SEA LAMPREY SPAWNING RUNS
IN THE GREAT LAKES 1951
SPECIAL SCIENTIFIC REPORT: FISHERIES No. 68
Marine Biological Laboratory
MAY15 1952
WOODS HOLE, MASS.
UNITED STATES DEPARTMENT OF THE INTERIOR
FISH AND WILDLIFE SERVICE
SEA LAMPREY SPAWNING RUNS
THE GREAT LAKES 1951
SPECIAL SCIENTIFIC REPORT: FISHERIES Na 68
Marine Biological Laboratory
L I B R yi. :R -ST
MAY15 1952
WOODS HOLE, MASS.
UNITED STATES DEPARTMENT OF THE INTERIOR
FISH AND WILDLIFE SERVICE
Explanatory Note
The series embodies results of investigations ^ usually of
restricted scope, intended to aid or direct management or
utilization practices and as guides for administrative or
legislative actioHo It is issued in limited quantities Tor the
official use of Federal, State or cooperating agencies and in
processed form for economy and to avoid delay in publication.
Washington, Do C<
March 1952
Special Scient,ific Repori. - Fisheries
No. 68
SEA LAMPREY SPACING RUNS IN THE GREAT LAKES, 1951
By
Vernon C. Applegate_, Bernard R, Smith, and
Alb er ton Lo McLain
Fishery Biologists
and
Matt Patterson
Super^i/isor, Comnercial Fishing
Wisconsin Conservation Department
UONTENTc
Page
Installation and operation of sea laxaprey
control structures in 1951 <, . 1
Lake Huron ^ ......... o ......... . 1
Lake Michigan ,,,.........,. U
Lake Superior .,.......,.,. U
Numbers of lampreys taken by control devices It
Relative abundance of sea laimpreys . . . , . 5
Lake Micl'dgan ,.,..,..........,... 5
Lake ijupc'-.^c.'r , ... ^ .,.........,.<, . b
Other species of fish taken in the weirs and traps and
degree of scarring among them, «...,,... 9
Some biological char-acteristics of the sea lamprey runs. . 9
New developments and further evaluation of mechanical
control devices, . , . o . . c 31
Barrier dams. .............. 31
Portable-type weirs and .traps ............ 33
Operating costs in 19pO and 1951™ ..... 33
Literature cited „.,„....,......,..... 36
List of common arid fr-cientlfic names of fishes mentioned
in this report .,.,.,,...... 37
ILXA.iSTRATIOKS
Figure Page
1. Map of upper Gi-eat Lalces showing the location
of sea ianprey contrcl cie:vi(^s£ operated in 1951 2
2. Rate of increase of sea lanp^r-eys in Lalces H''Ji-on
and Michigan as i"if Iecte;d hj -iireir and trap
catches in the Ocqueoc River, P;:'esqu.''^ Isle
Coixatj,, Michigan (LsJ^e Pfjron bas-in.) and Hibbard's
Creek. Door Co"anty, Wisconsin (Lake I,Iichigar>
basin) ........ 7
3. Experimental sea laaiprsy bai'/rier dam in -the Black
Ri.ver, Mackina-ar Co'onty, Miciiigan 32
U. Black River bai-rier dam. '.■:iose--'ap of bai'rier dam
shomng overhanging, o-uc-ved lip of sheet steel
attached to "wall of dam. , 32
$. Portable-type WBir and trap in Carp Creek, Presque
Isle CoTonty, Ifi-chigaii, after inst.allation of
permanent sills, ■'M'ap base, ard abutmenbs of
concrete . . . . <. 3h
6. Diagramatic plans of concrete sills, trap base, and
abutments used to protclde per:ianent base for
poi'table-tj'pe sea lampj'ey weir and trap. . o ..... . 35
Since the inauguration of the sea-lamprey investigations as a
part of the Ser^Jioe 's Great Lakes Fishery Investigations, in October
I9I1.9, consic.erabi'^. prcgr'e-rss has been made axi the long-term prcgr*am
for the d.evelc.pmer_t of methods of suppi'esof-ng or controlling the
pai-asitie sea lamprey. The sea-lamprey investigations may be divided
broadly into the folloiving phases: development and testing of con-
trol devices and procedirr-es, inelioding the acc-umulation of reasonably
exact data on costs of iD.stallation and operation of various struc-
tiiresj exir-ension cf studies on the life history and habits of the sea
lampi'ey -with a view tovra^d detei-mining better the vulnerable stages
of tiie life histo:cyj sunreys of streams to ascertain the distribution
of sea-lamprey runs and the extent of available spamiing grounds; and,
studies of species subject to attacks hj sea lampreys to learn the
incidence of attacks and the effects on abxmdance.
Selected from the pre:;ed±ng investigative program for inclusion
herein az'e summarizations of data collected in 19^1 concerning: a
second yeai' of experimental control operations in Control Zone H-1
(in nort.hem Lak:e H\iron) and in the Wisconsin -maters of Lake Michiganj
abundaiice of sea lampreys in the thi-ee upper lakes; a comparison of
the biological chai'acteristics of the sea-lamprey spatvning runs of
19>1 with these of pi'evious years; and developments and further eval-
uation of mechanical de'/ices for sea-lamprey control.
Similar data for the 19^0 season ar.d information basic to this
report have been presented by Applegate and Smith (19^1).
These particular operations and investigations were conducted as
in the previous year, W3.th the cooperation of the Wisconsin Conserva-
tion Departmenb and the Michigan Department of Conservation.
Installation and operation of sea-lamprey-control structures in 19^1
Lake Huron. — In nort-hera Lake Ku-ron 12 trapping devices "vvere
operatea in," Conti-ol Zone H-1 "srfiich was established in 19^0; 11 of
these sti-uctui-es were operated in the same streams as in the previous
year* (Applegate and Smith, 1951). In addition, a trap was installed
in the bottom compartment of the fish ladder at the paper-mill power
dam on the Cheboygan. River (fig. 1). All installations were the stan-
dard, po;rtable-type sea-lamprey weir and traps with the exception of
the pe:o.nanent"type Ocqueoc Ri.ver installation and the aforementioned
Cheboygan River trap which ivas a de-'rlce manufactured especially to
fib a fish-ladder Gompar-'&ment. Complete runs were captured in all but
tvro streams, and in one of these only a minor escapement occurred. In
the Cheboygan River only a small part of the -bobal run was taken be-,
cause the trap could n.)t be located near the main spill of the water
and consequently the majority of the lampreys were attracted away from
the trap. However j the dam prevented the upstream movement cf the
lampreys not taksn by the trap.
Figure 1. — Map of upper Great Lakes showing the location of sea-lamprey control devices
operated in 1951.
Table 1. — llyiinljc-' of spaTcning-CTin sea lampreys talcen by
control devices during the 1951 season
[Structures listed belotv may be located on map in figure 1]
Stream Number taken
Lake Hui'on tributardesi
(Control Zone H-1)
Car-p Lake River^ Eranet County, Michigan Ij.3 913
Li.ttla Black River 3, Cheboygan County, Mich. 909
Cheboygan Rive^', Cheboygan Coiinty, Mich. 2,368
Elliott Creek, Cheboygan Coui:^ty, Mich 70
Green Creak, Cheboyg.an Coixnty, Mich 78^
Lone Pine Creok, Presque Isle County, Mich 0
Mi.lliga:i Creek, Presque Is3.e Ccimty, Mich 5^7
Cedar Creek, Presque Isle Co^jjity, Mich 0
Grace Harbor Creek, Pi'esque Isle CoTinty, Mich 32
Carp Greek, Presque lole Coanty. Mich 1^ 266
Ocqueoc River, Pr.jsque Isle County, Mich 19,393
Trout River, Fre.'sque Isle County, Mich 1,903
Total, Lake Hiiron. 32, I7I
Lake Michigan tributaries:
Hibbard's Creek, Door County, lis , . 12,6i;0
lily Bay Creek, Door County^ Wis 128
Three Mile Creek, Ke-sra-onee County, Wis 2,l407
Kewaunee River, Kewaunee County, Wis 3, 270
Mishicot River. Msnitowoc County, Wis 21,080
Fischer Creek, MaaitoTroc County, Wis , 3jl4.5>
Total, Lake Mich?.gan. l!.2,980
Lake Superior tributaries;
Pendill > s Crselc, Chj-ppeiva County, Mich 20
Ghocolay River, Ma'rquette County, Mich 3 01
Total, Lai'Te Superior. 321
GRAND TOTAL. 75j,U72
An electromechanical iveir -was i:i£"'oalled below the Ocqueoo Tdver
Trveir and was operated and tested contiiriously for 6 weeks during the
height of the upstream migration of ser. lampreys, l/
The objectives in operating thic Control Zone another yeai' are
sttrnmarized briefly as follo-vTs:
(1) To gain additional experisnos in the operation of this
type of control and to obtain information on ajdmin-
istrative and operationcJ. p:'obleras and costs.
(2) To ascertain further the e/Veots of the prevention of
reproduction by sea lampreys in the streams tributrr;'"
to a limited area of shorel:_."ic .
(3) To continue the development and testing of improvements
in design and construction of meclianical control struc-
tures ; and
(U) To provide sites where adcqr.ate checlcing devices (T/eirs
and traps) were present foi 'costing other equipment,
primarily of an electrical nature.
Lake Michigan. — Six pc)rtable-t3n.)G r/cirs and traps were agaiji in-
stalled and operated by the Wisconsin Conservation Department in
streams tributary to Lalce Llichigan. Eocause of high waters these de-
vices were installed late and conseqv.<:)is.t?^'- some escapement occiirred,
but the majority of the sea lampreyo entsx'ing these streams ivere cap-
tured. Two structures were relocated to eliminate spawning which
occurred in areas below v/eir locationt: used in 1950.
The checking weir and traps luxLv. in the Black River, Mackinaw
County, Michigan, was installed at a :ic?a- location above the barrier
dam in that stream for operation by personnel of the Michigan Depart-
ment of Conservation. The purpose of these structures was to deter-
mine the effectiveness of a special!;''' dojiigned, low-head barrier dam
in blocking upstream movement of spai.rning-run sea lampreys. The oper-
ation of this unit was continuous tiu-cughout the season. Although a
large run entered the river, no lamr)i'c-)7s vrere taken in the checking
weir. The barrier dam was completely effective in blocking the migrants.
Lalce Superior. — In the Lalce Superior basin the weir and trap in
PendiHTs Creek, Chippewa County, viras operated for the second yeai" and
captured the entire run. An electrical fish screen and a portable-type
weir and trap (checking vreir) were operated in the Chocolay River,
Marquette County, Michigan.
Numbers of lampreys talcen by co:-trol devices. — A total of 7i^j^72
spawning-run sea lampreys was captui'Ou in 1951 in 21 control devices.
In nearly all streams, the entire spasming runs ivere captured. Of the
1/a detailed report of the developmeu'; of electrical and electromechan-
ical sea-lanprey-control dev:.ces T.i.11 be presented elsevriiere.
total catchj 32,171 individuals were taken in Control Zone H-1,
I|.2,930 were captured in the Wisconsin control devices, and the remain-
ing 321 lampreys were taken from the two streams tributary to Lake
Superior. Biological data were reviorded for many of these lampreys 5
all individuals were subsequently dcjstroyed. These catches ccre siim-
marized in table 1 iiriiere the jJidividvAi totals by strean and by iaK:e
basin are given.
Relative abundance r.f sea lampreys
Lake Huron. — The sea-lamprey population in northern Lalce Hurori., as
Indicated by the size of the spawnlig runs captured, apparently conbin-
ued to maintain itself at the pealc level of its abundance ior a:tiother
season. The total run in the Ocqueoc River was 19,393 sea lampreys in
the 1951 season as compared to 18,822 in 1?50 and 2l|.,6l4.5 in 19U9. As
in 1950, a considerable number of migi'ants from the adjacent lake ar-ea
was "siphoned-off" by trappijng operations in other streams in the virgin-
ity; this reduced the total catch in the Ocqueoc River to a cer-^-.ain ex-
tent. Consideration of all factors would indicate that, numerically
spealcing, the runs in the three seas./ns were of comparable sise.
Most of the catches in the snail .streams of Control Zone H-l were
considerably less than for the previ.ou3 year. At first thought this de-
crease would seem to indicate a declij.ne in the sea-lamprey population.
Actually, these small catches were the I'esult of the blcoking of the
stream mouths by sand ba::'s sevei-al t?jr.es during the period of upstreejn
migration. High lake levels, ?-ow stream disch.arges, and strong w3.nds
all contributed to unusual bari'ier-ba:'" formations during the 19^1 season.
All available records of spaivirLig runs of sea lampreys into the
streams of northern Lake Huron (Uriited States -viraters) arre assembled in
table 2. Those records for the Ocqucioc River demonstrate the phenomenal
increase in the population in the yea:;'f 19W-i- to 19li.9 and the st!.bsequent
leveljjig-off of that population -when fish stocks in the nci'^/heiT. areas
of the lalce were reduced almost to the point of disappeai'anca (fig. 2).
Lalce Michigan. —In the streams ti'ibutaj:y to noi'thwfistem La.ke Mich-
igan, ~tvelr"~an3~trap catches coutinuec. to ?.'eflect the explosive increase
of the species in these waters. In 19^1, sea-lamprey spaTjning runs in
six Wisconsin streams were nearly thiree times as lai-ge as those dntei--
ir.g the same streams the previous yea::'. In 19^0, I6jli.l0 spawning mi-
grants were taken in six control devices; in 1951^ U2,980 inoAi^-iduals
vrere captured. All available records of spawning runs entering thes'5 six
streams are presented in table 3« The spa'swriing runs captur^sd :ji Hibbards
Creek, Door County, Wisconsin demonstrate most dramatically the enonnous
increase in the nujabers of sea lampreys in the lalce since 191;.'? (babla 3
arid fig. 2).
The data collected in 19^1 give no indication that the sea lamprey
population in Lake Michigan has yet c>tf.ained the peak of its ab-o:adancft,
Maxjjn'om abundance and a leveling-off in tiumbers of the lam-prey popula-
tion in northern Lake Hui-on followed by several years the vix'^mal
Table 2. — Nura'ber of spawxiing-run sea lampreys oaken in 7?eirg and traps
in strsains tributary to Lalce Huron, '~l9^ - '1951
Strean
Year
19hk
19kS
19lt6
1917
-lo;,;!
19U9
19502/
1951
O'-queoc River
1/ 3,366
1/U,603
iAo^OOO
i/i3,ooo
i/2l;,6.U3
13,622
19,393
C.arp Creelc
• • •
2/ 1,617
2/ 2,939
2/ 2,763
1,161
1,266
Trout River
• « •
• • •
1,702
1,903
Grace Harbor Creek
52
32
Cedar Creek
• • •
0
0
Mill'' gaia Creek
700
527
Lone Pine Greek
0
0
Green Creek
l,9l;<
785
E'^liott Creek
266
70
Little Black River
• • 9
9$3
909
Cheboygan River
• * d
• • c
• • w
2,368
1/ Shetter (19U9)s partial capture of run; examination of Shetter's data suggests
~ tloat these catcLes represent about three-quc,rters of the run entc-:ring the stream
each year.
2/ Applegate (1950): data for Ocqueoc River for 19U7 and 19l;3 aa'e estimates based
~ on counts of total number of nests in watershed mth consideration given for
observed sparmji-ng habits a:id sex ratio in those years; other data are based on
entire runs captured in -weirs and traps.
3/ Applegate and Smith (1951): all data based on entire runs captuz'ed in iveirs
~ ajid traps.
000
-
600
-
400
—
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Table 3. — tlumber of spavTning-rua s'iZ. Israpreys talcen in '•^i£f> ^^-
traps in streams tributa:-"/ -'co_ north-western Lal-ce
lEchigan (19ii?~- 1951}
Tear
Stream
19Ub^
19l;6
I9U7
19.'+6
I9ij.9
19<0
193'1
Hibbard Greek
1/25
125
^96
S^9
1.579
5, 1' 22
12s6L.O
lily Bay Creek
• • •
• • «
0 « e
a A 9
16
128
Three Mile Creek
• • •
• • «
• • •
• 0 •
e • 4
1,051
2,1j.07
Kewaunee River
• • •
• • •
• • •
a • «
I5353
3,270
ICshicot River
• • •
• • «
0 • 0
» e «
7,712
21,080
Fischer Creek
e • •
« • «
81:7
'^s'S't.'
1/ The number of lampreys trapped in 1.9.45 is nc^t the complete laxi. into
~ the streams trapping operations ware intermittent in that year.
disappearance of the lake trout (a preferred prey species) froiT: the com-
mercial fisherj^. Maintenance of the population in Lake Huron at peak
abundance is attributed to the ability of the remaining fish stocks to
support, at least temporarily, the peak lamprey population. Those spe-
cies to which the lamprey has transferred its attentions are currently
suffering a severe decline. This same situation may apply to Lake Mich-
igan.
Lake Superior. — The sea-lamprey population in Lake Superior continues
to increase, A recheck of streams, tributary to the eastern third of
Lake Superior,, in which evidence of spawning activity was noted in 1950
revealed considerable increase in spawning activity in the 1951 seasonj
several streams in which no activity was observed in 1950 contained evi-
dences of spaivning in 1951.2/ The niimber of spavming mLgrants taken in
at least one of the two experimental control structures operated in Lake
Superior tributaries indicates that ■y^rj effective (productive) spax-raing
runs are even now present in the most suitable tributaries! the progeny
2/ A comprehensive report of a siorvey of the streains tributary to the
south shore of Lake Superior which was made in 1950 and 195l is now in
press.
of these runs, when they enter the lake some years hence, will be numer-
ous. Further surveys of tributaries of the lake conducted in 19^1 indi-
cate that extensive^ but as yet unused, spawning grounds of something less
than optimum quality exist for the species at least on the south shore of
the lake. A considerable expansion of the population, therefore, appears
imminent. Adequate warning of the effects of such an expansion upon the
lake trout and other commercially valuable species in the lake may be
found in the present condition of fish stocks in Lakes Huron and Michigan.
Other species of fish taken in weirs and traps
and degree of scarring among them
Counts by species were made of fish entering 10 of the weirs and
traps in Control Zone H-1 and in Pendill's Creek which flows into Lake
Superior. In addition to the sea lampreys captured, a total of 79,091
fish was taken in 10 streams in Control Zone H-I3 307 fish were captured
in Pendill's Creek (table I4) . Data were also collected on the numbers
of lamprey-scarred suckers of several species taken in 7 streams in Zone
H-1 (table 5) . Records of scarring were collected for other food and
game species but these records were generally incomplete or based on too
few individuals to warrant inclusion here.
From the data available it is difficult to say whether the food and
game species are still declining in northwestern Lake Huron. Trap records
indicate a stabilized condiLion might have been reached. 3_/ However, com-
mercial fishermen report that fewer suckers and other food species were
taken in their nets in 19^1 as compared with 1950. Furthermore, the inci-
dence of scarring at least among the suckers, continues to rise. For
example^ in 195l, 3U.6 percent of the suckers entering the Ocqueoc trap,
as well as those collected in o\ir nets, were scarred. This compares with
30.0 percent in 19^0 and 25«5 percent in 19ii9 =
Some biological characteristics of the
sea lamprey runs
Nearly all of the sea lampreys taken in eight streams in Control
Zone H-1 were examined to detennine the sex of the individuals (table 6);
similar records were made for all sea lampreys entering one tributary of
Lake Suoerior, Examination of these data collected in 19^1 indicates
that the sex ratio of entire runs in northern Lake Huron continues to
3/ It might be observed here that any further decline of, for example, the
"suckers below the levels indicated by the weir and trap catches in the pre-
ceding year^ 19^0, would have required the near disappearance of this
species from adjacent areas of the lake; see Applegate and Smith (195l) •
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11
Table 6.- — Sex ratio of sea-latnprey runs in eight trib-gtaries
of nortHem Lalce Huron during 1951 season
Stream
Total
catch
Total for
which sex
detem' ned
Number
of
males
Kuniber
of
females
Ratio
of males
to
females
Trout River, Presque
Isle County
1,903
1,903
l,37i^
528
260:100
OcqueoG River, Presque
Isle County
V^,393
19,322
13,9li9
5,373
26C';100
Carp Creek, Pr-esq\ie
Isle Go-'jnty
1,266
1,266
901
365
21;?: 100
Grace Harbor Creek^
presque Isle County
32
32
2$
7
357:100
Milligan Creek, Presque
Isle Ooiiiity
527
'621
367
160
229:100
Green Creel", Cheboygan
County
785
783
$12
211
271:100
Elliott Greek, Cheboygan
County
70
70
52
18
239:100
Little Black River,
Cheboygaii County
913
913
S\6
268
2lil:100
To sal
2U,889
2li,8l6
17,086
6,930
258:100
12
shift toward a higher percentage of males. The rate of change^ however^
has dimirdshed appreciably from that displayed in the two preceding yeears
This point is illustrated in the following records of the sex ratios of
entire sea-lamprey runs entering tributaries of northern Lake Huron dur-
ing the past 5 years k/t
Sex ratio
19li7o = o . o o l65 males s 100 females
19i;8. ...... o .... - o 169 males s 100 females
19^9.... 211 males ; 100 females
1950.............. 252 males s 100 females
1951=... .^..o 258 males i 100 females
The sex ratio of the run entering Pendill's Creek in the Lake Super-
ior basin was 110 males s 100 females j the run in that stream in 1950
displayed a ratio of 111 males ? 100 females. This proportion of males
to females among the spawning runs appears indicative of a rather re-
cently established population „ Judging from what has occurred among the
sea lampreys in lake Huron^ it is likely that this ratio will shift in-
creasingly in favor of the males if the population increases to the
levels of overabundance attained by the species in Lakes Huron and Mich-
igan « The reasons for these striking shifts in sex ratio with increas-
ing population density are a mystery,
Indiiddual lengths and weights of sea lampreys were recorded accord-
ing to a predetermined sampling schediole from the runs in Carp Creek and
the Ocqueoc Rivers 149.7 percent of the Carp Creek run and 22.0 percent of
the Ocqueoc River run were measured and weighed 'tables 7j 8^ 9s ^1^"^ 10).
The range in length of U,899 migrant sea lampreys, sexes combined^
that were measured in 1951 was 10,7 to 23.7 inches. The range in weight
for the same specimens was 32 to uOO grams (1.1 to lli.l ounces) . The
average size, sexes combined^ differed slightly between the two runs stu-
died. The average total length was l5.8 inches for the Carp Creek indivi-
duals and 16,2 inches for the Ocquecc River sample. The mean weight of
sea lampreys taken in Carp Creek was ll5,6 ^^rams (U.l ounces) while mi-
grants from the Ocqueoc River averai,ed 132.5 grams (Ii.»6 ounces).
Comparison of the preceding averages with similar data collected
since 19U7 shows a definite diminution in the size of mature spawning
migrants in northern Lake Huron tributaries (table 11) , For example^ the
k/ Where data for runs in more than one stream are available in any year^
an average has been obtained for the combined runs.
13
Table 7."-Lengi^
h frequen
;i53 01 se;.
. lampieys
collen-oed.
DjTi Carp Cr
eel-c ana
the Ocqueoc Rive.r, PreLque
Isle GoTino
y, Michigan, in 1951
Midpoint of
len^.h group
(inches )
Carp Greeic
Ocqueoc River
Males
Females
Total
Males
Females
ToT.al
10.7
• • •
-
« • e
e 0 «
• • «
.9
• • «
• • «
1
• f e
1
11.1
• • o J
• e e
• 9 «
• • o
* • «
.3
. 0 . ... I
« • •
• e c
• • e
.5
• ■ o
» <. <«
1
1
2
.7
• • «
• • a
1
X
.9
• ■ o
• • O
2
• • «
2
12.1
1
J_
2
5
• • •
5
.3
2
• • •
2
1
2
6
.5
1
1
p
li
1
15
.7
1
...
, '^
17
2
19
o9
)•
',■)
6
26
1;
30
13.1
6
2
3
he
12
58
.3
9
12
h$
7
52
.5
19
1
20
62
11
73
.7
12 1 \
.'. S
70
25
9$
.9
15
2
17
92
20
112
ll;.l
13
J
21
125
36
161
21
^ 1
118
hh
162
.5
2L
i
31
162
W;
206
•-7
* [
26
10
'-"o
118
lil;
162
.9
"7
1
2u
123
35
158
15.1
21
12
' ■■1
-■J
r,2
56
228
.3
18
"
25
131
U7
178
28
12
1.0
165
54
219
.7
36
12
I:.8
121
49
170
.9
11
13
^1
104
I;2
l)i6
16.1
13
7
20
97
U7
llih
.3
12
1
19
116
Uo
156
.5
2C
3
1C7
1.3
150
.7
16
h
20
76
111
117
.9
6
6
l;^
66
33
99
17.1
15'
6
83
38
126
.3
12
2
71
41
112
.5'
Ul
-'
12
82
li6
128
.7
7
li|
49
37
86
.9
3
1.
>^
49
32
81
18.1
U
9
$9
•-,2
91
.3
U
2
6
57
27
81;
.5
li
3
7
61
35
96
.7
1.
9
k9
21
13
.9
2
£
S
34-
1
25
S9
lU
Table ? (continued)
Midpoint of
Carp Cree
k
Ocqueoc River
leneth eroup
(inches)
Mal(
2S Females
Total
Males
Females
Total
19.1
1
i 3
7
53
25
78
.3
I
1 U
8
26
25
51
.5
t
? 1
6
37
19
56
.7
L 2
3
25
18
li3
.9
• •
k • • o
tt • 9
25
12
37
20.1
? 1
3
19
lii
33
.3
• •
3
3
13
8 •
21
.5
009
2
18
10
28
.7
• •
2
15
7
22
.9
1
2
6
8
m
21.1
• • 0
1
3
5
8
.3
• • «
« • •
1
I4
5
.5
• • •
1
1
2
.7
• OB
2
2
u
.9
■ « •
3
1
li
22.1
• • •
• • «
.3
e • •
.5
• • •
.7
.9
• • •
23.1
• • 0
.3
• • 0
.5
« • •
.7
• • •
1
1
Total
w.
> 18U
629
3,033
1,237
1,270
Mean length
is.t
S 16,2
15.8
16.0
16.6
16.2
Standard
deviation
+ IM
3 + 1.7li
+ 1.70
+ 1.87
+ 1.93
+ 1.91
15
Tabj-e S. — ^Yifeight frequency of sea lamprey.- collect eo. in 0££p G^eek and
the~0cque"oc River ~Fresqtxe Isl^ County, Michigan in 1951
Weight
Carp
Greek
/•
Dcquecc River
1 'rTr.p''*'i;'p ■
(g-.'am.s) Us
Lies Feir
lal
.es Tot;:'2
Mal-;s
Females
Total
30 - 39
1
• «
-
• • a
• • •
• 6 0
ho - h9
3
• •
'."*,
2
7
SO -59
6
2
u
33
'7
1
Uo
60 - 69
27
6
33
122
21
li-U
70 - 19
56
9
65
259
74
333
80 - 89
66
2lj
\ 90
337
87
U2I:.
90 - 99
5U
23
11
373
ni
U8li
100 - 109
53
23
8.:
333
102
435
TLO " 119
39
li
55'
233
85
318
120 - 129
3$
in
ii6
22k
81
305
130 - 139
23
11
3^
165
86
251
]i;0 - lLv9
lU
s
23
135
66
201
150 " 159
13
16
121..
57
181
160 - 169
3
I
t 12
i:.o
50
160
170 - 179
5
e
13
86
60
:iii6
180 - 189
9
c
' lo
100
56
156
19c - 199
1
4
I
1- !-•
70
L9
119
200 - 209
2
i 5
60
U
101
210 " 219
u
■/
' 11
59
33
92
220 - 229
3
■
1,
52
"7
89
250 - 239
9
J 5
38
33
^i
2li.0 - 21^9
i
c
> 6
32
24
5S
25c - 259
2
]
3
28
27
55
260 -- 269
h
• 3
* +
23
13
36
270 - 279
a «
]
1
15
9
24
28c - 289
• •
]
L 1
-3-
ri
15
290 - 299
* •
]
L 1
5
5
11
300 - 309
• e 4
• «
• £ 0
2
• • •
2
310 " 319
« «
• ,
3
''-
5
"20 -• :-.29
• • *
, 4
1
• * 0
-1
33c - 339
• • <
• 1
n_
2
3
3hO " 3h$
• «
0
• • •
• • •
350 - 3S9
» • •
« <
. 0
• • •
• • •
360 - 369
> e •
» • <
• «
• • •
» ■ •
370 - 379
k • •
1 • <
.. ,
1 • • •
« v «
330 - 339
» • «
•
• • •
390 - 399
» • •
» •
. .
• « •
• •* •
Uoo - 409
» • •
> •
k 0 •
1
1 ^
<. . .
Total.3 \
4Jt2 :
L8:
I 624
3,033
1237 i U,270
IU6.9 132.5
Mear: -yjeight 11(
3.8 12(
3.:
3 115.6
123.3
303X1(13111
' \
de7^.ation + U
i.U + h\
?.<
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+ 51.3
1+ 57.0 5 + 53.1
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CMr^_:^"LAMDr-;CqONpCM
iHHHHrHHrHHCvJ
■-3
-3
rHCMrni_:j\AMDt^coCTsOrHr^
CMOJC^OJCMCMCMCMCMr^fA I
H
CM
•-3
XA O VAIA-LAIA •
iH rH O CO ij\ c:>
rH CM r^_^XAvO I
+3
to
+3
CO
5
<
-3;
vO
CM
fA
•
CM
r-i
H
fA
O
CO
(U
ho
CT3
u
!>
U
O
CO
H
fi3
+^
o
25
Table 11. — Average lengths aiid average weights of samples of
sea lampreys taken in Carp Creek and the Qcqueoc RLver.,
Pre s que Isle County, "^^higan, by years, 19U7-19J1
Stream and year
MaZ
-63
Females
<^cf and ?2
Average
Average
Average
Average
Average
Average
length
weight
length
weight
length
■weight
Carp Creek:
19i;7y
17. U
131.6
17. U
186.6
17..^
• • •
19181/
16.7
« • •
16.9
• • •
16.8
• • «
19U9i/
16.9
• • •
17»U
• « •
17.1
• • »
1950
I6,h
• « •
16,9
* • d
16„5
• • «
1951
15.6
110.8
16.2
128.3
15.8
115.6
Qcqueoc River:
19U7i/
£/l6.2
« • •
i''l6.3
« • «
• e •
o • »
19U9i/
17,0
• • •
17.2
• • •
17.1
« • «
1950
16. U
• • •
16 = 7
• • •
16.5
r • •
1951
16.0
123.3
16.6
m6.9
16.2
132.5
1/ Applegate, 195C.
2/ Sample selective for smaller individuals; see Applegate (1950).
26
Table 12, — Daily minimum, maximum, and mean water temperatures (°F.) and water
gauge readings (feet) fc
r the Ocqueoc River
• (Presque Isle County, Michigan)
ivith mean air
temperature and wind ax.d we
ather records for one looa±xo7.
April 12 ■- August"
6, 1951
Date
Water temperature^:/
Water
0 /
Mea_n air
tempera-
Siqr
Weather
1951
Min,
Max.
Wind
Mean
gauge£/
tare
April 12
iiO
U0.5
Ui.
1.7
ii2,5
Overcast
Rain
Light
13
liO
Jil,0
ii2
2.U
ii3.0
II
Lt. rain
ft
Hi
lo
lil.O
li2
2.8
Lo.o
II
Fair
11
15
Uo
Ui.o
li2
2.5
kl,0
M
Snow
Moderate
16
33
39.5
liO
2.2
38.0
It
II
Li^t
17
39
39.5
liO
1.9
40,0
II
II
11
18
38
38,5
39
1.7
3S.S
II
Fair
II
19
38
39.S
hx
1.6
L;6,0
Ptly. o^cast
II
Calm
20
37
3Q.S
iiO
1.7
h2.S
Clear
II
II
21
37
38.0
39
1.7
y.^s
Overcast
Snow
Light
22
39
39.0
39
1.5
IiJ.O
11
Lt. rain
ft
23
38
UO.O
U2
1.9
hS6
Ptly. o'cast
Fair
Strong
2U
39
iiO.O
lil
1.7
iut.O
II
II
Light
25
Uo
iiO.5
Ul
1.5
i£.v9^
Overcast
Fog
Calm
26
iiO
U2,5
U5
1.7
5o.o
Clear
Fair
Light
27
JiO
ii3.o
U6
1.8
S3S
II
II
If
28
li5
li8.5
52
1.7
71.5
II
II
It
m
li8
50.0
52
1.7
60.5
Ptly. o'cast
11
ft
30
51
53.5
56
1.5
SS.o
Clear
It
Moderate
May 1
S3
SS,S
58
1.5
62. S
Clear
Fair
Light
2
9x
56.0
58
1.5
60. S
Ptly, o'cast
II
II
3
53
51i.o
SS
i.U
50.0
Overcast
Lt. rain
It
U
S3
SS^S
58
1.5
55.0
Clear
Fair
II
c'
S3
55.0
57
1.5
56.5
Overcast
It
Calm
6
S3
ss.s
58
1.5
56.0
Ptly. o'cast
II
Light
7
52
SS-S
S9
1.7
60.5
Clear
II
Moderate
8
S3
Sl.S
62
1.7
67,0
Ptly. o'cast
II
light
9
Sh
57.0
60
1.6
Sk.S
Overcast
It
It
10
S3
56.0
S9
l..)4
iiS.O
Clear
It
"
11
S3
56.0
S9
1.5
53,0
Ptly. o'cast
H
It
12
52
56.0
60
1,5
S2,S
Overcast
II
II
-
13
51
5ii.5
60
1.5
ii7.0
Clear
tl
II
Hi
S3
58,5
Sh
1.5
63.5
Clear
II
II
15
58
61.5
es
1,5
67.5
Ptly. o'cast
tr
Moderate
16
57
59.0
61
1.5
51.5
Overcast
Lt. rain
Calm
17
56
60.0
61i
i.li
h.9.0
Clear
Fair
Light
18
S6
60.5
es
i.li
55.0
Ptly, o'cast
II
It
19
S9
63.5
68
i.ii
66.0
Clear
11
II
20
62
61i.o
66
1.5
64.5
Ptly, o'cast
II
II
21
61
65.5
70
1.5
63.0
II
II
It
22
S9
61.0
63
1.5
50.5
Overcast
II
Calm
27
Table '12, (continued
Date
Water
temperabrjT'sV
Water
Mean air
tempera-
±v:rz-.
tir_ . j_ t
Wind
1551
te-r_.
Mean
Wiir.
SKy weaklier
May 23
57
62.0
67
h9.$
' L/lo<ir
Fair-
Light
2li
60
63.5
67
1.?
66.b
Overcast
(!
!l
25
60
65.0
70
1.6
70.5
Clear
f!
Moderate
26
63
66.5
-L.O
t'i.O
Pt>lyo o'cast
II
li^t
27
63
65.0
o7
J. 6
c-r. ::"
Over:;ast
•'
It
28
62
63.0
6U
1.6
52.0
Overcast Lt. rain
Cfl'iTn
29
60
6Ii.O
68
1.6
ec'.o
Pt'ly. o'cast Fai.x-
Light
30
60
66.0
7i
1.5
6$.0
Clea:r' _
tJ
IT
31
61
65,0
69
1.0
$Q.$
Fcly. o'cast
tl
t(
June 1
61
63.0
65
1.6
Overcast
-
Fai'"
Calm
2
60
62.0
6k
l.£
53.C
«<
It
Moderate
3
60
62.5
6$
1,6
i!
1;
Light
1
4
58
6C.5
63
l.V
k:.^:
-
II
It
5
56
61.5
67
- c
-i-. ■ ^
51.0
G1&9V.
tl
«f
6
58
62.5
67
1.7
5:^.5
1:
•1
tl
7
58
62,$
67
-—• i
50.0
Ptay. 0 ' cast
tl
It
8
59
64.0
69
1.6
51.5
Cle.d::' .
tl
It
9
60
6i.0
62
1.6
17.5
O^i^ercasT.
ft
Calm
10
59
63.5
68
$e.$
Clear
!l
Moderate
IT
57
62.5
68
J. 7
^l.O
n
Ij
Li gilt
12
^9
63.0
67
•
Ptly. o'cast
It
"
13
61
6$.''.
70
1.6
57.5
Ov&rc-ast
II
IJ
"III
60
65.5
71
1.0
53.5
Cle^^r
"
ii
15
61
66.5
72
1 "
61-.5
tl
IJ
tl
16
63
67.5
72
1.6
67. C
"
"
ii
17
65
69.5
71'.
1.6
'■:-| c*
■ — e -.
Hazy 1 "
It
18
67
70.5
7ii
1.7
72.5
n \ V.
tl
19
68
69.0
70
-. •-.
6r:.0
Overcast '.'
Ca'lm
20
67
69,$
72
1.9
l''7 u
If {{ ii
Mcxisrate
21
6U
66.5
69
2,1
60.0
Cle:^::- j "
Light
22
6)4
66.5
69
... ,
57.C
Overc'-ast | I'
fi
23
63
68.0
73
-; P
65.^^
Gleai" "
tl
21;
6^
63.0
71
1.9
4.5
Glc-aJy Raiu
I!
25
63
67.5
72
1,8
Clear Fai v
'A
26
66
67. c
68
• • •
60 0^
... . . -
. 0 .
27
65
69.0
73
1.8
64.0
Clear Fai-.'
Light
28
66
68.0
70
1.9
62.5
0^7 er cast
CI
It
29
6U
68.0
72
1.9
63.0
Cl^aa-
ft
Moderate
30
66
69.$
73
1.9
6,.--
5!
:i
Light
July 1
6^
68.5
72
Clear
Fa,i-.-
Ii.ght
2
6ii
61;. 5
•v^
'^-^
63.5
n
.1
•1
3
6L
66.$
69
i
.1. 8
.^■7.5
ft
fi
n
26
Table
Date
Wat ex
temper
atureV
Water
Mear- air
Weather
Wind
1951
Mir.
Mear.,
Ma;:,
gauge^/
temptira-
toj'e
Sky-
Ju]y k
63
6h,^
66
1.9
5i-.o
Overcast
Lt, rain
Ught
^
61
6i;.0
67
?..l
60.0
Clear
Fair-
:t
6
61
65.5
70
2.1
67.0
ti
It
(I
7
6^
67.0
70
2.1
67,0
Overcast
It
1!
8
66
68,5
71
■?'.., 0
II
It
it
9
67
71.0
75
?:.3
71.0
It
II
It
10
6''
70.0
73
2.0
63,5
It
!l
n
11
66
68.0
70
2.0
57.0
Overcast
Fair
CaJjn
12
66
70,0
7ii
2.0
6l.O
It
It
It
13
66
71.0
76
2.1
(6.0
Clear
II
Light
11;
68
73.5
79
2.1
■;U.5
It
II
Cfl'im
15
70
74.5
79
2.1
73.5
It
II
li^t
16
69
71.5
7k
2.1
99.S
Overcast
tt
n
17
67
71.0
75
2,0
01.5
Clear
It
81
18
69
72.0
75
2.0
69.0
Ptly. 0 ' cast
It
tl
19
66
68,5
71
2.0
61,5
It
It
tl
20
6U
69,0
71
2.2
63.5
Clear
It
II
21
6?
69o0
71
2.2
60 0 5
Overcast
Rain
Strong
22
67
70,5
Ik
2.1
65.5
Ptly. o'cast
Fair
Moderate
23
65
70,5
76
2.1
63,$
Clear
II
light
21;
68
73.0
78
2,1
69.5
II
It
II
25
71
7i;.5
78
2.2
-{^■^^
Ptly. o'cast
It
Moderate
26
73
76,5
80
2.3
7^.5
Clear
II
Light
27
69
71.5
7)4
2.1
60.0
Ptly. o'cast
It
ft
28
68
73.0
78
2.2
6k. 5
Clear
II
II
29
70
7U.0
78
2.2
73.5
Ptly. o'cast
n
II
30
72
7U.5
77
2.2
7U.5
11
It
•1
31
72
75.0
78
2.1
YCO
It
II
II
Aug. 1
67
71.5
?6
2.2
^r-''
Clear
Fair
Moderate
2
69
71.0
7?
2.1
68 a 5
Ptly. 0 ' cast
It
Light
J'
67
70,5
7U
2.3
56.0
«
It
Strong
h
61
68,5
73
2.2
9j,o
ti
tt
Calm
K
65
69.5
7li
2,2
Clear
tt
II
6
67
68.5
70
2.3
6Jl.p
Overcast
tt
Light,
1/ Thermograph station at -vveir
2/ Water-gauge readings are absolute depths in feet across the deck of weir
TaLle 13."
jAStailLation BX'd 'jpei'-ation of five tuiits of e>q?erir-entaJ
control stru'c^ui'3F~'i^~-at9d~in 1950 and 1951 1/
'IW'
i9Fr
Operational -unit
IrJ.-bial I;
i installLation i Annual
acd repa^.r jj operation
Reinstallation
and renair
Annual
operation
I
.1 " Trout River gi-oup
(1 control struct'^-ire)
2 - Ooquuoo - Gai'p Croek g:.-oup
(2 r.r.ntrcl staructuT'e--;)
3 •■ Cheboygan group
(8 oontrcl sti.-'u.ct.ir^es)
U - Gaz-p La>:e Rive::- group
(1 control stn".ctui-e.)
Control Zone R-H.
(12 control stractures )
Sub'-tota-L
S " Pendills Creek gjroup
(1 stinKAvx-s)
2/
- lJ-,721
1,951
17,i;25
529
^1,053
3,172
2,793
82U
7,8ij.2
856
$321
1,096
583
220
2,220
130
Grarid total
(13 control structu;:'e3 )
17, 95U
8,698 2,350
$1,061
2,6l!l
2,260
835
6,797
55i4
7,351
1/ Does net include ccst of engineering supe:.'vision or administrative overhead
2/ Includes $12,800 for construction of perrn>iiient-type Ocqueoc River iveir and
~ traps which -w-as in;? tailed ir-. 19l!.8
30
average, to-'cal length of the runs in Carp Creek has decreased 9 percent
(1,6 inches) from a maximum of 17.1!. inches in 19U7 to 1^.6 inches in
1951. In samples from both Carp Greek and the Ocqueoc River, the aver-
age total length declined betvreen O08 sxid 0.9 inch in the period I9I1.9 to
19.5l-» The average \veight of migrar.ts entering Cai'p Creek has decreased
about 33 percent (approximately 70 g.cmi3) during the ^--year period.
Arjy further decline in the size of mature spawning migrants -will
profoimdly affect any proposed control program based on the operation of
T/eirs and traps. Further reduction of 'tveir screen or grate aperatures
below the l/2-inch spacing now required will create e:5rtremely difficult
operational problems during spring flox.s.
The spaivning runs in Carp Creek and the Ocqueoc River in 19^1 did
not differ in character or in their response to cer-t.ain factors in the
environraent vary from these vane occu.';"::';j.ig in the same str-eans in pru-
vioiis years. Data pertaining to the runt, in these tvro streams in 19^1
are presented in tables 9} 10, and 12; s^jnilar information for the runs
occurring in 1950 has been presented by Applegate and Smith (19^1) and
for the years 19l;7, 19U8, and I9U9 by Applegate (19^0) . Strict compai-i-
sons of the character of the Ocqueoc River run in 19bl in relation to
time of migration and response to variov.s environmental factors should
not be made T/ith those runs of former years. Daily and periodic catches
in this i-iver (as detailed in table IC) were strongly influenced b3- the
operation of an e^rper-imental electromechanical weir and trap located below
the permanent Ocqueoc River installatioxi. Experimentation with this new
de\ace was carried on intermittentlly fi'om May 1 to June l5« During the
periods of effective operation of the eJ.ectromechanical iveir, many or all
lampreys "vvBre blocked below the electrodes and did not enter the traps in
the pe;:Tiianent installation until the ele^jtrical device became 5_noperative .
New developments and further evaluation of mechanical
control devi'c'-fis 5/
Barrier dams. — The experimental bar:-ier dam in the Black River,
Mackinaw Coimty, Michigan, which was designed to block aiid divert spawn-
ing runs of sea lampreys was rebuilt by the Michigan Department of Conser-
vation during the winter of 19^0-51 (fig';"'. 3 and i;)o A trap, which was
installed in the wall of the original c'am, was removed and the cur-vod
steel lip attached to the face of the dan was extended further across the
strear.1. These changes enabled the str-.ir/jure to handle ivith greater facil-
ity the large discharge of the Black Riv'?-;r during the spring runoff.
5/Fi.ve types of mechanical control de'Ticos have been developed to date:
Xl) large, permanent type weirs and tri.ps for capturing spawning i-uns,
(2) and (3) portable-type iveirs for use in medium- and small-sized streams
for capturing spawning runs, (li) dams ai:d inclined-screen trap units for
captiiring 7/^oung, downstream migrants, aiid, (5) barrier dams for blocking
and diverting spa-vming runs. The essential characteristics and the liiTiita-
tions of these devices have been described in an earlier report (Applegate
and Sirdth, 19^1).
31
Figure 3. — Experimental sea- lamprey barrier dam in the Black River,
Mackinaw County, Mich.
Figure ^. — Glose-up of harrier dam showing overhanging, curved
lip of sheet steel attached to wall of dam.
32
The dam functioned satisfactorily thi-oiighout the spring of 19pl..> blocked
all laiiipreys entering the stream from reaching the spawning groxmds in
that river J and, offered no significant barrier to the \ipstream migra-
tions of game fishes.
Control structures of this type vrl.ll be especially usefxtL in many
streams on the southwestern shore of Lal:e Superior indaich ax's cha:;'acter-
ized by steep gradients and very stable substrata and -where poor access-
ibility precludes the installation of devices which must be serviced
daily.
Portable-type iveirs and traps. — S'^reen, trap, and bracirig units of
the several portable-type weirs were operated in the streams of Control
Zone H-1 and in Lal-ce Superior tributa;:;'ies in 19^1 ivith no major structural
changes. Wearing quality of the original units, as designed, has been
found to be excellent. Most portable-t;rpe screen and trap units appai'-
ently will give from I4. to 5 years of service under reasonable stream con-
ditions before any replacement becomei:- necessary.
One innovation tested in Carp Creek, Presque Isle Co-ant}'', Mich,, was
the use of permanent sills, trap base, aizd abutments which were con-
structed of reinforced concrete (figs. 5 and 6). This stable substruc-
ture proved extremely effective. It pL'actically eliminatec' danger of
undercutting or bank-cutting and provided continuous troubld-free opera-
tion through a spring season of unusually high floods.
Similar concrete sills and abutments were installed in Hibbarda
Creek, Door County, Wis., by the Wiscor.sin Conservation Department. This
installation likewise proved to be much more effective than the wooden
substructures used in previous yearr.
The specific advantages of these v:;oncrete substinictui-es appear to
be as follows: (1) elimination of occasional replacement of substruc-
tm-'e; (2) reduction of maintenance of substructure to a minim-om,; (3)
reduction of wear on portable screen .and trap units j and, (u) reduction
of operating costs through increased e.ose of -weir operation (fe-#ei' man-
hours required for inspection and servicing). It would, seem, advisable,
therefore, in a long-term control program to install this mo:."e stable
weir and trap base in all streams where the portable-type str^ictu;.-es sz'e
to be used. Although initial capital outlay vrould obviously be g:?eater
tlian for similar wood substructiireE, tie advantages indiuatsd above
should effect more than compensating cr/ings over a period of years.
Operating costs in 1950 and 19^1 fo"-£ Control Zone H-1 and one stream
tributary to Lal-:e Superior. — Detailed records have been kept tlii'ough two
seasons of~operations of the costs ino--:.rred in installing^, operating, and
maintaining the 12 weirs and traps of Control Zone H-1 and the Pendills
Creek iveir. Briefly, these 13 conti-ol .structures were installed at an
aggregate cost of $17,95U and operated, successfully during the 19^0 sea-
son for $8,698. They vrere reinstalled in 19^1 at a cost of ^2,350 and
operated throughout that season for $7,35l. The cost of reinstallation
in 1951 is not typical of a normal season since it includes fi,mds expended
33
in the experimental installation of reinforced concrete sills and abut-
ments in Carp Creek (Unit 2). Had this additional construction been
omitted, reinstallation costs would have been approximately $1,000.
The figures presented above are broken down in Table 13 where
they are presented by operational units. An operational un.t is any
weir and trap or group of such structures which, when geography and
work load are considered can be most economically and efficiently oper-
ated by a single crew of men. Unit crews consist of night and day
shifts of one to four men per shift depending on the season aid the size
of the unit.
The expenditures indicated here for individual operational units
are believed to be representative of the costs of installing and operat-
ing such units (comprised of one or more mechnical control devices)
in any other similar areas in the Lake Huron and Lake Michigan basins.
Gross costs in other unit geographic areas such as Control Zone H-1
will vaiy widely from the costs indicated for that Zone depending on
the number of large, permanent- type weirs and traps required, the extent
of the area (control zone), and the dispersion of all required control
structures within the area. Operating unit costs in Control Zone H-1
will not apply, for example, in the Lake Superior basin where the
accessibility of most streams requiring control devices is very poor.
No data are available concerning installation and operating costs in
streams in the more remote and wild areas bordering on that lake.
Figure 5. --Portable -type weir and trap in Carp Creek,
Presque Isle County, Mich. , after installa-
tion of permanent sills, trap base, and
abutments of concrete.
34
5
3003 nrsviS
z
_l ~'0
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Literature cited
Applegatoj Vomi'-;:.
1950. Natural history of the sea lamprey (Peti-oriyzon marinus)
in Michigan. U. S. Dept. of Iiit,, Fish aid midl, Serv,,
Spec. Sci. Rept»: Fisheries IIo, 55: 237 PP.
Applegate, Vernon C. and Bernard R, Smith
1951. Sea lamprey spavming runs in the Great Lakes in 1950,
U. S. Dept. of Int., Fish and Wildl. Serr., Spec, Sci,
Rept.: Fisheries No, 6I: U9 pp.
Shetter, Da-'/id So
I9U9. A brief history of the sea lamprey problem in MLchigaii
waters. Trans « Amer. Fish. Soc, Vol. 76 (?9U6),
pp. I6O--I76.
List of ccmmon and scientific
Black bull head
Brook stickleback
Brook trout
Brown trout
Bxirbot
Carp
Comrnon shiners
Creek chubs
Great Lakes longnose dace
Lalce chub
Lake herring
Lake trout
Logperch
Longnose sucker
Muddler
Mudniinnow
Northern pike
Pumpkinseed
Rainbow trout
Rock bass
Sea lamprey
Silver lamprey
Smallraouth bass
Smelt
Yellow perch
Walleye
Wl'iite sucker
names of fishes raent-ioned in this report
iUneiurus m .. me las
Eucalia i noons cans
Salvelinus i - lontinalis
Salmo trutta
Lota 1. maculosa
Cyprinus carpio
Notropis cornutus frontalis
Seiiiotilus a. abromaculatus
Rh;michthys c, cat aractae
Couosius plunbeus
Leucichthys artedii
Salvelinus (Cristivome:-) n« naiiiaycush
Percina oaprodes
Catostomu? c. catostomus
Cottus b, bairdi
Umbra limi
Esox lucias
Lepomis gibbcsus
Saimo gairdneri
Ambloplites rupestris
Pe^tromy^on mai'inus
I.''.hthyom)'3on unicuspis
Micropteriis d_^. dolonieui
Osmerus morday
Perca f laves "ens
Stizostedion Vj. vitreum
Catostonius Co commersoni
37
I562li
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^ WHSE 01049
*