A stereo-atlas of ostracod shells

A Stereo-Atlas of Ostracod Shells

edited by R. H. Bate, D. J. Horne, J. W. Neale,

and David J. Siveter

Volume 14, Part 1; 30th May, 1987

Published by the British Micropalaeontological Society, London

Editors

Dr R.H. Bate, SSI (UK) Ltd., Tannery House, Tannery Lane, Send, Woking, Surrey GU23 7EF.
Dr D.J. Horne, Department of Geology, City of London Polytechnic, Walburgh House, Bigland Street,
London El 2NG.

Prof. J.W. Neale, Department of Geology, The University, Hull HU6 7RH.

Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.

Editorial Board

Dr G. Bonaduce, Stazione Zoologica, 80121 Napoli, Italy.

Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles, France.
Dr P. De Deckker, Research School of Pacific Studies, Australian National University, PO Box 4,
Canberra ACT 2600. Australia.

Dr D. van Harten, Universiteit van Amsterdam, Geologisch Instituut, Nieuwe Prinsengracht 130,
Amsterdam. The Netherlands.

Dr I. Purper, Departamento de Paleontologia e Estratigrafia, UFRGS, 90 000 Porto Alegre RS, Brazil.
Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse
55, D 2000 Hamburg 13, West Germany.

Officers of the British Micropalaeontological Society

Chairman Dr A.C. Higgins, BP Research Centre, Chertsey Road, Sunbury-on-Thames, Middlesex
TW16 7LN.

Secretary Dr P.P.E. Weaver, Institute of Oceanographic Sciences, Brook Road, Wormley, Godaiming,
Surrey GU8 5UB. Tel: 0428-79 4141.

Treasurer Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD. Tel: 01-589 6323.

Journal Editor Dr. L.M. Sheppard, SSI (U.K.) Limited, Chancellor Court, 20 Priestly Road, Guildford,
Surrey GU2 5YL. Tel: (0483) 506605.

Newsletter Editor Dr R.L. Austin, Department of Geology, University of Southampton, Southampton
S09 5NH. Tel: (0703) 559122/557941

Conodont Group Chairman Dr R.J. Aldridge, Department of Geology, University of Nottingham,
University Park, Nottingham NG7 2RD.

Secretary Dr P.M. Smith, Department of Earth Sciences, University of Cambridge, Downing Street,
Cambridge CB2 3EQ. Tel: (0223) 355463 (or 276121).

Foraminifera Group Chairman Dr P. Copestake, Britoil, 150 St. Vincent Street, Glasgow G2 5LJ.

Secretary Dr D.J. Shipp, Robertson Research Int. Limited, Ty’n-y-Coed, Llanrhos, Llandudno LL30
ISA. Tel: (0492) 81811.

Microplankton Group Chairman Dr G.L. Eaton, BP Research Centre, Chertsey Road, Sunbury-on-
Thames, Middlesex TW16 7LN.

Secretary Dr A.J. Powell, BP Research Centre, Chertsey Road, Sunbury-on-Thames, Middlesex TW16
7LN. Tel: (09327) 62818.

Ostracod Group Chairman Dr D.J. Horne, Geology Department, City of London Polytechnic,
Walburgh House, Bigland Street, London El 2NG.

Secretary Dr C. Maybury, Department of Geology, University College of Wales, Aberystwyth, Dyfed
SY23 3DB. Tel: (0970) 3111.

Palynology Group Chairman Dr M.C. Boulter, N.E. London Polytechnic, Romford Road, London E15
4LZ.

Secretary Dr J.E. A. Marshall, Department of Geology, The University, Southampton S09 5NH. Tel:
(0703) 559122.

Calcareous Nannofossil Group Chairman Mr M. Jakubowski, Robertson Research Int. Limited,
Ty’n-y-Coed, Llanrhos, Llandudno, Gwynedd LL30 ISA.

Secretary Dr J. Crux, BP Research Centre, Chertsey Road, Sunbury on Thames, Middlesex TW16 7LN.
Tel: (09327) 63062.

Instructions to Authors

Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited.
Format should follow the style set by the majority of papers in this issue. Descriptive matter apart from
illustrations should be cut to a minimum; preferably each plate should be accompanied by one page of
text only. Blanks to aid in mounting figures for plates may be obtained from any one of the Editors or
Editorial Board. Completed papers should be sent to Dr David J. Siveter.

The front cover shows a left valve of Neolimnocy there hexaceros Delachaux, 1928, from Quaternary
Deposits at Lago Junin, Peru. Photograph by Dr P. De Deckker, University of Monash, Victoria,
Australia.

Printed in the UK by BPCC Northern Printers Ltd., Stanley Road, Blackpool FY1 4QN

Stereo-Atlas of Ostracod Shells 14 (1) 1-4(1987) Cathaycythere reticulata ( 1 of 4)

595.337.14 (119.4) (265.72 : 161.108.21 + 161.109.21) : 551.351 + 552.51 + 52

ON CATHAYCYTHERE RETICULATA WHATLEY & ZHAO gen. et sp. nov

by Robin Whatley & Zhao Quanhong

(University College of Wales, Aberystwyth, UK & Tongji University, Shanghai, China)

Derivation of name:

Genus CATHAYCYTHERE gen. nov.

Type-species: Cathaycythere reticulata gen. et sp. nov.

From Cathay (China) + cythere; with reference to the type locality of the type-species in the South
China Sea.

Diagnosis:

Medium sized; subrectangular in lateral view, highest anterodorsally and longest subventrally;
anterior margin broadly rounded; posterior margin truncate above and bluntly pointed below.
Carapace laterally compressed at both end margins. Eye-tubercle and internal ocular sinus absent.
Surface coarsely and irregularly reticulate with a prominent subcentral node surrounded by a
subcircular sulcus. Hinge holamphidont with high conical anterior tooth, bulbous posterior tooth
and locellate median element in right valve. Inner lamella wide with a marked oval excavation
posteriorly aligned parallel to the postero-dorsal slope of the posterior margin; very narrow
vestibulae at each end; radial pore canals few, long, thin and simple. Adductor scars relatively
small, consisting of a vertical row of four; frontal scar single, oval.

Explanation of Plate 14, 2

Fig. 1, RV, ext. lat. (holotype, 1986.404, 710/xm long); fig. 2, LV, ext. lat. (paratype, 1986.405, 665/xm long); fig. 3, LV, ext. lat.

(paratype, 1986.406, 665/xm long).

Scale A (100/xm; x90), figs. 1-3.

Stereo-Atlas of Ostracod Shells 14, 3 Cathaycythere reticulata (3 of 4)

Remarks:

The excavation on the wide posterior inner lamella is not known to occur in any other genus and
can be used to distinguish it from Sinocythere Hou, 1978. The latter is closest in ornament and
muscle scar pattern to the present genus, but differs in hingement (hemiamphidont with clearly
crenulate posterior tooth) and in lacking the posterior excavation on the inner lamella. Kritlie
exhibits a similar excavation posteriorly but it occurs distal to the selvage whereas in Cathaycythere
it is proximal. The familial status of Cathaycythere is uncertain and possibly a new family is
required to accommodate this genus and Sinocxthere (see Whatlev & Zhao, Stereo-Atlas Ostracod
Shells 14, 5-8, 1987).

Cathaycythere reticulata Whatley & Zhao gen. et sp. nov.

Holotype:

British Museum (Nat. Hist.) no. 1986.404; RV.

Type locality:

[Paratypes: British Museum (Nat. Hist.) nos. 1986.405-409.)

Off Guangsi Province of China, Gulf of Tonkin, South China Sea; lat. 21° 29' 09"N, long. 108° 44'
46"E. Recent, water depth; 14m.

Derivation of name:
Figured specimens:

With reference to its reticulate surface.

British Museum (Nat. Hist.) nos. 1986.404 (holotype, RV; PI. 14, 2, fig. 1), 1986.405 (paratype,
LV: PI. 14, 2, fig. 2), 1986.406 (paratvpe, LV: PI. 14, 2, fig. 3), 1986.407 (paratvpe, car.: PI. 14. 4,
fig. 1), 1986.408 (paratype. LV: PI. 14, 4, fig. 2), 1986.409 (paratype, RV: PI. 14, 4, fig. 3). Nos.
1986.404-406 are from the type locality. Nos. 1986.407-409 are from lat. 21° 15' 46"N, long. 109°
24' 57"E; Recent, water depth: 14m.

Diagnosis:

Distribution:

As for the genus. Monotypic.

In four bottom samples off Guangsi Province, China, northern Gulf of Tonkin, South China Sea.
Water depth: 10 - 16m; substrate: mud to fine sand.

Explanation of Plate 14, 4

Fig. 1, car., ext. dors, (paratype, 1986.407, 635/xm long); fig. 2, LV, int. lat. (paratype, 1986.408, 635/xm long); fig. 3, RV, int. lat
(paratype, 1986.409, 645/xm long).

Scale A (100/xm; x90), figs. 1-3.

Cathaycythere reticulata (2 of 4)

Stereo-Atlas of Ostracod Shells 14, 2

Stereo-Atlas of Ostracod Shells 14, 4

Cathaycythere reticulata (4 of 4)

Stereo-Atlas of Ostracod Shells 14 (2) 5-8 (1987) Sinocy there sinensis (1 of 4)

595.337.14 (119.9) (510 : 161.120.34) : 551.351

ON SINOCYTHERE SINENSIS HOU

by Robin Whatley & Zhao Quanhong

(University College of Wales, Aberystwyth, UK & Tongji University, Shanghai, China)

Genus SINOCYTHERE Hou, 1982
Type-species (by original designation): Sinocythere sinensis Hou, 1982
Diagnosis: Medium sized; subrectangular in lateral view with parallel dorsal and ventral margins, prominent

posterior cardinal angle and obtusely rounded posterior margin. Eye-tubercle weakly developed
but internal ocular sinus not developed. Surface reticulate with an anteromedian node surrounded
by a subcircular sulcus. Hinge hemiamphidont: anterior tooth in right valve conical, posterior
tooth curved and distinctly dentate; in left valve, anterior socket enclosed ventrally by an anterior
extension of the anteromedian conical tooth, posterior socket with an anti-slip toothlet
ventromedianly, and median bar denticulate. Inner lamella relatively wide with shallow anterior
vestibule; radial pore canals few, thin and simple. Adductor muscle scars small, consisting of a
vertical row of four scars all in contact; frontal scar single, oval.

Remarks: This genus is close to Spinoleberis Hanai, 1961 in many features except that the latter has a
triangular outline and much narrower, more acute posterior margin in lateral view, and three
longitudinal ribs. In external characters Sinocythere is somewhat similar to Palmenella Hirschman,
1916, but the latter bears a schizodont hinge. Cathaycythere Whatley & Zhao, 1987 ( Stereo-Atlas
Ostracod Shells 14, 1-4) has similar ornament and also the circular sulcus surrounding the
anteromedian node. The two genera differ in hingement and Sinocythere lacks the excavated
posterior inner lamella so typical of Cathaycythere. Sinocythere and Cathaycythere are probably
worth including in a new family of Cytheracea.

Explanation of Plate 14, 6

Fig. 1, cf RV, ext. lat. (1986.410, 570/u.m long); fig. 2, § car., rt. lat. (1986.411. 540/u.m long); fig. 3, 9 car.. It. lat. (1986.411, 540/u.m
long). Scale A (100/u.m; xllO), figs. 1-3.

Stereo-Atlas of Ostracod Shells 14, 7

Sinocythere sinensis (3 of 4)

Sinocythere sinensis Hou, 1982

1982 Sinocythere sinensis sp. nov. Hou in Hou et al., Cretaceous-Quaternary ostracode fauna from Jiangsu, 242, text-fig. 77; pi. 87,
figs. 16-19.

1985 Sinocythere sinensis Hou; Zhao, Acta Oceanologica Sinica, pi. 2, fig. 10.

1985 Sinocythere sinensis Hou; Wang and Zhao, in Wang et al., Marine Micropaleontology of China, pi. 18, fig. 4.

Holotype:
Type locality:
Figured specimens:

Diagnosis:

Remarks:

Distribution:

Nanjing Institute Geology & Paleontology, Academia Sinica; no. 4107. 9 LV. Not figured herein.
Jiangsu Province, Eastern China; Dongtai Formation, Quaternary.

British Museum (Nat. Hist.) nos. 1986.410 (cf RV: PI. 14. 6, fig. 1; PI. 14. 8, fig. 3). 1986.411 (9
car.: PI. 14, 6, figs. 2, 3; PI. 14, 8, fig. 1), 1986.412 (9 LV: PI. 14, 8, fig. 2). All Recent, collected
from the littoral of Jiangsu Province, China; approx, lat. 34° 17'N, long. 120° 17'E.
Irregularly polygonal surface reticulation with superimposed narrow, oblique posterodorsal-
anteroventral rib, and a bifid rib running from the anterior cardinal angle obliquely to
mid-anterior. Female carapace strongly laterally compressed posterodorsally. posteriorly and
ventrally; the male is inflated in these areas.

This species is very close to S. dongtaiensis Chen, 1982 in outline and overall ornamentation, but
the latter is much more weakly reticulate.

Pliocene to Recent, Eastern China. Recent specimens most abundant in littoral zone and inner
shelf shallower than 20m, rare in estuaries and water depths from 20 to 200m.

Explanation of Plate 14, 8

Fig. 1, 9 car., ext- dors. (1986.411, 540jum long); fig. 2, 9 LV, int. lat. (1986.412, 490/xm long); fig. 3, cf RV, int. lat. (1986.410,
570/u.m long).

Scale A (100/u.m; xllO), figs. 1-3.

Stereo- Atlas of Ostracod Shells 14, 6

Sinocythere sinensis (2 of 4)

Stereo-Atlas of Ostracod Shells 14, 8

Sinocythere sinensis (4 of 4)

Stereo-Atlas of Ostracod Shells 14 (3) 9-12 (1987) Albileberis sinensis (1 of 4)

595.337.14 (119) (510 : 161.120.34) : 551.313.1 + 551.35

ON ALBILEBERIS SINENSIS HOU

by Zhao Quanhong & Robin Whatley

(Tongji University, Shanghai , China & University College of Wales, Aberystwyth, UK)

Genus ALBILEBERIS Hou, 1982
Type-species: Albileberis sinensis Hou, 1982

Diagnosis: Small to medium, laterally compressed; subovate to rectangular in lateral view with greatest height

anteromedianly. Bluntly truncate posterior and well rounded anterior margins. Surface smooth,
pitted or reticulate. Hinge between paleomerodont and holomerodont; all positive elements in
right valve, long ridge-like anterior tooth thickened anteriorly with a prominent square terminal
toothlet, median bar smooth anteriorly and faintly crenulate posteriorly, posterior tooth curved
and denticulate. Complementary negative elements in left valve with an anti-slip bar below median
groove and anterior socket, and two small terminal anti-slip toothlets respectively at anterior and
posterior ends. Inner lamella moderately wide with large vestibulae and narrow fused zone; radial
pore canals moderate in number and simple. Adductor muscle scars a vertical row of four; frontal
scar V-shaped; fulcral point crescentic and prominent.

Remarks: Although the generic name Albileberis was first published in Guan et al. ( Paleontological Atlas of

Central & S China (4): Ostracoda, 1978), who attributed the genus to Hou from an earlier MS
name, its type-species was not published until 1982 (Hou in Hou et al.). This genus is easily
identified by its laterally compressed carapace, outline and peculiar hingement, and probably
belongs to the Cytherideinae based on its internal characters.

Explanation of Plate 14, 10

Fig. 1, $ RV, ext. lat. (1986.413, 525/u.m long); fig. 2, 9 LV, ext. lat. (1986.414. 525pm long); fig. 3, cf LV. ext. lat. ( 1986.415. 495/im
long).

Scale A (lOO/um; xl20), figs. 1-3.

Stereo-Atlas of Ostracod Shells 14, 1 1 Albileberis sinensis (3 of 4)

Albileberis sinensis Hou, 1982

1982 Albileberis sinensis sp. nov. Hou, in Hou et al. , Cretaceous-Quaternary ostracode fauna from Jiangsu., 240-241, text-fig. 75. pi.
88, figs. 1-7.

1985 Albileberis sinensis Hou; Zhao, Acta Oceanological Sinica, pi. 1, fig. 9.

1985 Albileberis sinensis Hou; Wang & Zhao, in Wang et al., Marine Micropaleontology of China, pi. 7, fig. 2, text-fig. 5.

Holotype:

Type locality:
Figured specimens:

Diagnosis:

Remarks:

Distribution:

Nanjing Institute Geology & Paleontology, Academia Sinica; no. 41062; 9 LV. Not figured
herein.

Jiangsu Province, E China; Dongtai Formation, Quaternary.

British Museum (Nat. Hist.) nos. 1986.413 (9 RV: PI. 14, 10, fig. 1), 1986.414 (9 LV: PI. 14, 10,
fig. 2), 1986.415 (cf LV: PI. 14, 10, fig. 3), 1986.416 (9 car.: PI. 14, 12, fig. 1), 1986.417 (9 LV: PI.
14, 12, fig. 2), 1986.418 (9 RV: PI. 14, 12, fig. 3). All Recent, collected from the littoral of Jiangsu
Province, China; approx, lat. 34° 17'N, long. 120° 17'E.

Subovate in lateral view with vertical truncated posterior margin, surface smooth with few very
weak reticulae around the margins.

Markedly differs from the other two species in this genus. A. sheyangensis Chen, 1982 is much
more elongate and lower with obliquely truncated posterior margin, and A. asperata Guan, 1978
has an ornamentation of coarse reticulation.

China, Quaternary to Recent. At the present day this species is abundant and widespread in
brackish and nearshore waters along the coast of the East China and Yellow Seas with a salinity
range of 3%o to normal sea water, including marshes, estuaries, littoral and the inner shelf
shallower than 50m.

Explanation of Plate 14, 12

Fig. 1, 9 car., ext. dors. (1986.416, 535,u,m long); fig. 2, 9 LV, int. lat. (1986.417, 515pm long); fig. 3, 9 RV. int . lat. (1986.418.
515jU,m long).

Scale A (lOOpun; xl20), figs. 1-3.

Stereo-Atlas of Ostracod Shells 14, 10

Albileberis sinensis (2 of 4)

Stereo-Atlas of Ostracod Shells 14, 12

Albileberis sinensis (4 of 4)

Stereo-Atlas of Ostracod Shells 14 (4) 13-16 (1987) Sinocytheridea impressa (1 of 4)

595.337.14 (118.22 + 119.9) (512.317 + 510 : 161.118.39) : 551.313.1 + 551.35

ON SINOCYTHERIDEA IMPRESSA (BRADY)

by Zhao Quanhong & Robin Whatley

(Tongji University, Shanghai, China & University College of Wales, Aberystwyth, UK)

Genus SINOCYTHERIDEA Hou, 1978
Type-species (by subsequent designation) : S. latiovata Hou & Chen, 1982
(= Cytheridea impressa Brady, 1869; see below)

Diagnosis: A genus of Cytherideidae characterized by its modified antimerodont hingement with a

conspicuous anti-slip toothlet anteriorly in the left valve, by which it can be readily distinguished
from such similar genera as Cyprideis Jones, 1857, Neocyprideis Apostolescu, 1965 and
Sarsicytheridea Athersuch, 1982.

Remarks: Sinocytheridea was named by Hou in manuscript more than 20 years ago but remained

unpublished until Guan et al. (1978) first applied this name for the genus in a published work,
attributing the genus to Hou and using Hou's original description. 5. latiovata Hou & Chen, 1982
was designated by Hou and Chen in Hou et al. (1982) as the type-species of Sinocytheridea. The
present authors, however, have recently studied Brady’s material from Hong Kong which is
deposited in the Hancock Museum and consider 5. latiovata and Cytheridea impressa Brady, 1869
to be conspecific. We therefore consider C. impressa to be the type-species of Sinocytheridea.

Explanation of Plate 14, 14

Figs. 1, $ car., rt. ext. lat (paralectotype, 1.23.44, 740 pm long); fig. 2, 9 LV. ext. lat. (1986.419, 720 pm long); fig. 3, 9 car., ext-
dors. (1986.421, 750 pm long).

Scale A (100 p.m; x 85), figs. 1-3.

Stereo- Atlas of Ostracod Shells 14, 15 Sinocytheridea impressa (3 of 4)

Sinocytheridea impressa (Brady, 1869)

1869 Cytheridea impressa sp. nov. G.S. Brady in L. De Folin and L. Perier (eds.). Les Fonds de la Mer., 158, pi. 16, figs. 13, 14.
1978 Cyprideis yehi Hu & Yeh, Geol. Soc., China (Taiwan), Proc., 21, 157-159, text-fig. 5, pi. 3. figs. 10-13.

1978 Sinocytheridea sinensis Hou; Hou in Guan et al., Paleontological Atlas Central & S China (4): Ostracoda, 240, pi. 65, figs. 1-5.

1982 Sinocytheridea latiovata sp. nov. Hou & Chen in Hou et al., Cretaceous-Quaternary ostracode fauna from Jiangsu, 164-165,
text-figs. 26a-c, pi. 72, figs. 10-20.

Sinocytheridea longa Hou & Chen, ibid. 165-166, text-figs. 27a-c, pi. 72, figs. T9.

1982

Lectotype:

Type locality:
Figured specimens:

14, 16, fig.
14, 14, fig.

Hancock Museum, Newcastle-upon-Tyne, England, no. 1.24.37, 9 LV.

[Paralectotypes: Hancock Mus., nos. 1.24.38, 9 RV; 1.23.44, 9 car.]

Hong Kong Harbour; Recent.

Hancock Museum, Newcastle-upon-Tyne, England, nos. 1.24.37 (lectotype, 9 LV: PI.

2) , 1.24.38 (paralectotvpe. 9 RV: PI. 14, 16, fig. 3), 1.23.44 (paralectotvpe, 9 car.; PI.

1), Brit. Mus. (Nat. Hist.) nos. 1986.419 (9 LV: PI. 14, 14, fig. 2). 1986.421 ($ car.: PI. 14. 14. fig.

3) , 1986.420 (cf LV: PI. 14, 16, fig. 1). Nos. 1.23.44, 1.24.37 and 1.24.38 belong to the Brady
Collection in the Hancock Museum and are from the type-locality; nos. 1986.419-431 are from the
Pohai Bya, China, lat. 39° 50'N, long. 118° 46'E, water depth: 15m.

Elongate-oval without obvious trace of angle in lateral view'. RV larger than and slightly
overlapping LV along the periphery except anterior margin. Surface smooth with rounded shallow'
pits (openings of sieve-type normal pore canals). Avestibulate, radial pore canals few, simple.
Adductor muscle scar a vertical row of four elongate scars; frontal scar V-shaped.

Apart from the more elongate outline, 5. longa Hou & Chen is identical in carapace features to S.
impressa. The authors believe that the former species is based on males of S. impressa.
Pliocene to Recent, China. The modern representatives occur widely in shelf, littoral, estuaries,
marshes, tidal pools and channels of the supralittoral zone along the entire coast of China w'ith a
salinity distribution ranging from about 2%o to normal sea water and a w'ater depth ranging from
middle shelf (50-f00m) to supralittoral.

Explanation of Plate 14, 16

Fig. 1, cf LV, ext. lat. (1986.420, 660 pm long); fig. 2, 9 LV, int. lat. (lectotype, 1.24.37, 775 pm long); fig. 3, 9 RV. int. lat.
(paralectotype, 1.24.38, 740 yum long). Scale A (100 /im; x 85), figs. 1-3.

Diagnosis:

Remarks:

Distribution:

Stereo- Atlas of Ostracod Shells 14, 16

Sinocytheridea impressa (4 of 4)

Stereo- Atlas of Ostracod Shells 14, 14

Sinocytheridea impressa (2 of 4)

Stereo-Atlas of Ostracod Shells 14 (5) 17-20 (1987)

595.337. 14 (118.22) (798 : 162.153.70) : 551.351 + 552.52

Pterygocythereis vannieuwenhuisei (1 of 4)

ON PTERYGOCYTHEREIS VANNIEUWENHUISEI BROUWERS sp. nov.

by Elisabeth M. Brouwers
(U.S. Geological Survey, Denver)

1986 Pterygocythereis
Holotype:

Type locality:

Derivation of name:
Figured specimens:

Diagnosis:

sp
U.S

nov.

Pterygocythereis vannieuwenhuisei sp. nov.

C. A. Repenning, E. M. Brouwers, et al., Bull. U.S. Geol. Surv., 1687, pi. 1, fig. 1.
National Museum no. 410130, cf RV.

[Paratypes: U.S. National Museum nos. 410131-410134].

Cutbank on a tributary of the Kalikpik River, Arctic coastal plain. North Slope, Alaska (lat. 70°
26.7'N, long. 152° 09.4'W); Pliocene. Outcrop consists of 1.8m of late Pliocene marine clay and
sand overlain by 5.5m of Pleistocene fluvial and eolian sands. Deeper inner sublittoral to middle
sublittoral water depths; cold temperate to subfrigid marine climate.

In honour of Don Van Nieuwenhuise, research geologist at Amoco Production, Houston.
U.S. National Museum nos. 410130 (holotype, cf RV: Text-fig. 1), 410131 (paratvpe, 9 LV: PL
14, 18, fig. 1), 410132 (paratype, cf RV: PI. 14, 18, fig. 2), 410133 (paratype, cf LV: PI. 14, 20. fig.
1), 410134 (paratype, LV: PI. 14, 20, fig. 2). All from the type locality and horizon (locality
83-EB-187, 188, collected by E. Brouwers, 1983).

Short, high, rectangular lateral outline; large size; weak dimorphism. Three pairs of spines in
median valve area; large, strong marginal spines. Wide marginal flange, continuous along anterior
and venter. Numerous ventral marginal spines. Spinose anterodorsal margin with weak underlying
flange. Strong posteroventral spinose prolongation. Left valve hinge has anterior socket with
ventral rim; elongate, U-shaped posterior socket; cylindrical posteromedian tooth; weakly
crenulate median bar.

Explanation of Plate 14, 18

Fig. 1, $ LV, ext. lat. (paratype, 410131, 1380/um long); fig. 2, cf RV, ext. lat. (paratype, 410132. 1460/am long).
Scale A (100/um; xl25), figs. 1, 2.

Stereo-Atlas of Ostracod Shells 14, 19

Remarks:

Pterygocythereis vannieuwenhuisei (3 of 4)

Distribution:

Pterygocythereis ranges from the Paleocene-Holocene, occurring commonly throughout the
subtropical and temperate N Atlantic Ocean and rarely in the subfrigid Norwegian Sea. In the
northwestern Atlantic Ocean, Pterygocythereis occurs in the southern cold temperate zone, but
does not live in the northern cold temperate or subfrigid zones of the western N Atlantic.
Pterygocythereis vannieuwenhuisei is related to the European P. mucronata-P. jonesii species
complex and not to the more temperate NW Atlantic P. americana-P. inexpectata lineage.
?Late Miocene, early-late Pliocene (to 2.48 Ma): NE Alaska, three localities in Colvillian-aged
sediments of the Gubik Fm. (Fish Creek, Kalikpik R., Miluveach Creek; 2.48 - 3.0 Ma, late
Pliocene; Repenning et ah, op. cit.), three localities in the upper Nuwok Member of the
Sagavanirktok Fm. (Carter Creek, Barter Is., Manning Pt.; ?late Miocene, lower to middle
Pliocene).

P. vannieuwenhuia

°

icPm

°

a,

~ d

°

Text-fig. 1. Holotype (cf RV, USNM no. 410130),
camera lucida drawing, seen in transmitted light.

Text-fig. 2. Plot of length vs. height for 26 specimens
from the type locality.

Explanation of Plate 14, 20

Fig. 1, cf LV, ext. lat. (paratype, 410133, 1280/um long); fig. 2, LV, int. lat. (paratype, 410134, 1300/um long).
Scale A (100/am; xl25), figs. 1, 2.

Stereo- Atlas of Ostracod Shells 14, 18

Pterygocythereis vannieuwenhuisei (2 of 4)

Stereo-Atlas of Ostracod Shells 14 (6) 21-24 (1987) Muellerina hazeli (1 of 4)

595.337.14 (119.1/119.9) (261.4 : 162.082.24) : 551.351/352

ON MUELLERINA HAZELI COLES & CRONIN sp. nov.

by Graham P. Coles & Thomas M. Cronin
(University College of Wales, Aberystwyth & U.S. Geological Survey, Reston, Virginia)

Muellerina hazeli sp. nov.

Holotype:

Type locality:
Derivation of name:
Figured specimens:

British Museum (Nat. Hist.) no. OS 12971, 9 LV.

[Paratypes: British Museum (Nat. Hist.) nos. OS 12972-OS 12975. Four additional
paratypes have been deposited in the U.S. Museum of Natural History: USNM nos.

409239-409242).

Off the Florida Keys, United States continental slope; approx, lat. 24° 26'N, long. 81° 38'W;
Recent, water depth 107m.

In honour of Joseph E. Hazel, in recognition of his studies on Muellerina from the Atlantic Coastal
Plain and shelf.

British Museum (Nat. Hist.) nos. OS 12971 (holotype, 9 LV: PI. 14, 22, fig. 1; PI. 14, 24, fig. 1).
OS 12972 (paratype, 9 RV: PI. 14, 22, fig. 2), OS'l2973 (paratype, cf LV: PI. 14. 22, fig. 3). OS
12974 (paratype, cf RV: PI. 14, 24, fig. 2), OS 12975 (paratype, cf car.: PI. 14, 24, fig. 3). All from
the type locality and horizon.

Explanation of Plate 14, 22

Fig. 1, 9 LV, ext. lat. (holotype, OS 12971, 580/um long); fig. 2, 9 RV, ext. lat. (paratype. OS 12972. 570/am long); fig. 3, Cf LV. ext.

lat. (paratype, OS 12973, 550/u.m long).

Scale A (100/am; X130), figs. 1-3.

Stereo-Atlas of Ostracod Shells 14, 23 Muellerina hazeli (3 of 4)

Diagnosis:

Remarks:

Distribution:

A species of Muellerina characterised by small size, relatively thick shell, simple subovate outline
in dorsal view, and delicate ornament consisting of numerous discrete circular to ovate fossae with
several sharply defined narrow muri extending both above and below the muscle scar platform and
into the anterior field of the valve.

M. hazeli most closely resembles M. ohmerti Hazel, 1983, but is distinctly smaller, being
equivalent in size to the A-l instar of M. ohmerti , and has a distinctive, more delicately developed
ornament. M. hazeli rarely occurs sympatrically with M. ohmerti north of Cape Hatteras (above
35°N), but is more abundant in deeper water on the upper continental slope, whereas M. ohmerti is
a typical shelf species, most common at depths between 25 and 175m (Hazel 1970). M. hazeli is
both the smallest and the most southerly distributed extant species of Muellerina. It is believed to
have evolved from its parent species, M. ohmerti , during a high sea level stand in the late Pliocene
(Cronin & Coles, in prep.), and has since undergone little morphological change.

Recent of the Atlantic continental shelf and slope from the Florida Keys (24° 25'N) to off New
York at the head of Lydonia Canyon (40° 30'N). M. hazeli lives on the outer shelf and upper slope,
most commonly between 75 and 250m, having a maximum present-day depth range of 35 to 382m.
It is also present in Pleistocene sediments in cores off the eastern United States from 32° 04'N to
38° 22'N, and in Pleistocene outcrops in the Norfolk and Wilmington submarine canyons.

Explanation of Plate 14, 24

Fig. 1, 9 LV, int. lat. (holotype. OS 12971. 580/xm long); fig. 2, cf RV, int. lat. (paratype, OS 12974, 530/u.m long); fig. 3, cf car.,
dorsal (paratype, OS 12975, 570/um long).

Scale A (100/u.m; X130), figs. 1-3.

Stereo-Atlas of Ostracod Shells 14, 24

Muellerina hazeli (4 of 4)

Stereo-Atlas of Ostracod Shells 14, 22

Muellerina hazeli (2 of 4)

Stereo-Atlas of Ostracod Shells 14 (7) 25-28 (1987) Healdianella ? aremorica (1 of 4)

595.337.21 (113.51) (44 : 161.002.47) : 551.351 + 552.54

ON HEALDIANELLA ? AREMORICA CRASQUIN sp. nov.

by Sylvie Crasquin
(University of Lille, France )

Healdianella ? aremorica sp. nov.

University of Lille; France, ostracode Collection (COUL) no. 860, cf carapace.

[Paratypes: COUL nos. 861, 862, 863, 865, 2155].

Port Etroit Quarry (sample no. 85 MA 1), Laval syncline, Armorican Massif. France; lat. 47° 50'
54" N, long. 2° 39' 02"E. Sable Limestone, uppermost Tournaisian, Carboniferous.

From the latin aremoricus , Armorica, western province of Gaul.

University of Lille, France, ostracode collection (COUL) nos. 860 (holotype, cf car.: PI. 14, 26.
fig. 1), 862 ($ car.: PI. 14. 26, fig. 2), 863 (9 car.: PI. 14. 26, fig. 3). 865 (9 car.: PI. 14. 28, fig. 1),
861 (juv. car.: PI. 14, 28, fig. 2), 864 (juv. car.: PI. 14, 28, fig. 3), 2155 (cf car.: PI. 14, 28, fig. 4).
All from the Sable Limestone of type locality; uppermost Tournaisian, lower Carboniferous.

Holotype:

Type locality:

Derivation of name:
Figured specimens:

Explanation of Plate 14, 26

Fig. 1, Cf car., rt. lat. (holotype, COUL 860. 0.58 mm long); fig. 2, 9 car., rt. lat. (paratype. COUL 862. 0.58 mm long); fig. 3, 9 car..

rt. lat. (paratype, COUL 863, 0.60 mm long).

Scale A (200 fim; x 140), fig. 1; scale B (200 yu.m: x 85), figs. 2, 3.

Stereo-Atlas of Ostracod Shells 14, 27 Healdianella ? aremorica (3 of 4)

Diagnosis:

Remarks:

Distribution:

Small, smooth species (adults 0.52-0.63 mm long) doubtfully assigned to Healdianella.
Anterodorsal border straight; anterior border with a maximum of convexity located between 12
and lower 1/3 of valve height; ventral border is concave, with maximum concavity located in the
anterior 1/3 of valve length; posterior border broadly rounded with maximum convexity located
slightly below mid-height. In dorsal view, the carapace is laterally compressed in the medial
region. Overlap is weak.

Sexual dimorphism: heteromorphs have a more obtuse dorsal angle in lateral view, and in
dorsal view are wider behind a more pronounced median stricture. Tecnomorphs have a more
acute dorsal angle in lateral view and in dorsal view are virtually of equal width throughout, the
median stricture being poorly developed.

This species looks like Healdianella linevensis Tschigova, 1958 from the upper Tournaisian of the
Saratov-Leningrad area ( Trudy V.N.I.G.R.I. , 14). H. linevensis differs in having a smaller
length/height ratio, a more convex anterodorsal border and an anterior border which is not
laterally compressed.

H. ? aremorica is assigned to Headianella with doubt because in dorsal view its carapace is
laterally compressed in the medial region, a characteristic not observed in other species of that
genus.

Laval syncline, Armorican Massif, France: uppermost Tournaisian-lower Visean. lower Carbo-
niferous.

Explanation of Plage 14, 28

Fig. 1, 9 car., dors, (paratype. COUL 865, 0.58 mm): fig. 2. juv. car.. It. lat. (paratype, COUL 861. 0.47 mm long); fig. 3, juv. car., rt.

lat. (paratype, COUL 864, 0.40 mm long); fig. 4, cf car., dors, (paratype, COUL 2155. 0.55 mm long).

Scale A (200 fim; x 125), figs. 1, 3, 4; scale B (200 yum: x 100), fig. 2.

Healdianella ? aremorica (2 of 4)

Stereo-Atlas of Ostracod Shells 14, 26

Stereo-Atlas of Ostracod Shells 14, 28

Healdianella ? aremorica (4 of 4)

Stereo-Atlas of Ostracod Shells 14 (8) 29-32 (1987)
595.337.14 (116.331) (65 : 161.004.35) : 551.35

Maghrebeis tuberculata ( 1 of 4)

Derivation of name:
Diagnosis:

ON MAGHREBEIS TUBERCUEATA MAJORAN gen. et sp. nov.

by S. Majoran

(Department of Historical Geology and Palaeontology, University of Uppsala, Sweden)

Genus MAGHREBEIS gen. nov.

Type-species: Maghrebeis tuberculata sp. nov.

From the North African province of Maghreb (including Morocco, Algeria and Tunisia).
Carapace small, subtriangular, inequivalved. Left valve larger, overhanging posterior, ventral and
anterior margins of right valve. Ventral margin convex, converging posteriorly with straight dorsal
margin. Thick, swollen ridge runs along evenly rounded anterior margin, denticulated prominently
only on right valve, seldom and only feebly on left valve. Caudal process triangular, pointed at
mid-height and armed with a swollen ridge. Ornament polymorphic. Ventromedian and
dorsomedian areas bear pronounced lobate tubercles. Lateral surface variously pitted, ventral
surface with 4-5 fine longitudinal ribs. Hinge ear of left valve is large, forms a thick, hook-like
protuberance that overlaps right valve. Circular eye tubercle and ovate adductor muscle tubercle
prominent. Hinge amphidont/heterodont; right valve has large posterior tooth and strong anterior
tooth with a large, spherical distal part fused to a smaller proximal part.

Similar outline to Veenia Butler & Jones, 1957 and Veeniacythereis Griindel. 1973. which differ by
being larger, having 3 longitudinal ridges and lacking the curved, left hinge ear of Maghrebeis. The
left hinges of all three are similar, but the right hinge of Maghrebeis differs in having a modified
anterior tooth. Veenia also differs by its usually more pointed caudal process, and Veeniacythereis
by its feeble or absent subcentral tubercle.

Cythereis? sp. of Rosenfeld & Raab (Bull. geol. Surv. Israel, 62. pi. 2. figs. 45-46. 1974;
upper Cenomanian, Israel) probably belongs to Maghrebeis since it differs only by its smooth

Remarks:

Explanation of Plate 14, 30

Fig. 1, 9?car., It. lat. (holotype, PMAL1, 500/u.m long); fig. 2, 9? car., dors.. (PMAL2, 530/xm long); fig. 3. 9? RV, int. lat. (PMAL3.
500/j.m long); fig. 4, 9 ? LV, int. lat., (PMAL4, 510/xm long). Scale A (lOO^m; xl30), figs. 1-4.

Stereo-Atlas of Ostracod

Remarks: (cont.)

Holotype:
Type locality:
Derivation of name:
Figured specimens:

Diagnosis:

Distribution:

Shells 14, 31 Maghrebeis tuberculata (3 of 4)

surface and possibly having one less dorsal tubercle. Also possibly congeneric is Cythereis
lindiensis Bate, 1969 as reported by Grosdidier ( Revue Inst. fr. Petrole., 28. pi. 13. fig. 104. 1973),
which appears to be slightly larger and has more irregular dorsomedian and ventromedian
tuberculation, a less pronounced subcentral tubercle and a smooth surface. Further differences are
revealed by the original description and re-illustrated type material of C. lindiensis (Bate &
Mellish, Stereo-Atlas Ostracod Shells, 13, 59-62. 1986). Another externally similar, considerably
larger species is Grosdidier's Cythereis gr. malzi Bischoff. 1963 (Revue Inst. fr. Petrole, 28, pi. 14.
fig. 105, 1973).

Maghrebeis tuberculata sp. nov.

Palaeontological Museum, University of Uppsala, Sweden, no. PMAL1, 9? carapace.
Approx. 14km SW of Tocqueville, Algeria (approx, lat. 35° 52'N, long. 4° 55'E); Cenomanian.
Latin, from the prominent dorsomedian and ventromedian tubercles.

Palaeontological Museum, University of Uppsala. Sweden, nos. PMAL1 (holotype. 9? car.: PI.
14, 30, fig. 1), PMAL2 (9? car.: PI. 14, 30, fig. 2). PMAL3 (9? RV: PI. 14. 30. fig. 3). PMAL4 (9?
LV: PI. 14, 30, fig. 4), PMAL5 (9? car.: PI. 14, 32, fig. 2). PMAL6 (9? car.: PI. 14, 32, fig. 1),
PMAL7 (cf? car.: PI. 14, 32, figs. 3, 4). All from the type locality and horizon.

Maghrebeis with fine network of small pits, and 3 smooth, lobe-like tubercles respectively on
dorsomedian and ventromedian regions. Two additional, smaller tubercles vertically arranged on
posteromedian area. Swollen anterior and posterior ridges, pronounced eye tubercle, and ovate,
adductor muscle tubercle are all smooth as are also some narrow, longitudinal fields on ventral
surface. Ornament polymorphic with respect to size and configuration of dorsomedian and
ventromedian tubercles, strength of anterior and posterior ridges, and presence of denticles along
ventral section of caudal process. Shape differences might reflect sexual dimorphism: one type
being dorsoventrally and laterally more compressed (= cf?)-
Uppermost Albian (or lower Cenomanian) to middle Cenomanian of N Africa.

Explanation of Plate 14, 32

Fig. 1, 9? car., rt. lat. (PMAL 6, 500/um long); fig. 2, 9? car., vent., showing pitted surface with smooth, narrow, longitudinal fields
(PMAL5, 500/u.m long); figs. 3-4, cf? car. (compressed morph), (PMAL7, 500/xm long): fig. 3, car., rt. lat.; fig. 4. detail of
dorsomedian tuberculation. Scale A (100pim; x!30), figs. 1-3; scale B (lOOpun; x280), fig. 4.

Stereo-Atlas of Ostracod Shells 14, 32

Maghrebeis tuberculata (4 of 4)

Stereo-Atlas of Ostracod Shells 14 (9) 33-36 (1987) Howeina camptocytheroidea ( 1 of 4)

595.337.14 (118.22 + 119.9) (520 : 161.140.41 + 520 : 161.140.42) 551.351

ON HOWEINA CAMPTOCYTHEROIDEA HANAI

by Noriyuki Ikeya & Ellen Compton-Gooding
(Shizuoka University, Shizuoka, Japan & U.S. Geological Survey, Reston, V A)

Genus HOWEINA Hanai, 1957

Type-species (by original designation): Howeina camptocytheroidea Hanai, 1957
Diagnosis: Ovate Cytheruridae, right valve overlapping on dorsal margin, left valve overlapping on ventral

margin. Greatest height anterior, ventral margin nearly straight with a slight alate projection, eye
tubercle indistinct. Inner margin has modified S-shape along posterior margin.

Remarks: Howeina resembles Semicytherura Wagner, 1957 (see Whittaker, Stereo-Atlas Ostracod Shells, 2,

69-92, 1974); some might consider them synonymous since both have S-shaped posterior inner
margins, but the validity of this criterion for recognising Semicytherura is questionable. In any
case, right valves of Howeina have a large elongate anterior tooth, a knob-like posterior tooth and
a smooth median element; in Semicytherura anterior and posterior teeth of the right valve are
crenulate or have 2-3 knob-like projections, and the median element is smooth in the center with
sockets at its ends.

Howeina camptocytheroidea Hanai. 1957

1957 Howeina camptocytheroidea sp. nov. T. Hanai, J. Fac. Sci. Tokyo Univ., sec. 2, 11, 22-23, pi. 3, figs. 4a-c, text-figs. 5a, b.

1961 Howeina camptocytheroidea Hanai; T. Hanai, ibid., 13, 358, text-fig. 2, figs. 5a, b.

1971 Howeina camptocytheroidea Hanai; K. Ishizaki, Tohoku Univ. Sci. Rept., 2nd ser., (Geol.), 43, 79-80, pi. 2, fig. 21.

1977 Howeina camptocytheroidea Hanai; T. Hanai et al., Bull. Univ. Mus. Tokyo, 12, 56. pi. 3, figs. 1-7.

Explanation of Plate 14, 34

Fig. 1, cf LV, ext. lat. (IGSU-0-122, 618 pm long); fig. 2, cf car., ext. dors. (IGSU-0-126. 653 yum long); fig. 3, cf RV, ext. lat.
(IGSlJ-0-123, 613 pm long); fig. 4, 9 car., ext. vent. (IGSU-0-121. 605 yum long); fig. 5, 9 RV, ext. lat. (IGSU-0-124, 625 pm
long). Scale A (100 yum; x 100), figs. 1-5.

Type locality:
Figured specimens:

Diagnosis:

Remarks:

Stereo-Atlas of Ostracod Shells 14, 35 Howeina camptocytheroidea (3 of 4)

Holotype: University Museum, University of Tokyo, Tokyo, Japan, no. UMUT-CA-2612. 9 right valve.

[Paratypes: nos. UMUT-CA-2613-2615]

Upper Pliocene Setana Formation at Kaigarazawa, about 500m W of Nishinosawa, Kuromatsu-
nai, Suttsu-gun, Hokkaido (lat. 42° 39' 37"N, long. 140° 17' 37"E).

Institute of Geosciences, Shizuoka University (IGSU) nos. 0-121 (9 car.: PI. 14, 34, fig. 4), 0-122
(Cf LV: PI. 14, 34, fig. 1), 0-123 (cf RV: PI. 14, 34, fig. 3; PI. 14, 36, fig. 7), 0-124 (9 RV: PI. 14,
34, fig. 5; PI. 14, 36, figs. 4-6), 0-125 (9 LV: PI. 14, 36, figs. 1-3), 0-126 (cf car.: PI. 14, 34, fig. 2).
0-121 is a Recent specimen from Mutsu Bay, northern Honshu (lat. 41° 20'N, long. 140° 55'E).
0-122-126 are from the type locality; 0-122,123 are disarticulated valves of the same individual.
The valve surface has a pattern of pits that run parallel along the posterior and dorsal margins and
somewhat longitudinally in the center of the valve. The anterior and posterior areas have a pattern
of irregular polygons delineated by fine ridges. The flat ventral margin has a series of ridges that
run parallel to it. The posteroventral area is slightly depressed behind the alate projection.
H. higashimeyaensis Ishizaki, 1971, H. leptocytheroidea (Hanai, 1957), and H. neoleptocytheroidea
(Ishizaki, 1966) each possess a distinctive pattern of prominent ridges and varying degrees of
reticulation. They also have caudal processes that are more obvious than that of H.
camptocytheroidea. Specimens illustrated by McDougall, Brouwers & Smith (Bull, U.S. Geol.
Surv., 1598, 56, pi. 10, figs. 1,2, 1986) from Prudhoe Bay, Alaska, as Cytherura sp. B and
Cytherura sp. C., appear to be very similar to H. camptocytheroidea.

A cold water species, H. camptocytheroidea is currently living in Suttsu and Uchiura Bays,
southern Hokkaido; Aomori and Mutsu Bays, Aomori Prefecture; and Otsuchi Bay, Iwate
Prefecture. Late Pleistocene occurrences: the Nopporo Fm. Hokkaido; Shibikawa and Anden
formations, Akita Prefecture; Hashidate Fm. Ishikawa Prefecture; and Jizodo, Yabu and
Kiyokawa formations in Chiba Prefecture. In the late Pliocene, it occurs at the type locality; the
Tomikawa Fm. Hokkaido; the Hamada Fm. Aomori Prefecture; and the Junicho Fm. Toyama

Prefecture.

Explanation of Plate 14, 36

Fig. 1-3, 9 LV (IGSU-0-125, 650 yu.rn long): fig. 1, int. lat.; fig. 2, post, hinge; fig. 3, ant. hinge; figs. 4-6, 9 RV (IGSU-0-124): fig. 4,
inf. lat., fig. 5, ant. hinge; fig. 6, post, hinge; fig. 7, cf RV, int. muse, sc., dorsal is to right (IGSU-0-123).

Scale A (100 yum; X 100), figs. 1, 4; scale B (100 yum; x 160), figs. 2, 3, 5, 6; scale C (10 yum; x 540), fig. 7.

Distribution:

Stereo-Atlas of Ostracod Shells 14, 36

Howeina camptocytheroidea (4 of 4)

Stereo- Atlas of Ostracod Shells 14, 34

Howeina camptocytheroidea (2 of 4)

Stereo-Atlas of Ostracod Shells 14 (10) 37-40 (1987)
593.337.14 (116.333.3) (492:161.005.50) : 551.35

Spinoleberis eximia ( 1 of 4)

ON SPINOLEBERIS EXIMIA (BOSQUET)

by J. F. Babinot & J. P. Colin

(Universite de Provence, Marseille and Esso Production Research- European Lab., Begles, France)

Genus SPINOLEBERIS Deroo, 1966
Type-species: Cythere eximia Bosquet, 1854 (by original designation).

Diagnosis: Small-sized trachyleberidid (less than 650/am) characterized by a well-marked hemispherical,

sub-central tubercle; ventral ridge reduced to a strong posterior spinose tubercle and a median
lamellar spine; strong spinose tubercle present at the postero- dorsal angle and a vertical spine on
the middle part of the dorsal margin. A ridge connects the eye-tubercle and the sub-central
tubercle; a weak longitudinal median ridge may be present; anterior margin bordered by two rows
of strong spines. Surface of the valves smooth to very finely reticulate mostly on the anterior half.
Sexual dimorphism distinct, males being longer than females. Amphidont hinge. Anterior
marginal zone of medium width with about 20 straight pore canals. Muscle scars: three small scars
disposed in a V-shape or a V-shaped scar with an additional round scar above the posterior
branch; four adductor scars, the upper one being divided into two, the one below into three.

Remarks: The genus Spinoleberis is relatively common in the late Cretaceous of Western and Central

Europe. Typical species are restricted to the Campanian-Maastrichtian. Cenomanian and
Turonian species such as S. petrocorica (Damotte, Rev. Micropal , 14, 1, 1973), S. krejcii Pokorny
( Acta Univ. Carolinae Geoi, 4, 1968) and S. ectypus Babinot ( Geobios , 6, 1, 1973) have a rather
different morphology; they are deeply reticulate and do not display the characteristic spinose
tubercles. Species attributed to the genus Spinoleberis by Donze (1970), have been recently placed
in the newly errected genus Navarracy there Colin & Rodriguez-Lazaro ( Stereo-Atlas Ostracod
Shells, 13, 63-66, 1986).

Fig. 1, $ car., ext. rt. lat. (20648-49, 560 /xm long)
560 p.m long). Scale A (250 pm: xllO), figs.

Explanation of Plate 14, 38

fig. 2, cf LV, ext. lat. (20646-47, 550/u.m long); fig. 3, cf RV, ext. lat. (20642-43.
1-3.

1854

1936

1958

1966

1966

1983

Type locality:
Figured specimens:

Stereo-Atlas of Ostracod Shells 14, 39 Spinoleberis eximia (3 of 4)

Spinoleberis eximia (Bosquet, 1854)

Cythere eximia n. sp. J. Bosquet, Verhandel. geol. beschr. kaart Nederland , 2, 106, pi. 7. figs. 6a-d.

Cythereis eximia (Bosquet); J. E. van Veen, Nat. hist. Maandbl., 25, 11-12, 26, pi. 7, figs. 1-6.

Cythereis eximia (Bosquet); H. Howe & L. Laurencich, Introduction to the study of Cretaceous Ostracoda , 196-197.
Spinoleberis eximia (Bosquet); G. Deroo, Meded. geol. Sticht., C, 2, 2, 165-166, pi. 6, figs. 72-74, pi. 26, figs. 822-824.
Cythereis eximia (Bosquet); E. Herrig, Paldont. Abh., A, 2, 801-802. pi. 18, figs. 1-10, pi. 19, fig. 1.

Spinoleberis eximia (Bosquet); B. Clarke, Mitt. Geol. -Paldont. Inst. Univ. Hamburg, 54, 110-111, pi. 7, figs. 11-12.
Holotype: Material deposited in the collections of the Institut Royal des Sciences Naturelles de Belgique,

Brussels under the reference “Cretace Ostracodes 87 Arthr. Sec. I, Cret.”, slide no. 44.
Late Maastrichtian of St. Pietersburg, near Maastricht, southern Limburg, the Netherlands.
These are deposited in the collections of Esso Production Research - European Laboratories at
Begles, France and the numbers all carry the prefix EPR-E. EPR-E 20648-49 (2 car. : PI. 14. 38.
fig. 1), 20646-47 (Cf LV: PI. 14, 38, fig. 2), 20642-4 3 (cf RV: PI. 14, 38, fig. 3). 20650-51 (2 LV:
PL 14, 40, fig. 1), 20654-55 (cf car.: PI. 14, 40, fig. 2), 20816-17 (2 RV: PI. 14, 40, fig. 3). All
figured specimens are from Puits Maurits (250.5m), Maastricht, southern Limburg, The
Netherlands; late Maastrichtian.

As for the genus. The surface of the muri of the reticulation is very finely pitted. A little knob
occurs in the middle of each mesh.

Deroo (1966), illustrated several species of the genus Spinoleberis in the type Maastrichtian. Most
of the species are very similar to S. eximia and therefore extremely difficult to differentiate.
Whether they are different species or merely ecotypes is highly questionable. These are
S. eximioides (van Veen) and S. pseudoeximia Deroo. Cythereis symmetrica van Veen, 1936 (Nat.
Hist. Maandbl., 25, 1 1—12) is considered by Deroo (1966) and Clarke (1983) to be a juvenile of
S. eximia, by Howe & Laurencich (1958). (1965) to belong in S. tuberosa (Jones & Hinde) and by
Szczechura (1965) to belong in S. spinifera (van Veen).

Late Maastrichtian of the Netherlands and Belgium. Early to late Maastrichtian of Germany.

Diagnosis:

Remarks:

Distribution:

Explanation of Plate 14, 40

Fig. 1, 2 LV, int. lat. (20650-51, 575/xm long); fig. 2, cf car., ext. dors. (20654-55, 555p.m long); fig. 3, 2 RV, int. lat. (20816-17,
545/u.m long). Scale A (250/u.m; XllO), figs. 1-3.

Spinoleberis eximia (2 of 4)

Stereo-Atlas of Ostracod Shells 14, 40

Stereo-Atlas of Ostracod Shells 14, 38

Spinoleberis eximia (4 of 4)

Stereo-Atlas of Ostracod Shells 14(11) 41-44 (1987) Kovalevskiella caudata (1 of 4)

(118.21) (44 : 162.001.44) : 551.312.4 4- 552.54

ON KOVALEVSKIELLA CAUDATA (LUTZ)

by P. Carbonel, J.-P. Colin & L. Londeix
( University of Bordeaux, Talence & Esso Production Research, Begles, France)

Kovalevskiella caudata (Lutz, 1965)

1965 Gomphocythere caudata sp. nov. A. K. Lutz, Geol. Jahrb., 82, 311, text-fig. 27, pi. 13, figs. 1, 2.

1969 Cordocythere caudata (Lutz); G. Carbonnel & S. Ritzkowski, Arch. Sci. (Geneve), 22(1), 60.

1980 Kovalevskiella caudata (Lutz); J.-P. Colin & D. Danielopol, Paleobiol, continent., 11(1), 32, 37, fig. 17.

1985 Kovalevskiella caudata (Lutz); P. Carbonel, Bull. Centres. Rech. Explor.-Prod. Elf- Aquitaine, Mem. 9, pi. 90, figs. 7-10.

1986 Kovalevskiella caudata (Lutz); P. Carbonel, J.-P. Colin, D. L. Danielopol & L. Londeix, Geobios, 19(6), pi. 1, figs. 4-7.

Holotype:
Type locality:

Figured specimens:

Diagnosis:

Bundesanstalt fur Bodenforschung, Hanover, no. 5421, LV. [Paratype: no. 5420, LV]

Road cut between Undorf and Nittendorf near Regensburg. Bavaria, Federal Republic of
Germany; Tortonian, Late Miocene; freshwater molasse.

Dept. Geol. & Oceanography, Univ. Bordeaux I, CO nos. 5103 (LV; PL 14, 42. figs. 1-3; PI. 14.
44, fig. 1), 5104 (RV: PI. 14, 42, figs. 4-6), 5105 (car.: PI. 14, 44, figs. 4, 5), 5106 (LV juv.: PI. 14.
44, fig. 2), 5107 (RV juv.: PI. 14, 44, fig. 3), 5108 (LV juv.: PI. 14, 44. fig. 6). Aquitanian,
Miocene, of Le Moras, near Labrede, Gironde, France; lat. 44° 41 'N, long. 0° 34'W. Original,
German material could not be photographed.

Carapace subrectangular, rounded anterior and posterior extremities; ornament typical of genus:
regularly disposed pustules. Well developed sulcus; large brood pouch. RV larger than LV; both
cardinal hinge elements on LV trilobate. No sexual dimorphism. Tw'o strong denticles on the
posteroventral part of LV.

Explanation of Plate 14, 42

Figs. 1-3, LV (CO 5103, 430/xm long); fig. 1, ext. lat.; fig. 2, int. lat. hinge; fig. 3, int. lat. Figs. 4-6. RV (CO 5104, 433/am long); fig. 4,
int. lat.; fig. 5, ext. lat.; fig. 6, int. lat. hinge. Scale A (200/luti; xl35), figs. 1, 3-5; scale B (200/i.m; x205), figs. 2, 6.

Stereo-Atlas of Ostracod Shells 14, 43 Kovalevskiella caudata (3 of 4)

Remarks: Like other Kovalevskiella, K. caudata is parthenogenetic. It differs by the presence of 2 strong denticles on

the posteroventral part of the left valve. Denticles are also present in larval stages, but only in the right valve.
In the French locality studied, this species lived in a lagoonal to lacustrine environment (in oligo-to
mesohaline waters) on very fine grained, marl bottom sediment. K. caudata is associated with Neocyprideis
aquitanica Moyes or with Candonopsis and Limnocy there , and always with poorly diversified faunas. When
the waters become fresh and more stable, Kovalevskiella disappears. Lutz (1965) described K. caudata from
coaly marls in a freshwater molasse deposit. Its epibenthic life-style is very different to the hypogean or
interstitial habitats of Recent Kovalevskiella (Colin & Danielopol 1980; Carbonel et al., 1986).

Distribution: Miocene (Tortonian) near Regensburg, Germany (Lutz 1965); Miocene (Aquitanian) near Bordeaux, France

(Carbonel 1985; Carbonel et al. 1986).

Text-fig. 1. Size dispersion of 116 left and
right valves of K. caudata from Le Moras,
near Labrede, Gironde, SW France.

Explanation of Plate 14, 44

Fig. 1, LV ext. dors. (CO 5103, 430p,m long); fig. 2, LV juv.-l, ext. lat. (CO 5106, 340/u.m long); fig. 3, RV juv. -2, ext. lat. (CO 5107
285/xm long). Figs. 4, 5, car. (CO 5105, 420/u.m long): fig. 4, ext. dors.; fig. 5, ext. vent. Fig. 6, LV juv. -3, ext. lat. (CO 5108, 215pim
long). Scale A (200/xm; xl35), figs. 1-6.

250

Adults

’ ..-U*

I •>

¥ •

A:'

200 '

150 -

LV

RV

300

350

Length (um)

450

Stereo-Atlas of Ostracod Shells 14 (12) 45-48 (1987) Calocaria maurae (1 of 4)

595.339.1 (113.333) (64 : 162.008.31) : 551.35 + 552.52

ON CALOCARIA MAURAE VANNIER gen. et sp. nov

by Jean Vannier
(University of Rennes, France)

Derivation of name:

Genus CALOCARIA gen. nov.

Type-species: Calocaria maurae sp. nov.

From the Greek karia , a walnut and kalos, beautiful; alluding to the shell shape and lateral
ornament. Gender feminine.

Diagnosis:

Myodocopid ostracode, oval in outline. Length 3.5mm; adults have a length:height ratio of
approximately 1.1. Prominent anterior rostrum and rostral incisure. Composite external
ornament: pattern of coarse ridges running obliquely to the ventral and dorsal margins, converging
towards the middle of the valve and connected ventrally to a continuous marginal ridge;
posteriorly to mid-length, ornament consists of a linear alignment of more and less coalescent
tubercles laterally merging with corrugations. Small arcuate muscle scar impression.

Remarks:

This new genus shares many similarities (rostrum, simple muscle scar impression, oval outline)
with other Silurian ‘cypridinids' and is tentatively included within this group. Nevertheless, the
composite ornament of Calocaria is comparable to that of Silurian ‘bolbozoids’ (see Siveter,
Vannier & Palmer, Palaeontology , text-fig. 4, in press , 1987) but neither the distinctive
anterodorsal bulb nor sulcus typical of that group occur in Calocaria. By its well developed
ornament Calocaria is distinguished from any other Silurian ‘cypridinids', which are mainly
smooth (see Siveter, Vannier & Palmer, Palaeontology , pis. 2, 3, 5, in press , 1987). As with
numerous myodocopids from the Silurian of Europe, most specimens of Calocaria maurae show
‘plastic' deformation of the valves, suggesting a rather thin, flimsy shell.

Explanation of Plate 14, 46

Figs. 1-3, RV (holotype, IGR 33100, 3055/u.m long): fig. 1, ext. lat. ; fig. 2, ext. vent, obi.; fig. 3, ornament of lateral surface.
Scale A (750/iun; xl8), figs. 1, 2; scale B ( 100/am; x75), fig. 3.

Stereo-Atlas of Ostracod Shells 14, 47 Calocaria maurae (3 of 4)

Calocaria maurae sp. nov.

Holotype:

Institut de Geologie, University of Rennes (IGR), France, coll. no. 33100; RV.
(Paratypes: IGR coll. nos. 33101, LV; 33103, RV; 33104, LV; 33106. LV],

Type-locality:

Siltstones and mudstones in the Talmakent section near Talmakent (sample TA 84452 of J. J.
Cornee collections, University of Aix-Marseille, France), Haut-Atlas, Morocco; lat. 31° 52’N,
long. 7° 45'W. Upper part of the Silurian; as determined by J. J. Cornee (work in progress).

Derivation of name:
Figured specimens:

From maura , Moorish, alluding to the region where this species occurs.

Inst, de Geologie, Univ. Rennes (IGR), coll. nos. 33100 (holotype, RV: PI. 14, 46, figs. 1-3; PI. 14,
48, figs. 1, 2) and 33101 (paratype, LV: PI. 14, 48, figs. 3-6). From type locality; latex casts.

Diagnosis:

Remarks:

As for the genus. Monotypic.

C. maurae is the first Silurian myodocopid to be described from Africa. In the type locality, it is
associated with numerous other myodocopid ostracodes, both ‘bolbozoids" and ‘cypridinids-. In
this respect, this fauna is comparable to that recently documented from organic-rich. Silurian
sediments of Britain and France (Siveter, et al ., op. cit . , in press). As a prelude to their further
systematic studies of Silurian myodocopids the latter authors have noted the occurrence of similar
myodocopid faunas in the same type of deposits (black to terrigenous mudstones) from
northwestern Europe (e.g. Ludlow Series in Wales and the Armorican Massif, France), eastern
Europe (Ludlow Series of Bohemia, Czechoslovakia) and now North Africa (herein). An outer
shelf to shelf margin or even shelf slope environment is inferred, from faunal and sedimentological
evidence, for the myodocopid occurrences in Europe (see Siveter. et al., op. cit., in press) and is

Distribution:

Acknowledgements:

also likely in the case of the Moroccan material.

At present, known only from the type locality.

To J. J. Cornee (University of Aix-Marseille) for allowing me to study his material and to the
Humboldt Foundation (Bonn) for my Research Fellowship at Hamburg University.

Explanation of Plate 14, 48

Figs. 1. 2. RV (holotype, IGR 33100, 3055/j.m long): fig. 1, ext. ant. obi.; fig. 2, ext. post. obi. Figs. 3-6, LV (IGR 33101, 3050/xm
long): fig. 3, ext. lat.; fig. 4, ext. vent, obi.; fig. 5, ext. ant. obi.; fig. 6, ext. post. obi. Scale A (75(Vm; xl8), figs. 1-6.

Calocaria maurae (2 of 4)

Stereo-Atlas of Ostracod Shells 14, 46

Stereo-Atlas of Ostracod Shells 14, 48

Calocaria maurae (4 of 4)

Stereo-Atlas of Ostracod Shells 14 (13) 49-56 (1987) Spinohippula esurialis (1 of 8)

595.336.13 (113.312) (437 : 161.013.49) : 551.35 + 552.52

ON SPINOHIPPULA ESURIALIS VANNIER, KRUTA & MAREK gen. et sp

nov.

by Jean Vannier, Miroslav Kruta & Ladislav Marek
(University of Rennes, France; Academy of Sciences, Prague, Czechoslovakia)

Derivation of name:
Diagnosis:

Genus SPINOHIPPULA gen. nov.

Type-species: Spinohippula esurialis sp. nov.

Alluding to velar spines and ressemblance with species of the tribe Hippulini. Gender feminine.
Medium sized glossomorphitine (adults < 1.2mm long). Lateral surface virtually lacks lobation: sulci only
expressed dorsally as two very poorly marked depressions, presumably representing S2 and S3. Strong velum
extending from near anterior cardinal corner to posteromedium or posterodorsal part of valve and bearing a
distinctive coarse denticulation along inner margin bordering a deep (maximum depth ventrally and
anteriorly) fissum-like laterovelar furrow. Dimorphism mainly expressed in females by a broad crescent-like
velar flange and a wide concave subvelar area (dolonal antrum) both ends of which connect with lateral
surface of valve. Inner velar spines converging towards middle of domicilium tend to reach the lateral surface
over the laterovelar furrow. Tecnomorphs have narrower velum reduced to a row of radiating velar spines,
with laterovelar furrow more open than in females; marginal sculpture unknown.

Remarks:

The inclusion of Spinohippula within the Glossomorphitinae (see R. Schallreuter. Palaeontographica A, 180.
1983) is justified by the occurrence of a strong velar sculpture in both females and tecnomorphs, and well
marked velar dimorphism. The shape of the velar flange in females is its most significant glossomorphitine
feature, consisting of a massive adventral sculpture (PI. 14, 52. fig. 2) high the domicilium. Comparable
features are in typical glossomorphitines such as Collibolbina collis collis (Schallreuter, 1964) (see R.

Explanation of Plate 14, 50

Figs. 1-4, § LV (holotype, NM L26073, 1188/um long): fig. 1, ext. lat. ; fig. 2, ext. vent. obi. (tilted 75°): fig. 3, ext. dors. obi. (tilted
45°), antero-vent. part of the valve. Scale A (300/u.m; x69), figs. 1-3; scale B (200/am: xll5), fig. 4.

Stereo- Atlas of Ostracod Shells 14, 51 Spinohippula esurialis (3 of 8)

Remarks: (cont.)

Schallreuter, op. cit., 1983) from the middle Ordovician of Baltoscandia or Gracquina hispanica (Born, 1918)
(see J. Vannier, Palaeontographica A, 193. 1986) from the Llandeilo of France and Spain.

Schallreuter {op. cit., 1983) divided the subfamily Glossomorphitinae into three tribes on the basis of
different types of velar dimorphism. Spinohippula shares strong similarities with representatives of the tribe
Hippulini, especially Hippula and Parahippula. Although no typical torus is observed (in addition to the velar
flange) in Spinohippula , the general morphology of its adventral region is strikingly comparable to that of
Hippula (subgenera H. ( Hippula ) and H. (Cetona)) and Parahippula. Schematically, the shape (crescent-like
sculpture), position and nature (main adventral sculpture with hollow spaces and/or radiating structures) of
the velum (Text-fig. 2) are three important characteristics similar in Spinohippula from the middle Ordovician
of Czechoslovakia, Parahippula from the middle Ordovician of United States, and numerous species of
Hippula from the Ordovician of Europe and North America (see R. Schallreuter & M. Kruta, N. Jb. Paldont.
(Mil), 8, 1980). In Spinohippula the row of velar denticulation (26 spines in holotype) (PI. 14. 50, fig. 4; PI.
14, 52, fig. 4) associated with a deep laterovelar furrow forms a semi-open peripheral groove (Text-fig. la) at
the junction of the lateral, marginal and velar surfaces. By comparison, H. (Cetona) (Text-fig. lb), an
example of a typical unitoral hippuline, exhibits a row of short, flattened tubule-like spaces (= part of torus)
connected to the velum, exactly at the same place as the semi-open groove in Spinohippula. Moreover,
openings of these hollows (13 in females of H. (C.) cetona cetona = half the number of velar spines in the
holotype of S. esurialis) are also impressed on the velar flange surface (cf. Text-figs, la & lb). Similar
comparisons could also be attempted with H. ( Hippula ) characterized by two tori (Text-fig. 2d). The velar
flange of Parahippula (Text-fig. lc) is considered by Kraft (Mem. geol. Soc. Am, 86, 1962) as a "hollow velate
frill formed of two layers continuous with the outer layer of shell wall’'. The interpretation of this “hollow
structure” is problematic, bearing in mind that two-layered structures frequently observed on silicified
specimens (see D. J. Siveter, Stereo-Atlas Ostracod Shells, 12 (10), pi. 54, fig. 4) may be the result of
diagenetic processes. Nevertheless, as stated by Kraft, "the logical structure of the carapace wall” of
Parahippula is a convincing argument for true hollow spaces within the velum (as reconstructed in Text-fig.

Explanation of Plate 14, 52

Figs. 1-4, $ LV (holotype, NM L26073, 1 188/u.m long): fig. 1, ext. dors. obi. (tilted 45°); fig. 2, ext. post. obi. (tilted 75°); fig. 3, ext
ant. obi. (tilted 75°), postero-vent. part of the valve; fig. 4, ext. post. obi. (tilted 55°). antero-vent. part of the valve.
Scale A (300/u.m; x69), figs. 1, 2; scale B (200/u.m; x 115), figs. 3, 4.

Stereo-Atlas of Ostracod Shells 14, 50

Spinohippula esurialis (2 of 8)

Stereo-Atlas of Ostracod Shells 14, 52

Spinohippula esurialis (4 of 8)

Stereo-Atias of Ostracod Shells 14, 53 Spinohippula esurialis (5 of 8)

Remarks: (cont.) lc). In this case: 1) the hollow spaces (13 in females; Text-fig. 2c) within the velar flange of Parahippula , and
2) its deep laterovelar furrow showing 12 secondary radiating tiny furrows on the velar flange, may represent
homologous structures of the 13 toral hollows of H. (Cetona) (Text-fig. 2b) and the semi-open laterovelar
groove of Spinohippula (Text-fig. 2a) respectively.

Text-fig. 1. Reconstructions of the adventral sculpture in three genera of the tribe Hippulini Schallreuter, 1983: A, Spinohippula
gen. nov.; B, Hippula-, C, Parahippula. All views represent medioventral cross-sections of valves (see Text-fig. 2). d = dolonal
antrum; do = domicilium; If = laterovelar furrow; Is = lateral surface; ms = marginal sculpture (row of spines); t = torus; v = velar
flange; x = probable hollows within the velar flange.

Spinohippula esurialis sp. nov.

National Museum, Prague (NM), Czechoslovakia, coll. no. L26073; 9 LV.

[Paratypes: NM, Prague, coll. nos. L26074, tecnomorph RV; L26075. 9 TV]. Casts of the holotype
and paratypes are in the Institute of Geology, University of Rennes, France.

Ejpovice (borehole), 10km E of Plzen, WSW of Prague, Bohemia, Czechoslovakia; approx, lat. 49° 47'N.
long. 13° 38'E. Sandstones, Skalka quartzite Dobrotiva (Llandeilo ?) ‘series’, Ordovician.

Latin, esurialis , hungry; referring to the teeth-like velar spines.

Holotype:

Type locality:
Derivation of name:

Explanation of Plate 14, 54

Figs. 1-3, tecnomorph RV (paratype, NM L26074, 838/Atn long): fig. 1, ext. lat.; fig. 2, ext. vent. obi. (tilted 45°); fig. 3, ext. post. obi.,
part of the valve. Scale A (300/u.m; x93), figs. 1, 2; scale B (200/u.m; xll5), fig. 3.

Stereo-Atlas of Ostracod Shells 14, 55

Spinohippula esurialis (7 of 8)

Text-fig. 2. Three genera of the tribe Hippulini Schallreuter, 1983. All lateral views of female right valves. A, Spinohippula esurialis
gen. et sp. nov., from the Dobrotiva series (Llandeilo ?) of Czechoslovakia, approximately x95. B. Hippula (Cetona) cetona cetona
(Schallreuter, 1964), from Backsteinkalk erratic boulders of northern Germany, middle Ordovician, approximately x80 (after
Schallreuter 1983, op. cit. pi. 3, fig. 1). C, Parahippula ventrospina (Kraft, 1962), from the middle Ordovician of Virginia, United
States, approximately x65 (after Kraft 1962, op. cit.. pi. 12, fig. 5). D. Hippula (Hippula) latonoda (Schallreuter, 1964), from the
Upper Viru series (Caradoc) of Baltoscandia, approximately xlOO (after Schallreuter 1983, op. cit., pi. 5, fig. 1).

Figured specimens: National Museum, Prague (NM), Czechoslovakia, coll. nos. L26073 (holotype, 9 LV: Pi. 14, 50, figs. 1-4; PI.

14, 52, figs. 1-4), L26074 (paratype, tecnomorph RV: PI. 14, 54, figs. 1-3), L26075 (paratype, 9 LV: PI. 14,
56, figs. 1-3). Silicone rubber casts of topotype specimens.

As for the genus. Monotypic.

The laterovelar furrow represents a major concavity, widely extended ventrally, but not connected to the
domiciliar cavity and protected from the outside by a row of spines; it could be interpreted as an external
botulus-like brood concavity. More likely, this groove is homologous to a cavum (see R. Schallreuter in R.
Maddocks (Ed.), Proc. 8th Int. Symp. on Ostracoda, Houston, Texas , 1983) or a fissum. Some tvaerenellids
exhibit an arcuate cavum which tends to be closed by spines in the same way as the laterovelar furrow of 5.
esurialis. Huckea huckea (see R. Schallreuter. Palaeontographica A, 149, pi. 9, figs. 1-4, 1975) shows a ventral
fissum positioned similarly to the furrow in S. esurialis. The exact function of the cavum (buoyancy control?)
or fissum is still unknown but might represent an attempt to lighten the shell.

At present known only from type locality.

To the Humboldt Foundation (Bonn) for my Research Fellowship at Hamburg University.

Diagnosis:

Remarks:

Distribution:
Ac know l edgem ents:

Explanation of Plate 14, 56

Figs. 1-3, 9 LV (paratype, NM L26075, 1107/Am long); fig. 1, ext. lat.; fig. 2, ext. dors, obi., postero-vent. part; fig. 3, ext. post, obi.,
medio-vent. part. Scale A (300/Am; x76), fig. 1; scale B (200/Am; xll5), fig. 2; scale C (100/Am; x250), fig. 3.

Stereo-Atlas of Ostracod Shells 14, 54

Spinohippula esurialis (6 of 8)

Stereo-Atlas of Ostracod Shells 14, 56

Spinohippula esurialis (8 of 8)

Stereo-Atlas of Ostracod Shells 14 (14) 57-64 (1987)
595.336.11 (113.331) (481 : 161.010.59) 551.35 + 552.54

Beyrichia siveleri ( 1 of 8)

ON BEYRICHIA (SAGENABEYRICHIA) SIVETERI POLLICOT

subgen. et sp. nov.

by Paul D. Pollicott
(University of Leicester, England)

Derivation of name:
Diagnosis:

Remarks:

Subgenus BEYRICHIA ( SAGENABEYRICHIA ) subgen. nov.

Type-species: Beyrichia (Sagenabeyrichia) siveteri sp.lnov.

Latin sagena , fish-net; alluding to the reticulate ornament of the lobes + the genus Beyrichia.
Beyrichia with reticulo-tuberculate lobal ornament. Crumina elongate and relatively well
assimilated with lobal area. Syllobium weakly cuspidate; anterior cusp slightly more prominant.
posterior often lacking. Syllobial groove low and often above a well developed callus. Zygal arch
lacking.

The lobal reticulation of B. ( Sagenabeyrichia ) is unique within Beyrichia. Moreover, the
occurrence of reticulation in an otherwise typical beyrichiine species has significance for
beyrichiacean phyllogeny, particularly in the relationship between amphitoxotidines and
beyrichiines. Henningsmoen ( Geol . Foren. Stockh. Forh., 86, 387-9, 1965) thought that
amphitoxotidines, with their typically tubulose velar frill, evolved from beyrichiines ( Beyrichia
subgenera). In contrast, Martinsson (Bull. geol. Instn. Univ. Uppsala, 42. 56. 1963) thought that a
stabilized surface reticulation within the amphitoxotidines was entirely foreign to typical
beyrichiines (although reticulation is known in atypical Beyrichiidae such as Bingeria: see A.
Martinsson Bull. geol. Instn. Univ. Uppsala., 41, 1962), a subfamily which, furthermore, he

Explanation of Plate 14, 58

Fig. 1-3, cf RV (PMO 116.231, 2.00 mm long): fig. 1, ext. lat. ; fig. 2, ext. vent.; fig. 3. ext. vent. obi. Figs. 4, 5, cf LV (PMO 116.232.

1.32 mm long): fig. 4, ext. dors, obi.; fig. 5, ext. lat.

Scale A (370 /xm; x 28), figs. 1-3; scale B (260 gm; x 40), figs, 4, 5.

Stereo-Atlas of Ostracod Shells 14, 59

Beyrichia siveteri (3 of 8)

1954

Remarks: (cont.) considered more ‘advanced’ by lacking a tubulose velum. Henningsmoen (op. cit.) thought that
reticulation in the beyrichiines was an undeveloped possibility and. if found, its occurrence would
indicate a possible beyrichiine derivation for the amphitoxotodines. The lobal reticulation of
Beyrichia (Sagenabeyrichia) supports his idea.

B. (Sagenabeyrichia) further differs from many typical B. (Beyrichia) species by its better
assimilated crumina, a feature which it has in common with species of B. (Simplicibeyrichia) ,
especially B. (S.) callifera and B. (S.) duplicicalcarata (both Martinsson op. cit., 1962). B.
(Sagenabeyrichia) differs markedly from B. (S.) globifera Martinsson, 1962 by its reticulation, lack
of a calcarine spine, its often well developed syllobial groove/callus and in having a long, better
defined preadductorial sulcus.

Beyrichia (Sagenabeyrichia) siveteri sp. nov.

Beyrichia ( Beyrichia ) cf. kloedeni McCoy 1846; G. Henningsmoen, Norsk, geol. Tidsskr.. 34, 40-43 (pars), pi. 2, fig. 7, 10-18,
pi. 3, figs. 2-7; ? pi. 2, fig. 9.

Holotype: Paleontologisk Museum, Oslo, Norway, PMO 116.233; 9 RV (broken posteriorly).

Coastal section, southern tip of Kommerspya (east side), Holmestrand, Norway. Steinsfjorden
Formation, ‘9c|T of Kiaer (Skr. Vidensk. Selsk. Kristiania I Mat. - Naturv. Kl. 1906 II, 596 pp.);
Wenlock Series, Silurian. Approx, lat. 59° 32'N, long. 10° 18'E.

After Dr. David J. Siveter, University of Leicester, England.

Paleontologisk Museum, Oslo, nos. PMO 116.231 (cf RV: PI. 14, 58, figs. 1. 2, 3), PMO 116.232
(Cf LV: PL 14, 58, figs, 4, 5, PI. 14, 62, fig. 1), PMO 116.233 ($ RV: PI. 14, 60. fig 1) PMO 116.234
(9 RV: PI. 14, 60, figs. 2, 3). PMO 116.235 (9 LV: PI. 14. 60, fig, 4, 5), PMO 116.236 (cf LV: PI.
14, 62, fig 2, 5), PMO 116.237 (cf LV: PI. 14, 62, fig. 3), PMO 116.238 (cf LV: PI. 14, 62, fig, 4),
PMO 116.239 (Cf LV: PI. 14, 64, fig. 1), PMO 116.240 (cf LV: PI. 14, 64, fig. 2), PMO 116.241 (cf
RV: PI. 14, 64, fig. 3).

Type locality:

Derivation of name:
Figured specimens:

Explanation of Plate 14, 60

Fig. 1, 9 RV, ext. lat. (holotype PMO 116.233, 2.00 mm long). Figs. 2, 3, 9 RV (PMO 116.234, 2.50 mm long): fig. 2, ext. vent.; fig.

3, ext. vent, detail of crumina. Figs. 4, 5, 9 EV (PMO 116.235, 2.48 mm long): fig. 4, ext. vent, obi.; fig. 5, ext. lat.
Scale A (390 /x m; x 25), fig. 1; scale B (286 /xm; x 35), fig. 3; scale C (470 /xm; x 22), figs. 2, 4, 5.

Stereo- Atlas of Ostracod Shells 14, 58

Beyrichia siveteri (2 of 8)

Stereo-Atlas of Ostracod Shells 14, 60

Beyrichia siveteri (4 of 8)

Stereo-Atlas of Ostracod Shells 14. 61

Beyrichia siveteri (5 of 8)

Figured specimens:
(com.)

Diagnosis:

Remarks:

Distribution:

All figured specimens are from the Steinsfjorden Formation (9c), Sjorvoll, Ringerike, except for
PMO 116.233 (holotype) and PMO 116.238, which are from the type horizon and locality. All
specimens are prepared by mechanical preparation techniques from limestone slabs.

As for the subgenus. B. (Sagenabeyrichia) is monotypic.

B. (S.) siveteri exhibits wide variation in both lobal reticulation and tuberculation. Most valves are
reticulate over the entire lobal area, but in a few specimens, reticulation is lacking on the anterior
lobe (possibly a feature of preservation?). Tuberculation varies from forms with extensive cover
(mostly adults) to those in which it is lacking (small tecnomorphs). Reticulation is relatively
smaller in larger forms, and tubercles are commonly restricted to a supra-velar field (PI. 14. 58.
figs. 1, 5).

Size variation of female adults is common within a single sample (see Text-fig. 1). This is
thought to reflect mixed populations (chronodemes and/or ecodemes) rather than a possible case
of precocious dimorphism (unknown in Beyrichiacea).

The Wenlock Series, Silurian of Norway. Collected from localities in the Steinsfjorden Formation
(see Worsley, D. (ed.), Nor. geol. unders. 384, 1982) at Ringerike (9b-9e of Kiaer, op. cit.) and
Holmestrand (9b-9c of Kiaer, op. cit.).

Explanation of Plate 14, 62

Fig. 1, cf LV, reticulation on syllobium (PMO 116.232, 1.32 mm long). Figs. 2, 5. cf RV (PMO 116.236. 2.32 mm long): fig. 2. ext.
lat . ; fig. 5, reticulation and tuberculation on syllobium. Fig. 3. Cf LV. ext. lat. (PMO 116.237, 2.48 mm long). Fig. 4, cf LV. ext.
lat. (PMO 116.238, 1.20 mm long).

Scale A (21 /urn; x 460), fig. 1; scale B (455 /um; x 23), figs. 2, 3; scale C (230 /urn; x 40). fig. 4; scale D (62 /urn; x 150). fig. 5.

Stereo-Atlas of Ostracod Shells 14, 63 Beyrichia siveteri (7 of 8)

All specimens from a 2cm band in 9cJ. Wenlock Series.
Kommersaya, Holmestrand.

.♦V

• • •

• % •

. .*r

• %

_l . I , 1 , L_

Tecnomorphs (83 specimens).
Females (4 specimens).

, I I I

600 800 1000 1200 1400 1600 1800 2000 2200 2400

Hinge length Cum)

Text-fig. 1. Size variation within B. (Sagenabeyrichia) siveteri from the Steinsfjorden Formation (9c of Kiaer op. cit.), Wenlock Series
at Kommers0ya, Ffolmestrand, Norway.

Explanation of Plate 14, 64

Fig. 1, cf LV, ext. lat. (PMO 116.239, 2.72 mm. long). Fig. 2, cf LV, ext. lat. (PMO 116.240. 2.68 mm. long). Fig. 3 Cf RV, ext. lat.

(PMO 116.241, 2.68 mm. long).

Scale A (545 /um; x 18), figs. 1-3.

Beyrichia siveteri (6 of 8)

Stereo-Atlas of Ostracod Shells 14, 62

Stereo-Atlas of Ostracod Shells 14, 64

Beyrichia siveteri (8 of 8)

Stereo-Atlas ofOstracod Shells 14 (15) 65-68 (1987)
595.337.14 (119.9) (415 : 162.011.51) : 551.351

Bythocythere intermedia ( 1 of 4)

ON BYTHOCYTHERE INTERMEDIA ELOFSON

by David J. Horne

(Geology Department, City of London Polytechnic)

Type specimens:
Type locality:

Figured specimens:

Bythocythere intermedia Elofson, 1938

1868 Bythocythere constricta Sars; G. S. Brady, Trans. Linn. Soc. Lond. , 26, (pars), 451-452, pi. 35. figs. 48-52 only (non pi. 35. fig.
47) (non Sars, 1866).

1938 Bythocythere intermedia sp. nov. O. Elofson, Ark. Zool., 30A, 10, text-figs. 14-21.

1983 Bythocythere intermedia Elofson; J. Athersuch. D. J. Horne & J. E. Whittaker, J. micropalaeontol. , 2, 72-73, text-figs. 1, 2,
3a-g, 4r-t, 5b; pi. 2, figs. 1-4.

The whereabouts of Elofson’s type material is not known.

The Mittskaren, outside the mouth of Gullmar Fjord, W. Sweden, approx, lat. 58°15'N, long.
11°30'E; Recent, marine, sublittoral.

British Museum (Nat. Hist.) nos. 1982.345 (cf LV; PI. 14, 68, fig. 1; copulatory appendage:
Text-fig. 1), 1982.346 ($ LV: PI. 14, 66, fig. 2; RV: PI. 14, 66, fig. 3), 1982.347 (cf LV: PI. 14,68.
figs. 2, 3), 1982.348 (cf LV : PI. 14, 66, fig. 1). All from Valentia, SW Ireland (approx, lat.
51°55'N, long. 10°20'W), taken from slides labelled “B. constricta" in the Norman Collection at
the British Museum (Nat. Hist.); nos. 1982.345-347 are from slide 1900-3-6-379, no. 1982.348 is
from slide 1911.11.8 M3725.

Diagnosis: Moderately large (750-850/T.m long) species of Bythocythere ; carapace moderately inflated,

greatest width a little behind mid-length. Greatest height well behind mid-length. Dorsal margin
convex in female, almost straight in male; ventral margin weakly sinuous in both sexes. Posterior
margin denticulate. Dorsomedian sulcus weak. Male copulatory appendage with a relatively large,
subtriangular distal process.

Explanation of Plate 14, 66

Fig. 1, cf LV, ext. lat. (1982.348, 840^m long); figs. 2, 3, $ (1982.346, 810 /am long): fig. 2, LV, ext. lat.; fig. 3, RV. ext. lat.

Scale A (lOO^um; x80), figs. 1-3.

Stereo-Atlas of Ostracod Shells 14, 67

Remarks:

Distribution:

Bythocythere intermedia (3 of 4)

Early records of B. constricta Sars from British waters are now believed to be referable to either B.
intermedia or B. zetlandica Athersuch, Horne & Whittaker, 1983 (see Horne, Stereo-Atlas
Ostracod Shells , 14, 69-72, 1987). neither of which possesses the deep median sulcus which is
characteristic of Sars’ species. B. zetlandica has a smooth posterior margin and is less elongate with
a generally less rounded lateral outline than B. intermedia. A closely similar Miocene species, B.
neerlandica Kuiper, 1918, is less elongate, less tapered anteriorly, and has a deeper dorsomedian
sulcus than B. intermedia.

Fairly common in sublittoral marine waters around British coasts, the southern North Sea, S
Norway and Sweden, and as far south as the Bay of Biscay.

Text-fig. 1 Bythocythere intermedia , male copulatory appendage (1982.345).

Explanation of Plate 14, 68

Fig. 1, cf LV, dors. (1982.345, 820 pm long); figs. 2, 3, cf LV (1982.347, 790 p.m long): fig. 2, int. lat. ; fig. 3, central muscle scar field.
Scale A (100/um; x80), figs. 1, 2; scale B (50p.m; x400), fig. 3.

Stereo-Atlas of Ostracod Shells 14, 66

Bythocythere intermedia (2 of 4)

Stereo-Atlas of Ostracod Shells 14, 68

Bythocythere intermedia (4 of 4)

Stereo- Atlas of Ostracod Shells 14 (16) 69-72 (1987) Bythocythere zetlandica (1 of 4)

595.337.14 (119.9) (411 : 162.002.61 + 415 : 162.011.51) : 551.351

ON BYTHOCYTHERE ZETLANDICA ATHERSUCH,
HORNE & WHITTAKER

by David J. Horne

(Geology Dept, City of London Polytechnic)

Bythocythere zetlandica Athersuch, Horne & Whittaker, 1983

1868 Bythocythere constricta Sars; G. S. Brady, Trans. Linn. Soc. Land., 26, (pars), 451, pi. 35, fig. 47 only ( non pi. 35, figs 48-52)
( non Sars, 1866).

1983 Bythocythere zetlandica sp. nov. J. Athersuch, D. J. Horne & J. E. Whittaker, J. micropalaeontol., 2, 73, text-figs 41-n, 5c, pi.
2, figs 5-8.

Holotype: British Museum (Nat. Hist.) no. 1982.350, 9 carapace and appendages.

|Paratype, no. 1982.351, cf carapace and appendages.)

Type locality: Unst Haaf (fishing grounds off Unst), Shetland, approx, lat. 61° 00'N, long. 1° 30'W; Recent,

marine, sublittoral.

Figured specimens: British Museum (Nat. Hist.) nos. 1982.350 (holotype, 9 LV: PI. 14, 70, fig. 2; RV: PI. 14, 70, fig.

3), 1982.351 (paratype, cf LV: PI. 14, 70, fig. 1), 1982.352 (cf LV: PI. 14, 72, fig. 1; copulatory
appendage: Text-fig. 1), 1982.353 (cf LV: PI. 14, 72, figs 2-3). All taken from slides labelled “ B .
constricta ” in the Norman Collection at the British Museum (Nat. Hist.): the holotype and
paratype are from slide no. 1900-3-6-379; nos 1982.352 and 1982.353. both from Valentia, SW
Ireland (approx, lat. 51° 55'N, long. 10° 20'W), are from slides 1900-3-6-379 and 1911.11.8. M3725
respectively.

Explanation of Plate 14, 70

Fig. 1, Cf LV, ext. lat. (paratype, 1982.351, 770 pi m long); figs. 2, 3. 9 (holotype, 1982.350, 790 jotm long): fig. 2. LV, ext. lat.; fig. 3,
RV, ext. lat.

Scale A (100 pun; x 80), figs 1-3.

Stereo- Atlas of Ostracod Shells 14, 71 Bythocythere zetlandica (3 of 4)

Diagnosis:

Remarks:

Distribution:

Moderately large (750-800 pm long) species of Bythocythere ; carapace strongly inflated, greatest
width a little behind mid-length. Dorsal and ventral margins virtually straight, converging
anteriorly, greatest height well behind mid-length. Posterior margin smooth. Dorsomedian sulcus
weak. Distal process of male copulatory appendate relatively long, with a convex anterior margin
and an almost straight posterior margin.

B. zetlandica was formerly confused with B. constricta Sars, which does not live in British waters;
Sars’ species has a characteristically deep dorsomedian sulcus, and the distal process of its male
copulatory appendage is more symmetrical and slender than that of B. zetlandica. A similar NW
European species, B. intermedia Elofson, 1938 (see Horne, Stereo-Atlas Ostracod Shells 14, 65-68,
1987), is more elongate than B. zetlandica and has a denticulate posterior margin and a more
rounded outline in lateral view.

A marine species found in sublittoral waters around British coasts, particularly in the north.

Text-fig. 1 Bythocythere zetlandica, male copulatory appendage (1982.352).

Explanation of Plate 14, 72

Fig. 1, cf LV, dors. (1982.352, 800 pun long); figs 2, 3 cf LV (1982.353, 780 pun long): fig. 2, int. lat.; fig. 3, central muscle scar field.
Scale A (100 pun; X 80), figs 1, 2; scale B (50 pun; X 400), fig. 3.

Stereo-Atlas of Ostracod Shells 14, 70

Bythocythere zetlandica (2 of 4)

Bythocythere zetlandica (4 of 4)

Stereo-Atlas of Ostracod Shells 14, 72

Stereo-Atlas of Ostracod Shells: Vol. 14, Part 1

CONTENTS

On Cathaycythere reticulata Whatley & Zhao gen. et sp. nov.; by R. C.
Whatley & Zhao Quanhong

On Sinocythere sinensis Hou; by R. C. Whatley & Zhao Quanhong
On Albileberis sinensis Hou; by Zhao Quanhong & R. C. Whatley
On Sinocytheridea impressa (Brady); by Zhao Quanhong & R. C. Whatley
On Pterygocythereis vannieuwenhuisei Brouwers sp. nov.; by E. M.
Brouwers

On Muellerina hazeli Coles & Cronin sp. nov.; by G. P. Coles & T. M.
Cronin

On Healdianella? aremorica Crasquin sp. nov.; by S. Crasquin
On Maghrebeis tuberculata Majoran gen. et sp. nov.; by S. Majoran
On Howeina camptocytheroidea Hanai; by N. Ikeya & E. Compton -
Gooding

On Spinoleberis eximia (Bosquet); by J. F. Babinot & J. P. Colin
On Kovalevskiella caudata (Lutz); by P. Carbonel, J. P. Colin & L. Londeix
On Calocaria maurae Vannier gen. et sp. nov.; by J. Vannier
On Spinohippula esurialis Vannier, Kruta & Marek gen. et sp. nov.; by J.
Vannier, M. Kruta & L. Marek

On Beyrichia (Sagenabeyrichia) siveteri Pollicott subgen. et sp. nov.; by
P. D. Pollicott

On Bythocythere intermedia Elofson; by D. J. Horne
On Bythocythere zetlandica Athersuch, Horne & Whittaker; by D. J. Horne

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