the Inferior Oolite Limestone comes the zone of 
Harpoceras Murchisone. It rests on a bed of blue stone, which 
probably belongs properly to the sands. 

The bed in which H. Murchisone occurs is mostly of a ight brown 
colour, sometimes well filled with iron grains, sometimes almost 
altogether lacking them; it is generally very hard. At Bradford 
Abbas it is about 1 foot thick, but on Corton Down from 3 to 4 
feet. 

Above this comes the zone of Harpoceras Sowerbyz. ‘This bed is 
sometimes light yellow with iron grains, sometimes dark blue with 
similar grains, the light yellow being soft and the dark blue hard. 
Its range is about 3 feet at Bradford Abbas; and at Halfway House, 
&e. it is about the same thickness. 

Above this comes the zone of Stephanoceras Humphriesranum, 
which is almost entirely, if not quite, absent at Bradford Abbas. 
At Oborne, however, its thickness is about 5 feet, while at Louse- 


ad 


THE INFERIOR OOLITH OF DORSET. 589 


Hill and Wyke quarries this zone is only represented by two thin 
layers very much charged with iron, the two being only about 
6 inches thick, but containing nearly all the species that one finds at 
Oborne in the Humphriescanwm-zone. 

Above this comes the zone of Cosmoceras Parkinsoni, which varies 
very much in different positions. Near Sherborne it is about 15 to 
20 feet thick, but very unfossiliferous. At Bradford Abbas it is 
about 6 feet thick, the upper part having probably been washed 
away; it is rather unfossiliferous. At Clifton Maybank, however, 
about 2 miles from Bradford Abbas, a small opening in this zone 
was made; and it was found highly fossiliferous, while at Broad 
Windsor, and near Bridport, it contains a large number of species and 
specimens. 

I here give sections of three quarries—one at Oborne, the other 
at Wyke, and the third at Bradford Abbas—to show the different 
beds clearly. 


Fig. 1.—Section at Oborne, near Sherborne. 


Zone of Ammonites Parkin- 4 
sont. 


i 


| Light-coloured stone, 2 feet. 


bo 


Soft sandy stone, 2 feet. 


—————-—-~ -—-— 


| 


Zone of Stephanoceras Hum- 


phriesianum, 
3 | Harder stone, iron grains, 3 feet. 
Sere RS ; ‘ 
Bed with Spheroceras Sauzei \ 4 White marl, with green grains, 
&e. 6 inches. 
5 Hard blue and yellow stone, about 
| 2 feet. 
| 
Probably zone of Harpoceras 
Sowerbyt. 
| 
| Unseen. 
Bed with Rhynchonella rin- t | 
gens. \ | 
| Soft yellow sandstone. 


590 S. 8. BUCKMAN ON AMMONITES FROM 


Fig. 2.—Section at Wyke Quarry. 


( 
i) 
Whitish stone, perhaps 20 feet, 
extending up the hill. 
Zone of Cosmoceras Parkin- 1 
sont. | 
Zone of Stephanoceras Hwin- 9 Dark red stone, filled with iron 
phriestanum. t __| &e., 7 inches. 
f Bluish stone, sometimes yellow, 
Zone of Harpoceras Sewer eat 3 with plenty of iron grains, 
3 feet 10 inches. 
| * HHH X KH ee 
| ; Lighter yellow stone, 2 feet. 
Zone of Harpoceras Murchi- 4 
sone. | 
| 


Lower part hidden. 


Bed No. 2 contains Stephanoceras Humphriesianum, S. Blagdeni, S. Braiken- 
ridgui, Cosmoceras Garantianuwm, and many others peculiar to the zone of 
Steph. Humphriesianum at Oborne. 

xxx Bed with Rhynchonella ringens, Hérault. 


Pa 


THE INFERIOR OOLITE OF DORSET. 591 


Fig. 3.—Section at Bradford Abbas. 
{| 


ER en assras Parkin- 1 White oolite, 6 feet 6 inches. 
soni. 
| 
| 
2 | Marl bed, 3 inches. 
Humphriesianum zone. g Bluish stone, 6 inches. 
c Wr Trony stone, 6 inches. 


| 
| 


Zone of Harpoceras Sowerbyi. 5 | Vellow atonese f2ck 


/ 


6 | Paving bed, 1 foot. 


Zone of Harpoceras Murchi- 


HCE Hard blue stone, 1 foot 2 inches. 


| 
| 
| 
| 
) 
| 
| 
| 
| 
Perhaps representative of S.- ' 
( 
| 
| 
( 
| 


| 
Yellow sands. | 
Bed No. 4 is perhaps the representative of the Spheroceras-Sauzei bed at Oborne. 
Bed No. 2 contains Zerebratula Morieri and Rhynchonella parvula. 


The zone of Harpoceras Murchisone in this district contains :-— 


Harpoceras Murchison (Sow). 
, var. bradfordiense, S. S. 


Rhynchonella subangulata, Dav. 
Terebratula perovalis, Sow. 
Buckm. Waldheimia anglica ( Oppel). 
Rhynchonella subtetraedra, Dav. Terebratula Etheridgii, Dav. 
subdecorata, Dav., young. | —— simplex, Buckm. 


The zone of Harpoceras Sowerbyt contains :— 


Harpoceras Sowerbyi (Miller). Lytoceras confusum, 8. S. Buckm. 
adicrum ( Waagen). Amaltheus subspinatus, S. S. Buckm. 
Levesquei (@ O70.). Astarte excavata, Sow. 

fissilobatum (Waagen). elegans, Sow. 

—— cornu, 8S. S. Buckm. Terebratula Eudesii, Oppel. 


The Spheroceras-Sauzei bed at Oborne contains :— 


Spheroceras Sauzei (7 Ord.). Sphzroceras meniscum (Waagen) ? 
Stephanoceras polymerum (Waagen). | Harpoceras Sowerbyi (Miller), var. 


592 S. S. BUCKMAN ON AMMONITES FROM 


The zone of Stephanoceras Humphriesianum contains :— 


Stephanoceras Humphriesianum (Sow.). | Stephanoceras Deslongschampsii 


Harpoceras cycloides (d’ Ord.). (Defrance). 
Cosmoceras subfurcatum (Schloth.). Cosmoceras Caumontii (@’ Orb.). 
Garantianum (d’ Oro.). Spheeroceras Wrighti, S. S. Buckm. 
Haploceras oolithicum (@ Orb.). Oppelia subcostata, J. Buckman. 
Harpoceras Edwardianum (d’Orb.). Ammonites cadomensis, Defrance. 
Lytoceras Hudesianum (@’ Orb.). Perisphinctes Davidsoni, S. S. Buckm. 
Stephanoceras Blagdeni (Sow.). Terebratula spheroidalis, Sow., 
Stephanoceras Braikenridgii (Sow.). abundant. 
linguiferum (d’Ord.). Astarte obliqua, Desh. 
Spheeroceras Brongniarti (Sow.). Rhynchonella senticosa, von Buch. 
—— Gervillii (Sow.). Terebratula Buckmani, Dav. 


This Humphriestanum zone is the * Fossil Bed” in quarries round 
Sherborne, where it is so well developed. It has been put down 
as the equivalent of the Sowerbyz zone or “ Fossil Bed ” (misleading 
term) of Bradford Abbas and neighbourhood under the name of 
‘the Cephalopoda bed of the Inferior Oolite ;” and the extraordinary 
difference in the species has been attributed to the difference in the 
locality. 

The fact is that the Humphriestanwm zone is almost absent 
at Bradford Abbas, but is represented near Halfway House by a 
thin band of ironstone, about 6 inches in thickness, which overlies 
the Sowerbyt zone, and which contains a list of species which agrees 
with those from the Humphriescanum zone at Sherborne ; while the 
Sowerby zone at Sherborne is often not quarried, and therefore has 
escaped notice, as it was supposed the quarrymen had reached 
the sands, as they do at Bradford Abbas. The term “fossil 
bed,” too, was probably answerable for part of the confusion, and 
shows the necessity for calling beds by the name of some character- 
istic fossil, and not by some local appellation which may often denote 
beds at entirely different horizons. 


The zone of Cosmoceras Parkinsoni contains :— 


Cosmoceras Parkinsoni (Sow.). Stephanoceras zigzag (d’Orb.). 
Garantianum (d’Ord.). Terebratula Phillipsi, Morris. 
Oppelia Truellii (d@’ Ord.). spheroidalis, Sow. 
subradiata (Sow.). Stephani, Dav. 
Spheroceras polymorphum (d’O7r9.). Rhynchonella spinosa (Schloth.). 

dimorphum (d@ Orb.). Terebratula globata, Sow., variety ? 
Perisphinctes Martinsii (@’Ord.). 


Besides the Ammonites here mentioned from these four zones, I 
have separated about 50 more species, which, so far as | am aware, 
have not yet been described. ‘These rocks are also well stocked 
with Gasteropoda, of which there are about 150 different species ; 
and Lamellibranchiata are very abundant; of Brachiopoda we have 
about 40 species made out. 

It may be as well before describing the species to explain a few 
terms made use of in these descriptions. 

The Inner portion of whorl is that which is nearest to the centre 
of the Ammonite, otherwise called the dorsal part. 


tat racoen 


THE INFERIOR OOLITE OF DORSET. 593 


Outer portion is the opposite, otherwise called the ventral area. 

Shoulder is where the inner portion of the whorl meets the pre- 
ceding whorl, and is an important point in the diagnosis of Ammo- 
nites ; some shoulders are square, as in Harpoceras Tessonianum 
some concave, see Harp. Murchisone ; some convex, some merely 
sloping, de. 

Termination or mouth-border, i.e. the completion of the body- 
chamber. 

The termination is variously shaped: some have a plain semi- 
lunar band, as Stephanoceras Humphriesianum; others, ears of 
various shapes, set up either nearly altogether on the ventral area, 
as in Steph. Braikenridgzt, or on the sides and projecting straight, as 
in Spheroceras Sauzei (d’Orb.); others have a termination like an §, 
as Harpoceras concavum; this is called the double bend; others 
have this double bend with a horn projecting from the middle 
others have a single bend, like Amaltheus spinatus (Bruguicre). 

In the following Tables I have roughly classified the Ammonite 
according to their variously shaped terminations :— 


1. Semilunar termination. 


Stephanoceras Humphriesianum(Sow.).| Stephanoceras Deslongschampsii (De- 
Cosmoceras Garantianum (@’ OQrd.).- france). 
Stephan. polymerum (Waagen). | Perisphinctes Davidsoni, S. S. Buckm. 


2. Semilunar termination, with a deep furrow first, and, sometimes, 
a raised lip. 


Spheroceras Mansellii, J. Buck. 


Spheeroceras Gervillii (Sow.). 
dimorphum (d’ Oré.). 


Brongniarti (Sow.). 
—— Wrightii, SS. Buckm. 


3. Spathulate ears, projecting from a little on each side of the 
ventral area. 


Stephanoceras Braikenridgii (Sow. ys | Perisphinctes Martinsii (D’ Or0.). 
Blagdeni (Sow.). 


4. Kars like No. 3, but projecting from the side. 


Spheeroceras Sauzei (d’ Ord.). | Stephanoceras linguiferum (d’ Ord.). 
5. Termination, double bend without any horn. 
Harpoceras concavum (Sow.). | Harpoceras cycloides (@’ Ord.). 
Murchisonex, var. bradfordiense, adicrum ( Waagen). 


S. S. Buckm. 
— Moorei (Lyc.). 


Haploceras oolithicum (d@’Ord.). 


6. Termination, double bend with horn. 


Oppelia subradiata (Sow.). 


Harpoceras cornu, S. S. Buckin. 
Harpoceras Levesquei (d’O7b.). 


Cosmoceras Parkinsoni (Sow.) *. 
Harpoceras Edwardianum (d’ Orb.). 


* T have placed it here on the authority of d’Orbigny’s figures, my own spe- 
cimen not showing the termination exactly enough. 


Q.J.G.8. No. 148. 2R 


O94 Ss. S. BUCKMAN ON AMMONITES FROM 


7. Termination like No. 6, but not produced on ventral area. 


Cosmoceras subfurcatum (Schloth.). | Cosmoceras Caumontii (d@ Ord.). 


8. Termination, a spathulate ear each side, and one on the ventral 


are€a. 
Ammonites cadomensis, Defrance. 


9. Termination, a plain single bend, more or less produced on 


ventral area. 
Amaltheus spinatus (Brugwiére) in the Lias. 
Inf. Oolite. Amaltheus subspinatus, S. S. Buckm. 
Lytoceras confusum, S. S. Buckm. 


We will now proceed to give a short notice of the various species 
of Ammonites recognized in this district. 


STEPHANOCERAS HuMPHRIESIANUM (Sow.). 


1825. Ammonites Humphriesianus, Sow. Min. Conch. pl. 500, 
middle figure. 

1830. A. Humphriesianus (Sow.), Zieten, Petref. pl. 67. fig. 2. 

1845. A. Humphriesianus (Sow.), d’Orb. Pal. Franc. Terr. Jurass. 
pls. 133, 134, 185. figs. 1, 2. 

1849. A. Humphriesianus (Sow.), Quenstedt, Ceph. tab. 14. 
figs. 7, 11; Quenstedt, Jura, pl. 54. figs. 2, 3, 4. 

1849. A. coronatus oolithicus, Quenstedt, Ceph. tab. 14. fig. 4. 

1856. A. Humphriesianus (Sow.), Oppel, Juraformation, p. 376, 
no. 40. 

1856. A. subcoronatus, Oppel, Juraformation, p. 376, no. 39. 

1856. A. Bayleanus, Oppel, Juraformation, p. 377, no. 43. 

1854. A. Humphriesianus (Sow.), Morris, Catalogue, p. 292. 

1878. Stephanoceras Humphriesti (Sow.), Bayle, Explic. de la 
Carte Géol. de la France, vol. iv. pl. 51, figs. 2, 3. 

1881. A. Humphriesianus (Sow.), J. Buckman, Quart. Journal 
Geol. Soc. vol. xxxvii. p. 65, fig. 6. 


Bayle in ‘ Explication de la Carte Géologique de la France,’ vol. 
iv. plate 51. figs. 2, 3, shows what I take to be a rather peculiar 
variety of Stephanoceras Humphriesianum. 

Locahittes. Oborne, and near Sherborne, Burton Bradstock, and 
other places: rather common. 

Quenstedt, under the name of Amm. coronatus oblithicus, figured the 
young form of the thick variety of this species, which Oppel also 
mentioned by the name of subcoronatus. Oppel also separated the 
thin form, under the name of Bay/eanus, observing that it occurred 
lower than Humphriesianus proper. My own observations, however, 
do not agree with this, as I have found both the thin and thick 
varieties together; and Mr. D. Stephens assures me that they both 
occur together at Milborne Wick, the thin variety being the com- 
moner. I have, however, seen the necessity for distinction, and 
have kept Oppel’s names for the varieties, the thick variety being 
Stephan. Humphriesianum, var. subcoronatum (Oppel), and the thin 
Stephan. Humphriesianum, var. Bayleanum (Oppel). 


THE INFERIOR OOLITE OF DORSET. 595 


SrEPHANOCERAS DxEstonescHampsit (Defrance). 


1845. Ammonites Deslongschampsi (Defr.), d’Orb. Terr. Jurass. 
pl. 138. figs. 1, 2 

1856. A. Deslongschampsu (Defr.), Oppel, Juraform. p. 378, 
no. 44. 

1880. Stephanoceras Deslongschampsu (Defr,), Wright, Lias Amm. 
Palzont. Soc. 1880, p. 224, fig. 116. 

Locality. Humphriesianum zone, Oborne: very rare. 

The termination of this species is a fine semilunar border. One 
beautiful specimen showing this was obtained from Oborne by E. 
Cleminshaw, Esq., F.G.S. 


SrEPHANOCERAS BragpENt (Sow.). 


1818. Ammonites Blagdent, Sow. Min. Conch. pl. 201. 
1818. A. Banks (Sow.), Min. Conch. pl. 200. 
1830. A. coronatus (Schloth.), Zieten, Verstein. Wirttemb. tab. i. 
fig. 1. 
1845. A. Blagdeni (Sow.), @Orb. Terr. Jurass. tab. 132. 
? A. coronatus, Quenstedt, Der Jura, tab. 51. fig. 1. 
1849. ? A. coronatus, Quenstedt, Ceph. tab. 14. fig. 1. 
1854. A. Blagdeni (Sow.), Morris, Catalogue, p. 290. 
A. Blagdent (Sow.), Morris and Lycett, Great Ool. Moll. 
tab. 14. fig. 3. 
1856. A. Blagdeni (Sow.), Oppel, Juraformation, p. 374, no. 38. 
1880. Stephanoceras Blagdent (Sow.), Wright, Lias Ammonites, 
Palzont. Soc. 1880, p. 251, figs. 157, 158. 
Localities. This species occurs in the Humphriesianum zone of 
Oborne, Sherborne, &e. | 
Stephan. (Ammonites) Banksii (Sow.) is merely a very large 
variety of this species. 


STEPHANOCERAS BRAIKENRIDEII (Sow.). 


1818. Ammonites Braikenridgui, Sow. Min. Conch pl. 184. 
1845. A. Br OAL (Sow.), @Orb. Terr. Jurass. pl. 135. 
figs. 3-5. 
We Braikenridgu (Sow.), Quenstedt, Der Jura, tab. 54. fig. 5. 
A. Braikenridgit (Sow.), ? Morr. & Lyc. Great Ool. Moll. pl. 14. 
fig. 1. 
” 1854. A. Braikenridgii (Sow.), Morris, Catalogue, p. 290. 
1856. A. Braikenridgiu (Sow.), Oppel, Juraformation, p. 377, 
no. 42. 
1880. Stephanoceras Braakenridgu (Sow.), Wright, Lias Ammo- 
nites, Palzont. Soc. p. 251, figs. 159, 160. 
1881. A. Braikenridqu (Sow. ), J. Buckman, Date Journ. Geol. 
Soc. vol. xxxvii. p. 62, fig. 3. 
Localities. Humphriesianum zone at Oborne, and othise quarries 
near Sherborne. 
The termination of this species is a fine ear on each side, as is 
well shown in @’Orbigny. The length of ears varies very much; 
2R 2 


596 S. 8S. BUCKMAN ON AMMONITES FROM 


and I have some specimens which very much resemble Steph. 
Humphriesianum, var. subcoronatum (Oppel), both in the number of 
ribs and in other respects, but have short ears. 


STEPHANOCERAS LINGUIFERUM (d Orb.). 


1845. Ammonites linguiferus, d’Orb. Terr. Jurass. pl. 136. 

1856. A. linguiferus (d’Orb.), Oppel, Juraform. p. 376, no. 40. 

Locality. Zone of Stephan. Humphriesianum at Oborne: very 
rare. 

I possess one specimen with the termination ; it has a small plain 
ear projecting from the side. 


STEPHANOCERAS ziezaé (d’Orb.). 


1845. Ammonites zigzag, d’Orb. Pal. France. pl. 129. figs. 9, 
MO, Wk 

1856. A. zigzag (d’Orb.), Oppel, Juraformation, p. 378, no. 45. 

Of this species there are two forms, a thick and a thin one. They 
are found, however, in the same bed in the same localities, and are 
similar in all other respects. 

The thick primary ribs, so conspicuous in the small specimens, 
gradually disappear as the fossil becomes larger, and give place to 
small rounded primary ribs, which come much closer together, some- 
times dividing into secondary ribs, and sometimes passing straight 
round the ventral area without dividing at all. 

Localities. Parkinsoni zone at Broad Windsor, Crewkerne Sta- 
tion, &e. 

I have no specimen with the termination ; but the body-chamber 
is very long, as I possess one specimen with a body-chamber nearly 
one whole whorl in length, and yet it does not show even a sign 
of the termination. 


STEPHANOCERAS POLYMERUM (Waagen). 

1867. Ammonites polymerus, Waagen, Geogn. Pal. Beitrage, 
p- (99) 605. 

1845. A. Brongmarti, VOrb. (non Sowerby), Terr. Jurass. pl. 137. 
figs. 1, 2. 

1881. A. Brong gmart, J. Buckman (non aa Quart. Journ. 
Geol. Soc. vol. xxxvii. p. 64, fig. 5. 

Locality. This species occurs at Oborne, in a whitish sort of 
marl, with Spheroceras Sauzet. 

In the young state it very much resembles the thin form of 
Stephan. Humphriesicanum; but when adult it is far more like a 
Spheeroceras. 


Spymroceras Savuzer (d’Orb.). 


1845. Ammonites Sauzei, d’Orb. Terr. Jurass. tab. 149. 

1856. A. Sauzei (d’Orb.), Oppel, Juraformation, p. 375, no. 37. 

1867. A. Sauzec (d’Orb.), Waagen, Geogn. Paldont. Beitrage, 
p- (100) 606. 


THE INFERIOR OOLITE OF DORSET. 597 


1878. Spheroceras contractum, Bayle (non:Sowerby), Explication 
de la Carte Géol. de la France, vol. iv. pl. 53. figs. 1, 2. 

1881. A. Sauzec (VOrb.), J. Buckman, Quart. Journ. Geol. Soc. 
vol. xxxvil. p. 62, no. 6. 

Localities. Marly bed below the zone of Stephan. Humphriesianum 
at Oborne and near Sherborne: rather scarce. 


SpH#=ROcERAS Bronenrarri (Sow.). 


1817. Ammonites Brongniarti, Sow. Min. Conch. tab. 184 a. fig. 2. 

1845. A. Gervilliz, dV’ Orb. (non Sow.), Terr. Jurass. pl. 140. figs. 3-8 
(non 1, 2, non Sowerby). 

1849. A. Brongniarti (Sow.), Quenstedt, Ceph. tab. 15. fig. 9. 

1856. A. Brongniarti (Sow.), Oppel, Juraform. p. 375, no. 35. 

1867. A. Brongniarti (Sow.), Waagen, Geogn. Pal. Beitriige, 
p- (96) 602. 

1878. Spheroceras Brongniarti (Sow.), Bayle, Explic. de la Carte 
Géologique de la France, pl. 53. figs. 3, 4, 5. 

Localities. Humphriesianum zone at Oborne, near Sherborne, &c. 
Rather common. 

The species that I have placed under this name is the one figured 
by Sowerby with the closed umbilicus ; and the other I have called 
Spher.Gervillii. I do this because most authorities have so taken it ; 
d’Orbigny, however, transposed them, whether rightly or not I am 
unable to say without seeing Sowerby’s original specimens. There 
seems to have been some misprint or confusion with regard to them 
in Sowerby’s work. 


SPH#ROCERAS GERVILLII (Sow.). 


1817. Ammonites Gervilli:, Sow. Min. Conch. pl. 1844. fig. 3. 

1849. A. Gervillu (Sow.), Quenstedt, Ceph. tab. 15. fig. 11. 

1856. A. Gervillit (Sow.), Oppel, Juraform. p. 375, no. 36. 

1867. A. Gervillu (Sow.), Waagen, Geogn. Palaont. Beitrage, 
p- (99) 605. 

1878. Spheroceras Gervilla (Sow.), Bayle, Explic. de la Carte 
géologique de la France, pl. 53. figs. 6, 7. 

1881. A. Gervillii (Sow.), J. Buckman, Quart. Journ. Geol. Soc. 
vol. xxxvil. p. 63, fig. 4. 

Localities, Humphriesianum zone at Oborne, near Sherborne, &c. 
Not so common as Spher. Brongnarti. 


SPH HROCERAS MENISCUS (Waagen) ? 


1867. Ammonites meniscus, Waagen, Geogn. Paliont. Beitrage, 
p- (96) 602. 

1875. A. Gervillii, d’Orb. pars (non Sowerby), pl. 140. figs. 1, 2. 

Mr. E. Cleminshaw found one specimen at Oborne in the Sauzei 
bed, which I have referred to this species. 


SpuHz@rRoceras Mawsetiir (J. Buckman). 


1881. Ammonites Mansellii, J. Buckman, Quart. Journ. Geol. 
Soc. vol. xxxvil. p. 64, no. 18. 


598 S. 8. BUCKMAN ON AMMONITES FROM 


This species, when young, is extremely like Spher. Brongniarti 
(Sow.), plate 184 a. fig. 2, but when larger is easily distinguished 
by its very fine and numerous bifurcating ribs and its very square 
ventral area, especially just by the termination, and its far greater 
breadth. It also attains a larger size than Spher. Brongniarti. The 
termination is also peculiar—first a small lip, then a deep furrow, 
then another lip and band. 

This species was named but not figured by Mr. J. Buckman in 
his paper read before the Geological Society. 

Dimensions. Diameter 1°75 inch; umbilicus 0°33; aperture 
across 1:20; aperture back to front 0°45. 

Locality. Clatcombe, near Sherborne. Two fine specimens were 
obtained from this place by T. C. Maggs, Esq. I do not know of 
any other specimens. 


SPHHROCERAS POLYMORPHUM (d’Orb.). 


1845. Ammonites polymorphus, @Orb. Terr. Jurass. pl. 124. figs. 
1, 2, 3, 4, 5? 6? 

1849. A. Parkinsoni inflatus, Quenstedt, Ceph. Tab. iu. figs. 
6 and 7. 

1854. A. polymorphus (d’Orb.), Morris, Catalogue, p. 294. 

1856. A. polymorphus (d’Orb.), Oppel, Juraform. p. 382. 

We have undoubtedly found in this neighbourhood specimens 
which exactly correspond with those figured by d’Orbigny, ‘ Ter- 
rains Jurassiques,’ pl. 124. figs. 1, 2, 3,4; but we have not found 
any to correspond with figs. 5 and 6; and I have my doubts about 
figs. 5 and 6 really being the adult form of the others. Oppel, too, 
in quoting A. polymorphus, d’Orb., leaves out nos. 5 and 6 (see 
Oppel, ‘ Juraformation,’ p. 382, no. 54). 

Localities. Zone of Cosmoceras Parkinsoni at Broad Windsor, Bur- 
ton Bradstock, and other places. 


SPH#ROCERAS DIMORPHUM (d’OrD.). 


1845. Ammonites dimorphus, dOrb. Terr. Jurass. pl. 141. 

1854. A. dimorphus (d’Orb.), Morris, Catalogue, p. 291. 

This species varies somewhat in the width of its umbilicus. Some 
specimens have the umbilicus nearly closed, while others of the same 
size have the umbilicus much larger, so that a portion of the 
inner whorls can be seen. 

In the adult form the umbilicus widens rather quickly, as is 
well shown by d’Orbigny, pl. 141. fig. 1. 

The variety represented by d’Orbigny (pl. 141. figs. 3 and 4) 
has, so far as I know, not yet been obtained from this district. 

I possess only one specimen with the termination. It is a 
form with a rather wide umbilicus. The termination is merely 
a small furrow and a semilunar lip beyond. The specimen came 
from Bradford Abbas. 

Localities. Zone of Cosmoceras Parkinsont at Halfway House, 
Broad Windsor, Stoford, and other places. 


THE INFERIOR OOLITE OF DORSET. 599 


Spom{RocerRAS Wrieutir, S. 8. Buckman, n. sp. 

1849. Ammonites microstoma, Quenstedt (non dOrbigny), Cepha- 
lopoda, tab. 15. fig. 6.. 

Localities. Humphriesianum zone, Oborne, and near Sherborne: 
common. ; 

This species was figured by Quenstedt under the name of A. au- 
crostoma; but it does not at all agree with the figure of A. micro- 
stoma given by d’Orbigny (who named it) in ‘ Terr. Jurassiques.’ 

Dimensions. Diameter, adult, 1°59 inch; umbilicus 0:55 inch; 
umbilicus, omitting the last chamber, 0°22 inch; breadth, same 
place, 0°85 inch; breadth across termination 0°79 inch; termina- 
tion, from back to front, 0°50 inch. 

Length of body-chamber very nearly one whole whorl. 

Named in compliment to Dr. Wright, of Cheltenham, who is so 
ably working at the Lias Ammonites. 


Cosmocrras Parxinsont (Sow.). 

1821. Ammonites Parkinsoni, Sow. Min. Conch. pl. 307. 

1845, A. Parkinsoni (Sow.), d’Orb. Ter. Jurass. pl. 122, variety. 

1849. A. Parkinson gigas, Quenstedt, Ceph. tab. ii. fig. 1 (adult 
form). 

1854. A. Parkinson (Sow.), Morris, Catalogue, p. 294. 

1856. A. neuffensis, Oppel, Juraformation, p. 378, no. 48. 

1856. A. Parkinsoni (Sow.), Oppel, Juraformation. 

1849. A. Parkinsoni depressus, Quenstedt, Ceph. tab. i. fig. 5. 

1878. Parkinsonia Parkinsoni (Sow.), variety, Bayle, Exp. de la 
Carte géol. de la France, vol. iv. pl. 67. figs. 2, 3. 

1878. Parkinsonia neuffensis, Bayle ?, pl. 67. fig. 1. 

This species is somewhat variable, and has been much misunder- 
stood. The figures given by d’Orbigny (‘ Terr. Jurass.’ pl. 122. figs. 1 
& 2) are evidently of a variety of Sowerby’s species. This same 
variety is again figured by Quenstedt under the name of A. Par- 
kinsont depressus ; and as such it was quoted by Oppel (‘ Juraforma- 
tion’) as asynonym for Parkinson. It was also figured by Bayle 
under the name of Parkinsonia Parkinsoni (see synonyms). 

This variety differs from Sowerby’s type in having far fewer ribs, far 
less inclusion (consequently the whorls are not so broad from front 
to back), and in possessing rather marked tubercles. 

To this variety I give the name of Cosmoceras Parkinsoni, var. 
rarecostatum, 8. S. Buckman. 

The fragment figured by Quenstedt (Ceph. tab. ii. fig. 1) under 
the name of Ammonites Parkinsoni gigas is merely the very adult 
form of the true Parkinsoni; and this adult form occurs at Half-way 
House of very large size, one specimen measuring 19 inches across, 
and yet lacking a large portion of the outer chamber. This adult 
form was named by Oppel Ammonites neuffensis, he giving Quen- 
stedt’s figure as a reference. Bayle has also figured a specimen as 
Am. neuffensis ; but I consider that his figure most probably repre- 
sents merely a rather fine-ribbed variety of a medium-sized true 
C. Parkinsons. 


600 S. S. BUCKMAN ON AMMONITES FROM 


The true C. Parkinsoni occurs in great numbers at Halfway House, 
also at Bradford Abbas, Burton Bradstock, and Sherborne. 

The variety rarecostatum, 8. 8. Buckm., occurs in the same beds 
at Sherborne, Bradford Abbas, Burton Bradstock, Broad Windsor, 
&c., but is not so common. 

Cesmoceras Parkinsoni is characteristic of the upper beds of the 
Inferior Oolite. The very adult forms lose all their ribs and also 
the division on the ventral area, and are very different from the 
ordinary young form which was figured by Sowerby. 

Bayle has separated this species from the genus Cosmoceras under 
the generic name of Parkinsonia. I am rather of opinion that it, 
and also Cosm. Caumontu (d’Orb.), should be so separated. 

The termination of this species is somewhat produced on the ven- 
tralarea. I possess one specimen with the termination on the outer 
half of the whorl well shown, but am unable to say whether the 
species possesses a horn on the side, or merely a plain double curve. 
Length of body-chamber = of a whorl. 


Cosmocrras Caumontit (d’Orb.), 
Ammontes Caumontu, @Orb. Terr. Jurass. pl. 138. figs. 3, 4. 
Locality. It occurs at Oborne in the zone of Stephan. Humphriesi- 


anum, but 1s scarce. 
The termination is merely a small, very thin, narrow ear. 


Cosmoceras GaRantranum (d’Orb.). 

1845. Ammonites Garantianus, @Orb. Terr. Jurass. pl. 123. 

1849. A. Parkinsoni dubius, Quenstedt, Ceph. tab. ii. fig. 9. 

1854. A. Garantianus (d’Orb.), Morris, Catalogue, p. 292. 

1856. A. Garantianus (d’Orb.), Oppel, Juraformation, p. 381, 
no. 93. 

Localities. Humphriesianum zone, and less commonly in the lower 
part of the Parkinsona zone, Oborne, Louse Hill, Wyke: rather 
common. 

The termination is a semilunar band. 


CosmocERAS suBFURCATUM (Schlotheim). 

1830. Ammonites subfurcatus (Schlotheim), Zieten, Verst. Wiurt- 
temb. pl. vii. fig. 6. 

1845, A. niortensis, d’Orb. Terr. Jurass. pl. 121. figs. 7-10. 

1849. A. Parkinsons bifurcatus, Quenstedt, Ceph. tab. 11. fig. 11. 

1856. A. subfurcatus (Zieten), Oppel, Juraform. p. 381, no. 52. 

Zicten figures this species, giving Schlotheim as the authority for 
the name. 

Zieten’s figure shows a fossil witha large number of bifurcating 
ribs, which is the exception and not the rule, this species usually 
having very few bifurcating ribs, as is shown by d’Orbigny, who 
figured it under the name of A. mortensis. 

[There is also a fossil, of which a few specimens have occurred 
at Oborne, which is probably a variety of this species. It has very 


THE INFERIOR OOLITE OF DORSET. 601 


much the shape &c. of Cosmoceras subfurcatum, and is ornamented 
with four rows of sharp spines, Just where they occur in Cosm. 
subfurcatum ; but it is entirely destitute of ribs. | 

Locality. Higher part of the Humphriesianum zone at Oborne. 

The termination is a spathulate ear, slightly wider towards the 
extremity. 

Morris, in his Catalogue, put this species down as a synonym 
of Cosmoceras Parkinson; but it is quite distinct from it, espe- 
cially in its mode of volution. It also occurs lower, viz. in the 
zone of Stephan. Humphriesianum. 


Lytocrras Evpsstanum (d’Orb.). 


Ammonites Hudesianus, d’Orb. Terr. Jurass. pl. 1238. 

A, Hudesianus (d’Orb.), Quenstedt, Der Jura, tab. 54. fig. 8. 

A, Eudesianus (d’Orb.), Oppel, Juraformation, p. 373, no. 29. 

This species was for a long time put down for the Inferior 
Oolite of this district by the name of A. cornucopiw, Young. 

Locality. It occurs in the zone of Stephan. Humphriesianum at 
Oborne, but is rare. 


LytocrRas ToRULOSUM (Schubler) ? 

Ammonites torulosus (Schibler), d@Orb. Terr. Jurass. pl. 102. 
figs. 1, 2. 

I have one fragment from the sands near Bridport, but am not 
quite certain if it belongs to this species. 


Lytoceras conrusum, 8. 8. Buckman, n. sp. 

Syn. ? Ammonites jurensis, Morris, Catalogue, p. 292. 

This species approaches nearly to Lytoceras jurense (Zieten), and 
has been quoted from this district under that name. It, however, 
differs from it in a great number of points—namely, the shape of 
the aperture, which is nearly trianglar, in the very square shoulder, 
in the number of whorls, and the lobes, which are more compli- 
cated than in L. jurense (Zieten). Inclusion is small. The test 
is quite plain, but hasafew marked lines of growth in the young 
form, which also has not the peculiar square shoulder much marked. 

In occurs in the Sowerbyi zone at Bradford Abbas, Halfway 
House, Bradford Abbas railway-cutting, &c. 

At Half-way House the specimens are of an enormous size, one 
measuring as much as 17 inches in diameter. 

The species is somewhat abundant; but small specimens are very 
rare. 

[ have one small specimen, from Bradford Abbas, with the ter- 
mination. It is a plain bend forward towards the ventral area. 

Dimensions. Young form, diameter 1:25 inch; umbilicus 0:43 
inch; aperture, length 0-47 inch, breadth 0-40 inch ; inclusion about 
0:03 inch. 

Adult form, diameter 15:90 inches ; umbilicus 7-20 inches ; aper- 


602 S, 8S, BUCKMAN ON AMMONITES FROM 


ture, back to front, 4:50 inches, across about 53 inches, inclu- 
sion 0-80 inch. 

The aperture is, as nearly as possible, an equilateral triangle 
with a small piece out, caused by the inclusion. The ventral area, 
however, is a good deal rounded. 


PrRISPHINCTES PYemmuM (d’Orb.). 
Ammonites pygmeus, d’Orb. Terr. Jur. pl. 129. figs. 12, 13. 
I have one specimen of this species from the neighbourhood. 


PerispHinctes Marrtinsit (d’Orb.). 


1845. Ammonites Martins, d’ Orb. Terr. Jurass. pl. 125. 

1854, A. Martinsa (d’Orb.), Morris, Catalogue, p. 293. 

1856, A. Martinsic (d’Orb.), Oppel, Juraformation, p. 387. 

Localities. Lone of Cosmoceras Parkinsoni at Halfway House, 
Bradford Abbas, Sherborne, and Burton Bradstock, but is rather 
scarce. 

[At Oborne, in the Yumphriesianum zone, we find a very thin flat 
Ammonite with the furrows well marked. It is far more involute 
than Perisphinctes Martinsii, the amount of involution being nearly 
half the preceding whorl. It has a semilunar mouth-border, which 
projects well forward on the ventral margin. 

It has hitherto been put down as the same as P. Martinsw; but 
I feel certain that it should be separated as distinct from it, both on 
account of its stratigraphical position and also its lobes and other 
features. I propose forit the name Perisphinctes Davidsont, in com- 
pliment to Thos. Davidson, Esq., F.R.S. | 

Ihave one specimen of Perisph. Martinsii, d’Orb., with the termi- 
nation. It has two long ears, the inner edges of which project well 
forward till they touch the inner whorl. This termination is well 
represented by d’Orbigny, Terr. Jurass. pl. 125. fig. 1. The termi- 
nation represented in fig. 3 may possibly have been taken from a 
specimen of Perisph. Davidsoni, 8.8. Buckm. 


Harpoceras Sowrrsyt (Miller). 

1821. Ammonites Sowerbyr (Miller), Sow. Min. Conctt tab. 213. 

1821. A. Brown, Sow. Min. Conch. tab. 263. figs. 4, 5. 

1845. A. Sowerby: (Miller), var., d’Orb. Terr. Jurass. pl. 119. 

1849. A. Sowerbyi_(Miller), Quenstedt. Ceph. p. 374. 

1854. A. Sowerbyz (Miller), Morris, Catalogue, p. 295. 

1867. A.Sowerbyi (Miller), var., Waagen, Geogn. Palaont. Beitriige, 
pl2ie ties 2. 

1856. A. Sowerby: (Miller), Oppel, Juraformation, p. 369, no. 20. 

The figure given by d°Orbigny, pl. 119, represents a variety of 
this species, and it differs from it in several points. Waagen has 
also figured, pl. 27. fig. 6, a peculiar variety. 

Localities Ke. Harpoceras Sowerbyt is a scarce fossil, the species 
which has been put down by that name as occurring in this district 
being really Harp. adicrum (Waagen). ‘The chief characteristics of 
Harp. Sowerbyt are a very large keel, and sides which slope in gra- 


THE INFERIOR OOLITE OF DORSET. 603 


dually to join the inner whorl. It occurs at Sherborne and Bradford 
Abbas, but is rare. 

The variety figured by d’Orbigny occurs at Bradford Abbas, but 
is also rare. 


HARPOcERAS ADICRUM (Waagen). 


1867. Ammonites adicrus, Waagen, Geogn. Pal. Beitrage, pl. 25. 
hie. 7. 

Localities. Zone of Harpoceras Sowerbyi at Bradford Abbas, Half- 
way House, Louse Hill, and many other places; rather abundant. 
This species often attains a very large size. 

Dr. Waagen has very properly separated this species from Harpo- 
ceras Sowerby2 (Miller). It is sometimes rather difficult to separate 
it from some of the varieties of that species; but it is really distinct 
In possessing a small rounded keel, as opposed to the large sharp 
one of Harp. Sowerbyt, also a larger umbilicus and large distinct ribs, 
when adult, at a time when Harp. Sowerbyi is smooth. 


H4RPOCERAS FISSILOBATUM (Waagen). 


Ammonites fissilobatus, Waagen, Geogn. Pal. Beitrige, pl. 27. 
ie, db 

Locality. This species occurs at Sandford-Lane quarry near Sher- 
borne. I believe it is in the zone of Harpoceras Sowerbyi; but I 
have never seen this quarry worked. It is a common fossil there. 


HARPOCERAS CONCAVUM (Sow.). 


1812. Ammonites concavus, Sow. Min. Conch. pl. 94. fig. 2. 

1881. A. concavus (Sow.), J. Buckman, ‘“‘ Terminations of Ammo- 
nites,” Quart. Journ. Geol. Soc. vol. xxxvii. p. 60, fig. 1. 

Localities. This species is common in the Sowerbyt zone at Brad- 
ford Abbas, Halfway House, Sherborne, &c. 

The termination is a curve like an §; that is, it rises from the 
inner whorl, curving forward, then rounded and curving back, and 
then produced on the ventral area. It has no horn, as was sup- 
posed when it was figured in the Geological Journal. The species 
that d’Orbigny has figured by this name (Terr. Jurass. pl. 116) 
does not seem to me at all to agree with Sowerby’s figure. 


Harrocreras Murcutson x (Sow.). 


1829. Ammonites Murchisone, Sow. Min. Conch. tab. 550. 

1825. A. corrugatus, Sow.? Min. Conch. tab. 451. fig. 3. 

1830. A. Murchisone (Sow.), Zicten, Verstein. Wirttemb. tab. vi. 

1845. A. Murchisone (Sow.)?, d’Orb. Terr. Jurass. pl. 120. figs. 1, 
7. Be, Ville ts 

1854. A. Murchisone (Sow.), Morris, Catalogue, p. 293. 

1856. A. Murchisone (Sow.), Oppel, p. 368, no. 18. 

1867. A. Murchisone (Sow.), Waagen, Gevogn. Pal. Beitrage, p.(92) 
598, § 31. 

1878. Ludwigia Murchisone (Sow.), Bayle, Carte. Géolog. de la 
France, pl. 85. 


604. S. S. BUCKMAN ON AMMONITES FROM 


Localities. This species marks a distinct zone, which is just on the 
top of the sands or passage-beds. This zone is about a foot thick at 
Bradford Abbas, but about 3—4 feet at Corton and Hawthorn Downs. 
Harpoceras Murchisone occurs at Bradford Abbas, Marston Road, 
Corton and Hawthorn Downs, and other places, and also rather 
plentifully, but badly preserved, at Haselbury. 

There is a variety(?) which I am inclined to think should be 
separated from Harp. Murchisone. It has a smallish umbilicus and 
very fine ribs, and is far thinner. I have given it the name of 
Harpoceras Murchisone, var. bradfordiense, 8.8. B. 

Dimensions of a medium-sized specimen of Harp. Murchisone :— 
Diameter 2°85 inches, umbilicus 0°96, breadth of aperture 0°70 ; 
outer whorl, back to front, 1:20 inch. 


HARPOcERAS CycLorpus (d’Orb.). 


1845. Ammonites cycloides. @Orb. Pal. Frang. Terr. Jurass. pl. 121. 
figs. 1-6. 

1856. A. cycloides (d’Orb.), Oppel, Juraformation, p. 370. 

1867. A. cycloides (d’Orb.), Waagen, Geogn. Pal. Beitriige , Pp. (92) 
598. 


Localities. Humphriesianum zone at Oborne and quarries near 
Sherborne, somewhat plentiful; also at Wyke Quarry, but scarce. 

A variable species, some specimens being very thick, with a small 
umbilicus and coarse ribs, and others thin, with larger umbilicus and 
smaller ribs. The two varieties, however, merge one into the other. 

D’Orbigny has well represented the two extreme forms. 

This species was formerly quoted by the name of Amm. Cado- 
mensis, on account of the misprint underneath d’Orbigny’s plate. 


Harpoceras Levesquai (d’Orb.)? 


Ammonites Levesque, @Orb. Terr. Jurass. pl. 60 (misprinted 
solaris, Phillips). 


I quote this species with some hesitation, as our specimens seem 
to have fewer whorls and somewhat more marked ribs. 

Our specimens are from the zone of Harpoceras Sowerbyi at Brad- 
ford Abbas, Halfway House, &c. ; not very common. 


Harpoceras Epovarpranum (d’Orb.) ? 


1845. Ammonites Edouardianus, d’Orb. Terr. Jurass. pl. 180. figs. 
3-5. 

1856. A. Hdouardianus (d’Orb.), Oppel, Juraformation, p. 370. 

1881. A. Edouardianus (d’Orb.), J. Buckman, Quart. Journ. Geol. 
Soc. vol. xxxvii. p. 61, no. 3. 


Our specimens do not seem to agree very well with d’Orbigny’s 
figures, and are perhaps not the same. They have a more open 
umbilicus and far less inclusion; consequently the whorls are less 
QCross. 

Locality. Humphriesianum zone at Oborne: scarce. 


THE INFERIOR OOLITE OF DORSET. 605 


HARPOCERAS DISPANSUM (Lycett). 


My authority for this name is a specimen sent by Dr. Lycett, 
labelled “* Am. dispansus, Lyc., Frocester Hill.” 

I have one specimen from near Crewkerne which agrees very well 
with it. 


Harpoceras Mooret (Lycett). 


Ammonites Moorer, Lyc. Cotteswold Hills, pl. 1. fig. 2. 

A, Moorei (Lyc.), J. Buckman, Quart. Journ. Geol. Soc. vol. xxxvil. 
p- 65, fig. 7. 

Localities. Found in layers of comminuted shells in the sands at 
Bradford Abbas, Yeovil Junction, &c.: not very common. 

Termination. A slight double curve produced on the ventral area. 


Harpoceras cornu, 8. 8. Buckman, n. sp. 


Ammomites subradiatus, J. Buckman (non Sowerby), “Terminations 
of Ammonites,” Quart. Journ. Geol. Soc. vol. xxxvii. p. 61, fig. 2. 

This species is compressed, with strongly marked ribs, which are 
slightly reflexed and not so prominent close to the termination. 
They seldom bifurcate; but now and then a rib comes in between 
which is not continued all across. 

Keel distinct, with very sloping sides. 

Aperture sagittate. 

The umbilicus varies somewhat in width; it is, however, rather 
wide, with strongly marked ribs, showing also a certain portion of 
each whorl. 

Shoulder concave. 

Termination. Like that of Harpoceras concavum (Sow.); but it has 
a very long horn on each side. 

Difference. This species resembles Harp. concavum, but is distinct 
on account of the horn which the termination possesses, and which I 
I have never seen in that species. The ribs are also far more 
marked, and the umbilicus is far wider. 

This species never attains the same size as H. concavum; but very 
small specimens of these two species are hard to separate. 

This species might possibly be mistaken for the young of Harpo- 
ceras Murchisone (Sow.); it, however, never attains the size of 
H. Murchisone, is found at a higher level, has a more prominent 
keel, is far thinner towards the keel, is more acute, and its ribs are 
less prominent and rather more rounded. 

Dimensions. Diameter 2°50 inches, umbilicus 0:60, breadth of 
aperture 0°48, outer whorl from back to front 1°15. Another spe- 
cimen measured—diameter 2°70 inches, umbilicus 0°82, breadth of 
aperture 0°50, outer whorl from back to front 1:06, length of 
_horns 0°65. 

IT have never seen a specimen of this species which exceeded the 
diameter of about 22 inches. 

Compare these measurements with those of a medium-sized Harp. 
Murchisone (Sow.). 


606 S. S. BUCKMAN ON AMMONITES FROM 


Remarks. This species was figured in the ‘Quarterly Journal of 
the Geological Society ’ for February 1881, under the name of Am- 
monites subradiatus ; but it is very distinct from that species, the 
umbilicus alone being sufficient to distinguish it. 

Localities. It is found in the zone of Harpoceras Sowerbyi at 
Bradford Abbas, Halfway House, and many other places. It is 
tolerably common. 


AMALTHEUS SUBSPINATUS, S. S. Buckman, n. sp. 


This species is so peculiar and distinct from all others which I 
have met with in the Inferior Oolite, that I think it worth while to 
mention it here, although it has not been figured. 

It is very nearly allied to Amaltheus spinatus (Bruguiere) of the 
Marlstone. It possesses the peculiar crenulated keel, which is dis- 
tinct; but the crenulations are not very prominent. It has a large 
number of whorls, and very small inclusion, but does not increase in 
breadth at all rapidly. It has many fine angular ribs, ornamented 
With two spines on each rib, one set being in much the same place 
as those on Amaltheus spinatus, while the other set is on the inner 
portion of the whorl. 

Aperture. Quadrangular. 

Dimensions. Diameter 3 inches, umbilicus 1°55, aperture from 
back to front 0-75, across 0°67, inclusion barely any. 

Localities. It occurs in the Sowerbyt zone at Bradford Abbas, 
Halfway House &c., but is scarce. 

The termination is like that represented by d’Orbigny(Terr. Jurass. 
pl. 52. fig. 1) for Amaltheus spinatus—a plain single bend, much 
produced on the ventral area. 


OppEtia T'RUELLI (d’Orb.). 


1845. Ammonites Truellii, dOrb. Terr. Jurass. pls. 117, 129. 

figs. 1, 2. 
1849. A. Truellii (V’Orb.), Quenstedt, Jura, tab. 54. 

1854. A. Truellic (d’Orb.), Morris, Catalogue, p. 295. 

1878. Oppelia Truellai (d’Orb.), Bayle, Géologie de la France, 
vol. iv. pl. 89. figs. 1, 3, 4 (2?, 52). 

Occurs in the zone of Cosmoceras Parkinson at Halfway House, 
Wyke Quarry, and Burton Bradstock. 

The longitudinal lines well distinguish this species. It is also 
ornamented with longitudinal furrows, which vary slightly in depth, 
as also does the species in thickness. 


QpPELIA SUBRADIATA (Sow.). 


1825. Ammonites subradiatus, Sow. Min. Conch. pl. 421. fig. 2. 

1845. A. subradiatus (Sow.)?, d’Orb. Terr. Juras. pl. 129. fig. 3. 

1854. A. subradiatus (Sow.), Morris, Catalogue, p. 295. 

1856. A. subradiatus (Sow.), Oppel, Juraformation, p. 372, fig. 26. 

Localities. Zone of Cosmoceras Parkinsont at Broad Windsor, 
Crewkerne Station, Bridport, &c.: rather common. 


THE INFERIOR OOLITE OF DORSET. 607 


Termination from the inner whorl nearly straight, then a rather 
fine horn, then a curve back, and produced on the ventral area. 


OppELia suscostata (J. Buckman). 

1881. Ammonites subcostatus, J. Buckman, Quart. Journ. Geol. 
Soc. vol. xxxyli. p. 63, no. 8. 

1845. A. subradiatus, d’Orbigny (non Sowerby), Terr. Juras. 
pl. 118. figs. 1, 2. 

This species was figured by d’Orbigny as the large form of Oppelia 
subradiata (Sow.); but I am convinced that it is distinct. It has a 
far larger umbilicus, coarser ribs, hardly a distinct keel, and is far 
thicker. 

Localities. Zone of Stephan. Humphriesianum at Oborne &e.: 
rather scarce. 


HapiocERAs oonitHicum (d’Orb.). 

1845. Ammonites oolithicus, d’Orb. Terr. Jurass. tab. 126. 
figs. 1-4. 

1854. A. oolithicus (d’Orb.), Morris, Catalogue, p. 294. 

1856. A. oolithicus (d’Orb.), Oppel, Juraformation, p. 573, no. 32. 

Localities. It occurs in the Humphriesianum zone at Oborne and 
Milborne Wick. Our specimens have a slightly smaller umbilicus 
than those figured by d’Orbigny, but are exactly similar in all other 
respects. 


AMMONITES cADOMENSIS, Defrance. 


1845. Ammonites cadomensis (Defrance), dOrb. Terr. Jurass. 
pl. 129. figs. 9-11. 

T am not certain to what genus to attribute this species. 

Localities. This species occurs in the zone of Stephan. Humphrie- 
sianum at Oborne. It israre. A few specimens possess the peculiar 
termination. I have one specimen with it from Wyke Quarry, and 
others from Oborne. 


It may be also interesting to add that we have found the following 
species of Ancyloceras &c. in the Inferior Oolite :— 


Ancyloceras annulatum, d’Orb. Terr. Jurass. pl. 225. figs. 1-7. 
Humphriesianum zone at Oborne. 


Anc. subannulatum, d Orb. Terr. Jurass. pl. 225. figs. 12-15. 
Humphriesianum zone at Oborne. 


Ane. bifurcatum, Quenstedt: syn. Hamites bifurcatus, Quenstedt, 


Ceph. tab. 11. fig. 15. 
Humphriesianum zone at Oborne. 


Ane. bispinatum, VOrb.? Terr. Jurass. pl. 228. figs. 6-9. 
It much resembles this species, but is from the top of the Hum- 
phriesianum zone near Halfway House, and may be new. 


608 ON AMMONITES FROM THE INFERIOR OOLITE OF DORSET. 


Toxoceras Orbignyt (Baugier and Sauzé), d’Orb. Terr. Jurass. 
pl. 232. figs. 1, 2. 
From near Sherborne. 


Since the above paper was written, we have made out the Yeovil 
sands more clearly from some fresh sections. Just below the Mur- 
chisone zone (the equivalent of the Gloucestershire Pea-grit) come 
about 30 feet of sands. Then comes a marly bed about a foot thick, 
containing Harpoceras opalinum, Lytoceras torulusum, Waldheimia 
anglica, and Rhynchonella cynocephala. Below this come about 
100 feet, sometimes more, of sands in which Lytoceras jurense (Zieten) 
occurs along with Harpoceras Moore: (Lyc.) &e. These lower sands 
therefore belong to the zone of ZL. jurense. 

On the vexed question as to whether these latter belong to the 
Lias or Oolite I have not formed an opinion. 


Discusston. 


The Prestpent bore testimony to the great industry and skill 
shown by the young author in collecting from the richly fossiliferous 
beds of the Inferior Oolite of Dorsetshire. He pointed out the 
importance of having sections of perfect specimens of Ammonites 
made, so as to show the relation of the body-chamber to the others 
in these shells. 

Mr. Huptestron referred to the difference between the views of 
Mr. 8. Buckman and his father, Prof. Buckman, as to the value of 
the zones of the Inferior Oolite and the geological age of the Yeovil 
Sands. 

Mr. CuarteswortH remarked on the absence of phragmocones of 
Belemnites in Chalk rocks, and of the opercula of Ammonites from 
many Jurassic rocks. 

Prof. Srprey denied the universal absence of phragmocones of 
Belemnites in the Chalk, and stated that it might be accounted for 
by the condition of preservation of the fossils. In the same way the 
absence of aptychi might be accounted for by the general explana- 
tion of such facts given by Mr. Sorby, who showed that fossils 
composed of the unstable substance aragonite were rarely preserved. 

Prof. Brake thought that many of the so-called species of Ammo- 
nites would prove to be only varieties of well-known ones. He 
confirmed the views of the President as to the value of determining 
the proportion of the body-whorl. 

The PrestpEent stated that Mr. Buckman had adopted the views 
of classification which he had himself long ago insisted on. He 
accounted for the frequent absence of aptychi by the fact that the 
body-chamber is not usually broken open. He also agreed with Mr. 
Charlesworth as to the general but not universal absence of the 
phragmocone in Belemnitella, and with Mr. Blake that many of 
these Amamonite-forms could not be regarded as distinct species. 


E. J. DUNN ON THE SOUTH-AFRICAN DIAMOND-FIELDS. 609 


40. Nores on the Diamonv-FIELDs, Sour Arrica, 1880. By EH. J. 
Duyn, Esq. (Communicated by Prof. Ramsay, F.R.S., F.G.8.) 
(Read June 22, 1881.) 


THE mining-operations carried on during the last few years at the 
diamond-fields, South Africa, have brought to light some additional 
- facts bearing on the formation of diamonds*; the most interesting 
is the exposure, at all the old mines (Kimberley, De Beer’s, Du Toit’s 
Pan, and Bultfontein), of considerable deposits of black carbonaceous 
shale underlying the surface beds of grey shale. 

By the removal of the diamond-bearing ground of the old vol- 
canic “‘ pipes” constituting the above mines, the wall or rim is left 
unsupported ; after rain immense masses of this shale, or ‘“ reef,” 
as miners term it, fall into the excavated gulf, leaving excellent 
clean sections of the horizontal strata; for the bedding of the shales 
is horizontal, except where locally disturbed by intrusive rocks, or 
at the sides of the “pipes,” where they are turned upwards for a 
few feet, perhaps by the gabbro in the “ pipes.” 

At Kimberley mine the surface-shales, grey or, in places, pink or 
yellow, which contain remains of small Saurians, are from 40 to 50 
feet thick; underneath are black carbonaceous shales, for the most 
part arenaceous, and more than 100 feet thick. So combustible 
are these shales that in a part of the mine where they were acciden- 
tally fired they have smouldered on for more than eighteen months 
and are still alight. 

Time did not admit of a search for plant-remains; but a diligent 
search, especially in the finer and more argillaceous beds, would 
be almost certain of success. Thin seams of very impure coal full of 
pyrites occur in the black shales ; and here and there a long flattened 
piece of pure coal is found, probably the stem of some plant altered 
to coal and flattened by compression. 

At De Beer’s mine, on the north side, a somewhat different sec- 
tion is laid bare. First there is, from the surface down, about 50 feet 
of dolerite, then about 12 feet of yellow thinly laminated shales ; 
beneath these are the black carbonaceous shales, corresponding with 
those in Kimberley mine, and also containing thin seams, up to 
1 inch in thickness, of impure coal. 

The precise depth of these carbonaceous shales has yet to be 
determined; but that they extend horizontally over the whole 
country at no great depth below the surface there is no reason to 
doubt; for wherever wells have been sunk in the neighbourhood of 
Kimberley, De Beer's, Du Toit’s Pan, or Bultfontein, these black 
shales have been encountered at depths varying from 40 to 60 feet 
from the surface. These shales are to be traced cropping out on the 
banks of the Modder river}, some forty miles from Kimberley, and 


* [See E. J. Dunn’s previous papers, Quart. Journ. Geol. Soc. vol. xxx. 
p. 54; and vol. xxxili. p. 879. ] 
+ [Quart. Journ. Geol. Soe. vol. xxx. p. 582.] 


Q.J.G.S. No. 148. 25s 


610 E. J. DUNN ON THE SOUTH-AFRICAN DIAMOND-FIELDS. 


again on the Riet river, still further away. How much further they 
extend can only be determined by boring ; for the level nature of the 
country prevents one from obtaining any knowledge of the beds 
100 feet below the general level. 

That the old mines are volcanic “ pipes,” and that they have 
burst through these carbonaceous shales is evident. Is it not reason- 
able to infer that the carbon, indispensable in one form or another to 
the formation of diamond, was supplied by these shales ? 

It is well known that at and near the surface, or while the work- ~ 
ings in the “pipes” were bounded by grey shale, the mines were 
not so productive as at lower depths ; when the zone of black shales 
was reached the diamonds were more plentiful and also of better 
quality. 

At Kimberley the improved yield as depth was attained was well 
recognized ; and at Bultfontein mine so notably is this the case that, 
though the surface-ground scarcely paid for working, the yield at a 
depth of from 60 to 80 feet is most satisfactory. Jagersfontein, in 
the Free State, again, is an instance of a mine poor at the surface, 
but very profitable to work lower down. 

So far as experience can be drawn on as a guide, it appears that 
the yields of diamonds in these “pipes” are greatest when mining 
is carried on in the portions of the “pipes” surrounded by carbo- 
naceous shales, rendering it probable that these shales suppled the 
element necessary to the formation of diamond. 

A practical question of serious importance as regards the diamond- 
mining industry here suggests itself: if the black shales supplied 
the carbon of which the diamonds are formed, it is to be expected 
that some diamonds would be found higher in the “ pipes” than 
the black shale ; for the tendency of the molten rock would be to 
rise, and this would also be the case with carbon in the state of 
vapour; but is it probable that diamonds will be met with below 
these shales ? 

Already in three separate localities shafts sunk at the edge of 
Kimberley mine have, at a depth of 300 feet, struck remarkable 
intrusive rocks—amygdaloids, breccias, &c. that differ essentially 
from any thing found penetrating the surface-shales. Whether these 
are older intrusive rocks on which the shales were laid down, or 
whether they are later than the shales and penetrate them is not yet 
decided. ; 

That the “pipes” will continue down for a vast depth there can 
be no doubt; but if beneath the black shales the “pipes” have 
traversed rocks devoid of carbon, are diamonds still to be expected 
in them ? 

Koffyfontein mine, on the road between Kimberley and Jagers- 
fontein mines, has so far not proved rich at the surface. Many other 
undoubted “pipes” have been opened and prospected near Kimber- 
ley ; but subsequently they have been abandoned as unremunerative, 
although some diamonds were obtained. It is very probable that 
by deeper sinking in these localities richer yields would be found, 
as the black shales are almost certainly below. 


E. J. DUNN ON THE SOUTH-AFRICAN DIAMOND-FIELDS. 611 


Should it prove to be the case that these carbonaceous shales pro- 
vided the carbon for the diamonds, the conclusion is forced on us 
that the original source of the diamond is the atmosphere ; for the 
plants absorbed carbonic acid gas from the air, and in course of 
time were entombed, and thus provided a store of carbonaceous 
matter in the shales. Later on these shales were shattered and 
engulfed in the molten rock. The carbon was then liberated in the 
state of vapour by the intense heat; but, being under great pressure 
in the ‘ pipes,” instead of escaping, it crystallized out as sparkling 
diamonds. 

The disclosure of such extensive deposits of carbonaceous shale 
has other bearings quite as important as on the formation of dia- 
monds. Hitherto nosuch deposits were known to exist in the Karroo 
beds or Dicynodon series of rocks that cover such an immense area 
in Cape Colony and the Free State. Vast plains broken by hills, 
either isolated in groups or in long ranges, characterize the country 
occupied by these rocks; the river-beds are generally shallow; and 
thus it happens that until the southern limit of the Karroo beds is 
reached, which is 200 miles direct from Kimberley, no section, even 
to a moderate depth below the general level is obtainable. 

‘At their southern termination these horizontal beds abruptly 
cease, exposing their edges through a thickness of more than 3000 
feet, and form the Nieuwveldt, Camdeboo, and Winterberg ranges 
of mountains. 

No seam of coal has been found along these well-exposed edges, 
although well searched for. Very insignificant thin seams of black 
shale do exist, also occasional plant-remains in the form of Glosso- 
pteris and Hqwisetum, but no such shales as at the diamond-fields. 
The remarkably pure anthracite found vertically intersecting the 
rocks (Karoo beds) at Butfel’s Kloof, Camdeboo* has been proved 
by boring to be the result of distillation, perhaps ensuing from the 
action of intrusive rocks on the vegetable remains included in the 
shales. At the above locality a large dyke underlies the outcrop 
of anthracite, though at the surface it is several hundred yards 
distant. 

The trough of the great basin occupied by the Karrooy beds lies 
south of the Orange river, and about east and west. Is there not a 
possibility that these carbonaceous shales cropping out near the 
northern limit of the Karroo beds may develop into true coal-seams 
further south, or nearer the centre of the basin? A couple of bore- 
holes would settle the point; and the pressing need for fuel at the 
diamond-fields will ensure the trial being made. 

In previous notes some dyke-like masses at De Beer’s mine were 
described as noticeable near the surface; now that the workings 
have reached a depth of over 100 feet these dykes are less equivo- 
eal, as the rock forming them appears in almost its original con- 
dition, and not decomposed as at the surface; it is of dark bluish- 

* [See E. J. Dunn’s Report to the Colonial Parliament, 1879.] 


+ |This name, as used by Mr. Dunn, excludes the Uppermost Karroo or Storm- 
berg beds. | 
28 2 


612 E. J. DUNN ON THE SOUTH-AFRICAN DIAMOND-FIELDS. 


grey colour and somewhat erystalline texture. It appears to differ 
from the general mass filling the “ pipe” only in being fine-grained 
and less liable to decompose. These dykes arefrom 2 to 3 feet thick ; 
they cut through the gabbro filling the ‘‘ pipe,” and the carbonaceous — 
shales, grey shales, and dolerite that form the sides of the ‘‘ pipe.” 
No similar dykes were observed in any of the other mines. 

At Bultfontein mine the shales on the east side are much disturbed, 
and are tilted in places at an angle of 60° away from the ‘“ pipe.” 
There are numerous blocks and small pieces of dolerite in this 
mine that have been so rounded by attrition against one another, 
probably caused by the heaving of the mass when in a molten state, 
that they resemble boulders and pebbles. These stones are not so 
noticeablein any of the other mines as they are at Bultfontein. 

As an instance of the value of the ground filling these “ pipes,” 
the following (on good authority) is of interest :— 

At Du Toit’s Pan 7000 loads, 16 cubic feet each, yielded, on an 
average, diamonds to the value of £2 12s. per load. 


Discussion 


Prof. Ramsay said the facts mentioned were remarkable; and 1% 
was extremely difficult to say what the circumstances were under 
which diamonds were developed. | 

Mr. J. Evans said that the author had in this paper gone further 
than he had done in his previous communication. ese year small 
diamonds had been shown at the Royal Society made, it was said, 
artificially. It would be an experiment worth while for Mr. Hannay 
to repeat, in the form of heating together pieces of carbonaceous 
shale and of fusible igneous rock. 

Prof. Srptry said the view of the author was a plausible one. He 
himself had suggested that carbonic acid might have been carried 
down by water, and then decomposed by the heat of the volcanos, 
so that the carbon, when liberated, might become crystallized. The 
general principle of the author’s theory might be true, though, 
perhaps, not the precise application of it. 


G. R. VINE ON STOMATOPORA AND ASCODICTYA. 613 


41. Srvrran Untsrr1at Sromatoporz and Ascopicrya. By Grorcr 
Rosert Vine, Esq. (Communicated by Professor P. Marri 
Duncan, F.R.S., F.G.8.) (Read June 22, 1881.) 


Tue genus Alecto was founded by Lamouroux in 1821 for a group 
of adherent Polyzoa. In 1814 Leach had used the word Alecto for 
a genus of Hchinoderms ; and Mr. Hincks says that it is still employed 
in connexion with the Crinoidea. On this account its further use 
for species of Polyzoa is objectionable. In 1825 Prof. Bronn used 
the word Stomatopora, and in 1826 Goldfuss used Aulopora, as 
names for individuals of the same genus as that founded by Lamou- 
roux. For uniserial species d’Orbigny employed Prof. Bronn’s 
name; but Blainville, Johnston, Milne-Edwards, Busk, and Defrance 
used the original word “ Alecto” for species described by them in 
their various writings. 

The generic characters of Stomatopora have been given by various 
authors ; and additions have been made from time to time. ‘The 
rather full description given by Goldfuss* of Awlopora dichotoma, 
together with figures of the species, renders identification compara- 
tively easy. But somehow there has been a confusion in later 
identifications, and the Awlopora intermedia t type of Minster has 
been mixed up with Goldfuss’s type. Both of these are present in 
the Jurassic formation; and it is, I will admit, rather a difficult 
matter to say where the one ends and the other begins. If, how- 
ever, authors would distinguish between the two types, we should 
be able to get at the true range of the species, because each has an 
individual facies of its own. Hall, in describing the species found 
in the Trenton Limestone of Americat (Trenton Falls, Oneida 
county), makes this distinction:—In <Alecto inflata we have the 
tubes short and individually separate; whilst in Aulopera arachno- 
adea the tubes are not distinct or separated from the general con- 
sistence of the branch. Jules Haime, in his descriptions of the 
fossil Bryozoa of the Jurassic rocks, places the whole of his species 
under one genus; and I prefer this method rather than object to it. 
Thus, Stomatopora antiqua from the Inferior Lias of Valicre, S. 
Terqueni from the Inferior Oolite, and S. Bouchard: from the Ox- 
ford Clay are of the Aulopora intermedia type; and this holds good 
with species found in our own country. Stomatopora dichotoma, 
S. dichotomoides, D’Orb., and S. Waltoni are of the same type as 
that given by Goldfuss as Awlopora dichotoma. In S. Desondona, 
Haime, from the Inferior Oolite of Longwy, we have a passage- 
form between Aulopora intermedia and the genus Proboscina, and 
then species of Proboscena passing by gradations, with a tendency 
on the one hand to the [dmonew, and on the other to the larger 


* Petrefacta Germania, p. 218, pl. 65. f. 2. 
+ Ibid. p. 218, pl. 65. £1. 
{ Palzontology of New York, vol. i. p. 77. 


614 G. R. VINE ON SILURIAN UNISERIAL 


Stomatopore. There are in all the species and genera named indi- 
vidual characters, if isolated, that would indicate affinities in the 
whole group; and Mr. Hincks is wise in grouping all these genera 
under one family name, that of the Tubuliporide. 

Prof. H. Alleyne Nicholson, in working out material snbmitted to 
him for examination by Mr. U. P. James, of Cincinnati, and also 
material collected by himself, saw fit to rename the species of 
Hall Hippothoa inflata, and, according to his description, shifted 
the species from the suborder Cyclostomata to that of the Cheilo- 
stomata of Busk. I could not from the first agree with the Pro- 
fessor; but I was unable to dispute the point raised by him other- 
wise than by the mere expression of opinion ; for up to the present 
time no record has been given of species of Stomatopora found in 
our own Paleozoic rocks. I am now able to carry back the true 
uniserial Stomatopore to the Lower Wenlock Shales of Shrop- 
shire. 

It may be as well to say a few words about the material used by 
me for this and other papers (to follow) on Silurian Polyzoa. It is 
now pretty generally known that, for the purpose of assisting Mr. 
Thomas Davidson, F.R.S., in his labours on the Silurian Brachio- 
poda, Mr. George Maw, F.L.S. and F.G.8., of Benthall Hall, Shrop- 
shire*, has had washed and carefully picked, for Brachiopoda, about 
18 tons of Wenlock shales. The débris of these washings were after 
this laid aside for the use of other specialists. Some time since I 
applied to Mr. Davidson, and afterwards to Mr. Maw (Mr. Davidson 
supporting my request), for some of this refuse, for the purpose of 
working out stratigraphically the Polyzoa and smaller Actinozoa. 
My request being granted, Mr. Maw sent me on the 19th of March 
over two hundredweight of the débris for this purpose. I intend 
to use the whole of this material honestly ; for I feel convinced that 
it can be only by labours such as these that a true idea of the 
abundance of the Polyzoal life of former epochs can be obtained ; 
and, though picking out fragments from such a mass, by the aid of 
a hand-glass, may be both painful and tedious, I shall prefer to 
work on different groups, as material accumulates, rather than delay 
writing till the whole has been picked. I have already gone over 
about thirty pounds of the débris from the eleven localities and ho- 
rizons ; and it may be interesting, as showing the difference between 
the shales of the Carboniferous and the shales of the Wenlock series, 
when I say that a single pound of unwashed Hairmyres clay would 
yield me in the washing more individual specimens than I have been 
able to get from the thirty pounds of the Wenlock débris. In the 
Carboniferous the fragmentary organisms are tolerably well pre- 
served and perfect; in the other the Polyzoal remains seem to have 
been much waterworn, but, with the exception of one locality, not 
sufficiently injured to prevent identification. | 

We are indebted to Prof. Hall for the first indication of the ex- 
istence of uniserial Stomatopore in Silurian rocks. It is quite 


* See Geological Magazine, Jan., March, &. 1881. 


STOMATOPOR AND ASCODICTYA. 615 


possible that Lonsdale and other workers on the Silurian organisms 
may have had a previous knowledge of the fact of their existence in 
these rocks; but no detailed account was furnished. For the work- 
ing-out of these and other forms of Stomatopora we are equally in- 
debted to Prof. Nicholson, M.D., F.G.S., &e. 


Silurian Stomatopore. 
1. SroMATOPORA INFLATA. 


Alecto inflata, Hall, Paleont. of New York, vol. i. p. 77, pl. xxvi. 
figs. 7a, 6, = Hippothoa inflata, Nicholson, Ann. & Mag. Nat. Hist. 
Feb. 1875, pl. x1. figs. 1, la. 

“ Zoarium* attached, arachnoid; zowcia short, much expanded 
above, contracting at the aperture and narrowing rapidly below: 
orifice large, opening obliquely upwards.” 

This is Hall’s description of his species. Nicholson says that the 
branches of his specimens are linear, and the “cells uniserial and 
pyriform, each springing by a contracted base directly from the cell 
below ; about four cells in the space of one line.” There is nothing, 
however, in his description that would ally the species with the 
Hippothoe ; but in working out my own Upper-Silurian types I have 
given prominence to every feature that had any tendency to a Hippo- 
thooid character. 

The geological position of Hall’s species is the Trenton Limestone. 
Nicholson’s specimens are from the Hudson-River Formation, Cin- 
cinnati Group. 


2. StoMaToPoRA DissIMiLis, mihi. Figs. 1-8 (p. 616). 


Zoarium adnate, branching, generally attached to stems of Cri- 
noidea, very rarely to broken shells; branches lmear, sometimes 
wavy and anastomosing. Zoecia invariably uniserial, and, in the 
best preserved, very finely ribbed transversely ; the oral extremity 
slightly raised ; orifice circular or subcircular. Occial cells rather 
ventricose and strongly ribbed (?). Each normal zocecium about 
half a line; average about 6 to 34 lines. 

Loe. Upper Silurian; “ Buildwass beds,” Harley, near Wenlock, 
rather rare; also base of Wenlock shale, Buildwass Bridge, Shrop- 
shire, rather common. 

I have not found any specimens of this species in any other of 
the eleven localities which I have searched for Polyzoa. In search- 
ing the material from these two localities, I found it to my interest 
to examine on both sides every fragment of shell and Crimoid that 
came under my glass; the consequence of this is that I have speci- 
mens, more or less perfect, of about fifty colonies. The drawings 
are made from three of these, because they afforded me better facies 
than the others. What I have given are characteristic of the 
whole. 


* To present a uniformity in the descriptions, I change the exact words of 
authors to those in present use : thus, Hall's word “Coral” is changed to Zo- 
arium, “ Polyzoary” of authors to the same. 


G. R. VINE ON SILURIAN UNISERIAL 


Figs. 1-8.—Stomatopora dissimilis, Vine, from the Buildwass beds, 


Upper Stlurian, Shropshire. 


| 7 


jan 


| 


(cree / 


Yp 
I} 


Ups 
>" 
AW 

Te 
a 
Ce i 
IN 


1. Colony adherent to Crinoid stem. \ Hach of these colonies has been drawn 
| with the Camera lucida, from three 
5 es f separate specimens on Crinoid stems, 
the habit of each colonial growth be ing 
somewhat dissimilar in character. 


2. 99 99 
aed 


% 


99 


STOMATOPORZ AND ASCODICTYA. 617 


Fig. 4. Fig. 5. 


4. Profile of two cells from No. 3 colony, only lower down on specimen. 

5. Magnified cell on same colony (fig. 3, a). 

6. Three cells from same colony (fig. 3, 0). 

7. Cell from same colony, with caudate elongation (fig. 3, c). 

8. Cell from another colony. The stoloniferous processes on specimen No. 3 
do not belong to the Polyzoal colony, but to Ascodictyon. Upper Silurian, 
Buildwass beds, base of Wenlock shale, Shropshire. 


The habit of the colonial growth, as given in fig. 1, is similar 
to that of Aulopora dichotoma, Goldf. Generally speaking, about 
every second cell gives origin to a fresh one; and this is the be- 
ginning of a new branch. I cannot, however, give this as a cha- 
racter, on account of its variableness. The origin of fresh colonies 
of this beautiful species is a most interesting study. Without 
speaking positively on this point, I have in one small fragment 
probable evidence that clusters of cells are developed from one of the 
‘“‘ rosettes” otf Ascodictyon stellatwm, Nich. and Ether.* Around 
this cluster primary cells of various colonies are disposed ; some 
colonies have a linear arrangement of from three to five cells; the 
primary cells are also disposed singly on different parts of the frag- 
ment of broken shell; a larger mass of cells clustered in one spot 
give origin to several linear branches of what I am disposed to 
believe are new colonies. It may, however, be possible to explain 
this feature by stating that one rosette gives origin to several linear 
branches, and the whole clustered together would be the parental 
nucleus of one colony variously disposed. There is sufficient evi- 
dence to show that some colonies at least sprang from an inde- 
pendent primary cell. 

Amongst living Stomatopore a most remarkable feature is shown 
in the figures of S. fasciculata, Hincks, pl. lix. figs. 4,57. In his 
descriptive text (p. 441) Mr. Hincks separates this from all other 


* Ann. & Mag. Nat. Hist. June 1877. 
+ Hincks, Brit. Marine Polyzoa, vol. i1.; text, vol. i. 


618 G. R. VINE ON SILURIAN UNISERIAL 


known species, and places it in a division by itself (C. Colony 
clustered). In the figure there are several peculiar clusters of cells, 
including from two to five or seven cells. There are no stoloni- 
ferous processes. In comparing the feature given here by Hincks 
with that of Nicholson’s figure of Ascodictyon stellatum, and both of 
these with my own specimen, I cannot arrive at any other con- 
clusion than that some of the Ascodictya of the Paleozoic rocks 
are in some way homologous with the cluster found upon Hincks’s 
unique and solitary specimen of S. fasciculata. 

The profile of two cells, fig. 4, shows the true Stomatoporous de- 
velopment. There are some cells in fig. 3 that are of a most pecu- 
liar character. They differ in a few particulars from other cells ; 
and these I have ventured to suggest may be the ocecia of the 
colonies. I may, or I may not, be right in my conjecture on this 
point. Unless these be ocecia, I have not been able to trace in any 
other cells the least indication of ovarian chambers. 5 is a good 
illustrative example of the cell referred to. Other points of struc- 
ture are alluded to in the description of the figures. 

In the Annals and Magazine of Nat. History for June 1877, Messrs. 
Nicholson and Etheridge, Jun., described and figured a most peculiar 
and ‘‘anomalous genus of Paleozoic fossils.” The name given to 
the group was Ascodictyon; and several species were described as 
found in Devonian and Carboniferous rocks. The systematic po- 
sition and affinities of the fossils were not established by the authors 
when the paper was written. From material in my own cabinet I 
ventured to suggest, in a letter to Prof. Nicholson, what, judging | 
from the Carboniferous fossils, I believed to be the probable affini- 
ties. JI have now discovered in the Silurian shales of Shropshire 
several specimens of the species given by Nicholson; and so eare- 
fully are the characters of the Devonian fossils made out, that I 
can trace in the Silurian specimens a most remarkable resemblance. 
In the stellate rosette and stoloniferous processes there are differ- 
ences so slight that I was inclined to place my own fossils under the 
same generic and specific names, distinguishing one only with a 
varietal term *. 


3. ASCODICTYON sTELLATUM, Nich. & Eth., Jun. 


I have only two specimens of this type. There are a few differ- 
ences, which it may be well to indicate by giving it the varietal 
name, siluriense, mihi. 

Colony composed of calcareous clusters of ovoid cells, having a 
somewhat stellate character ; each cluster containing from four to 
seven cells, which are connected together by creeping filamentous 
cords, some of which anastomose at intervals. 


Loc. Buildwass beds, near base of Wenlock Shale, Shropshire. 


* Since this was written, I have been able to work out fuller details of this 
most remarkable group ; and I may add that Professor Nicholson has furnished 
me with specimens of his so-called Hippothoa inflata, particulars of which will 
be given in a future paper on the Polyzoa of the Wenlock Shales. 


STOMATOPORZH AND ASCODICTYA. 619 


4, Ascopicryon RADIANS ?, Nich. & Ether. (Provisional placement.) 


I have several colonies of this beautiful type apparently similar 
to those found in the Carboniferous rocks of Scotland. The colonies 
are not so prolific, however, in the Silurian as they are represented 
to be in the Kast Kilbride district. In the Silurian the clusters 
rarely exceed two or three ; in many cases there is only one stellate 
group of “elongated vesicles.” For the present I merely record 
their discovery, reserving more detailed description for some future 
time when my material is better worked. 

Loc. Buildwass beds, near base of Wenlock Shale, Shropshire. 

Hab. On stems of Crinoidea and fragments of shell. 

Without committing myself to any systematic classification (other 
than that suggested in the text) of these peculiar fossils, I think it 
would be unwise and ungenerous on my part to conclude this paper 
without speaking most approvingly of the labours of Prof. Nicholson 
and Mr. Robert Etheridge, Jun., in the same direction as my own. 
Prof. Nicholson remarks, in the paper on Ascodictyon™, that this 
“‘ genus, so far as our present knowledge goes, is confined to the 
Devonian and Carboniferous periods.” I am now able to extend 
its range. 


Ascopictyon, Nich. & Ether. Jun. 


Upper Silurian...... A. stellatum, var. siluriense, mihi, ............ Shropshire. 
Middle Devonian... A. stel/atwm, Nich. & Eth. ..................... Hamilton, Ontario. 
Fs cd PAL USUTOTIME, | ., Ss NS ee 7": 55 
Upper Silurian...... A. radians?,  ,, »» my own cabinet.. Shropshire. 

Carb. Limestone ... A. radians, Ps 3 ‘ s Scotland. 
33 32 A. stellatum 99 9? 29 De) 29 


SroMaTopora, Bronn, uniserial species. 


Lower Silurian...... IS) SIGE: ol 1 AR Ree oe Trenton Limestone, 

America. 
_ ay Oleoere S. inflata, (Hippothoa inflata, Nich.) ......... Hudson-River For- 

mation. 

Upper Silurian...... S. disstemilis, Vine; my own cabinet ......... Buildwass_ beds, 
Shropshire. 

12/:7170 1710 eee S. Voigtiana, King. Humbleton, York- 

shire. 
Discussion. 


The Presrpent stated that very important results were being ob- 
tained from these washings of Mr. Maw’s of Upper Silurian rock. 
Some of those obtained by Mr. Davidson were of the highest value. 
Aulopora had been made a receptacle for very various forms. 

Prof. Duncan said that the value of Mr. Vine’s researches was. 
very great. The numbers of Polyzoa produced were very great; and 
some of the Heteropore were singularly recent in aspect. 


* Ann. & Mag. Nat. Hist. June 1877. 


620 PROF. H. G. SEELEY ON THE REPTILE 


42. The Reprite Fauna of the Gosav FORMATION preserved in the 
Gurotogican Musrum of the University of Vinnna. By Prof. 
H. G. Susruy, F.R.S., F.G.S., &c., Professor of Geography in 
King’s College, London. With a Nore on the Guotoetcan 
Horrzon of the Fosstts at Nrvz WEtt, west of WrENER Nuvstapt, 
by Enpw. Svzss, Ph.D., F.M.G.S., &c., Professor of Geology in 
the University of Vienna, &c. (Read June 8, 1381.) 


[Puates XXVII.—XXXT.] 


ConrTENTS. 
INTRODUCTION. 


Historical review ; condition of specimens. 
Dinosavrtia. 


Dentary bone and teeth of Mochlodon Suessvi (Bunzel). 
Skull of Struthiosaurus austriacus, Bunzel; with a note on the base of the 
skull of Acanthopholis horridus, Huxley. 
On the genus Cratzomus, an armoured type with powerful fore limbs. 
Mandibles and teeth, probably referable to Cratgomus. 
Crateomus Pawlowiischti, Seeley, vertebral column, ribs, dermal armour, 
scapula, humerus, femur, tibia. 
Crateomus lepidophorus, Seeley, coracoid, scapula, humerus, femur, tibia, 
fibula, metatarsal bone, claw-phalange, dorsal vertebra. 
Tooth of Megalosaurus pannoniensis, Seeley. 
Femur of Ornithomerus gracilis, Seeley. 
Lower jaw and maxillary bone of Doratodon carcharidens (Binzel). 
Femur and humerus of Rhadinosaurus alcimus, Seeley. 
Scapula, humerus, and femur of Oligosaurus adelus, Seeley. 
Humerus, scapula, vertebrae, and armour of Hoplosaurus ischyrus (Seeley). 
CrocopiLia. 
Vertebral column, femur, fibula, ulna, radius, &c. of Crocodilus proavus. 
CHELONIA. 
Costal plates, postfrontal bones, and scapula of Plewropeltus Suessii, Seeley. 
Costal plates and plastron of Hmys Neumayri, Seeley. 
LAcervinia. 
Vertebra of Argosaurus gracilis, Seeley. 


ORNITHOSAURIA. 
Ornithocheirus Binzeli, Seeley. 


INTRODUCTION. 
Historical Review. 


Tur Gosau formation, nearly corresponding in age to the Upper Green- 
sand of this country, is represented at Neue Welt, near Wiener Neu- 
stadt, by freshwater deposits full of such freshwater shells as Melania 
and Unio, and land-plants such as Banksia and Pecopteris. The for- 
mation and its fauna have been described by Profs. Suess, Zittel, and 
many others ; but, although the late Dr. Stoliczka detected a tooth 
imbedded in the coal of the formation, no important knowledge was 
obtained of the vertebrate fauna of the Gosau beds until Prof. Suess 


—o a 


REMAINE 


C Berjeau lith | GOSAU REPTILIAN 


, 

) 
4 

a 
f 
} i 
| ’ 
; 


C.Berjeau tith 


REPTILIAN RE 


FAUNA OF THE GOSAU FORMATION, 621 


was so fortunate as to obtain the assistance of Bergverwalter Paw- 
lowitsch in conducting excavations. These were carried on with 
admirable skill; timber drift-ways were driven into the rocks, with 
the result that they penetrated into a perfect cemetery of the remains 
of Cretaceous reptiles. The remarkable collection thus obtained 
was intrusted for description to Dr. Emanuel Bunzel, whose memoir 
upon it was published in 1871 in the ‘ Transactions of the Imperial- 
Royal Geological Institution.’ Subsequently more specimens were 
discovered ; and in Easter 1879 my honoured friend, Prof. Suess, 
invited me to visit Vienna to examine these specimens, with the 
object of making them available for the advancement of knowledge 
by publication. With the assistance of the Royal Society I gladly 
undertook this work, and spent a month in Vienna studying the 
thousands of fragments which had been obtained. The great mass 
of these, mere comminuted bones, proved of but little value; or, 
rather, the time that I could give to their study enabled me to piece 
together but few specimens that were likely to prove interesting. 
There were, however, other important remains, which Prof. Suess 
had already reconstructed and pieced together with great patience 
and perseverance, that had produced many indications of lost animal 
forms out of a chaos of débris. I soon found that Biinzel’s 
views and my own presented certain differences. His memoir, 
which extends to eighteen quarto pages, and is illustrated by eight 
plates, describes the following species—Crocodilus carcharidens, 
Iguanodon Suess, Struthiosaurus austriacus, and Danubiosaurus 
anceps. Other remains are referred to the genera Hylcosaurus, 
Scelidosaurus, and Lacerta; while certain specimens are classed as 
‘“‘Crocodili ambigui,” Chelonians, and indeterminate remains. All the 
specimens which he described are figured; but the artist has so im- 
perfectly appreciated the details of character of the fossils represented 
in Bunzel’s plates, that it is impossible to form from them a just 
opinion of these fossil reptiles. After examining the specimens, I have 
come to the conclusion that some of Bunzel’s identifications may be 
modified. Jam unable to recognize Scelidosawrus, of which Bunzel 
figures a claw-phalange, tail-vertebre, and dermal armour. Hyl«o- 
saurus is another genus doubtfully cited, resting upon a single scute, 
which it may be well to discard. Lacerta is a genus that certainly 
_ cannot be recognized, although the author refers to it parietal and 

postfrontal bones, the articular element of the lower jaw, and the 
right side of the lower jaw, vertebra, fragments of ribs, humerus, 
radius, and femora. But the genus Lacerta could here only be used 
in the sense of animals of the Lizard group. 

For reasons that will be adduced, the Crocodilus carcharidens, 
founded upon a fragment of the lower jaw, cannot be referred to the 
genus Crocodilus; while the Danubiosaurus anceps was founded in 
error, and the remains, instead of being lacertilian, belong to other 
orders and other parts of the skeleton than those identified. Stru- 
thiosawrus being founded on a single specimen, remains an interesting 
type; but I feel constrained to refer the Iquanodon Suessir to a 
distinct genus. 


622 PROF. H. G. SEELEY ON THE REPTILE 


The vertebre, plate i. figs. 24-26, regarded as crocodilian, per- 
tain to a small Dinosaur ; figure 27 in the same plate, regarded as 
the dorsal rib of a Crocodile, I interpret as the cervical rib of a 
Dinosaur. The vertebra, regarded as crocodilian, which are figured 
in pl. 11. give evidence of a second and larger species of Dinosaur, 
and exemplify its cervical, caudal, and dorsal vertebre. 

On plate in. fig. 1, the specimen regarded as the right side of a 
hinder dorsal rib of a Teleosaur I regard as the shaft of the femur 
ot a new Dinosaurian genus. Figures 2—4, described as a crocodilian 
femur, is the femur of the larger Dinosaur. Figures 5, 6, called 
fragment of lower jaw of a Lizard, is certainly neither a fragment 
of jaw nor a Lizard-bone, but the proximal end of a large rib of a 
Dinosaur. Figures 12, 13, called the upper half of a crocodilian 
humerus, I regard as the proximal part of a Dinosaurian fibula. 

In plate iv. figs. 1, 2, classed as dermal bones of a Crocodile, I refer 
to one of the large new Dinosaurs. Figure 3, described as the right 
ium of Jguanodon Mantelli, is certainly a coracoid of a large Dino- 
saur. The tail-vertebre on the same plate, referred to Scelidosaurus, 
are caudals of the same Dinosaurian genus already referred to. The 
figures 11, 12, called phalange of crocodilian, is a Dinosaurian meta- 
carpal or metatarsal; and the claw-phalange (figs. 4, 5), referred to 
Scclidosaurus, probably belongs to the same animal. 

The figures of the remarkable skull of Struthiosaurus, represented 
on plate v., are all unsatisfactory, since they give but a vague idea 
of its structure. Figures 7-9, described as the rib of a lacertilian 
(Danubiosaurus anceps), represent the scapula of the larger Dinosaur. 
Figure 10, termed claw-phalange of Danubiosawrus, was shown by 
Prof. Suess to be a piece of Dinosaurian armour, since he fitted it to 
a remarkable horn-like scute of the larger new Dinosaur. 

On plate vi. figs. 1-3, 1s represented another example of the large 
Dinosaurian scapula, there interpreted as the rib of Danubiosaurus 
anceps. Figures 4,5 are said to represent the left ilium of this 
imaginary animal; but they are really the costal plate and blended 
rib of a large and remarkable new Chelonian type. Figures 8-10, 
termed bodies of vertebree of Lizard, are vertebree of the same species 
of Crocodile represented on plate i. Figures 6, 7, described as the 
articular part of the lower jaw of a Lizard, are really the articular 
end of the lower jaw of a Pterodactyle of the genus Orithocheirus. 
JT concur with the identification of fig. 11, as vertebra of a Lizard. The 
bone represented in figures 12, 13, termed dorsal rib of Lizard, is the 
fibula of a Crocodile. JI am unable to recognize satisfactory Lizard- 
characters either in the humerus figured in this plate or in any of the 
bones represented in plate vi., while that represented in figs. 22 and 
23, termed a rib, seems to me to be a femur of a new Dinosaurian 
genus. The Dinosaurian dermal armour in this plate, referred to 
Scelidosaurus, must be associated with the bones of one or the other 
of the large Dinosaurs already referred to. 

All the specimens on plate viii. are Dinosaurian ; and I should only 
differ from Dr. Buinzel in referring them, together with fig. 1 (which 
he terms tail-vertebre of a crocodilian) to the principal Dinosaur. 


FAUNA OF THE GOSAU FORMATION. 623 


Figures 2-4 are termed by the author vertebra of a foetal Dinosaur ; 
but I am not aware of any evidence which enables us to determine 
a matter of that kind, and I refer it to the same animal as the so- 
called Lizard-bones (pl. vi. figs. 14, 15, pl. vii. figs. 1-4). 

Indicating these and some other differences of opinion from Prof. 
Suess, and arranging the material, old and new, into species accord- 
ing to my interpretation, | was invited to deal with the remains in 
such a manner as my conclusions made necessary. As the time 
available did not suffice for description of the whole collection, I was 
generously permitted to borrow, from the museum of the University 
of Vienna, the more important specimens, which required further 
study or to be figured. The results I now offer to the Geological 
Society. The subject confessedly presents great difficulties ; and in 
the following memoir I have dealt with it to the best of my ability. 
As already stated, the bulk of Dinosaurian vertebre, scutes, and 
limb-bones are referable to two species of the same genus differing 
in size and other characters. This genus is certainly new. But when 
we come to examine the corresponding skull-fragments, there are two 
species indicated (by lower jaws) which are both of about the same size. 
There is also the somewhat smaller Dinosaur indicated by Biinzel 
as Iguanodon Suess; and there are teeth that appear to be referable 
to two other Dinosaurs, one resembling Lelaps or Megalosaurus, and 
the other somewhat approaching the Scelidosaurian pattern. Hence 
there is great difficulty in referring the right jaws to the skeletons ; 
and there is absolutely no evidence to show whether the hinder skull- 
fragment, called Struthiosaurus austriacus, belonged to one of these 
species, or is the only specimen of the animal hitherto discovered. I 
have therefore some doubt whether, in the endeavour to make the 
subject clear, a synonym or two may not be introduced, which can 
only be got rid of by the discovery of additional materials ; and I put 
my views forward with some diffidence. 

Of the new Chelonian indicated by costal plates which were sepa-~ 
rate from each other at the lateral margins, I find no other evidence 
except postfrontal bones indicating a skull covered with an elaborate 
pattern of minute scutes, and a strong but imperfect coracoid bone. 
Both these latter remains, however, are so typically Chelonian, 
although the skull-bones joined by squamose overlap instead of by 
suture, that I have no doubt of the propriety of including the costal 
plates in the Chelonian order, singular as is their form. This remark- 
able animal is associated with Emydian types which differ in no 
important respect from existing genera. The Pterodactyles are very 
impertectly represented, and badly preserved, and require but brief 
notice. The Crocodilians, however, are more curious, partly from their 
remarkable resemblances to types previously known in the Green- 
sand of New Jersey and Cambridge, and partly from displaying new 
characters in the vertebre. 

All the species hitherto discovered are peculiar to the deposit, and, 
with the exception of those temporally referred to Crocodilus, Emys, 
Ornithocheirus, and Megalosaurus, must, as it seems to me, be located 
in new genera. The most important new type is the Dinosaur Cra- 


624 PROF. H. G. SEELEY ON THE REPTILE 


twomus, represented by two species; the other Dinosaurian genera 
are Doratodon, Rhadinosaurus, Mochlodon, Ornithomerus, Oligosaurus, 
and Hoplosaurus. 


Condition of the Specimens. 


Almost all the bones are in fragmentary condition, and somewhat 
distorted by the effects of pressure. Being hollow, they have some- 
times become greatly crushed ; and were it not that both right and 
left bones are usually preserved, it would often be difficult to avoid 
being misled by appearances which result from conditions of fossili- 
zation. Unfortunately almost all the long bones have lost their 
articular extremities, and although in some instances this may 
perhaps be the result of fracture, yet in most cases it is certainly a 
consequence of decay of the bones before they were covered up in the 
deposit. I do not speculate as to whether the articular ends may 
have been eaten off by large carnivorous contemporaries ; for there 
are no indications of tooth-marks or other evidences of animals 
which might thus have mutilated the specimens. No doubt to some 
this condition would be proof that the remains were derived from 
an older deposit; but since the Wealden and Cambridge Greensand 
and Stonesfield Slate all have a number of bones in a not dissimilar 
condition, it seems to me less hypothetical to find an explanation 
of their condition in prolonged maceration, coupled with the litho- 
logical and petrological modifications which the deposit has since 
undergone. ‘There is, however, no record of natural association of 
any of the remains; yet, as they have mostly come from the same 
locality, it is probable that remains which agree in size and anato- 
mical characters may, in most cases, with certainty be referred to 
the skeleton of the same individual, since duplicate parts of the 
skeleton are almost unknown in each species. There may perhaps 
be a certain amount of doubt as to the correct association of the 
remains which I have ventured to put together; but this is a doubt 
which the anatomist will best appreciate who can realize the nature 
of the studies which have led me to group the bones as here set 
forth. 


Mocutopon Svurssi (Binzel). (Pl. XX VII. fig. 1.) 
See Biinzel, J. c. p. 8, pl. ii. figs. 7-11. 


One of the most beautifully preserved specimens is a right dentary 
bone of a small Dinosaur which at first sight exactly reproduces in 
miniature the characters of the Jguanodon of the Weald; but it 
differs in a character so remarkable that, had it occurred in a living 
animal, no hesitation would have been felt in relegating the jaw to 
a distinct genus. Anterior to the teeth, the symphysial extremity 
of every /guanodon-jaw bends round so that the rami form a U- 
shaped curve ; but this specimen is straight, and the anterior inward 
inflexion is scarcely appreciable, so that the snout was evidently 
sharply pointed, and therefore indicative of a new form of head. 
The fragment is little more than 73 centimetres long, and the tooth- 
bearing part of the jaw 17 or 18 millimetres; the height at the 


hart. Journ Geol.Soc Vol. XXXVILPI. XXX. 


a 


Ws) 


ie 


quart Journ Geo 


ore 


Hanhart imp 


PYILIAN REMAINS 


aay ere See leriae 


tan 


Pt imp 


a 


anh 


or 
ast 


iL 


Quart Journ Geol. Sc 


© Berjeau ith 


GOSAU REPTILIAN REMAINS 


FAUNA OF THE GOSAU FORMATION. 625 


posterior end to the top of the coronoid process, which is imperfectly 
preserved, is 34 centim. The external surface is smooth, rounded 
at the base, with a strong rounded ridge descending from the ante- 
rior margin of the coronoid process, and extending downward and 
forward along the jaw, dying away in front, and placed well above 
the middle of the side. Above this ridge the area of the jaw, ex- 
tending inward to the alveoli, is flattened almost horizontally at 
the back; but the area becomes more oblique anteriorly, and undi- 
stinguishable as the ridge subsides. Along its upper and outer 
margin towards this ridge is a series of foramina which are elongated 
or ovate—four of them larger, and half-a-dozen smaller. The area 
below the ridge is flattened towards the posterior limit of the dentary 
bone; anteriorly it is flattened and pointed, being bevelled on the 
inferior border for its union with the other ramus, and on the supe- 
rior border terminating in an oblique area, which is compressed from 
side to side and channelled by a somewhat deep groove. Whether 
this groove is merely vascular, or whether it may have contained a 
few premaxillary teeth, is a matter upon which I have no evidence. 
It is about 16 millim. long, wider and deeper behind than in front, 
and, as in the Wealden Jguanodon, has the inner border more ele- 
vated than the outer border. Below it are three or four vascular 
foramina. Both the inner and outer extremities of the jaw below 
are roughened, and indicate that the symphysis was loose, but held 
together by ligamentous union. As usual, the external outline of 
the bone, when viewed from above, is moderately convex, and its 
thickness from within outward continues to increase from before 
backward almost to the coronoid process. The inner side, which is . 
slightly crushed, displays ten alveoli. Portions of five or more teeth 
are seen in the jaw; and there are impressions of others and empty 
sockets, indicating ten in all. The first tooth, which is unfortu- 
nately imperfect (wanting the extremity of the crown), is remarkable 
for the smoothness of its inner surface, which, however, is elevated 
into a very strong median ridge, leaving the sides slightly concave. 
The serrations visible on the anterior margin are slight, and do not 
extend down the tooth. It is not sufficiently elevated to have come 
into wear. All the succeeding sockets are empty, owing to the teeth 
having dropped out; but most of them show successional teeth 
coming up, which have not yet reached the level of the outer alveolar 
margin. ‘The second and third teeth are broken away on their ex- 
ternal part, and not recognizable. The fourth tooth, also wanting 
the extremity of the crown, still shows the same enormously deve- 
loped median ridge; but external to it are, on each side, about half- 
a-dozen fine parallel ridges which have sulcations behind them of. 
about their own width. The fifth tooth only just shows the top of 
its crown coming up low down in the alveolus. The sixth toothgis 
’ the best developed, was apparently the largest, and occurs near the 
thickest part of thejaw. Its pattern is like that of the last described ; 
only the strong median ridge or keel is much sharper, and the lateral 
concayities deeper, in accordance with the width of the tooth. The 
lateral ridges run up and terminate in the sharp rounded lanceolate 


Q.J.G.8. No. 148. 20 


626 PROF. H. G. SEELEY ON THE REPTILE 


margin, and give it a crenulate appearance, which is due to their 
elevation. The median ridge terminates in a point, which is rounded 
and does not project beyond the tooth-margin. The tooth is slightly 
displaced, and leans backward towards the seventh socket. Low 
down in the seventh socket the crown of another tooth is seen. The 
eighth socket is empty. The ninth socket has lost the successional 
tooth, but displays the external impression, and shows it to have 
been marked with a median ridge and lateral finer ridges somewhat 
radiating upward. The tenth socket appears to have been small; it 
is imperfectly preserved, and there is no evidence as to its form or 
character; but a groove, which is smooth, is placed behind the last 
socket mentioned, and just in advance of the coronoid process. 
Hence these teeth appear to differ from those of Jguanodon in the 
persistent development of a powerful median ridge and in the stri- 
ation of the external surface. Behind the alveolar border the bone 
becomes squamous and thin, having overlapped the surangular bone, 
though there is no trace of a separate coronoid element, from the 
suture entering into the coronoid process. An opercular bone, or 
its representative ossification, appears to have extended along the 
broad subdentary groove at the base of the bone margining its upper 
part, while the angular bone, if it were distinct from the surangular, 
would appear to have reached far forward along the base of this 
groove, and to have rested on a thin ledge of the dentary. The 
submaxillary groove, in its anterior third, becomes shallow, but 
persists to the symphysis. ‘The region of the symphysis has no defi- 
nite outline. 

Two separate teeth, both such as may belong to this species, have 
been found. They are of small size, and may have belonged to 
this individual specimen. One belongs to the lower jaw, and might 
be the eighth tooth of the specimen described. The other is an upper- 
jaw tooth. The tooth from the lower jaw (Pl. XX VII. fig. 2) shows 
that the crown curves outward at a considerable angle to the fang; its 
outer margin is worn, showing that the teeth worked together with 
a scissors-like action, the lower-jaw teeth being, as usual, internal 
to those of the upper jaw. ‘The external surface is marked with 
about half-a-dozen primary ridges; between these, in the middle of 
the tooth, are finer ridges; and across them run transverse lines of 
growth. There is no median external ridge. The internal aspect 
of the crown is essentially the same as in the specimens described 
in the jaw itself. The median ridge, however, is not prolonged 
down the fang; and hence there is a slight constriction at the base 
of this ridge; and the elevated lateral ridges sharply define this 
side of the crown from the smooth lateral areas. _ 

The upper-jaw tooth (Pl. XX VII. figs. 3, 4) has the crown simi- 
larly curving inward from the fang. The fang is compressed from 
side to side, so as to give a subquadrate section. There is a slight 
constriction between the crown and the fang on the outer or cutting 
edge, but no constriction on the inner edge. ‘The fang is imperfect 
at the base; but the total length of fang and crown, as preserved, is 
18 millim. The worn surface (fig. 4), like that aspect of the tooth 


FAUNA OF THE GOSAU FORMATION. 627 


itself, is convex from side to side. The unworn part of the crown 
below is vertically striated; but the ribs are fainter than on the lower- 
jaw tooth. The inner surface of the crown is subquadrate, marked 
with eight vertical ridges, which are moderately elevated, stronger 
and wider apart on one side of the tooth than on the other. The 
height of the crown is about 8 millim., and its width about 7 millim. 
Its thickness at the base is about 4 millim. 


Parvetal bone of a small Dinosaur (probably Mochlodon). 
(See Binzel, pl. v. fig. 11, p. 14.) 


The parietal bone of a small Dinosaur (PI. XXX. fig. 1), which was 
regarded by Bunzel as a Lizard, shows, as I take it, the parieto-frontal 
suture in front, and an indication that the postfrontal bone was given 
off from the expanded anterior outer corner, much as in Jyuanodon. 
The under and interior surface of the bone, however, is much more 
lizara-like in some respects, seeing that it did not enclose a brain- 
case after the pattern demonstrated in Jguanodon, Hypsilophodon, 
Struthiosaurus, and other genera. The bone was relatively thin, but 
appears to have been united by a not very intimate suture to bone 
below, which formed the lateral wall of the brain-case. The bone is 
imperfect posteriorly, being fractured ; superiorly it is divided into 
three areas :—a median triangular area with concave sides, which 
becomes narrower posteriorly till it disappears at about the line of 
fracture (this surface is slightly convex from side to side in front) ; 
and two lateral areas for the attachment of muscles working the 
lower jaw, which converge posteriorly, and in converging are more 
highly inclined to each other. Their superior limit is sharply de- 
fined by a ridge, which becomes elevated posteriorly, and is appa- 
rently passing into a median crest, and is also elevated anteriorly at 
the point where the postfrontal suture is visible. The length of 
the fragment is 2 centim.; its width in front, as preserved, is 2 
or 3 millim. more; its width posteriorly is 14 centim. There is 
no foramen parietale. The characters are certainly such that the 
bone might well be referable to the skull of a Lacertilian; but it 
would be hazardous to determine absolutely on such evidence whether 
the bone really pertained to Mochlodon Suessii, as is rendered pro- 
bable by its Iguanodont form. 


Scapula (probably of Mochlodon). 


The imperfect proximal end of a small scapula (Pl. XXVIII. fig. 1) 
presents somewhat Crocodilian characters. The fragment is only 
4 centim. long. It shows the humeral articular surface and part of 
the sutural surface for the coracoid. The character which especially 
distinguishes it from Crocodiles is the extraordinary lateral position 
of the humeral articulation, in consequence of the sutural surface for 
the coracoid being prolonged beyond it. This articular surface is 
24 centim. long, 15 centim. wide proximally, and narrower towards 
the sutural surface. The bone is a little crushed, but was concave 
from above downward, and flattened in the antero-posterior direc- 

27 2 


628 PROF, H. G. SEELEY ON THE REPTILE 


tion. Hence the surface is narrower, more vertical, and more 
elongated, and especially more.concave than in the Crocodile; but 
there is a slight angle rising as a short ridge from the hinder exte- 
rior corner of the articulation, directed upward and forward, repre- 
sented in Crocodiles by a similar fainter ridge. Only the posterior 
part of the coracoid suture is preserved. It makes an angle of 45° 
with the humeral surface when seen from the front, and an angle of 
90° when seen from the side. The bone is narrower in front and 
behind than in the middle, where it is 12 millim. thick. So much as 
is preserved is 24 centim. long. Its external margin is convex; the 
internal margin appears to have been straighter. At the angle above 
its union with the humeral surface there is a small depression. 
Owing to the fact that the bone thickens externally with the humeral 
surface, the area anterior to that surface is concave and smooth. 
The concavity is directed obliquely downward and forward. There 
are indications that the anterior margin of the bone was deve- 
loped into an angular ridge, which may have corresponded with 
that of the Crocodile. The visceral surface of the bone was concave 
from above downward, and, though crushed, appears to have been 
flatter from side to side, more rounded on the anterior margin, and 
more compressed on the posterior margin about the humeral articu- 
lation than in Crocodiles. The blade of the bone, however, was 
similarly constricted at the fracture, where it is less than 2 centim. 
wide and about 1 centim. thickras’ preserved, convex in front and 
flattened behind. Though this bone is on the whole Crocodilian in 
its characters, it is also Dinosaurian, and perhaps makes the nearest 
approximation to the scapula figured by Prof. Marsh in the fore 
limb of his five-toed Dinosaurian Camptonotus dispar. 


SrRUTHIOSAURUS AUSTRIACUS, Biinzel. 
(See Bunzel, pl. v. figs. 1-6, p. 11.) 


The hinder part of the skull of a Dinosaur figured by Biinzel 1s some- 
what difficult to describe, on account of the obliteration or obscurity of 
the sutures ; and yet the anterior surface of the roof of the brain-case 
is margined by a well-marked transverse suture limiting the front of 
the parietal bone—a suture similar to that which persists in the skull 
of the Fowl long after other sutures have become obliterated in the 
hinder part of the cranium. The specimen certainly presents a remark- 
able resemblance to the back of the skull of a:bird; but I believe that 
Biinzel has attached more than due importance to this similitude, 
owing to the circumstance that the true nature of the Dimosaurian 
skull was even less perfectly known when he wrote than it is now. 
He has supposed his specimen to be more complete than, in truth, 
it is, being unaware, or unmindful, of the evidence that, external 
to the parietal bone, the Dinosaurian skull has-an upper arch or 
bar, like that so common in reptiles and unknown in birds, and 
that, as a rule, there is also a lower malar arch, more or less deve- 
loped, behind the orbit; and therefore it happens that the bone which 
he regarded as tympanic or quadrate, and interpreted as Crocodilian, 
is the paroccipital or opisthotic of modern anatomists, as, indeed, 


FAUNA OF THE GOSAU FORMATION. 629 


was long since appreciated by Mr. Hulke in describing his skull of 
Iguanodon*, Therefore Bunzel’s Avian and Crocodilian affinities of 
the skull both fall to the ground, owing to this fundamental miscon- 
ception of its characters. Hence it appears to me desirable to describe 
the specimen anew, in order to render its structure clearer. ‘The 
specimen exhibits superior, lateral, inferior, posterior, anterior, and 
cerebral aspects; and on each of these I propose to offer a few remarks. 
The fossil, as preserved, is 63 millim. broad behind, and 5 centim. 
high, owing to the downward direction of the occipital condyle ; for, 
although the skull obviously increases in height as it passes forward, 
the height from the base to the fronto-parietal suture is only 43 
centim. The presphenoid bone is broken away; but the length 
from the fracture or suture to the back of the occipital condyle is 
43 centim., and to the back of the supraoccipital bone is 5? centim. 
Superiorly and externally the cranial region is moderately convex 
from side to side, and also exhibits a slight convexity from front to 
back (PI. XX VII. fig. 6), especially towards the outer borders, indi- 
eative, I think, of the parietal bone just reaching the margin of the 
temporal fosse on the right side. The surface of the bone is rough, 
with slight and irregular close-set elevations, not so distinct as 
those of a Vrionyx, but certainly suggesting the surface that is 
sometimes seen when the scutes are removed from a Chelonian 
carapace. There is also a transverse furrow running across the 
bone, rather behind its middle, nearly parallel to the convex posterior 
border, and therefore curving backward. The width of this superior 
surface, as preserved, is nearly 53 centim.; but then the bone is 
broken on both sides, though it has become thin and separated late- 
rally from the brain-case. Its antero-posterior extent in the middle 
line where greatest is just over 3 centim. I have no doubt that 
the transverse groove (Pl. XX VII. fig. 5) indicates the limit of the 
parietal bone ; for the suture defining it is seen on the left side of the 
cerebral surface and on the external lateral surface; but I cannot 
trace it ucross the upper surface, and it may be that the suture is 
obliterated by ossification, consequent upon a scutal covering. The 
groove recalls those which occur on the skulls of Lizards such as 
Trachydosaurus, while the texture of the bone is not dissimilar; and 
hence it is also possible that we have here an explanation of the 
absence of sutures, in the circumstance that they are covered up by 
a layer of dermal ossifications. The parietal bone at the frontal 
suture (Pl. XXVII. fig. 6) is 9 millim. thick; but at the transverse 
groove the thickness is reduced to 7 millim., owing to cerebral exca- 
vation beneath it. The area behind the groove terminates posteriorly 
in a margin which is rounded, but suggests the idea that a plate 
4 millim. thick in the middle, and becoming thinner laterally, was 
superimposed upon the cranial bones. This region posterior to the 
groove I suppose to be occupied by the supraoccipital bone. 

The posterior aspect of the skull is chiefly remarkable for the 
elevated border above the foramen magnum, which was evidently 


* Hulke, Quart. Journ. Geol. Soc. vol. xxvii. p. 206. 


630 PROF. H. G. SEELEY ON THE REPTILE 


in somewhat close contact with the neural arch of the atlas, and for 
the transverse grooves and muscular rugosities, which run between 
this border and the slight groove defining the supposed cranial scute. 
Hence the back of the skull is not vertical, like that of some Croco- 
diles, and its superior margin is far from being as well rounded as in 
very young Crocodiles ; and on the whole there is nothing to call for 
remark as affihating this region to what is seen in either Crocodiles 
or Lizards. The surface ascends somewhat obliquely, but im two 
terraces ; that immediately above the ridge bordering the foramen 
magnum is divided by it into two lateral portions. These lateral por- 
tions are channels extending outward and downward, and widening 
as they go. There are, on each side of the median vertical dividing 
ridge in these channels, three large tubercles. Above the channels, 
and as nearly as may be of corresponding size, is a pair of convex 
surfaces, which are undistinguishable in the middle from the median 
bar just referred to. They are margined above by the supposed 
cranial scute, and, as they extend outward, widen and curve obliquely 
upward; and a muscular ridge appears dividing the outer part of 
this wedge longitudinally into two nearly equal parts. Hence the 
pattern of the back of the skull as preserved is very like a capital 
letter K placed transversely, so that the two diverging limbs corre- 
spond to the ridges above the foramen magnum. The height from 
the top of the foramen magnum to the summit of the back of the 
skull is 2 centim. The transverse width over the supraneural ridges 
is 4centim. The width of each ridge at its outer third, before its 
upper border becomes concave, is 9 millim., the upper margin extend- 
ing over the concave channel above it; higher up its width is little 
more than half. The median connecting ridge between the two 
transverse ridges at the back of the head is about 12 millim. wide. 
Though the back of the head as a whole is convex from side to side, 
it is concave from above downward in the median line. The foramen 
magnum is slightly elliptical, being 17 millim. wide and about 15 
millim. high. The skull presents the unusual condition that the 
basioccipital condyle retreats below and in front of the upper border 
of the foramen magnum, so that, placing the back of the skull verti- 
cally, which puts the base of the skull horizontal, the back of the 
brain-case projects for a centimetre behind the basioccipital condyle. 
The areas at the outer and upper corners of the occipital condyle 
are concavely notched, and at first convex from side to side, but 
more flattened as they pass outward and upward. The depth from 
the back of the skull to the base of the occipital condyle is 4 centim. 

The base of the skull, as preserved, is triangular, 54 centim. 
from the hinder border of the foramen magnum to the front of 
the basisphenoid, or a fracture in front of the sella turcica. The 
hinder border of the triangle is convex, and the lateral border is 
concave, though all the borders are irregular. There is no sutural 
distinction between the basioccipital and basisphenoid, any more 
than between the basioccipital, exoccipitals, and supraoccipital bones. 
The basioccipital condyle probably is formed to some extent by the 
exoccipital bones, much as in the Crocodile, since foramina occur 


FAUNA OF THE GOSAU FORMATION. 631 


some little distance in advance of the basioccipital, which may be 
presumed to penetrate the exoccipital bones and give passage to the 
pneumogastric and hypoglossal nerves. The occipital condyle is 
well rounded ; but its outline is subtrapezoidal ; its greatest width in 
the upper third is 2 centim., its greatest depth 16 millim. The 
under surface is deeply channelled, so that the thickness of the bone 
behind the articular surface is one millim. ‘The region in front of 
the occipital condyle is about 357 centim. long, fairly smooth, but 
concayely excavated in the middle, both in length and breadth, rising, 
however, to a rounded margin at the sides internal to the lateral 
foramina. ‘The width across the pneumogastric foramina is 3 centim. 
The lateral margin of the triangle is divided by a median convexity 
into two concavities: the shorter, in front, is about 2 centim. long; 
and the longer, behind, is a ttle more; while posterior to this, on the 
left side, is a surface which appears to have been laterally sutural 
and nearly vertical, while a suture on the opposite side shows that 
the upper part of this mass consists of a small bone, which readily 
comes away. Hence I interpret the lateral masses of bone external 
to the foramen magnum as being the exoccipital bones, as in Croco- 
diles, while the small bone above the outer border of the exoccipital 
is the paroccipital of Owen; and I suppose the exoccipital to extend 
forward so as to form the side of the wall of the brain-case; so that 
no portion of the posterior lateral structure preserved can be the 
quadrate bone, as supposed by Biinzel, and hence the analogy 
attempted to be made out in this region of the skull with the 
Crocodile can have no foundation. 

I would next note the characters of the lateral aspect of the skull. 
(Pl. XXVII. fig. 5). Here all the bones which are connected with 
the roof of the brain-case are more or less broken, and the bones have 
disappeared which formed the external suspensory arch for the lower 
jaw, so that nothing remains but the internal part of the head, 
which may be likened, perhaps, to that of a Crocodilian type in 
which neither were the quadrate bones blended with its lateral walls 
nor the pterygoid bones connected with its base. I fail altogether 
to recognize a Lizard-like type, although, as at present used, the 
term Lizard is almost large enough to include any thing. Forms 
like Amphisbeena, which have the quadrate bone firmly wedged into 
the skull and no trace of either of the postorbital arches, might well 
be regarded as a distinct ordinal type; and there are some Dinosaurs 
towards which the structures of the hinder part of such a brain-case 
somewhat approximates ; but the cranial bones in ordinary Lizards, 
like Iguana, form a part of the skull that is very imperfectly con- 
nected with its roof, and very different from the structure seen in 
Crocodiles and Dinosaurs, though other Lizards, ike Cnemidophorus 
for instance, have a better union between the brain-case and the 
surrounding bones; but I do not recall any type of Lizard that so 
far corresponds in the characters of the bones covering the brain 
with what is seen in Dinosaurs as to justify us in affirming that this 
skull is lacertilian. Turning our attention first to the basal part of 
the brain-case, it will be seen that the articular head of the basiocci- 


632 PROF. H. G. SEELEY ON THE REPTILE ~ 


pital is directed almost vertically downward, showing, I think, that 
the head must have been carried upon the neck as in Deer, Kangaroos, 
and other animals in which the position of the neck is vertical. The 
part of the basioccipital posterior to the sella turcica is flattened on 
its cranial surface, gently concave from side to side, with a slight 
median ridge. Then at a distance forward of about 3 centim. from 
the occipital condyle, the brain-case thickens from below upward, 
but does not present the cup-shaped depression seen in Crocodiles 
and many birds, though its edge is obviously destroyed by fracture, 
and must have extended some millimetres higher than the 13 milli- 
metres preserved. At the border of this ridge, on each side, is a 
large perforation for the second nerve. The inner and anterior wall 
of the perforation is broken away ; and the transverse width between 
the outer walls of these foramina is2 centim. Below these foramina, 
and extending forward, are several others. First, on the right side 
is a large foramen that runs obliquely outward and forward, pene- 
trating into the brain-case. On the opposite side, instead of one 
large foramen, there appear to be two, divided by a considerable 
intervening piece of bone. I have no doubt that this foramen gives 
passage to the fifth nerve. On the left side it is prolonged backward 
and outward in a horizontal groove; and although there is a groove 
on the right side, and though it is smaller and shallower, it does not 
impress the margin of the bone. In advance of this foramen, and 
below the sella turcica, in the anterior concavity of the bone, are 
two other foramina, which appear to be vascular. The anterior one, 
owing to fracture, is seen on the left side to curve obliquely down- 
ward and forward, and open upon the base of the skull. Between 
the outlets of these foramina, in the median line, is a portion of a 
conical foramen, the anterior wall of which is removed by the 
fracture. 

The vertical fracture of the sphenoid (Pl. X XVII. fig. 6), where it 
terminates, is triangular, 2 centim. wide, and about as high, the sides 
being compressed so as to meet superiorly in a crest which rises in front 
of the pituitary fosse, as in some birds, and not at all as in Crocodiles. 
Its walls are concave at the sides; and posteriorly a slight longi- 
tudinal ridge rises, which becomes directed at an angle inward and 
upward to the ridge bordering the pituitary region behind. ‘The pit 
for the pituitary body is about 12 millim. long and 9 millim. wide, 
is concave from side to side, and margined by a sharp elevated ridge. 
Its anterior part is overhung by the process of the sphenoid, which 
rises above it. The posterior portion of the lateral aspect of the 
skull consists of a small superior area, subtriangular, formed appa- 
rently partly of the parietal and partly of the supraoccipital bones. 
It is a smooth internal surface below the fractured roof of the brain- 
case, directed obliquely outward, downward, and forward, and 
traversed by a groove which probably indicates a suture between 
the two bones. The posterior border is a sharp knife-edge, concave 
from before backward. Under this edge a deep excavation extends, 
penetrating to within half a centimetre of the brain-case. The ex- 
cavation is smoothly concave, extends longitudinally, and is pro- 


FAUNA OF THE GOSAU FORMATION. 633 


longed backward and outward above the otic bones; but the sur- 
face is prolonged obliquely outward and downward, so as to form a 
smooth quadrate area, which rounds towards the base of the skull, 
and terminates backward on the left side, where best preserved, in 
a vertical lunate surface, convex behind and concave in front, formed 
by the exoccipital and otic bones. In front these spaces slightly 
converge, especially below and above; and behind the middle there 
is a foramen a millimetre or two in diameter, apparently largest on 
the left side, but smaller than might have been expected if it is the 
entrance to the auditory chamber, which probably lies in the depres- 
sion above it. From the lateral lunate surface to the inner wall of 
the brain-case above the occipital condyle is 24 centim. 

Finally there remains the interior cavity of the skull which held the 
brain(Pl. XX VII. fig. 6). This does not present any great contraction 
in the auditory region. Its extreme width behind is 17 millim., where 
the auditory bones bulge inward after the manner of Crocodiles. The 
transverse width of the brain-case is thus reduced to 13 millim. ; but 
at the same time its height increases from 14 millim. behind to about 
2+ centim. in the region of the auditory prominences, though the 
extreme height of the brain-case is somewhat in advance of this 
point, where it becomes 3 centim. ‘The width continues to increase 
from behind forward to the parieto-frontal suture. It is greatest 
in the upper third of the outline of the brain, where it amounts to 
22 millim. A bone which in a bird might be regarded as the ali- 
sphenoid, which lies above the sphenoid, appears to meet the parietal 
and exoccipital by a well-defined suture, visible externally and in- 
ternally, and running obliquely downward and backward. It is 
difficult to speak with confidence of the limit of this bone on the 
external surface, since as it extends backwards it is only preserved 
on the left side. The suture from which it has come away is well 
defined on the right side. Its anterior border is sharp; and the 
external surface is concave from within and outward. This sharp 
border appears to show that in this Dinosaur the brain-case was not 
completely closed in front in the middle line. Anterior to the 
highest point of the upper wall of the brain-case, which lies under 
the transverse scute-like groove crossing the external surface, the 
bone makes an angular bend forward; but though there are many 
little irregularities of outline in the internal surface, there is nothing 
so important as the bending inward and downward of the lower 
part of the alisphenoids, which must have made the transverse 
section of the cerebrum nearly circular at the parieto-frontal 
sutures. 

Imperfect as this description is, it will suffice to show that we 
have here a Dinosaur of a type so different from that indicated by 
the skull referred by Mr. Hulke to Jquanodon, as only to be classed 
in a separate suborder; and if the base of a skull figured by myself 
under the name Craterosaurus be, as 1 believe, also Dinosaurian, 
that also indicates a subordinal type, and is totally distinct from 
either of the others. These great differences of skull-structure lead 
me to suspect that the Dinosauria are a far more important group 


634 PROF. H. G. SEELEY ON THE REPTILE 


than has hitherto been suspected *; and it may well be that different 
genera present modifications which affiliate representatives of the 
group towards Crocodiles on the one hand and birds on the other. 
But I cannot believe that any order, however homogeneous, could 
have spanned the interval between the Crocodile and the bird, 
though there can be no doubt that this skull of Struthiosaurus makes 
a nearer approach towards the bird than does the skull of any living 
reptile ; its differences from the bird-skull are precisely those which 
distinguish it from the Crocodile, little as we know or can infer con- 
cerning the suspensory arches for the lowerjaw. In the base of the 
skull not being covered with pterygoids there is a notable difference 
from Crocodiles of the surviving type; but then the base of the skull 
is not bird-like, any more than it is like that of any other animal. 
It is one of the most distinctive points of the Dinosaurian skeleton. 
Itwould bedesirable to compare this specimen with other Cretaceous 
genera ; but, with the exception of Acanthopholis, none of these have, 
as yet, ylelded any evidence of the brain-case. One fragment, found at 
Folkestone at the base of the Chalk with the remains of the Acantho- 
pholts horridus, is briefly referred to by Prof. Huxley; and on in- 
spection it proves, though clearly allied, to belong to a different genus, 
a fact that will be best demonstrated by a description of the specimen 
and comparison of the figures (Pl. XXVIL. figs. 6 & 8 and 5 & 7). 


Nore on THE Base oF THE SKULL oF Acanthopholis horridus, Huxley. 


Professor Huxley’s account of the skull of Acanthopholis is so 
brief that it would be difficult to be sure from it of the identity of 
the specimen, especially since Prof. Huxley describes characters 
which we are now unable to recognize, though it is, of course, 
possible that the specimen is in a less perfect condition than when 
originally noticed. I therefore reproduce Prof. Huxley’s original 
remarks (Geol. Mag.1867, vol.iv., Huxley on Acanthophols horridus, 

. 66). 

ae Of the skull I possess only a very much mutilated fragment, 
showing the basioccipital and basisphenoid. ‘The occipital condyle 
measures 1-4 transversely, or has about the same diameter as that 
of the skull of a Crocodilus biporcatus which measures 16 inches in 
length from snout to occiput. But itis more elongated transversely 
and excavated above than in the Crocodile, and the exoccipitals 
enter more largely into its composition. The Crocodilian disposi- 
tion of the Eustachian tubes is absent ; and the carotids run up the 
side of the basisphenoid in Lacertilian fashion. The sella turcica has 
a, well-developed posterior plate.” 

This fragment (Pl. XX VII. figs.7 &8) comprises the base of the skull, 
and includes the basioccipital and basisphenoid, which are completely 
ankylosed, and give no indication whatever of suture. Iam similarly 
unable to detect any sutural evidence of the exoccipital; nor can I re- 
cognize the basioccipital condyle, which I believe to have been directed 


* Professor Marsh, since this was written, has published a classification of 
American Dinosaurs (Amer. Journ. Se. vol. xxi. p. 423). 


FAUNA OF THE GOSAU FORMATION. 635 


downward, and to have been largely removed by attrition, so that no 
idea can now be formed of its relative depth. The presphenoidal part 
of the specimen is broken away and terminates anteriorly in a trian- 
eular transverse fracture (fig.8). The base of the skull, as preserved, 
is 54 centim. long. In the middle it is 4 centim. wide, but narrows 
posteriorly to the region of the condyle, where the bone is 33 millim. 
wide. Auteriorly it also appears to contract a little; but at both 
ends the external white film of bone has scaled off, leaving the dark 
phosphatic substance below—a condition reminding one curiously 
of the pale and dark mineralization of bones in the Cambridge Green- 
sand. ‘This inferior region is concave in length, with a rounded 
median ridge, and lateral concavities on each side of it in the middle. 
Posterior to the middle area the bone is fractured inferiorly for a 
length of 24 centim.; and this fracture I suppose to have removed 
the lower half and characteristic form of the occipital condyle. 

The basioccipital bone(P1. XX VII. fig. 7) terminates posteriorly in a 
mass which, as preserved, is convex below and concave above, so as to 
have a crescent outline, and is also moderately convex from side to side. 
Above it is the brain-case, which certainly extended somewhat further 
backward than the present limits of the occipital condyle, as is shown 
by the form and character of the lateral walls. The superior surface, 
however, of the occipital bone appears to curve convexly downward 
as it extends backward ; and, as preserved, the bone is little over 
1 centim. deep, and about 3 centim. wide at the origin of the con- 
dyle, which is, as usual, defined by a lateral constriction that can 
only be detected by careful examination, and is some distance pos- 
terior to the lateral notches on each side of the base of the brain- 
case. 

The cranial cavity (fig. 8)is imperfectly defined, because there is no 
portion of the roof of the brain-case preserved, and its lateral walls 
are imperfect. It is evident, however, that it is higher than wide; 
the posterior width in the region of the foramina for the hypoglossal 
nerves is 28 millim., while the height appears to have been not less 
than about 4 centim. On the right side a part of the inner wall of 
the brain-case is exposed, showing that it is smooth, bulges inward 
a little in the auditory region, and is inclined a little inward as it 
extends upward. On the left side the bone is fractured, so as to 
show that the hinder wall of the brain in the auditory region is 2 
centim. thick, and extends outward transversely at a little higher 
level than the base of the skull. It shows a horizontal semicircular 
canal, which extends from the wall of the brain-case outward and 
forward for a length of about 15 centim., as exposed, and is 
about 3 centim. wide. The curve cannot be followed round; nor 
can its relations to the other semicircular canals be definitely made 
out. 

Along each side of the floor of the brain-case, and under the 
transverse jutting of its lateral walls, which extend out horizontally 
behind, is a row of foramina which extends in a curve, just sepa- 
rated from each other by bony interspaces. Six are visible on the 
left side; on the right side there were certainly five, and may have 


636 PROF. H. G. SEELEY ON THE REPTILE 


been six (fig.’7). ‘These foramina are different from those of Struthio- 
saurus or Iguanodon, and furnish a marked character, defining Acan- 
thopholis. It may be difficult to correlate them with the foramina in 
the back of the skull of a Crocodile; but since those perforations are, 
for the most part, in the exoccipital bones, and extend downward at 
the back of the skull, it is obvious that we have here in the longitu- 
dinal arrangement something more nearly paralleled by Lizards,where 
the twelfth, eleventh, tenth, eighth, fifth, and second nerves are given 
off in more or less longitudinal series. The hindermost foramen 
may be referred to the hypoglossal nerve, the next, perhaps, to the 
pneumogastric nerve, then perhaps a vascular foramen. ‘The two 
hinder foramina are much smaller than the third; and the third 
foramen may probably be for the eighth nerve; the fourth is small; 
the fifth is so large that it might well correspond to the fifth nerve. 
The anterior direction of the sixth makes it probable that we have 
here the foramen for the optic nerve; for though it is somewhat 
smaller than might have been expected, it is given off from the most 
anterior part of the side of the brain-case behind the sella turcica. 

The anterior extremity of the basisphenoid is massive and wedge- 
shaped, broken away on the compressed inferior lateral margins, as 
well as in the front. A strong vertical plate rises in the middle, so 
as to form the anterior border of the brain-case (fig. 8). The upper 
margin is 34 centim. above the base of the skull, and it is nearly 
2 centim. behind the anterior fragment of the basisphenoid preserved. 
This plate therefore seems to me to be exactly in the position of the 
posterior border of the sella turcica; but if so, the anterior border, 
such asis seen in Struthiosaurus (figs. 6, 5), is entirely broken away. 
What remains of the sella turcica is a concave base in front of the 
plate, terminating anteriorly in two diverging concave streaks of 
bone-surface, which probably represent the channels of the caro- 
tids (fig.8). They extend downward and outward, making an angle 
of 90° with each other, and do not appear to reach forward. On 
each side of the posterior plate of the sella turcica there is a con- 
cave notch in the skull-wall. 

The skull diverges so far from both the Crocodilian and Lacertian 
types that it may be as well to recognize it as equally distinct from 
both. It resembles Struthiosaurus in the downward direction of 
the occipital condyle, in the extension of the lateral wall of the 
brain-case posterior to the condyle, in the transverse horizontal ex- 
pansion of the exoccipital region in front of the occipital condyle, 
in the massiveness of the bone in the auditory region, and in the 
grouping of the foramina, so that the posterior three are inferior, 
while the anterior three have a more anterior and lateral position ; 
but the convex form of the base of the occipital bone, the immense 
thickness of the basisphenoid bone, are matter for distinction, as is 
the form of the alisphenoid in Acanthopholis. The resemblances, 
however, are so remarkable as to show that these two genera are 
near allies; and though we cannot infer with certainty the roof of 
the brain-case of Acanthopholis from that of Struthiosaurus, or the 
teeth of Struthiosaurus from those of Acanithopholis, yet they seem to 


FAUNA OF THE GOSAU FORMATION. 637 


me to show that Struthiosaurus was probably a Scelidosaurian, and to 
open up a suggestive possibility of its claim to the jaws and teeth 
which have a Scelidosaurian character. Future researches may 
possibly demonstrate it to be the skull of Cratewomus; but as the 
back of the skull of Siruthiosaurus is so different from that of Sceli- 
dosaurus, I have not felt justified in adopting such a view. 


CRATHOMUS. 


The dermal armour of this genus presents a remarkable resem- 
blance to that of the Scelidosaurian Dinosaurs. ‘The large supra- 
vertebral scutes of the caudal region are compressed, and terminate 
upward in a sharp knife-like edge. They are, perhaps, more like 
the similar scutes of Scelidosaurus than those of Acanthopholis. The 
flat dorsal scutes which were carried on the ribs were also keeled; but 
the keel was relatively lower, and the plates were more or less ovate. 
This, too, is a character paralleled in Scelidosaurus; but there are 
also scutes without any ridge at all, and marked with deep vas- 
cular grooves. These I regard as probably ventral. Coming probably 
from the region of the shoulders are two remarkable scutes which 
are quite unlike any thing at present figured. These plates, which 
are excavated on the underside, terminated in a sharp spine at each 
end; and the middle of the scute bore upon its surface a number of 
conical ossifications, which have much the appearance of a group of 
limpets packed close together. These ossifications have exactly the 
appearance of the scutes of Hylwosaurus, so much so as to suggest a 
doubt whether the armour hitherto referred to Hyleosaurus may 
not be unankylosed scutes separated from the plate which carried 
them, and really referable to Polacanthus, in which Mr. Hulke has 
found an armour closely approximating to that seen in this genus. 
Finally, there is a scute bearing a bone exactly like the horn-core 
of an ox; this | am also disposed to refer to the fore quarters. The 
distinctive features of this armour are the sharpness of the caudal 
scutes and the form and patelloid incrusting of the cervical scutes ; 
but in other characters it approximates to the genera already 
named. 

The vertebral column is remarkable for the forward extension of 
the neural arch in the neck and the deep gap between the anterior 
and posterior zygapophyses, the shortness of the neural spine, and 
the biconcave form of the vertebrze, while the dorsal vertebree are 
remarkable for the great strength of the ridge below the transverse 
processes, the distinctness of the facets to which the ribs were arti- 
culated, and the broad rounded base to the vertebre. The caudal 
vertebre have somewhat the form characteristic of Acanthopholis, 
having a groove in the middle line of the base ; but the single lateral 
ridge is a point of distinction, though the renee bigs were ‘obviously 
nearly allied to those of that genus. The ribs, in having the superior 
margin flattened and widened to a greater extent than the depth of 
the bone, present a character that is found in all reptiles which 
carried heavy armour, but is especially characteristic of this form, 
though, according to Mr. Hulke, met with also in Polacanthus. The 


638 PROF. H. G. SEELEY ON THE REPTILE 


fragments of jaw with the teeth, if rightly referred to this genus, 
present a character similar to that of Priodontognathus, Scelidosaurus, 
and Acanthopholis; but while nearly resembling Scelidosaurus, the 
tooth-structure is distinctive in the character of the serrations, just 
as the lower jaw is distinctive in its angularity and vascular foramina. 
But it is the hmb-bones which best define Cratwomus. The scimitar- 
shaped scapula, with its powerful acromion process, is altogether 
distinctive, while what remains of the coracoid appears to indicate 
an equally unusual form. The humeri are remarkably powerful, and 
indicate an animal strong in its fore limbs, evidently a quadruped, 
and therefore presumably carnivorous, since the herbivorous forms 
have the fore hmbs feebly developed. The humerus, with a general 
resemblance to that of Anoplosaurus, is far more robust, and indicates 
a heavier animal: no bone anterior to the humerus is known. The 
femur is distinguished from that of the Iguanodonts by wanting the 
separate external trochanter at the proximal end. It has the arti- 
cular ends powerfully developed, and, perhaps, most closely resembles 
in general form that of Cryptosaurus eumerus of the Oxford clay. The 
tibia is remarkable for the extremely compressed form and forward 
development of the cnemial crest. The fibula, so far as preserved, 
is very like the fibula of a bird, and bears a similar relation of size 
to the tibia. The metatarsal and phalangial bones, if belonging to 
this genus, rather indicate a flattened foot,terminating in claws which 
were broad rather than sharp. Taken asa whole, far more difference 
from Cratcwomus is found in described genera in the structure of the 
internal skeleton than would have been inferred from either the 
armour or the teeth; and it is quite possible that the armour, espe- 
cially in Dinosaurs, may have undergone as little change as the 
feathers of birds or scales of lizards, so as to be common to several 
families. 


Mandibles and Teeth probably referable to Crateeomus. 


Three fragments of the anterior extremities of Dinosaurian lower 
jaws have been found which indicate two species, though the re- 
mains are so fragmentary that they cannot be defined with the 
detailed accuracy which is desirable. Both specimens are of about 
the same size, and belong to a genus which is closely related to 
Priodontognathus. I will describe the more perfect specimen first. 

This species is represented by a dentary bone (Pl. XX VII. figs. 9, 
10), the anterior extremity of which is unfortunately not preserved ; 
nor is the fragment complete on the hinder or lower border, though 
it probably gives indications of the whole of the teeth. The alve- 
olar border is bent in a sigmoid flexure (fig. 9); and the bone itself is 
bent so as to present a flattened lower part at right angles to the upper 
part of the side behind, but sloping more and more outward in front. 
The lateral contour of the alveolar border is convex, rising higher in 
the middle and descending to near the level of the base; it has 
a width of about 6 centimetres. As usual with Dinosaurs, it is 
higher on the external than on the internal margin. The teeth 
were placed in sockets defined and separated by narrow bony inter- 


FAUNA OF THE GOSAU FORMATION. 639 


spaces. The sockets were circular, and indicate larger teeth in the 
fore part of the jaw than in the hind part. ‘The alveoli for twenty 
teeth are shown (fig. 9); they did not reach to the extremity of the 
jaw; nor apparently was there any bony union between the rami; but 
the small fragment anterior to the termination of the alveolar 
margin is broken away. The length of the alveolar margin is 
about 8 centimetres; and the extreme length of the fragment is 
under 9 centimetres. The internal aspect of the jaw has at its 
base a deep groove, which widens from before backward, and 
passes close to the base of the jaw (fig. 9), though it appears 
to slightly ascend, and no doubt lodged the opercular bone. 
The basal margin below this groove is rounded; the surface above 
the groove is smooth, and forms an obliquely twisted area, which 
maintains a depth of about 13 centimetre, so far as it is preserved. 
It is very slightly convex in length, but concave from above down- 
wards, the concavity increasing forward owing to the increasing 
twist in the bone. The depth of the jaw at the first alveolus is 
18 millimetres. At the tenth alveolus it is nearly 2 centimetres ; 
but the depth cannot be given further back, as the base of the jaw 
is broken away. The hinder part of the inner alveolar border shows 
indications of a squamous bone having come awey. This would pre- 
sumably be part of the opercular bone. 

The external surface (fig. 10), as already remarked, is traversed, 
at least in part, by a strongly elevated ridge, which inclines a little 
downwards as it extends forward, and dies away towards the anterior 
end. The surface below this ridge is flattened, but very slightly 
convex from above downwards, and, so far as preserved, is straight. 
It shows a few deep narrow vascular grooves and markings for 
vessels. The superior part of the side is obliquely twisted, be- 
coming more and more horizontal behind, and more and more ver- 
tical in front. In the middle of the side, both in length and depth, 
are four large foramina (fig. 10), placed close together in a line, 
seven millimetres below the alveolar border, and 7 millimetres above 
the longitudinal angle in the middle of the bone. These foramina 
and their interspaces extend over a length of about 28 millimetres. 
The anterior one descends vertically ; the three posterior ones enter 
the bone obliquely, being directed downwards, forwards, and inwards. 
From the hinder and inferior corner of the last foramen a slight 
ridge is prolonged backwards, which makes an angle in the upper 
margin of the jaw. ‘The surface anterior to these foramina rounds 
convexly from above downwards; and below the third to sixth 
alveoli there are about four minute foramina, and below the seventh 
and eighth, only much nearer to the alveolar margin, two others. 
It would thus appear as though a series of foramina had extended 
along the bone, of which the middle four had become greatly de- 
veloped. ‘The thickness of the jaw from within outwards augments 
along the line of the median lateral ridge; in front it is about six 
millimetres, in the middle 12 or 13 millimetres, and obviously in- 
creases aS it extends further backwards. What remains of the 
inferior margin, the anterior 4 centimetres, is concave; and the 


640 PROF. H. G. SEELEY ON THE REPTILE 


margin curves downwards and inwards as 1 extends forward. The 
fourth alveolus is the only one which displays an indication of tooth- 
structure. It is the extremity of a compressed arrow-shaped suc- 
cessional tooth with serrated border, more after the pattern of that 
seen in Priodontognathus than in any other genus, but too imper- 
fect to demonstrate the generic characters. It appears to be more 
elongated than the teeth of any genus hitherto described. It may 
pertain to one species of Cratwomus. 

The other fragmentary pieces of lower jaws may or may not belong 
to one individual. A terminal fragment pertains to the anterior end 
of the right ramus. Two other and smaller fragments belong to the 
left ramus; but they are so imperfectly preserved as to be scarcely 
worth notice, although they are apparently quite distinct from the 
species just described, if I may judge from the flatness of the in- 
ferior surface of the jaw and the flatness of the lower part of the 
side which was vertical. 

The anterior extremity of the right ramus was loosely attached, 
by a rough lunate surface about 17 millimetres deep and 8 milli- 
metres wide, to the ramus on the opposite side. Its extremity is 
bent a little inwards and downwards—the basal margin being con- 
cave from in front backward, and the prolongation of the alveolar ~ 
margin convex. The fragment is 4 centimetres long; and though 
upwards of 3 centimetres of the alveolar margin are preserved, 
I do not recognize with certainty any tooth-sockets. If such 
exist, they are three in number, and are indicated by small round 
sockets placed just behind the symphysis; but as the whole anterior 
end of the bone is covered with vascular foramina, and there are 
corresponding foramina external to these possible sockets, it is not 
improbable that they are foramina also, since they present no dis- 
tinctive alveolar characteristics. The jaw thickens a little in 
front here; it is bevelled, looks obliquely forward, upward, and, 
perhaps, outward, and has the appearance of having utilized the 
foramina in the nutrition of a pad. This surface is about 32 cen- 
timetres long, and above the symphysis is 13 millimetres wide, but 
becomes narrower posteriorly, where it terminates on the inner edge 
of the jaw in a sharp margin, external to which two large oblique 
foramina appear at intervals, the second of which seems to be ex- 
ternal to the first tooth-socket, which is compressed from side to 
side, if it really be a socket, of which there is some doubt. At this 
point the depth of the jaw is about 23 centimetres. Below the an-- 
terior area described, the upper portion of the side which is smooth 
begins to be concave from above downwards ; and the lower part of 
the side which is rugose is here convex, though it may, perhaps, as 
indicated by a fragment already referred to from the other side of 
the jaw, become flattened in its posterior extension. The specimen 
shows no trace of the groove on the inferior margin of the inner 
side seen in the specimen already described; and the appearance of 
a groove in the upper part probably results from fracture. The 
internal surface is smooth and concave in length. The thickness of 
the jaw at the posterior fracture is about 11 millimetres; the base, 


FAUNA OF THE GOSAU FORMATION. 641 


which is flattened, is nearly at right angles to the internal and ex- 
ternal surfaces, and altogether unlike the narrow rounded base 
which characterizes the species previously described. This, with 
the less extension of the alveolar margin forward and greater 
thickening of the extremity of the jaw to form the parrot-like ter- 
minal surface, constitute specific distinctions. 


Teeth. 


The teeth have very much the aspect of having been eaten 
(Pl. XXVIT. figs. 11-16), or at least exposed to some solvent which 
may have slightly dissolved their surfaces; but the contours are 
sharp and well preserved ; and though the fangs are in some cases 
broken, the teeth do not exhibit the indications of ordinary wear. 
It is very difficult, since they only number nine, to judge whether 
the differences which are to be detected result from relative position 
in the jaw, or whether one modification at least is not, as I am 
inclined to believe, of specific importance. 

These teeth have a triangular crown and a compressed fang. 
There is a cingulum at the base on the outer side only; but it 
merely serves to give a compressed aspect to the base of the 
crown and to thicken the top of the fang. Even these teeth 
exhibit certain modifications. First, there is one with the fang 
perfect ; and this shows that it 1s closed ; and on the inner side at 
the base it curves a little and shows an impressed area, as though 
a successional germ had rested there; the fang in its upper part is 
slightly concave from side to side. The tooth is bevelled off obliquely 
on each side by the cutting-edge of the crown (Pl. XXVII. fig. 13). 
The other side has a transverse cinguloid ridge, considerably lower 
in position than the bevellings (fig. 14). It extends up towards 
the bevelled corners at the sides. The crown is convex from side to 
side; but the median longitudinal ridge is not distinctly defined. 
Below the cinguloid ridge the tooth contracts from side to side. 
The extreme length of the tooth is over 9 millimetres, the extreme 
width of crown is about 53 millimetres, and its length down to the 
base 5 millimetres. The width of the fang becomes reduced to 
between 3 and 4 millimetres. In a second specimen the crown 
presents the sume characters, only that it is flatter on the cinguloid 
side. A third specimen has the bevellings on the attached side of 
larger extent, so as to reach further down the tooth; but all have 
the crown perfectly smooth, without the slightest trace of serrations 
on either side. It is quite possible that the bevellings may be 
produced by wear, though there is nothing to indicate such an ex- 
planation. 

Then there are two teeth very similar in character, only rather 
broader in the crown, being fully 8 millimetres wide. These spe- 
cimens want the bevellings, but have the inner side of the tooth 
marked, with a narrow middle surface which may be flat or con- 
cave, external to which the tooth is bevelled vertically on one side 
and has a thickening at the base of the crown on the other. The 


Q. a G. S. No. 148. » U 


642 PROF. H. G. SEELEY ON THE REPTILE 


base of the cinguloid thickening on the opposite side is convex in 
the middle and concave at the sides. Both these teeth are marked 
with slight rough ridges (Pl. XXVII. figs. 15, 16), which are not 
continuous to the cutting-edge, and are vertical, and much more 
marked on one tooth than on the other. Of the two other speci- 
mens which have the attached side of the crown flat, one, though 
but badly preserved, is remarkable for showing a few faint and 
vertical serrations, which are equally marked on both sides (fig. 12). 
They did not exceed five in number on each of the cutting-edges, 
though only one of these is preserved. All these teeth, I suppose, 
may belorg to one species. 

There remain two other teeth, which, perhaps, may belong to a 
second species or may be worn down. They are characterized 
by the same general features as those already described, but had 
the crown remarkably low, relatively broad, and hardly making 
any approach to a triangular form. ‘The tooth is very thick at 
the base of the crown; and the cinguloid thickening extends along 
both sides. ‘The crown is smooth, and shows no trace of serration. 

I am inclined to refer these teeth to Cratcwomus; they pro- 
bably belong to the species described. 


Craraomus Pawrtowirscuu, Seeley. 
Vertebral Column. 


The vertebral column which I refer to Cratewomus is chiefly re- 
presented by the tail, of which there are about eighteen vertebrae 
preserved ; and the series is very imperfect. ‘There are slight differ- 
ences of mineralization in these specimens, some being red, others 
brownish, and some nearly black; and there are slight differences in 
preservation, since some have the articular margins of the vertebre 
rubbed away, and the processes more or less broken, and others 
are better preserved but somewhat crushed. Still, when the series 
is arranged in sequence there is a perfect continuity of character 
and no evidence to suggest that the remains belong to more than 
one species, or indeed that they may not all have pertained to a 
single individual. A curious feature, also observed in some of the 
English Cretaceous Dinosaurs, is the circumstance that these caudal 
vertebrae scarcely vary in absolute length, though the centrums 
diminish in size. Hitherto no trace of the sacrum has been found. 
The dorsal region is represented by two vertebree, which show the 
forms of the processes ; while the cervical region is represented by 
a vertebra from the hinder part of the neck. In the absence of 
evidence of another vertebral column, it may be legitimate to refer 
these vertebre to the same species as the tail; and from the 
similarity of size it is not unlikly that the whole of these vertebral 
remains are the spoils of a single animal, the Crateomus Pawlo- 
wrtschia. 

Cervical Vertebra. 
(See Binzel, pl. ii. figs. 9, 10.) 


The centrum and neural arch are both preserved; but the neural 


FAUNA OF THE GOSAU FORMATION. 643 


spine and transverse processes are broken away. The centrum has 
the articular faces somewhat oblique; but though this may be to 
some extent natural, it is probably augmented by crushing, since the 
form of the centrum has become in this way a good deal distorted. 
Its length along the base 1 a) 2,2, inches, while the measurement along 
the neural zone: is about ~ inch less. The posterior articular face, 
as preserv ed, is ee 13 inch deep, and about 15% inch wide. 
It is considerably excavated by a saucer-shaped depression. The an- 
terior articulation was probably as deep, and 1,5 inch wide; but it 
does not appear to have been so deeply excavated as the posterior 
face. The base appears to have been flattened, and margined on 
page side by an angular ridge. In the middle these ridges are about 

5, inch apart ; and they diver ge towards both anterior and posterior 
PE The sides of the centrum are distinctly defined from the 
neural arch by the deeply marked horizontal suture, below which 
in front is the oblong articular face for the ED; which is about 
za inch long and z ce deep. It is about 74 nen behind the 
articular ace It rises as a slight pedicle ; and the transverse mea- 
surement over these parapophyses is 2,1, inches. The centrum is 
compressed from side to side below these } processes, so that a median 
cavity divides the side into a highly convex upper portion and a 
comparatively flat lower portion. The articular margin of the cen- 
trum is moderately sharp, thickened, and rounded. The neural 
arch has an aspect of leaning forward obliquely, which is more 
marked than that of the centrum, and may probably be taken as 
evidence that the neck of the animal was carried in a somewhat 
raised position. The pedicles lean forward at an angle of nearly 
45°, have their anterior margins concave, and are compressed from 
side to side, but especially pinched in the middle. The greatest 
width of the arch in front at its union with the centrum is 1,6 inch, 
while behind and above the middle of the centrum its w vidth is 
reduced to 4% inch, again to become expanded to 13 inch near 
the posterior articulation. This median depression extends up to 
the side of the neural arch, being margined above and behind by a 
sharp buttress, which widens laterally and extends outward so as 
to underprop the transverse process, and form with it the upper 
head for the rib. Where fractured this process is 1 inch above the 
capitular seecuuation, and has a triangular outline pointed in front 
and about ;4, inch deep. There is a triangular area which is concavely 
excavated behind these transverse processes and in front of the poste- 
rior zygapophyses. The posterior zygapophyses are div ed from each 
other throughout their length of an inch by a notch 54, inch wide 
behind and rather wider in front, where it terminates in the vertical 
wall of the neural spine, which in the middle has a slight sharp 
ridge. These processes have their inner sides subparallel, are placed 
obliquely, and are convex superiorly from below outward. ‘The 
articular facets are large, subovate, flat, and look downward and out- 
ward soas to make with each other an angle which is more than a 
right angle. The anterior zygapophyses extend entirely in front of the 
articular face of the centrum. They are similarly divided Ag 

au 2 


644 PROF, H. G. SEELEY ON THE REPTILE 


to a level with the centrum, and are thick strong processes which 
have the articular faces somewhat rounded and convex, as though 
to allow of considerable play. Behind the facets the bone is a good 
deal compressed, so as to be concave in length and concave from side 
to side. The base of the neural spine, as preserved, is a square 
pillar, rather more than 2 inch in diameter, which appears to rise 
vertically. In front of this and between the transverse processes is 
a deep excavation about 2 inch long and wide. The neural canal is 
large, and formed almost entirely by the neural arch, the neura- 
pophyses converging so as to almost unite in the middle line of the 
base of the neural cane The width of the neural canal is greatest in 
front, where it is 58, inch; and its height is greatest behind, where 
it is about the sen, or a little more, the canal being depressed in 
front and compressed posteriorly, the width of the canal behind 
being +2 inch. The height from the base of the centrum to the 
upper surface of the posterior zygapophyses 18 37> 1_ inch; the width 
over the outer margins of the posterior zyg gapophy ses is 1 inch ; 
the length, from posterior to anterior zygapophyses is 34, inch; the 
width over the anterior zygapophyses was about 2>4 inch. 


Dorsal Vertebre. 
(See Bunzel, pl. 11. figs. 1-3, pl. vii. fig. 24.) 


Of the two dorsal vertebre the more anterior has the lower half 
of the centrum badly preserved,:but shows the anterior zygapo- 
physis, transverse process, and neural:spine completely. In this 
the transverse processes extend more horizontally outward, while in 
the later vertebra they are directed more obliquely upward To begin 
with the latter (Pl. XXX. fig. 3), the centrum is 2,4, inches long, and 
has the anterior face subquadrate, 2,1, inches deep, “and 2,5 wide. It 
is moderately concave. The posterior face is badly preserved a the 
margin, but appears to have been much smaller, since it is 147 inch 
deep, and, as preserved, is rather wider. It has a deep pit just below 
the eat canal, while the remainder of the face is convex from above 
downward, slightly convex from side to side, and smooth. The 
neural canal is 2 inches long. The base of the centrum is flattened, 
margined by rounded lateral ridges. The upper parts of the sides of 
the centrum are concavely compressed, as though squeezed with the 
finger and thumb; and here, in the middle of the centrum, the trans- 
verse measurementis | inch a little below the neural canal. The neural 
arch is lofty, the height to the origin of the transverse process from the 
base of the centrum being 3,8, inches, and from its base 2,2, inches. 
The buttress which supports the transverse process is flattened at 
the side, since it is formed by pillars which arise from the anterior 
and posterior margins of the centrum, and converge as they ascend 
so as to form a sharp angular ridge beneath the transverse process, 
which is flattened and expanded above. There is a deep excavation 
in front of the vertical A-shaped masses which support the trans- 
verse processes ; and these were placed behind the anterior zyga- 
pophyses. There are much larger but similar posterior excayations, 
which are subtriangular and in front of the posterior zygapophyses, 


FAUNA OF THE GOSAU FORMATION, 645 


which were divided from each other, and looked obliquely outward 
and downward so as to form with each other an angle which was 
much less than aright angle. The neural spine is compressed from 
side to side, and originates from a base about 1,4, inch long. The 

angle enclosed superiorly by the diverging transverse processes 1s 
more than a right angle. The neural canal is subquadrate in front, 
and about +7 inch wide, Posteriorly its height becomes 1,3,, and 
its width about ;° inch. The neural arch has the aspect of being 
placed vertically on the centrum rather towards its anterior 
part. 

In the other dorsal vertebra the height from the base of the neural 
arch to the top of the neural spine is 3; inches. ‘T'he neural spine 
is greatly compressed from side to side, be about 15 inch above the 
platform of the transverse processes, is 52; inch in its greatest poste- 
rior thickness, has an antero- oe “measurement, of 1, inch, 
and swells out at its free end to a width of morethan 3 inch. This 
inflated mass is convex from side to side, and tapers forward in a 
wedge. The platiorms of the transverse processes, which are flat 
above and triangular im section, are given out horizontally ; the one 
preserved measures 243 inches from the median line to its free end, 
which is compressed from above downward, and is rounded from 
back to front. The base of this process occupied the whole space 
between the posterior and anterior zygapophyses. Its anterior 
margin is slightly concave, and terminates in a sharp thin edge. 
The posterior side is similarly thin ; but its hinder part is somewhat 
broken. ‘The width towards the free end is 1Z inch. On the under- 
side is the usual strong median buttress compressed from side to 
side, and terminating tonne in an ovate articular facet for the 
rib, which is 1 inch long, looks downward and is placed towards 
the anterior margin; while it terminates inward abruptly on the 
neural arch above in a nearly circular facet, which is large, vertical, 
slightly concave, and gave attachment to the head of the rib; half 
the facet is above the neural canal. There is the usual superior 
concave excavation in front, behind the zygapophyses, while poste- 
riorly the concavity which runs along the posterior side of the 
transverse process below terminates a an enlarged excavation ; 
and these excavations approximate so as to be separated only by a 
sharp vertical ridge which is placed above the neural canal, with which 
its outline helps to form an §-shaped curve on the right posterior 
aspect. ‘The zygapophyses present no peculiarities, the facets being 
flat and oblique ; the anterior excavation between the transverse pro- 
cesses in front is small; and the interspace between the two facets for 
the rib on the sides of the neural arch is about 1} inch. What 
remains of the posterior face of the centrum appears to be slightly 
concave from side to side, and slightly convex from above down- 
ward, though this conditicn has probably resulted from compression. 
There is here no sharp line dividing the neural arch from the cen- 
trum, though the separation can be easily traced, and it is at the 
middle point of the suture that the compression is greatest; there 
the transverse measurement is less than Linch. The lower portion 


646 PROF. H. G. SEELEY ON THE REPTILE 


of Biinzel’s figure of this vertebra is the centrum; the transverse 
process is lettered d. 


Caudal Vertebre. _ 
(See Biinzel, pl. ii. figs. 4-8, pl. iv. figs. 6-9, pl. viii. figs. 1, 7, 8, 16.) 


There are eighteen caudal vertebre preserved. The earlier ones have 
strong transverse processes, which, however, are more or less broken 
away, are compressed from above downward, and appear to have been 
short ; and the vertebre differ from each other in passing backward in 
the suppression of these transverse processes, which become represented 
by sharp ridges in the middle part of the series, while towards the end 
of the tail all trace of their existence is lost, and the centrum, which 
has become gradually reduced in vertical and transverse measurements, 
assumes a constricted or dicebox-like outline. None of the caudal 
vertebra, except the earliest, appear to have possessed a prominent 
neural spine; for the neural arch has well-developed zygapophyses 
and a concave outline from front to back. The arch, however, soon 
becomes reduced in size, and in the middle of the series is greatly 
compressed from side to side, and the articular zygapophysial facets 
are lost, while towards the end of the tail the neural arch is a mere 
rudiment. The chevron bones were at first apparently large, and arti- 
culated with large oblique facets at the hinder margin of the base of 
- the centrum; but these facets do not appear to have been quite distinct 
from each other, though they were partly divided by the median groove 
on the base of the centrum. They soon become relatively small, and 
near the end of the tail are quite separated from each other, though 
(it may be by an injury received during life) they appear to have become 
united to the centrum. Two of the hinder caudal vertebrae are 
fractured through the centrum, and show the bones to have contained 
central hollow spaces, which, however, were not clearly defined by a 
smooth bony lining, but are rather like the medullary cavities of the 
long bones of mammals. The articular edges, where preserved, are at 
first somewnat rounded, but terminate in a sharp outer margin. 
They are very slightly concave, and later on in the series become 
almost flat, showing that the tail possessed but little flexibility. 

The earliest vertebra preserved has the centrum leaning slightly 
forward. It is fully 1; inch long, and the same depth to the 
chevron facets on the hinder basal margin, which, however, is badly 
preserved. The width of the centrum in front, at the base of the 
transverse processes, is 1,J, inch. ‘The corresponding width behind 
is a trifle less. The anterior articular faco is nearly flat, but had 
the margin rounded. ‘The posterior articular face is more concave, 
and the rounding of the margin is less marked. The upper borders 
of the transverse processes are on a level with the base of the 
neural canal. They are placed nearer to the anterior than to the 
posterior articular face, are transversely oblong where broken close 
to the centrum, and measure ;°, inch in length, ;4 in depth. Be- 
low them the centrum is compressed ; the base is broad, ill defined, 
1 inch in width, rounds into the sides, and is divided longitudinally 


by a shallow groove about ;3, inch in width, which is most marked 


FAUNA OF THE GOSAU FORMATION, 647 


posteriorly. The neural canal is high and narrow: the sides of the 
neural arch converge upward ; and the anterior zygapophyses have 
the facets looking inward; they are concave in depth. 

The second and third vertebre of the series only differ in having 
slight tubercles adjoining the anterior and posterior articular margins 
on the middle of the sides, and in the decreasing dimensions of the 
centrum and processes, though the length still remains the same. 
The fourth vertebra is distorted by vertical compression, and the 
fifth by lateral compression. It, however, has the neural arch well 
preserved, and shows the length from the anterior to the posterior 
zygapophyses to be 213 inches. There is also an indication of a slight 
neural spine broken away, which rose above the posterior zygapo- 
physes. The greater part of both anterior and posterior facets pro- 
jects beyond the centrum. The anterior pair of facets is divided 
from each other ; but there is only a slight notch at the hinder ex- 
tremity of the posterior pair. In the sixth vertebra the neural 
arch is seen to taper posteriorly, when seen from above, in a trian- 
gular outline slightly compressed in the middle; and in the seventh, 
in which the centrum is 1,8, inch long, 1,4, inch deep posteriorly, 
and slightly wider, the neural arch is 2,4, inches long. There is a 
distinct concave compression below the anterior zygapophyses, from 
which faint ridges extend backward longitudinally towards the pos- 
terior zygapophyses. ‘The facet from which the transverse process 
has come away is still ovate, about ;4 inch long, and is placed in the 
middle of the side of the centrum, just below the neural arch. The 
neural spine is seen to be a slight sharp ridge. The anterior zyga-_ 
pophyses are 41 inch apart, while the posterior zygapophyses, which 
have smaller facets, are 54, inch long. The groove on the base of the 
centrum has become somewhat narrower and more sharply defined. 
Here several vertebrae appear to be missing ; and in the next of the 
series the transverse process has become much smaller, is placed lower 
on the side of the centrum, is margined by a vascular groove in front, 
and is prolonged backward by a sharp ridge towards the articular 
margin. The vertebre now begin to elongate a little ; and the ninth 
of the caudal series is 1,9, inch long; the transverse processes have 
disappeared, and are only marked by a sharp median ridge in the 
middle of the centrum, margined in front by an oblique vascular 
groove. Above these lateral ridges the centrum is compressed from 
side to side; the basal groove has become much shallower and 
best marked towards the extremities. The tenth and eleventh show 
the neural arch to be greatly compressed from side to side, and to 
rise very much higher behind than in front, owing to the greatly 
diminished size of the anterior zygapophyses. The posterior zyga- 
pophyses have disappeared; and the centrum is a good deal constricted 
in the middle. The twelfth centrum shows a much greater reduc- 
tion in size of the neural arch, which leaves the posterior third of the 
centrum free. The underside of the centrum is similarly compressed 
to the upper part, though the median basal ridges become rounded. 
The facets for the zygapophyses are distinctly marked at both ends, 
and divided by a groove, of which there is no trace in the middle of 


648 PROF. H. G. SEELEY ON THE REPTILE 


the base. In the thirteenth and fourteenth vertebra the centrum 
is 1,5 inch long, with the articular face 1,4, inch wide in front and 
rather less behind, while the depth in front is 1 inch. The neural 
arch is small, andits superior outline horizontal. The measurement 
from the middle of the base of the centrum to its upper border is 
1, inch. The neural canal indents the centrum concavely at both 
back and front articulations ; and from the hinder limit of the neural 
arch a vascular groove impresses the sides, descending slightly for- 
ward to the middle of the base. The centrum here is most con- 
stricted, and measures 55, inch from side to side, and has a rather 
less depth. The chevron bones are preserved in the thirteenth ver- 
tebra, with the posterior margin of which they appear to be blended; 
they unite below in a Y-shape, and have facets for the succeeding 
vertebra. They are $2 inch wide, $} inch deep, and, with the 
groove at the terminal end of the centrum, enclose the vascular 
canal. Of the fifteenth vertebra only half the centrum is pre- 
served. ‘The sixteenth and seventeenth are blended together, and 
the chevron may not have united in the median line. The vertebree 
now become roughened with many slight longitudinal muscular 
ridges, indicative of the near approach of the end of the series; and 
the articular faces of the centrum appear to have central con- 
cavities. 


Ribs. 
(See Bunzel, pl. iii. figs. 5, 6, pl. viii. figs. 14, 15, pl. 1. fig. 27.) 


One cervical rib is preserved ; there are about half a dozen tolerably | 
perfect dorsal ribs, and a multitude of fragments of dorsal ribs. The 
majority of these obviously belong to one animal; and I refer them 
to the smaller of the two large Dinosaurs, Crateomus lepidophorus ; 
but there are a few slightly larger fragments which possibly pertain 
to the larger Dinosaur. They are, however, too imperfect to yield 
any characters for description; and as they are doubtful remains, I 
prefer to leave their elucidation to future discoveries. They are 
larger and stouter than the bulk of the specimens. ‘The only ex- 
ample which shows the proximal end is represented in Pl. XXX1. 
fig, 12, and may be compared with the corresponding part of the 
smaller rib, fig. 17, Pl. XX VII. This proximal end is the speci- 
men figured by Biinzel as the lower jaw of a lizard, a determination 
which is presuniwviy due to its imperfect condition and the cireum- 
stance that the transverse platform is only developed on one side. 
I owe its identification to Mr. Hulke, who recognized its resemblance 
to the smaller specimens when examining the collection. 

The cervical rib wants the articular head of the lower tubercle. 
The interspace between these two heads was nearly 13 inch, the 
outline between them being deeply concave; the ventral outline is 
flat, the dorsal outline concave. The length of the rib, as preserved, 
is 3,2, inches. After being directed outward for half its length, it 
curves concayely backward and tapers at the same time. There is 
in front, in the middle of the rib, a slight ridge. The articular head 


which is preserved is 58; inch long and ,% inch wide. When perfect 


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marine Sok 15 


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FAUNA OF THE GOSAU FORMATION, 649 


the rib must have had a Y-shaped form ; and just below the fork 
of the Y on the posterior side is a moderately large vascular 
foramen. 

The dorsal ribs are all imperfect at the distal end; and some of them 
have a much greater curve than others. The longest fragments are 
imperfect, proximally and distally, and measure round the outside 
curve about. 1] inches, and across the chord a little over 10 inches. 
But all the ribs agree in distinctive character, which is seen in side- 
to-side compression along the proximal half of the visceral surface 
and lateral expansion on the dorsal surface (Pl. AXVII. fig. 17), 
so that in this haif of the rib the transverse section is shaped 
like a 7 (Pl. XXVIII. fig. 18); but distally the rib expands 
on the visceral surface, and the elevated ridge on the ante- 
rior border disappears, the direction of the rib becomes some- 
what oblique, and its section has a compressed ovate outline. One 
consequence of this remarkable dorsal expansion is to form an ex- 
panded table-like external surface which is convex in the direction 
of the length of the rib, and nearly flat in the antero-posterior 
direction ; so that the ribs closely resemble the combined rib and 
costal plate of a Chelonian, and in the living animal the interspaces 
between the ribs must have been as small as in many mammals, such 
as the Buffalo and the Lesser Anteater. It is, of course, possible that 
this expansion of the dorsal margin of the rib may be homologous 
with the costal plates of Chelonians, since representatives of the 
structure are also met with in Crocodiles, Hatteria, and birds. I 
have no doubt that this table structure carried the heavy dermal ar- 
mour with which these animals were weighted. One specimen (fig. 17), 
which has the articular head preserved, has an interspace of 13, inch 
between the capitulum and tuberculum. The capitulum curves 
slightly forward, is $+ inch in depth, and gradually widens on the 
ventral surface towards the articulation, where it is 4 inch in dia- 
meter. The tuberculum is relatively small, } inch from back to front, 
and about 4 inch in width. It is somewhat reniform and rounded, 
like the outside of a kidney. Immediately beyond the tubercle the 
bone begins to widen ; and the anterior ridge extends in the speci- 
men 4 inches, while in other ribs it extends 6 inches, and in 
some only 3. Its antero-posterior width is also variable, but, 
where widest, in no specimen measures more than | inch, and is 
usually about #inch. The compression of the rib just below the 
tubercle gives a measurement of less than ;%, inch, though in some 
specimens it may be a little more and in others less ; while the depth 
of the rib from dorsal to ventral surface is at first ;?, ich, and be- 
comes gradually reduced as the rib extends and loses its T-shaped 
section. ‘The side-to-side compression extends close under the ex- 
ternal platform, so that both sides of the rib are concavely chan- 
nelled. Several specimens show some amount of muscular roughness 
on the transverse platform and a part of the rib distad to its termina- 
tion ; and this is probably correlated with the muscular attachment of 
dermalarmour. The longest ribs preserved do not indicate a greater 
length when complete than 14 inches. When they become obliquely 


650 PROF. H. G. SEELEY ON THE REPTILE 


flattened they maintain a remarkable uniformity of width, and taper 
almost imperceptibly towards the distal extremity. 


Dermal Armour. 
(See Btinzel, pl. iv. figs. 1, 2, pl. vii. figs. 20, 21, pl. vin. figs. 9-12.) 

The Dinosaurian dermal armour which I refer to the genus 
Cratwomus presents many remarkable modifications, such as have 
not been met with in any genus hitherto described. Some of the 
plates are remarkably similar to those of Scelidosaurus and <Acan- 
thophols; others are large scutes with a median longitudinal ridge 
and numerous vascular impressions on the carinate surface, as 
though they were imbedded in the skin. These plates are all 
thin and may have been abdominal, while the more elevated plates 
may have been dorsal and caudal. A third type of plate appears 
to be greatly compressed from side to side with a sharp cutting 
surface in front, terminating in a spike superiorly, and with a 
rounded posterior margin. The articular bases of these plates are 
not preserved. A fourth kind of plate of large size appears to have 
terminated at each end in a great triangular spike, while across the 
intermediate space there extended rows of conical tubercles some- 
what resembling in outline those attributed to Hyleosaurus. A fifth 
kind of armour is represented by an immense conical spine, like the 
horn-core of an ox, which rises from a bony base. 

None of these pieces of armour are symmetrical, hardly any of them 
can be grouped in pairs ; altogether there are fully fifty well-defined 
plates, besides a large number of fragments. It is quite possible 
that the remains may have belonged to more than one species. But 
seeing that the vertebral column of one species is well preserved in its 
hin dey portion, andthatto this species the bulk of the limb bones may 
reasonably be ‘relegated, and that in many points of osteology there 
is an approximation of the animal towards Scelidosaurus, we may 
be justified in considering that the larger Austrian Dinosaur possessed 
armour as varied in character as that seen in its English prototype ; 
and in the absence of a second and larger vertebral column, I am 
unwilling to attempt to divide the scutes between the two different 
animals, merely on the ground of their contours. Yet it may be 
acknowledged that the horncore-like scute is larger than would 
have been expected, and that the whole armour is heavy even 
for an animal with such strongly marked muscular development as 
is shown on the bones of the limbs of both the species. 

I proceed to describe the armour according to the varieties it 
presents. As Prof. Suess had noticed, there is, besides the horn- 
like scute, a second base, from which the horn-like spine has been 
broken away. There thus appear to have been at least a pair. 
The base from which the horn rises (Pl. XX VIII. fig. 4) 1s 55 inches 
long, of irregular oblong shape with roughened edge, a little 
oroken at one end. It is nearly 33 inches wide where widest, 
and narrows to about 23 inches. It is concave in length on the 
under surface in the middle, convex at the sides. The margin is 
full of vascular perforations, and appears to have had strong union 


FAUNA OF THE GOSAU FORMATION. 651 


with the skin. On the broad side this bony base is 1,8, inch thick ; 
on the narrow side the greatest thickness is 1,4, inch. The horn- 
like spine is placed obliquely upon it, and rises vertically, curving a 
httle backward or outward. It is 53 inches high. Its base is ra- 
ther more than 23 inches long, and 2,1) inches wide. ‘The spine is 
slightly flattened on the convex and concave sides. ‘The right 
and left sides are nearly straight. The bone is covered with close- 
set irregular vascular perforations similar to those on a horn-core. 
(See Biunzel, pl. v. fig. 10.) 

Two other plates of quite as remarkable character are dissimilar in 
form, one being twice the width of the other; but both had smooth 
bases for attachment to the skin. The base is rounded at its lateral 
margins, as though it were a bone distinct trom the extraordinary der- 
mal ornament which rises fromit. The larger specimen (PI. XXVIII. 
fig. 2) is 8 inches long and imperfect at one end. The articular base 
appears to have been about 43 inches long and 2 inches wide, while 
the greatest width of the plate is 3} inches towards each end of the 
articular space, and in the intermediate area it becomes contracted 
to about 24 inches. The spine which existed at the other end of this 
contracted area has been almost entirely broken away ; so that the 
plate was originally probably a central oblong mass with constricted 
sides terminating at each end in a large triangular spine, which was 
directed upward from the body of the plate. The one spine which 
is preserved is on its upper surface about 4 inches long, and at 
the base 3z inches wide. It is slightly convex from side to side, 
and terminates in a sharp cutting-edge on each side, which is lon- 
ger and more convex on one side than on the other; and the longer 
edge is reflected a little upward. There are a few longitudinal sub- 
parallel vascular grooves in the middle of this part of the plate. On 
the under surface of this region the bone is flattened on the two 
sides, which converge towards a rounded ridge in the middle line, 
which helps to give strength to the sharp dagger-like extremity in 
which the bone terminates. The greatest thickness of this part 
of the plate in the middle is 1/ inch. Its base terminates 
abruptly, perhaps owing to some crushing on the underside. The 
middle oblong portion of the plate is studded over with conical tu- 
bercles, the bases of which are pretty clearly defined, and the cones 
are low. ‘They are arranged across the bone in three rows with 
four low conical tubercles in each of the two outer rows and 
two larger tubercles in the middle, 1 inch in diameter, with three 
on the external margin of each, making in all 6 in the middle. 
These tubercles make an elevated border abutting against the tri- 
angular spine. Their surfaces are roughened with close-set irre- 
gular vascular punctures. The smaller plate (Pl. XXVIII. fig. 3), 
of similar character, is rather better preserved, its total length 
64 inches, length of the articular base 54 inches. It carries a 
vertically elevated spine, and the base beneath this is deeply con- 
eave. The margins of the base are smooth and well rounded as 
already described in the larger specimen. The width of the base 
is 13,inch. At one end the compressed spine rises at an angle 


652 PROF. H. G. SEELEY ON THE REPTILE 


from the part of the plate on which it is situate; it is somewhat 
fractured ; but its height as preserved is nearly 23 inches, and the 
length of its base rather less; it terminates towards the free extre- 
mity in a sharp cutting-edge. Its thickness in the middle of the base 
is 5% inch; and it tapers upward and outward towards both margins. 
It is defined at the base by a constriction which appears to separate 
it from the plate from which it rises.* It is scored with somewhat 
irregular vertical vascular furrows. The corresponding plate at the 
other end is much smaller, and is defined from the under articular 
surface by a furrow; and a similar furrow appears to mark its mit 
on the upper surface, as though it did not completely cover the bone 
upon which it rests. It is of ovate outline, 2,4, inches long and 1} 
inch wide in the middle. Its surface is undulating, as though the 
free extremity, growing against another plate, had been forced up 
into an elevation. It has the aspect of projecting on one side be- 
yond the bone on which it rests, and is then sharply compressed, and 
terminates in a cutting-margin which is convex in length. The in- 
terspace between these terminal plates is rhomboidal, about 1,% inch 
in length, and is covered with conical tubercles, the largest of which 
is +5 inch long and about 4 inch high. These tubercles are about 
© in number, the 3 largest being on one side. 

The next series of dermal bones are all longitudinally carinate. 
They may, perhaps, be divided into such as have the base angularly 
excavated, as though they were median bones of the dorsal or 
caudal region, and such as have the base comparatively flattened ; 
and in these latter the keel becomes greatly reduced in height: 
these bones are probably lateral. Judging from the example of 
Stagonolepis, I am inclined to believe that most of these plates 
pertain to the tail. There are four plates, each about 2%, inches 
long, with an ovate base having a rough margin, rising into a sharp 
cutting median keel about 2 inches in height, which has a ver- 
tical sharp margin behind and a convex margin over the length 
of the plate (Pl. XXX. fig. 2). The sides of these plates are con- 
cave from above downward, and convex in length; but they are 
all somewhat distorted by pressure. They thin away at the free 
margin to about ;4 inch in thickness. our other plates, also an- 
gular on the underside, are much more elongated, and clearly over- 
lap each other at one end, which may be presumed to be posterior. 
The largest of these plates is 6? inches long, 14% inch wide where 
widest behind, and 24 inches high in the highest part of the sharp 
compressed keel. One side of this plate is moderately concave from 
above downward; the other side is plano-convex;. and posteriorly the 
underpart of the bone has the aspect of being obliquely truncated— 
a character which results from the posterior 23 inches rising free 
from the basal attachment so as to terminate in an upward and 
backwardly directed spine, which overlapped the next succeeding 
plate. The crest of the median ridge has a very slight sigmoid 
flexure. Attached to this bone on one side is a small fragment 
which appears to be a broken portion of the proximal end of the 
dorsal rib. Other plates are somewhat flatter and relatively broader ; 


FAUNA OF THE GOSAU FORMATION. 653 


one which measures nearly 5 inches in length has a subrhomboidal 
outline, two long sides converging in front, and a short pair of sides 
converging behind. The greatest width of the plate is 23 inches. 
The greatest length of the flat part of the base is 32 inches ; and 
the posterior 14 inch rises into a strong spine, which terminates the 
median crest, is 1,%, inch high, compressed behind and above. The 
crest gradually diminishes in height from this spine forward till it 
dies away at the anterior end. The outline of the crest is very 
slightly sigmoid. The crest has a compressed aspect, as though it 
had been naturally squeezed from side to side in its upper half. 

There are numerous smaller sharply carinate plates of a some- 
what ovate outline, with the keel placed nearer towards one margin 
than the other, and always becoming a little more elevated towards 
one end, where it is truncated. And these plates, though mostly flat 
on the underside, always have the end on which the ridge is highest 
bent a little upward, as though to overlap the next succeeding plate. 
These plates vary in size: one is 2,8, inches long, 1,5, inch wide, 
and has the keel $4 inch high ; another is 2-3, inches long, 1,4, inch 
wide, and has the keel 55, inch high posteriorly. 

Another remarkable series of plates is distinguished by extreme 
thinness. They appear all to have been subrhomboidal and to have 
had the keel scarcely elevated. 

The largest is about 2,5 inches long, and more than 2,%, inches 
wide. The under surface is smooth and slightly convex. The su- 
perior and inferior margins converge to a sharp but irregular edge ; 
the thickness of the body of the plate is about 4, inch, though many 
of the plates are much thinner; and the thickness in the line of the 
median ridge is about ;#; inch. This slight keel does not extend to 
either extremity of the plate; but the margin of each plate is turned 
up towards one of the posterior sides, as though they still obliquely 
overlapped (Pl. XXXI. fig. 3). The surface of these plates is 
slightly concave on each side of the median ridge, and there scored 
with vascular markings which ascend towards the ridge and ramify 
and interlace. ‘Their prevailing direction in the plates with more 
elevated keels is towards the posterior spine. There are a few 
slightly thicker plates which have no trace of keel, but are flat 
below and gently convex above (Pl. XXVIII. fig. 5), with a deep 
Y-shaped vascular groove on each, and a sharp margin. The 
smallest and best-preserved is 1,4 inch long, and 1,1; inch wide. 

Some fragments of crest-spines, broken away from the bases, in- 
dicate plates of a larger size than any thing here described. The 
plates appear to have been remarkable for their great side-to-side com- 
pression, the posterior elevation of the crest, and the sharpness of 
the spine, which, in fragments preserved, extended to a height of 5 
inches where the antero-posterior measurement is only about 3 
inches, and the greatest thickness of the spine from side to side is only 
42 inch (at the interior fracture). 

There is also a fragment indicating that the plates in which the keel 
is almost suppressed, in some regions attained a larger size than has 
here been described. 


654 PROF. H. G. SEELEY ON THE REPTILE 


One such fragment as preserved is about 5 inches long and rather 

wider, with apparently two slight keel-like ridges parallel to each 
other. The greatest height of the crest in this specimen is about 
45 inch. 
* If all this armour is correctly referred to the genus Cratwomus, 
it furnishes one of the most distinctive generic characters of this 
type. I do not remember any described genus in which large 
tubercled plates such as are here figured have been found, though 
an isolated plate was described from the Wealden of the Isle of 
Wight many years ago* as showing a not dissimilar ornament. 
Other plates are so similar to armour of Scelidosaurus, especially the 
median-keeled caudal plates, as to enable us to concur with Biinzel 
in recognizing a strong affinity to that genus, which, however, does 
not amount to identity. Cratewomus was more heavily armoured. 
It is difficult to say whether its armour has more in common 
with the bony tubercles which occur on the limbs and tail of many 
Chelonians, or approximates better to the bony scutes of certain 
lizards and crocodiles; for it is so distinct that no near parallel 
can be drawn between the armour of Dinosaurs and that of living 
reptiles; nor if the comparison were possible would it have much 
weight as a mark of organic affinity. 


Scapule. 


Three specimens of Dinosaurian scapulex have been obtained ; two 
are larger than the other, and belong to a distinct species. The two 
larger specimens were figured by Bunzel, and regarded by him as 
left ribs of his imaginary Lacertilian genus Danubiosaurus and the 
type of his species D. anceps. ‘They are left scapule. The smaller 
specimen obtained subsequently is a right scapula. I refer the. 
larger bones to the animal indicated by the larger limb-bones (Cra- 
teomus Pawlowitschir), though, as the smaller specimen is little more 
than half the size, the disproportion in the scapule is much greater 
than would have been anticipated in the two species. 


Left Scapula. 
(See Bunzel, pl. v. figs. 7-9, pl.vi. figs. 1-3.) 


The scapula (fig. A, p. 656) is remarkable for its great breadth, its 
curved form, its compressed aspect, and the remarkable acromial pro- 
cess in which its slight spine terminates. The specimen has been a 
little crushed, and is not quite perfect at its distal end; the whole 
surface for union with the coracoid is destroyed by decomposition, and 
slightly injured by fracture. What remains of the articular surface 
for the humerus is a semiovate surface 54 centim. wide and rather 
longer. Itis margined by an elevated ridge, has the usual roughness 
of cartilaginous surfaces, andis more concave than usual, both in length 
and breadth. The inner or visceral margin of the bone appears 
to have been more convex than the external margin. Both are 
somewhat inflated; and the external surface and posterior margin 


* J. H. Lee, Ann. & Mag. Nat. Hist. vol. xi. p. 5, reprinted in his ‘ Note 
Book of a Geologist.’ 


FAUNA OF THE GOSAU FORMATION. 655 


especially are roughened with ligamentous attachments and mus- 
cular rugosities. The thickness of the bone rapidly “diminishes 
above the articulation; and it continues to become thinner towards 
the free end, where the thickness does not exceed a centimetre 
and a half. The posterior margin is compressed and rounded, 
the rounding becoming more conspicuous as the surface ap- 
proaches the humeral articulation, owing to the increasing thick- 
ness of the bone. The length of the posterior side of the bone is 
about 25 centim.; its outline is concave; the concavity, as preserved, 
may be indicated by the fact that the chord joining its two ex- 
tremities is 2] centim. long, and the abscissa 6 centim. high. The 
corresponding anterior margin is not quite parallel, since the bone 
is wider in its upper third than in its lower third; and hence the 
‘anterior margin is more convex. ‘The least transverse measurement 
above what may be termed the spine of the scapula is 63 centim. 
In the upper third of the bone the width has increased to upwards 
of 8 centim.; it then contracts a little to less than 8 centim., so as 
to make the anterior termination of the superior margin concave; 
for the bone widens once more, so as to become broader than ever 
at its free end. The posterior corner of the free end appears to 
be curved a little outward. The external surface is smooth, con- 
vex in length, and more convex in breadth at the distal end than 
proximally. The visceral surface exhibits corresponding characters. 
The anterior margin of the bone is thicker, better rounded than 
the posterior margin, and rougher with muscular attachments. As 
preserved, the measurement from acromion to the anterior distal 
margin is 23 centim. in a straight line. In about the middle of 
the anterior margin the bone becomes appreciably thickened on 
the inner side with muscular attachments, and the thickness in- 
ereases until a vertical anterior shoulder is formed almost at right 
angles to the spine of the scapula, extending downward and inward 
towards the coracoid area. Only a small portion of this trian- 
gular space is preserved; but so much of it as is seen below the 
acromion is 4 centim. deep. The spine of the scapula only runs for 
a short distance along the proximal part of its surface, and is diffi- 
cult to define, because the bone is obviously compressed so as to 
make the surface posterior to the spine appear more concave than 
it really was; but the spine may be considered to originate in 
the thickening of the anterior margin of the bone already alluded 
to; and it becomes most distinct a centimetre or two above the 
acromion, where it is 13 centim. wide, flat above, and margined at 
the sides by rounded ridges. It is prolonged into a free process or 
acromion, which was directed forward and outward. This process is 
a little crushed, is nearly 5 centim. long, 33 millim. broad, and, as 
preserved, about 2 centim. thick, though before crushing it was 
thicker ; the corners and angles of its free end are rounded; and 
the inferior or internal surface is concave, since it rises from the 
anterior coracoid border of the bone. The plane of the acromion is 
parallel to that of the blade of the scapula, and makes an angle of 45° 
with the direction of the humeral articular surface. The distance 


656 PROF. H. G. SEELEY ON THE REPTILE 


Scapule of Crateeomus Pawlowitschii and C. lepidophorus. 


Wi 


/ 


A. Left scapula of Crateomus Pawlowitschii. 

B. Right scapula (drawn reversed for comparison) of Crateomus 
lepidophorus. 

1. Humeral articulation. 2. Acromion process. 


FAUNA OF THE GOSAU FORMATION, 657 


of its inferior border from the humeral articular surface is between 
6 and7 centim. From the inferior margin of the acromion descends 
a strong rounded ridge which divides the part of the bone above the 
humeral articulation into anterior and posterior areas, and, but for 
the intervening acromion process, would look like a continuation of 
the spine. This ridge dies away before reaching the humeral arti- 
culation. Posterior to the acromion the surface of the scapula is 
broadly channelled. In its singular curvature and development of 
the acromion it is unlike the scapula of any other Dinosaur. 

‘he second specimen, though evidently belonging to the same spe- 
cies, is a little smaller; it has been reconstructed out of even a larger 
number of fragments than the specimen described. The proximal 
part of the bone is wanting, the fracture having removed the arti- 
cular region and acromion process ; but this portion was compressed 
from side to side; the anterior margin of the bone, however, appears 
to be thicker, and the cavity in front of the acromion is more 
marked, than in the specimen here figured. Distally a portion of the 
terminal free margin of the bone is preserved, showing that it was 
obliquely truncated and somewhat thickened and roughened with 
muscular attachments, especially towards the posterior border; the 
specimen, as preserved, is 27 centim. long and about 76 millim. 
wide in the blade, where it is widest. The details of the two bones 
otherwise present the closest agreement. 


Distal End of Humerus. 


A large humerus is unfortunately only known from its distal end 
(Pl. XXIX. fig. 4), which has decayed in the manner so frequent 
with these fossils, and as though indicating that the terminal end 
had been a distal epiphysis similar to that which characterizes the 
long bones in the order Sauropterygia, which may have separated 
from the shaft absolutely or decayed in consequence of the less per- 
fect ossification of its cartilaginous substance. 

This fragment is little more than 4 inches long. It exhibits at 
the proximal end a natural fracture, made during extraction from 
the rock; and here the bone is 1,8, inch thick, 1,5, inch wide, ovate 
in outline, with an angular bulge towards the middle of the superior 
surface—a bulge which indicates a ridge similar to that referred 
to in the more perfect specimens of the other species (Pl. XXIX. 
fig. 1). I regard this specimen as the distal half of a left humerus 
of Cratwomus Pawlowitschi. 

The fracture is somewhat interesting as showing the existence 
of a central medullary cavity (Pl. X XIX. fig. 5). This cavity is 
= inch long and more than 3 inch wide; so that the bone round 
it is about half an inch thick; and this contrasts remarkably with 
the thinness of the terminal and irregular edges of the distal margin 
of the shaft, which is nowhere much more than ;y inch thick, though 
becoming somewhat thicker as it extends proximally. The cavity 
is relatively smaller in the second species. The superior surface is 
unfortunately somewhat crushed; and the whole specimen has been 
pieced together, like so many of these remains, with great patience, — 


Q.J.G.8. No. 148. 2x 


658 PROF. H. G. SEELEY ON THE REPTILE 


skill, and success. The shaft does not widen quite so rapidly towards 
the distal end as in the smaller species, since the bone is 24 inches 
wide only at 24 inches further towards the distal extremity. The 
surface is remarkably smooth and free from muscular markings, 
except on what I regard as the outer side. This is roughened; and 
rather above its middle there is a chain of strong muscular eminences 
(which appear to be much in the position of the inner straight ridge 
described in Crateomus lepidophorus, in which, however, the ridge 
was only developed proximally). Here it is strong distally, and 
would seem to have extended to the terminal articulation. The 
middle of the inferior aspect is marked by a straight vascular groove 
about an inch long in the upper half of the fragment; and at the 
extreme distal margin towards the inner side there is a vertically 
ovate muscular pit about 1 inch long. The distal extremity appears to 
be curved downward and outward rather more than usual, while the 
surface is more than usually convex. The sides, which are slightly 
concave in length, are comparatively straight. The extreme width, 
as preserved distally is 3 inches; but this is fully 1 inch short of 
what must have been the end of the specimen. 


Femur. 


(See Bunzel, pl. iii. figs. 2-4.) 


The shaft of the right femur which I refer to this species is very 
well preserved ; but there is no trace of the articular extremities, 
which disappeared before the bone was imbedded in the matrix ; 
and towards the articular ends the bone is crushed, on the posterior 
aspect proximally, and at the distal end in front. There is, how- 
ever, quite enough preserved to indicate a very distinct animal from 
that referred to Cratwomus lepidophorus (Pl. XXXI. fig. 5), to 
which, however, it was nearly related in femoral character. The 
fragment (PI. XX XI. fig. 1) is 11 inches long and 1,4 inch wide 
in the lower third of the shaft, is remarkably cylindrical, has the 
muscular ridges on the anterior surface strongly developed, while 
the inner middle trochanter of the shaft would not be recognized as 
such, so feebly is it developed, were it not for the characters of the 
other species. As preserved, the appearance of the bone is remark- 
ably mammalian. When perfect, it may have been 15 inches long. 

The fragment of the corresponding left femur is scarcely at all 
compressed, but was so far destroyed before mineralization that 
only 7 inches of its length now remain, showing the lower half of 
the shaft to be subtriangular in section, being flattened behind and 
somewhat compressed towards the median muscular ridge in front. 

The following description is drawn from the representative of the 
right limb (Pl. XXXI. fig. 1). The bone, which is most constricted 
in the lower third, widens in the usual way towards both proximal 
and distal ends; and its most remarkable feature is the inflation of 
the proximal half of the dorsal or anterior half of the shaft in the 
line of the median longitudinal muscular ridge. This ridge is strong ; 
and its crest is broken into short lengths of from half to three quarters 


FAUNA OF THE GOSAU FORMATION. 659 


of an inch; it originates distally in the part of the shaft which is most 
constricted, runs rather nearer to the external than to the internal 
margin, and increases in strength proximally till it becomes 52, inch 
wide, where the shaft is 3 inches in diameter. All that part of the 
shaft which is external to the ridge is obliquely flattened, with a 
slight increase of inflation towards the proximal end, but without the 
slightest indication of the formation of a proximal trochanter, which 
presumably was not developed, though the absence of this struc- 
ture may be due to mutilation. The inner side of the shaft is rather 
more convex than the outer side ; but on its upper side there curves 
round from where the lateral trochanter should be, a muscular ridge, 
which is rather stronger but less well defined than the principal 
median ridge, towards which it very slowly converges proximally. 
At first the width between the ridges is nearly 14 inch: but at 
3 inches nearer to the proximal end it is narrowed to 1,1, inch; and 
as it narrows, the area thus defined between the muscular ridges, 
which is at first flat, becomes very markedly concave. Proximally 
the bone curves inward as though approaching the terminal arti- 
cular head, and the space external to the inner ridge is fairly well 
rounded. The proximal half of the posterior aspect of the bone is 
somewhat crushed, and appears to have been more convex than 
usual. It may have terminated towards the outer side in a slight 
ridge, and shows but very slight or uncertain indications of the 
posterior muscular ridge seen in the second species. The lateral 
trochanteroid muscular indication is placed a little higher up than 
is usual with the lateral trochanter ; itis about 1} inch long, slightly 
elevated and rounded; its proximal end inclines slightly towards 
the anterior face of the bone. The lateral outline of the bone is 
here markedly convex. 

The distal end of the bone gives no indication of the widening 
on the outer side of the articulation which is so often met with, 
since it is flattened and straight externally. Posteriorly there is 
a moderately deep broad channel inclined a little outward; it was 
evidently prolonged between the condyles, and shows the outer 
condyle, as usual, to have been small, while the inner condyle was 
large. The bone appears to have been quite as much thickened as 
usual at the distal end, though only the backward curve of the shaft 
and no part of the articulation itself is preserved. The whole sur- 
face of the shaft is remarkable for the longitudinal muscular rough- 
nesses, which are more marked than in any reptile bone that I have 
ever seen. 


Tibia. 


A pair of large limb-bones, both 8 inches long, as preserved, but muti- 
lated before fossilization, so that no trace is shown of either proximal 
or distal articulations, present, however, characters which unmista- 
kably show them to be the tibial bones (Pl. XXXT. fig. 2). The frag- 
ments are straight on the inner side. The shaft bends inward a little at 
the distal end, has a long anterior crest immensely developed forward 
at the proximal end, and sends out a compressed process on the outer 

2x 2 


660 PROF. H. G. SEELEY ON THE REPTILE 


side. The bone measures from back to front at the proximal end 38 
inches, while the middle of the shaft measures 18 inch, and the distal 
end 1finch. The whole inner surface is remarkably flattened, and, 
except for the usual distal widening, shows no character that calls 
for remark. ‘The posterior aspect is badly defined, somewhat flat- 
tened towards the distal end, where the bone is rough with muscular 
markings, and 17 inch from side to side. The thickness in the 
middle of the shaft is apparently less, though the bone may be some- 
what compressed ; and the thickness varies in the two specimens. In 
length, the posterior outline is slightly concave in the middle of the 
shaft, and slightly convex in its upper portion, where it is well 
rounded from side toside. The outer surface of the bone is convex 
from back to front, and slightly concave in length. It becomes 
compressed proximally ; so that it is divided by an elevated median 
ridge into two portions——_the posterior somewhat flattened and look- 
ing obliquely outward and backward, while the anterior half is 
deeply concave, the concavity resulting from the natural compres- 
sion of the shaft anteriorly, so as to form an immense patelloid 
crest something after the pattern of that figured by Leidy as cha- 
racterizing Celosaurus. The anterior margin is somewhat sharp, 
and is defined at the distal end by a slight angular ridge, which above 
the middle of the shaft extends inward; so that the proximal por- 
tion of the anterior outline becomes convex from side to side, though 
the side-to-side compression increases, and the bone, where frac- 
tured, is again increasing in width from side to side, and measures 
4 inch. The upper part of the shaft has a subtriangular section, 
owing to the elevation of the external or fibular ridge. Fibule of 
two sizes occur; but the larger specimen is so small that I have 
noticed it under the next species. 


CRATHOMUS LEPIDOPHORUS, Seeley. 


In.grouping together the remains which are now to be described 
I have been influenced partly by their anatomical characters, partly 
by size, and partly by the fact that I have no evidence of the limb- 
bones of a third species of the same genus to which any of the 
bones might be referred. The specific distinctness of this smaller 
Dinosaur will be found well indicated by the characters of the 
scapula, humerus, femur, and vertebra. ‘The armour is probably 
undistinguishable from that of the larger species; and at present 
there is no sufficient ground for saying how much of that already 
described belonged to Cratwomus lepidophorus. 


Left Coracoid. 
(See Bunzel, pl. iv. fig. 3.) 

The left coracoid is very imperfectly preserved, giving no indica- 
tion of the outline of the bone, no trace, or even indication of direc- 
tion, of its union with the scapula, and showing the articular surface 
for the humerus but imperfectly. The bone, however, is perfectly 
recognizable; it has been figured by Biinzel as the right side of 
the ilium of Zguanodon Mantel. Since no other example of a 


FAUNA OF THE GOSAU FORMATION. 661 


Dinosaurian coracoid occurs in this formation, it may be useful to 
record the few indications and characters which it displays. The 
length of the fragment is about 113 centim.; its breadth is 44 
centim. I infer it to have been a bone, however, fashioned on 
the plan of the coracoid of Hyleosaurus or Scelidosaurus. The bone 
thickened considerably towards the articular surface for the humerus, 
where the greatest transverse measurement is 36 millim., though, 
being eroded, this may not have been its widest point. The length 
of the articulation is about 65 millim.; but, from the state of preser- 
vation of the specimen, this can only be given approximately. There 
are some irregularities on the surface which would suggest car- 
tilaginous covering, such as is indicated by the articular end of 
the scapula. In length, the surface is slightly concave. Below 
the articular surface the bone is excavated concayely in length, 
though the excavation is not very deep. This inferior surface is ob- 
liquely compressed on the inner side, so that a slight and rounded 
ridge extends downward from the articular surface on the outer 
margin of the bone. The excavation extends slightly under the 
articular surface. ‘The visceral surface is so eroded as to be almost 
unrecognizable, only one or two patches of unworn bony surface being 
preserved. The external aspect towards the scapular articulation is 
roughened with longitudinal ridges. The middle part of the bone 
appears to be smooth, but carries a row of seven or eight vascular 
pits close to the elevated and compressed margin of the humeral 
articular surface, in front of which is a broad shallow furrow, as 
though the thumb had been drawn over a plastic substance. This 
furrow becomes wider as it extends downward, and at its distal termi- 
nation is margined by shght muscular rugosities. Distant 3 centim. 
from the upper part of the humeral articulation, and rather more, 
apparently, from the margin of the scapula, was the coracoid foramen, 
which was about 15 millim. in length and probably ovate or pear- 
shaped, though its outline is imperfectly preserved. The external 
surface of the bone appears to have been convex both in length and 
breadth. The size of the bone can only be inferred from comparison 
with allied genera. From its imperfect condition I do not feel 
assured that this bone may not belong to the larger species. 


Right Scapula. 


This comparatively small bone, though corresponding in a general 
way with the larger specimens described (see fig. B, p. 656), presents 
remarkable differences, which enforce the conviction that it belongs 
to a very distinct species. The blade of the bone is flat, and presents 
no curvature of plane; its anterior margin is relatively straighter ; the 
acromion was smaller and differently placed, and approached almost 
to the margin of the humeral articulation. As in the other species, 
the surface for union with the coracoid is eroded, though apparently 
to no great extent ; and though the bone is imperfect at the oppo- 
site free end, there is no reason to suppose that it extended appre- 
ciably beyond the part preserved. The extreme length of the spe- 


662 PROF. H. G. SEELEY ON THE REPTILE 


cimen is 184 centim. The length of the posterior margin to the 
humeral articulation is about 15 centim., the length of the chord of 
the arc of the posterior curvature is 134 centim., and the abscissa 
is about 28 millim.; so that the curvature 1s much less than in the 
large species. The posterior margin is also more inflated; it is 
similarly sharp at its distal end; but the bone thickens steadily 
towards the humeral end, where it rapidly expands, chiefly on the 
inner side, to form the humeral articular surface. The posterior 
margin is more rounded on the external than on the internal sur- 
face, giving the effect of an obscure ridge along the visceral border 
of this outline of the bone. The anterior border is also well rounded 
and thicker than the posterior border, the thickness in the middle of 
the blade being about 12 millim. The outline divides itself into a 
proximal part, which is concave, and a distal part, which is straight. 
The middle convexity of the outline is much less pronounced than in 
the larger species. The concavity towards the proximal end is due 
to the prolongation forward of the comparatively thin process for 
union with the coracoid. The middle part of the anterior margin is 
marked with fine parallel muscular ridges; and from this region 
the spine of the scapula is prolonged downward obliquely across the 
bone, so that it terminates at about the middle of the proximal end 
of the bone, which is 9 centim. wide. The least width of the blade in 
the middle of the concavity on the anterior side of the margin is less 
than 5 centim. The width at the origin of the spine of the scapula 
is about 53 millim.; and the width at the distal end, as preserved, is 
63 centim. The spine is remarkably straight; and even the length 
to its acromial termination is nearly 10 centim.; it does not so 
much suggest the form of spine in a mammalian scapula as that of 
Hatteria, existing as a broad rounded ridge, which divides the 
proximal end of the external surface of the bone into two areas, 
which are both concave in length, though the outer subtriangular 
area is rather wider and shallower. The acromial process is imper- 
tectly preserved proximally ; and hence the spine appears to termi- 
nate in a rounded ridge which is about 2 centim. high and ap- 
proaches to within a centim. and a half of the humeral articulation. 
The thickness of the bone from the internal to the external surface 
at the worn or eroded termination of the spine of the scapula is 4 
centim. The width of the process is less than 14 centim.; and the 
measurement from its outer border to the anterior coracoid margin 
is about 5 centim., or over 4 centim. from its inner margin to the 
posterior humeral articulation. The visceral surface is remarkably 
flat; but beneath the region occupied by the spine the base was 
somewhat concave. The humeral articular surface obliquely trun- 
cates the inner half of the proximal end. Itis much roughened and 
grooved with the marks of a cartilaginous epiphysis, and was broader 
in proportion to its length than in the larger species. It shows 
some sign of crushing, and is fully 4 centim. wide and 47 millim. 
long. Its posterior outline is much broader than in the larger spe- 
cies ; and the axis of the articular surface was not materially different 
from the plane of the blade. The thickness of the anterior coracoid 


FAUNA OF THE GOSAU FORMATION, 663 


process was but little more than 15 centim. where widest, and became 
somewhat narrower as it extended outward. 


Humerus. 


Both the mght and left humeri are strong bones (Pl. X XIX. 
figs. 1-3) which, previously to fossilization, had lost both proximal 
and distal articular ends. They are of exactly the same length as pre- 
served, and are mutilated in almost the same manner, the distal ends 
especially being obliquely truncated from behind forward; and they 
show a subquadrate section. The right humerus is slightly the 
more perfect ; and neither bone is distorted by pressure. 

The fragments are 8 inches long; and, as preserved, the right 
humerus is 47 inches wide at the proximal end ; the left humerusis 
4 inches wide; the shaft is most constricted in the middle, where it 
measures 1, inch from side to side; there is no corresponding con- 
striction from back to front; but in this position the antero-pos- 
terior measurement is 14 inch. The bone widens distally ; but, as 
preserved, the distal measurement from side to side is only 2,3, 
inches. The proximal articular head was nearly in the same plane 
as the distal end. The inner lateral outline is gently concave; 
the external outline is deeply concave, owing to the expansion out- 
wards of the large thick deltoid process, which is bent at a consider- 
able angle with the shaft. The antero-inferior and postero-superior 
outlines are both nearly straight, though very slightly concave ; and 
they converge slightly from behind forward, owing to the slight distal 
twist giving to the bone an appearance of thickening to that end. 
At the distal end the shaft becomes flattened both in front and 
behind ; and these surfaces are nearly parallel; and, where fractured, 
the bone is here 1,6 inch thick on the outer side, and somewhat 
thinner on the inner side. The remainder of the inferior surface 
is anterior to, and makes a slight angle with, the subtriangular 
flattened distal area. It also may be said to be a long triangle ex- 
tending from the inner corner of the distal articulation forward to 
the divergent elements of the proximal end (fig. 2). Nearly its whole 
length is straight ; and its middle portion is more rugose with mus- 
cular attachments than the rest of the bone: the markings have the 
appearance of slightly impressed ovate pits, which extend for a length 
of nearly 3 inches. This inferior area is defined by faint lateral 
ridges, and proximally, beyond the muscular markings, becomes 
somewhat deeply concave from side to side, and compressed towards 
the superior aspect, so that in length it is convex. This area is a 
little inflated on the inner portion, and terminates laterally in a 
slight sharp ridge. The outer expanded wing has a well-rounded 
margin. 

Superiorly the bone is highly convex from side to side (fig. 1), 
though somewhat flattened on the inner and posterior side, and 
also on the expanded external process, which is smooth and 
slightly concave in length, and slightly convex from within outwards. 
The upper aspect of the bone is divided into two portions by an 


664 PROF. H. G. SEELEY ON THE REPTILE 


oblique moderately elevated muscular ridge, which extends for about 
32 inches across the middle of the shaft from near the inner side 
proximally towards the outer side distally ; and though the whole 
superior surface, except the expanded crest, is roughened with mus- 
lar lines, another fainter ridge may be traced simane straight dis- 
tally from the proximal termination of the oblique ridge. Imme- 
diately below the proximal articulation in the middle of the shait is 
an elevated muscular boss about 14, inch in diameter, which, as 
preserved, is subcircular. The thickness of the shaft at its inner 
margin is 1,5, inch, and at its outer margin 1-3, inch. The radial 
crest hickeane towards the proximal surface, “and curves a little 
upward. The outline of the fractured proximal end is somewhat 
boat-shaped and compressed (fig. 3). This humerus is quite distinct 
in character from any form of which I have any knowledge. 


Femur. 


The right femur (Pl. XXXI. fig. 5), found in 1876, is 10,2, inches 
long. The left femur (Pl. TOOL fig. +), found in 1877, 18 hardly 
more than 10 inches long. This difference i is apparently dns to the 
different ways in which the bones are compressed. They are both 
in the same state of mineralization, of a rich chocolate-brown colour, 
and quite free from matrix, which has been removed by Professor 
Suess. The bones belong to a somewhat distinct type; they offer many 
resemblances, as Professor Suess pointed ‘out to me, to Cr yptosaurus 
of the Oxford Clay, but are more slender. They therefore show the 
typical characters of Dinosaurs, though there is a difference from 
all English genera in the proximal anterior trochanter not being 
separated from the shaft ; and there is a remarkable development of 
muscular ridges on the bone, one of which extends on the proximal 
posterior face (fig. 4)in a curve upward and outward from the small 
middle trochanter on the inner margin of the shaft to the outer 
and external margin of the proximal articulation. It is impossible 
not to recognize the similarity of this strong muscular ridge to the 
ridge seen on the corresponding aspect of the mammalian femur ; 
and if this coincidence be admitted, it goes far to prove that the 
middle trochanter, which is the most distinctive mark of the femur 
of a Dinosaur, is homologous with the inner or lesser trochanter of 
man; and so far it would seem rather to imply a foreshadowing 
of a mammalian plan of muscle-arrangement. A similar muscular 
attachment to this may be observed in Crocodiles above the middle 
of the shaft. From it an intertrochanteric muscular ridge extends 
to the position in which the proximal trochanter of Dinosaurs 
is seen when it is developed. The shaft is most constricted in its 
distal third, where there is a slight flexure bending the distal arti- 
culation backward ; both articular ends appear to have been highly 
cartilaginous, since they are marked with ramifying furrows and 
occasional pits. As is usual with cartilaginous surfaces, the arti- 
cular margin is sharply defined. 

The proximal articular surface is best preserved in the right femur, 


— - 


FAUNA OF THE GOSAU FORMATION. 665 


and, as usual, consists of a subcircular head (fig. 5), which is directed 
inward and forward, and a narrower external area. The posterior 
border of the articulation is nearly straight ; but the anterior out- 
line is deeply excavated between the head of the bone and the 
external trochanter. The globose head measures 1; inch from front 
to back; and may be considered to be 15% inch from within outward; 
but the entire length of the proximal articular surface was about 
3,2; iach. The articular surface beyond the head contracts to less 
than | inch from front to back, but widens again to fully 1,3, inch 
at the border of the external trochanter; as in Cryptosaurus ewme- 
rus, the narrower external part of the articulation is concave from 
within outward, and does not extend so far proximally as the 
convex head by half an inch. The length of the bone to the distal 
border of the head on its inner margin is not more than 9 inches. 

The anterior proximal trochanter (fig. 5) is about 13 inch long, and 
convex in length: so that it dies away both distally and proximally, 
where it merges in the articular surface. It is rough with oblique 
muscular markings, and rounds into the flattened bnt slightly con- 
vex external surface, which is also roughened, with a triangular area 
of muscle-marking 25 inches long, which tapers distally. Below 
this area the external side loses its flattened aspect, and becomes 
rounded from front to back. 

The proximal half of the shaft is considerably compressed from 
above downward, and is flattened on both aspects: its width from 
within outward in a line with the distal limit of the proximal tro- 
chanter is about 24, inches, and just above the lateral inner trochanter 
about 1,4 inch, while just below the lateral trochanter the width 
is about 1,3, inch ; and there, as the shaft becomes narrower, it 
grows more convex from side to side. The superior or anterior 
aspect of the bone is marked with a strong longitudinal muscular 
ridge, which originates towards the hinder part of the articular ball, 
curves a little outward and then inward, and extends as nearly as 
possible in the middle line of the shaft for a length of 65 inches; 
it is moderately elevated, strongest proximally. and dies away where 
a distal flattening of the bone gives an aspect of flexure to the lower 
part of the shaft. There is also a second muscular ridge, which 
originates at the same point, below the outer limit of the proximal 
head, and, running obliquely inward, curves round the convex inner 
side of the bone and becomes merged in the proximal limit of the 
trochanter. Both these ridges are iess marked than in the larger 
species. 

The posterior aspect of the shaft(fig.4)is much more flattened. The 
strong muscular ridge to which I have already referred as defining 
an area homologous with the obturator-region of mammals, extends 
distally for nearly 3 inches, and then curves more sharply inward 
to merge in the inner trochanter. rom this ridge extend obliquely 
inward, so as to cross each other, two series of narrow straight 
linear muscular markings. Parallel to the sigmoid curve of the 
proximal articular margin, and about a quarter of an inch below it, 
is a line of about six or eight circular vascular perforations. 


666 PROF. H. G. SEELEY ON THE REPTILE 


The lateral trochanter is placed, as usual, at the angle between 
the internal and posterior aspects’ of the bone (fig. 4). It is moderately 
elevated, about 14 inch long, 6) inch wide proximally, and tapers 
distally ; it is placed exactly midway between the proximal and 
distal articular ends. Below the trochanter the section of the shaft 
becomes subtriangular, being flattened on the internal aspect and 
posteriorly, and rounded on the external and anterior aspect. 

The distal articulation is chiefly noticeable for the inflated expan- 
sion of the bone at the external margin, and for the relatively large 
size of the condyles (fig. 4). The articular surface is 3 inches long in 
the left femur, which has this region best preserved ; it is very mode- 
rately convex from behind forward, and very slightly concave from 
within outward, and rounds gently into the anterior surface of the 
bone, where a concave natural impression divides the anterior 
margin into a larger internal area and a smaller area which is 
external. ‘There is the usual ill-defined gently concave pit for liga- 
mentous attachment just above the articulation on the flattened in- 
ternal surface of the bone, which looks obliquely upward, much as 
in Cryptosaurus. The internal condyle is the larger of the two ; it 
is about an inch wide, and curves round considerably on the paste 
aspect of the bone, so as to cause the articulation to measure 2,2 To 
inches is om front to back. The interspace between the condyles is 
about ;% inch; and in this region the articulation measures, from 
front to back, 1> inch. This depression becomes prolonged up the 
middle of the posterior side of the shaft towards the inner part for 
about 14 inch. The smaller condyle is more compressed, about 
= inch wide, and gives an antero-posterior measurement to the arti- 
cular end of 2 inches; and external to this condyle is a concave area or 
eroove, + an inch wide, which defines it from the well-rounded broad 
external margin. The small part of the articulation external to this 
condyle makes a considerable angle with the major part of the sur- 
face. The anterior® half of the articular surface is nearly smooth ; 
but the posterior half is deeply scored with about eight comparatively 
straight grooves, six of which lie between the condyles. These 
grooves appear, from their corresponding development at the anterior 
part of the proximal articulation, to be in the positions of greatest 
pressure and greatest condylar growth, and may be regarded as evi- 
dence that the bone was carried in an oblique position, as among 
mammals. 

Tibia. 

A smaller pair of tibial bones are much less perfectly preserved 
than those of the large species, only exhibiting about 6} inches of 
the middle of the shaft (Pl. XXVII. fig. 19). Their ends are 
decayed in the usual way; and distally the fractured outline was 
subtriangular, but formed a triangle in which the anterior and two 
converging posterior elements rounded into each other, and were 
subequal. Here the extreme antero-posterior measurement 1s 
about 1,2; inch, and the extreme width from side to side at the 
distal end is the same. In the middle of the shaft the antero-posterior 


FAUNA OF THE GOSAU FORMATION. 667 


measurement remains unaltered, but the side-to-side measurement 
is reduced to 42 inch; where the specimen is fractured proximally, 
the antero-posterior measurement is $/ inch, while the measurement 
from side to side in the middle of the shaft is 1,1, inch. The right 
tibia (fig. 19) does not appear to be crushed ; but the left specimen 
is somewhat fractured at its distal end. ‘The inner side in both is 
flattened, though not quite so flat as in the larger species. The outer 
side 1s convex, but divided into two portions by a median ridge, 
which in its upper 34 inches is strongly muscular, though the 
markings appear to be stronger on the left tibia than on the right— 
a condition the reverse of that which obtains in the larger species. 
The muscular ridge is made up of three or four close parallel ridges. 
The posterior half of this side of the bone appears to be more flattened 
than in the larger species, while the anterior half shows indications 
of a similar longitudinal concavity, though the specimens are frac- 
tured too low down for more than the beginning of it to be detected. 
The proximal fracture displays a triangular outline with a long 
straight base formed by the inner side and two shorter converging 
sides which form the outer side. Here the bone is less than twice 
as deep as it is wide. On the posterior side, as compared with the 
larger species, the side-to-side compression is greater towards both 
the proximal and distal ends, while anteriorly the bone is rather 
more rounded from side toside. ‘There is about as much difference in 
size between the two types of tibie as there is between the two kinds 
of femora, though, so far as can be judged from the fragments 
preserved, the differences in essential characters in the tibize were 
less important than those of the femora. 

There have also been found fragments of patelloid ridges of 
tibiee which appear to belong to a species slightly larger than either 
of these; but the materials are too imperfect for description, or even 
for absolutely certain osteological identification. 


Fibula. 


The specimen which Bunzel (pl. i. f. 12, 138) regarded as the 
upper half of the left humerus of a Crocodile is undoubtedly a 
somewhat obscure fossil. It, however, presents nothing in common 
with any crocodilian humerus with which I am acquainted, espe- 
cially differing in its remarkable compression, in wanting all trace 
of a radial crest, in the lateral compression of the shaft at right 
angles to the supposed head of the bone, and in the inflation of 
the inferior side of the head. While, therefore, [ have no hesitation 
in affirming that the specimen is not crocodilian and not a humerus, 
the loss of the terminal articular end and the evidence of a certain 
amount of crushing makes any other determination a matter re- 
quiring some caution. The contour, however, of the bone is so 
similar to that of the tibia of Cratcwomus, and one aspect, and especi- 
ally the posterior margin, is so roughened with muscular attach- 
ments, that I have little hesitation in affirming that we have here 
the fibula of one of the Dinosaurs—a view which is further sup- 


668 PROF. H. G. SEELEY ON THE REPTILE 


ported by the circumstance that the tibia has a strong ligamentous 
ridge that would correspond to the rough side of the fibula. 
Moreover I have found a further small portion of the supposed shaft, 
not absolutely continuous, but showing that the bone retained the 
same characters for some length further, and did not expand at its 
distal end. It therefore may be well to state that this specimen 
(Pl. XXVII. fig. 20) is to be regarded as a right fibula imperfect at 
the proximal end, but, as preserved, 44 centim. wide. As pre- 
served, the main piece of bone is over 10 centim. long, and, with the 
additional fragment and the lost interspace, would indicate a length 
of about 14 centim. The distal end is imperfect, having decomposed 
before fossilization. The anterior margin is concave, the posterior 
margin straight and rugose. The thickness of the proximal end, as 
preserved, is about a centimetre, while the fractured distal end is 
7 millim. thick and about 17 millim. wide. As remarked by Bunzel, 
the distal fracture is semiovate, but the flattened side is towards the 
transversely convex head of the bone, while the convex distal outline 
is towards the transversely concave or external surface of the head. 
Other fragments of similar character, also presumably fibular, but too 
imperfect for detailed description, may be referred, one to a larger 
and one to a smaller Dinosaur. 


Metatarsal Bone. 
(See Biinzel, pl. iv. figs. 11, 12.) 


The specimen regarded by Biinzel (t. iv. f. 11, 12) as the phalange 
of a Crocodile is almost too imperfect for accurate determination ; 
but since it is certainly either the second or third metatarsal of a 
Dinosaur, probably the former, it requires a shght notice. As pre- 
served, it is little more than 64 centim. long and about 3 centim. 
wide in front. What I take to be the inferior surface is the best 
preserved ; only a small portion remains of the anterior articular 
end, which was unusually convex from above downward. The lateral 
outlines are concave; and the bone measures only 2 centim. from 
side to side in the middle, where most constricted. ‘The superior 
surface was compressed, so as to form an obscure broad rounded 
ridge on the outer side, with a very shght broad channel below it on 
the inner side. The depth of the bone in the middle of the shaft is 
21 centim. The under surface is concave in length, flattened from 
side to side, with a slight twist in the plane, which is directed a little 
inward as it extends forward. The posterior fractured end is sub- 
triangular, owing to the flattening of the base. 


Claw-Phalange. 
(See Bunzel, pl. iv. figs. 4, 5.) 


A claw-phalange (Pl. XXIX. fig. 6), probably pertaining to the 
second or third digit of the left hind limb, as indicated by Bunzel, 
is ascribed by that writer to a species of Scelidosaurus, but may 
be referred to Cratwomus lepidophorus. It is 33 millim. long, 25 
millim. wide behind, and 22 millim. wide in front, is compressed 


FAUNA OF THE GOSAU FORMATION. 669 


from above downward, and, though blunt anteriorly, is most com- 
pressed on the right border. The articular surface is imperfectly 
preserved, but was concave from above downward, straight trans- 
versely, and inclined obliquely to both superior and inferior aspects. 
The superior surface, which is slightly convex from side to side, 
is margined on each side by a strong groove, which is not very 
deep, and extends forward for more than half the length of the 
bone. Between these grooves are a number of slight parallel 
ridges. At the sides the bone is excavated for the attachment of 
powerful ligaments, more so on the left side than on the right; and 
on the left side the irregular excavations extend further forward. 
The under surface is comparatively flat, but concave in length, convex 
from side to side, and marked in the middle with two irregular longi- 
tudinal grooves,which have a tendency to branch as they pass forward. 
These grooves are very imperfectly indicated in Biinzel’s figure, which 
gives no indication of their dendroid character. The posterior outline 
of the bone is ovate, and less than 2 centim. deep. 


Dorsal Vertebra. 
(See Bunzel, pl. 1. figs. 24, 25.) 


A dorsal vertebra with the neural arch fairly well preserved is the 
best vertebral evidence of this species(Pl. XXX. fig.5). The centrum is 
4 centim. long, flattened on the under side, where it is 14 centim. broad 
in the middle, where most constricted ; it is concave from front to 
back. The sides are compressed and somewhat concavely excavated 
below the neural arch, where the least transverse width is 1+ centim. 
Thus the body of the vertebra in section would be subquadrate. 
The anterior articular surface of the centrum is concave, 32 millim. 
broad, and 27 millim. deep. ‘The posterior end is not concave, but 
somewhat flattened and convex from above downward. The margin 
is a little worn; the greatest transverse width is less than 3 centim., 
the greatest depth 25 centim. The neural arch is high, though not 
unusually so for a Dinosaur. Its base is nearly as wide as the 
anterior face of the centrum. It extends the whole length of the 
centrum, and has the aspect of being compressed from side to side 
below the transverse processes. The borders of the neural arch are 
excavated (fig. 5), moderately in front, and more deeply behind, to 
form the intervertebral passage for the nerves. The least length of 
the neural arch from front to back, in the middle of the neural canal, 
is 25 millim. The length from the anterior to the posterior zyga- 
pophyses is about 54 centim.; and the upper border of those facets 
is fully that height from the base of the centrum. The anterior 
zygapophyses are directed upward, forward, and inward. ‘Their 
external surface is rounded. There is a V-shaped notch between 
them in front; and they form the anterior border of a transverse 
cup-shaped depression in front of the neural spine, the hinder 
borders of which cup are contributed to by the transverse pro- 
cesses, which are directed upward and outward, and placed between 
the zygapophyses in the middle region of the vertebra. The neural 


670 PROF. H. G. SEELEY ON THE REPTILE 


spine is narrow and short, and placed on the hinder half of the 
neural arch; only its base is preserved. The posterior zygapophyses 
are circular facets placed below and somewhat beyond its hinder 
termination. They converge downward, but are separated by a broad 
groove. The height of the vertebra to the base of the neural spine, as 
preserved, is 6£centim. The extreme width over the posterior zyga- 
pophyses is 24 centim. The width of the groove between them is 
about 4 centim. ‘The side of the neural arch is of the usual cha- 
racter, with ridges ascending from the anterior and posterior ends of 
the arch which converge upward; and the posterior ridge, which is the 
better marked, passes into the strong ridge which extends under the 
base of the transverse process (broken away on the left side) The 
area between these two lateral ridges, which is unusually deep and 
narrow and ill-defined, appears to be the capitular articulation for 
the rib. The space posterior to the lateral ridges is concavely ex- 
cavated. The transverse process is broken off short. I infer this 
to have been an early dorsal vertebra. The posterior convexity of 
the centrum shows it to have been full-grown. This, no less than 
the long narrow articulation at the side, and the other characters 
of the neural arch described, show it to indicate a distinct species 
from the vertebre referred to Cratcomus Pawlowitschit. 


MEGALOSAURUS PANNONIENSIS, Seeley. 


There are two teeth of a carnivorous Dinosaur (Pl. XX VIL. figs. 
21-23) which present some resemblance to the teeth of Megalosaurus 
and Lelaps, differing in no character of importance except size, the 
fineness of the serrations, and shortness and breadth of the crown. 
One specimen is a crown tolerably perfect, fractured just above the 
base and before the commencement of the fang (fig. 21). The other 
is the lower half of the crown of a somewhat larger but similar tooth. 
The more perfect specimen is 21 millim. long, curved backwards, 
quite straight, convex on both sides, though rather more so on the 
inner side, and with the inflation towards the convex anterior margin 
of the tooth. The posterior margin is relatively straight, but 1s con- 
cave. he surface of the tooth is marked with microscopic longitu- 
dinal wrinkling and faint parallel transverse lines of growth, only 
to be detected by the way in which they reflect light. The posterior 
margin throughout its length is marked with perfectly regular 
transverse serrations, which extend along the tooth like a fringe. 
Towards the extremity of the crown the serrations become slightly 
shorter. On the anterior border (fig.22)the serrations are of a similar 
character, but only reach down the tooth for 13 millimetres, becom- 
ing smaller as they disappear. There are about forty-five of these 
minute chisel-like serrations in all this margin. Where they ter- 
minate, the tooth is just appreciably narrower and the anterior 
margin is rounded, so that the transverse section (fig. 23) is exactly 
the same asin Megalosawrus. The serrations of the posterior margin 
are larger than those of the anterior margin ; so that there are only 
about forty in the entire length of the tooth. The antero-posterior 


FAUNA OF THE GOSAU FORMATION. 671 


measurement is over a centimetre, and the thickness 6 millimetres. 
There are no bones that I could refer to this species; and when 
they are discovered the teeth may prove to belong to an animal as 
different from Megalosaurus as is Leelups. 


ORNITHOMERUS GRACILIS, Seeley. 


The specimen figured by Bunzel, pl. vii. figs. 22, 23, and regarded 
(p. 15) as the middle of the dorsal rib of a lizard, is the distal half of 
the shaft of the femur of a remarkable new Dinosaurian. From the 
circumstance that Biinzel has figured the external instead of the 
internal aspect of the bone, it would have been difficult to make this 
interpretation without examination of the specimen. The fragment 
(Pl. XXVIII. figs. 6,7) is only 53 centim. long, has a nearly circular 
shaft 13millim.in diameter at the proximal fracture, becoming a little 
more compressed distally from above downwards and slightly more 
expanded from side to side. The specimen shows no trace of the 
distal articular end; but distally the bone is a little flattened on the 
inferior and posterior surface, and slightly compressed towards the 
outer border. There is a distal curve in the bone, rather more 
marked, perhaps, than in the crocodilian femur. Towards the proxi- 
mal end of the fragment the transverse fracture passes through a 
longitudinal muscular pit, margined below by an elevated muscular 
ridge, which is prolonged further distally than the muscular pit, 
and appears to have terminated in a free process, though the ex- 
tremity of this is broken away. ‘This is the internal trochanter of 
the Dinosaurian femur (figs. 6, 7). What remains of the muscular 
impression is about 12 millim. long and half a centimetre wide. 
What remains of the sharp ridge bordering it is 17 millim. long. 
I am not acquainted with any Dinosaur in which the femur has this 
cylindrical bird-like form. ‘The shaft is formed of dense bone with 
a large medullary cavity about 7 millimetres in diameter (fig. 7), 
and has, at first sight, rather the aspect of the bone of a bird than 
of a Dinosaur. Though the fragment is so imperfect, it is so cha- 
racteristic that I have ventured to refer it to a new genus. 


DoRATODON CARCHARIDENS (Bunzel). 


The sculpturing of the outer surface of the jaw, no less than its 
general form, would seem to have weighed with Biinzel in referring 
the specimen represented in his plate 1. figs. 29-32 to the genus 
Crocodilus. I find myself unable, however, to accept this generic 
determination, partly because the teeth are such as indicate a dit- 
ferent genus, and partly because I am led to refer the maxillary 
bone represented in the same plate, figs. 5-5, to the same genus 
and probably the same species as the lower jaw; and this shows, 
though the fragment is very imperfect, characters which are not 
met with in the genus Crocodilus. But whether its affinities 
are stronger with Crocodiles or with Dinosaurs is a matter 
far from easy to determine. The lower jaw consists of slender 
rami, having a length, as preserved, of about 137 centim. with- 


672 PROF. H. G SEELEY ON THE REPTILE 


out reaching back to the articular region. The jaws converge 
forward to the symphysis, where the contraction ceases, and there 
is a slight anterior expansion before the lanceolate anterior termi- 
nation. The greatest width of this slight expansion is 24 centim.; 
and the width of the diverging rami at 10 or 11 centim. from 
the anterior termination is over 5 centim.; thus the jaw is re- 
markably pointed. The lateral contour of the alveolar margin 
is convex from before backwards in the region of the symphysis and 
concave in length behind the symphysis. The symphysis is 3 cen- 
timetres long, and is made up in the anterior and inferior part by the 
dentary bone, and in the posterior and superior part by the opercular 
bone, which, on the alveolar aspect, forms half the symphysis, while 
inferiorly it only constitutes the hinder fifth. In Crocodiles the oper- 
cular bone does not enter.into the symphysis. The anterior part of 
the jaw in the symphysial region is excavated in a spoon-shape, 
owing to the remarkable and vertical elevation of the alveolar margin, 
an elevation which appears to have relation to the straightness and 
vertical position of the teeth ; so that the fangs could not have made an 
angle with the crown. The whole inner side of the ramus is formed, 
as in crocodiles, by the opercular bone, which, as Biinzel remarks, 
is smooth and slightly rounded at its inferior and superior margins, 
and extends back beyond the alveoli, only showing in its anterior 
part two or three nutritive foramina. The external surface of the 
bone is formed of the dentary element, except, it may be, towards 
the hinder superior border, where a suture appears to indicate on 
the inner side a coronoid bone. On the under surface the jaw is 
flattened at the symphysis, but the flattened area rounds up an- 
teriorly to the alveolar margin; but, where the rami begin to diverge, 
there is a distinct sharp angle between the base of the jaw and its 
side, and this ridge is prolonged backwards for a large part of the 
region through which the teeth extend: and here the base of the 
ramus is slightly convex from side to side till, with the fading of 
the angle into the upper surface, it becomes more rounded and nar- 
rower. ‘The suture for the opercular bone runs along the inner 
third of the base. In ornament, the anterior expanded end of the 
snout is pitted much after the pattern of crocodiles, and evidently 
with similar relation to a vascular condition; but inferiorly the 
ramus is marked with rough, short, irregular longitudinal ridges, 
which extend round onto the side but do not risé to the alveolar 
region, Which is comparatively smooth and marked with a row 
of relatively large foramina, about seven’ or eight in number, 
and placed above the middle of the lateral margin in a concave 
line or groove. As they extend backwards they rise nearer to the 
alveolar margin, and form the basis of a slight compression of the 
bone above them. Posterior to the symphysis the depth of the 
jaw increases: it is about 9 millim. in front, 17 millim. at the 
last tooth- socket, and 3 centim. in the coronoid region; so that 
it increases in depth more rapidly behind the teeth. It is 
difficult to count the exact number of sockets; for some of 
the fangs are preserved, and in other cases the teeth have fallen 


FAUNA OF THE GOSAU FORMATION. 673 


out: but there were not fewer than fifteen; and as the alveolar 
groove is carried back and becomes very shallow beyond this 
point, it is possible there may have been five teeth more, of small 
size, in the hinder part of the jaw, making a total of twenty. The 
teeth are largest in the middle of the jaw; but only the tenth on 
the right side has the crown preserved in situ. The crown is 8 mil- 
lim. high, and nearly 6 millim. wide at its base. It is trian- 
gular in lateral outline, is curved inwards and directed upwards, 
inwards, and backwards. Its base is rather less than 3 millim. 
thick. Hach surface is convex, terminating in a sharp cutting-edge, 
which is very finely serrated along the margin. In front and be- 
hind there is a constriction, so as to separate the sides of the crown 
from the elliptical fang; but this constriction is not appreciable on 
the interior or the exterior aspect of the tooth. This form of tooth 
is entirely Dinosaurian. Four teeth have been found separately 
which show the same character. ‘Two of these are crowns broken 
off directly from the fang, and show the constricted oval base of the 
crown where the lateral ridges become pinched in. These teeth 
are sharply pointed, and have the surface smooth to the naked eye. 
There are also two teeth which have the same general form, except 
that the crown is broader and shorter; and, owing to this circum- 
stance, the serrations, which are transverse to the cutting-edge, 
have an appearance of being directed obliquely upward. These, 
however, are probably successional teeth, it may be from another 
part of the jaw, or from the upper jaw. A certain amount of varia- 
tion is obvious, because the fang of the eleventh tooth on the left 
side shows that the base of the crown was marked with blunt 
parallel ridges. 

There is a fragment of the anterior end of a right dentary bone 
from which the opercular element has come away, and which clearly 
belongs to the same genus. It may indicate another and smaller 
species, since the rami appear to diverge more rapidly, to have con- 
tained more numerous teeth, with smaller and more circular fangs, 
to want the anterior elevation of the jaw in its presymphysial region, 
to be devoid of the ridge between the base of the jaw and the side, 
and to have the side convexly inflated instead of flattened, especially 
external to the alveolar margin. The ornament also appears to be 
slightly different ; but as no teeth are preserved I have not thought 


_ it necessary to give a name to this fragment. The length of the 


dentary symphysis is 12 centimetre, and the length of the fragment 
43 centimetres. The corresponding length of the dentary symphysis 
in the larger specimen exceeds 23 centimetres. 

The fragment of upper jaw briefly described by Biinzel, p. 6, pl.1. 
figs. 3-5, I have, as already mentioned, identified with the lower 
jaw of the large species just described. Notwithstanding the circum- 
stance that Biinzel remarks on its close resemblance to existing 
erocodiles, he places the nasal aperture immediately in front of the 
orbit, which alone would suffice to show that the type differed from 
crocodiles fundamentally. In fact, the perforation of orbit and 
nares in the maxillary bone would be a modification of old-fashioned 


Q.J.G.8. No. 148. : 2¥ 


674. PROF. H. G. SEELEY ON THE REPTILE 


anatomy of no ordinary kind, since the maxillary bone does not 
enter into either the orbit or the anterior nares of the crocodile, and 
it certainly does not enter in any known Dinosaur into the external 
wall of either of these vacuities. Agreeing with Biinzel, that the 
margin which he regards as the anterior border of the orbit is cor- 
rectly identified, I regard the perforation which he terms nasal as 
the preorbital vacuity characteristic of Teleosaurs, and more or less 
developed in various Dinosaurs. The bone between the orbit and 
the preorbital vacuity is always the lachrymal; and I therefore 
identify the lachrymal bone as united by suture with the maxil- 
lary. The length of its base is 2 centim.; but it is fractured 
superiorly, and therefore its outline cannot be stated, further than 
that it appears to have been triangular. The posterior margin 
is concave, rounded and thickened, with an indication of a groove, 
which may have had relation to the lachrymal canal. The surface 
is sculptured with somewhat oblique ridges, which are short and 
irregular, and deeper than the sculpturing on the maxillary. The 
suture with the maxillary is straight but slightly oblique, so that it 
laps a little further down on the inner than on the external surface. 
The preorbital vacuity only shows a small portion of its basal 
margin, which is rounded. The lachrymal bone in front of it is 
thin, and gives the aspect of the vacuity having penetrated ob- 
liquely inwards and forwards. A small portion is preserved of a 
suture on the superior surface, which is straight and parallel to the 
alveolar surface, or but slightly inclined forwards. Hence it may 
reasonably be identified as the suture for the nasal bone. The 
depth of the bone from the nasal suture to the alveolar border 
is 22 millim. Anteriorly the bone is fractured, so that there is no 
indication either of its length or of the length of the nasal suture, 
or of the nature of its relation to the premaxillary bone. ‘The pos- 
terior end is also fractured ; but just below the orbital border there 
is a minute indication of a suture, evidently indicating the malar 
bone, and showing that its relations were the same as in Dinosaurs. 
The surface of the maxillary bone is marked with an indefinite 
rough sculpturing, which, in the upper part, has a tendency to 
assume a linear character; and the hinder part is somewhat lightly 
pitted. The internal surface is necessarily irregular ; and its appear- 
ances may be passed over in so far as they relate to the region 
above the palate ; but above the alyeoli the bone evidently developed 
a horizontal palatal plate, which has been almost entirely broken 
away. It appears to have been notched out posteriorly into a post- 
maxillary vacuity, such as is seen in the crocodile, since the hinder- 
most 14 centimetre is a smooth, sharp, somewhat concave margin 
bordering the alveoli. 

It is very difficult to understand the alveolar structure from an 
inspection of Biinzel’s plate, since it gives the appearance of a double 
row of tooth-sockets: this is due to the circumstance that while the 
tooth-sockets (in which most of the teeth still remain) are placed 
close to the outermost alveolar border, there is, internal to them, a 
parallel series of pits which are broad and shallow, and are, I think, 


FAUNA OF THE GOSAU FORMATION. 675 


produced by the circumstance that the teeth of the lower jaw were 
received scissor-like between the teeth of the upper Jaw; and these 
pits I regard as excavations which have resulted from the pressure 
of their crowns—a view which is especially supported by the cir- 
cumstance that they are deeper posteriorly, where the palatal border 
is but little above the outer alveolar border, and are less marked 
anteriorly, where the palatal border rises about 8 millim. above 
the alveolar border. The teeth extend along the whole alveolar 
length, which is 44 centim., and were eight in number in the 
fragment. ‘They are larger in front than behind, and mostly ap- 
pear to have been successional teeth not fully cut. The first, 
where broken, is 7 millim. long and 4 millim. thick. The second 
is less than 4 millim. long, and is clearly a section of the fang; 
but one of the later teeth shows the sharp serrated cutting-edge, 
compressed form, and smooth enamel characteristic of the teeth of 
the lower jaw; and it is on this evidence that I have felt justified 
in referring it to the same species. Both fang and crown appear 
to have been hollow ; but as the cavities are filled with iron pyrites, 
I have not been able to excavate them. 

There are a few other unimportant fragments, chiefly of the lower 
jaw, which exhibit similar sculpture, and presumably belong to this 
genus, but too imperfect to be worthy of description. 


There is a small claw-phalange (Pl. XXVII. fig. 26) which, 
perhaps, for the present may be noticed here, seeing that it is quite 
impossible to say with certainty to which of the animals it belonged. 
It is 17 millim. long, curved downward and to the right. It tapers 
to a point, and is subtriangular, being flattened on the under side, 
on the left side, and obliquely on the right side, which is large. 
But these three surfaces round into each other, except where they 
are divided by the sharp lateral ridges which margin the base. The 
articulation is 7 millim. deep, a little narrower, concave from above 
downwards, and convex from side to side. 


RHADINOSAURUS ALCIMUS, Seeley. 


I found this genus upon the femora, which are quite distinct from 
any thing hitherto discovered. The humeri are such as would be 
associated with those bones, though there is no proof beyond simi- 
larity of character that they belong to the same species. The same 
remark applies to the vertebra, which are such as might be expected 
in an animal of this kind; but there is no evidence of natural asso- 
ciation. I have placed this genus next in succession to Doratodon, 
because that genus is founded upon a head, while this is formed for 
limb-bones ; and though there is no evidence to justify their bemg 
thrown together, there is a possibility that Doratodon belongs to 
one of the animals of which the head cannot be identified. 


Humerus. 


Two specimens of humeral bones (Pl. XXXI. figs. 8-10) exhibit 
characters indicative of an animal in many respects unlike any other 
2y 2 


676 PROF. H. G. SEELEY ON THE REPTILE 


in the deposit, especially in the slenderness of the shaft and relatively 
small size of the articular ends, as well as in the circumstance that 
the articular extremities were at right angles to each other. Unfor- 
tunately both proximal and distal ends are lost by decomposition. 
The left humerus is best preserved proximally, while the right . 
humerus extends further distally. The length indicated by the two 
bones without reaching the articular ends is 14 centim.; so that 
the entire length of the bone, when perfect, was at least 2 centim. 
more. The left fragment (fig. 8) is fully 12 centim. long, and, when 
placed with the distal end uppermost, shows a slight convexity on 
one lateral margin and a corresponding concavity on the other, 
the concavity facing the anterior and inferior aspect of the bone, 
the convexity being superior. Owing to the circumstance that the 
proximal end is absolutely at right angles to the distal end, it hap- 
pens that the shaft of the kone widens distally. The transverse 
measurement at the base of the radial crest (fig. 9) is 14 centim., and 
atthe distal fracture 23 millim. (fig. 10); and it is still widening. 
The superior distal surface is convex from side to side, with a slight 
ridge towards the radial side of the bone, which is really a prelon- 
gation of the muscular ridge of the radial crest, which in crocodiles 
never extends along the superior aspect of the bone. This ridge 
produces a flattened radial aspect, a slight approximation to which 
may be observed at the distal articular end of the crocodilian hu- 
merus ; but here it gives a somewhat compressed and sharp aspect 
to the inferior radial margin of the bone along the distal half of the 
shaft, while the ulnar side is relatively flattened or rounded. The 
bone is marked, on the superior aspect especially, with strong longi- 
tudinal strie or slightly elevated ridges. _ Its thickness at the distal 
end, as preserved, is about 13 millim. ; but the right bone is a 
trifle stouter. The inferior aspect is flattened distally with a slight 
longitudinal depression. The proximal end of the bone necessarily 
widens, while the shaft remains comparatively uniform. The su- 
perior surface is smooth, and convex from side to side, but slightly 
channelled towards the expanded radial process, which is necessa- 
rily placed on the middle of the aspect which, when viewing the 
distal end, would be regarded as superior. The width of the 
proximal end appears to have been small. The lower part of the 
_ radial crest gives a transverse width, as preserved, of less than 
21 centim. The inferior aspect of the proximal end ‘is longitu- 
dinally channelled, and thus divided into a compressed anterior 
process and a rounded and somewhat inflated inferior and posterior 
part. The thickness of the bone here at the fracture is just over a 
centimetre. The whole inferior surface, not only of the radial crest 
but of the adjacent region of the bone, is roughened with powerful 
muscular attachments. The right fragment substantially repeats 
these characters; only the bone is appreciably stronger, with its 
muscular ridges more marked, and in length shows a decided sig- 
moid curve, like that which marks the humerus of a crocodile. I 
am led to refer these bones to the same species as is indicated by 
the femora next described. In this case they would indicate an 


Low 


FAUNA OF THE GOSAU FORMATION. 677 


animal with remarkably small anterior limbs, suggesting the pro- 
portions of a Teleosaur. I may state that this identification is 
founded chiefly on similarity of general aspect, superficial texture, 
condition of preservation, and colour. 


Femur. 
(See Biinzel, pl. ui. fig. 1.) 


A pair of remarkable slender bones, somewhat crushed, and with- 
out trace of an articular end, which, at first sight, have the aspect 
of being portions of ribs, I am disposed to regard as the femora of 
a small Dinosaur, having probably Teleosaurian affinities. Yet such 
an identification is necessarily somewhat conjectural; and I would 
therefore state that | am led to the conclusion that the bones are 
femora (Pl. XXXI. figs. 6,7) by their slightly curved form, by 
the widening of what I take to be the proximal end, by the 
slightly smaller size of what would be the distal end, but chiefly 
by a large somewhat oblique muscular scar with an elevated border 
in its lower part, which is situate in the same position as the 
middle trochanter on the femur of a Dinosaur, which also looks 
inwards, backwards, and downwards. And I do not recognize so 
many probabilities in favour of any other interpretation, especially 
as only two bones have been found, which are right and left, 
while nearly all the other limb-bones are similarly represented 
by pairs. The longer specimen measures about 6 inches; it is 
1-3, inch wide proximally ; ; and the trochanteroid scar reaches to 
within 13 inch of the proximal end. At its upper ne the bone is 
1% inch ae The scar is 1,5 inch long, 
in its greatest width in the middle (fig. 6). It causes the son 
to swell out in thickness, so that the internal border is much 
thicker than the external border, which appears to be somewhat 
compressed, and which is slightly convex in length; while the inner 
border of the bone, but for the trochanteroid bulge in its lower 
part, would be slightly concave. As it is, it is divided into two con- 
cavities. ‘The widening of the bone at the proximal end I take to 
indicate the base of the proximal articulation. The specimen is 
there slightly incrusted with pyrites; but no trace is preserved of 
either articular head or external trochanter. The long oval of the 
lateral trochanter is defined by a ring, which is slightly elevated in 
its proximal and internal part, and much more elevated on its distal 
and anterior portion, which gives a width to the bone of 1,3, inch, 
while the width at the distal end is $f inch. The thickness of the 
bone at the distal end may be slightly diminished by accidental 
compression. It is $4 inch on the inner side, which is less than 
the thickness at the proximal end (fig. 7), where, in a corresponding 
position, the bone measures 43 inch. The inner border, both proxi- 
mally and distally, is convexly rounded ; the external border appears 
to be flattened obliquely externally in the distal half, and com- 
pressed and rounded proximally. But as the specimen is crushed, 
this point remains obscure. These femora are among the most re- 
markable bones that the Gosau formation has yielded. 


678 PROF. H. G. SEELEY ON THE REPTILE 


Dorsal Vertebra. 


There are two dorsal vertebree, both a little crushed and without 
the neural arches, which belong to such a Dinosaur. From the 
more perfect of these I draw the following characters. The cen- 
trum is 44 centim. long, with the articular ends flat, and about 
2+ centim. wide. The anterior face appears to be rather the 
flatter and larger ; but both articular margins are a little injured by 
fracture. Their edges are defined by a narrow bevelled area; the 
body of the centrum is smooth, regularly constricted, so as to be 
concave from back to front in every position below the neural arch, 
and devoid of ridges. It is more constricted at the base of the 
neural arch than elsewhere, and, except in being much more slender 
and less deep, recalls the dorsal vertebra of Anoplosaurus. The 
neural canal is similarly narrow. 


OLIGOSAURUS ADELUS, Seeley. 
(See Biinzel, pl. vi. figs. 14, 15, pl. vu. figs. 1-4.) 


The bone which Binzel figures (pl. vi. figs. 14, 15) and regards as. 
the right humerus of a lizard, I regard as the right scapula probably of 
a Dinosaur; while the specimens (pl. vii. figs. 1, 2, and 3, 4) regarded 
as right femur and fragment of humerus of lizard, I regard, from 
their correspondence in character and size, as probably referable 
to the same animal, though the femur entirely wants the tro- 
chanter which is usually seen in Dinosaurs. Taken by themselves, 
these two bones have enough in common with lizards to account for 
Biinzel’s determination ; but if the scapula is rightly associated with 
them, there can, I think, be no doubt concerning their Dinosaurian 
affinities. 

Scapula. 


The scapula is a slender compressed bone 44 centim. long, and 
imperfect at both ends, but not so much injured as materially to 
affect its characters. The inner or visceral side is slightly concave 
in length, and makes no approximation to the concave form the 
bone has in the crocodile. This inner surface is gently convex from 
side to side. The posterior margin is nearly straight, becoming 
slightly concave towards the proximal end. In adopting this no- 
menclature I have followed the crocodilian analogies rather than 
those of certain Dinosaurs. This posterior margin, as preserved, is 
33 millim. long; it is rounded, rather compressed towards the free 
end, and somewhat obliquely flattened towards the proximal end, 
where a slight ridge becomes developed, which extends towards the 
articulation, giving the bone a thickness of about 3 centim. The blade 
of the scapula is moderately concave on its anterior border, which, 
as preserved, is 3 centim. long. The width of the distal end is 
11 or 12 millim; in the middle the blade becomes constricted to a 
width of 1 centim., and then expands proximally. The proximal 
width cannot be given, on account of fracture; but the specimen, as 
preserved, is 17 millim. wide. The anterior margin is more com- 


FAUNA OF THE GOSAU FORMATION. 679 


pressed than the posterior margin. There is an indication of a ridge 
towards the proximal end, similar to that on the opposite side of 
the bone; and between these ridges, which are 13 millim. apart, 
the proximal end of the bone is concave from side to side. The bone 
does not greatly thicken at the proximal end, the greatest thickness 
towards the anterior border is 6 millim. There is a general re- 
semblance in character to the scapula attributed by Professor Owen 
to Iguanodon Mantelli (Pal. Soc. 1854, pl. xiv. fig. 1); but in this 
form the blade is not so constricted, and it may be doubted whether 
the anterior process there so marked attained any corresponding de- 
velopment in this fossil; nor is the resemblance closer to Scelido- 
saurus; and, indeed, in the straightness of its posterior margin the 
bone rather suggests the scapula of an Ichthyosawrus, in which, how- 
ever, the straight margin is anterior. 


Humerus. 


The bone which Biinzel identifies as belonging to the right side 
of the body seems to me to be referable to the left side. The 
reason for this determination is that the articular head of the bone 
is on the right side, as proved by the thickening, and the radial 
crest on the left side. Like most of the other specimens, this is im- 
perfect at both the articular ends, though enough remains to convey 
an idea of the form and length of the bone before it was mutilated. 
The shaft, which is greatly expanded proximally, is 5 centim. long. 
The distal end has the superior and inferior surfaces parallel; and 
the bone is 8 millim. thick and nearly 12 millim. wide, more 
rounded on the anterior than on the posterior border. From the 
superior outer border a faint rounded ridge extends up the shaft 
towards the middle of the articular head; and an impression occurs 
in its upper half which appears to be muscular. The width of the 
bone at the proximal end, just above this ridge, is 22 millim. In- 
ternal to the ridge the bone is compressed, so as to contribute to 
form the articular head. External to the ridge it is transversely 
expanded, somewhat flattened, and marked with strong longitudinal 
srooves at the outer extremity of the radial border. The radial 
border is more concave than the ulnar border. The thickness of the 
radial expansion is about 6 millim., but diminishes proximally. 
The thickness of the base of the proximal articulation where frac- 
tured is about 9 millim. The inferior surface is regularly concave 
from side to side at the proximal end, and convex in length, cor- 
responding to the transverse convexity and longitudinal concavity 
of the superior surface. The shaft where most constricted, in its 
lower third, is less than 1 centim. wide, while its thickness steadily 
diminishes from the distal end to the radial margin at the proximal 
end. This bone possesses none of the typical characters of a lizard- 
humerus, but all those which are usually found in Dinosaurs, though 
I am unable to name any genus in which the form of this bone is 
so closely paralleled as to suggest generic identity or even affinity. 
The distal fracture, which is transverse, is so irregular as to suggest 
an epiphysial surface from which the epiphysis has come away. 


680 PROF, H. G. SEELEY ON THE REPTILE 


Femur. 


Biinzel, though referring this specimen to the right side of tne 
body, has given no indication as to its proximal and distal ends. 
These determinations are a matter of some difficulty; but it is well 
known that, asa rule, a Dinosaurian femur is thicker on the inner 
than on the external margin, which is commonly compressed: and 
there is usually a certain compression of the bone on the posterior 
surface. I have hence been led to regard this specimen as probably 
the shaft of a left femur which has lost both the proximal and distal 
ends. The fragment is 6 centim. long; the shaft is curved more 
after the crocodilian than the Dinosaurian pattern; only it is a 
simple convex curve, without any trace of a sigmoid flexure. The 
shaft is nearly cylindrical, 1 centim. in diameter in the middle, 
where most constricted ; and there it is about 9 millim. thick. What 
I take to be the distal end expands transversely to 13 millim., is 
flattened on the superior and inner borders, flat or concave on the 
inferior border, somewhat compressed towards the external side. 
The bone is 9 millim. thick on the inner margin, and 6 millim. 
thick on the outer margin.. Proximally there is a nearly similar ex- 
pansion of the bone; only the inner border is directed well inwards, 
the external border is more flattened, and the inferior surface is 
flattened. The width of the proximal end, as preserved, is 13 centim. 
It is impossible, in the absence of more distinctive characters, to 
form any opinion as to the affinities of this specimen. A curious 
circumstance concerning it is that, while indubitably a left femur, 
either end could be regarded as proximal. 


Vertebre. 


Two vertebre are figured by Biinzel, pl. vii. figs. 2-4, which he 
regarded as indicating a foetal Dinosaur. It is difficult to discover 
any evidences that would enable one to confirm this remarkable 
speculation. The vertebre are beyond all doubt Dinosaurian; but 
except in the fact that the neural arch is lost, there is no sign of 
immaturity. The articular ends are perfectly ossified and almost 
flat; and though the specimens are a trifle worn from rolling, 1 have 
no doubt that they must be referred to a fairly well-developed 
animal. ‘The larger and better-preserved vertebra has the centrum 
14 centim. long, with the sides somewhat converging, but well 
rounded below. There is the base of a transverse process, seen in 
the middle of the side, just below the pit for the neural arch. This 
circumstance determines the vertebra as an early caudal. The 
transverse measurement over the bases of these processes is 14 millim. 
The anterior articular surface is nearly circular, 13 centim. deep, 
and nearly as wide, with the margin rounded and the articular sur- 
face nearly flat. The posterior articular end appears to have been 
rather small and rather more concave. The neural arch did not 
extend the whole length of the centrum. The neural canal is con- 
cave from front to back. There are no signs of facets for chevron 
bones. 


ee 


FAUNA OF THE GOSAU FORMATION. 681 


The second vertebra is about 1} centim. long, but had a keel on 
the base of the centrum. It appears to have been smaller than the 
other ; but, as the dorsal half of the centrum is not preserved, it may 
be that this is a similar caudal vertebra. 

There is no evidence that these vertebre belong to the animal 
which possessed the limb-bones; and I have noticed them here be- 
cause they are not obviously referable to any of the other species 
described. 

HopiosavRvs IscHyrvs, Seeley. 


In June 1880 I received word from Prof. Suess that another 
collection of Cretaceous reptile remains from the Gosau beds existed, 
which was discovered long before the publication of Dr. Biinzel’s 
paper. Eventually this small series was secured by Prof. Suess, 
and forwarded to me. ‘The specimens are imbedded in a hard 
calcareous clay, and are in a bad state of preservation, having 
been fractured in the contortion of the rocks to an extent that 
often makes identification of the fragments.extremely difficult. 
Prof. Suess remarks of the collection, “It is not very much—a few 
broken articular ends of limbs, a number of those thick roof-shaped 
dermal plates well known to you, and a few fragments of vertebree— 
the whole imbedded in a number of hard fragments of calcareous 
clay evidently once united together, and clearly belonging to the 
same individual; so they may be of some interest as showing to 
which limb-bones the dermal plates belong.” They are certainly 
the most unpromising set of fragments that I ever examined with a 
view to the study of a new type of life; and yet they certainly in- 
dicate a different species from any of which the other collections give 
evidence. I have removed the matrix, and offer a few notes on some 
of the more characteristic fragments. ; 


Proximal end of right Humerus. 


This fragment indicates a humerus very different from any thing 
with which I am acquainted, but perhaps makes a near approxima- 
tion to that reptilian humerus described by Mr. Hulke as probably 
referable to Hylwosawrus. It has lost the radial crest, which appears 
to have been reflected downward more conspicuously than in Mr. 
Hulke’s specimen, and to have presented more of the Crocodilian 
conformation, though it extended up to the articular head of the bone, 
as is usual in Dinosaurs. The superior surface has been fractured, 
so that the entire thickness of the humeral articulation is not de- 
monstrated ; but it was evidently oblong, and measures 63 centim. in 
width, and, as preserved, is 4 centim. thick just intermal to the 
radial crest, and over 3 centim. thick above the ulnar tuberosity. 
It appears to have been comparatively flat, and is remarkably pitted 
with the evidences of a cartilaginous epiphysis. It appears to be 
inclined at a considerable angle towards the radial side of the bone. 
Towards the ulnar side it is rounded. On the ulnar margin it is con- 
stricted ; and a portion of a tuberosity remains on the inferior sur- 
face which is 2 or 3 centim. below the articular head, and increases 


682 PROF. H. G, SEELEY ON THE REPTILE 


the transverse width of the bone to about 8 centim., thus giving to 
the head of the bone a blunt wedge-shaped aspect ; a concavity com- 
pletely separates the tuberosity on the inferior surface from the 
articular head. Unfortunately I have not found it possible to clear 
away the matrix from the inferior surface, except so far as was 
necessary to show that there was a concavity behind the radial crest. 
The radial side of the bone is flattened, but a little concave from side 
to side on its external aspect. It forms a considerable angle with 
the superior face of the bone, from which it is divided by the ridge 
or tuberosity which formed the ball-like part of the articulation ; 
though this ball is fractured, it was placed conspicuously towards 
the radial side; and beyond it is a longitudinal concavity.. The 
fragment is only about 74 centim. long, and the fracture is not sharp; 
but it shows that the shaft was becoming remarkably flattened and 
compressed, especially towards the ulnar side. 

The specimen thus presents a very marked difference from the 
humerus of Anoplosaurus, and is unlike that of any other genus, 
especially in the constriction which defines the ulnar tuberosity and 
in the angle which it makes with the head of the bone. 

There is a small fragment which might well be a part of the 
middle of the shaft of this specimen, too imperfect for description ; 
but if it really pertained to this bone, it may be interesting as show- 
ing a thickness of nearly 23 centim. 

A third fragment, very imperfectly preserved, I am disposed to 
regard as the distal end of the right humerus, though it is so imper- 
fect that I cannot speak confidently on the matter. It only shows 
one condyle, which is almost globular, with the flattened lateral 
margin of the bone and a small adjacent part of the superior surface 
of the shaft. This condyle, however, is remarkably massive, is 
rounded, less than 4 centim. thick, and 43 centim. wide, with a 
rugous articular surface extending internally parallel to the external 
lateral surface of the bone, and indicating, if I have correctly guessed 
the nature of the fragment, that the ulna developed a process received 
between the condyles in an unusual manner. The bone was evidently 
concave from side to side in front and greatly compressed between 
the condyles, very much as Mr: Hulke has represented in Hylco- 
saurus; it was thick on the ulnar side, and evidently more com- 
pressed on the radial side. 


Articular ends of Scapule. 


The scapule are fractured so that the entire blade of the bone is 
lost as well as the articular surface for the coracoid ; so that nothing 
remains but the articular region. The left scapula is rather more 
perfect than the right. The specimens have, at first sight, rather 
the appearance of articular ends of caudal vertebre than of scapulee. 
The left specimen shows the articulation to have been an elongate 
ovoid with the internal margin of the articulation convex and the 
external margin more flattened. The extreme width is 43 centim., 
and the length about 63 centim., as preserved. The articular sur- 


FAUNA OF THE GOSAU FORMATION. 683 
face was moderately rough, with the margin slightly rounded; and 
it was concave from front to back. Just above the posterior rounded 
termination of the articulation the bone is thickened so as to form a 
very slight tuberosity, most conspicuous on the internal surface, 
where a groove divides it from the articular head. This inferior 
surface of the bone is convex from side to side, but soon becomes 
compressed, so that the blade, where fractured, at 4 centim. from the 
articulation, is less than 2 centim. thick. On the external surface 
the specimen is concave above the articulation, partly because there 
is a Slight rounded ridge on the posterior edge, not, however, separable 
from the rounded posterior border, and partly because the bone is 
becoming thickened anteriorly, evidently in relation to the develop- 
ment of a strong spine or crest at about 2 or 3 centim. above the 
articular surface. The articular margin is marked with longitudinal 
lines of ligamentous attachment. It is of course impossible to speak 
of the affinities presented by this form of scapula; but the bone ap- 
pears to me to be distinct from the scapula of Cratcwomus, Anoplo- 
saurus, Iguanodon, Scelidosaurus, and other types with which it 
might be supposed to be allied. 

A flat expanded bone, very imperfectly preserved, appears to be 
a portion of a coracoid. It has but one margin remaining, which is 
straight for a length of upwards of 5 centim. and indicates an arti- 
cular surface at right angles to the bone, such as may have adjoined 
the scapula. The specimen is upwards of 7 centim. broad and 
10 centim. wide, as preserved, but is too imperfect for description. 


Vertebre. 


The vertebral fragments have shared in the general fracturing. 
There are two portions of sacrum which had thoroughly decomposed 
before fossilization. One fragment indicates two vertebree anky- 
losed together; the other fragment I regard as showing the base of 
the centrum of another vertebra, and on this evidence should infer 
that there were at least three, and probably four, sacral vertebree, 
though the second specimen is imperfect in every direction, so that 
it has to be regarded carefully to be accepted as a vertebra at 
all. This fragment, as preserved, is 5 centim. long, broad, rounded 
on the under side and somewhat flattened, with a shallow median 
depression which is fully 1 centim. wide. 

The same characters are shown in one of the ankylosed vertebre. 
These also are imperfect at both ends, so that it is impossible to judge 
what the length of each centrum may have been; but, as preserved, 
the length of the two together is a little over 5 centim.; and 3 centim. 
of this belong to what I regard as the more anterior of the two. This 
vertebra is very slightly concave from front to back, and convex from 
side to side, with the aspect of being flattened on the under side and 
with a very slight median depression, which becomes more marked in 
the second vertebra. ‘The sides are rounded and concavely impressed 
between the transverse processes. This appears to have been given 
off at the suture between the two vertebre, which is somewhat ele- 


684 PROF, H. G. SEELEY ON THE REPTILE 


vated and expands laterally—though, from the fractured state of the 
specimen, it is impossible to state the width across the vertebra, 


which, as preserved, is only 5 centim., but may perhaps have been — 


somewhat more. The centrum appears to have been depressed, as 
usual in the sacral region, but there is no fracture showing its exact 
thickness. This form of vertebra is very different from that referred 
to Iguanodon by Prof. Owen (Pal. Soc. 1854, pl. 3), in which the base 
of the centrum is a sharp ridge. It is also unlike, but nearer to, the 
centrum represented by the same author, pl.7, and referred to a young 
Iguanodon, in which the base of the centrum is flat, with a sharp 
ridge margining each side. 

There are two vertebre which appear to be caudal—one from 
the early part of the tail, the other from the later part of the tail. 
Both appear to have been equally long; and both are crushed. The 
earlier caudal vertebra, as preserved, has the centrum about 45 centim. 
long, and 22 millim. wide in the middle of the base. The anterior 
face appears to have been concave, and, as preserved, is 43 centim. 
deep; the depth of the posterior end of the bone is somewhat less. 
The base is flattened, though concave in length, and separated from 
the sides by a sharp angular ridge. The sides of the bone are con- 
cave. ‘The transverse process appears to have been placed on the 
hinder part of the centrum, below the neural arch, and to have had 
an antero-posterior extent of no more than 17 millim. The neural 
arch is compressed from side to side and constricted from back to 
front, with a rounded ridge margining the straight outer border of 
the neural arch, which is inclined obliquely backward. The zyga- 
pophyses are imperfectly preserved. The posterior zygapophyses 
extend beyond the centrum ; of the anterior zygapophyses no trace 1s 
preserved. The later caudal vertebra has a constricted dicebox-like 
form. Its contour appears to have been five-sided at the articular 
ends, with a median ridge on the base and lateral ridges at the middle 
of the sides. The length of the centrum is about 5 centim. Its 
depth appears to be about 3 centim., and its width 34 centim. This 
I take to be at the posterior end of the bone. Only an indication 
of the neural arch is preserved ; the width of the canal increases pos- 
teriorly as usual. Another vertebra, apparently dorsal, is imbedded 
in the matrix ; but I have not felt justified i in excavating it so as fully 
to display its characteristics. 


Dermal Armour. 


The dermal armour comprises plates of two patterns :—first, more 
or less circular plates of moderate thickness; and angular plates 
which are thick at one margin, thin at the opposite margin, and have 
the surface concave in one direction and convex in the other. They 
are flat on the under side. One or two of the small biscuit-like 
plates present that curious angular combination of fibres that is cha- 
racteristic of the armour referred to Hyleosawrus, a character which 
rather suggests the etched surface of a meteorite than the structure 
of ivory. I have been led to think that this armour of Hyleosaurian 


FAUNA OF THE GOSAU FORMATION. 685 


character is probably abdominal, or at least was not placed in a po- 
sition so exposed on the-sides of the body as the larger and thicker 
scutes. One of the plates, as preserved, is 6 centim. “long, less than 
5 centim. wide, has the margin rounded, and not more than 3 centim. 
thick (Pl. ROX L fig. 11). It shows the fibres crossing somewhat 
obliquely, so as to define slightly rhombic areas. This structure is 
suggestive of the transverse crossing of the fibres in ivory; but I 
have no evidence whether it extends into the substance of the bone. 

Another plate of about the same size appears to be rather 
thicker, and, though retaining traces of the cross angular mark- 
ings, has the surface of the scute a good deal pitted and marked in 
lines with fine vascular perforations. The larger scutes appear to 
have lost the cross striping and to show a more porous texture. 
They are remarkable for having the angle on the scute placed so 
near towards one margin as sometimes to make that side all but ver- 
tical to the base. The other side covers nearly the whole superior 
surface of the scute, and presents the curious saddle-shaped form to 
which attention has already been drawn. One of these scutes is 
63 centim. long, and has the angular crest 23 centim. high, while 
the average thickness of the plate is not much over 1 centim. In 
another specimen the crest appears to rise considerably higher, and 
gives evidence of median compression and oblique overlapping ; the 
base is slightly concave. 

Besides these remains there are portions of ribs, which are slender, 
Straight, flattened, or slightly grooved on one side and convex on the 
other, with the anterior margin of the bone rounded and the posterior 
margin compressed to a sharp ridge. ‘These fragments are all short 
enough to appear straight, and have a width of about 13 centim. 
There are a few fragments which give some information about the 
characters of the articular ends of some of the larger limb-bones ; but 
they are so obscure that I have not felt justified in attempting to 
describe them; and there is a fragment which may be a portion of 
an ilium, of unusual shape. 


CrocoDILus PRoAvus, Seeley. 


The remains of a proccelous Crocodile, some of which were figured 
by Biinzel (pl. i.) comprise 2 cervical vertebrae, 8 dorsal, one lumbar, 
one sacral, an early and a late caudal, portions of two iliac bones 
(which unfortunately exhibit no characters of value), the proximal 
end of a femur showing the typical Crocodilian characters in a very 
pronounced manner, a the ulna and radius. There are Crocodilian 
teeth which have lateral ridges and striated crowns, and small suc- 
cessional teeth, and a renwal bone. 

There is also evidence of a minute Crocodile in the articular head 
of a femur. The fragment is too small for description; but it 
obviously indicates a distinct species from the larger remains, as 
shown by the greater forward curvature of the head, its relatively 
greater width, atid the stronger development of the saath crest on 
ihe under fide of the anverles surface. 


686 PROF. H. G. SEELEY ON THE REPTILE 


Femur. 


The proximal fragment of the femur (Pl. XXIX. figs. 7, 8), 
which may have belonged to the same animal as the vertebrae, 
is 6 centim. long, exhibits the usual sigmoid flexure, and is 
compressed on the external or superior surface as in living Cro- 
codiles. Unfortunately the posterior margin of the head is not 
quite complete; but from the region of the great trochanter a 
powerful muscular ridge, defined by a groove on its anterior side, 
runs obliquely downward, outward, and forward, exactly as in the 
living Alligator; only the muscular power appears here to have been 
greater. The shaft of the bone is similarly subcylindrical ; and the 
ridges are distributed in exactly the same way. The inferior surface 
of the bone shows the articular head to have been rather better de- 
veloped than in the Alligator, apparently a little broader, and cury- 
ing a little more inward. The median process is more developed, 
and extends as a distinct ridge for a short distance down the shaft. 
The muscular tuberosity which represents the lesser trochanter in 
Mammals and the trochanter of Dinosaurs generally, seen in the 
middle of the shaft, is better developed than in the recent type. 
There is an oblique oval depression with an elevated ridge behind 
it, and another ridge extending above it proximally. The posterior 
ridge runs proximally towards the articular head, and is separated 
from the ridge above the muscle by a groove. LHvery thing which 
distinguishes the living Crocodile is here intensified. 


Proximal end of Right Fibula. 


The specimen which Biinzel regarded (pl. vi. figs. 12, 18) as the rib 
of a lizard presents all the characters of a small crocodilian fibula, 
in which, as compared with living forms, the crocodilian attri- 
butes are somewhat intensified. But the bone-tissue is so dense 
and brilliant that it has more the aspect of the bone of a bird, Or- 
nithosaur, or Lacertilian. The fragment (Pl. XXVIII. figs. 10, 11) 
is about 32 centim. long, and has been obliquely fractured below 
the expanded head, with a minute displacement which slightly aug- 
ments the curvature of the bone. 3 

The articular head is relatively thicker than in the Mississippi 
Alligator, measuring fully 11 millim. from front to back, and 
7 millim. from side to side. The articular surface is somewhat 
saddle-shaped as in the living animal, being concave in length and 
convex in width. It is widened a little posteriorly, owing to the 
development of an oblique muscular ridge, which in the existing 
crocodile does not approach so near to the articulation and is less 
internal. The internal aspect of the bone is less flattened than in 
the Alligator, is similarly marked below the articulation with short 
muscular ridges; but they are not defined by a V-shaped area, and 
hence the longitudinal tibial ridge, which is well marked proximally, 
does not originate in a Y-shaped form. The anterior margin of the 
bone is convex from above downwards ; and there is a corresponding 
concavity in the length of the posterior outline. The external sur- 


FAUNA OF THE GOSAU FORMATION. 687 


face was presumably convex from side to side, though (owing, I think, 
to crushing) it has a longitudinal groove. There is a small muscular 
tuberosity on the outer anterior margin, as in the Alligator, just 
where the compressed and curved proximal end merges into the 
straight and nearly cylindrical shaft. The shaft at the fracture is 
7 millim. long by 6 millim. wide, and has a medullary cavity 
which is over 2 millim. in diameter (fig. 11). 


UOlna. 


With the radius figured by Biinzel, pl. vu. figs. 7, 8, occurs an 
ulna. The radius was regarded as that of a lizard. The ulna, which 
is slender and compressed, with a somewhat expanded proximal end, 
is exactly paralleled by some of the living short-snouted Crocodiles. 
The ulna (Pl. XXIX. figs. 9,10) is 63 centim. long, and referable to the 
left side of the body; it has the shaft smooth, compressed from side 
to side, and contracting and curving aseit descends from the humeral 
to the carpal surface. The articular head of the bone (fig. 10) is 
remarkably large, being in antero-posterior measurement over 17 
millim., while the width is 11 millim. towards the radial border, and 
about 1 centim. at the posterior border. This surface is somewhat 
concave from front to back. Immediately below the articulation at 
the proximal end is a deep impression, which may possibly be due 
to pressure, but may also indicate the entrance of a large vessel into 
the bone. On the superior aspect there are natural impressions in 
the middle of the side and below the inner margin of the articular 
surface. The thickness of the shaft just below the articular head is 
5 centim.; and though this thickness becomes reduced a little in the 
middle of the shaft, there is a slight thickening again towards the 
distal end. ‘The width of the shaft just below the articulation is 
1 centim.; and this width diminishes to about 6 millim. just above the 
distal articulation. On the superior surface of the distal end of the 
shaft a ridge appears at the hinder margin, which extends obliquely 
across the bone. ‘The distal end is a little crushed and twisted, sub- 
reniform, more than | centim. wide, and 6 millim. thick, and makes 
an angle of about 45° in the plane of its direction with the proximal 
end. The form of the articulation appears to include three regions— 
first a deep middle groove, external to which is a small spherical 
ball, while internal to it is a concavity. The median groove and the 
compression of the bone on the inferior margin give to the articula- 
tion somewhat of a broad V-shape. 


Radius. 


The radius is represented by both right and left bones. The speci- 
men figured by Biinzel (pl. vil. figs.7, 3) appears to me to be left. Itisa 
straight bone, 6-2 centim. long, corresponding in length with the ulna. 
The proximal end (PI.X XIX. fig. 13) of the bone is expanded trans- 
versely, with a thickened margin on the inferior border. This end of 
the bone is fully 1 centim. wide, but, as preserved, is only: 4 centim. 
thick, probably owing to crushing of the superior articular border. 


688 PROF. H. G. SEELEY ON THE REPTILE 


The shaft (fig. 11) rapidly contracts, so that at a little below the 
articulation it is hardly more than 3 centim. wide; but it gradually 
expands distally to 1 centim. The bone is compressed from above 
downward ; its thickness in the upper half of the shaft is 4 millim.; 
but it increases in the lower half of the shaft to over 5 millim., 
owing to the development of a median ridge which divides this part 
of the bone nearly equally into internal and external areas, which 
are margined laterally by ridges. The inferior surface is also 
marked towards the distal end by a slight longitudinal median ridge ; 
and the articular end of. the bone 1s subreniform, convex from side 
to side. The width of the distal end (fig. 12) is 1 centim.; and its 
thickness in the middle is about 3 centim. The distal ridges are an 
important distinctive character not found in living Crocodiles. 


Cervical Vertebre. 


The cervical vertebra are probably about the third and fourth. 
What I take to be the third is only the centrum, and, except in less 
development of the tubercle for the ribs, differs in no essential point 
from the better-preserved bone. ‘The fourth vertebra (Pl. XXX. 
figs. 6, 7) might, indeed, be the fifth, sixth, or seventh, since it pre- 
sents all the characters seen in the vertebre of the Alligator in the 
middle of the neck. It is about the size of the vertebra figured by 
myself as that of Crocodilus cantabrigiensis, but is shown to belong 
to a distinct species by the less antero-posterior length of the neural 
arch, its greater inclination forward, the stronger development of the 
hypapophysial spine, and minor characters. 

The centrum is 17 millim. long, terminates posteriorly in a well- 
rounded articular ball which is about 4 millim. long, and is margined 
below by a sharp ridge and above by an equally sharp incised groove 
(which may perhaps indicate a certain incompleteness of ossification). 
The ball is circular and as nearly as may be a centimetre in diameter. 
The cup in front is of corresponding size and form. The hypapo- 
physis is a strong compressed process which rises just in front of the 
ridge bounding the posterior articulation, and is directed forward 
and downward in front of the anterior cup, much as in the fifth 
cervical of the Mississippi Alligator. On each side of this process 
and below the middle of the side of the centrum, but rather higher 
than the corresponding process in the Alligator, is situate the parapo- 
physis or articulation for the rib. It blends with the margin of the 
anterior cup, and is a strong, compressed, triangular process with a flat- 
tened facet which looks obliquely outward and forward. Itis sepa- 
rated from the hypapophysis by the usual concave channel (fig.6); and 
a similar channel divides it from the diapophysial process on the neural . 
arch ; but in the middle of this channel, or, rather, towards its upper 
part, is a slight ridge which extends from the upper margin of the 
anterior cup backward towards the ball. The diapophysis, as usual, 
extends further outward than the parapophysis, and is a smaller 
facet supported on a process of which the hinder margin is no more 
free than in the third vertebra of the Alligator ; but the process is 


FAUNA OF THE GOSAU FORMATION. 689 


broad, and defined below and in front by an oblique groove. The 
vertical measurement across the two facets for the cervical rib is 
1 centim. The neural arch has a decided aspect of being inclined 
forward, owing to the way in which the slight rounded ridge from 
the diapophysis ascends obliquely forward towards the prezygapo- 
physis, coupled with the fact that the posterior border of the neural 
canal is straight and parallel to it. The antero-posterior measure- 
ment of the eee arch below the zygapophyses is less than a centi- 
metre. The sides of the neural arch are concave from above down- 
ward, and obliquely channelled. The preezygapophyses are strong pro- 
cesses compressed into a wedge-shape in the usual way (fig. 7), directed 
upward, forward, and outward, with circular facets, which are divided 
from each other by a broad V-shaped notch. The processes are 
united by a thin platform above the neural canal, the anterior half 
of which is excavated by a longitudinal pit which almost, or quite, 
perforates the forked depression between the zygapophyses. From 
the hinder half of the neural platform, which has a remarkably four- 
cornered appearance owing to the lateral constrictions, rises the 
neural spine. It is triangular, being compressed in front and wider 
behind, where it is vertically channelled; and the channel descends 
‘so as to divide the posterior zygapophyses from each other. The 
antero-posterior measurement of the base of the spine is } centim. 
Its height, as preserved, from the base of the zygapophysial facet is 
1 centim. A slight ridge extends from its anterior border to the 
posterior border of the prezygapophysial facet, while in the Alli- 
gator the corresponding ridge, when it exists, is carried forward ; 
behind this ridge the platform at the base of the neural arch rounds con- 
vexly on the side of the centrum. The posterior zygapophyses (fig. 6) 
are strong wedge-like processes directed outward, only slightly ex- 
tending behind the neural spine, and not entirely behind it as in the 
Alligator. They are remarkable for the well-defined notch below 
them, something like that seen in the third and fourth vertebre of 
the Alligator, and for the tubercle above the posterior margin of the 
ovate facets, which gives the back of the zygapophyses a sce 
channelled appearance, to which the Alligator offers no parallel. 

The measurement from back to front over the zygapophyses is 
2centim. The transverse measurement over the preezygapophyses 
is about 18 millim. The side-to-side measurement of the constric- 
tion between the anterior and posterior zygapophyses is 12 millim. 

The posterior zygapophyses were evidently narrower than those in 
front by a millimetre or two. The neural canal is transversely ovate 
behind, but in front appears wider and more depressed (fig. 7), with 
a flat sail: the floor of the centrum is concave, and shows the usual 
pair of nutritive foramina. 


Dorsal Vertebre. 


There are 8 dorsal vertebre, which mostly belong to.the hinder 
part of the series. Reckoning as the first dorsal that vertebra in 
which the rib is for the first time entirely carried on the transverse 
process, I should regard the earliest vertebra preserved of this 


Q.J.G.8. No. 148. 22 


690 PROF, H. G. SEELEY ON THE REPTILE 


series as probably not later than the 6th dorsal, while it might have 
been as early as the second, which is likely to have been its true 
position. Then succeeds a centrum which may have belonged to 
about the fifth dorsal. The remaining six vertebree appear to be in 
sequence, and belong to the hinder part of the vertebral column; 
and if the number of vertebre was the same in this animal as in the 
Alligator, they might be the 7th to 12th dorsal vertebrae. The 
13th, or last dorsal or lumbar vertebra, if found separately, might 
have been referred to another genus, and perhaps to a Dinosaur. 
The first sacral vertebra is impertfectiy preserved. 

The early dorsal vertebra is distinguished, as in existing Croco- 
diles, by the sight development of the ridge, which ascends from the 
hinder border of the preezygapophysis inward and backward towards 
the neural spine. The centrum has a length of from 18 to 19 millim.; 
so that there is but little increase of length as compared with the 
cervical region. The transverse process is compressed and directed 
outward horizontally or a trifle upward. It is thickened on the in- 
ferior posterior border by a ridge which extends downward towards 
the centrum and constitutes a marked specific character, especially 
as there is a deep conical pit behind it immediately below the notch 
which separates the transverse process from the postzygapophysis. 
The antero-posterior width of the transverse process is 11 millim. 
Resting upon it in front, rather more horizontally than in existing 
Crocodiles, so far as regards its lateral direction, is the preezygapo- 
physial facet (fig. 8), which i is about 11 millim. long and 6 millim. wide. 
The transverse process is perhaps a little lower in position than the 
transverse process in the early dorsal vertebre of the Alligator, being 
on a level with the upper part of the neural canal. The ridges on 
the posterior zygapophyses are rather narrow, so as to give a some- 
what pinched aspect to the concave area on each side at the base of 
the neural spine (fig. 8). The neural spine extended backward a 
little between the diverging ridges of the posterior zygapophyses ; and 
at its base there is a pit. The centrum is evenly rounded (fig. 9), 
and both cup and ball well developed. As in the cervical region, 
the ball is marked on its upper margin by an incised groove (fig. 8) 
similar to that seen in the vertebra of the Alligator. 

The later dorsal vertebrae, as in existing Crocodiles, are charac- 
terized by the deeper depression between the anterior and posterior 
zygapophyses, and by the increased elevation of the zyg apophyses 
above the transverse process, consequent upon this process acquiring 
a somewhat lower position relatively to the neural canal. The pre- 
zygapophyses come to extend a little further forward and to look a 
little more upward and inward. The centrums increase in length 
so that what I regard as the tenthis 23 millim. long. The articular 
cup in front, perhaps, becomes a little wider; and “the base of the 
centrum is rather more flattened. ‘The compressed appearance at 
the base of the transverse process gives place to a regular concavity 
from above downward ; and the antero-posterior extent of the neural 
arch is increased, while the size of the neural canal is diminished 
and its height lessened. There also comes to be a less development 


FAUNA OF THE GOSAU FORMATION. 691 


of the inferior margin of the cup of the centrum, which gives a slight 
appearance of leaning forward to the bodies of the vertebre. The 
twelfth centrum is 22 miilim. long, and has the cup 13 millim. wide 
in front, showing an increase in the depression of the centrum simi- 
lar to that which is observed in the later dorsal vertebree of existing 
Crocodiles. 


Lumbar Vertebra. 


The lumbar vertebra (Pl. XXX. figs. 10, 11) may not have been 
the only one of its kind; it certainly did not immediately succeed 
the last dorsal, if it is supposed to have pertained to the sams 
animal; for it has the centrum much more depressed, with the zyga- 
pophysial facets perceptibly higher, but still retaining a much less 
inclined position than is seen in the Alligator. The cup in front of 
the centrum (fig. 10) is 1} centim. wide, and, as preserved, 1 centim.in 
vertical diameter ; so that it has a transversely oval form, which gives 
a broad aspect to the base of the centrum. The posterior ball has a 
similar pterodactyle-like transverse extension, but, as in the earlier 
vertebree of the series, is margined above by the characteristic groove. 
The transverse process is only 7 millim. wide at the base, and was 
therefore small, hke the corresponding process in the Alligator. 
In transverse section (fig. 11) it is somewhat T-shaped, owing to the 
development of an inferior sharp vertical ridge which descends to 
the base of the neural arch, widening as it comes downward. This 
ridge curiously resembles that which is so characteristic in the neural 
arch of a Dinosaur; but it is clearly comparable to that which I have 
referred to in the early dorsal vertebra, and constitutes one of the 
distinctive characters of the species. The neural canal here enlarges 
again as though in anticipation of the sacral expansion. The notch 
between the posterior zygapophyses is wider in this vertebra than in 
the dorsal series; and the zygapophyses appear to be somewhat 
stronger and broader. 


Sacral Vertebra. 


The first sacral vertebra is only a fragment, the greater part of the 
neural arch being broken away, only a portion of it remaining at the 
anterior end of the left side. The centrum is remarkably oblique; it 
is 2 centim. long, and flattened on the under side, with a small median 
grooye in the anterior part. ‘The sides are rounded; the anterior cup 
is imperfectly preserved, but appears to have been more depressed 
than in the lumbar vertebra. The posterior end of the centrum is 
oblique, flattened, with a depression below the neuralsurface. It is 
narrower than the anterior end, being about 11 millim. wide, as pre- 
served, and semicircular in outline. The basal surface of the neural 
canal is smooth, straight, and concave from side toside. The neural 
arch enters into the anterior cup as in the Alligator, and gives oif at 
the anterior end of the side a massive transverse process, the frac- 
tured base of which is 13 millim. deep and 1 centim. wide, and has 
a somewhat reniform outline owing to the anterior concavity. Only 
the base of the anterior zygapophysis is preserved. 


222 


692 PROF. H. G. SEELEY ON THE REPTILE 


Caudal Vertebre. 


The early caudal vertebra was probably about the fifth or sixth. 
The late caudal vertebra would correspond with about the 23rd or 
24th in the Alligator, though the form of the centrum might be 
taken to indicate that the tail was relatively short, and probably 
contained fewer vertebrae, while, from the absence of any continued 
series, its identification as belonging to this species is open to some 
doubt. The early caudal vertebra (Pl. XXX. figs. 12-14) is of 
elongated form, compressed at the sides, narrow and somewhat flat- 
tened on the underside. The length of the centrum is 31 millim. 
The cup in front (fig. 14) is circular and 11 millim. in diameter; 
below it is an oblique hypapophysial facet. The cup is slightly 
oblique ; and the length of the base of the centrum is 22 millim. Pos- 
teriorly the outline is subquadrate, owing to the flatness of the sides 
and to the development of two facets divided by a groove below the 
articular ball (fig. 13). Unlike the Alligator, the centrum is widest 
over these facets. There is the incised margin on the upper surface 
of the ball which characterizes all the vertebree. The width of the 
posterior end of the centrum is | centim., its extreme depth 13 millim. 
The centrum has a pinched-in appearance (fig. 13) below the trans- 
verse process (fig. 12), which, as usual, was given off about the middle 
of the side, was horizontal, compressed from above downward, and, 
though convex on the underside, shows no indication of a ridge. 
The antero-posterior extent of its base is about 12 millim. The 
neural arch appears to have had avery short neural spine; and the 
ridges from the zygapophyses, instead of being directed downward to 
join the transverse process as in the Alligator, converge inward and 
backward to form the base of the neural spine in front. The ante- 
rior zygapophyses projected in front of the centrum ; the facets (fig. 12) 
looked more upward and less inward than in the Alligator ; and there 
was no notch between them exposing part of the neural surface of 
the centrum. 

The late caudal vertebra, supposing it to belong to this species, 
concerning which I feel some doubt, is about 17 millim. long, has a 
ridge in place of the transverse process, has the narrow base mar- 
gined by two sharp parallel ridges, and the posterior cup deeper 
than the anterior cup. 


Teeth. 


There are 5 Crocodilian teeth. The most important of these 
(Pl. XXVII. fig. 25) is fractured, but appears to indicate an ovate 
crown curved slightly inward, terminating on each side in a strong 
ridge, and having the exterior face marked with a number of faintly 
elevated blunt strie, with finer striz on the inner face; the crown, 
which is a little worn, is 1 centim. high, and gives indication of the 
beginning of the fang. There are two successional teeth; one of 
these has a circular base, the other an oval base; both are short and 
blunt, marked with lateral ridges and covered with numerous fine 
parallel striz. The largest is nearly 4 centim. in diameter (fig. 24). 


FAUNA OF THE GOSAU FORMATION. 693 


Another tooth has the same general character; but the crown 
appears more curved, with sharper lateral ridges and finer and more 
numerous striations. 

In the possession of lateral ridges, as well as in the striated sur- 
face of the crown, the teeth of this species approximate to those re- 
ferred by Leidy to Hyposaurus; but there is no correspondence in 
vertebral characters. ‘he vertebre referred by the same author to 
Thoracosaurus correspond in being proceelous, but differ in the posi- 
tion and character of the tubercles for the articulation of the ribs, as 
well as in the form and direction of the neural spine in the cervical 
region. Itis difficult to determine accurately the relation of the 
species to Holops ; but the absence of the hypapophysis from the early 
dorsal vertebre will exclude comparison with that form, equally with 
the contour of the neural canal. Holops, however, appears to have 
had the sharp ridge margining the parietal bone which is seen in 
the specimen figured by Biinzel. 

It is impossible to determine whether the parietal bone of Croco- 
dilian character (figured by Bunzel, pl. 1. figs. 1-2), is referable to 
this species or to some other animal. Its distinctive features are the 
sharp lateral ridges margining the temporal fosse, the fineness of 
the circular pits on the bone, and the remarkable thinness of the 
bony substance. 

That this species can remain in the genus Crocodilus is impro- 
bable; but at present I see no grounds on which to separate it. 


PLEUROPELTUS SuEssII, Seeley. 


Postfrontal Bones of a Chelonian Skull. 
Two fragmentary bones (Pl. XXVIII. figs. 8, 9), right and left, 


present on their external surface impressions of scutes, and on their 
under surface portions of a large cavity which I believe to be orbital ; 
therefore I interpret these fragments as being postfrontal bones of a 
large Chelonian of Emydian affinities; and as the skull, when com- 
plete, was obviously of large size, it may well have belonged 
to the animal indicated by the large costal plates presently to 
be described. The remarkable feature about this bone is the 
character of its sutures; for while a groove runs round the thin 
margin in which the edge of the bone terminates, and evidently 
received a sharp ridge from the adjacent bone, the margin on the 
under side is bevelled, and thus demonstrates a union of the bones 
by squamous overlap. The under surface of the bone (fig.9) is crossed 
by an oblique ridge, which divides it, as preserved, into two nearly 
equal portions. This ridge helps to form a part of the posterior 
boundary for the eye-ball, and is characteristic of Chelydra and 
allied genera. ‘The orbital surface is concave, and more resembles 
that of Chelydra serpentina than that of T'rachydoglossus; and the 
resemblance extends even to the position of the vascular foramen 
at the back of the orbit, though there may, perhaps, in the fossil be 
several small foramina in aline. This surface, as preserved, mea- 


694 PROF. H. G. SEELEY ON THE REPTILE 


sures about 4 centim. in its greatest length, and upwards of 
3 centim. in its greatest width. The bone probably extended for- 
ward no further than the middle of the orbit; and in that case but a 
small portion is lost. The area posterior to this ridge bounding the 
back of the orbit is excavated, 4 centim. long and 2 centim. wide. 
It consists of an inner oblong area, truncated externally by the 
suture for the malar bone, and margined by the bevelled edge re- 
ferred to, which is about 6 millim. wide behind and 8 millim. on the 
inner side. The superior surface of this subquadrate fragment (fig. 8) 
is convex from within outward, showing that the top of the head 
was flattened, but that it rounded into the lateral area. It is divided 
into seven areas by unusually wide and well-marked scutal grooves. 
The bone between these grooves has a roughened, somewhat pitted 
appearance, indicative of vascular structure. The scutes are espe- 
cially remarkable from their number and small size; that over the 
orbit is largest, being about 3 centim. long and more than 14 centim. 
wide. The other scutes will be best understood from the figure. 
They are irregular, subquadrate or subtriangular figures, varying in 
measurement from 1 centim. to about 14 centim. The grooves which 
define the scutes are from 2 to 3 milliim. wide. The articular sur- 
face for the malar bone (fig. 9) 1s 27 centim. long, and over a centi- 
metre deep. It is a concave shallow groove, which involves the ter- 
mination of the postorbital ridge. There is every reason to suppose 
that the squamosal articulation was a small area just behind the 
malar articulation, and that it did not extend for more than a centi- 
metre. Hence the inner and posterior bevelled articular surface, 
which becomes very thin behind, is related to union with the parietal 
bone. The posterior portion is oblique, and makes an angle of 45° 
with the interior portion, which is straight and parallel to the 
orbital margin. Three scutes are crossed by the parietal suture, 
which, indeed, is the case in the skull of the Common Turtle. In 
front of the parietal suture, commencing with the most anterior 
inner scute, is a distinct suture, which prolongs the line of the 
parietal, and therefore gave attachment to the frontal. The portion 
of the right postfrontal bone includes four scutes and a fragment 
of the inner part of the postorbital ridge, extending as far forward 
as the termination of the parietal suture. 


Oostal Plates. 


Dr. Biinzel figured a remarkable specimen, which he regarded as 
the left ilium of an animal named Danubiosaurus anceps, the sup- 
posed rib of which proved to be a Dinosaurian scapula. I find a 
second specimen of this supposed ilium, less perfect, but similar in 
character, evidently coming from the opposite side of the body. For 
reasons to be mentioned, I have no hesitation in interpreting these 
specimens as ribs to which dermal plates are ankylosed. But there 
is something very different here from what is observed in ordinary 
Chelonians ; and though I have no doubt that we have to do with 
a pair of Chelonian costal plates, the type indicated by these re- 
mainsisnew. It would, indeed, have been less startling to refer the 
specimens to any part of the skeleton of a Dinosaur or lizard than 


FAUNA OF THE GOSAU -FORMATION. 695 


to the Chelonia; for these fossils, if rightly interpreted, indicate an 
animal with a relatively larger vertebral column than any Chelonians 
now known, or at least with a vertebral column constructed upon a 
different plan. The ribs were wide, as in the marine Chelonia, and 
extended distally far beyond the limits of the plates which covered 
them. The superimposed plate is developed chiefly behind the rib ; 
its anterior margin is smooth and rounded; the posterior margin is 
not preserved, but obviously became thin. ‘The free articular end 
of the rib was massive ; and the superimposed plate extended beyond 
it proximally for an unusual distance, indicating great width for the 
intervening vertebra. ‘his portion of the dermal plate, which extends 
mesially beyond the rib, has the external surface well preserved, but 
wants the smooth ossified internal surface; and as this is absent in 
both specimens, it is possible that m the living animal this internal 
part of the plate may have been cartilaginous or united to other 
bones. Neither of the specimens gives the slightest indication either 
of external scutes or ornament, or of union to adjacent bones; and 
the external surface is such as would suggest that the bones were 
probably contained, if not beneath a muscular covering, at least 
beneath a skin which had not become specialized ; so that we have 
here an animal that in some respects recalls the Protostega of Prof. 
Cope, but differs essentially in the dermal plates being blended 
with the ribs. An allied but undescribed type from the Cambridge 
Greensand also has the costal plates separate from each other, but 
differs in having them marked with scutes. 

A short description of each of these specimens appears neces- 
sary. First, the less perfect of the two (Pl. XXX. fig. 15) 
shows a smooth external surface gently convex in length and 
somewhat convex from side to side. It consists of a thick dense 
dermal plate superimposed upon a rib, this plate probably being 
a representative of the supracostal cartilages and ossifications 
found in birds, crocodiles, and Hatterra. Immediately above the 
articular expansion of the rib, at the proximal end, the plate is a 
centimetre thick, and is defined by the density of its texture from 
the osseous matter of the rib beneath. This bone on the interior 
suriace is much eroded, but presumably extended much further to- 
wards the median line of the animal’s body, since the dermal plate 
is prolonged with a rough under surface, due to this bone having 
adhered to the bone with which it is blended; and as the plate is 
prolonged mesially, its thickness becomes reduced to one half, though 
the fractured specimen is imperfect at its margin. ‘The transverse 
width of the plate, as preserved above the expanded head of the rib, 
is about 94 centim. About 9 centim. further away from the 
middle line of the animal, a length of about 4 or 5 centim. of the 
margin of the plate is rounded; and here its union with the thin 
underlapping, transversely expanded margin of the rib is distinctly 
seen. On the opposite or posterior margin there are some faint 
indications of a lateral rounding of the margin of the dermal plate. 
At 18 centim. from the proximal end of the rib the dermal plate 
comes to an end, being broken away, and.allows the rib to project 
freely from under it, showing, I think, that although there was 


696 PROF. H. G. SEELEY ON THE REPTILE 


bony union between these two parts of the skeleton, it was a union 
established somewhat late in the animal’s life. At this point the 
bony plate is somewhat thinner than at its proximal fracture; the 
plate tapers apparently outwards, and is somewhat broken away 
from the anterior proximal margin of the rib. The plate is hence, 
as preserved, of a lanceolate form; but its outline cannot be accu- 
rately stated. 

The rib in this specimen differs from the ribs attached to the 
costal plates of living Chelonians in being coextensive with the plate 
—that is, as wide as the plate, and, as shown by the other specimen, 
as long; hence its limits are not defined distinctly in any part of 
its extent, least so in the proximal part of the bone, where the sur- 
face is convex from side to side. This convexity becomes narrower 
and relatively more elevated as the bone proceeds outward, and is 
confined to its anterior border; where the rib terminates it is over 
3 centim. wide, over 1 centim. thick, and, though compressed on 
the anterior border, is more compressed on the posterior border. 
The thickness of the combined rib and plate is about 24 centim. ; 
but even as preserved, the thickness at the fractured articular end is 
7 centim. The underpart of the bone, which is slightly displaced by 
a fracture consequent upon a minute dislocation, is convexly rounded 
from side to side, and concave from within outwards, so as to pre- 
sent a saddle-shape. ‘The transverse width of the fragment pre- 
served is about 64 centim. The anterior border is notched, as 
though for the passage of an intervertebral nerve, or from the head 
of the rib being free, as in ordinary Chelonia, though the conforma- 
tion would rather suggest the former interpretation. There is one 
curious character in evidence that the expanded lateral part of the 
rib extended further towards the middle line than the articular 
head, showing that the ribs had attained an unusual transverse de- 
velopment consequent upon the expansion ot the superimposed 
plates. The transverse width of the head of the rib is 4 centim. ; 
and here the fracture gives it an almost semicircular outline. 

The second specimen, already figured by Bunzel, pl. vi. figs. 4,5, has 
a length, as preserved, of 48 centim.; of this 39 centim. is occupied 
by the blended rib and its superimposed dermal plate, the remaining 
9 centim. consists of the extension of the plate alone beyond the head 
of the rib towards the middle line of the body. The plate, where it is 
presumed to have extended over the vertebra, is thin, and becomes 
thinner the further it extends, both mesially and laterally. Its mar- 
gins are imperfectly preserved, but have a wedge-shape. The costal 
region wants a small portion in the middle of the plate in front ; 
but throughout it has the anterior margin preserved rounded from 
below upwards, and roughened in the proximal half, as though with 
ligamentous attachment to the adjacent plate. On the upper surface 
these lines at first run in a transverse direction, and then run for- 
wards and inwards, just as on the under side they run downward 
and outward. This oblique direction is strongly suggestive of the 
oblique crossing of ordinary intercostal muscles. In length the 
anterior margin of the bone is gently concave. The posterior margin 


FAUNA OF THE GOSAU FORMATION. 697 


is, unfortunately, not preserved; and the plate is remarkably con- 
vex from before backwards above, though this convexity is more 
marked on what I take to be the posterior side than on the anterior 
side. The extreme width of the plate below the articular head of 
the rib is about 133 centim.; and it gradually tapers as it extends 
outwards. JDistally the thickness of plate and rib diminish; and at 
the extreme distal end, where the transverse measurement, as pre- 
served, is under 3 centim., the thickness of the combined plate and 
rib is less than a centimetre. The posterior half of the plate, while 
smoother externally than the anterior half, is marked with several 
short, parallel, straight, vascular grooves, which are very narrow ; 
each is about 2 centim. long. This condition leads me to suspect 
that the anterior part of the rib may have been imbedded in muscle, 
while, owing to its curvature, the middle of the plate may have had 
only a dermal covering. 

The interior or under side of the specimen has the expanded head 
of the rib broken away ; and while it was placed in the middle of 
the width of the plate as in the other specimen, the rib soon be- 
comes developed on the anterior border, being limited by a concavity 
which runs down the length of the bone, dying away with the ele- 
vation of the rib at the distal end. This principal part of the rib 
becomes narrower as it extends further outward; but the fractured 
condition of the posterior margin appears to indicate that the margin 
of the rib was prolonged as a thin film towards the adjacent plate. 

No similar remains which are referable to an animal of this kind 
have been discovered. 


Scapula of large Chelonan. 


The fragment which | identify as a portion of a right Chelonian 
scapula, indicates an animal of somewhat large size. It shows no 
trace either of the preecoracoid or the articular end of the bone, which 
had decomposed prior to fossilization, or of the distal end ; so that it 
is not a fragment giving valuable information concerning the affini- 
ties of the animal. The fragment 1s 9 centim. long, 4 centim. 
wide at the proximal end, as preserved, and 2 centim. wide at the 
distal end. The surface which { take to be posterior is smooth, 
convex from side to side, but more flattened at the distal end than 
at the wider proximal end. In length the surface is almost straight. 
The internal margin is concave and sharp, and looks as though it 
might have been produced into a preecoracoid. The external margin 
is shghtly convex, except towards the proximal end, where it is 
modified, owing to the bone bending outwards and downwards, as 
though for the formation of the articular surface. The anterior 
aspect of the bone is much rougher; and there is a ridge which 
becomes stronger towards the proximal end, thickening the bone 
on its outer part, and making it concave in length, and dividing the 
anterior aspect into a broad, flattened, inner area and a narrower 
external area. The thickness of the bone towards the proximal end, 
where fractured, is 2 centim., and the thickness towards the distal 


698 PROF, H, G. SEELEY ON THE REPTILE 


fracture 13 millim. This is not an identification in which I feel 
absolute confidence, on account of the smoothness of one side of the 
bone and the roughness of the other, which [ had not noticed in any 
Chelonian scapula. 


Emys Neumayert, Seeley. 


There occur many remains of several Chelonians of moderate size. 
I only brought to this country a selection of some of the more cha- 
racteristic fragments, which all belong to the carapace and plastron. 
It does not seem to me desirable to determine the genera from these 
specimens ; but from their general character rather than from any 
distinctive characteristics, | regard them as being Emydian. The 
specimens are as imperfect as any of the other reptile remains, and, 
as they do not differ much in size or character, are difficult to deal 
with. ‘There are, however, certain differences of texture and form 
which justify me in indicating the existence of several species. 
These are all referred provisionally to the genus Hmys, pending 
better evidence of their generic characters. ‘The species are best 
distinguished by the characters of the plastron; for the hyo- and hypo- 
plastral bones preserved may indicate, even in their fragmentary 
condition, four species. The bulk of the remains I refer to the 
largest species, which was fully 25 centim. across the carapace. 
This species is marked by the depth of the grooves which define the 
areas of the scutes, and frequently by their elevated borders. Another 
marked feature is the exceedingly fine subgranular condition of the 
bone on its external surface—a character difficult to define, but 
altogether peculiar. Of this species the plastron is represented by 
portions of the hyoplastral and hypoplastral bones, though from their 
fragmentary condition, it is not always easy to distinguish between 
these. One fragment (Pl. XXX. fig. 16) is only 73 centim. broad and 
54 centim. long ; it does not show a single sutural surface, but exhibits 
the axillary region crossed in its lower part by an oblique scutal im- 
pression which runs forwards and a little inwards till it reaches the 
inner margin of the preeaxillary scute, which is prolonged forwards 
on the superior edge as a strongly marked groove. The usual trans- 
verse scutal impression on the basal part of the hyoplastral plate 
runs a little in advance of the axilla, and, as it nears the lateral 
margin, is directed angularly forwards in the last centimetre of its 
length. This scutal impression is strongly elevated; the ascending 
axillary process was compressed and directed obliquely upwards, 
forwards, and apparently a little outwards. The thickness of the 
plate varies from 3 centim. in the inner part to 1 centim in front of 
the axillary notch. 

A second specimen, showing the anterior part of a similar right 
hyoplastral plate, may, perhaps, belong to the same species though 
to another individual: the thickness of the scute is the same; the 
elevated ridge at the scutal suture is the same; though fractured in 
front, it measures upwards of 44 centim. anterior to the trans- 
verse suture. The lateral margin of the plate is sharp, being be- 


FAUNA OF THE GOSAU FORMATION. 699 


velled above, with the bevelled area also defined internally by a 
sharp ridge, interior to which runs a slightly impressed broad prolon- 
gation of the supraaxillary impression. In the carapace no neural 
plate is preserved. The first costal plate (Pl. XX XI. fig. 13) on the left 
side is nearly perfect; it is slightly arched, rather less than 10 centim. 
long, and more than 33 centim. wide at the lateral impression of the 
first vertebral scute. It shows the oblique sutural surface for the 
nuchal plate, which has a concave border about 35 centim. long ; the 
width of the union with the first neural plate is 22 centim., but was 
probably more, as the posterior border of the moat 1s imperfect. 
The rib is not visibly distinct from the plate upon which it is sup- 
ported, as it is in some species of V’estudo; it has a well-elevated 
compressed head, 7 millim. deep and 4 millim. wide; it is placed 
obliquely, so as to look forward and outward. In the middle of 
the plate the rib has become so depressed as to be only just recog- 
nizable; it is there 1 centim. wide; but at the outer part of the 
plate its extremity is prolonged beyond the plate, to unite appa- 
rently with the marginal plate. Anterior to it, on the under 
side, the plate is excavated (Pl. XXVII. fig. 27), and the side 
of the rib roughly striated, owing to attachment of the supra- 
axillary process from the hyoplastron. This excavation extends 
inwards from the extremity of the rib for 44 centim. A fragment 
of the right plate shows its depth where it joins the first neural 
plate to be 34 centim., and the greatest depth of the bone to be 
about 4 centim. ‘There are several fragments of costal plates ; but 
they can only be identified by the scutal markings. What appears 
to be a third costal plate of the left side is impressed with the trans- 
verse border dividing the second and third vertebral scutes for a 
length of 32 centim.; and since at this point the antero-posterior 
measurement of the plate is only 3 centim., it shows that the ver- 
tebral scutes were extremely broad relatively to their length, since 
the length could not have exceeded 6 centim., while the breadth 
could hardly have been less than 10 centim. The plate is arched, 
showing that the carapace was as much elevated as in a testudinate 
Chelonian. Its extreme length, without reckoning the curve, is 
upwards of 11 centim.; following the curve, the length is nearly 
13 centim. The breadth of the plate, towards the outer margin, 
is about 34 centim. Its thickness at the proximal part is 4 millim., 
and at the distal end 3 millim. On the under side the head of the 
rib is moderately elevated; but its course down the plate is only just 
perceptible, and marked by a smoother condition. It does not ap- 
pear to have been prolonged at the distal margin. A plate from 
the hinder part of the carapace, which is imperfect, also shows the 
arched character strikingly. It is remarkable for its antero-posterior 
extent of nearly 33 centim., and appears to be the last costal plate. 
If so, it is impressed on its outer part with a vertebral scute. Hence 
T infer this animal to have had a nearly circular outline, and to 
have had the shield greatly elevated. It may be distinguished as 
Emys Newumayrt. 


700 PROF. H. G. SEELEY ON THE REPTILE 


OTHER SpEcIEs oF Emmys. 


Another species, represented by the remains of more than one 
example, is, however, known chiefly from the hypoplastral plates. 
It appears to have been a smaller species than the last, though it is 
not easy to estimate its size from the distance between the axilla, 
or from the breadth of the abdominal scutes. The hypoplas- 
tron shows some indication of the median and anterior sutural 
margins, which would indicate a broad species, after the pattern of 
the foregoing. The length of the fragment is 64 centim., and its 
breadth about 6 centim. The transverse scutal impression is from 
24 to 3 centim. behind the anterior suture. The bone is compressed 
to a sharp margin, which is prolonged as an elevated ridge for about 
1 centim. beyond the inguinal notch. The margin is nearly straight. 
I regard this ridge as indicative of a well-marked species. 

A third species is distinguished by the way in which the axillary 
and inguinal processes are obliquely overlapped. A fourth species, 
of small size, is represented by many parts of the carapace and 
plastron. 


ARMOSAURUS GRACILIS, Seeley. 
Vertebra. 


The vertebra of a lizard, figured by Bunzel, pl. vi. fig. 11, 1s very 
imperfect, and so badly drawn as to give no just idea of its cha- 
racters. It is remarkable for the perfectly globular form of the 
posterior articular ball, which is nearly 6 millim. in transverse 
measurement, and nearly 5 in vertical measurement. It is margined 
by an impressed groove, which extends further forward on the 
neural margin than on the visceral margin. The length of the cen- 
trum in the middle line is 13 millim. What remains of the an- 
terior cup is deeply excavated to correspond with the articular 
ball, with a sharp margin conspicuous on the inferior border. The 
inferior interarticular surface of the centrum is 1 centim. long; on 
its base run two parallel blunt ridges, divided by a median groove ; 
external to these ridges are two oblique impressed concave lateral 
areas, which are broad in front and narrow behind, margined 
superiorly by an oblique rounded ridge, which ascends from the 
upper margin of the articular ball towards the middle of the arti- 
culation for the rib on the anterior part of the vertebra. This arti- 
culation for the rib is a strong process, extending laterally further 
than the width of the articular cup of the centrum, is concave from 
above downwards in front, looking obliquely downward and out- 
ward, long and narrow, rounded from fore to back, and most 
elevated proximally. It carries superiorly the prezygapophysis, 
which was a large oval surface, looking upwards and a little inwards, 
placed just above the articulation for the rib, and considerably above 
the intervertebral articulation. The zygapophysis is only preserved 
on the left side, the portion which had existed on the right side 


FAUNA OF THE GOSAU FORMATION. 701 


haying disappeared before the bone came into my possession. It is 
impossible from this slender evidence to determine the affinities of 
this animal. 


OrnitHocHErRus Bunzett, Seeley. 


The remains of Ornithosaurians are unsatisfactory, being, for the 
most part, either small portions of shafts of bones, or else bones 
which have been greatly crushed. ‘The fragments of phalangeal 
bones throw no light on the structure of the animals to which they 
belong, and give no clue to specific characters. The bone-tissue, how- 
ever, is somewhat thicker than in English specimens ; and I have no 
doubt the fragments belong to a peculiar species. There is an in- 
teresting crushed proximal end of a humerus, showing the form of 
the head, the immense radial crest, and the ulnar expansion of the 
bone at the humeral articulation: and this, with some other frag- 
ments, characterized by thin texture of the bone, may, perhaps, 
indicate a second species. But although of great local interest as 
demonstrating the presence of these animals in a period of time 
in which they were so plentiful in England, these fragments are 
of no importance to the anatomist. The only specimen of import- 
ance is the articular end of the lower jaw, already described by 
Biinzel (pl. vi. figs. 6,7). This bone is obliquely fractured just 
in front of the articular end, and shows the articular surface 
and the characteristic keel beyond it. The length of the fragment 
is 34 millim. The bone is compressed from side to side; and the 
sides converge downward into a narrow rounded ridge. The ex- 
ternal surface is flattened like the internal suriace, which latter 
shows a suture with very irregular margin, nearly parallel to the 
base, and near to it, indicating that the articular bone was re- 
ceived into the angular bone. The area in front of the articula- 
tion contracts from side to side, and is rounded; but on the inner 
margin there is a large pit partly fractured through, indicating a 
pneumatic foramen. ‘The articular surface is transversely ovate in 
area, with a median ridge running obliquely backwards and outwards 
from the hind margin of this foramen. This divides the articular 
surface into a triangular concave area in front and towards the 
outer side, and a posterior groove which is best developed towards 
the inner side of the jaw. ‘This articulation perceptibly widens the 
bone at each side. Its width is 13 millim., its length 8 millim. 
Behind the articulation there is no defining border, like the sharp 
elevated ridge in front, but the surface is flattened, with the sides 
slightly converging till they terminate in the rounded extremity. 
This posterior area is directed obliquely downwards to the base of 
the bone. The inner half of its surface consists of a pneumatic 
foramen, which is 13 millim. long, and reaches forwards to the 
posterior articular groove. This jaw seems to be well distinguished 
from the species already described. 


702 PROF. H. G. SEELEY ON THE REPTILE 


CoNncLUSION. 


From this survey it appears that Dinosaurs were well represented 
in the Gosau beds. Most of the remains belong to two species of a 
quadrupedal carnivorous genus Cratwomus, which in many respects 
resembles Scelidosaurus. It is just possible that Struthiosaurus 
may prove to be the same genus, or may have possessed the teeth 
referred to Cratwomus. The genera Hoplosaurus, Oligosaurus, Rha- 
dinosaurus, and Ornithomerus are only known from a few bones 
each ; Meyalosawrus merely from teeth. It is just within the limits 
of possibility that Doratodon may prove to be the jaw of Rhadino- 
saurus; but it is not likely to belong to the Crocodile, because true 
Crocodilian teeth occur. Hence there are certainly, with the Moch- 
lodon, seven Dinosaurian genera, while there may be as many as ten 
genera. Of Crocodiles, Lizards, and Pterodactyles there are certainly 
at least one each. The Chelonians are represented by two genera 
and five species, two only of which are described. Thus the Gosau 
fauna includes in all fourteen genera and eighteen species of reptiles ; 
and there is every reason to suppose that these formed but a part of 
the Reptilia living when the deposits were formed. 

I can scarcely hope that my efforts have been in every case suc- 
cessful in determining the species to which these disjointed and 
often fragmentary bones should be referred; but I have throughout 
worked on the basis of anatomical structure, and indicated only 
such species and genera as the organization of the animals made 
inevitable. 

I have now only to express my gratitude to Professor Suess for 
his kindness in allowing me to study this collection and retain the 
specimens so long in this country; and I would also express my 
thanks to Prof. Ramsay for permission to figure the skull of Acan- 
thopholis ; and to the Council of the Royal Society for assistance in 
carrying on this research. 


APPENDIX. 


Norte on the Gosav Buns of the Nrvz Wert, Wusr of WIENER 
Nevsrapt. By Prof. Epwarp Suzss, F.M.G.S. 


Tue Gosau beds have been deposited in preexisting valleys of the 
Triassic and Rheetic portion of our North-eastern Alps, and have 
suffered so much subsequent folding and dislocation that in the 
valley of the ‘‘ Neue Welt,” the spot where the bones were gathered 
which I sent to you, several shafts pass twice through one and the 
same seam of coal. The Gosau beds usually form green slopes at 
the foot of the great mural precipices of Triassic and Rheetic limestone. 
In the Gosau valley, near Halstatt, exposures are offered by a series 


FAUNA OF THE GOSAU FORMATION. : 703 


of ravines; in the ** Neue Welt ” (south of Vienna, west of Wiener 
Neustadt) a number of coal-mines give the opportunity of following 
the succession of beds, although they are highly disturbed here; and 
I believe that the succession is not very different in the two valleys, 
notwithstanding their distance apart. 

The base of the Gosau beds is formed by a calcareous breccia of 
variable thickness, evidently the consolidated débris of the surround- 
ing mountains. 

Then follows a series of freshwater beds, sandstones, marls and a — 
few seams of coal, accompanied by freshwater Mollusca such as Mela- 
nopsis, Dejanira, Boysia, Tanalia, Cyclas, and Unio, and the remains 
of a highly heterogeneous flora, comprising a true Palm, together 
with Pecopteris Zippi, Microzama, Cunninghanuites, and leaves of a 
dicotyledonous tree resembling Magnolia, &c., evidently the ming- 
ling of the younger dicotyledonous type with a number of survi- 
ving older types. It is this horizon which has yielded the reptilian 
bones. 

Deposits of a brackish character, with Cerithiwm, Omphalia, and 
Acteonella, begin to appear above the freshwater beds, sometimes 
apparently intercalated with them and accompanied by gravel 
beds and conglomerate, sometimes also by the first true marine 
strata, usually characterized by Hippurites organisans and Nerinwa 
breincta. 

The next group is formed by a loose marly limestone ora cal- 
careous marl crammed with reef-building corals and with masses of 
Hippurites cornu-vaccnum, Hipp. sulcatus, Caprina Aguillona, 
Spherultes organisans, and a good number of highly ornamented 
Gasteropoda. This is the true French Turonian zone of Hippurites 
cornu-vaccinum. 

This zone is succeeded by a series of loose grey and marly sand- 
stones, likewise very fossiliferous. ‘The reef-building corals and 
Rudistez have disappeared or are very rare, corals being represented 
by a few species of Cyclolites, by Diplochenium lunatum and especially 
by Troechosmilia complanata. Here the first Ammonites appear. 
Natica bulbiformis, Cardium productum, Protocardia Hillana, 
Trigoma limbata, and Janira quadricostata are some of the most 
characteristic fossils. 

In some places rose-coloured limestone beds with Orbitoides and 
the remains of a small Decapod are seen, which seem to succeed 
directly to this zone, which I have sometimes named the zone of 
Trochosmilia complanata. 

The last and highest member of the Gosau beds is a series of 
sandy loose sandstone beds, containing no fossil except great 
numbers of Jnoceramus Crispi. 

I cannot, therefore, say positively that the age of the reptiles 
which you have had the kindness to study is quite exactly that of 
your Cambridge phosphate-beds ; but it is certain that they are older 
than the true Turonian deposits, and especially older than the zone 
of Hippurites cornu-vaccinum. 


704 


PROF. H. G. SEELEY ON THE REPTILE 


EXPLANATION OF PLATES XXVII.-XXXI. 


(All the figures are of the natural size, unless an enlargement is mentioned.) 


Or 


Puare XXVII. 


. Dentary bone of right ramus of lower jaw of Mochlodon Suessii (Bunzel) 


seen from above, showing tooth-sockets, symphysial curvature, and 
ascending coronoid process. 


. Separate tooth of Mochlodon Suessii from the lower jaw, showing the 


internal aspect; enlarged twice. 


. Tooth referred to the upper jaw of Mochlodon Suessiz, showing the 


ribbed external face of the crown; enlarged twice. 


. Side view of the same tooth, showing the worn internal edge of the 


crown and curved fang. 


. Left side of hinder portion of skull of S¢ruthiosawrus austriacus (drawn 


reversed for comparison with fig. 7), showing downward direction of 
occipital condyle, foramina at base of skull, plate in front of the sella 
turcica, transverse groove on roof of skull, &e. 


. The same skull seen from the front, showing the parieto-frontal suture, 


form of the parietal bone, cerebral cavity, form of the basisphenoid 
and sella turcica, &e. 


. Right side of hinder part of base of skull of Acanthopholis horridus, 


Huxley, showing the united basioccipital and basisphenoid bones, 
with the line of large nerve-foramina. (Original m Museum of 
Practical Geology.) 


. Anterior aspect of same specimen, showing posterior plate of sella turcica. 
. Dentary bone of right ramus of lower jaw referred to Cratgomus. The 


specimen is seen from above, and shows tooth-sockets along the alve- 
olar margin. 


. External aspect of same specimen, showing the large foramina below the 


alveolar margin and above the longitudinal angle. 


. Tooth referred to Cratgomus, probably from the lower jaw, showing 


cinguloid ridge at the base of the crown; enlarged twice. 


. Similar tooth, less worn, showing serrations on the right margin ; en- 


larged twice. 


. Tooth referred to the upper jaw of Crat@omus, showing bevelled edges, 


probably due to wear; enlarged twice. 


. External aspect of same specimen ; enlarged twice. 
. Tooth probably of the larger species of Crat@omus. 
. Anterior aspect of same tooth, showing cinguloid thickening on both 


sides of the crown ; enlarged twice. 


. Dorsal rib from the right side, referred to Crateomus lepidophorus, 


showing articular surfaces. 


. Transverse section from the proximal third of the same rib, showing 


transverse expansion of the superior plate and Jateral compression of 
the body of the rib. 


. Middle of shaft of left tibia referred to Crateomus lepidophorus, showing 


muscular ridges on the fibular aspect and commencement of proximal 
expansion. 
Proximal portion of right fibula of Crat@omus, showing convex tibial 


aspect. 


. Tooth referred to Megalesaurus pannoniensis; one and a half times 


natural size. [The serrations are not directed upward so much as in 
the figure. | 


. Anterior aspect of the same tooth, showing limit of the serrations. 
. Transverse section of base of same tooth, showing posterior compression. 
. Tooth of a Crocodile, with slight lateral ridges and worn crown; en- 


larged twice. 


. A smaller more compressed and curved Crocodilian tooth, showing one 


of the lateral ridges; enlarged twice. 


. Claw phalange, probably of Rhadinosaurus. 
. Internal surface of first left costal plate of Emys Newmayrt. 


14. 


ON 


FAUNA OF THE GOSAU FORMATION. 70 


Puate XXVIII. 


Proximal portion of left scapula, showing humeral articulation, pro- 
bably referable to Mochlodon Suessiz. [The articular surface is longer 
than in the figure. | 


. Dermal plate referred to Cratgomus, terminating at each end in a free 


spine. 


. Another dermal plate, with free spines at the ends and similar tubercles 


in the middle portion. 


. A dermal plate bearing a horn-like spine, also referred to Cratgomus. 


A small scute referred to Crateomus, probably from the ventral region. 


. Distal portion of right femur of Ornithomerus gracilis, showing part 


of the lateral trochanter on the inner side of the shaft. 


. Transverse section of the same bone at the proximal fracture, showing 


medullary cavity. 
Right postfrontal bone of a Chelonian, seen from above, showing the 
cranial scutes, referred to Plewropeltus Suessit. 


. Internal aspect of the same specimen, showing postorbital ridge and sur- 


faces for union with adjacent bones. 


. Proximal end of right fibula of Crocodilus proavus. 
. Transverse section of the same bone at the distal fracture. 


Puate XXIX. 


. Superior aspect of left humerus of Crateomus lepidophorus. 
. Inferior aspect of right humerus of the same species. 
. Proximal surface of left humerus, showing expansion of the radial 


crest. 


. Inferior aspect of distal end of a humerus referred to Crat@omus Paw- 


lowitschit. 


. Transverse fracture of proximal end of the same specimen, showing 


medullary cavity. 


. Side view of claw-phalange of Crateomus. 


Internal aspect of proximal end of left femur of Crocodilus proavus. 


. Outline of proximal articular surface of the same specimen. 
. Ulna of Crocodilus proavus. 

. Proximal articular surface of the same specimen. 

. Radius of Crocodilus proavus. 

. Distal end of the same bone. 

. Proximal end of the same bone. 


Pirate XXX. 


. Superior surface of parietal bone of a small Dinosaur, probably Moch- 


lodon Suessit. 


. Side view of an angular truncated dorsal piece of dermal armour of 


Cratgomus. 


. Posterior aspect of dorsal vertebra of Cratcomus Pawlowitschii, showing 


transverse processes and fractured base of the neural spine. 


. Left side of eariy caudal vertebra of Crateomus Pawlowitschii. [An 


earlier caudal exists with the short caudal rib unankylosed. | 


. Left side of dorsal vertebra of a Dinosaur, referred to Crateomus lepido- 


phorus. 


. Right side of mid cervical vertebra of Crocodilus proavus. 
. Anterior aspect of the same vertebra. 

. Posterior aspect of dorsal vertebra of Crocodilus proavus. 

. Left side of dorsal vertebra of Crocodilus proavus. 

. Anterior aspect of lumbar vertebra of Crocodilus proavus. 
. Left side of the same vertebra. 

. Left side of an early caudal vertebra of Crocodilus proavus. 
. Inferior aspect of the same vertebra. 


Anterior aspect of the same vertebra. 


SeeGes. No. 148. BUN 


706 


PROF. H. G. SEELEY ON THE REPTILE 


Fig. 15. Side view of rib and part of superimposed plate, showing the rib free 


Fig. 


from the plate at the number 15, and the great expansion of the costal 
articulation at the other end. Plewropeltus Suesstt. 


16. Right hyoplastral element of Emys Newmayri. 


Oo Ot oon fF wWhH + 


— 
iS) tS 


fit 
(se) 


Puate XXXII. 


. Anterior aspect of shaft of right femur of Crateomus Pawlowittschii, 


showing the muscular ridges. 


. Antero-external aspect of right tibia of Crateomus Pawlowitschit. 
. Thin slightly keeled dermal plate, probably lateral, of Crateomus lepi- 


dophorus. 


. Posterior and inferior aspect of left femur of Crateomus lepidophorus. 


The figure 4 is placed opposite the small lateral trochanter. 


. Anterior and superior aspect of right femur of Crateomus lepidophorus. 
. Posterior and inferior aspect of shaft of left femur of Rhadinosaurus 


alcimus. The figure 6 is placed against the lateral trochanter. 


. Outline of the proximal fracture of the same bone. 
. Antero-inferior aspect of shaft of left humerus referred to Rhadinosau- 


rus aleimus. 


. Outline of proximal fracture of the same bone. 
. Outline of distal fracture of the same bone. 
. One of the flat dermal plates of Hoplosaurus ischyrus, showing the 


cross-fibre structure. 


. Proximal end of a rib of Cratgomus Pawlowittschit, for comparison with 


fig. 17, Pl. XXVIT. Compare Bunzel, pl. iii. fig. 5. 


. Superior aspect of first costal plate of carapace of a Chelonian, Emys 


Neumayri. 


Synopsis of the Bones figured in these Plates, arranged under the 


Species to whach they are referred. 


Mocutopon Surssi (Bunzel),. 


Dentary bone, Pl. XXVII. fig. 1; teeth, figs. 2-4; scapula, Pl. XXVIII. 
fig. 1; parietal bone, Pl. XXX. fig. 1. 


STRUTHIOSAURUS AUSTRIACUS, Bunzel. 


Hinder portion of skull, Pl. XXVII. figs. 5, 6. 


ACANTHOPHOLIS HorRIDUS, Huxley. 


Hinder portion of base of skull, Pl. XXVII. figs, 7, 8. 


CraT&omMUus (species uncertain). 


Dentary bone, Pl. XXVII. figs. 9, 10; teeth, figs. 11-16. 


Cratzomus Pawtowitscui, Seeley. 


Femur, Pl. XXXI. fig. 1; tibia, fig. 2; fibula, Pl. XXVII. fig. 20; humerus, 
Pl. XXIX. figs. 4,5; dorsal vertebra, Pl. XXX. fig. 3; caudal vertebra, iowa 
dorsal rib, Pl. SOO, fig. 12; dermal armour, Pl. XXVIII. figs. 2-4. 


CRATHOMUS LEPIDOPHORUs, Seeley. 


Femur, Pl. XXXI. figs. 4, 5; tibia, Pl. XXVII. fig. 19; humerus, Pl. XXIX. 
figs. 1-3 ; dorsal vertebra, Pl. XXX. fig. 5; dorsal rib, Pl. XXVILI. figs. 17,18; 
claw-phalange, Pl. XXIX. fig.6; dermal armour, Pl. XXX. fig. 2, Pl. XXXI. 
fig. 3, Pl. XXVIII. fig. 5. 


Hopiosaurus 18cuyRus, Seeley. 


Dermal scute, Pl. XXXI. fig. 11. 


FAUNA OF THE GOSAU FORMATION. 707 


MEGALOSAURUS PANNONIENSIS, Seeley. 
Tooth, Pl. XXVII. figs. 21-23. : 


ORNITHOMERUS GRACILIS, Seeley. 
Femur, Pl. XXVIII. figs. 6, 7. 


RHADINOSAURUS ALCIMUS, Seeley. 

eee Pl. XXXI. figs. 6,7; humerus, figs.8-10; claw-phalange, Pl. XXVII. 
fig. 26. 

Crocopibus PROAVUS, Seeley. 

Vertebre, Pl. XXX. figs. 6-14; femur, Pl. XXIX. figs. 7, 8; fibula, Pl. 
XXVIII. figs. 10,11; ulna, Pl. XXIX. figs. 9,10; radius, figs. 11-13; teeth, 
figs. 24, 25. 

PLEUROPELTUS SuEssil, Seeley. 


Postfroxntal bone, Pl. XXVIII. figs. 8,9; rib, Pl. XXX. fig. 15. 


Emys Neumayrt, Seeley. 


Costal plate, Pl. XXVII. fig. 27; Pl. XXXI. fig. 13; hyoplastral plate, 
Pl. XXX. fig. 16. 


The species described which are not figured are Doratodon carcharidens, 
Binzel, Oligosaurus adelus, Seeley, Argosaurus gracilis, Seeley, and Ornitho- 
cheirus Biinzeli, Seeley. 


Discussion. 


Mr. Hurxe considered Prof. Seeley’s paper a very valuable com- 
munication, throwing, as it did, fresh light upon an important group 
of fossils the true nature of which had before been but imperfectly 
apprehended. So far as he had been able to judge from a cursory 
inspection of the fossils, he did not doubt the accuracy of Prof. 
Seeley’s interpretations. He called attention to the anterior extre- 
mity of the mandible of Mochlodon, which had sutural indications of 
a preedentary ossification, such as he thought he had seen in Hypsilo- 
phodon ; and he mentioned the difficulty which the downward exten- 
sion of the Dinosaurian inner trochanter appeared to him to offer 
to the hypothesis of its homology with the human trochanter minor, 
an extension which suggested that it might rather be homologous 
with an outgrowth of the middle part of the linea aspera to which 
the short head of the biceps is attached. 

Mr. Cuar.LeswortH remarked on the difference between the teeth 
in the upper and lower jaw of Mochlodon. 

Dr. Murti pointed out that the work of Prof. Seeley showed that 
much caution must be exercised in accepting hurried descriptions of 
genera and species from fragments, 

Prof. Boyp Dawxrns stated that his examination of the American 
collections of Secondary Saurians proved that the so-called Megalo- 
saurian type of teeth was exhibited by forms belonging to very 
different genera. : 

The AurHor agreed with Prof. Dawkins’s views concerning the 
Megalosaurian teeth, and agreed that teeth were not sufficient alone 
. for generic determinations. 


3A 2 


708 ON A CETACEAN FROM THE LOWER OLIGOCENE OF HAMPSHIRE. 


* 


43. On the Occurrence of the Remains of a Curacnan in the Lowzr 
OxieocrenE Strata of the Hampsuire Basin. By Prof. Jonn W. 
Jupp, F.R.S., Sec. G.S.; with an Apprnpix by Prof. SuEtzy, 
FE.R.S., F.G.8. (Read June 22, 1881.) 


Remains of the marine mammalia have been so seldom recorded 
from the Lower Tertiaries of Britain, that the discovery of a new 
form is of considerable interest to the geologist. Up to the present 
time the only species noticed was the Zeuglodon Wanklyni, described 
by Prof. Seeley* in 1876. The remains of this species, which were 
found in the Barton Clay, would appear to have been unfortunately 
lost. ‘The form which I have now the honour of laying before the 
Society is represented by a caudal vertebra only ; but this appears to 
present some very interesting peculiarities. 

The locality from which this specimen was obtained is Roydon, 
about a mile and a half south of Brockenhurst, in the New Forest. 
The brickyard at this place is almost the only locality in the New 
Forest at which the very interesting marine fauna of the Brockenhurst 
Series can now be collected. The beds exposed at this brickyard 
consist of sandy clays crowded with marine fossils ; they have been 
exposed to the depth of 25 feet ; but as no overlying freshwater beds 
have been seen in conjunction with them, the total thickness of the 
Brockenhurst Series cannot be determined. Judging, however, from 
the wide area over which the beds of this age have been found to 
be exposed, that thickness must be considerable. These thick 
marine strata are seen at Roydon to rest directly upon freshwater 
clays of a bright green colour and crowded with specimens of Unio 
Solandri, Sow., which doubtless belong to the Headon Series. 

That the Roydon beds belong to the same great marine series as 
the beds of Brockenhurst and Lyndhurst appears to be clearly 
proved by a comparison of the abundant fossils from the three lo- 
calities. On this point Von Konen and the late Mr. F. Edwards, 
who collected so assiduously at all those places, appear to have en- 
tertained no doubt whatever. Recently, however, an attempt has 
been made to separate the Roydon beds into two formations, and 
to assign each of these and the Brockenhurst beds to different 
geological horizons‘, on the ground that certain forms which are 


rare at one locality are abundant at the others, and wice versd. 


The fauna of all these beds is so unmistakably that of the Lower 
Oligocene or Tongrian that it is impossible to find any valid grounds 
for such a subdivision. 

Terrestrial mammalia are so abundant in the overlying Bem- 
bridge beds, as well as in the underlying Headons, that the disco- 
very of this marine form in the Brockenhurst Series is of much 


* Quart. Journ. Geol. Soc. vol. xxxii. p. 428, 1876. 
Tt Quart. Journ. Geol. Soc. vol. xxxvi. p. 113. 


mr 
= oe 


= 


a eee ey es ae eS ee a ee 


ee ot 


ON THE CAUDAL VERTEBRA OF A CETACEAN, 709 


interest, affording, as it does, such an important confirmation of the 
ection that a great change i in the physical conditions of the area 
must have pecurned at the time when these marine strata were 
deposited. 


APPENDIX. 


Note on the Cavpat VurrEesra of a Curacean discovered by Prof. 
Jupp im the Brocxenuurst beds, indicative of a NEw TyPE allied 
to BALZNoPTERA (BaL#nopreRs JUDD). By Prof. H. G. Szexey, 


RSs GS. 


Tuer yertebra submitted to me by Professor Judd belongs to the 
caudal region ; and although in a new type there may he some doubt 
concerning the exact place in the series, it may be affirmed to have 
been about the eighth caudal, and certainly not later than the 
twelfth. ‘The vertebra is probably distinct from all recent and 
fossil genera; but its characters are altogether in harmony with the 
Baleenide ; and the specimen indicates a genus much more closely re- 
lated to Balenoptera than to Balena, so far as can be judged from 
a single vertebra. This affinity is especially shown in the character 
of the base of the centrum, which had the facets for the chevron 
bones very small, and also in the general character of the neural 
arch, which is much less massive than in Balena. The differences 
from the great Balenoptera musculus on the other hand are clearly de- 
fined, and appear to consist in the remarkably forward position of the 
neural arch (fig. 2), sicein Balenoptera it does not usually reach the 
anterior sutural margin. The depressed transversely ovate outline of 
the neural canal (fig.3)is another distinctive character, since in Bale- 
noptera the neural canal is higher than wide. The neural spine also 
appears to be much less developed than in the recent genus, though, 
as the hinder part of the neural arch is broken away, this character 
cannot be positively affirmed. The centrum is relatively much shorter 
(fig. 2); the facets and ridges on the base connected with the chevron 
bones are less developed. The transverse width of the zygapophyses 
was relatively greater in the fossil. The transverse process is re- 
markable for extending the entire width of the centrum (fig. 2), and 
for having the vertical perforation rather behind the middle of the 
centrum (fig. 1), since in Balenoptera it is placed further forward. 
Balenoptera laticeps, however, a species of the North Sea, ranging, 
perhaps, to Java and Japan, approaches nearer to the fossil, espe- 
cially in having the epiphyses separate; the neural arch, too, is 
placed almost as far forward as in the fossil, but is higher, and the 
neural spine is larger; but otherwise its characters are very similar. 
Perhaps when we reflect that the contemporaries of this old whale 
were Palewothzrium, Anoplotherium, and their allies and congeners, 
all of which have passed away, it will not seem improbable that the 
marine mammals also should give evidence of subgeneric differences. 


710 PROF. H. G. SEELEY ON THE CAUDAL VERTEBRA 


Fig. 1.— Caudal Vertebra of Balenoptera Juddi, seen from above. 


Anterior border. 


Posterior border. 


{The vertebra does not lean forward as represented in the figure. | 


Fig. 2.—Right side of eighth (?) Caudal Vertebra of Balenoptera 
Juddi, showing anterior position of the neural arch and vertical 
perforation of the transverse process. 


i, 


i 
My, 
4, 


OF A CETACEAN FROM THE BROCKENHURST BEDS. Flat 


Fig. 3.—Anterior Aspect of Caudal Vertebre of Balenoptera Juddi, 
showing the depressed form of the neural arch and sutural sur- 
face, from which the vertebral epiphysis has come away. 


A tail vertebra of a whale, however, is about the last thing which an 
anatomist would select to furnish characters for a new genus. I have 
not thought that attention is more likely to be called to the fossil by 
thus dealing with it than would be the case by adopting the safer 
course of shrinking from the responsibility of indicating a genus 
which could not at present be sustained. 

The centrum has lost its epiphyses (fig. 3); each end is hexagonal, 
and marked with the usual radiating longitudinal rugosities. The 
centrum is longer at the superior margin, which measures nearly 
7 centim., than at the inferior margin, where the measurement is 
6:2 centim.; so that there is a conspicuous leaning forward of the 
superior margin of the vertebra (fig. 2). - Among vertebrates this cha- 
racter recalls a characteristic condition of the vertebrae of Pliosaurus. 
As already remarked, the face of the centrum is hexagonal; it is 
10-2 centim. wide, and 8-7 centim. deep. ‘The superior surface of the 
centrum is short, since the transverse outside measurement of the 
neural arch is only 4 centim.; and the width of the base of the cen- 
trum is probably no more, though its anterior and posterior margins 
are abraded, so that the exact characters cannot be stated. The lateral 
surfaces are unequally divided by the transverse processes (fig. 3), 
the superior pair, 5°3 centim. long, being the longest. Their mar- 
gins are moderately convex ; the inferior lateral borders were under 
5 centim. long. The posterior surface of the centrum is broken on 
the superior, inferior, and left lateral margins. The lateral margins 


AMD ON THE CAUDAL VERTEBRA OF A CETACEAN. 


appear to have been more convex behind than in front, and the 
transverse width of the centrum appears to have been greater, about 
10°6 centim. The base of the centrum is narrow, with a median 
longitudinal depression more marked beneath than in front, but 
not very conspicuous, and yet sufficient to give the base the appear- 
ance of being formed of two rounded lateral portions. The basal 
surface is concave from front to back: it is impossible to state its 
trausverse width in the middle, because there are no lateral limiting 
lines ; but it exceeds 25 centim. ‘The inferior lateral surfaces are 
nearly flat from above downward, the concavity being scarcely ap- 
preciable, but are markedly concave from front to back. The supe- 
rior lateral surfaces are convex from above downwards (fig. 2), the 
convexity being least marked in the middle and more marked on the 
posterior border than on the anterior border. The length of the 
transverse processes cannot be given, because they are fractured ; but 
since the fractured end is only half a centim. thick, and 4:7 centim. 
wide, it was presumably short. This process is perforated vertically 
by an oval foramen (fig.1) which is 7 millim. long and 5 millim. wide. 
Its anterior margin is distant 3:4 centim. from the anterior border 
of. the vertebra ; and the posterior margin is 2°5 centim. from the 
posterior border. On the left side the foramen appears to bea little 
smaller, and nearer to the middle of the centrum. 

The neural arch is small, depressed, placed anteriorly so as somewhat 
to overhang the centrum (fig. 1). It has a scarcely appreciable neural 
spine(fig.2); butits posterior margin is broken away. The sides of the © 
neural arch conyerge posteriorly, so that the transverse measurement 
behind, as preserved, is 3 centim. The antero-posterior measurement 
of the base of the neural arch is rather more than 4 centim. (fig. 2). 
The anterior zygapophyses, if such existed, are broken away; and the 
extreme transverse width of the neural arch in front (fig. 3), which is 
obviously less than its width when perfect, is about 5 centim. ‘The 
sides of the arch are directed upward and outward ; and the neur- 
apophyses are compressed. The canal for the spinal cord is wider in 
front than behind, and higher behind than in front. The transverse 
measurement in front is 2°7 centim. The height in front is 1:4 
centim. The height behind is about 1:6, as preserved, and the width 
fully 2 centim. The neural canal is concave from front to back, 
and contains several nutritive foramina. Two are placed near 
to the anterior margin, and one in the middle line behind the 
middle of the centrum. The superior surface of the neural arch was 
probably notched in front, where it was concave from side to side ; 
it was inclined backward, where the sides converge, and where it is 
divided mesially by the faint indication of a neural spine (fig. 1). 


&. H. HOLLINGWORTH ON A PEAT BED AT OLDHAM. FAS. 


44. Description of a Prat Bev interstratified with the Bov.pER- 
Drirt at OrpHam. By Geo. H. Hotiineworrs, Esq., F.G.S. 
(Read June 22, 1881.) 


In the early part of September last Mr. James Nield, of Oldham, 
called my attention to a bed of peat which he had discovered on the 
site of the railway-extensions then being made by the London and 
North-Western Railway Co., at Rhodes Bank, Oldham. 

The length of the section exposed to view was 53 feet in a north- 
to-south direction, and the maximum depth 14 feet; but the depth 
of the drift, as proved at the colliery about 100 yards to the west, 
is 344 feet. 


Section of Boulder-clay with interstratified Peat, near Rhodes Bank, 
Oldham. (Scale about 14 feet to 1 inch.) 


N. ae 


& 
SUS Sees 


SSe 


aie ieee 
ee ate, "08 ee 


e ies Ee a Sh Reb Sane Manet crea nm reer a . 
oe ty hepa) ee Goa —— = > a en F450 Se Se TS ee 
mee : S x a? . 9: : ao ene . . > 
Pe a Oa 2S patocls Ages Aes A EN noe fae ee, £ o g ole. 
. . a ee . K co s - e = 
ONC Ie ne On Nan) ga Qi ieee (Vidar wn Sitar On ala. A" a.7 2). 0S VZo Sy 
Gray io ao RG! Ae Tae a ies ER toe &-0 962% a DO Sees ay of = 
Cele omeyn et <2 S79) Silo Ao 8 4 atin See CNOA Ca po eres aed 
° ai) 5 ea ae - Bie . ° 
Dix anon fo} tescve Y Sean = See a02: S.2% Ge ay vs cftec ROY > fe “9 Co 2 =o cy ts) 9 . =~ 
es TOR MOR ae O14) Oe eles fel fal ann ISS OA! Kye SIC Soa S26 Ae NSO - ec Ses 
= ania S OE) Cheat. crala oa Sen pT eOLD ee oe — ° 
: "0 J -. ° 2 > pa 5 ec ont BID Sars . °o_ ogo - seenreh*) Cr awe 
Pe Sete SO a SE ie eNOS On OI NO CP Scio Ere GoM ec i Bic 
° 5 
a. Soil. 6. Boulder-clay. c. Fine blue clay. 
d. Peat. e. Sand. f. Ochre. 


The following is the description of the beds, proceeding down- 
wards :— 


No. 
il SGU scope eanee ee eae ee BREE Ce OTS CREO eee 8 to 10 in. 
2. Boulder-clay, sometimes sandy, with beds and | . : 
2 to 6 ft. 
BUREN DORO ADEA ey esac tasctsa ssdevnserceesSeaeea duane s « 


3. Main bed of peat containing mosses, exogenous 
stems, and beetles. At one point there was en- | 2 in. to 1 ft. 9 in. 
closed in the peat a deposit of iron ochre (f) about { Average 15 in. 
6 inches in maximum thickness and 6 feet long... 

4. Blue clay, mostly very fine, but stony where] 5 ,, fouleee 


thickest, forms the floor clay or soil ............... 

5. Current-bedded coarse sand and fine gravel, red- Aen oo ce 
BMP EVEMOW-COLOULCH eteer aise. 625 sins wceceae cutee ees 

Bee OUIGder-ClAY, LOVED) 5.2 fo< tec. nto dcacecnmecsssnsecens 6 ft. 


The deposit is situate in the valley of the Medlock, about 25 feet 
above its present level; but it has not beeen exposed by that brook, 
at this point, where there is from 2 to 6 feet of the upper clay still 
resting upon the peat. The upper clay, peat, and lower clay have 
probably been denuded by the action of the brook some yards to the 


Q.J.G.S. No. 148. 3B 


714 G. H, HOLLINGWORTH ON A PEAT BED AT OLDHAM. 


west. The upper clay is continuous to the east for at least 200 
yards, and was proved to be about 4 feet thick for 15 yards to the 
east of the section by the operations of the Railway Company in 
search of sand. At that point the sand was more than 6 feet in 
thickness, the bed of peat having run out. The boulders are mostly 
from a distance (porphyries, granites, &c.); but a few are of local 
origin, one picked up being ganister from the lower coal-measures. 

The section, which is now covered up, was measured by me when 
about 6 yards of the strata had been excavated from the face at 
right angles to the section. 

The peat bed terminates on the right of the section; but on the 
left or north the upper bed is going forward. ‘The floor clay (4) is 
of a fine silty nature generally, and is very similar to the floor clay 
of the Ashton Moss (Ashton-under-Lyne) as exposed in the railway- 
cutting. 

The mosses &c. have been placed in the hands of the Rev. J. 
Ferguson, of Brechin, N. B., for examination ; but his final report has 


not yet been received. They are, however, of decidedly northern 
character. 


eal 


GENERAL INDEX 


TO 


THE QUARTERLY JOURNAL 


AND 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


[The fossils referred to are described ; and those of which the names are printed 
in italics are also figured. | 


Abbey Town, section through, 296. 

Aberaeron, section from Llandeilo to, 
158. 

Aberffraw Sands, coast section from 
Porth Nobla to, 216. 

Aberystwith, constitution of grit from, 
8, 25; section from, to the Devil’s 
Bridge, 143; section from, through 
Pont Erwyd to near Builth, 158; 
vertical section of Silurian rocks at, 
164. 

— grits, 144, 149, 164. 

Abnormal geological deposits, Mr. C. 
Moore on, in the Bristol district, 
67. 

Acanthograptus ramosus, 174. 

Acanthopholis horridus, 634. 

Adriosaurus Suessti, 55. 

eee Jelinus, Prof. Owen on, 

61 


Adolian Sands, 19. 

Aitheotesta devonica, 306. 

Africa, South, Mr. Dunn on the Dia- 
mond-fields of, 609. 

African Desert, sand of the, 19. 

Age of Archzan rocks of Anglesey, 
229. 

Alderley Edge, cupreous sandstones 
of, 15. 

Alum Bay, sands from, 18. 

Amaltheus subspinatus, 605. 

Amlwch, contorted rocks near, 223; 
slates, 223; structure of slaty rock 
from near, 235. 

Ammonites, Prof. Buckman, on the 
terminations of some, from the In- 


ferior Oolitesof Dorsetand Somerset, 


Ammonites, Mr. 8. 8. Buckman on 
some, from the Inferior Oolite of 
Dorset, 588. 

Ammonites boscensis, 65. 

Braikenridgii, 61. 

— Brongniarti, 64. 

cadomensis, 607. 

— concavus, 60. 

—— Edouardianus, 61. 

— Gervillii, 63. 

—— Humphriesianus, 64. 

linguiferus, 62. 

—— Manselii, 64. 

— Martinsii, 63. 

—— Moorei, 65. 

Sauzei, 62, 

—-- subcostatus, 63. 

subradiatus, 61. 

Analysis of chloritic rock and talcose 
schist from Plas Goch, 45, note; of 
serpentine from Ty Newydd, 46, 
note; of iron-ore from Rio Tinto, 4. 

Andesite or basalt from Skomer Is- 
land, 411. 

Andrews, Rev. W. R., note on the 
strata of the Purbeck Beds of the 
vale of Wardour, 248. 

Anglesey, age of Archean rocks of, 
229. 


——, Mr. C. Callaway on the Archean 
geology of, 210. 

——, Prof. T. G. Bonney ona boulder 
of Hornblende Picrite near Pen-y- 
Carnisiog, 137. 


716 


Anglesey, Prof. T. G. Bonney on the 
microscopic structure of some rocks 
from, 282. 

Anniversary Address of the President, 
Proc. 37. See also Htheridge, R., 
Esq. 

Annual Report for 1880, Proce. 8. 

Anodontacantius, Mr. J. W. Davis on, 
427. 

Anodontacanthus acutus, 428. 

fastigiatus, 428. 

obtusus, 428. 

Anomodont reptile (Platypodosaurus 
robustus), Prof. Owen on the skeleton 
of an: Part II. Pelvis, 266. 

Antedon tmpressa, 135. 

Arabia Petrza, sand from, 19. 

Arzosaurus gracilis, 700. 

Archean geology of Anglesey, Mr. C. 
Callaway on the, 210. 

Archeopteris sp., 305. 

Ardennes, Upper Jurassic rocks of 
the, 501. 

Arenig-felstone and Hskdale-granite 
dispersions, junction of the, 361. 

Armangon, map of the valley of the, 


Ascodictya and Stomatopore. Mr. 
Vine on, 613. 

Ascodictyon stellatum, 618. 

radians, 618. 

Ashdown Sands, constitution of, 16. 

Ashton, vein at, 77. 

Assynt and Durness, Mr. C. Callaway 
on the Limestone of, 239. 

Asteropteris noveboracensis, 299, 307. 

Astroconia Granti, Mr. W. J. Sollas 
on, anew Lyssakine Hexactinellid 
from the Silurian formation of 
Canada, 254. 

Athabasea river, British Columbia, 
276. 

Aube, Upper Jurassic rocks of the, 
525 ; map of the valley of the, 526. 

Augill, near Brough, Sandstone from, 
12 


Aust, near Bristol, Mr. J. W. Davis 
on the Fish-remains of the Bone- 
bed at, 414. 

Australia, Devonian plant from, 306. 

Auxerre, map of the Yonne valley 
south of, 534. 

Avon, view of the gorge of the, near 
Clifton, 76; Conglomerates of the 
gorge of the, 76. 


Bagilt, Flintshire, Glacial sands from, 
19. 
Balenoptera Juddi, 709. 


Barmouth, Cambrian Grits of, 7, 25. 
Barrah Hill, Section through, 194. 


GENERAL INDEX. 


Barrois, Dr. C., Award of the Bigsby 
Medal to, Proce. 32. 

Basalt or Andesite from Skomer Is- 
land, 411. 

Bay de Chaleurs, Devonian plants 
from, 307. 

Beddgelert and Snowdon, Mr. F. 
Rutley on devitrified rocks from, 


——, devitrified spherulitic rock from, 
03 


Benches in British Columbia, 274. 

Bigsby Medal, Award of the, to Dr. 
C. Barrois, Proc. 32. 

Blake, Rev. J. F., on the correlation 
of the Upper Jurassic rocks of 
England with those of the Continent, 
497. 

Bluen y ciel, section through, 159. 

Bodafon Mountain, gneissic rocks of, 
219. 

——, quartz-schist from summit of, 
233. 

Bodorgan, section on the railway from 
Ty Croes to, 217. 

——, structure of igneous rock from 
a green shale near, 236. 

Bodwrog, structure of 
limestone from near, 236. 

Bolonian of the Boulonnais and HEng- 
land, 581. 

Bone-bed, Mr. J. W. Davis on the 
fish-remains of the, at Aust, near 
Bristol. 

Bonney, Prof. T. G., on a boulder of 
hornblende picrite near Pen-y- 
Carnisiog, Anglesey, 137. 

, on the microscopic structure of 
some Anglesey rocks, 233. 

——, on the serpentine and associated 
rocks of Anglesey, with a note on 
the so-called serpentine of Porth- 
dinlleyn (Caernarvonshire), 40. 

Bootle bore-hole,constitution of Bunter 
sandstone from, 13. 

Borth Saint, structure of quartz-schist 
from, 233. 

Boulder of Hornblende Picrite near 
Pen-y-Carnisiog, Anglesey, Prof. T. 
G. Bonney on a, 137. 

Boulder-clay of West Cumberland and 
North Lancashire, 29. 

Boulder-drift, Mr. Hollingworth on a 
peat-bed interstratified with the, 
713. 

Boulders, position of, in the Moel- 
Tryfan deposits, 354; position of, 
in Frondeg gravel-pits, 360. 

Boulonnais, Upper Jurassic rocks 
of the, 557; map of part of the, 
559. 


crystalline 


GENERAL INDEX. 


Bourges, map of country round, 540, 

Bovey Heathfield, sands from, 19. 

Bradford, constitution of Sandstone 
from Spinkwell Quarry near, 11. 

Bradford Abbas, vertical section of 
Inferior Oolite at, 590. 

Braich, section of drift-deposits be- 
tween, and the summit of the Ridge, 
Frondeg, Denbighshire, 367. 

Brampton, section from the Solway to 
near, 296. 

Brigham, Sandstone from, constitution 
of, 11; analysis of, 21. 

Bristol district, Mr. C. Moore on ab- 
poseal geological deposits in the, 


Bristol Reptilia, age of the, 80. 

British Columbia, Dr. G. Dawson on 
the superficial geology of, 272. 

Brockenhurst fauna, affinities of the, 
110; relation of Colwell marine 
fauna to, 112. ; 


Brockenhurst zone at Whitecliff Bay - 


and in the New Forest, 109; fossils 
of the, 115. 

Brodie, Rev. P. B., on certain quartzite 
and sandstone fossiliferous pebbles 
in the Drift in Warwickshire, and 
their probable identity, lthologi- 
cally and zoologically, with the true 
Lower-Silurian pebbles with similar 
fossils in the Trias at Budleigh Sal- 
terton, Devonshire, 430. 

Bryn-y-Carnau quarry, fossils from. 
145. 

Bryozoa, fossil Chilostomatous, from 
South-west Victoria, Mr. A. W. 
Waters on, 309. 

Buckman, J., Hsq., on the terminations 
of some Ammonites from the Infe- 

rior Oolite of Dorset and Somerset, 

57. 

Buckman, S. S., Esq., on some species 
of Ammonites from the Inferior 
Oolite of Dorset, 588. 

Budleigh Salterton, Lower-Silurian 
pebbles in the Trias at, 430. 

Bunter sandstones, constitution of, 12. 


Caberea rudis, 322. 

Caernaryonshire, high-level gravel and 
sand in, 357. 

Caerwen Farm, section through, 159. 

Callaway, Dr. C., on the Archeean geo- 
logy of Anglesey, 210. 

, on the limestone of Durness and 
Assynt, 239. 

Calyptograptus digitatus, 174. 

plumosus, 173, 

Cambrian grits and sandstones, 7. 

Canada, Prof. W. J. Sollas on Astvo- 


717 


conia Granti from the Silurian for- 
mation of, 254. 

Canda fossilis, 322. 

Caradoc Waterfall and Traws Coed, 
section in railway-cutting between, 
Gy 

Carboniferous Fenestellidz, Mr. G. 
Shrubsole on the, 178. 

grits and sandstones, 11, 25. 

Cardigan, 162. 

Cardiganshire, South-west, vertical 
section of Silurian rocks in, 164. 

Cardiopteris ertana, 305. 

Cardita-Beaumonti beds of Sind, 193 ; 
corals of the, 197. 

Carlisle Basin, Mr. T. V. Holmes on 
the Permian, Triassic, and Liassic 
rocks of the, 286. 

Carpenter, P. H., Hsq., on two new 
Crinoids from the Upper Chalk of 
Southern Sweden, 128. 

Carrall, J. W., Esq., on the locality of 
some fossils found in the Carboni- 
ferous rocks at T’ang Shan, China,83. 

Carreg-winllan, structure of igneous 
rock from, 236. 

Carruthers, W., Esq,, identification of 
plants in brown iron-ore from Rio 
Tinto, 4. 

Carstone, of Hunstanton, constitution 
of, 17; analysis of, 18. 

Cas Clock, characters of chlorite- 
schists from, 234. 

Catenicella alata, 317. 

ampla, 317. 

cribriformis, 317. 

elegans, 317. 

flexuosa, 317. 

— mternodia, 318. 

—— marginata, 317. 

solida, 318. 

Caulopteris Lockwoodi, 307. 

Cefn Coch, fossils from, 152. 

Hendre, fossils from, 145. 

Cellaria fistulosa, 319. 

globulosa, 321. 

malvinensis, 321. 

ovicellosa, 321. 

Cellepora fossa, 343. 

yarraensis, 343. 

sp., d44. 

Celluloxylon primevum, 302. 

Cemmaes, structure of crystalline 
limestone from near, 236. 

limestones, 223. 

Central Wales, Mr. W. Keeping on 
the geology of, 141. 

Central zone of Anglesey, 218. 

Cerrig Ceimwen, slaty group of, 215; 
structure of slaty rock from near, 
234, 


> 


718 GENERAL INDEX. 


Cerrig ddwyffordd, structure of slaty 
rock from, 234, 235. 

Ceryg Moelion, serpentine near, 42 ; 
section of quarry at, 43; micro- 
scopic character of rock from, 46. 

Cetacean, Prof. Judd on the remains 
of a, in the Lower Oligocene strata 
of the Hampshire basin, 708. 

Chalk, Upper, of Southern Sweden, 
Mr. P. H. Carpenter on two new 
Crinoids from the, 128. 

Charente, Upper Jurassic rocks of the, 
541. 


Charmouth, Mr. Sollas on a new 
species of Plesiosawrus from the 
Lower Lias of, 440. 

Cher, Upper Jurassic rocks of the, 
538. 


Cheshire, Bunter sandstones of, 12. 

Chilostomatous Bryozoa from South- 
west Victoria, Mr. A. W. Waters on 
fossil, 309. 

Chlorite schists of Anglesey, Prof, 
Bonney on the, 234. 

Chloritic marl and Upper Greensand, 
Isle of Wight, Mr. Parkinson on, 
370. 

schists of Mynydd Mechell, 222. 

Cladodus curtus, 422. 

Cladophora, Mr. C. Lapworth on the, 
in the Llandovery rocks of Mid 
Wales, 171. 

Clevedon, 78. 

Cliff End, Osborne beds at, 104. 

Clifton, Rhetic bone-bed near, 75; 
view of the gorge of the Avon near, 
76. 

Clogwyn dur Arddu, devitrified ob- 
sidian or rhyolite from, 405. 

Clwydd valley, vertical section of Si- 
lurian rocks in, 164. 

Coal-measures, Mr. J. W. Davis on 
Anodontacanthus, a new genus of 
fossil fishes from the, 427. 

Coal-measure sandstone, from Spink- 
nell Quarry, near Bradford, consti- 
tution of, 11 ; analysis of, 21; from 
Stonyhough, constitution of, 11. 

Coast of British Columbia, 277 ; map 
of the, 278. 

Columbia, British, Dr. G. Dawson on 
the superficial geology of, 272. 

Colwell Bay, Mr. H. Keeping and Mr. 
H. B. Tawney on the beds at Hea- 
don Hill and, in the Isle of Wight, 
85. 

, Lower Headon of, 101; Middle 
Headon of, 101. 

Colwell marine fauna, relation of, to 
the Brockenhurst fauna, 112. 


Colwyn Bay, Middle Glacial drift of, 
19. 


Comén, Prof. Seeley on the remains of 
a small Lizard from the Neocomian 
rocks of, 52. 

Conglomerates of the gorge of the 
Avon, 76. 

Constitution of grits and sandstones, 
Mr. J. A. Phillips on the, 6. 

Contortions in railway-cutting near 
Traws Coed, 155. 

Coppinger, Dr. R. W., on Soilcap-mo- 
tion, 348. 
Corallian of the Paris basin and Eng- 

land, 570. 

Coralliferous series of Sind, Prof. P. 
M. Duncan on the, 190. 

Corals of Sind, 197. 

Corris area, 162. 

Corwen, N. Wales, Dr. Hicks on some 
remains of plants in the Denbigh- 
shire grits at, 48. 

, section from Nant Llechos, near, 
to Moel Morfydd, 487. 

Cosmoceras Caumontii, 600. 

Garantianum, 600. 

Parkinsoni, 599. 

subfurcatum, 600. 

Craig fawr, 215. 

Craig Lluest, section through, 148. 

Crateomus, 637. 

lepidophorus, 660. 

Pawlowitschii, 642. 

Cretaceous sandstones, constitution of, 


16. 

Cribrillina dentipora, 326. 

suggerens, 327. 

terminata, 326. 

Crinoids, Mr. P. H. Carpenter on two 
new, from the Upper Chalk of 
Southern Sweden, 128. 

Crocodilus proavus, 685. 

Crogan goch, character of chlorite 
schist from, 234. 

Crossgates, interglacial deposits at, 
33, 37 

Cruglas, serpentine near, 42, 45; mi- 
croscopic character of, 46. 

Ctenoptychius Ordii, 422. 

pectinatus, 424. 

Cumberland, Mr. J. D. Kendall 
on interglacial deposits of West, 
29. 

Cummersdale, 289. 

Cwm, fossils from quarry at, 144. 

Cwm Elan, section through, 159. 

Cwm Maenydd, 163. 

Cwm Symlog, fossils from, 154. 

Cwm Ystwyth, section from, to Gwas- 
taden, 148. 


GENERAL INDEX. 


Cyclopteris sp., 305. 
Cyclostigma affine, 301. 


Dalston, section from Solway Moss to, 
296. 

Davis, J. W., Esq., on the fish-remains 
of the bone-bed at Aust, near Bristol, 
with the description of some new 
genera and species, 414. 

——, on Anodontacanthus, a new 
genus of fossil fishes from the coal- 
measures, with descriptions of three 
new species, 427. 

Dawson, Dr. G. M., on the superficial 
geology of British Columbia and 
adjacent regions, 272. 

Dawson, Dr. J. W., award of the 
Lyell Medal to, Proc. 30. 

——, on new Hrian (Devonian) plants, 
299. 

Denbighshire grit, constitution of, 9. 

——, Dr. Hicks on some plant-re- 
mains discovered at the base of the, 
Corwen, N. Wales, 482. 

Dendroid Graptolites, Mr. C. Lap- 
worth on the, in the Llandovery 
rocks of Mid Wales, 171. 

Depolarization, areas of, from strain 
within perlitic bodies in obsidian 
tuff, 396. 

Devil’s Bridge, section from Aberyst- 
wyth to the, 148; fossils from, 
147; great inversion of rocks at, 
146. 

Devitrified rocks, Mr. F. Rutley on, 
from Beddgelert and Snowdon, 
403. 

Devonian grits and sandstones, 9. 

plants, Principal Dawson on new, 
299. 

Diamond-fields of South Africa, Mr. 
Dunn on the, 609. 

Diastopora, Mr. F. D. Longe on some 
specimens of, from the Wenlock 
limestone, 239. 

Diastopora cricopora, 387. 

oolitica, 386. 

stomatoporides, 384. 

ventricosa, 385. 

Diastoporide, Mr. G. R. Vine on the, 
from the Lias and Oolite, 381. 

Dicranophyllum australicum, 506. 

Detyonema corrugatellum, 172. 

delicatulum, 172. 

venustum, 171. 

Dictyophyton, 303. 

Dolomitie conglomerate near Clifton, 


Doratodon carcharidens, 671. 
Dorset, Mr. J. Buckman on some Am- 


719 


monites from the Inferior Oolite of 
Somerset and, 57. 

Dorset, Mr. S. S. Buckman on some 
Ammonites from the Inferior Oolite 
of, 588. 

Dovey valley, 162; section through, 
163. 


Drift-zones, Mr. D. Mackintosh on the 
existence of, 351. 

Drigg, interglacial deposit at, 34. 

Dulas Bay, 220. 

Duncan, Prof. P. Martin, presentation 
of the Wollaston Medal to, Proc. 28. 

, on the coralliferous series of 
Sind and its connexion with the 
last upheaval of the Himalayas, 
190. 

Dunn, EH. J., Esqg., on the diamond- 
fields of South Africa, 609. 

Durdham Down, conglomerate of, 67 ; 
section at the edge of, 70; Theco- 
dontosawrus-bed of, 73. 

Durness and Assynt, Dr. C. Callaway 
on the limestone of, 239. 

Durness area, map of the, 240. 

Dymoke, Keuper sandstone from, 15. 


Easton, section through, 296. 

Kecles, J., Esq., on the mode of occur- 
rence of some volcanic rocks of 
Montana, U.S. A., 399. 

Egton, constitution of moor grits 
from, 16. 

Hquisetides Wrightiana, 301. 

Emys Newmayrt, 698. 

Hppelsheim, 206. 

Hrian (Devonian) plants, Principal 
Dawson on new, 299. 

Erratic stones in the Moel-Tryfan de- 
posit, 353. 

Eruptive rocks of Skomer Island, Mr. 
F. Rutley on, 409. 

Etheridge, R., Hsq. (President), Ad- 
dress on presenting the Wollaston 
Medal to Prof. P. Martin Dunean, 
Proc. 28; Address on presenting 
the Murchison Medal to Prof. 
Archibald Geikie, 29; Address on 
handing the Lyell Medal to W. W. 
Smyth, Esq., for transmission to 
Dr. J. W. Dawson, 30; Address on 
handing the Bigsby Medal to Prof. 
Morris for transmission to Dr. 
Charles Barrois, 32; Address on 
handing the Wollaston Donation 
Fund to Prof. J. W. Judd for trans- 
mission to Dr. R. H. Traquair, 33 ; 
Address on presenting the balance 
of the Murchison Donation Fund to 
Frank Rutley, Esq., 34; Address 


720 


on presenting one moiety of the 
Lyell Geological Fund to G. R. 
Vine, Hsq., 35; Address on handing 
one moiety of the Lyell Geological 
Fund to Prof. H. G. Seeley for 
transmission to Dr. Anton Fritsch, 
36; Anniversary Address, February 
18, 1881. Obituary Notices of De- 
ceased Fellows:—Searles Valen- 
tine Wood, 37; John Jeremiah 
Bigsby, 39; Rev. J. Clifton Ward, 
qa) Prot Dy avAnsted, 14:31) Prot 
William Hallowes Miller, 44; Mr. 
H. Ludlam, 47; Mr. Robert Clut- 
terbuck, 48 ; Dr. Edward Meryon, 
48; Mr. Elijah Walton, 43 ; Dr. W. 
P. Schimper, 49; Dr. J. A. H. 
Bosquet, 50; and M. P. H. Nyst, 
50. Address on the Analysis and 
Distribution of the British Palzo- 
zoic Fossils, 51. 

Etheridge, R., Esq., on a new species 
of Trigonia from the Purbeck beds 
of the vale of Wardour, 246. 

——, Appendix to Dr. Hicks’s paper 
on some plant-remains in North 
Wales, 482. 


Fault at Porth-y-defaid, 
Porth-yr-Ogof, 226. 

Felin bont, halleflinta of, 217. 

Felsite schist from near Pont-y-Grom- 
lech, 405. 

Fenestella crassa, 186. 

-halkinensis, 187. 

—— membranacea, 181. 

nodulosa, 183. 

—— plebeia, 179. 

polyporata, 185. 

Fenestellide, Carboniferous, Mr. G. 
Shrubsole on the, 178. 

Wishes, fossil, Mr. J. W. Davis on 
Anodontacanthus, a new genus of, 
from the Coal-measures, 427. 

Fish-remains of the bone-bed at Aust, 
near Bristol, Mr. J. W. Davis on 
the, 414. 

Flagey gneiss of Durness, 239. 

Flintshire, high-level deposits on 
Halkin mountain, 357. 

Fonthill Giffard, Portlandian sand 
from, 16. 

Forests, so-called submerged, of West 
Cumberland and North Laneashire, 
oT. 

Fossils, Mr. J. W. Carrall on the 
locality of some, m the Carbonife- 
rous rocks at T’ang Shan, China. 
83. 

Fossils of the Middle Headon at Hea- 
don Hill and Colwell Bay, 104-106 ; 


224; at 


GENERAL INDEX. 


from the Brockenhurst zone at W hit- 
ley Ridge, New Forest, 111; of the 
Middle Headon series and Broc- 
kenhurst beds,. table of, 115; of 
Central Wales, table of, 170; of 
the Manchhar series, 204; of the 
Upper Greensand and Chloritic 
Marl of the Isle of Wight, 371- 
373. 

Fritsch, Dr. Anton, award of one 
moiety of the Lyell Geological 
Fund to, Proc. 36. ~ 

Frodsham, Keuper sands at, 15. 

Frondeg  gravel-pits, 360; shells 
found in, 360; boulders in, 360, 
361. 


Gaerwen, gneiss of, 212. 

Gaj river, section on the north bank 
of the, 194. 

series, 195; corals of the, 201. 

Sipser: River, quartz rhyolite from, 
396. 

Garthen valley, fossils from, 156. 

Geikie, Prof. A., presentation of the 
Murchison Medal to, Proc. 29. 

Geyser basin, Lower, spherulite rock 
from the, 395; vitreous tuff from 
the, 396. 

Glanrhyd, grey gneiss of, 219. 

Gneiss, grey, of the Llangefni area, 
218; of Glanrhyd, 219. 

Gneissic series of Anglesey, geogra- 
phical and stratigraphical distribu- 
tion of, 227, 228. 

of the Menai anticline, 211; 
of the Llangefni syncline, 216; 
of the central zone of Anglesey, 
219. 

Gosau beds of the Neue Welt, Prof. 
Suess on the, 702. 

formation, Prof. Seeley on the 
reptile fauna of the, 620. 

Granitoid gneiss, structure of, from 
Pen Bryn yr Hglwys, 235. 

Granitoidite of Llanerchymedd, 219. 

Graptolites, dendroid, Mr. ©. Lap- 
worth on the, in the Llandovery 
rocks of Mid Wales, 171. 

Gravel and sand, arrangement of, in 
the Moel-Tryfan deposit, 353 ; high- 
level, between Minera and Llan- 
gollen vale, Denbighshire, 359; 
high-level, near Llangollen, 363; 
of Macclesfield forest, 363; high- 
level, in Caernarvonshire, 307 ; in 
Ireland, 357 ; on Halkin Mountain, 
O07. 

Great Orton, section through, 296. 

Green-quarter Fell,constitution of grit 
from, 9. 


GENERAL INDEX, 


Greensand, Upper, and Chloritic 
Marl, Isle of Wight, Mr. Parkinson 
on, 370. 

Grey gneiss of Glanrhyd, 219; of the 
Llangefui area, 218. 

Grits, Mr. J. A. Phillips on the con- 
stitution and history of, 6. 

Gwalchmai, microscopic structure of 
quartz- -schists fr om, 232, 233. 

Gwastaden, section from Cwm Ystw yth 
to, 148; section from Strata Flor ‘ida 
and the Teifi pools to, 159. 

Gwrthya, 42.” 

Gwyn Llyn, section through, 145. 

Gypseous shales of Carlisle basin, 288, 
295. 


Hahn, Dr. Otto, remarks on micro- 
scopic sections of meteorites, Proc. 


7: 

Hala range, 192. 

Halkin Mountain, high-level deposits 
on, 357. 

Halleflinta of the Llangefni area, 217; 
near Llangwyllog church, 219. 

Hampshire basin, Prof. Judd on the 
remains of a Cetacean in the Lower 
Oligocene strata of the, 708. 

Haploceras oolithicum, 607. 

Harpoceras adicrum, 602. 

concayum, 603. 

cornu, 605. 

—— cycloides, 604. 

—— dispansum, 604. 

—— Hdouardianum, 604. 

fissilobatum, 603. 

Leyesquei, 604. 

— Moorei, 604. 

Murchisonez, 603. 

Sowerbyi, 602. 

Haute-Marne, Upper Jurassic rocks 
of the, 516. 

Headon beds, list of fossils from the, 
105. 

—— Hill, Mr. H. Keeping and Mr. 
EH. B. Tawney on the beds at, and 
at Colwell Bay, Isle of Wight, 85. 

, survey section east of, 86; 
vertical section at north-east corner 
of, 88. 

Hempstead series, sands from, 18. 

Heppenheim, 206. 

Hertfordshire puddingtone, 18. 

Hicks, Dr. H., on the discovery of 
some remains of plants at the base 
of the Denbighshire Grits near 
Corwen, North Wales, 482. 

High House, sandstone of, 15. 

Himalayas, Prof. P. M. Duncan on 
the connexion of the Coralliferous 


Q. J.G.S. No. 148. 


721 


series of Sind with the last HE 
heaval of the, 190. 

Himalayan upheaval, general conside- 
rations regarding the age of the 
last, 205. 

Hollingworth, G. H., Hsq., on a peat 
bed interstratified with the boulder- 
drift at Oldham, 713. 

Holmes, T. V., Esq., on the Permian, 
Triassic, and Liassic rocks of the 
Carlisle basin, 286. 

Holyhead district, 226. 

Mountain, quartz-schist from, 
226; structure of quartz-schist from, 
233. 

Holywell, Flintshire, 
Boulder-clay at, 19. 
Holwell, 67; Postpliocene, Liassic 

and Rheetic deposits of, 69. 

Hoplosaurus ischyrus, 681. 

Hordwell, constitution of sand from, 
18. 

Hornblende picrite, Prof. T. G. Bon- 
ney on a boulder of, near Pen-y- 
Carnisiog, Anglesey, 137. 

Hunstanton, constitution of Carstone 
of, 17; analysis of, 18. 

Hybodus austiensis, 416. 

punctatus, 417. 


sands from 


Igneous rocks of Anglesey, Prof. Bon- 
ney on the structure of some, 
236. 

Il-ga-chuz Mountain, British Colum- 
bia, 274. 

Inchnadamff church, 
242. 

, plan of limestone and quartzite 
at, 243. 

Inferior Oolite of Dorset, on some 
Ammonites from the, 57, 588. 

Interglacial deposits of West Cumber- 
land and North Lancashire, Mr. J. 
D. Kendall on, 29. 

Ireland, high-level gravel and sand 
in, 357. 

Tron Mountain, 
272, 274. 

Tron-ore, Mr. J. A. Phillips on the 
occurrence of remains of recent 
plants in brown, l. 

Tron-ores, pisolitic, 18. 

Isle of Wight, Mr. Parkinson on the 
Upper Greensand and Chloritic Marl 
of, 370. 


section above, 


British Columbia, 


Judd, Prof. J. W., on the occurrence 
of the remains of a Cetacean in the 
Lower Oligocene strata of the 
Hampshire basin, with a note by 
Prof. Seeley, 708. 

3c 


722 


Jurassic rocks, Upper, of England 
and the continent, Rev. J. KF. Blake 
on the, 497. 

sandstones, constitution of, 16. 


Kamloops, 
279. 

Keeping, H., Esq., and HE. B. Tawney, 
Esq., on the beds at Headon Hill and 
ee Bay in the Isle of Wight, 

Keeping, W., Esq., on the geology of 
Central Wales, with an appendix an 
some new species of Cladophora by 
C. Lapworth, Hsq., 141. 

Kendall, J. D., Esq., interglacial de- 
posits of West Cumberland and 
North Lancashire, 29. 

Beran: sandstones, constitution of, 


British Columbia, 278, 


Khirthar range, geology of the, 193; 
diagram section of the, 194. 

series, 195; corals of the, 199. 

Kimmeridgian of the Paris basin and 
England, 577. 

Kirklinton sandstone, 290, 295. 


La Charité, map of the Loire between 
Sancerre and, 538. 

Ladock, constitution of grits from, 10. 

Laignes, map of part of the valley of 
the. 528. 

Lake district, vertical section of Silu- 
rian rocks in the, 164. 

Laki range, geology of the, 193. 

Lampeter, fossils from, 155. 

Lancashire, Bunter sandstones of, 12. 

, Mr. J. D. Kendall on interglacial 

deposits of North, 29. 

, Mr. T. M. Reade on the date of 
the last change of level in, 436. 

Lapworth, Prof. C., on the Cladophora 
or dendroid Graptolites collected 
by Prof. Keeping in the Llandovery 
rocks of Mid Wales, 171. 

Large-flag series, 151. 

Lepidodendron primevum, 302. 

Lepralia cleidostoma, var. rotunda, 
306. 

corrugata, 330. 

—— monilifera, 335. 

spatulata, 335. 

Lias, Mr. Sollas on a new species of 
Plesiosaurus in the Lower, Char- 
mouth, 440. 

in the Carlisle basin, 293, 297. 

and oolite, Mr. G. R. Vine on 
Diastoporide from the, 381. 

Liassic remains, in alluvial veins, in 
the Carboniferous of Durdham 
Down, 73. 


GENERAL INDEX. 


Liassic rocks of the Carlisle basin, Mr. 
T. V. Holmes on the, 286. 

vein in the quarry, Durdham 
Down, 75. 

Limestone of Durness and Assynt, 
Dr. C. Callaway on the, 239. 

Limestones, crystalline, of Anglesey, 
Prof. Bonney on the structure of, 
235. 

Lindal, borings in Boulder-clay at, 
30, 37. 

Lisburne, fossils from, 156. 

Litherland, Keuper sandstone from, 
15. 

Little Orton, section through, 296. 

Lizard, Prof. Seeley on the remains of 
a small, from the Neocomian rocks 
of Comén, near Trieste, 52. 

Llanbrynmaer, 161; vertical section of 
Silurian rocks at, 164. 

Llandeilo, section from, to Aberaeron, 
108 ; vertical section of Silurian 
rocks west of, 164. 

Llandovery rocks of Mid Wales, Mr. 
C. Lapworth on the Cladophora of 
Ofte isle 

Llanerchymedd and Llangwyllog, sec- 
tion between, 219. 

Llanfadog uchaf, section through, 159. 

Llanfechell grits, 223; structure of 
slaty rock from, 254. 

Llangefni syncline, 213, conglomer- 
ates and shales, 214; conglomerate, 
structure of a fragment of grit i, 
235. 

Llangollen, high-level gravel and sand 
near, 363. 

Llangollen valley and Minera, high- 
level deposits between, 359. 

Llangrannog, constitution of grit from, 
8 


Llangristiolus grits and slates, 214. 

Llangwrig, section through, 159. 

Llangwyllog church, Halleflinta near, 
219. 

—-— and Llanerchymedd, section be- 
tween, 219. 

Lianwenllwyfo, 220. 

Lian y Mowddwy, vertical section 
across the Dovey valley at, 163 ; ver- 
tical section of Silurian rocks at, 164. 

Loire, map of the, between Sancerre 
and La Charité, 538. 

Longe, F. D., Esq., on some specimens 
of Diastopora and Stomatopora from 
the Wenlock limestone, Proc. 239. 

Lower Charente, Upper Jurassic rocks 
of the 543. 

Lower Geyser basin, spherulite rock 
from the, 395; vitreous tuff from 
the, 396. 


2 eae 


GENERAL INDEX. 


Lower Headon beds, 94; of the cliffs 
between Weston and Widdick 
Chines, 96; of Warden Cliff, 97 , 
of Colwell Bay, 101. 

Lower Lickey Hills, Upper Llando- 
very sandstone from, 9. 

Lower Mottled Sandstones, 12, 14. 

Lower Oligocene strata of the Hamp- 
shire basin, Prof. Judd on the re- 
mains of a Cetacean in the, 708. 

Lower Permian of the Carlisle basin, 
287. 

Lunulites cancellata, 344. 

guineensis, 344. 

Lyell Geological Fund, presentation 
of one moiety of, to G. R. Vine, 
Esq., Proc. 35 ; award of one moiety 
of the, to Dr. Anton Fritsch, Proc. 


36. 

Lyell Medal, award of the, to Dr. J. 
W. Dawson, Proc. 30. 

Lyssakine Hexactineliid, Prof. W. J. 
Sollas on a new, from the Silurian 
formation of Canada, 254. 

Lytoceras confusum, 601. 

—— Hudesianum, 601. 

-—— torulosum, 601. 


Macclesfield Forest, high-level gravel 
and sand of, 363. 

Machynlleth, fossils from Morben 
quarry, 153. 

Mackintosh, Mr. D., on the precise 
mode of accumulation and deriva- 
tion of the Moel-Tryfan shelly 
deposits and similar high-level de- 
posits along the eastern slopes of 
the Welsh mountains, and on the 
existence of drift-zones, showing 
probable variations in the rate of 
submergence, 351. 

Mammalian jaw, Mr. E. W. Wil- 
lett on a, from the Purbeck beds at 
Swanage, 376. 

Manchhar series of Sind, 196; equi- 
valence of the, and the Sivalik group 
of the Himalayas, 202. 

Map of country near Neuvizy, 502; 
round Verdun, 507 ; round St. Mi- 
hiel, 511; between the Meuse and 
Ornain, 514; of part of the valley 
of the Rognon, 518; of part of the 
Marne valley, 520; of the valley of 
the Aube, 526; of part of the val- 
ley of the Laignes, 528; of the val- 
ley of the Armangon, 551, of the 
Yonne valley south of Auxerre, 534 ; 
of the Loire valley between San- 
cerre and La Charité, 538; of 
country round Bourges, 540; of the 
coast round the Pertuis d’Antioch, 


723 


544; of the coast near Trouville, 
548; of part of the Pays de Bray, 
506; of part of the Boulonnais, 
559. 

Map of part of British Columbia, 278. 

eee part of the Yellowstone Park, 
400. 

Map of the coast at Blundellsands, 
near Liverpool, 436. 

Map of the Durness area, 240. 

Market Rasen, pisolitic iron-ore from, 
18. 

Marl, Chloritic, and Upper Green- 
sand, of the Isle of Wight, Mr. Par- 
kinson on, 370. 

Marne valley, map of part of the, 520. 

Maryport, interglacial deposit at, 36. 

May-hill sandstone, constitution of, 8. 

Megalosaurus pannoniensis, 670. 

Melin Carnau, 42. 

Melin pant y gwyda, structure of 
chlorite-schist from, 234. 

Membranipora argus, 324. 

—— catenularia, 323. 

concamerata, 324. 

— cylindriformis, 328. 

—— geminata, 325. 

lineata, 323. 

——. lusoria, 324. 

—— macrostoma, 323. 

maorica, 325. 

Menai anticline, 211. 

Mesocrinus suedicus, 130. 

Metalliferous slate group, 145, 152. 

Metalliferous slates, 164. 

Meteorites, Dr. Otto Hahn’s remarks 
on microscopic sections of, Proc. 7. 

Meuse, Upper Jurassic rocks of the, 505. 

Meuse and Ornain, map of country 
between the, 514. 

Micklethwaite, section through, 296. 

Microlestes quarry at Holwell, 69. 

Micropora patula, 326. 

Microporella enigmatica, 331. 

clavata, 332. 

—— coscinopora, 331. 

—— elevata, 330. 

—— ferrea, 330. 

—— symimetrica, 332. 

—- violacea, 329. 

yarraensis, 331. 

Microscopic characters of volcanic 
rocks from Montana, 398. 

Microscopic structure of some Angle- 
sey rocks, Prof. T. G. Bonney on 
the, 233. 

Middle Headon beds, 92; of Warden 
cliff, 100 ; of Colwell Bay, 101; of 
Whitechff Bay, 108. 

Millet-seed beds, 12, 26. 

Millstone-grit Sandstone, from Brig- 


724 


ham, constitution of, 11; analysis 
of, 21, 

Minera and Llangollen, high - level 
deposits between, 350. 

Mochlodon Suessti, 624. 

Moel Morfydd, section from Nant 
Llechos, near Corwen, to, 487. 

Moel Tryfan, shelly deposits of, 351 ; 
deposition of, during submergence, 
353; identification of local and 
erratic stones in, 353; arrangement 
of the gravel and sand in, 353; 
position of boulders in, 354; bent 
and shattered edges of slaty laminz 
in, 854, 368; origin of shells of, 
355; section of laminated sand and 
bent slates on, 355. 

Moel-Tryfan shelly deposits and simi- 
lar high-level deposits on the Welsh 
mountains, Mr. Mackintosh on the, 
Bol, 

Montana, U.S. A., Mr. F. Rutley on 
the vitreous rocks of, 391. 

, Mr. J. Eecles on some volcanic 

rocks of, 391. 

, microscopic characters of vol- 
eanic rocks from, 398. 

Moore, C., Esq., on abnormal geologi- 
cal deposits in the Bristol district, 
67. 

Morben quarry, Machynlleth, fruits 
from, 153. 

Mucronella duplicata, 328. 

elegans, 329. 

mucronata, 328. 

Murchison Fund, presentation of the 
Balance of, to Frank Rutley, Esq., 
Proc. 34. 

Murchison Medal, presentation of, 
to Prof. Archibald Geikie, Proc. 


29. 

Mynyda Liwydiarth, gneiss of, 212. 

Mynydd Mechell, chloritic schists of, 
222. 

Nant Cader, section through, 159. 

Nant Llechos, near Corwen, section 
from, to Moel Morfydd, 487. 

Nari series, 195; Corals of, 200. 

Nebo, 220. 

Nemacanthus minor, 419. 

Nematophycus Hicksti, 494. 

Neocomian rocks, Prof. Seeley on a 
small Lizard from the, of Comén, 
near Trieste, 52. 

Nettlebury Quarry, 78. 

Neue Welt, Prof. Suess on the Gosau 
beds of the, 702. 

Neuvizy, map of country near, 502. 

New Brunswick, plants from the De- 
yonian of St. John, 303. 


GENERAL INDEX. 


nee Forest, Brockenhurst zone in the, 

109. 

New York, plants from the Devonian 
of, 299, 303. 

Nicols River, British Columbia, 273. 

Niévre, Upper Jurassic rocks of, 537. 

Normandy, Upper Jurassic rocks of, 
DAT. 


Northern area of Anglesey, 221. 
North-western area of Anglesey, 224. 


Oborne, near Sherborne, vertical sec- 
tion of Inferior Odlite at, 589. 

Obsidian, devitrified, 405, 411. 

Obsidian from the Yellowstone dis- 
trict, 392; black, 391; felspar crys- 
tals in, 392; black spherulitic, 393 ; 
spherulitic-banded, 393 ; black por- 
phyritic, 394. 

Obsidian from Tolcsva, Hungary, 406. 

Obsidian tuff, areas of depolarization 
fromstrain in perlitic bodies in, 396. 

Odontocaulis Keepingii, 176. 

Odontopteris squamosa, 305. 

Okanagan valley, British Columbia, 
278, 279. 

Oldham, Mr. Hollingworth on a peat 
bed interstratified with the Boulder- 
drift at, '713. 

Oligosaurus adelus, 678. 

Oolite, Mr. J. Buckman on some Am- 
monites from the Inferior, of Dorset 
and Somerset, 57. 

, Inferior, of Dorset, Mr. S. S. 

Buckman on some Ammonites from 

the, 588. 

and Lias, Mr. G. R. Vine on 
Diastoporide from the, 381. 

Oolitic remains at Westbury-on-Trym, 


(de 

Oppelia subcostata, 606. 

—— subradiata, 606. 

Truellii, 606. 

Ormskirk, constitution of Sandstones 
from Town Green, near, 14. 

Orne and Sarthe, Upper Jurassic rocks 
of the, 547. 

Ornithocheirus Binzeli, 701. 

Ornithomerus gracilis, 671. 

Osborne Beds at Cliff End, 104. 

Owen, Prof., on the parts of the Ske- 
leton of an Anomodont Reptile 
(Platypodosaurus robustus, Owen), 
Part II. The Pelvis, 266. 

, on the Order Therodontia, with 
a description of a new genus and 
species (Ailurosaurus felinus, Ow.), 
261. 

Oxfordian of the Paris basin and 
England, 567. 

Pachytheca, 484, 492. 


GENERAL INDEX, 


Paleosaurus Stricklandi, 420. 

Paris Basin, Rev. J. F. Blake on the 
Upper Jurassic of the, +97. 

Paris Mountain, area south of, 220; 
Mr. Allport on rocks from, 220; 
volcanic group of, 221. 

Parkhead, sandstone from, 12. 

Parkinson, C., Esq., on Upper Green- 
sand and Chloritic Marl, Isle of 
Wight, 370. 

Patagonian archipelago, Soil-cap mo- 
tion in the, 348. 

Pays de Bray, Upper Jurassic rocks of 
the, 554; map of part of the, 556. 

Peace River, British Columbia, 276. 

Peat-bed interstratified with the 
Boulder -drift at Oldham, Mr. 
Hollingworth on a, 713. 

Pelvis, Prof. Owen on the, of an Ano- 
modont Reptile, 266. 

Pen bryn’r Eglwys, section from 
Porth-y-defaid to, 224; structure of 
granitoid gneiss from, 235. 

Pen Rhiev Wen, section through, 143. 

Penrith Sandstone, 287. 

Pentewan, shore-sand from, 19, 25. 

Pentraeth, gneiss south of, 211; 
structure of crystalline limestone 
from near, 235, 236. 

Pen-y-Carnisiog, Anglesey, Prof. T. G. 
Bonney on a boulder of hornblende 
picrite near, 137. 

Perisphinctes Martinsii, 601. 

pygmeeum, 601. 

Permian, Lower, of the Carlisle basin, 
287. 

Permian, Upper, of the Carlisle Basin, 
286. 


Permian rocks, of the Carlisle Basin, 
Mr. T. V. Holmes on the, 286. 

Permian Sandstones, constitution of, 
22: 

Perranzabuloe, constitution of grit 
from, 10, 

Pertuis d’Antioch, map of the coast 
round the, 544. 

Phillips, J. A., Esq., on the occurrence 
of remains of recent plants in 
brown iron-ore, 1. 

, on the constitution and his- 
tory of grits and sandstones, 6. 

Picrite, hornblende, Prof. T. G. 
Bonney on a boulder of, 137. 

Pikermi, 206. 

Pisolitic iron-ores, 18. 

Pian of limestone and quartzite at 
Inchnadamff, 245; of part of Rio 
Tinto, 3. 

Plants, Dr. Hicks on the remains of 
some, near Corwen, North Wales, 
482. 


725 


Plasgoch, section in quarry at, 44; 
analyses of rocks from, 45, note ; 
microscopic character of rock from, 
46. 

Plastremys lata, 370. 

Plas uchaf, 220. 

Platypodosaurus robustus, Prot. Owen 
on the pelvis of, 266. 

Plesiosaurus brachycephalus, 476. 

conybeari, 440. 

megacephalus, 440. 

Pleuropeltus Suessti, 693. 

Plynlimmon, vertical section of Silu- 
rian rocks at, 164. 

grits, 162. 

group, 156. 

Point Atlianus, structure of slaty rock 
from, 235. 

Pont Cletwr Yspytty, constitution of 
erit from, 9. 

Erwyd, fossils from near, 152; 
section from, through the Plyn- 
limmon axis to Llangwrig and 
Rhyader, 159. 

Pont ur Elan, 
148. 

Pont-y-Gromlech, felsite schist from 
near, 405. 

Popty, section through, 143. 

Porella denticulata, 336. 

emendata, 336. 

Porina clypeata, 332. 

columnata, 334. 

coronata, 332. 

Porthdinlleyn, Prof. Bonney on the 
so-called serpentine of (Caernarvon- 
shire), 40. 

Porth feiin, structure of 
schist from, 234. 

Porth lygan, 220. 

Porth Nobla, coast-section from, to 
Aberffraw sands, 216. 

Porth-y-defaid fault, 224; section 
from, to Pen bryn’r Eglwys, 224; 
structure of chlorite-schist from, 
234. 

Porth-yr-Ogof fault, 226; structure 
of chlorite-schist from, 234. 

Portlandian of the Paris basin and 
England, 585. 

Post-Tertiary sands, constitution of, 
18). 

Psaronius textilis, 307. 

Purbeck beds, Mr. R. Etheridge on a 
new species of Zrzgonia from the, 
of the Vale of Wardour, 246. 

Purbeck beds at Swanage, Mr. E. W. 
Willett on a mammalian jaw from 


the, 376. 


section through, 


chlorite- 


Quartzite and sandstone fossiliferous 


726 GENERAL INDEX. 


pebbles in the drift in Warwick- 
shire, Rey. P. B. Brodie on certain, 
430. 

Quartzite of Durness, 242. 

Quartz rhyolite, from Yellowstone 
Canon, 395; from Gardiner’s River, 
396. 

Quartz-schist from Holyhead Moun- 
tain, 226; of the Llangefni area, 
PAT 

of Anglesey, Prof. Bonney on 
the characters of, 232. 

Queen-Charlotte Islands, glaciation of 
the, 280. 


Ranikot series, 193; corals of, 197. 

Reade, T. M., Esq., on the date of the 
last change of level in Lancashire, 
436. 

Recent plants in brown iron-ore, Mr. 
J. A. Phillips on the occurrence of 
remains of, |. 

Reptile fauna of the Gosau formation, 
Prof. Seeley on the, 620. 

Reptilia, age of the Bristol, 80. 

Retepora marsupiata, 342. 

rimata, 3438. 

Rhadinosaurus alcinus, 675. 

Rheetic bone-bed near Clifton, 75. 

Rheetic remains in alluvial veins in 
the Carboniferous of Durdham 
Down, 73. 

Rheda, Cumberland, constitution of 
St.-Bees sandstone of, 12. 

Rhodes Bank, Oldham, section of 
boulder-clay with interstratified 
peat near, 713. 

Rhosbeirio shales, 225. 

Rhoscolyn, serpentine from near, 40, 


42. 


church, quartz-schist from, 233. 

district, contortiou of rocks in, 
227. 

Rhyader, section through, 143; sec- 
tion from Pont Erwyd through the 
Plynlimmon axis to Llangwrig and, 
159; vertical section of Silurian 
rocks at, 164. 

Rhyolite, quartz, 
River, 396; 
Canon, 395. 

from Clogwyn @ur Arddu, 405. 

Rio Tinto, remains of recent plants in 
brown iron-ore at, 1. 

, plan and section of part of, 3. 

, analysis of iron-ore from, 4. 

Rocks, Anglesey, Prot. 'T. G. Bonney 
on some, 232. 

, devitrified, Mr. F. Rutley on, 

from Beddgelert and Snowdon, 

405. 


from Gardiner’s 
from Yellowstone 


Rocks, eruptive, Mr. F. Rutley on the, 
of Skomer Island, 409. 

, Serpentine and associated, Prof. 

Bonney on the, of Anglesey, 40. 

, vitreous, Mr. F. Rutley on the, 

of Montana, U.S. A., 391. 

, volcanic, Mr. Hecles on some, 
of Montana, U.S. A., 399. 

Rognon, map of part of the valley of 
the, 518. 

Roydon brickyard, section in, 113, 
note. 

zone, 113. 

Runcorn station, constitution of sand- 
stone from, 15. 

Rutley, Frank, Esq., presentation of 
the balance of the Murchison Fund 
to; Proes 3a. 

, on the microscopic characters 

of the vitreous rocks of Mon- 

tana, U.S. A., with an appendix by 

Mr. J. Eccles, 391. 

, on the microscopic structure of 

devitrified rocks from Beddgelert 

and Snowdon, with an appendix on 
the eruptive rocks of Skomer Island, 

403. 


Beare constitution of grit from, 

0. 

St. Austell, constitution of grits from 
near, 9, 25. 

river, sands of, 21. 

St. Bees, interglacial deposits at, 35. 

sandstone, 286; constitution of, 


12. 

St. Ewe, constitution of grits from, 9. 

St. John, New Brunswick, plants from 
the Devonian of, 303. 

St. Mihiel, map of country round, 
511. 

Sancerre, map of the Loire between 
La Charité and, 538. 

Sands, water-borne, examination of, 
21. 

Sandstone and quartzite fossiliferous 
pebbles in the drift inWarwickshire, 
Rev. P. B. Brodie on, 430. 

Sandstones, Mr. J. A. Phillips on the 
constitution and history of, 6. 

Sango Bay, gneiss of, 241. 

Sarthe and Orne, Upper Jurassic rocks 
of the, 547. 

Schist, dark, of the Llangefni area, 
218; of Glanrhyd, 219. 

Schists of Holyhead island, micro- 
scopic characters of, 47. 

Schizoporella amphora, 341. 

australis, 341. 

conservata, 340. 

— excubans, 341. 


GENERAL INDEX. aL 


Schizoporella fenestrata, 339. 

phymatopora, 338. 

spiroporina, 340. 

—— submersa, 340. 

—— ventricosa, 338. 

—— vigilans, 338. 

sp., 339. ; 

Scotland, Devonian plant from, 306. 

, South, vertical section of Silu- 
rian rocks in, | 64. 

Sections :—of part of Rio Tinto, 3; of 
junction of schist and serpentine, 
41; of quarry at Ceryg Moelion, 
43; of quarry at Plas Goch, 44; 
at the edge of Durdham Down, 70: 
of Thecodontosaurus-bed of Durdham 
Down, 73; vertical, at north-east 
corner of Headon Hill, 91; vertical, 
of Lower Headon (freshwater) beds 
exposed between Weston Chine and 
Warden Cliff, 98; vertical, of Up- 
per Headon beds in Colwell Bay, 
103; from Aberystwith to the Devil’s 
Bridge, 143; from Cwm Ystwyth, 
through Craig Lluest, to Rhyader 
and Gwastaden, 143; in railway- 
cutting between Traws Coed and 
Caradoc waterfall, 155; from Pont 
Erwyd through the Plynlimmon 
axis to Llangwrig and Rhyader, 
159; from Strata Florida and the 
Teifi pools to Rhyader and Gwas- 
taden, 159; showing general struc- 
ture of Central Wales, 160; through 
the Dovey valley at Llan y Mowd- 
dwy, 163; vertical, of Silurian 
rocks in the English lake-district, 
South Scotland, South-west Cardi- 
ganshire, at Rhyader, west of Llan- 
deilo, at Aberystwyth and Plynlim- 
mon, Llanbrynmaer, and Llan y 
Mowddwy, and in the Clwydd val- 
ley, 164; through Barrah Hill, 194; 
of the Khirthar range, on the north 
bank of the Gaj river, 194; above 
Inchnadamff church, 242; of Pur- 
beck beds in a railway-cutting west of 
Dinton Station, Vale of Wardour, 
252; from the Solway, north of 
Allonby, to near Brampton, 296; 
from Solway Moss to Dalston, 296 ; 
from Tordoff Point, Dumfriesshire, 
to the fault south of Wigton, 296; 
of laminated sand and bent slates 
on Moel Tryfan, 355; of drift-de- 
posits between Braich and the sum- 
mit of the ridge, Frondeg, Denbigh- 
shire, 367; of Upper Greensand 
and chloritic marl, St. Lawrence 
and Ventnor, Isle of Wight, 374; 
at Blundellsands, near Liverpool, 


437; from Nant Llechos, near Cor- 
wen, to Moel Morfydd, 487; of 
boulder-clay with  interstratified 
peat, near Rhodes Bank, Oldham, 
718. 

Seeley, Prof. H. G., on remains of a 
small lizard from the Neocomian 
rocks of Comén, near Trieste, pre- 
served in the Geological Museum of 
the University of Vienna, 52. 

—, on the reptile fauna of the 
Gosau formation, preserved in the 
Geological Museum of Vienna, 620. 

, on the caudal vertebra of a Ce- 
tacean, discovered by Prof. Judd in 
the Brockenhurst beds, indicative of 
a new type allied to Balenoptera 
(Balenoptera Juddi) 709. 

Selenaria alata, 344. 

marginata, 344. 

Serpentine, Prof. Bonney on the so- 
called, of Porthdinlleyn (Caernar- 
vonshire), 40. 

, Prof. Bonney on the, of An- 
glesey, 40. 

Sevenoaks stone, constitution of, 16. 
Shalk Beck, Yoredale grit from, con- 
stitution of, 11; analysis of, 21. 
Shells, of Moel-Tryfan deposit, origin 

of the, 355. 

Sherborne, vertical section of Inferior 
Oolite at Oborne, near, 589. 

Shrubsole, Mr. W., on the Carboni- 
ferous Fenestellide, 178. 

Silurian grits and sandstones, 7, 25. 

formation, Mr. W. J. Sollas on 

Astrocoma Granti, from the, of 

Canada, 254. 

uniserial Stomatoporee and Asco- 
dictya, Mr. Vine on, 613. 

Sind, Prof. P. M. Duncan on the co- 
ralliferous series of, 190. 

, Western, Table of formations in, 
192. 

Sivalik group, equivalence of the 
Manchhar series of Sind with the, 
202. 

Skomer Island, Mr. F. Rutley on the 
eruptive rocks of, 409. 

Slaty laminz, bent and shattered edges 
of, at Moel Tryfan, 354, 368. 

rocks of Anglesey, Prof. Bonney 

on the structure of, 234. 

series of Anglesey, geographical 

distribution of, 225. 

of the central zone of 

Anglesey, 218. 

of the Menai anticline, 213; 
of the Llangefni syncline, 214. 

Smith, Dr. William, presentation of 
portrait of, Proc. 2. 


—— 


728 


Smittia anceps, 337. 

centralis, 337. 

, var. levigata, 337. 

Tatei, 337. 

Snowdon and Beddgelert, Mr. F. 
Rutley on devitrified rocks from, 
403. 

Soilcap-motion, Mr. R. W. Coppinger 
on, 348. 

Sollas, Prof. W. J., on Astroconia 
Granti, a new Lyssakine Hexacti- 
nellid from the Silurian formation 
of Canada, 254. 

—, on a new species of Plesiosaurus 
(P. Conybearz) from the Lower Lias 
of Charmouth, with observations on 
P. megacephalus, Stutchbury, and 
P. brachycephalus, Owen, 440. 

Solway, section from the, to near 
Brampton, 296. 

Solway Moss, section from, to Dalston, 
295. 

Somerset, Mr. J. Buckman, on some 
Ammonites from the Interior Oolite 
of Dorset and, 57. 

South Thompson valley, British Co- 
luinbia, 2795. 

Spheroceras Brongniarti, 596. 

dimorphum, 9598. 

—— Gervillii, 597. 

—— Mantellii, 597. 

—— polymorphum, 598. 

Wrightii, 598. 

Sphenonchus obtusus, 420. 


nt 


Spherulitic rock, devitrified, from. 


Beddgelert, 403. 

from the Lower Geyser 

basin, 395. 

from the Yellowstone dis- 
trict, 394. ; 

Spinkwell quarry, constitution of 
Coal-measure sandstone from, 11 ; 
analysis of, 21. 

Spirophyton, 303. 

Stanwix, section through, 296. 

Stanwix marls, 292, 295. 

Stedd fa Gurig, section through, 159. 

Stephanoceras Braikenridgii, 595. 

Deslongchampsi, 594. 

—_— Humphriesianum, 594. 

linguiferum, 595. 

—— polymerum, 596. 

zigzag, 596. 

Stiper stone, constitution OI, Uf, UD: 

Stomatopora, Mr. F. D. Longe on 
some specimens of, from the Wen- 
lock limestone, Proc. 239. 

Stomatopora dissimilis, 615. 

inflata, 615. 

Stomatopore and Ascodictya, My. 
G. R. Vine on, 615. 


GENERAL INDEX, 


Stonyhough, constitution of coal- 
measure sandstone from, L1. 

Stow-on-the-Wold, constitution of 
Liassic sand from, 16. 

Strata Florida rock, 154. 

Struthiosaurus austriacus, 628, 

Submerged forests, so-called, of West 
Cumberland and North Lancashire, 
oT. 

Submergence, deposition of Moel-Try- 
fan shelly beds during, 353. 

——, Mr. D. Mackintosh on drift- 
zones and probable variations in the 
rate of, 357. 

, rate of, in Denbighshire, 362. 

Suess, Prof. E., on the Gosau beds of 
the Neue Welt, west of Wiener 
Neustadt, 702. 


_ Superficial geology of British Colum- 


bia and adjacent regions, Dr. G. 
Dawson on, 272. 

Swanage, Mr. H. W. Willett on a 
mammalian jaw from the Purbeck 
beds at, 376. 

Sweden, Mr. P. H. Carpenter on two 
new Crinoids from the Upper Chalk 
of Southern, 128. 


T’ang Shan, Mr. Carrall on some fossils 
found in carboniferous rocks at, 83. 

Tan y graig, Pentraeth, structure of 
crystalline limestone from, 236. 

Tau isa, Cemmaes, structure of crys- 
talline limestone from, 236. 

Tawney, H. B., Esq., and H. Keeping, 
Esq., on the beds at Headon Hill and 
Colwell Bay in the Isle of Wight, 
85. 

Teifi pools, section from, to Gwasta- 
den, 159, 

Terraces in British Columbia, 274. 

Tertiary sandstones and grits, consti- 
tution of, 18. 

Thames valley, sands of the, 19. 

Thecodontosaurus-bed of Durdham 
Down, 73. 

Theriodontia, Prof. Owen on Aluro- 
saurus felinus, a new species and 
genus of, 261. 

Thornbury railway, Secondary veins 
on, (9: 

Thornby, section through, 296. 

Tilgate sandstone, constitution of, 16. 

Tolesya, Hungary, deep red obsidian 
from, 406. 

Tonnerre, map of the valley of the 
Armancon, north of, 531. 


- Tordoff point, section from, to the 


fault south of Wigton, 296. 
Town Green, near Ormskirk, consti- 
tution of sandstone from, 14. 


GENERAL INDEX. 


Trachyte, quartz-oligoclase, from Sko- 
mer Island, 411. 

Traquair, Dr. R. H., award of the 
Wollaston Donation Fund to, Proc. 


33- 

Traws Coed and Caradoc waterfall, 
section in railway-cutting between, 
155. 

Triassic rocks of the Carlisle basin, 
Mr. T. V. Holmes on the, 286. 

Triassic sandstones, constitution of, 
12, 25, 26. 

Triconodon mordax, 378. 

Trieste, Prof. Seeley on the remains of 
a small Lizard from the Neocomian 
rocks of Comén, near, 52. 

Trigonia, Mr. R. Etheridge on a new 
species of, 246. 

— densinoda, 246. 

Trouville, map of the coast near, 
548, 

Tubutella ambigua, 71, 80. 

Tuff, vitreous, from the Lower Geyser 
basin, Yellowstone district, 396. 

Ty Croes, section on the railway from, 
to Bodorgan, 217 ; structure of rock 
from, 233. 

Tyddyn Gob, 41, 47. 

Ty Newydd, 40; microscopic character 
of rock from, 46; analysis of ser- 
pentine from, 46, ote. 

Ty Ucha, 41; microscopic characters 
of rock from, 45. 


Upper Greensand, Mr. Parkinson on 
the, and chloritic marl, Isle of 
Wight, 370. 

Upper Headon beds, 92; of Colwell 
bay, 102. 

Upper Jurassic rocks of England, Rev. 
J. HF. Blake on the correlation of the, 
with those of the continent, 497. 

Upper Mottled Sandstones, 12, 14. 

Upper Permian of the Carlisle basin, 
286. 


Veins, alluvial, with Liassic and Rhx- 
tic remains, in the carboniferous of 
Durdham Down, 73. 

Verdun, map of the country round, 
HUM 

Vertebra, caudal, of a Cetacean, Prof. 
Seeley on the, 709. 

Victoria, Mr. A. W. Waters on fossil 
Chilostomatous  Bryozoa from 
South-west, 309. 

View of perched gravel-mound on 
Halkin Mountain, 358. 

Vine, G. R., Esq., presentation of one 
moiety of the Lyell Geological Fund 
to, Proc. 35. 

Q.J.G.8. No. 148. 


729 


Vine, G. R., Hsq., on the Family Dias- 
toporide, Busk:—species from the 
Lias and Oolite, 381. 

, on Silurian uniserial Stomato- 
pore and Ascodictya, 613. 

Vitreous rocks, Mr. F. Rutley on, of 
Montana, U.S. A., 391. 

Voleanie group of Paris Mountain, 


= 


rocks from Montana, microscopic 
characters of, 398. 

of Montana, U.S. A., Mr. 
Eccles on some, 399. 


Wales, Central, diagram of the general 
structure of, 160. 

; , Mr. W. Keeping on the 

geology of, 141. 

, North, Dr. Hicks on some 
remains of plants in the Denbigh- 
shire grits at Corwen, 482. 

Walney, interglacial deposit at, 34. 
Warden cliff, Lower Headon beds of, 
97; Middle Headon beds of, 100. 
and Weston chine, verti- 
cal section of Lower Headon (fresh- 

water) beds exposed between, 98. 

Wardour, Mr. R. Etheridge on a new 
species of Trigonia from the Pur- 
beck beds of the vale of, 246. 

Warwickshire, Rev. P. B. Brodie on 
quartzite and fossiliferous pebbles 
in the drift in, 480. 

W ater-borne sands, examination of, 21. 

Waters, A. W., Esq., on fossil Chilo- 
stomatous Bryozoa from South-west 
Victoria, 309. 

Waterstone beds, constitution of, 15. 

Welsh mountains, Mr. Mackintosh on 
Moel-Tryfan shelly deposits and 
similar high-level deposits on the, 
ool. 

Wenlock limestone, Mr. F. D. Longe 
on some specimens of Dzastopora 
and Stomatopora from the, Proc. 


239. 

Westbury-on-Trym, carboniferous 
limestone with minerals and oolitic 
remains, 77. 

Western area of Anglesey, 225. 

Weston chine, Lower Headon beds of 
the cliff between, and Widdick 
chine, 96. 

and Warden cliff, vertical 
section of Lower Headon (fresh- 
water) beds exposed between, 98. 

Whidborne, G. F., Hsq., on the genus 
Plesiosaurus, 440. 

Whitecliff bay, Middle Headon of, 
108; Brockenhurst zone at, 109. 
Widdick chine, Lower Headon beds of 

3D 


730 


the cliffs between, and Weston chine, 
96 


Wigton, section from Solway Moss to 
the fault south of, 296. 

Willett, EK. W., Esq., on a mammalian 
jaw from the Purbeck beds at 
Swanage; with a note by H. Wil- 
lett, Esq., 376. 

Wollaston Donation Fund, award of 
the, to Dr. R. H. Traquair, Proc. 33. 

Wollaston Medal, presentation of, to 
Prof. P. Martin Dunean, Proc. 28. 

Wyke quarry, vertical section of In- 
ferior Oolite at, 589. 


Yate rock, 78. 


GENERAL INDEX. 


Yeathouse, sandstone from, 12. 

Yellowstone cafion, quartz rhyolite 
from, 395. 

district, vitreous tuff from the 

Lower Geyser basin, 396; spheru- 

litic rock from, 394; obsidian from, 

391-594. 

park, map of part of, 400. 

Y Foel, section through, 159. 

Ynyslas, 42. 

Yonne, Upper Jurassic rocks of the, 


valley, map of the, south of 
Auxerre, 534. 

Yoredale grit, from Shalk Beck, con- 
stitution of 11; analysis of, 21. 


END OF VOL, XXXVII. 


Printed by TAYLOR and Francis, Red Lion Court; Fleet Street. 


PROCEEDINGS 


OF THE 


GEOLOGICAL SOCIETY OF LONDON. 


SESSION 1880-81. 


November 3, 1880. 


Rosert Erneriper, Esq., F.R.S., President, in the Chair. 


Bernard Barham Woodward, Esq., 19 Stowe Road, Shepherd’s 
Bush, W., was elected a Fellow of the Society. 


The List of Donations to the Library was read. 


The following specimens were presented to the Museum :— 


Twenty specimens of Carboniferous Limestone Fossils from Flat- 
head River, Rocky Mountains (49th Parallel N.W. America), pre- 
sented by H. Bauerman, Esq., F.G.S. 


The President announced that the original portrait of Dr. William 
Smith, painted by M. Fourau in the year 1837, had been presented 
to the Society by William Smith, Ksq., of Cheltenham. 


The following communications were read :— 


1. “On the Serpentine and Associated Rocks of Anglesey, with a 
Note on theso-called Serpentine of Porthdinlleyn (Caernarvonshire).”’ 
By Prof. T. G. Bonney, M.A., F.R.S., Sec. G.S. 


2. “ Note on the Occurrence of Remains of Recent Plants in 
Brown Iron-ore.” By J. Arthur Phillips, Esq., F.G.S. 


3. “ Notes on the Locality of some Fossils found in the Carboni- 
ferous Rocks at T’ang Shan, situated, in a N.N.E. direction, about 
120 miles from Tientsin, in the province of Chih Li, China.” By 

VOL, XXXVII. a 


2 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


James W. Carrall, Esq., F,G.S. With a Note by Wm. Carruthers, 
Esq., F.R.S., F.G.S. 


The following specimens were exhibited :— 


Rock-sections and specimens of rocks from Anglesey and Porth- 

dinlleyn, exhibited by Prof. T. G. Bonney in illustration of his 
aper. 

y ate of recent plants in brown iron-ore, exhibited by J. 
Arthur Phillips, Esq., in illustration of his paper. 

Carboniferous plant-remains, from T’ang Shan, Chih Li, China, 
exhibited by J. W. Carrall, Hsq., in illustration of his paper. 

Sections of Devonian Corals, Agates, and Mocha Stones, exhibited 
by S. H. Needham, Esq., to illustrate a new method of mounting 
large sections of fossils, and minerals, and other natural-history 
objects (such as Lepidoptera), so as to exhibit both the upper and 
under surfaces at one view. 


November 17, 1880. 
Rosert ErHeripesr, Hsq., F.R.S., President, in the Chair. 


Prof. Joseph Henry Tompson, of the Auckland College, Auckland, 
New Zealand, was elected a Fellow of the Society. ‘ 


The names of the following Fellows in arrear to the Society were 
read out by the President for the first time, in accordance with 
Section VI. B, Article 6, of the Bye-laws:—W. H. Peacock, Esq., 
and W. Salmon, Hsq. 


The List of Donations to the Library was read. 


The Presipent called attention to the portrait of Dr. William 
Smith, presented to the Society by his grand-nephew, Mr. W. Smith, 
of Cheltenham, which was then suspended behind the chair, and 
expressed his great satisfaction at this most interesting picture being 
in the possession of the Society. 

Mr. W. W. Smyrta expressed the satisfaction that all must feel 
in possessing a genuine relic of this eminent stratigraphical geologist. 
Now this one, which had been so liberally presented to the Society, 
was a most indubitable portrait of the most conspicuous founder of 
English geology. That portrait was painted by M. Fourauin 1837, 
and was certainly an admirable likeness. The Society was deeply 
indebted to the donor, Mr. W. Smith, the cousin of the late valued 
Prof. Phillips. The portrait now hanging on the wall was engraved 


in Prof. Phillips’s ‘ Life’ of his uncle. He proposed a hearty vote 
of thanks to the donor. 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 3 


Mr. Evans rose with great pleasure to second the vote of thanks 
proposed by Mr. Warington Smyth. The portrait was indeed 
replete with interest, not only to English geologists but to all 
geologists in the world. An additional interest attaching to the 
portrait was that we had the whole history of it from Dr. Smith’s 
own hand, an extract from which Mr. Evans read. The portrait 
was an admirable one. He hoped that in the future Mr. Smith’s 
example would be followed, and that we should see many other por- 
traits of eminent geologists on the Society’s walls. The Society 
was also deeply indebted to the President for the interest which he 
had taken in the matter. 

The vote of thanks was carried by acclamation. 


The following communications were read :— 


1. *¢ On abnormal Geological Deposits in the Bristol District.” 
By Charles Moore, Esq., F'.G.S. 


2. “ Interglacial Deposits of West Cumberland and North Lanca- 
shire.” By J. D. Kendall, Esq., C.E., F.G.S. 


Specimens from deposits in the Bristol District were exhibited by 
Mr. Moore in illustration of his paper. 


December 1, 1880. 
Rosert Erueriper, Esq., F.R.S., President, in the Chair. 


William Heward Bell, Esq., Hast Shefford House, Hungerford ; 
William Jackson, Esq., Vernon Terrace Schools, Northampton ; 
Peregrine Propert Lewes, Esq., M.A., L.L.M., 84 Kensington Gar- 
dens Square, W.; William Libbey, Hsq., Jun., M.A., D.Sc., Princeton 
College, Princeton, New Jersey, U.S.A.; David Morgan Llewellin, 
Esq., Bryn Gomer, near Pontypool, Monmouthshire; John Mar- 
shall, Esq., Sowerby Bridge, near Halifax; Cyril Parkinson, Esq., 
Rock Cottage, Ventnor, Isle of Wight, and Farnsfield, Southwell, 
Notts; Cornelius McLeod Percy, Esq., The Grove, Standish, Wigan ; 
Thomas John Robinson, Esqg., Melbourne Street, Longton, Stoke- 
on-T'rent; Rey. Alfred Rose, M.A., Emmanuel College, Cambridge ; 
Beeby Thompson, Esq., F.C.8., Abington Street, Northampton; and 
Stuart Crawford Wardell, Esq., Doe Hill House, Alfreton, Derby- 
shire, were elected Fellows of the Society. 


The names of the following Fellows in arrear to the Society were 
read out by the President for the second time, in aceordance with. 


4 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Section VI. B, Article 6, of the Bye-laws:—W. H. Peacock, Esq., 
and W. Salmon, Esq. 


The List of Donations to the Library was read. 


The following communications were read :— 


1. “On Remains of a small Lizard from the Neocomian Rocks of 
Comén, near Trieste, preserved in the Geological Museum of the 
University of Vienna.” By Prof. H. G. Seeley, F.R.S., F.G.S. 


2. “On the Beds at Headon Hill and Colwell Bay in the Isle of 
Wight.” By H. Keeping, Esq., and HK. B. Tawney, Esq., M.A., 
F.G.S. 


Specimens were exhibited by Prof. Seeley and Messrs. Tawney and 
Keeping in illustration of their papers. 


December 15, 1880. 
Roser Erueriper, Esq., F.R.S., President, in the Chair. 


William Elijah Benton, Esq., Assoc. R.S.M., Highfield House,. 
Heather, near Ashby-de-la-Zouch ; Rev. George Clements, 26 St. 
Martin’s Road, Stockwell; J. Kerr Gulland, Esq., C.E., 6 a Victoria 
Street, S.W., and Ball’s Pond Road, N.; Francis T. 8. Houghton, 
Ksq., B.A., Wynnstay, Balsall Heath, Birmingham ; George Bingley 
Luke, Esq., Northbank House, Prestonpans, N.B.; and William 
Mansell MacCulloch, Esq., MD., 12 Cork Street, Burlington Gardens, 
W., were elected Fellows; and Professor Luigi Bellardi, of Turin, 
and Dr. M. Neumayr, of Vienna, Foreign Correspondents of the 
Society. 


The List of Donations to the Library was read. 


The following communications were read :— 


1. “On the Constitution and History of Grits and Sandstones.” 
By John Arthur Phillips, Esq., F.G.S. 


The Chair was then taken by J. W. Hutxz, Esq., F.RS., 
V.P.G.S. 


2. “ On a New Species of T'rrgonza from the Purbeck Beds of the 
Vale of Wardour.” By R. Etheridge, Esq., F.R.S., President. With 


a Note on the Stratigraphical position of the Fossil by the Rev. W. 
R. Andrews. 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 5 


The following specimens were exhibited :— 

Specimens exhibited by Messrs. J. Arthur Phillips and R. Ethe- 
ridge in illustration of their papers. 

Specimens of Crinoids from the Carboniferous Limestone, and a 
pair of candlesticks turned out of Carboniferous Limestone, exhibited 
by Prof. James Tennant, F.G.S. 


January 5, 1881. 


Rozert ErHeriner, Esq., F.R.S., President, in the Chair. 


George C. Crick, Esq., 9 Gwyn Street, Bedford; and Arthur S$. 
Reid, Esq., B.A., 12 Bridge Street, Canterbury, were elected Fellows 
of the Society. 


The List of Donations to the Library was read. 
The following communications were read :— 


1. “The Archean Geology of Anglesey.” By C. Callaway, Esq., 
M.A., D.Sc., F.G.S. With a Note on the Microscopic Structure of 
some of the Rocks, by Prof. T. G. Bonney, M.A., F.R.S., Sec.G.5. 


2. “The Limestone of Durness and Assynt.” By C. Callaway, 
Esq., D.Sc., F.G.8. 


3. “Ona Boulder of Hornblende-Picrite near Pen-y-Carnisiog, 
Anglesey. By Prof. T. G. Bonney, M.A., F.R.S., Sec.G.8. 


The following specimens were exhibited :— 


Specimens of the results of some experiments in the formation 
of Agates &c. exhibited by EH. A. Pankhurst, Esq., and James 
TAnson, Esq., F.G.S. 

Rock-specimens and microscopic sections exhibited by Dr. Calla- 
way and Prof. Bonney, in illustration of their papers. 


January 19, 1881. 
Rosrrt Ernzriper, Esq., F.R.S., President in the Chair. 
Jabez Church, Esq., M.Inst.C.E., 178 Great George Street, West- 


minster, 8.W.; George Augustus Freeman, Esq., B.Sc. Lond., 6 
VOL. XXXVII. b 


6 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Macduff Terrace, Danby Street, Peckham, 8.H.; Charles Horsley, 
Ksq., C.H., 174 Highbury New Park, N.; Edwin Simpson-Baikie, 
Ksq., F.L.S., United University Club, Pall Mall Hast, S.W.; and 
Charles John Wood, Esq., M.Inst.C.E., 2 Selbourne Terrace, Bradford, 
Yorkshire, were elected Fellows of the Society. . 


The List of Donations to the Library was read. 


The President called attention to the valuable donation made to 
the Library by Dr. J. J. Bigsby, F.B.S., F.G.8., of the set of six 
volumes of the ‘ Reports of the Geological Survey of Illinois,’ the 
last four volumes of which are extremely rare on this side of the 
Atlantic, 1f not almost unique. A special vote of thanks was passed 
to Dr. Bigsby for his present. 


The following communications were read :— 


1. ‘Further Notes on the Family Diastuporide, Busk.” By G. R. 
Vine, Esq. Communicated by Prof. P. Martin Duncan, M.B.Lond., 
PSs, EGS. 


2. ‘Further Notes on the Carboniferous Fenestellide.” By G. 
W. Shrubsole, Esq., F.G.S. 


The following specimens were exhibited :— 
Specimens of Opal, Chalcedony, &c., from Australia, exhibited by 
Prof. James Tennant, F.G.S. 


Specimens of Fenestellide, exhibited by Mr. Shrubsole in illus- 
tration of his paper. 


ie ce 


February 2, 1881. 
Roserr Ernertper, Ksq., F.R.S., President, in the Chair. 


Joseph Groves, Esq., B.A., M.B. Lond., Carisbrooke, Isle of Wight ; 
George Lewis, Esq., Green Hill, Derby; Rev. Edouard Méchin, 8.J., 
St. Beuno’s College, St. Asaph, North Wales ; James Osborne, Esq., 
Rio Tinto, Province of Huelva, Spain; and the Rey. William Shar- 
inan, 20 Headland, Plymouth, were elected Fellows of the Society. 


The List of Donations to the Library was read. 
The following communications were read :— 


1. “ On the Coralliferous Series of Sind, and its connexion with 
the last Upheaval of the Himalayas.” By Prof. P. Martin Duncan, 
M.B.Lond., F.R.8., F.G.S. 


2. “On two new Crinoids from the Upper Chalk of Southern 
Sweden.” By P. H. Carpenter, Esq., M.A. Communicated by 
Prof. P. Martin Duncan, M.B.Lond., F.R.S., F.G:S. 


PROCEEDINGS OF THE GEOLOGICAL SOCIDTY. 7 


The following specimens were exhibited :— 


A specimen of a new Crinoid, exhibited by Mr. Carpenter in 
illustration of his paper. 

Microscopic sections of Meteorites, exhibited by Dr. Otto Hahn, 
of Reutlingen. 

In explanation of the latter the following remarks were addressed 
on behalf of Dr. Hahn to the President and Fellows present :— 

“Dr. Hahn, in inviting you to examine the microscopical speci- 
mens of meteorites which he has prepared, and in order to assist 
you in determining the character of the forms and structures which 
you will find exhibited in them, desires to present a short summary 
of the negative considerations which forbid that such structures 
should be classed among crystalline forms. 

* As is well known, the chondrites, the species of meteorites 
from which his specimens are prepared, consist, besides the metals 
which they enclose, of the minerals enstatite and olivine. 

“‘ Tn his work on the meteorites and their organisms, lately pub- 
lished, Dr. Hahn has given photographs of 130 different forms and 
structures. Now if these structures are crystalline, the two 
minerals in question would present themselves in at least 130 dif- 
ferent forms and structures, although the absence of all structure is 
recognized as a fundamental principle of the theory of minerals. 

“ Again, the structures exhibited by the chondrites cannot be 
due to slaty cleavage, since olivine has no slaty cleavage, and that of 
enstatite and of uther minerals does not appear under the micro- 
scope, or else presents itself there under totally different forms. 

“The greatest importance, however, is to be attached to the total 
absence of all polarized light exhibited by the two minerals as oc- 
curring in the meteorites. The contained forms and structures do 
not polarize the light at all, or only very feebly, although the same 
minerals, under ordinary circumstances, polarize light very strongly. 
The absence of all aggregate polarization is especially noticeable, as 
proving that these objects are not aggregates of crystals. 

‘“¢ Should we still feel inclined to regard the enclosures as mineral 
forms, and not as organisms, we must be struck by the utter absence 
of all crystalline forms, especially in those very minerals which 
always, and occasionally also in meteorites, appear in a crystallized 
form. 

“ Further, the external forms, and consequently the outlines of the 
enclosures, harmonize so perfectly with their internal form and 
structure, that we cannot entertain the idea that these enclosures 
had been rolled about and ground down before they became finally 
imbedded in the chondrites. 

“The idea of an aggregate of crystals, if still looked upon with 
favour, would be contradicted by the fact that the enclosed balls 
or globes are all constructed excentrically, whereas all terrestrial 
erystallites are formed concentrically.” 


62 


8 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


ANNUAL GENERAL MEETING, 
February 18, 1881. 


Rozert Ernrriper, Esq., F.R.S., President, in the Chair. 


REPORT OF THE CouNnciIL FoR 1880. 


In presenting their Report for the year 1880, the Council of the 
Geological Society have much gratification in announcing to the 
Fellows that the affairs of the Society are in a much more satis- 
factory position than at the dates of their last two Reports. 

The number of new Fellows elected during the year is 60, of 
whom 49 paid their fees before the end of the year, making, ah 
14 previously elected Fellows who paid their fees in 1880, a total 
accession during the year of 63 Fellows. Against this we have to 
record the loss by death of 34 Fellows, and by resignation of only 
9 Fellows, whilst 5 Fellows were removed from the list for non- 
payment of Contributions, making a total loss of 48 Fellows. On 
the year, therefore, we have an increase of 15 Fellows. But as of 
the 34 Fellows deceased, 9 were compounders, and 9 con-contri-. 
buting Fellows, the number of contributing Fellows is actually 
increased by 25, being now 769. 

The total number of Fellows and Foreign Members and Corre- 
spondents was 1415 at the end of the year 1879, and 1432 at the 
end of the year 1880. 

During the year 1880 intelligence was received of the death of 
1 Foreign Member and of 2 Foreign Correspondents. As two 
vacancies existed in the list of Foreign Members at the end of the 
year 1879, three Foreign Members were elected in 1880; and the 
vacancies caused by these elections and by death in the list of 
Foreign Correspondents, were all filled up during the year. 

The total Receipts for the year 1880 were £2706 1s. 7d., being 
£130 15s. 7d. more than the estimated Income for the year. The 
total Expenditure, on the other hand, was £2520 18s. 9d., or 
£154 4s. 1d. less than the estimate for the year. The actual excess 
of Income over Expenditure was thus £185 2s. 10d.; and of this 
amount a sum of £98 10s. was invested in the purchase of £100 
Consols. 

The Council have to announce the completion of Vol. XXXVI. of 
the Quarterly Journal and the commencement of Vol. XX XVII. 

The Council have further to announce the completion of the 


a a a 


ANNUAL REPORT. 9 


printing of the Catalogue of the Library, which is now ready for 
distribution to subscribers. It forms a volume of over 620 pages, or 
about 100 pages more than was originally estimated ; but the Council 
have not thought it desirable to increase the price of the volume. 
It will accordingly be furnished to the Fellows at the price of five 
shillings. 

The Council have much pleasure in announcing that William 
Smith, Esq., of Cheltenham, the grandnephew of Dr. ‘William Smith, 
and cousin of the late Professor Phillips, has most liberally presented 
to the Society the portrait of Dr. Smith which was formerly in Prof. 
Phillips’s possession, and from which was engraved the small portrait 
illustrating his ‘Memoir of William Smith.’ This picture, which is 
admirable in itself, is of special interest to the Fellows of the 
Geological Society, not only as an excellent portrait of one of the 
great founders of their Science, but also from its historical associa- 
tions, as its origin and mode of production have been described in 
Dr. Smith’s own words in the ‘ Memoir’ just mentioned, as it bears 
his handwriting on its back, and as it is the original of the engraved 
portrait. For the due preservation of this valuable picture the 
Council have ordered it to be suitably framed and covered with plate 
glass, and to be suspended in a place of honour in the Meeting-room 
of the Society. They hope that it may not long remain there in 
solitude. In the mean-time four of the marble busts in the possession 
of the Society (those, namely, of Greenough, Sedgwick, Murchison, 
and Macculloch) have been arranged upon pedestals at the end of the 
Meeting-room, where they may serve to cherish the memory of some 
of those who have contributed the most to advance the Science of 
Geology. 

The Council have awarded the Wollaston Medal to Professor P. 
Martin Duncan, F.R.S., F.G.8., in testimony of their high apprecia- 
tion of his numerous and valuable paleontological memoirs, especially 
on the Fossil Corals, as well as of his contributions to physical and 
stratigraphical Geology. 

The Murchison Medal, with the sum of Ten Guineas from the 
proceeds of the Fund, has been awarded to Professor Archibald 
Geikie, F.R.S., F.G.S., in recognition of his valuable contributions to 
physical and stratigraphical Geology, particularly in relation to the 
Old Red Sandstone of Scotland, and the Voleanic Geology of the 
vicinity of Edinburgh and the Firth of Forth. 

The Lyell Medal, with a sum of Twenty-five Pounds from the 
proceeds of the Fund, has been awarded to Principal Dawson, 
F.R.S., F.G.S., in recognition of his important contributions to the 
Geology of Canada, and more especially of his investigations into the 
Fossil Flora of the Devonian and Carboniferous Periods in America. 

The Bigsby Medal has been awarded to Dr. Charles Barrois, of 
Lille, as a testimony to the importance of his contributions to strati- 
sraphical Geology, particularly with relation to the Cretaceous 
system. 

"The balance of the proceeds of the Wollaston Donation Fund 
has been awarded to Dr. Ramsay H. Traquair, F.G.S., in recogni- 


igo) PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


tion of the value of his researches upon the paleontology of Fossil 
Fishes, and to aid him in pursuing his investigations. 

The balance of the proceeds of the Murchison Donation Fund 
has been awarded to Frank Rutley, Esq., F.G.8., as a token of 
appreciation of his valuable memoirs on various petrological sub- 
jects, and to assist him in carrying on his researches. 

The balance of the proceeds of the Lyell Donation Fund has 
been awarded in equal parts to G. R. Vine, Esq., in recognition of 
the work done by him on the Carboniferous Polyzoa, and to assist 
him in continuing his investigations ; and to Dr. Anton Fritsch, of 
Prague, as a mark of the estimation in which his valuable contri- 
butions to Palzontology are held, and to aid him in the production 
of his important memoirs. 

During the last Session, a subscription was set on foot among the 
Fellows of the Society, which enabled the Assistant Secretary to 
purchase four small plain Microscopes, for use, when required, at 
the Evening Meetings of the Society. As the procuring the means 
of illumination for the microscopes was attended with considerable 
inconvenience and expense, the Council have decided to expend a 
small portion of the proceeds of the Barlow-Jameson Fund in the 
purchase of four of How’s Microscope Lamps. 


REPORT OF THE LIBRARY AND Museum Committee, 
Lnbrary. 


Since the last Anniversary Meeting a great number of valuable | 
additions have been made to the Library, both by donation and by 
purchase. é 

As Donations the Library has received 105 volumes of separately 
published works and Survey Reports, and about 220 Pamphlets and 
separate impressions of Memoirs; also about 102 volumes and 205 
detached parts of the publications of various Societies, and 14 
volumes of independent Periodicals presented chiefly by their re- 
spective Lditors, besides 11 volumes of Newspapers of various 
kinds. This will constitute a total addition to the Society’s Library, 
by donation, of about 272 volumes and 220 pamphlets. 

A considerable number of Maps, Plans, and Sections have been 
added to the Society’s collections by presentation from various Geo- 
logical Surveys, from the Ordnance Survey of Great Britain, and from 
the French Depot de la Marine. Several Geological Maps of parti- 
cular districts have also been presented by the authors. They 
amount altogether to 348 sheets, and among them may be noted, as 
especially valuable to the Society, 131 sheets of the Map of the 
Geological Survey of Great Britain and Ireland. 

The Books and Maps just referred to have been received from 


ANNUAL REPORT, Mt 


128 personal Donors, the Editors or Publishers of 15 Periodicals, 
and 131 Societies, Surveys, or other Public Bodies, making in all 
274 Donors, 

By Purchase, on the recommendation of the Standing Library 
Committee, the Library has received the addition of 46 volumes of 
Books, and of 53 parts (making about 10 volumes) of Periodicals, 
besides 12 parts of works published serially, the earlier portions of 
which were obtained in previous years. Five Sheets of the Geolo- 
gical Survey Map of France, and 4 sheets of a Map of the Bohe- 
mian Coalfield, by Dr. Wolff, have been obtained by purchase. 

The cost of Books and Periodicals during the year 1880 was 
£73 2s. 3d., and of Binding £76 10s. 1d. The total expenditure 
on account of the Library was thus £149 12s, 4d. 

The Books in the Society’s Library are generally in good con- 
dition; and a considerable number of old serial and other works, 
of which the binding had been damaged by long use, have been 
rebound or repaired. The Library continues to be much used by 
the Fellows of the Society. : 


Museum. 


The Collections in the Museum remain in much the same con- 
dition as at the date of the last Report of the Committee, the 
Foreign Collections being all available for reference. The Egerton 
Collection of Arctic Fossil Shells has been named and arranged by 
Dr. J. Gwyn Jeffreys. 

The following Donatisns have been made fo the Museum during 
the year 1880 ;—A specimen of rock from the top of the Asnai 
Heights, Cabul, presented by Lieut. F. Spratt, R.K., and 20 speci- 
mens of Carboniferous-limestone Fossils from Flathead River, Rocky 
Mountains, presented by H. Bauerman, Iisq., ¥.G.5. 


12 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


CoMPARATIVE STATEMENT OF THE NUMBER OF THE SOCIETY AT THE 
CLOSE OF THE YEARS 1879 anp 1880. - 


Dee. 31, 1879. Dec. 31, 1880. 

Compoundersyee es ae DOs eee: 312 
Contributing Fellows... .. TA” ibe ae. 769 
Non-contributing Fellows . . 21s POON E 268 

1334 1349 
Honorary Members ...... Siny de en ee 3 
Foreign Members ........ BOs he Mien 40 
Foreign Correspondents... . AO seis cody te tone tie 40 

1415 1432 


General Statement explanatory of the Alterations in the Number of 


Fellows, Honorary Members, &c. at the close of the years 1879 and 
1880. 


Number of Compounders, Contributing and Non- 1384 
contributing Fellows, December 31, 1879.... } 

Add Fellows elected during former year and paid 
: 14 
in 1880 

Add Fellows elected and paid in 1880 ....... : 49 


@:er7\0) ‘©) (a (e (e) © “0. ce) 1@ @ © (ee ie » © [ele « « (0 (e¢ je « 0) wy lene 


1397 
Deduct Compounders deceased .............. 


Contributing Fellows deceased ........ 
Non-contributing Fellows deceased .... 
Contributing Fellows resigned 
Contributing Fellows removed 


eecee ee eee 


lacoae 


1349 


Members, and Foreign Correspondents, 
December 31, 1879 
Deduct Foreign Member deceased .. ..... 1 
Foreign Correspondents deceased .. 2 
Foreign Correspondents ee | 


Number of Honorary Members, nets 


Foreign Members 


~I 


Add Foreign Members elected .......... 3 
Foreign Correspondents elected 


| 
| 


ANNUAL REPORT. 


Decrasep Frtiows. 


Compounders (9). 


Lord Belper. ) W. Gillespie, Esq. 

T. Bell, Esq. | W. H. Holloway, Esq. 
Col. W. G. Boyle. | EK. Walton, Esq. 

Sir R. Burdett. S. V. Wood, Esq. 


W. W. Collins, Esq. 


Resident and other Contributing Fellows (16). 


Prot. D. T. Ansted. D. Llewellin, Esq. 

T. D. Bott, Esq. _ iH. Ludlam, Esq. 
Lieut.-Gen. W. EK. D. Broughton. | Dr. E. Merypn. 

E. W. Cooke, Esq. E. J. Smith, Esq. 

J. Cope, Esq. T. Parry, Esq. 

J. G. H. Godfrey, Esq. W. W. Stoddart, Esq. 
J. Hamilton, Esq. : Rev. J. Clifton Ward. 
W. G. Kell, Esq. E. B. Webb, Esq. 


Non-contributing Fellows (9). 


C. L. Bradley, Esq. | J. R. Logan, Esq. 
R. Davey, Esq. Rey. 8. Lucas. 

T. C. Eyton, Esq. | Prof. W. H. Miller. 
W. Gray, Esq. | Dr. Jones Quain. 


Very Rey. H. P. Hamilton. 


Foreign Member. 
Profyb ws Niyst 


Foreign Correspondents. 


Prof. W. P. Schimper. | M. J, A. H. Bosquet. 
Fellows Resigned (9). 

Rey. W. H. Allen. _ Lieut.-Col. C. Manby. 

F. Campion, Esq. _ Lieut.-Col. E. B. Sladen. 

S. N. Carvalho, Esq., Jun. _ H. B. Whitehead, Esq. 

E. C. H. Day, Esq. | Capt. H. T, Wing. 


H, Fox, Esq. 


14 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Fellows Removed (5). 


Jonathan Harrison, Esq. W. Harrison Peacock, Esq. 
J. Trubshaw Johnson, Esq. William Salmon, Esq. 
Lewis Thomas Lewis, Esq. 


The following Personages were elected from the List of Foreign Cor- 
respondents to fill the vacancies in the List of Foreign Members 
during the year 1889. 


Professor Gustave Dewalque of Liége. 
Professor Adolf Eric Nordenskiold of Stockholm. 
Professor Ferdinand Zirkel of Leipzig. 


The following Personages were elected Foreign Correspondents during 
the year 1880, 


Dr. Ferdinand von Hochstetter of Vienna. 
Professor Leo Lesquereux of Columbus. 
M. Alphonse Renard of Brussels. 


After the Reports had been read, it was resolved :— 


That they be received and entered on the Minutes of the Meeting, 
and that such parts of them as the Council shall think fit be printed 
and distributed among the Fellows, 


It was afterwards resolvea :— 


That the thanks of the Society be given to Sir P. de M. Grey- 
Egerton, Bart., and Prof. A. C. Ramsay, retiring from the office of 
Vice-President. , 


That the thanks of the Society be given to Dr. H. Hicks, Prof. 
T. M‘Kenny Hughes, Prof. T. Rupert Jones, Prof. J. Prestwich, and 
Prof. A. C. Ramsay, retiring from the Council. 


After the Balloting-glasses had been duly closed, and the lists 
examined by the Scrutineers, the following gentlemen were declared 
to have been duly elected as the Officers and Council for the ensuing 
year :— 


ANNUAL REPORT, 15 


OFFICERS, 


PRESIDENT. 
R. Etheridge, Esq., F.R.S. 


VICE-PRESIDENTS. 


J. Evans, D.C.L., LL.D., F.B.S. 
J. W. Hulke, Esq., F.R.S. 
Prof. J. Morris, M.A. 

Tals Ob sont oni, dllb Dy Ine iigse 


SHCRETARIES. 


Prof. T. G. Bonney, M.A., F.R.S, 
rote Wierd udde aks. 


4 


FOREIGN SECRETARY. 
W. W. Smyth, Esq., M.A., F,R.S. 


TREASURER. 
J. Gwyn Jeffreys, LL.D., F.R.S. 


COUNCIL. 


H. Bauerman, Esq. W. H. Hudleston, Esq., M.A. 
Rev. J. F. Blake, M.A. J. W. Hulke, Esq., F.R.S. 
Prof. T. G. Bonney, M.A., F.R.S. |J. Gwyn Jeffreys, LL.D., F.R.S. 
W. Carruthers, Esq., F.R.S. Prof. J. W. Judd, F.R.S. 


Prof. P. M. Dune 


an, M.B.,F.R.S. | Prof. N. 8. Maskelyne, M.P., 


Sir P. de M. Grey-Kgerton, Bart., M.A., F.R.S. 


M.P., F.RS. 


Prof. Morris, M.A. 


R. Etheridge, Esq., F.R.S. J. A. Phillips, Esq. 
J. Evans, D.C.L., LL.D., F.R.S. | F. W. Rudler, Esq. 
Lieut.-Colonel H. H. Godwin- | Prof. H. G. Seeley, F.R.8. 


Austen, F.R.S. 
J. C. Hawkshaw, 


W. W. Smyth, Esq., M.A., F.R.S, 
Esq., M.A. H. C. Sorby, LL.D., F.R.S. 


Rey. Edwin Hill, M.A, ; |H. Woodward, LL.D., F.R.S, 


16 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


LIST OF 


THE FOREIGN MEMBERS 


OF THE GEOLOGICAL SOCIETY OF LONDON, tn 1880. 


Date of 
Election. 


1827. Dr. H. von Dechen, Bonn. 

1829. Dr. Ami Boué, Vienna. 

1844, William Burton Rogers, Esq., Boston, U. S. 
1848. James Hall, Esq., Albany, State of New York. 
1850. Professor Bernhard Studer, Berne. 

1851. Professor James D. Dana, New Haven, Connecticut. 
1853. Count Alexander von Keyserling, Raykiill, Russia. 
1855. Professor L.G. de Koninck, ee 

1854. M. Joachim Barrande, Prague. 

1856. Professor Robert Bunsen, For. Mem. R.S., Heidelberg. 
1857. Professor H. R. Goeppert, Breslau. 

1857. Professor H. B, Geinitz, Dresden. 

1857. Dy. Hermann Abich, Vienna. 

1859. Professor A. Delesse, Paris. 

1859. Dr. Ferdinand Roemer, Bresiav. 

1860. Dr. H. Milne-Edwards, For. Mem. R.S., Paris. 
1862. Professor Pierre Merian, Basle. 

1864. M. Jules Desnoyers, Paris. 

1866. Dr. Joseph Leidy, Philadelphia. 

1867. Professor A. Daubrée, Paris. 

1870. Professor Oswald Heer, Zurich. 

1871. Dr. 8. Nilsson, Lund. 

1871. Dr. Henri Nyst, Brussels. (Deceased. ) 

1871. Dy. Franz Ritter von Hauer, Vienna. 

1874. Professor Alphonse Favre, Geneva. 

1874. Professor E. Hébert, Paris. 

1874. Professor Edouard Desor, Neuchdtel. 

1874. Professor Albert Gaudry, Paris. 

1875. Professor Fridolin Sandberger, Wiirzburg. 

1875. Professor Theodor Kjerulf, Christianta. 

1875. Professor F. August Quenstedt, Tiibingen. 

1876. Professor E. Beyrich, Berlin. 

1877. Dr. Carl Wilhelm Giimbel, Munich. 

1877. Dr. Eduard Suess, Vienna. 

1879, Dr. F. V. Hayden, Washington. 

1879. Major-General N. von Kokscharow, St. Petersburg. 
1879. M. Jules Marcou, Salus. 

1879. Dr. J. J. S. Steenstrup, For. Mem. R.8., Copenhagen. 
1880. Professor Gustave Dewalque, Liége. 

1880. Professor Adolf Eric Nordenskidld, Stockholm. 
1880. Professor Ferdinand Zirkel, Lezpzzg. 


ANNUAL REPORT. 


LIST OF 


THE FOREIGN CORRESPONDENTS 
OF THE GEOLOGICAL SOCIETY OF LONDON, ry 1880. 


Date of 
Election. 


1863. Dr. G. F. Jager, Stuttgart. 

1863. M.S. Lovén, Stockholm. 

1863. Count A. G. Marschall, Vrenna. 

1863. Professor G. Meneghini, Pisa. 

1863. Signor Giuseppe Ponzi, Rome. 

1865. Signor Quintino Sella, Rome. 

1865. Dr. F. Senft, Evsenach. 

1864. M. J. Bosquet, Maestricht. (Deceased.) 
1864. Dr. Charles Martins, Montpellier. 

1866. Professor J. P. Lesley, Philadelphia. 

1866. Professor Victor Raulin, Bordeaux. 

1866. Baron Achille de Zigno, Padua. 

1870. Professor Joseph Szabd, Pesth. 

1870, Professor Otto Torell, Lund. 

1871. M. Henri Coquand, Marseilles. 

1871. Professor Giovanni Capellini, Bologna. 
1872. Herr Dionys Stur, Vienna. 

1872. Professor J. D. Whitney, Cambridge, U. S. 
1874, Professor Igino Cocchi, Florence. 

1874. M. Gustave H. Cotteau, Aaverre. 

1874. Professor W. P. Schimper, Strasburg. (Deceased. 
1874. Professor G. Seguenza, Messina. 

1874. Dr. J. S. Newberry, New York. 

1874. Dr. T. C. Winkler, Haarlem. 

1875. Professor Gustav Tschermak, Vienna. 
1876. Professor Jules Gosselet, Lille. 

1876. Professor Ludwig Riitimeyer, Basle. 

1877. Professor George J. Brush, New Haven. 
1877. Professor A. L. O. Des Cloizeaux, For. Mem.R.S., Paris. 
1877. Professor KE. Renevier, Lausanne. 

1877. Count Gaston de Saporta, Adx-en-Provence. 
1879. Professor Pierre J. van Beneden, For.Mem.R.8., Louvain. 
1879. M. Edouard Dupont, Brussels. 

1879. Professor Guglielmo Guiscardi, Naples. 
1879. Professor Franz Ritter von Kobell, Munich. 
1879. Professor Gerhard vom Rath, Bonn. 

1879. Dr. Emile Sauvage, Paris. 

1880. Dr. Ferdinand von Hochstetter, Vienna. 
1880. Professor Leo Lesquereux, Columbus. 

1880, M. Alphonse Renard, Brussels. 


18 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


AWARDS OF THE WOLLASTON MEDAL 


UNDER THE CONDITIONS OF THE ‘‘ DONATION FUND” 
ESTABLISHED BY 


WILLIAM HYDE WOLLASTON, M_D., F-.BS., F.G.S., &e. 


“To promote researches concerning the mineral structure of the earth, 
and to enable the Council of the Geological Society to reward those 
individuals of any country by whom such researches may hereafter be 
made,”—“ such individual not being a Member of the Council.” 


1831. Mr. William Smith. 1959, | Herr Hermann von Meyer. 
1835. Dr. G. A. Mantell. ' (Mr. James Hall. 
1836. M. L. Agassiz. 1859. Mr. Charles Darwin. 
1837, ae T. P. Cautley. 1860. Mr. Searles V. Wood. 

Dr. H. Falconer. 1861. Professor Dr. H. G. Bronn. 
1838. Professor R. Owen. 1862. Mr. R. A. C. Godwin- 
1839. Professor C. G. Ehrenberg. Austen. 


1840. Professor A. H. Dumont. 1863. Professor Gustav Bischof. 
1841. M. Adolphe T. Brongniart. 1864. Sir R. I. Murchison. 


1842. Baron L. von Buch. 1865. Mr. Thomas Davidson. 
1843. {M. Elie de Beaumont. 1866. Sir Charles Lyell. 

\M.P. A. Dufrénoy. 1867. Mr. G. Poulett Scrope. 
1844. The Rev. W. D. Conybeare. | 1868. Professor Carl F. Naumann. 
1845. Professor John Phillips. 1869. Dr. H. C. Sorby. 
1846. Mr. William Lonsdale. 1870. Professor G. P. Deshayes. 
1847. Dr. Ami Boué. 1871. Professor A. C. Ramsay. 


1848. The Rey. Dr. W. Buckland. | 1872. Professor J. D. Dana. 
1849. Professor Joseph Prestwich. | 1873. Sir P. de M. Grey-Kgerton. 


1850. Mr. William Hopkins. 1874, Professor Oswald Heer. 
1851. The Rev. Prof. A. Sedgwick. | 1875. Professor L. G. de Koninck. 
1852. Dr. W. H. Fitton. 1876. Professor T. H. Huxley. 
1853. M. le Vicomte A. d’Archiac. | 1877. Mr. Robert Mallet. 

M. E. de Verneuil. 1878. Dr. Thomas Wright. 
1854. Sir Richard Griffith. 1879. Professor Bernhard Studer. 
1855. Sir H. T. De la Beche. 1880. Professor Auguste Daubrée. 


1856. Sir W. HE. Logan. 1881. Professor P. Martin Duncan. 
1857. M. Joachim Barrande. re 


ANNUAL REPORT. 


AWARDS 


OF THE 


Ng 


BALANCE OF THE PROCEEDS OF THE WOLLASTON 
“ DONATION-FUND.” 


1831. 
18383. 
1854. 
18368. 
1836. 
1838. 
1839. 
1840. 
1841. 
1842. 
1848. 
1844. 
1845. 
1846. 
1847. 


1848. 


1849. 
1850. 
1851. 
1852. 
1853. 
1854, 
1855. 
1856. 
1857. 


Mr. William Smith. 

Mr. William Lonsdale. 
M. Louis Agassiz. 

Dr. G. A. Mantell. 
Professor G. P. Deshayes. 
Professor Richard Owen. 
Professor G. C. Ehrenberg. 
Mr, J. De Carle Sowerby. 
Professor Edward Forbes. 
Professor John Morris. 
Professor John Morris. 
Mr. William Lonsdale. 
Mr. Geddes Bain. 

Mr. William Lonsdale. 
M. Alcide d’Orbigny. 


} Cape-of-Good-Hope Fossils. 


M. Alcide d’Orbigny. 
Mr. William Lonsdale. 
Professor John Morris. 
M. Joachim Barrande. 
Professor John Morris. 


Mr. 8. P. Woodward. 
Drs. G. and F’. Sandberger. 
Professor G. P, Deshayes. 
Mr. 8S. P. Woodward. 


| 1868. 
1859. 


1860. 


1861. 
| 1862. 
_ 1868. 
| 1864. 
— 1865. 
1866. 
| dushovi, 
| 1868. 
| 1869. 
1870. 
1871. 
S72: 
| 1878. 
— :1874. 
| 1875. 
elisiG: 
| Men 
1878. 
| 1879. 
1880, 
1881. 


Professor L. G. de Koninck. | 


| 


Mr. James Hall. 

Mr. Charles Peach. 
Professor T. Rupert Jones. 

sae W. K. Parker. 

Professor A. Daubrée. 

Professor Oswald Heer. 

Professor Ferdinand Senft. 

Professor G. P. Deshayes. 

Mr. J. W. Salter. 

Dr. Henry Woodward. 

Mr. W. H. Baily. 

M. J. Bosquet. 

Mr. W. Carruthers. 

M. Marie Rouault. 

Mr. R. Etheridge. 

Mr. James Croll. 

Professor J. W. Judd. 

Dr. Henri Nyst. 

Mr. L. C. Miall. 

Professor Giuseppe Seguenza. 

Mr. R. Etheridge, Jun. 

Mr. W. J. Sollas. 

Mr. 8. Allport. 

Mr. Thomas Davies. 

Dr. R. H. Traquair. 


AWARDS OF THE MURCHISON MEDAL 


AND OCF THE 


ROCEEDS OF “THE MURCHISON GEOLOGICAL FUND,” 
ESTABLISHED UNDER THE WILL OF THE LATE 
SIR RODERICK IMPEY MURCHISON, Banr., F.R.S., F.GS. 


“To be applied in every consecutive year in such manner as the Council 
of the Society may deem most useful in advancing geological science, 


20 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


whether by granting sums of money to travellers in pursuit of know- 
ledge, to authors of memoirs, or to persons actually employed in any 
inquiries bearing upon the science of geology, or in rewarding any 
such travellers, authors, or other persons, and the Medal to be given 
to some person to whom such Council shall grant any sum of money 
or recompense in respect of geological science.” 


1873. My. William Davies. Medal. | 1877. Rev. W. B. Clarke. Medal. 


1873. Professor Oswald Heer. 1877. Rev. J. F. Blake. 

1874. Dr. J. J. Bigsby. Medal. 1878. Dr. H. B. Geinitz. Medal. 
1874. My. Alfred Bell. 1878. My. C. Lapworth. 

1874. Mr. Ralph Tate. 1879. Professor F. M‘Coy. Medal. 
1875. Mr. W. J. Henwood. Medal. | 1879. My. J. W. Kirkby. 

1875. Prof. H. G. Seeley. 1880. Mr. R. Etheridge. Medal. 
1876. Mr. A.R.C. Selwyn. Medal. | 1881. Professor A.Geikie. Medal. 
1876. Mr. James Croll. 1881. Mr. F. Rutley. 


AWARDS OF THE LYELL MEDAL 
AND OF THE 


PROCEEDS OF THE “LYELL GEOLOGICAL FUND,” 


ESTABLISHED UNDER THE WILL AND CODICIL OF THE LATE 


SIR CHARLES LYELL, Barrt., F.RS., F.G.S. 


The Medal ‘‘to be given annually” (or from time to time) “as a mark of 


honorary distinction as an expression on the part of the governing 
body of the Society that the Medallist has deserved well of the 
Science,” —‘‘not less than one third of the annual interest [of the 
fund] to accompany the Medal, the remaining interest to be given in 
one or more portions at the discretion of the Council for the encou- 
ragement of Geology or of any of the allied sciences by which they 
shall consider Geology to have been most materially advanced.” 


1876. Professor John Morris. | 1879. Professor H. A. Nicholson. 


Medal. 1879. Dr. Henry Woodward. 
1877. Dr. James Hector. Medal. 1880. Mr. John Evans. Medal. 
1877. Mr. W. Pengelly. 1880. Professor F. Quenstedt. 
1878. Mr. G. Busk. Medal. 188]. Professor J. W. Dawson. 
1878. Dr. W. Waagen. Medal. 


1879. Professor Edmond Hébert. | 1881. Dr. Anton Fritsch. 


Medal. 1881. Mr. G. R. Vine. 


nt 


ANNUAL REPORT. 21 


AWARDS OF THE BIGSBY MEDAL, 
FOUNDED BY 


Dee I biG SBN B.S. E. Gas: 


To be awarded biennially “as an acknowledgment of eminent services 
in any department of Geology, irrespective of the receiver’s country ; 
but he must not be older than 45 years at his last birthday, thus 
probably not too old for further work, and not too young to have done 
much.” 


1877. Professor O. C. Marsh. 1881. Dr, Charles Barrois. 
1879. Professor E. D, Cope. 


VOL. XXXVII. C 


22 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Estimates for 
INCOME EXPECTED. 
Ls. 8... Sane 
Due for Subscriptions for Quarterly Journal... 3 5 4 
Due for Arrears of Annual Contributions ...... AO) © 
Due for Arrears of Admission-fees ........... 5 4 Do 
257 7 4 


Estimated Ordinary Income for 1881 :— 


Annual Contributions from Resident Fellows, and Non- 


residentsiof lSo9 tonlSGla ee seen 1400 0 O 
Admission=fees.. oii. csin «Us wise candela 252 0 0 
Compositions... a! als fuk Jo8s Srenaa2 eabene nancies een 241 10 0 
Annual Contributions invadvancer). 25h e eee 1212 0 

Dividends on Consols and Reduced 3 per Cents .......... 238 10 2 
Advertisements in Quarterly Journal..............0..... 7 10°70 


Sale of Transactions, Library-catalogues, Orme- 
rod’s Index, and Hochstetter’s New Zealand... 30 0 0 


Sale of Quarterly Journal, including Longman’s 


DECOUNE| 25.52 Cinnge a te a milecn cpa ace teialleuor ata keno 215 0 0 
Sale of Geological Map, including Stanford’s 
QCCOUNEM Ht ier crete laytelo i ciicoe 1 orice emer 25 0 0 
— 270 0 0 
sale of Stock for Library Catalogue) 73.5. i100 eee 268 10 0 


£2947 19 6 


J.GWY¥N JEFFREYS, Treas. 
14 Hiding Were 


FINANCIAL REPORT, 23 


the Year 1881. 


EXPENDITURE ESTIMATED. 


House Expenditure: 


eRe sand ENSUTANCOT cans ssececisd-ssewscascedes es 33 10 O 
ROE oe on Rea ea ene vce angels ctesaredoedieiesiae 25 0 0 
ie rae oer tonics de datciousaiona ce otanetweuins 40 0 0 
PISSED GLEE OR cr ee a )oloe cise saldniiclesiice eclecaenioees 740) Wa) 
House-repairs and Maintenance.................. 35 0 0 
PeeMeROE MCLE AMMAG oan cssjosrascvseecactesedeocisae 2070) 0 
Washing and sundry small Expenses ......... 40 0 0 
12D 2) MIGHT) LaseAndansossenocopseagenoLosuacoocs 20 OF 50 
233 10 0 
Salaries and Wages: 
BS AAMT ESCCECLATY) Gi si\-sessiccsensssceesvceseowieas 350 0 0 
re ee anes cect enue cacste denies Sawsesacte’s 140 0 0 
Assistant in Library and Museum ............... MOTO 0 
VOR SLOW ATO: iio. cece segeect eevee abeeastedeaces 105 0 0 
1S UO UISCUTE TG Ue RSA Ae Rae ee ee ee 40 0 O 
Ere el Ls Oper coe acta: cae cratic le sve sins aaisroaleSie oe 32 10 O 
Charwoman and Occasional Assistance......... 50 0 0 
Attendants at Meetings ...............000+Rsceoee 810 0 
PMCCOMM ATU Seresetinit nace sna nsicisesecweasinsadscseetee tS) a) 


Official Expenditure : 


SIDE OIILGIAY oe SRB eEE ROAR Ene cere eee rier 2a nO @ 
Miscellaneous Printing  ..0.s.cle.sceessceees ccna 2a 0) 10 
Diaeramistat Meetings .....2.....ss-0-0reeeeeeeneins DO 50 
Postages and other expenses ............ceeeeees com OwEO 
129) 05-0 
LL ETE 900060 6 UR COCO IOC Cer nienCI Cn ia TEz aren ir ne sa NE 150 0 0 
Aine PTIMPMEM ASE Seo Foye 5 oo o\iuo, 6-5 514 4 oie, sore = ald. olezeye olinye! aye SG 
Publications: 
SeeOO A CAIMMIA NS. cance oebab see ineis cetenascnaisniesine uses 50 0 0 
@uarterly Journal ../..........:- £915 0 O 
a - Commission, 
Postage, and Addressing... 85 0 0 
—- -1000 0 0 
si eOMBCMOWSS.- oon. -ceceesscssecnseevevagesmsessens 36 0 0 
Abstracts, including Postage ............0+.c000 95 0 O 
Wibtanyp Catalogue. ..2..-.n2.cececeecesoasnsennci 268 10 0 
1449 10 0 
Balance in favour of the Society........ ene LOO leas 


£2947 19 6 


Ne ne nl 


24 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Income and Expenditure during the 


RECEIPTS. i 
eg ds Eset 
Balance in Bankers’ hands, 1 January 1880. 14 4 1 
Balance in Clerk’s hands, 1 January 1880. 111 8 
— 15 15 9 
Compositions aces: « . Someta menor we ranratieiete easton 241 10 OU 
Arrears of Admission-fees.............. 88 4 0 
Admuissiom=tceslSS On) coch-pae une etait 308 14 0 
a 396 18 0 
Arrearaor Annual Contrlbubionsi nen ceeia ele 189 11 6 
Annual Contributions for 1880, viz. :— 
Resident Fellows ......... £1329 16 6 
Non-Resident Fellows ... 26 15 6 
——_—_— 1356 12 0 
Annual Contributions in advance...... Bebe ObSoSccK RS Ney (G) 
Journal Subscriptions mm advancela-....- era ieee 016 4 
Dividends ony Consols) 5 ee 20fa= MeO 
. Reduced 3 per Cents. ...... 30 Sine 
ee 


Taylor & Francis: Advertisements in Journal, Vol.35.. 717 O 
Publications : 


Salevordournaly Violsal=3nmice eee eeeces 126 13 8 

Ms VOLTS Rees aan neers SB) yy ak 
SalevorMl:ansachions ence eee OMA 9 
Salevon lnbrany Catalogel cuss. scascscecesoes my 0 
Salevol Geological: Menpiienecncenescaacseaee 26 0 10 
SalevormOnrmerod’slindexis. 0 aera 2 210 
Sale of Hochstetter’s New Zealand ......... 016 O 


*Due from Messrs. Longman, in addition to the 
above, on sournals Viola 30) sce sue ee seen eee 6b Seat 
tDue from Stanford on account of Geological Map SelOel 


£69) 1392 


LM 17 4 


We have compared the Books and 
Accounts presented to us with this 
statement, and we find them to agree. 


(Signed) J. CLARKE HAWKSHAW, 


F, G. HILTON PRICE, \ Auditors. 


27 Jan. 1881. 


FINANCIAL REPORT. 25 
Year ending 31 December, 1880. 
EXPENDITURE. 
General Expenditure: CSA / Ly) Ges de 
SCR ete sre t ate isceme en eebwhiayineaatvor is ousiedes So LO 
MBE UMSUPATICO I ye crs.e cc worsens seis scienwohie eer 12050 
PE UISK nd GUN Clee Reels ict cn ncnctee ese a rancedease acces 20 15 6 
EVOQUSE-FEMAITA Ns (een ciathialeces aceiee santa ee teeioc ve sees 33 16 8 
BETO ee ee ncaa cei etic te cocci asvacees oanhcs 388 15 O 
Ti EING, cae SSeS stein BAO CeCE ae eR ice eee ene By IIA 
Miscellaneous House-expenses.........0..ceee0s 7, 
SITILOTIGIAY | dbangetBoroakaeeueee Cho bdsnee nebecenper nce 20 10 4 
Miscellaneous Printing. cecc cs.) -cedescscesensne 2, dG 
Menta Meetim SS. .c..5- 0 cistvaviace ese sce Hapieusiorane US IG) 
—— go2 924 
Salaries and Wages : 
Assistant Secretary (five quarters) ............ 437 10 O 
CLSTES axed pe RS HOSE IRSA I ae ge nates eee 140 0 0 
Library and Museum Assistants ............... 140 8 4 
MIMO LSC MSEC WAL) nieces ececoniseaerisentiscese siecle 105 0 O 
MOUSE MIEA IG tye. ccs Sogo eso eee cc caide e's pees 40 0 0 
Wecasional Attendants: oi. so. ce .edesrccensercnaee 8 10 O 
BARC UMELETILS) wanes casey ie oetacls snus cincas access COMER 
——— 878 15 4 
LLIN? oe ne eee SOR eee 149 12 4 
Miscellaneous Expenses, including postages ........ Wl ie 
Wiierans at Meevings 22.25. 6 sc ieee ec bee ees 314 0 
exesamentsim 6100 Consol, . 2 ..0..0e0c2 sere ee 98 10 0 
Publications : 
reolOmed le Map rrcesccarscctssssossescanoss cca: Ih igs © 
OMEN aeVOlS lO) is agendas case tele eases eee 7 
3 Wo lithon ttaeca cece ashes £877 10 5 
Commission, 
Postage, and Addressing... 82 15 11 
6 4 
PMS ACU NH saat nal taete sna ts ing slala oc urisenitiacoasise 95) on okt 
—— 106418 0 
Balance in Bankers’ hands, 31 Dec. 1880.. 96 0 6 
Balance in Clerk’s hands, 31 Dec. 1880 .. 6 8 1 
——__—_ 1092 8 7 
AEA OM Lal 4: 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


26 


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28 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


AWARD OF THE WoLtAston MEDAL. 


The Reports of the Council and of the Committees having been 
read, the President, Roserr Eraerrmer, Esq., F.R.S., presented 
the Wollaston Gold Medal to Prof. P. Martrmy Duncan, M.B. 
Lond., F.R.8., F.G.8., and addressed him as follows :— 


Professor Duncan,— 


It is with no ordinary pleasure that the Council have awarded to 
you the Wollaston Medal, the highest honour that it is in their 
power to bestow, in recognition of the valuable services which you 
have rendered during so many years to the advancement of Geology, 
and especially of Paleontology; and I may add that it is equally 
productive of gratification to me that this honour is to be formally 
conferred upon you by my hands. Since the year 1863 paleonto- 
logists have been indebted to you for no fewer than twenty-six 
memoirs relating to the history, structure, and distribution of the 
fossil Actinozoa, a group which you have made peculiarly your 
own by long-continued and most careful researches. Further, you 
have enriched the publications of the Paleontographical Society 
with several most important treatises on the British Fossil Corals, 
supplementary or, rather, perhaps, complementary to the classical 
Monograph of MM. Milne-Edwards and Haime. 

These labours alone, and the value of their results, might have 
justified the Council in awarding you the Wollaston Medal; but 
besides your researches upon the Actinozoa, we have to point to 
several important papers upon the fossil Echinodermata, to others 

relating to subjects of Physical Geology (also freely touched upon 
in your more special memoirs), and particularly to your exceedingly 
important work in connexion with the Geological Survey of India, 
in describing the fossil corals of that Peninsula, and discussing the 
questions of both zoological and geological interest which naturally 
arise out of the study of those organisms. Few, mdeed, of our 
Fellows are in a better position to appreciate your valuable labours 
than myself; scarcely a day passes that I have not occasion to 
consult one or more of your contributions; and the more I consult 
them the more I am convinced of their value. Patiently and un- 
obtrusively, for nearly twenty years, you have followed out the 
line of research necessary for the fulfilment of your self-imposed 
task; you have sacrificed the advantages of professional life to 
devote your energies to the advancement of science; for seven 
years (from 1864 to 1870) you gave the Society the benefit of your 
services as one of its Honorary Secretaries, and for two years 
(1876, 1877) you worthily occupied the Presidential Chair. Such 
considerations as these would not alone, perhaps, have warranted 
the award of the Council; but the recollection of such services 
rendered to the Society is hardly out of place, as supplementing 
those more generally appreciable merits upon which the award was 


ANNIVERSARY MEETING—-MURCHISON MEDAL AND FUND. 29 


really founded. On all accounts it is with much pleasure that I | 
hand to you the Wollaston Medal. 


Professor Duncan, in reply, said,— 
Mr. PrustpEnt,— 


The gift which you have presented to me, in the name of the 
Geological Society, I receive with feelings of great respect and 
thankfulness. ‘This Medal comes to me bringing a twofold pleasure ; 
for it is a distinction which has been hallowed and ennobled from 
its reception by a long succession of illustrious men, amongst whom 
were the founders of our science, our teachers, and many of our 
best friends ; and because, in presenting it to me, you have spoken 
so sympathetically in appreciation of my scientific work—work 
which I have been enabled to bring before the world in consequence 
of the advantages which this Society has placed within my reach. 
Cheered by this expression of your approbation, I shall labour on 
in this our common and much-loved science, endeavouring always 
to merit the esteem of the Fellows of this Society. 


AWARD oF THE Murcuison MeEpAt. 


The Presipent then presented the Murchison Medal to Prof. 
_ ArcuipaLp Gernie, F.R.S., F.G.S., and addressed him as follows :— 


Prof. Gerx1E,— 


If any one Fellow of our Society more than another could be 
selected to receive the Murchison Medal for his valuable contri- 
butions to Geology, it would be yourself; since no man living has 
contributed more to the advancement of that science which it is the 
special object of our Society to cultivate and diffuse. Your labours 
in the field connected with your duties as Director of the Geolo- 
logical Survey of Scotland, your learned and valuable contributions to 
the Journal of our Society, the Transactions of the Royal Society 
of Edinburgh, the Glasgow Geological Society, and other pub- 
lications too numerous to mention, eminently qualify you to be the 
recipient of the Medal founded by your late chief and friend Sir 
Roderick Murchison. To enumerate your contributions to the 
literature of the geology of Scotland, or your many important 
writings connected with our science, would lead me too far 
—some thirty papers, besides educational works, have resulted 
from your industry and knowledge. Your able paper on the “ Old 
Red Sandstone of Scotland,” published in the Transactions of the 
Royal Society of Edinburgh, would of itself entitle you to the highest 
consideration of the Society. Able, indeed, are other contributions, 
especially those “On the Chronology of the Trap Rocks of Scot- 


VOL. XXXVII. d 


30 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


land,” “On the Date of the last Elevation of Central Scotland ” (in 
vol. xviii. of our Journal), ‘‘On the Phenomena of Succession 
amongst the Silurian Rocks of Scotland” (Trans. Glasgow Geol. 
Soc. vol. ii.), and “ On Earth Sculpture.” The Council believed, 
too, that it weuld be gratifying to you to receive as a mark of their 
esteem and sense of your untiring labours, the Medal founded by 
one with whom in earlier life you were closely associated, and 
whose endowed Chair of Geology in the University of Edinburgh 
you have been the first to fill. 


Prof. Grrcin, in reply, said,— 
Mr. PREsIDENT,— 


If any thing could add to the gratification with which I receive 
this honour from the Geological Society, it would be the very kind 
and flattering terms in which you, Sir, have made the award, and 
the Fellows have been pleased to receive the announcement. The 
Geological Society of London has always seemed to me to be truly 
the Geological Society of the British Empire, electing its Fellows 
and bestowing its rewards, not in a local, but in a truly catholic and 
generous spirit. This conviction was renewed and strengthened in 
my mind on receipt of the unexpected intimation of the bestowal of 
one of the Society's Medals upon myself, as my contributions to 
science have but rarely appeared in the Society’s publications, and 
I am so seldom able to be present within the Society’s walls. I 
receive the Medal with peculiar pleasure ; first, as a valuable mark 
of the Society’s recognition, and next, as another link of association 
with the memory of Murchison, which is one of the most precious 
possessions of my life. 


AWARD oF THE LYELL MEDAL. 


The Prestprent next handed the Lyell Medal to Mr. Wartneron 
W. Suyre, F.R.S., F.G.S., for transmission to Dr. J. W. Dawson, 
F.R.S., F.G.8., of Montreal, and addressed him as follows :— 


Mr. Warineton SuytH,— 


Sir Charles Lyell, in founding the Medal that bears his name, 
intended that it should serve as a mark of honorary distinction, 
and as an expression on the part of the governing body of the 
Society of their opinion that the Medallist has deserved well of 
science. I need hardly say that the Council, in awarding the Lyell 
Medal to Principal Dawson, have done so with a sincere apprecia- 
tion of the high value of his truly great labours in the cause of 
Paleontology and Geology. When I refer to his published papers, 
I find that they number nearly 120, and that they give the results 
of most extensive and valuable researches in various departments of 
geology, but more especially upon the paleontology of the Devonian 


ANNIVERSARY MEETING-——-LYELL MEDAL AND FUND. 31 


and Carboniferous formations of Northern America. No fewer than 
30 of these papers have appeared in the pages of our own Quarterly 
Journal. Considering the nature of these numerous contributions, 
the Council would have been fully justified in awarding to Dr. 
Dawson one of its Medals, upon the sole ground of the value of their 
contents ; but these are far from representing the whole of the results 
of his incessant activity in the pursuit of science. His ‘ Acadian 
Geology, ‘ Post-pliocene Geology of Canada,’ and ‘ Fossil Plants of 
the Devonian and Upper Silurian of Canada’ are most valuable con- 
tributions to our knowledge of North American Geology ; whilst in 
his ‘ Archaia,’ ‘The Dawn of Life,’ and other more or less popular 
writings, he has appealed, and worthily, to a wider public. We are 
indebted to his researches for nearly all our knowledge of the fossil 
flora of the Devonian and other Precarboniferous rocks of America, 
and of the structure and flora of the Nova-Scotian coal-field; and, 
finally, I must refer especially to his original investigation of the 
history, nature, and affinities of Hozoon. These researches are so 
well known that they have gained for Dr. Dawson a world-wide 
reputation ; and it is asa slight mark of their esteem, and their high 
appreciation of his labours, that the Council have awarded to him 
this Medal, which I will request you to forward to him, with some 
verbal expression of the feeling with which it is offered. 


Mr. Warineron W. Suytu, in reply said— 


That it gave him much pleasure to receive this Medal for Dr. 
Dawson, who much regretted that he was unable personally to be 
- present, but had addressed a letter to the President expressing his 
sense of the honour conferred upon him, in the following terms :— 


*“[ regret that distance and the claims of other duties prevent 
me from appearing in person to express to the Geological Society 
my sense of the honour conferred upon me by the award of the 
Lyell Medal. 

“This expression of approval on the part of those whose good 
opinion I value so highly is doubly grateful to one who is so deeply 
sensible of the imperfection of scientific work done in circumstances 
of isolation from the greater centres of scientific life, and under the 
pressure of the severe demands made in a new and growing country 
on those engaged in educational pursuits. 

“Tt is further especially gratifying to me that this token of your 
kindly recognition is connected with the illustrious and honoured 
name of Sir Charles Lyell. Forty years ago the foundation of my 
geological education was laid by the late Prof. Jameson and other 
able educators in natural science, his contemporaries, in Edinburgh ; 
but in so far as I have been able to build any thing worthily on 
this substructure, the credit is due to the study of the ‘ Principles 
of Geology, and to the personal friendship and generous kindness 
of Sir Charles Lyell more than to any other cause.” 


32 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


AWARD oF THE Biespy MEDAL. 


The Presrpent then handed the Bigsby Medal to Prof. Morris, 
F.G.S., for transmission to Dr. Cuartrs Barrors, and addressed him 
as follows: 


Prof. Morr1s,— 


The Council of this Society have selected Dr. Charles Barrois to be 
the recipient of the Bigsby Medal, and have awarded it to him 
for his numerous papers and contributions to geological science. 
Dr. Barrois’s chief or most important work (written in the year 
1876, and published at Lille) is entitled ‘ Recherches sur le terrain 
erétacé supérieur de l’Angleterre et de l'Irlande,’ a production almost 
exhaustive in its description of the Cretaceous rocks of England 
and Ireland, and of the utmost value to English students of 
geology. Dr. Barrois in this work has been the first to attempt to 
arrange the English Cretaceous rocks in Paleontological zones, and 
eminently has he succeeded in defining and correlating the horizons 
of France and Britain. He is also the author of a ‘ Mémoire sur 
le terrain crétacé du Bassin d Oviedo, Espagne,’ with a paleonto- 
logical description of the Kchinodermata by Gustave Cotteau. His 
great industry and untiring zeal for geological science entitle him 
to the consideration of the Council; and I therefore beg you to for- 
ward to him the Bigsby Medal as our recognition of his services, 
and, according to the wishes of the founder, we look forward to 
other and equally valuable contributions. 


Prof. Morris, in reply, said :— 
Mr. PRESIDENT, 


I have much pleasure in receiving the Bigsby Medal for transmis- 
sion to Dr. C. Barrvis in recognition of his labours for the promotion 
of geological science. I feel assured that while he will fully appre- 
ciate the award, it will be also satisfactory to his geological col- 
leagues in the north of France, and especially to that energetic and 
accomplished geologist, Prof. Gosselet, under whose tuition and en- 
couragement he has been partly stimulated to prosecute those 
researches which have this day been recognized by the Council of 
the Geological Society. 

Not only, Sir, as you have justly stated, has Dr. Barrois contri- 
buted to our knowledge of’ the physical and paleontological con- 
ditions of the Cretaceous rocks of the north of France and of this 
country, but among other researches he has been occupied with the 
Paleeozoic strata of Brittany and also of Northern Spain, on which 
latter, he informs me, a memoir will shortly be published. In con- 
lusion I may read the following extract from a letter received from 
Dr. C. Barrois, who regrets exceedingly that a previous engagement 
rendered it impossible for him to express personally his grateful 
sense of the honour done to him by this award. He says :— 


ANNIVERSARY MEETING——-WOLLASTON FUND. 33 


‘Will you please present my best thanks to the Society for 
having judged my labours for the advancement of geological science 
worthy of the Bigsby Medal. 

“Tt is a great and quite unexpected honour for me to have my 
name on the list of all the good geologists who have received the 
Medals of the old and illustrious London Geological Society. By 
the award of the Bigsby Medal I feel myself honoured in the highest 
degree, as nothing can be more precious to a scientific man than 
the recognition of his endeavours by the masters and leaders of his 
own special science. I was already greatly indebied to English 
geologists as having so often used and profited by their works during 
my travelsin Great Britain. I now incur a new debt to them, as 
the Bigsby Medal conferred upon me is given, according to the 
donor’s bequest, ‘to those not too old for further work.’ Iam thus 
mightily stimulated to work for the performance of my duty ; I will 
endeavour to discharge it and to become worthy of so high an en- 
couragement.” 


AWARD OF THE WoLLASTON DoNATION Funp. 


In handing to Prof. J. W. Jupp, F.R.S., Sec. G.S., the balance of 
the Wollaston Donation Fund for transmission to Dr. Ramsay H. 
Traquair, F.G.S., the Presrpznt said :— 


Professor J upp,— 


In handing to you, to be forwarded to Dr. Traquair, the balance. 
of the proceeds of the Wollaston Donation Fund, I have to request 
that you will inform him of the feeling of the Council, that it is 
rarely that they can have the opportunity of awarding this fund to 
a more able and accomplished naturalist than himself. Huis long- 
continued researches upon the Ganoid Fishes of the Carboniferous 
formation have rendered his name eminent in this department of 
Paleontology. As an accomplished anatomist and zoologist, we 
must have every confidence that his treatment of these Vertebrates in 
the memoir which he is contributing to the publications of the 
Palzontographical Society will be of the most careful and judicious 
description, whilst the value of this and his other works is vastly 
enhanced by the beautiful figures with which he illustrates them. 
Under these circumstances it affords me much pleasure to place in 
your hands, for transmission to Dr. Traquair, the balance of the 
Wollaston Fund, which I hope he will receive as some recognition 
on the part of the Society of the value of his researches, and, at the 
same time, as a small aid to him in further prosecuting them. 


Prof. Jupp, in reply, read the following communication, received 
from Dr. Traquair — 


“Mr. PresiDENT,— 


“Permit me cordially to thank the Geological Society for the 


34 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


honour which it has conferred upon me by adding my name to the 
list of those who, in former years, have received this award. 

“Tf the subject of Fossil Ichthyology is not only fascinating, but 
essential to the student interested in the great problems of Verte- 
brate Morphology and Descent, it cannot fail also to be of import- 
ance to the geologist, who, calling to his assistance every available 
branch of natural knowledge, seeks to arrive at a scientific concep- 
tion of the past history of the earth and its living inhabitants. It 
is, therefore, very gratifying to me to receive from the Geological 
Society of London this very unexpected token of sympathy and 
recognition, considering that the amount of work which I have been 
able to achieve in this field is as yet comparatively small. It will, 
however, stimulate me to further exertions; for the field is itself a 
large one, and in spite of the magnificent labours of Agassiz and 
the accumulated researches of many other writers, still affords room 
for plenty of work.” 


AWARD oF THE Murcuison GrotoeicaLt Funp. 


The PrestpEnr next presented the balance of the proceeds of the 
Murchison Donation Fund to Franwx Rvurzey, Esq., F.G.8., and 
addressed him in the following words :— 


Mr. Ruritzy,— 


For many years you have devoted your time and attention to the 
microscopical structure of rocks and rock-forming minerals, a branch 
of scientific research of the highest importance to the petrologist 
and geologist; and now that our attention is being so much drawn 
to the structure of the metamorphic and igneous rocks, with a view 
to a better nomenclature and a revision of old and obsolete views, 
the Council of our Society believed that in your hands good work 
would still be carried on; they, therefore, have awarded to you 
the balance of the Murchison Fund, which I have much pleasure in 
handing to you in recognition of your past researches, the results of 
which you have from time to time communicated to the Journal of 
the Society. Few are more aware than myself of the interest you 
take in this branch of study, and it affords me much gratification 
to be the medium of conveying to you the appreciation of the Council 
and the accompanying fund. 


Mr. Rurtry, in reply, said that he felt deeply the honour thus 
conferred upon him, which was additionally gratifying to him per- 
sonally, as it was associated with the name of a former distinguished 
Director-General of the Geological Survey. With regard to his own 
special work, to which the President had alluded in such kind terms 
of praise, he remarked that in all such work the investigator is 
particularly liable to fall into error, and that all special microscopical 
researches upon petrological subjects need to be constantly checked 
by observations in the field. It was a further gratification to him 
to receive the award of the Council, for which he wished to express 


ee ee eee 


ANNIVERSARY MEETING——-LYELL FUND. 35 


his gratitude, from the hands of an old teacher, to whose instructions 
he was indebted for the habits of minute and careful investigation 
inculcated in his lectures. 


AWARD oF THE LYELL GroLogicaAL Funp. 


In presenting to G. R. Vinu, Esq., one moiety of the balance of 
the proceeds of the Lyell Donation Fund, the President addressed 
him as follows :— 


Mr. Vinzt,— 


A moiety of the balance of the proceeds of the Lyell Geological 
Fund has been awarded to you by the Council of the Geological 
Society. In making this award the Council were actuated in part 
by the wish to express their sense of the value of your researches 
upon the fossil Bryozoa of the Paleozoic rocks, as evinced especially 
by your published writings on the Diastoporide, an exceedingly 
dificult group, and in part by their desire to assist you in the 
further prosecution of your investigations. I have much pleasure 
in handing to you this small testimony of the appreciation of the 
Council. 


Mr. Vins, in reply, said :— 
Mr. PrrstpEnt,— 


I receive through you this token of recognition, on the part of the 
Council, of my humble labours, with mingled feelings of pleasure 
and pride. I cannot, at present, understand the reason why I 
should be selected as one of the recipients of the Lyell Fund. It 
cannot be for what I have done for science ; for, as yet, my name is 
young; it must be that, having looked upon my labours with favour, 
the Council desires to stimulate me to further exertions. If such be 
the motive, I shall do my utmost to fulfil the higher promise which 
my hitherto crude efforts have awakened. When, some years ago, I 
began to study the Carboniferous Polyzoa, I wrote to Prof. Duncan, 
F.R.S., asking for certain information respecting these Polyzoa; he 
wrote me back word saying that he did not know much about the 
species himself, and he did not think that there were many men in 
England who did; but he counselled me to go to work and find out 
for myself, and then he and others would be glad to know. Accept- 
ing this advice, I set to work, with results which you and the 
Council so kindly acknowledge to- day by this award. 

From your hands, Mr. President, I feel also proud in accepting 
the award. Many years ago, when poor and unknown, I sought 
information on this and other things from the cases of the Museum 
of Practical Geology. On that occasion you helped me by word and 
deed to name my private series of fossils—a kindness on your part 
that I never have forgotten or shall forget. 


36 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


To the Fellows of the Geological Society I also tender my thanks 
for the manner (judging by the discussions which have followed the 
reading of my papers) in which they have received my criticisms on 
a most difficult group of fossils. I may differ from many of you in 
the future, but I shall hope to do so in such a manner as to show 
that I do not forget that we are fellow-students and at the same 
time gentlemen. 


The PresipEent then handed to Prof. H. G. Szerezy, F.R.S., F.GS., 
for transmission to Dr. Anton Frirson, of Prague, the second moiety 
of the Lyell Donation Fund, and said :— 


Professor SEELEY,— 


The Council has awarded a portion of the Lyell Geological Fund 
to Dr. Anton Fritsch, Professor of Zoology in the University of 
Prague, in recognition of his valuable contributions to paleontology. 
Dr. Fritsch is an accomplished zoologist, who has enriched his: 
studies of many groups of fossils, invertebrate and vertebrate, with 
admirable knowledge of existing life. During the last thirty years 
Dr. Fritsch has published about one hundred and twenty memoirs, 
many of which relate to paleontology and geology. Besides scattered 
papers on Hozoon, Callianassa, and other subjects connected with the 
fossil fauna of Bohemia, Dr. Fritsch has also published some standard 
works monographing the fossils of his native land. These comprise 
memoirs on the Cretaceous Cephalopods (1872), the Cretaceous Rep- 
tiles and Fish (1878), and his great work on the Fauna of the Per- 
raian Rocks (still in progress), of which two volumes, devoted to Am- 
phibia, have been issued. ‘These volumes are excellent examples of 
descriptive work, illustrated worthily, and this award is especially 
intended to mark the sympathy of the Council with Dr. Fritsch in 
his endeavours to adequately make known the Permian fauna, and 
in the hope that the fund may assist him in completing a work which 
has already taken high rank among paleontological monographs. 


Prof. SEELEY, in reply, said :— 


Mr. PrestpEnt,— 


Dr. Fritsch will, I believe, appreciate the honour of the award 
which you have made, no less than the words in which you have ex- 
pressed the esteem in which his work is held in this country. I 
have seen the materials upon which his studies of the Amphibia are 
founded, and doubt whether any more important work has been 
done in paleontology during the past year, or whether it could have 
been accomplished at all by any one less skilled in the methods of 
zoology than Dr. Fritsch. His enthusiasm for research has given 
to his works a wealth of illustration that will help materially in the 
advance of knowledge, while by electrotyping the specimens he has 
rendered them easily available for study by all anatomists. I shall 
have great pleasure, Mr. President, in informing Dr Fritsch of the 
kindly way in which your award has been received by the Society. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. iy) 


THE ANNIVERSARY ADDRESS OF THE PRESIDENT. 
Rosert HruEriver, Hsq., F.R.S. 


In accordance with the usual practice, 1 must commence my 
Address with brief Obituary Notices of some of the more promi- 
nent Fellows and Members of the Society whose loss we have 
had to deplore since the last Anniversary Meeting. 


SEARLES VALENTINE Woop, born Feb: 14, 1798, died Oct. 26, 
1880. He became a Fellow of the Geological Society in 1839. 

Mr. Wood was born on St. Valentine’s day; hence his name. 
He went to sea as a midshipman in the ‘ Thames’ (one of the East- 
India Company’s mercantile fleet) in 1811, and continued in that 
service until the year 1826, when, being disappointed in obtaining 
the command of a ship that had been promised him, he retired 
from a maritime life and devoted himself to paleontological 
studies. Settling in his native place in Suffolk, he gave the 
larger part of his attention to the Crag: but he also collected ex- 
tensively from the Hampshire Tertiaries ; and for the purpose of 
working out the relations of these to the beds of the Paris basin, 
he formed an extensive collection of the French Eocene Mollusca. 

From these materials, and from correspondence with Deshayes 
and other French savans, he was prepared to have taken up the 
description of the English Eocene Mollusca long before he actually 
did so, circumstances having determined his undertaking the de- 
scription of the Mollusca from the Upper Tertiaries first. He 
also formed a considerable collection of recent Mollusca for com- 
parison in working out the relations of the Mollusca from Tertiary 
formations. Having left Suffolk from ill health, and settled in 
London, he was in 1837 introduced to Sir Charles (then Mr.) 
Lyell, and was associated with him in the endeavour in which 
Lyell was then mainly engaged, to work out a better knowledge of 
the Tertiary formations, which up to a period not long before that 
time had been regarded as of small account in comparison with 
the “Secondary” group. In this task Lyell relied principally on 
S. V. Wood and the late G. B. Sowerby for the determination of 
the identity of the Molluscan remains from various countries with 
those found fossil in England, and with the Molluscan fauna lying 
in existing seas, so far as these were then known. Mr. Wood 
also, for a few months about this time, acted as Curator of the 
Museum of the Geological Society. 

Urged to the task by Lyell, he commenced (with the coopera- 
tion of the present Mr. G. B. Sowerby as engraver and intended 
publisher) the description of the ‘ Crag Mollusca; ’ and considerable 
progress had been made with the manuscript and plates of the 
first or “ Univalve” part of this work, when the Paleeontographical 
Society was formed, in 1847; this part formed the first volume of 
the magnificent series of scientific publications which have been 


VOL. XXXVII. @ 


38 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


issued by that Society. The rest of the ‘ Crag Mollusca’ followed 
in subsequent years ; and upon the completion of this work, and in 
recognition of his labours generally in connexion with the Tertiary 
Mollusea, the Council of the Geological Society awarded to Mr. 
Wood in the year 1860 the Wollaston Medal. A large supple- 
ment to this work, embodying the discoveries which had subse- 
quently accumulated, was prepared by Mr. Wood; and this, ac- 
companied by an introduction describing geologically the formations 
from which the remains embraced by the work had been obtained, 
from the pen of his son and of Mr. F. W. Harmer, was issued by 
the Paleontographical Society in 1871 and 1873. <A second sup-. 
plement followed this in 1879; and Mr. Wood was actively en- 
gaged, up to the day of his seizure with fatal illness, on another 
small addition. On the completion of the description of the Mol- 
luscan remains from the Crag, Mr. Wood presented the unrivalled 
collection of them, which he had been forming during thirty years, 
to the nation, in order that, by being preserved intact in the 
British Museum, the types of all the forms which had been de- 
scribed and figured by him in his work (save two or three which 
belonged to other persons) might be available for examination and 
comparison by naturalists engaged in similar labours. 

He also presented to the nation the valuable collection of ver- 
tebrate remains (including among them the unique jaws of Alligator 
hantomensis and Microcherus erinaceus) which he had, in 1843-5, 
extracted from the Hocene Freshwater beds of Hordle Cliff, 
from which beds up to the time when he commenced forming this 
collection no such remains had been known. These he partially 
fioured and described in the London Geological Journal; but the 
stoppage of that publication brought this part of his labours to an 
unexpected termination. In 1858, having the advantage of an 
unrestricted manipulation of the more extensive collection of 
Eocene Mollusca which had been formed by his friend F. E. 
Edwards, he commenced the description of the Hocene Bivalvia, 
Mr. Edwards taking upon himself (and having commenced before 
this) the Cephalopoda and Gasteropoda. Several parts of this 
work were issued by the Paleontographical Society. Failure of 
health put a somewhat premature period to Mr. Edwards’s share 
in this work ; but Mr. Wood continued his labours for some years 
longer, relinquishing them only when, shortly before Mr. Edwards’s 
death, his collection was acquired by the British Museum. During 
the time in which it remained in Mr. Edwards’s possession he was 
accustomed to place in Mr. Wood’s care, for study at his leisure, 
all specimens he possessed which in any way illustrated the subject 
in hand; but this the transfer of the collection to the British 
Museum rendered impracticable ; and as it was Mr. Wood’s feeling 
that, in addition to that opportunity for careful study, all forms as 
to which any doubt existed ought to be carried abroad and com- 
pared with those in the museums of France and Belgium, if justice 
was to be done to the subject, and as at his advanced age he was 
unable to accomplish this, he on the issue of the part in the volume 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 39 


of the Paleontographical Society for 1870 relinquished the further 
prosecution of the ‘ Hocene Bivalvia,’ although he subsequently 
added a small contribution (both to Mr. Edwards’s and his own 
portion) on special groups of Eocene Mollusca, which he was able 
to do from resources afforded by his own collection and the collec- 
tions of some friends. 

He maintained his activity both of mind and body up to the 
day of his seizure with fatal illness, which took place on the 21st 
and terminated with his death on the 26th of October last. He 
was buried in the churchyard of Melton, near Woodbridge, in 
view of the Crag of which the study had occupied so much of his 
life. 


JOHN JEREMIAH BiasBy, M.D., F.R.S., one of the oldest of our 
Fellows, and a man endeared to all who knew him by many amiable 
qualities, died at his residence in Gloucester Place on the 10th 
February in the present year. He was theson of Dr. John Bigsby, 
and was born at Nottingham on the 14th August, 1792; so that at 
his death he was in his 89th year. 

Deciding to follow his father’s profession, he took his degree 
of Doctor of Medicine at Edinburgh; and soon afterwards he 
must have entered the military service, as, from a chance note 
in one of his books, it appears that he was at the Cape in 1817. 
In 1818 or 1819 he went to Canada as the Medical Officer to a 
large detachment of a German Rifle Regiment in the British 
Service; and in Canada he remained for several years performing 
a variety of commissions intrusted to him by the Government. 
Thus in the summer following his arrival in the Colony he was 
sent to the Hawkesbury Settlement, where a severe epidemic of 
typhus fever had broken out among the miserable Irish immigrants ; 
and in the next year his marked taste for Geological studies led 
to his being commissioned to travel through Upper Canada and 
make a general report upon its Geology. In 1822 he was ap- 
pointed British Secretary and Medical Officer to the Boundary 
Commission, which had already been in existence for three or four 
years. In the journeyings to and fro necessitated by these em- 
ployments Dr. Bigsby had many opportunities of making himself 
acquainted with the geological phenomena of regions then very 
little known ; and it is chiefly to these that his published papers 
relate. In 1823 he was elected a Fellow of this Society. 

He appears to have returned to England about the year 1827, 
and then proceeded to practice his profession at Newark in Not- 
tinghamshire, remaining there until 1846, when he came to London, 
where he afterwards resided until the end of his life. R 

Dr. Bigsby was elected a Fellow of the Royal Society in 1869, 
and received the Murchison Medal from this Society in 1874. 

His published papers are about twenty-seven in number; the 
earliest, entitled, ‘‘ Remarks on the Environs of Carthage Bridge, 
near the mouth of- the Genesee river,’ appeared in Silliman’s 
American Journal in the year 1820. It was followed by several 

e2 


40 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


other papers on geological subjects in the same periodical. In 
1823, the year of his election, he forwarded a paper to this Society 
‘“*On the Geography and Geology of Lake Huron,” which was 
published im 1824, in the first volume of our ‘Transactions.’ 
Until his return to England his contributions to scientific litera- 
ture made their appearance chiefly in American publications ; and 
for some years after his return he seems to have done but little, 
his medical practice probably occupying his time very fully. After 
his removal to London, however, several papers from his pen were 
communicated to the Society ; and among them may be noticed 
especially his paper ‘“‘ On the Paleozoic Basin of the State of New 
York,” which appeared in the Quarterly Journal for 1858, and 
was followed by “An Inquiry into the Sedimentary and other 
Natural Relations of the Paleozoic Fossils of the State of New 
York,” published in 1859; his note “ On the Erratics of Canada,” 
printed in the Journal for 1851; and his last communication to 
our ‘ Proceedings,’ ‘‘On Missing Sedimentary Formations,” pub- 
lished in 1864. Other memoirs of his appeared in various perio- 
dicals, such as the ‘ Philosophical Magazine’ and the ‘ Annals and 
Magazine of Natural History,’ the last of them bearing date 1867. 

The narrative of his Canadian travels appeared in 1850, in two 
octavo volumes, under the fanciful title of ‘ The Shoe and Canoe,’ 
Although evidently written by the light of knowledge subsequently 
obtained, it furnishes an interesting-account of the condition of 
Canada some sixty years ago, and is renderedamusing by the dis- 
play of that naive geniality mingled ‘with ‘shrewdness which cha- 
racterized the author to the end of his life. It contains many 
scattered popular notes on geological and other natural-history 
subjects. 

During the last twenty years of his life Dr. Bigsby was engaged 
in a work of great labour and research, the first outcome of which 
was the ‘Thesaurus Siluricus,’ published, with the aid of a grant 
from the funds of the Royal Society, in 1868. It is a list of 
described Silurian fossils, classified both zoologically and in accord- 
ance with their distribution in time; and although, no doubt, errors 
and defects may be detected in it, it will remain for a long time a 
most useful aid to the student of Silurian Paleontology, and a 
monument of the untiring industry of its author.’ The’ same 
things may be said, perhaps in still stronger terms, of ‘a ‘second 
publication of the same nature, the ‘Thesaurus Devonico-Carbo- 
niferus,’ which appeared in 1878, a wonderful example of industry 
and research in a man eighty-five years old; and not content with 
these labours, Dr. Bigsby, to the close of his life, was hard at work 
upon.a Permian Thesaurus, the MS. of which is left in an ad- 
vanced state. The last books borrowed for the compilation of this 
work were only returned to the Society’s Library when Dr. Bigsby 
took finally to his bed. 

The memory of our late lamented Fellow will be worthily kept 
alive by the Medal which he founded in the year 1877. The 
greater part of Dr. Bigsby’s fortune having accrued to him by 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 41 


marriage, he gave himself only a life-interest in it, and decided 
that the whole amount at his decease should go to the relations of 
the ladies whom he had married and survived. Hence, from de- 
lhieacy of feeling, he decided that instead of bequeathing a sum of 
money to the Society for the purpose of founding a Medal, he 
would provide the necessary funds out of his income during his 
life ; and accordingly, in the above year, he gave the Society a sum 
of money the interest of which was to be devoted to providing a 
Medal to be given biennially, preferentially to those who had studied 
American geology, with the further stipulation that the receiver 
should be not more than forty-five years old, and *‘ thus probably not 
too old for further work, and not too young to have done much.” 
At first the Medal was to have been of bronze and accompanied by 
a sum of money, the balance of the proceeds of the fund; but 
subsequently the founder increased the amount of his donation suf- 
ficiently to enable the Council to give a Gold Medal. In 1877 the 
Medal was awarded to Prof. O. C. Marsh, and in 1879 to Prof. E. 
D. Cope; this year it goes to Dr. C. Barrois. 


Rev. James Crirron Warp. The announcement of the death 
of Mr. Ward must have been to most of his many friends wholly 
unexpected, both on account of the early age at which he passed 
away, and the very brief illness which preceded his decease. 

After a weakly boyhood he entered the Royal School of Mines 
as a student in 1861, entirely through my suggestion, and gained 
the Edward Forbes Medal and prize of books in 1864. In the 
following year he joined the Geological Survey, and was sent down 
to Yorkshire. He worked there on the Millstone-grit and Lower 
Coal-measures in the neighbourhood of Sheffield, Penistone, Hud- 
dersfield, Halifax, and Leeds. Though Ward was never of robust 
appearance, he obviously improved in health after leaving the School 
of Mines ; so well did the laborious but healthy work of the Survey 
agree with him. 

In 1869 he was transferred to Keswick ; and the change from a 
colliery district to a locality not only devoid of coal-pits, but one 
in which wild Nature puts forth all her charms, was in the highest 
degree pleasing to him. At Keswick his activity became twofold. 
His Survey work and its results are now represented by his Geo- 
logical Survey memoir on the Geology of the Northern part of 
the English Lake District (published in 1876), and by numerous 
maps and sections. He also contributed to the Geological Society, 
and to various periodicals, many papers bearing on the structure 
of the Lake Country. Of these may be mentioned, in the first 
place, two on its glaciation, entitled, “The Origin of some of the 
Lake Basins of Cumberland ” (Quart. Journ. Geol. Soc. 1874), and 
“The Glaciation of the Southern Part of the Lake District” &c. 
(Quart. Journ. Geol. Soc. 1875). In both papers the origin of 
the lakes is discussed ; and (as regards the English Lake country) 
the original investigations of the author tend to confirm the views 
so long held by Prof. A. C. Ramsay. ‘These papers are illustrated 


42 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


with sheets of sections of the highest interest and value; and to make 
the work more complete, the results of a series of soundings care- 
fully taken on most of the lakes by this indefatigable worker are 
also given. 

In the years 1875 and 1876, and more recently, microscopical 
examination of the rocks of the Lake District occupied much of 
his time. Of papers on this subject I may here note one “On 
the Granitic, Granitoid, and Associated Metamorphic Rocks of 
the Lake District,” the first part of which appeared in the Quart. 
Journ. Geol. Soc. for 1875, and the second in the volume of the 
same periodical for 1876. Another paper is entitled, ‘“ Notes on 
the Comparative Microscopic Rock-Structure of some Ancient and 
Modern Volcanic Rocks,” and appeared in the Quart. Journ. Geol. 
Soc. for 1875. Among his latest contributions to geological litera- 
ture may be mentioned “ Notes on the Geology of the Isle of Man,” 
which appeared in the Geological Magazine for January, 1880. The 
following is by no means an exhaustive list of his productions :— 

1. In 1868, “ Internal Fluidity of the Harth.” Geol. Mag. v. 

p- 581, 582. 

2. In 1869, “ Suggestions as to Geological Time.” Geol. Mag. 
vi. pp. 8-13. 

3. In 1869, “On Beds of supposed Rothliegende Age near 
Knaresborough.” Quart. Journ. Geol. Soc. xxv. pp. 291-297. 

4. In 1870, “On the Denudation of the Lake District.” Geol. 
Mag. vii. pp. 14-17. 

5. In 1871, “‘The Development of Land.” Geol. Mag. vin. 
)0o TALS 

6. In 1872, “On Rock-staming.” Geol. Mag. ix. pp. 389-391. 

7. In 1873, “ On the Scenery of the English Lake District, 
geologically considered.” Brighton, Proc. Nat. Hist. Soc. xx. 
pp- 39-44. 

8. In 1873, “On Rock-fissuring.” Geol. Mag. x. pp. 245-248. 

9. In 1873, “ The Glaciation of the Northern Part of the Lake 
District.” Quart. Journ. Geol. Soc. xxix. pp. 422-441. 

10. In 1870, “* Coral Reefs and the Glacial Period.” Quart. 
Journ. Sci. 1. pp. 170-178. 

But the most characteristic side of his untirmg energy, and 
perhaps its most important one, was the zeal with which he worked 
for the diffusion of scientific knowledge while in Cumberland. 
Before leaving Yorkshire he had written a small elementary book 
on Physics ; and one of the firstfruits of his educational activity 
at Keswick was a similar work on Geology, composed of nine 
lectures delivered in the first place before a school audience, and 
secondly before the Keswick Literary Society. Being simple, 
clear, and free from unnecessary technicalities, his lectures soon 
became popular, and the lecturer himself acquired influence. 

As the originator and main support of the Cumberland As- 
sociation for the Advancement of Literature and Science, and 
of most of the local societies connected with it, he accomplished a 
work which it may be hoped will not now be suffered to languish, 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 43 
but will remain a lasting monument of his beneficent activity. A 
elance at the outer cover of the ‘ Transactions of the Cumberland 
Association’ (Part IV. was published at the beginning of this 
year) shows the date at which each of the associated societies was 
founded, and discloses the fact that only one of them (at White- 
haven) existed before Ward’s appearance in the county. The dates 
of the others vary from Keswick 1869 to Silloth, the latest, 
1879. 

He married at the beginning of*the year 1877, and very shortly 
aiter left the Lake country to do field-work in the lone barren 
district of Bewcastle, on the Lower Carboniferous rocks, wintering, 
however, in Keswick as before. But on finishing the Bewcastle 
work he made preparations for taking Holy Orders, and was 
licensed to the curacy of St. John’s, Keswick, in December 1878. 
He was as successful in his new duties as he had been as a geo- 
logical surveyor, and was appointed, at the beginning of last year, 
to the vicarage of Rydal. But he was scarcely established in his 
new home when a brief illness, which only at the last seemed 
dangerous, caused his departure, at the age of thirty-seven years, 
leaving behind him a widow and two children. 

His genial disposition, and the absence in him of the least 
approach to the temper of the dogmatist, caused him to number 
among his friends men of every shade of speculative opinion. It 
was this amiability, in addition to his ability as a lecturer and the 
single-mindedness of his desire for the spread of knowledge, which 
made him so successful in connexion with the Cumberland Asso- 
ciation, when the simple fact of his not being a Cumbrian by birth 
would have been fatal to any merely active and zealous man. 


Davin Tuomas ANsTED was born in London on the 5th Feb- ° 
ruary, 1814, and, after education at a private school, entered the 
University of Cambridge as a member of Jesus College. He was 
32nd Wrangler in the year 1836, and proceeded to the degree ot 
M.A. in due course in 1839. About four years afterwards he was 
elected to a fellowship on the Ley Foundation of Jesus College, 
which he retained for about eight years. In 1840 he was elected 
Professor of Geology at King’s College, London, which office he 
resigned in 1853. For some time, from 1845, he held the 
Lecturership on Geology at Addiscombe, and was also Professor 
of Geology at the College of Civil Engineers, Putney. From 1844 
to 1847 he was Vice-Secretary of the Geological Society and Editor 
of the Quarterly Journal. He became a Fellow in 1838. He was 
elected to the Fellowship of the Royal Society in 1844. In 1868 
he was appointed Examiner in Physical Geography to the Science 
and Art Department. 

By degrees his attention became diverted from the theoretic to 
the practical aspect of his favourite study ; and for the last thirty 
years of his life he acted professionally as a consulting geologist 
and mining-engineer. For some time before his death he was in 
failing health; and he expired on the 13th May, 1880, at his 
residence, Melton, near Woodbridge, Suffolk. 


44 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


Professor Ansted, however, when systematic teaching ceased to 
be a part of his regular duties, did not abandon his interest in 
geology as an educational subject, but not unfrequently lectured 
and took part in examinations on this and allied sciences. His 
pen also was rarely for long idle. He contributed three papers to 
the Journal of the Geological Society, two of them containing 
descriptions of remarkable mineral veins, published in 1856 and 
1857; his third paper was upon the geology of Malaga and the 
southern part of Andalusia (vol. xv. 1859). Besides these contri- 
butions to the ‘ Quarterly Journal of the Geological Society,’ 
he published memoirs in:—the ‘Transactions of the Cambridge 
Philosophical Society’ (vol. vii. 1842), “On a Portion of the 
Tertiary Formations of Switzerland ;’ in the ‘ Annals and Maga- 
zine of Natural History, “On the Zoological Condition of Chalk 
Flints and the probable Cause of the Deposit of Flinty Strata 
alternating with the Upper Beds of the Cretaceous Formation ” 
(vol. xu. 1844), and other like publications. He was whole or 
part author of a considerable number of volumes, the majority 
of which were of a somewhat popular character ; and more than one 
obtained a considerable share of success. He wrote lucidly 
and pleasantly, whether upon matters directly scientific or upon 
the incidents of travel. Among these works may be mentioned 
the following :—‘ Geological Gossip,’ first published in 1860; ‘A 
Short Trip to Hungary and Transylvania,’ in 1862; ‘ The lonian 
Islands,’ in the year 1863; and ‘ The Great Stone Book of Nature,’ 
published in the same year. He also published a work on Geology, 
in two volumes, in 1844, and more than one smaller book on this 
subject, Physiography, or Geography. He was joint author (with 
. Dr. R. G. Latham) of a work on the Channel Isles, and wrote, 
in 1866, upon the Physical Geography and Geology of Leicester, 
in the ‘ History of Leicestershire,’ entering largely into the litho- 
logy and chemistry of the Charnwood rocks, as well as devoting 
much space to their pure and speculative geology. In the Great 
Exhibitions of 1851 and 1862 he contributed much to the ‘ Reports.’ 
His latest, and one of his most important works, was on Water 
and Water Supply, published in 1878. Though for several years, 
owing to pressure of business and the advance of age, he had 
ceased to take an active part in the proceedings of many of the 
scientific societies of which he was formerly an energetic member, 
he did not lose that cordiality of manner and kindliness of dis- 
position which have left a pleasant impression on the memory of 
his friends. 


In Wittram Hatnowrs Mirier, Professor of Mineralogy in 
the University of Cambridge, our Society has lost a member who, 
though never a contributor to our Journal, was among the most 
eminent mineralogists in Europe, and the author of valuable works 
and papers on that science. He was born April 6th, 1801, at 
Velindre, near Llandovery, in Caermarthenshire, at which place his 
father, Captain Miller, had a few years previously fixed his resi- 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 45 


dence. The associations of the family were military, his father 
having served through a part of the American war, by which he 
had been a heavy loser, and other relatives having held commissions 
and obtained distinction in the British army. W. H. Muller was 
educated at private schools, and afterwards entered as a student 
at St. John’s College, Cambridge, graduating in the year 1826, 
when he was fifth among the Wranglers in the Mathematical Tripos. 
In 1829 he was elected to a Fellowship at his College, and sub- 
sequently proceeded to the degrees, first of M.A., and afterwards 
(in compliance with the statutes by which it was then governed) 
of M.D. ‘For some time he filled the office of a College Tutor ; 
and his first literary work was a ‘Treatise on Hydrostatics,’ published 
in 1831, and followed a few years subsequently by one on Hydro- 
dynamics. At this time the chair of Mineralogy at Cambridge 
was occupied by Dr. Whewell ; and under his guidance Mr. Miller 
devoted himself to the study of Crystallography, with so much 
success that, on the resignation of Dr. Whewell, he was elected to 
the Professorship, the duties of which became the chief work of 
his long and arduous life. In 1830 he was elected a Fellow of our 
Society, and in 1838 a Fellow of the Royal Society. Six years 
afterwards he vacated, by marriage, his Fellowship at St. John’s 
College; but im 1874 he was again elected under the statute 
(granted in 1860) empowering that Society to elect as Fellows 
persons eminent for science or learning, though otherwise techni- 
cally disqualified. Not long after this his health began to fail; 
and in the autumn of 1876 a course of lectures, which he had 
announced, was interrupted by a slight stroke of paralysis. This 
proved the beginning of the end. He was never able to meet his 
class again. Very slowly, but surely, his vital powers declined, a 
torpor stealing alike over body and mind, until at last he fell 
asleep on the 20th May, 1880. 

Professor Miller’s name is inseparably connected with two im- 
portant branches of scientific work. The first of these belongs, as 
might be expected, to mineralogy. “Crystallography,” as has 
been said by Professor Maskelyne, ‘was Miller’s science.” Other 
workers, indeed, had preceded him in laying the foundation and 
contributing important materials; but Professor Miller, ‘“ taking 
the important memoir by Professor Whewell ‘ On the Geometrical 
Treatment of Crystal Forms’ (published in the ‘Transactions of 
the Cambridge Philosophical Society’), and Naumann’s treatise of 
1823 (‘ Beitrage zur Krystallonomie’) as his starting-point, pro- 
ceeded to develop a system of crystallography, which was not pub- 
lished till 1838, but which was the most important work of his 
life.” The especial feature of this book and of his labours in this 
science ‘‘ consisted in working out into a beautiful system the 
indicial method of notation and calculation in crystallography, 
and obtaining expressions adapted for logarithmic calculations by 
processes of great elegance and simplicity. Miller’s system, then, 
gave expressions for working all the problems that a crystal can 
present ; and it gave them in a form that appealed at once to the 


46 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


sense of symmetry and appropriateness of the mathematician.” 
He thus, as it has been well said, ‘“‘ placed the keystone into the 
arch of the science of crystallography ;” and the “future develop- 
ment of that science, there can be little doubt, will follow on the 
lines laid down by Miller.” 

Professor Miller’s shorter communications on mineralogy and 
physics are numerous and valuable, and; in addition to them and 
to his original treatise, he published, in 1863, a tract on crystallo- 
graphy. In 1852 a work appeared entitled a new edition of the 
‘Elementary Introduction to Mineralogy, by the late William 
Phillips, by H. J. Brooke and W.H. Miller. It is, however, no 
_ disparagement to either the original author or his fellow editor to 
say that Professor Miller made this volume almost his own. 

But Professor Miller’s reputation does not rest only upon his 
work as a mineralogist, great though that was. His name is no 
less inseparably connected with the difficult and delicate experiments 
and investigations connected with the restoration of the standards 
of measurement and weight, and with the subsequent labours of 
the International Metric Commission. 

After the fire which, in 1834, consumed the Houses of Parlia- 
ment, it was found that the standards of measurement and weight 


were hopelessly ruined. Professor Miller was not a member of . 


the Commission appointed to consider the questions connected 
with their restoration, but lent the Commissioners much friendly 
assistance. ‘Then, in 1843, a Committee was appointed to super- 
intend the construction of the new Parliamentary standards of 
length and weight, of which Professor Miller was a member ; and 
to him was confided the construction of the new standards of 
weight. In the ‘ Philosophical Transactions ’ for 1856 he describes 
at length ‘the operations for restoring the value of the old 
standard of weight, for constructing the new standard of a 
different value, for constructing various derived standards, and 
for establishing the relative value of the kilogramme,” a paper 
which (to quote the words of the Astronomer Royal, indorsed by 
a former President of the Royal Society, Sir Edward Sabine) “ will 
long be cited as a model of accuracy.” 

He was subsequently a member of a new Royal Commission for 
“examining into and reporting on the state of the secondary 
standards, and for considering every question which could affect 
the primary, secondary, and local standards.” 

In the year 1870 he was appointed a member of the “Commission 
Internationale du Métre.” 

He was appoimted Foreign Secretary of the Royal Society in 
1856, a post for which he was eminently fitted by his accurate and 
extensive knowledge of French, German, and Italian, his methodical 
habits, and unvarying courtesy, as well as by his extensive scientilic 
knowledge. 

He received in 1865 the degree of LL.D. from the University 
of Dublin, and in 1876 that of D.C.L. from Oxford. In 1870 he 
was awarded a Royal Medal by the Royal Society. He was a 


ANNIVERSARY ADDRESS OF THE PRESIDENT. A] 


Knight of the Order of St. Maurice and St. Lazaro of Italy, and 
of the Order of Leopold of Belgium. He was also an honorary 
member of the Royal Society of Edinburgh and of numerous 
foreign societies. 

Three characteristics especially distinguished Professor Miller. 
One was the remarkable extent, depth, and accuracy of his know- 
ledge, not only on those questions which he had made peculiarly 
his own, but also on all branches of physical science, only 
equalled by the liberality with which he opened his rich stores 
of learning to others, and especially to younger students. | Another 
was the simplicity of his disposition. This, combined with a re- 
markable inventiveness of mind, was shown even in his laboratory, 
where the most homely odds and ends were utilized in the con- 
struction of instruments capable of performing delicate measure- 
ments. The third characteristic was the remarkable combination 
which he exhibited of independence of thought and freedom of 
opinion with gentleness of temper and speech, with forbearance, 
courtesy, and respect for the opinion of others. 

Blessed with much domestic happiness, and beloved by all who 
knew him, he lived a tranquil and blameless life, devoted to earnest 
work and the faithful performance of every duty, and passed away 
lamented by a large circle of friends and admirers in this and other 
countries. 


Henry Luptam. We have to record the loss of Mr. Henry 
Ludlam, F.G.S., who died on the 23rd of June, 1880, at the age 
of 58. 

Early a student of the natural sciences, Mineralogy became his fa- 
vourite subject, in the cultivation of which he was greatly assisted by 
the study of chemistry, which occupied many of the leisure ev ening 
hours which the care of an important business allowed him. 
Aided by an unusually critical judgment, he gathered together a 
large and valuable collection of minerals, in which some of the 
specimens were conspicuous for their perfection of crystalline form. 
To these he subsequently added the well-known and important 
collection which belonged to the late Mr. Charles Hamden Turner, 
of Rooksnest, Surrey, and also that made by the late Mr. William 
Nevill, F.G.S., of Godalming. The whole forms certainly the most 
complete and probably the finest collection of minerals ever made 
by a private collector. 

But Mr. Ludlam was nota mere collector. He had long had 
in view the desirability of preparmg a complete descriptive and 
crystallographic catalogue of that which it had been his good 
fortune to accumulate. This great work, to which he looked 
forward as to a labour of love, was actually commenced by him 
shortly before the illness which ended in his premature death. 

Although Mr. Ludlam never contributed a paper to the ‘ Journal 
of the Geological Society, he nevertheless took much interest in 
its proceedings. His great liberality in the support of every thing 
. bearing upon the kindred sciences to geology, and his munificence 


48 PROCEEDINGS OF THE GEOLOGICAL SOCTETY. 


as a collector possessing great knowledge of his subject, entitles 
him to notice in the obituary list of our Fellows. He left by will 
the whole of his superb collection of minerals to the museum in 
Jermyn Street for the use of all students. It is now, therefore, 
the property of the nation. Science, through the early death of 
Mr. Ludlam, has lost an earnest student, while his personal friends 
have to lament the loss of a warm-hearted and true English 
gentleman. 


Ropert CruTrersucK, F.G.S., was the eldest son of Robert 
Clutterbuck, the historian of Hertfordshire. Mr. Clutterbuck was 
educated at Harrow, being head boy of that School in 1817. He 
entered as a commoner at Exeter College, Oxford, where, previous 
to taking his degree, he gained University honours. He then 
entered as a law student under the tuition of the late Sir William 
Hayter. Mr. Clutterbuck spent some time on the continent, gain- 
ing an unusual proficiency in the Italian and French languages. 
During his residence at Watford he acted as a magistrate, and was 
instrumental in raising funds for the restoration of the abbey 
or.cathedral of St. Alban’s. As an author Mr. Clutterbuck 
was well known through his works on the monsoons &c., and on 
the rotatory action of storms; he also published accounts of 
his journeys over the great desert from Aleppo to Bassora, and 
the passage by Suez thr ough Egypt to determine the possibility of 
an overland communication with India. Mr. Clutterbuck never 
contributed to the ‘Journal of the Geological Society ;’ but he 
wrote a paper on the Coprolite beds at Hinsworth, which was 
published in the ‘Transactions of the Watford Natural History 
Society, vol.1. 1878. He died September 15th, 1879. 


Dr. Epwarp Meryon, F.R.C.P., F.G.S., &c., was many years 
Fellow of the Geological Society, and some time member of the 
Council, taking much interest in the working of the Society. He 
never contributed any paper to the Journal. He, however, was the 
author of several important works bearing upon his profession, among 
which may be mentioned especially ‘The History of Medicine,’ ‘The 
Physical and Intellectual Constitution of Man,’ ‘ The Functions of 
theSympathetic System,’ and ‘ Practical and Pathological Researches 
on the various Forms of Paralysis. Dr. Meryon was an accom- 
plished and practical physician of great professional experience. 
He was an accomplished scholar and true-hearted gentleman. Dr. 
Meryon died November 8, 1880. 


In Eriszan Watton, who died at his house near Bromsgrove on 
August 25, 1880, in his forty-eighth year, the Society has lost an 
artist who has been equalled by few, perhaps surpassed by none, 
in his power of rendering faithfully the forms of mountains and 
the structure of rocks. Huis studies of clouds and of the camel 
prove the versatility of his genius and the fidelity of his execution ; 
but it is in his pictures of the mountains of Sinai, Norway, and 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 49 


the Alps that his excellence in rock-drawing of every kind is con- 
spicuous. Overwork and anxiety first clouded and then terminated 
a life which once seemed full of brilliant promise. 


Dr. WitnEetM Purripe Scurmerer, Professor of Geology in the 
Faculty of Sciences, Director of the Museum of Natural History, 
and Professor in the University of Strasburg, a Corresponding Mem- 
ber of the French Academy of Sciences and of the Academies of 
Munich, Lisbon, and Philadelphia, elected a Foreign Correspon- 
dent of the Geological Society in 1874, died of disease of the heart 
on the 20th March, 1880. 

Dr. Schimper was descended from a family living in the 
Palatinate, the connexion of which with Alsace was at all times 
close, and especially so during the first French empire. His 
father at this time became the Protestant pastor of Dossenheim, 
near Saverne, in Alsace; and here Schimper was born in 1808. 
He was educated at first by his father, and from an early period 
showed a great taste for natural history, which continued to 
manifest itself after he had temporarily left the paternal roof, at 
the age of 14, to pursue his studies at the College of Bouxwiller. 
Here he remained until 1826, when he went to Strasburg, and 
there, in 1833, took his degree as Bachelor of Divinity. He sub- 
sequently acted for a time as curate ‘to his father, who was then 
pastor at Offwiller. 

_ he young Schimper, however, seems to have devoted his atten- 

tion more to Natural History, especially Botany, than to theologi- 
cal studies ; and he had acquired such a knowledge of the Mosses, 
that as early as 1834 the eminent botanist Bruch proposed to 
him that they should produce a joint monograph on the European 
forms of that class of plants. Schimper acceded to the proposal, 
and, with the view of devoting himself exclusively to natural- 
history studies, resigned his ecclesiastical functions, not without 
opposition on his father’s part, and in 1835 accepted the position 
of preparator in the Museum of Natural History at Strasburg. 
With that establishment he remained connected until the day of 
his death, having been reinstated in his various offices by the Ger- 
man authorities after their annexation of Alsace. 

The work on European Mosses above mentioned was pro- 
duced by Schimper alone. It is entitled ‘ Bryologia Huropzea, seu 
genera Muscorum Huropeorum, Monographia illustrata,’ and is a 
monument of the author’s industry and botanical attainments, 
extending to six 4to volumes, with an atlas of 641 plates. Its 
publication was commenced in 1836, but was not completed until 
nearly twenty years afterwards (namely, in 1855). The author's 
bryological publications did not cease then, however: he afterwards 
produced several supplements to his great work, besides other 
memoirs on Mosses, and a ‘Synopsis Muscorum Europeorum,’ of 
which the second edition appeared in 1874. 

In procuring materials for the above work, Schimper travelled a 
good deal in various parts of Hurope; and on his journeys he was in- 


50 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


defatigable in collecting specimens of all kinds for the enrichment of 
the museum under his charge. He acquired a very wide knowledge 
of general natural history, and paid much attention to geological 
phenomena, which, in conjunction with the direction of his special 
botanical studies, early led him to the investigation of fossil plants, 
upon which he was destined to become one of the chief authorities 
of our day. He associated himself with Kochlin-Schlumberger in 
the preparation of a great memoir upon the Terrain de Transition 
des Vosges, published at Strasburg in 1862; to this Schimper con- 
tributed the monograph of the fossil plants. 

In 1849 he married a Swiss lady who had been long an ardent 
student of botany, and who afterwards assisted him in his studies, 
and especially in the accumulation of the vast stores of materials 
upon which he founded his greatest work, the ‘ Traité de Paléon- 
tologie Végétale,’ published in three large 8vo volumes, with a 4to 
atlas of plates, of which the first volume appeared in 1869. This 
important treatise upon the fossil flora will long be a standard 
work of reference. Subsequently Schimper commenced the 
botanical section of Prof. Zittel’s admirable ‘ Handbuch der Pali- 
ontologie,’ which, however, he did not live to complete. 


JosrpH Aveustin Husrerr Bosqunr, of Maestricht, Doctor of 
Sciences, Pharmacien, Member of the Royal Academy of Sciences 
of Amsterdam, was elected a foreign Correspondent of the Geological 
Society in 1864. In 1868 the Council awarded the balance of the 
proceeds of the Wollaston Donation Fund to Dr. Hubert Bosquet 
as a reward for his valuable researches on the Tertiary and Cre- 
taceous strata of Holland and Belgium. Dr. Bosquet worked out 
with great industry and ability the fauna of the Maestricht beds, 
a division of the Upper Chalk not present in England. His re- 
searches amongst the Tongrian beds greatly extended our know- 
ledge of these Tertiary deposits, both as regards their paleontology 
and physical conditions. Bosquet’s labours were undertaken and 
carried on amidst business duties, and with zeal and ability rarely 
excelled. He was the author of twelve or fourteen contributions 
both to geological and paleontological science. His chief papers 
are :—‘¢ Description des Entomostracés fossiles de la craie de Mae- 
stricht” (Mém. Soe. Sei. Liege, vol. vi. 1847); and “ Description 
des Entomostracés fossiles des terrains tertiaires de la France 
et de la Belgique” (Mém. Couronn. Bruxelles, xxiv. 1850-51). 
His papers entitled “ Les Crustaecés fossiles du terrain Crétacé 
du Limbourg” (Nederland. Geol. Kaart Verhand. 11. 1854) and 
‘¢ Recherches Paléontologiques sur les terrains tertiaires du Lim- 
bourg Néerlandais ” (Amsterdam, Verhand. vii. 1859), are of much 
importance. Dr.Bosquet died on the 28th of June, 1880, aged 
66 years and 10 months. 


Prerre Henri Nyst, Conservator at the Museum of Natural 
History at Brussels, was elected a Foreign Correspondent of 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 51 


this Society in 1863, and in 1871 was elected a Foreign Member. 
He was a Member of the Royal Academy of Sciences of Belgium, 
and was also a Chevalier of the order of Leopold. 

In 1874 the Council awarded him the balance of the proceeds ot 
the Wollaston Donation Fund for his admirable, extensive, and 
original researches upon the Crag Mollusca of Belgium. His chief 
work is the ‘ Description des Coquilles et des Polypiers fossiles 
des terrains tertiaires de la Belgique’ (4to, Brussels, 1843). Up 
to the year 1873, Nyst had written about thirty-six papers, mostly 
upon recent and fossil concholog ey. His connexion with the national 
museum at Brussels, and obliging manner, rendered him at all times 
accessible to students or those desiring information upon Tertiary 
paleontology. Nyst died on April 6th, 1880, aged 67 years. 


On tHe ANALYSIS AND DIsTRIBUTION OF THE British PALmozOIC 
FossItzs. 


Tux history of so progressive and practical a science as Geology, 
and its kindred study Paleontology, especially during the past 
decade, not only in Britain, but in Europe, America, and our Colo- 
nies, now needs some revision and analysis, arising from the 
amount of research, and the progress that has been made in these 
and collateral sciences during this period. 

The history of the Lower Paleozoic rocks of the British Islands 
is almost the history of Geology; for it extends over forty years. 
The same may be said of Scandinavia and Bohemia. And parallel 
to these researches in Kurope may be recorded the great progress 
made in the western hemisphere; for the history of the Lower 
Paleozoic or oldest rocks of N. America and Canada is but that of 
Britain and Northern Europe almost repeated ; homotaxially they 
are the same. 

The study through many years of the distribution of life through 
the stratified rocks of the British Islands enables me to lay before 
the Society some results arrived at by long and patient research. 
I have, however, only taken advantage of my position on the 
Geological Survey, which has afforded me facilities otherwise 
almost impossible of access, and enabled me to carry on practically, 
both in the field and the study, those branches of our science which 
bear so intimately upon the progress of Physical Geology and Geo- 
graphy, besides elucidating some of the laws that have governed 
the distribution of life through time and space. 

De la Beche, Lyell, Edward Forbes, Hamilton, Phillips, Huxley, 
Ramsay, Prestwich, and Duncan, in their learned addresses delivered 
from this chair, have one and all dealt with questions bearing 


52 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


especially upon stratigraphical geology as based upon the range and 
distribution of organic remains, phenomena which since the time of 
William Smith have been studiously kept in mind, teaching us the 
succession of life and the relation in time of one formation to 
another—each successive sedimentary group containing undoubted 
records of its life and deposition, thus rendering clear and definite the 
changes in, and the advance of life through time. 

No more difficult problem exists cr remains to be solved than the 
first appearance or commencement of life on the globe. Is the Lau- 
rentian of North America, with its one known solitary form of life, 
the oldest sedimentary rock existing? is Hozoon canadense the oldest 
form of life? Research up to the present time has not revealed to us 
one of higher antiquity ; neither has it even a truly associated form. 
No true Annelide, Plant, or Protozoon accompanies this still mys- 
terious progenitor of Paleozoic life. 

The Hozoic or Laurentian gneiss of Britain or Europe, has not yet 
yielded a semblance of any thing approaching Protozoic affinities, 
although in the Hebrides, Norway, Sweden, Bohemia, and Bavaria 
the Laurentian rocks have been recognized. 

That the Laurentian rocks are not the oldest is manifest ; others 
of infinitely greater age yielded sediment to the Laurentian sea, 
and pabulum for the sustenance and material for the shelly structure 
of its supposed only inhabitant ; but whether Hozoon had precursors 
or not, time will probably tell. 

Dr. Dawson, however, suggests that vegetable life preceded 
Eozoon, and may thus have accumulated previous stores of organic 
matter. If any older forms of animal life did exist, they cannot 
have belonged to much simpler types; “naked Protozoa would 
have left no sign of their existence, except probably minute traces 
of carbonaceous matter.” Dr. Dawson in the year 1865 discussed 
the question of associated organic structures, and what share, if 
any, they may have had in the accumulation of the Laurentian 
Limestone. Microscopic examination exhibited evidence of calca- 
reous and carbonaceous fragments of organic origin. The contents 
of the organic limestone, as shown by Dr. Dawson, were “‘ Remains 
of Kozoon, other calcareous bodies probably organic, objects im- 
bedded in the serpentine, carbonaceous matters, perforations or 
worm-burrows.” Dr. Dawson strongly and fairly argues for asso- 
ciated life. 

The presence of graphite in large deposits occurring both in 
beds and veins in the Laurentian rocks, clearly determines that its 
origin and deposition were contemporaneous with the mass or con- 
taining rock; the graphite, again, 1s associated with calcite, quartz, 
and orthoclase. It is not improbable that the ‘ vein graphite ” was 
introduced as a liquid hydrocarbon. Dr. Sterry Hunt believes it 
possible that it may have been produced in a state of aqueous solu- 
tion*. In the lower Laurentians the quantity is enormous. Dr. 
Hunt also believes that the origin of the graphite was due to the 
deoxidation of carbonic acid by living plants. That the graphitic 


* Hunt, Report of the Geological Survey of Canada, 1866. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 53 


matter of the Laurentian rocks was laid down or accumulated in 
beds like coal is improbable, no evidence whatever tending to show 
that there existed terrestrial vegetation. On the other hand, the 
hydrocarbon may have been due to diffused bituminous matter 
closely resembling “‘ our bituminous shales and bituminous and oil- 
bearing limestones.” Research hitherto has failed to find traces of 
any organism save the Hozoon ; no cryptogam has yet occurred ; and 
we cannot imagine that, if of vegetable origin, the organic matter 
could have been so completely disintegrated and bituminized prior 
to being changed into graphite. Dr. 8. Hunt believes he has de- 
termined the presence of terrestrial vegetation in the great beds of 
Laurentian iron-ore which show subaerial decay, thus implying the 
‘reducing and solvent action of substances produced in the decay 
of plants.” Dr. Dawson long ago (1875) believed that he had found, 
in the compact graphitic limestones of Clarendon, traces of fibrous 
structure due to segregation which may be the remains of plants, 
‘and in some specimens vermicular lines ” which he believed to be 
“‘ Hozoon penetrated by matter once bituminous, but now in the 
state of graphite.” At the utmost we can only speculate upon the 
presence or condition of vegetation during the Archeean, or Pre- 
Cambrian, Laurentian, or Eozoic time. 

Dana employs the term Archzean in time, to express in full mean- 
ing that era in the physical development of the Earth which was 
‘“‘incompatible with the existence of life,” when life was not, so far as 
we know. Little, however, is known of that vast group of rocks 
we call Pre-Cambrian, adopting this term as used in this country, 
on the Continent, and in America, for rocks affording us no history 
or recognizable records. 

Dana again assumes and endeavours to show that four eras pre- 
ceded the Laurentian period :— 

A first or molten era subsequent to that of the presumed original 
nebulous state. 

A second era, one of solidification and consolidation through 
cooling, when the Earth became solid at the centre; later on, at- 
mospheric vapours became condensed and probably universal ; water 
covered all. The cooling and contracting of the sphere resulted in 
oceanic depressions in special areas, and our continents were sha- 
dowed forth and contoured, but as yet no life can be chronicled. 

The third era may have given us surface reliefs; stratified deposits 
were formed. 

The fourth era probably saw the beginning of life, which occurred 
when the oceanic waters may have stood at 200° F. With this 
fourth era we may associate the Hozoonal serpentines. 

In the British Islands we have yet to find that type of Laurentian 
rock which yields either of the great limestones of Canada,— 
the so-called Fundamental Gneiss of Scotland, Malvern, and Ire- 
land having no affinities with the Eozoonal and graphitic group 
of the North-American continent, and being doubtless of vastly 
younger date. 

To summarize the results of the labours and views of those who 
VOL. XXXVII. ie 


54 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


have carefully investigated and thought out any great physical 
question is at all times difficult ; and perhaps there are few subjects 
on which men have theorized and differed more than on the history of 
the globe. For time and life are two subjects that at once arrest the 
attention of all earnest students. Our knowledge of the commence- 
ment of either is as indefinite now as in the days of the earliest 
investigators. With the succession of sedimentary rock-masses 
in the outer framework of the globe we are perhaps partly familiar, 
certainly so for given and known areas; there are, however, exten- 
sive regions yet unknown and unexplored, and remaining to be corre- 
lated with the known; and yearly in the Transactions of our Society, 
through the researches of our Foreign Members and others, are we 
reminded how insecure and uncertain is our base, how doubtful our 
succession, when attempted to be universally applied. Yearly some 
new light is thrown upon the obscure history of the earliest rocks with 
which we believe ourselves acquainted. Both theearly metamorphic 
and the lowest Paleozoic rocks, even in our own small area, are 
still waiting for final position and classification; the same may be 
said of much of Europe, America, Canada, India, and Australia. 

Paleozoic time in Europe and Britain may have commenced with 
the deposition of the so-called earliest Cambrian rocks; but where 
geographically, we know not—probably far to the west of Ireland 
and the British Islands, under what.is now the deep Atlantic. The 
north-western coast of Scotland, and Ireland, much of the north-west 
of England, and North and South Wales all point to a region where 
we may believe that the earliest known sedimentary rocks of Western 
Europe, and their life contents had their commencement or origin. 
Little can be said here of Archean time and its rocks as developed 
on the American continent; possibly we may recognize and correlate 
the Archean system of Dana with our gneissose schists and so-called 
Laurentian rocks of the north-west coast of Scotland and the 
Hebrides. 

It would be mere speculation here to attempt to define any strict 
contemporaneity. We know from the labours of Dana, and the re- 
searches of Sir William Logan in the field, and of Billings in the 
study, that two periods or eras of Archean time are fully represented 
in North America and Canada:—Ist, the older or Laurentian; 
2nd, the Huronan, this latter in all probability represented by 
our lowest Cambrian, or those beds underlying the Menevian of St. 
David’s, and also constituting the rocks of Harlech, Llanberris, Ban- 
gor, and the Longmynds. We have hitherto discovered no life-remains 
in the Archean rocks of the British Islands, our so-called Lauren- 
tians having none of the prevalent limestones, Eozoonal or otherwise, 
of the New-York and Canadian rocks. To trace out the conditions 
of the northern hemisphere during Lower-Cambrian times, both 
zoologically and physically, has been and is still one of the most 
difficult and important problems of modern geological research*. 
We have long known that the. Huropean area was of great extent, 


* Vide Hicks, Geol. Mag. dec. 2, vol. iii. p. 876, ‘‘ On the probable Conditions 
under which the Paleozoic Rocks were deposited over the Northern Hemisphere. ’ 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 55 


even before the Cambrian rocks were accumulated : remnants of this 
early land or continent are still visible and traceable in Spain, 
France, Scandinavia, Ireland, Scotland, England, and Wales. Gneiss, 
eranitoid rocks, and occasional limestones formed the mass of this: 
Pre-Cambrian land, now exhibited to us in a greatly metamorphosed 
state, stratigraphically unconformable to the overlying Cambrian, 
and with a discordant strike north of latitude 380°. These rocks are 
visible on both sides of the Atlantic as far as the Arctic regions, 
representing portions of two continents, Hurope and America, 
now separated by the North Atlantic. 


Pre-Camprian Rocks. 


it may be received as an axiom that all the known older sedi- 
mentary rocks were deposited more or less in a similar manner, 
or under similar conditions, to those of modern times. Allowing 
this to be the case, it is most probable that those sedimentary 
deposits which are the lowest, or have the earliest position, have 
undergone most change; and it must be admitted that the erystal- 
line, semicrystalline, and metamorphosed state in which they now 
appear has been subsequently induced through various agencies 
(beat, pressure, and chemical change) exerted through countless 
ages. Research seems to prove that most of the rocks now recog- 
nized as Pre-Cambrian were originally sedimentary strata, which 
have undergone, since their deposition, alteration or metamorphism. 
Few men have paid more attention to the physical history, distri- 
bution, succession, and character of this most ancient group of rocks 
than Dr. Hicks, ably followed by Professor Bonney, Professor Hughes, 
Mr. Tawney, Dr. Sorby, Professor Hull, and Mr. Hudleston ; each of 
these observers has contributed largely to the elucidation of the 
history of the Cambrian rocks in his own particular way. Three 
if not four groups or systems have been determined by Hicks, having 
definite bearings or discordant strikes one to the other through 
certain portions of the British Islands. They occur in ascending 
order, and in the order of time, as follows :— 


1. Lewisian. Hebrides and North-west Highlands. 
2. Dimetian. St. David’s, Caernarvon, and Anglesey. 
3, Arvonian. Pembrokeshire, Caernarvonshire, Anglesey, Shropshire, 
and the Harlech Mountains. 
4. Pebidian. South Wales, Shropshire, Charnwood Forest, Anglesey, 
and Ireland. 


Lrewistan.—Murchison gave the name Lewisian to the crystal- 
line rocks largely entering into the structure of the Hebrides and 
North-western Highlands; probably these constitute the oldest group 
of rocks known or recognized in the British Islands. Red felspar, 
hornblende, and quartz are the prevailing ingredients in these mas- 
sive gneisses ; occasionally, as at Cape Wrath and on the coast to the 
south, almost pure hornblende rock occurs. The strike of the 
Lewisian group is usually east and west, or ranging between that 
and north-west and south-east. 

ee 


56 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


The Malvern chain may also represent the Lewisian ; and also the 
highlands of North Ireland as developed in Donegal. 

Dimer1an*.—This group is extensively developed in many parts of 
South and North Wales ; it oceurs in Shropshire, and, according to Dr. 
Hicks, is known in the Worcester Beacon of the Malvern chain; it is 
also said by Hicks to occur at Ben Fyn, Loch Maree, and near Gaer- 
loch in Ross-shire, and in other areas in the N.W. Highlands. 

Unlike the Lewisian type, the Dimetian contains impure chloritic 
limestones and serpentinous bands (not of organic origin so far as we 
know). ‘The chief minerals are flesh-coloured and white felspar, and 
chloritic and hornblendic bands ; and the rocks are highly quartzose. 
Fragments of the older Lewisian gneiss are occasionally found im- 
bedded in the gneiss, thus determining the unconformability of the 
Dimetian group, through derivation and denudation. Viridite occurs 
more or less throughout, giving a tinge to the rock. At St. David’s 
the ridge formed by these rocks, according to Dr. Hicks, averages from 
2000 to 8000 feet in thickness. ‘‘ Cambrian conglomerates rest im- 
mediately upon the Dimetian rocks ; and the highest members of the 
Harlech group strike up against the Dimetian ridge;” and both 
sides are flanked throughout by the uncomformable Pebidian group. 
At St. David’s the rocks chiefly composing the Dimetian series are 
compact crystalline quartz, chloritic schists, and indurated shales ; 
the quartz in places occurs almost perfectly crystalline. Massive 
beds of calcareous shale and dolomitic limestone occur. “ As muchas 
20 per cent. of carbonates have been determined on analysis by 
Mr. Hudleston, and in addition 0°5 per cent. of phosphoric acid ; 
hence it is more than probable that the lime was deposited by 
organic aid” (Hicks). The Pebidian rocks are strongly contrasted 
by their bedded and shaly character. Dr. Hicks has not discovered 
any traces of organic life in the inferior limestones, of which five 
beds occur. 

Dr. Hicks estimates the thickness of the St. David’s Dimetian at 
15,000 feet. Several intrusive dykes traverse the series ; and their 
injection took place before metamorphism had commenced. They are 
fine-grained altered dolerites, columnar in structure, with the columns 
at right angles to the plane of the dykes. 

Arvontan.—In 1878 Dr. Hicks discovered in Pembrokeshire new 
areas of Pre-Cambrian rocks of a totally different character from 
the Dimetian and Pebidian which he had previously described. 
The rocks of this Arvonian group are marked on the Geological 
Survey map as intrusive felstones; they occur in three or four 
isolated masses of considerable extent. Their strike is from north 
to south, and discordant to those of the newer rocks and the under- 
lying Dimetian. 

The wild mountain-region of Plymstone is mainly composed of 
the Arvonian group. Dr. Hicks finds Lower Cambrian and Lingula- 
flag rocks resting unconformably upon the “ Arvonian” along their 


* «Dimetia,” the ancient name for a kingdom which included this part of 
Wales. Vide Hicks, “On the Pre-Cambrian (Dimetian and Pebidian) Rocks of 
St. David’s,” Quart. Journ. Geol. Soc. vol. xxxiti. p. 229 (1877). 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 57 


N.W. edge. The texture of this new group is hard and flinty, 
some portions having an imperfect conchoidal fracture. The chicf 
character or peculiarity consists in the manner in which the quartz 
is separated or segregated into nests, giving the rock a pseudopor- 
phyritic appearance. 

The Arvonians, both generally and microscopically, closely re- 
semble the Swedish ‘‘ Hilleflintas ;” nevertheless associated with 
them are true quartz-felsites, probably old lava-flows (rhyolitic). 
The evidence as to the position which the Arvonian occupies in the 
Pre-Cambrian group is conclusive, as determined by the way in 
which the Cambrian rocks are faulted down against it. It is cer- 
tainly Pre-Pebidian, as fragments of the Hiilleflinta occur in the 
overlying Pebidian conglomerates; its rocks occupy a large pro- 
portion of the Dimetian axis of St. David’s; and, according to the 
researches of Hicks, that part coloured as syenite and felstone in 
the Geological map to the N.E. of St. David’s must be assigned to 
the “‘ Arvonian” group, only the lower portion being Dimetian. 

Like the Dimetian these rocks are greatly altered, consisting of 
highly metamorphosed indurated porcellanitic shales. The beds 
resting upon the Dimetian axis are hard compact conglomerates, 
composed of quartz and altered shale derived from the Dimetian. 

Prprp1an*.—The Pebidian rocks are unconformable to the Dime- 
tian, extending along both sides of the Dimetian ridge. They 
strike from §8.W. to N.E., nearly parallel to the ridge, or in 
accordance with the overlying Cambrian Rocks; they are irregular 
in thickness and greatly metamorphosed, composed of porcellanitic 
shales; “the narrow dark lines” of stratification and closely 
approximating and intersecting joints distinguish the Pebidians 
from the Dimetian, against which they rest. ‘The “lower beds are 
hard compact conglomerates, composed of masses of quartz and 
altered shales,” derived from the underlying rocks and closely con- 
nected together. ‘These all immediately lie on the Dimetian axis. 
They are to be seen at Nun’s Well, south of St. David’s, on both 
sides of the Caerbuddy valley to the east of the city, and north and 
south of the cathedral in the valley. This Pebidian group supports 
the true Cambrian rocks at many places in the neighbourhood. 
Little more than 3000 feet are exposed; and the strike is nearly 
identical with that of the overlying Cambrian. The large area 
coloured as intrusive greenstone upon the Geological Survey map, and 
extending parallel to Ramsey Sound, is composed of the Pebidians. 
A considerable portion of the 8.W. part of Ramsey Island consists of 
the compact porcellanites which characterize this series. The 
Harlech conglomerates rest unconformably upon or along the N.K. 
edge of the Pebidianst. Basic lavas and breccias predominate over 
the Rhyolites. 

It is stated by Professor Ramsay, in the ‘ Geology of North Wales’ 


* So named from ‘ Pebidiauc,” the name of the division or hundred where 
these rocks are exposed near St. David’s. . 

+ See Dr. Hicks, Quart. Journ. Geol. Soc. vol. xxxiii. pp. 229--239, for map 
and sections &e. 


55 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


(Mem. Geol. Surv. vol. iii. p. 8), that the oldest rocks in Wales and 
Shropshire appear at the surface in six districts—elevation, denuda- 
tion, and relative changes haying caused to be exposed as six inland 
islands as many masses, chiefly mountainous, and composed of the 
oldest sedimentary strata, in Britain, whose true history has yet 
to be written. In Ireland we may recognize three or four such 
areas also. 

These and others, of the history of whose masses we are still 
ignorant, constitute the basis and foundation (mostly unfossilife- 
rops) of all succeeding sedimentary strata. They have been of late 
and still are being subjected to critical examination, both chemically 
and microscopically. Large areas hitherto believed to be of igneous 
origin, and long ago mapped as such during the progress of the 
Geological Survey, are now found under rigorous examination to 
be sedimentary rocks which have undergone complete metamor- 
phosis since deposition. Such determinations are all-important 
when we come to consider the earliest appearance of life and its 
distribution within the British area; for recent investigation has 
resulted in the removal of certain rocks, hitherto believed to be 
igneous, to the sedimentary or stratified series. This has been the 
case, as we have seen, with several important masses in North and 
South Wales, now designated Pre-Cambrian, assuming them to be 
a group of rocks of higher antiquity than those in which undoubted 
traces of life have hitherto been found in Britain. 

To the influence of microscopic investigation and research is due 
the right determination and history of these and other doubtful 
rocks. Dr. Hicks has described some areas in the Lleyn peninsula 
along the N.W. shore which he believes to be representative of his 
Dimetian, Arvonian, and Pebidian groups; and Professor Bonney 
doubts not, from microscopical examination, the presence here of a 
considerable Pre-Cambrian series, with at least two very distinct 
groups of rocks. 

Dr. Hicks during the past summer determined the presence of 
an older series of rocks than the Cambrian of the Harlech or 
Merioneth anticlinal near the centre of the Harlech Dome. These 
rocks evidently underlie the Harlech sandstones, and constitute 
part of a pre-existing formation. The Cambrian conglomerates at the 
base of the Harlech grits contain fragments of rock identical with 
- this older formation. 

No discovery of late has equalled this in importance ; it has proved 
the existence in North Wales of a group or system hitherto un- 
known, although expected or anticipated. This enables us to compare 
the thickness of the Cambrian rocks of North Wales with that of 
those of the same age at St. David’s in South Wales, and at the same 
time to realize and compare the physical conditions of thetwo. This 
may enable us to measure the thickness of the Harlech Cambrians, 
and probably to arrive at the strike of the subjacent rocks. The 
broken and denuded anticlinal has exposed these Pre-Cambrian 
rocks, north and south ; they are marked in the maps of the Survey 
as intrusive felstones, but appear to be a highly metamorphic 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 59 


series of schists, alternating with bands of purplish slate, and 
are believed by Dr. Hicks to be of Pebidian age: the strike is from 
N.E. to S.W. 

Between Caen Cochion and Pen-maen east of Traws-fynnyld road, 
the Cambrian conglomerates are seen resting unconformably upon 
the older series; and masses of the Pre-Cambrian occur plentifully 
in the conglomerate. It is to be hoped that Dr. Hicks will continue 
his researches over this wild and difficult region; it is highly im- 
portant to know the precise nature of the axis and underlying rocks 
of the anticlinal. The researches of Professor Ramsay and the 
Survey detail all that we really know of the great mass of the 
Harlech Dome or Merioneth anticlinal*. Caernaryonshire, Angle- 
sey, Shropshire, the Malvern Hills, and Scotland have received critical 
examination from Dr. Hicks; but his results are too full of detail to 
be entered upon here. 

The physical structure and life-history of the classical promon- 
tory of St. David’s has of late years, at the hands of Salter and 
Hicks, received an amount of research and attention of which 
scarcely any other district can boast. 

The rocks composing the ancient headland of Menapie are, 
perhaps, the oldest known rocks containing organic remains in the 
British Islands, if not in the world; nevertheless the characters 
presented by the contained organisms are such that we cannot 
imagine that we have here traces of the earliest manifestations of 
life upon the earth. 

In treating of the various successive formations embraced in the 
recognized Paleozoic divisions of the British sedimentary rocks, 
1 shall omit altogether any definite classification into great systems. 
For while we may define one period as characterized by the 
presénce of a certain fauna, which in the next succeeding so- 
called epoch, is replaced by a different one, there will always be 
found in some part of their geographical distribution a region 
where, in some form or condition, the two faunas commingle, 
and where the old one gradually disappears as the new one comes 
in or makes its appearance. To assign, therefore, any definite or 
precise boundary-lines or limitations to our stratified rocks and their 
contents, when the record of past life is of necessity so incomplete 
and obscure, is, to say the least, at present premature, if not unphi- 
losophical, however convenient it may to a certain extent be; but 
the progress of so boundless and progressive a science as Geology 
with its associated subjects, demands that we should lay down no 
hard or definite lines, no brackets or definitely constructed tables. 

Each year witnesses the breaking-down of arbitrary divisional 
lines in classification, in Zoology and Pale econtology as well as in 
Geology. Any attempt, therefore, to establish geological divisions 
or horizons upon either stratigraphical or paleontological breaks 
must be temporary and local only; probably there exists no 
break in life, any more than in time or in sedimentation; for some- 


* Vide Ramsay, Mem. Geol. Survey of Great Britain, vol. iii. Geol. of North 
Wales, pp. 17-19. 


60 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


where there will be found continuous and conformable succession 
both physically and paleeontologically. We have no visible chrono- 
logical scale in Geology but such as is afforded by the relative 
magnitude of zoological changes; in other words, the geological 
duration and importance of any system is in strict proportion to the 
comparative magnitude and distinctness of its collective fauna. 

It would be out of place here to enter upon the question which 
has been long at issue as to the claims of two, if not three, schools 
of thought and research in relation to the divisions or classification 
of the Lower Palzeozoie rocks. 

“¢ To Professor Sedgwick must be conceded the credit of determining 
and assigning the limits and sequence of the larger subdivisions; for ~ 
single-handed he laid down with masterly precision the succession 
and true stratigraphical arrangement of the Lower Paleozoic rocks 
of Wales, from the Bangor beds to the summit of the Bala group.” 

‘To Sir R. Murchison must also be awarded the high credit of 
having been the first to distinctively assign to many of these rocks 
their sequence, distinctive fossils, and palecontological value”*. Con- 
flicting views led to the formation of a third school of research, in 
which Sir Charles Lyell, Mr. Salter, and Dr. Hicks exercised great 
influence. These last two observers assigned the term Silurian to 
all the strata ranging from the top of the Ludlow to the base of the 
Arenig, and restricted the term Cambrian to all between the base 
of the Lower Arenig and the lowest known beds of the Harlech and 
Llanberris group. The Tremadoc group, by its fossils, however, has 
hittle in common with the underlying Lingula-flags and Menevians, 
only sixteen of the Lingula-flag and other forms, out of one hundred 
and eighty-two that range below the Tremadoc passing up to the 
latter horizon. The great and almost total break at the top of the 
Tremadoe lends strong evidence in favour of the division. being recog- 
nized here. 

The recognition of a tripartite grouping of the faunas and 
strata, between the base of the Old Red Sandstone and the 
Harlech series, cannot be disputed; each is characteristic, and 
possesses a broadly-mark i ial Cam- 
brian, or first fauna, the “Ordovician” system, or second fauna, 
and the Silurian system, or third fauna, according to their succession 
in time. Sedgwick named his system after, the entire principality 
in which his rocks were typically developed, his. title of Cambrian 
being comprehensive enough to embrace the whole of the Palecozoic 
rocks. Murchison selected the term Silurian, associating the rocks 
of his system with the tribe of the Silures. 

Mr. C. Lapworth, in his able paper upon the tripartite division of 
the Lower Palmozoic rocks, has suggested, and with good reason, 
the name “Ordovician” for all that croup of strata in the Great 
Bala district called Upper Cambrian of Sedgwick or Lower Silurian 
of Murchison—from the ancient tribe of the Ordovices, who occupied 
the country now called Montgomeryshire, Merionethshire, Caernar- 
vonshire, Denbighshire, and Flintshire. So long, he remarks, as the 

* Geol. Mag. decade 2, vol. vi. pp. 1-15, 1879. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 61 


present system of nomenclature exists nothing can disturb the 
application. of the title Cambrian to the rocks of the primordial series, 
and of Silurian to the strata of the third fauna, or that series of 
strata intervening between the Old Red Sandstone and the Lower 
Llandovery. With the intermediate series from the Llandovery to the 
Lower Arenig ‘‘ we have had incessant differences respecting nomen- 
clature, proper limits of the groups, and sequence of fossils; they 
are still designated as Upper Bala, Caradoc, Middle Cambrian, and 
Lower Silurian. This central system of the Lower Paleozoic may 
therefore well receive a name equally euphonious and significant of 
the area where its strata are typically developed ” *. 

No division of the British sedimentary rocks has given rise to so 
much controversy as the so-called Cambrian strata. At the present 
moment they have no fixed or definitely-assigned horizon, either 
base or summit, beginning or end; their time-history, necessa- 
rily their space-development also, rests upon no agreed or recog- 
nized determination. ‘Two, if not three, schools of research in these 
lowest groups of rocks differ even among themselves as to the 
uppermost limit that should be assigned to the term Cambrian, or 
where in the field the line of demarcation should be drawn between 
it and the Silurian, or at the base between it and the Pre-Cambrian 
or metamorphic series. It is no part of my duty to enter ito 
details relative to the history of the controversy which has so long 
occupied the minds and attention of the respective advocates of the 
schools of Sedgwick and Murchison. 

The past ten years have witnessed great changes both in the 
nomenclature and classification of the Cambrian rocks. The name 
Cambrian, given by Sedgwick in 1838 to the whole group of strata 
below the May-Hill Sandstone, has of late years again given rise 
to much controversy. This critical research has been equally 
important in its bearing upon the investigation of European and 
American geology and the establishment of a corresponding no- 
menclature for the succession and history of these oldest known 
fossiliferous rocks on the globe. Whether we can rightly refer 
the Dimetian, Arvonian, and Pebidian groups of Britain to the 
Laurentian and Huronian of Canada is yet a question; that they 
occur below the recognized Cambrian, there can be no doubt, both 
from discordancy of strike, petrological differences, and supposed 
total absence of organic remains. Probably the great groups of rocks 
comprised under the above names (Laurentian and Huronian), the 
Archean of Dana, may be the equivalents in time of our St.-David’s 
and North-Wales Pre-Cambrian; and his Primordial or Cambrian 
system (embracing the two series Acadian and Potsdam) may repre- 
sent the lower portion of our fossiliferous Cambrian rocks, or those 
so well defined and developed at St. David’s, viz. the red shales and 
flaggy beds. with Lingulella ferruginea and L. primeva, the red, 
purple, and grey grits, the Plutonta-beds, the red, grey, and purple 
flagey sandstones, and the succeeding grey flaggy series, all five of 
which are fossiliferous. Not only so, but these Pembrokeshire Cam- 

* Loc. cit. p. 14. 


62 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


brian rocks seem to have no equivalents in North Wales, unless they 
represent in time the Llanberris, Harlech, and Bangor sandstones, 
grits, and flags, which, however, have as yet yielded no trace of organic 
remains. Again, very few species are common to these grey, red, and 
purple sandstones and the succeeding well-developed Menevian 
group, which has yielded nearly fifty species* ; yet in the Harlech, 
Barmouth, and Dolgelly areas the Menevians immediately succeed, 
are conformable to the underlying barren unfossiliferous grits, and 
exhibit the first traces in North Wales of a definite and defi- 
nable fauna. In other words, the representatives of the Harlech 
and Llanberris beds in the St.-David’s promontory contain a 
well-marked fauna, highly characterized by the Crustacea (7 
genera and 14 species) and by the still more important fact 
that no species occurring in the Longmynd and Harlech rocks of 
St. David’s are known in North Wales. It is only in the succeeding 
Menevian (Lowest Lingula-flags) that there is a well-marked com- 
munity. Of the 12 Pembrokeshire species (9 of which are 
Trilobita, viz. Agnostus cambrensis, Conocoryphe bufo, C. Lyellir, 
C. solvensis, Microdiscus sculptus, Paradowides aurora, P. Hicksw, 
P, Harkness, and Plutonia Sedgwickit) not one is known out of 
the area; whereas of the 28 Menevian species known in the St.- 
David’s area, 12 are common to it and North Wales and 5 are con- 
fined to North Wales, viz. Agnostus reticulatus, Ang., Anopolenus 
ampar, Hicks, Conocoryphe coronata, Barr., C. Homfrayi, Salt., 
and Hrinnys venulosa, Salt.; 7 are common to both areas, and 
16 peculiar to the Menevian promontory. These will be referred 
to under the Menevian group. 

This non-occurrence of fossil remains other than Annelide-burrows 
(Chondrites and Cruziana) in the Cambrian of the Geological Sur- 
vey has much significance when they are compared with groups of 
rocks equivalent in time, and not far removed geographically, meta- 
morphism not haying affected their original condition so as to have 
obliterated all traces of life. The sequence in both areas (North and 
South Wales) is the same, as proved or determined by the position 
of the Menevian beds, which in South Wales have a fossiliferous 
base in grey, red, and purple sandstones, these, again, resting upon 
a Pre-Cambrian foundation of highly metamorphosed rocks, divisible 
into three systems or formations, which Dr. Hicks has denominated 
Dimetian, Arvonian, and Pebidian, all having different strikes or 
bearings. We must remember that the Menevian beds of North 
Wales rest (so far as is at present known) on the unfossiliferous 
Bangor, Llanberris, and Harlech grits throughout their known ap- 
pearance and range. 


Lower CAmBRIAN Rocks. 


The fossils which occur in the lowest known part of the Harlech 
and Bangor group at St. David’s are Lingulella ferruginea, Salt., L. 
* The Middle Cambrian or Lower Lingula-flags of Sedgwick, the Lingula- 


flags of authors, and the Upper Cambrian of Lyell and Salter; the Menevian 
group of Salter and Hicks. 


. ANNIVERSARY ADDRESS OF THE PRESIDENT. 63 


primeva, Hicks, Discina, sp., D. caerfaiensis, Hicks, and Leperditia 
cambrensis, Toles: these all occur in the lowest red shales below the 
sands at Castell, on the east side of Ramsey Sound. About 1000 feet 
of unfossiliferous red and purple grits and sandstones succeed this 
first known fossiliferous group; then, in yellowish-grey flagg 

sandstones, between Porth-Clais Harbour and Caerbuddy, we meet, 
for the first time, with the gigantic Crustacea (Trilobita) Plutonia 
and Paradowides, and associated with them two genera, Microdiscus 
and Agnostus, almost microscopic in size, and at the same time, or 
in the same beds, Protozoa belonging to two species of Hexacti- 

nellid sponges, Pr otospongra fenestr ata, Salt., and P. major, Hicks, 
and also two species of Pteropoda, Theea antigua, Hicks, and .. 
penultima, Salt. I particularly mention this first assemblage of 
life, as being, so far as I know, the earliest in the earth’s aston "y 
on record; and it consists nevertheless of forms whose organiza- 
tion was not embryonic; thus distinctly pointing to a line of an- 
cestors belonging to this or some other area yet to be determined. 
The entire fauna of these Plutonia-beds consists of 8 genera and 
9 species, 5 of which are Trilobita (viz. Plutonia Sequickii, Hicks, 

Paradoxides Harknessii, Hicks, Oonocoryphe Lydellii, Hicks, Micro- 
cascus sculptus, Hicks, ’ Agnostus cambrensis, Hicks), two Protozoa 
(Protosponyia fenestrata, Salt., and P. major, Ticks), with Lingulella 
ferruginea, Salt., and Theea antiqua, Hicks; three of these, Para- 
dowides Harknessri, Agnostus cambrensis, and Protospongia fenestrata, 
pass to the Menevian group. Plutoma Sedgwicku, Conocoryphe 
Lyellia, and Microdiscus sculptus have not been detected in any 
higher horizon, and P. Harknessii only in the succeeding red, 
grey, and purple flagey sandstones, where it is associated with 
Conocoryphe solvensis, which represents Conocoryphe Lyellic below. 
Grey and purple sandstones, 1500 feet thick, but with a scanty 
fauna of only five species, including the doubtful Hophyton, succeed 
the Plutonia-sandstones. Finally the Harlech and Llanbezris group 
at St. David’s terminates with grey flaggy sandstones, which contain 
another form of Paradoxides (P. aurora), Conocoryphe bufo, Hicks, 
and Agnostus cambrensis, Hicks; in all eight species are known to 
occur. 

This first incoming of life in the British rocks and in the St.- 
David’s area exhibits a singular assemblage and grouping of 
species, doubtless owing to our limited acquaintance with’ the then 
existing fauna, rendered obscure through the present state of our 
collections. Numerically the fauna of the Lower Cambrian com- 
prises 61 genera and 182 species, ranging from the lowest red 
shales of the Longmynd group to the top of the Tremadoc. The 
Longmynd and Harlech beds yield 18 genera and 32 species, the 
Menevian group of South Wales 24 genera and 51 species, the 
Lower Lingula-flags of all localities 17 genera and 36 species, the 
Upper Lingula-flags 14 genera and 41 species, the Lower Tremadoc 
29 genera and 59 species, and the Upper Tremadoc 20 genera and 
33 species. Of this Lower-Cambrian fauna (61 genera and 182 
species) only 9 genera and 12 species pass to the Arenig; of these, 


64 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


5 are Trilobita (Asaphus affims, M‘Coy, A. Homfrayz, Salt., Chewrurus 
Frederici, Salt., Dionede atra, Salt., and Ogyqua scutatria, Salt.). It 
will be observed that these are all Tremadoc species, none of the earlier 
or more pronounced Longmynd, Menevian, and Lingula-flag forms 
ever appearing above the Lingula-flags; but assuming that the 
Tremadoe rocks terminate the lower division of the Cambrian series, 
they must be acknowledged to be of value. 

The species of Brachiopoda, as we should expect, tell much the 
same tale. They are Lingulella pruncva, L. ferruginea, var. ovalis, 
and Obolella maculata, which do not pass out of the lowest divi- 
sions; while Lingula petalon, L. lepis, Lingulella Davis, Obolella 
plicata, and Orthis Carausi, O. lenticularis, and O. menapice unite 
the Tremadoc to the Arenig. The remaining species are pelagic 
Mollusca. No Gasteropoda occur, and no form of Lamellibranch until 
we reach the Lower Tremadoc, where 12 species are met with, thus 
showing this earliest known fauna to have been highly specialized 
and of long duration ; this would be expected from the great physical 
development of the rocks, as well as through the large crustacean 
fauna, which numbers 28 genera and 108 species, only 5 of which 
passed to the succeeding horizon, or the Arenig, as before stated ; 
none of the great Olenide (Paradowides, Plutonia, and Neseuretus) 
passes to the higher division of the Cambrian or Cambro-Silurian 
rocks. 

The Pelagic fauna, 5 genera and 18 species, as exemplified by the 
Pteropoda and Heteropoda, was, and is still, the largest known, in- 
cluding Cyrtotheca 1 species, Theca 14, Stenorenn 1, Conularia 1, 
and Beller ‘“ophon 1. Nine of these are St.-David’s forms ; and only 
2 species of Pteropoda, Theca simplex and Conularia Homfrayi, 
with Bellerophon multistriatus, pass to the Arenig ; they will also be 
noticed under that group. 

Pranra.—Granting that the Oldhamiw might be caleiphites, or cal- 
careous corallines, resembling i in habit the Melobesice and Nullipore 
of modern seas, or the group of corallines so abundant in the seas 
of warmer latitudes, we are still at a loss as to their true nature, 
even if organic at all. Gdppert refers them to sea-weeds, and com- 
pares O. antigua with the living /zagora ramellosa of Kiutzing 
from Teneriffe ; but Goppert makes two genera of Oldhania; Mur- 
chisonites and Oldhamia, the O. antigua of Forbes being his Mur- 
chisonites antiqua. Professor Kinahan and Edward Forbes both 
referred. them to the class Hydrozoa as having affinities with the 
Sertulariide*. The Rev. Mr. Berkeley long ago suggested a resem- 
blance to the genus Acetabularia, one of the Ghiarospenmes (Chloro- 
sporex). No traces whatever of these singular remains have been 
found in any of the Cambrian rocks, either in North or South Wales. 
Mr. Salter well searched the Cambrian erits near Bangor and Harlech 
for Oldhanie ; but nothing approaching them was ever detected. 
Probably their place, in the absence of better evidence, is amongst 
the Hydrozoa. It is singular that, amidst these grandly developed 


* Vide Baily, ‘Figures of Characteristic Brit. Foss.’ vol. i. Paleozoic, p. 1, 
t. | (1875). ‘This book should be in the possession of all students of Palxozoic 
paleontology. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 65 


rocks of Wales, both North and South, no traces of this doubtful 
body have occurred. | 

Prorozoa.—Four species of Hexactinellid sponges (by no means 
uncommon in the Longmynd and Menevian rocks of St. David’s) 
have been described. ‘Three of the four species belong to the Har- 
lech beds, below the Menevian; and all four occur in the Menevian. 
Protospongia fenestrata, Salt., is also from the Lower Lingula-flags 
proper. Zittel places these sponges in the family Euretide with the 
Hexactinellide, and in the group Dictyonina. Mr. Carter believes 
them to be the remains of sarco-hexactinellid sponges, which Mr. 
Sollas has confirmed*. Mr. Salter was the first to notice these Amor- 
phozoa in the lowest and oldest rocks of Wales. ‘The original form of 
these sponges we know not; it may have been fiat and incrusting, 
like Grantia. Mr. Sollas suggests that they may have been “ sacci- 
form ” and anchored in the slimy ooze of the sea-bottom by a tuft of 
anchoring-spicules?. The spicules were originally siliceous, but are 
now converted into iron pyrites. The fact also that the spicules are 
separate, not being “ enveloped in a common coating, or united by 
ankylosis, clearly places or assigns them to the group Lyssakina of 
Zittel, nearly equivalent to Carter’s division of the Sarco-hexacti- 
nellide.” The species known are Protospongia diffusa, Salt., P. fene- 
strata, Salt., P. major, Hicks, and P. flabellata, Hicks. No traces 
of these have occurred either in the Harlech, Longmynd, Menevian, 
or Lingula-flags of North Wales. They may be sought for round 
the Harlech Dome, amidst the black shales and flags of the Menevian 
beds. 

AwnetipA.—The Longmynd group yields four genera and five 
species, Arenicolites uricomiensis, Call., A. sparsus, Salt., Scolrtes 
socialis?, Haughtoma pecila, Kin., and Histioderma hibernica, Kin. 
If the quartzites of the Wrekin in Shropshire are of the age assigned 
to them by Dr. Calloway, then the Arenicohtes uricomiensis of that 
author is, with the exception of Hozoon canadense, the oldest known 
fossil. Beyond the doubtful Annelidan tracks, and one species (Are- 
nicolites sparsus) from the Longmynd group of England and Wales, 
we have no determined species. ‘l'wo of the five species (Haughtona 
pecla, Kin.) and Histioderma hibernica, Kin., are Irish, from near 
Bray Point. Nine out of ten of the so-called fucoidal or plant- 
remains in the rocks below the Devonian are only Annelide-burrows. 

CrustAcEA.—Seven genera and 14 species occur in the lowest 
Cambrian or Harlech rocks of St. Davids, 10 of which are confined 
to them; the remaining + species (Agnostus cambrensis, Conoco- 
ryphe bufo, Paradoxides aurora, and P. Hicksw) connect the Har- 
lech and Longmynd Crustacea with the Menevian. The 7 genera 
are <Agnostus, Conocoryphe, Leperditia, Microdiscus, Paradoxides, 
Plutoma, and Paleopyge: 2 species of Conocoryphe (C. Lyell and 


* See Quart. Journ. Geol. Soc. vol. xxxvi. pp. 362-367, Sollas “On the 
Structure and Affinities of the Genus Profospongia.” 

t+ This has been confirmed through the occurrence of another form in the 
same family, lately described by Mr. Sollas from the Silurian of Canada. 


66 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


C. solvensis), 3 species of Leperditia (L. ferruginea, L. cambrensis, 
and L. primceva), Microdiscus sculptus, and Plutonia Sedgwicki 
are all confined to the Harlech and Llanberris group of St. 
David’s. They constitute the Crustacean fauna of that horizon, 
out of 103 species known in the six divisions of the Lower Cam- 
brian rocks, and up to the close of the Upper Tremadoc. The Me- 
neylan group possesses the largest Crustacean fauna in the eleyen 
classes, and, except the Brachiopoda, the largest number indi- 
vidually. 

Hyprozoa.—No form known below the Upper Lingula-flags or 
the Lower Tremadoc, from which Dr. Callaway has obtained the 
earliest known species, Bryograptus Callavei, Lapw. Mr. Lap- 
worth’s genus occurs in the Shineton shales of Shropshire. The 
same genus occurs in the Olenus-beds of Westrogothia, in Sweden, 
Linnarsson having detected these oldest Rhabdophora in that area. 

Bracutopopa.—Only 6 species are essentially Lower-Cambrian 
or Harlech forms, Discina pileolus, D. caerfaiensis, Lingulella ferru- 
ginea and var. ovalis, L. primeva, and Orthis sagitialis; this last 
species is doubtful. Five genera and 20 species range through the 
six horizons, or from the Longmynd group to the Upper Tremadoce ; 
but, as we have seen, only the 6 just named occur in the lowest 
horizon ; 4 of these belong to the Tretenterata (non-articulate group). 
It is doubtful if the others occur in the Longmynd group. 

LAMELLIBRANCHIATA.—No bivalve mollusk is known to occur 
below the Lower Tremadoc. In that formation for the first time 
in Britain we recognize 5 genera with 12 species. Davidia and 
Glyptarca are new ; the remaining 3 are the well-known genera 
Modvolopsis, Palearea, and Ctenodonta. This may be regarded as the 
first well-determined appearance of the Pelecypoda in any region. 

GastEropops.—None known in the whole of the series of forma- 
tions below the Arenig, where four genera seem to be recognized, 
both in South Wales (St. David’s) and in the Stiper-stones area. 
Pleurotomaria Uanvirnensis, Hicks, Huomphalus corndensis, Sow., 
and forms of Ophileta and Rhaphistoma occur. 

Prpropops.—The six horizons of the Lower Cambrian yield 4 
genera, Cyrtotheca, Stenotheca, Theca, and Conularia, and are repre- 
sented through the six divisions by 11 species; but only 2 occur in 
the Longmynd group of St. David’s under analysis (Theca antiqua, 
Hicks, and 7’. penultima, Salt.). The Menevian beds of the same 
area have yielded 6 species, to be noted under that group of rocks, 
and the Lower and Upper Tremadoc 9 species. During the Arenig 
and Caradoc periods only, have we to record so many pelagic Mol- 
lusca; 8 species occur in the Arenig, and 10 in the Caradoc. 

Hursroropa.— Bellerophon is the only form that occurs in these 
lowest rocks; but no species has been recorded lower than the so- 
called Middle Lingula-flags, where, as B. cambrensis, Belt, the genus 
first BU Nee, associated with Hymenocaris vermicauda and Cono- 
coryphe in the Upper Ffestiniog rocks ; but four of the five known 
species are Tremadoc, where, specifically, the genus becomes of im- 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 67 
portance. B. nultistriatus, Salt. (an Upper-Tremadoe species), unites 
the Heteropoda of the Lower with those of the Arenig or Middle 
Cambrian group. 

CrrHaLopopaA.—No form or species known until we reach the 
horizon of the Tremadoc rocks, where for the first time we find 
both Cyrtoceras and Orthoceras, one species of each genus. Cyrto- 
ceras precox, Salt., is, I believe, the earliest Cephalopod known. The 
Lower Tremadoc rocks of Llanerch, west of Portmadoc, yielded the 
Cyrtoceras ; and the Upper Tremadoc of Garth, and Llanvirn, St. 
David’s, O. sericeum. The latter passes to the Arenig rocks. 

We have thus seen how unequally distributed (as we should ex- 
pect) are the zoological groups through the lowest Cambrian rocks, 
and how distinctively certain genera characterize them, and, as will 
be found upon a separate analysis of the Menevian, Lingula-flags, 
and Tremadoc beds, how distinct a period is represented by all below 
and up to the close of the Tremadoc age. The six divisions of the 
Lower Cambrian represent the earliest history of the British rocks ; 
for out of the 61 genera and 182 species known to range through 
them, only 11 genera and 16 species pass to the Arenig or base 
of the Lower Silurian, or Middle Cambrian (see Table VII.). 


Taste 1.—Longmynd, Harlech, and Llanberris. 


Classes. Genera. | Species. ne oe | 
LPTe ance ave te ee eS Qin are 
PAROLOZOM raise secs tele Sesotecs 1 3 x 
Hydrozoa. 
Actinozoa. 
Kehinodermata. 
Ja CILIC ERS UR Ce MRR ees 4 | 4. 
Wisistaccan cee ee seen sce: i 14 z 
Bryozoa. | 
Brachiopoda ...............+.- 3 6 a 
Lamellibranchiata. 
Gasteropoda. 
HREEROWOOAL. tlie. cisecsccavosc 1 2 i 
Heteropoda. 
Cephalopoda. 

18 32 Be 


Dr. Hicks has favoured me with a new classification for the 
Lower Cambrian (Longmynd and Harlech) rocks of St. David’s: 
the geographical significance of the names employed renders them 
of value; and nothing of the kind has been previously attempted 
below the Menevian group :— 


68 PROCHEDINGS OF THE GHOLOGICAL SOCIETY. 


( Up bee Sandstones and shales, with Orthis Hicksii &e. 


VOOR: 
op hel at 
ee re e \ Flags and slates, with Paradoxides Davidis. 
al i 
{ ae \ Grey flags, with Paradoxides Hicksit &c. 
( ca oa Grey rocks, with Paradowides aurora &e. 
| Grey, purple, and red rocks, with Paradowxides solvensis 
group.) 1500 ft. | (Hicks, MS.), Conocoryphe solvensis, &c., also Hophy- 
{ 


Solva 4 Middle, 
i __ ton and large fucoids. 
| Lower, { Yellowish grits, sandstones, and flags, with Paradoxides 
| 150 ft. | Harknessii, Plutonia Sedqwickit, &c., also Eophyton. 


( Upper, \ Purple sandstones, with Annelids. 


1000 ft. 
ee | se Red shales and schists, with Leperditia cambrensis, 
OEE Middle, Lingulella primeva, Discina caerfaiensis (Hicks, 
group. | 50 fi. | MS), & 
| 


Lower, { Conglomerates and green flaggy sandstones, with An- 
520 ft. | nelids. 


Below the Caerfai group in South Wales, at St. David’s, occur the 
Dimetian and Pebidian rocks, Pre-Cambrian in age, unconformable 
in position, and discordant in strike ; they constitute the well-defined 
axis of the St.-David’s promontory. | 


MENEVIAN*® AND LINGULA-FLAGS. 


In 1865 Messrs, Salter and Hicks gave the name to and established 
this group of strata in the St.-David’s promontory, Pembrokeshire rf ; 
it constitutes the lowest division of the Middle Cambrian of Sedg- 
wick, and equals his Lower Lingula-flags, and is embraced in the 
Lingula-flags of authors and the Upper Cambrian of Lyell and Salter ; 
it is also recognized as being equivalent to Barrande’s Ktage C in 
Bohemia ; it occurs in Sweden, has been recognized in North Ame- 
rica and Canada, and is the St.-John’s group in Newfoundland. 

The Menevian is now recognized as a separate division or sub- 
group, distinguished from the Lingula-flags proper above, as the 
Ffestiniog of Sedgwick was applied to the Middle and Upper groups 
of the Lingula-flags in North Wales. In South Wales the Menevian 
is about 700 feet thick, and more or less fossiliferous throughout. 
Probably the base of the Menevian cannot be separated from the 
underlying grits and sandstones of the Harlech and Bangor series, 
passing as they insensibly do into them, both in North and South 
Wales, and also into the true Lingula-flags above. No fossils occur 
or have yet been detected in the grits and sandstones of the imme- 
diately underlying Harlech rocks, which comprise the structure 
of the Harlech Dome (Merioncthshire anticlinal of Sedgwick); the 
same may be said of the great and exposed masses of these Lower 
Cambrian rocks of the Bangor area, obscure traces of Annelide- 


* Meneevia is the classic name for St. David’s. ; y 
+ First announced at the British Association, Birmingham, 1865, ‘ Report, 
p. 281. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 69 


tracks (Chondrites) and burrows (Arenicolites) being all that has re- 
warded patient research ; but below the primordial Menevian, as 
we have seen, in the promontory of St. David’s, there exist greenish, 
grey, and red flaggy or shaly sandstones, with species of Lingulella 
differing from the typical LZ. Daviswz of succeeding age. In the 
yellow, grey, and purple sandstones and flags we have the first 
known Trilobites; and these are of gigantic size—the two genera 
Plutona and Paradoaides, with Conocoryphe, demanding from us 
the belief that they were by no means the first of Cambria’s Crus- 
tacea. 

The earlier highly metamorphosed rocks below the Caerfai group 
and the Plutonia-sandstones in all probability contained the pro- 
genitors of these early giants ; nowhere else in the British islands, 
Kurope, or America have species been found of such high antiquity. 
To the untiring energy of Dr. Hicks we owe the discovery of these 
rocks and the description of the species contained in them. 

The Menevian of the §$t.-David’s area rests upon and passes 
insensibly down into the fossiliferous Harlech and Bangor rocks, 
which contain an earlier fauna, which is not at all represented in 
North Wales, either at Harlech or Bangor. Only one of the five 
species of the great Paradoaides found in the pre-Menevian and 
Menevian beds of South Wales is known in North Wales. The 
long-known and solitary specimen called P. Forchhammeri, from 
an unknown locality, is now determined to be P. Hicksii, which 
occurs sparingly in the Menevian beds on the Camlan river north 
of Dolgelly, but more abundantly with its two congeners, P. aurora 
and P. Davidis, in South Wales, the fourth species, P.’ Harknessii, 
characterizing only the Harlech group at St. David’s*. Associated 
with these large Menevian Paradowxides, other forms, equally sig- 
nificant, occur, viz. Holocephalina inflata, Hicks, H. primordialis, 
Salt., and Carausia menevensis, Hicks; and they range no higher, 
being distinctively typical or characteristic species of this horizon. 

Two species of Anopolenus (A. Henrici, Salt., and A. impar, Hicks) 
are common to the two areas, the former occurring at Rhaidr-ddu 
valley, Tyddyngwladis, near Dolgelly, the latter at the Maentwrog 
Waterfall. 

No less than 6 species of the genus Conocoryphe occur in the 
Menevian group: 3 characterize the St.-David’s, and 2 the North- 
Wales beds; and 1 species, C. applanata, is common to both 
areas. Conocoryphe humerosa, Salt., C. bufo, Hicks, C. solvensis, St. 
Davyid’s only ; C. coronata, Barr., C. Homfrayi, Salt., North Wales, 
Maentwrog; C. applanaia uniting the two areas. Hrinnys venu- 
losa, Salt., and Arionellus longicephalus, Hicks, are both found in 
the Waterfall-valley at Maentwrog, and the latter also at St. David's. 
I draw attention to this group of Crustacea because they are strati- 
graphically important, and constitute by far the most characteristic 


* Dr. Hicks has added another species from the Trelewr beds below the 
Menevian, P. solvensis, associated with Conocoryphe solvensis and the so-called 
Eophyton. 

VOL. XX XVII. g 


7O PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


portion of the fauna of the lowest Cambrian rocks. No less than 
9 genera and 26 species occur in the Menevian rocks alone of North 
and South Wales; or, in other words, more than half the known 
fauna is Trilobitic. In the Menevian beds occur the first known 
Cystidean, a few fragments having been found by Dr. Hicks, in 1872, 
associated with Pteropoda (Theca) and Entomostraca. The finding of 
Paradoaides Davidis and P. Hicksvi at Dolgelly, on the same horizon 
as that of St. David’s, immediately above the “‘ Cambrian” sandstones 
(Harlech and Barmouth beds), in bands of uncleaved Menevian slate 
and ashes, aids greatly in the correlation of the two areas. This, with 
other evidence, justified Hicks and Salter in regarding the Menevian 
and the Lower Lingula-flags as a well-marked division or subgroup 
differing from the Upper Lingula-flags, quite as much so as do the 
Upper Lingula-flags from the overlying Lower Tremadoc slates (as 
now understood) ; for, with few exceptions, the species in each are 
peculiar, and they possess many peculiar genera. I may mention 
the following as genera that first appear in the Menevian beds; and 
those marked with an * are confined to that horizon :— 


Cystideas 2 see . &Protocystites. 
Amelia eee eee .. xArenicolttes. 
Crusticcas eee x Arionellus. 
Bee Un rprt ah bie ene + Anopolenus. 
Pa ate aaa ne eid OS * Hronnys. 


BENE ANU ee * Holocephalina. 
* Carausia. 


AOA neers Hare Primitia. 
Br achiopoda PNT eee ? Orthis 
net ee Obolella. 
Pteropoda SN asatet cae aes x Oyrtotheca. 


MAUR we ietisn ac, Ry x Stenotheci. j 


Although four oe of Protozoa (Protospongia. ee Salt., 
P. fenestrata, Salt., P. major, Hicks, and P. flabellata, Hicks) 
occur at St. David's, the three first named species commenced in 
the Harlech beds ielow, P. fiabellata being Menevian only. 

No Protozoa have yet been detected in the North-Wales area. 
Prior also to the year 1863 we had no knowledge of the exist- 
ence of the gigantic Olen, revealed to us by the researches of 
Salter, who disentombed the great Paradowides (P. Davidis) from 
the black slates of Porth-y-rhaw and Solva Harbour, thus showing 
the occurrence side by side in the same beds of the largest 
Trilobite known with Agnostus the smallest. Both these genera 
are remarkable ; and their extremes widely depart from the general 
type. meas 
Mh valuable paper was communicated to our Society in 1867, 
vol. xxiii., by Prof. Harkness and Dr. Hicks, “On the Ancient Rocks 
of St. David’s Promontory, South Wales, and their Fossil Contents.” 
The authors dealt fully with both the seology and the paleontology 
of that remarkable promontory. The chief portion of the labour and 


ANNIVERSARY ADDRESS OF THE PRESIDENT, rie 


research, however, was due to Dr. Hicks, whose intimate acquaint- 
ance with the area and knowledge of the fauna of the Cambrian 
and Silurian rocks enabled him to add largely to the palzontology 
of the district. The authors described the great series of red, purple, 
green, and grey rocks below the fossiliferous grey Menevian beds. 
The history of these variously coloured sandstones, as revealed by 
the researches already made in them, has an important bearing 
upon our knowledge of the distribution of life through the Cambrian 
and Silurian series of North Wales, with which they must be 
correlated, the St.-David’s series yielding a fauna not yet known 
in Monmouthshire, Caernarvonshire, or Montgomeryshire, or beneath 
the Menevian and Lingula-flags of North Wales, which contain 
organic remains older than, and different in species from any hitherto 
discovered in Britam. The underlying Cambrians repose upon a 
conglomerate, composed of quartz and other pebbles cemented in a 
purple or red arenaceous matrix, which occurs on both sides of the 
so-called Pre-Cambrian ridge or axis. That this quartziferous, 
metamorphosed, crystalline, and unconformable mass underlies and 
is older than the whole series of the Lower Cambrian rocks of that 
area is certain; and, as such, it is the key to the physical structure 
of the ancient headland of St. David’s. Subsequent research by 
Hicks has determined that this extensive and exposed Pre-Cambrian 
area is composed of three distinct groups of metamorphosed sedi- 
mentary rocks of different ages, and having different or discordant 
bearings or strikes*. Higher still, and also unconformably, succeed 
the rocks of the Harlech or Longmynd group, which lie both to the 
east and west. Until the Longmynd, Harlech, and Meneyian fauna 
was discovered and worked out at St. David’s, and the last named 
subsequently developed in North Wales, the lowest sedimentary 
rocks of Britain then known were believed to be almost devoid of 
life or “‘ barren in fossils.” Now, however, a remarkable fauna has 
rewarded the researches of many distinguished labourers, and the 
two areas have been carefully compared and correlated. Consider- 
able differences exist, of which the causes are as yet unknown; 
much has yet to be done in the paleontology of both these classical 
districts. The non-discovery or absence of the genus Olenus in the 
Menevian and Lingula-flags or primordial rocks of St. David’s is 
singular, and at present inexplicable. The genus is abundantly 
represented in both the Lower and Upper Lingula-flags of North 
Wales, where no less than 12 species are known, 4 in the lower 
division and 8 in the upper; and 4 distinct forms (viz. O. bisuleatus, 
O. scarabeeoides, O. humilis, and O. pauper) occur in the Upper Lin- 
gula-flags of Malvern, overlying the Hollybush sandstones. The 
species O. scarabeoides (O. spinulosus, Phill.) is found also in the 
Upper Lingula-flags of Carreg-Wen, west of Portmadoc. 

Comparison of the earliest known faunas in Hurope and America 
with that of St. David’s shows conclusively that they are identical as 
to age or time of deposition and in genera also. This Angelin has 

* See p. 56 ef seq. 
g2 


12 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


demonstrated through his researches into the alum shale of Sweden ; 
and the primordial fauna of Barrande has yielded Paradowxides, Cono- 
coryphe, Agnostus, and Ellipsocephalus, the first three of which are 
characteristic British genera. 

Sweden has afforded P. Hicksw and other forms like our own. 

The Spanish primordial rocks yield genera identical with those of 
our Menevian and Lingula-flags—Puaradowides, Conocoryphe, Ario- 
nellus, and Agnostus. America, has its Acadian group, with Para- 
doxides &c. The Protozoic schists of Bohemia (Region C, Barrande), 
also contain Paradoaides (P. bohemicus), Sao, Conocoryphe, and 
Agnostus. The same conditions prevail in Canada and New Bruns- 
wick. It would thus appear that all the regions in Europe and 
America north of 30° of latitude to the polar regions contain these 
primordial rocks and fossils. 

I regard the Menevian of St. David’s as being the most typical, 
as regards both physical development and the abundance of organic 
remains (individually and specifically). The intricacy of the geo- 
logical structure of North Wales renders both the mapping and the 
collecting of fossils in the Menevians there far more difficult than 
in the St.-Dayid’s promontory, the strike of the beds being more 
disturbed in continuity ; but I doubt not that an extensive fauna 
is yet to be obtained from the Menevian of North Wales, where the 
typical Lower Lingula-flags mantle round the still older Cambrian 
masses. 

The paleontological value of any group of rocks can only be 
arrived at through such analysis as I have here attempted to give ; 
it is the census at a given time or age, and from such may be cast or 
determined (approximately) the true and absolute value of the zoolo- 
gical groups and their distribution through their respective formations. 

PLanta.—No traces whatever of plant-remains have occurred in 
the Menevian. 

Protozoa.—The St.-David’s beds of this age contain all the known 
4 species of Protospongia, viz. P. diffusa, P. fenestrata, P. major, 
and P. flabellata, the first three of which, as we have seen, are also 
Longmynd and Harlech forms. At St. David’s P. fenestrata passes 
to the Lower Lingula-flags proper. None have occurred either in the 
Menevian or Lingula-flags of North Wales. 

Hyprozoa.—No traces. : 

Actrnozoa.—Totally unrepresented. 

EcurnopERMATA.—Cystidean remains were detected by Mr. Salter 
in 1866 at St. David’s; to these he gave the generic name Proto- 
cystites. Dr. Hicks subsequently named these fragments P. mene- 
vensis, after the horizon in which they occur. These obscure fossils 
consist of arm-ossicles and body-plates ; it is the first Cystidean re- 
corded. Dr. Hicks has also determined the presence of Dendro- 
crinus cambrensis in the Tremadoe rocks of St. David’s, the oldest 
Crinoid known in the British rocks. 

Aynetipa.—Arenicolites didymus, Salt., A. sparsus, Salt., and Ser- 
pulites fistula, Hall, appear to be all that are known of the Anarthro- 


ANNIVERSARY ADDRESS OF THE PRESIDENT. ks 


poda in the Meneyian beds. ‘The first two are confined ue this 
horizon ; S. fistula passes to the Upper Lingula-flags. 

Orusracra. —The largest crustacean fauna of the s1x ieaeear of 
the lowest Cambrian “dee occurs here. 12 genera and 32 species 
characterize the Menevian beds both of North and South Wales. 
Agnostus is represented by 7 species, Anopolenus by 3, Conocoryphe 
by 7, Paradowides by 3, Holocephalina by 2, and Leperditia by 4 
species. ‘These 6 genera are the chief and most largely represented. , 
Arionelius, Erinnys, Microdiscus, and Carausia, each with one species, 
are confined to the Menevian beds, and generically characterize them. 
The black slaty flags of St. David’s yield the gigantic Paradoaides P. 
aurora, Salt., P. Davidis, Salt., and P. Hicksi, Salt.; this last-named 
species has been recorded from the Lingula-flags of North Wales under 
the name P. Horchhammert. 14 of the 32 species pass to the Lower 
Lingula-flags, but none higher; and the Menevian beds are con- 
nected below with the Harlech beds and the Longmynd group 
through 4 species only, viz. Agnostus cambrensis, Hicks, Conoco- 
ryphe bufo, Hicks, Paradoaxides aurora, Salt., and P. Hicksi, Salt. 
No Olent occur in the Menevian rocks, although 13 species belong 
to the Lingula-flags proper. Olenusis essentially an Upper Lingula- 
flag genus, 9 species occurring in that series; and none pass out of 
the Lower Cambrian rocks or above the Tremadoc group, where 
4 species occur (O. alatus, O. umpar, O. triarthrus, and O. Saltert). 

Bryozoa.—No remains of this class occur in the Menevian ; they 
first appear in the Upper Lingula-flags under the form of Dictyonema? 

Bracuiopopa.—Only 4 genera and 6 species range through the © 
Meneyvian beds, Orthis Hicksvi, Salt., being the only peculiar form. 
Lingula ferruginea, Salt., and var. ovalis, Hicks, Obolella maculata, 
Hicks, Discina pileolus, Hicks, and Orthis sagitialis, M‘Coy, are asso- 
ciated either with the Harlech beds below or the Lower Lingula beds 
above. very species occurs in the St.-David’s Menevian beds. It 
is not until we rise into the Llandeilo and Caradoc rocks that the 
class becomes numerically so distinguished and important a factor 
in the Cambrian or Silurian rocks. 

LAMELLIBRANCHIATA.—No form known in the Menevian beds. 

GastEropopa.—No vestige known in the British rocks of this age. 

Preropopa.—Cyrtotheca hamula, Hicks, Stenotheca cornucopia, 
Salt., Theca corrugata, Salt., T. penultima, Salt., 7’. menevensis, Salt., 
and J’. stiletto, Salt., or 3 genera and 6 species, are known. 4 of 
the 6 species are Menevian only, viz. C. hamula, S. cornucopia, 
7’. menevensis, and T’. stiletto. T. corrugata is the only form that 
passes to the Lower Lingula-flags above. 

Herrropopa.— None known in the Menevian of either South or 
North Wales. 

CrpHaLopopa.—No species has yet occurred in the British rocks 
of this horizon, the only two species in the Lower Cambrian being 
those before mentioned, Oyrtoceras precox and Orthoceras sericeum, 
both of which are Tremadoc. 

Thus we see that, out of the 52 species that constitute the fauna 


74 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


of the Menevian rocks, 32 are Crustacea—the only other classes 
fairly represented being the Brachiopoda, 6 species, and the Ptero- 
poda, 6 species. A marked feature is the appearance of Cystidea 
(Protocystites) and Crinoidea (Dendrocrinus). Hight whole classes are 
entirely unrepresented : the Plante, Hydrozoa, Actinozoa, Bryozoa, 
Lamellibranchiata, Gasteropoda, Heteropoda, and Cephalopoda are all 
absent. The genera that first made their appearance in the British 
Menevian sea were Protocystites, Dendrocrmus, Serpulites, 7 Crus- 
tacea (Holocephalina, Entonus, Anopolenus, Arionellus, Leperditia, 
Erynnis, and Carausia), Orthis among the Brachiopoda, and 2 Pte- 
ropoda (Cyrtotheca and Stenotheca). The other genera appeared for 
the first time in the lowest Cambrian of the St.-David’s promontory. 


Taste 11.—Menevian. 


Oe 
4 me Pass to 
6 WS Classes. Genera. | Species. | Lower 
a & a Lingula. 
Plante. 
a Br ObOZOa encase ant 1 4 1 
Hydrozoa. 
Actinozoa. 
Echinodermata...... I 1 
Nina Shi Ee okbeebodnas 3 3 
4 Crustacea Wy... ee 12 32 Vy 
Bryozoa. 
3 Brachiopoda......... 4 6 3 
Lamellibranchiata. 
Gasteropoda. 
‘ Pteropodarss...-4-c 3 6 4 
Heteropoda. 
Cephalopoda. 
eR iiaileey PRE esto ecoesae 24. 52 a 


LINGULA-FLAGS. 


The fossiliferous strata lying both above and below the Lingula- 
group have of late years been well described and their fossil contents 
marked out, the fauna of each group carefully investigated, com- 
pared, and catalogued. The Harlech and Llanberris and also the 
Menevian beds of South Wales have been exhaustively illustrated 
by Hicks and Salter. Upon the Tremadoc group, in North Wales, 
the labours of Mr. Homfray, Mr. Ash, and the late Mr. Salter leave 
little to be done; subsequently Dr. Hicks investigated the Tre- 
madoc and Arenig groups in the St.-David’s promontory, with what 
success I hope to show. It must not be forgotten that since the 
Geological Survey (thirty years ago) mapped much of North and 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 75 


South Wales, a great amount of new work has been done by eminent 
geologists and paleontologists. Much of the work done by the 
Survey has had to be readjusted and reconstructed ; the maps then 
prepared with so much skill by Ramsay, Selwyn, Jukes, Aveline, 
and others, and which contributed so much to our knowledge, and to 
the elucidation of the structure of these intricate regions, are now 
in some areas behind the requirements of the age, through the pro- 
gress of modern research and nomenclature. The horizons of many 
of the lowest fossiliferous groups of rocks as now recognized are 
not expressed or delineated on the Survey maps; the now well- 
determined Menevian, Tremadoc, and Arenig rocks of North and 
South Wales—in other words, all the fossiliferous deposits below 
the Llandeilo flags, or between that formation and the base of the 
Tremadoc, are not yet delineated on the maps of the Survey. It 
may be said by some that the groups of the Harlech, Menevian, and 
Lingula-flags may be subdivided and placed upon the Survey maps ; 
but the aspect of these beds and the way they occur in the field will 
ever prevent this on al-inch scale. ‘Transcendentalism in mapping 
has been, and still is, carried to a greater extent and perfection by 
the officers of the British Survey than by any other government survey 
in the world; and although neither the Tremadoc nor Arenig rocks 
are recognized on the maps of the Survey, I believe it will be found 
that these formations have received the fullest recognition and atten- 
tion in the forthcoming new edition of Prof. Ramsay’s ‘ Geology of 
North Wales.’ 

The Lingula-flags of the Survey and authors generally are equiva- 
lent to the Middle Cambrian of Sedgwick, and the Upper Cambrian 
of Lyell and Salter; the divisions into Lower, Middle, and Upper 
have long troubled systematists in Britain; and the extensive 
and almost hypercritical subdivisions adopted have led to much 
misunderstanding. ‘The obscurity of these beds in the field, the 
smallness of the fauna, and the difference in physical condition in 
areas widely separated often render it a matter of opinion where the 
lines of demarcation should be drawn, or what should be embraced 
by the terms Lower, Middle, and Upper Lingula-flags. The so-called 
Middle group has no value whatever, its fauna consisting of five 
species only. Three of these are Brachiopoda—Kutorgina cingulata, 
Lingulella Davisii, and Lingula squamosa. There is one Trilobite 
(Conocoryphe? bucephala, Belt), and also Bellerophon cambrensis, Belt. 
These last two forms occur in the upper part of the Dolgelly beds 
of that area, and are said to have been associated with Hymenocarts 
vermicauda, elsewhere known only in the Lower Lingula-flags. 

We must also remember that the ‘‘ Lower Lingula-flags” of Sedg- 
wick are the ‘‘ Menevian group” of Salter and Hicks, and that the 
“« Wfestiniog group” of Sedgwick constitutes the Middle and Upper 
Lingula-flags of Salter. Thus the greatest care is required when 
analyzing or correlating these groups, either through their literature 
or by research in the field. In the year 1867 the Jate Mr. Belt* con- 


* Geol. Mag. vol. iv. 


76 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


tributed much to our knowledge of the fauna and geology of the 
Lingula-flag series. He was then engaged in investigating the 
Dolgelly and other districts, especially with reference to the divi- 
sions of the Lingula-flags called the Maentwrog, Ffestiniog, Dolgelly, 
and Tremadoc groups, lying between the Menevian and Tremadoc 
beds, and occupying extensive areas near Dolgelly and in the 
Mawddach valley, also at Maentwrog near Ffestiniog &c. 

In 1847 Prof. Sedgwick, on paleontological evidence, removed 
the previously named Tremadoe series from the “ Lingula-flags,” 
and designated the latter the Ffestmiog group. Salter divided the 
Lingula-flags, in 1866, into Lower, Middle, and Upper. Again, 
Salter and Hicks, in the same year*, included the Lingula-fiags of 
South Wales only in the Ffestiniog group, describing them as being 
characterized by “‘ hard siliceous sandstone with grey flaky slate”’ con- 
taining “‘Lingulella Davisi.” In character the North-Wales group 
of rocks differs altogether from those of South Wales, the arenaceous 
flags and shales with Z. Davisi forming only a subordinate part of 
a series of fine-grained dark blue and black slates containing many 
genera of Trilobites. Mr. Belt restricted the Ffestiniog beds to the 
flags containing Lingulella Davisii and the Phyllopod Crustacean 
Hymenocaris vermicauda, as originally applied by Sedgwick, and 
proposed to name the slates and flags below them the Maentwrog 
group, characterized by peculiar Olent. These Maentwrog beds are 
exhibited to great perfection at Maentwrog, 8.W. of Ffestiniog. 

The blue and black slates occurring above the Ffestiniog series 
Belt named the Dolgelly group, with reference to the circumstance 
that it is only in this area that both members have been found. 
This upper member of the Lingula-flags is characterized by peculiar 
forms of Oleni, comprising the subgenera Spherophihalmus, Para- 
bolina, &c. These three groups—the Maentwrog, Ffestiniog, and 
Dolgelly—are paleontologically and lithologically distinct ; none of 
the Trilobita passes from one group to another, peculiar genera occur- 
ring in each. Each group also has well-defined lithological charac- 
ters. The Maentwrog is readily distinguished by its dark-blue 
jointed ferruginous slates; the Ffestiniog by hard micaceous flags 
with abundant Lingulella Davis, Hymenocaris, and Conocoryphe ; 
and the uppermost or Dolgelly, by species of Agnostus, Olenus and 
its subgenera, which are abundant in the soft black slate, which 
shows a black streak when scratened. Mr. Belt divided the Maent- 
wrog slates into a lower and upper series: the lower are ripple- 
marked, and have many Annelide-tracks. They are conformable to 
the blue-black Menevian slates which mantle round the Merioneth- 
shire anticlinal. This lower group is about 700 feet thick. Olenus 
gibbosus, Wahl., and Agnostus pisiformis, Linn. (A. princeps, Salt.), 
var. obesus, Belt, occur in the slates. The range between the Eden 
and the Mawddach, and near Dolmelynllyn, in the Mawddach, are 
the chief localities for fossils. The upper Maentwrog series is 
nearly 2000 feet thick. Agnostus princeps is abundant in the flaky 


* Brit. Assoc. Reports, 1866. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. Th 


beds near the summit above Dolgoed ; in thin slaty shales it occurs 
in thousands, associated with Olenus truncatus, Ang.; the higher 
beds contain O. cataractes, Salt., which replaces A. princeps. ‘This 
eroup is completely developed in the Maentwrog district, fine sec- 
tions occurring in the Waterfall valley and in the valley running 
from Taffarn-helig to Caen-y-coed. Olenus cataractes abounds here. 


Lower Frestrinioe Bens. 


Thick micaceous grey flags, 2000 feet thick, conformably overlie 
the upper Maentwrog beds; the lowest series are bluish-grey slates, 
and contain Lingulella Davisu, M‘Coy, with Annelide-tracks ; and 
the highest part of the Lower Ffestiniog slates yields Hymenocaris 
vermicauda, only known in this division of the Lingula-flags, 


Urrrr Frestrnioe. 


This group succeeds the Lower series, and is not more than 50 feet 
thick, but distinct in fossil contents. The characteristic forms here 
are Olenus micrurus and Bellerophon cambrensis. 'This thinly deve- 
loped division occurs at Gwern-y-barend, in the Mawddach near 
Craig-y-dinas and on Mynydd-Gader. 

The river Mawddach cuts through the whole of these beds 
between Rhiufelyn and Hatod-fraith, thence ranging across the east 
end of Moel Hafodowen and by Pen-y-bryn. 


DoLegELLy GRovp. 


This uppermost division of the Lingula-flags is physically divided 
into two series, lower and upper, the former composed of hard 
blue slates abundantly filled with Olenus (Parabolina) spinulosus. 
The thickness of this series is about 300 feet. 

The upper division differs essentially from the lower. It consists 
of soft black slates, with black streak, intensely cleaved and often 
pisolitic (fine grains of pisolitic iron-ore). The constant character 
of the black streak distinctively marks this group, and in the most 
disturbed areas may be relied on as a never-failing criterion of the 
upper series. To Mr. Belt is due the determination of this character 
over a large area—a fact of inestimable value in the identification 
of strata so disturbed and intricately faulted as those round Dolgelly, 
Rhobell-fawr, and on Mynydd-Gader. The black shales of Malvern, 
overlying the Hollybush Sandstones, and containing Olenus bisulcatus, 
Phill., O. scarabeoides, Wabl., O. pauper, O. humilis, Conocoryphe 
malvernensis, Agnostus princeps, A. M‘Coyr, &c., belong here; they 
underhe the Dictyonema-shales, and are exposed in the valley of the 
White-leaved Oak. The upper Dolgelly beds are by far the most 
fossiliferous ; and nearly every species in any of the areas is confined 


78 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


to this division. The Trilobites in the Dolgelly area are Conocoryphe 
abdita, Salt., C. Williamson, Belt, C. longispina, Belt, Olenus 
scarabeordes, Sphcerophthalmus (Olenus) alatus, S. (Olenus) humalis, 
Phill., Agnostus princeps, Salt., A. trisectus, Salt., and <A. obtusus, 
Belt; and, as before stated, the Malvern species are equally restricted. 
No one of these occurs below in the Middle and Lower Lingula-flags. 
Orthis lenticularis, Dalm., and Obolella follow the same rule all 
through the Maentwrog, Ffestiniog, and Dolgelly deposits; the Tri- 
lobita that successively appear are closely allied, or belong to the 
Agnostide and Olenide; no new types of structure come in, the 
fauna being distinctive and similar and homogeneous as a whole. 
Dikelocephalus, Conocoryphe, Olenus, and Paradoaides constitute the 
natural group or family Olenide ; and the single genus Agnostus is 
rich in species. The succeeding Tremadoc and Arenig rocks evidence 
great and sudden change, faunal and otherwise, the large Asaphide, 
Cheiruride, Trinucleide, and Calymenide then first appearing ; 
sixteen genera at once occur in the Arenig, and two in the Tremadoe 
(Niobe and Psilocephalus); and we have no proof of any unconformity 
in either the North or the South Wales areas: these large forms 
come in at once and in vast numbers; and the old forms die out. 
Nor have we as yet any evidence of these new and distinct types 
having had prior existence in some other area, and migrated into 
the Tremadoc or Arenig seas; we know not at present where to look 
for evidence of this. Ireland possesses no rocks or fossils of this age 
or earlier, Scandinavia none, Western Europe none. America then 
possessed gigantic Asaphi ; but in which direction dispersion or dis- 
tribution may have taken place we have no trace; although the 
evidence tends to show that it must have been towards the east, 
or Europe, the main strike of the oldest American rocks being N.H. 
and §.W.; while the older groups are greatly developed on the 
eastern side of the North-American continent, and there is con- 
siderable affinity between the faunas of the two areas, the facies 
strongly resembling each other. No migration of an earlier fauna 
from one area would fully account for the disappearance of the 
earlier gigantic Trilobites (such as Paradoaides, Plutonia) and other 
equally characteristic genera (such as Hrinnys, Holocephalina, &c.), 
all of which characterize the lowest rocks of Britain, ranging 
from the Harlech to the top of the Lingula-flags, call them what 
we will. 

Salter, as far back as 1853, most carefully described the two groups 
(Lower and Upper) of the Lingula-flags in North Wales; he ex- 
amined them in their most typical localities, selecting Maentwrog, 
Tremadoc, Ffestiniog, Dolgelly, Carnedd Ffiliast, Bangor, &c. 
as those places where the Lower division could be best studied. 
This he divided into two sections, the lowest consisting of black 
pyritous slates, with numerous beds of intercalated sandstones near 
the base. These probably in part were Menevian; but the prevail- 
ing fossils are Agnostus princeps, Olenus cataractes, and Lingulella 
Davisii. (Salter’s species A. princeps is the A. pisiformes, Linn., 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 79 


of the Alum slates of Sweden and Norway, which holds the same 
position there as in our own area.) 

The second division or upper part of the Lower Lingula-flags of 
Salter (the typical Lingula-flags) are finely developed in North Wales, 
near Tremadoc and Ffestiniog, at Maentwrog, Borth, Marchllyn-Mawr, 
Llanberris, Carnedd Ffiliast, Dolgelly, and Trawsfynydd. Cruziana 
semiplicata, Hymenocaris vermicauda, Olenus nucrurus, Lingulella 
Davisii, and Microdiscus are the leading types of this upper part of 
the lower series. Physically these are hard, grey, thick, sandy 
beds and flags. 

The Upper Lingula-flags of Salter differ essentially from the Lower 
Division, consisting of dense black pyritous slates, or black slates with 
pyritous bands near the base, with a large fauna differing almost en- 
tirely from that of the Lower Lingula-flags; for out of 35 species oc- 
curring in the lower group, only 5 pass from the lower to the upper— 
namely Agnostus princeps, Microdiscus punctatus, Lingulella Davisir, 
L. lepis, and Obolella sagittalis, and this in the same district. At 
Tremadoc, Maentwrog, and west of Ffestiniog the beds are rich in 
fossils—Olenus alatus, O. scarabeoides, Agnostus primceps, Conoco- 
ryphe depressa, C. invita, Orthis lenticularis, and Dictyonema thickly 
crowding the beds. 

Tt is well known that in 1851 Barrande visited England pur- 
posely to compare his fine series of fossils with our types, when he 
at once recognized the “ Lingula-flags ” of Sedgwick as the precise 
equivalent of his primordial stratum (Etage C). lBarrande also 
carried out his comparison and correlation not only with the fauna 
of our country, but also between those of the Kuropean and American 
continents, with what success the literature and progress of geological 
science has shown. 

The comparison of the Arenigs of Shropshire (Stiper stones) and 
Westmoreland (Skiddaw) will form an important feature for our 
consideration. 

The close analyses of the Lower and Upper Lingula-flags must be 
taken separately. The faunas of the two differ remarkably ; and only 
8 species connect them—namely 5 species of Trilobita (Asaphus 
princeps, A. trisectus, A. Salter, Microdiscus punctatus, and Para- 
doxides Hicksii) and 3 of Brachiopoda (Lingulella Davisu, L. lepis ; 
and Orthis sagittalis). The Middle Lingula-flags of some authors con- 
tain also only 5 species :—1 Trilobite, Conocoryphe bucephala, Belt ; 
3 Brachiopoda, Kutorgina cingulata, Belt, Lingula squamosa, Holl, 
and Langulella Davisii; with Bellerophon cambrensis, Belt. Two 
of the five species appear to be confined to this horizon, viz. Lingula 
squamosa and Bellerophon cambrensis. This fauna is too small to 
be of any value for our present purpose. 

Pranta.—No trace of any kind. 

Prorozoa.—Thesponge Protospongia fenestrata, Salt., passes up from 
the Menevian and Harlech beds at St. David’s to the Lower Lingula- 
flags. ‘This species has the longest range of the four known forms ; 
but none of the four ranges higher. 


80 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


Hyprozoa.—No appearance; none occur before the Arenig rocks 
were deposited. 

Actinozoa.—No traces of Coelenterata in any form. 

KcuinopERMATA.—N one occur in the Lingula-flags. The cystidean 
Protocystites menevensis is Menevian only; and the first Crinoid 
appears in the Lower Tremadoc of Ramsey Island. 

ANNELIDA.—Three genera and three species. The characteristic 
Cruziana semiplicata, Salt., hitherto has only been found in the Cam- 
brian grits of the Bangor area, near Cwm Grainog and Carnedd- 
ffiliast, in North Wales (Lower Lingula-flags of the Survey). 

Specimens of this Cruziana or an allied species have occurred 
below the Stiper stones in the Shelve country, W. of the Longmynds. 

Scolicoderma tuberculata, from Pentre-felen quarry, near Wem 
Gate, Caernarvonshire, and Helminthites from the Maentwrog sand- 
stones. None of these forms occurs out of the Lower Lingula-flags. 

CrustacEa.—This class is nearly equal in zoological value to what 
it is in the Menevian. 9 genera and 25 species have been described : 
14 of the same species appeared before in the Menevian, and therefore 
occur in both formations ; 6 species (Agnostus princeps, A. trisectus, 
Anopolenus Salterr, Microdiscus punctatus, Paradoaides aurora, and 
P. Hicksw) pass to the Upper Lingula-flags. Like the Menevian 
beds it is the home of Agnostus, 6 species of that genus occurring ; 
but only 2 are peculiar or confined to the Lower Lingula-flags, 
viz. Agnostus limbatus and A. nodosus; besides these Olenus cata- 
ractes, Salt., O. micrurus, O. gibbosus, and O. truncatus? are also 
Trilobites confined to this horizon; the Phyllopod Hymenocaris 
vermicauda essentially characterizes the Lower Lingula-flags; so 
that, large as the Crustacean fauna appears (25 species), there are 
only 8 that are really Lower-Lingula species; the remaining 17 
ally the Lower Lingula either with the two formations below or 
with the Upper Lingula and Tremadoc above. The 7 typical spe- 
cles are given above. The genus Conocoryphe, which is very largely 
represented in the Lower Cambrian rocks (by no less than 21 spe- 
cies), has no distinctive or peculiar form in the Lower Lingula-fiags ; 
4 occur, but none are restricted. They are Conocoryphe applanata, 
C. bucephala, C. humerosa, and C. variolaris. 9 species of the 
genus Conocoryphe occur in the Upper Lingula-flags; and, as we 
shall see, this division possesses also more peculiar forms and a 
larger specific Crustacean fauna than the Lower Lingula-flags, but 
fewer genera. ‘The species of Conocoryphe and other genera are as 
follows—Agnostus obtusus, A. princeps, A. trisectus, A. Turneri, 
A. venulosus, Ampyx prenuntius, Anopolenus Salteri, Conocoryphe 
abdita, C. bufo, C. depressa, C. invita, C. malvernensis, C. longispina, 
CO. Plantii, C. simplex, C. Williamsoni, Microdiscus punctatus, Olenus 
bisulcatus, O. flagellifer, O. (Sphceerophthalmus) humilis, O. obesus, 
O. pauper, O. pecten, O. (Peltura) scarabeoides, O. serratus, O. spi- 
nulosus, Paradoxides aurora, and P. Hicksiit, also Primitia sol- 
vensis, 

Bryozoa.—None have yet appeared. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 8I 


Bracutopopa.—Lingulella, Obolella, and Orthis are the only 3 
genera in the Lower Lingula-flags. The 5 species are Lingulella 
Davisu and L. lepis (both ranging to the Arenig), L. ferruginea, 
Obolella maculata, and Orthis sagittalis. The last named is also 
Tremadoc. 

LAMELLIBRANCHIATA.—None known below the Lower Tremadoc, 
where 3 new genera and 12 species suddenly appear; they will be 
noticed under the Lower Tremadoc beds. 

GasrERopopa.—None occur beneath the Arenig rocks. 

Preropopsa.—Theea corrugata, Salt.,and 7’. obtusa, Salt., are the 
only 2 species that occur. The last named is confined to the Lower 
Lingula-flags. 

Hereropopsa.—None below the Middle Lingula-flags ; and the only 
species occurring therein is Bellerophon cambrensis, Belt. 

CrpHatopopa.—None have yet appeared below the Upper Tremadoc, 
where the oldest forms known occur. 

The fauna of the Lower Lingula-flags is expressed in the follow- 
ing Table; and it will be seen in the passage column that only 8 
species pass to the Upper Lingula-flags. They are Agnostus prin- 
ceps, A. trisectus, Anopolenus Salieri, Microdiseus punctatus, and 
Paradoxides Hicks, Lingullela Davis, L. lepis, and Orthis sagittalis. 
Only 2 species are believed to connect the Middle and Lower Lin- 
gula-flags—Oonocoryphe bucephala, Belt, and Lingulella Davisn, 
M‘Coy. Indeed the whole fauna of the middle division only amounts 
to 5 species—the two just named, and Kutorgina cingulata, Bil., 
Lingula squamosa, Holl, and Bellerophon cambrensis, Belt. 

Nine entire classes are unrepresented in the Lower Lingula-flags, 
as shown in the Table. 


Taare IT1.—Lower Lingula-flags. 


Pass up Pass to | Pass to 
from Classes. Genera. | Species. | Middle | Upper 
Menevian. Lingula-| Lingula- 

flags. flags. 
Plante. 
7 IPROUOZOR) Suresh sca ceewe nes if 1 
Hydrozoa. 
Actinozoa. 
Echinodermata. 
Annelida ....... Rag Wane 3 3 
rie (AAU TUSUACEA cose ceo hesasse cess 9 25 s a 
Bryozoa. 
: iBrachiopodace: 8. o.<..5- 3 5 a : 
Lamellibranchiata. 
Gasteropoda. 
Pteropodarss se.) 0see=- 1 2 
Heteropoda. 
Cephalopoda. 
1 nase gh aM Cee ee ta 17 36 z z 


82 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Upper LINGULA-FLAGS. 


Prantz.—None known. 

Prorozoa.—None occur. 

Hyproz0a.—Dretyonema sociale occurs abundantly in the grey and 
dark shales of Keys End Hill, Malvern, andin North Wales. Whether 
Dutyonema should be placed with the Hydrozoa or Bryozoa is still 
a doubtful question. Salter in 1857 separated this genus from 
Kjerulf’s genus Fenestella and also from Hall’s genus Graptopora. 

Actiyozoa.—None. 

Koutyopermata.—None. 

ANNELIDA.—2 genera, Scolicoderma antiquissimum, Salt., and Ser- 
pulites fistula, Holl. The Hollybush sandstone of the Malverns is 
characterized by the two Annelides above named; the Hollybush 
beds of the Wrekin also contain Annelide remains. S. fistula first 
appears in the Menevian beds. Whatever may be the age of the 
Hollybush sandstones, these two species occur in the Malvern area, 
at the base of the Upper Lingula-flags. 

Crustacea.—No less than 30 species represent the following eight 
genera—Agnostus 5 species, Ampyw 1, Anopolenus 1, Conocoryphe 9, 
Dikelocephalus 2, Microdiscus 1, Olenus 9, Paradowides 2*. 4 species, 
viz. Agnostus princeps, Ampyx prenuntius, Conocoryphe depressa, and 
Olenus alatus, pass from the Upper Lingula-flags to the Lower Tre- 
madoc’; and 20 species are confined to this horizon. Thus the Upper 
Lingula-flags through their Crustacea become of much importance, 
and the few (4) that pass to the Tremadoe clearly show the strati- 
graphical value of the Lingula group and its distinctness. It may be 
worth stating that 4 species in the Upper Lingula-flags also occur 
in the Menevian; they are Anopolenus Salteri, Microdiscus punctatus, 
Paradoxides aurora, and P. Hicksiv; only 2 species of the 4 that 
pass from the Upper Lingula-flags to the Tremadoc extend up into 
the Upper Tremadoc, namely Agnostus princeps and Ampyx pre- 
nuntius. 

Bryozoa.—Dictyonema sociale, Salt., is the only Bryozoon known 
in the Upper Lingula-flags ; it passes to the Lower Tremadoce, both in 
the North and South Welsh areas, at Tremadoc and St. Dayid’s; it is 
highly characteristic of the pale Dictyonema-shales that overlie the 
black shales in the valley of the White-leaved Oak at the southern end 
of the Malvern ranget. These black and pale greenish Upper Lin- 
gula shales are about 500 feet thick, and contain a singular fauna 
of Crustacea and Brachiopoda. Prof. Phillips first noticed these 
shales and their fossil contents. The Crustacea are Conocoryphe 
malvernensis, Phill., Olenus bisulcatus, Phill., O. humilis, Phill., O. 
spinulosus, Phill., and O. pauper, Phill., Spherophthalmus pecten, 
Salt., Agnostus M‘Coyr?, Salt., and A. princeps ; the Brachiopoda are 


* For list of the species see p. 80, last paragraph in the Lower Lingula-flag 
division. 

+ It is possible that Kjerulf’s Fenestella socialis from Scandinavia may be our 
Dictyonema; Eichwald’s Gorgonia flabelliformis, although much larger, is almost 
identical; Mr. Salter, however, established its generic distinctness. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 83 


Langula pygmeea, Salt., Obolella Salteri, Holl, and Spondylobolus, sp. ; 
many of these are peculiar to the Malvern area. Dr. Holl* in 1865 
described the geological structure of the Malvern Hills and adjacent 
districts, placing these shales in the Upper Cambrian, the Hollybush 
sandstone being the base or lowest known sedimentary rock in the 
Malvern area. Whether Dictyonema should be classed with the 
Bryozoa or Hydrozoa is still a doubtful question. 

Bracurorpops.—Only 5 genera and 8 species occur in the Upper 
Lingula beds—-Kutorgina cingulata, Lingulella Davisii, L. lepis, 
Langula pygmea, Obolella plicata, O. Saltert, O. sagittalis, and Orthis 
lenticularis. 6 pass tothe Tremadoe rocks—Kutorgina cingulata, 
LInngulella Davisti, L. lepis, Obolella plicata, O. sagittalis, Orthis 
lenticularis ; only 2 species are really restricted to the Upper Lin- 
gula-flags, Lingula pygmea and Obolella Saltert. No zoological 
value can therefore be attached to the Brachiopoda, as all the species 
but 2 occur both below and above the Upper Lingula-fiags. 

LAMELLIBRANCHTATA.—None have as yet been detected. 

Gastrropopa.—None known below the Arenig. 

Prrropopa.—Of the 4 genera and 17 species that range from the 
Longmynd group of St. David’s to the Upper Tremadoc no form has 
yet been discovered in the Upper Lingula-flags. 

Hzreropopa.—None have occurred, although 5 species range 
through the Lower Cambrian rocks. Theso-called Middle Lingula- 
flags have yielded one species, Bellerophon cambrensis, Belt, which is 
the first known species on record. 

CrpHatopopa.—None of any type. No species occur below the 
Lower Tremadoc. The entire Cephalopod fauna of the Lower Cam- 
brian rocks consists only of 1 genus and 2 species below the Arenig 
rocks, Orthoceras precow, Salt., and O. sericeum, Salt.; in the Upper 
Tremadoc these increase to 5 species. 

Between the close of the deposition of the Menevian rocks and the 
close of the Upper Lingula-flags, 10 new genera appeared, viz. 3 An- 
nelida, 1 Bryozoon, 4 Crustacea, and 2 Brachiopoda+, succeeded by 
13 new species in the Tremadoc(Hchinodermata 2 species, Crustacea 4, 
Lamellibranchiata 5, Cephalopoda 2 species). The known newly intro- 
duced genera in the Lingula-flags were, Oruziana, Scolicoderma, Hel- 
minthites, Dictyonema, Parabolina, Dikelocephalus, Peltwra, Hymeno- 
caris, Lingula and Kutorgina. Only 1 of these (Lingula) lived on 
after the deposition of those sediments we term Lingula-flags ; this 
genus has appeared in almost every successive marine formation. 

The Lower Lingula beds have yielded 17 genera and 36 species, and 
the Upper Lingula-flags 16 genera and 41 species, showing a generic 
loss or dying out of 7 genera in their upward succession; they ap- 
pear to be Protospongia, Cruziana, Helminthites, Anopolenus, Holo- 
cephalina, Hymenocaris, and Leperditia. 


* Quart. Journ. Geol. Soc. vol. xxi. 1865, pp. 72-102. 

t Cruziana (semiplicata), Scolicoderma (tuberculatum and antiquissimum), 
Helminthites (sp.), Dictyonema (sociale), Hymenocaris (vermicauda), Olenus (cata- 
ractes), Dikelocephalus (celticus and discoidalis), also Carausia (menevensis), with 
Lingula and Linguletla. 


84 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


The two accompanying small Tables (III. & IV.) relate to the dis- 
tribution of the fauna of the Lower and Upper Lingula-fiags, with the 
number of genera and species that pass to the Lower and Upper Tre- 
madoc. Nine whole classes are unrepresented in each division ; only 
2 classes, indeed, in either have real value, the Crustacea and Brachio- 
poda. I regard the Annelida as being wholly doubtful, so far as 
regards species: their presence and influence is manifest every where 
through the slates and flags of the lowest Paleozoic rocks ; but the 
fragmentary condition of the remains, as well as the uncertainty 
relative to their development and history, almost sets aside their 
numerical value as species in evidence. 

In the Lower Lingula-flags the 25 species of Crustacea stand alone ; 
and, with the exception of Hymenocaris vermicauda, Leperditia 
Hicksii, and Primitia solvensis, all are Trilobita. Five species, 
all Trilobites, pass to the Upper Lingula-flags. The Upper Lingula- 
flag table is even more significant ; for every one of the 30 species of 
Crustacea belongs to the group of Trilobita, and only 2 species pass 
to the Upper Tremadoc, Agnostus princeps and Ampyx prenuntius, 
and not one to the Arenig, from the Lingula-flags. 

The left-hand column in all the tables shows the number of genera 
and species that pass up from the next underlying or older formation, 
the upper figure enumerating or expressing the number of genera, 
and the lower denoting the number of species in each class; the 
same plan is appled to the genera and species that pass up into the 
several formations as the case may be, and as expressed in the head- 
ings of the columns. The column headed classes and the two suc- 
ceeding, marked genera and species, speak for themselves. 


Taste 1V.—Upper Lingula-flags. 


cau Pass to | Pass to 
Lower Classes. Genera. | Species. pels: a s be 
ew madoc. | madoc. 
| Plante. 
Protozoa. 
Hydrozoa. 
Actinozoa. 
Echinodermata. 
Ammelicda; Wa osandermascer eee: a: 2 2 
g C@ristacen era.csesecte cect 8 30 4 Z 
SEV OZOAR ind oenatissemein ae 1 1 7 
3 iBrachiopodda):.n.5-ses-5).26ees 9) 8 i 2 
Lamellibranchiata. 
Gasteropoda. 
Pteropoda. 
Heteropoda. 
Cephalopoda. 
Zech br oh gigi apne tee tar 16 41 nie 7 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 8 5 


Lower AND Upper TREMADOC. 


Prior to the year 1865 little or nothing was known of the Tre- 
madoc rocks in the promontory of St. David’s; but in North Wales 
they had been well studied by Sedgwick in 1847, and by Salter in 
1857; and in 1866 Salter and Hicks catalogued all the species 
known up to that time*. Doubtless the Lower Tremadoc series 
are a continuation and natural close of the Upper Lingula-flags or 
Ffestiniog series (the Middle Cambrian of Sedgwick). ‘The strict 
agreement, however, between the North- and South- Wales Tremadoc 
faunas was not thoroughly understood until Dr. Hicks in 1873 care- 
fully described and correlated the paleontological relations that 
exist between the typical North- and South-Welsh areas. The 
researches of Dr. Hicks into the Tremadoc fauna of the mainland 
of St. David’s and that of Ramsey Island made us then familiar 
with the physical relation and paleontological connexion between 
the two areasy. The prior and incomplete observations of Hicks 
and Salter in 1866 at Ramsey Island relative to the presence 
there of the Tremadoc rocks, Dr. Hicks in 1872 fully confirmed, 
being able to show their succession to, and conformable position 
with, the hard siliceous flaggy Lingula-flags. In this research he 
was assisted by three able and competent observers, Messrs. Hom- 
fray, Lightbody, and Hopkinson, the last-named naturalist sub- 
sequently contributing largely to the Arenig Hydrozoa from the 
same area. Dr. Hicks’s labours further resulted in Mr. Davidson’s 
learned paper upon the earliest British Brachiopoda, which con- 
tains a valuable table showing the distribution of all the known 
species of that class in these oldest British rocks (Harlech to Tre- 
madoc). 

Since the labours of Dr. Hicks in Wales, another important ad- 
dition has been made to our knowledge of the distribution and 
fauna of the Lower Tremadoc rocks of the Wrekin area, through the 
careful research of C. Callaway, Hsq., F.G.S. Mr. Callaway cor- 
rected errors made by the Survey, Sir R. Murchison, and Mr. Salter 
in the reading of the Harnage and Shineton beds in Shropshire. He 
described the Lower Paleozoic rocks that range from Wellington 
to Kenley (N.E. to 8.W.). He paid no critical attention to rocks 
newer than the Caradoc; but he greatly changed the previous views 
as to the reading and structure of the county§. Mr. Callaway 
clearly showed that the Shineton Shales are of Lower-Tremadoc 
age, and that the “quartzite” rock that les between the Shine- 
ton Shale and the Wrekin represents the Hollybush Sandstone of 
Malvern, which there underlies the black shales with Olenz, the 
equivalents of the Upper Lingula-flags. The determination by 
Mr. Callaway of the presence of a Tremadoc fauna in England is of 
high importance; and, singularly, most of the Shineton Tremadoc 


* British Association Report, 1866. 

Tt Quart. Journ. Geol. Soc. vol. xxix. p. 39 (1873). 

+ Geol. Mag. vol. v. (1868) pp. 3803-315. 

§ Callaway, Quart. Journ. Geol. Soe. vol. xxxiii. pp. 652-672. 
MOt, XXX VIT. 4 h 


36 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


species are new, and also many of the genera. The species as- 
signed by the author as having most zoological value are Conocoryphe 
monile, Salter, Olenus Saltert, Call., Olenus triarthrus, Call., Agnostus 
dux, Call., Lingulella Nicholsoni, Call., Obolella sabrine, Call., 
Asaphus (Asaphellus) Homfrayi, Salt., Asaphus (Platypeltis) Crofti, 
Call. 

Mr. Callaway also draws comparisons between the Shineton, 
Pedwardine, and Malvern series, showing their relations and affinities. 
All the species in the Shineton Shales are new except three, viz. 
Conocoryphe monile, Salt., Asaphus (Asaphellus) Homfrayi, and 
Dictyonema sociale; the fauna is therefore nearly unique. Mr. 
Callaway suggests that the Shineton Shales may be passage-beds 
between the Lingula-flags and the Lower Tremadoc: the presence 
of Dictyonema sociale strengthens this view; it is an Upper Lin- 
gula-flag species at Malvern, Pedwardine, and North Wales in 
many places, also a well-known Tremadoc form in the latter 
area. Mr. Callaway ably discusses the physical features of the 
Shineton area. He describes fourteen new species, and establishes 
two subgenera of Trilobites, Asaphellus and Platypeltis, and two 
genera, Conophrys and Lichapyge ; he also determines and describes 
a remarkable Cystidean, possessing a slender crinoidal-like stem and 
simple calicular pinnule, which he names Macrocystella Marie. At 
present we know of no true Tremadoc rocks in the Stiperstones area ; 
nor, indeed, are we sure of the presence of the true Lingula-flags in 
that district west of the Longmynds, unless the Stiperstones them- 
selves represent them; but of this we have no organic proof, the 
series succeeding being Arenig or Skiddaw, formerly mapped and 
named Lower Llandeilo by Sir Roderick Murchison and the Survey. 
Mr. Callaway believes from recent research that the quartzites of 
the Stiperstones are of Arenig age, and therefore distinct from the 
quartzites of the Wrekin, deducing his opinion from the fact that 
when they are conglomeratic the pebbles in the Arenig quartzites 
are mainly quartz, while the included fragments of the Wrekin 
quartz rocks are felsitic. Again, Mr. Callaway believes that he has 
detected the Shineton Shales (Lower Tremadoc) at the base of the 
escarpment of the Stiperstones dipping under the Arenig series in 
the same direction as the rocks of that formation, 2.e. W.N.W. 

There is little or no specific paleontological affinity or com- 
munity between the Tremadoc and Arenig groups in any known 
area in North or South Wales, although they are everywhere con- 
formable. No Tremadoc rocks have yet been proved to occur above 
the Lingula-flags in the passes of Llanberris and Nant Francon or 
the adjoining regions, where the latter rocks are so finely and fully 
developed ; neither is it yet quite certain that they occur in An- 
glesey. <A few fossils lately found by Professor T. M‘K. Hughes 
lead to a strong suspicion that the northern half of the island may 
yet yield a Tremadoc fauna. The singular Trilobite, Neseuretus 
ramseyensis, Hicks, found by Professor Hughes at Ty-hen near Llan- 
erchymaedd, has not hitherto occurred in North Wales, but only on 
Ramsey Island and at Tremanhire (St. David’s); and other remains 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 87 


occur with this form, furnishing strong evidence that the Tremadoc 
rocks extend as far N.W. in Wales as Anglesey: the above-men- 
tioned species has not occurred in the classical Tremadoc area, either 
in the lower or the upper division of the group. The presence of 
the Arenig beds in Anglesey favours the belief that the Tremadoc 
may also occur in the island. 

The Tremadoce slates, now so well known in the two classical 
Welsh localities, viz. Tremadoc in North, and St. David’s in 
South Wales, are so rich in conspicuous forms and well-determined 
species that we readily recognize and place them in two distinct or 
separate horizons or zones. ‘heir true position in North Wales, 
between the Lingula-flags below and the Arenig group above, was 
clearly shown and fixed by the distinguished Cambridge Professor 
in 1851. Their base was an easier line to trace than their passage 
above into what was then a group of rocks believed to be almost 
unfossiliferous. Small indeed was the fauna known to Sedgwick 
and Salter in 1843, when, at Tai-hirion, west of Arenig Mountain, 
they collected the two characteristic fossils Ogygia Selwyn and 
Calymene parvifrons. These two species formed the mental basis 
of Sedgwick’s Arenig, and determined the top of his Tremadoc 
group till better worked out. Now the Tremadoc fauna of North 
Wales consists of 42 species, of South Wales 30, and of Shropshire 
(Shineton) 16. The Arenig group numbers 62 genera and 149 
species; or 97 species occur at St. David’s, 74 in the Skiddaw beds 
of Westmoreland, 36 in the Stiper-stones area, 34 in North Wales. 
These have nearly all been described since the far-seeing eye of 
Sedgwick determined the stratigraphical place of the groups on 
physical grounds. We owe the description of the species to Dr. Hicks, 
Mr. Salter, Professor M‘Coy, and Messrs. Hopkinson, Lapworth, 
Nicholson, and Davidson. Guided by mineral or physical characters 
alone, the line of separation between the Lingula-flags and Lower 
Tremadoc would probably be drawn immediately above the hard 
grey arenaceous so-called Middle Lingula-flags, in the typical Tre- 
madoe area, where Lingulella Davisit ceases to be an abundant and 
characteristic fossil ; but zoological conditions forbid this ; the ‘ black 
slates’ that succeed the “hard grey series,” possess a recurrent 
generic fauna from the lower zones, including Conocoryphe depressa, 
Dikelocephalus?, and Agnostus princeps, with Orthis lenticularis, 
the two latter in millions. The cliff of Ogof-ddu, near Criccieth, 
and the jutting promontory of Craig-ddu exhibit a complete section 
from the Lingula-flags to the Upper Tremadoc slates, the character- 
istic Orthis lenticularis marking by myriads the junction of the 
Middle Lingula-fiags and the Upper Black pyritous slate. No Upper 
Lingula-flag section in Wales is more prolific in life: Conocoryphe 
abdita, Salt., and two other species, Dikelocephalus celticus, Salt., 
D. discoidalis, Salt., Agnostus princeps, and Orthis lenticularis 
dispel all doubts as to age or horizon; but the Tremadoc fossils do 
not occur at Ogof-ddu; the fossiliferous zones must be sought for in 
the sections at Penmorfa, Borthwood, Moel-y-gest, dc. near Port- 
madoc (places rich in characteristic species), and also across the 
estuary from Aber-ai, by Dudreath, to Cae-lago, near Maentwrog. 

h2 


88 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


These are the chief localities whence the history of the Tre- 
madoe rocks of North Wales has been drawn and deduced, from 
which also their remarkable fauna has been disentombed, and that 
mainly through the instrumentality of Mr. David Homfray and Mr. 
Ash of Portmadoe, the late Mr. Salter, and the old collector for the 
Survey, Mr. Gibbs. 

The careful and thorough examination of North and South Wales 
that has of late been made by so many able observers, has clearly 
shown the perfect sequence and agreement in stratigraphical suc- 
cession between the whole of the groups ranging from the Har- 
lech and Longmynd strata to the Llandeilo; this is complete for 
South Wales in the St.-David’s promontory as one continuous and 
ascending section, with no break in continuity. It is not so clear 
or complete a sequence in North Wales, where the chief localities 
for the Tremadoc and Arenig groups are more widely separated, 
both formations being only developed in two counties in North 
Wales, viz. Merionethshire and Caernarvonshire—the Lower Tre- 
madoce series chiefly at Penmorfa, Borthwood, and Llanerch in 
Caernarvonshire, and in Merionethshire at Cae lago near Maentw- 
rog and at Tyn-y-llan. The Upper Tremadoc rocks, on the contrary, 
are best developed in Merionethshire at Tuhunt-y-bwlch, Penclo- 
gwyn, and Garth, whilst Moel-y-gest is the only locality in Caernar- 
vonshire where they occur. The Arenigs, which also occur only in 
the same counties, I shall have occasion to refer to at considerable 
length when speaking of the distribution of the species. 

The discovery near St. David’s, by Hicks and Salter, of Tremadoc 
slates equivalent to those of North Wales threw new light upon the 
history of that remarkable promontory. Not only are these slates 
the same as those of Merioneth and Caernarvonshire, but a large 
number of additional species of much interest have been added to the 
whole fauna. In 1866* Messrs. Hicks and Salter published the results _ 
of their researches at St. David’s, giving a list of the species dis- 
covered by them. Since then Dr. Hicks has reaped a large harvest 
of new forms from these rocks, thus adding to the history and 
paleontology of that area—no less than thirty species having 
occurred to him. These were obtained from three distinct loca- 
lities—the first on the north-eastern coast of Ramsey Island ; 
the second at the northern extremity of Whitesand Bay, and ex- 
tending 4 miles in a north-easterly direction beneath the Arenig 
rocks; the third at Tremanhire, an extensive area or tract 5 miles 
east of St. David’s, thus showing extensive geographical distri- 
bution. The Ramsey-Island section is stratigraphically complete, 
having for its conformable base the Lingula-flags full of Linguiella 
Davisit ; and upon the intervening Tremadoc rocks, here 1000 feet 
thick, succeed the Arenigs, thus here, as at Whitesand Bay, affording 
clear proof of the intercalated position of the Tremadoc group. In 
North Wales no Arenig rocks are known to rest directly upon the 
Tremadoc anywhere. Dr. Hicks obtained in all 29 species from 
the Ramsey-Island section, 24 of which were new; and 15 occurred 
at Tremanhire, inland, east of St. David’s. 

* Brit. Assoc. Report, 1866, p. 182. 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 89 


The Tremadoc rocks of Whitesand Bay also rest conformably 
upon the Lingula-flags, and underlie the Arenigs, as at Ramsey 
Island; they strike north-east for 4 miles towards Abereiddy Bay. 
The third exposure, at Tremanhire, shows the same stratigraphical 
relations and succession, and has yielded fifteen out of the 30 
new species described by Hicks as occurring at Ramsey Island. 
This patch is now an isolated inlier surrounded by Lingula-flags and 
Menevian beds, and with a large area of Longmynd rocks to the west. 
The Pre-Cambrian ridge and Longmynd series now divide or sepa- 
rate the two Tremadoc areas, which must have been once conti- 
nuous. Dr. Hicks believes that the conditions under which these 
rocks at St. David’s were deposited were intermediate between those 
of the shoal and shallow water in which the Lingula-flags were 
deposited, and those of the deeper sea from which the finer muddy 
deposits of the Arenig slates were thrown down, and that this was 
probably one of the causes of the appearance in them of so varied a 
group of organisms. 

There is some difficulty in comparing the Tremadoc rocks of 
North with those of South Wales. At St. David’s they are so inti- 
mately connected with the underlying Lingula-flags and overlying 
Arenig series that the boundary-line is almost arbitrary. Palseon- 
tologically they appear to be on the same horizon as the Lower 
Tremadoc series of North Wales, the chief trilobite Niobe Homfrayt 
with Lingulella Davisti and L. lepis connecting them. Yet the 
mass of the faunas are entirely distinct, for out of the 42 North- 
Wales and 30 South-Wales forms only the 3 above named are com- 
mon to both. 

Thus out of the 4 North-Wales and 7 South-Wales Trilobita 
only 1, Mobe Homfrayi, connects the two areas. Of the 4 
genera and 9 species of Brachiopoda occurring in both areas only 
2 species connect them, viz. Lingulella Davisii and L. lepis; and 
more remarkable still is the occurrence in Ramsey Island of the 
earliest known Lamellibranchs in Britain, if not in Hurope. The 
Lower Tremadoc beds in that island have yielded 5 genera and 
12 species, the first representatives known of that order; 6 of 
the same species occur inland at Tremanhire. ‘Three of these genera 
belong to the Arcide, and one probably to the Anatinide ; prior to 
this discovery Jtedonia, Ribieria, and a species of Palearca were the 
oldest forms known, and were obtained from the Arenig beds at Lord’s 
Hill, Shelve. The last-named genus has been extracted from Lower 
Silurian rocks in Spain, Bohemia, France, and Devonshire. These 
bivalves are also associated with the earliest known Crinoid and 
Starfish, Dendrocrinus cambrensis and Paleasterina ramseyensis. Still 
more significant zoologically is the presence of a new genus with 
four species of Trilobita, which Dr. Hicks has named Neseuretus. 
This large and singular genus seems to have affinity with Drke- 
locephalus through its pygidium, with Calymene and Homalonotus 
through the thoracic segments, and with the Conocephalide 
through the glabella. The four species and one variety are all 
confined to the Lower Tremadoc rocks of Ramsey Island and 
Tremanhire on the St.-David’s promontory. None had occurred in 


go PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


North Wales until Prof. Hughes discovered portions of one in Angle- 
sey. If any other Trilobite were required to establish paleontological 
relationship between the Tremadoc rocks of North and South Wales 
it would be Angelin’s genus WMiobe ; and Dr. Hicks has obtained, both 
from Ramsey Island and Tremanhire, a new and splendid species 
(Niobe menaprensis), greatly exceeding in size the characteristic 
Niobe Homfrayt from Penmorfa and Tremadoc. Niobe is not 
known out of the Tremadoc group. The South-Wales beds do not 
possess so numerous an assemblage of pelagic Mollusca as those of 
North Wales: 9 species of Pteropoda (Theca Davisii, T. 
opercula, Lower Tremadoc only, 7. arata, T. bijugosa, T. cuspi- 
data, Lower and Upper, and 7’. alata, T. simplex, and T’ tri- 
lineata, with Conularia Homfrayr) are Upper Tremadoc; and 5 
Heteropoda (Bellerophon arfonensis, B multistriatus, B. ramsey- 
ensis, B. solvensis, and B. shinetonensis) range through the Trema- 
docs; but Theca Davisii, Bellerophon ramseyensis, Hicks, and B. 
solvensis, Hicks, are South-Wales species, the mass occurring in the 
typical Tremadoc area in North Wales. Associated with Vzobe 
menaprensis and the Nesewretr no less than 19 species occur, 
including the 12 Lamellibranchs and 5 Brachiopoda, viz. Lin- 
gulella Davisu, Lingula petalon, Obolella pheata, Orthis Carausit, 
and O. menapiw, 29 of the 31 being described by Hicks. 
Dr. Hicks believes that the lower portion of the St.-David’s 
Tremadoc rocks and the upper black slates of the Lingula-flags of 
North Wales were deposited contemporaneously. This may well be, 
looking at the remarkable difference existing in the faunas of the 
two areas, so few species being common to or uniting the two groups, 
and difference of depth at the time of deposition. Again, the 
absence of the fine black slates in the Upper Lingula-flags of St. ° 
David’s, so characteristic of the North-Wales and the Malvern beds, 
may be due to depth, shallow in one region and deep in the other. 
The Tremadoc series is divided into three groups by Dr. Hicks, 
the lowest at St. David’s consisting of grey, fissile, or flaggy sand- 
stones. Exposed at Ramsey Island, Tremanhire, and Llanveran, a 
peculiar assemblage of fossils occurs in this area, new and different 
from that of North Wales:—Davidia, Glyptarca, and Palcearca, 
being the earliest bivalves known; the earliest Crinoid, Dendrocrinus 
(unless Mr. Callaway’s genus Macrostella from the Shineton area 
should be earlier in time); and the first known Asteroid, Paleasterina 
ramseyensis, Hicks. The Middle Tremadoc of Hicks is equivalent to 
the passage-beds between the Lower and Upper series of Salter, con- 
taining Cheirurus Frederici, Conocoryphe verisimilis, C. vewata, and 
Asaphus Homfrayi,—the upper member, consisting of iron-stained 
slates and flags at Penclogwyn, Garth, Dudreath Tuhwnt-yr- 
bwlch, &c., in the Portmadoc area being characterized by Angelina 
Sedgwickit, Lingulocaris lingulecomes, and Conularia Homfrayt. 
The distinctness of the Tremadoc from the Lingula-flags below is in 
all areas the same; only 8 species out of 65 known forms in the 
former pass up to the Tremadoc, which possesses a fauna of 34 
species; these are Agnostus princeps, Ampyw prenuntius, Conocoryphe 
depressa, Lingulella Davisit, L. lepis, Orthis lenticularis, Obolella 


ANNIVERSARY ADDRESS OF THE PRESIDENT. gt 


sagittalis, and Diciyonema sociale. 42 species are known in North 
Wales, and 30 in South Wales; yet, as before stated, only 3 species 
are common to both districts—Mobe Homfrayi, Lingulella Davisi, 
and ZL, lepis. 

It must be observed that no species of true Graptolite is known 
in the Tremadoc rocks, none being really known below the Arenig 
in Britain, Europe, or America. Doubtless the whole Arenig series 
is more closely allied to the succeeding and overlying Llandeilo than 
to the underiying Tremadoc; it is certainly so as exhibited in the 
South-Wales promontory, where both groups are typically developed 
and succeed each other in one continuous and conformable section ; 
yet there is perhaps no greater change or paleontological break be- 
tween any two conformable British groups than between the 
Tremadoc and Arenig, the rich Hydrozoal fauna coming in at the 
close of the former and commencement of the latter. The change 
in the physical conditions of the sea and sea-bed at the close of the 
Tremadoc period was favourable to the development of this division 
of the Coelenterata. 

The most characteristic of all the fossils of the Lower Tremadoc 
are Miobe Homfrayi, N. menapiensis, Psilocephalus wnnotatus, with 
Angelina Sedgwicku and <Asaphus affinis in the Upper Tremadoc; 
these five species are met with wherever the Tremadoc rocks are 
well seen. <Agnostus princeps, so abundant in the Upper Lingula- 
flags of North Wales, is not known in the St.-David’s Tremadoc 
sections, and is of the rarest occurrence in the North-Welsh classical 
localities ; with Agnostus the characteristic Lingulella (L. Davisir) 
dwindles away, and seems to be replaced by the small but equally 
gregarious L. lepis. 

The true Upper Tremadoc series is characterized by many new 
species. Niobe Homfrayt, Psilocephalus innotatus, and P. inflatus 
no longer exist as species, but are replaced by Asaphus Homfrayt 
and Angelina Sedgwicki; Cherurus Predericc at Garth, Llanerch, 
Penclogwyn, Portmadoc, &c., ranges from the top of the Lower, 
through the passage-beds, and all through the Upper Tremadoe into 
the Arenig. Olenus impar adds another to the characteristic species 
of the Upper Tremadoc; and Cyrtoceras precox, which occurs at 
Llanerch, is the earhest knewn Cephalopod in the British rocks. 
The first Orthoceras (O. sericeum) known* also occurs in the passage- 
beds at Garth and Tuhwnt-yr-bwlch. 

It is clear that there is no stratigraphical or paleontological 
break between the Lingula-flags and the Tremadoc: both areas 
prove the former by direct evidence; and 9 species connect them 
paleontologically, viz. Dictyonema sociale, Salt., Agnostus princeps, 
Salt.. Ampyw prenuntius, Salt., Conocoryphe depressa, Salt., ? Olenus 
ampar, Salt., Lingulellu Davis, M‘Coy, L. lepis, Salt., Obolella sagit- 
talis?, Salt., Orthis lentrcularis, Wahl. Three of these same species 
also connect the Tremadoe and Arenig: they are Lingulella Da- 
visit, L. lepis, and Orthis lenticularis. All paleeontological evidence 
goes to prove “ that the Tremadoc rocks of the St.-David’s area are 
closely allied to, if not identical with, the lower portion of the 

* Originally in the cabinet of Mr. D. Homfray, of Portmadoc, 


g2 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Tremadoc rocks of North Wales.” Mr. Homfray, of Portmadoe, than 
whom no one has had greater experience or more carefully studied 
the Tremadoc rocks in their original localities, finds no difficulty in 
recognizing and correlating the rocks and fossils of the two districts, 
affirming the St.-David’s beds to be the equivalents of the lower 
portion of the series in the Portmadoc region. Mr. Homfray is 
also of opinion that the Upper Tremadoc rocks are represented at 
St. David’s by part of the Arenigs, which contain several Upper- 
Tremadoce fossils, in addition to the rich fauna of Graptolites (42 
species) now known to occur through the entire Arenig series. 
The whole Tremadoc fauna consists of 33 genera and 86 species, 
and no Hydrozoa; the Arenig includes 62 genera and 149 species, 
with 42 species of Hydrozoa; the succeeding Llandeilo has 230 
species, 90 of which are also Hydrozoa; yet in each formation the 
Graptolites appear mostly to be of distinct species. They also play 
an important part in the faunas of the Llandeilo, Caradoc, and 
Llandovery rocks of Scotland. 

Table VII. (p. t00) is a complete numerical analysis of the 
fossils in the Lower Cambrian rocks, from the Harlech and Long- 
mynd to the close of the Upper Tremadoc, in which it will be seen 
that the twelve classes (and Plante?) are represented by 61 genera 
and 182 species, as the known commencement of life in the British 
area. They are as follows :— 


The Longmynd group contains 18 genera and 33 known species. 


The Menevian _,, - 24 5 Sy Ba a 
The Lower Lingula-flags ,, 19 Bs SS) hie P 
ThevWippers. 7s, va moae 12 45 AQ Fie be 
The Lower Tremadoc _,, 26 . 515 Na Were “1 
The Upper * # 21 $3 34 Ci, an 


11 genera and 16 species pass to the Arenig, this small number 
clearly determining the individuality of the Lower Cambrian fauna. 
It is only through the 6 species of Crustacea out of 103, and the 
6 Brachiopoda out of 20, that the passage-forms are thus numerous. 
The remaining 3 species are pelagic, 1 Pteropod, 1 Heteropod, and 
1 Cephalopod. 

Lower TREMADOC. 

Pranrxz.—None. ProtozoA.—None known. 

Hyprozoa.— Dr. Callaway discovered in the Shineton Tremadoe 
rocks two species previously unknown. One belongs to the genus 
Clonograptus of Hall; the other is the Bryograptus Callavei of 
Lapworth, both belonging to the family Dichograptidee. These are 
the oldest Rhabdophora known. 

Actinozoa.—None, so far as we know, below the Llandeilo rocks. | 
EcHINODERMATA. Wind ocrinus Hann ensis, Hicks, and Paleaste- 
rina ramseyensis, Hicks, are both from the Lower Tremadoc beds 
of Ramsey Island, St. David's ; they are the first known species of 
the orders Crinoidea and Asteroidea; and they stand with numerous 
other discoveries as a testimony of the value of the researches of 
Dr. Hicks in these early Cambrian rocks, to which Palzeozoic palee- 

ontology owes so much. 

ANNELIDA.—None, either in the rocks of South or North Wales. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 93 


Crustacea.—Only 2 new genera appear with the coming-in of 
the Lower Tremadoe (Psilocephalus and Neseuretus); but the whole 
Crustacean fauna comprises 13 genera and 24 species, of which 10 
genera and 19 species especially characterize the Lower Tremadoc. 
They are so essential to a right understanding of the Lower Tremadoc 
rocks and their entire distinctness from the Upper Lingula-flags that 
tT enumerate them :—Agnostus Barlowu, Belt; A. dua, Call. ; Cono- 
phrys salopiensis, Call.; Asaphus Croft, Call.; Diomde atra, Salt. ; 
Lichapyge cuspidata, Call. ; Neseuretus (5 species, all by Dr. Hicks) ; 
Niobe Homfray?, Salt.; N.menaprensis, Hicks; NV. solvensis, Hicks ; 
Conocoryphe verisimilis, Sait.; Olenus triarthrus, Call.; O. Salter, 
Call. ; Psilocephalus inflatus, Hicks ; and P. ennotatus, Salt. None of 
the above species occurs below or above the Lower Tremadoc horizon. 
The remaining five occur also in the Upper Lingula-flags, and unite the 
two formations. The chief species named occur at Penmorfa, Borth- 
wood, and Llanerch in Caernarvonshire, and Dudreath and Tyn-y- 
ilan in Merioneth; and the same species occur in the St.-David’s 
promontory at Whitesand Bay, &c. Nowhere do the species above 
mentioned transgress the lower beds; the Lower and Upper Tre- 
madoc are connected only by Agnosius princeps, Ampyx preenun- 
tius, and Ogyqua scutatriv. Nothing that I could adduce would be 
stronger evidence of the value of the Lower Tremadoc as a well- 
defined zoological group: it contains more genera than any other 
division of the Lower Cambrian stages ; and the species are essen- 
tially characteristic. 

Bryozoa.—Dictyonema sociale, Salt., as in the Upper Lingula- 
flags. This genus occurs near Tremadoc in the passage-beds. 

Bracutopopa.—The Lower Tremadoc Brachiopoda are Kutorgina 
emgulata, Lingula petalon, Lingulella Nicholson, L. lepis, Obolella 
Beltu, O. sabrine, O. plicata, and O. sagittalis. Orthis carausit, 
O. lentecularis, and O. menapie. 4 of these 5 genera and 6 of the 
11 species pass to the Upper Tremadoc. ‘They are Lingula petalon, 
Lingulella lepis, Obolella Belin, O. plicata, Orthis carausw, and 
O. lenticularis. 4 genera and 5 species appear from the Upper Lin- 
gula flags, so that the true Lower Tremadoc species are few, and only 
2 species are peculiar, viz. Lingulella Nicholsont and Opolella sabrine. 

LaMeELLiBRANcHIATA.—The first appearance of this class, or the 
earliest known in the British rocks. Dr. Hicks, through his 
researches upon the Tremadoc rocks of Ramsey Island, has obtained 
these first evidences of Pelecypod or bivalve molluscan life. He 
refers them (and, I think, rightly) to 5 genera and 12 species— 
Dandia (2), Glyptarca (2), Modiolopsis (4), Palearca (2), and 
Cicenodonta (2); none of the species is known out of or above 
the Lower Tremadoc. 

GastERoPpopA.—None known below the Arenig rocks. 

PrEeropopa.—The genus Theca is represented by 5 species in the 
Lower Tremadoe, viz. Theca ovata, Salt., T. bijugosa, Salt., 7. cuspi- 
data, T. Davisw, and T. (Cleidotheca) operculata. The 3 first 
named pass to the Upper Tremadoc. 7. Davisz is the only species 
in the Tremadoe rocks of St. David’s. The remaining 4 are North- 
Welsh Tremadoc species. 


94. PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Herrroropa.—bellerophon ramseyensis, Hicks, B. solvensis, Hicks, 
and B. shinetonensis, Call. ; the former both Menapian forms, cha- 
racterizing the Tremanhire or Solva and Ramsey-Island beds, St. 
David's. 

Crpnatopopa.—None below the Upper Tremadoc. 

The analytical Table No. V. shows that 28 genera and 58 species 
occur in the Lower Tremadoc beds of North and South Wales. Three 
classes only are of numerical importance :—the Crustacea with 13 
genera and 24 species: the Brachiopoda, with 5 genera and 11 
species; the Lamellibranchiata, 5 genera and 12 species. These 
3 groups constitute the mass of the known genera and species, or 
23 out of 29 genera and 47 out of 59 species. 


Taste V.—Lower Tremadoe. 


ee rom Pass to 
Uopes Cl Genera. | Species. | Uppe 
Lingula- asses, enera. | Species. | Upper 
flags. Tremadoc. 
| Plante. 
Protozoa. 
Hydrozoa. 
Actinozoa. 
Hehinodermata...... 2 2 | 
Annelida.., 
5 Crustacea) Wi... sas: 18 24 x 
i Bryozoa. if 1 
3 Brachiopoda ......... 5 ll 4 
Lamellibranchiata... +5) 12 
Gasteropoda. 
Pieropoda: fs... 1 5 x 
Heteropoda ......... I 3 
| Cephalopoda. 
WT Meecéusbhooacoon 28 58 8 


Uprer Tremapoc. 

Praniva.—None. 

Prorozoa.—None. 

Hyprozoa.—None. 

EcuinopERMATA.—None. 

ANNELIDA.—Tracks only. 

Crusracra.—1l1 genera and 15 species constitute the Crusta- 
cean fauna of the Upper Tremadoc; 12 of the 15 species are Tri- 
lobita ; the remaining 3, Ceratiocaris (2 species) and Lingulocaris, 
are Phyllopoda, and the first of this order known in the British rocks. 
Asaphus affinis, M‘Coy, A. Homfrayi, Salt.,Cheirurus Frederici, Salt., 
Dionide atra, Salt., Ogygia scutatria, Salt., and O. Selwyni, Salt., 
pass to the Arenig, and, with 6 Brachiopoda and 3 Pteropoda, 
constitute the 13 uniting forms out of a total of 182 species in the 
Lower Cambrian rocks and 150 in the Arenig. 

The following 5 Crustacea only are confined to the Upper Tremadoe 
— Angelina Sedgwickii, Conocoryphe olenoides, Olenus ampar, Ceratio- 

ris laius, and Lingulocaris lingulocomes. Lam careful to name these 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 95 


as the first zoological or paleontological break between the Lower and 
Upper Cambrian, or the Cambrian and Silurian, takes place here, 
and that without any stratigraphical or physical ‘unconformity. No 
greater faunal change is known to take place through the whole of 
the Paleozoic rocks than at the close of the Tremadoc or Lower 
Cambrian. 

Bryozoa.—None known. 

Bracutorpopa.—Only 4 genera and 7 species occur ; and 6 of these 
pass to the Arenig. ‘They have therefore no stratigraphical value. 
Obolella Belti is the single remaining or restricted form. 

LAMELLIBRANCHIATA.—None yet known. 

GastrRopopA.—None have yet occurred below the Arenig. 

Prerovopa.—Conularia and Theca—the first with 1 species, C. 
Homfray?, which is also Arenig ; the latter contains 6, one of which 
(Theca simplex) also passes to the Arenig. 

HerEeropopa.— Bellerophon arfonensis, Salt., and B. muliistriatus, 
Salt., are the only Upper-Iremadoc species. The last-named spe- 
cies is Arenig also. 

CupHaLopopa.—Cyrtoceras precox, Salt., and Orthoceras sericeum, 
Salt.; the last also passes to the Arenig; they are the first Cephalo- 
pods known ; and it is doubtful if Cyrtoceras precox is not Lower 
Tremadoc also. The rich locality Garth yields O. sericewm from the 
upper part of the Upper Tremadoc. The Arenig of Llanvirn at 
St. David’s has yielded the same; it is therefore both a North- 
and South-Wales species, from the Tremadoc in one area, the Arenig 
in the other. 


TasLe VI.— Upper Tremadoc. 


From 
Lower Classes. Genera. | Species. 
Tremadoc. 


———— | | 


Plante. 


Pass to 
Arenig. 


Protozoa. 
Hydrozoa. 
Actinozoa. 
Echinodermata. 
Annelida. 
‘| Crustacea ....ss..06+- ll 15 


| 
| 
Bryozoa. 


Pe 


uN 
J 


op 


Brachiopoda ......... 
Lamellibranchiata. 
Gasteropoda. 


Pheropods -2203.... 52: 2 C 


Qe 


Heteropoda ........ e 1 
Cephalopoda......... 2 


96 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Description of the first six Tables. 


I have in this oldest Cambrian series (from the Longmynd rocks 
to the Upper Tremadoc) given an analytical table for each of the 
6 formations or groups of strata. They cannot be otherwise tho- 
roughly comprehended, taken as a whole, without wearisome de- 
scription, owing to the non-representation of some classes, and - 
the unequal representation and distribution of others. Neverthe- 
less I have constructed Table No. VII. for the whole, to show 
the unequal distribution, the Table delineates every thing, its 
accuracy being tested through the six smaller tables (Nos. I. to 
VI.), wherein are given the analyses of all the groups sepa- 
rately. More than this, these six small tables enable the student 
to see immediately the numerical value of each division, and the 
number of species that pass up or live on to the succeeding period 
or formation. Again, there is much difficulty in following some 
authors through the many subdivisions and local names adopted by 
them for the strata between the Longmynd group and the close 
of the Upper Tremadoc. The succeeding Arenig, in which some of 
the classes are largely represented, while the formation is widely dif- 
fused geographically, enables me to show the distribution in one 
Table (No. VIII.); the same is also done for every succeeding epoch. 

Tur Lonemynp anp Hartecs Grovrs.—Table I. enumerates 
the fauna of the Longmynd and Harlech beds only, chiefly those 
of St. David’s, and the first traces of life in the British Islands. 
In this Table 8 if not 9 of the 14 classes have no representa-— 
tives, no species of them having as yet occurred; and, with the 
exception of the Crustacea, the remaining 5 classes are but feebly 
illustrated. This Table, however, at once lays bare the fact that we 
are not tabulating the commencement of life even in our own area; 
both the rock masses and the fossils that do occur, notably the great 
Paradoxides, bid us search still deeper in time for their ancestors as 
well as those of the pelagic Mollusca, as also of the Brachiopoda 
(Lingule, Obolelle, and others). All these and the Lyssakine hexac- 
tinellid sponges had their progenitors, which we have yet to deter- 
mine. I doubt not that much light will yet be thrown upon them. 
through patient research. 

This first Table shows that the Longmynd and Harlech rocks are 
known at the present time to possess 18 genera and 33 species—- 
the commencement of life, so far as we know. The Crustacea are 
the most abundant class; 7 genera and 14 species occur: these 
earliest rocks of St. David’s yield Agnostus cambrensis, Hicks, 
Conocoryphe bufo, Hicks, C. Lyellii, Hicks, C. solvensis, Hicks, 
Microdiscus sculptus, Hicks, Paradowides aurora, Salt., P. Hicksi, 
Salt., P. Harknessti, Hicks, P. solvensis, Hicks, MS., Plutonia Sedq- 
wickii, Hicks, and the Longmynd form Paleopyge Ramsayii, Salt. ; 
the Ostracoda are Leperditia ferruginea, L. cambrensis, Hicks, and 
L. primeva. 4 of the above species pass to the Menevian—Agnostus 
cambrensis, Conocoryphe bufo, Paradoxides aurora, and P. Hacksu. 
Out of the 6 species of Brachiopoda, 4 also pass to the Menevian— 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 97 


Lingulella ferruginea, Salt., and var. ovalis, Hicks, Orthis sagittalis, 
and Discina pileolus. The special or restricted forms are only 2— 
Discina caerfaiensis, Hicks, MS., and Lingulella primeva, Hicks. 
The Plantz ? and Protozoa are mentioned under those heads in the 
analysis. The table shows that 8 out of the 14 classes are not repre- 
sented in the Harlech group. That they will remain so under re- 
newed search and scrutiny I much doubt. Hach year adds some 
new form to these once believed to be unfossiliferous strata. North 
Wales has yet to yield up from these rocks a fauna equal to that 
of South Wales. 

Menevian.—Table IT. gives us all that is known of the Menevian 
fauna and its relation to the Harlech series and the succeeding 
Lower Lingula-flags, of which, indeed, it is but the base; for out 
of the 24 genera and 51 species in the Mevevian, 13 of the former 
and 19 of the latter are common to the two. Of the Crustacea 8 of 
the 12 known genera, including 14 species, are the same—viz. 
Agnostus (A. Davidis, A. scutahs), Anopolenus (A. Henrici, A. Sal- 
teri), Conocoryphe (C. applanata, C. humerosa, C. variolaris), Holo- 
cephalina (H. primordialis), Leperditia (L. Hicksir), Primitia (P. 
solvensis), Microdiscus (M. punctatus), Paradowides (P. aurora, 
P. Davidis, and P. Hicksu); and 3 of the 4 genera of Brachiopoda, 
Lingulella (L. ferruginea), Obolella (O. maculata), and Orthis (O. 
sagittalis). Of the 6 species of Pteropoda, only 1 is common to the 
2 groups, namely Theca corrugata. The Menevian is united to 
the Harlech and Longmynd rocks through 8 genera and 12 species. 
The Crustacea by 3 genera with 4 species, viz. Agnostus (cam- 
brensis), Conocoryphe (bufo), Paradoxes (P. aurora and P. Hicksir) ; 
and the Brachiopoda the same, Discina (pileolus), Lingulella ( fer- 
ruginea and var. ovalis), and Orthis (sagittalis). The Lyssakine 
sponge (Protospongia) and Theca penultima complete the alliance. 
- 8 of the 14 classes are not represented. The first Cystidean, 
Protocystites menevensis, Hicks, occurs in the Meneyians of St. David’s. 
11 new genera (not known in the Harlech group) first occurred in 
the Meneyian sea ; 6 are Crustacea, 2 Pteropoda, 1 Cystidean, 1 An- 
nelide, and 1 Brachiopod? No species pass from the Menevian to 
the Middle or Upper Lingula-flags. 

Lower Lineuna-riaes.—Table III. But for the Crustacea, the 
Lower Lingula-flags would have no value as a subformation ; they 
are but the upper, continuous development of the Menevian. Only 
one genus of Crustacea, the Phyllopod Hymenocaris, is new to the 
Lower Lingula-flags ; the remaining 8 are also Menevian; and 14 of 
the 25 crustacean species also occur in the Meneyian below. Still 
further, of the 17 known Lower Lingula-flag genera, 13 are Mene- 
vian ; and this close association is more marked still from the fact 
that only 2 species pass to the Middle and 8 to the Upper Lingula- 
beds; so that there is a greater difference between the Lower and 
Upper Lingula-beds themselves than between the former and the 
underlying Menevian. The Upper Lingula-flags, through their 
Crustacea, 8 genera and 30 species, stand alone. 

Urrrr Linevra-riacs.—Table LV. Out of a community of 17 


98 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


genera and 36 species in the Lower, and 16 genera and 41 species in 
the Upper Lingula-fiags, only 8 genera and 8 species unite the two 
groups: they are 5 Trilobita and 3 Brachiopoda—namely Anopolenus 
Saltert, Hicks, Microdzscus punctatus, Salt., Agnostus princeps, Salt., 
Paradoaides Hicksir, Salt., P. aurora, Salt., Lingulella Davisii, L. 
lepis, and Orthis sagzitalis. Only 4 classes are represented in the 
Upper Lingula-flags, the Annelida, Crustacea, Bryozoa, and Brachio- 
poda. The remaining 10 have no representative whatever. 16 
genera and 41 species occur in the Upper Lingula-beds ; 8 genera and 
30 species are Crustacea, and 5 genera and 8 species Brachiopoda ; 
there are 2 Annelida, and 1 Bryozoon, Dictyonema. 9 genera and 
10 species pass to the Lower Tremadoc. 

Lower Tremapoc.—Table V. The Lower Tremadoc fauna nume- 
rically consists of 28 genera and 58 species; and the fauna is 
compact and characteristic. Only 8 genera and 9 species appear 
from the Upper Lingula-flags below: 4 of them are Crustacea; and 
5 are Brachiopoda, and, with Dictyonema, complete the incoming 
species. The one remarkable feature in the group is the presence 
and first appearance of 5 genera and 12 species of Lamellibranchiata. 
Dr. Hicks discovered them in the Upper Tremadoe beds on Ramsey 
Island. They mark an epoch in the history of the class. Davidia 
and Glyptarca are new genera, Modiolopsis, Palearca, and Cteno- 
donta receive Mr. Hicks’s 8 other species. Until this class is 
found in lower beds, these 12 species identify the Lower Tre- 
madoc of Ramsey Island as an important horizon in time and space. 
Only 4 of the 13 genera and 4 of the 24 species of Crustacea pass 
to the Upper Tremadoc; they are Agnostus princeps, Ampyaw pre- 
nuntius, Olenus alatus, and Dikelocephalus furea ; and 4 genera and 
6 of the 12 species of Brachiopoda also connect the Upper with 
the Lower Tremadoc; they comprise the following—Lingula pe- 
talon, Lingulella lepis, Obolella Belti, O. plicata, Orthis Carausii, 
and O. lenticularis. These and Theca ovata, T’. bejugosa, and T. cus- 
pidata, in all 9 genera and 13 species, constitute the transgressing 
fauna. 

Urrrr Tremapoc.—Table VI. Nine whole classes are wanting 
in this uppermost division of the Lower Cambrian rocks. Only 20 
genera and 33 species compose the fauna of the Upper Tremadoe. 
We have seen that 9 genera and 13 species pass from the Lower Tre- 
madoe, thus leaving only 11 genera and 20 species as truly belong- 
ing to the Upper Tremadoc. More than 50 per cent. of the species 
pass to the Arenig, or 11 genera and 16 species. These few species 
little affect the question of the paleontological break that takes 
place here ; for 42 new genera and 133 new species make their ap- 
pearance in the succeeding Arenig, through some physical changes 
accompanying the zoological, which we have not yet been able to 
satisfactorily discover. Unconformity between this group and the 
Arenig is not known in the typical areas of South and North Wales; 
but at no time in the history of the lowest Paleozoic rocks has appa- 
rent extinction on the one hand, and migration from some unknown 
area in the other, taken place so markedly. We have yet to learn or 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 99 


trace whence came for the first time the 17 genera of Hydrozoa, the 
17 genera of Crustacea, 5 Annelida, 3 genera of Brachiopoda, and 5 
of other Mollusca, &c., all presenting a different facies, or an aspect 
having little affinity with the previously existing fauna. The advent of 
the 17 genera and 42 species of Hydrozoa and the 35 species of Crus- 
tacea, all new forms and widely spread, has still to be explained. 
The 13 species connecting the Upper Tremadoc with the Arenig 
belong chiefly to the Crustacea and Brachiopoda, these two classes 
yielding 13 out of the 16 species. The 6 Crustacea are Asaphus 
affimis, M‘Coy, A. Homfray2, Salt., Checrurus Frederici, Salt., Dionide 
atra, Salt., Ogygua, scutatria, Salt., and O. Selwyni, Salt.; and the 
Brachiopoda, Lingula petalon, Hicks, Lingulella Davisti, M‘Coy, L. 
lepis, Salt., Obolella plicata, Hicks, Orthis Carausit (Hicks), and 
O. lenticularis. With these are Theca simplex, Salt., Conularia 
Homfrayt, Salt., and Bellerophon multistriatus, Salt. 

The Table numbered VII. embraces or shows the numerical value 
and stratigraphical distribution of the species through all the Lower 
Cambrian divisions, or from the Longmynd and Harlech group to 
the close of the Tremadoe, and also shows in the last column, headed 
“‘ Pass to Arenig,” the number of species that pass to that forma- 
tion, or the base of the Silurian as now recognized by many syste- 
matists. It will be seen that 11 genera and 16 species (14) pass 
to the Arenig ; and this mode of expressing the connexion between 
the lower and succeeding formations is carried through all the 
Tables. Thus the right-hand column shows the number of species 
passing up to the succeeding formation, and the left-hand column 
(in all the Tables but this) those that came from an older or lower 
series. As this special Table, and also Table No. I., shows the 
commencement of life in the lowest rocks of the British Islands, 
there are no forms older than those expressed by the number 61 genera 
and 182 species. Their ancestors we know not; neither do we 
know whence was derived the fauna given under the 11 classes. The 
6 smaller Tables distinctly show the more immediate relation of the 
several divisions of the Lower Cambrian rocks: nevertheless this 
completer Table is of value, showing, as it does, the whole range 
from the Longmynd rocks to the top of the Tremadoe. 

I have endeavoured in my analysis of this Table, as in all the 
subsequent ones, to express in words the result of the figures, 
without which the Tables would merely enumerate the facts of 
occurrences of no value to the student of geology and paleontology ; 
and although of necessity tautology must be frequent, yet the 
explanation, it is believed, will be clear. Throughout the whole 
of the Tables (and there is one for each epoch) the genera 
and species are given in the simplest form—the upper ficure 
in each square (and under each formation) enumerating the num- 
ber of genera in the respective horizons, and the lower figure 
4 genera 
20 species * 
gives the census both zoologically and stratigraphically ; z.¢. there are 
4 genera and 20 species in the given horizon or formation. In this 


demonstrating the number of species, thus : This at once 


I0o PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Table or record of the earliest rocks known, the numerous blanks 
show that the classes were not represented or had not appeared 
in the British Isles; but as we proceed through the formations we 
find that in time, or as life progresses, all the classes become repre- 
sented. The study of these Tables with the description may be 
useful ; and it is hoped they will be found not far from the truth, 
and exhibit the present state of our knowledge of the British species 
and their distribution in time. 


Taste VII. 
| , 1 
5 eS 
B Sas 
Shee | iso |a 
= g cI or al 3 ws 
Classes. é -| BR] 8 AA Sasi. & 
: 8 ES la 2la Blade, 25) 5 
5 | +3 5 6 la Sz Giesjasle3 
2) 8/8) 8 (6h em|6 A Bale & 
SGlalH|/a HH P HH Pp [AS 
ee SSS ania maa > 
Plambee ese. ee ceceivwcsee 2d 
Protozoal .m.h-secmer.<oek bod g.tde tie | z 
Actinozoa. 
Echinodermata......... 33s hace Jel oodadl eet 4 odode | eee = 
{ 
AMMEN Ga, 2 eeancs edoosaces STP Oe gS oll Rani s 
| 
Crustacea, .....2.-..-<.28| 20 (03 47, || 2 al eae alist eben teen ie 
BLyOZOARe eee ee sees Dy reel erases emcee leeeoee} 2 [7 
Brachiopoda ............ 54 20 bs Soh See eC 
Lamellibranchiata ...| 5 | 12 }...... sere grease lasode ae 
Gasteropoda. 
! | 
Pheropodai 2.5. wacmeems- lds || gsc toa Be leer 2 : : 
Heteropoda lesdce.cee Na aa ol ers | “aliod| a deeaal eed 2h Leweilegaa 
Cephalopoda ............ FV lees liceeosd|2s.i00¢ [ctw tone 2 ee 
ee ere es eoumee Peer Ee 
2 8S | 24 17 16 28 20 14 
61 182 3 besa 36 Al 58 33 16 | 


Arentce Rocks. 


The Arenig or Skiddaw group of Sedgwick, immediately under- 
lying the Llandeilo flags, forms the base of the true Silurian rocks. 
From Arenig-fawr Prof. Sedgwick and Mr. Salter obtained two of 
the characteristic fossils, Calymene parvifrons and Ogygia Selwyni. 
Sedgewick recognized these beds as being different from the Llandeilo 
above, and called them Arenig slates, believing them at the time to 
be the top of his Ffestiniog group. The group is distinctly recog- 
nized in the lead-mining district of the Stiperstones area, east of 
the Longmynd, underlying the true Llandeilo flags of Shelve and 
Corndon Hill, the Lower Llandeilo of Murchison. Salter first recog- 
nized the Arenig group near St. David’s, passing upwards into the 
Llandeilo flags, which are so finely shown at Abereiddy Bay. The 


ANNIVERSARY ADDRESS OF THE PRESIDENT. IOI 


distinctness of the Arenig from the succeeding Llandeilo is clear, 
both physically and palzontologically. The following table exhibits 
most of the type forms occurring in the Arenig and the Llandeilo :— 


Lower 
Llandeilo. 
Upper 

landeilo. 


Ir 


| 
| 


BPI CMAGAL 6.4 2c .eecescianesie ads x |... | ... | St. David’s, Whitesand Bay. 

SELLA 1 ee x |... | ... | Stiperstones and N. Wales. 

Tai-hirion,near Arenig-fawr, 
Shropshire. 


Calymene parvifrons .........46. Mull ossejareo { 
Afgiina binodosa ..............006- * 


——— PTANIS ...........2ceceeeeeeees % 

—— caliginosa.................000 * 

Trinucleus Murchisoni ......... Koh assed eee { Pee Ae teams (1S 
stones). 

(EULTLDSTT. | Seo ets eee x |... | ... | .Whitesand Bay. 

Orthoceras Avelinii ............... * |... |... | Stiperstones. 

Obolella plumbea .........s0...-... * | x2 | x 5 

Cucullella anglica ..........0.... oe ha | eee 5 

IBICRIMIGCS CSD: 5. .s-s-0-2ceceeees * 


Asaphus tyrannus...............06. 
POCA SKCS Ps caiiowiones'sincsonee - 
Ogygia Buchii, var. convexa ... 
Calymene cambrensis ............ 
SRPIMWELCHS LAVUS® ..........0...00 06 
Lichas patriarchus ............... 
Bellerophon bilobatus ............ 
EUCHECO FOUR was i... van caessneeen se 
iimeulaeranulata ..............- 
AELCWMALA wos... sescccececcceee 
Orthisctriatula.... 0s. 6. 
GnllioramMa  .2.%-.4.0s.00: 
Crenadonta,.? Sp. ...05.0...00-s00 abe 


* 


x 


| Rrenyn in L. Llandeilo. 


*K OK 


Fairfach, near Llandeilo. 


* OK 2K OK OK 


KOK OK OK 


Gilwern, near Builth. 


Ogygia (Asaphus) corndensis .. 
il Abereiddy Bay. 


15-716 ST Re cs ae 
Barrandia (Ogygia) radians ... 
(COR LPI Cae ate eee eee 
Calymene duplicata ............... 
Trinucleus fimbriatus ............ 
Cheirurus Sedgwickii ............ 
PANE MUMANIS © oo... 60.2... .ceeeseees 
Aenostus Maccoyi °...........:... 
Lingula Ramsayi .................. 
Bellerophon perturbatus ......... 
Murchisonia simplex ............ 
Modiolopsis inflata ............... 
Didymograptus Murchisoni ... 
@rthis calligramma ............... 


*« 


1k 


Builth and Abereiddy Bay. 


*K kK OOK xk OK 


—-——4- 


Abereiddy. 


KK Kok 


Builth. 


* kK OK 


12 | 14?) 16 


«The great break in organic life between the Tremadoc slates 
and the Arenig or Skiddaw group determined Salter, after working 
VoL. XXXVII. 4 


I02 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


out the two faune in the Tremadoc area in 1853, to regard the 
Arenig as the base of the Upper Cambrian of Sedgwick” *. Phillips 
regarded the Arenig as the upper or terminal member of the Middle 
Cambrian. Upon fossil evidence we may commence the Upper 
Cambrian of Sedgwick with the Arenig series, which is represented 
in the ‘“Stiperstones” district. It was not until 1859 that Mur- 
chison and Salter described the fossils of that area, although Sede- 
wick in 1843 established the Arenig group, and subsequently 
obtained the same fossils from the Skiddaw Slates, which were 
described by M‘Coy previous to 1851. Salter, from the majority of 
the fossils in the T'remadoc and Arenig groups, considered (and, I 
believe, rightly) that the Tremadoe is the ‘natural termination ” 
of the Ffestiniog or Middle Cambrian series, and the Arenig the 
base of Sedgwick’s Upper Cambrian. Lyell adopted this view in his 
manual; and Hicks and others have followed Salter in thus placing the 
two groups. In the Cambridge Catalogue the Arenig is treated partly 
as an intermediate group; and although it is 4000 or 5000 feet in 
thickness, yet the number of known species is few—about 60, all 
named, in the Woodwardian Catalogue. | 

That there is a perfect, continuous, and conformable succession 
from the Tremadoc into the Arenig rocks, admits of no doubt. It 
is true that this is seldom seen, even in the areas where they are 
typically developed. In the Shelve and Stiperstones districts, where 
36 species are known, and in the Skiddaw area, yielding 76, no 
downward stratigraphical succession has been truly or definitely de- 
termined. The Arenigs seem to stand alone, especially in Westmore- 
land, where no older stratified rocks occur with which to compare 
them. In South Wales, however, at St. David’s, their base rests upon 
the Tremadoc, and their summit is overlain by the Llandeilo beds of 
Abereiddy Bay, both clearly defined. The three divisions here have 
yielded no less than 96 species, 40 of which are Hydrozoa, and 31 
Crustacea. Every Graptolite here makes its first appearance in 
time ; none are known below the Arenig rocks. Six Crustacea out 
of the 31 are derived from the Tremadoc, and only 3 pass to the 
Llandeilo ; therefore 22 species are peculiar to the Arenig. Indeed, 
out of the whole Areriig fauna, comprising 149 species, all but 38 
are restricted to it. No more distinct group occurs in the British 
Islands. 


ArEnIG Rocks oF Norta WALES. 


Professor Sedgwick, in 1852, described the Arenig slates and 
porphyries of North Wales as forming a distinct and well-marked 
subgroup in his previously named “ Ffestiniog group,” and as rest- 
ing upon the underlying Tremadoc slate. It is also the “ great 
eroup of roofing-slate and contemporaneous porphyry ” described ds 
occurring in the chains of Arenig, Arran Mowddwy, and Cader Idris, 
and in the Ffestiniog mountains, or the western base of the Arenigs 
and Arran, and probably the Stiperstone rocks. For many years 


* Catalogue of the Collection of Cambrian and Silurian Fossils in the Geolo- 
gical Museum of the University of Cambridge, p. 18. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 103 


the elucidation of the true position of the Arenig group in North 
Wales remained a doubtful matter; probably the slates of Angers 
are of the same age. Mr. Salter did much in 1853 to unravel the 
structure of the Portmadoc area, and was enabled to correct the 
nomenclature and distribution of the so-called Lower Llandeilo 
rocks of the Penrhyn promontory, between Traeth-bach and Traeth- 
mawr, and of the country to the east, towards the Arrans. The 
succession across the Penrhyn promontory, from the Lingula-flags to 
the Arenig beds, is complete ; and itis here that the Tremadoc slates 
and Arenig beds are typically developed in North Wales, the close 
of the Tremadoe and base of the Arenig being everywhere marked 
by the presence of a well-defined band of “ grit,” varying in thick- 
ness. ‘The position of this grit had long been fixed by the late Mr. 
Salter in his Penrhyn section ; but it was left for Professor Ramsay, 
in 1874, with Professor Hughes, Mr. Homfray, and myself, to trace 
this zone from Ogof-dua, near Criccieth, to Drws-dwgoed, under 
the Dolbenmain alluvium, and on to T'ai-hirion ; and in 1875 the late 
Mr. Ward and Mr. Herbert continued and completed the survey of 
it west of the Arenig and the Arrans on to Cader Idris, until last 
seen passing under the sea at Towyn. This line is the base of 
Sedgwick’s Arenig slates and porphyries, and of the great roofing- 
slate series of the Ffestiniog area. Although occasionally obscured 
by faults, it is never lost, and must be traced by all who would 
understand the succession in North Wales between the Tremadoc and 
Arenig series. The fossils, when seen, at once determine the posi- 
tion—Psilocephalus innotatus, Niobe Homfrayi, Asaphus Homfrayt, 
Lingulella Davisir, and Dictyonema sociale illustrating the lower part 
of the Tremadoce, and Angelina Sedqwicku, Asaphus Homfrayi, and 
Ogygia scutairia at Garth and Penclogwyn the upper ; whereas, suc- 
ceeding the grit-band above named, at Ty-obry, Tai-hirion, &c., Caly- 
mene Murchisom, Asaphus affinis, Dionde atra, and Aiglina caliginosa 
are the characteristic forms, being Arenig fossils. Only about 16 
species connect the two groups. This physical line, when once deter- 
mined, is a marked feature at the base of the Arenigs, dividing the 
two formations. Beneath the grit at Garth Hill and Deudreath oceur 
the upper beds of the Upper Tremadoe slates (hard grey flags), rich 
in fossils, including Asaphus Homfrayi, Angelina Sedqwicku, Ogygia 
scutatriv, Lingulocaris lingulecomes, Cheirurus Frederici, and Co- 
nularia corvum. These hard grey beds at the top, some would 
separate from the Tremadoc group, purely upon paleontological 
grounds, or because certain species above named are common to the 
two horizons. Some regard must, however, be paid to stratigra- 
phical evidence. 

There is no distinct passage-bed containing fossils between the 
Upper Tremadoc and the Arenig. Through the Portmadoe district 
the two groups are separated by a peculiar band of felspathic grit, 
which intervenes to cut off the upper (volcanic) series from the true 
Tremadoc slates below. ‘This felspathic grit, variously conditioned, 
may be traced along Yr-allt-wen, above Tremadoc, and across the 
estuary of Treath Maw; it constitutes the brow of the Garth, and 

22 


I04 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


ranges under the foot of Moelwyn to the Manod mountains on the 
east. This zone, all along its course, indicates and ushers in a new 
set of conditions, physical and paleontological. No Graptolites 
occur below this line anywhere; but in the black slates immediately 
overlying it at Garth &c. they occur in plenty, associated with cer- 
tain species of Homalonotus, Asaphus, Calymene, Aiglina, and Dio- 
nide, accompanied by Conularia and Lamellibranchs; and among the 
lower beds of the volcanic rocks of the Manods Calymene parvifrons 
and Trimucleus Murchisons occur, accompanied still further eastward 
by Ogygia Selwynu, the species, of all others, which characterizes the 
Arenig rocks of Shelve and Skiddaw, and occurs at Llanfaelrhys, in 
South Caernarvonshire, associated with Lingula attenuata and Grap- 
tolhithus Murchison. ‘This classification is borne out by the persis- 
tent position of the grit which everywhere separates the two hori- 
zons palezeontologically as well as stratigraphically, the mass of the 
Arenig with its peculiar fauna not occurring until the grit is 
passed. No species of Rhabdophora is known below this grit (their 
first appearance being in the Arenig rocks); and although the 
Arenig Graptolithide are rare in North Wales (4 species), as com- 
pared with Skiddaw (27 species), or with South Wales, St. David’s. 
(18 species), still the distinct Crustacea must be held as significant. 
Again, comparing the value of the same formation in North and 
South Wales, it is important to remember that, of the whole Arenig 
fauna in South Wales (97 species), only 5 north-west forms are 
common to the two areas: these are Calymene parvifrons, Ogygua 
Selwynu, Aighna caliginosa, Lingulella Davis, and Orthis lenticu- 
laris. 26 Trilobites, 8 Brachiopoda, 38 Rhabdophora, and 3 Ptero- 
poda are peculiar and confined to South Wales. Thus amongst the 
same group of rocks the correlation of species is a difficult question, 
even over so small a geographical area as Wales. — 

The dark slates at Ty-obry, which immediately overlie the grit, 
contain Calymene parvifrons, diglhina caliginosa, and Diomde atra, 
associated with Diplograptus mucronatus, Climacograptus confertus, — 
and Gilossograptus ciliatus, all Lower-Arenig species. Palewarca 
socialis and a species of Ctenodonta are the only North-Welsh 
bivalves, and Conularia cortwm and C. margaritifera the only two 
Arenig Pteropods known in North Wales. None of these occur in 
the Arenig series of St. David’s, and none pass to the Llandeilo 
group im any area. 

The researches of Dr. Hicks at St. David’s have resulted in greatly 
advancing our knowledge of Sedgwick’s group in that area*. In 
his paper upon “the Succession of the Ancient Rocks of St. David’s,” 
Dr. Hicks draws attention to the history of the names Arenig and 
Llandeilo groups, and explains the manner in which the two series 
were confounded by one of the authors who have written upon them, 
and how the slates above and below the Arrans and Arenigs were 
by him and the Geological Survey classed as equivalent to Llandeilo 
flags of Builth &e. 

The St.-David’s Arenigs consist mostly of black slates, resting 

* Quart. Journ. Geol. Soc. vol. xxxi. pp. 167-194. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 105 


conformably upon the Tremadoe group on the N. and N.E. side of 
the anticlinal, and they may be continuously traced for many miles. 
They occur also at Ramsey Island, and are there 4000 feet thick, 
chiefly deep-sea deposits, and must have been of great duration, as 
proved by their varied and distinct faunas. 

Dr. Hicks divides the Arenig group, through its zonal contents, 
into three subgroups, the Lower, Middle, and Upper Arenig. The 
lower, or black slaty Arenig, strikes E.N.E. and W.S.W., and has 
a vertical dip. The cliffs at Whitesand Bay and Road-uchaf, in 
Ramsey Island, yield the chief fossils. Numerous species of Grap- 
tolites have been obtained from this lower subgroup ; the Trilobites 
Asaphus Homfrayi, and Ogygia scutatrix, with Conularia Homfrayt 
and Lingulella Davisi, especially characterize this lower division, and 
are associated with Phyllograptus stella, Trigonograptus ensiformis, 
Callograptus radiatus, and Piilograptus Hicks. Dr. Hicks enume- 
rates 25 species from this division, 28 from the middle, and 33 from 
the upper*. Asaphus Homfrayi, Ogygia scutatrivc, and Conularia 
Homfrayi are Upper-Tremadoc as well as Lower-Arenig forms, and 
connect the two formations. I am not aware that Lingulelia Davisit 
ever again appears above this horizon in the Cambro-Silurian rocks. 
The middle slaty and flaggy groups are also best seen at Whitesand 
Bay. In 1860 Mr. Gibbs, the late fossil-collector for the Survey, 
obtained a few fossils here. The characteristic Trilobite Trinucleus 
Gibbs was first obtained by him at Whitesand Bay, and subsequently 
in the Skiddaw beds. This locality has furnished Ogygia bullina, O. 
peliata, Aiglina grandis, Trinucleus Gibbsu, T. Sedqwicku, Ampyx 
Saltert, Lingula petalon, and Orthoceras sericewm, with 6 genera and 
11 species of Rhabdophora; all the above are essentially character- 
istic of, and confined to the middle subdivision. Of the Graptolites 
the chief are Tetragraptus crucifer, T. serra, T. Hicksu, T. Halli, 
Clematograptus vinplicatus, Callograptus elegans, and C. Salterc. 

Lithologically the Upper Arenigs resemble the Lower, being fine- 
grained dark shales, 1500 feet thick ; they underlie the true Lower 
Llandeilo near Abereiddy Bay, and are conspicuous for the new and 
distinctive fauna and the first appearance of many genera of Trilo- 
bites, distinctly marking a complete and progressive change from the 
fauna of the Tremadoc below towards the introduction of succeeding 
Llandeilo forms. Llanvirn quarry has yielded to Dr. Hicks a rich 
group of species ‘distinct from any previously discovered in any 
part of the Arenig series at St. David’s”?. Most of the genera are 
new ; and several genera which appear for the first time, culminate 
in the Llandeilo and Caradoc rocks; these are J//enus, Illenopsis, 
Barrandia, and Phacops, each having a representative species, and 
Placoparia, this last a genus new to Britain, hitherto only known in 
France, Bohemia, and Spain. The occurrence here of this genus is 


* These numbers are doubtless in excess;"many forms must give way on 
eritical examination, numerous species being made upon fragments only. 
Mr. Lapworth has already reduced them considerably. The Arenig of St. 
David’s may yield 25 species; the whole Arenig probably about 50 species. 

t Dr. Hicks, Quart. Journ. Geol. Soc. vol. xxxi. p. 174. 


106 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


important as bearing both upon distribution and correlation. Again, 
the Gasteropoda first appear in Britain in the Upper Arenigs of St. 
Dayid’s (Pleurotomaria llanvirnensis, and Ophileta or Raphistoma). 
Yet, strangely, we have no record of any Lamellibranchs through 
the whole of the Arenig group in South Wales; 4 species are known 
from the Stiperstones area, and 2 from Ty-obry, in North Wales. It 
will be remembered that no less than 12 species occur below in the 
Lower Tremadoc rocks of the same area. The value of this upper 
division of the Arenig group in South Wales is enhanced by its re- 
semblance to Barrande’s Etage D1 (Bohemia), and its close relation 
to the fauna found in the Arenig slates in North-west France. The 
recognized presence of this stage in Britain enables us to correlate 
our beds with those on the continent, and thus establish a general 
succession for the Western-European and Western-British rocks of 
this age. 

This result is due to the researches of Dr. Hicks, as detailed in 
his valuable papers upon the history and succession of the ancient 
rocks of St. David’s. Carefully constructed tables, prepared for the 
purpose of testing the value of the Arenig group and its relation to 
the Tremadoc below and Llandeilo above, clearly show that it stands 
almost alone ; for of the 55 known species of Hydrozoa in the Arenig, 
only 2 species pass to the Llandeilo (Didymograptus Murchisont and 
Climacograptus confertus). The first appearance and the abundance 
of the Rhabdophora in the Arenig rocks would alone justify the posi- 
tion now assigned to it as the base of the Lower Silurian rocks; only 
1 species is known to occur in the Tremadoc group (Dendrograptus, 
from the Shineton Shales of Shropshire). The crustacean fauna is 
equally conclusive ; out of 50 Arenig forms known, only 2 (7rinu- 
cleus Ramsayi, Hicks, and Homalonotus bisulcatus, Salt.) unite the 
Arenig and Llandeilo through the Trilobita. Out of the 18 species 
of Brachiopoda, only 3 pass to the Llandeilo, viz. Lingula brevis, 
L. attenuata, and Orthis calligramma. No Lamellibranch, Pteropod, 
Heteropod, Gasteropod. or Cephaloped (of which united, there are 
30 species) passes to the Llandeilo. ‘Thus, out of the known fauna 
of 149 species, only 16 pass upwards into or are common to the 
Llandeilo beds. 150 species make up the entire known fauna of 
the Arenigs; their distribution is as follows :—125 species occur 
in the North and South Wales Arenigs, 34 from various localities 
in North Wales (Tremadoc, Ty-obry, and Tai-hirion), and 94 in 
South Wales; the Skiddaw species number 50 and the Stiper- 
stones 36. ‘These two latter localities are important ; they are widely 
separated, and very few species occur in common. Of the 50 Skid- 
daw species of all classes, only 9 occur in the area west of the 
Stiperstones in Shropshire ; this is due chiefly to the rich Grap- 
tolite fauna, which group is largely represented, no less than 52 
species occurring in the Skiddaw beds alone. West of the Stiper- 
stones only 6 species are known; and they have an important bear- 
ing upon distribution: they are Didymograptus patulus, Hall, D. 
geminus, D, hirundo, D. constrictus, D. Murchisoni, and Clematograp- 
tus implicatus. Diplograptus dentatus also occurs at Tremadoc and 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 107 


Tai-hirion in North Wales, and ranges to both Llandeilo and Caradoc; 
Didymograptus patulus, D. hirundo, and Clematograptus vmplicatus 
are also present at St.-David’s*. Few Hydrozoa occur in the North- 
Wales Arenigs; Ty-obry, Tremadoc, and Tai-hirion yield the few 
that have been met with. 23 species range through the St.-Dayid’s 
series, 2 of which only are common to the Stiperstones area, viz. 
Didymograptus patulus and D. hirundo; and 5 or 6 ally them to 
Skiddaw. There is thus little in common between the Arenigs of 
Skiddaw and North Wales, few species connecting them. The rela- 
tion, however, between the Skiddaw species and those of St. David’s 
is closer, 5 or 6 being Graptolites, and 2 Trilobites (Ogygia Selwyni 
and Trimucleus Gibbsiz). These, we must remember, are the two 
largest groups, and have been most searched for, at St. David's 
especially, by Dr. Hicks, and in the Skiddaw beds by Dr. A. Nichol- 
son, Mr. Dover, and the Survey (through the late Mr. Ward). 

The value of the Arenig group is still more apparent when we 
consider the development of life that accompanied those physical 
conditions and changes which took place at the close of the Lingula- 
flag and Tremadoc epochs. We know that no less than 40 genera 
make their first appearance in the Arenig rocks in the British Islands. 
These 40 genera belong to the following 6 classes :-— 

Hyprozoa 16: Callograptus, Dendrograptus, Phyllograptus, Di- 
dymograptus, Azygoyraptus, Ptilograptus, Trigonograptus, Tetra- 
graptus, Climacograptus, Nemagraptus, Dicellograptus, Clemato- 
graptus, Diplograptus, Glossograptus, Dichograptus, and Logano- 
graptus. 

AnneLipa 4: Helmintholithes, Stellascolites, Nereites, and Palco- 
chorda. 

Crustacea 11: “glina, Trinucleus, Barrandia, Calymene, Pha- 
cops, Placoparia, Illenus, Illenopsis, Homalonotus, Beyrichia, and 
Caryocaris. 

Bracuiopopa 2: Dinobolus and Siphonotreta. 

LAMELLIBRANCHIATA 2: Ribeiria and Redonia. 

GastERopops 3: Ophileta, Pleurotomaria, and Rhaphistoma. 

The succeeding Llandeilo, as we shall see, was equally prolific, no 
fewer than 48 new genera accompanying the complete change of 
conditions at the close of the Arenigs and commencement of the 
Llandeilo. This change was bridged over by orly 8 genera and 9 
species passing to the Llandeilo in any area; 4 are Hydrozoa, 2 
Crustacea (Zrinucleus Ramsay: and Homalonotus bisulcatus), 3 Bra- 
chiopoda (Lingula brevis, L. petalon, and Orthis calligramma) ; and 
only 3 range into the Caradoc. As before stated, the relation of 
the Arenig to the Tremadoc is through 16 species, 6 of which are 
Brachiopoda, 6 Crustacea, and 4 bivalve Mollusca. Thus 122 
species out of 150 are strictly Arenig forms or confined to that 
horizon. 


* Five other Skiddaw species occur west of the Stiperstones, and, with the 
6 Graptolites, tend to connect the Arenigs of Shropshire with those of West- 
moreland. They are the Annelida Scolithus linearis, Salt., and Helmintholithus, 
with Agnostus Moret, Salt., Ogygia Selwynit, Salt., and Zglina binodosa. 


108 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Nothing can be more conclusive as to the value of the Arenig 
group than the distinctive part played by its characteristic fauna 
and its distinct stratigraphical position. Since the determina- 
tion by Hicks of the St.-David’s fauna, Mr. Marr of Cambridge 
recognized some fossiliferous shales south-east of Caernarvon, in | 
three localities on the banks of the Sciont; these shales contain 
Arenig species. The Trilobita and Brachiopoda indicate Arenig 
affinities; and the Orthoceras caercesiense occurs only in the Arenigs 
of Llanvirn, near St. David’s. The genus Caryocaris (a new spe- 
cies of which has occurred here, C. Marri) hitherto has not occurred 
out of the Skiddaw Slates of Cumberland (C. Wrightw being the 
type). The associated Graptolites are those of the Skiddaw rocks 
also, viz. Didymograptus bifidus, D. indentus, and the Llandeilo form 
D. Murchison. The genus Barrandia would help to place these 
beds either in the Arenig or the lower part of the Llandeilo group. 
This locality is suggestive of the Arenig or Llandeilo beds striking 
from Caernarvon to Bangor and Aber, west of the great fault that 
runs from Dingle on the 8.W. to Aber on the N.E. 


ARENIG oF SoutaH Wategs At St. Davip’s. 


The following table enumerates all the species at present known 
in the Arenig rocks of St. David’s ; it is convenient to subdivide the 
group into 3 series, both on lithological and palecontological grounds. 
It will be seen how distinct the fauna of each division or series ap- 
pears. The total number of species at St. David’s is 70. The lower 
Arenig has yielded 16 species, the middle 24, and the upper 33. 
14 species are peculiar or confined to the lower series, 21 to the 
middle, and 30 to the upper. No species out of the 70 occurs in all 
three divisions. We thus see how distinctive a group of fossils 
each subdivision contains. For the construction of the Arenig Table 
(No. VIIL., p. 115) I have selected 18 localities from which the Arenig 
species have been obtained, and tabulated 8 of the most important. 
The four type areas, where the Arenig rocks are present and well 
developed, are :— 


1. Westmoreland: Skiddaw. 

2. Shropshire: Shelve area. 

3. North Wales: Tremadoe, Tai-hirion, &e. 

4, South Wales: Ramsey Island and St.-David’s area. 


The chief localities for fossils are the following 18, comprised in 
the above 4 areas :— 


1. Westmoreland: Skiddaw, Longside, Keswick, Outerside, Scaafel, White- 
side, and Braithwaite River. 4 

2. Shropshire: Shelve area (west of Stiperstones), Mytton Dingle, Perkins 
Beech. 

3. North Wales: Tremadoc, Garth, Ty-obry, Tai-hirion. é 

4. South Wales: Ramsey Island, Llanvirn, Whitesand Bay, and St. David's. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 


Areng Species, St. David's. 


109 


Species. Lower. | Middle. | Upper. 
RHABDOPHORA. 
Didymograptus extensus, Hall ...............08 % 

LETTE, ING Oe, WO anaes Ole pon eR ea eer au * 

[SECIS 2 (01 Hoes DAM ee eee Re * * 

bien ALOU T ees os title cakk schist oStiele tat mses’ * 

PeMa GUUS PAU ess casaslectoeincie set eeee sess % x 

BMAESUS, eLLODemewa. scene va dcticms cusstice've vide * 

NOW CMEUS AONE ath... oe idvolaiicndasenessccs * 
———INGCMOlsont, Lap .iscsecccesseseceeereeees: Ste * 
Phyllograptus stella, Hall .......00....0ececeeseee * 

PUI SIA Lice aeaica cto notiscleaie sSsiseaiscissebieis d's * 

Trigonograptus truncatus, Lapw. .............4. % 

emsionnis, Halles. .scasnssasesi cs sastae seeks * 

eiostapuus EMCkKsi .2..5....2:20sstssacessea0s 0 * 
Dicellograptus divaricatus, Hall ............... x 
Diplograptus dentatus, Brongn...........0...066- * 
Climacograptus confertus, Lapw. ............... See * 
Clematograptus implicatus, Hopk. ............ * 
Tetragraptus crucialis, Sal¢. ............0.0eee ee * 

EINE O DICK Toes. 8 ote ae dun sin das ce saree cules x 
LIP SIM OPK io loc Us wcnieavenedercscs senses % 

eyouordes, PGI... c.%as.2esebasenses sce os * 

quadribrachiatus, Hall ...........-.c.000--- % 

SMTP OMG a. stun onus cee naduscees escac uses % 
Glossograptus ciliatus, Hii.........cecseeeeeeeee * 

CrusTACEA. 
PSAnMISPELOMUPAYT, SALE. vo. lesnsienosccstt sense * 
RTOS US MIUN GON SCUC.0 i cevsscsceesese-deeeesacc % 
PAM SALCON, LLICKS oo nos cwevnascee MH acscene cast: * 
PBeliMaeeRAMCIS, SAUL. 2... 0... cc ee sceesncsensceess % 

LOT, JE GO AS BAS apee te ye SNC a eae Aer een at i * 

Obtusicaudata, Aicks . 508... eels * 
iBasranGtasclomibirayl, A2cks ...0...5..sene ase: * 
Calymene Hopkinsoni, Hicks................0006+ * 

PML OMS USOUG shee cach Geis ele soja esto ecco % 
— Pyare MuUrchisoni Saleen a 43 % 

», SID: Boot des Sak MANOR ae OSE eae oF nas Ace * 
POEM BUSS i785 Cece Uh eek leldigoSae deta Mela kiele Hae * 
Oevedarsemtatrix, SAC... s6c.cassncecssnescieetees * 

MUG A ISL Ca, fo serivo ag Jgsesittic nnceearis ceeoe * 

"DAW UTa Bis “PSYOMAES Meck Soares tem Soa Re a Rs * 
Mlcemmomtughesti, AUCKS... fo .65.2.e6 le seveks cee: * 
Illenopsis? acuticaudata, Hicks ............... * 
Placoparia cambrensis, Hicks.............0.0+++0 af * 
Phacops Ilanvirnensis, Hicks .........0..0..s0++: sie aN *% 
Mrimteleus GalbDsit, SAC. ccoc.cc. ve sceseerecesneees * 

Dem AWC KI WOR sno esate eaticscasenscacee * 
EMCO COR, SOLE. « saceceues souls doudee soeedeees * 
EVI SLY, MELICHSH. clic ccciseais aes décieleioa Rsv % 

MSE Saisie Saisie oct aaa be cttasndasen aodeyiaes x 
IE YNEIC NVA AB YDeg 7. 5 vsiae,s Seis siogin oataeleintininais som date te oiler * 

Carried forward.....,...... 11 18 22 


IIo PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Aremg Species, St. David’s (continued). 


Species. Lower. | Middle. | Upper. 
Brought forward) 11 18 22 
BRACHIOPODA. 
Lingulella Davisii, M‘Coy ........... Bor Nee erin * 
Wine wlaypetalonne chs ene are ecm eee ere er % * 
ingulayattenuata SOU: yeeeseene serene ecaeece 08 ea % 
Discobolusv tacks eave nee eee eee * 
DIS CIMAS SPE teste ie cia se oe Se an se icine rr wae % 
Oris lenticularist)0l 7a ee ee earn * 
iS Dus accilecceactcaseistrclmasct et cmamreec a tace ence at ve * 
Ai) Ohne ars ie Seba Ue Tiel brane sneer a * 
Obolellapplicata PECs aces eenesee ete eeeeent * 
Siphonotmetayspycscaset ecco eee eeae eereea ter 60 % 
GASTEROPODA. 
Oplulletal spy. togs cs. cttbionecs wae toaenece toate 500 ane * 
Pleurotomaria llanvirnensis, Hicks ...........- % 
PTEROPODA. 
Conularia dlonatraya Salt. scseeeee reset etceneee * 
Manvatnensis" H7CS: ss aaeee ce oe ears * 
Mhreca Cacreesiensisy ea7CHS 0% «cesacaeeee eee Bei bee * 
lar knessil AICHhS cote te ce eh Renee fis % 
HETEROPODA. 
Bellerophon multistriatus, Salt. ............6.. siete * 
=== AMVATTICNSISML CHS” a) Jneeceecoaeceeeence see ae * 
CErPHALOPODA. 
Orthoceras sericeum, Salt. ...::::.0.5..522c8sn.8ee a * 
caereesiense, Wicks 2h. de ia, See ssc Bie % 
ANNELIDA. 
Buthotreplisisp.\.07lntecsesedsscece soccer sea e 400 Be * 
16 24 30 


The intimate stratigraphical conformity at St. David’s between 
the Tremadoc and Arenig rocks, and between the latter and the 
overlying Llandeilo series, is clear and determined, both on the 
mainland and on Ramsey Island, where both the Tremadoc and the 
Arenig occupy a considerable area. Much of the centre of the 
island is composed of the Arenig series, and is prolific in fossils. The 
slates of St. David’s are black and probably of deep-sea origin. The 
subdivision here by Dr. Hicks into Lower, Middle, and Upper is 
based upon the distribution of the organic remains and the relation 
to the Llandeilo above. This comparison is important in its bear- 
ing upon the value and meaning of the terms Arenig and Lower 
Llandeilo as applied to at least four areas and groups of rocks in 
England and Wales, viz.:—Westmoreland, or the Skiddaw group ; 
Shropshire, or the Stiperstones series; North Wales, or the Tremadoce 
and Ffestiniog areas; and South Wales, or the St.-David’s beds. 
Formerly the Geological Survey regarded three of the groups in the 
areas above mentioned as belonging to rocks termed Lower Llan- 
deilo by Sir R. Murchison; they were mapped as Llandeilo simply, 


ANNIVERSARY ADDRESS OF THE PRESIDENT. Ii! 


the Survey not recognizing the two divisions termed Lower and Upper 
on the maps, as they are not separable or distinguishable in the field 
over large areas. The exact position of the Arenig group was, and 
has for many years been, a doubtful question ; and it is due to the 
continued researches of Messrs. Salter and Homfray in North Wales, 
and of Dr. Hicks in South Wales, that the name and position assigned 
to these beds by Prof. Sedgwick in 1843 is now revived and defi- 
nitely established, and that the Arenig group of rocks and fossils 
has now had assigned to it its true stratigraphical place above the 
Tremadoe group and below the Llandeilo proper, and with a fauna 
recognized as peculiarly its own. The Llandeilo of Murchison and 
the Survey (first named in 1843) did not really include any of the 
Arenig proper as determined in North Wales in 18438, 1846, and 
1852, or as occurring in the chain of the Arrans, Arenig, Cader Idris, 
and in the Ffestiniog region (typical districts). The term ‘‘ Lower 
Llandeilo” of Murchison included Prof. Sedgwick’s “ Arenig,” not 
as first intended ; and Prof. Ramsay, in the first edition of his ‘ Geo- 
logy of North Wales’*, states that “since 1848 the Survey con- 
sidered the slates close below and above the Arans and Arenigs as 
equivalent to the Llandeilo fiags of Builth and Shelve” 7. 

Mr. Salter, however, in the appendix to the same work in 1866, 
pp. 293-257, under the section “‘ Lower Llandeilo” and plates 8-12, 
clearly showed the importance of truly correlating the Arenig group, 
eliminating it partly from the Upper Tremadoc below and from the 
so-called Llandeilo above. 

There is now no doubt whatever about the horizon or position 
of the Arenig rocks. They are entirely distinct from the Tremadoc 
eroup of Sedgwick, and were claimed as Lower Llandeilo by 
Murchison. They are the “ Arenig and Skiddaw” group of Sedg- 
wick, established by him in 1834, and confirmed through subse- 
quent research in the Skiddaw area, and of which the fossils were 
described by M‘Coy before 1851. Priority therefore under all heads 
is due to the researches of Prof. Sedgwick. Finally Salter described 
the majority of the fossils of both the Arenig and Tremadoc groups, 
showing in the clearest manner that the Tremadoc rocks were the 
natural termination of the Ffestiniog or ‘“‘ Middle Cambrian ” series, 
and the Arenig group the base of the “Upper Cambrian” or 
‘¢ Lower Silurian ” of Murchison ; these rocks through their fossils, 
have their equivalents, as shown by Salter, in the Quebec group of 
Canada. 

Prantra.—None. 

Prorozoa.—None. 

Hyprqz0a.—Subsequent to the change of conditions which ter- 
minated the Tremadoc period, or at the coming-in or commencement 
of the Arenig deposits, no less than 18 genera and 42 species of 

* Mem. Geol. Surv. of Great Brit. vol. ii. p. 6 (ed. 1, 1866). 

t Sir R. Murchison, in 1834, termed the Llandeilo flags ‘“ Trilobite schists ” 
in his description of the Shelve country, the Carneddau (Builth), and the 
neighbourhood of Llandeilo; and under this name were included (near Shelve) 


the strata as low as the base of the Stiperstones, the whole of these rocks being 
older than the Caradoc sandstone. 


I12 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Hydrozoa first appeared in the British Arenig sea, and that with- 
out any known previously-existing Ceelenterate fauna. We have no 
clue to the area or region whence they migrated ; possibly Canada 
may be selected as one, at least, of the probable areas for their 
origin and dispersion. We regard the Quebec group of rocks as the 
_ equivalent in time of our Arenig series; and they contain the same 
species of Graptolites in great profusion. We cannot look to Scan- 
dinavia for such an assemblage; and although the Bohemian Grap- 
tolitic fauna was extensive, still, on the whole, I should regard the 

Canadian, through certain genera and species, as being the source 
of dispersion and migration. There is intimate agreement between 
the faunas of many of the American and our own Paleozoic rocks ; 
and this is notably the case with the Arenig and its Graptolites. 

Thirteen genera and 28 species of Graptolites occur in the 
Skiddaw group, showing that more than two thirds of the known 
genera and species are in the almost barren slates of the Keswick 
area; for, with the exception of 10 species of Annelida (all tracks 
and burrows), 9 species of Crustacea, and 2 Brachiopoda, no 
other forms are known—neither Lamellibranch, Gasteropod, Pte- 
ropod, or Heteropod, so far as we know, ever having appeared 
in the Skiddaw series west of the Stiperstones in the Shelve area*. 
The Arenig rocks are there clearly exhibited, but yield only a 
small series of Hydrozoa. Only 3 genera and 6 species have yet 
been collected ; they are Diplograptus pristis?, Didymograptus pa- 
tulus, D. geminus, D. hirundo?, D. constrictus, and Clematograptus 
amplicatus. The Crustacea eminently characterize the Shelve 
Arenigs; and it is through the 6 genera and 9 species of Trilobita 
and the stratigraphical position that the age of the beds is clearly . 
determined: 3 of the 7 genera that occur here are also in the Skid- 
daw rocks—Agnostus Moret, Ogygia Selwynu, and Aiglina binodosa. 
Next to North Wales the Shropshire Arenigs contain the smallest 
number of species; but they are distinctively characteristic. 

North Wales, from the Tremadoc and Ffestiniog areas at Tre- 
madoe, Ty-obry and Tai-hirion, has yielded only 4 genera and 4 
species. 

: Next to the Skiddaw rocks and the Westmoreland area the chief 
development of the Arenig group occurs in South Wales on the St.- 
David’s promontory, at Llanvirn, Whitesand Bay, Trwyn-hwrd- 
dyn, and Porth Melgan, and the rich locality of Ramsey Island. 
These several localities have yielded, to the researches of Dr. Hicks, 
Mr. Hopkinson, Mr. D. Homfray, and others, about 8 genera and 
18 species of Graptolites. Only 3 species are common to North and 
South Wales, namely Diplograptus dentatus?, Clumacograptus con- 
fertus, and Glossograptus ciliatus ; 6 show the relation of the Skiddaw 
rocks with South Wales, viz. Didymograptus affinis, D. brfidus, D. 
patulus, Tetragraptus quadribrachiatus, Phyllograptus typus, and 
Diplograptus dentatus?; and 7 are common to the Stiperstones area 
and South Wales, viz. Didymograptus patulus, D. affinis, D. hirundo, 
D. bifidus, Diplograptus dentatus, Clematograptus implicatus, and 

* Lords Hill, Bog Mine, Dingle, Ritton Castle, Corndon, &e. . 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 113 


Climacograptus confertus. Out of the entire fauna of the Arenig 
Rhabdophora only 2 or 3 genera and species pass to the Llandeilo 
rocks. ; 

KcutnopErmMsta.—No record whatever of this class in the Arenig 
rocks. 

ANNELIDA.—The whole group, of which 9 genera and 11 species 
are described, possesses no zoological value ; they are simply worm- 
burrows, worm-tracks, or trails of Mollusca. Ten of the 11 so-called 
Annelide species occur in the Skiddaw Slates, the one unrepresented 
form being Buthotrephis from St. David’s. Scolithus linearis and 
Helmintholithes are in the Stiperstone beds also. No species has 
occurred in North Wales; and no species pass up from the Tremadoc 
and Lingula-flags below. 

Crustacea.—The Arenig rocks have unequally distributed through 
them 14 genera of Trilobites and 48 species, Ostracoda 2 (Beyrichia 
and Primitia) and the Phyllopod Caryocaris Wrightt increasing 
the Crustacean fauna to 17 genera and 51 species. The special 
Trilobita, however, number 41 species (6 coming up from the 
Tremadoc and 2 passing to the Llandeilo, namely TZrinucleus 
Ramsayi and Homalonotus bisulcatus). No group in any of the 
British rocks is so specialized as the. Arenig Crustacea. The 
Skiddaw rocks hold 9 species, representing 7 genera *; the Shelve 
Arenigs have 10 species and 7 generat, both areas showing either 
imperfect collecting, or poverty in representation (which is by 
no means likely). The few species (mostly single) known or 
occurring in each genus clearly shows how large a fauna may 
be expected if the more fossiliferous beds could be found in these 
somewhat barren rocks. This applies equally to North Wales, 
where 9 genera are only represented by 11 ‘species. In South 
Wales (St. David’s) the three areas have yielded 15 genera and 

39 species. Circumstances have favoured collecting in this re- 
gion; and the known fauna is increased thereby. The Northern 
or Skiddaw fauna is allied to the Shelve or Shropshire group through 
3 species only (Agnostus More, Ogygia Selwynu, and dAiglina 
benodosa). The special Skiddaw forms are 4 only (Caryocaris 
Wrightu, Phacops Nicholson, Niobe Dovert, and Aglina sp.); but 
9 occur in these beds. The special Shelve species are Trinw- 
cleus Murchison, Illenus perovalhs, I. Thomsont, and Chewrurus 
sp.; but 10 species occur in the Shelve beds. The special North- 
Wales species are also only 2, Asaphus affimms and Caryocaris Marrit. 
The special South-Wales species are Agnostus hirundo, Asaphus 
Homfray2, Ogygra scutatria, O. peltata, O. bullina, Asaphus menapie, 
Trinucleus Sedqwickun, Ampyx Salterr, Aiglina bora, 4. grandis, Ah. 
obtusicaudata, Barrandia Homfrayr, Calymene ultima, and C. vewata ; 
the occurrences in South Wales, however, are 35 species in 15 

* Caryocaris Wrightu, Agnostus Moret, Ogygia Selwynit, Trinucleus Gibbsii, 
Aiglina caliginosa, Salt., A. binodosa, AL. sp., Phacops Nicholsoni, and Niobe 
Doveri. 

t Agnostus Moret, Oyygia Selwynti, Trinucleus Murchison, Ziglina binodosa, 


Calymene parvifrons and var. Murchisont, Illenus perovalis, I. Thomsoni, Chei- 
rurus Frederict, and C. pectinatus. 


II4 PROCEEDINGS OF THE GEOLOGICAL SOCIETY, 


genera. Eleven new genera of Crustacea, 10 of which are Trilobita, 
appeared in the Arenig sea: they are Afglina, Angelina, Trinu- 
cleus, Barrandia, Calymene, Phacops, Placoparia, Illenus, Ille- 
nopsis, Homalonotus, and the Phyllopod Caryocaris, which, with 
Placoparia, never passed to higher rocks. 

Bracuioropa.—7 genera, Lingulella, Lingula, Orthis, Obolella, Dis- 
cina, Stphonotreta, and Dinobolus, represented by only 18 species, 
are all that can be recorded of this class through the Arenig rocks. 
All the genera and 12 of the 18 species occur in the St.-David’s area. 
Only 2 species are known in the Skiddaw rocks, Lingula brevis and 
a Discina; and 5 in the beds west of the Stiperstones ; they are 
LIingula attenuata, Orthis alata, O. calligramma, O. striatula, and 
Obolella plumbea. The 5 North-Wales forms are Lingulellu Davisu, 
L. lepis, Orthis lenticularis, O. caligramma, and Obolella plicata, 
Tai-hirion yielding 4 of the 5. Lengula petalon, L. brevis, and 
Orthis calligramma are all that pass to the Llandeilo rocks; but 9 
come from the Tremadoc ; therefore the special forms are only 6— 
Siphonotreta micula, Dinobolus Hicksu, Dav., Discina sp., Orthis 
striatula, O. remota, and O. alata. The new genera are Dinobolus 
and Siphonotreta. ; 

LAMELLIBRANCHIATA.—Ribeiria complanata, Modiolopsis trapezi- 
formis, Palearca amygdalus, and Redona anglica occur in the 
Shelve Arenigs. Palwarca socialis and a species of Ctenodonta occur 
only at Ty-obry in North Wales. None are known either in the 
Skiddaw rocks or in South Wales, 6 species being the entire bivalve 
fauna. ibeiria and Redonia first appear in the Arenig rocks. 

GastERoPoDA.—Only 4 genera and 4 species are at present known. 
Plewrotomaria Uanvirnensis, Hicks, and Ophileta sp., occur at 
Llanvirn, St. David’s ; Rhaphistoma, Skiddaw only; and the Stiper- 
stone form is Huomphalus corndensis, Sow. No species known in 
North Wales. 

Prrropopa,—4 species of Conularia and 4 of Theca constitute 
the Arenig Pteropod fauna. ‘The Shelve species are U'heca sumplea, 
Salt., and 7. vaginula, Salt. Those occurring in North Wales are 
Theca vaginula, Salt., Conularia corvum, Salt., and C. margarite- 
fera,Salt. Theca Harknessii, Hicks, T. caereesiensis, Hicks, Conularia 
llanvirnensis, Hicks, and C. Homfrayt, Salt., range through the St.- 
David’s Arenigs. 

Hersropopa.—4 species of Bellerophon in the Stiperstone beds 
and 2 in the St.-David’s are all that are known. ‘The Stiperstone 
species are B. hippopus, Salt., B. bilobatus, Sow., B. carimarioules, 
and B. perturbatus, Sow. The South-Wales species are B. multi- 
striatus, Salt., and B. llanvirnensis, Hicks. No species yet known in 
North Wales. 

CrpHALoropaA.—Orthoceras Avelinit and O. encrinale occur in the 
Corndon area (Shropshire).  O. sericewm and caereesiense are only 
known at St. David’s, South Wales; an undeterminable form occurs 
at Ty-obry (Tremadoc, North Wales). None in the Skiddaw beds. 

The distinctive value of the Arenig fauna is shown in the fact 
that only 8 genera and 9 species pass to the Llandeilo; and 4 


ANNIVERSARY ADDRESS OF THE PRESIDENT. II5 


of these species are Hydrozoa, 2 are Crustacea (Trinucleus Ram- 
sayi and Homalonotus bisulcatus), and 3 Brachiopoda (Lingula 
brevis, L. attenuata, and Orthis calligramma). Looking, therefore, 
at the number of genera (63) and species (150) in the whole Arenig, 
the change or extinction at the close of the period has scarcely 
any parallel in the Paleozoic rocks. The accompanying Table 
shows the numerical value of the Arenig species, the 63 genera and 
150 species being distributed through four well-determined areas, 
and showing the fauna to be, in the 


Skiddaw beds ...... 31 genera and 49 species. 
Stiperstones area .. 26 i Sa ae 
North’ Wales’ 202"... 21 A DOr te 
South Wales ...... 3o7 i HON a hes 


Only 8 genera and 9 species transgressed or passed to the suc- 
ceeding formations; and 11 genera and 16 species came in with 
the advent of the group from the Lower Cambrian series. The 
true numerical faunal value of the Arenig group is therefore 51 
genera and 133 species; those forms, or the 9 species, which pass to 
higher formations do not affect the Arenig beyond linking it to the 
next group in succession. 


Taste VIIL—Arenig. 


Geographical 
Distribution. 
: : Pass to 
5 . as aR g n | 
(o) = [<D) 
a5 Ee Sas ie 
ro) i) ea 2) : 
a 3s . = Wea = iS =a S) 
| | eS nD = on (2) oO 
Ss SS | eirsh WR Supers ieee i sin | 
1/5 ® on ‘2 ‘Ss o 5 a oS 
ea Slmia Folalela Oo 
Beam ose eet oh Rene a> teers aah alaral Sele 
PEO OZOR Aad tassadesoaeat onesies 
Ti OZOA Oks v.05. .clotos setae SA Aa sce Mean eae le kee 
PACHIMOZOM SG ssc ose sssscheccess soe 
Hiehimodermata..........secsss- 
9 8 2 
PARIMVE NI G reieciS5stsi sic sv eiselddsenieick CS) TE ee es Z 
PU RUMCEACEA. 9... ccc acseaeasieesleecec LZ) UG) |) ete ea eee I 
BGVOZOA Mec eccctvactnctesddecet. 
3 © ee 2 2 3 7 2 
ale btaehiOpOda.<........7-.-r0+- +s: Ne Wsh |) A : Bollietael le A 
Bamellibranchiata ..........:. A | Or Besee. | lhag 2 
| 1 1 2 
GaAsKerONOCS .. 05 a.icasde es dear. 4; 4] 3 z meh 
2 1 2 2 
Peimeecopodari2.. :sstikeeeii:. DAN fo) Z z e 
1 i 
Te ELC LCLOPOGa4....2:... sesc.08eeees 1| 6 ae ilpcee aleges 
MIC CPUMOPOUA 3.:...0c.-25 +e. 52 Missa line 5) 3 zt A 
real - A 31 26 
Te (PSG 8 ces aed a a G33 ISD) ea ea ee I) a 


116 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


LiANDEILO. 


It may be thought by some superfluous to discuss the history of 
the Llandeilo rocks proper; but it is part of my purpose to do so, 
for two reasons :—first, to show their true relations (affinities and 
differences) to the Arenigs, which they overlie and with which they 
have long been confounded ; and, secondly, to show the geographical 
distribution of .the Llandeilo species and their paleontological rela- 
tions to the Caradoc rocks above and the Arenigs below. The three 
classical localities for the Llandeilo rocks and fossils are Llandeilo, 
Builth, and Abereiddy Bay north of St. David’s. Each of these areas 
possesses a characteristic and definite fauna ; on the whole the species 
in each locality are much the same; still the facies is peculiar in 
each. It is also of much interest to clearly understand the true 
position of the great band of dark roofing-slate of the ‘ Ffestiniog 
quarries” &c. that underlies the Caradoc or Bala series, occupying 
the position of the Llandeilo flags. “It is probable that the lowest 
portion of the Llandeilo group may occur in the Arenig and Ffesti- 
niog mountains, and also in the Arrans ;” but the evidence is slight 
and the materials (in the shape of fossils) are scanty. Salter obtained 
Bellerophon perturbatus in dark slates near Bangor, and also other 
fossils in the slates, identical with those of the Llandeilo rocks, 
‘‘ which overlie the Arenig porphyries.” It is doubtful if the 
black slates of Anglesey are the equivalents of the dark earthy 
slates that range east of the Arenig mountain. The Anglesey slates 
contain Asaphus Powis and Phacops apiculatus, both rare in the 
Llandeilo group; but the Asaphus occurs both in North and South 
Wales, Phacops apiculatus only n Anglesey. The Graptolites, through 
Climacograptus celatus(?) and Didymograptus Murchison, distinctly 
enable us to refer the beds to the Llandeilo group. 

The Llandeilo rocks and their fossils in North Wales are confined to 
few localities, and are but feebly exposed anywhere; and the whole 
fauna of this formation in that area numbers 47 species. Highteen 
of these belong to the prolific class Brachiopoda, 15 to the Crustacea. 
The Llandeilo rocks of South Wales contain 88 species; and about 
40 are common to North and South Wales. Itis important to show 
those species which unite the two areas ; but for my purpose I name 
only the Crustacea (Trilobita), Hydrozoa, and Brachiopoda. Of the 
former class 8 species are common, viz. Asaphus tyrannus, A. 
Powisii, Ogygia Portlocku, Calymene cambrensis, C. Blumenbachii, 
Trinucleus concentricus, T. favus, and the Ostracode Beyrichia 
complicata. | | 

The Hydrozoa are not less important; they are Duplograptus folr- 
aceus, D.dentatus, Didymograptus Murchison, Leptograptus flaccidus, 
Dicranograptus ramosus, D. formosus, D. Nicholsom, Climacograptus 
bicornis, C. celatus, C. confertus, CO. Sharenbergr, and Dicellograptus 
sextans. 

The Brachiopoda number still more, as we should expect from 
their habit and relation to the sea-beds upon which they live. They 
are all common and well-known species; and 8 of the 13 forms 


ANNIVERSARY ADDRESS OF THE PRESIDENT. Le? 


belong to the genus Orthis: they are Crania divaricata, Siphonotreta 
nucula, Orthis vespertilio, O. turgida, O. Actonie, O. calligramma, 
O. striatula, O. elegantula, O. insularis, O. bifoveata, Strophomena 
rhomboidalis, Leptena tenussimistriata, and L. sericea. 

Cardiola interrupta is the only Lamellibranch out of the known 6 
species, and Bellerophon perturbatus and B. bilobatus the only 2 Hete- 
ropoda out of 7, which, with Favosites fibrosus, help to make up the 
41 species connecting the North and South Wales Llandeilos; so 
that, of the 47 North-Wales species, 40 are common to both areas. 
No less than 88 species occur in South Wales. The whole Llan- 
deilo fauna for the British Islands numbers 80 genera and 175 
species ; of these ] may mention that Ireland has yielded 51 species, 
Scotland 66, and Shropshire only 26: these latter are chiefly from 
the Shelve district, and are mostly Crustacea and Brachiopoda. Our 
knowledge of the number of species occurring in Scotland is due to 
the researches of Mr. Lapworth in the Moffat area. 

In the St.-David’s area and throughout Pembrokeshire, Brecon- 
shire, and Caermarthenshire, the typical localities of Llandeilo and 
Builth &c. contain a special fauna. At Abereiddy Bay the black 
slates and argillaceous limestones are interstratified with felspathic 
tuff; they rest upon the Arenig group, having a distinct fauna; a 
thick bed of tuff divides them. The immediate presence and abund- 
ance of Didymograptus Murchisom and Dicellograptus, Cryptograptus, 
&¢e. in the slates on the south side of Abereiddy Bay above the fel- 
spathic tuff, at once clearly marks or determines the base of the 
Llandeilo beds. 

Dr. Hicks divides the true Llandeilo series of St. David’s into 3 
subgroups*, the lowest containing most Rhabdophora or Graptolites, 
associated with Trinucleus Ramsayi, Calymene Murchisone, and 
Theca caereesiensis, which especially characterize this lower division. 
The middle subgroup consists of dark “calcareous shales, with 
compact limestone at the upper part;” Asaphus tyrannus, A. pel- 
tastes, Calymene cambrensis, Ogygqia convexa, and Trinucleus Lloydu 
are the typical Crustacea, with Lingula granulata, Halysites catenu- 
latus, and 6 Graptolites. In addition to the fauna, the calcareous 
nature of the beds lithologically separates this middle subgroup 
from the, over- and underlying series. Everywhere it is charac- 
terized by the forms just enumerated ; many of the same occur at 
Llandewi Velfrey, Lampeter Velfrey, and Musclewick Bay. The 
Upper Llandeilo is also distinguished by special forms of Trilobita 
—Barrandia Corda, B. longifrons, Cheirurus Sedqwickit, Ogygia 
Buchu, Calymene duplicata, Ampyx nudus, and Agnostus M‘Coyit. 
It will be seen that each division is specialized or can be discri- 
minated by its Crustacea; the well-known Ogygia Buchu and 
Cheirurus Sedgwicki, here as elsewhere, are the typical forms in 
the upper black argillaceous slaty flaggy sandstones. The middle 
yields Asaphus tyrannus, A. peltastes, and Ogygia convexa, &c. ; and 
the Lower Calymene Murchisome and Trinucleus Ramsayi; and 
this grouping holds good wherever the Llandeilo beds occur. Three 

* Quart. Journ. Geol. Soc. vol. xxxi. pp. 177-180. 

VOL. XXXVI. 


118 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


species only of Trilobita have occurred in the Scotch Llandeilo rocks, 
viz. Lllenus Bowmannu, Salteria involuta, and S. primeva. Ire- 
land has 5; but probably many more will occur. I enumerate them 
so far as we know them—Acidaspis James, Ampyx mammillatus, 
Ogygia Portlocku, Phacops truncato-caudatus, and Avglina mirabilis. 
I know no others, although in all the three Llandeilos we have 45 
species of Crustacea. Shropshire has only yielded 11 species, and 
these chiefly from the Llandeilo rocks of Shelve: they are all marked 
species. In the Builth and Llandeilo areas no species occurs below 
the tuff-beds of the Lower Llandeilo. Few species of Graptolites 
are common to the Llandeilo and Arenig rocks of St. Dayid’s, al- 
though here the succession is so complete, and no less than 18 
species there occur in the Arenig and 6 in the Llandeilo. Only 16 
species of all groups are common to both formations, although the 
Arenig fauna numbers 149 and the Llandeilo 175, thus clearly 
showing the faunal differences. 

Llandeilo beds are known in Merionethshire, Caernarvonshire, 
Denbighshire, and Anglesey. Fifteen species have been determined 
from Merioneth; 13 from Garn, east of Arenig, by far the largest 
fauna in North Wales. Anglesey from five localities yields 15 
species and 18 occurrences, or only 3 species for each locality ; 
Caernarvonshire, from four localities, 17 species (the chief loca- 
lity, Teddyn-Dicum, has produced 8 species); and Craig-y-Glyn, 
near Llanrhyader in Denbighshire, 6 species. No one can doubt, 
from the above results, that careful research would greatly add to 
the faunal value of the Llandeilo rocks of North Wales. Prof. 
Hughes has lately succeeded in ascertaining the presence of Tre- 
madoc fossils in Anglesey ; and further search at Garn, east of the 
Arenigs, could not fail to throw much light upon both the Arenig 
and Llandeilo faunas, and to substantiate still more the relation 
between North and South Wales. As yet, out of 13 Garn fossils, 
we know only 1 Trilobite (Ampya mammillatus). In Anglesey, 
near Llanerch-y-medd, out of 5 species collected 3 are characteristic 
Trilobites—Phacops apiculatus, Asaphus Powisir, and Calymene cam- 
brensis. It is the same with Treiorwerth in Anglesey,—clearly 
showing that a rich harvest of Llandeilo species is yet to be ob- 
tained in North Wales. 

The elaborate researches of Mr. Lapworth, in Scotland*, into the 
history of the Rhabdophora of that and other areas, has greatly 
modified our views respecting their distribution and range. 

The oldest fossiliferous strata known in Scotland are the Grap- 
tolite shales forming the well-known Moffat series, a group of black 
shales about 600 feet in thickness, and separated by Mr. Lapworth 
into three distinct paleontological divisions. The lowest (Lower 
Moffat or Glenkiln Shales) he determines to be of Upper-Llandeilo 
age; the Middle Moffat or Hartfell Shales above he correlates with 
the Bala group of North Wales “as an attenuated representative ;” 
and the third or uppermost (the Upper Moffat or Birkhill Shales) 
represents the Lower Llandovery. This last determination has an 

* Quart. Journ. Geol. Soc. vol. xxxiv. pp. 240-346. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 119g 


important bearing upon the distribution of the Graptolithina or 
Rhabdophora, it being well known that in the Llandovery rocks of 
Wales no Graptolites occur. The Lower Llandovery of Cardigan and 
Merioneth, of such great thickness and extent (yet without Grap- 
tolites), has ‘‘ dwindled down to the thickness of the Birkhill Shales 
(about 70 feet) in the intervening Lake-district, where it constitutes 
the Coniston Mudstones—a group of beds almost identical in thick- 
ness, lithology, and paleontology with the equivalent Scottish deposit 
of the Birkhill Shales” (Lapworth). The conclusion arrived at by 
Mr. Lapworth, that the oldest beds of the south of Scotland (the 
Glenkiln Shales) are the equivalents of the highest Llandeilo rocks, 
is borne out through his researches amongst the Moffat series 
in Southern Scotland; and, with him, we must come “to the 
conclusion that the Lower Silurian rocks of the southern uplands 

can be arranged in two distinct formations, viz. a lower and very 
- thin group of fine-grained Graptolite shales, and an upper and com- 
paratively massive series of arenaceous strata. In their mineralo- 
gical features and paleontological characteristics the Moffat series 
differ from any of the typical Silurian rocks of the principality.” 
Unlike the rocks of Wales, they are almost exclusively Graptolitie 
in their fossil contents, scarcely any Brachiopoda or Crustacea being 
known throughout the series. The Llandeilo age of the Lower 
Moffat shales is further determined through the total absence of those 
complex Arenig forms of “ Dichograpti, Tetragrapti, and Phyllo- 
grapti, so characteristic a feature of the Graptolitic fauna of the 
Skiddaw, Shelve, St. Dayid’s, and Canadian equivalents.” 

Mr. Lapworth, “ arranging the Moffat strata in the order in 
which they are displayed, and distinguishing each chief band by its 
peculiar fossil,” gives the following Table :— 


(Upper Birkhill | Zone of Rastrites peregrinus. \ 
ir or Grey Shale | Zone of Monograptus spinigerus. 
Soa group. Subzone of Diplograptus cometa. | Lower 
Birkhill oe 4 \ 
State Lower Birkhill |) Zone of Monograptus gregarius. Llandovery. 
f or BlackShale + Zone of Diplograptus vesiculosus. | 
group. Zone of Diplograptus acuminatus. _) 
pees eel Zone of Dicellograptus anceps. | 
irae Zone of Barren Mudstones. ; 
Hartfell 4 stone group. _ { Bal 
. Sh, i. \ Lower Hartfell ) Zone of Plewrograptus linearis. r i 
a“es- | or Black Shale | Zone of Dicranograptus Clingani. | 
{ group. Zone of Climacograptus Wilsoni. }} 
Upper Glenkiln 
a Fe tiesto | Didymograptus beds. Upper 
Tteq, | Lower Glenkiln ) p. Llandeilo. 
Shales. SPAlos Ribbed mudstones. 


The facies of the Glenkiln shales is distinctively that of the Llan- 
deilo of South and North Wales, the assemblage being the same: 
differences occur dependent upon locality and the mineral composition 
of the beds; but the common Glenkiln forms of Scotland, and those 
species collected by observers in North Wales (Messrs. Salter, Hop- 
kinson, Homfray, Gibbs, &c.), serve at once to connect and correlate 
the faunas of the two areas. 10 species from South Wales and 12 
from North Wales are all Glenkiln species and of Llandeilo age; 

k 2 


I20 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


and, adopting the views of Mr. Lapworth, from the absence of 
Didymograptus Murchison and the genus Phyllograptus in the black 
shales of the Tremadoc area, and the presence of some succeeding 
Bala forms, we may infer that these dark Tremadoc shales and the 
Glenkiln beds are probably of higher-Llandeilo age. This inference 
is further strengthened by researches made by Linnarsson in the 
Llandeilo strata “of Central Sweden, where the greater portion of 
the Llandeilo formation of Britain is represented by the Orthoceras- 
limestone.” ‘‘ Upon this limestone rests the sheet of dark shales 
named by Dr. Linnarsson the ‘ Middle Graptolite schists; the 
lowest part of these schists, according to Dr. Tornquist, affords 
abundant examples of Phyllograptus and the Murchisoni form of 
Didymograptus. The highest Swedish beds assigned or referable to 
the Llandeilo formation afford both the peculiar Scotch or Glenkiln 
and North-Wales forms, such as Didymograptus swperstes, Lapw., 
Cenograptus gracilis, Hall, Climacograptus Scharenbergi, Lapw., C. 
perexcavatus, Lapw. 

Mr. Lapworth further discusses the relations and affinities of the 
Llandeilo species of South Scotland (Glenkiln Shales) with the dark 
shales and flagstones of the Cincinnati group which bound the valley 
of the Hudson near the. city of Albany and everywhere underlie 
the Trenton limestone; he shows reason for believing that we 
may regard them as forming the ‘“ highest division of the Quebec 
group (Arenig) which emerges unconformably from below the hori- 
zontal Trenton limestones. ‘The Hudson-River shales may therefore 
represent the higher Llandeilo beds of Britain.” Lapworth names 
14 species collected from Norman’s Kiln in the valley of the Hudson, 
on the Marsouin river, &c., as being identical with the Glenkiln and 
Welsh forms. He still further establishes representative affinity 
between the peculiar Glenkiln species which are absent from the 
American strata and others which occur in them. Thus :— 


BrirTisu. AMERICAN, 
Glossograptus Hincksti is represented by Glossograptus ciliatus, Hmm. 
Dicranograptus ziczac bs »  Dicranograptus furcatus, Hall. 
Climacograptus Scharenbergi  ,, »  Climacograptus scalaris, His. 
Clathrograptus cuneiformis  ,, »  Clathrograptus Geinitzianus, Hall. 


‘‘ All evidence, therefore, tends to prove that the Glenkiln shales 
belong to the age of the Upper Llandeilo series, and are closely allied 
to the North-Wales grouping.” From the absence of certain Dzdy- 
mograptt and Phyllograpti from the Glenkiln division, and their 
abundance in the lower beds of the Llandeilo of Wales and Sweden, 
we are obliged to assign the Scotch Glenkiln Rhabdophora “ to the 
highest division of the Llandeilo, and not far below the base of the 
Caradoc or Bala’’*. 

It is of the Hydrozoa, Crustacea, and Brachiopoda, which together 
make up the 123 out of the 175 known species, that the mass of 
the fauna of the Llandeilo is composed. The same classes largely 
transmit their genera and species to the succeeding Caradoc or Bala 

* For an exhaustive paper upon the Graptolites of the Moffat series, by 
C. Lapworth, Esq., F.G.S., see Quart. Journ. Geol. Soe. vol. xxxiv. pp. 240-343. 


The general characters of the Lower Silurian rocks of Scotland and the strata of ~ 
the Moffat district are also ably discussed. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 121 


rocks, viz. 16 species of Hydrozoa, 17 Crustacea, and 23 Brachio- 
peda, or, together, 56 out of the whole 76 that connect the two 
formations. It is interesting to know also those areas that are spe- 
cifically most prolific in the above groups, and thus help us to draw 
conclusions as to the nature of the sea and its bathymetrical condition 
during the time of the deposition of the Llandeilo and succeeding 
Caradoc, which are more intimately related than is generally believed. 
United, the Llandeilo fauna of North and South Wales is constituted 
by 94 species; or this is the known number for Wales, 88 occurring 
in South and 47 in North Wales, and 41 being common to both, 
chiefly among the Hydrozoa and Crustacea. Upon whatever hypothesis 
we may account for the physical relationship between the Llandeilo 
rocks of Scotland and Ireland, the connecting species are few (only 
27); and no Crustacea are common to these two countries—which, 
out of a total fauna of 45 species, is remarkable. ‘The Irish species 
number 6, 3 of which are also South-Wales forms—Ogygia Port- 
locku, Calymene duplicata, and Aiglina mirabilis. As before stated, 
Scotland has hitherto yielded only 4 genera and 5 species of Crus- 
tacea—lilenus Bowmann, Salteria wmvoluta, S. primeva, Peltocaris 
aptychoides, and Discinocaris Browniana. 

Praxta.—None. 

Proroz0a.—Ischadites antiquus, Salt. No other Protozoon has 
been recognized in the Llandeilo; it is not a common genus in the 
Lower Silurian rocks. 

Hyprozoa.—The Llandeilo beds of Scotland appear to be the most 
prolific in Rhabdophora. Our knowledge of them is due to the 
researches of Nicholson, Lapworth, and Carruthers in Scotland, all 
of whom have done able work in elucidating the history and struc- 
ture of this obscure class. Forty-four species illustrate 18 genera ; 
but it is chiefly through the larger genera that this Graptolitic fauna 
is represented. South Wales, as we should expect, is also represen- 
tative, although poor, as compared with Scotland, having only 18 
species, or about half the number. North Wales has 7 species, and. 
Treland 28. The Welsh and Irish genera have not the same specific 
value; generally they are much the same, but illustrated by fewer 
species. This may be due to want of systematic search in Wales, 
such as the rocks of Scotland have undergone by the above-men- 
tioned authors. The genera Didymograptus, Diplograptus, Climaco- 
graptus, and Dicellograptus are those chiefly occurring in South 
Wales; and Diplograptus, Dicellograptus, and Climacograptus in 
North Wales, &c. The Llandeilo rocks of Ireland contain species 
of nearly all the above genera. ‘Thus of the 44 known Llandeilo 
Rhabdophora we have 


Species. 
Ouse WMeS ee. 6.4. Sooke craye Bue 18 
INO rb NWiale sues cc Bac a ee o 
SCO bam sees tie eR Bu 
Nereida See wi,.. & wa vets 28 


85 occurrences ; 
and 16 species pass to the succeeding Caradoc. 


122 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Actrnoz0a.—Halysites catenulatus, Linn., Favosites fibrosus, Goldf., 
and Monticulipora favulosa, Phill., complete the list of the Llandeilo 
Actinozoa. No English form is known. North Wales yields Favo- 
sites fibrosus. All three occur in South Wales, and Fuvosites fibrosus 
in Ireland. Scotland has no representative. All three species pass 
to the Caradoc. These are the first corals occurring in the British 
rocks ; but they expand to 20 genera and 42 species in the succeeding 
Caradoc. We cannot admit that change of condition only thus 
favoured the growth and development of the Actinozoa. It is true 
little limestone occurs in the Llandeilo, and that little in the upper 
division; but the organic remains in the Upper Llandeilo limestones 
are chiefly Crustacea. The increase of limestones and calcareous 
matter in the Caradoc rocks being due to life agency, we may 
attribute much of it to the secretion of carbonate of lime by the 
Coelenterata. The only notable or appreciable form is Halysites, 
which was abundant. 

Ecutnopermata.—We know of only 4 genera and 4 species of 
this class, which, like the Actinozoa, may be said to first appear in 
the Llandeilo. None are known in the Arenig below; but 4 species 
(1 Menevian and 3 Tremadoc) in fragments have been preserved in 
older formations. The 4 Llandeilo species are Actinocrinus Wynne, 

ichinospherites granulatus, Paleasterina Kinahan, and Glypto- 
crinus basalis. The first 3 are Irish; Glyptocrinus occurs in the 
South-Wales Llandeilos. Two of them passed to the Caradoc. As 
with the Actinozoa, this group became greatly developed in the 
Caradoc, no less than 15 genera and 32 species enriching that for- 
mation. ‘These, like the corals, must have lived in a moderately 
deep sea well charged with calcareous matter. 

ANNELIDA.—It is questionable if more than 6 species (although 
9 are placed in the Llandeilo) occur: Paleochorda teres, P. major, 
and P. minor may be Arenig; they are very sparingly distributed, 
only 1 species occurring in England, South Wales, and North Wales, 
and 2 in Scotland. I know of no Irish Llandeilo Annelida. 

Crustacea.—In this class the Trilobita are of high significance, 
in both the Llandeilo and Caradoc formations, especially the latter, 
in which we know of more than 100 British species. The Llandeilo 
rocks have yielded 18 or 20 genera and only 45 species, or upon 
the © igs only about 2 species to each genus. Many genera are 
represented only by 1 species; these are Homalonotus (H. bisul- 
catus, Salt.), Lichas (L. patriarchus, W.-Kdg.), Stygina (S. Murchi- 
sonic, Murch.), Oheirurus (C. Sedgwickit, M‘Coy), and Acidaspis (A. 
Jamesti, Salt.). Much has yet to be done by collectors to increase 
our knowledge of the specific crustacean fauna of the Llandeilo rocks. 
South Wales being the classical locality for the Llandeilo group, it is 
natural that we should expect to find there the largest and most 
typical fauna, crustacean or otherwise. In one group (Graptolites) 
we have seen that this is not the case, Scotland being most repre- 
sentative. Here, however, the Crustacea of South Wales (29 
species) are double those of North Wales (15), and ten times more 
than those of Scotland, which has yielded only 3 species, conditions 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 123 


being more favourable for the support and distribution of certain 
forms of life in one area than the other. The only 3 Scotch Llan- 
deilo species known to me are Jilwnus Bowmanii, Salt., Salteria 
wnvoluta, Thom., and S. primeva, Thom. The 6 Irish species 
are Acidaspis Jamesii, Ampyx mammillatus, Ogygia Portlocki, 
Phacops truncato-caudatus, Calymene duplicata, and Aiglina mirabilis. 
The English forms are chiefly from the Shelve area; and 8 out of 
the 10 are South- Wales species. 

The following occupy and characterize definite areas; they are 
single representatives of the genera named :— 


ae aoe England.| Scotland.) Ireland. 


Homalonotus bisulcatus...... * 
menostus Mi Coyi) s....-1.-..- Pe * 
Stygina Murchisone ......... : * 
Cheirurus Sedgwickii......... soo * 
Acidaspis Jamesii ............ Sas a wh Acts % 
Lichas patriarchus ............ "stg * 
1 4 il 


Thus 6 Llandeilo genera are illustrated by only 1 species each, 
England and Scotland having no really marked or characteristic 
form. Nevertheless there are important Llandeilo species which 
are entirely restricted to that formation, not ranging either lower 
or higher. The most important are :— 


North | South | 
Wales. Wales, Hie Tana (Secktind | Ireland. 


Illznus Murchisoni ......... 
Ogygia angustissima ......... 
—— Portlockii ............... 
——= COINCeENSIS .........0.000. 
AD WCHM ays slag ost eee aides 
COMVOMAN io. oe cc cece: 
Asaphus tyrannus ............ 
EMGASTES) seo askc.dsectewe- 
datveostatus.....-.sc-.9a<s: ; 
Calymene cambrensis ...... * 
Trmucleus Lioydu............ 
IUAMING AWG eicline seen delice 
MUSEUM ES . castscce. ves 
VENTE! Gate GOns ee ermprerere 


KK OK 


kK KK Ss 


* OK KOR: 


5 re 


KK OK OK kK KS ft 
KOK 


Ks ° 


* 
Kk 


% 
AGIIAMGP sere aeecd sew elelse we 3 
* 


I24 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


All the above 20 species are confined to the Llandeilo and to the 
geographical areas assigned to them, and they are essentially cha- 
racteristic ; the remaining 15 species pass to the Caradoc, and are 
therefore of no value stratigraphically. To complete the analysis 
of the distribution and value of the Crustacea I name the 15 species 
that connect the Llandeilo and Bala formations :— 


Acidaspis Jamesii. Calymene Blumenbachii. 
Homalonotus bisulcatus. brevicapitata. 
IWenus Bowmanni. Trinucleus fimbriatus. 
Ampyx mammillatus. —— concentricus. 
rostratus. Phacops apiculatus. 
nudus. truncato-caudatus. 
Asaphus Powisii. /églina mirabilis. 


Calymene duplicata. 


The singular Phyllopod Peltocaris aptychoides occurs both in Scot- 
land and North Wales, and is not known out of the Llandeilo beds. 

Bracutopopa.—This class is an important factor in the Llandeilo 
rocks; 10 genera and 34 species are known. Five new genera appear, 
Acrotreta, Orania, Rhynchonella, Strophomena, and Leptena; the 
first 4 with a single known species only; but Leptena has yielded 
4 species, all of which occur in South Wales, and 2 of them in North 
Wales. As we should anticipate, the species are most numerous in 
South and North Wales. The former area gives us 26 species, the 
latter 18 ; Scotland 9, and England 6. Although there are 20 species 
in the Arenig group (and 34 in the Llandeilo) yet no species 1s common 
to the two formations, no Arenig form passes up; but of the 34 
Llandeilo forms, 23 pass to the Caradoc. ‘This is chiefly through 
the genera Lingula, Orthis, and Leptena—A4 out of 6 in the former 
genus, 11 out of 13 in Orthis, and all the known 4 in Leptena. 
We have already seen that of the whole fauna of the Arenig (150 
species) only 17 pass up into the Llandeilo; whereas no less than 72 
out of 175 Llandeilo species are found in the Caradoc, viz. Hydrozoa 
16 species, Crustacea 17, Brachiopoda 23, Bivalvia 3, Gasteropoda 4, 
Heteropoda 5, and Cephalopoda 2. The remaining forms are dis- 
tributed through smaller classes. But for the distinctive character 
of the Abereiddy-Bay and Caermarthenshire faunas, there is much 
that is common between the Llandeilo and Caradoc. 

LAMELLIBRANCHIATA.—With the exception of the Irish species 
Pleurorhynchus calcis, Baily, allthe Bivalye Mollusca known are from 
the South-Wales Llandeilos. We know of no Llandeilo Pelecypod 
mollusk either in England or Scotland. Cardiola interrupta occurs 
in both North and South Wales; Modiolopsis expansa, M. inflata, 
Ctenodonta varicosa, and Palearca amygdalus only in the South- 
Welsh beds. Three pass to the Caradoc. Pleurorhynchus and Car- 
diola are new genera. 

GastERopopa.—South Wales has hitherto only given us 1 species, 
Murchisonia simplex, M‘Coy; Ireland 2, Huomphalus parvus, Portl., 
and Twi'bo tritorguatus, M‘Coy; the Shelve area and North Wales 
Cyclonevva crebristria, M‘Coy, and Ewomphalus corndensis, Sow., the 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 125 


remaining 7 being from the Durness limestone of Scotland (they may 
be Arenig). The Scotch species are Huomphalus matutinus, Hall, 
Murchisoma angulocincta, Salt., MW. angustata, Hall, WM. bellicincta, 
Salt., MZ. gracilis, Hall, Ophileta compacta, Salt., and Pleurotomaria 
thuli, Salt. Three of the 12 pass to the Carodoc. Huomphalus, 
Cyclonema, and Murchisonia are new genera, and first appear in 
the Llandeilo. 

Prrropona.—Three species of T’heca—T. cometoides, Baily (Irish), 
T. reversa, Salt., and 7’. caereesiensis, Hicks (from St. David’s). No 
species occurs in England, North Wales, or Scotland. 

Hurrrorops.—sScotland possesses 5 of the 7 species known ; and 
4 of the 5 are Macluree. These are associated with the Gastero- 
poda in the Durness limestone. Bellerophon perturbatus is a North 
and South Welsh species ; and B. bilobatus, English, North and South 
Welsh also. Huomphalus scoticus is from the Ayrshire rocks, The 
3 last-named species and Heculiomphalus pass to the Caradoc. 

CrpHALopopa.—Of the 3 genera and 7 species none occurs either 
in North Wales or Ireland. The only English species is Orthoceras 
Avelani?, Salt., from the Shelve area (Shropshire) ; Endoceras eoum, 
Edgell, is from South Wales. The remaining 5 of the 7 known 
species are Scotch, including the singular genus Piloceras (P. in- 
vaginatum) from the Durness limestone. 

The accompanying Table (p. 126) gives the following results. 
The 12 zoological groups occurring in the Llandeilo rocks are re- 
presented by 80 genera and 175 species, which are numerically 
distributed as follows :— 


England .... 18 genera and 26 species. 
North Wales; 30. ieee 3 
South Wales.. 47 ,, RS OR: =. 
Scaulandiy hey sok 
itrelandiy ss 24: SOs SAU Pg 


38 genera and 73 species of the 175 pass to the Caradoc. It 
will be seen that it is through the Hydrozoa, Crustacea, and Bra- 
chiopoda that the community of species largely occurs between the 
Llandeilo and Caradoc generically and specifically—through the 
Hydrozoa 9 genera and 16 species, the Crustacea 9 genera and 
17 species, and the Brachiopoda 7 genera and 23 species. ‘The re- 
maining passage numbers are seen in Table IX. 


From Arenig. 


AL 


© bo 


© 0 


126 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Taste [X.—Llandeilo. 


Geographical Distribution. 


n a 3 
oS oi ro a a a) cs = 
: 2 = > | & = a se 
Classes. © 5 = a = Ss ae 
; 2 ep = o Ay & 
5) 2, = =| pics| ° LY ‘S) 
O wD = 3 = = 
cS B Es R = ° 
i) - 
Zz mM 
Plante. 
(PEOLOZOd: Wiaewae eee 1 1 a 
eerseeceeoree 4 5 1 4 1 3 Z 4 2 
Hydrozoa 18 44 a : 18 aS 28 16 
. 1 3 3 
ACHINOZOR 2. e2s se 3 3 ae : 3 A 3 
Echinodermata...... i 4 1 is } cae d 2 
j 1 1 i 2 
AMMeli Gay seacoast 6 9 2 i 1 3 
sta 6 ie) 12 4 9 
Crustacea ele Seiwidiolalai ovate 20 45 10 ; 5 29 5 é ry 
Bryozoa. 
ng ahi 2 6 8 4 4 Cf 
Brachiopoda ......... 10 34 r 18 25 9 9 23 
Lamellibranchiata. . 5 6 fh A 4 1 3 
2 1 2 2 
Gasteropoda ......... 6 12 2 1 i 7 2 3 
1 1 1 
Pteropoda ....d24.065% 1 3 de B. A ies 7 : 
1 1 1 2 2 
Heteropoda ......... 3 a ; 2 2 3 i 4 
Cephalopoda......... 7 ; 1 é 1 
Wh 18 27 47 31 31 38 
aogooosas 80 1 (Oo 26 47 87 66 51 73 


Carapoc or BAta. 


The rocks overlying the Arenig and Llandeilo groups, and under- 
lying the May-Hill Sandstone, have received much critical attention 
from Sedgwick, Murchison, M‘Coy, Salter, and the officers of the 
Geological Survey. The facies of the fauna is that of the Llandeilo, 
but greatly developed. The increased fauna of the Caradoc is 
scarcely recognized until subjected to critical research and analysis. 
The difference in the fossil contents of the two groups is due chiefly 
to age and the conditions under which they were deposited. Sedg- 
wick, in 1853, placed the Bala group in his ‘‘ Upper Cambrian ” 
division, dividing it into Lower, Middle, and Upper Bala; it in- 
cludes also the ‘Cambro- Silurian of some authors, a term used or 
adopted by Prof. Jukes in his ‘ Manual of Geology.’ 

From the year 1832 to 1853 Sedgwick included the Caradoe in 
the term “ Lower Bala,” not ieee then the existence of the 
term Caradoc. Sir H. de la Beche, Prof. J. Phillips, and the Survey, 
from 1840 to the present date, designate this formation the ‘Caradoc 
or Bala.” Murchison, from 1831 to 1859, used the term “‘ Caradoc 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 127 


Sandstone.” The term Llandeilo was not admitted by Sedgwick 
either in the ‘ Synopsis’ or the Woodwardian Catalogue ; therefore 
great difficulty was and is still felt as to the identification of species 
long ago collected in certain localities, especially those having 
reference to the Lower “ Llandeilo of Murchison ” and the Arenig 
of Sedgwick; the faunas of all three groups require careful revision, 
in consequence of old or early errors, which of necessity demand it. 

Sir Roderick Murchison, in 1833, first noticed the ‘ Caradoc 
Sandstone ”’*; in 1834 the same rocks were described by him under 
the name of the “ Horderley and May-Hill Sandstone.” In the 
‘Silurian System,’ subsequently published, these strata were called 
‘Caradoc Sandstone,” the name being based upon their being highly 
developed in the neighbourhood of Caer Caradoc. 

In the early history of the Caradoc rocks the fossils of the 
Pentamerus-beds were included in its lists. The Survey subsequently 
corrected this, by separating the upper part or highest beds of the 
Caradoc. In 1852 Prof. Sedgwick and Mr. Salter in part corrected 
this error, and showed that the fossils of the May-Hill group were 
very distinct from those of the Caradoc Sandstone. Again, the 
upper Pentamerus-beds rest unconformably on the true Caradoc 
Sandstone, and the whole pass under the Wenlock Shale. In 
Shropshire and at Builth the unconformity is visible; and in the 
Malvern area, west of the Herefordshire and Worcestershire Beacons, 
the Upper Llandovery beds lie directly upon the Upper Lingula-flags 
or “‘ Dictyonema-shales.” Again, the Upper Llandovery beds on the 
banks of the Onny river lie on the upper part of the Caradoc 
or Bala beds. West of Wenlock Edge they cover the nearly vertical 
edge of the Cambrian or Longmynd rocks. “ Probably,” says Prof. 
Ramsay, “there is no unconformity so complete yet observed in 
other members of the British Silurian strata.” Besides the researches 
and large collections of fossils made by the Survey from these 
rocks, the labours of Sedgwick are preeminently associated with the 
Caradoc and Bala beds of Wales, through the great memoir on the 
British Paleozoic Fossils, in which he was so well aided by Prof. 
M‘Coy, and the “ Woodwardian Catalogue” ft prepared by Salter 
from the great store of materials in the Woodwardian Museum at 
Cambridge, and which, many years previously, he had helped 
Sedgwick to collect, and name, from the classical localities in Wales. 
To Prof. Ramsay, for his valuable memoir on the Geology of 
North Wales+, every student is deeply indebted. In this great 
treatise every detail relative to the physical history and distribu- 
tion of the Caradoc and Bala rocks and fossils is treated upon ; 
and to the appendix, originally compiled by Mr. Salter, I have 
greatly added, especially in that portion treating of the distribution 
of life, not only for the Caradoc, but through the whole of the for- 


* Proceedings of the Geological Society, 1833, vol. 1. p. 476. 

t ‘A Catalogue of the Collection of Cambrian and Silurian Fossils contained 
in the Geological Museum of the University of Cambridge.’ By J. W. Salter, 
F.G.S. With a Preface by the Rev. A. Sedgwick, LL.D., F.R.S. 

¢{ Mem. Geol. Surv. of Great Britain, vol. iii. Geology of North Wales. 


128 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


mations of North and South Wales. The portion of the memoir by 
Prof. Ramsay devoted to the Caradoc is of necessity large, arising 
from the extent, magnitude, and importance of the Caradoc rocks 
as developed in North and South Wales, Westmoreland, Ireland, 
and Scotland. I therefore, as in the case of the older groups, com- 
pare or attempt to correlate other portions of Britain with the 
typical area in Wales, so as to show the present aspect or distribu- 
tion of the Caradoc fauna, which is so largely developed in the 
British Islands. To devote much space to the purely geological 
aspect of the Caradoc would be needless under present circum- 
stances, as so much has already been done by able observers in the 
pages of our Journal, as well as in the exhaustive memoir on North 
Wales by Prof. Ramsay. My tabular results in the appendix to the 
above memoir, relative to the distribution of the Caradoc fossils both 
in time and space, will, when published, embody almost all the in- 
formation known. ‘The result, however, will be referred to here, as 
being a complete analysis of the 600 species. It is scarcely neces- 
sary for me to discuss the views of authors as to very minute sub- 
divisions of the Caradoc group in any given area or under any 
peculiar condition ; such will always arise under critical examina- 
tion, extended knowledge, or constant research ; and large as we know 
the fauna to be, owing to the rocks being so extensively worked in 
the days of Sedgwick and Murchison, and through the long-con- 
tinued labours of the Geological Survey, yet many of the zoological 
groups are still being added to. This is notably the case with the 
elaborate memoir upon the Girvan fossils by Prof. A. Nicholson and 
Mr. Etheridge, jun., the first volume of which, containing this 
addition to our knowledge of the Caradoc fauna of Scotland, is just 
completed and published. In this work the authors describe 41 
genera and 65 species, many of which are new. Perhaps, with the 
exception of Mr. Lapworth’s paper on the Graptolites of the Moffat 
series at Girvan and Glenkiln* (mostly Lower-Bala or Llandeilo 
_ forms) no more important addition to our knowledge of the Caradoc 
fauna has been made since M‘Coy completed his great work upon 
the British Paleozoic fossils. 

The legends attached to the published maps of the Geological Survey, 
and the explanation of the colours employed to designate the horizons 
or formations surveyed, show that no attempt was made by the Survey 
to divide the Caradoc rocks into subgroups, or into Lower, Middle, and 
Upper Caradoc or Bala; neither does Prof. Ramsay, in that part of 
his memoir devoted to the Caradoc rocks, attempt any subdivision, 
but masses the whole group between the top of the Llandeilo and 
the overlying Llandovery. In the Catalogue of the Cambrian and 
Silurian fossils in the Woodwardian Museum, Cambridge, Prof. 
Sedgwick has divided the Bala beds into three subgroups, placing 
them in his Upper Cambrian stage (the Cambro-Silurian of some 
authors), the Lower Silurian of Murchison. The grouping adopted 
in this catalogue is such as to eliminate the true Llandeilo fossils, 


* Quart. Journ. Geol. Soc. vol. xxxiy. pp. 249-346. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 129 


and place them in his Lower Bala group. The Bala group, or 
Upper Cambrian of Sedgwick, consists therefore of 


( Lower Bala= Llandeilo flags (Upper Llandeilo of the 
| Geological Survey, the Arenig being the Lower). 
Upper | Middle Bala=the Caradoc Sandstone and Bala rocks 
Cambrian *. § (Geological Survey and Sir R. Murchison). 
| Upper Bala=the Caradoc Shales, Hirnant Limestone, 
| and Lower Llandovery rock (Geological Survey). 


The Upper Llandovery or May-Hill Sandstone unconformably over- 
lies these. 

Sedgwick’s Middle Bala, and part of the Upper, are therefore 
equivalent to the true Caradoc of the Geological Survey; and as 
such I treat that group here; or the Middle Bala of the Cambridge 
Catalogue will embrace the whole of the 600 species known as 
Caradoc forms. I have already discussed the Llandeilo formation 
proper in the sense now used by the Geological Survey, having 
relegated those species hitherto called Lower Llandeilo to the Arenig 
group—the Middle Cambrian of Sedgwicky. The Lower Bala 
includes certain dark earthy slates and bands of limestone occurring 
on the east flank of the Arenigs, Mynydd Tarw and Craig-y-glyn 
above Llanarmon in the Berwyns, the black slates on the flanks of 
Snowdon, also the arenaceons deposits on the west side of Bala Lake 
below the Bala Limestone, and that limestone also. 

The Middle Bala group of the Woodwardian Catalogue embraces 
the Bala Limestone and its associated sandstones and slates in North 
and South Wales. In Shropshire it is the Caradoc Sandstone, with 
its Horderley Limestone. The Coniston and Kildare limestones, and 
the Craig-Head and Peebles limestones, are all of this aget. 

Sedegwick’s ‘“‘ Upper Bala” comprehends the Aber-Hirnant beds 
above the Bala Limestone (with peculiar fossils), the lower part of 
the Coniston Flags (that conformable to the hmestone), and all beds 
above the Bala Limestone and beneath the May-Hill Sandstone. 
Again, the ‘‘ Upper Bala” includes all the beds, whether near 
Meifod or Welchpool or Llanwyddyn, which lie above the Bala 
Limestone and under the unconformable cover of the Denbighshire 
grit and flag. In ascending order the Upper Bala includes 


(1. The Hirnant Limestone and slate=Coniston Flags, 
| lower part only (Ashgill, Coldwell, &c.), above 
the Coniston Limestone. 
(Sedgwick). | 2. Llandovery beds (Lower Llandovery of Murchison) 
2 eh =the Mathyrafal Limestone, near Meifod, of 
\ Sedgwick. 

“Tt is the great fossiliferous group of Haverfordwest. The 
Daiquorhan and Mullock beds in Ayrshire, the fossiliferous rocks of 

Maume and Cong in Galway belong in part to it’’§. 


* Vide Woodwardian Catalogue, Cambridge, pp. 25, 26, &e. 

T Consult the table of equivalents of the strata underlying the May-Hill 
Sandstone, Woodwardian Catalogue, p. 25. 

t Vide Woodwardian Catalogue, pp. 26, 39. § Loe. cit. pp. 26, 72. 


130 PROCEEDINGS OF THE GEOLOGICAL SOCIETY, 


It has been stated that the fauna of the Llandeilo rocks num- 
bers 80 genera and 175 known species. The Caradoc or Bala 
contains 179 genera and 613 species; or 88 new genera and 375 
new species make their appearance either at the close of the 
Llandeilo or during that period which ushered in or brought about 
those changes of conditions under which the deposits of the Cara- 
doc sea were accumulated and its species developed and multiplied, 
covering as it does so extensive an area in Kurope and Britain, also 
extending to America, whether homotaxially in that region or con- 
temporaneously with the deposition in the British area it is difficult 
to say. That the Trenton and Hudson-River group of North 
America, and nearly the whole of the Ktage D of Barrande (Bohemia) 
and Regio D of Angelin (Sweden), are the equivalents of our great 
Middle Caradoc and Bala, or the true Caradoc, is now not doubted. 
The fauna of the Bala group, especially in its zoological groupings, 
differs much from the preceding Llandeilo, although many species 
(85) are common to the two formations. 

I purpose treating of the Caradoc rocks and their fossils at some 
leneth—their stratigraphical place, the geographical distribution of 
their fossils (especially with reference to the succeeding Lower and 
Upper Llandovery)—questions which demand critical examination 
or analysis. 

The three classes having the greatest number in common are the 
Hydrozoa (Rhabdophora), the Crustacea (Trilobita), and the Bra- 
chiopoda. The number of known species in the Llandeilo in each 
of the above classes is 


Ey droZz 0a; tert ven.rteee ee 44 
Crugtaces ene ee 45 
Brachiopeda 1.232% .): 34 


In the Caradoc fauna the same three classes number, including 
those from the Llandeilo, 


Hydrozoay 2a 38 
Crustacea y cee mnne 146 
Brachionodan @siosaec. 109 


Numerically, the remaining ten classes have little value for compa- 
rison, although the Actinozoa, Echinodermata, Lamellibranchiata, 
and Gasteropoda are well illustrated (see Table X.). It is through 
the above three extensively developed groups, which are mainly 
moderately deep-sea forms, that the two formations are united. 

The fact of 73 species out of 175 passing from the Llandeilo to 
the Caradoc is highly suggestive, and is confirmatory of the views of 
Sedgwick in uniting the Llandeilo to the Lower Bala; and the 
distinctiveness of the great Middle Bala is confirmed by the fact 
that, out of 610 species in that formation, only 102 pass to the Lower 
Llandovery. Therefore, on paleontological data, we may regard the 
Llandeilo and Caradoc as being most closely allied. The 102 trans- 
gressive Caradoc species are distributed through the following 
classes :— 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 131 


12h: Y.-C) 1 out of 10 known. ) 

FIVOEOZOG. 2. i nus ly PETG) 

Colenterata ...... Die eri GOs, | 

TES Cree Often tN: 

iRehinedcrmata ucts 721) 496 (O20 ols, 

Weushacka, uiis.6 NG. De eA Pass to TL, 

12 \1(07 20 er eee ee Al Dial 3 = ‘ d ie 
Pe brachiOpodan sca 93°. 55) LOD) «3, BNET 

Hamellipranchiata... 3 -,,« 76 «,, 

Gasteropoda ...... LO eS 08,5 

Pieropoda: 4.5.0. - thin ates hy D7 net 9 25 

Heteropoda ...... Lp pestn whois 

Cephalopoda ...... Tie nA] lt. 


ee 


OGIO’ wiz: 


The important zoological groups in the Caradoc are therefore the 
Hydrozoa, Coelenterata, and Echinodermata—the latter especially 
through the Cystideans, no less than 8 genera and 23 species illus- 
trating thisrare group. The Crustacea number 146 species, of which 
106 are peculiar; 20 species are from the Llandeilo and 16 pass 
upwards to the Lower Llandovery, leaving thus for the Caradoc 
and Bala beds the largest Crustacean fauna known in the Lower 
Palzozoic rocks. Only 4 species of Bryozoa out of the 21 appear to 
pass to the Lower Llandovery; and they all commence in the 
Caradoc: they are Fenestella subantiqua, d’Orb., Ptilodictya dicho- 
toma, Portl., P. coslellata, M‘Coy, and P. lanceolata, Goldf. We 
should expect more species in common in this group, from the fact 
of their being usually a moderately deep-sea family, and less subject 
to vicissitude or change than littoral or sublittoral species. The - 
Brachiopoda of the Caradoc and Bala group number 109 species, 
representing 16 genera; they are more numerous than in other of 
the Paleozoic rocks. 33 of the 109 pass to the Lower Llandovery ; 
and we have seen that the connexion with the Llandeilo below is 
through 26 species, leaving, therefore, 50 as Caradoc proper. As 
regards geographical distribution, North-Welsh forms greatly pre- 
dominate, owing probably to the larger amount of research to which 
the rocks of the four counties of Caernaryon, Denbighshire, Mont- 
gomeryshire, and Merioneth have been subjected. Ireland also has 
yielded 60 species, and Scotland 40. Their more gomrighee distribu- 
tion will be further discussed under that head. 

The littoral or shallow-sea condition of the Caradoc is further 
illustrated by the occurrence of a large number of bivalve Mollusca. 
No less than 76 species are known; and of these only 3 species pass 
to the Lower Llandovery, viz. Pterinea retroflewa, Wahl., Orthonota 
suleata, His., and Mytilus mytilimeris, Cony. ; and 2 of these 3 forms 
pass to the Ludlow and the Upper Llandovery, viz. Pterinca retroflexa 
and Mytilus mytilimeris ; so that 66 species of Lamellibranchata be- 
long to the Caradoc exclusively ; and, strange as it may appear, these 
3 are the only species known im the whole of the Lower Llandovery. 


132 PROCEEDING OF THE GEOLOGICAL SOCIETY. 


The Bivalvia of the Caradoc exceed in number those of any other 
known formation below the Carboniferous Limestone. Careful analysis 
shows that the Arenig rocks contain only 6 species, the Llandeilo 6, 
the Caradoc 76, Lower Llandovery 3, Upper Llandovery 29, Wen- 
lock 45, and the Ludlow 71 species; about 13 species range from 
the Caradoc upwards or through to the Ludlow, 7 to the Upper 
Llandovery, 3 to the Woolhope, 11 to the Wenlock, and 12 to the 
Ludlow—this of course being inclusive, as some of the same forms 
appear more than once in their range. It is not improbable that 
errors as to species occur, especially when we have to do in many 
instances with mere casts in these arenaceous deposits. In both the 
Lower and Upper Llandovery groups the fossils are badly preserved, 
being casts only. If the Lamellibranchs are largely represented in 
the Bala group, the Gasteropoda are almost equally so, by 14 genera 
and 53 species; 2 only (Murchisona simplex, M‘Coy, and Turbo 
tritorquatus, M‘Coy) are common to the Llandeilo below; 10 pass to 
the Lower Llandovery ; 9 direct to the Upper Llandovery ; 7 species 
are common to the Caradoc, Lower Llandovery, and Upper Llan- 
dovery ; they are Cyclonema crebristria, M‘Coy, Holopella cancellata, 
Sow., H. tenuicincta, M‘Coy, Murchisonia cancellata, M‘Coy, MM. 
pulchra, M‘Coy, Trochonema triporcatum, M‘Coy, and Rhaphistoma 
lenticulare, Sow. Only 3 species pass to the Wenlock rocks: one 
of these is Holopella cancellata, Sow. ‘The Gasteropod fauna there- 
fore is as significant as the Lamellibranchs, no fewer than 32 of 
the 53 species being confined to the Caradoc or Bala group. 16 of 
the 53 are peculiarly Irish forms, and 5 Scotch, leaving 21 for dis- 
tribution through the North- and South-Welsh and English beds, of 
which 9 occur in the Caradoc of Shropshire. I have no determined 
species from Westmoreland ; North Wales yields 26 species, South 
Wales only 3, viz. Cyclonema crebristria, Holopella cancellata, and 
Patella saturni, and these only in Caermarthenshire. This group, 
like the bivalves, indicates shallow-water conditions. 

The Pteropoda (4 genera and 12 species) are, with 4 exceptions, 
confined to the Caradoc. Conularia Sowerbyt passes to the Lower 
Llandovery, Wenlock, and Ludlow ; ELeculeomphalus scoticus, M‘Coy, 
is Llandeilo and Upper Llandovery ; 9 of the known species are 
Irish, and 3 of them Scotch; so that the Lower Llandovery is only 
directly allied to the Caradoc by one form (C. Sowerby) out of 
the 12 occurring. This is even more strongly manifested by the 
associated’ class Heteropoda, of which, as in the Pteropoda, only 1 
species in 15 passes to the Lower Llandovery—Bellerophon carinatus, 
Sow., being, so far as I know, the only form in this group connecting 
the two formations. Bellerophon bilobatus, Sow., B. perturbatus, Sow., 
and Maclurea macromphala, M‘Coy, are also Llandeilo. The Upper 
Llandovery has 3 in common, Woolhope 4, Wenlock 2; and 2 
pass to the Ludlow. Specifically, 6 pass up, leaving 8 as be- 
longing to the horizon. The Pteropoda, being strictly or essen- 
tially pelagic, give us little clue as to bathymetric conditions at 
the time of deposition. Neither do the Cephalopoda; but no single 
class is so preeminently Caradoc: of the 47 known species, only 1 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 133 


(Orthoceras arcuoliratum, Hall) unites it with the Llandeilo group 
(and this I believe to be a doubtful species); and only 7 connect it with 
the Lower Llandovery, and 3 of the 7 do not range higher. The 7 
species are Jituites cornu arietis, Sow., Orthoceras ibex, Sow., O. 
vagans, Salt., O. annulatum, Sow., O. Barrandu, Salt., O. politum, 
M‘Coy, and O. tenuistriatum, Minst. The 3 species that directly 
unite the Caradoc and Lower Llandovery are three of the above— 
O. vagans, Salt., O. annulatum, Sow., and O. Barrandii, Salt. 9 
range to the Upper Llandovery, 4 to the Woolhope, 9 to the Wenlock, 
and 3 to the Ludlow. These are the appearances of species that 
ascend through and into the higher formations. 30 of the 47 species 
are confined to the Caradoc, not ranging higher, whereas 15 species 
pass to the rocks above. Ireland is represented by 24 of the 47 
species, 18 of which do not leave the Caradoc horizon, and 17 are 
strictly confined to the Irish deposits. 

Of the Scotch Caradoc Cephalopoda we only know of 6 restricted 
species, viz. Orthoceras audax, Salt., O. arcuoliratum, Hall (also 
Llandeilo), O. vaginatum, Schloth., O. politum, M‘Coy, O. prumevum, 
Forbes, and O. tenwistriatum, Munst.; and only 15 species out of the 
known 47 occur in the Scotch rocks. 


GEOGRAPHICAL DISTRIBUTION. 


I have now to treat of the geographical distribution of the 
extensive series of species occurring in the Caradoc and Bala rocks, 
the large fauna (614 species) being an additional reason for so doing, 
more especially considering the widely spread area occupied by the 
Lower and Middle Bala groups. It would be impossible to give ali 
the localities whence our knowledge of their distribution has been 
_derived; but the chief in North Wales number between 30 and 40, 
and in South Wales about 10. The Shropshire and Westmoreland 
localities are less numerous, but are prolific in species. 

The following 10 areas, including about 50 localities in the British 
Islands, have yielded the 613 species known. Their occurrences or 
appearances number 1555. 


- ( Caernarvonshire......... 105 species, distributed in 10 chief localities. 
North } Denbighshire ............ 131 af ° Ghees - 
Wales. | Montgomeryshire ...... 1388 4 Be is A 3 

Merionethshire ......... 162 Fe i LOweaes $3 

South { Pembrokeshire ......... 93 es Ms Rn iss Pe 

Wales. Neer ccchirs mere 72 fs a pes 6 
end leeresie ara Poe aces os: 123 
8 * | Westmoreland ...... 123 
SUC ENING | Goh SAS COREG REEMA aEEe Bere 302 
[iyailninG! CL EAe seer 306 
1555 


The actual number of species (so far as we know) occurring in 
North Wales is 270, in South Wales 134, in Shropshire 123, in 
Westmoreland 123, in Scotland 302, and in Ireland 306. 

The intimate relation of the Caradoc to the succeeding Lower 
VOL. XXXVII. 


134 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Llandovery rocks obliges me to show their specific affinity, for the 
purpose of ascertaining the paleontological value of the Lower Llan- 
dovery (the Upper Bala of Sedgwick). Analysis shows that, of the 
613 Caradoc species, 103 pass to the Lower Llandovery ; but, strange 
as it may appear, 107 are common to the Caradoc and Upper Llan- 
dovery, although we know of the stratigraphical break that occurs 
between the two formations. This relation is chiefly through the 
Actinozoa, Hydrozoa, and Brachiopoda, as we might expect from their 
bathymetrical position and habitat. 

Of the 40 known corals, 22 (or more than one half) range to the 
Llandovery rocks, 20 occurring in the Lower and 17 in the Upper 
Llandovery; or all but 3 are both Lower and Upper Llandovery ; 
and 37 species of Brachiopoda, out of the 109 known, pass to the 
Llandovery, 33 of which are Lower Llandovery. 

Of the 16 species of Annelida, only 2 pass to the Lower and 
Upper Llandovery, Tentaculites anglicus and Cornulites serpularis ; 
they range also to the Ludlow. 

The class Echinodermata, represented by 32 species, is remark- 
able for the occurrence of 8 genera of Cystidea and no less than 
23 species. They essentially characterize the Caradoc and Bala 
rocks ; with the exception of one species (Hchinospherites arach- 
noidea, Forbes), which passes up into the Lower Llandovery, all 
are peculiar. Itis only in the Caradoc and Wenlock strata that the 
Cystidea are a well-developed and characteristic group of the Kchi- 
nodermata; they are replaced in the Carboniferous rocks by the 
Blastoidea (Pentremates and Codonaster) and do not appear again. 

The Crustacea are the largest and most important class in the 
Caradoc. The species number 146, representing 37 genera—Ireland 
yielding 90 (the largest number of species), Scotland 78, North 
Wales 50, and South Wales 30; or taking North and South Wales 
as one area, it has 59 species, 16 genera and 21 species being 
common to North and South Wales. Shropshire and Westmoreland 
are nearly equal, the former having a known crustacean fauna of 
31 species and the latter 35. The genera most largely represented 
are Calymene 8 species, Phacops 14, Jilenus 13, Remopleurides 8, 
Inchas 6, Stawrocephalus 4, Acidaspis 8, Ampyx 6, Asaphus 6, 
Chewurus 6, and Trinucleus 5. The remaining 26 genera (many 
of which are Phyllopods) are represented by one or few species. 
The 37 genera of Crustacea appear or occur 217 times in the 
geographical areas named in the table of distribution and the 
horizons they pass to above, and the species appear 433 times. 
This Table clearly shows us how much has yet to be done before 
we can obtain reliable zoological data, or be assured that the 
several classes and genera have been even yet fairly ulustrated, 
feeling at the same time that, except by exhaustive and careful col- 
lecting, we never shall arrive at even a fair illustration of the fauna 
of any given group. For example, the 9 genera of Caradoe Pro- 
tozoa are represented by only 10 species, or a fraction more than 1 
species for each genus. 8 of these genera belong to the Spongida ; 
and only among these do any 4 of the genera, illustrated by 1 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 135 


Species each, occur in any one locality, as in Shropshire, Scot- 
land, and Ireland. North and South Wales have yielded only 2 
genera, each with 2 species, in any of the six counties. The 
Actinozoa even afford us stronger evidence still of the importance 
of specific evidence. In this class we know through the fullest 
research that 20 genera and 40 species occur, thus averaging only 
2 species for each genus. Many genera (13) have as yet yielded 
only 1 species; they are Plasmopora, Syringophyllum, Halysites, 
Stylarea, Omphyma, Thecostegites, Alveohites, Aulacophyllum, Calo- 
stylis, Cyathophyllum, Pistulipora, Tredadium, and Prosopora. We 
cannct suppose for one moment that these genera are not repre- 
sented by more species; it is a matter of research only. The 
genus Havosites has 8 species, Heliolites 6, Petraia 6, and Monti- 
culipora 4. The 20 genera make 90 appearances through the 
10 areas, and the species 188; 32 of the generic and 68 of the 
specific appearances occur in the formations above the Caradoc, 
-and show the relation of one group of strata to another, as deter- 
mined through paleontological research. The Lower Llandovery, 
which succeeds or is the natural termination of the Caradoc, con-~ 
tains 9 genera and 20 species of the whole fauna (7°); and the 
Upper Llandovery 6 genera and 17 species of the whole. The 
coral fauna of the Caradoc at once makes its appearance underived, 
the older Llandeilo possessing only 3 species of Actinozoa, Halysites 
catenularvus, Favosites fibrosus,and Monticulipora frondosa; whereas, 
next to the Wenlock, the Caradoc possesses the largest Coclenterate 
fauna of the Lower Palzozcic rocks. Nowhere, either in Europe 
or America, does this class appear with so large a generic grouping. 
20 of the 40 species pass to. the Lower Llandovery ; most of the 
same appear in the overlying and unconformable Upper Llandovery. 
The Crustacea pre-eminently characterize the Caradoc and Bala 
rocks, and constitute the largest group in the whole of the Palo- 
zoic series; 37 genera and 146 species have been collected, de- 
scribed, and registered through the labours of the Geological Survey. 
of the three countries, England, Ireland, and Scotland, as well as 
by the researches of Sedgwick, Salter, M‘Coy, Baily, Sharpe, Prof. 
Hughes, &e. No zoological group is better understood, none more 
important to the student of stratigraphical geology. The Lower, 
Middle, and Upper Bala beds could with difficulty be read or under- 
stood without a minute acquaintance with the Trilobita of this vast 
middle series of Lower Paleozoic deposits. They are to this group 
of rocks what the Ammonitide and Kchinide are to the Mesozoic 
series. The order Trilobita is illustrated by 27 genera, the re- 
maining 10 are mostly Ostracoda; among them is Z'wrrilepas. 


Caernarvonshire has yielded 11 genera and 17 species. 


North Denbighshire i ei ee Bow US. 
Wales. }) Montgomeryshire id Sie ars) 29s Le 
| Merionethshire Fe 1G) a ae DO ee 
South f Pembrokeshire % BG S26 24e oe 
Wales. { Caermarthenshire Mi DSH § 200 os 


12 


I 36 PROCEEDINGS OF THE GHOLOGICAL SOCIETY. 


Rochad Shropshire has yielded 15 genera and 31 species. 
Oni ‘etaecie ie ist B86 
Scotland as ol ie, 78% ee 
Treland ee Silene 90a 


The most important genera, or those of chief stratigraphical value 
and in which the species are numerous, are Acidaspis (8 species), 
Ampyx (6), Agnostus (5), Asaphus (6), Calymene (8), Cheirurus (6), 
Homatlonotus (4), Illenus (13), Lachas 6, Phacops (13), and Remo- 
pleurides (8). The genera essentially characterizing the Caradoc’are 
Harpes, Salteria, Remopleurides, Cyclopyge, Dionide, Trresias, and 
Cyphonscus, most of which are only represented by one or few 
forms. Scotland and Ireland possess the richest assemblage of species ; 
26 genera and 63 species occur in the former, and 23 genera and 
77 species in the latter area. Only 15 of the 123 species of Trilo- 
bita pass to the Lower Llandovery; they are Acidaspis Brighti, 
Calymene Blumenbachu, C. brevicapitata, C. caractacr, C. Allportiana, 
Cheirurus bemucronatus, OC. clavifrons, Encrinurus punctatus, Cy- 
phaspis megalops, Cybele verrucosa, Illenus Rosenbergi, I. Bowmanni, 
I, Thomson, Lichas laxatus, and Phacops Brongnarti. 8 of 
these same also pass up into the Upper Llandovery, 6 to the 
Wenlock, and 5 to the Ludlow. The long-ranged species are 
chiefly those illustrating the largest genera, such as Calymene 
Blumenbachui, C. Allportiana, Cheirurus bumucronatus, Encrinurus 
punctatus, Cyphaspis megalops, and Phacops caudatus, all of which 
species appear in the Ludlow and then cease to exist, the Devo- 
nian rocks having none in common; yet Bronteus and Harpes are 
repeated from the Caradoc in the Middle Devonian, both in Britain 
and on the continent. We must remember, however, that the 
marine Devonian nowhere visibly overlies the Silurian rocks in Great 
Britain, and the Old Red Sandstone contains no true marine form 
anywhere. 

The order Ostracoda, illustrated by Beyrichia, Leperditia, Cythere, 
Primitia, and Entomzs, needs little more than notice here; they 
have received at the hands of Professor Rupert Jones the closest 
scrutiny both zoologically, paleontologically, and stratigraphically ; 
few men have so largely added to our knowledge of the orders 
Ostracoda and Phyllopoda. 

Bracutoropa,—And next to the Crustacea in force and classifica- 
tory value we must place this group of Mollusca or Molluscoida. 
Numerically in the whole of the Cambrian and Silurian rocks the 
Crustacea include the largest number of species, 550 being known ; 
whilst of the Brachiopoda we know 456. Individually no class sur- 
passes the Brachiopoda through all the Palzeozoic and Mesozoic rocks, 
many genera being, however, far more richly represented than others. 
In the Caradoc this is notably the case. The genus Orthis has in 
Britain alone, we know, through the large collections that have been 
made and the elaborate researches of Davidson, no less than 110 
species, and culminates in the Caradoc. In tracing the numerical 
and stratigraphical value of the genus, we find that in the Cambrian 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 137 


group up to the close of the Tremadoc the genus Orthis possesses 
(so far as we know) only 4 species, viz. Or this Hicks, O. Carausi, 
O. lenticularis, and O. Menapie; in the Arenig 10 species, the 3 
last-named forms being in common with the horizon below; in the 
Llandeilo 13 species, 4 of which are Arenig also, viz. Orthis alata, 
O. calligramma, O. remota, and O. striatula. 

In the Caradoc and Bala group the species have increased to no 
less than 41, all having extensive geographical distribution; 7 are 
peculiarly Irish. In the Lower Llandovery we know of 20 species, 
17 of which are Caradoc, the only 3 peculiar Lower Llandovery 
forms being Orthis Bouchardi, O. reversa, and O. mullochensis. Thus 
through this one genus alone we see the close alliance of the so- 
called Lower Llandovery (the Upper Bala of Sedgwick) with the 
Caradoc. The moderate depth at which they lived is clearly indicated 
by the coarse and varied arenaceous nature of the deposits, for there 
is no evidence to show, even by the zoological grouping, deep-sea 
conditions. No Lingule are known in the Lower Llandovery rocks ; 
although 6 species occur in the Caradoc and 5 in the Upper Llan- 
dovery. The fauna generally may be regarded as one accumulating 
under decreasing depth or slow elevation over given areas. 8 of the 
20 Lower Llandovery species pass to the Upper Llandovery or May- 
Hill beds. To show still further the decline in specific as well as 
individual members of the genus Orthis, I may mention that in the 
Upper Llandovery there are 10 species, but not a single form 
belongs truly to that horizon ; Orthis rustica, the only true Upper 
Llandovery species, passes to the Wenlock rocks, so that no form of 
Orthis is special to the Upper Llandovery. The species of Orthis in 
the Wenlock rocks number 16, of which 8 come from the Llan- 
dovery and Caradoc and 4 pass to the Lower Ludlow (Orthis biloba, 
O. crassa, O. elegantula, and O. hybrida), so that Orthis Hdgeliana, 
Salt., O. Lewrsi, Dav., O. Hughesw, Dav., and O. mullochensis, var., 
or variety of O. reversa, are new and restricted forms. None pass 
to the Devonian. I have selected this genus in the Caradoc for com- 
parison with the other Silurian groups on account of its magnitude 
and stratigraphical value. Two other genera, Leptena and Stropho- 
mena, Which first appear in the Llandeilo, also characterize the Cara- 
doc by greatly increased specific development. These are Leptena 
with 4 species and Strophomena with 2, both individually numerous 
in North and South Wales, In the Caradoc Leptena yields 8 species 
and Strophomena 19, and for the first time we meet with Rhyncho- 
nella with 8 species ; but no form is known to occur in South Wales, 
and only 3 have been found in North Wales. Ireland and Scotland 
yield 7 of the 8. The following Table shows the specific and stra- 
tigraphical value of these four important genera from the Cambrian 
to the Upper Silurian :— 


138 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


E 
ie} 
7S 
= 
A : 
7 
1 
a 
5 


. bo] ob 

: 0 5 | 
Z qd = S) S tite 6 : 
Be 28 dvley Wiese lice. Ise dec Snel 
Bes ts |. ae | een en es eo eal 
Oe cue bese) 2 le 
See ewes koe | =) = 
Orthismas ctcuteeelt eee Pe 8) | 14s AY 20s Oe Gs 
Heptenarc...oc-cececen| Sec sine Pieepie Nicene cf agin, ta Atle ore eke | mm oe 
Strophomena.........] ... ihe 2) .19: |) EE a 
Rhynchonella ...... Lh 8.) | BOs eo 


O; 1) L| 8] 7 | 21°) 76 | 42 1 (34 aoe 


Those genera having only one or two species are doubtless of 
equally high stratigraphic significance. Such are Orthisina, Stipho- 
notreta, Porambonites, Triplesia, Merista, and Meristella. 

LAMELLIBRANCHIATA.—The known species of Bivalvia in the 
Caradoc and Bala rocks are more numerous than in any other 
Lower Paleozoic group. ‘Thus the Tremadoe rocks of St. David’s 
yield to us the earliest Bivalve fauna, consisting of 5 species; the 
Arenig has 6, the Llandeilo 6, the Caradoc 76, the Lower Llandovery 
only 3, the Upper Llandovery 29, the Wenlock 45, and the Ludlow 
71. These numbers help us to see, by comparison, the zoological 
value of the class Conchifera in the several strata. 

Ctenodonta, Orthonota, Modiolopsis, Pterinea, Ambonychia, and 
Palearca are the largest genera, or those: haying the most species, 
by comparison with the older and younger‘fortiiations. The Caradoe 
Conchifera excel in number all beneath the Carboniferous. The 
following Table shows this, through the 6 Caradoc genera above 
named :— 


B| B 
. Bil eS 
Sp mo | os 
Z 4 = ) 6 i 8 8 ‘ 
mB | LS a = A S eS a hae = 
S last = ls a ra an ee 3 
Gos ao | q q < ge esecalere hl |) 
ale |e)?|2\s | See eeeecn ee 
eens fora) Ss )e <= 
Ctenodonta ......... a Dei he Tile Eee 6 3 4 
Orthonotay eee loe. MIS | ee 5 1 6 3 | 16 
Modiolopsis ......... 4) 1| 2:) 16 | 029) SSc em mes 
IPterinvesc ee ere wae Albers | aes 6 1 6) TE we 
Ambonychia ......... dae See gee cee SOT eee 
iPaleearca, | eeeete mene shies Tae ee 2 a STAR MS 438 
<=. "| —_ |J "|_| | || |_| | |__| 
4 | 57 2 D1 Dae ares 


Cardiola with 3 species, Mytilus with 4, Pleurorhynchus, Ano- 
dontopsis, Cucullella, Megalomus, Clidophorus, Arca, and Lyroderma 
having each 1 species, complete the Conchifera of the Caradoc. Their 


/ 
ANNIVERSARY ADDRESS OF THE PRESIDENT. 139 


geographical distribution is significant, but clearly shows the im- 
perfection in collecting, and how much has yet to be done to 
demonstrate their relation to the extensive areas over which they 
ara distributed :— 


Caernarvonshire has yielded 6 genera and 16 species. 


Denbighshire a a (RE 
Montgomery shire ,, 8 eS 14. —SC««, 
Merionethshire __,, 6 “ Los 
Pembrokeshire _,, none none 
Bo thenskite s 2 i. 2 We 
Shropshire ut 5 if TOe ees 
Westmoreland 7 3 i Ouhaktss 
Scotland a ih R Bye 
Ireland ‘5 10 55 Side ee 


Nothing short of strict analysis could impress upon us the fact 
of such unequal distribution as the above examination shows. No 
species of bivalve is known in Pembrokeshire, a district rich in 
other groups. Only 1 genus and species in Scotland (Pleurorhyn- 
chus dupterus, Salt.), 2 only in Caermarthenshire (Ambonychia triton 
and Ctenodonta varicosa, Salt.), and 3 in Westmoreland (Ctenodonta 
anglica, D’Orb., Pterinea tenwstriata, M‘Coy, and Cardiola inter- 
rupta, Sow.). The physical geography of the area, either through 
barren or interrupted coast-line, rather than movements of land or 
depth of sea, would most probably account for this unequal distri- 
bution and poverty of species in one area, and their comparative 
abundance or fair representation in another. Looking at the horizons 
which the above 6 genera illustrate, and through which they pass, 
it seems hardly explicable that there should be only 3 species of 
Lamellibranchiata known in the Lower Llandovery, viz. Pterincea 
retroflexa, Mytilus mytilimeris, and Orthonota sulcata ; and these are 
Caradoc and Upper Llandovery also, showing us that not a single 
species belongs or is confined to the Lower Llandovery. This 
result, compared with the large bivalve fauna in the Caradoc or 
Lower Bala beds (57 species), and under conformable stratification, 
is scarcely to be accounted for, except through elevation of the sea- 
bed and change in bathymetrical conditions sufficient, with all 
other conditions, to cause the extinction or removal to another area 
of the Caradoc Lamellibranchs. Similar results are obtained with 
the Gasteropoda; the 53 Caradoc and Bala forms dwindle down to 
16 in the Lower Llandovery. The Llandovery species will be noticed 
in their place or order subsequently. As before, Ireland yields the 
largest number of species (37). . 

GastERopopa.—Like the three preceding classes, the group 
Gasteropoda is largely represented in the Caradoc strata ; 14 genera 
and 53 species have been determined. No species has yet occurred 
in Pembrokeshire, and only 3 in Caermarthenshire (Holopella can- 
cellata, Sow., Patella saturni, Goldf., and Cyclonema crebristria, 
Moy). Neither have we evidence of any species from Westmore- 


I40 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


land, although, considering the somewhat abundant forms in Shrop- 
shire (9) and Scotland (11), and that the Westmoreland area inter- 
venes, we might expect some to occur. The genus Huomphalus in 
Ireland (with 7 species), Holopewa in Merioneth (7), Murchisonia and 
Rhaphistoma in North Wales generally (12), constitute nearly one 
half the univalve fauna—T'rochonema, Turbo ?, Trochus?, and Pleu- 
rotomaria having but few species in each. 8 genera and 19 species 
actually pass up to the succeeding Llandovery, Wenlock, and Ludlow 
rocks above. The Lower Llandovery receives 10 species, and the 
Upper 16; the Woolhope 1 (Turbo tritorquatus, M‘Coy), the Wen- 
lock 2 (EHuomphalus alatus, His., and . sculptus, Sow.). 3 species 
range into the Ludlow—Huomphalus alatus, Holopella cancellata, and 
H. conica. ‘Three species of Gasteropoda are common to the Llan- 
deilo below—Cyclonema crebristria, Murchisoma simplex, and Turbo 
tritorquatus. The relationship of the two Llandoveries is marked ; 
the Lower has 103 species of all classes in common and the Upper 
107, Ireland and Scotland possessing the highest numbers, being re- 
spectively 304 and 206. 

PrreropopA.—Ten species of this class occur, the largest pelagic 
Pteropod fauna known in the Paleozoic rocks ; they occur in nearly 
equal numbers through the 10 areas, Ireland still yielding the 
highest number, nearly three times as many as Shropshire and 
Merioneth. No species is known in Pembrokeshire. 

Hetreropopa.—This order is represented in the Caradoc and Bala 
rocks by 11 species of Bellerophon, 2 of Kcculiomphalus, and 3 species 
of Maclurea. In the British area the two last-named genera are 
restricted in their geographical range to Ireland and Scotland, and 
stratigraphically to the Arenig or Llandeilo and Caradoc. Maclurea 
is known only in Scotland; MM. Logani, Salt., M. Peach, Salt., 
M. macromphala, M‘Coy, and M. magna are said to be (in Scotland) 
of Llandeilo age. The Durness species, M. Logam and M. Peachia, 
with their associated forms, I believe to be Arenig. Piloceras in- 
vaginatum and Orthoceras mendax occur in the same beds. Maclurea 
abounds in the Chazy Limestone of the United States and Canada 
—hbeds probably to be correlated with our Lowest Llandeilo, the 
Upper Cambrian or Lower Bala group of Sedgwick. J. M‘Coyz, 
M. macromphala, and M. magna are admitted to be with us of 
Caradoc age, and certainly occur low down in the series. Clonmel, 
Stinchar river and Bugon &c. in Ayrshire, and the Durness Lime- 
stones yleld M. Peachu and M. Logan: abundantly. 

I have preferred to place Kcculiomphalus with the Heteropoda 
rather than the Pteropoda, although Portlock’s genus may be one 
of the latter group. 

CrpHatoropA.—No less than 47 species have been determined 
from the Caradoc series ; their maximum development is in Ireland, 
where 6 genera and 24 species occur; no species is known in Shrop- 
shire. Westmoreland and Scotland register 12 each. The Irish 
genera are Cyrtoceras, Koleoceras, Lituites, Poterioceras, Trocholites, 
and Orthoceras. As a generic fauna this is by far the largest in 


ANNIVERSARY ADDRESS OF THE PRESIDENT. I4I 


the Paleozoic rocks. Only one species (Orthoceras arcuoliratum, 
Hall) is common to the Caradoc and Llandeilo; but 15 species 
range upwards—7 to the Lower Llandovery, 9 to the Upper Llan- 
dovery, 4 to the Woolhope, 9 to the Wenlock, and 3 to the Ludlow ; 
or the 15 make their appearance or occur 32 times in their range 
through the higher divisions of the Silurian rocks ; 32 species are 
therefore essentially Caradoc or Bala forms. The numerical distri- 
bution of the Caradoc species through North and South Wales is re- 
markably uniform. 12 species occur in the four North-Welsh coun- 
ties, and 8 in the two South-Wales counties. Shropshire possesses 
no Cephalopod, yet 57 genera and 1283 species range through 15 
zoological classes occurring in that county; but even this number 
represents numerically only a little over 2 species for each genus. 
Table X. is prepared on geographical data, the universality 
or extension of the Caradoc rocks and species not admitting of 
any other mode of expressing their distribution; and to analyze 
the group under subdivisions of the Caradoc or Bala, as given by 
various authors, would end in confusion, owing to the application 
of the terms Lower, Middle, and Upper Caradoc, or Bala, not being 
the same inall. Text-books have not yet defined the limits of these 
divisions, either zoologically or geographically ; in the former sense 
the greatest difficulty would be felt over extended areas, through 
want of true succession and continuity in strike. A careful study 
of the Woodwardian Catalogue demonstrates the difficulty of treating 
the Caradoc or Bala group as a whole in any other way. The 
column headed ‘ Localities and Numbers” in that Catalogue at once 
shows that it is only through space distribution that we can realize 
the value of the subdivision. In the Lower Bala group of the Upper 
Cambrian, as established by Sedgwick, Mr. Salter has placed 45 
genera and 82 species; in the Middle Bala 82 genera and 212 
species; and in the Upper Bala group 43 genera and 100 species. 
That the fossils of the “‘ Lower Bala” of the Woodwardian Cata- 
logue represent the Llandeilo proper of the Geological Survey there 
can be no doubt ; and out of the 82 species catalogued, only 17 pass 
to the Middle Bala group, and 8 to the Upper Bala. The Middle 
Bala of the Cambridge Catalogue receives the fine assemblage of 
fossils collected from the Bala limestone, sandstones, and slates of 
North Wales, the Caradoc sandstones and the Horderly limestones 
of Shropshire, or the true Caradoc. It includes also the fossils of 
the Coniston Limestone, the Kildare, Craig-Head, and Peebles Lime- 
stones. The Hirnant Limestone is not included in Sedgwick’s 
Middle Bala, but constitutes the base of his Upper Bala group. The 
212 species largely illustrate the characteristic fauna of the Caradoc, 
and the group clearly shows the value of the division (especially 
with the elimination of the Llandeilo forms) and its separation 
from the Lower Llandovery (the Upper Bala of Sedgwick), into 
which only 33 of the 212 species passed. These proportions in the 
Cambridge collection (as catalogued), showing the community of 
forms between the 3 divisions, show how well the collection was 


I42 -PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


made, and also the care bestowed upon the stratigraphical grouping 
of the fossils. The Upper Bala group of Sedgwick includes the 
Aber-Hirnant Limestone, which occurs above the Bala Limestone, 
also the lower part of the Coniston Flags, the beds above the Bala 
Limestone, and below the May-Hill Sandstone; it therefore includes 
the Lower Llandovery or Mathyraful Limestone near Meifod, and 
also the rich fossiliferous series at Haverfordwest, the Dalquorhan 
and Mulloch beds of Scotland, and the Maume and Cong beds of 
Galway *. The Woodwardian Catalogue enumerates 43 genera and 
100 species from this group (the Lower Llandovery of the Geolo- 
gical Survey), 18 genera and 36 species of which also occur in the 
Middle Bala below. 


Lower LLANDOVERY. 


In my analysis of the Lower Llandovery species from the rocks 
of that age in North and South Wales, Scotland, and Ireland, I have 
brought together and tabulated all that is known relative to the 
range and distribution of life through this Middle Silurian or Upper 
Bala group of Sedgwick. ‘The table shows the value of this upper 
member of the Caradoc or Bala beds, and its relation to the transi- 
tional Upper Llandovery, or the group so called, which stands be- 
tween and connects the Lower Silurian of the Survey with the well- 
defined succeeding Wenlock and Ludlow series. Careful analysis of 
the species in both their stratigraphical and geographical distri- 
bution shows how small a specifically characterized group the Lower 
Llandovery appears to be, for only 115 species out of 204 are really 
Lower Llandovery. The total number of genera and species known 
for all Britain is 68 genera and 204 species; but 50 genera and 
104 of these species transgressed or came from the Caradoc and Bala 
beds below, thus reducing the actual Llandovery fauna to 18 genera 
and 100 species. The intimate connexion with the Upper Llan- 
dovery is chiefly through the Actinozoa (20 species), Brachiopoda 
(38), Crustacea (10), and Gasteropoda (8). No form of Protozoa, 
Echinodermata, or Pteropoda is common to the two Llandoyeries 
in any area in Britain. 

No species of Echinodermata occurs in North Wales, Ireland, or 
Westmoreland. No Lower Llandovery Annelid is known in North 
Wales. Neither do we know of any Lamellibranch, Pteropod, 
Heteropod, or Cephalopod in the same area. I will now discuss 
through tabular analysis the distribution of the chief zoological 
groups in the Lower Llandovery, viz. the Echinodermata, the Crus- 
tacea, Brachiopoda, and Gasteropoda. The remaining 7 of the 13 
groups (there being no plants) are conspicuous through feeble re- 
presentation, due doubtless to geographical changes towards the 
close of the Caradoc and Bala period. 

Hyprozoa.—The careful researches and generalizations of Lin- 
narsson in Sweden, Lapworth, Nicholson, Carruthers, and Hopkin- 


* Vide Cat. Coll. Cambrian and Silurian Fossils in Mus. Univ. of Cambridge, 
pp. 25, 26, 39, & 72, for greater details. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 


143 


A 2 TE Sm aD ee 


=o ee AD 4H won on a 52 mn wo =e | From Llandeilo. 

Q bg Q kg 

wae ese Soe ee a 

[Sect |) 2 Bee Sy See Ess eh ey 3 

= Oa a OO AR Eee os 

wets 2 Se S Sie eo 6 § 

Seo OE o Ss 2 eS SN io 
= si «63S fen en erie a of Ee Go £ 
: Bee 6S. oO: & Se os fo} 
: a © - Seale coe J het el laden = ar baa aie ie SS; 
: g9 3) OLAS of aCe ge ae = 

. wm 

2 6 By Qe NARI = algae ail iees eal la R a a 
: | 2 SGT SR <p hn dae as enc @ 
. . . . . oc . . . . . . . . . 
: es ops ere ee ea eS 
— 
~I — et co — — b> bO 
Cc aw worroroaoantwnare oo Oo Ke 


| 
| 
| 
i | Genera. 
Ip 
| 
| 
| 
: 


~ . 
OF | wp oe ee < ao be we Ue be AR BAR AA HE Caernarvonshire. 
uo 
: iA 
‘ ; ° 
G2 [ow an ue wa sa So oe EY we ca Zo : =~ | Denbighshire. a 
5 
eg [ee an ae oe Ee So Ta Sy we xe Gx : sw | Montgomeryshire. = 
| ss Q 
ne =i COS Lec) ARC cole ee : Meri thshir S 
aS ai) tm 4P oF LG B& Go AA GX OR > erlonetnsnire. ae 
Lar} 
OVE & 
TM 
Sl oe Sr : 5 Ss qo BE AS Fe Bos bop Pembr k hir g E. 
EN : 5 a RO wo ~~ : roKkesoaire. = Q 
S 
| RO rR oe we nm Bo re TO AA ww on wes | Caermarthenshire.| = ae 
! | fh Fy 
; S 
=) ; = 
we > oH on oa Da oe we CF AA A® SA AA abo Shropshire. = . 
| F | | 8 
Se eo ee ee Oe exes, Sm nono) com he W t r i d B 
os = pon VS ee ae TS) estmoreland. or 
| | 
TS an _ ) Ae a9 : De wr 
Pegiemo se oF SS ee OH ge OF 2 ae wo aa | Scotland. 
| | 
se | 
oe 2 Of won iS as os aw =i oh 82 oa Pa EN | Treland. 
| | 
= 
og | We He He So we Bo aw 55 ww ww So x» ee | Lower Llandovery. 
| | 
<A 
os | or au we Zo an Yo ow TT ww < : | tipper Manidevary. 
| | 
Fg 
. = 
ae | Aw aH fue we iS Ny CQ we an ae Woolhope 2 
| | S 
an | ow pe wr we To a? bn ©@ Aw fico ne Wenlock. 
| | 
nS wh Ne Me top ee TO me @r po ah ee Ludlow. 


“0PD.WI—'X ATAV I, 


144 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


son in Britain, Hall in America, and Barrande in Bohemia, upon 
the history, morphology, and distribution of the Rhabdophora 
through the Lower Paleozoic rocks of Britain, Europe, and North 
America, have greatly added to our knowledge of their stratigraphical 
value and distribution both in time and space. To no author, how- 
ever, are we so much indebted as to Mr. Lapworth for his exhaustive 
researches in this department of Paleontology. Until his careful 
analysis and descriptions of the Rhabdophora, as well as the physical 
relations, of the Moffat series in Scotland, and his able papers in the 
Ann. & Mag. Nat. Hist. ser. 5, vols. i11.-vi., we had little clear know- 
ledge either of their zoological grouping or stratigraphical distribution 
and relations to other geographical areas, either in Britain, Europe, 
or America. That the southern Highlands or uplands of South Scot- 
land yield rocks of a peculiar character is well known. They consist 
of two types—coarse greywacke, grey, green, or purple in colour, 
associated with fissile flagstones, which either alternate with the grey- 
wacke, or are arranged separately in zones of considerable thickness. 
These two types preponderate. ‘On the west coast, near Girvan, 
limestones, shelly sandstenes, and mudstones yield fossils in great 
profusion, and both mineralogically and paleeontologically remind us 
of the most prolific areas of Wales and Shropshire.” 

Certain beds of black carbonaceous shales and mudstones occupy- 
ing “long lenticular areas occur in the great mass of barren grey- 
wacke; they form extended moniliform lines of great extent, and 
range at intervals throughout the northern half of the Southern 
Uplands, from the Irish Channel to the North Sea; these black 
shales everywhere swarm with Rhabdophora.” It is this group of 
Graptolitiferous strata of the south of Scotland (the Moffat series), 
and their physical and zoological relations to deposits in other areas, 
that Mr. Lapworth has made classical. We know of few or no Lower 
or Upper Llandovery Hydrozoa in Wales. The Coniston Mudstones 
(Westmoreland) yield 6 genera and 25 species; and these probably 
belong to the Lower Llandovery series. The value, however, of the 
Scotch deposits, and their Graptolitic fauna of 50 or more species, is 
very great; they clearly show us that the paucity of species in 
the Welsh rocks is probably due to geographical and other physical 
changes. 

South of Westmoreland this group had scarcely a representative 
after the deposition of the Caradoc, none having survived the close 
of the Upper Bala period ; for the Lower Llandovery in North Wales 
has yielded few or no species; the Upper Llandovery and the suc- 
ceeding Wenlock group, through the Tarannon beds, 5 genera and 
23 species ; North Wales 4 genera and 8 species, and South Wales 
8 genera and 11 species; Westmoreland 3 genera and 12 species; 
Ireland none. These several numbers are as near as I can bring 
them, knowing the difficulty of understanding the Llandovery ques- 
tion, and omitting in North Wales the sandy or arenaceous Llan- 
dovery, which (in that area) does not appear to have furnished a 
congenial habitat for the growth and development of the Rhabdo- 
phora. Highly favourable, however, the conditions seem to have 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 145 


been during the formation of the Arenig, Llandeilo, Caradoc, and 
Lower Llandovery deposits, as shown in the following table :-— 


eREMEOM ee Pee, ee Sse ee 17 genera and 24 species. 
MbmmeNOme rie oes Sl 18 e3 44, 
JADLLIG | oe ole eee 21 a Srey be 
Lower Llandovery of South Wales 5 es ly Guar 
Lower Llandovery of Westmoreland 6 cs 20 ee 
Lower Llandovery of Scotland... 7 3 Sy bie 
Mippersilandoyery .2...-.....- + HE d iage 
1 ESET G] Rie ap ge a Re 9 " Zee 
_ LUD ab ei eer 3 Be oie Pe 


I mention these facts and results here rather than under the 
Ludlow section, as the group of the Graptolitide are conspicuous 
here, and they are of little value in classification above the Llando- 
very rocks. It may be remembered that in 1872* Prof. H. A. 
Nicholson published, in the Journal of the Geological Society, a 
paper of great originality and highly suggestive of the mode of mi- 
gration and distribution of the Graptolites in time and space, or 
the vertical and lateral range of this group of fossils. In this paper 
Prof. Nicholson endeayours to show the peopling of one area as 
derived from another and from points widely separated, showing 
that under such circumstances they can never be truly contempo- 
raneous. ‘In the present state of our knowledge, however, it must 
be more or less provisional and tentative” to attempt to trace the 
migrations of any given set of species or groups of fossils; but pro- 
bably the time will arrive when definite groups under given conditions 
may receive elucidation for lateral range, especially when the original 
area of occurrence and dispersion is known. Dr. Nicholson argues, 
and justly, that “‘ if we were thoroughly acquainted with the range 
of any given fossil or species vertically, and were conversant with 
all the details of its geographical distribution, we should then be 
able to lay down with some degree of accuracy the lines along which 
it must have migrated when the condition of its original area be- 
came unsuitable for its further existence therein.” Prof. Nicholson 
endeavours to show in his paper certain facts relating to the distri- 
bution of the Graptolites, so as to enable us to sketch out the mi- 
erations of these and other organisms from their first appearance in 
time ; in the case of the Rhabdophora from the Arenig to their final 
disappearance at the close of the Ludlow deposits. Five areas are 
selected to illustrate the distribution of the Graptolites:—I1st. The 
Skiddaw group and its Arenig fauna, as being the earliest in which 
the Graptolites appeared, and probably the first area, for we have no 
reason to believe that the Canadian or Quebec deposits much preceded 
in time our own Cumberland series; the large number of genera 
and species not only common to the Skiddaw Slates of the north of 
‘England and the Quebec group, but exclusively confined to them, 
conclusively and distinctly prove that the area was no small one. 


* Quart. Journ. Geol. Soc. vol. xxviii. pp. 217-2382 (1872). “ Migrations of 
Graptolites,” by H. A. Nicholson, M.D., D.Sc., &e. 


146 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


The differences, indeed, between the Graptolitic faunas of the Skid- 
daw and Quebec regions are not greater than what may be due to 
imperfect observation, or the effect of difference of station, without 
either supposing that one region was peopled by migration from 
the other*. 

The second area discussed by Dr. Nicholson is that of Wales and 
Scotland, containing the Llandeilo Graptolites. The auther believes 
that the north of England was dry land during the early portion of 
the Llandeilo period, and also that the close of the Skiddaw 
(Arenig) period was signalized by the upheaval of the Lake district ; 
this, or the unfitness for life of the succeeding seas, in conse- 
quence of igneous activity, may have been the cause of no deposits 
containing Graptolites being superimposed upon the Skiddaw Slates, 
until we reach the higher or later portion of the Caradoc or Bala 
period (the Lower Llandovery), at which time an immigration of 
Graptolites must have taken place from neighbouring seas. On such 
hypotheses, supported by details too minute to discuss here, we may 
realize the absence in certain areas of zoological groups whose 
existence necessitates the deposition of sedimentary matter con- 
genial to their growth and development. Why the Lower and 
Upper Llandovery rocks of North and South Wales do not possess a 
larger Graptolitic fauna, when in the north of England and Scotland 
such a fauna is well developed, can only be explained through 
physical changes, either movements of land and sea, or interference 
through sedimentary agency, of which there may be many kinds. 

Dr. Nicholson further enters into the consideration of the second 
great Graptolitic period of the north of England, during which the 
mudstones of the Coniston series were deposited. In the Coniston 
Limestone proper, which corresponds to the Bala Limestone, no 
species have occurred, it being wholly barren of Hydrozoa; but the 
so-called Graptolitic mudstones succeeding it abound in species, 6 
genera and 25 species having been recorded from them. Many of 
these (16) Nicholson believes have been derived from the Upper 
Llandeilo of the south of Scotland, the remaining 9 being peculiar 
to the Coniston Mudstones. The fourth area Dr. Nicholson calls the 
Gala area of the south of Scotland, in Dumfriesshire ; Mr. Lapworth 
applied to these deposits the name “Gala Group.” Hight species in 
this Gala series are derived from the Upper Llandeilo of the Scotch 
area, and the remaining 5 (of the 13) are importations from the 
Coniston Mudstones and Coniston area of the Lake district. 

The Hudson-River shales and Utica Slates Nicholson believes to 
have been peopled by a great migration of the Upper Llandeilo Grap- 
tolites of the south of Scotland, which appear to have taken a 
westerly course, and ultimately to have reached the United States, 
there forming the Graptolitic fauna occurring in the Caradoc or 
Hudson-River shales and Utica Slates described by James Hall. | 
This view of Nicholson’s, that the Graptolites of the Hudson-River 
group are derived through migration from the Upper Llandeilo of the 
south of Scotland, he supports or illustrates by means of a table of 

* Nicholson, Quart. Journ. Geol. Soe. vol. xxviii. pp. 218, 219. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 147 


24 species of Graptolites, 13 of which are so derived. These are 
associated with 1 from the Skiddaw and 8 peculiar to the Hudson- 
River deposits. Nicholson, in his fifth, or Saxon and Bohemian area, 
endeavours to show that two great migrations of Graptolites took 
place from the area of the south of Scotland at the close of the Upper 
Llandeilo period, one westward through Ireland to America, the other 
southward into the north of England; and he suggests a third migra- 
tion from the same area in a south-easterly direction, or into the Silu- 
rian seas of Saxony and Bohemia. Geinitz, in his “Grauwackenfor- 
mation” of Saxony, believed the Graptolitiferous rocks to be the 
summit of the Lower Silurian series (Llandovery). Of 5 genera and 
12 species given, 4 genera and 10 species are derived from the Upper 
Llandeilo of the south of Scotland, and 2 from the Coniston Mud- 
stones. Barrande has recognized the British derivation of the Bohe- 
mian Graptolites (‘ Défense des Colonies, 1870); he recognizes two 
chief Graptolitic zones in Bohemia, viz. Etage D at the summit of 
the Lower Silurian series, and Htage E at the base of the Upper. 
The Bohemian Graptolites derived from Britain number 5 genera 
and 14 species, and are believed to be derived through migration from 
the Coniston Mudstones or Upper Llandovery beds of the north of 
England. These are identical species with those which lived in the 
mudstones named*. I have thus shown that it is only in Scotland 
and Cumberland that we obtain a Graptolitic fauna in the Llando- 
very rocks (probably Lower), and that none occur in either of the 
two Welsh areas‘. 

Acrinozoa.—The species of this class in the Lower Llandovery 
number a fraction more than 2 to each genus, of which there are 12 
(and 26 species). The genera Favosites, Heliolites, and Petrara abound 
most in species—Fuvosites 5, Heliolites 4, Petraia6. The distribution 
for North Wales is 5 genera and 8 species; for South Wales 8 genera 
and 16 species; and for Ireland the same number. Singularly just 
as many species pass to the Upper Llandovery (20) as came up from 
the Caradoc and Bala beds (20), and in each case 9 of the same 
genera; so that in reality the Lower Llandovery rocks possess only 
6 species of Actinozoa peculiar to them, so closely allied are the two 
formations through this, as through other groups of fossils. 7 of the 
9 genera have as yet only yielded 1 species each. Nothing can be 
more unsatisfactory than the evidence afforded by such scanty mate- 
rials; close research would result in yielding numerous and more 
definite species. 

EcuinopERmMATA.—No species is known either in North Wales or 
Ireland, and two species only in South Wales—Schinospherites 
arachnoidea and Glyptocrinus, sp. (stem-ossicles). This last genus 
occurs in Scotland also; neither pass to higher formations, but the 
same two species occur in the Caradoc. 


* Nicholson, loc. cit. pp. 228-230. 

+ Recent researches by Lapworth into the specific value and history of the 
Rhabdophora have tended to reduce the number of species, great numbers of so- 
called species being established on mere fragments of other species. ‘Their 
elimination will be of great value to the student of this difficult class of the 
animal kingdom. 


148 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Crustacea.—The Crustacea of the Lower Llandovery reveal to us 
numerically only 2 species in each genus. We have much yet to 
learn of the history and zoological relations of the Trilobita of the 
Llandovery rocks. The 13 genera and 25 species of this order 
mostly characterize the rocks of South Wales and Scotland ; 2 genera 
and 4 species only are known in North Wales, Calymene (3) and 
Trinucleus (1); and 7 genera and 8 species in South Wales ; Scotland 
has 9 genera and 16 species. Eighteen of the Trilobita are also Cara- 
doc ; and 8 genera and 10 species pass to the Upper Llandovery ; so 
that in the Lower Llandovery one of its largest groups has little 
or no stratigraphical value in classification. 

Bracutopopa.—This class, like the Crustacea, has little or no classifi- 
catory value ; for of the known 10 genera and 59 species in the Lower 
Llandovery, 8 genera and 35 species are also Caradoc and Bala, and 
10 genera and 38 species pass to the Upper Llandovery. The charac- 
teristic species are Meristella subundata, Orthis Bouchardi, Rhyncho- 
nella tripartita, Spirifera exporrecta, Strophomena arenacea, Strick- 
landima lens, Pentamerus oblongus, and P. undatus. The first ap- 
pearance of Pentamerus and Stricklandima is a marked and im- 
portant feature in the Lower Llandovery beds. Three species of 
Pentamerus (P. oblongus, P. globosus, and P. undatus) are Lower and 
Upper Llandovery forms; the 2 species of Stracklandima, S. lens 
and S. lirata, are common to both horizons. In North Wales, 8 
genera and 22 species occur; and in South Wales 10 genera and 48 
species ; Scotland has yielded 8 genera and 29 species, and Ireland 
9 genera and 27 species. The South-Wales beds are enriched 
chiefly through the genus Orthis with 20 species, Strophomena with 
11, and Rhynchonella with 7. 

LaMELLIBRANCHIATA.—No species of this extensive group of Mol- 
lusca occurs either in North Wales or in Scotland; and we are only 
acquainted with 3 genera and 3 species in the Lower Llandovery, 
viz. Pterinea retrofleca, Mytilus mytilimeris, and Orthonota suleata ; 
these 3 are South-Wales forms, and the Pterinea occurs in the Irish 
Llandovery. This meagre representation of so large a class in these 
beds, as compared with the Caradoc with 16 genera and 76 species, 
clearly shows that at the close of the Caradoc and Bala period great 
changes took place with regard to the relations of land andsea. The 
Caradoc sea evidently shallowed and reduced its area, and this was 
accompanied by corresponding sedimentary changes which governed 
the life, habits, and distribution of the succeeding Llandovery fauna. 
This is borne out by the Brachiopoda and other groups which 
reached such perfection in the Caradoc seas (109 species), Crustacea 
(146 species), the Lamellibranchs (76 species), Gasteropoda (53 
species), and Cephalopoda (47 species); whereas in the Lower Llan- 
dovery many of these groups are scarcely represented ; for example, 
the group we are now treating of has only 3 species, the Kchino- 
dermata 2, theGasteropoda 16, and the Cephalopoda only 3, with 
only 1 species of Pteropod and 1 Heteropod. 

Doubtless the strictly arenaceous nature of the deposits and the 
somewhat shallow sea constituted the main cause of this extremely 


ANNIVERSARY ADDRESS OF THE PRESIDENT. I49 


small bivalve fauna. Two of the three species of Conchifera are 
Caradoc, and the same two passed to the Upper Llandovery, so that 
in reality only | species is to be regarded as Lower Llandovery. 

GastERopopaA.—Al]though this class includes a much greater number 
of species than the Lamellibranchiata, yet the value of the fauna for 
classificatory purposes is equally unsatisfactory. 9 genera and 13 
species are known; but of these 6 genera and 10 species are Caradoc 
and Bala forms, and 6 genera and 8 species are common to the two 
Llandovery series. Only 1 species is known in North Wales (Mur- 
chisonia pulchra, M‘Coy), 5 genera and 9 species in South Wales, 
and 6 genera with 6 species in Scotland and with 7 in Ireland. Thus 
only 6 species of Gasteropoda are truly of Lower Llandovery age— 
Murchisona Prycew, M. sulcata, Platyschisma Williamsoni, Acroculia 
haliotis, Trochus Moore, and 7’. multitorquatus. 

Prrropopa.—The only Pteropod occurring in the Lower Llando- 
very is Conularia Sowerby, Det., and this has only appeared in South 
Wales and Scotland; none are known in North Wales or Ireland. 
This species bridges over the Upper Llandovery, Denbighshire Grits, 
and Woolhope series, and reappears in the Wenlock Shales and 
Limestones in South Wales and Scotland. 

Hzrrrorops.— Bellerophon carinatus, B. bilobatus, and B. dilata- 
tus? are the chief Lower Llandovery species. The same 3 are Caradoc, 
and B. carinatus and B. bilobatus also occur in the Upper Llando- 
very of South Wales and Scotland, B. dilatatus being Irish also. 

The species of these two nocturnal and pelagic groups, the Hetero- 
poda and Pteropoda, from their habit, had a wide geographical dis- 
tribution, and consequently a long range in time also. 

CrepHatoropa.— We have seen that the Lower Caradoc and Bala 
rocks have yielded no less than 8 genera and 47 species ; the Lower 
Llandovery, the highest member of the Bala formation, has, on the 
contrary, only 3 genera and 8 species distributed through the 6 areas 
in which this formation is represented. The genera are Lituites, 
Tretoceras, and Orthoceras. ‘The first-named genus has 2 species, 
L. cornu-arietis and L. undosus ; the second (Tretoceras) 1 species, 
T. bisiphonatum ; and Orthoceras contains 5 species, O. annulatum, 
O. ibex, O. politum, O. tenwistriatum, and O. vagans(?). Of these no 
species occurs in North Wales. South Wales, however, in the Llan- 
dovery area, has yielded 5 species, Scotland 2 (Orthoceras politum 
and O. tenuistriatum), and Ireland 1 species (Orthoceras ibex). Four 
of the 8 pass to the Upper Llandovery, and 7 came from the Caradoc. 
Tretoceras bisiphonatum and Lituites undosus are the only restricted 
Lower Llandovery species out of the 8 known. 

The accompanying table shows the complete census of the Lower 
Llandovery rocks, and their relationship to the Caradoc below 
and the Upper Llandovery above, and at once reveals the fact that 
this upper member of the Bala group has no claim to separation 
from it; for 50 genera and 105 species unite it with the Caradoc 
and Bala proper, and 45 genera and 104 species pass to the 
Upper Llandovery ; and we shall find, on casting the life-history of 
the Upper Llandovery also, that nearly one half of both genera 

VOL, XXXVI. Mm 


150 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


and species are also so-called Llandovery; or out of 91 genera and 
240 species known in the Upper Llandovery, the above 45 genera 
and 104 species occur in the lower group also. Only 16 genera and 
101 species are therefore really Lower Llandovery. 


Taste X1.—Lower Llandovery. 


Geographical Distribution. 


S mm eae 
S) wa wR ra Oo > 
o F i 
S Classes. = 4 Sf ao 
o Z| 2) S 
eS e : = = oc : 6 jts 
2. s g a fe = = | Oe 
g 2 3 = 5 a & |) aad 
= © 8, S 5 8 2 Sai 
Fy to) a A D D 5 Ee ia 
Plantae ses seteceiee 
Fil) LEIROLAOVAOE YS “AU onae mace pane 3 A 3 a 
1 5 7 7 6 
Ep ||| LelylxoyZory eessabeddser 7 50 A we te & 3 
5 ¢ 5 8 7 8 
BP || EGBUIVOYACE opodaaoaves 12 26 an A ao oh 
2 | Echinodermata...... 2 2 3 E 
2 1 9 2 1 1 
2) Atameladays tereenee tees 2 3 Be: z iu u 2 
10 1D im 4 8 10 7 
+ Ore@rustaceaiwscsencees 13 25 P ae 1¢ Z o 
AaB RY OZOBiasceemelorec oe 2 5 a 2 z 4 2 
8 7 8 10 
3 | Brachiopoda ......... 10 59 Bo ae ne a 10 
3 | Lamellibranchiata..) 3 3 a Z a 2 2 
a 1 6 6 
> | Gasteropoda ......... 3) 15 u : 2 5 g 
1 | Bteropodarsscvadaces- 1 1 i 1 
1 ° 1 a 1 
1 | Heteropoda ......... 1 6 3 x 4 2 
2 | Cephalopoda ......... 3 8 2 3 1 1 
50 22 52 5 4 ; 
68 204 43 120 130 98 a5 nice 


Uprrer Luanpovery, on May-Hitt Grovp. 


Perhaps no formation in the Lower Palseozoic series is more diffi- 
cult to understand, either physically or zoologically, than the “ Upper 
Llandovery,” or May-Hill group. Its place, stratigraphically, was 
long ago settled by Sedgwick, who was the first to point out the 
necessity for separating these beds from the Lower Silurian (his 
Cambrian), and removed them from the Caradoc in 1853, proposing 
the name of the ‘‘ May-Hill Sandstone ” for these beds, being above 
all his Cambrian Rocks; and this is so throughout the northern 
hemisphere. 

On the west flank of the Malverns, at Woolhope, May Hill, and 


ANNIVERSARY ADDRESS OF THE PRESIDENT. I51 


Tortworth, in many localities in Shropshire (Norbury, Cherbury, 
Church Stretton, &c.), and at Llandovery, Presteign, &c. in South 
Wales, the Upper Llandovery or May-Hill beds are typically exposed. 

Many names have been proposed for these variable beds, such as 
Upper Caradoc, Pentamerus-beds, Wenlock Grit, &c.; but, through 
their chief development at Llandovery and May Hill, in Gloucester- 
shire, the double name of the ‘‘ Upper Llandovery,” or ‘‘ May-Hill 
Sandstone,” has been given to them. In North Wales the Upper 
Llandovery rocks are absent ; m South Wales they first appear in 
Marloes Bay, appearing here and there across Pembrokeshire. Near 
Llandeilo, in Caermarthenshire, they lie at the base of the Upper 
Silurian rocks, being most variable in thickness. They transgress 
or lie indifferently and unconformably on Lower Llandovery, Caradoe, 
or Llandeilo beds. In Shropshire they are markedly unconformable 
to the Caradoc. In the Longmynd area, under the condition of a 
caleareous conglomerate, they rest upon all the older rocks exposed. 
That a great physical break takes place between the two Llandoveries 
there can be no doubt. They rise from under the Wenlock Shale at 
Noeth Grug, strike to near Llandovery and Pen-y-lan; again they 
range uninterruptedly from Marloes Bay, in Pembrokeshire, where 
they are highly fossiliferous (28 species occurring there), through 
Caermarthenshire to Builth, and on to the Longmynd and typical 
Silurian country of Shropshire. They everywhere rest unconform- 
ably upon the older rocks, sometimes lying on the denuded edges of 
the Lower Llandovery or on the Caradoc Sandstone ; at Builth and 
the Longmynd on the contorted and denuded Arenig, Llandeilo, and 
Cambrian strata. 

The absolute unconformity of the Upper Llandovery beds to the 
strata below, coupled as it is with changes of species, is doubtless 
connected with a lapse of unrepresented time. Whether that time 
be of great duration or not we have no means of judging; but, 
looking at the intimate connexion between the fauna of the Lower 
Llandovery and that of the Upper, we are led to suppose that it 
was not of sufficiently long duration to cause either the extinction 
or migration of the older fauna or the wmtroduction of a new one 
(only 4 genera seem to have appeared); for we have seen that the 
Lower Llandovery transmitted 45 genera and 104 species, out of 
its fauna of 68 genera and 204 species, to the Upper Llandovery. 
It is therefore evident that upheaval and denudation must have 
been of comparatively short duration, and little physical change 
could have taken place in the area occupied by the Lower Llan- 
dovery after upheaval ; this the physical geography and paleontology 
of the two groups help to show. | 

The preponderance of individuals in species of the genera Pentc- 
merus and Stricklandinia, especially P. oblonqus, S. lens, and S. lirata, 
which are the best known and most widely spread, is a marked 
feature in the Upper Llandovery rocks, and imparts to them peculiar 
and distinct facies. These species occur, though not so plentifully, 
in the Lower Llandovery. The Malvern and May-Hill form is 
S. lens, and it oceurs in profusion. Looking at the grouping of the 

m 2 


152 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Upper Llandovery and the distribution of its fauna, we are surprised 
to find that, of the 261 species which occur in the formation, only 
136 species really constitute the Upper-Llandovery fauna. The fact 
that 104 of the species are Lower Llandovery goes very far to unite 
the two so-called formations into one, or a Middle-Silurian group, 
abolishing the terms Lower and Upper. Granting the high value of 
the unconformity as of paramount stratigraphical importance, yet 
the community of fossils and general facies of both horizons lead us 
to regard the species as belonging to neither; for certainly the Lower 
Llandovery was an expiring close to the Caradoc and Bala, through 
elevation of the Caradoc sea-bed, and the Upper through continuity 
of certain species ; and the introduction of new forms commenced at 
the base of the Upper Silurian of Sir R. Murchison and the Geological 
Survey, from which, in the Wenlock rocks, were derived 58 genera 
and 125 species; for out of the 523 Wenlock, 125 are Upper Llan- 
dovery, 136 species only being the true Upper Llandovery fauna 
proper. 

Pranta#.—One species only, a fucoid (Pucordes gracilis), is known 
in this group of strata. This is probably only one of the many forms 
of Annelide or molluscan tracks or burrows, which simulate the 
habit of some of the marine alge. 

Pror0z0a.—Clathrodiciyon vesiculosum, Nich., and Cliona ( Vioa) 
gracilis constitute the only two Protozoa known. The former occurs 
in the Llandovery rocks of Scotland; the latter appears to be a 
dichotomizing, burrowing sponge, found in the shelly structure 
of a Pterincea (P. demissa) from the Upper Llandovery of the Mal- 
vern Hills. This is probably the oldest known burrowing sponge ; 
its habit is quite that of the modern genus Cliona. 

The Welsh Llandovery rocks, although so carefully searched, have 
yielded no traces of Spongide. We should, however, quite expect 
to find the group Silicipongie, through the Hexactinellide, repre- 
sented in these sediments or formations accumulated in moderately 
deep water, and associated with an extensive Coelenterate and 
Brachiopodal fauna, such as we have in the Upper Llandovery. The 
Analytical Table (p. 159) shows that only in two of the ten 
(Worcestershire and Scotland) are the Protozoa known. 

iypRozoa.—The genera Monograpius, Oyrtograptus, Duplograptus, 
and fetiolites alone seem to represent the Rhabdophora in the Upper 
Llandovery rocks ; but so many species appear to be common to the 
Lower and Upper Llandovery, that it is with difficulty I am enabled 
to realize the distinctness of the two faunas. The Irish, Scotch 
(Girvan, Gala, and Mulloch beds), and the Coniston flags and mud- 
stones have so many species in common with the Lower Llandovery 
that only through intimate and practical acquaintance with the two 
groups of species can they be separated. In the Valentian or 
Llandovery-Tarannon series of Lapworth, including the Birkhil 
series and its several zones, so many species of Monograptus, Deplo- 
graptus, and Cyrtograptus appear to be common, that it is no easy 
task to determine numerically or statigraphically, especially when 
geographical distribution is also taken into consideration, their true 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 153 


history. Probably the Upper Llandovery rocks of Ireland yield 6 
or 8 species, the Girvan beds about 30, the Gala group the same, the 
Mulloch beds 25 species, and the Coniston beds 25 species; but many 
also belong to the Lower Llandovery belowandelsewhere. The species 
in two of the genera above named, AMonograptus and Cyrtograptus, 
increase in the Wenlock; in which, if we include the species which 
occur in the Tarannon Shales lying at the base of the Upper Silurian 
(Lower Wenlock group), some 9 genera and 20 species may occur. 
As in the Caradoc, it is extremely difficult to arrive at the number 
of true species that range through the areas, owing to not having 
correct knowledge of the species. 

Actinozoa.—This is, zoologically, an important group in the Upper 
Llandovery, from the number of genera occurring and the numeri- 
eally few species, 11 of the 16 genera being represented only by 
1 species each. These are Alveolstes, Coenites (probably one genus), 
Halysites, Labechia, Omphyma, Plasmopora, Ptychophyllum, Lind- 
stroma, Propora, Pinnacopora, and Streptelasma. Thus only 5 
genera constitute a specific fauna, viz. Favosites 4 species, Heliohtes 5, 
Paleocyclus 2, Petraia 7, and Syringopora 3. Their dispersion or 
distribution is equally significant, showing that the two chief com- 
pound genera, Mavosites and Heliolites, are, with one exception 
(Peiraia), the most widely distributed geographically. Petraca 
occurs in all the 14 localities* ; and perhaps no single genus has so 
wide a range or such persistent representative forms, commencing 
in the Caradoc with 6 species, having the same number in the 
Lower Llandovery, and 7 species in the Upper Llandovery. It is 
reduced to 5 in the Wenlock, and 1 (P. bina) in the Lower Ludlow. 
We know also of 3, if not 4, Petracw in Lower, Middle, and Upper 
Devonian ; they are, however, distinct species, being P. celtica, 
Lonsd., P. gigas, M‘Coy, and P. plewradialis, Phil. As regards the 
number of genera and species numerically of value, or important 
in certain areas, we find that in Pembrokeshire and Cardiganshire 
these are but feebly represented. The former county, at Marloes 
Bay, has given us 3 genera and 4 species, viz. Mavosites asper, 
Omphyma turbinata, Petraia bina, and Petraia subduplicata; and 
Cardiganshire only 1, Petraia elongata. Radnorshire has yielded 
2 genera and 3 species, Heliolites interstinctus, Petrara elongata, 
and P. subduplicata. These few and local species are significant 
and suggestive as to the cause of their restricted numbers. 
Worcestershire, through the Malvern area, is richest in species, its 
7 genera and 18 species being the highest known in the nine areas— 
Caermarthenshire and Caernarvonshire having 12, Gloucestershire, 
Scotland, and Ireland each 10 species, and South Wales generally 14. 
Of the 16 genera and 32 species that range through the Upper Llan- 
dovery beds, 14 genera and 22 species pass up into the Wenlock. The 
3 genera peculiar to this horizon are Propora (P. Edwardsu), Pina- 
copora (P. Anderson), and Lindstromea (L. levis). The remaining 
genera and species all pass to the Wenlock series, the Coelenterate 
fauna of which numbers 76 species. 

* P, subduplicata has recently been discovered in Scotland. 


154 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Ecuinopermata.—lt is clearly shown that this group requires to 
be more carefully collected and studied. We have 5 genera and 5 
species, or only | representative species to each genus. None occur 
in Shropshire or Scotland. Two species, representing 2 genera, 
have been met with in Pembrokeshire. The species are Actino- 
crinus pulcher?, Cyclocystoides Davisu, Salt., Paleaster coronella, 
Periechocrinus monliferus, and Paleechinus Phillipsie, Forbes— 
the last being the first known representative of the Kchinoidea in 
time. The Perischoechinide, to which family this genus belongs, 
attained its maximum development during the Carboniferous 
period, both in Britain and America. No fewer than 7 genera 
have been recognized in America and 4 in Britain. In the test 
of Paleechinus the interambulacral plates abut against each other, 
not being bevelled for overlapping articulation, as is the case with 
Perischodomus, Oligophorus, Melonites, and other genera. The family 
Archeocidaridee, of which the Carboniferous genus Archwocidaris 1s 
the type, does not occur below the Carboniferous rocks in any known 
area. The modern representative, ‘‘ Calveria” (C. hystrix) of Prof. 
W. Thomson, has been dredged in 445 fathoms water (2670 feet) in 
lat. 59° 38’ N., long. 7° 46’ W. Calveria differs from all known 
recent Kchinoidea through the structure of the plates of the test, 
which overlap, instead of abutting against each other, as in all other 
genera. Again, the plates composing the interambulacral areas over- 
lap from the apical disk towards the mouth, and those of the ambu- 
lacral area from the mouth towards the apical disk, or in the reverse 
direction. This ancient type appears in the Chalk as Echinothuria 
(E. floriformis); and now, through the dredgings of the ‘ Porcupine,’ 
we have proof of ifs continuity from the Cretaceous epoch. 

AnnrLIpA.—Three genera, with four species, two of which claim 
attention here, viz. Ventaculites (ornatus, Sow.), and Cornulites (serpu- 
laria, Schloth.). They have occurred in almost every locality since 
their first appearance in the Caradoc and Bala beds. Tentaculites 
ranges through the Caradoc everywhere except in Scotland ; it is 
abundant in the Lower Llandovery of South Wales; but its maxi- 
mum of development takes place in the May-Hill Sandstones, where 
it is a marked feature in the fauna. It is sparingly exhibited in the 
Wenlock shales and sandstones, Cornulites having replaced it indi- 
vidually in those beds. TYentaculites anglicus, T. ornatus, and 
7’. tenwis are also Ludlow species; and the long-lived 7. anglicus 
either passes to the Middle Devonian slightly modified, or is replaced ~ 
by Schlotheim’s species 7’, scalaris. 

The 3 genera and 4 species (Cornulites serpularius, Tentaculrtes 
anglicus, T. ornatus, and Spirorbis Lewis) pass to the Wenlock. 

The Cephalobranchiate or Tubicolar Annulosa (Annelida) play an 
important part in the life and distribution of the class through 
time. Their habits insure persistency, their structure security, and 
their bathymetrical ranges are such as to lessen their chances of 
modification through relative changes of land and sea. No form of 
the order Errantia, or Dorsibranchiate Annelida, has been detected 
in the Llandovery rocks, although, doubtless, they existed, as many 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 155 . 


of the so-called fucoidal remains are only tracks of these creatures ; 
and their presence has been demonstrated by Mr. G. J. Hinde 
through his discovery of the jaws of many species of the Poly- 
cheta in the Silurian, Devonian, and Carboniferous formations. 
In the Wenlock alone he has discovered the remains of 6 genera 
and 24 species; and in the Ludlow 2 of the same genera and 6 
species. No one can doubt that, by careful sifting and preparation, 
the clays and shales of the still older rocks may yield a rich harvest 
of the remains of the Annulosa and soft-skinned Echinoderms. This 
process is now bearing fruit through the energetic measures em- 
ployed by Mr. G. Maw, in having large quantities of the Wenlock 
Shales washed, sifted, and examined. Numbers of new forms have 
been discovered. The same result has been obtained through the 
determined and persistent search made by Mr. Bennie (the Collector 
for the Scotch Survey) for remains of the Holothuride in the Car- 
boniferous shales of Scotland, and also for the Chitonide. ‘The 
results obtained by this patient worker, and the species collected, are 
now being described and figured for the Glasgow Geological Society 
by Mr. Etheridge, Junior. 

It is now the age of microscopical investigations, and ere long 
microscopic paleontology will stand on the same footing with petro- 
logical and crystallographic investigations. 

Crustacea.—The distribution of some of the species of the Crus- 
tacea through the Lower Llandovery is more constant and uni- 
versal than in any other Silurian deposit. Calymene Blumenbachu, 
Eincrinurus punctatus, Ilenus Thomsom, Proétus Stokesw, and Pha- 
cops Stokesi occur in almost every locality, and all 5 pass to the Wen- 
lock. 7 of the known 24 species do not pass to any higher horizon than 
Upper Llandovery; they are Illenus emuius, I. Bowmanm, I. Mac- 
callunn, Phacops Weaveri, P. obtusicaudatus, Lichas laxatus, and Tri- 
nucleus concentricus. .'To show how equally the species are distri- 
buted, I may state that in Caermarthenshire 7 genera and 12 species 
occur ; in Pembrokeshire 7 genera and 11 species ; in Worcestershire 
7 genera and 13 species ; in Shropshire 7 genera and 15 species; and 
in Ireland 7 genera and 14 species; and, singularly also, 7 genera and 
14 species pass to the Wenlock. As we have before seen, 10 species 
are Lower Llandovery and Caradoc—thus, as regards the Crustacea, 
rendering the value of the Upper Llandovery as a distinct paleon- 
tological group more questionable still. 

Bracuiopopa.—tThe species in this class are more than double the 
number of those of any other in the Upper Llandovery rocks. We 
recognize 65 species and 13 genera. The geographical distribution 
of Atrypa (5 species), Leptena (4), Orthis (10), Pentamerus (4), 
fihynchonella (9), Stricklandinia (2), Spirifera (5), and Strophomena 
(11) is complete through South Wales, Gloucestershire, Worcester- 
shire, and Shropshire. These 8 genera include the mass of species 
(50 of the 65) that range through the fourteen localities I have 
selected, and whence I have drawn the materials for analysis, and 
I believe I have omitted none. Lingula with 5 species, Crania and 
Discona each with 1, Chonetes with 3, and Meristella with 6 species 


156 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


are sparingly and irregularly distributed ; but we should expect this 
from the habits and habitats of 3 at least of the genera. No less 
than 36 species come up from the Lower Llandovery (22 of the same 
are Caradoc); and 41 of the 65 range into the Wenlock beds. As 
among the Crustacea, few species belong to the Upper Llandovery 
proper, showing how slight is the evidence afforded us to separate the 
Lower and Upper Llandovery. The stratigraphical unconformity is, 
however, completely borne out by a corresponding paleontological 
break. This is most decisively seen from the Actinozoa, the 
Crustacea, the Brachiopoda, and the Lamellibranchiata; indeed Car- 
diganshire has as yet only yielded | species of Actinozoa, Petraia 
elongata, M‘Coy. The Upper Llandovery rocks of the nine chief 
areas given below have yielded the following number of genera and 
species of Brachiopoda :— 


Counties. Genera. Species. 
Remibrokeshines 7.40 ae. 11 19 
Caermarthenshire ........ a, 38 
Cardiganshirenc. seen oe none. none, 
IRGOGESOS So esessacece 6 De 
Gloucestershire .........-. 9 it 
Wrorcestershires 4 ae att 35 
Shmopsiune yeti ae ae 13 13 
SCOGlAMGs 2s cc ct ee aiee mee 9 De, 
Terelamndiy: case ee eo oun 10 19 


No less than 11 genera and 41 species pass up to the Wenlock 
group, thus showing that a greater number of species in this class 
pass up than belong to any even of the most prolific areas. 

LAMELLIBRANCHIATA.—Considering the few species known in the 
Lower Llandovery (3), we have here a marked change in the greatly 
increased fauna of bivalve Mollusca, which have increased from 
the 3 genera and 3 species of the Lower Llandovery (Pterincea retro- 
flexa, Mytilus mytilimeris, and Orthonota sulcata) to 12 genera and 
29 species. 2 of the 3 species above named pass to these upper beds, 
so that the specific fauna is essentially new; in the end, however, 
it gives to the Wenlock 7 genera and 15 species; or 50 per cent. 
pass to the Wenlock. The Worcestershire area is most prolific, 
12 species occurring. Only 3 species are known in Scotland, 
Anodontopsis bulla, Pterinea Sowerbyt, and Cardiola striata, and 3 
in Ireland, Anodontopsis bulla, M‘Coy, Pterinwa bullata, M‘Coy, 
and Ctenodonta subcylindrica, M‘Coy. No species is known either 
in Radnorshire or Caermarthenshire. (Vide Table No. XII.) 

GastEropops.—Kqually balanced with the above group, the dis- 
tributional value being through species of Holopella, Acroculia, ha- 
plistoma, and Turbo. These species, too, have a wide geographical 
distribution, Worcestershire, as before, possessing the greatest num- 
ber of species (9), although here only equal to Ireland, which has 
also 9 species; while Caermarthenshire has 7, the remainder of 
the 13 genera and 28 species being equally distributed. 6 genera 
and 8 species pass to the Wenlock; they are Acroculia halts, 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 157 


Euomphalus alatus, E. frenatus, E. sculptus, Holopella obsoleta, 
Loxonema sinuosum, Murchisonia articulata, and Turbo tritorquatus. 
No form occurs in the Upper Llandovery of Cardiganshire. 

Preropops.—Only | species is known in the Upper Llandovery. 
Conularia cancellata, Sandb., occurs in Gloucestershire and Shrop- 
shire, and is known in no other formations or localities in Britain. 
The more abundant C. Sowerbyi is Lower Llandovery, Wenlock, 
and Ludlow. It is questionable if C. cancellata is really a British 
species or only a variety of C. Sowerbyt. 7 of the 10 areas have 
no represesentative of this group. 

Herzroropa.—Leculiomphalus (Cyrtolites) and Bellerophon, the 
. former having 2 species (#. levis, Sby., and £. scoticus), the latter 
9, range pretty equally through the Upper Llandovery rocks, 
B. dilatatus, B. trilobatus, and B. carinatus haying the widest or 
most extended geographical distribution. The rarer forms, Hecul- 
omphalus scoticus, M‘Coy, Bellerophon subdecussatus, M‘Coy, B. wen- 
lockensis and B. obtectus, Phill., have as yet only occurred each in 
one locality ; 5 of the 11 have appeared before in the Caradoc; but 
only one species, B. carinatus, seems to have been Lower Llandovery. 
These Pelagic Mollusca, especially the family Atlantide, have no fixed 
habitat, their distribution being quite independent of the nature of 
the sea-bottom, so that the Bellerophons of the Silurian, Devonian, 
and Carboniferous periods occur indifferently in mechanical deposits 
of the most varied petrological or lithological characters. The great 
Poreellia (P. Woodwardit, Sby.) of the Middle Devonian, or the two 
species P. striata and P. Symondsii of Phillips, occur in all sorts of 
deposits all through the history of the genus, of which 12 or 14 
species range from the Devonian to the Trias, Belgium having 
afforded many forms. Conrad’s genus Cyriolites, probably the 
Ecculiomphalus of Buckland, is represented in Britain by 3 species 
—H. Bucklandi, Caradoc only; £. levis, Upper Llandovery, Wen- 
lock, and Ludlow; and Z. scoticus, Llandeilo, Caradoc, and Upper 
Llandovery. Of the 11 Upper Llandovery species 6 pass to the Wen- 
lock, 5 being species of Bellerophon, and 1 Eeculiomphalus (EL. levis). 

CrpHatopopa.—Of the 6 genera occurring in the Upper Llan- 
dovery, viz. Actinoceras, Cyrtoceras, Lituites, Phragmoceras, Treto- 
ceras, and Orthocerus, one (Actinoceras nummularium) appears for 
the first time, and the rare forms Phragmoceras compressum and 
Cyrtoceras upproximatum have but local geographical distribution. 
The last named is from the Upper Llandovery of Malvern only; 
P. compressum the same, being of doubtful occurrence in the Ludlow 
rocks. We have seen that the Caradoc rocks of Shropshire have not 
yielded a single species of Cephalopod; but here, in the Upper Llan- 
dovery of the same county, we have 3 genera and 9 species; yet 8 
genera and 47 species occur in the Caradoc rocks. This fact, coupled 
with others to be arrived at through the tables of distribution, 
tends to show the shallowing of the Caradoc sea and slow elevation 
of the land through the latter part of the Caradoc period, and the 
time represented by the deposition of both the Lower and Upper 


158 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Llandovery series. 4 genera and 9 species pass to the Wenlock. 
Ireland has only 2, Orthoceras coralliforme, M‘Coy, and O. subgre- 
garium, M‘Coy. Radnorshire has 1 species, Litwites cornu-arietis, 
Sow. Taking South Wales generally, only 6 species are known 
there, and these represent 4 genera—Actinoceras 1 species, Lituites 
1 species, Orthoceras 3 species, and T'retoceras 1 species. 

The Upper Llandovery rocks of Scotland give us 4 species, 
3 Orthocerata and 1 Phragmoceras (P. compressum),. 

We therefore find that, out of the whole fauna of the Upper 
Llandovery rocks, numbering 240 species, only 91 are confined to 
them and do not pass up. The following list shows the special 
Upper Llandovery fauna :— i“ 


RObOZ08) 44 Weinents ee 2 species. 
iNctamOzoa,! Joe aa are. Ae 10 “3 
Kehinodermata.......... 3 a 
Crustacea vik ae. ee 8 ee 
TTY OZ OA cphisual.-p senses net eenae evel igs 
Brachiopodarn es -.ntiee ee 22a ae 
Lamellibranchiata ...... 11 Be 
Gasteropoda)j2 OV isin kek ening 
Pteropodas. secigisiat wick & il: < 
Meteropodauc...u ee. aes RO ee 
Cephalopoda ss. er Dit.) U5 
i 
WENLOCK. 


I have stated that 58 genera and 125 species are common to the 
Wenlock rocks and the Upper Llandovery; in other words, these 
125 species pass up from the Llandovery to the Wenlock formation. 
Next to the Caradoc and Bala, the Wenlock group, embracing the 
Tarannon Shales, Denbighshire Grits, and Woolhope beds, possesses 
the largest Silurian fauna known; it numbers 168 genera and 530 
species. Hight chief groups or classes out of the fourteen furnish the 
majority of the species, in each case attaining their maximum in the 
Wenlock Limestone. ‘They are the following :— 


said now oa yi ats ees 6 genera and 30 species. 
2s Achimozoa nts dea a 25 45 ROr es 
3. Echinodermata ...... 28 genera and 68 species. 
4: Orustacealys)) 28). 210.8 27 3 Vestn Ones 
oi) brachiopodaniee ase 21 F Oss ees 
6. Lamellibranchiata .... 16 He AB yk 
Ge (Gasveropoda (ia. s en: 9 : PATRIA 
&. Cephalopoda =) .. 78 5 y DOT es 


I may add the Bryozoa, for at no period in the history of the 


159 


THE PRESIDENT. 


S OF 


ES 


SARY ADDR 


ANNIVER 


Taare XII.— Upper Llandovery. 


Geographical Distribution. 
net 
= o i 
w o Pat ot ats) P: 
i re q a a | 4 ae ass 
Se Classes. a g a = S g a g Os to 
oir : \ ng :: A rz 2 a =) 08 ag rd Wenlock. 
wb) 3 n ia &o Oo a4 o © n m 5 a 
o 
ES Ce Wee en coh ge erate eegin Sh calgon er enemas 
Sg | 3 Ee 3 = ae: = 3 3 be i= 5 
es B) a, ) @ @ S A oa! te ae So co) 
By a) MD or Oo o) ete wn O A) HO RQ 
Plone atcmamectsrakealt acl 1 oa 6 oe : 1 
EOUOZO 8 ax the ates catewete, 2 UE) Vee a vee ves wee ves te 1 “ vee i 
Heled |) HAV GIOZOA:. vac hes ccavtevcescule — 4 33 *F ... on ie ve tee oe nee vee 2's 30 
9 : " 3 5 3 10 6 7 4 7 
a0 UNGEINOZOBN. cate: A ctor oul RHO 32 4 12 i 3 14 10 18 9 10 22 
Eechinodermata..............., 5 i) 5 vee oC 1 5 I 2 vee 2 
/NOIMIOES coemaooddgecorcodecon|=. & 4 3 “ ves 1 3 3 3 3 vee ' 4 
8 7 7 6 7 6 7 7 6 
a Onustacode. ncn all Pie || art 13 : 8 13 & 13 15 vee 7 14 
2 . é N 2 2 BY 1 1 1 < 
2 BUYOZOS ayaacnaieesianescsecl 2 7 vee 3 vee ves 4 3 1 2 : 1 3 
10 . 7 tC 11 6 9 11 13 
PU EMECHIONOCay ins. cme: LG 65 Ae an im As ate 27 35 13 000) 12 44 
2 Lamellibranchiata .........) 12 29 7 4 vee see 3 8 2 é ee 1 15 
6 : ‘ 4 3 9 3 4 4 3 ; 
g 1) Grasteropoda...c.c.e.0 LO 28 3 7 3 11 é 9 5 nee 4 8 
UREGHOMOG Esta dew. tevteenccw cs 1\ Tit exe fo BEC Guie ade A eee i 
2 1 1 1 1 1 1 1 1 2 
? Re terO pode. .ce.cmeacnnl wee Hel IY PE i . 1 3 1 3 7 os 4 6 
1 | Cephalopoda ...... cenceualh Ome ie Ong 4 aa 1 é : 3 ve 4 9 
A5 38 42 22 54 41 48 41 6 35 59 
104 thse eeeeees 91 261 H 658 90 I 40 106 70 VOR ke 25 74 126 
| 
i ae iS a 2) eS Np I Sa Le lA Nhe al ee ee al eee eel ee ARR ge 


* It is difficult to decide with regard to these species which are Upper and which Lower Llandovery. 
t L also give these 6 genera and 25 species in the Upper Llandovery as well as the Lower, for I really know not whose view to adopt as to their 


stratigraphical position, Mr, Avyeline and Prof, Hughes place them in the Tarannon Shales, Mr. Lapworth and Dr. Nicholson in the 
Lower Llandovery. 


160 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Lower Paleozoic rocks were they so largely represented, and all the 
known forms in the Wenlock group (11 genera and 24 species) 
occur in the Wenlock Limestone and Shale, and in those of England 
alone, only 1 species occurring either in Scotland or Ireland. 

Protozoa.—Chona prisca and Spongarium Hdwardsii are both 
Denbighshire-grit species, the latter being also Wenlock and Lud- 
low; Cnemidium tenue occurs in the Wenlock only, and Ischadites 
Grindrodi in the Woolhope; the other well-known species, J. 
Keng, Murch., has a wider range both in time and space, and has 
occurred in the Wenlock rocks beneath the Cretaceous series at 
Ware, in Herefordshire, in the Silurian cores brought up from a 
depth of 1000 feet, during trials for the better supply of London 
with water. 30 Wenlock species were obtained here from a few 
feet of cores; they are identical with the Wenlock species of Dudley 
and Wenlock Edge. (Vide page 229, in the Chapter upon the 
extension of the older rocks under the London area.) The singular 
and still doubtful genus Stromatopora (NS. striatella) occurs plentifully 
in the shales and limestones of the true Wenlock beds, and is also 
found in the Aymestry Limestone. This genus comprises a large 
number of Silurian and Devonian fossils of every size and form. The 
affinity of the Stromatoporids is still doubtful; by some authors they 
are placed with the Hydrozoa, in the subclass ‘“‘ Hydrocorallina.” 
The doubtful Devonian genus Caunopora suggests much research, 
both from its abundance and peculiar structure. In Britain Stroma- 
topora first appears in the Caradoc and Bala group, ranging upwards 
into the Middle Devonian of Devonshire. 

Hyprozoa.—The Wenlock rocks (assuming the Tarannon and 
Denbighshire beds to be at their base) have yielded 6 genera and 30 
species. Stratigraphically they occur chiefly in the Tarannon beds, 
or probably the Denbighshire Grits. So far as | know there are 
none in the Woolhope beds, and I omit them from the column 
headed Wenlock, as they occur chiefly at Builth and in North Wales, 
and in this case would be repeated. The T'arannon or Denbighshire 
beds have yielded 5 genera and 23 species; North Wales 1 genus 
(Monograptus) and 8 species; South Wales 3 genera and 15 species ; 
Westmoreland 3 genera and 16 species; Scotland 3 genera and 5 
species. 

Acrinozoa.—At no period during the progress of Palseozoic time 
was there so large and rich a Coralliferous fauna as during the 
Wenlock period; numerically the Middle Devonian about equalled 
the Wenlock species in number, the genera and species being in both 
deposits much the same. The Wenlock rocks of Britain yield 25 
genera and 76 species, the Devonian 24 genera and 52 species, 
and the Carboniferous 36 genera and 141 species. 

I believe every known Wenlock species (there are 76 of them) 
occurs in the Wenlock Limestone and its subordinate shales. In the 
lower division, especially the Tarannon Shale and the Denbighshire 
Grits, the species are few, not more than five being known in the 
former and four in the latter. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 161 


The Tarannon species are Havosites asper, I’. fibrosus, F. gothlan- 
dicus, Petrara bona, and P. subduplicata. The four Denbighshire-grit 
species are also Favosites aspera and I’. fibrosus, Petraia subduplicata 
and Syringopora serpens. The upper member of the Lower Wenlock 
(the Woolhope beds) yields 16 genera and 26 species, most equally 
distributed generically. The North-Welsh Wenlocks yield 7 genera 
and 11 species, and the South-Wales beds 10 genera and 18 
species ; Scotland 8 genera and 11 species; and Ireland 14 genera 
and 23 species. 

We must not forget that 14 genera and 22 species came from the 
Llandovery rocks, and 18 pass to the Ludlow group, 13 of which 
were also Llandovery ; thus 41 species are peculiarly Wenlock. 

Ecutnoprermata.—As compared with the Upper Llandovery, with 
only 5 genera and 5 species, and the Lower Llandovery only 2 
species, the Wenlock sea must have been highly favourable to the 
development of the Echinodermata ; no greater development of life 
is known in any formation, for 60 of the 68 known species of 
Echinodermata were new appearances in the British area. Many 
are American ; 65 of the 68 are Wenlock Limestone; no species is 
known in Scotland, and only 1 in Ireland (Acéenocrinus Wynne, 
Baily); only 1 species occurs in N. Wales in the Tarannon Shale 
(Actinocrinus pulcher), and only 2 in the Woolhope (Lucalyptocrinus 
polydactylus, M‘Coy, and Pisocrinus pilula, De Kon.). 

We should expect that out of so large a fauna many species would 
pass to the higher divisions of the Upper Silurian, but only 4 genera 
and 6 species pass to the Lower Ludlow. These are of the Crinoidea, 
Actinocrinus pulcher, Ichthyocrinus pyriformis, and Ichthyocrinus 
M‘Coyanus ; of the Cystidea, Pseudocrinites magnificus, P. quadrifas- 
ciatus; and of the Asteroidea, Rhophalocoma pyrotechnica. Ichthyo- 
crinus pyriformis is the only species occurring in the Aymestry 
Limestone ; the Upper Ludlow only holds 1 species in common with 
the Wenlock, the long-lived Actinocrinus pulcher. No less than 20 
new Kchinodermal genera made their appearance in the Wenlock 
sea, and 17 of them are confined to the Wenlock rocks; the 4 genera 
that pass to the Ludlow are Ichthyocrinus, Taxocrinus, Pseudocri- 
nites, and fhophalocoma. ‘This marked, sudden, and important 
addition to the 3 orders of the Echinodermata can only be explained 
upon the theory of migration from some prolific area; whence it is 
most difficult to suggest ; but looking at the great generic and specific 
development of the Crinoidea, Cystidea, &c. in N. America, and 
the great resemblance, if not identity, of the faunas in the two 
areas, would lead one to surmise that it was from the west rather 
than from Europe that the Wenlock sea derived these HKchino- 
dermata. 

Awnetipa.—No other British Palxozoic strata possess so many 
species of Annelida; they number 35. The Upper Llandovery 
possesses only 4 species, the Lower Llandovery 3, the Caradoc 16, 
the Llandeilo 3, and Ludlow 17 species. 

This exuberance in the Wenlock strata is due to the researches of 


162 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


one naturalist *, through whose careful investigations no less than 6 
new genera and 24 new species have been added to the Annelidan 
group of the Wenlock. We know of but one Wenlock species in 
Scotland or Ireland. All except one species occurs in the true 
Wenlock Limestone and shales: 4 species pass to the Lower Ludlow 
(Helminthites, sp., Serpulites dispar, S. longissemus, and Sprrorbis 
Lewis) and 8 to the Upper (viz. Cornulites serpularius, Serpulites 
dispar, S. longissimus, Tentaculites anglicus, T’. ornatus, Tr ach yderma 
squamosa, and 2 species of Arabellies). 

CrustacEA.—Ten new genera appear here for the first time; it is 
important to name them—Turri lepas, Aichmina, Cyphaspis, Deiphon, 
Eurypterus, Hemiaspis, Pterygotus, Thlipsura, Cythere, and Cythe- 
rellina. 'The entire Crustacean fauna consists of 27 genera and 78 
species; the Upper Llandovery transmits to it 7 genera and 14 
species: 74 of the 78 species occur in the Wenlock Limestone ; and the 
4 wanting species are Homalonotus cylindricus, Ceratiocaris ludensis, 
Primitia excavata, and P. lenticularis. The Woolhope beds hold a 
remarkable Crustacean fauna, comprising 13 genera and 24 species : 
this class and the Brachiopoda are the only two marked zoological 
groups in this division of the Lower Wenlock series ; 12 of the Trilo- 
bita are Upper Llandovery, and only 1 is distinctly a Woolhope 
species, viz. Homalonotus cylindricus, Salt. The remaining 3 species 
of Crustacea in the Woolhope are Primitia excavata, P. lenticularis, 
and Cythere Grindrodiana, Salt., all belonging to the Ostracoda ; 
they are of no value. We thus rather unexpectedly find that the 
Woolhope formation possesses groups of fossils almost equal to 
those of the Wenlock Limestone; such as the Actinozoa with 26 
species, 18 of which are also Upper Llandovery; the Crustacea 
with 24 species, 19 of which are Trilobites; the Brachiopoda 
with 17 genera and 56 species out of the 21 genera and 96 species 
known in the Wenlock Limestone. The poverty of the Crustacea 
in the Tarannon and Denbighshire Grits might be expected from the 
lithological characters of the rocks; only 8 of the 78 species occur 
in the former, and 5 in the latter horizon. North Wales has only 
yielded 5 genera and 11 species, and South Wales 9 genera and 14 
species ; only 3 species occur in Scotland—Turrilepas Wirightianus, 
Beyrichia Kleedeni, and Phacops Stokes. 'The Irish species number 
8—Calymene Blumenbachit, var. Allportiana, Cheirurus bimucronatus, 
fnerinurus punctatus, EL. variolaris, Phacops caudatus, P. nudus, P. 
Stokesti, and Proétus latifrons. The order Merostomata makes its 
first appearance in the Wenlock through Hurypierus punctatus, 
flemiaspis horridus, and Pterygotus problematicus, all in the Wenlock- 
Limestone series. This group of Crustacea has been extensively 
and critically worked out and illustrated in a masterly manner 
by Dr. H. Woodward, F.R.S., in the volumes of the Palzonto- 

graphical Society, the Quarterly Journal of the Geological Society, 
and the ‘Geological Magazine.’ The suborder Eury pterida i is repre- 


* G. J. Hinde, Quart. Journ. Geol. Soc. vol. xxxvi. pp. 368-378. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 163 


sented by 5 genera, 3 of which commence or first appear in this 
division of the Upper Silurian strata. No less than 12 species of 
Primitia, besides other Ostracods, have been determined by Prof. 
Rupert Jones, F.R.S., two of them being Woolhope (Cythere Grin- 
drodiana and Primitia lenticularis). Of the whole group of Crus- 
tacea, 13 genera and 18 species pass up to the Lower Ludlow, whose 
Crustacean fauna numbers 18 genera and 47 species. 

Bryozoa.—Of the 11 known genera that occur, 6 are new in 
the Wenlock. No less than 24 species are known in the shales 
and limestones; none have occurred in the Tarannon Shales, Den- 
bighshire Grits, or anywhere in North Wales or Scotland. In South 
Wales 2 species are known, Drastopora consimilis, Lonsd., and Péilo- 
dictya lanceolata; the same species occur in Ireland, and, with Escha- 
rina angularis, Lonsd., pass to the Lower Ludlow.  Fenestella 
assimilis, F. subantiqua, and Ptilodictya lanceolata were also Lower 
Llandovery species. The 6 genera not known in the older rocks, 
and which first appear in the Wenlock, are Cellepora, Ceriopora, 
Diastopora, Discopora, Hscharina, and Polypora. 

Most of the Paleozoic genera belong to the order Gymnolemata, 
suborder Cyclostomata. We know of no Paleozoic genera extending 
into the Secondary Period, where this class attains its maximum 
paleontological development, the Cretaceous system alone having 
yielded more than 200 species. 

BracwropopA.—In the Wenlock rocks, like the Caradoc, the Bra- 
chiopoda greatly predominate over most of the other groups. We 
have determined 15 genera and 109 species in the Caradoc; and now 
in the Wenlock are enabled to show that 22 genera and 101 species 
occur, being an increase of 7 new genera (namely, Athyiis, Cyrtia, 
Fiichwaldia, Nucleospira, Obolus, Orbiculoidea, and Retzia), the species 
being fewer so far as we know through the literature of the class. 
21 genera and 96 of the 101 species belong to the Upper Wenlock 
or Wenlock Limestone and shale*; all the known 22 genera are 
also represented in the same horizon. The Woolhope beds yield 17 
genera and 56 species, every Woolhope shell being also Wenlock. 
Orthis mullochensis, Dav., or its variety O. reversa, may stand alone 
as a Woolhope species, but every other form is equally Wenlock. 
Certain horizons in the Wenlock rocks are richer generically and 
poorer in species than others; this is shown in the Tarannon Shales 
and Denbighshire Grits, where, in the former, 10 genera occur 
with only 15 species; it is the same with the Denbighshire Grits, 
in which we have 14 genera and only 19 species. No genus or 
species is peculiar to either the Tarannon or Denbighshire beds; 
all are good Upper Wenlock forms. ‘Those species having the 
longest range in time in the Wenlock group number about 11; they 
also have a correspondingly wide range in space; they are :— 


* The researches of Mr. Davidson, F.R.S., upon the collection made by 
Mr. G. Maw in these beds has enabled him to add many new species and one 
or two new genera to the fauna of the Wenlock rocks. 


164 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Atrypa reticulata. 


Rhynchonella nucula. 
Chonetes striatellus. 


Spirifera elevata. 


Crania implicata. exporrecta. 
Leptzena transversalis. Strophomena pecten. 
Meristella tumida. rhomboidalis. 


Orthis elegantula. 


All those named have appeared in the Llandovery or Caradoc 
before, and all pass to the Ludlow; but their universality through 
the Wenlock renders them conspicuous or long-lived, though not 
perhaps the best witnesses in determining the age of any particular 
horizon through the Wenlock deposits; those species which are 
short-ranged, or essentially typical or confined to particular beds, 
should be named, as contradistinguished from those given above. 
No species is peculiar to the Tarannon Shale, although 10 genera 
and 15 species range through this series of beds. No species essen- 
tially marks, distinguishes, or is confined to the Denbighshire Grits ; 
yet 14 genera and 19 species also occur in this horizon. 

Although 17 genera and 56 species are known in the Woolhope 
beds, yet only 2 species are restricted to them; these are Orthis 
mullochensis, Dav., and Pentamerus undatus, Sby. This last species 
occurs in the Woolhope (?) of Ireland. 

Every Tarannon species except Lingula Symondsw had previously 
occurred, or appeared in one or other of the two Llandovery horizons 
or in the Caradoc. All the species in the Denbighshire Grits except 
three, Discina Morrisi, Meristella tumida, and Rhynchonella navicula, 
also have appeared in and passed up from the same deposits. Thus 
we find that the restricted Brachiopodal fauna (so far as we at pre- 
sent know) in these beds at the base of the Wenlock is extremely 
scanty. 

Out of the 101 known Wenlock species, 56 had not occurred in 
any earlier formation, but first appeared in the Woolhope and 
Wenlock Limestone; these newly introduced Wenlock forms repre- 
sent 20 genera. I deem it important to name them and give the 
numbers of the species in each genus; they are :— 


VNIgwAAt) Godgsosdacgac0504900000. o.species: §|) Ololtisy so peusasceses eee eeeee 3 species. 
Athy Dain cccssase care sleeniands-bi- Deities Orbiculoideayeac.-ceeeree is 
Orantas pecccen tee cosas ccc So One Orthig.:....Jssoshe ee eee Ons frat 
(ON Acti codaudocanaddeqoqoqouandee A Re Pentamenus ys -cseee ee eeeee Sar 
Diseina! tkvcerc teers coanees ge aa BRetzial since Waetenceeeeeeeeene Pap 
iWichiwaldiaeeece eee ree EN 4 Rhynchonellal) -s-ceeseeee Aner, 
LDS Oh AE), ma pdcagsussuscoso0coo Zieteee Siphonotretayy... cases eerie 1 ye 
IGHAGWIEY, Soncssgnuebsosreooods000 oa Spirilera. 2.) wcckece eee Subs 
Meristellapea tee 4 , Sirophomlenals- ceca eeeeeeee Sis ee 
INucleospiraie. a--ccnsde ser gael ir foes PTriplesia..c..25. ds. saan EO Fes 


Out of so large a fauna as 101 species, 96 of which occur in one 


division (the Wenlock Limestone and Shale), we should expect to 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 16 5 


find a large proportion absolutely restricted to that horizon. Of 
such restricted species in the true Wenlock rocks (limestones and 
shales) we have 24 or 25 per cent. None of the following species 
occurs in any other horizon; and they may therefore be accepted as 
definitely typical Wenlock species—<dAitrypa Barrandii, Day., Cra- 
nia Grayu, Day., C. Sedquicku, Lewis, C. siluriense, Dav., Cyrtia 
nasuta, Lewis, Leptena segmentum, Ang., Meristella nitida, Hall, 
Nucleosptra pisun, Sow., Obolus Davidsoni, Salt., Orbiculoidea 
Becketiana, Dav., Orthis equivalvis, Dav., O. basalis, Dalm., O. 
Lewis, Day., O. Hughes, Dav., Retzia Salteri, Dav., Siphonotreta 
anglica, Mor., Sprrifera sulcata, His., Strophomena Dayi, Dav., S. 
Fletchert, Dav., S. Henderson, Dav., S. Orbignyr, Dav., S. Walin- 
stedti, Linds., S. Waltont, Dav., and Triplesia MacCoyana, Dav. 

The Wenlock Brachiopoda of Scotland number 11 genera and 20 
species, most of them belonging to the long-range species, the more 
restricted being eristella Maclaren, Hasw., Nucleospira pisum, 
Sow., Orthis Lewisti, Dav., O. wenlockensis, Day., O. polygramma, 
Sow., and Strophomena Henderson, Day. 

The Irish Wenlocks contain 14 genera and 31 species; the most 
important are Obolus Davidsoni, Salt., Pentamerus undatus, Sow., 
Lthynchonella Beltiana, Dav., R. deflexa, Sow., Orthis rustica, Sow., 
Spirifera bijugosa, M‘Coy, and Strophomena Dayi, Dav. The genera 
numerically rich in species are Orthis (16 species), Rhynchonella 
(10), Sperefera (8), Strophomena (16), Pentamerus (6), Meristella 
(5), and Atrypa (5). The rarer genera are but feebly represented. 
12 genera and 32 species pass to the Lower Ludlow, 10 genera and 
20 species appear in the Aymestry, and 11 genera and 17 species in 
the Upper Ludlow. Again, the Wenlock is allied to the Caradoc 
and Llandovery groups through 11 genera and 41 species, 59 being 
Upper Llandovery, 22 Lower Llandovery, and 14 Caradoc species ; 
5 Llandeilo species lived on through all five periods, viz. Lepiena 
sericea, Orthis biforata, O. calligramma, O. elegantula, and Stropho- 
mena rhomboidalis. 

LAMELLIBRANCHIATA.—Out of the 16 genera and 44 species of 
bivalve shells ranging through the Wenlock rocks, the upper divi- 
sion or Wenlock Limestone yields 43 species, the one absent or 
uncertain form being doubtfully Lunulacardium aliforme, Sby. 

The Tarannon Shale contains, so far as we know, only 2 species, 
and both are Upper Llandovery shells, Pierinwa planulata, Conrad, 
and P. retrofleca, Wahl.; both pass to the Ludlow beds. The Den- 
bighshire Grits from all available sources yield 7 genera and 10 
species. The Woolhope only 3 genera and 5 species: these, with 
the Denbighshire-grit species, are all Upper Wenlock also. The 
local stratigraphical position of the ‘“* Woolhope or Barr” Limestone 
carries with it occasionally a peculiar fauna, and, although strictly 
of Wenlock age, there is always uncertainty as to the occurrence of 
few or many species, but it is next in numerical value to the Upper 
Wenlock. The Wenlock rocks of North Wales furnish 8 genera 
and 14 species, Pternea, Cardiola, and Cucullella being the chief 
genera. Pterinea has yielded 6 species, Cardiola 3, and Cucullella 

VOL. XXXVI. n 


166 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


3: other genera (Avicula, Arca, &c.) only one or two species 
each. 

In South Wales 11 genera and 26 species have been collected. 
The genera are the same, but with specific additions to Modiolopsis 
and Mytilus. 

Of the 3 Irish genera the 6 species are—Pterinea lineata, 
Goldf., P. planulata, Conr., and P. retrofleza, Wahl., Cardiola 
Jibrosa, Sow., and C. interrupta, Brod., with Goniophora cymbeformis, 
Sow.; none are confined to the Irish beds. Only 5 genera and 8 
species of Lamellibranchiata are really peculiar to the Wenlock 
series. 7 genera and 15 species came in from the Upper Llandovery; 
3 of these 15 are Lower Llandovery also, viz. Pterinea retroflexa, 
Cardiola interrupta, and Mytilus mytilimeris; 2 of the same are 
Caradoc, and 3 species came direct from the Caradoc; they are 
Pterinea tenuistriata, M‘Coy, Arca (Palearca) edmondiuformis, 
M‘Coy, and Ctenodonta anglica, D’Orb. Thus the Wenlock is united 
to the Upper Llandovery below through 17 species, and to the Lower 
Ludlow above by 21 species, and these all pass to the Upper Ludlow, 
so that the actual number of peculiar Lamellibranchs, or the 
Bivalve fauna, is reduced to the following 8 species :—Ambonychia 
tumida, Sollas, Pterinea asperula, M‘Coy, and P. laminosa, Goldf., 
Cleidophorus planulatus, Conr., Modiolopsis acutipora, Sollas, M. 
inflata, var., and M. chemungensis?, Conr., with Mytilus ungweu- 
latus, Salt. Nevertheless, assuming that the transgressional species 
also strictly belong to the rocks in which they occur, the Wenlock 
Bivalve fauna consists of 25 species by retaining those forms that 
pass up to the Ludlow. 

GastERoPoDA.—Looking at the splendid Ccelenterate, Echino- 
dermal, Crustacean, and Brachiopodal faunas in the Wenlock series, 
the distribution and ranges of which we have analyzed and tabu- 
lated, we should have expected this class to have been more 
largely represented, especially as during the- Caradoc and Upper 
Llandovery periods a larger molluscan fauna existed, although 
apparently the nature of the sea-bed was not so favourable as that 
of the more calcareous and argillaceous condition of the Upper 
Wenlock, in which the great mass of the Wenlock fauna occurs. 
All the genera belong to the order Prosobranchiata and division 
Holostomata. The Caradoc Gasteropoda, as we have seen, number 
14 genera and 53 species, the Wenlock 9 genera and 27 species ; 
the 9 genera represent 5 families. One species only occurs in the 
Tarannon Shale (Acrocula haliotis), and ranges through to the 
Wenlock Limestone and Ludlow. The Denbighshire Grits contain 
Acroculia haliotis, Holopella gracilior, Loxonema elegans, Murcha- 
sonia articulata, and M. Lloydi, or 4 genera and 5 species. ‘The 
Woolhope exhibits but a small fauna, only 8 species, 4 of which are 
Euomphali and 2 Acroculie, with Trochus (Cyclonema) exaltatus and 
Turbo tritorquatus. In this, as in all the other classes, nearly every 
species known occurs in the Upper Wenlock; thus the 9 known 
genera and 24 out of the 27 species belong to that horizon. 7 genera 
and 17 species occur in South Wales, but only 1 in Scotland 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 167 


(Euomphalus furcatus). Ireland yields 4 genera and 5 species 
(Murchisonia articulata, M. Lloydii, Euomphalus alatus, Cyclonema 
corallii, and Acroculia haliotis). We must remember that 5 genera 
and 6 species range from the Upper Llandovery; these are Huom- 
phalus alatus, Sow., HE. furcatus, Sow., E. sculptus, Sow., Holopella 
obsoleta, Loxonema sinuosa, Sow., and Murchisonia articulata, Sow., 2 
of the same occurring in the Caradoc, namely—Luomphalus alatus, 
Sow., 2. sculptus, Sow., and probably Turbo tritorquatus, M‘Coy ; and 
5 of these same 6 range to the Ludlow. Only 17 species out of the 
27 strictly belong to the Wenlock rocks proper. At present we 
know of no species other than those in the division Holostomata 
in the Silurian rocks. Hunema is the only new genus that appeared 
in this formation; £. cirrhosum is a South-Wales species. 
Preropops.—Only 3 species, Conularia Sowerbyu, Defr., Theca 
anceps, Salt., and J. Forbesii, Sharpe, have as yet appeared in the 
Wenlock group; none of the 3 occurs in the Tarannon Shale, in the 
W oolhope beds, or in Ireland. Theca Forbesii has been found in 
the Denbighshire Grits and Wenlock Limestone, and passes upwards 
to the Upper Ludlow. JZ. anceps is here Wenlock only, but is a Cara- 
doc species also. Conuwlaria Sowerby: first appeared in the Caradoc, 
and, with the exception of the Upper Llandovery, is present in all 


‘horizons up to the top of the Upper Ludlow; it also occurs in the 


Scotch beds. The only form of Thecosomatous Pteropod that trans- 
gresses the Silurian rocks is Conularia, which is Devonian and 
Carboniferous; but in Britain Conularia has not occurred in the 
Devonian series, although its associate, Bellerophon, is represented by 
5 British Devonian forms, 2 occurring in the Lower, 1 in the 
Middle, and 4 in the Upper Devonian. Conularia Sowerby: and 
Theca Forbesiz both pass to the Upper Ludlow. 

Hereropopa.—This pelagic order is well represented in the Wen- 
lock beds through Eecculiomphalus levis and 7 species of Bellerophon ; 
but no species is confined to or specifically characteristic of the 
Wenlock beds. All had previously occurred either in the Caradoc 
or Llandovery rocks. Hcculiomphalus levis, Bellerophon trilobatus, 
and B. carinatus range into the Upper Ludlow beds ; none occurs in 
the Tarannon Shale; 3 species are known in the Denbighshire Grits 
(Bellerophon trilobatus, B. expansus, and B. carinatus), and 3 in the 
Woolhope beds (B. bilobatus, B. dilatatus, and B. trilobatus). In 
the Wenlock Limestone we know of 4 species of Bellerophon and 
Ecculiomphalus levis. B. dilatatus is the only form known in 
Ireland; and Scotland yields none. The 3 species passing up or 
ranging into the Ludlow series are Ecculiomphalus levis, Bellerophon 
trilobatus, and B. subdecussatus ; there are none in the Aymestry 
Limestone. 

CEPHALOPODA.—5 genera and 30 species of the order Tetra- 
branchiata, illustrating the Nautilide and Orthoceratide, are known 
in the Wenlock rocks: all but 3species are represented in the Wen- 
lock Limestone and Shale ; these 3 are Woolhope species ( Actinoceras 
baccatum, H. Woodw., Lituites cornu-arietis, Sow., and Orthoceras coni- 
cum, Sow.). Actinoceras baccatum is essentially a Woolhspe species, 

R2 


168 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


described by Dr. H. Woodward, and the only one known from the 
Woolhope beds of Little Hope. Lituctes cornu-arietis, Sow., and 
Orthoceras conicum, Sow., although not ranging higher than the 
Woolhope beds, appeared in the Llandovery, and help to connect 
the Lower Wenlock with the Upper and Lower Llandovery. In 
the Denbighshire Grits we have almost unexpectedly 3 genera and 
7 species, only 2 of which are of Llandovery age (Orthoceras tenui- 
cmctum, Portl., and @. subgregarwum, M‘Coy). The remaining 5 
species pass from the Denbighshire rocks, where they are first known, 
to the Upper Wenlock ; they are O. primeevum, Forbes, O. tracheale, 
Sow., O. ventricosum, Phragmoceras nautilewm, and Cyrtoceras ibea. 
26 of the 30 species occur in the Wenlock Limestone and Shale; 13 
range through South Wales, and 12 through North Wales, 7 belonging 
equally to North and South Wales. The only 3 Irish species known 
are Orthoceras (Creseis) primevum, Forbes, O. subundulatum, Portl., 
and O. tenuiconctum, Portl.; and the 3 Scotch forms are O. Maclareni, 
Salt., O. subundulatum, Portl., and O. tenwemctum, Portl. 11 species 
pass to the Lower Ludlow; 2 of these, and the only 2 known 
(latuites giganteus, Sow., and Orthoceras angulatum, Wahl.), pass 
to the Aymestry Limestone; and 3 genera and 8 species range to the 
Upper Ludlow; thus only 4 genera and 9 species out of the 30 
are really Llandovery, or these 9 species come up from the Llan- 
dovery and Caradoc. 

The accompanying Table shows the numerical history of the 14 
classes. The first four columns illustrate the stratigraphical distri- 
bution of the species through the Tarannon beds, Denbighshire 
Grits, Woolhope beds, Wenlock Shales and Limestone, indeed all the 
formations above the Upper Llandovery. The succeeding five 
columns show the geographical distribution of the species through 
North Wales, South Wales, Westmoreland, Scotland, and Ireland ; 
and to show still further the connexion between the Wenlock and 
Ludlow fauna, I have stated in the remaining or last three columns 
the number of species that pass to the three divisions of the Ludlow 
group, in which it will be seen that out of the 171 genera and 536 
Wenlock species, 71 genera and 126 species pass to the Lower 
Ludlow, 38 genera and 52 species to the Aymestry, and 51 genera 
and 87 species to the Upper Ludlow; any higher transgression from 
the Ludlow into the Devonian is given under the Ludlow table. 
There are only 20 species, including the 6 Fishes from the passage- 
beds and the 8 species of Merostomata, that range through the 
Ludlow series. The oldest fish-remains occur in the Lower Ludlow, 
Scaphaspis ludensis, Salt., being the oldest species known. 


169 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 


| From Upper Llandovery. 


Plante. 


Classes. 


JEARO UB OVAOIEN © So ndedeAeaonanabaneses ea. 
Hydrozoa ...... Secuene 
ACtInIOZ02 \...¢.--. ate a CAGE 
HKehinodermata..... Arata Teohea 
Annelida 


Crustacea ........... SI or eee ate 


beoseersvesarsess ree eercsees 


Bryozoa 
IB GAC AIO DOC ames me ease 
Lamellibranchiata .. 


2200S ea eser 


Gasteropoda......... Pesea ean Cnataa 


Picropodae cae. 


etOLO MOM Gir cacenmtacanetasneacm aes 


Caphbalopodas, «.csj<avscs.eastacrse 


Coes eoeeeeessoseae 


Taste XITI.— Wenlock. 


Stratigraphical Geographical 
Distribution. Distribution, 
a 
at 
"5 
re O cS cn ; . rd 
rb) 2 2 rm f wo nD S| 
“2 oe} De eee |e 
=| a 3, aoe nee ee © 
Sas Se) ee Piss ton Ge 
S i= — —s Ff = ts S 
S| Se) ese] Swe ps 
rH q fe) @). |} > bs S ) Oo 
eS o = [es a) i) iS 5 
HA Fi4liale ia 
2 1 3 1 1 1 
2 1 3 1 1 1 
5) 1 3 3 3 
23 : r 0 8 15 16 5 
2 2 16 25 7 10 8 
5 3 26 76 11 1&8 sxe 11 
1 2 28 1 4 
* 1 2 65 1 5 00 
1 3 13 4 4 
sf 1 3 34 4 6 © 
6 3 14 2.7 5 9 s 
8 5 2:5 74 11 14 3 
3 10 2 
4 24 eee 2 e 
10 14 17 21 9 12 11 
15 19 06 96 23 39 20 
1 7 3 15 8 11 ? 
2 10 5 43 14 26 
1 4 4 9 E 7 1 
1 5 8 24 8 al te " 1 
l 3 1 1 1 
ee 1 ee 3 1 i « 1 
1 1 2 1 1 
Mins 3 3 5 4 3 9 
3 3 5 4 3 1 
vi 4 26 12 13 3 
26 39 67 1G1l R47 68 3 29 
54 57 137 473 98 160 16 45 


* In the column headed Wenlock beds I omit the Rhabdophora; they appear in the localities, 


Pass to 
Lower Aymestry Upper 
Ludlow. Limestone. Ludlow. 
3 2 2 
3 2 2 
1 
2 
9 ee 4 
16 8 7 
4 1 2 
6 1 2 
3 3 4 
4 3 6 
13 5 8 
18 6 10 
2 1 
2 1 
12 10 iil 
32 20 17 
13 3 9 
21 3 24 
6 4 4 
9 6 6 
2 
Be Uh 2 
2 2 
Pee ne {he tens. 2 
TPs 2 3 
11 2 3 
71 38 51 
126 562 817 


170 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Luptow. 


392 species, representing 137 genera, is, as nearly as can be ascer- 
tained, the numerical value of the Ludlow fauna. No true Ludlow 
rocks are certainly known to occur in North Wales, excepting 
perhaps in a small part of Montgomeryshire, south-east of Newtown. 
The Ludlow formation is zoologically allied to the Wenlock through 
129 species. The chief classes, or those most represented numeri- 
cally, are the Actinozoa with 12 genera and 17 species, the Echino- 
dermata with 12 genera and 21 species, the Crustacea with 29 genera 
and 97 species, the Brachiopoda with 13 genera and 48 species, the 
Lamellibranchiata with 17 genera and 71 species, the Gasteropoda 
with 9 genera and 33 species, the Cephalopoda 6 genera and 37 species, 
and lastly the Vertebrata, which first appear through the Fishes, 
represented by 10 genera and 14 species, omitting the genus Onchus, 
the remains of which may be “telsons” or tail-spines of Crus- 
tacea. The comparatively small fauna of the Aymestry Lime- 
stone (53 genera and 84 species) is, although local, a remarkable 
one, and clearly shows the nature of the sea-bed and sedimentary 
material in relation to the habits of life of the species represented. 
Mostly it is a recurrent fauna from the Wenloek, 57 species being 
common to the two, these 57 representing 38 genera, clearly showing 
the rarity and paucity of species illustrating the genera, and the 
lenticular or accidental nature of the Aymestry Limestone between 
the Lower and Upper Ludlow beds. The classes that appear to be 
most fully represented are the Coelenterata (Actinozoa), the Crusta- 
cea, Brachicpoda, Gasteropoda, and Cephalopoda. No fish-remains 
are known, although the two Ludlow series contain them*. The 
feeble representation of the Ludlow rocks in North Wales, as before 
stated (only 19 species), by comparison with Westmoreland (127) 
calls for critical examination into the physical rather than the pale- 
ontological aspect of the question; and this is still more prominently 
seen through the still larger fauna in South Wales and Shropshire, 
the former yielding 163 species, the latter 187; and further through 
the intermediate areas termed Herefordshire (where 114 species are 
known) and Worcestershire (121). Could we but see and examine 
the continuity of the old Wenlock and Ludlow sea-bed and its 
deposits between Denbighshire, Westmoreland, and the south of 
Scotland, now under the St.-George’s Channel (or denuded away), 
we should doubtless find traces of the Lower Wenlock and Ludlow 
rocks, uniting the two now disunited areas. We cannot doubt that 
this continuity once existed, though probably the rocks of the two 
areas were deposited under very different and local conditions. 
Petrologically they essentially differ ; but paleeontologically or speci- 
fically in certain groups the agreement is close and decisive, omit- 
ting the largely represented Annulose or Crustacean orders Meros- 


* Mr. Salter included the Aymestry Limestone with the Lower Ludlow, as 
being a calcareous condition of it. Mr. Lightbody was always impressed with 
the fact that the calcareous nature of the beds above the Aymestry Binatone 
influenced the fossils to a marked degree. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. I71I 


tomata and Phyllopoda in the Ludlow rocks (45 species), which 
stand alone as a peculiar and local group; their presence there- 
fore need not enter into the calculation*. The great extension of 
the Wenlock promontory to Wellington, running parallel to the 
Dudley beds, and the overlap and cover of the Triassic rocks to the 
north on to the Mersey, Liverpool Bay, Lancaster and Morecambe 
Bays, go far to lead us to believe that the missing evidence of 
the agreement between the Ludlow of the two areas of Wales and 
Westmoreland, as well as the unsatisfactory correlation of the Den- 
bighshire Grits and Tarannon Shales with the rocks of the Lake 
country, can be accounted for ; for it must be admitted that much has 
yet to be done with these Lower Wenlock rocks of Denbighshire, 
Westmoreland, &c. The Kendal group (or the Ludlow rocks) and 
the Ireleth Slates, which are the equivalents of the Wenlock of the 
south, can certainly be correlated paleontologically, dissimilar as they 
appear physically. The Scotch uppermost Silurians are exhibited 
only in three localities; to the south of Kirkcudbright and in the 
Pentland Hills both the Wenlock and Ludlow groups occur. The 
American series termed the Lower Helderberg group are the true 
equivalents of our Lower and Upper Ludlow, and contain a number 
of species in common. ‘The exact equivalent of the Oriskany Sand- 
stones may be our lowest Devonian. In North America their place 
is either at the top of the Upper Ludlow or between that and the 
lowest Devonian rocks, or ‘‘Corniferous beds” of the American 
geologists. I now attempt to analyze the extensive fauna of the 
Ludlow group, accepting the triple division unconditionally ; or, in 
other words, retaining the Aymestry Limestone as a distinct subfor- 
mation, not attached either to the Lower or the Upper Ludlow; its 
accidental position carries with it certain fossils having an important 
bearing upon physical geology. 

Prantra.—For the first time above the Wenlock we meet with what 
may be termed plants proper; yet the two genera Chondrites and 
Fucowdes are doubtful. 5 genera and 5 species are known; 4 
species occur in the Upper Ludlow, and 3 of the same in the 
passage-beds between the Upper Ludlow and Lower Old Red Sand- 
stone ; none are known in the Lower Ludlow or the Aymestry beds ; 
nowhere in the Ludlow of either South or North Wales, Westmore- 
Jand, or Ireland are plant-remains known. Actinophyllum plicatum 
occurs in Shropshire and Herefordshire; Chondrites verisiemilis 1s an 
Upper Ludlow form in Scotland. The seed-like bodies of doubtful 
affinities (Pachytheca spherica) occur in the tilestones and bone-bed 
of the uppermost Ludlow of Shropshire, Worcestershire, and Here- 
fordshire. Actinophyllum and Pachytheca here first occur, and do 
not pass to the higher formations. The passage-beds contain Acti- 
nophyllum plicatum, Chondrites verisimilis, Pachytheca spherica, 
and Pachysporangiwm pilula. 15 species of Crustacea, 19 species 
of Lamellibranchiata, 6 species of Gasteropoda, 4 species of Hetero- 
poda, and 6 species of Fish, or nearly 70 species, illustrate 10 classes 


* Phyllopoda (Ceratiocaris 14 species), Merostomata (Hurypterus 10 species, 
Hemiaspis 6, Pterygotus 9, Slimonia 3, and Stylonurus 3). 


172 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


in this borderland between the Upper Ludlow and the Old Red 
Sandstone. 

Prorozoa.—A series of sponges (Calcispongie) numbering 7 genera 
and 10 species occur in the Ludlow rocks, 3 of which are also 
Wenlock. 6 genera and 8 of the species belong to the Upper 
Ludlow; Amphispongia, Favospongia, and Tetragonis are new 
forms, and appear for the first time, and, with Mavospongia Ruthven, 
Ischadites Kenigr, and 4 species of Spongariwm, die out in the 
Upper Ludlow. 3 genera and 5 of the species occur in the Ludlow 
beds of Westmoreland; they are Favospongia Ruthven, Spongarium 
equstriatum, S.imterlineatum, S. interruptum, and Tetragonis Danbyt. 
The conformable Old Red Sandstone in the Ludlow area, or anywhere 
along the extended strike and junction of the two formations, con- 
tains nothing in common with the Upper Ludlow save a few fish 
and crustacean remains, all ceasing with the Downton beds and 
tilestones. The Eurypterids (Sty Wlowan us megalops and Hurypterus 
pygmeus) pass to the lower beds of the Old Red Sandstone; but 
through all the Old Red of South Wales (or the Silurian area) or 
Gloucestershire or Somersetshire, from its base to its junction with 
the Lower Carboniferous, nothing has occurred save fragments of 
Cephalaspis and Holoptychius, no other class in the animal kingdom 
being represented. 

Hyprozoa.—Only 1 genus and 8 species of this class seem left to 
bring to a close the history of the Graptolite group of the Hydrozoa. 
Monograptus has 8 species, which are all Lower Ludlow and occur 
in the Ludlow area. In no region of the globe where the Silurian 
rocks have been deposited have any species of this class trans- 
eressed or passed into higher formations. About 210 species belong- 
ing to 30 genera have lived during the Silurian epoch, ranging from 
the Arenig with 42 species, the Llandeilo 44, Caradoc 38, Lower 
Llandovery 50, Upper Llandovery 12, Wenlock 23, to the Ludlow 
with 8, their maximum development being at the commencement 
of the Lower Silurian of certain systematists. They came into 
existence with unexampled prodigality and suddenness, but gradu- 
ally died out as conditions grew less and less favourable for their 
sustenance and development. America, Bohemia, Scandinavia, 
Australia, and Britain have all largely possessed the same hydrozoal 
fauna, illustrating the same or similar deposits. Whether homo- 
taxially or not, most of the same genera have occurred in rocks 
of the same age as those of the british Islands, and always under 
the same physical conditions. The labours and researches of Hall, 
Barrande, Geinitz, Linnarsson, Nicholson, Lapworth, Hopkinson, 
Carruthers, and M‘Coy in this field of zoological history will ever 
reflect honour on their memory. To Mr. Lapworth graptologists 
owe much; no other author has given such attention to the Rhab- 
dophora, whether we regard his specific work or that devoted to 
their history and geological distribution through time, and his phi- 
losophical views relative to their affinities, structure, and systematic 
classification. 

Actinozoa.—Not a single species out of the 17 occurring in the 


ANNIVERSARY ADDRESS OF THE PRESIDENT. DRS 


Ludlow beds strictly belongs to them or is confined to the Ludlow ; 
all have previously occurred in the Wenlock. 17 species are Lower 
Ludlow; 8 of the same occur in the Aymestry Limestone, and 
7 in the Upper Ludlow, chiefiy the Favosites (5 species) ; 13 species 
occur in Shropshire, 6 in Westmoreland, and 12 in Worcestershire. 

Monticulipora papillata is the only Scotch species known. 6 species 
occur in Ireland (Fwosites asper, F. cristatus, F. fibrosus, Alveolites 

Labechu, Cr yathophyllum truncatum, and Monticulipora papillata) 
only 1 species is known in North Wales (Heliolites tubwlatus) ; and. 
6 genera and 9 species in South Wales. ‘The accompanying Table, 
showing their geographical distribution, will be clearer than descrip- 
tion, where the species are so irregularly distributed. 


oO . 
3 eeliccs 
Ri es see ob ve 
o © e oS “3 3S 
Cet] Se a il 
O + S 5 
Ludlow Species. Peal ete Hee eral Et resem ee 
le eaten = | ts 
ete ara SS ath By eles 
as) = (e) S © on) Gs | & 
S B = S a On| Wasa eees 
S) Sees) is = So | = 
aD |) 24 |p| Eo es ca | 
Alveolites Labechii .................. % % % % * 
Coomites mbertextus .c.2..<.5 .csecss. * 
Cyathaxonia siluriensis ............ ea al wae 
Cyathophyllum truncatum ......... “3 % ae * 
Favosites GDSTTE “yscqssoandodnosdaroce % % * Benalla % 
cristatus SE TaN ota eae baa. || sc00 * ae * x 
Bee =A DNOSUG CS AS ticle osu -wetdc dev doek % x * % % 
—— gothlandicus ....... pactasaets * x | * | x 
Horbesi. 
Fistulipora decipiens. 
Halysites catenularius............... See ae le % 
Heliolites interstinctus ............ Sean |e 0) |e ¥ % x 
fubulatus.....s..:. EC or ne AOA ae asst * * % 
Monticulipora papillata ...........- Stay eeu kere Ana Deen fe IE gh 9G- hea 
Ompbyma turbinata ..............- Ree ee Sea aes | aE * 
Pabraley OYA. ......+csceey ese Repth arr Ser diese malts | Fae 
Syringopora bifureata.......... oobl|.cok: |leodoin lesion lle we 
Be MPCUISE waht vssiaeshiewies sacievee * % * 
9 US Aa 76 6 1 6 


EcurnopERMATA.—This group in the Ludlow comprises 12 genera 
and 21 species. There are 3 genera and 4 species of Crinoidea 
(Actinocrinus pulcher, Ichthyocrinus pyriformis, I. M‘Coyanus, and 
Taxocrinus @Orbignyt). Of the remaining 17 species 4 are Cys- 
tideans (Hchinocystites pomum, H. uva, Pseudocrinites maguificus, 
and P. quadrifasaatus), all Lower Ludlow; and 13 Asteroidea. 
2 species of Palwaster (P. Ruthvent and P. hirudo) and Pale- 
asterina primeva, with Protaster Sedgwicku, occur in the Upper 
Ludlow of Kendal, Westmoreland. 4 Palwocome (P. Colvini, P. 
cygnipes, P. Marston, P. vermiformis) are all Lower Ludlow from 
the Leintwardine beds near Ludlow. Palwodiscus ferox, Protaster 
leptosoma, P. Miltom, and P. Sedgwicku are from the same horizon 
and locality, as well as Rhophalocoma pyrotechiica—Shropshire thus 


174 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


yielding 8 species of Lower Ludlow starfishes and Westmoreland 
4 Upper Ludlow. The Lower Ludlow Echinodermata therefore 
number 11 genera and 19 species, the Upper Ludlow 6 genera and 
7 species. None occur in Herefordshire, Scotland, or Ireland. 4 
genera and 7 species (3 Crinoidea, 1 Cystidean, and 3 Asteroidea) 
came from the Wenlock beds. In the Ludlow promontory and all 
along the line that marks the junction of the Old Red Sandstone 
with the Upper Ludlow all Silurian life ceased. The slow elevation 
of that region caused the Silurian Annuloid fauna to dwindle and 
pass away almost entirely. No vestige of it again appeared in the 
true Old Red area anywhere; and the want of evidence to show 
what rocks may be below the Lower Devonian of the Devonshire 
and Somersetshire areas leaves the question of stratigraphical or 
zoological continuity one of doubt and in the region of the unknown. 
Ireland or North Devon may yet solve the problem, for it is here 
we should expect it. The new Ludlow genera were Hchinocystites, 
Paleasterina, and Tetragonis, none of which leaves any successors 
behind, none being known either in the Devonian or Carboniferous 
rocks. 

ANNELIDA.—The Annelida are a singular group in the Ludlow 
rocks; 9 genera and 17 species have been obtained from the several 
horizons (4), and their geographical distribution is wide also. 15 
species occur in the Upper Ludlow; the characteristic Cornulites 
serpularvus is the most abundant, occurring at all 4 horizons, and 
in all localities except Shropshire and Ireland. The Lower Ludlow 
has yielded Cornulites serpularius, Serpulites dispar, S. longissimus, 
Spirorbis Lewisv, and Trachyderma squamosa. The Aymestry Lime- 
stone contains 4 species, all of which are also in the Lower Ludlow. 
4 of the 17 species are also Wenlock. 3 species of Afnonites and 
3 species of Arabellites, belonging to the order Polycheta, are 
determined from jaw-remains by Mr. G. J. Hinde*; they are Upper 
Ludlow, and help to swell the Annelide fauna. 3 species of the 
order Tubicola occur in the Passage-beds. 9 species range through 
South Wales, North Wales has 4, Westmoreland 6, Shropshire 5, 
Worcestershire 7, Herefordshire 4, and Scotland 3; but none, so far 
as I can ascertain, occur in Ireland. 

Crustacra.—The two great orders of the class Crustacea in the 
Ludlow rocks are the Merostomata and the Phyllopoda. Of the 
former we know 32 species—Lurypterus 10, Pterygotus 9, Himan- 
topterus 1, Slimona 3, Stylonurus 3, and Hemiaspis 6. Of the 
latter, through Ceratiocaris and Dictyocaris, we have 16 species ; and 
there are possibly one Amphipod (Necrogammarus Salweyr) and a 
Peecilopod (Neolimulus falcatus, Woodw.). These 32 species of 
Merostomata and Peecilopoda swell up the Crustacean fauna at the 
expense of the Trilobita, which number only 10 genera and 20 
species, 2 genera only of the 10 (Homalonotus and Phacops) passing 
to or occurring in the Devonian rocks. The Ostracoda number 
5 genera and only 13 species. The Crustacean remains termed Asta- 
coderma by Dr. Harley, number 14 so-called species; all are in the 

* Quart. Journ. Geol. Soc. vol. xxxvi. pp. 368-376. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 175 


Lower Ludlow. The whole Crustacean fauna, therefore, of the 
Ludlow rocks is 29 genera and 97 species. The Lower Ludlow 
contains 18 genera and 47 species, the Aymestry Limestone 9 genera. 
and 12 species, and the Upper Ludlow 23 genera and 71 species. 
The Passage-beds into the Old Red Sandstone have yielded 9 genera 
and 15 species, 4 being Ostracoda, 1 Phyllopod (Ceratiocaris decorus), 
and 10 Merostomata (Eurypteride). No Trilobite occurs in the 
Passage-beds. This Passage-bed Crustacean fauna is what we 
should have anticipated ; the Brachiopoda and Mollusca proper show 
the same indications of a change from a deeper to a shallow coast- 
line, and a fauna struggling to maintain life against new and adverse 
conditions. Shropshire through the Ludlow area possesses 18 genera 
and 46 species, Worcestershire 17 genera and 25 species, Hereford- 
shire 14 genera and 26 species, Westmoreland 8 genera and 15 
species. Scotland, chiefly through the Phyllopoda and Merosto- 
mata, has 12 genera and 22 species. Ireland only possesses 2 
(Calymene Blumenbachu and Phacops caudatus). This deficiency of 
the Crustacea in Ireland, as indeed of all the classes except the 
Brachiopoda, bears out the fact of the smallness of the Ludlow 
fauna in Ireland. I am obliged to carefully detail the number of 
genera and species that range through and enter into the distri- 
bution of the Crustacean fauna on account of its peculiarities. 
13 new genera made their appearance during the Ludlow period. 
4 Ostracoda, Cypridina, Entonus, Kirkbya, and Moorea ; 4 Merosto- 
mata (Hurypterida), Himantopterus, Slimona, Stylonurus, and 
Parka; Xiphosura 1, Neolumulus; Phyllopoda 1, Dictyocaris ; Am- 
phipoda 1 ?, Necrogammarus ; Crustacean remains, Astacoderma (14 
species). 3 genera (Stylonurus, Cypridina, and Parka) occur in 
the Old Red Sandstone, but not in connected areas. 16 genera and 
26 species are common to the Wenlock and whole of the Ludlow, thus 
reducing the true Ludlow Crustacean fauna to 71 species instead of 97. 

Bryozoa.—Heteropora crassa, Cerropora sulcata, and Ptilodictya 
lanceolata are the only 3 Ludlow Bryozoa. The first and last are 
Wenlock also. They occur in the Lower Ludlow and Aymestry 
Timestone of Worcestershire and Shropshire. Ceriopora sulcata 
occurs in the Ludlow of Ireland. No species occurs in the Upper 
Ludlow or Passage-beds, either in South or North Wales, West- 
moreland, or Scotland. The nature of the Ludlow shales and mud- 
stones doubtless was the cause why the Actinozoa, Hydrozoa, Kchi- 
nodermata (Crinoidea &c.), and Bryozoa, all mostly dwellers in clear 
water, are so sparingly developed and distributed through the Ludlow 
rocks, the Aymestry Limestone being of little palaeontological value 
owing to its lenticular disposition, mode of occurrence, and uncertain 
continuity. 

Bracuropopa.—Although by no means a small fauna, the Ludlow 
Brachiopoda numbering only 48 species and 13 genera are less 
than half as numerous as those of the underlying Wenlock rocks. 
41 of these 48 had hved in the Wenlock seas and passed to the 
Ludlow ; therefore only 7 species are Ludlow proper; and only 3 
species passed to the Devonian—Altrypa reticularis, Strophomena 


176 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


rhombordalis, and Lingula cornea. The last named in the Ludlow 
country passes into the lowest beds of the Old Red Sandstone. The 
others named are “ Devonian” of the North and South Devonshire 
areas, and not known in the Old Red proper. In the Passage-beds 
into the Old Red of Herefordshire, Shropshire, and Worcestershire 
some 5 or 6 species occur; they are Chonetes striatellus (latus), Dis- 
cma rugata, Lingula cornea, Lingula nanima, and Strophomena 
rhomboidalis. They are associated in places with a few Asiphonida 
of shallow-water habits, Pterinea, Modiolopsis, Orthonota, &e. ‘The 
Lower Ludlow has yielded 13 genera and 38 species, 33 of which 
are Wenlock. 12 genera and 25 species occur in the Aymestry 
Limestone, and 11 genera and 24 svecies in the Upper Ludlow; but 
no genus is peculiar to either the Aymestry or Upper Ludlow beds, 
the whole being represented in the Lower zone. 

‘The only species in the Ludlow not Wenlock are Lingula cornea, 
L, lata, Orthis canahculata, O. lunata, Rhynchonella pentlandica, and 
Strephomena ornatella. In other words, only these 6 species of 
Brachiopoda are really Ludlow forms. No species is known in 
North Wales, but 28 have been collected in South Wales. Here- 
fordshire, Worcestershire, Shropshire, and Scotland have each yielded 
11 genera, and respectively 23, 27, 30, and 15 species; Westmore- 
land has 8 genera and 14 species ; Ireland 9 genera and 18 species. 

LAMELLIBRANCHIATA.—Both the Lower and Upper Ludlow pos- 
sess a large Molluscan fauna; the Lower 14 genera and 54 species, 
and the Upper 15 genera and 56 species; the intermediate Ay- 
mestry Limestone only 4 genera and 7 species—Pterinea lwans, 
P. retrofleca, P. Sowerbyi, Cardiola striata, Grammysia cingulata, 
Orthonota rotundata, and O. semisulcata. The chief genera are Pte- 
rincea with 12 species, Anodontopsis 7, Orthonota 16, Modiolopsis 5, 
Ctenodonta 4, Cucullea 4, and Grammysia 4 species. The Passage- 
beds contain 9 genera and 19 species, yet no single species occurs in 
the Lower Old Red Sandstone; with the change of lithological 
characters all life seems to cease also, or no record is left. For the 
life-history of those rocks, equivalent in time to those of the Old Red 
Sandstone in the classical Silurian area, we have to consult the De- 
vonian series of North and South Devon and Cornwall, where a 
fauna unsurpassed in perfection, if not in magnitude, in all the in- 
vertebrate classes exists—a group made famous in British geological 
history through the labours and researches of Sedgwick, Murchison, 
De La Beche, and Phillips. Westmoreland and Cumberland, through 
the researches of Sedgwick and M‘Coy, Harkness and Nicholson, in 
the Kendal area, exhibit a large Ludlow Lamellibranchiate fauna ; 
12 genera and 37 species are known. Shropshire has 9 genera and 
16 species, Worcestershire 8 genera and 17 species, Herefordshire 6 
genera and 12 species, Scotland 6 genera and 13 species, and Ireland 
only 2 genera and 3 species. North Wales hitherto has yielded only 
1 genus and 4 species; 13 genera and 21 species ranged from the 
Wenlock into the Ludlow, thus altering the numerical value of the 
Ludlow bivalves proper to 50 species, none of which passed the 
Passage-beds between the Upper Ludlow and Old Red Sandstone. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. Maa 


Extensive patches of Upper Silurian rocks on the eastern side of the 
Silurian area and border of the Old Red, such as Woolhope, Usk, 
the long strike of the Malverns on to May Hill, thence across the 
Severn at Purton Passage to Tortworth, indicate great physical 
changes in the borderland between the fossiliferous Upper Silurian 
and the unfossiliferous Old Red; and whatever change brought in 
the marine Devonian of North Devon, south of the Mendip axis and 
on the same strike, and all the South-Wales Old Red beneath the 
South-Wales coal-field to the Bristol Channel, certainly must ac- 
count for the loss of all the Ludlow species ; that the Devonian area 
was one of long and continuous depression south of the latitude of 
the Mendips there cannot be any doubt, and probably the mass of 
Old Red Sandstone occupying Caermarthenshire, Monmouthshire, 
Breconshire, Herefordshire, and Worcestershire was being at the 
same time slowly elevated. 

GasTEROPODA.—9 genera (six of which had appeared in the 
Wenlock) and 33 species, 11 of which were also Wenlock, range 
through the Ludlow rocks; the Lower Ludlow has yielded 8 genera 
and 15 species, the Aymestry Limestone 5 genera and 11 species, 
and the Upper Ludlow 9 genera and 21 species. 3 genera and 
6 species occur in the Passage-beds into the Old Red Sandstone ; 
they are Holopella conica, H. gregaria, Holopea obsoleta, Mur- 
chisonia torquata, Platyschisma helicites, and P. Williamsi. They, 
however, cease here, none living into the red beds of the Old Red 
Sandstone. North Wales has yielded only 2 species, Holopella 
gracilior and H. gregaria ; Scotland only 4 species, Acroculia anti- 
quata, Huomphalus funatus, Platyschisma semulans, and P. helicites ; 
Ireland has only registered 1 species, Huomphalus alatus; Here- 
fordshire 7 genera and 11 species, Worcestershire 6 genera and 9 
species, Shropshire 7 genera and 14 species, and Westmoreland 6 
genera and 14 species. 6 genera and 13 of the 33 species come up 
from the Wenlock and range through the Ludlow rocks; they are 
Acroculia haliotis, Cyclonema coralli, C. octavia, Kuomphalus alatus, 
E. carinatus, E. funatus, H. rugosus, Holopella gracilior, H. obsoleta, 
Loxonema elegans, L. sinuosa, Murchisonia Lloydi, and Mu. ar Beclaties. 
9 species only are Lower Ludlow. All become extinct in the Tile- 
stones and Passage-beds. 

Preroropa.—Conularia subtilis and C. Sowerbyi, with Theca 
Forbesi, are all the species (3) illustrating this pelagic group. ‘The 
two last named are Upper Ludlow. Neither the Aymestry Lime- 
stone, Passage-beds, North Wales, or Worcestershire yield either 
species, nor ‘do we know of any species in Ireland. C. Sowerbya 
occurs in Scotland; all 3 species in Westmoreland and 2 (Theca 
Forbes and Conularia Sowerby yt) in Shropshire. These last are also 
Wenlock species. 

Hereroropa.— Bellerophon (6 species) and Heculiomphalus levis 
constitute the Ludlow stock of Heteropoda. Heculiomphalus levis, 
Bellerophon expansus and B. dilatatus occur both in the Lower and 
Upper Ludlow. 3B. Murchison, B. obtectus, and B. trilobatus are 
Upper Ludlow only ; 4 species occur in the Passage-beds, B. cari- 


178 PROCEHDINGS OF THE GEOLOGICAL SOCIETY. 


natus, B. exepansus, B. Murchison, and B. trilobatus; 6 species 
occur in the Upper Ludlow of Westmoreland; Shropshire and 
Worcestershire possess 3 of the same species. None are known in the 
Ludlow rocks of North Wales, Herefordshire, Scotland, or Ireland. 
B. carinatus, B. trilobatus, and B. dilatatus are also Wenlock species. 

CrpHaLopopA.—No less than 24 species of Orthoceratites, 6 species 
of Phragmoceras, and 3 of Litutes, with other forms, make up the 6 
genera and 37 species known in the Ludlow. Ascoceras Barrandii 
constitutes a new generic type. The Lower Ludlow has yielded 
4 genera, Latuites (3 species), Orthoceras (15 species), Phragmo- 
ceras (6 species), and EHxosyphonites (2 species). 

The Aymestry Limestone has yielded 3 genera and 6 species— 
Tntuites giganteus, Orthoceras 4 species, and Phragmoceras ventricosum. 
The Upper Ludlow 5 genera and 22 species—Ascoceras Barrandi, 
Litwites giganteus, Orthoceras 17 species, Phragmoceras nautileum, 
and Tretoceras semipartitum. 

In the Passage-beds we have noted Orthoceras bullatum and Tre- 
toceras semipartitum. 16 species of the genus Orthoceras occur in 
Westmoreland, mostly Upper Ludlow, and no other genus has yet 
occurred there. Shropshire possesses 4 genera and 22 species, em- 
bracing most of the species in the Upper and Lower Ludlow divisions ; 
Herefordshire 5 genera and 21 species, Ascoceras Barrandu, Intwites 
3 species, Orthoceras 11, Phragmoceras 3, Exosiphonites 2. Worces- 
tershire has 10 species. Orthoceras Maclareni and O. subgregarium 
are the only Scotch forms known ; and O. subgregarium is the only 
Irish species. 11 species come from the Wenlock rocks to the Lower 
Ludlow, but 16 Wenlock species range through the Ludlow group ; 
Ascoceras is the only new genus; and none ranged above wmto erther 
Old Red Sandstone, Devonian, or Carboniferous rocks. 

Pisces.—No Vertebrata have yet been discovered in earlier rocks 
than the Lower Ludlow. Scaphaspis (Pteraspis) ludensis, Salt., 
is the first fish known, and the only species in the Lower Ludlow; 
none have yet been detected in the Aymestry Limestone, but every 
known Ludlow form occurs in the Upper Ludlow, and five of them 
in the Passage- or junction-beds, through which about 20 species of 
the Silurian fauna pass to the Lower Old Red Sandstone, 7 of which 
are Crustacea (Merostomata), 6 Fishes, and 2 Cephalopoda, &e. 

The Crustacea common to the Ludlow and Old Red, and all be- 
longing to one order, are Hurypterus abbreviatus, HL. acumimatus, and 
E. pygmeus, Pterygotus problematicus, Stylonurus Powriet, S. me- 
galops, and Parka decipiens. The fishes that connect the two for- 
mations, but only in the Silurian area along the frontier of the Old 
Red, are Auchenaspis Salteri, Cephalaspis Murchison, C. ornatus, 
Pteraspis Banksii, Scaphaspis ludensis, and Eukeraspis pustuliferus. 
All belong to the Ganoidei. 


DEVONIAN. 


Prant#.—We have no clue as to the region in which the 
Devonian Plants first appeared; and yet small as is the flora of 
the British Devonian as compared with that of North America it 


6éeT 


~t i) Ne We tt i) n 
x nO OM NY OM Fe oy YY gay PY FA Do we woo From W enlock. 


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180 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


shadowed forth the prolific flora of the Carboniferous epoch, which 
in the British Islands numbers 82 genera and 330 species; but the 
Devonian, which preceded it, has, up to the present time, yielded 
only 12 genera and 18 species. In America, however, about 95 
genera and 160 species have been described through the researches 
of Dr. Dawson, Lesquereux, Vanuxem, Hall, Hartt, &c., especially 
the first-named author, who has devoted a long life to the eluci- 
dation of the Devonian and Carboniferous flora of the American 
region. 7 of the 12 British Devonian genera occur in America, 
viz. Calanuites, Lepidodendron, Psilophyton, Sagenaria, Spheno- 
pteris, Stigmaria, and Trichomanites ; but our ill-preserved speci- 
mens scarcely admit of specific determination, in fact even their 
affinities are often difficult to make out; and the poverty of our 
flora is clearly seen from the few species (18) occurring to illustrate 
the 12 genera. On such slender evidence it is difficult to conceive 
from what area our Devonian and Old Red Sandstone flora was 
derived; whether migration took place from Central Europe or 
America. The facies appears to be American rather than European, 
this being the case with the Scotch and Irish floras, especially 
-through Psilophyton, Paleopteris, and Sagenaria. 

The Lower Devonian (Lower Old Red) has yielded 2 species, 
Lepidodendron nothum, Unger, and Psilophyton Dechenianum, Carr., 
with coniferous remains from Caithness and Wick in Scotland. The 
Middle Old Red contains Caulopteris Peachi, Salt. (=? Psilophyton 
robustus, Dawson), and the two mentioned as occurring in the Lower 
beds. The Upper Old Red species (12) approach closely to the Car- 
boniferous. 7 genera and 12 species range through the Upper divi- 
sion—Advantites 1 species, Calamites 1, Filicites 1, Sagenaria 5, 
Sphenopteris 2, Trichomanites 1, and Anorria 1. Thus only 18 
species occur in the widely extended and thickly developed deposits 
of the Old Red of Scotland, the Silurian Old Red, and the rocks 
of the Irish area. 

Prorozoa.—Scyphia turbinata, Spherospongia tessellata, and 5 
species of Stromatopora all occur in the Middle Devonian; Cawno- 
ora, Phillips, Sparsispongra, D’Orb., and Coscinopora of Goldfuss are 
probably synonyms of Stromatopora, The Middle Devonian lime- 
stones of Torquay and Newton Abbot abound in the ever-varying 
forms of these incrusting Protozoa. The Devonian forms differ 
essentially from the Wenlock and Ludlow species, the type in those 
rocks being S. striatella, whereas the characteristic species in the 
Middle Devonian are S. concentrica, Goldf., and S. placenta, Lonsd., 
or the Caunopora of both Phillips and Lonsdale. No Protozoa occur 
either in the Lower or Upper Devonian, being strictly confined to 
the massive limestones of South and North Devon. S. placenta, S. 
ramosa, and S. verticillata are strictly British. 

Actinozoa (Actinoidea, Dana; Coralliaria, M.-Kdw.).—Perhaps 
during no period in the physical history of the British Islands have 
we had such a remarkable assemblage of Actinozoa as that which 
so essentially and specifically characterizes the Middle Devonian 
rocks of South and North Devon. Out of the 24 known genera and 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 181 


52 species, no single form passes to the Carboniferous, and none are 
common to the Silurian rocks in any area; they stand alone, and 
are sufficient in themselves to maintain the integrity of the Devonian 
system. This is equally definite and distinct throughout the Kuro- 
pean or American areas, or wherever the Devonian rocks are deve- 
loped. 15 genera illustrate the Zoantharia rugosa and 8 the Z. tabu- 
lata. The genus Acervularia numbers 7 species, Alveolites 4, Cya- 
thophyllum 12, Favosites 5, Smithia 3, Endophyllum 2, and Petraia 2. 
The remaining 18 genera are only represented by 1 species each ; 20 
species are common to the rocks of North and South Devon, and 25 
occur on the continent (Rhenish Prussia, Belgium, and France) ; 
none pass to the Carboniferous rocks in any region. ‘The Lamelli- 
branchiata, Gasteropoda, and Cephalopoda are all of equal strati- 
graphical value. The Middle Devonian group contains all the 24 
genera and 48 of the 52 species. The Lower Devonian has hitherto 
only yielded 4 genera and 7 species; the latter are Alveolites sub- 
orbicularis, Cyathophyllum helianthoides, Petraia celtica, P. gigas, P. 
pleuriradialis, P. bina?, and Pleurodictyum problematicum ; and the 
Upper Devonian also 4 genera and 7 species, viz. Amplexus tortuosus, 
Cyathophyllum cespitosum, C.ceratites, Pistulipora eribrosa, Michelinia 
antiqua, Petrara celtica, and P. pleuriradialis. The researches of Dr. 
Nicholson upon the Devonian Corals of North America have thrown 
much light upon their history, habits, and mineralization ; and itis 
to be hoped that his labours will be embodied in a volume upon the 
Rugose Corals, equal in interest and value to his late contribution on 
the Tabulata. Doubtless the Paleeozoic Actinozoa of the British 
rocks now require critical revision, especially the Silurian and Car- 
boniferous groups. 

The Upper and Lower Devonian rocks are chiefly composed of 
slates, with here and there impure limestones. They therefore 
possess no coral fauna; whereas the highly developed masses of 
limestone around Torquay and Newton Abbot are simply Devonian 
coral reefs of great magnitude. In North Devon, between Lfra- 
combe and Linton, the limestones are lenticular, thin, and im- 
pure; yet more than one half of the entire known Devonian Acti- 
nozoa have occurred in them, and we have evidence of beds below 
at low-water level near Llfracombe yielding even finer specimens 
than at Torquay. They have to be searched for in North Devon 
and West Somerset ; patient working over that extensive area be- 
tween Barnstaple and the Foreland with unbiassed views and a 
knowledge of the fauna which exists altogether independent 
either of the Silurian or Carboniferous, would readily convince 
those who have never examined the county that the rocks of North 
Devon between Baggy and the Foreland have nothing whatever to do 
with theCarboniferous system. ‘The fossils alone, setting aside strati- 
graphical evidence and succession, determine the relation of this 
area to Belgium, the Rhine, and France; and their continuity under 
Somersetshire, Wiltshire, Berkshire, and Middlesex on to the continent 
is no longer matter of doubt or speculation, for the philosophical 


and far-seeing views and hypotheses of Mr. Godwin-Austen and Prof. 
VOL. XXXVIL. 0 


182 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Prestwich have been fully realized, through the determination of 
the Devonian and Silurian rocks in Middlesex and Hertfordshire, 
immediately below the Cretaceous series. 

EcHINopERMATA.—/ genera of Crinoidea, 2 Asteroidea, and 1 
Blastoid are all that are known of this class in the British Devo- 
nians. 80 genera have been described from the Devonian rocks of 
Europe and America. By comparison, therefore, the British fauna has 
scarcely any claim to recognition, and 5 of the 7 Crinoidea are also 
Carboniferous in Britain. It cannot be said, therefore, that the species 
in this division of the order are representative in Britain. The 10 
genera and 24 British species are of necessity unequally distributed. 
Actinocrinus tenwmstriatus, Phill., Cyathocrinus megastylus, Phill., 
and C. pinnatus, Goldf., are the only forms known in the Lower 
Devonian ; 6 genera and 12 species occur in the Middle Devonian, 
viz. Actinocrinus 1 species, Cupressocrinus 2, Cyathocrinus 3, Hexa- 
crinus 3, Platycrinus 2, and Taxocrinus 1 species. 

The Asteroidea all belong to the upper division, and are mostly 
from the North-Devon beds, which with them have also yielded 7 
Crinoidea. We have therefore 8 genera and 14 species in the Upper 
Devonian beds of North Devon—Protaster 2 species, Paleaster 2, 
Helianthaster 1, Pentremites 1, Adelocrinus 1, Actinocrinus 1, Cyatho- 
crinus 5, and Taxocrinus 1 species. Of the whole fauna (24 species) 
3 genera and 6 species pass to the Carboniferous; they are Actino- 
crinus triacontadactylus, Cyathocrinus ellipticus, C. geometricus, C. 
pinnatus, C. variabilis, and Pentremites ovalis. Three of these are 
European Devonian. 

AnnELIpa.—Serpula advena, Salt., occurs in the Upper Old Red 
Sandstone of Caldy Island, and Yentaculites annulatus?, Schloth., 
in the Middle Devonian of North Devon; the last named is abundant 
(in places) in the limestones near Ilfracombe. 

CrustaceA.—All the 4 orders, illustrated only by 20 genera and 
37 species, are represented in the British Devonians ; 45 genera and 
290 species, however, have been described from Bohemia, Germany, 
Spain, Asia, Africa, America, &c. 20 of the foreign genera and 200 
species are Trilobita ; the remaining 90 are Merostomata, Phyllopoda, 
and Ostracoda. As in the case of the Echinodermata our Crustacean 
fauna is any‘thing but representative, yet it has a characteristic 
facies that carries with it the conviction of distinctness. Only 6 
genera of Trilobita are known, viz. Bronteus (flabellifer), Cheirurus 
(articulatus), Harpes (macrocephalus), Homalonotus (elongatus), Pha- 
cops (Sspeciesincluding the subgenera 7rimerocephalusand Crypheus), 
and Phillipsia (Brongniarte); the doubtful genera have been relegated 
to their supposed places. 9 genera and 24 species of all the orders 
are Lower Devonian (6 genera are Trilobita), 5 genera and 6 species 
are Middle, and 7 genera and 9 species are Upper Devonian. 
Eurypterus (6 species), Stylonurus (6 species), and Pterygotus (4 
species) are all, with one or two exceptions, Lower Old Red Sand- 
stone types. None of these are known to occur in the typical Devonian 
area, being either Scotch or in the Silurian region. The singular 
genus Prearcturus (P. gigas, Woodw.) of the family Idoteide is 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 183 


from the Old Red of Herefordshire; this and Proricaris M‘Henrici 
of Baily form the only 2 new British genera introduced into the 
Devonian fauna. Phillipsia Brongniartii is the only crustacean that 
occurs in common in the Devonian and Carboniferous rocks ; it is the 
Asaphus obsoletus and A. granuliferus, Phill.; so that, small as the 
Devonian Crustacean fauna appears to be, it is nevertheless distinc- 
tive. A. Romer, Sandberger, Dalman, Brongniart, Richter, Minster, 
Beyrich, Steining, Roualt, &c. on the continent, with Salter, Phillips, 
Woodward, Jones, R. Etheridge, jun., and Baily in Britain, have all 
greatly enriched our knowledge of the Devonian Crustacea. 
Bryozoa.—The Tubuliporide through Ceriopora, the Retioporide 
through Henestella, Henntrypa, Polypora, Ptylopora, and Retepora, 
and the Hscharide through Glauconome, are represented through the 
Devonian rocks by the above 7 genera with 11 species; all the 
genera are equally Carboniferous, but only 4 species are common to 
both formations-—Ceriopora similis, Phill., Fenestella plebeia, M‘Coy, 
Glauconomne bipinnata, Phill., and Polyporalaxa,Sandb. The Lower 
Devonian possesses 2 species only, viz. Henestella antigua, also oc- 
curring in the Middle and Upper, and Retepora repisteria, which is 
also Middle Devonian. 6 of the 7 genera and 7 species are Middle 
Devonian—Polypora, through its representative species (P. lawa), 
being Upper and Carboniferous. 5 genera and 6 species are Upper 
Devonian ; they are both South- and North-Devon forms—Cerio- 
pora (Millepora) gracilis, Phill, Fenestella antiqua, Goldf., F. 
prisca, Goldf., #. plebera, M‘Coy, Glauconome bipinnata, Phill., 
and Polypora lava. The known Devonian Bryozoan fauna (Kuro- 
pean, American, and British) consists of 26 genera and 115 species ; 
of these we have only 7 genera and 11 species. These species 
range through North and South Devon and South Cornwall; they 
are rarely well preserved and always difficult to determine. Critical 
analyses of the species have been undertaken by Messrs. Shrubsole 
and Vines, who in time will revise the entire group. 
Bracuropopa.—With the exception of the fishes of the Old Red 
Sandstone (125 species) this is the largest group in the British 
Devonian rocks. We should expect this when we know that no 
less than 61 genera and over 1100 foreign species have passed 
through the hands of European, American, and British zoologists and 
palzontologists, and all have been described ; of these 1100 species 
only 116 are British; and of the 61 known genera we possess 26. 
Calceola (1), Davidsonia (3), Cyrima (A), Rensseleria (11), Camaro- 
phoria (1), Stringocephalus (1), and Uncites (1) are the genera new to 
Britain, none of which made their appearance in our area until the 
Middie periodof the Devonian deposits, the most marked and prolific 
of the three horizons. With the exception of Cyrtina, which exhibits 
_4 species, each of these genera is represented only by 1 species. 
Hall’s genus Rensseleria yields 10 as the total value of the genus ; 
but only 1 species occurs in Britain (Rensseleria stringiceps, Rom.) ; 
8 of the rest are American, and 1 species is Coblentzian. JI call 
attention to these 7 genera because they are essentially Devonian ; 
the remaining 19 appeared in the Silurian rocks, and lived on 
through the Carboniferous. 
02 


184 PROCEEDINGS OF THE GEOLOGICAL SOCINTY. 


The Lower Devonian yields 9 genera and 21 species—Athyris 1, 
Atrypa 3, Chonetes 3, Leptena 1, Or this 3, Rhynchonella 3, Spirifera 
3, Spur iferi ina 1, Sér eptorh ynchus 3. The Middle division contains 
23 genera out of the 26, and 80 of the 116 species. The Upper 
contains 14 genera and 37 species; of these 10 genera and 16 
species pass to the Carboniferous. It is essential to the history of 
the Devonian fauna that I name the genera—Athyris 2 species, Cho- 
neétes 1, Discina 1, Lingula 1, Productus 1, Rhynchonella 3, Spirifera 3, 
Streptorhynchus 1, Strophomena 1, Terebratula 2. The large genera 
are Lthynchonella 16 species, Spirifera 20, Streptorhynchus 6, Orthis 
6, Cyrtina 4, Productus 4, and Terebratula 4. Regarding the 
great discrepancy or smallness of this peculiar fauna as compared 
with that of the continent and America, we must have regard to the 
smallness of the area now exposed in England as compared with the 
original area occupied by the Devonian sea, the accumulations of 
which are now covered by the Secondary and Tertiary rocks of the 
west and east of England, the Devonian floor or old surface being 
hidden east of the Quantocks, North Devon, and Torquay. We have 
proof of a rich Upper Devonian fauna in the rocks under London 
and Turnford, at the depth of 1000 feet, and below the Cretaceous 
series ; between these two places and North Devon we can at pre- 
sent only surmise the plane they occupy. 

The volume of the Paleontographical Society’s publications by 
T. Davidson, Esq., F.R.S., &c., devoted to the British Devonian 
Brachiopoda, is worthy of the fame of its distinguished author; in it 
are described 116 species, the arguments for and against their 
genuineness being impartially and consummately reasoned out, 
figures of all the species and doubtful forms are given; and when 
we know the patience and skill required to adjust and discriminate so 
dismembered and small a group as the British Devonian Brachiopoda 
out of such a mass of material occurring in the European and Ame- 
rican fauna (1100 species), we may well be thankful that there are 
men who have the required leisure and knowledge and who devote 
their lives to one subject. Associated with Davidson in the Devo- 
nian fauna, the names of Von Buch, Schlotheim, Schnur, Dalman, 
Sandberger, Hall, Billings, Dall, Conrad, Romer, Vanuxem, Bar- 

rande, De Verneuil, Roualt, Phillips, Sowerby, King, and M‘Coy 
must be prominently noticed. 

LAMELLIBRANCHIATA.—The mass of the species of the class Conchi- 
fera occurring in the Upper Devonian, and especially abundant, 
are Asiphonida, through Avieulopecten, Pterincea, and the Mytilidee. 
20 genera and 39 species are known, and 29 species illustrating 11 
genera are in the Upper division ; yet of the whole bivalve fauna 
only 4 genera and 5 species pass to the Carboniferous series in 
North Devon, viz. Aviculopecten granosus, Sow., A. plicatus, Sow., 
Pterinea damnonensis, Sow., CuculleaGriffithu, Salt., and Curtonotus 
unio, Salt. The Lower Devonian is even poorer, only 3 genera and 
4 species occurring; these are Aviculopecten polytrichus, Phill., 
Pierinea anisota, Phill., P. spimosa, Phill., and Otenodonta Kratche, 
Rom.; the last mentioned is the only bivalve species actually con- 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 18 5 


fined to the Lower Devonian. ‘The Middle Devonian is represented 
by 13 genera exhibiting 23 species—<dviculopecten 2 species, Péeri- 
nea 6, Anodonta Jukes, Cledophorus ovatus, Corbula Hennahe, 
Cucullea Hardingu, C. trapezium, Ctenodonta lineata, Leptodomus 
constrictus, Megalodon 3, Modiola scalaris, Pleurorhynchus 2, Pul- 
lastra antiqua, and Sanguinolaria elliptica. Megalodon, Pleurorhyn- 
chus, Corbula, and Clecdophorus are generically and specifically essen- 
tially Middle Devonian. Cucullwa 6 species, Curtonotus 6, Cteno- 
donta 4, are the larger genera of the Upper Devonian beds of North 
Devon, south of the latitude of Baggy Point; but we have seen how 
few pass to the Carboniferous south of Barnstaple and Pilton, or along 
the strike of the two formations from Barnstaple to Braunton. The 
South-Petherwin and Land-lake beds in North Cornwall are Upper 
Devonian, and have yielded 65 species ; they cannot be affiliated with 
the Carboniferous, the Clymenie and species of Orthoceratide for- 
bidding it. 

Any comparison of the insignificant British fauna with the splendid 
series of Devonian species in Kurope and America is useless : no fewer 
than 90 genera and 900 species of Lamellibranchiata have been de- 
seribed ; 260 are Monomyarian and 640 are Dimyarian species. 

GasrEropopa.—13 British genera and 45 species constitute the 
entire list of the Odontophora. No species occurs in the Lower Devo- 
mian. ‘The Middle, as in the case of all the other classes, contains the 
largest number of species (36) and 12 of the 13 genera—Acrocula 
3 species, Huomphalus 5, Lowonema 5, Macrocheilus 6, Murchisona 4, 
Nerita 1, Pleurotomaria 6, Scoliostoma 1, Trochus 1, Turbo 2, and 
Vermetus 2 species. The Upper Devonian species are 14, and occur 
mostly in the Pilton, Brushford, and Petherwin areas; they represent 
7 genera—Aecroculia 1, Huomphalus 1, Loxonema 4, Macrocheilus 1, 
Murchisona 1, Natica 2, and Pleurotomaria 4. 5 species pass to the 
Carboniferous—Acroculia vetusta, Lovonema rugiferum, L. tumidum, 
Murchisoma angulata, and M. spinosa. 

The South-Devon area has yielded 37 of the 45 Middle Devonian 
species, North Devon only.5, viz. Huomphalus serpens, E. radiatus, 
Acroculia vetusta, Macrochelus brevis, and Natica meridionalis, 
and 11 Upper Devonian ; the Petherwin beds in North Cornwall 
contain 7 Gasteropods, all Upper Devonian. Complete comparison 
of the two areas will be made at the end of the Devonian fauna. 

Preropopa.—No Conularia or Theew has appeared in the Devonian 
rocks of either South or North Devon. 30 species are known in 
the Devonian rocks of Germany and America. 

Hureropopa.—The order Nucleobranchiata through the Atlantidee 
is represented in all three divisions of the Devonian rocks by Belle- 
rophon (5 species) and Porcellia (3). Bellerophon bisulcatus occurs in 
the Lower and Upper divisions; B. striatus, Lower and Middle; 
B. hiulcus, B. subglobatus, and B. Urii in the Upper; B. hiulcus and 
B. Urii appear also in the Carboniferous rocks. B. subglobatus is also a 
Coomhola species (Irish). Porcellia Woodwardi, Sow., and P. striata, 
Phill., occur in the Middle Devonian of South Devon; but P. Symondsiz 
has hitherto only been found in the Upper division, or Pilton and Barn- 


186 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


staple beds south of Baggy. Thus these 2 genera and 8 species have 
a wide range, due to their pelagic habit; and regarding B. Uri, B. 
hiulcus, and B. subglobatus as Carboniferous, 3 occur in the Lower, 
3 in the Middle, and 5 in the Upper Devonian. 5 genera and 77 
species are Continental and American. 

CrpHatopopa.—Clymenia, Cyrtoceras, Goniatites, Nautilus, Ortho- 
ceras, and Poterioceras are the 6 British Devonian genera with 60 
species. Other foreign genera and 500 species are noma We can 
hardly attempt comparison through such an extensive and widely 
disseminated group. (The genus Goniaiites is illustrated by 168 
species, Clymenia 50, and Orihoceras 130; but Bactrites with 9 
species, Cyrtoceras 60, Phragmoceras 12, and Trochoceras 6 species, 
are not known in the British Devonians, besides many other smaller 
genera.) Strange as it may appear, only one species occurs in the 
Lower Devonian, Cyrioceras bdellahies, Stutchb.; yet this genus is 
represented by 12 Middle and 2 Upper Devonian species. I am 
disposed to believe that we have not in Britain any Lower Devonian 
form at all, this single species from one locality being doubtful. 
The 11 other forms are all Middle Devonian. 

Clymenia, the essential genus in the Devonian rocks of Britain 
(and elsewhere), numbers 11 species, and all occur in the Upper 
Devonian of Petherwin in North Cornwall. This genus is repre- 
sented in Europe by 50 species, and, in the same stratigraphical 
position, in the Upper division of the Devonian serics. America 
yields scarcely any Clymenie; and no Clymenia occurs in the Devo- 
nian rocks of North Devon, either Middle or Upper. 

Gonratites—In the Upper Devonian of Petherwin this genus is 
represented by G'. biferus, G. linearis, G. subsuleatus, G. striatus, 
and G. vinctus; and 12 species occur in the Middle Devonian of 
Torquay ; only 2 forms occur in the Upper beds of North Devon, 
G. vinctus and G. spirorbis; 4 of the Devonian species occur in the 
Carboniferous rocks, G. carbonarius, G. cxcavatus, G. serpentinus, 
and G. subsuleatus? As compared with the Carboniferous Gonia- 
tites, the Devonian group is of slight importance ; in Britain alone 
we have 72 Carboniferous species, and 160 Devonian species have 
been described from European rocks, and only 6 or 8 from America. 

Cyrtoceras—— With one exception the 15 species of this genus 
occur only in the Middle Devonian of Torquay. The single departure 
from this is Cyrtoceras rusticum, the only Upper Devonian form in 
the Petherwin beds ; this species is probably the C. arcuatum of Stein- 
meyer. 60 species occur in the Devonian rocks of Central Europe ; 
and, as in the two genera before noticed, few are American. 

Orthocer as.—No species of Orthoccras has yet occurred in the 
Lower Devonian beds, either of North or South Devon. The Middle 
group at Torquay has yielded 8 species, and the Upper Devonian of 
North Devon 13 ; and 4 of these North-Devyon species pass to the Car- 
boniferous south of Pilton and Barnstaple, viz. O. cinctum, O. lineola- 
tum, O. striatum, and O. undulatum. 130 species occur in the Deyo- 
nian rocks of Rhenish Prussia and Central Europe, and less than 20 
in America. Our Carboniferous rocks contain 47 species; but, as we 
have seen, only 4 species are common to the two formations. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 187 


Nautilus.—The European area, so rich in the 4 genera previously 
noticed, is unexpectedly poor in Nautilz, not more than 11 species 
haying been described therefrom, and only 4 or 5 from America. 
We possess only 2 species, VV. germanus and NV. megasipho; the 
former Middle Devonian, Torquay, the latter Upper Devonian, from 
Petherwin. 

Poterioceras.—Only in the Upper beds at Petherwin is this genus 
represented, and by 1 species (P. fusiformis) ; it also passes to the 
Carboniferous rocks. The analysis of the Devonian Cephalopoda and 
their distribution show that the British species number 60, belonging 
to the 6 genera I have separately discussed. Only 1 is Lower Devyo- 
nian ; the Middle Devonian has yielded 4 genera and 33 species, and 
the Upper 5 genera and 33 species. 

Looking, then, at our small Devonian Cephalopod fauna (6 genera 
and 60 species) as compared with that of Europe (20 genera and 500 
species), which alone nearly equals all the species in the 14 classes 
in the British Devonian rocks (544), we readily see the value of 
this fauna in Europe, Belgium, the Eifel, the Harz, Saxony, 
the Fichtelgebirge, Russian Poland, Russia in Europe, and Spain. 

In Belgium, France, the Hifel area, the Fichtelgebirge, and Saxony, 
the Devonian group, as with us, readily falls under the 3 divisions of 
Lower, Middle, and Upper, andin the main the grouping corresponds 
or may be correlated with ourown. In Belgium (south of the Condros) 
the Psammite of Condros, the schists of Famenne, and the limestone 
of Frasne are the equivalents of our Upper Devonian as exhibited 
at Petherwin in North Cornwall, and the Baggy, Pilton, Brushford, 
and Marwood beds. ‘The ‘“ Givet Limestone” at the summit of the 
Middle group is the equivalent of the Stringocephalus-beds of North 
Devon, but with us stratigraphically lower. 

The Calceola-schists and the (Spirifer) “ Cultrijugatus-Stufe ” 
of Belgium are our Calceola-beds of Torquay and Newton in South 
Devon, and the typical Middle Devonian. The schists of Burnot, 
Ahrian, and the Coblentzian and Gadinian correspond with the 
Tinton and Lower slates and grits of North Devon—our Lower 
Devonian (the lowest probably not seen). In the Eifel area, the 
beds of which, like the deposits of Belgium, correspond so closely 
with our own, the Goniatite-schists and (Rhynchonella) Cuboides- 
limestone are, like those of North Cornwall and Devon, the upper 
division; and, as in Belgium, the Calceola-schists and Cultrijugatus- 
Stufe are the equivalents of the South-Devon limestones or Middle 
Devonian: the Vichter schists, Ahrian (of Dumont), and the Coblentzian 
beds represent the mass of the lower shales, slates, and grits below 
the Hangman beds in North Devon. M. Gosselet*, in his succession 
of the Devonian rocks of North France and Belgium, divides the 
whole into 6 divisions or beds. The corresponding North-French 
beds to our Upper Devonian are probably the Psammites of Condros, 
the slates of Famenne, and the Boulonnais beds. 

The Givet Limestone and;Calceola-slates are our Middle Devonian, 
and the systeme Rhénan the Lower. 


* Bull. Soe. Géol. de France, vol. xviii. p. 18. 


188 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


M. Gosselet* again divides the Devonian of the Franco-Belgian 
region into Upper, Middle, and Lower, but with differences which 
have paleontological value. 

His Upper Devonian contains the Psammites du Condros in 2 divi- 
sions; the Schistes de Famenne in 4 divisions. 

The Middle Devonian one large group (caleaire de Givet) divided 
into 9 beds or bands. 

In the Lower Devonian Gosselet places (1) the schistes & Calecoles 
(in 3 divisions), (2) Poudingue de Burnot, (3) the Grauwacke 4 
Leptena Murchisoni (in 2 divisions), (4) the Couches de Gedinne and 
conglomerates and schists (in 3 divisions). 

In the departments of the Loire Inférieur and Maine-et-Loire, 
according to M. Caillinaud, the Lower Devonian occupies large areas 
in the west of France; and in the same departments M. Bureau 
has determined the presence of the Upper Devonian. 

In Rhenish Prussia, in the Eifel district, the Devonian strata were 
long studied by Ferd. Romer, of Breslau, and Dr. Emanuel Kayser. 
They determined the following horizons and successions f :— 

MherGonvatitelschists ) esses. ee 


Cuboides (Rhynchonella) limestone ... Marly limestone, (Uo ee 
Stringocephalus-schist 
Calceola- schists ie pean hy cre ec inerene Marl, Middle Devonian. 
Cultriyjugatus-Stufe (Spirifer) 
Vachteriscistee teri ene cca 


Ahrian schist (Dumont) ............... Sandy schists, Lower Devonian. 
Coblentzian iia ae ere een ee 


eeeeeesecsecesrese 


F. A. Romer, of Clausthal, has correlated the parallel horizons or 
deposits of the Harz and the Eifel. In the Hifel he gives 7 divisions, 
in the Harz 8; they are compared with Aix-la-Chapelle and Couvin. 


In the ‘ Paleontographica’ (for 1854), F. A. Romer divides the 
Devonian rocks of the Harz into 9 portions :— 


. Amay schists (jiingere Grauwacke), 
. Cypridina-schist. 

. Goniatite-schist. 

. Iberg limestone. 

. Receptaculites-schist. 

. Stringocephalus-limestone. 

. Orthoceratite or Wissenbach schist. 
8. Calceola-schist. 


9. Spirifera-sandstone (altere or Rhenish Grauwacke). 
Characteristic fossils are given with each, and the chief localities 
where the beds are exposed. F 
The Fichtelgebirge, through the researches of Prof. C. W. Gumbel, 


exhibit 4 divisions or groupings :— 


JWoOuUp WN eH 


1. Cypridina-schist......... Upper Devonian (in 3 stages). 


2. Calamopora-schist ...... ere re 
3. Tentaculite-schist ...... J tibelally Dereuten 


4, Nereite-schist ...........- Lower Devonian. 


* Annales des Mines, 6° sér. tom. xii. p. 595. 

+ Zeitschrift d. deutschen geol. Gesell. Jahrg. 1871, p. 375. bang 

t ‘Paleontographica,’ Dunker and Meyer, x1. p. 109, “ Ueber Clymenien in 
den Uebergangsbilden des Fichtelgebirges.” 


ANNIVERSARY ADDRESS OF THE PRESIDENT, r89 


I have thus referred to some of the chief European localities and 
systems to show the importance and development of the continental 
Devonian rocks with their enormous fauna, as compared with the 
clearly determined but comparatively poor British series, our whole 
fauna only numbering 550 species at most. Unless some reference 
had been made to the American and European Devonians no force 
could have been given to the system; and its geographically dis- 
jointed state (even on the continent) tends to detract from the original 
unity of the whole, but at the same time to show that it was, 
and has been, one of the most extensive systems on the two conti- 
nents, and even in the British Islands could we but uncover the 
extensive area between Belgium, the Boulonnais, and Devon and 
Cornwall, now deeply buried under the Secondary and Tertiary 
systems. 

I cannot pass over the Devonian formation of North America 
and the Canadas, either physically or paleeontologically, for we know, 
through the labours and researches of Hall, Bigsby, Newberry, 
Meek, Shumard, Winchell, Dawson, and others, how great and widely 
extended is the Devonian system in North America, and how persistent 
are the conditions and successions; the state of New York and the 
Canadas, with the Gaspé region in the east, have as yet exhibited 
the Devonian rocks in greatest development, all the three great 
horizons being present. The Hamilton and Upper Helderberg 
groups, with the Catskill, Chemung, and Portage groups above, 
occur in the States of Pennsylvania, Ohio, Llinois, Missouri, and 
Michigan in full development. In lowa and Tennessee the Upper 
Devonian only is present. In New-York State the 11 recognized 
divisions make up a mass of limestones, sandstones, shales, flag- 
stones, and conglomerates upwards of 8000 feet in thickness. In 
Pennsylvania the Devonians are 12,000 feet thick. The great 
Middie or Hamilton group is developed in 6 of the 8 States, the 
Lower or Helderberg in 4. Principal Dawson, in his ‘ Acadia,’ has 
shown that the Devonians of the St.-John’s area, New Brunswick, 
exhibit peculiar features different from those of the Western States 
before mentioned, but nevertheless are divisible into three stages 
or horizons*. 

The above generalizations relative to the development and dis- 
tribution of the European and American Deyonians are given to 
show their great value and extent as compared with the really 
or comparatively poor fauna of the British Islands. That the whole 
formation or system once extended universally over the area thus 
roughly traced there cannot be any doubt, nor that probably during 
one general age one or other of its three divisions may have occurred 
homotaxially ; but as a whole it occupied all that time or epoch 
extending from the close of the Upper Ludlow to the base of the 
Carboniferous. In North Devon its upper member passes insensibly 
and conformably into the Carboniferous, which is itself in an abnormal 
eondition. 

The Scotch and Irish beds of this age differ from all others, the 

* © Acadia, 2nd ed. pp. 503-505. 


Igo PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Trish Coomhola beds, with Anodonta and Paleopteris or Adiantites, 
being undoubtedly of Upper Devonian or Upper Old Red Sandstone: 
age. Mr. Kinahan would class the Old Red Sandstone of the 
Dingle and Cork area as Lower Carboniferous, and corresponding 
in the Limerick area to the Lower Carboniferous sandstones and 
shales. Kinahan further states that in no place in Ireland has 
the Old Red Sandstone a defined upper boundary, one group gra- 
duating into the other. Furthermore, he says, only in Munster 
and in the hills between Lough Erne and Pomeroy, in the counties of 
Fermanagh and Tyrone, is the Old Red Sandstone at the absolute 
base of the Carboniferous formation, as in all other places the 
rocks so called and described are on different geological horizons, 
ranging up to the base of the Coal-measures*. 

As far back as 1863 Jukes and Salter stated that the yellow lime- 
stone of the South of Ireland is the upper part of the Old Red Sand- 
stone, with the Coomhola grits between it and the Carboniferous 
slate with Avicula damnoniensis and Cucullea &e. 

Piscrs.—50 genera and 125 species of fish range through the 
Old Red Sandstone. It can hardly be said that ichthyic life com- 
menced prior to the commencement of the deposition of Old Red Sand- 
stone or the Devonian age. The few genera and species that made 
their appearance in the British seas at the close of the Silurian 
period (8 genera and 12 species, 10 of which passed to the Old Red 
Sandstone) can only be traced into the very lowest beds of the Old 
Red in the Silurian area of Ludlow, Kington, and Ledbury, and 
scarcely passing beyond the passage group; we know them not 
higher in the system. Inthe North of Scotland and in the Orkneys, 
Caithness, and Cromarty area, the Lower Old Red Sandstone is 
known to have furnished 18 genera and about 60 species ; no prior 
stratigraphical relation or origin is known. They occur in that 
region, like the strata or rocks which contain them, fully developed 
and in vast numbers. Unlike the Old Red Sandstone of the Silu- 
rian area the formation has there no fossiliferous base. ‘The Silurian 
rocks are absent, the sandstones and conglomerates lying uncon- 
formably upon the metamorphic rocks. The only other organisms 
are Phyllopod and Eurypterid Crustacea, with remains of 10 or 11 
genera of plants. 

To no living geologist are we so much indebted for our intimate 
knowledge of the Old Red of Scotland as to Prof. A. Geikie. His 
paper in the ‘ Quarterly Journal of the Geological Society’, on the 
Old Red Sandstone of the South of Scotland, and his last elaborate 
essay upon the Old Red Sandstone of Western Europe, are ex- 
haustive so far as they have gone. Prof. Geikie shows that there 
is no middle Old Red Sandstone in either North or South Scotland, 


* Vide Kinahan, ‘ Geology of Ireland,’ pp. 50-94, for his views and much 
valuable information. 

+ “On the Old Red Sandstone of the South of Scotland.” By A. Geikie, Kisq., 
F.G.S. Quart. Journ. Geol. Soe. vol. xvi. pp. 312-328. 

t+ “The Old Red Sandstone of Western Europe.” By A. Geikie, LL.D., 
F.R.S. Trans. Roy. Soc. Edinburgh, vol. xxviii. 1878. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 1gt 


but that it consists only of two great divisions—a lower, passing down 
conformably into the Upper Silurian shales, and an upper, graduating 
upward into the Lower Carboniferous sandstones, with a complete 
discordance between the two series. 

Prof. Geikie in his memoir traces the series of changes in the 
physical geography of Western Europe which took place between 
the close of the Upper Silurian and the commencement of the Car- 
boniferous period. Viewed in a large sense the Old Red Sandstone 
of Great Britain groups itself stratigraphically into two divisions, 
physically and paleontologically distinct, a lower and an upper. 

Prof. Geikie regards the Old Red as a great lacustrine accumula- 
tion, and treats of the separate basins of deposit as lakes, to which 
he has assigned different names :— 


1. Lake Orcadie.-—* Kmbracing all the Old Red Sandstone to the 
north of the Grampian range, including all the Orkney 
Islands.” 

2. Lake Caledoma or the Mid-Scottish Basin.—* Occupying the 
central valley of Scotland between the range of the High- 
land mountains on the north and that of the Silurian 
pastoral uplands of the southern counties. This basin was 
probably prolonged across the Firth of Clyde into the north 
of Ireland.” 

3. Lake Cheviot.— A portion of the south-east of Scotland and 
the north of England, extending from near St. Abb’s Head 
south-west along the base of the Silurian hills to the head of 
Liddesdale, and including the area of the Cheviot Hills.” 

4, The Welsh Lake.—‘ The Old Red Sandstone region of Wales, 
bounded on the north and west by the Cambrian and Silu- 
rian rising grounds, but its eastern and southern extension 
obscured by later formations.” 

Lake of Lorne-—*“ A district in the North of Argyllshire ex- 

tending from the south-east of Mall to Loch Awe, and 
perhaps northward up the line of the great Glen.” | 


Ca 


OV 


At p. 374 of his memoir Mr. Geikie gives the order of succession 
among the strata, the thickness, and typical localities of the Old Red of 
Caithness, which he makes 16,200 feet thick, and then treats of all 
the subdivisions separately. Pp. 406-414 treat of the Orkney 
Islands paleontologically and physically; and pp. 414-421 of 
the Shetland Islands in the same manner. The description of the 
Basin of the northern Firths occupies 26 pages. 

I commend this great addition and acquisition to the history of 
the Old Red Sandstone of Scotland to every student of geology. 

Professor Hull has lately ably discussed the vexed question of 
the geological relations of the rocks of the south of Ireland to those 
of North Devon and other British and continental districts*. 

Mr. Hull had previously discussed, in his paper upon the Dingle 
beds and Glengariff grits and slates, the relations of the Upper 


* Quart. Journ. Geol. Soc. vol. xxxvi. pp. tell (1860). 
Tt Quart. Journ. Geol. Soc. vol. xxxv. pp. 699-728 


ig2 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Silurian series of the south-west of Ireland to those of the Silurian 
region of Engiand, and also the relation of the Old Red Sandstone 
to the Dingle beds, and shown that, as in the north of Scotland, 
all through the south and centre of Ireland the Old Red is every- 
where unconformable to the rocks on which it reposes, while it 
passes up conformably into the Carboniferous series. 

Prof. Geikie has suggested that the Dingle beds or Glengariff 
grits may represent the Old Red Sandstone of Scotland (Joc. cit.). 
Should this be the case they would be the marine representatives 
of lacustrine deposits. Both, according to Mr. Hull, are uncon- 
formably overlain by Upper Old Red conglomerate and sandstone. 

Mr. Ralph Tate held the view that the Dingle beds are the 
equivalents of the Tilestones or Passage-beds of England and Wales*, 
or the lowest Old Red. Prof. Hull endeavours to show that between 
the Glengariff beds and the succeeding formation, be it what it may, 
either Ola Red Sandstone or Carboniferous, there is a great blank, 
or, in other words, ‘unrepresented time.” This long period Mr. 
Hull believes was filled up in the south and south-west of England 
and in Belgium by the extensive series known as the “ Middle” 
and ‘“‘ Lower Devonian beds,” lying between the “ Foreland grits ” 
on the one hand and the “ Pickwell-Down Sandstone ” on the other. 
Prof. Hull also believes that ‘“‘while a deep sea, in which were 
deposited the Middle and Lower Devonian beds, overspread the 
south of England and adjoining continental areas, land conditions 
prevailed in the south of Ireland during the same period.” 

The analogy between the OJd Red Sandstone of the south of Ireland 
and its supposed representatives in North Devon, Belgium, and 
Scotland receives critical analysis from Mr. Hull. That author 
endeavours to demonstrate that the Middle and Lower Devonian 
rocks of North Devon are totally unrepresented in the Irish area, no 
correlation being possible above the Foreland beds. I believe no 
one who has ever examined the area south of the Pickwell-Down 
Sandstone can doubt that the Baggy, Marwood, Croyde, Barn- 
staple, and Pilton beds are upon the same general horizon as the 
Carboniferous slate and Coomhola grits of Ireland, and that the 
Pickwell-Down Sandstones represent the Upper Old Red Sandstone 
of the south of Ireland. Both on physical and paleontological 
grounds these two groups seem to agree. Although the freshwater 
shell Anodonta Jukesit has not occurred in the Baggy or Pilton series 
in North Devon, its plant associate, Adiantites (Palwopteris) hibernicus, 
has been found in the Baggy beds, or immediately above the Pick- 
well-Down Sandstone (Upper Old Red). Ido not despair of finding 
the Irish shell in North Devon, especially after its occurrence in 
Northumberland in the lowest Carboniferous rocks of that area. We 
may well believe that the “ lacustrine ” conditions of the south of 
Ireland gave place to marine conditions over the Devonian area 
during the same time. 

Mr. Hull proposes the following tabulated form as representative 
tor two areas, viz. the south of Ireland and North Devon :— 

* Weale’s series. Geology. Portlock and Tate, p. 72. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 193 


South of Ireland. North Devon. 
( Carboniferous Limestone. Carboniferous Limestone. 
eee Osrboni- | Carboniferous Slate. Beas beds. 
ferous beds. Coomhola Grit and Slate (pas- ST en eae 
eos ee Baggy and Marwood beds 
\ 8 of ( Cucullea-zone). 
Old Red Sand- ( Kiltorean beds. Upcot beds ? 
stone or Upper } Old Red Sandstone and Conglo- =Pickwell-Down Sand- 
Devonian. merate. stone. 


Prof. Hull then enters minutely into the question of the equiva~ 
lency and relationship of the North-Devon, Lower and Middle 
Devonian series to beds, if any, in the South-Irish area. The Irish 
Upper Old Red Conglomerate does not appear to have had, in that 
area, any immediate predecessor; and Mr. Hull suggests and con- 
tends that the unconformity between the Glengariff beds and 
the Old Red or Carboniferous beds proves that certain strata are 
absent or were never deposited over the South-Irish area. If Mr. 
Hull’s correlation be correct, it may possibly follow that the Middle 
and Lower Devonian of North Devon may occupy the place of the 
missing strata in the south of Ireland. Furthermore, Mr. Hull 
regards the Foreland grits and slates (the base of the North-Devon 
beds) as representing in part the Glengariff beds, which he believes 
to be of Upper Silurian age. The Foreland grits and sandstones, the 
recognized base of the Devonian rocks in Devonshire, Mr. Hull would 
correlate with the uppermost Silurian or Passage-group of Sir R. Mur- 
chison; and he also infers that the great gap which appears to exist 
in Ireland between the Glengariff beds and the succeeding Old Red 
Sandstone and Carboniferous series was filled up in North Devon by 
the Mortehoe slates, the Ilfracombe series, and the Lynton beds, or all 
the strata that exist between the base of the Pickwell-Down Sand- 
stones and the top of the Foreland grits. Mr. Hull’s views as regards 
the representative beds in the south of Ireland, North Devon, and 
Belgium are carefully tabulated on p. 266 of the ‘ Quarterly Journal of 
the Geological Society,’ vol. xxxvi., 1880, the relations of each group 
being given. The Belgian and Boulonnais sections are also referred 
to and tabulated. Scotland, through Geikie’s latest memoir (Joc. 
cit.), is compared with both areas, Devon and North and South Ire- 
land. On page 273 is also given a table of representative forma- 
tions for Ireland, North Devon, South Wales, Scotland, and Belgium. 
These two contributions, by Professors Geikie and Hull, to the his- 
tory of the Old Red and Devonian rocks of Scotland, Ireland in 
part, and North Devon have materially added to our knowledge of 
their physical history and condition. 

The distribution of the 49 or 50 genera and 125 species of fish 
through all the known deposits of Old Red in the British Islands 
seems to result in there being no middle fish-bearing group, either 
in England, Scotland, or Ireland. Only 2 genera and 2 species are 
regarded as Middle Old Red, Kiwcephalaspis (Cephalaspis) Agassiz, 
Lank., and Holoptychius Murchison, Ag.; but it would hardly 
appear that these two species are rightly placed stratigraphically. 


r9O4 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


On paleontological grounds this fully confirms the most recent re- 
searches relative to the absence of the Middle Old Red Sandstone 
in Scotland and Ireland, and probably in the Welsh area also. 

8 genera and 12 species of Elasmobranchii (Placoidei) range 
chiefly through the Lower Old Red—many with only 1 representa- 
tive, viz. Byssacanthus, Conchodus, Cosmacanthus, Cienacanthus, 
Ctenoptychius, and Parexus; also Onchus with 4 and Ptychacanthus 
with 2 species. The remaining 114 species represent 9 families and 
41 genera, all Ganoidei :— 


Genera. Species. 


ies @colacamthanh < Soler. 2 2 
2) ePlacodermitS.. 00 ero. 3 10 
3. Cephalaspide .......... 15 40 
ANeChemolepimmyern sais os 1 4. 
Sy, (Cahqnwodbyomeim 44 S66 ooc 8 23 
6. Sauredipterimi 9.) cis) a4 3 9 
MeN CLEMOGIpLETIMI «oye cr ee see 2 ve 
8. Phaneropleurini .......- 1 L 
DUNC aria dices lo) ata as cart 6 23 

41 114 


The Lower Old Red is represented by 36 genera and 85 species, 
and the Upper by 15 genera and 25 species; and I know of no 
spectes that is common to the Old Red Sandstone proper and the 
Carboniferous group above. 

Those genera that seem to specially characterize the Upper Old 
Red Sandstone, chiefly Scotch, are Actinolepis, Asterolepis, Bothrio- 
lepis, Cosmacanthus, Dendrodus, Glyptolemus, Glyptopomus, Holopty- 
chius (some species), Homothorax, Lamnodus, Pamphractus, Phanero- 
pleurus, and Phyllolepis*. The 2 genera(?) and 2 species (?) in the 
so-called Middle Old Red tend also to show, through physical re- 
search, that no such grouping exists. 

56 genera and 150 species, other than British, have been recog- 
nized in Europe and America, but are allied through about 40 
genera, or that number is common to all three areas, America, the 
British Islands, and Europe. 

Of the entire British Devonian and Old Red fauna (544 species) 
we have seen that 52 genera and 51 species pass to the Carboniferous. 
ft give the number of species in the classes that connect the two 
epochs :— 


* Dr. Traquair is now engaged in carefully revising and describing the 
Ganoid fishes of the Old Red Sandstone. They could not be in better hands ; 
few men are so competent to undertake so large a task. Many names are only 
provisional. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 1Q5 


Genera. Species. 

ARUN LTS 2 ek.a, of" lash ma etoeeas 2 
PP OLOZOM UO fh sd Iolo: sich acdodet a none pass. 
ENG EUMOZO isp. 40s) i scdeusia)eue alors 12 
ichimodermata. 6)... 3 6 
EATIITCING GY 7 5 ean ey cae nae ene none. 
GUISE ACC AE ro 5 Ais oa ci ocnk «uty ay: 1 1 
IBIEVIOLORS. 9 SB gle Be oer enre + 4 
I ACHTON OMS cre ey suev-yersy a ces 10 16 
Lamellibranchiata ........ 4 5 
Gasteropoda.. 2... es 3 5 
NCR OO arch alee) cre alejet ays ® none. 
IEvetenopoda: ..i).).. 70:0. os 5 1 2 
Cephalopoda. . py -icussworen<dcesi- 3 9 
WBS CORE fe Arc ech tvei sista! ceive ee none. 

32 51 


It will be seen that, out of the 51 species, the two large connect- 
ing groups are the Brachiopoda and the Cephalopoda. 

Professor Hull, in May 1880, communicated a paper to the Royal 
Dublin Society ‘‘On the Relations of the Carboniferous, Devonian, 
and Upper Silurian Rocks of the South of Ireland and those of 
North Devon”*; and although resembling his previous paper ‘“ On 
the Geological Relation of the Rocks of the South of Ireland to those 
of North Devon &c.,” it treats of the subject-matter in a different 
light. Prof. Hull enters more into the paleophysical geography of 
the south of Ireland and North Devon, as indicated by the relation 
of the formations and areas respectively. In the south-west of 
Treland, north and south of Dingle Bay, the rock-groups which rise 
into high elevations belong to the Dingle beds or Glengariff grits and 
slates of Professor Jukes. The lowest beds of the Dingle promontory 
do not reach the surface amidst the mountain-ranges to the south of 
Dingle Bay. 

The Glengariff beds may measure about 10,000 feet in thickness. 
There is a total absence of the Old Red Sandstone at the base of the 
Carboniferous beds+. Contortions in the Glengariff beds are of earlier 
date than the deposition of the Old Red Sandstone or Carboniferous 
beds ; this therefore implies that between the Glengariff beds and 
the Old Red Sandstone there is a vast amount of wanting strata, 
indicative also of unrepresented time—in other words, a wide gap 
and “prolonged interval of time actually separates the two formations. 
It is the representation or filling up of this gap that Prof. Hull en- 
deavours to show through the uninterrupted sequence in the Middle 
and Lower Devonian beds of North Devon, or through the whole 
series from the lowest Devonian up to the base of the Carboniferous 


* Proc. Royal Dublin Soe. vol. i. new series, 1880, pp. 135-150. 
t Hull, “ On the Geological Age of the Rocks forming the Southern High- 
lands of Ireland,” Quart. Journ. Geol. Soc. vol. xxxv. p. 699 e¢ seg. (1879). 


Fa 


1g0 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


formation in that area. In no other district do we find evidence to 
account for such loss, or beds that can be correlated with those of the 
south of Ireland. Mr. Hull tabulates the succession of the Carboni- 
ferous and Devonian rocks of North Devon and the south of Ireland, 
and endeavours to show their relation and probable contempo- 
raneity as follows* :— 


North Devon. South of Ireland. 


( Harthy limestones, with Posidono- Carboniferous limestone. 
| mya (Venn Quarry). 


Lower Carboni- 4 Barnstaple slates. Lower Carboniferous slate. 


OES Sea: | Pilton beds. Coomhola grits and slates 
| Marwood beds ( Cucullea-zone). (with Cucullea &c.). 
zy Upcot flagstones. Kiltorcan beds (with Ano- 
Pe ee | donta Jukesit). 
; leiclvell- Down sandstones. Old Red Sandstone and 
conglomerate. 
= Mortehoe slates. 
Ne Ilfracombe limestone group. Strata absent over the area 
a Hangman grits(Martinhoe beds). in the south of Ireland. 
eaten vera Lynton shales and limestones. 
NA eee Foreland grits (base invisible). Glengariff beds, passing 


down into the Upper 
Silurian beds. 


The above table is probably the true reading of the areas under 
notice. This table is followed by a concise description of the North- 
Devon formations and their fossils, in descending order, from the 
Carboniferous series south of Barnstaple to the Foreland grits. It 
is worthy of attentive study; and Mr. Hull has marked with an 
asterisk the identification by Mr. Baily of those species also occurring 
in the Carboniferous and Coomhola beds of the south of Ireland, 
thus showing still further, on paleontological grounds, the rela- 
tionship between the now separated, but probably once united, 
areas. 

Mr. Hull’s views “ that the missing chapter between the Silurian 
and the Carboniferous, in the paleontological history of Ireland, is 
supplied by the rocks of Devonshire with their marine organisms” 1s 
probably near the truth ; and the proportions assumed in the geolo- 
gical series by the Devonian rocks “ offers a solution to one of the 
problems of Irish geology.” 

Professor Hull enters into geographical considerations of a minute 
character, to which I must refer, and which bear significantly upon 
the relative changes of sea and land from the Silurian to the close of 
the Carboniferous period. The successive phases of the physical 
conditions are given through four diagrams of the British Islamds on 
plate v., which show the relations of land and sea, or elevation and 
depression. 

He believes that there is only one true Old Red Sandstone in North 
Devon, which is represented by the Pickwell-Down Sandstone, and 
that the Upper Old Red of Scotland and Ireland is not the equivalent 
of the Devonshire marine Devonian strata, which are newer and 


* Hull, loc. cit. p. 144. 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 197 


higher. If Prof. Geikie is right in saying that the Scotch Old Red 
Sandstone represents the Irish Glengariff beds, then Prof. Hull may 
be right in concluding that the Scotch beds are the lacustrine equi- 
valents in time of the marine uppermost Silurian strata, the Upper 
Scotch Old Red being the equivalent of the same in Ireland. 


TABLE X V.—Devonian. 


: a : ; 
a 8 Ba | oe 
OI gq — 

Se EI 5 a eu 
ae > e & a 
oS Classes. © S © S53 
=) 3 n 2) ° 5 

Qj oO H mt foal eS 
Sen feece me Sees By aae 
my O D = = =) py 
2 2 
ffpitlernitea Weeecewereeese-- 12 18 : 3 12 2 
WOCOZOR, <sccsses ca cl son 3 3 
Hydrozoa. 
* € 4 
PNCEIMOZOA) catone sc teeece- 24 52 : ab 5% ql 
Wehinodermata......... 10 24 2 eA By 3 
E J:\10 0016) 110 ER Sere 2 2 1 1 
3 @rustacea ....c..s.00cs.. 20 o7 a F q 7 
ROW Z Oa acc ms sencouseos- 7 il Z S ss 
3 Brachiopoda............ 26 116 ay 28 a ate 
Lamellibranchiata ...| 20 39 + 23 29 3 
Gasteropoda............ 13 45 36 14 3 
Pteropoda. 
a 2 
Eleteropoda <.2.5...--.- 2 8 3 3 5 2 

suit Cephalopodas.......:..- 6 60 : Se 33 3 

A WISCES! © cela cece de HOR 25 ae 2 ae 
3G Ol ee 195 544 era 268 Toe BA 


Contents: 195 genera, 544 species. 
Pass to Carboniferous: 32 genera, 51 species. 


CARBONIFEROUS SYSTEM. 


We must go back beyond the present decade for new matter 
bearing upon the history and distribution of the British Carboni- 
ferous species—not that any significant change has taken place in 
the nomenclature or division of the groups, but speculative and 
philosophical views have been propounded by many authors upon 
the distribution and redistribution of the Carboniferous land- 
surfaces and their relation to still older areas. No one has con- 
tributed more than Professor Hull to the elucidation of the Carbo- 
niferous strata in England, Scotland, and Ireland. This is notably 


VOL. XXXVII. p 


198 -PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


the case with his paper on iso-diametric lines representing the distri- 
bution of sedimentary strata as distinguished from calcareous strata — 
with reference to the Carboniferous rocks of Britain*, also his paper 
upon the evidence of a ridge of Lower Carboniferous rocks crossing 
the plain of Cheshire beneath the Trias &c.t Another and equally 
important communication was brought by him before the Society 
«‘ On the Thickness of the Carboniferous Rocks of the Pendle Range 
of Hills in Lancashire, illustrating the author’s views regarding the 
South-easterly Attenuation of the Carboniferous Sedimentary Strata 
of the North of England”t. This was followed by another paper 
entitled, ‘‘ Observations on the Relative Ages of the leading Physical 
Features and Lines of Elevation of the Carboniferous District of Lan- 
cashire and YorkshireӤ. Prof. Hull again in 1876 still further 
enriched the literature of the Carboniferous system through a valuable 
and exhaustive paper “On the Upper Limits of the essentially Marine 
Beds of the Carboniferous Group of the British Isles and adjoining 
Continental Districts, with Suggestions for a fresh Classification 
of the Carboniferous Series”’||. I consider these contributions of Mr. 
Hull of the highest importance, as bearing upon theoretical geology, 
and tending to excite us to closer research and broader views as to the 
physical geography and distribution of life during the Carboniferous 
period—certainly one of the most prolific during Paleozoic times, 
governed by the distribution of land and water or their relations one 
to the other, under marine, estuarine, and freshwater conditions. 

The nature of sediments, whether physical (such as inorganic 
sandstones, clays, and shales) or organic (as calcareous marine or 
freshwater limestones and their relation to life during deposition), 
has hardly received the attention it demands. Little has been 
written relative to these since Dr. Bigsby, in his memorable papers 
in 1858-59, entered into the history and nature of the sediments 
laid down over the sea-bottoms, and the immediate relations of 
animal life to the strata which the fossil remains now occupy. He 
made comparisons of the State of New York and Wales for obvious 
reasons. Bigsby showed, through his elaborate researches, that, 
owing to a determinate law (with few exceptions), the accumulated 
‘sediments graduate into each other, clearly seen in Wales, Scan- 
dinavia, Russia, and America.” The change in sedimentation usually 
takes place slowly and tranquilly, and the majority of life in the 
uppermost or terminated section perishes, thus showing that destruc- 
tion or change can take place without any marked disturbing force. 
This is perhaps more manifest in the accumulation of the Carbonife- 
rous strata throughout Britain than probably in any other formation. 
The marked difference, yet, on the whole, agreement between the 
Scotch and border formations, or the Calciferous Sandstone, north of 

* “On Iso-diametric Lines as means of representing the Distribution of 
Sedimentary Clay and Sandy Strata as distinguished from Calcareous Strata, 
with Special Reference to the Carboniferous Rocks of Britain.” Quart. Journ. 
Geol. Soc. vol. xviii. pp. 127-146 (1862). 

Loc. crt -volwscxixen ps Will { Loe. cit. vol. xxiv. p. 319. 

§ Loe. cit. vol. xxiv. p. 223 (1868). 

|| Loc. cit. vol. xxxil. pp. 618-651 (1876). 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 199 


the Cheviots to the latitude of Burnt Island, and those same strata 
south of the Tweed to the latitude of Harbottle and on towards 
Rothbury &¢., would seem to be due to comparatively small changes 
of conditions; and from the mapping by the Survey, Prof. Hull’s sug- 
gestion as to the origin of and difference between the calcareous and 
true sedimentary or argillo-arenaceous strata, especially as exhibited 
in the Upper Silurian, Devonian, and Carboniferous formations, appear 
to be confirmed. Mr. Hull regards the true calcareous strata as dif- 
ferent in origin and distribution from the other stratified rocks with 
which they are associated,—not, indeed, properly coming under the 
term sedimentary, this term being restricted to “gravels, sandstones, 
shales, and clays,” the presence of both classes of rock in the same 
geological group being no argument in favour of their similarity. 
Whenever interstratifications occur the limestones may be regarded as 
occupying neutral ground between the respective areas of dispersion 
of the sedimentary series. Could it be possible to trace the sources of 
the ‘‘sedimentary” strata of any formation on the one hand and of the 
limestones on the other, they would be found expanding in opposite 
directions, this arising from the difference in the origin of the two 
classes of stratified rocks, the calcareous being essentially organic and 
the sedimentary essentially mechanical. We may, I believe, regard 
the predominance of ‘“‘ sedimentary strata” as highly unfavourable 
to the development of calcareous deposits in the same group of 
rocks, from the fact of interference with the development of 
life. 

Looking at the physical geology of Beran prior to the deposition 
and distribution of the Carboniferous strata it is probable that “a 
barrier of land existed, stretching from Wales eastwards, touching 
the southern ends of the South-Staffordshire and Warwickshire 
coal-fields, and including the Cambrian rocks of Charnwood Forest.” 
This barrier Prof. Hull believes was possibly an extension of 
the Scandinavian promontory, stretching across the Irish Sea to 
embrace the Cambro-Silurian districts of Wicklow and Carlow, and 
dividing the Carboniferous rocks of South Wales, Somersetshire, and 
Dean Forest from the coal-tracts of Central and Northern England 
and Scotland, the strata on each side belonging to two distinct 
systems of distribution; and their origin is therefore ‘ due to two 
different sets of oceanic currents ” (Hull). 

The correlation of the Carboniferous rocks of Britain and Ireland 
is even now a matter of difficulty and of difference amongst 
those best able to decide. Even the continuous series between 
England and Scotland from the Calciferous Sandstones of Fife, 
south of the Cheviot, to the Yoredale series of Durham are most 
difficult to correlate—much that has been called Carboniferous Lime- 
stone being calciferous in one area and even Yoredale in the other. 
The literature of the whole question is scanty and unsatisfactory 
in the extreme; and in the construction of my Tables these dif- 
ficulties have met me everywhere. The wide difference that exists 
in the mode of derivation and accumulation of the materials com- 
posing the sedimentary series of the northern and southern groups, 


P 


200 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


as well as their zoological differences, are matter for much research. 
Materials have not long been obtained whereby we may institute 
comparisons between the different members even of the British 
group of Carboniferous rocks, setting aside our intimate connexion 
on the one hand with the European, and on the other with that of 
the American continent, especially New Brunswick and Nova Scotia, 
whose relation to Scotland through the Coal-measures can 
hardly be doubted. Those beds which overlie the Carboniferous 
Limestone in the south of Ireland have hardly yet had assigned 
to them their true place by correlation with those of England; 
neither has it been satisfactorily settled what is the precise strati- 
graphical commencement of the Lower Carboniferous series of the 
North-Devon area, or where the line should be drawn between the 
Upper Devonian and Lower Carboniferous. Both horizons are 
fossiliferous, and it is only in the south of Ireland that we have 
similar conditions, or where a comparison can be instituted be- 
tween the two. Again, as regards the Coal-measure “shales and 
flags ” immediately overlying the Carboniferous Limestones of the 
south of Ireland, there can be no doubt that they represent the 
Millstone Grit of the English Coal-fields. This was the opinion of 
the late Professor Jukes, who, in the explanatory memoir to sheet 
No. 137 of the Irish Geological Survey maps, says, ‘‘ Doubtless the 
whole of the Coal-measure series of Central Ireland is contempo- 
raneous with the lower part only of that of Central England, in- 
cluding the Millstone Grit in that lower part.” Prof. Hull also 
doubts not that through the identification of the Gannister beds in 
the Leinster Coal-field, which, in the north of England, overlie 
the Millstone Grit, that contemporaneity, to a large extent, occurs 
between the English and Irish Carboniferous beds, and this especially 
on physical grounds. 

Through the close research of the Irish Survey in the Leinster 
and Munster Coal-fields they are now enabled to trace out and 
distinguish four divisions:—1, the Yoredale beds; 2, the Millstone 
Grit ; 3, the Gannister beds; and 4, the Middle Coal-measures as 
determined in England. Hitherto they have simply been classed 
under the general term ‘‘ Coal-measures.’’ 

Professor Hull endeavours to show to what extent the British 
Carboniferous rocks have their representatives in Ireland; and he 
also proposes to establish a ‘‘ Middle Carboniferous series,’ which 
shall include all the strata lying between the Carboniferous Lime- 
stone and the Middle Coal-measures, including the Yoredale beds, 
Millstone Grit, and Gannister beds. Mr. Hull is careful to impress 
upon us the fact that the Yoredale beds of Ireland are more inti- 
mately associated with the Millstone Grit and Gannister beds than 
with the Carboniferous Limestone; hence his reason for proposing 
the new classification. In England the Yoredale beds are most 
closely allied to the Carboniferous Limestone, upon which they con- 
formably rest. 

Mr. Hull minutely describes the main features of the Southern 
and Northern Irish Coal-districts and their subordinate Coal-fields. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 201 


In the southern coal-fields of Carlow, Queen’s County, Kilkenny, &c. 
the succession of the strata above the Carboniferous Limestone is 
remarkably uniform and constant; any one may be taken as the 
type of the whole, having reference to an English standard for 
comparison *, 

The Northern Coal-district in Ireland includes the Leitrim and 
Tyrone Coal-fields, which are representative of the southern in all 
their successive beds and stages. Isolated geologically occurs the 
Ballycastle Coal-field of Co, Antrim, the Irish representative of the 
Clyde Coal-basin, of Lower Carboniferous age, a conclusion estab- 
lished and confirmed through its fauna. The Carboniferous Lime- 
stone here is only a few feet in thickness, its place being taken by 
sedimentary or mechanical deposits. This is paralleled in North 
Britain ; but the same beds thicken to the south-west in Derbyshire 
and attain a thickness of 5000 feet of solid limestone. 

The English and Scotch Carboniferous districts and their respective 
coal-fields receive from Prof. Hull terse but careful analysis in his 
paper, each coal-field being divided into recognized stages varying 
from A—G, or from the Calciferous Sandstone series of the Scottish 
beds (A) to the Upper Coal-measures of any coal-field (G). 

I deem it important for my analysis of the Carboniferous system 
to state concisely the subdivisions or stages into which they are 
divided. I adopt the classification clearly given by Professor Hull, 
which is mainly that of the English, Scotch, and Irish surveys. I 
omit the physical characters of the beds, reference to his paper being 
better +. 


The British Carboniferous Series in descending order, with 


localities. 
Names of Formations. Localities. 
(Stage G. Upper Coal-measures. Manchester, Stoke-on-Trent, 
Thin coal-seams and lime- Newcastle-under-Lyne, 5S. 
| stones. part of Dudley Coal-field ; 
Fossils (freshwater or ma- banks of the Dee near 
i rine): Fish; Crustacea, Ruabon; Hamilton and 
Essentially | Cythere inflata; Annelida, Ayrshire, Scotland. 
Freshwater | Spirorbis carbonarius. 
and Stage EF. Middle Coal-mea- Central portions of all the coal- 
Estuarine 4 sures. 'Thick coals. fieldsof England and Wales ; 
Beds. Fossils (freshwater or es- Upper Ooal-measures of 
tuarine): Fish ; Mollusca, Scotland. 


Anthracosia, Anthraco- 
mya; Crustacea, Beyri- 
chia, Estheria; Annelida, 
Spirorbis. Marine species 
rare, 


alco 


* Prof. Hull describes the Castlecomer and Killenaule Coal-fields. Vide 
explanations of Sheets 136 and 137 of maps of the Geological Survey of Ireland ; 
also the Slieveardagh Coal-field, Co. Tipperary. 

t Quart. Journ. Geol. Soc. vol. xxxiii. pp. 613-616. 


202 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


British Carboniferous Series §¢. (continued). 


Names of Formations. 


( Stage E. Gannister Beds, or 
Lower Coal-measures. 
Thin coals with hard sili- 
ceous floors; flagstones 

and shales. 

| Fossils (marine): Fish (mi- 
gratory); Mollusca, Goni- 
atites, Discites, Orthoceras, 
Posidonomya,Monotis, Ari- 
culopecten, Anthracosia, 
Lingula, &e. 

Stage D. Millstone- Grit Series. 
Coarse grits, flagstones, 
and shales, a few thin 
coal-seams. 

Fossils: similar to those 
of the Lower Coal-mea- 
sures. 

Stage C. Yoredale Series. 
Shales and grits, passing 
down into dark shales 
and earthy limestones. 

Fossils (marine): including 
Goniatites, Aviculopecten, 
Ctenodonta, Chonetes, 
Discina, Posidonomya, 
Productus, &e. 


(Stage B. Carboniferous Lime- 
stone. Massive limestone 
in many beds with inter- 
vening shales and grits 
(thick in the south, thin 
in the north). 

Fossils: Fish, Crustacea, 
Crinoids, Corals, &e.; all 
marine. 

Stage A. Lower Limestone 
Shale and Calciferous 
Sandstone. Dark shales 

| in some places; grits, 

conglomerates, and red 
| sandstone and shales in 
the northern districts. 

| Fossils (marine). 


Basis. Upper Old Red Sand- 
stone. Yellow sandstones 
and conglomerates. 

Fossils (freshwater): not 
well represented in Eng- 
land. 


Essentially y 


Marine. 


OOO 


Kssentially | 
Marine | 
(except 
Stage A 4 

in 

Scotland). 


Localities. 


South Lancashire, North Staf- 
fordshire, Nurth and South 
Wales. 


Uplands of Yorkshire, Lanca- 
shire, and Derbyshire, N. 
Staffordshire, and N. and 
S. Wales. 


Uplands and valleys of Lanca- 
shire, Yorkshire, Derby- 
shire, North Staffordshire, 
Wales, &e. 


North and South Wales, Derby- 
shire, Yorkshire, Cumber- 
land; in Scotland the 
Lower or Main Limestone. 


South Wales, Gloucestershire 
and Somersetshire, Nor- 
thumberland and Durham ; 
in Scotland, Calciterous 
Sandstone series. 


South Wales, Northumberland, 
Scotland (Dura Den), Ire- 
land (Kiltorcan). 


The above stages contain, in all, about 500 genera and 2400 


species. 


I purpose showing their numerical value and relation one 


to the other in the passage of the species through the 8 horizons 
or stages to the close of the Carboniferous period. The connexion 
of the Permian rocks with the Carboniferous is not an easy task, 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 203 


small as the fauna is known to be; but I-shall reserve this as the 
basis of my next address, if permitted to do so. 

The recent work of the Geological Survey has proved that the 
greater part of the Calciferous Sandstone of Scotland is the equivalent 
in time of the.true Carboniferous Limestone of England. Most of 
the Carboniferous Limestone of Scotland is the equivalent in time 
of the Yoredale rocks of Yorkshire; so also is most of that 
part of the Carboniferous Limestone series of Northumberland 
from which lists of fossils have been published. In fact the terms 
Calciferous Sandstone, Lower Limestone Shales, Carboniferous 
Limestone, and Yoredale Rocks must really be regarded as repre- 
senting conditions of depth and deposit, and not, when comparing 
different areas, as representing relative age. I have had great 
difficulty in dealing with the Yoredale species, as well as those 
from the Lower Limestone Shales, arising from the terms ‘‘ Lower 
Limestone” of the north of England and Scotland and Lower 
Limestone Shale of the South-west of England and South Wales not 
being always applied to the same horizons by different authors. 
Physically in South Wales, Gloucestershire, and Somersetshire they 
can be definitely traced and mapped, carrying with them an unmis- 
takable suite of fossils, differing essentially from the Great, or Scaur, 
or Carboniferous Limestone above; entirely destitute of many 
zoological groups, especially the compound Actinozoa, the Bryozoa, 
and the Cephalopoda; abounding in fish-remains, chiefly Placoidei 
(teeth and spines). 

The comparison of the Northumberland, Durham, Yorkshire, - 
and Lancashire rocks amongst themselves, and further correlation 
with the Scotch series, has yet to be effected; but the three groups 
in the table at p. 226 (viz. the Lower Limestone shales, the Carbo- 
niferous Limestone, and the Yoredale rocks) present, as nearly as 
possible, the census of species as now received and recorded. 

In the Pendle and Clitheroe area the Yoredale rocks are of great 
thickness, 3000-4000 feet ; they are scarcely represented in Scotland, 
although so near; whilst the great or ‘“‘ Scaur-limestone” of Derby- 
shire is represented in Northumberland. The probable arrangement 
and agreement of the North-of-England and Central-Scottish stages 
have been arranged or correlated by Mr. Hull, through a tacit 
agreement with Professor Geikie and Mr. Lebour, both on strati- 
graphical and paleontological grounds. The stages are as follows :— 


North of England. Central Scotland. 


Stages HE. Gannister beds. EK. Slaty-black-band series. 

D. Millstone Grit. D. Moorstone-rock series. 
“Great Limestone.” Upper Limestone series. 

C C (Yoredale beds.) 

’ | Flagstone and Shales. * ) Lower Coal-and-Ironstone 
series. 

B. “ Scaur-Limestone series.” B. Lower Limestone series. 
Sometimes absent. ; t : 

A. { («“ Tuedian.”) \ A. Calciferous Sandstone series. 


Mr. Lebour, in corroboration of this, compared lists of fossils from 
the “ Great Limestone” of Northumberland (‘‘the most marked 


204 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Yoredale bed”) with the prepared Scotch list, and found that 32 
species Were not known in Scotland, and that about 60 species ran 
through or were common to the three Scotch divisions, and also 
that 28 were found in the Scotch upper and middle series only, 10 
being found only in the lower series. This community of 28 species 
corroborates Mr. Hull’s views, who believes that geographical posi- 
tion, through an interposed Silurian barrier, separated the Scotch 
and English marine areas. ‘There can be no doubt also that over 
the whole of the north of England and the borderland of Scotland 
shallow-sea, estuarine, and land conditions largely prevailed during 
the earlier, if not the middle, Carboniferous stages. It is by such 
conditions that correlation is rendered difficult over extended areas; 
and only by patient, long-continued work with a definite end or pur- 
pose, changes both on the dip and strike being carefully mapped 
or recorded, can the subdivision of one region be actually compared 
with that of another. IJtis from this want of uniformity in working 
that various horizons are adopted for the same beds by different 
authors, and that ranges of fossils are made to differ also. A very 
important table is appended to Mr. Hull’s paper, giving the census 
or known fauna and vertical range of the marine genera and species 
of the Gannister beds (stage E). All these species are embodied in 
the numerical estimate in my own table. 36 genera and 69 species 
are enumerated by Mr. Baily; all the genera and 40 species ascend 
upwards or come from the underlying Carboniferous Limestone. 
Mr. Baily determines that only 6 species of the 69 pass to Stages F 
and G. They are Goniatites Listerr, G. Looneyt, Conularia quadri- 
sulcata, Aviculopecten papyraceus, Discna nitida, and Productus 
scabriculus. No marine shells occur in the Upper Coal-measures in 
the table, Spirorbis carbonarius and Anthracosia only characterizing 
the Stage G. Baily also states that 18 species are peculiar to Stage 
E, or the Gannister beds *. 

Professor Hull in his paper gives a table of the “ Continental 
equivalents of the British Carboniferous divisions,” and a table also 
of ‘‘ Representative Carboniferous formations,” both of great use in 
classification. A fourth table is added, showing the vertical range 
of the marine genera and species of Stage F (Middle Coal-measures). 

Prant#.—Large as the Devonian flora of America appears to 
have been, it bears no comparison with the succeeding Carboniferous 
group, either in America, Britain, or HKurope. 95 genera and 
about 320 species constitute the whole of the known Devonian 
Plante, through the universality of their distribution. That much 
of the Carboniferous flora was derived from the Devonian is evident. 
Of the 84 British Carboniferous genera, 36 occurred in the De- 
vonian, and these are also the most typical or marked in the 
Carboniferous system. 

We owe our chief knowledge of the Devonian and Carboniferous 
flora of North America to Dr. Dawson, of Montreal, Vanuxem, 
Hartt, Rogers, Lesquereux, Newberry, and others, who have largely 
added to the literature of the fossil flora of the American Continent. 

* Quart. Journ. Geol. Soc. vol. xxxiii. pp. 613-651. 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 205 


The British Devonian flora is small, including only 12 genera and 
18 species, as stated in the Devonian table. The American 
genera and species of the Carboniferons flora number, according to the 
published lists &c., 94 genera and 695 species; and the European 
Carboniferous flora, including the British, 176 genera and 1370 
species ; the species occurring in the 8 divisions of our Carboniferous 
strata number 339, illustrating 84 genera. The stratigraphical 
distribution is as follows :— 


Calciferous Sandstone ...... 10 genera and 12 species. 
Lower Limestone Shale .... 22 ze 44, 
Carboniferous Limestone.... 13 Se DAY nce 
Moredale Rocks ......... HONEY p55 none ,, 
Malistone Grit: is... ce 3 6 ws 10 = 254s 
Lower Coal-measures ...... 54 ss A i lege Ps 
Middle stele. (uteri bn hide 34 = 1 
Upper ce i eee ee 33 a 168 Fx. 


Ten genera are largely represented in Britain, and contain 192 
species out of the 339, forming the whole flora: they are Alethopteris 
9 species, Asterophyllites 11, Calamites 14, Lepidodendron 21, Lepido- 
strobus 15 (but for the number given, these cones of Lepidodendron 
should be taken as that genus ; probably we may never know to what 
species of Lepidodendron all these Lepidostrobi belong), Neuropteris 
23, Pecopteris 28, Szgillaria 27, Sphenopteris 32, Ulodendron 12. All 
the Alethopterides are from the Coal-measures. The Asterophyllites, 
with two exceptions (A. foliosus and A. longifolia), are also Coal- 
measure fossils. The Calamites range from the Calciferous Sandstone 
to the Upper Coal-measures—4 species, C. canneformis, Schloth., 
C. cultranensis, Haught., C. dubius, Artis, and C. Lindleyz, Sternb., 
occurring in and below the Carboniferous Limestone; the remaining 
10 are Coal-measure forms and mostly belong to the Lower series. 
The 21 species of Lepidodendron, with 5 exceptions, are all from 
the Coal-measures, as are 13 of the 15 Lepidostrobi; 2 species 
(L. comosus, L. & H., and L. variabilis, L. & H.) are Lower Limestone 
Shale species. With one exception in the genus Newropteris (NV. 
Toshi), the species are from the Coal-measures ; although we know 
that NV. cordata, N. flexuosa, and NV. gigantea are Millstone-Grit 
forms. No Pecopteris is known below the Lower Coal-measures ; 
therefore the large number of species (28) are essentially Upper 
Carboniterous. Of the 27 species of Szgillarza only 2 occur below the 
Lower Coal-measures, viz. S. dichotoma, Haught., and S. pachyderma, 
Brongn., which occur in the shales at the base of the Carboniferous 
limestone. 3 species of Sphenopteris (S. affinis, S. linearis, and S. tri- 
foliata) range from the Calciferous Sandstone to the Coal-measures ; 
4A others (S. bifida, L. & H., S. crassa, L. & H., S. elegans, Brongn., 
and S. obovata) occur in the Carboniferous shale and limestone; 
the remaining 25 species irregularly range through the Lower, 
Middle, and Upper Coal-measures. Ulodendron minus and U. par- 
matum range from the Carboniferous shale to the Upper Coal- 


206 PROCEEDINGS CF THE GEOLOGICAL SOCIETY. 


measures. U. Landleyanum, U. majus, and U. minus are the Mill- 
stone-Grit species. The above 10 genera were selected on account 
of their being largely represented specifically. Regarding the 
distribution of the Carboniferous flora, we know that many species 
occurring in Pennsylvania and Nova Scotia are identical with Euro- 
pean and British species. This points to a greater similarity and 
equality of climate than now prevails, and also to the possible con- 
nexion of the coal-hearing areas of Europe with that of America, 
either by continuous land or by groups of islands. 

It is difficult in all cases to determine from the often somewhat 
vague descriptions of authors the true geological horizon of the 
many species in the true Coal-measures, or whether they are 
Upper, Middle, or Lower Coal-measure forms. Every care, however, 
has been taken to determine this when possible, not only for the 
Plantee but for the whole fauna also. 

Protozoa: SponerpaA.—The 10. genera and 15 species known 
are confined to the Carboniferous Limestone and shales below. 
The genus Paleacis of Haime (1860) now embraces Phillips’s, 
Hydriopora, M‘Coy’s Astreopora, Seebach’s Paleacis, and the genus 
Sphenopoterium of Meek and Worthen. ‘The 3 British species, 
P. cuneiform, J. Haime, P. cyclostoma, Phill., and P. obtusa, Meek 
and Worthen, have a wide geographical range, being almost uni- 
versally distributed ; but none of the 10 genera and 15 species have 
occurred above the Carboniferous Limestone in any area. 

Protozoa: FoRAMINIFERA.—14 genera (11 belonging to the Im- 
perforata and 3 to the Perforata), with 48 species, range through the 
lower portion of the Carboniferous group, but none have yet occurred 
higher than the Yoredale rocks, where 7 genera and 18 species are 
. found, and only 1 genus (Stacheia) is known in the Calciferous 
Sandstone ; 6 species illustrate Stacheia, and all range from the Cal- 
ciferous beds to the Carboniferous Limestone. Our knowledge of 
this order (Reticularia) of the Rhizopoda in the Carboniferous rocks is 
due to the labours and researches of Mr. H. B. Brady. 40 species are 
known from the Carboniferous Limestone. 43 of the 48 species in 
these rocks are named and described by Mr. Brady in his learned 
monograph upon the British Carboniferous and Permian Fora- 
minifera in the volume of the Paleontographical Society for 1876. 

Hyprozoa.—Arbusculites and Palwocoryne are the only 2 known 
genera in this class. <Arbusculites argenteus, with Palwocoryne 
radiata and P. scotica (all three from the Carboniferous Limestone), 
complete the list. 

ActinozoA.—Both the Tabulata and Rugosa are well repre- 
sented in the Carboniferous rocks, especially the Rugosa. 21 
genera fall under that order, and 14 in that of the Tabulata. 
The genus Cladochonus of M‘Coy (Jania of the same author) and 
Aulopora are the only representatives of the Tubulosa. Aulopora, 
which is probably the young state of Syringopora, usually referred 
to the Tubulosa, must be placed with the Tabulata if so deter- 
mined. 9 families, 36 genera, and 141 species constitute the 
Actinozoal fauna. The only 2 families of numerical value are the 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 207 


Favositide, with 5 genera, and Cyathophyllide, with 20; the 
remaining 7 families have only 1 or 2 genera each and but few species. 
No form of Coral is known above the Yoredale rocks, and only 
2 genera and 2 species occur in the Yoredale series, viz. Mavosites 
tumidus, Phill., and Zaphrentis Phillipsu, M.-Edw. The Calciferous 
Sandstones yield 3 species, Alveolites septosus, Fleming, Mavosites 
tumidus, Phill., and Lithostrotion junceum, Fleming. 

The Lower Limestone Shales (mostly those of the west of England) 
have yielded 13 genera and 30 species; the nature of the deposit 
greatly influenced and determined the presence of the species. 
Nearly all the Corals in the shales are simple forms and dwarfed 
in habit, certainly more so than those species that lived in 
clearer and deeper water, where lived most of the compound 
forms which occur so abundantly in the limestone beds. This 
difference, under petrological association, is manifest on carefully 
examining the stratigraphical position of the corals in many locali- 
ties. The Carboniferous Limestone is coralliferous throughout, 
and the fullest evidence may be obtained of large and extensive 
reefs composed essentially or mainly of the Rugosa, Lithostrotion, 
Lonsdaleia, species of Cyathophyllum, and numerous other species. 
The Carboniferous Limestone of the British Islands yields every 
known genus and species catalogued; in other words, 36 genera 
and 141 species (all the Carboniferous Actinozoa known) are found 
in the Carboniferous Limestone in one locality or other in Great 
Britain and Ireland. The whole class culminates in the Mountain 
Limestone, for only 2 genera and 2 species pass to the Yoredale beds 
(favosites tumidus and Zaphrentis Phillipsu). This sudden ter- 
mination in time of the Actinozoa was undoubtedly due to those 
physical changes which took place in consequence of the elevation of 
the sea-bed and prior to the deposition of the Permian. Our lists, 
tables, and collections of the Carboniferous Corals have not been 
studied sufficiently to enable us to construct proper distribution 
tables ; but the accompanying table of 19 of the chief British genera 
is intended to show their geographical distribution in Britain, also 
in the chief locality in Europe, and further their correlation with 
America. 


208 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Table showing the numerical value of the species in the chief genera 
of Actinozoa in Great Britain, Belgium, and America, the 
former for European, the latter for American comparison. 


{D) 

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tg = : ; 3 5 

eH el |= 

eS le | ee ae 

el ee gia | aes 

Amiplexusien ees daoe teaetececlerads 6 “ 4 12 9) 21 
JMO) Nay AblTIO Goo sscnasqaceanosadoc 2 2 2 wae ie 6 
@hestetesy vssyemeecscatceercs sence 4 2 4 2 3 15 
Cladochonuseaereercan cect 4 a 2 2 Ms 8 
Clisnoyolaylknwan » .sssoondacodosdaoose 4 5 4 3 O0 16 
Cyathaxomlanecoseuosncasccrea es 2 1 a 3 6 12 
Cyathoplayllunayeeasceeseee ea ilo 5 6 7 3 32 
Cycloplivlllmmam ena cence 2 2 1 235 ee 5 
I AVOSICES Ie lacersus as seleciene a match: 6 i 3 5 3 18 
leterop nwvilliaige cee reece a eee 2 6 ss shee ee 8 
JUNI NOSHROELO sonsccosnaeccoossn0s 16 10 12 4 5 47 
Thonsdaleran ii wissen nese 4 2 ay 1 1 8 
Miche liniajaeeepeeccseeeee tate 4 2 5 4 1 16 
Pallgeacisth veces asus coctetlestane 1 2 ae 2 ~cieie tite 
ehilllijpsastreca ycpemencdee csc: 2 cS. ie Hf Bee 2 
SH ABUTYERO) COTES) BGs anasaeodoodsqunanesqs 5 3 6 5 4 23 
ZL PATemuis ie se acces sce eneesce es 8 10 5 21 18 62 
83 57 54 al 49 | 304 


Ecuinoprrmata.—tThe class Echinodermata is represented in the 
Carboniferous rocks by 4 orders—the Crinoidea, the Blastoidea, the 
Perischoechinoidea, and the Echinoidea. The Holothuroidea through 
the Synaptz occur also in a few localities. 30 genera and 163 
species constitute the entire fauna, and the whole are in the Car- 
boniferous Limestone division. Only 1 species (Archeocidaris Uret) 
seems to be known in the Calciferous Sandstone. No species 
appears in the Yoredale rocks; indeed the entire class ceases to 
appear above the Carboniferous Limestone, the 5 succeeding 
divisions being utterly void of this class and the Coelenterata. The 
Lower Limestone Shales through 11 genera yield 31 species—Actino- 
crinus 5, Archeocidaris 3, Astrocrinus 1, Atocrinus 1, Cyathocrinus 2, 
Palechinus 3, Platycrinus 10, Poteriocrinus 3, Rhodocrinus 2, and 
Taxocrinus 1 species. Only 1 of the above is special or confined 
to the Lower Carboniferous Limestone of Carlops, Peeblesshire, 
Astrocrimtes (Zygocrinus) Benner, R. Ether., jun. 

The characteristic genera, and those containing the largest 
number of species, are :—Actinocrinus 21 species, 5 of the 21 occur- 
ring in the Lower Limestone Shale ; Cyathocrinus 10 species, 2 of 
the 10 are also Lower Limestone Shale; Platycrinus 25 species, 
10 of which are in the Lower Limestone Shale ; /hodocrinus 12 
species; and <Archwocidaris 10 species, The order Blastoidea, 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 209 


through the Pentremitide, 16 species, and Pentaphyllum 1 species, 
are all confined to the Carboniferous Limestone proper. The sin- 
gular group of the Perischoechinoidea, with Palwchinus 7 species, 
Perischodomus and Melonites 1 each, also essentially characterize the 
Carboniferous Limestone proper. 

ANNELIDA.—4 species of this class, belonging to 13 genera, wander 
through the 8 divisions. Spzrorbis numbers 14 species and Serpula 6; 
the remaining 11 genera have only 1 or 2 species each. Serpulites 
carbonarius, Spirorbis pusillus, S. caperatus, and S. helicteres are the 
only 4 species that range above the Carboniferous Limestone and into 
the Coal-measures where the argillaceous shales and clays are of 
marine or estuarine origin. The Calciferous Sandstones of Scot- 
land, through the researches of Bennie, have yielded Serpulites 
carbonarius, Spirorbis Kichwaldi,S. globosus, S. helicteres, S. pusillus, 
and 2 varieties. 16 species occur in the Lower Limestone Shale ; 
10 genera and 24 species in the Carboniferous Limestone. Serpulites 
membranaceus is the only Yoredale form ; and 4 species, Crossopodia 
Embletone, C. media, Serpulites carbonarius, and Spirorbis pusillus, 
occur in the Millstone Grit ; and Serpulites carbonarius is the only 
Lower Coal-measure form, also associated with Arenicola carbonaria. 
The same 2 species are Middle and Upper Coal-measures, which 
exhausts the Annelidan fauna. 

Crustacua.—The remarkable Crustacean fauna that occurs in the 
Silurian rocks numbers about 320 species, chiefly Trilobites, all of 
which, with the exception of 5 or 6 genera, died out entirely at the 
close of the Silurian period. The Crustacea of the Old Red Sandstone 
belong chiefly to the order Merostomata—Hurypterus, Pterygotus, and 
Stylonurus being the characteristic genera. The Trilobita, comprising 
only 6 genera (Bronteus, Cheirurus, Harpes, Homalonotus, Phacops, 
and Phillipsia), with 11 species, are Devonian, no Trilobite occurring 
in the Old Red proper. The Carboniferous system in Britain yields 
35 genera and 225 species, 130 of which are Ostracoda, 59 Phyllopoda, 
2 Merostomata, 6 Pcecilopoda, 2 Stomapoda, 5 Macrura, and 13 Tri- 
lobita, illustrating the 3 genera, Brachymetopus (2), Griffithides (5), 
and Phillipsia (6), which group entirely disappears with the close 
of the Carboniferous Limestone. Nowhere in Europe or America 
have we evidence of the continuity of this group above the rocks 
named. Beyond the Ostracoda and Phyllopoda, which constitute 
the 2 important orders in the Carboniferous rocks (together num- 
bering 189 species out of the 225 known), there is little to discuss. 
The genera of Ostracoda number 85, the Phyllopoda 8, the Trilobita 
only 3. 

The CatcrrErovs Sanpstone (Tuedian) has yielded 9 genera and 
20 species, mostly Ostracoda, and 2 species of Anthrapalemon, the 
first known appearance of the group or tribe Macrura. (38 other 
species of the same genus occur in the Coal-measures.) 

Lower Limxstonn Suates.—17 genera and 48 species, also chiefly 
Ostracoda and Phyllopoda, are individually abundant in the Lower 
Carboniferous beds (below the Carboniferous Limestone). 

CaRBONIFEROUS LimEstoNE.—159 species are recognized and de- 


210 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


scribed from the Carboniferous Limestone proper, illustrating 28 
genera out of 33 (or nearly every occurring genus is represented), 
and the above number of species out of the known 225. The 
4 genera wanting in the Limestone are Anthrapalemon, Belinurus, 
Pygocephalus, and Candona. The Ostracoda number no less than 
15 genera and 107 species; the Phyllopoda 8 genera and 36 
species ; the Trilobita 3 genera and 8 species. 

YorEDALE Brps.—The Ostracoda through the genus Bairdia 
(6 species) and Kirkbya (2); the Phyllopoda through Beyrichia (5), 
Estheria (1); and Griffithides mucronatus (Trilobita), complete the 
Crustacean fauna of this horizon. 

Mittsrone Grir.—No Crustacean remains known. 

Lower CoaL-mMEasuRes.—13 species of Ostracoda, 8 of Phyllopoda, 
and 4 Peecilopoda, with 3 species of the genus Anthrapalemon, a 
Macrurous Decapod, 2 species of which first appeared in the 
Calciferous Sandstone (A. Maconochii, Ether., and A. Wood- 
wardi, Ether.). A. dubius, Prestw., A. Grossarti, Salt., and A. Rus- 
sellianus, Salt., belong to the Lower and Upper Coal-measures. This 
is the earliest known appearance of the order Decapoda. This ancient 
type has been found both in the Coal-measures of Europe and 
North America. Anthrapalemon is related to the living Galathea. 
Of the suborder Xiphosura or Pecilopoda we possess 2 genera, 
Bellinurus and Prestwichia, this last allied to Neolimulus of the 
Upper Silurian. The Penny-stones of Coalbrook Dale yield Prest- 
wichia rotundata (Limulus). All the thoracic and abdominal seg- 
ments in Prestwichia are ankylosed. The earliest appearance in 
time or commencement of existence of the Xiphosura is in the 
Ludlow rocks; no Xiphosuran has yet been detected in the Deyo- 
nian. Pygocephalus Cooperi, Huxl., and P. Huxley, Woodw., occur 
in the Lower Coal-measures. These Podophthalmatous Stomapods, 
with the Decapod Anthrapalemon (3 species), carry back in time 
the higher order of the Crustacea. 

MippLe Coan-mEAsvrEs.—Anthrapalemon dubwus, Bellinurus bel- 
lulus, Konig, Beyrichia arcuata and B. Binneyana, Hstheria Bei- 
nertiana and H. striata, are all the Crustacea known in the Middle 
Coal-measures. 

Upper Coat-mMrasvrEs.—10 genera and 16 species compose the 
Crustacean fauna of the Upper Coal-measures—Ostracoda 6, Phyllo- 
poda 7, Poecilopoda 1, and Anthrapalemon Grossariv. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 211 


Number of Species in the six Paleozoic orders of the Crustacea 
occurring in England, Scotland, and Ireland; also in Belgium 
and America. ‘The two latter areas for comparison as betore. 


cs} se i FS 
| q oS =| Ss 
Orders. a = 2 5 o 

cold) | 
S @ oO S 
ey oD) = aa < 
SRI OMG ta acacs afelscecawicae odes ead 45 68 38 21 9 
HSPN O POCA) acs. ctdenra coe ecscess: Ils} = 48) 14 9 11 
PREMMOSUMA Fi t)44 class seadeeceeessee ees 8 1 5) Bs 2 
Bina AbOTIG A! 4). 5.0550. 0cs- accesses eae 1 2 che se Ze 

Ble GAMOU A) cp ton sae cease sinc sobiesnias 1 7 

Tsong eee ee ee 18 7 14 15 26 
ae g5 |il4 | 71 | 45 | 50 


Bryozoa.—The Carboniferous Bryozoa as a group constitute by far 
the largest series in any division of the Paleeozoic rocks. 77 species 
range through the 3 lower horizons of the Carboniferous series, 74 
belonging to the true Carboniferous Limestone, 28 to the Lower Lime- 
stone Shales or Lower Limestones, and 4 to the Calciferous series ; not 
a single species passes to or occurs in the Yoredale, Millstone Grit, or 
either one of the three divisions of the Coal-measures. The whole 
group essentially belongs to and characterizes the Calcareous rocks 
and shales at the base of the formation; the large genera are— 
Ceriopora 5 species, Fenestella 26, Glauconome (Acanthocladia) 8, and 
Polypora 8. 20 other genera of great zoological value occur, but the 
species are few in each genus(1 to 4). Actinostoma, Carinella, Dras- 
topora, Gomoclacia, Hemitrypa, Ptilopora, Rhabdomeson, Synocladia, 
and Vineularia are important genera in the Carboniferous rocks, 
and distinctively determine the age of the beds in which they occur. 
Be it remembered that none of the Paleozoic genera live on or 
extend into the Mesozoic period, and the Fenestellide become 
extinet in the Permian rocks. 24 genera and 77 species at present 
represent the fauna of this division of the Molluscoida; doubtless 
the species will be much reduced through extended research and 
eritical examination*; but as Mr. Shrubsole is carrying on research 
in the Fenestellide, and Mr. Vine investigating the Diastoporide, it 
is better to give the census of the species as they now stand without 
incomplete modification. Mr. Shrubsole’s last paper is not yet pub- 
lished; I will therefore notice their distribution, subject to the 
anticipated work of these two authors. 

CaLCIFEROUS SANDSTONE.—Archeopora nexilrs, Cerropora similis, 
Fenestella Morrisui, F. plebera, F. tuberculato-carinata, and Glauco- 


* Vide the able papers by Mr. Shrubsole, F.G.S., and Mr. G. R. Vine in the 
‘Quarterly Journal of the Geological Society,’ vel. xxxvi.; also Vine, ‘ Report of 
the British Association,’ 1880, “On the Carboniferous Polyzoa,” for important 
matter relative to the history and classification of the Polyzoa. 


212 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


nome gracilis appear to be all known in these lowest beds of the 
northern Carboniferous rocks. 

Lower LimestonE AND SHALES.—13 genera and 26 species illustrate 
these shaly limestones at the base of the Carboniferous; the 3 
chief genera are—Venestella 12 species (far too numerous), Glau- 
conome 3, and Pustulipora 2 species; the remaining 9 genera have 
only 1 species each; many of these, necessarily, also occur in the 
succeeding Carboniferous Limestone. 

CsRBoniFEROUSs LimEstonE.—72 species occur, belonging to 19 
genera; the most important of the latter are—Fenestella with 
25 species, Glauconome 8, Polypora 8, Rhabdomeson 3, Pustulipora 
4, and Cercopora 5 species. The 13 remaining genera, although 
poor in species, are highly characteristic; amongst them may be 
named Sulcoretepora, Synocladia, Vincularia, Actinostoma, and Dias- 
topora. No species passes to higher strata or above the Carboni- 
ferous Limestone. 

ARACHNIDA.—Only in the Lower Coal-measures have we detected 
the Arachnida, 5 species representing 3 genera—Architarbus sub- 
ovalis, H. Woodw., Hophrynus Prestvicit, Busk, Hoscorpius anglicus, 
H. Woodw., and &. carbonarius, Meek and Worth.; an unde- 
scribed form occurs, which, however, may be H.anglicus. E. carbo- 
narius 1s also an American species, occurring in the Coal-measures 
of Illinois; and Architarbus is also a North-American form. The 
occurrence of two of the three genera and one species in two 
areas so widely separated is significant, with many other facts, of 
the probability of the continuity of land between Britain and 
America during the long period required for the deposition of the 
Coal-measures and the accumulation of coal. 

Myrropopa.—The 2 genera of Chilognathous Myriopoda, Hupho- 
beria and Xylobius, which occur in our Coal-measures are also 
American, and, with the Arachnida, again add strong presump- 
tion to land-continuity, densely covered with that peculiar flora 
known to have flourished during the Coal-measure period. We 
possess 3 species of Huphoberia, viz. HL. anthrax, Salt., LE. Brownia, 
H. Woodw., and £. ferov, Salt. Dr. Dawson’s genus Xylobius, also 
the known American species ‘“ Stgillarue,” lends additional interest 
to the attempt to correlate our fauna and flora of the Coal-measures 
with those of the American continent; for many species of plants 
are identical, and with them associated faunal groups. 

Insecra.—Both the Orthoptera and Coleoptera are represented in 
our Coal-measures. The Mantide through the genus /ithomantis 
of Dr. H. Woodward, the Locustide through Giryllacris, Swinton ; 
these two genera belong to the Orthoptera. The genus Curculiordes 
of Dr. Buckland belongs to the Coleoptera. These all occur in our 
Lower Coal-measures. Their history and zoological affinities have 
been ably described by Dr. H. Woodward, F.R.S., in the ‘ Journal ’ 
of our Society (vol. xxxii.); doubtless every fresh investigation into 
the rocks that hold and yield the extinct flora of our coal-fields will 
bring to light the remains of the three last groups, the Arachnida, 
Myriopoda, and Insecta; they are, or were, intimately associated 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 213 


with the vegetation of the Carboniferous epoch, and, with other 
remains in the same rocks, show that the Atlantic ocean covers a 
vast area of once continuous or contiguous land. 


Eyery one who has studied the varied conditions of the Carbo- 
niferous series, the deposition of its strata, the distribution of its 
fauna, will at once admit that its history is only to be written 
through the great and reliable group of the Mollusca (Brachiopoda, 
Lamellibranchiata, Gasteropoda, Pteropoda, Heteropoda, and Cepha- 
lopoda), whose 6 divisions or classes number 116 genera and 1020 
species, or nearly one half the entire known Carboniferous fauna. So 
many of the genera have come down to the present day, that infe- 
entially, through observation, we know their habits in Carboniferous 
times; they show us that, during or throughout the accumulation 
of the Carboniferous Limestone, moderately deep-sea conditions 
prevailed, and the Molluscan fauna then surpassed all subsequent 
numerical development during the Carboniferous period. Ultimately 
physical changes caused the sea to shallow, thus altering all bathy- 
metrical conditions at the same time: littoral species died out, 
through altered circumstances; and the habits of the deeper-sea 
fauna changed also on slow elevation, the zonal lines of life under- 
going modification through adaptation. Plainly are these modifi- 
cations and changes to be observed through the several stages, from 
the Calciferous Sandstones to the summit of the Coal-measures. 
Zoologically, only the last five stages of the Carboniferous group 
have to be considered, as so great a change took place at the close of 
the Gannister group or Lower Coal-measures, the succeeding beds 
illustrating terrestrial conditions through the rich and earliest exten- 
sive flora known (vide Table at end of Carboniferous system, p. 226). 
No one has felt the difficulty of showing the distribution of life 
through the Carboniferous group more than myself, the rich fauna 
that characterizes the series from the Calciferous Sandstones to the 
top of the Gannister (or Lower Coal-measure) beds being most diffi- 
cult of analysis. 

Every extended sea-bed has its own conditions and its own fauna; 
the exact nature of that sea-bottom it is difficult to predicate; it 
can only be learnt through examination. Admitting that the fauna 
of any group of rocks is, or was, as local as the physical condi- 
tions during deposition, and the maximum of life is usually local, 
it will show itself in any area, in any part of an epoch or of a 
stage; it is governed chiefly by the nature of the sediment and 
by temperature. The questions of appearance of species, duration, 
migration, extinction, recurrence, and many other conditions that 
govern marine life in all its phases are to be studied through a 
careful investigation of the physical history of the Carboniferous 
rocks ; and through no other zoological group do we receive so much 
instruction as through the Mollusca. 

Bracuropopa.—Both the Tretenterata, or “‘ Inarticulata,”’ and the 
Clistenterata, or “ Articulata,” are largely represented in the Carbo- 
niferous rocks; and many genera appear for the first time, and 


VOL. XXXYVII. gq 


214 PROCEEDINGS OF THE GROLOGICAL SOCIETY. 


essentially characterize the lower members of the Carboniferous 
group: such are Hypodema and Syringothyris, neither of these 
appearing in older strata. 

The specific development of Productus has no precedent or parallel 
in any other group of British strata. 44 species occur in the deposits 
from the Calciforous Sandstone to the Lower Coal-measures, 5 species 
in the Calciferous series, 16 in the Lower Limestone Shale, 41 in 
the Carboniferous Limestone, 7 in the Yoredale, 5 in the Millstone 
Grit, 3 in the Lower Coal-measures, 1 in the Middle Coal-measures 
(P. scabriculus), and none in the Upper Coal-measures; the genus 
(but no known Carboniferous species) passes to the Permian rocks, 
and is represented by P. horridus, its last appearance. 

The genus Spirifera attains its highest development in this 
horizon aleor ; 30 species range side by cide by with the Producta; and 
it is well to state also their numerical value in the several horizons : 
Calciferous Sandstone 3 species, Lower Limestone Shale 17, Carbo- 
niferous Limestone 30, Yoredale rocks 6, Millstone Grit 4, Lower 
Coal-measures 4. 2 of the 30 species of Spirfera are Devonian, 
Sp. lineata and Sp. Urii; thus 28 species are new to Britain; and 
no less than 185 species (British, European, and American) have been 
described. The Lhynchonelle of our Carboniferous rocks number 
21 out of the 30 European species; and 18 of them are new; the 
3 derived from the Devonian are 2. plewrodon, R. pugnus, and 
KR. remformis. 

The genus Zerebratula begins here to be numerically abundant, 
no species being known in the Silurian strata, and the Devonian 
rocks yielding only 4 species, whilst 7 species are Carboniferous. 
The great development of this genus in the Jurassic rocks surpasses 
that of all other Clistenterata; in our own area they apparently 
nearly died out or migrated after the Carboniferous epoch, none 
occurring in the Trias. The genus Athyris ranges from the Cal- 
ciferous Sandstones to the Lower Coal-measures ; A. ambigua and 
A, planosulcata are the only species that transgress the Carboni- 
ferous Limestone. The genus Atrypa was formerly recognized as 
a Carboniferous genus with $ 26 species, which are now relegated to 
Rhynchonella, Retzia, and Athyris. Camarophoria, with its 5 species, 
belongs to the Carboniferous Limestone ; only C. crumena passes to 
the Permian, omitting the 5 intermediate stages or horizons. 
Chonetes, only in one instance (under the species hardrensis), ranges 
above the Carboniferous Limestone into the Lower Coal-measures, 
the whole 15 species being otherwise confined to the limestone 
and the shale below. 

The significance or importance of Productus as a Carboniferous 
genus cannot be overlooked when determining through its species 
definite horizons in these rocks. It is ubiquitous; in no region on 
the globe, where the Carboniferous rocks are developed, do we not 
find “this characteristic shell and in vast abundance—in the Polar 

regions, Australia, New Zealand, Van Diemen’s Land, India, Ame- 
rica (in 15 States), throughout Europe, and in Africa. 

The Carboniferous Brachiopoda consist of 20 genera and 175 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 215 


species; and 7 of the 8 horizons possess them: 9 genera and 18 
species occur in the Calciferous Sandstones of Scotland; 11 genera 
and 380 species range through the thin-bedded Carboniferous Lime- 
stone Shales, or the Lower Limestones ; 14 genera and 172 species 
have been collected and named from the Carboniferous Limestone 
alone; 10 genera and 31 species are from the Yoredale rocks and 
Upper Limestone Shales; 11 genera and 24 species are Millstone 
Grit; the Lower Coal-measures yield 9 genera and 18 species, and 
the Middle 3 genera and 3 species. We know of no Brachiopoda in 
the Upper Coal-measures. The whole group, with every species, has 
been exhaustively worked out by T. Davidson, Esq., F.R.S., in the 
volumes of the Palewontographical Society; nothing is left undone 
by that distinguished brachiopodist, whose labours in this field of 
paleontology are universal. To Mr. Davidson we are indebted for 
the highest research into the morphology, history, and classification 
of this most difficult class of Mollusca. 


S ce} 3 a 

Se te ahve | Ae ene 

“eb S Si op g 3 

= S @ oO q 3 

eo oD) i FL <q oe) 

Wamarophoria ..... .....-.-.0.00+. 4 I 3 1 3 12 
(CIMOIEUES) GARR agROsO SHORE EE Cn Ea rOnnE 12 10 13 5 28 68 
aa asl ccfeine suc es atevernasecs 2 2 3 2 3 12 
(CONTTHENT noo SU OSB ESB LORE Eee eee 2 Eve if we 3 
IDET) scShSe algae Ree ee eee ae 2 1 2 11 14 30 
HUMAN s cacecs vcveeecanese 5 4 3 3 8 23 
USHER eae as cel sbeaeeke adlead 7 4 8 8 25 52 
FEPROCHUICEUSW  grced tes eececs otaneine 45 32 28 47 | 109 | 261 
ulaymich Onell se sscce see sencsecees 17 3 11 11 36 78 
IRCIVAG) ” Goh hee pe Cen EeEe eae Se acer 3) I 1 3 12 20 
SUMMON Ree ccaccecvcses-<c seen 40 21 30 48 | 108 | 247 
OPUMECEINA fcc edcc es ccleuer eset ee 3 5 5 3 9 29 
Streptorhynchus .................. 4 7 5 3 10 29 
BRET EMAL UN AN aici. a's cesses credo +a 4. 3 2 8 26 43 
150 94 | 115 | 153 | 891 | 908 


The preponderance of the two genera Productus and Spirifera is 
a great feature in the Carboniferous rocks. Chonetes, Ahynchonella, 
and Orthis follow in importance so far as the number of species lends 
significance. Chonetes and Productus make their first appearance in 
these rocks, and, with Leizia, Streptorhynchus, and Cyrtina, dis- 
appear at the close of the period. 5 genera and 6 species of Bra- 
chiopoda pass to the Permian; they are Camarophoria globulina, 
Phill., C. Schlotheimt, V. Buch, Discina Konincku, Geinitz, Lingula 
Oredneri, Geinitz, Spirifera Clannyana, King, and Spuriferima cris- 
tata; and beyond 2 species of Ostracoda I know no other connecting 
or transgressive species than the above; in other words, only 8 
species of all orders pass to higher rocks out of the Carboniferous 

q 2 


216 . PROCEEDINGS OF THE GEOLOGICAL SOCTETY. 


system. Even the Rhizopoda (Foraminifera), which have so wide a 
distribution in time, occur not beyond the limits of this formation. 

Lamerirsrancuiata.—Monomyaria*. This group and the Dimy- 
aria united number no less than 54 genera and 415 species. The 
Monomyaria, up to the present time, are represented by 10 genera 
and 179 species, 103 of which belong to the genus Aviculopecten, 28 
to Avicula, Pinna 6, Posidonomya 12, Pterincea 4, Pteronites 9, Stre- 
blopierra 3, Gervillia 2, Inoceramus 4, Loma 2, and Pinna 6. The 
Calciferous Sandstones of Scotland and Northumberland have yielded 
3 genera (Avicula, Aviculopecien, and Pteronites) and 13 species; 
the Lower Limestone Shales, or Lower Limestones, 5 genera and 64 
species; the Carboniferous Limestone 11 genera and 151 species; 
the Yoredale rocks 4 genera (Avicula, Aviculopecten, Pinna, and 
Posidonomya) and 14 species; the Millstone Grit 3 genera (Avcula, 
Aviculopecten, and Posidonomya) and 11 species; the Lower Coal- 
measures 3 genera (the same) and 13 species; 4 species of Aviculo- 
pecten occur in the Middle Coal-measures ; and 1 (Aviculopecten papy- 
raceus) is the only form in the Upper Coal-measures. The species 
having the longest range are Aviculopecten alternatus, A. gentilis, A. 
granosus, A. papyraceus, A. quadratus, A. scalaris, and A. variabilis, 
Posidonomya Becheri, P. lateralis, and P. membranacea. The species 
of the genera Pteronites, Pterinceea, Streblopterra, Posidonomya, Pinna, 
Avicula, and most of the Aviculopectines are chiefly from the Car- 
boniferous Limestone. We may assume that the largely represented 
eroup of the Aviculopectinidz possessed much the same bathymetrical . 
range as that of the genus Pecten in our modern seas; the same with 
Pinna and Posidonomya. Indeed it would appear, from the per- 
sistency of the vertical range of the group Monomyaria, that no 
general depth governed their habit ; for only 14 species out of the 178 
range above the Carboniferous Limestone and Yoredale beds. 10 are 
named above ; and the following 4 may complete the species—Avicu- 
lopecten fibrillosus,A. obtusus, Posidonomya Gibsoni,and Pinna spatula. 
The Table also expresses the number of species that are in the Mill- 
stone Grit and Coal-measures included here. The magnitude of the 
Monomyarian group is sufficient reason for treating it separately, 
especially considering that one genus alone (Aviculopecten) is repre- 
sented by 103 species. 


* T retain this term here, including in it only the Asiphonida, represented by 
the Ostreidee, Aviculide, and Pectinide. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 217 


| 
~ | : 
Gen MEE < A S 
S S a 1] ay 
Beco ties mi <4 
A STCITT S E Se Oa 2 ed | 
Piriatilen ete Re ie 25h dg 16 10. i 2S ulna ni moe: 
EVICBIGNECEEM | conc cne-neneeecse> 2 48 50 80 | 33 40 
PD YETSTTLTIES Re ae + its 2 lsc 5 
IRRECCEATINUS, cc cteccs cs icc sciessecest ae 4 1 1 
LUTTE O) Seinen i OC 2 | nef} Al 
LEGO. GOk ae eRe Eee eee , 1 ee ees 
"PATER Rtn aae ene ae 5 5 3 | 6 
EGRIGOHOMYS  ....,..0.sncereceese 8 2 8 5 J. 
ETRE et ecm cccoes snes cuecete i x2) aeelitysee 9 
PETEPOMICES Sones ec sscsccbcveccotess 3 6 7 be 2 


85 74 | 186 67 


pa 
eS 
Oo 


The preponderance of species in one genus, as in Aviculopecten, 
is unexampled in any other British formation. 150 species occur in 
Britain and Europe; 50 have been described from America, and all 
differ from our forms. The species occurring in the Belgian Car- 
boniferous rocks are known to us through the researches of De 
Koninck. In Ireland, mainly through the researches of M‘Coy 
and Baily, the Integropallialia appear to have been exhaustively 
recorded. The genus Avicula, in the Irish Carboniferous rocks, 
numbers about 23 species, Aviculopecten 80, Pecten 5, Pteronites 5, 
Posidonomya 8, Pierinea 3, Pinna 5. Scotland: Avicula 10, Avicu- 
lopecten 50, Pinna 4, Posidonomya 2, Pteronites 6. England: Av- 
cula 16, Aviculopecten 48, Pinna 5, Posidonomya 8, Pieronites 3, and 
Pterinea 1. These comparisons are important, as showing geogra- 
phical distribution as well as age. The collective fauna illustrating 
this group in all Kurope beyond Belgium does not exceed 50 species, 
showing either want of research or the small development of the 
Carboniferous rocks east of Rhenish Prussia. 

LAMELLIBRANCHIATA.— Dimyaria*.—No less than 43 genera and 
245 described species occur in the Carboniferous rocks, the largest 
number being in the Carboniferous Limestone (80 genera and 182 
species); the underlying Lower Limestone and Shales yield 23 genera 
and 103 species; and the Calciferous beds of the north of Kngland 
and Scotland 17 genera and 38 species; only 9 species are Yoredale. 
Many species are necessarily common to the three lower horizons ; 
but the faunal contents are as stated. The Coal-measure species will 
be noticed in their place. 

Thus the united British Lamellibranchiate (or Pelecypod) fauna 
numerically reaches 54 genera and 424 species, 179 being Mono- 
myarian Asiphonida (Integropallialia) and 245 Siphonida (Sinu- 
pallialia). Hitherto I have not mentioned the Coal-measure Dimy- 


* 18 families, including the Arcidz, Trigonid, and Unionida. 


218 PROCEEDINGS OF THE GEOLOGICAL SOOIETY. 


aria, or those occurring in the Millstone Grit and Lower, Middle, 
and Upper Coal-measures. With the exception of the Lower Coal- 
measures, which have yielded 12 genera and 42 species, the three 
remaining horizons are comparatively poor, as would be anticipated 
from the estuarine condition or nature of the deposits. Anthracosia, 
Anthracoptera, Aximus, Ctenodonta, Edmondia, Modiola, Myacites, 
Myalina, and Schizodus are the genera that characterize the Coal- 
measures, or beds above the Yoredale series. 

6 genera and 9 species are all that are known from the Millstone 
Grit; the genera are Anthracosia 2 species, Aanus 1, Ctenodonta 3, 
Edmondia 1, Lunulacardium 1, Myacites 1. 

The genera in the Lower Coal-measures, 14 with 44 species, are 
still more estuarine in habit, many allied to the Unionide and 
Myadz—Anthracomya 5 species, Anthracosia 6, Anthracoptera 2, 
Axinus 2, Conocardium 1, Ctenodonta 6, Edmondia 3, Leptodomus 1, 
Modiola 4, Myacites 4, Myalina 6, Pleurophorus 1, Pullastra 1, 
Schizodus 2. The Middle Coal-measures have, as yet, only yielded 
4 genera and 16 species—Anthracomya 6 species, Anthracosia 6, 
Anthracoptera 2, and Myalhna 2 species. The Upper Coal-measures 
contain the same 4 genera with 11 species. The above, with the 
table of distribution, clearly shows the changes from the deeper-sea 
fauna of the Limestone series to the shallow and estuarine accumu- 
lations of the Coal-measures, the gradual dying-out of those genera, 
essentially dwellers in clear and deep water and with associated 
sedimentary matter, as well as the almost total extinction of the 
molluscan fauna with the elevation of the sea-bed, a few genera 
only living on into the Permian sea. 

The accompanying Table gives the numerical value of the species 
of 18 of the chief Dimyarian genera out of 43 for England, Scot- 
land, and Ireland; also Belgium, for comparison, which is the only 
country in Europe where the Carboniferous rocks (Lower series) are 
well developed. The researches of Prof. De Koninck, of Liege, have 
afforded me data for this column. I have before stated the small 
number of known species in the European area beyond Belgium ; to 
show, however, the relationship between the American fauna and 
our own, I have appended a column also, which shows excess in the 
number of species in most of the genera. Such holds good with the 
older Paleozoic genera also; but, through the flora and Mollusca of 
the Coal-measures, America and Scotland are intimately associated, 
and, in a similar but less degree, Ireland, 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 219 


Table of 18 of the chief Carboniferous Genera, showing their specific 
or numerical value through the five areas or countries named. 


| | [mel 
oe dies : 5 Oo | 
ElElglelgig. 
Genera. eh ot So lcen ine lee 
q s 9 | 'o ee 
Bla ae |Alsai6 
Allorisma (Myacites) ...... 6} 3] 4 28 | 41 
ATC APACOINYA.......00ss-000- Salas 2 | 13 |Chiefly Coal-measures 
(6 species). 
RIUBEAEORIG) <.650.5 00.0 20200- 10; 9 23 | 8 | 50 | Ditto (6 species). 
Lacs, 2 ee AT SOn StOn ke ht 44 
LAShit =e Rel ae 100 Sta Et 
Cardiomorpha ............... 9} 6| 5]} 86} 18 | 69 
Conocardium .........:..... Gullean Ol (Ss) 39 
EGtemodouta, <.:.......<...-.-. 11 | 15 | 15 | 9 | 34 | 84 | 6 Coal-measures. 
Cypricredia. ....-............. eo OM 1Gy TL os 
JSC ES Nit 12 eee 10 | 10 | 12} 2 | 25 | 59 | 8 Coal-measures. 
LG) A 10; 8} 12]... | 5 | 35 |4 Coal-measures. 
BURY EMBs soa. -cas-..eesees 3; 2] 3] 380] 6} 44 
WAM eee. 32-2.--.02.-.00- 6; 9; 1| 3] 18 | 37 |6 Coal-measures. 
Sanguinolites................ LOv) 20) | 1645327 |-68 
SCLNIZ00 si 3| 8] 7]... | 13 | 31 | 2 Coal-measures. 
BCAGWHEMED -20.2.2000s00.000- ees | Bani ak 6 | 15 
i Timrce. ——————— Were era omni orl oul t4 
SL. TUT 3A Ease eens LST PRG at ha bern 
| 106 /102 115 [171 [222 716 


Sorznoconce1a.—6 species of Dentaliwm are known ; but none pass 
above the Carboniferous Limestone. D. priscwm is the only species 
that occurs in 3 horizons—the Calciferous Sandstones, Lower Lime- 
stones, and Carboniferous Limestone. JD. scoticum is only found in 
the Calciferous Sandstones; the remaining 4 belong to the Carboni- 
ferous Limestone, and do not range higher. 

GaAsTEROPODA.—223 species and 29 genera, all belonging to the 
division Holostomata of the order Prosobranchiata, constitute the 
univalve fauna of the Carboniferous rocks. All the genera (29) and 
202 species occur in and range through the Carboniferous Limestone ; 
or nearly every species occurs in this horizon. 16 of the 29 genera 
are distributed through the Carboniferous rocks of Europe and 
America, and are therefore of zoological as well as stratigraphical 
value. I give the following Table of these genera to accompany 
that showing the distribution of the Lamellibranchiata (Dimyaria) ; 
by comparison with the molluscan fauna of Europe and America 
we may hope to obtain some clue relative to the migration and dis- 
persion of the Mollusca from some original area. These 16 chief 
genera are represented in England by 176 species, in Scotland by 
90, in Ireland by 111, in Belgium (chiefly through De Koninck) by 
176, and in the American Carboniferous rocks by nearly 200 species. 
This generic relationship with America is important as determining 
similarity of bathymetrical conditions, temperature, and food, and con- 
nexion through coast-line or land now lost. 9 genera and 21 species 


nt 


220 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


occur in the Calciferous Sandstone—Huomphalus 4 species, Lacuna 
1, Inttorina 2, Loxonema 2, Macrocheilus 4, Murchisonia 3, Natica 1, 
Naticopsis 1, Pleurotomaria 3. 

The Lower Limestone Shales contain 50 species, which, with one 
exception (Turbo appropinquans), are also those of the Carboniferous 
Limestone above; or 49 of the 50 occur also in the massive lime- 
stones. 

The Yoredale beds (the Upper Limestone Shales of some authors) are 
poor in Gasteropoda; only 7 genera and 10 species appear to be known. 
Clearly this must be the result of imperfect collecting rather than 
of almost total absence of species. We know that both the Lower and 
Upper Limestone Shales are impure argillaceous limestones, and not 
favourable to the presence of the Mollusca at the time of deposition 
or to the preservation of organic structure afterwards: but we see here 
the 7 genera only represented by 10 species; they are :—Huomphalus 
catillus, Sow.; Loxonema constricta, Sow.; Macrocheilus curvilinea, 
Phill.; Jf. imbricatus, Sow.; M. rectilinea, Phill.; Murchisonia 
fusiformis, Phill.; Naticopsis plicistria, Phill.; Plewrotomaria lim- 
bata, Phill.; P. twmida, Phill.; and Yurritella tenwistria, Phill. 
The Millstone Grit, as we should expect, contains but a small 
Gasteropod fauna; such arenaceous deposits suit not the habits of 
this group of Mollusca; only 3 genera and 3 species are known— 
Natica variata, Phill.; Pleurotomaria limbata, Phill.; and Murchisonia 
fusiformis, Phill. The Pelagic Cephalopoda in the same beds, as we © 
shall see, number no less than 33 species; but the sandy and muddy 
beds of the shore was not their habitat; they are not such good wit- 
nesses in the beds in which they are found as the ordinary Lamelli- 
branchiata and Gasteropoda. The Lower Coal-measures alone yield 
Gasteropoda; none are known either in the Middle or Upper. 7 genera 
and 19 species are recorded from this division :—Huomphalus Glover, 
Brown; Juttorina obscura, Sow.; L. solida, De Kon.; Loxonema 
galvam, Baily; L. minutissema, Baily; L. Owen, Brown; L. reticu- 
lata, Brown; Macrocheilus, 4 species ; Natica vetusta, Sow.; Nats- 
copsis plicistria, Phill.; Plewrotomaria limbata, Phill.; P. usocona, 
Sow.; Zurritella, 4 species. 

Middle Coal-measures none. 

Upper Coal-measures none. 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 221 


Geographical Distribution of the Gasteropoda through 
16 characteristic Genera. 


| | | 
ne] ne) ra osteo 
| Ae es ieeeceices | 
SPE. Nishi 1 oes oc Ree ae | = 
<3) 2) Ss ea bales 6 
“ESET TOL a eae ins ee 6 5 8 13 22, 54 
CClifiae, | go ree 3 3 4 15 2, 27 
BU OMIDMALUES Jt enscnvesseeeses 20 Wi 20 20 23 | 100 
LORS ae a a 2 1 os 2 yes 5 
WMERCROCMOIUG! { 1. ..0--cc0--00 lesvses 17 8 12 5 19 61 
EOWA A oo ea. = cvnon noses sens 12 Na os 5 2 19 
RECIEVE)... soc sorcccceccsseees 15 O 6 19 20 67 
MUSE TURN coco es ceacccwswecnose 13 9 9 14 16 61 
PE AVICOSIS mod. ais o's ox a'view oneiecnt 2 1 3 a mE 6 
[ESE 12) ° 5 Seen eee 7 ae 5 6 ay 18 
IPMAMICTORMUS, -coceccsecccecccsecsee 4 ee 3 “ss 2 9 
IIABYRCHISWID, 2... 05-2 cecc econo sae 9) 2 4 1 + 16 
IEIOMEGLOMIATIG «.-5ccscccneeecnseces 46 24 16 59 65 | 210 
MIME TOUS 2s oo coc la occecccccevscaecs 6 1 4 aan 2 13 
TNS 19)” GA ae i 6 1 4 fy af, 16 
| SOS TEL GSIEE: Gea Seo ere 12 i] 13 12 20 68 
176 90 | 111 WG. tb 197-2 F750 


Preropopa.—The genus Conularia first appears in the Carboni- 
ferous rocks in the Lower Limestone Shale, and ranges through all 
but the Upper Coal-measures. This genus dies out in the Coal- 
measures. Many of the nodules in the “ Penny-stone” of Coal- 
brookdale contain fine examples of C. quadrisulcata, Sow., the only 
determined species known. An undetermined species occurs in the 
Calciferous Sandstones of Woodhall, Scotland. America yields 17 
species of Conularia to our 2; our C. quadrisulcata is not known 
out of Britain. 

Herrrovopa.—Bellerophon and Porcellia (the former with 27 
species and the latter 4) exhaust the species in the Carboni- 
ferous rocks. At no period in Paleozoic times did so many species 
of Bellerophon exist in one horizon. 24 of the 27 species are 
found in the Carboniferous Limestone; and 6 of these 24 species 
range higher; they are :—JBellerophon apertus, Sow.; B. decussatus, 
Flem.; B. Dumonti, D’Orb.; B. hiulcus, Mart.; B. Oldham, Portl. ; 
and B. Uru, Flem. 4 species (B. costatus, Sow., B. decussatus, 
Flem., and var. undatus, Kther., and B. Urii, Flem.) are Calciferous 
or Tuedian species. B. decussatus, B. apertus, B. hiulcus, and B. Urii 
range up to the Middle Coal-measures. 5 species occur in the 
Lower Limestone Shale; but none are peculiar to those beds. 17 
Species are essentially Carboniferous Limestone. The Yoredale 
beds have yielded 4 species—B. apertus, B. decussatus, B. hiulcus, 
and B. Urn, all species having a long range. The Millstone 


222 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Grit contains the same forms. The Lower Coal-measures yield the 
same, with the addition of B. navicula, B. Oldhamu, and B. 
Dumontic; none occur in the Upper Coal-measures—this order, 
like all others in the Carboniferous series, dying out at its close, not 
to appear again. 

The genera Bucania of Hall and Huphemus of M‘Coy are synony- 
mous with Bellerophon. Ireland has yielded 21 species of Bellero- 
phon, Scotland 15, England 17, Belgium 25, and America 39 species ; 
only 19 species range through Europe, 8 of them being British; and 
14 of these 19 are Russian. 

CrpHatopopa.—No fewer than 169 species are distributed through 
the British Carboniferous system. 4 genera and 6 species occur in 
the Calciferous or Tuedian beds, 5 genera and 22 species in the Lower 
Limestone Shale, 8 genera and 140 species in the Carboniferous Lime- 
stone ; 5 genera and 33 species are Yoredale ; 3 genera and 30 species 
are Millstone Grit; 3 genera and 24 species occur in the Lower Coal- 
measures, 3 genera and 6 species in the Middle Coal-measures, and 
1 genus and 2 species in the Upper Coal-measures. These 169 species 
represent 8 genera—Actinoceras 1 species, Cyrtoceras 3, Discites 17 
(subgenus of Nautilus), Goniatites 59, Nautilus 36, Orthoceras 48, 
Poterroceras 3, and Tirigonoceras 2 species. 

The accompanying Table of the 8 British genera shows the number 
of species in each country, including, as in the two prior Tables, 
Belgium and America for comparison, in which it will be seen that 
the 169 species, through the 500 occurrences, are thus distributed :— 
the number of species in the 8 genera in England is 126, in Scot- 
land 63, in Ireland 107, and in Belgium 90. Comparison carried 
beyond Europe to America, as before, shows generic affinity through 
114 species, hardly a single form being British. All Europe, only 
yields 95 species through 11 areas; research and the accident of our 
possessing these rocks highly developed is the reason why our mol- 
luscan fauna is so extensive. 


8 

é : : q 

rd oo ro 3 

BP lik Os lly ee ca ees a 

a) 3) 5 a0 ee 

WA CEIMOCELAS Ye face sean oe ekeseeenee 1 1 1 1 2 6 
Oyrtoceras.:, hiss sc. seeasenlesdwcess 2 3 aie 8 5 18 
DIS Cites, Sees. f ee Ne asset eek 10 3 LS idee 1 32 
Goniatites 2047 tcacnaneccteeties 59 15 32 25 32 | 163 
Nautilus: tin tscccscsecec neuen ea ea: 18 14 21 22 54. 129 
Orthoceras /eeerk eee cee Reena eee 33 24 36 29 20 142 
IPoteriocerasin. econ ee eee 2 3 2 au Fad a 
Tri gOnOGeLrasnnc:t pa scmeedennesee 1 a 1 1 on 3 


126 63 | 107 90 | 114 | 500 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 223 


VERTEBRATA. 


Piscrs.—120 genera and 290 species of fish and 26 genera and 
33 species of Amphibia compose numerically the Vertebrate fauna 
of the Carboniferous rocks. The comparatively rich assemblage of 
ichthyic remains in the Old Red Sandstone bears no proportion to the 
great increase and development of this class in the Carboniferous 
system. Only two orders are represented in the Carboniferous 
rocks, viz. the Ganoidei and Elasmobranchi (Placoidez, Ag.), the 
Ganoidei through the suborder Crossopterygide by 4 of the 6 recog- 
nized families, the Saurodipterini, Glyptodipterini, Ctenodipterini ? 
(Dipnoi)*, and Celacanthini. The Elasmobranchi mainly corre- 
spond to the Chondropterygide or the Cartilaginous Fishes of Cuvier 
(the Holocephali and Plagiostomi of Owen and the Selachia of 
Miller). 

The distribution and appearance in time of the two orders appears 
to have been much the same; the Plagiostomi first appeared in the 
Ludlow rocks, the Holocephali through the Chimeroids in the 
Devonian rocks of N. America (genus fhinodus of Newberry). The 
section Cestraphori of the order Plagiostomi 1s abundantly represented 
in the Carboniferous rocks from base to summit, spines and teeth of 
the several genera occurring everywhere, although very locally, the 
Cienacantht, Gyracanth, Homacantht, Oracanthi, Onchi, and Lepta- 
canthi being the most important, and occurring as spines (Ichthyo- 
dorulites). 

The genera or groups illustrated by palatal and other teeth are 
Cochliodus, Deltodus, Psammodus, Petalodus, Otodus, Ctenoptychius, 
Cladodus, Glossodus, Diplodus, Helodus, &e. Many of these generic 
groups will be greatly reduced under stricter and more complete 
research. My duty here is not to criticise, but rather to bring 
together for a special purpose the labours of others as at present 
received. 

The lowest recognized rocks of the Carboniferous system, the 
“‘Calciferous Sandstone” of the Scotch geologists, or Tuedian of 
Northumberland, confined to the north of England and south-east- 
ern Scotland, contains 13 genera and 14 species—Cladodus 1 species, 
Cienacanthus 1, Ctenodus 1, Ctenoptychius 1, Diplodus 1, Hurynotus 1, 
Megalichthys 1, Nematoptychus 1, Pecilodus 1, Pygopterus 1, Rhadi- 
nichthys 1, Rhizodus 2,and Wardichthys 1. Thus, with one exception, 
each genus is represented only by 1 species. Surely much has to be 
done both with the Ganoidei (8) and Placoidei (5) when better 
materials come to hand ; for many of the genera must be established 
upon slender grounds. 

THe Lower Limestonzs anD SHatEs have yielded 29 genera and 
50 species ; and if this lower member be associated with the thick or 


* T am aware that Dr. Traquair would remove the Glyptodipterini from the 
Crossopterygious Ganoids and place them sectionally in the order Dipnoi (vide 
Trans. Roy. Soc. Edinb. vol. xxvii. for the sections and families proposed by 
Dr. Traquair). 


224 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


true Carboniferous Limestone, the two together would represent 80 
genera and 170 species, 17 genera and 22 species being peculiar to 
the Lower Limestone Shales. No one can doubt that these dark, 
impure, argillaceous limestones were the commencement of the purer 
calcareous series above; and paleontologically they cannot be sepa- 
rated. Closer research will probably show that the 17 genera now 
known to occur only in the Lower Limestones exist in the succeed- 
ing series, 

Carponirerous Limestonz.—This finely developed group of rocks 
in the British Islands, which has received so much attention both 
under geological and paleontological aspects, is possibly better 
understood than any other division of the Carboniferous system. 
The limestone beds known as the Mountain Limestone have yielded 
no less than 63 genera and 147 species of fishes; but the Yoredale 
series which succeed them yield none ; and only 3 genera and 3 species 
are known in the Millstone Grit (Megalichthys Hibberti, Rhizodus 
Hrbberti, and Acanthodes Bronni). This sudden cessation or non- 
occurrence can scarcely be accounted for on lithological grounds, or 
original sedimentary accumulation, such as would be applicable to 
the habits of certain genera of the Mollusca. Bathymetrical con- 
ditions may have influenced this paucity in the whole fauna of the 
Yoredale and Millstone Grit, which is so apparent on inspection of 
the Table of Distribution. 38 genera of Placoidei are confined to 
the Carboniferous Limestone; and 12 genera of the same order are 
from both Lower Limestone Shale and Carboniferous Limestone: on 
the other hand, it is remarkable that only 6 genera of Ganoidei are 
strictly or essentially confined to that horizon ; they are Acrolepis, 
Asterolepis, Coccosteus ?, Cycloptychius, Phyllolepis, and Platycanthus. 
(I give them as near the truth as possible, as the species of Ganoidei in 
the Carboniferous system are in as much confusion as the Placoidei*.) 
As before stated, the Yoredale rocks(Upper Limestone Shale of some 
areas) have not yielded remains of fishes. We are in doubt as to the 
real position of the Upper Limestones of Durham and Northum- 
berland, or what, if any, should be assigned to the Yoredale group. 
Nevertheless [ am not aware of any Vertebrata occurring in the 
Yoredale beds. The Millstone Grit, as before stated, has only (so 
far as I know) yielded 3 genera and 3 species, and these in the 
form of scales. | 

Lowrr CoaL-MEAsuRES.— Whatever conditions caused so complete 
a break in the distribution of the fishes in our area zoologically, 
stratigraphically, and geographically during the deposition of the 
rocks mentioned, the return of the same genera and many of the 
same species in the Lower Coal-measures is equally important. We 
have evidence of the remains of 52 genera and 112 species, the 
whole composed nearly equally of the orders Placoidei and Ga- 
noidei. The numbers may be thus expressed :—In the Lower Coal- 


* The Ganoid Fishes are under revision by Dr. Traquair, whose accurate 
knowledge of the order will be brought to bear upon their structure and classi- 
fication. 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 225 


measures 32 genera and 50 species of Ganoidei occur, in the Middle 
Coal-measures 5 genera and 6 species, in the Upper Coal-measures 
6 genera and 8 species. The Placoidei are represented in the Lower 
by 22 genera and 40 species, in the Middle by 10 genera and 14 
species, and in the Upper by 6 genera and 8 species. Regarding 
the Coal-measure fishes as a whole, however, they stand in the Table 
as Lower Coal-measures 52 genera and 112 species, Middle 14 
genera and 20 species, and the Upper 12 genera and 14 species. 
The numerical distribution for Britain and Belgium is— 


[ST EG oe 136 species. 
seaulands | oss Socey 5... P52) es 

Mrclanieey ps 2 ee ad 21) miner 

ibelstunyeS a eee ou S. ist) Wagers 


‘The Carboniferous system is finely developed in Belgium. 37 of 
the same genera occur in the American rocks, 10 being Ganoidei 
and 27 Placoidei, the two orders there yielding 240 species. 

Amputpra.—Of the 4 orders in the class Amphibia, the Labyrin- 
thodontia alone occur in the Carboniferous rocks. 26 genera and 
33 species are known in Britain. Without exception they all 
occur in one or the other of the divisions of the Coal-measures. 

Prof. De Koninck, in his great work (‘ Ann. du Mus. Roy. d’Hist. > 
Nat. de Belg.’ tome ii., Faune du Calc. Carb. de la Belg. pt. i.), has 
described 29 genera and 44 species, all from the Carboniferous 
Limestone. The fauna of the Coal-measures in Belgium is small 
compared with that of Britain. 

Tue Lowsr-CosL-MEASURE genera are Ainphicelosaurus, Amphi- 
saurus, Anthracerpeton, Brachyscelis, Batrachiderpeton, Discospon- 
dylus, Dolichosoma, Erpetocephalus, Ichthyerpeton, Keraterpeton, 
Lepierpeton, Loxomma, Ophiderpeton, Labyrinthodontosaurus, Lepto- 
gnathosaurus, Macrosaurus, Megalocephalus, Orthosaurus, Parabatra- 
chus, Pholidogaster, Streptodontosaurus, and Urocordylus. 

Tar Mippre-Coat-MpasvRE genera are only 3—Anthracosaurus, 
Megalerpeton, and . holiderpeton. 

Loxomma and Pieroplax occur in both Lower and Upper Coal- 
measures. 

We are chiefly indebted to Professors Huxley and Miall for 
elaborate researches into the structure and affinities of the extinct 
order Labyrinthodontia, a group eminently characteristic of Carbo- 
niferous time; for, with the exception of RAinosaurus from the Lias 
and the Jurassic Brachyops, no remains of this order have hitherto 
been discovered in rocks younger than the Trias. 


226 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


TaBLE XVI.— Carboniferous. 


——— 


*UBIMLIOg | 
07} 03 ssed an ne oe 

yey} soroodg | 
*soansvoul-[v0D | .@ © 
soddq ae aN =e OO AR v0 NS av on 
"SQITISVOTU-[VOD) | ~@ a o® 


AA TO art Oo Av SF ae sa MO 


eTPPIAL a mc =e 


*SOUNSVOT-[VOD) | ga Glo an 
JOMOTT 2% rie SE OA COD Ore may coke Se re nin am mg on an ae 

e e . e e pr) 

“ITH OUOISTTNAL | aa SS og yeaa HUES Re Pe 22 me an nt a, ae 3 | oe 
a 3 Q . ° a 

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“QUO SOUT Oa 6 0o ot Ho ot @ : ° > ov 8 At of os te ” ae 
snoIE;IUOGIeO nn a ome SN ot ae rie ae 2 S 7 Nr nr ne ae ae 8 ret) (GU OS asi ae 
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-OUI'T -ToMOorT NN aN Ca Cs cb : 2 rN nD “NO a Dale) ; mM e¢ | le 
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ANNIVERSARY ADDRESS OF THE PRESIDENT. 227 


Tre EXTENSION oF THE OLDER OR Patmozorc Rocks BELOW THE 
NEWER oR SECONDARY FORMATIONS. 


The position and extension of the Paleozoic rocks beneath the 
newer formations of the British Islands is a problem of deep interest, 
and is now occupying much attention in consequence of the facts 
brought to light by the numerous trials lately made either for the 
supply of water or in search of minerals. It may be said that ever 
since the remarkable trials for water at Harwich and Kentish Town, 
and also the still more remarkable generalizations (almost predic- 
tions) of R. Godwin-Austen, Esq., in 1856*, and Prof. Prestwich in 
18727, with reference to the possible extension of the Coal-measures 
beneath the south-eastern part of England, the minds of pure geolo- 
gists have been excited by speculative views, and desires to arrive 
at some knowledge of the extension or distribution of old land or 
Paleozoic surfaces, so as to restore to the eastward in England the 
physical geography of those groups of rocks which now constitute so 
grand a feature along the western side of England, Wales, and Scot- 
land, but which are lost or covered up beneath the unconformable 
newer or Secondary rocks. West of long. 1° 30’ the greater part of 
the exposed rocks are Palxozoic, ranging from the Cambrian to the 
Coal-measures, their general strike being about N.E. and 8.W. 
East of this meridian are Secondary and Tertiary rocks of great 
thickness, which doubtless cover the easterly extension of the Pale- 
ozoic series towards the European continent. The geographical 
changes of land and sea must have been numerous from the time 
of the consolidation of the Cambrian sea-bed to the close of the 
Carboniferous epoch, the rocks of the latter period being deposited 
in depressions and valleys of the older, with succession or position 
due to the removal of the subjacent rocks. With the old and wide 
extension of these earliest-formed masses we are becoming daily more 
familiar. Looking at the physical structure of the south-western 
and north-western parts of the British Islands, and the great mass of 
the older Paleozoic rocks of North and South Wales, it is evident 
that from the Cheviots to Cornwall the oldest rocks in Kurope are 
exposed, their eastern extension being hidden. The Northumber- 
land and Yorkshire coal-fields down to the latitude of Notting- 
ham are covered and deeply buried by the Triassic, Jurassic, and 
Cretaceous rocks. South of Nottingham these old land areas are 
again exposed; the Charnwood rocks of unknown age, the associated 
coal-field of Ashby-de-la-Zouch, with the Warwickshire and South- 
Staffordshire coal-fields stand out like islands in the midst of the great 
Triassic plain of mid-England; they are the last isolated exposures 
or remnants of Paleozoic land seen south of the great Penine axis. 
A line drawn from the Malvern range, due south to the Mendips, 
and thence to Torquay, will define absolutely the exposed line of 
demarcation between the Paleozoic and Mesozoic rocks. The mass 
of North and South Wales stands out in bold relief westward 


* Quart. Journ. Geol. Soc. vol. xii. pp. 838-4 
+t Popular Science Review, vol. xi. p. 241 (1872). 


228 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


of the Severn valley. The Old Red mountains and older Silurian 
rocks which border the northern edge of the South- Welsh coal-field, 
as well as the Devonian promontory of Cornwall and the mass of 
North Devon, isolated as they appear to be from the unconformity of 
the Secondary rocks, are only apparently so through the great over- 
lap. Could we uncover and expose the old Paleozoic floors or land- 
surfaces with all their irregularities, doubtless we should find 
that the eastern face of the Paleozoic plain would stretch away 
under the north-eastern and south-eastern counties and the 
German Ocean, the newer rocks filling up the irregularities in the 
old land-surface—this denuded plain being either produced by the 
eroding agency of the Secondary seas during the slow depression 
of the area they then occupied, or previously sculptured and fashioned 
into hills and valleys prior to the deposition of the Secondary or 
Mesozoic rocks. ‘The Irish Sea, the English Channel, and the 
German Ocean are only hollows in the land occupied by the several 
seas around the British coasts, any important change in which 
would alter our relative position to the continent either to the north, 
the east, or the south, and in case of upward movements, would 
reveal those accumulations which have gone on since the close of the 
Glacial epoch. Could the valleys of the North Sea or St. George’s 
Channel, the English Channel, and the German Ocean become again 
dry land, we should again be restored to and form part of the great 
European plain or plateau, and those stratigraphical masses that are 
now abruptly cut off at the coast all round the British Islands would 
be traced in broken continuity over their once continuous or origi- 
nally connected area. Further, could we strip off all the Secondary 
and Tertiary rocks, and reveal or expose the extension of the older 
or Paleozoic series towards Germany on the east, and France on 
the south, then the vexed question of the old physical geology and 
geography (paleography) of Britain and the relation and correlation 
of our area with that of Europe would be revealed; the once 
continuous terrestrial surface joining us to Europe, and probably 
America, on which grew and flourished the flora which furnished 
the materials of our coal, could be determined; the probable 
relation of the underlying or partly contemporaneous Devonian to 
both the Silurian and Carboniferous; the reason for the isolation of 
the Old Red Sandstone in different geographical areas, marine in 
one area, freshwater in another—the one with a well-defined base 
and top, the other having as yet no discovered base, but having a 
well-defined passage into the Carboniferous; such and a hundred 
other problems would be solved could this old floor be ours to 
examine. A rise of a thousand feet would reveal much of all the hidden 
older land east of long. 2° W., or all eastward of that which extends 
from the southern termination of the Penine chain and Charnwood 
Forest ; for we now know that as far south as Northampton, and at 
less than 1000 feet*in depth, the Carboniferous Limestone occurs. 
Still further south, and but little deeper (1184 feet), the Coal- 
measures have been proved at Burford, in each case yielding the 
characteristic fossils, the limestone at Northampton being crowded 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 229 


with 2 species of corals, Lithostrotion irregulare and Lonsdaleia flo- 
riformis ; the Coal-measures at Burford with Oyclopteris orbicularis, 
Neuropteris, and Pecopteris. By degrees we are arriving at sufficient 
data to enable us to judge somewhat of the physiography of these 
older accumulations or formations and arriving at their distribution. 
The sources of the oldest sedimentary strata will probably ever 
remain a mystery; the materials that supplied these oldest British 
strata have wholly disappeared ; the extent, distribution, and dimen- 
sions of the Paleozoic series all afford indications and proofs of the 
vast regions in the north and west which have been thus denuded or 
washed away. 

This brings me to the consideration of the proof of the extension 
of the Upper Silurian and Devonian rocks eastward of the Malvern 
chain, the Staffordshire beds (Dudley), and the Bristol Coal-basin, 
obtained at no less than five places, if not six; the seventh was 
unfortunately never completed. 

The oldest rocks yet touched are the Wenlock; they were deter- 
mined at Ware, in Hertfordshire, at 795 feet, immediately beneath 
the Gault, during the process of boring for an extra supply of water 
for the New River Company. These beds dip at an angle of 40° to 
the S.E.; consequently their strike is from the N.E. to the S.W.; and 
probably the Devonians rest upon them in succession, as they are 
known to occur at Turnford, 7 or 8 miles to the south of Ware, and 
continuously on to London. No rocks, however, except 1 foot of 
Lower Greensand (‘“ Carr stone”), came in between the Silurian and 
the Gault—the Devonian, Carboniferous, and all the Lower Secondary 
rocks being unrepresented. This feature in the paleogeography 
of the eastern region of England, long ago anticipated in some form 
by Austen, Prestwich, and Hull, has now been verified, and the age 
of the rocks determined. The line occupied by the Wenlock rocks 
may be higher or nearer to the north than was hypothetically believed 
by the authors above mentioned, but not more so than the 
course or strike of the Silurians and Devonians would probably 
take, having regard to the position of the Malvern, Woolhope, May- 
Hill, and Tortworth Silurians to the west, although the Ware beds 
can hardly be referred to the type of Silurian rocks that exists at 
the places above named. It will therefore be asked whether this 
Wenlock at Ware is of British or continental type; in other words, 
ean it be correlated with our Welsh or English Upper Silurian, or is 
it of the Ardennes type? Do they constitute a portion of the Staf- 
fordshire (Dudley) or Shropshire (Wenlock) Silurians, spreading away 
eastwards towards Belgium? or are they a prolongation of the Silu- 
rians of Belgium to the west, or a western extension from the con- 
tinent? In other words, does this Wenlock at Ware belong to the 
edge of another basin or coast-line, an extension from Western 
Europe or Scandinavia, or an easterly expansion of the Upper 
Silurian of the Silurian area? The facies of the fossils and the 
characters of the rock in all essentials are decidedly British; yet 
there is much resemblance to the Scandinavian fauna, a prolongation 
of the rocks containing which is by no means impossible or impro- 


VOL. XXXVII. if 


230 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


bable. Onthe whole, however, I am inclined to regard the Wenlocks 
under Ware as part of the continuous and old denuded Upper Silurian 
surface of those rocks now conspicuously exposed in Herefordshire, 
Shropshire, and Worcestershire. In direct E. and W. line, and latitu- 
dinally, the Wenlock rocks of Malvern, Woolhope, and May Hill may 
(under the view of continuity) claim connexion; but I regard the 
Ware fossils as having more affinity with the Wenlock-Edge group, 
or that series which underlies the western side of the great mass of 
the Old Red Sandstone. 

The fossils also equally resemble those of the Wren’s Nest at 
Dudley, where the dull earthy limestones as well as the crys- 
talline ones (as at Ware) occur. The 33 species of fossils noticed 
in the cores at Ware are species for species identical with those of 
the Wenlock Edge or the Wren’s Nest (Dudley). Could we remove 
the overlying Mesozoic series between Ware and Burford, and again 
expose the Coal-measures known to occur there at the depth of 
1180 feet, then should we better understand the thinning-out of 
the Triassic and Jurassic series eastwards towards this Silurian 
ridge or plateau. We can hardly .now doubt the extension of 
the Burford Coal-measures, in all ‘probability terminating against 
the Silurian or Devonian series in the: Ware region, at Turnford, 
and near London. 

To what distance rocks of these or older date may occur north of 
Ware further research alone can:decide. We have, therefore, under 
the so-called London basin an axis of Palzeozoic rocks, two divisions 
of which are known, the Upper Silurian and Devonian. The Lud- 
low rocks may be expected to occur under or a little south of 
Hertford ; for, as we shall see, the Devonians set in between Ware 
and Turnford, and probably occupy the entire area between that 
place and London, where it is now well known they occur under 
Tottenham Court Road. The boring at Ware was carried down 
797 feet and into the Wenlock beds to the distance of nearly 
50 feet, or, to give particulars :—Gravel 14 feet, ‘Chalk 416 feet, 

halk-marl 128 feet, Upper Greensand 77 feet, Gault 160 feet, and 
Lower Greensand (Carr stone) 1 foot; the boring was continued 
for 50 feet in the Wenlock Limestone, and without the intervention 
or occurrence of the Devonian. We are thus justified in stating 
that the old Paleozoic land-surface composed of Devonian and 
Silurian rocks occupies much of Middlesex and Hertfordshire at the 
mean depth of 970 feet; they probably extend westward to the 
exposed Silurian, Devonian, and Carboniferous areas of Cornwall, 
Devon, and Wales. Harwich, which lies 80 miles to the eastward 
and 10 miles further north, and probably on the same strike, has 
revealed Paleozoic rocks at a depth of over 1000 feet ; but the age of 
these Harwich dark clays or shales was never clearly made out. 
This question of the depth and geographical extension of these oldest 
rocks is seldom practically tested ; for no mineral wealth of sufficient 
value occurs below the Coal-measures (Upper Palseozoic) to induce 
trial or experiment; and but for the purpose of obtaining water 
pure and in quantity, it is questionable if this problem would have 


ANNIVERSARY ADDRESS OF THE PRESIDENT, 231 


been solved (at present at least); had the Lower Greensand oc- 
curred in its normal condition as a water-bearing stratum, or even 
had the Upper Greensand (above the Gault) yielded water in quan- 
tity, neither at Messrs. Meux’s, at Turnford, nor at Ware should we 
have touched the undoubted and unequivocal Devonian and Silurian 
strata. Few give thought to or are aware of the difference that 
exists in the thickness of rocks of the same age in different yet not 
very remote localities. In Britain the Cambrian and Lower Silurian 
deposits are from 20,000 to 30,000 feet in thickness, whilst in 
Sweden and Russia their representatives or equivalents in time 
rarely if ever exceed 1000 feet. 

This difference is and was probably due to the form and nature of 
the Pre-Cambrian land on which the newer Cambrians and Silurians 
were deposited; for there cannot be any doubt that such Pre- 
Cambrian rocks did and do exist, and were and are widely extended, 
although concealed over the present known European area, and that 
their existence was connected with a probable great geographical ex- 
tension westwards of the British Islands. The plateau governed by 
the 100-fathom level that surrounds the British Islands is part of 
this extension, on which all our physical changes have taken place. 
Probably the crystalline rocks of Scandinavia, parts of North Wales, 
North-west Ireland, St. David’s, and the Hebrides are exposed areas 
of this Pre-Cambrian stage of the highest antiquity, and were covered 
on their submerged and denuded masses by the Longmynd, Harlech, 

ad St.-David’s rocks, which in their turn were succeeded in 
some areas by the Lingula-flags, the Tremadoc, and Arenig, life- 
groups of antiquity so high that we have no formula to express their 
age, or when life first appeared in the seas of the British Islands. 
As yet we have no evidence relative to these formations occur- 
ring eastward of the Penine chain, of which Charnwood, Ashby- 
de-la-Zouch, and the Warwickshire coal-field are the most southerly 
exposures ; but we have lately unexpectedly determined the presence 
of the Carboniferous Limestone below Northampton at the depth of 
890 feet*. This has carried still further south the Penaine axis, 
and would lead us to expect that between Northampton and the 
exposed Coal-measures of Atherston and Nuneaton an extended coal- 
field may occur. What relation the Burford coal on the south may 
have is conjectural only; but I am disposed to regard the whole as 
one greatly extended coal-tract. We must now regard the Carbo- 
niferous Limestone of Northampton in longitude 1° 30’ W. as the 
most easterly known in England; and no Triassic rocks cover this 
limestone, a few feet only of undeterminable rocks occur between it 
and the Lower Liast. Whether the New Red hasthinned away entirely 
here, or it happens to be accidentally absent, is a question; but at 
Burford on the same strike and relative position there is a consider- 
able thickness of Triassic rocks, and to the N. and N.W. also every- 

* Jn the cores brought up I determined the presence of Lithostrotion junceum 


and Lonsdaleia floriformis in abundance. 
+ The cores brought up are 15 inches in diameter. This great undertaking, 
as well as those at Turnford and Chatham, were carried on by Messrs. Docwra 


and Gulland with their new machinery. 


232 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


where. The Upper Devonian beds were first determined by myself 

at Messrs. Meux’s ; they are of the true N. Devon and N. Cornwall 
Devonian type—dark chocolate-coloured semi-slates containing the 
characteristic shells Spirifera disjuncta and Rhynchonella cuboides 
&c. This discovery solved the problem of the existence of Paleeozoic 
rocks at an accessible depth under London, and of the absence of 
the Jurassic series. Immediately above the Devonian occurred the 
Lower Greensand, abnormal in all conditions save the never-failing 
test of fossil remains, and what few occurred left no doubt as to the 
age of this at first somewhat doubtful rock. Trigonia aleformis, 
Cardium Hillanum, Trochocyathus Harveyanus, and Cerithia, &e., 
added to the general facies, immediately settled the question of age ; 
neither of the other four borings exhibited any signs of the true 
Lower Greensand. At Ware and Turnford the Gault rested upon 
from 8 to 10 inches of theCarr stone, and this was all that represented 
the Neocomian rocks beyond the 65 feet of chalky oolitic subcal- 
careous abnormal Lower Greensand; all the borings show that the 
Neocomian (Lower Greensand) is interrupted on the north by the 
underground Paleozoic ridge. The geographical extent or surface- 
area occupied by the Devonian of North Devon between the Pilton 
beds on the south and the Linton beds on the north (15 miles), 
where we believe we have the full thickness of the whole series, is 
as near as possible that known to occur between London and Turn- 
ford, assuming that the strike of the North-Devon beds is directly 
towards the London area, and thence on towards Belgium and the 
Rhine (Eifel and Coblentz). The Devonian cores brought up at 
both localities (Tottenham Court Road and Turnford) yielded the 
same characteristic fossils; and the beds dipped at the same angle 
at both places, 30° $.H.—the plane of the old land-surface being 
1148 feet below London, and 980 feet below Turnford, showing a dif- 
ference in level of 168 feet, due either to dip or denudation. I should 
believe the difference was due rather to denudation along or over a 
given plane striking from 8.W. to N.K. Where the junction of the 
Upper Devonian and Wenlock rocks may take place between Turn- 
ford and Ware it is difficult to say ; but if we infer that the Ludlow 
and Lower Devonian beds occur above the Wenlock, dipping south 
between Ware and Turnford, then there is little room for them, and 
the Upper Devonian must cease at no great distance north of Turnford. 
Looking at the thickness of the Lower Devonian in North Devon, and 
regarding it as maintaining the same in its strike under Wiltshire, 
Buckinghamshire, Middlesex, and Hertfordshire, there is room in 
the 8 miles from 8S. to N. at the dip of 30°S. for the presence of the 
Lower Devonian and Ludlow. We need not assume that the Lud- 
low beds do cecur, any more than we should expect to find the great 
sandy and gritty group of the Foreland, whatever it may repre- 
sent. The old land-surface, therefore, of Devonian and Old Red 
Sandstone of the southern half of England must have extended from 
near Yarmouth (lat. N. 52° 50’), descending by a gentle southerly 
curve to about Leighton Buzzard, thence 1ising to the Wenlock pro- 
montory, and again sharply deflected S.W. to Milford Haven, and 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 233 


on under the Bristol Channel to Cornwall, all east of the Mendip 
Hills being now covered by the Secondary and Tertiary strata— 
thus, I think, showing that a widely spread floor of these Paleozoic 
rocks occurs under the extensive region of the southern half of 
England, and which could as easily be shown to occur in the north 
and in Ireland, but under different conditions. 

Now that the Ware Upper Silurians have been proved to dip to 
the south, we may anticipate the occurrence of the more ancient or 
lower series further north towards Cambridge, Huntingdon, and 
Peterborough. Whether in their extension northwards they may 
be of Scandinavian type we can only surmise, unless the Westmore- 
land and Cumberland Silurians pass under the unconformable and 
overlapping Carboniferous group of the Penine chain as a con- 
tinuous floor or old surface to the south. 

Since the determination of the existence of the Devonian rocks 
north of the Thames to Turnford, and of the Upper Silurian at Ware, 
below the Cretaceous rocks at the depth of 800 feet, our views rela- 
tive to the distribution of the Coal-measures have materially altered. 
It is clear that all the stratified rocks between the Silurian and Gault 
and the Devonian and Gault are wanting or not represented ; in other 
words, the whole of the Carboniferous, Triassic, and Jurassic rocks, 
and the Purbecks, &c. are missing north of the Thames to lat. 52° 10’. 
This shows the great unconformable overlap upon the Silurian and 
Devonian floor to the west. Had the sub-Wealden boring been 
completed, the problem as to extension would have been solved. 
Unfortunately for science this undertaking was never completed, 
and the problem remained unsolved; the ancient land was not 
discovered. We must rest content with the hypothesis of Mr. 
Godwin-Austen as to the area where we should expect to find the 
Coal-measures, probably ranging, as he suggests, under or north of 
the North Downs. Life must indeed have been most abundant and 
prolific in the Wenlock sea of the Ware locality; for so rieh in 
fossils are these Ware beds that no less than 33 species were 
obtained, all belonging to the Wenlock series, furnishing sufficient 
data in themselves to establish and determine the age of the rocks 
in which they occur. 

The five borings in the London area, or within the London basin, 
within a radius of 20 miles, Ware being the furthest removed, taking 
them in the order in which they were sunk, may be thus enume- 
rated :—No. 1. Kentish Town, 1300 feet deep; the London clay 
passed through was 350 feet thick, the Reading beds 50 feet, and 
the Thanet sands 15 feet, Upper Chalk 250 feet, Chalk-marl 30 feet, 
Upper Greensand 10 feet, Gault 60 feet, and 190 feet of red sandy 
rock believed to be Old Red Sandstone or a condition of the Devo- 
nian; the depth passed through was 1300 feet. 

No. 2. The Crossness new ‘well bore-hole about 1030 feet deep. 
No London c¢lay proper occurred in this sinking; the alluvial clay 
and gravel immediately below the ordnance level. was 20 feet thick, 
and rested upon the Woolwich and Reading and Thanet beds, here 
about 100 feet thick, the Chalk 620 feet, the Upper Greensand 


234 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


33 feet, the Gault 135 feet, succeeded by loose red, coarse, and fine 
sand, having all the appearance of the New Red Sandstone, and 
resembling the Kentish-Town Red series. 

The boring No. 3, the most important in geological results, 
was that at Messrs. Meux’s, Tottenham Court Road. The London 
Tertiaries here attained to 156 feet, the Upper Chalk 440 feet, 
the Lower Chalk and Chalk-marl 215 feet, the Gault and Upper 
Greensand 190 feet, the Neocomian (of peculiar type) 64 feet ; 
beneath this occurred the Upper Devonian shales at the depth of 
1064 feet, with characteristic fossils, Spirifera disjuncta, Hdmondia, 
Rhynchonella cuboides, Orthis sp., &e. This was the first indication 
of the presence of rocks older than the Neocomian east of North 
Devon and north of the latitude of London, clearly showing the 
easterly extension of the Paleozoic rocks from the western side of 
England, and extending towards Holland and Belgium. This boring 
therefore may be considered classical; it has revealed to us what 
was surmised intuitively by Mr. Godwin-Austen and Prof. Prest- 
wich. ‘Turnford and Ware have revealed other facts of nearly equal 
significance, and at the depth of 940 feet and 800 feet. 

The 4th boring is that at Turnford, 12 miles north of London, 
where the London Tertiaries are 100 feet thick, the Chalk 620 feet, 
the Upper Greensand 15 feet, the Gault 135 feet, the Neocomian, 
of the Carr-stone type, about 12 inches. We here again prove the 
position of the dark chocolate-coloured Upper Devonian rocks 
crowded with characteristic fossils; at the depth of 940 feet I 
obtained Spirifera disjuncta, Rhynchonella cuboides, Rhodocrinus, 
Strophomena rhomboidalis, Edmondia, Pterineea, <Aviculopecten, 
Modiola, Avicula damnoniensis, A. texturata, Fenestella, Tenta- 
culites, &c. 

The 5th trial or boring for water was at Ware, east of Hertford, 
and due north of the Turnford boring. ‘The boring commenced 
here in the Upper Chalk, which is 416 feet thick, followed by the 
Chalk-marl 125 feet, the Upper Greensand 77 feet, and the Gault 
160 feet, the Neocomian a trace only (8 inches), and of the Carr- 
stone type, resting upon an eroded surface of Upper Silurian Lime- 
stone. The probable relation of these beds to the typical Wenlock 
series of Wales and Shropshire I have discussed in the earlier 
portion of this section. No less than 20 species of Brachiopoda 
alone, with 13 other species, occur and are given in the list; and 
all were extracted from a core less than 3 feet in length and 1 foot 
in diameter. 

The species collected were the following :— 

I. Protozoa: Ischadites Kenigu, Murch. II. KcurnopErmata : 
Periechocrinus moniliformis and Taxocrinus sp. III. Annevipa: Ten- 
taculites ornatus, Sow. IV. Crustacta: Phacops caudatus, Brongn. 
MottuscaA Bracutopopa: Orthis canaliculata, Dalm.; O. elegantula, 
Dalm.; Meristella tumida, Dalm.; Cyrtia exporrecta, Wahl. ; Spirifera 
elevata, Dalm.; S. plicatella, Linn. ; Athyris sp.; Orania implicata, 
Sow. : fihynchonella cuneata, Dalm.; Atrypa reticularis ; Pentamerus 
faleatus, Dalm.; P. linguifer, Sow.; Strophomena euglypha, Dalm. : 


ANNIVERSARY ADDRESS OF THE PRESIDENT. 235 


S. reticulata, M‘Coy; S. depressa, Dalm.; S. rhomboidalis, Wahl. ; 
S. antiquata, Sow.; Chonetes sp.; Leptena sericea, Sow.; L. trans- 
versalis, Dalm. Concnirera: Ctenodonta sp.; Pterinea sp. ; 
Mytilus mytilimeris, Conr.; Orihonota rigida, Sow. GASTEROPODA : 
EHuomphalus rugosus, Sow. Crpnatopopa: Orthoceras attenuatum, 
Sow.; O. angulatwm, Wahl.; and Orthoceras sp. 

These five borings within the metropolitan area are of infinite 
scientific as well as economic value; and in addition the equally 
important trials at Burford due west of Ware, and that of North- 
ampton N.W. of London, open up important questions as to the 
deeply-seated structure of the triangular area having Ware and 
Burford at its base and Northampton at its apex. Have we within 
the rolls and folds of the Devonian rocks as they strike across 
England between latitude 51° and 52° any still undiscovered coal- 
tracts? is the Bristol coal-field repeated to the east and N.E. to 
Burford, with the underlying Devonian in place to the south-east ? 
It would appear so when we regard the bearing and strike of the 
older rocks towards the continent. The geological importance and 
interest of these borings is their revelation to us of the old Paleo- 
zoic land-surfaces, whether coast-lines or tablelands, theoretically 
enabling us to reconstruct the physiography of those land-masses, 
concealed, yet connected to now exposed areas, whether British, 
European, or even American. 


February 23, 1881. 


Rosert Etuerines, Esq., F.R.S., President, in the Chair. 


William Henry Goss, Esq., Rode Heath, Cheshire, and Stoke- 
upon-Trent, Staffordshire, was elected a Fellow of the Society. 


The following communications were read :— 


1. A letter from Dr. John Kirk, communicated to the Society by 
the Right. Hon. Earl Granville, dated :—- 


‘*H.M Agency and Consulate General, 
Zanzibar, December 20, 1880. 
‘¢ My Lorp, 

‘‘ It may be of interest to record the occurrence here of an 
earthquake-shock felt in the island of Zanzibar at 6.58 a.m., mean 
time, on the morning of the 18th inst. 

« Although the shock was very distinct, no damage appears to 
have been done to any buildings in town. 

‘Tt is now twenty-four years since a similar shock has been here 
noticed ; but on the mainland, especially in the vicinity of Ujiji, 
they are both more common and more severe than at the coast. 

‘Shortly after the cable was laid between Mozambique and 


236 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


Delagoa Bay, the communication was suddenly interrupted after 
one of these earthquake-shocks, which seems to have caused the 
falling in of rocks by which the cable was crushed. 
‘“¢ T have the honour to be &c., 
“ Joun Kirk, 

H.M. Agent and Consul-General, Zanzibar.” 
“ The Right Honourable 
Earl Granville, &c. &e., 

London.” 


2. “The Permian, Triassic, and Liassic Rocks of the Carlisle 
Basin.” By T. V. Holmes, Ksq., F.G.S. 


3. 6 On Astroconia Grant, a new Lyssakine Hexactinellid from 
the Silurian Formation of Canada.” By Prof. W. J. Sollas, M.A., 


The following objects were exhibited :— 


Specimens illustrating the transformation of Spodumene into 
Felspars, from Branchville, Connecticut, exhibited by H. Bauer- 
man, Esq., F.G.S. 


Canine of Deuterosaurus biarmicus, Eichw., from the Upper Per- 
mian of Kargalinsk Steppe, Bere exhibited by W. H. Twelvetrees, 
Ksq., F.G.S. 


Plates prepared for the illustration of the next part of Dr. A. 
Fritsch’s ‘Fauna der Gaskohle und der Kalksteine der Permfor- 
mation Bohmens.’ 


March 9, 1881. 
Rosert Erueriper, Esq., F.R.S., President, in the Chair. 


Robert Thompson Burnett, Esq., Irlams o’th’Height, near Man- 
chester; William Erasmus Darwin, Esq., B.A., Basset, South- 
ampton ; Charles James Fox, Ksq., 26 South Molton Street, W. ; 
and the Rev. T. Granger Hutt, M.A., Sedbergh, Yorkshire, were 
elected Fellows of the Society. 


The following communications were read :— 


1. “ Description of Parts of the Skeleton of an Anomodont Reptile 
(Platypodosaurus robustus, Ow.).—Part I. The gens 7%) By Prom 
Owen, C.B., F.R.S., F.G.8., &e. 


2. “On the Order Theriodontia, with a Description of a new 
Genus and Species (4lurosaurus felinus, Ow.).” By Prof. Owen, 
CoBS Ban Se EeGes: 


3. * Additional Observations on the Superficial Geology of British 
Columbia and adjacent regions.” By G.M. Dawson, Esq., D.Sc., 
F.GS. 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 237 


The following specimens were exhibited :— 


Skull of dlurosaurus felinus, exhibited by Prof. Owen, in illus- 
tration of his paper. 


Incisor tooth of Deuterosaurus biarmicus, Kichw.; and a Repti- 
lian canine from the Upper Permian Cupriferous Sandstones of 
Kargalinsk Steppe, near Orenburg, exhibited by W. H. Twelvetrees, 
Esq., F.G.S. 


March 23, 1881. 


R. Erxeries, Esq., F.R.S., President, in the Chair. 


Rey. Daniel Dutton, Sydney Street, Wellington, New Zealand ; 
and Capt. George Ernest A. Ross, 170 Cromwell Road, 8. W., were 
elected Fellows of the Society. 


The following communications were read :-— 


1. “The Upper Greensands and Chloritic Marl of the Isle of 
Wight.” By C. Parkinson, Esq., F.G.8. ~ 


2. “On the Flow of an Ice-sheet, and its connexion with 
Glacial Phenomena.” By Clement Reid, Esq., F.G.S.* 


3. “ Soil-cap Motion.” By R. W. Coppinger, Esq. Communicated 
by the President. 


The following specimens were exhibited :— 


A large pebble from the Gault of Folkestone, exhibited by J. 8. 
Gardner, Esq., F.G.S. 


Some recent Centipedes, killed near Upata, 60 miles 8.E. of Las 
Tablas, on the Orinoco River, exhibited by G. Attwood, Esq., 
NG:S. 


April 6, 1881. 
J. W. Horxs, Esq., F.R.S., Vice-President, in the Chair. 


Edward F. Boyd, Esq., Moor House, Leamside, Fence Houses ; 
Herbert de Haga Haig, Esq., Lieut. R.E., Staff College, Camberley, 
Farnboro’ Station; J. C. Margetson, Esq., 2 Spring Hill, Kings- 
down, Bristol; Edward David Price, Esq., Collegiate School, 
Hounslow; and James Tonge, Esq., Woodbine House, West- 
houghton, Bolton-le-Moors, were elected Fellows of the Society. 


* This paper has been withdrawn by the author with the permission of the 
Council. 
VOL. XXXVII. § 


238 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


The following communications were read :— 


1. “ The Microscopic Characters of the Vitreous Rocks of Montana, 
US.” By F. Rutley, Esq., F.G.S.; with an Appendix by James 
Eccles, Esq., F.G.8. 


2. “On the Microscopic Structure of Devitrified rocks from Bedd- 
gelert, Snowdon, and Skomer island.” By F. Rutley, Esq., F.G.S. 


3. “The Date of the last Change of Level in Lancashire.” . By 
T. Mellard Reade, Esq., C.E., F.G.S. 


The following specimens were exhibited :— 


Specimens of arsenical silver-ore from Gunnison, Colorado, ex- 
hibited by H. Bauerman, Esq., F.G.S. 


Rocks and rock-sections, exhibited by F. Rutley, Esq., in illustra- 
tion of his papers. 


April 27, 1881. 


R. Erxeriner, Esq., F.R.S., President, in the Chair. 


Samuel Gerrard Kirchhoffer, Esq., M.A., Yately Grange, Farn- 
boro’ Station; Arthur Henry Shakespere Lucas, Esq., The Leys, 
Cambridge ; and Frederick Thomas Nelson Spratt, Esq., Lieut. R. E. 
Clare Lodge, Tunbridge Wells, were elected Fellows of the Society. 


The following communications were read :— 


1. “On the precise Mode of Accumulation and Derivation of the 
Moel Tryfan Shelly Deposits ; on the Discovery of similar high-level 
Deposits along the Eastern Slopes of the Welsh mountains ; and on 
the Existence of Drift-zones showing probable Variations in the Rate 
of Submergence.” By D. Mackintosh, Esq., F.G.S. 


2. “On the Correlation of the Upper Jurassic rocks of England 
with those of the Continent.” By the Rev. J. F. Blake, M.A., 
F.G.S. Part I. The Paris basin. 


3. “On Fossil Chilostomatous Bryozoa from the Yarra-Yarra, 
Victoria, Australia.” By Arthur William Waters, Esq., F.G.S. 


The following specimens were exhibited :— 


Upper molar of an extinct species of Elephant from the Sivélic 
Older Pliocene (?), and Plant-remains in ironstone nodules from Coal- 
brook Dale, exhibited by E. Charlesworth, Esq., F.G.S. 


Specimens from Moel Tryfan and Frondeg gravel-pit, Denbigh- 
shire, exhibited by D. Mackintosh, Ksq., in illustration of his paper. 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 239 


May 11, 1881. 
R. Erusrivger, Esq., F.R.S., President, in the Chair. 


Joseph Deeley, Esq., Ruabon, North Wales; George Kilgour, Ksq., 
C.E., F.R.A.S., Dutoit’s Pan, Griqualand West, South Africa; and 
Roderick William MacLeod, Esq., Bengal Staff Corps, 55 Parliament 
Street, W., were elected Fellows of the Society. 


The List of Donations to the Library was read. 


The following communications were read :— 


1. “ Notes on the Fish-remains of the Bone-bed at Aust, near 
Bristol, with the Description of some new Genera and Species.” By 
James W: Davis, Esq., F.S8.A., F.G.S. 


2. “On some Fish-spines from the Coal-measures.” By J. W. 
Davis, Esq., F.S.A., F.G.S. 


3. “On some specimens of Diastopora and Stomatopora from the 
Wenlock Limestone.” By Francis D. Longe, Esq., F.G.S. 


[ Abstract. ] 


Mr. Longe showed and described some specimens of Bryozoa from 
the Wenlock Limestone of Dudley, which he compared with corre- 
sponding forms from the Oolites and later periods, and pointed out 
the close similarity of the Silurian with the later forms, in respect 
of the shape and dimensions of the cells, as well as in the habit of 
ccencecic growth. 

Alluding to some other Palzozoic forms, assigned to the Bryozoa 
under the generic names of Berenicea and Ceramopora, he pointed 
out the difference between the shape of the cells in these forms and 
those which he had described, and expressed a doubt whether they 
should be classed as Bryozoa at all. 

On the other hand, he referred to some specimens described by 
Professor Nicholson (Ann. & Mag. Nat. Hist. vol. xv. 1875) under 
the names of Alecto auloporoides &c. as having the true Bryozoan 
cell, and furnishing additional evidence of the existence in the 
Silurian seas of forms of Bryozoa which, though very abundant in 
the Oolites and at all subsequent periods, were not generally sup- 
posed to have existed in the Paleozoic period. 


DiscussIon. 


The Prestpenr stated that the genus Aulopora had never been 
referred to the Bryozoa before, but to the Actinozoa. He demurred 
to the use of the term “‘ plant” as applied to the Bryozoa. 

The Avruor stated that the object of his paper was to prove that 
in the Wenlock beds he had found forms quite undistinguishable 
from Stomatopora or Alecto, and Diastopora or Berenicea of later 
periods. He had not confounded Aulopora with the Bryozoa. 


240 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


4, “On a new Species of Plesiosaurus (P. Conybeart) from the 
Lower Lias of Charmouth, with Observations on P. megacephalus, 
Stutchbury, and P. brachycephalus, Owen.” By Prof. W. J. Sollas, 
M.A., F.R.S.E., F.G.8., &e., Professor of Geology in University 
College, Bristol ; accompanied by a Supplement on the Geological 
Distribution of the Genus 2 Plesiosaurus, by G. F. Whidborne, Esq., 
M.A., F.G.S. 


5. “On certain Quartzite and Sandstone Fossiliferous Pebbles in 
the Drift in Warwickshire, and their probable identity with the true 
Lower-Silurian Pebbles, with similar fossils, in the Trias at Bud- 
leigh Salterton, Devonshire.” By the Rey. P. B. Brodie, M.A., 
E.G.8. 


The following specimens were exhibited :— 


Fish-spines and Teeth, exhibited by J. W. Davis, ple F.G.S., in 
illustration of his papers ; and 

Specimens of Diastopora and Stomatopora, onunead by F. D. 
Longe, Esq., F.G.S., in illustration of his paper. 


May 25, 1881. 
R. Ernermer, Ksq., F.R.S., President, in the Chair. 


Rey. Tom Bullock Hardern, M.A., LL.M., Burnham Overy, Lynn, 
Norfolk, was elected a Fellow of the Society. 


The following specimens were presented to the Museum :—The 
type specimens illustr ating the ‘* Note on the ‘Tubulations Sableuses’ 
of the ‘ Etage Bruxellien? in the Environs of Brussels” by H. J. 
Carter, Esq., F.R.S. (Ann. & Mag. Nat. Hist. May 1877), who 
presented them. 


The following communications were read :— 


1. “On the Discovery of some Remains of Plants at the base of 
the Denbighshire Grits, near Corwen, North Wales.” By Henry 
Hicks, M.D., F.G.8. With an Appendix by R. Etheridge, Esq., 
F.R.S., Pres. Geol. Soe. 


2. “Notes on a Mammalian Jaw from the Purbeck Beds at 
Swanage, Dorset.” By Edgar W. Willett, Esq. Communicated by 
the President. 


The following specimens were exhibited :— 


Specimens of the Reptile Fauna of the Gosau Formation, exhibited 
by Prof. H. G. Seeley, F.R.S., F.G.8.; and 

Specimens exhibited by Messrs. Hicks and Willett in illustration 
of their papers. 


a eee 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 241 


June 8, 1881. 
R. Erurriver, Esq., F.R.S., President, in the Chair. 


The Meeting was made a Special General Meeting for the election 
of a Member of the Council in the room of the late Sir P. de Malpas 
Grey-Egerton, Bart., M.P., F.R.S., F.G.S. 


The Presipent announced that the late Sir Philip Egerton had 
bequeathed to the Society all the original drawings made from 
specimens in the collection of the Earl of Enniskillen for the illus- 
tration of Prof. Agassiz’s works on Fossil Fishes. The Society had 
long possessed the drawings made for the same purpose from the 
Earl of Ellesmere’s collection, and some years ago the Earl of 
Enniskillen presented those which had been prepared from speci- 
mens in the possession of Sir Philip Egerton. Sir Philip Egerton’s 
kind bequest would complete this interesting series. 

Mr. Joun Evans remarked that on this, as on so many other 
occasions, the Society was deeply indebted to the kindness of the 
late Sir Philip Egerton, and proposed that the President and Secre- 
taries should be instructed to communicate with Sir Philip’s repre- 
sentatives, and to express the gratitude of the Society for this 
bequest and their condolence with his relatives on the loss they 
have sustained. 

Prof. W. Boyp Dawxrys seconded this proposal, which was carried 
unanimously. 


The ballot fora new member of Council was kept open till 9 
o'clock, when Sir John Lubbock, Bart., M.P., F.R.S., was declared 
to have been elected. 


Grenville A. J. Cole, Esq., Mayland, Sutton, Surrey, and J. L. 
Roberti, Hsq., 92 Malpas Road, New Cross, 8.H., were elected 
Fellows; and I] Commendatore Quintino Sella, of Rome, a Foreign 
Member of the Society. 


The following names of Fellows of the Society were read out for 
the first time in conformity with the Bye-laws, Sect. VI. B, Art. 6, 
in consequence of the non-payment of the arrears of their contribu- 
tions :—J. Entwisle, Esq.; R. Koma, Hsq.; W.H. Le Feuvre, Esq.; 
C. 8. Mann, Esq. ; Joseph Thompson, Esq. 


The following communications were read :— 


1. “The Reptile-Fauna of the Gosau Formation, preserved in the 
Geological Museum of the University of Vienna.” By Prof. H. G. 
Seeley, F.R.S., F.LS., F.G.S8.; with a Note on the Geological 
Horizon of the Fossils, by Prof. Edward Suess, F.M.G.S. 


2. “On the Basement-beds of the Cambrian in Anglesey.” By 
Prof. T. McKenny Hughes, M.A., F.G.8. 
VOL. XXXVII. t 


242 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 


3. “* Description and Correlation of the Bournemouth Beds.— 
Part II. Lower or Freshwater Series.” By J. 8S. Gardner, Esq., 
F.G.S. 


Specimens were exhibited by Messrs. Seeley, Hughes, and Gardner, 
in illustration of their papers. 


June 22, 1881. 
R. Erurrives, Esq., F.R.S., President, in the Chair. 


Thomas Hart, Esq., Richmond Terrace, Blackburn; and David 
William Jones, Esq., Coronel, Chili, South America, were elected | 
Fellows of the Society. | 


Specimens of Tertiary Brachiopods from South Australia, and 
three specimens of Mesozoic Belemnites from Central Australia, were 
presented to the Museum by Prof. R. Tate, F.G.S. 


_ The following names of Fellows of the Society were read out for 
the second time in conformity with the Bye-laws, Sect. VI. s, Art. 6, 
in consequence of the non-payment of the arrears of their contribu- 
tions :—J. Entwisle, Esq.; R. Koma, Esq.; W.H. Le Feuvre, Esq.; 
C. 8. Mann, Esq. ; Joseph Thompson, Esq. 


The following communications were read :— 


1. “ Description of a new Species of Coral from the Middle Lias 
of Oxfordshire.” By R. F. Tomes, Hsq., F.G.S. 


2. “ Note on the Occurrence of the Remains of a Cetacean in the 
Lower Oligocene Strata of the Hampshire Basin.” By Prof. J. W. 
Judd, F.R.S.; Sec. G.S. With a Note by Prof. H. G. Seeley, F.R.S., 


3. ‘Description of a Peat-bed interstratified with the Boulder- 
drift at Oldham.” By G. H. Hollingworth, Ksq., F.G.S. 


4. “Silurian Uniserial Stomatopore and Ascodictya.” By G. R. 
Vine, Esq. Communicated by Prof. P. Martin Duncan, F.R.S., 


5. “Note on the Diamond-fields of South Africa.” By. EH. J. 
Dunn, Esq. Communicated by Prof. Ramsay, F.R.S., F.G.S. 


6. “On a new Comatula from the Kelloway Rock.” By P. H. 
Carpenter, Esq., M.A., Assistant Master at Eton College. Commu- 
nicated by the President. 


PROCEEDINGS OF THE GEOLOGICAL SOCIETY. 243 


7. “Descriptive Catalogue of Ammonites from the Sherborne 
District.” By Sydney 8. Buckman, Esq. Communicated by Prof. 
J. Buckman, F.G.S., F.L.S., &c. 


The following specimens were exhibited :— 


Prehistoric remains (pottery &c.) found by J. B. Andrews, Esq., 
of Mentone, at St. Vallier, near Grasse, in the neighbourhood of 
dolmens &c. 

Specimens exhibited by Messrs. Tomes and Judd in illustration of 
their papers. 


t2 


ADDITIONS 


TO THE 


LIBRARY AND MUSEUM OF THE GEOLOGICAL SOCIETY. 
Session 1880-81. 


I. ADDITIONS TO THE LIBRARY. 
1. PreriopiIcaLts AND Pusiicatrions oF LEARNED SOCIETIES. 


Presented by the respective Societies and Editors, if not otherwise 
stated. 


Academy, The. Nos. 425-451. 1880. 
The British Association at Swansea, 157, 176, 190. 


Nos. 452-478. 1881. 


Adelaide. Philosophical Society. Transactions and Proceedings and 
Report for 1878-79. 1879. 

R. Tate. Anniversary Address, xxxix.—G. Scoular. The Geology of 
the Hundred of Munno Para, Part 1, 60—O. Tepper. Introduction to 
the Cliffs and Rocks at Androssan, Yorke’s Peninsula, 71.—R. Tate. 
The Natural History of the Country around the Head of the Great Aus- 
tralian Bight, 94.—R. Tate. Zoologica et Paleontologica Miscellanea, 
chiefly relating to South Australia, 129. 


——. Royal Society of South Australia. Transactions and 
Proceedings and Report. Vol. i., for 1879-80. 1880. 

R. Tate. Anniversary Address, 39.—J. E. Tenison-Woods. On some 
recent and fossil Australian Selenariads, 1—Otto Tepper. ‘The “ Bay 
of Biscay’ Soil of South Australia and its Formation, 91.—J. Hi. Tenison- 
Woods. Onsome new Corals from the Australian Tertiaries, 99.—R. Tate. 
Description of a new Species of Belemnite from the Mesozoic Strata of 
Central Australia, 104.—G. Scoular. The Geology of the Hundred of 
Munno Para, Part 2, 106.—R. Tate. On the Australian Tertiary Pal- 
hobranchs, 140.—R. Tate. Rock-formations and Minerals in the vicinity 
of Peake, C. A., 179.—W. Fowler. Sections of Strata traversed by two 
wells at Yarrow, in the Hundred of Clinton, 181. 


Albany. New-York State Museum of Natural History. Annual 
Reports, Nos. 28 (Museum edition, 1879), 29-31. 1878-79. 
Presented by James Hall, Esq., F.M.G.S. 


Analyst, The. Vol. v. Nos. 52-57. 1880. 
Vol. vi. Nos. 58-63. 1881. - 


Annals and Magazine of Natural History. Ser. 5. Vol. vi. Nos. 
31-36. 1880. Purchased. 

C. Lapworth. On the Geological Distribution of the Rhabdophora, 16. 

—A. W, Waters. Note on the Genus Heteropora, 156.—C. Lapworth. 

On the Geological Distribution of the Rhabdophora, 185.—H. J. Carter. 


=i 


ADDITIONS TO THE LIBRARY. 245 


On fossil Sponge-spicules from the Carboniferous Strata of Ben Bul- 
ben, near Sligo, 209.—G. Cotteau. On the Tertiary Echinida of 
Beleium, 246.—R. Etheridge, Jun. Notes on the Gasteropoda contained 
in the Gilbertson Collection, British Museum, and figured in Phillips’s 
‘Geology of Yorkshire,’ 289.—A. J. Jukes-Brown. The Chalk Bluffs of 
Trimmingham, 305.—H. Alleyne Nicholson. On the Minute Structure of 
the recent Heteropora neozelanica, Busk, and on the Relations of the 
Genus Heteropora to Monticulipora, 329, 414.—A. Agassiz. On Paleon- 
tological and Embryological Development, 348.—J. W. Davis. On a 
Species of Gyracanthus, a fossil Fish from the Coal-measures, 372.—W. 
J. Sollas. On the Flint Nodules of the Trimmingham Chalk, 384, 437. 


Annals and Magazine of Natural History. Ser. 5. Vol. vu. Nos. 
37-42. 1881. Purchased. 

H. A. Nicholson. On some new or imperfectly-lnown Species of 
Corals from the Devonian Rocks of France, 14.—R. Etheridge, Jun. 
Descriptions of certain peculiar Bodies which may be the Opercula of 
small Gasteropoda, discovered by Mr. James Bennie in the Carboniferous 
Limestone of Law Quarry near Dalry, Ayrshire, with Notes on some Si- 
lurian Opercula, 25.—W. J Sollas. Note on the Occurrence of Sponge- 
spicules in Chert from the Carboniferous Limestone of Ireland, 141.—G. 
C. Wallich. On the Origin and Formation of the Flints of the Upper or 
White Chalk; with Observations upon Prof. Sollas’s Paper in ‘ The Annals 
and Magazine of Natural History’ for December 1880, 162.—S. H. Scudder. 
Relation of Devonian Insects to Later and Existing Types, 255.—G. C. 
Wallich. On Siliceous Sponge-growth in the Cretaceous Ocean, 261.—P. 
H. Carpenter and R. Etheridge, Jun. Contributions to the Study of the 
British Paleeozoic Crinoids, 281.—H. J. Carter. On the Kunker Formation 
of the Alluyium in India compared with the Flint Formation in the Chalk 
of England, 308.—H. Filhol. The Bears of the Cavern of Lherm, 428.— 
J. W. Davis. On Paleospinax priscus, Egerton, 429.—S. H. Scudder. 
The Structure and Affinities of Euphoberea, Meek and Worthen, a Genus 
of Carboniferous Myriopoda, 437.—O. C. Marsh. Discovery of a fossil 
Bird in the Jurassic of Wyoming, 488. 

Atheneum (Journal). Nos. 2748-2774. 1880. 

A. C. Ramsay. Address delivered at the Swansea Meeting of the 
British Association for the Advancement of Science, August 25, 1880, 263. 
—British Association at Swansea, 509. 

- Nos. 2775-2799. 1881. 
Sir Philip de Malpas Grey Egerton, Bart., M.P., 494.—The Hamad, 817. 


Parts 630-636. 1880. 
Parts 637-640. 1881. 


Barnsley. Midland Institute of Mining, Civil, and Mechanical 
Engineers. Transactions. Vol. vii. Part 50. 1880. 
T. W. Embieton. On an Outburst of Carbonic Acid Gas in the Roche- 
belle Coal Mine, France, 89. 


Volo vis Rarts 5Jand 53. “lee: 


Basel. Schweizerische palaontologische Gesellschaft. Abhandlungen. 
Vol. vu. 1880. Purchased. 

Forsyth Major. SBeitrige zur Geschichte der fossilen Pferde——Ph. 
de la Harpe. Htude des Nummulites de la Suisse et révision des espéces 
éocénes des genres Nummulites et Assilina.—L. Rutimeyer. Beitrage zu 
einer natiirlichen Geschichte der Hirsche——F. Koby. Monographie 
des polypiers jurassiques de la Suisse.—P. de Loriol. Monographie pa- 


246 ADDITIONS TO THE LIBRARY. 


léontologique des couches de la zone & Ammonites tenutiobatus (Badener 
Schichten) d’Oberbuchsitten et de Wangen (Soleure).—P. de Loriol. 
Description de quatre échinodermes nouveaux. 


Belfast Natural-History and Philosophical Society. Proceedings for 
the Sessions 1878-79, 1879-80. 1880. 
‘J.J. Murphy. Lunar Volcanoes, 124.—J. J. Murphy. On the Origin 
of Mountains and Volcanoes, 148.—W. H. Patterson. Flint Implements 
&e. found at Ballymisert, Co. Down, 165. 


Berlin. Deutsche geologische Gesellschaft. Zeitschrift. Band xxxii. 
Heft 2-4 (1880). 1881. 

W. Pabst. Untersuchung von chinesischen und japanischen zur Por- 
zellanfabrication verwandten Gesteinsvorkommnissen, 223.—A. Heim. 
Zum ‘ Mechanismus der Gebirgsbildung, 262.—F’. Noetling. Die Ent- 
wickelung der Trias in Niederschlesien, 300.—F. Sandberger. Ueber die 
Bildung von Erzgangen mittelst Auslaugung des Nebengesteins, 350.— 
Hi. Dewitz. Ueber einige ostpreussische Silurcephalopoden, 371.—G. 
Stemmmann. Mikroskopische Thierreste aus dem deutschen Kohlenkalke 
(Foraminiferen und Spongien), 394.—W. Branco. Beobachtungen an 
Aulacoceras v. Hauer, 461.—F. Klockmann. Ueber Basalt-, Diabas- und 
Melaphyr-Geschiebe aus dem norddeutschen Diluvium, 408.—P. Grigo- 
riew. Der Meteorit von Rakowska im Gouvernement Tula in Russland, 
417.—Rothpletz. Radiolarien, Diatomaceen und Spharosomatiten im 
silurischen Kieselschiefer von Langenstriegis in Sachsen, 447.—A. Neh- 
ring. Uebersicht tiber vierundzwanzig mitteleuropaische Quartar-Faunen, 
468.—E. Geinitz. Der Jura von Dobbertin in Mecklenburg und seine 
Versteinerungen, 510.—Fr. Pfaff. Hinige Beobachtungen tiber den Loch- 
seitenkalk, 536.—Fr. Pfaff. Hinige Bemerkungen zu Herrn Heim’s Auf- 
satz ‘ Zum Mechanismus der Gebirgsbildung,” 542.—G. Holm. Bemer- 
kungen tiber Illenus crassicauda Wahlenberg, 559.—H. Credner. Ueber 
Glacialerscheinungen in Sachsen, nebst vergleichenden Vorbemerkungen 
uber den Geschiebemergel, 572.—W. Branco. Ueber die Verwandtschafts- 
verhaltnisse der fossilen Cephalopoden, 596.—Huyssen. Uebersicht der 
bisherigen Ergebnisse der vom Preussischen Staate ausgefuhrten Tief boh- 
rungen im norddeutschen Flachland und des bei diesen Arbeiten verfolgten 
Planes, 612.—Jentzsch. Uebersicht der silurischen Geschiebe Ost- und 
Westpreussens, 623.—W. Dames. Ueber Cephalopoden aus dem Gault- 
quader des Hoppelberges bei Langenstein unweit Halberstadt, 685.—M. 
Hoyer. Ueber das Vorkommen von Phosphorit- und Grunsand-Geschieben 
in Westpreussen, 698.—H. Kayser. Dechenella, ee devonische Gruppe 
der Gattung Phillipsia, 703.—C. Rammelsberg. Ueber die Vanadinerze 
aus dem Staat Cordoba in Argentinien, 708.—Max Bauer. Dioptas aus 
den Cordilleren von Chili, 714.—Max Bauer. Nochmals die Krystallform 
des Cyanit, 717.—C. Struckmann. Ueber die Verbreitung des Renthiers 
in der Gegenwart und in alterer Zeit nach Maassgabe seiner fossilen Reste 
unter besonderer Beriicksichtigung der deutschen Fundorte, 728.—F. 
Wahnschaffe. Ueber Gletschererscheinungen bei''Velpke und Daundorf, 
774.—Otto Lang. Ueber den Gebirgsbau des Leinethales bei Gottingen, 
799, : 


——. Gesellschaft naturforschender Freunde. Sitzungsberichte, 
1880. 1880. 

K. A. Lossen. Vorlegung und Besprechung von Handstucken und 
Diinnschliffen metamorphosirter Eruptiv- bezw. Tuff-Gesteine vom Schma- 
lenberg bei Harzburg, 1.—O. Reinhardt. Ueber die zum Subgenus Or- 
cula, Held, gehorigen Pupa-Arten und deren geographische Verbreitung, 
12.—yv. Martens. Ueber vorspringende Linien an der Innenflache einiger 


ADDITIONS TO THE LIBRARY. 247 


Muschelschalen, welche die Lage der Kiemen markiren, 22, 59.— 
S. Schliiter. Ueber Zoantharia rugosa aus dem rheinischen Mittel- und 
Ober-Devon, 49. 


Berlin. K6niglich preussische Akademie der Wissenschaften. Mo- 
natsbericht. March to December 1880. 1880-81. 
Rammelsberg. Ueber die Zusammensetzung des Descloizits und der 
naturlichen Vanadinverbindung en itberhaupt, 652. — Rammelsberg. 
Ueber die Zusammensetzung des Pollucits von Hlba, 669. — Websky. 
Ueber die Krystallform des Descloizit, 672. 


d . September and October 1880. 1881. 
Websky. Ueber die Krystallform des Vanadinits von Cordoba, 779. 


Zeitschrift fiir die gesammten Naturwissenschaften. Band lili. 
1880. 

K. Pettersen. Scheuerungserscheinungen in der gegenwartigen Lit- 
toralzone, 247.—P. Kaiser. F%coxylon bohemicum: ein neues fossiles 
Laubholz, 309.—L. Besthorn. Der Kohnstein bei Nordhausen, 340.— 
O. Luedecke. Ueber einen Anorthithasalt vom Fujino-yama, in Japan, 
410.—O. Luedecke. Ueber Roemerit, Grailich, 417—H. Pohlig. Zur 
Beantwortung der Frage nach Entstehung der “krystallinischen Schiefer,” 
445.—K. Pettersen. Terrassen und alte Strandlinien, 783.—R. Collett. 
Glaciale Mergelknollen mit Fischrest-Hinschlissen aus Beieren im nord- 
lichen Norwegen, 839. 


Bern. Schweizer Alpenclub. Jahrbuch, XV. Jahrgang (1880), and 
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H. vy. Fellenberge. Geologische Wanderungen im Rhonegebiet 1878 
und 1879, 268. 


Birmingham. Mason Science College. Calendar for the Session 
1880-81. 1881. 


Birmingham Philosophical Society. Proceedings, 1878-79. Vol. 1. 
No. 3. 1879. Presented by W. Whitaker, Esq., F.G.S. 

H. W. Crosskey. Notes on some of the Glacial Phenomena of the 
Vosges Mountains, with an Account of the Glacier of Kertoff, 58.—T. G. 
Bonney. The Structure and Distribution of the Pre-Cambrian Rocks of 
Britain, 140. 

Bordeaux. Société Linnéenne. Actes. Série4. Tome ii. Livr. 3 et 
4, 1879. 


Boston. American Academy of Arts and Sciences. Proceedings. 
Vol. xv. Parts 1 & 2. 1880. 


Boston Society of Natural History. Memoirs. Vol. i. Partl. No.3. 
1879. 
S. H. Scudder. Palzeozoic Cockroaches: a complete Revision of the 
Species of both Worlds, with an Essay toward their Classification, 23. 


Occasional Papers. III. 8vo. 1880. 


W. O. Crosby. Contributions to the Geology of Eastern Massachu- 
setts, 1. 


Proceedings. Vol. xx. Parts 2 & 3 (1878-80). 1879-80. 
N.S. Shaler. Evidences of a Gradual Passage from Sedimentary to 
Volcanic Rocks in the Brighton District, 129. == ©. Crosby. Occur- 
rence of fossiliferous Boulders in the Drift of Truro, Cape Cod, 136.— 
T. Sterry Hunt. Remarks on the Precambrian Rocks of Great Britain, 


248 ADDITIONS TO THE LIBRARY. 


140.—W. O. Crosby. A possible Origin of Petrosiliceous Rocks, 160.— 
G. F. Wright. The Kames and Moraines of New England, 210.—W. 
Upham. Glacial Drift of Boston and Vicinity, 220.—S. Kneeland. On 
the Mineralized Phosphatic Guanos of the Equatorial Pacific Islands, 
935.—N. 8. Shaler. Notes on the Submarine Coast-shelf, or Hundred 
Fathom Detrital Fringe, 278.—M. EH. Wadsworth. Danalite from the 
Tron Mine, Bartlett, N. H., 284M. E. Wadsworth. Picrolite from a 
Serpentine Quarry in Florida, Mass., 286.—W. H. Melville. Analysis of 
the above Picrolite, 287.—J. H. Huntington. On the Iron Ore of Bart- 
lett, N. H., 288.—W. O. Crosby. Evidences of Compression in the Rocks 
of the Boston Basin, 308.—S. H. Scudder. Probable Age of Haulover 
Beach, Nantucket Harbour, 329. 


Brighton and Sussex Natural-History Society. Annual Report, 
1879-80. 1880. Presented by W. Whitaker, Esq., F.GS. 


Bristol Museum and Library. Report of Proceedings at the Tenth 
Annual Meeting. 1881. 


British Association for the Advancement of Science. Report of the 
Fiftieth Meeting, held at Swansea, 1880. 1880. 

Everett. Thirteenth Report of the Committee appointed for the pur- 
pose of investigating the Rate of Increase of Underground Temperature 
downwards in various Localities of Dry Land and under Water, 26.— 
W. Pengelly. Sixteenth and concluding Report of the Committee ap- 
pointed for the purpose of exploring Kent’s Cavern, Devonshire, 62.— 
H. G. Seeley. Report on the Mode of Reproduction of certain Species of 
Ichthyosaurus from the Lias of England and Wurtembere, 68.—G. R. 
Vine. Report of the Committee appointed for the purpose of reporting 
on the Carboniferous Polyzoa, 76.—C. E. De Rance. Sixth Report of the 
Committee appointed for investigating the Circulation of the Underground 
Waters in the Permian, New Red Sandstone, and Jurassic Formations 
of England, and the Quantity and Character of the Water supplied to 
Towns and Districts from those Formations, 87.—W. H. Baily. Second 
Report of the Committee appointed for the purpose of collecting and 
reporting on the Tertiary (Miocene) Flora, &c., of the Basalt of the North 
of Ireland, 107.—H. W. Crosskey. Eighth Report of the Committee 
appointed for the purpose of recording the Position, Height above the Sea, 
lithological Characters, Size, and Origin of the Erratic Blocks of England, 
Wales, and Ireland, reporting other matters of interest connected with 
the same, and taking measures for their Preservation, 110.—First Report 
of the Committee appointed for the purpose of exploring the Caves of the 
South of Ireland, 209.—C. H. Perkins. On the Anthracite Coal and 
Coal-field of South Wales, 220.—Geddes. Report of the Committee for 
conducting Palzontological and Zoological Researches in Mexico, 254.— 
J. Gwyn Jeffreys. The French Deep-sea Exploration in the Bay of 
Biscay, 378.—W. Whitaker. List of Works on the Geology, Mineralogy, 
and Palsontology of Wales (to the end of 1875), 397.—H. C. Bolton. 
On the Application of Organic Acids to the Examination of Minerals, 505. 
—H.C. Sorby. Presidential Address to Section C, 565.—W. Boyd Daw- 
kins. On the Action of Carbonic Acid on Limestone, 573.—F. C. J. 
Spurrell. On the Site of a Paleolithic Implement Manufactory at Cray- 
ford, Kent, 574.—G. H. Kinahan. On the Hiatus said to have been 
found in the rocks of West Cork, 574.—W. H. Dalton. Note on the 
Range of the Lower Tertiaries of Hast Suffolk, 575.—J. P. O'Reilly. On 
the Relation to be established between Coast-line Directions represented. 
by Great Circles on the Globe, and the Localities marked by Earthquakes 


= ae 


ADDITIONS TO THE LIBRARY. 249 


in Europe, 576.—W. J. Sollas. On the Island of Torghatten, 576.—W. 
J. Sollas. On a Fragment of Mica Schist, 577.—W. T. Blanford. On the 
Geological Age and Relations of the Sewalik and Pikermi Vertebrate and 
Invertebrate Faunas, 577.—E. Wethered. On the Sandstone and Grits 
of the Lower and Middle Series of the Bristol Coalfield, 579.—J. Prest- 
wich. Ona Raised Beach in Rhos Sili Bay, Gower, 581.—J. Prestwich. 
On the Geological Evidence of the temporary Submergence of the South- 
west of Europe during the early Human Period, 581.—C. Moore. Proofs 
of the Organic Nature of Zozoon canadense, 582.—W. J. Sollas. On a 
striated Stone from the Trias of Portishead, 586.—W. J. Sollas. On the 
Action of a Lichen on Limestone, 586.—W. J. Sollas. On Sponge- 
spicules from the Chalk of Trimmingham, Norfolk, 586.—G. M. Dawson. 
Sketch of the Geology of British Columbia, 588.—H. H. Godwin-Austen. 
On the Post-Tertiary and more recent Deposit of Kashmir and the Upper 
Indus Valley, 589.—R. B. Foote. Notes on the Occurrence of Stone Im- 
plements in the coast Laterite south of Madras, and in high-level Gravels 
and other Formations in the South Mahratta Country, 589.—C. E. De 
Rance. On the Pre-Glacial Contours and Post-Glacial Denudation of the 
North-west of England, 590—Schaffhausen. On Prehistoric Times in 
the Valley of the Rhine, 624.—H. Stopes. On a Paleolithic Stone Imple- 
ment from Egypt, 624.—H. Stopes. On a Paleolithic Flint Implement 
from Palestine, 624. 


Brussels. Académie Royale des Sciences. Annuaire. 1879, 1880, 
and 1881. 1879-81. 


: . Bulletins. Série 2. Tome xlvi. (1878). 1878. 
C. Malaise. Découverte de Brachiopodes du genre Lingula dans le 
Cambrien du massif de Stavelot, 58.—C. Malaise et L. de Koninck. 
Rapport sur le mémoire de M. A. Renard: “La diabase de Challes, 
prés de Stavelot,’ 186, 188.—C. Malaise. Rapport sur le mémoire 
de M. L. de Koninck: “Sixiéme notice sur les minéraux belges,” 189, 
—A. Renard. La diabase de Challes, prés de Stavelot, 228.—L. L. 
de Koninck. Sixiéme et septiéme notices sur les minéraux belges, 240 
et 245.—H. Dupont. Rapport sur le mémoire de M. A. Renard: “ Re- 
cherches lithologiques sur les phthanites du calcaire carbonifére de la 
Belgique,” 325.—E. Dupont. Sur la découverte d’ossements d’Iguanodon, 
de poissons et de végétaux dans la-fosse Sainte-Barbe du charbonnage de 
Bernissart, 387.—A. Renard. Recherches lithologiques sur les phthanites 
du calcaire carbonifére de Belgique, 471.—E. Dupont. Sur les alluvions 
torrentielles qui se déposent de nos jours sur les plateaux de l’ Entre-Sambre- 
et-Meuse et du Condroz, 643.—C. Malaise. Sur une espéce minérale 
- nouvelle pour la Belgique: l’Arsénopyrite ou Mispickel, 881. 


: : : Tome xlvu. (1879). - 1879. 

C. Malaise. Sur l’Arsénopyrite ou Mispickel et sur l’eau arsénicale de 
Court-Saint-Etienne, 29.—P. J. Van Beneden. Sur un enyoi d’ossements 
de Cétacés fossiles de Croatie, 183.—C. Malaise. Rapport sur le mémoire 
de M. A. Renard: ‘ Des caractéres distinctifs de la dolomite et de la cal- 
cite dans les roches calcaires et dolomitiques du calcaire carbonifére de 
Belgique,” 492.—A. Renard. Des caractéres distinctifs de la dolomite 
et de la calcite dans les roches calcaires et dolomitiques du calcaire car- 
bonifére de Belgique, 541.—L. L. de Koninck. Huitiéme et neuviéme 
notices sur les minéraux belges, 564 et 568. 


: : ? Tome xlviii. (1879). 1879. 
Gilkinet. Du développement du régne végétal dans les temps géolo- 
giques, 814. 


es 


250 ADDITIONS TO THE LIBRARY. 


Brussels. Académie Royale des Sciences. Bulletins. Série 2. 
Tome xlix. (1880). 1880. 

Stas. Sur la découverte par le professeur Scacchi, de Naples, d’un 
corps simple nouveau dans la lave du Vésuve, 93.—L. G. de Koninck et 
Cornet. apports sur le mémoire de M. E. Vanden Broeck “Sur les 
phénomenes d’altération des dépots superficiels par V’infiltration des eaux 
météoriques, étudiés dans leurs rapports avec la géologie stratigraphique,” 
615.—P. J. Van,Beneden. Rapport sur le mémoire de M. G. Cotteau: 
“Description des Kchinides tertiaires de Belgique,” 619. 


Brussels. Académie Royale des Sciences. Bulletins. Série 2. 
Tome |. (1880). 1880. 

P. J. Van Beneden. Les Mysticétes & courts fanons des sables des 
environs d’Anvers, 11.—L. G. de Koninck. Rapport sur le mémoire de 
M. A. Renard “Sur la composition chimique de l’épidote de Quenast,” 
80.—A. Renard. Sur la composition chimique de l’épidote de Quenast, 
170.—P. J. Van Beneden. Sur deux Plésiosaures du lias inférieur du 
Luxembourg, 308. 


Mémoires. Tome xliii. 1° Partie. 1880. 


A. Briart et F. L. Cornet. Description des fossiles du calcaire grossier 
de Mons. 38° partie. Supplément aux deux premiéres parties. 


——., ———. Mémoires Couronnés. ‘Tome xxxix, 2° Parnes liso: 


: . . Tome xli. 1879. 
G. Cotteau. Description des Hchinides du calcaire grossier de Mons. 


Tome xlii. 1880. 


G. Cotteau. Description des Hchinides tertiaires de la Belgique. 


: ‘ . (8vo.) Tome xxix. 1880. 
F. V. Rysselberghe. Note sur les oscillations du littoral belge, No. 5. 


=) ome nxxex loos 


ee ee ee Lome maxon aalioole 


Tables des mémoires des membres, des mémoires 
couronnés et de ceux des savants étrangers, 1816-57. 1858. 


AEE Gee Sg SO: 


——. Société Malacologique de Belgique. Annales. Tome xi. 
ke iil 

C. Paul. Considérations nouvelles sur les systémes Bolderien et 
Diestien, 7.—G. Dollfus. Valvata disjuncta, espéce nouvelle des meu- 
liéres supérieures des environs de Paris, 27.—-T. Lefévre et A. Watelet. 
Additions 4 la faune tertiaire du bassin de Paris. Description de deux 
Solens nouveaux, 29.—J. de Cossigny. Tableau des terrains tertiaires 
de la France septentrionale, 37. A. Rutot, G. Dollfus et J. Ortlieb. 
Rapports sur le mémoire précédente, 40.—T. Davidson. Liste des prin- 
cipaux ouvrages, mémoires ou notices qui traitent directement ou indi- 
rectement des Brachiopodes vivants et fossiles, 55.—A. Rutot. Quelques 
observations relatives aux conclusions de M. Lefévre dans son rapport sur 
mon travail intitulé: “ Description de la Rostellaria robusta Rutot,” xi.— 
T. Lefévre. Rapport sur le travail de M. Vincent, intitulé: Description 
de la faune de l’étage landenien inférieur de la Belgique, xxvii.—J. Deby. 
Nécrologie. David Forbes, xxx.—A. Rutot. Rapport sur le travail de 
M. G. Vincent, intitulé: Description de la faune de l’étage landenien 
inférieur. J¢ partie: Massif du Brabant, xxxvi—H. Vanden Broeck et 


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Paul Cogels. Observations sur les couches quaternaires et pliocénes 
de Merxem prés d’Anvers, Ixviiii—T. Lefévre. Sur la “Note sur la 
Rostellaria robusta” par M. Rutot, lxix. 


Brussels. Société Malacologique de Belgique. Procés-verbaux des 
Séances. Tome vill. 1879. 

A. Rutot. Note sur des fossiles du Tongrien inférieur, v.—G. Vincent 
et A. Rutot. Observations nouvelles relatives 4 la faune du systéme 
Bruxellien et a celle de Vancien Laekenien supérieur, actuellement 
systeéme Wemmielien, xii—E. Vanden Broeck et Paul Cogels. Diluvium et 
Campinien, réponse a M. C. Dr. Winkler, xviii—A. Rutot. Sur la plus 
belle coupe de terrains qui ait jamais été visible aux environs de Bruxelles, 
xlvii—k. Vanden Broeck. Compte rendu de l’excursion faite 4 Anvers 
les 27 et 28 Juillet 1879, Ixvii. 


: : Tome ix. 1880. 
P. Cogels et O. Van Ertborn. Nouvelles observations sur les couches 
quaternaires et pliocénes de Merxem, v. 


: : Tome x. pp. i-xxxvi. 1881. 

A. Rutot. Compte rendu d’une course dans le quaternaire de la vallée 
de la Somme, aux environs d’Abbeville, vii—A. Rutot. Note sur les 
découvertes paléontologiques faites dans ces derniers temps aux environs 
d’Erquelennes, xvilii—Stevens. Liste des fossiles recueillis dans les ter- 
rains éocénes des environs de Bruxelles, xx.—T. Lefévre. La Rostellaria 
ampla, Sol. et ses variétés, xxiv. 


Budapest. K. ungarische geologische Anstalt. Jahrbuch. Band iv. 
pe bterg 18s. 
J. Bockh. Geologische und Wasser-Verhaltnisse der Umgebung der 
Stadt Funfkirchen. 


Literarische Berichte aus Ungarn. Band iv. Heft 1-4. 
1880. 

F. Schafarzik. Ueber das Erdbeben in Sud-Ungarn, 642. (For the 
continuation of this journal see “ Leipzig. Ungarische Revue.”) 


——. Magyar Tudomdnyos Akadémia. Ertekezések a Mathema- 
- tikai Tudomdnyok Korébol. Kotet vii. Szdm 3-18. (1879-80). 
1879-80. 


: . Ertekezések a Természettudomanyok Korébol. Kotet 
ix. Szam 20-25. (1879). 1879-80. 

G. Stollar. Az Alsé-kekedi gyogyforras chemiai elemzése, Szim 21, 
. 3.—L. Solymosi. A Fels6-rakosi savanyuviz valamint a székely- 
udvarhelyi hidee sdésfiirdo chemiai elemzése, Sz4m 21, p. 19.—W. A. 
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Szabo. A granat és Cordierit (Dichroit) szereplése a magyarorszagi 
trachytokban, Szam 25. 


: : Kotet x. Szim 1-18 (1880). 1880-81. 

T. Ortvay. A magyarorsziei Dunaszigetek foldirati csoportosulésa s 
képzodésok tényezoi, Szim 3.—J. Bockh. Adatoka Mecsekhegység és 
domboidéke jurakorbeli lerakodasainak ismeretéhez. I. Stratigraphiai 
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: . Mathematikai és Termeszettdoményi Kézlemenyek. 
Kotet xvi. 1881. : 
S. Roth. Szepesmegye nehany barlangjanak leirasa, 618. 


252 ADDITIONS TO THE LIBRARY. 


Buenos Ayres. Sociedad Cientifica Argentina. Anales. Tomo x. 
Entrega 1-6. 1880-81. 
J. J.J. Kyle. La Boronatrocalcita de la Provincia de Salta, 169.— 
J.J. J. Kyle. Observaciones sobre un depésito fosfatico en la Patagonia, 
226. 


——. ——. Tomoxi. Entrega 2-5. 1881. 


Caen. Société Linnéenne de Normandie. Bulletin. Ser. 3. Vol. i. 
(1876-77). 1877. 

G.de Tromelin. Etude de la faune du erés silurien dans le Calvados, 5. 
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dévonien du Cotentin, 96.—J. Pépin. Note sur une nouvelle espéce de 
Sténéosaure, 115.—L. Crié. Considérations sur la flore tertiaire de Fyé 
(Sarthe), 121.—L. Crié. Note sur le Carpolithes Decaisneana, nob., des 
erés éocénes de la Sarthe, 123. 


: —. : - Vols. 877-78) ome 

Moriére. Sur le grés de Bagnoles, 5.—G. de Tromelin. Sur les 
terrains paléozoiques de la Basse-Normandie, 6.—Lodin. Note sur le 
niveau occupé a Honfleur par Vargile de Kimméridge, 15.—J. Moriére. 
Note sur le grés de Bagnoles (Orne), 20.—L. Crié. Note sur les Morinda 
de la flore éocéne du Mans et d’Angers, 46.—J. Moriére. Note sur un 
tronc fossile paraissant se rapporter au genre Cycadeomyelon, Saporta, 51. 
—J. Moriere. Note sur une Astéride fossile nouvelle trouvée dans 
Voxfordien des Vaches-noires entre Dives et Villers-sur-Mer, 75.— 
Letellier. Deuxiéme excursion de la Société Linnéenne a Alengon, 270. 
—Lodin. Du mode de formation et de la classification des terrains supé- 
rieurs, 315.—Drouaux. La géologie et l’Exposition géologique au con- 
erés du Havre, 525.—Letellier. Note géologique sur les terrains traversés 
parle chemin de fer d’Alencgon & Domfront, 340.—Lionnet. Phosphates 
de chaux fossiles, géologie et origine, application en agriculture, 351. 


Calcutta. Asiatic Society of Bengal. Journal. N.S. Vol. xlvii. 
Extra number to Part 1 for 1878. 1880. 


: . -——. Vol. xlix. Part 1. Nos. 1—4, and 
extra number. 1880-81. 


: . Vol. xlx. Part 2. Nos. 1-4. 1880-81. 
R. Lydekker. A Sketch of the History of the fossil Vertebrata of 
India, 8. 


ae 


——, ——. ——., Vol.l. Partl. No. fy 133an 
~ Volol. Part: 23, Noxieeame Sis 


V. Ball. On the Identification of certain Diamond-mines in India, 
which were known to and workefl by the Ancients, especially those which 
were visited by Tavernier: with a note on the History of the Koh-i-nur, 
31.—List of Harthquakes recorded in Assam during the years 1879 and 
1880, 61. 


. ——. Proceedings. 1880, Nos. 2-10. 1880. 
Rey. C. Swynnerton. On a Celt of the Paleolithic Type found at 
Thandiani, Punjab, 175. 


: 1881, Nos. 1-3. 1881. 

V. Ball. On the Identification of certain Diamond-mines in India 
which were known to and worked by the Ancients, especially those which 
were visited by Tavernier, 2. 


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Cambridge Philosophical Society. Proceedings. Vol. ui. Parts 
ace &-" 18380; 

O. Fisher. On the Implement-bearing Loams in Suffolk, 285.— 
T. McK. Hughes. On the Transport of fine Mud and Vegetable Matter 
by Confervee, 339.—T. M‘K. Hughes. On the Altered Rocks of Anglesea, 
341.—W. Keeping. On the included Pebbles of the Neocomian Deposits 
of Potton and Upware, and their bearing upon the Physical Features of 
the Lower Cretaceous Period, 377. 


——. ——. Vol.iv. Part l. 1881. 

EH. Tawney and H. Keeping. On the Beds at Headon Hill and Colwell 
Bay in the Isle of Wight, 59. 
Transactions. Vol. xiii. Part 1. 1881. 


Cambridge, U.S.A. Museum of Comparative Zoology at Harvard 
College. Annual Report for 1879-80. 1880. 


bulketme se Volsvie. Noss oli asso: 


——, ——. ——. Vol.vu. No.1. (Geological Series, Vol. i. 
Noss) 9 1830- 
M. E. Wadsworth. Notes on the Geology of the Iron and Copper 
Districts of Lake Superior, 1. 


: : Vol. viii. Nos. 1-4. 1880. 

C. D. Walcott. The Trilobite: new and old Evidence relating to its 
Organization, 192.—H. A. Hagen. The Devonian Insects of New Bruns- 
wick, 275. 


—_—— 


: Memoirs. Vol. vi. No.1. 1880. 
J.D. Whitney. The Auriferous Gravels of the Sierra Nevada of 
California, 289. 


: : Wok vm,  IN@s 1, ihesl0- 
L. Agassiz. Report on the Florida Reefs, 1. 


: : ae olen Now. sRartols, 1880. 
J.D. Whitney. The Climatic Changes of later Geological Times, 1. 


Canterbury. East-Kent Natural-History Society. 12th Report, 
1870. Presented by W. Whitaker, Esq., F.GS. 


——. ——. 20th Report, 1878. Presented by W. Whitaker, 
Esq., F.G.S. 

G. Dowker. On Flint Stones, with an account of Banded Flints, 20.— 
McDakin. Structure of the Sea-cliffs at Folkestone, 29—M‘Dakin. On 
some Fossil Wood from EHastwear Bay near Folkestone, 55.—J. Reid. On 
the various Iron-stone Bands in the Strata of Hast Kent, 40.—M*‘Dakin. 
On the Tertiaries of the London and Hampshire Basins, 44. 


Cardiff Naturalists’ Society. Report and Transactions. Vol. xi. 
MOO SoU. 
W. J. Sollas. On the Silurian District of Rhymney and Pen-y-lan, 
Cardiff, 7—T. Wright. On the Geological Features of the Landscape 
seen from Symond’s Yat, 70. 


Cassel. Palsontographica. Band xxvii. Lief. 1-6. 1880-81. 
Purchased. 


F, Romer. Ueber eine Kohlenkalk-Fauna der Westkiiste von Sumatra, 1. 
—W. Branco. Beitrige zur Entwickelungsgeschichte der fossilen Cepha- 


254 ADDITIONS TO THE LIBRARY. 


lopoden, 12.—F’. Sandberger. Ein Beitrag zur Kenntniss der interpleisto- 
canen Schichten Englands, 82.—Hans Pohlig. Maritime Unionen, 105.— 
M. Neumayr und V. Uhlig. Ueber Ammonitiden aus den Hilsbildungen 
Norddeutschlands, 129. 


Cassel. Paleeontographica. Supplement iii. Lieferung 8 und 9. 
1880. Purchased. | 
R. D. M. Verbeek, O. Boettger und K. von Fritsch. Die Tertiar- 
formation von Sumatra und ihre Thierreste, 1. 


Chemical News. Vol. xli. Nos. 1074. 1880. 


. Vol. xlii. Nos. 1075-1101. 1880. 

J.A.Church. The Heat of the Comstock Lode, 42,52.—A. C. Ramsay. 
Presidential Address to the British Association, 99.—F. A. Massie. On 
the Composition of Dufrenite from Rockbridge Co., Va., 181.—F. A. 
Massie. Analysis of Amphibole (var. Actinolite) from Amelia Co., Va., 
194.—W. T. Page. Analysis of a Highly Aluminous Pyroxene from 
Amhurst Co., Va., 194.—A. L. Baker. Analysis of Ferruginous Mineral, 
probably referable to Amphibole, from Amhurst Co., Va., 194.—W. T. 
Page. Examination of Livingstonite from a New Mexican Locality, 195. 
—W.T. Page. Examination of a altered Livingstonite from Guadalcazar, 
S. Luis Potosi, Mexico, 195.—A. Liversidge. Water from a Hot Spring, 
New Britain, 324.—A. Liversidge. Water from a Hot Spring, Fiji 
Islands, 324. 


——. Vol. xl. Nos. 1102-1125. 1881. 


H. Leffman. Analyses of some Geyser Deposits, 124.—R. Romanis. 
On the Hot Spring at Natmoo near Maulmain, British Burmah, 191. 


Chemical Society. Journal. No. 200. 1879. 
G. Attwood. Ona Gold Nugget from South America, 427. 


: . Nos. 212-217. 1880. 
H. Baker. On a Crystal of Diamond, 579. 


: . Nos. 218-223. 1881. | 
R. H. Davis. The ancient Allum Well at Harrogate, 19.—A. Liver- 
sidge. Analyses of Queensland Soils, 61. 


es 


—— 


——. Indexes to vols. xxxvii. & xxvii. 1880. 


Cincinnati Society of Natural History. Journal. Vol. in. Nos. 2 
& 3. 1830. 

S. A. Miller. North-American Mesozoic and Czenozoic Geology and 
Paleontology, 79.—S. A. Miller. Description of four new species of 
Silurian Fossils, 140.—A. G. Wetherby. Remarks on the Trenton Lime- 
stone of Kentucky, with descriptions of new fossils from that formation 
and the Kaskaskia (Chester) Group, Sub-Carboniferous, 145.—S. A. 
Miller. North-American Mesozoic and Ceenozoic Geology and Paleeon- 
tology, 165.—S. A. Miller. Descriptions of four new species and a new 
variety of Silurian Fossils, and remarks upon others, 282. 


Clausthal. Berg- und hiittenminnische Vereins Maja. (Late Natur- 
wissenschaftlicher Vereins.) Neue Folge. Heft 2. 1880. 
Purchased. 

F. Wunderlich. Beitrag zur Kenntniss der Kielschiefer, Adinolen und 

Wetzschiefer des nordwestlichen Oberharzes, 1.—H. L. Bridgman. Das 

Erzvorkommen im Leadville District, 119. 


ADDITIONS TO THE LIBRARY. 255 


Copenhagen. Kongelige Danske Videnskabernes Selskab. Natur- 
videnskabelige og Mathematiske Afhandlinger. Reekke 5. 
Bind xu. No. 6. 1880. 


—. ——. ——. Rekke 6. Bindi. Nos. 1&2. 1881. 
——. ——. Oversigt. 1880, Nos.2&3. 1889. 


1881, No.1. 1881.. 


Colliery Guardian. Vol. xl. Nos. 1018-1044. 1880, 

Hxcursion of the Edinburgh Geological Society to Linlithgow and 
Borrowstounness, 95.—The “Lamb-bottom Caves at East Harptree, 
Somerset, 186.—British-Association Meeting at Swansea, 346, 375.— 
The Coal Industry of the Lower Rhine and of Westphalia, 426,—W. 
Saise. The Kurhurballee Coalfields, with some remarks on Indian Coals, 
454.—Coal in Cape Colony, 461.—Coal deposits in Natal, 664, 856.— 
H. Woodward. The Temperatnre of strata in Coal mines, 903.—W. 
Bailes. On Mineral Veins, 1065. 


Vol. xli. Nos. 1045-1068. 1881. 

John Brown. A description of some borings on the Clifton Estate, and 
of the sinking at Clifton Colliery, Nottimgham, 93.—The deepest Coal-pit 
in Britain, 387.—The St. Etienne Coalfield, 415.—Coal in Venezueia, 
493, 785. 


Cracow. Akademija umiejetnosci w Krakowie. Sprawozdanie, 
1879. 1880. 
Jana Trejdosiewicza. O porfirze w Krélestwie Polsliém, 258. 


Darmstadt. Vereins fiir Erdkunde. Notizblatt. 1880. Folge 4. 
Heft 1. 1880. 


Delémont. Société Jurassienne d’Emulation. L’Emulation Juras- 
sienne. 1877, Jan.—Juillet. 1877. Presented by H. B. Wood- 
ward, Ksq., F.GS. : 


Dorpat. Naturforscher-Gesellschaft. Archiv fiir die Naturkunde. 
Ser. 2. Biologische Naturkunde. Band ix. Lief. 1 u. 2. 
1880. 


— EE 
° ° 


Sitzungsberichte. Band vy. Heft 3, 1880. 1881. 

C. Grewinek. Uebersicht der altquartaren und auspestorbenen neu- 
quartaren Saug ethiere Liv-, Est- und Kurlands, 339.—C. Grewingk. 
Ueber eylindrische Strudel- und Sickergruben in deyonischen Gypslager 
bei Diinhof, oberhalb Riga, 359.—M. Ostwald. Ueber Moorseen, 455.— 
C. Grewingk. Ueber zwei Geschiebehugel der Westkuste Hstland’s und 
deren Entstehungsweise, 439. 


Dresden. Naturwissenschaftlichte Gesellschaft Isis. Sitzungs- 
berichte. (1880.) 1881. | 

Deichmiller. Ueber eine Abhandlung von Walter Keeping “On some 
Columnar Sandstone in Saxon Switzerland,” 1—A. Jentzsch. Briefliche 
Mittheilungen tiber seme Sammlung sachsischer Diluvialvorkommnisse, 4. 
—H.Engelhardt. Ueber den tertiiren Stisswassersandstein von Grasseth, 5. 
—E. Zschau. Ueber Mineralvorkommnisse im Granit, 6.—H. B. Geinitz. 
Ueber die Fortschritte der geologischen Forschungen in Nordamerika, 59. 
—H. Engelhardt. Ueber Pflanzenreste aus den ‘Tertidrablag erungen von 
Liehotitz und Putschirn, 77. 


‘ 


256 ADDITIONS TO THE LIBRARY. 


Dublin. Royal Geological Society of Ireland. Journal. N.S. Vol. v. 
Part 3. 1880. 

T. Mellard Reade. A Problem for Irish Geologists in Post-Glacial 
Geology, 173.—S. Haughton. Notes on the Annual Water-discharge of 
large Rivers, with Indications of some New Methods of Calculation, 177. 
—J. P. O'Reilly. On the Occurrence of Microline Feldspar in Dalkey 
Granites, 189.—V. Ball. On Spheroidal Jointing in Metamorphic Rocks 
in India and elsewhere, producing a Structure resembling Glacial “ Roches 
Moutonnées,” 193.—S. Haughton. On Rossetti’s Law of Cooling, applied 
to the consideration of the relative effects of Sun-heat, Harth-heat, Star- 
heat, and Atmospheric Conditions upon Climates during Geological Time, 
199.—G. H. Kinahan. Anniversary Address, 212.—V. Ball. On the 
Evidence in favour of the belief in the existence of Floating Ice in India 
during the deposition of the Talchir (Permian or Permio-Triassic) Rocks, 
223.—V. Ball. On the Coalfields and Coal Production of India, 230.— 
V. Ball. On the Mode of Occurrence and Distribution of Gold in India, 
258. 

——. Royal Irish Academy. . “Cunningham Memoirs.” No. 1. 
1880. 

——. ——. Transactions. Vol. xxiv. Antiquities. Part 1. 
1864. 

——. : . Vol. xxvi. Science. Part 22. 1879. 

J.P. O'Reilly. Explanatory Notes and Discussion of the Nature of the 

Prismatic Forms of a Group of Columnar Basalts, Giant’s Causeway, 641. 


: Vol. xxvii. Polite Literature and Antiqui- 
ties. Part 3. 1879. 
——, Irish Manuscript Series. Vol. 1. Part i 


1880. 
— . Proceedings. Ser. 2. Vol. i. No. 1, 1869-70. 


1870. 

W. K. Sullivan. On the Formation of Thenardite in connexion with 
the date of the Glacial Period, and the Temperature that prevailed during 
it, as deduced from the Influence of the Eccentricity of the Karth’s Orbit 
on the Length of Summer and Winter in Aphelion and Perihelion, 2.— 
W. K. Sullivan. Note on the Hornblende and Augite Groups of Mine- 


rals, 37. 


Ser, 2. Vol. i. No. 11, 1874-75. 1875. 
pa yf Ce eT Ser) D2) Viol Si NiO egos 
ee aan ye eS Oe iolman mmiNoniss )  LSGO, 


Eastbourne Natural-History Society. Papers, 1879-80. Presented 
by W. Whitaker, Esq., F.G.S. | 
A. Ramsay. On the Local, Stratigraphical, and Paleontological Evi- 
dence as to the conditions of the Cretaceous Ocean. . 


Fast-India Association. Journal. Vol. xii. Nos.1&2. 1880-81. 
Easton. American Institute of Mining Engineers. ‘Transactions, 
Vol. i. 1871-73. (Philadelphia.) Presented by W. Whitaker, 


Esq., F.G.S. 
J. F. Blandy. Topography, with especial reference to the Lake Su- 
perior Copper District, 75.—J. W. Harden. The Brown Hematite-ore 


oe oo 
° e 


ADDITIONS TO THE LIBRARY. 257 


deposits of South Mountain, between Carlisle, Waynesborough, and the 
South-eastern Edge of Cumberland Valley, 136.—B. 8. Lyman. On the 
Importance of Surveying in Geology, 183.—R. N. Clark. The Tertiary 
Coal-beds of Canyon City, Colorado, 293.—T. 8. Hunt. The Geoenos- 
tical History of the Metals, 331.—O. J. Heinrich. The Midlothian 
Colliery, Virginia, 346.—B. Silliman. On the probable Existence of 
microscopic Diamonds, with Zircons and Topaz, in the Sands of Hydraulic 
Washings in California, 371.—T. 8. Hunt. Remarks on an Occurrence of 
Tin-ore at Winslow, Maine, 372.—T.S. Hunt. The Origin of Metalli- 
ferous deposits, 415. 


Easton. American Institute of Mining Engineers. Transactions. 
Vol. ii. 1873-74. Presented by W. Whitaker, Esq., F.G.S. 

T. S. Hunt. The Geology of the North Shore of Lake Superior, 58.— 
R. W. Raymond. Remarks on the Occurrence of Anthracite in New 
Mexico, 140.—R. W. Raymond. Remarks on the Occurrence of South- 
African Diamonds, 143.—R. P. Rothwell. Alabama Coal and Iron, 145. 
—T. S. Hunt. The Coals of the Hocking Valley, Ohio, 273.—J. C. 
Smock. The Magnetic Iron-ores of New Jersey, their Geographical Dis- 
tribution and Geological Occurrence, 514. 


: Vol. vi. (1877-78). 1879. Presented by W. 
Whitaker, Esq., F.G.S. 

A. J. Bowie, jr. Hydraulic Mining in California, 27.—O. J. Heinrich. 
The Manhattan Salt Mine at Goderich, Canada, 125.—-C. M. Rolker. The 
late operations on the Mariposa Hstate, 145.—J. C. Smock. The Fire 
Clays and Associated Plastic Clays, Kaolins, Feldspars, and Fire Sands 
of New Jersey, 177.—W.H. C. Hustis. The Nickel Ores of Orford, 
Quebec, Canada, 209.—A. L. Holley. Notes on the Iron Ore and An- 
thracite Coal of Rhode Island and Massachusetts, 224.—O. J. Heinrich. 
The Mesozoic Formation in Virginia, 227.—T. Kgleston. Copper mining 
on Lake Superior, 275.—W. 5S. Keyes. The Eureka Lode of Kureka, 
Eastern Nevada, 344.—R. W. Raymond. What is a Pipe Vein?, 393,— 
J. P. Carson. Iron Manufacture in Mexico, 598.—P. Frazer, jr. Clas- 
sification of Coals, 480.—G. H. Cook. On the Southern Limit of the last 
Glacial Drift across New Jersey and the adjacent parts of New York and 
Pennsylvania, 467.—E. P. Jennings. Analyses of some Tellurium Mine- 
rals, 506.—P. Frazer, jr. Missing Ores of Iron, 531.—W. P. Blake. The 
Ore-deposits of Hureka District, Hastern Nevada, 554. 


Edinburgh. Geological Society. Transactions. Vol. ii. Part 3. 
1880. 

A. Taylor. On the Crag Structure on the South Slopes of Queen’s 
Park, 279.—W. 1. Macadam. On the Chemical Composition of Rocks 
from Salisbury Crag (south side), 288.—W. M‘Diarmid. On the North- 
east coast of Norfolk, 292.—T. Stock. On a Section recently exposed 
near Straiton, 294.—J. Young. On Scottish Carboniferous Microzoa, 
999.—H. M. Cadell. On the Volcanic Rocks of the Borrowstounness 
Coalfield, 304.—J. Melvin. On the Evidence the Vegetable Soil affords 
as to Geological Time, 326.—T. D. Wallace. On the Geology of Rathven 
and Enzie, 331.—D’A. W. Thompson. On the Ulodendron and Halonia, 
341.—J. Henderson. On some recently discovered Fossiliferous Beds in 
the Pentlands, 353.—D. Milne Home. Valedictory Address, 357. 


——. Royal Society. Proceedings. Vol. x. 1879-80. 
A. Geikie. On the Geology of the Rocky Mountains, 426.—J. Geikie. 
The Geology of the Faroe Islands, 495.—J. Murry. On the structure and 
VOL. XXXVII. U 


258 ADDITIONS TO THE LIBRARY. 


origin of Coral Reefs and Islands, 505.—A. Geikie. Rock-weathering, as 
illustrated in Edinburgh Churchyards, 518.—Sixth Report of the Boulder 
Committee, 577.—R. H. Traquair. Report on Fossil Fishes collected by 
the Geological Survey of Scotland in Roxburgshire and Dumfriesshire. 
Part I. Ganoidei, 710.—B. N. Peach. On some new Crustacea from the 
Cementstone Group of the Calciferous Sandstone Series of Eskdale and 
Liddesdale, 711. 


Edinburgh. Royal Society. Transactions. Vol. xxix. Part 2. 
1880. 


Epping Forest and County of Essex Naturalists’ Field Club. 
Transactions. Vol.i. Parts 1-3. 1880-81. 
H. Walker. A day’s Elephant Hunting in Essex, 27. 


Falmouth. Royal Cornwall Polytechnic Society. Forty-eighth 
Annual Report. 1880. 


Geneva. Société de Physique et d’Histoire Naturelle. Mémoires. 
Tome xxvii. Partie 1. 1880. 
P. de Loriol. Monographie des Echinides contenus dans les couches 
nummulitiques de ’ Egypte, 59. 


Geological and Natural-History Repertory and Journal of Prehistoric 
Archeology and Ethnology. Edited by S.J. Mackie. 1865-67. 
Purchased. 


Geological Magazine. Dec. II. Vol. vii. Nos. 7-12. 1880. 

W.H. Hudleston. Contributions to the Paleontology of the Yorkshire 
Oolites, 289, 391, 481, 529.—T. G. Bonney. Classification of Welsh Pre- 
Cambrian Rocks, 298.—R. Etheridge, jun. Contributions to British Tubi- 
colar Annelides, 304.—E. Wilson. On the Unconformability of the Keuper 
and Bunter, 308.—W. J. M‘Gee. The “ Laterite” of the Indian Penin- 
sula, 310.—A. B. Wynne. The Paleozoic Rocks of the Northern Punjab, 
313.—T. Davidson. On the Brachiopoda that characterize the Beds of 
Brittany and South Devon, 837.—F. Romer. Note on the Genus Cau- 
nopora of Phillips, 345.—W. Davies. On the Bones of the Lynx from 
Teesdale obtained by Mr. James Backhouse, of York, 546.—J. J. Harris 
Teall. Influence of Earth Movements on the Structure of the British 
Islands, 349.—J. Croll. Aqueous Vapour in Relation to perpetual Snow, 
357.—A.Champernowne. Upper Devonian in Devonshire, 359.—R. Ethe- 
ridge, jun. Contributions to a Study of Carboniferous Tubicolar Annelides, 
362.—J. G. H. Godfrey. Association of Stibnite and Cinnabar in Mineral 
Deposits, 369.—T. M. Reade. Oceans and Continents, 585.—T. G. Bonney. 
Pebbles in the Bunter Beds of Staffordshire, 404.—H. H. Howorth. The 
Mammoth in Siberia, 408, 491, 550.—W. Keeping. On the Pebbles of 
the Upper Neocomian of the South-east of England, 414.—T. Rupert 
Jones. Note on a Well lately sunk at Wokingham, 421.—E. T. Newton. 
Notes on the Vertebrata of the Pre-Glacial [orest-bed of the East of 
England, 424.—C. T. Clough. The “Whin Sill” of Teesdale, as an 
Assimilator of the surrounding Beds, 433.—E. T. Newton. Notes on the 
Vertebrata of the Pre-Glacial Forest-bed Series of the Kast of Hngland, 
447,—E. B. Tawney. Woodwardian Laboratory Notes: North-Wales 
Rocks, 452.—Thoroddsen, Thorvald. Volcanic Eruptions and Harth- 
quakes in Iceland within Historic Times, 458.—H. Hicks. Pre-Cambrian 
Volcanos and Glaciers, 488.—G. R. Vine. On the Carboniferous Polyzoa, 
501.—R. J. Ussher. On the Caves and Kitchen-Middens at Carrigagower, 


ADDITIONS TO THE LIBRARY. 259 


Co. Cork, 512.—T. G. Bonney. On some Serpentines from the Rheetian 
Alps, 538.—R. B. Foote. Notes on the Occurrence of Stone Implements 
in the Coast Laterite south of Madras, 542.—A. Liversidge. Analyses of 
Moa Eege-shell, 546.—C. Reid. Classification of the Pliocene and Pleis- 
tocene Beds, 548. 


Geological Magazine. N.S. Dec. IL. Vol. viii. Nos. 1-6. 1881. 

T. Dayidson. On Spiral-bearing Brachiopoda from the Wenlock and 
Ludlow Shales of Shropshire, 1—H.G. Seeley. Ontwo Ornithosaurians 
from the Upper Greensand of Cambridge, 18.—O. Fisher. Oblique and 
Orthogonal Sections of a folded Plane, 20.—F. D. Longe. On the Oolitic 
Polyzoa, 23.—R. H. Traquair. On New Fish-remains from the Blackhand 
Ironstone of Borough-Lee, near Edinburgh, 34.—W. H. Hudleston. Contri- 
butions to the Paleeontology of the Yorkshire Oolites, 49, 110.—J. D. Dana. 
Metamorphism of Massive Crystalline Rocks, 59, 110, 162.—B. N. Peach 
and J. Horne. The Glaciation of the Shetland Isles, 65.—J. Smith and 
T. Rupert Jones. Upper Silurian Entomostraca, 70.—T. Mellard Reade. 
Oceanic Islands, 75.—T. Stock. On anearly entire Rhizodus at Wardie, 
77.—W. J. Sollas. On Striated Triassic Pebbles, 79.—A. B. Wynne. 
On the Travelled Blocks of the Upper Punjab and a supposed Glacial 
Period in India, 97.—T. Davidson and George Maw. On the U. Silu- 
rian Rocks of Shropshire, with their Brachiopoda, 100.—W. A. E. 
Ussher. Pre-historic Europe and the Cornish Forest-beds, 151.—G. H. 
Kinahan. ‘ Laccolites, 134.—T. Davidson. New Upper Silurian Bra- 
chiopoda from Shropshire, 145—G. M. Dawson. Geology of British 
Columbia, 156, 214.—W. H. Herries. Woodwardian Laboratory Notes: 
Part 5. The Bagshot Beds, 171.—G. W. Lamplugh. Ona Shell-bed under 
the Drift at Speeton near Filey, Yorkshire, 174.—T. Rupert Jones. On the 
Carboniferous System in Britain, 181.—H. Woodward. Note on a Head- 
shield of Zenaspis Salwey?, 193.—R. D. Roberts. Position of the Twt-Hill 
Conglomerate, 194.—T. Mellard Reade. AXjolian Sandstone, 197.—H. H. 
Howorth. The Mammoth in Europe, 198, 251.—D. Milne Home. Glaci- 
ation of the Shetlands, 205.—W. G. Lock. _ Volcanic History of Iceland, 
212.—J. S. Gardner. Subsidence and Elevation, 241.—J. H. Marr. 
Classification of the Cambrian and Silurian Rocks, 245.—K. 'T. Newton. 
The Vertebrata of the Forest-bed Series of the East of England, 256.— 
C. Lapworth. Correlation of the Lower Paleozoic Rocks of Britain and 
Scandinavia, 260. 


eV Olivia NOS (oe LOoOn weeunchascd. 
VO) Vili NOSa lO.) el GOlionn eeeneRased. 


Geologists’ Association. Proceedings. Vol. vi. Nos. 7-9. 1880 
and 1881. 

Visit to the Museum of Practical Geology, Jermyn Street: Demon- 
stration on the Minerals in the Horse-shoe Case; reported by W. H. 
Hudleston, 311.—J. S. Gardner. Excursion to the Hampshire Coast, 
316.—R. Owen. Visit to the British Museum: Demonstration on the 
Elephantine Mammals, 321.—T. Rupert Jones, A. Irving, and C. Cooper 
King. Excursion to Camberley, 529.—W. Whitaker. Excursion to 
Upnor, 336.—W. H. Hudleston. Excursion to Oxford.—W. H. Hudle- 
ston. Excursion to Aylesbury, 344.—A. Taylor. On the Origin and 
Petrology (specially of the South side) of Salisbury Crags, Edinburgh, 
303.—J. W. Davis. On the Fish Fauna of the Yorkshire Coal-tield, 359. 
—Excursion to Kew Gardens, Richmond, and Kineston Hill, 370.—Ex- 
eursion to Croydon and Riddlesdown, 372.—Excursion to Redhill, 373. 

u 2 


260 ADDITIONS TO THE LIBRARY. 


W. J. Sollas. On the Geology of the Bristol District, 375.—Excursion 
to Maidstone, 392.—Excursion to Leith Hill and Dorking, 393.—Excur- 
sion to Bristol, 396.—G. 8. Boulger. On the Geological and other 
Causes that Affect the Distribution of the British Flora, 403.—J. F. 
Blake. On the Classification of Rocks, 418.—EH. Renevier. A Petro- 
graphical Classification of Rocks, 426.—T. Rupert Jones. Onthe Geology 
and Physical Features of the Bagshot District, 429. 


Geologists’ Association. Proceedings. Vol. vii. No.1. 1881. 
_ T. Rupert Jones. Presidential Address, 1—H. Hicks. On some 
Recent Researches among Pre-Cambrian Rocks in the British Isles, 59. 


Giessen. Oberhessische Gesellschaft fiir Natur- und MHeilkunde. 
Neunzehnter Bericht. 1880. 
A. Nies. Vorlaufiger Bericht tiber zwei neue Mineralien von Grube 


Eleonore am Diinsbere bei Giessen, 111.—A. Streng. Ueber die Phosphate 
von Waldgirmes, 151. 


Gloucester. Cotteswold Naturalists’ Field Club. Proceedings. 
1879-80. i880. 

Sir W. V. Guise, Bart. Presidential Address, 1879, 94.—K. Witchell. 
On a Section of Stroud Hill and the Upper Ragstone Beds of the Cottes- 
wolds, 118.—Sir W. V. Guise, Bart. Presidential Address, 1880, 157.— 
T. Wright. On the Modern Classification of the Ammonitide, 170. 


Haarlem. Société Hollandaise des Sciences. Archives Néerlandaises 
des Sciences Exactes et Naturelles. Tome xv. Livr. 3-5. 
1880. 


Halifax, N.S. Nova-Scotian Institute of Natural Science. Pro- 
ceedings and Transactions. Vol. v. Part 2. 1880. 

D. Honeyman. Nova-Scotian Geology, Annapolis County, 119.—D. 
Honeyman. Geological Waifs from Mag dalen Islands, 1836,—D. Honey- 
man. Nova-Scotian Geology : Notes on a New Geological Progress Map 
of Pictou County, 192. 


Havre. Socicté Géologique de Normandie. Bulletin. Tome v. 
1878. (1880 ?) 

G. Lionnet. Réunion de la Société Linnéenne de Normandie a Alen- 
gon les 15 et 16 Juin 1878 (premiere journée), 40.—G. Drouaux. Réunion 
de la Société Linnéenne de Normandie a Alencon les 15 et 16 juin 1878 
(deuxiéme journée), 53.—G. Drouaux. La Géologie et ’ Exposition gé0- 
logique au Congrés du Havre, 74.—H. Savalle. Note sur les sables néo- 
comiens de la Héve et sur quelques espéces f fossiles qui y ont été recueil- 
lies, 94.—E Savalle. Note sur une espéce de crustacé fossile trouvé dans 
les sables micacés de la Héve, 98.—H. Savalle. Note sur les sables mi- 
cacés de la Héve, 100.—F. Prudhomme. Rapport sur le Tome V. des 
Annales de la Société Géologique du Nord, 106.—C. Brylinski. Note 
sur l’Hozoon canadense, 114.—M. Gourdon. Note sur quelques blocs 
erratiques de la Vallée de Valencia de Arreu, Catalogne, 119.—(G. Lion- 
net’). Notes sur quelques substances minérales servant aux Arts ou a 
VIndustrie, 120.—G. Biochet. Note sur le dolmen troué d’Aizier, Kure, 
137. 


; Tome vi. 1879. 1880. (Exposition Géo- 
logique et Paléontologique du Havre en 1877.) 

Description de l’ixposition, 1—G. Lennier et G. Lionnet. Résumé 

sur la Géologie Normande, 29.—Letellier. Les terrains des environs 


ADDITIONS TO THE LIBRARY. 261 


d’Alencon, 525.—W. A. E. Ussher. Etude sur les roches triasiques de 
Normandie et sur les dépdts limitrophes, 531.—A. Durand. L’étage co- 
rallien sur le littoral du Calvados, 563.—E. Savalle. Note sur les sables 
micacés de l’étage néocomien, ala Héve, et sur quelques espéces fossiles 
découvertes récemment a ce niveau, 577.—G. Dollfus. Essai sur l’ex- 
tension des terrains tertiaires dans le bassin anglo-parisien, 584.—G. 
Cotteau. Catalogue des Kchinides jurassiques de Normandie, 606.—G. 
de Saporta. Notice sur les végétaux fossiles de la craie inférieure des 
environs du Havre, 640.—G. Drouaux. Combustibles minéraux en Nor- 
mandie, 662.—C. Marchand. De la composition chimique des calcaires 
exposés par la Société Géologique de Normandie, 692.—Besnou. Consi- 
dérations sommaires sur quelques produits sousmarins, et analyses de ces 
produits, 704.—E. Marchand. Considérations générales sur la constitu- 
tion des eaux potables et en particulier des eaux du terrain crétacé dans 
les arrondissements du Havre et d’Yvetot, 742.—H. Meurdra. Etude 
sur le regime des sources du Havre, 777.—E. T. Hamy. Notes d’an- 
thropologie paléontologique, prises a l’Exposition du Havre, 791.—D. 
Bourdet. L’archéologie préhistorique 4 l’Exposition du Havre, 807.—C. 
Beaugrand. Note sur quelques silex taillés des environs d’Yvetot et du 
Havre, 846. 


Heidelberg. Naturhistorisch-medicinischer Verein. Verhandlungen. 
Neue Folge. Bandii. Heft 5. 1880. 
A. Schmidt. Die Zinkerz-Lagerstitten von Wiesloch, Baden, 369. 


Institution of Civil Engineers. Minutes of Proceedings. Vol. lx. 
1880. 


: Nola eS ssv: 

J. Lucas. The Hydrogeology of the Lower Greensands of Surrey and 
Hampshire, 200.—G. Attwood. The Chile Vein Gold Works, South 
America, 244, 


Vol. lxu. 1880. 
——. ——. Vol. lxii. 188]. 


Kansas. Kansas City Review of Science and Industry. Vol. i. 
No. 10. 1879. Presented by W. Whitaker, Esq., F.G.S. 


: Vol. 1. No. 6. 1879. 

B. F. Mudge. Botany and Evolution, 321.—E. P. West. Following 
the Pick and the Spade, 328.—W. H. R. Lykins. The Stone Age in 
Kansas, 331. 


: Vol. 11. No. 9. 1880. 
J. A. Smith. Some Objections to Mcdern Geological Teachings, 740. 
—The Attractions of the Yellowstone National Park, 745. 


: oP Voleiv= No: I 7kS80: 
T. M‘K. Hughes. Antiquity of Man Questioned, 19. 


Lausanne. Société Vaudoise des Sciences Naturelles. Bulletin. 
Sér. 2. Tome xvii. No. 84 1880. 

E. Schmidt. Analyse de Veau de Vernex-dessus, 15.—G. Maillard. 
Nouveau gisement de feuilles fossiles, 32.—P. de la Harpe. Note sur les 
Nummulites Partschi and Oosteri, de la H., du calcaire du Michelsberg, 
pres Stockerau (Autriche) et du Gurnigelsandstein de Suisse, 33.—G. 
Maillard. Notice sur la molasse dans la ravine de la Pandéze au Moulin 
de Belmont, 81.—W. Fraisse. Note sur les érosions de Vallamand, 157. 


262 ADDITIONS TO THE LIBRARY. 


Leeds. Philosophical and Literary Society. The Annual Report 
for 1879-80. 1880. 3 


——. Yorkshire Geological and Polytechnic Society. Proceedings. 
Vol. vu. Part 3, 1880. 1881. 

J. W. Davis. On the Distribution of Fossil Fish in the Yorkshire 
Coal-fields, 228.—G. W. Lamplugh. Ona Faultin the Chalk of Flambro’ 
Head, with some Notes on the Drift of the Locality, 242._J. R. Dakyns. 
On Glacial Deposits North of Bridlington, 246.—J. Magens Mello. Ona 
Short History of the Creswell Caves, 252.—J. W. Davis. On a Group 
of Erratic Boulders at Norber, near Clapham, in Yorkshire, 266.—W. H. 
Wood. Ona Chemical Method of Distinguishing Black Obsidian from 
Black Blast Furnace Slag, 274.—W. Y. Veitch. On the Geology of the 
District around Middlesborough, 284.—T. Allison. Notes on the Geology 
of the Cleveland District, 285.—L. C. Miall. On some Bones of Ctenodus, 
289.—J. W. Davis. Report of the Raygill Fissure Exploration Commit- 
tee, 800.—J. W. Davis. Summary of Geological Literature relating to 
Yorkshire, published during 1878, 1879, and 1880, 319. 


Leipzig. Naturforschende Gesellschaft. Sitzungsberichte. Jahr- 
gane vi. 1879. 1880. 


: : . Jahrgang vi. 1880. 1881. 
H. Credner. Ueber die geologischen Resultate einer Tiefbohrung am 
Berliner Bahnhofe zu Leipzig, 1. 


——, Ungarische Revue. 1881. Hefte 1-3. 1881. 


——. AZeitschrift fiir Krystallographie und Mineralogie. (Groth.) 
Band iv. Hefte5&6. 1880. Purchased. 


H. Laspeyres. Mineralogische Bemerkungen,433.—L. Calderon. Ueber 
die optischen Eigenschaften der Zinkblende von Santander, 504. 


(——.) Band v. Hefte 1-6. 1880-81. Pur- 


chased. 

G. vom Rath. Mineralogische Mittheilungen, 1—A. Damour und G. 
vom Rath. Ueber den Kentrolith, eine neue Mineralspecies, 32.—A. 
Bertin. Ueber eie Farben von Krystallplatten im elliptisch polarisirten 
Lichte, 36.—V. von Zepharovich. Mineralogische Notizen, 96.—A. Brun. 
Mineralogische Notizen, 103.—W. Voigt. Ueber den Einfluss einer 
Kriummung der Prismenflachen auf die Messungen von Brechungsindices 
und tiber die Beobachtungen des Herrn Calderon an der Zinkblende, 113. 
—W.Websky. Ueber die Berechnung der Elemente einer monoklinischen 
Krystallgattung, 168,—G. J. Brush und E. 8. Dana. Ueber Krystalli- 
sirten Danburit von Russell, St. Lawrence County, New York, 183.—G. 
J. Brush und E. 8. Dana. Ueber die Mineralfundstatte von Branchville, 
Connecticut: der Spodumen und seine Zersetzungsprodukte, 191.—A. 
Grosse-Bohle. Ueber das optische Verhalten des Senarmontils und der 
reguliren arsenigen Saure, 222.—A. Damour und G. vom Rath. Ueber 
den Trippkeit, eme neue Mineralspecies, 245.—M. Guyot. Ueber einen 
ungewohnlich grossen Huklaskrystall, 250.—F. Sansoni. Hin neues Vor- 
kommen von krystallisirten Manganspath, 250.—F. Sansoni. Pyrit vom 
Binnenthal, 252.—P. Groth. Beitrag zur krystallographischen Kenntniss 
des Wismuthglanzes, 252.—P. Groth. Hine Pseudomorphose aus dem 
Binnenthal, 253.—G. vom Rath. Mineralien von Zoptau, 253; Baryt 
in Basalt, 256; Mineralien von Copiapo in Chile, 256; Ueber Fahlerz 
von Horhausen, 258; Diaspro vom Greiner, 259.—D. M. Kramberger. 
Pilarit, ein neues Mineral aus der Gruppe des Chrysocolla, 260.—C. Délter. 


ADDITIONS TO THE LIBRARY. 263 


Krystallographisch-chemische Studien am Vesuvian, 289,—G. A. Konig. 

‘ Jarosit von einer neuen Fundstatte, 317.—G. A. Konig. Beegerit, ein 
neues Mineral, 322.—A. von Lasaulx. Mineralogische Notizen, 326.—H. 
Forstner. Cossyrit, ein Mineral aus den Liparitlaven der Insel Pantel- 
laria, 348.—H. Forstner. Ueber kunstlichen Wurtzit, 363.—C. Vrba. 
Mineralogische Notizen, 417.—J. Beckenkamp. Ueber die Ausdehnung 
monosymmetrischer und asymmetrischer Kvystalle durch die Warme, 436. 
—G. J. Brush. Ueber amerikanische Sulfoselenide des Quecksilbers, 467. 
—A. Arzruni und 8. Koch. Ueber den Analcim, 483.—G. vom Rath. 
Quarz und Feldspathe von Dissentis, 490.—G. vom Rath. Lamellare 
Zwillingsverwachsung des Augit nach der Basis, 495.—P. Groth. Hinige 
Erwerbungen der Strassburger Universitatssammlung, 496.—J. Lehmann. 
Krystallographische Mittheilungen,529.-M. Websky. Ueber Descloizit 
und Vanadinit aus La Plata, 542.—H. Bamberger und K. Feussner. So- 
dalith von Tiahuanaco, 580. 


Lille. Société Géologique du Nord. Annales. Tome yii.(1879-80). 
1881. 

Ladriére. Documents nouveaux pour l'étude du terrain dévonien des 
environs de Bavai, 1—Ladriére. Le terrain quaternaire du Nord, 11.— 
A. Rutot et EH. Vanden Broeck. Les phénomenes post-tertiaires en Bel- 
gique dans leurs rapports avec Vorigine des dépdts quaternaires et 
modernes, 33.—A. Potier. Lettre a M. Gosselet au sujet de l’argile a 
silex, 53.—Ortlieb. Compte-rendu d’une excursion géologique a Renaix, 
Belgique, 67.—De Lapparent. Note sur l’argile 4 silex du Nord de la 
France, 79.C. Barrois. Note sur les alluvions de la Serre, Aisne, 82. 
—A. Rutot. Note sur une coupe de terrain observée dans la Gare de 
Frameries prés Mons, Belgique, 92.—Gosselet. Notes sur les sables ter- 
tiaires du plateau de l’Ardenne, 100.—Potier. Resultats de deux sondages 
de Sangatte, 112.—F. Romer. Le genre Oldhama, Forbes, 115.— 
Gosselet. De Vusage du droit de priorité et de son application aux noms 
de quelques Spiriféres, 122.—Gosselet. Roches cristallines des Ardennes, 
132.—C. Barrois. Sur les recherches géologiques de Mr. G. K. Gilbert 
dans les monts Henry, 160.—Gosselet. Division a établir dans le terrain 
diluvien de la vallée de la Somme, 165.—C. Barrois. Note sur l’étage 
turonien de l’Irlande, 175.—C. Barrois. Note sur la fauna quaternaire 
de Sangatte, 181.—Ladriére. Observations sur le terrain crétacé des envi- 
rons de Bavai, 184.—Gosselet. Sondage de Menin, 188.—Van Ertborn. 
Sur la position du Diestien, et lage des sables blancs de Herenthals, 191.— 
J. Ortlieb. Lettre a M. le Maire de Tourcoing au sujet de 1’établisse- 
ment d’un Cimetiére, 192.—Gosselet. Troisiéme note sur le Famennien: 
tranchée du chemin de fer du Luxembourg; les schistes de Barvaux, 195. 
—Jannel. Note sur la présence de phosphates dans le lias des Ardennes 
et de la Meuse, 201.—Gosselet. Quatriéme note sur le Famennien: 
divisions a établir dans les schistes et les Psammites des environs de 
Maubeuge, 206.—Ladriére. Note sur les tranchées des chemin de fer 
d’Henin-Lietard a Carvin, 211.—P. Hallez. Discours présidentiel, 217. 
—T. Barrois. Rapport sur les travaux de la Société en 1878-79, 229.— 
Gosselet. Compte-rendu de l’excursion aux environs de Saint-Omer, 235. 
—N.de Mercey. Note sur la confusion résultant de l’emploi de la déno- 
mination d’argile a silex appliquée a deux dépéts placés, l’un a la base, 
et autre au sommet de la série tertiaire du Nord de la France, 237.—_N. 
de Mercey. Observations a occasion de quelques travaux publiés dans 
les Annales de la Société géologique du Nord sur le quaternaire ancien, 
246.—C. Barrois. Sur le terrain silurien supérieur de la presqwile de 
Crozon, 258.—Gosselet. Description géologique du canton de Berlaimont, 
270.—J. Ladriére. Etude sur les limons des environs de Bavai, 302.— 


264 ADDITIONS TO THE LIBRARY. 


P. Duponchelle. Compte-rendu de l’excursion du 29 aott au 7 sep- 
tembre 1879 dans les terrains primaires de l’Ardenne et de |’Hifel, 319. 
—C. Maurice. Compte-rendu de lexcursion du 3 au 7 septembre 1879 
dans les régions volcaniques de |’Hifel, 331.—C. Maurice. Compte-rendu 
de l’excursion géologique du 29 mars au 1° avril 1880 dans le Boulon- 
nais, 300.—P. Duponchelle. Gris-Nez et environs de Marquise, 360.— 
J. Coroénne. Compte-rendu de l’excursion du jeudi 29 avril 1880 4 
Sainghin, 369.—C. Maurice. Compte-rendu de l’excursion du 2 mai 
1880, 572.—J. Coroénne. Compte-rendu de l’excursion du 6 juin 1880, 
a Ath et a Lens, 376.—Trachet. Compte-rendu de excursion géologique 
du i6 au 18 mai 1880, dans V’arrondissement d’Avesnes, 379. 


Linnean Society. Journal. Zoology. Vol. xv. No. 82. 1880. 


. Nos. 83-85. 1880-81. 

EH. Ray Lankester. On the Tusks of the fossil Walrus found in the 
Red Crag of Suffolk, 144.—P. H. Carpenter. On the Genus Solanocrinus, 
Goldfuss, and its Relations to recent Comatule, 187. 


Botany. Vol. xvii. Nos. 106-112. 1880-81. 


——. Transactions. 2nd Ser. Botany. Vol. i. Parts 8 & 9. 
1880. 


Lisbon. Academia Real das Sciencias. Jornal de Sciencias Mathe- 
maticas, Physicas e Naturaes. Vol. vi. No. 24. 1878. 


, : Vol. vii. Nos. 25-28. 1879-80. 

J.F.N. Delgado. Correspondance relative a la classification des schistes 
siluriens a Néreites découverts dans le sud du Portugal, 103.—J. C. B. 
Cotter. Fosseis das bacias terciarias marinas do Tejo, do Sado e do 
Algarve, 112. 


——_., —_—_——-=-, 


Vol. viii. No. 29. 1880. 
——, —-—. Sessdo publica em 9 de Junho de 1880. 1880. 


: Memorias. Classe de Sciencias Mathematicas, Phy- 
sicas e Naturaes. Ser. 2. Tomov. Parte 2. 1878. 

J. Filippe Nery Delgado. ‘Terrenos paleozoicos de Portugal, sobre a 
existencia do terreno siluriano no Caixo Alemtejo.— Carlos Ribeiro. 
Estudios prehistoricos em Portugal. Noticia d’algumas estacoes e monu- 
mentos prehistoricos. Noticia da estagao humana de Licéa. 

. Sociedade de Geographia. Boletin. Ser. 2. Nos. 1-4. 
1880-81. 


Liverpool. Geological Society. Proceedings. Session 21st (1879-80). 
Vol. iv. Part 2. 1880. 

W.Semmons. Presidential Address, 98.—C. Ricketts. On the Car- 
boniferous Limestone near Skipton and in North Derbyshire, 132-—T, 
M. Reade. The Glacial Beds of the Clyde and Forth, 159.—R. A. Esk- 
rigge. Notes on human Skeletons and Traces of human Workmanship 
found in a Cave at Llandudno, 153.—W. Boyd Dawkins. Memorandum 
on the Remains found in the same Cave, 156. 


. Naturalists’ Field Club. Proceedings for the year 1879-80. 
1880. Presented by W. Whitaker, Esq., F.GS. 


London, Edinburgh, and Dublin Philosophical Magazine. Series 5. 
Vol. x. Nos. 59-64. 1880. Presented by Dr. Francis, F.L.S., 


——— 


ADDITIONS TO THE LIBRARY. 265 


London, Edinburgh, and Dublin Philosophical Magazine. Series 5. 
Vol. xi. Nos. 65-71. 1881. Presented by Dr. Francis, F.GS. 


London. Hertfordshire Natural-History Society and Field Club. 
Transactions. Vol.i. Part1l. 1880. 
J. E. Harting. Animals which have become extinct in Britain within 
historic times, 5. 
London Iron-Trades Exchange. Vol. xxvi. No. 1098. 1880. 
. Vol. xxvii. Nos. 1099-1124. 1880. 
The Minerals of New South Wales, 7, 66, 134, 169, 196, 237, 268. 
Vol. xxviii. Nos. 1125-1199.. 1881. 


Petroleum in Venezuela, 397.—Extinct Monsters in a Coal Mine, 536. 


Manchester. Geological Society. Transactions. Vol.xv. Parts 14—- 
18. 1880. 

J.B. Perrin and John Plant. On coucretionary Nodules of Limestone 
and foliated Mouldings in the Permian Marls near Leigh, 305.—W. Watts. 
Geological Strata, and Mode of Procedure in driving a Tunnel at Saddle- 
worth, 352.—J. D. Kendall. The Formation of Rock-basins, 368.—C. HE. 
De Rance. Further Notes of Triassic Borings near Warrington, 388. 


4 Vol. xvi. Parts 1-5. 1881. 

J. Dickinson. On the Discovery of Rock-Salt at Preesal, Lancashire 
26.—H. A. Woodward. On the Temperature of Strata and Seams of 
Coal taken in the new Sinkings of the Clifton and Kersley Coal Company 
at Newtown Collieries, Clifton, 29.—G. Wild. On marine fossil Shells 
at Ashton Moss Colliery Sinking, near Ashton-under-Lyne, 37.—W. 
Watts. On “ Arctic Peat Bog,” Rhodes Bank, Oldham, 43.—G. H. Kina- 
han. Jron-masked Rocks, 77.—J.T. Boot. Sections of Collieries and 
Bore-holes, 83. 


——. Literary and Philosophical Society. Memoirs. Series 3. 
Wolk vi. £379. 

Rooke Pennington. A Descent into Elden Hole, Derbyshire, 1.—A. 
W. Waters. Inquiries concerning a Change in the Position of the Earth’s 
Axis, 110.—C. A. Burghardt. On the Origin of some Ores of Copper, 171, 
253.—E. W. Binney. Notice of a fossil Plant found at Laxey, in the 
Isle of Man, 214.—J. B. Hannay. On Siliceous Fossilization, 234. 


. Proceedings. Vol. xvi. (1876-77). 1877. 
W. Boyd Dawkins. Address, 34.—J. Plant. Wealden Fossils from 
Columbia, 8. America, 50.—M. Stirrup. The Raised Beaches of the 
County Antrim, their Molluscan Fauna and Flint Implements, 51.—M, 
M. Pattison Muir. Note on a Manganese Ore from New South Wales, 
and on a Specimen of native Silver from New Zealand, 58.—H. Grim- 
shaw. On a Mineral Water from Humphrey Head, near Grange Sands, 
North Lancashire, 63.—E.W. Binney. A Notice of some organic Remains 
from the Schists of the Isle of Man, 102.—A. W. Waters. Inquiries 
concerning a Change of Position of the Earth’s Axis, 171.—John Plant. 
Footprints of Dinornis, Bones of Dinornis and Dodo, 181.—M. Stirrup. 
Upon Specimens from the Fresh Water Deposits of La Limagne d’Auvergne 
in Central France, 182.—E. W. Binney. Note on the Upper Coal-Measures 
of Canobie, Dumfriesshire, 192. 
——. ——. ——. Vol. xvii.(1877-78). 1878. 

A. W. Waters. Table of Effect of Movements of the Surface of the 
Globe on the Shifting of the Axis of the Earth, 2.—E. W. Binney. On 


266 ADDITIONS TO THE LIBRARY. 


some Coal-Measure Plants and other organic Remains from Spain, 5.— 
M. M. Pattison Muir. Note on an Edible Clay from New Zealand, 6. — 
C. A. Burghardt. On the Origin of some Ores of Copper, 27, 151.—E. 
W. Binney. Notice of a large Boulder Stone at Old Trafford, Manchester, 
54.—W. C. Williamson. On the Microscopic Conditions of a Slab from 
the Mountain Limestone of Bolland, 70.—K. W. Binney. Notice of a 
fossil Plant found at Laxey, in the Isle of Man, 85.—J. B. Hannay. On 
Siliceous Fossilization (Part 1), 97. 


Manchester. Literary and Philosophical Society. Proceedings. 
Vol. xvi. (1878-79). 1879. 

HK. W. Binney. On Boulders of Clay from the Drift, 40.—C. Grimshaw 
and H. Grimshaw. Ona further Analysis of the Water of the Mineral 
Spring at Humphrey Head, 45.—A. Bell. The Area of the Middle Drifts 
as determined by their Contents, 51.—J. B. Hannay. On Siliceous Fos- 
silization (Part 2), 75.—C. A. Burghardt. Mineralogical Notes, 111.— 
EK. W. Binney. On the Marine Shells found in the Lancashire Coalfields, 
117. 


——, ——. ——. Vol. xix. (1879-80). 1880. 

i. W. Binney. Notes on some Fossils from the Iron Mines of Furness, 
5.—J. Plant. Bog-butter (Butyrellite), from Co. Galway, Ireland, 70,— 
EK. W. Binney. Notes on a Bore through Triassic and Permian Strata 
lately made at Openshaw, 98.—J. Plant. On Flexible Sandstone, 103.— 
J. Plant. On the Use of Infusorial Earth, 106.—Watson Smith. The 
Castel Nuovo Lignite Deposit, near San Giovanni, Tuscany, 135. 


Melbourne. Royal Society of Victoria. Transactions and Proceed- 
mares) Wolk cyly Were 
A. W. Howitt. The Diorites and Granites of Swift’s Creek and their 
Contact Zones, with Notes on the Auriferous Deposits, 11.—J. Cosmo 
Newbery. Some new Localities for Minerals in Victoria, 144.—N. Taylor. 
Notes on the Geology of the West Tamar District, Tasmania, 155. 


Middlesbrough. Cleveland Institution of Engineers. Proceedings, 
1879-80. No.6. Presented by G. Barrow, Esq., F.GS. 
G. Barrow. On the Cleveland Ironstone (Part 2), 180. 


Mineralogical Society. Journal. Vol.iv. Nos. 17-19. 1880-81. 
J. H. Collins. On some Cornish Tin-stones and Tin-capels, 1, 103.— 
Heddle. The Geognosy and Mineralogy of Scotland, County of Caith- 
ness, 21.—J. P. Howley. List of Newfoundland Minerals, 36.—A. French. 
On a peculiar pasty Form of Silica from a Cavity in Gold-bearing Quartz, 
42.—Heddle. On anew Face on Crystals of Stilbite from two Localities, 
43.—T. A. Redwin. Further Notes on Mineral Growth, 96.—Heddle. 
Preliminary Notice of Substances which may prove to be new Minerals, 
117.—W. Terrill. On certain crystallized Products formed in Smelting 
Operations, 133.—Heddle. On the Geognosy and Mineralogy of Scotland : 
Sutherland, 185.—H. C. Bolton. Action of Organic Acids on Minerals, 
181.—Heddle. On Substances which may prove to be new Minerals, 189. 
—C.0O.Trechmann. On crystallized Olivine from Slag, 192. 


Montreal. Canadian Naturalist. N.S. Vol.ix. Nos. 6-8. 1880-81. 

J. W. Dawson. New Facts respecting the geological Relations and 
fossil Remains of the Silurian Ores of Pictou, Nova Scotia, 352.—G. F. 
Matthew. ‘Tidal Erosion in the Bay of Fundy, 368.—T. Sterry Hunt. 
Geological Notes, or Abstracts of recent Papers, 429.—C. U. Shepard. 
A peculiar Mineral of the Scapolite Family, 437.—J. W. Dawson. 
Revision of the Land Snails of the Paleozoic Era, with Descriptions of 


ADDITIONS 'TO THE LIBRARY. 267 


new Species, 449.—F. Bain. Note on Fossils from the Red-Sandstone 
System of Prince-Edward Island, 463. 


Montreal. Canadian Naturalist. N.S. Vol. x. No.1. 1881. 

J. W. Dawson. Paleontological Notes, 1—A. W. Wright. On the 
gence Substances contained in the Smoky Quartz of Br anchv ille, Conn., 

2.—G. M. Dawson. Note on the Geology ‘of the Peace-River Recion, 20. 
YS F, Whiteaves. Ona new Species of Ptertchthys, allied to Bothrio- 
lepis ornata, Kichwald, from the Devonian Rocks of the north side of the 
Baie des Chaleurs, 25. a F’, Whiteaves. Description of a new Species 
of Psammodus from the Carboniferous Rocks of the Island of Cape 
Breton, 36.—R. Chalmers. On the Glacial Phenomena of the Bay- 
Chaleur Region, 37.—C. Hoffmann. Recent Analyses of Canadian 
Minerals and River-waters, 55. 


Montrose. Natural-History and Antiquarian Society. Report of 
the Directors, 1881. 1881. Presented by W.S. Dallas, Esq., 
FLS. | | 

R. Barclay. Mineralogy of the Arbroath and Montrose Railway-cut- 

ting, 8. 

Moscow. Société Impériale des Naturalistes. Bulletin. Tome liv. 
(1879). No.4. 1880. 


Tome lv. Nos. 1 & 2. 1880. 

V. Kiprijanoff. Ueber fossile Fische des Moscauer Gouvernements, 1. 
—W. H. Twelvetrees. On a Labyrinthodont Skull (Platyops Rickardi, 
Twelvetr.) from the Upper Permian Cupriferous Strata of Kargalinsk 
near Orenbure, 117.—W. H. Twelvetrees. On Theriodont Humeri from 
the Upper Permian Copper-bearing Sandstones of Kargalinsk near Oren- 
burg, 123.—H. Trautschold. Zur Frage uber das Sinken des Meeres- 
spiegels, 174. 

Munich. K6niglich-bayerische Akademie der Wissenschaften. Ab- 
handlungen der mathematisch-physikalischen Classe. Band xii. 
Abth. 3. 1880. 


. Sitzungsberichte der mathematisch-physikalischen 
Classe. 1879, Hett 4. 1879. 


. 1880, Hefte 1-4. 1880. 

C. W. Giimbel. Geognostische Mittheilungen aus den Alpen, 164, 
542.—C. W. Giimbel. Ueber die mit einer Flissigkeit erfullten Chalce- 
donmandeln (Enhydros) von Uruguay, 241.—F. Pfaff. Petrographische 
Untersuchungen iiber die eocenen Thonschiefer der Glarner Alpen, 461. 

‘ . ——. 1881, Hefte lund 2. 1881. 

V. Kobell. Ueber Polarisationsbilder an Zwillingen zweiaxiger Kry- 
stalle, 199.—K. Haushofer. Ueber das Verhalten des Dolomit gegen 
Fissigsaure, 220. 

Nancy. Société des Sciences. Bulletin. Série2. Tome iv. Fasc. 10 
Gis79). 18380. 

Fliche. Sur les lignites quaternaires de Jarville, 8.—Braconnier. 
Description des terrains qui constituent le sol du département de Meurthe- 
et-Moselle, 12. 


_—_-,. 


Tome iv. Fasc 11 (1880). 1880. 


Nature. Vol. xxii. Nos. 556-574. 1880. 
J.S. Gardner. A Chapter in the History of the Conifere, 199,—J. 


268 ADDITIONS TO THE LIBRARY. 


Milne. Seismology in Japan, 208.—T. S. Humpidge. New Metals, 
232.—The Caribbean Sea, 242.—N.S. Maskelyne. Prof. W. H. Muller, 
247.—Artificial Diamonds, 255.—A. Geikie. North-American Geology : 
Idaho and Wyoming, 268.—J. S. Gardner. Observations on Arctic 
fossil Floras with regard to Temperature, 341.—J. Murray. The Struc- 
ture and Origin of Coral Reefs and Islands, 8351.—A. C. Ramsay. Address 
to the British Association: On the Recurrence of certain Phenomena in 
Geological Time, 383.—H. C. Sorby. Opening Address to the Geological 
Section of the British Association, 390.—A. Geikie. A Fragment of 
Primeval Europe, 400.—A. Agassiz. Paleeontological and Embryological 
Development, 424.—British Association: Reports of Committees, 442 ; 
Section C, Geology, 449.—H. B. Guppy. The Yang-tse, the Yellow 
River, and the Pei-ho, 487.—Landslips, 505.—Richard Owen, 577.—A. 
Zittel. The Geology of the Libyan Desert, 587. 


Nature. Vol. xxiii. Nos. 575-600. 1880-81. 

A. Geikie. The Lava-fields of North-western Europe, 3.—A. Murray. 
Mineral Resources of Newfoundland, 46.—Curious Impressions in Cam- 
brian Sandstone near Loch Maree, 93.—J. Thomson. Notes on the 
Geology of East Central Africa, 102.—C. J. Merriman. Subterranean 
Forest in India, 105.—British Earthquakes, 117.—A. Geikie. The 
Mineralogical Society of Great Britain and Ireland, 150.—A. Geikie. 
Geology of Bosnia and Herzegovina, 224.—J.S. Gardner. A Chapter 
in the History of the Conifer, 251, 412.—_J. 8. Gardner. Geologising 
at Sheppey, 293.—C. E. De Rance. The Blackheath Holes, 865.—Inter- 
esting new Crinoids, 377.—Dr. J. J. Bigsby, 389.—The recent Discovery 
of the Body of Rhinoceros Merckti in Siberia, 466.—Gold in Newfound- 
land, 472.—V. Ball. On the Identity of some ancient Diamond-mines 
in India, especially those mentioned by Tavernier, 490.—H. J. Johnston- 
Lavis. The Earthquake in Ischia, 497.—C. E. De Rance. The Inter- 
national Geological Congress, 510.—Achille Delesse, 535.—The new 
Museum of Natural History, 549.—Dunes amd Moving Sand, 569.—Sir 
Philip de Malpas Grey Egerton, 579.—C. A. Stevenson. Harthquake of 
November 28, 1880, in Scotland and Ireland, 591. 


. Vol. xxiv. Nos. 601-607. 1881. 
T. G. Bonney. A Geologist’s Notes on the Royal Academy, 86. 


Neuchatel. Société des Sciences Naturelles. Bulletin. Tome xii. 
Cahier 1. 1880. 


E. Desor. Sur la mer saharienne, 16.—Ritter. De l’action des vagues 
sur les sables des bords du lac de Neuchatel, 114. 


Newcastle-upon-Tyne. Natural-History Society &c. Natural- 
History Transactions. Vol. vii. Part 2. 1880. 

R. E. Hooppel. Presidential Address to the Members of the Tyneside 
Naturalists’ Field Club, 187—W. Howchin. Notes on a Find of Pre- 
historic Implements in Allendale, with Notices of similar Finds in the 
surrounding District, 210.—J. Wright. Short Memoir of the Life of 
Thomas Belt, F.G.8., 235.—Hugh Miller. Tyneside Hscarpments: their 
Preglacial, Glacial, and Postglacial Features, 285.—R. Howse. Pre- 
liminary Note on the Discovery of old Sea-caves and a raised Sea-beach 
at, Whitburn Lizards, 8361.—R. Howse. Note on the Priority of Discovery 
of Anodon Jukesti, Forbes, in the Lower Carboniferous Rocks of North 
Northumberland, 376. 


—. North-of-England Institute of Mining Engineers. Trans- 
actions. Vol. xxix. (1879-80). 1880. 


ADDITIONS TO THE LIBRARY, 269 


New Haven, Conn. American Journal of Science. Ser. 3. Vol. xx. 
Nos. 115-120. 1880. 


J.D. Dana. Geological Relations of the Limestone Belts of West- 
chester County, N. Y., 21, 194, 359, 450.—C. A. White. Antiquity of 
certain subordinate Types of Fresh-water and Land Mollusca, 44.— 
C. U. Shepard. Mineralogical Notices, 54.—J. M. Stillman. Bernar- 
dinite; its Nature and Origin, 93.—G. J. Brush and E. 8. Dana. On 
Crystallized Danburite from Russell, St. Lawrence, New York, 111.— 
J. F. Whiteaves. On a new Species of Prerichthys, allied to Bothrio- 
lepis ornata, Eichwald, from the Devonian Rocks of the North Side 
of Baie des Chaleurs, 132.—C. D. Wallcott. The Permian and other 
Paleozoic Groups of the Kanab Valley, Arizona, 221.—O. C. Marsh. 
Notice of Jurassic Mammals representing two new Orders, 235.—G. J. 
Brush and E. 8. Dana. Mineral Locality at Branchville, Connecticut : 
(fourth paper) Spodumene and the Results of its Alteration, 257.—A. 
Agassiz. Paleontological and Embryological Development, 294, 375.— 
J. W. Dawson. Revision of the Land Snails of the Paleozoic Era, with 
Descriptions of new Species, 403—W. O. Crosby and G. H. Barton. 
Extension of the Carboniferous Formation in Massachusetts, 416.—E. T. 
‘Cox. Discovery of Oxide of Antimony in extensive Lodes in Sonora, 
Mexico, 421.—H. 8. Willams. Abstract of some Paleontological Studies 
of the Life History of Sprrifer levis, H., 456. 


ooo 


: : . Vol. xxi. Nos. 121-126. 1881. 

G. W. Hawes. The Albany Granite, New Hampshire, and its Contact 
Phenomena, 21.—C. Barrois. Review of Professor Hall’s recently pub- 
lished Volume on the Devonian Fossils of New York, 44.—Earthquake 
at the Philippine Islands, of July 1880, 52.—W. H. Dall. Notes on 
Alaska and the Vicinity of Behring Strait, 104.—S. H. Seudder. Relation 
of Devonian Insects to later and existing Types, 111.—C. U. Shepard. 
Meteoric Iron of Lexington Co., 5. C., 117.—G. F. Wright. Date of the 
Glacial Era in Eastern North America, 120.—P. Collier. A remarkable 
Nugget of Platinum, 123.—R. P. Whitfield. A new Genus and Species 
of Air-breathing Mollusk from the Coal-measures of Ohio, 125.—J. L. 
Smith. Hiddenite,a Variety of Spodumene, 128.—S. W. Ford. Remarks 
on the Genus Obolella, 13i.—H. M. Chance. The Millstone Grit in 
England and Pennsylvania, 134.—O. C. Marsh. Principal Characters 
of American Jurassic Dinosaurs, 167.—S. H. Scudder. Structure and 
Affinities of Zuphoberia of Meek and Worthen, 182.—C. G. Rockwood. 
Recent American Earthquakes, 198.—G. W. Hawes. Liquid Carbon 
Dioxide in Smoky Quartz, 203.—A. W. Wright. Gaseous Substances 
contained in Smoky Quartz of Branchville, Conn., 209.—W. C. Kerr. 
Origin of new Points in the Topography of North Carolina, 216.—W. P. 
Blake. Occurrence of Realgar and Orpiment in Utah Territory, 219.— 
O. C. Marsh’s Monograph on tht Odontornithes or Toothed Birds of 
North America, 255.—W. E. Hidden. Whitfield County, Georgia, 
Meteoric Iron, 286.—J. EK. Hilgard. The Basin of the Gulf of Mexico, 
988.—E. A. Smith. The Geology of Florida, 292.—G. J. Brush. On 
American Sulpho-Selenides of Mercury, with Analyses of Onofrite from 
Utah by W. J. Comstock, 312.—H. 8. Williams. Channel-fillings in 
Upper Devonian Shales, 318.—O. C. Marsh. New Order of extinct 
Jurassic Reptiles (Cceluria), 539.—O. C. Marsh. Discovery of a fossil 
Bird in the Jurassic of Wyoming, 341.—O. C. Marsh. American Ptero- 
dactyles, 342.—W. C. Kerr. Action of Frost in the Arrangement of 
superficial Earthy Material, 345.—N. H. Winchell. Dall’s Observations 
on Arctic Ice, and the Bearmeg of the Facts on Glacial Phenomena in 
Minnesota, 358.—J. P. Cooke. William Hallowes Miller, 379.—G. M. 


270 ; ADDITIONS TO THE LIBRARY. 


Dawson. Geology of Peace-River Region, 891.—O. C. Marsh. American 
Jurassic Dinosaurs, 417.—J. D. Dana. Geological Relations of the 
Limestone Belts of Westchester County, New York: South Westchester 
County and Northern-New- York Island, 425.—J. L. Smith. Nodule of 
Chromite in the Interior of compact Meteoric Iron from Cohahuila, 461. 

J. F. Whiteaves. Fossil Fishes from the Devonian Rocks of Scaumenac 
Bay, in the Province of Quebec, 494.—O. C. Marsh. New Jurassic 
Mammals, 511. 


New Haven, Conn. Peabody Museum of Yale College. Memoirs. 
Moluis) S30: 
O. C. Marsh. Odontornithes: a Monograph on the extinct Toothed 
Birds of North America, 1. 


New York. Lyceum of Natural History. Annals. Vol. xi. No. 
13 (Index and Contents). 1876. 


: . N.S. Volii.) Nos. 9-135) te7gSsu: 
A. A. Julien, On Spodumene and its Alterations, 318. 


— 
r) 


New York. The Engineering and Mining Journal. Vol. xxxi. 
Nos. 1-11, 18-22. 1881. Presented by Dr. C. Le Neve 
Foster, F.GS. 

AC? Campbell. The Copper and Iron-bearing Rocks of Lake Su- 
perior, 20.—W. B. Devereux. Gold and its aeeO aves Minerals at 
King’s } Mountain, North Carolina, 59. The Mica Veins of 
North Carolina, 211.—A. Murray. The Gold Ree near Brigus, New- 
foundland, 252. “The Gold Mines of the Rhetian ‘Alps, 281.—J. 8. New- 
berry. The Genesis of our Iron Ores, 286, 298.—I’. M. F. Cazin. The 
Triassic Age of Sand-Rock carrying ores oe Copper and Silver, and the 
origin of these Ores, 300.—The History of Tin, 313.—The Geological 
formation, Tombstone, Arizona, 316. 


Norwich Geological Society. Proceedings. Vol.i. Part 5. 1881. 

H. B. Woodward. Ona Disturbance of the Chalk at Trowse, 109.— 
J. Reeve. Additions to List of Fossils from the Norwich Crag at 
Bramerton, 110.—H. Norton. On the Paleontology of the Ancients 
(Greeks and Romans), 110.—A. J. Jukes Browne. ‘The Subdivisions of 
the Chalk, 113.—J. H. Blake. Well-boring at Hast Dereham Water- 
works, 126, —C. Reid. Well-boring at Cromer Waterworks, 129.—H. 
Norton. Derivation of the Word “ Paramoudra,” 132.—J. H. Blake. On 
the Age and Relation of the so-called Forest-bed of the Norfolk and 
Suffoll: Coast, 137. 


Norwich. Science Gossip Club. ‘Report of Proceedings at the 
Annual Meeting, June 11, 1874. 1874. Presented by H. B. 
Woodward, Esq., F.GS. : 


. Report of Proceedings at the Annual Meetings 
June 6, 1877, and May 14, 1878. 1878(?). Presented by 
Ed Woodward, Esq., PGS. 

S. ©. Sothern, Presidential Address, 6.—H. B. Woodward. Presi- 
dential Address, 29. | 


Nottingham. The Forester, or the Nottingham-High-School Maga- 
zine. Vol. i. 1878. Presented by W. Whitaker, Esq., £.GS. 


ADDITIONS TO THE LIBRARY. 271 


Nottingham. The Forester, or the Nottingham-High-School Maga- 
ame,” Vol. it. 1879. ‘Presented “by W. Whitaker, Esq., 
GS. 

The Sandstones of the Carboniferous and Triassic Rocks, 68.—Geo- 

logical Excursion to Charnwood Forest, 94. 


. —. Vol. ni. 1880. Presented by W. Whitaker, Ksq., 
F.GS. 
The Origin of Sandstones, 36. 


Paleontographical Society. Monographs. Vol. xxxv. (1881). 1881. 
(Two copies.) 

T. Wright. The Cretaceous Echinodermata, vol. i. part 9.—T. David- 
son. Supplement to the Fossil Brachiopoda, vol. iv. part 4.—J. Lycett. 
The Fossil Trigoniz : Supplement.—T. Wright. The Lias Ammonites, 
Part 4—R. Owen. The Reptilia of the Liassic Formation, Part 3.— 
L. Adams. The Fossil Elephants (£. primigenius and meridionalis), 
Part 3. 


Paris. Académie des Sciences. Comptes Rendus. Tome xcii. 
Nos. 1-24.. 1881. 

W. L. Green. Nouvelle irruption du Mauna-Loa (iles Hawai), 48.— 
A. Daubrée. Substances cristallines produites aux dépens de médailles 
antiques, immergées dans les eaux thermales de Haracci, commune 
d’Olmeto (Corse), 27.—A. Daubrée. Production contemporaine du soufre 
natif dans le sous-sol de Paris, 101.—B. Studer. Le contact mécanique 
du gneiss et du calcaire, dans i’Oberland bernois, observé par M. A. 
Baltzer, 169.—A. Daubrée. Hxamen de matériaux provenant de quelques 
forts vitrifiés de Ja France; conclusions qui en résultent, 269.—D. Col- 
ladon. Sur le tremblement de terre qui a été ressenti en Suisse, le 27 
janvier 1881, 530.—S. Meunier. Examen lithologique et géologique de 
la météorite tombée le 13 Octobre 1872 aux environs de Soko-Banja, en 
Serbie, 331.—F. Fouqué et A. M. Lévy. Reproduction artificielle des 
basaltes, 367.—A. Daubrée. Sur les réseaux de cassures ou diaclases qui 
coupent la série des terrains stratifiés; nouveaux exemples fournis par 
les couches crétacées aux environs d’Htretat et de Dieppe, 393.—A. Cara- 
ven-Cachin. Ancienneté de lElephas primigenius (Blum.) dans le bassin 
sous-pyrénéen, 475.—Desor. Ossements humains trouyvés dans le dilu- 
vium de Nice; examen de la question géologique, 746.—Niepce. Osse- 
ments humains trouvés dans le diluvium de Nice; description des 
ossements, 749.—Quatrefages. Ossements humains trouvés dans le 
diluvium de Nice; détermination de la race, 750.—A. Gaudry. Sur un 
nouveau genre de poisson primaire, 752.—A. Julien. Sur l’existence et 
les caractéres du terrain cambrien dans le Puy-de-Déme et dans |’ Aller, 
754.—Dieulafait. Loi générale de formation des eaux minérales salines ; 
application au cas particulier de Gréoux (Basses-Alpes), 756.—L. Crié. 
Sur la découverte, a Noirmoutiers (Vendée) de la flore éocéne a Sabalites 
andegavensis, Sch.,759.—A. Julien. Sur la nature et Vordre d’apparition 
des roches éruptives anciennes que l’on observe dans la région des volcans 
a cratéres du Puy-de-Dome, 799.—Des Cloizeaux et Damour. Note sur 
la chaleoménite, nouvelle espéce minérale (sélénite de cuivre), 837.— 
J. L. Smith. Anomalie magnétique du fer météorique de Sainte-Cathe- 
rine, 843.—Mayencgon. Sur la bismuthine produite par les houilléres 
incendiées, 854.—F’. Fouqué et A. Michel Lévy. Reproduction arti- 
ficielle des diabases, dolérites et météorites 4 structure ophitique, 890.— 
A. Julien. Sur le terrain dévonien de Diou (Allier) et de Gilly (Sadéne- 
et-Loire), 891.—H. Filhol. Sur les différentes espéces d’Ours dont les 


272 ADDITIONS TO THE LIBRARY. 


débris sont ensevelis dans la caverne de Lherm (Ariége), 929.—Le Cha- 
telier. Production d’un silicate de baryte hydraté en cristaux, 931.— 
EK. Mallard. Sur la production d’un phosphure de fer cristallisé et du 
feldspath anorthite, dans les incendies des houilléres de Commentry, 933. 
De Pellissier. Sur le tremblement de terre de Chio, 956.—G. Rolland. 
Sur les grandes dunes de sable du Sahara, 968.—A. Daubrée. Examen 
de matériaux provenant des forts vitrifiés de Craig Phadrick, prés Inver- 
ness (Ecosse) et de Hartmannswillerkopf (Haute-Alsace), 980.—A. 
Daubrée. Météorite tombée & Louans (Indre-et-Loire) le 25 janvier 
1845 et dont la chute est restée inédite, 984.—Lawrence Smith. Nodule 
de chromite dans Vintérieur du fer météorique de Cohahuila (Mexique) 
(météorite de Butcher), 991.—F. Fouqué. Sur la série stratigraphique 
des roches qui constituent le sol de la haute Auvergne, 1089.—F. Fouqué 
et M. Lévy. Hxamen de quelques produits artificiels de James Hall, 
1040.—E. Filhol. Sur quelques feldspaths de la vallée de Bagnéres-de- 
Luchon (Haute-Garonne), 1059.—A.Gaudry. Sur les plus anciens rep- 
tiles trouvés en France, 1145.—H. Fayol. Etudes sur le terrain houiller 
de Commentry, 1172.—G. de Saporta et A. F. Marion. Sur les genres 
Williamsonia, Carruth. et Goniohna, VOrb., 1185, 1268.—F. Pisani. Sur 
un vanadate de plomb et de cuivre du Laurium, 1292.—A. Julien. Sur 
Vexistence du terrain cambrien a Saint-Léon et Chatelperron (Allier), 
1293.—H. Fayol. Sur le terrain houiller de Commentry; expériences 
faites pour en expliquer la formation, 1296.—De Lesseps. Sur le rap- 
port de M. le commandant Roudaire, relatif 4 sa derniére expédition dans 
les chotts tunisiens, 1309.—K. Hébert. Observations sur les résultats 
géologiques fournis par les missions de M. le commandant Roudaire dans 
les chotts tunisiens, 1510.—A. Damour. Nouvelles analyses sur la jadeite 
et sur quelques roches sodiféres, 1312.—C. Friedel et E. Sarrasin. Sur 
la reproduction par voie aqueuse du feldspath orthose, 1874.—A. Julien. 
Sur la faune carbonifére de Régny (Loire) et ses relations avec celle de 
V’Ardoisiére (Allier), 1451. 


Paris. Annales des Mines. Série 7. Tome xvii. 2° et 3° livr. de 
1880. 1880. 


Delesse et de Lapparent. LExtraits de géologie pour les années 1877 
et 1878, 209.—Petiton. Note sur les mines de l’archipel des Ferde (iles 
de Sudérée et de Naalsde), 305.—F. Rigaud. Notice sur les travaux 
exécutés 4 Bourbonne-les-Bains, 349, 


Tome xviii. 4°-6° livr. de 1880. 1880. 

E. Boutan. Note sur la constitution géologique de Visthme de Panama, 
au point de vue de l’exécution du canal interocéanique, 5.—H. Sauvage. 
Notice sur les sources minérales des départements de Seine-et-Oise, de 
Seine-et-Marne et du Loiret, 102.—Ferrand. Notice sur les mines 
d’anthracite de la Mure, 121.—A. Carnot. Note sur deux variétés de 
diadochite (phosphosulfate de fer) trouvées dans la mine d’anthracite de 
Peychagnard (lsére), 148.—G. Rolland. Mission transsaharienne de 
Laghouat, El-Goléah, Ouargia, Biskra: géologie et hydrologie, 152.— 
Domeyko. Note sur des cristaux épigénes de cuivre métallique prove- 
nant des mines de cuivre de Coro-Coro en Bolivie, 531.—Domeyko. 
Note sur les minéraux de bismuth de Bolivie, du Pérou et du Chili, 
538. 


: 4 Tome xix. 1%e livr.de 1881. 1881. 

Villot. Note sur le régime de la source de Camoins-les-Bains (com- 
mune de Marseille), 5, 157.—M. Du Chatenet. Htat actuel de Vindus- 
trie minérale dans le Cerro de Pasco, 61. 


ADDITIONS TO THE LIBRARY. 273 


Paris. Annales des Sciences Géologiques. Tomex. 1879. Pur- 
chased. 

C. Barrois. Mémoire sur le terrain crétacé du bassin d’Oviédo, No. 1.— 
Cotteau. Notice sur les Echinides urgoniens recueillis par M. Barrois 
dans la province d’Oviédo (Hspagne), No. 2.—H. Filhol. Etude des 
mammiféres fossiles de Saint-Géraud le Puy (Allier), No. 3. 

: Tome xi. Nos.1&2. 1880. Purchased. 

H. Filhol. Etude des mammiféres fossiles de Saint-Géraud le Puy 

(Allier), 1, 49. 


Pe Hobart. 


——. Annales des Sciences Naturelles. Série 6. Zoologie et Pa- 
léontologie. Tomeix. Nos. 1-6. 1880. Purchased. 


«6 Lome x. Nos. 1-6. 1880. Purchased. 


——. Annales Hydrographiques. Série 2. Tomei. 2° semestre 
1879. 1879. Presented by the Dépét de la Marine. 


a  ~=—index to vols. xxix. to xit. 1879, 


Nome) xa. Nos) Po& 2.) 1830.- Presented by 


——. Association francaise pour lVAvancement des Sciences. 
Compte-rendu de la 8° Session (Montpellier, 1879). 1880. 
Purchased. 

Garrigou. Des sources minérales mercurielles de la France, analyse 
des eaux minérales, 474.—A. Toucas. Notice sur le terrain crétacé des 
Corbiéres, 574.—V. Lemoine. Sur les ossements fossiles des terrains 
tertiaires inférieurs des environs de Reims, 585.—E. Riviére. De quelques 
hyperostoses de poissons trouvées dans les grottes quaternaires de Menton 
en Italie, 595.—F. Pommerol. Le mouflon quaternaire, 600.—A. Jean- 
jean. tude sur l’oxfordien supérieur, le corallien et le néocomien infé- 
rieur, dans les Cévennes, 610.—P. de Loriol. Les crinoides fossiles de 
la Suisse, 627.—C. Grad. Sur la formation des charbons feuilletés gla- 
ciaires de la Suisse, 636.—Cazalés de Fondouce. Erosion de cailloux 
quaternaires due 4 l’action du vent et du sable, 646.—P. de Loriol. Note 
sur les échinides recueillis dans les expéditions du ‘Challenger’ et du 
‘Blake, 650.—G. Cotteau. Considérations stratigraphiques et paléon- 
tologiques sur les échinides de l’étage cénomanien de l’Algérie, 655,— 
Moriére. Note sur une station de silurien 4 la Bréche-au-diable (Cal- 
vados), 663.—C. Barrois. Sur l’étendue du systéme tertiaire inférieur 
dans les Ardennes et sur les argiles a silex, 666.—C. Barrois. Sur le 
marbre griotte des Pyrénées, 668.—C. Barrois. Sur la faune troisiéme 
silurienne du Finistére, 669.—E. Riviére. Le pliocéne de Castel d’Appio 
en Italie, nomenclature des fossiles quwil renferme, 670.—Ollier de 
Marichard. Les troglodytes de ’Ardéche ou premiére page inédite de 
Vhistoire du Vivarais, 797.—Pommerol. Le gisement quaternaire de 
Sarliéve, 301—F. Daleau. Les stations préhistoriques des étangs 
d@Hourtin et de Lacanau (Gironde), 807.—P. Fiére. Grottes préhis- 
toriques du Dauphiné, 852. 


Paris. Dépot de la Marine. Annuaire des Marées des cétes de 
France pour ’an 1881. 1880. 


——. Journal de Conchyliologie. Série 3. Tome xx. Nos. 3 et 4. 
1880. Purchased. 
R. Tournouér. Description d’un Ostrea fossile de la mollasse miocéne 
VOL, XXXVII. x 


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de Forcalquier, Basses-Alpes, 256.—H. Douvillé. Sur la forme de 
Vouverture de ? Ammonites pseudo-anceps, 355. 


Paris. Journal de Conchyliologie. Serie 3. Tome xxi. No. 1. 
Purchased. 


——. Muséum d’Histoire Naturelle. Nouvelles Archives. Série 2. 
Tomenium.)/Hascylet 24) isso: 

S. Meunier et J. Lambert. Recherches stratigraphiques et paléon- 

tologiques sur les sables marins de Pierrefitte prés Htampes, Seine-et-Oise, 


235.—A.T.de Rochebrune. Révision des ophidiens fossiles du Muséum 
de histoire naturelle, 272. 


Revue Scientifique de la France et de l’Etranger. Série 2. 
Tome xix. Nos. 1-26. 1880. 

Fouqué. De la pétrologie, de la place qu’elle doit occuper dans les 
programmes universitaires et des modifications principales a apporter 
dans ces programmes en ce qui regarde les sciences géologiques, 18.— 
G. Rolland. La mission transsaharienne El Goleah, 53.—C. Velain. 
La chorologie des sédiments et sa signification pour Ta géologie et la 
théorie de la descendance, 204. —Friedel. La reprog ern des minéraux, 
242.—C. Barrois. Paléontologi ie de |’ Etat de New York, 257.—J. Thoulet. 
Contribution a V’étude des propriétés physiques et chimiques des miné- 
raux microscopiques, 283.—J. Roche. La mission d’exploration trans- 
saharienne, 505.—J. Bayol. La mission du haut Niger, 601. 


. Tome xix. Nos. 1-25. 188ie 

Faye. Des voleans de la lune, 130.—Les sources de naphte dans la 
région du Caucase, 337.—C. Velain. L’Algérie et le pays des Kroumits,, 
545.—G. Rolland. - Les orandes dunes de sable du Sahara, 609. 


——. Société Géologique de France. Bulletin. Série 3. Tome vi. 
(US 73)S Nosy Iréc LO rSss0: 

A. Cordella. Note sur les mines du Laurium et sur les nouveaux gites 
de mineral de zine (Smithsonite), 577.—Loustau et Belhomme. Note 
sur un sondage exécuté a Monsoult (Seine-et-Oise), 581.—G. Dollfus. 
Observations sur le sondage de Monsoult, 585.—Bonneau du Martray. 
Note sur un bloc erratique situé dans la yallée de la Dragne, prés de 
Moulins-Engilbert (Niévre), a 2 kilométres environ de la faille occiden- 
tale du Morv van, 098.—Torcapel. Les glaciers quaternaires des Cévennes, 
600.—Daubrée. Expériences sur la production de déformations et de 
cassures par glissement, 608.—Potier. Sur la direction des cassures dans 
les corps isotropes, 609.—H¢hert et Munier-Chalmas. Recherches sur 
les terrains tertiaires du Vicentin, 610, 619.—R. Zeiller. Sur une nou- 
velle espéce de Decranophyllum, 611 —Leymerie. Observations sur le 
mémoire de M. Peron sur les calcaires a Echinides des bains de Rennes, 
616.—Peron. Réponse aux observations de M. Leymerie, 616.—Cotteau. 
Sur Vexposition géologique et paléontologique du Havre, 618.—Tournouer. 
Sur les cérites des marnes a Happarion du puits Kharoubi, prés Oran, 
618.—Tournouer. Sur les coquilles marines trouvées dans la région des 
chotts sahariens, 619.—K. Riviére. Note sur la grotte de Grimaldi, 621. 
—Sauvage. Note sur les poissons fossiles, 625.—Tardy. De la limite 
entre le Crétacé et le Tertiaire aux environs de Vitrolles (Bouches-du- 
Rhone), 637.—C. Vélain. Compte- -rendu de l’excursion 4 Meudon, 654. 
—Tournouér. Compte-rendu de l’excursion 4 Etampes, 663. —De Lap- 
parent. Hxcursion dans le pays de Bray, 675.—De Mercey.. Compte- 
rendu de Vexcursion a Maigneley, 679.—O. Vélain. Compte-rendu de 
Vexcursion de la Frette a Sannois, 687.—H. Douvillé. Compte-rendu de 
Vexcursion & Vernon (1° partie), 694.—De Chancourtois. Compte-rendu 


ADDITIONS TO THE LIBRARY. 275 


de Vexcursion a Vernon (2° partie), 697.—De Chancourtois. Sur les 
alionements géologiques relevés dans les environs de Vernon, 703.—H. 
Douyillé. Résumé de l'état de la question des sables dits éruptifs, 706. 
—C. Vélain. Compte-rendu de l’excursion a Cuise-la-Motte, 711. 


Paris. Socicté Géologiquede France. Bulletin. Série 3. Tome vii. 
(1879). Nos. 5-8. 1880. 

C. Vélain. Sur la constitution géologique des iles Seychelles, 278.— 
A. Falsan. Sur la position stratigraphique des terrains tertiaires supéri- 
eurs et quaternaires & Hauterives (Dréme), 285.—A. Locard. Observa- 
tions paléontologiques sur les couches a Ostrea Falsani dans les environs 
de Hauterives (Drome), 307.—A. de Lapparent. Note sur l’ouvrage de 
M. Struckmann intitulé “ Le Jura supérieur des environs de Hanovre,” 
315.—G. Dollfus. Les dépdots quaternaires du bassin de la Seine, 318.— 
A. de Lapparent. Sur la disposition générale des reliefs du globe ter- 
restre, 346.—N. de Mercey. Remarques sur la classification du terrain 
crétacé supérieur, 355.—Tournouér. Sur l'étude du bassin de Visan par 
M. Fontannes, 587.—C. Vélain. Notes géologiques sur la Haute-Guy- 
ane, d’aprés les explorations du Dr. Crevaux, 388.—V. Lemoine. Oi- 
seaux fossiles des terrains tertiaires inférieurs des environs de Reims, 
398.—F. Delafond. Note sur l’existence du terrain néocomien aux envi- 
rons de Tournus (Saéne-et-Loire), 403.—E. Hébert. Sur la position des 
sables de Sinceny, 408.—A. Boué. Sur la vallée de la Soukava, 412.— 
C. Vélain. Ile de Pagues, 415.—Delage. Etude du calcaire de Lorman- 
diére, 426.—Meugy. Observations sur une note de M. Barrois, relative au 
terrain crétacé des Ardennes, 445.—Lebesconte. Sur le bassin tertiaire 
des environs de Rennes, 451.—Tournouér. Etude sur les fossiles de 
létage tongrien (d’Orbigny) des environs de Rennes en Bretagne, 464.— 
E. Benoit. De Vextension géographique et stratigraphique du purbeckien 
dans le Jura, 484.—F. Fontannes. Note sur la découverte de deux 
espéces nouvelles du genre Antedon dans les terrains tertiaires supérieurs 
du bassin du Rhéne, 497.—Tardy. Le dernier diluvium quaternaire, 
500.—Tardy. Deuxiéme note sur le chronométre de la Sadne, 514.— 
A. Gaudry. Allocution présidentiel, 517.—KH. Jannettaz. Notice nécro- 
logique sur M. G. Delafosse, 524.—L. Lartet. Vie et travaux d’Alex- 
andre Leymerie, 530.—V. Lemoine. Note sur les ossements fossiles des 
terrains tertiaires inférieurs des environs de Reims, 558.—Rey-Lescure. 
Carte géologique de Tarn-et-Garonne, 562. 

: : i Tome vii. Séance Generale An- 
nuelle et Célébration du Cinquantenaire de la Société. 1880. 


Penzance. Royal Geological Society of Cornwall. Transactions. 


Wolke: Part 3. 1331. 

A. P. Vivian. Presidential Address, Ixxxvi.—W. A. E. Ussher. 
Pleistocene Notes on the Cornish Coast between Plymouth and Looe, 68. 
—W. A. E. Ussher. The Devonian Rocks between Plymouth and East 
Looe, 70.—W. W.Smyth. On the Occurrence of Feather Ore (Plumosite) 
in Foxdale Mine, Isle of Man, 82.—C. W. Peach. On Fossils’ from the 
Rocks of Cornwall, some of them new to the list, 90.—J. H. Collins. Note © 
on the Occurrence of Stanniferous Deer-horns in the Tin-gravels of Corn- 
wall, 98.—J. H. Collins. Note on the supposed Serpentine of the Parish 
of St. Veep, 101.—R. N. Worth. Notes on the Geology of the South-east 
Border of Cornwall, 103. 

Philadelphia. Academy of Natural Sciences. Journal. Series 2. 
Vol. viii: Part 4. 1881. 

W.M. Gabb. Description of Caribbean Miocene Fossils, 8337.—W. M. 

Gabb. Description of new Species of Fossils from the Pliocene Clay Beds 
“2 


276 ADDITIONS TO THE LIBRARY. 


between Limon and Moen, Costa Rica, together with Notes on previously 
known Species from there and elsewhere in the Caribbean Area, 349. 


Philadelphia. Academy of Natural Sciences. Proceedings, 1880. 


Parts 1-3. 1880. 

A. Heilprin. On the Stratigraphical Evidence afforded by the Tertiary 
Fossils of the Peninsula of Maryland, 20.—A. W. Vogdes. Description 
of a New Crustacean from the Upper Silurian of Georgia, with Remarks 
upon Calymene Clintoni, 176.—T. D. Rand. Serpentine Belts of Radnor 
Township, Delaware Co., 225.—H. C. Lewis. A Garnet with inverted 
Crystallization, 241.—T. D. Rand. Change of Serpentine into Quartz, 
241.—H. C. Lewis. Magnetite Markings in Muscovite, 242.—H. C. 
Lewis. The Optical Characters of some Micas, 244.—H. C. Lewis. On 
an exfoliated Talc, 252.—H. C. Lewis. Tin in North Carolina, 253.— 
H. C. Lewis. On Siderophyllite, a new Mineral, 254.—H. C. Lewis. 
On Sterlingite and Damourite, 256.—H. C. Lewis. Vanadium in Phila- 
delphia Rocks, 256.—H. C. Lewis. The Surface Geology of Philadelphia 
and Vicinity, 258.—T. D. Rand. On Randite, 274.—T. D. Rand. Some 
microscopic Enclosures in Mica, 276.—H. C. Lewis. On the Bryn-Mawr 
Gravel, 277.—H. C. Lewis. On some Enclosures in Mica, 278.—H. C. 
Lewis. On Dendrites, 278.—H. C. Lewis. On a Jurassic Sand, 279.— 
H. C. Lewis. The Minerals of Surry Co., N. C., 280.—T. D. Rand. On 
a, peculiar Stratification in Gneiss, 280.—H. C. Lewis. A new Locality 
for Lignite, 281.—H. C. Lewis. On Serpentine in Bucks Co., 281.— 
H. C. Lewis. The Iron Ores and Lignite of the Montgomery-Co. Valley, 
282.—H. C. Lewis. An Enclosure in Quartz, 292.—W. W. Jefferis. 
Menaccanite and Tale from Maryland, 292.—W. W. Jefferis. Sandstone 
in Labradorite, 292.—A. E. Foote. On a probable Pseudomorphism of 
Gumuite and Uranotite after Uraninite, 292.—H. C. Lewis. Ona new Fu- 
coidal Plant from the Trias, 293.—T. D. Rand. The northern Belt of Ser- 
pentine in Radnor Township, 295.—H.C. Lewis. The Trenton Gravel and 
its Relation to the Antiquity of Man, 296.—H. C. Lewis. Note on Phila- 
delphite, a new Mineral, 310.—R. Haines. Analysis of Philadelphite, 
310.—F. A. Genth, jun. The so-called Emery-ore from Chelsea, Bethel 
Township, Delaware Co., 311.—H. C. Lewis.—On Philadelphite, 313.— 
H. C. Lewis. A Potsdam-Sandstone Outcrop on the 8. Valley Hill of 
Chester Valley, 329.—G. A. Konig. Notes on Jarosite, 331.—J. Leidy. 
Bone Caves of Pennsylvania, 546.—A. Heilprin. On some new Lower- 
Eocene Mollusca from Clarke Co., Alabama, with some Points as to the 
stratigraphical Position of the Beds containing them, 364. 

Philadelphia. American Philosophical Society. Proceedings. Vol. 
xvii. Nos.105 & 106. 1880. 

C. A. Ashburner. On the Oil-sand of Bradford, M‘Kean County, 419. 
—C. E. Hall. Relations of the Crystalline Rocks of Pennsylvania to the 
Silurian Limestones, and the Hudson-River Age of the Hydromica Schists, 
435. 

Photographie Society of Great Britain. Journal and Transactions. 
NESS Wolves wNoycamlcod: 


: Vol. v. Nos. 1-7. 1880-81. 
Physical Society. Proceedings. Vol. ii. Part 4. 1880. 
WVoliive catty en loole 


Pisa. Societa toscana di Scienze Naturali. Atti. Processi verbali. 
Vol. ii. pp. 89-232. 1880-81. 


S. de Bosniaski. La formazione gessoso-solfifera e il secondo piano 


ADDITIONS TO THE LIBRARY. 27 


mediterraneo in Italia, 90.—G. Meneghini. Nuovi fossili delle Alpi 
apuane, 102.—M. Canavari. Di alcuni ammoniti del lias medio rac- 
colti nelle vicinanze diS. Antonio nel gruppo montano di Tivoli, 109.— 
D. Pantanelli. NRadiolarie nei calcari, 111.—A. Manzoni. Ciottoli im- 
prontati, 112.—C. de Stefani. Carte geologiche delle Calabrie meridio- 
nali, 113.—C. de Stefani. Il Tortoniano dell’ alta val di Tevere, 114.— 
C. de Stefani. I fossili di Dicomano in Toscana e della Porretta nel 
Bolognese, 115.—C.de Stefani. Natura dei battrilli, 116.—C. de Stefani. 
Il gault e la creta superiore nell’ Apennino settentrionale, 117.—C. de 
Stefani. Ordinamento cronologico dei terreni delle Alpi apuane, 118.— 
F, Castelli. Delfino fossile, 131.—C. de Stefani. Il miocene di Canipa- 
rola, 140.—C. de Stefani. Osservazioni ad alcune pubblicazioni geolo- 
giche del R. Comitato geologico italiano sulle Alpi apuane, 141.—C. 
de Stefani. Pieghe costituenti le Alpi apuane, 156.—A. D’Achiardi. 
Su di alcuni minerali della miniera del Frigido presso Massa nelle Alpi 
apuane, 171.—D. Zaccagna. In risposta alle osservazioni del De Stefani 
sopra alcune pubblicazioni geologiche del R. Comitato geologico italiano 
sulle Alpi apuane, 179.—B. Lotti. In risposta alle osservazioni del De 
Stefani sopra alcune pubblicazioni geologiche del RK. Comitato geologico 
italiano sulle Alpi apuane, 186.—G. Meneghini. Ammoniti del lias 
medio, 188.—C. de Stefani. Di nuovo sui lavori del Comitato geologico 
nelle Alpi apuane, 189.—M. Canavari. Alcuni nuovi Brachiopodi degli 
strati a Terebratula aspasia Meh. nell’ Appennino centrale, 197.—D. Pan- 
tanelli. Su alcune rocce della Montagnola senese, 197.—G. Meneghini. 
Nuovi trilobiti di Sardegna, 199.—C. de Stefani. Studi microlitologici 
pel paleozoico e pel trias delle Alpi apuane, 202.—C. de Stefani. Il 
macigno di Porretta ed i terreni corrispondenti, 206.—C. de Stefani. 
Origine degli strati pontici intorno al Mediterraneo, 209.—C. de Stefani. 
Sui terreni marini dell’ epoca postpliocenica, 212.—C. de Stefani. Le 
pieghe dell’ infralias nelle Alpi apuane, 216.—S. de Bosniaski. Una 
pianta fossile del Verrucano dei monti pisani, 219.—S. de Bosniaski. Le 
argille da stoviglie del Camerinese, 221.—S. de Bosniaski. L’eta geolo- 
gica dei monti della Tolfa, 222.—Forsyth Major. Sgualodon quaternarium, 
227.—Forsyth Major. Studii sugli avanzi pliocenici del genere Sus (Sus 
Strozzt Menegh.), 227. 


Pisa. Societa toscana di Scienze Naturali. Atti. Memorie. Vol. iv. 
fase. 2. 18380. 

M. Canavari. Sui fossili del lias inferiore nell’ Appennino centrale, 
141.—G. Grattarola e F.Sansoni. Studii chimici sulla heulandite e sulla 
stilbite di S. Pietro (Elba) etc., 173.—G. Grattarola. Beccarite, varieta 
di zircon di Ceylon, 177.—R. Lawley. Nuovi denti fossili di Motedanus 
rinvenuti ad Orciano pisano, 196.—G. Grattarola. Orizite e pseudona- 
trolite, due nuove specie del sott’ordine delle zeoliti, 226.—A. D’Achiardi. 
Coralli giurassici dell’ Italia settentrionale, 233.—F. Sansoni. Sulle 
zeoliti dell’ isola d’Elba, 311.—A. Manzoni. Echinodermi fossili plioce- 
nici, 8327.—G. Meneghini. TF ossili oolitici di Monte Pastello nella pro- 
vincia di Verona, 336. 


Plymouth. Devonshire Association for the Advancement of Science, 
Literature, and Art. Report and Transactions. Vol. xii. 
1880. 

W. A. E. Ussher. Physical Features of Devonshire, 251.—A. R. 
Hunt. Notes on the Submarine Geology of the English Channel off the 
Coast of South Devon, 291.—W. Pengelly. Notes on Boulders and 
Scratched Stones in South Devon, 304.—N. Worth. Recent Geological 
Discoveries in the Neighbourhood of Plymouth, 361.—W. Downes 


27 8 ADDITIONS TO THE LIBRARY. 


Blackdown, 420.—W. Pengelly. Notes on recent Notices of the Geology 
and Paleeontology of Devonshire, 591. 


Plymouth Institution and Devon and Cornwall Natural-History 
Society. Annual Report and Transactions. Vol. vii. Part 2. 
1880. | 

W. Sharman. Darwinism, 218.—F. J. Webb. Natural History of 

Coal, 293.—F. Brent. The Stone Implements, 295. 


Popular Science Review. New Series. Vol. iv. Nos. 15 & 16. 
1880. Purchased. 

P. H. Carpenter. Feather-Stars, recent and fossil, 193.—J. F. Blake. 
The Portland Building Stone, 205.—M. F. de Castro. On the Influence 
which a molecular Movement, due to Electricity, may have exerted in 
certain Geological Phenomena, namely the Metamorphism of Rocks and 
the Formation of Metalliferous Deposits, 230.—P..M. Duncan. On the 
Opinions of Voltaire and Laplace regarding Geology, 310.—J. Milne. A 
large Crater, 336. 


; . Vol.v. Nos. 17 &18. 1881. Purchased. 
J.S. Gardner. The Permanence of Continents, 117. 


Quekett Microscopical Club. Journal. Nos. 44-46. 1880-81. 


Ray Society. A Monograph of the Free and Semiparasitic Copepoda 
of the British Islands. By G. 8. Brady. Vol. ii. 8vo. 
1880. | 


-——. Monograph of the British Aphides. Vol. iii. By G. B. 
Buckton. 8vo. 1881. 


Riode Janeiro. Museu Nacional. Archivos. Vol.ii. Trimestres 
3 e4, 1878. 

O. A. Derby. A geologia da regiao diamantifera da Provincia do 
Parana no Brasil, 89.—O. A. Derby. <A bacia cretacea da bahia de Todos 
os Santos, 185.—R. Rathbun. Observacdes sobre a geologia aspecto da 
Ilha de Itaparica, na bahia de Todos os Santos, 159. 


Rome. Reale Accademia dei Lincei. Atti. Serie 3. Transunti. 
Wolves Haseaielss 0: 

Capellini e Meneghini. Sopra la memoria del Dott. D. Pantanelli, in- 
titolata: I diaspri della Toscana e i loro fossili, 198.—Capellini e Meneghini., 
Sulla memoria del Dott. M. Canavari, dal titolo: I Brachiopodi degli 
strati a Terebratula Aspasia, Mgh. nell’ Appennino centrale, 199.—Tara- 
melli e Meneghini. Sulla memoria del Dott. C. F. Parona, intitolata: Il 
calcare di liasico di Gozzano, e i suoi fossili, 201.—Cornalia e Capellini. 
Sulla nota del Dott. A. Incoronato avente per titolo: Sopra uno scheletro 
umano dell’ epoca della pietra della provincia di Roma, 201.—Ponzi. Sui 
lavori del Tevere e sulle variate condizioni del suolo romano, 203.—G. 
Terrenzi. II lias superiore nel versante orientale della catena montuosa 
Narnese, 209.—Struever. Sulla Perowskite di Val Malenco, 210.—A. 
Piccini. Analisi di un’ augite del Lazio, 224.—F. Mauro. Ricerche 
chimiche sulle lave di Montecompatzi, del Tusculo, di Villa Lancellotti e 
di Monte Pila, 226.—Cossa. Sulla diffusione dei metalli della Cerite, 
232.—Cossa. Sultungstato di didimio, 233.—Cossa, Sopra un granato 
verde di Val Malenco, 254. 


=. Vol. vi Fase) 122 iiseme 


R. Meli. Sulla natura geologica dei terreni incontrati nelle fondazioni 
tubulari del nuovo ponte in ferro costruito sul Tevere a Ripetta, e sull’ 


ADDITIONS TO THE LIBRARY. 279 


Unio sinuatus, Lamk. rinvenutovi, 21.—Cornalia. Due piccoli imenotteri 
fossili dell’ ambra siciliana, 80.—A. Cossa. Sulla stilbite del ghiacciaio 
del Miage (Monte Bianco), 86.—A. Cossa. Sulla Ollenite, roccia anfibo- 
lica del monte Ollen, 88.—Meneghini e Capellini. Sulla Memoria del 
Prof. Seguenza “ Lo stretto de Messina,” 113.—Capellini. Resti di Tapiro 
nella lignite di Sarzanello, 131.—Ponzi. Sui tufi vulcanici della Tuscia 
romana, a fine di togliere qualunque discordanza di opinione emessa sulla 
loro origine, diffusione, ed eta, 1382.—Pigorini, Cossa e Capellini. Sulla 
memoria del Dott. D. Lovisato “Cenni critici sulla preistoria calabrese,” 
195. 


Royal Agricultural Society of England. Journal. Second Series. 
Vol. xvi. Part 2. No. 32. 1880. 3 


Vol. xvii. Part 1. 1881. 


Royal Asiatic Society of Great Britain and Ireland. Journal. N.S. 
Vol. xii, Parts3 & 4. 1880. 


: Wolltoaire (Barts i 2) 1S3i: 
Royal Astronomical Society. Memoirs. Vol. xlv. 1879-80. 1880. 


Royal Geographical Society. Journal. Vol. xlix. (1879). 1880. 

R. F. Burton. Itineraries of the Second Khedivial Expedition: Me- 
moir explaining the new Map of Midian made by the Egyptian Staff- 
officers, 1—R. F. Burton. A Visit to Lissa and Pelagosa, 151.—R. C. 
Temple. An Account of the Country traversed by the Second Column 
of the Tal-Cho’tia’li Field Force in the Spring of 1879, 190.—G. Rol- 
leston. The Modifications of the external Aspects of Organic Nature 
produced by Man’s Interference, 320. 


——. Proceedings. Vol. ii. Nos. 7-12. 1883. 


; . N.S. Vol. ii. Nos. 1-6. 1881. 
W.-M. Crocker. Notes on Sarawak and Northern Borneo, 193.—J. 
Stewart. Lake Nyassa and the Water Route to the Lake Region of 


Africa, 257. | 
Royal Institution of Great Britain. Proceedings. Vol. ix. Part 3. 
No. 72. 1880. 


W. B. Carpenter. Sea and Land in relation to Geological Time, 268. 


Royal Microscopical Society. Journal. Vol. iii. Nos. 4—-6a. 1880. 


H. Stolterfoth. On the Diatomaces in the Llyn Arenig Bach Deposit, 
913. 


Second Series. Vol: i. Parts1-3. 1881. 


Ww. H. Shrubsole and F. Kitton. The Diatoms of the London Clay, 
381. 


Royal Society. Philosophical Transactions. Vol. clxxi. Parts 1-3. 
1880-81. 

W.C. Williamson. On the Organization of the fossil Plants of the 
Coal-measures, Part X., including an Examination of the supposed Ra- 
diolarians of the Carboniferous Rocks, 493.—G. H. Darwin. On the 
Secular Changes in the Elements of the Orbit of a Satellite revolving 
about a tidally-distorted Planet, 713.—R. Owen. Description of some 
Remains of the gigantic Land-lizard (Megalania prisca, Owen) from 
Australia, Part IT., 1037. 


——, ——. Vol. clxxi. Part1. 1881. 


280 ADDITIONS TO THE LIBRARY. 


Royal Society. Proceedings. Vol. xxx. No. 205. 1880. 

W.C. Williamson. On the Organisation of the fossil Plants of the 
Coal-measures, Part XJ., 550.—O. Heer. On the Miocene Plants disco- 
vered on the Mackenzie River, 560. 


: Vol. xxxi. Nos. 206-212. 1880-81. 

C. Schroter. Note on the Microscopic Examination of some fossil 
Wood from the Mackenzie River, 147.—J. W. Hulke. Polacanthus Foxit, 
a large undescribed Dinosaur from the Wealden Formation in the Isle of 
Wight, 336.—R. Owen. Description of some Remains of the Gigantic 
Land-lizard (Megalania prisca, Owen) from Australia, 380.—W. B. Pauli. 
Notes on the Earthquakes of Julv 1880 at Manila, 460.—S. Haughton. 
Notes on Physical Geology, No. 7. On the Secular Inequalities in Ter- 
restrial Climates depending on the Perihelion, Longitude, and Eccentricity 
of the Earth’s Orbit, 473. 

Rugby-School Natural-History Society. Report for the year 1880. 
1881. 

EK. H. Acton. Subdivisions of the Carboniferous Limestone in Den- 
bighshire, North Wales, 4.—A. Percy Smith. Holyhead, 20.—Seabroke. 
The ‘Model’ (of the neighbourhood), 24—H. Y. Oldham. On some 
Sections lately exposed near Rugby, 39.—T. K. Worthington. Report of 
the Geological Section, 48. 


St. Petersburg. Académie Imperiale des Sciences. Bulletin. 
Tome xxvi. Nos. 2 & 3. 1880. 


F, A. Forel. Les échantillons de limon dragués en 1879 dans les lac 
d’ Arménie, 571. 


: 4 Tome xxvii. Nos. 1&2. 1881. 
N. v. Kokscharow. Beryll-Krystalle eines neuen Fundortes, 35. 
: . Mémoires. Tome xxvil. Nos. 2-14. 1879-80. 
J. Schmalhausen. LBeitrage zur Jura-Flora Russlands, No. 4.—V. vy. 
Moller. Die Foraminiferen des russischen Kohlenkalks, No. 5.—L. v. 


Schrenck. Der erste Fund einer Leiche von Rhinoceros Merchi, Jag., No. 

7.—O. Heer. Nachtrage zur Jura-Flora Sibiriens, No. 10. 

——. ——. ——. Tome xxviii. Nos.1 & 2. 1880. 

Sanitary Institute of Great Britain. Transactions. Vol.i. 1880. 
G. F. Symons. Address to Section 3, 173.—A. Haviland. Geology 

in relation to Sanitary Science, 190.—J. Lucas. On the Quantitative 

Elements in Hydrogeology, 195.—W. B. Kinsey. Particulars of an Ar- 

tesian Well at Thames Haven, Essex, 203.—W. F. Stanley. Conditions 

of the Water Supply of Croydon in relation to its Rainfall and Geology, 

with Suggestions for its Sanitary and profitable Improvement, 225. 

Scientific Roll and Magazine of Systematized Notes. Nos. 1-3. 

1830-81. . 
Shanghai. North-China Branch of the Royal Asiatic Society. 
Journal. Vol.i. No.3. 1859. 

._——. By Oly ails, INO: Iho eG: 

W. H. Shock. Notes on the Mineral Resources of Japan, 92. 

NeS aiNo 9s Sijos 

—. 1. ——. ——. No. 12. 1878. 

—. #——. —. —. No. 13. 1879. 

—. «——. ——. —. No. 14. 1879. 


ADDITIONS TO THE LIBRARY. 281 


Sherborne. Dorset Natural-History and Antiquarian Field Club. 
Proceedings. Vol.in. 1879. 

J.C. Mansel-Pleydell. Anniversary Address of the President, 1.—A. 
U. Kent. The finding Yerebratula Morierei at Bradford Abbas, 39.—J. 
F,. Walker. The Occurrence of Terebratula Morieret in England, 42.—J. 
C. Mansel-Pleydell. On the Dorset Trigone, 111.—J. Buckman. On 
.a Series of Sinistral Gasteropods, 185.—J. Buckman. On the Belemno- 
teuthis Montefiore, 141. 
Society of Arts. Journal. .Vol. xxvii. Nos. 1440-146 

C. G. W. Lock. Pandermite: a new Boracic Mineral, 76 
: . Vol. xxix. Nos. 1461-1491. 1880-81. . 
B. Latham. Croydon Bourne Flow, 84.—Hyde Clarke. Goldin India, 
244.—Gold-mining in Japan, 286.—E. Whymper. Ascent of Chimborazo 
and Cotopaxi, 353.—James Tennant, 366.—Marble Quarries of Algeria, 


1880. 


y 
he 


Society of Biblical Archeology. ‘Transactions. Vol. vii. Part 1. 
1880. 


Society of Medical Officers of Health. Annual Report 1878-79. 
1879. Presented by W. Whitaker, Esq., F.GS. 


——. Transactions. Session 1880-81. 1880. Presented by W. 
Whataker, Esq., F.GS. 


Stuttgart. Neues Jahrbuch fiir Mineralogie, Geologie und Palion- 
tologie. 1879, Hefte 8 and 9. 1879. 

A. Losch. Ueber Kalkeisengranat (Demantoid) von Syssertsk am 
Ural, 785.— F. M. Stapff. Zur Mechanik der Schichtenfaltungen, 792.— 
L. van Werveke. Beitrag zur Kenntniss der Gesteine der Insel Palma, 
815.—F. Wohler. Bemerkungen itber das gronlandische Gediegen-Hisen, 
832. S.M. Babcock. Ueber den Colestin aus dem Muschelkalke von 
Juhbnde bei Gottingen, 835.—P. Herter. Ueber die Silicatgesteine im 
EKocangebirge yon Massa maritima, 839.—C.A.Tenne. Ueber Hpistilbit, 
840,—O. Schmidt. Bemerkungen zu “ Osc. Schmidt, Die Spongien des 
Meerbusens von Mexico,” 841.—A. Arzruni. Cerithiwm corallense (Bu- 
vignier) von Salatau, 842.—U. Stulz. Geologische Notizen aus den Alpen. 
Ueber das Erstfelder Thal, 842.—K. Martin. Ueber das Tertiar von Java, 
3800.—R. Richter. Diluvium bei Saalfeld, 850.—G. Berendt. Riesen- 
kessel auf dem Riidersdorfer Muschelkalk bei Berlin, 351.—C. Struckmann. 
Sowerbya Dukei in hannover’schen Pteroceras-Schichten, 853.—H. Fischer. 
Ueber die Bezeichnung yon Farbenabstufungen bei Mineralien, 854.—T. 
Fuchs. Ueber Solfataren in Serpentinstocken bei Kalamaki, 857.—E. 
Cohen. Kersantit von Laveline. 858.—C. W. Giimbel. Vulkanische 
Asche des Aetna von 1879, 859.—E. Cohen. Ueber einen Hklogit, 
welcher als Hinschluss in den Diamanteruben von Jagersfontein, Grange 
Freistaat, Std-Afrika vorkommt, 864. 


é ») 18380) Bandi, 1380. 

KE. Kalkowsky. Ueber die Erforschung der archaischen Formationem, 
1.—H. Kalkowsky. Ueber Gueiss und Granit des bojischen Gneisstock- 
werkes im Oberpfalzer Waldgebirge, 29.—C. A. Tenne. Ueber den Epi- 
stilbit, 43.—F. Klocke. Ueber Doppelbrechung regularer Crystalle, 
52.—F. Zirkel. Ueber den Zirkon als mikroskopischer Gesteins- 

emengtheil, 80.—E. Dathe. Gletscherschliffe bei Lommatzsch in 
achsen, 92.—A. Sauer. Titanmineralien in Amphiboliten, 94.—von 
Konen. Tertiar zwischen Guntershausen und Marburg, 95.—H. Cohen. 
Mandelstein aus den Maluti-Bergen, 96.—A. Inostranzeff. Ein neues 


282 ADDITIONS TO THE LIBRARY. 


ausserstes Glied in der Reihe der amorphen Kohlenstoffe, 97.—C. Struck- 
mann. Vorlaufige Nachricht uber das Vorkommen erosser vogelahnlicher 
Thierfahrten (Ornithoidichnites) im Hastingssandsteine von Bad Rehburg 
bei Hannover, 125.—G. Seligmann. Krystallographische Notizen L, 129. 
A. Weisbach. Zur Keuntniss des Leucits, 143.—G. Steinmann. Die Mu- 
mien des Hauptrogensteins, 151—C. Klein. Ueber Goldstufen von 
Vorospatak, 155.—A. Cossa. Rutil in Gastaldit-Eklogit von Val Tour- . 
nanche, 162.—-A. Kenngott. Ueber Topas, Pyrrhotin und Pseudobrookit, 
164.—H. Kayser. Ueber Barrande’s Brachiopodenwerk, 166.—A. Pichler. 
Beitrage zur Geognosie Tirols, 172.—H. Fischer. Ueber Jadeit, 174.—F. 
Schalch. Die Gliederung der Liasformation des Donau-Rheinzugs, 177.— 
Des Cloiseaux. Ueber Pyrenien-Mineralien, 267.—T. Wolf. Geolovische 
Arbeiten im Staate Ecuador, 268.—O. Fraas. Glaciales, 268.—Val. von 
Moller. Schadel von Elasmotherium Fischert, Desm., 273.—G. vom Rath. 
Contactverhaltnisse zwischen Kohle und einem basischen Eruptivgestein 
bei Funfkirchen, 274.—A. Kenngott. Ueber Barytplagioklas, 278.—A. 
Sauer. Rutil als mikroskopischer Gesteinsgemengtheil, 279.—-C. Klein. 
Erwiderung, 281.—A. Koch. Ueber das Tertiar in Siebenbiirgen, 283.— 
C. A. Tenne. Ueber ein neues Vorkommen von Epistilbit, 285.—C. 
Klein. Berichtigung, 286.—E. Hussak. Eruptivgesteine von Schemnitz, 
Augitandesit von St. Heidi, 287. 


Stuttgart. Neues Jahrbuch fir Mineralogie, Geologie und Palion- 
tologie. 1880, Bandu. Hefte 1-3. 1880. 

C. E. Weiss. Die Krystallisationsgesetze seit C.S. Weiss, insbesondere 
die Lehre von den Hemiédrieen, erlautert am Diamant, 1.—E. Cohen. 
Ueber Laven von Hawaii und einigen anderen Inseln des grossen Oceans 
nebst einigen Bemerkungen iiber glasige Gesteine im Allgemeinen, 23.— 
M. Bauer. Beitrage zur Mineralogie. I. Reihe, 63.—A. Streng. Ueber 
die Hinschlusse von Pflanzenresten in dem Hisensteinslager am Dunst- 
berge bei Giessen, 83.—H. R. Goppert. Ueber die versteinerten Holzer 
des Kytffhauser, 89.—A. Nehring. Ueber Fossilreste von Lemmingen, 
93.—Herbich. Geologisches aus Bosnien und der Hercegowina, 94.— 
H. H. Reusch. Die Alaunschieferscholle von Bikkelaget bei Christiania, 
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felsauren Blei, 97.—O. Lang. Giebt es Gletscherspuren im Harz ?, 99.— 
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A. Streng. Erklirung, 107.—A. Weisbach. Mineralovische Notizen L., 
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Kenntniss fossiler Kalkalgen (Siphoneen), 130.—L. van Werveke. Ueber 
den Nephelin-Syenit der Serra de Monchique im stidlichen Portugal und 
die denselben durchsetzenden Gesteine, 141.—W. OC. Brogger. Ueber 
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Neu-Seeland, 192.—A. Baltzer. Ueber Bergstiirze, 197.—F. Sandberger. 
Sycidien aus dem Devon am Sjass, 199.—O. Liidecke. Ueber Skolezit, 
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und Geologie, 203.—V. Goldschmidt. Unterscheidung von Arsenkies 
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ADDITIONS TO THE LIBRARY. 283 


schieferscholle yon Biakkelaget, 290.—A. Pichler. Aus Tirol, 292.—O. 

Miigge. Glimmerporphyrit vom Steinacher Joch, 293.—A. Koch. Ueber 

siebenbiirgisches Tertiaér, 294.—A. Wichmann. Turmalin als authigener 

Gemengtheil von Sanden, 294.—A. Nehring. Neue Notizen iiber fossile 

Lemminge. Jin Losslager bei Mariaspring unweit Gottingen, 297. 

Stuttgart. Neues Jahrbuch fir Mineralogie, Geologie und Palion- 
tologie. 1881, Bandi. Heft 1. 1881. 

G. Werner. Mineralogische Mittheilungen, 1—F. Klocke. Ueber die 
optische Structur des Gletschereises, 23.—G. Steinmann. Die Forami- 
niferengattung Nummoloculina, n. g., 31.—H. von thering. Die Aptychen 
als Beweismittel fiir die Dibranchiaten-Natur der Ammoniten, 44.—E. 
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mikroskopischen Structur von Mineralien und Gesteinen, 93.—P. Merian. 
Zur Gattung Graphularia, M.-Edw. & Haime, 96.—R. D. M. Verbeek. 
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: Beilage-Band i. Heft 1. 1880. 

F. Maurer. Palaontologische Studien im Gebiet des rheinischen Devon. 
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——. Verein fir vaterlindische Naturkunde in Wurttemberg. 
Jahreshefte. Jahrgang 26. 1880. 

C. Dorn. Anwendung der gelegentlich der Tiibinger Wasserversor- 
gung gewonnenen Hrfahrungen fir die Wasserversorgung von Stuttgart, 
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spathe im Basalttuff des Owener Boller, 74. 


Swansea. South-Wales Institute of Engineers. Proceedings. 
Vol. xu. Nos. 1-4. 1880. 
H. Cossham. Some of the geological Problems in the Bristol Coalfield, 
84.—T. Joseph. Observations on the Clayband-Ironstone, or “ Mine,” of 
the Northern Outcrop of the South-Wales Coal-field, 255. 


Sydney. Linnean Society of New South Wales. Proceedings. 
Vol. iv. Part 4. 1880. 


C. Jenkins. On the Geology of Yass Plains, 404.—C. 8. Wilkinson. 
Notes on the Abercrombie Caves, 460. 


: : Oven Area e OOO: 
J. EH. Tenison-Wood. On a Fossiliferous Bed at the Mouth of the 
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Royal Society of New South Wales. Journal. Vol. xii. 
13879. 1880. 
J. Hector. On the Geological Formations of New Zealand compared 
with those of Australia, 65.—C.S. Wilkinson. Notes on the Occurrence 
of remarkable Boulders in the Hawkesbury Rocks, 105. 


Taunton. Somersetshire Archeological and Natural-History Society. 
Proceedings. Vol. xix. (1873). 1874. Presented by H. B. 
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H. B. Woodward. Notes on the Geology of the Neighbourhood of 

Wells, Somerset, 50. | 


284 ADDITIONS TO THE LIBRARY. 


Tokio. University of. Memoirs of the Science Department. Vol. 1. 
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EH. 8. Morse. Shell-mounds of Omori, 1. 


: : Noli 2539) (379): 
C. Netto. Mining and Mines in Japan, 1. 


—_—_——— 


. The Calendar of the Departments of Law, Science, 
and Literature. 2539-40 [1879-80]. 


Troms6 Museums Aarshefter. Heft ii. 1880. 
Karl Pettersen. Terrasser og gamle strandlinjer, 1—R. Collett. 


Glaciale Mergelboller med indesluttede Fiskelevninger fra Bejeren 1 
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Turin. Osservatorio della Regia Universita. Bollettino. Anno xiy. 


(1879). 1880. 


—. —. —. Anno xy. (1880). 1881. 
——. Reale Accademia delle Scienze. Atti. Vol. xv. Disp. 1-8. 
1879-80. 


A.Cossa. Sui cristalli microscopici di rutilo contenuti in una eclogite 
di Val Tournanche, 21.—A. Portis. Intorno ad alcune impronte eoceniche 
di vertebrati recentemente scoperte in Piemonte, 221.—G. Berruti e L. 
Bellardi. Relazione sopra una memoria del Professore Martino Baretti, 
intilolata: Il ghiacciaio del Miage, 299.—M. Baretti. Sui resti fossili 
di Rinoceronte nel territorio di Dusino, circondario d’Asti, provincia 
d’ Alessandria, 678, 751.—G. Spezia. Nota sul calcare albitifero dell’ Ar- 
gentera (Cuneo), 785. 


: 2 Vol. xvi. Disp. 1-5. 1881. 

A. Cossa. Nota su alcune roccie sepentinose del Gottardo, 71.—G. 
Berruti e M. Lessona. Relazione sulla memoria presentata dal Dottore 
Alessandro Portis col titolo: Sui terreni stratificati di Argentera (Valle 
di Stura), 199.—A. Cossa. Nota sopra alcune roccie serpentinose 
dell’ Appennino bobbiese, 296.—A. Cossa e E. Mattirolo. Sopra alcune 
roccie del periodo silurico nel territorio d’ Iglesias, Sardegna, 385.— 
G. Piolti. Nota sopra alcune pietre a sccdelle dell Anfiteatro morenico 
di Rivoli, Piemonte, 403. 


University College. Calendar (Session 1880-81). 1880. 


Victoria Institute. Journal of the Transactions. Vol. xiv. Nos. 
A= jon lesOLell. 


T. McK. Hughes. On the Evidence of the later Movements of Eleva- 
tion and Depression in the British Isles, 248. 


eS )6OWOL aa, INO, SSB IL 


Vienna. Archiv fiir practische Geologie. Bandi. 13880. 

F. Posepny. Die Goldbergbaue der Hohen Tauern mit besonderer 
Beriicksichtigung des Rauriser Goldberges, 1—F’. PoSepny. Die Erzla- 
gerstatten von Kitzbiihel in Tirol und der angrenzenden Theile Salzburgs, 
257.—F. PoSepny. Die Erzlagerstatten am Pfundererberge bei Klausen 
in Tirol, 441.—H. Hofer. Die Edelmetall-Production Karntens, 490.— 
F. Pogepny. Ueber den alten Bergbau in Tirol, 520.—F. Posepny. Geo- 
logie und Bergbau in ihren gegenseitigen Beziehungen, 531. 


ADDITIONS TO THE LIBRARY. 285 


Vienna. Beitraige zur Paliontologie von Oseterreich-Ungarn und den 
angrenzenden Gebieten. Bandi. MHefte 1 & 2. 1880-81. 
Purchased. 

H. Zugmayer. Untersuchungen tiber rhatische Brachiopoden, 1.—A. 
Bittner. Beitrage zur Kenntniss alttertiarer Hchinidenfaunen der Siid- 
alpen, 43, 73.—V. Uhlig. Die Jurabildungen in der Umgebung von 
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——. Kaiserliche Akademie der Wissenschaften. Anzeiger. 1880, 
Nos. 11-16. 1880. 
F. Toula. Geologische Untersuchung des westlichen Balkan, 86.— 
J. Wold¥ich. Diluvialfauna von Zuzlawitz bei Winterbere im Bohmer- 
walde, 89. 


——, ——. ——. 1880, Nos. 20-28. 1880. 
J.C. Labhart. Ueber die im Monate Juli d. J. [in Manila] stattge- 
funden Erdbeben, 195. 


: . ——. 1881, Nos. 7-13. 1881. 
F. Toula. Grundlinien der Geologie des westlichen Balkan, 62.—A. 
Brezina. Ueber die Meteoreisen von Bolson de Mapimi, 103. 


: Denkschriften. Band xl. 1880. 

A. Bittner. Der geologische Bau von Attika, Bootien, Lokris, und 
Parnassis, 1—M. Neumayr. Der geologische Bau des westlichen Mittel- 
Griechenland, 91.—Fr. Teller. Der geologische Bau der Insel Eubéa, 
129.— Fr. Teller. Geologische Beschreibung der studoéstlichen Thessalien, 
183.—M. Neumayr. Ueber den geologischen Bau der Insel Kos und 
tiber die Gliederung der jungtertiaren Binnenablagerungen des Archipels, 
213.—M. Neumayr. Geologische Beobachtungen im Gebiete des thes- 
salischen Olymp, 315.—L. Biurgerstein. Geologische Untersuchungen 
im sudwestlichen Theile der Halbinsel Chalkidike, 321.—M. Neumayr. 
Geologische Untersuchungen uber den nordlichen und ostlichen Theil der 
Halbinsel Chalkidike, 328.—F’. Teller. Geologische Beobachtungen auf 
der Insel Chios, 340.—F’. Calvert und M. Neumayr. Die jungeren Abla- 
gerungen am Hellespont, 857.—A. Bittner, M. Neumayr, und Fr. Teller. 
Ueberblick tiber die geologischen Verhaltnisse eines Theiles der agaischen 
Kustenlander, 379. 


: . ——. Band xli. 1879. 

T. Fuchs. Ueber die von Dr. E. Tietze aus Persien mitgebrachten 
Tertiarversteinerungen, 99.—L. Szajnocha. Die Brachiopodon-Fauna 
der Oolithe von Balin bei Krakau, 197. 


; : Band xli. 1880. 

Hofer. Die Erdbeben Karntens und deren Stosslinien, 1—Hornes. 
Materialien zu einer Monographie der Gattung Megalodus mit besonderer 
Beriicksichtigung der mesozoischen Formen. 91.—Manzoni. Kchino- 
dermi fossili della molassa serpentinosa e supplemento agli Echinodermi 
fossili dello schlier delle colline di Bologna, 185. 


. Sitzungsberichte. Band lxxix. Hefte 1-5. 1879. 

J. Niedzwiedzki. Geologische Untersuchungen im westlichen Theile 
des Balkans und in den angrenzenden Gebieten zur Kenntniss der 
Eruptivgesteine des westlichen Balkans, 139.—V.v. Zepharovich. Halo- 
trichit und Melanterit von Idria, 183,—A. Boué. Ueber die Oro-Potamo- 
Limne- (Seen) und Lekanegraphie (Becken) des Tertiaren der europaischer 
Tirkei und Winke zur Ausfullung der Liicken unserer jetzigen geogra- 
phischen und geognostischen Kenntnisse dieser Halbinsel, 26].—Liebe. 


286 ADDITIONS TO THE LIBRARY. 


Die fossile Fauna der Hohle Vypustek in Mihren nebst Bemerkungen 
betreffs einiger Knochenreste aus der Kreuzberghohle in Krain, 472. 


Vienna. Kaiserliche Akademie der Wissenschaften. Sitzungs- 
berichte. Band Ixxx. Hefte 1-5. 1880. 

J. V. Janovsky. Ueber Niobit und ein neues Titanat vom Isergebirge, 
34,—J. Rumpf. Ueber den Krystallbau des Apophyllits, 84.—F. Becke. 
Ueber die Zwillingsbildung und die optischen Kigenschaften des Chabasit, 
90.—F. Berwerth. Ueber Nephrit aus Neu-Seeland, 102.—F. Berwerth. 
Ueber Bowenit aus Neu-Seeland, 116.—M. Slabnetier. Ueber die optische 
Orientirung der Plagioklase, 192. —V. Uhlig. Ueber die liasische Bra- 
chiopodenfauna yon Sospirolo. bei Belluno, 259.—F. Hochstetter. Ergeb- 
nisse der Hl plectiosel anita sien : im Jahre 1879, zweiter Bericht, 526. 


ao SS Sin Io, Lele IES, S80, 

T. Fuchs. Weber einige tertiaire Hchiniden aus og 97.—A. Boué, 
Ueber den ehemaligen und jetzigen Stand der Geologie und Geogenie 
und die Untersuchung en und Methoden in diesen Richtungen, 148.—Leo 
Birgerstein und Franz Noé. Geologische Beobachtungen im sudlichen 
Calabrien, 154.—F. Toula. Geologische Untersuchungen im westlichen 
Theile des Balkan und in den angrenzenden Gebieten (1X.), 188. 


Band Ixxxi. Hefte 1 & 2. 1880. 

J. N. Woldfich. Diluviale Fauna von Zuzlawitz bei Winterberg im 
Bohmerwalde, 7.—J. Sieber. Zur Kenntniss der nordbéhmischen Braun- 
kohlenflora, 67.—V. Bieber. Ueber zwei neue Batrachier der bohmischen 
_Braunkohlenformation, 102.—C. v. Ettingshausen. Beitrage zur Erfor- 
schung der Phylogenie der Pflanzenarten, 133.—G. Tschermak und L. 
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zur Kenntuiss der Er uptivgesteine der U mgegend von Schemnitz, 164. 


——. Kaiserlich-konigliche Bergakademien zu Leoben und Pribram 
und die konighch ungarischen Bergakademie zu Schemnitz. 
Berg- und huttenminnisches Jahrbuch. Band xxviii. Hefte 
2-4, 1880. 

K. Reyer. Zinn in Cornwall, 153. 


——. ——. ——. Band xxix. Heft 1. 1881. 

HK. Reyer. Ueber die “‘Granitstocke,” 15.—H. Reyer. Was versteht 
der Bergmann unter dem Worte “ Stock” 2, 20.—M. v. Lill, Analysen 
ausgefiihrt im Laboratorium des k. k. General-Probiramtes in Wien im 
Jahre 1880, 27, 

Kaiserlich-konigliche geologische Reichsanstalt. Abhand- 
lungen. Band xii. Heft 2. 1880. 

R. Hornes und M. Auinger, Die Gasteropoden der Meeresablage- 
rungen der ersten und zweiten miocinen Mediterranstufe in der dster- 
reichisch-ungarischen Monarchie, I1., 53. 


Jahrbuch. Band xxx. Hefte3 &4. 1880. 

F. vy. Hauer. Grundlinien der Geologie von Bosnien-Hercegovina 
(Vorwort), 160.—E. v. Mojsisovics. Grundlinien der Geologie von 
Bosnien-Hercegovina. I. West-Bosnien und Turkisch-Croatien, 167.— 
KE. Tietze. Grundlinien der Geologie von Bosnien-Hercegovina. II. 
Das ostliche Bosnien, 266.—A. Bittner. Grundlinien der Geologie von 
Bosnien-Hercegovina. III. Die Hercegovina und die sudostlichsten 
Theile von Bosnien, 353.—C. v. John. Grundlinien der Geologie von 
Bosnien- Hercegovina. IV. Ueber krystallinische Gesteine Bosnien’s und 
der Hercegovina, 439.—M. Neumayr. Grundlinien der Geologie von 
Bosnien-Hercegovina. V. Tertiire Binnenmollusken aus Bosnien und 


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der Hercegovina, 465.—M. Vacek. Neocomstudie, 493,—I. Reyer. Die 
Bewegung im Festen, 543.—KH. Kayser. Zur hercynischen Frage, 556.— 
D. Kramberger. Die fossilen Fische von Wurzenege bei Prassberg in 
Steiermark, 565.—V.v. Moller. Ueber einige Foraminiferen-fiihrenden 
Gesteine Persien’s, 573.—M. Lomnicki. Die galizisch-podolische Hoch- 
ebene zwischen dem oberen Laufe der Fliisse Gnila, Lipa und Strypa, 587. 
—R. Scharizer. Mineralogische Beobachtungen, 593.—A. Cathrein. Die 
Dolomitzone bei Brixlege in Nordtirol, 609.—H. Walter. Ein Durch- 
schnitt in dem Mittelkarpathen von Chyréw tiber Uherce und den un- 
garischen Grenzkamm bis Sturzica, mit Beriicksichtigung eniger Paral- 
lel-durchschnitte, 635.—R. Hornes. Die Trilobiten-Gattungen Phacops 
und Dalmamnites und ihr vermuthlicher genetischer Zusammenhang, 651. 
—K.M. Paul. Ueber die Lagerungsverhiltnisse in Wieliczka, 687.— 
HE. von Mojsisovics. Ueber heteropische Verhiltnisse im Triasgebiete 
der lombardischen Alpen, 695.—J. Schmidt. Ueber die Fossilien des 
Vincaberges bei Karlstadt in Croatien, 719.—E. Tietze. Zur Geologie der 
Karsterscheinungen, 729. 


Vienna. Kaiserlich-konigliche geologische Reichsanstalt. Jahr- 
bouche Band xxxi. Nov. 13881. 

HE. Reyer. Predazzo, 1.—H. Reyer. Ueber Tuffe und tuffogene Sedi- 
mente, 5/.—E. Tietze. Ueber einige Bildungen der jiingeren Epochen 
in Nord-Persien, 67.—C. M. Paul. Die Petroleum- und Ozokerit-Vor- 
kommnisse Galiziens, 131. 


. ——. Verhandlungen. 1880, Nos. 10 and 11. 1880. 

A. Heim. Ueber die Glarner Doppel-Falte, 156.—R. Hornes. Mas- 
todon angustidens von Oberdorf nordlich von Weiz, 159.—V. Bieber. 
Ueber zwei neue Batrachier aus dem Diatomaceenschiefer bei Sulloditz 
in Bohmen, 160.—E. Hussak. Ueber Eruptivgesteine von Gleichenberg, 
160.—T. Fuchs. Ueber ein neues Vorkommen von Stisswasserkalk bei 
Czeikowitz in Mahren, 162.--R. Hornes. Tertiar bei Derwent in 
Bosnien, 164.—E. Suess. Ueber die vermeintlichen saécularen Schwan- 
kungen einzelner Theile der Erdoberflache, 171.—C. von Hauer. Krys- 
tallogenetische Beobachtungen, 181.—M. Vacek. Hrwiederung auf die 
Mittheilung des Herrn Prof. A. Heim in Nr. 10 der Verhandlungen, 189.— 
R. Hornes. Die Stosslinie des Villacher Erdbebens von 1348, 193.—R. 
Hornes. Amphiope nov. sp. vom Seckauer Berg bei Leibnitz, 194. 


é ——. 1880, Nos. 13-17. 1880. 

Karl Ritter von Hauer, 229.—H. Reyer. Ueber die Tektonik der gra- 
nitischen Gesteine von Predazzo, 231.—A. Bittner. Die Sedimentgebilde 
in Judicarien, 253.—V. Hilber. Reisebericht aus Ostgalizien, 238.— 
R. Hornes. Das Auftreten der Gattung Terebra in den Ablagerungen 
der ersten und zweiten miocainen Mediterran-Stufe der 6sterreichisch- 
ungarischen Monarchie, 245.—W. Jicinsky. Basalt in der Jaklowetzer 
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tallinische Centralmasse und die palaolithischen Randzonen der Alpen 
vom Gailthaler Gebirge uber das Tauernkreuzjoch nach dem Innthal bei 
Woregl, 249.—G. Stache. Der krystallinische Gebirgs-Abschnitt zwischen 
dem hinteren Ulten-Gebiet und Unter-Sulzberg, 250.—G. Stache. Aus 
den Randgebieten des Adamellogebirges, 252.—E. Tietze. Die Gegend 
von Rospucie in Galizien, 255.—F. Teller. Verbreitung und Lagerung 
der Diorite in der Umgebung von Klausen und Liisen, 261.—V. Hilber. 
Reiseberichte aus Ostgalizien, 264.—R. Hornes. Das Erdbeben vom 
9. November in Steiermark, 269.—M. Lomnicki. LEiniges tiber die Gyps- 
formation in Ostgalizien, 272.—V. Uhlig. Zur Gliederung des rothen 
Ammonitenkalkes in der Umgebung von Roveredo, 275.—C. W. Gumbel. 


288 ADDITIONS TO THE LIBRARY. 


Rothkalk. Magnesit von Elmen, 276.—G. C. Laube. Pflanzenreste aus 
dem Diatemaceenschiefer in Sulloditz im bohm. Mitteleebirge, 277.— 
G. Starke. Notizen uber Bol und Polyhydrit, 278.—KH. Tietze. Zur 
Geologie der Karsterscheinungen, 281.—F’. y. Hauer. Bouteillenstein von 
Trebitsch, 282.—J. N. Woldrich. Beitrage zur diluvialen Fauna der 
mihrischen Hohlen, 284.—G. Stache. Ueber das Vorkommen von 
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suches tiber Bewegung im Festen, 288.—D. Kramberger. Die jung- 
tertiare Fischfauna Croatiens, 297.—A. Rzehak. Ueber die Gliederung 
und Verbreitung der alteren Mediterranstufe in der Umgebung von Gr. 
Seelowitz in Mahren, 300.—F. Teller. Vorlage des Blattes Klausen, 303. 
—E. Reyer. Ueber Predazzo, 304.—L. Szajnocha. Vorlage der geolo- - 
gischen Karte der Gegend von Gorlice, 304.—A. Biezina. Ueber ein 
neues Mineral, den Schneebergit, 313.—K. Hussak. Umeeschmolzene 
Basalte und Granite von Hdersertin bei Karlsbad, 314.—J. Kusta. Zur 
Geologie und Palaontologie des Rakonitzer Steinkohlen-Beckens, 317.— 
G. Sebisanovic. Hiniges tiber die Erdbeben von Karlstadt in Kroatien, 
325.—R. Hornes. Vorlage emer geologischen (Manuscript-) Karte der 
Umgebung von Graz, 326.—H. von Mojsisovics. Ueber heteropische 
Verhaltnisse im Triasgebiete der lombardischen Alpen, 330.—C. M. Paul. 
Geologische Karte der Gegend von Przemysl, 380. 


Vienna. Kaiserlich-kénigliche geologische Reichsanstalt. Verhand- 
lungen. 1881, Nos. 1-11. 1881. 

Fr. Ritter v. Hauer. Jahresbericht, 1—M. v. Hantken. Arbeiten 
der k. ungarischen geologischen Anstalt, 15.—v. Loeffelholz. Hinige 
geognostische Notizen aus Bosnien, 23.—A. Bittner. Bemerkungen zu 
voranstehender Mittheilung, 27.—F. Kreutz. Ueber den Ursprung des 
Erdols in der galizischen Salzformation, 28.—EH. Tietze. Das Alter des 
Kalkes von Steinbergen bei Graz, 34.—G. Stache. Ueber die Gesteine 
des Adamello-Gebirges, 37.—EH. Tietze. Ueber die geologische Aufnahme 
der Gegend von Lemberg und Grodek, insbesondere tiber den Loss dieser 
Gegend, 37.—M. Vatek. Ueber die Schichtenfolge in der Gegend der 
Glarner Doppelfalte, 48.—V. Uhlig. Zur Kenntniss der Malm- und 
Tithonstufe in der Umgebung von Steierdorf im Banat, 51.—A. Bittner. 
Mittheilungen aus dem Aufnahmsterrain, 52.—D. Stur. Gebirgshub 
und Gebirgsschub, 58.—EH. Tietze. Bemerkungen zu den Ansichten 
von F, Kreutz uber das Erdol der galizischen Salzformation, 59,— 
E. v. Mojsisovics. Zur Karstgeologie, 65.—S. Kontkiewicz. Kurzer 
Bericht uber die von ihm ausgefuhrten geologischen Untersuchungen 
im stidwestlichen Theile vom Konigreich Polen, 66.—F. Teller. Zur 
Tektonik der Brixener Granitmasse und ihrer nordlichen Umrandung, 
69.—E. Reyer. Ueber die Tuffe der massigen Euptivgesteine, 74.—T. 
Fuchs. Chalicotheriwm von Siebenhirten, 77.—-A. Rzehak. Die Fauna 
des mihrischen Rothliegenden, 78.—C. Dolter. Von den capverdischen 
Inseln, 79.—v. Lorenz. Ueber terra rossa, 81.—H. von Dunikowski. 
Geologische Verhaltnisse der Dniesterufer in Podolien, 82.—H. Reyer. 
Ueber Predazzo, 85.—v. Loffelholz. Hin Beitrag zur Feststellung des 
Alters der Lossbildung bei Wien, 89.—J. Wentzel. Fossile Pflanzen aus 
den Basalttuffen von Warnsdorf in Bohmen, 90.—G. Laube. Neue 
Knochenfiinde aus dem Lehm der Umgebung von Prag, 93.—C. M. Paul. 
Ueber Petroleumvorkommnisse in der nordlichen Walachei, 93.—V. 
Hilber. Vorlage geologischer Karten aus Ost-galizien, 95.—F. Kreutz. 
Erklarung zu Dr. Tietze’s ‘‘ Bemerkungen zu den Ansichten von F. Kreutz 
uber das Hrdol der galizischen Salzformation, 101.—E. Kittl. Ueber einen 
neuen Fund von Listrzodon, 103.—H. v. Mojsisovics. Ueber die Cephalo- 
podon-Fauna der Trias-Schichten von Mora d’Ebro in Spanien, 105,— 


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K. M. Paul. Ueber das Ozokerit- und Erdél-vorkommen von Boryslaw, 
107.—F. Kreutz. Ueber die Bildung und Umbildung von E rdwachs und 
Erdél in Galizien, 115.—F. Puen: “Ucher den Ursprung des Steinsalzes 
am Rande der Karpathen, a. Pseudometeorit, gefunden 
in Cista, Pilsener Kreis, A. H. Schindler. Neue Angaben 
liber die Mineralreichthiimer Bees und Notizen tiber die Gegend 
westlich von Zendjan, 122 2.—V. Hilber. Die Stellung des oste: alizischen 
Gypses und sein V erhiltniss zum Schlier, 123.—H. y. Foullon. Krys- 
tallogenetische Beobachtungen, 131.—H. Kittl. Ueber die Mineralquellen 
Nordbéhmens, 149.—F. Wurm. Limonitconecretionen in der Umgebung 
von Béhmisch-Leipa, 153.—H. Engelhardt. Dritter Beitrag zur icennt. 
niss der Flora des Thones von Preschen bei Bilin, 154.—D. ‘Kramberg er. 
Studien tiber die Gattung Sawocephalus, 155.—W. Dames. Ueber “die 
Cephalopoden aus dem Gaultquader des Hoppelberges, 155.—C. Dolter. 
Spuren eines alten Festlandes auf den capverdischen Inseln, 156.—M. 
Vacek. Vorlage der geologischen Karte der Umgebung von Trient, 157. 
—L. Szaynocha. Das “Petroleumvorkommen von Sloboda rungur ska, 162. 


Vienna. Kaiserlich-kénigliche zoologisch-botanische Gesellschaft. 
Verhandlungen, 1880. Band xxx. 1881. 


—. Mineralogische und petrographische Mittheilungen. Neue 
Folge. Bandi. Hefte 1-6. 1880-81. Purchased. 

Curt von Eckenbrecher. Untersuchungen tber Umwandlungsvorgange 
in Nephelingesteinen, 1.—C. W. C. Fuchs. Die vulkanischen Hreignisse 
des Jahres 1879,35.—J. Becke. Hypersthen von Bodenmais, 60.—A. 
Penck. Die pyroxenfiihrenden Gesteine des nordsachsischen Porphyrge- 
bietes, 71.—P. Jannasch und J. H. Kloos. Mittheilungen tiber die laystal- 
linischen Gesteine des Columbia Flusse in Nordamerika und die darin 
enthaltenen Feldspathe, 97—Max Schuster. Ueber die optische Ori- 
entirune der Plagioklase, 117.—A. Frenzl. Mineralogisches aus dem 
ostindischen Archipel, 289.—F. Becke. Ueber den Hessit (Tellursil- 
berglanz) von Botes in Siebenburgen, 301.—G. Tschermak. Der Boden 
und die Quellen von Slanik, A Ja Pohlig. Die een ce 
im Siebenbirger Trachyt von der Perlenkardt bei ae <0. W. 
Cross. Studien iiber bretonische Gesteine, 369.—G. ‘Stein Die 
Melaphyre der kleinen Karpathen, 411.—H. Dietrich. ee Unter- 
suchung der drei neuen Mineralquellen von Krynica, 439.—J. Blaas. 
Petrographische Studien an jiingeren Eruptivgesteinen Persiens, 457.— 
A. Frenzel. Mineralogisches, 504.—A. v. Lasaulx. Ueber sogenannten 
kosmischen Staub, 517.—K. Geinitz. Der Phyllit von Rimognes in den 
Ardennen, 533. 


Washington. Smithsonian Institution. Annual Report of the Board 
of Regents for 1878. 1879. 


——, ——. Annual Report of the Board of Regents for 1879. 


1880. 
W. Gesner. Mica Beds in Alabama, 382. 
——. ——. Contributions to Knowledge. Vol. xxii. 1880. 
-——, ——. Miscellaneous Collections. Vol. xvi. 1880. 
——. ——. Vol. xvi. 1880. 


Watford Natural-History Society and Hertfordshire Field-Club. 
Transactions. Vol. ii. Part 8. 1880. 
VOL. XXXVII. y 


290 ADDITIONS TO THE LIBRARY. 


Watford. Hertfordshire Natural-History Society and Field-Club. 
Transactions. Vol.i. Parts 1-4. 1880-81. 

J. Gwyn Jeffreys. Anniversary Address, 84.—H. Gilbertson. Notes . 
on Sponges, recent and fossil, 97.—J. V. Elsden. The Post-Tertiary 
Deposits of Hertfordshire, 1053. r 
Wellington, N. Z. New-Zealand Institute. Transactions and Pro- 

ceedings. Vol. xii. (1879). 1880. 

W. Colenso. On the Moa, 63.—J. C. Crawford. On Wind-formed 
Lakes, 415.—J.C. Crawford. On Bidwill’s Front Hills, 416.—W. Collie. 
Remarks on Volcanoes and Geysers of New Zealand, 418. 

Wiesbaden. Nassauische Vereins fiir Naturkunde. Jahrbiicher. 
Jahrgang 31 u. 32 (1878 u. 1879). 1880. 

R. Fresenius. Chemische Analyse der Mineral-Quelle bei Biskirchen 
im Lahnthale, 1—R. Fresenius. Analyse der Wappen-Quelle zu Bad 
Ems, 17.—R. Fresenius. Analyse des Kaiser-Brunnens zu Bad Ems, 
32.—R. Fresenius. Chemische Untersuchung der warmen Quelle zu 
Schlangenbad, 49.—R. Fresenius. Chemische Analyse der Wilhelms- 
Quelle zu Kronthal, 70.—Fr. Wenckenbach. Uebersicht tiber die in 
Nassau aufgefundenen einfachen Mineralien, 147. 

- York. Natural-History Journal. Vol. iii. Nos. 3,4,6&7. 1879. 
Presented by W. Whitaker, Esq., F.G.S. 

J. EK. Clark. ae Bog I., 115. 

—. —-. Vol. Cee 23-31. 1880. Presented by W. 
Whitaker, Esq., r. G. Se 

J.P. Drewett. St. Bee’s Head, 11, 24. 

—. Yorkshire Philosophical Society. Annual Report for 1880. 
1880. 

T. S. Noble. An Account of the Scientific Work of Prof. HK. E. 
Deslongchamps, 53.—T. 8. Noble. M. J. Barrande and the Darwinian 
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Zoological Society. Proceedings. 1880, Parts 2-4. 1880-81. 

T. H. Huxley. On the Application of the Laws of Evolution to the 
Arrangement of the Vertebrata, and more particularly of the Mammalia, 
649. 

———, | ssi Part esis 


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2. Booxs. 


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Achepohl, L. Das niederrheinisch-westfalische Steinkohlengebirge. 
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Angelin, N. P., et G. Lindstrém. Fragmenta silurica e dono Caroli 
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Anon, Canada: a Hand-book of Information for intending Emi- 
grants. 8vo. Ottawa, 1879. Presented by W. Whitaker, Esq., 
HGS. 


—. Notes on the Geology of Ilfracombe and its Neighbourhood 
by a late Resident. 12mo. Ilfracombe, 1880. 


——. Scepticism in Geology and the Reasons for it, by Verifier. 
2nd edition. 8vo. London, 1878. Presented by Prof. P. M. 
Duncan, F.BRS., F.GS. 


Ansted, D. T. A short Trip in Hungary and Transylvania in the 
Spring of 1862. 8vo. London, 1862. Purchased. 


Water and Water-supply, chiefly in reference to the British 
Islands. 8vo. London, 1878. Purchased. 


Atkinson, E. T. Economic Mineralogy of the Himalayan Districts 
of the North-western Provinces. 8vo. Calcutta, 1881. 


Attwood, G. A Contribution to South-American Geology. With an 
Appendix by the Rev. Prof. 7. G. Bonney. 8vo. London, 1879. 


Attwood, G. Practical Blowpipe Assaying. 8vo. London, 1880. 


Ball, V. On Spheroidal Jointing in Metamorphic Rocks in India 
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Dublin, 1880. 


On the Evidence in favour of the Belief in the Existence of 
floating Ice in India during the Deposition of the Talchir 
(Permian or Permio-Triassic) Rocks. 8vo. Dublin, 1880. 


. On the Mode of Occurrence and Distribution of Gold in 
India. 8vo. Dublin, 1880. 


Barrande, J. Du maintien de la nomenclature établie par M. 
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Barrois, C. Note on the Rey. J. F. Blake’s paper on the Chalk of. 
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——. Note sur les alluvions de la Serre (Aisne). 8vo. Lille, 
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Sur le terrain silurien supérieur de la presqwile de Crozon. 
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Baverman, Hilary. Text-book of Systematic Mineralogy. 8vo.. 
London, 1881. 


y 2 


292 ADDITIONS TO THE LIBRARY. 


Belgium. Commnussion de la Carte Géologique de la Belgique. Texte 
explicatif du levé géologique de la planchette d’Aerschot, par M. le 
baron O. van Ertborn, avec la collaboration de P. Cogels. 8vo. 
Brussels, 1880. 


Texte explicatif du levé géologique de la planchette 
@ Anvers, par M. le baron O. van Ertborn, avec la collaboration 
de P. Cogels. 8vo. Brussels, 1880. 


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de Beveren, par M. le baron O. van Ertborn et P. Cogels. 8vo. 
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Brussels, 1880. 


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de Boom, par M. le baron O. van Ertborn, avec la collaboration de 
P. Cogels. S8vo. Brussels, 1880. 


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de Heyst-op-den-berg, par M. le baron O. van Ertborn, avec la 
collaboration de M. P. Cogels. 8vo. Brussels, 1880. 


; Texte explicatif du levé géologique de la planchette 
de Lierre, par M. le baron O. van Ertborn, avec la collaboration 
de M. P. Cogels. S8vo. Brussels, 1880. 


Texte explicatif du levé géologique de la planchette 
de Lubbeek, par M. le baron O. van Ertborn et P. Cogels. 8vo. 
Brussels, 1880. 


Texte explicatif du levé géologique de la planchette 
de “Malines, par M. le baron O. van Ertborn et P. Cogels. 8vo. 
Brussels, 1880. 


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de Putte, par M. le baron O. van Ertborn, avec la colloboration 
de M. P. Cogels. 8vo. Brussels, 1880. 


Benecke, E. W., und E. Cohen. Geognostische Beschreibung der 
Umgegend von Heidelberg. Hefte 2 & 3. 8vo. Strassburg, 
1880-81. ° 

Binney, HE. W. On some Marine fossil Shells in the Middle Coal 
Measures of Lancashire. S8vo. Manchester, 1880. 

Bittner, A., L. Burgerstein, F. Calvert, Fr. Heger, V. Hilber, 1. 
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Blake, J. H. Address at the Anniversary Meeting of the Norwich 
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Blake, W. P. Report of a Geological Reconnaissance in California, 
made in connexion with the Expedition to survey Routes in 
California to connect with the surveys of routes for a Railroad 
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men. Archiv, Band LV. No. 2(1880). Erliuterungen zur geolo- 
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On Traces of Glacial Action near Llandudno. 8yvo. London, 


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—. On supposed Pholas Burrows in Derbyshire. 8vo. London, 
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ixey al 


——. On the Formation of ‘“ Cirques,” and their Bearing upon 
Theories attributing the Excayation of Alpine Valleys mainly to 
the Action of Glaciers. 8vo. London, 1871. 


Ice Scratches in Derbyshire. 8vo. London, 1872. 


On certain Lithodomous Perforations in Derbyshire. 8vo. 
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Notes on the Roslyn-Hill Clay-pit. 8vo. London, 1872. 


Lakes of the North-eastern Alps and their Bearing on the 
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On the Occurrence of a Quartzite Boulder in a Coal-seam in 
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1876. 
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and Felsites in Arran. 8yo. London, 1877. 


On Mr. Helland’s Theory of the Formation of Cirques. 8vo. 
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hove 


On the Serpentine and associated Rocks of the Lizard 
District. With Notes on the Chemical Composition of some of 
the Rocks of the Lizard District. By W. H. Hudleston. 8vo. 
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The Lherzolite of the Ariége. 8vo. London, 1877. 


On some Specimens of Gabbro from the Pennine Alps. 8vo. 
Truro, 1878. 


On the Serpentine and associated Igneous Rocks of the 
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204 ADDITIONS TO THE LIBRARY, 
Bonney, T. G. Notes on some Ligurian and Tuscan Serpentines. 
8vo. London, 1879. 


Notes on the Relations of the Igneous Rocks of Arthur’s 
Seat. 8vo. London, 1878. 


On Professor Dana’s Classification of Rocks. 8vo. London, 


1879. 


On the Quartz-felsite and associated Rocks at the base of 
the Cambrian Series in North-western Caernarvonshire. S8vo. 
London, 1879. 


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mingham, 1879. 


David Thomas Ansted, obituary notice. 8vo. London, 1880. 
——. In Memoriam: W. H. Miller. 8vo. Cambridge, 1880. 


——. Onsome Serpentines from the Rhetian Alps. 8vo. London, 
1880. 


Petrological Notes on the Vicinity of the Upper Part of 
Loch Maree. 8vo. London, 1880. 


——. William Hallowes Miller, obituary notice. 8vo. London, 
1880. 


——. In Memoriam: W.H. Miller. 8vo. Cambridge, 1881. 


. On the Serpentine and associated Rocks of Anglesey. 8vo. 
London, 1881. 


and F.T.S. Houghton. On some Mica-traps from the Kendal 
and Sedbergh Districts. 8vo. London, 1879. 


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narvon) and Port Dinorwig. 8vo. London, 1379. 


Bosniaski, Sigismondo de. La formazione gessosa e il secondo 
piano mediterraneo in Italia. S8vo. Pisa, 1880. 


British Museum. Index to the Collection of Minerals, with references 
to the Table Cases in which the Species to which they belong are 
exhibited. 8vo. London, 1866. Presented by W. Whitaker, 
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——. A Guide to the Collection of Minerals. 8vo. London, 1867. 
Presented by W. Whitaker, Esq., F.G.S. 


Brongniart, A. Recherches sur les graines fossiles silicifiees. Pré- 
cédées d’une notice sur ses travaux par J.B. Dumas. 4to. Paris, 
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Brongniart, C. Notice sur quelques poissons des lignites de Ménat. 
8vo. Caen, 1880. 


. Rapport sur excursion faite 4 Gisors et aux environs les 
16 et 17 mai 1880. 8vo. Rennes, 1880. 


et Maxime Cornu. Observations nouvelles sur les épidémies 
eévissant sur les insectes. 8vo. Paris, 1830. 


ADDITIONS TO THE LIBRARY. 295 


Brown, John. A Description of some Borings on the Clifton Estate, 
and of the sinking at Clifton Colliery, Nottingham. 8vo. London, 
1881. 


Brown, Thomas. Illustrations of the Fossil Conchology of Great 
Britain and Ireland, with Descriptions and Localities of all the 
Species. 4to. London, 1849. Purchased. 


Bryce, J. Geology of Clydesdale and Arran. 8vo. London, 1859. 
Purchased. 

Canada. Geological Survey. Catalogues of the Silurian Fossils of 
the Island of Anticosti, by E. Billings. 8vo. Montreal, 1866. 
Paleozoic Fossils, vol. 1., 1861-65, by E. Billings. 

8vo. Montreal, 1865. 

——. -——. Report of Progress for 1878-79. S8yvo. Montreal, 
1880. 

Capellin, G. Gli strati a Congerie o la formazione Gessoso-solfifera 
nella Provincia di Pisa e nei dintorni di Livorno. 4to. Rome, 
1880. 

Cassell’s Natural History. Edited by P. M. Duncan. Vol. IV. 
4to. london, 1880. Presented by Messrs. Cassell, Petter, Galpm, 
and Co. 

Castro, M. F. de. Discurso leidos ante la Real Academia de 
Ciencias Exactas, Fisicas y Naturales en la recepcion publica, 
2 de Junio de 1878. 8vo. Madrid, 1878. 

On the Influence which a molecular Movement due to 

Klectricity may have exerted in certain Geological Phenomena, 

namely, the Metamorphism of Rocks and the Formation of 

Metalliferous Deposits. 8vo. London, 1880. 


Chambers, G. F. A Handbook for Eastbourne &c. 5th edition. 
8vo. London, 1873. Presented by W. Whitaker, Esq., F.G.S. 


Chaper, Maurice. Note sur le région diamantifére de l Afrique 
australe. 8yo. Paris, 1880. Purchased. 


Choffat, Paul. Etude stratigraphique et paléontologique des ter- 
rains jurassiques du Portugal. Premiére livraison. Le Lias et 
le Dogger au Nord du Tage. 4to. Lisbon, 1880. 


Church, J. A. The Heat of the Comstock Lode. S8vo. London, 
1880. 


Clarke, C. B. A Review of the Ferns of Northern India. 4to. 
London, 1880. 


Cohen, EH. Sammlung von Mikrophotographien zur Veranschau- 
lichung der mikroskopischen Structur von Mineralien und Ges- 
temen. Lief. 1-4. 4to. Stuttgart, 1881. Purchased. 


Collingwood, C. Rambles of a Naturalist on the Shores and Waters 
of the China Sea. 8vo. London, 1868. Purchased. 


206 ADDITIONS TO THE LIBRARY. 


Congrés International de Géologie, tenu 4 Paris, du 29 au 31 aoiit 
et du 2 au 4 septembre 1878. Comptes Rendus. 8vo. Paris, 
1880. Two copies. Presented by the Foreign Office. 


Cope, E. D. On the Extinct Cats of America. 8vo. Salem, 1880. 


Catalogue of Vertebrata of the Permian Formation of the 
United States. Svo. Salem, 1881. 


——. Mammalia of the Lower Eocene Beds. 8vo. Salem, 1881. 


On some new Batrachia and Reptilia from the Permian Beds 
of Texas; on a Wading Bird from the Amazon Shales; on the 
Nimravide and Canidze of the Miocene Period; and on the 
Vertebrata of the Wind-River Kocene Beds of Wyoming. 8vo. 
Washington, 1881. 


—-. Paleontological Bulletin. No. 32. Second Contribution to 
the History of the Vertebrata of the Permian Formation of 
Texas. Figures. 8vo. Philadelphia, 1881. 


The Systematic Arrangement of the Order Perissodactyla, 
with a Note on the Structure of the Foot of TYoxodon. S8vo. 
Philadelphia, 1881. 


Cornu, M., C. Brongniart et Dumas. Sur une épidémie dinsectes 
diptcres causée par un Champignon. 8vo. Paris, 1880. 


Craven, A. H. Description of three new Species of Land and 
Freshwater Shells from Nossi-Bé Island (N.W. coast of Mada- 
gascar). Svo. London, 1880. 


Monographie du genre Sinusigera, dOrb. 8vo. Brussels, 


1880. 


Credner, H. Geologischer Fiihrer durch das sichsische Granulit- 
gebirge. S8yvo. Leipzig, 1880. Purchased. 


Credner, H. Ueber die Vergletscherung Norddeutschlands wahrend 
der Kiszeit. S8vo. Berlin, 1880. 


Damour, A., und G. vom Rath. Ueber den Kentrolith, eine neue 
Mineralspecies. 8vo. Leipzig, 1880. 


; . Ueber den Trippkeit, eine neue Mineralspecies. 8vo. 
Leipzig, 1880. 


Dana, J.D. Appendix (I.) to the 5th edition of Dana’s Mineralogy, 
by G. J. Brush, and Appendix II., by E. be Dana. 8vo. New 
2 1876. Purchased. 


Daubrée, A. Descartes, l’un des créateurs de la cosmologie et de la 
géologie. 4to. Paris, 1880. 


——, Hxamen minéralogique et chimique de materiaux provenant 
de quelques forts vitrifiés de la France. 8vo. Paris, 1881. 


Davidson, T. Liste des principaux ouvrages mémoires ou notices 
qui traitent directement ou indirectement des Brachiopodes 
vivants et fossiles. 1606-1876. 8vo. Brussels, 1880. 


ADDITIONS TO THE LIBRARY. 297 


Davis, J. W. Report on the Raygill Fissure Exploration Com- 
mittee. 8vo. Leeds, 1881. 

Dawson, J. W. Sketch of. 8vo. ——, 1875. 

Dawson, J. W. Revision of the Land Snails of ‘the Paleozoic Era, 
with Descriptions of new Species. 8vo. New Haven, 1880. 


Delesse, FE. Sur les études de géologie agronomique aux Etats-Unis 
et en particulier sur celles de M. G. H. Cook dans le New-Jersey. 
8vo. Paris, 1880. 


Delesse, H., et A. De Lapparent. Extraits de Géologie pour les années 
1877 et 1878. 8vo. Paris, 1880. 


. Revue de Géologie pour les années 1877 et 1878. 
Tome XVI. 8vo. Paris, 1880. 


Dewalque, G. Prodrome d’une description géologique de la Belgique. 
Seconde édition. S8vo. Brussels, 1880. 


Sur Vuniformité de la langue Géologique. 8vo. Liége, 


1880. : 


Doyle, P. Petroleum: its History, Origin, and Use, with reference 
to its Advantages and Perils as an Illuminator. 8vo. Brisbane, 
1880. 


_ Duff, P. A. Ramble among the Fossiliferous Beds of Moray. Fol. 
Elgin ?, 1859. Presented by H. B. Woodward, Esq., F.GS. 


Duncan, P. M. Scientific Results of the Second Yarkand Mission, 
based upon the Collections and Notes of the late Ferdinand 
Stoliczka. Syringospheeride. 4to. Calcutta, 1879. Presented by 
the India Office. 


Duncan, P. M. An Abstract of the Geology of India. Third 
edition. Fol. London, 1881. 


, and W. Percy Sladen. A Memoir on the Kchinodermata of 
the Arctic Sea to the West of Greenland. 4to. London, 1881. 


Eichwald, E. De pecorum et pachydermorum reliquiis fossilibus, 
in Lithuania, Volhynia et Podolia repertis. 4to. Bonn, 1835, 
Purchased. 


England and Wales. Geological Survey. Memoirs. Figures and De- 
scriptions illustrative of British Organic Remains. Decade XIII. 
Ato. London, 1872. 


English Channel. Pilote de la Manche. Cote nord de France, par 
K. de Courthille avec le concours de F. Hedouin. Tome l. 8vo. 
Paris, 1880. Presented by the Dépot de la Marine. 


Etheridge, R., jun. A Catalogue of Australian Fossils (including 
Tasmania and the Island of Timor), stratigraphically and zoolo- 
gically arranged. 8vo. Cambridge, 1878. Purchased. 


Evans, John. The Ancient Bronze Implements, Weapons, and Orna 
ments of Great Britain and Ireland. 4to. London, 1881. 


298 ADDITIONS TO THE LIBRARY. 


Evington Coal-Boring Company, Limited. Prospectus. [Containing 
Reports by R. Etheridge and James Plant.| Fol. Leicester, 
1878? Presented by W. Whitaker, Esq., F.GNS. 


Falsan, A., and E. Chantre. Monographie géologique des Anciens 
Glaciers et du Terrain Erratique de la partie moyenne du Basin 
du Rhone. Text, 2 vols, Svo., 1879-80. Atlas, fol., 1875: 
Lyon. 


Favre, Ernest. Revue géologique suisse pour Vannée 1880. XI. 
8vo. Geneva, 1881. 


Festenberg-Packisch, Hermann. Der metallische Bergbau Nieder- 
schlesiens unter bentitzung amtlicher Quellen in geognostischer, 
historischer und technischer Beziehung. S8vo. Vienna, 1881. 
Purchased. 


Figari, Antonio. Studii scientifici sull’ Egitto e sue adiacenze com- 
presa la penisola dell’ Arabia Petrea. 2 vols. S8vo. Lucca, 1864— 
65. Presented by H. Bauerman, Esq., LGN. 


Finlands Geologiska Undersokning. Beskrifning till Kartbladet 
No. 2, af K. Ad. Moberg. 8vo. Helsingfors, 1880. 
Fleming, John. The Lithology of Edinburgh. Edited, with a Me- 


moir, by the Rey. John Duns. 8vo. Hdinburgh, 1859. Pur- 
chased. 


Fontannes, F. Description des Ammonites des calcaires du Chateau 
de Crussol, Ardéche. 4to. Lyon, 1879. Purchased. 


Foote, Rk. Bruce. On the Geology of parts of the Madras and North 
Arcot Districts lying north of the Palar River, and included in 
Sheet 78 of the Indian Atlas. S8vo. Calcutta, 1873. 


. The Auriferous Rocks of the Dambal Hills, Dharwar Dis- 
trict. 8vo. Calcutta, 1874. 


The Geological Features of the South Mahratta Country and 
the adjacent Districts. Svo. Calcutta, 1876. 


Notes on the Representatives of the Upper Gondwana Series 
in Trichinopoly and Nellore-Kistna Districts. 8vo. Calcutta, 
1878. 


. On the Geological Structure of the Eastern Coast from Lati- 
tude 15° northward to Masulipatam. 8vo. Calcutta, 1879. 


Rough Notes on the Cretaceous Fossils from Trichinopoly 
District, collected in 1877-78. 8vo. Calcutta, 1879. 


Sketch of the Geology of North Arcot District. 8vo. Cal- 
cutta, 1879. 


Forrest, Alex. North-west Exploration. Journal of expedition 
from De Grey to Port Darwin. 4to. Perth, Western Australia, 
1880. Presented by the Governor of Western Australia. 


ADDITIONS TO THE LIBRARY. 299 


France. Carte géologique detaillée de la France. Mémoires. Miné- 
ralogie Micrographique: Roches Eruptives Francaises, par F. 
Fouqué et A. M. Lévy. 4to. Paris, 1879. Text and Atlas. 
Purchased. 


: . ——. Le pays de Bray, par A. de Lapparent. Ato. 
Paris, 1879. 
Purchased. 


—. —. —. Notice explicative. Feuilles 9, 23, 34, 50, 
67, 69, 78, 93, 113, 122, 1387, 213 bis, 225 bis, 237. (In slips.) 
1881. Purchased. 


Fritsch, A. Fauna der Gaskohle und der Kalksteine der Permfor- 
mation Bohmens. Band I. Hefte 2 und 3.°4to. Prag, 1880-81. 


Gaudry, A. Sur un Reptile trés perfectionné, trouvé dans le terrain 
permien. 4to. Paris, 1880. 


Sur un nouveau genre de poisson primaire. 4to. Paris, 


1881. 


Geikie, A. Outlines of Field Geology. 8vo. London, 1876. Pre- 
sented by W. Whitaker, Hsq., F.GS. 


Geikie, J. Prehistoric Europe: a geological sketch. Svo. London, 
1881. Purchased. 


Gemitz, H. B. Nachtrige zur Dyas. I. Mit Beitragen der Herren 
Drude, Vetter und Weiss. 4to. Cassel, 1880. 


Geyler, H. T. Phytopaleontologie. 8vo. ——-,1878. Presented 
by W. Whitaker, Esq., F.GS. 


Giebel, C.G. Die Saugethiere in Zoologischer, anatomischer und 
paliontologischer Beziehung. 2nd edition. 8vo. Leipzig, 1859. 
Purchased. 


Gilpin, H. The Mines and Mineral Lands of Nova Scotia. 8vo. 
Halifax, N.S., 1880. 


Goodyear, W. A. Harthquake and Volcanic Phenomena, December 
1379 and January 1880, in the Republic of Salvador, Central Ame- 
rica. Svo. Panama, 1880. (Iwo copies.) 


Gosse, Philip Henry. Omphalos: an attempt to untie the geological 
knot. 8vo. London, 1857. Purchased. 


Gosselet, J. Constitution géologique du Cambresis. Svo. Cambrai? 
1865. Presented by W. Whitaker, Esq., F.GN. 


Esquisse géologique du département du Nord et des contrées 
voisines. Fasc. 1 et 2. 8vo. Lille, 1873-76. Presented by W. 
Whataker, Esq., 2.GS. 


Gosselet, J. Le terrain dévonien des environs de Stolberg. 8vo, 
Lille, 1876. 


——. Relations des sables d’Anvers avec les systemes Diestien et 
Boldérien. S8vo. Lille, 1876. 


"aco ADDITIONS TO THE LIBRARY. 


Gosselet, J. Compte-rendu de lVexcursion dans les Ardennes du 23 
aout au 5 septembre 1876. 8vo. Lille, 1877. 


Quelques réflexions sur la structure et lage du terrain 
houiller du Nord de la France 4 Voccasion du Mémoire de M. 
Breton et de celui de M. ’Abbé Boulay. 8vo. Lille, 1877. 


——. Sur quelques fossiles trouvés dans le silurien des Ardennes. 
8vo. Lille, 1877. 


Apercu sur la constitution géologique de la Forét de Mormal. 
8vo. Lille, 1878. 


Excursion dans les tranchées du chemin de fer de Cambrai 
au Quesnoy. 8vo. Lille, 1878. 


——. le marne de la Porquerie (€océne inférieur). 8vo. Lille, 
1878. 


——. Le calcaire dévonien supérieur dans le N.-E. de larrondisse- 
ment d’Avesnes et documents pour l’étude des schistes de Famenne. 
8yvo. Lille, 1878. 


——. Documents nouveaux pour l’étude du Famennien. Tran- 
chées du chemin de fer entre Féron et Semeries. Schistes de Sains. 
8vo. Lille, 1879. 


Le calcaire de Givet. 3™° et 4™° parties, suivies de consi- 
dérations sur la terminaison orientale de la grand faille. 8vo. 
Lille, 1879. 


——. IT /argile a silex de Vervins. 8vo. Liaille, 1879. 


——. Laroche aTfépin. Contact du terrain silurien et du terrain 
dévonien sur les bords de la Meuse. 8vo. Lille, 1879. 


Notice necrologique sur Jean-Baptiste-Julien d’Omalius 
d’Halloy. 8vo. Lille, 1879. 


——. De Vusage du droit de priorité et de son application aux 
noms de quelques Spiriferes. 8vo. Lille, 1880. 


——. Les roches cristallines des Ardennes. 8vo. Lille, 1880. 


——. Notes sur les sables tertiaires du Plateau del’Ardenne. Lille, 
1881. 


——. Terrain diluvien de la Vallée de la Somme. 8vo. Lille, 
1880. 


——. Troisiéme note sur le Famennien. Tranchée du chemin de 
fer du Luxembourg. Les schistes de Barvaux. 8vo. Lille, 1880. 


Esquisse géologique du Nord de la Frauce et des contrées 
voisines. 1% fascicule. Terrains primaires. Text and Plates. 8vo. 
Lille, 1880. 

—— et Henri Rigaux. Mouvement du sol de la Flandre depuis les 
temps géologiques. 8yvo. Lille, 1878. 

Gotthard Tunnel. Geologische Tabellen und Durchschnitte uber den 


Grossen Gotthardtunnel. Lief.4—-6. 4to. Zurich, 1877-79. Pur 
chased. 


ADDITIONS TO THE LIBRARY. 301 


Gregorio, Antono de. Fauna di 8. Giovanni [larione (Parisiano). 
Parte 1°. Cefalopodi e Gasteropodi. Fasc. 1. 4to. Palermo, 
1880. | 

Giimbel, C.W. Geognostische Mittheilungen aus den Alpen. VII. 
8vo. Munich, 1880. 

Nachtrage zu den Mittheilungen iiber die Wassersteine 
(Enhydros) von Uruguay und ber einige stid- und mittel-ameri- 
kanische sogenannten Andesite. 8vo. Munich, 1881. 

Ginther, A.C. L.G. An Introduction to the Study of Fishes. 8vo. 
Edinburgh, 1880. Purchased. 

Gutch’s Literary and Scientific Register and Almanack for the year 
1867. (Geology, by J. B. Jukes.) 12mo. London, 1866. Presented 
by W. Whitaker, Esq., F.G.S. 

Habenicht, H. Die Grundzige im geologischen Bau Europa’s, mit 
einer Karte, “‘ Die Verbreitung der Kruptiv- und Uebergangs- 
gesteine in Europa,” und funf Nebenkarten. 8vo. Gotha, 1881, 

.Hahn, Otto. Die Meteorite (Chondrite) und ihre Organismen. 4to. 
Tubingen, 1880. Purchased. 

Haniel, John. Die Flotzlagerung in der Stoppenberger und Horst- 
Hertener Mulde des Westfilischen Steinkohlengebirges. to. 
Essen, 1881. Purchased. 

Hann, J., F. vy. Hochstetter und A. Pokorny. Allgemeine Erdkunde. 
Ein Leitfaden der astronomischen Geographie, Meteorologie, Geo- 
logie und Biologie. 8ve. Prague, 1872. Purchased. 

Hawes, G.W. The Albany Granite, New Hampshire, and its con- 
tact phenomena. S8vo. New Haven, 1881. 

Hébert, E. Histoire géologique du canal de la Manche. 4to. Paris, 
1880. 


Recherches sur la craie superieure du versant septentrional 
des Pyrénées. 4to. Paris, 1880. 


Nomenclature et classification géologiques. 8vo. Paris, 


1881. 

Heer, O. Flora fossilis Artica. Band 2. 4to. Winterthur, 1871. 
Purchased. 

Band 6. 1 Abth. 4to. Zurich, 1880. Purchased. 

Hicks, Henry. On some recent researches among Pre-Cambrian 
Rocks in the British Isles. 8vo. London, 1881. 

Hill, E., and 7. G. Bonney. The Precarboniferous Rocks of Charn 
wood Forest. Part 1. S8vo. London, 1877. 

Part 2. 8vo. London, 1878. 

——. Part3. 8vo. London, 1880. 


Hinde, G. J. Fossil Sponge-spicules from the Upper Chalk, found 
in the interior of a single flint-stone from Horstead in Norfolk. 
8vo. Munich, 1880. 


302 ADDITIONS TO THE LIBRARY. 


Hitchcock, E. The Religion of Geology. 8vo. London, 1851. 
Purchased. 

Hornes, R. Die Trilobiten-Gattungen Phacops und Dalmanites 
und ihr vermuthlicher genetischer Zusammenhang. 8vo. Vienna, 
1830. 

——. Materialien zu einer Monographie der Gattung Megalodus 
mit besonderer beriicksichtigung der mesozoischen Formen. 4to. 
Vienna, 1880. 

und M. Auinger. Die Gasteropoden der Meeres-ablagerungen 
der ersten und zweiten miocinen Mediterran-stufe in der oster- 
reichisch-ungarischen Monarchie. Parts 1&2. 4to. Vienna, 
1879-80. 

Holm, G. Anteckningar om Wahlenbergs Illcenus crassicauda. 8vo. 
Stockholm, 1880. 

Bemerkungen iiber IJlenus crassicauda. 8vo. Berlin, 1880. 


Hudleston, W. H. Contributions to the Paleontology of the York- 
shire Oolites. S8vo. London, 1881. : 

Hull, E. On the Geological Structure of the Northern Highlands of 
Scotland ; being notes of a recent tour. 8vo. Dublin, 188]. 

The Coalfields of Great Brirain; their History, Structure, 
and Resources. Fourth edition. 8vo. London, 1881. 

Idria. K.-k. Bergdirection. Das k.-k. Quecksilberwerk zu Idria in 
Krain. 4to. Vienna, 1881. Presented by the K.-k. Ackerbaumt- 
nisterium, Vienna. 

Tllinois. Geological Survey. Report. Vol. 3, by A. H. Worthen, 
H. Engelmann, H.C. Freeman, H. M. Bannister, and F. B. Meek. 
Ato. Springfield, Ill., 1868. Presented by the late Dr. J. J. Bigsby, 
JBL TiS, JEG IS 

. —. Vol. 4, by A. H. Worthen, H. M. Bannister, 
ae H. Bradley, H. A. Green, J. 8. Newberry, and Leo Lesquereux. 
Ato. Springfield, Ill.,1870. Presented by the late Dr. J. J. Bigsby, 
JBI gShy, HAAG HS 

——. : . Vol. 5, by A. H. Worthen, J. Shaw, and F. 

B. Meek. 4to. Springfield, Ill., 1873. Presented by the late 
Dr. J. J. Bigsby, LLS., F.GS. 

as, ——. Vol. 6, by A. H. Worthen, G. C. Broadhead, 
KE. T. Cox, 0. St. J ohn, and F. B. Meek. to. Springfield, IIL, 
1875. Presented by the late Dr. J. J. Bigsby, P_BS., EGS. 

India. Geological Survey. Memoirs. Vol. xv. Part 2. 1880. 

Vol. xvi. Part 1. 1880. 

. Paleontologia Indica. Ser.10. Indian Tertiary and 

Post-tertiary Vertebrata. Vol. i. Part 4. Supplement to Crania 

of Ruminants, by R. Lydekker. 1880. 

Ser. 10. Vol.i. Part 5. Siwalik and Narbada 

Proboscidia, by R. Lydekker. 1880. 


ADDITIONS TO THE LIBRARY. 303 


India. Geological Survey. Paleontologia Indica. Ser. 13. 
Salt-Range Fossils, by W.Waagen. Part2. 1880. 


ee. Records, Vol) xi. /Parts 3&4. 1880, 
Molt xaiveinanismivanece Toole 


Indiana. Department of Statistics and Geology of the State of In- 
diana. First Annual Report, 1879. 8vo. Indianopolis, 1880. 
Jack, R. L. Report on the Geology and Mineral Resources of the 
District between Charters Towers Goldfield and the Coast. 4to. 

Brisbane, 1879. 


Report to the Honourable the Minister for Mines on the 
Bowen River Coalfield. 4to. Brisbane, 1879. 

Jeffreys, J. Gwyn. The French Deep-sea Exploration in the Bay 
of Biscay. 8vo. London, 1880. 

—. Deep-sea Exploration. 8vo. London, 1881. 

Jervis, G. Dell’ Oro in Natura. 8vo. Turin, 1880. 


Jones, T. Rupert. Unio and Paludine from the Wealden Beds of 
Kent, in the collection of Mr. W. Harris of Charing. 8vo. London, 
1860. 


Note on the Well lately sunk at Wokingham, Berkshire. 
8yo. London, 1880. 


Report of the Excursion of the Geologists’ Association to 
Camberley, April 24, 1880. 8vo.. London, 1880. 


Fossil Foraminifera of the Carboniferous Limestone. (Review 
of Memoirs by V. von Moller.) Slip. London, 1881. 


On the Geology and Physical Features of the Bagshot Dis- 
trict. 8vo. London, 1881. 


Julien, A. A. On Biotite as a Pseudomorph after Olivine. 8vo. 
New York, 1878. 


——. On the Geological Action of the Humus Acids. 8vo. Salem, 
1880. 


Karsten, G. Beitrige zur Landeskunde der Herzogthumer Schleswig 
und Holstein Reihe2. Heftelund2. 4to. Berlin, 1869 & 1872. 
Purchased. 


Kobell, V. Obituary notices of Johann von Lamont, Carl Friedrich 
Mohr, Heinrich Wilhelm Dove, Aug. Heinrich Rudolph Grisebach, 
Bernhard v. Cotta, and J. F. Brandt. 8vo. Munich, 1880. 


——-. Ueber Polarisationsbilder an Zwillingen zweiaxiger Kry- 
stalle. S8vo. Munich, 1880. 


Kuntze, O. Ueber Geysirs und nebenan entstehende verkieselte 
Baume. 4to. Stuttgart, 1880. 


Leidy, J. Bathygnathus borealis, an extinct Saurian of the New 
Red Sandstone of Prince Edward’s Island. 4to. Philadelphia, 
18381. 


304 ADDITIONS TO THE LIBRARY. 


Leidy, J. Remarks on Bathygnathus borealis. 4to. Philadelphia, 
1881. 
Locard, A. Nouvelles recherches sur les argiles lacustres des ter- 
rains quaternaires des environs de Lyon. 8vo. Lyon, 1880. 
Purchased. 


Loretz, H. Ueber Schieferung. 8vo. Frankfurt a. M., 1880. 


Macpherson, J. Uniclinal Structure of the Iberian Peninsula. S8vo. 
Madrid, 1880. 


Maestre, Amaleo. Descripcion geologica industrial de la cuenca car- 
bonifera de San Juan de las Abadesas en la Provincia de Gerona. 
Ato. Madrid, 1855. Purchased. 


Mallet, F. R. On the Ferruginous Beds associated with the Basaltic 
Rocks of North-eastern Ulster, in relation to Indian Laterite. 
8vo. Calcutta, 1881. 


Marcou, Jules. Sur les colonies dans les roches taconiques des bords 
du lac Champlain. 8vo. Paris, 1881. 


Marsh, O. C. Notice of Jurassic Mammals representing two new 
Orders. S8vo. New Haven, 1880. 


—. The Sternum in Dinosaurian Reptiles. 8vo. New Haven, 
1880. 

A new Order of extinct Jurassic Reptiles (Ceeluria). 8vo. 

New Haven, 1881. 


Discovery of a fossil Bird in the Jurassic of Wyoming. 8vo. 
New Haven, 1881. 


——. Note on American Pterodactyls. &vo. New Haven, 1881. 


——., Principal Characters of American Jurassic Dinosaurs. Part 
IV. Spiral Cord, Pelvis, and Limbs of Stegosaurus. 8vo. New 
Haven, 1881. 


Martin, K. Die Tertiirschichten auf Java. 4to. Leiden, 1880. 
Purchased. 


Mathews, W. The Flora of Algeria, considered in relation to the 
Physical History of the Mediterranean Region and supposed Sub- 
mergence of the Sahara. 8vo. London, 1880. 


Mawe, John. Travels in the Gold and Diamond Districts of Brazil. 
Svo. London, 1825. Purchased. 


Meli, Romolo. Sui dintorni di Civitavecchia. 4to. Rome, 1879. 


Meneghini, G. Fossili oolitici di Monte Pastello nella provincia di 
Verona. 8vo. Pisa, 1880. 


——. Nuovi fossili siluriani di Sardegna. 4to. Rome, 1880. 


Merccy, N. de. Sur un sondage exécuté a Saint-Blimont (Somme). 
Svo. Amiens, 1879. 


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Mercey, N. de. . Composition des sables de Bracheux et mode d’origine 
de l’argile plastique, premier produit d’une émanation terminée par 
le dépot du caleaire de Mortemer, d’aprés des coupes du chemin 
de fer de Compicgne (Oise) & Roye (Somme). 8vo. Paris, 1880. 


Compte-rendu du course de la Soci¢té géologique de France 
& Maignelay le samedi 7 septembre 1878. S8vo. Paris, 1880. 


Note sur la confusion résultant de ’emploi de la dénomina- 
tion @argile a silex appliquée 4 deux dépots placés, Pun a la base, 

- et l'autre au sommet de la série tertiaire du Nord de la France. 8vo. 
Lille, 1880. 


Observations a loccasion de quelque travaux publiés dan 
les Annales de la Société géologique du Nord sur le quaternaire 
ancien. 8vo. Lille, 1880. 


Remarques sur la classification du terrain crétacé supérieur. 
8vo. Paris, 1880. 


—. Sur les couches de Sinceny. 8vo. Paris, 1880. 


Minnesota. Geological and Natural-History Survey. Sixth Annual 
Report, for the year 1877. 8vo. Minneapolis, 1878. Presented 
by Prof. N. H. Winchell, State Geologist. 


Seventh Annual Report, for the Ne 1878. 8vo. 
Minneapolis, 1879. Presented by Prof. N. H. Winchell, State 
Geologist. 


=——, 


Eighth Annual Report, for the year 1879. 8vo. 
Saint Paul, 1880. Presented by the State Geologist. 


Moyjsisovics, HL. v., H. Tietze, und A. Bittner. Grundlinien der Geo- 
logie von Bosnien-Hercegovina. S8vo. Vienna, 1880. 


Ueber heteropische Verhiltnisse 1m Triasgebiete der lom- 
_bardischen Alpen. S8vo. Vienna, 1880. 


Morris, John. Coal; its Geological and Geographical Position. 8vo. 
London, 1862. Presented by Prof. T. Rupert Jones, P.RS., 
areeGsS. 


Morton, G. H. The Carboniferous Limestone and Cefn-y-fedw Sand- 
stone of the Country between Llanymynech and Minera, North 
Wales. 8vo. London, 1879. Purchased. 


Mourlon, Michel. Géologie de la Belgique. 2 vols. 8vo. Brussels, 
1880 & 1881. 


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Murray, Alex. Report on the Gold Region near Brigus, Conception 
Bay, Newfoundland. Fol. St. John’s, 1880. 


Nathorst, A. G. Berattelse, afgifven till Kongl. Vetenskaps-Akade- 
mien, om en med understod af allminna medel utford vetenskaplig 
resa till England. 8vo. Stockholm, 1880. Presented by (. Reid, 
Esq., F.GS. 

VOL. XXXVII. ze 


306 ADDITIONS TO THE LIBRARY. 


Naumann, C. F. Lehrbuch der Geognosie. Paliontologischer Atlas. 
70 Tafeln. 4to. Leipzig,n.d. Purchased. 


New Jersey. Geological Survey. Annual Report for the year 1880. 
8vo. Trenton, N.J., 1880. (G. H. Cook, State Geologist.) 


New South Wales. Department of Mines. Annual Report for 1878. 
Ato. Sydney, 1879. 


for 1879, and Maps. 4to. Sydney, 1880. 


Report upon certain Museums for Technology, Science, and 
Art, also upon Scientific, Professional, and Technical Instruction 
and Systems of Evening Classes in Great Britain and on the Con- 
tinent of Europe. By A. Liversidge. Fol. Sydney, 1880. 


New York, Natural History of. Paleontology. Vol. V. Part 2 
(Text and Plates). By James Hall. 4to. Albany, 1879. 


New Zealand. Colonial Museum and Geological Survey Department. 
Manual of the New-Zealand Mollusca. By F.W. Hutton. 8vo. 
Wellington, 1880. 

Palaontology of New Zealand. Part IV. Corals and 


Bry ozoa of the Neozoic Period in New Zealand. By J. KE. Tenison 
Woods. 8vo. Wellington, 1880. 


. Colonial Musewm. Meteorological Report, 1880: including 
returns for 1877, 1878, 1879, and averages for previous years. 
8vo. Wellington, 1882. 

——. -——. 15th Annual Report. 8vo. Wellington, 1880. 


New-Zealand Court, International Exhibition, Sydney, 1879. Ap- 
pendix to Official Catalogue. 8vo. Wellington, 1880. Pre- 
sented by the Colonial Government. 


Nicholson, H. A. On the Structure and Affinities of the Genus 
Monticulipora and its Subgenera, with critical Descriptions of 
illustrative Species. 8vo. Hdinburgh, 1381. 

** Norwegian North-Atlantic Expedition.” Den Norske Nordhavs- 
Expedition, 1876-1878. Chemi af H. Tornée. 4to. Christiania, 
1880. Presented by the Meteorological Institute, Christiania. 


——. =. Zoology. Fiske, af R. Collett. 4t0. Christiama, 
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Ombon, G. Denti di Ippopotamo da aggiungersi alla fauna fossile 
del Veneto. 4to. Venice, 1880. 


Ormathwaite, John Lord. Astronomy and Geology compared. 8vo. 
London, 1872. Purchased. 


Paléontologie francaise. 1" Série. Animaux invertébrés. Terrain 
Jurassiqué. Tome III. Livr.3 et 9. Gastéropodes. 8vo. Paris? 
Purchased. 


——, —, ——. Livr. 44-46. Echinodermes nesuliens 
par G.Cotteau. 8yvo. Paris, 1880-81. Purchased. 


ADDITIONS TO THE LIBRARY. 307 


Parker, J. Map and Sections of Strata south of Oxford, prepared 
for the meeting of the Excursion of the Geological Association 
May 17, 1880. 8vo. Oxford, 1880. Presented by W. Whitaker, 
Esq., F.GS. 


Pengelly, W. Notes on Boulders and Scratched Stones in South 
Devon. Part 3. 8vo. Plymouth, 1880. 


Notes on recent Notices of the Geology and Paleontology of 
Devonshire. Part 7. 8vo. Plymouth, 1880. 


Sixteenth and concluding Report of the Committee, consist- 
ing of John Evans, Sir John Lubbock, Bart., Edward Vivian, G. 
Busk, W. Boyd Dawkins, W. A. Sanford, J. E. Lee, and W. Pen- 
gelly, appointed for the purpose of exploring Kent’s Cavern, De- 
vonshire. 8vo. London, 1880. 

Percy, C. M. Winding and Overwinding. 8vo. Wigan, 1880. 

Pettersen, Karl. Lofoten og Vesteraalen. 8vo. Christiania, 1881. 


Piret, F. M. La fontaine ardente de Saint-Barthélemy (Isére). Gas 
naturel, Charbon, Pétrole. 8vo. Grenoble, 1881. 


Ponzi, G. I terremoti delle epoche subappennine. 8vo. Rome, 
1880. 


Sui lavori del Tevere, e sulle variate condizioni del suolo 
romano. 4to. Rome, 1880. 


Preussen und die thiiringische Staaten. Geologische Specialkarte. 
Erliuterungen. Gradabtheilung 67. Nos. 53, 54, 59, 60. 8vo. 
Berlin, 1880. Presented by the Prussian Minster of Manu- 
fauctures, Trade, and Aris. 


Gradabtheilung 68. Nos. 49 & 55. 8vo. 
Tediine 1880. Presented by the Prussian Minister of Manufactures 
fc. 


—_——— ed 
e 


Gradabtheilung 80. Nos. 19, 20, 25, 26, 
31,32. 8vo. Berlin, 1880. Presented by the Prussian Minister 
of Manufactures &c. 


Quenstedt, F.A. Petrefactenkunde Deutschlands. Abth.1. Band VI. 
Heft 5 (Korallen). Text, 8vo; Atlas, 4to. Leipzig, 1880. Pur- 
chased, 


. Abth. 1. Band VI. Heft 6 (Korallen, Heft 11). 
Text, 8vo; Atlas, fol. Leipzig, 1880. Purchased. 


Reade, T. M. A Problem for Irish Geologists in Postglacial Geology. 
8vo. Dublin, 1879. 


—. Oceans and Continents. 8vo. London, 1880. 


——. The Glacial Beds of the Clyde and Forth. 8vo. Liverpool, 
1880. 


Reinsch, Paulus Friedrich. Neue Untersuchungen iiber die Mikro- 


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Renault, B. Cours de botanique fossile fait au Muséum d’Histoire 
naturelle. Premiere Année. 8vo. Paris, 1881. Purchased. 


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comprise entre le Rhone et le Rawyl (groupes des Diablerets et 
du Wildhorn). 8vo. Lausanne, 1880. 


——. Rapport sur la marche du Musée géologique vaudois en 
1879. 8vo. Lausanne, 1880. 


Seconde compte-rendu de la Commission géologique interna- 
tionale pour l unification des procédés graphiques. 8vo. Lausanne, 
1881. 


Reyer, H. Allgemeine Geschichte des Zinnes. 8vo. Vienna, 1880. 
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-——. Zinn: Eine geologisch-montanistisch-historische Mono- 
grafie. 8vo. Berlin, 1881. 


Ricketts, C. On the Carboniferous Limestone near Skipton and in 
North Derbyshire. 8vo. Liverpool, 1880. 


Riviére, Emile. Paléoethnologie. De lantiquité de ’homme dans 
les Alpes-Maritimes. Livr. 7. 4to. Paris, 1879. Purchased. 


——, = Livr. 8. -4to! Paris) 1Seilieercnoscoe 


Ruskin, J. Deucalion. Collected Studies of the Lapse of Waves 
and Life of Stones. Part VII. 8vo. Sunnyside, Orpington, 1880. 


Rutot, A., et H. van den Broeck. Les phénoménes post-tertiaires en 
Belgique dans leurs rapports avec lorigine des dépéts quaternaires 
et modernes. 8vo. Lille, 1880. 


Sandberger, F. Kin Beitrag zur Kenntniss der unterpleistocinen 
Schichten Englands. 4to. Cassel, 1880. 


. Ueber die Bildung yon Erzgiingen mittelst Auslaugung des 
Nebengesteins. Svo. Berlin, 1880. 


Saporta, G. de, et A. F. Marion. L’évolution du régne vegetal. 
Les Cryptogames. 8vo. Paris, 1881. 


Sartorius von Waltershausen, W .Der Aetna. Herausgegeben, selbst- 
stindig bearbeitet und vollendet von A. von Lasaulx. Band I. & 
II. 4to. Leipzig, 1880. Purchased. 


Sauvage, H. E. Recherches sur les poissons fossiles du terrain 
erétacé de la Sarthe. S8vo. Paris, 1872. 


——. Notes sur les reptiles fossiles. Svo. Paris, 1873. 


-——. Notice sur les poissons tertiaires de VAuvergne.  §8vyo. 
Toulouse, 1874. 


Notes sur les reptiles fossiles. 8vo. Paris, 1876, 


ADDITIONS TO THE LIBRARY. : 309 
Sauvage, H. H.. Sur les Lepidotus palliatus et Spherodus gugas. 
8vo. Paris, 1877. 


Etude sur les poissons et les reptiles des terrains crétacé 
et jurassique supérieur de l’Yonne. 8yo. Auxerre, 1880 (?). 


Etude sur les poissons des faluns de Bretagne. 4to. Chalon- 
sur-Sadne, 1880. 


Saxony. Geologische Landesuntersuchung des Kongreichs Sachsen. 
Erlauterungen zur geologischen Specialkarte. Blatt 28,75, & 115. 
8vo. Leipzig, 1880. 


Schmid, E. E. Die quarzfreien Porphyre des centralen Thiringer 
Waldgebirges und ihre Begleiter. 4to. Jena,1880. Purchased. 


Schmidt, A. Die Zinkerz-Lagerstatten von Wiesloch (Baden). 8vo. 
Heidelberg, 1881. 


Schmidt, Oscar. Die Spongien des Meerbusen von Mexico (und des 
earaibischen Meeres). Heft 2. 4to. Jena, 1880. Purchased. 


Schédler, F. The Treasury of Science, Natural and Physical. 
Translated by H. Medlock. 8vo. London, 1862. Presented by 
W. H. Dalton, Esq., F.GS. 


Scrope, G. Poulett. On the supposed Influx of Water to the Interior 
of the Globe as the Cause of Volcanic Action. 8vo. London, 1869. 
Presented by Prof. J. W. Judd, F.RS., Sec. GS, 


On the Character and Composition of Lavas. 8vo. London, 
1870. Presented by Prof. J. W. Judd, F.RS., Sec. GS. 


——. Notes on the late Eruption of Vesuvius. 8vo. London, 
1872. Presented by Prof. J. W. Judd, F.RS., Sec. GS. 

. On “ Blocky” Rock-surfaces, and the Theory of the Shrink- 

ing Nucleus of the Globe. 8vo. London, 1873. Presented by Prof. 

HaWeadudd, Fis. 


Scudder, S. H. The Devonian Insects of New Brunswick. en. 
Boston, 1880. 

. The Structure and Affinities of Hwphoberia, Meek & Worthen, 

a Genus of Carboniferous Myriapoda. 8vo. New Haven, 1881. 


Sheafer, P. W. The Anthracite Coalfields of Pennsylvania, and their 
Exhaustion. 8vo. Haston, 1880 (?). 


Simpson-Baikie, H. The International Dictionary for Naturalists 
and Sportsmen, in English, French, and German. 8vo. London, 
~ 1880. 


Sollas, W. J. On the Ventriculite of the Cambridge Upper Green- 
sand. S8vo. London, 1873. 


——. On Evolution in Geology. 8vo. London, 1877. 


——. On Pharetrospongia Strahani, Sollas, a fossil Holorhaphidote 
Sponge from the Cambridge ‘‘ Coprolite”-bed. 8vo. London, 
1877. 


310 ADDITIONS TO THE LIBRARY. 


Sollas, W. J... On Stauronema, a new Genus of fossil Hexactinellid 
Sponges, with a Description of its two Species, S. Carteri and 
S. lobata. 8vo. London, 1877. 


On the Changes produced in the Siliceous Skeletons of cer- 
tain Sponges by the Action of Caustic Potash. 8vo. London, 
1877. 


On the Perforate Character of the Genus Webbina, with a 
Notice of two new Species, W. levis and W. tuberculata, from the 
Cambridge Greensand. 8vo. London, 1877. 

On the Structure and Affinities of the Genus Stphonia. 8vo. 
London, 1877. 

. On the Structure and Affinities of the Genus Catagma. 8vo. 
London, 1878. 

On two new and remarkable Species of Cliona. 8vo. London, 


1878, 
Observations on Dactylocalyx pumiceus (Stutchbury), with 
a Description of a new Variety, Dactylocalyx Stutchburyi. 8vo. 
London, 1879. 

On Plectronella papillosa, a new Genus and Species of Echi- 
nonematous Sponge. 8vo. London, 1879. 


On Plocamia plena, a new Species of Echinonematous Sponge. 
8vo. London, 1879. 

——. On some Three-toed Footprints from the Triassic Conglo- 
merate of South Wales. 8vo. London, 1879. 


——. On the Silurian District of Rhymney and Pen-y-lan, Cardiff. 
8vo. London, 1879. 


——. On some Eskimos’ Bone-Implements from the East Coast 
of Greenland. 8vo. London, 1880. 

. On the Flint Nodules of the Trimmingham Chalk. 8vo. 
London, 1880. 

——, On the Structure and Affinities of the Genus Protospongia 
(Salter). 8vo. London, 1880. 

——. The Sponge-fauna of Norway: a Report on the Rev. A. M- 
Norman’s Collection of Sponges from the Norwegian Coast. 8vo. 
London, 1880. 


Sollas, W. J., and A. J. Jukes-Brown. On the included Rock-frag- 
ments of the Cambridge Upper Greensand. 8yvo. London, 1873 


South Australia. Public Works Report. Report from the Public. 
Works Department for the Half Year ended 30th June, 1878. 4to. 
Adelaide, 1879. Presented by the Colonial Government. 

Spain. Comision del mapa geologico de Espana. Boletin. Tomo VII- 
Cuadernol1 &2. 8vo. Madrid, 1880. 


Spencer, J. W. Discovery of the Preglacial Outlet of the Basin of 


ADDITIONS TO THE LIBRARY. 3I1 


Lake Erie into that of Lake Ontario; with Notes on the Origin 
of our Lower Great Lakes. 8vo. Philadelphia, 1881. 


Spencer’s Illustrated Leicester Almanack dc. for 1880. 8vo. 
Leicester, 1880. Presented by W. Whitaker, Esq., F.G.S. 


Spratt, T. A. B. Report on the Present State of the Navigation of 
the River Mersey (1880). 8vo. London, 1881. 


Struckmann, C. Die Wealden-Bildungen der Umgegend von Han- 
nover. 8yvo. Hannover, 1880. (Two copies.) 


Ueber den Parallelismus der hannoverschen und der 
euglischen oberen Jurabildungen. 8vo. Stuttgart, 1881. 


Swansea. British Association. The Official Guide and Handbook 
to Swansea and District, by 8. C. Gamwell. 8vo. Swansea, 1880. 
Presented by W. Whitaker, Esq., F.GS. 


Switzerland. Comnussion géologique suisse. Materiali per la carta 
geologica. Vol. XVII. Il canton Ticino meridionale ed 1 paesi 
finitimi, per Torquato Taramelli. Appendice ed Indice. 4to. 
Bern, 1880. 


: Beitrige zur geologischen Karte. Lieferung 20. Der 
mechanische Contact von Gneiss und Kalk im Berner Oberland 
von A. Baltzer. 4to. Bern, 1880. 


Tate, R. The Anniversary Address of the President to the Adelaide 
Philosophical Society. S8vo. Adelaide, 1879. 


——. The Natural History of the Country around the Head of the 
Great Australian Bight. 8vo. Adelaide, 1879. 


——. Zoologica et Paleontologica Miscellanea, chiefly relating to 
South Australia. 8vo. Adelaide, 1879. 


Taylor, J. E. A Guide to the Ipswich Museum. 8vo. Ipswich, 
1871. Presented by W. Whitaker, Esq., F.GS. 


Terquem, O. Essai sur le classement des animaux qui vivent sur 
la plage et dans les environs de Dunkerque. 8vo. ——, 1880 (?). 


Thomson, James. Contributions to our Knowledge of the Rugose 
Corals from the Carboniferous Limestone of Scotland. 8vo. Glas- 
gow, 1880. 


Thureau, G. Synopsis of a report on Mining in California and 
Nevada, U.S. A. 8vo. Melbourne, 1879. 


Trafford, F. W. C. Souvenir de ’amphiorama, ou la vue du monde ~ 
pendant son passage dans une comete. 8vo. Zurich, 1880. 


=== Homnelny Weel. 


Tribolet, Maurice de. Sur le gault de Renan. 8vo. Delémont, 
1877. Presented by A. J. Jukes-Browne, Hsq., F.G.S. 


Tullberg, S.A. Om Agnostus-arterna i de kambriska aflagringarne 
‘vid Andrarum. 4to. Stockholm, 1880. 


312 ADDITIONS TO THE LIBRARY. 


Tullbery, S.A. Om lagerfoljden ide kambriska och siluriska aflag- 
ringarne vid Rostanga. 8vo. Stockholm, 1880. 


——. Tvenne nya graptolitsligten. Svo. Stockholm, 1880. 


Ueber Versteinerungen aus den Aucellen-Schichten Novaja- 
Semljas. 8vo. Stockholm, 1881. 


Twelvetrees, W. H. Ona Labyrinthodont Skull (Platyops Rickardt, 
Twelvetr.) from the Upper Permian Cupriferous Strata of Karga- 
linsk near Orenburg. 8vo. Moscow, 1880. 


On Theriodont Humeri from the Upper Permian Copper- 
bearing Sandstones of Kargalinsk near Orenburg. 8vo. Moscow, 


1880. 


United States. Geographical and Geological Survey of the Rocky- 
Mountain Region. (J. W. Powell, in charge.) Report on the Geo- 
logy of the High Plateaus of Utah, by C. E. Dutton. 4to; Atlas 
fol. Washington, 1880. 


Geological and Geographical Survey of the Territories. Bul- 
let Voli Voy Nowa ess: 


——. ——. ——. Vol. VI. No.1. 1881. 


. Miscellaneous Publications. No. 12. History of North 
American Pinnipeds: a Monograph of the Walruses, Sea-lions, 
Sea-bears, and Seals of North America, by Joel Asaph Allen. 8yo. 
Washington, 1880. 


. Report. Vol. XII. Fresh-water Rhizopods of North 
America, by Joseph Leidy. 4to. Washington, 1879. 


Ussher, W. A. E. On the Geology of Parts of Devon and West 
Somerset north of South Molton and Dulverton. 8vo. Taunton, 
1881. 

Van den Broeck, EK. Du role de Vinfiltration des eaux météoriques 
dans l’altération des dépéts superficiels. 8vo. Paris, 1880. 


——. Mémoire sur les phénomeénes d’altération des dépdts super- 
ficiels par Vinfiltration des eaux météoriques étudiés dans leurs 
rapports avec la géologie stratigraphique. 4to. Brussels, 1881. 


Observations nouvelles sur les sables diestiens et sur les 
dépots du Bolderberg. 8vo. Brussels, 1881. 


Van den Broeck, E., et H. Miller. Wes Foraminiféres des couches 
pliocenes de la Belgique. 1" Partie. Hsquisse géologique et palé- 
ontologique des dépdts pliocénes des environs d’Anvers, par E. 
van den Broeck. Fascicule 2. Les sables moyens et les sables 
supérieurs d’Anvers. 8yo. Brussels, 1878. 


Vetter, B. Die Fische aus dem lthographischen Schiefer in Dres- 
dener Museum. 4to. Cassel, 1881. 


Victoria. Mineral Statistics of Victoria for the year 1879. 4to. 
Melbourne, 1880. Presented by the Minister of Mines. 


ADDITIONS TO THE LIBRARY. 313 


Victoria. Geological Survey. Report of Progress, No. VI. 8vo. 
Melbourne, 1880. Presented by the Minister of Mines. 


Report of the Chief Inspector of Mines to the Honourable the 
Mimster of Mines for the year 1879. 4to. Melbourne, 1880. 


Reports of the Mining Surveyors and Registrars. Quarter 
ended 31st March, 18380. 4to. Melbourne, 1880. Presented by 
the Minister of Mines. 

Quarter ended 30th June, 1880. Fol. Melbourne, 
1880. Presented by the Minster of Mines. 


Quarter ended 30th September, 1880. 4to. Mel- 
bourne, 1880. Presented by the Minister of Mines. 


. Quarter ended 3lst December, 1880. 4to. Mel- 
bourne, 1880. Presented by theMimster of Mines. 


Vogdes, A. W. Description of a new Crustacean from the Upper 
Silurian of Georgia, with Remarks Tipe Calymene Clintom. ,8vo. 
Philadelphia, 1880. 


Vogt, C. Archeopterya macrura, an intermediate Form between 
Birds and Reptiles. Translated by Prof. A. Newton. 8vo. Cam- 
bridge, 1880. 


Vom Rath, G'. Mineralogische Mittheilungen. Neue Folge. 12. Die 
Quarzkrystalle yon Zoptan in Mahren. 13. Ein neuer Beitrag 
zur Kenntniss der Krystallisation des Cyanit. 8vo. Leipzig, 


1880. 
——. Vortrige und Mittheilungen. 8vo. Bonn, 1880. 


. Ueber einen sehr kleinen schwarzen Krystall wahrscheinlich 
yon Orthit. 8vo. Bonn, 1881. 


. Ueber das St. Gotthardgebirge und die Gotthardbahn. vo. 

Bonn, 1881. 

. Ueber einige neue oder seltene Mineralvorkommnisse. 8vo. 
Bonn, 1881. 

Waagen, W. On the Geographical Distribution of fossil Organisms 
in India. (Translated by &. B. Moote.) Svo. Calcutta, 1878. 


Wagener, G., and kK. Krpping. On a new Seismometer, with a 
Summary of Observations. 4to. Tokio, 1880. 


Wallich, G. C. On the Origin and Formation of the Flints of the 
Upper White Chalk; with Observations upon Prof. Sollas’s Paper 
in the ‘Annals and Magazine of Natural History’ for December 
1880. 8vo. London, 1881. 


Wardle, T. The Wild Silks of India, principally Tusser. Folio. 
London, 1880. 


Warring, C. B. The three Climates of Geology. 8vo. Philadel- 
phia, 1880. 


Waters, A.W. Quelques roches des Alpes vaudoises étudiées au 
VOL. XXXVII. 2a 


314 ADDITIONS TO THE LIBRARY. 


microscope. 8vo. Lausanne, 1880. Presented by Prof. E. Rene- 
vier, F.C.GAS. 


Wiebel, K. W. M. Die Insel Helgoland. Untersuchungen iiber 
deren Grosse in Vorzeit und Gegenwart vom Standpunkte der 
Geschichte und Geologie. 4to. Hamburg, 1848. Purchased. 


Wigan Free Public Library. Reference Department. Index Cata- 
logue of Books and Papers relating to Mining, Metallurgy, and 
Manufactures, by H. T. Folkard. 8vo. Southport, 1880. 


Wigner, G. W. Sea-side Water: an Examination into the Character 
of the Water Supply at the Watering-places of England and Wales. 
Svo. London, 1878. Presented by W. Whitaker, Esq., F.GS. 


Williams, J. J. The Isthmus of Tehuantepec: being the Results of 
a Survey for a Railroad to connect the Atlantic and Pacific Oceans, 
made by the Scientific Commission under the direction of Major J. 
G. Barnard; with a Résumé of the Geology, Climate, Local Geo- 
graphy, Productive Industry, Fauna and Flora of that Region. 
8vo. New York, 1852. Purchased. 

Wilson, E. Introductory Lecture on Geology. 12mo. Nottingham, 
1872. Presented by W. Whitaker, Esq., F.GS. 

Wolf, H. Begleitworte zur geologischen Gruben-Revier-Karte des 
Kohlenbeckens von Teplitz-Dux-Briix. S8vo. Vienna, 1880. 
Purchased. 

Wood,C.J. Tunnel Outlets from Storage Reservoirs. 8vo. London, 
1880. , 

Woodward, H. A Catalogue of British Fossil Crustacea, with their 
Synonyms and the Range in Time of each Genus and Order. 8vo. 
London, 1877. Purchased. 

Woodward, H. B. A Sketch of the Geology of Kent. 8vo. London,: 
1881 (?). 

The Origin of Mountains. 8vo. Birmingham, 1881. 

Zittel, Karl A. Handbuch der Paliontologie. Band I. Lief. 3. 8vo. 
Munich, 1880. Purchased. 

——. ——. Band II. Lief. 2 (unter Mitwirkung von W. P. 
Schimper). 8vo. Munich, 1881. Purchased. 

——. Ueber den geologischen Bau der libyschen Wiiste. 4to. 
Munich, 1880. Presented by the Munich Academy. 


35. Mars &e. 


The names of Donors in Ltalies. 


Belgium. Commassion de la Carte géologique de la Belgique. Carte. 
Planchettes Aerschot, Anvers, Beveren, Boisschot, Boom, Heyst-op- 
den-berg, Lierre, Lubbeck, Malines, and Putte. 1880 & 1881. 


ADDITIONS TO THE LIBRARY. 315 


Botella y De Hornos, Federico De. Mapa geologico de Espana y 
Portugal. La parte geogrifica por Francisco Coello. Escala de 
somo): Madrid, 1879. Purchased. 


Dépét dela Marine. Twenty-nine charts and plans of various coasts 
and ports. 


Figari Bey. Etudes gcographiques et géologiques de l’Egypte, de la 
péninsule de lArabie Pétrée et de la Palestine. 1864. (Six sheets.) 
Presented by H. Bauerman, Esq., F.GS. 


Finlands geolog giska Undersoking. Kartbladet No. 2. 1880. 


France. Carte géologique detaillée de la France. Feuilles 9, 23, 
34, 50, 67, 69, 78, 93, 113, 122, 213 bis, 225 bis, 237. Scale sou" 
1880. Purchased. 


Jack, R. L. Geological Sketch Map of the district between Charters 
Towers Goldfield and the Coast. Brisbane, 1878. 


Ordnance-Survey Maps. Presented by the First Comnussioner of 
Works. 
One-inch General Maps. 
England and Wales. (New Series.) Quarter sheets 269, 284, 
301, 308, 314, 319, 329, 332, 333, and 334. 
Treland. Hills. Sheets 115, 140, 141, 160, 171, 184, 186- 
189, and 194. 
Scotland. Hills. Sheets 37, 53, and 103. 
Six-inch County Maps. 
Argyll. Sheets 70, 85, 97, 129, 137, 148, and 159. 


Ce 

a (Isle of Canna.) Sheet 53 and 54. 

. (Isle of Coll.) Sheet 21. 

45 (Isle of Colonsay.) Sheet 145. 

an (Isle of Muck.) Sheet 74. 

a (Isle of Mull.) Sheets 39, 53, 108, and 109. 

. (Isle of Oigh-sgier.) Sheet 64. 

(Treshnish Isles.) Sheet 80. 

Cheshire. Sheet 2. 
Denbigh. Sheets 3, 4, 6, fe 1 LU, | 12 Bay PE Wo. Ak, aici, Se). 


—— 
ead sy and 4H and 46. 
Essex. Sheets 1-8, 10-12, 18, 19, 22, 29, 31, 38-40, 42, 
45, 46, 48, 56, 58, 61-64, 69-72, 78, and 86. 


Flint and Denbigh. Sheet 7 and 8. 

Inverness. (Barra.) Sheets 59, 65, 68-70, 
3 (Isle of Burra.) Sheets 66 and 67. 
. (Isle of Harris.) Sheet 26. — 


So 
He (Isle of Skye.) Sheets 6, 8-12, 14, 22, 32 and 32, 
42, 48, and 58. 
- (North and South Uist.) Sheets 24, 25, 32, 36, 
41, 54, 56, and 61. 
Orkney and Shetland. Sheets 1, 6, 9, 10, 23, 26, 27, 28, 30, 
31, 33, 34, 36, 37, 38; 44, 49, 50, 54, and 58. 


3216 ADDITIONS TO THE LIBRARY. 


Ross. Sheets 27, 5°, 10, 11, 113, 18, 18% 24, 27, 30, 38, 39, 
AO, 43; 66,91) 917. 1002 10225 176 22 26) eos 
and 134. 

Ross and Cromarty. Sheets 5, 16, 17, 25, 26, 41, 52-55, 

pee 


a ee 
65, 76, 77, 67 and 79,92, 93, 101; 104, 1091097 andialias 
MO; DIL, 116, 113,119) 120, 123, 124. 1285 tiene 
—V—$—$——— 


and 133 and 130. 
Ross, Cromarty, and Nairnshire. Sheet 89. 
Surrey. Sheet 3. 
Sussex. Sheets 11, 13, 22, 34, 36, 47-49, 61, 62, and 66. 


: ti 
Wilts and Hants. Sheets 76 and 54. 


Preussen und die thtiringische Staaten. Geologische Specialkarte. 
Gradabtheilung 67, Nos. 53, 54, 59, & 60; Gradabtheilung 68, 
Nos. 49 & 55; und Gradabtheilung 80, Nos. 19, 20, 25, 26, 31, 
& 32. 1889. Presented by the Prussian Ministry of Manufac- 
tures, Trade, and Arts. 


Saxony. Geologische Landesuntersuchung des Komgreichs Sachsen. 
Geologische Specialkarte. Blatt 28, Grimma; Blatt 75, Langen- 
leuba; Blatt 115, Zschopau. 1880. 


Sheafer, Messrs. Diagram of the Progress of the Anthracite Coal 
Trade of Pennsylvania, with Statistical Tables &&. 1879. Pre- 
sented by P. W. Sheafer, Esq. 


—. ——. Presented by Dr. F. V. Hayden, PMLGS. 


Siedamgrotzky. Flotz-Karte des Aachener Steinkohlen-Beckens. 


Scale sa" In 7 sheets. Purchased. 


Switzerland. Commission géologique pour la Carte géologique de la 


Suisse. Carte géologique, feuilles I1V.& V. Scale jou 


Wolf, H. Geologische Grubenrevier-Karte des Kohlenbeckens von 
Teplitz-Dux-Briix im nordwestlichen Bohmen. Blatter 1-16. 
Scale .+,. Vienna, 1880. Purchased. 


Il. ADDITIONS TO THE MUSEUM. 


Specimens of Carboniferous Limestone Fossils from Flathead River, 
Rocky Mountains, 49th parallel, N.W. America. Presented by H. 
Bauerman, Esq., F.GS. 


The Type Specimens of the “ Tubulations sableuses ” of the “ Etage 
Bruxellien ” from the environs of Brussels, descrihed by H. ./. 
Carter, Esq., in the ‘Annals and Mag. Nat. Hist.’ for May 1877. 


Specimens of Tertiary Palliobranchs from South Australia, and three 
specimens of Belemnites from Central Australia. Presented by 
Prof. 2. Date, H.G.s: 


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o<}s 
7s 


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