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STUDIES OF
CENTRAL AMERICAN PLANTS III
BY
PAUL C. STANDLEY
CURATOR OF THE HERBARIUM, DEPARTMENT OF BOTANY
AND
JULIAN A. STEYERMARK
ASSISTANT CURATOR OF THE HERBARIUM, DEPARTMENT OF BOTANY
BOTANICAL SERIES
FIELD MUSEUM OF NATURAL HISTORY
VOLUME 23, NUMBER 1
JANUARY 14, 1943
PUBLICATION 524
STUDIES OF
CENTRAL AMERICAN PLANTS III
BY
PAUL C. STANDLEY
CURATOR OF THE HERBARIUM, DEPARTMENT OP BOTANY
AND
JULIAN A. STEYERMARK
ASSISTANT CURATOR OF THE HERBARIUM, DEPARTMENT OF BOTANY
BOTANICAL SERIES
FIELD MUSEUM OF NATURAL HISTORY
VOLUME 23, NUMBER 1
JANUARY 14, 1943
PUBLICATION 524
THE LIBRARY OF THE
AUG 1 - 1049
UNIVERSITY OF ILLIfiOiS
PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS
580-5
STUDIES OF CENTRAL AMERICAN PLANTS III
PAUL C. STANDLEY AND JULIAN A. STEYERMARK
Most of the new species described on the following pages were
collected by the authors during recent exploration in Guatemala,
and are published here preparatory to their inclusion in the descrip-
tive Flora of Guatemala, whose preparation is now well advanced.
Because of present economic conditions, the descriptions and dis-
cussions of these new plants have been somewhat abbreviated but
they will be extended, if deemed necessary, when treated in that work.
PINACEAE
Juniperus Standleyi Steyermark, sp. nov. Frutex prostratus
vel depressus vel erectus, ramulis ultimis numerosis brevibus densis
confertis 1.1-1.4 mm. crassis 5-20 mm. longis; folia adulta ovata
vel ovato-rotundata cucullata, apice obtusa vel rotundata, 1.5-1.7
mm. longa, basi 0.8-1 mm. lata, medio depressa, dorso glandulosa,
marginibus crassis incurvis minute erosis vel subintegris; folia
juvenilia lanceolata subulata acuminata 4-5 mm. longa; bacca
coerulescens 7-9 mm. diam., pedicello brevi recurvo pendulo;
semina 3-4. Guatemala: Dept. San Marcos: Upper slopes of Volcan
de Tacana, alt. 4,100-4,400 meters, February, 1940, Julian A.
Steyermark 36137 (type in Herb. Field Mus.). Collected at various
localities in the departments of San Marcos and Huehuetenango,
and extending into Chiapas, Mexico.
This has been confused with J. mexicana Spreng., also native in
Guatemala, which is a tall or medium-sized tree with 1-2-seeded
fruit.
ARACEAE
Philodendron Rojasianum Standl. & Steyerm., sp. nov.
SubgenusEuphilodendron, Sect. Pteromischum. Epiphytica scandens,
ramulorum internodiis superioribus ca. 4 cm. longis et 8 mm. crassis;
petiolus ca. 16 cm. longus crassus, vagina ca. 1 cm. lata foliacea
persistente ca. 1-2 cm. infra apicem soluta atque ca. 1.5 cm. longe
protracta, apice obtusissima; lamina tenuiter coriacea rigidula late
oblongo-ovata ca. 21 cm. longa et 11-13.5 cm. lata, apice obtusa
vel subrotundata et breviter cuspidato-acuminata, basi lata breviter
lateque cordata, nervis primariis utroque latere ca. 10 angulo
3
4 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
latissimo abeuntibus tenuibus quam secundariis et tertiariis solem-
niter crassioribus; pedunculus crassus vix ultra 1.5 cm. longus;
spatha oblongo-linearis ca. 14 cm. longa prope basin 2 cm. lata
viridis, spadice viridi. Guatemala: Climbing on tree along a small
stream, Finca Pirineos, lower southern slopes of Volcan de Santa
Maria, between Santa Maria de Jesus and Calahuache", Dept.
Quezaltenango, alt. 1,300-1,500 meters, December, 1939, Julian A.
Steyermark 33244 (type in Herb. Field Mus.).
This may be only an extreme form of P. guttiferum Kunth, but
in that the petioles usually are relatively shorter and the ligule of
the petiole sheath much less developed. Also related is P. rigidi-
folium Krause of Panama, but that also has a less developed petiole
sheath. The ligule of the sheath in P. Rojasianum is sometimes as
much as 2.5 cm. long. The new species is named for Professor
Ulises Rojas, Director of the Jardin Botanico de Guatemala.
AMARYLLIDACEAE
Agave huehueteca Standl. & Steyerm., sp. nov. Subgenus
Littaea. Folia numerosa rosulata oblongo-ovata vel obovata ca.
35 cm. longa atque 10 cm. lata, vel minora majorave, suberecta vel
adscendentia, subincurva, plus minusve concava, laevia, crassa,
fibrosa, griseo-viridia vel glaucescentia, rigida; spina terminalis
conica vel elongato-conica 2.5-4 cm. longa basi ca. 6 mm. crassa
griseo-brunnea, interdum lucida, superne teres, inferne late sulcata;
margen folii late corneo-incrassatus griseus, dentibus plerumque
5-11 mm. longis complanatis griseis latissimis saepe curvis vel
recurvis; inflorescentia 3-5-metralis simplex spicata erecta, spica
densa elongata, bracteis numerosis, superioribus e basi lata fere
setaceis elongatis, floribus fere sessilibus fasciculatis; ovarium
fusiforme ca. 12 mm. longum et 4 mm. crassum; perianthii tubus
obconicus 5 mm. longus, segmentis anguste oblongis 2-2.5 cm.
longis obtusis; stamina longiexserta, antheris fere 2 cm. longis;
capsula late ovalis 2 cm. longa 12 mm. lata viridis breviter sulcata;
semina semiorbicularia ca. 4 mm. longa. Guatemala: Dept. Huehue-
tenango: Open limestone hillside, along Aguacatan road east of
Huehuetenango at km. 13-14, altitude about 1,950 meters, January
2, 1941, Paul C. Standley 82039 (type in Herb. Field Mus.). Dry
rocky hills north of Chiantla, 2,000-2,250 meters, January 6, 1941,
Standley 82588. Near crossing of Rio San Juan Ixtan, east of San
Rafael Pe"tzal, 1,730 meters, January 9, 1941, Standley 82857.
Perianth dark purplish green; filaments and style dark purplish
red. Only one other species of the subgenus Littaea has ever been
STUDIES OF CENTRAL AMERICAN PLANTS III 5
reported from Guatemala, A. dasylirioides Jacobi & Bouche", and
that is an altogether different plant with unarmed leaves. It is
quite possible that ultimately A. huehueteca will have to be united
with one of the Mexican species, but we have been unable to refer
it satisfactorily to any of the species of this subgenus listed by
Trelease from Mexico.
POLYGONAGEAE
Gymnopodium floribundum Rolfe, var. antigonoides (Rob.)
Standl. & Steyerm., stat. nov. Millspaughia antigonoides Rob. Bot.
Jahrb. 36, Beibl. 80: 14. 1905.
Triplaris melaenodendron (Bertol.) Standl. & Steyerm., comb,
nov. Vellasquezia melaenodendron Bertol. Fl. Guat. 40. pL 11. 1840.
T. Macombii Donn. Smith, Bot. Gaz. 19: 257. 1894.
NYCTAGINACEAE
Mirabilis pulchella Standl. & Steyerm., sp. nov. Herba vel
frutex erectus 30-100 cm. altus ramosus, ramis vetustioribus plus
minusve glaucis ad nodos incrassatis, novellis dense pilis plurilo-
cularibus villosis; petioli usque 1.5 cm. longi, foliorum superiorum
multo breviores, dense longivillosi; lamina late ovata usque oblongo-
ovata vel deltoideo-ovata 3-8.5 cm. longa 2.5-5.5 cm. lata acuta vel
acuminata, basi plus minusve inaequali truncata usque obtusa,
crassiuscula, utrinque subsparse pilis patentibus villosa, subtus paullo
pallidior; pedunculi axillares et in cymulas vel racemos terminales
parvos dispositi, usque ad 1.5 cm. longi sed saepius multo breviores
dense villosi; involucrum campanulatum vel in statu maturo fere
rotatum 7-10 mm. longum viride triflorum villosum, profunde
5-lobum, lobis triangulari-ovatis acutis vel acuminatus ciliatis;
perianthium coralloideum roseo-purpureum 1 cm. longum infundibuli-
formi-campanulatum glabrum vel glabratum; stamina 3 exserta;
fructus obovoideus ca. 5 mm. longus et 2.5 mm. latus obtuse 5-
costatus, basin versus angustatus, sat dense pilosulus, costis crassis
laevibus vel plus minusve corrugatis, sulcis rugulosis. Guatemala:
Dept. Zacapa, brushy rocky slope, near divide on the road between
Zacapa and Chiquimula, alt. 500-660 meters, October, 1940, Paul C.
Standley 73841 (type in Herb. Field Mus.). Dept. Chiquimula, near
the same locality as the preceding collection, Standley 73748. Dept.
Chiquimula, between Ramirez and Cumbre de Chiquimula, on road
between Chiquimula and Zacapa, 400-600 meters, Standley 74516.
6 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
The plant is frequent in the vicinity of the type locality, growing
in damp thickets or on brushy rocky slopes. It is rather handsome
and showy because of the abundance of rose-purple flowers, which
close around noon, as in other species of the genus. Except for the
widespread tropical Mirabilis violacea (L.) Heimerl, this is the first
Mirabilis of the subgenus Oxybaphus to be reported from Central
America. Its relationship is with M. suffruticosa Standl., of Oaxaca,
which has merely puberulent branches and rounded or obtuse invo-
lucral lobes.
PHYTOLACCAGEAE
Stegnosperma scandens (Lunan) Standl., comb. nov. Tri-
chilia scandens Lunan, Hort. Jam. 2: 320. 1814. S. halimifolium
Benth. Bot. Voy. Sulph. 17. 1844.
BERBERIDACEAE
Mahonia volcania Standl. & Steyerm., sp. nov. Frutex erectus
pauciramosus 1-2-metralis; folia breviter petiolata, foliolis 2-6-jugis
approximatis; foliola rigide coriacea sessilia ovalia usque oblongo-
ovalia vel late oblonga 2-4 cm. longa 1-1.7 cm. lata apice rotundata
vel late obtusa et spinoso-mucronata, basi rotundata vel late obtusa,
in toto margine spinoso-mucronato-serrata, serraturis parvis sub-
adpressis, glabra, supra lucida, venulis prominulis pallidis arete
reticulatis, subtus pallida, nervis venisque prominentibus reticulatis;
flores racemosi, racemis sat dense multifloris ca. 3 cm. longis breviter
pedunculatis, bracteis bracteolisque parvis inconspicuis acuminatis
vel interdum obtusis et denticulatis, pedicellis gracilibus sed rigidulis
usque 10 mm. longis; sepala exteriora late ovata 2 mm. longa;
petala lutea ca. 6-7 mm. longa; baccae immaturae sessiles glabrae.
Guatemala: Pine forest, slopes of Volcan de Agua along the trail
above Santa Maria de Jesus, alt. about 3,000 meters, February,
1939, Paul C. Standley 65221 (type in Herb. Field Mus.); same
region, January, 1937, J. R. Johnston 580. Region of Che"mal,
Huehuetenango, 3,300 meters, December, 1940, Johnston 1690.
This plant has been confused heretofore with M. fascicularis DC.,
which ranges from southern California southward to Veracruz. It
is closely related to that species but apparently uniformly distinct
in its small compact leaves, whose leaflets have small and appressed
teeth rather than the large spreading ones characteristic of M.
fascicularis.
Mahonia Johnstonii Standl. & Steyerm., comb. nov. Berberis
Johnstonii Standl. & Steyerm. Bot. Ser. Field Mus. 22: 140. 1940.
STUDIES OF CENTRAL AMERICAN PLANTS III 7
ANNONAGEAE
Annona primigenia Standl. & Steyerm., sp. nov. Arbor usque
10-metralis vel frutex altus, ramulis gracilibus glabris vel primo
sparse pilis brevibus adscendentibus indutis; folia modica breviter
petiolata membranacea in sicco fuscescentia, petiolo glabro 7-14 mm.
longo; lamina elliptica usque lanceolato-oblonga vel obovato-oblonga
8-14 cm. longa 3-6 cm. lata, acuta vel abrupte acutata, interdum
acuminata, basi subrotundata usque subacuta, supra glabra sub-
lucida, costa anguste impressa, subtus fere concolor saltern in statu
adulto glabra in axillis nervorum obscure domatiata, nervis laterali-
bus utroque latere ca. 10 teneris; inflorescentiae e medio internodii
exeuntes pluriflorae, pedicellis fructiferis glabris 1.5-3 cm. longis
rectis; fructus subglobosus 1.5-3 cm. diam., fere laevis vel interdum
distinctius areolatus, sparse puberulus vel fere glaber; semina pauca
vel numerosa lucida fusco-brunnea ca. 8 mm. longa. Guatemala
(Dept. Pete'n) : Gavilan, Fallabon-Yaxha road, March, 1933, C. L.
lAindell 2213 (type in Herb. Field Mus.), 2215. Uaxactun, April,
1931, H. H. Bartlett 12474. British Honduras: San Antonio, high
ridge, P. H. Gentle 569. Mexico: Tuxpena, Campeche, December,
1931, Lundell 1128.
Called Anonillo in Pete'n. The fruit is described as russet-
colored and edible. It is probable that the species is represented also
by Lundell 6651 from San Agustin, British Honduras. This speci-
men is in flower, the flowers being 1.5 cm. long, and like those of
A. reticulata L., except that they are less densely pubescent.
The relationship of A. primigenia is with A. reticulata but in
that, one of the important fruit trees of all Central America, the
fruits are 10 cm. or often much more in diameter. It seems reason-
able to assume that the small-fruited tree of the Yucatan Peninsula
is a distinct species.
Sapranthus megistanthus Standl. & Steyerm., sp. nov.
Arbor 9-metralis, ramulis novellis dense pilis mollibus subpatentibus
ochraceis tomentosis; folia crasse 5-6 mm. longe petiolata mem-
branacea, petiolo dense ochraceo-piloso; lamina oblongo-elliptica
10-14 cm. longa 5.5-7 cm. lata acuta vel obtusa, basi obtusa, supra
viridis molliter atque sat dense pilis albidis velutino-pilosa, subtus
paullo pallidior ubique dense velutino-pilosa, reticulo venularum
eleganter prominulo-reticulato; pedunculi oppositifolii crassi 1.5 cm.
longi tomentosi; sepala utrinque tomentella anguste lanceolato-
oblonga 2-2.5 cm. longa basi 7 mm. lata nervosa; petala maxima
atropurpurea, intus sparse puberula extus subsparse tomentella,
8 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
oblanceolato-oblonga 17-19 cm. longa ca. 7 cm. lata, apicem sub-
obtusum versus angustata, basin versus longe attenuata. Guate-
mala: On roadside bank, Estancia Grande, Dept. Guatemala, alt.
about 600 meters, December 8, 1938, Paul C. Standley 59219 (type
in Herb. Field Mus.).
Only one tree was observed. The flowers are pendent, dark
purple-brown, and have a very strong and offensive, carrion-like
odor. The type was determined originally as S. nicaraguensis Seem.,
but more careful examination shows that probably it represents a
perfectly distinct species. In that the petals are only half as large,
and the sepals are much shorter and relatively broader. The flowers
are twice as large as those of any other member of the genus.
MENISPERMACEAE
Hyperbaena vulcania Standl. & Steyerm., sp. nov. Frutex
scandens, ramulis subteretibus puberulis vel dense breviter patenti-
pilosulis, glabrescentibus; folia longipetiolata subcoriacea, petiolo
2-7 cm. longo et ultra pilosulo vel puberulo, serius glabrato, apice
paullo incrassato; lamina integra ovalis vel rotundo-ovalis usque
ovata vel rotundato-ovata vulgo 9-20 cm. longa atque 5.5-13 cm.
lata, apice obtusa vel rotundata et acuminata vel breviter acuminata,
basi rotundata usque breviter cordata, interdum obtusa, supra lucida
glabra, subtus brunnescens ad nervos pilosula vel puberula vel fere
omnino glabra, ima basi 5-nervia, costa supra basin utroque latere
nervos laterales vulgo 3 emittente, nervis venisque elevatis gracilibus,
venulis prominentibus laxe reticulatis, ultimis reticulum densum
prominulum efformantibus; flores feminei racemosi, racemis axillari-
bus pedunculatis vel sessilibus 7-12 cm. longis laxe paucifloris,
rhachi pedicellisque densiuscule pilosiusculis vel brunneo-puberulis,
pedicellis crassiusculis 6-15 mm. longis; sepala interiora 5 mm. longa
late ovata obtusa dense minute puberula, apicibus recurvis, exteri-
oribus minutis ovatis; carpella densissime breviter pilosa. Guate-
mala: Moist forest, a large vine over a tree, near Barranco Hondo,
above Las Lajas, Dept. Escuintla, alt. about 1,200 meters, January,
1939, Paul C. Standley 63878 (type in Herb. Field Mus.). Dept.
Quezaltenango, between Finca Pirineos and Finca Soledad, lower
southern slopes of Volcan de Santa Maria, 1,300 meters, Steyermark
33517. Dept. San Marcos, vicinity of Finca El Porvenir, southern
slopes of Volcan de Tajumulco, 1,300-1,500 meters, Steyermark
37944, 37343, 37535.
Related to H. hondurensis Standl., which occurs in northern
Guatemala but differs in having pistillate inflorescences with pedicels
STUDIES OF CENTRAL AMERICAN PLANTS III 9
only 2-4 mm. long or even shorter. Most of the collections cited
above are sterile, but probably belong to H. vulcania.
ROSACEAE
Hirtella media Standl., sp. nov. Arbor glabrata 10-20-metralis,
trunco usque 50 cm. diam., ramulis strigillosis vel fere glabris;
folia inter minora breviter petiolata coriacea, petiolo ad 4 mm.
tan turn longo; lamina elliptico-obovata usque elliptico-oblonga
saepius paullo supra mediam latissima, 4.5-10 cm. longa 2-5 cm.
lata, apice obtusa vel subacuta, basi cuneato-angustata et acuta,
supra glabra in sicco fuscescens glabra, costa prominente, subtus
fere concolor fere glabra sed saltern ad costam sparse strigosa;
inflorescentiae terminales thyrsoideo-paniculatae sat dense multiflorae
foliis paullo longiores, ramis dense minute sericeis vel glabratis,
interdum elongatis et recurvis, secundifloris, floribus breviter pedi-
cellatis vel sessilibus, bracteis ovatis parvis acutiusculis persistentibus
conspicuis; hypanthium obovoideum adpresse pilosulum, sepalis
reflexis late ovatis vel rotundo-ovalibus intus minute sericeis obtusis-
simis; petala late obovata alba vel lavendulacea glabra; stamina
3 longissime exserta; ovarium dense pilosum; drupa juvenilis oblongo-
obovoidea apice late rotundata sparse breviterque strigosa.
Honduras: Guarunta, Wispernini Camp, Dept. Colon, alt. 15-23
meters, March, 1938, Christine & Wolfgang von Hagen 1378 (type
in Herb. Field Mus.). Mexico: Lobani, Oaxaca, 1841-43, Liebmann
1751. Ubero, Oaxaca, alt. 30-90 meters, June, 1937, Llewelyn
Williams 9480.
Related to H. triandra Swartz, a common species of Central
America, which has larger long-acuminate leaves. H. media is
noteworthy for the large size the tree attains, according to data
supplied with three collections. Most species of the genus are shrubs
of only 2-3 meters, and but few of the species attain such a height
as 15 meters. The vernacular names used in Oaxaca are Jicaco
and Jicaco prieto.
LEGUMINOSAE
Aeschynomene eriocarpa Standl. & Steyerm., sp. nov.
Annua prostrata, caulibus dense breviter albido-pilosis, pilis paucis
longioribus viscidis patentibus intermixtis; stipulae parvae basi
non productae late ovatae acutae striato-nervosae; folia parva,
foliolis 3-4-jugis late oblongo-obovatis 5-8 mm. longis 3-5 mm.
latis, apice late rotundatis, supra glabris, subtus pilis longis albidis
subadpressis pilosis; racemi foliis longiores pauciflori, interdum
10 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
elongato-racemosi, rhachi valde geniculata dense albido-pilosa et
viscido-pilosa, bracteis late ovatis obtusis pilosis; calyx 3 mm. longus
dense breviter pilosus; corolla flava 7 mm. longa, vexillo sparse
pilosulo; legumen vulgo 3-articulatum densissime albo-tomentosum,
articulis isthmo angustissimo junctis semiorbicularibus. Guate-
mala: Zacapa, rocky slopes, Santa Rosalia, 2 miles south of Zacapa,
alt. 200 meters, October 7, 1939, Julian A. Steyermark 29313 (type
in Herb. Field Mus.).
Similar in most details to A. biflora (Mill.) Fawc. & Rendle,
but distinct from that and the several other related species in its
densely white-tomentose fruits, which bear no viscid hairs.
Aeschynomene tricholoma Standl. & Steyerm., sp. nov.
Annua erecta ramosa ca. 60 cm. alta, caulibus dense viscido-hispidis;
stipulae lineari-attenuatae basi productae striatae pectinato-
dentatae hispidulae; foliola ca. 23-juga lineari-oblonga 5-10 mm.
longa, obtusa vel subacuta et mucronata, apice denticulata, glabra,
costa valde excentrica, nervis subtus prominentibus; racemi elongati
laxi foliis subaequilongi, ramis hispidulis, bracteis lanceolato-ovatis
longiacuminatis grosse dentatis, dentibus longe setoso-mucronatis;
flores 7 mm. longi, calyce bilabiate glabro; vexillum glabrum;
legumen marginibus sparse ciliatis exceptis glabrum, uno latere fere
rectum, altero profunde constrictum, articulis vulgo 4, interdum 5,
semiorbicularibus, laxe et subprominente reticulato-venosis, stipite
calyce vix longi ore. Guatemala: Dept. Chiquimula, brushy rocky
slope, near the divide on the road from Zacapa to Chiquimula, alt.
660 meters, October, 1940, Paul C. Standley 73714 (type in Herb.
Field Mus.). Dept. Jutiapa, region of El Tablon, northeast of
Jutiapa, alt. 900 meters, damp thicket, October 31, 1940, Standley
75840.
In general appearance as well as in most details this is exactly
like A. americana L. but it differs in having uniformly much broader
pods, of somewhat distinctive appearance, and it is believed that a
good species, perhaps of very local distribution, is represented.
Bauhinia Herrerae (Britt. & Rose) Standl. & Steyerm., comb,
nov. Schnella Herrerae Britt. & Rose, N. Amer. Fl. 23: 206. 1930.
Canavalia munda Standl. & Steyerm., sp. nov. Frutex
scandens, caulibus glabris, petiolis foliolis brevioribus; foliola ob-
longo-elliptica vel oblongo-ovata 8-13 cm. longa 4-6 cm. lata acumi-
nata vel anguste longiacuminata, basi rotundata vel obtusa, longi-
petiolulata, glabra, subtus sparse albido-punctata; racemi nodosi
STUDIES OF CENTRAL AMERICAN PLANTS III 11
folia aequantes vel longiores; calyx campanulatus 10-12 mm. longus
subdense sericeus, labio superiore tubo multo breviore emarginato,
labio inferiore brevi, lobis 3 ovatis obtusis; corolla lilacina fere 3 cm.
longa, vexillo lato ad anthesin reflexo; legumen 13-16 cm. longum
3 cm. latum, dense pilis fusco-brunneis velutino-pilosum, breviter
stipitatum, valvis 3-costatis, costa media a sutura ventrali 7-8 mm.
remota; semina late ovalia compressa castanea lucida 12-15 mm.
longa 9-12 mm. lata, hilo fusco 5 mm. longo. Guatemala: Dept.
Quezaltenango: Finca Pirineos, low southern slopes of Volcan de
Santa Maria, between Santa Maria de Jesus and Calahuache",
alt. 1,300-1,500 meters, December, 1939, Julian A. Steyermark 33179
(type in Herb. Field Mus.). Several other collections seen from the
departments of Quezaltenango and San Marcos, western Guatemala.
Related to C. villosa Benth., in which the leaflets usually are
abundantly pubescent and not punctate.
Climacorachis guatemalensis Standl. & Steyerm., sp. nov.
Herba annua erecta usque 80 cm. alta simplex vel ramosa, interdum
suffrutescens, praeter inflorescentiam glabra; stipulae angustae 1 cm.
longae prope medium affixae; foliola ca. 20-juga 6-8 mm. longa 1.5
mm. lata oblique acuta 3-nervia sessilia; racemi pauciflori flexuosi
folia aequantes vel longiores, rhachi viscido-setulosa, pedicellis longis
gracilibus; bracteae ovatae setuloso-ciliatae; calyx 3 mm. longus
viscido-setulosus; legumen oblongum 1.5-2 cm. longum 5-7 mm.
latum parum compressum, apice oblique rotundatum vel truncatum,
densiuscule viscido-setulosum, pilis basi dilatatis et nigrescentibus;
semina globosa 2 mm. diam. opaca nigro-olivacea. Guatemala:
Dept. Huehuetenango: Pine-oak forest, Rio Pucal, about 14 km.
south of Huehuetenango, alt. about 1,800 meters, January, 1941,
Paul C. Standley 82330 (type in Herb. Field Mus.) ; also no. 82370
from the same locality.
Two other members of this genus are known, each from a single
collection made in Jalisco, Mexico. Both differ from the Guatemalan
plant in their narrower fruit which is more or less constricted along
the ventral margin.
Inga Skutchii Standl., sp. nov. Arbor 9-metralis, ramulis
novellis pilis mollibus brevibus fulvis adscendentibus sat dense
pilosis; folia inter minora ca. 12-13 cm. longa, rhachi dense pilis
subadpressis fulvis pilosa, inter paria foliolorum glandula parva
subglobosa elevata onusta; foliola parva 6-8-juga membranacea
lanceolato-oblonga vel anguste elliptico-oblonga, terminalia paullo
12 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
majora, infima valde reducta, media et terminalia 3.5-5.5 cm. longa
1.2-2 cm. lata, apicem versus breviter vel longius angustata, apice
ipso anguste obtuso et mucronato-apiculato, basi subinaequali obtusa
vel anguste rotundata, supra ad costam breviter pilosa, aliter glabra,
subtus paullo pallidiora sparse, ad costam densius, pilis parvis
rigidis subadpressis hirtello-pilosa, costa nervisque lateralibus pro-
minulis; flores albi spicati, spicis solitariis vel fasciculatis 10-11 cm.
longis ca. 3 cm. longe pedunculatis remote laxeque multifloris, rhachi
dense breviter adpresso-pilosa; calyx anguste tubulosus 5 mm.
longus sparse hirtellus, lobis ovatis acutiusculis tubo bene brevioribus;
corolla ca. 13 mm. longa dense strigosa, lobis ovatis acutis vix ultra
2 mm. longis; stamina ca. 3.5 cm. longa. Costa Rica: By a forest
stream, basin of El General, Prov. San Jose", alt. 675-900 meters,
March, 1940, Alexander F. Skutch 4823 (type in Herb. Field Mus.).
An exceptionally distinct species for the genus; easy of recogni-
tion among Costa Rican ones because of the combination of numerous
small leaflets and elongate, remotely flowered spikes.
Pachecoa Standl. & Steyerm., gen. nov. Frutices erecti ramosi
pilosi; stipulae lineari-subulatae striato-nervosae; folia imparipinnata,
foliolis paucis integris mucronatis exstipellatis herbaceis; flores flavi,
pedunculis axillaribus rigidis plerumque 2-floris; bracteae parvae
lanceolatae rigidae, bracteolis subulatis vel fere filiformibus denticu-
latis; calyx basi attenuatus et stipitiformis, prpfunde 5-lobus, lobis
lanceolato-attenuatis, 4 subaequalibus, infimo paullo longiore et
angustiore; vexillum suborbiculare extus pilosum unguiculatum, alis
oblique obovatis, carina incurva obtusa; stamina in tubum clausum
connata, antheris ovalibus uniformibus; ovarium sessile pauci-
ovulatum, stylo elongate, stigmate minuto terminali; legumen lineari-
oblongum subtetragonum, suturis rectis, articulis quadratis dense
breviter pilosis. Type species, Pachecoa guatemalensis Standl. &
Steyerm.
The genus is dedicated to Don Mariano Pacheco Herrarte,
Director General de Agricultura de Guatemala, who for many years
has labored with signal success to improve agricultural conditions
in this republic. His celebrated gardens, and particularly their
fine collection of orchids, are one of the most interesting sights to
be seen in Guatemala City. His exhibits of orchids and other
ornamental plants and of agricultural products of Guatemala supply
each November one of the most attractive and instructive sections
of the Feria Internacional that draws so many visitors from all
Central America as well as from Mexico and the United States.
STUDIES OF CENTRAL AMERICAN PLANTS III 13
This genus is obviously related to Chapmannia Torr. & Gray,
confined to Florida. Chapmannia is a herbaceous plant with similar
leaves but with a long, terminal, wand-like, almost naked inflores-
cence; the calyx is closed in bud, and has very short, broad teeth;
the legume is glandular-muricate and deeply constricted into ovoid
subterete joints. Pachecoa is a tall shrub with hard stems and small
axillary inflorescences; the calyx is open in bud and has long narrow
lobes; the legume is merely pilose, not at all constricted between
the seeds, the joints being quadrate and in cross section somewhat
tetragonous.
Pachecoa guatemalensis Standl. & Steyerm., sp. nov. Frutex
erectus 2-3-metralis, ramis striatis dense pilosis et sparse pilis viscidis
luteis patentibus setulosis; stipulae persistentes adpresso-pilosulae
5-8 mm. longae; folia parva breviter petiolata; foliola vulgo 3-5
alterna breviter petiolulata late elliptica vel ovalia 13-25 mm. longa
7-14 mm. lata, apice rotundata et mucronata, basi obtusa vel
rotundata, supra viridia pilosa et sparse setulosa, subtus dense pilis
albidis subadpressis laxis pilosa; pedunculi folia aequantes vel
breviores, pilosi et setulosi; calyx ca. 8 mm. longus pallide viridis
adpresso-pilosus et sparse supra medium setulosus, lobis tubum
fere aequantibus angustis attenuatis setoso-ciliatis; petala pallide
aurantiaca, vexillo ca. 15 mm. longo extus dense piloso; legumen
1.5 cm. longum apice rostro 4-5 mm. longo auctum, 3-4 mm. latum,
sessile, dense pilis brevibus non viscidis subpatentibus pilosum.
Guatemala: Dept. Jutiapa: Damp thicket near Jutiapa, alt. 850
meters, October, 1940, Paul C. Standley 75307 (type in Herb. Field
Museum); also no. 75032 from the same locality. Mexico: "Rio
Taba" (locality not certainly decipherable), 1841-43, Liebmann.
The plant is abundant in thickets at the northeastern edge of
Jutiapa, where it forms dense stands of considerable extent on some
of the hillsides.
Dichapetalum Brenesii Stand!., sp. nov. Frutex vel arbor
4-8-metralis, trunco usque 20 cm. diam., cortice laevi griseo, ramulis
brunnescentibus sat gracilibus densiuscule sericeis vel strigosis,
glabrescentibus, internodiis plerumque brevibus; stipulae parvae
cito deciduae; folia modica breviter petiolata in sicco fusca sub-
lucida crasse membranacea, petiolo crassiusculo vulgo 5-7 mm.
longo; lamina elliptico-oblonga vel lanceolato-oblonga media vel
paullo supra mediam latissima vulgo 7-15 cm. longa et 3.5-6 cm.
14 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
lata subabrupte longiacuminata, basin acutam versus angustata,
supra ad costam nervosque strigosa, aliter glabra vel glabrata, venulis
prominulis, subtus concolor ad nervos venasque strigosa vel fere
omnino glabra; cymae parvae vulgo dense multiflorae, petiolo fere
omnino cum pedunculo adnato, ramis dense sordide sericeis, floribus
breviter crasseque pedicellatis congestis; bracteae parvae lineares
extus tomentellae; sepala 2-2.5 mm. longa extus dense albido-tomen-
tosa; drupa clavata vel anguste obovato-oblonga 2.5-3.5 cm. longa
dense sordide adpresso-tomentulosa vix compressa. Costa Rica:
Zarcero, alt. 1,950 meters, deep forest, April, 1937, Austin Smith
4120 (type in Herb. Field Mus.). La Palma de San Ramon, March,
1929, A. M. Brenes 6766. La Ventolera, southern slope of Volcan
de Poas, 1,700 meters, wooded quebrada, Standley 34723, 34674.
Zarcero, Austin Smith 4104, 366519. Las Lajas de Zarcero, 1,575
meters, Smith 746. La Brisa de Zarcero, 2,200 meters, Smith H971.
Panama: Progreso, Chiriqui, Cooper & Slater 172.
Heretofore the Costa Rican tree has been identified with D.
Donnell-Smithii Engler, of Guatemala. It is, however, more closely
related to D. chiapasense Standl., which ranges from Chiapas to
British Honduras and Honduras, but is sufficiently distinct in its
acute-based leaves and larger, narrower, minutely tomentulose fruit.
D. Donnell-Smithii, confined so far as now known to Guatemala,
has much larger and relatively broader, thin, densely pilose leaves.
D. Brenesii is called Blancito in Panama. The corolla is described
as white to cream-colored or pale yellow.
SAPINDACEAE
Serjania lobulata Standl. & Steyerm., sp. nov. Frutex alte
scandens, ramis subteretibus, novellis minute cano-puberulis; corpus
lignosum simplex teres; folia pinnatim 5-foliolata, petiolo ut rhachi
nudo puberulo; foliola inferiora longiuscule petiolulata, superiora
sessilia, rhombea vel rhombeo-ovata, interdum oblongo-elliptica,
4-10 cm. longa 2-6 cm. lata, acuta vel obtusa, basi truncata usque
acuminata vel cuneatim attenuata, grosse remoteque crenulata vel
fere lobata, supra sat dense puberula, subtus pallidiora ubique
dense breviter pilosula; thyrsi foliis breviores dense ochraceo-
tomentulosi, cincinnis longipedunculatis paucifloris, bracteis minutis,
pedicellis brevibus; sepala dense cano-tomentulosa 3.5-4 mm. longa
suborbicularia apice rotundata; petala alba 5 mm. longa glabra
intus glandulis parvis conspersa; fructus late cordatus ca. 2 cm.
longus atque 2.5 cm. latus superne parum angustatus, loculis duris
STUDIES OF CENTRAL AMERICAN PLANTS III 15
subglobosis dense hirtellis rugoso-venosis, dissepimentis latis, alis
tenuibus dense puberulis infra loculos non constrictis. Guatemala:
Dept. Chiquimula, near the divide on the road from Zacapa to
Chiquimula, brushy rocky slope, alt. about 660 meters, October,
1940, Paul C. Standley 73715 (type in Herb. Field Mus.). Dept.
Zacapa, rocky slopes near Santa Rosalia, 250-300 meters, October,
1939, Steyermark 29019; vernacular name Chilmecate.
Related to S. Schiedeana Schlecht. of Mexico, which has a different
stem structure, much denser and more abundant pubescence, smaller
flowers, and larger fruits.
Serjania macrocarpa Standl. & Steyerm., sp. nov. Frutex
scandens, ramis obtuse trigonis dense pilis subpatentibus brunneo-
pilosis vel subtomentosis; corpus lignosum e centrali magno subtereti
et periphericis 3 parvis com posi turn; folia biternata, petiolo nudo,
rhachi anguste marginata; foliola lanceolato-oblonga usque obovato-
elliptica vel oblongo-elliptica 5-13 cm. longa 2.5-5.5 cm. lata acumi-
nata vel abrupte acuminata, basi attenuata ad acuta, sessilia, sub-
grosse remoteque crenato-serrata, supra sublucida tantum ad nervos
puberula, subtus pallidiora sat dense velutino-pilosa; flores non visi;
fructus ambitu late cordatus ca. 4 cm. longus et 4.5 cm. latus ubique
dense molliter pilosulus vel ad loculos tomentosus, loculis triangulari-
bus dorso acutissimis et subalatis, dissepimentis latissimis, alis
tenuibus infra loculos non angustatis conspicue venosis. Guate-
mala: Los Amates, Dept. Izabal, February 23, 1908, W. A. Keller-
man 7532 (type in Herb. Field Mus.). British Honduras: Vaca,
El Cayo District, on hill slope, March, 1938, P. H. Gentle 2291.
The fruits are distinctive and not similar to those of any Mexican
or Central American species of which we have seen material. They
are unusually large and remarkable for the large hard seed-bearing
portion, which is triangular in cross section and almost winged
dorsally, the partition walls being relatively broader than in most
species of the genus.
Pseudabutilon inornatum Standl. & Steyerm., sp. nov.
Frutex 1.5-3 m. altus ramosus, ramis pallidis dense pilis stellatis
valde inaequilongis tomentosis; folia longipetiolata membranacea,
petiolo 2.5-11 cm. longo; lamina late ovata vel rotundo-ovata
6-17 cm. longa 5-12 cm. lata acuminata vel longiacuminata, basi
breviter cordata, crenato-dentata, inferioribus interdum angulatis
vel breviter trilobis, supra viridis sparse minute stellato-puberula,
16 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
subtus paullo pallidior sparse vel subdense stellato-hirtella; flores
in racemos elongates interruptos paniculatos dispositi, plerumque
aggregati, breviter pedunculati, pedunculis calyce vulgo brevioribus
stellato-pilosis; calyx in statu fructifero campanulatus carpidia
arete amplectens 5 mm. longus dense stellato-hispidulus, breviter
5-lobatus, lobis latissime ovatis acutis; fructus 6-8-carpidiatus ca.
5 mm. altus et 6 mm. latus, carpidiis apice late rotundatis dorso
densissime stellato-hirtellis; semina uniseriata; carpidia dissepimento
a dorso oriente fere perfecte 2-locularia. Guatemala: Near Ama-
titlan, Dept. Guatemala, alt. 1,170 meters, December, 1938, Paul C.
Standley 61425 (type in Herb. Field Mus.) ; also no. 61400 from the
same locality, growing in dry forest; John R. Johnston 1714.
Mexico: Rio de Santa Maria, near Zacuapan, Veracruz, rocky soil,
November, 1906, C. A. Purpus 2239.
Evidently referable to the subgenus Abutilastrum, and related
to P. paniculatum (Rose) R. E. Fries of western Mexico. In that
the fruit consists of 10-11 carpels which are acute or sharply angulate
at the apex, while in P. inornatum they are broadly rounded, without
any indication of an angle, and fewer.
GARRYACEAE
Garrya corvorum Standl. & Steyerm., sp. nov. Frutex 2-
metralis, ramulis crassis lucidis, novellis dense pilis albidis subad-
pressis indutis; folia parva rigide coriacea breviter petiolata, petiolo
3-5 mm. longo piloso; lamina oblonga vel elliptico-oblonga 2.5-4.5
cm. longa 1-1.5 cm. lata, apice obtusa vel subacuta et apiculata,
basi obtusa vel anguste rotundata, supra lucida fere glabra, costa
nervisque manifeste impressis, subtus opaca sat dense pilis albidis
subadpressis pilosa, costa prominente, nervis lateralibus parum
prominentibus inconspicuis, venis obsoletis; racemi masculi 3-3.5 cm.
longi breviter pedunculati simplices terminales solitarii ubique dense
pilis longis albidis pilosi; bracteae latissime ovatae usque 8 mm.
longae alte connatae, cuspidato-acutae. Guatemala: Dept. Huehue-
tenango, dense rocky (limestone) Juniperus forest, along the road
in the region of Che'mal, Sierra de los Cuchumatanes, at km. 36,
altitude about 3,300 meters, December, 1940, Paul C. Standley
81651 (type in Herb. Field Mus.).
Only one shrub of the species was found. The only other species
known from Central America, Garrya laurifolia Hartweg, differs in
having much narrower bracts in the racemes, and much larger leaves
in which the venation is not impressed or scarcely so.
STUDIES OF CENTRAL AMERICAN PLANTS III 17
CLETHRACEAE
Clethra bimatris Standl. & Steyerm., sp. nov. Arbor 12-
metralis, trunco 30 cm. diam., ramulis densissime pilis longis laxis
sordide brunneis tomentosis; folia magna petiolata membranacea,
petiolo crasso 1.5-2.5 cm. longo dense piloso; lamina oblongo-obovata
usque anguste oblanceolato-oblonga 12-21 cm. longa 3-8 cm. lata,
supra medium latissima, apicem versus plus minusve angustata,
apice ipso obtuso usque acuto, basin versus late cuneato-angustata,
basi ipsa obtusa vel anguste rotundata, margine basali utroque
anguste recurvo, integra vel apicem versus remote serrata, supra
viridis in sicco fusca primo sparse hirsuta glabrescens, sublucida,
subtus glaucescens atque sordido-brunnescens, ubique tomento laxo
non adpresso induta, costa nervisque brunneis atque tomentosis,
nervis lateralibus utroque latere ca. 17 valde obliquis et adscenden-
tibus fere rectis vel leviter curvis prominentibus, venis teneris
manifestis; racemi numerosi elongati laxi paniculam amplam usque
28 cm. longam efformantes, rhachi pedicellisque dense brunneo-
stellato-tomentosis, pedicellis rectangule divaricatis 4-5 mm. longis
gracilibus; calyx extus dense brunnescenti-tomentosus ca. 3 mm.
longus, lobis ovalibus obtusissimis; petala calyce paullo longiora
glabra, apice eroso-dentata et ciliata. Guatemala: Entre Rios,
Dept. Izabal, July, 1927, Henry Kuylen 151 (Yale School of
Forestry No. 10744; type in Herb. Field Mus.). British Honduras:
Temash River, in secondary forest, altitude 30 meters, August,
1935, William A. Schipp 1343.
This has been reported from Guatemala as well as from British
Honduras under the name Clethra lanata Mart. & Gal., a Mexican
species, not definitely known to occur in Guatemala or British
Honduras. In that the flowers are conspicuously larger and the
leaves relatively broader and generally rounded or broadly rounded
at the apex.
Clethra licanioides Standl. & Steyerm., sp. nov. Arbor 9-
metralis, ramulis crassis dense brunneo- tomentosis; folia modica
breviter petiolata rigide coriacea, petiolo crasso 1-1.5 cm. longo
dense brunneo-tomentoso; lamina rotundo-obovata 8-10 cm. longa
5-7 cm. lata, apice latissime rotundata vel truncata et minute
emarginata, basin versus paullo angustata, basi ipsa anguste rotun-
data vel obtusa, supra sublucida in sicco sublutescens ubique pilis
minutis vix perspicuis depresso-stellatis conspersa, costa nervisque
manifestis sed non elevatis, subtus viridis ubique pilis majusculis
sat dense velutine stellato-pilosa, nervis lateralibus utroque latere
18 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
ca. 13 fere rectis angulo semirecto adscendentibus, venis prominulis;
racemi usque 25 cm. longi laxiflori numerosi paniculam magnam
latamque efformantes, rhachi dense brunneo-tomentosa, pedicellis ca.
5 mm. longis divaricatis vel subadscendentibus, floribus saepius
nutantibus; calyx 4-5 mm. longus dense stellate- tomentellus, seg-
mentis late ovatis vel ovalibus obtusissimis; petala glabra magna
sepalis paullo longiora apice laciniata et erosa. Guatemala: Dept.
Jalapa, Potrero Carrillo, 13 miles northeast of Jalapa, altitude 1,500-
1,700 meters, December 12, 1939, Julian A. Steyermark 33124 (type
in Herb. Field Mus.).
Local name, Zapotillo. The flowers are fragrant, the petals white,
the sepals buff with a slight tinge of pink. The relationship of the
Guatemalan tree is with C. lanata Mart. & Gal., which has smaller
flowers and relatively narrower leaves.
SOLANACEAE
Athenaea physocalycia (Donn. Smith) Standl. & Steyerm.,
comb. nov. Brachistus physocalycius Donn. Smith, Bot. Gaz. 90:
8. 1905.
Athenaea viscida (Standl.) Standl. & Steyerm., comb. nov.
Capsicum viscidum Standl. Carnegie Inst. Wash. Publ. 461: 84. 1935.
Lycianthes cuspidata (Morton) Standl. & Steyerm., comb,
nov. Solatium cuspidatum Morton in Lundell, Contr. Univ. Mich.
Herb. 4: 25. 1940.
Lycianthes hondurensis (Morton) Standl. & Steyerm., comb,
nov. Solanum hondurense Morton in Lundell, Contr. Univ. Mich.
Herb. 4: 26. 1940.
Physalis campanula Standl. & Steyerm., sp. nov. Herba ut
videtur perennis laxe ramosa procumbens, caulibus pilis longis viscid is
albidis paten tibus dense villosis; folia longipetiolata, petiolo 1-4 cm.
longo viscido-villoso; lamina late ovata 3-7.5 cm. longa 2-5 cm. lata,
subacuta usque longiacuminata, basi late rotundata, integra vel
undulata, interdum remote sinuato-dentata, supra viridis sparse
breviter villosa, subtus concolor sparse breviterque villosa; pedicelli
floriferi gracillimi 1.5-2 cm. longi dense viscido-villosi; calyx ad
anthesin 1 cm. longus campanulatus dense pilis paten tibus viscido-
villosus, basi subtruncatus, dentibus tubo lato paullo brevioribus
triangularibus longe anguste acuminatis; corolla tubuloso-campanu-
lata 18 mm. longa apice 8 mm. lata, extus breviter viscido-villosula,
intus glabra, limbo remote obtuso-dentato; stamina prope basin
STUDIES OF CENTRAL AMERICAN PLANTS III 19
corollae inserta, filamentis filiformibus paullo inaequalibus, antheris
paullo ultra 3 mm. longis; calyx fructifer late ovoideus immaturus
2 cm. longus et 16 mm. latus, ut videtur teres et vix angulatus dense
viscido-villosus, basi non depressus, apice abrupte contractus, lobis
ca. 5 mm. longis fere subulatis. Guatemala: Dry banks along
upper part of Quebrada Canjula, between Sibinal and Canjula,
Volcan de Tacana, Dept. San Marcos, alt. 2,200-2,500 meters,
February, 1940, Julian A. Steyermark 36067 (type in Herb. Field
Mus.).
The species is noteworthy for its narrowly campanulate corolla,
slightly contracted at the apex and altogether unlike the corolla
usual in this genus. The fruit, however, i.e. the fruiting calyx, is
quite typical for the genus, and there is no good reason for attempting
to refer the plant to some other group, which could be done on the
basis of corolla form alone.
Physalis carnosa Standl. & Steyerm., sp. nov. Annua prostrata
dense ramosa fere glabra, caulibus crassis ca. 15 cm. longis carnosis
subgeniculatis, novellis scaberulis; folia parva carnosa, petiolis crassis
usque ad 15 mm. longis; lamina late ovata usque ovato-oblonga
plerumque 1-2 cm. longa et 5-12 mm. lata, obtusa vel subacuta,
basi cuneata vel obtusa, saepe valde inaequilateralis, primo minute
scaberula, cito glabrata, nervis lateralibus utroque latere ca. 3 crassis
subtus prominentibus; pedicelli floriferi 3-4 mm. longi minute
puberuli, fructiferi vix longiores; calyx florifer 2.5 mm. longus
obscure puberulus, dentibus anguste triangularibus acuminatis tubo
paullo brevioribus; corolla ut videtur campanulata vix ultra 3 mm.
longa; calyx fructifer ca. 12 mm. longus et 8 mm. latus late ovoideus
basi non depressus glabratus arete reticulato-venosus, obtuse
pentagonus, dentibus brevibus conniventibus triangularibus acutis.
Guatemala: On sand dunes along the seashore, Ocos, Dept. San
Marcos, March, 1940, Julian A. Steyermark 37766 (type in Herb.
Field Mus.).
In technical characters, particularly its small flowers and fruiting
calyces, this evidently is close to P. Lagascae Roem. & Schult. and
P. micrantha Link. Those, however, are normally erect or ascend-
ing plants with delicate and very thin leaves. The leaves of P.
carnosa as pressed are exceedingly irregular in outline, so much so
that it is difficult to ascribe to them any normal leaf shape.
Physalis lassa Standl. & Steyerm., sp. nov. Ut videtur vulgo
perennis sed interdum annua, erecta vel procumbens vel decumbens,
20 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
ramosa, caulibus dense pilis brevibus vel longiusculis eglandulosis
patentibus pilosa vel subvillosa; folia petiolata, petiolo usque 2 cm.
longo dense villosulo; lamina late ovata vel ovata 3-6 cm. longa
2-4.5 cm. lata acuta vel interdum obtusa, basi late rotundata vel
subcordata, supra dense pilis brevibus mollibus villosula, subtus
pilis paullo longioribus pilosa vel interdum fere puberula; pedicelli
floriferi ca. 5 mm. longi dense villosi, fructiferi usque ad 13 mm. longi;
calyx florifer ca. 9 mm. longus dense albido-villosus, lobis tubo
paullo longioribus anguste triangularibus acutis vel acuminatis;
corolla late campanulata ca. 14 mm. longa, extus sparse villosula
vel fere glabra; calyx fructifer 3-3.5 cm. longus 2 cm. latus dense
villosulus vel breviter pilosus, basi vix depressus, obtuse pentagonus,
dentibus ca. 8 mm. longis triangularibus acutis vel acuminatis; bacca
fere 1 cm. diam. Guatemala: Dept. Jalapa, grassy thickets, between
Jalapa and Montana Miramundo, alt. 1,500-2,000 meters, December,
1939, Julian A. Steyermark 32868 (type in Herb. Field Mus.).
Dept. Guatemala, Ignacio Aguilar 213. Sierra de las Minas, near
San Geronimo, Dept. Baja Verapaz, 1,140 meters, W. A. Kellerman
6656. Dept. Zacapa, grassy places, lower slopes of Sierra de las
Minas, along trail above Rio Hondo, 250-900 meters, Steyermark
29545.
This species bears some relationship to the Mexican one that has
been referred to P. peruviana L. That South American species,
however, does not occur in Mexico or Central America, so far as
the available specimens show. P. peruviana differs from the Guate-
malan plant here described in its much broader and conspicuously
cordate leaves and in its larger and relatively broader fruiting
calyx.
Solatium decurtatum Standl. & Steyerm., sp. nov. Frutex ut
videtur omnino inermis, ramis sat gracilibus, vetustioribus ochraceis,
novellis dense pilis stellatis ochraceis sessilibus vel breviter stipitatis
aequaliter ramosis tomentosis; folia solitaria integra vel vix undulata
membranacea, petiolo 1-2 cm. longo dense stellate- tomentoso;
lamina elliptico-oblonga vel ovali-oblonga prope mediam latissima
5-8 cm. longa 2-3.5 cm. lata apice rotundata vel obtusa, interdum
breviter acutata, basi plus minusve inaequali obtusa vel subacuta
abrupte in petiolum contracta, supra viridis sat dense pilis parvis
aequalibus stellatis tactu asperulis induta, subtus grisea dense
molliterque pilis intertextis stellato-tomentosa, nervis lateralibus
utroque latere ca. 7; inflorescentia terminalis vel pseudolateralis
cymosa, interdum densa atque umbelliformis 3 cm. tantum lata,
STUDIES OF CENTRAL AMERICAN PLANTS III 21
saepius dichotomo-ramosa et usque 6 cm. lata, multiflora, ramis
plus minusve elongatis et secundifloris dense griseo-stellato-tomen-
tosis, pedicellis fructiferis usque 8 mm. longis; calyx fere 3 mm.
longus densissime stellato-tomentosus, dentibus triangularibus sub-
acutis brevibus; corolla ut videtur alba extus dense stellato-pilosula
ca. 7 mm. longa stellata, lobis angustis sensim attenuatis intus glabris;
stamina distincta glabra, antheris fere 5 mm. longis oblongo-lineari-
bus sursum vix angustatis; bacca glabra globosa lucida 5-6 mm.
diam. Guatemala: Lago de Yaxha, Pete"n, March, 1933, C. L.
Lundell 2012 (type in Herb. Field Mus.). British Honduras:
Corozal District, 1931-32, P. H. Gentle 121, 253.
A relative of S. racemosum Jacq. of the West Indies, which has
usually much longer, mostly acute or acuminate leaves and anthers
7-9 mm. long.
RUBIACEAE
Borreria vegeta Standl. & Steyerm., sp. nov. Herba annua
erecta 30-60 cm. alta et ultra simplex vel pauciramosa, caulibus
crassis subteretibus sparse laxeque albido-pilosulis vel glabratis;
stipularum vagina laxa 5 mm. longa, setis paucis erectis aequilongis
onusta laxe pilosa; folia herbacea, interdum usque 1 cm. longe
petiolata, petiolo lato marginato; lamina lanceolato-oblonga usque
ovata vel oblongo-elliptica 3.5-7.5 cm. longa 1.2-2.8 cm. lata atten-
uato-acuminata, basi cuneatim contracta, supra sparse scabra vel
glabrata tactu aspera, costa nervisque subimpressis, subtus pallidior
ad costam nervosque scabra vel hispidula, nervis lateralibus 6-7
valde obliquis prominentibus; capitula plerumque terminalia, in-
terdum quoque ex axillis superioribus nascentia 2-2.5 cm. lata dense
multiflora, floribus breviter pedicellatis; bracteae capitulorum
terminalium foliis similes et aequimagnae, vulgo 4; hypanthium
oblongum saltern superne villosulum ; sepala 4 ovalia usque suborbic-
ularia vel rotundato-obovata viridia apice rotundata dense pectinato-
ciliata glabra vel glabrata; corolla alba ca. 2.5 mm. longa extus
glabra, lobis ovatis acutiusculis; antherae bene exsertae; capsula
ovalis vel late oblonga 3 mm. longa superne villosula vel glabra
membranacea calyce persistente coronata; semina teretia 2 mm.
longa fusco-ferruginea profunde transverso-rugosa. Guatemala:
Dept. Jutiapa, between Jutiapa and La Burrera, northeast of Jutiapa,
alt. 800 meters, November, 1940, in damp thicket, Standley 76007
(type in Herb. Field Mus.) . Vicinity of Jutiapa, 850 meters, Standley
75041. Near El Molino (Dept. Santa Rosa), Dept. Jutiapa, 600
meters, wet thicket, Standley 78447. Dept. Santa Rosa, along the
22 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
Avellana road south of Guazacapan, 150 meters, weedy field,
Standley 79475.
Because of the transversely rugose seeds as well as habit and
general appearance, it is obvious that this plant is related to B. laevis
(Lam.) Griseb. That, however, is a less stout plant, usually decum-
bent or procumbent, with conspicuously smaller flower heads and
narrow, usually much smaller sepals.
Bouvardia Pachecoana Standl. & Steyerm., sp. nov. Frutex
metralis glaber; vagina stipularis ca. 4 mm. alta setis paucis 2-2.5
mm. longis aucta; folia opposita membranacea, petiolo 4-8 mm.
longo; lamina ovata vel oblongo-ovata 5-10 cm. longa 2.2-4 cm.
lata acuta vel acuminata, basi obtusa vel acuta et interdum breviter
decurrens, utrinque in statu sicco pallida, nervis lateralibus utroque
latere vulgo 3 prominulis, margine adpresso-ciliato; inflorescentia
terminalis cymoso-corymbosa dense multiflora, bracteis pro parte
foliaceis, pedicellis usque 6 mm. longis; hypanthium obovoideum
1.5 mm. longum basi obtusum; sepala ca. 3 mm. longa erecta tri-
angulari-linearia attenuata viridia; corolla salmoneo-coccinea, tubo
16-18 mm. longo extus glabro sursum sensim dilatato ad faucem
usque 3 mm. lato, lobis suberectis ovatis vel oblongo-ovatis ca.
5 mm. longis obtusis; stylus et stamina inclusa. Guatemala: Dept.
San Marcos, in a wet, densely forested quebrada, Barranco Emi-
nencia, along the road between San Marcos and San Rafael Pie
de la Cuesta, in upper part of the barranco between Finca La Lucha
and Buena Vista, alt. 2,500-2,700 meters, February, 1941, Paul C.
Standley 86226 (type in Herb. Field Mus.).
A well-marked species, not very closely related to any other of
Guatemala or southern Mexico. From B. dictyoneura Standl. it
differs in its obviously petiolate leaves, acute at the base or contracted
and decurrent, and not reticulate- veined. The plant is a particularly
handsome one because of its abundance of bright salmon-red flowers.
It is named for Don Mariano Pacheco Herrarte, Director General
de Agricultura of Guatemala, to whom we are indebted for many
favors.
Crusea setosa (Mart. & Gal.) Standl. & Steyerm., comb. nov.
Borreria setosa Mart. & Gal. Bull. Acad. Brux. 11, pt. 1: 131. 1844.
Spermacoce setosa Hemsl. Biol. Centr. Amer. Bot. 2: 60. 1881.
Crusea cruciata Wats. Proc. Amer. Acad. 25: 152. 1890. B. chiapensis
Brandeg. Univ. Calif. Publ. Bot. 10: 417. 1924. While no specimen
of Galeotti 2627, type of Borreria setosa, has been seen, there is in
the Herbarium of Field Museum an excellent photograph of a
STUDIES OF CENTRAL AMERICAN PLANTS III 23
specimen of this number from the Paris herbarium. This, in associa-
tion with the original description, shows clearly that the species
represented is the same as the one that has passed as Crusea cruciata
Wats., a plant of wide distribution in Mexico and Central America.
Hillia macrocarpa Standl. & Steyerm., sp. nov. Frutex epi-
phyticus omnino glaber, ramis crassis ochraceis; folia in sicco coriacea,
petiolo crasso ca. 4-7 mm. longo; lamina elliptico-oblonga vel
oblanceolato-oblonga 6-8.5 cm. longa 2-3.5 cm. lata, acuta vel
subacuminata, basi cuneato-angustata, prope vel paullo supra
mediam latissima, supra plana, nervis subtus obscuris planis, laterali-
bus paucis angulo angustissimo adscendentibus; capsula breviter
crasse pedicellata anguste cylindracea 8-9 cm. longa, medio fere
1 cm. crassa, apice breviter angustata, basin versus longe sensimque
angustata. Guatemala: Dept. Quezaltenango, lower south-facing
slopes of Volcan de Santa Maria, between Santa Maria de Jesus
and Calahuache", along the great barranco between Finca Pirineos
and Patzulin, alt. 1,300-1,500 meters, January, 1940, Julian A.
Steyermark 33667 (type in Herb. Field Mus.).
Local name Jazmin. Leaves fleshy, coriaceous, rich grass-green
above, paler beneath, with whitish green at base in the center;
capsule pale green. Growing on a ridge in a cafetal. The relation-
ship is with Hillia loranthoides Standl. of Costa Rica, which has
capsules less than half as long.
Psychotria Aguilarii Standl. & Steyerm., sp. nov. Frutex
1-1.5 m. altus ramosus, ramis gracilibus, novellis glabris; stipulae
caducae ca. 1 cm. longae membranaceae ferrugineae glabrae, primo
folia calyptratim involventes, serius fissae; folia membranacea,
petiolo 5-12 mm. longo glabro fere ad basin marginato; lamina
oblanceolata vel oblongo-oblanceolata 8-12 cm. longa 2-3.5 cm.
lata longe angusteque attenuato-acuminata, basin versus longe
anguste attenuata et longe decurrens, supra viridis glabra, nervis
manifestis sed non elevatis, subtus pallidior glabra sed in axillis
domatiatis nervorum sparse breviter barbata, nervis lateralibus
pallidis utroque latere ca. 11 angulo semirecto vel paullo latiore
adscendentibus, ad costam interdum sparse hirsuta; inflorescentia
terminalis sessilis laxa pauciflora usque 4.5 cm. longa e basi pauci-
radiata, ramis glabris, bracteis parvis caducis, floribus ut videtur
fere omnibus sessilibus dense ad apices ramulorum aggregatis;
calyx cum hypanthio vix ultra 1 mm. longus glaber truncatus et
remote obscure dentatus hypanthio obovoideo-conico paullo brevior;
corolla alba extus glabra infundibuliformis 5 mm. longa fauce non
24 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
barbata, lobis 1.5 mm. longis obtusis ovato-ovalibus suberectis
intus glabris; fructus intense ruber globosus 6 mm. diam., pyrenis
dorso grosse granulatis ecostatis. Guatemala: Dept. Chimaltenango:
Damp wooded barranco along the road between Chimaltenango
and San Martin Jilotepeque, alt. 1,500-1,700 meters, December,
1940, Paul C. Standley 80900 (type in Herb. Field Mus.). Dept.
Sacatepe"quez: Near Barranco Hondo, southeast of Alotenango,
1,000-1,250 meters, Standley 65024. Dept. Guatemala: Without
locality, in 1939, Jose Ignacio Aguilar 215. Near Finca La Aurora,
1,500 meters, Aguilar 126.
The species is related to P. nervosa Swartz, to which it bears much
superficial resemblance, but in that the fruits are conspicuously
costate and longer than thick. In that species, too, there is usually
ample pubescence on some parts of the plant.
Psychotria erecta (Aubl.) Standl. & Steyerm., comb, nov.
Ronabea latifolia Aubl. PI. Guian. 134. pi. 59. 1775, not Psychotria
latifolia Humb. & Bonpl. 1819. R. erecta Aubl. op. cit. 156. 1775.
P. axillaris Willd. Sp. PI. 1: 962. 1798. The genus Ronabea is
maintained as distinct by Bremekamp in the Flora of Surinam, a
treatment that has much justification, but on the whole it seems
more practical to leave the single species in Psychotria where most
authors have been content to place it.
Psychotria Schippii Standl. & Steyerm., sp. nov. Subgenus
Mapouria. Frutex vel arbor saepe 5-9-metralis, trunco interdum
5-10 cm. diam., ramulis subteretibus glabris viridescentibus vel
brunnescentibus interdum lenticellis magnis elongatis conspersis;
stipulae caducae usque 1.5 cm. longae attenuatae glabrae firmae;
folia majuscula breviter petiolata crasse membranacea vel sub-
coriacea, petiolo crasso 1-2.5 cm. longo, saepe fere ad basin margi-
nato; lamina elliptica vel oblongo-elliptica 11-19 cm. longa 5-9.5
cm. lata abrupte acuminata vel caudato-acuminata, basi acuta
vel subito contracta et longe decurrens, supra in sicco lucida glabra,
subtus paullo pallidior glabra sed in axillis nervorum saepius breviter
sparseque barbata, nervis lateralibus utroque latere ca. 14 promi-
nentibus angulo lato adscendentibus arcuatis; inflorescentia termi-
nalis sessilis cymoso-paniculata copiose ramosa in statu fructifero
plus minusve laxa multiflora, ca. 6 cm. longa et 8 cm. lata, ramis
basalibus 3-4 crassiusculis ferrugineis glabris, bracteis caducis,
floribus sessilibus; fructus ruber globosus glaber 4-5 mm. diam. basi
et apice rotundatus, pyrenis dorso obtuse costatis; calyx ad apicem
fructus persistens breviter dentatus, dentibus obtusis vel rotundatis
STUDIES OF CENTRAL AMERICAN PLANTS III 25
minutis. British Honduras: In advanced forest, limestone valley,
Valentin, El Cayo District. June-July, 1936, C. L. Lundell 6193
(type in Herb. Field Mus.). Columbia-Toledo, in 1927, G. H.
Donald 2. In forest, Guatemalan boundary, 630 meters, occasional,
Schipp S640. Guatemala: Uaxactun, April, 1931, Bartlett 12534.
In most of its characters Psychotria Schippii approaches the
common and widespread P. nervosa Swartz (P. undata Jacq.),
but in that the leaves have much fewer nerves and the fruits are
oval or ellipsoid, and evidently longer than broad.
Randia habrophlebia Standl. & Steyerm., sp. nov. Frutex
2-3-metralis, ramis crassis, lateralibus brevibus crassis late divergen-
tibus, foliis ad apices ramulorum dense aggregatis; stipulae brunneae
glumaceae extus strigosae, intus dense sericeae; folia breviter
petiolata herbacea, petiolo crasso 6-15 mm. longo densissime pilis
longis ochraceis subadpressis piloso; lamina suborbicularis usque
rotundato-ovata vel late elliptica 6.5-9.5 cm. longa 5.5-7.5 cm.
lata, apice late rotundata usque obtusa, basi late rotundata vel
obtusa, supra ubique densissime velutino-pilosa, ob nervos venasque
plus minusve impressas bullata, subtus grisea vel ochracea ubique
dense molliter tomentosa, nervis lateralibus utroque latere ca. 9
angulo acutissimo adscendentibus, venulis nervisque omnibus in-
signiter prominentibus et reticulatis; bacca terminalis solitaria
sessilis globosa 3.5 cm. diam., fere laevis, ecostata, sat dense pilis
subadpressis pilosa; semina compressa numerosissima. Guatemala:
Dept. Jutiapa, in thicket along river, vicinity of Jutiapa, alt. 850
meters, October-November, 1940, Paul C. Standley 76321 (type
in Herb. Field Mus.).
Closely related to R. monantha Benth., of the same region, but
in that the leaves at maturity are hirsute or pilose with straight,
not at all matted hairs; the leaves of the type specimen of R. habro-
phlebia are very old, yet still densely tomentose beneath with matted
hairs. The venation, too, is unlike that shown in any specimens
of R. monantha. It may be expected confidently that flowering
specimens of R. habrophlebia will afford other characters for separat-
ing the two species. The type specimen is almost destitute of spines,
but it is assumed that normally the spines are in 4's at the ends of
the branches, as in related species of the subgenus Basanacantha.
Rondeletia brachistantha Standl. & Steyerm., sp. nov.
Arbor, ramulis gracilibus sed rigidis teretibus, novellis pilis adpressis
fulvidis sparse pilosis; stipulae ca. 14 mm. longae reflexae brunneae
oblongo-deltoideae obtusae, extus strigosae; folia subcoriacea, petiolo
26 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. 23
usque 4 mm. longo; lamina ovata vel ovato-oblonga vel elliptico-
oblonga 5-8 cm. longa 2.5-3.5 cm. lata acuta vel subacuminata, apice
ipso obtuso, basi rotundata vel anguste rotundata, glabra vel subtus
secus nervos sparse hirtella, subtus paullo pallidior, nervis lateralibus
utroque latere ca. 5 prominentibus pallidis angulo semirecto adscen-
dentibus; inflorescentia terminalis cymoso-corymbosa dense multi-
flora 2-5 cm. longe pedunculata 3-6 cm. longa 4-9.5 cm. lata, e basi
triradiata, ramis dense fulvido-strigosis, bracteis minutis, floribus
sessilibus vel subsessilibus; hypanthium globosum 1.5 mm. altum
dense minute adpresso-hispidulum et primo paullo tomentosum,
calyce 0.7 mm. alto, dentibus minutis remotis obtusis; corolla rosea
extus ubique dense minuteque strigillosa, tubo 4-5 mm. longo
sursum paullo dilatato fauce dense luteo-barbato, lobis late rotundatis
vix ultra 1.5 mm. longis patulis intus puberulis; stylus glaber exsertus,
antheris inclusis; capsula globosa 4^4.5 mm. lata basi rotundata
glabrata loculicide dehiscens; semina 0.7 mm. lata subcompressa
fusco-brunnea puncticulata. Guatemala: Dept. San Marcos, dry
upper slopes, Rio Vega, near San Rafael and Guatemala-Mexico
boundary, Volcan de Tacana, alt. 2,500-3,000 meters, February,
1940, Julian A. Steyermark 36286 (type in Herb. Field Mus.).
Differing from R. cordata Benth. chiefly in the very small corollas,
those of that species being almost twice as large. Further material
will be necessary to decide whether the plant here described can be
maintained as a distinct species.
Rondeletia Cordovana Standl. & Steyerm., sp. nov. Arbuscula
fere glabra, ramulis subteretibus, novellis sparse minute strigillosis;
stipulae e basi triangulari subulatae ca. 3 mm. longae erectae; folia
parva membranacea, petiolo gracili 5-10 mm. longo sparse minuteque
strigilloso vel fere glabro; lamina lanceolato-oblonga 5.5-7.5 cm.
longa 1.7-2.5 cm. lata longissime attenuato-acuminata, basi acuta,
supra ad costam prominentem sparse minute pilosula vel omnino
glabra, nervis non elevatis, subtus pallidior, in axillis nervorum
barbata, aliter glabra, nervis lateralibus utroque latere ca. 6 arcuatis
angulo lato adscendentibus; inflorescentia terminalis 1.5 cm. longe
pedunculata erecta cymoso-corymbosa pauciflora ca. 3 cm. alta et
4.5 cm. lata basi trichotoma, floribus ad apices ramorum primariorum
dense cymoso-aggregatis sessilibus vel usque 3 mm. longe pedicellatis,
bracteis inferioribus foliaceis foliis conformibus sed parvis, usque
2 cm. longis, superioribus subulatis minutis; hypanthium obovoideum
1.5-2 mm. longum dense minute pallido-strigillosum; calyx maxime
inaequaliter 4-lobus, lobis 3 linearibus vel lineari-subulatis ca.
STUDIES OF CENTRAL AMERICAN PLANTS III 27
3 mm. longis, quarto lanceolato-oblongo 5-6 mm. longo obtuso,
omnibus sparse strigillosis vel fere glabris; corolla in alabastro apice
obtusissima extus dense minuteque strigillosa. Guatemala: Dept.
Zacapa, cloud forest in ravine bordering Quebrada Alejandria,
summit of Sierra de las Minas, vicinity of Finca Alejandria, alt.
2,500 meters, October, 1939, Julian A. Steyermark 29906 (type in
Herb. Field Mus.).
The most closely related species is R. aetheocalymna Donn. Smith,
of Alta Verapaz, in which the much larger leaves are oval, and the
small calyx lobes are oval. The present species is dedicated to
Don Alejandro Cordova, distinguished journalist of Guatemala, and
proprietor of Finca Alejandria, where the junior author was a guest
when the type material was collected.
COMPOSITAE
Liabum tajumulcense Standl. & Steyerm., sp. nov. Frutex
4.5-5 m. altus, ramulis arachnoideo-tomentosis; folia non visa;
capitula discoidea in paniculas multifloras contractas thyrsoideas
disposita, ramis dense ochraceo-tomentosis; bracteae ovatae vel
lanceolatae 4-6 mm. longae extus dense lanuginosae intus glabrae,
pedicellis dense tomentosis; involucrum campanulatum 4-seriatum
14-16 mm. altum, 17-20 mm. latum, bracteis 5-13 mm. longis
gradatis ovato-lanceolatis usque lineari-lanceolatis acuminatis, extus
ochraceo-lanuginosis; discus luteus 1.5-2 cm. altus 1.5-2 cm. latus;
corollae 12-13 mm. longae; achaenia immatura 5-costata compressa
4 mm. longa 1.5 mm. lata, dense sericeo-hirsutula; pappi setae in-
aequales, interiores 35-40 corollam fere aequantes 10-11 mm.
longae, exterioribus brevioribus. Guatemala: Dept. San Marcos:
Moist thicket bordering stream, barrancos southwest of Tajumulco,
northwestern slopes of Volcan de Tajumulco, alt. 2,300-2,500 meters,
February, 1940, Julian A. Steyermark 36543 (type in Herb. Field
Mus.).
Apparently related to L. Deamii Rob. & Bartl., but differing in
its much larger heads and flowers; distinguished from L. Klattii Rob.
& Greenm. by the densely lanuginose and more numerous involucral
bracts and broader heads.
Stevia vulcanicola Standl. & Steyerm., sp. nov. Frutex 2-3-
metralis, caulibus glutinosis; folia opposita petiolata elliptico-ovata
usque ovata viscido-glutinosa pinnatinervia, 15-30 cm. longa 7-12
cm. lata, arete crenato-dentata, supra puberula, acuta vel acuminata,
basi abrupte cuneatim contracta, petiolo anguste alato 2-9 cm. longo;
28 FIELD MUSEUM OF NATURAL HISTORYBOTANY, VOL. 23
inflorescentia corymbosa terminalis dense fastigiato-ramosa, ramis
glabratis vel villosis; capitula 11-14 mm. alta sessilia vel breviter
pedicellata; involucrum pallide viride, bracteis linearibus acuminatis
vel cuspidatis sparse dorso hispidulis; corolla rosea vel pallide lilacina
6.5-7 mm. longa glandulosa; achaenia linearia ad angulos sparse
hispidula fuscescentia 5.5-7 mm. longa, corona brevi acute dentata;
setae nullae. Guatemala: Dept. Quezaltenango: Pine-fir forest,
Volcan de Santo Tomas, alt. 2,500-3,700 meters, January, 1940,
Julian A. Steyermark 34757 (type in Herb. Field Mus.); also numer-
ous other collections from the departments of Quezaltenango and
San Marcos.
Similar to S. polycephala Bertol., but differing in its larger broader
leaves, more contracted at the base, the margins with more salient
and regular serrations.
UNIVERSITY OP n-un =
3 01 12 049897595
